eat ta ah PPA eon 5 A = : Pas s ’ Y Bs. a ail GL) i: SOSH Aa? . - < » 6 2 + + s Pea a me Stee sine . 8. A. longistipitatus Boriss.5 ..... sitet acana Ciih emi Ailes INS Mpa Section 5. Lithophilus Bge. 9.9A. ihthophilus)'Kareiet Kirs\/.2 3) 2 8) Ht ens BIeBianaty grate LOMAS chronanthusiMaihopetel: ch sho) bss lie ven Win) et We Velie lee Heels MTA. OlgaciBye:jeweliti te tel et sche ooh at ey eine 5650.10.00 0.0 Section 6. Stipitella G. Grig. 12. A. Massagetovii B. Fedtsch.. . . 2.2. +--+ 2+. eee ese Section 7. Coluteocarpus Boiss. 13. A. coluteocarpus Boiss... ..- 202+ + ee ce Poqecsnciade lle 14. A. dshimensis Gontsch. ........ gaa oMe Tartelta: ters Section 8. Nueuliella Gontsch. 15, A. chinensis L. f. «2 2 se ee eee ee ee ee eee eee “ [Russian page numbers appear in the left-hand margin of the text. ] 12 13 21 22 BB 24 fe sto 7320, 7330. 7340. Section 9. Galegiformis Gohtsch. 16, A. galegiformis L, . 2.2 eee ee secre erece Section 10. Sesbane lla Bge. 17. A. badachschanicus Boriss. . . - + 0 + eee ees Section 11. Cenanthrum Koch Subsection 1. Ellipticea Gontsch. 18, A. aksuensis Bge. OB! a) fen ies oe: ey 6). 01 0 7b) DOO. 0) re 19. A. dictamnoides Gontsch.. ». . . 2 2 eo ee we eo 20. A. tecti-mundi Freyn eceeee et e& © © © © © © 23, Astrigidus: (L.) Bye.) cic) suse) sons.» ey euelre > 24. A. Grigorjevii B. Fedtsch .... eciisisl weorents 25. A. saralensis Gontsch. . . . - «oo « paras tig A fh 26. A. umbellatus Bge. . eeeee«eeeeee 6 © #@ 6 Subsection 2, Semilunaria Gontsch, 27. A. ishigensis Maxim. - . . - .. « OM LCR MOOR IOLS 6 Oa 28. A. mongholicus Bge.. . .- °°... -- 2 PM rein COG. 5 ac 29. A. membranaceus (Fisch.) Bge. . 2 2 2 20 1 ee ee we ee ww 30. A. propinquus B, Schischk.. ........- : © ellie eames 31. A. sericeocanus Gontsch, ..--.-. 2 ee Bie vias | ska coeee eheemas 5 Section 12, Komaroviolla Gontsch. Subsection 1. Dolichanthos Gontsch. 32. A. salicetoruam Kom. . - 2.2.2. - 2-2 ees sgh 35. AS alpinus be a aicet goua eiiei cou torcel ov mienets steers 34. A. subpolaris B. Schischk. et Boriss.. . . . . «© Subsection 2, Macrotropis Boriss. 35. A. mongutensis Lipsky ........----.-- 36. A. Kuschakevitschii B, Fedtsch,. ....... oie 87. A. aksaicus B. Schischk. . . 2... ... 2. ee eee Section 13, Orobella Gontsch. 38. A oroboides Horn. ...... shld Seperate coe 39. A. pseudobrachytropis Gontak Sil ton oil (eile oto uae istoma 40. A. brachytropis (Stev.) C. A.M. 2... ee eee. 41. A. Politovii Kryl e 6 ° ° e OO, eosn en 40> 0m '¢ e@ . e e e 42, A. luxurians Bge.. . 2 2 2 2 2 eo ew AO oO O66 Section 14, Vesicularia Gontsch QS. A. Atlasovil Kom. 66. seve. of on cutive eucmelvs vi 25 REG sae 7350. 7360. 7370. 7380. Section 15, Brachyearpus Boriss, 44... A.omelanostachys: Bye... ©.) e's o.0,0ue 0 0 0 0 ec 45. A. schugnanicus B. Fedtsch.. ........0.¢. 46. A, subscaposus M. Pop, ........22.000. AT Ascjagnobicus) Lipskyis.. . 6 6 i ‘6 ee ee ve 5 4 48. A, densiflorus Kar. et Kir. ....... 4.2... 49. A. Fetissovii B. Fedtsch . ....... 5 ba el eee SNe: el rele er heaton he. 50. A. Tulinovii B. Fedtsch. 2... 2.20.20 .0--0% ce ie ec SIA; “imetensis/Boriss:. «tse. «sce 6 ease 52. A. heterodontus Boriss.......2..e.. Oe een Section 16, Hemiphragmium Koch 53. A.jvaginatus Palliw. so. 3 3 ey ce Shel NGWE 54. A. trigonocarpus (Turcz.) Bge. . .. 2-0 oe ee 55. A. australis (L.) Lam... ... Saami, 56. A. pseudoaustralis Fisch. et Mey.- ........ 57. A. sarchanensis Gontsch, .........024 ou. Ac tschiensis Bye. .t .: .* o! o* o ot o tke auneles 59. A. Beketovii (Krassn.) B, Fedtsch, ........ GOA“ chorinensis Byes uty. ss sc oe ke ee we Section 17, Hemiphaca Bge. Subsection 1. Hemiphaeopsis Gontsch. 61. A. sachalinensis Bge.. . . . . . oe eine ee ere 62. A. vorsicolor Pall ........ Sol aleus vemos ten Beye ss 63. A. puberulus Ldb. ..... or i; 105 ie) oi onsets, folk uci aleyuze 64. A, olchonensis Gontsch. .... . wtixetoel® sine 65. Avymulticanlispidb: .. cs. bs 2 6 cs «ews 66. A. vallicola Gontsech. . 2. 2. 0 2 ee eee eee Gi: Asemacropterus: DG. xs! o5 6 6: eye 6 66 6s Se 6 we 68. A. rytidocarpus Ldb...... Shine women tn eee. Je : Subsection 2. Euhemiphaca Goatsch. 69. A. hemiphaca Kar. et Kir. ..-...-.. «Hone ass 70. A. Abramovii Gontsch,. ....... 3 ernys ako ecue: 71. A. kenkolensis B. Fedtsch .......0..00- dd, AStauczilikensis Golosk. -... 9. << « sue us 4 Sx Ab@laceratustieipsky) im « % % 4 -declinatus Wilds: 3 2s 4 9 2 3 3 281% Se cme wee BGM dee eh. 183 151. A. polyphyllus Bgee --.....- Ne, » © © 2 7 ee ww 163. A. andaulgensis B. Fedtsch.. . .. - «Hatem. cults 164. A, sarytavicus M. Pop. . 165. A. apiculatus Gontsch, ... . 166. A. sericeopuberulus Boriss. . - » 2 + + ce ee ew oe 167. A. Alexeenkoi Gontseh,. . 168. A. utriger Pall. e es: e 169. A. pseudoutriger Grossh,. . . 170. A. volgensis Bge. 171, A. buchtarmensis Pall. . 172, A, Henningii (Stev.) Boriss.. . 1-2-2. ee eee 173, A. chlorodontus Bge. . 175. A. Titovii Gontsch. 176. A. varzobieus Gontsch. . 177. A. Linezevskii Gontsch, . 178. A. atrovinosus M, Pop... . . 179. A. Syreitschikovii Pavl. . 180. A. substipitatus Gontsch. . A. myriophyllus Bge. . . . A. alaicus Freyn . . A. dignus Boriss. . A. alatavicus Kar. et Kir . A. Nevskii Gontsech. .. . 181. A. Kusnetzovii M. Pop... ....-. eile Act sistas 182. A. terekliensis Gontschh ...... Se eitaioe tate 2 183. A. rotundus Gontsch. ...... - ROR, of OR 184, A. macronyx Bge. . . = SACU 2 eR, BE 185. A, Lipskyi M. Pop... ....+2+++0-+-cees 186. A. nephtonensis Freyn ... - da ew Se ae 187. A. exeapusils 28 21s ee | ce el ceo oe a tote 188. A. Jarmolenkoi Gontseh,.. . . - 2.2 2 222+ > 189, A. Alexeji Gontsch, .. . oe other Wetetelite 190, A. farctissimus Lipsky ....... JS aren 191. A. subspinescens M. Pop... . - - oc) a eterna 192. A balchaschensis Sumn, .......--- BES atte 193. A, farctus Bge. Bye) Sa Sb) lor ey io once) 194. A. abbreviatus Kar. et Kir. . ae se 195. A. charguschanus Freyn rec omcrc oe 196. A. kuschkensis Boriss. . 2... 22-2 eee 5 197, A, aktauensis Gontsch.. . .....-.. Sol 6 5 198. A. angustidens Freyn et Sint... . 2.2.2. +--+ 199. A. Barrowianus Aitch. et Baker .. ......- 200 A;curvipes ‘Trauty,) a. s+ 2 « s «© * © « oe 201. A. trachycarpus Gontsch, ...-..-. © © sikenterte 202. A. subverticillatus Gontsch.. . 2. - 0+. 2 ee . A. patentivillosus Gontsch. . . . . . - 0 ee eo « . A. kaschkadarjensis Gentsch. - 2... 2. 2--- . A. adpressipilosus Gontsch.. . . . 2.2. +e ees e ® e ® e i} e e 7520 e 7530. 7540. Section 31 Macroseminm Bge, Pil LAs) PAarAGOXUS.Bge.. 6, \s, 0, «, 19s, ©, 8, 6, eka suleWlantoneauenie: wh oFalls ¢ « Section 32, CampaneJla Gontsch. 212. A. exasperatus N. Basil. .. 2... Section 33. Chronopus Bgo, 213. A. spineseens Bee. . Section 34. Astenolobium (Nevski) Gontsch. 214. A. Bobrovii B. Fedtsch.. . ..... Section 35. Aegacantha Bge. 215. A. Bornmiillerianus B, 216. A. nigrocalyx Slob. . 217. A. Antoninae G. Grig 218. A. lejosemius (Lipsky) M. Pop...... 219. A. lasiosemius Boiss. Fedtsch. ... 220. A. roschanicus B, Fedtsch. ..... 221. A. dsharfi B. Fedtsch. Oe) 4.6 Fee 46 He «o Section 36. Acanathophace Bge. 222. A. sachendi Buhse . eo) a> Ve) ee 223. A. coaretatus Trautv.. . -..2e«cececcece 224. A. Beckerianus Trautv. . . ... 225. A. euoplus Trautv. . 226. A. sangesuricus Boriss Subgenus IIL Hypoglottis Bge. e ° e e es e © ie, fe. ie) (oe) Ke! Je ° ee e ® s ee e eo ® © © e e e e 8 e OLS OO. 1e: Je: ee. fe). fe) POs): ser) .@)) 18) 1:@) fe @) e730 le) a) Ko Section 37. Eu-Hy poglottis Bge. 227. A. cicer L. 2 we 228. A. Freynii Alb. . . siicu cmc emsutedey Gusset Higioco to 60 229. A. oreades C. A.M. ..... ii ay a tie Si el eA A Sa Si alk ee HA 2305 As flaceidtis Ma B, o".) se os er oF a 2* oe SONRREL een ee UPDRS 231. A. viciifolius DC. ..... Sie Soins. co ues ARC icaae aaa 4.5 252: A. kubensis Grossh. . . > . © + + 0 6 «© s 6 6 Soo ere ee 233. A. chordorhizus Fisch. . ... ~ Sine Ladtetenteneseivemems hc . 234, A. bachmarensis Grossh. . .. . whe: eter 2h Senses oie 235. A. saganlugensis Trautv. - - - . - « oe chet ciiedtaltetes woh iene: 236.,A., supinus Bge., <. «, «, « «, «. +, « 4 % 16 oo dolelietisiis sel oltlne © 6 237. A. Klopotovskii Sosn. . 2. 2 2. ee cee te ee ew ike! es! 6 238s Ganicus Retz. <, oo se eleteVe ic} vibe teil ouetteletic ., sens 239. A. dasyglottis Fisch. . 2 - + +202 2222 ce cee cere rece ZaU wATEtbetanus (OOnth- cs, sil ci ouied s) (oo) ole. (0) o)) sire! «lo! eleie : Section 38 Stereothrix Bge. 241. A. brachypetalus Trautv. .... . ae 242. A. setosulus Gontseh. ~ eeeeseeere#eekee#eF 8 Bee 6 X1 231 232 7550. 7560. 7570. Section 39. Malacothry»x Bge. 243. A. macrostachys DC... 2. 2 eee ee eee ee eee eee vee 244. A. podocarpusC. A.M. 2 2 s+ ee eee ee ere ere e cee 245. A. Rawlinsianus Aitsch. et Baker... -..-.. eh: Gutaguioucy oh rams 246. A. aznabjurticus Grossh. - - + +--+ eee eee esr eee ee 247. A. macrourus Fisch. et Mey. - 2 2 - +++ -2 ees Suiich auton easel 248. A. Eugenii Grossh.. » 2 2 2 ee eee ee ee ee ee ee ees : 249. A. suluklensis Freyn et Sint. . ..-- +--+ 2s oy cs) Jay Seo 250. A. protractus Boriss.. . - +--+ ++ e+ ee ee eee ote hab oy om fe 251. A. elogans Bge. . - 2 + 2+ +e +e ee ee ee eee oo a4 /smnencoane 252. A. kabristanicus Grossh, ..-- +--+. +++ ++ SS ay! aio an Mey eres 253. A. megricus Grossh. . - -+.--. Kee eT ee Ome oe cle DSA A. molligh Mons, «oc 6) 0: <) sy toy ar sh epyew oll oe ee sil'enl ee ec Gc 255. A. Schischkinianus Gontsch. .... . eet Rae Me no Co 256. A. Rzaevii Grossh,- . - +--+ +--+ ++ Sy) Sy ley a hat) oh ee 257. A. Takhtadzhjanii Grossh, . - 2. +--+ s+ 2 eee eee By ghis 958° A-:albanicus:Grossh’ «<6 2 9 2 8 2 See. ea See Subgenus IV. Trimeniaeus Bge. Section 40. Cyeloglottis Bge. 259. A. contortuplicatus L. . .-..- - one, ees, A. Pape SM ees She Section 41. Ophiocarpus Bge. 260. A. ophiocarpus Benth. . . 2 0 --- 222+ eee eee ee eee e Section 42. Heterodontus Bge. 2612,As dahuricus DG) 2) «ey oy) of ep ef ok et nego” sieicle Wiemann Section 43. Aulacolobus Bge. 262. A. striatellus Pall...... fo Nenana ee le: 6: 6-6) \e) | ienlel Mote malite Section 44. Campylotrichon Gontsch. 263. A. campylotrichus Bge.. . ...-+-+--- oie ai) epi th -eeeenle Section 45. Drepanodes Bge. 264. A. ujalensis Gontsch? . 5) t,o suey 4s cu cy + Ah oe eyiencaly mere 68 Section 46. Harpilobus Bge. 265. A. campylorrhynchus Fisch. et Mey... ..- +--+ «Waite ai ie 266. A. Drobovii M. Pop. et Vved. . ... 2. 2-2 2 ee ee ee ee 2 267. A. arpilobus Kar. et Kir. eos © © © ee ee e@ e@ © © © es ew ew ew 8 268., A. reticulatus »M., B..ui\\c% 6) a oc om ob ob ob oy watel'e) Seles, omental 269:, A..corrugatus ;Bert: 2ciies o» oh a oy o> o vemeins AeMCRCCR als, ita Em elie 270. A. umnodus M. Pop. et Vved. .. 2... 2222 ee ce ecece 2712, A. bacaliensis .Bge., oo. sh oo 0: 0 or oilaille’” -fell ei ieeils, ie Mentel ole Section 47. Sewerzowia Bge. 272. A z Schmalhausenii Bge. ee «8 © © 6 6 8 6 6 fe) es 66 ee Se Ue ee 6) 6 273:, A. compositus: Payliiichie, c, so & sativev’’ sneiopenegens Anemenene 274. A. vicarius Lipsky eeeee ee >» © © © © © © © © © © © ee 8 8 XAT 287 7580. 7590. 7600. 7610. 275. 276. 277. 278. 279, >PDP>P>pP PDS p> Section 48. Thlas pidium Lipsky AS thiaspiiltpsky .¥ 0" 158 "ehh ote! ot ot cMe teh Man tie. beh Sic Section 49. Ankylotus (Stev.) Bge. AQ commixtus (Bge.t.* 90) /eh chia" oh oP ch oc? ee TOR B heals AeNinag, Gontseh.t)." cto 3) ofc 6 ot ch of oh ch a Me An PEM, APY SIG A. ankylotus( Fisch::et ‘Mey; °.*./2*.".*. Seeing Bete Wh Kh Sic A. brachymorphus !Nikif, ) 0.4.6." 4.0%. Se2ts BORD Ogi, LY. Section 50. Platyglottis Bge. fA-.Bungei, Cy Winkl: et) B.\Fedtsch: <0. 300.5 5 «6 6 se es ee Section 51. Oxyglottis Bge. «A. Sesamoides, BOIsS:, «ys: \s+. cn «<5 o» o edhe doumabeeat. A atte”. . A. filicaulis Fisch. et Mey. . . 2... 2.6. a aS (ee aes ES . A. rytilobus Bge.. . 2.2... . athe? ac ancient. Wh) BRE. - WA oxmglottis Steve 6 6 =< so 3 0 6 0 BERS Etaenmee e Ee.”. As cornu-bovis Lipsky . . « «« « ssmreel cmtabaivewen were ite. Sy OEE.” . A. cruciatus Link... ...... o = Satedh -petetigod. A 2 eae . A. sinaicus Boiss... . . 2... 2 « « ciraelawaycmeyicmeney. A, soe abe . A. tribuloides Delil. ..... ao o.e e+ e-+eee 318. A. paliurus Boriss.....-.--- Pt. Ae ae 319, A. caucasicus Pall... .-- 2-2 ees Sere 320. A. caspicus Ms Bo... -- - Sao Oe Oo ee 321. A. oltensis Grossh. « « » « «6 « « eile ie «0 5) ous 322. A. transoxanus Fisch. - ..-.--.- EES oo Peimeuie 323, A. prominens Boriss. . .. +--+ e+e ce-s Section 54. Polyanthos Boriss. 324. A. asaphes Bge.... - 5 325. A. rubens B. Fedtsch. et N. Ww Bt eee wa ycsne 326. A. bactrianus Fisch. ev bo Sere) Oo 66% ot, 6) Oka iorale hed emiemie 327. A. Hilarfae Boriss. - © Oe (Ob cen yednel ce pielkke) Je enkuiite: De Belite 328. A. Alexeenkoanus B. Fedtsch. et N. Ivan. ...... 329. A. Sommierl Freyn. . - 2-22 se eee eee 330. A. eutranscaucasicus Boiss. - - .- 1-2-2 e+e 331. A. imbricatus Boriss.. . 2. 6. «2.2.22 ee se 332. A. Voronovianus Boriss. . -- «++. 383, A. Meyerl Boiss... 6 0 6 6 os 5) 4 etal tele 334. A. lasiostylus Fisch, . 2 2... - 2-2 ee ees 335. A. maecranthoides Boriss. ... . cote OG. eabwets, iis 336. A. nuratavieus Boriss. « - 2. + 2. e+ ee ° 337. A. plumatus Boriss, © . - ++ eee 22s e-- 338. A. pterocephalus Bge. .-..-.- e 339. A. pycnanthus Boriss. 2... .- 2+ .e2es 340, A. flexilispinus Boriss, . . 341. A. stipulosus Boriss.. . ..- 20-2 eee ree 342, A. cerasocrenus Bge.. » 2-2 2 2 2 2 see eee 343, A. strictifolius Bolss.. . . .. 2. 20 ee eee 344. A. compactus Willd. .... 2... 2.222222 345. A. condensatus Ldb. . .....---- ee eae 346: A... ferox,Borisss, s\ Bic-) « «, « so pee ofits amare, & 347, A, Karjaginii Boriss.. ....-+-+-+--- 348. A., vedicus .Takht, ©. 2 «6 « © o/s sauntehi 349, A. karabagensis Bge.. . - - . 350. A. aureus Willd... ....... Section 55. Maerophyllium Boiss. 351. A. Lagowskyi Trautv, 2 2 «+6 ec eee 352. A. oleifolius DC.. ....... Subgenus VJ. Calycophysa Bge. Section 56. Alopeeiodes Gontsch. 353. A. terrae-rubrae Butkoy .... . cee ew # &@ @ Section 57, Alopeeias (Stev.) Bge. Subsection 1. Microtropi Bge. 354. A. ponticus Palh. ..... 355. A. alopecias Pall, ..... e e o e e e s e 8 e e e ° eC. 4a) sex! xe> 8) lap ciie ee e@ ee e e © @ eer (0 (6, een ie: a hep ie , fol ef eye © jotvie: + se, aiver « e a oe e-7 382 387 387 7660. 356. A. alopecurus Pal... . 7670. 7680. 7690, SST MAM MAXIMUS WIG. 5 666) 6 9 6. 2 6 eyes 6) 3 6 ee 6 whe S58 AMeramius|/Brestitus 3, 3 3. 3) 6 8 6,6, i ee te Op 0) (oh 68 010, Ko 359. A. vulpinus Willd. eeeeee Oy 0). Oimi@ie re.) (einen .e) se: ee @ ° e 360. A. trichocalyx Trautv. ... .~ D0 HOO NOROY O80 VEO MS ane 361. A. kulabensis Lipsky .........-.. Sooo oe eehrs 362. A. mogoltavicus M. Pop. ......-+e.essesee . one S63HAmGristophi Trautve: oo. oo, 0) oo ee: 0 smaiiell etnentep uence shite, sy oftte SoAmAwischachrudensis (Bye: <, <. <1 : o: 6) oi G} 6) 6) sl 0s 0) oe ms ee K-66 S65:- Aw finitimus Og esyil- ud couch ooh elena) okie Molec) «| So 8 & 6 6 366. A. sphaerocephalus Stev.. .....-+....-. ayer eons lac tatheoy ue 367. A. megalotropis C. A.M. ....... Ee Meh a 368. A. turbinatus Bge. s 5 2 ee ce et tt tw SOS nai in Oe ac$ yt Subsection 2. Bibracteolati Bge. SOSA MRegeliitmrautvite te otere vee hey elles otc) ete) 6. ey @: ws 0) ‘oh Os on 10 370. A. agameticus Lipsky. . . . 2... 002+ -+s-++s6 6 seichiskcok cious Sil MAN globiceps Bre. 7s.) or oh of oto) al ct eh 0) oh eet ° AO 10 Section 58. Eremophysa Bge. Subsection 1. Dolichantha Gontsch. Sizes Vatianae Lincz. << ss se 6 6 6 ss ep eee Se eee hee ae ears 373. A. subkahiricus Gontsch. . . . 2... 2 2-2 oe pe eames eae eepaesnae 3745A. Maximowiczil Trauty. . 0. 6 2. «2 ee tw el iels Aes. cancuas 375. A. Litvinovii Lipsky eo. le) 1@, e @;) We) erie 'e) fe. \.0) Kel (ei sce;s 1) “ele ei omen erne. Subsection 2. Micrantha Gontsch. 376. A. Lehmannianus Bge. .* . -- 2-2 +222 es ° 6 SiiwAckelifiglipsloyile. o.0) 6. > 0 eo: eie.1+ je, o's 5 OS iaiG xo 378. A. Winckleri Trautv. le! te! “e}_ ce) te “0 ‘6 eee @ © @) (©, (01 © Je) s (© ee 379. A. sphaerophysa Kar. et Kir. . - «© . -. + © See eee OO ae uae 380. A. lelophysa Bge. - .. 2. - +--+ tke) Ub oiioy eiconnct ae SS Saw MChIVENSISBDQE:g 64 cue: p<, s 2,1, + poop sine pp eyellek chs ce. 6 ous Section 59. Eremophysopsis Gontsch. 382. A. scricopetalus Trautv. .... - OE Ceo GO cme ae A Section 60. Halicaeabus Bge. 383. A. mesites Boiss. et Buhse . ......-... ea NS op 5 B84 A suhalicacabusmWams «> ob oP en cite), SACRA ae 443 Section 67. Craecina (Stev.) Bge. 410. A. leptocaulis Ldb.. . 2... -+--+2---+-+-+5e- @ oe eee ao 411. A. mailiensis B. Fedtsch.. . .-.- +: > oe ee at ot at mecely atten aan ae 412. A. managildensis B. Fedtsch . . . - +++ -+-2-- es Pe ama a rh? 17) 413. A. Petkoffii B. Fedtsch, . .. 2-2 ee ee eee ee eee eee 448 AlV4V AS austriacus Ley oat hal et loll ef) oll oo) oho ist cho oh ofl ta ot (oh oa 449 415. A. Clerceanus Iljin et H. Krasch, - - > - 2+ - 22 2 ee ee - ee 450 4720; “4 627A: sulcatus tie. Sow = 0, 1S ue) o's hae) een ai) ier rPrrp sweancellatus Byers <2. « « «6 « ee 6 me Tat LE ee eS . pseudocancellatus Grossh.. 2... .-- 2-2 - +. eee e 466. A. onobyychioides M. B. . . 2... 2.2.26, TELE RO NETS oS ORR 467. A. kadshorensis Bge.. . .....-. os a cGS Saipan. UA 468. A. lasioglottis Stev. 469. A. brevidens Freyn et Sint’ ..--.-...--2-.. SRRTSTMS 0s : 470. A. adzharicus M. Pop. 471. A, arguricus Bgen . - 2-2-2 2. 2 es DEM STIS NNER ak BES ADEA lunatus Falls. tosis soko) le uae Yee. euyal’ 3 aM el tates one ae fe eh ines 473. A. mcskhcticus Manden. 474. A. Kozlovskyi Grossh. . . 475. A. umilateralis Kar. et Kir... 2... 2.2. eee. at NISL aati Ai XVil 7780. 7790. 7800. 7810. 476. A. sarbasnensis B. Fedtsch.. . - +» 477. A. Tranzschelii Boriss. . « « e e 478. A. djilgensis Franch. . . - - = « 479, A. adsurgens Pall. . . - + - © « 480. A. austrosibiricus B. Schischk.. 481. A. inopinatus Bnriss. - . . +. + - = « Oo A482. A. marinus Boriss., «. «. <: «. « «+: +. o: ep aiteite Section 72. Hololeuce Bge. 483. A. incertus Ldb.. ....-.- Se Daas BSS o Bas ee ara 484. A. Ruprechtii Bge.. - ..~ - AONE OM SOKO Sob ouietas 485. A. hyalolepis Bge. -.-- 2-2. +s es 0s oe se ss me ae ASGe A--Overiniivbees< <-> cle) sii cies gel eo) ye a oe ibd 487. A. bicolor Lam. . ....- - ei telnet eerie enter ve o aley cation eee 488. A. schuschensis Grossh, . 2. 2-6 .2-s- etc cnaiopie. on feroune : 489. A. dzebrailicus Grossh.. . 2. 2... 0 se ee ee ee orn we hte 490. A. psoraloides Lam, ..-...... ECS anata ate Bar ee 491. A. zuvanticus Grossh.. . - . 2. - 2+ Scehae te cor een 492. A. alyssoides Lam... .. . CoC aitA, niio les Me ee eve ee 493. A. elbrusensis Boiss... . - 2. . ee ee ee ee ee ; 494. A. Sayer AS OCIS BF Ge Qeuciaomol io ol clo clllolo Nol ML NG Shy a ae ee 495. A. Schischkinii Grossh.. . . . . .« « cf ee ee Section 73. Chlorosphaerus Bge. 496. A. stenanthus Bge.. . . 2 --- 22.2 ee o of .cHingene, 497. A. dolichopodus Freyn. .-..-....2..6-. AD ar he eae Section 74. Acmothrix Bge. 498, A. fragrans Willd ...... © =e @ © aileedke Chee : 499. A. Levieri Freyn. ...... ap ap ok lap eR ses eke 2 es © Bilcatites tation kr aati 3 i 554. A. salsugineus Kar. et Kir... -- > et aie Pa eR ttc 555. A. subauriculatus Gontsch - - - © - © e ee ee ofitne i" PRR 556. A. glomeratus edb sy (eho. jelve! uel ie ue’ ee Pot OU ROLIO TO OM OME eS oc 5 557. A. brevifolius Ldb, - - 2 2 © + 2 +e ee ee ° Ales 5. eae 558. A. galactites Pall .. - 2 ee 222s ee ee > Shae” ot ta ee 559. A. scaberrimus Bge. - - -een+*e-ee- 2 o's : 560. A. dolichophyllus Pall... - - 2 ++ cess . es its Be 561. A. humilis M. B.. . - - - - > - scale B os Mor heeds 569) (Al salareticusbOrisS.i« fap

ZA. punctatus. Bye: insti) ex ye ye 16 fo se) 1s HONE UINSROVOR ts, bic. 4 B00" AS longicuspis Byes: Siieyc 0 as jc se so paht ee erriehemiabiey. SPA. 601. A. fodinarum Boiss. et Ne&. ..... ae sineMauts, dhe 602A. campylosema; Boiss. ,.; 2j.« Poi siialiesalnols walsucvdevaulisuk ahs.% 603. A. ackerbergensis Freyn. .......02s siticthanxethe: oh weer 604. A. brachyrachis M. Pop. .......- Sruborendett ea ester 3 605: A: brachycarpusyMoiB. . 2 i 4 + 0 5 < otpot Mei Se a ee 606. A. robustus Bge.. .. 2... 5 wie Se 4 sis 607. A. subrobustus Boriss. . . 2. 2... 2 ee eee eae ot 8 “ 608. A, cinereus Willd. ........0. StRSiUS ronive heeeneuta Ns cel 609. A. taschkendicus Bge. . 2... 0. ee tw wee te ws : 610. A. platyphyllus Kar. et Kir.. . oie : hn Ce ine Oli. A. ‘coelestis’Boisss)\sh.. 2 2 «= a 6 3 s cnt end ° see 612. A. elongatus Willd) ......... cama eS 613. A. hissaricus Lipsky ...... s wee VuGOR amass A ROSS 614. A. namanganicus M. Pop.. ....-....2.. bivelae . : Section 91. Myobromopsis Boriss. 615: A. EricktiiBeed. &.. . 2's & 2 oe suplyeth vovenematateenh Ss 5 Section 92. Xiphidium Bge. 616. A. variegatus Franch, ...-....--. . Ltealhoaionl ote Sieh (6 617. A. neo-Lipskyanus M. Pop. ....... oMehopeapustt «fe cite) O18); A: leucocalyx: M. Pop: 2737. 5) .0 oo eBe Silo wsedstuostiomson «ifs , 2 : 619. A. urgutinus Lipsky .....-....... BS Te BP Sion in Sees 620. A. Kudrjaschovii A. Kor.. ......2..-.-. Se teimcu ode aehe: 621. A. cisdarvasicus Gontsch. ect gl. giao Bhi a ASN ailG” ¢ sue 622. A. kabadianus Lipsky ..... - Dares Tg ee one SLE ING. PBI O23: 7A. curchobivGontschs /s % 2°20 6 Fe ty whe tte s fe abo 624. A. xiphidium Bge. -- ---- +22 --e- bet eee EP wae s 625. A. Fedtschenkoanus Lipsky. . ...-.-----+eecees.e a 626. A. ischnocarpus Gontsch.. ...-.-----s Be eee eae ae aes ue 627. A. acanthocarpus Boriss. .-......-- SMe hat a Geo TS) 2 628: A: Spryginii M2 Pop.-). 2. = <6 os + eens ous sine fe. sta = « 629. A. nigriceps M. Pop... .--.-+--+-+-, Set a eC et Sass teas 630. A. arbuscula Pall . ..... LE eR MIAES Wey Tae ome Sik. Ree 631. A. temirensis M. Pop. .... . CNM rile Mea, OB aE "5 lear 632. A. tscharynensis M. Pop... .-.-..- a yen Cece nec ie : 633. A. duanensis Saposhn. . .. . - sao et ONG SOT SAT cans 5 eae 634, A. polyceras Kar. et Kir.. 2... .- 2 2 eee suche fe steeds : 635. A. Gontscharovii Vass... ....---.-> he teem: ke IY ous 636. A. falcigerus M. Pop. . « s 2 ¢ © + 2 05 sss + 2 ele 637. A. aulieatensis M. Pop... ....-- ics. Se: woulemteuie 42 36 6 638. A. scleroxylon Bge. . - 22 --- 2+ 35 Sip ayle, chet « 639. A. Lorinserianus Freyn. ..-.-.-.--+.- - sptderithiaule, SUP ss 640. A. maverranagri M. Pop... -..-+-+.++-++-eeee-e Meo OC 641. A. lancifolius Gontsch. . . - 2 2 2 2-2 eee ee 5 Oud Dare 642) (A. exilisyAcuKor:,. Meyie 0; ©, 2) 9: in) ayiteieiieb iol site! elect eMayile, ey%er 0} Xl 7950. 643. A. subexcedens Gontsch. - « « - » « . A. transhyreanus M. Pop.. « « « © « . A. xiphidioides Fr. et Sint. . . ~ « . A. xanthoxiphidium Fr. et Sint... . argyroides G, Beck.» - - +2 2» Sachokianus Sosn,. - - + 26» ese sumbari M. Pop. - - » - «© «© « = Neilreichianus Fr. et Sint... 2-2... stenoceroides Boriss. dsharkenticus M. Pop... . - 2 ee © e+ e @ © macrotropis Bge. .- +--+ 2-2 -« amabilis M. Pop. macroceras C. A.M. .... ortholobiformis Sumn... . . angarensis Turez .....-2.+2eee-s . A. ortholobus Bge,. . . 2. 2. - e+ eee 659. A. karkarensis M. Pop... .. 660. A. tephrolobus Bge. - - . 2... 0. 661. A. compressus Ldb. . . .....-e2es 662. A. chaetolobus Bgea. . .. 2... 2-2. te 663. A. ceratoides M. B. ........ - A. polytimeticus M. Pop....... . A. stenoceras C. A. M.. . A. substenoceras Boriss. : As Karelinianas (M> Pop. 5, 2,0 jee oy,s, 35 8) ce) ee . A. artvinensis M. Pop. ..... . A. haesitabundus Lipsky ......... . A. Reverdattoanns Sumn.. .........-. ee oa” eae oean rene 671. A. corniculatus M. B. .. . ee es eee ee y subulatas Ms Bs. 6: 5S ee Mees ce eee one 6s - ucrainicus M. Pop. et M. Klok, .. . , pseudotataricus Boriss. . sogotensis Lipsky . . . « pischtovensis Gontsch.. . Michaelis Boriss. . leptopoides M. Pop... ..... ° cornutus Pall... . brachylobus DC. .... fruticosus Pall. iskanderi Lipsky ugamfcusuMibop.-). (60s, © sie ee suds uby 2g 6.43, cui KO CnC Komarovii Lipsky . Virldis Byes. . . . . eR a el es es) Oe col ts ho etm ae . mugodsharicus Bge.. ..... . Gealbatus Pall) » .. 2... . Zingeri Korsh. ae ae 689. A. pycnolobus Bgo.. . .......4. St te ee oi ore: (eras) ere one 690. A.-lonae »Palib. 60203". 4°. 48, ge ae, Se ee 691. A. pskemensis M. Pop... ...... aOR. OT Se heron ey eae 692, A. badamensis M. Pop.. ... . 693. A. Baranovii M. Pop.. ... . 694. A. fissuralis Alex. . .. 2... 695. A. sattectoichus Gontsch.. . . 7960. © “ten (eid Frere we ey te Be eo) .6) WOnens) Cemme PP PPPPP >>> 7970. 7980. o & we is sie) ee ie! eee) oe eo e 2 ee @ e @. 8 © gs Shape js) ie 7990. HN a a a ee ee XX1l1 8000. 8010. 8020. 8030. 8040. 696. A. chorgossicus Lipsky . .. ... & hediewes ics Me 697, pASikendyriyki MouPop. << 1. (6 io ic c0 isi gs do» ane ole bsiane aie : 698. A. sogdisnus Bgew . - - - 2-2 2 ees oh tyatowmes sthenwlectend A 699. A. angreni. Lipsky .... . O oo awa 00.5 cliente me 5 Oo 700. A. karataviensis Pav ...... 5 0N6 Bi ictaistichiccl amber GO As marguzaricus Lipsky, {6 \s\.s i) i) is io ee eo ole lie SOD a6 702. A. baldschuanicus M. Pop. .-...-.. ot RS ial oS putitcheiebeiay i= ° 703. A. Kossleri Trautv. ....-..-.. Sas ave ete te Setae ee Bes 704. A. pendulinus M. Pop. et B. Fedtsch.. . .. 2. - e+ ee eee : MOS VAs subtermatuss avis so yo fe se (ve oo! ie ote tonic) alineebesWettuch cohol ke 706. A. karabilicus M. Pop. . ..-..-. » ame Susala : 707. A. psilophus Schrenk. . .. . SF ouclates? tee dee Swayetssi: 5 708, A. speciosissimus Pavl.. .......-. of usleee & 5 5 709. A. tumescens M. Pop. ........ sites > ot snchtycectstrenls ahha. % WOW ACeLipschitzin Ravi ts: oi c( 0 cc se? oh on 4 0 shen oe are cues 711. A. bossuensis M. Pop. . -.-2+-+..-+-+-.-. soretodicetcry othig GMAW Ase CianthusmB een tel clts ce! chke) cli cluctvelselsckackiolech CM MbemiNNenne? aM . 713. A. dianthoides Boriss. . . 2. 2 2 0 2 eee ee te ee eee ewe 714, A. Knorringianus Boriss. . . 2... +++. ++e- by Bekeowia ° 715. A. Mokeevae M. Pop. .... 2c eee cece ec eee ° : Ml Omarcnvingatus Fallotcmcis-) 1 ©) je! s! etic! ouje® o) yet iv" ot tet lo! ee? oll el) el a! dildeeAcekustanaicuseM. (Pop...) s+)! of «| 0! she! 6) of 0! «lof 0 8 oF ot 0) of © 718. A. macropus Bge. ... . 56 Bis Ob hot lo 6 55 cille! (sie IOWA. nallescens) M; B: . «= «5 sf @ eof 5 6 6 see ls © © = 6) «\ 6 720. A. Petunnikovii Litw. ...... +--+ -2ese-s- ay detenstsbns2 eels 721. A. eremobius M. Pop. ... - Ne Oo Ge oo 6 B56 55 722, A. rariflorus Ldb. «2. - 2 2 2-2 ee eo es ° GA ol ol od Section 93. Cytisodes Bge. 723. A. cytisoides Bge. . . - © - 2 es : 54 sibeuaceittihy weighens eons 724, A. pseudocytisoides M. Pop. .---+e-+-+---+- shust eet ote syst 725. A. promontoriorum Gontsch. . . . 2. +--+ 2 ee se ee ee oe 726. A. Krassnovianus Gontsch. 2. - - 2 2 ee 2 eee ee ee Shieh che 727. A. dolichocarpus M. Pop... - +. +--+. + see eee eee ° 728. A. stenocarpus Gontsch. . - -- 22+ ++ eee ° 5h See 729. A. Nikitinae B. Fedtsch,. .-..-.-> ee eaaeeine laneice Pome uetion te oe 730. A. Georgii Gontsch, . - 2. 5-52 7 eee te ew tw Booey ie 731. A. macropetalus C. A.M. ..... ciiah op aha: ROD. BR 56 732. A. ornithorrhynchus M. Pop. -...+-+-+-+++se-+e+--s 733 A. xipholobus M. Pop. .....+.+-e-- MS th Nee seh state Ws4ee mucleosus MipRop..° <) Woldemari Juz. 3)... 5. . - sie) daimieli sic ialeiticimelee . - 821. A. dendroides Kar. et Kir... .- +--+ 2 eee, : Mes lke aha iens 822. A. kopalensis Lipsky . ..-.- 2222+ ese e Sire rer ens : Section 103. Laguropsis Bge. Subsection 1. Alabugocystis M. Pop. 823.;A-/alabugensis’ B: Fedtsch. =) <) 4) s1<) 6 ss 1s) 6) s) © ee =)! fe 824. A. subalabugensis M. Pop. . - . 2 - - 222222222 se 0 Subsection 2. Sphaerocystis M. Pop. 825. A, Krasnovii M. Pop. e e e e e e e . e e e e e e e e e e ° e e es B26 vAnailutus Byes s:Vel/els) 1/16, fe) couse) ous) we) ie XXV 849 850 8140. 8150. 8153. 827. A. ellipsoideus Ldb. . . 828. A. petropylensis Bge.. . 829. A. arkalycensis Bge. - - 831. A. transiliensis Gontsch. 832. A. kazymbeticus Saposhn. 830. A. kurtschumensis Bge. - - © » 833, A. sphaerocystis Bge. « - - 2 oo « Subsection 3, Lagurocystis M. Pop. 834. A. laguroides Pall... -..- 22s quo. Oo oo oe obs: gels tls) 2 835. A. lupulinus Pall. fe Weve) hel ube. Ke: uc) Kel) Maule © © @ Helxel (ele ey 0's 6 Ve) 8 © Subsection 4. Globicalyx M. Pop. 836. A. calycinus M. B..~. «= o 837. A. follicularis Pall... . . . 838. A. zaissanensis Sumn. ....--.-.. Rites ewe Fee suirseth a >0 ee Subsection 5, Psammocystis M. Pop. 839. A. sabuletorum Ldb. © ela 0 g.0 ete) ge mene eeeee er © © @ © © © © Subsection 6. Microcystis M. Pop. 840. A. kisyIkumi Boriss. . . . « 841. A. pulcher E, Kor. . 2 2 2» 2 es ee oo Spe,itene wihewtn Fe) 2a 842. A. inaequalifolius N. Basil. . 2. 2 ee se ee ee te ee ww ewe 843. A. Schrenkianus Fisch. et Mey. . . - 2-2-2 see ees baie 844. A. violaceus N. BasiL. ..... 5 ° a 8&3) eee 845. A. schachimardanus N. Basil. . . 2 - 2 2 2 2 ee eo e anne 846. A. subschachimardanus M. Pop.. . « - oe useipelmeuioils a6 847, A. nobilis Bge.. . 2. - 2 eo ORC ERO MON Ome Sos 848. A. pseudonobilis M. Pop... - 2 «© ees ese eee ee ee eevee 849. A. plumbeus (Nevski) Gontsch, - . « © « «Ke o,6, eho ge sebemol ie XXvl 859 860 861 861 862 865 866 866 867 867 868 PREFACE The present volume, No. XII of the ''Flora'' contains the treatment of a single genus, Astragalus, comprising 849 species. This large number does not nearly exhaust the wealth of species of this genus, the greatest in our flora. The treatment of most species of Astragalus is by N.F. Goncharov, and only a few sections have been written jointly with M.G. Popov. A most important part of the work was prepared by A.C. Borisova, while a relatively small portion of the treatment has been shared by I.T. Vasil'chenko, S.G. Gorshkova and A.A. Grossheim. The present publication is of greater interest because the systematic treatment of the genus, which had remained unchanged since the studies of A. Bunge, was considerably revised by N.F. Goncharov. A great number of new sections have been established, and radical changes have been introduced in the arrangement of the material. As opposed to earlier volumes of the ''Flora"' all illustrations in volume XII have been drawn by a single artist, S.P. Korovin. This greatly enhances the value of the treatment and facilitates the identification of Astragalus species. Most of the work concerned with the setting up of the manuscript was carried out during the siege of Leningrad by the German army. For the workers of the Botanical Institute, as indeed for all inhabitants of Leningrad, those were days of hunger, ruin and privation, but at the same time they were days of heroic struggle against the enemy. Among the many losses suffered by Russian botanists during the blockade, one of the most grievous was the death of N.F. Goncharov, the principal author of the work on Astragali. The present work being, as already indicated, largely due to N.F. Goncharov, its publication will serve as a most fitting memorial to his devoted toil. He nearly succeeded in completing the manuscript, working on it until the last hours of his life. The few items which had not been completed in Goncharov's manuscript, were supplemented by the editors with the active assistance of A.G. Borisova. The Editors XXVil Explanatory List of Abbreviations Used for Designation of Phytogeographical Regions Afr, — Africa Alt, — Altai Am,— America Amu D,— Amu Darya An,— Anadyr Ang.-Say.— Angara-Sayans Ar,-Casp, — Aralo-Caspian Arc,— Arctic Arm,— Armenia As,— Asia Atl, — Atlantic Aust, — Australia Bal.-As. Min.— Balkans-Asia Minor Balkh. — Lake Balkhash Bat, — Batum Bes, — Bessarabia Bl, — Black Sea region Casp.— Caspian Centr, — Central Cisc, — Ciscaucasia Chuk, — Chukot Coast Crim, — Crimea Dag.— Dagestan Dau, — Dauria Dnp.— Dnieper Dv.-Pech, — Dvina-Pechora Dzu.-Kash,— Dzungaria-Kashgaria Dzu,-Tarb,— Dzungaria-Tarbagatai E,— East, Eastern 5 Eur, — Europe, European Gr,— Great, Greater Ind,-Him.— India-Himalayas Ilm,— Ilmen Irt, — Irtysh Jap.-Ch, — Japan-China Kamch,— Kamchatka Kar,.-Lap.— Karelia-Lapland Kara K,— Kara Kum Kash, — Kashgaria Kop. D.— Kopet Dagh Kurd, — Kurdistan Kyz. K.— Kyzyl Kum L,— Lower Lad,-Ilm.— Ladoga-Ilmen Laz, — Lazistan Lena.-Kol, — Lena-Kolyma M.— Middle Med,— Mediterranean Mong.— Mongolia Mtn. — Mountainous N,— North, Northern; Nov. Z.— Novaya Zemlya Okh,. —Okhotsk Pam.-Al,— Pamir-Alai S.— South, Southern Sakh. — Sakhalin Scand.,— Scandinavia Sib, — Siberia Syr D.— Syr Darya T.Sh.— Tien Shan Tal, — Talysh Tob.— Tobol Transb, — Transbaikalia Transc, — Transcaucasia Transv.— Trans-Volga Turk, — Turkish Turkm, — Turkmenistan U,— Upper Uss. — Ussuri V.— Volga V.-Ka.— Volga-Kama W.— West, Westem, Yenis,— Yenisei Ze,-Bu. —Zeya-Bureya XXVili Genus 809. ASTRAGALUS*#¥ L. L. Sp. Pl. ed. 1(1753) 755. Annual or herbaceous perennial, suffruticose, rarely fruticose plants, with pubescence composed of simple and basifix or bifurcate hairs; leaves imparipinnate, rarely paripinnate, trifoliolate or simple (with a single ter- minal leaflet); stems developed or strongly reduced (plants acaulescent or nearly so). Flower in racemes, these often contracted, capitate or spicate, in the axils of bracts, often with 2 bracteoles at the calyx base; calyx cam- panulate or cylindric, often somewhat inflated or bladdery-inflated in fruit, ruptured by the pod, or not ruptured and then enclosing the pod; corolla papilionaceous, the keel obtuse or acutish; stamens diadelphous. Fruits varying in shape, fully or partially bilocular, rarely unilocular, sessile or stipitate, membranous or coriaceous, occasionally cartilaginous, sometimes bladdery-inflated, dehiscent, with non-inflexed or, more rarely, slightly inflexed valves. A holarctic genus, penetrating along mountain ranges into the Paleo- tropics and the Neotropics. Key to Sections fm Anniaisuwith a, slender taproot... 1s). fee cnet re lene tear ele el cise OS De + Perennials from a woody root crown or, more commonly, from an ab- breviated buried caudex with short divisions, suffrutescent or low- BEGLEIS COMER AetTMenls Nona mitre tera ren clei hottest GMe re ch cemcelt el ear aw teae tanta: ot atectie 3. 2. Pubescence simple (basifix) ........ Subgenus Trimeniaeus Bge** Pubeseence Consisting iol 2. —pronged Mairs’. %esie tet suet ete. Subgenus Epiglottis Boiss. (section 96. Buceras D.C.) Sep Ubescenceysimplel(basitix)! 2 Ss sce ee eek ee ot ce eke meee eee 4, Eubescencerconsisting or 2-pronged hairs. . 4 over. <1 seule rete cualte 64. » + Spiny -frutescent or frutescent plants from a distinct woody caudex, the leaf petioles and rachises at length lignified and turning into spines; flowers in short axillary few-flowered racemes; calyx cylindric or cylindric-eampanulate; not becoming inflated r ery... ae same +s 2 3. + Perennial herbs or spineless suffrutescent, rarely diminutive frutes- cent plants, devoid of distinct annual shoots; calyx becoming at length Arde Ga aeons id Ws Layee isin. eek! chet PEMA es Seteh ohdaNen,” Alacadderye. “etodtcl 6 Near IGE From a name for a leguminous plant mentioned by Dioscorides. The subgenus Trimeniaeus comprises, in our flora, 13 sections of annual astragali —sections 40 to 52. As opposed to the treatment of other subgenera, the subdivision of the subgenus Trimeniaeus into sections has been left out of this key. In due course, beginning on page 208, keys are pro- vided for direct determination of species belonging to this subgenus, while sectional subdivision is fur- nished with the actual exposition of the material. 14, 16. Ie Calyx turbinate, pointed at base... (Subgenus Tragacantha Bge.) 6. Calyx cylindric or campanulate-cylindric, not pointed at base, sacci- RON G00, Pe OA ee iC rat er Manne Caen ns - Canin scala Fane a pane. 4.0 8. Caulescent shrubs (stems from 20-30 to 150cm long); calyx-teeth ex- ceeding, equaling or shorter than the tube, but never less than 2/3 -1/2 its length; occasionally plants subcaulescent and then corolla violet, rose or red, 12-18mm long and calyx-teeth (as indicated above) rela- (any, cully iain (0) ol osama SPS nnd oer OUcen nO ONAG. Ol5 3 8G ont 'D 6.0 Do C\SMo Ob nO ic hs Nearly acaulescent plants (stems to 7-10cm long, simple, with sub- radical leaves); corolla white or whitish-yellow, 20-28 mm long; calyx-teeth (1/2-1/3) 1/4-1/5 the length of the tube............... se Bact cm ash bibs «0%, tof omhogioms: ai saan chil gate Section 55. Macrophyllium Boiss. Hlowersminwe2 iSsOGousPin thenlelat) axallisiiSesiciiliclacs sitll ek ae ae Sia cue Bel i els Omisn'opshas oA OT Mog © Section 53. Oliganthos Boriss. Flowers in (3's) 4's-15's in the leaf axils, borne on very short (1-4mm) SLOME Mattes DeCICCUS ais) telecnis| «liane ase Section 54. Polyanthos Boriss. Weav.eS IMP ae PINMACS ae Wiel a #5; age, epee IR schg Rye Cher ote sa iets oo) as Jo 3) Leaves imparipinnate in juvenile stages and paripinnate in age .. .10. Flowers subsessile, in 2's or 3's in the leaf axils.............. SOE Rm eee ee ae Section 34. Astenolobium (Nevski) Gontsch. Blowers in loose racemes borne on axillary peduncles, . ~ 373 see Swen. SNES reg call Aaron ge pmol) Pi np co Bcuasincs ox * cadens Section 6. Stipitella G. Grig. Bractsrsmally inerbaceous,, at lengthymiarcescent jarenat. 2)--ean ene Eo Ose CMM ST 8 oe: ts, veyed. Levine!" i) ike: Go y's at wt aie AOR Section 35. Aegacantha Bge. Bractsmmembranous. .o— i mm lone es peiesistenty.sssenune. cl) one Beto. 0 Co Gert lo, GE Cuas meme eeie erie Section 36. Acanthophace Bge. Standardeypubescent, osc. ce eis ase) deco dms ava eae, 0 IPAs Standardvelabwous 3) 6 5 4s 4 ee se GoPets gee sv hehe eee ee ee 15. Shrubs vested with white and black hairs, from a rather long woody caudex; flowers in compact capitate inflorescences borne on long axil- les prscneloe eo 8 6 Cee dg Mids Section 26. Bungeastrum Gontsch. PerRennialls vested exclusive liye waithiswianibe yh aie Sere) lade 13. Flowers in compact capitate spherical inflorescences borne on rather long axillary peduncles ....... Section 57. Alopecias (Stev.) Bge. Subacaulescent or caulescent plants with abbreviated loosely few -flow - ered often subradical racemes, or tall plants with well developed stems ofsthe year and dense axillary sessile intlonescences. -.. i aieeee 14. Beside the bract, calyx subtended by 2 bracteoles. Large plants, with long stems and compact axillary sessile racemes; calyx ultimately bladdeicsy—intlateds a. sa cueseabs « Section 59. Eremophysopsis Gontsch. Bracts none; inflorescences loosely few-flowered, often subradical; calyx not becoming inflated. Caulescent, shortly caulescent or sub- ACAUIESCEMEMOLANES peice oie »\s «) + Ge lle) itis aoe 44, 44, Wings exceeded by the keel; stems 20-40 (50)cm long ..... : Seat A cite osha lac Gwe alot mia) a anno Section 8. Nuculiella Pantech. + Wings longer than the keel; stems upto 50-100cm long....... 45. 45. Leaflets 11-16 pairs, 9-20 (25)mm long; pods 10-16mm long..... Ri Ae SMe ESR uns Sere iC AM. eT AY Ba) 8 Section 9. Galegiformis Gontsch. + Leaflets 4-8 pairs, 20-70mm long; pods 30-40mm long......... oh ahah as i oy I, HONRURNE. RMR rie CE Section 19. Glycyphyllus (Stev.) Bge. 46. > Wing=petal's' bilobate* or bitidiatjapexs. 2/08 "hae .@eees eke 47. 837, 1117 54. Wang petals entire or slightly emarginate)l). .atiinle iieakelueoaA. . 48. Ovary glabrous, borne on a stipe 2-4mm long, rarely 1mm long and then the ovary pubescent...... Section 16. Hemiphragmium Koch Ovary sessile or nearly so, rarely on a gynophore up to 1mm long, PAD EOUSH Menon Edmul\cMeinceeOdeireiar . «5 aeys Section 17. Hemiphaca Bge. ZOGS HVA ROUS! 6 yy siuucice oil eiilorplaiys «io ye pemtthe le: Gare Section 17.. Hemiphaca Bge. Oe Samana ate welt mtr wtih Laetoli kie dor mbar inmate. 49, Corolla 20/>22)immylonieyliyhs) ee) sites: Section 13. Orobella Gontsch. Conoblagmueh, SHOTESL | oi) < yo) ois vege geeey bb oka sy 84ley ou eps. Rigremmcd silt roapehes ge 50. Corolla 7-8mm, calyx 2.5-3mm long; if corolla 10-14 arid calyx 4- 6mm long, then keel equaling the standard or nearly so; leaflets 8- i painscandy podsionia stipe;d<6mam, longy. Gisjne se 4 bat eeeecl. Ua. Sag Vali ch tC © Lenin COR MERLEE eR TS §. Oars tame Section 12. Komaroviella Gontsch. Corolla 11-14mm, calyx 5-6mm long; leaves 4-8 pairs; pods ona stipe 0.5-1.5 mm long; keel markedly shorter than the standard. . 51. Pods subsessile (stipe absolescent); leaflets small, 3-6 mm long; bigactsylenain OMG, iy tye hfe. cep site 'aue Section 14. Vesicularia Gontsch. Pods on a slender stipe 1-1.5mm long; leaflets larger; bracts from ED SAC LO KOMI TOMG TL. bes Payrois, 6 «apie, eos. 6 Section 13. Orobella Bge. Petioles and leaf rachises becoming lignified and transformed into spines; corolla 30-35mm long, yellow. Section 33. Chronopus Bge. Petioles and leaf rachises not becoming lignified or transformed into spines, and if at length spinescent then flowers much smaller... 53. Stems developed, about 40cm long, erect; corolla red, 18-19mm long. ca le Rn i Mees a Bel BA Section 27. Erionotus Bge. (A. arianus Gontsch. ) Corolla differently colored, not red; if corolla yellowish or whitish fading red (or actually red), then longer and the plants commonly BEAMESSCONURA MMe LUT A cnc od ha, COlsp atte Leah he FORA Ss Lahey DSS IMSL 6's 54. Plants acaulescent or nearly so (very rarely with stems up to 20cm long); stipules adnate to calyx; leaves imparipinnate or verticillate - pinnate; racemes loose, commonly few-flowered, nearly radical; calyx cylindric or cylindric-campanulate; wings and keel not united with the staminal tube,...... Section 30. Myobroma (Stev.) Bge. DitferentehT OM, 2D ON Cys inp ciel cirew +sush sy sly’ tyseu saan: soueu Spastic lel euieniskisplo cule since BO), The axillary racemes sessile or subsessile, few-flowered, orflowers madieale(or nearly so)iisteaic Ge milarscoee diay Lied: mie Gb of-ebien 56. The axillary racemes borne on rather long peduncles......... 62. Wing- and keel-petals united through 1/2 their length with the staminal tube; standard linear-oblong, (30) 45-50mm long. Dwarf plants, 4-7 (15) cm tall, with very short, at length lignified, stems; leaflets 4-5 pairs, obovate or suborbicular, glabrous; flowers subradical..... Bato ch ONO: lowe] OMe ete) Ott [Okc ae Cane Omer Section 31. Macrosemium Bge. Ditlerent: fromyabOVient.. (>) <,.,.s) 2, ste, 0) meMeeR PehCied PRMD PL eth bre ke, tse ale Flowers pale violet, tinged with yellow; pods 35-50cm long and 15- 20mm broad, shortly pubescent, glabrate in maturity .......... oe Worl aeneery. Ribas hl Meera Hel ste Porys Section 24. Macrocarpon Gontsch. Flowers white or yellow; pods smaller, 30mm long and 15-18mm broad ulanatelpanmos er sr micuteuley cule picd oo) leper ton ob ieleey oh ounce cu reniies oy RESIN > 58. Flowers white (sometimes fading reddish) ................ 59): OW eESE YC IMOWiiallcuansn cueiiey cy ch Cuteuiehiedonial ay leg: opllse ob pu ofy opneg) Suieg sures va eRe tite 60. Plants with developed long erect stems; racemes axillary........ AVS Ve oke toi Samal CMa Chad oe Bm Section 22. Cartilaginella Gontsch. 62. 63. 64. 70. Acaulescent plants; flowers nearly radicalis. Vitae ALN es Belly. ba 2. OAL a eS SERRE. Section 29. Pendulina Gontsch. (A. pauper Bge. ) Leaflets large, 35-40 (50)mm long; flowers subsessile in axillary clusters of 3-5, large (35-40mm), ochroleucous; pods lanate-pannose. meee RON AS FOE, ORES! Bae Section 23. Lithoon (Nevski) Gontsch. Meatlets*smallerynceie' mm Jonge. Sirs Pa ee ee ee ee 61. Standard-limb oblong-elliptic, many times the length of the claw; flowers in subsessile apparently fasciculate racemes........... TEU W a ede Eb ied eb wa ower eRe Nawte Ramee Hee ahem Section 20. Phacodes Bge. Standard-limb broadly oblong obovate or suborbicular; flowers in short loosely few-flowered racemes . .Section 21. Christianopsis Gontsch. Leaflets 20-35 pairs; calyx campanulate, the teeth exceeding the tube; corolla 2l=24arim longe eee 4... Section 32. Campanella Gontsch. Leaflets 2-9 pairs (occasionally 15-20); calyx cylindric or cylindric - campanulate, the teeth distinctly shorter than the tube, rarely as long, and then the plants glabrous and corolla 30mm long.......... 63. Corolla yellow, 32-36mm long; calyx cylindric, 15-20mm long; bracts 7-10mm long; peduncles of racemes 7-12cm long. Hairy plants.... SP ears, AMET MRS Cal ti ccalltes Sins Section 28. Albertoregelia M. Pop. Corolla differently colored and, if yellow, not more than 20-24mm long, or attaining 30mm and then the plants glabrous or, if hairy, calyx 10mm long and peduncles 4-7cm long; bracts 5-7mm long... Zaye EDR BITRE STN IIR A We as SER Sara Be Section 29. Pendulina Gontsch. Petioles and leaf rachises becoming indurated and transformed into spines; leaves 2-foliolate. Squat, strongly branching, subspherical, Spiny shrubs; ‘calyx cylindrie-campanulatex). aan eik Rie eee Sea tie aia, fea sega. Real . bbs Section 95. Bulimiodes Bge. Petioles and leaf rachises not becoming transformed into spines or rarely) so; andi thenthe shrubs ratheritall ons 2 eee: eee 65. SUEVelolcb@e hal WGcnvene tant Kune Ueto oe heb cabs ce Or Gr oc 66. Star araw slab ROWS Hyg Cerone ay await hee NS Sa NSS gears Re Ot. All petals Ceniceouss. 29) n0k: COSA aA rhe Aa a Section 87. Tanythrix Bge. Appressed=silky hairs confined to the standard). s).).7s0- sen) -aeemeene nes bas aia etun a> debe e® se ae ab ew BhtekSy cw ee ah Seed ... Section 71. Onobrychium Bge. The large keel broader and longer than the wings. Diminutive plants up to 10cm tall, with ascending or prostrate stems; the small flowers in a loose 3-6-flowered raceme ...... Section 68. Irinaea Boriss. Keélvexcecdedsby the-wingsare 20.298 #1 28G.74. 0.04 Cn eee 68. Fruticose or suffruticose psammophytes, the aerial caudex and stems of the year well developed; flowers in loose, rather long-peduncled racemes, the calyx cylindric-campanulate or campanulate ....... aes aia a a RO NA Section 94. Ammodendron Bge. Plants different-from above. 3. 4. Sea eee A eee 69. Flowers in loose rather elongated racemes, purple or violet, rarely yellowish and then the upper stipules connate .............. 70. Flowers in capitate or abbreviated subumbelliform racemes, or spici- form loosish racemes and then the corolla yellow and upper stipules ih | a ee ee ee mT eM a ey 80. Calyx at length more or less inflated, oblong, ovaloid or ovate -glo- BOSE. oh Gries ose ae eh ey eiliavian ib ahi witar empdieagyh Sn Saeelowteedee bona eee ele Ee ar el, Calyx not becoming inflated or modified in shape............ 62 rile 80. 84, 85. Calyx ruptured by the pod, becoming slightly inflated........... Bare etka ketittetees be sk ay ca Ne ehey aye Section 98. Macrocystodes M. Pop. Calyx not ruptured by the pod and included in its cavity... .:..:.. [ob OM DaBS “belong id o> Aah Can ie Abc. ots Fan a Bi Section 99. Macrocystis M. Pop. Rlanis a caulescentiOmMe an lyr SOmr ce eitys criss ot ch lier gh ep or hueeeenbe. maids Ua FiAnts with adevielOped) SUCMUS sat Shs Wel weir av ior oo ow oh et et eh Geol eta a ke My tamu io), Each flower furnished with a bract and 2 bracteoles; calyx cylindric. Pt Se Ce? 4 » vate... . Section 90. Proselius (Stev.) Bge. BWA cteOleis enone, =e, HIST LUE ty eA IES « SeRe oe oe ee 74, Calyx cylindric; leaflets orbicular, ovate or oblong ovate, glabrous above, sparsely strigulose; peduncles greatly exceeded by the leaves. Section 91. Myobromopsis Boriss. Calyx campanulate or cylindric-campanulate; leaflets varying from filiform-linear to lanceolate, with richer vesture ............. PMN Asi @ NMG oRiet cRictia te ck stir hek ot otior ch et ah e®s Section 69. Corethrum Bge. CayeolOnoteCwlIMEar VE Se. x, | EMRE ACMA Ee SED Rem OS STA! Meg eet | hy, 76. Calyx campanulate or campanulate-cylindric .. 2...) oe. ele Leaflets vested with appressed hairs. ... Section 92.. Xiphidium Bege. Leaflets vested with bifurcate spreading or ascending hairs........ Me asda RAMS RE A te hy TERA Tee oe wt hat aha’ Section 77. Tamias Bge. Pods narrowly linear, long, curved. Section 70. Ornithopodium Bge. Pods short, ovoid or oblong, rarely linear-oblong....,.....-. 78. Flowers yellowish or greenish, the standard occasionally suffused with purple; racemes loosish oblong ...... Section 65. Euodmus Bge. EMOWwersh pur pletor wiOlet Ms eee tes ee ela eas Me eye atone) 6 79. Glabrous plants, only the young calyx sparsely vested with appressed hairs; leaflets orbicular or orbicular-oval, rarely oval, 3-4 (7) pairs, wleshy: lowers le2 Gimm lone yirose colored. jiey. s- -l- 1 ee © nhs 5s Al UGLY GO Bean pret Let Aan nace aan oF Section 79. Picrophace Bge. Haimyaplanvs leavesidiiterent from ADOVE.. 26.1.8 a) tetas en) sf fete eli-tir © Peete weds Weed ested UR APMED, % Section 67. Craccina (Stev.) Bge. Dwarf pulvinate perennials; stems of the year abbreviated, procum- bent; leaflets 2-4 (5) pairs, subflabellate, sericeous; flowers solitary or paired in leaf axils, subsessile; stipules completely connate into a sheath. Ammophilous plants. Section 75. Ammodytes (Stev.) Bge. TE LEUNIGS), CHUGISISSANy ANSON YB] OXON AS sre. G coo Gledeeniec oculouyceo “cleturas) Of Cleo celdlines ole Fruticose or suffruticose plants with a well developed aerial caudex. 6: 01-0 On BiG OAD GRDS se Biscieelah ial aie irae eda ik at daria falc are dnc ar Su re a ae le a2, Perennial or suffruticose plants or small shrubs with a very short AST LAN, CAUCIE tne natn soelcletriaia warake ae iChat ear ira alc Tartare ir Alien earn rine bc 84, Calyx at length bladdery-inflated..... Section 102. Cysticalyx Bge. Calyx not changing its shape or at length scarcely to slightly inflated EVOL WP UYOMUASC ll OS AN CEL OOS bt ecuhicd orca i cltoineaecidl air cae ny MlicerAN cuties i. Huemeticlic 83. Calyx at length slightly inflated; standard-limb auriculate-angular Oram wat OaAse wa ss eee) cae) ot laces Section 97. Cystodes Bge. Calyx not changing its shape or at length barely to slightly inflated pil, TWO UB el Oy Mel GVEN YOKG | ancuadwemiomiae Gn One 1O woleoeoiolecmomte SN Seclotowolotandiis 103. Plants acaulescent, subcaulescent or with very short stems .... 93. Piantsswithe well developed stems <.".1:%.%.'.i.'. .'etats se 2 0 +s ee es 85. Leaflets beset with ascending or spreading bifurcate hairs ....... PEEP BO, er RK a, kar ta Ns hy Oe He Sauk Mel My Mocha Section 100. Chaetodon Bge. meatletssbesetawith appressed hatwsSe cers wes. iste e «mle ae ets me 86. Wane Metals, Clenivatwe POX teers the te Mote os Note Garts Meats Tense ee she e Ne 87. 11 88. 89. 94. 95. Wing-petals entire or emarginate at apex; inflorescence capitate. 88. Calyx at length bladdery-inflated. . . . Section 101. Hypsophilus Bge. Calyx at length but slightly inflated, oblong, ultimately ruptured bythe pod Re "Arve wceeety. tee. B Section 81. Pseudoammotrophus Gontsch. Stipulleiss connate ., =<. cf s) cic, sick ft ch cs oy te oe OMI et Reon ete nen ae. eeu 89. Stipule’s Tree. ey yh a a, EO AW ol ich Re IEDs er Hetey Ahae eeae 90. Calyx'campanulate or icylandriescampanulates emails vents nen ene ssiee a Spee esas god. Lb Ae eee! oY Section 71% 'Onobrychiumy Beer (Caillyipe CydliincimlG 5560000000 0D 00 Section 80. Ammotrophus Bge. Flowers yellow, in many-flowered compact racemes............ sali ait oes Gaaginesae te adele evden Section 66. Pedina (Stev.) Bge. Flowers purple or violet, rarely yellow and then racemes abbreviated, Subumbellitorm sstews-t lowered cncric acne ire) obese ecu me ee 91. Pods bladdery-inflatedwije «opens aiew - Section 82. Leucophysa Bge. JEAO SMS Taro, Jollee lel even ae " h 85 bid ooOYOWBUO VO SGANO MONO DMO“ IOC yONCo a daorib oo 6 92. Pods shaggy with spreading stiff hairs. . Section 76. Erioceras Bge. Pods wooly with appressed) or ascending hairs i.) fa) cient ete ok eee ste siteieh OPA EM. one eI OGa. ss Sa Svs Section 92. Xiphidium Bge. Each flower furnished with a bract and 2 bracteoles............. OE oa i) eM A RE UR HES ot, a Section 90. Proselius (Stev.) Bge. BracteOles.-MOMNGy iy. he sdiay bopreuseds eyutelrep tte ls Woplot 6. black hairs, the supulate equal teeth as long as or shorter than the tube; corolla pale lilac drying yellowish or whitish; standard 5-6 (7) mm long, the oblong-oval, round-tipped limb gradually attenuate into the claw; wing- petals 1mm shorter than the standard and exceeding the keel, the oblong limb 1.5 times as long as the claw, with an auricle ca. 1mm long; keel ca. 4-5mm long, the ovate limb equaling the claw; pods inflated, ovaloid-glo- bose, ca. 4mm long, sessile, beakless or minutely beaked, covered with mixed black and white hairs, rarely black or white alone, deeply grooved, 44 bilocular, 2-seeded; seeds oblong-oval, subreniform, 2mm long, smooth, lustrous brown. Fl. June-July; fr, July-August. (Plate DMV ethic S73) Occurring at an altitude of 2000-2850 m, in subalpine meadows and in the upper part of the zone of woodland and scrub formation.- Soviet Centr. Asia: Pam. Al. (Turkestan, Hissar and Zeravshan ranges). Endemic. De- scribed from Yagnob River. Type in Leningrad. Series 2. EROSTRATI Boriss.- Pods round in Outline, beakless, obtuse at apex. ; 48. A. densiflorus Kar. et Kir. in bull. Soc. Nat. Mosc. XV (1842) 329; Bge. Astrag. geront. II, 22. Perennial, 7-20cm tall, branched from base; stems many, weak, re- curved, vested with appressed hairs; stipules (3) 4-5mm long, ovate-tri- angular, acute, membranous, the lower connate the upper nearly free, her- baceous; leaves 3-7cm long, the rachis 1.5 times the length of the petiole; leaflets distant, 4-6 pairs, oblong or ovate -oblong, 3-8mm long, 1-3mm broad, covered beneath with appressed hairs, glabrous above, borne on very short petiolules, subobtuse, rounded-cuneate at base; peduncles axil- lary, 3-10cm long, appressed-tomentose, white hairs predominant in infl. ; infl. densely spicate or capitate, spherical or oblong, 10-30mm long and 10-13mm broad; bracts linear-lanceolate, 3-4mm long, long-acuminate, covered with black hairs; flowers many, sessile; calyx shorter than corolla ca. 5mm long, hirsute with black hairs, the linear teeth 2mm long; corolla dry, dingy violet; standard 7mm long, the rounded limb ca. 4mm long, scarcely retuse, gradually attenuate at base into the claw; wing-petals 6mm long, the oblong apically enlarged limb twice as long as the claw; keel ca. 4mm long, the ovate limb slightly exceeding the claw; ovary stipitate glo- bose; style recurved, ca. 2.5mm long; pods circular in outline, ca. 3mm in diam., beakless, beset with white and black hairs, smooth, conspicuous- ly cross-wrinkled. July-August. Alpine zone, alt. 2800-3200 m, on stony slopes and sandy courses of mountain streams.- Soviet Centr. Asia: Dzu.-Tarb. (Dzungarian range), Tien Shan. (Terskei Ala Tau, Kirgiz range). Endemic. Possibly growing in Dzu.-Kash. Described from Dzungarian Ala Tau, between the rivers Lepsa and Sarkhan. Type in Leningrad. a 49. A. Fetissovii B, Fedtsch. in A.H.P. XXV (1905) 202. Perennial, 15-25cm tall from a strong taproot; stems many, ascending, slender, curved, profusely branched from base, glabrate with sparse white hairs; stipules oblong- or ovate-triangular, the lower membranous, the upper herbaceous and more elongate, 5-10mm long, 2-5mm broad; leaves 5-6cm long, the rachis 1.5 times to twice the length of the petiole; leaflets distant, 5-7 pairs, oblong to lanceolate, 8-18mm long, 2-4mm broad, acute, rounded-cuneate at base, short-petiolulate; peduncles axillary, ex- ceeding the leaves, 5-10cm long, beset with black spreading hairs beneath the infl.; inflorescence densely many-flowered, Spicate globose or oblong, 10-30mm long and 10-12 mm broad; bracts 6-7mm long, linear or lanceo- late, acuminate, white -membranous, thinly covered with black hairs; calyx ca. 6mm long, densely beset with spreading hairs, the teeth equal and equaling the tube; corolla whitish; standard 7mm long, the orbicular limb attenuate into a claw ca. 1.5mm long, scarcely retuse; wing-petals 7mm 45 61 long, the oblong apically enlarged limb twice the length of the claw; keel 5mm long, 2mm broad, the limb inequilaterally oval, the claw 1.5mm long; ovary stipitate, globose, beset with white hairs, surmounted by a filiform recurved style ca. 3mm long; pods circular in outline, 2mm in diam., beakless, densely covered with white and black hairs. June-August. Screes in the Alpine zone, alt. 2400-3500 m.- Soviet Centr. Asia: Tien Shan (centr. part, Atbashi and Narynranges). Endemic. Described from be - tween Bolshoi Naryn and Atbashi. Type in Leningrad. 50. A. Tulinovii B. Fedtsch. in A.H.P. XXI (1903) 315; Fl. Vadzghik: Ve 2148: Perennial, 8-20cm tall; stems branched, glabrous, ascending, 1.5-9cm long; stipules ovate-triangular 3-6 mm long; leaves 2-4 (5)cm long, short- petioled; leaflets 3-5 (6) pairs, elliptic-oblong, 5-8 mm long, 1-2.5mm broad, obtuse, glabrous or nearly so; peduncles axillary, nearly twice the length of the leaves, 5-8cm long, covered with black hairs beneath the infl. inflorescence a densely capitate oblong or ovoid many-flowered raceme, 1.5-2.5cm long, with subsessile flowers; bracts lance-oblong, exceeding the calyx, ca. 4mm long, sparsely covered with black hairs; calyx cam- panulate, ca. 5mm long, densely black-villous, the equal lance-subulate teeth as long as the tube; corolla whitish; standard 7-8 mm long, the sub- orbicular retuse limb 5mm broad, the distinct claw 1.5-2mm long; wing- petals ca. 7 mm long, the limb oblong-obovate, 2mm broad, slightly re- tuse, attenuate at base, twice the length of the claw; keel 5mm long, the limb 1.5mm broad, 1.5 times as long as the claw, the lower margin strong- ly incurved, the upper margin straight; pod circular in outline, ca. 3mm long, beakless, smooth, tomentose with white and black hairs, bilocular, 2-seeded; seeds flattened, unevenly reniform, ca. 2mm long, smooth, brown, lustrous. Fl. June-August; fr. July-August. Banks of mountain streams, alt. 4000-4260 m.- Soviet Centr. Asia: Pam.-Al. (Pamir). Gen. distr. : Dzu.-Kash. (Chinese Turkestan). De- scribed from Pamir. Type in Leningrad. 51. A. imetensis Boriss. in Not. Syst. Inst. Bot. As. Sc. URSS X (1946). Perennial from a strong thick taproot, 20-30cm tall, branching from base, glaucous-green; stems many, 10-15cm long, sparingly covered with short appressed hairs, slender, sulcate, clothed in remnants of distant stipules from earlier years; stipules cauline, connate only in lower part, green, broadly triangular -lanceolate, ca. 4mm long; leaves 5-7 cm long, short-petioled; leaflets 5-6 pairs, oblong or oblong-lanceolate, obtuse, glabrous above, covered beneath with appressed hairs, 10-15mm long, 2-4mm broad; peduncles 2-3times as long as the leaves, axillary, 7-15 cm long, covered with white and black hairs, the rachis sulcate, covered with spreading chiefly black hairs; inflorescence a densely cylindric, many- flowered raceme, 2-8cm long, the lower flowers distant; flowers on short pedicels ca. 1mm long; bracts narrowly lanceolate, acuminate, ca. 3mm long, beset with spreading white hairs; calyx broadly campanulate, ca. 4mm long, hirsute with spreading mostly black and some white hairs; corol- la drying white; standard ca. 6-7mm long, obtuse, scarcely retuse, orbi- cular, broadly cuneate at base, without a distinct claw, ca. 3mm broad in lower third, 5mm at the middle; wing-petals ca. 6mm long, about equaling the standard, the obtuse apically enlarged limb ca. 1.5mm broad, the claw 46 62 1-1.5mm long; keel 4.5-5mm long, the limb ovate, 1.5mmbroad, twice the length of the claw; ovary short-stipitate, surmounted by a recurved style ca. 2mm long; pods ovaloid-globose, ca. 3mm long, beakless, beset with predominantly white hairs and an insignificant proportion of black hairs or pure white, bilocular, sometimes faintly cross-wrinkled; seeds 2.5-3 mm long, Smooth, ovaloid, slightly flattened, olivaceous. Fl. July; fr. July. Sandy banks.- Soviet Centr. Asia: Tien Shan (Naryn river valley, near the mouth of Imet River). Endemic. Described from Naryn River. Type in Leningrad. 52. A. heterodontus Boriss. in Tr. Tadzhik. bazy Akad. Nauk II (1936) ole Capri Ladzhikw Vie tabe 27. Perennial from a stout long taproot, 10-25cm tall, the root-crown much divided, 3-5cm long, the divisions rather stout, clothed in infundibularly connate membranous whitish stipules; stemsmany, slender, 10-25cm long, ascending, simple, sparingly covered with short appressed hairs; stipules triangular, 2-4mm long, herbaceous, the upper subacute nearly free, the lower subobtuse connate at base; leaves 2-4cm long, the peduncle 0.5- 1.5cm long; leaflets (4) 5-7 (8) pairs, elliptic or oblong, 5-12 mm long, 3-4mm broad, obtuse, rather thick, covered beneath with short appressed hairs; peduncles axillary, 2-8cm long, sparsely covered in upper part with appressed short hairs; inflorescence subcapitate, 12-18mm broad, (10) 14-20mm long, elongating infr. up to 30mm; bracts small, linear, 1-2 mm long, hyaline, exceeded by the calyx; calyx campanulate, 2-3mm long, beset with scattered short black occasionally interspersed with white ap- pressed hairs, the tube twice as long as the unequal teeth, of these three linear-subulate and two slightly shorter, triangular from a broad base; corolla drying an intense violet color; standard rounded-obovate, 5-8mm long, scarcely retuse, gradually attenuate at base, without a distinct claw; wing-petals 5-7mm long, the limb entire oblong or oblong-oval, twice as long as the claw; keel 4-5mm long; ovary stipitate; pods angled-globose, 3-4mm long, beakless, deeply grooved, cross-wrinkled, bilocular, tomen- tose with white and black hairs. Fl. July-August; fr. August. Confined to the high-mountain zone, alt. 3400-3500 m., on shores of rivers and lakes, sandy banks and meadows, often in saline soil.- Soviet Centr. Asia: Pam.-Al. (E. Pamir). Gen. distr.: W. Tibet. Described from Pamir. Type in Leningrad. Section 16. HEMIPHRAGMIUM Koch, Syn. Fl. Germ. (1843) 200; Bge. Astrag. geront. I (1868) 21, p.p.- Perennials with more or less de- veloped stems, rarely subacaulescent, vested with simple (basifix) hairs; stipules connate, not united with petiole; racemes initially compact be- coming lax, borne on rather long peduncles; bracteolesnone; calyx campan- ulate;, corolla lilac-violet or whitish (rarely yellowish-white), the tip of keel violet; wings deeply cleft at apex or bilobed; ovary ona stipe (1)2-4mm long; pods on a stipe equaling or exceeding, rarely shorter than the calyx and then not less than (2) 3-4mm long, cernuous, oval, ovate,’ oblong-el- liptic or linear-oblong in outline, unilocular or nearly so, rarely incomplete - ly bilocular, glabrous (very rarely sparingly covered with short appressed hairs). An Arctic-Alpine, andtoa lesser extent mountain-taiga section of Amer- ican and European plants distributed in the mountains of Europe, in Arctic 47 and mountainous Europe and to a lesser extent in North America and in Soviet Central Asia (only in extreme northeast in Dzungarian Ala Tau). Key to Species in Flowering Condition 1. Ovary and pod loosely or sparsely covered with very short appressed hairs; ovary on a stipe ca. 1mm long. (Dzungarian Ala Tau)....... Say ciate, 4h vot! rath Steals ayerg corey Mesute Loo a 4 Ais aah Reco 57. A. sarchanensis Gontsch. + Ovary and pod glabrous; ovary on a stipe 2-4mm long. (Ural, Siberia). 2. Leaflets oblong-ovaloid or ovaloid, rarely oblong-ovoid, not founda at apex, rarely acutish, 10-23 (29)mm long, 5-15mm broad, glabrous, (5) 6 pairs. Tall plants, with stems 15-22cm long. (Transbaikalia: Barewucin Distimilet))) Mwy a eee 54, A. trigonocarpus (Turcz.) Bge. 63 + Leaflets linear-lanceolate to elliptic, acute, rarely round-tipped or re- tuse and then the plants subacaulescent (stems 0.5-1.5cm long)... 3. 3. Calyx teeth slightly shorter than the tube; stems none or short, not execedinglOl5 Siemans ost ms! Le UN Rew sie yeh Pace ee ee 4. + Calyx-tube 2-3 times as long as the teeth, rarely 1.5 times as long and then stems well developed),,,10-45¢em) long... ws). 695 26). sie «ee 5. 4, Stems very short, not exceeding 0.5-1.5cm, thickly coated in approxi- mate black-hirsute stipules 6-9mm long; wings shorter than the stan- dard; stipe of ovary 2mm long. 56. A. pseudoaustralis Fisch. et Mey. + Stems diffuse; ascending, 1-5cm long, vested with white appressed hairs; wings equaling the standard; stipe of ovary 3-3.5mm long. URED CURR TTL Se CERSS RDS MRR ROME SRRS SA OE CM Se 7 Be 58. A. tschuensis Bge. 5. Leaflets lanceolate or linear-lanceolate, 4-5 (6) pairs; stems short, ascending, (1) 2-5 (10) cm long; bracts lanceolate, 3-4mm long. (Trans- Datileal raver re OER: RRR: aS eae tees Loin 60. A. chorinensis Bge. + Leaflets lance-oblong lance-ovate or, if lance-linear, stems longer, (10) 15-45cm long; bracts linear, rarely lanceolate; leaflets (4) 5-9 jOkEB CSCO OMoLe Gino lia ons Sid OU leks ol oMOl Ds lol Gig ote ually Sale to) olG: oo 1c 0 69:9 ¢ 6. 6. Wing-limb as long as the claw; peduncles (1.5) 2-3 times as long as the leaves; racemes oblong, commonly many-flowered, 3-5 (7) cm long; stems erect 15-45cm long. (S. Ural, Altai, Sayany, Lena river valley). Oar athre abe cous iouiniiay ott Dojo boeipsine (a. B/0k ca 18 ley) «he QOS OA ay olen a rai meres + Wing-limb 1.5-2 times the length of the claw...........-..... the Plants (5) 8-15 (17) cm tall, withcommonly prostrate stems 1-7 (10) cm long; leaves 1.5-5cm long; corolla pale violet with a dark spot on the keel (Soviet Centr. Asia, Pam.-Al.. central it. sli)) cue eee SEE GIRL i TN OR SHE ath 59. A. Beketovii (Krassn.) B. Fedtsch. + Plants (10) 15-40cm tall, with erect or ascending stems 10-22 (30) cm long; leaves (2) 4-10cm long; corolla ochroleucous or whitish, the tip of keel dark violet (mountains of Centr. andS. Europe, Pechorabasin, Ural, Siberia to Altaiand N. Mongolia) ..55. A. australis (L.) Lam. =—J Key to Species in Fruiting Condition 1. Pods bladdery-inflated, ovoid or ovaloid subglobose, inequilaterally ovaloid or inequilaterally oblong-ovaloid °).). 2. 2%. 2). 2 2). eis 2. 48 64 65 + Pods inequilaterally linear-oblong, oblong-ellipsoid or lance -oblong, more or less inflated, but all compressed laterally........... 4. 2. Leaflets narrowly lanceolate or linear-lanceolate, (7) 8-20 (25)mm LOT er tedster. mee. A TANS SB, ela ee, 60. A. chorinensis Bge. Leaflets oblong-ovate or ovate-lanceolate, (3) 4-8 mm long ..... 3 Calyx campanulate, the tube about as long to 1.5 times as long as the teeth; leaflets oblong-ovate or elliptic...... 58. A. tschuensis Bge. + Calyx cylindric-campanulate, the tube 2-3 times as long as the teeth; leaflets ovate-lanceolate or lance-oblong, rarely oblong-ovate...... Rate Her eete Le) . Wheels Taphey ey bes 59. A. Beketovii (Krassn.) B. Fedtsch. 4. Pods inequilaterally oblong, carinate ventrally, grooved dorsally, tri- angular in cross section, almost fully bilocular. 53. A. vaginatus Pall. + Pods oblong-ellipsoid or lance-oblong, inflated, but more or less com- pressed laterally, not grooved either dorsally or ventrally or scarcely o + narrowly grooved dorsally, unilocular or nearly so........... 3. SEEOU Slab TOUS! 2.15) <<. aol sto al cine Mevyedho ted Pos aad teh» PMeaMes PRGA, «cake alts 6. Pods sparsely covered with very short appressed hairs .......... 2.5) 5 4d..ol Gus’ Gch 0. OWED (Sbnmets Acie Vein ELPA. a 07. A. sarchanensis Gontsch. 6. Stems very strongly reduced, 0.5-1.5cm long, thickly coated with ap- proximate stipules 56. A. pseudoaustralis Fisch. et Mey. + Stems well developed, erect or ascending, (10) 15-40cm long.... 7. J Leaflets oval or oblong-oval, round-tipped, rarely subacute, glabrous; pods lance-oblong, scarcely grooved ventrally, subunilocular (the in- complete septumyl-125 mim! broad)i. Cav S)l) OY. Oy aren a, eI. AES 3) Guat iA eNO ott ei a 54, A. trigonocarpus (Turcz.) Bge. + Leaflets narrowly elliptic or lanceolate, rarely elliptic, acute; pods semiovaloid or oblong-ellipsoid, carinate dorsally and ventrally, fully unilocular or rarely subunilocular (with a very narrow obsolescent flange from the ventral suture)........ 55. A. australis (L.) Lam. Series 1. TRIGONI Gontsch.- Pods inequilaterally linear-oblong, cari- nate ventrally, g-ooved dorsally, triangular in cross-section, nearly fully bilocular. 53. A. vaginatus Pall. Astrag. (1800) 46; Bge. Astrag. geront. I, 22; II, 23 yhrylemls Zapjsib. Vil, 1648.— "A. versicolor !p..majonoDC./As- trag. (1802) 111.- A. semibilocularis DC. Astrag. (1802) 136; DC. Prodr. II, 291.- A. phacaeformis Bge. Enum. alt. (1835) 69.- Phaca australis Ldb. Fl. alt. III, 270 non L.- Ph. australis Bp. altaica Fisch. in DC. Prodr. II (1825) 274. Perennial, 20-50cm tall, stems 15-45cm long, erect, simple or slightly branched, tomentulose with appressed white or mixed white and black rare- ly spreading curly hairs; stipules beset with white hairs or merely white - ciliate on the margin, rarely covered with white or mixed white and black curly hairs, the lower connate into a scarious sheath, the upper free lan- ceolate acuminate; leaves (4.5) 6-10cm long, the short petiole and the rachis thinly covered with short white appressed hairs or glabrate, rarely covered with curly hairs; leaflets (5) 6-9 pairs, elliptic or lanceolate, rarely the upper ones linear-lanceolate, acute, 9-22mm long, glabrous above, appressed-puberulent beneath, rarely covered beneath or on both sides with spreading curly hairs; peduncles (1.5) 2-3 times as long as the leaves, finely sulcate, beset with short appressed white hairs; racemes 49 66 rather loose, oblong, 3-5 (7)cm long, the flowers subapproximate; bracts lanceolate or linear, (2) 3-4mm long, 1.5-3 times the length of the pedicels, covered with short white or mixed white and black hairs; calyx campanulate, (4.5) 5-6 mm long, rather densely covered with short black subappressed hairs or with these interspersed with few white hairs, the tube 1.5-2 times the length of the linear-subulate teeth; corolla whitish, the tip of keel violet; standard 11-14mm long, the limb obovate or rounded-oval, rounded -bilo- bate at apex, 2.5-4 times as long as the claw; wing-petals 9-11 mm long, the limb oblong-spatulate, apically cleft, equaling to slightly exceeding the claw; keel 7-9mm long, the limb nearly semicircular, slightly incurved, as long as the claw; ovary glabrous, the body as long as the stipe; pods on a stipe equaling or slightly exceeding the calyx, cernuous, gathered ina loose raceme, asymmetrically linear-oblong, pointed at both ends, 15- 18mm long, 3mm broad, carinate ventrally, grooved dorsally, compressed - triquetrous, indurated-membranous, glabrous, nearly fully bilocular, Fl. June; fr. June-July. (Plate V, fig. 1). Stony and gravelly steppe mountain slopes.-European part: V.-Ka. (Mias in Southern Ural); W. Siberia: Alt.; E. Siberia: Ang.-Say. (Sayany, on Lena River). Endemic: Described from the Uba and Yenisei rivers, Type in London. Series 2. SUBCOMPRESSI Gontsch.- Pods semioval or oblong-elliptic in outline, acute at both ends, inflated but more or less compressed lateral- ly, carinate dorsally and ventrally, rarely scarcely grooved dorsally, uni- locular or nearly so. 54. A. trigonocarpus (Turcz.) Bge. Astrag. geront. I (1868) 22; II (1869) 23.- Phaca tri onocarpa Turcz. in Bull. Soc. Nat. Mosc. XV (1842) (Ol Aepinac ae or mis Murez. Hil bale. calc i (1842-45) 317, non Bge. Perennial, 30-45cm tall; stems 15-22cm long, erect, rarely ascending at base, terete, finely sulcate, glabrous; stipules free, broadly ovate or ovate, the upper often oblong, 7-10mm long. short-acuminate, glabrous thinly white -ciliolate on the margin; leaves (5) 7-10cm long, sessile, the rachis glabrous; leaflets (5) 6 pairs, oblong-oval or oval, rarely oblong - ovate, round-tipped rarely subacute, 10-23 (28)mm long, 5-12 mm broad, both faces glabrous; peduncles 1.5-2 times the length of the leaves, finely sulcate, glabrous, 10-17cm long; racemes lax, oblong, 4-6cm long, rather many-flowered, elongating in fruit-up to 10cm; bracts lance-linear, slightly exceeding to 1.5 times as long as the pedicel, 4-5 mm long, thinly black~ hirsutulous or merely black-ciliate on the margin; calyx campanulate, 5- 6mm long, 2.2-2.5mm broad, hirsutulous with short ascending black (rare - ly mixed with few white) hairs, the tube twice as long as the subulate -linear teeth; corolla whitish, the tip of keel violet; standard 12-14mm long, the oval retuse limb 3 times as long as the claw; wing-petals 9-10mm long, the limb oblong, slightly enlarged upward and shortly bilobate at apex, some- what longer than the claw; keel 8-8.5mm long, the limb nearly semicircu- lar subobtuse equaling the claw; ovary glabrous linear, ona stipe ca. 2mm long; pods cernuous, as long as the slender stipe, linear-oblong, pointed at both ends, slightly compressed laterally, carinate ventrally, scarcely grooved dorsally, 15-18mm long, 4-5mm in diam., glabrous, subunilocu- lar, the incomplete septum 1-1.5mm broad. Fl. June; fr. July. 50 = : SN : I? / ~ Colas aS es 4 t Ge, l TOV WN SAWN IMG >» A>h' Lip PLATE V 1. Astragalus vaginatus Pall. —2. A. Beketovii (Krasn.) B. Fedtsch.—3. A. australis (L.) Lam. ol 69 Mountain forests, alpine (woodless Siberian) meadows, often on rather stony ground.- E. Siberia: Dau. (mountains in Barguzin river basin on the eastern shore of Lake Baikal). Endemic. Described from the Barguzin River. Type in Leningrad. 55. A. australis (L.) Lam. Fl. Fr. II (1778) 637.-Phaca australis L. Mant. (1767) 103.- Colutea australis Lam. Encycl. I (1783) 354. Phaca Halleri Vill. Prosp. 45; Hist. Pl. Dauph. III (1789) 473. - Phaca atrata Turcz. in Bull. Soc. Nat. Mosc. XV (1842) 760.-Astra- galus Tugarinovii Basilevsk. in Not. Syst. ex Herb. Hort. Petrop. V (1924) 69.-A. Kaufmannii Kryl. in Animadv. Syst. ex Herb. Univ. Tomsk. No. 3 (1932)2; Kryl. Fl. Zap. Sib. VII, 1650.-A.uralensis Litw. in Bull. Soc. Nat. Mosc. (1893) 501. Perennial, (10) 15-40cm tall; stems ascending or erect, 10-22 (30)cm long, simple, glabrous or vested with short appressed rarely spreading hairs; stipules more or less connate and united with petiole, triangular - ovate or elliptic, acute, 4-10mm long, glabrous or covered with black or mixed white and black hairs; leaves (3) 4-10cm long, both the petiole and the much longer rachis more or less pubescent; leaflets (4) 5-6 (9) pairs, narrowly elliptic or lanceolate, less often oblong, rarely elliptic, (6) 8-25 (40)mm long, (2) 3-7 (12)mm broad, acute, glabrous or covered beneath rarely on both faces with appressed or shaggily ascending hairs; peduncles equaling to rarely 1.5-2 times as long as the leaves, glabrous or puberulent; bracts linear, rarely lance-linear, 3-6 (7)mm long, covered with short or very rarely white hairs; calyx campanulate, 5-7mm long, densely hirsute with semiappressed black or very rarely white hairs, the tube (1.5) 2-2.5 times the length of the linear-subulate teeth; corolla yellowish-white or whitish, the tip of keel dark violet; standard 10-15mm long, the broadly oval retuse limb 3-4 times as long as the claw; wing-petals 8-13 mm long, the limb enlarged apically, rather deeply cleft, 1.5times as long as the claw; keel 7-12 mm long, the semicircular acutish limb equaling the claw; ovary on a short stipe 3-4mm long; pods as long to twice as long as their slender stipe, subcernuous, semioval or oblong-elliptic in outline, pointed at both ends, (11) 15-30mm long, 5-9mm in diam., acuminate intoa straight subulate beak 1-1.5mm long, carinate dorsally and ventrally, inflated, membranous, glabrous, unilocular (sometimes with a very narrow flange from the ventral suture). Fl. June-August; fr. July-August. (Plate V, fig. 3). Deciduous forests, scrub and treeless heights (in the alpine tundra), snady river banks, rocks, rarely in arctic tundra, occasionally (European part) in pine woods and on gypsum.- Arctic: Arc. Sib. ; European part: Dv. - Pech. (Ust'-Pinega), V.-Ka. (Ural near Zlatoust and Kungur, along Chuso- vaya River, and Berezov District); W. Siberia: Alt.; E. Siberia: Lena-Kol., Ang.-Say., Dau. Gen. distr.: Centr. Eur. (E. and Centr. Carpathians, Alps, Apennines, and Pyrénées); Mong. (E. Mongolia). Note: Exceedingly variable inanumber of characters, suchasvesture, shape of leaflets, size and shape of calyx, length of pod-stipe, pod shape, etc. The species will undoubtedly have to be divided into a number of geo- graphic races, but the division is at present impossible in view of the in- adequate collections. Considering the impracticability of a full treatment of the entire complex of closely related forms which have been here gathered underthename A.australis (L.) Lam., we have indicated allthe species described by various authors as well as related species ina list of synonyms. 52 70 Besides these species, the North American A. abori genorum Rich, has been reported by A.I. Tolmachev for Eastern Siberia. Comparison of Amer - ican specimens with specimens from Siberia has shown that the latter differ materially in shape and size of calyx, rather resembling in this respect the Siberian Phaca atrata Turcz. 56. A. pseudoaustralis Fisch. et Mey. in Suppl. ad Indicem XI Sem. Hort. Petrop. (1843) 16; Bge. Astrag. geront. I, 22; II, 24; Kryl. Fl. Zap. Sib. VII (1933) 1649-1650. Perennial, subacaulescent plants, 7-15cm tall, with a stout taproot and creeping, branched, buried offshoots; aerial stem 0.5-1.5cm long, clothed in stipules; stipules membranous, approximate, beset with black hairs, connate through 1/3-1/2 their length, 6-9mm long, sheathing the petiole, the lower obtuse, broadly vaginate, the upper lance-ovate subobtuse; leaves (2) 4-10cm long, the petiole (0.6) 1.5-3cm long, both petiole and rachis thinly covered with short white subappressed hairs; leaflets 6-9 pairs, ellip- tic or oblong-ovate, 4-9mm long, subobtuse, rarely orbicular or scarcely retuse, glabrous above, covered beneath with sparse appressed short white hairs; peduncles slightly longer than the leaves, rarely as long, thinly ap- pressed-hairy, 4-9cm long; racemes rather loose, short, 4-10-flowered, 2-3cm long, the flowers divergent, the lower ones nodding; bracts lanceo- late, (3) 4-5mm long, subobtuse, beset with black hairs; pedicels 1-2 mm long; calyx campanulate, 7-8mm long, black-hirsute, the linear-lanceolate teeth slightly shorter than the tube; corolla violet; standard 12 mm long, the limb obovate retuse, the claw broad cuneate 3mm long; wing -petals 10- 11mm long, the limb spatulate-oblong unevenly bilobate, the claw 4mm long; keel 9mm long, the limb semicircularly concave dorsally, straight ventrally, obtusish, the claw 4mm long; ovary linear-lanceolate, glabrous, on a stipe 2mm long, the glabrous style shorter than the ovary; pods thinly membranous, triquetrously ovoid, 12-16mm long, obtuse or very short - acuminate, glabrous, on stipe 4-5mm long, unilocular (a septum not more than about 1 mm broad issuing from the dorsal suture), divergent, the lower ones cernuous. Fl. June; fr. June-July. Stony slopes of the Alpine zone, inthe gravelly and the moss -lichen mountain tundra.- W. Siberia: Alt. Endemic. Described from the vicinity of Ridder- skii mine. Type in Leningrad. Series 3. INFLATI Gontsch.- Pods ovate, oval, asymmetrically oval or asymmetrically oblong-oval in outline or subglobose, bladdery-inflated, fully unilocular or nearly so. 57, A. sarchanensis Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, (10) 15-22cm tall; stems 3-11cm long, ascending or nearly prostrate, thinly vested with very short spreading hairs; stipules connate in lower 1/3 to 1/2, the upper sometimes free, ovate, the lower oblong, acute, (3) 5-7mm long, the margin minutely white- rarely black-ciliolate; leaves 3.5-7cm long, the petiole and the much longer rachis covered with short appressed rarely spreading hairs; leaflets 5 pairs, oblong -ovate rarely ovate-lanceolate, acute, (5) 8-14mm long, (2) 3.5-6mm broad, cov- ered on both sides or only beneath with very short ascending hairs; pedun- cles twice as long as the leaves, sparsely covered with white and, be- neath the infl., also with black hairs; racemes lax but with approximate 53 72 flowers, ovaloid or oblong-ovaloid, 2.5-3cm long, many-flowered; bracts lanceolate, 2.5-3mm long, twice the length of the pedicel, beset with short black and white or exclusively white hairs; calyx campanulate, 5mm long, pubescent with mostly black and some white subappressed hairs, the tube 1.5 times as long as the linear-subulate teeth; standard 9-10mm long, the limb ovate, bilobate to 1/4 from apex with subtriangular obtusish lobes, 3- 4 times as long as the claw; wing-petals 7-8 mm long, the limb enlarged upward, cleft to 1/3-i/2 from apex, slightly longer than the claw; keel 5.5- 6mm long, the semicircular obtuse limb as long as the claw; ovary on a stipe 1mm long; pods (immature) on a stipe 2mm long, sparsely and mi- nutely hirsutulous with appressed hairs. Fl. July. Rocks in the high-mountain zone. -Soviet Centr. Asia: Dzu.- Tarb. (Dzun- garian Ala Tau). Endemic. Described from the sources of Sarkhan River. Type in Leningrad. 58. A. tschuensis Bge. Astrag. geront. I (1868) 22; II (1869) 24; Kryl. Fl. Zap. Sib. VII, 16951. Perennial, (5) 7-10cm tall, with a stout much divided root-crown; stems diffuse ascending, 1-5cm long (rarely shorter), vested with white appressed hairs; stipules cauline, 4-5mm long, connate to 1/3-1/2 (2/3) the length, the lower obtusely triangular, broadly vaginate, the upper ovate-lanceolate; leaves 3-5cm long, the petiole 1-2 mm long, both petiole and rachis covered with white appressed hairs; leaflets 7-8 pairs, oblong-ovate or elliptic, obtusely acuminate, (3) 4-7mm long, glabrous above, cinereous beneath by appressed to somewhat spreading hairs; peduncles 3-6cm long, slightly longer thanthe leaves, rarelyas long, covered with short white hairs; racemes abbreviated, (4) 6-10-flowered; bracts membranous, lanceolate, 3-4mm long, 3(4) times as long as the pedicel, but exceeded by the calyx, beset with black hairs, ca. 1mm long; calyx 7-8mm long, campanulate, black- hirsute, the lance-linear teeth slightly shorter than the tube; corolla violet; standard 12-15mm long, the limb obovate, broadly emarginate; wing-petals equaling the standard, the limb deeply bilobed (the lobes divergent), twice as long as the claw; keel shorter than the wings, 9-12 mm long, the limb semicircular dorsally straight ventrally; ovary on slender stipe 3-3.5mm long, glabrous, 14-18-ovuled; pods spreading or cernuous, bladdery-in- flated, broadly ovoid, thinly membranous, 12-17mm long, 8-12 mm broad, glabrous, unilocular (septum practically absent), the stipe shorter than the calyx (3-3.5 mm long). Fr. June-July. Gravelly slopes of the Alpine zone.-W. Siberia: Alt. (rarely in Chuiskie Belki, upper course of Taldura River, Chegan~-Burgaza river valley and Kalgutta valley). Endemic. Described from Chuiskie Belki. Type in Lenin- grad. 59, A. Beketovii (Krassn.) B. Fedtsch. in Bull. Herb. Boiss. sér. II, V, (1905) 316.- Oxytropis Beketowii Krassn. in Script. Hort. Univ. Petrop. A, 1 (1889) 15.- A. polychromus Freyn in Bull. l'Herb. Boiss. sér. 2, IV (1904) 454.-Ic.: Fl. Tadzh. V, tab. 29. Perennial, (5) 8-15 (17) cm tall; stems many, mostly prostrate, 1-7 (10) cm long, glabrate; stipules connate in lower part, broadly ovate or the upper ones lanceolate, 3-5mm long, more or less covered with white rare- ly mixed black and white hairs or merely ciliate on the margin; leaves 1.5- 5cm long, the petiole much shorter rarely as long as the rachis, both peti- ole and rachis covered with short appressed white hairs; leaflets 5-7 pairs, ovate, lance-oblong or oblong-ovate, 5-8mm long, 1-3mm broad, obtuse 54 —---~ —_ ———————————— to subacute, covered on both sides with appressed or ascending grayishhairs or glabrate above; peduncles 2-7 cm long, covered with white and beneath the inflorescence with mixed white and black hairs, sometimes glabrate; racemes loosely contracted, (5) 8-17-flowered, spherical or rounded-ovaloid, 1.5- 3cm long, elongating infr.; bracts lance-linear, 3-4mm long, beset with short black and white hairs, half as long as to exceeding the calyx-tube; calyx campanulate, 6-8 mm long, hirsutulous with black and fewer white hairs, the tube 2-3 times the length of the lanceolate teeth; corolla lilac- violet, with a dark spot at the tip of the keel; standard 14-16 mm long, the rounded -obovate deeply emarginate limb twice as long as the claw; wing- petals 12-14mm long, the limb enlarged upward, bilobed, twice as long as the claw; keel 9-11 mm long, acutish; pods cernuous, on slender stipe ca. 3mm long, bladdery-inflated, asymmetrically ovaloid rarely inequilaterally oblong -ovaloid, 1.5-2.5cm long, glabrous, lustrous, red-spotted, mem- branous, unilocular. Fl. June-July; fr. July. (Plate V, fig. 2). Stony or fine-earth stone-covered mountain slopes inthe alpine vegeta- tion zone, alt. 3500-4300 m.- Soviet Centr. Asia: Pam.-Al. (E.), Tien Shan (E.). Endemic. Described from Sary-Yazy River, near the city Khan- tengri. Type in Leningrad. 60. A. chorinensis Bge. Astrag. geront. II (1869) 24.- Phaca are- momioe all Its App. (Li7 6) 48 wel dbigk eRossielynd hie wbunczs EY. balcedaheal 293,.n0n, A. arenarius jauct, al. = Ics: Palléstabei@sies flee. Perennial, (5) 6-15 (17)cm tall, from a woody root-crown; stems rather many, more or less ascending, (1) 2-5 (10) cm long, vested with short ap- pressed white hairs or glabrous, rarely villosulous with spreading white hairs; stipules connate to 1/3-1/2, sheathing the petiole, scarious, white ciliate on the margin, rarely villous throughout with short spreading white hairs, ca. 4mm long, ovate-triangular subobtuse in the free portion, the lower semicircular rounded-obtuse; leaves 2-5 (6) cm long, the petiole 0.5-1 (2) cm long, both petiole and rachis covered with short appressed white hairs or glabrous; leaflets 4-5 (6) pairs, lanceolate or linear-lanceo- late, acute, (7) 8-20 (25) mm long, glabrous, rarely covered beneath with scattered appressed white hairs or villosulous with spreading white hairs; peduncles about equaling to exceeding the leaves, 2-5cm long, glabrous, very rarely covered with short appressed white hairs; racemes short, densely 6-12 (14)-flowered; bracts lanceolate, 3-4mm long, scarious, the margin black- and white-ciliate; pedicels ca. 1mm long; calyx campanulate, 6-7mm long, covered with mostly black and some white hairs, the teeth triangular -lanceolate or lance-subulate, 1.5-2mm long; corolla violet (?); standard 14-15mm long, the limb broadly oval or broadly obovate, retuse, the claw broad cuneate 4mm long; wing-petals 12-13mm long, the limb Spatulate-oblong, unequally bilobate, the broadly oblong auricle ca. 2mm long, the claw ca. 5mm long; keel 10-11 mm long, the limb semicircular - concave dorsally, straight ventrally, the claw 5-6cm long; ovary glabrous, linear-lanceolate, on a stipe ca. 3mm long, the glabrous style shorter than the ovary; pods bladdery-inflated, thinly membranous, (9) 12-17mm long, ovaloid to subglobose, on a stipe 3mm long, glabrous, devoid of septum (fully unilocular); seeds reniform-oval, compressed, smooth, brown, ca. 2mm long. Fl. May-June; fr. June. Stony mountain slopes, steppe (mostly stony) slopes, among thickets of steppe scrub, in open pine woods, and on riverside pebbles.- E. Siberia: Dau. Endemic. Described from Transbaikalia. Type in Leningrad. 59 Section 17. HEMIPHACA Bge. Astrag. geront, I (1868) 20.- Peren- nials with well developed stems, vested with simple hairs; stipules partial- ly connate, or theupper or all stipules free; leaves imparipinnate with mar- cescent petiole; racemes lax or more or less compact, borne onrather long axillary peduncles; bracteoles none; calyx campanulate; wings exceeding the keel, cleft at apex, more rarely emarginate or.entire; pods sessile or subsessile, rarely on a stipe up to 1.5mm long, semiovate or semioval in outline, globose -ovoid or asymmetrically ovaloid-globose, mostly erect, rarely spreading or cernuous, membranous, glabrous, very rarely slightly hairy, bilocular, subbilocular or unilocular. 74 An Asian section covering Eastern Siberia, Mongolia and Soviet Central Asia, reaching southward as far as Himalayas and Tibet. Mesophytes. Key to Species in Flowering Condition 1,, Ovary and pod glabrous), . -..4.6,-5- i, 2 © -.9 sean Gee: dial) wee 2. + Ovary and pod hairy; pods small, wrinkled-nervose, inequilaterally rounded-ovoid; leaflets linear-lanceolate or linear, 8-20 mm long, 2- D iiraaioa’ [oreo (laraiarate os clio o Ole ooo 51d Ow c 68. A. rytidocarpus Ldb. 2. Limb of wing-petals entire at apex, obtuse. ....-.-+--+++++ee-: 3. + Limb of wing-petals emarginate or cleft. ......---+-+++sseee- 6. 3. DStipulestireensh.jj. “sysop 2784 a8 Pl 8B onich o SO Kemet tains ce Ree 4. + All stipules or only the lower ones connate...--.-++++++s+e-> 5. 4. Racemes elongated, loosely many-flowered; corolla 9-11 mm long; calyx 2.5-3mm long....-.-+-.-+-+++-+>> 67. A. macropterus DC. + Racemes short, rather loosely capitate, 8-15-flowered; standard ca. 15mm long; calyx 4-5mm long........-- 61. A. sachalinensis Bge. 5. Tall plants, up to 40-60cm in height; leaflets 13-25 (30)mm long; co- rolla 11-14mm long; stipules lance-linear or linear, ca. 3mm long. eS ER EB ta) ea EM cite its. kell 69. A. hemiphaca Kar. et Kir*. + Low plants, 11-12cm in height; leaflets (5) 8-11 mm long; corolla ca. 9mm long; stipules lanceolate, ca. 5mm long ...--+-+++-++++-+-> cia Ri Ne MN DDS Rh aa, SR ONECMLEHGY toreleY oi 71. A. kenkolensis B. Fedtsch*. 6. Calyx-tube equaling the’ teeth). 6/0) 28's 22 oe ee ea i a fic + Calyx-tube 1.5-3 times as long as the tSSEH YC, 29. HM aie career 9. 7. Standard 9-11 mm long, the limb oblong-elliptic; calyx-tube 1.5-2 time the length of the teeth; bracts linear (N. Pam )=Aly) 8, “OR Pee LAAT SPUN SEN AE es YE LORY RCE SPR ENS alae yet, citar Dai eh te 75. A. kokandensis Bge. + Standard 13-16mm long, the limb ovate; calyx-tube equaling the teeth or nearly so; bracts ovate, oblong-ovate or oblong-oval....-...---> 8. 8. Plants (23) 25-40 (45) cm tall; stipules free; limb of wing-petals cleft to 1/3-1/2 the length (Kulja, Centr. Tien Shan). 73. A. laceratus Lipsky + Plants 20-23cm tall; stipules connate in lower part; limb of wing - petals short-cleft (rarely subentire). (Chotkal'skii range) .......-- TED EY EAES oh) AES CAAA Ws" og can atte, Ete: Be 74. A. aflatunensis B. Fedtsch. 9, Leaflets glabrous. ...-.---...-.-0+-++08- 62. A. versicolor Pall. Leaflets hairy at least on lower side... 2-1-1 ese es ee ee 10. 10. Racemes remotely flowered, 5-15cm long.. .63. A. puberulus Ldb. 75 + Racemes rather contracted, ovoid or subglobose, 1.5-3cm lone ave: a See also 70. A. abramovii Gontsch., p. 652. *® See also 72. A. tauczilikensis Golosk., p. 652, 56 76 La. 1UPate atowot pw Leaflets shaggy on both sides with short spreading curly white hairs. He Ia: em eemmnirgh Mat aiahiawercanide SRReel J aki. ela ee 64, A. olchonensis Gontsch. Leaflets glabrous above, covered with appressed hairs or glabrate beneath; pods horizontally spreading or slightly cernuous...... 12. Leaflets oblong-oval, commonly retuse, rarely orbicular; standard SOiaMMlOn ees Ls) d aie ies ak Rech nae hice 65. A. multicaulis Ldb. Leaflets lanceolate, acute; standard 11-12mm long............ Pa Molcet oe taiea SLMS NEN aa dik eit meena beatae. 6 Sh, amenay oi. 66. A. vallicola Gontsch. Key to Species in Fruiting Condition Bods elabrousiey. (bi oY LON Re TT NS: WES ATER MERLE SOO tere: SORT MU het De Pods minutely hirsutulous with appressed white hairs, asymmetrical- ly rounded-ovoid, 4-5mm long, transversely rugose; leaflets lance - iineateLonunear, 6-20 mim lone, 22,0 mind TOad: 5 sc swe sre ce ctl a PEER AT sia Yat cin ais 6) RUMMY LO G1EY JY -+..... 68. A. rytidocarpus Ldb. VOGSMUMTLOCULAT oe ee esl ee e eoe te ahem RR SRS LE yey ane eer 3 Podcaollocwlar Onmearly SOn cia aie a icedae aioe Laelia ect thot a) eens Ms Bracts ovate or lanceolate, 5-14mm long................-. 4, BGacts ance Linear or ineanr, i2-o mim LONG weve semen). vamalcns a 6. Stipules free; rather large plants, with stems 15-30cm long...... od! BS: Bibb aIyAiie RE ASCIG OMEN Ata Sts ecu Ane RIS Mees An id 73. A. laceratus Lipsky Stipules connate in lower part or merely at base; larger plants, with SUCTIS PA OTCTONVONS aia te sitet ah tee eM tate ce Nay te acc etm Man ge Tati re Naina ta ct Ra 3. Leaflets 6 (7) pairs, glabrous; calyx 7-8mm long............. PMetemrame te Welis feels poner stoulelre ets 74. A. aflatunensis B. Fedtsch*. Leaflets 4-5 pairs, glabrous above, covered beneath with scattered appressed hairs; calyx 4mm long.. 71. A. kenkolensis B. Fedtsch. Plants 40-60cm tall; pods broadly ovoid, almost bladdery-inflated, scarcely carinate dorsally and often ventrally, 8-12mm long...... Meee Serie. clone Ps! eulapie grey suuel veel scene te 69. A. hemiphaca Kar. et Kir**. Plants lower, 15-35cm tall; pods ovoid, carinate ventrally, concave dorsally, 6-8mm long, subtriangular in cross section.......... PR Uke i MLV (oi sisteiigcib, suusm c's | sky Tap ssh lenets: es dom, e4 bonis 75. A. kokandensis Bge. Least Joby BUSKON aoe BNUMSNy In alien igo iy oes cucecioiears tamer o untae sna 8. Meatlersrolabmousvabove, sat. 1) site © Ge bie ie io a She ueihebielle: ola lle Me 9. Leaflets covered on both sides with short appressed hairs; stems 15- 40cm long; racemes lax, 5-15cmlong..... 63. A. puberulus Ldb. Leaflets shaggy on both sides with short spreading curly hairs; stems 8-13 cm long; racemes contracted, ovoid, 2-2.5cm long......... BF oo Bi Toto Bert BARS Sabah ok ARB CURL Oey ae es Legacies RO 64. A. olchonensis Gontsch. EOdSISEE CORP ASCENGIN GH ier cul: PNA atid Loulalas o (albbl isi. a) Joh al outset tail at?olile 10. Podsvhorizontally spreading OF Cernuous) ooasind ysinekil vo feasts 113. A. floccosifolizs Sumn. Calyx covered with appressed soft hairs; pod rounded dorsally .... 3a: Oi Oh Cat GOO cw aOnemenesy chides: same 114. A. ephemeretorum Gontsch. Leaflets silvery-velutinous above ........-.-2++6-+++22+eee- 20. Leaflets villous, appressed-hairy, sericeous or glabrate above. 26. Stipules lanceolate or broadly lanceolate; calyx campanulate; flowers IES) sag aa\l Jo yaten MeN Ee eS Io Ne SSM ele 106. A. pseudoamygdalinus M. Pop. Stipules linear-subulate; calyx cylindric; flowers ca. 30mm long ... eR eae aioghs fo! (o WPk eR cage Tati te 117. A. juniperetorum Gontsch.. Leaflets orbicular or oval-orbicular, 3-6 (9)mm long; cespitose acaulescent plants, 7-l12cmtall........ 101. A. sisyrodites Bge. Leaflets oval, oblong-oval or oblong ovate ................ 27. 81 110 itis 28. 29. co + co t Ovary subsessile; inflorescence capitate, spherical or ovaloid, com- pact or rather looSé: .a.qmiaiamos, Men eie- ta VPS ea bos eae 28. Ovary long-stipitate, the stipe as long as the body or nearly so; in- floréscéence loose) oreloosish we) ihiyannes siisttinn Cloehampig® « 29. Standard-limb oblong, oval-dilated in lower part; claw of wing-petals halt the length ofthe litt.) © p5 cute oe aye rue 95. A. lasiopetalus Bge. Standard-limb oval-oblong, broadly oblong or suboval-quadrangular, scarcely attenuate in lower half; claw of wing-petals equaling the limb Or Hearily: SOy{ 0 Ms Mae le shoes ew elesy os 96. A. dasyanthus Pall. Leaflets (10) 15-20 pairs; peduncle together with infl. 13-22 cm long, slightly shorter than the leaves; racemes very loose; flowers 25- ZS OMTMIGLON Sere, suey teary. ~Meee mare ce fares 104. A. Litvinovianus Gontsch. Leaflets 12-14 pairs; peduncle together with infl. (2:5) 5-7cm long, much shorter than the leaves; racemes loosish; flowers 17-20 (22)mm Noy eleh Mo ROW 0 ey a ccagririete cere mr Mane GRAM oo ia 105. A. amygdalinus Bge. Key to Species in Fruiting Condition Stipules green, herbaceous; stems long, branched; racemes axillary; pods oblong, 14-15mm long, compressed, carinate ventrally, rounded or flattened dorsally, incompletely (to 1/2-2/3) bilocular......... Sis oo aay ss ci. CR, AGIs EMME Ree aioe Se) ub 84/6 94. A. orbiculatus Ldb. Stipules whitish=sCarious or menmibranous. 2. a. 3 s+ «1s cece Don Corolla clabrous .. vam. oc) ct 1s 6 + ely eousenns 93. A. arianus Gontsch. Corolla more or less pubescent 2... sca. aos oes Ge © = bogs eee ae Pods nuciform, rigidly coriaceous, compressed, asymmetrically ovaloid or asymmetrically oblong-ovoid, carinate ventrally (with a ' hard thick suture), inflated upward and convex or shallowly grooved dorsally, incompletely to nearly fully bilocular.. .. 2.0 0 ss 4. Pods coriaceous, hirsute, villous or lanate.............4... hs Plants with stems (8) 25-50cm long; leaflets covered on both sides with, Short appressed Nags. cost sus cps spe 3, MR s,s ool sea yams as) ie Oe Plants acaulescent or nearly so (rarely with stems up to 6mm long); leaflets silvery-velutinous on both sides with appressed hairs... .6. Leaflets (10) 15-20 pairs; racemes very loose with distant pods; pods (stipe and beak excluded) 22-25mm long, grooved dorsally, the stipe TROT OMe ts meee ee ee a ee 104. A. Litvinovianus Gontsch. Leaflets 12-14 pairs; racemes rather loose with approximate pods; pods 17-20 (22)mm long, flattish dorsally, the stipe 2.5-3mm long. 5, Re Cet: Me sees oc SERN e. AP ae Tenn MONS} Feces 105. A. amygdalinus Bge. Bracts lanceolate to broadly lanceolate; calyx campanulate........ Bc: ch ac ciara a ~ ses... ..106. A. pSeudoamygdalinus M. Pop. Bractspinear-subulate; veallyx) cy lindmic 2% 00) aes eee SORE! Gc EAD: 6 GRORORERSS: COCREMERSE A Urecuemcmr ticle 107. A. juniperetorum Gontsch. Pods carinate, rounded or flattened dorsally (rarely scarcely con- CAVE) co cy a ts we ec dvate li cite age) sta ls a ROMO so etpeer ah one «ten (ental eon 8, Pods more or less distinctly prooved or suleaté 220 5 Ss Wels Leaflets velutinous with rufous hairs, (2) 3-4.5 (5)cm long....... SUA ERUSEAe alu eMCey. Chan tue tel trees aiouley ca lucd Cepe ea COee 103. A. holosericeus M. Pop. Leaflets floccose, glabrate or hirsute, sericeous or villous .... 9. Leaflets lanceolate, densely canescent-hispid on both sides, (5) 7- 10mm long, acute; pods sessile, subradical. 102. A. Tekutjevii Gontsch. 82 a4, 25. Leaflets appressed-pilulose, glabrate, white-floccose or sericeous. ome cE e MON OW fo SES AEP eh ty le tote te We tee be eRe on me Ee o 10, Standard-limb oblong, more or less ovally dilated in lower part... . SEA, EMEP CM TOT Ske Ny CAE gh te ke ey 95. A. lasiopetalus Bge. BEandardalimbiditfenent! inishapel, suse l ts a ae ate ole eee oat Iie Bullies exh oben Gry IAA ile seewoay leyaVet a A eae ein Gite ee Ze Calyx campanulate or cylindric-campanulate, 6-11 (15) mm long .15. Upper face of leaflets sericeous with soft reddish hairs ......... a Melee EN se ee ee eRe 100. A. Nicolai Boriss. Upper face of leaflets glabrate or more often floccose with soft white hairs (at least on the midvein) mod cron lony) On/Ove2loid=lanceolatet aaa fy Ae heen ee 14, Pods ovaloid or oblong-ovaloid.......,... 199. A. mucidus Bge. Calyx narrowly cylindric, the teeth half the length of the tube; a ces- pitosersuiinuticose'plant . 4 tisk). 2 ak 115. A. ochranthus Gontsch. Calyx tridentate, the teeth slightly shorter than the tube; a herba- COOUSAOLAMCN. fosae hs thoes te, 21. A. ephemeretorum Gontsch. Leaflets villosulous on both sides; calyx-teeth twice the length of the LIOLO 60 of Mere scan) OP, a Se eS es a 101. A. sisyrodites Bge. Leaflets floccose above with soft white hairs (at least on the midvein), Pods with a narrow partial septum (imperfectly unilocular)..... 18. Calyx campanulate, 9mm long; standard-limb obovate.......... ED alias feu) Mott cus key Eh ote) St Ak eS 118. A. densus M. Pop. Calyx cylindric-campanulate, 10-11 mm long; standard-limb ovaloid- VMOTAE eine a GI aia Sa oie 119. A. bischkendicus Gontsch. HMOROVE SEMA CESCONt i.e is % acs b4e 6 5 cou 120. A. Korolkovii Bge. Petioles becoming indurated, partly or fully persistent.......... SUM Me Eee OLE oP cy aioe susie, ale) 8 pots 117. A. babatagi M. Pop. Pods covered with short soft hairs; calyx hirsutulous with more or lesstaporessed Wars. 6 cues be 6 & sos be 120. A. karategini Gontsch. OMIsROndec al yxavAll OUS Pia. (ek: jon op Seis home eases, s 6 eae ee ro a An Leaflets glabrous above, 9-14 pairs, obovate SQUIOV BSS BY 5 Gla clo wbee Zilye Leaflets pubescent above, rarely glabrous or nearly so and then (14) ISB (AM}) PERSIANS Nes. SAPS eta tN at nN. teat ava el en iN eaten PAS) Sera resales Last LOMO Sg ec cous) aistaie ses) ae 110. A, oeroilanicus M. Pop. (CEISTES (UD) BOE CAE NTo a) a1 Ko 0 aR lee eae Daa at inet) opt. la 22. Calyx broadly cylindric, ca. 20mm long, 4-5mm broad......... 57 BUGGIES Rein ema ngn 109. A. rubromarginatus E. Czernjak. Calyx narrowly cylindric, (15) 20-25mm long, ca. 3mm broad.... 20 6 81006 "SV SSR ene eS ne al aaa UE yaa 108. A. grandiflorus Bge. Upper face of leaflets glabrate or more often floccose with soft white LLERITES) 5 es 6 “Tae tS es ae ela Sa A teh cane ale ae arly ay. 24, Upper face of leaflets sericeous with appressed and rather long hairs, lanateeiuNlous Orimappressed-VillOUS . . i. 74 sca cs he te ee 26. Standard-limb oblong, dilated in lower part. 95. A. lasiopetalus Bge. Standard-limb obovate or obovate in upper part and oblong-attenuate aera ia sy utc ol Gi Ned Mc ere Saha Mo che, Spas ono amathelee Siew ious Syl 29. Pod nearly fully bilocular; standard-limb obovate-dilated in upper part, oblong-attenuate below, angular at base..............s... O1ChG O-RCh O po Og On CREM OF OUP ce RO Ree 1 eT ee 111. A. lanuginosus Kar. et Kir. 83 + Pod bilocular merely through 1/2-2/3; standard-limb obovate...... eee ide Loui atl Geateacaniald alias ewe cyuslailicue akmaas at ee 113. A. floccosifolius Sumn. 112 96, Calyx-teeth eaueinne to 1.5-2 times the length of the tube; leaflets i glabrate or white-villous above......... 99. A. anisomerus Bge. + Calyx teeth shorter than the tube, rarely as long and then leaflets red- dish-SericeOuSabDOVE® «isi ise yeti esac? SR he eae Sr ec ithe 27. Plants with more or less developed stems; pods in compact capitate or ovaloid, rather many-fruited, racemes borne on peduncles 5-15cm longa’ tedee tet ugh OL cts ae tigello ss dedia do! ME ee 96. A. dasyanthus Pall. + Acaulescent plants; pods in loose or compact racemes, in the former case the racemes subsessile (subradical) or more rarely short-pe- duncled) (peduncles) upito 2\em long) mie sai-e ee cee eh cee 28. 28. Fruiting racemes loose, 6-20-fruited, the peduncles 3-7 (10)cm long. ‘cei th of AP Oath Be Wine. eet Mee ei en 97. A. tanmaiticus C.Koch + Fruiting racemes 4-8 (9)-fruited, sessile (radical) or nearly so (pe- duncles normally not more than 2cm long). ..98. A. pubiflorus DC. Series 1. NUDIFLORI Gontsch.- Stems long; stipules free, cauline, not united with petiole, scarious; racemes few-flowered, loose, borne on axillary peduncles; corolla glabrous. 93. A. arianus Gontsch. nom. novum.-A. Michelsoni Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS. VII, t: 7 (1938)'150; non B. Fedtsch. Perennial, 40-52cm tall; stems 37-48cm long, lanate with white spreading hairs; stipules triangular, 5-7 (10)mm long, ciliate; leaves ca. 5cm long, the petiole obsolete, the rachis covered with spreading hairs; leaflets 7 pairs, oblong to oblong-lanceolate, (8) 10-14 (15) mm long, round- ed-obtuse, mucronulate, glabrous above, densely covered beneath with short appressed hairs; peduncles 2-2.5cm long, shaggy with woolly spread- ing pubescence, together with infl. equaling the leaves or nearly so; ra- cemes (4) 5-6-flowered; bracts narrowly lanceolate, 4-5mm long, shaggy- hispid pedicels ca. 2mm long, villous; calyx 17mm long, shaggy-lanate, the subulate-linear teeth 5-6 mm long; corolla red; standard 18-19mm long, the limb oval, retuse, angled at base, the claw 7-8 mm long; wing-petals ca. 14mm long, the limb oblong, the claw 8mm long; keel 14mm long, the limb rounded, strongly convex beneath and slightly so above, the claw 10mm long; ovary on a stipe ca. 1mm long, hairy; style glabrous. Fl. April. River valleys, on pebbles. - Soviet Centr. Asia: Mtn. Turkm. (Badkhyz). Endemic. Described from Kushka River. Type in Leningrad. Series 2. ORBICULARES Gontsch.- Stems long; stipules free, cauline, scarcely united at base with petiole, green, herbaceous; racemes few- flowered, borne on axillary, more or less nodding peduncles; pods thinly coriaceous, oblong, of medium size, carinate ventrally, rounded or flat- tish dorsally, compressed laterally, terete to 1/2-2/3, bilocular, sub- sessile. 94. A. orbiculatus Lbd. Fl. Alt. III (1831) 311; Fl. Ross: I, 623; Bee. Astrag. "geront. Il,” 39; Boiss. Fl "Ori, 278; Bee. Astras. tune. aces Kryl. (Rl. (Zap! Sib: VIE, 1659;"R i Padzhiks Vi"s4— lew sb darter mia emma Ross. Ill, tab. 290. -Exs.: H.F.R. No. 1863; Herb. Fl. As. Med. No. 362. 84 114 Perennial, 20-40cm tall; stems (7) 10-35cm long, branched, becoming lignified at base, villous; stipules ovate, 6-10 (15)mm long; leaves 7-10 (18)cm long, the petiole obsolete, the rachis covered with spreading hairs; leaflets 6-11 pairs, oval, 5-10 (15)mm long, round-tipped, covered sparse - ly above and densely beneath with shaggy pubescence, rarely with scattered hairs on both sides; peduncles slender, 2-4cm long, nodding; racemes 2- 3cm long, loosely 4-7-flowered; bracts linear, 8-12mm long, acute, com- monly hairy; pedicels 1-2 mm long; calyx campanulate, 9-10mm long, more or less shaggy, the tube slightly exceeding to rarely twice as long as the linear teeth; corolla yellow; standard 14-16 mm long, the limb oblong, round -tipped or subobtuse, angular at base, 4-5 times as long as the broad claw; wing-petals 13-15mm long, the limb oblong, silky outside, round- tipped, 1.5-2 times the length of the claw; keel ca. 9mm long, glabrous, obtuse, the limb convex beneath, straight above, equaling the claw; ovary villous, stipitate; pod 14-15mm long, subacute at both ends, villous, the beak 1mm long; seeds compressed, reniform, 2.5mm long, smooth, red- dish. Fl. April-May; fr. May-June. River valleys.-W. Siberia: Irt.; Soviet Central Asia: Aralo-Casp. (shores of AralSea), Balkh. region, Amu D. Gen. distr. : Afghanistan. De- scribed from Bukon River (tributary of Irtysh). Type in Leningrad. Series 3. LASIOPETALI Gontsch. - Plants ranging from long-stemmed to acaulescent or nearly so, mostly reddish-villous; stipules united at base with petiole; racemes many - or few-flowered, compact or loose, sessile or more often distinctly pedunculate, corolla yellow; pods oblong or ovaloid, slightly compressed laterally, carinate and convex ventrally, rather broad- ly grooved dorsally, fairly large, sessile, coriaceous, incompletely biloc- ular. 95. A. lasiopetalus Bge. Ind. Sem. H. Dorp. (1839 et 1841) 7: Lab. Fl. Ross. I, 651; Bge. Astrag. geront. 40; Bge. Astrag. turk. 226; Kryl. Fl. Zap. Sib. VII, 1660.-A. lasianthus C.A.M. in Bong. et Mey. Suppl. fl. Alt. (1841) 27.-? A. ulacholensis B. Fedtsch. in A.H.P. XXIV (1905) 205. -Ic.: Bong. et Mey., l.c., tab. 6 et 7. Perennial, 20-40 (47)cm tall; stems commonly developed, shaggy with white woolly pubescence, (1) 3-15 (24) cm long, the plants rarely subacaules- cent; leaves (11) 15-25 (27)cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis shaggy with spreading hairs; stipules oblong - oval, the upper lance-oblong, sparsely hairy outside, ciliate-margined; leaflets (7) 10-15 pairs, ovate -elliptic or oblong-oval, more or less round- ed at apex, mostly scarcely mucronulate, (12) 16-28 (40)mm long, glabrate above, appressed-villous beneath; peduncles 1-3 (5)cm long, rarely infl. subsessile; racemes compact, glabrate, spherical or ovaloid, 8-12-flow- ered; bracts lanceolate, scarious, hairy on the outside, 8-14mm long; pedi- cels 2-3 mm long, villous; calyx campanulate, villous, (10) 12-15 (18)mm long, the linear teeth twice the length of the tube; standard(14) 15-20 (23)mm long, sericeous on the outside, the short claw 3-4 (5)mm long; wing-petals (11) 13-16 (20)mm long, the limb linear, sericeous on the outside, rounded - obtuse, twice the length of the claw; keel 9-11 (13)mm long, the limb ob- tuse, bifid at apex, hairy, about as long as the claw; ovary subsessile, woolly, witha short hairy style; podovaloid, 12-15mm long, densely villous, the beak 1-1.5mm long; seeds ovaloid, emarginate, compressed, ca. 3mm long, smooth, palebrown. Fl. May-June (July); fr. June-July. (Plate X, Figure 1). 85 be Banks of rivers and canals, meadows, rarely slopes of waterless val- leys; often in rather saline soil. - Soviet Centr. Asia: Balkh. region, Syr- D. (E.), T.Sh. (foothills), Pam.-Al. (foothills). Endemic. Described from Zaisan depression. Type in Leningrad. 96. A. dasyanthus Pall. Spec. Astragal. (1800) 79; Ldb. Fl. Ross.I, 623; Bee. Astrag. geront, Il, 41.—lce-s Pall. spec. Astragal.,, tabwooe Perennial, (10) 16-35cm tall, commonly with well developed rufous - villous stems 4-20cm long, rarely subacaulescent; stipules triangular -lan- ceolate or oblong, 16-23mm long, subulate-acuminate, beset outside with soft rufous hairs; leaves 12-20cm long, the petiole much shorter than the rachis, both covered with spreading, mostly rufous, hairs; leaflets (11) 12- 14 (17) pairs, oblong-oval or lance-oblong, (6) 12-18 (20)mm long, silky- shaggy on both sides with appressed hairs; peduncles 5.5-15cm long, most- ly shorter than the leaves, villous with spreading hairs; racemes compact, capitate, spherical or ovaloid, (6) 10-20-flowered, (2.5) 3.5-6 (6.5) cm long; bracts as long as or slightly shorter than the calyx, lanceolate, acuminate, covered with reddish hairs; pedicels 1-1.5mm long; calyx campanulate, villous, (12) 13-15 (17)mm long, the subulate-linear teeth 5-6 (8)mm long; standard (17) 18-25 (26)mm long, lanate on the outside, the broad claw (4) 5-7 (8)mm long; wing-petals (13) 15-18 (19) mm long, the limb oblong, round-tipped or scarcely retuse, lanate onthe outside andinupper part; keel (12) 13-15 (17) mm long, the limb obtuse, hairy, convex beneath, more or less straight above, 1.5 times the length of the claw; ovary long-villous; style hairy in lower part, 10-11 mm long, rarely villous, round-based, sub- truncate or Subacute, the beak 2-3mm long. Fl. June. steppes. - European part: U. Dnp. (S.), M. Dnp., U. Don. (S.); Bes. ; Bl., L. Don, L.V. (N.). Endemic. Described from the vicinity of Tsari- tsyn (Stalingrad)*. Type in Leningrad. 97. A. tanaiticus C. Koch in Linnaea XXIV (1851) 94; Bge. Astrag. geront. II, 59. Perennial, acaulescent plants, 15-27cm tall; stipules oblong-ovate or lance-oblong, up to 18mm long, rufous-villous; leaves 15-27cm long, the petiole 4-9cm long, both petiole and rachis shaggy with rufous woolly pu- bescence; leaflets 12-18 pairs, ovate to oblong-ovate, round-tipped or sub- acute, mucronulate, beset with long appressed hairs onboth sides, (12)15-25 (30)mm long; racemes oblong, 5-10 (12) cm long; bracts lance-linear, long- acuminate, thinly scarious, 8-11mm long, rufous-villous outside; pedicels 2-2.5 mm long, shaggy with woolly reddish hairs; calyx cylindric-cam- panulate, 13-18 mm long, densely rufous-villous, the linear acute teeth o9-7mm long; standard 22-28 mm long, the limb retuse, angular at base, sericeous outside, the claw 9-12mm long; wing-petals 18-22 mm long, the limb linear-oblong, round-tipped, lanate outside in upper part, 9-12 mm long; keel glabrous, 14-16mm long, the limb convexbeneath, nearly straight above, subobtuse, slightly bifid, the claw 7-9mm long; ovary sessile, rufous-lanate; style hairy (glabrous beneath the stigma); pod oblong -oval- oid, ca. 15mm long, rufous-villous, the beak 2-3mm long. Fl. May-June; fr. June (July). Steppes, in chernozem. - European part: L. Don, Bl. Endemic. De- scribed from the Don. Cotype in Leningrad. * [Now Volgograd. ] 86 NOD yy) b aE ya, a Le AS Eo\\ AN a f ‘ " ey if PLATE X 118 119 98. (At pubiflorus’ DSC: Astrag. (802) "7 3; db. Ele Ross. 1.) G04: Bge. Astrag. geront. II, 59.-Ic.: Pall. Astrag. tab. 64, sub nom. A. ex - capo. Perennial, acaulescent plants, 10-25cm tall; stipules oblong or lanceo- late, acute or acuminate, 12-26mm long, more or less covered outside with reddish hairs, ciliate-margined; leaves 6-25cm long, the petiole (1.5) 2-16 cm long, petiole and rachis villous with spreading reddish hairs; leaf- lets 8-14 pairs, oblong-oval or oblong-ovate, rarely oblong, round-tipped and mucronulate, (7) 9-25 (33) mm long, covered on both sides with ap- pressed long rather scattered reddish hairs, sometimes sericeous above; peduncles mostly none, rarely up to 2 (4) cm long; bracts subulate-linear, long-acuminate, ciliate, 10-20mm long; pedicels 3-6 mm long, hairy; calyx cylindric-campanulate, 18-25mm long, shaggy with long reddish hairs, the subulate-linear teeth equaling the tube; standard (19) 23-30mm long, the limb oblong, retuse, slightly angular at base (rarely subobovate), more or less pubescent, the claw 7-10mm long; wing-petals (16) 20-24mm long, the limb linear-oblong, round-tipped, hairy, equaling the claw; keel (12) 16-20mm long, glabrous, the limb convex beneath, above convex apically and straight downward, equaling the claw; ovary subsessile, lanate; style hairy below; pod oblong-ovaloid or oblong, subsessile, 13-15 mm long, rufous-villous, the straight beak 2-3mm long. Fl. April-June; fr. May-July. Steppes. - European part: L. Don, Bl., Crim. (N.). Endemic. Described from Siberia* (?). Type in Geneva. Series 4. ANISOMERI Gontsch.- Plants acaulescent, very rarely short- stemmed; stipules scarious, united at base with petiole; racemes loose, many-flowered, short-peduncled (peduncles 0.5-2-3cm long); corolla yel- low; limb of wing-petals 6-8 times the length of the claw and 3-4 times as long as the keel; pod sessile, oblong, carinate ventrally, broadly grooved dorsally, coriaceous, imperfectly bilocular. 99. A. anisomerus Bge. Astrag. turk. (1880) 228. Perennial, 7-23 (30) cm tall, from a short, branched, buried caudex, acaulescent or nearly so; stems obsolescent or strongly reduced, 1.5-3cm long, angled-sulcate, densely hispid with spreading hairs; stipules united with petiole at base, membranous, whitish, 10-15mm long, densely ciliate - margined, the lower triangular-ovate, the upper oblong in the free portion, acute or acuminate, rounded at apex; leaves 7-20 (30) cm long, the rachis many times, rarely 3 times the length of the petiole, both rachis and petiole » densely covered with reddish and yellowish hairs, sometimes almost woolly, , stiffish; leaflets (10) 16-25 pairs, oval or oblong-ovate, (5) 7-12 (20)mm long, (2.5) 4.5-7 (10)mmbroad, obtuse, often mucronulate, beset on both sides with appressed, silky, rather dense, ascending, hairs; peduncles obsolescent, 0.5-1 (3) cm long, covered with rather dense, spreading, stiff hairs; racemes short, lax, 2-2.5mm [sic] long, (rachises 5-7 mm long), (3) 5-7 (10)-flowered; bracts linear or lance-linear, 7-1l1mm long, acumi- nate, scarious, rather densely long-ciliate on the margin; pedicels 3-6mm long; calyx infundibular-campanulate, (8) 10-12mm long, white-villous with | spreading hairs, the teeth narrowly linear, finely pointed, equaling the tube, rarely 1.5 times as long; corolla yellow; standard (15) 20-26 mm long, the limb sericeous outside, oblong-obovate, in upper 2/3 obcordate, narrowed * The report for Siberia by the author of the species is probably erroneous. 88 by half below, slightly angular at base, 6-8 times as long as the claw; wing- petals (13) 16-20mm long, the limb sericeous outside, narrowly linear, entire, obtuse, ca. 2mm broad, 5-6 times the length of the claw; keel 6- 8mm long, the limb woolly-villous at apex, somewhat asymmetrically oblong-ovate, strongly attenuate and short-cleft at apex, obtusish; 2.5times the length of the claw; ovary raised on a very short stipe, woolly-villous; pods sessile, semioval-oblong in outline, 12-16mm long, 6mmbroad,ter- minating in a reflexed beak ca. 1mm long, slightly inflated, carinate ven- trally, grooved dorsally, coriaceous, shaggy with soft ascending hairs, semibilocular. Fl. June-July; fr. July-August. Stony and gravelly mountain slopes from 1000 m and up to the Subalpine zone. - Soviet Centr. Asia: T.Sh. (W. — from S. Karatau (Burnoe) to Talas- skii Ala Tau and Korzhantau mountains). Described from the vicinity of Tashkent. Type in Paris. Series 5. SONGORICI Gontsch.- Acaulescent, loosely cespitose plants; stipules scarious, united at base with petiole; racemes loosely few-flowered, short-peduncled; corolla sordid; pods sessile, oblong-ovaloid, slightly compressed, carinate, thickened or scarcely concave dorsally, nearly fully bilocular, coriaceous. 100. A. Nicolai Boriss. nom. nov.-A. songoricus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Acad. Sc. URSS, t. VII, fasc. 7 (1938) 152, non JEU, Sikes 2 Maen) uilisys)4 Perennial, 15-28 (35) cm tall, from a short, woody, branched, buried caudex; stipules dimorphic, the lower ovate or oblong more or less round- tipped, the upper lanceolate, acute, (6) 10-16 (20)mm long, hairy on the outside; leaves (7) 15-29 (39) cm long, the rachis 2-4 times the length of the spreading-villous petiole; leaflets (14) 15-22 (29) pairs, oval or elliptic, obtuse or subacute, (8) 10-18 (24) mm long, covered on both sides with ap- pressed reddish silky hairs; peduncles 1.5-3 (6) cm long, together with infl. much shorter than the leaves, (2) 3-6 (7)-flowered; bracts lance-linear, 7- 12 (20)mm long, scarious, acute, white-ciliate; pedicels 3-5 (7) mm long, villous; calyx cylindric, 13-17 (19) mm long, villous, the teeth (3) 4-6 (9)mm long; standard (20) 23-28mm long, sericeous on the outside, the oval or oblong-oval limb twice the length of the claw; wing-petals 19-23mm long, the oblong sericeous round-tipped limb about equaling the claw; keel glabrous, 15-18mm long, the subtriangular acutish limb 1.5 times the length of the claw; ovary sessile, sericeo-lanate with white hairs; style hairy; pod 15-17 mm long, rather sparsely white -villous. Mountain slopes. - Soviet Centr. Asia: Dzu-Tarb. (Dzungarian Ala Tau). Endemic. Described from Dzungarian Ala Tau. Type in Leningrad. 120 Series 6. CAESPITOSI Gontsch. - Acaulescent low plants, forming rather compact tufts. Stipules scarious, united at base with petiole; ra- cemes short, 2-4-flowered, subsessile, subradical; corolla yellow; pods ovaloid or oblong-ovaloid, carinate ventrally, rounded dorsally, imperfect- ly unilocular (with a narrow septum not more than 3mm broad), sessile, coriaceous. 101. A. sisyrodites Bge. Astrag. geront. II (1869) 40; Astrag. turk. (1880) 227. 89 121 Perennial, 7-12cm tall, from a strong, woody, much-divided, buried caudex; stipules oblong, acute, 6-11mm long, faintly hairy in upper part, ciliate; leaves (4) 6-12 cm long, the petiole firm, persistent, 2-4 (5) cm long, petiole and rachis villous; leaflets 8-14 pairs, villosulous on both sides; flowers subsessile, partly concealed at the bases of crowded leaves; racemes borne on peduncles not more than 3mm long, 3-4-flowered, the infl. rachis not exceeding 7mm, long-hairy; bracts narrowly lance-subulate or linear-subulate, hyaline, white-villous outside, 10-15mm long; calyx cylindric-campanulate, 10-15mm long, densely villous, the teeth setiform, feathery-ciliate, (5) 8-10mm long; standard (13) 15-17mm long, the limb oblong-quadrangular or oblong, slightly angular at base, the claw 4-6mm long; wing-petals (11) 13-15mm long, the limb linear-oblong, round-tipped, sericeous cutside in upper part, the claw (3) 4-5mm long; keel (6) 9-10mm long, the limb concave beneath, slightly incurved above, bifid and ciliate at apex, the claw (2) 4-5mm long; ovary subsessile, villous; style hairy nearly throughout; pod ovaloid or oblong-ovaloid, ca. 11mm long, white- villosulous, the beak ca. 4mm long. Fl. May-June. Gravelly and stony mountain slopes. - Soviet Centr. Asia: T. Sh. (Karatau range). Endemic. Described from Karatau Mountains. Type in Leningrad. 102. A. Tekutjevii Gontsch. in Not. Syst. ex Herb. Inst. Bot. Acad. Sc. URSS VII, 7 (1938) 148. Perennial, ca. 16cm tall; stipules lanceolate, ca. 15mm long, strigose; leaves 9-14cm long, the petiole and the much longer rachis indurated per - sistent hispid; leaflets 10-14 pairs, lanceolate acute; bracts narrowly line - ar, 5-7mm long, acuminate, scarious, hispid outside and on the margin; pedicels 2mm long; calyx campanulate, 14-15 mm long, the tube glabrous, the teeth filiform, long-villous, unequal, the longer 7-9mm long; standard 14-15 mm long, the limb oblong, scarcely retuse, gently angular at base, the claw 4-5mm long; wing-petals 13-14.5mm long, stipules linear, round- tipped, lanate outside in the upper half, the claw 4-5.5mm long; keel 10- 10.5mm long, glabrous, obtusish, cleft at apex, the claw 5.5mm long; ovary subsessile, barbate-woolly; style sparsely hairy; pod oblong-oval, ca. 12mm long, more or less white-villous. Stony and gravelly mountain slopes. - Soviet Centr. Asia: T.Sh. (Karatau range). Endemic. Described from Karatau Mountains. Type in Leningrad. Series 7. HOLOSERICEI Gontsch. - Acaulescent velutinous plants; stip- ules membranous, united with petiole in lower part; racemes compact, capitate, many-flowered, short-peduncled; corolla yellow; pods sessile, rigidly coriaceous, ovaloid, slightly compressed laterally, inflated, cari- nate ventrally, rounded subcarinate dorsally, imperfectly bilocular. 103. A. holosericeus M. Pop. In Append. ad. Fl. Tadzhik. V (1937) 662. Perennial, 35-46 cm tall; stipules oblong or lanceolate, acute or short- acuminate, 20-25mm long, densely villous outside; leaves 20-33 (41)cm long, the petiole 9-13 (17) cm long, petiole and rachis villous; leaflets 9- 13 pairs, oblong-ovate, acute; flowers up to 3.5 (4) cm long, white -villous; bracts linear-filiform, 10-15mm long, white-villous; pedicels not exceed- ing 1mm, villous; calyx cylindric-campanulate, 14-16mm long, densely white-villous, the linear-filiform teeth equaling to 1.5 times the length of the tube; standard 19-20mm long, the limb oval, rounded-obtuse, much 90 1122 longer than the claw; wing-petals 17-18 mm long, the limb oblong, silky- villous outside, round-tipped, equaling the claw; keel 15-16mm long, the limb oblong-lanceolate, rounded at summit, deeply cleft, equaling the claw; ovary Sessile, densely white-lanate; style glabrous; pod 15-16mm long, densely villous with long hairs, the beak 3-4mm long. Fl. June-July; fr. July-August. Rosaria and Exochorda. - Soviet Centr. Asia: Pam.-Al. Endemic. De- scribed from vicinity of Kangurt (Tadzhikistan). Type in Leningrad. Series 8. AMYGDALINI Gontsch. -Long-stemmed or acaulescent plants; stipules scarious, united at base with petiole; racemes loosely few -flowered, short- or long-peduncled; corolla yellow; pods short- or long-stipitate, asymmetrically subquadrangular-ovaloid, compressed laterally, inflated up- ward, rigidly carinate ventrally with thickened suture, slightly grooved or rather flattened dorsally, rigidly stony-coriaceous, bilocular or nearly so, covered with appressed short hairs. 104. A. Litvinovianus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Acad. Sea URS) VIEX fase. 7) (1 938) 42). Perennial, (20) 35-50cm tall, the stems (8) 22-32cm long, with shaggy pubescence; stipules linear-lanceolate from a broad base, subacute, 12- 17mm long, sparingly hairy outside, white-ciliate on the margin; leaves (12) 16-25cm long, the petiole obsolete, the rachis villosulous; leaflets (10) 15-20 pairs, oblong-oval or oblong-ovate, (15) 18-25 (30)mm long, round- tipped, rarely subacute, mucronulate, sparsely covered on both sides with appressed hairs; peduncles axillary, together with infl. only slightly ex- ceeded by the leaves, 6-11cm long excluding infl., peduncle and inflores- cence villosulous with spreading hairs; racemes remotely 5-7-flowered; bracts linear-subulate, 7-9mm long, scarious, more or less hairy outside, ciliate-margined; calyx short-cylindric, 15-20mm long, villous, the teeth lance-linear, subulate-tipped, unequal, the longer 10-12 mm long; standard- limb oblong, slightly constricted at the middle, deeply emarginate, angular at base, sericeous outside, the claw 5-6mm long; wing-petals 23-24mm long, the limb linear, scarcely enlarged upward, round-tipped, outwardly sericeous upward, the claw 7mm long; keel 15-16 mm long, the limb slight - ly bifid at apex, sericeous and ciliate-margined, slightly longer than the claw; ovary silky-villous with appressed hairs; style sparsely hairy; pod 22-25mm long, on a stipe 7-9mm long, grooved dorsally, nearly fully bilocular, the beak 3-3.5mm long. Fl. May-June; fr. June-July. (Plate X, iitoe2 ye Meadows and forest glades in the lower part of the walnut zone in eastern Fergana; of isolated occurrence in Kirgiz Ala Tau. - Soviet Centr. Asia: T.Sh. (S.W. slopes of Fergana range and Kirgiz Ala Tau). Endemic. De- scribed from the vicinity of Khazret-akob in Andizhan district. Type in Leningrad. 105. A. amygdalinus Bge. Astrag. turk. (1880) 228.-Exs.: Herb. Fl. As. Med. (1928) No. 351. Perennial; related to the preceding, distinguishing characters being: leaflets 12-14 pairs, mostly smaller, 10-17 (18)mm long, peduncles to- gether with inflorescence shorter than the leaves, flowers smaller, pods smaller and flattened dorsally, (16) 18-21 mm long, the stipe 2.5-3 mm long. oi WY, UuEE SERS MS PREY > JMR Wc Oo LL Alba We de ge ae AR PLATE XI 1, Astragalus grandiflorus Bge.—2. A. Korolkovii Bge. 125 Upper layer of the bluegrass-sedge ephemeroid meadows (montane semi- deserts) and the quack-grass steppe zone. - Soviet Centr. Asia: T. Sh. (the western part, foothills from Chimkent to Tashkent). Endemic. Describedfrom the vicinity of Tashkent. Type in Paris. 106. A. pseudoamygdalinus M. Pop. in Appendix Fl. Tadzhik. V (1937) Bola wes: bis ladznik. Vi tal.3 1. Perennial, acaulescent or nearly so, (13) 15-25cm tall; stipules dimor- phic, the lower triangular-ovate, the upper oblong-triangular -ovate, acute, 7-10mm long, ciliate; leaves (12) 18-22 (26)cm long, the petiole (1.5) 3-4cm long, firm, shaggy with woolly white hairs; leaflets (10) 13-15 pairs, elliptic to ovate-elliptic, round-tipped or subacute, mucronulate, (10) 14- 25mm. long; peduncles 1-3 (4)cm long, slender, villous; racemes (5) 10- 13cm long, very remotely (5) 8-9-flowered; bracts scarious, acute, 6- 9mm long, villous outside; pedicels 2-3 mm long, villous, the petiole 11- 14mm long, woolly-villous, the teeth linear-subulate, unequal, the longer 7-9mm long; standard sericeous outside,the limb oblong, scarcely con- stricted at the middle, (3) 5-6 mm long; wing-petals 15-16 mm long, the limb linear-oblong, round-tipped, sericeous outside in upper part, the claw 4-4.5 mm long; keel 11-12 mm long, obtuse, cleft at apex, the claw 4mm long; ovary short-stipitate, white-villous; style hairy nearly throughout; pod 16-21 mm long, 9-11mm broad, shaggy-pubescent with more or less appressed hairs, imperfectly bilocular, the stipe 2-3mm long, the beak 1.5-2 mm long. Fl. June-July; fr. July. Zone of grasS-wormwood mountainous semidesert.- Soviet Centr. Asia: T.Sh. (Angren river basin, Kuraminskii range). Endemic. Described from Kuraminskii range. Type in Leningrad. 107. A. juniperetorum Gontsch. in Not. Syst. ex Herb. Inst. Bot. Acad. Se. URSS, VII, fase. 7 (1938) 145. Perennial, acaulescent or nearly so, 29-30 cm tall; stipules glabrate, the lower oval round-tipped 25-18 [sic] mm long, the upper narrowly trian- gular-lanceolate, long-acuminate, 18-20 mm long; leaves 20-32 cm long, the petiole much shorter than the rachis, soft-woolly; leaflets (8) 10-21 pairs, ovate or broadly oval, obtuse or scarcely retuse, (11) 13-20 mm long; peduncles 8-15mm long, lanate; racemes loosely 4-6-flowered; bracts 8-13 mm long; pedicels 3-5mm long, lanate; calyx ca. 2cm long, villous, the teeth unequal, the longer equaling the tube; standard sericeous outside, the limb oblong retuse angled at base, the claw ca. 6mm long; wing-petals ca. 23mm long, the limb linear-oblong, round-tipped, sericeous, slightly longer than the claw; keel 20mm long, round-tipped, the claw 10-11mm long; ovary on a stipe ca. 1mm long, densely white-lanate. Fl. May. Juniper woods. - Soviet Centr. Asia: Pam.-Al. (Uzbekistan, vicinity of Baisun). Endemic. Described from Ketmen'-Chapty Mountains. Type in Leningrad. Note: Inclusion of this species in the series Amygdalinae is pro- visional, as its fruits are unknown. Series 9. GRANDIFLORI Gontsch. - Long-stemmedor acaulescent plants; stipules scarious, briefly united with petiole; flowers 2-4 on short pe- duncles; leaflets glabrous above; corolla yellow or pinkish-white; standard sericeous; wings and keel glabrous; pods ovaloid or oblong, more or less compressed, carinate ventrally, grooved dorsally, coriaceous, partially bilocular, stipitate. 93 126 127 108. A. grandiflorus Bge. Astrag. turk. (1880) 229; Fl. Tadzhik. V, Sail Perennial, acaulescent, (8) 10-15 (35) cm tall, rarely with stems up to 20cm long; stipules oblong or lanceolate, rarely oblong-oval; leaves (6) 8-14cm long, the short petiole and the rachis villous with spreading hairs; leaflets 10-12 pairs, ovate-elliptic, (6) 8-12mm long, 5-8mm broad, sub- obtuse and often scarcely retuse, glabrous above, covered beneath with soft appressed hairs, densely long-ciliate on the margin; peduncles 5-12 mm long, 2-3-flowered; bracts linear, villous, exceeding the pedicels, these 2-3mm long; calyx narrowly long-cylindric, (15) 20-25mm long, ca. 3mm broad, villous with long soft hairs, the tube 2.5-3 times the length of the lance -subulate teeth; standard (30) 35-40mm long, the limb obovate -oblong, dilated in upper part, oblong-attenuate in lower part, angled at base, the claw 6-8 mm long; wing-petals 28-35mm long, the limb shorter than the claw, gently incurved at apex, cultriform, acute, the claw 1.5 times as long as the limb; keel 27-34mm long, the limb convex beneath, nearly straight above, subacute, about half the length of the claw; ovary on a stipe 5-6.5mm long, silky-villous; pod on a stipe 4-5mm long, ovaloid to ovaloid-oblong, 15-20mm long, mucronulate, villous. Fl. April-May (June); fr. June. (Plate XI, fig. 1). An ephemeroid perennial, associated with submontane plains and foot- hills, in bluegrass-sedge ephemeroid meadows (loess desert and, in the south, semidesert).