AF 1 a a alii il hk et he POW MR Ae ee SW Eh et aw ee ee “t : & AA a < oe ar — \ am . a in i _ li fa if 7 ft ~ 2 Ag lua i ~~ HARVARD UNIVERSITY. LIBRARY OF THE MUSEUM OF COMPARATIVE ZOOLOGY. ane ar UA 4 t i F , URAC aye, ae a aye . ive ) Nv hs ; Ti Webs Ne eae egg aed a eh OS Ae t sO) Can ¥ =! tenes) 1 Vy on eg Laue > 7 Pei ane bey, ae Ye ih uy Te Ba ate 2 Many me 8 i" er ar it " ue on rae ed - Wn Fi EXTINCT ANIMALS Extinct Animals By BE) RAY LANKESTER, M-A., LL.D... F.R.S, Director of the Natural History Departments of the British Museum; Correspondent of the Institute of France WiTH 218 ILLUSTRATIONS NEW YORK HENRY HOLT AND COMPANY 1905 BUTLER & TANNER, THE SELWOOD PRINTING WORKS, FROME, AND LONDON, PREFACE HIS volume is a corrected shorthand report of the course of lectures adapted to a juvenile audience given by me during the Christ- mas holidays 1903-4 at the Royal Institution, London. The lantern slides which I used in the lectures have been converted into process blocks. Many of these were photographs specially prepared under my direction for the lectures, and are from specimens in the Natural History Museum. My desire was, as far as possible, to illustrate what I said by photo- graphs taken from actual specimens. Some of these have come out fairly well as process- blocks. For several of the slides and figures I have to thank my friend and colleague Dr. Arthur Smith Woodward, Keeper of the Geological Department of the Museum, to whom I am greatly indebted for kind help in many ways in regard to these lectures. Ihave Vv PREFACE also to thank other friends for the loan of lantern-slides and consequent process-blocks, viz.. Mr. R. lLydekker, Dr. lBather, Dr. Andrews and Mr. Pyecraft of the British Museum, and Professor Sollas of Oxford. Jam also indebted to the Trustees of the British Museum for permission to use several figures of extinct animals taken from the guide-books to the Natural History Museum, published by their order, to Messrs. Macmillan & Co., and to Mr. John Murray. I trust that this volume will not be regarded as anything more ambitious than an attempt to excite in young people an interest in a most fascinating study, and that it will be understood that it does not profess to give more than a peep at the strange and won- derful history of extinct animals. EK. RAY LANKESTER. 1905. CONTENTS CHAPTER I ANIMALS WHICH HAVE LATELY BECOME EXTINcCT— THE STRATA OF THE EArtTuH’s CRUST CHAPTER II STRATA AND LAanD SuRFACES—TEETH AND BoNES— Extinct Men—F tint IMpLEMENTS—THE Mam- MOTH ELEPHANTS AND MaAstoDON—CLASSIFT- CATION OF ANIMALS CHAPTER III THE ANCESTRAL HistoRY oF ELEPHANTS—EXTINCT HorsEs AND RHINOCEROSES—THE ARSINOITHE- RIUM , , 5 CHAPTER IV EXTINCT GIRAFFES AND THE OKAPI—THE GIANT SLOTHS OF SOUTH AMERICA AND THE GIANT Kan- GAROOS OF AUSTRALIA vii PAGE 59 103 CONTENTS CHAPTER V PAGE THe Great Extinct REpTILES—DINOSAURS FROM THE OOLITES—THE PARIASAURUS AND INOSTRAN- SEVIA FROM THE TRIAS OF NorTH RUSSIA AND South ArricAa—MariIneé REPTILES ; . 100 CHAPTER VI Extinct FisHES—BELEMNITES — LINGULA — TRILO- BITES—SCORPIONS AND STONE LILIES . . 245 vill Mist. .OF 1LEUST RATIONS Portrait of the Author b ‘ : . Frontispiece Fic. PAGE 1. A number of bones of extinct animals embedded in rock, from Pikermi near Athens : : , 2 2. Head of an Ichthyosaurus, from the Liassic rocks of Lyme Regis : ; ‘ : : 5 6 3. The skeleton of the Megatherium found in the alluvial sands of the Argentine Republic . : ii 4, The skeleton of a gigantic extinct rat-like animal— the Toxodon—from the Argentine Republic : 8 5. Photographs of two skulls of Rhinoceroses in the Natural History Museum ; : ‘ : 10 i=) . Photograph of the thigh-bone of the great extinct reptile, Atlantosaurus, from the oainnecic rocks of the United States of America . : ‘ 11 7. The Common Grey Wolf (Canis lupus) of eS once common in England é : 14 8. Photograph of a mounted specimen of the Beaver 15 9. Skull of the great extinct Bull, the Bos Primigenius or the Urus, or Aurochs . Wy 1X 15a. Egg of the Great Auk, of the natural size 16. 17. 18. 19: 20. LIST OF ILLUSTRATIONS Photograph of two Giraffes from life . . Photograph of the living Quagga (Equus quagga), . Photograph of the living Zebra (Equus birchelli) . Steller’s drawing of the Sea-cow discovered by him and called Rhytina stelleri Photographed with its egg Reproduction of a picture of the Dodo, mii ve Roland Savery from life, in 1626 ; . Photograph of a skull of Steller’s Sea-cow . The Great Auk, or Gure-fowl (Alca impennis). A nearly complete skeleton of the Dodo, put to- gether from bones collected by Mr. George Clark in a marshy pool in Mauritius The living Giant Tortoise of the Court House, Mauritius F The ruins of the ancient Roman public buildings at Puzzuoli (Puteoli) near Naples One of the three columns of the “ Puzzuoli . . . 5 Empire (third century) . . Puzzuoli at the present day . Puzzuoli in the ninth century . temple ” at . Puzzuoli or Puteoli in the time of the Roman . Imaginary view of Spanish sailors carving an inscription on rocks at sea-level in 1600 4.D., on the Chilian coast . The same rocks as they would appear in 1900, raised 150 feet above the sea-level by an imperceptible movement of six inches a year . x 26 27 29 32 39 40 LIST OF ILLUSTRATIONS Fic. 26. 28. 29. 30. 31. 32. 33. 34. 35. 36. 37. 38. 39. 40. Map to show the effect of elevation of the earth’s surface on the distribution of land and water in Western Europe . The real test of Geology : an attempt to determine the distribution of land and water in past ages. Photograph of a slab of Bognor Rock (Lower Eocene) showing embedded marine shells Skeleton of a tapir-like animal (Palzotherium) as found embedded in ealcareous rock at Montmartre, Paris Wings of a Dragon-fly preserved in the ancient limestone of the Carboniferous period or Coal- bearing rocks . Pterodactyle skeleton preserved in Lithographic limestone, showing the impression of the mem- brane of the wings . A jelly-fish (similar to the recent Awrelia auriia) preserved in Lithographic limestone Alternate layers of hard and soft rock (“‘ strata ’’) forming the sea-cliff at Lyme Regis Tilted strata of the Chalk at Seaford, Sussex. Strata of the cliff at Lyme Regis Diagram to show the effect of the bending or un- dulation of the earth’s crust : : : Ripple-marks preserved in ancient Triassic strata . Bird-like footprints on a slab of Triassic rock from Connecticut, U.S.A. Three-toed~ footprint (probably of Iguanodon) from the ,;Wealden Sandstone of the Isle of Wight Slab of Triassic rock from Cheshire, showing hand- like five-fingered footprints . : ° . x1 PAGE 4] 43 45 46 47 47 ou ou LIST OF ILLUSTRATIONS Fic. PAGE 41. A tabular view of the strata of the earth’s crust, showing the relative thickness of each “‘ system”’ or group of strata, and the position in which important animal remains have been discovered. 60 42. Map of the World, showing its division into great provinces and regions characterised by the presence of different kinds of animals : , 63 43. Photograph of the original piece (seven inches long) of a thigh bone of a gigantic bird, from the examination of which Sir Richard Owen inferred the former existence of a ee flightless bird in New Zealand . : : : - 68 44, Photograph of Sir Richard Owen standing beside the restored skeleton of the New Zealand Moa (Dinornis maximus) 3 - : - : 69 45. Photograph of the skeleton of Manand Horse from a group, prepared under the direction of Sir William Flower for the Natural History Museum 71 46. Photograph of the back of a skull of an Ox ,. ‘ie 47. Photograph of the back of a Crocodile’s skull . 74 48. Drawing of the auditory organ or internal ear of man . ‘ : é : i : 75 49. Photograph from a section through the bone in which the soft internal ear is lodged . 5 75 50. Photograph from preparations of the upper and lower jaw of a Pig, to show the teeth in position. 78 51. Photograph of a preparation of the teeth of the upper and lower jaw of a Pig. = - 79 52. Photograph of a preparation of the upper and lower jawbone ofman : ; : ; - 80 53. Skull of the Clouded Tiger ° . ° o) anol 54. Photograph of the skull of the Coypu Rat . - 82 xii LIST OF ILLUSTRATIONS Fig. 55. Jaws of the Gharial, an Indian crocodile 56. Photograph of the skull and lower a of a true Crocodile 5 57. Enlarged representation of the lower jaw of a small mammal (Amphitheriwm prevostii) from the ‘Stonesfield slate of Jurassic (Oolite) age near Oxford . 58. Photographs of two flint Se Ee of the’ Palzo- lithic age : : ‘ 59. Photograph of the top of the skull or “ calvaria ”’ of=the so-called Monkey-man, P2thecanthropus, discovered in Java . 594. Photograph cf a human skull of modern to aes race . - 60. Engravings on ivory and boneXjmade by ancient men, who lived in caves in the South of France at the time when the mammoth, reindeer, bear and hyena inhabited Europe 61. Engraving on ivory found in a cave in the South of France 62. The skeleton of the Mammoth found frozen in Siberia . 63. Skeleton of a male of the giant Irish deer (Cervus giganteus) dug up from peat in Ireland 64. An imaginary picture of the Mammoth (Elephas primigenius) as it appeared in life : 65. Photograph from life of the Indian tas (Zlephus maximus), incompletely grown 66. Photograph of a young specimen of ‘the African Elephant (Elephas africanus) from life 67. Two tusks of Elephants photographed from speci- mens in the Natural History Museum : Xili 84 86 89 91 92 93 94 96 97 98 99 LIST OF ILLUSTRATIONS Fie. 68. Skeleton of the American Mastodon (Mastodon americanus), from a drawing by the late Professor Marsh : : ' : 69. Skeleton of Indian Elephant (Elephas maximus) 70. Skull of an adult Indian Elephant 71. Photographs of skulls of a Bull-dog on the left and of a Greyhound on the right to show the shortening of the bones of the face in the first . 72. Photograph of the skull of the American Mastodon (Mastodon americanus), frcm the specimen in the Natural History Museum 73. Skull of a new-born Indian Elephant 74. Section of the skull of a young Indian Elephant 75. Section of a half-grown Indian Elephant’s skull 76. Lower jaw of an Indian Elephant . 76a. The last molar of the lower jaw of a Mammoth 77. Lower jaw of an adult African Elephant 78. Lower jaw of the American Mastodon 78a. Molar teeth of Mastodon arvernensis, photographed from specimens found in the Red Crag of Suffolk. 79. Photograph of the complete skeleton of Mastodon (Tetrabelodon) angustidens, from the Miocene strata of the South of France 80. Restored representation of the skull and lower jaw of Mastodon (Tetratelodon) angustidens, from a drawing prepared by Dr. Henry Woodward, F.R.S. 81. The skull of Dinotherium giganteum, Kemp, from the Miocene of Eppelsheim, near, Worms, on the Rhine ‘ : ; X1V PAGE 100 101 104 105 106 107 108 109 110 111 112 113 115 116 118 LIST OF ILLUSTRATIONS Fic. 82. 83. 84. 85. 86. 87. 88. 89. 90. 91. 92. 93. 94. 95. 96. oT. Drawing representing the probable appearance in life of the Tetrabelodon angustidens A drawing of the head of Tetrabelodon anqgustidens with open mouth and uplifted “ trunk.’’. : Drawing of the head of the African Elephant, with uplifted trunk A scene in the Fayum Desert, showing the remains of silicified trees, embedded in the sands Profile views of a series of Elephant ancestors, from drawings by Dr. Andrews Lower jaws of extinct Elephants, from drawings by Dr. Andrews Profile and palatine views of the skull of Meri- therium Lyonsi, as restored by Dr. Andrews The Meritherium, discovered by Dr. Andrews Photograph of a model of a ee ed Sit horse, by Vashtag . Hind- and fore-foot of an English cart-horse Hind-foot and fore-foot of the horse-ancestor, Hyracotherium The hind- and the fore-foot of Hipparion, one of the three-toed ancestors of the horse The skeleton of Hyracotherium, an ancestor of the modern horse, found in Eocene strata. Restoration of the probable appearance of the Hyracotherium : ‘ Skeleton of the Phenacodus, a five-toed Eocene ani- mal, related to the ancestor of the Horse Cheek-teeth or molars of the upper and lower jaw, left side, of Mesohippus Bairdii, from the Middle Oligocene of South Dakota XV PAGE 119 137 138 139 140 141 141 LIST OF ILLUSTRATIONS Fic. 98. 99. 100. 107 108. 109. 110. cB UUIE 112. 113. 114. Upper molar tooth of a recent Horse . The skeleton of Rhinoceros antiquitatis, the Woolly Rhinoceros of the late Pleistocene period in Europe and Siberia Photograph of a stuffed specimen of the Square- mouthed African Rhinoceros (R. simus F . Skeleton of Titanotherium (Brontops) robustum, from the Lower Miocene of Dakota 2. Photograph of a skull of Titanotherium . Side-viewJof the skull of Titanotherium . Skeleton of Déinoceras mirabile . Probable appearance in life of the Dznoceras mirabile of North America . Photographs of plaster casts of the brain-cavity of (A) Dinoceras, (B) Hippopotamus, (C) Horse, and (D) Rhinoceros : : : ; : : Drawing of the skull of Arsindithertum Tittelli (Beadnell) ; : 3 : ; A drawing showing the probable appearance in life of Arsinditherium . 5 : ‘ Drawing of the head of the Five-horned Giraffe Photograph of the skull of the Five-horned Giraffe Front teeth of the lower ou of the Giraffe and allied animals : : Photograph of a restored skull of the Sivatherium Photograph of the skull of the Samotherium . Restored skeleton of the giraffe-like animal Hella- dotherium Xvi PAGE 142 143 144 145 146 147 148 149 150 152 161 162 LIST OF ILLUSTRATIONS Fic. PAGE 115. Photograph of the specimen of the Okapii(Okapia erichsoni) obtained by Sir Harry Johnston near the Semliki river in Central Africa. : : . 163 116. Photograph of a skull of a male Okapi . : a = 164 117. Photograph of the two “* bandoliers ”’ cut from the striped part of the skin of an Okapi . ; . 165 118. Photograph of a stuffed specimen of the two-toed Sloth (Cholepus Bue): hanging from a branch of atree. : ; : . 168 119. Photograph of a stuffed specimen of the Hairy Armadillo or Peludo (Dasypus villosus) . . 169 120. Drawing of the skeleton of the great extinct armadillo-like animal called Glyptodon . LTO 121. Probable appearance in life of the Giant Ground Sloth, the Megatherium giganteum . : a lll 1225 the ekeleton of Mylodon robustus, one of the giant Ground Sloths of the Argentine. 5 ‘ - 173 123. View, looking outwards, from the mouth of the cavern on the fiord of the Ultima Speranza in Southern Patagonia, in which have been found the skin and hair and the bones with cartilage, blood and tendon and the dung of the Mylodon and other animals . : E : ; : . (174 124. Photograph of a piece of the skin of the Mylodon 175 125. The under side of the same piece of skin as that shown in Fig. 124 7 : : ; fy l6 126. Photograph of various specimens found with the remains of the Mylodon in the Ultima Speranza cave : : a : 5 : . ne ia 127. Photograph of remains of Mylodon from the cave of the Ultima Speranza . 3 : ; + 78 128. Photograph of a “ barrel-full of bones”? obtained by prospectors from the cave of Ultima Speranza 179 XVil b Fic. 129. 130. 131. 132. 133. 134. 135. 136. 137. 138. 139. 140. 141. 142. 143. LIST OF ILLUSTRATIONS Photograph having the same history as that shown in Fig. 128 Photograph of three pellets of the dung of the Mylodon from the cave of Ultima Speranza Photographs of the leg-bone (tibia) of Mylodon, from the cave of Ultima Speranza . : : Drawing of the skull of the Giant Australian Marsupial, Diprotodon The restoration of the skeleton of Diprotodon, as drawn by the late Sir Richard Owen Photograph of the morass or lake in South Australia in which the remains of several specimens of Diprotodon have been recently discovered View of the upper surface of the right hind-foot of Diprotodon, as discovered by Professor Stirling of Adelaide, South Australia Lower jaws of the ancient Mammals, Dromatherium (upper—Triassic), and Dryolestes (lower—Juras- sic) : : Photograph of a cast taken from life of the New Zealand lizard Tua-tara, known as Sphenodon punctatus Phrynosoma orbiculare (Mexican Horned Lizard, or Horned Toad) Chlamydosaurus kengi, from Queensland, Australia Zonurus giganteus (Great Girdled Lizard) Drawing of the skeleton of Iguanodon bernissar- tensis. : Probable appearance of the Iguanodon in its living condition : : Two teeth of Iguanodon mantelli . XVili PAGE 180 186 187 188 189 193 194 195 196 197 198 199 Fig. 144. 145. 146. 147. 143. LIST OF ILLUSTRATIONS A portion of the upper jaw of the recent lizard Iguana . : : Photograph of the skull of an Iguanodon . Drawing of the skeleton of a carnivorousiDinosaur, the Megalosaurus Drawing of a completely restored skeleton of the Brontosaurus Probable appearance of the Ceteosaurus (and of the closely similar Divlodochus and Brontosaurus) in life , . Drawing of the appearance in life of the three- horned Dinosaur, Triceratops . . Probable appearance in life of the Jurassic Dinosaur, Stegosaurus . Photograph of the skeleton of Parisaurus . . Probable appearance in life of the Theromorph Reptile, Dimetrodon . View of one of the dark patches in the cliffs of the river Dwina . One of the nodules showing the form of the em- bedded skeleton . Peasants working on the face of the cliff near Archangel and removing nodules containing the skeletons of great reptiles . 2 . Professor Amalitzki’s workshop in Warsaw . . A series of skeletons of Parisaurus removed bit by bit from Archangel nodules and mounted as de- tached specimens by Professor Amalitzki . Photograph of a skeleton of Pariasaurus, removed from an enveloping nodule and mounted by Professor Amalitzki ab: PAGE 200 202 203 205 206 207 208 211 212 213 214 215 216 217 218 LIST OF ILLUSTRATIONS Fie. PAG! 159. Photograph by Professor Amalitzki on a larger seale of askull of a Pariasaurus from an Archangel nodule 219 160. Skeleton of a huge carnivorous beast of prey, the reptile named Inostransevia 220 161. Skull of the vigantic Theromorph Carnivorous Reptile, Inostransevia 221 162. Photograph of another skull of Inostransevia 222 163. Photograph of a skeleton of a Plesiosaurus . 223 164. Plesiosaurus as it probably appeared when alive 224 165. Photograph of a skeleton of the large- Pas Ichthyosaurus : 225 166. Drawing to show the probable appearance of an Ichthyosaurus swimming beneath the surface of the sea . 226 167. Photograph of the upper surface of the skull of an Ichthyosaurus 3 228 168. Side view of the skeleton of an Ichthyosaurus 229 169. Photograph of a restoration of the skeleton of the great Pterodactyle (Pteranodon longiceps) 230 170. The great Ha eae Pteranodon as it pass : in flight . : 231 171. Photographs of three wings for comparison of their structure A 233 172. Probable appearance in life of two kinds of Jurassic Pterodactyles (Dimorphodon and Khampho- rhynchus) ‘ : : : : . 235 173. Restored skeleton of the toothed Bird Ichthyornis 237 174. The Berlin specimen of the Archwopteryx litho- graphica 238 xX Fie. Lizhss 176. 186. 187. 188. 189. 190. LIST OF ILLUSTRATIONS Photographs to one scale of the South American Cariama and the skull of the gigantic extinct Phororachus Photographs to one scale of the Apteryx, the Ostrich and the giant Moa of New Zealand, each with its ees - The hard bony scales of a Ganoid Fish . Photograph of a dried skin of the Polypterus of the Nile . A fossil Ganoid Fish, as discovered embedded in rock . Outline drawing of the extinct Ganoid Fish Osteo- leps . The Australian Lung-fish Ceratodus . The extinct Devonian Fish Dipterus . . Outline drawings of the extinct fish Pterichthys . Photograph of a cardboard model of Pterichthys . The upper figure is a restored outline of the curious Devonian fish Coeceosteus Photograph from the original specimen of Cephalas- pis lyeli, preserved in the Natural History Museum : , Drawings of the head-shield of the fossil fish Pteraspis Photograph (of the natural size) of a specimen showing parts of the upper and lower head-shields of Pteraspis crouchii, with ten rows of lozenge- shaped scales attached Photographs of models of the Devonian Fish Drepanaspis, in the Natura] History Museum Outline drawing of the Silurian fish Birkenia XX1 PAGE Fic. 191. 192. LIST OF ILLUSTRATIONS Outline drawing of Lasanius Photograph of the jaws of a large recent Shark (Carcharodon rondeletiz) 1924. Photograph of the natural size of a tooth of the 193. great Shark, Carcharodon megalodon Ammonites (Aegoceras) capricornus . 1934. Shell of the Pearly Nautilus 194. 198. 199: Divided shell of the Pearly Nautilus . The shell of Ancyloceras matheronianum . Belemnites hastatus, from the Oxford Clay (Jurassic) . Restored drawing of the animal in which the ‘** Belemnite ’’ is formed . Loligo media, a cuttle-fish or squid now living in British seas Lingula (Lingulella) davisit, of the natural size, embedded in the slaty rock of Port Madoc, North Wales. ‘ ‘ ‘ : : : ; . One of the most ancient Trilobites known (Cono- coryphe lyellc) . Drawing of Triarthrus becki . The Desert Scorpion (Buthus australis) . Drawing of the remains of a Scorpion (Palewophonus huntert) . Completed drawing of the Scotch Silurian Scorpion (Paleophonus hunteri) . Completed drawing of the Silurian Scorpion of Gothland (Paleophonus nuncius) XXli BIG: 206. 207. 208. 209. LIST OF ILLUSTRATIONS View of the anterior part of a recent Scorpion from below View from below of the anterior part of the great Silurian Scorpion-like creature Pterygotus osilien- Sis 5 3 : Photograph of a restored model of Stylonurus lacoanus) Eurypterus fischeri, a marine Scorpion-like animal from the Silurian rocks of Rootzikul . Dorsal view of the King-Crab (Limulus polyphemus, Linnzeus), one-fourth the size of nature. . Diagram of the dorsal surface of a King-Crab 2. Diagram of the ventral surface of the same King- Crab . Dorsal view of the eighteen segments and post-anal spine or sting . Slab containing Pentacrinus hemeri . Photograph of a block of Limestone of the Car- boniferous, showing several kinds of stone-lilies or Encrinites . . Encrinus fossilis, of Blamenbach, the original ‘“* Stone-lily.”’ . . The living British Encrinite, the minute young of the Feather Star-fish (Comatula or Antedon), . Drawing by Mr. Berjeau from an actual specimen of the Feather Star-fish (Comatula or Antedon TOSaCeQ) . : : XXiil PAGE 279 bo Ne) bo a CHER i ANIMALS WHICH HAVE LATELY BECOME EX- SEN Ci TEE STRATA OF THE EARTH'S CRUST XTINCT animals are animals which no longer exist in a living state. Of course a vast number of individual animals, and men too, become extinguished, or extinct, in the course of every year, every month and every day. But the extinct animals of which I wish to speak in these lectures are extinct kinds of animals, kinds of animals which no longer exist on the surface of the globe in a living state, although once they flourished and held their own. We know of some of them by tradition. The records of men of past ages who have seen some animals, now extinct, and have written about them, and even drawn them, have by human care been passed on to the present day. We I B EXTINCT ANIMALS know of other extinct animals by finding their bones buried in the ground, some quite near the surface, others deeper in the rocks, far down in the depths of the earth. Such bones may be dug out. There is a sample of such bones Fic. 1.—A number of bones of extinct animals embedded in rock, from Pikermi near Athens. Photographed from a specimen in the Natural History Museum. found buried in the earth, photographed as our first illustration (Fig. 1). Many of these bones have been so big, so huge, that they have led to the notion of the existence of giants in former days, it not having occurred, apparently, to those who 2 GREAT AGE OF THEIR REMAINS found them, that they were the bones of extinct animals and not of a great race of men. The indications given by buried remains of a condition of the world which has passed away, as, for instance, in the great buried town of Pompeii, and some of the buried cities of Egypt, excite, when they are dug up, the greatest interest. From the records still preserved to us, we try to find out what was the meaning of the particular objects found, what were the nature and the life of the men to whom they belonged. The same kind of interest belongs to the remains of extinct animals that we dig up, only that many of them are far older than any remains of man ever found. We speak of the remains of an ancient Egyptian city as being some thousands of years old; but the remains of many animals to which I shall have to refer in these lectures have to be estimated, not by thousands of years, but by millions of years ; so many years in fact that no numbers with which we are familiar will suffice to bring the facts to the minds of my readers. Far down in the depths of the earth we find the remains, in a well-preserved condition, of the bones and teeth of such animals; we are 3 EXTINCT ANIMALS able to tell what kind of animals they were, where they lived, what they fed upon, how they moved, and, in fact, their whole general appear- ance. It is urged by some educationists—I myself do not agree with them—that we should present knowledge to young people in a logical order ; and that before talking to young or uninstructed people about extinct animals you ought to ad- minister to them a complete course of instruc- tion concerning living animals; that beginners must learn the nature of the structure of living animals, and must study the geography and history of the crust of the ground in which the remains of extinct animals are found, before they can look with any intelligence on extinct animals. That is an opinion which exists. But I do not believe in such a method. The logical method of instruction or study is in my judg- ment a mistaken one. The whole art of educa- tion consists in exciting the desire to know. By showing something wonderful, mysterious, as- tonishing and marvellous, dug from the earth beneath our feet we may awaken the desire to understand and learn more about that thing. The strangeness of the bones and teeth of 4 A FASCINATING SKULL extinct animals will lead a boy or girl on to learn- ing about the bones and teeth of living animals in order to make a comparison, and thus to learn- ing more concerning the strange remains dug up. I believe that is usually the case. It certainly was the case with myself. When I was very young, younger than, or as young as any of my readers, I used to be taken by a very kind lady, my governess, to the Natural History Museum of the day, which was then in a remote part of London called Bloomsbury, whence it has been removed to Cromwell Road, Kensington. I was absolutely fascinated as a child with the remains I saw of strange extinct animals. And it is my hope that the boys and girls who read these pages may share some of this interest and fascination, and that they will pass from these lectures to see the actual specimens which are placed on view at the Natural History Museum. These lectures are indeed little more than a sort of invitation to you all to go and see the real things at Cromwell Road, of which I can only show you photographs in this book. I will now show you a portrait of a creature which has always fascinated me with its stony stare. It is the head of an Ichthyosaurus dug oO EXTINCT ANIMALS out of the rock in the South of England, at Lyme Regis, many years ago (Fig. 2). The eye is peculiarly well preserved. The circle of bony plates, similar to those found in the eyes of birds, give an expression of interest which few fossils can boast of. It was dug out of the rocks by a wonderful lady, Miss Anning, who at the beginning of the last century secured a Fic. 2.—Head of an Ichthyosaurus, from the Liassic rocks of Lyme Regis. Photographed from the original specimen in the Natural History Museum. The head is three feet six inches long. great number of such remains in the cliffs on the sea-shore. For many years the front part of this specimen was missing, but eventually it was found and dug out of the rocks. I shall have more to say later about creatures of this kind, Another creature which fascinated me is shown here as it is exhibited in the East court 6 THE GIANT SLOTH Fic. 3.—The skeleton of the Megatherium found in the alluvial sands of the Argentine Republic, South America. Photo- graphed from the cast in the Natural History Museum. The skeleton stands fourteen feet high. 7 ‘(uMosN YS oy} Jo sooysnay, oy Aq yuoyT st ons sty) ‘oof ouru [rez oy} OF yous oY} WLOAT yj3ue'T += “BoLoUlyYyNog ‘oulpUOS.1y oy} UIO1J—UOpOoxO [, OYJ —[VULLUe O¥I[-7B4 YOUTZXO OLYUBDIS B JO UOYI[OYS OUL—F “OW iw A LONDON RHINOCEROS of the Museum (Fig. 3). It is similar in structure and nature to the sloth. But instead of living on a tree it stood on the ground, and pulled the tree down to it, in order to feed on the young branches. The skeletons of a great many of these huge sloths have been found in the gravel of South America. Another strange great creature is revealed to us by this skeleton (Fig. 4), like a huge guinea pig with tremendous chisel-like teeth in front. It also is found in South America. This is the Toxodon. The next picture (Fig. 5) I have here shows the skulls of two rhinoceroses. The lower one is the skull of an African rhinoceros, a living beast known as the square-mouthed or white rhinoceros—called white apparently, not because he is black, but in spite of the fact that he is black. As a matter of fact he sometimes has a number of white patches. But it suffices to know him as the square-mouthed rhinoceros. The upper specimen is the skull and lower jaw of a rhinoceros, dug up last year in the City of London in Whitefriars, under the office of the well-known newspaper the Daily Chronicle. Digging in the mud and clay there, S EXTINCT ANIMALS the workmen came upon this rhinoceros skull. Many such have been found in English river Fic. 5.—Photographs of two skulls of Rhinoceroses in the Natural History Museum. The upper one was dug out of the Thames clay in Whitefriars, London, and is that of the species known as Rhinoceros antiquitatis. The lower one is that of the living African square-mouthed Rhinoceros (R. stmus), which is more like the fossil one than is any other living rhinoceros. gravels, and we know accordingly that such animals used to exist on the banks of the 10 Fic. 6.—From a cast in the Natural History Museum. Photo- graph of the thigh-bone of the great extinct reptile, Atlantosaurus, from the Jurassic rocks of the United States of America. The thigh-bone is six feet in length : that of a very big elephant is barely four feet. el EXTINCT ANIMALS Thames many thousands of years ago. That specimen also is in the Natural History Museum. Here (Fig. 6) you have a thigh bone; you can see how enormous it is from the figure of the full-grown man beside it. That is the thigh bone of a huge kind of reptile, bigger than the ordinary elephant, or the biggest African elephant, without counting the rep- tile’s tail, Such remains have been found in England; but the largest have been found in the United States. These are just a few samples of the remains of extinct animals, and indicate the kind of creatures I want to tell you about. Of course I cannot in these pages refer to all the many thousands of kinds of extinct animals which are known ; I can only hope to show you pictures of a few samples of these things, which, how- ever, I hope will suffice to induce you to look further into the matter, to look at the real specimens, and to read more elaborate books, and thus come to feel the same interest and pleasure in examining them that I do myself. The world upon the surface of which we live has been for millions of years always changing. Nothing is to-day as it was even one hundred 12 INCESSANT CHANGES years ago. A thousand years brings about enormous changes, quite a different state of things in fact. There are now cities where forests were growing. Animals which existed a thousand years ago have altogether gone. And this history of change has been going on, not merely for a thousand years, but for hundreds and thousands and millions of years. The changes have been incessant, and have been very great. The difficulty in this study of extinct animals and in the geology connected with it is to think of long enough lapses of time. If you look at that clock you cannot see the hand moving, and yet it is moving. And thus even in a human lifetime you will hardly notice any difference in the rivers and the sea-shore and the cliffs. But if you range over a long enough time, say a thousand years or several thousands of years, and compare the condition which existed a thousand years ago with what exists to-day you will be able to observe great change. The difficulty is to realize this change, for it comes about too slowly for our short lives to give us any real definite experience of it, just as we fail to see the hands of the clock moving when 13 EXTINCT ANIMALS we glance at them for a second. Throughout these lectures I want you always to bear that in mind. We know of animals even now which are becoming extinct. In this country we have Fic. 7.—The Common Grey Wolf (Canis lwpus) of Europe, once common in England, but now extinct there. historical records of animals that have become extinct. I will show you one which used to exist in this country. This creature, the grey wolf (Fig. 7), existed 14 THE BEAVER ‘umMosnyy ATOISTET [RANQVN oY ULOTZ ¢ £1 N GION IOABI_ 94} JO Uoutloods pozUNnOW vB Jo YdeaIsojoyq—'s “YI EXTINCT ANIMALS in England till the time of Henry VIII., at the end of the fifteenth century, and 150 years later in Scotland and Ireland. But it was entirely exterminated by human beings, on account of its rapacious and dangerous habits. Though it is extinct in England, it still exists in France, Spain, Germany and Russia. Here is another animal (Fig. 8), the beaver, which used to exist in England, and was found as late as the sixteenth century in Wales. It still exists in France, on the banks of the streams at the mouth of the river Rhone ; also in Russia and Scandinavia. In America, in Canada, beavers are still more abundant. Another creature which, records tell us, existed all over Europe, and which has ceased to exist, is the great bull or Urus of Julius Caesar (Fig. 9). He mentions it as existing wild in different parts of Europe, and says it was nearly as big as an elephant. Well, no such great wild ox now exists in Europe. The last was killed near Warsaw in 1627. All we have now are the breeds derived partly from this, partly from other kinds of bulls, which, are quite changed in their general appear- ance. Some of the more or less wild cattle 16 THE GREAT BULL OF CASAR in different parts of England, for instance those on Lord Tanqueray’s and the Duke of Hamil- ton’s estates, are supposed by some persons to be the remains of this race of wild oxen. But this is probably a mistake. They are really Fic. 9.—Skull of the great extinct Bull, the Bos primigenius, or the Urus, or Aurochs. The measurement from one horn- tip to the other taken round the curves, was in some cases eight feet. The Urus stood in rare instances as much as seven feet at the shoulder ; a fair-sized Elephant stands nine feet. the remains of cattle introduced by the Romans, and have run wild. They are not the Urus of Julius Caesar, which was a good deal bigger than the largest domesticated cattle, even bigger than the white oxen of Umbria. This (Fig. 10) is another animal which has 7, Cc EXTINCT ANIMALS become extinct. But it is not a zebra, as no doubt some of you thought it must be. This is the quagga, which differs from the zebra in being striped in front only. The quagga lived £ edie at ss Casts = Fie. 10.—Photograph of the living Quagga (Hquus quagga) in the gardens of the Zoological Society in 1875, now extinct. in South Africa, and was quite common there until forty years ago. This photograph was taken from a specimen which lived in the 18 THE ZEBRA Zoological Gardens some twenty-five years ago. Fic. 11.—Photograph of a living Zebra (Equus burchelli). Its stuffed skin is preserved in the Natural History “Museum. This creature has now en- 19 EXTINCT ANIMALS tirely ceased to exist, owing to the fact that the country over which it ranged has been taken up and cultivated by white men. There are no more living quaggas anywhere. This animal has become extinguished in our own lifetime. Zebras (Fig. 11), however, are still common enough in Africa, with their beautiful stripings on the head, and on the fore as well as on the hind regions of the body and legs. Here is an animal which, it is feared, is becoming extinct—the giraffe (Fig. 12). In South Africa it has become extinct already. But sportsmen now seek it in Equatorial Africa. It is still existing in great numbers in that region, and we hope now will be properly protected by Government. Two new and well-mounted specimens have recently been put in the Natural History Museum. The neck of the giraffe is often represented as growing up from the body with a graceful curve, as is seen in the neck of the swan. But the true position of the neck is as you see here (Fig. 12). The specimens in the Natural History Museum shows this properly. This is a picture (Fig. 13) of a curious creature, an animal known as the sea-cow, found in the Aleutian Islands, between North America and 20 THE GIRAFFE Asia. It was discovered by the traveller- Fic. 12.—Photograph of two giraffes from life, showing the natural carriage of the head and neck. naturalist Steller in the eighteenth century. It was no sooner found than sailors went to the 21 EXTINCT ANIMALS islands where it existed, knocked it on the head and ate it, and in about ten years it ceased to Fic. 13.—Steller’s drawing of the Sea-cow discovered by him, and called Rhytina Stelleri. The animal was twenty feet long. exist. This picture is from Steller’s drawing. It is an enormous creature, some twenty feet Fic. 14.—Photograph of a skull of Steller’s Sea-cow, from a specimen in the Natural History Museum. long, and in shape something like a seal. But it is not in reality a seal or a whale, but belongs 22 THE GREAT AUK to a peculiar group of vegetable-feeding marine animals, the Sirenians. It has a small head, flipper-like fins, no hind limbs, and a fish-like tail. Fic. 15.—The Great Auk or Gare-fowl (Aica ampennis). Photo- graphed with its egg, from the specimens in the Natural History Museum. The skull of the same animal is shown in Fig. 14. It has no teeth, but instead bony plates. This is the picture of a celebrated animal (Fig. 15)—for you must understand that birds are animals. You will have been handed a list of the groups of animals (see the end of this chapter). 9 2 eo) EXTINCT ANIMALS T shall not have space to explain it at any length, but it gives the division of animals into groups and their relation one to another. It shows how they are classified, so that I need not refer to the classification again. This picture (Fig. 15) is the portrait of an interesting bird, the Great Auk. It is only about 23 feet high. It is like the penguin in appearance, but it is really related to the puffin and albatross. Fig. 154 shews the egg, which from time to time in the newspapers, we read of as being sold to enthusiastic egg- collectors for as much as £300. Nearly a hundred specimens of the egg of this bird are known, for it only became absolutely extinct some sixty years ago. It used to be found on the rocky islands off the North of Scotland, Shetland, Iceland and Greenland. But it has now absolutely ceased to exist. It is very difficult to say why it died out, for it had not been hunted down. Since it has become extinct we have been able to get to know about it by finding its skeleton buried in sand and guano in certain places on the coast of Newfoundland. Here (Fig. 16) is another creature, the dodo, a bird which, like Steller’s sea-cow, became 24 AUK GREAT OF THE GG E oy} UL uouTToeds wor poydeisojyoyg ‘umnosnyy AdoystFT Tengen ‘OZIS [VINZVU OYY JO SyNW yeoIH 9Yy JO SsnJ—‘ve]| ‘pILT EXTINCT ANIMALS extinct almost as soon as it became known. It was found in the island of Mauritius by the earlier explorers, first the Portuguese and then the Dutch. The bird was incapable of flying, Fic. 16.—Reproduction of a picture of the Dodo, painted by Roland Savery from life, in 1626. The bird was about three feet long from beak to tail. as it was too fat for its little wings to lift it from the ground. It was knocked on the head by the sailors and worried by the pigs they intro- duced, and was soon exterminated. About the beginning of the seventeenth century, between 26 THE DODO 1610 and 1620, specimens were brought alive to Europe and were exhibited as a show. We once possessed at Oxford a stuffed specimen, secured by that ingenious and worthy gentleman Fic. 17.—A nearly complete skeleton of the Dodo, put together from bones collected by Mr. George Clark in a marshy pool in Mauritius. In front is seen the dried foot of a specimen which was brought alive to Europe about the year 1600. The foot andjthe skeleton are in the Natural History Museum. Mr. Elias Ashmole, who gave his collections to the university 250 years ago. But asit became mouldy and eaten by insects, it was ordered, a hundred years ago, by the Vice-Chancellor and 27 EXTINCT ANIMALS Proctors of the University of Oxford, that the specimen of the dodo should be destroyed. They do not like mouldy things at Oxford. But the curator cut off the head and one foot, and kept them. This head and foot, together with another foot in London, anda skull in Copenhagen, are about all we have left of dodos seen in the living state by Europeans. But since the dodo _be- came extinct, by digging in the mud of a lake in Mauritius skeletons and bones of it have been found (Fig. 17). This (Fig. 18) is another interesting creature, whose kind is on the way to extinction. It is probably the oldest living terrestrial animal. It was brought from the Seychelles, where its kind is rapidly becoming extinct. In different oceanic islands such tortoises have been found of large size. This specimen was brought in 1764 to the island of Mauritius, and is still alive there. Thus it has been 140 years in captivity in the Court House Garden, in the Mauritius ; and how old it was when brought there it is impossible to say. A question of great interest is—** What makes animals become extinct ?”’ It is obvious in many cases that another animal, Man, inter- 28 WHY DO ANIMALS BECOME EXTINCT? feres. He either kills and eats animals, or takes their food from them, or occupies their ground, or cuts down the forests in which they live, and so on. But before man appeared on the scene there were changes going on, and different Fie. 18.—The living Giant Tortoise of the Court House, Mauritius, more than 150 years old. kinds of animals succeeded one another. We know this by finding the remains of different animals at different depths in the crust of the earth, in the different strata which have suc- ceeded one another. The cause of these changes, the cause of the extinction of animals, 29 EXTINCT ANIMALS is a very elaborate and difficult question, and one which I do not propose to deal with at any length. It is connected, of course, with the whole doctrine of the origin of the different kinds of animals. We all recognize now that there has been a gradual development of the different forms of animals by natural birth, from ancestral forms more or less like themselves. But the more remote we get from the present day, in the line of descent, the less like are the ancestors to the present form. The original parental forms have given rise to very different branches of “descent. The descendants of one ancestral form‘ have branched out in different directions: just in the same way as some person named Smith at the time of the Con- queror has ‘given rise to all sorts of Smiths. Some of them perhaps are still actually metal- workers, others have become Ministers of State and Right Honourable judges; others have great possessions; but they can all be traced back to the one original Smith. So many living animals of various appearance and form can be traced back to one ancient ancestral form, and these again to other more primitive ancestral forms. CHANGES OF LAND AND WATER The reason why the ancestral forms died out is really connected with the general change in the surface of the earth. New forms have gradually taken the place of the old forms—for no piece of land remains the same for many years. A thousand years, as I have said, in this matter is merely nothing, but even in a thousand years we get great changes in the surface of the land. Land may rise far above the sea, and what was an island become part of a continent. And what was part of a continent may partly sink, and become an island—that is, the connexion between it and the continent may become covered with water; and then the conditions of life for the animals are very much changed. Such currents as the Gulf Stream are affected by this alteration in land and water. Were certain changes to take place, the warm water of the Gulf Stream would no longer warm certain land; the climate would become colder than the animals have been accustomed to. The animals that could not stand the cold would die out, whilst those that could stand the cold would flourish.t All I + A fish—the Tile-fish—living in the Atlantic, near the North American coast, was destroyed in this way a few 31 EXTINCT ANIMALS would say is that changes in the disposition of land and water have been a great cause in changing the forms of animals and in bringing about the extinction of one set and the flourish- ing of another set. That this rising and sink- Fria. 19.—The ruins of the ancient Roman Sanne buildings at Puzzuoli (Puteoli) near Naples. The three celebrated columns are seen on the left. ing of the surface of the land really takes place I will try now to give you evidence. Here (Fig. 19) is the photograph of the Temple at Puzzuoli, near Naples, on the shore years ago by millions. It was feared it might have become extinct, but the cold current having again changed, its numbers have increased once more. Lr. Of THE ROMAN REMAINS AT PUZZUOLI —___ Fic. 20.—One of the three columns of the “‘ temple” at Puzzuoli showing (a) the portion eaten into by boring marine clam-shells, (6) the upper part, which was not submerged, and (c) part which was probably covered up by sea-sand and mud during submersion. EXTINCT ANIMALS of the Gulf of Naples. This has been celebrated for something like eighty years, ever since Mr. Babbage carefully examined and described it, and thus caused it to be largely visited by geologists. In common with most geologists, I have had the pleasure of visiting it. The three standing columns have marks of dis- coloration up to a certain height. The lower part, as shown in the diagram (Fig. 20), is full of little holes in which tiny sea creatures have burrowed holes in which there are small shells. This is so defined that it is certain these columns have stood in sea water up to that line. The evidence of that is quite complete. These columns formed part of a Temple or public palace in the great Roman town of Puteoli, which had in front of it a Roman road along by the sea-shore. Between the temple and the sea was the road. Now in Roman times that temple stood complete and very much in the same position relatively to the sea- level that it does to-day, but rather higher up. Mr. Giinther, of Oxford, examining the shore- line carefully, has found covered over by the sea the remains of the Roman road, and the remains of great blocks to which ships were 34 THE ROMAN TOWN PUTEOLI moored when they brought their wares to the town of Puteoli. I have here made a drawing of the town and the great public palace as it must have appeared in Roman times (Fig. 21). In the distance is the island of Nicida; in the foreground we have the palace and the town, the 7 Fie. 21.—Puzzuoli or Puteoli in the time of the Roman Empire (third century). The dock and public buildings are repre- sented. quay and harbour. Things existed thus in the days of the Roman empire, in the third century of our era. Then earthquakes occurred, the columns were broken, the city sank beneath the sea. Wehave no written history of this town. But it is known that in the Middle Ages, in the 39 EXTINCT ANIMALS eighth or ninth century, the whole of the coast of this part of Italy had sunk many feet, and the columns were broken and standing in the sea. This is the appearance then presented by Puteoli (Fig. 22). The coast had sunk; the remains of the road were covered by sea, and also the Fig. 22.—Puzzuoli in the ninth century, showing the sub- mergence of the land and the columns of the ruined temple or palace standing up in the sea. remains of the columns up to the height marked a on the diagram (Fig. 20). The whole land must have sunk as much as torty feet, since the temple or palace stood on high ground originally. Then it was that, while they were under water in the ninth century, the columns 6 iS) THE MODERN PUZZUOLI were bored into by sea-shells. Now some more centuries have elapsed; the ground has risen again until we have the condition shown in the photograph (Fig. 23), which gives a general view of the same region as that drawn by the use of the imagination in Figs. 21 and 22. The Fic. 23.—Photograph of Puzzuoli at the present day showing the three columns of the so-called temple of Serapis, as now seen after the retreat of the sea due to the re-elevation of the land. land rose again from the water. But the sea left its mark on the columns, showing exactly how deeply they were merged in the intervening centuries. This is considered one of the clearest and most direct proofs of the changes which take ays D4 EXTINCT ANIMALS place in the level of the ground. The change need not be a continuous or a rapid one. It took some two or three centuries for that temple to sink into the water, and a few more centuries for it to come out again. Such movement is always going on. It does not occur very obviously on our own coast. It can be seen to some extent on the Devonshire coast at Plymouth. You get evidence of it in what are called raised beaches above the level of the ocean. In Norway this kind of thing is very obvious. In South America it is going on, and has been going on at an enormous rate for the last thousand years. Probably a great part of the height of the Andes has been acquired within the last few thousand years by rapid rising. When the original sailors landed on the coast of Chili in the sixteenth century or thereabouts they are said in one spot to have chiselled on the rocks an inscription. Here you see an imaginary sketch of them doing so (Fig. 24). It is said, but I cannot find any accurate record of it, that such inscriptions have been discovered now, raised high up on the cliff (see Fig. 25). We know that many kinds of sea- shells are found 200 and 300 feet up the cliffs 38 “ THE COAST OF CHILI in this part of the world. According to the observations that have been made, the original inscription which we see the sailors cutting in Fig. 24 would, after 300 years, be found high and dry some 150 feet up the face of the cliffs. On the coast of South America there is good Fic. 24.—Imaginary view of Spanish sailors carving an inscription on rocks at sea-level in 1600 a.p., on the Chilian coast. reason for believing that a movement upwards goes on at the rate of half a foot to a foot a year. If such a rising continued for a thousand years we should find that the original shore-line had risen 500 feet above the sea-level. 39 EXTINCT ANIMALS What, then, is the general result of such move- ment? I will show you what would be the result of elevating the shore of England (the whole of this part of the world) 600 feet. From this map (Fig. 26) you will see that if the Fic. 25.—The same rocks as they would appear in 1900, raised 150 feet above the sea-level by an imperceptible movement of six inches a year. floor of the ocean were raised 600 feet, the cross-shaded area would become dry land, and we should be brought by land into contact with the neighbouring continent and islands. And if the land were raised 3,000 feet we should have a still greater extension of dry 40 ny Jerbishire & Slantfora, lo ine Usford Geog Institut Fic. 26.—Map to show the effect of elevation of the earth’s surface on the distribution of land and water in Western Europe. The dowbly-cross-shaded area shows what would become dry land if the sea-bottom were raised 600 feet. The Channel, the German Ocean, the Baltic and the Irish Sea, cease to exist. The smaller dotted area would become dry land if the sea-bottom rose another 2,400 feet. Men could then walk from Scotland to Iceland by way of the Shetlands and Faroe Islands. Most noticeable is the great change which would be brought about by the comparative- ly small rise of 600 feet, and the much greater elevation required to change any further the contour of the land. 41 EXTINCT ANIMALS land. Even the smaller change would make Eng- land part of the Continent of Europe. The study of extinct animals found in the various strata of the earth enables one to arrive at a notion of the distribution of land and water in past time. Here is an arrange- ment of land and water which we are able to conclude must have existed in Europe in what is called the Middle Tertiary period (Fig. 27). All this darker part is the sea, and the pale part land: in fact, the distribution is quite different from what it is at the present time. The whole surface of the earth has been shifting and changing all through time. During the millions and millions of years of past ages, different seas have arisen, different continents, different dry land and different animals,—changed by the various influences of the land and climate. And all this movement is accomplished by the slow cracking and ‘‘ curl- ing” of the earth’s crust, by the continual wash- ing of the surface of the land by rain and rivers, by the eating away of the edge of the land by the waves of the sea. This “eating away” of the land by the sea—quite apart from any sinking of the land-level—has caused and is yearly 42 ANCIENT LANDS AND SEAS causing great loss of land on the east coast of England, especially in Suffolk, where the great city of Dunwich has been swallowed up by the sea. In other parts the sea throws up sand and iN ¢ ; AU Al q Fic. 27.—The real test of Geology: anattempt to determine the distribution of land and water in past ages. The period here shown is the Oligocene or Middle Tertiary, the area is that of our present Europe. The sea is shaded, the land areas are left white. Central and Southern Europe was a sea, with a few large isiands in it. North Europe formed a continent including the British Islands and Iceland. (After Lapparent.) adds miles of new land to the coast. The immense quantity of stuff which is carried off the surface by rains and rivers is difficult at first to imagine. Taking the river Thames at Kingston, it is found that something like 500,000 43 EXTINCT ANIMALS tons of solid salts of lime in solution is carried every year past that spot. Now a cubic block of limestone measuring a yard in each dimension weighs about two tons. Accord- ingly, 250 thousand solid cubic yards of rock are carried past Kingston every year by this little river! Enough to build a new St. Paul’s Cathedral every year! Think, then, what must be the enormous quantities of solid matter dissolved and carried away by such rivers as the Mississippi and the Amazon. And remember that in addition to this dissolved lime- stone there is almost as large a bulk of fine sand and mud carried along by most rivers! What becomes of it? It is deposited in layers, and forms what we call stratified rock. You see it, some of it, on the seashore when the tide goes back, in the form of layers of sand, but most of it is deposited far out in the deep bottom of the sea—the lime being taken out of solution by shell-making plants and animals. But where the land is rising, the sand or ground which is ex- posed when the tide goes back, would after afew years have been raised away from the sea and become hard rock. Layer after layer is imposed. and raised from the sea bottom. Without 44 STRATIFIED DEPOSITS going into detail we may accept as a fact that this formation of layers by stuff brought down from the land by rivers and washed from the coast-line by the sea waves gives rise to what are called “ stratified de- posits.” I will now show you some pictures of this stratification. Here (Fig. 28) are shells a. — ra <= i = 5 - ~- = = y Fic. 28.—Photograph of a slab of Bognor rock (Lower Eocene) showing embedded marine shells. embedded in the Bognor rocks deposited some thousand million years ago; there are many sorts of shells, whelk-like shells and _ volute shells embedded here. Whole skeletons of animals are sometimes 45 EXTINCT ANIMALS found in the stratified deposits. This one (Fig. 29) is from stratified rock which forms the hills round Paris, the calcareous rocks of Mont- martre. Next let us see what fine mud will do in Fic. 29.—Skeleton of a tapir-like animal (Paleotherium) as found embedded in calcareous rock at Montmartre, Paris. preserving the impression of delicate structures, such as the wings of insects. Here are the wings of a dragon fly (Fig. 30), preserved in very ancient stratified rock, the Carboniferous. Here (Fig. 31) are the wings of the Pterodactyle 46 FOSSILIZED WINGS Fre. 30.—Wings of a Dragon-fly preserved in the ancient lime stone of the Carboniferous period or Coal-bearing rocks. preserved in fine sandy limestone of Oolitic age. Here (Fig. 32) we have a jelly-fish preserved ; Fic. 31.—Pterodactyle skeleton preserved in Lithographic limestone, showing the impression of the membrane of the wings. 47 EXTINCT ANIMALS you see its seal stamped as it were on the sand. It is many millions of years old, from Oolitic rock. Now let us look at the layers or the strati- fication of rocks. This picture (Fig. 33) shows part of the cliff at Lyme Regis, where the Ichthyosaurus-head, which I showed you just Fic. 32.—A Jelly-fish (similar to the recent Aurelia aurita) preserved in Lithographic limestone. now, was found. We see the layers of harder and softer material lying one over the other. The next figure (Fig. 34) shows how the layers of the surface of the earth may be bent. With- out digging far into the earth you may reach a deep layer of stratification or “ stratum ” brought near the surface by the general tilting. 48 TILTING OF STRATA This (Fig. 34) is part of the chalk cliff at Sea- ford, showing the strata tilted, so that the deeper layers come to the surface. Here (Fig. 35) is part of the shore of Lyme Regis, showing the strata exposed by the action Fig. 33.—Alternate layers of hard and soft rock (‘‘ strata ’’) forming the sea-cliff at Lyme Regis. Photographed by Messrs. Dollman Bros. of the sea. A long series of superimposed layers one on top of the otheris seen. They are slightly tilted, so that the deeper strata come to the surface near the observer. The tilting of the strata of the earth’s crust 49 E EXTINCT ANIMALS is the rule and not the exception. It is rare for the strata to lie in a strictly horizontal position. The crust of the earth is continually being slowly pushed up or down, and as it were ‘“crumpled”’ or thrown into wave-like folds. The cause of this crumpling is to be found in graphed by Messrs. Dollman Bros. the shrinking of the earth and the movements of subterranean steam—causing earthquakes and other earth movements. The “crust” of the earth is a mere skin. If we bored twenty miles into it we should come to immensely hot 50 FOLDING AND CRUMPLING OF STRATA molten material, and on this the crust is sup- ported. It cannot be said to “rest’’ on the deeper matter, for it is always, though very, very slowly, shifting and crumpling. Con- sequently, according to the height and depth of Fic. 35.—Strata of the ele at Lyme eee! Photographed by Messrs. Dollman Bros. the folds of the crust, we find that deeper, even very deep-lying strata may be brought to the surface, and as the upraised folds get worn away by sea and rain and rivers, the deepest layers may be exposed on the surface. Thus it is that we are able to examine the oldest rocks - 51 ANIMALS EXTINCT s of the immensely- and to search for the remain ‘saoXeB] eyUOZLIOY VOU 9Y}Z JO UOTZB_NpuN 40 Sutpueq ayy Aq dovyjans ayy 0} FYsnoaq ‘sdeAv_ Jodoep UsAS puB *G PUB g BPBIIS Puy Ayqeqoad plmoys aM ‘uredserp oy} Jo Fruty ety puodoeg our, cures oy} Suole uorye10[dxe Ino ponuryUos OM F[ “MOTA 0} 1 ‘ON UWoA Sutsodxe sjuoutdagose ,, 10 ,, SYTO ,, JO UOTy -eulof ayy Aq puv oovjans oY} UO g ‘ON SB Coop SB ByRA4S Sutsodxe snyy ‘ABMB poysBar usog SBY YOM BzBVQs poavoydn oyZ JO UOIZBNUTZUOD JOULLOF OY} YxeUL Sout, Poop e"uUL °6 ‘8 ‘L ‘9 ‘GS ‘F ‘ES ‘S% ‘IT SpresuMOp 9A0GB WOdy poyIwut o1B BIBIJS BATSSID -ons oy, ‘eoBjans s.yjave oy} UO AoT[BA B JO W0}}0G AO [OAV] OY} SYIVUr (7) eullL ¥ “qsndo S$ ‘ ‘ SSeS SS NN Y4rBvo 94 jo uorzyByNpun LO sutpusq oy} Jo Joo}yo oy} MOYS O04 uIBIsel([— ‘9g “OI The In. t creatures which they conta anclen N ite) RIPPLE-MARKS AND RAIN-DROPS diagram (Fig. 36) will help to make it clear how the pushing and crushing of the earth’s crust into wave-like folds such as you may see when a tablecloth or carpet is not spread flat, results in bringing the deep-lying strata to the surface, so that we can walk along a cutting or cliff and come to deeper and older rocks as we walk along. Here (Fig. 37) is a specimen which shows Fic. 37.—Ripple-marks preserved in ancient Triassic strata. ripple-marks still preserved as we see them nowadays on the shore at low tide. The marks of raindrops are also often preserved on such slabs of rock, which once were soft wet sand. On such surfaces we often find footprints, the footprints of birds and of reptiles. In some cases we do not know the animal itself (Fig. 38), 33 EXTINCT ANIMALS but we see its footprints in the ancient rocks now far removed from the sea and covered over by thousands of feet of later rocks. Here (Fig. 39) are the footprints of a great creature Fig. 38.—Bird-like footprints on a slab of Triassic rock from Connecticut, U.S.A. from the Isle of Wight which has left its impress in the sandstone. In Fig. 40 we have drawn a slab of Triassic Fig. 39.—Three-toed footprint (probably of Iguanodon) from the Wealden Sandstone of the Isle of Wight. 54 FOOTPRINTS ON ANCIENT SANDS rock, showing the five-fingered hand-like foot- prints of the Cheirotherium (as it was once called), a huge salamander-like animal. Fie. 40.—Slab of Triassic rock from Saxony, showing hand- like five-fingered footprints, each seven inches long, probably due to a Labyrinthodon which walked over this substance when it was soft wet sand. These footprints occur also in rocks of the same age in Cheshire. TABULAR LIST OF THE GREAT GROUPS OR BRANCHES OF THE PEDIGREE OF LT: IIT. XI, ANIMALS VERTEBRATA. (Back-boned Animals.) Class 1. MAMMALS. BIRDS. REPTILES. AMPHIBIANS. . FISHES. 