- Soviet Centr. Asia: Syr. D. (western part). Pam.-Al. (from Kulyab to Kurgan-tyube). Endemic. Described from the vicinity of Tashkent. Type in Leningrad. 109. A. rubromarginatus E. Czernjak. in Not. Syst. Herb. Hort. Bot. Petrop. II (1921) 69.-A. oeroilanicus M. Pop. in Not. Syst. Herb. Hort. Bot. Petrop. IV (1923) 156, ex p. Perennial, closely related to the preceding; distinguished by smaller flowers, 27-30 (35) mm long, whitish, at length rose color, the broadly cylindric saccate-based calyx, and the shape of pod; plants with stems up to 30cm long, acaulescent or nearly so; leaflets 12-16 pairs, obovate or rarely orbicular, 5-10mm long, 5-10mm broad, subobtuse or slightly re- tuse; calyx ca. 20mm long, 4-5mm broad, villous with long spreading hairs, the tube equaling or 1.5 times the length of the lance-linear acumi- nate teeth; pods semioval-oblong in outline, carinate ventrally, grooved dorsally, (1.5) 2-2.5cm long, 0.7-0.9cm broad, rather sparsely white - villous, mucronulate, the thick stipe 2-3mm long. Fl. April-May. Sandhills and consolidated sands, submontane desert plains, and loess, often gypseous, foothills. - Soviet Centr. Asia: Kyz. K. (eastern border), Mtn. Turkm. (W. Kop. D.), Pam.-Al. (Guzar District). Endemic. De- scribed from western Turkmenia (Songu-dag Mts. and surroundings of Dzhalila station). Type in Leningrad. 110. A. oeroilanicus M. Pop. in Not. Syst. ex Herb. Hort. Petrop. IV. (1923) 156.-A. rubromarginatus var. oeroilanicus M. Pop. in Sched. ad Herb. Fl. As. Med. fasc. XV (1928) No. 365.-Exs.: H. B. A.M: No. 365. Perennial, 13-35cm tall; stems obsolescent, 2-3 cm long, or up to 28cm long, thinly white-villous with spreading hairs; stipules united in lower part with petiole, 5-6mm long, hyaline, thinly villous with spreading hairs, the lower triangular -ovate, the upper lanceolate, acuminate; leaves 94 (8) 10-15 cm long, the rachis 3-4 times as long as the petiole, both petiole and rachis thinly villous with spreading white hairs; leaflets 13-16 pairs, obovate to suborbicular, retuse, 4-8mm long, 3-6mm broad, glabrous above, covered beneath with soft spreading hairs; peduncles greatly exceed- ed by the leaves, 0.5-2cm long, thinly covered with spreading white hairs; racemes loose, short, 3-4cm long, 4-8-flowered; bracts lance-linear, 6- 8mm long, acuminate, hyaline, white-villous, 1.5-2 times the length of the pedicel; calyx short-cylindric, 11-15mm long, ca. 3mm broad, white- villous with soft spreading hairs, the tube 1.5-2 times as long as the lance- linear, finely pointed teeth; corolla pinkish-white; standard 19-22 (24)mm long, the limb oblong-obovate, scarcely retuse, 7-8 (9)mm broad, rather abruptly angled-attenuate at base, 2.5-4 times as long as the claw; wing- petals 18-22 mm long, the limb linear-oblong, entire, round-tipped, equaling or shorter than the claw; keel 14-15 mm long, the limb ob- long, acutish, slightly concave beneath, as long as the claw; ovary on a stipe 3-4mm long, densely shaggy with woclly pubescence; pods on a stipe 2-3mm long, spreading to often subcernuous, semioval-oblong in out- line, subobtuse, 20-25mm long, ca. 8mm broad, terminating in a subulate, reflexed, indurated beak, 2.5-4mm long, carinate ventrally, grooved dor- sally, compressed laterally, coriaceous, thinly villous with white spreading hairs, bilocular. Fl. April; fr. (April) May. Consolidated sands and sandhills. - Soviet Centr. Asia: Kara K. (south- east), Mtn. Turkm. (Badkhyz). Endemic. Described from the vicinity of Lake Eroilan. Type in Tashkent. Series 10. TUBULATI Gontsch. - Plants with long stems or subacaules- cent; stipules scarious, united at base with petiole; upper face of leaflets white-floccose, rarely glabrate or glabrous; racemes small-flowered, loose, short-peduncled; calyx tubular; corolla yellow, all the petals pubescent outside; pods oblong, carinate ventrally, grooved or rounded dorsally, coriaceous, short-stipitate, ranging from subbilocular to unilocular. 111. A. lanuginosus Kar. et Kir. in Bull. Soc. Nat. Mosc. XIV (1841) 409; Bge. Astrag. geront. II, 41; Bge. Astrag. turk., 231, ex parte; Ldb. Fl. Ross.1, 651.-A. mucidus Bge. Astrag. turk. (1880) 230 ex parte. - 28 A. mucidiformis Sumn. in Animadv. Syst. ex Herb. Univ. Tomsk. fasc. 9-10 (1926) 2.-A. larvatus Sumn., l.c., 3. Perennial plants with stems (5) 13-15 (29) cm long, rarely subacaules- cent; stems lanate with spreading hairs; stipules lanceolate, finely long- pointed, 7-10mm long, villous outside; leaves (7) 10-14 (16) cm long, the very short petiole and the rachis rather thinly villous with spreading hairs; leaflets (16) 20-25 (26) pairs, oval to oblong-oval, (4) 5-11 (12) mm long, subobtuse or scarcely retuse, the upper surface floccose, rarely glabrous or nearly so; peduncles (0.3) 0.5-1.5 (2) cm long, spreading-villous; ra- cemes short, 2-5 (6)-flowerei; bracts linear, acuminate, 5-6 (7) mm long, villous outside; pedicels (3) 4-5 (6) mm long, villous; calyx 15-23 mm long, the teeth subulate from a triangular base, (4) 5-8 mm long; standard white- lanate, scarcely retuse, the claw about half the length of the limb; wing- petals 18-25mm long, the limb white-villous, round-tipped or scarcely retuse, about twice the length of the claw; keel 14-22 mm long, the limb obtuse or subobtuse, more or less white-villous at apex, the claw 8-13mm 95 long; ovary long-stipitate, white-villous; style hairy in lower part; pods oblong, 14-20mm long, more or less inflated, carinate ventrally, broadly and shallowly grooved dorsally, white-villous, terminating in a beak (1) 2-3mm long, acute at base, borne on a stipe 2-5mm long, almost fully bilocular. Fl. June-July; fr. July-August. Plains, foothills, mountains, wormwood desert and semidesert, con- solidated sands, occasionally a weed among crops, and along river banks.- Soviet Centr. Asia: Balkh. region, T. Sh. (northern foothills), Dzu.- Tarb. (foothills). Endemic. Described from the rivulet Burgan at the foot of Tar- bagatai. Type in Leningrad. 112. A. mucidus Bge. Astrag. turk. (1880) 44; Fl. Tadzhik. V (1937) 339.-A. serafschanicus Freyn. in Bull. Herb. Boiss. IV (1904) 767. Perennial, with developed densely villous stems 3-20 (31) cm long (rarely subacaulescent), (9) 12-25 (35) cm tall; stipules lanceolate or triangular - lanceolate, 6-7 mm long, acuminate, white-villous; leaves 9-15 (18) cm long, the petiole very short; leaflets (14) 18-26 pairs, oblong-oval to oval, 5-7 (12) mm long, floccose above; peduncles 1-1.5 cm long, villous; ra- cemes (2) 3-6 (7)-flowered; bracts subulate-linear, 4-5 (6) mm long, equal- ing or barely exceeding them [?], white villous outside; pedicels (2) 3-4mm long, villosulous; calyx cylindric, 12-15mm long, densely villous, the tube slightly exceeding the linear teeth (rarely 1.5 times as long); standard lanate outside, retuse, the claw (6) 7-9mm long; wing-petals 13-16 mm long, the limb more or less hairy outside, somewhat irregularly emarginate, as long as or scarcely longer than the claw; keel 12-14mm long, the claw as long to 1.5 times as long as the hairy limb; ovary long-stipitate; pod ovaloid or 129 oblong-oval, 8-9 (10) mm long, borne on a stipe 1-2 mm long, terminating in a beak ca. 1mm long, inflated, thickened or scarcely grooved dorsally, white-villous, partially bilocular. Fl. April-June; fr. May-July. Submontane plains and foothills, in the zone of bluegrass-sedge, ephem - eroid meadows (loess desert and semidesert). - Soviet Centr. Asia: T. Sh. (west from Chimkent to Tashkent and Mogoltau), Pam.-Al. (west from Samarkand to Dzhizak). Endemic. Described from Mogoltau. Type in Leningrad. 113. A. floccosifolius Sumn. in Anim. Syst. ex Herb. Univ. Tomsk., fasc. 9-10 (1936) 2.-A. pseudolanuginosus Gontsch. in Append. ad. Fl. Tadzhik. V (1937) 657. Perennial, 10-25 cm tall, with short densely villous stems 1-8 cm long, rarely subacaulescent; stipules lanceolate or (the lower) triangular -lanceo- late, 7-10 (11) mm long, villous outside; leaves (5) 10-19cm long, the peti- ole much shorter than the rachis; leaflets 18-25 (30) pairs, oval-oblong or oblong-elliptic, rarely ovate or lanceolate, round-tipped, (5) 6-10 (13)mm long, the upper face white floccose, at length oftenglabrescent; peduncles (0.5) 1-2.5 (4) em long, villosulous; racemes (3) 4-6 (10)-flowered; bracts subulate-linear, (3) 4-6 mm long, villous; flowers 3-5mm long, villosu- lous; calyx (12) 13-15 mm long, densely villous, the tube 1.5 times the length of the linear teeth; standard (18) 19-23mm long, the limb lanate, scarcely retuse, the claw (7) 8-10mm long; wing-petals (17) 18-22mm long, the limb oblong, scarcely dilated in upper part, retuse, lanate, about equaling the claw; keel 12-17 mm long, obtuse, lanate, the claw 7-11mm long; ovary long-stipitate; pod oblong, (11) 12-16 mm long on a stipe 1.5- 3mm long, more or less grooved dorsally, white-villous, bilocular through 1/5-2/3. Fl. June-July; fr. July-August. 96 Foothills, high-mountain desert and desert-steppe valleys. - Soviet Centr. Asia: T.Sh. (eastern part of Fergana kettle hole and central part), Pam.- Al, (Alai range and Surkhob river valley). Endemic. Described from west- ern Tien Shan (southwestern slopes of Fergana range). Type in Tomsk. 114, A. ephemeretorum Gontsch. in Append. ad Fl. Tadzhik. V (1937) 658. Perennial, related to the preceding and replacing it in the central part of western Pamir-Alai. The entire plant less hairy: stems vested with short spreading hairs; calyx covered with appressed soft hairs; wings and keel thinly hairy, sometimes glabrous; leaflets lanceolate to elliptic -lanceolate, rarely oblong, (17) 20-35 (43) pairs, (6) 10-14 (16) mm long; pods oblong, (12) 14-17 mm long, rounded dorsally (rarely scarcely carinate), more or less villous, bilocular through 1/2-2/3. Fl. May-June; fr. June-July. Foothills and more rarely the intermediate mountain zone; among ephem- eral and rarely woodland and scrub vegetation at 900-2300 m alt. - Soviet Centr. Asia: Pam. -Al. (from Kulyab to Stalinabad and Karatag). Endemic. Described from the vicinity of Stalinabad. Type in Leningrad. 115. A. ochranthus Gontsch. in Appendix ad Fl. Tadzhik. V (1937) 659. Perennial, suffruticose, 10-14 (15) cm tall, from a short, woody caudex with short, diffuse, erect or ascending ligneous braches, densely ciothed in upper part in old petioles and stipules, forming loose tufts; annual stems 1-2cm long, villous, concealed by approximate leaves; stipules 6-8 mm long, ovate or oblong (the upper often lanceolate), acute, subapressed - lanate; leaves 5-11 (12) cm long, the very short petiole and the rachis sparsely villosulous with spreading hairs; leaflets (15) 20-34 pairs, oval, rarely broadly lanceolate, 3-6 (9) mm long, subacute, white-floccose above; peduncles short, 0.8-1cm long, villous; pedicels 2-4mm long, villosulous; calyx 13-15 (18) mm long, densely villous-hirsute with subappressed hairs, the teeth linear-subulate; standard 17-22 mm long, more or less silky, re- tuse, the claw 6-9 mm long; wing-petals 15-19 mm long, the limb linear- oblong, subretuse, slightly shorter than the claw, rarely about as long; keel 13-16 mm long, the claw 1.5 times the length of the glabrous obtuse limb; ovary long-stipitate, appressed-villous; style short -hairy below; pod oval- lanceolate in outline, 11 mm long, inflated, subacute, dorsally flattened, white-villous, unilocular, the beakca. 2mm long, the stipe ca. 2mm long. FI. May-June. Low mountains (1200-1800 m alt.), on gravelly slopes. - Soviet Centr. Asia: Pam.-Al. (Babatag, Karatau and Khodzha Mumyn mountains). Endem- ic. Described from Karatau mountains (S. Tadzhikistan). Type in Lenin- grad. Series 11. CAMPANULATI Gontsch. - Short-stemmed or acaulescent plants; stipules scarious, united at base with petiole; leaflets white-floccose above; flowers small in few-flowered, short-sessile or short -peduncled racemes; corolla yellow; standard pubescent; wings and keel glabrous; calyx campanulate or tubular-campanulate, the long teeth equaling or slight - ly shorter than the tube; pod ovaloid or oblong-ovaloid, borne ona very short stipe or subsessile, carinate ventrally, flattened, rounded or carinate dorsally, partially bilocular or unilocular, coriaceous, more or less com- pressed laterally. 97 N31 116. A. Korolkovii Bge. Astrag. turk. (1880) 230; Fl. Tadzhik. V (1937) 342. Perennial, 10-17 (22) cm tall, loosely cespitose, acaulescent or with short (3-6cm long), sparsely villous stems; stipules lanceolate, 8-11mm long, long-acuminate, villous; leaves 5-12 (17) cm long, the very short petiole and the rachis villosulous, soft; leaflets (11) 14-22 pairs, oblong- oval or oblong, 4-10 mm long, round-tipped or subobtuse, covered beneath withappressed hairs; peduncles 0.7-1.5 (2.0)cm long, hairy; racemes short, (3) 5-8-flowered; bracts linear-subulate, 3-5 mm long, hirsute with spreading hairs; pedicels 2-4 mm long, hairy; calyx cylindric-campanulate, 8-10 (12) mm long, hirsute with spreading hairs, the teeth linear-subulate; stand- ard 11-13mm_long, obovate, scarcelyretuse, the claw 3-4(5)mm long; wing- petals 9-12 mm long, the limb linear-oblong, scarcely incurved, scarcely retuse, twice as long as the claw; keel about equaling the wings, obtusish, the claw 3.5-4mm long; ovary short-stipitate, white-villous; pod ovaloid to oblong-ovaloid, ca. 8mm long, flattened dorsally, thinly coriaceous, sparse - ly white-villous, bilocular somewhat beyond the middle, 3-4-seeded, the stipe and the beak ca. 1.5mm long. Fl. May-June (August); fr. June-July (August). (Plate XI, fig. 2). Low and intermediate mountain zones; witch-grass steppes, fescue- needlegrass dry steppes, thinned-out juniper woods. - Soviet Centr. Asia: Pam.-Al. (western - chiefly in Zeravshan basin), Mtn. Turkm. (western Kop. D.). Endemic. Described from Dzhamskii canyon. Type in Leningrad. 117. A. babatagi M. Pop. in Schedis ad Herb. Fl. As. Med. XVI(1928) 41; Fl. Tadzhik. V, 343.-Exs.: Herb. Fl. As. Med. No. 377. Perennial, closely allied to the preceding species; distinguished chiefly by the leaf-rachises becoming indurated in fruit; plants (10) 15-23 (30) cm tall, subacaulescent, rarely with stems up to 5-1lcm long; stipules trian- gular-lanceolate, acuminate, 5-7 mm long; bracts oblong-lanceolate, ca. 2mm long, about equaling the pedicels; calyx cylindric-campanulate, green- ish, 6-7 mm long, covered with scattered appressed to subascending hairs, the teeth linear -subulate from abroadbase; corolla yellow, commonly green- nerved; standard 9-11 (12) mm long, the limb obovate, angular -auri- culate at base, 2.5-3 times as long as the claw; pod 9-10mmlong, more or less flattened or carinate dorsally, subunilocular, the stipe very short, the beak 1-2mm long. Fl. May-June. Foothills and the low and intermediate mountain zones; zone of ephemer- al vegetation, more rarely in the open part of the woodland and scrub belt; clay and gravel slopes. - Soviet Centr. Asia: Pam.-Al. (from Kulyab to . Babatag). Endemic. Described from Babatag range. Type in Tashkent. 118. A. densus M. Pop. in Schedis ad Herb. Fl. As. Med. XV (1928) in adnot. ad No. 377 (nomen solum).-Myobroma densa (M. Pop.) Nevski in Acta Inst. Bot. Acad. Sc. URSS, ser. I, 4 (1937) 256. Perennial, acaulescent, 4-16 cm tall, ,compactly cespitose; stipules 132 lance-oblong to lanceolate, subacute or acuminate, beset with white hairs; leaves 4-15 cm long, the petiole very short, shaggy with white spreading hairs; leaflets oblong-oval, (3) 4-9 mm long, densely covered beneath with appressed hairs; peduncles 4-8 (10) mm long, villosulous; racemes many, very short, 3-5-flowered, fasciculate, axillary, forming at the leaf base a rather compact, moré or less spherical "inflorescence"; bracts linear - lanceolate, 3-5 mm long, hyaline, villous; pedicels 1.5-3mm long; calyx 98 campanulate, 9mm long, villous, the linear-subulate teeth about equaling the tube; standard 11-12 mm long; wing-petals 10-11mm long, the limb oblong, 1.5-2 times as long as the claw; keel 10-11 mm long, the limb ob- tusish, equaling the claw; ovary short-stipitate, hirsute; pod ovaloid, 7- 8mm long, compressed, carinate dorsally, white-villous, unilocular, 2-3- seeded, the stipe very short, the beak 1.5-2 mm long. Fl. June-July; fr. June-July. Zone of ephemeral vegetation in the lower part of the juniper formation. - Soviet Centr. Asia: Pam.-Al. (southwestern spurs — Kugitang range). En- demic. Described from the Kugitang range. Type in Tashkent. ~ 119. A. bischkendicus Gontsch. in Append. ad Fl. Tadshik. V (1937) 660. Perennial, subacaulescent, suffruticose, compactly cespitose, 16-21 cm tall; stipules lanceolate, subulate-pointed, 11-12mm long, villous; leaves (9) 11-20 cm long, the very short petiole and the rachis indurated, spread- ing-villous, persistent but not spiny; leaflets 18-27 pairs, oblong, (3) 5- 9mm long, round-tipped or subobtuse, densely covered beneath with ap- pressed hairs; peduncles very short, 2.5-5mm long, villous, 2-3-flowered; bracts narrowly linear-subulate, 4-6 mm long, villosulous; pedicels 2-3mm long; calyx cylindric-campanulate, 10-11mm long, the subulate teeth equal- ing the tube; standard 15-17mm long, slightly retuse, the claw 5-6 mm long; wing-petals 14-15mm long, the limb oblong, dilated at apex, slightly retuse, nearly twice the length of the claw; ovary stipitate, villous; pod ovaloid, 6-8mm long, carinate dorsally, more or less villous, unilocular, 2-seeded, the beak and the stipe ca. 1.5mm long. Fl. April-May; fr. May. Low mountains; stony slopes. - Soviet Centr. Asia: Pam.-Al. (interior part — southern part of the Babatag range, Khodzha-Kadyyan mountains near Kabadian). Endemic. Described from the Babatag range. Type in Lenin- grad. Series 12. MICRODONTI Gontsch.- Acaulescent or nearly so; stipules hyaline, united at base with petiole; leaflets covered above with appressed short hairs; racemes loosely few-flowered, short-peduncled; calyx cylin- 135 dric-campanulate, the tube 2-3 times the length of the short teeth; corolla greenish-yellow; pod ovaloid-oblong, carinate ventrally, grooved dorsally, subsessile, thinly coriaceous, partially bilocular. 120. A. karategini Gontsch. in Fl. Tadshik. V Appendix (1937) 660. Perennial, acaulescent or nearly so (rarely stems up to 4 cm long), glabrous, 6-10 cm tall, loosely cespitose, from a tufted, much-divided caudex, stems abbreviated, pubescent; stipules lance-subulate or oval, ca. 5mm long, ciliate; leaves (3) 4-8 (14) cm long; the petiole 1/4-1/2 the length of the rachis, slender, marcescent, puberulent; leaflets oblong- oblanceolate or suboblong, (7) 8-10 (15) pairs, (3) 4-6 (10) mm long, round- ed-obtuse, densely appressed-canescent beneath, more thinly covered with short hairs above; peduncles 2-10 (20)mm long, pubescent, 1-3 (5)-flow- ered; bracts lanceolate, 4-5mm long, hirsute, scarious; pedicels slender, 1-2mm long, hirsutulous; calyx 8mm long, densely appressed-hirsutulous, the teeth linear-subulate; standard 13-15 mm long, the limb rounded-oval to suborbicular, slightly retuse, often angular at base, the claw 5-6mm long; wing-petals 12-13mm long, the limb oblong, round-tipped or obtuse, equaling the claw; keel 10-11 mm long, the limb obtuse, slightly shorter 99 \Y =, Ve) GY ip. SEMBAWS a> SANNA NNN é | OOIP a ; | J G, N ; ™ Py i ZS = pa, \ . tsk a y, } i Faw ae > Wy \\) 23 oO et Kir. Kar. PLATE XII N. Basil.—2. A. lithophilus rmosus 100 LAY Lip LZ 1, Astragalus aco 837, 1117 136 than the claw; ovary short-stipitate, sericeous; pod 10-11 mm long, bilocular to 1/3, tomentose. Fl. July-August; fr. August. Mountainous and high-mountain zones; rosaria of the upper layer of the wood- land and scrub formation and in subalpine xerophilized meadows. - Soviet Centr. Asia: Pam.-Al. (Peter I range, Darvaz region). Endemic. Described from the Peter I range. Type in Leningrad. Section 28. Albertoregelia M. Pop. in Sched. ad Herb. Fl. As. Med. HaSeP OV NOs 0° (L928) bso. xcle Aly ts chim oanieus. MePop.)). — Large herbaceous perennials, vested with white and black hairs; stems well developed or more or less reduced; stipules united at base with petiole; leaves imparipinnate, the petiole marcescent; racemes loose, borne on long axillary peduncles; bracts lacking; corolla glabrous, yellow, early fading; calyx cylindric; pods long-stipitate, more or less pendulous, tri- angular-oblong, large, fleshy-coriaceous, imperfectly bilocular. A monotypic Soviet Central Asian section, distributed on gypseous mot- tled layers in southern Pamir-Alai pyatirech'e. 121. A. Albertoregelia C. Winkl. et B. Fedtsch. in Bull. Jard. Bot. Pétersb. V, fasc. 1 (1905) 42; Fl. Tadzhik. V (1937) 308-309. Perennial, (27) 30-50 cm tall; stems reduced or well developed, (2.5) 5-20 (25) cm long, firm, glabrous; stipules united at base with petiole, ovate, 8-12mm long, acute, glabrous or nearly so; leaves (11) 20-36 (40) cmlong, the petiole obsolete, the rachis finely sulcate, commonly hispid; leaflets (5) 8-10 (13) pairs, ovate or oblong-ovate, rarely suborbicular or oval, (10) 16-30 (37) mm long, round-tipped or scarcely retuse, succulent, ap- pressed-hairy on both sides or glabrous; pedicels hispid at length glabrate, the early 3-6 cm, the later 7-10 (12) cm long; racemes loose, the early 2-6-flowered, 4-8 cm long, the later many-flowered, 12-20 cm long; bracts lanceolate, rarely oblong-ovate, 7-10mm long, acute, villous outside or merely ciliate-margined; pedicels pubescent, 2mm long; calyx cylindric, 15-20mm long, densely covered with black or mixed white and black hairs, the teeth unequal, two triangular-lanceolate, 2.5-5mm, the other three lance-linear, 4-7mm long; standard 32-36mm long, the limb oblong, slight - ly retuse, slightly angular at base, the claw broad, ca. 10mm long; wing- petals 30-32 mm long, the limb lance-oblong, round-tipped, about equaling the claw; keel 28-30mm long, the limb slightly gibbous, straight above, acutish, slightly cleft, the claw ca. 17mm long; ovary long-stipitate, to- mentose; pod 21-32mm long, ca. 9mmbroad, nearly flat and straight, cari- nate and rather straight ventrally, very thinly puberulent, imperfectly biloc- ular, borne on a stipe 10-15mm long, abruptly contracted into a beak (2) 3-6 mm long; seeds oblong-ovaloid, irregularly emarginate, ca. 4mm long, smooth, pale brown. Fl. April-June; fr. May-July. (Plate XVI, fig. 2). Foothills and low mountains in the zone of ephemeral and xerophytic woody vegetation in the multicolored-layer region. In areas with sparse plant-cover on clayey (mostly red) and sandy (detritus of gray sandstone) slopes and river banks. - Soviet Centr. Asia: (S. Tadzhikistan: Kulyab, Bal'- dzhuan, Karshitau mountains). Endemic. Described from the vicinity of Bal'dzhuan. Type in Leningrad. Section 29. PENDULINA Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946). -Herbaceous perennials, acaulescent or short-stemmed, 101 Wait 138 vested with simple black or mixed black and white hairs, rarely glabrous; stipules cauline, united with petiole or not so, free, rarely connate at base; petiole and leaf rachis marcescent; leaves imparipinnate, racemes rather loose, borne on axillary peduncles, rarely sessile subradical; bracts per- sistent; bracteoles none, rarely present; calyx campanulate-cylindric, rarely cylindric, hirsute with black or mixed black and white hairs; corolla deciduous, glabrous; wings shorter than the standard, exceeding the keel; ovary stipitate; pod stipitate, cernuous, thinly coriaceous, elongated, sub- triquetrous, straight and broadly grooved dorsally, carinate and convex ventrally, partially or fully bilocular.- A Soviet Central Asian section dis- tributed in W. Tien Shan and in Pamir-Alai. Mesophytes and mesomorphous ephemeroids. ; 1. Plants glabrous, large; a aan racemes Short, few-flowered, borne on axillary peduncles 3-4 cm long... 122. A. tschimganicus M. Pop. +, Plants ‘More Or JESS NAGY cee. Meee aoe 5 Ne «ate elec a ue) Rete Meet ele eee Ze 2. Leaflets orbicular, 4-6 pairs; stems 8-15 cm long; pods sparsely cov- ered with long spreading white hairs ....123. A. nuratensis M. Pop. + Leaflets linear-lanceolate, elliptic or ovate, rarely suborbicular and then. stems ?not developed). i wtnnhs hoe ei Sh. (PAR te Feed eee ee ees 3. 3. Leaflets 2-3 (4) pairs, ovate or oblong-ovate, (3.5) 4-7 (8) cm long .. SoA SPE ey nek OMT Bae ies els bie eee Pe le 124. A. acormosus Basil. wolieatlets (5)"6220 pairs) & eh Syl GeRekT eh desi ee See sien ye ne ee 4, 4. Corolla 4-5 cm long, bright green; leaflets linear or linear-lanceolate, 14-20 pairs; pods (23) 25-35 mm long, borne on a stipe 15-18 mm long, minutely white-puberulent and white-hirsute with spreading hairs; Cay XMAS AO LCrAOO MeN Me NaN Meal) 2) WRT 126. A. viridiflorus Boriss. + Corolla whitish, greenish-white, violet-white or yellowish-lilac, not - more than 23 mm long; calyx 8-12 mm long; pods 10-22 (25) mm long, borne on.avstipe) Sisli2ormmvlon gee! «a elon ee 5. 5. Plants 4-8 (9) cm tall; leaves (3) 4-8 (10) cm long; peduncles none or not exceeding 3-5 cm; racemes 3-5-flowered; leaflets oval or rounded- eval! '4=8'(0)imimrlohe) Aiea te haa Le Bee? eh On! 2 ee eee eee 6. + Plants (6) 15-45 cm tall; leaves (12) 15-35 cm long; peduncles (5) 8- 15 cm long; racemes 8-20-flowered; leaflets 1-5 (6.5) mm long, dif- ferentlyishapedic. (“arise . nei aon wena! atoll ch?) «arn Saeeepen ieee he 6. Ovanyrandjpod)plabmousi leit im cere) aya). clemal: ps 128. A. pauper Bge. + Ovary and pod spreading-villous. . 129. A. pauperiformis B. Fedtsch. 7. Corolla greenish-white, rarely slightly suffused withviolet; pods minute - liv white =pube rulentey ay. ew ably) Give) smal bans 127. A. mirabilis Lipsky + Corolla lilac-yellowish; pods covered with short black hairs loosely interspersed with long spreading white hairs. .......+.2+.++.s.2-+4:-. a, Sa aks AP aee « AUR OEE se) «reget ho. Sh 125. A. vegetior Gontsch. Series 1. GLABERRIMI Gontsch. - Plants devoid of pubescence. Western Tien Shan. 122. A. tschimganicus M. Pop. in Sched. ad Herb. Fl. As. Med. fasc. SOV S28) soy Hse) eer -w A Wil IO. oO. Perennial, 40-76 cm tall, completely glabrous; stems stout, erect, ribbed-sulcate, 18-38 cm long; stipules 12-14 mm long, the lower oblong- ovate, the upper ovate, acute; leaves 20-28 cm long, the petiole 5-6 cm long, sulcate; leaflets 8-9 pairs, ovate, subobtuse or round-tipped, 20-35 102 39 (40) mm long, thin, herbaceous; peduncles 3-4 cm long, 4-6-flowered; bracts linear-lanceolate, 5-8 mm long, hyaline, whitish; pedicels ca. 4mm long; calyx ca. 19mm long, the teeth linear, subulate-tipped, equaling the tube; standard ca. 30mm long, the limb oblong-obovate, slightly retuse, scarcely auriculate at base, the claw broad, ca. 8mm long; wing-petals ca. 25mm long, the limb oblong, round-tipped, the claw ca. 11mm long; keel 20 mm long, the limb slightly convex above and beneath, round-tipped and shortly cleft, equaling the claw; ovary linear, glabrous, ona stipe 4mm long; pods 20-30mm long, straight or scarcely concave, convex [?], acute at both ends, the stipe 10-limm long. Fl. andfr. June-July. Stony clay slopes, among shrubs. - Soviet Centr. Asia: T.Sh. (W.). En- demic. Described from Mt. Malyi Chimgan. Type in Tashkent. Series 2. PILOSI Gontsch. - Plants more or less hairy; pods covered with spreading long hairs. Nuratau mountains. 123. A. nuratensis M. Pop. in Not. Syst. ex Herb. Inst. Bot. Ac: Sc. Wits Vil, f) 5. (1938) 111. Perennial; stems 8-15 cm long, thinly vested with spreading hairs; stip- ules membranous, triangular-lanceolate (the lower ovate or oblong), gla- brous and sparsely ciliate, 5-10mm long; petiole about half the length of the rachis; leaflets 4-6 pairs, orbicular, retuse, subobcordate, 1-2cm long, long-ciliate on the margin and beneath, at length glabrescent; pe- duncles glabrate, 4-7 cm long; racemes secund, 5-10-flowered; bracts lanceolate, 5-7 mm long, sparingly black-ciliate; gynophore ca. 3mmlong; calyx cylindric, 10mm long, thinly covered with spreading white hairs, the linear teeth ca. 7mm long; corolla pale yellow (?); standard ca. 30mm long, narrow retuse, the claw long; wing-limb short, about equaling the claw, 15mm long; keel 25mm long, the limb acute, the claw 15mm long; pods 10-20mm long, borne on stipe 10-13 mm long, oblong pendulous acute coriaceous, fully bilocular mucronulate, thinly covered with ascending long white hairs. Fr. June. Stony slopes. - Soviet Centr Asia: Pam.-Al. (Nuratau mountains). En- demic. Described from Yang-ogla landmark in the Nuratau mountains (Ak- tau). Type in Tashkent. Series 3. PUBESCENTES Gontsch. - Plants more or less hairy; pods minutely puberulent as well as often covered with scattered, long, spreading hairs. Western Pamir-Alai. 124. A. acormosus N. Basil. in Not. Syst. Herb. Hort. Petr. IV (1923) 42; Fl. Tadzhik. V, 315-316. Perennial, (17) 30-45 (47) cm tall; stems (2) 10-23 (26) cm long, firm, angled, simple; stipules cauline, free or connate at base, membranous, whitish, glabrous or, when young, black-ciliate at apex, 20-35mm long, the lower oval-oblong, acute, the upper lanceolate from a broad base, acuminate; leaves 16-25 cm long, the petiole (5) 7-10 cm long, beset with short spreading hairs; leaflets 2-3 (4) pairs, ovate to oblong-ovate, (3.5) 4-7 (8) cm long, obtuse or acute, densely covered on both sides (more sparsely above) with short soft appressed hairs, sericeous when young; ra- cemes (7) 8-15 cm long, rather compact, 10-flowered; the peduncles axil- lary, covered with short spreading hairs, (10) 17-21 cm long, the flowers 103 140 divergent, at length nodding; bracts linear-lanceolate, 5-12 mm long, beset with spreading black and fewer white short hairs; calyx campanulate - cylindric, 11-13mm long, hirsute with white and black hairs, the teeth linear -lanceolate, slightly or scarcely shorter than the tube; corolla yellow- ish, often reddish in upper part; standard 20-21 mm long, the limb oval- quadrangular, scarcely retuse, slightly angular at base; wing-petals 17- 18mm long, the limb oblong, slightly dilated upward, round-tipped, barely longer than the claw; keel 10-11 mm long, the limb strongly convex dorsal- ly, scarcely so ventrally, equaling the claw; ovary short-stipitate, hirsute; pods cernuous, asymmetrically and triquetrously lance-oblong, 22-28 mm long, imperfectly bilocular, shortly black-pubescent and white-villous (rarely destitute of black hairs), the stipe 8-9mm long, the beak 3-5mm long. Fl. May-July; fr. May-August. (Plate XII, fig. 1). Scrub and witch-grass steppes, rosaria and subalpine mixed grass as- sociations (in the upper zone of the woodland and scrub formation to the lower limit of the Subalpine zone). - Soviet Centr. Asia: Pam.-Al. (W. Zeravshan, central part of Hissar range). Endemic. Described from Zerav- shan basin. Type in Leningrad. 125. A. vegetior Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS, IX (1946) 114. Perennial, 20-25cm tall, from a short buried woody caudex; stems (2) 5-15 cm long, prominently angled, glabrous; stipules free, herbaceous, 8-13 mm long, the lower ovate or oblong-ovate-triangular, the upper lanceo- late, acuminate; leaves (12) 15-20cm long, the rachis 1.5-2 timesthe length of the petiole, both petiole and rachis sulcate, at first thinly covered with spreading or ascending short white hairs, at length glabrescent; leaflets (6) 7-8 pairs, lanceolate, acute to subobtuse, sometimes truncate, rarely obovate, (2) 3-5cm long, deeply and broadly emarginate, glabrous, bright green above, thinly strigulose with appressed or subappressed short hairs; peduncles slightly shorter than the leaves (by not more than 1/3), together with inflorescence or fruiting racemes equaling the leaves, sulcate, rough with spreading or ascending short hairs; racemes lax, oblong, 6-7cm long, many-flowered, secund; bracts lanceolate, 2-3 times the length of the pedicel, covered with white andblack hairs; calyx cylindric -campanulate, 11-12 mm long, hirsutulous with short black spreading hairs, the tube 2- 2.5 times the length of the linear (lance-subulate) teeth; corolla lurid in bud, approaching lilac in anthesis, the standard and the tip of wings and keel lilac; standard 18-21 mm long, the limb oval-quadrangular, 7-8 mm broad, slightly retuse, rounded-angular at base, 2.5-3 times the length of the claw; wing-petals 16-19mm long, the limb oblong, slightly dilated upward, entire, round-tipped, equaiing the claw; keel 13-15 mm long, the limb gibbous, subtriangular, broad, subacute, 2/3 the length of to equaling the claw; ovary on a stipe 3-4mm long, densely hirsute; pods on a slender stipe 7-10mm long, asymmetrically lance-oblong, 17-22mm long, 5-6mm broad, acute at both ends, sharply carinate ventrally, broadly grooved dorsally, thinly coriaceous, sparsely white -hirsute with spreading hairs and more densely covered with minute black spreading hairs, imperfectly bilocular. Fl. June-July (August); fr. July (August). On more or less gravelly mountain slopes, chiefly in ephemeroid tall- grass associations of Prangos pabularia inthe lower part of the sub- alpine vegetation zone and inthe upper part of the zone of mesophilic woody vege - tation at 2300-3000 m alt. - Soviet Centr. Asia: Pamir-Alai (southern slope of 104 141 Hissar range, in Varzob river basin; passing over the Anzob pass onto the northern slope which forms the Varzob river valley). Endemic. Described from the northern slope of Anzob pass. Type in Leningrad. 126. A. viridiflorus Boriss. in Acta Inst. Bot. Acad. sce. URSS, ser. Ty 3 (1936) 207; Bl. Tadzhik, V, 315. Perennial, stems 1-1.5 ecm long; stipules cauline, free, lanceolate, long-acuminate, 2.2-2.5cm long; leaves 25-30 (the lower 8-9) cm long the petiole 4-5cm long, petiole and rachis sparsely hairy; leaflets 14-20 pairs, linear to linear-lanceolate, acute, 3-4 (5) cm long, glabrous above, sparsely covered beneath with appressed hairs; peduncles 22-27 cm long (shorter than the leaves), sparsely hairy; racemes loosely 7-9-flowered, 10-13 mm long, membranous, acuminate, covered with white and black hairs; calyx cylindric, 2-2.5 cm long, hirsute with white and predominantly black hairs, the tube 2-3 times the length of the lanceolate teeth; corolla bright green, large; standard 4-5 cm long, the limb oblong, narrow, grad- ually attenuate into a short claw; wing-petals 3.5-4.5 cm long, obtuse, the limb half the length of the claw; ovary oblong-lanceolate, ca. 1 cm long, white-hirsute, on a stipe ca. 7mm long; style glabrous; pods on a long stipe 15-18 mm long and equaling the calyx-tube, lance-oblong, acute, (23) 25-35 mm long, inflated, slightly carinate ventrally, broadly grooved dor- Sally, terminating in a gently reflexed beak 1.5-3mm long, coriaceous, minutely puberulent with fine white down interspersed with spreading stiff white hairs. imperfectly bilocular. Fl. May-June; fr. June. Loess foothills in the ephemeral vegetation zone. - Soviet Centr. Asia: Pam.-Al. (southern slopes of Hissar range, central and western parts). Endemic. Described from the vicinity of Stalinabad. Type in Leningrad. 127. A. mirabilis Lipsky in A,H.P. XXVI (1907) 155: Fl. Tadzhik. Venloe Perennial, (6) 15-30 (40) cm tall, acaulescent or with short stems 1- 4cm long; stipules cauline, free, oblong or linear-oblong, finely pointed, glabrous, herbaceous, 10-18mm long, sparsely ciliate on the margin and at apex; leaves (6) 16-30 (35) cm long; leaflets (3) 6-15 (20) pairs, varying from filiform to lanceolate, elliptic or ovate, acute or obtuse, rarely re- tuse, 10-30 (65) mm long, covered with scattered appressed hairs, some- times glabrous above; peduncles (5) 8-10 cm long, sparsely covered with appressed hairs or glabrate; racemes 8-20-flowered, 5-8 cm long, together with peduncle shorter than the leaves; bracts membranous, narrowly linear, 5-9mm long, beset with white and black hairs; flowers divergent, at length nodding; calyx cylindric, 8-12 mm long, hirsute with appressed black and fewer white hairs, the linear-subulate teeth about half the length of the tube; corolla whitish, the standard suffused with greenish-azure; standard 18- 23mm long, the limb broadly oblong, mostly slightly retuse, abruptly at- tenuate into and 3 times the length of the cuneate claw; wing-petals 16- 19mm long, the limb oblong or spatulate -oblong, round-tipped, somewhat longer than the claw; keel 13-15mm long, the limb strongly convex be- neath, slightly concave above, obtusish, about equaling or slightly shorter than the claw; ovary long-stipitate, minutely puberulent; pods cernuous, triquetrously ellipsoid-lanceolate, up to 2-2.5cm long, 10-15 mm thick, minutely white-puberulent, subbilocular, the stipe 5-12mm long exserted from the calyx, the beak 1-2 (3)mm long; seeds reniform, pitted, 4-5mm long, reddish-brown. Fl. May-June (July); fr. June-July (August). 105 142 Widespread in the low and intermediate mountain regions, at 900-2700 m alt., inthe woodland and scrub formation and in the upper part of the ephem- eral vegetation zone, chiefly in scrub and woodland and scrub associations. - Soviet Centr. Asia: Pam.-Al. (interior part, to the south of the Hissar range). Endemic. Described from S. Tadzhikistan (Bal'dzhuan, Kulyab, Darvaz). Type in Leningrad. 128. A. pauper Bge. Astrag. turk. (1880) 224; Fl. Tadzhik. V, 312.- A. kohistanus Franchet in Ann. Sc. Nat. sér. VI, XV (1883) 254. Perennial, acaulescent, 4-8 (9) cm tall; stipules free, cauline, united at base with petiole, oblong, 6-10mm long, membranous, ciliate, the lower obtuse, the upper acute; leaves (3) 4-8 (10) cm long, the petiole muchshorter than to nearly equaling the rachis, sparsely covered with spreading hairs or glabrous; leaflets (5) 10-15 pairs, rounded-oval or oval, (4) 6-8 (10)mm,. long, obtuse or slightly retuse, covered on both sides with scattered ap- pressed hairs or glabrous; peduncles scarcely developed or up to 3-5cm long, 3-5-flowered; bracts linear, more or less membranous, 4-8 mm long, sparsely black-ciliate or glabrate; calyx campanulate-cylindric, 8-10mm long, black-hirsute, the lance-linear teeth half the length of the tube; corol- la white or whitish, sometimes fading reddish; standard 20-24mm long, the limb obovate-oblong, slightly constricted at the middle, 4-5times as longas the claw; wing-petals 17-20mm long, the limb oblong, slightly dilated up- ward, slightly retuse, somewhat longer than the claw; keel 13-15 mm long, the limb subrectangularly convex beneath, 2/3 the length of the claw; ovary glabrous or sparsely hirsutulous; pods subcernuous, asymmetrically ellip- soid or lance-oblong, 10-15 (18) mm long, partially bilocular (septum pro- jecting from dorsal suture), the stipe 3-5mm long, the very short beak 1- 2mm long; seeds ovaloid, emarginate, smooth, 2-2.5 mm long, brown, Fl. June-July (August); fr. July-August. High-mountain zone, 2800-3500 m, subalpine meadows and steppes, rarely in the Alpine zone (up to 4200 m).- Soviet Centr. Asia: Pam.-Al. (Hissar, Alai, Peter I, and Darvaz ranges). Endemic. Described from Saratag river valley in the Zeravshan basin. Type in Leningrad. 129. A. pauperiformis B. Fedtsch. in A.H.P. XXIV (1905) 207. Perennial, very closely related to the preceding species; differing in the shaggy ovary and pod; corolla yellow. In subalpine coarse-grass association (Cousinieta) and in xerophyll sub- alpine meadows. - Soviet Centr. Asia: Pam.-Al. (Darvaz region). Endemic. Described from Darvaz. Type in Leningrad. Note: All available material concerning this species relates to the Darvaz region (Khazret-sultan, Kugifrush, and some other mountains), except for the one specimen from which A. pauperiformis was de- scribed. The label of this specimen does not record any of these locations, but indicates the mountains of Chimgan and W. Tien Shan. A careful exam- ination of all the material showed a perfect identity between the type and the specimens collected by Regel in the Khazret-sultan mountains which should indeed be considered the ''locus classicus"' of the species, since the description from W. Tien Shan was due to the mixing up of labels while arranging the herbarium. Thus A. pauperiformis is a minor geo- graphical race, replacing A. pauper Bge. inthe Darvaz region, and both species belong to Pamir-Alai. 106 143 144 Section 30. MYOBROMA (Stev) Bge. Astrag. geront. I (1868) 33; genus Myobroma Stev. in Bull. Soc. Nat. Mosc. IV (1832) 268. - Acaulescent or subacaulescent perennials, very rarely with stems up to 20cm long, from a short more or less branched buried caudex; stems of the year greatly re- duced and undeveloped, concealed under a thick coat of approximate petioles and stipules, vested with simple basifix white hairs; stipules united with petiole, not connate; leaves imparipinnate or verticillately pinnate, the petiole and rachis marcescent, rarely indurated and persistent but not spi- nescent; racemes loosely few-flowered, commonly subradical; bracteoles wanting (very rarely present); calyx cylindric, rarely campanulate-cylin- dric; pods sessile or stipitate, unilocular, rarely fully bilocular. Ephemeroids, with a growth cycle adapted to a brief wet and warm peri- od. Greatest differentiation centered on areas with a pronounced drought period which brings about a rest period. The plants display very marked features of adaptation of the mesophytic constitution to a specific climatic regime: secondary acaulescence, a shortened growth period, etc. The distribution area of the section embraces the Mediterranean region, Near Asia (Asia Minor, Armenia, Kurdistan, Iran, Afghanistan, Arabia, N.W. India) and Central Asia, reaching in the east as far as Altai, in the west to southern Europe, and in the north to central Europe and the steppe and forest-steppe regions of the U.S.S.R. Species of Near and Soviet Central Asia are most numerous. Key to Species in Flowering Condition PICA VE SMM OAT PIMMALE Tce ere ceil e Lesa atie ce ue ete euNIes sovelie erate anasmtag an mi) Phe CVC Sve LUlCLLlALe LY OUMMALE hs Roti hts Moe eco MG kM UNecr lhe sta eM aMe 5s 82. 2. Upper face of leaflets hairy (at least at the edge of the lamina).... 3. wpe race, Of learlets plabrouists'.'. 24s he he Mile) s a etee te anit he, mer sh ys ce 20. 3. Upper face of leaflets white-floccose (the vesture consisting of slender, Solty scuLteduhalis) Or aArACHNOldw rs 4 cms Wak eae: She Rn, SCREEN Gu, 4, + Upper face of leaflets covered with appressed or appressed-curly hairs SAD, STA Re dere itt ae RE Re aE Se sthe Mach Gl duc i en i bea Re RAs URS as 6. 4. Corolla red; standard 26-34mm long; leaflets white-floccose above, oval or rounded-oval, (2) 3-5 (5.5) mm long. (Kopet Dagh, chiefly OTMEE eI) erin oe ar ty ete et ee Mane nce a inte Mat, es 200. A. curvipes Trautv. + Corolla yellow or whitish; standard 16-22mm long ........... 5. 5. Leaflets 16-23 (25) pairs, rounded-elliptic, rounded-oval or oblong- ovate, rarely broadly oblong, arachnoid-pubescent above; corolla yel- low (Kopet Dagh, chiefly western)...... 198. A. angustidens Freyn. + Leaflets (22) 26-36 (40) pairs, oblong or oblong-oval, white -floccose; corolla whitish. (Badkhyz, S.W. Pam.-Al., Kopet Dagh)......... 21 Sou aie Sb. SNE HONE EN Bolte ae Si onoie 199. A. Barrowianus Aitch. et Hemsl. 6. Calyx-tube equaling the teeth or nearly so; standard-limb oblong-oval Ormoblong (sometimes nearly quadranpular). . 3.2). hes ca tiene eve) aus Ue + Calyx-tube 2-4 times as long as the teeth, rarely 1.5 times as long and LHe testandar ad —Limily \OWOVACCI A \uetalis cl sift 6) (a0 > iRewtoll «ve Mlle aetna emDey oll lat elie Zi ++ Calyx-tube 1.5-2 times as long as the teeth; standard oblong....... 5 TTS Mae acetal tandem Mabel mbes 172. A. Henningii (Stev.) Boriss. 7. Wing-petals round-tipped; leaflets sericeous above (Bessarabia, Bl., central and rarely western and southern Europe). 187. A. excapus L. 107 145 Il. 16. Wife Wing -petals retuse, rarely obtuse and entire and then leaflets covered above with scattered, long, ascending hairs .............6... 8. Leaflets acute, terminating in a short whitish cartilaginous point; ealys 12-13 mim long s(Panin= Al > «Daiey az CCID) non nica: amen ee eB, ke dee Here repicd cn aphasia syeehth cn tere aeaeniaend dows 165. A. apiculatus Gontsch. Leaflets round-tipped, rarely retuse or subacute, without a cartilagi- nNOwWS} point; eadyse te on the margin; calyx glabrous; standard obleng-obovate, gently angular (S.W. Altai, Irtysh river basin from Zaisan kettle hole) ...... 144. A. altaicus Bge. Leaflets 20-22 pairs, covered beneath with scattered appressed hairs; calyx hirsute on the teeth; standard oval or obovate (Dzungarian Ala Raum Ng Bienishan) ne. ae wih es wee 173. A. chlorodontus Bge. Standard 30-35 mm long; calyx-tube 3.5-5 times the lengthof the teeth. 31. 109 ol. 32. 147 39. 40, 148 41. Standard 17-28 (30) mm long, rarely reaching 33mm and then calyx- tube_2.5 timesithew@eneth ofjthe teeth = mmacrials tema aie ils to kalba eee a2 Leaflets 25-33 pairs, covered beneath with scattered appressed hairs (foothills and low mountains of Tien Shan and Pamir-Alai)........ wo abou Pts, WU As SUG SRLS TAD ten. cet ae 184. A. macronyx Bge. Leaflets 8-16 pairs, villosulous beneath (sands of Lower Volga region and Kazakhstan, eastward to the Zaisan depression)............ oo Seer Aah CN Ole abe ts cow ck A Rete Ream ct 8 145. A. longiflorus Pall. Leaflets (1) 1.5-3 (4) mm long; a low compactly cespitose plant, 8- 11cm tall; leaf petioles and rachises becoming indurated and trans- formed into spines (W. Pamir-Alai: basins of Kashka Darya, San- gardak and Tupalang rivers),,....... 161. A. indurescens Gontsch. Leaflets 5 and more mm long (rarely 4mm long); leaf petiole and Hachis NOUANAUGES SCENES fe eyiowiardd Veit ys 4 Postale 0 He ba tor rhea Maphauple Me mammonlameemee 33. Leaflets,and calyx/completely glabrows|(:-4)% 292’ ..0-niencee mere etree 34, Leaflets and calyx, or at least the leaflets, more or less hairy (some - timesisparsélys0).4 (On08 UO). She SOUR, OB Oe uke 35. . Leaflets 4-6 pairs; standard broadly ovate; wings linear-oblong (E. TYHARSCD SRA Alaiye: alee antowehie hatuethanente 134. A. schemachensis Karjagin Leaflets (12) 18-26 (28) pairs; standard oval; wings rounded, spatu- lately dilated upward (Badkhyz)...... 196. A. kuschkensis Boriss. Leaflets villous beneath with rather long hairs (E. and S. Transc., INFLAM) NA. MPI, ITED RG RETR Os 133. A. fabaceus M.B. Leaflets covered beneath with short appressed hairs, sometimes hispid on! thes midvein, + .§ .: frien ... Vee. a os eeeeeeee 88. + “otipules merely ciliate=margined 27.22") Sl VOey 2S ne ee 89. 88. Leaflets in whorls of 6-8 (10), lance-oblong; calyx 12-14 mm long, the tube 2-3 (4) times the length of the teeth; pods lance-oblong, 13- 17 mm long, carinate ventrally, finely and broadly grooved, coria-. ceous, densely covered with short ascending hairs, bilocular to 2/3- (Upper Pyandzhybasiny ‘earmiii)) ene 207. A. dignus Boriss. + Leaflets in whorls of 3-4, lanceolate to lance-obiong, shaggy with soft and rather long hairs; calyx 14-15 mm long, the tube equaling to slightly longer than the tube; pods ovaloid, 9-12 mm long, inflated, coarsely carinate ventrally, flattened or broadly sulcate dorsally (with a raised dorsal suture), rigidly coriaceous, densely shaggy with long soft spreading hairs, bilocular to 1/2-2/3 (northern Pamir-Alai: mountainous Zeravshan, northern slopes of Turkestan range) he BREN BOM Ra RL OBE ap ae tpi ett i te iL ata 208. A. patentivillosus Gontsch. 89. Leaflets linear to linear-oblong, (3) 7-11 mm long, 1.5-2.5 (3) mm broad; bracts 3-4 mm long; calyx shaggy with divergent hairs, the tube (2) 3 times the length of the teeth (W. Tien Shan, N. and N.E. 124 Pamir-Alai: northern slopes of Alai and Turkestan ranges, mountain- ous Zeravshan, Karategin, Vakhsh)....... 205. A. alaicus Freyn. 166 + Leaflets lanceolate to lance-oblong, 4-8 mm long, 2-3.5 mm broad; bracts 8-9 mm long; calyx covered with soft spreading long hairs, the tube 1.5-2 times the length of theteeth (S. Pamir-Alai: Kugitang range). eas Re terme Tbe ce Bi re bay bees qaeke * ceyerpeg th. dh pmarl Soe 206. A. Nevskii Gontsch. Series 1. RIGIDI Gontsch. - Acaulescent or subacaulescent plants; ra- cemes few-flowered, short-peduncled; calyx cylindric, corolla yellow; wing -petals oblong or linear-oblong, entire; pods sessile or subsessile (stipe not exceeding 1 mm), large, rigidly cartilaginous-cocriaceous, cari- nate ventrally with a thickened suture, rather broadly sulcate dorsally, in- completely bilocular, glabrous or sparsely covered with spreading, rather long hairs. S. Altai, Tien Shan, Pamir-Alai, Kopet Dagh, Caucasus, N. Iran, Ar- menia, Asia Minor. This series includes the species: A. anatolicus Boiss. (Asia Minor), A. Fraxinella Bge. (Armenia, N. Iran), A. remotijugus Boiss. et Hoh. (N. Iran), A. lobophorus Boiss. (N.W. Iran), A. samamensis Boiss. et Buhse, etc. 130. A. angustiflorus C. Koch in Linnaea XV (1841) 72; Bge. Astrag. geront. II, 39; Boiss. Fl. or. II, 289; Grossh. Fl. Kavk. II, 317. Perennial, acaulescent plants (5) 12-20 cm tall; stipules 10-14 mm long, the lower ovate, the upper lanceolate, ciliate; leaves (5) 10-20 cm long, the petiole (1) 2-5 (6) cm long, covered with spreading hairs, rarely gla- brate; leaflets 8-18 pairs, ovate-oblong, (4) 10-25 mm long, round-tipped, rarely subacute, ciliate on the margin and on the midvein beneath, rarely glabrous; peduncles 0.5-1.5 cm long, glabrous, 3-5-flowered; bracts line- ar-lanceolate, 5-7 mm long, ciliate; pedicels glabrous, 3-4 (6) mm long; calyx cylindric-campanulate, 9-11 (13) mm long, the tube glabrous, the teeth barbate 3-4 mm long; corolla yellow; standard 21-33 mm long, op- long, retuse, the claw 4-7 mm long; wing-petals 18-24 mm long, the limb linear-oblong obtuse, the claw 7-10 mm long; keel 14-19 mm long; ovary subsessile, shaggy; pods sessile, oblong, acute, 15-23 mm long, inflated dorsally, more or less shaggy, bilocular to 1/2-2/3, the beak 3-6 mm long. Fl. May-June; fr. June-July. Stony mountain slopes.- Caucasus: E. and S. Transc., Tal. Gen. distr. : Arm.-Kurd. Described from Kars and Shuragel. Type in Berlin. 131. A. aegobromus Boiss. et Hohen. Diagn. Ser. I, IX (1849) 74; Bge. Astrag. geront. II, 39; Boiss. Fl. or. II, 295; Grossh. Fl. Kavk. II, 318. Perennial, glabrous, acaulescent plants, 20-40 (50) cmtall; stipules di- morphic, the lower ovate, the upper lance -oblong acuminate; leaves 15-28 (38) cm long; the petiole 6-7 cm long; leaflets (7) 11-12 pairs, ovate, rarely lanceolate, obtusish or round-tipped, mucronulate, 13-22 (30)mm long; pe- duncles 1-2 cmlong; racemes 3-7-flowered, ca. 4cm long; bracts lanceolate to ovate-lanceolate, 9-10(14)mm long; pedicels 4-5 mm long; calyx cylindric - campanulate, 11-12 mm long, glabrous; the subulate -lanceolate teeth unequal, the longer ones slightly exceeded by the tube; corolla yellow; standard 20- 21mm long, the limb oval to oblong oval, round-tipped or slightly retuse, some- times scarcely angular at base, twice the lengthof the claw; wing-petals 17- 19 mm long, the limb linear -oblong, equaling the claw; keel 14-16 mm long; 125 ovary sessile or nearly so; pods oblong, subquadrangular -ovaloid, 20mm long, inflated dorsally, glabrous, rigidly cartilaginous -coriaceous, slightly rugose, almost fully bilocular, thebeak4-5mm long. Fl. June-July; fr. July-August. Lake shores and river banks in the forest zone; riverside pebbles. - Caucasus: S. Transc. (Nakhichevan A.S.S.R., Yerevan, Nor-Bayazet). Gen. distr.: N. Iran. Described from the vicinity of Derbent and El'borus Type in Geneva. 132. A. torrentum Bge. Astrag. gerontII (1869) 55; Boiss. Fl. or II, 29S eGrossh. while weak ll ole) lo SOF Perennial, related to the preceding, differing chiefly in the shaggy ovary and the pod hispid with spreading hairs; standard-limb subquadrangular - oval, angular at base. Fl. June-July; fr. July-August. Pebbles. -Caucasus: S. Transc., Tal. (according to Grossheim's indi- cation in the Flora of the Caucasus). Gen. distr.: N. Iran. Described from the surroundings of Nakhichevan-chai. Type in Leningrad. 133. A. fabaceus M.B.Fl. Taur.-Cauc. III (1819) 496; Ldb. Fl. Ross.I, 653, excl. syn. Gand.; Bge. Astrag. geront. II, 53; Boiss. Fl. or. II, 289; Grossh. Fl. Kavk. II, 317.-A. tumidus M.B.Fl. Taur.-Cauc. II (1808) 198, non Willd.-Exs.: Herb. Fl. Cauc. No. 373. Perennial, acaulescent plants 8-20 (25) cm tall; stipules white -hyaline, long-ciliate on the margin, acuminate, united at base with petiole, 10-15mm long, the lowest ovate, the upper lance-oblong; leaves (4) 12-23 (28) cm long, the petiole firm, (2) 3-7 (9) cm long, both petiole and rachis sulcate, beset with spreading reddish long hairs; leaflets 7-9 (11) pairs, oval to rounded-oval, rarely elliptic, round-tipped, rarely slightly retuse, rounded at base, (6) 12-30 (37) mm long, glabrous above, shaggy beneath (especial- ly on the midvein and sometimes almost solely on it) with long mostly red- dish hairs; peduncles firm, sulcate, covered with divergent reddish long stiff hairs, 2-8 cm and with infl. 6-16 cm long; racemes loosely branched, oblong-oval in outline, 7-12-flowered, the flowers spreading to somewhat 168 drooping; bracts narrowly lanceolate, long-acuminate, thinly membranous, whitish, (6) 7-10 mm long, shaggy outside with long hairs; pedicels 1-3 (4) mm long, covered with rufous long spreading hairs; calyx cylindric, (12) 15-18 mm long, shaggy with long spreading hairs, the teeth lanceolate or linear-subulate (3) 4-5 mm long; corolla pale greenish-yellow; standard 25-28 (33) mm long, the limb oblong-ovate, rarely oblong-oval, scarcely retuse, scarcely angular at base, the broad claw 7-10 mm long; wing-petals 22-25 (31) mm long, the limb oblong, obtuse or subobtuse, slightly incurved, more or less gibbous above the auricle, the basal auricle 2.5-3 mm long, the claw 10-14 (17) mm long; keel 20-24 (28) mm long, the limb acutish, triangular -obovate, the lower margin rounded-convex at a right or an obtuse angle, the upper margin slightly concave in its middle part, the basal au- ricle short semicircular, the claw 10-14 (16) mm long; ovary subsessile, slenderly fusiform, glabrous; style glabrous; pods subsessile, oblong, triquetrous, 25-40 (50) mm long, acute at both ends, rigidly coriaceous, glabrous, bilocular, many-seeded, the beak 2-3 mm long; seeds ovaloid, asymmetrically reniform, 5-6 mm long. Fl. April-May (June); (May) June. Dry and stony slopes and precipitous clay mountainsides, more rarely | on sands.- Caucasus: E. and S. Transc. Gen. distr.: N. Iran. Described from the Caucasus. Type in Leningrad. 134. A. schemachensis Karjagin in Not. Syst. ac Geoga. Inst. Bot. | Tphilis. f. 13 (1946). | 126 Perennial, subacaulescent, glabrous plants, 15-23 cm tall; stems 2- 3 cm long; stipules united at base with petiole, thinly membranous, tri- angular-ovate, short-acuminate, 12-13 mm long; leaves 13-22 cm long, the rachis 1.5-2 times the length of the sulcate petiole; leaflets 4-6 pairs, ovate, 20-30 mm long, 11-23 mm broad, subobtuse; racemes subsessile, Subradical, strongly abbreviated, 2-3-flowered; bracts lanceolate, ca. 9mm long, thinly white-membranous; pedicels 5-6 cm long; calyx13-14mm long, the linear-subulate teeth ca. 4 mm long; corolla pale yellow; stand- ard ca. 24 mm long, the limb broadly ovate, scarcely retuse, twice the length of the claw; wing-petals 20 mm long, the limb linear-oblong, round- tipped, as long as the claw; keel ca. 18 mm long, the limb acutish, slightly shorter than the claw; style-glabrous; pods sessile, asymmetrically oblong, 3-3.5 cm long, 1.1-1.2 cm broad, carinate ventrally, broadly sulcate dor- sally, subtriquetrous, short-acuminate to a subulate beak 3-4 mm long, rigidly cartilaginous-coriaceous, almost fully bilocular. Fr. May. Clay and stony mountain slopes. - Caucasus: E. Transc. Endemic. De- scribed from the Gil'gin-chai river valley in the Divichi region of Azerbai - jan, the only location from which it is known. Type in Leningrad. 135. A. Trautvetteri Bge. Astrag. geront. II (1869) 54; Astrag. turk. 238. Perennial, acaulescent plants, (10) 17-35 cm tall; stipules 12-20 mm long, the lower triangular-ovate, the upper oblong-triangular, all acumi- nate; leaves (8) 13-35 cm long, the petiole covered with spreading hairs, (3) 5-12 cmlong; leaflets (10) 12-20 pairs, oblong-ovate, obtuse to acute, (6) 12-28 mm long, sparsely ciliate beneath on the midvein and on the mar- gin; peduncles (3) 4-8 cm long, covered with spreading hairs; peduncles 5-7-flowered; bracts linear or lance-subulate, 8-12 mm long, covered with spreading hairs; pedicels 6-7 mm long, covered with spreading hairs; calyx cylindric, 11-13 mm long, sparsely covered with spreading hairs, rarely glabrous, the teeth subequal, more or less subulate, 2.5-4 mm long; corolla yellow; standard 24-26 mm long, the limb broadly oblong, angular at base, the claw 6 mm long; wing-petals 22-24 mm long, the limb oblong, somewhat longer than the claw; keel 17-18 mm long; ovary glabrous, ses- sile or raised on a peduncle ca.1mm long; style glabrous; pods triquetrous- oblong, ca. 20 mm long, glabrous, bilocular, the beak 4-6 mm long, FI. June-July; fr. July. Mountain steppes and mixed-grass meadows.-W. Siberia: Irt. (extreme south); Soviet Centr. Asia: Dzu.-Tarb. (Tarbagatai). Endemic. Described from Tarbagatai. Type in Leningrad. 136. A. Schanginianus Pall. Astrag. (1800)77; DC. Prodr. II, 303; Ldb. Fl. Alt. III, 332; Ej. Fl. Ross. 1, 652; Bge. Astrag. geront. II, 54; Bge. Astrag. turk. 237; Krylov, Fl. Zap. Sib. VII, 1664.-Ic.: Pallas, As- trag.)t) 63) Perennial, acaulescent or short-stemmed (stems 1-5 cm long) plants (10) 20-43 cm tall; stipules scarious, ciliate, ovate-lanceolate; leaves (10) 20- 40 cm long, the petiole much shorter than the rachis, sparsely covered with Spreading hairs; leaflets 12-17 pairs, ovate, round-tipped, (10) 15-30 mm long, covered beneath and on the margin with scattered long soft hairs; pe- duncles 1.5-6 cm long; racemes loosely (5) 8-12-flowered; bracts linear, scarious , long-ciliate, 2-3 times the length of the pedicel; pedicels 2-4 mm long; calyx cylindric, 13-19mm long, rather sparsely covered with soft 127 hairs, the teeth lance-linear, acute, 2/3 as long to as long as the tube; corolla yellow; standard 20-26 mm long, the limb obovate, round-tipped, gradually attenuate into and 2-3 times as long as the claw; wing-petals 18-22 mm long, the limb oblong, round-tipped, equaling the claw; keel 16-18 mm long, the limb convex beneath, nearly straight above, ob- tuse, ca. 2/3 the length of the claw; ovary hairy, raised on a glabrous stipe up to 2mm long; pods borne on a very short stipe (ca. 1 mm long), oblong to oblong-ovate, 15-22 mm long and ca. 7 mm broad, carinate ventrally, sulcate dorsally, somewhat compressed laterally, coriaceous, rather sparsely villous with soft white hairs, triangular in cross-section, bilocu- lar. Fl. June-August; fr. July-August. TO) Foothills and chiefly the mountain zone. Scrub and meadows, screes, and stony slopes. In Altai — on exposed stony and gravelly slopes of steppe hills and mountains.- W. Siberia: Alt. (S.E. and S.); Soviet Centr. Asia- Dzu.-Tarb., Tien Shan (Centr., N. and more sparsely W.). Gen. distr. : Kulja. Described from Altai (Katun' river basin). Type in Leningrad. 137. A. Severzovii Bge. Astrag. geront. II (1869) 54; Ej. Astrag. turk. 238. Perennial, acaulescent or rarely subacaulescent, (7) 10-25 (40) cm tall; stipules dimorphic, the lower ovate, the upper lance-oblong, 10-15 (20) mm long, glabrous; leaves (7) 11-30 (40) cm long, the mostly glabrous petiole much shorter than the rachis; leaflets oval or ovate, rarely suborbi- cular, (7) 10-15 (45) mm long, round-tipped; racemes sessile or nearly so, 4-8 (16)-flowered; bracts lanceolate, (7) 13-20 mm long, acuminate or acute, glabrous; pedicels (3) 4-6 mm long; calyx short-cylindric, 15-19 mm long, the linear teeth as long as or slightly shorter than the tube; corolla yellow; standard 20-27 (32) mm long, the limb round-tipped or scarcely retuse, the claw 5-8 mm long; wing-petals 18-25 (30) mm long, the limb about as long as to slightly longer than the claw; keel 15-20 (23) mm long; ovary on a stipe 1-1.5 mm long, hairy, rarely glabrous; pods subsessile, ovaloid-oblong to ovaloid, 10-17 mm long, inflated (dorsally), incompletely bilocular, covered with spreading hairs, rarely glabrous, the beak 2-4 mm long. Fl. June-July; fr. July-August. From foothills to the Subalpine zone.-Soviet Centr. Asia: Tien Shan (W.), Pam.-Al. (N.). Endemic. Described from the Karatau range. Type in Leningrad. Note. Inthe area of contact with A. Schanginianus Pall. forms of transition towards this species occur. These forms have hairy calyx teeth and more or less hairy or ciliate leaf margins. In W. Tien Shan (in the Tashkent area) hybrids occur between this spe- cies and A. lasiopetalus Bge. These are characterized bythe standard - limb which is narrowly oblong, sometimes slightly dilated below and mostly hairy, as well as the pods which are small, ovaloid and more compressed. These forms of hybrid origin were probably described by Bunge as A. sub - barbellatus Bge. (Astrag. turk. p. 227, 1880). 138. A. tupalangi Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 118. Perennial, acaulescent, ca. 35 cmtall, from an abbreviated, stout, short-branched, buried caudex; stipules united at base with petiole, large, ovate, often acuminate, rounded-auriculate at base on one side, thinly membranous, strigose on the margin; leaves (15) 20-30 cm long, the petiole 128 and the much longer rachis rigid, sulcate, glabrous; leaflets 12-13 pairs, ovate or subovate, obtusish, (15) 20-27 mm long, rather thick, glabrous 171 above, thinly covered with stiff hairs on the midvein beneath and sparsely 172 strigose on the margin; racemes subradical, loose, 6-7 cm long, rather many-flowered; bracts oblong-ovate to ovate-oblong (the upper ones broad- ly lanceolate), (9) 10-14 mm long, slenderly long-acuminate, whitish hya- line, strigose-margined; pedicels 2-3 mm long; bracteoles small, lanceo- late, ca. 3mm long, acuminate, whitish, strigose, inserted at the middle of the pedicel; calyx cylindric, ca. 20 mm long, glabrous, merely the nar- rowly linear acuminate teeth strigose; corolla bright yellow; standard 30- 33 mm long, the limb obovate-oblong, retuse, slightly angular at base, 10- 11 mm broad at apex; wing-petals 24-31 mm long, the limb oblong, round- tipped, 2/3 the length of the claw; keel 25-26 mm long, the limb gibbous, curved into a short broad obtuse rostrate apex, half as long as the claw; ovary on a stipe ca. 1 mm long, strigose; pods subsessile (the stipe ca. 0.5 mm long), oblong-ovaloid, 11-12 mm long, ca. 7 mm broad, slightly compressed laterally, obtusish, carinate, rounded or slightly flattened dorsally with a convex dorsal suture, coriaceous, hispid with spreading hairs, bilocular, the straight rigid beak ca. 3mm long. Fl. May; fr. (May) June. On granite outcrops. - Soviet Centr. Asia: Pam.-Al. (Tupalang river basin). Endemic. Described from Tupalang river valley, at the mouthof Kshtut River. Type in Leningrad. 139. A. cartilagineus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX, 1 (1941) 99. Perennial, acaulescent, 10-17 cm tall; stipules triangular-ovate, acute, 8-10 mm long, strigose; leaves 8-17 cm long, the petiole 1.5-3 cm long, more or less covered with spreading bristles, persistent; leaflets obovate, (6) 20-38 mm long, broadly cuneate at base, sparsely hispid beneath (some - times merely on the prominent veins), rather sparsely bristly -ciliate on the margin; peduncles 0.6-1 cm long, 2-3-flowered; bracts lanceolate, acute, ca. 7mm long, stiffly ciliate on the margin; pedicels 3-5 mm long; calyx cylindric, ca. 13 mm long, hispid, the teeth subulate-lanceolate, ca. 3 mm long; corolla yellow; standard ca. 22 mm long, the limb obovate, round-tipped, the claw 8 mm long; wing-petals 21 mm long, the limb oblong, 7mm long; keel ca. 19 mm long; ovary on a stipe ca. 2 mm long, woolly with white-hairs, the style glabrous; pods sessile, quadrangularly oblong- ovaloid, (12) 15-23 mm long, hispid with scattered spreading hairs, in- completely bilocular, the beak 3-4 mm long. Fr. June. Gravelly mountain slopes. - Soviet Centr. Asia: Mtn. Turkm. (Kopet Dagh). Endemic. Described from the surroundings of Kheirabad. Type in Leningrad. Series 2. PHYSOCARPI Gontsch.- Acaulescent or subacaulescent plants; racemes short, loosely few-flowered, subsessile or borne on a peduncle up to 10 cm long; calyx cylindric; corolla yellow or purple-tinged; wing- petals dilated in upper part; keel slightly cleft at apex; pods on a stipe 4- 18 (25) mm long, spherical to oblong-ovoid, large, strongly inflated, more or less rounded dorsally and ventrally with prominent sutures, coriaceous, glabrous, fully or partially (to 1/3) bilocular. W. Tien Shan, Pamir-Alai. 129 140. A. glabrescens Gontsch. in Fl. Tadzhik. V (1937) 664. t. 39. Perennial, acaulescent plants, (12) 17-25 (33) cm tall, fromavery short, woody, buried caudex; stipules 8-17 mm long, united at base with petiole, glabrous or sparsely ciliolate-margined, scarious, the loweroval, the up- per oblong to lanceolate; leaves (8) 13-18 (35) cm long, the petiole obsolete, the rachis glabrous; leaflets 18-25 pairs, oblong-ovate or oval, rarely triangular -lanceolate, obtusish or round-tipped, (8) 11-18 (24) mm long, (6) 7-10 mm broad, glabrous above, covered beneath all over or merely on the midvein with scattered appressed hairs or glabrous, always more or less ciliate on the margin; peduncles (5) 7-10 cm long, elongating up to 12-15 cm in fruit, glabrous; racemes short, loosely (3) 4-6-flowered; bracts lance-linear, 7-8 (10) mm long, mostly ciliate-margined; pedicels 2-4 mm long; calyx cylindric, 13-16 mm long, glabrous, ciliate only on the margin of the teeth, the tube 2.5-3.5 times the length of the teeth; co- rolla pinkish-yellow fading red; standard 23-28 mm long, the limb obovate or oval, scarcely retuse, about twice the length of the claw; wing-petals 22-26 mm long, the limb oblong, round-tipped, 2/3 the length of the claw; keel 20-23 mm long, the obtusely cuneate limb 2/3 the length of the claw; ovary glabrous, on avery long stipe; pod long-stipitate, the stipe exserted from the calyx and curved above its throat, 15-18 (25) mm long, the body oblong-ovaloid, 25-33 (37) mm long, inflated, rounded dorsally and ventral- ly, or slightly and broadly concave dorsally, coriaceous, glabrous, partial- ly bilocular (the septum 4mm broad), the acute beak 3-4 mm long. Fl. April-June; fr. May-July. Sporadically in the woodland and scrub formation, mostly in clearings of maple (Acer turkestanicum) woods or in rosaria. - Soviet Centr. Asia: Pam.-Al. (central: from Kulyak to Tupalang river basin and Gazi- mailik range). Endemic. Described from the Farak range near the bank of Pyandzh River. Type in Leningrad. 141. A. kirghisorum Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1941) 131. Perennial, 15-25 cm tall, from a strongly abbreviated, short-branched, buried caudex; stipules 5-7 mm long, membranous, sparsely white-ciliate, the lower triangular-ovate, the upper lanceolate, acuminate; leaves 12- 173 25 cm long, the petiole and the much longer rachis finely sulcate, covered with sparse spreading hairs; leaflets (13) 15-19 pairs, oval to oblong-oval, round-tipped or slightly retuse, glabrous with sparse cilia confined to mid- vein underneath and the margin; peduncles 1-2 cm long, glabrous; racemes short, loosely 3-6-flowered; bracts linear, 5-8 mm long, sparsely ciliate - margined; pedicels 5-7 mm long, glabrous; calyx 13-17 mm long, sparse- ly hirsute with spreading hairs and ciliate on the margin of the teeth, the teeth linear, 3-4 mm long; standard 22-30 mm long, the limb obovate, as long as the claw; wing-petals 21-28 mm long, the limb oblong, slightly di- lated upward, rounded-obtuse, 2/3 the length of the claw; keel 16-23 mm long, obtusely and broadly cuneate, half as long as the claw; ovary ona stipe ca. 3 mm long; pods on a stipe 4-10 mm long, not exceeding the calyx, ovoid to spherical-ovoid, strongly inflated, rounded dorsally and ventrally with prominent sutures, 2-3.5 cm long, glabrous, incompletely bilocular, the straight beak 2-5 mm long. Fl. May; fr. June. In fine-grained soil of mountain slopes, in the deciduous forest zone and in the upper part of the zone of witch-grass steppes.- Soviet Centr. Asia: 130 Tien Shan. Endemic. Described from the Ak-bura river valley. Type in. Leningrad. 142. A. virens Pavl. in Bull. Soc. Nat. Mosc., sér. Biol. XLIV, 1-2 (1935) 33. Perennial, acaulescent or nearly so, 11-22 cm tall; stipules united at base with petiole, scarious, scarcely ciliate, 5-8 mm long, the lower ovate- oblong, acute, the upper lanceolate, subulately long-acuminate; leaves (5) 8-22 cm long, the petiole 0.5-2 (3) cm long, sparsely covered with white hairs; leaflets (10) 12-17 pairs, oval or oblong-ovate, round-tipped or sub- obtuse, 7-15 (18) mm long, glabrous above and beneath or ciliate on the midvein; peduncles obsolescent, 0-1.5 cm long, glabrous; racemes short, 3-4-flowered; bracts linear-lanceolate, acuminate, 6 mm long, scarious, ciliate; pedicels 4-5 mm long, glabrous, rarely hairy; calyx cylindric, 14- 15 mm long, sparsely hirsute (especially on the teeth), rarely rather pro- fusely hirsute with spreading hairs, the teeth lance-subulate, 2-3 mm long; corolla yellow; standard 20-28 mm long, the limb oblong-obovate, round- tipped, about twice as long as the claw; wing-petals shorter than the stand- ard, the limb oblong-spatulate, half as long as the claw, with a long au- ricle at base; keel somewhat shorter than the wings, the limb cleft at the apex; ovary glabrous; pods oval, subspherical to oblong-ovaloid, inflated, with prominent sutures, 20-30 mm long, coriaceous, bilocular, the rigid stipe shorter than the ovary (5-11 mm long), the cuspidate beak 3-5 mm long; seeds smooth, ovaloid, asymmetrically emarginate, 6 mm long, com- pressed, brown, covered with dark red speckles. Fl. May; fr. June. 174 Foothills and intermediate mountain zone. - Soviet Centr. Asia: Tien Shan (Karatau range). Endemic. Described from Karatau range. Type in Moscow. Series 3. PSEUDOEREMOPHYSA Gontsch.- Plants large, acaulescent or with short stems (not more than 5 cm long); racemes loosely many- flowered, short-peduncled; calyx long-cylindric; flowers yellow; standard- limb oval upward, oblong-attenuate below; wing-petals oblong, entire; pods on a stipe 3-7 mm long, ovaloid, strongly inflated toward the back, rigidly carinate ventrally, broadly sulcate dorsally, rigidly coriaceous, velutinous-pubescent, almost fully bilocular.-S. Pamir-Alai. 143. A. pseudoeremophysa M. Pop. in Fl. Tadzhik. V (1937) 671.-Ic.: IAC. Stabues Oey fr ok Perennial plants 15-30 (40) cm tall, subacaulescent or with short stems (not more than 7 cm long), from a stout, woody, branched, buried caudex; stipules united high up with petiole, lanceolate, 5-8 mm long, white, shaggy; leaves (13) 15-30 (38) cm long, the petiole and the much longer rachis pu- bescent, rigid; leaves 8-14 pairs, orbicularornearly so, (10)13-20(30)mm long, rounded-obtuse or scarcely retuse, occasionally mucronulate, gla- brous above, densely pubescent beneath, densely pubescent-ciliate on the margin; peduncles much shorter than the leaves, densely pannose with soft pubescence, together with raceme 7-11 (17) cm long; raceme short, rarely elongated (up to 15 cm), loosely 6-10 (15)-flowered; bracts lanceolate, acute, 5-7mm long, shaggy; pedicels 2-3 mm long, shaggy; calyx 18-22 mm long, densely shaggy, the lanceolate teeth about half as long as the tube; standard 27-29 mm long, the limb oval in the upper part, oblong-attenuate below, bilobed at apex, gently angular at base, the claw ca. 6-8 mm long; 131 PLATE XIII 1, Astragalus pseudoeremophysa M. Pop.—2, A, trachycarpus Gontsch. 837, 1117 132 177 wing -petals 26-28 mm long, the limb oblong, round-tipped, about 2/3 the length of the claw; keel 24 mm long, the limb obtuse, half the length of the claw; ovary long-stipitate, shaggy; pods on a stipe 3-7 mm long, sub- cernuous, ovaloid, 17-22 mm long, thickly indurated-carinate ventrally, broadly grooved dorsally, with strongly convex valves, rigidly coriaceous, velutinous, nearly fully bilocular, the beak rigid, 3-5 (7) mm long, curved below. Fl. May-June; fr. May-July. (Piate XIII, fig. 1). Outcrops of mottled layers, gypsous detritus of red sandstone and lime- stone, outcrops of red clay, in the area of ephemeroid vegetation, both in pure composition and in composite xerophytic and ephemeroid tall-grass associations. - Soviet Centr. Asia: Pam. -Al. (S.W., foothills and low moun- tains of the watersheds of rivers Surkhan, Kafirnigan, Vakhsh and Tair- su). Endemic. Described from Sarsaryak range. Type in Leningrad. Series 4. ARENICOLAE Gontsch.-—Acaulescent plants; racemes remote - ly flowered, more or less elongated, borne on rather long peduncles; calyx cylindric; coroila yellow, at length becoming red or violet-red; standard- limb oblong or obovate-oblong, more or less angular at base, rarely ob- long-oval; wing-petals mostly rather enlarged toward the apex; pods on a rather long stipe, ovoid, spherical-ovaloidor ovaloid-oblong, convex ven- trally and dorsally, more or less shaggy, bilocular or incompletely so. Sands of plains in the north of Soviet Central Asia, S. W. Siberia, S. E. of the European part of the USSR, and western coast of the Caspian Sea, from Ciscaucasia and Volga in the west to Irtysh and the Zaisan depression in the east. 144. A. altaicus Bge. Astrag. geront. I (1868) 37; II (1869) 46; Bge. Astrag. turk. 235; Kryl. Fl. Zap. Sib. VII, 1663.-A. longiflorus Pall. Astrag. (1800) 173, pro parte, excl. pl. alt. "pr. Buchtorma, ubi magis glabrescit". Perennial, acaulescent, 15-30 (37) cm tall; stipules united in lower part with petiole, scarious, glabrous or ciliate -margined, 10-15 mm long, the lower ovate, the upper ovate -lanceolate, long-acuminate; leaves (8) 11-25 (37) cm long, the petiole 4-8 (10) cm long, both petiole and rachis gla- brous or covered with spreading hairs; leaflets 8-16 pairs, ovate-oval or obovate, (6) 10-20 (23) mm long, subobtuse, rarely slightly retuse, gla- brous or sparsely ciliate on the margin; peduncles 5-8 cm long, more or less covered with spreading hairs; racemes remotely (6) 8-14-flowered, (4) 6-10 cm long; stipules lance-linear, 7-11 mm long, whitish, scarious, slenderly pointed, ciliate-margined; pedicels glabrous, 2-3 mm long; calyx cylindric, glabrous, 13-17 mm long, the teeth linear, 3-4 mm long; co- rolla yellow; standard 22-26 mm long, the limb oblong-obovate, attenuate below; scarcely angular at base, the claw broad, 5-6 mm long; wing-petals 20-23 mm long, the limb oblong, round-tipped, the gibbous claw 12 mm long; keel 17-19 mm long, the limb gibbous below, slightly incurved above, obtusish, the claw 11-12 mm long; ovary on a stipe 3-4 mm long, glabrous; pods on a stipe (3) 4-5 mm long, ovoid, inflated, 17-22 (26) mm long, coriaceous, smooth, glabrous, bilocular (with a thin hyaline septum), shortly cuspidate - beaked, the beak ca. 2 mm long. Fl. June; fr. June-July. Sandhills, pine woods and their margins, rarely in steppe meadows. - W. Siberia: Alt. (S.W.), Irt. (Irtysh basin); Soviet Centr. Asia: Balkh. region (N.E. Zaisan depression). Gen. distr.: N. W. Mongolia. Described 133 from the Irtysh river valley in its upper reaches close to Altai, as far as Lake Zaisan. Type in Leningrad. 145. A. longiflorus Pall. Astrag. (1800) 73; Ldb. Fl. Ross. I, 650 excl. var. ; Bge. Astrag. geront. I, 36; Il, 46; Boiss, Fl. Or, Il, 283; Bge. Astrag. turk. 234; Fl. Yugo-Vost. II, 593; Kryl. Fl. Zap. Sib. VII, 1662 - Ic: Pall. Astrag. (1800) tab. 60. Perennial, acaulescent, (11) 15-25 (35) cm tall, from a short, short- branched buried caudex; stizules united in lower part with petiole, scarious, ciliate-margined, 10-13 mm long, the lower ovate to oblong, the upper broadly lanceolate-acute; leaves (5) 10-25 (35) cm long, the petiole (2) 3-5 (8) cm long, shaggy with more or less spreading hairs; leaflets 8-16 pairs, ovate to rounded-ovate (rarely oblong-ovate), (4) 8-17 (21) mm long, gla- brous, subobtuse or slightly retuse, shaggy beneath with subappressed hairs; peduncles (2) 3-11 (15) cm long, shaggy with spreading hairs; racemesre- motely 6-10-flowered, 5-7 cm long; bracts linear to lance-linear, white- hyaline, 8-11 (12) mm long, more or less hairy, profusely ciliate on the margin; pedicels 3 (5) mm long, beset with spreading hairs; calyx cylindric, 15-20 mm long, glabrous or with hairy teeth, rarely sparsely covered with spreading hairs, the teeth linear subulate-tipped 4-6 mm long; corolla yellow; standard 30-35 mm long, the limb obovate-oblong, retuse, attenuate in lower part, angular at base, the claw 6-7mm long; wing-petals 26-30mm long, the limb oblong, slightly dilated in upper part, the claw 13-16 mm long; keel 22-25 mm long, the limb convex beneath, scarcely convex above, obtuse, the claw 12-14 mm long; ovary glabrous or covered with isolated hairs, the stipe 5-9 mm long; pods on a stipe 4-6 mm long, ovate, inflated, (17) 20-23 mm long, 15-17 mm broad, convex ventrally and dorsally, with prominent sutures, nearly smooth (scarcely nervulose), rigidly coriaceous, glabrous, bilocular to 1/2-2/3, the straight cuspidate beak 2-3 mm long. FI. May-June; fr. June-July. Sands. - European part: L. V. (valleys of Volga and its tributaries), Transv. (southern part), L. Don (E.); Caucasus: Cisc; W. Siberia: U. Tob. (S.), Irt. (southern part); Soviet Centr. Asia: Ar.-Casp. (northern part, southward to Caspian Sea coast), Balkh. region (eastward to the sands of Kurman-it-kum). Endemic. Described from Volga. Type in London. 146. A. bakuensis Bge. Astrag. geront. I (1868) 37; II (1869) 44; Boiss. Fl. or. Il, 283; Grossh. Fl. Kavk. II, 317. Perennial, acaulescent, 12-20 cm tall; stipules 10-15 mm long, white- hyaline, sparsely covered with short hairs, united to 1/3-1/2 with petiole, acuminate, densely ciliate-margined, the lower ovate, the upper oblong; leaves commonly ascending, (5) 10-17 (23) cm long, the petiole much shorter than the rachis, 1-4 (7) cm long, both petiole and rachis shaggy with spread- ing hairs; leaflets 14-20 pairs, oval to suborbicular, rarely oblong-oval, (4) 5-10 mm long, round-tipped or scarcely retuse, glabrous above, softly 179 shaggy beneath; peduncles 1.5-3 (6) cm long, (2) 4-6-flowered, together with infl. 1/3-1/2 the length of the leaves, shaggy with spreading hairs; bracts rather longer than the pedicel, rarely as long, (6) 8-13 mm, linear, hispid-villous; pedicels 3-5 (6) mm long, shaggy with spreading hairs; calyx cylindric, 16-18 mm long, shaggy with spreading hairs, the linear- subulate teeth 3-4 mmlong; corolla violet (?); standard 28-30 mm long, the limb oval or obovate-oblong, 9-11 mm broad, retuse, attenuate in lower part and angular at base, the claw ca. 10 mm long; wing-petals 27- 134 80 28 mm long, the limb oblong, somewhat dilated upward, round-tipped (scarcely concave), the claw 13-15 mm long; keel 23-25 mm long, the limb obovate, rounded-obtuse, the claw 13-14 mm long; ovary on a very long stipe (7-8 mm), hairy; pod on a stipe 7-11 mm long and equaling the calyx- tube, ovaloid-obleng, inflated, dorsally and ventrally convex, subcoriaceous, 17-25 mm long, 11-13 mm broad, acute at both ends, with thick su- tures, villous with soft hairs, at length glabrous, fully bilocular, the beak ca. 2mm long. Fl. April; fr. May. Sands and more or less sandy sites. - Caucasus: Dag., E. Transc.(Apsheron Pen. and islands of the Caspian Sea). Endemic. Described from Apsheron Peninsula and islands of the Caspian Sea. Type in Leningrad. 147. A. pulposus M. Pop. in Not. Syst. Inst: Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent, up to 24-25 cm tall; stipules united with petiole, hyaline; leaves 10-25 cm long, the petiole 4-10 cm long, the petiole and rachis firm, subterete, beset with soft white short hairs; leaflets 10-14 pairs (the pairs distant), oval or obovate, short-petiolulate, obtuse, 10- 12 mm long, 7-10 mm broad, glabrous above, canescent shaggy beneath with short divergent hairs; peduncles erect, barely shorter than the leaves, together with infl. 10-15 cm long, covered with spreading white hairs; ra- cemes loosely 5-10-flowered; bracts soft, membranous, sublinear, acumi- nate, 7-9 mm long, covered with short hairs; pedicels 2-3 mm long; calyx cylindric, green, 10-12 mm long, rather sparsely covered with short white hairs, the teeth triangular-subulate, 1/5-1/4 the length of the tube, ca. 2mm long; corolla yellow, fading red; standard ca. 26 mm long, the limb oblong- oval, scarcely retuse, abruptly attenuate into theclaw, this 9 mm long; wing-petals ca. 22 mm long, the limb linear oblong, entire and rounded at apex, the claw 14 mm long; keel ca. 21 mm long, the limb strongly convex beneath (subgibbous), slightly concave above, terminating in an obtuse apex, the claw 13 mm long; ovary somewhat hairy, on a stipe 2.5 mm long; pods on a stipe 5-7 mm long, broadly oval, 20-25 mm long, not grooved, containing pulp, very sparsely hairy, mucronulate, bilocular, Fl. and fr. July. Sands. - Soviet Centr. Asia: Tien Shan (Sogoty Mountains between rivers Chilik and Charyi). Endemic. Described from Sogota. Type in Alma-Ata, cotype in Leningrad. 148. A. rubellus Gontsch. in Not. Syst. Herb. Inst. Bot. Ac. Sc. URSS X (1941) 94. Acaulescent or nearly so, (15) 30-35 cm tall; stems up to 4 cm long, angled-sulcate, white-shaggy; stipules united below with petiole, scarious, ciliate, acute, the lower triangular, the upper lanceolate, 7-12 mm long; leaves (15) 20-35 cm long; the petiole (3) 5-7 cm long, both petiole and rachis shaggy with short spreading hairs; leaflets (9) 12-25 (30) pairs, ovate to round- ed-ovate, round-tipped or slightly retuse, glabrous above, shaggy beneath with short appressed hairs; peduncles (3) 8-15 (17) cm long, rather sparsely shaggy with short spreading hairs; racemes remotely flowered, oblong, 11-18 cm long, 8-15-flowered; bracts linear, acuminate, (7) 8-10 mm long, shaggy; pedicels 2-4 mm long, shaggy with more or less spreading hairs; calyx cylindric, 17-18 mm long, more or less white-shaggy, the teeth linear, acute, 4-5 mm long; corolla red; standard 32-35 mm long, 183 the limb oblong, rounded at apex, angular at base, the claw 8-9 mm long; wing-petals 30-33 mm long, the limb oblong, rounded and spatulately di- lated upward, 2/3 the length of the claw; keel 27-30 mm long, the limb con- vex beneath, slightly concave above, obtuse, 2/3 the length of the claw; ovary on a slender stipe 9mm long, shaggy; pods inflated, ona stipe 5- 8 mm long, coriaceous, 15-20 mm long, convex dorsally and ventrally, with prominent sutures, more or less shaggy, incompletely bilocular, the cuspidate beak 1.5-2 mm long; Fl. and fr. May. Sandhills. - Soviet Centr. Asia: Syr D. (eastern part). Endemic. De- scribed from the village Kara-Kalpak in Fergana. Type in Leningrad. Series 5. BREVIALATI Gontsch.- Acaulescent plants; racemes rather many-flowered, lax, borne on developed peduncles 5-12 cm long; calyx cylindric; corolla yellow; standard oblong-obovate; wings shorter than the keel, the limb lance-oblong to linear-oblong; keel obtuse and deeply cleft, hence stamens and pistil exserted; pods on a stipe 5-8 mm long, oval-ob- long, inflated, carinate ventrally, rounded or scarcely carinate dorsally, rigidly coriaceous, incompletely bilocular. Plains of Soviet Central Asia from Balkhash region to Kara-Kum, Iran, Beludzhistan. Psammophytes. 149. A. flexus Fisch, in Bull. Phys. Mat. Ac. Pétersb. III (1844) 307 inadnot.; Bge. Astrag. geront. II, 46; Boiss. Fl. Or. II, 283; Bge. Astrag. turk., 235; Krylov, Fl. Zap. Sib. VII, 662.-A. pentapetaloides Bge. in Mém. Sav. Etr. Pétersb. VII (1851) 274.-A. stenanthus Freyn in Bull. Herb. Boiss. sér II, IV (1904) 761.-A. aquae rubrae B. Fedtsch. in Beih. Bot. Centralbl. XXII, II (1908) 352. Perennial, 18-37 cm tall; stipules 8-13 mm long, glabrous or ciliate, the lower ovate, the upper oblong; leaves (8) 12-30 (37) mm long, the peti- ole (3) 4-7 (9) cm long, glabrous or covered with spreading hairs; leaflets 7-12 pairs, rounded-ovate or rounded-obovate, (6) 8-18 (20) mm long, sub- obtuse or scarcely retuse, glabrous above, glabrous or white-shaggy be- neath; peduncles 5-12 cm long, glabrate; racemes loosely (10) 15-25- flowered, 7-15 cm long; bracts lanceolate to linear-lanceolate, 5-9 mm long; pedicels glabrous, 2-3 mm long; calyx narrowly cylindric, 14-17 mm long, glabrous or scarcely hairy, the teeth subulate-linear, 2-3 mm long; corolia yellow; standard 32-38 mm long, the limb broadly oblong, scarcely retuse, the claw 16-20 mm long; wing-petals 20-26 mm long, the limb en- tire, 1.5-2 times the length of the claw; keel 24-29mm long, the limb straight above, rounded at apex, the claw 13-17 mm long; ovary glabrous or hairy, on a stipe 6-7 mm long; style glabrous; pods (14) 16-24 mm long, slightly or scarcely rugose, obtusish, glabrous or very sparsely covered with spreading hairs, bilocular to 2/3, the beak 2-3 mm long. Fl. May-June. (Plate XIV, fig. 1). Sands and sandy deserts.- Soviet Centr. Asia: Ar.-Casp., Balkh. region, Kyz.-K., Kara-Kum, Syr-D. (sands along the Syr-D.), Pam.-Al. (sands along the Amu-D.). Gen. distr.: Iran, Beludzhistan. Described from "Skifi'', according to Pander's collection. (plant apparently collected in the sandy deserts of the Aral area). Type in Leningrad. Series 6. COMPRESSI Gontsch.- Acaulescent plants; racemes sessile or borne on short peduncles (usually not exceeding 3 cm), loosely few- flowered; calyx cylindric or campanulate-cylindric; corolla yellow or red; 136 wb YS 5 A OC? SRS ESN QA IY, Ny Ll Zs Lf OY . Sy i a) U DIRE rs 0 3 ENO Ore, =) LN Smo oD O 2 D O PLATE XIV 137 184 standard obovate or oval, rarely oval-oblong; angular at base; limb of wing- petals linear -oblong, rarely spatulately dilated in upper part; pods ovaloid or oblong, small, strongly compressed laterally, carinate ventrally, narrowly flattened or narrowly grooved (with a prominent suture) dorsally, straight, sessile or nearly so, coriaceous, often more or less shaggy (rarely ap- pressed -pubescent), incompletely bilocular. Pamir-Alai, Kopet Dagh, Caucasus, N. Iran, Turkish Armenia, Syria, Asia Minor, Crete. The section includes a number of species: A. modestus Boiss., A. pinetorum Boiss., A. nummularius Lam., etc., from Near Asia and eastern Mediterranean. 150. A. declinatus Willd. Sp.; Pl. I (1800) 1294; Bge. Astrag. geront. Il, 56; Boiss. Fl. or. Il, 295; Grossh. Fl. Kavk. II, 318 (exclusively var. suprahirsutus Freyn). Perennial, (6) 12-17 cm tall; stipules 5-8 mm long, ciliate, the lower ovate or oblong, the upper lanceolate, acuminate; leaves (3) 7-16 (19) cm long, the petiole (1) 2-4cm long, both petiole and rachis covered with spreading hairs; leaflets (14) 16-24 (32) pairs, oval, rarely oblong-oval, covered be- neath with appressed hairs, ciliate-margined, (3) 5-10 (13) mm long; peduncles (0.1) 0.5-1.0cm long, covered with spreading hairs; raceme 4-8-flowered; bracts lance-linear, 6-7 mm long, ciliate; pedicels 4-5 mm long; calyx cam - panulate-cylindric, 9-13mm long, covered with spreading hairs, the tube 2-3 times the length of the linear teeth; corolla yellow; standard 19-20 mm long, the limb obovate, retuse, 1.5-2 times the length of the claw; wing -petals 18-19 mm long, the limb linear -oblong, equaling the claw; keel 15-16mm long; ovary subsessile, shaggy; style short-hairy beneath the stigma; pods oblong-ovaloid, 10-13 mm long, shaggy with soft hairs, thickened and faint- ly sulcate dorsally, the beak ca. 2mm long. Fl. June-August; fr. July - September. Dry mountain slopes at 1800-2400m — Caucasus: E. and S. Transc., Dag. Gen. distr. : Turkish Armenia. Described from Armenia. Type in Berlin. 151. A. polyphyllus Bge. Astrag. geront. I (1868) 35; IL (1869) 59; Grossh. Fl. Kavk. II, 316. | Perennial, closely related to A. declinatus Willd. , from which it differs chiefly in the leaflets being covered above with appressed hairs and ciliate on the margin. Caucasus: Cisc. (central part), E. Transc. (Shemakha District), S. Transc. (Armenia). Endemic. Described from Altyagach. Type in Paris. Note. This species has been erroneously classified by Bunge together with A. abbreviatus Kar. et Kir., on account of the absence of pu- bescence beneath the stigma. As he notes himself "Floris unius tantum rudimenta examinavi"', the pubescence may have been rubbed off. Later collections from the classic site — Altyagach, have shown the presence of short hairs on the inner side of the style beneath the stigma, such as char- acterizes A. declinatus Willd. 152. A. talyschensis Bge. Astrag. geront. I (1868) 40; Il (1869) 57; Boiss. Fl. or. II, 297; Grossh. Mivikaviewerl lo icwne nummularius c.A.M. ex Bge. l.c. non Lam.-A. tragacanthoides Hohen. Enum. Tal. (1837) 111, non Lam. 138 Perennial, (7) 10-16 cm tall; stipules lanceolate, acute, 6-10 mm long,. hairy; leaves 5-16 cm long, the petiole (1.5) 2-3 cm long, shaggy with é short spreading hairs; leaflets (12) 18-25 pairs, oblong (rarely oval), round- ed at apex, 4-10 (14) mm long, aensely covered above with short ap- pressed hairs; peduncles 0.5-1.5 cm long, shaggy with short spreading hairs, 3-6-flowered; bracts lanceolate, 7-8 mm long, shaggy with spread- ing hairs and densely ciliate; pedicels 3-5 mm long, shaggy with short hairs; calyx cylindric, ca. 13mm long, shaggy with spreading hairs, the tube 1.5-2 times the length of the subulate-lanceolate teeth; corolla yellow; 185 standard 12-20 mm long, the limb obcordate, 2.5-3 times the length of the claw; wing-petals 18-19mmlong, the limb linear-oblong, equaling the claw; keel 14-15 mm long; ovary subsessile, shaggy; style covered with short hairs beneath the stigma; pods oblong to oblong-ovaloid, (10) 12-20mm long, narrowly grooved dorsally, shaggy, the beak 1-2 mm long. Fl. June-July; fr. (June) July-August. Stony and gravelly slopes at altitudes of 2300-3050m.-Caucasus: Tal., E. Transc. (?). Gen. distr.: N.W. Iran, Arm.-Kurd. Described from Talysh. Type in Leningrad. 153. A. Kopetdaghi Boriss. nom. nov.-A. glabriusculus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1941) 97, non Gray. Perennial, acaulescent or nearly so (stems glabrous, not more than 2cm long) 13-25 cm tall; stipules glabrous, the lower oblong or ovate, the upper lanceolate, acute, 6-9 mm long; leaves 15-25 cm long, the petiole 3-5 cm long, both petiole and rachis glabrous; leaflets 17-18 pairs, oblong or lance-oblong, acute, (8) 12-22 mm long; peduncles glabrous, 1-3 (4) cm long; raceme 6-10-flowered; bracts linear-lanceolate, 5-6 mm long, cili- ate; pedicels 5-8 mm long, glabrous; calyx cylindric, 14-16 mm long, glabrous, the teeth linear, 5-6 mm long, more or less hairy; corolla yel- low; standard 20-25 mm long, the limb broadly oblong, retuse, angular at base, 2-2.5 times the length of the claw; wing-petals 18-21 mm long, the limb linear-oblong, equaling the claw; keel 15-19 mm long; ovary on a stipe ca. 1 mm long, shaggy; style hairy above [?] the stigma; pods ob- long, 13-16 mm long, grooved dorsally, villous with soft hairs, the beak 1-2 mm long. Fl. June; fr. June. Soviet Centr. Asia: Mtn. Turkm. (W. Kopet Dagh). Endemic. De- scribed from Mount Massinev. Type in Leningrad. 154. A. balchanensis Boriss nom. nov.-A.rubriflorus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1941) 98, non Bge. Perennial, 10-14 cm tall; stipules dimorphic, the lower ovate, the upper oblong, round-tipped, sparsely ciliate; leaves 6-14 cm long, the petiole 1-2 cm long, covered with short spreading hairs; leaflets (15) 20-35 (38) pairs, oval or oblong (rarely rounded-oval), (3) 5-7 mm long, glabrous above, covered beneath with short hairs; bracts lance-linar, ca. 6 mm long, ciliate; pedicels glabrous, 3-5 mm long; calyx narrowly cylindric, 13-15 cm long, covered with scattered spreading hairs, the teeth linear - lanceolate, 3-3.5 mm long; corolla red; standard 22-25 mm long, the limb oval, entire, the claw 8-10 mm long; wing-petals 21 mm long, the limb oblong, slightly dilated above, equaling the claw; keel 20 mm long; ovary on a stipe 1-1.5 mm long, shaggy; style glabrous; pods oblong, (10) 12- 15 mm long, flattened and sulcate dorsally, rather sparsely villous with white hairs, the beak ca. 1mm long. Fl. June; fr. June. 13/9 Mountain steppes, occasionally stony slopes. - Soviet Centr. Asia: Mtn. 186 Turkm. (Bolshie Balkhany Mountains). Endemic. Described from Mount Dyuineg. Type in Leningrad. 155. A. gaudaneusis B. Fedtsch. in Not. Syst. Herb. Hort. Petrop. II (1921) 51. Perennial, acaulescent, 9-16 cm tall; stipules ovate, the upper oblong- ovate, glabrous, 7-9 mm long; leaves 7-16 cm long, the petiole (2)3-5cm long, both petiole and rachis covered with short spreading hairs; leaflets 14-20 pairs, lance-oblong or oblong-oval, round-tipped, covered beneath with short hairs, (4) 5-12 mm long, peduncles 0.2-0.4 (1) cm long, 2-3- flowered; bractslance-linear, 4mmlong, ciliate; pedicels 4-5 mm long, glabrous; calyx cylindric, 12-13 mm long, the teeth more or less hairy, subulate, 4-5 mm long; corolla red; standard 18-19 mm long, the limb ob- ovate, scarcely retuse, the claw 8-9 mm long; wing-petals 16-18 mm long, the limb rounded-spatulate, the claw 9-10 mm long; keel 15-16 mm long; ovary on a stipe ca. 1mm long, covered with appressed hairs; style gla- brous; pods ovaloid, 8-10 mm long, rounded dorsally, rather sparsely ap- pressed-pubescent, the beak ca. 1mm long. Fr. May-June. Soviet Centr. Asia: Mtn. Turkm. (Centr. Kopet-Dagh). Endemic. De- scribed from Gaudan. Type in Leningrad. 156. A. schutensis Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 126. Perennial, acaulescent, 15-16 cm tall; caudex woody, buried, divided into short ligneous branches 1.5-5 cm long; stipules united with petiole to 1/2-1/3, 6-10 mm long, membranous, glabrous, the lower ones ovate in their free part, round-tipped, the upper oblong, subacute; leaves 9-16 cm long, the rachis 2-4 times the length of the petiole, both petiole and rachis slender, angled-sulcate, covered with scattered spreading hairs; leaflets 14-25 pairs, oblong-oval, round-tipped or scarcely retuse, (4) 5-10 mm long, glabrous above, covered beneath with subappressed soft hairs; ra- cemes short, subradical, of 2-4-flowers; bracts lanceolate, ca. 4mm long, acute, thinly scarious, sparsely furnished with long white hairs; calyx cylindric, ca. 15 mm long, covered in upper part, especially on the teeth, with spreading hairs, the teeth lance-linear, half the length of the tube; corolla yellow; standard ca. 23 mm long, the limb obovate, retuse, gently angular at base, twice the length of the claw; wing-petals ca. 20 mm long, the limb oblong, slightly dilated above, rounded-obtuse, slightly shorter than the claw; keel ca. 15 mm long, the limb obtuse, half the length of the claw; style glabrous; pods sessile, lance-oblong, 12 mm long, 4-5 mm broad, acute, carinate ventrally, deeply sulcate dorsally, appressed later- ally, coriaceous, rather sparsely white-villous with spreading hairs, near- ly fully bilocular, the straight beak 3-4 mm long. Fl. June-July; fr. July- August. The lower part of the subalpine zone and the upper part of the juniper zone (chiefly in shrub thickets — rosaria). - Soviet Centr. Asia: Pam.-Al. 187 (W. Kashka Darya river basin). Endemic. Described from the village of Shut. Type in Moscow. 157. A. subrosularis Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 127. Perennial, acaulescent, (2) 5-9 cm tall, from a woody buried caudex divided into slender lignified offshoots; stipules united to the middle with 140 petiole, 4-5 mm long, white-hyaline, the free portion of the lower ovate subobtuse, of the upper oblong, acute, ciliate; leaves (1.5) 5-9 cm long, the petiole slightly shorter than the rachis, both petiole and rachis covered with spreading or ascending hairs; leaflets (10) 15-23 pairs, oblong, rarely oblong -obovate to obovate, obtusish or slightly retuse, (2) 4-6 mm long, glabrous above, softly tomentose beneath with spreading or ascending hairs; peduncles 2-4 mm long, glabrous, 2-4-flowered; bracts lanceolate, rarely linear-lanceolate, acuminate, 3-6 mm long; pedicels 1.5-3 mm long, gla- brous or thinly pubescent with spreading hairs; calyx cylindric, 9-14 mm long, glabrous or sparsely covered with spreading hairs, the tube 2-3 times the length of the narrowly linear acuminate ciliate-margined teeth; corolla yellow, often fading red; standard 18-21 mm long, the limb rounded-oval, obsoletely retuse, 1.5- 2 times the length of the claw; wing-petals 16-18 mm long, the limb oblong, entire, round-tipped, equaling or slightly shorter than the claw; keel 12-16 mm long, the limb obtuse, 1/2-2/3 the length of the claw; ovary on a stipe 1-1.5mm long, densely shaggy; pods subsessile, asymmetrically ovaloid-oblong, 8-10 mm long, 4-4.5 mm broad, inflated towards the back, carinate ventrally, rather shallowly sulcate dorsally, coriaceous, softly villous with short hairs, nearly fully bilocular, the straight beak 1-2 mm long. Fl. June. Short-grass meadows in the subalpine vegetation zone, rarely in juniper groves. — Soviet Centr. Asia: Pam.-Al. (W.) Endemic. Described from the mountains S.E. of Guzar. Type in Tashkent. 158. A. aphanassjievii Gontsch, in Fl. Tadzhik. V (1937) 665.-A. pa- miro-alaicus Lipsky in A.H.P. XXVI (1907) 147, ex parte.- A. pa- miro-alaicus M.Pop. in Sched. ad Herb. Fl. As. Med.fasc. XV, 29, 1928, non Lipsky.-—Ic.: Gontsch. l.c. tab. 39, f.6.-Exs.: H. F. A. M.No. 363. Perennial, acaulescent, (2) 4-10 cm tall, the wooden buried caudex up to 5 mm in diameter, profusely divided into more or less lignified under - ground branches 1.5-3 (5) cm long, these covered with subdistant scalelike stipules and above ground leaf-bearing and floriferous; stipules 6-8 mm long, united high up with petiole, whitish, subhyaline, the free portion ob- long-lanceolate, acute, the margin furnished with rather scattered cilia; leaves 5-8 cm long, the petiole obsolescent, both petiole and rachis covered with scattered rather spreading hairs; leaflets 12-17 pairs, linear-oblong, rounded-obtuse or slightly retuse, rounded or broadly cuneate at base, 4-7 (13) mm long, 2-4 (5) mmbroad, thickish, with a prominent midvein, gla- brous above, covered beneath with appressed hairs, ciliate-margined, rare- ly glabrate on both sides; peduncles very short, not exceeding 2-3 mm, bearing 5-6 crowded flowers; bracts lance-linear, 5-6 mm long, glabrous or ciliate-margined; pedicels glabrous, 2.5-4 mmlong; calyx cylindric, 11-12 (15) mm long, the tube glabrous, twice (rarely 1.5 times) the length of the teeth, these sparsely ciliate, rarely glabrate; corolla yellow; stand- ard 17-20 mm long, the limb oblong-obovate, scarcely retuse, gradually attenuate into a broad claw; wing-petals 16-19 mm long, the limb oblong, round-tipped, slightly shorter than the claw, rarely as long; keel 13-15 mm long, the limb obtuse, half the length of the claw; ovary short-stipitate, glabrous or white-shaggy; pod ovaloid to ovaloid-oblong, ca. 8mm long, on a stipe ca. 1 mm long, carinate ventrally, grooved dorsally, coriaceous, glabrate, or pubescent, nearly fully bilocular, the indurated beak ca. 3mm long. Fl. May-June (July); fr. June-July. 141 High mountain regions; fescue-steppe belt (Turkestan range) and the highland xerophyte vegetation belt (Hissar range). - Soviet Centr. Asia: Pam.-Al. (western part of Turkestan and Hissar mountain ranges). Endem- ic. Described from the Guralash natural border. Type in Leningrad. Series 7. INDURESCENTES Gontsch.-Cespitose, acaulescent or sub- acaulescent plants; caudex aerial, strongly abbreviated, branched; leaf petioles indurescent, partially or fully persistent, sometimes subspines- cent; leaves paripinnate, with small leaflets; flowers yellow, in few-flow- ered, subradical loose racemes; calyx cylindric; pods sessile, ovaloid to oblong-ovaloid, carinate ventrally, broadly sulcate or flat dorsally, in- completely bilocular.-W. Tien Shan, Pamir-Alai. 159. A. subinduratus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS DS, (946) 124° —lenie Iie. fig..4. Perennial, ca. 25cm tall; stems of the year short, froma short branched woody buried caudex, forming loose tufts; stems of the year 3-5 cm long, whitish, glabrous; stipules united with petiole to 1/2-1/3, 5-7 mm long, white hyaline, glabrous, the lower ovate-triangular, the upper lanceolate, finely pointed; leaves 16-20 cm long, the petiole glabrate, firmly strawlike, be- coming slightly indurated and partially persistent, but fairly flexible and sometimes not spinescent; leaflets 21-25 distant pairs, linear-oblong, (3) 5-8 mm long, subobtuse, glabrous above, sparsely and minutely appressed - puberulent, thickish; peduncles many, axillary, 1-3 cm long, slender, gla- 189 brous, the racemes loosely 3-4-flowered; bracts lance-linear, finely pointed, white-hyaline, glabrous, 5-6 mm long, about equaling the pedicel; calyx cylindric, 11-12 mm long, glabrous, the lance-subulate teeth ca. 3mm long; corolla yellow; standard 18-20 mm long, the limb subquadrangular -oval, 7 mm broad, angular at base, 3-4 times the length of the claw; wing-petals 17-19 mm long, the oblong limb slightly dilated upward, obtuse, equaling the claw; keel 14-16 mm long, acutish; ovary on a stipe 1.5 mm long, gla- brous. June. Stony river banks in the lower subalpine vegetation belt, at altitude ca. 2600 m, somewhat above the upper juniper limit.- Soviet Centr. Asia: Pam.- Al. (Kashka Darya river basin, northern slopes of Turkestan mountain range). Described from the upper reaches of river Tankhyz. Type in Leningrad. ; 160. A. talassicus M. Pop in Sched. ad Herb. Fl. As. Med. fasc. XV (1928) 21. - Perennial, acaulescent or nearly so, 6-14 (20) cm tall, the short wood caudex up to 2 cm in diameter, profusely short-branched, forming tufts; stems of the year obsolescent or strongly reduced (not exceeding 2 cm), glabrous; stipules united in lower part with petiole, ovate to broadly ovate, acute, white-hyaline, sparsely ciliate-margined, the free part 2-3 (4) mm long; leaves (2) 5-10 (16) cm long, the petiole much shorter than the rachis; petioles and rachises persistent, soft, rarely subspinescent, glabrous or scarcely short-hairy; leaflets (13) 16-22 pairs, obovate oblong-obovate or broadly oblong, rarely oblanceolate, 2-5 (10) mm long, 1.5-2.5 (3) mm broad, round-tipped or slightly retuse, almost round or broadly cuneate at base, thickish, glabrous above, covered beneath with scattered short ap- pressed hairs, very readily deciduous; peduncles (including infl. rachis) short, glabrous, not exceeding 2-4mm long; racemes loosely (1) 2-3-flowered; 142 bracts triangular-ovate, rarely ovate-lanceolate, 1.5-2 mm long, white- hyaline, acuminate, sparsely ciliate-margined; pedicels slender, glabrous, 2-3 mm long; calyx 9-10 mm long, covered with scattered appressed short white hairs or glabrate, the teeth linear-subulate, 2.5-3 mm long; corolla yellow; standard 14-17 mm long, the limb obovate to oblong-obovate, slight - ly retuse, the cuneate claw 5-6 mm long;- wing-petals 13-16 mm long, the limb oblong, round-tipped, slightly shorter than the claw; keel 12-13 mm long, the limb obtusish, ca. 4mm long, arcuately and strongly convex dor- sally, slightly convex to nearly straight ventrally; ovaryon a stipe ca.1mm long, white-hairy; pod sessile, triquetrously ovaloid-oblong, 9-12 mm long (excluding the beak), coriaceous, broadly grooved dorsally, carinate ven- trally, covered with appressed soft white hairs, bilocular to 1/2, 2-6 (8)- seeded, the beak ca. 3mm long; seeds ovaloid-reniform, 3-4 mm long, 1.5- 2mm broad, smooth, brown, lustrous. Fl. July; fr. August. 190 Gravelly and stony mountain slopes in high-mountain and more rarely in the intermediate mountain zone.- Soviet Centr. Asia: Tien Shan (Kura- minskii, Talass, Chatkal' and Uzun-Akhmatskii mountain ranges). Endemic. Described from Talass Ala Tau. Type in Tashkent. 161. A. indurescens Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. Whsse VIE £075) (1938) N61: Perennial, compactly cespitose, 8-15cm tall; caudex buried, strong, short, profusely short-branched; stipules (6) 7-9mm long, ovate or oblong- ovate, the upper lance-oblong, white-hyaline, glabrous, united with petiole in the lower 1/3-1/2; leaves (3) 5-15 (25) cm long, the very short petiole and the rachis sulcate, sparsely covered with short appressed hairs, at length lignified, subspinescent; leaflets (12) 15-25 pairs, oblong to oblong-obovate, rarely broader, suborbicular, (1) 1.5-4 (7) mm long, obtuse, conduplicate, glabrous or covered with sparse very short appressed hairs; peduncles very short, up to 5 mm long, concealed in the leaf axils, glabrous, bearing 1-3 flowers; bracts narrowly lance-linear, acuminate, 4-6 mm long, white- hyaline, sparsely ciliate-margined, exceeding the pedicel, very rarely as long; pedicels 1.5-2 (4) mm long, glabrous; calyx 10-12 mm long, glabrous or nearly so, the subulate-linear teeth 2-3 mm long, sparsely ciliate; co- rolla yellow; standard 16-18 mm long, the limb broadly oblong, rarely scarcely dilated upward, gently angular at base, 2-3 times the length of the claw; wing-petals 15-17 mm long, the limb oblong, entire, equaling the claw; keel 10-13 mm long; ovary sessile or nearly so, glabrous; pods sessile, ovaloid, rarely oblong-ovaloid, ca. 8 mm long and ca. 5 mm broad, coria- ceous, carinate ventrally, flattened dorsally, semibilocular, few-seeded, the beak 1-2 mm long. June. (Plate XV, figure 1). Stony and gravelly fine -earth slopes and screes in the subalpine vegetation zone. - Soviet Centr. Asia: Pam.-Al. (westernriver basins of Kashka- Darya, Tupalang and Sangardak). Endemic. Described from the upper reaches of Tupalanga. Type in Leningrad. Series 8. BICARINATI Gontsch. - Acaulescent plants; racemes sub- sessile or borne on Short peduncles, mostly strongly abbreviated and few- flowered, crowded at the leaf base and forming a kind of a hemispherical "inflorescence"; standard-limb oblong or broadly oblong, rarely oblong- obovate, mostly angular at base; wing-limb oblong, rarely dilated upward, 191 mostly retuse; calyx short-cylindric or campanulate-cylindric, the teeth 143 ZO yyy awe ieee “BSE MWNVIS ee 285 eZcap NN Y S DA \ A 4 és =p, 3. A. Glabrescens PLATE XV indurescens Gontsch.— 2, A, Jarmolenkoi Gontsch.— 1. Astragalus 144 93 equaling the tube or nearly so; pods sessile, ovoid or ovaloid, moreor less pointed, strongly compressed laterally, carinate dorsally and (more strongly) ventrally, unilocular or semibilocular. Western Tien Shan. Pamir-Alei. 162. A. Zaprjagaevii Gontsch. in Fl. Tadzhik. V, Appendix (1937) i 670.=Ic.:lic., tab. 41. Perennial, 30-42 cm tall; stipules (18) 20-24mm long, acute, glabrous, rarely whitish, ovate to lance-oblong; leaves 22-35 (40) cm long, the peti- | ole 11-14 cm long, both petiole and rachis covered with scattered spreading hairs; leaflets (12) 14-19 pairs, lanceolate to lance-ovate, mostly acumi- nate acute or subobtuse, (22) 25-44 (53) mm long, glabrous above, sparsely hairy beneath, rarely covered on both sides with scattered long divergent hairs, long-ciliate on the margin; peduncles 0.5-1.3 cm long, glabrous, racemes rather loose, 3.5-6.5 (7.5) cm long including peduncle; bracts linear, acute, 6-8 mm long, white-ciliate; pedicels slender, 6-18 (20)mm long, covered with scattered long spreading hairs; calyx campanulate, 14- 17 (18) mm long, the tube covered with scattered hairs, the teeth linear- subulate, (5) 6-7 mm long, densely barbate; corolla yellow, becoming red; standard 18-23 mm long, the limb broadly oblong, obtuse or scarcely re- tuse, the claw 3-5 mm long; wing-petals 17-20 mm long, the limb broadly oblong, slightly dilated upward, round-tipped or scarcely retuse; keel 15- _ 18mm long; ovary short-stipitate, densely hairy; pods 13-15 mm long, _ villous with long white hairs, nearly unilocular (the septum not exceeding 1 mm in length), the beak 4-7 mm long. Fl. June-July; fr. July-August. Rosaria and open woods of the intermediate mountain zone. - Soviet Centr. Asia: Pam.-Al. (Darvas region, upper Yagnob River). Endemic. Described from mount Vakmikuk. Type in Leningrad. 163. A. andaulgensis B. Fedtsch. in A.H.P. XXIV (1905) 213; Fl. Tadzhik. V, 367.-A. pamiroalaicus Lipsky in A.H. P. XXVI (1907) 147 (ex parte).-Ic.: Fl. Tadzhik, tab. 39, 40. Perennial, acaulescent, 7-15 (38) cm tall; stipules oblong, obtusish or (the lower) ovate, abruptly attenuate toward apex, united through rather much of its length with petiole, 13-15 mm long, glabrous or ciliate-mar- gined; leaves (6) 8-15 (36) cm long, the very short petiole and the rachis covered with scattered spreading hairs; leaflets 12-15 (20) pairs, oval or elliptic, (4) 8-16 (39) mm long, the upper face glabrous or sparsely cov- ered with appressed long hairs, the lower face covered with scattered long appressed hairs, the margin long-ciliate; racemes subsessile, strongly abbreviated, 6-8-flowered, the flowers approximate, subradical; bracts _ linear, slightly exceeding the pedicel, ciliate-margined; pedicels 5-8 mm 94 long; calyx 14-18 mm long, shaggy with rather scattered long hairs, the _ teeth linear, slightly shorter than to as long as the tube; corolla yellow; standard 20-23 mm long, the limb broadly oblong, slightly or scarcely constricted at the middle, angular at base, rarely auriculate-angular, 3 times as long as the claw; wing-petals 18-20mm long, the limb oblong, re- tuse, about equaling the claw; keel 14-16 mm long, obtuse; ovary short- stipitate; pods subsessile, ovaloid, 8-10 mm long, rather thinly villous with more or less appressed hairs, unilocular, the erect subulate beak 3mm long. Fl. June-Juiy; fr. July-August. The upper part of the woodland and scrub formation, chiefly in the high- mountain zone in subalpine meadows and mixed grass and fescue steppes. — 145 195 Soviet Centr. Asia: Pam.-Al., Tien Shan (W.). Endemic. Described from W. Tien Shan (Andaul'gen). Type in Leningrad. Economic importance. A good pasture plant, readily eaten by cattle andby horses. In flowering stage it contains 17.44% protein, 3.16% fats, 7.22% minerals, 15.94% cellulose, and 48.40% nitrogen-free extract- able substances. 164. A, sarytavicus M. Pop. in Not. Syst. Inst. Bot. Ac. Sec. URSS X (1946). Perennial, acaulescent, 12-15 cm tall; stipules oblong, ca. 10 mm long, ciliate; leaves (7) 10-13 cm long, the petiole as long as the rachis, rarely half to two-thirds as long, hispid with spreading hairs; leaflets 6-12 pairs, elliptic, rarely lanceolate, obtuse to subacute, (5) 8-13 mm long, glabrous above, hispid beneath with long hairs; flowers subradical; bracts linear, 10-14 mm long, hispid outside and on the margin with long stiff hairs; pedicels’2-3 mm long; calyx 17-18 mm long, covered with spreading stiff hairs, the tube 2-3 times as long as the linear-subulate teeth; corolla yel- low; standard 23-29 mm long, the limb oblong to oblong-obovate, slightly retuse, the claw 7-10 mm long; wing-petals 21-26 mm long, the limb line- ar, obtuse, slightly shorter than the claw; keel 19-23 mm long; ovary sessile, hairy; pods 11-14 mm long, ovoid, hispid with spreading hairs, rigidly cartilaginous-coriaceous, semibilocular, the beak ca. 4 mm long. Fl. June; fr. July. Stony mountain slopes, chiefly in the juniper zone. - Soviet Centr. Asia: Mtn. Turkm. (Kopet-Dagh). Pam.-Al. (northern slopes of Turkestan range). Endemic. Described from Sarytau mountains. Type in Tashkent. 165. A. apiculatus Gontsch. in Fl. Tadzhik. V (1937) 668.-Ic.: l.c. tab. 39,245 9: Perennial, (3) 6-8 (12) cm tall; stipules oblong, obtuse, 7-13 mm long; leaves (6) 9-10 (23) cm long, the rachis leaflet-bearing nearly from base, covered with scattered divergent hairs; leaflets (18) 20-26 (30) pairs, ovate, | rarely ovate-lanceolate or elliptic, (3.5) 6-8 (11) mm long, acute, termi- nating in a short, whitish, cartilaginous point, shaggy on both sides with soft short hairs; flowers subradical; bracts linear, 3-6 mm long, ciliate; pedicels 2-5 mm long; calyx campanulate-cylindric, 12-13 mm long, shag- gy with long rufous hairs, the teeth filiform-linear; corolla yellow; stand- ard 18-20 mm long, the limb oblong, gently constricted at the middle, retuse; wing-petals 16-18 mm long, the limboblong, dilated upward, re- tuse, 1.5-2 times the length of the claw; keel 11-12 mm long; ovary short- - stipitate (the stipe 1.5-2 mm long), shaggy; pods 6-7 mm long, rather sparsely white-villous, unilocular (the septum ca. 1 mm broad), the beak ca. 2mm long. Fl. June-July; fr. August-September. Subalpine zone, at altitude of 2700-3100 m and the upper part of the woodland and scrub formation. ~- Soviet Centr. Asia: Pam.-Al. (Darvas region, Khazret-sultan moutains, Kugifrush, Lyulichan, Imam-Asker). Endemic. Described from Khazret-sultan mountains. Type in Leningrad. 166. A. sericeopuberulus Boriss. nom. nov.-A. mollissimus Gontsch. in Fl. Tadzhik. V, tab. 39, Appendix (1937) 668. non Torr. Perennial, 7-12 cm tall; stipules oblong, 9-17 mm long, rounded at apex or acutish; leaves (7) 10-17 cm long, the petiole obsolescent or great-- ly exceeded by the rachis, both petiole and rachis shaggy with spreading hairs; leaflets 17-21 pairs, oval, rarely ovate, round-tipped or slightly 146 =~ ) | retuse, rarely subacute, (4) 8-14 mm long, sericeous on both sides with appressed hairs; flowers subradical; bracts linear, 10-12 mm long, long- ciliate; pedicels 3-4 mm long, glabrous; calyx campanulate-cylindric, 15- 18 mm long, more or less shaggy, the linear teeth equaling the tube; co- rolla yellow; standard 22-25 mm long, the limb oval-oblong, gently con- stricted about the middle, slightly retuse, angular at base, 2-3 times the length of the claw; wing-petals 20-21 mm long, the limb oblong, retuse, equaling the claw or nearly so; keel ca. 18mm long; ovary short-stipitate, shaggy; pods 10-12 mm long, white-villous, bilocular to 1/2-2/3, the beak 2mm long. Fl. May-June; fr. June. High-mountain steppes. - Soviet Centr. Asia: Pam.-Al. (Zeravshan river basin, upper reaches of Karatag River). Endemic. Described from Chandar river valley in Zeravshan basin. Type in Leningrad. 167. A. Alexeenkoi Gontsch. in Fl. TadzhikV (1937) 669. Perennial, 10-16 cm tall; stipules oval-oblong, subobtuse, glabrous, ca. 15 mm long; leaves 10-17 cm long, the rachis 1.5-2 times the length of the petiole, both petiole and rachis covered with spreading hairs; leaf- lets (17) 20-25 pairs, oblong-oval to oval, ovate or obovate, 4-6 (3) [sic] mm long, round-tipped or retuse, densely shaggy on both sides with spreading hairs; flowers subradical; bracts linear-filiform, 9-10 mm long, long- villous; pedicels 3-5 mm long, white-villous; calyx cylindric-campanulate, ca. 17 mm long, rather sparsely villous, the linear-subulate teeth equaling 196 the tube, densely long-barbate; corolla becoming red; standard 21-26 mm long, the limb oblong-pandurate, retuse, 5 times as long as the claw, the upper part narrower than the lower dilated part; wing-petals 17 mm long, the limb oblong, retuse, slightly longer than the claw; keel 13 mm long, deeply cleft at apex; ovary stipitate, white-villous; pods ca. 11 mm long, villous with long white hairs, unilocular, the beak ca. 2 mm long. Fl. June- August; fr. (June) July-August. Soviet Centr. Asia: Pam.-Al. (W. Pamir). Endemic. Described from Gushkhon canyon. Type in Leningrad. Series 9. UTRIGERI Gontsch.- Acaulescent plants; racemes strongly abbreviated, loose, borne on abbreviated peduncles 1-5 cmlong; calyx cam - panulate -cylindric; corolla yellow, fading red; standard limb ovate -orbicular; wing-limb oblong, slightly dilated upward, round-tipped, entire; podsona stipe 3-10mm long, coriaceous, strongly inflated, ovaloidto oblong-ovaloid, rounded or broadly carinate ventrally, flattened or broadly sulcate-concave dorsally, loosely villous with spreading hairs, bilocular or partially so. - Southern coast of Crimea, Black Sea coasts, Transcaucasia, Kurdistan. 168. A. utriger Pall. Astrag. (1800) 75; Ldb. Fl. Ross. I, 652 pp. ; Bge. Astrag. geront. I, 36; II, 44 pp. (specim. taurica); Grossh. Fl. Kavk. Mralie sehmalhe Flt, 274, pp. \(specim: taurica),-A. utriger fs. tay - ricus Boiss. Fl. or. II (1872) 282. Perennial, acaulescent, (10) 14-30 cm tall; stipules united below with petiole, the lower oblong-ovate, obtuse, the upper lanceolate acute, 8-15mm long, shaggy; leaves (6) 8-20 cm long, the petiole and the much longer rachis sparsely shaggy with spreading hairs; leaflets 10-15 pairs, oblong- ovate or lance-oblong, 6-15 (25) mm long, 4-6 (10) mm broad, subobtuse, glabrous above, ciliate beneath on the midvein and on the margin; peduncles 147 1-5 cm long, shaggy with stiff spreading hairs; racemes very short, loose- ly 4-6-flowered; bracts lance-linear, acuminate, 9-13 mm long, 2-4 times the length of pedicels; pedicels covered with long spreading hairs; calyx campanulate cylindric, 12-14 mm long, shaggy with spreading hairs, the tube 2-4 times as long as the lance-linear acuminate teeth; corolla yellow, at length becoming red; standard 21-22 mm long, the limb ovate-orbicu- lar, round-tipped, scarcely retuse, somewhat longer than the claw; wing- petals 19-20 mm long, the limb oblong, slightly dilated upward, round-tipped, entire, equaling the claw; keel 15-18 mm long, the limb gibbous, curved into a very short broad obtuse rostrate apex, 4-4.5 mm broad, slightly shorter than the claw; ovary on a stipe 3-4 mm long, lanate-shaggy; pods on a stipe (6) 7-10 mm long, equaling to slightly exceeding the calyx-tube, ovaloid, (1.5) 2-3 cm long, 1.2-2 cm broad, strongly inflated, nearly blad- 197dery, obtuse at both ends, abruptly attenuate into the stipe and into the straight indurated rather thick subulate (5) 6-9-mm-long beak, rounded ven- trally, mostly somewhat broadly and sulcately concave, rarely flattened dor - sally, coriaceous, sparsely spreading-villous, bilocular. Fl. April-May; fn, Way June. European part: Crimea (south coast); Caucasus: Ciscaucasia (Black Sea Region). Endemic. Described from Mount Golodnaya in Crimea. Type in London. 169. A. pseudoutriger Grossh. in Izv. Azerbaijan. fil. Akad. Nauk SSSR I (1941) 43.-A. utriger Bge. Astrag. géront. I (1868) 36; II (1869) 44, p.p;" Boiss. Fl. ‘or. I) (1872) 282°non Pall; Grossharl Kavkiei si: Perennial, acaulescent, 10-20 cm tall; stipules united in lower part with petiole, 8-11 mm long, the lower oblong-ovate rounded-obtuse, the upper lanceolate, acute, sparsely shaggy with spreading hairs; leaves (10) 11- 20cm long, the rachis several times the length of the petiole, both petiole and rachis sparsely shaggy with spreading hairs; leaflets (15) 18-25 pairs, oblong -ovate to lance-oblong, subobtuse, 8-15 mm long, glabrous above, sparsely shaggy beneath and on the margin with spreading hairs; peduncles abbreviated, 1-2 (4) cm long, sparsely shaggy with spreading hairs, loose- ly (2) 3-5-flowered; bracts lance-linear to linear, 6-7 mm long, somewhat exceeding the pedicel, shaggy with spreading hairs; calyx campanulate - cylindric, 12-14 mm long, shaggy with spreading hairs, the tube 2-3 times the length of the lance-linear acuminate teeth; corolla yellow, fading red; standard 23-28 mm long, the limb rounded-ovate, slightly and broadly emarginate, equaling the claw; wing-petals 19-23 mm long, the limb oblong, slightly dilated upward, round-tipped, equaling the claw; keel 18-22 mm . long, the limb semicircular-obovate, extending into a short obtuse rostrate beak, 2/3 the length of the claw; ovary on a stipe 3-4 mm long, lanate- shaggy; pods on a stipe 3-4 (5) mm long and exceeded by the calyx-tube, ovaloid or oblong-ovaloid, strongly inflated, 17-27 mm long, 12-18 mm broad, broadly carinate ventrally, flattened or concavely flattened dorsally, with prominent sutures, coriaceous, sparsely villous with spreading hairs, obtusish, abruptly contracted into a subulate beak 4-7 mm long, semi- bilocular. Fl. (April) May; fr. May-June (July). Stony mountain slopes (in Armenia, according to data of V. A. Transhel', in stony wormwood semidesert).-Caucasus: E. and 8. Transc., Tal. Gen. distr.: Kurdistan. Described from the surroundings of Avan, near Erevan. Type in Leningrad. 148 Series 10. LUTEIFLORI Gontsch. - Acaulescent plants; racemes strong - ly abbreviated, few-flowered, loose, subradical or borne on very short peduncles rarely reaching 5 cm in length; calyx cylindric; corolla yellow; standard oval or obovate, rarely ovate; wings slightly dilated upward; pods 198 subsessile or borne on very short stipes not more than 2 (3) mm long, ob- | | | long or oblong-oval, rarely oval, broadly carinate ventrally, flattened or slightly sulcate, rarelynearly rounded dorsally, coriaceous, villous, rarely glabrous, semibilocular to nearly fully bilocular. Steppes in the south of the European part of the U.S.S. R., in south of W. Siberia, Altai, Tarbagatai, Dzungarian Ala Tau and N. Tien Shan. 170. A. volgensis Bge. Astrag. geront. I (1868) 36; II (1869) 47; Fl. Yugo-Vost. V, 592. Perennial, acaulescent, 10-20 (30) cm tall; stipules scarious, the lower ovate, the upper lanceolate, acuminate, 8-16 mm long; leaves (7) 10-20 (30) cm long, the petiole 2-3 (5) cm long, covered with short spreading hairs; leaflets 12-17 (20) pairs, oval-oblong, oblong-ovate or oblong-oval, subobtuse, ciliate on the margin and on the midvein, glabrate, 9-14 (18)mm long; peduncles (1) 3-4 (5) cm long, 3-6-flowered; bracts hyaline, linear, acuminate, equaling the calyx-tube, 8-12 mm long; pedicels 5-6 mm long; calyx cylindric, 10-15 mm long, sparsely hairy (chiefly on the teeth), the tube 2-3 times the length of the lanceolate teeth; corolla yellow; standard 20-24 mm long, the limb ovate, slightly retuse, abruptly contracted into and twice as long as the claw; wing-petals 18-21 mm long, the limb oblong, slightly dilated upward, not gibbous above the auricle, round-tipped, equal- ing the claw; keel ca. 16 mm long, the limb gibbous beneath, concave above, obtusish, 2/3 to nearly as long as the claw; Ovary on a stipe 2-2.5 mm long, glabrous; pods subsessile (the stipe 2-3 mm long), ovaloid to oblong -ova- loid, 13 (20-27) mm long, strongly inflated, mostly rounded-convex rarely broadly carinate ventrally, mostly rounded rarely slightly flattened dorsal- ly, with prominent sutures, subobtuse, coriaceous, glabrous, nearly cir- | | | 19 } | cular in cross-section, 2/3 to nearly fully bilocular, the beak 2-3 mm long. Fl. May-June; fr. June. Steppes (mixed-grass, meadow, rarely dry), forest margins, and aspen groves in the forest-steppe zone. -European part: Zavolzh., V. Ka. (S. ). L. Don (Krasnogvardeisk, Novocherkassk); W. Siberia: U. Tob. (Cisura- lian part). Endemic. Described from the Volga and Ural basins. Type in Leningrad. 171. A. buchtormensis Pall. Astrag. (1800) 76; Ldb. Fl. Ross. I, 652; Bge. Astrag. geront. I, 36; II, 45; Bge. Astrag. turk. 234; Fl. Yugo-Vost. V, 592; Kryl. Fl. Zap. Sib. VII, 1661.-JIc.: Pall. Astrag. tab. 62, fig. A.- Exs.: HFR No. 915. Perennial, acaulescent, 12-22 cm tall; stipules scarious, ciliate, ovate, subobtuse, the upper lanceolate acuminate 7-13 mm long; leaves (8) 10- 23 cm long, the petiole 2-3 cm long, both petiole and rachis covered with short spreading hairs; leaflets (11) 18-28 pairs, elliptic to oblong, obtuse or slightly retuse, 6-13 (16) mm long, the upper face glabrous, the lower face hairy throughout or merely on the midvein; peduncles slender, 0.5-2 (5) cm long, mostly 2- rarely 3-flowered; bracts linear, exceeding the pedicel, 8-13 mm long, ciliate; pedicels 5-8 mm long, covered with short spreading hairs, rarely glabrate; calyx cylindric, 10-14 mm long, covered with white hairs throughout or merely on the teeth, the tube 2-4 times the 149 length of the linear-lanceolate acute teeth; corolla bright yellow; standard 20-24 mm long, the limb rounded-ovate, scarcely retuse, abruptly con- tracted into the claw, the claw 1.5-2 times the length of the limb; wing- petals 18-22 mm long, the limb oblong, slightly dilated upward, round- tipped, equaling the claw, not gibbous; keel 17-19 mm long, the limb acu- tish, 2/3 the length of the claw, white-hirsute, rarely glabrous, the stipe 2-3 mm long; pods subsessile (the stipe 1.5-2 mm long), oblong, rarely ovaloid, inflated, acuminate, (13) 15-20 mm long, carinate ventrally, flat or slightly and broadly sulcate dorsally, triangular in cross-section, co- riaceous, sparsely white-villous, rarely glabrous (var. gymnocarpus Trautv.), the straight beak 3-5 mm long. Fl. April-May, fr. May-June (July). Steppes.-W. Siberia: U. Tob., Irt., Alt.; Soviet Centr. Asia: Dzu. Tarb. Endemic. Described from Altai (from river Bukhtorma). Type in Leningrad. Note. An exceedingly polymorphic species with a wide distribution area. In its typical form, widespread from southern Altai to Kazakh area of low rounded isolated hills. The leaflets 17-28 pairs. A naked-fruited form (var. gymnocarpus Trautv. ) occurring in Kazakh areas of low, rounded, isolated hills, differs from the type in the glabrous ovary and fruit; from A. vogensis it differs in leaflets being 20-28 pairs and in pod shape. 172. A. Henningii (Stev.) Boriss. comb. nova.-Myobroma Hennin- gii Stev. in Bull. Soc. Nat. Mose. XXIX (1856) 151.-A. buchtormen- sis auct. Fl. Ross. europ., non Pall.-A. utriger auct. Fl. Ross. europ. non Pall. Perennial, acaulescent, (15) 20-30cm tall; stipules membranous, lanceo- late to linear-lanceolate, often filiform-acuminate, 15-18 mm long; leaves (15) 20-30 cm long, weak, disintegrating (in A. buchtormensis Pall. - leaves subobtuse, erect); leaf petioles 5-10 cm long, the petioles and rachises covered with long spreading hairs; leaflets (11) 18-22 pairs. lance- oblong to oblong, acute, sometimes subobtuse but mucronulate, hairy on both sides; leaves 2-3 times as long as the peduncles; bracts linear-fili- form, exceeding the pedicel, 8-10 mm long, hairy; calyx covered with spreading hairs, 12-15 mm long, the tube 1.5-2 times the length of the fili- form teeth; corolla yellow; standard 20-22 mm long, oblong, gradually attenuate into the claw; wing-petals ca. 18 mm long, the limb curved, with a large auricle at base; pods on a stipe 3-5 mm long, oblong, inflated, rigidly coriaceous, acuminate, densely covered with long hairs. Fl. April- may; fr. May. : 200 Steppes.- European part: Bl., L. Don., Transv. Endemic. Described from the Don. Type in Leningrad. 173. A. chlorodontus Bge. Astrag. turk. (1880) 235. Perennial, acaulescent, 15-27 cm tall; stipules scarious, ciliate, the lower ovate, subobtuse, the upper linear-oblong to linear, 6-16 mm long; leaves (11) 15-28 cm long, the petiole 2-7 cm long, the petiole and the rachis covered with soft spreading hairs; leaflets 20-22 pairs, oblong-ovate, rarely oval, round-tipped or subobtuse and mucronulate, (5) 8-18 (23) mm long, glabrous above, covered beneath with appressed hairs; peduncles (1) 2-3 (5) cm long, glabrate; racemes loosely 5-6-flowered; bracts white - hyaline-margined, medially green, lance-linear, acuminate, 6-8 mm long, about equaling the pedicel; pedicels 5-8 mm long, glabrous; calyxcylindric, 150 201 10-15 mm long, the teeth hairy, 2-5 mm long, lanceolate to linear-lanceo- late, acuminate, the tube 2-4 times the length of the teeth; corolla yellow; standard 25-29 mm long, the limb oval or obovate, scarcely retuse, abrupt- ly contracted into and 1.5-2 times as long astheclaw; wing-petals 22-24mm long, the limb oblong, slightly dilated upward, round-tipped, gibbous above the auricle, as long as the claw; keel 18-20 mm long, the limb gibbous be- neath, concave above, acute, 2/3 the length of the claw; ovary on a stipe 2- 3 mm long, glabrous; style long, curved in upper part, glabrous; pods on a stipe 2 mm long, oblong in side view, ovaloid from above, strongly in- flated, obcompressed, ca. 20 mm long, slightly carinate ventrally, slightly and broadly sulcate dorsally, straight, coriaceous, glabrous, nearly fully bilocular, the beak ca. 2mm long. Fl. April-May (June); fr. May-June. Loess foothills. - Soviet Centr. Asia: Dzu.-Tarb., Tien Shan (vicinity of Alma-Ata). Endemic. Described from Dzungarian Ala Tau. Type in Leningrad. Series 11. RADICIFLORI Gontsch. - Acaulescent radical plants; flowers subradical in strongly abbreviated 1-2-flowered racemes borne on obsoles- cent peduncles; calyx cylindric; corolla red or yellow at length turning red; standard-limb obovate; wing-limb slightly dilated upward; pods sessile, ovaloid to spherical, rarely oblong-ovaloid, rather strongly inflated, slight - ly carinate, rounded or flattened, rarely scarcely sulcate dorsally, bilocu- lar or semibilocular. Tien Shan, Pamir -Alai. 174, A. mendax Freyn in Bull. Herb. Boiss. 2 sér. IV (1904) 770. - A. pamiroalaicus Lipsky in A.H.P. XXVI (1907) 147, ex parte. - Perennial, acaulescent, 5-12 cm tall; stipules scarious, ciliate, 5- 12 mm long, the broad portion united with petiole, the lower triangular - ovate acute or rounded-ovate, the upper oblong acute; leaves (4) 6-10(12)cm long, the petiole 1-1.5 cm long, both petiole and rachis covered with short spreading hairs; leaflets (11) 14-20 pairs, oval to rounded-oval, 4-8 (10)mm long, round-tipped or scarcely retuse, glabrous above, covered beneath with scattered short appressed hairs (rarely merely ciliate on the midvein), ciliate-margined; flowers radical, in 2's, on obsolescent peduncles; bracts linear, acute, 5-6 mm long, scarious, more or less hairy; calyx cylindric, 13 mm long, covered with rather scattered spreading hairs, thetube 3times the length of the lance-linear 3-3.5-mm-long teeth; corolla yellow becoming red, or red; standard 22-24 mm long, the limb obovate, retuse, 1.5-2times the length of the claw; wing-petals 20-22 mm long, the limb oblong, slight - ly dilated upward, round-tipped, slightly gibbous above the auricle, equaling the claw; keel 17-18 mm long, the limb gibbous beneath, straight above, acutish, about half the length of the claw; ovary on a stipe not exceeding 2mm, hairy; style glabrous; pods sessile, coriaceous, white-villous, ova- loid, 11-14 mm long, carinate ventrally, flattened dorsally, inflated, bilocular or semibilocular, the beak 3-4mm long. Fl. (May) June-July; fr. July-August. Juniper groves, stony and grassy mountain slopes at altitudes of 2600- 3200 m.-Soviet Centr. Asia: Tien Shan (chiefly central), Pam.-Al. (N.). Endemic. Described from Alai (Ol'gin meadow). Type in Geneva. 175. A. Titovii Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. WSs, Villy’. 7 (1938) 156; Perennial, acaulescent, (3) 5-10 (28) cm tall, from a strong woody tap- root; stipules scarious, united below with petiole, 4-10 mm long, the lower 151 ovate round-tipped, the middle ones oblong acute, the uppermost lanceo- late, acuminate, ciliate; leaves (2) 4-13 (22) cm long, the petiole 0.5-1.5cm long, both petiole and rachis shaggy with short spreading hairs, mostly prostrate or ascending; leaflets (8) 12-19 (22) pairs, rounded -ovalto oblong - oval, (3) 4-8 (10) mm long, obtuse or round-tipped, densely shaggy beneath and above toward the margin, medially glabrate above; peduncles strongly abbreviated, bearing 1-2 flowers (flowers subradical); bracts linear -lanceo- late or subulate-linear, 5-10 mm long, acute, scarious, ciliate; pedicels 3-4 mm long, shaggy with spreading hairs; calyx cylindric, 11-14 mm long, covered with white hairs, the lanceolate teeth 3-3.5 mm long; corolla red or yellow turning red; standard 15-21 mm long, the limb obovate, round- tipped, shortly attenuate and angular at base, 1.5 times the length of the claw; wing-petals 14-20 mm long, the limb oblong, dilated upward, round- tipped, equaling or slightly shorter than the claw; keel equaling the wings, the limb gibbous beneath, slightly concave above, gently curved into a broad 202 rostrate apex, obtuse; ovary sessile or nearly so, densely villous; style hairy in lower part; pods rounded-ovaloid, rarely oblong-ovaloid, 10-15 (17) mm long, coriaceous, strongly inflated, sessile, prominently sutured, villous with rather spreading white hairs, partially or fullybilocular, slight- ly carinate ventrally, rounded or just concave dorsally, the beak 1-2 mm long. Fl. June-July; fr. June-July. Juniper groves and the subalpine zone, often on gravelly sites. - Soviet Centr. Asia: Pam.-Al. (Kugitang mountain range, Chal'bair, Kashka Darya river basin, northern slope of the Turkestan mountain range, Zeravshan mountain range), Tien Shan (Angren river basin and upper Chatkal river, Parkents region). Endemic. Described from Parkentskii district. Type in Leningrad. Series 12. RUBESCENTES Gontsch. - Short-stemmed or subacaulescent plants; racemes short, loosely few-flowered, borne on strongly abbreviated peduncles (rarely up to 9 cm long); calyx cylindric; corolla yellow, at length becoming red; standard oblong-obovate; wing-petals rounded and spatulate - ly dilated upward; pods on a short stipe 2-3 mm long, oblong-ovaloid, rare- ly ovaloid, strongly and thickly carinate ventrally, flattened and prominent - ly sutured, rarely nearly rounded dorsally, rigidly coriaceous, more or less villous, partially bilocular or nearly unilocular. Pamir-Alai. 176. A. varzobicus Gontsch. in Fl. Tadzhik. V (1937) 666.-A. ey pa - roalaicus Lipsky in A.H.P. XXVI (1910) 147, ex parte. Perennial, acaulescent, rarely with short stems up to 3-4 cm long, (10) 15-25 (33) cm tall; stipules lanceolate, 8-16 (20)mm long, ciliate -margined, the lower scalelike oval acute, shorter than the upper; leaves (11) 17-25 (35) cm long, the stipules and the longer rachises covered with rather scat- tered spreading hairs; leaflets (18) 20-22 (26) pairs, broadly oval, very rarely oblong-oval, (5) 7-14 (16) mm long, (4) 5-8 (12) mm broad, retuse and subobtuse or nearly round-tipped, glabrate above with scattered ap- pressed short hairs toward the margin, covered beneath with scattered ap- pressed short hairs, densely ciliolate-margined; peduncles 1.5-3 (9) cm long, together with infl. 5-7 (15) cm long, pinnate [?], loosely 3-4-flow- ered; bracts linear-lanceolate, acuminate, 8-10 mm long, ciliate; pedicels (5) 6-7 (8) mm long, pubescent; calyx cylindric, 14-18 mm long, sparsely covered with spreading hairs, the tube 2-2.5 times the length of the lance- 152 203 linear teeth; corolla yellow becoming red, or reddish; standard 24-25 mm long, the limb obovate to oblong-obovate, scarcely retuse, about 2.5 times the length of the claw; wing-petals 21-23 mm long, the limb oblong-obovate, round-tipped, about equaling the claw; keel 19-22 mm long, the limb nearly semicircular, obtusely cuneate-tipped, somewhat less than 2/3 the length of the claw; ovary stipitate, white -villous; pods on a stipe ca. 3 mm long, oblong-ovaloid, 15-20 mm long, rigidly carinate ventrally, rounded or near- ly flat and convexly sutured dorsally, rigidly coriaceous, villosulous, biloc- cular to 1/3-1/2, the subsubulate beak 2-3 mm long. Fl. May-June; fr. June- July. % Coppices and clearings in open maple woods and more rarely juniper woods in the woodland and scrub formation, descending into the area of contact with ephemeral vegetation; occasionally on outcrops of red sand- stone. - Soviet Centr. Asia: Pam.-Al. (southern slopes of Hissar mountain range from Varzob river basin to Ketman! -Chapty mountains, Sarsaryk range toward left bank of the Vancha River, northern extremity of Babataga). Endemic. Described from vicinity of Kharangon settlement and Varzob river basin. Type in Leningrad. 177. A. Linezevskii Gontsch. in Fl. Tadzhik. V (1937) 667, tab. 38. Perennial, subacaulescent, compactly or loosely cespitose, 17-27 cm tall, from a branched buried caudex; stipules united in lower part with peti- ole, lanceolate, subulately acuminate, 8-12 (18) mm long, white -lanate; stems strongly abbreviated, partly buried, 2-3 cm long, concealed by stip- ules and petioles of the current and (beneath) the preceding year, (10) 15- 26 cm long, the petioles and the much longer rachis covered with scattered appressed or spreading white hairs; leafets (11) 15-24 pairs, obovate, rarely obovate-oblong, more or less retuse, broadly cuneate to nearly round at base, (4) 6-12 (14) mm long, (2) 3-7 mm broad, glabrous above, covered beneath with scattered short appressed or ascending whitish hairs; peduncles 7-20 mm long, more or less pubescent, bearing 1-3 flowers; bracts linear, 5-10 mm long, hairy; calyx cylindric, 17-18 mm long, white- Shaggy, the teeth linear-subulate, ca. 4mm long; corolla yellow at length becoming red; standard ca. 30 mm long, the limb oblong-obovate to sub- oval slightly retuse, the limb 11-12 mm long; wing-petals ca. 25 mm long, the limb spatulate -oblong round-tipped entire, the claw ca. 15mm long; keel ca. 24 mm long, the limb nearly semioval, slightly concave above, attenuate to a rostrate apex, ca. 9 mm long; pods on a stipe ca. 2 mm long, obiong-ovaloid, rarely ovaloid, 16-18 mm long, prominently carinate ven- trally, more or less flattened dorsally with a broad convex suture, more or less white -villous, rigidly coriaceous, nearly fully unilocular, the sub- ulate beak ca. 1 mm long. Fr. June. Outcrops of red tertiary clays. - Soviet Centr. Asia: Pam.-Al. (S. Kulyab region, top of Khodzha-Mastan in the Karatau mountains on the right bank of the Vakhsh river, Babatag mountains). Endemic. Described from vicinity of Tirkoni Kulyab village. Type in Leningrad. Series 13. ATROVINOSI Gontsch. - Acaulescent piants; racemes sub- 04 radical, strongly abbreviated, few-flowered; calyx cylindric; corolla red or initially yellow, at length becoming red; standard obovate rounded-oval or orbicular; wings round-tipped, Spatulately enlarged; pods born on a 153 short or a long stipe, ovoid oblong-ovoid or subspherical, strongly inflated, more or less carinate ventrally, convex flattened or scarcely and broadly concave, coriaceous or rigidly coriaceous, more or less shaggy, bilocular or partially so. Foothills and low mountains of Tien Shan and Pamir -Alai. 178. A. atrovinosus M. Pop. in Journ. Turkest. Branch. Russ. Geogr. Soc. XVII (1924) 13, nomen; Pavlov in Bull. Soc. Nat. Mosc. Sér. Biol. XLIV, 1-2 (1935) 34.-A. atrovinosus Gontsch. non M. Pop. in Fl. Tadzhik. V (1937) 355, ex parte.-A. macronyx Bp. purpurinus Lipsky in A.H.P. XXVI (1907) 241. Perennial acaulescent plants (5) 7-15 (20) cm tall, from an abbreviated woody short-branched buried caudex; stipules united at base with petiole, 8-20 mm long, the free portion of the lower ones ovate or oblong, obtuse, of the upper ones lanceolate acuminate shaggy; leaves 5-16 (20) cm long, the petiole much shorter than the rachis, both petiole and rachis sparsely covered with spreading hairs; leaflets 12-25 pairs, ovate -lanceolate to lanceolate, (3) 6-12 (24) mm long, (1) 3-5 mm broad, obtuse to subacute, glabrous above, rather densely covered beneath with soft hairs; peduncles strongly abbreviated, 0.5-1 (4) cm long, sparsely covered with spreading hairs, bearing 2-3 flowers; bracts narrowly linear, 8-11 mm long, beset with short white hairs; pedicels 3-5 (6) mm long, covered with spreading hairs; calyx (18) 19-22 mm long, sparsely villous with spreading hairs, the tube (3) 4-5 times the length of the triangular -lanceolate teeth; corolla red or yellow becoming red; standard (22) 27-35 mm long, the limb rounded - oval, scarcely retuse, equaling the claw; wing -petals (20) 25-30 mm long, the limb obovate-spatulate, 2/3 the length of the claw; keel equaling the wings. | the limb incurved to a short obtuse cuneate apex, about half the length of the claw; ovary on a stipe 5-7 mm long, densely covered with appressed white hairs; pods on a stipe up to 7-8 mm long, cernuous, subspherical, 12-20 mm long, convex dorsally and ventrally with prominent sutures, coriaceous, villous with scattered long divergent hairs, bilocular, the beak 3-5 mm long. Fl. April-June. Foothills and (in the south) mountain slopes, mostly on fine -grained soils. -Soviet Centr. Asia: Tien Shan (from Karatau mountains to vicinity of Tashkent). Endemic. Described from Karatau. Type in Moscow. f 179. A. Syreitschikovii Pavl. in Bull. Soc. Nat. Mosc. Sér. Biol.XLIV, , 1-2 (1935) 3-4. Perennial acaulescent plants 5-8 cm tall, from an abbreviated short - branched buried woody caudex; stipules united at base with petiole, 4-10mm) long, scarious, the lower ovate or oblong, obtuse, the upper lanceolate, 205 ciliate-margined; leaves 4-12 cm long, the petiole much shorter than the rachis, both petiole and rachis covered with short spreading hairs; leaflets 12-22 pairs, ovate to ovate -lanceolate, 4-10 mm long, 2-6 mm broad, ob- tuse to subacute, glabrous above, densely canescent beneath with appressed white hairs; peduncles abbreviated, 0.5-1 cm long, covered with spreading hairs, bearing 2-4 flowers; bracts linear-lanceolate, acuminate, 6-7 mm long, beset with long hairs; pedicels 3-4 mm long, beset with spreading hairs; calyx cylindric, 11-12 mm long, densely hirsute with long ascending white hairs, the tube 5-7 times the length of the lance-subulate teeth; corol- la red, rarely yellow becoming red; standard 12-20 mm long, the limb ob- ovate, scarcely retuse, equaling the claw; wing-petals 11-19 mm long, the || 154 206 limb obovate-spatulate, 2/3 the length of the claw; keel equaling the wings, the limb gibbous, incurved to a short obtuse cuneate apex, 2/3 the length of the claw; ovary on a stipe 4-5 mm long, densely shaggy; pods on a stipe (4) 6-10 mm long, strongly inflated, ovoid or more often subspherical, 15- 26 mm long, rounded dorsally and ventrally, coriaceous, finely speckled with red, villous with long soft spreading hairs, incompletely bilocular, the beak 2-3mm long. Fl. April-May; fr. June. More or less gravelly or stony mountain slopes, at altitudes of 1200- 1900 m. - Soviet Centr. Asia: Tien Shan (W. Tien Shan, from Karatau Moun- tains to Little Chimgan). Endemic. Described from Uch-uzen' springs in Karatau mountains. Type in Moscow. . 180. A. substipitatus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 122. Perennial acaulescent 7-15 (20) cm tall; from a strongly abbreviated short-branched woody buried caudex; stipules united in lower part with petiole, (7) 10-16 mm long, shaggy, the lower ovate or oblong, subobtuse, the upper lanceolate acute; leaves 10-15 (20) cm long, the petiole much shorter than the rachis, both petiole and rachis sparsely covered with spreading hairs; leaflets 16-22 pairs, ovate to ovate-oblong, obtusish or round-tipped, 6-15 (20) mm long, glabrous above, shaggy beneath with sub- appressed hairs; peduncles obsolescent, 0.5-1 (2) cm long, shaggy with spreading hairs; bracts linear, 10-13 mm long, shaggy; pedicels 2-5 mm long, shaggy with spreading hairs; calyx cylindric, 15-17 (18) mm long, sparsely shaggy with spreading hairs, the subulate-linear teeth 2.5-4 mm long; corolla red or yellow becoming red; standard 23-27 mm long, the limb quadrangular-oval, scarcely retuse, equaling the claw; wing-petals 21-24 mm long, the claw obovate-spatulate, not retuse, equaling the claw; keel equaling the wings, the limb gently gibbous, incurved to an obtuse short cuneate apex, 2/3 the length of the claw; ovary on a stipe 3 (4) mm long, shaggy; pods on a stipe 1-3 mm long, ovoid to broadly oblong, obtusish, 18-27 mm long, 9-14 mm broad, indurated and thickly carinate ventrally, applanate dorsally, rigidly and thickly coriaceous, villosulous with spread - ing hairs, finely speckled with red, partially bilocular. Fl. March-April; fr. April-May. Soviet Centr. Asia: Pam.-Al. (western spurs of Hissar mountain range, Shakhimardan River basin to northern slope of Alai mountain range), Tien Shan (Mogoltau and Kuraminskii mountain ranges). Endemic. Described from Mogoltau. Type in Leningrad. 181. A. Kusnetzovii M. Pop. in Trans. Sc. Soc. Turk. I (1923) 21 in obs. Perennial acaulescent 12-35 cm tall, from a strongly abbreviated woody caudex and a strong taproot; stipules united to 1/4-1/3 with petiole, 15- 20mm long, whitish-scarious, shaggy with spreading hairs, the free por- tion of the lower ones oblong, of the upper ones linear-lanceolate, long- acuminate; leaves (7) 10-26 mm long, the petiole much shorter than the rachis, both petiole and rachis sparsely shaggy with spreading hairs, at length becoming somewhat indurated; leaflets (12) 15-18 pairs, oblong, ob- tuse, 10-16 (19) mm long, 3-5mm broad, glabrous above, densely shaggy beneath with soft ascending hairs; peduncles 1-2 cm long, beset with spread- ing hairs, bearing 2-4 distant flowers; bracts narrowly linear, acuminate, hispid; pedicels ca. 5 mm long, more or less covered with spreading hairs, 155 207 calyx cylindric, 18-23 mm long, densely villous with ascending hairs, the linear-subulate teeth 3-5 mm long; corolla yellow at length becoming red; standard 30-35 mm long, the limb suborbicular, ca. 14 mm broad, scarce- ly retuse, abruptly contracted at base into and equaling the claw; wing-petals 27-32 mm long, the limb obovate, shaggy, entire, 1/2-2/3 the length of the claw, the subovate basal auricle ca. 4mm long; keel 26-32 mm long, the limb broad, obtuse, incurved to a broad cuneate apex, half the length of the claw; ovary on a stipe 3-5 mm long, white-shaggy; pods on a stipe 3- 4 mm long, ovate, 19-25 mm long, strongly inflated, slightly obcompressed, 8-10 mm broad, 12-15 mm thick, hence oblong in side-view, ovate from beneath, carinate ventrally, flattened or broadly subconcave and prominent - ly sutured dorsally, terminating in a cuspidate beak 4-5 mm long, rigidly coriaceous, densely and minutely velutinous as well as villous with long spreading hairs, partially bilocular. Fl. April; fr. May. Outcrops of mottled layers (chiefly red sandstone, less often gypsum outcrops) in foothills and low mountains. - Soviet Centr. Asia: Pam.-Al. (southwestern spurs of Hissar and Kugitang mountain ranges to the source of the Khodzha-Ipak). Endemic. Described from the foothills of the Kugi- tang range. Type in Tashkent. 182. A. terekliensis Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS ial, tee) (OSS) PGs tioy a2. Perennial acaulescent, 15-20 cm tall; caudex woody, varying in length (up to 25 cm long), buried, clothed in remnants of old petioles and stipules; stipules united in lower part (up to 6-7 mm) with petiole, scarious, white - shaggy outside, glabrous inside, the lower oblong-ovate, 8-10 mm long, the upper lance-linear from a broad base and linear upward, 12-16 mm long, the free portion up to 10-11 mm long; leaves (10) 12-20 cm long, the peti- ole and the much longer rachis sulcate, covered with soft white spreading hairs; leaflets (19) [?] mm long and 2-3 (4) mm broad, round-tipped or re- tuse, glabrous above, shaggy beneath with soft white hairs; peduncles short (1-3 cm long in fruit), bearing 2-3 flowers; bracts lanceolate below, fili- form-linear upward, densely white-shaggy; calyx cylindric, ca. 20mm long, shaggy, the teeth linear from a triangular base, ca. 5 mm long; corolla reddish (?); standard ca. 32 mm long, the limb oblong-ovate, ca. 10 mm broad, nearly round at apex, attenuate to a claw ca. 12 mm long; wing- petals ca. 28 mm long, thelimb oblong, slightly dilated upward (by about 3 mm), entire and rounded at apex, the claw ca. 17 mm long; keel ca. 26 mm long, the limb somewhat rostrately incurved, rounded-obtuse, the claw ca. 12 mm long; pod rigidly coriaceous, ovoid to oblong-ovoid, in-- flated, 22-25 mm long, 14-15 mm broad, 16-18 mm thick, shallowly con- cave rarely flat ventrally, rigidly carinate dorsally, canescent-villous with subappressed hairs, nearly fully unilocular (septum along ventral suture ca. 2-3 mm broad), 20-26-seeded, borne on a stipe 3-5 mm long, much shorter than the calyx-tube, the indurated beak 3-4 mm long. Fl. April (May); fr. May-June. Occurring sporadically in the foothills and low mountains of S.W. Tad- zhikistan, inthe areaofbluegrass and sedge ephemeroid vegetation and tall xerophytic grasses (mountainous semidesert), chiefly in pistachio groves. Apparently associated with outcrops of mottled layers. Expectable occur - rence in Babatag (Bukhara region). - Soviet Centr. Asia: Pam.-Al. (Tere- klitau mountains and Khodza-Kadian). Endemic. Described from Tereklitau. Type in Leningrad. 156 Series 14. EPHEMEROIDEI Gontsch, - Acaulescent plants; racemes subradical, strongly abbreviated, few-flowered; calyx cylindric; corolla yellow or greenish-yellow; standard obovate; wings round-tipped or sub- obtuse and spatulately enlarged; pods short-stipitate, ovaloid or ovaloid- oblong, mostly carinate ventrally, flattened or scarcely sulcate rarely some - what convex dorsally, rigidly coriaceous, more or less shaggy or glabrate, bilocular or incompletely so; Loess foothills and low mountains (Tien Shan, Pamir-Alai, Badkhyz, Kopet Dagh, N. Iran). 183. A. rotundus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 120.-Ic.:1.c. fig. 3 : Perennial, acaulescent, 15-20 cm tall; stipules united to 1/3-1/2 with petiole, 10-16 mm long, scarious, the lower ovate-triangular subobtuse, 208the upper lanceolate acuminate, ciliate-margined; leaves (10)13-28cm long, the petiole and the much longer rachis sparsely covered with spreading short hairs; leaflets (7) 10-15 pairs, orbicular to rounded-oval, slightly broad- retuse, (10) 15-20 (25) mm long, glabrous above, rather densely shaggy beneath with appressed short hairs; peduncles obsolescent, 0.5-2 cm long, bearing 2-5 distant flowers, beset with ascending hairs; stipules linear, shaggy with spreading hairs, 10-12 mm long; pedicels 4-7 mm long, sparse- ly shaggy with spreading hairs; calyx cylindric, 13-15 mm long, densely shaggy with spreading white hairs, the linear-subulate teeth ca. 3 mm long; corolla yellow (?); standard ca. 24 mm long, the limb obovate, round- tipped, gradually attenuate at base into and 1.5 times as long as the claw; wing-petals ca. 23 mm long, the claw obovate-spatulate, entire, half the length of the claw; keel equaling the wings, the limb semioval, slightly sub- obtusely cuneate, 2/3 the length of the claw; ovary on a stipe ca. 2 mm long, shaggy; pods on a stipe 3-5 mm long, ovoid or oblong, 25-37 mm long, strongly inflated, slightly obcompressed, rounded-applanate dorsally and ventrally, 12-15 mm broad, 15-20 mm thick, rigidly coriaceous, sparsely villous with ascending hairs, bilocular. Fl. April-May; fr. May. Outcrops of mottled layers (red clays and sandstone) in foothills and low mountains. - Soviet Centr. Asia: Pam.-Al. (Surkhan river basin). Endemic. Described from the vicinity of Khodzha-Ipak. Type in Tashkent. 184. A. macronyx Bge. Astrag. turk. (1880) 236.-A. samarkandi- nus Freyn in Bull. Herb. Boiss. 2, IV (1904) 763. Perennial, acaulescent, 7-22 cm tall, from a sturdy woody strongly ab- breviated and sometimes short-branched caudex; stipules united to 1/3 with petiole, 10-16 mm long, lanceolate or (the lower) oblong, acute, commonly ciliate-margined; leaves 15-25 cm long, the rachis beset with scattered spreading hairs; leaflets thin, green, (5) 8-15 (22) mm long, ovate to ovate- oblong, linear toward the apex, subobtuse or slightly retuse, rounded to broadly cuneate at base, glabrous above, covered beneath with scattered appressed hairs, commonly sparsely ciliate-margined; peduncles very short, barely 3 mm long, bearing 1-3-flowers; bracts lance-linear, slightly exceeding the pedicel; pedicels 5-9 mm long; calyx cylindric, 15-19 mm long, glabrous or covered with scattered spreading hairs, the tube 4-5 times the length ofthe lanceolate teeth; corolla yellow, fading green; standard 32-35mm long, the limb obovate, more or lessretuse, angular atbase, about equaling to slightly exceeding the claw; wing-petals 27-33 mm long, the limb rounded -spa- tulate; keel 25-32 mm long, the limb slightly incurved intoa rostrate apex, obtuse, 2/3 as long to nearly as long as the claw; ovary short-stipitate, 157 rarely long-stipitate, glabrous or somewhat hairy; pods sessile or borne 2090n a short stipe not exceeding 1-3 mm, oblong, acutish, 24-35 mm long, strongly inflated, carinate ventrally, more or less flattened and prominent - ly sutured dorsally, cartilaginous-coriaceous, glabrous'or sparsely covered with spreading hairs, partially bilocular. Fl. April-May; fr. May. (Plate XIV, Figure 2). Perennial-ephemeroid. Foothill slopes (submontane plains in the north) in the area of ephemeroid vegetation or ephemeroid and wormwood semi- desert, penetrating in the south into the area of ephemeroid vegetation and pistachio groves. - Soviet Centr. Asia: Pam.-Al., Tien Shan. Endemic. Described from the vicinity of Tashkent. Type in Leningrad. 185." "A. Lipskyis Me Pop. in’ Not!) Syst:"ex Herbs HUBS wiv, dee (1923) 158, in adnot.-A. farctissimus Lipsky in A.H.P. XXVI (1906) 150-152, ex parte. Perennial, acaulescent, 16-25 cm tall, from an abbreviated short- branched woody buried caudex; stipules united in lower part with petiole, 7-12 mm long, shaggy with spreading hairs, the lower ovate-triangular, the upper lanceolate, acuminate; leaves (10) 14-25 cm long, the very short petiole and the rachis beset with scattered spreading hairs; leaflets 18-28 pairs, oblong lance-oblong or linear-oblong, (6) 10-15 mm long, 3-5 (6) mm broad, subobtuse or round-tipped, thin, the upper face bright green, gla- brous and sparsely hairy along the margin, the lower face sparsely covered with ascending hairs, the midvein prominent; peduncles 1-2 cm long, gla- brous, bearing 2-3 flowers; bracts linear, 8-9 mm long, shaggy; pedicels in fruit 4-5 mm long; calyx cylindric, 16-17 mm long, shaggy with ascend- ing hairs, the linear-subulate teeth 2.5-3mm long; standard ca. 28 mm long, the limb obovate to oblong-obovate, scarcely retuse, rather gradually attenuate into and about twice as long as the claw; keel ca. 22 mm long, the limb gibbous, incurved into a short obtuse cuneate apex, about half the length of the claw; pods on a stipe 4-6 mm long, ovoid to oblong-ovoid, 17- 27 mm long, more or less convex and prominently sutured dorsally and ventrally, rigidly coriaceous, glabrate (with sparse ascending long hairs), nearly fully bilocular, the straight indurated beak ca. 3 mm long; seeds ear-Shaped, compressed, ca. 5 mm long, smooth, reddish- or greenish- brown. Fr. May-June. Southern mountain slopes, often in gravelly fine-grained soil, at alti- tudes of 950-1200 m. Scarce. - Soviet Centr. Asia: Pam.-Al. (western spurs of Hissar mountain range near Shut village, Chu!'bair foothills, Tupalang river basin). Endemic. Described from the vicinity of Shut village. Type in Leningrad. 186. A. nephtonensis Freyn. in Bull. Herb. Boiss. sér. 2, IV (1905) 760.-A. diversifolius Trautv. in A.H.P. IX (1884) 445, non A. Gray (1864). 210 Perennial, acaulescent, 17-22 (25) cm tall, from a very short, short- branched, buried caudex; stipules united at base with petiole, the lower oblong to oblong-ovate subacute 8-9 mm long, the upper lanceolate from a broad base acuminate 10-15 mm long, hyaline ciliate-margined; leaves (8) 15-25 (28) cm long, the very short petiole and the rachis beset with soft spreading hairs; leaflets 10-15 (18) pairs, rarely approximate in 4's, broadly ovate, rarely oblong-ovate, scarcely retuse or more rarely round- tipped or subobtuse, (7) 10-17 (24) mm long, glabrous above, shaggy 158 13 beneath, long-ciliate on the margin; peduncles obsolescent or up to 2 cm long, glabrous or sparsely hairy, bearing 1-3 flowers; bract linear -lanceo- late, acuminate, 6-10 mm long, ciliate-margined; pedicels 3-5 (6) mm long, glabrous or covered with scattered spreading hairs; calyx cylindric, 14- 19 mm long, thinly white-villous, the tube 4-5 (6) times the length of the linear 2-4-mm-long teeth; corolla yellow; standard 23-31 mm long, the limb suborbicular to rounded-obovate, retuse, abruptly contracted into the claw, this 10-17 mm long; wing-petals 22-28 mm long, the limb spatulate -oblong, subobtusely rounded at apex, the claw 16-17mm long; keel 21-26 mm long, the limb nearly semioval, more orless concave above, obtuse, the claw 13- 19 mm long; ovary on a stipe 6-7 mm long, densely white-shaggy; pods on a stipe (3) 5 mm long, oval to oval-oblong, inflated, (20) 24-37 (40) mm long, (8) 13-17 mm broad, carinate ventrally, slightly sulcate dorsally, coriaceous, thinly white-villous, bilocular to 1/3, the beak 3-5 mm long. Fl. March-April; fr. April-May. Slopes of foothills and mountains in the upper part of the semidesert and lower part of the steppe belt. - Soviet Centr. Asia: Mtn. Turkm. (Kopet Dagh, Malye Balkhany, Badkhyz). Endemic. Described from the vicinity of Neftonovka. Type in Geneva. Series 15. EXSCAPI Gontsch.- Subacaulescent plants, rarely witha short stem not exceeding 10 cm, densely covered with silky pubescence; racemes abbreviated, few-flowered, subradical or borne ona peduncle up to 5cmlong, numerous, aggregated at the base of leaves into a sort of capitate "inflorescence''; calyx campanulate-cylindric; corolla yellow; standard oblong to obovate -oblong; wings linear-oblong; pods subsessile, oval or oblong- oval, carinate ventrally, broadly applanate or slightly sulcate dorsally, subtriquetrous, shaggy, incompletely bilocular. Southwestern European part of the USSR, Centr. Europe, and, toalesser extent, southern Europe. 187. A. excapus L. Mant. (1771) 275; DC. Astrag. 220, No. 116; Pall. Astrag. 78, ex parte; Bge. Astrag. geront. I, 35; I, 59 (excl. specim. turkest. ). Perennial, acaulescent or nearly so; stems shaggy, 1, rarely2cm, up to 10 cm long; stipules scarious, ciliate, 8-12 mm long; leaves (10) 15- 25 cm long, the petiole and rachis shaggy with spreading soft hairs; leaflets 12-19 pairs, elliptic-ovate, round-tipped, (5) 10-20 (25) mm long, seri- ceous on both sides; racemes sessile or borne on apeduncle upto 5cm long, shaggy with spreading hairs, 3-9-flowered, sessile racemes aggregated at the base of leaves into a kind of ''capitate inflorescence'' 2.5-4 (6) cm long; stipules lanceolate, shaggy; calyx campanulate-cylindric, 17-19mm long, rather densely villous with long hairs, the linear teeth equaling to slightly exceeding the tube; standard 18-28 mm long, the limb oblong to obovate-oblong, retuse, the claw 5-7 mm long; wing-petals 16-20 mm long, the limb linear-oblong, round-tipped, slightly longer than the claw; keel 14-17 mm long, the limb acutish, cleft at apex, equaling or rarely shorter than the claw; ovary on a stipe ca. 1 mm long, lanate; style woolly below; pods subsessile, ovaloid to oblong-ovaloid, 11-15 mm long, inflated, cari- nate ventrally, broadly flattened and somewhat broadly sulcate dorsally, coriaceous, softly villous, triangular in cross-section, incompletely biloc- ular, the beak 2-3 mm long. Fl. May-June: fr. June-August. 1959 {i oye r PSS 160 PLATE XVI 1, Astragalus spinescens Bge.— 2, A, Albertoregelia C, Winkl. 214 European part: Black Sea region (surroundings of Odessa). Gen. distr.: a greatly disjunct distribution in Europe, penetrating into W. and S. Europe (Spain). Described from Thuringia. Type in London. Economic importance. The roots are used locally (Elba) as a pur- gative. They contain oil, sugars and an aromatic resin. The seeds are used in Valais as a coffee substitute. Series 16. ELEGANTIFLORI Gontsch.- Small acaulescent plants; ra- cemes loose, short, few-flowered, borne on long peduncles; calyx short- cylindric; corolla yellow becoming reddish; standard suborbicular; wing- petals rounded-spatulate, constricted above the auricle; pods ovaloid or oblong-ovaloid, strongly inflated, rounded dorsally and ventrally, nearly fully unilocular, thinly coriaceous, shaggy with rather short hairs, borne on a stipe ca. 2-5 mm long.-Kopet Dagh, N. Iran. This series includes A. chrysanthus Boiss. et Hohenack. from N. Iran. 188. A. Jarmolenkoi Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Se. URSS, t. VU, f. 7 (1938) 158. Perennial, (3) 8-16 cm long; stipules 5-8 mm long, the lower ovate or oblong, obtuse, the upper lanceolate acuminate, glabrous; leaves (4) 6-14 (16) cm long, the petiole (1) 2-3 (4) cm long, covered with rather short spreading pubescence; leaflets (11) 13-19 (21) pairs, oval to oblong-oval, (2) 3-7 (10) mm long, round-tipped or scarcely retuse, covered on both sides (on the upper face chiefly toward the margin) with short curly hairs; peduncles beset with short curly hairs; racemes 2-5 (6)-flowered; bracts lanceolate, equaling to slightly exceeding the pedicel, more or less shaggy with short hairs; pedicels 2-3.5 mm long, covered with short hairs; calyx 10-13 mm long, covered with short curly hairs, the lanceolate teeth 2- 3 min long; standard 17-21 mm long, slightly retuse, the claw 5-9 mm long; wing-petals 14-29 mm long, the claw 7-9 mm long; keel 11-14 (18) mm long; ovary on a stipe ca. 3-5 mm long, hairy; pods 12-14 mm long, the beak 2mm long. Fl. June-July; fr. July. (Plate XV, Figure 2). Area of mixed-grass, needlegrass and fescue steppes and juniper groves. - Soviet Centr. Asia: Mtn. Turkm. (Centr. Kopet Dagh). Endemic. De- scribed from the summit of Chapan-Dag. Type in Leningrad. Series 17. ALEXEJANI Gontsch.- Acaulescent plants; racemes loose, short, few-flowered, borne on rather short peduncles not exceeding 4-5 cm; calyx cylindric; corolla yellow, at length becoming reddish; standard quad- rangular-oval, the base of the limb gently angular; wings slightly en- larged upward; pods on a stipe 4-5 mm long, lance-oblong, carinate ven- trally, somewhat broadly sulcate or nearly flat dorsally, rigidly coriaceous. villous, partially bilocular. Pamir-Aiai. 189. A. Alexeji Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946), Perennial, acaulescent, 20-30 cm tall; stipules united to 1/3-1/2 with petiole, 9-10 mm long, densely shaggy with woolly pubescence, the lower lanceolate, the upper linear-lanceolate, acuminate; leaves 18-30 cm long, the petiole densely velutinous-shaggy with spreading hairs; leaflets 16-20 pairs, obovate-orbicular, broadly and slightly retuse, (5) 7-12 mm long, glabrous above, densely shaggy beneath with short subvelutinous pubescence, densely ciliolate on the margin; peduncles 4-5 cm long, densely pannose- 161 shaggy with spreading hairs; racemes 3-4 cm long, remotely 3-5-flowered; bracts linear-lanceolate, acuminate, 5 mm long, villosulous, barely ex- ceeding the pannose-shaggy pedicel; calyx cylindric, 12-13mmlong, dense- ly shaggy with long spreading hairs, the tube 3 times the length of the subulate-linear teeth; corolla yellow, at length becoming reddish; standard ca. 25 mm long, the limb quadrangular-oval, scarcely retuse, obtusely angular at base, equaling the claw; wing-petals ca. 23 mm long, the limb oblong, slightly dilated upward, entire, equaling the claw; keel ca. 21 mm long, the claw 2.5 times the length of the obtuse limb; pods horizontally 215 spreading tocernuous, on a stipe 4-5 mm long, lance-oblong, 17-20 mm long, 5-6 mm broad, strongly inflated, ca. 7 mm thick, acuminate to a straight subulate beak 2-3 mm long, densely shaggy with spreading hairs, rigidly coriaceous, carinate ventrally, broadly and finely sulcate to sub- applanate dorsally, partially bilocular. Fl. April; fr. May. Outcrops of mottled layers. - Soviet Central Asia: Pam.-Al. (between Baisun and Denau). Endemic. Described from Tash-kak winter settlement. Type in Tashkent. Series 18. DOLICHANTHI Gontsch.-Acaulescent plants; leaf petioles and rachises persistent, becoming lignified; racemes short, rather loose, few-flowered, short-peduncled; calyx cylindric; corolla reddish or yellow turning reddish; standard-limb angular at base, oblong or subquadrangularly obovate; wing-petals more or less enlarged upward; pods coriaceous, fully or partially bilocular, more or less softly villous, short-stipitate, ovaloid, strongly inflated, rounded ventrally and dorsally, prominently sutured, or subapplanate dorsally. Western Pamir -Alai. 190. A. farctissimus Lipsky in A.H.P. XXVI (1910) 150.-A. Janis - chewskyi M. Pop. in Not. Syst. Herb. Hort. Petrop. IV (1923) 158. Perennial, acaulescent, 13-26 cm tall, the buried caudex giving rise to short stout branches clothed in remnants of petioles and old stipules; leaves many, 12-23 (25) cm long, the petioles and rachises persistent, in- durated, but sometimes not spinescent, profusely covered with spreading hairs at length glabrescent; stipules united at base with petiole, 10-13 mm long, the free part lanceolate acuminate densely shaggy; leaflets glauces- cent, (20) 28-36 pairs, oblong to lance-linear, rarely broader, oblong to oblong-ovate, (7) 8-10 (13) mm long, 2.5-3.5 (4) mm broad, glabrous above, covered beneath with appressed soft hairs, rather thick, obsoletely veined except for midvein slightly raised beneath in lower part; peduncles elongated (notably in fruit), (3) 5-7 cm long, together with infl. (6) 8-10cm long, about equaling or exceeded by the leaves, hairy; racemes short, 3-5-flowered; pedicels 3-5 mm long, villosulous, bracts linear-lanceolate, acuminate, subsubulate, about equaling to exceeding the pedicel, membranous, ciliate; calyx cylindric, 15 mm long, densely covered with soft hairs, the lance- subulate teeth half the length of the tube; corolla yellow, at length becoming reddish (at least the standard); standard 22-29 mm long, the limb oblong, somewhat constricted at the middle, slightly retuse, gradually attenuate at base to a short claw; wing-petals 19-27 mm long, the limb oblong-spatu- late, entire, the claw 10-15 mm long; keel exceeded by the wings, 18-24mm long, cuneately incurved, the claw 10-14 mm long; pod on an indurated stipe, ovaloid-oblong, inflated, ca. 2-3 cm long (inf. macrocarpa M. Pop. up to 5cm long), truncate with a long point, fully bilocular (indurated dorsally), 216 carinate ventrally and dorsally, shaggy with long white soft hairs; seeds 162 217 5-7 per locule, asymmetrically emarginate-ovaloid, 4.5-5 mm long, smooth, brown. Fl. May; fr. June. Mountain slopes (often stony) in the lower part of the juniper belt, de-- scending into theupper part of the witch-grass steppes. - Soviet Centr. Asia: W. Pam.-Al. (from the Guzar district and Kashka-Darya river basin to the western slope of the Zeravshan mountain range). Endemic. Described from western mountainous Zeravshan. Type in Leningrad. 191. A. subspinescens M. Pop. in Sched. ad Herb. Fl. As. Med. XV (1928) 21 in adnot.-Myobroma subspinescens Nevski in Acta Inst. Bot. Ac. Sc. URSS, sér. 1, 4 (1937) 256.-Exs.: H. F. A.M. No. 354a. Perennial, acaulescent or nearly so, 15-23 cm tall; stipules united in lower part with petiole, scarious, 13-15 mm long, ovate-lanceolate, white- hirsute or villous outside, subacute; leaves 12-20 cm long, the petiole 2- 4 cm long, both petiole and rachis covered with spreading white hairs, at length becoming lignified and subspinescent; leaflets 15-20 pairs, lance- oblong, subobtuse, becoming somewhat thickened, 6-13 mm long, glabrous above, covered beneath with sparse white short hairs; peduncles 3-6 cm long, bearing 2 flowers, glabrous, initially slender, at length becoming lignified; bracts linear to lance-linear, 5-6 mm long, white-hyaline, ciliate; pedicels 5-6 mm long, glabrate; calyx cylindric, thinly white hirsute (gla- brate), 14-16 mm long, the linear-lanceolate teeth 5-6 mm long; corolla sordid, at length becoming reddish; standard 20-22 mm long, the limb ob- ovate, scarcely retuse, angular at base, twice the length of the claw; wing- petals 18-20 mm long, the limb oblong, slightly dilated upward, round- tipped, equaling the claw, the basal auricle oblong; keel 17-18 mm long, the limb obtusish, about half the length of the claw; ovary glabrate, ona stipe 2-3 mm long; pods on a stipe 3-4 mm long, ovaloid-oblong, inflated, 22-25 mm long, carinate ventrally, somewhat broadly sulcate dorsally, thinly coriaceous, sparsely villous with rather long spreading hairs, at length glabrescent, incompletely bilocular, the indurated beak 4-5 mm long. he uly: ire July: Mountain slopes in the juniper zone. - Soviet Centr. Asia: Pam.-Al. (Kugitang range). Enaemic. Described from Kugitang range. Type in Tash- kent. Series 19. FARCTI Gontsch.- Acaulescent or nearly so; racemes ab- breviated, loose, few-flowered, borne on a short peduncle not exceeding 6 cm; calyx campanulate-cylindric; corolla reddish or yellow turning red- dish; standard oval ovate or obovate; wings mostly enlarged upward; pods short-stipitate, spherical-ovaloid to oblong-ovaloid or oblong, carinate ventrally, flattened nearly rounded or slightly sulcate dorsally, coriaceous, villous, partially bilocular. Sands of Soviet Centr. Asia (Balkhash region, Kyzyl-Kum, Syr- Darya valley). 192. A. balchaschensis Sumn. in Animadvers. Syst. ex Herb. Univ. Tomsk. No. 1-2 (1937) 4. Perennial, acaulescent, 8-16 cm tall, from a woody branched caudex; stipules white-hyaline, ciliate-margined, 8-12 mm long, the lower lanceo- late, the upper linear; leaves (5) 8-15 cm long, the petiole 2-3 (3.5) cm long, firm, white-shaggy; leaflets 13-18 pairs, rounded-ovate to rounded- obovate, rarely oblong-oval, (4) 5-8 mm long, round-tipped or scarcely 163 218 retuse, glabrous above, shaggy beneath with short white hairs; peduncles 3-4 cm long, white-shaggy, bearing 2-3 distant flowers; bracts linear, 5-6 mm long, white-hyaline, shaggy outside with white hairs; pedicels 4- 5 mm long, white-shaggy; calyx campanulate-cylindric, 12-14 mm long, shaggy with short white hairs, the linear-subulate teeth 4-5 mm long; co- rolla yellow, at length sometimes turning reddish; standard 20-23 mm long, the limb obovate, scarcely retuse, abruptly contracted into the claw, this 6-7 mm long; wing-petals 17-20 mm long, thelimb oblong, slightly dilated upward, round-tipped; keel 16-18 mm long, the limb slightly incurved to an obtuse apex, the claw 11 mm long; ovary on a stipe 2 mm long, shaggy with white hairs; pods on a stipe ca. 2 mm long, cernuous, oblong-ovaloid, rigidly coriaceous, villosulous, inflated and somewhat compressed lateral- ly, 14-18 mm long, carinate ventrally, flattened dorsally with prominent suture, bilocular, the beak 3 mm long. Fl. April-May; fr. May-June. Sandhills and sandy steppes. - Soviet Centr. Asia: Balkhash region. En- demic. Described from Balkhash region. Type in Tomsk. 193. A. farctus Bge. in Mem. Sav. Etr. Pétersb. VII (1851) 276; Bge. Astrag. geront. I, 36; II 44 pp; Astrag. turkm., 233 pp. Perennial, subacaulescent, compactly cespitose, 10-17 cm tall, from a thick abbreviated short branched buried caudex; stems strongly ab- breviated, 1-2 cm long, concealed by a tight coating of the current year's stipules as well as old stipules and petioles; stipules united in lower third with petiole, the lower triangular -ovate 6-8 mm long, the upper lanceolate to linear-lanceolate acuminate 10-15 (18) mm long, white-hyaline, shaggy with rufous hairs; leaves 7-13 cm long, the petiole one-third to half the length of the rachis, both petiole and rachis indurescent, partially persis- tent, covered with short rufous spreading hairs; leaflets 16-24 approximate pairs, oblong-oval to oblong-obovate, rarely obovate. round-tipped or slight ly and broadly emarginate, (5) 7-12 mm long, glabrous above, densely covered beneath with spreading short rufous hairs; peduncles 1-6 cm long (excluding flowers), shaggy with more or less spreading short rufous hairs, loosely (1) 2-3-flowered; bracts narrowly linear, acuminate, 8-10 mm long, rather sparsely shaggy with short spreading hairs, exceeding the pedicels; pedicels beset with spreading rufous hairs, 6-7 mm long; calyx campanulate- cylindric, 15-17 mm long, 4 mm broad, densely villosulous with spreading hairs, the subulate-linear teeth half the length of the tube; corolla ochro- leucous, at length turning reddish (?); standard 25-27 mm long, the limb ovate, round-tipped and slightly retuse, 12-13 mm broad and 1.5 times as long as the claw; wing-petals 23-25 mm long, the limb oblong-obovate, spatulate, rounded and entire at apex, 4.5-5 mm broad, equaling the claw; keel 21-22 mm long, the limb semicircular-triangular, slightly convex above, incurved to short obtuse rostrate apex, 2/3 the length of the claw; Ovary on a stipe 3-4 mm long, shaggy; pods on a stipe 3-4 mm long, strong- ly inflated nearly bladdery, spherical-ovaloid, ca. 18mm long and1.5mm broad, prominently sutured ventrally and dorsally, obtuse, coriaceous, villous with spreading soft rufous hairs, partially bilocular, the subulate beak ca. 2 mm long. Fl. May; fr. June. Sands. - Soviet Centr. Asia: Kyzyl-kum, Syr-D. (along river Syr-Darya). Endemic. Described from Kyzyl-Kum sands. Type in Leningrad. 194. A. abbreviatus Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 343; Bge. Astrag. geront. I, 35; II, 59; Bge. Astrag. turk. 231. 837, 1117 164 219 Perennial, acaulescent, 14-16 mm [sic] tall; stipules linear to lance- linear, scarious, acuminate, 9-12 mm long, shaggy with rufous hairs; leaves 10-16 cm long, the petiole 3-4 cm long, shaggy with short hairs; leaflets 14-17 pairs, oval-oblong, rarely oblong-ovate, round-tipped sub- obtuse or scarcely retuse, (5) 7-11 mm long, sericeous sparsely above and densely beneath with mostly rufous appressed hairs; peduncles shaggy, 1-3 cm long; racemes short, loosely 2-4-flowered; bracts subulate-linear to lance-linear, 3-5 mm long, scarious, more or less shaggy; pedicels 5-10 mm long, mostly shaggy with short hairs; calyx campanulate-cylindric, 13-15 mm long, rather sparsely shaggy, the linear teeth half the length of the tube; standard 18-22 mm long, the limb oval, entire, about equaling to 1.5 times the length of the claw; wing-petals 15-18 mm long, the limb line- ar-oblong, entire, about equaling the claw; keel 14-16 mm long, the limb acute, 2/3 the length of the claw; ovary lanate, ona stipe 1-1.5 mm long; pods on a stipe 1-1.5 mm long, oblong, 14-18 mm long, inflated, slightly carinate ventrally, flattened or somewhat broadly sulcate dorsally, acute, coriaceous, villous, incompletely bilocular; the beak 2-3 mm long. Fl. May. Soviet Centr. Asia: Balkhashregion. Endemic. Described from the river Ai. Type in Leningrad. Series 20. CHARGUSCHANI Gontsch. - Acaulescent dwarf plants; ra- cemes very short, 3-6 (8)-flowered, sessile or borne on a very short pe- duncle not exceeding 1 cm, subradical; calyx cylindric; corolla yellow, tending toward reddish; standard rounded-oval; wings subulately dilated toward apex; pods sessile, lance-oblong, finely carinate ventrally, broadly and deeply grooved dorsally, thinly coriaceous, villous, incompletely bilocular. Pamir, Shugnan, eastern part of Peter I mountain range. 195. A. charguschanus Freyn in Bull. Herb. Boiss. sér. II, IV (1904) 764; Fl. Tadzhik. V, 364. tab. 39.-A. pamiricus B. Fedtsch. inBull. Herb. Boiss. (1905) 315. Perennial, cineraceous, acaulescent, (3) 4-8 cm tall, from a buried caudex, the lignified elongated branches covered below with bladeless rounded-oval obtuse scales 1-5 (7) mm long; stipules united high up with petiole, oval-oblong subobtuse or oval-lanceolate subacute 10-13 mm long, the margin beset with long stiff cilia; leaves (3) 4-10 (17) cm long, spread- ing or ascending rarely erect, the petiole about equaling the rachis, both petiole and rachis beset with scattered spreading hairs, rarely glabrous; leaflets (8) 10-15 pairs, oval to elliptic-oval, retuse or round-tipped, (4) 5-7 (8) mm long, rather densely covered onbothsides with somewhat spread- ing woolly pubescence; racemes subradical or borne on a very short pe- duncle not exceeding 1 cm, 3-6 (8)-flowered; bracts linear, 5-9 mm long, hispid with rather long bristles, greatly exceeding the long glabrous pedicels; calyx cylindric, 12-17 mm long, the tube 2-2.5 times the length of the linear-lanceolate hirsute teeth; corolla grayish-yellow at length becoming reddish; standard ca. 22 mm long, the limb rounded-oval, re- tuse, angular at base, equalint the claw; wing-petals about 20 mm long, the limb spatulate-oblong; keel ca. 18 mm long, slightly incurved at apex; ovary sessile; pod sessile, lance-oblong, 16-20 mm long, acuminate to a straight slightly incurved beak 3-5 mm long, thinly carinate ventrally, broadly and deeply grooved dorsally, thinly coriaceous, villous with long hairs, incompletely bilocular. Fl. June-July (August); fr. July-September. 165 More or less stony or gravelly substrata of the high-mountain region, up to 4300 m above sea level. - Soviet Centr. Asia: Pam.-Al. (Pamir, Shugnan, eastern part of Peter I mountain range). Endemic. Described from Khargush canyon. Type in Geneva. Series 21. OOCARPI Gontsch. - Acaulescent plants; stipules united at base with petiole; leaflets lanceolate to lance-linear, rarely ovate, acu- minate; peduncles abbreviated, bearing few flowers; calyx cylindric; wings rounded and spatulately enlarged upward; pods short-stipitate, strongly inflated, spherical-ovaloid, rounded dorsally and ventrally, thinly coria- ceous, unilocular.-S. Pamir-Alai, Badkhyz, Kopet Dagh, Afghanistan. Besides the species reported, this section includes A. gompholobi- um Benth. of Afghanistan. 196. A. kuschkensis Boriss. nom. nov.-~ A. nudus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1941) 95, non Wats. Perennial, 18-24 cm tall, glabrous; stipules glabrous, sparsely cilolate, 220the lower ovate acuminate 6-8 mm long, the upper linear-oblong acuminate 11-20 mm long; leaves (8) 15-24 cm long, the petiole 3-5 cm long; leaflets lanceolate to broadly lanceolate, commonly attenuate acute, in lower leaves triangular or ovate-oblong rarely ovate, (4) 8-20 (22) mm long; peduncles 0.5-1 cm long, bearing 2-3 flowers; bracts linear-lanceolate, acuminate, 8-12 mm long; pedicels 4-7 mm long; calyx 12-17 mm long, the lanceolate acuminate teeth 3-4 mm long; corolla green; standard 28-29 mm long, the limb oval entire 11-12 mmlong; wing-petals 27-28 mm long, the claw 15- 17 mm long; keel equaling the wings, acutish, the claw 16-18 mm long; ovary on a stipe ca. 2 mm long, smooth; pods on a stipe ca. 2 mm long, spherical-ovaloid, ca. 19 mm long, glabrous, the beak ca. 2 mm long. Fl. April-May. ‘Region of ephemeroid semideserts; pistachio groves on the slopes of mountains and hills, often in more or less sandy soil.-Soviet Centr. Asia: Mtn. Turkm. (Badkhyz, Kopet Dagh). Endemic. Described from the vicinity of Kushka. Type in Leningrad. 197. A. aktauensis Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX, 3 (1941) 96. Perennial, (16) 20-30 cm tall; stipules 11-15 mm long, glabrous, the lower lanceolate, the upper linear, acute; leaves (15) 20-30 cm long, the the petiole (3) 4-6 cm long, both. petiole and rachis velutinous with short pubescence, rarely glabrous; leaflets lance-linear, minutely velutinous - beneath, rarely glabrous; peduncles bearing (2) 3-4 flowers, (2) 3-5 cm long, minutely velutinous, rarely glabrous; bracts linear-lanceolate, subu- late-tipped, 7-10 (13) mmlong, sparsely ciliate; calyxca. 14 mm long, the teeth subulate from a triangular base, 4-5 mm long; corolla yellow; standard ca. 26 mm long, the limb oval, entire, the claw ca. 10 mm long; wing-petals ca. 24 mm long, the limb slightly shorter than the claw; keel ca. 23 mm long, the limb acutish, about 2/3 the length of the claw; ovary velutinous with short hairs, on a stipe ca. 2 mm long; pods on a stipe 2mm long, ovaloid to rounded -ovaloid, (12) 15-20 mm long, rounded-obtuse at both ends, very minutely velutinous with white hairs, beak 1-2 mm long. Bla sune. In the juniper zone, on stony fine-textured slopes sparsely covered with juniper and maple (Acer Fedtschenkoanum Krysht). - Soviet Centr. 166 2 ? 21 22 Asia: Pam.-Al. (S.W., Aktau range). Endemic. Described from Aktau mountains (Tadzhikistan). ‘ype in Leningrad. Series 22. ERIONOTOPSIDES Gontsch. - Acaulescent or subacaulescent plants; leaves floccose or arachnoid above; racemes abbreviated few-flow- ered, sessile or borne on short peduncles (usually not exceeding 3 cm); co- rolla yellow, whitish or red; calyx cylindric; standard obovate to oblong- obovate; wing-petals oblong, more or less enlarged upward, more or less retuse; pods subsessile, oblong rarely ovaloid or oblong-ovaloid, inflated, slightly compressed laterally, carinate ventrally and dorsally, coriaceous, villous or shaggy-pubescent, unilocular.-S. Pamir-Alai, Badkhyz, Kopet Dagh. 198. A. angustidens Freyn et Sint. in Bull. Herb. Boiss. sér. II, IV (1904) 758.-Exs.: Sintenis "Iter 1900-1901" Nos. 243, 1708. Perennial, (8) 15-25 cm tall; stipules 8-16 mm long, ciliate, the lower ovate, the upper lanceolate to lanceolate-linear; leaves (5) 10-28 cm long, the petiole 1.5-6.5 cm long, both petiole and rachis shaggy with soft spread- ing hairs; leaflets 16-23 (25) pairs, rounded-elliptic, oval or oblong-ellip- tic, rarely broadly oblong, (4) 6-12 (15) mm long, covered beneath with more or less appressed hairs, obtuse; peduncles 0.5-2 (3) cm long, shaggy; bracts linear, subfiliform upward, long-ciliate, (3) 4-5 (7) mm long; ped- icels 4-5 (8) mm long, shaggy with short hairs; calyx cylindric, 11-16mm long, covered with scattered spreading hairs, the tube equaling the fili- form -linear teeth, rarely 1.5 times as long; standard 18-22 mm long, the limb obovate to oblong-obovate, more or less retuse, attenuate and angular below, the claw 4-6 mm long; wing-petals 16-21 mm long, the claw 7-8mm long; keel 11-15 mm long, obtuse; ovary stipitate, sericeous, the style hairy below; pods ellipsoid-oblong, rarely ellipsoid, 9-13 mm long, more or less villous, the beak 2-3 mm long. Fl. May; fr. June. Mountain slopes in the witch-grass steppe belt. - Soviet Centr. Asia: Mtn. Turkm. (W. Kopet Dagh penetrating into Centr.). Endemic. Described from the surroundings of Ashkhabad. Type in Geneva. Cotype in Leningrad. 199. A. Barrowianus Aitch. et Baker in Trans. Linn. Soc. sér. II (1886) 50. Perennial, cespitose, (9) 15-22 (30) cm tall; stipules ovate, the upper lance-linear, ciliate; leaves (4) 8-21 (29) cm long, the petiole (0.6) 1-3 (4) cm long, both petiole and rachis beset with spreading rufous hairs, at length indurescent and persistent; leaflets densely covered beneath with appressed hairs, (22) 26-36 (40) pairs, oblong to oblong-oval, (2) 3-8 (10)mm long, obtuse or round-tipped; peduncles 1-3 (5) cm long, shaggy with spreading hairs; bracts (3) 4-6 (7) mm long, linear, filiform upward, cili- ate; pedicels 3-5 mm long, shaggy; calyx short-cylindric, densely villous - hirsute, the tube equaling or 1.5 times as long as the linear-subulate teeth; standard 16-22 mm long, the limb ovate, retuse, attenuate and angular below, the claw 4-7 mm long; wing-petals 14-20 mm long, the limb linear- oblong, scarcely enlarged upward, about equaling to slightly exceeding the claw; keel 11-16 mm long; ovary lanate, on a stipe 3-4 mm long, the style hairy below; pods oblong to ellipsoid-oblong, 13-20 mm long, acute, dense- ly pubescent, the beak 1-2 (3) mm long. Fl. April; fr. (April) May. Stony, gravelly or clay mountain slopes. - Soviet Centr. Asia: Mtn. Turkm. (Badkhyz), Pam.-Al. (S.W.). Endemic. Described from Badkhyz. Type in London. 167 223 200. A. curvipes Trautv. in A.H.P. IX (1886) 446.-A. supralana- tus Freyn in Bull. Herb. Boiss., sér. II, IV (1904) 768.-Exs.: Sintenis "Iter 1900-1901", No. 602 a-b (sub A. supralanato Freyn). Perennial, compactly cespitose, 8-20cm tall; stipules dimorphic, the lower ovate-lanceolate, acute, the upper linear from a broad base, subfili- form-acuminate, densely hairy; leaves (5) 8-15 (18) cm long, the petiole 1.5-2 (5) cm long, both petiole and rachis shaggy with spreading short hairs, at length glabrescent, indurescent and persistent; leaflets (22) 24-32 (38) pairs, oval to rounded-oval, (2) 3-5 (5.5) mm long, subobtuse, retuse or mucronulate, shaggy beneath with appressed hairs; peduncles 2-5 (10) mm long, densely covered with spreading hairs, bearing 1-2 (3) flowers; bracts lanceolate, 5-6 mm long, filiform-acuminate, covered with scattered spread- ing hairs, ciliate; pedicels 3-5 mm long, beset with spreading hairs; calyx narrowly cylindric, 16-24 mm long, the teeth filifofm, villous with spread- ing hairs, 4-10 mm long, thetube 1.5-3 times the length of the teeth; stand- ard 26-34 mm long, the limb oblong-obovate, round-tipped, scarcely re- tuse, attenuate and scarcely angular below, the claw 5-8 mm long; wing- petals 23-32 mm long, the claw 9-16 mm long; keel 23-30 mm long; ovary short-stipitate or subsessile, hairy; style hairy below; pods ovaloid to ob- long-ovaloid, 9-16 mm long, white-villous, the beak 2-5 mm long. Fl. May; fr. June. Stony or gravelly mountain slopes. - Soviet Centr. Asia: Mtn. Turkm. (Centr. Kopet Dagh). Endemic. Described from Kopet Dagh. Type in Lenin- grad. Series 23. TRACHYCARPI Gontsch. - Large plants with short stems; racemes loosely few-flowered, subsessile or short-peduncled; calyx cy- lindric; corolla compressed, large; standard oblong; wing petals oblong; pods subsessile, strongly compressed laterally, quadrangular -ovate or rhomboid-ovate, rarely rhomboid-oblong, carinate dorsally and ventrally, rigidly coriaceous, strigose-villous incompletely bilocular. - Pamir -Alai. 201. A. trachycarpus Gontsch. in Fl. Tadzhik. V, Appendix (1937) 672. Perennial; stems 1-16 (22) cm long, densely white-pannose; stipules broadly ovate, the upper oblong-ovate, ciliate, 15-22 mm long; leaves 24- 36 cm long, the very short petiole andthe rachis hispid; leaflets (10) 14- 17 pairs, rounded-oval to lanceolate, round-tipped, rarely retuse or sub- acute, covered with scattered appressed hairs; racemes (6) 8-12 (rarely more)-flowered, 6-12 cm long; bracts lanceolate, 7-10 mm long, ciliate; pedicels 2-3 mm long, shaggy; calyx strigose-villous, 14-19 mm long, the linear teeth half the length of the tube; standard-limb oblong, obtuse or scarcely retuse, 3 times the length of the claw; wing-petals 30-35 mm long, the limb oblong, equaling the claw; keel 26-28 mm long; ovary subsessile, white-lanate; pods (15) 16-19 (22) mm long, subsessile, the beak (2) 3-6 (7) mm long; Fl. May-June (July); fr. June-July (August). (Plate XIII, Fig- ure 2), Stony and pebble-covered terraces at altitudes of 1400-2700 m. - Soviet Centr. Asia: Pam.-Al. (Centr. and E. Tadzhikistan). Endemic. Described from Kichik-Terai range. Type in Leningrad. Series 24. SUBVERTICILLATI Gontsch. - Leaflets verticillately pinnate; standard-limb oval-oblong, angular at base. 168 202. A. subverticillatus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, acaulescent, 10-20 cm tall; stipules united to 1/3-1/2 with petiole, 10-16 mm long, thinly scarious, white-shaggy, the lower oblong- ovate subobtuse, the upper narrowly lanceolate acuminate; leaves 7-20 cm long, the petiole and the much longer rachis shaggy with scattered (rarely dense) short hairs; leaflets in (12) 15-20 distant verticels of 4 (6), oval to oblong-oval, round-tipped or scarcely retuse, 5-10 (13) mm long, glabrous above, villous-hirsute beneath with appressed short hairs; peduncles 1.5- 3 (4) cm long, sparsely (rarely densely) shaggy with short spreading hairs, bearing 2-4 distant flowers; bracts lance-linear, acuminate, 5-6 mm long, shaggy with ascending hairs; pedicels ca. 5 mm long, shaggy with short spreading hairs; calyx cylindric, 17-20 mm long, densely spreading -villous, the subulate-linear teeth (4) 4-7 mm long, half the length of the tube; co- rolla reddish-yellow, at length red; standard 23-28 mm long, the limb sub- quadrangularly oval-oblong, round-tipped and scarcely retuse, angular or auriculate-angular, 9-10 mm broad, slightly exceeding to 1.5 times the length of the claw; wing-petals barely shorter than the standard, the limb obovate -spatulate, entire, 2/3 as long as to nearly as long as the claw, the basal auricle oblong ca. 3 mm long; keel 21-23 mm long, the limb nearly semicircular, curved into a broad shortly cuneate apex, 1/2-2/3 the length of the claw; ovary on a stipe 3-4 mm long, densely shaggy; pods on a stipe 2-3 mm long, ovate in ground-view, strongly inflated, oblong in side-view, (17) 20-27 mm long, slightly obcompressed, 8-10 mm broad, 14-15 mm thick, coriaceous, thinly villosulous with scattered soft hairs, at length glabrescent, bilocular, the straight beak 2-3 mm long. Fl. April-May; fr. (April) May-June. Gravelly or stony mountain slopes, often on sandstone, in the lower and intermediate mountain zones. - Soviet Centr. Asia: Pam. -Al. (southwestern spurs of Hissar mountain range, Kugitang range, Chul'bair and Ketmen!'- Chapty mountains). Endemic. Described from Chul'bair mountains. Type in Tashkent. Series 25. SEMIGLABRI Gontsch. - Leaflets verticillately pinnate, gla- brous above; standard-limb auriculate-angular at base, more or less at- tenuate in lower part. 203. A. alatavicus Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 344; Bge. Astrag. geront. II, 42; Bge. Astrag. turk. 232. Perennial, acaulescent or nearly so, (4) 7-12 (20) cm tall, forming small tufts; stipules united in lower part with petiole, membranous, red- dish, glabrous, ciliate-margined, (5) 8-12 mm long, the lower ovate ob- tuse, the upper oblong-lanceolate acuminate; leaves (3) 4-12 (22) cm long, the rachis 1.5-4 times as long as the petiole, both petiole and rachis rather densely shaggy with short spreading hairs; leaflets in 10-17 verticils of 4-8, oblong to oval-oblong, very rarely oval, round-tipped, (2) 3-8 (10)mm long, (1.5) 2-3 mm broad, glabrous above, sparsely shaggy beneath with spreading hairs; peduncles 2-6 (15) mm long, bearing 3-5 flowers, shaggy with spreading hairs; bracts lance-linear, acuminate, 4-8 mm long, ex- ceeding or rarely equaling the pedicel, ciliate; calyx 10-14 mm long, gla- brate or more or less densely villous withdivergent hairs, the tube 2-4 times the length of the linear teeth; corolla yellow, at length becoming reddish; 169 227 standard 20-26 mm long. the limb obovate to oblong-obovate, round-tipped and retuse, constricted above the lower third, acutely angular toward base, the claw 4-8 mm long; wing-petals 19-24 mm long, the limb oblong, more or less enlarged in upper part, asymmetrically emarginate or nearly en- tire, 2/3 the length of the claw; keel 18-22 mm long, obtusish; ovary on a stipe 2-2.5 mm long, densely woolly; pods oblong, ca. 10 mm' long, ona stipe ca. 2 mm long, carinate ventrally, broadly grooved dorsally, puberu- lent with ascending hairs, thebeak2mmlong. Fl. June-July; fr. July-August. In the forest region and chiefly in the subalpine and alpine zones, mostly on more or less gravelly or stony slopes. - Soviet Centr. Asia: Dzu-Tarb. (Dzungarian Ala Tau), Tien Shan, (except extreme western), Pam.-Al. (Alai valley). Endemic. Described from the Sarkan and Lepsa rivers in Dzun- garian Ala Tau. Type in Moscow, cotype in Leningrad. 204. A. myriophyllus Bge. Astrag. turk. (1880) 233.-A. pamirensis Franch. in Bull. Mus. Hist. Nat. Par. II (1896) 344. Perennial, subacaulescent, rarely with stems up to 5 cm long, (6) 10- 22 cm tall; caudex strongly abbreviated, short-(rarely long-) branched, buried, forming rather compact tufts; stipules united in lower part with petiole, scarious, villous-ciliate on the margin, 7-12 mm long, the lower ovate obtuse, the upper oblong to lance-oblong subacuminate; leaves (5) 8-21 cm long, the petiole 2-5 cm long, the rachis (1.5) 2-3 times as long as the petiole, both petiole and rachis villous-tomentose with spreading hairs; leaflets in (8) 12-20 verticels of 4-6, oval-oblong or oblong-oval, round-tipped, 5-9 mm long, 2-4.5 mm broad, glabrous pale green above, appressed-villous canescent beneath; peduncles (1) 2-5 cm long, densely villous-tomentose with spreading hairs, rarely obsolescent; racemes short, 4-6 cm long, loosely 5-10 (15)-flowered; bracts narrowly lanceolate, long- acuminate to a fine point, 4-6 mm long (rarely lance-linear and up tol10Omm long), villous-ciliate on the margin; pedicels 3-4 mm long, densely villous - tomentose with spreading hairs; calyx 13-14 mm long, densely villous- tomentose, the linear teeth 3-4 mm long; corolla yellow, at length becoming reddish; standard 21-22 mm long, the limb subquadrangular-obovate, round- tipped and retuse, somewhat attenuate in lower third, angularly dilated and auriculate at base, the claw 4-5 mm long; wing-petals 20-21 mm long, the limb oblong (rarely somewhat dilated in upper part), round-tipped, entire, 2/3 to nearly as long as the claw; keel 17-18 mm long, obtuse; pods ona stipe ca. 2 mm long, semioval-oblong, 14-17 mm long, asymmetrically acute, carinate ventrally, grooved dorsally, erect, thinly coriaceous, densely villous-tomentose with spreading hairs, partially bilocular, 8-10- seeded, the beak 2-3 mm long. Fl. June. Sands, rarely pebbles, or more or less sand-covered sites in the high- mountain zone. - Soviet Centr. Asia: Pam.-Al. (Alai valley, E. Pamir). Endemic. Described from Alai valley. Type in Leningrad. Series 26. PILOSISSIMI Gontsch. - Leaves verticillately pinnate; leaf- lets hirsute or villous on both sides with ascending hairs; standard-limb more or less constricted in lower part, auriculate-angular at base. 205. A. alaicus Freyn.in Bull. Herb. Boiss., sér.,II, 4 (1904) 766. - A. canifolius Gontsch. in herb. Perennial, acaulescent, 6-10 cm tall, forming small tufts; stipules united with petiole in lower part, scarious, glabrous, ciliate-margined, 170 , 4) Sys Bvt WES 1) WHE, Wf LAR SAN ZA A. ne V Ai. WZ / NALA Wf OA x0 PLATE XVII 1, Astragalus Bobrovii B. Fedtsch.—2. A. nigrocalyx Bge.—3. A. Massagetovii B. Fedtsch. ik rfal 228 5-10 mm long, the lower rounded-ovate to oblong-ovate, the upper lanceo- late; leaves 5-14 (17) cm long, the rachis equaling or 1.5-2 times as long as the petiole, both petiole and rachis villosulous with divergent hairs, slender; leaflets in 7-15 approximate verticils of 6, linear to linear -oblong, subobtuse, (3) 7-11 mm long, 1.5-2.5 (3) mm broad, densely covered on both sides with spreading hairs; flowers many, subradical (peduncles up to 3, rarely up to 20 mm long, bearing 2-3 (5) flowers); pedicels shaggy with spreading hairs, ca. 3-4 (5) mm long; bracts linear to linear-lanceo- late, 3-4 mm long, villous-ciliate; calyx 11-13 (15) mm long, villous with more or less divergent hairs, the linear teeth 2-5 mm long, the tube 3 (rarely 2) times as long as the teeth; corolla yellow, at length becoming reddish; standard 21-26 (28) mm long, the limb obovate to oblong-ovate, retuse, constricted above the lower third, obtusely angular -auriculate be- low, the claw 4-7 mm long; wing-petals 20-24 (26) mm long, the limb ob- long, rounded-obtuse, very rarely subretuse, 2/3 the length of the claw; keel 17-18 (21) mm long, the limb obtusish, 2/5 the length of the claw; ovary on a stipe 1-2 mm long, lanate. June. Stony slopes in the high-mountain zone, occasionally descending into the juniper belt.- Soviet Centr. Asia: Tien Shan (Tashkent area), Pam.-Al. (northern slopes of Alai and Turkestan ranges, Zeravshan, Karategin and Vakhsh mountains). Endemic. Described from ''Ol'gin lug" natural land- mark in Alai mountain range. Type in Geneva (?). 206. A. Nevskii Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946) 131.-Myobroma alatavica Nevski in Acta Inst. Bot. Ac. Sc. URSS, ser. I, IV (1937) 256, non A. alatavicus Kar. et Kir. Perennial, acaulescent, compactly cespitose, 3-15 cm tall; stipules united in lower part with petiole, oblong, obtuse, 6-10mm long, ciliolate- margined; leaves 3-15 cm long, the rachis equaling to 1.5-2 times as long as the petiole, both petiole and rachis sparsely covered with spreading hairs; leaflets in 10-12 verticils of 4-8, lanceolate to lance-oblong, 4-8 mm long, 2-3.5 mm broad, densely canescent on both sides with ascending long soft hairs; peduncles obsolescent to 1 cm long, bearing 2-3 flowers; bracts linear, acuminate, 8-9 mm long, ca. 1 mm broad, equaling to slightly ex- ceeding the pedicel, ciliate-margined; calyx cylindric, 11-14 mm long, densely villous with soft spreading long hairs, thetube 1.5-2 times the length of the linear teeth; corolla yellow at length becoming reddish; stand- ard 20-25 mm long, the limb obovate, entire, scarcely auriculate -angular, half the length of the claw; wing-petals 18-23 mm long, the limb linear, entire, equaling the claw; keel 16-21 mm long, obtuse; ovary densely shag- gy, ona stipe 1.5-2.5 mm long. Fl. June-July. Stony slopes in the high-mountain zone. - Soviet Centr. Asia: Pam.-Al. (Kugitang range). Endemic. Described from the vicinity of Markushi and Khodzhafil'ata. Type in Leningrad. 207. A. dignus Boriss. Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent or nearly so, (5) 8-20 cm tall; stipules united in lower part with petiole, membranous, hairy outside, densely ciliate on the margin, 7-10 mm long, the lower ovate obtuse, the upper oblong- lanceolate acuminate; leaves 6-19 cm long, the rachis equaling to slightly exceeding rarely 1.5 times as long as the petiole, both petiole and rachis villosulous with spreading hairs; leaflets in 7-10 approximate verticils of 6-8 (10), lance-oblong, obtuse, (3) 4-7 mm long, 1.5-2 (2.5) mmbroad, 172 229 lanate -villous on both sides with short ascending hairs; peduncles 2-10 rarely up to 25 mm long, 3-4 (5)-flowered, villosulous with spreading hairs; bracts lance-linear, acuminate, 3-6 mm long, about equaling the pedicel; calyx 12-14mm long, villous with more or less spreading hairs, rarely glabrate, the tube 2-3 (4) times the length of the linear teeth; corolla yel- low at length becoming reddish; standard 20-24 mm long, the limb obovate to oblong-obovate, round-tipped, nearly entire, constricted above the lower third, angular-auriculate at base, the claw 5-7 mm long; wing-petals 19- 22mm long, the limb oblong, entire at apexor nearly so, 2/3 the lengthof the claw; keel 18-20mm long, obtusish; ovary ona stipe ca. 2mm long; pods ona stipe ca. 2mm long, lance-oblong, inflated, 13-17mm long, 4-5 mm broad, carinate ventrally, finely and broadly sulcate dorsally, short-acuminate toa beak 2-3 mm long, coriaceous, densely villous with short ascending hairs, bi- locular to 2/3, 10-12-seeded. Fl. June-July; fr. July-August (September). Gravelly mountain slopes, rarely on pebbles in the high-mountain zone up to 4000m above sea level. - Soviet Centr. Asia: Pam.-Al. (upper Pyandzh basin, Pamir). Endemic. Described from Mats river valley. Type in Leningrad. 208. A. patentivillosus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 129. Perennial, acaulescent, 5-15 (17) cm tall, from an abbreviated buried branched caudex; stipules oblong-triangular to oblong, subobtuse, 10- 13 mm long, shaggy; leaves (5) 7-15 (17) cm long, more or less prostrate, the petiole 2.5 cm long, both petiole and rachis shaggy with spreading hairs; leaflets lanceolate to lance-oblong, subobtuse, (3) 6-11 mm long, in (10) 15-25 verticils of 3-4, densely shaggy with spreading soft hairs; flowers subradical (peduncles not more than 4 mm long, bearing 2-3 flowers); ped- icels shaggy with divergent hairs, 3-4 mm long; bracts linear, 6-8 mm long, shaggy; calyx 14-15 mm long, villous with long divergent hairs, the teeth linear-subulate 6-8 mm long, the tube 1.5 times as long as, rarely equaling the teeth; standard 18-20 mm long, the limb broadly obovate, con- stricted above the lower fourth, gently auriculate-angular below, slightly retuse, the claw 5-6 mm long; wing-petals 16-19 mm long, the limb oblong, slightly dilated upward, round-tipped, equaling the claw; keel 15-18 mm long, obtuse; pods on a stipe ca. 2 mm long, inflated, 9-12 mmlong, 5- 6 mm broad and thick, deeply carinate ventrally, flattened or broadly sul- cate dorsally (with a convex suture), indurated, coriaceous, densely villous with long soft divergent hairs, the long straight beak about 1.5 mm, bilocular to 1/2-2/3, Fl. June-July (August); fr. (June) July-August. Subalpine steppe zone and the upper part of the juniper zone. - Soviet Centr. Asia: Pam.-Al. (upper Zeravshan basin, northern slope of Turkes- tan range). Endemic. Described from region of Kshtut village in Zerav- shan. Type in Leningrad. 209. A. kaschkadarjensis Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. IX (1946) 130. Perennial, acaulescent, (3) 4-7 mm tall [sic}; stipules united below with petiole, 5-7 mm long, the lower ovate, the upper lanceolate, hispid with Spreading hairs; leaves (3) 4-11 cm long, the rachis 1.5-2 times as long as the petiole, both petiole and rachis sparsely covered with spreading hairs, more or less prostrate; leaflets in 10-15 verticils of 4-6, suborbicular, rarely obovate, slightly retuse, (2) 3-5 mm long, sparsely villous above 173 231 with subappressed hairs, villosulous beneath with spreading hairs; flowers subradical (peduncles not more than 4 mm long, bearing 2-3 flowers); ped- icels ca. 3 mm long; bracts linear, 5-6 mm long, white-hyaline, cilitate- margined; calyx 10 mm long, hispid with spreading hairs, the linear teeth 3-3.5mmlong; corolla ochroleucous, at length turning reddish; standard ca. 20 mm long, the limb obovate, constricted above the lower third, angu- lar-auriculate below, scarcely retuse, 2.5 times the length of the claw; wing-petals about equaling the standard, the limb oblong, slightly dilated upward, round-tipped, barely shorter than the claw; keel ca. 16 mm long, obtusish; ovary on a stipe ca. 3 mm long, densely lanate. Fl. June. Stony slopes of the high-mountain zone. - Soviet Centr. Asia: Pam.-Al. (western part of Hissar range from Varzob in the east to Khyrt-sultan mountains, Kashka-Darya river basin and Chul'bair range in the west). En- demic. Described from Chul'bair mountains. Type in Tashkent. Series 27. ADPRESSIPILOSI Gontsch. - Leaves verticillately pinnate; leaflets appressed-pilose on both sides; standard-limb narrowed in lower part, angular-auriculate at base. 210. A. adpressipilosus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 128.-A. alatavicus auct. fl. turk. ex parte. Perennial, acaulescent, 6-14 (20) cm tall, forming small tufts, rarely with short stems beset with short appressed hairs, not more than 3 cm long; stipules united in lower part with petiole, membranous, 7-10 mm long, sparsely ciliate, the lower ovate or oblong obtuse, the upper lanceolate acute; leaves 5-14 (20) cm long, the petiole equaling to slightly exceeding the rachis, both petiole and rachis covered with short appressed hairs; leaflets in 6-9 verticels of 4-6, oblong, rounded-obtuse, rarely lance-ob- long, mucronulate, (3) 6-15 (25) mm long, 2-3.5 (4) mm broad, covered on both sides with appressed hairs (silvery beneath); peduncles 3-10 (20)mm long, bearing 2-3 flowers; bracts lance-linear, long-acuminate to a fine point, strigose-margined, 5-6mm long, slightly exceeding the pedicel; calyx glabrous or covered with isolated or sparse appressed hairs, the lance -linear acuminate sparsely ciliate teeth 3-4 mm long; corolla yellow, becoming reddish; standard (18) 22-24 mm long, the limb obovate, slightly retuse or round-tipped, constricted at the lower fourth, auriculate -angular at base, the claw 5-8 mm long; wing-petals 19-22 mm long, the limb ob- long, round-tipped, 1/2-2/3 the length of the claw; keel 17-19mm long, ob- tusish; pods on a stipe ca. 15 mm long, oblong-oval or oval-oblong, 12- 15 mm long, carinate ventrally, rounded dorsally with a prominent suture, inflated, 5-8 mm broad, 3.5-5 mm thick, densely villosulous with very short appressed hairs, subunilocular (the narrow partial septum barely 1.5mm broad), obtusely beaked, the beak 2-3 mm long. Fl. June-July; fr. July- August. More or less gravelly mountain slopes in the juniper zone. - Soviet Centr. Asia: Pam.-Al., Tien Shan (W.). Endemic. Described from the surround- ings of lake Azorchashme. Type in Leningrad. Section 31. MACROSEMIUM Bge. Astrag. geront. I (1868) 63. - Subacaulescent, covered with straight white hairs; stipules free (not con- nate), united to high up with petiole; flowers subradical, borne on very short few-flowered peduncles; bracteoles none; calyx long-cylindric, slightly 174 32 infundibuliform -dilated upward; wings and keel united to half the length of the claw with the marcescent staminal tube; standard very long, linear - oblong; pods sessile, small, coriaceous, subunilocular, few-seeded. A monotypic section, distributed in Iranian Azerbaijan and Armenia. 211. A. paradoxus Bge. Astrag. geront. I (1868) 63-64; II, 108: Boiss. Fl. or. H, 264; Grossh. Fl. Kavk. II, 315.-Exs.: Pl. or. exs. No. 39. Perennial, acaulescent, 4-7 (15) cm tall, the branched buried caudex and the very short woody stem densely clothed in longitudinally splitting fibrillose remnants of petioles and leaves; stipules oblong-lanceolate, 13- 16 mm long, subhyaline, ciliate-margined; leaves 3-7 (9) cm long, the rachis equaling to 1.5-2 times as long as the petiole, both petiole and rachis glabrous; leaflets 4-6 pairs, obovate, rarely suborbicular, obtuse and minutely mucronulate, 4-8 (10) mm long, glabrous; peduncles very short, bearing 2-3 flowers; bracts ovate, 2-3 times the length of the pedicel, 4- 5 mm long, white-hyaline, ciliate-margined; calyx 13-20 mm long, the glabrous tube 4.5-5 times the length of the linear-lanceolate ciliolate-mar- gined teeth; standard (30) 45-50 mm long, the limb linear-oblong, retuse, gently angular at base, 2-3 times the length of the claw; wing-petals (20) 30-35 mm long, their limbs linear-oblong, entire, 2/3 the length of the claw; keel (18) 27-32 mm long, the limb oblong, gently curved, obtuse, half as long as the claw; pods sessile, spherical-ovoid, 7-10 mm long, inflated, rounded and prominently sutured dorsally and ventrally, glabrous, mucron- ulate. Fl. May; fr. June. Hills and river banks.-Caucasus: S. Transc.: N.W. Iran. Described from Iranian Azerbaijan. Type in Leningrad. Section 32. CAMPANELLA Gontsch. sect. n. in Addenda XI, p- 660.— Herbaceous, subacaulescent perennials (a distinct stem of the current year absent), with pubescence consisting of simple (basifix) white hairs; stipules adnate at base to petiole, not connate; leaves paripinnate, the petioles mar- cescent; racemes loosely many-flowered, borne on a long subradical pe- duncle; bracteolesnone; calyx campanulate, the teeth somewhat longer than the tube; corolla deciduous; pods long-stipitate, cernuous, inequi- laterally lance-oblong, carinate dorsally, canaliculate ventrally, coria- ceous, bilocular. A monotypic central Asian section, most closely related to section Pendulina Gontsch. 212. A. exasperatus N. Basil. Not. Syst. Herb. Hort. Petrop. IV (1923) 45; Fl. Tadzhik. V, 363. Perennial, acaulescent, (16) 20-40 cm tall, from a stout short mostly branched buried caudex; stipules oblong to oblong-lanceolate, acute, scari- ous, 15-20 mm long, 5-8 mm broad, shaggy with long hairs; leaves (18) 25-45 cm long, the petiole and the rachis rigid sulcate covered with long spreading hairs; leaflets 20-35 pairs, broadly ovate to oblong-ovate, slightly retusely truncate, ciliate on the midvein beneath and on themargin, (12) 15-30 (34) mm long, 8-17 (22) mm broad, slender, bright green; pe- duncles (9) 17-30 cm long (about equaling or slightly exceeded by the leaves), stout, covered with spreading hairs and bristles; racemes (4) 6-13 cm long, loosely many-flowered; bracts shorter than, to equaling the calyx, 8-15 (18) mm long; pedicels 1-2.5 mm long, shaggy with spreading rufous hairs; calyx campanulate, 12-16 mm long, the teeth slightly longer than the tube, 175 233 triangular-lanceolate below, acuminate; standard 21-24 mm long, reflexed above, oblong, broadly bilobate at apex, the spreading angular sides abrupt- ly contracted into a broad claw ca. 5 mm long; wing-petals slightly exceed- ed by the standard, the spatulate-oblong limb somewhat longer than the claw; keel obtuse, the lower margin incurved at a right angle; pods lance-oblong, ca. 25 mm long (excluding beak and stipe), densely long-villous, carinate dorsally, grooved ventrally, rigidly coriaceous, the stipe 5-7 mm long, the beak ca. 5mm long. Fl. May; fr. May-June. Mountain slopes in the steppe and juniper zones or (in the south) in the zone of rosaria. - Soviet Centr. Asia: Pam.-Al. (Zeravshan and Kashka Darya basins, southwestern spurs of Hissar range, S. Darvas region). Endemic. Described from Zeravshan basin. Type in Leningrad. Section 33. CHRONOPUS Bge. Astrag. geront. I (1868) 41. - Peren- nials, becoming partially lignified at base, with developed stems or sub- acaulescent, vested with simple pubescence; stipules free (not connate), shortly adnate to petiole; leaves imparipinnate, the petiole marcescent or indurescent, turning into spines; racemes axillary few -flowered abbreviated; bracteolesrarely present; calyx cylindric, not becoming inflated; corolla yellow, glabrous, deciduous; pods oblong to linear-oblong, fleshy or lig- nified, spiny-beaked, bilocular. 213. A. spinescens Bge. Astrag. geront. II (1869) 64. Perennial, 15-30 cm tall, branched, spinescent; stems becoming lig- nified at base, densely leafy, covered with appressed white hairs; stipules free, triangular-lanceolate, acuminate, 5-7 mm long, hyaline, covered with white hairs; leaves 8-16 cm long, the rachis 1.5 times as long as the petiole, both petiole and rachis densely beset with appressed to semi- appressed white hairs, persistent as rigid stout spines; leaflets 20-25 (30) ‘pairs, ovate to ovate-oblong, obtuse or scarcely retuse, glabrous above, shaggy beneath with white hairs; peduncles 1-1.5 cm long, bearing 1-3 flowers, densely covered with appressed white hairs; calyx cylindric, 12- 16 mm long, hirsute with scattered spreading white hairs, the linear teeth 3-5 mm long; corolla yellow; standard 30-35 mm long, the limb oblong- elliptic, obtuse or scarcely retuse, 15-28 mm long, 8-10 mm broad; wing- petals ca. 30 mm long, the limb linear, ca. 18 mm long; keel 28 mm long, the limb obtusish, 11 mm long; pods oblong, straight, 4.5-5cm long, 8- 10 mm broad, compressed laterally, carinate ventrally and dorsally, grad- ually attenuate into a 10-12 mm long beak, indurated, coarsely rugose, rather sparsely hirsute with spreading white hairs, bilocular, many-seeded. Fl. andfr. April-June. (Plate XVI, Figure 1). Dry gravelly soils, often on riverside pebbles in the region of saline wormwood deserts. - Soviet Centr. Asia: Syr-Dar. (Kanibadam, Ashlyk). Gen. distr.: Iran. Described from Iran. Type in Paris. Section 34. ASTENOLOBIUM (Nevski) Gontsch.-Gen. Asteno- lobium Nevski in Acta Inst. Bot. Ac. Sc. URSS ser. I, 4 (1937) 256. - Spinescent shrubs, the branches clothed in approximate recurved hard spines, the leaf petioles and rachises becoming lignified; pubescence con- sisting of simple white and partly black hairs; stipules adnate at base to petiole and partially connate, white-hyaline; leaves imparipinnate, the terminal leaflet early deciduous, the rachis becoming lignified and apically 176 234 237 Spinescent; flowers subsessile, in 2's-3's in leaf axils; bracteoles none; calyx cylindric; corolla yellow; pods subsessile, small, oblong-ovaloid, thinly coriaceous, bilocular.- Related to section Aegacantha, from which it differs in the imparipinnate leaves and the fully bilocular pods. These characters place it as a more primitive section. 214. A. Bobrovii B. Fedtsch. in Addenda XI, p.663.—Astenolobium Bobrovii Nevski in Acta Inst. Bot. Ac. Sc. URSS, ser. I, fasc. 4 (1937) 257, nom. Perennial, 15-27 cm tall, profusely and loosely branched; shoots of the year abbreviated, 2-5 cm long, densely lanate with spreading black and predominantly white hairs; stipules connate to 1/3, glabrate,- (5) 6-7 mm long, triangular, acuminate, the margin beset with white and black cilia; leaves 2-4 cm long, the petiole 0.3-0.7 cm long, both petiole and rachis beset with subappressed short white hairs; leaflets 7-10 pairs, obovate to oblong-obovate, obtuse, 2-5 mm long, broadly cuneate at base, the upper face glabrous or covered with scattered hairs toward the margin, the lower face covered with subappressed short white hairs; bracts white-hyaline, narrowly lanceolate, ca. 4mm long, the margin beset with white and fewer black hairs; calyx ca. 12mmlong, densely villous with spreading soft white hairs interspersed with scattered short black hairs, the tube 1.5-2 times the length of the linear-subulate teeth; standard 17 mm long, the limb ob- ovate, scarcely retuse, somewhat longer than the claw; wing-petals 15mm long, the limb oblong, slightly retuse, somewhat shorter than the claw; keel13mmlong, the limb rhomboidal-oval, acute, 2/3 the length of the claw; ovary short-stipitate, furrowed, villous; pods 9-10 mm long, acute, com- pressed laterally, rounded-applanate ventrally, grooved dorsally, densely villous with long spreading white hairs, commonly 2-seeded, the beak 3- 4mm long. Fl. July; fr. August. (Plate XVIII, Figure 1). Stony mountain slopes and rock crevices in the upper part of the juniper zone and higher up in the zone of mountainous xerophytes. - Soviet Centr. Asia: S.W. Pam.-Al. (Kugitang range). Endemic. Described from the Kugitang range. Type in Leningrad. . Section 35. AEGACANTHA Bge. Astrag. geront. I (1868) 42. - Rather low, profusely branched shrubs, vested with simple hairs; stipules hyaline, adnate at base to petiole, connate or free; leaves paripinnate, the rigid apically spinescent rachises persistent on perennial branches after the shedding of leaflets; flowers 1-3 on very short or more often obsolescent peduncles; bracts hyaline; bracteolesnone; calyx cylindric; flower of medium size, mostly yellow, often finally turning reddish; pods ovaloid or semiovaloid, coriaceous, indurated, subunilocular (a narrow septum pro- jecting as a flange from the ventral suture) or partially bilocular, dehiscent by more or less contractile valves, few-seeded. A section of western Himalayas, Hindu Kush and Central Asia, distributed through the western mountain fringes of middle Asia, from western Tien Shan to the western Himalayas, the maximum number of species occurring in Pamir-Alai, Hindu Kush, and northwestern Himalayas. A section pre- dominantly associated with high mountains. As regards growth habit, it occupies an intermediate position between tragacanth forms on one hand and the pulvinate high-mountain xerophytes on the other. The structure of flower and pod discloses genetic links with the interrelated sections Myo- hroma and Erionotus. LET 1, Standard pubescent (throughout or at the middle).............. 2. + /Standandisilabrouwss yim see te kei) aoe enon ye ne) omen enw een ar ee 4. 2. Standard densely sericeous outside; calyx densely covered with white Hairs vee oes Ge) WS Ree eae, ey RUE fer Pyle ice ee eed oe? eee 3. Standard rather sparsely covered outside about the middle with black hairs (with a varying admixture of longer white hairs); calyx rather sparsely covered with short black hairs interspersed with longer white EES es sae ror te so tel ene eT homer telten amet 217. A. Antoninae G. Grig. QUIS ) ONL hh N G NS SENN SS 1 QAY <= fie . » SA , i pp =e. | | ND SN ’ WAY YL Ale Wn YF y 4 oy ANZ Vi “e {4 — 9 Lhe PLATE XVIII 1. Astragalus sangesuricus Boriss.—2. A. euoplus Trautv., the flower and its parts. 178 238 3. Shoots of the year ca. 2-5 cm long, the internodes 1-3 (15) mm long; peduneles upitorormimy lone offtentobsolescent wren. 2 ayaa Siewiien. eter. RTRs RIUM. STS RU ERR BEATS Ps Mae CREE AS SOU Lads 219. A. lasiosemius Boiss. + Shoots of the year up to 30-35 cm long, the internodes 5-12 cm long; peduncles 8-30 (40) mm long ..... 220. A. roschanicus B. Fedtsch. 4. Standard-limb oblong, angular at base, much longer than the claw; co- Bolla, whitish watwengthturmingiyellowlsheie, Ay aeiich cial) lieve fence s-luele ania ee peed, Sc EE I la OE LS AA, Osseo & 218. A. lejosemius M. Pop. + Standard-limb oval, rather gradually attenuate into the claw...... 3. 5. Calyx covered with white and predominantly black hairs........... Ree eh ne. Me eteO RES Hay Namaig! ta aM ahied ot NARs Moen to.! alle, a BEG a a\c nigrocalyx Slob. ec alys Slabroussor covered with white hairs: iAH aie ante she i ceaee 6. 6. Calyx glabrous; corolla yellow. 215. A. Bornmiullerianus B. Fedtsch. Calyx7covered withywhite hairsecorollarredimsr. wr enews h-iemamemclae! erhemrenieme Es dt Ren oF ot, ato Ae a nee ee 2215 .A. dsharfi’B.'Fedtsch. Series 1. GLABRATI Gontsch. - Standard-limb oval; corolla glabrous; feebly pubescent plants, with a glabrous calyx; stipules connate. 215. A. Bornmiillerianus B. Fedtsch. nom. nov.-A. Bornmulleri B. Fedtsch. in Fl. Tadzhik. V (1937) 674, non Freyn.-Ic.: Fl. Tadzhik. Ver eras Perennial, densely branched, spinescent, shrubby, 10-20 cm tall, sub- pulvinate; spines on the lignified branches 2-5 (6) cm long; shoots of the year glabrous, short, densely leafy, the internodes 1-2 mm (rarely 6 mm) long; stipules ovate, 5-8 mm long, glabrous; leaves 2-5 cm long, the ra- chis glabrous; leaflets 5-7 (8) pairs, suboblong to obovate, 3-7 mm long, glabrous or very sparsely hairy; flowers 1-3 on peduncles 1-3 mm long; pedicels 1-2 mm long; calyx 8-13 mm long, glabrous, the tube 3-4 times the length of the linear-subulate teeth; corolla greenish-yellow; standard 18-21 mm long, the limb equaling or exceeding the claw; wing-petals 17- 19 mm long, the limb suboblong, round-tipped, as long as the claw or near- ly so; keel 15-18 mm long, the limb half the length of the claw, acute, sometimes acuminate; pods semiovaloid, 14-17 mm long, flattened or shal- lowly grooved dorsally, subtriquetrous, attenuate into a beak, coriaceous, partially bilocular, more or less covered with white hairs. Fl. July-August (September); fr. August-September. High-mountain region, chiefly in the alpine (more rarely subalpine) zone at altitudes of 3100-3600 m. Low-grass alpine meadows, foxtail and bent- grass meadows, more rarely steppes. - Soviet Centr. Asia: Pam.- Al. (Zeravshan mountains, Hissar range, Darya region, Darvas and Pamir region). Endemic. Described from Darvaz region. Type in Leningrad. Series 2. PUBESCENTES Gontsch. - Standard-limb oval; corolla gla- brous or standard hairy about the middle; pubescent plants, the calyx hir- sute; stipules connate. 216. A. nigrocalyx Slob. in Fl. Tadzhik. V (1937) 674.-A. lasio- semius var. lejosemius Lipsky in A.H.P. XXVI (1907) 235, exparte. A low shrub, loosely branched, subpulvinate, spinescent; shoots of the year glabrous or covered with scattered hairs, densely leafy; internodes very short (2-6 mm long); stipules ovate, 5-8 mm long, glabrous; leaves (2) 3-5 cm long, the rachis rather sparsely vested with appressed or ILS) 239 spreading hairs; leaflets 5-8 pairs, oblong-elliptic to obovate, (3) 4-10mm long, rather sparsely hairy on both sides or merely beneath; flowers 1-3 on peduncles 0.5-3 mm long; pedicels 1-2 mm long; calyx 9-12 mm long, rather densely hirsute with black and white hairs, the tube 2.5-3.5 times the length of the linear-subulate teeth; corolla yellow; standard 14-17 mm long, the limb slightly retuse, equaling to somewhat exceeding the claw; wing-petals 12-16 mm long, the limb oblong-elliptic, round-tipped, equaling the claw or nearly so; keel 12-14 mm long, the limb acute, half the length of the claw; pods semiovaloid, 10-13 mm long, flattened or shallowly grooved dorsally, carinate ventrally, subtriquetrous, short-beaked, coria- ceous, indurated, hemispherical, more or less white-hirsute. Fl. July- September; fr. (July) August-September. (Plate XVII, Figure 2). The subalpine mountain zone at altitudes of 2750-3500 m. Commonly in subalpine xerophyll meadows, among coarse-grass xerophytic high-moun- tain vegetation (Cous nieta). Occasionally in Prangoseta thickets and in the upper part of the woodland and scrub formation. - Soviet Centr. Asia: Pam.-Al. (Hissar range, Darvaz region, Darvaz, Pamir region). Endemic. Described from western Darvaz. Type in Leningrad. 217. A. Antoninae G. Grig. in Fl. Tadzhik. V (1937) 673. A densely branched spiny low shrub, 7-15 cm tall, subpulvinate; spines on the lignified branched (2) 3-5 cm long; shoots of the year short, densely leafy, rather sparsely puberulent, the internodes 1-3 (4) mm long; stipules ovate to ovate-lanceolate, 4-5 mm long, covered with scattered hairs or merely ciliate-margined; leaves 2-5 cm long, the rachis rather sparsely vested with appressed or spreading hairs; leaflets 4-7 pairs, oblong or oblanceolate, 2-6 mm long, hairy on both sides; flowers 1-3 on short pe- duncles 2-6 mm long; pedicels 2-3 mm long; calyx 10-13 (15) mm long, rather sparsely hirsutulous with black hairs interspersed with longer white hairs, the tube 3-4 times the length of the linear-subulate teeth; corolla yellow; standard 18-22 mm long, the limb about as long to 1.5-2 times as long as the claw, rather sparsely covered outside about the middle with black hairs interspersed with white hairs, glabrous-margined; wing-petals about equaling or slightly shorter than the standard, the limb suboblong, round-tipped, as long as the claw; keel 16-19 mm long, the limb acuminate, as long as the claw or nearly so; ovary black-hirsute. Fl. July-August. Dry steppes in the high-mountain zone, at altitudes of 3888-4150 m— Soviet Centr. Asia: Pam.-Al. (replaces the preceding species inW. Pamir). Endemic. Described from Vakhan. Type in Leningrad. Series 3. DOLICHANTHI Gontsch.-Standard-limb oblong; corolla gla- brous. 218. A. lejosemius (Lipsky) M. Pop. in Schedis ad Herb. Fl. As. Med. fasc. XVI (1928) p. 50; in Fl. Tadzhik. V, 379.-A. lasiosemius var. leiosemius Lipsky in A.H.P. XXVI (1907) 235, ex parte. - Exs. : Herb. Fl. As. Med. No. 388. A densely-branched unarmed low shrub, 4-10 cm tall, compactly sub- pulvinate, the spines on the lignified branches 3-6 cm long; shoots of the year 3-6 cm long; shoots of the year 0.5-1 (rarely 3)cmlong, densely leafy, more or less hairy or glabrate; stipulestriangular-lanceolate, 4-6 mm long, not connate, adnate at base to petiole, more or less hairy or merely ciliate -margined; leaves (2) 3-6 cm long, the rachis rather sparsely covered 180 240 with spreading hairs; leaflets 6-10 pairs, oblong-elliptic to suboblong, ob- tusish and mucronulate, hairy on both sides but more densely so beneath; flowers 1-2 on peduncles 1-2 mm long; pedicels 1-2.5 mm long; calyx 8- 11 mm long, white-hirsute, the tube 1.5-2 times the length of the lance- subulate teeth; corolla white; standard 18-22 mm long, retuse, the claw 3-4 mm long; wing-petals 16-20 mm long, the limb linear, obtuse, twice as long as the claw; keel 11-14 mm long, cleft at apex, the limb oval-oblong, obtuse, 1.5 times the length of the claw; pods ovaloid, 10-14 mm long, com- pressed laterally, carinate ventrally and dorsally, coriaceous, indurated, white -hirsute, prominently sutured, unilocular, the septum consisting of a narrow 1-1.5 mm broad flange, the beak 2-3 mm long. Fl. June-July; fr. June-August. The subalpine mountain zone, on more or less stony slopes. Aconstituent of mountainous xerophytic, pulvinate and coarse-grass vegetation, more rarely of gravelly high-mountain steppes. Rarely penetrating into the upper fringes of the juniper belt. - Soviet Centr. Asia: Pam.-Al. (W. from Kugi- tang through Khodzha-gurgur-ata range, upper Saigardak and Kashka- Darya rivers to central part of Zeravshan). Endemic. Described from the Kugitang range. Type in Tashkent. Series 4. SERICOPETALI Gontsch. - Standard-limb oval, exterior dense - ly covered with white silky hairs. Pubescent plant with hirsute calyx; stip- ules not connate. 219. A. lasiosemius Boiss. Diagn. ser. I, IX (1849) 96; Bge. Astrag. geront. II, 71; Boiss. Fl. or II 304; Fl. Tadzhik. V, 387. A densely branched spiny low shrub, 10-25 cm tall, loosely or rather compactly pulvinate; spines on the lignified branches (2) 3-5 cm long; shoots of the year 2-5 cm long, densely leafy, rather densely hairy, the internodes 1-3 (5) mm long; stipules lanceolate, 5-8 mm long, long-acuminate, more or less hairy; leaves (2) 3-5 (6) cm long, the rachis more or less hairy; leaflets 6-11 pairs, obovate to oblong-lanceolate, subobtuse to subacute, commonly mucronate, (2) 3-8 (9) mm long, rather densely covered with hairs on both sides; flowers 2-3 on very short peduncles (ca. 5 mm long); pedicels 2-3 mm long; calyx 12-16 mm long, densely white-hirsute, the tube 2-3 times the length of the linear-subulate teeth; corolla yellow; stand- ard 19-25 mm long, the limb equaling to 1.5 times as long as the claw; wing-petals 18-22 mm long, the limb oblong, round-tipped, glabrous or par- tially pubescent, about equaling theclaw; keel 15-20 mm long, the claw 1.5- 2 times the length of the limb, the limb glabrous or covered with hairs on the lower margin; pods semiovaloid, 14-17 mm long, flattened or shallowly grooved dorsally, carinate ventrally, subtriquetrous, beaked, sessile, coriaceous, indurated, partially bilocular, rather densely hirsute with white hairs. Fl. June-July (August); fr. July-September. Intermediate and high-mountain zones, at altitudes of 2400-3400m, in the deciduous woodland and scrub, juniper (in the higher parts) and subalpine belts. Commonly in dry steppes, sparse Ferula and Prangos as- sociations; also thinned-out rosaria and juniper groves, among xerophytic mountain vegetation, on more or less stony or gravelly slopes. - Soviet Centr. Asia: Tien Shan (W. and S.), Pam.-Al. Gen. distr.: Afghanistan. Described from Afghanistan. Type in Geneva. 220. A. roschanicus B. Fedtsch. in Fl. Tadzhik. V (1937) 673. Frutescent, very closely related to the preceding species; differing chiefly in the elongated shoots of the year. 181 241 242 Stony slopes of the intermediate mountain zone. Replacing A. lasio- semius Boiss. in enclosed desert-invaded valleys. - Soviet Centr. Asia: Pam.-Al. (Zeravshan, Karategin, Vakhsh, Vanch, W. Pamir). Endemic. Described from Darvaz. Type in Leningrad. Series 5. RUBRIFLORI Gontsch. - Standard-limb oval; corolla glabrous, red; pubescent plants with hirsute calyx; stipules not connate. 221. A. dsharfi B. Fedtsch. in Fl. Tadzhik. V (1937) 675. A densely branched spiny shrub ca. 15-25 cm tall, subpulvinate; spines on the lignified branches 5-8 cm long; shoots of the year 2-7 (10) cm long, densely leafy, more or less hairy or glabrate, the internodes 2-6 (10) mm long; stipules lance-subulate, long-acuminate, 7-12 mm long, hairy or glabrate, 1.5-8 cm long, the rachis densely or sparsely vested with ap- pressed or spreading hairs; leaflets 5-8 pairs, oblong elliptic or oblanceo- late, acute, 5-10 mm long, glabrous above or nearly so, rather densely hairy beneath; flowers 1-2 on peduncles 5-15 mm long; pedicels 2-4 mm long; calyx 17-22 mm long, rather sparsely covered with white hairs, the tube 2-3 times the length of the linear-subulate teeth; standard 21-28 mm long, the limb oblong-oval, 1.5 times the length of the claw; wing-petals equaling standard or nearly so, the limb suboblong, slightly curved, round-tipped, half as long as the claw; keel 21-26 mm long, the claw 2- 2.5 times the length of the obtusish limb; pods inequilaterally semiovaloid, flattened or shallowly grooved dorsally, carinate ventrally, subtriquetrous, 13-16 mm long, partially bilocular, white-hirsutulous, the beak 3-5 mm long. Fl. and Fr. June. Stony dry slopes in the intermediate mountain zone. - Soviet Centr. Asia: Pam.-Al. Endemic. Described from the vicinity of Dzharf (Darvaz). Type in Leningrad. Section 36. ACANTHOPHACE Bge. Astrag. geront. I (1868) 45.- flowers sessile on a stout, hairy peduncle 2-8 mm long; calyx cylindric or campanulate, scarious, pubescent with short or long hairs; corolla violet or yellow, the petals free; pods sessile or short-stipitate, rigidly coriaceous, inflated, unilocular, many-seeded; branched low shrubs, often pulvinate; leaves paripinnate, the petiole spinescent; stipules imbricated, adnate to petiole. -Iranian and Caucasian section. 1. Plants 2-3 cm tall; branches crowded, very densely covered with erect overlapping thick spines; calyx hirsutulous, the tube 4-5 times the length of the teeth; style 9 mm long (Talysh). 222. A. sahendi Buhse + Branches more distant; less densely covered with divergent slender spines (2-4 cm long); calyx hirsute with long or short hairs, the tube about equaling or 1.5-2.5 times as long as the teeth; style 10-18 mm OMG ear. warble he ce! sow Een omy cto gfe Apa teuee is kete wt kon aii ee 2. 2. Calyx-tube 2-2.5 times the length of the teeth; calyx covered with rather lone white hains. (cay el —1s5 hmm lone)! 5 2. ese. een we + Calyx-tube about equaling to 1.5 times the length of the teeth; calyx covered with short white hairs (less than 0.5 mm long).......... 4. 3. Leaflets 4-6 pairs, 2 mm broad, hairy on both sides; spines 2-3 cm long; bracts ca. 17 mm long; calyx 14 mm long; style 18 mm long (MounteArarat)> xray ga ata, 2h) cote tit 223. A. coarctatus Trautv. 182 243 + Leaflets 6-8 pairs, 1-1.5mm broad, glabrous above; spines 3-4 cm long; bracts 7-8 cm long; calyx 11-13 mm long; style 13-16 mm long (DIS SS Earl) oi sragt soetaivan clon aby ays: Farpce thm s, opis) ike. c 224. A. Beckerianus Trautv. 4. Flowers 23-30 mm long, in 2's-3's in leaf axils, on peduncles 2-6 mm long; calyx 14-16 mm long; style 14-16 mm long; leaflets 6-8 pairs, 8 mm long, 2 mm broad, glabrous above, pubescent beneath with scat- tered appressed hairs; bracts 13 mm long. . 225. A. euoplus Trautv. + Flowers 18-21 mm long, in 3's-5's in leaf axils, onpedunclesca. 8mm long; calyx 10-12 mm long; style 10-12 mm long; leaflets 6-9 pairs, 4-6 mm long, 2mm broad, glabrous on both sides; bracts 7 mm long. 2 CORN G SEMEN EES eS a ANIA SRO ie SrA 226. A. sangesuricus Boriss. 222. A. sahendi Buhse in Fisch. Syn. Astrag. Trag. (1853) 102; Bge. Astrag. geront. II (1869) 72; Boiss. Fl. or. II, 311; Grossh. Fl. Kavk. Hews Oo lee risch isc, tabe. £.)10%>—Exs, SH; PyR. Nomi714: A densely branched shrub, pulvinate; branches short, leaf rachises and petioles densely clothed in erect to somewhat divergent spines; spines firm, straight 2-3 cm long, beset with spreading hairs when young, at length glabrescent; stipules membranous, ca. 10 mm long, the lower third con- nate to petiole, the upper portion free acuminate glabrous ciliate-margined; leaflets 5-7 pairs, oblong to oblong-ovate, 3-8 mm long, 1-2 mm broad, long-acuminate and spine-tipped, conduplicate, covered on both sides with spreading hairs, the upper ones much shorter than the free upper part of the spine; flowers axillary in 2's, subsessile on a short peduncle ca.5mm long; bracts membranous, ca. 5mm long, oval, long-acuminate, exceeding the short pedicel; calyx ca. 11-12 mmlong, cylindric, pubescent withblack hairs, the tube 4-5 times the length of the subulate teeth; standard ca. 22 mm long, oblong-obovate, gradually attenuate toward base, 7-8 mm broad, obsoletely retuse; wing-petals ca. 18 mm long, the oblong limb 8mm long, 2 mm broad, the claw filiform; keel 14 mm long, the limb ovate, rounded on one side, 5 mm long, 4 mm broad, with a bluish-violet speck at apex; ovary 4mm long, covered with short white hairs; style 9 mm long, curved at the end, the lower 1/3 hairy; pods unilocular, oblong, hirsute, firm, 1cm long, 5 mm broad. Fl. May; fr. June. Mountains, on rocks and stony slopes, at altitudes of 1800-2400 m (ac- cording to Grossheim).- Caucasus: Tal. (Dyman, Mount Kyz-Yurdy, ac- cording to Grossheim). Gen. distr.: N.W. Iran (Savelan and Sakhend mountains). Described from Sakhend mountains. Type in Leningrad. 223. A. coarctatus Trautv. in A: H.P. IV (1876) 365; Boiss. Fl. or. Suppl. 183. A low shrub, up to 7 cm tall, densely branched, forming small short- stemmed cushions; branches short, crowded, clothed in imbricated stip- ules and spiny leaf rachises; spines 2-3 cm long, slender, pubescent when young at length glabrous, straight, erect; stipules ca. 13 mm long, linear long-acuminate from an oblong base, glabrate with sparse hairs on the back, Scarious, connate to 7 mm, ciliate; leaflets 4-6 pairs, oblong, acuminate, without apical point, hairy onboth sides, greenor glaucescent by pubescence; up to 7 mm long, 2 mm broad, conduplicate, thickish, about equaling the terminal spine; flowers large, axillary, commonly paired, sessile, per- sistent on the fruit; bracts yellow, membranous, linear, filiform long- acuminate, conduplicate, up to 17 mm long, sparsely hairy on the back and at apex; calyx cylindric, 14 mm long, hirsute with scattered long white 183 244 .maturity. Fl. June-July; fr. July-August. hairs, prominently nerved, scarious, attenuate toward base but not conic, slightly inflated in fruit the teeth unequal, lance-subulate, straight, ca. 4 mm long, 2/5 the length of the tube; standard oblong-lanceolate, 28-29mm long, obtuse or retuse, straight, twice the length of the calyx’, glabrous, ca. 15 mm long, 6 mm broad, narrowed to 5 mm toward the claw, the claw 6 mm broad in upper part, cuneately attenuate to 2 mm at base, with adnate stamen; wing-petals 25 mm long, the lanceolate limb half the length of the claw, auriculate at base; keel ca. 20 mm long, the ovate limb 2/5 the length of the claw; stamens fused at base with wings and keel; ovary oblong, pu- bescent; style ca. 18 mm long, pubescent merely at base, naked upward; pods sessile, oblong-ellipsoid, acute, densely appressed-hirsute, unilocu- lar, many (4)-seeded. August. Mountains. - Caucasus: South Transc. (Mount Ararat, Lake Kyup-gel!'). Endemic. Described from Mt. Ararat, Lake Kyup-gel'. Type in Leningrad. 224. A. Beckerianus Trautv. in A.H.P. IV (1876) 363; Grossh. FI. Kavk. II, 305. A shrub up to 17 cm tall, densely branched, pulvinate, the short branches clothed in imbricated stipules and spiny leaf rachises; spines firm, 3-4cm long, sparsely appressed-pubescent at length glabrescent, straight, diver- gent; stipules 10-14 mm long, membranous, adnate to petiole up to the middle, ciliate-margined, glabrous, oblong-ovate at base, the free part linear-lanceolate; leaflets 6-8 pairs, 5-8mm long, 1-1.5 mm broad, ob- long-lanceolate, subobtuse, minutely mucronulate, green, glabrous above, sparsely appressed-pubescent beneath; flowers axillary in 2's-3's, borne on a stout hairy peduncle 2-3 mm long, crowded in capitate "inflorescences" 3-4 cm in diameter; bracts membranous, scaphoid, linear to linear-lanceo- late, covered with white hairs on the back or glabrous, ciliate-margined, equaling the calyx-tube; bracts filiform-linear, 5-6 mm long; calyx scari- | ous, cylindric, densely villous with white divergent rather long (1-1.5mm) | hairs, slightly inflated in fruit, 11-13 mm long, the subulate-lanceolate teeth (3) 4-5 mm long; corolla persistent, pink; standard 23-30 mm long, 7-9 mm broad in upper part, oblong, obtuse, straight, twice as long as | the calyx, glabrous, the limb prominently or obsoletely auriculate at base, attenuate to a broad cuneate claw; wing-petals somewhat exceeded by the standard, 20-27 mm long, the oblong limb 8-11 mm long, 2-2.5 mm broad, the filiform claw 12-16 mm long; keel shorter than the wings, 18-23 mm long, the ovate obtusish limb 6-7 mm long, ca. 3 mm broad; ovary white- hairy, suboblong-lanceolate; style 13-16 mm long, filiform, naked, curved; pods sessile, acute, oblong, subacutely attenuate toward base, covered with white silky hairs, unilocular, many-seeded, equaling the calyx in | Stony and gravelly slopes, at altitudes of 2500-3500 m.- Caucasus: Dag. (surroundings of Kurush and Ishir), East. Transc. (region of Kubinskii). Endemic. Described from 5. Dagestan, surroundings of Kurush. Type in Leningrad. 225. A euoplus Trautv. in A.H.P. IV (1876) 364. A shrub up to 25 cm tall, densely branched, pulvinate, the decumbent elongated branches clothed in imbricated stipules and spinescent leaf ra- chises; spines slender, weak, 3-5 cm long, divergent, straight, sparsely appressed-pubescent, at length glabrous; stipules 12 mm long, ca. 4-3mm is broad, membranous, adnate to petiole up to the middle, the free part “a 184 245 linear-lanceolate, acuminate, glabrous, ciliate-margined; leaves termi- nating in a spine the length of leaflet; leaflets 6-8 pairs, small, up to 8mm long, 2 mm broad, oblong-ovate, commonly flat, green, glabrous above, pubescent beneath with scattered appressed hairs, subacute, minutely mucronulate; flowers sessile, in 2's-3's in the leaf axils on a very short pubescent peduncle (2-6 mm long), crowded in a capitate "inflorescence" ca. 3 mm in diameter; bracts membranous, linear, ca. 13 mm long, con- cavely curved, scaphoid, white-hairy on the midrib, ciliate at apex, long- acuminate, exceeded by the calyx-tube; calyx 14-16 mm long, cylindric, densely puberulent with white divergent hairs ca. 0.5 mm long, scarious, dilated in fruit, subacutely attenuate toward base, the lance -subulate herba- ceous appressed-puberulent teeth exceeding half the length of the tube, 5- 7 mm long; corolla violet (in drying), persistent; standard oblong, obtuse, retuse, straight, 23-30 mm long, 7-8 mm broad, twice as long as the calyx, glabrous, attenuate at base to a broad claw; wing-petals exceeded by the standard, 22-26 mm long, the lanceolate limb 9-11 mm long, auriculate at base, the claw filiform; keel shorter than the wings, 18-21 mm long, the ovate limb 6-7 mm long; ovary oblong, appressed-hairy; style 14-16mm long, naked, filiform; pods sessile, oblong, acute, densely covered with white silky hairs, unilocular, many-seeded. Fl. June-August. (Plate XVII, Figure 2). Dry stony slopes, alt. ca. 2,800 m.-Caucasus: E. and S. Transc. Endemic. Described from Karabakh (Okhchi). Type in Leningrad. 226. A. sangesuricus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). A rather low shrub, 10-15 cm tall, densely branched from the base, pulvinate; branches short, clothed in imbricated stipules and spinescent leaf rachises; spines strong, 2-4 (6) cm long, straight, slightly divergent; stipules ca. 12 mm long, ca. 4 mm broad at base, attenuate toward apex, linear-lanceolate, acuminate, glabrous, ciliate-margined at base; leaves terminating in a short spine shorter than to equaling the leaflets; leaflets 6-9 pairs, 4-6 mm long, ca. 2 mm broad, oblong, green, glabrous, mu- cronulate; flowers axillary in 3's-5's on a pannose peduncle ca. 8mm long, crowded into a capitate inflorescence ca. 2.5cm in diameter; bracts mem- branous, filiform-linear, 7 mm long, scaphoid, acuminate, sparsely ciliate at base; calyx cylindric, pubescent with short appressed white hairs, 10- 12 mm long, dilated in fruit, the filiform teeth 4-5 mm long; corolla violet; standard 18-21.5 mm long, 7-8 mm broad, ovate-oblong, gradually cuneate - ly attenuate toward base, scarcely or markedly retuse; wing-petals 15- 19 mm long. the limb lanceolate, 8-9 mm long, auriculate at base, the claw filiform; keel 14-16 mm long, the limb oblong-ovate, truncate at a right angle, ca. 5mm long; ovary oblong, appressed-hairy; style 10-12 mm long, filiform, naked; pods oblong-ovoid, acute, rupturing the calyx, appressed- hirsutulous, unilocular, many-seeded (3). Fl. June; fr. July-August. (Plate XVIII, Figure 1). Mountains, alt. ca. 1800 m.-Caucasus: S. Transc. (Zangezur range, Ordubad village). Endemic. Described from the Caucasus, from the Zan- gezur range. Type in Leningrad. Note. A species akinto A. euoplus Trautv., from which it differs in a number of characters: the size of flower (18-21.5 mm as against 23- 30 mm long) and flower parts, the number of flowers in the leaf axil (3-5 185 246 as against 2-3), the length of peduncle (8 as against 2-6 mm), the shape, size (4-6 mm long and 2 mm broad as against 8 and 2 mm respectively) and pubescence of leaflets, as well as the size of stipules (7 mm as against 13 mm long). Subgenus III. HY POGLOTTIS Bge. Astrag. geront I (1868) 46. - Herbaceous perennials, subacaulescent or with more or less developed stems of the year, vested with simple (basifix) hairs; inflorescences compactly capitate or spiciform, borne on a long axillary peduncle; calyx cylindric or campanulate, sometimes attenuate toward base, not changing in shape, i.e. not becoming inflated, commonly ruptured by the pod; corolla violet or purple, rarely yellow, marcescent, not persistent in fruit; pods sessile or nearly so, oblong or semiovaloid, rarely oblong-ovaloid, coriaceous, rarely submembranous and bladdery-inflated, bilocular or nearly so. Section 37. EU-HYPOGLOTTIS Bge. Astrag. geront. I (1868) 47,50. - Perennials, with well developed stems, vested with white and black basifix hairs; stipules more or less connate; inflorescences compact, capitate or spiciform, borne on axillary peduncles; bracteoles none; calyx cylindric or cylindric-campanulate; corolla violet, purple or yellow; wings exceeding the keel; pods sessile or nearly so, erect, oblong to semiovaloid, more or less carinate ventrally, canaliculate dorsally, often bladdery-inflated, sub- spherical, coriaceous or membranous, bilocular or nearly so. The distribution area of the section Eu-Hypoglottis covers the temperate region of Eurasia and partially of North America, penetrating southward (in Asia Minor, Armenia and Syria) into the subtropical region. Species of this section also occur widely in the Caucasus. Key to Species in Flowering Condition 1...Cornolla ochroleucousiyy. says wer sees aytatehets Gil yelleticn ests) ope uomcnte Ne eee 2. Corolla azure, purple or violet, rarely ochroleucous and then the keel AZUEC OF, VOLS toh paigai oc foul oblolmsiacuhalhicl tows tawoayen of opie komt Releente tana 3. 2. Leaflets (8) 10-15 pairs, lanceolate to lance-oblong, covered on both sides with scattered short appressed hairs, rarely glabrate above... SS RE MED Ty nS eee ne nee mn Neen J, + Leaflets (5) 6-8 (9) pairs, elliptic to oblong-elliptic, glabrous above. Be NOTION eee toe ER CMe Ee OMe Or er oe bo 228. A. Freynii Alb. 3. Dwarf high-mountain plants, 3-10 cm tall, the short stems 1-4 (8) cm long, more rarely obsolescent; corolla yellowish with azure or violet keel, or azure; inflorescences few (2-8)-flowered or flowers subradi- SEM TE" olla 6 ee s aviietivw, sO, «« Pitas sicadndirauis, Asptele are Rly Uae i ako fae a oS 4. + Plants taller, with more developed stems, not growing in high moun- CELTS ie csp puss copes Sey 0; Voy setae gue a0 | on vemee been cue its gion’ owe Gaye iRcpide) on lap co ate 5. 4. Inflorescences subumbellate, 5-8-flowered, borne on peduncle 2-5 (9) cm long; stipules herbaceous, green... 229. A. oreades C.A.M. 247 + Flowers subradical, in 2's-3's on very short (1-2 mm long) obsolescent peduncles; stipules whitish, subhyaline; pods pubescent with scattered Pa Sas thy to ha eek Sacaksascn bY cei oe cue pice cnes: noe code demcets 236. A. supinus Bge. 186 ' : Parblowersuniaise ss pods) clabnousmmoblong we ibe V.4h kee NS Hiocee WRT OR EE Wi Mee, 8 ad ye wed dld 237. A. Klopotovskii Sosn. 5. Calyx covered with exclusively white hairs, the teeth setiform not ex- ceeding half the length of the tube. . . 235. A. saganlugensis Trautv. + Pubescence of calyx always including some black hairs......... 6. 6. Calyx-tube equaling to barely exceeding the teeth; calyx villous with long white hairs and less profusely black-hirsutulous............ PT EIR te Seater sew: its wine L, gmk Aenean 9 ead 239. A. dasyglottis Fisch. calyx -tuberc oO) timesuas)longsassthesteeth ia eal) See hee. Sebo es 7. Leaflets glabrous above and faintly pubescent merely on the-margin. 6 oho on doer OIG O: Cla OMBaet CRASPELSEEDTT bet hE RAREL CL Saas Bhs LARS op ee mares Cuero ar ne 8. 2eeecanlets pubescent ion both) sides Lycra. eed tnaes woned.) oPatie em 10. 8. Calyx-tube 5-6 times the length ofthe teeth; corolla whitish with vio- let keel; inflorescences loosely capitate, subumbellate.......... 5. Oe Ef RIOD IG) CeiC) CeO Om CRERIen Seine Ce ewer a 233. A. chordorhisus Fisch. + Calyx-tube 2-3.5 times the length of the teeth; corolla violet or azure - waoletaeintlonescences icompactlyicapitatertw.ype ) awlacs tie needs oF 9. Standard-limb oblong-ovate, attenuate in upper 1/3.............. >. oF 6 Bo bueto o aulous GONE CRCMAE G5 NenemEE Onteee 234. A. bachmarensis Grossh. + Standard-limb oblong-obovate, attenuate in lower part........... RUE PoRE PIO AR PUES SAA Reh) FG thm nba ated te 240. A. tibetanus Benth. lOeeotandard-limb ovateior rhomboid-ovate: . 9.) .4.0... 25 yee. i Wie eeotancdard—limbropovater On Oval: las 5 WAM Teh, Or See 12 11. Leaflets appressed-hispid; wing-petals entire at apex.......... TN ert etal Ne a ce LE ae Re 231. A. viciifolius DC. * + Leaflets more or less puberulent with spreading hairs.......... erent ese Pee Beas ee aS See 230. A. flaccidus M.B. 12. Standard-limb oval; bracts oblong-ovate ....238. A. danicus Retz. + Standard-limb oblong-obovate, attenuate below; bracts lanceolate, nVaNaTIe ee Maun fh. tod able teh steers, fe geht tal, Whe Se SPB es. 240. A. tibetanus Benth. Key to Species in Fruiting Condition 1. Pods spherical or ovoid, bladdery-inflated, membranous, grooved WE HOBIUIK? Ea ClOreSaiys Ssh ols ch BkGNENG oie Sale Seo ia! oN 6a SeG Ge6 oe 2. + Pods not bladdery, more or less carinate or subapplanate but not MOVE Cuvier ably rages si Jol ime keke, Sits kas etka Nar alirebeUatver iy», le ted Gina o! « Uorkowi: 4. 2. Rather large plants, 20-80 cm tall, the stems 15-60 cm long; stip- ISS COMME Mo TWAS] Bi lOAESis Gow She oters S Slane alera Giallo GNoT Hid ooo a as 248 + Squat plants, 4-10 cm tall, the strongly abbreviated branches 1-3 (8) cm long; stipules connate high up... 229. A. oreades C. A. Mey. 3. Leaflets (5) 6-8 (9) pairs, elliptic to oblong-elliptic; pods densely villous with blackish-brown hairs (with a slight admixture of white hairs) . i ee, Selo oee Ae Ere yrinic eA + menilers (8) 10- 15 sais, levasecletia to lance-oblong, rarely oblong- oval; pods densely villous often with black commonly long hairs. 0.8) a0 WSR a goad nb, ‘te et Ate Ra, couche AIRS RSI oR NS zat. A. cicer I. 4. Flower and pods in 2's-3's on a peduncle 1-2 mm long; pods sessile, semiovaloid-oblong, 7-13 mm long, thinly membranous-coriaceous, puberulent with scattered hairs........... 236. A. supinus Bge. * See also 232. A. kubensis Grossh., p. 653. 187 + . Pods in 4's-5's on a very:short, peduncle, wslabrate \.ge ha sabe TOMER tera A ee Re: Eee Mere Ree Neuen Art 237. A. Klopotovskii Sosn. ++ Pods capitately crowded, several at the end of a long peduncle. ... 5. 5. Pods strongly inflated, obcompressed, ovoid (oblong in side-view), coriaceous, canaliculate dorsally, rounded-carinate or subapplanate ventrally i 2ase0e wekien tone Pace ane eget ce tae enh See es 6. Pods compressed laterally, semiovaloid to linear-oblong....... We Calyx-tube twice (rarely 1.5 times) as long as the teeth; calyx hirsute with black and fewer white appressed or subappressed hairs...... ERD SR Ae ERR Ly TSR MRLs Rs A fa hs 238. A. danicus Retz. + Calyx-tube equaling tobarely longer than the teeth; calyx villous with long white hairs and less profusely hirsute with black spreading hairs. Shite Eek Wad Silk WAM Mb OEE CORE RND.S SNL TG Oe th 239. A. dasyglottis Fisch. Tt POdS. SESS] VHA. LOLI Be, 6 Bal ALM DUNC MSI Old Oi REA Re a 8. Pods on a stipe (1) 2-4 mm long, oblong to linear-oblong, more or less compressed laterally, gently curved. . 240. A. tibetanus Benth. 8. Pubescence of calyx consisting exclusively of white hairs; calyx-tube twice as long as the subsetiform teeth; pods ca. 7mm long........ 2 MPS a UNL A MRIS TEED ater. SUMS ta ay aa 235. A. saganlugensis Trautv. + Pubescence of calyx consisting of white and black hairs; calyx-tube 3-6 times as long as the subulate-linear teeth, rarely twice as long and a + then, pods!-ca’. "Hamm long. Stawi-f sdsytae s eo ere Seer eee 9. 9. Pods linear to linear-oblong, falcate, 4-4.5 times as long as broad. Bole ae hai iia abl se PG. BULA oe pane i 230. A. flaccidus M.B. + Pods semiovaloid, semiovaloid-oblong or ovaloid-oblong, 2-3 times as long ais broad} Uy aie Miva ear el aie r slelaly sels he aS Diy, Se a 10. 249 10. Pods covered exclusively with white hairs. 231. A. viciifolius DC. * + Pods covered with black hairs, occasionally interspersed with some . White whaiss! demir « Gyieleeie 284. A. oxyglottis Stev. Ridges of the pod-valves noe with teeth or prickles; pods disposed int OOS eA CERMOT yy cj i sitsa SPihte) sicies ‘eeu tests) Hor. A.M. Now 17a, b- Annual, 8-40 cm tall; stems erect, branched or nearly simple, 4-35cm long, sulcate-angled, sparsely vested with appressed short hairs; stipules free, adnate at base to petiole, triangular-lanceolate, 3-4 mm long, beset with mixed white and black or pure black cilia; leaves 3-5 cm long, the petiole and the much longer rachis covered with white or rarely mixed black and white appressed hairs; leaflets 5-7 pairs, oblong, oblong-elliptic or sublinear, 10-14 (20) mm long, subobtuse or slightly retuse, sparsely cov- ered on both sides (sometimes densely beneath) with appressed hairs, some- times glabrous above; peduncles somewhat shorter than to rarely equaling or somewhat exceeding the leaves, sulcate, covered with ascending and beneath infl. sometimes mixed black and white hairs; inflorescence com- pact, capitate, ca. 1 cm long, 5-10-flowered, the flowers subsessile; bracts narrowly linear, beset with white or mixed white and black cilia, 231 307 308 2-3.5 mmlong; calyx 6-7 mm long, black- or white-hirsute, the finely subu- late teeth somewhat longer than the tube; corolla violet; standard 6-8 mm long, the limb oblong-elliptic, slightly retuse, rather gradually attenuate into and twice as long as the short claw; wing-petals 5.5-6 mm long, the limb oblong, rounded-obtuse, equaling the claw; keel 4.5-5 mm long, the limb broad, obtuse, equaling to 1.5 times as long as the claw; pods some- what obliquely upright, tightly appressed to each other, linear-oblong, near - ly straight, 12-20 mmlong, submembranous, carinate ventrally, grooved dorsally, prolonged into an acute reflexed beak 1-1.5 mm long, hirsute with subappressed white hairs. Fl. and fr. May-June (August). Loess foothills in the meadowgrass-sedge and wormwood-sedge vegeta - tion containing xerophytic tall grasses. In the south (Hissar range) ascend- ing into the mountains along river valleys, penetrating into a part of the woodland and scrub formation. In the north, on river valley sands and stony mountain slopes. - Soviet Centr. Asia: Balkh. region (rarely in the southern part), Tien Shan, Pam.-Al., Mtn. Turkm. (Balkhyz). Gen. distr. : Afghan- istan. Described from Afghanistan (Kabul distr.). Type in Geneva. 282. A. filicaulis Fisch. et Mey. in Bull. Soc. Nat. Mosc. XII (1839) 152 (nomen); Ldb. Fl. Ross. I, 637.-A. leptodermus Bge. Astrag. turk. (1880) 208.-A. agrestis Freyn in Bull. Herb. Boiss. 2 sér IV (1904) 446.-Ic.: Fl. Tadzhik. V, t. 24, 8. Annual; stems erect, 5-40 cm long, branched, sparsely vested with ap- pressed hairs; stipules free, partly adnate at base to petiole, lance-subu- late, 2-3 (4) mm long, beset with white or rarely mixed white and black cilia; leaves (3) 4-7 (8) cm long, the petiole and rachis sparsely covered with short ascending hairs; leaflets 6-7 pairs, oblong, oval-oblong or line- ar, 9-10 mm long, subobtuse, sometimes slightly retuse, sparsely covered on both sides with appressed hairs; peduncles somewhat shorter to some- what longer than the leaves, sparsely pubescent with ascending white hairs; flowers in heads of 6-20; bracts lanceolate to narrowly lanceolate, acumi- nate, 1-1.5 mm long; calyx 3-4 mm long, hirsute with appressed white and black hairs, the linear-subulate teeth equaling the tube; corolla pale violet; standard 6-8 mm long, the limb oblong, 5.5-7.5 mm long, retuse, attenuate and imperceptibly passing into the claw; wing-petals 5-6 mm long, the limb oblong, slightly retuse, 1.5 times the length of the claw; keel 4-5 mm long, the limb broad, obtuse, equaling the claw; pods stellately spreading, ob- long, slightly recurved, 7-10 mm long, 2-3 mm broad, carinate ventrally, grooved dorsally, thinly coriaceous, densely appressed- or subappressed- pubescent and hirsute with longer spreading hairs, finely rugose, few -seeded- Fl. (March) April-May (June); fr. May-June (July). Desert plains in the north of Soviet Centr. Asiaand submontane loess plains and foothills of Soviet Central Asian mountains, onclay and loam fine - grained or stony fine-grained soils, rarely on sands. Ascending in the south into the mountains up to the witch-grass steppe zone. - Soviet Centr. Asia: Ar.-Casp. (W. of Inders lake), Balkh. region, Syr D., Tien Shan (desert valleys of Centr. Tein Shan, ascending to alt. of 2300 m), N. Pam.- Al, Mtn. Turkm., Kyz. K. (rarely). Gen. distr.: Cin'sheyan' (Kulja), Afghani- stan (?). Described from the east coast of the Caspian Sea. Type in Berlin. Economic importance. Avery valuable forage plant, suitable for production of spring hay and for improvement of spring pastures. Resem- bling A. rytilobus Bge. in chemical composition. 232 309 283. A. rytilobus Bge. Astrag. turk. (1880) 209.-Ic.: Fl. Tadzh. V, t. 20,024) -Pxs. HUF SASM. No: 386, sub nom. A. filicaulis. Annual, very closely related to the preceding species; differing in pods, these capitately gathered, ovoid to oblong-ovoid, 6-8 mm long, 3-4 mm broad, coarsely reticulate-rugose, terminating in a curved beak. Deserts in the north of Soviet Central Asia and submontane plains, in the foothills and lower levels of Central Asian mountains. Chiefly as- sociated with loess plains and foothills, ascending in the south up to the witch-grass steppe zone. Distributed more to the south than A. filicaul- is, it is more prevalent in the southern regions of Soviet Central Asia, while A. filicaulis is largely confined to the northern part. - Soviet Centr. Asia: Ar.-Casp., Balkh. region, Tien Shan, Pam.-Al. (except E. Pamir and Shugnan), Mtn. Turkm., Syr D., Amu D. Gen. distr.: Iran. Described from the surroundings of Tashkent and Samarkand. Type in Leningrad. Economic importance. Avery valuable forage plant, widely dis- tributed in the foothill and low-mountain pastures of Soviet Central Asia. Suitable for production of spring hay and for undersowing of pastures. The hay contains about 3 percent fats, 19.44 percent protein, 4.54 percent starch, 25.56 percent cellulose, and 11.5 percent ash. In favorable condition the plant attains 40 cm in height. Series 2. CARINATI Gontsch. - Flowers whitish-pink, in compact heads borne 1-2 on short axillary peduncles; pods crowded and stellately spread- ing in practically one plane, lance-oblong, with prickles along the center of the valves, subquadrangular in cross-section, thinly coriaceous. 284. A. oxyglottis Stev. in M.B. Fl. taur.-cauc. II (1808) 192; Bge. Astrag, seront. 1, 11910) 9; Boiss’ FL. ori, 229; Ldb. Fl. Ross.’ 1, 636; Bge. Astrag. turk. 208; Grossh. Fl. Kavk. II, 296; Schmalh. Fl. I, 270; Fl. Tadzhik. V 258.-A. psiloglottis Stev. ex DC. Prodr. II (1825) 288; Grossh. Fl. Kavk. II 296.-Ic.: Fl. Tadzh. Vt. 24, 2.-Exs.:H.F.R. No. 1864. Annual, 5-15 cm tall, appressed-pubescent; stems erect or ascending, commonly branched from base, 7-12 cm long, puberulent with appressed or subappressed white hairs, rarely plants subacaulescent; stipules free, ad- nate at base to petiole, triangular-lanceolate, acuminate, 2-3 mm long, white - or black-ciliate; leaves 3-8 cm long, the petiole 1/2-2/3 as long as the rachis, both petiole and rachis sparsely appressed-puberulent; leaflets 4-6 (8) pairs, cuneate-oblong, retuse, 6-12 mm long, appressed-hairy beneath, glabrous or rarely appressed-hairy above; peduncles axillary, shorter than to rarely equaling the leaves, (0.4) 1-2 (4) cm long, puberulent with appressed white hairs; flowers small, 4-8 in heads, often in 2's-3's on same peduncle; bracts ovate to oblong-ovate, 0.5-0.7 mm long, white- or black-ciliate; calyx campanulate, ca. 3 mm long, minutely puberulent with appressed white and black or pure white hairs, the tube 3-4 times the length of the linear teeth; corolla whitish-pink; standard 5-7 mm long, the limb oval to oblong-oval, faintly distinct from and (3) 4-5 times as long as the broad claw; wing-petals 4-6 mm long, the limb oblong, slightly dilated upward, 1.5 times the length of the claw; keel 3-4 mm long, the broad ob- tuse limb 1.5 times the length of the claw; stigma glabrous; pods crowded, stellately spreading, lance-oblong, slightly arched-recurved, nearly straight, (8) 10-12 (16) cm long, 2.5+3 mm broad, acute, bluntly carinate ventrally, finely sulcate dorsally, carinate along the center of valves with ridge - 233 310 margins tuberculate by prominent nerves or smooth, subquadrangular in cross-section, thinly coriaceous, glabrous, lustrous or appressed-puberu- lent. Fl. April-June (July); fr. May-July (August). (Plate XXI, Figure 6). Clayey or gravelly wormwood or rarely sandy deserts in foothills and low mountains; in Crimea and Caucasus, stony slopes and often more or less sandy sites. - European part: Crimea, L. V. (lower Volga); Caucasus: S. and E. Transc.; Soviet Centr. Asia: Ar.-Casp., Balkh. region, Kyz.K., Kara Kum, Mtn. Turkm. (Kopet Dagh, B. Balkhany), Amu D., Syr D., Pam.- Al., Tien Shan (W.). Gen. distr.: E. Med. (Syria), Bal.-As. Min. (Asia Minor), Arm.-Kurd. (N. Iran), Iran. (Iran). Described from surroundings of Sudak in the Crimea. Type in Helsinki. Series 3. EGIBBOSI Gontsch.- Flowers yellow, in rather compact heads borne on very short peduncles; pods approximate in a short raceme, usually arranged in a single plane in 2-3 whorls, oblongly linear-lanceolate, acute, without distinct lateral tubercles at base. 285. A. cornubovis Lipsky in A.H.P. XVIII (1901) 23. Annual, 6-17 cm tall, branched from base; stems prostrate or ascending,- (2) 5-23 cm long, densely vested with short ascending white hairs; stipules free, adnate at base to petiole, triangular, acute, 2-3 mm long, white- ciliate; leaves 3-5 cm long, the petiole as long as the rachis, both densely shaggy-pubescent; leaflets 3-4 pairs, oval, broadly emarginate, broadly cuneate or nearly round at base, 5-10 mmlong, glabrous above, very dense- ly shaggy canescent beneath with short appressed white hairs; peduncles somewhat exceeded by the leaves, 4-8 mm long, densely pubescent, be- coming thick and elongating in fruit up to 10 mm; flowers on densely white - hairy pedicels ca. 1 mm long, gathered in (3) 4-6-flowered rather compact racemes 5-10 mm long; bracts broadly lanceolate, ca. 1 mm long, white- hyaline, white-hairy; calyx campanulate, 3 mm long, densely hirsutulous with subappressed hairs, the lanceolate teeth 0.6-0.7 mm long; corolla yellow; standard 5.5-6 mm long, the limb ovate, round-tipped, 4-5 times as long as the broad claw; wing-petals ca. 5 mm long, the limb oblong, entire, 2/3 as long as to equaling the claw; keel 4.5 mm long, the limb in- equilaterally obovate, rounded-obtuse, 1.5 times as long as the claw; ovary sessile, white-pubescent; pods sessile, gathered in heads of 4-6, sub- horizontally spreading, linear-lanceolate to lance-oblong, acute, 1.2-1.4cm long, slightly arched-recurved, 3.5-4 mm broad, compressed laterally (especially toward the ventral suture), more or less inflated toward the dorsal suture, carinate ventrally and concave along the suture, deeply grooved dorsally and convex along the suture, densely white pubescent, terminating in a straight beak ca.1mm long, nearly bilocular (the narrow septum ca. 1.5 mm broad), 6-8-seeded. Fl. April-May; fr. May-June. Sands and sandy clay soils, in the desert plain region, rarely in the foothills. - Soviet Centr. Asia: Kyz.K. (inundated forests of Amu Darya and residual mountains near its lower reaches), Kara Kum (lower reaches of Amu Darya), Amu D. (surroundings of Kara-kul'), Mtn. Turkm. (Pere- val'naya). Endemic. Described from Turkmenia (Pereval'naya). Type in Leningrad. Series 4. CRUCIATI Gontsch.