6, 7 and 8. Lancelets, Ascidians and Acorn-Worms. MOLLUSCS. (Mussels, Oysters, Clams, Snails, Slugs, Whelks and Cuttle-fish.) APPENDICULATES. (Insects, Crabs, Shrimps, Spiders, Scorpions, Centipedes, and Annulate Worms and Wheel-animalcules. ) ECHINODERMS. (NStarfishes, Sea Urchins and Sea-Cucumbers. ) FLAT WORMS. (Flukes, Tape-Worms, Water- flukes, etc.) NEMERTINES. (Cord-like Sea-Worms.) NEMATODS. (Parasitic Thread-Worms. ) ot ge to CORAL-POLYPS and SEA-ANEMONES. , HYDRA-POLYPS and JELLY FISH. SPONGES. PROTOZOA. (Microscopic Unicellular Animalcules, Amcebex, Gregarines, Flagellates, etc.) N.B.—The list does not contain the less important great groups, and is purposely made more simple than are the tables of classification used in scientific text-books. 56 TABULAR LIST OF THE CHIEF ORDERS OF THE VERTEBRATE CLASS Order tm GC bo — 14 15 16 MAMMALS. Primates Man, Apes and Monkeys. Insectivors Hedgehogs, Shrews and Moles. Chiroptera Bats. Carnivors Dogs, Bears, Cats and the extinct Creodonts. Pinnipedes Seals. Ungulates Hoofed Animals: even-toed and odd-toed. Elephants Elephants and their extinct ances- tors. Amblypods Dinoceras and Arsinditherium. Toxodonts Toxodon. Rodents Rats, Rabbits, Beavers and Porcu- pines. Hyracoids The Syrian and African ‘‘ Coney.” Sirenians The Manatee, Dugong and Steller’s Sea-Cow. Edentates Sloths, Armadilloes, Ant-eaters. Cetaceans Whales and Porpoises. Marsupials Kangaroos, Opossums, Tasmanian Wolf. Cloacals The Egg-laying Platypus and Echidna of Australia. N.B.—The list is not complete 57 TABULAR LIST OF THE CHIEF ORDERS OF THE VERTEBRATE CLASS Order 1 REPTILES. Dinosaurs Huge extinct creatures, often as big as elephants. Crocodiles Crocodiles and Alligators. Chelonians Turtles and Tortoises. Lizards \ar Closely allied to one another, and - having the epiderm moulded Snakes | to form ‘ scales.” Pterodacty!ls Extinct: they had great flying wings supported by one finger. Theromorphs Extinct: often with teeth resem- bling those of Mammals. Plesiosaurs Extinct: swan-necked aquatic forms, with four paddles. Ichthyosaurs Extinct: short-necked marine forms, closely representing among Reptiles, the Whales and Porpoises of the Mamma- lian series. N.B.—This does not profess to be a complete enumeration. 58 CHAPTER II STRATA AND LAND SURFACES— TEETH AND BONES — EXTINCT MEN — FLINT IMPLE- MENTS—THE MAMMOTH, ELEPHANTS AND MASTODON—CLASSIFICATION OF ANIMALS EFORE giving you further accounts of extinct animals, I wish to point out to you that what I have to say is true, and not mere imagination. Some people talk about the “fairy tales of science.”’ There never was a more inappropriate phrase: it is altogether wrong to speak of fairy tales having anything to do with science. The wonderful things which science reveals to us are altogether remote from fairy tales, for in regard to the tales of science you can test what you are told, you can see the things of which I speak, you can ascertain the truth of what is asserted. That is the great pleasure of this study; one knows that the things one examines, however astounding and 59 TABLE OF SFRATIFIED ROCKS SHOWING APPROXIMATE THICKNESS. TOTAL DEPTHS. QUATERNARY YO .EMS TYPICAL a DOr RECENT SSS ee MAN, MAMMOTH, MASTODON 450°" PLEISTOCENE 200 ft == SS THREE TOED HORSE 1430. — ; =50 GS TETRABELODO 2850 - purosene all EOCENZ SOO feet _—*&=_ARSINOITHERIUM DINOCERAS ———— = ANCESTORS OF ELEPHANTS : > ——~---CRETACEOUS —~ HORSES & MAMMALS GENERALLY Be a Fe em 53506 nae ~ ~— 2.800 ft ~— LAST ICHTHYOSAURS zo 2G 0) 1a) 0) oO) Ol Ol OF.oO) Oo) (oslo bu GREAT DINOSAURS SSS a3 ey | OME OO N Oe ORNS Har re ZNOIO 1050 .0 (0) (0) 8 010: © Ove |e we z$ BuO. ors Oot lomomegoe Er 1aa5o ae CH oO! Towoolble ©o: OsoN_ol OLOLelig ' a= a « n THEROMORPHS ‘ace < PARIASAURUS 14.850 2 = LAST TRILOSITES = < i < w « o SCORPIONS Ex 26850 - ae a TTT / x DEVONIA ny x 5,000 ft /) = wi 31,850 ~ Gu = FIRST FISHES = MARINE SCORPION re) z < 38,850 « = < e a =) « =) o oO o 20 3 = a ae) = 1) Ww F < 53.850 " : =] & BOGS SE0E2S S08 S82 Sea SS0 000808 | i GEREBERERERERBS Soe is 1 || ipo SOD Seeee c GS SSP SERRE SS0RR w Pa SEBSERSEEGERCRESEEEE > — GUBESEEEEOEESa - z < rrtTiTTrTiCAMBRIAN[TTTT ITT] eB ean ERHEERSROR Ft GERREEER S PTT Ty yy i2,000 BRRSRRER HGEBRs008 GEESHEaR BEBESERLaaS =! t HOGS FIRST TRILOBITES 65850 Fig. 41.—A tabular view of the strata of the earth’s crust, showing the relative thickness of each “‘ system ” or group of strata, and the position in which impor- tant animal re- m5659 mains have been discovered. The rocks are never found lying hori- zontally like this, which is a diagram. In nature they are tilted and ecrum- pled, but we can make out their thickness and the order in which they lie one over the other. THE SUCCESSION OF STRATIFIED ROCKS incredible they seem, really exist, and are not mere imagination or fancy. I want now to refer to these large diagrams (Figs. 41 and 42). Fig. 41 should be carefully examined. It represents what has been dis- covered with regard to the succession of de- posits, those stratified deposits of which I spoke in my last lecture. On the left-hand side is stated the thickness of each deposit, so far as it has been ascertained. Most of the extinct animals, all the great extinct animals I have to speak about, come within the upper part. We have an enormous thickness of stratified rock beneath, which contains only marine things, fishes, a few crustaceans, and things of that kind. But all the more interesting great animals have left their bones in the higher strata. The upper- most layer (the recent and Pleistocene) is only some 200 feet in thickness, yet it indicates a period of something like 500,000 years. This being so, you can judge by the thickness of subjacent deposits what an immense lapse of time is represented. Before we get to the chalk we get down nearly 3,000 feet. The thickness of the chalk itself is another 2,500 feet. 61 EXTINCT ANIMALS The estimate thus given probably does not fully represent the time which has elapsed. If you take a thousand years for each foot, you only get an approximate measure of the time represented, because a great deal more time has passed than is actually shown by the permanent deposits or strata. Strata have been broken up by the sea and water, and have been deposited again and again; and it is probable that a much longer time has elapsed than one thousand years for each foot of the deposits which form the stratified crust of the earth. An important general fact, which I cannot dwell on further, is that whilst it is true that the great animals occur in the later stage of the world’s history, there is a gradual succession from simpler to more complex forms of life. We get fishes at the top of the Silurian; and we get in the Carboniferous great amphibians ; and the first reptiles in the Permian; and then we get birds and crocodiles in the Triassic ; and the first hairy warm-blooded quadrupeds in the Jurassic. Thus the different kinds of animals succeed one another in the order of increasing complexity of structure so that the highest animals are the latest to appear. 62 THE ZOO-GEOGRAPHICAL MAP This map of the world (Fig. 42) has special ‘[eUoAGQ oy (Ff) pue ‘ustdoryyT aya (g) ‘peotdoay-ooN oY} (Z) ‘ormoaepoFyT oy} ({) *ZIA ‘suotsoa aMmofF oY} OFUL PoOplATp SI Bliay, UMITeQyUeDR[q oy} AYseyT “Bildoy, WMITeJUooR[_ oY} PUB BITBASNY 0JUL paprArp si: Bvesowoyy, Usyy, “Besos, (q) pue purleoz MON (”) OJUT ST UOISIAIP JSay oy, ‘“speuuTUe JO spuTy qUetoyTp yo oouesead oy Aq postiojoRreyo suoIsor pue sooutAodd yvoad OFUT UOISTAIP s}yt Surmoys ‘ppxoA4 OY} JO Aey\—'sF 199n7/4SU/ Boa) pyoyxO PY] “OL "PLT (PIOZ LETS P As/YSIQJEQ ce” é A See BS : fee s§ x yet ex » ' x SL. gee 2 . = : oe Surah pice : e+ ty y AGE Ae ae interest, for it shows the present position of It is meant to show different kinds of animals. 63 EXTINCT ANIMALS something of the history of the migrations or movements over the surface of the earth of the large animals which have lived upon it. The line which separates New Zealand from the rest of the world, which we call Theriogzea—the land of big animals—shows that the large islands of New Zealand have no such animals upon them. Till man went there some thousand years ago there were no large animals. The largest animals were great birds. There were no cattle, or cats and dogs—not even mice. Thus this piece of land seems to be separated from the history of the movement of animals in the rest of the world. It is an old and detached land-surface. Then you will see a second line between Australia and the rest of the world. Australia is distinguished by its marsupials (kangaroos, wombats, phalangers, etc.). The young of the marsupials are very small when born and are placed by their mother in a pouch of skin overlying her teats. Those animals which are nourished inside the mother before they are born are of a much larger size at birth. They are the Placentals, and there are no aboriginal Placentals in Australia. The greater part of the world (the rest when 64 THE ZOOLOGICAL PROVINCES Australia is cut off) may be divided into the great Holarctic surface, the northern strip which comprises North America, Europe and the Northern part of Asia, while projecting downwards are three other regions, South America or the Neo-tropical, the great African or Ethiopian region, and the Oriental or Indian region. The animals of which fossil remains are found in the Holarctic region have migrated into these projections of the land which sub- sequently became of their present shape, at different times in the world’s history. The Neo-tropical region of South America. was at one time a separate mass of land, and upon it lived very peculiar animals, such as the great sloths and armadilloes, and strange birds. In the Holarctic region we find, either still living or buried in recent strata, the great hairy mammals, elephants and cattle, antelopes, deer, camels, horses, rhinoceroses, tapirs, pigs, hippopotami, tigers and lions, and such forms. When we dig down only to the depth of a few feet, in river gravels and comparatively modern deposits, we find all the big creatures in this region as shown by their fossilized bones. But, owing to some change of climate and other con- 65 F EXTINCT ANIMALS ditions not very clear, most of them left this region and migrated to the southern projecting regions. One of the most curious results of this emigra- tion is that at the present day the tapir is found alive in the island of Sumatra and that it is found alive also in Central America. At one time naturalists were much surprised to find a tapir in the new world like the tapir in the old world, and nowhere else but in these limited spots, remote from each other. But now we know that tapirs existed all over the Holarctic region, for we find there their fossil remains ; we recognize them by the shape of their teeth and bones which we dig up. Even in England, in Suffolk, we find the tapir in the deposit known as the Red Crag, and again in different parts of Germany, France and Greece, and even in China and in North America, tapirs are found buried in the sands of Pliocene and Miocene age. The present race of tapirs existing in the East Indies and in Central America are as it were the outlying survivors of those which existed formerly all over the great Holarctic region. Such facts as these about the tapir indicate the importance of knowing where particular 66 HOW TO RECOGNIZE BONES fossil animals are found; for thus we are enabled to come to some conclusion as to the former connexion of different land surfaces of the world with one another. The question must have occurred to many of you,—How do we recognize fragments of bones found in the earth? How do I know that a fragment I may find is the lower jaw of a creature like the horse? or that bones I may dig up are the bones of a tapir? How do I know that a given skull is that of areptile ? and that a given shell was inhabited by a creature like the nautilus ? We are able to know these and like matters because the shape of different parts of each kind of animal is very constant. The kinds which are like one another in other respects are like one another in the details of their bones and teeth, even in such minute points as the microscopic texture of the bones. An immense mass of facts about such things is known, and when set out in orderly fashion is termed the science of comparative anatomy or animal morphography. The first photograph I have to show in this chapter is of a piece of bone which was sent 67 EXTINCT ANIMALS fifty years ago to Professor Owen by a gentle- man in New Zealand who had lately arrived there, and who had found it in his garden. Professor Owen, on examination, was able to say from the general make and structure of the bone that it was the bone of a bird. It was about seven or eight inches long (Fig. 43). On examining the ridges and various Fic. 43.—Photograph of the original piece (seven inches long) of a thigh bone of a gigantic bird, from the examination of which Sir Richard Owen inferred the former existence of a gigantic flightless bird in New Zealand. The specimen is preserved in the Natural History Museum. (Original.) marks on the bones, Owen was able, from his knowledge of the character of bones, to say that it was identical with the middle part—the ends were broken off—of the thigh bone of an 68 SS : Fic. 44.—Photograph of Sir Richard Owen standing beside the restored skeleton of the New Zealand Moa (Dinornis maximus). From a memoir by Owen. 69 EXTINCT ANIMALS ostrich. He ventured then to publish that this bone was a proof that there existed formerly in | New Zealand a huge terrestrial bird like the os- trich, only bigger. After a few years, more bones were sent to Owen from New Zealand, which entirely confirmed what he had said: and in the course of a few years he was able to put to- gether from the bones sent a skeleton with enormous legs and neck, the skeleton of the ostrich-like bird the Moa of New Zealand. In Fig. 44 you see Professor Owen himself at the side of the restored skeleton. Since that time a great number of these birds have been found buried in the morasses and comparatively recent deposits of New Zealand, showing that many of them existed alive some five or six hundred years ago, and that they were then probably hunted out of existence by the an- cestors of the present Maoris. I shall have a few more words to say about the giant birds of New Zealand in a later chapter. In Fig. 45 we have the photograph of a very fine preparation in the Natural History Museum, showing the skeleton of a man and a horse side by side. The main object of this com- parison is to show that, though so different in 70 MAN AND HORSE Fic. 45.—Photograph of the skeletons of Man and Horse from a group, pre pared under the direction of Sir William Flower for the Natural History Museum. Sh. Shoulder-bone. W. Wrist-bones (so-called knee of horse’s fore- leg). E. Elbow process (olecranon). IK. Knee joint (Stifle of horse). P. Hip-bones. TZ. Tail-bones. H. Heel-bone (caleaneum of man), the hock of the horse. general bearing and form, all the bones of a man correspond in detail with those of the horse. The thigh bone of the horse and the thigh bone of the man, the knee (called the “stifle’’) of the 71 EXTINCT ANIMALS horse and the knee of the man, correspond. The man has a short foot, the horse a long one. The upstanding bit at the back of the horse’s leg called the “ hock” is really the heel, and cor- responds to the heel bone which you can dis- tinguish in the man’s skeleton. So also the fore-arm and shoulder-blade correspond in the two skeletons. Accordingly, as animals are alike or unlike in the details of their structure, so we can group them into divisions and sub-divisions (see the list of classes at the end of ChapterI). There are certain marks by which it is easy to recognize fragments of bone, dug it may be out of a quarry or railway cutting, and to know at once the division or kind of animals to which the owner of the fragments belonged. I have already alluded to the fact that the strata of the earth are revealed to us by cliffs on the sea-shore, by exposed rocks and. by river banks; and I would add by such activities of man as the digging of quarries and railway cuttings. Suppose that you find a skull in such a digging—there are marks by which you can tell whether it belongs to a mammal or reptile. THE OCCIPITAL CONDYLES In Fig. 46 I have photographed the whole back part of a skull which contained the brain, and you see where the spinal cord entered the skull to join the brain. In this creature (an ox) there are two bony surfaces (marked Ex, Ex) £03. aa 99.3. 124. eee ee ee ee ee - = wen 0 oie eater 9 Fic. 46.—Photograph of the back of a skull of an Ox, to show the two occipital condyles, Hx, Ex. forming the joints or condyles of the skull by which the first neck-bone or vertebra was fastened to it whilst allowing a rotating move- ment. All mammals’ skulls are provided with this pair of knobs or “condyles.” But in the crocodile’s skull (Fig. 47) you will see below =9 /o EXTINCT ANIMALS the aperture for the spinal cord only one large condyle (marked Bas). From such a fragment of the skull then you can at once tell whether to place the creature to which it belonged among the hairy warm-blooded quadrupeds called mammals, or with the reptiles. A bird Fic. 47.—Photograph of the back of a Crocodile’s skull to show the single occipital condyle Bas, lying below the hole or foramen by which the spinal cord enters the skull to join the brain. is like a reptile in having a single joint or knob at the back of the skull. As an example of the definite marks by which bones can be referred to their proper classes, the following is a curious point. Fig. 48 is a drawing of the internal ear of man—the soft 74 THE SNAIL-LIKE COCHLEA part of the ear bedded in bone. three loop-like canals and a snail-like coil. mals have that snail-like con- struction of the internal ear. It consists of All hairy mam- inside, em- In Fig. 49 is photographed the pre. 48 —Drawing of ear-bone of a mammal’s skull cut through, and you can see the place for the snail-like soft ear—the cochlea or spiral of the ear, as it is called. No other animals except the Fic. 49.— Photograph from a section through the bone in which the soft inter- nal ear is_ lodged, showing the coils of the snail-shaped space in which the spiral cochlea lies. the auditory organ or internal ear of man. A the coiled tube known as the helix or cochlea. B the three tubu- lar arches or semi- circular canals. internal mammals are known to possess a_ spiral internal ear, and all known If, therefore, you discovered a mammals do possess it. fragment of bone showing this spiral-like space you would know that the bit of bone must in all probability belong to a mammal. At the beginning of the nineteenth century a portion skull was brought from America 75 of a great elongated EXTINCT ANIMALS to Europe, dug out of the sands of Florida. It was thought to belong to a reptile like the crocodile, and was called Basilosaurus. But the naturalist in whose care it was, on showing the specimen to a friend (Herman von Meyer) dropped it on the stone floor of his museum and cracked the back of the skull. The crack exposed the spiral cavity or cochlea of the ear, and thus it was shown that the specimen was the skullof a mammal. Sure enough, it turned out later to be the skull of a kind of whale (Zeuglodon). Teeth are of great help and importance in determining the sort of animal to which a fragment belongs. Fig. 50 is a photograph from a specimen prepared in the Natural History Museum. The wild boar or pig occupies in regard to teeth a sort of central position among mammals (hairy warm-blooded quadrupeds). Its teeth are such that to them you can refer, as to a standard pattern, the teeth of all other mammals. There are three middle teeth in front in the upper and lower jaw, chisel-like teeth, the incisors. Be- yond these are the great canine teeth: then the cheek teeth follow. These are seven in 76 THEETH- OF THE PIG Fria. 50.—Photograph from preparations of the upper and lower jaw of a Pig, to show the teeth in position. The bone has been cut away so as to show the roots or fangs of the teeth. (1, 2, 3) the three upper incisor teeth of the left side ; (4) the upper canine tooth of the left side ; (5, 6, 7, 8) the four front molars or cheek teeth, called the premolars, of the left side of the upper jaw ; (9, 10, 11) the three back molars (not preceded by “ first’? teeth) of the left side of the upper jaw; (12, 13, 14) the three lower incisor teeth of the left side; (15) the canine of the lower jaw, left side: note its enormous root; (16, 17, 18, 19) the four front molars (premolars) of the left side of the lower jaw ; (20, 21, 22) the three back molars (not preceded by “first ” teeth) of the left side of the lower jaw. HLF EXTINCT ANIMALS number, four in front which are replaced —that means that second teeth come to take the place of the first—and three hinder ones, which are never replaced. If you look at the surface of these cheek teeth you will find they are broad, with many tubercles, fitted for grind- ing great varieties of food. There are seven of these cheek teeth on each side in each jaw, upper and lower, one canine, and _ three incisors, so that eleven on each side in upper and in lower jaw or forty-four teeth in all is the complete number, the typical number—the most characteristic number in the group of hairy mammals. Many have less, but among the immediate ancestors of those mammals with ‘reduced dentition ’’ we find a larger number of teeth, and in their remote ancestors the com- plete typical number is discovered. It is important to notice that whereas the front teeth have a single fang by which they are implanted in the jaw the cheek teeth have two fangs, as shown in Fig. 51. Teeth with two fangs appear to be peculiar to mammals. Other animals have only single farigs to all their teeth, as mammals have for their incisors and canines (as a rule). TEETH The human teeth (Fig. 52) are reduced in number. There are only two incisors above and below on each side; then the small canine Ftc. 51.—Photograph of a preparation of the teeth of the upper and lower jaw of a Pig. The small teeth between the upper and lower row of large teeth are the milk teeth or ‘first’? teeth which are shed. Note how small the predecessors (15 and 23) of the great tusks are, and also that the foremost molar (7 and 33) in both upper and lower jaw has no successor or predecessor, as is also true of the three back molars. or dog-teeth, one on each side ; then five cheek teeth or ‘“‘ molars,” two smaller and three bigger. From a single tooth we could tell whether a piece of jaw-bone belonged to a man 79 EXTINCT ANIMALS or not. Though like a monkey’s, a man’s tooth can be distinguished from it and from all other Fic. 52.—Photograph of a preparation (in the Natural History Museum) of the upper and lower jaw-bone of man, the bone cut away so as to show the fangs of the teeth. The pattern of the crowns of the molars is well seen in the upper and lower figures. teeth. In the next figures we have photographs showing certain modifications in the teeth of 80 TEETH OF TIGERS AND OF RATS mammals. You see in the Clouded Tiger (Fig. 53) that the teeth are few in number, and are sharp, for cutting or tearing flesh, whilst the canine teeth are very large. In Fig. 54 the skull of a great rat, as big asa beaver or fair-sized dog, is photographed. The Fic. 53.—Skull of the Clouded Tiger, to show the large canine teeth and the few but pointed and cutting molars, two above and three below. front teeth (only one on each side above and below) are chisel-like, and very large, to enable the rat to gnaw wood. In reptiles you no longer get complex cheek teeth. All the teeth are peg-like. They have SI G EXTINCT ANIMALS no grinding teeth with big surfaces, and all the teeth have a single fang (Figs. 55 and 56). Fic. 54.—Photograph of the skull of the Coypu Rat, to show the greatly enlarged incisor teeth or “ rodent ”’ chisel-like teeth in front, the absence of canines, and the flat grinding molars behind. The large gap in the row of teeth between the incisors and the molars is very characteristic. The fossil jaw shown in Fig. 57 came from Fic. 55.—Jaws of the Gharial, an Indian Crocodile, to show the peg-like teeth. The bone is removed, showing that the teeth have only a single fang each. Stonesfield in Oxfordshire. It is embedded in 82 A FOSSIL JAW FROM STONESFIELD hard Jurassic slate, and is one of the most ancient evidences of the existence of a mammal. The sight of its double fangs at once rendered it almost certain that the teeth must be those of a mammal: the whole shape of the jaw is Fic. 56.—Photograph of the skull and lower jaw of a true Crocodile. The numerous peg-like teeth of different sizes, firmly implanted in the jaw-bones, are shown. like that of a small mammal, such as the hedge- hog. We must now take up again the general story of extinct animals; and to do so we will first of all go back, so to speak, a little way into the strata deposited on the earth’s surface—just far 83 ‘yySueay ur yout ouo Ajuo Aye -nyow st Met oyy, ‘ouoq-ael oyy Jo Ave Suryeorq oyy Aq posodxa YY204 Yyooyo oy} Jo [eteAes JO SSuvy o[qnop oy} SMOYS oMSY OY, “pOJXO aBvou OSB (OJI[OD) oIssBIne Jo a4yR[S p[eyseuoyg oy woul (~soaaid Uniwaynydup) yeutureut [peuis B jo Mel JaMol oYy JO UOIVkZUeSeIdoa posaepuyf—"LG “O91 YT PREHISTORIC REMAINS OF MAN enough to take us beyond the range of written history or record, which barely reaches further than four thousand years. Recent explorations in ancient cities, Egypt and other parts of the East have brought out from layer after layer of rubbish and mud the different remains of man, different instru- ments, utensils and works of art. As one gets deeper one finds remains showing different habits and ways of life. But all are practically within the historic period. Beyond that we come to a period of which there is no tradition or written record, but of which we have evi- dence only by the remains we find; flint instruments, carvings, and even occasionally some human bones. The most important of the prehistoric remains of man take us back, to judge from the position in which they are found, some 150,000 years. These are the remains, found in river gravels in England and France and other countries, proving that man lived here in a savage state with the Mammoth, the Rhinoceros, Hyena, Cave Lion and Cave Bear. Fig. 58 shows two flint implements which these men manufactured and used. A flint instru- 85 EXTINCT ANIMALS ment of this kind was found more than a hundred years ago in the gravel in Gray’s Inn Lane, in Fic. 58.—Photographs of two flint implements of the Paleo- lithic age, obtained from the gravel-pit at St. Acheuil near Amiens, by the author, in 1870. London, and was figured and described ; but its ereat antiquity was not recognized at that time. In the middle of the last century, attention 86 PREHISTORIC MAN was drawn to these flint instruments found in the gravel of the river Somme by a French antiquarian, M. Boucher de Perthes. He got immense quantities of these worked flints from the neighbourhood of Abbeville and Amiens, and he maintained they were the work of men. They were clearly, from the depth of gravel under which they were found, of enormous antiquity. The matter was gone into carefully at the time; geologists and naturalists took keen interest in it, and the great antiquity of man in Europe was established. And besides these implements in the gravel others have been found in caves associated, as in the gravel, with the remains of animals which*have long ceased to exist in this part of the world. 4 These are such mammals as the reindeer, the hairy rhinoceros, the great Irish stag, the cave bear, the cave hyena and the lion. Huge wild cattle, such as the Aurochs or Urus of Caesar, and the Bison, existed then in quantity. In some places the actual bones and skulls of these primitive men have been found with the bones of extinct animals. The skulls of primitive men and of modern men show a certain difference in shape. If we o6) 7 EXTINCT ANIMALS take two skulls, that of a man and a monkey (Fig. 59), and draw a line from the region just over the nose, between the ridges of the brow, and run it back to the occipital ridge at the back of the skull, there is left above the line a Fic. 59.—Photograph of the top of the skull or “ calvaria ” of the so-called Monkey-man, Pithecanthropus, discovered in Java. On the left is the skull of a Chimpanzee and on the right that of a modern man, for comparison. A line is drawn from the point between the eyebrows to the occipital ridge at the back of the skull, showing how much shallower the dome of the skull (the part above the line) is in the ape than in the man, and that the Javanese skull is nearly as shallow as that of the ape. (Original.) great hemispherical dome in the human skull, whereas in the monkey the space left above is much flatter, much shallower. In ariver gravel in Java the imperfect skull of the so-called Pithecanthropus, or monkey- 88 HUMAN SKULLS man, was lately discovered. It is really, in its main features, a human skull. A_ photo- graph of it is seen in Fig. 59—the middle one Fie. 59a.—Photograph of a human skull of modern Juropean race. of the three figures. It has a shallow upper region, much like that of a monkey. Other shallow skulls of primitive men have been found in caverns, such as those of Spy in Belgium and 89 EXTINCT ANIMALS in the sand of the Neanderthal on the Rhine. It seems certain that primitive man had a shal- lower brain than the more recent man. But some of the prehistoric men seem to have been able to draw, and to have exhibited great skill in that art. It is difficult to say whether there was more than one race present at this time, and whether the men of the shallow skulls were the same men who made the drawings. In one of the caves of France inhabited by prehistoric men, and thickly strewn with their chipped flints and with the bones of extinct animals eaten by the men, a piece of a mammoth’s tusk has been found with a mammoth carved upon it (Fig. 60) evidently by the men who lived there. We also find the heads of reindeer, carved upon pieces of bone. The photographs reproduced in Figs. 60 and 61 are from drawings of the actual specimens. In Fig. 61 is shown a piece of an antler upon which a reindeer is cleverly outlined. The tuft of hair below the chin is shown, and the great feet and the extra toes are correctly pictured. Clearly the men who drew this reindeer lived with the reindeer. And besides the reindeer, living with those men in the South of France was the great mammoth. 90 THE MAMMOTH The mammoth was like an Indian elephant, but with a coarse hairy pelt and its tusks had a slightly different curvature from that seen in the Fic. 60.—Engravings on ivory and bone made by ancient men, who lived in caves in the South of France at the time when the mammoth, reindeer, bear and hyzena inhabited Europe. The uppermost figure is that of a mammoth, the others represent reindeer. Indian elephant. It was a little bigger than the biggest Indian elephant. The mammoth has left its remains all over the Holarctic region. Even in our own country QI EXTINCT ANIMALS we are continually coming across tusks and teeth. In the Natural History Museum there is a whole skull, with enormous tusks, which was dug up in a brick-field at Ilford in East London about twenty-five years ago. From this brick- Fic. 61.—Engraving ona piece of an antler found in a cave in Switzerland. It represents very accurately a Reindeer. field I used to get many remains of mammoth, rhinoceros and hippopotamus when I was a boy. When this mammoth’s skull was found by the workmen, the authorities of the British Museum of that day sent down a man to remove the specimen with the oreatest care, and it is now at the Natural History Museum. 