-Flowers yellow, in compact heads borne on long peduncles; pods crowded and stellately spreading in a single plane, bigibbous at base inthe shape of an obliquely truncate horseshoe, coriaceous. 234 311 312 286. A. cruciatus Link, Enum. Hort. Berol. II (1822) 256; Bge. Astrag. pexont. i) 11> ll, 6; DE. Prodr. Il; 268; ‘Boiss: Flor: Il})225; Schmalh. Fl. I, 271.-A. Asterias Stev. ex Ldb. Fl. Ross. I (1842) 636; Grossh. Fl. Kavk. II, 296.-A. stella M.B. Fl. taur.-cauc. III (1819) 191, excl. syn., non Gouan.-Exs.: Herb. Fl. Cauc. No. 372. Annual, 15-20 cm tall, branched from base; stems ascending to sub- decumbent, 10-15 cm long, rather densely covered with short white hairs, these often interspersed beneath the nodes with black hairs; stipules free, adnate at base to petiole, triangular to triangular-lanceolate, acute, ca. 4 mm long, covered with white and predominantly black hairs; leaves 2.5- 4 cm long, the very short or obsolescent petiole and the rachis densely cov- ered with short white hairs; leaflets 6-8 pairs, lanceolate, acute to sub- obtuse, 6-9 mm long, the lower ones elliptic, 2-4 mm broad, covered on both sides with subappressed white stiff hairs; peduncles shorter than or rarely equaling the leaves, 1.5-3.5 cm long, densely covered with short white and upward mixed white and black hairs; racemes compactly capitate, ca. 1 cm long, 3-5 flowered; bracts lanceolate, 1.5-2 mm long, white- hyaline, covered with white and predominantly black hairs; flowers sub- sessile; calyx campanulate, ca. 4mm long, densely hirsute with black hairs; interspersed with few isolated white hairs, the linear-subulate teeth about equaling the tube; corolla yellow; standard ca. 8 mm long, the limb oblong- rhomboid, attenuate upward, round-tipped, slightly retuse, 2.5-3 times as long as the broad claw; wing-petals ca. 4.5 mm long, the limb oblong, en- tire, twice the length of the claw; keel equaling the wings, the limb in- equilaterally oval, obtuse, as long as the claw; pods sessile, commonly in 3's, stellate, linear-lanceolate, more or less inflated, bigibbous at base, somewhat arched-incurved, 1-1.5 cm long, rounded ventrally, deeply grooved dorsally, rigidly coriaceous, pubescent with ascending white hairs, nearly fully bilocular (the septum ca. 1 mmbroad), 6-10-seeded. Fl. April; fr. May. (Plate XXI, Figure 3). Dry, often stony, mountain slopes and foothills. - Caucasus: Dag., S. and E. Transc.; Soviet Centr. Asia: Mtn. Turkm. (W. Kopet Dagh). Gen. distr.: Med., Bal.-As. Min. (Asia Minor), Arm.-Kurd., Iran, Mesopota- mia, Arabia. Described from cultivated specimens of the Berlin Botanical Garden without indication of origin. Type in Berlin. Series 5. TRIBULOIDES Gontsch.- Flowers violet or whitish, in com- pactly capitate racemes, these sessile or borne on very short peduncles; pods oblong to lance-oblong, bigibbous horseshoe-shaped at base, stellately Spreading in a single plane, coriaceous. 287. A. sinaicus Boiss. Diagn. ser. I, IX (1849) 57. Annual, 3-8 cm tall, mostly branched at base; stems ascending or pros- trate, 2-8 (15) cm long, rarely obsolescent, densely vested with ascending white hairs; stipules free, adnate at base to petiole, ovate-triangular to lance -triangular, acuminate, 3-4.5 mm long, green, covered with white and black hairs; leaves (2.5) 3-5 (7) cm long, the petiole andthe much longer rachis densely covered with ascending white hairs; leaflets (7) 8-10 pairs, oblong-oval, round-tipped or slightly retuse, (2) 4-8 (11) mm long, densely covered on both sides with appressed hairs (sometimes sparsely above); inflorescence compactly capitate, axillary, sessile or borne on a very short peduncle (up to 0.5 cm long), ca. 1 cm long, few-flowered; peduncles dense - ly covered with ascending white hairs; flowers subsessile; bracts ovate, 235 313 acuminate, ca. 3 mm long, covered with white or mixed white and black hairs; calyx campanulate, 4-5.5 mm long, hirsute with subappressed black and white hairs (black hairs sometimes few, confined to the teeth), the sub- ulate-linear teeth somewhat shorter than the tube; corolla violet; standard 7-8 mm long, the limb ovate to oblong-ovate, retusely obtusish, twice the length of the claw; wing-petals 6-6.5 mm long, the limb oblong scarcely asymmetrically retuse, somewhat longer than the claw; keel 4-5 mm long, the limb broad, obtuse, somewhat shorter than the claw; ovary sessile, densely hairy; pods sessile, approximate, in (3's) 4's-5's, stellate, lance- oblong, scarcely recurved, (8) 10-13 mm long, 2.5-3 mm broad, 3-3.5 mm thick, acuminate to a subulate beak ca. 1 mm long, progressively attenuate from base, not distinctly gibbous, rounded or scarcely carinate ventrally, deeply grooved dorsally, coriaceous, puberulent with ascending hairs and densely hirsute with spreading white hairs, bilocular, few-seeded.F1. April; fr. May. Dry, often stony slopes. - European part: Crimea (south coast). Gen. distr.: Bal.-As. Min. (Greece, Crete, Anatolia, N. Syria). Describedfrom Mount Sinai. Type in Geneva. 288. A. tribuloides Delil, Fl. Aegypt. Illustr. (1813) 70; Bge. Astrag. geront. II, 7; Bge. Astrag. turk. 208.-A. kirghisicus Schtschegleev in Bull. Soc. Nat. Mosc. XX VII (1854) 161.-Exs.: H.F.A.M. No. 21.- levee Mad ze ayiate wZiGes ate Annual, (1.5) 3-8 (13) cm tall; stems branched nearly from base, de- cumbent or ascending, (0.5) 2-15 (24) cm long, covered with subappressed or spreading gray hairs; stipules free, adnate at base to petiole, lanceolate or rarely lance-ovate, green, white-ciliate on the margin; leaves (1.5) 3- 6 cm long, the petiole 1/4-1/2 as long as the rachis, both petiole and rachis covered with ascending white hairs; leaflets 5-7 (8) pairs, oblong-elliptic, lanceolate or sublinear-lanceolate, (2) 5-10 mm long, acute, rarely sub- obtuse, covered on both sides with appressed hairs; flowers subsessile; racemes not exceeding 1 cm in length, capitate, loosely 2-6-flowered, ses- sile or borne on a very short peduncle (not exceeding 1 cm); peduncles axillary, covered with ascending white hairs; bracts lanceolate, acuminate, 1.5-2 mm long, black- and white-ciliate; calyx narrowly campanulate, 4- 5 mm long, black- and white-hirsute, the tube 1.5 times as long as orrare- ly equaling the linear teeth; corolla whitish, rarely violet; standard 6-8 mm long, the limb oblong-elliptic or elliptic-oblong, slightly retuse, 5-7 mm long, gradually attenuate at base into the claw; wing petals 5-7 mm long, the limb narrowly oblong, entire, obtuse, equaling to somewhat exceeding the claw; keel 4.5-5 mm long, the limb broad, obtuse, 2/3 the length of the claw;npods sessile, stellate, oblong, (5)6-10mm long, obtuse, slightly arched -incurved, carinate ventrally, grooveddorsally, subtriquetrous, obtusely bigibbous at base, pubescent and hirsute with more or less spreading rarely appressed hairs. Fl. March-June; fr. April-July (August). (Plate XXI, Figure 5). Clay and saline clay, chiefly wormwood deserts in plains, stony foothills and low mountains, rarely sands; in the Caucasus on dry stony slopes. - European part: L. V. (surroundings of Astrakhan); Caucasus: E. and S. Transc.; Soviet Centr. Asia: Ar.-Casp. (except the northern part), Kyz.K., Kara-Kum, Mtn. Turkm. (flatlands region at foot of Kopet Dagh), Amu D., Syr D., Pam.-Al. (ascending on Zeravshan up to 1800 m), Tien Shan (Mogol- tau). Gen. distr.: W. Med. (Canary Islands, Algeria, Tunisia, Tripolita- nia, Egypt, Syria), Arm.-Kurd. (Azerbaijan), Iran, Arabia, Mesopotamia, N. W. India. Described from Egypt. Type in Paris. 236 Note. Erroneously reported for Crimea. The western limit of A. tri- buloides does not reach beyond Lower Volga, whereas the clearly dis- tinguishable A. sinaicus Boiss. is distributed in the Caucasus. Series 6. AMMOPHILI Gontsch.- Flowers violet, in compact, sessile or short-peduncled heads, one peduncle often bearing two heads; pods crowd - ed in compact heads, obliquely upright, inequilaterally ovoid-oblong or ob- long ovoid, not gibbous at base, thinly coriaceous, In addition to species occurring in the USSR, this series includes a num- ber of Iranian species. 289. A. ammophilus Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 335; Bge. Astrag. geront. II, 10; Boiss. Fl. or Il, 228; Bge. Astrag. turk. 209; Grossh. Fil. Kavk. II, 296.-A. persepolitanus Boiss. Diagn. sér. I (1849) 60. Annual, 4-9 cm tall; stems branched from base, prostrate, 4-20 cmlong, puberulent with appressed or rarely spreading hairs; stipules free, adnate at base to petiole, triangular-lanceolate, 1-2mm long, white-ciliate; leaves 3-6 cm long, the petiole 1/2-2/3 as long as to as long as the rachis both petiole and rachis puberulent with spreading hairs; leaflets 4-6 pairs, cune- ate-oblong, (3) 4-8 mm long, retuse, appressed-hairy beneath; peduncles axillary, shorter to rarely somewhat longer than the leaves, 0.5-3 cm long, puberulent with white appressed or rarely spreading hairs; flowers sub- sessile, 3-10 in compact heads, one peduncle often bearing 2 heads; bracts ovate, ca. 1 mm long, white-hyaline, white-ciliate; calyx 2.5 mm long, white-puberulent, the tube 1.5-2 (3) times the length of the linear teeth; corolla violet; standard ca. 6-16.5 [?] mm long, the limb oblong-ovate, slightly retuse, twice the length of the claw; wing-petals 4.5-5.5 mm long, the limb oblong, scarcelyretuse, 1.5-2 times the length of the claw; keel somewhat shorter than the wings, 3.5-4.5 mm long, the broad rounded-ob- tuse limb equaling the claw; pods sessile, in compact heads, obliquely up- right, inequilaterally oblong, slightly arched-incurved, 7-9 mm long, 3- 3.5 mm broad, carinate ventrally, deeply grooved dorsally, subtriangular in cross-section, acuminate, faintly cross-wrinkled or smooth, glabrous or appressed-puberulent, nearly straight or slightly convex on the ventral side, strongly convex on the dorsal side (especially in lower part), few- seeded. Fl. April-May; fr. May-June. (Plate XXI, Figure 4). Gravelly slopes of foothills and low mountains, rarely sands, in the plain region of Soviet Central Asia. - Caucasus: Dag., E. and S. Transe. ; Soviet Centr. Asia: Balkh. region, Kyz.K., Kara Kum (lower reaches of Amu Darya), SyrD., Pam.-Al.(S.), Tien Shan (Mogoltau), Gen. distr. : Iran, (Iran, Afghanistan, Beludzhistan). Described from the area between Arganaty mountains and source of the Sasyktau at eastern extremity of Lake Balkhash. Type in Moscow, cotype in Leningrad. 290. A. vachanicus Boriss. et A. Kor. in Fl. Tadzhik. V (1937) 655. - eo Wik madzh. Vi, t.°24;° 4. Annual, stems ascending, branched, 13-15 cm long, vested with white hairs; stipules linear-subulate, ca. 2 mmlong; leaves ca. 5 cm long; leaf- lets 6-7 pairs, cuneate-oblong, 6-7 mm long, 2-3 mm broad, deeply retuse, glabrous above, covered beneath with appressed hairs; flowers 5-6 in com- pact heads; peduncles long, exceeding the leaves, bearing 2-3 distant heads; bracts linear, hyaline, very small; calyx campanulate, 3-4 mm long, 237 oe | white -hirsute, the tube twice the length of the linear teeth; corolla reddish; standard elliptic, slightly retuse, ca. 5 mm long and 2 mm broad, the claw obsolescent; wing-petals ca. 4 mm long; keel barely shorter than the wings; pods sessile, ovoid-oblong, obliquely upright in heads, ca. 8 mm long, acuminate to a short beak, carinate ventrally, deeply grooved dorsally, sub- triquetrous, thinly coriaceous, glabrous, nearly smooth, bilocular. Fl. and fr. July. Gravelly screes at alt. ca. 2950 m.-Soviet Centr. Asia: Pam.-Al. (Bakhan). Endemic. Described from Katta-Kharov river valley. Type in Leningrad. 291. A. cinerascens M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Annual, 2-4 cm tall, branched from base; stems rather stout, prostrate, 1-2 cm long or shorter, rarely up to 8 cm long, shaggy-pubescent with white spreading hairs; leaves 2-3.5 cm long, the petiole and the longer ra- chis shaggy-puberulent with spreading hairs; leaflets 4-5 pairs, cuneate- oblong or obovate, broadly emarginate, 4-7 (9) mm long, glabrous above, densely shaggy-pubescent beneath with spreading hairs; flowers sessile or nearly so; heads compact, small, (3) 5-8 (9)-flowered, sessile or nearly so, the very short peduncles not exceeding 2-3 mm, densely shaggy-pubes- cent with spreading hairs, firm, axillary; bracts small, ca. 1 mm long, ovate, white-hyaline, white-ciliate; calyx ca. 1mm long, white-pubescent, the lance-linear teeth equaling the tube; corolla pale violet; standard ca. 5 mm long, the limb oblong, ca. 1-2 mm broad, broadly and shallowly truncate -emarginate, the claw obsolete; wing-petals 3.5 mm long, the limb oblong or spatulate-obovate, round-tipped, 1.5 times the length of the claw; keel ca. 3 mm long, the limb broad, rounded-obtuse, somewhat longer than the claw; ovary sessile; pods crowded in heads at the ends of branches, oblong-ovoid, straight, triangular in cross-section, carinate ventrally, broadly sulcate dorsally, acute, 5-6 mm long, shaggy-puberulent with spreading hairs, gray, 7-seeded; seeds ear-shaped, ca. 5mmlong, smooth, brown. Fl. April; fr. May. Gravelly slopes of residual mountains. - Soviet Centr. Asia: Kyz.-Kum (mainly Sultanuizdag and Bukantau residual mountains). Endemic. De- scribed from Sultanuizdag mountains. Type in Leningrad. Section 52. DIPELTA (Rgl. et Schm.) Bge. in A.H.P. VII (1880) 36. - Genus Dipelta Rgl. etSchmalh. in A.H.P. V (1878) 578.-Didymope- lia Taubert in Engl. und Prantl. Pflanzenfam. III, 3 (1894) 284. - Annuals, vested with basifix hairs; racemes loose, borne on long axillary peduncles; bracteoles none; calyx campanulate; corolla violet; keel equaling the wings; pods sessile, double, separating in maturity along the sutures into two rounded-ellipsoid or nearly rectangular components, submembranous, cov- ered with fine pubescence interspersed with long spreading uncinate hairs. - A monotypic Soviet Central Asian section. 292. A. dipelta Bge. in A.H.P. VII (1880) 368.-Dipelta turkestan- ica Rgl. et Schmalh. in A.H.P. V (1878) 578 cum icone. -Didymo - pelta turkestanica™ Tauber Wcr—Ic2: Fl. "Padzh. Vit. 20, oe eee H.F.A.M. No. 383. Annual, 5-20 (30) cm tall; stems erect or ascending, (3) 8-30 cm long, branched, vested with appressed white hairs; stipules free, adnate at base 238 318 319 to petiole, linear-subulate, 2-3 mm long, white-ciliate; leaves 2-3 cm long; leaflets 3-5 pairs, oblong-elliptic to linear, 6-10 (12) mm long, subobtuse or slightly retuse, canescent on both sides with short appressed hairs; peduncles 2/3 as long to as long as the leaves, slender, covered with short appressed white hairs; pedicels covered with white and black hairs, ca. 0.5 mm long; racemes loosely and distantly (2) 3-10-flowered, (0.5) 1-4 cm long; bracts lanceolate to narrowly linear, equaling the pedicel, covered with white and black hairs; calyx campanulate, ca. 3 mm long, black- and white -hirsute, the linear-subulate teeth about equaling the tube; corolla pale violet; standard 3.5-4 mm long, the limb broadly ovate, 3-3.5 mm long, retuse, abruptly contracted at base into and 5-6 times as long as the claw; wing-petals 3-3.5 mm long, the limb oblong, rounded-obtuse, 1.5 times the length of the claw; keel equaling the wings, the broad rounded-obtuse limb twice the length of the claw; stigma naked; pods sessile, double, 3-4 mm long, 4-5 mm broad, the valves (halves) strongly flattened, rounded-elliptic to nearly square, submembranous, covered with short pubescence inter- spersed with long spreading uncinate hairs, 2-4-seeded. Fl. May-June; fr. May-July. (Plate XX, Figure 4). Loess foothill slopes in the ephemeroid vegetation and xerophytic tall- grass belt in the south; elevated foothill and mountain slopes in the witch- grass steppe and ephemeroid vegetation belt and the lower zone of the wood- land and scrub formation. Rare in the lower part of the juniper zone. - Soviet Centr. Asia: Mtn. Turkm. (W. Kopet Dagh), Pam.-Ai., Tien Shan. Endemic. Described from the surroundings of Karaguz in Karatau Moun- tains. Type in Leningrad. Subgenus V. TRAGACANTHA* Bge. Astrag. geront. (1869) 120. - Gen. Tragacantha Mill. The Gard. Dictionary, ed. VI (1752); Stev. in Bull. Soc. Nat. Mosc. IV (1832) 269; ibid. XXIX (1856) 152; Wallr. ex. Endl. Gen. (1837) 370. Flowers sessile, persistent, 2-15 in the leaf axils, sometimes borne on a short stout hairy peduncle 1-4 mm long, approximate and hence forming an apparent inflorescence, this elongate-oblong or spherical; each flower subtended by a bract adnate to calyx, the bracts of the same inflorescence varying in breadth; bracteoles sometimes present; calyx mostly obconic, pointed at base, soft or indurated, sometimes readily splitting nearly to base, densely lanate with long white hairs, often glabrous at base, the teeth woolly-villous onbothsides; corolla white, yellow, violet, red or reddish; standard commonly pandurate, constricted at the middle, rarely without marked transition between limb and claw, the limb base often auriculate; wings, keel and staminal tube fused in lower part, rarely nearly free; sta- mens diadelphous, nine united through much of their length, the free sta- men adnate to standard base; anthers spherical; ovary sessile, hairy, the style naked or hairy, long, filiform; pod included in calyx, rounded or ob- long, 5-6 mm long, densely hairy, unilocular, commonly 1-seeded; seeds reniform, mostly ca. 3 mm long. - Spiny frutescent or suffrutescent plants, at most 1 m tall, mostly 20-50 cm or less, commonly branched nearlyfrom base or from somewhat higher up, rarely unbranched; stems weakly de- veloped, 7-10 cm long; vesture composed of simple hairs; branches densely * From the Greek tragos = vipers’ grass, and acanthus = spines, spiny plant. 239 320 leafy, clothed in imbricated stipules; leaf rachises spinescent; stipules acuminate, connate and adnate to petiole, shaggy, especially when young, or glabrous; leaves paripinnate, the spinescent rachises becoming indurated and persistent together with stipules on old branches; leaflets always ter- minating in a prickle. All species of the subgenus Tragacantha are characterized by the presence of gum and mucilaginous cell walls in the medullary rays and in the pith (the broad rays containing 6-20 cell layers), numerous conducting vessels and narrow annual rings, cellulose -coated fibers, wide cortex, and thick-walled epidermis cells. Economic importance. Inview of their gum content, many species of tragacanth are exploited for this highly valuable product which is indis - pensable for various light industries. The greatest demand comes from the textile industry in which the gum is used as a thickening in preparation of boiled dyes for calico printing, in the application of textile dyes, and in the dressing of silk fabrics and lace. Gum tragacanth is used in the con- fection industry as a thickening agent, in tanning industry for leather dress- ing, in the printing trade, in production of matches, and in preparation of plastic materials. It is also used in the production of glue, water colors and ink (to supply the gloss); in perfumery, in pencil manufacture, and in the paper industry, as a binding agent. For pharmaceutical purposes, the gum is employed in the making of pills, tablets, and certain medicaments; another use is in the production of superior sorts of soap. In areas of natur- al occurrence, in Soviet Central Asia and in the Caucasus, tragacanths are used as forage plants; they are eaten by camels in fall and winter and the plant tips are grazed by sheep in spring; the spines are burned off and then the bushes are softened by beating or are milled into powder, and are fed to livestock in case of fodder deficiency. Tragacanths are sometimes used as fuel. Gum tragacanth is used for the preparation of nutrient media in bacteriological laboratories, as an agar substitute. It is reported (Haus - knecht) that, inIran, sugar extracted from certain tragacanth species is used in food. Note. Tragacanth is known by various names, derived from many languages and apparently referring to different species and various kinds of gum; thus, tragakant, tragant, adragant, dragant, gummi- tragakant, [Russian]; tragacanth (English); gomme adragante, gemma-dragant (French); Traganth or Tragakant (h) (German); dragant (Swedish); tragacanth (Dutch); tragakanta (Greek); go- mo tragacantha (Spanish); gomma agragante, gomma dra- gante (Italian). Names current in Iran, India and Afghanistan: katira, katad, kettira, kutera, khitira, khaluzia; In Iran tragacanth shrubs are called kon or gabina, while the gum is referred toas keti- ra-ye-arrebor, which means incised tragacanth, ketira-ye-gol', ketira-ye-nich, ketira-ye-kharmeni — inmarket parlance; Kurds call tragacanth guini; in Turkey the plant is called ketereh ortirnak- kitre, whereas the name given to it by the Turkish population of Armenia is gyavan; inArmeniaitis called gaz, in the Karachai language kipri; in Asia Minor tragacanth shrubs go by the name ketre ortragakanti- nos. Inferior sorts of tragacanth are known in Afghanistan as anzerus. The Turkmenian names for tragacanth are teken, guvan, or guven. 240 321 Stems poorly developed, 7-10 cm long; unbranched shrubs; leaves subradical, large, 15-35 cm long; leaflets large, 13-45 mm long, 7- 14 mm broad; flowers crowded in large subradical rounded or oblong solitary heads 4-6 cm broad (Caucasus, Asia Minor).......... Ze Stems developed; shrubs 8-100 cm tall, densely branched; leaves 1-10 (15) cm long; leaflets 5-30 mm long, 2-11 mmbroad; flowers crowded in the leaf axils in various inflorescences 0.5-4 cm broad....... 3. Leaflets 7-9 pairs; calyx 12-15 mm long, the tube 4-5 times the length of the thickish teeth; standard 20-22 (25) mm long, the limb twice as long as the claw, with broad obtuse auricles at base; inflorescence SPNEEICAURIW ED isder OLE t MN iaticms. ON etic hraihet Me 351. A. Lagowskyi Trautv. Leaflets 8-11 (14) pairs; calyx 13-18 mm long, the tube 2-3 times the length of the teeth; standard 20-28 mm long, the limb about equaling to somewhat exceeding the cuneately attenuate claw, with acutish auricles at base; inflorescence ovoid....... 352. A. oleifolius DC. Bracts conspicuous, large, navicular, 10-12 mm broad when expanded, ovate or broadly ovate; flowers in 2's in the leaf axils; calyx shorter togslightly longer ithanithercorolla engine tis awstats es 4, Bracts more or less concealed among the calyces, 1-3 mm broad when expanded, carinate or not so, filiform, linear, or lanceolate to oblong; flowers (3) 4-12 in the leaf axils and then calyx mostly equaling or very slightly shorter or longer than the corolla, or flowers in 2's or 3's in the axils and then calyx much shorter than the corolla ......... 3. Bracts 12 mm broad when expanded, navicular, firmly united with calyx and covering it up to above the middle, overlapping the bract of the twin flower, obliquely truncate, rounded at base, glabrous except the long-ciliate margin; calyx 20 mm long, the filiform teeth partial- ly glabrous; stipules glabrous, ciliate-margined; leaves covered be- neath with scattered hairs.......... 323. A. prominens Boriss. Bracts 5-6 mm broad when expanded, imbricated and not exserted from the leaf axil; calyx 12-15 mm long, the teeth hairy; stipules and leavestpannose: Seo husienoe PS oa. 322. A. transoxanus Fisch. Flowers 2-3 in the leaf axil, covered by stipules adnate to spinescent leaf rachis; calyx commonly much shorter than the corolla and com- monly splitting down to base, or not splitting and then calyx teeth much SOMES Ladnh. ieMetlelka Leketne Rome’ cris | ) LN AA) eee ee Piet See sf 52. Leaflets white-pannose, 8-12 mm long, 3-4 mm broad; leaf-rachises white-pannose; stipules 11 mm long, pannose with white spreading hairs; inflorescence 2.5-3 cm in diam. ; flowers 4-5 in the leaf axil; calyx 12-14 mm long; corolla 15-18 mm long (Caucasus) ........ ee ere ie Shc cts, Met ets: ded ee an aloe te ie te Soh 344. A. compactus Willd. Leaflets silvery with appressed hairs, 5-15 mm long, 1-3 mm broad; leaf-rachises silvery with appressed hairs; stipules 9-10 cm [sic] long, silvery on the back with appressed hairs when young, becoming gla- brous; inflorescence 1.5-3 cm in diam. ; flowers 5-7 in the leaf axil; calyx 15-18 mm long; corolla 18-20 mm long (Kopet Dagh)....... PMS Se ylehcs Poke had oul tsp Pare 34 de. c0 ged ois core of Eke fe 342. A. cerasocrenus Bge. (48) Stipules large, 25-30 mm long; leaflets 20-23 mm long, 5-7 mm broad, glabrous above, sparsely hairy mainly on the midvein; inflor- escence 4.5-5 cm broad, 5-7 cm long; calyx 22-25 mm long, the teeth 3-4.times, the length of the-tube (PamssAl.):). 0. 06). ees - 6 8G hoe @ eb! Pats eoeeoee: feepeedae ced eetd «fiers: lapedeue’ extern’ 335. A. macranthoides Boriss. Stipules 14-20 mm long; leaflets 10-30 mm long, 2-7 mm broad; in- florescence 3-5 cm in diam.; calyx teeth 13-20 mm long, equaling, scarcely exceeding or twice as long as the tube............. 54. Leaflets silvery with silky appressed or spreading hairs, rarely be- coming somewhat glabrescent; stipules 15-20 mm long........ 56. Leaflets green, glabrous above, or sparsely appressed-hairy on both sides or merely beneath; stipules 10-15 mm long ........... 55: 247 330 55. 56. oT. 58. 59. Flowers 6-8 in the leaf axil; calyx ca. 18 mm long, the teeth equaling the tube; corolla ca. 21 mm long; spines 7-10 cm long, slender, slightly curved, all erect; leaflets 5-7 pairs, 10-15 mm long, 3-5mm BRO AG wh ioout tH peiaigr sh cee dpoeh ets aaa, se eerdee pines 339. A. pycnanthus Boriss. Flowers 10-12 in the leaf axil; calyx about 18-25 mm long, theteeth about twice as long as the tube; corolla ca. 17 mm long; spines 5-7cm long, straight, stout, somewhat spreading; leaflets 4-5 pairs, 15- ly mm jong,.3-4 (5) mmuybroad jase va 338. A. pterocephalus Bge. Inflorescence 3-3.5 cm broad, 4-8 cm long, cylindric; leaflets 3-5 pairs, (13) 15-30 mm long, 2-5 mm broad; calyx 13-15 mm long, gla- brous at base, the teeth twice the length of tube (Caucasus)....... REE Ree Tee ee On pee eae ae 343. A. strictifolius Boiss. Inflorescence 3-5 cm in diam., spherical; leaflets 4-7 pairs, 10-23 mm long, 3-11 mm broad; calyx 15-20 mm long, covered throughout with VORA WG So rors cpuei Sb cues oak oi satennewems od ieee ease iciia slay alts ala Chicas ee 57. Calyx-teeth twice as long as the tube; inflorescence 3-3.5 cm in diam., leaflets not more than 10-20 mm long, 3-7 mm broad......... 58. Calyx-teeth barely longer than the tube; inflorescence (3) 4-5 cm in diam. ; leaflets mostly large, (10) 15-30 mm long, 3-11 mm broad. Sst typaifairel cue? Cayaihe, le sspapgon an le. bischi lc, tabs A, 1. 6) (mala. ). A spiny shrub not more than 60 cm tall; branches stout, elongated, ca. 15 cm long, densely covered with old spinescent leaf rachises; spines ap- proximate and spreading at nearly right angles, firm, white-pubescent, slightly twisted, 3-5 cm long; stipules coriaceous, ca. 8 mm long, densely hairy outside becoming glabrous, ciliate-margined, ovate, broadly linear- acuminate toward apex; leaflets 6-8 (9) pairs, densely gray-pubescent or pannose, sometimes bluish-green, 8-10 mm long, oblong to linear-oblong, terminating in a short prickly point; flowers in 2's or rarely 3's in leaf axils, crowded at the base of glabrate shoots, forming an oblong-ovaloid, oblong-cylindric or capitate inflorescence ca. 1.5-2 cm in diam; bracts re- curved, acute, broadly linear, terminating in a short point, shaggy merely in upper half, glabrous below, equaling the calyx; calyx 6-7 mm long, gla- brous at base, white-hairy elsewhere, the teeth as long as the tube; corolla 10-12 mm long, yellow; standard yellow, the limb violet-nerved, attenuate, retuse in upper third, subacutely auriculate at the middle, cuneately at- tenuate into the claw; wing-petals somewhat shorter than the standard, about equaling the keel, the limb half as long as the claw, the same relationship applying to the keel; ovary shaggy; style naked; pod ovoid, hairy, 1-seeded. June-July. Stony dry slopes, at the altitude of up to 2200 m. - Caucasus: Cisc. (upper reaches of Kuban', Karachai), Dag., W. E. and S. Transc., Tal.; Soviet Centr. Asia: Turkm. (Bolshie Balkhanyrange). Endemic. Described from N. Caucasus (Lars). Type in Leningrad. Note. See note in connection with A.denudatus Stev. 299. A. multifoliatus Boriss. in Acta. Inst. Bot. Acad. Sc. URSS, Serle a (936) 219) —Tragacantha multifoliata Boriss. lc. ; Ibid., ser. V, 1 (1938) 506. A densely branched shrub, up to 50 cm tall; branches stout, rather short, densely spiny; spines approximate, 2-3 cm long, erect, slightly twisted, 253 338 339 rather thick, firm, commonly pubescent; stipules pannose-shaggy, pointed at apex, 5-8 mm long; leaflets (7) 8-12 pairs, approximate, small, 5-8mm long, ca. 2mm broad, conduplicate, erect, canescent with dense curly hairs, terminating in a prickle; flowers in 2's in the leaf axils, borne all along the branch, forming a spiciform inflorescence 8-10 mm broad; bracts equaling the calyx, glabrous below, with a bunch of white hairs at apex; calyx 5-6 mm long, glabrous from base to the middle, shaggy upward with short hairs, the teeth half as long as the tube; corolla yellow; standard 10mm long, the limb 3mm broad, half as long as the narrow, cuneately attenuate claw, constricted and obtusely auriculate above the claw; wing- petals 9 mm long, the limb not more than 1 mm broad, 3 mmlong; keel ca. 8 mm long, the oval limb 3 mm long, 1.5 mm broad; ovary covered with short hairs; style glabrous, ca. 6 mm long. June-July. Stony and gravelly slopes. - Soviet Centr. Asia: Mtn. Turkm. (Kopet Dagh), Bolshie Balkhany range). Endemic. Described from W. Kopet Dagh. Type in Leningrad. 300. A. meschchedensis Bge. Astrag. geront, I (1868) 82; II (1869) 140; Boiss. Fl.or.II, 338.—Tragacantha meschchedensis O. Ktze. Rey. gen. (1891) 946; Boriss. in Acta Inst. Bot. Ac. Sc. URSS, ser. 1, 3 (1936) 220; ser. V, 1 (1938) 506. A densely branched bush, up to 60 cm tall; branches long, with graybark, densely spiny, the spines 3-6 cm long, rather stout, firm, somewhat re- flexed, those of the preceding year gray, those of the current year strami- neous, fluffy; stipules broad, short-ovate, lanate at base, ciliate-margined, all white-pannose when young, ca. 8 mm long; leaflets 5-6 (7) pairs, nar- rowly lanceolate to oblong-lanceolate, 6-10 mm long, commonly carinate - conduplicate, at first pubescent beneath, glabrate above, at length becoming glabrous, green, prominently nerved, terminating in a prickle; flowers in 2's in the leaf axils, crowded into an oblong or cylindric inflorescence ca. 1 (1.5) cm broad, borne on the lower part of the branch; bracts pubescent, recurved, about equaling the calyx, oblong, navicular; calyx ca. 5 mm long, splitting down to base, covered with white hairs, acutely conic and glabrous at base, the teeth as long as the tube; corolla ca. 10 mm long, yellow; standard slightly retuse, constricted in upper third, ca. 3 mm broad, in broader parts, cuneately attenuate toward base beneath the acute spreading auricles; wing-petals equaling the standard, ca. 1 mm broad, the oblong limb half as long as the claw; keel 9 mm long, the limb 2/5 as long as the claw, ca. 1.5 mm broad; ovary shaggy, oblong; style pubescent at base. June-July. Stony and gravelly slopes. - Soviet Centr. Asia: Mtn. Turkm. (central part of Kopet Dagh range — Gaudan, Mt. Dushak). Gen. distr.: Iran (Mesh- khed). Described from vicinity of Turuk near Meshkhed. Type in Leningrad. 301. A. meanus Boriss. in Not. Syst. ex Herb. Inst. Bot. Acad. Sc. X (1946).-Tragacantha meana Boriss., l.c. A densely branched shrub with grayish bark; branches rather stout and fairly short, densely spiny; spines straight, somewhat spreading, stout, at first pubescent, 2-3 cm long; internodes pannose; stipules ca. 6 mm long, ovate at base, acuminate, at first densely covered withfelt-like pubescence, becoming glabrous on the margin; leaflets 7-8 pairs, 5-9 mm long, 1.5- 2mm broad, lanceolate, gray, densely covered on both sides with spread- ing felt-like pubescence, terminating in a prickle, carinate-conduplicate, 254 40 lowest pair quite close to stipules; flowers in 2's in the leaf axils, crowded into a capitate inflorescence; bracts navicular, ca. 2 mm broad (1 mm on either side), ca. 5 mm long, the upper side white-hairy down to the middle, the base and sides glabrous; calyx acutely obconic, white hairy except for glabrous 2 mm at base, 5 mm long, the teeth as long as the tube; standard 6-7 mm long, the limb ca. 2 mm broad, 2/5 as long as the claw, auriculate, the claw cuneately attenuate, somewhat enlarged about the middle; wing- petals about equaling the standard and the keel, the limb 2/3 the length of the claw; keel-limb oval, ca. 2.5 mm long, 1mmbroad; ovary ovoid, white - hairy; pods ovoid, ca. 3 mm long, puberulent in lower part, long-hairy up- ward; seeds 2 mm long, 1 mm broad, ovaloid, light-colored, yellowish. July. Soviet Centr. Asia: Mtn. Turkm. (Eastern part of Kopet Dagh range, Dag Bulag and Meany Mtn. Systems). Gen. distr.: Possibly in Iran. De- scribed from Dag Bulag mountains. Type in Tashkent. Note. Described from a single specimen. Description based on flowers and fruits preserved from the previous year. Most closely related to A. chorossanicus Bge. and A. pachystachys Bge.; differs from these species in smaller flowers, shape of flower and a number of other char- acters. 302. A. meracus Boriss. in Not. Syst. ex Herb. Inst. Bot. Acad. Sc. URSS X (1946).-Tragacantha meraca Boriss., l.c. A densely branched shrub, with grayish bark; branches short, recurved, densely spiny; spines 1.5-3.5 cm long, the lower convex toward the stem, arched-recurved on the outside, the upper spreading, at first pubescent, densely imbricated among old stipules; stipules pubescent at first, becoming glabrous, acuminate from ovate base, ca. 8-10 mm long; leaflets 5-6 pairs, oblong-lanceolate, 5-8 mm long, 1-1.5 mm broad, glaucous, covered with spreading pubescence, less so on the upper face, navicular, terminating in a prickle; flowers in 2's in the leaf axils, crowded in an ovaloid inflores- cence ca. 1 cm broad; bracts linear, ca. 1.5 mm broad, 5-6 mm long, navicular, white-pubescent in upper part and terminating in a prickle, pu- bescent on the back merely at base; calyx 7 mm long, white-hairy, splitting down to base, acutely conic and glabrous at base, the teeth shorter than the tube; corolla 10 mm long, yellow; standard round-tipped, the limb orbicu- lar, 2.5 mm broad, constituting the upper one-third, lower part of limb slightly constricted, the claw 3 mm broad in upper part, obtusely auricu- late, cuneately attenuate toward base; wing-petals equaling the standard, the limb ca. 1 mmbroad at base, attenuate toward apex, half as long as the claw; keel 10 mm long, the oval limb half as long as the claw; ovary and pod oblong, shaggy; style 6 mm long, hairy merely at base. July-August. Stony slopes. - Soviet Centr. Asia: Mtn. Turkm. (Pul'-i-Khatuma dis- trict; Gyaz-Gyadyk range, Rakhna-Tur). Endemic. Described from Gyaz- Gyadyk range. Type in Tashkent. Series 2. INTERMIXTI Boriss. in Acta Inst. Bot. Ac. Sc. URSS, ser. 1, 3 (1936) 221; Ibid. ser. V, 1 (1938) 508.-Flowers small, 9-10 mm long, ochroleucous; calyx 5-6 mm long, splitting down to base; gum yellowish, with fruit-drop consistency of the Macranthae type; flowers of the Micranthae type. 255 341 303. A. intermixtus Litv. ex Boriss. in Acta Inst. Bot. Ac. Sc. URSS, ser! 1,3 (1936)'222: Ibid. ‘ser. /V,'1°(1938)1508. >" Pragacant ha inters aswel, IXOALSSi5. I, Ce A densely branched spiny shrub, the spines slender, yellow, erect, 2- 4 cm long; stipules glabrous or nearly so, filiformly long-acuminate, cili- ate-margined, 7 mm long; leaflets 5-8 pairs, oblong-linear, green, pu- bescent at first, 5-7 mm long, 1-1.5 mm broad; flowers small, 2-3 in the leaf axils, borne all along the branch, forming an oblong inflorescence 1- 1.5 cm broad; bracts about equaling the calyx, glabrous, pubescent merely at apex; calyx ca. 6 mm long, covered with long woolly pubescence, the teeth longer than the tube, terminating in glabrous prickles projecting from the pubescence; corolla ochroleucous; standard 9 mm long, the obtuse di- lating auricles somewhat broader than the limb, the claw cuneate; wing- petals equaling the standard, the oblong limb 2/3 as long as the claw; keel 8 mm long, the oval limb 2/3 as long as the claw, ovary shaggy, oblong; style naked, 6 mm long. Fl. June. Rocks, at altitude ca. 1700-2000 m.-Soviet Centr. Asia: Mtn. Turkm. (Centr. Kopet Dagh). Endemic. Described from the Kopet Dagh range. Type in Leningrad. 304. A. karakalensis Freyn et Sint. in Bull. Herb. Boiss., ser. 2, IV (1904) 1110.-Tragacantha karakalensis Boriss. in Acta Inst. Bot. Acs Sey WRSS, ser.in3, (@936)0221 7 Mids sex: Iii (oss) 1 s0a: A densely branched shrub, not exceeding 1 m in height, often lower; branches rather stout, fairly short, densely spiny; spines slender, sub- horizontally spreading, glabrate or sparsely pubescent, 1.5-3 cm long; stipules glabrous, membranous, yellowish, ovate, ca. 0.4mm long, acuminate, leaflets 6-8 pairs, glabrous or very sparsely pubescent, rather thick, green, lanceolate, 4-5 mm long, ca. 1 mm broad, terminating in a prickle; flowers in 3's in the leaf axils, crowded into a capitate ovaloid or oblong inflores- cence 1-2 cm long and ca. 1 cm broad, borne at the base of the annual shoots; bracts linear, sparsely pubescent in upper part, shorter than the calyx; calyx 6 mm long, glabrous from base to the middle, covered in upper part with appressed upward projecting hairs, the hairy teeth longer than the tube; corolla yellow, the petals fused at base; standard with a broad cuneate claw, the limb ca. 2 mm broad in upper part, with rectangular auricles at the middle, constricted beneath the auricles; wing-petals ca. 8 mm long, the limb oblong, ca. 3.5 mm long, da. 1 mm broad, enlarged toward base; keel about equaling the standard, ca. 9 mm long, the obtuse limb ca. 3 mm long and 1.5 mm broad; ovary hairy; style naked, pods covered with long white erect hairs, 1-seeded. June-July. (Plate XXIII, Figure 2). Gravelly and stony slopes and screes. - Soviet Centr. Asia: Mtn. Turkm. (Kopet Dagh range, western part). Endemic. Described from the Kopet Dagh range. Type in Leningrad. 305. A. pulvinatus Bge. Astrag. geront. I (1868) 82; II (1869) 142; Boiss. Fl. or. I, 323.-Tragacantha pulvinata, O. Ktze. Rey. gen. (1891) 947; Boriss. in Acta Inst. Bot. Ac. Sc. URSS, ser. I, 3, 222; Ibid., Serie Ve lana 09s A pulvinate shrub densely branched from base, the short branches form- ing cushions 15-30 cm tall; bark dark brown; foliar spines slender, short, 2-2.5 cm long, at first divergent at an acute angle, at length strongly spread- ing, light stramineous; stipules glabrous, pubescent when young, acuminate, 256 densely imbricated, stramineous, sometimes reddish; leaflets sparsely pubescent, 5-6 pairs, oblong, mucronate, 5-6 mm Long) lily omm,: broad: flowers in 2's in the leaf axils, gathered in a small dense spherical capitate inflorescence; bracts linear, navicular, long-acuminate, glabrous at base, hairy on the back, about equaling the calyx; calyx 5-6.5 mm long, shaggy, glabrous at base, the teeth half as long as the tube; corolla yellow; standard violet nerved, 9-10 mm long, the limb short, emarginate, constricted at the middle, gradually passing into the straight elongated claw, somewhat enlarged at apex; wing-petals ca. 8 mm long, the narrowly oblong limb 2/3 as long as the claw; keel equaling the wings, the oval limb 1/3 as long as the claw; ovary shaggy; style naked; pods ovoid, ca. 8 mm long, pubescent, with long white hairs at the tip; seeds olivaceous, rounded-reniform, 2mm long, 1.5 mm broad. July-August. Mountain slopes at altitudes up to 2350 m. - Soviet Centr. Asia: Mtn. Turkm. (Mt. Massinev, Mt. Dalancha near Kheirabad). Gen. distr. : Iran, near Meshkhed. Described from Khorosan province in Iran. Type in Lenin- grad. Series 3. MACRANTH Boriss. in Acta. Inst. Bot. Ac. Sc. URSS, ser. I, 3 (1936) 221; Ibid. ser. V, 1 (1938) 508. -Flowers commonly violet, 10- 13 mm long; calyx 5-10 mm long, splitting down to base; standard-limb broad, subquadrangular; inflorescence 0.8-1.5 cm broad, usually elongate, oblong or cylindric, rarely capitate; gum translucent, of fruit-drop con- sistency, yellowish; plants growing at altitudes of (1500) 1700-2000 m. 306. A. turkmenorum Boriss. in Acta. Inst. Bot. Ac. Sc. URSS, ser. I, 3 (1936) 221; ibid., ser. V, 1, 508.-A. verus auct. non Oliv. ; Bge. Astrag. geront. II (1868) 139; Boiss. Fl. or. II, 321.-Tragacantha turkmenorum Boriss., l.c. A pulvinate shrub, densely branched from base, up to 60 cm tall; branch- es long, with gray bark, densely spiny; spines (3) 5-6 cm long, those of the preceding year grayish-yellow, those of the current year green, pu- bescent, erect, curved; stipules large, lanceolate, broad, convex, fili- formly long-acuminate, glabrous, light yellowish-green, lustrous, faintly pubescent or glabrous when young; leaflets dark green, oblong-lanceolate, 0.5-1.5 cm long, ca. 2 mm broad, long-acuminate, covered with spreading hairs when young, 5-6 rarely 7 pairs, distinctly nerved above; flowers in 2's-3's in the leaf axils, crowded along the entire length of the branch into an oblong spiciform inflorescence and on abbreviated branches into rounded inflorescences to 1.5 cm broad; bracts hairy, lanceolate, falcate, navicular, terminating in a point, equaling the calyx; calyx 8-9 mm long, glabrous at base, bluish-gray and shaggy throughout, the long-hairy teeth equaling the tube, green-tipped; corolla 12 mm long, lilac or yellow; standard 4.5 mm broad, the emarginate limb 2/5 as long as the claw, the claw short, attenu- ate in lower part, enlarged upward; wing-petals 11 mm long, the lanceolate limb 2/3 the length of the claw, ca. 1 mm broad; keel 10-11 mm long, the limb half as long as the claw, ca. 1.5 mm broad; ovary shaggy; style pu- bescent merely at base. Fl. June-July; fr. June-August. Steppe slopes, at altitudes of 1500-1800 m. - Soviet Centr. Asia: Mtn. Turkm. (near N. Sulyukli, Prokhladnoe; Mt. Almandzhikh). Gen. distr. : occurrence possible in adjoining parts of Iran. Described from Centr. Kopet Dagh. Type in Leningrad. 2907 343 344 Note. A. turkmenorum Boriss. is one of the species of importance for gum exploitation, Secondary fall flowering of this species has been re- ported. Hybrids A. turkmenorum XA. piletocladus occur (Centr. Kopet Dagh). 307. A. densissimus Boriss. in Acta. Inst. Bot. Ac. Sec. URSS, ser. I. 37(1936)' 222; ibid, ‘sers Vj" (1938)"509)-Dracacantha densirssimya BoORISSey alice eo alice panier ay ie A pulvinate shrub, branched from base, the cushions up to 50 cm tall; spines approximate, rather slender, pointing upward at an acute angle, straight, 3-5 cm long, yellow; stipules at first pubescent, becoming gla- brous, acuminate, ca. 5mm long; leaflets (5) 6-7 pairs, glabrous or pu- bescent, oblong, 5-7 mm long, 1.5-2 mm broad; flowers in 2's in the leaf axil, crowded on the branches into a capitate or oblong inflorescence; bracts linear, shorter than the calyx, terminating in a prickle, up to 5 mm long, all or merely the upper ones appressed-shaggy; calyx glabrous merely at base, 5-7 mm long, the teeth shorter than the tube, densely covered with very long spreading hairs up to 3 mm long; corolla violet; standard ca. 10 mm long, the limb half as long as the claw, constricted at base, the cune- ate claw cuneate-based, the auricles broad, spreading; wing-petals equaling ~ the standard, the narrow oblong limb 2/3 the length of the claw; keel barely shorter than the wings, the oval limb half as long as the claw; ovary covered with long hairs; style naked, pubescent only at base; pods ovoid, white - hairy, ca. 4-5 mm long; seeds olivaceous, ca. 3 mm long, 2 mm broad, reniform. Fl. June-July; fr. July-September. Steppe slopes, at altitudes of 1500-2000 m.- Soviet Centr. Asia: Mtn. Turkm. (central part of Kopet Dagh range). Endemic. Possibly occurring in the adjoining parts of Iran. Described from Kopet Dagh range. Type in Leningrad. ‘Note. Hybrids occurring: 1) A. densissimus XA. piletocladus (Kara-gura range and at Mt. Massinev), 2) A. densissimus XA. longi- ramosus (Mt. Arvaz). Economic importance. A. densissimus is of practical impor- tance, even though this species is less suitable for exploitation than A. piletocladus on account of the pulvinate shape which renders the making of stem incisions difficult. 308. A. longiramosus Boriss. in Acta Inst. Bot. Ac. Sc. URSS ser. I, 3 (1936) "223; ibid., ser) Ve" 510.—=Tragacantha None amos Boriss., l.c. : A spiny shrub, up to 1m tall, branching from base, the branches long, slender; spines very long, up to 5-7 cm, slender, slightly curved, yellow; stipules ca. 8 mm long, acuminate, always glabrous, only when very young sometimes pubescent, ciliate-margined, stramineous; leaflets glabrous, at first pubescent, 5-6 pairs, linear, 5-10 mm long, 1-2 mm broad; flowers in 2's in the leaf axils, crowded in a capitate or oblong inflorescence; bracts linear, glabrate, with sparse pubescence confined to apex, about equaling the calyx; calyx 9-10 mm long, glabrous at base, long-hairy elsewhere, the teeth as long as the tube; corolla yellow; standard violet nerved, 12- 13 mm long, the claw 2.5 times the length of the limb, cuneately attenuate toward base, broad upward; wing-petals about equaling the standard, the claw twice the length of the oblong limb; keel ca. 10 mm long, the claw twice the length of the oval limb; ovary oblong, white-hairy; style ca. 8mm 258 347 long, naked, with hairs merely at base; pod oblong-ovoid, hairy; seeds 2mm long, 1.5 mm broad, reniform, olivaceous. Fl. June-August. Steppe slopes, at altitudes of 1600-2000 m.- Soviet Centr. Asia: Mtn. Turkm. (central and partly eastern part of W. Kopet Dagh). Endemic. De- scribed from Por-Dere. Type in Leningrad. 309. A. consentaneus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).-Tragacantha consentanea Boriss., l.c. A densely branched shrub up to 50 cm tall, with gray bark; branches stout, short, densely spiny; spines erect, 3-4 cm long, stoutish, pubescent; stipules ovate, ca. 7 mm long, short-acuminate, at first pubescent, be- coming glabrous; leaflets 7-8 pairs, ovate or oblong, 5-6 (8) mm long, 2- 2.5 mm broad, covered on both sides with spreading hairs, less densely so to glabrate above, distinctly veined, often carinate-conduplicate, mucro- nate; flowers in 2's in the leaf axils, crowded in ovaloid inflorescence 1- 1.5 cm in diam. ; bracts ca. 5 mm long, scarious, point-tipped, glabrous merely at base, white-pubescent elsewhere, linear, navicular, ca. 2mm broad; calyx ca. 8 mm long, covered throughout with long white hairs ex- cept for the glabrous conic base, the teeth equaling the rupturing tube; co- rolla exserted from calyx, yellow (in dry condition); standard 11-12 mm long, the orbicular limb 4 mm broad, obtusely truncate, the broadly cuneate claw twice as long as the limb, 5 mm broad in upper part, 1 mm at base; wing-petals equaling the standard, the oblong limb half as long as the claw; enlarged and auriculate at base; keel 10 mm long, the oval limb 4 mm long; ovary ovoid, white-hairy; style naked, 8 mm long; pod ovoid, white-hairy, ca. 5mm long; seeds reniform, ca. 2-2.5 mm broad, ca. 3 mm long, oliva- ceous, black-punctate, smooth. Fl. andfr. July. Slopes. - Soviet Centr. Asia: Mtn. Turkm. (W. Kopet Dagh, at the Iran border, in Karakalli district; southern slope to river Sumbar, between Koina-Kosyr and Por-dere canyon. Gen. distr.: likely in Iran. Described from river Sumbar. Type in Leningrad. Series 4. DISSECTI Boriss. - Calyx 6-9 mm long, splitting down to base; corolla 9-13 mm long; inflorescence (1.5) 2-2.5 cm broad, commonly spheri- cal or ovaloid; leaflets spatulate, oblong or oblong-lanceolate, subobtuse, terminating in a short prickle. 310. A. chionocalyx (Nevski) Boriss. comb. nova.-Tragacantha chionocalyx Nevski in Fl. Tadshikist. V (1937) 492 et 681; B. Fedtsch. et N. Ivan, in Tr. Tadzhik. bazy Akad. Nauk II (1936) 148-149, pro parte. - Ic: Fl. Tadzhik. l.c. 60, 4. A branched shrub; branches stout, ca. 0.7 cm in diam., densely beset with spines and stipules; spines slender, erect, inflorescence divaricate, 3-4 cm long; stipules oblong-ovate, ca. 1 cm long, connate to 1/3, finely pointed, at first pannose with long spreading white hairs, at length sparsely hairy to glabrate, ciliate-margined; leaves approximate in tufts at the end of the current year's shoots; young leaf rachises pannose; leaflets 6-7 (11) pairs, 5-10 mm long, 1-2 mm broad, oblong to lance-oblong, hairy on both sides; flowers in 3's in the leaf axils, crowded in a rounded inflorescence 2-2.5 cm in diam.; bracts linear, ca. 8 mm long, about equaling the calyx, glabrous, with tufts of white hairs at apex; calyx obconic, ca. 9 mm long, lanate with long white hairs, glabrous at the acute base, splitting nearly down to base, the teeth subulate; corolla ochroleucous; standard violet - 259 Ag wh YJ = eS ye SE wy a PR = es. Boh iS i % £7 gS SS Nee 2s Sy s= SAP. Yes a . = Zam LAL —<— Ss S Sg, PLATE XXIV 1. Astragalus caucasicus Pall.—2. A. bactrianus Fisch.— 3. A. prominens Boriss. 837, Ll 260 348 nerved, 12-13 mm long, rounded, and ca. 3.5 mm broad in upper part, 4 mm broad about the middle, with rectangular auricles, the lower one- third cuneately attenuate at base into the narrow claw; wing-petals some- what shorter than the standard, the limb ca. 1 mm broad, half as long as the claw; keel about equaling the wings, the limb half as long as the claw; ovary white-pannose; style hairy merely at base. June. (Plate XXIII, Figure 1). Stony and gravelly mountain slopes at altitudes of 1250-1500 m, in the upper part of the ephemeroid vegetation zone. - Soviet Centr. Asia: Pam.-Al. (Hissar and Kugitang ranges). Endemic. Described from Kugitang range. Type in Leningrad. 311. A. kuhitangi (Nevski) Boriss. comb. nova.-Tragacantha ku- hitangi Nevski in Acta Inst. Bot. Acad. Sc. URSS, ser. I, 4 (1937) 258. A branched shrub, 16-25 (30) cm tall; branches densely beset withspines and old stipules; spines straight, spreading or erect, brownish, 2-3 (3.5)cm long; stipules stramineous, subulate-tipped, ca. 1 cm long, faintly ciliate at apex and on the margin, white-hairy and enlarged at base, more densely hairy when young; leaves approximate in clusters at the ends of the current year's shoots, 6-7 pairs, later deciduous; young leaf rachises pubescent; leaflets 6-9 mm long, 1.25-1.75 mm broad, terminating in a prickle 0.25mm long, canescent with soft hairs on both sides; flowers in 3's in the leaf axils, crowded in a spherical capitate inflorescence 2-2.5 cm in diam.; bracts ca. 8 mm long, oblong-linear, glabrous except for the white-hairy apex; calyx obconic, 8 mm long, splitting nearly down to base, glabrate at base, white -lanate throughout; corolla 12-13 mm long; standard pandurate, con- stricted at the middle, rounded-ovate and 3.5-4 mm broad in upper one- third, with round auricles, narrowly cuneate -attenuate and 1-1.5 mm broad in lower one-third, cream-colored violet-nerved; wing-petals yellowish, somewhat shorter than the standard, the limb ca. 1 mm broad, half as long as the claw; keel ca. 10 cm long, the limb half the length of the claw; anthers ca. 0.5 mm long; ovary densely white-hairy; style hairy merely at base. July. Near the lower limit of the juniper belt, among steppe associations. - Soviet Centr. Asia: Pam.-Al. (Kugitang range, opposite village of Khodza- fil'ata). Endemic. Described from Kugitang range, Type in Leningrad. Note. A species very closely akin to A. chionocalyx (Nevski) Boriss., from which it differs only in the smaller number of leaflet pairs and a somewhat different shape of standard. 312. A. proximus Boriss. nom. nov.-A. dissectus B. Fedtsch. et N. Ivan. in Tr. Tadzhik. bazy Akad. Nauk, II (1936) 148-149, pro parte. - Tragacantha proxima Boriss. in Fl. Tadzhik. V (1937) 492 et 682, tab. 60, 3. A branched shrub with gray bark; branches stout, 0.5-0.6 cm in diam., white-hairy, densely beset with spines and stipules; spines slender, di- vergent, 2-3 cm long; stipules ovate, ca. 1 cm long, acuminate, connate and adnate to petiole, at first white-pannose, at length glabrous above, coriaceous, light yellowish; leaf rachises pannose -pubescent, becoming glabrous, terminating in a yellowish naked prickle; leaflets 5-7 pairs, ob- long-lanceolate, 0.5-0.6 cm long, 1-2 mm broad, acute, prickle-tipped, at first covered with whitish short spreading hairs, at length glaucescent, sparsely hairy on both sides (more densely beneath) or glabrous; flowers 261 349 350 in 2's-3's in the leaf axils, crowded in a spherical or oblong inflorescence 1-2 cm long and 2 cm broad; bracts lanceolate or linear, somewhat shorter than the calyx, navicular, acute, membranous, with long white hairs at apex; calyx 6-8 mm long, densely white-hairy, the obconic tube glabrous at base, the subulate teeth as long as the tube; densely covered to the tip with white hairs; corolla ochroleucous; standard mostly with darker veins in upper part, 9-10 (11) mm long, rounded and ca. 2,5 mm broadin upper part, with rectangular acute auricles about the middle, the claw cuneate, half as long as the limb, navicular; wing-petals and keel about equaling the standard, 9-10 mm long, fused with staminal tube, the limb half as long as the claw; ovary oblong, shaggy with silky hairs; style pubescent merely at base; pods oblong-ellipsoid, white-pannose, ca. 5 mm long; seeds rounded-ovaloid, 3 mm long, 2 mm broad, smooth, light brown. Fl. June. Gritty and stony slopes, in rock crevices, at altitudes of 1500-1800 m, in the upper part of the ephemeroid vegetation and rosarium zone. - Soviet Centr. Asia: Pam.-Al. (Peter I range, Kuraminskii, Gazimailik and Sarsa- ryak ranges). Endemic. Described from Pamir-Alai. Type in Leningrad. 313. A. devius Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).- Tragacantha devia Boriss., l.c.-Ic. A spiny, densely branched shrub, with gray bark; branches pannose- hairy, covered with persistent spinescent leaf rachises and stipules adnate to their base; spines 3-4 cm long, pannose at first, at length less hairy, the terminal point equaling or shorter than the uppermost leaflets; stip- ules ca. 10 mm long, long-acuminate, ovate at base, densely white-pannose at first, at length merely white hairy at base and on the back, membranous, stramineous; leaf petioles very short; leaflets (8) 9-11 pairs, approximate, ovate to oblong or lanceolate, 5~11 mm long, ca. 2-2.5 mm broad, cine- reous on both sides by curly spreading dense hairs, somewhat enlarged in upper part, terminating in short prickle; flowers in 2's-3's in the leaf axils, crowded in a capitate inflorescence ca. 1.5 cm broad; bracts linear, less than 1 mmbroad, ca. 8 mm long, navicular, curved, white -hairy, sometimes glabrous at base, subulate-tipped; calyx 9 mm long, splitting, white-hairy, acute and glabrous at base, the hairy teeth equaling the tube; corolla 10mm long, violet, the keel yellow; standard 10 mm long, pandurate, the limb violet-nerved, 3-3.5 mm broad, rounded in upper part, enlarged about the middle to 4mm, with obtuse round auricles, attenuate at base to the claw, this ca. 4 mm long and 1-1.5 mm broad; wing-petals equaling the standard, the lanceolate limb 4 mm long and less than 1 mmbroad, auriculate at base, 2/3 as long as the claw; keel 9 mm long, the ovate limb 3 mm long; ovary white hairy, oblong; style hairy only on the lower 1/3, ca. 5mm long. June. Limestone rocks. - Soviet Centr. Asia: Pam.-Al. (S.W., Babatag Moun- tains, surroundings of village of Dzhida-bulak). Endemic. Described from Babatag Mountains. Type in Tashkent. 314. A. munitus Boriss. nomen nov.-A. dissectus B. Fedtsch. et N. Ivan. in Tr. Tadzhik bazy Akad. ‘Nauk II (1936) 148-149, pro parte. - Tragacantha tenuispina Boriss. in Fl. Tadzhik. V (1937) 492 et 682, tab. 59, non A. tenuispinus Bge. A branched shrub; branches stout, 0.6-0.8 cm in diam., densely beset with spines and old stipules turned brown; spines slender, curved, 3-5 cm long, divergent at an angle of 30-45°; stipules oblong-ovate, ca. 1.5cm long, membranous, stramineous, ciliate-margined; spines and stipules at first 262 (351 covered with scattered curly sparse hairs, becoming glabrous; leaves ap- proximate in clusters at the ends of young shoots; leaflets 6-7 pairs, spat- ulate-oblong, 7-9 mm long, 2-2.5 mmbroad (in upper part), subobtuse, prickle-tipped, green, glabrous above, prominently veined, covered beneath with scattered spreading hairs, more densely so when young; flowers in 2's in the leaf axils, crowded in an ovaloid-rounded compact inflorescence 1.5- 2.5 cm long and 1.5-2 cm broad, borne at the base of the current year's shoots; bracts linear, ca. 0.5 cm long, navicular, shorter than the calyx, whitish, glabrous in lower part, white-shaggy upward; calyx ca. 7 mm long, white-lanate with long hairs, acute and glabrous at base, splitting nearly down to base, the teeth subulate; corolla yellow; standard ca. 10 mm long, rounded in upper one-third, 2.5 mm broad, 3 mm broad about the middle, with rectangular auricles in the lower one-third, cuneately attenuate toward base into a narrow claw; wing-petals equaling the standard, the limb ca. l1mmbroad, half as long as the claw; keel somewhat shorter than the wings, the limb half as long as the claw; ovary white-pannose, style filiform, pu- bescent at base. Fl. July. Limestone rocks in the woodland and scrub formation. - Soviet Centr. Asia: Pam.-Al. (Kyzyl-su and Yakh-su pass, western slopes of Karimonok mountains in upper reaches of Darvaz-Pishau river). Endemic. Described from Karimonok Mountains. Type in Leningrad. Series 5. ARNACANTHI Boriss. Calyx 7-9 mm long, splitting down to base; corolla 12-19 mm long; inflorescence 1.5-2 cm broad, rounded; leaf- lets long-acuminate and prickle-tipped, linear. 315. A. insidiosus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).-Tragacantha insidiosa Boriss., l.c.-Ic.: l.c.f. A densely branched shrub; branches rather short, densely clothed in spines and stipules; spines covered with spreading hairs, slightly curved, divaricate, 2-3.5 cm long; stipules 14-15 mm long, filiformly long-acumi- nate from an oval base, glabrous, the margin long-ciliate at base; leaves terminating in a spine much shorter than uppermost leaflets, 2-5 mm long; leaflets linear, 8-16 mm long, 1-2 mm broad, long-acuminate to a prickle, bluish on both sides with appressed pubescence, (3) 4 pairs, often condu- plicate; flowers in 3's in the leaf axils, crowded in a capitate inflorescence 2-2.5 cm broad, borne at the base of the current year's shoots; bracts ca. 8 mm long, linear, less than 1 mm broad, glabrous, membranous, straight; calyx 11 mm long, splitting down to base, white-hairy throughout, with shorter hairs at base, the filiform teeth about as long as the tube; corolla drying yellow; standard 14-15 mm long, the limb 4.5 mm broad in upper part, constricted at the middle to 2 mm, auriculate and enlarged at base to 5 mm, the claw narrowly cuneate, 5 mm long and ca. 1 mm broad at base; wing-petals 12 mm long, the claw about 1.5 times as long as the narrowly oblong limb, ca. 1 mm broad; keel fused to 3 mm with wing-claws and the staminal tube, ca. 13 mm long, the claw 1.5 times as long as the ovate limb; ovary oblong, hairy; style 7 mm long, hairy up to the middle. Fl. July. Dry stony slopes.- Caucasus: S. Transc. (toward north of village of Ur- mir). Endemic. Described from the indicated locality. Type in Baku. Note. Relatedto A. arnacanthoides Boriss., from which it is distinguished by the somewhat larger stipules, more numerous pairs of leaflets, larger stipules, calyx 11 mm long (as against 8-9 mm), larger standard 14 mm long (as against 12 mm). 263 a2 393 316. A. arnacantha M.B. Fl. taur.-cauc. II (1808) 205, pro parte III, 500; Edb. Fl. Ross.)1/9640); pro, parte:.DE.. Prody lh «29, pRo. parte: ecer Astrageygeront: Il) (1353 Wisch;) Synaimag.e Ose Schmalh.eky.. 1, 26a. Poterium Pall. Astrag. (1800) I, non Vahl (Symb. I, 63) in Willd. Mém. sur les diverses espéces de Tragacanthe (1794) 6.-A. Tragacantha Habl. taur. (1789) 112.-A. criacantha Stev. ex Fisch. Syn. Trag. (1853) 18.-A. horridus Willd. herb.-Tragacantha arnacantha Stev. in Bull. Soc. Nat. Mosc. IV (1832) 269, pro parte; O. Ktze. Rev. gen. 943. - T. criacantha Stev. in Bull. Soc: Nat. Mose. XXIX (1856) 257: -Iec.- Pall. loca tabs 1s) Fisch dic.) tabr Bafind;dsoriss, iny Not. (Syst. Insta Bor Ac.w Seb VEY LOW 94 O)e1 Bly fa. A densely branched shrub, 30-50 cm tall; branches short, crowded, stout, ca. 7mm in diam., 3-8 cm long; internodes lanate, densely clothed in spinescent leaf rachises and stipules; spines divergent, theuppermosterect, 2.9-3 mm long, pubescent, the terminal spine much shorter than the upper - most leaflets; stipules lanceolate, long-acuminate from an ovate base, up to 18 mm long, pannose-shaggy on the back, long-ciliate on the margin; leaflets lanceolate, rarely narrowly oblong, cinereous on both sides with spreading hairs, rarely white-pannose, prickle-tipped, 4-5 pairs, (8) 10- 16 mm long, 2-3 mm broad, approximate, often conduplicate, erect; flow- ers sessile in 2's in the leaf axils, crowded in an oblong or capitate inflor - escence ca. 1.5-2 cm in diam., borne at the base of current year's shoots; bracts equaling the calyx, coriaceous, linear, ca. 2 mm broad, curved, navicular, covered with long hairs at apex and on the back; calyx white by long dense hairs, glabrous at base, 7-8 (9) mm long, the linear -filiform teeth equaling the splitting tube; corolla pink or white with pink nerves; standard 14-16 (18) mm long, pandurate, the oblong limb 10 mm long, 4- 5 mm broad in upper part, constricted at the middle to 4 mm, enlarged at base to 5-7 mm, the auricles acutish spreading, the narrow claw ca.1mm broad and 4-5 mm long; wing-petals 13-15 mm long, the linear limb 6-7mm long, 1-1.5 mmbroad, auriculate at base; keel somewhat shorter than the standard, the limb oblong-elliptic, convex on the back, 6-7 mm long, 2- 3 mm broad, the upper part gradually attenuate toward base; ovary oblong, covered with long white hairs; style ca. 10 mm long, naked, hairy merely at base, curved at the tip; pods 2 mm long, ovaloid or oblong, often 1- seeded, densely lanate or white-pannose. Fl. June-August; fr. July. Stony slopes of mountains and hills. - European part: Crimea (from Bala- klava to Koktebel and Feodosiya); _? Caucasus (Sredinskii's herbarium). Endemic. Described from Crimea. Type in Leningrad. Note. Reports on the occurrence of A. arnacantha inthe Caucasus refer to A. arnacanthoides Boriss. which is distributed in N.W. Cau- casus. In the herbarium of the Botanical Institute of the Academy of Sciences from the Caucasus there are still two specimens of typical T. [?] arna- cantha, and this apparently accounts for the confusion of labels. 317. A. arnacanthoides Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS VIII, 10 (1940) 180.-A. arnacantha auct. non M.B.-Tragacantha arnacanthoides Boriss., l.c., f.1.-Exs.: Herb. Fl. Cauc. No. 423. A densely branched shrub, forming cushions to 1 m in diam., branches short, crowded, 5-7 mm in diam., densely clothed in stipules and spines; spines arched-recurved or fairly straight, 2-3.5 cm long, at first sparsely shaggy-pubescent; internodes pannose; stipules 12-13 mm long, subulately long-acuminate from an oval base, glabrous, long-ciliate on the margin; 264 04 terminal spine of leaf shorter than the terminal leaflet; leaflets lanceolate, 6-12 mm long, 1-2 mm broad, villous-hirsute on both sides, bluish, prickle- tipped, 5-7 pairs, conduplicate; flowers in 3's in the leaf axil, crowded in a capitate inflorescence ca. 2 cm in diam., borne at the base of the current year's shoots; bracts 5-9 mm long, linear, ca. 1 mm broad, glabrous at base, white-lanate in upper part, yellow, membranous, navicular, con- duplicate; calyx 8-9 mm long, white-lanate, puberulent at base, pointed, the white -hairy teeth about equaling the tube; corolla drying yellow, the standard violet-nerved; standard 12 mm long, the limb 7 mm long, round- tipped, 3-4 mm broad, constricted at the middle to 2 mm, enlarged toward base to 4-5 mm with obtuse auricles, the claw 5 mm long, narrow, ca.1mm broad; wing-petals equaling the standard, the oblong limb 4 mm long, 1mm broad, the claw twice as long as the limb; keel about equaling the wings and the standard, the ovate limb ca. 3 mm long and 1.5 mm broad, the claw 2.5 times as long as the limb; ovary oblong, hairy; style 6 mm long, naked nearly to the middle; pods hairy, ovoid, ca. 5mm long. Fl. May-June; fr. July-August. Wooded mountain slopes, on marle limestone and chalk, and on seaside precipices.- Caucasus: W. Transc. (near Novorossiisk, Gelendzhik, Sukko, Tuapkhash range, Abrau, Markotkh range). Endemic. Described from the surroundings of Novorossiisk. Type in Leningrad. Series 6. INDIVISI Boriss. - Calyx 4-10 mm long, not splitting down to base, with distinct tube; corolla 10-16 mm long; inflorescence elongate, spiciform, 5-12 mm broad. 318. A. paliurus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946) f.-Tragacantha paliurus Boriss., l.c.-Exs.: HFR No. 1413 (sub Astragalo bactriano Fisch). A branched shrub; branches spiny; spines subhorizontally spreading, at first pubescent becoming glabrous, 1.5-5 cm long; stipules oblong-lanceo- late, ca. 1 cm long, hairy on young current year's shoots, at length gla- brous; leaflets 6-8 pairs, oblong-lanceolate to oblong-ovate, 5-10 mm long, 1-2 mm broad, carinate-folded, cineraceous on both sides with short shaggy pubescence, more densely pubescent when young, prickly-tipped or acute; flowers in 2's in the leaf axils, forming an oblong inflorescence 2-6 cm long, 1.5-2 cm broad; bracts oblong or oblong-lanceolate, 7-8 mm long, folded ca. 2-3 mm broad, clasping the calyx, navicular, membranous, yellow, sparsely white -hairy on the margin and on the back; calyx 8-9 mm long, white-hairy, glabrous at base, not splitting, the shaggy lanceolate teeth ca. 2 mm long; corolla drying yellow; standard 15-16 mm long, 5-7 mm broad in upper part, almost imperceptibly constricted at the middle, the ovate claw 10 mm long, ca. 6 mm broad; wing-petals 12-15 mm long, the oblong limb ca. 1.5 mm broad, 2/3 the length of the claw; keel 12-14 mm long, the oval limb 2/3 the length of the claw; ovary oblong, white-hairy; style ca. 10 mm long, curved at base and at the tip, hairy up to the middle. July. Mountains, at the altitude of 1100-1700 m.-Soviet Centr. Asia: Pam.-Al. (Zeravshan range, Nura-tau range, Koi-tash mountain). Endemic. Described from Takhta-Karagin mountain pass in Zeravshan range. Type in Leningrad. 319. A. caucasicus Pall. Astrag. (1800) 2; Willd. Sp. pl. Il, 1331; M.B. Fl. taur.-cauc. II. 205 (pro parte); Boiss. Fl. or. II, 327; Fisch. Syn. Trag. 26; Bge. Astrag. geront. II, 131; Ldb. Fl. Ross. I. 639; DC. 265 Prodr. II, 296; Grossh. Fl. Kavk. Il, 306.-Tragacantha caucasica Stev. in Bull. Soc. Nat. Mosc. IV (1832) 269; O. Ktze. Rev. gen. (1891) 9435 — loa alll lace amtabercselischeplaiehs talons Crea 0s sB Sens leone eace No. 389. A densely branched shrub, 40-50 cm tall; branches up to 20 cm long, with tomentose internodes, densely clothed in stipules and spinescent leaf rachises; spines divergent, pubescent, slender, 2-7 cm long; stipules lanceolate, acuminate, pubescent at first, becoming glabrous, stramineous, subcoriaceous-membranous; leaflets 6-8 pairs, linear-lanceolate to oblong, cineraceous with appressed hairs, glabrate above, 7-10 mm long, to 2mm broad, terminating in a strong prickle; bracts narrow, linear-navicular, acuminate, hairy on the back, about equaling or somewhat shorter than the calyx; flowers in 2's or 3's in the leaf axils, crowded in a cylindric spici- form inflorescence 1-2 cm broad; calyx 7-10mm long, lanate with white hairs, not splitting, with a distinct tube, commonly glabrous toward base, the pannose -hirsute lanceolate or subulate teeth about equaling or somewhat shorter than the tube, corolla about twice the length of the calyx, 12-14 ¢ (16) mm long, whitish or yellowish; standard oblong-spatulate, 4-6 mm broad in upper part, retuse, gradually attenuate towardbase, the claw broad, but narrower than the limb; wing-petals about equaling the standard, the oblong-lanceolate limb about equaling the claw, auriculate at base; keel- limb oval, 2/3 as long as the claw; ovary hairy, oblong; style 8 mm long, naked; pods lanate, oblong, 1-2-seeded. June-September. (Plate XXIV, Figure 1). Rocks and stony dry slopes. - Caucasus: Dag. (Chir-yurt, Derbent), Cisc. (mountains of Mozdok), E. and S. Transc. Gen. distr. : Arm.-Kurd. (Kars). Described from Terek. Type in London. 355 320. A. caspicus M.B.Fl. taur.-cauc. II (1808) 204; III, 500.-A. cas- pius M.B. ex Bge. Astrag. geront. II (1869) 132; Ldb. Fl. Ross. I, 641; Boiss. Fl. or. Il, 334; Grossh. Fl. Kavk. I, 307.-A. cretensis Pall. Astrag. (1800) 5, No. 6, excl. syn.-A. leptodendron Fisch. Syn. trag. (1853) 345.-A. Echinus C. A.M. Enum. (1831) No. 1266, non DC.-Tra- gacantha caspica Stev. in Bull. Soc. Nat. Mosc. IV (1832) 269; O. Ktze. Rev. gen. 941.-Ic.: Fisch, l.c., tab. C, f. 23, 28. A densely branched pulvinate shrub; branches divergent, slender, pan- nose, elongated, spiny; spines divergent, straight, strong, (1.5) 2-3 (5)cm long, often distant on elongated branches; stipules ovate-lanceolate, ciliate at base, glabrous or pubescent, thinly pubescent when young; leaflets (3) 4-8 pairs, oblong-lanceolate, subacute, mucronulate, hairy, greenish gla- brate above, sparsely appressed-pubescent beneath, small, 3-8 mm long, 1-2 mm broad; bracts equaling the calyx, linear, navicular, hairy or gla- brous at apex, ciliate-margined; flowers in 2's in the leaf axils, crowded in an oblong-cylindric spiciform inflorescence 8-10mmbroad; calyx4-6 mm long, white-lanate, glabrous at base, with a distinct tube, not splitting, the glabrous-tipped subulate-lanceolate subequal teeth shorter than the tube; corolla ochroleucous, about twice the length of the calyx, 10-11 mm long; standard oblong, obtuse, violet-nerved, retuse, constricted at the middle, the claw equaling to twice as long and sometimes broader than the limb, convex-margined; wing-petals somewhat shorter than the standard, the claw 1.5-2 times the length of the limb; keel equaling the wings, the claw about twice the length of the oval limb; ovary oblong, hairy; style ca. 10 mmlong, 266 356 naked; pods whitish-cineraceous with dense hairs, included in the calyx; seeds sometimes 2, reniform, ca. 2mm long. June-July. Stony and sandy dry mountain slopes, at altitudes between 500 and1500m, sometimes on compacted saline shell-rock at the sea shore. - Caucasus: Dag. (rare), E. Transc., S. Transc. (Nakhichevan! in vicinity of Bichenakh), Tal. Gen. distr.; Iran. Described from Shirvanskie mountains. Type in Leningrad. 321. A. oltensis Grossh. Fl. Kavk. II (1930) 306.-Tragacantha oltensis Boriss. comb. in manuscr. s A diminutive shrub, 10-20 cm tall; branches glabrate or sparsely pu- bescent, slender, 2-3 mm in diam., clothed in imbricated stipules and spinescent leaf rachises; spines slender, (1) 1.5-3 cm long, at first covered on the back with spreading hairs, at length glabrous, ciliate-margined, slightly divergent; stipules ca. 7 mm long, connate to 3-4 mm, membra- nous, the free part triangular-acuminate from an ovate base; leaflets (3) 4-5 (7) pairs, conduplicate, canescent, ca. 5mm long, 1-1.5 mm broad; flowers in 2's in the leaf axils, crowded in a spiciform inflorescence 5- 7 mm broad; bracts small, 3-4 mm long, acute, linear, navicular in upper part, the margin beset with white hairs; calyx 4-5 mm long, glabrous to the middle, the upper part covered with curly white hairs, the distinct tube 4-5 times as long as the lanceolate teeth; corolla exceeding the calyx, 8 (10) mm long, pale violet, with a violet spot on the keel; standard 8 (10) mm long, 3 mm broad in upper part, gradually attenuate into a broad obovate convex-margined claw; wing-petals 7-8 mm long, the limb less than 1 mm broad, half as long as the claw; keel 6-7 mm long, the ovate limb half as long as the narrow claw; ovary oblong, hairy; style 4 mm long, naked. June. Taluses. - Possibly growing in Transcaucasia. Beyond border of former Ol'tin settlement in neighborhood of Bardus (Sosnovskii). Described from former Ol'tin settlement. Type in Baku. Series 7. BRACTEATI Boriss.-Bracts large, 6-12 mm broad, ovate, covering the calyx; calyx 12-20 mm long, very slightly shorter to longer than the corolla; standard 17-22 mm long; flowers in 2's. 322. A. transoxanus Fisch. in Bull. Soc. Nat. Mosc. XXVI (1853) 338; Bge. Astrag. geront. I, 82; I] 142; Bunge, Astrag. Turk, 250; Boiss. F1. Or. Il, 326.-Tragacantha transoxana O. Ktze. Rev. gen. (1891) 948; BORISS. ln hi whadzhik., Vi, 488. —le.s.Buisch. lae., tab.) Buf. 183 hl, tad-— Zhik. I. c., tab. 60. A branched, densely leafy shrub; branches firm, lanate-pannose when young, older densely clothed in spines 4-5 cm long; stipules lance-oval to lanceolate, ca. 2 cm long, acute, coriaceous, light yellowish, adnate to petiole to above the middle, lanate at base; leaf rachises firm, pannose, terminating in a glabrous yellowish long spine exceeding the leaflet; leaflets 5-7 pairs, oblong to oblong-elliptic, 10-11 mm long, 2.5-3.5 mm broad, subacute, prickle-tipped, whitish-pannose on both sides; flowers in 2's in the leaf axils, crowded in an ovaloid-spherical or cylindric inflorescence 4-6 cm long and ca. 2.5 cm broad; bracts ovate, conduplicate, ca. 5-6mm broad, expanded, obliquely truncate, somewhat shorter than the calyx, navicular, membranous, yellowish, glabrous, ciliate-margined; calyx 12- 15 mm long, densely white-shaggy except for the lower half of the tube, the triangular -subulate teeth half as long to as long as the tube; corolla 267 357 358 white; standard 17-22 mm long, straight, the limb oblong-oval, somewhat constricted at the middle, navicular, ca. 4mm broad in upper part, slightly retuse, about as broad as the claw; wings and keel about equaling the stand- ard, their limb half the length of the claw, scarcely fused; ovary white- pannose, oblong; style hairy from base to the middle. June. Stony and gravelly south slopes, at the altitude of about 650m. - Soviet Centr. Asia: Pam.-Al. (Zeravshan range, Karatau mountains). Endemic. Described from Karatau mountains in Zeravshan. Type in Leningrad. 323. A. prominens Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).-Tragacantha prominens Boriss. l.c.-Ic.: l.c.f. A branched shrub; branches long, stoutish, densely clothed in spinescent leaf rachises and stipules; spines somewhat divergent, covered with spread- ing hairs, 4-6 cm long; stipules ca. 12 mm long, lanceolate, acutish but not pointed, stramineous, membranous, covered with short hairs merely on the back, long-ciliate on the margin at apex, glabrous elsewhere; leaves ter- minating in a long spine exceeding the terminal leaflet; leaflets 6-7 pairs, oblong to oblong-obovate, 8-12 mm long, 3-4 mm broad, bluish, glabrous above, short-hairy beneath, terminating in a short prickle, carinate; flowers in 2'sin the leaf axils, crowded in a spiciform inflorescence 6-7 cm long, ca. 2.5 cm broad; bracts firmly enclosing the calyx-tube on all sides, oh- liquely truncate at apex, rounded at base, glabrous, long-ciliate on the margin, conduplicate, ca. 12 mm broad (6 mm each flank), 12 mm long, carinate dorsally, the unfolded face turned toward the axillary twin flowers; calyx 20 mm long, covered with spreading long hairs, the filiform partially glabrous acute teeth equaling the tube; corolla drying yellow; standard 19mm long, the limb 6 mm broad in upper part, attenuate toward and grad- ually passing into the claw, this 4 mm broad in upper part, 2 mm at base; wing -petals equaling the standard, the claw 1.5 times as long as the oblong limb; keel-limb ovate, round-tipped, the claw 1.5 times as long as the limb; ovary white-pannose; style 14 mm long, the lower two-thirds hairy; pod ca. 6 mm long, oblong, white-hairy, unilocular. July-September. (Plate XXIV, Figure 3). | Mountain slopes, stony and gravelly places, in witchgrass-dominated rosaria and giantfennel associations; also hay plant associations in the up- per part of the woodland and scrub formation. - Soviet Centr. Asia: Pam.- Al. Endemic. Described from W. Karategin. Type in Leningrad. Section 54. POLYANTHOS Boriss.-Macrocalycina Fisch. in Bull. Soc. Nat. Mosc. XXVI (1853) 42, p.p.-Flowers (3) 4-15 in the leaf axils, on very short stout densely hairy peduncles not exceeding 1-4 mm; calyx 8-25 mm long, mostly above 15 mm. Shrubs. Series 1. INDIVISI Boriss.- Flowers (3) 4-9 in the leaf axil; calyx 8- 14mm long, with non-splitting tube, hairy throughout, much shorter than the corolla; corolla 12-15 mm long; claw of standard enlarged; inflores- cence oblong or rounded-oval, 1.5-3 cm broad. Species of Soviet Central Asia. 324. A. asaphes Bge. Astrag. Turk. (1880) 251.-Tragacantha asaphes Boriss. in Fl. Tadzhik. V (1937) 488, tab. 58, 4. A densely branched shrub; branches straight, short, densely spiny; spines slender, 2-3 (4) cm long; stipules lanceolate, ca. 1 cm long, acute, 268 59 membranous, faintly ciliate-margined at first, becoming glabrous; leaflets 5-7 pairs, linear-oblong, 5-10 mm long, 1.5-2 mm broad, prickle-pointed, glabrous above, sparsely appressed-hairy beneath; flowers 3-4 in the leaf axils, crowded in an ovaloid-spherical inflorescence (1.5) 2-2.5 cm indiam.; bracts lanceolate, about equaling the calyx tube, stramineous, ciliate at the apex or glabrate; calyx ca. 12 mm long, shaggy with white spreading hairs, the filiform colored teeth about equaling the tube; standard 14-15 mm long, the limb 5 mm broad, retuse, slightly constricted at the middle, about as long as the claw, cuneate at base; wings and keel nearly free, somewhat shorter than the standard, the claw twice the length of the limb; ovary ob- long, white-shaggy; style hairy at base, sometimes up to the middle, ca. 9mm long. Fl. June-July. Gravelly mountain slopes at altitudes up to 2400 m, chiefly in the steppe belt and in juniper groves. - Soviet Centr. Asia: Pam.-Al. (Karatau range in Zeravshan, Kuramskii range), Tien Shan. (W.). Endemic. Described from Karatau range in Zeravshan. Type in Leningrad. 325. A. rubens B. Fedtsch. et N. Ivan. in Tr. Tadzhik. bazy Akad. Nauk II (1936) 146.-Tragacantha rubens Boriss. in Fl. Tadzhik. V (1937) 487, tab. 58. A branched shrub; branches stout, long, clothed in spines and stipules; spines 3-5 cm long, erect; stipules lanceolate, large, exceeding the calyx, acute, membranous, stramineous turning brown in age, glabrate, ciliate - margined, adnate to petiole to about the middle; leaves aggregated in tufts at the ends of branches, the rachises firm, spinescent, faintly pubescent; leaflets 4-5 pairs, lanceolate, ca. 10 mm long and 2 mm broad, acute and prickle-pointed, sparsely appressed-hairy beneath, glabrate above, prom- inently veined; flowers 4-5 in the leaf axils, approximate in a spherical inflorescence 2.5-3 cm long and ca. 3 cm broad; bracts elliptic-lanceolate, ca. 10 mm long, acute, navicular, shorter than the calyx, membranous, glabrous or faintly hairy, ciliate-margined; calyx ca. 14 mm long, split to the middle, white-shaggy, the subulate acute teeth colored at the tips, less hairy than the tube; standard ca. 15 mm long, barely overtopping the teeth, purple in upper part, guitar-shaped to suboblong, the limb not clear- ly differentiated from and equaling or shorter than the claw; keel and wings purple or whitish, shorter than the standard, fused at base with staminal tube; ovary white-pannose; style hairy up to about the middle; pod lance- oblong, ca. 7 mm long, indurated, covered with short white hairs; seeds reniform, ca. 4mm long, 2-2.5 mm broad, smooth, brownish, black- spotted. Fl. June-July. Stony slopes at altitudes of about 2600 m, in the upper zone of the wood- land and scrub formation. - Soviet Centr. Asia: Pam.-Al. (Hissar range, Darvaz). Endemic. Described from Pamir-Alai. Type in Leningrad. 326. A. bactrianus Fisch. in Bull. Soc. Nat. Mosc. XXVI (1853) 354. Boiss. Fl. or. II, 329; Bge. Astrag. geront. Il, 167; Bunge. Astrag. Turk. 251.-Tragacantha bactriana O. Ktze. Rev. Gen. (1891) 943; Boriss. BY Padzhik: WeN4s4 ick: Fisch!) ike} tabs DIMf: 38; Fly Tadzhik: V'(1937) tab. 60, 6. A branched shrub up to 1.5 m tall, sometimes pulvinate; branches densely clothed in spines; spines long, divergent, pubescent at first, be- coming glabrous; internodes of old branches densely pannose; stipules ovate- Janceolate, ca. 1 cm long, acuminate, at first hairy at base, becoming 269 360 glabrous, wrinkled-nervose; leaflets 6 pairs, obovate-lanceolate, ca. 7 mm long and 2-2.5 mm broad, prickle-pointed, coriaceous, hairy at first, bluish, turning greenish, appressed-hairy beneath, glabrate above, prom- inently veined; flowers 7-9 in the leaf axils, forming an oblong inflor- escence ca. 5 cm long and2.5 cm broad; bracts oblong-lanceolate, sub- navicular, shorter than the calyx, yellow, hairy at apex and ciliate-mar- gined; calyx ca. 8 mm long, the tube turbinate-cylindric, hairy downtobase, not splitting, the teeth linear-lanceolate, straight, acute, half as long to nearly as long as the tube, hairy; standard exceeding the calyx, 10-12 mm long, the limb obovate or elliptic, somewhat constricted at the middle, re- tuse, the claw somewhat narrower than the limb, gradually attenuate toward base; wing-petals barely shorter than the standard, the limb obtusely auri- culate, half as long as the claw; ovary hairy, oblong; style twice as long as the ovary, hairy merely at base; pods oblong-ovaloid, 4-5 mm long, in- durated, white-hairy; seeds reniform, ca. 3 mm long and 1.5 mm broad, smooth, olivacous, minutely black-spotted. Fl. May-August; fr. June- September. (Plate XXIV, Figure 2). Stony and gravelly mountain slopes, on clay and sandy soils, and on mot- tled rock and gypseous outcrops, at altitudes of 1000-1700m. - Soviet Centr. Asia: Tien Shan. (western part — Kuraminskii range, Pam.- Al. (Turkes- tan, Zeravshan and Hissar mountain ridges). Endemic. Described from Karatau range in Zeravshan. Type in Leningrad. Series 2. PAMIROALAICI Boriss.- Flowers 3-6 in leaf axils; calyx ca. 10 mm long, much shorter than the corolla, splitting down to base; corolla 15-19 mm long, with a rather broad claw [?]; claw of standard gradually cuneate-attenuate; inflorescence spherical, 3-5 cm broad. 327. A. Hilariae Boriss. comb. nova.-Tragacantha Hilariae Boriss. Fl. Tadzhik. V (1937) 493, 683, tab. 90, 2. A densely branched shrub; branches stoutish, ca. 10cm [sic] in diam., recurved, densely beset with spines and stipules; spines stout, recurved, 5-6 cm long, sometimes nearly horizontally spreading, becoming brown; leaves aggregated in tufts at the ends of young shoots; stipules acute, oblong, ca. 1 cm long, imbricated, stramineous, connate and adnate to petiole to 2/3, subfiliform-acuminate, glabrous except for hairs at base when young; leaflets 7-8 pairs, oblong, 1U-12 mm long, 2.5-3 mm broad, prickle-tipped, white-pannose on both sides, less densely so and prominent - ly veined above; flowers 3-4 in the leaf axils, crowded in a dense sub- spherical inflorescence ca. 3 cm in diam., borne at the ends of current year's branches; bracts ca. 1 cm long, linear, navicular, equaling to some- what exceeding the calyx, stramineous, glabrous in lower part, white- shaggy upward; calyx ca. 10 mm long, shorter than the corolla, lanate with long white hairs, glabrous at base, splitting nearly down to base, the teeth subulate; corolla yellow; standard ca. 15 mm long, 3 mm broad in upper part, oblong, round-tipped, enlarged below the middle, exauriculate, grad- ually attenuate toward base; wing-petals about equaling the standard, the limb just above 1 mm broad, half as long as the claw; keel somewhat shorter than the wings, the limb half the length of the claw; wings and keel fused at base with staminal tube; ovary white-shaggy; style shorter than the corolla, pubescent in lower one-third. July. 270 Stony mountain slopes. - Soviet Centr. Asia: Pam.-Al. (Darvaz, and Viskharvi and Vanch river valleys). Endemic. Described from Darvaz. Type in Leningrad. 328. A. Alexeenkoanus B. Fedtsch. et N. Ivan. in Tr. Tadzhik, Bazy Akad. Nauk II (1936) 147.- Tragacantha Alexeenkoana Boriss. in Fl. Tadzhik, V (1937) 494, tab. 60. A low shrub, densely branched from base, ca. 50 cm tall; branches stout, ca. 10 mm in diam., white-pannose, densely spiny; spines thick, recurved, spreading, 5-6 mm long, becoming brown, often reflexed, cur- rent year's shoots 20-30 cm long; stipules elliptic, ca. 1.5 cm long, acu- 361minate, stramineous, connate and adnate to petiole to 2/3, glabrous in upper part, pannose-pubescent at base (young stipules and leaf rachises pubescent throughout); leaflets (5) 8-9 pairs, lanceolate or linear-lanceolate, some- what enlarged upward, 10-13 mm long, 1.5-3.5 mm broad, acuminate, pu- bescent beneath, glabrate above (at first pubescent on both sides), green, prominently veined; flowers 5-6 in leaf axils, crowded in a spherical in- florescence 4-5 cm in diam.; stipules linear, equaling the calyx, 12 mm long, finely pointed, navicular, pannose-villous with white hairs, glabrate at apex and at the very base; calyx ca. 10 mm long, covered with long white hairs, splitting nearly down to base, the subulate teeth equaling the tube; corolla yellow (or white); standard 15-19 mm long, round-tipped, 3-4 mm broad in upper one-third, enlarged at the middle to 6 mm and auriculate, the auricles sometimes slightly reflexed, the lower part of the limb cuneate- attenuate toward base into a narrow claw; wing-petals equaling the standard, the limb ca. 1 mm broad, the claw 2.5 times the length of the standard; keel barely shorter than the wings, the limb 1.5-2 mm broad, the claw 2-3 times the length of the limb, fused at base with the wings and the staminal tube; ovary white-shaggy; style pubescent at base. August-September. Stony slopes, at altitudes 1400-2000 m.- Soviet Centr. Asia: Pam.-Al. (between villages Dzharf and Kevron). Endemic. Described from Pamir- Alai. Type in Leningrad. Series 3. IMBRICATI Boriss. - Flowers in 3's-5's; calyx 9-17 mm long, markedly shorter than the corolla; corolla 12-20 mm long; claw of standard narrow; inflorescence 2-4.5 cm long, rounded. (Caucasus). 329. A. Sommieri Freyn. in Bull. Bot. Ital. (1893) 526; Ej. in Osterr. Bot. Zeitschr. XLII, No. 12, 418; Grossh. Fl. Kavk. II, 309.-Traga- cantha Sommieri Boriss. in manuscr.-Ic.: Somm. et Lev. in A.H.P. XVI (1900) tab. XV. A low, densely branched shrub 8-20 cm tall; stems ca. 5-7 mm thick, pubescent, densely clothed in long spines and stipules; spines twisted, flexible, 14-15 cm long, glabrate; stipules coriaceous, ovate, acuminate, at first long hairy, at length covered with white hairs merely in the middle part, ciliate-margined, stramineous, 12-15 mm long; leaflets 8-10 (12) pairs, elliptic or ovate, 5-7 mm broad, (7) 14-15 mm long, subobtuse and slenderly mucronate, round-based, green, glabrous or the young sparsely hairy, prominently veined; flowers 5 in leaf axils, sessile, crowded in an ellipsoid-ovoid dense head ca. 4-4.5 cm long, 2.5-4 cmbroad; bracts linear, slightly splitting, pointed at base, densely white-shaggy with long white 362 hairs, 8-9 mm long, 1.2 mm broad, the hairy teeth half the length of the tube, subulate-tipped, concealed in dense wool; calyx 10 mm long; corolla 271 365 pink, drying yellow to nearly white; standard 18-20 mm long, the oblong scarcely retuse limb 10 mm long, 5 mm broad, auriculate at base (the auricles acute reflexed), faintly constricted at the middle, attenuate into a narrow cuneate claw ca. 8 mm long; wing-petals somewhat shorter than the standard, the oblong-lanceolate limb half as long as the claw; keel shorter than the standard, ca. 16 mm long, the claw 3 times as long as the standard; stamens fused to high up with wings and keel; ovary ovoid, dense- ly hairy, glabrous at base; style naked, 16 mm long; pods shaggy, ca. 5mm long, unilocular; seeds reniform. May-July. Dry sunlit and stony slopes, rocks, and margins of pine forests. -Cau- casus: W. Transc. (between Batumi and Akhaltsikh, between villages Keda and Khela; near Khulo junction; near rural Charbiet). Gen. distr.: Arm. - Kurd. (former Artvin village). Described from Adzharia. Type in Leningrad. 330. A. transcaucasicus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).-Tragacantha transcaucasica Boriss., l.c. A densely-branched shrub; branches 4-5 mm in diam., densely clothed in remnants of old stipules and persistent spinescent leaf rachises; spines long, 4-8 cm, slender, somewhat twisted, glabrous, sparsely pubescent when young; stipules compact, coriaceous, ca. 13 mm long, slenderly long- acuminate from an oval base, green, initially covered with scattered hairs, at length glabrous, ciliate-margined; leaflets 8-12 (14) pairs, distant, ob- long-elliptic, (3) 5-10 mm long, 1.5-2 mm broad, terminating in a prickle, glabrous except sometimes for isolated hairs on the midrib beneath, prom - inently veined, cuneate at base; flowers 4 in leaf axils, crowded in a capitate inflorescence ca. 2-2.5 cm in diam., bracts filiform, 9 mm long, glabrous from base to the middle, pubescent upward; calyx ca. 16-17 mm long, white-shaggy down to base, the slenderly filiform long-hairy, gla- brous-tipped, brownish teeth 1.5 times as long as the tube; corolla violet, drying brownish; standard 15-19 mm long, the limb overtopping the calyx, oblong, round-tipped, 12 mm long, ca. 7 mm broad in upper part, scarcely constricted at the middle, 7 mm at base with reflexed auricles, the claw narrow, cuneate-attenuate toward base; wing-petals ca. 18 mm long, the lanceolate limb acutish, 1.5 mm broad at base, 2/3 as long as the claw; keel ca. 15 mm long, the oval limb ca. 5 mm long, 2 mm broad; stamens fused at base with wings and keel; ovary oblong, hairy; style 15 mm long, naked; pods oblong, 5 mm long, white-shaggy; seeds reniform, ca. 3 mm long, ca. 2 mm broad, tawny, black-punctate. May-June. Possibly occurring in the USSR in W. Transcaucasia. Described from former Artvin vicinity. Type in Leningrad. 331. A. imbricatus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946):-Tragacantha imbricata Bonriss., J. c.f. A shrub; branches densely clothed in old stipules and spinescent leaf rachises; spines sparsely appressed-hairy, slender, slightly curved to nearly straight, 4-8 cm long; stipules compact, coriaceous, shaggy atfirst, becoming glabrous, ciliate-margined, 12 mm long, acuminate from an ovate base; leaflets (5) 8-10 pairs, 5-9 mm long, 2-4 mm broad, ovate to elliptic, terminating in a short point green, initially hairy, at length gla- brous except for hairs on the midrib beneath, prominently veined; lowest pair of leaflets 2 cm from base, the upper leaflets more closely set than the two lowest pairs; flowers 3-4 in the leaf axils, crowded at the base of 272 Vin VY L ik SY S Sh PLATE XXV 1, Astragalus stipulosis Boriss. - 2. A. Cerasocrenus Bge.—3. A. aureus Willd. 366 current year's shoots into a capitate inflorescence ca. 3 cm in diam. ;bracts 7 mm long, linear, hairy at apex, glabrous at base; calyx 9 mm long, white- hairy throughout, attenuate toward base, the teeth densely hairy, scarcely perceptible among the hairs, half as long as the tube; corolla pinkish (?), drying yellowish; standard ca. 18 mm long, the oblong-ovate limb 6 mm broad, 12 mm long, twice the length of the claw, with obsolescent obtuse auricles at base, the claw 1.5 mm broad; wing-petals about equaling the standard, the oblong-lanceolate limb half as long as the claw, auriculate at base; claw of keel 2.5 times the length of the limb; stamens fused to high up with wings and keel; ovary oblong, hairy; style naked, ca. 12 mm long. July. Mountains. - Possibly in W. Transcaucasia. Described from the former Artvin vicinity near village of Kuchena. Type in Leningrad. 332. A. Voronovianus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).-Tragacantha Voronoviana Boriss. l.c. A densely branched shrub; branches rather slender, 3-4 mm in diam., clothed in imbricated stipules and spinescent leaf rachises; spines spread- ing, straight or curved, 3-4 (5) cm long, covered with scattered white curly hairs; stipules glabrous when young, long-ciliate on the margin, 7-9 mm long, acuminate, coriaceous, stramineous, oblong at base; leaflets 5-7 pairs, distant, the lowest pair 1.5 cm from base, small, ovate, 3-6 mm long, 1.5-3 mm broad, terminating in prickle, sparsely to rather densely hairy, cuneate at base, prominently veined; flowers 4 in the leaf axils, crowded in a rounded inflorescence ca. 2 cm in diam.; bracts narrowly linear, subfiliform, ca. 5mm long, 0.5 mm broad, glabrous at base, cov- ered in upper part with long white hairs; calyx ca. 11 mm long, acute, covered throughout with long white hairs, the filiform teeth equaling the tube; corolla lilac, drying yellowish, exserted from the calyx; standard ca. 17 mm long, the limb pandurate, nearly round at apex, 5-6 mm broad in upper part, constricted at the middle to 4 mm, 6-7 mm broad at base, with hooked auricles, 1.5 times the length of the claw, this narrow, 6-7 mm long; wing-petals 15-16 mm long, the limb lanceolate, 2/3 the length of the claw; obtusely auriculate; keel ca. 14 mm long, the oval limb 2/3 the length of the claw; stamens fused to high up with wings and keel, 13 mm long; ovary oblong, covered with short hairs; style ca. 11 mm long, naked, hairy mere- ly at base; pods oblong, shaggy, ca. 5 mm long. June-July. Dry slopes near pine forests. - Possibly growing in Western Transcauca- sia. Described from former Artvin vicinity near village of Tsitvet. Type in Leningrad. 333. A. Meyeri Boiss. Diagn. ser. I, 9 (1849) 88; Boiss. Fl. or. Il, 349; Bge. Astrag. geront. II, 150; Grossh. Fl. Kavk. II, 309.-A. com- pactus C.A.M. Enum. Casp.-Caus. (1831) 143, non Willd.; Ldb. Fl. Ross. II, 640, pro parte, non Willd.-Tragacantha Meyeri O. Ktze, in Rev. gen. (1891) 946.-Ic.: Fisch. Syn. trag. tab. E, f. 46 p.p. A spiny branched shrub, up to 30 cm tall; branches short, crowded, beset with persistent leaf rachises, these divergent, curved, brown, thinly pannose-pubescent, thickish, (1.5) 2-5 (7) cm long, the lower subhorizontal- ly spreading, the upper erect; stipules triangular-lanceolate on sterile, and broadly ovate on flowering branches, shori-acuminate, thinly pannose- hairy, ca. 7 mm long and broad; leaf rachises terminating in a short spine; leaflets 4-15 pairs, approximate, broadly elliptic, terminating in a short 274 67 68 point, whitish-cinereous by short densely pannose pubescence, 8-12 mm long, 4-6 mm broad, often conduplicate; flowers 4-5 in the leaf axils, form- ing a spherical or ovaloid inflorescence 2-3 cm in diam., borne at the base of the current year's branches; bracts very narrow, linear-spatulate, sub- ulate-based, dilated upward, somewhat less than 1 mm broad, navicular, glabrous at base, white-pannose upward, ca. 7mm long; calyx8-9mm long; attenuate toward base, pannose with white silky hairs, the linear curved teeth longer than the tube, this splitting nearly to base; corolla pink (pale red, according to Boissier), 12-14 mm long; standard oblong, obsoletely or distinctly constricted at the middle, nearly exauriculate, the limb ca. 8 mm long, gradually attenuate into a cuneate claw; wing-petals fused at base with keel and staminal tube, ca. 11 mm long, the limb lanceolate, en- larged at base, half as long as the claw; keel about equaling the wings, the oval limb half as long as the claw; ovary white-hairy; style naked. Fl. June-July. Gravelly dry slopes, at altitudes 1000-1800 m.-Caucasus: S. Transc. (Karabakh, Bogulty, Diabar). Tal. Gen. distr.: N.W. Iran. Described from Talysh. Type in Leningrad. Series 3. MACROCALYCINI Boriss. - Flowers 4-12 in the leaf axils; calyx large, 12-25 mm long, shorter than to markedly exceeding the corol- la, the long teeth equaling to considerably longer than the tube; corolla 15- 25 mm long; claw of standard commonly broad; inflorescence mostly large, (1.5) 3-5 cm broad. 334, A alopecuroides Pall) Spec. Astrag. (1800) 9.-A. Clausii C.A.M. in Goebel Reise in die Steppen des siidl. Russl. II (1838) 265; Ldb. Fl. Ross. I, 624.-A. lagocephalus Fisch. et Mey. in Bull. Phys.-Math. Acad. Pét., II (1844) 197.-Ic.: Pall. InCapecabe VIG GaA VEN ices tabi. a. Perennial; stem erect, more or less flexuous at the end, angled, finely sulcate, sparsely covered with straight simple long spreading hairs or sometimes glabrate, up to 35-40 cm long; stipules ovate-triangular or lan- ceolate, 1 cm long, acute, ciliate-margined; leaves 7-19 cm long, the petiole 2-3 cm long; leaflets 12-15 pairs, broadly ovate or obcordate, sub- obtuse, 0.8-2.8 cm long, 0.5-2 cm broad, covered beneath with short hairs, glabrate above; flowers sessile; inflorescence ovoid to ovoid-oblong, 4- 6 cm long, 3-5 cm broad, borne on a short peduncle or the upper ones sub- sessile; bracts linear-lanceolate to linear, acute, 1-2 cm long, ciliate, about 2/3 the length of the calyx; calyx campanulate, shaggy with long white spreading hairs, 1.5-2.3 cm long, 0.5 cm broad, exceeded by the corolla, the linear acute teeth about as long as the tube; corolla yellowish; standard 2.8 cm long, 1 cm broad, about equaling to somewhat exceeding the wings and the keel, the limb oblong-obovate, obtuse or scarcely retuse, glabrous or outside somewhat pubescent, gradually attenuate into and 1.5 times as long as the claw; limb of wings oblong, obtuse, about equaling the claw; keel angular, the limb 1.3 cmbroad, about twice as broad asthe wings, about equaling the claw, obtuse; ovary pubescent; style pubescent to half its length; pod ovaloid, densely covered with slender straight white hairs. - May-June. Sandy and clayey steppes, sandy shores, sandhills under plant cover, saline places, meadows and stony sites. -European part: L. V., Transv. ; W. Siberia: U. Tob., Irt., Alt., Soviet Centr. Asia: Ar.-Casp., Balkh., Dzu-Tarb. Endemic. Described from Siberia. Type in Berlin. 360. A. trichocalyx Trautv. in A.H.P. IV, 1 (1876) 362; Boiss. Fl. or. Suppl. 185; Grossh. Fl. Kavk. II, 303. Perennial; stem erect, terete, subsulcate, densely covered with ap- pressed simple straight white hairs; stipules lanceolate to linear-lanceo- late, long-acuminate, up to 3 cm long, densely ciliate-margined; leaves 10-30 cm long, exceeding the inflorescence, the petiole 3-5cm long; leaflets about 20 pairs, oblong to oblong-lanceolate, 1-3 cm long, 0.3-1 cm broad, subacute, glabrous, commonly ciliate-margined; flowers sessile; inflor- escence many-flowered, rather dense, short-cylindric to oblong, 6-12 cm long, 3-5 cm in diam., borne on peduncles 2-4 cm long in the axils of 2-6 uppermost leaves, pubescent with short straight appressed white hairs: bracts subfiliform, 1.5-1.7 cm long, equaling the calyx, covered with long simple white hairs; calyx cylindric-campanulate, 1.5-1.7 cm long, 4 mm broad, covered with spreading simple long white hairs; the subulate teeth equaling the tube; corolla ochroleucous; standard 2.2 cm long, somewhat exceeding the calyx, the wings and the keel, the limb ovate-elliptic, gla- brous, obtuse, gradually attenuate into and 3 times the length of the claw; 293 392 wing-petals 1.7-1.8 cm long, about equaling to somewhat exceeding the keel, the limb about as long as the claw; limb of keel somewhat shorter than the claw; podbroadly ellipsoid, obtuse, sessile, 7mm long, 4mm broad, glabrous, subcompressed, equaling the calyx-tube. June-July. (Plate XXVI, Figure 2). Stony places in the intermediate mountain zone. - Caucasus: E. Transc. Gen. distr.: Arm.-Kurd. Described from the vicinity of Abastuman. Type in Leningrad. Series 2. SPHAERANTHI Gorschk. - Inflorescence spherical; pods ovoid- lanceolate, oblong-ovoid, or obovoid, cartilaginously thickened, not oblique- ly truncate. 361. A. kulabensis Lipsky in A.H.P. XXVI (1910) 158; Fl. Tadzhik. V, 405-406. Perennial, densely vested throughout with long white hairs; stem erect, up to 1 m long, more or less flexuous at the end, densely leafy; stipules lanceolate, long-acuminate, 0.7-1 cm long; leaves paripinnate, 9-14.5 cm long; leaflets 15-20 pairs, elliptic to elliptic-lanceolate, subobtuse, 0.7- 1.7 cm long and 0.3-0.7 cm broad; flowers short -pediceled; inflorescence many-flowered, spherical to oblong-spherical, subsessile or borne on an abbreviated peduncle, 4-4.5 cm in diam., 1/4 to 1/3 as long as the leaf; bracts linear, 1 cm long, 1/2-2/3 the length of the calyx, acute, ciliate; calyx broadly cylindric, 1.6-2 cm long, 0.5 cm broad, densely covered with long white hairs, the tube about equaling to sometimes somewhat exceeding the narrowly linear tomentose teeth; corolla ochroleucous; standard 2.4- 2.6 cm long, exceeding the calyx, pubescent on the outside, the limb oblong, obtuse or somewhat retuse, attenuate toward base into and 1/4 as long as the claw; wing-petals equaling the standard, glabrous, the limb shorter than the claw; keel glabrous, about equaling or somewhat shorter than the wings, the limb half as long as the claw; pod ovoid-lanceolate to broadly lanceolate, terminating in a point, 0.9-1 cm long, densely shaggy with long white hairs; reticulate, compressed, bilocular, the locules 1-seeded; seeds ovaloid-reniform, light brown, 3.5 mm long and 2.5 mm broad. May-July. (Plate XXVIII, Figure 3). Mountain slopes and elevated foothills (up to 600-1500 m).- Soviet Centr. Asia: Pam.-Al. (S.). Endemic. Described from Kulyaba district at the Aksu river. Type in Leningrad. 362. A. mogoltavicus M. Pop. in Byull. Sr.-Az. Gos. univ. (Bulletin of the Soviet Central Asian State University) 4 (1926) 139; Fl. Tadzhik. V, 407. Perennial; stem erect, somewhat flexuous at the end, glabrous, up to 1 m long, terete, faintly angled, finely sulcate; stipules lanceolate, finely pointed, subsubulate, 1.5-2 cm long; leaves 15-28 cm long, the petiole 1-3 cm long; leaflets 13-31 pairs, oblong, obtuse or mucronulate, 0.7-1.4 (2) cm long, 0.3-0.5 (0.6) cm broad, sparsely covered beneath with ap- pressed white hairs; flowers subsessile; inflorescence many -flowered, spherical, 3-3.5 cm in diam., borne on a pubescent peduncle 1-2.5 cm long; bracts narrowly linear, 1.2-1.5 cm long, about equaling the calyx, hyaline, covered with long hairs; calyx cylindric-campanulate, densely white -shaggy with long white hairs, 1.3-1.5 cm long, the linear acute glabrous-tipped teeth equaling to somewhat exceeding the tube; corolla ochroleucous; stand- 395 ard 1.3-1.5 cm long, about equaling the calyx, the limb ovate, obtuse, pu- bescent on the outside, gradually attenuate toward base into the claw, this 294 Yes TaN Q PLATE XXVI 1, Astragalus alopecias Pall.—2. A. trichocalyx Trautv.—3. A. Kulabensis Lipsky.— 3. A. maximus Willd.—5. A. globiceps Bge.—6. A. agameticus Lipsky. 295 396 3-4 mm long; wing-petals incurved, glabrous, about equaling the standard, the limb oblong, somewhat longer than the claw; keel 1.1 cm long, somewhat exceeded by the standard and the wings, glabrous, the gibbous obtuse limb about as long as the claw; ovary oblong-ovaloid, subsessile, densely pu- bescent with white appressed hairs; pod obovoid, 1 cm long, glabrate except for white hairs at the top, gibbous, compressed, obtuse. May. Sodded gravelly mountain slopes and river valleys. - Soviet Centr. Asia: T.-Sh. (Mt. Mogoltau, Kuraminskii range). Endemic. Described from Mt. Mogoltau (Uch' bag river valley). Type in Tashkent. 363. A. Cristophii Trautv. in A.H.P. IX, 1 (1884) 378; Grossh. FI. Kavk. II, 304. Perennial; stem erect, more or less flexuous, terete, finely sulcate, covered with soft straight spreading white hairs; stipules linear-lanceolate, acute, 1-1.2 cm long, ciliate-margined; leaves 24 cm long, greatly exceed- ing the inflorescence, the petiole 1.5-3 cm long; leaflets 20 pairs, elliptic, obtuse, 0.7-1.8 cm long, 0.3-1.1 cm broad, glabrate or faintly pubescent above, densely covered beneath with soft white hairs; inflorescence many- flowered, spherical, 3.5-6 cm in diam., borne on a peduncle 4.5-7 cm long and covered with soft white hairs; bracts narrowly linear, 1 cm long, sub- ulate, acute, ciliate-margined, exceeded by the calyx; calyx turbinate - campanulate, more or less inflated, 1.8 cm long, 5 mm broad, devoid of bracteoles, covered with soft simple straight appressed long white hairs, the linear -filiform ciliate teeth about equaling the tube; corolla yellow, 1.5 times as long as the calyx; standard 2.4 cm long, about equaling the wings and the keel, pubescent above, the elliptic obtuse limb gradually attenuate into and twice as long as the claw; wing-petals and keel glabrous, the limb about twice the length of the claw; pod sessile, oblong-ovoid, covered with white hairs. May-June. Soviet Centr. Asia: Mtn. Turkm. (Akhal-Teke). Endemic. Described from Cristoph's specimen from Turkmenia. Since both Akhal-Teke (Turk- menia) and Karabakh were indicated on the label, this species has been erroneously reported also for the Caucasus. Type in Leningrad. 364. A. schahrudensis Bge. Astrag. geront. (1868) 61; (1869) 103; Boiss. Fl. or. Il, 416.-Exs.: H.F.A.M. No. 394. Perennial; stem glabrous, erect, somewhat flexuous at the end, finely sulcate, up to 1 m long; stipules broadly ovate, acute, long-ciliate on the margin, subauriculate, 1-2 cm long; leaves 13-25 cm long, the petiole 1.5- 3 cm long; leaflets 6-8 pairs, ovate-oblong, obtuse, glabrous, 1.5-3.7 cm. long and 0.6-2 cm broad; inflorescence many-flowered, axillary, subspheri- cal, 4 cm in diam., subsessile or borne on a glabrous or sparsely hairy - peduncle 1.3 cm long; bracts narrowly linear, covered with long hairs, 1.5- 1.8 cm long, about equaling the calyx; calyx campanulate, white-shaggy, 1.3-1.8 cm long and 0.5 cm broad, exceeded by the corolla, the linear acute pubescent teeth about equaling to somewhat exceeding the tube; corolla och- roleucous; standard 1.7-2.3 cm long, about equaling the wings and the keel, the suborbicular retuse limb 1.3 cm broad, attenuate toward base into and somewhat longer than the claw; wing-petals equaling the keel, the limb about as long as the claw; pod obovoid, covered with stiff hairs. May. Mountains (1200-1500 m), on stony slopes. - Soviet Centr. Asia: Mtn. Turkm. Gen. distr.: Iran. Described from Iran (Shakh-Rud). Type in Paris. 296 397 365. A. finitimus Bge. Astrag. geront. I (1868) 61, II, (1869) 102: Boiss. Fl. or. II (1872) 416 (excl. pl. e Kachetia); Trautv. in A.H.P. IV, 2 (1876) 362 et in A.H.P. VII, 202; Grossh. Fl. Kavk. II (1930) 304, ex parte, quoad pl. ex Armenia.-A. macrocephalus Ldb. Fl. Ross. I (1842) 635, ex parte quoad pl. ex Karabagh, non Willd.; Trautv. in A.H. P. II, 2 (1873) 521; Grossh. Fl. Kavk. II (1930) 304, ex parte quoad pl, ex Armenia et Nachitschevan. Perennial; stem erect, more or less flexuous, glabrous or faintly pu- bescent, terete, rather finely sulcate, up to 40 cm long; stipules broadly triangular to broadly lanceolate, acute, 2-3.5 cm long, ciliate-margined; leaves 23-28 cm long, the petiole 4.5 cm long; leaflets 8-15 pairs, oblong - lanceolate, acuminate, glabrous, 1-4 cm long, 0.4-1 cm broad, commonly ciliate -margined; flowers sessile; inflorescence many-flowered, axillary, dense, spherical, 5-6.5 cm in diam., borne on a glabrous peduncle 3-8 cm long; bracts linear to linear-lanceolate, acute, 1.7-2 cm long, ciliate - margined, equaling to sometimes exceeding the calyx; calyx campanulate, 1.5-2 cm long, 4 mm broad, covered with long straight white hairs, the linear-subulate teeth about equaling to exceeding the tube; corolla bright yellow; standard glabrous, 2.5 cm long, about 1.5 times as long as the calyx, about equaling the wings and the keel, the limb ovate to rounded-oval, entire or faintly retuse, attenuate toward base into and 2.5-3 times as long as the claw; wing-petals and keel glabrous, about equal, the limb Slightly exceed- ing to 1.5 times as long as the claw; pod obovoid -pyriform, 7 mm long, 5 mm broad, hairy, obsoletely grooved dorsally. May-June. Steppe foothills and dry slopes.- Caucasus: E. and S. Trans c., uals Gen. distr.: N. Iran. Description based on Shovits! collections from Azer- baijan (Shakhbulakh) and Bunge's and Binert's collections from N. Iran. Type in Leningrad. 366. A. sphaerocephalus Stev. in. Mém. Soc. Natur. de Moscou IV (1832) 266, nomen nudum (Manden. descr. ); Trautv. in A.H.P. VII (1883) 241.-A. macrocephalus MB. FI. taur.-cauc. II (1808) 183, non Willd., Ldb. Fl. Ross.I (1842) 635, ex parte quoad pl. ex Iberia.-A. finitimus Grossh. Fl. Kavk. II (1930) 304, ex parte quoad pl. ex Iberia. -Exs.: Herb. Fl. Cauc: No. 124; Pl. Or. exsicc. No. 209. Perennial; stem 15-50 cm long, glabrous, slightly flexuous; leaves 14- 18 pairs, ovate or oval, round-tipped, glabrous, 2.5-3 cm long, 0.8-1 cm broad; stipules ovate -oblong, hyaline-margined; flowers sessile; inflor- escence many-flowered, secund, dense, spherical, commonly 4 rarely 4.5 cm in diam., the peduncle rarely equaling to somewhat exceeding the inflorescence; calyx campanulate, 1-1.5 cm long, 5 mm broad, densely covered with long hairs, the linear-subulate teeth equaling to just exceed- ing the tube; corolla stramineous; standard glabrous, 1.8-2 to rarely 2.2 cm long, about 1.5 times the length of the calyx, exceeding by 3-4 mm the equal wings and keel; ovary pubescent; style pubescent on lower 1/3 -1/2; pod ellipsoid, 7-8 mm long, 5 mmbroad, densely pubescent with spreading hairs. May-July. Steppe foothills and dry slopes. - Caucasus: E. Transc. Endemic. De- scribed from the village Dzegvi in Georgia. Type in Leningrad. 367. A. megalotropis C.A.M. ex Bge. in Mém. Acad. Sc. Pétersb. VO, sér. XI, 1 (1868) 60; Bge. l.c. VII, sér. XV (1870) 99; Boiss. Fl. or. II, 415; Grossh. Fl. Kavk. II, 304.-A. narbonensis C.A.M. Ver- zeichn. d. Pflanz. (1831) 143 (non Gouan); Ldb. Fl. Ross. 1, 634. 297 398 Perennial; stem erect, more or less flexuous at the end, finely sulcate, covered especially in upper part with long white more or less spreading hairs; stipules broadly ovate, 2 cm long, acute; leaves 12-19 cm long, the petiole 2-4 cm long; leaflets 15-17 pairs, oblong, 0.5-2 cm long, and 0.3- 0.9 cm broad, subacute, densely covered beneath with short hairs, glabrate above; flowers sessile; inflorescence many-flowered, spherical, 3-5 cm in diam., dense, subsessile; calyx campanulate, covered with long divergent rufous hairs, 1.7-2 cm long, 0.6 cm broad, the elongate narrowly linear teeth 1-1.2 cm long, about equaling the tube; corolla yellow; standard2.7cm long, exceeding the calyx by 7-10 mm, somewhat (1-2 mm) longer than the wings and the keel, glabrous, the limb oblong, orbicular in upper part, at- tenuate at base into and 3 times the length of the claw; wing-petals about equaling the keel, the limb 3-4 mm longer than the claw; the limb of the keel equaling the claw; ovary ovaloid-oblong, densely covered with long rufous hairs. June. Dry mountain slopes.- Caucasus: Tal. Endemic. Described from Talysh. (Zuvant). Type in Leningrad. Note. According to Meyer, the plant has not been collected by anybody else. Series 3. TURBINATI Gorschk. - Inflorescence spherical or ovoid- spherical; pod obliquely obconic, the lower part membranous, the upper part cartilaginously thickened. 368. A. turbinatus Bge. in Mém. Acad. Sc. Pétersb. VII (1854) 269. Perennial, appressed-hairy throughout; stem erect, finely sulcate, sub- terete; stipules lanceolate, acute, 1 cm long, glabrous above; leaves 17- 26 cm long, the petiole 0.7 cm long; leaflets 16-30 pairs, oblong-obovate to linear-elliptic, sparsely hairy above, subacute or terminating in a short point, 0.6-1 cm long and 0.7 cm broad; flowers sessile; inflorescence axil- lary, ovoid-spherical, 4.5 cm long and 3.7-4 cm broad, the peduncle hairy; bracts lance-linear, acute or subulate-tipped, 1.2 cm long, covered with long hairs, about half as long as the calyx; calyx cylindric, thickened at base, 2-2.2 cm long and 0.3-0.5 cm broad, shaggy with long white hairs, the long setaceous long-hairy teeth barely longer than the tube; standard 2.4 cm long, pubescent on the outside, somewhat exceeding the calyx, the limb oblong-obovate, retuse, gradually attenuate toward base into and 3 times the length of the claw; wing-petals and keel glabrous equal, somewhat shorter than the standard, the limb equaling the claw; pod obconic, short- stipitate, obtuse, puberulent. April-May. Sand-dunes and fixed sands, slightly undulating sands. Soviet Centr. Asia: Ar.-Casp., Kyz K., Kara K., Mtn. Turkm., Amu D. Gen. distr. : Iran. Described from the Kyzyl-Kum Desert. Type in Paris. Subsection 3. BIBRACTEOLATI Bge. Astrag. geront. (1869) 103. - Calyx furnished with a bract and 2 lateral filiform or linear -filiform brac- teoles. 369. A. Regelii Trautv. in A.H.P. II (1873) 472, 522; Grossh. Fl. Kavk. eee Perennial; stem erect, terete, more or less finely sulcate, covered (especially in upper part) with short appressed straight white hairs; leaves 298 399 400 6-20 cm long, exceeding the inflorescence, the petiole 0.6-2 cm long; leaf- lets 10-20 pairs, oblong, attenuate at both ends, subobtuse, 0.8-2.5 cm long, 0.3-0.9 cm broad, subcinereous, densely covered on both sides with minute appressed simple straight white hairs; inflorescence many-flowered, dense, spherical, 4-5 cm in diam., in the axils of 2-3 uppermost leaves, the pe- duncle 2.5 cm long, covered with straight white appressed hairs; flowers sessile; bracts linear, acute, 1 cm long, equaling the calyx, densely pu- bescent; bracteoles linear-filiform, 8 mm long, exceeding the calyx-tube, densely covered, like the calyx, with simple straight appressed white hairs; calyx turbinate, 0.9-i1 cm long, 5 mm broad, the linear-subulate teeth twice as long as the tube; corolla yellow, glabrous, about twice as long as the calyx; standard incurved, 1.5-1.8 cm long, 1.5 times as long as the calyx and about equaling the wings and the keel, the limb ovate or oval, obtuse, attenuate into and 7-9 times as long as the claw, this 2 mm long; limb of wings about 1.5 times the length of the claw; limb of keel somewhat longer than the claw; ovary oblong-ovoid, densely covered with simple straight appressed white hairs. June. Dry places.- Caucasus: S. Transc. Endemic. Described from Armenia (near Belava). Type in Leningrad. 370. A. agameticus Lipsky in A.H.P. XVIII (1900) 38.-A. macro- sphaerus Freyn et Sint. in Bull. Herb. Boiss., sér. II, IV (1904) 1116. - A. Androssowi Litw. in Tr. Mus. Bot. Ac. Sc. VII (1910) 73.-A. glo - biceps ssp. agameticus M.Pop.in Byull. S.A.G.U. 14 (1926) 137. — A.globiceps ssp.agameticusf. euagameticus M. Pop. etf. sphaeroideus M.Pop., l.c.,137.-A. sphaeroideus Eug.Kor.nom. Perennial, vested with short white hairs; stem erect, up to 35 cm long; stipules lance-subulate, setaceously long-acuminate, 1.5-2 cm long; leaves 10-32 cm long, the petiole 0.6-3 cm long; leaflets 25 pairs, ovate-elliptic or ovate, 0.8-2.7 cm long, 0.6-1.4 cm broad, mucronulate, glabrous, green above, rather densely pubescent beneath with short white hairs; flowers sessile; inflorescence many-flowered, loosely spherical, (3-5) 3-6-7 cm in diam., half as long as the leaves; bracts linear-subulate, exceeded by the calyx; bracteoles setaceous, white-shaggy, 1.2 cm long, shorter than to equaling the tube; calyx campanulate, 2.2-3.4 cm long, 0.5-1 cm broad, sometimes exceeding the corolla, shaggy with silky white hairs, the unequal subulate to filiform teeth equaling to somewhat exceeding the tube; corolla ochroleucous, about equaling to somewhat exceeding the calyx; standard glabrous, 2.5-3.2 cm long, somewhat exceeding the wings and the keel (about equaling the latter), the limb oblong-obovate, slightly emarginate or obtuse, 4-5 times as long as the claw; limb of wings and keel equaling to somewhat exceeding the claw; ovary sessile, oblong-ovaloid, densely covered with soft white hairs. April-May. (Plate XXVI, Figure 6). Sands, sandy hills, and river banks. - Soviet Centr. Asia: Kara K., Mtn. Turkm. Endemic. Described from the Turkmen SSR (onthe roadto Agamet). Type in Leningrad. 371. A. globiceps Bge. in A.H.P. VII, I (1880) 372; Fl. Tadzhik. V, 406.-A. flavicomus Bge. in Izv. O-va lyub., est., antr. ietn. (Society of Nature Lovers, Anthropologists and Ethnologists) II (1880) 245.- A. ti- muranus Franch. in Ann. Sc. Nat. Sér. VI, XV (1883) 256.-A. jakka- bagi Lipsky in A.H.P. XXVI (1917) 160.-A. globiceps ssp. euglo- biceps M.Pop. in Byull. S.A.G.U. 14 (1926) 137, tab. XII.-Exs.: H.F. A.M. No. 9. 299 401 Perennial; stem erect, somewhat flexuous at the end, finely sulcate, up to 70 cm long, covered with short straight spreading hairs; stipules lance-subulate, 1-1.5 cm long, ciliate; leaves 20-27 cm long, the petiole 1-3 cm long; leaflets 20-24 pairs, oblong, faintly mucronulate, 1-2 (5) cm long, 0.2-1 cm broad, densely covered beneath with long white hairs, gla- brous or sparsely hairy above; flowers sessile; inflorescence spherical, 5-6 cm in diam., the pubescent peduncle 3-5 cm long; bracts linear -subu- late, exceeded by the calyx; bracteoles filiform, hairy, acute, 0.6-1 cm long, exceeded by the calyx-tube; calyx campanulate, 1.5-2.6 cm long, 0.4 cm broad, about 2/3 the length of the corolla, densely covered with long white hairs, the tube 2.5-3 times as long as the lance-subulate to linear - subulate ciliate teeth; corolla yellow or pale yellow; standard 2.3-3.5 cm long, about equaling the wings and the keel, the limb oblong-elliptic, slight- ly retuse or obtuse, attenuate into and 3 times the length of the claw; limb of wings and keel about as long as the claw; pod ellipsoid, 0.5-0.8 cm long, narrowed at apex into a beak, shaggy with long white hairs; seeds ovaloid- reniform. May-June. (Plate XXVI, Figure 5). Foothills and mountain slopes. - Soviet Centr. Asia: Amu D., Pam.-Al., Mtn. Turkm., Syr D. (vicinity of Tashkent). Endemic. Described from Soviet Central Asia, between Lus village and Pendzhikent. Type in Paris. Section 58. EREMOPHYSA Bge. Astrag. geront. I (1868) 62. - Pe- rennial plants, with a well developed stem, vested with simple white hairs; stipules cauline, often adnate at base to petiole, free (not connate), com- monly green, herbaceous; leaves imparipinnate, the petiole marcescent; racemes axillary, loose or compact, borne all the way up the stem or on the upper 1/2-2/3, subsessile; each flower subtended by a bract and 2 basal bracteoles; calyx initially cylindric or campanulate, becoming inflated, nearly bladdery, thinly membranous, profusely reticulate; corolla persist- ent, glabrous; pod borne on a slender short stipe, membranous or thinly coriaceous, carinate ventrally, grooved dorsally, bilocular or nearly so, few -seeded. This section included 12 species, distributed from the sands of the Lake Balkhash and Aralo-Caspian regions, through the desert sands of Soviet Central Asia, Iran, Afghanistan, and Arabia, to North Africa. A section of Soviet Central Asian origin, comprising 10 species in Soviet Central Asia and 2 outside that area. 1. Standard 23-40 mm long, the limb oblong or oval-oblong; pod bilocular. + Standard 12-21 mm long, the limb orbicular or ovate; pod partially bilocular or nearly fully bilocular (the septum not reaching and fused withithe venttalisuture) duct « .clnad cavim bre. lin whee qe 5. 2. Keel nearly straight (scarcely convex beneath, oblong or lanceolate in GUeLN| PFS See Ma eee wee ie cay ue ee) day eae a 3. +) KWeeligibbous DEneain. cel says ons aus e+ 6, 5 6 Ge «ls. euteene 4, 3. Standard (30) 35-40 mm long, narrowly oblong; keel acute; calyx 32- 40 mm long in fruit; bracts lanceolate to oblong-lanceolate, 15-20 mm long; leaflets 4-6 cm long, covered beneath with scattered hairs... . Guveuiaieh te’ hrlcdlaeh cle sie Milo) eateia ta sue Meat cunt ts oe... 372, A, Tatjanae Lincz. 300 102 + Standard 32 mm long, oval-oblong; keel obtuse; calyx 23-27 mm long in fruit; bracts linear-lanceolate, 5-6 mm long; leaflets 2-3.2 cm long, densely shaggy beneath with soft hairs. 373. A. subkahiricus Gontsch. 4. Stem, the underside of leaflets, petiole and calyx hairy .......... sired ees eadae cs itw cm erin: mead. tenthad 374, A. Maximowiczii Trautv. + Stem and the underside of leaflets glabrous; calyx glabrous except for fhesthbroatandsthe outer sunfacelorwhesteet haces feel. Cil-ieme. ile tet le ete petites wads stereicairsie: yer daer ardor tod ene dos 2 wl 375. A. Litvinovii Lipsky. Ovary and pod densely hairy ......... 376. A. Lehmannianus Bge. Ovary and pod glabrous, rarely sparsely hairy .w)..5). ya. 22 .. 6. mAaMlet Ss loa l8.(20-palRS)rote inthe cdee enthe che Ssh wAt Winkleri Trautv. Peabo c4—1.0.(d 16) ade Sine tiie eitensleaiertatel omaaeley woceliel ae haw ttre Ue Racemes long, loose, remotely many-flowered, (10) 15-20 (25) cm long; standard 12-13 mm long; corolla purple (rarely yellow or fading GES) eatery ccc iucye'= agile uaa aber yatt cide o Ste, Ryacepttuie ees Taek 381. A. chivensis Bge. Racemes compact or fairly loose, 3-8 cm long; standard 15-21 mm lononsconolla yellow (rarely fadime red)iy “se, siccsc-terare @ eve lee is, 2 eke 8. Wears) AO DATES vs cule essence vou 379. A. sphaerophysa Kar. et Kir. eC G MC ORO Ss ale Lint tA BrIY, ARE roee sued Bebe! OL en. esata s aye 2 oF Calyx cylindric; standard 20-21 mm long.. 377. A. kelifi Lipsky. Calyx short-cylindric (campanulate Se ndrie): standard 17-18 mm long. a volto: 2 Bldg a GMS Roleqeedactonaimeiies Shas tute Manta PRT Hpi hut oA an 380. A. leiophysa Bge. A+ Ot + + OF © Subsection 1. DOLICHANTHA Gontsch. subsect. nova. - Standard 23-40 mm long, the limb narrowly oblong or oval-oblong; pod bilocular. A most primitive section, comprising the most ancient species, distributed in the foothills of S. W. Pamir-Alai, in the southern deserts of Soviet Cen- tral Asia (Kara Kum), Iran, Afghanistan, Arabia, and North Africa. Series 1. RECTIFLORI Gontsch.-Keel nearly straight, scarcely or slightly convex beneath. Beside the Soviet Central Asian species, this series includes A. kahi- ricus DC. which is distributed in North Africa, Arabia, Mesopotamia and, probably, southern Afghanistan. 372. A. Tatjanae Lincz. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS Weinman @eaa) iva. le. : Ie. 1 1. Perennial, 40-65 cm tall; stems 70-100 cm long, prostrate or ascending, up to 30-50 cm long, subterete, stout (up to 1 cm indiam.), glabrous, cov- ered in upper part with scattered hairs; stipules herbaceous, cauline, par- tially amplexicaul, triangular, 1.5-2 cm long, acuminate. covered with scattered hairs; leaves sparsely covered with long white hairs, the Jower 22-30 cm, the upper 15-19 cm long, the petiole finely sulcate; leaflets 3-4 (6) pairs, rounded-ovate or broadly elliptic, obtuse, 5-8 cm long, 4-6cm broad, glabrous above, ciliate-margined; racemes loosely 15-40-flowered, 20-60 cm long; bracts herbaceous, green, rarely hairy, ciliate-margined; peduncles 2-4 mm long, densely hairy; bracteoles [?] subulate-linear, ca. 4mm long, hairy; calyx cylindric in flower, 18-19 mm long, loosely shag- gy with spreading long white hairs, the teeth lanceolate, 5 mm long, acu- minate, densely bearded-shaggy inside; fruiting calyx often purple -tinged, oblong-ovoid or bladdery oblong-ellipsoid, thinly membranous, profusely reticulately nerved, covered with scattered hairs, the teeth triangular, 301 403 acuminate, 6-9 mm long; corolla yellow; standard (30) 35-40 mm long, the limb entire, round-tipped, rather gradually attenuate into and twice as long as the claw; wing-petals (29) 34-38 mm long, the limb linear, obtuse, about equaling the claw; keel (27) 32-35 mm long, the limb slightly convex beneath, nearly straight above, about as long as the claw, somewhat in- equilaterally lanceolate in outline; ovary on a stipe ca. 1.5 mm long, gla- brous; style naked; pod borne on a slender stipe ca. 5 mmlong, lance- oblong, 18-20 mm long, inflated, carinate ventrally, grooved dorsally, acute, thinly coriaceous, glabrous, terminating in a beak up to 4 mm long. Fl. April-May; fr. May. Sandy loess hills in the foothills; entering into the composition of mead- owgrass-and-sedge ephemeroid meadows with xerophyll perennials, and in pistachio groves. - Soviet Centr. Asia: Pam.-Al. (S.: watershed of the rivers Tair su and Vakhsh). Endemic. Described from Mt. Utunchi in the Kurgantyubin District. Type in Leningrad. 373. A. subkahiricus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, up to 50 cm tall; stems well developed, up to 35 cm long, firm, ribbed, shaggy with spreading hairs; stipules cauline, often adnate at base to petiole, herbaceous, green, triangular, 6-9 mm long, shaggy on the outside; leaves 13-17 cm long; the petiole indistinct, 2.5-3 cm long, the petiole and the rachis shaggy with soft spreading hairs; leaflets 4-5 pairs, suborbicular, green and glabrous above; racemes sessile, compactly many-flowered, 14-15 cm long (partially in fruit); peduncles 2 mm long, shaggy with spreading hairs; flowering calyx narrowly cylindric, 16-20mm long, shaggy with soft spreading hairs, the lanceolate-linear teeth 4-5 mm long; fruiting calyx ovoid-bladdery, thinly membranous, profusely reticu- late-nervose, rather densely shaggy with soft white hairs, the triangular - lanceolate teeth 7-8 mm long; corolla yellow, round-tipped and scarcely retuse, gradually attenuate at base into the claw, this ca. 10 mm long; wing-petals equaling the standard, the limb linear-oblong, round-tipped, barely longer than the claw; keel ca. 31 mm long, the limb just convex be- neath, strongly convex above (nearly straight in general outline), about equaling the claw; ovary on a stipe 2 mm long, glabrous; style naked; pod borne on a slender stipe 3 mm long, oblong-ovaloid, 8-9 mm long, ca. 5 mm broad, terminating in a straight subulate slender beak 1-1.5 mm long, membranous, glabrous, inflated, carinate ventrally, grooved dorsally. Fl. April; fr. April-May. : Soviet Centr. Asia: Pam.-Al. (S.W. spurs of the Kugitang range). En- demic. Described from the foothills between Kelif and village Okuz-bulak. Type in Tashkent. Series 2. GIBBOSI Gontsch. -Keel gibbous beneath. Two species from Soviet Central Asia and one (A. gibbosus Gontsch. sp. nova) from N. and Centr. Iran and W. Afghanistan. 374. A. Maximowiczii Trautv. in A.H.P. IX (1884) 379.-Exs.: P. Sint. Iter transcasp.-pers. 1900-1901 No. 100 sub. nom. A. Lehmanniani Bge. Perennial, 50-70 cm tall; stems firm, angled-ribbed, 45-60 cm long, pubescent or velutinous; stipules cauline, free, herbaceous, green, glabrous, sparsely white-ciliate on the margin, broad, transversely subreniform, 302 404 407 clasping the petiole, abruptly acuminate, rarely triangular -oblong, 7-10mm long; leaves (10) 16-25 (30) mm long, the petiole (3-5 cm long) and the ra- chis pubescent or velutinous; leaflets 5-7 (10) pairs, ovate or rounded-ovate, rarely elliptic, subobtuse or round-tipped, 20-40 (50) mm long, finely mucron- ulate, borne on pubescent petiolules 1-3 mm long, rounded or broadly cuneate at base, glabrous above, green, densely pubescent beneath with short ap- pressed hairs, densely white-ciliolate on the margin; racemes sessile, loosely many-flowerered, 12-17 cm long; bracts linear-lanceolate, sub- acute, greenish, 4-8 mm long, ciliate-margined; peduncles, ca. 1 mmlong, hairy; calyx at the early flowering stage 12-13 mm long, white-velutinous, the teeth triangular, acuminate, 2-3 mm long, bearded-lanate inside and on the margin with white hairs; fruiting calyx inflated, ovoid-bladdery, 25- 30mm long, thinlymembranous, profusely reticulate-nervose, moreor less purple-tinged, rather profusely covered with short white hairs; corolla pur- ple, glabrous; standard 22-30 mm long, the limb oval to oblong-oval, round- tipped and slightly retuse, rounded or broadly cuneate at base, the claw 8- 10 mm long; wing-petals 20-25 mm long, the limb linear-oblong, slightly attenuate upward, round-tipped, about as long as the claw; keel about equal- ing the wings, the limb gibbous beneath, straight above, obtuse, as long as the claw; ovary ona short stipe 3-4 mm long, lanceolate, slightly hairy; style long, slender, naked; pod borne on a slender stipe 4-5 mm long, oval or oblong in side-view, 8-11 mm long, inflated (chiefly ventrally), some - what compressed laterally, grooved ventrally, carinate and prominently sutured dorsally, thinly coriaceous, finely cross-nervose, covered with scattered short white hairs, few-seeded, the slender beak 2-3 mm long. Fl. March-April (May); fr. April-May. (Plate XXVII, Figure 2). Hilly and flat consolidated sands. - Caucasus: (?) S. Transc. (reported by Trautvetter for the Caucasus); Soviet Centr. Asia: Kara K. Endemic. Described from Turkmenia (Akhal-teke) and from Transcaucasia (Karabakh). Type in Leningrad. 375. A. Litvinovii Lipsky in A.H.P. XVIII (1901) 39. Perennial, 40-60 cm tall; stems well developed, 33-35 cm long, firm, ribbed, glabrous; stipules caulescent, free, herbaceous, green, broad, transversely subreniform, clasping the petiole, abruptly acuminate, rarely triangular-oblong, 7-10 mm long, glabrous, sparsely white-ciliate on the margin; leaves (young 7) 15-27cm long, the petiole rather indistinct, 3-4 (5) cm long, both petiole and rachis glabrous, sulcate; leaflets (4) 5-7 (9) pairs, distant, ovate to subelliptic, rarely obovate, commonly acute, rarely more or less round-tipped and mucronulate, rather broadly cuneate at base, (8) 23-35 (41) mm long (large on the lower leaves, much smaller on the upper ones), glabrous on both sides, green, sparsely white-ciliolate on the margin, the petiolules 1-2 mm long; racemes sessile, loosely many- flowered, 12-16 cm long, the rachis glabrous, sulcate; bracts herba- ceous, green, linear-lanceolate, acute, 4-6 mm long, ciliate-margined; pedicels 1-2 mm long, glabrous; flowering calyx 9-13 mm long, sparsely white -hairy or glabrate, the teeth triangular, acuminate, 2-3 mm long, glabrous on the outside, barbate-lanate on the inside and on the margin with white hairs; fruiting-calyx inflated, ovoid-bladdery, 22-25 mm long, thin- ly membranous, profusely reticulate-nervose, glabrous, lustrous, more or less purple-tinged; corolla purple or violet (?), with a purple keel; standard 24-30 mm long, the limb oval to oblong-oval, round-tipped and slightly retuse, nearly round at base, the claw 8-12 mm long; wing-petals 303 i hie ae 5 Ret Fh gs: i = p>, i hee PLATE XXVII 1, Astragalus chivensis Bge.—2. A. Maximoviczii Trautv.—3. A. Lehmannianus Bge. 304 408 21-26 mm long, the limb entire, linear-oblong, slightly attenuate upward, round-tipped, about as long as the claw; keel 20-25 mm long, the limb gib- bous beneath, straight above, obtuse, about as long as the claw; ovary on a slender stipe 3-4 mm long, lanceolate, glabrous; style long, slender, naked; pod borne on a stipe 4 mm long, broadly lanceolate in side-view, 8-11 mm long, thinly coriaceous, inflated (especially toward the ventral side), more or less compressed laterally, grooved ventrally, carinate and prominently sutured dorsally, glabrous or covered with isolated white hairs, few-seeded. Fl. April-May; fr, May-June. Sands. - Soviet Centr. Asia: Kara K. Endemic. Described from Mt. Aga- met. Type in Leningrad. Note. A species closely related to the preceding; differing in the more pronounced absence of pubescence and mostly in the shape of leaflets. Subsection 2. MICRAN THA Gontsch. - Standard 12-21 mm long, the limb orbicular or ovate; pod not fully bilocular (partially to almost fully bilocular). Series 1. TUBULATI Gontsch. - Calyx cylindric, velutinous in flower, rarely densely shaggy. 376. A. Lehmannianus Bge. in Arb. Nat. Ver. Riga I (1847) 352; Rel. Lehm. (1851) No. 382; Bge. Astrag. geront. II, 106; Bge. Astrag. turk. 248; Boiss. Fl. or. II, 420; Grossh. Fl. Kavk. II, 302. Perennial, 35-65 cm tall; stems erect, stout, firm, 25-50 cm long, terete, sulcate, velutinous, at length glabrous; stipules cauline, free, par- tially adnate at base to petiole, triangular, acute or acuminate, 8-9 mm long, green, shaggy on the outside; leaves 15-27 cm long, the petiole rather indistinct (up to 2 cm long), both petiole and rachis firm, pubescent; leaf- lets (6) 7-10 (11) pairs, orbicular or nearly so (rarely broadly cuneate at base), rather thick, round-tipped, very rarely scarcely retuse, (10) 15-25 (30) mm long, green above, covered with scattered appressed hairs (es- pecially at the margins), shaggy beneath with silky pubescence; peduncles 0-2 cm long, firm, velutinous-pubescent; racemes rather firm, (7) 9-14 cm long, many-flowered; bracts linear, subulate-acuminate, 5-6 mm long, ~ green, Shaggy on the outside, profusely ciliate-margined; bracteoles simi- lar in shape, 3-4 mm long; flowering calyx 11-15 mm long, densely silky - velutinous, the teeth linear-acute, 4-5 mm long, half as long as the tube; fruiting calyx on an inflated stipe, ovoid to subspherical-bladdery, 17-22 mm long, velutinous, thinly membranous, profusely reticulate-nervose, the teeth linear acute from a triangular base; corolla yellow, glabrous; stand- ard 16-20 mm long, the limb ovate-orbicular, scarcely retuse, as long as the claw; wing-petals equaling the standard, the limb linear-oblong, entire, round-tipped, somewhat shorter than the claw; keel somewhat exceeded by the standard and the wings, the limb obtuse, convex beneath, straight above, about 2/3 the length of the claw; ovary on a stipe 2 mm long, shaggy; style naked; pod borne on a slender stipe 2-3 mm long, oblong-linear in side- view, 7-9 mm long, inflated (especially toward the dorsal side), carinate ventrally, grooved dorsally, thinly coriaceous, silky-velutinous, termina- ting in a beak 1-2 mm long, bilocular to 2/3, 2-4-seeded; seeds ovaloid to oblong-ovaloid, compressed, 4 mm long, scarcely minutely pitted (near- ly smooth), tawny. Fl. May-June (July); fr. June-July (August). (Plate XXVII, Figure 3). 305 409 Fixed sands, sandy river terraces, sand dunes.- Caucasus: Dag. ; Soviet Centr. Asia: Ar.-Casp., Kara K., Kyz. K., Balkh. (rare). Endemic. De- scribed from Aral Lake area. Type in Leningrad. 377. A. kelifi Lipsky in A.H.P. XXVI (1910) 158.-Exs.: Herb. Fl. As. Med. No. 387. Perennial, 32-52 cm tall; stems 20-40 cm long, angled-sulcate, veluti- nous with spreading soft hairs, firm, erect; stipules cauline, free, adnate at base to petiole, green, triangular, acuminate, 5-7 mm long, white -hairy on the outside and on the margin; leaves (10) 14-20 cm long, the petiole 2-3 cm long, both petiole and rachis shaggy with spreading short white hairs; leaflets 7-10 pairs, distant, orbicular, slightly emarginate, (5) 7-11 mm long, thickish, glabrous green above, shaggy-canescent beneath with ap- pressed silky hairs, densely white-ciliolate on the margin; racemes sessile, 5-8 cm long, 9-14-flowered, the rachis hairy-velutinous with white spread- ing hairs, the flowers sessile; bracts linear-lanceolate, acuminate, 4-5mm long, thinly membranous, white-shaggy on the outside and on the margin; bracteoles linear, ca. 3mm long, hyaline, white-shaggy on the outside and on the margin; flowering calyx cylindric, 12-13 mm long, densely velutinous- silky; fruiting calyx ovoid-bladdery, 16-18 mm long, thinly membranous, profusely reticulate -nervose, densely shaggy with soft and more or less spreading hairs, the teeth 3-4 mm long, initially subulate shaggy, at length triangular and subulately acuminate; corolla yellow; standard 20-21 mm long, the limb obovate, rarely oval, gently angular at base, scarcely re- tuse, as long as the claw; wing-petals equaling the standard, the limb ob- long, entire, round-tipped, barely shorter than the claw; keel 19-20 mm long, the limb obtuse, gibbous beneath, straight above, the claw 11-12 mm long; ovary on a stipe 3 mm long, glabrous or slightly white-hairy; style slender, long, naked; pod borne on a slender stipe 3 mm long, membranous, ovaloid to oblong-ovaloid, ca. 8 mm long, inflated (especially toward the back), slightly compressed, carinate ventrally (facing the standard), deeply grooved dorsally, sparsely covered with spreading hairs or glabrate, near- ly fully bilocular (septum just failing to reach the opposite suture and much more removed from it at the apex), few-seeded. Fl. April-May; fr. May. Wind -blown or less frequently fixed sands, in river valleys. - Soviet Centr. Asia; Pam.-Al. (S.W.). Endemic. Described from the vicinity of Kelif. Type in Leningrad. 378. A. Winckleri Trautv. in A.H.P. 1X (1884) 449. Perennial, 35-60 cm tall; stems firm, erect, 24-40 cm long, white-— pubescent, with appressed hairs toward base and spreading hairs upward; stipule cauline, free, adnate at base to petiole, herbaceous, green, triangu- lar-lanceolate to linear-lanceolate, acute, ca. 5 mm long, green, glabrous, white -ciliate on the margin; leaves (10) 15-23 (30) cm long, greatly exceed - ing the raceme, the petiole rather indistinct, 1.5-2 cm 1ong, both petiole and rachis densely white-pubescent; leaflets rather thick, orbicular (very rarely obovate), round-tipped or scarcely retuse, (5) 6-10 (12) mm long and broad, glabrous above, densely appressed-pubescent beneath; racemes sub- sessile, 9-13 cm long (up to 17 cm in fruit), loosely and remotely many- flowered, the rachis densely pubescent, the flower subsessile; bracts lance- linear, subulate-acuminate, 4-5 mm long, green, glabrous, white -ciliate on the margin; bracteoles same, but smaller, 2.5-3 mm long; flowering calyx cylindric, 12-13 (15) mm long, densely subpannose white-shaggy, 306 410 the teeth linear, 3-4 (5) mm long, the tube 2-3 times the length of the teeth; fruiting calyx inflated, ovoid-bladdery, 15-20 mm long, thinly membranous, profusely reticulate-nervose, shaggy-pubescent, the teeth triangular -lan- ceolate, long-acuminate; corolla yellow; standard 16-18 mm long, the limb rounded -ovate to suborbicular, round-tipped, the claw 7-8 mm long, wing- petals equaling the standard, the limb linear-oblong, round-tipped, as long as the claw; keel 14-17 mm long, the limb gibbous beneath, straight or scarcely concave above, obtuse, as long as the claw; ovary glabrous, ona stipe 2 mm long; style long, slender, glabrous; pods oblong-oval in side - view, inflated, membranous, 7 mm long, borne on a stipe 3-4 mm long, carinate ventrally, grooved dorsally, glabrous, terminating in a beak2mm long, bilocular to 2/3, 2-seeded; seeds oblong-ovaloid, 4 mm long, com- pressed, slightly notched laterally, finely pitted, brown. Fl. May; fr. May- June.. Sands and stony ''steppes''. - Soviet Centr. Asia: Kara K. (Krasnovodsk district). Endemic. Described from Kyzyl-Arvat. Type in Leningrad. Series 2. SUBCAMPANULATI Gontsch. - Calyx campanulate or cylindric - campanulate, shaggy. 379. A. sphaerophysa Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 338; Bge. Astrag. geront. II, 106; Bge. Astrag. turk. 246. Perennial, 27-50cmtall; stems 23-37 cm long, firm, ribbed, fully or mere - ly at base adnate to petiole, triangular -lanceolate or linear lanceolate, acumi- nate, 8-12 cmlong, white-shaggy onthe margin and mostly onthe outside; leaves (6) 10-15 (18) cm long, the petiole 2-3 cmlong, both petiole and rachis shaggy with white hairs, rarely glabrate; leaflets distant, orbicular to obovate - orbicular, round-tipped, (14) 20-25 (30) mm long, glabrous above, densely shaggy beneath with long soft hairs; racemes sessile, spherical or ovaloid (ovaloid or rarely oblong in fruit) 3-5 cm long, 5-8 cm in fruit, densely 12-25-flowered; bracts linear-lanceolate, subulate-acuminate, white villous- ciliate on the margin, 8-10 mm long; bracteoles narrowly linear, 5-7 mm long, acuminate, ciliate-margined; flowering calyx campanulate or cylin- dric-campanulate, 12-14 mm long, the upper part and the teeth moreor less shaggy with white hairs, the teeth linear-subulate, 4-5 cm long, half as long as the tube; fruiting calyx spherical- to ovoid-bladdery, thinly membranous, profusely reticulate-nervose, glabrate except for shaggy pubescence at the throat and on the teeth, 15-20 mm long, the teeth linear from a triangular base, ca. 5 mm long; standard 15-20 mm long, the limb orbicular or sub- orbicular, round-tipped and retuse, as long as the claw; wing-petals equal- ing the standard, the limb oblong, round-tipped, somewhat shorter than the claw; keel 14-18 mm long, the limb rounded-convex beneath, straight above, obtuse, 2/3 as long as the claw; ovary on a stipe 2-3 mm long, glabrous; style naked; pods oblong-oval in side-view, 5-9 mm long, inflated, thinly coriaceous, carinate ventrally, grooved dorsally, borne on a slender stipe 3-9 mm long, terminating in a beak 1-2 mm long, bilocular to 2/3-3/4. F1. May-June; fr. June-July. Fixed sands and sand dunes. - Soviet Centr. Asia: Balkh. Endemic. De- scribed from the region between Arganaty and the source of the Sassyk- pastau and between Arganaty and Ayaguz river. Type in Leningrad. 380. A. leiophysa Bge. in A.H.P. III (1875) 104; Bge. Astrag. turk. 247. 307 Perennial, (30) 35-80 (110) cm tall; stems erect, firm, 18-70 cm long, 411 terete, finely sulcate, hairy (lanate at base) or rarely glabrous; stipules cauline, often partially adnate at base to petiole, triangular -lanceolate, acuminate, 8-12 mm long, shaggy or glabrous on the outside, ciliate-mar- gined; leaves 17-29 (30) cm long, the petiole 3-4 (5) cm long, both petiole and rachis hairy, rarely glabrous; leaflets (6) 7-10 pairs, orbicular to ovate-orbicular, round-tipped, (8) 10-21 mm long, glabrous above or rarely appressed-shaggy toward the margin; shaggy beneath with appressed soft hairs or glabrate except for faint pubescence mainly on the veins; racemes axillary, sessile, densely 17-many-flowered, 5-7 cm long; bracts lance - linear, green acuminate, 6-7 mm long, shaggy-margined; bracteoles re- sembling the bracts, smaller, ca. 4mm long; pedicels ca. 1 mm long; flow- ering calyx campanulate to short-cylindric, 12-13 mm long, softly shaggy or rarely glabrate with hairs confined to throat and the teeth, the teeth lance - linear, 3-4 mm long, half as long as the tube; fruiting calyx ovoid-bladdery, 15-18 mm long, thinly membranous, profusely reticulate-nervose, shaggy with short spreading pubescence, rarely sparsely pubescent or glabrate except for hairs at the throat and on the teeth, the teeth lance-linear from a triangular base, 4-5 mm long; corolla yellow sometimes fading red; standard 17-18 mm long, the limb ovate to rounded-ovate, round-tipped entire, about as long as or somewhat longer thanthe claw; wing-petals some - what exceeding the standard, the limb oblong, attenuate upward, round- tipped, as long as the claw; keel equaling the standard, the limb gibbous beneath, straight above, obtuse, 2/3 to as long as the claw; ovary on a stipe ca. 2mm long, glabrous; style naked; pods borne on a stipe 2-3 mm long, ovaloid, 6-7 mm long, inflated, carinate ventrally, grooved dorsally, thinly coriaceous, glabrous, terminating in a beak 1.5-2 mm long, bilocular to 2/3. Fl. May-June; fr. June-July. Sands.- Soviet Centr. Asia: Kyz. K., Syr D. Endemic. Described from Kyzyl Kum sands. Type in Leningrad. 381. A. chivensis Bge. Astrag. turk. (1880) 247.-Exs.: P. Sint. Iter transcasp.-pers. No. 340 (sub. nom. A. Winkleri Trautv.); H.F.R. No. 10; ed. Horti Bot. Petrop. No. 38. Perennial, (20) 35-60 (70) cm tall; stems firm, erect, simple, scabrous - sericeous with scattered hairs, (10) 20-40 cm long; stipules cauline, par- tially adnate to petiole at base, free, triangular acuminate, 6-8 mm long, green, glabrous, ciliate; leaves (14) 18-25 (35) mm long, the petiole rather indistinct, 3-6 cm long, both petiole and rachis covered with scattered spreading white hairs or shaggy with spreading white hairs, slightly indur- escent; leaflets (4) 5-7 pairs, mostly scarcely or slightly retuse, rounded or rarely broadly cuneate at base, (10) 15-25 (28) mm long, glabrous above, shaggy beneath with appressed white hairs, densely white -ciliolate on the 412 margin; racemes long, loose, remotely many-flowered, (10) 15-20 (25) cm long, the rachis shaggy with spreading hairs; bracts linear-lanceolate, finely pointed, 4-6 mm long, white-ciliate on the margin; bracteoles linear- lanceolate, 3-4 mm long, white-ciliate on the margin; pedicels 1-2 mm long, covered with spreading hairs; flowering calyx campanulate, 8-i0mm long, densely shaggy with long spreading silky hairs, the lanceolate teeth half the length of the tube, 3-5 mm long; fruiting calyx inflated, ovoid- bladdery, narrowed but not narrowly constricted at the throat, 14-20mm long, thinly membranous, lustrous, profusely reticulate-nervose, shaggy 308 with spreading white hairs, the teeth triangular-lanceolate; standard-limb ovate -orbicular, round-tipped or scarcely retuse, the claw 5-6 mm long; wing-petals about equaling the standard, the limb oblong, entire, round- tipped, somewhat longer than the claw; keel 11-12 mm long, the limb in- equilaterally triangular, gibbous beneath, straight or just concave above, obtuse, about as long as the claw; ovary glabrous, ona stipe 2-2.5 mm long; style naked; pod oval to oblong-oval (rarely oblong) in side-view, borne on a stipe 3-4 mm long, inflated, membranous, carinate ventrally, grooved dorsally, 6-7 mm long, terminating in a slender recurved beak ca. 3 mm long, glabrous, bilocular to 1/2-2/3, 2 (4)-seeded; seeds ovaloid to oblong - ovaloid, slightly notched, 3.5-4.5 mm long, compressed, brown, sometimes red-spotted. Fl. (April) May-June (July); fr. (May) June-July. (Plate XXVII, Figure 1). Flat, ridged or dune sands. - Soviet Centr. Asia: Kara K., Kyz. K. (area adjoining Amu D.), Amu D, (near river Amu Darya). Described from former Khiva (according to specimens collected by Korol'kov and Krause between Karakhaty and Adam-Kyr-Ulgan and Ginakchik). Type in Leningrad. Section 59. EREMOPHYSOPSIS Gontsch. sect. nova in Addenda XI, p.660.—Perennials; stems long, erect, vested with simple basifix white hairs; stipules cauline, free (not connate), adnate in lower part to petiole; leaves imparipinnate; leaflets many; leaf rachis becoming indurated but not spines- cent; racemes axillary, sessile, capitate; each flower subtended by a bract and 2 bracteoles; flowering calyx cylindric; fruiting calyx bladdery-inflated, thinly membranous, profusely reticulate-nervose; corolla persistent, pu- bescent; pods sessile, small, carinate ventrally and dorsally, coriaceous, unilocular, 1-seeded. A monotypic section represented by a single species in Kopet Dagh. The plant was earlier included in the section Eremophysa, but it differs markedly from the other representatives of that section inthe sessile uniloc - ular bicarinate pod and the pubescent corolla. A. sericopetalus Trautv. is a mountain-steppe plant, while all species of section Eremophysa are psammophytes. The section Eremophysopsis displays a close relation- ship to the section Alopecias, from which it is distinguished only by the bladdery-inflated calyx. 382. A. sericopetalus Trautv. in A.H.P. IX (1884) 448.-Exs.: P. Sint. Iter transcasp.-pers. No. 1809. Perennial, (25) 30-45 cm tall; stems firm, angled, softly pannose- velutinous, (15) 20-35 cm long, stipules triangular, 10-12 mm long, sub- hyaline, more or less covered on the outside with spreading hairs; leaves (10) 15-25 cm long, the petiole 1-2 cm long, both petiole and rachis densely covered with soft spreading hairs; leaflets 20-27 pairs, oval to oblong-oval, round -tipped and scarcely retuse, glabrous above, sparsely covered beneath with more or less spreading hairs, (5) 8-14 mm long; racemes axillary, few (up to 12)-flowered, dense, up to 3 cm long, borne on the upper 1/2-1/3 of the stem and gathered in a sort of a spiciform compound "inflorescence"; bracts linear-lanceolate, ca. 6 mm long, hyaline, the margin broadly hairy; bracteoles linear, hyaline, woolly on the outside, 3-6 mm long; pedicels ca. 1 mm long; flowering calyx cylindric, 13-15 mm long, woolly-velutinous with white hairs, the linear-subulate unequal teeth 5-6 mm long; fruiting calyx inflated, spherical-bladdery, 20-24 mm long, lanate with spreading hairs; corolla yellow; standard 23-26 mm long, the limb obovate, acuminate 309 414 and scarcely retuse, somewhat longer than the claw, sericeous on the out- side; wing-petals usually equaling the standard and the keel, the limb linear - oblong, round-tipped, more or less sericeous on the outside; keel gibbous and silky beneath, nearly straight and glabrous above, the claw 1.5 times as long as the limb; ovary sessile, sericeous; pods ovaloid, strongly com- pressed, 8-9 mm long, white-velutinous, the beak 1-3 mm long. Fl. May- June (July); fr. June-July. Mountain steppes. - Soviet Centr. Asia: Mtn. Turkm. (Centr. and W. Kopet Dagh). Endemic. Described from Kyzyl Arvat. Type in Leningrad. Section 60. HALICACABUS Bge. Astrag. geront. I (1868) 74; II, 123.-Acaulescent or subcaulescent xerophytic perennials, from an abbrevi- ated woody short-branched caudex, vested with white and (in infl. ) black hairs, pulvinate; stipules adnate to petiole; leaves paripinnate, the petiole commonly persistent but not spinescent; racemes loose or compact, spici- form, commonly elongate, borne on rather long peduncles; each flower furnished with a bract and 2 persistent bracteoles; flowering calyx cylindric; fruiting calyx bladdery-inflated, thinly membranous, profusely reticulate - nervose; corolla persistent, glabrous; pods small, unilocular, 1-4-seeded. - A Near Asian section, distributed from Syria and Asia Minor to Iran and penetrating into the USSR in the xerophytic regions of Transcaucasia, Da- gestan, Kopet Dagh and S.W. Pamir-Alai. 1. Limb of standard rounded or broadly cuneate at base .......... 2. + Limb of standard auriculately truncate (hastate) or attenuate and more or lesisrangularatibase Fic. Tie ype, hited = (ot 10 a toik =, 4 fhe is tes een ae 4. 2. Pod glabrate, except for sparse white hairs on the ventral side; leaf- letsséqlsepainsy lancéolatessacute cuspidiakey =iywel ter.) cnen teenie ee + Pod profusely puberulent; leaflets 15-20 pairs, or 10 pairs and then oblong-elliptic; yobtuse and cuspidateyima.i/: ns ieee Cae, eee Bu 3. Leaflets 15-20 pairs, lanceolate, rarely lance-oblong, acute and mucron- ULE Sy vaayut ae da atures Dern ee hla ese le Mie Whe 383. A. mesites Boiss. et Buhse. + Leaflets 10 pairs, oblong-elliptic, obtuse,,.mucronulate.s). .f.s) aeee eS eee oer Soha os eee PEN on 385. A. macrophysus Somm. et Lev. 4. Standard-limb short-attenuate and auriculate-angular at base .... 5. + Standard-limb auriculate-truncate (hastate) at base........... Ms 5. Leaflets 15-22 pairs; calyx covered with appressed predominantly black hairs, sometimes interspersed with occasional white hairs; bracts - lanceolate; keel greatly exceeded by ithe wings — % <.p ee seen SEAN sik he aOR RG. salle Sa Ral AAA is MPSSHUNba a oe ice ves 386. A. kaghysmani Gontsch. + Leaflets 8-16 pairs; calyx persistently pubescent; bracts ovate or lance- OVS! OM BWR ORR CN I EIN LE I VEE SS Ae St 6. 6. Leaflets (8) 10-12 pairs, glabrous above or nearly so (with occasional appressed hairs); racemes elongate, (7) 10-15 (17) cm long, loose, re- motely flowered; fruiting calyx shaggy with raised stiff white hairs, sometimes interspersed (esp. on the teeth) with black hairs........ Eo LEC ge Lomi eth aul te oN Me au cy ch 387. A. pseudoanthylloides Gontsch. + Leaflets 8-10 pairs, sparsely pubescent with spreading stiff hairs; racemes 3-4 cm long, ovoid to broadly ovoid, loose, with more or less approximate flowers, but not spiciform; fruiting calyx rather densely shaggy with spreading soft white hairs and sparsely pubescent with very SHOGEUSOUU ae ke mitettts So Mime en sere fe 388. A. karakushensis Gontsch. 415 416 touweahlersjovali-obovatelli: (uhewasan }da we 391. A. daghestanicus Grossh. + Leaflets oblong, elliptic or lance- eee TAGS Y OMA cya alice stein ouch 8. 8. Wings somewhat exceeding the standard. . 389. A. Raddei N. Basil. + Wings shorter than the standard........ 390. A. Willisii M. Pop. Series 1. ANGUSTATI Gontsch. Racemes loose, long, remotely flow- ered; limb of standard rounded or broadly cuneate at base; leaf rachises becoming indurated, but unarmed. 383. A. mesites Boiss. et Buhse in Nouv. Mém. Soc. Nat. Mosc. XII (1860) 66; Bge. Astrag. geront. II, 126; Boiss. Fl. or. II, 404; Grossh. Fl. Kavk. II, 301.-A. halicacabus C. Koch ex Ldb. Fl. Ross. I (1842) 643. Perennial, 23-35 cm tall, loosely cespitose, aculaescent; stipules con- nate and adnate to petiole to 1/3-1/2, rigidly membranous, glabrous or cov- ered with white hairs, white-ciliate on the margin, 10-14 mm long, (the free portion) lanceolate acuminate; leaves (15) 20-28 cm long, the petiole 5- 6 cm long, both petiole and rachis covered with scattered short white appressed hairs or glabrate, not becoming lignified, unarmed; leaflets 15-20 pairs, lanceolate, rarely lance-cblong, acute, mucronulate, glabrous or sparsely covered with short appressed white hairs above, scattered-hairy beneath, (7) 10-17 (21) mm long; peduncles somewhat shorter than the leaves (some- what exceeding them together with inflorescence), 15-21 cm long, sparsely covered with short appressed black hairs (mixed black and white at base); racemes 9-18 cm long, 15-25-flowered; bracts ovate, acuminate, 3-5 mm long, hyaline-membranous, covered on the outside with black hairs (interspersed or not with white), black-ciliolate on the margin; pedicels ca. 1 mm long, covered with white and predominantly black hairs; bracteoles lanceolate, ca. 3mm long, more or less covered on the outside and on the margin with black hairs; calyx initially short-cylindric, 8-10 mm long, becoming blad- dery-inflated, spherical, 16-18 mm long, thinly membranous, reticulate by numerous nerves, covered at first densely then sparsely with short ap- pressed black hairs interspersed with few white ones, the triangular-subu- late teeth 2-3 mm long; corolla pale purple; standard 19-29 mm long, the limb ovate, short-attenuate and retuse at apex, rounded at base, the claw 8-10 mm long; wing-petals 18-20 mm long, the limb oblong, entire, short- auriculate, the claw 10-12 mm long; keel15-16 mm long, the limb obtuse, rounded -convex beneath, slightly concave above, the claw 9-11 mm long; ovary on a stipe ca. 1 mm long, covered with short black and few white hairs; style hairy below, naked upward; pods oblong-ovaloid, 8-9 mm long, inflated, somewhat prominently sutured, terminating in a point ca. 1 mm long, thinly coriaceous, sessile, sparsely covered with black and (few at maturity) white hairs, 1-seeded; seeds ovaloid, slightly notched, ca. 4mm long, slightly compressed laterally, smooth. Fl. June-July; fr. July-Au- gust. (Plate XXVIII, Figure 1). Stony, gravelly, and clayey gravel mountain slopes. - Caucasus: S. and E. Transc. (Armenia, Nakhich. ASSR, Gandzha). Gen. distr.: Turk. Arm. Described from 8S. Transcaucasia. Type in Geneva. 384. A. halicacabus Lam. Dict. I (1789) 320; DC. Astrag. (1802) 182; Bge. Astrag. geront. II, 126; Boiss. Fl. or. Il, 404; Grossh. Fl. Kavk. SO: Perennial, from a woody branching caudex, glabrous or very sparsely vested with appressed stiff hairs; stipuleslanceolate, ciliate; leaves 8-15cm 311 417 -long, covered with black hairs; pedicels 2 mm long; fruiting calyx thin- long, the petiole half the length of the rachis; leaflets 8-15 pairs, lanceo- late, acute and mucronulate, 8-12 mm long, 2-3 mm broad; peduncles equaling to exceeding the leaves; racemes 8-10-flowered, loose; bracts lanceolate, equaling the pedicel; bracteoles acuminate-ovate, short; flow- ering calyx cylindric, 13 mm long, covered with short appressed stiff hairs; fruiting calyx membranous, spherical-inflated, glabrate, nervose, the teeth triangular, acute, very short; corolla yellowish (?); standard ca. 27 mm long, the limb oblong-ovate, obtuse, cuneate at base, slightly retuse at apex, exceeding the wings and the keel; limb of wings oblong; keel obtuse; pod stipitate, oblong, compressed, pointed at both ends, cuspidate, -labrous or ventrally very sparsely covered with white hairs, scarcely and partially bilocular, 2-4-seeded; seeds reniform, compressed. Fl. May-June. Recorded from the border regions of the USSR and likely to be discovered | in the future in Armenia. Gen. distr.: Turkish Armenia (vicinity of Erze- rum, Mt. Ararat). Described from ''the East''. Type in Paris. 385. A. macrophysus Somm. et Lev. in A.H.P. XIII (1893) 190. | Perennial, subacaulescent, from a woody caudex clothed in vestiges of petioles and stipules; stipules adnate to petiole in lower 1/3, free (not con- | nate) or connate merely at base, membranous, profusely reticulate -ner - vose, densely white-ciliate at base, the free portion narrowly lanceolate, acuminate; leaves 30 cm long, long-petiolate, unarmed; leaflets 10 pairs, | oblong-elliptic, up to 2 cm long, obtuse and mucronulate, glabrate above, sparsely covered beneath with appressed white hairs; peduncles 35 cm long, | somewhai exceeding the leaves, beset with white and shorter black hairs; | fruiting raceme 15 cm long, loosely 20-flowered, the rachis finely black- hairy; bracts ovate-lanceolate, acute or acuminate, up to 7 mm long, mem- branous, covered with brown hairs; bracteoles ovate-lanceolate, 3-4 mm ly membranous, inflated, spherical, 2.5 cm long, sparsely covered with appressed black hairs, the teeth triangular, acute, ca. 3 mm long; calyx pink (?); standard 18-22 mm long, the limb attenuate in upper part, bilo- bate, strongly enlargedat the middle, attenuate at base into the claw, this 10-12 mm long; wings shorter than the standard, exceeding the keel, the limb retuse; keel pale pink in dry condition, 3/4 as long as the standard, obtuse; pod ovoid-conic,. rounded at base, attenuate upward, 8-9 mm long, thickened ventrally, carinate dorsally, covered with short black hairs, uni- locular, 4-6-seeded, the densely black-hairy stipe 2 mm long. Fl. June. Caucasus: S. Transc. Endemic. Described from Karabakh. Type in Florence. Note. Judging from the description, very similar to A. mesites Boiss. et Buhse. Comparison with authentic specimens may disclose synonymy. i | Series 2. SUBAURICULATI Gontsch. -Racemes loose, remotely flow- ered; limb of standard attenuate and angular at base; leaf rachises at length — becoming slightly indurated, but not spiny or lignified, scarcely spinescent. — 386. A. kaghysmani Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. i Sc. URSS, IX, 3 (1941) 100. " Perennial, acaulescent, from a woody short-branched caudex, loosely cespitose, (8) 12-25 cm tall; stipules connate and adnate to petiole to 1/3, i" rigidly membranous, 10-13 mm long, the free portion lanceolate, long- acuminate, ciliate-margined; leaves (4) 10-20 cm long, the petiole (1) i 312 3-4 cm long, both petiole and rachis covered with subdivergent short white hairs; leaflets 15-22 pairs, elliptic, rarely oblong-elliptic, commonly cuspidate, 4-11 mm long, slightly canescent on both sides (more diffusely above) with short appressed whitish hairs; peduncles shorter than to rarely equaling the leaves, (3) 7-10 cm long, rather densely covered with some- what spreading short white hairs (black and fewer white beneath infl.); ra- cemes elongate, (5) 8-15 cm long, (6) 20-28-flowered; bracts lanceolate, acuminate, 4-6 mm long, thinly membranous, black-ciliolate on the margin; pedicels ca. 1 mm long, hairy; bracteoles lanceolate, acute, 2-3 mm long, hyaline, ciliolate-margined; flowering calyx short-cylindric, ca. 10 mm long; fruiting calyx inflated, ovoid, 13-15 mm long, thinly membranous, at first densely then sparsely covered with short black appressed hairs, (these sometimes interspersed withfew longer white hairs), profusely reticulate - nervose, the triangular -subulate teeth 1.5-2 mm long; corolla purple; standard 16-20 mm long, the limb ovate, retuse, truncate and angular -auriculate at base, as long asthe claw; wing-petals 10-18 mm long, the limb oblong, entire, short- auriculate, the claw 9-11 mm long; keel11-15 mm long, the limb inequilaterally obovate -triangular, rounded-gibbous beneath, straight or slightly convex above; Ovary ona stipeca. 1mm long, covered with short white andblackhairs; style naked, curved in upper part. Fl. May-June. Stony and clayey stone-covered mountain slopes. - Caucasus: S. Transc. 418 (Armenia). Gen.distr.: Turkish Armenia. Described from Turkish Arme- nia (Kagyzman). Type in Leningrad. 387. A. pseudanthylloides Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Se. URSS IX 3 (1941) 101. Perennial, acaulescent, 15-25 cm tall, from a short branched buried caudex, the ligneous branched 3-10 (13) cm long, densely clothed at the end with remnants of petioles and stipules, forming large loose or loosish tufts; stipules 6-10 mm long, rigidly membranous, hyaline-bordered in upper part, whitish, adnate to petiole and connate to 1/3-1/2, covered at base with stiff white hairs, more or less white-ciliate on the margin, the lower ovate acuminate, the upper linear-attenuate from an ovate base, acute; leaves (4) 6-12 cm long, the petiole obsolescent, the rachis more or less white-seri+ ceous with appressed hairs; leaflets (8) 10-12 pairs, elliptic, rarely ob- ovate, mucronulate, stiff, veined above, cinereous, covered beneath with appressed white or silky hairs, (3) 4-9 (12) mm long, subdistant; peduncles equaling to exceeaing the leaves, 6-14 cm long, shaggy with short white hairs, these sometimes interspersed with short black hairs; racemes loose - ly and remotely (8) 18-25-flowered, (7) 10-15 (17) cm long, the rachis shag- gy with mixed white and black spreading short hairs; bracts ovate to lance- ovate, (3) 5-7 mm long, white-hyaline, acuminate, sparsely covered on the outside with white or rarely black hairs; bracteoles ovate or lanceolate, acute, 2-2.5 mm long, more or less covered with white hairs; flowering calyx cylindric, 9-12 mm long, densely shaggy with white silky hairs, the triangular teeth 1.5-2 mm long; fruiting calyx bladdery-inflated, spherical or ovoid-spherical, 13-18 mm long, whitish or purple-tinged, shaggy with spreading stiff white hairs (these sometimes interspersed, mainly on the teeth, with few black hairs), constricted at the throat, the triangular acu- minate teeth 2-3 mm long; corolla white, faintly pinkish; standard 14-18 mm long, the limb rounded-oval, retuse, the claw 6-9 mm long; wing-petals equaling the standard, the limb oblong-obovate, entire, short-auriculate, the claw 9-11 mm long; keel 13-16 mm long, the limb rounded-convex 313 419 420 beneath, scarcely convex above, obtusish, the claw 9-11 mm long; ovary on a very short stipe, densely covered with appressed white hairs; style long, the lower half covered with short hairs. Fl. May; fr. June. Stony and clay-and-gravel slopes of elevated foothills, the submontane and lower mountain zones, often in juniper groves. - Soviet Centr. Asia: Pam.-Al. (W., in districts of Baisun and Denau). Endemic. Described from the vicinity of Baisun. Type in Leningrad. 388. A. karakuschensis Gontsch. in Not. Inst. Bot. Tphil. 10 (1941) 87, cum ic. Perennial, dwarf plants, ca. 10 cm tall, from a strong woody much- branched root-crown and short buried woody caudices; stems prostrate, ligneous, clothed in old brown stipules and vestiges of petioles, 2-3.5 cm long; stems of the year obsolescent; stipules adnate to petiole to 1/3-1/2 and connate in lower part, forming an amplexicaul sheath 7-9 mm long, thinly but rigidly membranous, the free portion lanceolate, acuminate, glabrous, yellowish, ciliate-margined; leaves (1.5) 2-3 cm long, the petiole much shorter than the rachis, both vested with stiff spreading pubescence; leaf- lets 8-10 pairs, ellipticor obovate, 3.5-5 mm long, rounded-obtuse, rarely scarcely retuse, canescent sparsely above and densely beneath with spreading stiff pubescence; peduncles exceeding the leaves, 3-5cm long, densely shaggy with spreading white hairs and pubescent withvery short blackhairs; racemes short, ovoid to spherical-ovoid, 3-4 cm long, loosely 5-8-flowered, the flowers approximate (but not in a spiciform arrangement); bracts ovate, 6-8 mm long, acute, shaggy with white and black spreading hairs; pedicels ca. 1mm long, covered with spreading black and white hairs; calyx early inflated, ovoid in flower and in fruit, 15-18 mm long; fruiting calyx thinly membranous, densely shaggy with spreading soft hairs and sparsely pubes- cent with very short black hairs, the nerves mostly red-tinged, the teeth linear-lanceolate, acuminate, 3-5 mm long; corolla whitish, finally be- coming red; standard ca. 23 mm long, the limb rounded oblong -rectangular, 15 mm long, 10 mm broad, faintly and narrowly emarginate, narrowed in lower one-third to 8 mm between the parallel margins, auriculate-angular at base; wing-petals 22 mm long, the limb oblong, slightly enlarged and rounded at apex, somewhat shorter than the claw; keel 18 mm long, the limb inequilaterally obovate, truncate but point-tipped, the claw 2.5 times as long as the limb. Fl. May (June). ; Stony limestone mountain slopes.- Caucasus: S. Transc. (near Buzgov woodland and vicinity of Nakhichevan). Endemic. Described from vicinity of Nakhichevan. Type in Baku. Series DENSIFLORI Gontsch. -Racemes compact, capitate; standard- limb truncately hastate and angular-auriculate at base; leaf rachises at length lignified and subspinescent. 389. A. Raddei N. Basil. inNot. Syst. exHerb. H.B.P. IV, 6 (1923)116. Perennial, 9-25 cm tall, from a woody short-branched caudex, the branches clothed in indurated persistent petioles and vestiges of stipules, forming compact tufts; stipules adnate to petiole and connate to 1/3-1/2, 6-12 mm long, rigidly membranous, white-hairy at base and ciliate-mar- gined, the free portion lanceolate, long-acuminate; leaves (3) 5-14 cm long, the petiole 1-2.5 cm long, both petiole and rachis canescent; leaflets 10- 16 pairs, oblong, less often elliptic or rarely oval, canescent on both sides 314 423 with subappressed white hairs, mucronulate, somewhat indurated, (3) 4-9 mm long; peduncles (infl. excluded) 2-3 times as long as the leaves, pubescent with white hairs (mixed white and black above the infl.), (9) 12-24 cm long; racemes compact, capitate-ovoid, 4-6 cm long, rather many-flowered; bracts lanceolate, acuminate, 5-6 mm long, thinly mem- branous, covered on the outside with white and black hairs; pedicels ca. 1mm long, hairy; bracteoles linear, half as long as the calyx, hairy on the outside; flowering calyx cylindric, ca. 11 mm long; fruiting calyx inflated, 13-19 mm long, spherical or ovoid, more or less tinged with purple, reticulate-nervose, shaggy with soft white hairs, the-triangular acuminate teeth (2) 3-4 mm long; corolla purple; standard 14-18 mm long, the limb rounded-square, nearly entire at apex, round-tipped, truncate- hastate and short-auriculate at base, the claw 8-10 mm long; wing-petals 15-19 mm long, somewhat exceeding the standard, the limb entire, obovate minutely auriculate at base, the claw 9-13 mm long; keel 14-18 mm long, somewhat shorter than the wings, the limb inequilaterally obovate, obtuse, strongly convex beneath and slightly so above, entire, the claw 10-13 mm long; ovary on a very short stipe, white-hairy; style long, curved at the end, often sparsely hairy at base; pods ovoid, ca. 5 mm long, unilocular, rigidly membranous, 2-seeded, covered with short appressed hairs, ter- minating in a point ca. 1 mm long, slightly obcompressed, oblong in side view; seeds round to rounded-oval in outline, compressed, 3-3.5 mm long, distinctly notched, tawny. Fl. June-July; fr. July-August. (Plate XXVIII, Figure 2). Mostly stony steppe mountain slopes, at altitudes of 1950-2600 m. - Soviet Centr. Asia: Mtn. Turkm. (Centr. Kopet Dagh from Gaudan to Kheirabad, rarely W.). Gen. distr.: N. Iran. Described from Kopet Dagh. Type in Leningrad. 390. A. Willisii M. Pop. in Not. Syst. ex Herb. H.B.P. IV, 19-20 (1923) 155. Perennial, acaulescent, from a branched caudex, the branches stout, 4-10 cm long, densely clothed in vestiges of petioles and leaves, bearing at the ends leaf tufts and peduncles; stipules large, rigidly membranous, prominently nerved, adnate to petiole and connate to the middle, the free portion lance -acuminate, densely pannose at base, upward glabrous or ciliate-margined; petiole (including leaf rachis) 7-20 cm long, subceri- ceous -shaggy with subappressed hairs, the petiole proper much shorter than the rachis; leaflets very often alternating, 9-18 (19-37) pairs, elliptic or lance-oblong, conduplicate at apex, mucronate, 4-10 mm long, somewhat prominently veined, readily deciduous, the terminal leaflet mostly narrower and smaller; peduncles erect, equaling to exceeding the leaves, 7-20 cm long (excluding infl.), shaggy with subdivergent white hairs (chiefly black beneath the raceme); racemes compact, ovoid or more often plump-cylin- dric, 4-7 cm long; bracts oblong; bracteoles linear, less than half the length of the calyx; calyx soon becoming inflated, at length bladdery, ovoid to ovoid-spherical, thinly membranous, reticulate-nervose, whitish or purple-tinged, densely covered with ascending white hairs (often black on the teeth), 14-16 mm long in flower, up to 20 mm in fruit, the teeth lanceo- late, half as long as the tube; corolla violet or whitish (?); standard ca. 20 mm long, truncately and angularly hastate at base; wing-petals exceeded by the standard, 19 mm long; keel 14-16 mm long; podsunilocular, strongly 315 "a a = A SS \Naec= 7 JZ —S Sf PED J ABRAMS Ss SS SZ = Z \S N. \ ‘ A> PLATE XXVIII 1. Astragalus mesites Boiss. et Buhse.—2. A. Raddei N. Basil. 316 424 compressed laterally, oblong in side view, obliquely cuspidate-pointed, 8 mm long, commonly 1-2-seeded (ovary 6-8-ovuled), densely appressed - pubescent, subsericeous. Fr. May. Stony mountain slopes. - Soviet Centr. Asia: Pam.-Al. (S. W.-Kugitang Mountains). Endemic. Described from Kugitang. Type in Tashkent. 391. A. daghestanicus Grossh. Fl. Kavk. II (1930) 301. Perennial, nearly acaulescent plants, densely white-pubescent; leaflets 15-25 pairs, 5-6 mm long, pubescent on both sides, strongly so beneath; stipules hairy, adnate to petiole to the middle, lanceolate, acute; racemes ovaloid, 5-15-flowered, the peduncles exceeded by the leaves; calyx strong- ly inflated in fruit, 20 mm long, pale, reticulate-nervose, densely hairy, the tube 3 times as long as the teeth; corolla purple (?); pods unilocular, 1-seeded. June. Dry stony slopes. - Caucasus: Dag. Endemic. Described from Daghestan. Type in Baku. Note. This species is included in the series Densiflori conditional- ly, as it is known only in fruiting condition and its description is incomplete. Section 61. GRAMMOCALYX Bge. Astrag. geront. I (1868) 62. - Low perennials; stems short; caudices buried, often elongated, decumbent, slender; pubescence consisting of simple hairs; stipules not adnate to peti- ole or slightly so at base, free or connate; leaves imparipinnate, the petiole marcescent; racemes capitate, compact, borne on a short axillary peduncle; bracteoles none; calyx at first cylindric, at length bladdery-inflated; corolla glabrous, persistent; limb of standard exauriculate, attenuate at base into the claw; pods coriaceous, bilocular, few-seeded, included in the calyx. - Distributed in the mountains of Asia Minor, Armenia, and Iran. In the USSR known only from Southern Transcaucasia. 1. Bracts lanceolate to linear-lanceolate, 8-11 mm long; calyx shaggy with spreading soft yellow or rufous hairs, the teeth covered with few VG y ALIS bac onthe chastened chest uous 392. A. grammocalyx Boiss. et Hohen. + Bracts ovate, more or less acuminate, 4-8 mm long; calyx lanate with short black and white hairs, in fruit sparsely shaggy with spreading StidishysShortpblack andswhitey hai Siwe. pei fal e\e lols. ee ucy.s 9) eye)! © Ise. 2. 2. Calyx covered with spreading hairs; wing-petals entire ......... MNT Wes oa cis “Siemens cya cuisy ooh spe mcy iss # faa) sates 393. A. sphaerocalyx Ldb. + Calyx covered with ascending hairs; wing-petals slightly retuse..... REET gee oi) wrod met tein oe sdaey os se erect opae wee: shi 394. A. gezeldarensis Grossh. 392. A. grammocalyx Boiss. et Hohen. in Boiss. Diagn. Ser. I. IX (1849) 52; Bge. Astrag. geront. II, 107; Boiss. Fl. or. II, 421.-Exs.: Th. Kotshy Pl. Pers. bor. Ed. R.F. Hohenacker (1846) No. 320. Perennial, nearly acaulescent, dwarf plants, 6-9 cm tall, the strongly abbreviated buried woody caudex divided into slender woody underground branches 6-15 cm long; stems 1-2 cm long, densely pannose-lanate with soft white hairs; stipules adnate at base to petiole and connate in lower part into an amplexicaul sheath, ovate-triangular or rarely lanceolate -triangular, acuminate, 5-7 mm long, green, shaggy with white spreading hairs and covered with few black hairs at apex; leaves 2-5 cm long, the petiole much shorter than the rachis, both densely shaggy with short spreading white or rufous hairs; leaflets 10-15 approximate pairs, oblong-oval, round-tipped, oT 425 426 (3) 4-8 mm long, (2) 3-5 mm broad, shaggy on both sides (densely beneath) with ascending white or rufous silky hairs; peduncles equaling the leaves, densely pannose- shaggy with spreading white or rufous hairs; inflorescence ovoid, 3-5 cm long, containing 9-15 subsessile flowers; bracts lanceolate to linear-lanceolate, 8-11 mm long, acuminate, densely shaggy with soft black hairs, these interspersed with few white hairs; calyxat first cylindric, early inflated, ovoid, 12-18 mm long, shaggy with long spreading white or rufous hairs and bearing a few black hairs on the teeth, the teeth filiform - linear, 4-8 mm long; corolla violet; standard 18-22 mm long, the limb ovate, subligulately short-attenuate upward, notched-bilobate at apex, near- ly round at base, the claw equaling to 1.5 times as long as the limb; wing- petals 15-19 mm long, the oblong limb as long as the claw; keel 13-16 mm long, the limb nearly semicircular, acutish, half as long as the claw; ovary on a stipe ca. 1 mm long; pods (imperfectly mature) ovaloid, ca. 7 mm long, carinate ventrally, grooved dorsally, densely shaggy with rufous or white spreading soft hairs. Fl. May-June; fr. June-July. ; Stony limestone mountain slopes.- Caucasus: S. Transc. (vicinity Nakhi- chevan: between Aznabyurt wcodland and Buzgov). Gen. distr. : mountains of N. Iran (El'burs Range). Described from Mt. Demavend foothills. Type in Geneva; cotype in Leningrad. 393, A. sphaerocalyx Ldb. Fl. Ross. I (1842) 647; Bge. Astrag. geront. II, 107; Boiss. Fl.or. II, 422. Perennial, nearly acaulescent plants (stems rarely up to 5 cm long), the fairly numerous caudices slender, prostrate, up to 5 (15) cm long, forming loose tufts 5-9 (11) cm tall; stipules connate to high up, amplexicaul, ca. 4mm long, greenish, white-hairy onthe outside, the free portiontriangular, acutish; leaves (1.5) 2-4 (7) cm long, the petiole white-shaggy, 0.3-0.7 (2) cm long; leaflets oval to oblong, round-tipped, 8-12 pairs, 3-5 (10) mm long, covered on both sides with white silky hairs, rarely faintly pubescent above; peduncles (1) 1.5-3 (8) cm long, shaggy with short white hairs inter - spersed beneath the infl. with black hairs; inflorescence 2.5-3 cm long, 4-7-flowered; bracts ovate, acuminate, 4-8 mm long, covered with white and black hairs; pedicels ca. 1 mm long; calyx at first 11-13 mm, at length 12-14 mm long, ovoid to ovoid-spherical, more or less purple -tinged, shag- gy with black and white spreading hairs, the teeth subulate, 3 mm long; corolla azure; standard (16) 20-25 mm long, the limb ovate, broad and re- tuse at apex, round or broadly cuneate at base; wing-petals (14) 17-19 mm long, the limb oblong, entire at summit, somewhat longer than the claw; _ keel (11) 15-17 mm long, obtuse; ovary on a very short stipe, white -hairy; style naked; pods ovoid, ca. 10 mm long, ona stipe ca. 1 mm long, ter- minating in a beak ca. 1 mm long, grooved ventrally, carinate dorsally, white-shaggy. Fl. July-August; fr. August. Slopes and taluses in the high-mountain zone. - Caucasus: S. Transc. (Armenia). Gen. distr.: Turkish Armenia. Described from Ararat. Type in Leningrad. 394. A. gezeldarensis Grossh. in Beih. Bot. Centralbl. XLIV, II (1927) 219. Closely resembling the preceding species; differing in the ascending pubescence of the calyx and the slightly retuse wing-petals. Rocks. - Caucasus: S. Transc. (Nakh. A.S.S.R.). Endemic. Described from Gezel'dara range in Nor Bayazet district. Type in Leningrad. 318 Section 62. MEGALOCYSTIS Bge. Astrag. geront. I (1868) 72; II, 120. - Acaulescent or subcaulescent perennials, rarely tufted subshrubs; stipules adnate to petiole and mostly connate at base; leaves paripinnate, terminating in a spine, the petiole persistent, becoming indurated or rarely lignified; peduncles mostly exceeding the leaves, scapiform, bearing a loose or compact spikelike raceme; each flower furnished with a bract and 2 persistent bracteoles; calyx at first cylindric, becoming bladdery-inflated, thinly membranous, the teeth much shorter than the tube; corolla glabrous, persistent; limb of standard hastate or angular-auriculate rarely rounded at base; wing-petals entire; pods solitary, 1-4-seeded. y Species of this section are chiefly distributed in the mountains of Iran and, toalesserextent, inSyria and Mesopotamia. They are typical mountain xerophytes; in the subalpine and alpine regions they form a series (Sub - pulvinares Gontsch.) of subpulvinate tufted frutescent species. 1. Dwarf, loosely cespitose, subpulvinate subshrubs; branches prostrate, ligneous; leaves 2-4.5 cm long; leaflets densely cinereous -pannose on both sides; racemes capitate, reddish, 5-7-flowered............ Sia QROUME O) <0 eee WGP Shc Oh Capea HAT TAA AREND ag aamnGU SP Un Coa an 396. A. micracme Boiss. + Perennials of medium height, nearly acaulescent; leaves 7-42 cm long; leaflets covered on both sides with appressed white hairs; racemes mianv-T Lowered jicontractedvormilOOSe, wSPLCITOFM.: oo) se) cieiievor el st.cus @ < ZAG 2. Racemes loose, 20-35 cm long, remotely flowered; leaflets (4) 15- 18 mm long; limb of standard more or less attenuate and slightly angu- PEO MAT CMD ES Cais icn “hee poipeuitsiarin shes) «othdsibas 395. A. Fuhsii Freyn et Sint. + Racemes contracted, compact, capitate, ovoid or broadly oblong, 3- 7 mm long; leaflets 2-4 mm long; limb of standard hastate -auriculate PUNE S Oe aaa ci MIL iba Shoda Ga oa a 397. A. Szovitsii Fisch. et Mey Series 1. LAXIFLORI Gontsch. - Racemes somewhat compact or capi- tate, but always fairly loose; standard more or less rounded or attenuate and slightly angular at base; nearly acaulescent perennials. This series, containing the largest number of species, is widely dis- tributed in the dry mountains of Iran and comprises one species in Meso- potamia. 395. A. Fuhsii Freyn et Sint. In Bull. Herb. Boiss. 2 sér. IV (1904) 1114.-Exs.: Sint. Iter transcasp.-pers. No. 739. 427 Perennial, nearly acaulescent plants, 25-70 cm tall, loosely cespitose from a woody branched caudex; stipules adnate to petiole up to the middle, free, 14-22 mm long, subcoriaceous, nervose, ciliate-margined, the free portion narrowly triangular, long-acuminate, scarious-margined; leaves 20-42 cm long, the petiole (4) 5-15 cm long, sparsely covered with short white subappressed hairs; leaflets 12-17 pairs, obovate or elliptic, rarely oblong-obovate, (4) 15-18 mm long, cuspidate; peduncles ca. 22 cm long, rigid; racemes loose, remotely flowered, 20-35 cm long, sparsely covered with short subappressed white hairs; bracts subulate-lanceolate, acuminate, 6-9 mm long, covered with short black hairs; bracteoles lanceolate, cov- ered with short black hairs, 3-4 mm long; calyx at first 14 mm long, dense- ly and roughly velutinous with white hairs, at length ovoid inflated, 20- 25 mm long, covered with scattered white appressed hairs, the subulate- triangular teeth 5 mm long; corolla pink; standard 25-28 mm long, the limb broadly obovate, deeply retuse, more or less attenuate in lower part, slightly 319 428 angular at base, 1.5 times as long as the claw; wing-petals 25-26 mm long, the limb oblong, enlarged upward and obliquely truncate, somewhat shorter than the claw; keel equaling the wings, obtuse; ovary short-stipitate, silky; style hairy at base, naked upward; pods oblong, ca. 10 mm long, borne on a stipe 2 mm long, grooveless, pubescent with white appressed hairs, 4- seeded; Fl. June-July; fr. August. Gravelly wormwood-covered mountain slopes. - Soviet Centr. Asia: Mtn. Turkm. (Kopet Dagh: vicinity of Sulyuklyu and Prokhladnoe). Endemic. Described from vicinity of Sulyuklyu. Type in Geneva. cotype in Leningrad. Series 2. SUBPULVINARES Gontsch. -Racemes capitate, open; standard more or less rounded or broadly cuneate at base; low, cespitose, subpul- vinate subshrubs. In addition to A. micracme Boiss. which occurs in the USSR, this series includes A. coluteoides Willd. and A. murinus Boiss. It is associated with the high-mountain zone (alpine and subalpine vegetation belts) in Armenia, Kurdistan, Iran, and Lebanon. 396. A. micracme Boiss. Diagn. ser. 2, No. 5 (1856) 87; Bge. Astrag. geront. II, 121; Boiss. Fl. or. I, 398; Grossh. Fl. Kavk. II, 299. A dwarf, subpulvinate, suffrutescent perennial, 5-8 cm tall; branches ligneous, prostrate, clothed in divergent old spines; shoots of the year 1 or rarely 2 cm long; stipules connate to high up, white -pubescent, the lower ones glabrous; leaves approximate, triangular, free [?], the petiole 0.5- 2 cm long, both petiole and rachis pubescent, becoming lignified, termi- nating in a slender spine, this greatly exceeding the terminal leaflets; leaf- lets approximate, 7-10 pairs, conduplicate, obovate, orbicular, or broadly oval, subobtuse, (1) 2-4.5mm long, covered onboth sides with appressed white hairs; peduncles equaling to exceeding the leaves, pubescent; racemes capi- tate, rather loosely 5-7 flowered; bracts half as long as the calyx; bracteoles lanceolate; calyx cylindric, at length ovoid-inflated, ca. 18 mm long, more or less shaggy with gray hairs, the teeth subulate from a lanceolate base, the tube 4 times as long as the teeth; corolla about twice the length of the calyx; standard-limb attenuate upward, retuse; wing-petals somewhat ex- ceeded by the calyx, the limb oblong, round-tipped; keel obtuse, the limb gibbous beneath, concave above, about half as long as the claw; pods borne on a very short stipe, ovoid-spherical, subcompressed, membranous, cov- ered with short hairs. July. Stony slopes of the subalpine high-mountain zone. - Caucasus: S. Transc. (between Erevan and Kars). Gen. distr.: Turkish Armenia, Kurdistan. Described from the area between sree and Kars and from the surround- ings of Lake Van. Type in Geneva. Series 3. DENSIFLORI Gontsch.- Nearly acaulescent perennials; ra- cemes contracted, compactly capitate, ovoid or broadly oblong; standard hastate-auriculate at base. The series contains a single species. 397. A. Szovitsii Fisch. et Mey. in Fisch. Syn. Tragac. (1853) 160; Bge. Astrag. geront. II, 73; Boiss. Fl. or. II, 401; Grossh. Fl. Kavk. II, 301.-Exs.: Herb. Fl. Cauc. No. 427. Perennial, 14-30 cm tall, from a short-branched woody caudex; stipules 8-9 mm long, glabrous above, the free portion lanceolate, acuminate, 320 ciliate-margined, rigidly membranous, prominently nerved on the outside; leaves 7-18 mm long, the petiole (2) 3-5 cm long, both petiole and rachis indurescent, strongly spinescent, beset with short appressed hairs; leaf- lets 15-25 pairs, ovate or broadly oval, round-tipped, 2-4 mm long, thick- ish; peduncles somewhat shorter than to exceeding the leaves, (6) 10- 25 cm long, rigid, covered with short white appressed hairs; racemes con- tracted, capitate, ovoid or broadly oblong, 3-7 cm long; bracts lanceolate, acuminate, 4-5 mm long, white-hairy on the outside; bracteoles narrowly linear, very small; calyx becoming spherical-inflated, more or less purple - tinged, 12-14 mm long, covered with scattered lurid hairs, the lance -subu- late teeth 3-4 mm long; standard 16-17 mm long, the limb ovate, slightly retuse, hastate-auriculate at base, a trifle shorter than the claw; wing- petals equaling the standard, the limb oblong, round-tipped, the claw 9- 10 mm long; keel 15-16 mm long, the limb obtuse, gibbous beneath, very slightly concave above, the claw 10 mm long; ovary on a very short stipe, covered with appressed white hairs; style naked; pods borne on a very short stipe, membranous, ovoid, slightly compressed, appressed-hairy, 1-seeded. Fl. May-June. Dry hills.- Caucasus: S. Transc. (along the river Nakhichevanchai). Endemic. Described from Nakhichevanchai river valley. Type in Leningrad. Section 63. TRICHOLOBUS Bge. Astrag. geront. I (1868) 69. - Branched subshrubs, spiny-pulvinate; stipules adnate to high up with petiole, connate in lower part, vaginately amplexicaul, membranous, glabrous; leaves paripinnate, the petiole and rachis becoming lignified, terminating in a spine; racemes compactly capitate, borne on a long scapiform peduncle; bracts large, scalelike; bracteoles none; calyx at first campanulate, usual- ly becoming insignificantly inflated; corolla persistent, glabrous; standard- limb ovate or obovate; wing-petals entire; pods subbilocular, dorsally con- cave. A section associated with xerophytic mountain regions of W. and N. Iran; occurring in the USSR in the extreme southeast of Transcaucasia. 398. A. Hohenackeri Boiss. Diagn., sér. I, 2 (1843) 70; Bge. Astrag. eeront. If, 117; Boiss:,Fl. or! Il, 386; Grossh. Fl: Kavk. II, 298. —- A. tricholobus fp. minor Hohenack. Enum. pl. Talysch. (1837) 109; Ldb. Fl. Ross. I. 642. Perennial, strongly branched, spiny, pulvinate, suffrutescent, 10-17cm tall, the old lignified caudices densely clothed in divergent persistent spines and stipules, the strongly abbreviated shoots of the year not more than 1 (rarely up to 2) cm long; stipules 7-9 mm long, the free portion lanceolate, acuminate; leaves 1.5-3 cm long, the petiole and the rachis indurated, cov- ered with short appressed white hairs; leaflets 5-7 pairs, linear-lanceo- late, mucronulate, 3-7 mm long, conduplicate, glabrous or Sparsely cov- ered with short white hairs; peduncles 2-3 times as long as the leaves, slender, 4-6 cm long, covered with short appressed white hairs; racemes capitate, spherical, dense, 1.5-2 cm long; bracts ovate, acuminate, gla- brous, membranous, 7-9 mm long, subcarinate; deciduous; pedicels ca. 1 mm long, white-shaggy; calyx campanulate, becoming slightly inflated, 12-14 mm long, white-shaggy, the teeth setaceous, longer than the tube and somewhat shorter than to equaling the standard, 7-9 mm long; corolla purple; standard 14 mm long, the limb ovate, obtusish, round-based, the 321 430 433 claw 2-3 mm long; wing-petals 12-13 mm long, the limb oblong, round- tipped, the claw ca. 4 mm long; keel 11-12 mm long, the limb convex be- neath, straight above, obtuse, the claw 3-4 mmlong; ovary subsessile, white-shaggy; style naked. June-July. (Plate XXIX, Figure 1). Dry stony mountain slopes. - Caucasus: Tal. Gen. distr.: mountains of N. and W. Iran. Described from Talysh (vicinity of Zuvant Pass). Type in Leningrad. Section 64. HYMENOSTEGIS Bge. Astrag. geront. I (1868) 66; II, 108.-Cespitose small shrubs, devoid of a distinct stem of the year; leaf petioles and rachises abbreviated, becoming lignified, or branched spiny, vested with simple white hairs; stipules adnate to petiole to 1/3-1/2; leaves paripinnate, the rachis spinescent, persistent after the shedding of leaflets as rigid spines on the branches; inflorescences compactly spiciform, ovoid, or oblong, borne on a long strongly abbreviated axillary peduncle; bracts large, indurated, thinly coriaceous or soft subhyaline, caducous; bracteoles none; calyx at first short-cylindric, very slightly inflated, soon becoming ovoid-inflated, constricted at the throat; corolla persistent, glabrous; stand- ard straight, obtusely or acutely and angularly auriculate at base, sub- pandurate in outline, Pods sessile, soft, compressed, included in the calyx, unilocular, rarely bilocular, the valves scaphoid. About 25 species, distributed in N. and W. Iran, Armenia, Talysh Moun- tains, and rarely in Mesopotamia. Xerophytes, associated with dry, mostly stony, mountain slopes. A recent group, differentiated into minor geo- graphical races. 1. “Bracts hyaline, (small) nearly translucent 0.iicae 3 sek eels eee UL TRE Naha aT MERRIE My i SATE TREE | AN A TER ie 399. A. uraniolimneus Boiss. +. Bracts'thinly coriaceous, ‘rigidsvyellowishyn hilw 2s eee 2h Bracts broad, sharply long-acuminate, exceeding the calyx, equaling to somewhat exceeding the corolla, sericeous on the outside over the whole surface or merely on the midrib; calyx equaling the corolla or nearly so, the teeth as long as the tube ..... 401. A. lagurus Willd. + Bracts narrower than the calyx, exceeded by the corolla and the calyx; calyx markedly exceeded by the corolla, the tube 1.5-2 times as long BIS NS USSU, os) a alic. co loka ca dlonas Bho yond 400. A. persicus Fisch. et Mey bo Series 1. SUBSCARIOSI Gontsch.-Bracts hyaline, nearly translucent. 399. A. uraniolimneus Boiss. Fl. or. II (1872) 380; Grossh. Fl. Kavk. II, 300.-A. lagopoides Bge. Astr. geront. II (1869) 111, ex parte, non Lam. Perennial, cespitose, pulvinate, suffrutescent, 7-13 cm tall; branches abbreviated, ligneous, densely clothed in slender divergent spines; stipules hyaline, 9-14 mm long, densely ciliate on the margin, the free part ovate- lanceolate, acuminate, or[?]lanceolate; leaves 2-3.5cm long, the petiole 0.5- 1 cm long, both petiole and rachis pubescent with spreading hairs; leaflets 9-6 pairs, narrowly elliptic or short-lanceolate, cuspidate, 5-10 mm long, covered on both sides with ascending silky hairs; peduncles 1-4 (4-5) cm long, shorter than to equaling the leaves; inflorescence largely many-flow- ered, ovoid, 2.5-4 cm long; bracts hyaline, soft, nearly translucent, the lower broadly ovate, the upper oblong-ovate, acuminate, densely long- 322 ( wea) i Wy, pies Hi fi \ p s SLA, —=s— Zo ¢ —. ~~ 7. o— e — 7g = a, >= Le ROM —\ Nit SS a iY = A ‘Z D 6, Se eS, > —he as, WSs ~~ = \ é Z y LI / y ox A = I Nb tg, Fan eae Eye Se ee Za QA PLATE XXIX 1. Astragalus Hohenackeri Boiss.—2. A. persicus Fisch. et May 323 434 ciliate on the margin, 7-10 mm long; calyx softly shaggy, at first cylindric- campanulate, 11-18 mm long, becoming ovoid-inflated, the teeth subulate, as long as the tube; corolla purple; standard ca. 15-23 mm long, the limb broadly pandurate-oblong, obtusely angular-auriculate at base, round- tipped, the claw 2-4 mm long; wing-petals exceeded by the standard, 14- 19 mm long, the limb oblong, faintly ciliate at apex, round-tipped, some- what shorter than the claw; keel 12-15 mm long, the limb rounded -gibbous beneath, nearly straight above, obtusish, the claw 1.5-2 times as long as the limb; ovary subsessile, densely shaggy; style hairy at base, glabrous upward. Fl. July-August. Gravelly and stony southern mountain slopes, at altitudes of 2400-3000m. Caucasus: S. Transc. Gen. distr.; Turkish Armenia. Described from Armenia (Alagez and the shores of Lake Gokcha). Type in Geneva. Series 2, CORIACEI Gontsch.-Bracts thinly coriaceous, rigid. 400, A. persicus Fisch. et Mey. in Hohenacker Enum. Talysch. (1837) 109, nomen; Bge. Astrag. geront. Il, 113; Boiss. Fl. or. i, 382; Grossh: Fl. Kavk. II, 300. Perennial, dwarf, spiny, suffrutescent-cespitose, 8-13 cm tall; branches strongly abbreviated, ligneous, densely clothed in divergent spines; stip- ules hyaline-membranous, 15-18 mm long, the free portion lanceolate, acu- minate, ciliate-margined; leaves 3-6 (7) cm long, the petiole short, both petiole and rachis densely pubescent with subappressed hairs; leaflets 5-6 pairs, elliptic-lanceolate, prickle-tipped, 12-17 mm long, densely seri- ceous onboth sides with short silvery or grayishhairs; peduncles shorter than to equaling the leaves, 1-5 (6) cm long, densely pannose withshort soft hairs; inflorescence ovoid, 3-5 cm long, many-flowered; bracts indurated, thinly coriaceous, yellowish, hyaline-margined, narrower than the calyx, exceeded by the corolla and the calyx, glabrous on the outside, the lower ovate, the upper oblong, all subulate-acuminate, 10-12 (15) mm long, cili- ate-margined; calyx 11-16 mm long, markedly shorter than the corolla, softly shaggy, short-cylindric, ovoid-bladdery in fruit, the tube 1.5-2 times as long as the subulate teeth; corolla red; standard 16-25 mm long, the limb broadly pandurate-oblong, round-tipped andretuse, obtusely auriculate- : angular at base, 3-4 times as long as the claw; wing-petals barely exceeded by the standard, the limb lance-oblong, obtuse, the claw 1.5 times as long as the limb; keel 14-18 mm long, the limb gibbous beneath, scarcely convex above, the claw 1.5-2 times as long as the limb; ovary sessile, shaggy; style hairy at base, naked upward. Fl. (June) July-August. (Plate XXIX, Figure 2). Dry crevices; stony and rocky slopes at altitudes of 1500-2500 m.- Caucasus: Tal. Gen. distr.: N. Iran. Described from Mt. Zuvant (Tal.). Type in Leningrad. 401. A. lagurus Willd. in Act. Berol. (1794) 28; DC. Astr. (1802) 168; Bge. Astr. geront. II, 115; Boiss. Fl. or. II, 384; Grossh. Fl. Kavk. I, 299.-A. brachypodus Boiss. Diagn. sér.I, 2 (1843) 69.—Ie: = Willd: locwertabil, ff. 2: DC 1. es, taba 36s5E xs, 5 El. OF. exc NO mc oon Perennial, suffrutescent-cespitose, (12) 15-29 cm tall; branches strong- ly abbreviated, igneous, clothed in divergent rigid spines; stipules 10- 15 mm long, the free portion lance -oblong to lanceolate, ciliate -margined; leaves (4) 6-11 cm long, the petiole (1) 1.5-3 cm long, both petiole and 837, 1117 324 435 rachis densely and softly pubescent; leaflets 5-7 pairs, elliptic-oblong to lanceolate, 10-15 (20) mm long, sericeous on both sides with appressed silvery hairs, prickle-tipped; peduncles (5) 8-16 cm long, pannose with short white hairs, equaling to twice as long as (rarely shorter than) the leaves; inflorescence many-flowered, ovoid-oblong, (4) 5-9 cm long; bracts thinly coriaceous, indurated, yellowish, broad, sharply long-acuminate, exceeding the calyx, equaling to somewhat exceeding the corolla, sericeous on the outside over the whole surface or merely on the midrib, 15-20 mm long, the lower ovate, the upper oblong; calyx equaling the corolla or near- ly so, softly shaggy, at first subcylindric, 16-25 mm long, becoming sub- spherical-inflated, the subulate teeth as long as the tube; corolla whitish- carneous, the tip of keel often azure; standard 17-24 mm long, the limb broadly pandurate-oblong, round-tipped, subobtusely angular-auriculate at base, 2-3 times as long as the claw; wing-petals equaling the standard, the claw 1.5-2 times as long as the lanceolate obtuse limb; ovary sessile, white - sericeous; style hairy at base, naked upward. Fl. July-August. Dry stony mountain slopes, at altitudes of 1400-2400 m.- Caucasus; S. Transc. Gen. distr.: Asia Minor, Turkish Armenia, N. Iran. Described from Asia Minor. Type in Berlin. Subgenus VII. CERCIDOTHRIX Bge. Astrag. geront. I (1868) 94. - Perennials, subshrubs, or shrubs, vested with bifurcate hairs; calyx cyl- indric or campanulate, not inflated in fruit, ruptured by the pod, rarely scarcely inflated; pods coriaceous, rarely bladdery-inflated, membranous. Section 65. EUODMUS Bge. Astrag. geront. I (1869) 96; II, 168.-Her- baceous perennials; stems commonly long, erect; stipules more or less connate, rarely the upper ones free; leaves imparipinnate, the petiole mar- cescent; racemes loose or fairly so, mostly many-flowered, borne ona long peduncle; calyx cylindric or cylindric-campanulate, not becoming in- flated; corolla glabrous; wings shorter than the standard, exceeding the keel; pods oblong or lance-oblong, sessile, erect or cernuous, carinate or rounded ventrally, grooved dorsally, coriaceous, fully or partially bilocular. IER OCISRORO CT 1 sperm him sy cvenciiog, ats ooh btsiesersy «i= Tourspers Gomer eberencsseuenis Sys wes 2 Se OC SECE RNUOUSH Hl) ped Mornehts lols Gepleittdichte, db sf tylicmiel” Sofa)» ner eresysedthe, ahieiue sploue 4. Zee Ovary and ppodyGlabrouisil)...wes%, .. sepals ++ 230. eh 402. A. uliginosus L. am ONeiay, AnGypoduhar yarn. A .an5, specials sxe heesubind. enerey te sfoeeiaqwiney os's » Spence» 3. 3. Calyx and pod covered with black hairs. 403. A. Schelichovii Turcz. TS calysand pod covered with whitemnairs...s 6 alse se tere Me tte. ite te, ane le = SOMONE ERGOT b. Guor ae Cea oere aie 404, A. kamtschaticus (Kom.) Gontsch. 4. Upper stipules free; flowers drooping, 10-12 mm long; pods linear - oblong, (14) 17-22 mmlong, falcate........ 408. A. falcatus Lam. + All stipules more or less connate; pods oblong or lance-oblong, straight. OV SURE HG NGMS ROWAN Rolion Gn Moti omOn sia OU Lom McOmOBONO yole Bo, 0. btn G2 on +0 5. 5. Pods 8-9 mm long, lance-oblong; flowers drooping; standard 9-11 (12) mm long, the limb obcordate to oblong-obcordate .......... 5 BN aN Gee BUSN SES la Ayetol Suc OuoND ONO LON AMokotloyion 407. A. odoratus Lam. + Pods 11-15 (20) mm long, oblong; standard 15-16 mm long, or 11- 13 mm long and then flowers horizontally spreading..........-. 6. 325 436 437 6. Flowers drooping, 15-16 mm long..... 405. A. peduncularis Royle. + Wlowers*horizontally spreading) /11=13, mim lOnp a. eh). vee Mek ee eens see a ees WEN rai a Sh Tam eee eet aah ee 406. A. discessiflorus Gontsch. Series 1. ERECTOSTRICTI Gontsch. - Pods erect, straight; racemes rather compact; all stipules more or less connate. 402. A. uliginosus L. Sp. pl. ed. I (1753) 757; DC. Astrag. (1802) 138; Ldb. Fl. Ross. I, 604; Bge. Astrag. geront.II, 168; Krylov, Fl. Zap. Sib. VII, 1670.-A. nertschinskensis FreyninO.B.Z. (1902) 21.-Ic.: DC. tab. 26. Perennial, (25) 46-92 cm tall; stems erect, usually several, (20) 40- 85 cm long, rather sparsely covered with short white or at the nodes mostly mixed black and white hairs; stipules connate at base, 6-10 mm long, mem- branous, appressed-hairy, the lower triangular, the upper lanceolate, long- acuminate; leaves (7) 10-18 cm long, the petiole and the much longer rachis slender, thinly covered with short appressed or subappressed white or mixed black and white hairs; leaflets 10-13 pairs, oblong-elliptic, obtuse or mi- nutely mucronulate, (1) 2-4 cm long, 0.5-1.5 cm broad, glabrous above, cov- ered beneath with short appressed hairs; peduncles 1/2-2/3 as long as the leaves, about as long as the inflorescence, finely covered with appressed black and white hairs; racemes dense, oblong or cylindric, 3-6 cm long, many-flowered, the flowers subsessile, the lower drooping; bracts ovate - lanceolate, finely pointed, somewhat exceeding the calyx, 5-6 mm long, hyaline, sparsely black-ciliate; calyx ca. 9 mm long, covered with short appressed black hairs, the tube about twice as long as the linear teeth; co- rolla pale greenish-yellow, sometimes fading slightly reddish; standard 14-18 mm long, the limb oval, scarcely angular at base, slightly retuse, 1.5 times as long as the claw; wing-petals 11-14 mm long, the limb oblong, obtuse or scarcely retuse, about as long as the claw; keel 9-12 mm long, the limb rounded-gibbous, acutish, as long as the claw, violet-tipped; ovary glabrous, sessile; pods sessile, obliquely ascending, oblong-ovoid or ob- long, terminating in a divaricate beak 1.5-2 mm long, inflated, 9-13 mm long, 3.5-4 mm broad, 4-5 mm thick, rounded or slightly carinate ventral - ly, more or less grooved dorsally, coriaceous, glabrous, bilocular. Fl. June-July; fr. July-August. Inundated forest or steppe-like forest meadows, river banks, and mar- gins of dark coniferous forests; mostly in river vaileys. A typical meso- phyte.-W. Siberia: Ob (Tarskii district, Tobol'sk), Irt., Alt.; E. Siberia: Ang.-Say., Dau. Gen. distr.: Mong., Jap.-Chi. (N. Korea, Manchuria, N. China). Described from Siberia. Type in London. 403. A. Schelichovii Turcz. in Bull. Soc. Nat. Mosc. (1840) 68; Ldb. Fl. Ross. I, 605; Bge. Astrag. geront. II, No. 168. Perennial, (8) 22-40 (45) cm tall; stems (3) 15-35 (40) cm long, finely pubescent with appressed white hairs; stipules connate to the middle, 6- 8 mm long, the free portion triangular, acuminate, thinly scarious; leaves (6) 10-14 cm long, the petiole and the much longer rachis slender, pubes- cent with appressed white hairs; leaflets 9-12 pairs, oval to oblong-oval, round-tipped, rarely scarcely retuse, 10-20 mm long, thin, glabrous above, finely hairy beneath; peduncles 2/5-2/3 as long as the leaves, 4-10 cm long, appressed-pubescent with white and beneath the inflorescence mixed white and black hairs; racemes loosely ovoid, 3.5-4 cm long, at least the lower 326 438 flowers drooping; bracts lanceolate to ovate-lanceolate, acuminate, white - hyaline, finely black-ciliate; pedicels ca. 1 mm long; calyx cylindric-sub- campanulate, ca. 6 mm long, thinly black-hairy, the subulate teeth1-1.5mm long; corolla yellowish; standard 12-14 mm long, the limb oval, obtusish and scarcely retuse, 1.5-2 times as long as the claw; wing-petals 10-11 mm long, the limb linear-oblong, entire, as long as the claw; keel 9-10 mm long, the limb convex-gibbous, obtusish, as long as the claw; ovary sessile, covered with white and black hairs; pods sessile, oblong, somewhat oblique - ly ascending, aggregated in a compact raceme, 10-14 mm long, 3-4 mm broad, membranous -coriaceous, sparsely andthinly covered withappressed black and considerably fewer white hairs, carinate ventrally, grooved dor- sally, terminating in a slender divaricate beak 2-4 mm long, bilocular. Fl. June-July; fr. July-August (September). Sandy and pebbly banks in river valleys, wet meadows, osier-beds, rarely in larch taiga.- Arctic: Anad.; E. Siberia: Ang.-Say. (N.), Len.-Kol. (mainly in E. part); Far East: Kamch., Okh., Ze.-Bu., Uda, Sakh. En- demic. Described from the area between Yakutsk and Okhotsk. Type in Leningrad. 404. A. kamtschaticus (Kom.) Gontsch. comb. nova.-A. uliginosus L. var. kamtschatica Kam. Fl. Komtch. II (1929) 281. Perennial, 50-60 cm tall; stems erect, firm, 40-47 cm long, thinly cov- ered with appressed white hairs; stipules connate at base, 6-10 mm long, covered with white appressed hairs, the lower oblong, the upper lanceolate, acuminate; leaves 12-18 cm long, the petiole and the much longer rachis finely covered with appressed white hairs; leaflets (6) 8-12 pairs, ovate- oblong, subobtuse, 18-25 (34) mm long, (4) 6-13 mm broad, glabrous above, covered beneath with short appressed white hairs; peduncles shorter than to about as long as the leaves, firm, sulcate, thinly covered with appressed white hairs; racemes elongate, rather many-flowered, compact, 9-14 cm long in fruit, the flowers drooping; bracts linear, thin, whitish, 5-7 mm long, finely white -ciliate; calyx campanulate-cylindric, ca. 4mm long, densely covered with fine appressed white hairs, the subulate teeth ca.1mm long; corolla yellowish (?); standard 11-12 mm long, the limb obovate, gradually attenuate into and 1.5 times as long as the claw, scarcely retuse; wing-petals 10-11 mm long, the limb oblong, obtuse, as long as the claw; keel 8-9 mm long, the oblong limb as long as the claw; ovary sessile, dense- ly white-pubescent; pods sessile, oblong, 15-19 mm long, ca. 3 mm broad, slightly carinate ventrally, grooved dorsally, terminating in a divaricate beak 4-5 mm long, coriaceous, densely and finely pubescent with appressed white hairs, bilocular. Fl. June; fr. July. Osier-beds and riverside pebbles. - Far East: Kamch. Endemic. De- scribed from Kamchatka river valley. Type in Leningrad. Series 2. NUTANTES Gontsch. - Pods cernuous, more or less appressed to inflorescence rachis; all stipules more or less connate. 405. A. peduncularis Royle, Illustr. Bot, Himal. (1839) 199; Bgo. As- trag. geront. II, 168; Hook. Fl. Brit. Ind. Il, 136.-A. corydalinus Bge. in A.H.P. III (1874) 101; Bge. Astrag. turk. 261.- Exs.:H.F.A.M.No. 393.- Ic.: Fl. Tadzhik. V. tab. 46. Perennial, 25-70 cm tall; stems many, 20-60 cm long, angled-sulcate, erect or more often ascending in lower part, canescent withappressed hairs; 327 441 stipules not adnate to petiole, connate, at length mostly splitting, triangular - lanceolate, 6-8 mm long, acute; leaves 5-10 cm long, the very short petiole and the rachis covered with appressed white hairs; leaflets 7-12 pairs, el- liptic-oblong, rarely broadly sub-linear, (7) 11-19 (25) mm long, round- tipped and mostly terminating in a minute point, covered with scattered appressed hairs or glabrous above, canescent beneath with appressed hairs; peduncles axillary and terminal, 10-18 cm long, exceeding the leaves; ra- cemes rather loose, oblong, many-flowered, the flowers drooping; bracts lanceolate to linear-lanceolate, 2-3 mm long, exceeding the pedicel, white - hyaline, finely black-ciliate on the margin; calyx cylindric, 8-10 mm long, covered with long white or short black or mixed white and black: hairs, the triangular-subulate teeth 1-1.5 mm long; corolla greenish-white, the stand - ard sometimes more or less purple-tinged; standard 15-16 mm long, the limb rounded-obovate, entire or slightly retuse, 2-2.5 times as long as the claw; wing-petals 11-12 mm long, the limb linear-oblong, entire or slightly retuse, about as long as the claw; keel 10-11 mm long, the obtuse limb as long as the claw; pods cernuous, more or less appressed to inflorescence rachis, oblong, 11-15 (20) mm long, sessile, abruptly contracted into a beak 1-2 mm long, slightly carinate or nearly flat ventrally, more or less grooved dorsally, coriaceous, finely pubescent with appressed white or rarely black or mixed white and black hairs, incompletely bilocular, 6-10- seeded. Fl. (April) May-July; fr. (June) August-September. (Plate XXX, Figure 2). Mountain slopes, shrub thickets and open woods, in the deciduous wood - land and scrub belt, in fescue steppe and juniper belts. - Soviet Centr. Asia: Tien. Shan (W.), Pam.-Al. Gen. distr.: W. Tibet, N.W. India. Described from N.W. India. Type in London. 406. A. discessiflorus Gontsch. in Fl. Tadzhik. V (1937) 676. Perennial, canescent plants, 47-51 cm tall; stems subflexuous, 38-45cm long, 4-angled, branched, densely vested with appressed white hairs; stip- ules connate to the middle, 7 mm long, covered with scattered appressed white hairs, the free portion triangular-lanceolate; leaves 7-9 cm long (5.5- 7 cm on lateral branches), the very short petiole and the rachis covered with appressed white hairs; leaflets (6) 7-9 (10) pairs, oblong-oval, those of upper leaves oval-elliptic, 8-13 mm long, obtuse, glabrous or nearly so above, densely covered beneath with appressed white hairs; peduncles exceeding the leaves, 10-13 cm long, covered with white and in upper part with mixed white and black hairs; racemes oblong, (3) 6-8 cm long, (10) 15-30 (35)-flowered, loose, the flowers horizontally spreading; bracts linear-lanceolate, white-hyaline, 2.5-3 mm long; pedicels ca. 1 mm long; calyx campanulate-cylindric, ca. 5 mm long, densely covered with short white pubescence interspersed with longer black hairs, the triangular teeth ca. 1 mm long, profusely black-hairy; corolla greenish-white; standard 11-13 mm long, the limb rounded-obovate, entire or slightly retuse, the claw 4-5 mm long; wing-petals ca. 11 mm long, the limb oblong, obtuse, half as long as the claw; keel 40 mm long; pods sessile, oblong, cernuous and more or less appressed to inflorescence axis, 11-12 mm long, abrupt- ly contracted into a slender beak ca. 5 mm long, slightly carinate ventrally, deeply grooved dorsally, coriaceous, densely pubescent with appressed white hairs, bilocular, 4-6-seeded. Fl. andfr. June. Margins of rosaria and rosarium-derived mixed-grass and orchard- grass meadows in the rosarium belt. - Soviet Centr. Asia: Pam.-Al. 328 =— a lj f ppee le lf PLATE XXX 1. Astragalus asper Jacq.—2. A. peduncularis Royle. 329 442 443 Endemic. Known only from a single site on the western slope of Sanglak Mts. in S. Tadzhikistan. Type in Leningrad. 407. A. odoratus Lam. Encycl. I (1789) 311; DC. Astrag. (1802) 139; Ldb. Fl. Ross. I, 605; Bge. Astrag. geront. II, 168; Bge. Astrag. turk. (1880) 261.-Ic.: DC. Astrag. tab. 26. Perennial, 20-35 cm tall; stems 15-30 cm long, erect or in lower part ascending, slender, covered with fine appressed white hairs; stipules con- nate to the middle, thinly membranous, 7-9 mm long, the lower triangular, the upper lanceolate, acuminate, 7-9 mm long, sparsely white -ciliolate; leaves 8-12 cm long, the petiole obsolescent, the rachis slender, finely covered with appressed white hairs; leaflets 9-12 pairs, lanceolate or el- liptic-oblong, (10) 12-17 mm long, subacute to subobtuse, minutely mucron- ulate, 3-4 mm broad, glabrous above, very finely covered beneath with appressed white hairs; peduncles 2/3 as long as to slightly shorter than the leaves, together with inflorescence equaling to somewhat exceeding the leaves, finely covered with appressed white and beneath the inflorescence mixed white and black hairs; racemes oblong, 4-5 cm long, rather loosely many-flowered; bracts linear-lanceolate, ca. 2 mm long, black-ciliolate; pedicels 1 mm long; calyx campanulate -cylindric, 4-4.5 mm long, minutely puberulent with black appressed hairs, the subulate -triangular teeth ca. 1 mm long; corolla whitish; standard 9-11 (12) mm long, the limb obcordate to oblong-obcordate; 2-3 times as long as the claw; wing-petals 8-9 mm long, the limb oblong, obtuse, 1.5-2 times as long as the claw; keel 7-8 mm long, the limb semicircular, acutish, as long as the claw; ovary sessile, glabrous; pods cernuous in a compact raceme, sessile, lance-oblong, acu- tish, 8-9 mm long, ca. 3 mm broad, terminating in a slender beak 1.5- 2 mm long, inflated, rounded and prominently sutured ventrally, sulcate dorsally, coriaceous, minutely covered with scattered appressed hairs, bilocular. Fl. June-July; fr. July. Scrub steppes in the foothills and mountain meadows. —- Caucasus: Dag., S. Transc., Tal; Soviet Centr. Asia: Dzu.-Tarb. (Tarbagatai). Gen. distr. : Bal.-As. Min. (Anatolia), Arm.-Kurd. (Turkish Armenia), Iran (N. Iran). Described from the Levant. Type in Paris. Series 3. FALCATI Gontsch.- Pods cernuous, linear-oblong, falcate; stipules of upper leaves free. 408. A. falcatus Lam. Encycl. I (1789) 310; Pall. Astrag. (1800) 34; DC. Astrag. 141; Ldb. Fl. Ross. I, 619; Bge. Astrag. geront, II, 168; Schmalh. Fl. I, 277. Perennial, 55-85 cm tall; stems erect, 45-70 cm long, finely sulcate, sparsely and finely covered with appressed black and white hairs; stipules dimorphic, the lower connate at base, ovate -triangular, the upper free, lance -oblong to lanceolate, acuminate, 5-11 mm long, white -hyaline, sparse - ly covered with white and black hairs; leaves 10-16 cm long, the petiole very short, the rachis finely and sparsely pubescent with subappressed white hairs; leaflets 9-18 pairs, oblong, rarely oval-oblong, round -tipped and minutely mucronulate, 10-20 mm long, glabrous above, sparsely covered beneath with fine appressed hairs; peduncles equaling the leaves, firm, terete, finely sulcate, very finely covered with scattered appressed white and black hairs; racemes oblong, (8) 10-12 cm long, rarely ovoid, loosely many-flowered, the flowers more or less drooping; bracts linear -lanceolate, 330 444 4-6 mm long, finely covered with black and white hairs; pedicels ca. 1 mm long, covered with white and predominantly black hairs; calyx campanulate, 5 mm long, rather densely and finely covered with appressed black hairs, the triangular or linear -triangular teeth 1-1.2 mm long; corolla whitish, faintly suffused with purple; standard 10-12 mm long, obovate, retuse, 1.5- 2 times as long as the claw; wing-petals 9-11 mm long, oblong, obtuse, the limb somewhat longer than the claw; keel 8-5-10 [?] mm long, the limb semicircular, obtusish, as long as the claw; pods sessile, pointed at base, linear-oblong, (14) 17-22 mm long, 3.5-4 mm broad, slightly compressed laterally, falcate, carinate ventrally, deeply grooved dorsally, coriaceous, very finely and sparsely covered with appressed white and fewer black or rarely pure black hairs, bilocular, pointed to a straight subulate beak 1.5- 2mm long. Fl. June-July; fr. July-August. Open deciduous (oak) woods and their clearings, meadow scrub, and rarely needle-grass and mixed-grass steppes in the forest-steppe region. In the Caucasus in Persian walnut groves, mountain birch and pine woods, and river banks, ascending in the South up to 1400 m.- European part: V.- ka. (S.), V.-Don (N.), L. Don; Caucasus: Cisc., E. and S. Transc., Dag. Gen. distr.: introduced as a forage plant into Central Europe where it grew wild. Described from the European part of the USSR. Type in Paris. Section 66. PEDINA (Stev.) Bge. Astag. geront. I (1868) 96; II, 169. Gen. Pedina Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 144. - Herba- ceous perennials; stems long, erect, vested with bifurcate hairs; stipules free (not connate), adnate at base to petiole; leaves imparipinnate, the peti- ole marcescent; flowers erect; racemes rather compact, borne on a long axillary peduncle; bracteoles none; flowers yellow; calyx cylindric-cam- panulate; corolla glabrous, marcescent; pods sessile, linear-oblong, ex- ceeding the calyx, carinate ventrally, sulcate dorsally, coriaceous, biloc- ular.- A monotypic section. 409. A. asper Jacq. Ic. Plant. Rar. I (1781-86) 15; Bge. Astrag. geront. II, 169; Ldb. Fl. Ross.I, 619; Schmalh. Fl. I, 277.-A. chloran- thus Pall. Astrag. (1800) 30, tab. 25.-Pedina aspera Stev. inBull. Soc. Nat. Mosc. XXIX (1856) 144.-Ic.: Jacq. l.c., tab. 152. Perennial, (25) 40-60 cmtall, from a short woodyburied caudex; stems (12) 20-45 cm long, short, sulcate, sparsely covered with fine appressed white hairs; stipules triangular-lanceolate, rarely triangular-ovate, 3-4mm long, acute; leaves 6-9 cm long, the very short petiole and the rachis sul- cate, sparsely and finely covered with stiff appressed hairs; leaflets (7) 8-10 pairs, linear-lanceolate to lance-linear, short-acuminate, (12) 15- 25 mm long, the appressed hairs scattered above, sparse beneath; pedun- cles equaling to twice as long as the leaves, sulcate, sparsely covered with appressed white and above the inflorescence sometimes mixed white and black hairs; racemes rather compact, spiciform, erect, the flowers sub- sessile; bracts ovate-lanceolate, acuminate, 2-3 mm long, sparsely white - and black- or merely black-ciliate; calyx cylindric-campanulate, (8) 10- 12 mm long, sparsely and finely covered with appressed white and black or pure black hairs, the teeth narrowly linear, the tube 1.5-2.5 times as long asthe teeth; corolla yellow; standard 18-21 mm long, the limb oval-oblong, retuse, angular atbase, 3-4timesaslongasthe claw; wing-petals 16-19mm long, the limb oblong, retuse, as long as the claw; keel 14-16 mm long, the 331 445 obtuse limb 2/3 as long as the claw; pods sessile, erect, mostly appressed to the inflorescence rachis and mutually appressed in a mostly compact spike, linear-oblong, 12-20 (22) mm long, 2.5-3 mm broad, straight or scarcely curved, pointed to a divaricate beak 2-3 mm long, carinate ven- trally, sulcate dorsally, carinate, coriaceous, finely covered with ap- pressed white or rarely mixed white and black hairs, bilocular. Fl. May- June; fr. July (August). (Plate XXX, Figure 1). Steppes, more rarely limestone outcrops. - EHuropean part: M. Dnp., V. Don, Transv., Bes., Bl., Crim. (Kerch', Perekop), L. Don; Caucasus: Cisc. Gen. distr.: Centr. Europe (Austria, Rumania), Bal.- As. Min. (Dobruja). Described from Austria. Type in Vienna. Section 67. CRACCINA (Stev.) Bge. Astrag. geront. I (1868) 97; II, 169.-Gen. Craccina Stev. in Bull. Soc. Nat. Mosc. IV (1832) 266; l.c. XXIX (1856) 266. - Herbaceous perennials, with well developed rarely di- minutive stems, less often suffrutescent plants. Pubescence consisting of bifurcate white or mixed white and black hairs; stipules free or connate; leaves imparipinnate, the petiole marcescent; flowers mostly small; ra- cemes loose, elongate or more or less contracted, short, fairly loose, borne on axillary peduncles; bracteoles none; calyx campanulate or cylin- dric-campanulate; corolla violet or purple, rarely white; keel exceeded by the wings; pods sessile or short-stipitate, linear-oblong or linear-lan- ceolate, rarely oblong, grooved dorsally, thinly coriaceous, bilocular or nearly so. Xerophytes, rarely meadow mesophytes, associated with steppe vege- tation zone, and within it with limestone and sand. The distribution area extends from the Atlantic Ocean to Mongolia. With the exception of one spe- cies (A. bayonnensis Lois. from the maritime sands of S. W. France), all species of this section occur within the territory of the USSR. 1. Standard broadly or narrowly ligulate in upper part........... 2. 4-4 Standard NOtMOULALe . Akwcucw lm ani tole 7-¥ ceuch cd ©) fim louie, “po uceelcy icine 4. Zep lecarletSHODOVate. oes CNN tl -) el ace) Peet 3. + Leaflets narrowly linear.......... 413. A. Petkoffii B. Fedtsch. 3. Calyx covered with white hairs; pods pubescent with appressed yellow- USHGHaALES Me. aac) eeecmspeus 2 iam broads.) Seni aah s oueney oh 5 Re ss LEVIS, ya CRIES Rant aed OM OM CS Re me es ees es Rete ee 442. A. virgeus Boriss. + Stems 2-5, not flagelliform, 3-4 mm in diam., greenish or sparsely covered with white appressed hairs, 20-50 cm long; calyx 7-8 mm long, the teeth 2-3 mm long; corolla 17-22 mm long; pods falcate, 2- TIE LOMS SHC 14) (STI ROA | santas jhe hie, terheiiie) seieeol| sles’. sekecnls euwpye: jo uel cells 9. 9. Inflorescence a loosely few-flowered raceme, the flowers distant; calyx-tube 3 times the length of the teeth; standard oblong-lanceolate, CAA Talia Koy aVeTy Eaten Sa ae ra ven) enee nM Mee eae an eer 443. A. Stevenianus DC. + Inflorescence an ovoid or oblong dense many-flowered raceme; calyx- tube 1.5 times as long as the teeth; standard oblong-rhomboid, 17- 2al0): Tae Moyea 5) An avn ae UTS 444. A. applicatus Boriss. 10. Calyx ca. 5 mm long, shortly cylindric-campanulate, the tube 3-4 rmesiac ong ac the teeth, .corolla 3 —lGr mimi. oi sueqe ee) ye ole eueiene. 6 MT Ms! Sota cae? bus Neh ayia ate a wal gies ia) 445. A. ornithopodioides Lam. + Calyx 8-10 mm long, cylindric, the tube equaling to 1.5-2 times as lonceasitnewtectis (corolla G20 mai Ome. 5) cee nel 6.5) io) ol oly wien a) felsa) lal 6 SMCS RNC is Paleo iichi/al low oilec\ (1) /of le) shy ay | si)i61 1 446. A. achtalensis Conrath et Freyn. Series 1. MONILIFORMES Boriss.- Wings weak, thin; upper stipules free, podsflattened, cernuous, whiie-hairy, the teeth strongly outlined, round. 436. A. Shelkovnikovii Grossh. in Beih. Bot. Centralbl. XLIV, II (1928) AZO lon may k: Mis 23)0— Excel orivexs. (No! 2Z8i7. Perennial, weak plants, vested all over with appressed cinereous hairs; stems branching 10-30 cm long, slender, angularly flexuous; stipules herbaceous, triangular-lanceolate, 3-5 mm long, cauline, the upper ones free, sparsely hairy; leaves 3-7 cm long, the petiole 1-2 cm long; leaf- lets 4-7 pairs, narrowly linear, the upper linear, 6-20 mm long, 1- 1.5 mm broad, acute, sparsely hairy above, more densely covered beneath with appressed hairs; peduncles equaling or exceeding the leaves; inflores- cence a loosely 5-14-flowered raceme, 3-5 cm long; pedicels ca. 1 mm long; bracts 1-1.5 mm long, linear, white-hairy; calyx cylindric, pubescent with black and white hairs, 6-7 mm long, dilated in fruit, the tube 3-4 times the length of the subulate teeth; corolla purple, 14-16 mm long; standard oblong-rhomboid, enlarged at the middle, up to 6 mm long, obtuse, scarce- ly retuse, attenuate toward base and toward apex; wings ca. 11 mm long, 1-1.5 mm broad, the oblong limb about as long as the claw; keel 10 mm long, the limb inequilaterally oval, 5mm long, 2 mm broad, as long as the claw; ovary linear, glabrous; style ca. 6 mm long; pods linear-cylindric, flattened, cernuous, somewhat curved, 2.5-3 cm long, grooved on the back, Sparsely covered with appressed white hairs, the round outline of the seeds very distinct. Fl. andfr. May-June. (Plate XXXI, Figure 3). Dry stony slopes, at altitudes of 570-1400 m, sometimes as a weed of field crops.- Caucasus: S. Transc. (Erevan, Nakhichevan'). Gen. distr : Iran. Described from the vicinity of Dzhulfa. Type in Baku. Series 2. NIGRESCENTES Boriss.- Roots strong, ligneous; stipules connate at base; pods cernuous, predominantly black-hairy, cylindric. 351 472 437. A. glochideus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 45-50 cm tall, from a strong woody taproot; stems many, slender, ca. 2.5-3 mm in diam., sulcate, canescent by appressed white hairs (interspersed with black around the nodes and on the infl. rachis, angularly flexuous, few-branched, the internodes usually shorter than the leaves; stipules connate and adnate to petiole at base, membranous, shortly broad-triangular, 3-4 mm long, acute, glabrous or sparsely white -hairy; leaves 7-10 cm long, short-petiolate; leaflets 8-10 (11) pairs, oblong or lanceolate, subobtuse, 8-13 mm long, 2-4 mm broad, green, glabrous above, sparsely hairy beneath; peduncles exceeding the leaves, 12- 15 cm long, sparsely covered in lower part with short black hairs; ra- chis of inflorescence black-hairy; racemes loose, the distant flowers borne on black-hairy pedicels ca. 2 mm long; bracts ca. 1.5 mm long, linear; calyx ca. 6 mm long, sparsely covered with short black hairs, asymmetri- cal, the very short triangular teeth ca. 0.5 mm long; corolla drying dark violet; standard 22-23 mm long, oblong-rhomboid, cuneate-based, en- larged below the middle to 7 mm, gradually attenuate toward apex, acutish; wing~petals ca. 14 mm long, the lanceolate limb 2 mm broad, about twice as long as the claw; keel 10 mm long, the limb oblong, 2 mm broad, as long as the claw; ovary linear, sparsely hairy; style naked, ca. 6 mm long; pods cernuous, linear, hamate, 2-3 cm long, ca. 1.5 mm broad, acuminate, hairy, the black hairs interspersed with white. Fl. and fr. June-July. Grass~-covered and wooded slopes. —Caucasus: Tal. (N. Gelyadar, Bobola Baid mountain, toward south of Tongavan village). Endemic. Described from Talysh (northern Gelyadar). Type in Leningrad. Note. Resembling A. jodostachys Boiss. et Buhse, from which it differs inits larger size, shape of leaflets, the loose inflorescence, shape and length of calyx teeth, shape of standard and wings, and shape and position of pods. 438. A. jodostachys Boiss. et Buhse, in Nouv. Mém. Soc. Nat. Mosc. XII (1860) 60; Boiss. Fl. or. II, 427; Bge. Astrag. geront. II, 172. Perennial, many-stemmed, 25-30 cium tall; stems ascending, elongated, flexuous, cinereous by appressed bristly hairs; stipules membranous, acu- minate, triangular, connate at base, up to 5 mm long; leaflets 7-12 pairs, oblong-linear or linear-lanceolate to oblong, triangular or subacute; pe- duncles exceeding the leaves, thickish; racemes oblong, dense, at length becoming loose and elongating; bracts linear to lanceolate, setiform, ex- ceeding the very short pedicel; calyx campanulate-cylindric, ca. 8 mm long, sparsely covered with white and black hairs, the teeth subulate, ca. 2 mm long, the tube 2-3 times as long; corolla violet; standard 17-20 mm long, oblong-lanceolate, obtuse, sometimes mucronate; wing-petals 11-14 mm long, the oblong limb about as long as the claw; keel 9-11 mm long, the ovate limb somewhat shorter than the claw; ovary ovoid, glabrous; style 5-6 mm long; pods 2.5-3 cm long, cernuous or horizontally spreading, linear-cylindric, somewhat curved, covered over part of the surface with black appressed hairs, speckled, acuminate, grooved. Fl. andfr. June. Possibly growing in S. Transcaucasia. Gen. distr.: Iran (Sakhend mountains Eastward from Lake Urmiya), Arm.-Kurd. (former Kars territory). Described from Mount Sakhend. Type in Leningrad. 439. A. Conrathii Freyn in Bull. Herb. Boiss. III (1895) 182; Grossh. JaG sens J, BAB). 352 Perennial, many-stemmed, spreading plants, very sparsely vested with appressed hairs; stems ascending, branched, 20-25 cm long; stipules mem- branous, small, ovate-triangular, connate up to the middle, acute and scarcely divergent; leaflets 10-12 pairs, obtuse or obsoletely retuse, often mucronate, oblong or elliptic, up to 6.5 mm long, 2.5 mm broad; peduncles 8-12 cm long, straight, sulcate, scarcely or markedly exceeding the leaves; racemes few-flowered (upto12 flowers), cylindric, rather dense, at length elongating and becoming loose, 4-6 cm long; bracts divergent, exceeding the pedicel, membranous, triangular-ovate, ciliate, at length spreading; calyx cylindric, covered with appressed white and black hairs, 6-7 mm long, the tube 4 times as long as the triangular acute teeth; corolla red (?); standard ca. 20 mm long, 3.5 mm broad, rather shallowly emarginate, strongly re- curved, 3 times as long as the wings; wing-petals exceeding the keel; pods linear, ca. 16 mm long, 2.5 mm broad, densely covered with appressed black hairs, extending into a short conic beak, narrowly grooved dorsally, blunt ventrally. May. Caucasus: E. Transc. (Karagayadag). Described from the Akhtal border. Endemic. Type in Geneva. Series 3. ALBESCENTES Boriss.- Roots ligneous; stipules connate at base; pods erect or horizontally spreading, white-hairy or glabrous, cy- lindric. 440. A. trigonelloides Boiss. Diagn.I, 2 (1843) 49; Boiss. Fl. or. II, 425; Bge. Astrag. geront. II, 171. Perennial, ligneous from base, 2.5-5 cm tall, covered throughout with gray silky appressed hairs; stems erect, scantily branched; stipules mem- branous, small, connate; leaflets 4 pairs, oblong-elliptic, acute, 5-7.5mm long, 2-4 mm broad; peduncles terminal, shorter than the leaves; inflores- cence densely capitate, 6-8-flowered, not elongating in fruit; bracts lan- ceolate, hairy, barely exceeding the pedicel; calyx appressed-hairy, the subulate teeth as long as the tube; pods cernuous, curved, cylindric, ca. 2.5 cm long, 4mm broad, grooved dorsally, pointed, many-seeded, sub- tuberculate, 4-5 times as long as the calyx. Fl. June. Caucasus:S. Transc.(recorded for Armenia).Gen. distr.: Bal.-As. Min.(Asia Minor). Describedfrom Armenia (Aucher pl. exs. No. 1381). Type in Geneva. 441. A. ispirensis Boiss. Diagn. II, 2 (1856) 25; Boiss. Fl. or. I, 426; Bge. Astrag. geront. II, 171. Perennial, many-stemmed, ligneous at base, 14-30 cm tall; stems gla- brous, sulcate, erect, branched from base; branches ascending; stipules shortly connate into a single bidentate structure; leaves elongate-elliptic to oblong-linear, obtuse to subacute, cinereous by bristlelike appressed hairs; leaflets 5-7 pairs, 10-12 mm long; peduncles exceeding the leaves, glabrate; inflorescence terminal, oblong, rather loose, 8-12-flowered; bracts linear, setiform, not exceeding the very short pedicel; calyx covered with black appressed hairs, the tube twice as long as the lanceolate teeth; corolla 2.5 cm long, intensely violet, 4 times as long as the calyx; stand- ard linear, obtuse, greatly exceeding the wings and the keel; pods 3.4 cm long, glabrous, cylindric, curved through half a circle, constricted between the teeth, 5-6 times as long as the calyx. June. Possibly growing in S. Transcaucasia. Gen. distr.: Arm.-Kurd. (Turk- ish Armenia). Described from vicinity of Ispir (North of Turkish Armenia). Type in Geneva. 353 475 442. A. virgeus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 15-31 cm tall, from a strong ligneous root and many distant woody caudices, densely cespitose; stems many, erect, flagelliform, slen- der, flexuous, canescent by dense appressed hairs, usually exceeding the leaves, few-branched, sulcate, the internodes long; stipules small, mem- branous, short-triangular, 1-2 mm long, connate and adnate to petiole at base, sparsely covered with appressed white hairs; leaves 2-3 cm long, short-petioled; leaflets 3-5 pairs, distant, linear to oblong-linear, 4-12mm long, 0.5-1.5 (2.5) mm broad, subacute, appressed-hairybeneath, glabrous or sparsely hairy above; peduncles axillary and terminal, greatly exceeding the leaves; racemes loosely few-flowered, short-pediceied; bracts small, ca. 1mm long, membranous, triangular-lanceolate, white-hairy; calyx campanulate, round-based, 4-5mm long, whitish by appressed hairs, asym- metrical, the subulate teeth 0.5-1 mm long; corolla drying pale lilac; stand- ard oblong-rhomboid, 13-15 mm long, enlarged at the middle to 4 mm long, attenuate toward both ends, obtusish or scarcely retuse, wing-petals 8- 11 mm long, the limb oblong, somewhat longer than the claw; keel 7-9 mm long, the oval limb about as long as the claw; ovary linear, white -hairy; style naked, ca. 5mm long; pods erect, linear, straight or slightly curved, 1.5-2.5 cm long, 1.5-2 mm broad, pointed at the end, covered with white appressed hairs. Fl. andfr. May-July. Dry stony slopes, on rocks. - Possibly growing in S. Transcaucasia. Gen. distr.: Arm.-Kurd. (former Artvin territory). Described from the banks of river Chorokh. Type in Leningrad. Note. Resembling A. Stevenianus DC., from which it is distin- guished by the numerous white slender stems, the length of leaves, shape of calyx and petals, shape of pod, and other characters. 443. A. Stevenianus DC. Prodr. II (1825) 285; Boiss. Fl. or. II, 426; Bge. Astrag. geront. II, 171; Ldb. Fl. Ross. I, 609; Grossh. Fl. Kavk. I, 32047 A. ye Ta ciddis) stev..nex DO ywlic., expludbs ul..c. 4, non. Nuttall ceus Ldb. in Spreng. Syst. veg. III (1826) 297.-A. dahuricus C.Koch in Linnaea XV (1840) 719, non DC. Perennial, glabrous or appressed-hairy, from a long woody tap-root; stems 20-40 cm long, slender, few-leaved, erect, the internodes long; stipules small, triangular, shortly connate and adnate to petiole, sparsely covered with white and black hairs; leaves 4-10 cm long; leaflets dimorphic, the upper narrowly linear, the lower ranging to oblong-lanceolate, distant, 3-5 pairs, 5-11 mm long, 1-2.5 mmbroad, covered sparsely above and densely beneath with appressed hairs; peduncles exceeding the leaves, sparsely appressed-hairy; inflorescence remotely few-flowered, elongating in fruit, loose; flowers short-pediceled; bracts ovate-triangular or linear, very small, 1-1.5 mm long, membranous, hairy; calyx 7-8 mm long, cy- lindric-campanulate, appressed hairy, the tube 3 times as long as the sub- ulate teeth; corolla purple; standard 20-22 mm long, oblong-lanceolate, re- tuse; wing-petals 13-16 mm long, the oblong limb about equaling the claw; ovary linear, hairy; style naked, ca. 7 mm long; pods linear-cylindric, 2-4 cm long, slender falcate, erect, pointed, more or less covered with appressed hairs, grooved dorsally. Fl. April-June; fr. May-June. (Plate XXXI, Figure 2). Dry stony and clayey slopes.- Caucasus: E. Transcaucasia. Gen. distr. : Iran. Described from the Caucasus (Iberia). Type in Geneva. 354 476 444. A. applicatus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 20-50 cm tall, from a ligneous taproot and divided caudices; stems 2-5, erect or ascending, with rather long internodes, usually exceed ing the leaves, weakly branched, sulcate, covered with appressed white hairs; stipules connate and adnate to petiole at base, membranous, the free portion triangular-acuminate, 3-4 mm long; leaves 3-7 cm long, short- petioled; leaflets 3-6 pairs, lanceolate to linear-lanceolate, rarely oblong, 10-25 mm long, 2-4 mm broad, covered with appressed white hairs, more densely below, less so or glabrous above; peduncles twice as long as the leaves; inflorescence an ovoid or oblong, rather dense, many-flowered ra- ceme, 1.5-9 cm long; flowers borne on a pedicel ca. 1 mm long; bracts linear, white-hairy, ca. 2 mm long; calyx cylindric, ca. 8mm long, cov- ered with white appressed hairs interspersed with fewer black, the filiform teeth ca. 3 mm long; corolla dark violet; standard 17-20 mm long, broadly oblong-rhomboid, up to 7 mm broad at the middle, ca. 3 mm at apex, scarcely retuse; wing-petals ca. 13 mm long, the oblong limb 6-8 mm long; keel ca. 10 mm long, the oval limb ca. 5 mm long; ovary linear, appressed- hairy; style naked, 7 mm long; pods linear-cylindric, at firsterect at length horizontally spreading, sparsely covered with appressed white hairs, acu- minate. Fl. April-May; fr. May. Dry sandy, gravelly and stony slopes. - Caucasus: S. Transcaucasia (Nakhichevan', Vagarshapat). Gen. distr.: possibly growing in parts ad- joining Iran. Described from the Caucasus and from the vicinity of Vagar- shapat (Echmiadzin). Type in Leningrad. Note. Resembling A. Stevenianus DC., from which it is distin- guished by the dense many-flowered inflorescence, the shape of calyx and petals, color of corolla, and several other characters; A. Kochianus Sosn., described from S. Transcaucasia (Dokl. A.N. Arm. SSR, III, 2, 1945, 2) is apparently related to A. applicatus Boriss. 445. A. ornithopodioides Lam. Encycl. method. I (1789) 312; Boiss. Blom ll 4255) Boe. Astras. geront. 11, 171).—A. nigrostriaitus C: Koch in. Linnaea XV (1840) 719.-A. multijugus Grossh. Fl. Kavk. II (1930) 325.-A. Stevenianus var. multijugus Trautv. in herb. Perennial, 8-22 cm tall, from a strong ligneous taproot, branched and woody at base, cespitose, canescent throughout by gray appressed pubes- cence; stems many, prostrate-ascending, slender, short, appressed-hairy; stipules triangular-lanceolate, 2-3 mm long, shortly connate, herbaceous or membranaceous; leaflets 7-14 pairs; peduncles usually exceeding the leaves, rarely as long; inflorescence 10-15-flowered, dense, somewhat elongating in fruit, 1-3 cm long; bracts lanceolate, equaling the pedicel; calyx ca. 5 mm long, white hairy, shortly cylindric-campanulate, covered with appressed black and white hairs, the tube 3-4 times as long as the subulate teeth; corolla violet; standard 13-16 mm long, obtuse or scarcely retuse, oblong-rhomboid; wing-petals 10-12 mm long, the oblong limb about as long as the claw; keel 8-10 mm long, the oval limb as long as the claw; ovary linear, appressed-hairy; style naked; Pods 3-4 cm long, 2.5 mm broad, spreading, covered with appressed white hairs, slender, angularly curved through half a circle, pointed, somewhat constricted between the seeds, many times as long as the calyx, obsoletely grooved dorsally. Fl. April-July; fr. May-July. Dry gravelly and stony slopes, at altitudes of 900-1700 m. - Caucasus: E. andS. Transc., Tal. Gen. distr. : Arm.—-Kurd. (Turkish Armenia, former Ol'tinskii district), As. Min., Iran. Describedfrom Armenia. Type in Paris. 355 " Lh c i) aul NO Pz Hel WSFA es fs NOW My tes AZZ ais) / ‘ aot Chen newer al = PLATE XXXII 1, Astragalus eremospartoides Rgl.—2. A. nematodes Bge.—3. A. subscoparius Gontsch. 837, 1117 479 » 480 446. A. achtalensis Conrath et Freyn in Bull. Herb. Boiss. III (1895) 182; Grossh. Fl. Kavk. II, 325. Perennial, from a rather slender taproot, 15-35 cm tall, branched at base from distant caudices; stems ascending, 5-6, mostly arcuately curved, with rather long internodes, often exceeding the leaves, usually fairly stout and strong, covered with firmly appressed hairs, these pure white below, interspersed upward with short black hairs; stipules membranous, the up- per green, connate and adnate to petiole, 3-5 mm long, the free portion triangular-lanceolate; leaves 4-8 cm long, short-petioled; leaflets (7) 8-10 (14) pairs, ovate to oblong, 6-11 mm long, 2.5-3 mm broad, covered with appressed white hairs; peduncles axillary or terminal, equaling or slightly exceeding the leaves; racemes capitate or ovoid, densely many-flowered, loose in lower part; pedicels 1-1.5 mm long; inflorescence rachis predom- inantly white-hairy; bracts 2-3 mm long, linear-lanceolate, covered with predominantly white hairs; calyx cylindric, 8-9 (10) mm long, the tube equaling to 1.5-2 times as long as the filiform teeth, covered with predom- inantly black appressed hairs; corolla drying violet; standard 18-20 mm long, oblong-rhomboid, 3.5 mm broad, cuneate at base, gradually attenuate upward, shallowly retuse; wing-petals 15 mm long, the limb oblong-spatu- late, about twice as long as the claw; keel 11 mm long, the ovallimb about as long as the claw; ovary linear-lanceolate, covered with scattered ap- pressed white hairs; style naked, ca. 6 mmlong; pods unknown. Fl. April- May. Stony and sandy slopes, rocks, among wormwood associations. - Cauca- sus: E. Transc., (former Karyaginskii district, S. Transc. (Erevan, Vagar- shapat). Gen. distr.: possibly growing in parts of adjoining Iran. Described from Mount Karagayadag, near Akhtal. Type in Geneva. Section 71. ONOBRYCHIUM Bge. Astrag. geront. I (1868) 95, 100; II (1869) 173.-Gen. Macrosema Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 146, pro parte. - Racemes capitate, borne on long axillary peduncles; bracts small; bracteoles none; calyx campanulate, rarely cylindric; corolla yellow or violet or purple, rarely white, sometimes drying yellowish; pe- tals usually glabrous, free, deciduous; ovary sessile; pods ovate, elliptic or oblong, bilocular, rarely partially bilocular or subunilocular; perennial herbaceous plants, with long or rarely short stems; hairs bifurcate, white or mainly on calyx bracts and fruits predominantly black; stipules cauline, more or less connate; leaves imparipinnate. 1. Standard oblong-spatulate, broadest in upper one-third, bilobate at apex, attenuate toward base, usually twice as long as the calyx (Siberia). + Standard oblong, broadest about the middle, drawn out or merely eons uate or not narrowed at apex, obtuse or retuse, gradually attenuate LOW AEUmMDaA SEs be Ain. o., Ue Male Vaca ras Is) loll ah cay ohfeinni We Mey na pobe” vepirika iba ls dou! pels Ve 5. 2. Calyx 8-9 mm long; corolla white drying yellowish, (15) 17-19 mm long; pods covered with white and on the back with black hairs; leaf- lets mainly large, oblong or ovate, obtuse, 10-30 mm long, 5-12 mm broad, glabrate; plants 30-60 cm tall (Maritime Territory)........ 482. A. marinus Boriss. eo) ame re pe ef Her se) 0, "0 ~eviepioe: o% ep fore, (6) 106. oie) le vel' eo; eo fe! ee) a 357 Calyx 5-6 or 7-8 mm long; corolla yellow, or violet or purple, rarely milky white, 12-16 mm long; leaflets lanceolate to oblong-elliptic or ovate, 4-25 mn» long, (2; “Nb GIS 480. A. austrosibiricus Schischk. Calyx 5-6 mm long, the tube 3-5 times as long as the setiform teeth; corolla purple; standard 5-6 mm broad; pods oblong-cylindric to tri- angular-cylindric, covered with short brown hairs.............. cee ity Derehaed® arheeietesmetcia He BA. eine. 2 479. A. adsurgens Pall. Green glabrate plants, sparsely covered with firmly appressed hairs; leaflets 3-6 pairs, ovate-oblong to orbicular, 7-18 (30) mm long, (3) 8-15 (20) mm broad; calyx cylindric; pods 2-2.5 mm broad, 13-15 mm long (Gentri) Asia, Altai). tead .wak 475. A. unilateralis Kar. et Kir. Plants more or less covered with appressed hairs, often silky-gray or green; leaflets oblong to linear, commonly 6-16 pairs, rarely 1-5 pairs; calyx campanulate, rarely cylindric; pods different from above. Pods lanate with dense white hairs, rounded-ovoid, equaling the ca- lyx; ovary densely covered with long hairs; standard 17-20-25 mm long, deeply and broadly bilobate ....... 468. A. lasioglottis Stev. Pods glabrous or more or less short-pubescent or hispid, ovoid to lanceolate; ovary glabrous, puberulent or covered with short white hairs; standard 10-29 mm long, entire at apex or scarcely retuse. 7. Pods pendulous or horizontally spreading becoming cernuous; racemes mostly loose) or merely elongated inifruit. 2... . 2 0. Belt ie ees 8. Pods erect, rarely horizontally spreading; racemes dense or occa- sionally somewhat elongating in fruit. .......-.....c-..4:+:s-.-. 10. Calyx 3.5-6 mm long; standard 10-15 mm or 16-20 mm long; pods lunate, more or less curved; gray or grayish-green plants .... 9. Calyx 7-9 mm long; standard ca. 15 mm long; pods straight; plants covered with raised white hairs ...... 470. A. adzharicus M. Pop. Calyx (6) 8-10 mm long; standard (18) 20-25 mm long; pod straight; ~ green plants sparsely covered with appressed hairs............ AR Aicugiy ce hSuloe fa Sralo coNe Memmi ie Pellerin 5.0918) 5.6. tls (i 471. A. arguricus Bge. Calyx 3.5-5 mm long; flowers 10-15 mm long; racemes dense, many- flowered, 1.5-3 cm long, loose and elongating in fruit; pods slightly curved, twice as long as the calyx; leaflets 9-14 pairs.......... SMM Er es cs aml tee eM sic ist hajmerey tna | ative 474. A. Kozlovskyi Grossh. Calyx 5-6 mm long; flowers 16-20 mm long; racemes 5-8 cm long, remotely flowered; pods lunate, 2.5-3 times as long as the calyx, predominantly black-hairy; leaflets 10-15 pairs................-. Wiret e Totemerte. felch ter vei cerlce, a" eaten al yeu fental telmegt es Neukell eile ie aura tre 472. A. lunatus Pall. Calyx 5 mm long; racemes remotely flowered; pods slightly curved, 3-4 times as long as the calyx, predominantly white-hairy; leaflets SHSipains!: cH wie eee Meee Ae tee 473. A. meskheticus Mand. 358 10. iat 12. 13. 14. 15. 16. ei (12). Calyx long-cylindric, appressed-hairy; peduncles 2-3 cm long; inflorescence dense, capitate; leaflets 6-8 pairs; plants 8-18 cm taliMrascencding (Pam vA Vie G RO) oe OOM Riel! La, Sle Calyx campanulate or short-cylindric; plants differing in other characters (Caucasus, western part of Soviet Centr. Asia, Europe). PRPHER: sett WPaGREy ASR USO CLE VNR RN TUTE RL, Me ieee LORE Ne CLT Eh Pods spreading, the beak 4-5 mm long, reflexed; leaflets 1-5 (6) pairs, elongate-lanceolate or linear; large plants, 35-60 cm tall (Kopet Dagh range)........ 469. A. brevidens Freyn et Sint. Pods erect, the beak mostly short, rarely up to 5-6 mm long; leaflets 6-16 pairs, rarely (2) 4-6 pairs and then the plants dif- FoI OMin ObhersChaAracrer sy th. Las tans hanes Ta ew eben vents ea 12. Calyx covered with spreading hairs, the teeth long, about equaling the tube, covered with hairs or sometimes with bristles. ...13. Calyx covered with appressed or rarely spreading hairs and then whole plant white -pannose; calyx-teeth usually much shorter than BHR TUDOR a yee SHrtte ne kmeare ee urn AINE LUEN DNR IERT EAS RAM A es We Plants 15-30 cm tall; stems 3-8 cm long, mostly ascending, rare - IVROTE CHR Uma arene. seu eMea is calc wets et ateuloM alts 2h cya) Te ccyvenio “Seis ce ialuia 14. Plants 25-40 cm tall; stems well developed, 20-35 cm long, MOsSuKy, CHRECE Manel y1a SCEMGIN GM cy. Bie ements suena iy ait ethos ae 15. Bracts ca. 6 mm long, oblong or lanceolate, leaflets 8-15 mm long, 3-4mmbroad, oblong pods ca. 8mm long, the beak 2-3 mm long, covered with spreading soft hairs. . 467. A. kadshorensis Bge. Bracts ca. 8 mm long, narrowly lanceolate or subulate; leaflets 1.5-3 mm broad, 5-15 mm long, oval-oblong to lanceolate; pods ean, KOhmmilons, Seovered withbristlyhairsis 0712 sR seit ade. 5. PUP ANEAT SRP cre. cath wtoiden ta: Fale oi 96. Najtel ho, He 466. A. onobrychioides M.B. Calyx covered with soft hairs; corolla12-18 mm long; pods 4-8 mm long, ovoid or oblong-ovoid; stems fairly slender, weak, often EAU RATOULS el syitof ite Aa) gins ignsids Ale Wa shes Ws ye ites dno ipsyike: RT Magnscibs Vo, xa, No Nfs de ME MANE Se en 16. Calyx covered with bristly hairs; corolla 21-22 mm long; pods 11-12 mm long, oblong-lanceolate; stems firm, strong, sulcate. a iA LU Se ae gE FW 454. A. perrarus Boriss. Calyx 8-10mm long, theteeth 4-5 mm long; corolla 13-18 mm long; pods 6-8 mm long; leaflets oblong or oblong-linear, 5-10mm long, 1-2mmbroad; plants25-40cmtall... 464. A. cancellatus Bge. Calyx 5-8 mm long, the teeth 2-4 mm long; corolla 12-13 mm long, pods ca. 4 mm long; calyxnarrowly linear, 4-5 mm long, ca. i HAAMIEOSMe! 225) SMe hee 465. A. pseudocancellatus Grossh. Calyxteethvery slightly shorter thanthe tube; leaflets 3-5 pairs, the lower ovate -spatulate, the upper oblong; plants 15-25 (40) cm tall; stems flexuous (mainly S. Transc.). 458. A. conspicuus Boriss. Plants differing an various seharaeters Ws 2 er... le let ere ors) 6 18. Hairs of the calyx, especially those ofthe teeth, borne ontubercles; leaflets 4-6 pairs; plants7-15cmtall. 463. A. teheranicus Boiss, Hairs of the calyx not borne’on'tubercles 2 7 er ee 19. Plantsusually erect, sometimes ascending; stems stout, (20) 25- 80cm long (rarely shorter), greenor white-pannose; leaflets 6-16, mostly 8-14 pairs; flowers usually 16-29mm rarely 13 mm long. 20. Ascending plants; stems usually slender, flagelliform, erect, of- ten densely branched, (7) 10-25 (30) cm long, greenish or covered with appressed silvery-gray hairs; leaflets (2)4-8 or 7-14 pairs; 359 20. 483 22. 23. 24. 25 (19). 26. 484 + flowers 9-16 mm, rarely up to 20 mm long............. 25. Plants densely shaggy-pannose with short white hairs, 30-60 cm tall; leaflets 7-30 mm long, 5-10 mm broad, pannose-pubescent, ovate to oblong-elliptic (Sands of Kara Kum, Ar.-Casp.)...... CLA ehh eo, EOS Woke, oie ay ek ane 449. A. albescens Boriss. Plants greenish or greenish-gray by appressed hairs; leaflets 7- 25 mm long, 1-3 mm rarely up to 4 (7) mm broad, oblong or nar- rowly. Subulate -linear{. «1 sen)! 70. jeloennane eebaede o «aes 21. Pods, and sometimes flowers, subhorizontally spreading; corolla violet; the lower part white ..... 451. A. sevangensis Grossh. Pods and flowers erect; corolla bright violet ........... 22. Bracts setiform; leaflets narrow, subulate-linear, up to 12 (15) pairs, (7) 10-15 mm long, 1-2 mm broad; pods 7-9 mm long... = iss ee oe cok obes kecyoch Ghevcmin oes ject 450. A. circassicus Grossh. Bracts lanceolate; leaflets oblong and oblong-elliptic to linear - lanceolate, 8-25 mm long, 1-4 (7) mm broad ........... a3. Leaflets oblong-lanceolate or lanceolate; pods ovoid, very slight - lysexcecdingythevcallyact ss \egepee soe 455.. A. goktschaicus Grossh. Leaflets oblong-elliptic to linear-lanceolate; pods 11-15 mm long, l.o=2 timesias lonpyas the Calyx) 2.5, si.’ «ftw syemebemans ie eon 24. Leaflets oblong-elliptic; calyx 8 mm long, sparsely black-hairy, the teeth more than half the length of the tube; pods oblong, 13- 15 mm long, the beak 2-3 mm long (E. and S. Transc.)...... jth diceectead 2).0) Him MONG esas ae eee ee, ole 902. A. arcuatus Kar. et Kir. Acaulescent plants, 5-8 cm tall; peduncles equaling or slightly exceeding the leaves, 2.5-4.5 cm long... 510. A. arganaticus Bge. Stems 5-10 cm long; peduncles considerably exceeded by the leaves, AN D CIRM LOMNGs Moe . Vey eel “nays aietemen- hey ie 508. A. rubrigalli M. Pop. Key to Species in Fruiting Condition Stems geniculately bent, brittle at the geniculations, 6-14 cm long; leaflets 5-8 pairs, oblong-ovate or elliptic, 5-10 mm long, 2.5-4 mm broad, densely covered with ascending hairs; inflorescence compact, 2-10-flowered; peduncles shorter than to equaling the leaves; pods subsessile, fusiformly lance-linear, straight, 30-40 mm long, covered with appressed white hairs, partially bilocular............... Be et Ca Oy CEO CLC Sry EOL C6 EC: cou ot sky O 507. A. infractus Sumn. Stems not bent, not brittle; plants differing in their characters from theyprecedin ge iSpeCie’s fess) <: Setendl ela erret of Tapodceeis Weil uence itor 2. Leaflets 9-14 pairs; acaulescent or nearly acaulescent plants; peduncles long; pods oblong to linear-oblong, falcate, shaggy with short, whiteystiffishhainsisos.uLioer heueiok 504. A. reduncus Pall. Wéafletse lial) pairsic... mpage wl s Bah fe Cyeaet).) cepa eee 3. Leaflets 2-2.5 mm long, 1-1.5 mm broad, 5-6 pairs, rounded -obovate, covered with appressed hairs....... 514. A. innominatus Boriss. Leaflets (3) 4-25 (30) mm long; plants differing in their characters. Pods covered with black and white hairs............0+e+eee8-s 5. Pods covered merely with white hairs 2. .2)6 6 o siss0. ey 000m Boeial« « 7. Pods linear, 22-30 mm, long, 272.5 mm broad,| faleatec ey. aa = bo bac, o oa Ceo oS \owesipic (sy tapeiatey sl sme oc) D01.6 Ad petracnsgitdragek Kar. Pods broadly linear, linear, or lance-oblong, straight or scarcely CUPV.EO, «icy. satel gencptargrstls .eo Genuine aden ore Lees folie, ee 6. Leaflets lanceolate or elliptic, (3) 4-6 pairs; pods broadly linear - oblong, 20-24 mm long, sulcate dorsally. 512. A. rarissimus M. Pop. Leaflets obovate-oblong or oblong-obovate, 3(4) pairs; pods lance- oblong, 10-17 mm long, not sulcate dorsally. .............+-+-.+-s riety boleh eiteerh Oleh s sees arrapin alas beh bes 513. A. alitschuri B. Fedtsch. Leaflets covered with appressed hairs. ........0++2eeeeeee0 8 396 532 333 + Leaflets covered with more or less spreading hairs........... 1s 8. Leaflets 1-2 pairs; acaulescent or nearly acaulescent plants, 5-7 cm ae ee ci) sonal a hake Au el tae akc i ta alae cat i Rite 511. A. Subbijugus Ldb. —~aepmeat leusn(2)*3—Gipairs!e). Leia Nees Pee ban Ph.dhle. Brine 9. 9. Calyx 17-18 mm long; leaflets orbicular, rounded-ovate, or elliptic Sr BOG Oo te RCL ey a ee Ba 505. A. ferganensis (M. Pop.) B. Fedtsch. + Calyx 11-15 mm long; leaflets lanceolate, linear-lanceolate, or OlOnNSROVOV Aten ce 2.6. ee ee eds See Pete CMO ete ee ei mes nee Ad (08 HOke Wearlets) lanceolate or linear -lanceolate 2. es ge ee te oe SEMEN CROMER aH. cl eM. eee be Riigee & 506. A. erioceras Fisch. et Mey. + Leaflets oblong-obovate........ 509. A. erioceratiformis M. Pop. 11. Subshrubs with a short divided woody caudex............... 76 SEMPIBCTVCTATINA SY © dee fea iette Cavities Kove, ade tae veterans io SNe ta MPa ee eee ete a V3" 12. Pods linear-oblong, 14-19 mm long, 5 mm broad; ovary borne on a Staperca.sOlo mrmmulongt 32. ek. Sea 503. A. subarcuatus M. Pop. + Pods linear, 17-27 mm long, 3-3.5 mm broad; ovary borne on a stipe A= 75'5) Fealved) hol ovey eG: celal eeR NSE IH ecules! ois ES 502. A. arcuatus Kar. et Kir. 13. Peduncles equaling or somewhat exceeding the leaves, 2.5-4.5 mm forees podstsulcate dorsally. oo. ws 510. A. arganaticus Bge. + Peduncles greatly exceeded by the leaves, 1-1.5 cm long; pods rounded LO TES OME ar ea Paliogs) fs) suis, eh ste dteych ctelsle Net caked «.(t 508. A. rubrigalli M. Pop. Series 1. HARPOCARPI Gontsch. — Pods linear or oblong -linear, falcate, subshrubs with short or sometimes well developed aerial divided caudices; corolla violet or purple. 501. A. petraeus Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 333; Bge. Astrag. geront. II, 221; Bge. Astrag. turk. 286.— A. xylorrhizus Bge. Astrag. turk. (1880) 267.—A. irkeschtensis B. Fedtsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS, VII (1940) 168. Perennial, suffrutescent, with a strongly abbreviated often obsolescent stout short -branched aerial caudex 0.5-2 (3) cm long and a strong buried caudexupto2.5cm in diam.; stems of the year (0.5) 1-5 (6) cm long, vested with white appressed hairs; stipules ovate-triangular, 3-4 cm long, acute, covered with appressed white and black or almost exclusively white hairs; leaves (2) 4-7 (8) cm long, the rachis 3-4 times as long as the petiole, both rachis and petiole covered with appressed white hairs; leaflets 4-7 pairs, elliptic or oblong-elliptic or rarely lance-elliptic, (5) 8-13 mm long, hoary on both sides by dense appressed hairs; peduncles 1.5-2 times as long as the leaves, finely covered with white or mixed white and black (beneath the inflorescence always white and black) appressed hairs; racemes short, fairly compact, subcapitate, rounded-ovoidto ovoid, 3-4 (7) cm long, 10-18-flowered, the flowers approximate, subsessile; bracts lanceolate, ca. 2mm long, covered with white and blackor rarely only white hairs; calyx cylindric, 8-10 mm long, shaggy with spreading short black and longer white hairs, the tube 3-4 times as long as the finely subulate teeth; corolla violet; standard 18-22 mm long, the limb oblong-obovate, slightly attenuate toward apex, scarcely retuse, shortly and angularly attenuate in lower part, 4 times as long as the claw; wing-petals 17-20 mm long, the limb oblong, entire, as long as the claw; keel 14-17 mm long, obtuse; pods sessile, linear, 2.2-3.0 cm long, 2-2.5 mm broad, subcylindric, rounded ventrally 397 534 and dorsally, pointed, falcate, coriaceous, shaggy with ascending or spreading short black and longer white hairs, bilocular, Fl. June (July); fr. (June) July. Desert mountain slopes and stony desert river terraces. — Central Asia: Dzu.-Tarb. (Dzungarian Ala Tau), Pam. Al. (eastern Alai valley), T.Sh. (Centr. and E.). Described from Sarkan river valley and Dzungarian Ala Tau. Type in Moscow; cotype in Leningrad. 502. A. arcuatus Kar. et Kir. in Bull. Soc. Mosc. XIV (1841) 407; Ldb. Fl. Ross. I, 631; Bge. Astrag.geront. II, 190; Bge. Astrag. turk. 2665) heey her Aen olan Wille Lo siap. Perennial, suffruticose, 7-15 cm tall, from a short strong woody caudex with slender woody decumbent divisions, these coated with yellowish -red bark, 2-6 (8) cm long; shoots of the year 3-7 (10) cm long, white, densely hispid with spreading hairs, fairly slender; stipules adnate to petiole up to 1/2-1/3, oblong-triangular, 3-4 mm long, densely covered with short spreading white hairs; leaves 1.5-2 cm long, the petiole equaling to twice as long as the rachis, white-tomentose; leaflets 2-3 (4) pairs, linear- lanceolate, lanceolate or oblanceolate, round-tippedor subacute, subcuneate at base, 5-10 mm long, 1.5-3 mm broad, sparsely shaggy tomentose above and densely sobeneath with short spreading hairs; peduncles somewhat shorter than to equaling the leaves, densely white-hispid, 1-2 cm long; racemes short (the rachis 3-6 mm long), subcapitately pointed, 2-3 cm long, o-7-flowered; bracts ovate-oblong to lanceolate, obtusish, 1.5-2 mm long, sparsely covered with white and predominantly black hairs, 1.5-2 times as long as the pedicel; calyx cylindric, 12-13 mm long, densely tomentose- villous with white and black spreading hairs, the subulate-linear teeth 2-3 mm long; corolla purple (?); standard 18-25 mm long, the limb oblong- obovate, angularly attenuate in lower part, 7-9 mm broad in upper part, 3 times as long as the claw; wing-petals 17-22 mm long, the limb linear- oblong, scarcely retuse, 2/3 as long as the claw; keel 16-19 mm long, obtusish; ovary on a stipe 2-2.5 mm long, densely hairy; pods sessile, linear, 17-27 mm long, falcate, compressed laterally, 3-3.5 mm broad, 2 mm thick, carinate ventrally, rounded-carinate dorsally, coriaceous, densely lanate with spreading short white hairs, pointed at both ends, bilocular. Fl. May (June); fr. June. Clayey and stony slopes, sandy and gravelly plateaux, needlegrass~- and- fescue steppes, and in steppe meadows on hillsides. — W. Siberia: U. Tob. (S.), Irt. (northern and southern parts); Centr. Asia: Ar.-Casp. (N.), Balkh. (N.), Gen. distr.: Mong. (W.). Described from the Irtysh (between Omsk and Semipalatinsk) and from the area between Dzhartash and Arkat. Type in Moscow. 503. A. subarcuatus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 7-12 cm tall, branched; caudex stout, the woody divisions short, fairly slender, subdecumbent, coated with yellowish- brown peeling bark; shoots of the year (1) 1.5-4 cm long, white, rather slender, densely tomentose with white spreading hairs; stipules triangular, acute, 1-1.5 mm long, covered with white or mixed white and black hairs; leaves 1-2 cm long, the petioles slender, equaling or exceeding the leaves, hispid with spreading hairs; leaflets 2-3 pairs, oblong-lanceolate to linear- lanceolate, acute, long-cuneate at base, 5-10 mm long, 2-3 mm broad, 398 535 hoary~-hispid sparsely above and densely beneath with spreading hairs, the uppermost pair close to the terminal leaflet; peduncles equaling to 1.5 times as long as the leaves, slender, tomentose with white spreading hairs, these interspersed beneath the inflorescence with few black hairs; racemes short (the rachis 0.5-1 cm long), subumbellate-capitate, 2-2.5cm long, 5-10- flowered; bracts triangular-ovate, acute, ca. 1.5 mm long, membranous- hyaline, black-hairy, 1.5 times as long as the pedicel; calyx cylindric, 11-12 mm long, densely villosulous with spreading white and black hairs, the teeth linear-subulate, 1.5-2 mm long; corolla purple (?); standard 20-23 mm long, the limb oblong-obovate, 6.5-9 mm broad in upper part, retuse, angularly short-attenuate at base, 2.5-3 times as long as the claw; wing -petals 17-20 mm long, the limb oblong, slightly dilated, round-tipped, scarcely retuse, the claw 1.5 times as long the limb; keel 15-18 mm long, rounded-obtuse; ovary on a stipe ca. 0.5 mm long, hairy; pods sessile, linear -oblong, 14-19 mm long, falcate, compressed laterally, 5 mm broad, 2.5-3 mm thick, pointed, terminating in a beak 1.5-2 mm long, carinate dorsally and ventrally, coriaceous, densely lanuginous-shaggy with spreading white hairs, bilocular. Fl. April-May; fr. May. Steppes, often on gravelly slopes.— Centr. Asia: Ar.-Casp. (Mugodzhary). Endemic. Described from Mugodzhary. Type in Leningrad. Series 2. MULTIJUGI Gontsch.— Acaulescent or nearly acaulescent perennials, with linear falcate pods and relatively many leaflet pairs; corolla ochroleucous or slightly suffused with violet. 504. A. reduncus Pall. Spec. Astrag. (1800) 109; Bge. Astrag. geront. II, 190; Ldb. Fl. Ross. I, 649; Schmalh. Fl. I, 280.—A. hamosus Pall. Reise III (1776) 653, nomen, non L.— A. cinereus y.longipes DC. Astrag. (1802) 222.