92 THE MAMMOTH ) The remains of the mammoth are found in *s00} oyeredos ay} oprIy tory sped AdoyZROT YVIIS OY UL posBoU [IIS oAv JooF oYy ASTIYA SpBoy oYyZ UO 04%4S PoLIp 8 UL poAdosoid St ULYS pue YSo] oy} JO [Bap poos VW “Sanqsaojoq *}Q Jo wmMesnut oy} UL MOU pUB BIIEqIG UL UoZOA’y PUNOF YYOWIUIBT OY JO UOJIPOYS OY T—'S9 “YIYL enormous abundance throughout the Holarctic It is probable that this huge beast 93 region. EXTINCT ANIMALS existed later in Asia and Siberia than in our part of the world. In the north of Siberia complete carcases of the mammoth and also of the hairy rhinoceroses are found in a frozen condition, with the skin, hair, trunk and soft parts complete. Fic. 63.—Skeleton of a male of the giant Irish deer (Cervus giganteus) dug up from peat in Ireland. At the beginning of the last century one of these frozen carcases was removed to the Museum at St. Petersburg. It is from this specimen, drawn in Fig. 62, that we know that the mammoth had a hairy skin. 94 THE GIANT IRISH DEER It is an interesting fact that the newborn young, both of the Indian and the African elephant have a complete coat of fairly long hair, which disappears in a few weeks. So the mammoth is not really peculiar in this matter. In Fig. 63 is shown the skeleton of the largest and most beautiful of all the deer tribe; it is now extinct, but existed later in Ireland than anywhere else, and in great numbers. The bones are found in the moss and bogs of Ireland. It was co-existent with primitive man, and perhaps survived in Ireland till nearly historic times. Why it died out there is a difficult thing to explain. As our explorations into the river gravels of only twenty or thirty feet depth have brought us into contact with the mammoth, I propose now to say something more about recent and extinct elephants, and to take a glance at the past history of the elephant tribe. Fig. 64 gives a careful restoration of the hairy mammoth as it must have appeared in life, and in Fig. 65 we have a photograph from life of the Indian elephant. In the Indian elephant you should note the comparatively small ear and the high forehead. 95 EXTINCT ANIMALS Fig. 66 is a photograph from life of the African It has a longer head and much elephant. ‘squeydoyo Aueut ul sev ‘prveagno you ‘10y}0 YoRo spavMo} pousny ore Yor jo syutod ey} ‘sysng podAdano ATsnormno ey puB “yeoo Arey oy} BION oytT UL poavodde qt se (sruahrunud soyday) Yyyourmmeyy oy Jo oangotd Areurseut uy—'F9 “Oly ‘ a —_— = : @ Oo - =i io) i) 32 ~ nm O [ et — + Fia. dogged.”” The mastodon’s skull (Fig. 72) shows far more of a projecting bony face or snout fo) ey 105 EXTINCT ANIMALS than does that of the elephant, and this would lead us to suppose that the mastodons are more primitive, that is to say, more like the ancient ancestors of elephants, than are the true ele- phants. An interesting fact in this connexion is that the young new-born elephant has a more ; i PLA ] Fic. 72.—Photograph of the skull of the American Mastodon (Mastodon americanus), from the specimen in the Natural History Museum. ‘““snouty ” skull than the grown-up elephant, as is shown by Figs. 73, 74 and 75. It is often the case that very young animals show features in which they resemble their ancestors, which disappear as the young creatures grow to full size. 106 NEW-BORN ELEPHANT’S SKULL It is not only in having a more elongated face that the American mastodon is of a more primitive build than the true elephants. Its teeth also are less peculiar than those of true elephants and more like in number and shape Fic. 73.—Skull of a new-born Indian Elephant, photographed from a specimen in the Natural History Museum. to those of the ordinary, more central kinds of mammals, such as the pig (see Fig. 50 in the last lecture). The elephant has _ two enormous incisor teeth in the upper jaw, in front—the tusks. There are no corre- 107 EXTINCT ANIMALS sponding teeth in the lower jaw. Then there is a gap in the series, and we come to the cheek teeth, which are very strange. The jaws, both upper and lower, are so short, and the teeth Fic. 74.—Section of the skull of a young Indian Elephant, to compare with the section of a half-grown elephant’s skull given in Fig. 75, in which the face has become relatively shortened and upright. Note in this and in Fig. 75 the curious conical nasal bone, which is like a small bony horn. are so big, that there is only room for one tooth or a tooth and a half on each side above and below at one time. An elephant only ever has 108 ELEPHANT’S TEETH three full-sized cheek-teeth on each side above and below (twelve in all), and these push from behind forwards—the first getting worn out and pushed forwards as the second comes for- wards, and this again wearing out and dis- Fic. 75.—Section of a half-grown Indian Elephant’s skull, with the first and second molar teeth in position (therefore more than twenty and less than twenty-five years old). appearing as the third pushes itself into place from the back of the jaw. Three little “ milk teeth ’’ or first-teeth of the molar series precede these on each side above and below, and are lost one after the other—between the second 109 EXTINCT ANIMALS and fifteenth years of life. The first big molar comes into place in the fifteenth year, and lasts for ten years, when its place is taken by the second, which is already showing its crown in LFr Fic. 76.—Lower jaw of an Indian Elephant, showing two molars on each side, the front ones wearing away as the back ones come. up into position. The transverse ridges on the teeth are well seen. the twentieth year. The third comes forward in the same way about twenty years later. The molar teeth of the Indian elephant and of the mammoth have a great number of narrow transverse ridges set across the crown of the IIo RIDGES ON ELEPHANT’S TEETH tooth. As many as twenty-seven of these ridges are seen on the biggest molar tooth when it is in place, and the whole surface is worn by grinding. In Fig. 76 the ridges on the teeth are shown, but not to the full number, as the front tooth is reduced in size by wear, and the hinder one has not yet got all its crown into Fic. 764.—The last molar of the lower jaw of a Mammoth, in order to show the great number of transverse ridges or segments of the tooth (as many as twenty-two in this specimen), a feature in which the Indian elephant and the mammoth are closely similar. play. In Fig. 77 is shown a photograph of the lower jaw of an African elephant. Only one big molar tooth on each side is in position, and it has eleven transverse ridges. This is the most the African elephant ever has. It will be seen, by comparing the figures, that the ridges of the African elephant are much wider than those of the Indian. The corresponding tooth of the 1a tf EXTINCT ANIMALS Indian elephant, owing to the narrower shape of the ridges, would have twenty-seven of them in view when fully “cut.’? Now there is no doubt that the increase in the number of the ridges and their narrow form is a late and special 4 1FT Fic. 77.—Lower jaw of an adult African Elephant, showing molars with only eleven transverse ridges, or “ lozenges.” character of the elephants. Their cheek-teeth would be more like those of pigs, tapirs and bears, if they had fewer transverse ridges. Accordingly, in correspondence with the view that the mastodons are more primitive in their 112 RIDGES OF MASTODON’S TEETH characters than the true elephants, we find that their cheek-teeth have very few transverse ridges—from two to five (fig. 78)—and that the jaw is relatively longer, so that there is room, [eo I Fig. 78.—Lower jaw of the American Mastodon, with two molars on each side, completely cut, showing respectively three and four transverse ridges only. Note also the elongated form of the jaw. not only for two complete crowns of molars .to _be in position on each side at the same time, but even forthree. Thus we approach nearer to the central or “‘ typical ” condition of the mammals’ teeth which, as we have seen in the pig (fig. 50) 113 I EXTINCT ANIMALS shows seven cheek-teeth in position on each side in each jaw at once—of which the front ones are second-teeth and were preceded by milk-teeth —whilst the three big back ones are not pre- ceded. In tracing the ancestry of living mam- mals through extinct ancestors of different suc- ceeding geological ages, we expect to find even the strangest and most curiously modified creatures, such as are the elephants in regard to their teeth and jaws and the horses in regard to their toes—preceded by forms which bring us nearer and nearer, as we recede into the past, to a sort of common form or “type” of the mammalian group—a_hairy-coated creature, with five toes on each foot, the typical dentition or tooth series of three incisors, one canine, four front or fore-molars, and three back molars on each side of each jaw, with three or four tubercles or knobs on the crowns of the molar teeth. And we do not expect this remote an- cestor to be very big—not much bigger than a dog—since great size is a peculiarity implying long and special predominance. A further point in which the American masto- don is more like the ordinary run of mammals than are the elephants, is that it has front teeth II4 THE LONG-JAWED MASTODON —a single pair—in its lower jaw when it is quite young. These drop out in the American masto- dons, but we have here a photograph (Fig. 79) of the skeleton of a much older mastodon, the re- mains of which were dug up in strata of the Middle Miocene (not only below Pleistocene, but Fie. 784.—Molar teeth of Mastodon arvernensis, photographed from specimens found in the Red Crag of Suffolk. These molars have five transverse ridges : that on the left shows the bony fangs beneath the crown of the tooth. below Pliocene and below Upper Miocene) in France. This skeleton is preserved in the Museum of Paris, where the photograph was taken. You will see that its head differs in many ways from that of elephants and the late American masto- don. It has an extraordinarily long lower II5 EXTINCT ANIMALS jaw, with two tusks init. A drawing of the side view of the skull is shown in Fig. 80, and you (Tetrabelodon) see Ss esr LS =] gi Bh Y Aa? As is Ee Ege Are 79.—Photograph of the complete skeleton of Mastodon F ta. can see how the two horizontal lower teeth must have played between the two curious downwardly-bent tusks of the upper jaw. 116 angustidens, from the Miocene strata of the South of France; taken from the specimen as it now stands in the Museum: of the Jardin des Plantes in Paris. THE LONG-JAWED MASTODON By way of parenthesis I must here mention a mastodon-like creature of the same age— which had no tusks in the upper jaw, but two huge tusks in the lower jaw, which is bent downward. This is the Dinotherium, found in the Miocene in Germany and other localities. It seems to have left no modern representatives, - Fig. 80.—Restored representation of the skull and lower jaw of Mastodon (Tetrabelodon) angustidens from a drawing prepared by Dr. Henry Woodward, F.R.S. and is a sort of extinct side-branch of the ele- phant family. The big tusks of the lower jaw were probably used for raking up roots in the mud of rivers and lakes. The Miocene mastodon, with the long lower jaw, is known as T'etrabelodon angustidens. The examination of its skeleton some years ago led me to the conclusion (as, indeed, was inevitable) I17 EXTINCT ANIMALS that it could not have had a depending trunk like an elephant has and such as the short-jawed mastodons certainly must have had. Its “trunk ’? must have rested horizontally on the pu S Ae Reye, Lg SS i, re ae Fic. 81.—The skull of Dinotheriwm giganteum, IKKaup, from the Miocene of Eppelsheim, near Worms, on the Rhine. long lower jaw between the upper tusks—and was in fact not a “‘ trunk” at all, but an elon- gated upper lip (Fig. 82)—representing the middle part of the upper jaw in a soft, flexible 118 THE LONG-JAWED MASTODON condition. It seemed to me probable that the elephant’s trunk had originated in this way: (Original. ) Fia. 82.—Drawing representing the probable appearance in life of the. Tetrabelodon angustidens, namely by the great elongation, in the first place, of the lower jaw and upper lip and jaw, 119 EXTINCT ANIMALS and by the subsequent shrinking of the lower jaw and bull-dogging of the bones of the face. Thus the elongated mid-part of the face—no longer supported by a long lower jaw—would gradually drop as the lower jaw grew shorter and shorter in successive ages, and at last it would hang down as a perpendicular trunk. In Fig. 83 I have endeavoured to represent this long-jawed mastodon (Tetrabelodon) open- ing his mouth and rearing his flexible, boneless upper jaw as does the living elephant rear his trunk (Fig. 84). It is very difficult to form a definite idea as to how the Tetrabelodon made use of his tusks and horizontal “ trunk.” The upper tusks have a sharp edge along the inner face strengthened by enamel, so that it is probable that, working against the tough skin pads of the lower jaw, they would serve for cutting vegetable matter. My friend Rudyard Kipling has given a different account of the origin of the elephant’s trunk, which he declares was formed by the pulling of the nose of an unfortunate young elephant which, before the days of trunks, stopped to drink some water from a pool, and was seized by an enormous crocodile just about the nose. 120 ORIGIN OF THE ELEPHANT’S TRUNK (TemsO) ..yunay ,, poyydn pue yqnour uedo YIM suapysnbup Uopojagnjaf, JO peo, 944 Jo SUIMBIP Y—'ES “OI EZ, EXTINCT ANIMALS The elephant pulled, and the crocodile held firm, and the result was the stretching of the elephant’s nose till it became a trunk. This story was not told to Mr. Rudyard Kipling by the boy Mowgli of the Jungle Book, who I feel sure must have heard from the elephants the Fic. 84.—Drawing of the head of the African Elephant with up-lifted trunk. history as I have given it unless, as is not un- likely, they have forgotten all about the way in which their trunks grew, and would reject, as most men and women do, the notion that they have been derived by slow change in long ages of time from other and more simple animals. 122 THE ORIGIN OF ELEPHANTS The history, suggested above, of the gradual production of the elephant in the later ages of the world’s history from a long-jawed creature has been wonderfully confirmed by the dis- coveries made in Egypt within the past four years by my friend Dr. Andrews, who is one of the staff of the Natural History Museum. Dr. Andrews was in Egypt four years ago on account of his health and joined a party of the officers of the great survey of Egypt, organized by Lord Cromer, in a visit to the Great Western Desert, the rainless, sandy waste lying west of the Nile, not very far from what is now called the Fayum, and where in Roman days was the great Lake Meris—now dried up to a mere brine-pool, in the salt water of which the freshwater fishes of the Nile still live. The surveying party in- tended to determine the geological age of these sands, which stretch for hundreds of miles, often rising into cliffs which are cut sharp by the wind and show horizontal stratification. Some fragments of bone had been recorded from this region twenty years ago by the traveller Schweinfurth, and Dr. Andrews, who is a special expert and authority in the interpreta- tion of fossil bones, was hopeful of securing PID) 123 EXTINCT ANIMALS some specimens for the Natural History Museum. He was rewarded far beyond his expectations. The party had to travel into an absolute desert waterless region, establishing a staff of camels which daily brought up water as far as three days’ march into the sandy wilderness, return- Pe A Fic. 85.—A scene in the Fayum Desert, showing the remains of silicified trees, embedded in the sands. From a photo- graph by Dr. Andrews. ing with empty tanks on their backs to fetch more. In Fig. 85 is reproduced one of many photographs taken by Dr. Andrews. It shows the flat sandy desert with some fossilized lumps lying in the sand which are the remains of trees. The geologists determined that the sands in 124 THE FOSSIL ANIMALS OF THE FAYUM this region were of Upper Eocene and of Miocene age, and from them Dr. Andrews brought home some very interesting bones. These included remains of a more primitive mastodon than any as yet known and of an animal which he called Meritherium (after Lake Meris)—which is the connecting link between elephants and the central typidentate mammals. But the col- lection included also remains of great carnivores, of Hyrax of great size (like the Syrian coney), of Sea-cows (Sirenians), and of Tortoises, and a Snake sixty feet long. The Egyptian Survey has since in the most enthusiastic way sent further expeditions into this desert to collect the bones of the extinct animals half-buried there, and Dr. Andrews, by the direction of the Trustees of the British Museum and further assisted by a generous donation from Mr. de Winton, has twice again in succeeding years camped out in the desert and excavated the sands by the aid of a troop of native diggers. In regard to the history of elephants, the upshot of Dr. Andrews’ most important dis- coveries is that we find living here in the Upper Eocene period (older than the German or French Miocene with its Tetrabelodon) an 125 EXTINCT ANIMALS elephant ancestor of the mastodon kind to which Dr. Andrews has given the name Palzo- mastodon. The skulls and many limb-bones of Fig. 1 Fig. 4 Fic. 86.—Profile views of a series of Elephant ancestors, from drawings by Dr. Andrews. 1. The Indian Elephant. 2. The American Mastodon. 3. The Miocene Tetrabelo- don (France). 4. The Eocene Paleomastodon (Egypt). 5. The Eocene Meritherium (Egypt). this interesting creature have been obtained, and are now reposing, some in Cromwell Road 126 THE ANCESTORS OF ELEPHANTS and some far away in the fine Museum of the Egyptian Survey in Cairo. In Fig. 86 a drawing (No. 4) is given of the skull of this Paleomastodon. The _ figure includes several other elephant forms. We have the skull and lower jaw of Tetrabelodon (No. 3), of the American mastodon (No. 2), and of the Indian elephant (No. 1). It will be seen at once how completely the Palzeomastodon skull fills in the series leading back from the bull- dog-faced elephants with short jaws to or- dinary mammals. It has a fairly long skull and long bony face, with two large—but not _very large—downwardly directed tusks. The jaws are long, but the lower one not so exces- sively long as that of Tetrabelodon (No. 3), and the cheek-teeth are there in nearly full number—as many as five in each half of each jaw. These are well seen in the view of the lower jaw given in Fig. 87 (No. 2), where the condition of the lower jaw of Paleeomastodon is clearly contrasted with that of Tetrabelodon (Mastodon angustidens, No. 3). In Paleomastodon we have arrived, by passing as far back as the Eocene strata, at an ancestral elephant-like creature which serves 127 EXTINCT ANIMALS to join the elephant stock on to more ordinary “normal”? mammals. I should say that this beast was not so very big—about as large as a Fic. 87.—Lower jaws of extinct Elephants, from drawings by Dr. Andrews. 1. The lower jaw (and above it the upper jaw) of Meritherium, showing six molar or cheek-teeth in position. 2. The lower jaw of Paleomastodon. 3. The lower jaw of Tetrabelodon. (Compare with the lower jaws of more recent forms shown in Fig. 76, 77 and 78.) fair-sized horse. Dr. Andrews’ great triumph, however, is the discovery of a somewhat smaller animal in the same deposits, which is undoubtedly an elephant, and yet at first sight 128 THE EARLIEST ELEPHANT ANCESTOR has no resemblance to one and probably had no trunk at all, as certainly it had only small Fic. 88.—Profile and palatine views of the skull of Meritherium Lyonsi, as restored by Dr. Andrews. Note the elongated form of the skull and the normal development of teeth, viz. six incisors (above), a right and left small canine and six molars on each side (above and below). tooth-like tusks, unworthy of comparison with 129 K ‘]7n48 049 Aq usAId ooUaptAG 8yy Aq peyysnt st ueyy yuna quordiout ue oy, o1oUL pue JoSae,T se payuesorador ory st di soddn oy Alqeqoig ‘paempoo,, ssi Aq UMeap ‘[eUITUR 949 JO OUT[YNO PeIOyso. B :qdAsny ‘unABy oy} JO qAoSop UeYSeAA JBOIKH 9YY JO BYBIYS OUEDO7;T JoddgQ oYy Ut ‘ummesnyy ALojstp] [BANYVN 9Y9 Jo ‘smoapuy “aqZ Aq pedoAoostp ‘UMUEYILWe; OU T— 68 “Ol gstpapee- Ri “SRR 130 THE EARLIEST ELEPHANT ANCESTOR the great ivory columns of later elephants. This is the Meritherium seen in Fig 86 (No. 5), and more fully exhibited in Figs. 88 and 89. As is obvious at once, the skull of Meritherium does not suffer from “ bull-dogging”’ at all; there is a fine, well-developed facial region, and the teeth are neither deficient in number nor greatly exaggerated individually. The ‘“ den- tition” (that is to say, the enumeration of the complete series of teeth) approaches closely to that of the central mammals with typical den- tition. In the upper jaw (as shown in Fig. 88) there are six front teeth or incisors, and it is the second of these on each side which is enlarged and is (so to speak) going to become the great tusk of the elephants. In the lower jaw there are four front teeth (see Fig. 87, No. 1). In the upper jaw we also find a small canine or dog- tooth ; next the incisors and the cheek-teeth in both upper and lower jaw are fully repre- sented, namely six on each side in each jaw— only one short of the type-number. And yet. these cheek-teeth are quite obviously and recog- nizably mastodon teeth. They have the trans- verse ridges of the mastodon tooth (two or three) and are in other features like those of mastodons. 131 EXTINCT ANIMALS Here, then, we have arrived at a form which undoubtedly was closely related to the ancestors of all the elephants—if not itself actually that ancestor—and in it we see the origin of the ele- phant’s peculiar structure. From this com- paratively normal pig-like Meritherium, the wondertul elephant, with his upright face, his dependent trunk, and his huge spreading tusks, has been gradually, step by step, produced. And we have seen some, at least, of the inter- mediate steps—the elongation of the jaws and increase of the size of the incisors in Palzo- mastodon—carried still further in Tetrabelodon, and then followed by a shrinkage of the lower jaw and final evolution of the middle part of the face and upper jaw as the drooping, wonder- ful, prehensile trunk. 4 So much for the “ great sagacious elephant ” and his extinct relatives. Let us now turn for a few minutes to the most beautiful and the most helpful to man of all animals—the horse, nobler as he is bigger and stronger and more beautifully shaped, than man’s other animal companion, the dog. The horse is curiously different from the central typical mammals in that he has only one toe on each foot instead of five, and further, 2M} 132 THE ANCESTORS OF THE HORSE in the complex pattern of his teeth. But immense numbers of extinct horses and horse- like creatures have been dug up, and we now know quite clearly all the stages leading from living horses back to four-toed and ultimately to five-toed ancestors. First of all I will put Fic. 90.—Photograph of a model of a thoroughbred English horse, by Vashtag; one of a series in the Natural History Museum. before you a photograph of a very beautiful model of an English thoroughbred (Fig. 90). There are a set of these models, both of horses and cattle, in the Natural History Museum : each is carefully modelled to one-fourth the 133 EXTINCT ANIMALS size of nature. They were executed by a Hungarian artist for the exhibition at Buda- Pesth some years ago, and we ought to have such a series made now in England of samples of all the best breeds. It is the only way of keeping a really complete and satisfactory record, and such models of known horses and cattle, made to-day, would be of immense interest and value in fifty years’ time. But they are costly things to make, and can only be undertaken by the rich owners of race-horses and pedigree bulls. Fig. 91 shows us the fore and the hind foot of the horse. As is very usual with photo- graphers and those who prepare drawings and lantern-slides, the artist has placed the hind- foot in front and the front-foot behind. The hind-foot (that on the left) shows the heel-bone or “hock” (the caleaneum) standing forth at the top of the ankle. Below you see the three bones which constitute,as in our toes and fingers, what is called the digit. Then there is a long bone, which is the meta-tarsal bone. In the front-foot the similar bone is called the meta- carpal. At the top of these are several short bones jointed together; these are the tarsus or 134 Fic. 91.—Hind and fore-foot of an English cart-horse, to show the single toe of three pieces—or joints—and the small splint-bones on each side of the long metatarsal and meta- carpal bone. 135 EXTINCT ANIMALS ankle and the carpus or wrist (so-called “‘ knee ”’ of the horse’s front-leg) respectively. You see, the horse walks on the very last joint of its toes, and keeps the foot and the hand upright, so that the heel is right above the toe instead of behind it, as in ourselves and the bears. On each side of the long bone of both fore and hind- foot you will see a small long bone, narrow and delicate. The nearer one of these delicate bones is not very clearly shown in the photograph, but still can be made out. These delicate e) ‘* splint-bones,” as they are called, are all that remain in the modern horse of two additional toes. There was a time when horses had three toes—far back in the Miocene strata we find horses which had three well-developed toes, each with a hoof resting on the ground (the Meso- hippus and Anchitherium), and earlier than that we find a horse-like creature (Hyraco- therium) with three nearly equal-sized toes on the hind-foot and four on the front foot (Fig. 92). In the Pliocene we find a_ three-toed horse in Europe known as the Hipparion (and a similar kind is dug up in America), which had three toes on each foot; but the side toes were getting small, were in fact like the “ petti-toes ”’ 136 THE ANCESTORS OF THE HORSE of the pig, and of cattle, and of the reindeer. They did not touch the ground (Fig. 93), and Fic. 92.—Hind-foot (to the left) and fore-foot (to the right) of the horse-ancestor, Hyracotherium. The fore-foot is seen to have four toes in full development. Photographed from specimens in the Natural History Museum. ° were evidently on the way to disappearing, leading to the single-toed modern horse, with 137 EXTINCT ANIMALS its splint-bones, as the sole representatives of Fic. 93.—The hind- and the fore-foot of Hipparion, one of the three-toed ancestors of the horse. The side-toes were * pettitoes ’ and did not reach the ground. the two outer toes. Occasionally living horses ’ 13) THE ANCESTORS OF THE HORSE are born with two complete little toes provided with hoofs and attached to the splint-bones, one on each side of the big central toe, “ throw- ing back,” as the term is, to their three-toed ancestors. Beyond the stage, with four equal toes on the front foot and three on the hind- foot, which is exhibited by a quite small horse- Fic. 94.—The skeleton of Hyracotherium, an ancestor of the modern horse, found in Eocene strata. like creature—the Hyracotherium shown in Figs. 94 and 95—we can trace the pedigree of the horse to a five-toed ancestor, the Phenacodus (Fig. 96). The later stages of this history, from the Mesohippus to the modern horse, have been traced by very abundant fossil remains of many steps or stages in the gradual change. Not only 139 ANIMALS EXTINCT has there been a gradual change from the three- toed to the one-toed condition, but there has ‘y10K MON ‘uMosn{ UWeotrouty oy Aq ponsst Sut -MBIp B Joypy “Sop o5uey B UBYY JOSSIq YoU eVM 4 = “4OOJ-puTY ayy UO vary pue 4ooj-yuoay oy} UO S90Z MOF YALA ‘as1oy sYyy JO JOJSOOUB OUBDOT UB ‘untdeyyooRrAyyT oy} Jo aouvrvodde oiqeqoid oyy jo UuoTyRIOYSoY—'CGH “OTT been a great increase in size, and moreover the cheek-teeth have gradually become more and 140 THE ANCESTORS OF THE HORSE more complex in the pattern which they show when worn down. In Fig. 97 crowns of the Fic. 96.—Skeleton of the Phenacodus, a five-toed Eocene animal related to the ancestors of the horse. cheek-teeth of the Mesohippus are represented, and in Fig. 98 the crown of an upper molar of Fic. 97.—Cheek-teeth, or molars, of the upper and lower jaw, left side, of Mesohippus Bairdii, from the Middle Oligo- cene of South Dakota. ae a recent horse. There are a great number of I4I EXTINCT ANIMALS interesting details in the history of the changes of the teeth and toes of the ancestral series of horses which it is not within my scope to describe here, but they may be studied on speci- mesostyle parastyle metastyle metacone~ Ne , metaconule.+ pretoloph pete ph : Fic. 98.—Upper molar tooth of arecent horse. A, uncut and unworn ; PB, C, D, in successive stages of wear. mens of a variety of ancestral horses which have been set out for the purpose in the Natural History Museum. The rhinoceroses of to-day—the unicorn or 142 RECENT AND EXTINCT RHINOCEROSES Indian rhinoceros and the two-horned African rhinoceroses, one with a pointed upper lip and the other with a square, broad mouth—have been preceded by a whole regiment of extinct rhinoceroses, whose bones and skulls are dug up in the Pleistocene, Pliocene and Miocene strata. In Fig. 99 is represented the complete Fie. 99.—The skeleton of Rhinoceros antiquitatis, the woolly rhinoceros of the late Pleistocene period in Europe and Siberia. skeleton of the commonest kind of fossil rhino- ceros, the skull of which was dug up in London the other day and is shown in Fig. 5. This rhinoceros had a hairy coat like the mam- moth, and is found sometimes with the mam- moth in frozen gravel in Siberia. The living 143 EXTINCT ANIMALS rhinoceros most like it is the African square- mouthed rhinoceros or Burchell’s rhinoceros (Rhinoceros simus), misleadingly called some- times the white rhinoceros (Fig. 100). Many of the extinct kinds of rhinoceros had two horns, one behind the other like the African rhinoceros. The horn of the rhinoceros is truly horny in Fic. 100. —- Photograph of a stuffed specimen of the sue mouthed African Rhinoceros (Rhinoceros simus) preserved in the Natural History Museum. substance, and fibrous. It is not bone, as are the horns of stags, nor has it a core of bone as have the horns of sheep, cattle and antelopes. Some- times, however, there is a fairly big boss of bone, which forms a sort of base or pedestal for the horny horn. One great extinct beast (the Elasmotherium), allied to the rhinoceroses, had a 144 ET GaN wal it f “ay THE TITANOTHERIUM 145 EXTINCT ANIMALS great horn, carried on a huge boss on the middle of its head instead of on the nose, whilst in the Miocene of North America complete skeletons have been found of an enormous creature allied Fic. 102.—Photograph of a skull of Titanotherium in the Natural History Museum, showing the huge molar teeth. to the rhinoceroses, but having a pair of horns, perched side by side on the nose, instead of one in‘ the mid-line, or two placed one behind the other. The skeleton of this great beast, called 146 THE EXTINCT DINOCERAS Titanotherium, is shown in Fig. 101, and in Figs. 102 and 103 photographic views of the skull are given. As large as the rhinoceros, but having a very different arrangement of the bones of its wrists and ankles, and very different teeth and horns, Fic. 103.