—Euilus reduncus Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 149. Perennial, acaulescent or nearly so, 10-20 cm tall; caudex woody, buried, with strong short divisions; stems 0.3-1 cm long, densely lanate with spreading or ascending short white hairs; stipules adnate to petiole in lower 1/4-1/3, 5-7 mm long, ovate-lanceolate, acuminate, tomentose with spreading white hairs; leaves 6-12 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis finely hispidulous with spreading white hairs; leaflets 9-14 pairs, oblong-oval, rarely oval, slightly retuse or round-tipped, (3) 5-8 (10) mm long, shaggy tomentose sparsely above and densely beneath with spreading hairs; peduncles equaling or somewhat exceeding (rarely shorter than) the leaves, minutely tomentose with curly hairs; racemes loose, closely 8-20-flowered, nearly subulate or ovoid, 2.5-4 (6) cm long; bracts lance-linear, acute, 3-5 mm long, finely covered with white or mixed white and black hairs; calyx cylindric, 11-13 (14) mm long, villosulous with black and white short hairs, the tube 2-4 times as long as the linear-subulate teeth; corolla, ochroleucous or slightly suffused with violet; standard 19-22 (27) mm long, the limb oblong -obovate, retuse, attenuate and angular at base, 2.5-3 times as long as the claw; wing-petals 16-17 (21) mm long, the limb oblong, slightly retuse, equaling the claw or nearly so; keel 14 (19) mm long, obtuse; ovary ona stipe ca. 1 mm long, white-hairy; pods sessile, spreading, oblong or linear -oblong, 10-20 mm long, 3-5 mm broad, falcately upturned, carinate ventrally, 399 536 grooved or concavely sutured or rarely rounded dorsally, pointed, terminating in a straight rigid beak 1.5-3 (5) mm long, coriaceous, hispidulous with short white hairs, bilocular. Fl. April-May (June); fr. May-June. Steppes and semideserts. — European part: Bl., Crim., L. V. Endemic. Described from Lower Volga. Type in London; cotype in Leningrad. Series 3. ERIOCERATI Gontsch. — Nearly acaulescent or short-stemmed perennials with few leaflet pairs; pods linear-oblong, rarely oblong -oval or oblong-lanceolate, more or less curved, rarely almost straight or straight; corolla violet. 505. A. ferganensis (M. Pop.) B. Fedtsch. ex A. Korol. in Fl. Tadshik. V (1937) 414.—A. erioceras var. ferganensis M.Pop. in Sched. HeF. ALM. fase: I (1924) 13.— bhxs.- H. fF. A.M. No. 8. Perennial, 5-20 cm tall, appressed-canescent, froma short woody buried caudex; stems 1-3 (6) cm long, ascending or erect, vested with appressed white hairs, grayish-white; stipules triangular, rarely triangular - lanceolate, 2-4 mm long, covered with white and black hairs; leaves 2-6 cm long, the petiole (0.5) 1-2.5 cm long, both petiole and rachis slender, appressed-canescent; leaflets (3) 4-5 (6) pairs, orbicular, rounded-ovate, or elliptic, 6-14 mm long, 5-9 mm broad, acute or obtuse, minutely mucronulate, densely covered on both sides with appressed hairs; peduncles equaling to 1.5 times the length of the leaves, (2.5) 4-6 (8) cm long, covered with white appressed hairs; racemes 3-5 cm long, subumbellate, loosely 3-11-flowered; bracts lanceolate, 3-4 mm long, about equaling the pedicel, hyaline, covered with black and white hairs; calyx cylindric, subsaccate at base, 17-18 mm long, villosulous with white and black hairs, the tube 4-6 times as long as the linear -subulate teeth; corolla violet -pink; standard 30-32 mm long, the limb obovate, retuse, slightly constricted in lower part, 3 times as long as the claw; wing-petals ca. 29 mm long, the limb oblong-oval, slightly retuse, 8-9 mm long; keel 27 mm long, obtusish; pods borne on a very short stipe, subsessile, oblong to linear -oblong, 22-25 mm long, straight or subfalcate, more or less compressed laterally or subtriquetrous, 5-9 mm broad, slightly carinate or almost rounded ventrally, scarcely and narrowly grooved dorsally, terminating in a slender reflexed beak 1-1.5 mm long, coriaceous, densely shaggy with white hairs; bilocular, many-seeded. Fl. March-April; fr. April-May. Gravelly and stony sites on submontane plains and in the foothills. — Centr. Asia: Syr D., Pam.- Al. (Sarytau mountains, Shurab), T. Sh. (Mogoltau mountains). Endemic. Described from Santo in the northern _ foothills of the Turkestan Range. Type in Tashkent; cotype in Leningrad. 539 506. A. erioceras Fisch. et Mey. ex Ldb. Fl. Ross. I (1842) 626; Bge. Astrag. geront. II, 190; Bge. Astrag. turk., 567. Perennial, 6-15 (18) cm tall, from a strongly abbreviated woody buried caudex, the short divisions forming small blunt-topped mats; stems (4) 5-9 (12) cm tall, erect or subassurgent, rather slender, canescent with appressed hairs; stipules triangular-ovate, acutish, 2-3 mm long, covered with appressed white hairs; leaves 3-4 (6) cm long, the petiole as long as the rachis, slender, canescent withappressed hairs; leaflets (2) 3-4 (5) pairs, lanceolate to linear-lanceolate, 7-17 mm long, 2-3 mm broad, acute, canescent sparsely above and densely beneath with appressed hairs; racemes 400 ) . ma @ WW ish yk ce e NC MOE? . S B ayy" u ae ; ne iy Ss qT FC Moy 22>. aoa PLATE XXXV 1. Astragalus Borodinii Krassn.—2. A. Chomutovii B. Fedtsch.—3. A. ammodytes Pall. 401 540 short, loose, subumbellate, 4-6 (7)-flowered, 2.5-3 cm long, the rachis 5-10 (17) mm long; bracts oblong or the lower ovate, 1.5-2 mm long, subhyaline, sparsely black-ciliate; pedicels 1-1.5 mm long; calyx cylindric, 11-14 mm long, densely tomentose with ascending white and black hairs, the linear-subulate teeth 1.5-2.5 mm long; corolla violet (?); standard 20-25 mm long, the limb oblong-obovate, scarcely retuse, scarcely auriculate-angular in lower part, cuneately attenuate into and 1.5-2 times as long as the claw; wing-petals 19-24 mm long, the limb broadly oval- oblong, obtusish, the claw 3 times as long as the limb; keel 18-28 mm long, round-tipped; ovary on a stipe ca. 1 mm long, white-shaggy; pods borne on a stipe ca. 1 mm long, linear-oblong, rarely linear, 15-23 (30)mm long, compressed laterally, 4-5 mm broad, 2.5-3 mm thick, pointed at both ends, slightly arcuate (rarely straight), terminating in a reflexed beak 1.5-2 mm long, carinate ventrally, grooved dorsally, rigidly coriaceous, densely hispid-villous with spreading hairs, almost fully bilocular, many- seeded. Fl. May-June; fr. May-June. Region of white wormwood-deserts, on gravel-and-clay hillsides, sandhills, chalk, on gritty, gravelly and stony slopes of low mountains, and on outcrops of hard mottled rocks.— Centr. Asia: Ar.-Casp., Kyz. K. (lower course of Syr Darya). Endemic. Described from the eastern shore of the Caspian Sea. Type in Leningrad. 507. A. infractus Sumn. in Animadvers. Syst. Univ. Tomsk. 9-10 (1936) 4. Perennial, from a buried caudex 3-5 mm thick; stems geniculately bent, readily breaking at the geniculations, 6-14 cm long, 1-2 cm thick, densely appressed-tomentose, white; stipules free, triangular -ovate, 1.5-2 mm long, densely covered with white and black appressed hairs; leaves 3-5 cm long, the rachis 5-6 times as long as the petiole; leaflets 5-8 pairs, oblong ovate or elliptic, short-acuminate, 5-10 mm long, 2.5-4 mm broad, densely covered with ascending hairs; peduncles shorter than to as long as the leaves; inflorescence compactly 2-10 -flowered; bracts equaling or exceeding the pedicel, ovate-lanceolate, covered with white and black hairs; calyx cylindric, 13-15 mm long, faintly covered with ascending short subappressed black hairs, the teeth 2-3 mm long; standard 22-26 mm long, oblong-obovate, 7-9 mm broad, retuse, somewhat attenuate in upper one-third; wing-petals ca. 24 mm long, the claw 3 times as long as the limb; ovary on a stipe 1-2 mm long; pods subsessile, fusiformly oblanceolate or linear, straight, 30-40 mm long, partially bilocular, covered with white appressed hairs. Riverside pebbles. — Centr. Asia: Tien Shan (N.) Endemic. Weboribed from the vicinity of Mount Tomsk. Type in Tomsk. 508. A. rubrigalli M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 10-15 cm tall: stems herbaceous, 5-10 cm long, geniculately bent, densely hispidulous with short white spreading hairs; stipules triangular -lanceolate, acute, 2-3 mm long, slightly tomentose; leaves 5-7 cm long, the petiole 0.5-1 cm long, both petiole and rachis hispid; leaflets 5-7 pairs, obovate, (5) 8-14 mm long, 0.4-0.8 mm broad, obtuse or terminating in a very short point, covered above with scattered subappressed hairs and beneath with divergent stiff hairs, hoary; peduncles 1-1.5 cm long, hispid; racemes elongate, 3-4 cm long, rather compact; 402 C a bracts translucent hyaline, lanceolate, 2-3 mm long, sparsely covered with white hairs; calyx cylindric, 15-17 mm long, covered with white stiff and predominantly spreading hairs, the setaceous teeth 3-5 mm long; standard 25 mm long, the limb oblong-obovate, ovally dilated in upper part, 6.5 mm broad, angularly attenuate to 4.5 mm in lower part, retuse at apex, 1.5 times as long as the claw; wing-petals 23 mm long, the limb oblong, round -tipped, half as long as the claw; keel 22 mm long; pods sessile, broadly linear, 25-30 mm long, nearly round in cross-section, arcuate, 3-4 mm broad, terminating in an uncinate beak 3-4 mm long, coriaceous, hispid with ascending long white hairs, rounded dorsally and ventrally, bilocular. Fr. May. ’ Centr. Asia: AmuD. (vicinity of Kelif). Endemic. Described from the vicinity of Kyzyl-khuraz village. Type in Tashkent. 909. A. erioceratiformis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 6-7 cm tall, from a rather slender vertical root; stems 1-2 cm long, almost lanate with subappressed white hairs, angled; stipules broadly triangular, ca. 0.5 mm long, covered on the outside with stiff white. hairs; leaves 3-6 cm long, rachis as long as petiole, hoary with stiff appressed hairs; leaflets 3-4 pairs, oblong-obovate, 8-12 mm long, 4-6 mm broad, obtuse to subacute, rather sparsely covered on both sides with long appressed hairs, scabrous-hoary; racemes 1-2 cm long, 5-10-flowered, hispid with appressed white hairs; the rachis tomentose with white and black hairs; bracts lanceolate, ca. 5mm long, covered with black and white hairs; calyx cylindric, ca. 15 mm long, covered with subdivergent white and on the ventral face with black divergent hairs, the teeth filiform, ca. 3mm long, blackish; corolla probably purple; standard 25 mm long, the limb oblong-obovate, ovally dilated in upper part, 6 mm broad, retuse; wing-petals equaling the standard, the limb oblong, obtuse, half as long as the claw; keel 24 mm long; pods straight, broadly linear, 20 mm long, 4-5 mm broad, sessile, carinate ventrally, broadly grooved dorsally, terminating in a short oblique point, bilocular, sparsely pubescent with. subappressed hairs. Fr. July. Centr. Asia: Tien Shan (N. W.). Endemic. Described from Mt. Ul'kul burul' in the Dzhambul district. Type in Tashkent. 510. A. arganaticus Bge. ex Rgl. et Herd. in Bull. Soc. Nat. Mosc. XXXIX (1866) 111; Astrag. geront. II, 190; Astrag. turk., 266. Perennial, acaulescent, 5-8 cm tall, from a woody very short caudex, the weak short divisions forming small mats; stipules adnate to petiole up to 1/2-2/3, 3-6 mm long, densely white-hispid, the free portion oblong to broadly ovate, obtusish; leaves 2.5-4 cm long, the petiole as long as the rachis, both sparsely villosulous with short white hairs; leaflets 5-7 pairs, obovate, or oblong-obovate to rounded-oval, (4) 6-10 mm long, round -tipped and sometimes mucronulate, (2.5) 4-5 mm broad, subcuneate at base, approximate, hoary, villosulous-hispidulous sparsely above and densely beneath with spreading hairs; peduncles equaling to somewhat exceeding the leaves, 2.5-4.5 cm long, densely villosulous with ascending hairs; racemes short, subumbellate, rather compactly 6-8-flowered, the rachis ca. 0.5 cm long; bracts linear to linear-lanceolate, 3-4 mm long, covered with white and black hairs; pedicels 0.5 mm long; calyx cylindric, 13-15 mm long, densely villosulous with white and black spreading hairs, 403 542 the linear teeth 2-3 mm long; corolla purple (?); standard 22-28 mm long, the limb oblong -obovate, short~attenuate and retuse at apex, gradually attenuate below into and twice as long as the claw; wing-petals 20-23 mm long, the limb oblong, slightly curved at the tip, half as long as the claw; keel 19-22 mm long, rounded-obtuse; ovary on a stipe ca. 2 mm long, hairy; pods sessile, linear, 2.5-3 cm long, 3.5-4 mm broad, falcate in upper half, acuminate to a subulate beak 2-3 mm long, more or less compressed laterally, carinate ventrally, sulcate dorsally, coriaceous, villosulous with short spreading hairs, bilocular. Fl. May; fr. May-June. Rocky mountain slopes.— Centr. Asia: Balkh. (Arganaty mountains). Endemic. Described from Arganaty mountains. Type in Leningrad. 511. A. subbijugus Ldb. Fl. Ross. I (1842) 627; Bge. Astrag. geront. II, 190; Bge. Astrag. turk. 268.— A. cygneus Fisch., herb. —lIc.: Bge. Ic. Pl. Nov. Lehm. tab. 13 f. 1 (1849). Perennial, acaulescent or nearly so, 5-7 cm tall, from a stout strongly abbreviated buried caudex, the very short divisions forming compact mats; stems obsolete to 2 cm long, densely vested with appressed hairs, white, triangular, acute, 2-2.5 mm long, white-hairy; leaves 2-4.5 cm long, the petiole 1.5-2 times as long as the rachis, both petiole and rachis slender, densely covered with short appressed hairs; leaflets 1-2 pairs, elliptic, rarely obovate, short~acuminate to rarely obtuse, 7-15 (20) mm long, covered on both sides with appressed hairs, hoary; peduncles many, 0.5-1.5 cm long, densely covered with appressed hairs, white; racemes short (the rachis ca. 0.5 cm long), loose, 3-4 cm long, 3-5-flowered; bracts lanceolate, ca. 2.5mm long, covered with white and black hairs; pedicels 1.5-2 mm long; calyx long-cylindric, 17-18 mm long, densely hispid-villous with black and white hairs, the teeth subulate-linear, 4 mm long, the tube 3-3.5 times as long as the teeth; corolla violet; standard 30 mm long, the limb obovate, oblong, ovately dilated upward, 9 mm broad, angularly narrowed downward to 7 mm and then cuneately attenuate into and twice as long as the claw, retuse at summit; wing-petals 28 mm long, the limb oblong, round-tipped, the claw 2-3 times as long as the limb; keel 26 mm long; ovary on a stipe ca. 1 mm long, hairy; pods broadly oblong, 20 mm long, borne on a stipe ca. 1 mm long, compressed laterally, 7 mm broad, 5 mm thick, carinate ventrally, finely sulcate dorsally, terminating in a reflexed beak ca. 1.5 mm long, rigidly coriaceous, densely shaggy with white stiff hairs, incompletely bilocular. Fl. April-May; fr. May. ; Rock crevices and outcrops of mottled layers.— Centr. Asia: Kyz. K. (residual mountains of Khan-ata and Aktau and occasionally on sand-blown outcrops of yellowish-red layers of Torb-kuduk, Koz~-kuduk, Mount Karak). Endemic. Described from S. Kyzyl Kum. Type in Leningrad. Series 4. RARISSIMI Gontsch. — Acaulescent or nearly acaulescent perennials; flowers yellow; pods linear-oblong, scarcely curved. 512. A. rarissimus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent or nearly so, 15-18 cm tall; divisions of the short buried caudex stout, closely approximate, scaly by vestiges of petioles and white -pannose between the scales; stems obsolete or up to 1-2 cm long, stout, covered with pannose pubescence; stipules up to 10 mm long, rather 404 sparsely covered with white hairs, the free portion partially membranous, the lower ovate, the upper lanceolate and relatively long-acuminate; leaves 10-20 cm long, the petiole 2-4 cm long, both petiole and rachis hispid with spreading hairs, stoutish; leaflets (3) 4-6 very distant pairs, lanceolate to elliptic, acute, (10) 15-25 (30) mm long, densely covered on both sides with appressed white stiff hairs; peduncles 7-15 cm long, shorter than the leaves (together with inflorescence 10-20 cm long and equaling the leaves), covered with ascending white hairs and upward subvillous with white and black hairs; racemes subumbellate, ovaloid or spherical, 3-5 cm long; bracts lance-linear, 5-10 mm long, hispid-villous with white and black hairs; calyx broadly cylindric; 15-17 mm long, densely hispid-villous with black and white hairs, the filiform-linear teeth 5-6 mm long half the length of the tube; corolla yellow; standard ca. 27 mm long, the limb obovate, deeply retuse, 14 mm broad, shortly and angularly attenuate toward base, 4 times as long as the claw; wing-petals ca. 21 mm long, the limb oblong, slightly curved, obtuse, as long as the claw; keel 19 mm long, acute; ovary on a stipe ca. 1 mm long, hairy; pods broadly linear - oblong, 23-24 mm long, almost fully sessile, compressed laterally, 9-11mm broad, ca. 5 mm thick, carinate ventrally, sulcate dorsally, rigidly coriaceous, densely lanate-villous with white and fewer black long spreading hairs, terminating in a reflexed firm beak ca. 2 mm long, bilocular. Fl. June; fr. July. Mountain slopes.—Centr. Asia: Tien Shan (Dzhumgol' district). Endemic. Described from Kovaktau. Type in Tashkent. Series 5. BRACHYCARPI Gontsch.—Nearly acaulescent perennials; flowers yellow; pods short, lance-oblong, straight. 513. A. alitschuri O. Fedtsch. Pl. Pamir. (1904) 28; Fl. Tadzhik. V (1937) 415. Nearly acaulescent plants, (8) 10-20 cm long, from a short woody buried caudex, the short divisions forming loose mats; stems of the year long, flexuous, densely vested with short appressed hairs; stipules triangular to triangular -lanceolate; 1.5-2.5 mm long, acute, covered with black and white hairs; leaves (2) 3-5 cm long, the petiole as long as or longer than the rachis, canescent by short appressed hairs; leaflets (3) 4 pairs, obovate -oblong or oblong-obovate, rarely ovate or oblong, 4-7 (9) mm long, round -tipped or obtuse, canescent on both sides by short appressed hairs; peduncles (4) 5-10 cm long, covered with short appressed black and white hairs; racemes short, subumbellate, loosely (2) 3-5 (7)-flowered; bracts lanceolate, ca. 1.5 mm long, covered with white and predominantly black hairs; pedicels ca. 1 mm long; calyx cylindric, 12-13 mm long, densely covered with short black hairs, these sometimes interspersed with appressed white hairs, the linear-subulate teeth 1-2 mm long; corolla yellow; standard 19-20 mm long, the limb oblong-elliptic, subobtuse, 8-9 mm broad, gradually attenuate into a claw ca. 7 mm long; wing-petals 18-19 mm long, the limb broadly oblong, round-tipped, somewhat shorter than the claw; keel 14-15 mm long, obtuse; ovary subsessile, hairy; pods sessile, lance-oblong, 10-17 mm long, compressed laterally, 3.5-4mm broad, 2-2.5 mm thick, not grooved dorsally or ventrally, terminating in a mucronate beak 1-1.5 mm long, thinly coriaceous, densely lanate with long spreading hairs, these predominantly white on the sides 405 545 and black on the sutures, notably the dorsal one (hence the pod rather broadly striped on the back), bilocular. Fl. June-July; fr. August. Stony mountain slopes, infrequent on stony moraines and terraces. — Centr. Asia: Pam.-Al. (Pamir). Endemic. Described from Alichur river valley. Type in Leningrad. 514. A. innominatus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS WALL, io LOA 8 ss7)) AsO. Perennial, from a long strong woody root, ca. 10 cm tall, the divisions of the root-~crown forming dense mats; stems short, up to 5 cm long, vested with short appressed bifurcate hairs; stipules cauline, connate at base, small, ca. 2mm long, triangular, acutish, covered on the outside with white hairs; leaves 2-2.5 cm long, short-petioled; leaflets 5-6 pairs, rounded obovate, small, 2-2.5 mm long, 1-1.5 mm broad, scarcely subacute, cuneate-based, short-petioluled, covered on both sides with appressed hairs; peduncles axillary, twice as long as the leaves, covered with appressed white hairs; bracts oblong-ovate, acute, ca. 2 mm long, covered with predominantly black spreading hairs; flowers sessile; racemes densely capitate; calyx cylindric, ca. 12 mm long, at length becoming inflated and distally contracted, covered with spreading long white and black hairs, the teeth lance-triangular, unequal, ca. 1 mm long; corolla drying pale yellow; standard 18 mm long, the limb retuse, oblong-ovate, gradually passing into the cuneate claw; wing-petals equaling the standard, the limb 6 mm long, retuse and dilated at apex, short-auriculate at base; keel 16 mm long, the limb 4 mm long, obtusish; ovary sessile, lanceolate, white- shaggy, the style long, naked; pods unknown. Fl. July-August. Dry valleys, amidst cone debris, on slopes, among wormwood associations. — Centr. Asia: Pam.-Al. (Pamir). Endemic. Described from Pamir. Type in Leningrad. Section 77. TAMIAS Bge. Astrag. geront. I (1868) 110. — Caulescent herbaceous perennials, vested with simple and bifurcate spreading hairs; stipules adnate to petiole in lower part, not connate; leaves imparipinnate, the petioles marcescent; racemes loose, elongate; bracteoles none; calyx narrowly cylindric; ovary short-stipitate; pods short-stipitate, subhorizontally spreading, linear-oblong, compressed laterally, carinate dorsally and ventrally, coriaceous, bilocular.— Related to the section Erioceras, widely distributed on the sands of Central Asia. 515. A. Turezaninovii Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 342; Bge. Astrag. geront. II, 191; Bge. Astrag. turk. 282.— Exs.: He Ar Mia NOnm oo Shi oHegky NORE GiEG). Perennial, (11) 15-30 (40) cm tall; stems (3) 5-13 (20) cm long, firm, densely lanate with spreading short white hairs; stipules triangular - lanceolate, acuminate, (3) 5-7 (8) mm long, green, covered with stiff white hairs; leaves (5) 7-12 cm long, the petiole and the much longer rachis covered with short spreading stiff hairs; leaflets (5) 6-9 pairs, lanceolate or elliptic, acute and short-cuspidate, (5) 8-15 (20) mm long, rigid, glabrous above, covered beneath with stiff ascending hairs, the pairs distant; peduncles 2-8 cm long, 1/2-2/3 as long as the leaves, lanate with stiff spreading hairs; racemes loose, very remotely (6) 9-17-flowered, (6) 8-15 cm long, the flowers spreading, finally subcernuous; bracts 406 (546 ovate -lanceolate to lanceolate, acuminate, (2) 3-3.5 mm long, hispid with white or mixed black and white hairs; pedicels 1.5-2 mm long, hairy; calyx 15-18 mm long, densely hispid -villous with black and white or rarely exclusively white spreading hairs, mostly profusely covered with black hairs on the nerves, hence striped, the tube 3.5-4 times as long as the subulate-linear teeth; corolla yellow; standard 21-25 mm long, the limb obovate, 7.5-8 mm broad, slightly retuse, 1.5 times as long as the claw; wing -petals 20-21 mm long, the limb oblong, round-tipped, half as long as the claw; keel 19-20 mm long, the limb arcuately convex, very slightly concave above, rounded-obtuse, 2.5 times as long as the claw; ovary on a stipe 1.5-3 mm long, white-hairy; pods borne on a stipe 2-3 mm long, mostly slightly rarely falcately curved or straight, pointed (11) 15-23 mm long, 4-4.5 mm broad, 2.5-3 mm thick, terminating in a reflexed beak 2-4 (5) mm long, densely villous with spreading white hairs. Fl. April- June; fr. May-June. Fixed sands and sandhills, red sandstone and its derivatives, and sand- covered takyrs, in the desert or rarely semidesert region. — Centr. Asia: Aria Gasp.,; Balkh.'SyrjD,, Kyz. K.;Karak.,, Amu:D:))Pami7Al. (along river Amu Darya and in the lower reaches of its tributaries as far as river Vakhsh). Endemic. Described from E. Kazakhstan (between Sasyk-pastau and Arganaty Mountains). Type in Leningrad. Section 78. HELMIA Bge. Astrag. geront. I (1868) 111. — Acaulescent or nearly acaulescent cespitose dwarf perennials, vested with bifurcate hairs; stipules connate; inflorescence few-flowered or capitate-umbellate, borne on a rather short peduncle; calyx campanulate-cylindric; corolla yellow or purple; pods subsessile, oblong-ovoid to linear -oblong, compressed laterally, carinate ventrally, narrowly grooved or flattened dorsally, coriaceous, densely lanate villous with spreading hairs, bilocular. Ural, Kazakhstan region of low rounded hills, Altai, Tarbagatai, Dzungarian Ala Tau, central Tien Shan. 1. Pods linear-oblong, 18-20 mm long, 3-5 mm broad; leaflets (1) 2-4 UG) OTIS IS, dota lhe a cay Mura U ES ak eS i ae i eh eR a RTS MR Are eps 2p + Pods ovoid-oblong to ovoid, 10-15 mm long, 5-6 mm broad; leaflets ch) FONDUS OS aill alM OD Daa LN Na RRA OY 519. A. Helmii Fisch. Pre COLO larOochicoOleucCOusis | isis © cialis leer 516. A. depauperatus Ldb. SE OCGA MOLD OR (OUTS C tre Fon ect cays de alte ol «ea Poltonic (alpen ol velo urate, Cae, colle Si Seleeailens iic4 mim LOMO Ya. i ey ites ees 518. A. heptapotamicus Sumn. sp LoySEhSBESGelaw Onaaveoulleveven\ Aids Aisle ti Bug aaiewomo Me 517. A. Polakovii M. Pop. 516. A. depauperatus Ldb. Fl. Alt. III (1831) 314; Ldb. Fl. Ross. I, 612; Kryl. Fl. Zap. Sib. VII, 1681.— A. eriolobus Bge. in Mém. Sav. Etr. Acad. Pétersb. II (1835) 591. Perennial, forming loose small mats, (3) 6-11 (15) cm tall; buried shoots many, branched, prostrate, woody; aerial shoots of the year strongly abbreviated, 0.5-3 (4) cm long, densely leafy and densely vested with appressed white hairs; stipules adnate below to petiole, connate up to 0.6 mm, 4-5 mm long, membranous, covered with appressed white or mixed white and black hairs, the free portion lanceolate, acute; leaves 3-7 cm long, the petiole 1/2 to 2/3 as long as to equaling the rachis, both petiole and rachis densely covered with silvery appressed hairs; leaflets 407 547 2-4 (6) pairs, lanceolate to oblong-elliptic, 5-10 mm long, 1.5-3 mmbroad, acute, densely covered with silvery appressed hairs; peduncles 3-6 cm long, equaling or exceeding or rarely shorter than the leaves, sometimes elongating in fruit up to 10 cm, covered with appressed white hairs; racemes loose, 2-2.5 cm long, subumbellate, 3-5-flowered, the rachis 2-5 mm long; bracts ovate-lanceolate, acuminate, membranous, 2-2.5 mm long, covered with white hairs; pedicels 1-1.5 mm long, white-hairy; calyx cylindric or campanulate-cylindric, 9-12 mm long, covered with white and black appressed hairs, the tube 2-3 times as long as the linear -subulate teeth; corolla ochroleucous; standard 18-22 mm long, the limb oblong -obovate, retuse, 8-9 mm broad at apex, angularly short-acuminate toward base, 5-6 times as long as the claw; wing-petals 15-18 mm long, the limb linear - oblong, scarcely retuse, as long as the claw; keel 13-16 mm long, the limb obtusish, semicircularly gibbous beneath, densely hairy above, the claw 1.5 times as long as the limb; ovary on a stipe ca. 2 mm long, white- hairy; pods borne on a stipe ca. 1 mm long, linear-oblong, straight, rarely scarcely curved, pointed at both ends, 18-20 mm long, 4-4.9 mm broad, compressed laterally, 2-2.5 mm thick, carinate ventrally, narrowly grooved or canaliculate dorsally, coriaceous, densely hispid-lanate with spreading white hairs, bilocular, 2.5-3 times as long as the calyx. Fl. May- July; fr. June-July (August). Saline desert steppes and gravelly and stony slopes of hills and mountains.— W. Siberia: Alt.; E. Siberia: Ang.-Say. (Minusinsk, village Beiskoe). Endemic. Described from Altai. Type in Leningrad. 517. A. Polakovii M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, from a stout short divided root-crown, cespitose, nearly acaulescent; stems short, not more than 5 cm long, densely tomentose with appressed white hairs; stipules triangular-ovate, cauline, somewhat connate, acuminate, pubescent, hyaline; leaves thin, long-petioled, 2-4 cm long; leaflets 1-2 or rarely 3-4 pairs, silky canescent, oblong or oblanceolate, short-acuminate, or subobtuse and mucronate, ca. 10 mm long, the terminal leaflet equaling the others; peduncles exceeding the leaves or scarcely so, firm, together with inflorescence 5-7 (12) cm long, appressed-pubescent, bearing 2-4 flowers at the end; flowers strongly subumbellately approximate, erect; bracts short, ovate, acuminate, tomentose; calyx narrowly cylindric; commonly covered with appressed black pubescence, 9 (11) mm long, the subulate teeth 2-3 mm long; corolla pink; standard ca. 20-22 mm long, broad, scarcely notched, the claw broadly ovate; wing-petals subentire, minutely auriculate, the limb almost as long as the claw; keel 14-15 mm long; ovary 23-ovuled; pods bilocular, ca. 18 mm long, oblong-linear in side view, 3.5 mm broad, compressed, carinate ventrally, flattish dorsally, pointed at the end, attenuate at base into the stipe, covered with white hairs; seeds 2-2.2 mm long, angularly ovoid, compressed, olivaceous-green. June-July. Gravelly slopes and needle-grass steppes. — Centr. Asia: Dzu.~Tarb. : Dzhungarian Ala Tau, Sarkan river valley, between Kopal and Sarkand (eastward to Kopal). Endemic. Described from Mount Suok-tau. Type in Alma Ata. 518. A. heptapotamicus Sumn. in Animadvers. Herb. Univ. Tomsk (1933) No. 5-6, page 7. 408 Perennial, acaulescent, dwarf, compactly pulvinate-cespitose, 2-4 cm tall, from a strong profusely divided buried caudex; stipules connate to high up, 2-2.5 mm long, ovate, covered with appressed white hairs; leaves 0.5-1.5 cm long, the petiole about as long as the rachis, both tomentulose with appressed white hairs; leaflets 2-4 pairs, obovate to oblong-obovate, 2-3 mm long, densely covered on both sidés with silvery appressed hairs; peduncles equaling to 1.5-2 times as long as the leaves, white -tomentose; flowers (1) 2-5, approximate in an umbelliform inflorescence; bracts ovate, subhyaline, ca. 1.5 mm long, equaling the pedicel; calyx campanulate- cylindric, 8-10 mm long, densely covered with appressed black and few white hairs, the tube 3-4 times as long as the linear-subulate teeth; corolla pale purple; standard 17-25 mm long, the limb broadly elliptic, round- )48 tipped and scarcely retuse, 4-5 times as long as the claw; wing-petals 15-22 mm long, the limb linear-oblong, scarcely retuse, as long as the claw; keel 13-20 mm long the limb broadly semiovate, acute, the claw 1.5 times as long as the limb; ovary short-stipitate, 1.5-2 mm long; pods subsessile, gradually attenuate toward base into an obsolescent stipe ca. 1 mm long, narrowly linear-oblong, acuminate, ca. 20 mm long, 3 mm broad, slightly compressed laterally, carinate ventrally, narrowly canaliculate dorsally, gradually attenuate toward the subulate ca. 1-mm- long beak, coriaceous, densely villous with spreading white hairs, bilocular. Fl. June-July; fr. June-August. Stony and argillaceous mountain slopes.— Centr. Asia: Balkh. (northern shore of Lake Balkhash), Dzu. Tarb. (Tarbagatai, Dzhungarian Ala Tau). T. Sh. (Centr.). Endemic. Described from Mt. Suyun-tyube near Mt. Kopal, upper Lake Son~Kul!' and Bel' Bulak mountain range. Type in Leningrad. 519. A. Helmii Fisch. in DC. Prodr. II (1825) 301; Ldb. Fl. Ross. I, Ciisrakneviegkhls Zap. sib. Vil, 1682°— A. permiensis €. A.M. ex Rupr: Bre SoremUirals (lo0) G2. le. : Rupr. Ic. tab) ll. Perennial, acaulescent or nearly so, loosely cespitose, 6-13 cm tall, from a stout buried caudex with short divisions, stems 0.5-1.5 cm long, densely leafy, densely tomentose with white appressed hairs, silvery; stipules adnate at base to petiole and connate, 3-5 mm long, lanceolate, acuminate, densely covered with white or mixed white and black appressed fine hairs; leaves (3.5) 5-7 cm long, the petiole 1/3-1/2 as long as to equaling the rachis, both petiole and rachis slender, tomentose with appressed or subappressed hairs; leaflets 4-7 pairs, elliptic, acute, 7-15 mm long, 3-8 mm broad, covered on both sides with silvery appressed hairs; peduncles equaling the leaves, finely covered with appressed white hairs; inflorescence compactly 5-8-flowered; 2-3 cm long; bracts lanceolate to linear -lanceolate, 2.5-4 mm long, somewhat exceeding to twice as long as the pedicel, white- or white- and black~-ciliate; calyx campanulate- cylindric, 8-10 mm long, densely covered with white and fewer black short appressed hairs, the tube 2-2.5 times as long as the filiform -linear teeth; corolla ochroleucous; standard 21-26 mm long, the limb oblong -obovate or obovate -oblong, round-tipped and scarcely retuse, 4 times as long as the broad claw; wing-petals 18-22 mm long, the limb linear -oblong, entire, as long as the claw; keel 15-19 mm long, the limb obovate, obtusish, the claw somewhat more than 1.5 times as long as the limb; ovary on a stipe ca. 1 mm long; pods borne on a stipe ca. 1 mm long, compactly and capitately crowded, ovoid-oblong, or oblong-ovoid, 10-15 mm long, 409 549 550 5-6 mm broad, compressed laterally, carinate ventrally, narrowly flattened or subcarinate dorsally, pointed, terminating in a subulate beak 1-2 mm long, coriaceous, densely pannose and lanate-villous with spreading hairs, bilocular. Fl. June; fr. June-July. Stony slopes, exposed sites over limestone, mostly in stony steppe associations. — European part: Transv., V.-Ka. (from Ural to lower reaches of the Vishera in the north); W. Siberia: U. Tob., Irt. (Kokchetav, Borovoe, Loktevskoe). Endemic. Described from vicinity of Voskresenskii plant in the southern Urals. Type in Geneva. Section 79. PICROPHACE Bge. Astrag. geront. I (1868) 111; II, 192.— Gen. Picraena Stev. in Bull. Soc. Nat. Mosc. IV (1832) 265. — Perennials; stems of the year well developed; plants glabrous except for sparse pubescence on young petioles consisting of bifurcate hairs; stipules dimorphic, the lower fully connate, scalelike, the upper (foliar) green, herbaceous, connate merely at base; leaves imparipinnate, the petioles and rachises rather long-persistent; bracteoles none; calyx campanulate- cylindric, not becoming inflated in fruit; corolla pink; racemes axillary, long, loose; pods subsessile, inequilaterally oblong, broadly linear -oblong, inflated, carinate ventrally, finely canaliculate dorsally, rigidly coriaceous, medium sized, bilocular.— A monotypic section, associated with the Aralo-Caspian, mainly sandy, deserts. 520. A. amarus Pali. Astr. (1800) 8; Ldb. Fl. Ross. I (1842) 626; Bge. Astrag. geront. II, 192; Bge. Astrag. turk. 263:—Ic:.: Pall. Astr. tab. 6. Perennial, (20) 45-65 cm tall, from a strong short buried woody divided caudex; stems (10) 19-37 cm long, simple, firm, terete, clothed at base in scalelike hyaline stipules; foliar stipules not adnate to petiole, connate at base, lunate, acutish, (4) 6-12 mm long, thickish; leaves (6) 10-15 (17) cm long, the petioles firm, rigid, terete, much shorter than the rachis; leaflets 3-4 (7) pairs, orbicular to rounded-oval, rarely oval, short- acuminate, 5-11 mm long, thickish; peduncles equaling the leaves, rarely half as long, 7-14cm long; racemes (8) 15-20 cm long, very remotely (8) 14-20 flowered; bracts lanceolate, acute, 3-5mm long, hyaline-margined; pedicels 1.5-2.5 mm long; calyx 10-12 mm long, the teeth lance-subulate to subulate, 1-2 mm long, sparsely and finely fringed with hairs on the margin; standard 21-28 mm long, the limb ovate,-short-acuminate, obtusish and retuse, abruptly contracted into and longer than the claw; wing-petals 19-25 mm. long, the limb oblong, round-tipped and scarcely retuse, the claw 1.5 times as long as the limb; keel 17-23 mm long, the limb slightly convex beneath, very slightly concave above, round-tipped, half as long as the claw; ovary on a stipe 1-2 mm long, glabrous; pods 10-13 mm long, gently arcuate, divergent, 4-5 mm broad, 3-4 mm thick, pointed and terminating in a straight attenuate rigid beak ca. 2 mm long, few-seeded. Fl. May-June; fr. (May) June-July. (Plate XXXVII, Figure 3). Sands, sandstone outcrops, or rarely clayey slopes, often over gypseous substrates. — European part: L.V.; Centr. Asia: Ar.-Casp. Endemic. Described from Arsagar mountains (between Volga and Ural). Type in Leningrad. 410 {| j Section 80. AMMOTROPHUS Bge. Astrag. geront. I (1868). — Herbaceous perennials with a rather long stem, vested with bifurcate white hairs; stipules connate to rather high up, adnate to petiole only at base; leaves imparipinnate, with marcescent rather long-persistent, petioles; racemes capitate, borne on axillary peduncles; bracteoles none; calyx cylindric, not becoming inflated or scarcely so, always ruptured by the pod; corolla purple; pods sessile or subsessile, oblong, ellipsoid or ovoid- oblong, 7-12 mm long, coriaceous, more or less carinate ventrally, flattened or slightly canaliculate dorsally, bilocular, few-seeded. — A section endemic for Central Asia, distributed in Central Asian plains. 1. Leaflets 6-11 (14) pairs, glabrous or covered with isolated hairs above; standard 18-21 mm long, the limb ovate. ... 521. A. albicans Bongd. + Leaflets 4-6 pairs, sparsely covered above with appressed hairs; standard 15 mm long, the limb oval..... 022. A. ammotrophus Bge. 521. A. albicans Bongd. in Mém. Acad. Sci. Pétersb. Ser. 6, IV (1845) 177, tab. 2; Ldb. Fl. Ross. I (1842) 610; Bge. Astrag. geront. II, 192; Bge. Astrag. turk. 270. Perennial, 18-30 (40) cm tall; stems several, 10-20 (32) cm long, more or less flexuous, terete, densely appressed-tomentose, white; stipules connate to the middle and adnate to petiole at base, membranous, slightly hairy at apex, 3-5 mm long, the free portion triangular or triangular - lanceolate, acute; leaves 6-7 (11) cm long, the petiole obsolete, rachises canescent appressed-tomentose; leaflets 6-11 (14) pairs, obovate-orbicular, round -tipped, or slightly retuse (rarely subelliptic, acute), broadly cuneate at base, (3) 6-12 (15) mm long, sparsely covered beneath with short appressed hairs; peduncles 1.5-2 (3) mm long, finely sulcate, rather densely canescent-tomentose with appressed hairs; racemes 2-2.5 cm long (the rachis 0.5-1 cm long), rather loosely 6-10-flowered; bracts lanceolate, 2-2.5 mm long, covered with white hairs; pedicels ca. 1 mm long; calyx 9-10 mm long, densely tomentose with white subappressed hairs, the teeth narrowly triangular-lanceolate, ca. 1.5 mm long; corolla purple; wing- petals 16-18 mm long, the limb linear-oblong, half as long as the claw; keel 14-16 mm long, acutish; ovary on a stipe 0.5-1 mm long, hairy; pods sessile, ascending, oblong, 10-13 mm long, terminating in a mucronate beak ca. 1 mm long, inflated, slightly carinate or thickened ventrally, slightly canaliculate dorsally, rigidly coriaceous, densely villous with spreading long stiff hairs. Fl. June-July; fr. (June) July-August. Sands and pebbles. — Centr. Asia: Balkh. (vicinity of Lake Balkhash, Lake Alakul' and Arkaul mountains). Endemic. Described from Mt. Arkaul in Zaisan basin. Type in Leningrad. 522. A. ammotrophus Bge. Astrag. geront. II (1869) 192; Astrag. turk. 270. Perennial, (8) 10-25 cm tall, from a strong buried short woody caudex, the numerous divisions forming small loose mats; stems 4-20 cm long, densely appressed-tomentose, white; stipules triangular, acuminate, 3-4 mm long, scarious, sparsely white-hairy; leaves 4-8 cm long, the rachis 2-4 times as long as the petiole, both petiole and rachis densely canescent tomentose with appressed hairs; leaflets 4-6 pairs, orbicular (often broader than long), round-tipped or retuse, rounded or broadly 411 cuneate at base, 4-11 (16) mm long, thickish, canescent, covered sparsely above and densely beneath with appressed hairs; peduncles 2-4 cm long, firm, densely canescent by appressed hairs; racemes rather compact, ovoid or rounded-ovoid, (2) 3 cm long, (10) 12-17-flowered; bracts lanceolate to oblong-lanceolate, 1.5-2 mm long, covered with white hairs; pedicels ca. 1 mm long; calyx 9-10 mm long, densely shaggy with appressed white hairs, the linear -lanceolate teeth 1.5 mm long; corolla purple; standard-limb about twice as long as the claw; wing-petals 14 mm long, the limb linear-oblong, somewhat shorter than the claw; keel 13 mm long, obtusish; ovary subsessile, hairy; pods oblong, subsessile (borne on a stipe ca. 0.5 mm long), carinate ventrally, flattened or faintly canaliculate dorsally, more or less pointed, terminating in a reflexed beak 2-3 mm long, rigidly coriaceous, densely lanate-villous with white hairs. Fl. April; fr. (April) May. Gravelly and stony slopes, outcrops of mottled strata, or rarely clay- and-pebble soils. — Centr. Asia: Kyz K. (residual mountains and outcrops of bedrock in S.E. Kyzyl-kum, Nuratin mountains). Endemic. Described from Kyzyl-kum. Type in Leningrad. Section 81. PSEUDAMMOTROPHYS Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946) 135.— Perennial, with well developed stems, vested with white and black bifurcate hairs; stipules connate and not adnate to petiole, small; leaves imparipinnate; inflorescence umbellate or subumbellate, borne on a long axillary peduncle; bracteoles none; calyx 952 cylindric, at length scarcely inflated, ruptured by the pod; corolla violet; wing ~petals bilobate at apex; pods subsessile, oblong, compressed laterally, carinate ventrally, not grooved dorsally, coriaceous, bilocular or subbilocular, lanate with spreading hairs.— Pamir-Alai.— Related to the section Hypsophilus Bge. of the subgenus Calycocystis indigenous in the interior of Asia and in Central Asia. 1. Leaflets elliptic -oblong to oblong, (7) 9-12 mm long, (3) 3.5-5 mm broads limb of standard oblong’-obovate. .. 5... 6 6 5 9 +. 0 ean ee ce Cah 5 SON NB so I SRS sie til let i 523. A. cernuiflorus Gontsch. + Leaflets oval, elliptic, rounded-elliptic or suborbicular, 2-4 (7) mm long, 1.2-3 (4) mm broad; limb of standard oblong or oblong-elliptic. MMM eee Gent ge Us eee scat er st ae a a ae eet on cae. s 524. A. saratagius Bge. 523. A. cernuiflorus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, ca. 30 cm tall, from avery short buried caudex; stems 15-20 cm long, thinly covered with appressed white hairs; greenish; stipules connate up to 1/3-1/2, 4-5 mm long, covered with appressed white hairs, the free portion lanceolate, acuminate; leaves 5-7 cm long, the petiole and the much longer rachis thinly covered with appressed white hairs; leaflets 7-8 pairs, elliptic-oblong to oblong, (7) 9-12 cm long, (3) 3.5-5 mm broad, thinly covered on both sides with appressed hairs, round- tipped, broadly cuneate at base; peduncles 1.5 times as long as the leaves, thinly covered with appressed white and beneath the inflorescence with white and black hairs; racemes short, umbelliform, 7-10-flowered; flowers approximate, nodding, short-pediceled; pedicels covered with spreading 412 black hairs, ca. 1 mm long; bracts linear, 3-4 mm long, covered with spreading black hairs; calyx cylindric, at length scarcely inflated, oblong, ruptured by the pod, 13-14 mm long, densely covered with short white and black spreading hairs, the tube 3-4 times as long as the linear-subulate teeth; corolla violet; standard 23-25 mm long, the limb oblong -obovate, attenuate in lower part, angular at base, broadly short-attenuate at apex, retuse, much longer than the claw; wing-petals 18-19 mm long, the limb linear -oblong, bifid at apex, somewhat shorter than to as long as the claw; keel 15-16 mm long, the limb rounded-gibbous, obtusish, about half as long as the claw; ovary on a stipe ca. 1 mm long; pods subsessile, oblong, ca. 10mm long, 4mm broad, compressed laterally, carinate ventrally, convex to subcarinate dorsally, terminating in a beak ca. 3 mm long, coriaceous, densely covered with spreading white and, on the ventral side, mixed white and black hairs, bilocular. Fl. August-September; fr. September. Dry natural sedge meadows in the upper part of the juniper zone, at altitudes of ca. 2900m.—Centr. Asia: Pam.-Al. (Karavshin river basin, 553 northern slopes of the Turkestan range). Endemic. Described from Dzhaupaya river valley. Type in Leningrad. 524. A. saratagius Bge. Astrag. turk. (1880) 269; Fl. Tadzhik. V, A270 etc > El. dadzhik. V, Plate XLV III. Perennial, (9) 15-30 (36) cm tall, densely canescent with appressed white hairs; floriferous stems (7.5) 15-20 (27)cm long, bearing 1-3 flowers; stipules connate up to 1/3- 2/3, to 3 mm long, the free portion short - triangular to acuminate-linear; leaves 1.5-4.5 cm long, short -petioled; leaflets 6-9 (10) pairs, oval, elliptic, or suborbicular, 2-4 (7) mm long, 1.2-3 (4) mm broad, subobtuse or round-tipped, rather densely covered on both sides with appressed hairs; peduncles (1.2) 2.4-6 (9) cm long; inflorescence capitate, obovoid, 5-15 (22)-flowered; bracts linear -subulate to narrowly linear, 3-5 mm long, rather densely covered with black or mixed black and white hairs; calyx cylindric, at length scarcely inflated, ruptured by the pod, 10-14 mm long, covered with appressed or rarely slightly spreading white hairs, these interspersed, especially on the teeth, with shorter black hairs; the teeth subulate, 2-3 (5) mm long; corolla violet or pale violet, with a dark violet speckle on the keel; standard 19-24 mm long, the limb oblong to oblong-elliptic, 15-17 mm long, retuse, much longer than the claw; wing-petals 15-20 mm long, the limb oblong, bifid at apex, 7-9 mm long; keel 12.5-16 mm long, the limb obtuse, 4.5-6 mm long; ovary on a stipe up to 1 mm long; pods subsessile, inequilaterally oblong or oblong-elliptic, 7-10 (11) mm long, 2.5-3.5mm broad, more or less compressed laterally, slightly carinate ventrally, convex dorsally, terminating in a short upturned beak 1.5-2 mm long, coriaceous, covered with white and, chiefly on the ventral side, with mixed white and black hairs. Fl. May-July; fr. June-August. Stony slopes and taluses in the steppe and juniper zones and in the lower part of the subalpine zone, at altitudes of 2000-3000 m.— Centr. Asia: Pam.-Al. (Zeravshan mountains from Dukdon Pass and Saratag River to upper reaches of the Yagnob; northern slopes of the western part of Turkestan range). Endemic. Described from Saratag river valley. Type in Leningrad. 413 554 557 Section 82. LEUCOPHYSA Bge. Astrag. geront. I (1868) 12.— Herbaceous perennials, from a woody buried caudex, vested with bifurcate hairs; stems of the year developed; stipules free (not connate), partly adnate at base to petiole; leaves imparipinnate; inflorescence compactly capitate, few-flowered, borne on an axillary peduncle; bracteoles none; calyx cylindric; corolla pinkish-violet, glabrous; limb of standard angularly attenuate in lower part; pods sessile, bladdery-inflated, membranous, bilocular. Irtysh basin. Psammophytes of Zaisan depression and of Chernyi Irtysh river valley near the border of W. Mongolia. 1. Leaflets densely and finely pannose-lanuginous on both sides with ascending hairs; peduncles somewhat shorter than to equaling the leaves; bracts lanceolate; pods spherical, acuminate to a curved beak 4-7 mm long, densely pannose-lanuginous with spreading white hairs. dora We Kaui yate) Seah shes Wedeeese SE oh ol ee ators SY aeragy Neds lial 925. A. candidissimus Ldb. + Leaflets covered on both sides (sometimes thinly so above) with stiff appressed hairs; peduncles much shorter than the leaves; bracts linear to lance-linear; pods spherical, round-tipped, beaked by a short mucro 1-2 mm long; densely sublanuginous to hispid-villous and COMACTILO SC Hep sm civs FA AYERS St mani Rh ear aeee ede a 526. A. Steinbergianus Sumn. 525. A. candidissimus Ldb. Fl. Alt. III (1831) 309; Ldb. Fl. Ross.I, 623; Beet Astrag. seront. 11,193; ‘Kryl. Plt Zap. sib. Villy Hoos neem yar. pauciflorus ‘Kryl. "et Serg.— Ic.: Ldb. Ic. Fl Ross. ii (lao) tab. 237. Perennial, 15-26 cm tall, from a short buried woody caudex, with woody divisions, 4-10 cm long; stems erect or assurgent, 12-20 cm long, terete, white-pannose; stipules triangular, acuminate, 4-6 mm long, white-pannose; leaves (2.5) 4-8 cm long, the rachis 2-4 times as long as the petiole, both petiole and rachis white-pannose; leaflets 7-12 approximate pairs, broadly elliptic to suborbicular, rarely broadly obovate, subacute or mucronulate, 3-8 (13) mm long, light gray; peduncles firm, lanate with short hairs, white, sometimes with black hairs at the end; inflorescence 915-flowered, 2-3cm long, spherical; flowers subsessile; bracts 4-5 mm long, acuminate, white woolly-shaggy; calyx densely pannose with spreading white hairs, sometimes interspersed with scattered black hairs, the linear-subulate teeth 3-4 mm long; corolla pinkish-violet; standard 16-17 mm long, the limb obovate, retuse, 2-3 times as long as the claw; wing-petals 14-16.mm long, the limb oblong, round-tipped, somewhat longer than to 1.5 times as long as the claw; keel 13-14 mm long, the limb acutish, 2/3 as long as the claw; pods 12-17 mm long, deeply grooved ventrally, canaliculate dorsally; seeds angular, ca. 3mm long, scarcely pitted, yellowish-brown. Fl. June; fr. June-August. Sandhills and slightly fixed sand dunes.— Centr. Asia: Balkh. (Zaisan depression). Endemic. Described from Irtysh river valley (between Kurchum and Naryn rivers).— Type in Leningrad. 526. A. Steinbergianus Sumn. in Animadvers. Herb. Univ. Tomsk. Nos. 2-3 (1934) 3:— A. candidissimujs ;van., page 1 fl 0 uses ler Sero.siny Hl. Zap. Sib.) Vill (1933) 1634" Perennial, (8) 12-23 cm tall; stems erect or assurgent, rarely subdecumbent, (2) 4-8 (10) cm long, lanate with short white hairs; stipules 414 NN TX BN ‘ SIN \ QQ. Ne Wy Poses a Me PLATE XXXVI Boiss Lipsky.—4. A. isfahanicus 415 558 lanceolate, acuminate, 5-7 mm long, lanate with short subappressed white hairs; leaves (5) 8-18 cm long, the petiole and the much longer rachis densely strigulose with subappressed hairs; leaflets (8) 10-14 pairs, elliptic, acute, rarely broadly oval and round-tipped, (5) 7-12 (15) mm long; peduncles (2) 3-7 cm long, densely strigose with short subappressed hairs, hoary; inflorescence compactly capitate, (3) 5-8-flowered, ca. 2 cm long; bracts mostly finely pointed, (3) 4-7 mm long, covered with white hairs; pedicels ca. 1 mm long; calyx 11-12 mm long, finely lanate with subappressed white or black hairs, the linear-subulate teeth 1.2-2 mm long; corolla pinkish-violet; standard 18-19 mm long, the limb obovate, slightly retuse or subentire, 2-3 times as long as the claw; wing-petals 17-18 mm long, the limb oblong, round-tipped, somewhat shorter than the claw; keel 15 mm long, the limb obtusish, the claw 1.5-2 times as long as the limb; pods 13-15 mm long, grooved ventrally, canaliculate dorsally. Fl. June- July; Fr. July-August. Sand dunes, rarely sand and pebble deposits.— Centr. Asia: Balkh. (Zaisan depression). Gen. distr.: W. Mongolia, in Chernyi Irtysh river basin. Described from the vicinity of Buran, near the estuary of Alkabek River. Type in Leningrad. Section 83. CYSTIUM Bge. Astrag. geront. I (1869) 113 (pro maxima parte). — Nearly acaulescent perennials, vested with bifurcate hairs; stipules more or less connate, sometimes only at base; leaves imparipinnate, the petioles marcescent; racemes contracted, borne on a long peduncle; bracteoles none; calyx cylindric; corolla more or less violet-colored; pods sessile or nearly so, biovaloid or spherical-bladdery, grooved dorsally and ventrally, membranous, glabrous or very finely covered with scattered appressed hairs, bilocular. The distribution area of this section comprises Crimea, the southern steppe and semidesert region from Lower Volga to Altai, and the mountains of Central Asia and North Iran. 1 Weafletsgollabrouls above ect-h- 04 tier. Geet. lala atnely-t ote a mene 2. + Leaflets covered above with appressed hairs .......2.4..-e2-2ee% Ne 2. Corolla ochroleucous, faintly suffused with violet and with a violet keel, 22.5-26 mm long; leaflets ovate or oval, glabrous except for isolated appressed hairs on the midrib beneath and on the margin ............ Py UMA Yet, .. SSeS ees, RS ie. ee ee 527. A. physocarpus Ldb.. + Corolla violet, 16-18 (20) mm long; leaflets elliptic to oblong- elliptic, acute, covered beneath with scattered appressed hairs..... TS ee ee De, Cea Seer ee 528. A. physodes L. 32 Ovary and=podsralabrous)s. 2 Ay. (yi. yoy ee cineun) - een ee 4. + Ovary pubescent; pods covered with subappressed white hairs...... SOS A UTES La. OUT Sea fsa 531. A. Skorniakovii B. Fedtsch. 4. Standard 18-20 mm long, the limb rhomboid-oblong; pods 10-15 mm Origen sabre STA YSe. Tey es he ee 529. A. suprapilosus Gontsch. + Standard (20) 21-25 mm long, the limb oblong-oval to rhomboid -oval, subligulately attenuate toward apex .... 530. A. kurdaicus Saposhn. Series 1. SUPRAGLABRI Gontsch. — Leaflets glabrous above; ovary and pod glabrous. 416 59 527. A. physocarpus Ldb. Fl. Alt. III (1831) 336; Ej. Fl. Ross. I, 658; Kryl. Fl. Zap. Sib. VII, 1684.—A. physodes altaicus Pall. Astrag. (1800) 72.— A. physodes Bge. Astrag. geront. II (1869) 196, non L. ex parte. Perennial, nearly acaulescent, (10) 15-18 cm tall; stems strongly abbreviated, 0.5-1.5 cm long, vested with white hairs, tightly clothed in stipules; stipules ovate-lanceolate, 6-10 mm long, scarious, glabrous, the lower sometimes appressed-hairy; leaves 10-15 cm long, the rechis 1.5 times as long to as long as the petiole, both petiole and rachis glabrous; leaflets 8-13 pairs, ovate or oval, (8) 10-20 mm long, 5-12 mm broad, green, practically glabrous throughout, except for isolated appressed hairs on the midrib beneath and onthe margin; peduncles somewhat exceeded by the leaves, together with inflorescence equaling or somewhat exceeding them, glabrous, sulcate, stout; racemes rather compactly capitate, ovoid, 3-5 cm long, many-flowered; bracts lanceolate, 5-7 mm long, 3-4 times as long as the pedicel, rather sparsely covered with black hairs; calyx cylindric, 10-12 mm long, covered with appressed black hairs, the tube 3-4 (5) times as long as the linear-lanceolate teeth; corolla ochroleucous, faintly suffused with violet; standard 22.5-26 mm long, the limb oblong- rhomboid, dilated at the middle, gradually and broadly ligulate -attenuate upward, more or less retuse, abruptly narrowed and angular at base, 4-5 times as long as the claw; wing-petals 17-20 mm long, the limb lance -oblong, entire, obtusish, somewhat shorter than to as long as the claw; keel 15-17 mm long; ovary on a stipe 1.5-3 mm long, glabrous; pods spherically bladdery-inflated, thinly membranous, glabrous, (15) 20-30mm in diameter, grooved ventrally and dorsally, bilocular. Fl. May; fr. June. Steppes, often in dry and more or less saline soils; sporadically in the southern steppe region, from Orenburg to South Altai. — W. Siberia: U. Mopeaisn SeALt a(S); «Centr. Asia: Ar.-Casp. (N.),, Balkh. \(N..). Endemic. Described from Altai foothills. Type in Leningrad. 528. A. physodes L. Sp. pl. (1753) 760; Pall. Astrag. 71 (excl. var.); Ldb. Fl. Ross. I, 659, ex parte; Bge. Astrag. geront. II, 194, ex parte; Schmalh. Fl. I, 282, ex parte.— Xerophysa physodes Stey. in Bull. Soc. Nat. Mose. XXIX (1856) 150.—Ic.: Pall. Astrag. tab. 98,°B, £2) 1. Perennial, nearly acaulescent (stems not more than 1 cm long), 10-17cm tall; stipules 7-9 mm long, ovate-lanceolate, scarious, glabrous, the lower covered withappressed stiff white hairs; leaves 7-15cm long, the rachis 1.5- 3 times as long asthe petiole, both petiole and rachis covered with scattered appressed white hairs; leaflets 8-12 pairs, ellipticto oblong-elliptic, acute, rarely subobtuse, 6-12 mm long, glabrous above, covered beneath with scattered appressed hairs, very rarely the hairs confined to the midnerve and the margin; peduncles somewhat exceeded by the leaves, together with inflorescence mostly about equaling or exceeding them, covered with scattered white appressed hairs; these sometimes interspersed above [?] the inflorescence with black hairs; racemes rather compact, ovoid, 3-5cm long, mostly many-flowered; bracts lanceolate, 3-4 mm long, covered with black hairs; calyx cylindric, 10 mm long, villosulous with short ascending black hairs, the narrowly linear teeth 1.5-2 mm long; corolla violet; standard 16-18 (20) mm long, the limb rhomboid-oblong, abruptly narrowed in lower one-third, oblong-ovate upward, entire, 5-6 times as long as the claw; wing-petals 15-16 mm long, the limb linear-oblong, slightly retuse, somewhat shorter than to as long as the claw; keel 13-14 mm 417 560 long, obtuse; ovary on a stipe 1.5-2 mm long, glabrous; pods sessile, spherical, 15-25 mm long, grooved ventrally and dorsally, glabrous, bilocular. Fl. April (May); fr. May. Steppes and hillsides. — European part: L.V.; W. Siberia: U. Tob. (S.W.); Centr. Asia: Ar.-Casp. (N.W.). Endemic. Described from "Siberia''. Type in London. Note. A small-flowered form, with glabrate leaves, occurring in the Ural'sk region. It resembles the Siberian A. physocarpus Lab. Series 2. SUPRAPILOSI Gontsch. — Leaflets hairy above; ovary and pods glabrous. 529. A. suprapilosus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). A. physodes var.tauricus Pall. Astrag. (1800) “Zt lee: HPallapAstrage=ntabic.oojrf.. ale Perennial, nearly acaulescent, 6-17 cm tall; stipules dimorphic, the lower ovate, the upper ovate-lanceolate, 5-7 mm long, glabrous, the lower ones often covered with white hairs; leaves 4-10 (13) cm long, the rachis 1.5-3 times as long as the petiole, both petiole and rachis covered with scattered appressed hairs; leaflets (10) 12-15 pairs, lanceolate to oblong- elliptic, acute, 5-10 mm long, covered with appressed white hairs, these scattered above and sparse beneath; peduncles somewhat shorter than the leaves, together with inflorescence equaling or somewhat exceeding them; racemes commonly ovoid, rather loose, many-flowered; bracts lanceolate, ca. 3mm long, black-ciliolate, 2-3 times as long as the pedicel; calyx narrowly cylindric, 7-8 mm long, thinly covered with short ascending black hairs, the linear -subulate teeth 1-2 mm long; corolla violet; standard 18-20 mm long, the limb rhomboid-oblong, abruptly narrowed in lower 1/3-1/4, angular at base, 5-7 times as long as the claw; wing-petals 14-15 mm long, the limb oblong, slightly retuse, as long as the claw; keel 12 mm long, obtuse; ovary on a stipe 1-1.5 mm long, glabrous; pods spherical to ovoid-spherical, 10-15 mm long, rarely ovoid, 17-25 mmlong, grooved ventrally and dorsally, glabrous, bilocular. Fl. May; fr. May- June.. Mountain slopes. — European part: Crimea. Endemic. Described from the vicinity of Feodosiya. Type in Leningrad. 530. A. kurdaicus Saposhn. ex Sumn. in Animadv. Syst. univ. Tomsk, Nos.9-10 (1936) 7.—A. pseudophysodes Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, (8) 10-20 cm tall, from a short woody buried caudex with short divisions; stems of the year obsolescent or up to 2 cm long, densely vested with subappressed stiff white hairs; stipules dimorphic, the lower ovate, the upper lanceolate, acuminate, 6-9 mm long, white -hyaline, glabrous; leaves (5) 11-18 cm long, the petiole and the much longer rachis sparsely covered with appressed white hairs; leaflets (10) 13-20 (22) pairs, lanceolate, (5) 8-14 mm long, acute, covered with appressed hairs, these sparse beneath, scattered above; peduncles somewhat shorter than the leaves, together with inflorescence equaling to somewhat exceeding them, covered with isolated appressed white hairs; racemes subovoid, rather compact, 3.5-5 cm long, many-flowered; bracts lanceolate, the lower sometimes ovate-lanceolate, 4-6 mm long, acutish, black-ciliate, 3-4times the length of the pedicel; calyx cylindric, 10-13 mm long, covered with 418 561 black subappressed hairs, these often interspersed with isolated white hairs, the tube 2.5-3 times as long as the lance-linear teeth; corolla violet; standard (20) 21-25 mm long, the limb oblong-oval to rhomboid-oval, slightly and broadly ovate-attenuate upward, retuse, abruptly narrowed in lower 1/4-1/3, angular at base, 4-6 times as long as the claw; wing-petals 17-21 mm long, limb oblong, somewhat emarginate, equal to nearly as long as the claw; keel 16-19 mm long, obtuse; ovary ona stipe 1-1.5 mm long, glabrous; pods ovoid or spherical, sessile, 16-30 mm long, grooved dorsally and ventrally, glabrous, bilocular. Fl. April (May); fr. May-June. Stony and gravelly slopes of mountains and foothills, in the upper part of the mid-mountain zone and the peripheral zone of the Central Asian mountains. — Centr. Asia: Pam.-Al. (W.), T. Sh., Mtn. Turkm. (Kopet Dagh). Endemic. Described from Kirgiz (Kurdai River). Type in Leningrad. Series 3. PILOSISSIMI Gontsch.— Leaflets hairy above; ovary and pods hairy. 531. A. Skorniakovii B. Fedtsch. in A.H.P. XXIV (1904) 227.— A. physodes B. Fedtsch. in O. et B. Fedtsch. Fl. Pamir. (1903) 459, nonL.—A. Danieli Kochii Freyn in Bull. Herb. Boiss. V, 2 (1905) 558. Perennial, nearly acaulescent, 10-30 cm tall, loosely cespitose; stems of the year 1-2.5 (3.5) cm long, densely covered with white hairs; stipules connate to 1/3-1/2, 6-12 mm long, hyaline, the lower oblong triangular - ovate, the upper triangular-lanceolate, covered with white hairs; leaves 6-11 (18) cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis rather sparsely covered with appressed white hairs; leaflets 11-23 pairs, lanceolate to oblong-elliptic, acute, 6-10 mm long, rather sparsely covered on both sides with appressed hairs; peduncles 4-10 (15) cm long, sparsely covered with appressed white andinthe upper part sometimes mixed white and black hairs; racemes rather compact, 2-4 cm long, elongating in fruit up to 10-13 cm; bracts linear-lanceolate, 5-7 mm long, thinly covered with white and black hairs, 3-4 times the length of the pedicel: calyx cylindric, 10-13 mm long, covered with appressed black and white hairs, the tube 3-4 times as long as the linear teeth; corolla violet; standard 20-25 mm long, the limb elliptic, obtuse, 16-21 mm long; wing-petals 18-22 mm long, the limb oblong, 9-11 mm long; keel14-18mm long; ovary hairy; pods sessile, bladdery-inflated, ovaloid or spherical, 16-30 mm long, thinly membranous, covered with scattered subappressed short white hairs, bilocular, many-seeded. Fl. May-June; fr. June-July. (Plate XXXVII, Figure 2). Mountain slopes, often in gravelly soil; mountaindeserts and semideserts, in scrub of the mid-mountain zone. Centr. Asia: Pam.-Al., T. Sh., Mtn. Turkm. Endemic. Described from Arasan. Type in Leningrad. Section 84. PARACYSTIUM Gontsch., sect. nova in Addenda XI, p.661.—Nearly acaulescent or short-stemmed low perennials, vested with bifurcate hairs; stipules partially adnate to petiole, not connate; racemes loosely few-flowered; peduncles short, exceeded by or rarely longer than the leaves; bracteoles none; calyx cylindric; corolla commonly violet, glabrous; pods bladdery-inflated, rigidly membranous or coriaceous, sessile or short-stipitate, more or less pointed, grooved or rarely flattened 419 563 dorsally, villous with spreading hairs or pilosulous or glabrous, bilocular.— An endemic Turanian section, distributed in the deserts of Central Asia and the foothills of E. Transcaucasia and Tien Shan (Karatau Mountains). 1. Leaflets covered above with appressed hairs; pods appressed-pillose. 58, eer ee Ae) aiaclgaeime tatrerras, omnoktodeth bce eee 2. ‘= leeatlets labrousraboviecien. -)eewictencetial tetietoullts ue f. g-Ueiae) enna 3. 2. Leaflets 7-10 pairs, narrowly linear to subfiliform, subacute...... Va. 82 ive Jingk o.oo = ee 532. A. pachyrhizus M. Pop. + Leaflets 4-7 pairs, oblong to oblong-lanceolate, obtuse .......... MarR 6 SE) Ol B.S R. LRA Ss ate 5935. A. Kikodzeanus Sosn. gu Ovaryrandspod) collab OUus) 9A: aterieds 3-8 ices Menke 536. A. tetrastichus Bge. + Ovary and: pod hairy”. <4 tau.) SAGkr ay Ges CURR | seamen ae 4. ake ON BISY BD-SOHCnAMael ohn co lb oo bo dO Oo c 034. A. Biebersteinii Bge. ie Ovatay “10-6 0l-Ovul edi ei ee ene on) atten bet 933. A. lasiophyllus Ldb. Series 1. CYSTIMORPHI Gontsch.— Pods appressed -pilose. 532. A. pachyrhizus M.Pop. in Bull. Univ. As. Centr., fase. 14 (1926) 139. Perennial, 8-9 cm tall, from a strong woody root and a strongly abbreviated woody buried caudex, the numerous short divisions forming large loose mats; stems of the year (2) 4-7 cm long, simple, densely appressed-puberulent, white; stipules adnate to petiole in lower 1/3, free, ca. 2 mm long, thinly membranous, whitish, glabrous or nearly so, the free portion triangular, acute; leaves 4-6 cm long, the rachis 2-3 times as long as the petiole, the petiole very slender, whitish, densely covered with short appressed hairs; leaflets 7-10 pairs, narrowly linear, rarely linear-oblong or sometimes subfiliform, subacute, (2) 5-10 mm long, 0:5-1 mm broad, rather densely covered on both sides with appressed hairs, hoary, mostly conduplicate; peduncles mostly exceeded by the leaves, together with inflorescence shorter than to equaling the leaves, 2-4cm long, covered with short appressed white hairs; racemes subspherical, 2-2.5 cm long, loosely 3-5-flowered; bracts ovate, acutish, 1-1.5 mm long, white- hyaline, thinly covered with black and white hairs; pedicels 1.5-2 mm long, covered with appressed black hairs; calyx cylindric, 9-11 mm long, densely covered with appressed short black hairs, the tube 4 times as long as the narrowly linear teeth; corolla purple; standard 19-20 mm long, the limb oblong-ovate, attenuate at the tip and retuse, 1.5 times as long as the claw; wing-petals 16-17 mm long, the limb oblong, round-tipped, as long as the claw; keel 13-14 mm long, the limb straight above, convex beneath, rounded-obtuse, 2/3 as long as the claw; ovary subsessile; pods sessile, bladdery-inflated, ovoid, 2-4 cm long, 1-2 cm broad, acute, membranous, sparsely or rather densely appressed-villulose with white hairs (at first canescent). Fl. April-June; fr. May-July. Clay-and-gravel and stony mountain slopes.— Centr. Asia: Tien Shan (Karatau and Talass Ala Tau mountains). Endemic. Described from Karatau and Talass Ala Tau. Type in Tashkent. Series 2. TURANICI Gontsch. — Pods glabrous or villous with spreading hairs. SSipmllsly7, 420 964 533. A. lasiophyllus Ldb. Fl. Ross. I (1842) 627.—A. Pallasii Fisch. Cat. Hort. Gorenk. (1812) 71; non Spreng. (1807); DC. Prodr. II, 305, Ldb, Fl. Alt. III, 336; Ldb. Fl. Ross. I, 659; Bge. Astrag. geront. Pl ale Loa Bese. Astrag. turks 2785) Keyl. Pl. ’Zap. Sibe) Vil.) 16853— A. inderiensis Claus, Beitr. Pfl. Russ. Reich. VIII (1851) 64. Perennial, hoary, dwarf, acaulescent or short-stemmed, 5-10 cm tall, from a short buried woody caudex, the short divisions forming rather small mats; stems of the year scarcely developed, up to 5 cm or rarely up to 10-15 cm long, densely villous with appressed hairs or lanate with ascending hairs; stipules adnate to petiole up to the middle, not connate, ovate, the upper ovate-lanceolate, covered with white spreading or rarely appressed hairs; leaves 2.5-8 cm long, the rachis 2-3 times as long as rarely shorter or as long as the petiole, the petiole covered with short white appressed or ascending hairs; leaflets 4-7 pairs, linear-oblong to oblong- oval rarely oblong-obovate, (3) 4-8 (10) mm long, the upper face glabrous or covered especially on the margins with scattered hairs, the lower face densely covered with appressed or spreading hairs; peduncles half as long to as long as the leaves, densely covered with appressed or ascending white hairs; racemes short, 2-5-flowered; bracts ovate-lanceolate or ovate, ca. 1.5 mm long, half the length of the pedicel; calyx cylindric, 13-16 mm long, covered with black and white, more or less spreading hairs, the tube 5-6 times as long as the lanceolate teeth; corolla pale lilac; standard 23-28 mm long, the limb oblong-obovate, ovate in upper part, somewhat drawn out toward apex, abruptly narrowed in the lower 1/3-1/2, auriculately angular at base, 3-4 times as long as the claw; wing-petals 22-27 mm long, the limb oblong, inequilaterally emarginate, about half as long as the claw; keel 19-25 mm long, the limb broad, slightly curved, obtuse, the claw 2.5-3 times as long as the limb; ovary on a stipe 1-1.5 mm long, tomentose; pods subsessile (borne on a very short stipe), bladdery- inflated, broadly ovate to rounded-ovate, (15) 18-24 mm long, sparsely covered with spreading white hairs, rigidly membranous, short~acuminate, bilocular, Fl. (April) May; fr. May-June. (Plate XXXVII, Figure 1). In wormwood and saltwort-and-wormwood associations; rarely on sandhills in river valleys of the sandy ''steppe'’, on chalk and hard clay outcrops.— Centr. Asia: Ar.-Casp., Balkh., Syr D. (N.). Gen. distr.: Kulja, N.W. Mongolia. Described from Inders Lake. Type in Leningrad. 534. A. Biebersteinii Bge. Astrag. geront. II (1869) 194; Grossh. RUN davies Tyr s Sil}: Perennial, closely resembling the preceding species; distinguished by the smaller number of ovules. Dry slopes in the foothills and in the lower mountain zone. — Caucasus: E. Transc. (Kurt-bulak, Altyagach). Endemic. Described from Kurt -bulak and Altyagach. Type in Leningrad. 535. A. Kikodzeanus Sosn. in Trav. Inst. Bot. Tphilis. II (1937) 227. — A. Biebersteinii Boiss. Fl. or. II (1872) 453, non Bge. — A. physodes M.B. Fl. taur.-cauc. II (1808) 198, non L. Perennial, nearly acaulescent, peppery-gray by appressed pubescence; root -crown branched into many divisions; stipules linear -subulate, membranous at base, herbaceous-tipped, rather long-ciliate, almost free; leaflets 4-7 pairs, oblong to oblong-lanceolate, obtuse, covered on both sides notably beneath with nearly straight hairs; peduncles sessile; 421 racemes capitate, 3-6-flowered; bracts small, linear-lanceolate, one-third as long as the calyx, white-hairy on the margin, exceeding the pedicel; calyx covered with short appressed white hairs, these not interspersed with black, the tube 4 times as long as the subulate teeth; standard about 2.5 times the length of the calyx, the limb oblong, suberoded; ovary appressed- hairy; pod unknown. Fl. May. Dry hills. — Caucasus: E. Transc. (Georgia). Endemic. Described from Goriis district. Type in Tbilisi. 536. A. tetrastichus Bge. Astrag. turk. (1880) 279. Perennial, nearly acaulescent, 8-13 cm tall, from short buried woody caudices, cespitose; stems of the year scarcely developed, not more than 0.5 cm long, densely vested with spreading white hairs, tightly clothed in petioles and stipules; stipules adnate to petiole up to one-third, densely white-lanate, 4-5 mm long, the lower triangular, the upper lance-oblong; leaves 5-10 cm long, the petiole and the longer rachis sparsely covered with spreading hairs; leaflets 5-9 pairs, oblong-oval, round-tipped, (5) 6-9 mm long, glabrous above, densely shaggy beneath with short hairs; peduncles exceeded by the leaves, 1.6-7 cm long, sparsely villosulous with spreading hairs; racemes very short, 2-5 fruited; bracts triangular -ovoid, ca. 2mm long, sparsely villosulous with short spreading white hairs; pedicels ca. 4 mm long, beset with white and black hairs; calyx cylindric, 10-13 mm long, covered with ascending black and white hairs, the linear - subulate teeth 1.5-2 mm long; standard 20-25 mm long, the limb oblong, retuse; ovary glabrous; pods borne on an obsolescent stipe 2-3 mm long, bladdery-inflated, ovoid to oblong-ovoid, 14-20 mm long, pointed, terminating in a beak 1.5-2 mm long, slightly canaliculate dorsally and 565 ventrally, thinly coriaceous, glabrous, densely and alveolately reticulate - nervose, bilocular. Fr. May-June. ' Sands and more or less sandy slopes of low mountains. — Centr. Asia: Syr D. (Syr Dar river valley, at Chiili Station, in Dzhankil', eastern outskirts of Kyzyl Kum, Karakuz), Pam.- Al. (Sarytau mountains). Endemic. Described from Karakuz. Type in Leningrad. Section 85. POPOVIANTHE Gontsch. sect. novain Addenda XI, p. 661. Acaulescent plants, forming small mats, covered with bicuspidate hairs, from a short buried caudex with short divisions; stipules adnate to petiole; flowers in 2's or 3's on peduncles, these 1/3-1/2 as long as the leaves; bracteoles none; calyx cylindric; corolla purplish-violet, large; pods subsessile, oblong, carinate ventrally, complanate dorsally, rigidly membranous. — Perennial Central Asian section. 537. A. Popovii Pavl. in Bull. Soc. Nat. Mosc. sér. Biol. XLIV, Nos. 1-2 (1935) 36. Perennial dwarf plants, 3-6 cm tall, from a strong very short woody buried caudex, the divisions forming fairly compact mats; stipules adnate to petiole up to 1/3-1/2, 3-5 mm long, the free portion oblong or lanceolate, acute, thinly membranous, white-hairy; leaves (1.5) 2-4 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis slender, rather densely covered with short spreading white hairs; leaflets (5) 7-10 pairs, lanceolate or elliptic, conduplicate, (2) 3-6 mmlong, subacute, densely covered on both sides with spreading hairs; peduncles 1/3 -1/2 the length of the leaves; 0.5-1 (1.5) cm long, rather densely tomentose with 422 66 short spreading hairs, 2-3-flowered, the fruiting peduncles somewhat shorter thanto equaling the leaves; bracts lanceolate, acute, ca. 2 mm long, white- hairy, faintly white- and black-hairy; pedicels ca. 2 mm long, beset with black and white hairs; corolla purplish-violet; standard 27-30 cm long, the limb oblong-obovoid, round-tipped, retuse, abruptly narrowed in lower 1/3, angular at base, 2-3 times as long as the claw; wing-petals 23-27 mm long, the limb oblong to linear-oblong, slightly retuse, barely shorter than the claw; keel 25-28 mm long, somewhat exceeding the wings, the limb oblong or obovate, rounded-obtuse, somewhat shorter than the claw; ovary ona stipe 1.5-2 mm long, shaggy; pods subsessile (the stipe obsolescent), oblong, 18-21 mm long, short-acuminate, carinate ventrally, complanate dorsally, strongly inflated, subtriquetrous, coriaceous, densely villous with spreading soft white hairs, unilocular. Fl. (May) June-July; fr. (June) July-September. Stony or gravelly mountain slopes, at altitudes from 1750 m (Karatau) to 3200 m (Alai Valley).— Centr. Asia: T. Sh. (Karatau Mountains), Pam.-Al. (Alai Valley). Endemic. Described from the vicinity of Mindzhelke in the Karatau Mountains. Type in Moscow. Note. The occurrence of this species in Alai Valley is very doubtful. It is possible that the indication of the site by the author was due to an error in labelling, the more so considering that all the labels of the collections of S. Yu. Lipshits which provided material for the description of the species from Karatau Mountains and from Alai Valley, refertothe same year of 1930. Section 86. TROPIDOLOBUS Gontsch. sect. n.in Addenda XI, 881.— Nearly acaulescent perennials, often lignescent at base, vested with white and black bicuspidate hairs; stipules free (not connate), more or less adnate to petiole; leaves imparipinnate; racemes short, compact, borne on rather long peduncles; bracteoles none; calyx cylindric; corolla glabrous; pods sessile, ovoid to oblong-ovoid, rupturing the calyx, carinate by the valves, 3- or 4-angled, rigidly subligneous ~coriaceous, villous with spreading hairs, bilocular. An endemic Central Asian section, occurring in the eastern part of West Tien Shan and in northern Pamir- Alai. 1. Leaflets ovate or elliptic or oblong-elliptic, 18-34 mm long; peduncles 10-17 cm long; pods 20-22 mm long, 4-angled (Fergana Range)...... Le MSMR Rs kote hav ileihol Seta Ae Ack Map st 6 he ay oe ve 538. A. irisuensis Boriss. + Leaflets oblong-elliptic or lanceolate, 7-15 (17) mm long; pods 12-15 mm long, 3-angled or irregularly 4-~angled ...........-.. 2 2. Pods 3-angled, complanate dorsally; peduncles ca. 8 cm long, half as long as the leaves (Alai range, Shakhimardan river basin)....... 50 oth OE ROR AU GIRL BGs Cid c oie ats 2 9A) ge tees 540. A. Borissianus Gontsch. * + Pods irregularly 4-angled, angled-carinate dorsally; peduncles 3-4 cm long, much shorter than the leaves (Alai range, Sfawiam) oss ce cen 5 -0..08 Oh -ogRe we Senet ar le tar an ct 539. A. isphairamicus B. Fedtsch. 538. A. irisuensis Boriss. nom. nov.— A. sclerocarpus M. Pop. in Not. Syst. Herb. Inst. Bot. Ac. Sc. URSS VII (1938) 112, non A. Gray. Nearly acaulescent perennials, up to 35 cm tall; stems of the year 3-6 cm long, often obsolescent, densely vested with short stiff ascending * See also A. distentus Boriss., p. 654, 423 567 968 hairs; stipules shortly adnate to petiole, lance-linear, 5-9 mm long, covered with white hairs; leaves 15-30 cm long, the rachis 4-5 times as long as the petiole, both petiole and rachis angled, sparsely covered with appressed white hairs, indurescent and partially persistent; leaflets 7-12 pairs, ovate or elliptic to oblong-elliptic, short-cuspidate, 18-34 mm long, sparsely covered on both sides with appressed white hairs, green; peduncles half as long as to somewhat shorter than the leaves, angled, firm, 10-17cm long, appressed -pilosulous with white hairs; racemes crowded, becoming rather loose in fruit, 10-20-flowered; bracts linear -lanceolate to lanceolate, 3-5 mm long, finely covered with black hairs; calyx cylindric, about 12mm long, covered with white and black appressed hairs, the teeth linear-subulate, ca. 2mm long; corolla violet; standard 23 mm long; keel 18 mm long; pods sessile, oblong-ovoid, 20-22 mm long, carinate by the sutures and the valves, tetrangular, terminating in a curved beak 3-4 mm long, very rigid, ligneous~coriaceous, hispid-villous with spreading hairs, bilocular. Fr. August. Gravelly mountain slopes.— Centr. Asia: Tien Shan (southwestern slopes of Fergana Range: Iri-su, between Charvak and Arslanbob, between Malyi Urtak and Kyzyl-ungur river basin). Endemic. Described from Iri-su river valley. Type in Tashkent. 539. A. isphairamicus B. Fedtsch. in Not. Syst. Herb. Inst. Bot. Ac. Sc. URSS VIII (1940) 168. Perennial, nearly acaulescent, 20-24 cm tall, lignescent at base; stems of the year 2-3 cm long, tightly clothed in stipules and leaf petioles; stipules adnate to petiole up to the middle, 7-8 mm long, covered with appressed white hairs, the free portion linear-lanceolate, the upper stipules linear; leaves 10-20 cm long, the petiole and the longer rachis densely appressed- tomentose; leaflets 7-12 pairs, oblong-elliptic to lanceolate, acute, 7-15mm long, appressed-hairy on both sides; peduncles 3-4 cm long, covered with appressed white hairs; racemes congested, short, 7-9-flowered; bracts lanceolate, 2-2.5 mm long, black-tomentose; calyx cylindric, 11 mm long, pilosulous with appressed black and white hairs, the lanceolate-linear teeth 1.5 mm long; corolla pale violet (?); standard 26-27 mm long, the limb oblong-rhomboid, ligulately elongated upward, entire, slightly attenuate in lower 1/3, angular at base, 3 times as long as the claw; wing-petals 22 mm long, the limb linear-oblong, slightly curved, acutish, 2/3 as long as the claw; keel acute; pods sessile, ovoid, 12-14 mm long, angularly- keeled by the sutures and the valves, irregularly tetrangular, rigidly subligneous -coriaceous, hispid-villous with spreading white hairs, terminating in a straight rigid beak 2-2.5 mm long, bilocular. Fr. July. Rocks in the juniper zone.— Centr. Asia: Pam. Al. (Isfairamsai river basin on northern slopes of Alai range). Endemic. Described from Isfairam- sai, above Lyangar natural boundary. Type in Leningrad. 540. A. Borissianus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).— A. isphairamicus B. Fedtsch. in Not. Syst. Herb. Inst. Bot. Ac. Sc. URSS VIII (1940) 168, pro parte. Perennial, nearly acaulescent, suffruticose, 27-28 cm tall, from a short divided underground caudex, the old stems strongly abbreviated, clothed in old leaf petioles; stems of the year 1-7 cm long, vested with subappressed stiff hairs; stipules adnate to petiole upto the middle, ca. 10mm long, covered with appressed white hairs, the free portion of the lower ones triangular, 424 of the upper lance-linear, acuminate; leaves 10-27 cm long, the short petiole and the rachis covered with appressed white hairs, lignescent and persistent; leaflets 10-13 pairs, oblong-elliptic or lanceolate, acute, 7-15 (17) mm long, rigid, covered on both sides with appressed white hairs; peduncles half the length of the leaves, ca. 8 cm long, covered with appressed white hairs; racemes compressed, short, 6-8-flowered; bracts lanceolate, 3-4 mm long, black-pilulose; calyx cylindric, 11 mm long, covered with short white and black hairs, the subulate-linear teeth ca. 2mm long; pods sessile, oblong-ovoid, ovate in dorsal view, 14-15 mm long, obtusish, angled-carinate by the ventral suture and by the valves, complanate dorsally, triquetrous, rigidly subligneous-coriaceous, hispid- villous with spreading white hairs, bilocular. Fr. June-August. Stony mountain slopes in the juniper zone.— Centr. Asia: Pam.~Al. (Shakhimardan river basin on northern slopes of Alai range). Endemic. Described from the upper reaches of Shakhimardan River. Type in Leningrad. 541. A. distentus Boriss., see page 874. Section 87. TANYTHRIX Bge. Astrag. geront. I (1868) 114. — Perennials, with stems lignified at base, vested with spreading or ascending white bicuspidate hairs; stems well developed, prostrate; stipules not connate, adnate at base to petiole; inflorescence compactly capitate, spherical, borne on short axillary peduncles; bracteoles none; calyx cylindric-campanulate, ruptured by the pod; corolla sericeous; pods small, somewhat longer than the calyx tube, sessile, inequilaterally and subrectangularly oblong-ovoid, subtriquetrous, carinate ventrally, slightly and broadly canaliculate dorsally, rigidly coriaceous, partially bilocular, many ~-seeded. A monotypic section. Psammophytes, distributed along the Irtysh in S.W. Altai and in Zaisan depression. 542. A. roseus Ldb. Fl. Alt. III (1831) 330; Bge. Astrag. geront. II, 114; Bge. Astrag. turk., 271; Kryl. Fl. Zap. Sib. VII, 1686.— A. dasycephalus Bess. ex Stev. in Bull. Soc. Nat. Mosc. (1832) 264.— Ic.: Ldb. Ic. Fl. Ross. III (1831) tab. 300. Perennial, 10-20 cm tall; stems prostrate, well developed, (4) 7-15 (20) cm long, densely shaggy with short stiff spreading hairs, cinereous or white; stipules triangular -lanceolate in lower part, subulate-linear upward, 5-8 mm long, covered with stiff spreading hairs; leaves (5) 7-13 cm long, the rachis equaling to 1.5 times as long as the petiole, both petiole and rachis densely canescent by stiff spreading hairs; leaflets (5) 9-13 pairs, oval to oblong-oval, rarely more or less broadly lanceolate, rounded -obtuse or subobtuse, (6) 8-15 mm long, 3-6 mm broad, densely covered on both sides with stiff ascending white hairs, canescent; peduncles greatly exceeding the leaves, (0.5) 1-2 (4) cm long, densely covered with short stiff spreading hairs, white or cinereous; racemes 2-2.5 cm long, the flowers subsessile; bracts linear, equaling to somewhat exceeding half the length of the calyx, covered with stiff spreading white hairs; calyx 11-14 mm long, densely white-villous, the teeth filiform, somewhat longer than to 1.5 times as long as the tube; corolla pink; standard 13-17 mm long, the limb broadly linear-oblong, retuse, slightly auriculate-angular at base, the claw 3-4.5 times as long as the claw; wing-petals 11-15 mm long, the 425 570 limb linear -oblong, bilobate, as long as the claw; keel 9-11 mm long, the limb obtuse, somewhat shorter than the claw; ovary subsessile, densely shaggy; pods 7-9 mm long, slightly compressed laterally, 4 mm broad, ca. 3mm thick, terminating in a reflexed beak ca. 1 mm long, densely villous with spreading white hairs. Fl. May-June; fr. June-July. Shifting or more or less fixed sands.— W. Siberia: Alt.(S.W.), Irt. (S.E.); Centr. Asia: Balkh. (Zaisan depression). Gen. distr.: N. W. Mongolia. Described from Irtyshriver valley and vicinity of Ust'-Kamenogorsk. Type in Leningrad. Section 88. TRACHYCERCIS Bge. Astrag. geront. I (1868) 114, incl. sect. Borodiniana B. Fedtsch. in O. et B. Fedtsch. Consp. Fl. turk. I (1906) 227.— Acaulescent or nearly so, rarely short-stemmed, mostly cespitose low perennials, vested with bicuspidate hairs; caudex rather short, buried, divided, woody; stipules more or less adnate to petiole, free, rarely somewhat connate; flowers subradical, 1-4 in the leaf axils or in rather loose short-peduncled racemes; peduncles exceeded by the leaves; calyx cylindric or campanulate, not modified or rarely at length slightly inflated ovoid -oblong not ruptured by the pod; corolla whitish, pink, violet, or lilac, rarely yellow; standard oblong -obovate or oblong; pods sessile or borne on a very short stipe, spherical-ovoid to oblong, small, shorter than and included in or exceeding the calyx, rupturing it, coriaceous, bilocular or nearly so, rarely subunilocular. The steppe andrarely the semidesert zone of E. Europe, Caucasus, the northeru part of Centr. Asia and Siberia, from the Crimea and the Black Sea region in the west to Transbaikalia in the east. Altai, Tien Shan, Pamir-Alai, Kopet Dagh, Caucasus and Transcaucasia, Turkish Armenia, S. and N. Mongolia, Manchuria, and N. China. More or less petrophilous steppe plants; rarely psammophytes or subhalophytes. 1. Plants 2-3 cm tall; leaflets suborbicular, 7-8 pairs; pods oblong- ovoid (Caucasus) sf2% chim 516. A. involutivus Sumn. 5. Leaflets (2) 3-5 mm long, the calyx tube 3 times as long as the teeth; pods ca. 9 mm long, villous with soft white hairs; peduncles obsolescent, not more than 4 mm long. Acaulescent dwarf plants, 5-8 cm tall (Alai Walley) sedaign ken dat rorcbrieats ye etepehve oo bie 546. A. caespitosulus Gontsch. 426 + Leaflets (4) 5-15 (20) mm long, the calyx tube equaling to 1.5-2.5 times as long as the teeth; peduncles mostly very short, but all exceeding lcm; plants with strongly abbreviated stems, larger, (5) 10-20 cm Galles {pods lanaterwith stifishairs'., ec, <, <, «2; eeeokewed ihel deuce) eteoo. ba. 6. 6. Standard 20-27 mm long, the limb oblong-elliptic or oblong; claw of wings 1.5 times as long as the limb; racemes 4-8-flowered; calyx initially cylindric, at length inflated oblong-ovoid, 14-16 mm long (Steppes of S.E . and W. Siberia)....... 544. A. rupifragus Pall. + Standard ca. 33 mm long, the limb oblong-ovate, slightly ligulate- attenuate upward; claw of wings as long as the limb; racemes 5-12- flowered; calyx campanulate-cylindric, 11-12 mm long, at length slightly inflated (Tien Shan)..... 043. A. rupifragiformis M. Pop. imcalyx covered with, white, and black) hairs, 2.5. . 062 5 sb en sas « « 8. tamiGalyx covered with«purne, white hairs, soe tei o.oo feceueld 0 6 a, oslo s « « 9. 8. Pods ovaloid or rounded-ovaloid, pannose with spreading stiff white hairs, 9-18 mm long (Steppes of South Russia, Siberia, and Altai). MMM pee itil ok) sip oube Nisie oh oulpediep to. -olitsuken Sibine 548. A. testiculatus Pall. 571 + Pods obovoid-oblong, densely pannose with soft appressed white hairs (Se WieAltaiy Tarbagatai)..°. . Sie. «3. 6 >) 549. A. poliotes Bge. 9. Pods oblong-ovoid, the beak strong, subulate, as long as the body of the pod and exserted from the calyx (Balkh. sands and Ar.-Casp. }. g008 ONO 6 opcwis SUN ale tone CT CN ones Geng oe 565. A. scabrisetus Bongd. +epeakwmuchyshorter than the body of the pod .5. .........4... LO: 10. Calyx covered with appressed or subappressed hairs......... Mate + Calyx densely villous with spreading hairs................ whe 11. Leaflets glabrous above (Caucasus: Adzhariya)............... IMME SME's Scie oe sce see teens Se 562. A. salareticus Boriss. + weatlLetssimone on less*hairy above’ .") 3% se ec ne we eee we we ote Ze 12. Leaflets obovate, 4-6 pairs, obtuse; bracts ovate (S. Altai)...... soo o oo Gee > Cyaan een ceneineskeicmeah 556. A. glomeratus Ldb. + Leaflets lanceolate oval or elliptic, very rarely oblong-obovate. . 13. Ws eCOnOMAMDALERDUTDIC sacle s so sts so «0 ss se When ce ese allo ste et es 14. + Corolla yellow, rarely slightly suffused with violet .......... 15. 14. Calyx 8-9 mm long, the tube 2-3.5 (4) times as long as the teeth; standard 16-18 mm long, the limb oblong, angular at base; pods oblong, 11-13 mm long, densely lanate -villous with white hairs (Lake Balkhash area, eastern part of the Aralo-Caspian region)........ 3! ORG (bn AUS Sig) ORORC RS Atanas ae 954. A. salsugineus Kar. et Kir. + Calyx 10-11 mm long, the tube at least 5 times as long as the teeth; standard 18-19 mm long, the limb oblong-oval, attenuate in lower one- fourth, slightly angular -auriculate at base; pods ovaloid, 8-9 mm long, densely villous with soft spreading hairs (Fergana) ............. OO. Gaby D'S eon CD NEMReRG Oiich exe m deO cata ican 555. A. subauriculatus Gontsch. 15. Limb of standard ovate (Transbaikalia, Mongolia, Manchuria, N. Ching) reer ss ens RA Oe Tia Pa 559. A. scaberrimus Bge. Swim Or StandarG OOlONG Ss Areere ote sts sete ele ch etets eile beers ont 16. 16. Leaflets (2) 3-4 pairs; wings entire; pods oblong, slightly curved, 5-6 mm long (Caucasus, Transcaucasia) .. 557. A. brevifolius Ldb. + Leaflets (5) 7-12 pairs; wings shallowly retuse; pods ovoid, 6-7mm long (Caucasus, Transcaucasia)......... 561. A. humilis M.B. iveniweatlets* 4-24) pairs: 22 BAPE IO. «A. AG, nad oS PR BR 18. tee CALE E Sele TS NaN ana eit e ae eee ARENT cE Da MME MG ie cl mctta ltaM o Nar RiNe Sam otseneMs 21. 427 18. Leaflets glabrous above; calyx short-tubed, 8-10 mm long, the tube 1.5 times as long as the filiform -linear teeth (Transbaikalia, Mongolia). wa Sa aN EAR as TTY TS i, 24 SAO Aa Oe AR Ta 558. A. galactites Pall. 572) - +... leeaflets: hainy above: site a.) cid ll hNSCs 19. 19. Leaflets 8-12 (14) pairs (Steppes of South Russia, W. Siberia, Caucasus, N. Kazakhstan, and Crimea). 560. A. dolichophyllus Pall. + ‘Leaflets 4-6 pairs (Altai, Tarbapatai,) Tien Shan)? .)2).4>. Seis 20. 20. Leaflets covered with appressed hairs; limb of standard oblong- obovate (Tien Shan)! Vir. We ee AEA, By 24! 550. A. projecturus Sumn. + Leaflets covered with stiff spreading or ascending hairs; limb of standard oblonp (Altai). <0 ia a py 551. A. hypogaeus Ldb. 21. Leaves composed of a solitary lanceolate or lance~-elliptic leaflet; standard ca. 22 mm long, the limb linear; limb of wings 1.5-2 times as long as the claw; calyx-tube as long as the teeth (Tien Shan —Issyk- Kulyaradistricey Recl jal) i) Seed a Ue oY hacia, 552. A. Alberti Bge. + Leaves trifoliate, the leaflets oblong-obovate; standard 22-27 mm long, the limb obovate-oblong; limb of wings as long as the claw; calyx-tube 5-6 times as long as the teeth (Dzhungarian Ala Tau, Tien SHEN ANE TE ere Ne en LTR a a Le 553. A. Borodinii Krassn. Series 1. SUBTUMESCENTES Gontsch. — Calyx initially cylindric, at length slightly inflated, oblong, finally ruptured by the pod. 543. A. rupifragiformis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, (5) 10-20 cm tall, from a short buried woody caudex, the underground divisions short, more or less branched, ligneous, (1) 2-10 cm long, forming loose mats; stems mostly strongly abbreviated, 0.5-4 cm or rarely up to 11 cm long, densely covered with appressed hairs, nearly white; stipules adnate to petiole at base or up to 1/4, free or connate merely at base, 6-10 mm long, covered with subappressed white hairs, the free portion lanceolate to linear-lanceolate, acuminate; leaves (3) 7-10 (15) cm long, the petiole and the somewhat longer rachis covered with subappressed white hairs; leaflets 7-9 pairs, oblong-elliptic to oblong-oval, rarely broadly lanceolate, subobtuse or rounded-obtuse, (4) 6-15 (20) mm long, covered on both sides with appressed hairs, more densely so beneath; leaves (3) 4-6 times as long as the peduncles; peduncles 1.5-3 (6) cm long (rarely racemes subsessile), covered with subappressed white hairs; 575 racemes (3) 4-5 cm long, loosely 5-12-flowered; bracts linear -lanceolate, acuminate, exceeding the pedicel, 5-7 (10) mm long, covered with white or mixed white and black hairs; pedicels 2-3 mmlong, covered with white and black hairs; calyx campanulate-cylindric, at length slightly inflated, 11-12 mm long, covered with appressed or subappressed white and fewer black or nearly pure black hairs, ruptured by the pod, the tube 2-2.5 times as long as the linear-subulate teeth; corolla pale violet; standard ca. 33 mm long, the limb oblong-ovate, slightly ligulate-attenuate upward, not angular at base, obtusish and scarcely retuse, 2.5 times as long as the claw; wing-petals ca. 26 mm long, the limb linear-oblong, retuse, as long as the claw; keel ca. 20 mm long, the limb broadly semiobovate, obtuse, half as long as the claw; ovary ona stipe ca. 1 mm long; pods borne on a stipe ca. 0.7 mm long, oblong to lance-oblong, 11-16 mm long, 4-5 mm broad, carinate ventrally, canaliculate dorsally, triquetrous, 428 Bans Pi) aN : . We ry hI eee at, A: i j Viva SN \ ch : PLATE XXXVII 1. Astragalus lasiophyllus Ldb.—2, A. Skorniakovii B, Fedtsch.—3. A. Amarus Pall. 429 576 slightly compressed laterally, acutish or obtusish, terminating in a rigid beak 2-3 mm long, coriaceous, densely woolly with short stiff subappressed white hairs, bilocular. Fl. June-July; fr. June-July. (Plate XXXVIII, Figure 3). Mountain slopes, mostly stony.— Centr. Asia: Tien Shan (N. and Centr.). Endemic. Described from Turgen' in Trans-Ili Ala Tau. Type in Leningrad. 544. A. rupifragus Pall. Astrag. (1800) 86; DC. Prodr. II, 305; Ldb. Fl. Ross. I, 656; Bge. Astrag. geront. I, 116, II, 198; Bge. Astrag. turk. 277; \Schmalh. Fly 1, 280; Kryl., Fl. -ZapepsibaiVileeto ole = A. longipes Steud. Nom. ed. II, 1 (1842) 162.— A. sareptanus Beck. in Bull. Soc. Nat. Mosc. LVII, 1 (1882) 52.—Ailuroschia rupifraga Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 152.—Ic.: Pall. Astrag. (1800) tab. 70hes Exs. 3 Hor Rh. Now IO. Perennial, nearly acaulescent or more often short-stemmed, 8-15 cm tall; stems 0.5-5 (10) cm long, ascending, densely covered with stiff ascending white hairs; stipules free, lanceolate to linear-lanceolate; leaves 3-6 cm long, the petiole and the longer rachis densely covered with short white hairs; leaflets 6-12 pairs, oblong-elliptic to elliptic, 5-10 (15) mm long, 2-3 mm broad, covered on both sides with subappressed hairs; racemes rather loose, short, 4-8-flowered, subradical or more often short peduncled, the peduncles exceeded by the leaves; bracts lance -linear to linear, 5-8 mm long; calyx initially cylindric, slightly inflated in fruit, oblong-ovoid, 14-16 mm long, scabrous with white and sparse black hairs, the tube about equaling to twice as long as the linear -subulate teeth; corolla pale lilac; standard 20-27 mm long, the limb oblong-elliptic to oblong, retuse, cuneately attenuate toward base, 7-10 mm broad, longer than the claw; wing-petals 18-20 mm long, the limb linear-oblong, retuse, the claw 1.5 times as long as the limb; keel 17-18 mm long, the claw 1.5 times as long as the limb; ovary hairy; pods borne on a stipe ca. 2 mm long, bluntly triquetrous-ovaloid, 10-14 mm long, 7-8 mm broad, mucronulate ~beaked, densely white-lanate. Fl. May-June; fr. June-July. Steppes and stony slopes. — European part: Transv., Crim., L. Don (E.), L.V.; W. Siberia: U. Tob., Irt. (west of river Irtysh). Described from S. Crimea. Type in London. 945. A. caespitosulus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent, 5-8 cm'tall, from a short buried caudex with short diffuse divisions; stipules adnate to petiole up to one-third, 4-5 mm ~ long, densely covered with stiff white hairs; the free portion lanceolate; leaves 4-8 cm long, the rachis 2-4 times as long as the petiole, both petiole and rachis densely canescent with ascending stiff hairs; leaflets 7-10 pairs, oval, rarely suboblong-oval, (2) 3-5 mm long, round~-tipped, densely canescent on both sides by appressed hairs; peduncles obsolescent, 2-4 mm long, densely covered with stiff white hairs, loosely 3-4-flowered, the flowers subradical; bracts linear, 4-6 mm long, white-hairy; calyx cylindric, at length somewhat inflated, ruptured by the pod, 13 mm long, tomentose with white and black appressed hairs, the tube 3 times as long as the subulate teeth; standard ca. 28 mm long, the limb oblong, ca. 7 mm broad, retuse, angular at base, 3 times as long as the claw; wing-petals 24 mm long, the limb oblong, irregularly retuse, the claw 1.5 times as long as the limb; keel 20 mm long, the limb gibbous, obtusish, half as long as 430 oe eer a ee = 5 7 the claw; pods sessile, suboblong-ovaloid, subtriquetrous, carinate ventrally, scarcely and broadly grooved dorsally, ca. 9mm long, ca. 4.5 mmbroad, terminating in a cuspidate beak ca. 2 mm long, coriaceous, densely villosulous with spreading white hairs, bilocular. Fl. May-June; fr. June. Mountain slopes, on outcrops of red clays.— Centr. Asia: Pam.-Al. (Alai valley). Endemic. Described from Tarash River. Type in Moscow. 546. A. involutivus Sumn. in Animadvers. Syst. Univ. Tomsk. 9-10 (1936) 5. . Perennial, from a short buried caudex 5-10 mm thick, with many short offshoots; stems 4-10 cm long, herbaceous or in lower part ligneous, covered with silvery and white appressed hairs; stipules connate up to 1/4-1/3, ovate-lanceolate, covered with appressed white hairs; leaves 6-15 cm long, the petiole 2-6 cm long; leaflets 5-8 pairs, elliptic, 5-10mm long, 1-4 mm broad, covered with appressed white hairs, on sterile shoots, larger, oblong-obovate, ca. 13 mm long and 6 mm broad; peduncles firm, greatly exceeding the leaves, rarely as long, covered with appressed white and, in distal part, mixed white and black hairs; inflorescence capitately abbreviated; bracts lanceolate to linear-lanceolate, 4-5 mm long, ca. 1mm broad, exceeding the pedicel, covered with appressed black and white hairs; calyx cylindric, at length slightly inflated, 12-15 mm long, covered with more or less appressed black and white hairs, the tube 3 times as long as the subulate teeth; corolla whitish, sometimes pink; standard 25-26 mm long, the limb oblong-obovate, ca. 9mm broad, retuse, slightly constricted about the middle; wing-petals ca. 24 mm long, slightly retuse, the claw 1.5 times as long as the limb; keel shorter than the wings, intensely colored at the tip; ovary on a stipe ca. 1.5 mm long; pods sessile, erect, ovoid to ovoid-oblong, triquetrous, compressed laterally, carinate ventrally, rather shallowly canaliculate dorsally, 12-17 mm long, ca. 5-7 mm broad, rupturing the calyx, densely villosulous-lanate with spreading white and short black hairs, bilocular. Fl. and fr. May-June. Outcrops of Tertiary sandstone.— Centr. Asia: Tien Shan (northern slope of Terskei Ala Tau, Issyk-kul' lake region). Endemic. Described from the vicinity of Pokrovka. Type in Tomsk. 547. A. megalanthus DC. Astrag. (1802) 165; DC. Prodr. II, 288; Ldb. Fl. Ross. I, 627; Bge. Astrag. geront. I, 116; Ik, 198; Bge. Astrag. turk. 277; Kryl. Fl. Zap. Sib. VII, 1692.—Ic.: DC. Astrag. (1802) tab. 20. Perennial, 10-18 cm tall, from a buried woody caudex, the fairly short and rather slender divisions 2-12 cm long, forming loose mats; stems 2-6 cm long, more or less lignifed at base, densely scabrous by appressed hairs, nearly white; stipules free, adnate to petiole up to 1/4-1/3, 3-7 mm long, acuminate, covered with subappressed white hairs, the lower ovate- triangular, the upper lance-triangular; leaves (4) 6-12 cm long, the rachis somewhat longer than to twice as long as the petiole, both petiole and rachis rather sparsely covered with subappressed white hairs; leaflets 9-12 pairs, oblong-elliptic or lanceolate, rarely broadly oblong, subacute, rarely rounded-obtuse, 5-10 (15)mm long, covered on both sides with rather scattered appressed hairs, more densely so and mostly canescent beneath; leaves 1.5-3 times as long as the peduncles; peduncles 2.5 cm long, covered with short subappressed white hairs; racemes loose, (3) 4 cm long, 5-7-flowered; bracts lanceolate, obtusish, 3-4 mm long, covered 431 578 579 with white andfew black hairs; pedicels 2-3 mm long, white- and predominantly black-hairy; calyx notched-cylindric, at length narrowly inflated-ovaloid, 12-16 mm long, covered with short appressed white and black hairs (either color predominating), often longitudinally striped by predominance of black hairs on the nerves, the tube 3-5 times as long as the lance-linear teeth; corolla violet; standard 23-26 mm long, the limb oblong-obovate, retuse, slightly constricted below the upper 2/3, auriculate-angular at base, 2.5-3 times as long as the claw; wing-petals 19-22 mm long, the limb oblong, retuse, as long as the claw; keel 16-19 mm long, the limb gibbous- notched beneath, nearly straight above, acutish, the claw 1.5-2 times as long as the limb; ovary on a very short stipe, white-hairy; pods borne on a stipe ca. 1 mm long, ovaloid to oblong-ovaloid, 9-10 mm long, slightly compressed laterally, carinate ventrally, slightly canaliculate or complanate dorsally, terminating in a straight beak ca. 1 mm long, coriaceous, densely lanate with spreading white hairs, bilocular. Fl. May (June); fr. June. Steppe foothills and stony steppe-mounds.— W. Siberia: Alt. (southern part of Altai). Endemic. Described from ''Russia''. Type in Geneva. Series 2. TESTICULATI Gontsch. — Calyx cylindric, not becoming inflated, covered with black and white hairs; pods ovaloid to rounded~ovaloid or obovoid-oblong, pannose with spreading or appressed hairs; flowers subradical. 548. A. testiculatus Pall. Astrag. (1800) 82; DC. Prodr. II, 305; Ldb. Fl. Ross. I, 655; Bge. Astrag. geront. I, 116; Il, 197; Bge. Astrag. turk. 277; Kryl. Fl. Zap. Sib. VII, 1689; Schmalh. Fl. 1, 261; Grossh. Fl. Kavk.. Il, 387: — A. cinereus) a radiciflorus, DC. Astrag..4a02) 1718.i= Ag-lavelt if low us! pidbs) FL.calt.lll, (183) isa ipeptedan A. amygdaliger Less. in Linnaea IX (1834) 176; Ldb. Fl. Ross. I, 657.—A. radiciflorus Steud. Nom. ed. Il, 1 (1842) 163.— Ailuroschia testiculata Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 1522 ens Pally Astrage (QsO0) ita. Gite Perennial, acaulescent or nearly so, 5-12 cm tall; stems obsolescent and concealed beneath a dense cover of leaves, rarely up to 2-3 (6) cm long, subdecumbent; stipules free, linear-lanceolate, long-acuminate; leaves 2-10 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis densely covered with ascending stiff white hairs; leaflets 7-13 pairs, elliptic or oblong, 2-13 mm long, 1-4 mmbroad, densely scabrous on both sides by subappressed hairs; flowers subradical, crowded at the base of leaves in few-flowered inflorescences, these subsessile or borne on peduncles up to 2-6 cm long; bracts lanceolate, exceeding the pedicel but not reaching the middle of the calyx-tube; calyx cylindric, hispid-villous with ascending white and black hairs, the tube 2-4 times as long as the lanceolate teeth; corolla whitish or pinkish, or pale violet; standard 18-25 mm long, the limb oblong-obovate, 8-10 mm broad, scarcely constricted about the middle, retuse, much longer than the claw; wing- petals exceeded by the standard, the limb oblong, scarcely retuse, rather shorter than the claw; keel 14-19 mm long; ovary sessile, hairy, 15-23- ovuled; pods sessile, subradical, ovaloid to rounded-ovaloid, bluntly triquetrous, 9-18 mm long, 8-9 mmbroad, coriaceous, densely pannose with stiff spreading and rather long white hairs. Fl. April-June (July); fr. May-July (August). (Plate XXXVIII, Figure 2), 432 Steppes, on stony slopes (in Centr. Asia sporadically as a relict in montane and alpine steppes), rarely in sands and at the margins of southern pine woods. — European part: V.-Don (mainly in east), Transv., Crim., L. Don, L.V., Ural (S.); Caucasus: Cisc., Dag.; W. Siberia: U. Tob., Irt., Alt.; E. Siberia: Ang.-Say. (Krasnoyarsk and Minusinsk steppes, Balagansk district); Centr. Asia: Ar.-Casp. (northern part), Balkh. (northern part), Dzu.-Trab. (Tarbagatai), Mtn. Turkm. (Kopet Dagh, Bolshie Balkhany), Pam.-Al. (W.: western part of Mtn. Zeravshan), T.Sh. (W., Chatkal basin). Gen. distr.: N.W. Mongolia. Described from the Caspian Desert. Type in London. 549. A, poliotes Bge. Astrag. geront I (1868) 116; II (1869) 197; Bge. Astrag. turk. 276; Kryl. Fl. Zap. Sib. VII, 1691.— A. lactiflorus var. Ldb. Fl. alt. II (1831) 334; Ldb. Fl. Ross. I, 656. Perennial, nearly acaulescent, low, cespitose; stems very short, prostrate, densely pannose with white hairs; stipules free, subulate; leaves long -petioled; leaflets 10-12 pairs, oblong, acute, densely tomentose on both sides with ascending hairs; flowers subradical, subsessile, mostly solitary in the leaf axils; bracts broadly cordate, acute, very short; calyx cylindric, ca. 13 mm long, pannose with white and sparse black hairs, the tube 4 times as long as the teeth; standard ca. 23 mm long, the limb oblong, emarginate-bilobate at apex, rather faintly yellowish at base, much longer than the claw; wing-petals ca. 21 mm long, retuse; keel ca. 18 mm long, ovary 10-ovuled; pods obovoid-oblong, obliquely cuspidate, densely pannose with soft appressed white hairs, smallerthanthe pods of A.testiculatus Pall. Fl. May. Slopes of steppe foothills. — W. Siberia: Alt. (S.W.); Centr. Asia: Dzu- Tarb. (Kichkenetau mountains near Zaisan). Endemic. Described from the foothills between Bukhtarma and Talovka. Type in Paris. Series 3. HYPOGAEI Gontsch. — Calyx cylindric, not becoming inflated, densely villous with spreading white hairs; pods spherical or spherical- ovoid or ellipsoid-spherical, small, included in the calyx; flowers nearly subradical. 550. A. projecturus Sumn. in Animadvers. Syst. ex Herb. Univ. Tomsk. Nos. 9-10 (1936) 6. 580 Perennial, acaulescent, silvery-white, 8-13 cm tall; stipules free, broadly ovate, the outer densely covered with stiff appressed hairs, the inner mostly merely ciliate on the margin; leaves 3-7 cm long, the rachis 2-3 times as long the as petiole, both petiole and rachis densely covered with appressed white hairs; leaflets 4-6 pairs, oblong-obovate to elliptic, 6-10 mm long, 2-5 mm broad, short-acuminate, densely covered on both sides with appressed white hairs; flowers subradical, crowded at the leaf base; bracts linear-lanceolate, ca. 9mm long and ca. 1mm broad, hispid with long white hairs; calyx campanulate-cylindric, 13-15 mm long, villous with long spreading white hairs, the tube 3 times as long as the subulate teeth; corolla ochroleucous; standard 22-25 mm long, the limb oblong-obovate, sharply notched, 5-8 mm broad, constricted above the lower third, auriculate and angularly dilated below, much longer than the claw; wing-petals 20-22 mm long, the limb linear-oblong, slightly retuse, somewhat shorter than to as long as the claw; keel exceeded by the wings; ovary subsessile, covered with white hairs; pods spherical-ovoid, 433 981 5-6 mm long, 3-4 mm broad, coriaceous, covered with appressed white hairs, bilocular. Fl. May-June (July); fr. June-July. Gravelly mountain slopes and desert~-steppe terraces of mountain streams.—Centr. Asia: Tien Shan (Terskei Ala Tau, Issyk-Kul district and Naryn river valley). Endemic. Described from the vicinity of Pokrovka village and Irdyk-Dzhaty Oguzov district and Terskei Ala Tau. Type in Tomsk. 551. A. hypogaeus Ldb. lc. pl. Fl. ross. I (1829) 23; Fl. alt. III (1831) 329; Ldb. Fl. Ross. I, 657; Bge. Astrag. geront. I, 115; II, 195; Bge. Astrag. turk. 275; Kryl. Fl. Zap. Sib. VII, 1687.—Aulosema eximium Walp. Rep. I (1842) 634.—Ic.: Ldb. (1829) tab. 95. Perennial, acaulescent, compactly cespitosulous, the many divisions of the root-crown giving rise to stem-like offshoots; plants 6-10 cm tall, vested with grayish spreading crooked bicuspidate hairs; stipules lance- linear, slenderly long-acuminate, 12-24 mm long, hyaline, densely woolly on the outside and on the margin with long white hairs; leaves 5-12 (14) cm long, the rachis equaling to 2-3 times as long as the rachis, both petiole and rachis covered with spreading hairs; leaflets 4-6 pairs, obovate, cuneate at base, obtuse or round-tipped, 10-15 mm long, 5-9 mm broad, covered on both sides with stiff spreading or ascending hairs; flowers subradical, subsessile, borne in 2's-5's on very short axillary peduncles; bracts linear, 14-18 mm long, about equaling the calyx, hyaline, ciliate- margined; calyx cylindric-campanulate, 15-18 mm long, densely villous with long spreading white hairs, the linear teeth about as long as the tube; corolla ochroleucous; standard (27) 32-38 mm long, the limb oblong, 7-9 mm broad, retuse, auricled below the middle, the tooth-like auricles convergent; wing-petals 25-29 mm long, the limb linear-oblong, irregularly bilobate, 2/3 as long as the claw; keel somewhat shorter and broader than the wings, the limb inequilaterally obovate-oblong, slightly curved, obtuse, half as long as the claw; ovary on a stipe ca. 1 mm long, densely white- hairy, 10-12-ovuled; pods sessile, subspherical, 5-6 mm long, strongly inflated, obcompressed, complanate or scarcely concave ventrally and dorsally, slightly emarginate, terminating in a mucronate beak 1-1.5 mm long, densely pannose with short white hairs, bilocular. Fl. May-June; fr. June-July. Steppes, on bare stony slopes of hills and mountains, and in steppe valleys of mountain streams. — W. Siberia: Irt. (rare in Karkarali Mountains, Kulundin, Bel'agach steppes), Alt. Endemic. Described from Altai. Type in Leningrad. 552. A. Alberti Bge. in A.H.P. VII (1880) 375. Perennial, low, acaulescent, compactly cespitose, 3-4 cm tall; stipules adnate to petiole to high up, 7-10 mm long, clasping the scarcely developed stems and the petioles, hyaline, whitish, covered onthe back with appressed white hairs, white-ciliate on the margin, the free portion ovate triangular, short acuminate; leaves 1.5-2 cm long, the petiolule ofthe solitaty terminal leaflet as long as the blade; leaflets lanceolate to lance-elliptic, acute, 7-11 mm long, rigid, densely covered on both sides with stiff subappressed hairs, silvery; flowers subradical, the axils 2-6-flowered; calyx subinfundibular-cylindric, 10-11 mm long, densely villous with long spreading white hairs, the teeth linear-subulate, pinnately hairy, as long as the tube; corolla pink (?); standard ca. 22 mm long, the limb linear, 434 (582 3-3.5 mm broad, slightly retuse, the claw ca. 1.5 mm long; wing-petals ca. 17 mm long, the limb narrowly linear, slightly retuse, 1.5-2 times as long as the claw; keel ca. 10 mm long, the limb inequilaterally elliptic - oblong, acutish, 2/3 as long as the claw; ovary subsessile, white-tomentose. Fl. May. Valleys of mountain streams and mountain passes. — Centr. Asia: Tien Shan (Issyk-Kul district). Gen. distr.: Dzu.-Kash. (Kulja). Described from Kuyankuga. Type in Leningrad. 553. A. Borodinii Krassn. in Scripta Bot. Hort. Univ. Petrop. II (1889) 15.—A. Muschketovii B. Fedtsch. in Bull. Herb. Boiss. VII (1899) 825.— A. hypogaeus var. Borodinii Krassn. in Zap. Russk. geogr. o-va XIX (1888) 378. A. orophacoides Freyn in Bull. Herb. Boiss. 2, V (1905) 563. Perennial, dwarf, nearly acaulescent, mostly cespitulose, 3-7 cm tall, from a strong short buried caudex with short divisions; stipules more or less adnate to petiole, triangular -ovate, densely shaggy with soft hairs, ciliate-margined; leaves 2-5 cm long, trifoliate, the petiole slender, appressed-hairy; leaflets oblong-obovate, obtuse or minutely cuspidate, 8-20 mm long, silvery-canescent on both sides by appressed hairs; flowers subradical, the axils 2-8-flowered; bracts linear, about equaling the calyx, white-hairy; calyx cylindric, 12-15 mm long, densely white-villous, the linear -subulate teeth 2-3 mm long; corolla pink (?); standard 27-30 mm long, the limb obovate-oblong, narrowed above the lower one-third, angularly enlarged and auriculate below, ovate upward, retuse; wing-petals 22-27 mm long, the limb linear, round-tipped, as long as the claw; keel 21-23 mm_long, the limb convex beneath, nearly straight above, acutish, 7-8 mm long; ovary sessile, subpannose-tomentose with white hairs; pods sessile, elliptic-orbicular, 4-5 mm long, 3-4.5 mm broad, inflated, slightly compressed laterally, abruptly contracted at apex into a subulate beak ca. 1 mm long, slightly carinate ventrally, complanate or scarcely concave dorsally, densely lanate with subappressed white hairs, partially bilocular or subunilocular. Fl. May-June; fr. June-July. (Plate XXXV, Figure 1). Gravelly and stony slopes in the high-mountain zone (in Tien Shan in alpine steppes). — Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala Tau, Borborogussun), Pam.- Al. (Pamir), Tien Shan (Centr.). Gen. distr.: Kulja. Described from Charyn. Type in Leningrad. Series 4. SALSUGINEI Gontsch. — Calyx not becoming inflated, white- hairy; racemes compactly 6-15-flowered, short-peduncled; pods ovaloid or oblong. 544. A. salsugineus Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 941; Bge. Astrag. geront. I, 116; II, 197; Bge. Astrag. turk. 276. Perennial, (9) 12-20 cm tall, acaulescent or nearly so, from a short woody buried caudex, the short divisions forming fairly compact mats; stems obsolete or nearly so, rarely 1-2 cm long, densely vested with appressed hairs, whitish; stipules free, adnate at base to petiole, (3) 5-9 mm long, rather densely covered with stiff white hairs, acute, the lower triangular -ovate, the upper lance-oblong or ovate-lanceolate; leaves (8) 10-15 cm long, the petiole about as long as the rachis, both canescent by appressed hairs; leaflets 6-8 rather distant pairs, lanceolate to 435 583 984 oblong-elliptic, subobtuse, (6) 10-20 (22) mm long, (2) 4-8 mm broad, sparsely covered on both sides with appressed white hairs, glaucescent; peduncles 10 (40) mm long, densely canescent by subappressed white hairs; racemes 1.6-2.0 (2.5) cm long, 6-9-flowered, rather compact; bracts linear, 3-5 mm long, acute, green, rather sparsely covered with spreading white hairs; pedicels 1.5-2 mm long, covered with spreading white hairs; calyx cylindric, 8-9 mm long, densely covered with stiff appressed or subappressed white hairs, the tube 2-3.5 (4) times as long as the filiform ~- linear teeth; corolla pale purple (?); standard 16-18 mm long, the limb oblong, 4-4.5 mm broad, slightly retuse, angular at base, 3-4 times as long as the claw; wing-petals 14-15 mm long, the limb lance-oblong, entire at apex, 2/3 as long as the claw; keel 12-13 mm long, the limb semicircularly convex beneath, more or less straight above, obtuse, about 2/3 as long as the claw; ovary on a stipe 1-1.5 mm long, densely lanate; pods sessile, oblong, 11-13 mm long, exceeding the calyx, 5-6 mm broad, more or less compressed laterally, carinate ventrally, ca. 3 mm thick, thinly coriaceous, obtusish, terminating in a straight beak 1-2 mm long, densely lanuginous -villous with white hairs, almost fully bilocular, 16-20- seeded. Fl. May-June; fr. (May) June. More or less saline clay soils, near rivers, lakes, and draw-wells (in sands) in the desert region. — Centr. Asia: Balkh., Ar.-Casp. (chiefly E.). Endemic. Described from the Ai river valley. Type in Leningrad. 555. A. subauriculatus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent or nearly so (stems not more than 2 cm long, white by densely appressed hairs), 8-20 cm tall, from a short woody caudex, the short divisions forming rather compact mats; stipules adnate to petiole up to 1/3-1/2, free, covered with appressed white hairs, 7-8 mm long, the free portion lance-linear from a broad base, acuminate; leaves (3) 8-16 cm long, the petiole somewhat shorter than to equaling the rachis, rather slender, canescent by appressed hairs; leaflets 7-11 pairs, lanceolate or oblong-elliptic or oblong-obovate, 5-12 (15) mm long, subobtuse or rounded-obtuse, covered sparsely above and densely beneath with appressed hairs, subcanescent; peduncles 0.3-0.7 (2) cm long, densely covered with subappressed white hairs; racemes fairly compact, 2-3 cm long, 8~-15- flowered; bracts lance-linear, 2.5-3 mm long, acutish, densely white - hairy; pedicels 1-1.5 mm long, white-hairy; calyx cylindric, 10-11 mm long, densely covered with subappressed white hairs, the teeth filiform - linear, unequal, (1.5) 2 mm long; corolla pale purple; standard 18-19 mm long, the limb oblong-oval, 4.5-5 mm broad, slightly retuse, attenuate and slightly auriculate-angular in lower quarter, 3.5-4 times as long as the claw; wing-petals 15-16 mm long, the limb linear-oblong, entire, round -tipped, semicircularly convex beneath, obtuse, the claw 2.5-3 times as long as the climb; ovary short-stipitate, covered with white hairs; pods spreading or (the lower) cernuous, sessile, ovaloid, 8-9 mm long, 5-6 mm broad, compressed laterally, carinate ventrally, finely grooved dorsally, coriaceous, densely villous with soft spreading white hairs, bilocular, few-seeded, the short slender beak ca. 1 mm long. Fl. April; fr. April-May. In 'sazy'' [marshes] and ''aryks"' [irrigation canals].— Centr. Asia: Syr D. (Fergana Valley). Endemic. Described from the vicinity of Kara- Sakal village, formerly Margelan county. Type in Leningrad. 436 985 Series 5. GLOMERATI Gontsch. — Calyx cylindric, covered with appressed or spreading white hairs; flowers subradical; pods equaling or somewhat shorter (rarely much shorter) than the calyx, mostly oblong or ovaloid. 5956. A. glomeratus Ldb. Fl. alt. III (1831) 327; Ldb. Fl. Ross. I, 658; Bge. Astrag. geront. I, 116; II, 197: Kryl. Fl. Zap. Sib. VII, 1689.—Ic.: Edpe cpl bE Ross. Ill, tab?"299! Perennial, acaulescent, fromnumerous short buried stemlike branched woody shoots, forming small rather compact mats; stipules free, adnate to petiole up to 1/3-1/2, ovate, long “acuminate, 5-8 mm long, hyaline, appressed -hairy; leaves 4-8 cm long, elongating after flowering up to 15 cm, the petiole as long as or slightly longer than the rachis, both petiole and rachis sparsely covered with short appressed hairs; leaflets 4-6 pairs, obovate, obtuse or slightly retuse, 7-15 mm long, 4-9 mm broad, glaucescent with appressed short white hairs; peduncles 2-3 mm long, covered with short appressed white hairs; racemes 2-2.5 cm long, rather compactly 4-8-flowered; bracts ovate, obtusish, 1.5-2.5 mm long, barely exceeding the pedicel, hyaline, white-ciliolate; pedicels 1.5-2 mm long, covered with short appressed white hairs; calyx cylindric, 8-11 mm long, covered with short appressed hairs, subulate teeth equaling to somewhat exceeding half the length of the tube; corolla pinkish-lilac; standard 18-25 mm long, the limb oblong, 5-6 mm broad, bilobate, slightly constricted below the middle, only just auriculate-angular below, 4-5 times as long as the broad claw; wing-petals 14-32 mm long, somewhat shorter than the standard, the limb linear “oblong, scarcely retuse, erose or entire, about as long as the claw; keel 11-15 mm long, the limb rounded -gibbous beneath, nearly straight above, as long as the claw; ovary subsessile, covered with appressed white hairs, 14-18-ovuled; pods sessile, linear - oblong, ca. 13-16 mm long, 3-3.5 mm broad, pointed, carinate ventrally, broadly canaliculate dorsally, obtusely triquetrous, coriaceous, rather Sparsely covered with short appressed white hairs, bilocular. Fl. April- May; fr. May. Exposed gravelly steppe slopes. — W. Siberia: Alt. (S.), Irt. Endemic. Described from Irtysh River valley (between Voronina and Batamai). Type in Leningrad. 957. A. brevifolius Ldb. Fl. alt. III (1831) 196: Bge. Astrag. geront. II, IG.) ery el Zapaisibe Vil. 1688e— A” ga Lae tit es) "odb. Fl. Ross. i 656, ex parte, non Pall.—Ic.: Ldb. lc. plo ral. (ROSSaeLV. “tabs. SOmE Perennial, acaulescent, 3-6 cm tall, from numerous short buried branched woody stemlike shoots, forming rather compact mats; stipules free, adnate to petiole up to the middle, ovate-lanceolate, acuminate, 3-6 mm long, hyaline, covered with appressed white hairs and ciliate- margined; leaves 2-4 (5) cm long, the petiole 1.5-2 times as long as the rachis, both petiole and rachis covered with short appressed white hairs; leaflets (2) 3-4 pairs, oblong to lanceolate, subacute, 4-10 (12) mm long, 1-3 (4) mm broad, covered on both sides with scattered appressed hairs, cinerescent; peduncles 1-2 mm long; flowers 1-2 in the leaf axils, crowded at the base of leaves, the pedicels covered with appressed white hairs, 1-2 mm long; bracts lance-linear to linear, 4-5 mm long, not exceeding half the length of the calyx-tube, white -ciliolate; calyx short-cylindric, 8-10 mm long, membranous, densely covered with short appressed white 437 586 hairs, the tube 2-3 times as long as the filiform often broad-based teeth; corolla grayish-yellow; standard 20-30 mm long, the limb oblong, entire, oblong -obovate toward apex, slightly constricted below the middle, angular - auriculate below, 6-8 times as long asthebroad claw; wing-petals18-25mm long, the limb linear-oblong, slightly enlarged upward, entire, somewhat shorter than the claw; keel 16-21 mm long, the limb gently convex beneath, very slightly concave above, obtuse, somewhat broader than the wings, the claw 1.5 (2) times the length of the limb; ovary sessile, 19-21-ovuled, white-hairy; pods sessile, oblong in side view, slightly curved, 5-6 mm long, not exserted from the calyx, ovate to oblong-ovate in ground view, complanate-rounded ventrally, slightly canaliculate dorsally, coriaceous, covered with appressed white hairs, bilocular, mostly 4-seeded. Fl. June- July; fr. July. Desert steppes and desert -steppe valleys of mountain rivers, on gravelly slopes and ancient moraines. — W. Siberia: Alt. (S.E.). Endemic. Described from Chuya River valley. Type in Leningrad. 558. A. galactites Pall. Astrag. (1800) 85; Bge. Astrag. geront. I, ible 10.” bes. Perennial, acaulescent or nearly so (stems not more than 0.5 cm long, clothed in stipules of approximate leaves), (2:5) 4-6 cm tall, from a short buried woody caudex, the short divisions forming compact mats; stipules adnate to petiole up to the middle, 6-10 mm long, white-hyaline, rather densely covered on the outside and on the margin with long hairs, the free portion triangularly oblong-ovate to ovate-triangular, acutish, the upper lanceolate acuminate; leaves (2.5) 3-6 cm long, the petiole equaling to 1.5 times as long as the rachis, both petiole and rachis slender, appressed- pilulose; leaflets 6-8 pairs, lanceolate to lanceolate-oblong, 3-8 mm long, 1-2.5 mm broad, acute, rarely subobtuse, glabrous above, appressed- pilulose beneath; flowers subradical, 1-2 in the leaf axils, subsessile; bracts linear -lanceolate, often enlarged in lower part, 6-7 mm long, acuminate, white-hyaline, profusely white-ciliate on the margin, exceeding the calyx -tube but overtopped by the teeth; calyx short-cylindric, 8-10 mm long, densely villous with spreading white hairs, the tube 1.5 times as long as the filiform-linear teeth; corolla lurid; standard 22-28 mm long, the limb oblong, 6-7.5 mm broad, retuse, slightly constricted in lower third, angular-auriculate below, 2.5-4 times as long as the claw; wing- petals 20-22 mm long, the limb linear-oblong, round-tipped, somewhat shorter than to as long as the claw; keel 18-20 mm long, the limb obtusish, the claw 2-2.5 times the length of the limb; ovary sessile, hairy. Fl. : May-June. Steppes. — E. Siberia: Dau. Gen. distr.: Mong. Described from Transbaikalia. Type in Leningrad. 559. A. scaberrimus Bge. in Mém. Sav. Etr. Pétersb. II (1835) 91; Ldb. Fl. Ross. I, 649; Bge. Astrag. geront. I, 116; II, 197. Perennial, acaulescent or nearly so (stems not more than 1 cm long, clothed in approximate stipules), from a short buried woody caudex, the short divisions forming rather compact mats; stipules free, adnate to petiole up to 1/3-1/2, (3) 4-7 mm long, covered with appressed white hairs, ovate-triangular, acuminate, the upper often triangular-lanceolate; leaves (1.5) 5-10 (14) cm long, the petiole half as long as to equaling the rachis, both petiole and rachis slender, hoary with appressed hairs; leaflets 438 ofr, 4D Nx 4 ( bs y s : nee : % Ss ‘TG = 5 4 ~ a Oe Se Z IA y g \ 3 SSSI Q42 4 g aes. PLATE XXXVIII 1. Astragalus scabrisetus Bong.—2. A. testiculatus Pall.—3. A. rupifragiformis M. Pop. 589 590 5-8 pairs, oblong-elliptic, rarely lanceolate, acute or rarely subobtuse, covered on both sides with dense or rarely scattered appressed white hairs, hoary, (3) 4-12 (15) mm long; peduncles (2) 3-8 (17) mm long, covered with appressed white hairs; racemes often subsessile, loosely 3-5-flowered, 1.8-2.5 cm long; bracts lanceolate, acute or acuminate, 2-3 mm long, white-hairy; pedicels 1.1-5 mm long, covered with white hairs; calyx cylindric, 7-9mm long, densely hoary by appressed white hairs, the tube (1.5) 2-3 (4) times as long as the linear teeth; corolla yellow, the keel and standard often suffused with violet; standard 20-21 mm long, the limb ovate, retuse, 8-9 mm broad, abruptly contracted into and twice as long as the claw; wing-petals much shorter than the standard, 16-17 mm long, the limb obliquely oblong -obovate, slightly retuse, as long as the claw; keel 12-13 mm long, the limb acute, the claw 1.5 times the length of the limb; ovary sessile, covered with short white hairs; pods sessile, lance -oblong, falcate, somewhat curved, pointed, (8) 10-13 mm long, 3.5-4 mm broad, compressed laterally, carinate ventrally, grooved dorsally, terminating in a straight beak not exceeding 0.6 mm, coriaceous, densely covered with appressed white hairs, bilocular, few-seeded. Fl. May-July (August); fr. June-August. Steppes. —E. Siberia: Dau. Gen. distr.: Mong., Manchuria, N. China. Described from Transbaikalia. Type in Paris. 560. A. dolichophyllus Pall. Astrag. (1800) 84; Bge. Astrag. geront. T; 116;, 11; 3196; Schmath: Fle 1,“281;, Grossh. Fie Kavk. Hyiseen An diffius us) Willd) Sip.spl. MT (1800) 1321; Ldb. vel. Ross aihwodgome. Ailuroschia diffusa Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 152.— Ic.: Pall. Astrag. (1800) tab. 68. Perennial, acaulescent or nearly so (stems rarely up to 1 cm long), compactly cespitulose, 6-15 cm tall; stipules slightly adnate to petiole, linear-subulate to lance-subulate, 7-11 mm long, covered with stiff spreading white hairs; leaves (5) 7-12 (15) cm long, the petiole andthe much longer rachis covered with short stiff spreading or ascending hairs; leaflets 8-12 (14) pairs, oblong to lanceolate, subobtuse to acute, 5-15 mm long, 1.5-4 (5) mm broad, densely covered on both sides with stiff ascending or spreading hairs; racemes very short, few-flowered, subsessile, crowded at the base of leaves; bracts linear, equaling the calyx tube, hispid with white bristles; calyx cylindric, 10-13 mm long, densely villous with spreading white hairs, the linear -setaceous teeth as long as or somewhat longer than the tube; corolla ochroleucous; standard 25-30 mm long, the limb oblong, slightly retuse, slightly constricted in lower third, slightly auriculate -angular below, 3-4 times as long as the claw; wing- petals 19-25 mm long, the limb linear-oblong, retuse, the claw 1.5 times as long as the limb; keel 16-20 mm long, the limb semioval, rounded- obtuse, half as long as the claw; ovary subsessile; pods sessile, ovaloid to oblong-ovaloid, obtuse, 7-11 mm long, 4-5 mm broad, carinate ventrally, rounded-convex dorsally, coriaceous, densely hispid-villous with short ascending white hairs, partially or almost fully bilocular, the beak hard, subulate, recurved toward the ventral side, ca. 3 mm broad [?]. Fl. May; fr. (May) June-July. Sands in steppes of the semidesert region. — European part: M. Dnp., U. Don (S. W..); Transv.(S.); 'Bl., Crim., L: Don; Ls V0; Caucasus: Cisc., Dag.; W. Siberia: U. Tob. (S.); Centr. Asia: Ar.-Casp. (northern) part; in south as far as Emba River and sands of Bolshie Barsuki). Endemic. Described from the Aralo-Caspian Desert. Type in London. 440 591 561. A. humilis M.B. Fl. taur.-cauc. II (1808) 203; Ldb. Fl. Ross. I, 655; Bge. Astrag. geront. 1, 116; II, 147; Grossh. Fl. Kavk. II, 338.— Perennial, acaulescent, 4-10 cm tall, diffusely cespitulose, from a short buried caudex with short divisions; stipules adnate to petiole in lower 1/3-1/4, lanceolate to linear-lanceolate, 5-6 mm long, acuminate, more or less covered with white hairs; leaves (2) 5-10cm long, the rachis 1.5-4 times as long as the petiole, both petiole and rachis rather densely covered with short appressed hairs; leaflets (5) 7-12 pairs, oblong-oval or oblong- elliptic, subobtuse or rounded-obtuse, (2.5) 5-10 mm long, (1.5) 2-3.5 mm broad, appressed-hairy on both sides; racemes subradical, 3-5-flowered, strongly abbreviated, subfasciculate; bracts linear, 5-7 mm long, more or less covered with white hairs; calyx campanulate-cylindric, 7-10 mm long, densely covered with appressed white hairs, the tube 1.5-3 times as long as the linear-subulate teeth; corolla yellow; standard 22-25 mm long, the limb oblong, scarcely retuse, slightly attenuate in lower quarter and slightly enlarged below, 2.5~-3 times as long as the claw; wing-petals 17-20 mm long, the limb linear-oblong, rather shallowly retuse, the claw somewhat longer than to 1.5 times as long as the limb; keel 15-17 mm long, the limb semiobovate, obtusish, half as long as the claw; ovary on a stipe ca. 0.5 mm long; pods sessile, ovoid, 6-7 mm long, 5 mm broad, rounded- obtuse, inflated, carinate ventrally, canaliculate dorsally, coriaceous, densely villous with appressed white hairs, bilocular to 1/2, the beak subulate, obliquely bent toward the ventral side, 3 mm long. Fl. May-June; fr. June-July. Dry mountain steppes and clayey slopes of mountains and foothills. — Caucasus: Cisc. (Terek), Dag., E. and S. Transc. Endemic. Described from the Caucasus. Type in Leningrad. 562. A. salareticus Boriss. nomen novum.— A. Barbeyanus Freyn in Bull. Herb. Boiss. III (1895) 184, non Post (1893); Grossh. Fl. Kavk. ll, 333. Perennial, acaulescent, sparsely vested with appressed hairs, up to 10 cm tall, from a short woody buried caudex; stipules adnate to petiole to high up, sparsely appressed-hairy, the free portion short-ovate; leaves up to 11 cm long, the petiole rather long; leaflets 9-14 pairs, broadly elliptic, obtuse, up to 7 mm long and up to 4 mm broad, glabrous above, appressed-hairy beneath; inflorescence loosely capitate, subradical, 2.5-5 cm long; bracts lanceolate, hyaline, sparsely covered with stiff hairs, 1.5 times as long as the pedicel and somewhat shorter than the calyx-tube; calyx cylindric, 10-11 mm long, sparsely covered with stiff appressed black hairs, these thinly interspersed with white, the tube 4 times as long as the triangular-subulate teeth; corolla purple; standard 24 mm long, the limb linear-oblong, shortly bilobate, obtusely auriculate at base, exceeding the wings; ovary stipitate, woolly. Fl. May. Mountain slopes, at altitudes of about 2100 m.— Caucasus: W. Transc. (Adzhariya). Endemic. Described from the vicinity of Salaret. Type in Geneva. 563. A. Theodori Grossh. In Proc. Ac. Sc. Azerbaijan SSRI, 1 (1945) 33. Perennial, acaulescent, 2-3 cm tall, vested with appressed gray hairs; leaves 2-5 cm long; leaflets 7-8 pairs, small, 2-3 (5) mm long, ovate- orbicular or broadly ovate, round-tipped; inflorescence few-flowered, 441 592 sessile; calyx covered with appressed white hairs, the unequal teeth somewhat shorter than to as long as the tube; corolla undescribed, pods oblong-ovoid, 8-10 mm long, grooved dorsally, mucronulate, covered with appressed white hairs. Fr. July-August. Subalpine zone, on dry slopes, at altitudes of 1140-1500 m.— Caucasus: Dag. (Levashi), E. Transc. (eastern part of Greater Caucasus — Shemakha district; Gil'skii district, Gyumur). Endemic. Described from Shemakha district. Type in Baku. Note. Hardly justifying separation from A. humilis MB. 564. A. erivanensis Bornm. et Woron. in Monit. Jard. Bot. Tiflis, f. 34 (1914) 1.—A. pseudohumilis Grossh. in Tr. azerb. otd. Ak. Nauk SSSR 1 (1933) 54. Perennial, acaulescent, 6-10 cm tall, cinerescent or cinereous by dense spreading bicuspidate white hairs; caudex short, with many divisions; stipules lance-linear, 6-9 mm long, covered with long bristles; leaves covered with spreading white hairs, the petiole as long as the rachis; leaflets (8) 10-14 pairs, yellowish-green or gray, crowded, oblong-elliptic, obtuse to subacute, 10 mm long, 4 mm broad; racemes sessile, few- flowered; bracts linear, 7-10 mm long, equaling the calyx-tube; calyx covered with long spreading white hairs, 8-9 mm long, the subulate teeth half as long as to equaling the tube; corolla 2.5-3 times the length of the calyx, 25-26 mm long, yellow; standard oblong, ca. 6 mm broad; wing- petals ca. 4mm broad, bilobate at apex; ovary covered with appressed white hairs; pods densely white-tomentose, ovoid, triquetrous, acuminate. Bune ser siuliy;. Stony southern slopes, at altitude of 2300 m.— Caucacus: S. Transc. (Erevan and Surmalin districts, vicinity of Sami at Lake Sevan). Possibly growing in Iran. Described from Sami vicinity. Type in Tbilisi. “Note. The description of pods is taken from the description of A. pseudohumilis Grossh. which is undoubtedly synonymous with this species. Series 6. DOLICHORRHYNCHI Gontsch.— Calyx not becoming inflated, covered with white hairs; flowers subradical; beak of the pod as long as the body, exserted frorn the calyx. 565. A. scabrisetus Bong. Verzeichn. der am Saisang-Nor gesammelten Pfl. (1841) 26; Ldb. Fl. Ross. -I, 657; Bge. Astrag. geront. II, 195; Kryl. bls Zap. olbamvVill L601. eailGo we ONP el Come LAD me ; Perennial, acaulescent or nearly so, cespitulose, 5-15 cm tall; stems of the year strongly abbreviated or obsolescent, not more than 4 cm long, tightly clothed in vestiges of petioles and stipules; stipules adnate to petiole in lower 1/4-1/3, covered with stiff white hairs, the lower oblong-ovate, the upper lanceolate, acuminate; leaves (3) 5-11 (16) cm long, the petiole as long as the rachis, both petiole and rachis covered with appressed silvery hairs; leaflets (2) 3-8 pairs, elliptic or obovate, subobtuse, rarely acute, 5-10 (13) mm long, 2.5-5 (8) mm broad, densely covered on both sides with stiff appressed hairs, whitish; racemes many, compactly capitate, ca. 2 cm long, borne on very short peduncles at the base of leaves, forming a spurious compound ''inflorescence''; bracts lance-linear to linear, long-acuminate, 4-6 mm long, covered with spreading white hairs; calyx 442 193 cylindric, 8-11 mm long, densely covered with spreading white hairs, the tube 1.5-2 (3) times as long as the linear-subulate teeth; corolla white, the tip of the standard and of the keel violet; standard (14) 16-19 mm long, the limb oblong, 5-6 mm broad, round-tipped or scarcely retuse, somewhat auriculate and angularly enlarged below the middle, 4-5 times as long as the claw; keel (9) 12-14 mm long, the limb oblong-oval, obtuse, half as long as the claw; pods sessile, oblong-ovoid, 6-8 mm long, convex ventrally, slightly canaliculate dorsally, subtriquetrous, prolonged into a beak, covered with stiff ascending white hairs, bilocular, 2-10-seeded, the beak long, straight or slightly curved, firm, as long as the body of the pod, exserted from the calyx. Fl. May: fr. June-July. (Plate XXXVIII, Figure 1). Sands, rarely sand- and stone-covered sites or loams in semidesert and desert regions.— Centr. Asia: Ar.-Casp., Balkh. Endemic. Described from Zaisan depression. Type in Leningrad. Section 89. CHOMUTOVIANA B. Fedtsch. in O. et B. Fedtsch. Consp. Fl. turk. I (1906) 228.— Low, compactly cespitose, nearly acaulescent perennials, vested with bicuspidate hairs; stipules connate; stems strongly abbreviated and hardly discernible beneath the compact cover of leaves; leaflets few or 1 or 2 pairs; racemes short, rather compactly capitate, few-flowered, borne on short peduncles; calyx cylindric; corolla violet; standard oblong-elliptic; pods sessile, oblong, equaling or somewhat exceeding the calyx, inflated, prominently sutured, coriaceous, white-villosulous, almost fully unilocular (the septum confined to the base or the middle part of the pod). A section associated with Central Tien Shan and Pamir. High-mountain deserts and semideserts. 966. A. Chomutovii B. Fedtsch. in Bull. Herb. Boiss. VII (1899) 826; Fl. Tadzhik. V, 429.—A. Enaphae B. Fedtsch. et N. Bazil nom in herb. Perennial, dwarf, 2-8 cm tall, the many very short shoots forming compact mats; stipules few, connate, hyaline, covered with white hairs; leaves 1-4 cm long, trifoliate or imparipinnate with 2 (3) pairs of leaflets; leaflets oblong to oblong-elliptic, 5-15 mm long, densely covered on both sides with appressed silvery hairs; peduncles 1.5-4 (6) cm long, covered with appressed white hairs; racemes rather compact, 1-3 em long, 5515- flowered; bracts linear-lanceolate, 3-4 mm long, hyaline; calyx cylindric, (4) 6-8 (10) mm long, densely covered with appressed black and white hairs, the tube 4-5 times as long as the triangular or more often linear-lanceolate teeth; corolla violet; standard (10) 15-20 (22) mm long, the limb oblong- elliptic, retuse, gradually attenuate toward base; wing-petals (8) 17-18 (20) mm long, the limb oblong, as long as the claw; keel (6) 11-14 (16) mm long, the claw somewhat longer than to 1.5 times as long as the limb; pods sessile, oblong, slightly falcate, (6) 8-9 (10) mm long, strongly inflated, prominently sutured, terminating in a cuspidate beak 1-1.5 mm long, coriaceous villosulous, with white hairs, almost fully unilocular. Fl. June- August; fr. July-August. (Plate XXXV, Figure 2). High-mountain deserts and desert steppes.— Centr. Asia: Pam.- Al. (Pamir, Alai valley), Tien Shan (Centr.). Gen. distr. : Kashgaria. Described from Pamir. Type in Leningrad. 443 594 595 Section 90. PROSELIUS (Stev.) Bge. Astrag. geront. I (1868) AG, excl. A. Frickii Bge. II (1869) 198.—Gen. Proselias Stev. in Bull. Soc, Nat. Mosc. IV (1832) 268.— Gen. Euilus Stev. l.c. XXIX (1856) 149. Racemes dense or loose, sometimes spikelike, long-peduncled; bracts herbaceous, persistent and, in addition, 2 bracteoles attached at the base of calyx; calyx cylindric, commonly gibbous at base, not becoming inflated, sometimes at length ruptured; corolla deciduous or sometimes persistent; ovary subsessile or stipitate; pods bilocular, ovate or oblong to linear, tetrangular or cylindric, terete or laterally compressed, rigid, coriaceous or tightly-textured membranous, appressed-hairy, rarely villous with spreading hairs or glabrous. Perennials, acaulescent or nearly so, herbaceous or occasionally lignified at base, the short caudex sometimes branched, clothed with imbricated stipules and petioles; plants vested with bicuspidate white or sometimes black hairs, rarely glabrous; stipules petiolar, shortly adnate to petiole; leaves imparipinnate, sometimes consisting of a solitary leaflet, marscescent. The species of the section Proselius are chiefly distributed through the Mediterranean region, speading westward to North Africa and Spain; in the east certain species (10) reach as far as Pamir-Alai and Tien Shan. The widest range of species of the section Proselius is associated with the Caucasus, Asia Minor, and Iran. 1. Pods villous with spreading white hairs, ovoid or oblong; leaflets orbicular or lanceolate, large, 10-20 mm long, 6-13 mm broad (Centr. Asian lave, cA acaniedhte raat at eaartliiyeed te hdl seal eee MSRP rsh Bve) oc ae + Pods covered with appressed hairs or glabrous, ovoid to oblong or narrowly linear, cylindric or flattened; leaflets varying in size and shape(Cauecasus, ‘Centr: A’sia; Europe)... > .') |”. | 2) eee See 3. 2. Pods reflexed, disposed in a long loose raceme, the beak short, straight; leaflets orbicular, glaucescent; standard spatulate, enlarged in upper part, slenderly long-cuneate toward base........+2.+.e-. OF ea cea OR Ch ors Oe Coho cat nacienn tec ie hc 614. A. namanganicus M. Pop. + Pods erect, gathered in a dense oblong or ovoid raceme, the beak long, reflexed; leaflets lanceolate to oblong-lanceolate; standard oblong, broadest about the middle, broadly short-cuneate toward base. Sr bleh See ee ee eee ee ee eee ee 613. A. hissaricus Lipsky. 3. Leaflets 1-3 pairs, obevate= orbicular, 7-15 mm long, 5-10 mm broad; pods cernuous (Centr. Asia): ..... -- 604. A. brachyrachis M. Pop. + Leaflets 4-16 (20) pairs, ovate to linear, large, the lower leaves sometimes unifoliate or trifoliate, usually large, cylindric in fruiting, eneeh. aa: ape ee ONE LERENYCS, sts elie ie ete hat? ah De hk LOS PROMS ESS ope, 2 en et 4. 4. Leaflets solitary or 3-11 (13) pairs (leaves with number of leaflets ranging from 1 to 27 often to be found on the same plant); leaflets mostly large, 1-7 cm long, 1-5.5 cm broad, ovate-oblong to orbicular; pods narrowly cylindric ,w4-7/em long,ierect» 2... bec ae 29. +) Plantsidiffening fronajabome:-\iwas ic) is @ mle tein). Se 5. Pods linear-cylindric, 5-9 cm long (including beak), the beak subulate, (7) 10-15 (20) mm long; leaflets linear to oblong-linear, glabrate or Spansely,appnessed hairy abovieuwesg-i ewer us Bela a ee 6. + Pods 0.7-2 or 2-4.5 cm long, rarely up to 6 (7) cm and then the beak not more than 1-3 mm long or, if 4-6 mm, the flowers yellow and fruits pendulous; pods sometimes more or less flattened; leaflets ao 444 596 wo t 10. fl. 12. 13. 14, 15. linear to oblong or ovate or orbicular, mostly appressed~hairy above ie pollen ots) Pal ae aa ies ol oon em nen CaM mami teas ie eon tse een ie Leaflets 7-15 pairs; calyx-tube 2-3 times as long as the teeth; beak of the pod ca. 10mm long......... 573. A. zangelanus Grossh. Leaflets 6-8 (10) pairs; calyx-tube 4-6 times as long as the teeth; beak of the pod 10-15 (20) mm long.... 572. A. rostratus C. A.M. Pods oblong or ovoid, mostly firm, coriaceous, inflated, sometimes subtriquetrous, erect or pendulous;. ovary oblong or ovoid..... PAPA 6 Pods oblong-lanceolate or linear, cylindric, straight or curved, erect or pendulous, or oblong and more or less flattened, not inflated; Ovanyelhnéaninarely oblonga linear weit byaeecitievs tolsivabey ey. 8. Pods pendulous, sometimes in part horizontally spreading or erect and then the leaf petioles firm, partially persistent and becoming PSPC OULU SIS eifen Mike: olvloWlcls ole cd ciel sheleuh ety Woh cio thy olhioble wieulely aidan ofebiatt sWislie « sil/e 9. Pods erect; leaf petioles not becoming lignified at base ....... 18. Pods borne on a long stipe exceeding the calyx, oblong-lanceolate, dark-spotted; leaflets small, oblong, 4-10 mm long, obtuse, 4-8 pairs. ERRete Meine Raw cute Atel Bl Ce Siig) Recreoile iceman bareee cited esha ay ary 599. A. punctatus Bge. Pods borne on a short stipe included in the calyx, or sessile... .10. Some vewlows sy kay Wieser sh tna ehi owls sy LIN es seit Powel esis SOP ct lied eai/shiie ik Corolla purple or pale violet or azure, drying violet ......... 12. Leaflets 9-11 pairs, acute, oblong-lanceolate; pods triquetrously cylindric, 25 mm long, 3 mm thick, the beak 4-6 mm long....... Pees) cehis BUS SUITE woicue L E ee cementaep ys 596. A. refractus C.A.M. Leaflets 5-7 pairs, obtuse or subacute, ovate to ovate-oblong; pods Gbilong-cylindric,)15-lS8imm~ long; ‘short sbeaked sic. qaiie: aot sien oe) ne PAPANMA Soh terey CANN Sh k)) Wtetiretied ot ad Pasheniian lol voalres Yana 602. A. campylosema Boiss. Stipules glabrous; leaflets 15-16 pairs, oblong, obtuse, up to 2.5 cm TORTS PSeshe) wigs s SS a chet eS Roiem dh paltomeithe 6p 601. A. fodinarum Boiss. Stipules covered with appressed or sometimes partly with spreading hairs; leaflets 3-12 or rarely 14 pairs or sometimes solitary on the lower leaves, mostly acute or subacute, rarely subobtuse...... 3! Pods narrow, linear-cylindric, 2-3 mm broad and 18-35 mm long, straight; peduncles equaling or slightly exceeding the leaves; base of plant becoming lignified, clothed in persistent divergent vestiges of EA Sp RPE THOWE SUP Wi pB.: «cei eae sy os vielen e Pouce clbeiiel) o\labelp Ulett al. Jetells 14, Pods oblong-cylindric, straight or falcate, 14-30 mm long and 3-5 mm broad; peduncles about equaling the leaves or exceeding them and up to 10 cm long and then fruits slightly arcuate; petioles not INGUEESCERL o! 5,s,'5, ose alana oie Gs Micaela Sen 3: abit Spxsthretp eas aitena bead oil's. 15. Leaflets linear, 8-12 pairs, 3-12 mm long, 1-1.5 mm broad; corolla purples) pods 18-23) mmilong.. . 4 wise. « 595. A. fuliginosus Beck. Leaflets oblong-lanceolate or elliptic, 5-8 pairs, 2-10 mm long, 1.5-3 mm broad; corolla pale violet; pods 25-35 mm long....... SFT E ICN AN) Pus Co ob aePOyle EM A 594. A. ordubadensis Grossh. Plants ca. 10 cm tall; leaflets (7) 9-14 pairs, 4-6 mm long, 2-3 mm broad; bracts ovate or lanceolate ...... 600. A. longicuspis Bge. Plants (15) 30-40 cm tall; leaflets 3-10 pairs, 7-25 mm long, rarely up to 45 mm long, 5-13 mm broad, the leaves of young plants sometimes consisting of a solitary obtuse leaflet; bracts narrowly lanceolate or) Hnear Torfilifor riba | Sinan ds eee weenie nial eis fey eye oye 16. 445 GE Wile 597 18 (8). 19. 20. 21. 22 (7). Pods 14-18 (20) mm long, oblong-cylindric, gathered in a long dense raceme, strongly reflexed and appressed to peduncle; leaflets A-NOl pairs a Wierw.mewen cee 598. A. atripurpureus Boiss. et Heldr. Pods 20-30 mm long, narrowly lanceolate to lanceolate, disposed in a long loose raceme, reflexed but not appressed to peduncle, or spreading; leaflets 3-7 pairs, sometimes the lower leaves consisting of a solitary large leaflet ........ 22s cee cseves Lt Pods glabrous, speckled, straight or nearly so, reflexed; leaflets 3-5 pairs; bracts 2-3 mm long; calyx 9-10 mm long (Kopet Dagh). TOES, AUP TAR Ne tek ee rons 603. A. ackerbergensis Freyn. Pods appressed-tomentose with black and white hairs, faintly speckled when young, falcately curved in distal part, subhorizontally spreading; leaflets 4-7 pairs; bracts ca. 5 mm long; calyx ca. UZ; sooWaal Moreen GC bolo oad Omi 6.010 do 00 G6 597. A. cuscutae Bge. Pods compressed laterally, 10-25 mm long, 4-5 mm broad, slightly curved or straight, oblong, sometimes speckled; leaflets 7-15 pairs; inflorescence a short raceme.......-..++e-es+ese0-. 1S) Pods cylindric, linear or oblong-lanceolate, 14-60 (70) mm long, 2-5 mm broad, sometimes somewhat compressed laterally but 40-60 (70) mm long in fruit; leaflets 5-7 or (8) 10-20 pairs; inflorescence usually a long raceme, loose or becoming so in fruit. tS TOUS BOOG (OILS. SHSLT BOOLAO. PERI 2. RNG “io 32. Corolla whitish; pods rigidly coriaceous, slightly flattened, sometimes bluntly triquetrous, 10-15 mm long, and ca. 5mm DEGAAIO TILE, AF IR 4 BTR RD eS 586. A. salatavicus Bge. Corolla purple, violet, azure, or ochroleucous, with a dark purple blotch on the keel, drying yellowish; pods (15) 20-25 mm long, 4-6$immebroad .Wesisrhimm ie AIO e Gee Se Sir ocr nena cies 20. Calyx 10-15 mm long, the tube 3-5 times as long as the teeth; corolla violet or ochroleucous, with a dark violet blotch on the keel, 20-26 mm long; leaflets small, 1-4 (5) mm long, 1-4 mm broad; peduncles and leaf rachises slender, weak..........2+.266-. 21. Calyx 15-18 mm long, the tube twice as long as the teeth; corolla purple, 25-33 mm long; leaflets 5-10 mm long, 3-8 mm broad; peduncles and leaf rachises rather thick and firm ........... Cihelk sitre a o.com Malmo pa eee eee ee se «6983. As Schizopterus Boiss. Leaflets 5 or more often 7 pairs, 3-4 (5) mm in circumference, mostly orbicular, rarely ovate; stipules broadly lanceolate, barely adnate to petiole, not acuminate; pods carinate on one side and shallowly grooved on the other, stipitate ........-.-..+seee-. 3 oP T SE ILUERY Secirne: TF MARES «TIMER BR SN 582. A. czorochensis Charadze. Leaflets 10-15 pairs, greenish, 1-3 (5) mm long, 1-2 (3) mm broad, elliptic or ovate; stipules narrowly lanceolate, adnate to petiole nearly up to the middle, acuminate; pods carinate on both sides} Subsessille’: se! Win cree cr enems 584. A. sanguinolentus MB. Podstpendulows* 2 tes Se Te Rs Rel ottetks Tol te. eis MP MONNAT OR 23. Pods erect or sometimes merely some of the pods or flowers horizontally spreading or pendulous. ........+.+-+.2+seee-. 24. Pods oblong, covered with black bristles, subapplanate, acuminate toward both ends, obliquely mucronulate, rounded on one side, carinate on the other; leaflets 8-10 pairs, elliptic, whitish-gray. RS SA EDs) OE CRN NS ite ee cst, ie 611. A. coelestis Boiss. 446 598 499 24, 25. 26. 27. 28. 29 (4). 30. 31, Pods ovoid or oblong, appressed-hairy; mucronulate, resembling pistachio fruits; leaflets 4-6 pairs, elliptic, Whiteshairyene ne: Pt Mop ikret Mins rates o Redbolt. Ey cs Sete a eA NN 5, 612. A. elongatus Willd. Coxolllayencenishsyellow orluridal VAsekaoewwee ue os aRbe. 25. Corolla drak purple, violet, dingy violet, or whitish ....... 26. Corolla lurid; leaves 20-25 cm long; peduncles 25-45 cm long; pods oblong-ovoid, carinate, not grooved dorsally........... CPM kP A Calla: Oke eae SU Beet. ecole 606. A. robustus Bge. Corolla greenish-yellow; leaves 7-10 cm long; peduncles 15-20 cm long; pods oblong or ovoid, scarcely carinate ventrally, slightly emarginate dorsally, sometimes horizontally spreading, rarely PENGUWIOUSMH Ma Ba See RL a at 608. A. cinereus Willd. Flowers dark purple, 18-20 mm long; pods oblong-ovaloid, equally broad throughout, tomentulose to glabrate........... Eee PANE are IN ceer Kine heme iy RAPIER Mik TS ae 609. A. taschkendicus Bge. Flowers violet or dingy violet or whitish; 20-25 mm long; pods ovoid or oblong-ovaloid, attenuate distally or proximally, or equally broad throughout, appressed-hairy.............. Zits Pods oblong, somewhat flattened, terete, faintly wrinkled lengthwise; ovary not attenuate toward the base or covered with predominantly black hairs....... 607. A. subrobustus Boriss. Pods attenuate distally or proximally, ovoid-triquetrous, clavate, cross-wrinkled; ovary ovoid, attenuate upward or toward base. SMe COM: eerste ok CAT Corey Vole Meh LURE aD STA at co Liman Smee Eh! Bey kee Ye, 28. Pods attenuate toward base, clavate; corolla dingy violet or purple; leaflets 7-15 pairs, ellipsoid to oblong-ellipsoid; stipules 5-10mm loneq(Caucasus))i.w.«mave! eevee veka 605. A. brachycarpus M.B. Pods enlarged toward base, broadly ovoid-triquetrous; corolla violet or whitish; leaflets 4-8 or more often 5-6 pairs. ellipsoid, subspherical; stipules 10-13 mm long (Centr. Asia).......... 5-10 30) Sen Aol eae Ot LO MEMOR eee ora Ae 610. A. platyphyllus Kar. et Kir. Leaflets green, glabrous above or sometimes sparsely appressed- hairy beneath; calyx covered with few appressed hairs; bracts narrowly lanceolate, glabrous or sparsely ciliate-margined, (5) 7-10 mm long; leaves consisting of single leaflets or 3-11 (13) pairs of large leaflets....... 970. A. heteromorphus Boriss. Leaflets appressed-hairy on both sides; calyx densely covered with appressed hairs; bracts appressed-hairy, ovate, 1-2 mm long, orslanceolate wo 4immmongyn. et face ERs CMe Re IS Soee. 305 Calyx-tube 8-10 times as long as the triangular obtusish teeth; pods 3.5-4.5 cm long, the beak straight, firm, 10-15 mm long; inflorescence 2-5-flowered ........ 571. A. Rollovii Grossh. Calyx tube twice as long as the teeth; inflorescence many-flowered, elongated; leaflets 4-5 pairs ....... 569. A. Fedorovii Takht. Calyx-tube 2-5 times as long as the linear-lanceolate teeth; pods 3-8 cm long, the beak 2-4 mm long; inflorescence many-flowered, or 2-6-flowered and then leaves consisting of single leaflets or of Soo Veatlevs tw. itetat. seh ade Sebeeed, sbbeer ate. 5 didee 6 Gino 6.46 31. Leaves consisting of single leaflets or sometimes the lower leaves comprising 3-5 orbicular or orbicular-ovate leaflets 8-15 mm long and 6-18 mm broad, truncate-based; bracts ovate, small, 1-2 mm long; inflorescence 2-6-(rarely more)-flowered ............. ee Ne sas ee cos a eset Stay at 967. A. Candolleanus Boiss. 600 32 (18). 35. 36. 37. Sa 38 (32). 39. 40. Leaves consisting of a single leaflet or 1-6 (7) pairs; leaflets ovate to oblong-ovate, 5-20 (35) mm broad, 10-35 (50) mm long, cuneate-based; bracts lanceolate, 3-4 mm long; calyx 12-14 mm long; inflorescence many-flowered... 568. A. latifolius Lam. Limb of standard broadly quadrangular-ovate, mostly enlarged in upper and lower part; corolla yellow, often becoming red, mostly: drying darkured/(Centr.e Asia) “ico. .becye woeliia e ge 38. Limb of standard oblong, attenuate in upper part; corolla purple or violet or azure or whitish, with a dark blotch on the keel (Caucasus;iBurope)iacm Glocweas 4» bierm.eo eee eee 23, Peduncles shorter than the leaves; pods glabrous,........ 34. Peduncles equaling or exceeding the leaves; pods commonly appressed tomentose. narely:glabnate). . te cu sage bee 35. Stipules glabrous; calyx (10) 12-15 mm long, the tube 5-6 times the length of the teeth; pods falcate; leaflets 6-7 pairs, 5-10 mm long; moviateze lip tale uly ciipsiy a fiat oy supe) neon 980. A. cyri Fomin. Stipules densely appressed-hairy; calyx 15-18 mm long, the tube twice the length of the teeth; pods straight or slightly curved; leaflets 10-12 pairs, 15-20 mm long, ovate or orbicular...... dais ERAN oreorceticyikre 578. A. glaucophylloides Bornm. et Woron. Leaflets 5-7 pairs, small, 3-4 (5) mm long, broadly obovate; inflorescence 3-7-flowered; corolla whitish, with a reddish- mio letiblotcheacie:s ee eet hte 2 579. A. montis-aquilis Grossh. Leaflets 8-20 pairs, oblong-ovate or orbicular, 4-20 mm long; inflorescence many~flowered, mostly a loose, rarely dense, raceme: qin vineu tla amoinen ot biomes i awe ee a 36. Calyx 10 mm long, the teeth 1-2 mm long; pods 20-25 mm long, covered with appressed white and black hairs; bracts subulate, 3mm long; corolla dingy violet; leaflets 8-12 pairs........ SS ae PES! 3 pea) AIA eA UR He ek: 981. A. Prilipkoanus Grossh. Calyx 9-14 mm long, the teeth 3-5 mm long; pods (18) 20-25 or 30-55 mm long, glabrate or sparsely covered with white hairs; bracts lanceolate or subulate, 3-5 mm long; corolla purple or whitish or azure with a dark purple keel; leaflets10-20pairs. 37. Standard 23-25 mm long; corolla purple throughout; style 5-7mm long; pods 3-5.5 cm long, the beak 4-5 mm long (M. Dnp.).... Ah reel bt Tatectes Sy, re hats inet sey Dae vy cae roe 574. A. monspessulanus L. Plants: differing fromabovess aside ele al see eee 43. Pods 38-70 mm long, or 20-30 mm and then not more than 2.57 3\mm broad, narrowly linear! sacsaont).Ol8 Saeiemeaee 39. Pods 20-30 mm long, 3-5 mm broad, broadly linear or lanceolate. wie Piste Le ia os BR a tian nly ey os pe Mebane 41. Leaves 1.5-3 cm long; leaflets 3-4 (6) mostly 4 pairs, oblong to oblong-lanceolate, acute, rarely subobtuse; pods 20-30 mm long, 2.5-3 mm broad; plants 3-8 cmtall.. 590. A. maculatus Bge. Leaves 3712 cm long; leaflets (5) 6-11 pairs, oblong to oblong- lanceolate acute or ovate to orbicular subobtuse; pods 4-7 cm long, 2.5-3 mm broad; plants 4-10 or 12-17 cmtall....... 40. Leaves 10-15 cm long; leaflets 6-7 pairs, oblong; pods 3.8-4.5cm long, 7 mm broad (Badkhyz). 591. A. Holdihianus Aitch. et Baker. Leaves 3-7 cm long; leaflets (5) 6-7 (8) pairs, mostly orbicular- ovate, subobtuse, 3-7mm long, 2-5 mm broad; peduncles 4-10 448 (12) cm long; calyx 10-12 mm long, the tube 3-5 times the length of the lanceolate teeth. ..... 592. A. jolderensis B. Fedtsch. + Leaves 8-12 cm long; leaflets (8) 9-11 pairs, oblong to oblong- lanceolate, acuminate or acute, 7-13 mm long, 2-4 mm broad; peduncles 12-17 cm long; calyx 14-16 mm long, the tube twice the length of the linear teeth...... 993. A. confiniorum Boriss. 41. Leaflets (4) 5-7 pairs, 6-18 (23) mm long, 5-10 mm broad; bracts 6-12 mm long. glabrous or sparsely ciliate; corolla yellow, persistent in fruit; pods falcate, glabrous at maturity (Care AT aired la a ae eo ae. 989. A. Cottonianus Aitch. et Baker + Leaflets 6-11 pairs, 2-3 mm long, 2-5 mm broad; bracts 1-2 mm long, covered with appressed white hairs; corolla yellow, becoming red; pods straight or slightly curved (Kopet Dagh).. . 2 ots | OM GS VOOR ET Ben hy (e RERRCE Ce eee ee ene ne eer ne aS oa ene Ta TE Seder EE 42. 42. Bracts lanceolate; calyx white-hairy, the tube 1.5-2 times or rarely 3-4 times as long as the teeth; stipules narrowly lanceolate; pods; ta opMimyDROAG), 4:2) je Se CHA MON Gr ie spec Ahion of ucla ie nictlls > Aoele qicueie 3 2 SO ied ly stacy py A ic HOE cI oS ace 987. A. submaculatus Boriss. + Bracts ovate; calyx predominantly black-hairy, the tube 5 times 601 the length of the teeth; stipules mostly ovate; pods 3.5 cm long, ZEON DROAC it ee kM ee 088. A. velatus Trautv. Asma) standard ca. 2 i nim ‘long. Roe Nhe) aR SN Se A RR ae, 44. + Standard aaa Omari ONO! sh eens er ah ema tom, NMS MenritL” rateraian taken 45. 44. Corolla whitish, with azure-violet-tipped keel; pods 30 mm long, curved, glabrous, red or spotted (Ciscaucasia). ........... Me ME ee ieee deere shin esac ts 576. A. Demetri Charadze. + Corolla dark purple, drying azure-violet; pods 18 (25) mm long, straight, tomentose with appressed white hairs. (Dagestan, Ineushenlvay ice toe eee Ll 077. A. Alexandri Charadze. 45. Corolla whitish or azure; pods sparsely pubescent or glabrate, somewhat curved (H..and's. iranscaucasia). 0. 5. «oe 3 fsa OI GS ese Ah ar la a gt ala 575. A. polygala Pall. - Corolla whitish-azure, with dark purple-tipped keel; pods tomentose with appressed white hairs; straight, spreading (central part of Greater Caucasus). 585. A. kazbeki Charadze. Series 1. HETEROPHYLLI Boriss. — Leaves consisting of a single leaflet or 3-11 (13) pairs, all the types often appearing on the same plant; leaflets mostly large, 1-7 cm long, 1-5.5 cm broad, ovate-oblong to orbicular; pods narrowly cylindric, 4-7 cm long, erect.— Iran, S. Transc. 567. A. Candolleanus Boiss. Diagn. ser. I, 2 (1843) 80; Boiss. Fl. or. II, 461; Bge. Astrag. geront. II, 203; Grossh. Fl. Kavk. II, 334. — A. supervisus Sheld. in Miness. Bot. Stud. I (1894) 140. — A. Latitolius, DC. Astrag. (1802).235,, pro parte, non, Lam.— Jc. ;, DC. Ll Sepitab).wiA6,.4.. (2. Perennial, from a woody vertical taproot, acaulescent, canescent by appressed hairs; stipules triangular, short, sericeous, adnate to petiole to above the middle; leaves mostly consisting of a single terminal leaflet, this ovate to ovate-orbicular, obtuse, mucronulate, subtruncate-based, commonly silvery appressed-tomentulose, the lower leaves sometimes consisting of 3-5 leaflets, the lowest leaflets 8-15 mm long, 6-18 mmbroad, the upper ones up to 4 cm long, the petiole mostly longer than or rarely 449 602 603 equaling the rachis, 1-2.5 cm long; peduncles 1-2, exceeding or rarely equaling the leaves; racemes 2-6-(or more)-flowered; flowers subsessile or borne on pedicels 1-3 mm long; calyx cylindric, 10-15 mm long, tomentose with appressed white and black hairs, the tube 2-5 times as long as the linear-lanceolate teeth; corolla yellow (according to Boissier, drying lilac), (18) 20-24 mm long; standard entire, obtuse, the limb broadly ovate, 8-12 mm broad, gradually cuneate-attenuate toward base; wing-petals about equaling the standard, the limb oblong, enlarged at apex, retuse on the inner side, obtuse, the basal auricle oblong, slightly curved, the claw 1.5 times as long as the limb; ovary linear, subsessile, sparsely hairy; style naked, 11-13 mm long; pods straight or slightly curved, narrowly linear-triquetrous, flattish, 3-5 cm long, appressed-hairy, 2-2.5 mm broad, dark-spotted, bilocular, many-seeded; seeds brown, oblong-reniform, ca. 3 mm long, 1 mm broad. Fl. April; fr. April-May. (Plate XXXIX, Figure 4). Dry stony slopes of mountains and hills. — Caucasus: S. Transc. (Nakhichevan. Erevan). Gen. distr.: Arm. Kurd. (vicinities of Ol'tin and Kagyzman). Iran. Described from Iran. Cotype in Leningrad. 568. A. latifolius Lam. Encyl. meth. I (1789) 319; DC. Prodr. II, 304; Pall. Astrag. 58; Bge. Astrag. geront. II, 204; Boiss. Fl. or. II, 462; Grossh. Fl. Kavk. II, 334.— A. choicus Bge. Astrag. geront. II (1869) 204 aac. DC ANStrace tabby.) 4 Gehl: Perennial, from a thick straight woody taproot, acaulescent, prostrate, canescent by appressed pubescence, the thick caudex covered with vestiges of stipules and petioles; stipules petiolar, appressed-hairy, triangular- ovate, 3-5 mm long; leaves 5-15 cm long, simple and consisting of a single leaflet, or 1-5 (7) pairs, both simple, mostly subradical and compound leaves, with varying number of leaflets occurring on the same plant, the petiole 4-5 cm long, appressed-hairy, somewhat enlarged toward base; leaflets appressed-puberulent on both sides, glaucescent or greenish, mucronate, mostly cuneate-based, ovate to ovate-oblong, 1-3.5 (5) cm long, 5-20 (35) mm broad; peduncles equaling or exceeding the leaves, appressed-hairy, 5-16 cm long, erect or recurved and spreading; raceme many-flowered; flowers distant, subsessile; bracts lanceolate, 3-4 mm long, hairy; calyx cylindric, 12-14 mm long, densely appressed-hairy, the tube 2-3 times as long as the linear-lanceolate or linear teeth; corolla yellow (according to De Candolle, .reddish in dry condition); standard 20-23 mm long, 6-10 mm broad, spatulate-oblong and drawn out in upper third, gradually cunate-attenuate toward base, acutish, rarely shallowly retuse; wing-petals 18-20 mm long, the limb spatulate-oblong, 2/3 as long as the filiform claw, round-tipped, emarginate on the inner side, the basal auricle oblong, acutish; keel somewhat exceeded by the wings, the inequilateral limb 2/3 as long as the claw, acutish; ovary linear, sessile, appressed-hairy; style ca. 8 mm long, filiform, naked; pods narrowly linear, 5-8 cm long, straight or somewhat curved, attenuate into an acute beak 2-4 mm long, covered with appressed hairs, not spotted, bilocular; seeds many, brown, oblong-reniform. Fl. May-June; fr. May-July. Clayey and stony mountain slopes and dry hills. — Caucasus: S. Transc. (Nakhichevan'). Gen. distr.: Arm.- Kurd. (Khoi province, Kul'py, Kagyzman vicinity, Bayazet), Iran. Described from Armenia and Iran. Type in Paris. 450 604 Note. Specimens (var. transcaucasicus Boriss.), collected in Kagyzman vicinity and in Yerevanvicinity (Kul'py), differ from the typical material in having a retuse standard. Specimens with terminal leaves consisting of 3 or 5 leaflets were collected in the Kulja area. 969. A. Fedorovii Takht. in Not. Syst. Bot. Tphil. XK (1941).67.—Ic.: ey tose. Acaulescent plants, vested with appressed gray pubescence, from a thick vertical tuberculate hard woody root-crown; leaves oblong in outine, 10-15 cm long, 5-6 cm broad, imparipinnate, leaflets 4-5 pairs, rnhomboid- ovate, acuminate and cuspidate, covered on both sides with silky gray pubescence, 1-2-3 cm long and 0.7-1 cm broad; stipules triangular, sericeous on the outside, purple; peduncles about equaling the leaves; racemes many-flowered, elongated; bracts linear-lanceolate, greatly exceeded by the calyx-tube, covered with white and black bristlelike hairs; calyx covered with appressed white and black nairs, the tube twice the length of the teeth; standard twice as long as the calyx; wings folded; pods linear, somewhat curved, at first densely pubescent, mucronate. Fl. May. Stony slopes and hills.— Caucasus: S. Transc. (Daralagez). Endemic. Described from the vicinity of the Karavank monastery. Type in Yerevan. Note. Resembling the preceding species and chiefly distinguished by the smaller leaflets. 570. A. heteromorphus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, from a very firm long woody curved or straight root, acaulescent, 8-30 cm tall, the thick short caudex clothed in vestiges of stipules and petioles remaining from previous years; stipules triangular- lanceolate, acute, sparselyhairy, 7-12 mm long, leaves 4-15 cm long, simple or composed of 3-11 (13) leaflets; leaf petioles 2-7 cm long, leaflets the fewer the larger, green, glabrous or sparsely covered, mainly beneath, with appressed hairs, mucronate, sometimes obtuse, broadly cuneate toward base, short-petioluled, rounded-ovate, 1-7 cm long, 1-5.5 cm broad; peduncles 8-30 cm long, much longer than the leaves, straight; raceme many-flowered, 6-20 cm long, the distant flowers borne on pedicels 1-2 mm long; bracts narrowly lanceolate, (5) 7-10 mm long, glabrous or sparingly ciliate-margined; bracts linear, glabrous, 2-8 mm long; calyx cylindric, 12-14 mm long, the tube 2-3 times as long as the very thinly appressed-hairy acute linear teeth; corolla yellow; standard 22-25 mm long, 8-12 mm broad, obtuse, the limb oblong, bluntly angular at base and much narrower than in upper part, the cuneate claw shorter than the limb by one-third; wing-petals 18-20 mm long, the spatulate-oblong limb as long as the filiform claw, emarginate on the inner side, round-tipped, the basal auricle oblong; keel 16-19 mm long, 3-4 mm broad at the broadest part, the inequilateral limb a trifle shorter than the claw; ovary linear, sessile, thinly appressed-hairy; style filiform, naked, 9-10 mm long; pods narrowly linear, 4-5.5 cm long, 2 mm broad, straight, gradually acuminate toward the short beak, covered with isolated appressed hairs, pale green, striate-spotted with brown, bilocular, many-seeded; seeds oblong-reniform, brown, ca. 2mm long, 1 mm broad. Fl. April-May; fr. May-June. Mountains. — Caucasus: S. Transc. (Nakhichevan', Chananab salt mines). Gen. distr.: Arm.- Kurd. (surroundings of Lake Urmiya). Described from the vicinity of Nakhichevan'. Type in Leningrad. 451 605 Note. Resembling A. latifolius Lam., from which it differs in the very thinly hairy or glabrate developed leaflets, calyx, and bracts, and their size, the shape of standard, and the glabrate and spotted pods. 571. A. Rollovii Grossh. im Beihefte zum Bot. Centralbl. XLIV (1927) PAN Perennial, from a woody root, cinereous by appressed hairs, 5-10 cm tall; stipules hyaline, triangular, acuminate, ciliate; leaflets 4-5 pairs, 6-12 mm long, ovate-oblong or ovate, acuminate; raceme somewhat overtopping the leaves, loosely 2-5-flowered; bracts small, somewhat longer than the pedicel, 1-2 mm long; calyx cylindric, the tube 8-10 times as long as the triangular obtusish teeth; corolla lurid, 18-22 mm long; standard oblong-ovate, retuse; pods cylindric, somewhat curved, 35-45mm long, terminating in a straight firm point 10-15 mm long. Fl. andfr. June. Stony slopes, at altitudes of about 1400 m. Possibly occurring in the Caucasus. Gen. distr.: Iran (N. Iran). Described from the vicinity of Tabriz. Type in Baku. Series 2. SILIQUIFORMES Boriss.— Pods cylindric, linear or lanceolate, erect, the beak rather long; raceme long, often loose or becoming so in fruit. Standard limb oblong, constricted in upper part. 572. A. rostratus C.A.M. Verz. Pfl. Kauk. (1831) 144; Ldb. Fl. Ross. I, 648; Beet Astrag i 1l) 1983" Boiss lssorsHl» 458sq,Grossh® Win Khayk ie BIO c Perennial, acaulescent, 5-20 cm tall, cinereous by appressed hairs; caudex short, branched, clothed in remnants of old stipules and petioles; stipules petiolar, adnate to petiole up to the middle, narrowly lanceolate, acuminate, densely covered with appressed white hairs, these interspersed with black; leaves 2-9 cm long, the petiole shorter than the rachis; rachises slender, straight or curved; leaflets 6-8 pairs, distant, linear to lance-linear, acute, 7-22 mm long, 1-3 mm broad, green and glabrous or thinly hairy or glabrate above, densely sericeous with gray appressed hairs; peduncles shorter than to somewhat exceeding the leaves, 4-20 cm long, rather thick; racemes 2-3 cm long, densely 6-13-flowered, elongating in fruit up to 6-8 cm, the rachis nodose-flexuous; bracts small, ovate or lanceolate or linear, 2-4 mm long, thinly covered with predominantly black hairs; pedicels ca. 4-5 mm long; calyx cylindric, covered with predominantly black hairs, ca. 10mm long, the teeth converging on one side, the tube 4-6 times as long as the teeth; ovary bright purple (according to Grossheim); standard 21-25 mm long, very slightly retuse, ca. 2 mm broad, abruptly enlarged in upper part to 7 mm, constricted below the middle, broadly cuneate toward the base; wing-petals ca. 17 mm long, the limb oblong, inequilateral, ca. 2 mm broad, emarginate on the inner side, scarcely retuse and round-tipped, the basal auricle straight, the filiform claw somewhat longer than the limb; keel 16 mm long, the limb inequilaterally oval; 6 mm long, 3 mm broad; ovary linear; style filiform, naked, 7 mm long, somewhat curved at the end, pods 5-9 cm long (including beak), 3-4 mm broad, 2 mm thick, linear- cylindric, straight or somewhat curved, appressed-hairy, slenderly subulate-acuminate, terminating in a straight beak 10-15 (20) mm long, bilocular, many-seeded, sessile; seeds 2.5 mm long, 2 mm broad, angularly and obliquely reniform, nearly black, minutely pitted, naked. Fl. May-June. (Plate XXXIX, Figure 1). SS ede? 452 606 609 @ Dry stony or gravelly, woodless and barren mountain slopes. — Caucasus: Tal. (Zuvant, Diabar, Mistau, Barnazar). Gen. distr.: Iran. Described from Talysh, rural Zuvant. Type in Leningrad. 573. A. zangelanus Grossh. in Izv. Azerb. fil. Akad. Nauk No. 1-2 (1939) 114. Perennial, cinerescent by appressed hairs; stipules short-lanceolate, obtuse, tomentose; leaflets 7-15 pairs, oblong-linear, acute, profusely hairy beneath, faintly so or glabrate above; peduncles mostly exceeding the leaves; raceme 6-12 mm long; calyx covered with appressed white and black hairs, the tube 2-3 times as long as the subulate teeth; pods covered with appressed white hairs, erect, slightly curved, cylindric, 5-6 cm long (excluding the beak), the slender beak 7-10 mm long. Fl. May-June. Exposed clayey sites and earth creeps in the foothill timber zone. — Caucasus: E. Transc. (Zangelan district). Endemic. Described from Zangelan district, Azerbaijan. Type in Baku. Note. Strikingly resembling A. rostratus C.A.M. and differing chiefly in the length of the beak of the pod. 574. A. monspessulanus L. Sp. pl. (1753) 761; Willd. Sp. pl. III (1800) 1314; DC. Prodr. Il,,304; Ldb. Fl. Ross. 1, 647, pro parte; Schmalh. Fl. I, 227, pro parte.— A. praecox Baumg. Enum. stirp. Transsilv. II (Hedie)i 362. AN dieclinatus ‘Salish. Prodr# (1796) 337.— Exs: Fl. exs. austrohungarica No. 408; Fl. Hung. exs. No. 678.—Ic.: Hegi, Illustr. Fl. Ni feswy in LesoseRehbe lenmvEl. germ. XXL, 158. Perennial, from a firm woody root, cinerescent, sparsely covered with appressed hairs, the branched caudex and underground shoots clothed in stipules and remnants of old leaves; stipules petiolar, lance-acuminate, appressed-hairy; leaves many, 10-20 cm long, short-petioled; leaflets 10-20 pairs, oval or oblong, subobtuse, glabrous above, appressed-hairy beneath; peduncles equaling or exceeding the leaves; raceme loose, oblong; bracts lanceolate, 3-5 mm long, exceeding the pedicel; calyx cylindric, 11-13 mm long, glabrous except for isolated white and black hairs, the filiform teeth 3-5 mm long; corolla purple, not becoming yellow in drying; standard 23-25 mm long, oblong, 6-7 mm broad, scarcely retuse; wing-petals 17-18 mm long, the limb oblong, retuse at apex and on the inner side, the basal auricle curved, angularly round, the claw about as long as the limb; keel 14-17 mm long, obtuse, the limb oval; ovary linear, covered with isolated hairs; style filiform, naked, 5-7 mm long; pods linear, 3-5.5 cm long, mostly arcuate, appressed~hairy, brown, long-acuminate to a subulate point ca. 4-5 mm long. Fl. April-May; fr. June-July. Stony calcareous soils. — European part: M. Dnp. (Kamenets~Podol'sk to Mogilev). Gen. distr.: S. Europe (Spain, France, Switzerland), N. Africa (Algeria). Described from Europe. Type in London. Note. A species strikingly resembling A. polygala Pall. 575. A. polygala Pall. Astrag. (1800), 110, excl. syn. — A. monspessulanus var. polygala DC. Prodr. II (1825) 304. — A. monspessulanus Ldb. Fl. Ross. I, 647, pro parte; Schmalh. Fl. 1, 282, pro parte; Grossh. Fl. Kavk. Il, 333.— A. sanguinolentus auct. non M.B.— A. somcheticus C. Koch. in Linnaea XV (1841) 720.—Ic.: Palin acy talon ore). 453 (7 . RY ra r a a OK ACS A) Da) da ae, PLATE XXXIX 4. A. Candolleanus Boiss. 454 ; ‘ AAI ; " a : WOU aN BS SS Astragalus rostratus C.A.M.—2. A. fragrans Willd.— 3. A. jolderensis B. Fedtsch. if Perennial, from a firm woody vertical taproot, green, usually thinly appressed-hairy; caudex branched, the divisions 2-5 cm long, covered with remnants of old petioles and stipules; stipules petiolar, connate to about the middle, ovate-lanceolate, 5-10 mm long, covered with appressed white hairs, the free portion acuminate; leaves many, 5-25 mm long, the rachis slender and curved, equaling (in large specimens) to 2-3 times as long as the petiole; leaflets (10) 12-20 pairs, mostly 12-15, ovate-oblong to oblong- oval, glabrous or covered with isolated hairs above, thinly appressed-hairy beneath, the young leaflets more profusely hairy, 4-13 mm long,.3-6 mm broad, commonly obtuse; peduncles equaling or exceeding the leaves, curved; raceme loose, rarely dense, short, ovaloid or oblong, mostly few-flowered; pedicels 2-5 mm long; bracts oblong-lanceolate or subulate, 2.5-4 mm long; bracteoles small, linear; calyx cylindric, ca. 11 mm long, thinly appressed-hairy, often merely covered with isolated hairs, predominantly on the teeth, the teeth subulate-filiform, 3-5 mm long; corolla white or pale azure, with a darker bluish-purple keel (purple according to Pallas), often yellowish in dry condition; standard oblong- lanceolate, ca. 24 mm long, 6-9 mm broad, retuse or sometimes nearly bilobate, gradually attenuate toward base; wing-petals 21 mm long, the limb lanceolate, 2-2.5 mm broad, unequally bilobate, the inner margin emarginate and convex, the basal auricle curved, the claw about as long as the limb; keel 18 mm long, the limb inequilaterally oval, ca. 4mm broad in upper part, the claw 1.5 times as long as the limb; ovary linear, glabrous or covered with isolated hairs; style naked, curved at the end, 10-11 mm long; pods linear-cylindric, straight or somewhat arcuately curved, 2.6 cm long, ca. 3.5 mm broad, 2 mm thick, terminating in a point 2-3 mm long, minutely spotted, brownish or yellowish, glabrous or covered with isolated hairs, large, somewhat inflated, obsoletely sutured ventrally, indistinctly grooved dorsally, bilocular; seeds dark brown, reniform, 1.5-3 mm long, ca. 1 mm broad. Fl. April-June; fr. May-July. (Plate XLI, Figure 2). Rocks, stony and argilaceous or calcareous slopes in the subalpine and alpine zones, at latitudes up to 2700 m.— Caucasus: S. Transc. Gen. distr. : Arm.- Kurd. (in vicinity of Artvin and of Kars). Described from the Caucasus. Type in London. 576. A. Demetri Charadze in Bull. Ac. Sc. Georg. SSRV, IIINo. 7 (1942) Asa newline lentws auct:, non. Bi A monspessulanus AUC EMOnmWer ea es mil ec. TO44f: Il. Perennial, from a thick woody much-divided root crown, green, thinly covered with appressed white hairs, the divisions clothed in remnants of old petioles and stipules; stipules triangular-lanceolate, acuminate, hairy; leaves (8) 10 (15) cm long; leaflets 25 pairs, ovate-elliptic to elliptic, 12-15 mm long, 3-6 mm broad, the lower thinly covered with appressed hairs; peduncles equaling or somewhat exceeding the leaves; bracts ca 4mm long, lanceolate, acute, membranous, somewhat exceeding the pedicel, thinly covered with appressed white hairs; calyx cylindric, thinly covered with appressed black hairs, ca. 9mm long, almost red, membranous, faintly purple in fruit, the subulate teeth ca. 3 mm long; corolla ca. 22 mm long, white, the keel azure-violet-tipped; standard 22 mm long, 5 mm broad, gradually attenuate toward base, shallowly retuse at apex; wing-petals 17 mm long, bilobate, the lobes short unequal; keel 16 mm long; pods straight, 30 mm long, 4 mm broad, curved, 455 611 scarcely grooved dorsally, somewhat inflated, carinate ventrally, purple or red-spotted, terminating in a long beak; seeds reniform, brown or nearly black, 2 (2.5) mm long. Fl. May; fr. July. Rock slopes.— Caucasus: W. Cisc. (Kislovodsk, Beshtau, Voroshilovsk, Grushevka). Endemic. Described from the vicinity of Kislovodsk. Type in Tbilisi. 577. A. Alexandri Charadze in Bull. Ac. Sc. Georg. SSR V, III, No.7 (1942) 700.—A. sanguinolentus auct., nonM.B.—A. muonspessulanus auct., nonL.— A. salatavicus auct. non Bge.—Ic.: l.c! 705, 8. IV. Perennial, cinereous, thinly covered with short appressed white hairs, from a thick woody root-crown with many divisions; stipules triangular- lanceolate, acuminate, hairy; leaves 8-12 (20) cm long; leaflets 12-15 pairs, ovate-elliptic to ovate, 6-8 (10) mm long, 3-4 (8) mm broad, the lower thinly appressed-hairy; peduncles shorter to longer than the leaves, 10-12 (15) cm long; racemes 4-6 cm long, few-flowered; bracts 4-6 mm long, membranous, lanceolate, acute, exceeding the pedicel, thinly covered with white hairs; pedicels ca. 21.5 mm long, dark purple, drying azure- violet; calyx cylindric, thinly covered with appressed white and black hairs, membranous enlarged in fruit, red, up to 12 mm long, the subulate teeth ca. 3mm long; standard 21 mm long, elliptic, retuse; wing-petals 18 mm long; keel 16 mm long; pods straight, 18 (25) mm long, 8 (4) mm broad, cylindric, curved on the back, subsulcate dorsally, long-mucronate, carinate ventrally, brown, slightly rugose, covered with appressed white hairs; seeds reniform, brown, 1.5 mm long. Fl. May; fr. June. Dry stony slopes in limestone-~dominated areas, in the intermediate mountain zone. — Caucasus: Dag., E. Transc. Endemic. Described from Gunib. Type in Tbilisi. 578. A. glaucophylloides Bornm. et Woron. in Monit. Jard. Bot. Tiflis XXXIV (1914) 2; Grossh. Fl. Kavk. II, 332. Perennial, from a firm woody root, acaulescent, cinerescent, the thickish branched caudex clothed in imbricated remnants of stipules and petioles; stipules broadly ovate to broadly lanceolate, acuminate, densely appressed-hairy; leaves many, subradical, 20-25 cm long, the petiole covered with scattered appressed hairs, 5-8 cm long; leaflets 10-12 pairs, glabrous above, very sparingly hairy beneath, oblong or ovate to orbicular, subdistant, 1.5-2 cm long, ca. 8 mm broad, obtuse or mucronulate, rounded-cuneate at base, borne on very short petiolules; peduncles covered with scattered appressed hairs, curved, ascending or procumbent, 10-22cm long, together with inflorescence not exceeding the leaves; racemes many~ flowered, 4-10 cm long, somewhat elongating in fruit; pedicels 4-5 mm long; flowers large, 30-35 mm long; bracts membranous or herbaceous, lanceolate, thinly hairy, 5-6 mm long; calyx 15-18 mm long, long~-cylindric, whitish or violet, thinly covered with appressed hairs, the subulate teeth converging on one side, half as long as the tube; corolla purplish-violet, drying violet-blue, twice the length of the calyx; standard (18) 27-35 mm long, the limb oblong, 5-7 mm broad, obtuse, scarcely retuse; wing-petals exceeding the keel, 20 mm long, ca. 3 mm broad, the basal auricle incurved; keel round-tipped, ca. 18 mm long, the limb oval curved, the auricle distinct straight; ovary linear, borne on a very short stipe. subsessile, glabrous; style ca. 8 mm long, naked; pods erect, narrowly linear, 4-4.5 cm long, ca. 4mm broad, beaked by an upright persistent 456 612 style, straight or scarcely curved, glabrous, dark brown or faintly spotted, the pointed beak 2-3 mm long. Fl. and fr. June. Dry soils and rocks.— Caucasus: E. Transec. (Meskhetiya, Akhaltsikh), S. Transc. (Karabakh). Gen. distr.: Arm.-Kurd. (former Kars region, Kagyzman vicinity, Ol'ty). Described from Kagyzman. Type in Leningrad. 579. A. montis-aquilis Grossh. in Izv. Azerb. fil. Akad. Nauk. I (1940) 34. Perennial, from a strong firm many-headed rootstock, appressed-hairy throughout except for the upper surface of leaflets, green; stipules small, triangular-lanceolate, tomentose; leaves many, rather small, 5-6 cm long; leaflets 5-7 pairs, small, 3-4 (5) mm long, broadly obovate, indistinctly retuse, glabrate above; peduncles equaling or very slightly exceeding the leaves; racemes 3-7-flowered; bracts lanceolate, half the length of the pedicel; calyx 10-12 mm long, coveredwith appressed predominantly black hairs, the tube 4-5 times as long as the black subulate teeth; corolla whitish, the keel reddish-violet-tipped; standard 2-2.5 times the length of the calyx, the limb oblong, retuse; wing-petals narrow, a trifle shorter than the standard; pods straight, cylindric, covered with appressed gray hairs, ca. 20 mm long, twice the length of the calyx. Fl. andfr. May. Crevices of calcareous rocks in the foothills, at altitudes of 500-800 m. — Caucasus: E andS. Transc. (Azerbaijan, Nakhichevan A.S.S.R.). Endemic. Described from the vicinity of Aznav''yurt, Kara Kum mountain spurs (Orlinaya mountain). Type in Baku. 580. A. cyri Fom. in Delect, sem. Horti Bot. Tifl. (1905) 7, nom. ; Grossh. Fl. Kavk. II, 332; Gosnovskii in Monit. Jard. bot. Tifl. XXXII, 10.—A. purpureus Fom. inTr. Tifl. Bot. sada, No. VI, 2 (1902) 2, non Lam. Perennial, acaulescent, pruinose, very thinly appressed-hairy, from a short divided caudex; stipules glabrous, triangular-lanceolate, petiolar; leaves many, subradical, 6 cm long; leaflets rather thick, 6-7 pairs, ovate-elliptic, 5-10 mm iong, obtuse, glabrous above, the few appressed white hairs on the lower surface confined to the midrib, scarcely retuse, mucronate; peduncles much shorter than the leaves, procumbent; racemes 3-5-flowered; bracts ovate-lanceolate, glabrous, sparingly cililate- margined; bracteoles 2; pedicels exceeding the bracts, covered with black and white hairs; calyx bright purple (merely red spots persisting indrying), (10) 12-15 mm long, glabrous or very thinly covered with black hairs, the tube 5-6 times as long as the lance-subulate black-hairy teeth; corolla bright purple, 20-25 mm long; standard more than twice the length of the calyx, ovate-oblong, retuse; wing-petals unequally bilobate at apex; pods faleate, narrowly linear, 4-4.5cm long, 4-5mm broad, compressed laterally, glabrous, lustrous, smooth, prolonged into a short slender beak, red- spotted, shallowly grooved dorsally, short-stipitate, the stipe equaling the breadth of the pod. Fl. March; fr. April. Solonetz soils, in gullies.— Caucasus: E. Transc. (Tiflis-Mtskhet, Kartli). Endemic. Described from the vicinity of Mtsket. Type in Tbilisi. 581. A. Prilipkoanus Grossh. in Tr. Bot. inst.Azerb. fil. Akad. Nauk II (1936) 254. Perennial, from a firm woody vertical root, vested with appressed gray hairs or green, cespitose by the many divisions of the caudex covered with remnants of old stipules and petioles; stipules petiolar, linear-lanceolate, 457 covered with appressed white hairs, long-acuminate, 6-13 mm long; leaves 10-14 cm long, the appressed-hairy petiole half as long as the rachis; leaflets 8-12 pairs broadly ovate to oblong-ovate, round-tipped and mucronulate, 5-20 mm long, 3-8 mm broad; peduncles straight, 15- (40)cm long, exceeding the leaves; bracts subulate, ca. 3 mm long, thinly hairy; pedicels short, 1-2 mm long or the flowers subsessile; calyx ca. 10 mm long, cylindric, covered with appressed black hairs, the teeth 1-2 mm long; corolla dingy violet; standard attenuate in upper third along 6 mm, 3-5 mm broad, enlarged about the middle to 9-10 mm, gradually attenuate toward base into a cuneate claw, scarcely retuse at apex; 22-24 mm long; wing-petals ca. 18 mm long, the limb oblong-lanceolate, ca. 10 mm long, 2mm broad, sharply retuse, the basal auricle oblong, the claw somewhat shorter than the limb; keel 13-15 mm long, the limb oval obtuse, somewhat curved, short-auriculate at base, the claw 1.5 times the length of the limb; ovary oblong, sessile, hairy, notably so on the ventral side; style ca. 10 mm long; pods straight, slightly curved, 20-25 mm long, twice as long as the calyx, covered with appressed white and black hairs, acutish, mucronate, grooved dorsally, the long style persisting for a long time. Fl. July. The intermediate mountain zone, on gravelly or clay-and-gravel slopes, and on pebbles in the beds of mountain streams.— Caucasus: S. Transc. (Nakhichevan A.S.S.R.; Leninakan, Mt. Alagez), Tal. (Mt. Alashi). Gen. distr.: Iran. Described from Bilyav River and Diza, Nakhichevan A.S.S.R. Type in Baku. Note. According to Grossheim, this species is most closely related to A. Wulfenii Kochand A. Spruneri Boiss. It differs from A. Wulfenii inthe pod which is 2/3 as long, and the smaller and dull- colored flowers; the characters distinguishing it from A. Spruneri Boiss. are the taller and more upright growth, the short subulate (not oblong) bracts which are twice the length of the pedicel, the duller color 614 of corolla, and the shape of the pod which is not verrucose-bristly. In flowering condition there is a resemblance to A. robustus Bge., the distinguishing characters being shape of pod and color of corolla. 582. A. czorochensis Charadze in Bull. Acad. Sc. Georg. SSR V. III No. 5 (1942) 462.— A. septemjugus Boriss. in sched. (1940).— AV eS an oun Ohemturs) \Woron. wn sched. non VEE sa Leis Ui lms lenC cme lites EOS gu CalC he xs uN OmcOllc Perennial, from a firm woody root, acaulescent, canescent throughout by appressed hairs, 5-10 cm tall; caudex clothed in remnants of stipules and petioles, the many divisions forming a compact mat; stipules scarcely adnate to petiole, 8-9 mm long, appressed-hairy, the free portion broadly lanceolate, 1-2 mm broad, acute or subacute; leaves 2-9 cm long, the slender curved rachis 2-3 times as long as the slender petiole; leaflets 7 pairs, rarely 5-6 or 8, distant, 3-4 (5) mm in circumference, mostly orbicular, rarely ovate, minutely mucronulate or scarcely retuse, glabrous green above, appressed-hairy beneath; peduncles equaling to very slightly exceeding the leaves, 5-10 cm long, mostly arcuate, appressed-hairy; racemes many-flowered; pedicels ca. 4mm long; bracts lanceolate or oblong, appressed-hairy, exceeded by the pedicel, 2-3 mm long; bracteoles at the base of calyx barely discernible, subulate; calyx 12-15 mm long, cylindric, obliquely truncate, sparsely covered with appressed white hairs, 458 615 the subulate unequal teeth 2-3 mm long; corolla purple, drying violet; standard ca. 24 mm long, spatulately oblong-lanceolate, 8 mm broad in the broadest upper third, truncate, obtuse or slightly retuse, gradually attenuate toward base into a narrowly cuneate claw; wing-petals 21-23 mm long, the limb 10 mm long, retuse and ca. 2 mm broad at apex, the basal auricle oblong reflexed, the claw somewhat longer than the limb; keel 20 mm long, the inequilateral oval limb 7-8 mm long, ca. 3 mm broad, the basal auricle short obtuse; ovary oblong-lanceolate, glabrous, borne on a stipe ca. 2.5 mm long; style 12-13 mm long, naked; pods flattish, lanceolate, carinate on one side and slightly grooved onthe other, spotted. sparsely covered with appressed white hairs, 16-21 mm long, 2.5-3 mm broad, terminating in a pointed beak 2-3 mm long, obtuse at base, the stipe short and slender; seeds dark, reniform, 2-2.5 mm long. Fl. April- May; fr. from the end of April. Mountain rocks.— Possibly growing in the USSR. Gen. distr.: Arm. - Kurd. (Chorokh River of lower Artvin; Ol'tin vicinity). Described from the vicinity of Artvin. Type in Tbilisi. 583. A. schizopterus Boiss. Diagn. ser. 1, 2 (1843) 82; Boiss. Fl. or. II, 459; Grossh. Fl. Kavk. II, 332.— A. cuspidatus Bge. Astrag. II (1869) 198. Perennial, nearly acaulescent, sparsely hairy, from a woody vertical taproot and a short caudex densely clothed in old stipules and petioles; stipules densely white-tomentose, lanceolate, acuminate; leaflets 7-15 pairs, rounded-ovate to ovate, obtuse or minutely retuse, glaucous, covered sparsely above and more densely beneath with scattered appressed white hairs or glabrate, (3) 5-10 mm long, (2) 3-8 mm broad; peduncles shorter than to equaling the leaves, thickish, curved; raceme loosely few-flowered; bracts lanceolate, acute, exceeding the pedicel, 4-5 mm long; bracteoles half as long as the bracts; calyx cylindric, sparsely covered with short appressed white and black hairs, whitish or greenish, 15-18 mm long, the long subulate teeth half as long as the tube; corolla 25-33 mm long, purple; standard twice the length of the calyx 25-33 mm long, ca. 9 mm broad at the middle, the elongated lance-oblong limb cuneately attenuate at both ends, retusSe at apex; wing-petals unequally bilobate or almost entire, greatly exceeded by the standard, somewhat longer than the keel, ca. 23mm long, the limb elongate-oblong, ca. 2 mm broad, about as long as the claw, acutish, the basal auricle ovate; keel 20 mm long, the limb inequilaterally oval, round-tipped, 3 mm broad, the basal auricle short, the claw 1.5 times as long as the limb; ovary oblong-lanceolate, stipitate, hairy; style ca. 12 mm long, glabrous, nearly straight; pods 2-2.5 cm long, 4-5 mm broad, slightly curved or straight, compressed laterally, obtusish at apex and terminating in a slender beak 5-8 mm long, scarcely grooved dorsally, convexly carinate at base, sparsely hairy, spotted with purple. Dry rocky soils, over limestone.— Caucasus: Possibly growing in Transcaucasia. Gen. distr.: Arm.- Kurd. (Ol'tin vicinity). Described from Kilik. Type in Geneva. 584. A. sanguinolentus M.B. Tabl. prov. Casp. (1798) 117; M.B. Beschr. Land. zw. Terek und Kura (1800) 190; M.B. Fl. taur.-cauc. II, ZOUmiNe 497 Waillds: Spa voli 2 sel gid AN. Vierz. (btlz > eauc. Ces) 4 pCa Prodr: my 304; idb: PilssRoss. 1,648; sBge, Astrag. geront. II, 200; Boiss. Fl. or. II, 459; Grossh. Fl. Kavk. II (1930) 333.— 459 616 617 A. haematocarpus Bge. Astrag. geront. II (1869) 201; Boiss. Fl. or. II, 460; Grossh. Fl. Kavk. II. 333.— A. Kiapasi Woron. nom.— Kiapasia Schelkovnikovi Woronov in sched.; Grossh. in Izv. Azerb. Fil. AN 1-2 (1939) 115, I (1940) 35.—Ic : Pall. Astrag. (1800) tab. 84 (sub! Av ineanio);>sinpBulley) Ac.«Sc# Georgy ISSR Vil Nowa. (1942) aeaavile Perennial, from a firm woody taproot, nearly acaulescent, mostly 5-10 (15) em long, the caudex clothed in old stipules and petioles, the many divisions forming a compact mat; stipules lanceolate, acuminate, 3-5 mm long, tightly imbricated, compact, appressed-hairy; leaves 1-5 cm long, the rachis slender, the petiole short or the leaves subsessile; leaflets small, 1-3 (5) mm long, 1-2 (3) mm broad, 10-15 pairs, elliptic to ovate, obtuse, sometimes minutely retuse, rarely subacute, green, glabrous above, more or less tomentose beneath; peduncles exceeding the leaves, often procumbent, 5-15 cm long, appressed-hairy; racemes few-flowered, short-pediceled; bracts subulate, barely exceeding the pedicel; calyx cylindric, 10 (13) mm long, covered with appressed white and black hairs, the tube 3 times the length of the short subulate teeth; corolla violet or ochroleucous with a dark violet blotch on the keel, the color varying sometimes on the same plant and even on the same flower, emarginate, more than twice as long as the calyx, 20-26 mm long, the limb oblong, 8-10 mm broad, retuse, angular at base, 3 times the length of the short cuneate keel; wing-petals somewhat exceeded by the standard, white, broad at apex, commonly retuse, 10-20 mm long, the limb oblong-lanceolate, about as long as the claw, the basal auricle short; keel exceeded by the wings, 13-16 mm long, obtuse, white, dark violet-blotched, the claw 1.5 times the length of the oval limb; ovary linear, subsessile, sometimes short-stipitate, sparsely appressed-hairy or glabrate; style 8-10 mm long, naked; pods suberect, oblong or broadly lanceolate, (1.5) 2.5-3 cm long, 4-6 mm broad, flattened, thickly sutured, carinate, straight or slightly curved, terminating in a short rigid point, spotted, the spots often confluent dark red, bilocular many-seeded. Fl. May-July. (Plate XLI, Figure 4). Mountains, at altitudes of 1300-3150 m, on gypsiferous clays, on dry rocky slopes of the subalpine and alpine zones, often among Cobresia associations. — Caucasus: Dag., E. Transc. (Kuba, Shemakha, Nukha), S. Transc. (rare). Endemic. Described from the vicinity of Kurt-Bulaga (between Shemakha and Kuba, near Alty-agach village). Type in Leningrad. 585. A. kazbeki Charadze in Bull. Ac. Sec. Georg. SSR. V, III, No. 7 (1942) 702.—A. sanguinolentus auct., nonM.B.—A. monspessulanus auctoainon Wea ale co leniii0o ae iNe: Perennial, greenish, sparsely covered in upper part with appressed white hairs, from a thick many-headed woody root; stipules lanceolate, hairy, acuminate; leaves 12 cm long; leaflets 12 pairs, 8-12 mm long, 4-6 mm broad, ovate or oblong-elliptic, round-tipped, covered with appressed white hairs; peduncles a trifle shorter than to exceeding the leaves; racemes short, few-flowered; bracts 4-5 mm long, narrowly lanceolate, acute, herbaceous, sparsely covered with appressed black hairs, barely exceeding the pedicel; flowers 25.5 mm long, whitish-azure, the keel dark violet-tipped; calyx cylindric, 10-12 mm long, covered with appressed black hairs, the subulate teeth 3-4 mm long; standard 25 mm long, deeply notched, bilobate; wing petals 21 mm long; keel 17.5 mm long; pods 25-30 mm long, 4mm broad, straight, spreading, carinate on both 460 sides, falcate, covered with appressed white hairs, attenuate at apex to a point; seeds brown, 2 mm long, reniform. July. High-mountain zone, on rocky slopes.— Caucasus: W. Transc., Cisc. (The main range). Endemic. Described from Kazbek. Type in Tbilisi. Note. Var. megrelicus Charadze, collected on Megrelii limestone, differs from the type in the smaller pod size and details of flower structure. 086. A.salatavicus Bge. Astrag. geront. II (1869) 201; Boiss. Fl. or. II, 460; Grossh. Fl. Kavk. II, 333. Perennial, green, acaulescent, becoming lignified in lower part, from a short, thick, strongly branched caudex clothed in old stipules and petioles, cespitose; stipules lanceolate, herbaceous, imbricated, appressed-hairy on the outside; leaves 3-10 cm long; leaflets 9-12 pairs, glabrous above, obovate-elliptic to orbicular, approximate, 5-7 mm long, 3-4 mm broad; peduncles ascending, equaling or somewhat exceeding the leaves; racemes short, narrowly linear, 3-7-flowered; bracts lanceolate, exceeding the pedicel; calyx covered with appressed black and white hairs, the tube 3 times as long as the subulate teeth; corolla twice the length of the calyx, whitish; standard 22-24 mm long, the limb oblong, angular at base; wing-petals gibbous at apex, subequally and obtusely short-bilobate, ca. 20 mm long, the claw 1.5 times as long as the oblong limb; keel obtuse, 15-17 mm long, the claw 1.5 times as long as the oval limb; ovary 14-17-ovuled, oblong, hairy; style naked, ca. 12 mm long; pods oblong- ovaloid, 10-15 mm long, ca. 5 mm broad, straight, rather firm, flattish, bluntly triquetrous, carinate ventrally, slightly grooved dorsally, rounded at the top and abruptly contracted into a thick straight beak 2-3 mm long, covered with scattered appressed white hairs, brownish, cross-wrinkled, bilocular; seeds small, olivaceous, ca. 2 mm long, 1 mm broad, reniform. July. In the alpine zone of mounvains, on calcareous rocks, at the altitude of ca. 2500 m.— Caucasus: Dag. (Khanakoitau, Salataviya). Endemic. Described from Khanakoitau. Type in Leningrad. Series 3. VELATI Boriss.— Corolla persistent in fruit, yellow, sometimes fading and drying red; limb of standard broadly subquadrangular- ovate; pods linear or lanceolate. 618 087. A. submaculatus Boriss. in Not, Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, silvery gray by appressed white hairs, 6-20 cm tall, from a long woody taproot, the strongly divided caudex clothed in remnants of stipules and petioles; stipules petiolar, small, narrowly lanceolate, 2-3 mm long, long-acuminate; leaves 5-10 cm long, the petiole about as long as the rachis; leaflets 6-11 pairs, commonly distant, mostly orbicular, rarely ovate, 2-6 mm long, 2-5 mm broad, short-acute or obtuse; peduncles 6-20 cm long, mostly curved, slender, covered with appressed white and sometimes black hairs, 3-6-flowered; pedicels 3-5 mm long; bracts lanceolate, acute, somewhat exceeding the pedicel, appressed-hairy; bracts small, linear; calyx cylindric, 12-17 (20) mm long, 3-4 mm broad, appressed-hairy, the tube 1.5-2 (rarely 3-4) times as long as the lanceolate teeth; corolla yellow, at length becoming red; standard 20-24 mm long, arcuate, the limb ovate, angular at base, round-tipped and scarcely retuse, 10-11 mm broad, limb twice the length of the cuneate claw; wing-petals equaling the standard, the limb spatulate, 3-5 mm broad 461 619 in upper part, emarginate on the inner side and at apex; keel 18-20 mm long, the claw 1.5 times as long as the oval limb; ovary linear, stipitate, appressed-hairy; style hairy in lower third, pods straight or gently curved, 2.5-3 cm long, 4-5 mm broad, straight-beaked, covered with short appressed white hairs, reddish, bilocular; seeds brown, angularly reniform, ca. 2 mm long, 1.5 mm broad, smooth, short-funicled. Fl. April-June; fr. May-July. Stony and gravelly mountain slopes in the steppe region, among junipers, at altitudes of (700) 850-1950 m.— Centr. Asia: Mtn. Turkm. (centr. part of the Kopet Dagh range). Gen. distr.: ? Iran. Described from the Kopet Dagh range. Type in Leningrad. 588. A. velatus Trautv. in A.H.P. IX, 2 (1886) 450. Perennial, subacaulescent, appressed-hairy, (7) 10-20 (30) cm tall, the woody taproot ‘penetrating deep into rock fissures, the short branched caudex clothed in old stipules and petioles; stipules small, mostly ovate, rarely linear-lanceolate, herbaceous, greenish, covered on the outside with appressed white hairs, adnate to petiole at base; leaves all subradical or nearly so, together with petiole 3-9 cm long; leaf petioles and rachises beset with: appressed white hairs, firm at first, at length weaker; leaflets (8) 9-11 pairs, orbicular to broadly elliptic, rarely obovate, round-tipped, faintly mucronate or retuse, silvery, covered sparsely above and densely beneath with appressed white hairs, 2-5 mm long, 2-5 mm broad; scapes straight, elongating in fruit up to 30 cm, 10-11 cm long in flowers, covered with appressed white hairs; [bracts ?] 1-2 mm long; pedicels covered with appressed black hairs, up to 5 mm long; calyx 12-15 mm long, long- cylindric, gibbous at base, subtended by 2 barely discernible linear bracteoles, covered with short appressed black and fewer white hairs, the linear teeth 2-4 mm long; corolla 20-25 mm long, drying red, exceeding the calyx; standard 20-25 mm long, the limb broadly ovate to oblong, 8-10 mm broad, the limb acute, broadly cuneate toward base; wing-petals 18-21 mm long, curved, the limb oblong-spatulate, round-tipped, about as long as the claw, 3-5 mm broad in upper part, 2-3 mm below, the basal auricle curved gibbous; keel equaling the wings, the claw twice the length of the oval acutish limb; ovary linear, stipitate, thinly appressed-hairy; style 10-11 mm long, curved at the end, naked; pods 2-2.5 cm long, 3.5 mm thick, with persistent corolla, firm, smooth, divergent, linear, twice as long as the calyx, straight, more or less distinctly grooved dorsally, carinate ventrally, bilocular, sparsely covered with appressed white hairs, short-stipitate, the beak erect, very short. Fl. April-May, fr. April-June. Among stones, on gravelly and stony slopes, at altitudes of 600-1600m.— Centr. Asia: Mtn. Turkm. (Bolshie Balkhany range, western part of Kopet Dagh range). Endemic. Described from the vicinity of Kyzyl Arvat. Type in Leningrad. 589. A. Cottonianus Aitch. et Baker in Trans. Linn. Soc. ser. 2, III (1886) 51.—A. involucratus .Lipsky in,A.H.P. XVII, 1 (1900) 4655, A. Dilankuri. Lipsky in AJHYPs) xxvii) 2h(0907 1909) Ia perez V, 431.—Chondrocarpus Dilankuri Nevski in Acta Inst. Bot. Acad. Se. ser. 1, 4 (1937) 254. Perennial, acaulescent, green, at first sparsely appressed-hairy, at length glabrous, 14-30 cm tall, from a long thick firm fusiform root and 462 620 a short branched caudex imbricately clothed in old stipules; stipules dimorphic, the upper lance-ovate, the lower ovate, covered with soft white hairs, 5-8 mm long, the free portion long-acuminate, lanceolate; leaves 5-20 cm long, ascending, sometimes prostrate, equaling or exceeding the peduncles, the petiole finely sulcate, covered with appressed white hairs, 5-7.5 cm long; leaflets (4) 5-7 pairs, suborbicular to broadly obovate, obtuse or retuse, mostly mucronulate, 6-18 (23)mm long, 5-10mm broad, short-petioluled, covered beneath with appressed gray bicuspidate hairs, glabrate or covered with scattered appressed hairs on the greener upper face; peduncles straight, erect, thick, 5-30 cm long, glabrate; inflorescence (5) 10-20-flowered, the flowers at length distant; bracts oblong-lanceolate to linear, herbaceous, about twice the length of the pedicel, acute or acuminate, 6-12 mm long, glabrous or sparingly ciliate- margined; pedicels 3-5 mm long; calyx cylindric, glabrous, subtended by 2 linear bracteoles, 3-5 mm long, the tube 2-3 times as long as the subulate teeth, the margin often fringed with isolated black hairs; corolla yellow, persistent in fruit, ca. 2.5 cm long; standard large, up to twice the length of the calyx, ovate, the limb broad, oval-oblong, 8-14 mm broad, scarcely retuse, gradually cuneate-attenuate into and 2.5-3 times as long as the claw; standard orbicular in bud; wing-petals 20-21 mm long, the limb spatulate, emarginate on the inner side, 9-10 mm long, 2-4 mm broad in upper part, attenuate and auriculate at base, about as long as the claw; keel equaling the wings, the limb oval, curved, 4 mm broad, about as long as the claw; ovary glabrous; style naked, curved, ca. 10 mm long; pods short-stipitate, 20-30 mm long, 3-4 mm broad, broadly linear, subfalcate, compressed laterally, subtriquetrous, firm, rigidly mucronulate, carinate dorsally, grooved ventrally, glabrous or sparsely hairy in maturity especiallynear the groove, bilocular, dehiscent, the valves sometimes separating; seeds 4-5, angularly reniform, ca. 3 mm long, brown. Fl. April; fr. June. Outcrops of red clay and sandstone on mountain slopes, at altitudes of 700-1450 m.— Centr. Asia: Pam.-Al. (Zeravshan range. Baisun, Kugitang, Babatag range); Mtn. Turkm. (Badkhyz). Endemic. Described from Badkhyz. Type in Leningrad. Note. A. Cottonianus Aitch. et Baker was described from Badkhyz in 1886, but was erroneously referred by the authors to the section Myobroma, as they had overlooked the bicuspidate hairs. The mistake was repeated by a number of authors (Lipsky, Fedchenko). In 1900 Lipsky described A. involucratus from Baisun, which does not differ in any respect from specimens identified by Lipsky himself as A. Cottonianus but originating from the Zeravshan Range. Lipsky described this astragalus again in 1907, from plants collected at an earlier stage of flowering, under the name A. Dilankuri. A thorough examination of the latter discloses the stronger long and thick peduncles of preceding years. The shape and size of the standard and other flower parts was naturally somewhat different. The specimens described by Lipsky under the name A. involucratus were collected in fruiting condition and are not therefore readily comparable with the specimens obtained on the earlier occasion. Series 4. ECHINOCARPI Boriss.—Corolla yellow, sometimes drying red; limb of standard broadly ovate-quadrangular; pods narrowly linear, auricular, somewhat curved. 463 621 622 590. A. maculatus Bge. Astrag. gerontog. II (1869) 206. Perennial, acaulescent, 2-8 cm tall, silvery-gray throughout by appressed-white hairs; root long, woody, usually single, strong, rarely many roots, slender; caudex very short, branched with many divisions, clothed in remnants of old stipules and petioles; stipules triangular- lanceolate, firm, often tinged with red or pink, acuminate, appressed- hairy, 3-4 mm long; leaves 1.5-3 cm long, the petiole somewhat shorter than to about as long as the rachis; leaflets 3-4 (6), mostly 4 pairs, oblong or oblong-ovate to ovate, acute or acuminate, rarely triangular, commonly 4-5 mm long and 2-3 mm broad, appressed-hairy on both sides; peduncles slender, 3-8 cm long, equaling or slightly exceeding the leaves, appressed- hairy; raceme 2-4-flowered; pedicels ca. 2mm long; bracts oblong, acute to subobtuse, somewhat exceeding the pedicel; calyx cylindric, 10-12 mm long, ca. 3 mm broad, covered with appressed white and black hairs, the tube 3 times as long as the lanceolate teeth; corolla yellow, becoming red; standard 18-20 (23) mm long, 7-8 (10) mm broad, arcuately recurved, ovate, rounded and faintly retuse at apex, angular at base, 3 times as long as the broadly cuneate claw; wing-petals 17-19 mm long, the limb obovate, notched at apex and on the inner side, ca. 4 mm broad in upper part, ca. 2mminlower part, recurved-auriculate atbase, aslongas the claw; keel about equaling the wings, the claw 1.5 times as long as the oval 3 mm broad limb; ovary stipitate, linear, appressed-hairy; style naked, 6-7 mm long; pods linear, somewhat curved, ca. 2-3 cm long, 2.5-3 mm broad, attenuate into a short beak, covered with appressed white hairs, brownish or yellowish, bilocular; seeds elongate, ca. 4mm long and 1.5mm broad, quadrangular-reniform, smooth, borne on a placenta ca. 1 mm long, olivaceous. Fl. May-August; fr. July-September. Gravelly tablelands and slopes, and stony sites up to the alpine zone and snowfields, at altitudes of 1600-2600 m.— Centr. Asia: Mtn. Turkm. (centr. part of the Kopet Dagh Range). Gen. distr.: Iran. Described from E. Iran, between Nichapur and Meshkhed. Type in Leningrad. 591. A. Holdichianus Aitch. et Baker in Trans. Linn. Soc. ser. 2, II, (1886) 54. Perennial, acaulescent, clothed at base in persistent old petioles, from a long firm root; stipules ovate, acuminate, adnate at base to petiole; leaves densely covered with appressed white hairs, the petiole 2.5-5cm long, the rachis 7.5-10 cm long; leaflets 6-7 pairs, short-petioluled, acute or obtuse, oblong, 1-1.5 cm long; peduncles 5-15 mm long; racemes broad, 3-9-flowered; pedicels 4-5 mm long; bracts lanceolate, persistent; calyx ca. 17 mm long. covered with short appressed predominantly black hairs, the oblong tube twice the length of the lanceolate teeth; corolla 23 mm long, yellow; standard glabrous, somewhat exceeding the other petals; pods 3.8-4.5 cm long, 7 mm broad, linear, compressed laterally, curved, covered with short hairs, black-spotted, bilocular. April-May. Usually in sandy soils.— Centr. Asia: Mtn. Turkm. (Badkhyz). Gen. distr.: Iran (probably growing in Afghanistan). Described from Badkhyz. Cotype in Leningrad. 592. A. jolderensis B. Fedtsch. in. Not. Syst. Herb. Horti Bot. Petrop. II, 13 (1921) 52.— A. subalpinus Freyn in Bull. Herb. Boiss. sér. 2 (1905) 563, non Boiss. et Buhse. 464 623 Perennial, acaulescent, silvery by appressed hairs, 4-10 (12) cm tall, fromalong vertical woody taproot; caudex covered with crowded leaves and stipules, the many divisions forming a compact mat; stipules petiolar, ca. 3mm broad, lanceolate, white-hairy, imbricated, persistent in age; leaves many, crowded at base, 3-7 cm long, the grayish-silvery petiole somewhat shorter than the rachis; leaflets (5) 6-7 (8) pairs, ovate, mostly rounded-ovate to orbicular, subobtuse to obtuse, 3-7 mm long, 2-5 mm broad; scapes suberect to assurgent, 4-10 (12) cm long, appressed-hairy, covered in upper part with predominantly black hairs; racemes 2-5-flowered, the peduncles 1-2 mm long, together with calyx covered with black hairs; calyx cylindric, 10-22 mm long, the tube 3-5 times as long as the lanceolate teeth; corolla 22-26 mm long, yellow, the standard lilac; limb of standard ovate-oblong, (8) 10 mm broad, round-tipped or slightly retuse, distinctly auriculate at base, 3.5 times the length of the cuneate claw; wing-petals yellowish, 21 mm long, the limb spatulate, enlarged in upper part, 4-5 mm broad, 2 mm below, shallowly notched, the basal auricle oblong, the claw shorter than the limb; keel equaling the wings, incurved to a porrect apex, the claw about 1.5 times as long as the limb; ovary subsessile to short-stipitate, minutely and finely tomentulose, linear-filiform, ca. 9-10 mm long; style naked, ca. 10 mm long, somewhat curved at the end; pods narrowly linear, cylindric, mucronulate, 4-6 (7) mm long, 3 mm broad, flattened, bilocular, reddish, covered with appressed white hairs; seeds many, 2 mm long, ca. 1 mm broad, olivaceous. Fl. April-May; fr. May-July. (Plate XXXIX, Figure 3). Gravelly slopes and hills, on marle outcrops, at altitudes up to 1600 m.— Centr. Asia: Mtn. Turkm. (western part of Kopet Dagh Range). Gen. distr. : ? Iran. Described from loldere in vicinity of Kara-Kala. Type in Leningrad. 593. A. confiniorum Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent, (8) 12-17 cm tall, silvery by appressed hairs, from a dark brown woody contorted root, the much divided caudex clothed in old stipules and petioles; stipules narrowly lanceolate, long-acuminate, covered with appressed white hairs, often light brown, ca. 8 mm long; leaves 8-12 cm long, the petiole equaling the rachis; leaflets (8) 9-11 pairs, oblong to oblong lanceolate, rarely ovate, commonly acuminate, rarely acute, 7-13mm long, 2-4mmbroad, sericeous onboth sides with appressed hairs; peduncles rather slender, curved, equaling or slightly exceeding the leaves, covered with appressed black and white hairs; pedicels short, covered with predominantly black hairs; racemes loosely 5-8-flowered; bracts narrowly lanceolate, 4-6 mm long, appressed-hairy; bractoles filiform, ca. 2 mm long; calyx long-cylindric, 14-16 mm long, covered with appressed mostly white and sometimes black hairs, the tube twice as long as the linear teeth; corolla yellow, becoming red; standard 20-27 mm long, 8-10 mm broad, strongly curved, the limb ovate-orbicular to quadrangular, 3 times the length of the cuneate claw; wing-petals 19-24 mm long, the limb oblong-spatulate, emarginate on the side, the claw 1.5 times as long as the limb; keel equaling the wings, the claw 1.5 times as long as the inequilaterally ovate limb; ovary linear, short- stipitate, appressed-hairy; style 9-10 mm long, the lower third appressed- hairy; pods linear, 4-7 cm long, ca. 2.5 mm thick, acuminate to a beak 465 624 ca. 3mm long, brown, appressed-hairy, bilocular; seeds angularly reniform, brown, ca. 2 mm long, 1.5 mm broad, the naked placenta 1.5 mm long. Fl. May-June; fr. May-June. Mountain slopes.— Centr. Asia: Mtn. Turkm. (Mt. Massinev in Central Kopet Dagh). Gen. distr.: Iran (Astrabad Province). Described from Kopet Dagh Range. Type in Leningrad. Series 5. LIGNESCENTES Boriss.— Plants lignified at base, with persistent firm erect petioles; pods partlycernuous anderect; inflorescence a few-flowered raceme, the flowers distant. 594. A. ordubadensis Grossh. im Beih. zum Bot. Centralbl. XLIV Abt. Il (1927) 222; Grossh. Fl. Kavk. II, “339. Perennial, acaulescent, 10-20 cm tall, from a vertical woody taproot; caudex much divided, 2-6 cm long, clothed in old stipules and persistent petioles; stipules petiolar, lanceolate, connate up to the middle, covered with appressed white hairs, ca. 5mm long; leaves 10-16 cm long, the petiolesfirm, persistent, virgate, straight or slightly curved; leaflets 5-8 pairs, distant, small, 2-10 mm long, 1.5-3 mm broad, oblong~ lanceolate to elliptic, sometimes ovate, attenuate toward base, obtuse, rarely acute, greenish or grayish, covered with appressed white hairs; peduncles equaling or rarely somewhat exceeding the leaves; raceme remotely 3-4-flowered; bracts ovate-lanceolate, 1-2 mm long, about equaling the pedicel; calyx cylindric, covered with white and black hairs, the unequal teeth converging on one side, 1-1.5 mm long; corolla pale violet, 18-20 mm long; standard 18-20 mm long, ca. 6 mm broad in upper part, round-tipped and obtuse, the oblong limb minutely notched at base and minutely auriculate, 3 times as long as the cuneate claw; wing-petals ca. 16 mm long, the limb about as long as the claw, notched on the inner side, nearly round at apex, the reflexed auricle ca. 1 mm long; keel 15 mm long, the inequilaterally oval acutish limb 2/3 the length of the claw; ovary borne on a stipe ca. 3 mm long, linear, covered with short appressed hairs; style 9 mm long, naked; pods partly cernuous and partly erect, short~stipitate, spreading, linear-cylindric, 25-35 mm long, 2-2.5 mm broad, 2 mm thick, sparsely white-hairy, shallowly grooved dorsally, red-spotted, very narrowly carinate, the attenuate beak ca. 2 mm long; seeds ca. 2mm long, 1 mm-broad, olivaceous, angularly reniform. Fl. May; fr. May-June. Limestone on mountain slopes, at the altitude of about 800 m.— Caucasus: S. Transc. (Nakhichevan, Ordubad, Migry). Gen. distr.: ? Iran. Described from Ordubad. Type in Tbilisi; cotype in Leningrad. Note. Grossheim's description of A. ordubadensis does not entirely agree with the material at our disposal. According to Grossheim, this is a shrub up to 0.5 m tall, with oblong-linear or elliptic leaflets 4-6 mm long; peduncles bearing 2-5 flowers; calyx-tube 4-6 times as long as the teeth; retuse corolla 20-23 mm long; pods 32-35 mm long, 3-4 mm broad. 595. A. fuliginosus Beck. ex Stapf in Denkschr. Acad. Wien. II (1886) 2k Perennial, nearly acaulescent, 5-15 cm tall, from a vertical woody taproot, the caudex much divided, 1-2 cm long, densely clothed in old stipules and persistent petioles; stipules petiolar, lance-linear, covered 466 627 with appressed white hairs, ca. 5mm long; leaf petioles peristent, virgate, slender, curved, 3-10 cm long; leaflets distant, spreading, 8-12 pairs, linear, 3-12 mm long, 1-1.5 mm broad, covered with appressed silvery hairs; peduncles about equaling or slightly exceeding the leaves; racemes remotely 7-9-flowered; bracts small, ca. 1.5 mm long, oblong-lanceolate; pedicels ca. 2 mm long; calyx cylindric, ca. 10 mm long, covered with appressed predominantly black hairs, the subulate teeth 2-3 mm long, converging on one side; corolla purple; standard ca. 17 mm long, 8 mm broad at the middle, abruptly attenuate in upper part to 2 mm, broadly cuneate toward base; wing-petals 14 mm long, the oblong limb 2 mm broad at base; the basal auricle recurved elongate, the claw about as long as the limb; keel ca. 11 mm long, the triangular-oval inequilateral limb as long as the claw; ovary borne on a stipe 0.5-1 mm long, covered with short appressed hairs; style 7-9 mm long, sparsely hairy in lower part, glabrous elsewhere; pods sparsely spotted, cernuous or erect, 17-23 mm long, ca. 3 mm broad, carinate, sparsely covered on one side with appressed white hairs, the straight beak ca. 3 mm long; seeds ca. 1 mm long and broad, brown. Fl. May; fr. May. Mountain slopes, at the altitude of 300-400 m.— Caucasus: S. Transc. (vicinity of Ordubad). Gen. distr.: Iran (N. Iran). Described from N. Iran. Type in Vienna. Note. The form collected in the Obdurbad area of the Caucasus is not quite typical, being intermediate between A. fuliginosus Beck. and A. ordubadensis Grossh. As regards number of leaflet pairs, shape of leaflets and corolla, and number of flowers in inflorescence, it resembles A. fuliginosus; in dominant shape of leaflets it approaches A. ordubadensis Grossh. Series 6. REFRACTI Boriss.— Pods pendulous, sometimes partly spreading horizontally, oblong-lanceolate to linear, straight or curved, often spotted. 596. A. refractus C.A.M. Verz. Pfl. Cauc. (1831) 144; Ldb. Fl. Ross. I, 648; Bge. Astrag. geront. II, 207; Boiss. Fl. or. II, 464; Grossh. Fl. Kavi, IT, 335. Perennial, acaulescent, 20-35 (40) cm tall, canescent by appressed hairs, from a vertical taproot; stipules petiolar, adnate to petiole up to the middle, free part triangular-lanceolate, acute or subacute, 8-13 mm long, sericeous by appressed white hairs; leaves 12-18 cm long, arcuately curved, the rachis twice as long as the petiole; leaflets 9-11 pairs, oblong-lanceolate, acute, 10-22 mm long, 3-7 mm broad, broadest in lower part, round-based, covered on both sides with appressed white hairs, more profusely so beneath; peduncles exceeding the leaves, 20-35cm long, straight or somewhat curved, rather thick, angular, covered with scattered appressed white and black hairs; raceme 12-22 cm long, remotely 20-30-flowered; bracts ovate-lanceolate, small, 2-3 mm long, covered with scattered hairs; pedicels very short; calyx cylindric, ca. 9mm long, covered with appressed black hairs, obliquely truncate, the subulate teeth ca. 1 mm long, converging on one side; corolla yellow; standard ca. 20 mm long, 6-7 mm broad, oblong to ovate-oblong, arched~ recurved, cuneate-attenuate toward base, round-tipped; wing-petals 16-17 mm long, the lanceolate limb 9 mm long, 2 mm broad, the basal auricle elongated, the claw about as long as the limb; keel 14-15 mm long, 468 629 the limb inequilaterally oval, 7 mm long, 3 mm broad in upper part, short- auriculate, acutish, the claw somewhat longer than the limb; ovary stipitate, linear, covered with appressed white hairs, especially on the ventral side; style ca. 8 mm long, naked; pods straight or slightly curved, cernuous, borne on a stipe ca. 5 mm long, triangular-cylindric, ca. 25mm long, ca. 2mm broad, 3 mm thick, deeply grooved ventrally, carinate dorsally, terminating in a straight firm beak 4-6 mm long, bilocular, many-seeded, covered with scattered appressed white and black hairs; seeds reniform, almost black, 2 mm long, 1 mm broad. Fl. and fr. June- July. (Plate XL, Figure 3). Dry stony mounds and rocky slopes, at altitudes of 1000- 1200 IM. a Caucasus: E. Transc. (Kuba, Gyul'gan-chai), Tal. (Zuvant). Endemic. Described from Talysh. Type in Leningrad. 597. A. cuscutae Bge. Astrag. geront. II (1869) 207; Boiss. Fl. or. II, 466; Grossh. Fl. Kavk. II, 334. Perennial, grayish by appressed hairs, acaulescent; stipules lanceolate, acuminate, silky-tomentose, firm; leaves 7.5-15 cm long, long-petioled; leaflets 4-7 pairs, sometimes (probably in young plants) the lower ones consisting of a single leaflet, 4.5 cm long, ca. 3mm broad, or trifoliate, at first covered with white hairs; peduncles exceeding the leaves, up to 30 cm long, ascending, angular, appressed-hairy; racemes broad, many- flowered, 7.5-15 cm long, loose; flowers subsessile; bracts ca. 5mm long, narrowly lanceolate; calyx cylindric, ca. 12 mm long, covered with white and black bristles, the tube 3-4 times as long as the subulate teeth; corolla azure (according to Grossheim), drying violet; standard oblong, ca. 23mm long, ca. 9mm broad in upper part, gradually attenuate toward base, scarcely retuse; wing-petals ca. 20 mm long, the limb oblong, as long as the claw, broader in upper part, scarcely retuse, auriculate at base; claw of keel 1.5 times as long as the limb; pods lanceolate, 2.5-3 cm long, 4mm broad, far surpassing the corolla, slightly falcate, gray-tomentose with appressed hairs, carinate on one side, grooved on the other, sessile or nearly so, attenuate toward base, mucronulate at apex, faintly spotted at first, covered with stiff appressed black and white hairs, bilocular; seeds angularly reniform, smooth, olivaceous. Fl. May; fr. May-July. Stony or gravelly slopes, at altitude of about 400 m.— Caucasus: E. Transc. (Shemakha, Divichi district), Dag. (Kaitag-Tabasaran, Mt. Dvavan- dag). Endemic. Described from the Caucasus, Shemakha district. Type in Paris. Note. A.cuscutae Bge.resemblescloselyA. Prilipkoanus Grossh. in floral characters, the distinguishing characters being the smaller number of leaf pairs (4-7 as against 8-12 pairs), longer calyx, and cernuous pods. Both species need further study; it is possible that more ample material would show that there is no difference between the two species. 598. A. atripurpureus Boiss. Diagn. ser. I, 9 (1849) 54; Boiss. Fl. or Il, 470; Bge. Astrag. geront. II, 210; Grossh. Fl. Kavk. II, 336.— A. paphlagonicus Bge. Astrag. geront. II (1869) 210. Perennial, acaulescent or nearly so, vested throughout with appressed silky white hairs, (15) 30-40 cm tall, from a thick woody root; stipules triangular-lanceolate, acuminate or acute, adnate to petiole, covered with appressed white hairs, 10-12 mm long; peduncles exceeding the leaves, 469 630 elongating in fruit; racemes many-flowered, ovoid, at length elongating, rather dense, becoming loose in fruit; flowers spreading, finally cernuous; pedicels 1-2 mm long; bracts linear, 4-5 mm long, covered with appressed black and white hairs; calyx cylindric, 8-10 mm long, ca. 2 mm broad, covered with appressed white and black hairs, the filiform teeth ca. 2 mm long; corolla dark purple, 20-24 mm long, more than twice the length of calyx; standard oblong-lanceolate, 6-7 mm broad about the middle, recurved, 3 mm broad in upper part, slightly retuse, gradually cuneate- attenuate toward base; wing-petals 15-18 mm long, the limb lanceolate, curved, notched on the inner side, elongate-auriculate at base, as long as the claw; keel 13-14 mm long, oval, curved, ca. 6 mm long, 2.5 mm broad in upper part, the basal auricle small; ovary stipitate, lanceolate, ca. 4mm long, sparsely appressed-hairy; style naked, ca. 10 mm long; pods pendulous, bilocular, oblong-cylindric, somewhat curved at first, at length becoming straight, covered with appressed white hairs, dark- spotted, 14-18 (20) mm long, 3-4 mm thick, terminating in a point 2-3 mm long; seeds angularly reniform, small, ca. 2 mm long, brown. Fl. June; fr. June-July. Dry sandy and stony slopes.— Possibly growing in the Caucasus. Gen. distr.: Arm.-Kurd. (Turkish Armenia), Bal.- As. Min. (Pizidiya, Kariya, Anatolia). Described from Asia Minor. Cotype in Leningrad. 599. A. punctatus Bge. Astrag. geront. II (1869) 209; Boiss. Fl. or. II, 465; Grossh. in Beih. zum. Bot. Centralbl. XLIV (1927) II, 222.— Bxs. sel) on.exs: INO. i230. Perennial, acaulescent, cinereous, 10-20 cm tall, from a woody divided caudex; stipules membranous, small, linear, spreading; leaves commonly 6-8 cm long (according to Bunge 12-13 cm long); leaflets 6-8 (Bunge: 4-6 (7)) pairs, oval, ovate, or oblong, gray on both sides, 6 mm long (Bunge: 4-10mm); peduncles straight, exceeding the leaves, 12-15 cm long; racemes remotely 8-12-flowered; bracts ovate, membranous, exceeded by the pedicel; calyx 10 mm long, covered with short appressed black hairs, the tube 8-9 times (Bunge: 3 times) longer than the lanceolate teeth; standard broadly ovate, 23 mm long, 11 mm broad, 5-5.5 mm in upper part, truncate, attenuate toward base into a cuneate claw ca. 5 mm long; pods 2.5-3 cm long (Bunge: 3-4.5 cm), borne on a stipe 7 mm long (Bunge: more than 10 mm), cernuous,. oblong-lanceolate, straight, rarely very slightly curved, sharply carinate dorsally, appressed-hairy, spotted with dark red, together with the stipe and mucro 3-4.5 cm long, the valves navicular. Fl. May; fr. June. Stony slopes, at altitudes of 1350-1600 m.— Possibly occurring in the -southern frontier regions of the Caucasus. Gen. distr.: N.W. Iran, near Zeidabad and to the east of Tabriz. Type in Paris. Note. The description of this species has been somewhat modified and supplemented on the basis of specimens collected on the classical site. 600. A. longicuspis Bge. Astrag. geront. II (1869) 211; Boiss. Fl. or. Il, 477; Grossh. Fl. Kavk. II, 335; Grossh. in Beih. zum. Bot. Centralbl. SINE (URE) Mle 22e Perennial, acaulescent, whitish-gray throughout, from a woody root and a short caudex clothed in leaf petioles and stipules; stipules ovate, short, covered with gray hairs; leaves 5-6.5 cm long; leaflets (7) 9-14 pairs, small, approximate, oblong to oblong-elliptic, subobtuse to acute, 470 631 632 5-6 mm long, 2-3 mm broad, covered with white and gray hairs; peduncles ca. 10 cm long, exceeding the leaves, straight; racemes loosely 5-12- flowered, elongating in fruit; bracts ovate or lanceolate, exceeding the pedicel, 2-4 mm long; flowers finally nodding, subsessile; calyx 10-12 mm long, covered with appressed and predominantly black hairs, the tube 3 times as long as the lanceolate teeth; corolla purple-violet; standard 12-23 mm long, the limb lanceolate, 6-8 mm broad at the middle, attenuate at apex, scarcely retuse; wing-petals subentire, 15-18 mm long, the claw about as long as the oblong limb, short-auriculate at base; keel 11-13 mm long, the oval limb about as long as the claw; ovary sessile, 20-22-ovuled, oblong, hairy, gradually attenuate into the filiform style narrowing toward the end, hairy at the base; pods 20-25 mm broad, 3 times as long as the calyx, slightly arcuate, covered with appressed white hairs, attenuate at both ends, minutely beaked, carinate dorsally, rounded and grooved ventrally. Fr. May. Calcareous hills.—Caucasus: S. Transc. (Nakhichevan', Negram Station — Grossheim's indication). Gen. distr.: Iran (Lake Urmiya). Described from the Urmiya Lake region. Type in Leningrad. 601. A. fodinarum Boiss. et Noé ex Bge. Astrag. geront. II (1869) 209; Boiss bike or Il, 470. A. campylosema var. glabrescens Boiss. Diagn. ser. 2, II (1854) 30. Perennial, green appressed-hairy; stipules ovate-lanceolate, glabrous, elongated; leaves up to 30 cm long; leaflets 15-16 pairs, oblong, obtuse, large, up to 2.5 cm long; peduncles exceeded by the leaves; racemes long and broad, the flowers at length nodding; bracts lanceolate, exceeding the pedicel; calyx densely covered with appressed black hairs, ca. 10 mm long, the tube about twice as long as the subulate teeth; standard ca. 2.5 cm long, more than twice the length of calyx, oblong at base, the upper part furnished with a long broadly linear appendage, round-tipped; wing-petals slightly gibbous, unequally bidentate at apex; ovary short, glabrous, 16-ovuled. Fl. May. Possibly growing in S. Transcaucasia. Described from Turkish Armenia, from the vicinity of Bakermaaden. Type in Geneva. Note. We have not seen specimens of this species; the description has been composed by Bunge and Boissier. 602. A. campylosema Boiss. Diagn. ser. I, 2 (1843) 82; Boiss. Fl. or. II, 469; Bge. Astrag. geront. II, 209; Grossh. Fl. Kavk. II, 336. Perennial, acaulescent or nearly so, 20-30 cm tall, sparsely vested in upper part with white hairs and more profusely in lower part; root woody; stipules broadly lanceolate, appressed-hairy; leaves 15-30 cm long, the petiole stout, shallowly sulcate, at first profusely hairy finally glabrate, somewhat shorter than the rachis; leaflets large, 5-7 pairs, ovate-oblong to ovate, 13-30 mm long, 5-14 mm broad, obtuse to subacute, sparsely covered with appressed hairs, more profusely so at first; peduncles slightly exceeding the leaves; racemes oblong, many-flowered, at first rather dense, at length strongly elongating, the lower flowers remote, short-pediceled, 1-2 mm long; bracts ovate, very small, ca. 1 mm long; calyx cylindric, half the length of corolla, covered with appressed mostly black hairs, the tube 3-5 times as long as the unequal subulate teeth; corolla yellow, 20-23 mm long; standard recurved, oblong, ca. 8 mm broad at the middle, attenuate to 3-4 mm in upper part, scarcely retuse, 471 633 narrowed at base; wing-petals ca. 18 mm long, the filiform claw as long as the limb; wing-petals lanceolate, 2 mm broad at apex and 3 mm at the middle, angular, unequally bidentate above, auricle recurved at base; keel 15 mm long, the claw 1.5 times as long as the limb, the limb 3 mm broad curved ovaloid; ovary short-stipitate, lanceolate, covered with appressed white hairs, gradually prolonged into the style, thisfiliform, naked, 8mm long, curved at the end; pods pendulous, oblong-cylindric, finally twice the length of the calyx, 15-18 mm long, obtuse, mucronulate, grooved dorsally, white-hairy, at length glabrate, 15 mm long and 4 mm broad. El. June’ Possibly growing in the frontier regions of S. Transcaucasia. Gen. distr.: Arm.-Kurd. (Kurtlyu in Kagyzman district near region of Kars), Bal.- As. Min. (Tavr). Described from Tavr. Type in Geneva. 603. A. ackerbergensis Freyn in Bull. Herb. Boiss. sér. 2, V (1905) 562; Fedtsch. O. et B. Consp. Fl. Turk. I, 230.— A. pendulus Kar. ex Pedtscn.4 ©. et -B. ilve-non-De- Perennial, acaulescent ornearly so, silvery by appressed hairs, 30-40cm tall; root stout, woody; caudex short, clothed in persistent petioles and stipules; stipules petiolar, 7-12 mm long, covered with appressed and partly spreading hairs, the free portion elongate-lanceolate, long-acuminate; leaves ascending, 10-20 cm long, the petiole subradical, straight, just exceeding to 1.5-2 times as long as the rachis; leaflets distant, 3-5 pairs, ovate or elliptic, 7-22 mm long, 5-13 mm broad, green, sparsely hairy beneath with appressed silvery hairs, commonly acute or subacute, rarely obtuse or round-tipped, cuneate-based, borne on very short petiolules; peduncles 1-3, exceeding the leaves, straight, covered with appressed white and in the inflorescence black hairs, sulcate; racemes loosely 10-25-flowered, 5-12 cm long, the lower flowers remote; bracts filiform, ca. 3 mm long, about twice the length of the short pedicel; bracteoles ca. 0.5 mm long, filiform; calyx cylindric, 9-10 mm long, covered with short appressed black hairs especially on the teeth, the acute teeth2-2.5mm long; standard 22 mm long, 9 mm broad at the middle, attenuate to 4mm in the upper quarter, scarcely retuse, cuneate-attenuate toward base; wing-petals 19 mm long, the limb spatulate-oblong, round-tipped, 10mm long, ca. 3 mm broad in upper part, 1.5 mm at base, the auricle somewhat reflexed; keel 15-16 mm long, the limb 6 mm long, ca. 4 mm broad, curved on the outside; ovary ca. 6 mm long, oblong, glabrous, short~ stipitate; style somewhat curved, ca. 10 mm long, naked; pods cernuous in maturity, short-stipitate, bilocular, somewhat curved, narrowly oblong to lanceolate, 2-2.8 cm long, 3.7-4 mm broad, terminating in a rigidbeak 1.5-2 mm long, yellow, chestnut-brown-spotted, glabrous, somewhat appressed laterally, shallowly canaliculate ventrally; seeds angularly reniform, 3-4 mm long, 1-1.5 mm broad, chestnut, pitted. Fl. May-June; fry June-July. Mountains, in the zone of mixed~grass and needlegrass-and-fescue associations, at altitudes of 1500-1600 m.— Centr. Asia: Mtn. Turkm. Endemic. Described from Suluklyu in the central part of Kopet Dagh Range. Type in Leningrad. 604. A, brachyrachis M. Pop. in Tr. Turk. Gos. univ., No. 4 (1922), OZ) GH: 472 634 Perennial, nearly acaulescent, 20~30 cm tall, rather compactly cespitose; stems 1-2.5 cm long, densely covered with appressed hairs, white; stipules adnate to petiole, triangular-ovate, ca. 1.5 mm long, acute, densely white-hairy; leaves 10-15 cm long, the petiole rather stout, 1.5-2 times the length of the rachis, cinerescent by appressed black and white hairs, indurescent and partially persistent; leaflets 3 pairs, obovate- orbicular, obtuse, often mucronulate, broadly cuneate at base, 7-15 mm long, 5-10 mm broad, rigid, densely covered on both sides, more so beneath, with appressed hairs, prominently veined, distant; peduncles 12-30 cm long, rather stout, loosely covered with appressed white hairs, indurated, partially persistent; racemes 8-12 cm long, loosely and remotely 7-12-flowered, straight, firm; bracts triangular-ovate, acute, ca. 1 mm long, white-hyaline, covered with appressed white and fewer black hairs; pedicels 2-2.5 mm long, covered with appressed white hairs, flowers erect; calyx long-cylindric, 15-20 mm long, densely covered with short appressed white hairs interspersed with isolated black hairs, the tube 5-7 times as long as the subulate-linear teeth; corolla yellow; standard 22-25 mm long, 8-9 mm broad, the limb broadly oblong, rounded-truncate and slightly retuse, 1.5 times as long as the cuneate claw; wing-petals 22-25 mm long, the limb about half as long as the claw; keel equaling the wings, the limb obliquely truncate, obtuse, about half as long as the claw; ovary linear, glabrous, ona stipe 2 mm long, 20-ovuled; pods cernuous, linear, 15-40 mm long, straight or slightly curved, borne on stipe 2-3 mm long, carinate ventrally, deeply grooved dorsally, terminating in a straight beak 2-3 mm long, coriaceous, glabrous, bilocular. Fl. andfr. May-July Mottled outcrops and gravel- and stone-covered mountain plateaus. — Centr. Asia: Pam.- Al.: Karatau foothills, eastern part of the Turkestan Range, within limits of Isfarin (''Santo'' petroleum industries) and Leninabad (vicinity of Makhram settlement) districts and Tadzhikistan. Endemic. Described from Karatau foothills. Type in Tashkent. Series 7. BRACHYCARPI Boriss.— Pods oblong or ovoid, usually large, coriaceous, inflated, sometimes triquetrous, erect. 605. A. brachycarpus M.B.Fl. taur.-cauc. II (1808) 201; III, 498; Lidbyy Pl: Ross. 1,690; Bee. Astrag. geront. 11,212; Boiss. Fl.) or-)II, 475; Schmalh. Fl. I, 283; Grossh. Fl. Kavk. II, 335.— A. clavatus Ldb. Fl. Ross. I (1842) 648, non DC.— A. macranthus Willd. ex Ldb. eee650e— les :(Ma Bs Cent) pl. rar; tab; 58; Sims, Bot. Mag. tab. '2335.— BS ER No} 23) Perennial, acaulescent or nearly so, grayish-green, silky by appressed hairs, from a vertical woody taproot; caudex many-headed, the divisions up to 5 cm long, clothed in imbricated stipules and old leaf petioles; stipules lanceolate, acuminate, petiolar, adnate to petiole and connate up to about the middle, 5-10 mm long, covered with appressed white hairs; leaves 5-25 cm long, the petiole somewhat shorter than to equaling the rachis; leaflets 7-15 pairs, elliptic to oblong-elliptic. glabrous or sparsely covered with isolated appressed hairs on the upper face, sparsely appressed-hairy beneath, moredensely so when young, subobtuse to obtuse, rarely subacute, 10-20 mm long, 3-7 mm broad; peduncles 10-32 cm long, commonly exceeding the leaves, covered with appressed hairs, some of these black in upper part, ascending or erect; racemes many-flowered, loose, elongating in fruit; bracts 1-3 mm long, oblong to lanceolate, subtended by 2 minute bracteoles; calyx cylindric, covered with appressed 473 635 636 black and white hairs, 8-11 mm long, the tube 2-4 times as long as the subulate-lanceolate teeth; the teeth crowded together at one side; corolla dingy violet-purple; standard ca. 25 mm long, 8-9 mm broad at the middle, attenuate in upper part to 4mm, scarcely retuse, the lower third cuneate-attenuate toward base; wing-petals 16-19 mm long, the limb lanceolate, 9-10 mm long, 2 mm broad, the limb bidentate at apex, undulate on the inner margin, the basal auricle oval and somewhat reflexed, the claw about as long as the limb; keel 13-15 mm long, the limb 5-6 mm long, ca. 3mm broad, somewhat shorter than the claw; ovary sessile, oblong, ca. 5mm long, sparsely covered with very short hairs; style 8-10 mm long, glabrous; pods straight, obovoid-clavate, attenuate toward base, sometimes about the length of the calyx, 10-17 mm long, ca. 5mm thick, reddish, red-spotted, appressed-hairy, more densely at first, slightly curved, large, coriaceous, terminating in a slender beak 2-3 mm long, bilocular, scarcely carinate dorsally, shallowly grooved ventrally, sessile, few-seeded; seeds reniform, castaneous, 2.5 mm long, 1.5 mm broad, smooth. Fl. March-May; fr. May. Rocks, dry clay- and stone-covered hillslopes and foothills, at altitudes of 200-850 m.— Caucasus: Cisc., Dag., E. and S. Transc. Gen. distr.: Arm.-Kurd., Bal.- As. M. Described from Mt. Beshtau. Type in Leningrad. 606. A. robustus Bge. Astrag. geront. II (1869) 212; Boiss. Fl. or. II, 475; Grossh. Fl. Kavk. II, 336. Perennial, cinereous by appressed hairs, acaulescent, erect, 25-45 cm tall, from a long vertical woody root; stipules petiolar, ovate-triangular, acuminate, sericeous, prominently nerved, 8-14 mm long, adnate to petiole up to the middle; leaves 20-25 cm long, long-petioled, the petiole finely sulcate, appressed-sericeous, equaling or exceeding the rachis; leaflets 6-10 pairs, 7-25 mm long, 4-13 mm broad, oblong-ovate to ovate, obtuse to subacute, covered on both sides with appressed gray hairs, rounded at base; peduncles exceeding the leaves, 25-47 cm long; racemes rather dense, elongating in fruit, the distant flowers borne on pedicels ca. 2mm long; bracts exceeding the pedicel, lanceolate or ovate, 2-4 mm long, covered with appressed white hairs; bractoles small, up to 1 mm long, subulate, inserted at the calyx base; calyx 8-10 mm long, covered with short appressed black bristles interspersed with white hairs, the triangular teeth ca. 2 mm long; corolla lurid; standard 20 (25) mm long, the limb broad, ovate, 8 mm broad at the middle, attenuate in upper fourth to 4mm, truncately retuse, the claw broadly cuneate; wing-petals 14 (22) mm long, scarcely retuse, the oblong limb 2 mm broad, as long as the claw, subacutely auriculate at base; keel 12 (16) mm long, the limb inequilaterally oval, 3 mm broad, auriculate at base, acutish at apex; ovary 13-16-ovuled, oblong, 4mm long, white-hairy, short-stipitate; style hairy up to the middle, ca. 10 mm long, curved at the end; pods straight, oblong-ovoid, ca. 13 mm long, 5-6 mm broad, 4 mm thick, mostly straight, sometimes somewhat enlarged or narrowed toward base, rounded and not grooved dorsally, terete, carinate ventrally, covered with white and black bristles, cross-wrinkled, spotted, rigid, bilocular, borne ona stipe ca 1 mm long; seeds 8-10, reniform, 2 mm long, 1.5 mm broad, castaneous, smooth Fl. May-July; fr. June-August. (Plate XL, Figure 1). Mountain slopes.— Caucasus: S. Transc. (Ordubad, Migry, Nakhichevan). Gen. distr.: Iran. Described from Iranian Azerbaijan between Khoi and Koshadara. Type in Leningrad. 474 137 Perennial, silky silvery-gray by appressed hairs, acaulescent, erect, 20-40 cm tall, clothed at base in imbricated remnants of petioles and stipules, from a long woody root; stipules adnate to petiole, 5-7 mm long, broadly triangular at base, reddish, appressed-hairy, the free portion long-acuminate; leaves 15-17 cm long, the petiole as long as to longer than the rachis; leaflets 5-7 pairs, 6-20 mm long, 5-10 mm broad, oblong- ovate to ovate, obtuse or minutely mucronulate, covered on both sides with short appressed-hairs; peduncles commonly greatly exceeding the leaves, 20-40 cm long, covered with predominantly white hairs; racemes rather loosely many-flowered, elongating in fruit, the rachis covered with appressed predominantly black hairs; pedicels black-hairy, 1-1.5 mm long; bracts 2-3 mm long, lanceolate to ovate; bracteoles at the base of calyx obsolescent, less thanl mm long, filiform; calyx 9-10 mm long, sparsely covered with short appressed black bristles, the thickish teeth 1-1.5 mm long; corolla violet; standard 21-24 mm long, attenuate in upper part, 8-10 mm broad at the middle, cuneate-attenuate at base, slightly retuse; wing-petals 17-18 mm long, the limb oblong-lanceolate, wavy-margined, 2mm broad, somewhat longer than the claw, the basal auricle short and round; keel 13-15 mm long, the oval limb about as long as the claw; ovary oblong, covered with black and white hairs, sessile; style naked, ca. 10 mm long; pods oblong, 15-17 mm long, ca. 4.5 mm broad, 3 mm thick, somewhat flattened, faintly wrinkled longitudinally, not carinate or grooved, densely covered at first with appressed black hairs interspersed with white, sparsely hairy in maturity, terminating in an oblique beak ca. 4mm long, bilocular; seeds rounded-reniform, castaneous, smooth, ca. 2mm in diameter. Fl. May-July; fr. July-August. Slopes of mountain gorges, at altitudes of 1800-2300 m.— Caucasus: S. Transc. (vicinity of Lake Sevan, adjacent part of Zangezur range, Erevan vicinity). Endemic. Described from the vicinity of Lake Sevan. Type in Leningrad. Note. Resembling A. robustus Bge. from which it differs in violet and longer flowers (15-17 mm long and 4.5 mm broad, as against 13 and 5-6 mm, respectively), black-hairy sessile pods, and the distribution area. 608. A. cinereus Willd. Sp. III (1800) 1315; Bge. Astrag. geront. II, ZO borss wh lor. INN 4isoA. Clavatus Bp obrevibracteatus DEC: Astrag. (1802) 230, excl. syn. Tournef. Perennial, acaulescent, 15-20 cm tall; stipules oblong-acuminate, marcescent; leaves 7-10 cm long; leaflets elliptic to suborbicular, gray- sericeous on both sides, 7-8 pairs, obtuse, 10-13 mm long; peduncles straight, exceeding the leaves, 15-20 cm long, gray tomentose; inflorescence spiciform, oblong-elongate, the flowers partly nodding; bracts very small, the lower suborbicular, the upper elongate to lanceolate; flowers greenish-yellow; calyx cylindric, ca. 7mm long, covered with appressed white and black hairs, the subulate teeth ca. 1 mm long; corolla 23-25 mm long; pods 7-15 mm long, 5 mm broad, ovate to ovate-oblong, straight, obtuse and mucronate, slightly notched dorsally, carinate ventrally, appressed-tomentose, short-beaked. Caucasus: S. Transc. (Elenovka). Endemic. Described from Armenia. Type in Berlin. Note. A. clavatus DC., described from Armenia, is probably Synonymous with A. cinereus Willd. According to Willdenow'y, A. cinereus has nodding flowers; later authors (Bunge, Boissier) do not refer to this. 475 638 609. A. taschkendicus Bge. Astrag. Turk. (1880) 280.—Ic.: Fl. hNadzhik, Vii@lo3t)"tabe 0). bxse) (Hen AM Ii) No. SOT. Perennial, acaulescent, 20-50 cm tall, erect, green, from a stout straight deep-penetrating root; stipules adnate to petiole up to the middle or nearly so, 8-18 mm long, lanceolate, densely covered on the outside with white hairs, the free portion often subulately long-acuminate; leaves (7) 15-25 (32) cm long, the short petiole and the rachis covered with very short appressed hairs or glabrate; leaflets (7) 10-14 (17) pairs, large, (5) 10-18 (25) mm long, oblong-oval to oval, rarely suborbicular or ovate, round-tipped, mostly minutely mucronate, glabrous above, covered beneath with scattered very short appressed bristlelike white hairs; peduncles (9) 13-20 cm long, straight, elongated, rather sparsely covered with short appressed white hairs, angled-sulcate; racemes rather loosely 20-50- flowered, ovoid-oblong, finally elongating and becoming increasingly loose, (13) 15-30 (50) cm long, the flowers and fruit remote; bracts lanceolate, 2-4 mm long, acute, exceeding the flowers, hyaline-margined, glabrate; bracteoles caducous, small, ovate, translucently membranous, ciliate; pedicels ca. 1 mm long; calyx cylindric, 11-13 mm long, glabrate or covered with very short and rather scattered appressed black hairs, membranous, the teeth 4-5 mm long, linear, acute or subulate-tipped; corolla mostly dark purple; standard recurved through nearly half a circle, 18-20 mm long, the limb oblong-ovate or subrhomboid-oval, attenuate upward, broadest about the middle, obtuse or retuse, abruptly contracted into a narrow claw 3-5 mm long; wing-petals 15-18 mm long, the limb oblong, slightly curved, somewhat enlarged in upper part, the claw 6-8 mm long; keel 13-16 mm long, the limb oblong, curved, 2.5-3 mm broad in upper part; ovary short-stipitate, tomentulose, linear; style naked, ca. 11 mm long; pods erect, 10-15 (18) mm long, ca. 5 mm thick, oblong-oval, terete, terminating in a curved beak 1-2.5 mm long, thickly and convexly ribbed-carinate ventrally, straight and slightly concave-sulcate dorsally, rather sparsely covered with short white hairs, rarely glabrate, bilocular, few-seeded. Fl. April-June; fr. May-July. (Plate XLI, Figure 5). In the ephemeral vegetation zone, loess or loesslike grassy slopes in the witchgrass~steppe region and the lower part of the juniper zone, at altitudes of (450) 900-1800 (2200) m.— Centr. Asia: Pam.- Al., W. Tien Shan. Endemic. Described from the vicinity of Tashkent. Type in Paris. 610. A. platyphyllus Kar. et Kir. in Bull. Soc. Nat. Mose. XV (1842) 345; Bge. Astrag. geront. Il, 211; Ej. Astrag. Turk. 280; Fl. Tadzhik.. V, 431.— A. sykensis Freyn in Bull. Herb. Boiss. sér. 2, V (1905) 559. Perennial, acaulescent, vested with isolated appressed hairs, green, 10-35 cm tall; stipules adnate to petiole and connate to high up, linear and long-acuminate, 10-13 mm long, covered with appressed white hairs; leaves subradical, 10-25 cm long, the petiole shorter than the fachis; leaflets 4-8 pairs, elliptic to suborbicular, obtuse, subcanescent at first by appressed white hairs, finally green and sparsely appressed-hairy; peduncles equaling to twice as long as the leaves, mostly straight, sulcate; racemes short, ovaloid-capitate, 2-8 (17) cm long; bracts 3-4 mm long, lanceolate; bracteoles linear or filiform, 5-10mm long, thinly membranous; calyx ca. 8-13 mm long, the teeth lance-subulate, recurved at the tips, the lateral ones ca. 5 mm long, the others somewhat shorter; corolla violet; standard 20-25 mm long, lance-elongate, gradually attenuate toward 639 base and toward apex, 5-9 mm broad about the middle, retuse or entire 476 at apex; wing-petals 15-20 mm long, gibbous above the auricles, retuse, the limb as long as the claw; keel 10-17 mm long, the oval limb somewhat shorter than the claw; pods broadly ovate, triangular, acute, terminating in an incurved point, enlarged at base, 10-13 mm long, 4 mm broad, appressed-hairy, bilocular; seeds angularly reniform, brown, lustrous, 3-4 mm long, ca. 2mm broad. Fl. May-July; fr. July-August. (Plate XLI, Figure 1). Steppe slopes, in gravelly and stony soils, from low foothills to the alpine zone, at altitudes of 400-3000 m.— Centr. Asia: T. Sh., Pam.-Al. (the ranges Alai, Tukestan, Peter I) Zeravshan range, Dzu.- Tarb. (Tarbagatai range, Dzungarian Ala-Tau). Gen. distr.: Dzu.-Kashg. (Kuldja). Described from the rocks of Alatau Mountains, rivers Lepsa and Sarkan. Type in Leningrad. Note. A. platyphyllus is an exceedingly polymorphic species with an extended distribution area. The variable characters include the size of corolla and of its parts, size of calyx and its teeth, height of plants, and less frequently the shape and number of leaflets, the latter not usually exceeding 8 pairs, the number of 5-6 pairs being most prevalent. Series 8. ELONGATI Boriss.— Pods oblong, cernuous, sometimes slightly curved; flowers dingy violet or purple. 611. A. coelestis Boiss. Fl. or. II (1872) 470; Grossh. Fl. Kavk. II, 336.— A. pendulus Bge. Astrag. yeront., I. (1869) 211, non DC. Perennial, ca. 15 cm tall, cinereus by appressed white hairs; stipules triangular-lanceolate, sericeous, long-acuminate; leaflets 8-10 (11) pairs, elliptic, 8-12 mm long, 2-4 mm broad, subacute, sometimes glabrate above; peduncles ascending, somewhat shorter than to very slightly “exceeding the leaves; racemes capitate or ovoid, short, few-flowered, broad, at first dense, finally loose; bracts small, triangular-lanceolate, equaling or exceeding the pedicel; calyx covered with appressed white and black bristles, the tube 3-5 times as long as the teeth; corolla 18-20 mm long, dingy violet-purple; standard oblong, about twice the length of the calyx; limb of standard strongly attenuate upward, enlarged to 6 mm at the middle, cuneate-attenuate toward base; wing-petals acutish, sharply bidentate at apex, undulate on the inner margin, the auricle recurved; limb of keel ovate, about as long as the claw; ovary oblong, 10-14-ovuled; style naked; pods (10) 15-18 mm long, ca 5 mm broad about the middle, pendulous, covered with black bristles, rugose, oblong, subcompressed, more or less acuminate at both ends, rounded dorsally, carinate ventrally, terminating in an obliquely divergent point. July. 640 Dry stony slopes, at altitudes of 1800-2100 m.— Caucasus: S. Transc. (Lake Sevan, Alagez). Endemic. Described from Lake Sevan. Type in Geneva. Note. Resembling A. brachycarpus M.B., in flowering condition, the distinguishing characters being the pendulous somewhat flattened and obliquely point-tipped pods and the spreading flowers. The indications of Karelin and Fedchenko (Consp. Fl. Turk. I, 230) concerning the occurrence of A. pendulus in Turkmenia refer to A. ackerbergensis Freyn. 612. A. elongatus Willd. Sp. III (1800) 1314; DC. Prodr. II, 305; Bge. Astrage ceront. 1175209; Boiss.Pl:or? 1, (466.— A. pendu luis" DC: Astrag. (1802) 232. 477 641 Perennial, acaulescent; stipules ovate, acuminate, hairy; leaves ca. 14 cm long; leaflets elliptic, acute, covered with gray hairs, 4-7 pairs, [?] ca. 13 mm long, appressed-hairy; peduncles exceeding the leaves, straight, not sulcate, ca. 30 cm long; racemes elongated, covered with isolated hairs; flowers erect, finally nodding, purple; pods oblong, very firm, appressed-hairy, cernuous, bilocular, oblong or ovoid, ca. 13 mm long, 2-3-seeded, mucronate; pods, according to De Candolle, resembling pistachio fruit in shape and size. Fl. July. Possibly occurring inthe Caucasus: Recorded for ''Armenia'' (Tournefort). Gen. distr.: Bal.- As. M. (Galati, Kappadocki). Described from Galati. Note. According to De Candolle, resembling A. brachycarpus, but differing from it in the nodding fruits. Series 9. RECURVI Boriss.— Pods triquetrously rounded-ovoid, covered with long spreading white hairs, rigidly coriaceous, the beak long and reflexed. 613. A. hissaricus Lipsky in A.H.P. XXVI (1907); 173; Fl. Tadzhik. V, 432. Perennial, acaulescent, scabrous by bristlelike appressed white hairs, (17) 25-30 (50) cm tall, erect or subdecumbent to ascending; stipules adnate to petiole to high up, lanceolate, long-acuminate, 15-20 mm long, imbricated, glabrate or appressed-hairy on the outside; leaves 15-30 cm long, about equaling the pedicel; leaflets 8-11 pairs, elliptic-lanceolate, rarely narrowly lanceolate to linear, sometimes ovate, 10-30 mm long, 2-12 (15) mm broad, acute or acuminate, rarely subacute, mucronulate, covered on both sides with appressed bristlelike hairs; peduncles erect or slightly assurgent, curved, 20-30 (50) cm long, commonly somewhat exceeding the leaves; racemes densely capitate or oblong, 2.5-6.5 cm mostly 4-5 cm long, not elongating in fruit; bracts small, oval, membranous-hyaline, white, sparsely hairy, ciliate, about half as long as the calyx, ca. 5mm long; calyx cylindric, 9-12 mm long, covered with short appressed black and white hairs, the tube 2-4 times as long as the unequal subulate teeth; corolla twice the length of the calyx, 16-25 mm long, lurid with a lilac standard; standard 16-25 mm long, the limb lanceolate, somewhat enlarged in lower part, gradually attenuate into and 2-3 times as long as the claw; wing-petals 15-18 mm long, the limb lanceolate, attenuate upward, as long as the claw; keel 12-17 mm long, the claw 1.5 times as long as the limb; ovary oblong, sessile, hairy; style 8-10 mm long, naked; pods sessile, 7 mm long, subovoid, rigid, subtriquetrously terete, covered with rather long spreading white hairs, 2-4-seeded, the beak mostly arched-recurved and about as long as the body of the pod, the persistent style brittle; seeds ovaloid-oblong, ca. 5 mm long and 2 mm broad, inequilateral, minutely pitted, brown, mostly punctate with dark red. Fl. May-June; fr. June-July. (Plate XL, Figure 2). In close-grained soil on slopes of the deciduous woodland and scrub and the juniper zones, in rosaria, and on rosarium-dominated northern slopes of the ephemeral vegetation zone, at altitudes of 900-1500 m.— Centr. Asia: Pam.-Al. (Gazimailik mountains, Hissar and Darvaz ranges, Khodzha-Gungur-ata mountains, Babatag range). Endemic. Described from Hissar range. Type in Leningrad. 478 342 Series 10. ERIOCARPI Boriss.— Pods oblong, cernuous, densely shaggy with spreading hairs; stipules almost free, barely adnate to petiole. 614. A. namanganicus M. Pop. in Bull. Univ. As. Centr. 14 (1926) [20> ExstembinwA. Ma Now 391. Perennial, nearly acaulescent, 17-60cm tall, rather compactly cespitose, from the divisions of a short caudex; stems1.5-5cm long, angled, densely covered with white hairs; stipulesfree, barely adnate in lower part to petiole, triangular-lanceolate, 1-4mm long, acuminate, membranous, white-hiary onthe outside; leaves 6-22 cm long, the petiole and the somewhat longer rachis canescent by appressed hairs, indurated; leaflets 4-6 pairs, distant, broadly obovate-orbicular, 6-15 mmlong, 5-15mmbroad, round ~tipped or truncate, minutely mucronulate, mostly broadly cuneate, rigid, covered sparsely above and densely beneath with appressed hairs; peduncles firm, straight or curved; 13-60cm [sic]long, 1.5-3timesas long asthe leaves, canescent by rather dense appressed white hairs; racemes loose, 12-25 cm long, very GEMOtel yada o's flowered; bracts oblong-lanceolate, acute, 2-2.5mm long, thinly membranous, more or less covered with white hairs; bracteoles wanting; pedicels 1.5-2 mm long, white-hairy; calyx cylindric, 15-18 mm long, rather densely covered with short appressed white hairs interspersed withfew short black hairs, elongating infruit, the lanceolate acute teeth 2-3 mm long; corolla yellow, persisting in fruit for a longtime; standard 26-30mm long, spatulate-lanceolate, 7-8 mm broad at the broadest upper part, scarcely retuse, gradually prolonged into a narrowly cuneate beakca. 1 mmbroad at base; wing-petals 24-25 mm long, the limb half as long as the claw, linear-oblong, the basal auricle minute triangular; keel 21 mm long, the claw 1.5-2 times as long as the acutish limb; ovaryona stipe 3-4mm long, lanuginous; style straight, naked, ca. 14mm long; pods subcernuous, borne ona stipe 3-4 mm long, oblong, somewhat compressed, acutish, 10-15mm long, 3-5mmbroad, 2-4mmthick, carinate ventrally. shallowly grooved dorsally, subtriquetrous, rigidly coriaceous, densely lanuginous-villous with spreading white hairs, bilocular. Fl. April-May; fr. May-June. (Plate XLI, Figure 3). Dry stony gypseous loess slopes, conglomerates in dry river valleys and beds of runoff streams in the semidesert region.— Centr. Asia: Syr D. (vicinity of Namangan, in Andizhan and Margelan districts). Tien Shan. Endemic. Described from W. Tien Shan. Type in Tashkent. Note. According to Popov, resembling most closely A. brachyrachis, from which it differs in the more numerous leaflets, the looser raceme, and the shorter woolly pods. The free stipules scarcely adnate to petiole, distinguished A. namanganicus from all the other species of the section Proselius. Section 91. MYOBROMOPSIS Boriss. sect. nova in Addenda XI, p.662.— Peduncles much shorter than the leaves; leaves radical; racemes lax; calyx devoid of bracteoles, covered with black hairs, cylindric, not becoming inflated; ovary stipitate; pods large, inflated, 15-27 mm long, ovaloid or broadly ovoid, bilocular, firm, rugose. Green acaulescent plants, sparsely vested with appressed bicuspidate hairs; stipules very short, adnate to petiole merely at the very base, large, membranous. 615. A. Frickii Bge. Astrag. geront. II (1869) 201; Boiss. Fl. or. II, 461; Grossh. Fl. Kavk. II, 334.—A.Galagani Lipsky in A.H.P. XIV, 2, (1898) 258; Grossh. Fl. Kavk. II, 331.— A. Wilhelmsii Czerniajew in herb. Turcz. (ex Bge. ). 479 wel 1\( = Q QOS YD /, OTN hd ae, QS PLATE XLI r. et Kir.—2. A. polygala Pall.—3. A. namanganicus Ka s platyphyllus s M. B.—5. A. taschkendicus Bge. 480 Perennial, nearly acaulescent. green, 10-40 cm tall, with approximate internodes, from a woody root and the divisions of a short caudex 2-5 cm long, vested with bicuspidate hairs; stipules membranous, ovate to broadly ovate, 5-10 mm long, acute or acuminate, profusely and prominently nerved, appressed white-hairy, connate; leaves 10-35 cm long, long- petioled, commonly prostrate, petiole half as long as to sometimes equaling the rachis, both petiole and rachis ribbed, sparsely covered with appressed hairs; leaflets 3-20 mm long, 2-12 mm broad, 10-17 pairs, orbicular, ovate to oblong-ovate, the upper approximate, the lower distant, obtuse, obsoletely mucronulate, glabrous above, sparsely covered beneath with appressed bristles; peduncles much shorter than the leaves, 3-9 cm long, subradical, ribbed; racemes loosely 4-10-flowered, elongating; pedicels 2-5 mm long; bracts membranous, oblong to lance-linear, about equaling the pedicel, 5-6 mm long, covered with appressed black hairs; calyx 10-15 mm long, covered with appressed black hairs, the tube 4-5 times as long as the lance-subulate or lanceolate teeth; corolla pale purple, drying blue, twice the length of the calyx; standard 23-25 mm long, the limb elongate-oblong, bilobate at apex, ca. 8 mm broad, gradually attenuate toward base; wing-petals 18-23 mm long, 2 mm broad, entire, somewhat curved, the basal auricle round obtuse, the claw shorter than the limb; keel 15-19 mm long, the claw 1.5 times as long as the ovate limb; ovary 26°30-ovuled, stipitate, oblong, glabrous; style filiform, ca. 10 mm long; pods large, ovaloid or ovoid to broadly ovoid, 15-27 mm long, 6-13 mm broad, inflated, rigid, flattened and thickly carinate ventrally, broadly and deeply grooved dorsally, bilocular, scabrous by rugosity and sparse woolly hairs, terminating in a somewhat recurved rigid point 3-5 mm long, rounded at base, short-stipitate; seeds 6-10 per locule, pale brown, smooth, reniform, 3-4.5 mm long, 3 mm broad. Fl. June-July; fr. June-September. Grassy mountain slopes, alpine and subalpine meadows, and mixed forests, at altitudes of 1350-3000 m.— Caucasus: Cisc., W. and EF. Transc. Gen. distr»: Bal.- As. M. (former Artvin vicinity). Described from E. Transcaucasia — Racha in vicinity of Oni. Type in Leningrad. Note. Bunge's report relating to the occurrence of this species in Signakhi does not correspond to reality, as the material was collected by Frick much more to the west, in the area of Oni, near Rachi. This is a plant of the alpine and subalpine zones, descending into the timber zone chiefly in the northern and western parts of the Caucasus. Specimens of A. Frickii from Bredzhiiskaya desert and Mukhrovan (Lagovskii's collections) are recorded for parts not associated with A. Frickii, and were probably collected in some place much higher up the mountains. Section 92. XIPHIDIUM Bge. Astrag. geront. I (1868) 123; II, 216.— Frutescent, suffrutescent, and herbaceous perennials, vested with bicuspidate hairs; stipules quite free or partially adnate to petiole at base; leaves imparipinnate, the petiole marcescent; racemes lax, rarelycompact, subcapitate or abbreviated subumbellate, borne on a rather long peduncle; bracteoles wanting; calyxcylindric, not becoming inflated; corolla glabrous, or white or violet or otherwise colored (pleochroism); wings mostly shorter than the standard; pods stipitate or sessile, linear or oblong, coriaceous, bilocular, rather many~seeded. Over 100 species of the section Xiphidium grow inthe USSR. The principal distribution center of the section is associated with the mountains 481 of Central Asia, where the most ancient shrubby species occur. Specialized branches derived from this center spread far to the north, east, west, and, apparently, also to the south. 1. Pods short and broad, ovaloid-oblong or oblong to narrowly oblong, from 6-10 to 15-20 mm long and from 2-3 to 5-6 mm broad, rarely attaining the length of 25-30 (40) mm and then (3.5) 4-5 mm broad. 2. + Pods narrow, elongated, subulately linear, from 20-30 to 100-120mm long-and 1/23) mmibr dad «274 Opening eee” ae ee 44. 2. Corolla yellow (pure yellow, yellowish-brown, or yellowish-green). 3. 4+ ‘(Corolla otherwise colored) not *yellowt 7. 2s yes aan eee iL, 647 3. Flowering racemes short, sometimes finally elongating and then corolla ereenisheyellowe >. [haere 3 GRE GY |p REO e ORO ee se 4. + Flowering racemes very loose, elongating, the flowers very distant 9. 4. Caulescent plants; peduncles commonly exceeding or rarely equaling the leaves and then nevertheless long (6-9 cm)............... + Plants acaulescent or nearly so; peduncles shorter than the leaves or equaling them and then nevertheless short (1.5-5cm).......... 8. 5. Calyx-tube 4-5 times as long as the teeth; calyx hairy, but not DAMNOS SE ON genre aM ON OMe EEA ut ey Re eR anne seen aaa 6. + Calyx-tube 2-3 times as long as the teeth; calyx pannose......... er Pa, Be Ce, CAR! MOORS. 