—Side-view of the skull of Titanotherium, to show the two bony upgrowths of the nasal region which carried horns. Photographed from a specimen in the Natural History Museum. are the extraordinary creatures known as Dinoceras, whole skeletons of which have been disinterred from the Upper Eocene of Wyoming in the United States. As many as two hundred individuals were studied by Professor Marsh, who has written a large treatise on them. These creatures had three pairs of horns on the 147 EXTINCT ANIMALS top of the head (Fig. 104) and a pair of great tusks formed by the enlargement of the upper canine teeth. The horns are outgrowths of the bone of the skull and were probably covered by Fic. 104.—Skeleton of Dinoceras mirabile, from the Upper Eocene of Wyoming, U.S.A. hardened skin. The probable appearance of this creature in life is shown in Fig. 105. A very interesting fact has been observed about the brains of these most ancient big mam- 148 THE DINOCERAS mals, viz., the Dinoceras occurring so far back as in the Upper Eocene, and the Titanotherium Probable appearance in life of the Dinoceras mirabile of North America. Fig. 105. of the Lower Miocene. We can get castings from the interior of the skulls and compare them 149 EXTINCT ANIMALS with those of recent rhinoceros, hippopotamus and horse (Fig. 106), and it is found that although Dinoceras and ‘Titanotherium were _ bigger than the largest rhinoceros of to-day, yet they had quite small brains, not more than an eighth the volume of that of the recent big animals. The subject has not been so fully Z . oe § ry 6 a K : Fic. 106.—Photographs of plaster casts of the brain-cavity of A, Dinoceras; B, Hippopotamus; C, Horse; and D, Rhinoceros: to show the relatively very small size of the brain of Dinoceras. looked into yet as it deserves, but it seems that modern animals, the animals which have sur- vived, have much bigger brains than those which died out in the Eocene and Miocene times, and it is probable that they have survived to a large extent because of the value to them, in the struggle for existence, of the bigger brain. 150 THE SIZE OF BRAINS It seems that a small brain may serve very well to guide the great animal machine in established ways, but in order to learn new things in its own lifetime an animal must have a big brain— indeed, a very big brain. And the kind of animal which can learn—that is to say, can be educated —will, in the long run, beat the kind which has too small a brain to be capable of learning. This is the significance, not only of the big brains of recent rhinoceros and horse as com- pared with those of Titanotherium and Dino- ceras, but it is also the significance of the big brain of man, which is far bigger than that of any other animal in proportion to the bulk of his body and limbs. Another huge horned animal has quite lately become known which in some ways resembles Titanotherium and Dinoceras, but has to be kept apart from them on account of being really unlike them in its teeth and skull and feet-bones, although having a general resemblance to them in outline and bulk. This creature was found only three years ago in the same Upper Eocene sands of the Egyptian Fayum from which Dr. Andrews obtained the ancestors of elephants. The skull of this most strange animal is shown I5I EXTINCT ANIMALS in Fig. 107, and a representation of what we suppose it looked like in life is given in Fig. 108. This wonderful beast was discovered by Mr. / / eo, zy ig ea TVS If Lr, Wy iY / yy YY Uy Si x, af Y, € WH {{ ul oF iq (l K | AWS : ash Fie. 107.—Drawing of the skull of Arsinoitherium Zitteli (Beadnell), from the specimen preserved in the Natural History Museum. The skull was found in the Fayum Desert, and is nearly three feet in length. Beadnell of the Egyptian Geological Survey, and the name Arsinditherium was given to it 152 QUEEN ARSINOES GREAT BEAST by him because the Egyptian queens of Greek named Arsinde—had a palace near where race (Original ) Fie. 108.—A drawing, showing the probable appearance in life of Arsinoitherium. the bones were dug up. Two thousand years ago many parts which are now sandy desert 153 EXTINCT ANIMALS were well-watered and under cultivation. The drawing given in Fig. 107 is prepared from a skull in the Natural History Museum, where we have brought together portions of several other skulls and the complete set of bones of the skeleton dug up, some by Dr. Andrews and some by the energetic officers of the Egyptian Survey. The huge pair of horns are entirely bony out- growths of the nasal bones, and are hollow. aa = a Fic. 115.—Photograph of the specimen of the Okapi (Okapia erichsoni) obtained by Sir Harry Johnston near the Semliki river in Central Africa. The specimen is a female, not fully, grown, and is of the size of a very large donkey. Africa. As they live in these immense dark gloomy and damp forests they are very difficult to shoot or to catch, and moreover they are not abundant. The natives cut the striped skin into girdles and bands for ornament. Two of these were sent home by Sir Harry Johnston 163 EXTINCT ANIMALS before the animal was known, and were described as coming from a new species of zebra which was named Hquus Johnstoni by Dr. Sclater (see Fig. 117). 1% Fic. 116.—Photograph of a skull of a male Okapi, showing the simple pointed bony horns like those of the Samotherium. The horns were not enclosed in a horny case as are those of cattle, sheep and antelopes. Some people, on account of the Okapi being striped somewhat like a zebra, whilst it has the double hoofs of giraffes and also paired horns, have supposed that it might be a hybrid or ‘““mule ”’ between a zebra and a giraffe. This 164 NO HYBRIDS IN NATURE is, however, a supposition which every naturalist knows to be quite out of the bounds of remotest probability. It is a fact that “mules” or hybrids never are produced by animals living in their natural conditions, except in a few rare cases among aquatic animals whose eggs are fertilized in the water after they have been laid. Fic. 117.—Photograph of the two ‘‘ bandoliers ’’ cut from the striped part of the skin of an Okapi,which, when sent home by Sir Harry Johnston, were at first thought to have been cut from the skin of a new kind of zebra. And no one has ever produced, even in cap- tivity, a hybrid between any creatures so unlike each other as a double-hoofed and a single-hoofed mammal. There are a good many instances in which small living animals were represented in the past by gigantic forms very close in structure to the little living beasts, but of much greater 165 EXTINCT ANIMALS size. Hence it is concluded that these particular living animals are the reduced and dwindled representatives of a race of primeval monsters. There is some truth in this, as you will see from the history of the living sloths and armadilloes of South America, as compared with the giant extinct sloths and armadilloes dug up in that country. The same relation is true as to the kangaroos and wombats now living in Australia as compared with gigantic extinct creatures of the same kind (Fig. 182) which are dug up in Australia in sands and morasses of late geological date. Butitis a great mistake to conclude from this that it is a law of Nature that recent animals are all small and insignificant as compared with their representatives in the past. That is simply not true. Recent horses are bigger than extinct ones,and much bigger than the three- toed and four-toed ancestors of horses. Recent elephants are as big as any that have existed, and much bigger than the earlier elephantine ancestors. There never has been any creature of any kind—mammal, reptile, bird, or fish— in any geological period we know of, so big as some of the existing whales, the Sperm Whale, the Great Rorqual, and the Whale-bone whales. 166 BEASTS OF MONSTROUS SIZE It is true that there were enormous reptiles in the past, far larger than any living crocodiles, stand- ing fourteen feet at the loins and measuring eighty feet from the tip of the snout to the tip of the tail; but their bodies did not weigh much more than that of a big African elephant and were small compared with whales. So let us be under no illusions as to extinct monsters, and proceed to look at those of South America with simple courage and confidence in our own day. South America (see the map, Fig. 42) was not so long ago a vast island and connected at an earlier period with Australia. Later it has joined on to North America. Its own peculiar productions in the way of animals appear to be the members of the group of mammals called Edentata—very peculiar forms, with strange teeth, and none at all in the front of the jaws. From North America, when it joined on there, it received the mastodons, horses, tigers, tapirs, and other kinds produced in the Holarctic area. This seems to have led to the dying out of the big kinds of Edentata, and now there are only the small tree-sloths (Fig. 118), the small arma- dilloes (Fig. 119) and the strange-looking ant- eaters. But in quite late geological deposits in 167 EXTINCT ANIMALS South America we find the bones of gigantic armadilloes and of gigantic ground sloths, which oF Photograph of a stuffed specimen of the Two-toed Sloth (Chola- pus didactylus) hanging from a branch of a tree. Fria. 118. lasted on till the time when man appeared on the scene, though nowextinct. A great variety of large creatures of the kinds known as Edentata 168 ARMADILLO THE ‘(snsoypa sndfispq) opnjog 10 OT [[peulry hae ey Jo uourtoods poyyngs B jo ydeisojyoyg—'6IT “OT 169 ysiug oyy Jo seaqysnay, oy} Aq JUSy st onsy stp, “WMOYs SI jo dnowas ouog poyutofun oy], “Boltotuy YINOG UL e7BIS CUTFUO uopoyd Ap) poles [BuUtrUB O¥T][-OT[IpwUAIB your 4xo yeoid oy} JO UOJo[o “mInesny ‘peay oy Jo doy pur ‘[rey ‘Apoq oy4 Say oy} Jo oussozslo[q OY} Woy ys oy} jo Suraerq—'OZl “O17 EXTINCT GIANT ARMADILLOES preceded these in earlier geological times in South America. The Glyptodons (Fig. 120), of which there are For the skeleton of the same Megatherium giganteum, as big as an elephant, found in the Pleisto- animal, see the photograph on p. 7. cene gravels of South America. Fic. 121.—Probable appearance in life of the Giant Ground Sloth, the several different kinds, were enormous arma- dilloes, as big as an ox. Like the recent little armadilloes they carried a hard case formed by ity EXTINCT ANIMALS bones in the skin, but this was not jointed so that they could roll up into a ball, as can the living armadilloes. The Megatherium (Fig. 121) was nearly as big asanelephant, and was very closely similar in its skeleton and teeth to the little living sloths of to-day. But it stood on the ground and pulled the trees down in order to eat the tender young branches instead of climbing up into the trees and living there as the present sloths do. Not quite so big as the Megatherium was the Mylodon, which lived at the same time. The remains of both are found in the comparatively recent (Pleistocene) gravels of the Argentine Republic. The skeletons of these animals may be seen side by side in the Natural History Museum. | In Fig. 122 is represented the skeleton of the Mylodon, and just above it, for comparison, is placed the photograph of the skeleton of the two-toed sloth. The relative sizes of the two are shown and the sloth’s skeleton is placed in the same position as that of the extinct Mylodon, although in life it is always hanging from the branches of trees and never goes on all fours on the ground. 172 THE MYLODON The Mylodon had, we know, a number of little bony pieces scattered in its skin in the L Fic. 122.—The skeleton of Mylodon robustus, one of the giant Ground Sloths of the Argentine, about as big as a large bull. Above it is placed the skeleton of a recent Tree- Sloth for comparison. Both skeletons are reduced to the same scale. 173 EXTINCT ANIMALS region of the back, like the pieces forming the bony case of the armadilloes and Glyptodons, but not fitted closely together. It was sup- posed that the Mylodon, like all the gigantic Edentata of South America, had long ceased to exist and was extinct as long ago as the mammoth and the woolly rhinoceros of our Fic. 123.—View, looking outwards, from the mouth of the cavern on the fiord of the Ultima Speranza in Southern Patagonia, in which have been found the skin and hair and the bones with cartilage, blood and tendon and the dung of the Mylodon and other animals, proving its co-existence with man and its survival until a period estimated variously at fifty or a thousand years ago. own country. But about seven years ago a traveller (Dr. Nordenskj6ld) found in Patagonia, at the end of a fiord near the Chilian coast, a vast cavern (Fig. 123), and from this cavern the white settlers living in a farm close by had re- moved an enormous piece of skin (Fig. 124) covered with greenish-brown hair and studded on 174 FRESH REMAINS OF MYLODON its inner side with little knobs of bone (Fig. 125) ! The skin was dry, but undecomposed, and when soaked in water gave out the smell of decom- posing animal matter. It was evidently a piece of the skin of a Mylodon which had survived in Fic. 124.—Photograph of a piece of the skin of the Mylodon (also called Grypotherium darwini) showing the coarse greenish-coloured hair. From a specimen found in the cave of the Ultima Speranza in South-west Patagonia. this region until modern times! Further ex- plorations were made in the cavern by Dr. Moreno, of the Museum of La Plata, and by other persons, and as a result an immense quantity of bones were obtained and more portions of 175 EXTINCT ANIMALS the skin of Mylodon with the hair on. The cavern had been inhabited probably several centuries ago by Indians, and human bones as 99 well as “ forks ” made out of dogs’ bones (Fig. 126) were obtained. The remains of as many as Fic. 125.—The under side of the same piece of skin as that shown in Fig. 124. It shows the small oval ossicles scattered in the deep substance of the skin. twenty Mylodons have been obtained from the cavern, and many of the bones have been cut or broken by human agency, the inhabitants of the cave having fed upon the Mylodons and split the bones to obtain the marrow! Some of the Mylodon bones, skulls, jaw-bones, leg- 176 THE FRESH REMAINS OF MYLODON Fic. 126.—Photograph of various specimens found with the remains of the Mylodon in the Ultima Speranza cave. 1. The lower end of the humerus (upper arm bone) of a very large jaguar (felis onca). 2. Molar tooth of an extinct horse (Onohippidium). 3. End of femur of a huge rat (Megamys). 4. Upper jaw of guanacho (Auchenia). 5. Molar tooth of same. 6. Lower end of lowest leg-bone of the rhea. 7. Foot-bone of the jaguar. 8. Hoof-bone of the fossil horse. 9, 10, 11. Dung of the Mylodon. 12, 13. Two bones of a Dog, with ends sharpened by human agency. 14. Distorted human shoulder-blade, probably of a woman. bones, etc., are smeared with blood and the soft tendon and membrane are still attached. The cartilage at the end of the long bones is still 177 N EXTINCT ANIMALS in place, dried and cracked in the drying. Not only that, but great balls of dung were found made up of the remains of masticated grass, Fie. 127.—Photograph of remains of Mylodon from the cave of the Ultima Speranza. 1. Shaft of tibia. 2. Bone ofa claw. 3, 4, 5, 6. Claws (horny). 7, 8. Rudimentary toe bones. 9, 10. Cervical vertebrae. 11. Lower end of scapula. 12, 13. Broken bones. indicating that the Mylodons lived in the cave. Moreover, a very large quantity of cut grass was found in the cave, and it has been surmised 178 MYLODONS LIVING IN THE CAVE that the Indians kept the Mylodons alive in the cavern and fed them with hay brought in from the outside. Specimens of these objects and of others to be mentioned below are now in the 3 Fic. 128.—Photograph of a *‘ barrel-full of bones’? obtained by prospectors from the cave of Ultima Speranza, three years after the first finds, and offered for sale to the Natural History Museum. Unfortunately it was not possible to send a reply to the owners in time, and the collection was dispersed. Skulls, jaws, and other bones of Mylodon are to be seen as well as a large skull of a jaguar, and bones and teeth of horses. Natural History Museum, and some idea of their number and variety may be formed from the photographs reproduced in Figs. 126 to 131. Besides the remains of the Mylodons and of 179 EXTINCT ANIMALS man—all lying loosely covered by a greater or less depth of blown sand, and in some parts by chopped hay—the cavern has yielded bones and teeth and many horny hoofs of horses, appar- ently belonging to the extinct and very peculiar ne -* a+ tes “Leek ana rr ey 3 ; AF Fic. 129.—Photograph having the same history as that shown in Fig. 128. South American genus Onohippidium, the skull and bones of a very large kind of jaguar, the skull of a young lama, and bones of other kinds. We have not yet a full account of all that has been found in the cave, nor have the contents, 180 THE CAVE OF THE MYLODONS unfortunately, been removed with sufficient care to enable us to say which were lying more deeply in the sand and which were at a higher level and therefore more recently living. The cavern is in a very remote spot and seems to Fie. 130.—Photograph of three pellets of the dung of the Mylodon from the cave of Ultima Speranza. offer some peculiar difficulties to explorers, for neither Sir Thomas Holditch nor Mr. Hesketh Pritchard, the latter of whom started for the purpose, succeeded in reaching it. It is stated that there are other caverns of a similar nature 181 EXTINCT ANIMALS in the neighbourhood. A great peculiarity about the occurrence of the remains of animals in this cavern is due to the fact that it hasa dry sandy bottom. The bones are not embedded in Fic. 131.—Photographs of the leg-bone (tibia) of Mylodon, from the cave of Ultima Speranza, to show the dried and cracked cartilage on the ends (articular surfaces) of the bones. b) *‘ stalagmite ” as is the case in the bone-caves of England and France, and whilst they are quite unaltered and full of animal matter, the horny and tendinous parts of many of the animals, such as skin, hair, claws and hoofs, and 182 WHEN WERE THE MYLODONS ALIVE? the soft dung of the Mylodon, are preserved un- changed. It is quite certain that in any known cavern in Europe such remains would be destroyed in the course of fifty years by putre- factive bacteria, and were the conditions too dry for that process to continue, the remains would have been consumed by scavenger beetles and other insects within the like period. The climate of South Patagonia, where the cavern exists, is similar to that of Devonshire. It isa moist climate, although the cavern itself is not damp nor subject to inundation by streams. There is nothing in the sandy soil of a preser- vative nature, and it seems at first sight impos- sible to suppose that the soft dried remains, skin, claws, blood, etc., can be more than fifty years old. Yet the horses’ hoofs and bones seem to belong to the extinct Onohippidium, and there is no record or tradition among the present race of Indians (in spite of some statements to the contrary) of any huge beast corresponding to the Mylodon. Altogether the case is a very puzzling one, and excites a very eager desire for further exploration. A noticeable fact bearing on the matter is that the whole of the southern part of South America has been submerged 183 EXTINCT ANIMALS rapidly and has rapidly risen again and is still rising at the rate of two feet a year in some parts, within the late Pleistocene period. Possibly the rocks and high lands where the Mylodon cavern occurs formed an island during the submergence where a number of individuals of the earlier fauna took refuge and survived until the re-elevation of the land, and so lived on in the present condition of the land surface until fifty or a hundred years ago. Possibly, though by no means probably, the Mylodon is still living in similar caverns in this region, as yet unvisited by man. In Australia, the land of the marsupials or pouched mammals, the bones of gigantic crea- tures have been found belonging to that peculiar tribe. Giant kangaroos, twice as tall as any living kangaroos, are thus known. But there are also remains of some extraordinary animals, like wombats and koalas, only as big as the largest rhinoceros or a small elephant. One of these is the Diprotodon of Owen, known to him by its skull and the rest of the skeleton, excepting the feet. The skullis drawn in Fig. 132 with a human skull beside it to give ascale. In Fig. 133 is given Owen’s restoration 184 GIANT BEASTS FROM AUSTRALIA of the complete skeleton with the exception of the feet. These have now been found by Dr. Stirling, of South Australia. A number of com- plete skeletons of this huge beast were found embedded in the mud of a great lake or morass. (ij >») ¢ : _y!] oe \ engi Rou i Fie. 132.—Drawing of the skull of the Giant Australian Marsupial, Diprotodon, preserved in the Natural History Museum. By its side is placed a drawing, to the same seale, of a human skull. In the photograph (Fig. 134) the lake is shown, and one of the great skeletons is seen in the foreground. The bones were in a very friable state, but Dr. Stirling has succeeded in pre- serving them and has secured the complete feet. In Fig. 135 the right hind-foot is shown. It 185 EXTINCT ANIMALS is expected that the complete skeleton will be put together and exhibited in the Natural History Museum before very long. Fie. 133.—The restoration of the skeleton of Diprotodon, as drawn by the late Sir Richard Owen. It will be observed that the feet were not known when this drawing was made. The oldest remains of mammals, which we know of, are found in the Oolitic and Triassic strata and consist of very small lower jawbones with their teeth, embedded in very fine-textured 186 AUSTRALIAN MONSTERS rock. It is usually held, on account of the form of the angle of the jawbones, that they belonged to small marsupial mammals. They are very small, few of them as much as an inch in length, and one of them we have already seen in Fig. Fie. 134:—Photograph of the morass or lake in South Australia in which the remains of several specimens of Diprotodon have been recently discovered. One of, the skeletons is seen lying in the mud in the foreground. 57 enlarged to ten times its natural length. It is probably due to their density and hardness that the little jaw-bones- have been embedded and preserved in these ancient rocks, whilst the rest of the skeleton is lost to us. The first 187 EXTINCT ANIMALS specimens of jaw-bones of this age were obtained seventy years ago in the Stonesfield Slate near Oxford by two undergraduates of the University, and it was at first supposed, on account of their occurring in such ancient rock as the Oolite Fic. 135.—View of the upper surface of the right hind-foot of Diprotodon, as discovered by Professor Stirling of Ade- laide, South Australia. The left-hand figure has the astragalus (ankle-bone) removed, whilst it is in place in the right-hand figure. (see list of strata on page 60) that they must be jawbones of lizards. Soon, however, the fact was noticed that the teeth had double fangs, and it became clear from this, as well as the shape of the jaws and teeth, that they had 188 MAMMALS OF THE MESOZOIC PERIOD belonged to small mammals. In Fig. 136 two of these very ancient mammalian jaws are figured. Fic. 136.—Lower jaws of the ancient Mammals, Dromatherium (upper—Trias), and Dryolestes (lower—Jurassic), mag- nified about 24 times linear. CEP TER Wy THE GREAT EXTINCT REPTILES—DINOSAURS FROM THE OOLITES—THE PARIASAURUS AND INOSTRANSEVIA FROM THE TRIAS OF NORTH RUSSIA AND SOUTH AFRICA—MARINE REPTILES. N the next two chapters I propose briefly to bring before you a few examples of extinct reptiles, birds and fishes, and to take the very shortest glance at the host of invertebrate shell-fish, insects, star-fishes and such like extinct animals whose name is legion. We will proceed at once to the reptiles. You will see from the list of groups of reptiles which I gave to you in a former chapter (p. 58) that there are four big orders or groups of living reptiles: (1) the Crocodiles ; (2) the Tortoises (Chelonians); (3) the Lizards; and (4) the Snakes. The lizards and snakes are in their real structure so much alike that they are con- 190 EXTINCT REPTILES sidered as one double order. Extinct repre- sentatives of all these orders are found right away down through the Mesozoic strata to the Trias (see table of strata, p. 60). But there is nothing very astonishing about them excepting the large size of some of the extinct tortoises and snakes, and the fact that the older extinct crocodiles had the opening of the nose-passages into the mouth-openings, which we and all air- breathing vertebrates also possess, placed far forward as they are in the more primitive air- breathers, whereas living crocodiles have them pushed ever so far back to the very furthest recess of the long ferocious mouth, from which arrangement it results that the modern crocodile can have its mouth full holding the body of a victim under water whilst the air passes from the tip of its nose through the long nasal passage to the very back of its mouth and so to its lungs. This convenience was not enjoyed by primitive crocodiles. The great interest in regard to extinct reptiles centres in those which were so entirely different from the reptiles of to-day that naturalists have to make separate orders for them. Many of them were of huge size. They flourished in the 191 EXTINCT ANIMALS Mesozoic period and abruptly died out; at any rate their remains disappear from the rocks at the close of the Chalk or Cretaceous period (see the table of strata, p. 60). These extinct orders of reptiles are the Dinosaurs, the Thero- morphs, the Ichthyosaurs, the Plesiosaurs and the Pterodactyles. They are a prominent example of that kind of extinct animal which is not the forefather, so to speak, of living ani- mals, but of which the whole race, the whole order, has passed away, leaving no descendants either changed or unchanged. To begin with the Dinosaurs. They are a very varied group and mostly were of great size. They seem to have occupied in many ways the same sort of place on the earth’s surface which was filled at a later period by the great mam- mals, such as elephants, rhinoceroses, giraffes, giant kangaroos, etc. Preying on the vege- table-feeding kinds there were huge carnivorous dinosaurs, representing the lions and tigers of to-day. Yet the mammals I have mentioned are in no way descended from these great reptiles. They came from another stock, and only superseded them on the face of the earth by a slow process of development, in which the 192 NEW ZEALAND TUA-TARA THE ‘OZIS [BANZVU OY} JO pALYJ-o0UO SI oINSYy oy, ‘sngnj~ound uopouaydg se uMOUy ‘B1Eq-BNT, PlVZI'T puvlwoz MON oY} JO ofl] WOAy UOye, YSvo eV Jo yYdeadojoyG—/ET “DI 193 EXTINCT ANIMALS great reptiles disappeared and the great mam- mals gradually appeared and took their place. Some of the forms assumed by the great Photographed one half the natural size. Toad), _Fia. 138.—Phrynosoma orbiculare (Mexican Horned Lizard or Horned Dinosaurian reptiles are not unlike the forms of the small scaly lizards of to-day (see Figs. 137, 138, 139, 140); but on the whole the Dino- saurs were more like mammals in shape, stand- 194 THE CHLAMYDOSAUR -9U0 0} poydeasoqoy,g ‘OZIS [eInyBU OY} paIyy ‘BIyerysny ‘puBlsuson?) urory “bury snunvsophunjyo—6el ‘OIL 195 EXTINCT ANIMALS jo ozIs oy} jrey poydeasojoyg “( oanyeu prezry popparg yeory) snaquphrbh snanuoz—'oFT “OI 196 THE GREAT DINOSAURS ing well up on the legs. We do not know much about their skin; it was probably smooth and with only small horny scales on it, as in many living lizards, and often had great horns and crests growing out of it. But we know the Fic. 141.—Drawing of the skeleton of Iguanodon bernissar- tensis. From the ground to the top of the head as the animal is posed, is about fourteen feet. complete skeletons put together from bones chiselled out of the hard rock in which they are found, and we know that in important matters of shape and build the skeleton was different from that of living reptiles. The great size to which some of the Dinosauria attained is 197 EXTINCT ANIMALS shown by the thigh-bone of one found in the United States, and called Atlantosaurus—photo- Fra. 142.—Probable appearance of the Iguanodon in its living condition. graphed in Fig. 6, p. 11. This thigh-bone is one third as long again as that of the biggest elephant known. 198 THE IGUANODON In Fig. 141 is shown the complete skeleton of the Iguanodon. This great Dinosaur was one of the first to be discovered. As you see, it stood on its hind legs like a kangaroo, and in running occasionally went on those feet only, touching the ground now and then with its front feet. Footprints in slabs of sandstone, once soft wet sand, are found showing | this. The animal stood about fourteen feet from the head to the ground in the position shown in the figure. Its thigh bone was only three feet long and it was therefore only half the size, in linear measure- ment, of the Atlanto- Saurus. Fia. 143.—Two teeth of Igua- nodon mantelli of the natural size, showing the serrated margin. In Fig. 142 an attempt is made to show what the animal looked like when the skeleton was clothed with flesh and skin. The first bones and teeth of the Iguanodon were found seventy 199 EXTINCT ANIMALS years ago by a celebrated and most delightful collector and explorer of the earth’s crust, Dr. Gideon Mantell, in the strata known as the Wealden in Sussex, just below the Chalk and Greensand (see table of strata). Dr. Mantell found that the teeth, of which two are here represented of the natural size, were those of a Fic. 144.—A portion of the upper jaw of the recent lizard Iguana, showing the serrated edges of the teeth, similar to those of Iguanodon. herbivorous animal and like those of the little living lizard from South America, called the Iguana, in the fact that the broad chisel-like crown has a saw-like edge (Fig. 144). From this fact the name Iguanodon (Iguana-toothed) was given to the new fossil giant reptile. The bones found by Mantell and others were scat- tered and not in their natural position and the 200 THE IGUANODON form of the creature had to be cuessed at by fitting this and that together. But some twenty-five years ago a wonderful find was made near Brussels in a coal-mine at a village called Bernissart. The skeletons of no less than twenty-two huge Iguanodons were found com- plete, and embedded in a fairly soft clay-like rock! The authorities of the Government Museum took charge of the place and most care fully removed the rock containing the skeletons to the Museum workshops at Brussels, where the complete skeletons of seven were, with enormous difficulty and care, removed bit by bit from the rock and set up as entire skeletons in the Brussels Museum, where they may be seen. A cast of one of these seven isin our own Natural History Museum. The photograph of the skull of one of these specimens is given in Fig. 145, It shows not only the teeth in position, but in front the bony supports of a great horny beak, like that of a turtle. As you may see in the drawing of the skeleton (Fig. 142), the forefeet (or hands) were provided with five fingers, of which the thumb had a huge claw on it at least a foot long. The foot was very much like that of a bird and had only three toes, and the bones 201 EXTINCT ANIMALS of the pelvis or hip-girdle are extraordinarily like those of a bird. In fact it is now certain that reptiles similar to the Iguanodon were the stock from which birds have been derived, the front limb having become probably first a f ee . ‘ = : PRA i Fic. 145.—Photograph of the skull of an Iguanodon as dug out of the rock, showing the teeth of the lower jaw and the smooth bony supports for the horny beak of both upper and lower jaw. The specimen is three feet in length. swimming flipper or paddle, and then later an organ for beating the air and raising the creature out of the water for a brief flight. From such a beginning came the feather-bearing wing of modern birds. Fig. 146 shows the skeleton of a Dinosaur of 202 MEGALOSAURUS THE “SnINBSOl[B SS SOS) S . ‘MOpOURNST OY} JO OZIS OY} SpaTYyy-OMY Jog’ svM TeWUITUR Ol, Sow oy} ‘ANVsoulC] SNOAOATUIVO B JO UOJoTOHNS OY JO SUTMVIG— ‘OFT “OTT [o> ANG SS ENN ap) N EXTINCT ANIMALS somewhat less size but with the same kangaroo- like carriage, which was a beast of prey. It is the Megalosaurus, and had many tiger-like teeth in its jaws. It hunted down and fed upon the herbivorous Dinosaurs as lions and tigers hunt and eat antelopes and buffalo to-day. By no means all the Dinosaurs walked on their hind legs. There were enormous kinds which went on all fours. Here is the skeleton of the Brontosaurus (Fig. 147) and a sketch of its appearance in life (Fig. 148). The great Ceteo- saurus, of which the limb bones and most of the skeleton were found near Oxford, is similar to this,and Mr. Andrew Carnegie has presented to the Natural History Museum a complete re- construction of the skeleton of a closely allied Dinosaur—the Diplodocus—which was _ exca- vated in Wyoming and is now in the Carnegie Institute at Pittsburg. It is eighty feet long. Its head is very small, and a great part of the length is made up by the very long neck and the very long tail, but the body is bigger than that of the biggest elephant and the back was nearly fourteen feet from the ground. The immense profusion in which the bones of these huge creatures have been found in 204 THE BRONTOSAURUS ‘SuIdoo4s YNoyYIM yoou oy} Aopun ssoy oy} JO QUOAZ UT YTV p[Noo ueUL Vo tey pue peoy oy) jo YASUE, yeors oy} puR [[Nys oYyZ jo ozIs [[euIs AToWIOIyxO oy} 9J0N MoS ‘snainesoyuorg oyy"fo UoyI[oys poroysor AjToqoTduroo B JO SuIMBIGQU—'L FT “917 A Ue Ws oe MS x SS AN 28 Qe ue ee BS ile) N EXTINCT ANIMALS Mesozoic strata in the United States is astonish- ing; no less remarkable is the skill and success with which American naturalists—chief among whom have been Professor Marsh of Yale and Professor Cope of Philadelphia—have collected, Fic, 148. closely similar Diplodochus and Brontosaurus) in life. It has been suggested that the animal walked along the sea or river bottom keeping its head just above water. Speci- mens of over sixty feet in length have been found. fitted together and drawn every detail of more than thirty different kinds of these monsters. They have given such full evidence of the structure and build of the animals that we may with confidence accept the reconstructions of the appearance of the animals such as those 206 THE TRICERATOPS ‘SOLQDOUTYY SUTATT YSOSAB] 9YZ JO OzIS Oy JO SBM ayydea styy, ‘(Adoystp_ YeangeN Jo wmMosny UBoltoury oy} Aq ponsst Jopout B 40948) sdozes0LLy, ‘anesourqy pouroy-saayy ayy Jo oftT Ut soUBAvodde 944 Jo SuImMeIq@—'6FT oT 207 EXTINCT ANIMALS shown in Figs. 149 and 150, where the rhino- ceros-like Triceratops and the huge crested Stegosaurus are represented. Such crests and horns are bizarre and grotesque even when carried by little living lizards a few inches long, Fie. 150.—Probable appearance in life of the Jurassic Dinosaur Stegosaurus. The hind leg alone is twice as tall as a well- grown man. but it must be remembered that the Dinosaurs drawn in Figs. 149 and 150 were as big in the body as large elephants. A curious fact about these great Dinosaurs is that they had, as compared with big living reptiles such as the crocodiles, very tiny brains. 208 THE BRAIN OF DINOSAURS You will remember that the extinct mammals known as Titanotherium and Dinoceras have brains one-eighth the bulk of living mammals of the same size, such as rhinoceros and hippo- potamus. So it was with the huge extinct reptiles. In some the head itself was ridicu- lously small according to our notions of cus- tomary proportion, and even in others, such as Triceratops, where the bony and muscular parts of the head were big, as in a rhinoceros, yet the brain was incredibly small. It could have been passed all along the spinal canal in which the spinal cord lies, and was in proportion to bulk of body a tenth the size of that of a crocodile.‘ Very probably this small size of the brain of great extinct animals has to do with the fact of their ceasing to exist. Animals with bigger and ever increasing brains outdid them in the struggle for existence.” So much for the Dinosaurs, which might well occupy a complete course of lectures all to themselves. We will now turn to the Thero- morphs, which are an older group even than the Dinosaurs and flourished in the Trias period (see table of strata, p. 60). The Thero- morphs are so called because in some important 209 P EXTINCT ANIMALS parts of the structure of skull and jaw, and often also in the teeth, they resemble the mammals or Theria. They come near to a point in the history of terrestrial vertebrate beasts which is the common origin of Reptiles, Mammals and Batrachia or Amphibians (newts, salamanders and frogs). Their remains have been found in the Triassic sandstones and limestones of South Africa, of Russia, of India and of Scotland and the centre of England. One of the most striking of these is represented by a completely reconstructed skeleton from Cape Colony in the Natural His- tory Museum, photographed in Fig. 151. The skeleton is some eight feet long and looks like a gigantic pug-dog. This is the Pariasaurus, and is shown by its small teeth to have been herbivorous. From the same locality we have the Dicyno- don with two huge tusks, and the Cynognathus with a skull and set of teeth wonderfully re- calling those of a bear at first sight. Another strange crested form belonging here is the Dimetrodon from the Permian strata of ‘Texas, U.S.A. (Fig. 152). But I am now able to show you, through the 210 THE PARIASAURUS "q09J JYSIO Jno” ‘[1e} 07 yous Woy YYSUOETT ‘UMESN AIO4STFT [RANZVNY oy ur Ajoog tossojorg Aq du jos sv ‘snaneseleg Jo Uojofoys oYy yo Ydvasoqoyg—"[E] “pIy t . 211 EXTINCT ANIMALS kindness of Professor Amalitzky, of Warsaw, a set of photographs taken by him, showing the discovery and working out by him of a whole series of skeletons of these Theromorph reptiles, closely similar to those from the rocks of Cape [het oS awe OS eA eee Fie. 152.—Probable appearance in life of the Theromorph Reptile, Dimetrodon, from the Permian of Texas. As big as a large dog. Colony but belonging to a locality far removed from South Africa, namely, to the banks of the Northern Dwina near Archangel in North Russia. Professor Amalitzky has not yet finished his excavations nor published these 212 THE BANKS OF THE DWINA photographs, and it is therefore a great kindness on his part to allow me to show them here in London. First of all, we have the cliff of Permian strata on the banks of the Dwina (Fig. 153), from Fic. 153.—View of one of the dark patches in the cliffs of the river Dwina (the Northern of that name), where nodules containing the skeletons of extinct reptiles are found. which and from another similar spot the remains were extracted. At this point, where the colour is dark in the photograph, there is a peculiar “pocket” or accumulation of sandy matter with large hard nodules embedded in it. These nodules are removed and broken up for mending 219 EXTINCT ANIMALS the roads. The pocket seems to be in a fissure and of Triassic age, later, that is to say, than the Permian rocks on each side of it. However that may be, the great nodules are removed from it for road mending, and four or five years ago Professor Amalitzky on visiting the spot was astounded and delighted to find that when broken each nodule was seen to contain the skeleton or skull of a great reptile. Fig. 154 Fic. 154.—One of the nodules showing the form of the em- bedded skeleton, head to the right, tail to the left. shows such a nodule, some eight feet long, and in this specimen one can easily distinguish the skull, the four limbs and the backbone of a large animal. The Russian geologist determined to make a most thorough investigation of this wonderful deposit, and for some years now has spent a thousand pounds a year, obtained for 214 REPTILES FOUND IN NODULES the purpose through the Imperial Academy of St. Petersburg, in having the nodules dug out by the peasants after their farming work is over for the year, and in removing them to the University of Warsaw, where with the finest Fic. 155.—Peasants working on the face of the cliff near Archangel and removing nodules containing the skeletons of great reptiles. instruments and greatest care the nodules are opened and each bone removed in fragments is put together from its more or less broken parts, firmly cemented and set up in its natural position and relations as part of a complete skeleton. 215 EXTINCT ANIMALS Fig. 155 shows the peasants at work, protected by a shed from the fall of stones from above. Fig. 156 shows some of the nodules as yet unopened lying in the laboratory of the geologi- cal professor at Warsaw. Fig. 157 shows a Fre. 156.—Professor Amalitzky’s work-shop in Warsaw, showing skeleton-holding nodules ready to be broken open and others already under preparation. - number of skeletons of the huge but harmless vegetarian Pariasaurus which have been cleared out of the nodules and set up on iron supports, as more or less complete specimens. Of course it is not possible in every individual to get out 216 PARIASAURUS SKELETONS all the bones complete, especially those of the feet. Few of the individuals were complete even when originally embedded in the mud ages ago. When an animal’s body is carried away by ariver and floats in a decomposing state it tends to fall to pieces. a ale Fic. 157.—A series of siecle tons of Peas removed bit by bit from Archangel nodules and mounted as detached specimens by Professor Amalitzky. The cliff formed by the present river Dwina consists of rocks of immense, indeed of almost inconceivable, age, and existed as solid rock ages and ages before the surface of the earth had its present form. These deep-lying rocks have been brought near to the surface by bending of 257, EXTINCT ANIMALS the strata (as shown in Fig. 36, p. 52), and the cutting or cliff made by the comparatively modern river exposes them to our view and to easy excavation. The nodules are relatively to the age of the river-valley or cutting (which is probably some 150,000 thousand Fic. 158.—Photograph of a skeleton of Pariasaurus, removed from an enveloping nodule and mounted by Professor Amalitzky. years old), as much older than it is as are Roman coins older than the trench dug three hours ago which brings them to lght. HI you look at the position of the Trias and Per- mian in the table of strata you will get some idea of how immensely remote is the time when these great reptiles lived where now is Arch- 218 SPECIMENS OF PARIASAURUS angel, for whilst the thickness of a twentieth of an inch suffices to indicate the accumulations of strata since the mammoth lived in England, the Trias is a long way down the series, far below the Eocene, where the ancestral elephants of Egypt are found, far below the Chalk, and Fre. 159.—Photograph by Professor Amalitzky on a larger seale of a skull of a Pariasaurus from an Archangel nodule. older than the long Jurassic series of rocks in which the remains of the great Dinosaurs we have recently looked at, occur. In Fig. 158 one of Professor Amalitzky’s specimens of Pariasaurus is shown. There is no artificial completing of this skeleton: all that is seen is actual bone as cleaned out of a 219 EXTINCT ANIMALS nodule. Only one foot is preserved, but that of course tells us as to its fellow of the opposite side. The skull of another specimen of Paria- saurus is shown in Fig. 159. It is very remark- able that this species seems to be so closely similar to the one discovered far away in South Africa in beds of the same age. Fic. 160.—Skeleton of a huge carnivorous beast of prey-—the reptile named Inostransevia, discovered and _ photo- graphed by Professor Amalitzky of Warsaw. The skull alone is two feet in length. These Pariasaurs were about as big as well grown cattle, but not so high on the legs. In Fig. 160 we have the skeleton of another creature revealed by these nodules. It is an enormous and truly terrible carnivor, with a 220 INOSTRANSEVIA, THE CARNIVOR skull two feet long and enormous tiger-like teeth. This creature is named Inostransevia by Pro- fessor Amalitzky, and is larger than any of the carnivorous reptiles from South Africa. Speci- mens of its skull are shown in the Professor’s photographs reproduced in Figs. 161 and 162. Fic. 161.—Skull of the gigantic Theromorph Carnivorous Reptile, Inostransevia discovered by Professor Amalitzky in Northern Russia. It is allied to Lycosaurus found in Cape Colony in beds of the same age. No doubt the vegetarian herds of Pariasaurus, whose small peg-like teeth indicate clearly enough their inoffensive habits, were preyed upon by the terrible Inostransevia, as were their brethren in South Africa devoured by the Cynognathus, the Lycosaurus, the Cynodraco 221 EXTINCT ANIMALS and other carnivorous reptiles of that remote Triassic age. So we see the co-existence of blood- sucker and victim—of the destructive oppressor and the helpless oppressed—forced on our attention in these two localities, Russia and Fie. 162.—Photograph of another skull of Inostransevia. South Africa, when we study the immensely remote past of the Triassic age. We leave now these great extinct land-dwell- ing reptiles and take a glance at representatives of two extinct orders of huge aquatic creatures which must also be classified as reptiles. These are the Plesiosauria and the Ichthyosauria. Though some of them must have measured thirty feet from snout to tail, they do not equal 222 PLESIOSAURS ‘umnosnyy AdoqstpT [BIngeVN oYyy ut 0}7e9s ojoTdwWod AjTeoU B UL GUO Aq ouoq dn yos puv ‘Yysnodoqieqog aweou (ose orssBing¢ jo) Avo oy WoT 223 SARaie Pe le the great aquatic mammals of to-day, the in size whales. In Fig. 163 is shown the photograph of the skeleton of a large Plesiosaur, and in Fig. 164 EXTINCT ANIMALS is given a drawing showing how the creature appeared in life. It had a body like the hull of a submarine with four paddles attached, the fore- and the hind-legs. It had a long neck like that of a swan and an elongated head pro- Fre. 164.—Plesiosaurus as it probably appeared when alive, swimming near the surface of the water with its back showing and its neck and head raised above the surface.’ vided with powerful jaws armed with numerous pointed teeth. It probably could swim under water as well as on the surface, and when in the latter position could snap small lizards and birds from the land. The paddles have the definite structure of legs, with five toes, wrist or ankle 224 ICHTHYOSAURS and fore-arm or fore-leg and upper arm or thigh. A great number of kinds of these Plesiosaurs have been discovered, especially in the Lias rocks of the South of England, slabs containing whole skeletons being frequently obtained. They and the similarly embedded and flattened skeletons of different kinds of —Photograph of a skeleton of the large-paddled Ichthyosaurus preserved in Liassic rock. Fie. 165. Ichthyosauria may be seen in quantity on the walls of the gallery of fossil reptiles in the Natural History Museum. In Fig. 165 the flattened skeleton of an Ichthyosaurus is photographed. This particular species is remarkable for the great size of its fore-paddles. to to on ~O EXTINCT ANIMALS In Fig. 166 a drawing of an Ichthyosaurus, as it must have appeared in life, isgiven. The Ichthyosaurs are much more fish-like or rather whale-like in form than the Plesiosaurs. They were indeed singularly like the porpoises and Fic. 166.—Drawing to show the probable appearance of an Ichthyosaurus swimming beneath the surface of the sea. erampuses among living whales and stand in the same relation to land-living reptiles that the porpoises do to land-living mammals. Their fish-like appearance and fins are not primitive characters and do not indicate any closer blood- relationship to fishes than that possessed by 226 ICHTHYOSAURS other reptiles. They are the offspring of four- legged terrestrial reptiles which have become specially modified and adapted to submarine life. Like many whales they had a median fin on the back devoid of bony support. The bones of their legs have become greatly changed, much more so than those of the Plesiosaurs and form often more than five rows of nearly circular or polygonal plates fitted together as a flexible paddle. The tail is fish-like, but has the lower lobe bigger than the upper and the vertebral column bends down into the lower lobe instead of turning up into the wpper lobe as it does in fish. The details as to the fins are known from some wonderfully preserved speci- mens found in the fine hardened mud known as the lithographic slate of Solenhofen, where the soft bodies of jelly-fish, cuttle-fishes and the wings of flying reptiles also are preserved. As mentioned in the first chapter, the Ichthyo- sauria (see Fig. 2) had a ring of bony plates supporting the eye-ball (as birds also have), and these are often preserved in the fossil specimens. In Fig. 168 a view of the top of the skull of an Ichthyosaurus is given in order to show the round hole in the middle line of 227 EXTINCT ANIMALS the brain-case (on a level with the letter P). This is called the “ parietal foramen,” and is a fair-sized hole in which was lodged an eye, a Fia. 167.—Photograph of the upper surface of the skull of an Ichthyosaurus. Ona level with the letter P in the middle ~ of the skull is seen an oval pit, the ‘* parietal foramen ”’ in which was lodged the “‘ third ” or “‘ pineal ’’ eye. third eye called the pineal eye. This eye is found in some other reptiles also, and especially 228 ICHTHYOSAURS AND COPROLITES in some of the living lizards where its structure has been studied with the microscope. There is no doubt that the body filling this hole in living lizards is an eye, although it seems to have lost the power of sight in these recent forms. 21e) iut p eppoqui 9 podlOAOOST pue p se « (umoesny, Ysa oyy Jo iy, 949 Aq yuory) (‘Aapney toyFy) ‘]rey poptatp ATTenboun oy Auoq wepnsue gos AToeso[a oY} OATesSqGQ ‘“sr1oquTeZIN AA sisuayay snyopidayT st ysy ou, ‘Yst prouwy) [Issof W— 6LT ‘Ol It is known as Dipterus, and is shown Russia. It differs from Ceratodus in having 182. 12g. F strong bony scales (whence its preservation as in THE OSTEOLEPIS a fossil) and a triangular tail-fin. The true’ ‘suy ([e10qo0d a0) ;e107R] JOLIOQUB ) syoor ouo4spuRg por SuIMBIp ouTTINO—'OsST oy} JO oqo] URBIpoU oY} 9JON “puRTJOOg Jo (URTUOAZCT PIO 94} Ut punox ‘stdoyoo4sQ Ysty plouewsy youTyxe oy Jo OLA tail-fin has disappeared altogether in the living Dipterus has peculiar teeth just mud-fishes. 251 EXTINCT ANIMALS like those of Ceratodus, and its fins are similar in character to those of the latter. In the Devonian strata are found also the It grows to a length g as well as water by its gills. 3ritish Museum. ) 181.—The Australian lung-fish Ceratodus. of two feet and breathes air by its lun (Figure lent by the Trustees of the I ~ a IG.} F extraordinary fishes known as Pterichthys. They were compared by the wonderful Scotch quarry- man, Hugh Miller—who seventy years ago 22 252 THE DIPTERUS discovered them and cleaned out many speci- mens from the rocks of his native hills at Cro- (Figure lent by the , but has a more fully , found in Scotland and in It was in many respects like Ceratodus developed tail fin and other separate median fins. Trustees of the British Museum. ) Russia. Fria. 182.—The extinct Devonian Fish, Dipterus marty in Scotland—to a tortoise’s shield with a fish thrust into it. We have now gained from 253 EXTINCT ANIMALS Fic. 183.—Outline drawing of the extinct fish Pterichthys from the Devonian or Old Red Sandstone strata. A dorsal (34), ventral (35) and lateral view (36) are given. The various bony plates are numbered, The scaly body with dorsal fin and tail fin is shown. Note also the lateral leg-like anterior fins. The round orbits (4) are seen in Fig. 34 and the mouth in 35 between the plates 2and 3. (After Traquair.) the examination of a great number of specimens from Canada as well as Scotland a very detailed 254 THE STRANGE FISH, PTERICHTHYS knowledge of the curious bony plates which build up the case or “carapace” of the body Fic. 184.—Photograph of a cardboard model of Pterichthys made by Hugh Miller, the celebrated stone-mason and naturalist of Cromarty, preserved in the Natural History Museum. of Pterichthys (Fig. 183), and also of its soft scaly tail, and the two extraordinary paddles 255 EXTINCT ANIMALS or limbs which represent the anterior or breast fins of acommon fish. Hugh Miller puzzled this out with great skill and constructed a card- board model of the fish which we have still pre- served in the Natural History Museum. It will, I think, be interesting to those who have read the writings of Hugh Miller (The Testimony of the Rocks, My Schools and Schoolmasters, and other books) to see a photograph of the model of Pterichthys which he made with his own hands (Fig. 184). In the same rocks with Pterichthys occurs another very curious fish, the Coccosteus. This and Pterichthys were of small size only, about a foot long, but in Ohio in the United States the lower jaws and skulls of huge fishes allied to Coccosteus have been found, which must have been ten or twelve feet in length. The lower jaw of one of these (called Dinichthys), together with a restored outline of Coccosteus is shown in Fig. 185. Very strange and curious fishes (only a few inches long) are found in still older strata—in the oldest Devonian and the Upper Silurian. One of these is called the buckler-head or Cephalaspis (Fig. 186). Its head is of the shape 256 COCCOSTEUS AND DINICHTHYS of a saddler’s knife and the two eyes are placed near the centre. Another fish is known almost solely by the shields which covered the head or head and body, one above and the other below. 23 2 Fic. 185.—The upper figure is a restored outline of the curious Devonian fish, Coccosteus. It is about a foot and a half long. The lower figure is a photograph to the same scale of the lower jaw of a huge fish allied to Coccosteus found in the Devonian rocks of Ohio in the United States of America. Itis called Dinichthys, and must have been from ten to twelve feet long. The above jaw and nearly complete skulls are in the Natural History Museum. This is the Pteraspis (Fig. 187). The head or head-and-body shields of these fishes and those of Cephalaspis are found in immense numbers in the hard gritty ‘“‘ cornstones ’’ of Worcester- 257 S EXTINCT ANIMALS shire and Herefordshire, also in Scotland. The stone is quarried for road mending, and great quantities of specimens have been found, though no other fossils occur with these fish-heads. It used to be imagined that this rock was the deposit of a great fresh-water lake, but that is not likely, since Pteraspis heads are found with marine shells in the rocks of Galicia. The curious thing is that although occasionally a Fic. 186.—Photograph from the original specimen of Cephalas- pis lyelli, preserved in the Natural History Museum, one- third the natural size, showing the saddler’s-knife-shaped head and the scale-bearing body. tail or body of Cephalaspis covered with scales and provided with fins has been found attached to a head-shield, as in Fig. 186, yet the body or tail of Pteraspis remains unknown. The only specimen showing any trace of the hinder 258 THE SCALES OF PTERASPIS region of Pteraspis is one which I obtained when I was a boy (in 1864) at a quarry in Herefordshire, the workmen from whom I got it saying it was a fossilized fir-cone. As a Fic. 187.—Drawings of the head-shield of the fossil fish Pteraspis. A is the species Pteraspis crouchit. B is Pteraspis rostratus. C shows a view of the under surface of the fish’s head, which was protected by a peculiar oval plate (called Scaphaspis, when it was supposed to repre- sent an independent kind of fish). The probable position of the mouth in front of the oval shield is shown. (Original. ) little concession to my vanity, I have had this solitary specimen, which I gave long ago to the British Museum, photographed of the natural size (Fig. 188). It is not much to look at, but it is one of the most interesting specimens I 259 EXTINCT ANIMALS have myself had the pleasure of unearthing. The strange thing is that it is and remains unique. Fig. g. 189 is a photograph of the upper and Fie. 188.—Photograph (of the natural size) of a specimen showing parts of the upper and lower head-shields of Pteraspis crouchii, with ten rows of lozenge-shaped scales attached. This is the only specimen showing the scales of Pteraspis, and was obtained by the author at Cradley, near West Malvern, Herefordshire, in 1864, and subse- quently presented by him to the British Museum. under side of a model of the Drepanaspis, a most strange fossil fish of the same early .age, allied to Pteraspis. It is prepared from the 260 THE DREPANASPIS drawings of Professor Traquair, who has de- scribed the fish. Specimens of it in a crushed state preserved in the slate-rock of North Germany are in the Natural History Museum. ee : SS ws PR SGN S Boo ey SS YS Fic. 189.—Photographs of models of the Devonian Fish Drepanaspis, in the Natural History Museum, prepared after the drawings of Dr. Traquair. (Original.) Fishes resembling this in shape have recently been found in the Silurian strata of Lanark- shire, and they, together with the curious little fishes drawn in Figs. 190, 191, are 261 EXTINCT ANIMALS the oldest remains of fishes which have been discovered. These last two—Birkenia and Lasanius (Figs. 190, 191)—are very puzzling Fic. 190.—Outline drawing of the Silurian fish Birkenia from Scotland, described by Dr. Traquair. little creatures, with spines set in a row along the belly. It is difficult to make out back from belly or to distinguish eyes or mouth, yet they show characteristic fish tails and a scaly cover- Fie. 191.—Outline drawing of Lasanius, another genus of fish similar to that drawn in Fig. 190, and from the same locality, described by Dr. Traquair. ing of the body. These are among the most recent discoveries and come from the Upper Silurian strata of Scotland. Specimens of these are in the Natural History Museum, but the finest series are in the Edinburgh Museum, where Professor Traquair has made a special study of the most ancient fish remains, the 262 VAST EXTENT OF ANCIENT STRATA most ancient vertebrate remains, yet disinterred from the crust of the earth. Ancient, inconceivably ancient, as are these Upper Silurian rocks, there are yet immense thicknesses below them of stratified rock, con- taining fossils in which no fish remains have been discovered. We must not conclude that the very curious-looking fishes of the Upper Silurian are really the actual forefathers of all later fish and of all vertebrate life. They just happen to be preserved and dug up, but probably sof{t-bodied fishes existed then and before that time which had no bones inside and no hard scales outside, and so have left no sign, in the rocks, of their existence. The Upper Silurian strata are, as you will see by looking at the ‘Table of Strata on p. 60, just halfway down in the thicknesses of rocks, between the present river gravels above and the Cambrian beds with the oldest known fossils (certain Trilobites) below. We will revert to the Trilobites directly ; but before leaving the extinct fishes I wish to men- tion the great fossil sharks of the late Tertiaries (Miocene and Pliocene). These we know by their teeth ; enormous shark’s teeth are found which are three times the length of the teeth 263 EXTINCT ANIMALS of the biggest living sharks on record, as shown in Fig. 192. These teeth are found in beautiful preservation in Malta, in the Antwerp a a Fic. 192.—Photograph of the jaws of a large recent Shark (Carcharodon rondeletii), the largest specimen of the kind in the Natural History Museum. At a, a, right and left, is placed a single tooth of the great extinct Miocene shark for comparison. The space between the upper and lower jaw is two feet. The fossil teeth are six inches in length, and the largest in the jaw are two inches in length. sands, in Maryland, U.S.A., and in Suffolk in England. In Suffolk they occur in the same wonderful bone-bed of the Red and Coralline Crag (see Fig. 1924), from which we get the 264 Fic. 192a.—Photograph of the natural size of a tooth of the great shark, Carcharodon megalodon, from the bone-bed of the Red Crag of Felix- stowe, Suffolk. The specimen is in the author’s cabinet. It is three times the length of the largest living shark’s tooth, and the fish which bore it was probably 100 feet in length. A kind of sandstone is seen adhering to a part of the surface of the tooth, which shows that this tooth (like many others found in the Red Crag) had been embedded in an earlier sandy deposit (the Diestien sands) before it was washed into the Red Crag. 2605 EXTINCT ANIMALS teeth of mastodon, rhinoceros and tapir. It seems to be a correct conclusion that this huge shark (Carcharodon megalodon) was nearly one hundred feet in length, since its teeth were fully three times the length of an almost identi- cal recent shark (Carcharias rondeletii), which measures thirty feet in length. “Extinct animals’? include, as must be obvious at once, a vast number of smaller creatures besides the vertebrate Fishes, Am- phibians, Reptiles, Birds and Mammals. Rocks occur containing thousands, even millions, of shells of Molluscs (whelks, bivalves, etc.) crowded together in a space of a few feet. Remains of minute shrimpsare equally abundant, and whole mountains are built up of rock formed by the coral or calcareous skeleton of minute polyps resembling our sea-anemone. Many of these are very peculiar forms, unlike those now living. Others, again, are remarkable for the tact that though found in the most ancient rocks they yet closely resemble creatures still living to-day. We will now glance at a few of the more remarkable “ fossils ’’ of these lower or simpler kinds. (See the table of classes on p. 56.) 266 THE AMMONITES In the Jurassic strata and in the Greensand and Chalk wonderful coiled shells are very Fic. 193.—Ammonites (Aegoceras capricornus) from the Lower Lias of England. commonly found which have been compared by the country-folk to petrified snakes and to Fic. 193a.—The shell of the Pearly Nautilus, cut in half so as to show the air chambers in the coils of the shell. (Lent by the Trustees of the British Museum.) These are the so- the coiled horns of the ram. called Ammonites (Fig. 193), of which there 267 EXTINCT ANIMALS are a great number of different kinds, some as big as five feet in diameter. When cut across they are seen to be divided into a number of chambers internally. In fact, their structure is the same as that of the beautiful shells of the Pearly Nautilus (Fig. 1934), which to-day lives in the Indian and Pacific oceans. The chambers in the shell of the pearly nautilus contain gas Fic. 194.—The divided shell of the Pearly Nautilus, with the animal in place in the large front chamber. (Lent by the Trustees of the British Museum.) and act as a float, whilst the animal lives in the last chamber (Fig. 194). There are only some three or four species of pearly nautilus now living, and they represent a vast variety of extinct creatures which comprise not only the Ammonites but the more ancient Goniatites. Some of these extinct allies of nautilus, such as 268 ALLIES OF AMMONITES the Orthoceras, were not coiled but quite straight: others were loosely coiled, as is the Ancyloceras AN < LQ QA WN \ “SS EN « = Lill, \” Gi L, yi (ee UNET )Y Fic. 195.—The shell of Ancyloceras matheronianum, from the Neocomian (Lower Cretaceous) rocks of France. by the Trustees of the British Museum.) (Lent A similar shell is found in the Lower Greensand of the Isle of Wight. shown in Fig. 195, and others were twisted into elongated spires (Turrilites). The creature which lived in these shells was similar to a cuttle-fish (as we know from ex- 269 EXTINCT ANIMALS amination of the animal of Nautilus), and be- longed to the class Cephalopoda of the great group Mollusca. The Molluscs include, besides these, the whelks, snails, mussels, clams and oysters. Fic. 196.—Belemnites hastatus from the Oxford Clay (Jurassic). The left-hand figure represents a specimen cut in half and shows the conical cavity or phragmacone (rudimentary chambered shell). The right-hand figure is the ‘‘ thunder- bolt ”? as usually found. A celebrated fossil which is the internal shell or “ pen” of a kind of cuttle-fish is that known bythe name ‘“ Belemnite’’(Fig. 196). These fossils are called “ thunder-bolts”’ in some parts of England, where they are sufficiently common in the clay and shale to attract attention. They 270 THE BELEMNITE’S CUTTLE FISH are found only in the Jurassic and Cretaceous formations. In fine clay specimens occur showing the soft parts of the sort of cuttle-fish in which they were formed (see Fig. 197). They Fic. 197.—Restored drawing of the animal in which the *‘ Belemnite ’’ is formed. The dense pencil-like piece lies embedded near the hinder end. (From a drawing by Sir Richard Owen.) are of the same character as the “‘ cuttle-bone ” of the living cuttle-fish and the pen of the squid (Fig. 198), but are more solid and heavy. The oldest fossils which are known are found 2z7T EXTINCT ANIMALS in the Lower Cambrian rocks (see Table of Strata, p. 60), and are the remains of small marine creatures, which were, however, by no means very simple in structure. One of these is the Lingula davis (Fig. 199), from the Lingula Fic. 198.—Loligo media, a cuttle-fish or squid now living in British seas. On the left is seen the long horny “ pen,”’ which, like the Belemnite, is embedded in the animal’s back. (Lent by the Trustees of the British Museum.) flags of Wales. Only the simple oval shells are known, but they are almost exactly like the shells of a marine animal which is still found living in immense numbers on the shores of the warmer oceans. The living owners of these shells occur in great numbers burrowing in sand and 7 fe) 272 THE OLDEST FOSSILS OF ALL have a very highly complex structure and red- coloured blood. It is indeed a most remarkable fact that the remote fossil shells of the lower Fie. 199.—Lingula (Lingulella) davisii, of the natural size, embedded in the slaty rock of Port Madoc, North Wales. Cambrian strata should be identical with those of a living animal of a high rank in the scale of structure. Not only is that the case, but in all Fie. 200.—One of the most ancient Trilobites known (Cono- coryphe lyellit), from the Lower Cambrian of Nun’s Well, Wales. From a drawing by Professor Gaudry. This Trilobite is also called Conocephalites. the deposits above the Cambrian we find the shells of Lingula, so that we must conclude that Lingula has been existing in the seas of 272 7 EXTINCT ANIMALS this earth, with very little change in form, ever since the Lower Cambrian times. Another class of fossils which are equally ancient are the Trilobites (Fig. 200). These are well-marked forms with ringed or jointed bodies divided very often into three longitudinal lobes; hence the old name Trilobites. An immense number of different kinds of Trilobites are known and classified, but they ceased to exist in the Permian period (see Table of Strata, p. 60). For along time the legs of these creatures were unknown; they have only been found within the last ten years. Mr. Beecher, of the United States, discovered them in one particular kind— the Triarthrus becki (Fig. 201). Some people consider these animals to be allied to the wood- lice or other crustacean shrimp-like forms now living. But it seems most probable that they were a primitive marine group allied to the scorpions, spiders and king-crabs (the Arachnida). It is a fact of very great significance that the earliest fossils yet discovered are the remains of very highly developed animals, by no means near the beginning of animal life. It is indeed a reasonable supposition that the earliest forms of animal life must have preceded the Cambrian 274 THE LEGS OF TRILOBITES r : is aN AX AIS Aah Ax ss ree rats A | nae ZA ZZ, Z EE Zé S A 4B eet H Ee in| pet, Mig Fic. 201.—Drawing of Triarthrus becki, a Trilobite from the Silurian rocks (Ordovician) of New York, of which the legs and antennz are well preserved, although no other Trilobite has been found showing these parts. (Lent by Macmillan & Co.) 275 EXTINCT ANIMALS Trilobites and Lingula by as long a period as these latter precede the animals living to-day. Apparently the soft-bodied animals which pre- ceded the Cambrian fossils have not left any remains in the rocks below the Cambrian or their remains have been destroyed by chemical and Fic. 202.—The Desert Scorpion (Buthus australis). Drawn from a living specimen in the author’s laboratory. structural change in those most ancient deposits. The Scorpion itself (Fig. 202) isa very ancient and important animal which so far impressed the imagination of even the earliest civilized men, that they named one of the constellations after it. Some hundreds of distinct species of scorpions are known as living at the present day 276 EXTINCT SCORPIONS in various parts of the world. In the Car- boniferous strata we find fossil scorpions hardly differing at all from those now alive, and even in the Upper Silurian we find a scorpion (Fig. 203), which would be recognized at once by a child Fic. 203.—Drawing of the remains of a Scorpion (Paleophonus hunter?) from the Upper Silurian of Lesmahago, Scotland. as being a true scorpion. It, however, seems probable that whilst modern scorpions are terrestrial, and breathe air by means of lung- sacs, the Silurian scorpion was aquatic. This is indicated by its thick crab-like legs with strong pointed end-joints (Figs. 204 and 205). Besides the Silurian scorpion of undoubted 277 EXTINCT ANIMALS affinity to modern scorpions, we find in the Silurian and Devonian rocks remains of enor- mous aquatic scorpion-like creatures, sometimes Fic. 204.—Completed draw- Fic. 205.—Completed draw- ing of the Scotch Silurian ing of the Silurian Scorpion (Palewophonus Scorpion of Gothland hunteri), seen from be- (Paleophonus nuncius), low, so as to show the seen from above. attachments of the legs. four or five feet in length (Figs. 207, 208, 209). These are known as the Eurypterids (Ptery- gotus, Stylonurus, Eurypterus, etc.). They had 278 THE EURYPTERIDS six legs like the scorpion, of which the anterior carried nippers in some instances. The great a Saat 5 f \ Fic. 206.—View of the anterior part of a recent Scorpion from below, so as to show the attachments of the limbs, the genital plate (VII go), the combs (VIII p), and the lung- mouths (IX stg to XII stg). Note also the claws at the ends of the walking legs. spine at the end of the body is the representa- tive of the scorpion’s sting, whilst they agree with scorpions in the position and character of 279 EXTINCT ANIMALS the eyes and in the number of segments or rings which build up the body and the head. A very interesting animal which is still alive (but is also found in ancient rocks) connects the Fic. 207.—View from below of the anterior part of the great Silurian Scorpion-like creature, Pterygotus osiliensis (From Zittel’s Palwontology, lent by Messrs. Mac- millan. ) scorpions with the great extinct Eurypterids and also with the Trilobites. This is the King- crab (Figs. 210, 211, 212), which is not a true crab—that is to say, a member of the class 280 THE KING CRAB Crustacea—but is a sort of marine scorpion with shortened tail (though having a long sting-like spine at the end of its body)—a x Be Fic. 208.—Photograph of a restored model of Stylonurus lacoanus, from the Upper Devonian of Pennsylvania, U.S.A. Original about five feet in length. By Professor C. E. Beecher. member of the class Arachnida. Its legs, six in number (Fig. 212), are singularly like those of the scorpion, and in a great number of minute 281 EXTINCT ANIMALS details it agrees with scorpions (see Fig. 213) Fic. 3209.—Eurypterus fischeri, a marine Scorpion-like animal from the Silurian rocks of Rootzikul. Half the size of nature. (Cut lent by Macmillan’s Co., New York, from Zittel’s Paleontology.) and differs from crabs. It is the only surviving representative of the aquatic ancestors from 282 THE KING CRAB which the modern air-breathing scorpions and spiders have been developed. From amongst all the great variety of extinct Fic. 210.—Dorsal view of the King-Crab (Limulus polyphemus Linnzeus), one-fourth the size of nature. (Cut lent by Messrs. Macmillan from Parker and Haswell, T'ext-book of Zoology.) invertebrate animals, I select for our last illus- trations and descriptions a few of the beautiful stone-lilies or Pentacrini, or Encrinites as they 283 oo eee , ee ee noon ‘\ Fic. 211.—Diagram of the dorsal surface of a King-Crab, to show the head-shield carrying the central eyes (oc’) and the lateral eyes (oc), and corresponding to six segments I to VI); also the posterior shield, corresponding to twelve segments (VII to XVIII), and the terminal post- anal spine (PA), which is identical in position with the scorpion’s sting (see Figs. 210 and 213). 284 THE KING CRAB “Anus Fic. 212.—Diagram of the ventral surface of the same King- Crab, showing the six legs, the genital operculum (Op) and the branchial plates (Br. app.). Sfr, sub-frontal piece ; Cam, upper lip or camerostome ; M, mouth ; Pmst, anterior sternal plate ; Chi, the chilaria, which are the same parts as are seen in the pentagonal sternum of the Scorpion (Fig. 206, met) and in the oval plate of Pterygotus (Fig. 207, m). iS) je 2) Ou EXTINCT ANIMALS Fic. 213.—Dorsal view of the eighteen segments and post-anal spine or sting (PA) of a Scorpion’s body to compare with those of Eurypterus (Fig. 209) and of the King-Crab (Fig. 211). In each of these the head-shield corresponds to six segments, as indicated by the legs (see Fig. 206 for the legs of the Scorpion). were long ago called. They have a very inter- esting history, for they were known as fossils as 286 THE STONE LILIES long ago as the seventeenth century, many years before they were found in the living state. They are a kind of star-fish, with long delicate arms attached to a central cup or body a Rag ; a Pps 5 al Foe ee 4 ~~ ¢ > 4 ps ? Fic. 214.—Slab containing Pentacrinus hemeri. The stalks are sometimes eighteen feet in length. (Photograph lent by Dr. Bather). which is mounted on a jointed stem, which is often of great length. Several kinds are shown in the figures 214, 215, 216. The fossil remains which we find are the hard in- 287 EXTINCT ANIMALS ternal skeleton, consisting of carbonate of lime, which was a very prominent feature in their structure. It is of the same nature as the hard box-like skeleton of the sea-urchins, which, with Fig. 215.—Photograph of a block of Limestone of the Car- boniferous period from Iowa, United States of America, showing several kinds of Stone-lilies or Encrinites. They are lettered as follows: A. Rhodocrinus kirbyi, W. and Sp. B. Rhodocrinus watersianus, W. and Sp. C. Platycrinus planus, Ow. and Sh. D. Platycrinus symmetricus, W. and Sp. £. Dorycrinus immaturus, W. and Sp. F. Dicho- crinus tnornatus, W. and Sp. (Photograph lent by Dr. Bather). the star-fishes, form the great group called by naturalists the Echinoderma. In the eighteenth century a specimen of a living Encrinite or stalked star-fish was dis- covered in the deep water off Martinique in the West Indies, and was brought in a dried con- 288 THE STONE LILIES dition to Europe and described as a “‘ sea-palm- tree.” Fora long time such specimens were very rare and difficult to obtain, but now a great number have been dredged up in deep water in Fic. 216.—Encrinus fossilis of Blumenbach from rock of Jurassic age : the original ‘‘ Stone-lily.”” (Photograph lent by Dr. Bather.) different parts of the world. Still there are only a dozen or so of different kinds or species of the Encrinites still living, whereas in all the older rocks we find their remains often in great 289 U EXTINCT. ANIMALS profusion. Many hundreds of extinct kinds are known and they occur as far back as the Cambrian rocks and are wonderfully varied and abundant in the Silurian, Devonian and Car- boniferous (Fig. 215). Some of the finest are found in the Jurassic strata (Figs. 214 and 216). A very interesting discovery in regard to the Encrinites was made by a celebrated English naturalist, Vaughan Thompson, in 1836, who was an army surgeon and quartered at Cork, where he studied the marine animals of Queenstown har- bour. He found out many new and important things by watching the growth from the egg by means of the microscope of barnacles, star- fishes and sea-moss, which he kept alive in small glass vessels. Vaughan Thompson first of all discovered in the sea at Queenstown a minute Encrinite, not a third of an inch long (Fig. 217), and to this he gave the name Pentacrinus europeus. The large one from the West Indies was at that time the only other living Encrinite known, and was called Pentacrinus asteria. This was a sufficiently astonishing discovery, but more was to come. Vaughan Thompson found in the next place that the body of his 290 THE BRITISH ENCRINITE little Pentacrinus europeus grows larger and larger whilst the stalk shrivels and ceases to Fia. 217.—The living British Encrinite, the minute young of the Feather Starfish (Comatula or Antedon), greatly magnified. (Lent by the Trustees of the British Museum.) grow, so that the animal becomes detached and swims away freely by the movement of its 291 EXTINCT ANIMALS arms. It grows up to be a well-known star- fish, the feather-star or comatula (Fig. 218). At first this history was not believed, and the Royal Society of London refused to publish SS — Ss < BOS Fic. 218.—Drawing by Mr. Berjeau from an actual[specimen of the Feather Star-fish (Comatula or Antedon rosacea), showing the ten “pinnate ’”’ or feather-like arms rising from the edge of the disc or central body, and the small grasping ‘‘ cirrhi’’ by which the animal is clinging to a stone. Of the natural size. (Original.) Vaughan Thompson’s account of what he had seen. But it was soon fully established. The little Pentacrini were bred in glass jars by many observers from the eggs of the feather- 292 YOUNG OF THE FEATHER STAR star, and all of them were seen to proceed after acertain time to produce freely-swimming little star-fish. Thus it was proved, what indeed is clear enough from the structure of its feathered arms and other parts, that the feather-star or comatula is unlike other star-fishes and is a Pentacrinus (or stone-lily or Encrinite or Crinoid) which has lost its stalk. And all the time that the naturalists of 250 years ago were disputing as to the real nature of the stone-lilies found in the rocks, little stone-lilies a quarter of an inch long were being abundantly produced every year close to hand in the sea on the rocky shores of England and France and in the Mediterranean. Whilst the first recent unfossilized Pentacrinus seen by naturalists was brought all the way from Martinique, any number of a minute size were to be found living on our ownshores. But these European Pentacrini escaped observation on account of their minute size and the sudden dwindling and loss of the stalk. Only in its very young stage does the common feather-star of to-day retain the most remarkable character- istic of its remote Cambrian ancestors, the stalk. But it does for a brief week or so, whilst almost invisible to the unaided eye, possess a well- 293 EXTINCT ANIMALS grown stalk by which it is fixed just as were its remote forefathers by their splendid waving stems many feet in length. In these pages I have only been able to bring to the reader’s notice a few of the marvellous and delightful things which we know as “ fos- sils °*>—only a very small selection of what is known about extinct animals. I have avoided going into much detail and using more technical terms and long names than is absolutely neces- sary. It is impossible to speak of these things without mentioning their names; and though it is true that a fossil “called by any other name’’ would still be full of interest, yet we must have a definite name by which to speak of each kind of animal and each kind of rock. If one’s interest is aroused in these astoundingly ancient and curious remains of extinct creatures, it becomes after all no very difficult matter to remember their names and to distinguish them from each other as well as to recognize them when we come across the names in books or museums. To learn more than the few facts which I have so briefly stated in these pages, 294 THE FOSSIL-HUNTER’S SPORT the reader should visit many times the Natural History Museum, see the actual specimens, and by the aid of the illustrated guide-books get to know more details about them. And if he or she have the chance and can go and hunt in some of the quarries or cliffs which are so often full of fossils, an endless delight and a health-giving pursuit is the prospect before him or her. Fossil-hunting with hammer and chisel and a bag to be laden with speci- mens, is splendid exercise, and, if skilfully conducted, an exciting form of sport. Even within reach of a Londoner’s day there are the brickfields of Ilford and Grays, where I used to get remains of mammoth, rhinoceros and such beasts ; there are the chalk and tertiary strata of Charlton in Kent full of fossils; the Red Crag pits of Suffolk; the oolites of Oxford ; the sponge-gravel of Farringdon. A very little longer journey brings the fossil-hunter to the Isle of Wight, which used to be, and I doubt not still is, a magnificent preserve of Eocene, Greensand and Wealden fossils ; and not further off in length of journey are the Malvern hills, with Silurian and Devonian _ strata exposed in quarries and railway cuttings, 295 EXTINCT ANIMALS teeming with fossils. And there is always the chance—a good sportsman’s chance—of finding “something new” if one understands the business and is never wearied of digging in sand and clay, and hammering the rock, and hunting up quarrymen and those delightful people—rarer now than they were forty years ago—the local naturalists. Ihope that many, if not all of my readers, may be incited by the accounts and pictures of extinct animals which I have given in these pages, to become “‘ soldiers of the hammer,” as Sir Roderick Murchison used to call us, and collectors of fossils—and if blessed by good fortune, discoverers of things as yet unknown to man. 296 INDEX A Abbeville neighbourhood, flint implements found in, 87 Advice to those interested in extinct animals, how to obtain knowledge, fossil-hunter’s sport, ete., 294-295 Aegoceris capricornus, 267 Aipyornis (moa of Madagas- car), size of egg laid by, 243 Africa : Central Africa under the equator : Five-horned giraffe— specimen shot by Sir H. Johnston, 158 Giraffe still existing, pro- tection required, 20 Okapi—skin and_ skulls discovered by Sir Harry Johnston, 161, 163 Ethiopian region—zoologi- cal province, 63, 65 South Africa : Giraffe extinct, 20 Lycosaurus — reptile found in Northern Russia allied to, 221 Pariasaurus—skull, speci- men discovered in Rus- sia similar to one dis- covered in South Africa, 220 Africa: South Africa—contd. Quagga, a native of, 18 Theromorph reptiles found in rocks of Cape Colony, 210, 211, 212 Zebra common in, 20 African elephant refer to title Elephants African square-mouthed rhi- noceros, 144 Skull compared with that of Rhinoceros antiqui- tatis, 9, 10 Age of extinct animals—great age of remains, 3, 218 Air-breathing vertebrates— Nose-passages in living and extinct crocodiles, difference in position of, 191 Aleutian Islands, sea-cow found in, 20 Amalitzky, Professor, 212, 214, 216, 29; 2211 America, South: Animal remains found in, 167 Iguana — similarity of teeth to those of the iguanodon, 200 Megatherium, 7, 9 Mylodon, discovery of remains in cave in Patagonia see Mylo- don. o7 INDEX America, South: Animals found, ete.—contd. Phororachus — gigantic extinct specimen, photograph, etc., 239, 240 Size of recent animals compared with their ‘representatives in the past, 166 Toxodon, 8, 9 Coast level, changes in, 38, 39, 40 Fishes—mud-fishes allied to the ganoid fish, 248 Neatta breed of cattle, ** bull-dogging ” of skull, 104 Neo-tropical region :—zoo- logical province, 65 American mastodon Mastodons Amiens neighbourhood, flint implements found in, ' 86, 87 Ammonites—where found, structure of — shell, animal living in the last chamber of the shell, ete., 267, 268 Animals which lived inside, similarity to cuttle- fish, 269 Extinet allies, 268, 269 Amphibia animals : Labyrinthodonts, creatures allied to, 245 see Variety found in_ the carboniferous system, 245 Anatomy—science of com- parative anatomy, 67 Ancestors : Mammals, size and descrip- tion of original “ type,” 114 Young animals, features re- sembling ancestors, which disappear on at- taining full size, 106 particular animals see their names, Horse, Elephant, etc.) Anchitherium—three-toed an- cestor of the horse, 136 Ancyloceras, shell of, 269 (for Andes—height partly ac- quired by rising of coast, 38 Andrews, Dri, 123) 124s tebe 126 Animal life, earliest forms of, preceding the Cam- brian ‘Trilobites and Lingula—no remains of soft-bodied animals in the rocks, ete., 263, 275 Animal morphography, 67 Animals which are becoming extinct : Beaver, 15, 16 Giraffe, 20, 156 Tortoise, 28, 29 Wolf, 14 Anning, Miss, 6 Antiquity of man in Europe, 85-87 Antwerp, shark’s teeth found in, 264 Apteryx : Egg of the ostrich, giant moa and apteryx, size compared, 242, 243 298 INDEX Apteryx—contd. Wingless live bird found in New Zealand, 241 Aquatic creatures refer to titles, Reptiles, Scor- pions, Fishes, Shells, ete.; also names of creatures. Arachnida class, king crab member of, 281 Archeopteryx, toothed bird, 236 Berlin specimen, 236, 238 Fingers, three distinct fin- gers, 237 Form, shape, ete., with tail like lizard and _ true bird feathers, 238 Archangel, North Russia— Professor Amalitzky’s discoveries, 212-222 Argentine Republic : Glyptodon— Armadillo-like animal from Pleisto- cene, 170 Megatherium, skeleton of, found in alluvial sands, i) Toxodon, 8, 9 Armadillo-like animal, Glyp- todon—enormous _ ar- madilloes, ete., 170, 171 Armadilloes of South America: Hairy armadillo, photo- graph, 169 Size compared with repre- sentatives in the past, 166, 167 Arsinéitherium : Appearance in life, picture showing probable ap- pearance, 153 Arsinoitherium—contd. Discovery in Upper Eocene sands of Egyptian Fa- yum, 151 Horns and teeth, 154 Name, origin of, 152 Skull, 152 Ashmole, Mr. E., 27 Atlantosaurus—example of size to which some of the Dinosaurian rep- tiles attained, 197, 198, 199 Photograph of thigh-bone, ll Auk see Great Auk Australia : Fishes : Lung-fish Ceratodus, il- lustration, 252 Mud-fish found in rivers of Queensland, 248 Kinds of animals—no_ ab- original Placentals, ete., 64 Land of the Marsupials or pouched mammals: bones of gigantic crea- tures discovered, 184 (for particular animals see their names) Reptile Chlamydosaur from Queensland — photo- graph, 195 Size of recent animals com- pared with their repre- sentatives in the past, 166 B Babbage, Mr., 34 Basilosaurus, 76 Bather, Dr., 287, 288, 289 299 INDEX Bats’ wings—resemblance be- tween wings of a Ptero- dactyles and those of a bat, 232, 233 Beadnell, Mr., 152 Beaver extinct in England, still existing in Europe and America, 15, 16 Beecher, Prof. C. E., 274, 281 Belemnite’s cuttle-fish—fos- sils called ‘‘ thunder- bolts”? in parts. of England, etc., 270 Berjeau, Mr., 292 Bird-like footprints on slab of Triassic rock from Con- necticut, 54 Birds : Animals, birds constituting group of, 23, 56 Derived from reptiles, rep- tiles coming nearest to birds in structure, ete., 202, 235, 236, 239 Feather-bearing wing © of modern birds, begin- ning of, 202 Fossil remains, where found, etc., 236, 239 New Zealand—giant ‘birds see New Zealand Skull, single joint at back of, 74 Teeth, fossil remains of birds with full set of teeth like those of rep- tiles, 236, 237 Wingless birds : Loss of wings, causes, etc., 241, 243 Remains of, where found, etc., 240 Wingless birds—contd. Water-bird, Hesperornis, etc., 244 Wings : Fin-like organs, wings probably derived from, 234 Reptiles — wings of the flying reptile, Pterodac- tyle,compared with,233 (for particular birds see their names) Birkenia—oldest remains of fishes which have been discovered, 262 Bognor Rock — photograph of slab with shells em- bedded, 45 Bones : Age of remains discovered, 3 Buried remains indicating kind of animal, food, etc., 2-4 Recognition of—marks,ete., by which fragments of bone may be referred to their proper classes, 67, 72-76 (see also Skulls) Boucher de Perthes, M., 87 Brains, size of: Dinoceras and Titanothe- rium, brains much smaller than those of recent big animals, 148-151, 209 Size of, in proportion to body — tiny size of brains of Dinosaurian reptiles, probable effect on their ceasing to ex- ist, 209 300 INDEX Brontosaurus—skeleton and probable appearance in life, 204, 205, 206 Brussels—discovery of com- plete skeletons of huge TIguanodons in coal mine, 200 Buckle-head fish found in oldest Devonian strata, saddler’s knife-shaped head, etc., 256—illus- tration, 258 ‘** Bull-dogging’”’ of skull in elephants, pugs, etc., 103-105, 106 Bulls : Urus of Julius Cesar, 16, 17 Wild cattle still found in England, ancestry of, 16, 17 Burchell’s = rhinoceros — or square-mouthed Afri- can rhinoceros — see Rhinoceros Buthus Australis: Drawing of desert scorpion, 276 C Cambrian Rocks, shells of Lingula found in—fos- sil shells identical with those of a living animal of a high rank in scale of structure, 272, 273 Cape Colony, refer to Africa, South Carboniferous system, variety of amphibia found in, 245, 246 Carcharodon megalodon—tooth of the great shark, 265 Carcharodon rondeletii—jaws of large recent shark, 264 Cariama or Screamer—gigantic extinct South American Phororachus, etc., 240 Carnegie, Mr. Andrew, 204 Cattle : ** Bull-dogging ” of skull in Neatta breed, 104 Wild cattle still to be found in England, ancestry of Gs 17 Causes of extinction of ani- mals : Changes in the surface of the earth—conditions of life altered for the animals, 31 and note Development of ancestral form in different direc- tions, 29, 30 Man’s interference, 28 Caves : Engravings on ivory and bone found in, 90-92 Mylodon, remains _ dis- covered in cave in south-west Patagonia see Mylodon Cephalaspis—saddler’s knife- shaped head and scale- bearing body, 256—il- lustration, 258 Ceratodus—Australian lung- fish related to ancient extinct fishes, 248—il- lustration, 252 Ceteosaurus—remains found near Oxford, 204—pro- bable appearance in life, 206 > 301 INDEX Chalk—tilted strata at Sea- ford, Sussex, 50 Changes in the earth: Animals, effect on—change of form or extinction, 31 and note Difficulty of realizing changes, inability to think in long enough lapses of time, 13 Eating away of edge of land by sea waves, 42 Incessant and great changes, 12 Land added to the coast by the sea, 43, 44 Rising and sinking of sur- face of the land, changes in distribution of land and water, 31 Europe: Elevation of the sea- bottom, effect on distribution of land and water, 40, 41, 42 Middle Tertiary Period— map showing attempt to determine distri- bution of land and water, 42, 43 Fossils as a means of tracing former connec- tion of different land surfaces, 66, 67 Places where there is evi- dence of change in level, 38, 39 Roman remains at Puz- zuoli,with photographs showing the temple as it was and is now, 32— 38 Changes in the earth: Rising and sinking of sur- face, ete.—contd. Washing of material from surface of land by rains and rivers, 42, 43, 44 Charlton, J Ichthyosaurus—contd. Fish-like or whale-like ap- pearance, 226, 227 Head of, from Liassie rocks of Lyme Regis, 6 Large-paddled ichthyosau- rus preserved in Lias- sic rock, 225 Offspring of four-legged ter- restrial reptiles, 227 Size of, 222 Skeletons, 225, 229 Young, bringing forth alive, 231 Iguana—upper jaw showing serrated edges of teeth similar to those of the iguanodon, 200 Iguanodon : Bones and teeth, discover- ies made by Dr. Gide- on Mantell, 199, 200 Footprint, supposed, in Isle of Wight sandstone, 54 Foot like that of a bird— stock from which birds have been derived, 202 Size, shape, ete.—probable appearance in living condition, 198, 199 Skeletons : Complete skeletons dis- covered near Brussels, 201 Drawing of, 197 Skull—specimen discovered near Brussels, 201— photograph, 202 Teeth showing — serrated margin, 199 3T0 Ilford INDEX Iguanodon—contd. Similarity of teeth to those of the little South American lizard igu- ana, 200 brickfield, remains of mammoth, ete., found in, 92, 295 Illustrations : American mastodon, 101 Ammonites, 267 Ancyloceras, shell of, 269 Apteryx, ostrich and giant moa with eggs, 242 Archeopteryx, 238 Armadillo, 169 Arsinéitherium, 152, 153 Atlantosauros,thigh-bone, 11 Beavers, 15 Belemnite’ scuttle-fish speci- mens, 270, 271 Birkinia, Silurian fish, 262 Bognor rock, 45 Bones embedded in rock, from Pikermi near Athens, 2 Brain-cavity of Dinoceras, small size compared with that of the horse, ete., 150 Brontosaurus, skeleton, 205 Cephalaspis, 258 Ceteosaurus, Diplodochus and Brontosaurus, 206 Chilian coast, change in level ; alleged Spanish inscriptions on rocks, 38, 39, 40 Chlamydosaur from Queens- land, 195 Clouded tiger, teeth of, 81 Coccosteus : curious Devon- ian fish, 257 Tllustrations—contd. Coypu rat, teeth of, 82 Crocodile—fossil jaw, 82 Deer—skeleton of male of giant Irish deer, 94 Dinoceras, 148, 149 Dinosaur stegosaurus, 208 Dinosaur, Triceratops — three-horned dinosaur, 207 Dinotherium, skull of, 118 Diprotodon — skull, skele- ton, ete., 185-188 Dodo, 26, 27 Drawings by primitive men 91, 92 Drepanaspis, 261 Dromatherium, lower jaws of, ete., 189 Divinariver, Northern Rus- sia — Professor Ama- litzky’s discoveries,213 Ear of man, show spiral construction of inter- nal ear, 74-76 Elephant, mammoth, and mastodon— transverse ridges on molar teeth, 110-113, 115 Elephants : Head of African elephant, with uplifted trunk 122 Indian and African ele- phants, 97, 98 Skulls, 104, 107, 108, 109 Skulls and jaws of series of elephant ancestors, 126, 128 Tusks, specimens in Natural History Mu- seum, 99 311 INDEX Illustrations—conitd. Fayum Desert, remains of silicified trees, 124 Flint implements, 86 Footprints of animals in ancient rocks, 54, 55 Ganoid fish fossil, 250 Hard bony scales of, 247 Giraffe, 21 Five-horned giraffe, 156, 157 Teeth of lower jaw and allied animals, 159 Glyplodon—skeleton, 170 Great auk and egg, 23, 25 Horse : Hyracotherium, Eocene ancestor, 139, 140 Model of thoroughbred English horse, 133 Phenacodus, skeleton of, 141 Toes and foot of modern horse and of four-toed and three-toed ances- tors) aso: 137.138 Horse and man, skeletons compared, 70—72 Human teeth, 80 Ichthyornis—toothed bird, 237 Ichthyosaurus, 6, 225, 226, 228, 229 Iguanodon, 197, 198, 199, 200, 202 Inostransevia, skeleton and skull, 220, 221, 222 Jaw of mammal _ from Stonesfield slate, 84 Jelly fish preserved in litho- graphic limestone, 48 312 Illustrations—contd. King-crab, 281-286 Lasanius—Silurian fish, 262 Lingula, shell of, 272, 273 Lizard : Mexican horned lizard, 194 New Zealand lizard, Tua- tara, 193 Loligo media — cuttle-fish living in British seas, 22, Lyme Regis, strata of cliff at, 49, 51 Mammoth : Imaginary picture of, 96 Skeleton found frozen in Siberia, 93 Mastodons : Meritheriwm, 129, 130 Tetrabelodon angustidens, long-jawed Miocene mastodon, 116, 117, 119, 121 Megalosaurus — skeleton, - 203 Megatherium—skeleton, 7 Moa—New Zealand moa, 68, 69 Mylodon : Remains of, discovered in cave, Piece of skin of the mylodon, ete., 175, 176 Various specimens found with the remains” of the mylodon, 177-182 View from the mouth of the cave on the fiord of the Ultima Speranza in Southern Patagonia, 174 INDEX Illustrations—contd. Skeleton, 173 Nodules containing skele- tons of reptiles—Pro- fessor Amalitzky’s dis- coveries, 213-216 Occipital condyles in skulls of mammals and rep- tiles, 73, 74 Okapi, specimen of, dis- covered by Sir Harry Johnston, ete., 163, 164, 165 Osteolepis—extinct ganoid fish, 251 Pariasaurus—skeleton, 211, 218—skull, 219 Pearly nautilus, 267, 268 Phororachus, 240 Pig’s teeth, 77, 79 Plesiosaurus, 223, 224 Polypterus of the Nile, 249 Pteraspis, 259, 260 Pterichthys, 254, 255 Pterodactyles, 230, 231, 235 Puzzuoli, Roman remains at, 32, 33, 35, 36, 37 Quagga, 18 Rhinoceros : Skeleton of Rhinoceros antiquitatis, 143 Skulls of African square- mouthed — rhinoceros and of Rhinoceros anti- quitatis (found in Lon- don), 10 Stuffed specimen — of square-mouthed rhino- ceros, 144 Ripple marks preserved in Triassic strata, 53 Tllustrations—contd. Samotherium: skull, 161 Scorpions and scorpion-like creatures, 276-283, 286 Sea-cow discovered by Steller, 22 Shark—jaw and tooth of the great shark, 264, 265 Sivatherium, 160 Skeleton of animal found embedded in caleare- ous rock at Montmar- tre, Paris, 46 Skulls of monkey, primitive man, and modern man, 88, 89 Sloth, 168, 171 Stone-lilies, 287, 288, 289, 291 Tilted strata of chalk at Seaford, Sussex, 50 Titanotherium, 145, 146, 147 Tortoise of Court House, Mauritius, 29 Toxodon, 8 Trilobites from Silurian rocks of New York, 275 Urus or bull of Julius Cesar, 17 Wings—birds, bats, and pterodactyles com- pared, 233 Wings of dragon-fly and pterodactyle preserved in limestone, 47 Wolf, 14 Zebra, 19 India — remains of Thero- morphs found in, 210 elephant refer to title Elephants Indian 313 INDEX Indian or or-ental region— zoological province, 63, 65 Information concerning ex- tinct animals, sources of : Author’s advice to those seeking information, 294 Bones and teeth found in the earth, 2—4 Tradition, 1 Inostransevia — skeleton and skull of huge specimen discovered by Profes- sor Amalitzky, 220,221 Insects : Flying insects, 232, 234 Fossilized wings, preserva- tion in stratified rock, 46, 47 Trish deer—skeleton of male of giant Irish deer, 94, 95 Isle of Wight : Footprint of animal in the sandstone, 54 Fossils—where fossils are to be found, 295 J skull of monkey-man discovered in, 88 Jaws refer to Teeth Jelly fish preserved in litho- graphic limestone, 48 Johnston, Sir Harry, 156, 158, 161, 163 Julius Cesar, great urus of, 16, 17 Java bull or Ik Kangaroos—giant kangaroos: Bones of gigantic creatures found in Australia, 184 Living specimens in Aus- tralia, size of, com- pared with gigantic ex- tinct creatures of the same kind, 166 Kansas’ refer to United States of America King-crab Animal which connects the scorpions with extinct Eurypterids and Tri- lobites, 280 Diagrams of, 284, 285 Dorsal view of—illustra- tion, 283 Member of class Arachnida —scorpion-like crea- ture, 281 Only surviving representa- tive of aquatic ances- tors from which mo- dern air - breathing scorpions and spiders have been developed, 282 Segments and post - anal spine or sting of scor- pion to compare with the king-crab, 286 Kipling, Mr. Rudyard, 120, 122 Kiwi—wingless bird found in New Zealand, 241 Knowledge, imparting—logi- cal method v. exciting the desire to know, a7 314 {NDEX L Labyrinthodonts : Allied to creatures which form the class Amphi- bia, and essentially aquatic animals, 245 Size and shape of large alli- gators, 246 Lanarkshire—fishes found in Silurian strata, 261 Land, rising and sinking see title, Changes in the earth Land-dwelling reptiles refer to Reptiles, and names of reptiles Lasanius—oldest remains of fishes which have been discovered, 262 Leeds, Mr. A. N., 223 Lepidosiren — mud-fish — of South America, 248 Lepidosteus—specimen of ga- noid fish in North American lakes, 248 Lepidotus helvenis—fossil ga- noid fish—illustration, 250 Limestone — fossilized wings of insects, etc., pre- served in, 46, 47 Limestone in solution, amount carried past Kingston by the Thames every year, 44 Limulus Polyphemus—dorsal view of the king-crab, 283 Lingula, shells of, found in the Cambrian rocks, 272 Lingula—contd. Complex structure of living owners of these shells, Ces, VAP Palio Lizard : Chlamydosaur from Queens- land—photograph, 195 Flying lizards, 234 Great girdled lizard—photo- graph, 196 Jawbones found in Oolitic strata supposed at first to be those of izards, but afterwards found to belong to small mammals, 188 Mexican horned lizard or horned toad, 194 New Zealand lizard Tua- tara—photograph, 193 Local naturalists rarer now than they were forty years ago, 296 Loligo media—cuttle-fish liv- ing in British seas, 272 London—skull of rhinoceros found in Whitefriars, 9, 10 Lycosaurus — remains dis- covered in Cape Co- lony — Inostransevia allied to, 221 Lyme Regis : Ichthyosaurus head from Liassic rocks of, 6 Strata of cliff at, 48, 49, 51 M Madagascar, wingless birds found in, 240 Malta, shark’s teeth found in, 264 315 INDEX Malvern Hills—where fossils are to be found, 295 Mammals : Ancestry—size an descrip- tion of original “‘ type,” 114 Brains of ancient big mam- mals much smaller than those of recent big animals, 148-151 Classification of — tabular list of chief orders, 57 Ear, spiral construction of internal ear, 74-76 Oldest remains—fossil jaw found at Stonesfield, one of most ancient evidences of existence, 82, 84, 186, 188 Skulls provided with pair of condyles, 73 Teeth refer to title, Teeth Mammoth : Appearance in life, imagi- nary picture of, 96 Description of, 91 Drawings by prehistoric men, 90, 91 Hairy skin, 94 Remains of, found all over the Holarctic region, 91, 92, 93 Skeleton of mammoth found frozen in Siberia, 93 Teeth — transverse ridges 110, 111 Man : Prehistoric man see that title Man—contd. Skull, size of : Giant Australian Dipro- todon, size of skull compared with that of human skull, 185 Modern man compared with that of a monkey and of a primitive man, 88-90 Man and_ horse — skeletons compared, correspond- ence in details of struc- ture, 70 Mantell, Dr. Gideon, 200 Maps : Europe — elevation of the sea-bottom, effect on distribution of land and water, 40, 41, 42 Zoo-geographical map, 63— 66 Marine creatures’ refer to titles, Reptiles, Fishes, Shells, also names of creatures Marsh, Prof., 147, 206 Marsupials : Australia distinguished by, 64 Giant Australian marsupial, Diprotodon,184—skull and skeleton, 185, 186 Jawbones embedded and preserved in ancient rocks — specimen dis- covered in Stonesfield slate near Oxford, 186— 188 Mastodon-like creature found in the Miocene—Dino- therium, 117, 118 316 INDEX Mastodons : American mastodon, 101, 102, 106, 113 Skull more projecting than that of an ele- phant, 105 Survival later in America than in Europe, 102 Teeth less peculiar than those of true elephants — fewer transverse ridges, 107, 112-113, 114 100, Meritherium, Eocene (Egypt) : Description of, head, teeth, etc., 128-132 Elephant ancestry, con- nection with, 132 More primitive mastodon than any yet known, 125 Picture representing pro- bable appearance in life, 130 Palzeeomastodon, Eocene (Egypt), 126, 128 Description of—link in the series leading back from bulldog-faced ele- phants to ordinary mammals, 127 Size, 128 Skulls and jaws of series of elephant ancestors compared, 126, 127, 128 Tetrabelodon angustidens, long-jawed Miocene mastodon : Drawing representing probable appearance in life, 119 Mastodons—contd. Tetrabeloden angustidens— contd. Skeleton from Miocene strata of south of France, 115, 116 Trunk not a ‘ trunk,”’ but an elongated upper lip, 118 Tusks and _ horizontal Tnebinlie,” URE Gi, Elen, 120, 121 Mauritius : Dodo found in, 26 Giant tortoise living in Court House Garden, 28, 29 Megalosaurus : Skeleton, drawing of, 203 Teeth, tiger-like teeth, 204 Megatherium : Comparison with little liv- ing sloths of to-day, ete., 172 Photograph of skeleton, 7 Similarity to sloth, 9 Meritherium see Mastodons Mesohippus — three-toed an- cestor of the 136 Teeth, 141 Meyer, Herman von, 76 Middle Tertiary Period, see Oligocene Period Migration of Animals : Results of—Tapir found alive in Sumatra and also in Central Amer- ica, 66 Zoo-geographical map, 63— 66 Miller, Hugh, 252-256 horse, 317 INDEX Moa: Eggs of apteryx, ostrich and the giant moa— size compared, 242, 243 New Zealand giant bird see New Zealand Size of the Madagascar moa, 242 Models of horses and cattle : Set of, in the Natural His- tory Museum, 133 Value of models as a record of best breeds, 134 Monkey, monkey-man, and modern man-skulls compared, 88—90 Monstrous size—giants in for- mer days, theory of, 2, 165, 166 Montmartre, Paris—skeleton of animal found in stratified rock, 46 Moreno, Dr., 175 Mud-fishes allied to the ganoid fishes —mud-fishes of Australia and South America, 248 Mule — okapi as hybrid or ‘““mule”’ between ze- bra and giraffe theory, 164, 165 Murchison, Sir R., 296 Mylodon : Date of extinction — sup- posed date, 174, 182, 183 Remains discovered in cave of the Ultima Speranza in South-west Patago- nia—fresh remains, ete. Mylodon—contd. Alive in the cave—indi- cations that the mylo- dons lived in the cave and were fed by the Indians, 178 Bones, claws, ete., of the mylodon, 178 Inhabitants of the cave: probable Indian inha- bitants, 176 Pellets of dung of the mylodon, 178—photo- graphs, 177, 181 Position of the cavern : explorers’ difficulties, 181 Skin covered with green- ish-brown hair, 174; photograph, 175, 176 Skin, hair, etc., preserva- tion of, in original state—probable expla- nation, 182 Various remains of the mylodons and of man, 177-180 Skeletons—comparison be- tween the skeletons of the mylodon and two- toed sloth, 172, 173 N Neanderthal, skulls of primi- tive men found in sand of, 90 Neo-tropical region—zoologi- cal province, 63, 65 318 INDEX New Zealand : Animals—New Zealand dis- tinguished from the rest of the world, 64 Birds—giant birds : Moa — ostrich-like bird, 240, 241 Skeleton constructed by Sir R. Owen, 69, 70 Thigh bone, from which existence of bird was inferred, 68, 70 Wingless birds found in New Zealand, 240, 241 Lizard Tua-tara — photo- graph, 193 Nile — Polypterus, specimen of ganoid fish still liv- ing in the Nile and other African rivers, 248—photograph, 249 Nodules containing skeletons of great reptiles—Pro- fessor Amalitzky’s dis- coveries, 213-216 Nordenskjold, Dr., 174 Norway — changes in coast level, 38 O Object of the book—bringing to notice a few of the marvellous and de- lightful things which are known as “ Fos- sils,”’? 294 Occipital condyles — mam- mals distinguished by, from birds and rep- tiles, 73 Okapi—animal allied to the giraffe : Equus Johnstoni — name given to the okapi by Dr. Sclater, 164 Hoofs, paired hoofs, 161, 164 Horns, paired horns, 164 Skin and skulls discovered by Sir Harry John- ston, 161, 163 Skull of a male okapi— photograph, 164 Species — smaller larger species, 163 Specimen of the okapi— photograph, 163 Striped skin on legs and haunches, 162 Girdles and bands for ornament made out of skin by natives, 163 ‘* Bandoliers ’? cut from the striped skin ; pho- tograph, 165 Teeth—crown of tooth in lower jaw divided by slit into two halves, described as bi-foliate, 159, 162 Unknown species of animal— hybrid or mule be- tween a zebra and giraffe theory, 164 Oligocene or Middle Tertiary Period — distribution of land and water in Europe, map showing attempt to determine, 42, 43 Oriental region — zoological province, 63, 65 and 319 INDEX Orthoceras—extinct allies of pearly nautilus, 268 Osteolepis — extinct ganoid fish : Beautifully preserved speci- men found in the De- vonian strata, 248 Drawing, 251 Ostrich-like bird—New Zea- land moa see New Zealand Owen, Sir R., 68, 69, 70, 184, 186, 271 Oxen : Skull of ox, photograph showing occipital con- dyles, 73 Urus of Julius Cesar, 16, ly Wild cattle still to be found in England, ancestry om 16; 17 P Paleomastodon, Eocene (Egypt), 126, 127, 128 Paleophonus hunteri—draw- ing of the remains of a scorpion from Upper Silurian of Lesmahago, 277, 278 Paleophonus nuncius—Silu- rian scorpion of Goth- land, 278 Paleotherium — __ skeleton found in caleareous rock at Montmartre, Paris—photograph, 46 Paleozoic strata—no reptile, bird, or mammal found in, 245 Pariasaurus : Nodules containing skele- tons— Professor Ama- litzky’s discoveries, 216-220 Remoteness of the time when these reptiles lived, 218 Size of the reptile, 220 Skeleton set up by Pro- fessor Seeley, 211 Skeleton and skull removed from an archangel nod- ule, 218, 219 Skull of Pariasaurus dis- covered in Russia: species similar to one discovered in South Africa, 220 Pearly nautilus—structure of shell, species now liv- ing, etc., 267, 268 Penguins — use of wings as swimming organs, 244 Pentacrini see Stone-lilies Permian strata on banks of the Dwina, North Russia — Professor Amalitzky’s dis- coveries, 212—222 Peterborough—skeleton of a Plesiosaur removed by Mr. A. N. Leeds—pho- tograph, 223 Phenacodus — five-toed an- cestor of the horse, 139, 141 Phororachus of South Amer- ica—photograph, ete., 239, 240 320 INDEX Phrynosoma orbiculare (Mexi- ean horned lizard or horned toad)—photo- graph, 194 Pithecanthropus or monkey- man—skull compared with skulls of chimpan- zee and modern man, 88—90 Placentalium terra—zoologi- eal province, 63, 64 Plesiosaurs : Extinct order of reptiles, 192 Form and shape—probable appearance in living condition, 224 Number of kinds discovered in Lias rocks of the south of England, 225 Size of, 222 Skeleton of, 223 Plymouth—changes in coast level, 38 Polypterus—specimen of gan- oid fish still living in the Nile and other African rivers, 248— photograph, 249 Prehistoric man : Antiquity of remains in Europe, 85-87 Drawing, skill in — photo- graphs of engravings upon bone and ivory, ete., 90-92 Skull compared with that of a monkey and of a modern man, 87—90 Pritchard, Mr. Hesketh, 181 Protopterus — mud-fish — of Africa, 248 Pteraspis —fish known by its shields, which covered head and body, where found, ete., 257, 258 Hinder unknown, 258 Specimens obtained by the author in Hereford- shire — unique speci- mens, etc., 259, 260 region Pterichthys — _ discoveries made by Hugh Miller from rocks of his na- tive hills at Cromarty, 252 Cardboard model made by Hugh Miller, 255 Curious bony plates, soft scaly tail, etc., 255 Outline drawing of the fish, 254 Pterodactyles—flying reptiles Different kinds of Jurassic pterodactyles — prob- able appearance in life, etc., 234, 235 Extinct order of reptiles, 192 Form, size, etc., as it ap- peared in flight, 231 Skeleton, 230 Wings : Formation of — bat-like appearance, etc., 232, 233 Preserved in sandy lime- stone of Oolitic Age, 46, 47 Pterygotus — _ scorpion-like creature, 278, 280 Sant Y INDEX Puzzuoli or Puteoli, condition of Roman remains at: proof of changes that take place in the level of the land, 32-38 Q Quagga : Extinct, owing to country ranged over being oc- cupied by white men, 20 Photograph of specimen in Zoological Gardens in 1875, 18 South Africa, inhabitant of, 18 Queensland refer to Aus- tralia Queenstown — Encrinite dis- covered by Vaughan Thompson, 290 R Raindrops, marks preserved on rocks which were once soft sand, 53 Rains and rivers, quantity of material carried off surface of land by, 43 Raised beaches, 38, 43, 44 Rats—teeth of Coypu rat, 81, 82 Reindeer—drawings by pre- historic men, found in caves, 90, 91, 92 Reptiles : Atlantosaurus, thigh-bone of, from Jurassic rocks of UssAe Ils ise) Reptiles—contd. Birds derived from — rep- tiles coming nearest to birds in structure, ete., 235, 236, 239 Classification of — tabular list of chief orders, 58 Crocodile see that title Difference between living and extinct reptiles— separate orders made for living _ reptiles, 191 Extinct orders—disappear- ance of remains from rocks, ete., 192 Flying reptiles, 231 Groups, 190 Land-dwelling — reptiles— great extinct reptiles, 190-222 Marine reptiles—represen- tativés of extinct or- ders of huge aquatic creatures, 222 Pterodactyle skeleton pre- served in lithographic limestone, 47 Size of extinct reptiles— enormous sizes, 167, 191 Snake, fossil remains of, found in the Fayum 125 Teeth : Description of, 81 Peg-like teeth with single fangs, 81, 82, 83 (refer also to names of reptiles) INDEX Rhinoceros : Horns : Composition of, etc., 144 Creatures allied to the rhinoceros, horns of, 144, 146 Skulls compared — African square-mouthed rhino- ceros and Rhinoceros antiquitatis, 9, 10 Square-mouthed = African rhinoceros (white rhi- noceros), 144 Rhinoceros antiquitatis— woolly rhinoceros of late Pleistocene period in Europe and Siberia : Hairy coat, 143 Skeleton of, 143 Ripple marks preserved in Triassic strata, 53 Rising and sinking of surface of the land see title, Changes in the Earth Rivers and rains—amount of material washed from surface of land and carried away by, 43 Roman remains at Puzzuoli, condition of—proof of changes that take place in the level of the land, 32-38 Rootzikul—marine_ scorpion- like animal from Silu- rian rocks, 282 Russia—Theromorph reptiles, discovery and working out of skeletons near Archangelin North Rus- sia by Professor Ama- litzki,’ 210, 212-222 32 S Samos, Island of — Samo- therium, giraffe - like animal found in Mio- cene beds, 159, 160 Samotherium — giraffe - like animal : Skull—photograph, 161 Teeth—crown of tooth in lower jaw divided by slit into two halves, described as bi-foliate, 159 Saxony—Triassic rock from, showing footprints of Cheirotherium, 55 Scales of fishes see Fishes Schweinfurth, traveller, 123 Sclater, Dr., 164 Scorpions : Ancient and important ani- mal—number of dis- tinct species: extinct species, etc., 276, 277 Animal which connects scorpions with extinct Eurypterids and Tri- lobites—king-crab, 280 Desert scorpion—drawing, 276 Silurian scorpions and enor- mous aquatic scorpion- like creatures, 277—282 IXing-crab see that title Scotland—Fishes : Fishes with head and body shields found in ** corn- stones,’ 258 Miller’s, Hugh, investiga- tions relating to the Pterichthys, 252-256 3 INDEX Scotland—Fishes—contd. Recent discoveries from the Upper Silurian strata, 262 Sea Cow: Bony plates teeth, 23 - Description of, 22, 23 Discovered by Steller, 21 Fossils found in the Fayum, 125 Picture of, 22 Sirenian group, sea-cow be- longing to, 23 Skull, photograph of, 22 Seely, Professor, 211 Seychelles—tortoise becoming extinct in, 28 Sharks : Most ancient kind of fish known, 247 Probable size of the great shark—100 feet long, 266 Teeth — enormous teeth, where found, ete., 263, 264, 265 Shells and small marine ani- mals, ete. : Animals which lived inside these shells, similarity to the cuttle-fish, 269 Bognor rock with shells em- instead of bedded, photograph, 45 Coiled shells—ammonites, pearly nautilus, ete., 267 Cuttle-fish —— Belemnite’s cuttle-fish, ete., 270 Extinct allies of nautilus, 268, 269 324 Shells, ete.—contd. Lingula, shells of, found in the Cambrian rocks, PA, PAG: Mollusca group, classes in- cluded in, 270 Oldest fossils which are known — remains. of small marine creatures, Par ils YATIP4 Trilobites see that title. Vast number of smaller creatures included in ‘ Extinct Animals ’’— mountains built up of rock formed by the coral, etc., 266 Siberia—mammoth and rhi- noceros remains found in, 93, 94 Silver-scaled fish — varieties and comparatively re- cent origin, etc., 246 Sivatherium—extinct animal from India : Skull—photograph, 160 Teeth of lower jaw—crown divided by slit into two halves, described as bi- foliate, 159 Size : . Bones — giants in former days, theory of, 2 Mammals, remote ancestor not much bigger than a dog, 114 Recent animals, size of, compared with their representatives in the past— illus ons as to extinct monsters, 165, 166 INDEX Size—contd. (for particular animals see their names) Skulls : ** Bull-dogging”’ of skulls in elephants, pugs, etc., 103-105, 106 Primitive man, skull com- pared with that of a monkey and of a mo- dern man, 87—90 particular animals see their names) Sloths : Giant ground sloth, Mega- therium : Photograph of skeleton, 7 Probable appearance in life—illustration, 171 Living sloths of South America : Size compared with re- presentatives in the past, 166, 167 Mylodon and _ two-toed sloth, comparison be- tween: skeletons, etc., 172, 173 Two-toed specimens—pho- tograph, 168 Smaller creatures—vast num- ber included in “Extinct Animals ”’— mountains built up of rock formed by the coral, etc., 266 Snake : Fossil remains of, found in the Fayum, 125 Size of extinct snakes: large size, 191 (for Soft-bodied animals— no re- mains in rocks to show earliest form of animal life preceding the Cambrian Trilobites and Lingula, 263, 275 South Africa see Africa South America see America Sphenodon punctatus (New Zealand lizard, Tua- tara)—photograph,193 Spiders—surviving represen- tative of aquatic an- cestors from which modern air-breathing scorpions and spiders have been developed, 282, 283 Spiral fold on walls of intes- tine — skeleton with excrement of the ichthyosaurus, 229 Spy, Belgium—skulls of pri- mitive men found in caverns, 89 Squirrels—flying squirrels, 234 Star fish refer to Stone-lilies Stegosaurus — probable ap- pearance in life of the Jurassic Dinosaur Ste- gosaurus, 208 Steller, discoverer cow, 21, 22 Stirling, Dr., 185 Stone-lilies, or pentacrini, or encrinites : Block of limestone showing several kinds of stone- liles from Iowa, 288 British encrinite—Vaughan Thompson’s discovery, ete., 290 of sea- INDEX Stone-lilies, ete.—contd. Young of the feather- star — Vaughan Thompson’s account established, ete., 291, 292 Common feather-star of to- day—resemblance to its remote Cambrian ancestors, 293 Encrinus Fossilis of Blu- menbach from rock of Jurassic age, 289 Fossil remains, 287 Known as fossils before they were found in the living state, 286, 293 Number of, and various species, 289, 290, 293 Stalks, length of—photo- graph, etc., 287 Stonesfield, jaw of mammal found at, 82, 84, 188 Stratification of rocks : Hard and soft rock, alter- nate layers : Pictures showing strata of cliff at Lyme Regis, 48,.49, 51 Tilting of strata, 48, 49, 50, 51 Diagram showing effect of bending or undula- tion of earth’s crust, DOD Ripple marks preserved in Triassic strata, 53 Seaford, chalk at, 50 Time elapsed during form- ation of strata, esti- mate of, 61, 62 Stratified rocks : Footprints on slabs of Tri- assic rock, 53-55 Formation of stratified de- posits from material brought down from the land by rivers, 44 Fossilized remains found in: Jelly fish preserved in lithographic limestone, photograph, 48 Shells embedded in slab of Bognor rock, 45 Skeleton of animal found in calcareous rock at Montmartre, Paris, 46 Succession from simpler to more complex forms of lfe—diagram, etc., showing position in which animal remains have been found, 60—62 Wings of insects, impres- sion preserved in lime- stone, 46, 47 Ripple marks and _ rain- drops, preservation of marks, 53 Thickness of systems of strata, diagram, etc., 60, 61 Sturgeon—ganoid set of fishes sturgeon belonging to, 248 Stylonurus — scorpion - like creature, 278, 281 Succession of animal life from simpler to more com- plex forms — position in strata in which fos- silized remains have been found, 60—62 326 INDEX Suffolk : Fossil remains, 66, 295 Land swallowed up by the sea, 43 Shark’s teeth found in the bone-bed of the Red Crag at Felixstowe, 264, 265 Sussex : Bones and teeth of the iguanodon discovered by Dr. Gideon Mantell, 200 Tilted strata of chalk at Seaford, 50 a Tadpoles—young of the Laby- rinthodonts, 245 Tanqueray’s, Lord, estate, ancestry of wild cattle ata on, 17 Tapirs : Fossil remains found all over Holarctic region, 66 Migration, results of—liv- ing tapirs found at pre- sent day in Sumatra and Central America, 66 Teeth and jaws : Arsinoitherium, 154 Bi-foliate canine see sub- heading Slit Birds, fossil remains of birds with teeth, 236 Dromatherium and Dryo- lestes, lower jaws of, 189 327 Teeth and jaws—contd. Elephants : Description of 107-110 Ridges, 110-112 Fishes—Dipterus, peculiar teeth of, 251 Horse : Cheek-teeth of modern horse more complex teeth, than in ancestors, 140 Mesohippus, teeth of, 141 Upper molar tooth of a recent horse, 142 Human teeth : Distinct from all other teeth, 80 Photograph of upper and lower jaw bone, 80 Reduced in number, 79 Iguanodon—serrated mar- gin of teeth, 199, 200 Importance of, in deter- mining animal _ to which a fragment be- longs, 76 Mammals : Fossil jaw from Stones- field slate, 82, 84, 188 Modifications in teeth of mammals, 81 “* Reduced dentition,’’ 78 Typical number of teeth, 78 (see also sub-headings, names of animals) Mammoth, 110, 111 Mastodons, jaws of, 126, 127, 128, 129, 13] INDEX Teeth and jaws—contd. Pig’s teeth : Description of, number, arrangement, etc., 76, 78 Front teeth have single fang, cheek teeth two fangs, 78, 79 Photographs of, 77, 79 Standard pattern for teeth of all mammals, 76 Reptiles, teeth of, 81, 82, 83 Ridges — elephant, mam- moth and mastodon compared, 110-115 Sea-cow—bony plates in- stead of teeth, 23 Sharks — enormous teeth, where found, ete., 263, 264, 265 Slit—crown of toothin lower jaw divided by slit into two halves, described as bi-foliate — pecu- liarity of the giraffe and allied animals, 158, 159 Tusks see that title Two fangs peculiar to mam- mals, other animals only single fangs, 78 Tetrabelodon angustidens see Mastodon Texas, refer to U.S.A. Thames, river — amount of limestone, etc., carried past Kingston each year, 43 Theriogzea or land of big ani- mals—zoological pro- vince, 63, 64 Theromorph reptiles : Extinct order of reptiles, 192 Co-existence of, in the two localities, Russia and South Africa, 212, 221, 222 Older group than Dino- saurian reptiles—where remains had been dis- covered, ete., 209 Remoteness of the time when these _ reptiles lived, 218 Russia, North: Professor Amalitzky’s discover- ies, 212-222 Nodules containing skele- tons, 213-216 (for particular members of this group see their names — Pariasaurus, Dicynodon, Inostran- sevia, etc.) Thickness of each system of strata, diagram, etce., 60-62 Thigh-bone of Atlantosaurus, 1 ae Thompson, Vaughan, 290 * Thunder-bolts ’» — Belem- nite’s cuttle-fish fossils, 270 Tiger—teeth of clouded tiger, 81 Tile-fish, destruction of, owing to change in temperature of sea, near American coast, 31 note 328 INDEX Theromorph reptiles—contd. Tilting of strata, 48, 49, 50, 51 Diagram showing effect of bending or undulation of earth’s “* crust,’ 52, yes Seaford, Sussex, chalk at, 50 Time — stratified deposits, formation of — esti- mate of lapse of time, 61, 62 Titanotherium—creature al- lied to rhinoceros : Brains much smaller than those of recent big animals, 148-151, 209 Horns, 146 Skeleton, picture of, 145 Skull, Pictures of, 146. 147 Tortoise : Becoming extinct, 28 Fossil remains found in the Fayum, 125 Giant living tortoise of the Court House, Mauri- tius, 28, 29 Size of extinct tortoises— large size, 191 Toxodon, 9—picture of, 8 Tradition—information con- cerning extinct ani- mals handed down by, 1 Traquair, Prof., 260, 261, 262 Trees — fossilized remains found in sand of Fay- um Desert, 124 Triceratops : Brain, size of, 209 Drawing of, 207 Trilobites—ancient class of fossils : Animals which connect scorpions with the ex- tinct Trilobites—king- crab, 280 Number of different kinds— primitive marine group allied to the scorpions, etc., 274. Specimens in which legs and antennze are well pre- served, 275 Trunk of elephant, develop- ment of, from elon- gated upper lip of mastodon, 118—122 Tusks : Dinoceras, 148 Dinotherium, mastodon- like creature found in the Miocene, 117, 118 Elephants — Indian and African elephants com- pared — specimens of tusks in Natural His- tory Museum, 99, 100, 101 Tetrabelodon angustidens, 116, 117, 120, 121 Meritherium, 129 U Uganda—five-horned giraffe, specimen shot by Sir Harry Johnston, 158 Okap:-skin and skulls dis- covered by Sir Harry Johnston, 161, 163 INDEX United States of America: Atlantosaurus, thigh-bone of, from Jurassic rocks, ete 1 oa 198 Coccosteus found in De- vonian rocks of Ohio, 256, 217 Dimetrodon from the Per- mian strata of Texas, 210, 212 Dinoceros, skeletons found in Upper Eocene of Wyoming, 147 Didosaurian reptiles— pro- fusion in which bones have been discovered— skill and suecess of American naturalists, etc., 206 Ichthyornis, toothed bird from chalk of Kansas, 237 Mastodon remains found in bogs, etc., 102 Scorpion-like creature from Pennsylvania, 281 Shark’s teeth found in Maryland, 264 Stone-lilies from lIowa— photograph, 288 Trilobite from Silurian rocks of New York, DiAeono Urus or bull of Julius Cesar, 6s 7 Skull, picture of, 17 WwW Wales—shells of Lingula, dis- covered in Cambrian rocks, 272, 273 Warsaw—Professor Amalitz- ky’s discoveries at Archangel, workshop at Warsaw, 216 Water-birds— extinct Hesper- ornis, ete., 244 Whale-like reptiles—Ichthyo- saurus, 226 Whales : Size of, comparisons be- tween size of recent and extinct animals, 166, 223 Skull of, mistaken for that of a reptile, 76 White rhinoceros or square- mouthed African rhi- noceros see Rhinoceros Winged reptiles see Ptero- dactyles Wings : Birds see that title Insects — fossilized wings, preservation in strati- fied rock, 46, 47 Pterodactyles, flying rep- tiles—wings compared with birds and bats, PRIS Daye Bis Swimming organs, pen- guins use their wings as, 244 Winton, Mr. de, 125 Wolf—common grey wolf— extinct in England, still existing in Europe, 14, 16 Wombats—living specimens in Australias, size of, compared with gigan- tic extinct creatures of the same kind, 166 a0 INDEX Woodward, Miss, 130 Worcestershire — fishes with head and body shields found in “* cornstones,”’ 257 Wyoming, skeleton of the Diplodocus excavated at, 204 Yy Young animals—features re- sembling ancestor 5 which disappear on at- taining full size, 106 Zebra : Africa, zebra common in, 20 Okapi—hybrid or mule be- tween zebra and giraffe theory, 164 Photograph of living zebra, 19 Zonurus giganteus great gir- dled lizard — photo- graph, 196 Zoo-geographical map, 63-66 Butler & Tanner, The Selwood Printing Works, Frome, and London, 331 ‘ oa J ee ink ‘i ‘ ~ . — ; 5 —P - ¢ >< - pe . 7! 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