710. A. Lipschitzii Pavl.” 6. sBracts el bis2mamnlonoren a eae te anit 711. A. bossuensis M. Pop. + NBracts) 5=LObmim long head WPL SR POA LIOAS 4 DEEN. cee 7. 7. Meaflets sparsely hairy on. both sides,.green:).0. .¥) 227)... eee ae ee. | RO Ne SO Oe VR ae Seat 709. A. tumescens M. Pop. + Leaflets) profusely, hairy on both sides ;*canescent, (sii) \e--salan seem Sits NSC? BS Ce Be eS. SOR”. Sens 708. A. speciosissimus Pavl. 8. Leaflets 6-8 pairs; calyx-teeth filiform, 3-4 mm long; corolla 24-25 mm long; pods 30-40 mm long and 4-5 mmbroad.......... 2 oa, AHO, ie ae eae eter Cae id enn. Hane WY 713. A. dianthus Bge. + Calyx-teeth filiform, 5-6 mm long; corolla 25 mm long; plants differing) frommalone rs ia 21h uel “aire mete nee a. ee ee 103. 9. WeatletstGinc) pavysts Oval. Git) et -—acereL een: 721. A. eremobius M. Pop. + “Leafiletisi2-3 (455) pais, (linear 0s" Seema otek Cl. eee mene 10. 10. Pods straight; standard 7-9 mm broad . 720. A. Petunnikovii Litw. + Pods flacate; standard ca. 5mmbroad.... 722. A. rariflorus Ldb. 11. Pods narrow, linear-lanceolate, 2-3 mm broad, spreading or cernuous. WB LEP OL « ew, Dates See A Ree ee ee ee, See, ee 2h + Pods broad, oblong, large, from 3.5-4 to 5-6 mm broad, erect. . 23*. * In some instances, especially in the case of whitish tones of corolla coloring, asin A. pallescens, A. Zingeri, etc., yel'owing of the corolla may be noted in drying, or sometimes a yellow corolla tums blue (A. Petunnikovi). It is therefore necessary to indicate the color of corolla on the spot, at the time of collecting. Usually, however, with experience the yellow color can be distinguished from other colors turned yellow by drying, even in the herbarium (i.e., in dry condition), In case of difficulty, it is advisable to consult both stages of identification, the more so in view of the fact that, in some species, traces of pleochroism appearing as a faint blue tint, may be detected in association with the primary yellow color, especially at the tips of petals and, apparently, with greater frequency in the bud and in the opening flower. In composing the identification key, the the greatest difficulties presented themselves in connection with species described by the authors as fruitless in flowering condition (A, karabilicus, A, reverdattoanus); with all due reservations, these species had to be provisionally incorporated in the key. ** See also A. Reverdattoanus Sumn. on p. 522, the pods of which are unknown. 482 648 649 ++ Pods erect, sessile, oblong-linear, together with the elongated point 12. ifs. oaiiimplonca (Karatan)itio. | 2. es Bees. 678. A. leptopoides M. Pop. Corolla pink; pods glabrous or sparsely hairy, 6-14 mm long... 13. Corolla otherwise colored (white or violet); pods covered with white appressedvhairs orslabrous, up to 5-20 mm long’)... ). . 14. Coxnollagaoa nim dong; pods 13al4yimm@longi sat.) se es ees. SR. Pt ee ee ae teers cake es, MER Sele we 696. A. chorgossicus Lipsky. CGonoilastG min Tong; pods 6-10 murilong!s @2'e). ae |. eee ee ee eer, Nee ee, eens nee nee 697. A. kendyrlyki M. Pop. Wecilersmly ss (4) Dames fee eek Lone oe ent ee ee meee eae By Beatict sum ore muUmenrouls,(Can4 =i epalins) tant mene an cee ee ne Gr: Calyx tRomortOst-Ourmim LONG) 3. cutee ee. eee ale geet meen eter es oe 16. Galyxpinom el cptowa0 mn long. 5 8 Soy secs ieee scen, aos eet ee, ee Peduncles equaling to 1.5 times as long as the leaves............ > ob + oho seni aon at i da aad a ada el eal 707. A. psilophus Schrenk. Pedunclestoe+ times vacmonp as the leaves: ate ees el 2 9 (Bg Pn Nerve gh ti Ei its tell ei adi a ON Pedic ie 705. A. subternatus M. Pavl. Corotlakt)s20526 mmerlong > ser. * 47 701. A. margusaricus Lipsky. Corollastl4-167(£3)immilong. 7 4 2. 702. A. baldshuanicus M. Pop. Soroulaswintereac sO Tam Monga tae, Me Maer) mime en ce, kere terme |e If), Corolla violet of various shades, 14-20 mm long. ........... 20. Conollas22-235smmelong a 6 2 St eee eee 699. A. angreni Lipsky. CGornollag2 923 0smmy long, x as sya ee. 706. A. karabilicus M. Pop. * Standard 20 mm long; keel 16 mm long .. 703. A. Kessleri Trautv. Standard from 14 to 19 mm long; keel from 11 to 13 mm long.... 21. SUA haa deel OTT LONG a hg ef ed Nt Aedes od Bed Goes oe oe 22. Standard 14° min Jones (85% 2 are . & 704. A. pendulinus M. Pop. Calyx covered with black and white hairs. ..698. A. sogdianus Bge. Calyx covered exclusively with white hairs 7). 07.0 7 Sah as .93 7%. EE ee sk a ea eed a ee 700. A. karataviensis. Pavl. Pods enclosed by a double wall, the two layers separated by a spongy HISIGUCT Rea ores GEMS le Se ee 695. A. satteotoichus Gontsch. Pods thin-walled, the wall of normal constitution............ 24. Suffrutescent plant, the lower leaves trifoliate, stipules adnate to petiole, the upper leaves composed of 4-5 leaflets............. ci -b ih OB Aa Rr eM ie nr a Da a 686. A. mugodsharicus Bge. Teaticisus Or more pairs: Stipules free ss: . se aa ee a 20) Stems strongly branched, up to 15-25 (35) cm long, often abbreviated to 1-3 (5) em; racemes short, umbelliform-capitate; pods from 10 Loreopmimelongsand tromve-o-3 to 4 mmebroad sey) = = eee 26. Plante rithe mitten irOmrabOVie wu e ss eee Gye ale ee eet eee) Silks Leaflets large, from 7 to 25 mm long, linear or oblong ....... Zane Leaflets small, from 2 to 5 (7) mm long, oval or oblong-oval ... 32. Bractsrsmally tromy #15 tos mam TOM. ee oe eee li 28. TENE EUS) ETE SEO el el Aull esenalehka or acuinae cidieencec ncn © Somes comic Womcdennolucres the -< 30. Pods oblong, (10) 15-20 mm long and 3-4 mm broad; suffrutescent MlAnUS WAL OW. StGMUSHeren ta etc at. cee ete ewe ee tua bee ceca 78), Pods narrower, 10-20 mm long, 2.5-3 mm broad; herbaceous perennials, acaulescent or nearly so....... 690. A. Ionae Palib. * Pods of this species are unknown, hence their inclusion in the section Sogdianae is conditional. 483 PAS). Standard 17-18 mm, keel 13-14 mm long; calyx 12 mm long, covered with black and white hairs, the tube 2-3 times as long as the teeth (Wie, enemy Shami cnet «dy wea Ae eae. oe 691. A. pskemensis M. Pop. Standard (17) 20-23 mm long; keel (13) 15-18 mm long; calyx 9-11 mm long, covered mainly with white hairs, the tube 4 times as long as the teeth (Transvolga region)...... 688. A. Zingeri Korsh. Calyx-tube 2-3 times as long as the teeth; calyx covered with spreading white hairs (Crimea)......... 687. A. dealbatus Pall. Calyx-tube 4-5 times as long as the teeth; calyx covered with appressed or subappressed hairs (Altai). . 689. A. pycnolobus Bge. Pods elongated, 15=20) mimplonow. yo nd of yds uasusy oo creeen a Se 33. Pods short, LOST tram lomgy ei. lace 693. A. Baranovii M. Pop. Corollan Gy mam) Vomge 15) they ep. cleat acl spllogas 692. A. badamensis M. Pop. Corolla 2 = 22) crate LOM Cie aldactone obi ad cua des dt as 694. A. fissuralis Alex. Low shrubs with developed stems from 20-30 to 90-100 cm long; pods thickened, from 10 to 20 mm long and (3) 4-6 mm broad .. . 34. Herbaceous perennials with usually shorter stems (10-30 and up to 60 cm long); pods narrower, from 10 to 20 mm long and 2-3 mm ROA a ce icecpitlogtst et pe svete Us a cis Cutie os aed taal ae Oita gt 41. Racemes loose, up to 6-14 cm long...... 684. A. brachylobus DC. Racemes capitate or ovaloid, compact (closely flowered though sometimes, elongating in fruit) . 2... 1. atsiye! «payee Sah RR 35. Pedunelesrequalingwthenleaves .... 5. «4 Ris eRe ene lea ee eee 36. Peduncles 1.5-3 times as long as the leaves............... ae. Pod covered with short ascending hairs; bracts 3-5 mm long ..... OT SER ee aE en nn MUN ABE 6% | ¢ 679. A. cornutus Pall. Pod densely, white-villous; bracts 1-2 mm: long? s..9: 4! Sauer seepage Morini! Bhs. WER. aus cama het ia elietinne cbyicg th bE 681. A. fruticosus Pall. BractsySiak 2 mam: LO py) eedjscen oh HG ai yp sori qa Saey oh eens eee 38. Bractsfromel to. mimglon gen.) ayes, pice hehe cal 4 dae 40. Bracts 5=8mmelongeiy.o) cet wid. 2 lowe ehdvek aad te ete 39. Bractsedil sl) ammoy LOM QE 8, viyray | uae a 684. A. Komarovii Lipsky. Leaflets 3-5 pairs; all flower parts large (calyx 17-20 mm long, standard 23-27 mm, keel 20-23 mm long). 682. A. iskanderi Lipsky. Leaflets 6-11 pairs; flower parts much smaller (calyx 10-13 mm, standard,.1.9-24.mm,. keel. ll-14 mm long) 4) ee ee andyS O54. my DR OAGG a cndsihhh his push co) a bowndineripeas 681. A. fruticosus Pall. Calyx 15 mm long, the teeth 3-4 mm long; pods 15-20 mm long and SGM ROAM ete este cor a Tear «te ek” agate 685. A. viridis Bge. Corolla white (darkeningin drying), (Crimea)... ». 0.9 eae eee Me) ible gait nosh beagatebiaenlt Wa aiaie, MslnGstalen.a'r/ guia el pant Nght Sani 719. A. pallescens M.B. Corolla ‘violet; other regionsy.08) 20.)).) Ao Ue apo ca: B) eran ee 42. Plants densely leafy; leaflets filiform; stems of the year covered with white hairs, 5-7 cm long; peduncles long, 2-3 times the length of the leaves; pods narrow, 15-20 mm long, 2.5-3 mm broad..... a alle alten ar thats ch ncaa tin ouvoy ects nici Bed AN) Succes 717. A. kustanaicus M. Pop. Plants).differing From AWOVES aie, wiley fobiny cerns. sey iereinlaeey cash elas aie enna 43. Racemes short (4-8 cm long, elongating in fruit); peduncles 2-3 times the length of the leaves; corolla pale violet, 20-23 mm long; pods LOSS mm long and 3 mim) broad ue. au 718. A. macropus Bge. 4 837, 1117 484 651 652 53. 04. Racemes elongated (8-18 cm long); peduncles equaling to 1.5 times as long as the leaves; corolla violet, 16-20 mm long; pods 13-19 (29) mam long and 242)5) (3) mm broad tia ap-ane 716. A. virgatus Pall. Corolla yellows) ceyot one LL Week iat Me OS rN ee eee 45. Corolla otherwise colored, not yellow (see above on p. 646)..... 62. Shamu wart Oxo) Gmamba Les ros sie tee ae rm Mere mis A abt) Evieee et 46. Herbaceous perennials or low subshrubs with short stems...... 47. Corolla 21-23 mm long (Kopet Dagh)... 629. A. nigriceps M. Pop. Corolla 28-30 mm long (W. Tien Shan). 628. A. Spryginii M. Pop. Flowering racemes short, umbellate to capitate ...... AE tence 48. Flowering racemes loose, elongated, remotely flowered ...... 51. Peduniclessequaling omisiichtly exceeding thevleaves (jij). .6.- . . RRR tne aoa teas. ae tel edly 650. A. Neilreichianus Fr. et Sint. Peduncles (1.5) 2-4 times as long as the leaves (see also p. 85, Aveo cmievenumierivctyaiy Sm WIE RROD iyi oo ae eee ah es tuntnee eile! \ellete ter se, yn se 49, Leaflets 3-4 pairs, 10-20 mm long; pods relatively short (25-30 cm long and 2-3 mm broad), curved... 652. A. dsharkenticus M. Pop. Leaflets (4) 5-8 pairs, ca. 10 mm long; pods long (from 30 to 65 mm Konorand oc2 0 mam bcoad),(olabrOUS) .) -\-)2) m1 - meheneeh ree te nets 50. Bracts Ca.) 2 mim longs ipods) l5immubroad (Shusnan\iix. 26 ualie, . 4 RE AE cnthosne RLM. &. Aekpoeet Re as 651. A. stenoceroides Boriss. Bracts 2.5-3 mm broad; pods 2-2.5 mm broad (Kopet Dagh)...... a SSG AL dad fee) RS SO | ee ene er Pe 649. A. sumbari M. Pop. Rods) imciteirmm longyybracts O77 mim Lome Aten ys ii) eh k ceauiee clean 202 Er diet esti ah A. (ORO NT, THES, Say ..... 643. A. subexcedens Gontsch. Podstlongers! bracts shorter CMO O7 Sy. sa eine 34 eben ios ah ses Ueaneen tte BAe Leaflets very narrow and elongated, up to 30 mm long, and 1 mm broad; ecorolla,.25-28'mm long; pods up to 60/mm long. “a Aes. . BPW ide aed dh i te mm, ee toe hot 644. A. transhyrcanus M. Pop. Pa aneSaelitfe ringer Onda AWOVeS) ..¢ PRL i Mee MU UCihe eta tees eile. Seb Ra een sees ide 53. Pedunecles shorter than or equaling the leaves......:....... 54. RPedunclesvlioa;s times as long asi themeaves cis. ih ee wee 58. Galyxatube uwicelas, longuas thexteet hi) Ais \- MER. amity. 827A Wa 3s VATE RA aE RY ee ee 645. A. xiphidioides Fr. et Sint. Calyx-tube 4-10 (or more) times as long asthe teeth......... 55. Leaves 1.5-2 times as long as the peduncles; calyx-tube 9-10 times as long as the teeth; calyx 10-11 mm long; corolla 20-25 mm long; POisetOn oO) Malm sOM Sele elie el eee 647. A. argyroides G. Beck. Calyx 6-7 mm long, the tube 4-6 times as long as the teeth; corolla 15-18 mm long; pods 20-22 mm long... .648. A. Sachokianus Sosn. Blanisidiitecenttromitabowe Ast. ! seas eid. RG ak es Ott sk Mee 56. Leaflets faintly hairy or glabrous above, green..........+... oT. Meatlels appressed hairy onvbothiside's! 2 pe . feos -ms ey hoists odie tt) (oy te MRIS rer sense eo ee (minis: tense) uronic. teh ove 645. A. xiphidioides Fr. et Sint. Leaflets glabrous above; corolla 18-22 mm long.............. 94). Gash Ape Sakae’ Misa Cane a 673. A. ucrainicus M. Pop. et M. Klok. Leaflets faintly hairy above; corolla 25-27 mm long............ Suicli .N ne Eee ae a ae et 641. A. lancifolius Gontsch. Leaflets 2-3 pairs; stipules 1-2 mm long. 642. A. exilis A. Kor. Leaflets 4-9 pairs; stipules 3-5 mm long ...........+..... 59. Pods on a stipe 4-5 mm long ...... 639. A. Lorinserianus Freyn. Pods sessile or subsessile, the inconspicuous stipe ca. 1 mm long. PUPPET CON NSN let Mr el ng SS A el AH Saeed, Nadas sok gee tay ok Jain gl ns mga agay Mol ae va BeaeE SYN 60. 60.a0cBractsi lommglonga(SyneD:44W.. (Panut-vAl D> 6) etka s eae ee PPh wlohe. umceyl paces oa! etek. § 640. A. maverranagri M. Pop. +e Bractspins =2ymmi long. (KopetpDagh) mat. (feo. 4 oe Sere Bee 61. 61. Corolla 21-24 mm long; calyx 13-15mmlong; peduncles equaling to 1.5 times as long as the leaves... 645. A. xiphidioides Fr. et Sint. + Corolla 21 mm long; calyx 11-13 mm long; peduncles twice as long as the: leaves) tine. zeke 646. A. xanthoxiphidium Fr. et Sint. 62. Pods more or less spinescently curved, cernuous........... 63. + Pods straight or slightly curved, cernuous or obliquely ascending. 65* 63. Leaflets 3-5 pairs, very narrow, 5-15 mm long and 0.5-2 mm broad; corolla 18-20 mm long; pods narrow (1 mm broad) ............ » Medacahett. TORI Meee ae saxe wing 636. A. falcigerus M. Pop. + Leaflets 1-3 pairs, broader; corolla large; pods 2-3 mm broad. 64. 64. Calyx 10-11 mm long; leaflets 15-25 mm long, 4-10 mm broad. i danat: vata bie, on G4 hse wopcedd a Cadas ee tet 637. A. aulieatensis M. Pop. + Calyx 12-16 mm long; leaflets 5-15 mm long and 2-3 mm broad ... 5 pe Re OR sll ok OR «are ys, BI CE het 638. A. scleroxylon Bge. 65. Herbaceous perennials or subshrubs, with short usually ascending or decumbent, stems ./...... RROSARTS .. tosou Pi €.5"%¢. . See eee 66. + Shrubs or subshrubs with long developed stems............. 88. 66. Leaflets 2-3 pairs (see also A. compressus Ldb., p. 692)... 67. + , Leaflets; manyopairs (455 torl0=N3 pains) =beast! sive Ye. 8 eee 68. Gime CorolagaOpmam On Ce Rae aces oe ee els 675. A. sogotensis Lipsky. +. Corolla ol 4=Voynam ong, segs sea) oe 676. A. pischtovensis Gontsch. 68. Plants with many (up to 100) strongly developed stems (subpulvinate); leaflets 10-13 pairs, small (5-8 mm long and 1-1.5 mm broad); corolla pale pink, 15-17 mm long...... 677. A. Michaelis Boriss. + Plants not subpulvinate; leaflets 4-6 to 9-10 pairs; corolla from PUG al eomaGemimmelOne. BBE oe eos ie be vl cus qe ee 69. 553 69. Flowering racemes elongated, loose, remotely flowered; leaflets very narrow, filiform-linear, from 0.5 to 1 (1.5-2) mm broad... 70. + Flowering racemes short, subumbellate to capitate, 1.5-3 (5) cm long, or, if loose and elongated, then leaflets broad (from 2-3 to Spmunt bio ad) ppart behets toys PERS ces ree cates cde oo An: hcg ee 70. Corolla lurid (apparently tinged with other colors).............. want etpvtee: : & WG. 8 6 acy) 26729 A: isubulatusy M. Be (see also A. ucrainicus M. Pop. et M. Klok., p. 700). + e@orollaxdifierentiy, colored «sq. j.0\-l\. bt yen Lovee Bete Sie eee brs 71. Calyx 8-19 mm long, the teeth 1-1.5 mm long; corolla purple..... ; fe wars G ety: : BR cui Oe de ST) § 672. A. subulatus M.B. + Calyx 11-13 mm long, the teeth 1.5-2 mm long; corolla white or Violetiie chide. daacte ets ae 674. A. pseudotataricus (DC.) M. Pop. 72. Racemes elongazed: up to 6-10 cm long, loose, remotely flowered. ia Ne RR ey Mi BAL eg ge ee wae ee. 6 ee Eee Sy + Racemes short, subumbellate to capitate, from 1.5-2 to 3-4 (5) cm lone. 2g. <.sde eM. pNP BSB. ola ls cae © ap oie Alec ee 74. 73. Leaflets 5-7 mm long, 2-3 mm broad; corolla 20-24 mm long (Tien Shan) ealiatearel A INS oe eg 670. A. Reverdattoanus Sumn. + Leaflets (7) 10-20 mm long and 2-3 (5) mm broad; corolla 17-19 mm long (Daighestan)) ) ac aaah: even ',..669. A. haesitabundus Lipsky. * See also A. Reverdattoanus Sumn. onp. 522, the pods of which are unknown. 486 74. 84. 85. 86. Sit. Pod lanate with white spreading hairs, relatively broad (27 mm long, 3.5 mm broad); peduncles somewhat exceeding the leaves ....... TR es aie lay ere A fatal ak ce SEN 662. A. chaetolobus Bge. Pilantspitterentiiroml above. 4)... 008 Jeet. ie Ua oes Oe DR OR ws 75. Leaflets broad, not more than 2-3 times as long as broad...... 76. Leaflets narrow, 3-4 and more times as long as broad........ the Stipules 2.5-3 mm long; calyx-tube 4 times as long as the teeth... . COM Ua NAMES vs col cls tocttame 2 bain Fyn a tebe ae haa 654. A. amabilis M. Pop. Stipules 1-1.5 mm long; calyx-tube twice as long astheteeth..... eer amt Av btial S A) cet wey’ ba. 4 oe waged meee 668. A. artvinensis M. Pop. CorollayZ6-28immulong:) awings) 29527. mam Loney tis = aioe. loads |. Rane Peters, .OIBMONE PE inom: 656. A. ortholobiformis Sumn. Corolla not more than 24-25 mm long and, if longer, then wings IAAT OM OS, Ae. : BALE) An aT eee, ses Ee eet Be 78. Peduncles equaling or not more than 1.5 times the length of the leaves bec blo 4 by OUss RANen tol ee Me ae OG EN els keen BOA Rep else Lek d As eB 79. Peduncles (1.5) 2-3 times the length of the leaves ........... 84. Corollaginon: 6) toz22 mm: longoria. Ps. 278) seas .. ak GIS A oe, wee 80. Goroblay22-2ammplongy. ps Biel... ee.) Lopes) eR, ee Bee 82. Leaflets very narrow, 5-10 mm long and 0.6-1 mm broad; racemes ioe 2hocemmlone) armliusbans ea eu. § ad 671. A. corniculatus M.B. Leaflets 5-15 mm long and 1-4 mm broad; racemes 2-5 cm long. 81. Flowering racemes 2-2.5 cm long, 3-6-flowered; wings entire .... eran’ ff eal cilin ly. ere Ane 657. A. angarensis Turcz. Flowering racemes 4-5 cm long, 7-15-flowered; wings retuse...... p> oh a ATA te Wolke: hee a, eae Sane oh: 655. A. macroceras C.A.M. Corolla pink, sometimes suffused with green or yeilow.......... Me MPU ERS STAGE os te kG. bs Cae BLUE. TEE. 653. A. macrotropis Bge. Gorolllaywhitash 2.00)... 6.5 Be eo. Gee | aes. Sel eee 83. Stems of the year 15-20 cm tall; leaflets 8-18 mm long and 2-4 mm bod standard Zo mmo long andiica.ismm' broad! >. .71.e-earee ee PRI ie Siibek as as do 2B ss i AWN ES Gr) AD Ch SENS: I CNS 658. A. ortholobus Bge. Stems of the year 2.5 cm long; leaflets 5 mm long and 1.5 mm broad; standard 22-25 mm long and 9mm broad. 660. A. tephrolobus Bge. Wer-ollavnalenvAOlet eye Fh MN is) be yp eens Speier ey REN uw eae SPU 85. Conollagilaespursple sacks oyh es acti cores DR OA Ge. SM. TE BA 86. Calyx 10-12 mm long; racemes 2-4 (5)-flowered (Zeravshan)..... BPG ee ee ee ut REE S Ey see) Dr 664. A. polytimeticus M. Pop. Calyx 7-9 mm long; racemes 5-10-flowered (Centr. and S. Ural). Dae MOE AGA ee SPR NENE IER oo PEO BITES. STO. 667. A. Karelinianus M. Pop. Calyx 9-10 mm long; racemes many-flowered, containing (5) 8-14 flowers (Kungei Ala Tau)......... 659. A. karkarensis M. Pop. Calyx 6-7 mm long; stems and leaves rather sparsely hairy, green. Se RRE ANT Re Ch ete uc eae i eT SL Es Pa 663. A. ceratoides M.B. Calyx 8-10 mm long; stems and leaves whitish by dense hairiness. Fe TON Trae NES AR eas PTS RTP yet Re Tate SL OR AS i a Sele Ser ae 87. Leaflets 5-7 pairs; pods ca. 25mm long, 2mmbroad ......... AP ees MECH IET Es: Pah O, erdiio rere la 665. A. stenoceras C.A.M. Leaflets 8-11 pairs; pods ca. 20 mm long and ca. 1.5 mm broad... RM AES CES Aik, SN Re eM eed Fae 666. A. substenoceras Boriss. Leaflets commonly 3 pairs; pods 21-28 mm long, 1.5-3 mm broad. 2 ill Ce ae eC LED. mT Yael so ae 661. A. compressus Ldb. 487 88. 93). 94. Pods from 15-20 to 30 (38) mm long and 1.5-2.5 mm broad; leaflets 2-4 (5-6) pairs; corolla violet or purple (steppes and semideserts of (ORT oe AMET an ey Bled MMA Sia sham gyal A AR a aM Umea A, AORN egw 89. Pods 35-40 to 100-120 mm long and 1.5-2 (3) mm broad; leaflets 4-6 to 9 pairs; corolla commonly violet, tinged with other colors (pleochroism) (mountains of Central Asia and partly Transcaucasia). eee tS ee re rh ane tet Be obs UM bin ar Wey gas AN TERE ps Paty wes eek aaa Klowemnrusacemessremotely flowered)... 2/2). 2 9 a.s) lay a) eee EPR ARR eo? ET ae Tidy: taceu yO ee ode Gn ta 634. A. polyceras Kar. et Kir. Bileweringiracemies Shortepeanitate (a) et aus wi, aus gos say eee 90. Calyx-teeth ca. 171.2 mmfPlong; Ypeds straight i! oer Bae eo. eee 1). Calyx-teeth 1.5-2 mm long; pods somewhat arcuate, sometimes some of them/straight and thenjealyxal0ql? mmploney. Bene Beri. a. dee om. Leaflets 2-3 pairs, oblong-obovate to oblong; corolla 18-19 mm long. aces! afi o atetol mitt os ata. 2 cat 632. A. tscharynensis M. Pop. Leaflets 4-5 pairs, lanceolate or linear; corolla 15-18 mm long. BE oo lela fe be oe oy dy 5 ORL AS Obes GE 631. A. temirensis M. Pop. Plants 50-120 cm tall; leaflets 2-4 pairs; peduncles 2-3 times as long as the leaves (plains of Centr. Asia and W. Siberia) ........ peters. Riese wee Da ee wrtot fee OF 630. A. arbuscula Pall. Plants 20-40 cm tall; leaflets 2-3 pairs; peduncles 3-5 times as long as the leaves’ (valleys: of*Centr:) TieniShan)in 4 92a). iy. 12 0 eee vo hoe, aR ELISE Nie even tey iy aE Ri Fo Warned g 633. A. duanensis Saposhn. Plants up to 1m tall; leaflets 5-6 pairs; peduncles 1.5 times as long as the leaves (mountain meadows of Tien Shan and Tarbagatai)..... A A -rgh OPTI LPN. MN eee ons 635. A. Gontscharovii Vass. Pods from (50) 70 to 110 mm long and 1.5-2 mmbroad........ 94. Pods from 20-30 to 40 (55) mm long and (1.5) 2-3 mm broad... . 96. Racemes) loose; ‘elongated,, 8510 em long.) . 2... >) oR eee VIR a IR SUMO REEL Od ee ee ete 626. A. ischnocarpus Gontsch. Racemes short, closely flowered, up to 3-6 cm long ..:.... 5). 95. Standard entire at apex; pods sessile (Shugnan) .............. Bete syltey AS GA. usin. Bee et he Le 627. A. acanthocarpus Boriss. Standard retuse; pods borne on a stipe ca. 2 mm long (W. Tien Shan). 7) Sok ia tate an Re dk ta Raa Ae band 625. A. Fedtschenkoanus Lipsky. Goerollafrom (16 to 1s mimalong > py Ye i se SAE oe ea Sill 8 Corolla ca. 21 mm long; pods horizontally spreading, villous with ascending hairs, disposed in a-long loose raceme ............. oe SELLA ot ee "EP BOEY «Maloy aunt te Q) EA gs 623. A. surchobi Gontsch. Corolla from 20 to 25 mm long (or longer); plants different from abovek lr ‘yais.osis. at tweed Reig ety. etree leon Sys ery dl Scena 98. (see also A. karabilicus M. Pop., with corolla 26-30 mm long, p. 727) Leaflets linear, 8-20 mm long....... 622. A. kabadianus Lipsky. Leaflets oval or oblong-oval, 5-12 (15) mm long and. 2.5-5 (8) mm DROAGSTIALEN AS8ob. VG. Bal tina, eel. 621. A. cisdarvasicus Gontsch. Limb of standard rhomboid, enlarged about the middle, attenuate toward base and toward apex ........ 616. A. variegatus Franch. Limb of standard obovate or Sci sak Base not attenuate toward Peduncles shorter than to equaling the leaves ei. ec eres eee 100. Peduncles 1.5-2.5 times as long as the leaves............. MO. 488 656 1657 100. Racemes 4-6 cm long, rounded-ovaloid; calyx pannose with black amciavinate Naame thesteethr2 .om3) may VOM gee veut ieaeeteeu eer) ini trl. 6)8 pln TOR Re IRE RRR Ne 617. A. neo-Lipskyanus M. Pop. + Racemes loose, 6-14 cm long, remotely flowered; calyx covered withvappressed white hairs, the teeth4>4 mm long ..... . wn a. a: 3 Bleaiod inl tote RR lee DOT Re Oe 1 ES NEN 618. A. leucocalyx M. Pop. 101. Leaflets 10-20 mm long and 1-2 mm broad, covered on both sides with appressed white hairs; bracts ca. 2 mm long; corolla whitish- Poi ERY! (a = C2) ch MMS SMR a Se | CR 624. A. xiphidium Bge. + Leaflets 5-10 (12) mm long and 1-3 mm broad, hairy sparsely above and densely beneath; bracts 3-6 mm long; corolla dingy pale violet WERNER) a Sea nen A ppMnre meat > GMM ny Coster (ia WAL ey Wert a AMAIA a! 102). 102. Standard 23-25 mm long; keel 21-24 mm long; pods 25-35 mm long and 1.5-2 mm broad, suborbicular in cross section, gradually attenuate into a beak... sw Jncerd wvs! we 620. A. Kudrjaschovii A. Kor. + Standard 21-23 mmlong; keel18-20 mm long; pods (1.5) 20-30 mm long and 2.5-3 mm broad, cordate-orbicular in cross-section, abruptly iSy2iau ky SY Uehara ai ae Amr eee Pile men LANE PaO ea 619. A. urguticus Lipsky. 103. Leaflets 3-5 pairs; flowers 25-30 mm long; calyx 12-16 mm long. 2/91 pedal estrada aber ag 1 lame See irae LL Spit let bet ry bie 104, + Leaflets 7-9 pairs; flowers 33-36 mm long; calyx 20 mm long... eT on es wh bing clive ane oraperrane. ie al oe 714. A. Knorringianus Boriss. 104. A dwarf subshrub; calyx-teeth 5-6 mm long; corolla 25 mm long; pods ca. 20 mm long, 3-3.5 mm broad (Karatau Range) ........ MEW ee cook ccles) eye eka putea eu es Rha wise Aes 715. A. Mokeevae M. Pop. + Herbaceous plant, becoming woody at base; calyx teeth 1.5 (2) mm long; corolla 26-30 mm long; pods 20-25 mm long, ca. 3 mm broad REMAN ATA Nee et tee gtk ee MAR en 713. A. dianthoides Boriss. Series 1. VARIEGATI Vass. — Frutescent or tall suffrutescent plants; leaflets elongated, many pairs; racemes loose, sometimes abbreviated; corolla of middle size (from 16 to 25 mm long), violet suffused with other colors (yellowish-violet, dingy violet, etc. pleochroism). Pods sessile, subulate-linear, from (20) 30 to 40-55 mm long and (1.5) 2-3 mm broad, more or less spreading. — Representatives of the series are distributed through the midmountain zone of Central Asia. 616. A. variegatus Franch. in Ann. Sc. Nat. ser. VI, XV, (1883) 259. A. sarypulensis B. Fedtsch. in Trudy Tadzhik. bazy Akad. Nauk. II, (1936) 150. Perennial, suffrutescent, 50-70 cm tall, the fairly tall caudex covered with brownish or brownish-gray bark; stems of the year 2-6 cm long, densely covered with appressed black and white hairs; stipules triangular or lanceolate, 3-5 mm long, herbaceous, covered with appressed black and white appressed hairs; leaves 4-9 mm long, the very short petiole and the rachis covered with scattered appressed white hairs; leaflets 6-10 pairs, oblong or elliptic, obtuse and minutely mucronulate, covered on both sides, more densely beneath, with appressed white hairs, greenish above; peduncles 5-12 cm long, rather sparsely covered with appressed white hairs; racemes many-flowered, rather compact, mostly ovoid, 2-4 cm long; bracts oblong-lanceolate, 3-4 mm long, covered with black and white hairs, exceeding the pedicel; calyx cylindric, 11-16 mm long, rather sparsely covered with spreading white and appressed black hairs, 489 658 the tube 3-4 times the length of the subulate slender teeth; coroVla violet; standard 20-23 mm long, the limb rhomboidally oblong-ovate, rounded about the middle, broadly ligulate upward, obtuse, short-acuminate in lower part, angular at base, 2.5-3 times as long as the claw; wing-petals 16-19 mm long, the limb linear-oblong. obtuse or slightly retuse, 7-9 mm long; keel 12-17 mm long, the limb inequilaterally oval, 6-7 mm long; pods sessile, subcernuous, linear, slightly curved or straight, 2.2-4 cm long, ca. 2.5 mm broad, attenuate into a beak 2-5 mm long, strongly carinate ventrally, deeply and broadly grooved dorsally, subtriquetrous, coriaceous, rather sparsely covered with appressedblack hairs interspersed with isolated appressed white hairs, subbilocular, many-seeded. Fl. May- June; fr. June-July. (Plate XLII, Figure 2). More or less gravelly or stony slopes in the intermediate mountain zone and in low mountains. — Centr. Asia: T. Sh., Pam.-Al. Endemic. Described from Dzhadzhik (W. Zeravshan). Type in Paris. 617. A. neo~Lipskyanus M. Pop. in Sched. ad Herb. Fl. As. Med. Fasc. XV (1928) 113.— A. Lipskyanus M. Pop. l.c. 24, non Freyn (1904). — Exs.: H.F. A.M: No. 358. A frutescent plant 35-50 cmtall; caudexthick, upto 8 mm indiameter, often weakly branched, erect, 10-25cm tall, covered with longitudinally spitting castaneous~brown bark; branches of the year 5-15cm long, densely covered with appressed white hairs, adnate to petiole in lower part, triangular-ovate, the upper ones suboblong, acuminate or acutish, 2-3 mm long, covered with appressed white hairs; leaves (2) 3-6 cm long, the petiole and the much longer rachis slender, covered withappressed white or rarely mixed white and black hairs; leaflets6-7 pairs, lanceolate or oblong-oval, rarely linear-lanceolate, acute, rarely obtuse, (2) 3-10 mm long, (1) 2-4 mm broad, rather densely covered beneath and thinly above with appressed hairs or more often glabrous above; peduncles equaling the leaves, covered with appressed white and upward mixed white and black hairs; racemes short, rather loosely capitate (sometimes the fruiting racemes fairly loose), 4-6cm long, rounded-ovaloid or subspherical, 10-20-flowered; bracts narrowly lanceolate to linear-lanceolate, 2-2.5 mm long, covered with spreading black hairs interspersed with fewer white or pure black hairs; calyx cylindric, (11) 12-14 mm long, densely tomentulose with spreading black hairs and villous with longer white hairs, the teeth linear~-subulate, 2.5-3 mm long, the tube 4 times the length of the teeth; corolla whitish, with violet standard; standard 20-21 mm long, the limb obovate-orbicular, - rounded-obtuse, not attenuate, slightly angular at base, 2-3 times as long as the claw; wing-petals 17-18 mm long, the limb obovate, ca. 4 mm broad, entire, half as long as the claw; keel 16-17 mm long, the limb semicircular, acute, half the length of the claw; pods subsessile, on a stipe ca. 0.5 mm long, gathered in a rather compact short raceme, spreading horizontally or ascending, the lower subcernuous, subulate-linear, straight or barely curved, (3.5) 4-5.5 cm long, 2-2.5 mm broad, 2.5-3 mm thick, carinate ventrally, broadly grooved dorsally, trihedral in cross-section, acuminate to a straight subulate beak 1-3 mm long, coriaceous, densely covered with appressed white hairs, bilocular. Fl. May (June); fr. (May) June. Fine-grained and stony fine-grained steppe soils or meadow-steppe foothill slopes.— Centr. Asia: Tien Shan (W.: from Dzhambul district and Karatau mountain to Mt. Korzhantau). Endemic. Described from Ak-tash river valley in Korzhantau mountains. Type in Tashkent. 490 659 618. A. leucocalyx M. Pop. in Bull. Univ. As. Centr. XIV (1926) 140.— Exs.: H.F.A.M. No. 357. Perennial, suffrutescent, up to 50 cm tall; woody caudex up to 20 cm tall, fairly stout (up to 7 mm in diam.), covered with more or less peeling grayish-brown bark; stems of the year 6-15 cm long, firm, white, angled, densely covered with appressed white hairs; stipules adnate to petiole up to the middle, triangular, the upper lanceolate, acute, 3-5 mm long, covered with appressed white hairs; leaves 6-10 cm long, the petiole 1.5-2 times as long as the rachis, both petiole and rachis densely canescent by appressed white hairs; leaflets 5-6 pairs, linear, subacute, 7-15 mm long, 1-2mm broad, densely covered on both sides with short appressed white hairs, hoary; peduncles 2/3 as long as to equaling the leaves, densely covered with appressed white and in inflorescence sometimes mixed black and white hairs, hoary, (41) 5-8 cm long; racemes 6-14 cm long, loosely 10-17-flowered, remotely flowered and fruited, the flowers horizontally spreading; bracts lanceolate, acute, 13-17 mm long, by a whitish densely coat of short appressed hairs, the teeth narrowly linear, 4-5 mm long; corolla impure white; standard 21-22 mm long, the limb broadly obovate, emarginate, slightly auriculate-angular at base as long as the claw; wing- petals equaling the standard, the limb oblong-obovate, entire, the claw 1.5-2 times as long as the limb; keel ca. 20 mm long, the limb semicircular, acute, half as long as the claw; ovary ona stipe 2-3 mm long, white-hairy; pods borne on a thick stipe 2-5 mm long, spreading to subcernuous, narrowly linear, 4.5-5.5 cm long, 2 mm broad, subterete, prominently sutured dorsally and ventrally, acuminate to a straight stout beak 7-10 mm long, coriaceous, densely covered with appressed white hairs, hoary, unilocular, Fl. May-June (July); fr. June-July. Gypseous stone- and clay-covered slopes of foothills in the ephemeral short-grass vegetation associated with xerophytic tall grasses. — Centr. Asia: Tien Shan. Endemic. Described from W. Tien Shan. Type in Tashkent. 619. A. urgutinus Lipsky in A.H.P. XXVI (1907) 65.— Exs.: H.F.A.M. NOP Aiea oa lCay Els Padzhik..sVi,, tabi)9 1). Perennial, suffrutescent, 30-70 cm tall, the fairly thick caudex 10-30 cm long, covered with grayish bark; stems of the year 10-34 cm long, densely covered with short appressed hairs, slightly sulcate and angled; stipulesfree, triangular-lanceolate, acuminate, (2.5) 4-5mm long, covered with appressed white hairs; leaves 6-10 cm long, the petiole andthe much longer rachis covered with appressed white hairs; leaflets 5-9pairs, linear, (5) 7-10 mm long, 1.5-2 (2.5) mm broad, rounded-obtuse, covered sparsely above and denselybeneath with appressed hairs; peduncles 2-2.5times the lengthof the leaves, covered with appressed or ascending white and beneath the inflorescence mixed white and black hairs; racemes oblong, many-flowered, elongating to 7-12 (18) cm in fruit, the flowers subhorizontally spreading; bracts lanceolate to ovate-lanceolate, acuminate, 3-4 mm long, barely exceeding the pedicel, covered with black and white hairs; calyx cylindric, ca. 15 mm long, shaggy with very short black and long white spreading hairs, the teeth subulate-linear, 3-5 mm long, the tube 2-3 times the length of the teeth; corolla dingy whitish-violet; standard 21-23 mm long, the limb obovate, retuse, slightly angular at base, 1.5-2 times as long as the claw; wing-petals equaling the standard, the limb oblong-obovate, entire, half as long as the claw; keel 18-20 mm long, the limb semicircular, acutish, 491 half as long as the claw; pods sessile, linear, (1.5) 2-3 cm long, 2.5-3 mm broad, carinate ventrally, slightly canaliculate dorsally, subtrihedal, straight or barely curved, densely covered with appressed or subappressed ascending white hairs, these sometimes interspersed with scattered black hairs, abruptly contracted into a straight rigid beak 6-12 mm long, coriaceous, bilocular. Fl. May-June (July); fr. June-July. Mountain slopes in the steppe and juniper zones.— Centr. Asia: Pam.- Al. Endemic. Described from Ugrut. Type in Leningrad. 620. A. Kudrjaschovii A. Kor. in Fl. Tadszik. V (1937) 677. 1 Perennial, suffrutescent, 30-45 cm tall, the caudex 10-25 cm tall, covered with castaneous-brown bark; stems of the year 5-10 (18) cm long, finely sulcate, densely white-tomentose by appressed hairs; stipules free, triangular-lanceolate, 25-3 mm long, covered with appressed white hairs; leaves (3) 4-7 cm long, the petiole and the much longer rachis hoary by appressed white hairs; leaflets (4) 5-6 (7) pairs, linear, (4) 6-12 mm long, 1-1.5 (1-3) mm broad, obtuse, covered sparsely or scatteredly above and densely beneath with appressed hairs; peduncles twice as long as the leaves, densely covered with appressed white and, beneath the inflorescence, mixed white and black hairs; racemes loose, oblong, rather many-flowered, 10-18 cm long in fruit, subremotely flowered, the distant fruits horizontally spreading to subcernuous; bracts lanceolate, 3-5 (6) mm long, covered with black and white hairs, twice the length of the pedicel; calyx cylindric, 15-16 mm long, villous with very short black and predominantly white spreading hairs, the subulate-linear teeth 3-4 mm long, the tube 3-3.5times as long as the teeth; corolla dingy pale violet; standard 23-25 mm long, the limb rounded-obovate, subentire, scarcely angular at base, 1.5 times as long as the claw; wing-petals equaling the standard, the limb obovate, entire, half as long as the claw; keel 21-24 mm long, the limb semicircular, acutish, half as long as the claw; pods sessile, narrowly linear, 2.5-3.5 cm long, 1.5-2 mm broad, gradually attenuate to a straight beak 10-15 mm long, subterete, scarcely carinate ventrally, scarcely canaliculate dorsally, straight or scarcely curved, coriaceous, villous with short ascending white and fewer black hairs, bilocular. Fl. May-June; fr. (May) June-July. Centr. Asia: Pam.-Al. Endemic. Described from Shakhrisyabsa. Type in Leningrad. 621. A. cisdarvasicus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). ; A tall suffrutescent plant, 50-90 cm tall, the caudices 20-35 cm long,- rather stout, covered with castaneous-brown bark; stems of the year 5-18 mm [?] long, whitish by a dense coat of appressed white hairs; stipules free, triangular-lanceolate, 3-4 mm long, covered with appressed white and black or rarely almost pure white hairs; leaves 5-9 cm long, the petiole and the much longer rachis loosely covered with appressed hairs; peduncles twice as long as the leaves, covered with appressed white and in inflorescence mixed white and black hairs; racemes short, loosish or loose, rather closely flowered, mostly ovoid, 4-6 cm long in fruit, rather many-flowered; bracts lanceolate or linear-lanceolate, 2-2.5 mm long, covered with black hairs sometimes interspersed with few white ones; calyx cylindric, 9-11 mm long, villous with short spreading black and white hairs, the subulate-linear teeth ca. 2 mm long, the tube 4 times as long as the teeth; corolla dingy lilac-violet; standard 16-17 mm long, the 492 661 limb broadly obovate, scarcely retuse, short-truncate and angular at base, 2-2.5 times as long as the claw; wing-petals 15-16.5 mm long, the limb lance-oblong, rarely obovate-oblong, round-tipped, as long as the claw; keel 12-14 mm long, the limb semioval, acutish, 2/3 as long as to somewhat shorter than the claw; pods sessile, subapproximate, linear, 2-3.5 (4) cm long, 1.5-2 mm broad, gradually attenuate to a beak 3-6 mm long, rounded ventrally, slightly canaliculate dorsally, coriaceous, covered with short subappressed white and black hairs, bilocular. Fl. May; fr. May-June (July). Northern slopes covered with fine-grained soil, in the low and intermediate mountain zone, in the upper part of the ephemeral and xerophytic wood and scrub vegetation zone and in the lower part of the deciduous wood and scrub zone.— Centr. Asia: Pam.- Al. Endemic. Described from Vakhsh River (Tadzhikistan). Type in Tashkent. 622. A. kabadianus; Lipsky in A.H. P. XXVI (1907) 161.— Exs. : iM. Faas MNO. 3:56. Perennial, suffrutescent, 30-90 cm tall; caudices 10-30 cm tall, covered with brownish-gray bark; stems of the year 5-25 cm long, whitish by a dense coat of short appressed hairs; stipules free, triangular to lance-triangular, 3-4 mm long, covered with white or rarely white interspersed with fewer black hairs; leaves 4-10 cm long, the petiole and the much longer rachis covered with appressed white hairs; leaflets 6-10 pairs, linear, 8-15 (20) mm long, obtuse, glabrous or sparsely hairy above, hoary beneath by appressed white hairs; peduncles 1.5-2.5 times at long as the leaves, covered with appressed white hairs; these interspersed beneath the inflorescence with black hairs; racemes loose, ovoid, rarely oblong, the pods subapproximate; bracts ovoid-lanceolate to lanceolate, 2-3 mm long, covered with white and black hairs; calyx cylindric, 10-12mm long, densely villous with spreading white and mostly few short spreading black hairs, the tube 4 times as long as the linear-subulate teeth; corolla dingy yellowish-violet; standard 17-18 mm long, the limb oblong-obovate, scarcely retuse, angularly attenuate toward base, 2-4 times as long as the claw; wing-petals equaling the standard, the limb oblong, slightly enlarged upward, rounded-obtuse, somewhat shorter than the claw; keel 15-16 mm long, the limb semicircular, acutish, 2/3 as long as to slightly shorter than the claw; pods sessile, horizontally spreading to subcernuous, narrowly | linear, 3-3.3 cm long, 2 mm broad, straight or slightly curved, gradually | attenuate to a straight subulate beak 5-9 mm long, rounded ventrally, scarcely canaliculate or flattened dorsally, subterete, coriaceous, villous with short ascending white hairs, these interspersed with very short black hairs, bilocular. Fl. May; fr. May-June (July). (Plate XLII, Figure 2). 362 Fine-grained or somewhat gravelly soils on tableland slopes. — Centr. | Asia: Pam.~ Al. Endemic. Described from Vakhsh river valley. Type in | Leningrad. | 623. A. surchobi Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 40-60 cm tall; caudices 15-35 cm long, rather stout, covered with castaneous~brown bark; stems of the year 12-25 cm long, whitish by a dense coat of appressed hairs; stipules adnate at base to petiole, oblong or lance~triangular, covered with appressed mostly white and few black hairs, 3-4 mm long; leaves 7-9 cm long, | 493 663 the petiole and the much longer rachis slender, densely hoary by appressed white hairs; leaflets 6-8 pairs, linear to linear-lanceolate, obtuse or mucronulate, 8-15 mm long, 2-3 mm broad, covered sparsely above and densely beneath with appressed hairs; peduncles 1.5-2 times as long as the leaves, covered with appressed white hairs and about the inflorescence mixed white and black hairs; racemes oblong, loosely and remotely flowered and fruited, the flowers and fruits 7-15 cm long, horizontally spreading; bracts linear-lanceolate, 3-4 mm long, covered with black and white hairs, 2-3 times as long as the pedicel; calyx cylindric, 12-13 mm long, densely covered with spreading white and short black hairs, the linear- subulate teeth 4.5-5 mm long, the tube 1 .5-2 times as long as the teeth; corolla violet; standard 21 mm long, the limb oblong, slightly retuse, angular at base, gradually attenuate into a cuneate claw; wings 20 mm long, oblong, the round-tipped limb as long as the claw; keel 15 mm long, acutish, the semicircular limb about as long as the claw; pods sessile, linear, ca. 3cm long, 2 mm broad, gradually attenuate toward apex, scarcely canaliculate dorsally, subterete, scarcely carinate ventrally, nearly straight, indurated, villous with short ascending white and fewer black hairs, bilocular. Fl. andfr. June, Stony slopes.— Centr. Asia: Pam.- Al. (Karategin). Endemic. Described from vicinity of northern Kashka-terek. Type in Leningrad. 624. A. xiphidium Bge. Astrag. geront. I (1868) 125; II (1869) 218; Grossh. Fl. Kavk. II (1930) 321. Perennial, suffrutescent, 30-50 cm tall, the caudices rather short, woody, branched, up to 5 (7) mm thick, together with branches 7-16 cm long, covered with castaneous-brown bark; stems of the year 8-20 cm long, erect, rarely ascending, finely sulcate, shaggy by a dense coat of appressed white hairs; stipules short-lanceolate, often falcate, 2-3 mm long, herbaceous, covered with appressed white hairs; leaves 3-6 cm long, the rachis 1.5-3 times as long as the petiole, both petiole and rachis slender, covered with appressed white hairs; leaflets 4-5 pairs, narrowly linear, 1-2 cm long, 1-2 mm broad, subobtuse, covered on both sides with appressed white hairs; peduncles 1.5-2 (2.5) times as long as the leaves. sparsely covered with short appressed white hairs; racemes (7) 10-15 cm long, remotely (4) 6-10-flowered; bracts ovate-lanceolate, somewhat exceeding the pedicel, ca. 2mm long, covered with white and black hairs; calyx cylindric, 13-15 mm long, rather densely covered with appressed white and black or sometimes almost pure white hairs, the teeth 2-2.5 mm long, the tube 4-5 times as long as the teeth; corolla whitish, with a faintly pinkish-purple standard; standard 12-17 mm long, the limb oblong-obovate, obtuse or scarcely retuse, 1.5-2 times as long as the claw; wing-petals 20-26 mm long, the limb obovate-oblong, round-tipped, entire, the claw 1.5 times as long as the limb; keel 18-24 mm long, the limb semicircular slightly concave beneath, acutish, barely shorter than the claw; pods sessile, spreading horizontally or obliquely downward, subulate-linear, straight or scarcely curved, 3.5-5 cm long, 2-3 mm broad, acuminate to a straight subulate beak ca. 1 mm long, carinate ventrally, scarcely canaliculate dorsally (the dorsal suture slightly concave), bluntly quadrangular in cross-section, coriaceous, covered with appressed white hairs, bilocular. Fl. (May) June-July; fr. June-August. Dry clay- and stone-and-clay-covered mountain slopes. — Caucasus: E. Transc., Tal. Endemic. Described from the vicinity of Elizavetpol' (Kirovabad) and from Somkhetia. Type in Leningrad. 494 664 Series 2. FEDTSCHENKOANI Vass.-— Shrubs or subshrubs, with (4) 6-9 pairs of leaflets; racemes short, capitate, rarely elongated, loose; corolla violet with admixture of other colors (pleochroism). 20-25 mm long; pods subulate-linear, very elongated, from (50) 70 to 110 mm long and 1.5-2 mm broad.— Mountains in the south of Central Asia. 625. A. Fedtschenkoanus Lipsky in A.H.P. XXVI (1907) 192. A perennial subshrub, up to 60-80 cm tall; caudices up to 35 cm tall, stout, covered with castaneous-brown bark; stems of the year 4-6 cm long, simple or slightly branched, canescent by a dense coat of short appressed white and black hairs; stipules adnate to petiole in lower third, covered with appressed white and black hairs, the free portion broadly oblong, obtusish; leaves (2) 3-5 (7) cm long; leaf petiole and rachis covered with appressed white hairs; leaflets (5) 6-9 pairs, oblong-elliptic, subobtuse, 6-13 (17) mm long, covered scatteredly above and sparsely or fairly densely beneath with appressed white hairs; peduncles equaling to 1.5-2 times as long as the leaves, covered with appressed black and white hairs; racemes capitate, fairly compact, rather many-flowered, ovoid, 3-6 cm long; bracts linear, 4-5 mm long, black hairy; pedicels 1 mm long; calyx cylindric, 11-12 mm long, villous with spreading white and fewer black hairs, the linear teeth ca. 2 mm long; corolla pinkish-violet; standard 20-22 mm long, the limb oblong-ovate, slightly attenuate and angular toward base, broadly short-ligulate upward, slightly retuse, 3-4 times as long as the claw; wing-petals 17-18 mm long, the limb lance-oblong, entire, obtuse or scarcely retuse, Somewhat shorter than to as long as the claw; keel 14-16 mm long, the limb obtuse, semicircular, half as long as the claw; ovary on a stipe ca. 2 mm long; pods gradually attenuate to a stipe ca. 2mm long, erect, narrowly linear, (5)7-11cm long, obcompressed, ca. 3 mm thick and ca. 1 mm broad, with prominent ventral and dorsal sutures, coriaceous, covered with short appressed blackand white hairs, acuminate into a straight slender beak 5-11 mm long, unilocular. Fl. April-May; fr. May-June. (Plate XLIII, Figure 1). Slopes of the intermediate mountain zone, on rocks among shrubs. — Centr. Asia: Syr D., T. Sh. (W.). Endemic. Described from Dzhambulak (southern Kazakhstan). Type in Leningrad. ; 626. A. ischnocarpus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 20-40 cm tall, the caudices strongly abbreviated, 3-5 cm long, rather stout, branched, covered with grayish bark; branches of the preceding year obsolescent, rarely up to 5 cm long, covered with appressed white hairs; stems of the year 1-5 (11) cm long, whitish by a dense coat of appressed hairs; stipules free, triangular to triangular-oblong, ca. 3 mm long, covered with appressed white hairs; leaves 3-7 cm long, the petiole and the much longer rachis rather slender, covered with appressed white hairs; leaflets 5-8 pairs, oval to oblong-oval, (14) 9-10 mm long, round-tipped, covered sparsely above and densely beneath with appressed white hairs, fairly thick; peduncles 2.5-3 times as long as the leaves, erect, covered with appressed white and, beneath the inflorescence, mixed white and black hairs; bracts ovate to oblong- ovate, mostly acuminate, 1.5-2.5 mm long, covered with black and fewer white hairs, somewhat exceeding the pedicel; calyx cylindric, 12-13 mm long, densely villous with spreading short black and long white hairs, 495 665 666 the narrowly linear teeth 2.5-3 mm long, the tube 3-4 times the length of the teeth; corolla dingy pale-violet; standard 21-24 mm long, the limb obovate, scarcely retuse, slightly auriculate-angular at base, 1.5-2 times as long as the claw; wing-petals 20-22.5 mm long, the limb oblong~obovate, rarely obovate, round-tipped, half as long as the claw; keel 17-21 mm long, the limb semicircular, acute, half as long as the claw; ovary ona stipe 1.5-2 mm long, hairy; pods subsessile (the stipe ca. 1 mm long), narrowly linear, (5) 6-8.5 cm long, ca. 1.5 mm broad, straight or rarely somewhat curved, gradually attenuate into a straight subulate beak 3-5 mm long, subterete, not grooved, thinly coriaceous, covered with subappressed short black and more numerous long white hairs, bilocular. Fl. May; fr. June. More or less stony and gravelly mountain slopes, on taluses.— Centr. Asia: Pam.-Al. Endemic. Described from Guzar. Type in Leningrad. 627. A. acanthocarpus Boriss. in Not. Syst. Herb. Inst. Bot. Ac. Sc. URSS VII, fasc. II (1937) 294. A shrub 30-50cm tall, the caudices branched at base, covered withgray bark; stems covered at first with white and black hairs distributed in patches, later with pure white hairs; stipules adnate to petiole, lanceolate, acute, 3-4 mm long, covered with appressed black and fewer white hairs; leaves 2-4 cm long; leaflets (4) 5-6 pairs, oblong to lance-oblong, 5-11 mm long, 1-3 mm broad, the upper ones often narrowly linear, 8 mm long, 1 mm broad, subacute to subobtuse, hoary on both sides by appressed hairs, finally glabrate; peduncles 1.5 times as long as the leaves; racemes densely capitate, 8-12-flowered, the flowers subsessile; bracts lanceolate, ca. 3mm long, covered with spreading black hairs; calyx cylindric, 10-12 (13) mm long, villous with spreading white and fewer appressed black hairs, the teeth subulate, unequal, two of them 4 mm and the three others ca. 2mm long; corolla dingy violet; standard 20-25 mm long, the limb oblong-lanceolate, enlarged in upper third, entire at apex; wing~-petals 15-20 mm long, the limb oblong, somewhat attenuate upward, entire, the claw 1.5 times the length of the limb; keel 16-18 mm long, the limb half as long as the claw; pods sessile, subulate-linear, 8-10 cm long, ca, 2 mm broad, sparsely covered with appressed black and white hairs, quadrangular, acuminate toward apex, subbilocular. Fl. May-June; fr. June-July. ; Desert wormwood associations on high-mountain slopes, at altitudes of 2700-3280 m.— Centr. Asia: Pam.-Al. (Shugnan). Endemic. Described from the vicinity of Dilekh settlement in Vakhan-Ishkashimskii district. Type in Leningrad. Series 3. SPRYGINIANI Vass. — Shrubs with stems up to 30-50 cm long; leaflets (3) 4-7 pairs, elongated, fairly large; corolla yellow, from 21 to 30 mm long; pods erect, subulate-linear, 20-30 mm long and 2-3 mm broad. — Mountains in the south of Central Asia (W. TienShan, Kopet Dagh). 628. A. Spryginii M. Pop. inN.A. Dimo. Pochyv. eksp. v bass. rr. Syr-dar'i i Amu-dar'i I (1915) 51. A shrub up to 45 cm tall; caudex stout, up to 7 mm thick, covered with brownish-gray bark, up to 14 cm tall; branches of the preceding year 6-20 cm long, covered with appressed white hairs; stems of the year 5-8 cm long, densely covered with appressed hairs, all white except for 496 ‘667 isolated black hairs beneath the nodes; stipules oblong-triangular or sublinear, 2-4mm long, acuminate, herbaceous, covered withappressed black and white hairs; leaves 7-8cm long, the petioles andthe much longer rachis slender, covered with appressed white hairs; leaflets (3) 4-5 pairs, linear to oblong- elliptic, 10-25mmlong, 2-5 mmbroad, subobtuse to subacute, sometimes minutely mucronulate, covered scatteredly above and more densely beneath with appressed white hairs; peduncles axillary, 6-12 cmlong, rather sparsely covered with appressed white hairs; racemes loose, 5-8 cm long, remotely flowered and fruited, the rachis covered with appressed white and black hairs; bracts oblong-lanceolate, 2-3 mm long, black-hairy, covered with isolated white hairs, exceeding the pedicel; calyx cylindric, 17-19 mm long, covered with appressed white and black hairs, the tube 8-9 times as long as the subulate teeth; corolla yellow; standard 28-30 mm long, the limb oblong-obovate, slightly constricted about the middle, angularly auriculate below, cuneately attenuate at base into the claw; wing-petals ca. 25 mm long, the limb oblong, entire, 8 mm long; keel ca. 23 mm long, the limb semioval, obtuse, ca. 8 mm long; pods borne on a stipe 4.5-6 mm long, cernuous, linear, 2-2.8 cm long, subapplanate-obcompressed, 2.5-3 mm thick, slightly concave dorsally, prominently sutured on both sides, coriaceous, glabrous, bilocular. Fl. andfr. April-May. Stony slopes of low mountains. — Centr. Asia: Syr Darya. Endemic. Described from Mogoltau. Type in Leningrad. 629. A. nigriceps M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). A shrub woody to high up, 28-35 cm tall; caudices 4-7 cm long; branches 7-15 cm long, erect, rather slender, the older covered with castaneous-brown bark, those of the preceding year cinereous by a dense appressed white tomentum; branches of the year (2.5) 4-6 cm long, slender, terete, densely covered with appressed white hairs, nearly white except for black hairs at the nodes; stipules nearly free, adnate to petiole in lower part, herbaceous, triangular-oblong, acutish, 2-3 mm long, covered with white and black hairs; leaves (5) 6-7 cm long, the rachis 2-3 times as long as the slender petiole, both petiole and rachis covered with appressed white hairs; leaflets (4) 6-7 pairs, narrowly linear to lance- linear, acute, (6) 10-15 mm long, 1-1.5 (2) mm broad, densely covered on both sides with appressed white hairs, silvery-black; peduncles 2-3 (4) times the length of the leaves, (8) 10-14 cm long, rather slender but firm, densely covered with appressed white and, beneath the inflorescence, mixed white and black hairs; racemes short and broad, capitate, 3-4 cm long, (3) 5-12-flowered, not elongating in fruit; bracts ovate, 2-3 mm long, densely covered with black hairs; calyx cylindric, 12-15 mm long, densely tomentose-villous with white and black hairs, either predominating, the tube 4-5 times as long as the linear-subulate teeth; corolla stramineous, with a faintly lilac standard; standard 21-23 mm long, the limb obovate- orbicular, entire, slightly auriculate-angular at base, as long as the claw, 9-10 mm broad; wing-petals 20-22 mm long, the limb obovate-oblong, entire, rounded-obtuse, the claw 1.5-2 times as long as the limb; keel 20 mm long, the limb semicircular, acute, about half as long as the claw; ovary on a stipe 1.5 mm long; pods sessile (immature), erect, subulate- linear, acute, ca. 2.3 cm long and 2 mm broad, villous with ascending white and black or pure white hairs. Fl. June. 497 668 Mountain slopes in the steppe and juniper zone.— Centr. Asia: Mountainous Turkmenistan. Endemic. Described from Kulyar Mountain near Gaudan. Type in Leningrad. Series 4. ARBUSCULI Vass.— Shrubs and subshrubs; leaflets elongate, 2-3 (4-5) pairs; racemes capitate, short, rarely loose and remotely flowered; corolla violet or purple, 15-20 mm long; pods straight or slightly curved, (15) 20-30 (33) mm long and 1.5-2 mm broad. — Plains of Central Asia and desert-dominate high-mountain valleys of Central Tien Shan. 630. A. arbuscula Pall. Astrag. (1800) 19; DC. Prodr. II (1825) 286; Ldb. Fl. Ross. I (1842) 632; Bge. Astrag. geront. I (1868) 124; II (1869) 2175 Kryl. FR oZap. Sib.) Vil 2833) 1695!" A hori zont a bits! har eet Kir. in Bull. Soc. Nat. Mosc. XIV (1841) 406.— A. eremothamnus Kar. et Kir. l.c. XV (1842) 334 pp. — Exs.: H. Fl. URSS No. 3136. A branched shrub 50-120 cm tall; caudices erect, stout, up to 1.5 cm in diam, branching in upper part, (10) 30-80 cm tall, covered with castaneous~brown, longitudinally splitting bark; branches of the year simple, 5-20-(30) cm long, firm, densely covered with short appressed white hairs; stipules free, adnate, to petiole in lower part, triangular- ovate to ovate-lanceolate, (2) 3-4 mm long, often acuminate, densely covered with appressed white hairs; leaves (2.5) 3-5 cm long, sessile, the rachis 1-2 cm long, covered with appressed white hairs; leaflets 2-4 pairs, linear, (8) 15-40 mm long, (1) 1.5-3 mm broad, rarely linear- lanceolate or oblong-oval, up to 7 mm broad, acute, appressed~hairy on both sides; peduncles 2-3 times as long as the leaves, densely covered with appressed short white hairs; racemes compactly capitate, 2-3 (4) cm long, 8-20-flowered, obovoid, rarely oblong-ovaloid; bracts ovate or lanceolate, 1-2 (3) mm long, more or less covered with white and black hairs; calyx cylindric, 7-10 mm long, densely tomentose-villous with short ascending white and fewer black hairs, dense hairs sometimes confined to the teeth, the filiform subulate teeth 1.5-2 mm long; corolla purple; standard 15-19 mm long, the limb rhomboid, attenuate upward, scarcely retuse, abruptly contracted in lower fourth, angular at base, 4-5 times as long as the claw; wing-petals 14-17 mm long, the limb linear- oblong, rounded-ok.use, as long as the claw; keel 11-14 mm long, the limb inequilaterally oblong, -obtusish, somewhat shorter than the claw; ovary on a stipe 0.3-0.4 mm long; pods borne in a rather crowded capitate raceme, sessile, spreading, rarely cernuous, linear-subulate, straight, (1.7) 2-3 cm long, (1.5) 2 mm broad and thick, abruptly narrowed to a straight subulate beak 1-1.5 mm long, carinate ventrally, flat or rarely somewhat convex dorsally, triangular or rarely quadrangular in cross- section, coriaceous, covered with appressed or slightly spreading white and black or rarely pure white hairs, bilocular. Fl. June-August; fr. June- August. Steppes, semideserts, and wormwood deserts; much less frequent on sands. Often on stony slopes.— W. Siberia: U. Tob., Irt., Alt.; Centr. Asia: Ar.-Casp., Balkh., Dzu.-Tarb., T. Sh. Gen. distr. Kulja. Described from Chazil'-tash Mountains (E. Balkh). Type in London. 631. A. temirensis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). 498 669 Adensely leafy shrub, 20-40 cm tall; caudices rather stout, up to 6 mm thick, well developed, (5) 12-25 cm long, erect or ascending; bark covering the caudices and branches splitting longitudinally and peeling, grayish- brown, white-tomentulose with appressed hairs, smooth castaneous-brown when exposed; branches of the year 1.5-4 (10) cm long, slender, whitish by a dense coat of appressed white hairs; stipules free, herbaceous, adnate to petiole in lower part, triangular-ovate, 2-3 mm long, covered with appressed white or mixed white and black hairs; leaves 2-3 (4) cm long, the obsolescent petiole and the rachis slender, covered with appressed white hairs; leaflets 4-5 pairs, lanceolate or linear; 5-12 mm long, 1-3 mm broad, acute, rarely obtuse, covered on both sides (faintly above) with appressed hairs; peduncles 2-3 times the length of the leaves, covered with short appressed white and, in upper part, mixed white and black hairs; racemes umbelliform-capitate, 1.5-2 cm long, (6) 8-15- flowered; bracts ovate to oblong-ovate, 1-1.5 mm long, covered with short black hairs or, at least, black-ciliate on the margin; calyx short- cylindric, 7-8 mm long, densely covered with ascending white and black hairs, often striped black, the triangular-subulate teeth ca. 1 mm long; corolla violet (?); standard 15-18 mm long, the limb obovate, not attenuate upward, 7-8 mm long, round-tipped and scarcely retuse, slightly constricted above the lower 1/4-1/3, 4.5-5 mm broad below, angular at base; wing- petals 13-16 mm long, the limb oblong, entire, somewhat shorter than the claw; keel 12-13 mm long, the limb broadly semioval, acutish, the claw 1.5 times as long as the limb; ovary ona stipe ca. 0.5 mm long; pods crowded in an umbelliform~-capitate raceme, more or less obliquely ascending, sessile, subulate-linear, (1.5) 2-2.5 cm long, 1.5-2 mm broad, attenuate into a straight subulate beak ca. 1 mm long, straight of slightly curved, carinate ventrally, subcomplanate or scarcely canaliculate dorsally, trihedral in cross-section, coriaceous, covered with white and black or rarely pure black hairs, bilocular, Fl. May; fr. May-June. Steppes and grass-and-wormwood semideserts.— Centr. Asia: Ar.- Casp. Endemic. Described from the vicinity of Temir. Type in Leningrad. 632. A. tscharynensis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). A shrub 30-35 cm tall; caudices 5-10 cm long, branched, covered with castaneous-brown bark; stems of the year 5-10 cm long, densely covered with appressed white hairs; stipules adnate to petiole at base or up to 1/3, ovate-triangular, ca. 1.5 mm long, covered with white appressed hairs; leaves 2.5-3.5 cm long, the very short petiole and the rachis densely covered with appressed white hairs; leaflets 2-3 pairs, approximate, oblong-obovate or oblong, rounded-obtuse, rarely minutely mucronulate, rather densely covered on both sides with appressed white hairs; peduncles 13-19 cm long, greatly exceeding the leaves, covered with short appressed white hairs; inflorescence compactly capitate, subspherical, 10-18- flowered; bracts ovate-lanceolate, 1.5-2 mm long, hyaline, black-ciliolate; calyx cylindric, 8-9 mm long, densely covered with subappressed white hairs interspersed with very few black ones, the teeth linear~-subulate, 1-1.2 mm long, the tube 6-7 times as long as the teeth; corolla violet; standard 18-19 mm long, the limb rhomboid-obovate, obtusish, abruptly contracted in lower part, angular at base, 4-5 times as long as the claw; wing-petals 16-17 mm long, the limb linear-oblong, entire, as long as 499 the claw; wing-petals 16-17 mm long, the limb linear-oblong, entire, as long as the claw; keel 13-15 mm long, the limb semiobovate-oblong, the claw 1.5 times as long as the limb; ovary ona stipe 0.4 mm long; pods borne on a stipe ca. 0.7 mm long, spreading to subcernuous, subulate- linear, slightly arched upward, 25-30 mm long, ca. 2 mm broad, compressed laterally, carinate ventrally, acute and terminating ina straight subulate beak 1-1.5 mm long, coriaceous, densely covered with appressed black and profuse ascending white hairs, bilocular. Fl. May-June; fr. June-July. Mountain slopes, less frequently on sands.— Centr. Asia: Tien Shan. Endemic. Described from Charyn River (Alma-At. region). Type in Leningrad. 670 639. A. duanensis Saposhn. ex Sumn. in Animadv. Syst. Herb. Univers. Tomsk. 1-2 (1933) 6. Suffrutescent, 20-40 cm tall; caudex strong, 2-3 cm long, branched, the branches ascending, 3-6 (8) cm long, rather stout, covered with grayish-brown longitudinally splitting bark; branches of the year ascending, rarely erect, (3) 4-11 (13) cm long, rather numerous, white by a dense coat of short appressed hairs; stipules connate, in lower part adnate to petiole, lanceolate or oblong-ovate, 2-3 mm long, covered with appressed white or mixed white and black hairs; leaves 2-4 cm long, the petiole and the longer rachis covered with short appressed white hairs; leaflets (2) 3-4 pairs, lance-oblong, subobtuse, (5) 7-15 (20) mm long, 2-3 mm broad, densely covered on both sides with appressed hairs, silvery-gray; peduncles 3-5 times the length of the leaves, 10-23 cm long, firm, covered with appressed white and, in upper part, mixed white and black hairs; racemes compactly capitate, 2-2.5 cm long, 8-16-flowered; bracts oblong-ovate to ovate-lanceolate, 1-1.5 mm long, tomentose with dense black and very few white hairs; calyx cylindric, 7-8 (9) mm long, densely villosulous with spreading black and white hairs, cinerescent, the subulate-linear teeth 2mm long; corolla violet (?); standard 17-20 mm long, the limb rhomboid, gradually attenuate upward, slightly retuse, rather abruptly contracted in lower 1/4, angular at base, 4-5 times as long as the claw; wing-petals 14-17 mm long, the limb oblong, mostly retuse, as long as the claw; keel 11-13 mm long, the limb semicircular, obtusish, the claw slightly longer than to 1.5 times as long as the limb; ovary on a stipe ca. 0.5 mm long; pods sessile, obliquely ascending or horizontally spreading, rarely subcernuous, capitately approximate, subulate-linear, straight or barely curved, 1.8-3.3 cm long, 2 mm broad, carinate ventrally, flattened and slightly drawn out dorsally, triangular in cross-section, coriaceous, densely covered with short appressed black and ascending white hairs, cinerescent, bilocular. Fl. May-June; fr. June. Slopes of mountain valleys, at altitudes of 2100-2500 m.— Centr. Asia: Tien Shan (centr.). Endemic. Described from Alabugariver valley. Type in Leningrad. 634. A. polyceras Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 332; Bge. Astrag. geront.I (1868) 125; II, 219; Bge. Astrag. turk. 285. A shrub 30-35 cm tall, the weakly branched caudex 10-20 cm long and up to 6 mm thick, covered with grayish-brown more or less longitudinally splitting bark; branches of the year 10-25 (30) cm long, firm, terete, whitish by a dense coat of short appressed white hairs; stipules adnate 500 671 to petiole at base, free, triangular-lanceolate or triangularly oblong-ovate, acuminate, 4-5 mm long, densely covered with appressed white hairs; leaves 3-5 cm long, the rachis 2-3 times the length of the petiole, both petiole and rachis covered with appressed white hairs, indurescent; leaflets 2-3 pairs, linear to linear-oblong, 12-20 (30) mm long, (1) 2-4 mm broad, covered sparsely above and densely beneath with appressed hairs, hoary; peduncles 1.5-2 times as long as the leaves, covered with appressed white hairs; racemes 6-14 cm long, loose, remotely (4) 9-15-flowered; bracts linear-lanceolate, 3-4 mm long, covered with short white hairs; calyx cylindric, 10-12 mm long, 2.5mm broad, densely covered with short black mostly ascending hairs, subpannose, the teeth filiform-subulate, 2-2.5 (3) mm long, the tube 4 times as long as the teeth; corolla purple (?); standard 21 mm long, the limb rhomboid-oval, short-attenuate toward apex, scarcely retuse, abruptly contracted in lower 1/4, angular at base, 4 times as long as the claw; wing-petals 19-20 mm long, the limb linear, entire, obtuse, as long as the claw; keel 15 mm long, the limb semioval, acute somewhat shorter than the claw; ovary on a stipe ca. 0.5 mm long; pods sessile, subcernuous, subulate-linear, straight or slightly arcuate, 2.3-3.1 cm long, 1.5-2 mm broad, abruptly narrowed into a straight subulate beak 1-2 mm long, carinate ventrally, flattened or slightly convex dorsally, 3- or 4~angled, coriaceous, villous-tomentose with short appressed black and ascending white or merely ascending white hairs, bilocular. Fl. May; fr. June. Saline clayey soils. — Centr. Asia: Balkh. (between Lepsy River and the source of the Baskan, Sasyk-pastau in vicinity of Lake Alakul', northern shore of Lake Balkhash). Endemic. Described from Sasyk-pastau. Type in Moscow; cotype in Leningrad. 635. A. Gontsharovii Vass. sp. nova in Addenda XI, p. 666. Shrub up to 1 mm tall; stems branched from the middle, woody in lower part, the bark light brown, peeling; stems of the year vested with appressed white hairs; leaves 5-8 cm long; leaflets (4) 5-6 pairs, oblong, 7-8 to 20-25 mm long and from 2 to 7 mm broad, covered beneath with scattered appressed hairs, glabrous or covered with isolated hairs above, green on both sides; racemes short, capitate; peduncles 1.5 times as long as the leaves; bracts linear-subulate, 3-4 mm long, studded with white and black hairs; calyx 7-8 mm long, blackish by the hairs, the tube 5-6 times the length of the teeth; corolla pale violet to practically white; standard 15-18 mm long, oval, slightly retuse; keel 10-11 mm long, exceeded by the wings; pods linear, crowded, obliquely ascending, covered with appressed black and scattered white hairs, 20-30 mm long and 2-2.5 mm broad, abruptly narrowed to a readily breaking beak 3-5 mm long, i72 borne on a very short stipe (0.5 mm). Fl. May-June; fr. June-July. t Mountain meadows.— Centr. Asia: T. Sh., Dzu.-Tarb. Endemic. Described from Chatyrbai mountain pass near Lepsy. Type in Leningrad. Note. As disclosed by observations of the type specimen preserved in the Botanical Institute of Leningrad. A. eremothamnus Kar. et Kir. is synonymous with A. arbuscula Pall. Bunge erroneously referred to this species of plants collected by Kushakevich in Trans-Ili Ala Tau 30 June 1873. These plants should be included in the new species named after N.F. Goncharov. 501 673 Series 5. FALCIGERI Vass.— Subshrubs; leaflets 1-2 (3-5) pairs, narrow, elongated; racemes short, capitate; corolla pink or purple; pods cernuous, falcate, narrowly linear, 25-70 mm long and 1.5-2 (3) mm broad. From Alatau and Zeravshan westward to Karakalpak. 636. A. falcigerus M. Pop. in Bull. Univ. As. Med. fasc. 14 (1926) 143. A dwarf, loosely cespitose subshrub, 8-13 cm tall; caudex short, branched; branches rather slender, flexuous, ascending or subdecumbent, thinly canescent, 1-5 cm long; stems of the year 1-4 (6) cm long, erect, slender, whitish by a dense coat of appressed white hairs; stipules almost free or connate up to the middle, adnate in lower part to petiole, triangular- ovate or lanceolate, 1 mm long, sparsely covered with appressed white hairs; leaves 1-1.5 cm long, the short petiole and the rachis slender, canescent by appressed or less often spreading hairs; leaflets 3-5 pairs, filiform to narrowly linear, acute, 5-15 (20) mm long, 0.6-2 mm broad, covered on both sides with appressed or sometimes ascending hairs; peduncles 3-4 times as long as the leaves, 5-10 cm long, erect, firm, rarely abbreviated, hoary with white hairs; inflorescence compactly capitate, (2) 5-10-flowered, 2-3 cm long, the rachis andthe pedicels covered with appressed mainly black hairs; bracts ovate, 0.5-1 mm long, hyaline, covered with short black hairs or rarely glabrate; calyx cylindric, (6) 10-11 mm long, covered with appressed hairs, these black on the ventral and mainly appressed white on the dorsal side, the setaceous-linear teeth (1) 1.5-2 mm long, the tube 4-5 times as long as the teeth; standard 18-20 mm long, the limb rhomboidally oblong-obovate, broadly short- attenuate toward apex, retuse, constricted above the lower 1/3, slightly subauriculate-dilated below, abruptly and angularly contracted into and 4-5 times as long as the claw; wing-petals 16-18 mm long, equaling the standard, the limb linear-oblong, sometimes slightly enlarged toward apex, entire, as long as the claw; keel 14-15 mm long, the limb semioval, acutish, the claw 1.5 times as long as the limb; pods sessile, linear, arcuate or more often falcate, (2) 5-7 cm long, 1 mm broad, acuminate into a subulate beak 1 mm long, cylindric, rounded or slightly carinate ventrally, somewhat prominently sutured dorsally, rather sparsely covered with appressed white hairs, bilocular. Fl. May; fr. June. Stony mountain slopes.— Centr. Asia: Pam.-Al. (Zeravshan: Mogian). T.Sh. (Karatau Mts., Talass Alatau, Chu-Ili Mountains). Endemic. Described from Karatau, from upper reaches of Arystanda River near Kurok Dam and Almasai pass. Type in Tashkent. 637. A. aulieatensis M. Pop. in Bull. Univ. As. Med. fasc. 14 (1926) 141. Suffrutescent or frutescent; caudex short, buried, woody; branches short or long (up to 18 cm), erect or assurgent, covered with brown bark; stems of the year 10-30 cm long, stout, subflexuous, erect or assurgent, densely covered with short white hairs; stipules barely adnate to petiole, triangular-ovate, ca. 3 mm long, acute, membranous, covered with appressed white and black appressed hairs; leaves short, the petiole 0.5-2.5 mm long, indurescent and persistent but not spinescent, densely covered with appressed white hairs; leaflets 1-3 pairs, oblong to oblanceolate, 15-25 mm long, 4-10 mm broad, obtuse or subobtuse, covered on both sides with appressed white hairs; fruiting peduncles 10-14 cm long (the racemes 12-20 cm long), firm, ca. 1.5 mm thick, terete, sparsely 502 covered with appressed white hairs; racemes short, capitate, 5-15- flowered; bracts lanceolate, 1-2 mm long, membranous, covered with black hairs; calyx cylindric, ca. 10 (11) mm long, villous with spreading white hairs interspersed with some black hairs; the tube 3-4 times as long as the teeth; pods ca. 0.5 mm long, stipitate, linear, subarcuate, rarely almost straight, 25-43 mm long, 2-3 mm broad, 3-4 mm thick, cernuous, carinate ventrally, thickened or rounded dorsally, coriaceous, densely covered with ascending white and black hairs, bilocular. Fr. August (September). Gypsiferous conglomerates, in the foothills. — Centr. Asia: Tien Shan (W., at bases of northern slopes of Talass Ala Tau mountains). Endemic. Described from the vicinity of Romanovka village. Type in Tashkent. 638. A. scleroxylon Bge. in Arb. Nat. Ver. Riga 1 (1847) 241; Bge. Astr. geront. II (1869) 230; Bge. Astr. turk. (1880) 290. Suffrutescent, 50-100 cm tall, the caudex rather long (up to 30 cm?), woody, strongly (twofold or threefold) branched, the caudex and the woody branches covered with longitudinally peeling grayish-brown bark; shoots of the year (3) 5-13 cm long, terete, white by a dense coat of appressed hairs, slightly flexuous; stipules adnate to petiole in lower 1/3, 1.5-2mm 674 long, triangular, acutish, densely covered with white hairs; leaves 1-2 cm long, the petiole together with rachis 0.5-1.0 cm long, slender, densely covered with appressed white hairs, hoary; leaflets 1-2 pairs, oblong or oblanceolate, rarely oblong-obovate, cuneate at base, round-tipped, (5) 7-15 mm long, 2-3 mm broad, rather densely covered on both sides with appressed white hairs, hoary; peduncles equaling to twice as long as the leaves, 2-3 cm long, hoary, densely covered with appressed white and, beneath the inflorescence, mixed white and black hairs, terete; racemes rather compact, ca. 4 cm long, (3) 7-11 (16)-flowered, the rachis 0.7-1.2 (2) cm long, the flowers subumbellately approximate; bracts lance-oblong, acutish, 3-4 mm long, covered with short mostly white and fewer black hairs; calyx cylindric, 12-16 mm long, lanate with short spreading white hairs, these interspersed on the teeth with some black hairs, the linear teeth 2.5-3 mm long; corolla pink; standard 25-27 mm long, the limb oblong-obovate, round-tipped and scarcely retuse, gradually attenuate into a claw 5-7mmlong; wing-petals 24-25 mm long, the limb obovate to oblong-obovate, half as long as the claw; keel 22-23 mm long, the limb semioval, acutish, the claw 2.5 times as long as the limb; ovary on a stipe ca. 1 mm long, white-hairy; pods sessile, horizontally spreading, finally often subcernuous, 25-35 mm long, 2.5-3 mm broad, acute and terminating in a straight beak ca. 1 mm long, carinate ventrally, flattened or rounded dorsally, coriaceous, densely villous with ascending soft white hairs, arcuate, bilocular. Fl. May; fr. May-June. (Plate XLIII, Figure 3). Gravelly and stony or less frequently sandy slopes of mountains and low clayey or sandy buttes in the desert region. — Centr. Asia: Kyz. K., Syr. D., Balkh. Endemic. Described from Kyzyl Kum (Bakali). Type in Leningrad. Series 6. LORINSERIANI Vass. — Herbaceous perennials or subshrubs; leaflets linear or rarely oval-oblong; racemes loose, elongated; corolla yellow, fairly large; pods subulate-linear, from (20) 25 to 50-60 mm long and 1-2 (3) mm broad. Steppe foothills and mountain blocks in the southern part of Central Asia and Transcaucasia. 503 ZZ Y PIILE \ PLATE XLII Lipsky. 504 \ 639. A. Lorinserianus Freyn in Bull. Herb. Boiss. 2 Sér. 5 (1905) 565. | A low perennial subshrub, 20-35 cm tall: caudices very short, 677 sometimes almost obsolescent, or up to 12 cm long, covered withcastaneous bark; stems of the year 5-10 cm long, densely vested with appressed white | hairs, terete, grayish-white; stipules free, herbaceous, triangular, the upper lanceolate, acute, 3-4 mm long, covered with appressed white hairs; | leaves 4-5 (6) cm long, the rachis 3-4 times as long as the petiole, both petiole and rachis covered with appressed white hairs; leaflets 4-5 pairs, very variable in shape, from oval or elliptic to oblong and linear, obtuse or acute, (5) 8-15 mm long, covered scatteredly above and rather densely beneath with appressed white hairs, hoary; peduncles 1.5-2 times as long as the leaves, firm, erect, covered with white and, beneath the inflorescence, with mixed white and few black hairs, hoary; racemes remotely flowered. (4) 6-12 cm long, elongating in fruit up to 15 cm, the flowers obliquely ascending or horizontally spreading; bracts lanceolate to linear-lanceolate, 2-4 mm long, covered with white or mixed white and black hairs; calyx cylindric, 17-18 mm long, densely covered with appressed black and white hairs, the filiform-linear teeth 3-4 mm long, the tube 3-4 times as long as the teeth; corolla bright yellow; standard 24-27 mm long, the limb obovate, to oblong-obovate, scarcely retuse, slightly attenuate and dingy [ ?] at base, 2-3 times as long as the claw; wing-petals 23-27 mm long, the limb linear-oblong, entire, round-tipped, about half as long as the claw; keel 20-21 mm long, the limb almost semicircular, obtusish, the claw 2-2.5 times as long as the limb; ovary on a stipe 4-5 mm long, hairy; pods borne on a stipe 4-5 mm long, cernuous, linear, straight | (2) 2.5-3.5 cm long, subcomplanate-obcompressed, ca. 3 mm thick and ca. 1 mm broad, with prominent ventral and dorsal sutures, coriaceous, _ covered with short appressed white and black hairs. unilocular. F1. (May). June-July; fr. June-July. | Clayey bluffs, rocks and taluses, rarely in rather fine-grained soil overlying rocks in mountains and elevated foothills, at altitudes up to 2500 m.— Centr. Asia: T. Sh., Pam.- Al. Endemic. Described from _ Sufi-kurgana and Alai mountain range. Type in Geneva. Note. Fairly often the leaflets of lower leaves are elliptic, while those of upper leaves are linear. , | 640. A. maverranagri M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS _ & (1946). Frutescent, 13-30 (40) cm tall; stems 4-15 (30) cm long, ascending, slender but firm, angled, sparsely hispid with appressed hairs, subflexuous; stipules free, the lower ovate-triangular, the upper triangular-lanceolate, acuminate, 3-4 mm long, herbaceous, covered with appressed black hairs; leaves 5-8 cm long, the petiole long and slender; leaflets 5-9 pairs, narrowly linear, the upper subfiliform, acute, cinerescent sparsely above and more densely beneath by appressed hairs, 8-15 mm long, 0.5-2 mm 18 broad; peduncles exceeding the leaves; racemes remotely few-flowered; | bracts lanceolate, 1 mm long; calyx cylindric, 12-15 mm long, covered _ with appressed black and white hairs, the teeth subulate-linear; corolla yellow; standard 23-27 mm long, the limb subrhomboid-obovate, slightly _ fetuse, angularly attenuate in lower 1/4, twice as long as the claw; wing- | petals 21-22 mm long, the limb oblong, round-tipped, half as long as the __ claw; keel 19-20 mm long, the limb obtuse, half as long as the keel; pods _ Subsessile, on a stipe 1 mm long, subulate-linear, 2.5-5 cm long, 505 subfalcate, 1.5-2 mm broad, not grooved, with slightly convex sutures (the dorsal suture occasionally somewhat concave), coriaceous, covered with appressed white and black or rarely pure white hairs, bilocular, the subulate beak ca. 2 mm long. Fl. April (May); fr. May. Low foothills in the meadowgrass and sedge vegetation belt including tall xerophytic grasses, Centr. Pamir-Alai (Surkhandar'ia river basin), penetrating into the lower part of the ephemeral vegetation belt, rarely (in the north) in the upper part of the meadowgrass and sedge belt. — Centr. Asia: Syr D., Pam.-Al. (interior Pam.-Al. from Baisun district to Chul'bair foothills). Endemic. Described from Dzhilga. Type in Tashkent. 641. A. lancifolius Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 30-46 cm tall; stems 13-30 cm tall, erect or assurgent, slightly grooved, covered with appressed white hairs; stipules 3-7 mm long, white-hairy, the lower triangular, the upper lance-triangular; leaves 5-10 (14) cm long, the petiole and the much longer rachis covered with short appressed white hairs; leaflets (3) 4-7 pairs, oblong-elliptic or lanceolate, the upper ones rarely linear-lanceolate, 8-20 mm long, acute, green above with scattered appressed hairs, hoary beneath with sparse | appressed hairs; peduncles equaling the leaves, covered with appressed white and, beneath the inflorescence, mixed white and black hairs; racemes few-flowered (5) 6-8 cm long, (4) 6-9-flowered; bracts lanceolate, 2-3 mm > long, covered with black and white hairs, 1.5-2 times as long as the pedicel; calyx cylindric, 16-18 mm long, covered with appressed black and white | or rarely pure white hairs, the tube 4-5 times as long as the linear- | subulate teeth; corolla yellow; standard 25-27 mm long, the limb oblong- obovate, broadly ovate in upper 1/2-2/3, scarcely retuse, attenuate below and somewhat auriculately angular-dilated toward base, twice as long as _the claw; wing-petals 23-25 mm long, the limb oblong, round-tipped, the | claw 1.5-2 times as long as the limb; keel 20-22 mm long, the limb | obtusish, the claw 2.5 times as long as the limb; ovary ona stipe 2.5-3 mm | long; pods gradually attenuate into a stipe 1-2 cm long, narrowly linear, 679 3-3.5 cm long, 1.5-2 mm broad, almost straight or slightly curved, rarely falcate, slightly carinate ventrally, rounded dorsally with a convex or rarely concavesuture, subterete, coriaceous, covered with appressed mixed white and black or rarely pure white hairs, bilocular. Fl. April- June; fr. May-June. Dry needlegrass or rarely. witchgrass steppes or wormwood associations, on slopes and summit plateaus of mountain ranges, at the altitude of about 2000 m, descending at the northern limit into the foothills of the Vaksh river valley (vicinity of Kyzyl-Kalu). Often in more or less gravelly soil. — Centr. Asia: Pam.-Al. Endemic. Described from the vicinity of Khadzha- | beles in the Aktau Mountains. Type in Leningrad. 642. A. exilis A. Kor. in Not. Syst. ex herb. Inst. Bot. Ac. Sc. URSS Vil, fasc. 8 (1938) 171.—A. rubrovenosus M. Pop. in herb. i Perennial, 25-60 cm tall; caudex strong short buried, often raised up to 3-4 cm above ground level, the aerial divisions woody, 1-4 cm long, giving rise to annual stems; stems of the year (5) 12-26 (28) cm long, erect, firm, virgate, often branching in lower part, terete, densely covered with i short appressed silvery hairs; stipules adnate to petiole at base, triangulary) acute, 1.5-2 mm long, covered with appressed white hairs; leaves 506 680 (6) 9-18 cm long, the petiole about as long as the rachis, both firm, subvirgate, densely covered with short appressed silvery hairs; leaflets 2-3 pairs, linear, rarely linear-lanceolate, (8) 10-30 (50) mm long, subacute, canescent loosely above and densely beneath with appressed white hairs; peduncles 1.5-2 times the length of the leaves, 10-19 cm long, firm, virgate, terete, densely covered with short appressed white hairs, hoary, in upper part with admixture of few black hairs; racemes loose, 10-24 cm long, remotely (7) 10-20-flowered, the yellow flowers obliquely ascending; bracts ovate-lanceolate, acute, (1) 1.5-2.5 mm long, covered with white hairs, these sometimes interspersed with few black hairs; pedicels 1.5-2 mm long, densely covered with white hairs; calyx cylindric, 15-19 mm long, covered with appressed black and white hairs, the tube 3-5 times as long as the linear~subulate teeth; standard 18-23 mm long, the limb oblong-obovate, attenuate in lower 1/3-1/2, slightly angular at base, 2-2.5 times as long as the claw; wing-petals about equaling the standard, the limb oblong, entire, round-tipped, half as long as the claw; kell 18-20 mm long, the limb obtusish, the claw 2-2.5 times as long as the limb; pods borne on a stipe 1.5-2 mm long, narrowly subulate-linear, 5-6 cm long, 1.5 (2) mm broad, more or less falcate, not grooved, coriaceous, densely covered with appressed white hairs, rarely interspersed with very few black hairs; seeds oblong, ca. 3 mm long, bilocular. Fl. May-July; fr. June-August. Mottled outcrops, mostly red sandstone, in foothills and low mountains. — Centr. Asia: Pam.-Al. Endemic. Described from Mount Tash-Tipe of Hissar Range. Type in Leningrad. 643. A. subexcedens Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, 55-65 cm tall; stems erect, firm, 30-35 cm long, angled- sulcate, sparsely covered with appressed white hairs, scabrous; stipules adnate at base to petiole, almost free, 8-10 mm long, herbaceous, the lower lanceolate, subulately acuminate, the upper subulate-linear; leaves 6-11 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis sparsely covered with short appressed white hairs; leaflets 5-6 pairs, linear to lance-linear, 2-3.5 cm long, sparsely covered on both sides with appressed hairs, green; peduncles 2-3 times the length of the leaves, 15-19 cm long, firm, erect, angled-sulcate, sparsely covered with short appressed white hairs; racemes elongated, loose, remotely many-flowered, as long as the peduncle; bracts narrowly linear, 5-7 mm long, sparsely covered withwhite and black hairs; calyx narrowly. cylindric, 14-15 mm long, 2 mm broad, covered with white and black hairs, often black-striped, the tube 3-4 times the length of the subulate teeth; corolla yellow; standard 22-24 mm long, the limb orbicular in upper part, scarcely emarginate at apex, oblong-attenuate in lower 1/3, slightly angular at base, 3 times as long as the claw; wings 21-23 mm long, the limb oblong, entire, round-tipped, half as long as the claw; keel 18-19 mm long, the limb convex on both sides, obtuse, the claw 3-4 times as long as the limb; pods subsessile, erect or obliquely ascending, linear, straight, 17-18 mm long, 1.5mm broad, barely exserted from the calyx, carinate ventrally, rounded dorsally, sometimes with a scarcely concave dorsal suture, terminating in a slender subhamate to beak ca. 2 mm long, coriaceous, covered with appressed white hairs, bilocular. Fl. May; fr. May-June. 507 681 Clayey mountain slopes.— Centr. Asia: Pam.-Al. (Karatau Mts. and southern Tadzhikistan). Endemic. Described from vicinity of Sardoba- Kutal' Pass. Type in Leningrad. 644. A. transhyrcanus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, rather low, suffrutescent; caudex distinct but short, the rather slender divisions covered with brownish-gray bark; stems of the year 5-10 cm long, erect, slender, densely covered with short appressed white hairs; stipules slightly adnate to petiole, the upper ones free, triangular-lanceolate, small, covered with white or sometimes mixed white and black hairs; leaves 4-10 cm long, the rachis 3-4 times as long as the petiole; leaflets 3-4 (5) pairs, narrowly linear, up to 3 cm long, ca. 1mm broad, acute, conspicuously nerved; peduncles 12-15 cm long (together with raceme 15-25 cm), firm, covered with appressed grayish hairs; racemes remotely 3-7-flowered; bracts oblong-linear, 1-2 mm long, covered with appressed white and black hairs; calyx long~-cylindric, 15-18 mm long, covered with appressed white and black hairs, the subulate 2-4 mm long; corolla yellow; standard 25-28 mm long, the limb not attenuate at apex, retuse; wing-petals equaling the standard, the limb entire; keel by 2-3 mm shorter than the standard, the claw 3-4 times the length of the limb; ovary stipitate; pods short~-stipitate, pendulous, subulate-linear, up to 6 cm long, 2-3 mm broad, almost straight or somewhat curved, grooved dorsally, finely pointed at apex, appressed- hairy, bilocular. Fl. and fr. May. Mountain gorges.— Centr. Asia: Mtn. Turkm. (W. Kopet Dagh: Karakalinskii district). Endemic. Described from Bakhcha Canyon, near northern Yartykal village at Chandyr' River. Type in Ashkhabad. 645. A. xiphidioides Fr. et Sint. in Bull. Herb. Boiss. sér. 2, V (1905) 786. Perennial, 20-40 cm tall; stems 10-30 cm long, canescent by appressed white hairs; stipules triangular-oblong, or lanceolate (the lower ones sometimes triangular-ovate), 4-5 mm long, acuminate, covered with white and black hairs, adnate to petiole in lower part; leaves 3-8 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis covered with appressed white hairs; leaflets (4) 5-7 pairs, linear to linear-lanceolate, the lower mostly lanceolate or oval, acute or obtuse, 5-30 mm long, 1.5-5 mm broad, appressed-hairy.on both sides; peduncles equaling to 1.5 times as long as the leaves, densely covered with appressed white hairs; racemes remotely 5-12-flowered, (5) 7-13 cm long bracts lanceolate, 1.5-2 mm long, covered with mixed white and black or pure black hairs, 1.5 times as long as the pedicel; calyx cylindric, 13-15 mm long, densely covered with appressed white and black hairs, the subulate teeth ca. 2 mm long; corolla yellow; standard 21-24 mm long, the limb obovate, scarcely retuse, abruptly narrowed toward base, angular at base, 2.5-3 times as long as the claw; wing-petals mm long, the limb rather broadly oblong, entire, the claw 1.5-2 times as long as the limb; keel 18-21 mm long, the limb semiorbicular, obtusish, half as long as the claw; pods sessile, subulate-linear, horizontally spreading or cernuous, 3-5 cm long, 27>2.5mm broad, carinate ventrally, rounded or canaliculate dorsally, acute, terminating in a straight subulate beak ca. 2 mm long, straight or falcate, bilocular. Fl. April; fr. May. 508 682 683 Wormwood and ephemeroid deserts. — Centr. Asia: Mtn. Turkm. (Kopet Dagh, Badkhyz). Gen, distr.: Afghanistan. Described from Turkmenia. Type in Leningrad. 646. A. xanthoxiphidium Fr. et Sint. in Bull. Herb. Boiss. sér. 2, V CSO >) pi Gchiane Ee xsesi Sint wlts No.) 1 70Mmvar Perennial, suffrutescent, ca. 35 cm tall; caudices branched, 7-10 cm long, covered with a castaneous-brown bark, rather stout; stipules triangular-lanceolate, green, faintly covered with white hairs, 3-4 mm long; leaves 5-7 cm long, the petiole as long as the rachis, both slender, covered with few white appressed hairs; leaflets 4-6 pairs, linear, 7-30mm long, ca. 22 mm broad, greenish, sparsely covered on both sides with appressed hairs; peduncles twice as long as the leaves, more or less covered with appressed white hairs; racemes remotely 15-20-flowered, 10-12 cm long; bracts ovate-lanceolate, 1.5-2 mm long, covered with white and black hairs; calyx cylindric, 11-13 mm long, densely covered with appressed white and black hairs, the subulate teeth 2-2.5 mm long; corolla yellow; standard ca. 21 mm long, the limb obovate, slightly retuse, slightly attenuate in lower 1/3, angular at base, 2.5 times as long as the claw; wing-petals ca. 20 mm long, the limb oblong, entire, half as long as the claw; keel ca. 18 mm long, the limb semiorbicular, obtusish, half as long as the claw; pods sessile, subulate-linear, slightly curved, 2.5-3.5 cm long, ca. 2 mm broad, densely covered with appressed white and black hairs, terminating in a point ca. 2 mm long, bilocular. Fl. May; fr. May- June. Mountain slopes.— Centr. Asia: Mtn. Turkm. (W. Kopet Dagh), Kara K. (Krasnovodsk). Endemic. Described from Sundsodag Mountain (W. Kopet Dagh). Type in Leningrad. Note. Freyn and Sintenis (Freyn et Sint. in Bull. Herb. Boiss. sér. 2, V (1905) p. 788-799, 1012-1015) described a number of subspecies of A. xanthoxifidium, suchas A. accrescens from the vicinity of Ashkhabad; A. obscurus, A. latifoliolatus, A. holoxanthus, A. curvicaulis, from Karakal; A. campylopus, A. rectus from the vicinity of Krasnovodsk; A. Broterusii; also the species A. variifolius (l.c. 784) from the Kazandzhik district, A. ufraesis (l.c. 568) from the Krasnovodsk district; A. Juratzkanus (l.c. 567) from the Kyzyl-arvat vicinity, are sometimes presented as independent species (O. et B. Fedtsch. Consp. Fl. Turk. I, 234), but are apparently synonyms of the polymorphic A. xanthoxipidium. It is possible that some of them may prove to be distinct species, but for the time being this problem cannot be settled due to insufficiency of material. 647. A. argyroides G. Beck. ex Stapf in Denkschr. Acad. Wien. XL. (1886) 341.— A. angustatus Bge. in Boiss. Fl. or. II (1872) 483, non Boiss. in Diagn. 1849; Bge. Astrag. geront. II, 220, ex parte. Perennial, practically herbaceous, scarcely woody at base, 8-20 cm tall, lignified at base, with short divisions; stems of the year, 5-10 (13) cm long, stoutish, geniculate, covered with short appressed white hairs, grayish-white or gray, stoutish, terete; stipules triangular-ovate or linear, ca. 2mm long, appressed-hairy; leaves4-8 cm long, the petiole 1-2 cm long, covered with appressed white hairs; leaflets 4-5 pairs, elliptic, rarely obovate, 10-15 mm long, 3-7 mm broad, acute, covered on both sides with appressed white hairs, gray beneath, greenish above; peduncles 509 684 3-4 cm long, covered with appressed white hairs, the leaves 1.5-2 times as long as the peduncles; racemes loose, remotely 8-12-flowered, 3-5 cm long; bracts triangular-lanceolate, 1-2 mm long, hyaline, covered with white or mixed white and black hairs; calyx ca. 14 mm long, covered with appressed black and white or rarely almost pure white hairs, the linear- lanceolate teeth 1-1.5 mm long; corolla yellowish with a faintly violet standard; standard 22-24 mm long, the limb oblong-obovate, retuse, attenuate in lower 1/3, angular at base, 4 times as long as the claw; wing- petals 21-23 mm long, the limb oblong, entire at apex, the claw 1.5 times as long as the limb; keel 20-22 mm long, the limb semioval, half as long as the claw; acutish; pods subsessile, subulate-linear, spreading to cernuous, 30-40 mm long, ca. 2 mm broad, acuminate at both ends, compressed laterally, sharply carinate ventrally, rounded or rounded- carinate dorsally, coriaceous, covered with appressed white hairs, bilocular. Fl. April-May; fr. May-June. ; Clayey, often gypsiferous mountain slopes.— Caucasus: S. Transc. (vicinity of Nakhichevan'). Gen. distr.: N. Iran. Described from Manzhil (Iran). 648. A. Sachokianus Sosn. in Proceed. Ac. Sc. Azerb. S.S.R., No.I (1945) 34. Perennial, vested with appressed silvery gray hairs; stems many, woody at base, 10-20 cm long; leaflets 4-5 pairs, elliptic to rounded- elliptic, 6-12 mm long, 3-6 mm broad, acute, covered on both sides with silvery-gray hairs; racemes rather loosely 5-10-flowered; bracts 1-2 mm long, equaling the pedicel, hairy on the margin; calyx 6-7 mm long, covered with black and white hairs, the tube 4-6 times as long as the teeth; corolla 15-18 mm long, yellowish; pods 20-22 mm long, hairy, slightly curved, beaked, 3 times as long as the calyx, densely covered at first with appressed white hairs, finally glabrate. Fl. andfr. June. Caucasus: E. Transc. (Shemakha district). Endemic. Described from Shemakha district, between Azhidzha and Kushcha-gorai. Type in Baku. Series 7. SUMBARENSES Vass.-— Differing from the preceding series in the short subumbellate to capitate racemes. Pamir-Alai, Kopet Dagh. 649. A. sumbari M. Pop. in Not. Syst. Inst. Bot. Ac.Sc. URSS X (1946). ; Perennial, dwarf, suffrutescent, (10) 15-25 cm tall; caudices short, branched in lower part, the branches 2-6 (15) cm long, covered with pale castaneous-brown (longitudinally peeling bark; stems of the year (1) 3-7cm long, white by appressed hairs, often flexuous; stipules free, barely adnate at base to petiole, 2-2.5 mm long, herbaceous, densely covered with appressed white hairs, the lower triangular, the upper triangular- lanceolate; leaves (2) 4-6 cm long, the petiole 2/3 as long as to equaling the rachis, densely covered with appressed white hairs, hoary; leaflets 5-8 pairs, linear to lance-linear, 4-9 mm long, covered sparsely above and densely beneath with appressed hairs, hoary; peduncles 2-3 times as long as the leaves, firm, covered with white and, in upper part, with mixed white and black hairs; racemes subumbellately abbreviated, 2.5-3cm long, closely many-flowered; bracts lanceolate, 2.5-3 mm long, covered with white and black hairs, 2.5-3 times the length of the pedicel; calyx cylindric, 13-15 mm long, densely covered with short subappressed black 510 185 and white hairs; corolla yellow; standard 22-24 mm long, the limb oblong- obovate, scarcely retuse, attenuate in lower 1/3, angular at base 2-3 times as long as the claw; wing-petals 17-20 mm long, the limb oblong to linear- oblong, rounded-obtuse, the claw 1.5-2 times as long as the limb; keel 16-19 mm long, the limb semioval, acutish, barely shorter than the claw; pods approximate, obliquely ascending to suberect, sessile, narrowly linear, (3) 4-6.5 cm long, 2-2.5 mm broad, slightly compressed laterally, not grooved and carinate, subcylindric, gradually attenuate into a beak 2-2.5 mm long, coriaceous, rather densely covered with appressed white and black hairs, bilocular. Fl. May; fr. June. : Mountain slopes.— Centr. Asia: Mtn. Turkm. Endemic. Described from Kopet Dagh. Type in Leningrad. 650. A. Neilreichianus Fr. et Sint. in Bull. Herb. Boiss. sér. 2 V (1905) 571.— Exs.: Sint. Iter transc.- pers. 1900-1901, No. 1623. Perennial, 15-27 cm tall; shoots of the year spreading, simple, angularly flexuous, 8-20 cm long, densely covered with short appressed hairs, whitish, terete; stipules free, herbaceous, ovate or oblong- triangular, 2-3 mm long; leaves 5-8 cm long, the petiole shorter than the rachis, covered with appressed white hairs, cinerescent; leaflets 6-7 pairs, oval to oblong-oval, rarely elliptic-lanceolate or [?] curved, (4) 7-17 mm long, covered scatteredly above and rather densely beneath with appressed white hairs, round-tipped or acutish; peduncles equaling to somewhat exceeding the leaves, covered with appressed white hairs; racemes abbreviated, often subumbellate, loosely 5-7-flowered, 3-5 cm long; bracts ovate, short-acuminate, covered with white and black hairs, 1.5-2.0 mm long; pedicels 1.5-2 mm long; calyx cylindric, 14 mm long, densely covered with appressed white hairs, these interspersed on the nerves in the upper part of the tube and on the teeth with few black hairs, the subulate-linear teeth 1.5-2 mm long; corolla yellow; standard 20-24 mm long, the limb oblong-obovate; retuse, attenuate in lower 1/3, slightly angular at base, 3-4 times as long as the claw; wing-petals 19-22 mm long, the limb oblong, round-tipped, the claw 2-2.5times as long as the limb; keel 17-18 mm long, the limb semicircular, scarcely convex above, obtuse, the claw 3 times the length of the limb; pod subsessile (the stipe ca. 1 mm long), narrowly linear, 4-5 cm long, 2.5 mm broad, acute at both ends, carinate ventrally, rounded dorsally, more or less compressed laterally, coriaceous, densely covered with appressed hairs, whitish, bilocular, straight or slightly curved. Fl. April; fr. April, May. Clay-and-sand-surfaced desert takyrs, low mountains, and sands. — Centr. Asia: Mtn. Turkm. Endemic. Described from Kyzyl-Arvat. Type in Leningrad. 651. A. stenoceroides Boriss. in Not. Syst. ex Herb. Inst. Bot. Acad. Se URSo Ve tase. Il (1937) 232. Perennial, 10-25 (30) cm tall; the caudex very short, partly buried and partly aerial; stems (5) 7-15 (20) cm long, erect or in lower part ascending, densely covered with appressed white hairs, whitish; stipules free, the lower ovate, the upper oblong-ovate or lanceolate, 2-2.5 mm long, acuminate, covered with appressed white and black hairs; leaves 3-4 mm long, the short petiole and the rachis densely covered with appressed white and black hairs; leaflets (4) 5-7 pairs, lance-oblong to oblong-elliptic, 5-10 mm long, (1) 2-3 (4) cm broad, obtuse, or oblong-obovate, rarely 511 linear, cuneate at base, covered on both sides, densely beneath, with stiff appressed hairs, silvery-gray; peduncles 3-4 times as long as the leaves, densely covered with appressed white hairs; racemes umbellately capitate, 3-3.5 cm long, 8-10 (12)-flowered; bracts lanceolate, ca. 2 mm long, covered with appressed white and black hairs; calyx cylindric, 13-15 mm long, covered with short appressed black and fewer white hairs, the subulate teeth 3-4 mm long; corolla yellow; standard 22-25 mm long, the limb oblong7obovate, ca. 10 mm broad, broadly short-attenuate upward, abruptly attenuate in lower 1/3 to about half the breadth, 4-5 times as long as the claw; wing-petals 20-22 mm long, the limb linear-oblong, scarcely retuse, the claw about 1.5 times as long as the limb; keel 17-19 mm long, the limb obovate, acutish, half as long as the claw; ovary on a stipe ca. 1 mm long; pods borne on a stipe ca. 0.5 mm long, subulate-linear, (2) 3-5 cm long, 1.5 mm broad, straight or barely curved, obliquely ascending, terete, not carinate and not grooved, acuminate into a slender straight subulate beak 1-1.5 mm long, coriaceous, covered with appressed white and black hairs, bilocular. Fl. June-July; fr. (July) August. Gravelly and fine-grain-surfaced mountain slopes in the saltwort-and- wormwood semidesert belt, rarely in meadowgrass-wormwood-and- needlegrass steppes, ascending up to 2800-3000 m.—Centr. Asia: Pam.- Al. (upper course of the Pyandzh River basin: Shugnan, Vakhan-Ishkashim). Endemic. Described from the vicinity of Khorog. Type in Leningrad. 686 652. A. dsharkenticus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent at base, the caudex divisions slender, grayish- yellow, diffusely many~branched, subdecumbent, 3-7 cm long; stems of the year many, slender, ascending, terete, greenish, sparsely covered with appressed hairs; stipules shortly connate at base, covered with appressed white and black hairs 1-2 mm long, the free portion triangular - subulate; leaves 4-6 (7) cm long, the petiole half as long as or rarely longer than the rachis, green or scarcely canescent; leaflets 3-4 pairs, mostly oblong-lanceolate, acute, 10-20 mm long, 3-4 mm broad, appressed-hairy, greenish or scarcely canescent; peduncles 10-15 cm long, slender, greenish; racemes capitate, 5-10-flowered, the black~hairy rachis 5-10 mm long; bracts triangular-lanceolate, 1-1.5 mm long, membranous, sparsely covered with white and black hairs; calyx 12-13 mm long, covered with black (mostly-on the ventral side) and white hairs, the subfiliform teeth 2 mm long; corolla lurid; standard 20-22 mm long, broadly retuse; wing-petals 18-19 mm long, broadly retuse; keel 17 mm long, the limb 5 mm long; pods sessile, spreading, linear-subulate, arcuate, 25-30 mm long, 2-3 mm broad, carinate ventrally, grooved dorsally, densely covered with ascending black and longer white hairs, hoary. Fl. May-June. Steppe mountain slopes.— Centr. Asia: Dzu.- Tarb. (Dzhungarian Ala Tau). Endemic. Described from the upper reaches of Bol'shoi Usek River. Type in Alma Ata. Series 8. MACROTROPIDI Vass. — Herbaceous perennials; stems short, ascending or decumbent, rarely erect; leaflets many pairs, small; racemes short, capitate-umbelliform, rarely loose and elongated; corolla 20-27 mm long, varying in color; pods erect, linear-subulate, 20-35 (40) mm long and (1.5) 2-3 mm broad. 512 Steppes (in the north) and steppe-dominated mountain slopes (in the south). 653. A. macrotropis Bge. Astrag. geront. I (1868) 127; II (1869) 223; Astrag. turk. (1880) 286. Perennial, herbaceous, (12) 15-22 cm tall; caudex very short, woody, buried; stems 5-12 (18) cm long, slender, covered with short appressed white and black hairs; stipules free, triangular, 2.5-3.5 mm long, acute, covered with appressed white and black hairs; leaves (3) 4-7 (10) cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis slender, covered with short appressed white and black hairs; leaflets 7-9 pairs, 687 lanceolate to linear-lanceolate, 5-8 (11) mm long, acute to subobtuse, covered scatteredly above and densely beneath with short appressed white hairs; peduncles equaling to 1.5 (rarely 2) times as long as the leaves, covered with short appressed white and black hairs; racemes abbreviated, subumbellate, the flowers obliquely ascending, the fruits subhorizontally spreading; bracts lanceolate, 2-2.5 mm long, covered with short appressed white and black hairs; calyx cylindric, 11-15 mm long, covered with short appressed black and white hairs, the linear teeth ca. 2-2.5 mm long; corolla pink; standard 24-27 mm long, the limb oblong-obovate, auriculate- angular at base, broadly short-ligulate at apex, retuse, 2-2.5 times as long as the claw; wing-petals 17-21 mm long, the limb linear-oblong, retuse, the claw 1.5 times as long as the limb; keel 16-19 mm long, the limb semioval, obtuse, 2-2.5 times as long as the claw; pods subapproximate, subhorizontally spreading or scarcely cernuous; sessile, linear, (2.5) 3-4 cm long, 2-2.5 mm broad, acute, terminating in a slender straight beak 2-2.5 mm long, subterete, rounded ventrally, grooved dorsally, coriaceous, covered with short appressed white and black hairs, bilocular. Fl. May-June; fr. June-July. More or less stony and gravelly slopes of elevated foothills and mountains.— Centr. Asia: Pam.-Al., T. Sh., Dzu.- Tarb. Endemic. Described from Arganata. Type in Leningrad. 654. A. amabilis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 10-15 cm tall; stems (4) 7-10 cm long, ascending, slightly curved, densely covered with appressed white hairs, hoary; stipules adnate to petiole at base, ovate to oblong-ovate, 2.5-3 mm long, covered with appressed white and black hairs; leaves (2) 3-4 cm long, the rachis equaling te 2-3 times as long as the petiole, both petiole and rachis slender, covered with appressed white hairs, hoary; leaflets 5-6 pairs, elliptic to lance- elliptic, short-acuminate, (5) 7-9 mm long, (2) 3-4.5 mm broad, covered sparsely above and densely beneath with appressed hairs, cinerescent; peduncles equaling to 1.5 times as long as the leaves, densely covered with appressed white hairs; racemes abbreviated, subumbellate, ca. 4 cm long, 7-10-flowered; bracts lanceolate to ovate-lanceolate, 2-4 mm long, villous with ascending black and very few white hairs; calyx long-cylindric, 14-16 mm long, densely villous with ascending white and few black hairs, the subulate teeth half as long as the tube; corolla purplish-violet; standard 23 mm long, the limb oblong-obovate, abruptly narrowed in lower 1/3, slightly angular at base, short-attenuate toward apex, retuse, 4 times as long as the claw; wing-petals 21 mm long, the limb oblong, round-tipped, the claw 2.5 times as long as the limb; keel 19 mm long, the limb semioval, 688 obtusish, the claw about 3 times as long as the limb; ovary sessile; pods 513 689 sessile, more or less spreading, narrowly linear, 2.5-4cm long, 2.5-3 mm broad, scarcely curved, attenuate into a subulate beak 1.5-2 mm long, scarcely carinate ventrally, convex dorsally, slightly compressed laterally, rather densely covered with short appressed or ascending white hairs, biloculan: Sls Mavyawune : wer June: Onblack pebbles.—Centr. Asia: Tien Shan (submontane plain of Trans-Ili Ala Tau). Endemic. Described from the vicinity of Chilik settlement. Type in Leningrad. 655. A. macroceras C.A.M. in Bong. et Mey. Verzeichn. (1841) 24. — A. macrolobus M.B. Fl. taur.-cauc. III (1819) 493 pp. (quoad pl. ex Siberia); Bge. Astrag. geront. II (1869) 222 pp.; Kryl. Fl. Zap. Sib. VII (193°3) AkO0O" npr alice Pall: SAstracn tapre205 5. Perennial, 10-20 (50) cm tall, the buried caudices much divided; stems of the year 1.5-8 (20) cm long, mostly ascending at base, densely covered with appressed white hairs, silvery-white; stipules almost free or connate at base, 3-5 mm long, covered with appressed white or mixed white and fewer black hairs, broadly triangular, rarely the upper ones triangular- lanceolate, acute; leaves 3-7 cm long, the petiole half as long as the rachis; leaflets 5-7 pairs, lance-oblong to lance-elliptic, subobtuse, 5-12 mm long, 1.5-4 mm broad; peduncles equaling or rarely up to 1.5 times as long as the leaves, covered with appressed white hairs; racemes abbreviated, capitate, broadly obovoid, 4-5 cm long, rather loosely 7-15-flowered; bracts ovoid-lanceolate, ca. 2 mm long, equaling to 1.5 times as long as the pedicel, covered with black or mixed white and black hairs; calyx cylindric, 9-12 mm long, covered with appressed black and fewer white hairs, the tube 4-5 times the length of the lance~subulate teeth; corolla white or violet; standard 20-22 mm long, the limb oblong-obovate, short-attenuate toward apex, retuse, rather abruptly narrowed in lower 1/3, angular at base, 5-6 times as long as the claw; wing-petals 15-17 mm long, the limb linear-oblong, retuse, as long as the claw; keel 13-14 mm long, the limb broadly obovate, acutish, the claw 1.5 times as long as the limb; ovary borne on a stipe 1-1.5 cm long; pods sessile, obliquely ascending, linear, somewhat curved, 20-30 mm long, 2-2.5 mm broad, attenuate into a subulate beak ca. 1 mm long, slightly carinate ventrally, canaliculate dorsally, subtriquetrously terete, villous with appressed white hairs and also sparsely covered with short opr Ese eee black hairs, bilocular. Fl. May- August; fr. June-August. Often stony or gravelly steppe mountain slopes and steppe meadows. — W. Siberia: Altai. Endemic. Described from S. Altai. Type in Leningrad. Note. Forms occurring along Chulym River (Altai) in Balagansk district are transitional between A. Ionae Palib. and the Altaian A. macroceras C.A. M., with broader oblong leaflets, broader and often longer pods (up to 25 mm), and more developed stems up to 10 mm long. 656. A. ortholobiformis Sumn. in Animadyv. Herb. Univ. Tomsk. Nos. 9— 10 (1936), 8.— A. ortholobus auct. fl. turk., non Bge. Perennial, greenish, from a short buried caudex; stems ascending, 9-21 cm long, 1-1.5 mm thick, covered with appressed white hairs; stipules free, lanceolate, 2.5-3 mm long, 1/2-2/3 mm broad, covered with appressed white and black hairs; leaves 5-9 cm long, the petiole 1-2 cm long; leaflets 4-7 pairs, linear-oblong or lanceolate, short-acuminate, 5-11 mm long, 514 690 1-2 (3) mm broad, appressed-hairy; peduncles 1.5-3 times as long as the leaves; inflorescence umbelliform-capitate, 4-8-flowered; bracts lanceolate, about equaling or exceeding the pedicel, 1.5-3 mm long, ca. 0.5 mm broad; calyx cylindric, 13-15 mm long, covered with appressed black and white hairs, the subulate teeth 1.5-2 mm long, the tube 7-10times as long as the teeth; corolla whitish or pink; standard 26-28 mm long, the limb oblong-obovate, 9-10 mm broad, attenuate toward apex, retuse, slightly constricted about the middle; wing-petals 25-27 mm long, the limb retuse, the claw 3-4 times as long as the limb; ovary subsessile, 28-30- ovuled; pods spreading, sessile, subulate-linear, 25-33 mm long, 1.5-2mm broad, straight or slightly falcate, 2.5-3 times as long as the keel, bilocular. Fl. May; fr. May-June. Southern stony mountain slopes.— Centr. Asia: Tien Shan (Kurdai Mountains and elsewhere). Endemic. Described from the Tokmak district. Type in Tomsk. Note. According to the author: ''Habit identical with that of A. ortholobus Bge., from which it differs in more pointed (not obtusish) [calyx] teeth, the tube 7-10 times (as against twice) as long as the teeth, the oblong-obovate standard attenuate at apex and constricted near the middle (not oblong-obovate, not attenuate toward apex, waistless), the claw of wings 3-4 times as long as the limb". 657. A. angarensis Turcz. ex Bge. Astrag. geront. 1 (1868) 26; II (1869) 222. Perennial, 5-12 cm tall, the slender buried prostrate caudices up to 10 (20) cm long, the divisions ligneous, not more than 1 cm long, mostly slender, prostrate, covered with yellowish-brown bark; stems of the year 2-8 cm long, slender, erect or assurgent, densely covered with appressed white hairs sometimes interspersed with few black hairs; stipules triangular-ovate, 2-4 mm long, densely covered with black and fewer white hairs; leaves 2-5 cm long, the rachis equaling to 2-3 times as long as the petiole, the petiole rather sparsely covered with appressed white and sparse black hairs; leaflets 6-8 (10) pairs, linear to narrowly linear, 5-15 mm long, 1-2 (3) mm broad, scatteredly hairy or glabrate above, rather densely covered beneath with appressed hairs, cinerescent; peduncles somewhat exceeding to 1.5 times as long as the leaves, covered with appressed white and, beneath the inflorescence, black hairs; bracts ovate, 1-2 mm long, hyaline, densely black-hairy; inflorescence capitately umbellate, 3-6- flowered; racemes 2-2.5 cm long; calyx cylindric, 6-8 (10) mm long, densely covered with short black and rather long ascending white hairs, the teeth linear, 1.5-2 (3) mm long,the tube 3-4 times as long as the teeth; corolla violet; standard 16-18 (22) mm long, the limb obovate, gently short- attenuate at apex, slightly retuse, 3-5 times as long as the claw; wing- petals 14-16 mm long, the limb linear-oblong, entire, as long as the claw; keel 12-14 mm long, the limb obtusish, as long as the claw; pods sessile, subulate-linear, obliquely ascending, straight or slightly curved, 20-30 mm long, 2-2.5 mm broad, carinate ventrally, canaliculate dorsally, attenuate into a beak 1.5-2 mm long, rather sparsely covered with appressed white hairs, coriaceous, almost fully bilocular. Fl. May-June; fr. June-July. Dry clayey or stony steppe slopes; rarely riverside sands. —E. Siberia: Ang.- Say., Le.- Kol. (in Lena river basin as far as Yakutsk). Endemic. Described from Angara River in vicinity of Balagansk. Type in Leningrad. 515 691 Note. The species A. Miklaschevskii N. Basil., described by N. Bazilevskii (Not. Syst. ex Herb. Hort. Bot. Petrop. V, 5 (1924) 70), differs from A. angarensis Turez. inthe shape of leaflets (elliptic), size of flower, and shape of standard. In the absence of sufficient information, we are not in a position to accord to this plant the standing of a distinct species. 658. A. ortholobus Bge. Astrag. geront. (1868) 127; Kryl. Fl. Alt. I (1908) 323; Fl. Zap. Sib. VII (1933) 1702. Perennial; stems short, divided, prostrate, woody; annual stems 15-20 cm long, erect or more or less spreading, white by a compact cover of appressed hairs; stipules free, oblong-ovate, white-hairy, covered at base with white and black hairs; leaves 2-4 cm long; leaflets 4-8 pairs, linear-oblong, hairy, 8-18 mm long and 2-4 mm broad; peduncles up to twice as long as the leaves, covered with appressed white and partly black hairs; racemes abbreviated, subcapitate, ca. 2.5 cm long; bracts exceeding the pedicel; calyx cylindric, ca. 1.2 mm long, covered with black and sparser white hairs, the tube 4 times the length of the linear teeth; corolla whitish; standard elongate-obovate, ca. 28 mm long and 11 mm broad, subbilobate; wing-petals ca. 20 mm long, slightly retuse; keel 17 mm long; pods linear, straight, deeply canaliculate dorsally, subtriquetrous, 20-30 mm long and ca. 2-3 mm broad, profusely beset with appressed white hairs, these interspersed with few black hairs, Fl. July; fr. August. Valleys of mountain streams.— W. Siberia: Altai. Endemic. Described from Altai. Type in Paris. Note. Reports concerning the occurrence of this species near Chkalovsk (Orenburg) and in Tien Shan are unauthenticated. 659. A. karkarensis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, (7) 14-25 (35) cm tall; stems more or less developed, scarcely ligneous at base, slender, branched, (1.5) 6-15 (20) cm long, ascending to erect, angled-sulcate, covered with appressed white and black hairs; stipules lanceolate or triangular-ovate, acute, 2-3 mm long, thinly membranous, adnate at base to petiole, tomentose on the outside; leaves (2) 4-6 (9) cm long, the rachis 2-3 times as long as the petiole, the petioles weak, slender, rather sparsely covered with appressed black and white hairs; leaflets (5) 7-10 pairs, oblong, or oval-oblong or linear, (3) 5-12 (16) mm long, obtuse, greenish glabrate above with isolated hairs at the margin, cinerescent beneath with scattered appressed white hairs; peduncles firm, 1.5-2 times as long as or rarely equaling the leaves, sulcate, covered with appressed white and black hairs, (1.5) 6-11 (14) cm long; inflorescence compactly short-ovoid, 3-5 cm long, the (5) 8-14 approximate flowers erect or spreading; bracts lanceolate, acuminate, ovate-lanceolate, 25-3 mm long, covered with black and white hairs, exceeding the short pedicel; calyx cylindric, 9-10 mm long, covered with appressed black hairs, the teeth 1-2 mm long; corolla whitish-violet; standard 22 mm long; wing- petals 20 mm long, retuse; keel 16.5 mm long; ovary 22-ovuled; pods linear, slightly compressed laterally, ca. 35 mm long, narrowly carinate ventrally, erect to slightly spreading, straight or slightly curved, cuspidate, covered with appressed black and white hairs. Fl. May-July; fr. July- August. 837, 1117 516 692 693 Steppe associations and foothills, on alluvial deposits. — Centr. Asia: Tien Shan (Kungei-Ala Tau Range: Karkara River, Naryn river basin and Atbashi River). Endemic. Described from Karkara. Type in Leningrad. 660. A. tephrolobus Bge. Astrag. geront. I (1868) 187; II 222; Kryl. Bi. Alt. (908) 323); Fl. Zap. Sib: VIL (1933) 1701. Perennial, dwarf (up to 5 cm tall) grayish-white by appressed hairs, the cauline shoots stout, short, branched; aerial shoots very short, not exceeding 2.5 cm; stipules ovate, covered with appressed white hairs; leaves subsessile, up to 2.5 cm long; leaflets 5-6 pairs, oval or elongate- obovate, up to 5 mm long and 1.5 mm broad, commonly conduplicate, grayish-white on both sides by appressed hairs; flowering peduncles equaling the leaves, finally somewhat elongating; racemes abbreviated, umbelliform, 4-7-flowered; bracts equaling or exceeding the pedicel; calyx short-cylindric, ca. 10 mm long, covered with appressed white and black hairs the tube 4-5 times as long as the linear teeth; corolla whitish; standard elongate-obovate, broadly retuse, partly bilobate, 22-25 mm long and 9 mm broad; wing-petals 18-22 mm long, bilobate; keel 15-18 mm long; pods straight, linear, upturned, subterete, 25 mm long and 2 mm broad, cinerescent by appressed white and black hairs. Fl. July. River valleys and mountain steppes.— W. Siberia: Altai. Endemic. Described from Altai. Type in Paris. 661. A. compressus Ldb. Fl. alt. III (1831) 304; Fl. Ross. I (1842) Goze ery. il. Zap. sib. Vil (1933) 1697: — Ie: Ldb: leon. ‘pl Fl. Ross: III, (1831) tab. 289. Perennial; stems many, 6-30 cm long, decumbent, ligneous at base, covered with brown peeling bark, white above by a profuse cover of appressed hairs; leaves 1-2.5 cm long, the petiole 2-10 cm long; leaflets 3 or rarely more pairs, linear, 4-5 (20) mm long and 1-1.5 mm broad, covered on both sides but more densely beneath with appressed white hairs; racemes 2.5-3 cm long, 5-12-flowered; peduncles 2-4 times as long as the leaves; bracts somewhat exceeding the pedicel; calyx cylindric, 8-10mm long, covered with appressed black hairs interspersed with longer spreading white hairs, the linear-lanceolate teeth 1-1.5 mm long; corolla lilac-purple; standard obovate, retuse, 20-22 mm long and 8-9 mm broad; wing-petals somewhat exceeded by the standard, slightly retuse, oblong, the claw 1.5 times as long as the limb; keel ca. 17 mm long; pods linear, 20-28 mm long and 2-3 mm broad, gradually attenuate at apex, beset with tightly appressed white and black hairs, bilocular. Fl. May; fr. from June. Sands and gravelly sites in river and lake valleys.— W. Siberia: Irt., Alt. Endemic. Described from Semipalatinsk (estuary of Kurchum River). Type in Leningrad. 662. A. chaetolobus Bge. Supplem. Fl. Alt. (1836) 93; Astrag. geront., 126; II, 221; Kryl. Fl. Alt. I (1908) 320; Fl. Zap. Sib. VII (1933) 1697. — A. macrolobus Lab. Fl. alt. (1831) 307 et Fl. Ross. I (1842) 613, pp. Perennial, dwarf, grayish by appressed hairs; cauline shoots short, branched, decumbent, ligneous; aerial shoots short; leaflets 4-7 pairs, elongated; racemes abbreviated, subumbellate, few-flowered; peduncles Somewhat exceeding the leaves; calyx cylindric, the teeth subulate; corolla whitish; wing-petals exceeded by the standard, subentire at apex; pods erect, straight, linear, 27 mm long and ca. 3.5 mm broad, white-lanate by spreading hairs. Fl. May. O17 6 94 Plains.— W. Siberia: Irt., Alt. Endemic. Described from the Semipalatinsk district. Type in Paris. 663. A. ceratoides M.B. Fl. taur.-cauc. III (1819) 429; DC. Prodr. II, 284: )tLdbs FisRoss. ly G30; ‘Kryls hile (Zapesib). Vil, Loor A. stenolobus Bge. in Mém. Sav. Etr. Petersb. II (1835) 593. Perennial, from a woody crooked root; cauline shoots short, woody; stems ofthe year 5-30 cm long, erect or assurgent, rather sparsely covered with appressed bicuspidate hairs; stipules free, subovate, obtusely pointed, appressed-hairy; leaves short-petioled, green, sparsely covered especially beneath with appressed bicuspidate hairs, 3-6 cm long; leaflets 6-9 pairs, oblong obtuse or sublanceolate acuminate, 5-15 mm long and 1-4 mm broad; peduncles twice as long as the leayes; flowers 6-10, lilac- purple, gathered in a subumbellate or capitate raceme; bracts ovate, appressed-hairy, shorter than to about equaling the pedicel; calyx cylindric, 6-7 mm long or rarely longer, covered with appressed black and few white hairs, the tube 6 or more times as long as the short obtusish teeth; standard 18-22 mm long and 7-10 mm broad, oblong-obovate, retuse; wing-petals exceeded by the standard, the limb incised at apex, as long as the claw; keel shorter than the wings; ovary appressed-hairy, borne on a very short stipe, 20-30-ovuled; pods at first ascending, horizontally spreading to cernuous in maturity, linear, long-acuminate, 4-5 times the length of the calyx, 25-35 mm long and ca. 2 mm broad; shallowly canaliculate dorsally, bluntly triquetrous, bilocular, covered with appressed black and white hairs. May-June. Southern, mostly stony, mountain slopes and bluffs, rarely steppe meadows.— W. Siberia: Alt., Irt.; Centr. Asia: Dzu.- Tarb., Balkh. Endemic. Described from Altai. Type in Leningrad. 664. A. polytimeticus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 7-15 cm tall, shortly ligneous at base, silvery-white, the branches decumbent; stems of the year 1-5 (8) cm long, very short, densely covered with short appressed white hairs; stipules triangular, acute, 0.5-1 mm long, white-tomentose; leaves 1-3 (4) cm long, the rachis equaling to twice as long as the petiole, both petiole and rachis covered with appressed white hairs; leaflets (4) 5-6 pairs, linear, 4-7 (10) mm long, 0.5-1 mm broad, subobtuse, covered on both sides with appressed white hairs, silvery-gray; peduncles 2-3 times the length of the leaves, 5-10 cm long, slender, terete, covered with appressed white and upward sometimes mixed black and white hairs; racemes very short, subumbellate, 2-4 (5)-flowered; bracts oblong, acute, 0.5-1 mm long, covered with black and white hairs; calyx cylindric, (10) 11-12 mm long, covered with appressed black and white hairs; the linear teeth 1-2 mm long, black~hairy; corolla pale violet; standard 20-22 mm long, the limb oblong-obovate, short-attenuate upward, retuse, narrowed in lower 1/5, angular at base 4-5 times as long as the claw; wing-petals 18-20 mm long, the limb lanceolate-oblong, entire, acutish, the claw 1.5 times as long as the limb; keel 15-16 mm long, the limb semioval, acutish, half as long as the claw; pods sessile, subulate-linear, 20-22 mm long, 1.5-1.8 mm broad, straight or scarcely falcate, spreading or pendulous, carinate ventrally, canaliculate dorsally, terminating in a slender subulate beak 2-3 mm long, covered with appressed white hairs, these sometimes interspersed with few black hairs. Fl. May-August; fr. August-September. 518 695 In juniper groves, to the upper limit of the juniper zone.— Centr. Asia: Pam.- Al. (Upper Zeravshan basin). Endemic. Described from the vicinity of Kuli-kalon. Type in Leningrad. 665. A. stenoceras C.A.M. in Bong. et Mey. Verzeichn. (1841) 24; Ldb. Fl. Ross. I (1842) 629; Bge. Astrag. geront. II (1869) 221 et Astrag- turk. (1880) 287; Kryl. Fl. Zap. Sib. VII (1933) 1699.— A. subulatus altaicus Pall. Astrag. (1800) 23, var.B.—A. ceratoidesa. campestris Ldb. Fl. alt. III (1831) 306.—A. ceratoides Bge. Enumer. Alt. (1836) 71, non M.B.—Ic.: Pall. l.c. t. 20 A. Perennial, suffrutescent, 20-25 cm tall; caudices 1-5 cm long, mostly slender, covered with castaneous-brown bark, woody; stems of the year 3-12 cm long, densely appressed hairy, whitish, slender; stipules free, ovate-lanceolate, 1.5-2 mm long, covered with appressed white or sometimes mixed white and fewer black hairs; leaves 2-5 cm long, the petiole and the much longer rachis slender, covered with appressed white hairs; leaflets 5-7 pairs, oblong-elliptic or lanceolate or linear, obtuse to acute, 4-10 mm long, 1-2 mm broad, appressed-hairy on both sides; peduncles (1.5) 2 times as long as or rarely about equaling the leaves, covered with appressed white hairs; racemes abbreviated, umbelliform, 2-2.5 cm long, 4-8 (10)-flowered; bracts broadly ovate, 1-1.5 mm long, shorter than to about equaling the flowers, black-tomentulose; calyx cylindric, 9-10 mm long, densely covered with white and black hairs, the subulate teeth 1-1.5 mm long; corolla lilac; standard 20-22 (24) mm long, the limb oblong-obovate, retuse, rather abruptly narrowed in lower 1/3, slightly angular at base, 2-3 times as long as the claw; wing-petals 17-20 (22) mm long, the limb oblong, retuse or subentire, as long as the claw; keel 16-19 (20) mm long, the limb obovate, obtusish, the claw 1.5-2 times as long as the limb; pods borne on a stipe 0.5 mm long, approximate, obliquely ascending to spreading, subulate-linear, (2) 2.5-3.5 cm long, 2-2.5 mm broad, carinate ventrally, convex dorsally, straight or scarcely curved, acuminate into a straight beak 1.5 mm long, coriaceous, densely covered with appressed white and black hairs, incompletely bilocular. Fl. May-June; fr. June. Steppes and steppe-like meadows, rarely sands in the semidesert and desert regions; stony mountain slopes. W. Siberia: U. Tob. (S.), Irt. (S.), Alt.; E. Siberia: Ang.-Say. (Minusinsk, Balagansk and Verkholensk steppes); Centr. Asia: Ar.-Casp. (N.), Balkh (N.), Dzu-Tarb. (Tarbagatai). Endemic. Described from Zaisannor. Type in Leningrad. 666. A. substenoceras Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, ca. 30 cm tall, woody at base, from divisions of the caudex; stems 12-15 cm long, white, finely sulcate, 2-3 mm thick; stipules 1-2 mm long, the upper ones green, herbaceous, lanceolate, free, covered with appressed mainly white hairs; leaves 2-4 cm long, the petioles and the leaves white by a dense coat of appressed hairs; leaflets 8-11 pairs, oblong to oblong-ovate, 5-10 mm long, 2 mm broad, scarcely retuse or mucronulate; peduncles 2-3 times as long as the leaves; racemes capitate, 5-8-flowered; bracts linear, 1-1.5 mm long; pedicels ca. 1 mm long; calyx cylindric, ca. 11 mm long, the subulate teeth 2 mm long; corolla violet (?); standard 20 mm long, the limb oval, gradually attenuate toward base, round-tipped; wing-petals 17 mm long, the limb 7 mm long, entire, 919 696 the claw 1.5 times as long as the limb; keel 15 mm long, the limb 5 mm long, ovate, the claw twice the length of the limb; ovary linear; pods straight at first, appressed-hairy, linear, ca. 1-1.5 mm broad, acuminate, subulate-beaked. Fl. June; fr. June-July. Stony slopes, at altitudes of 2000-2300 m.— Centr. Asia: Pam.-Al. (Darvaz). Endemic. Described from Vanch. Type in Leningrad. 667. A. Karelinianus M. Pop. nomen novum.— A. virescens Kar. ex Ldb. Fl. Ross. 1 (1842) 630 (?).— A. macrolobus f.nigropilosus Bge. in Mém. Sav. Etr. Pétersb. VII (1851) 261; Bge. Astrag. geront. II, (1869) 222 (ex p. quoad BM Uralenses).— A. ortholobus Bge., l.c., 223 (quoad pl. Ural.). Perennial, 17-30 (40) cm tall, the caudex buried, its divisions ligneous aerial 1-4 cm long; stems of the year many, up to 10-20 cm long, erect or rarely ascending, rather sparsely covered with appressed white hairs; stipules free, lanceolate, the lower oblongly triangular-ovate, 3-4 mm long, covered with appressed white or mixed white and black hairs; leaves 5-10 cm long, the petiole 1.5 (2) cm long, the petiole and the rachis slender, covered with appressed white hairs; leaflets 5-7 pairs, lanceolate or lance~ linear or linear-oblong, acute, rarely subobtuse; peduncles 10-20 cm long, (1.5) 2 times the length of leaves, slender, angled, sparsely covered with appressed white hairs; racemes capitate, 2-3 cm long, rather loosely 5-10-flowered; bracts triangular-ovate, 0.5-1 mm long, hyaline, covered with white and black hairs; calyx cylindric, 7-9 mm long, covered with appressed mostly black and fewer or rarely more numerous white hairs, the subulate teeth ca. 1 mm long; corolla whitish or pale-violet or violet; rarely ochroleucous; standard 18-21 mm long, the limb oblong-obovate; slightly and broadly emarginate, rather gradually narrowed in lower ter scarcely angular at base, 6-7 times as long as the claw; wing-petals (14) 16-20 mm long, the limb oblong, scarcely retuse, as long as the claw; keel (11) 13-15 mm long, the limb broadly semiorbicular, obtuse, the claw 1.5 times as long as the limb; ovary borne on a stipe ca. 0.5 mm long; pods sessile, obliquely ascending to horizontally spreading, subulately linear- oblong, 20-30 mm long, 2.5-3 mm broad and thick, slightly curved, acute, terminating in a subulate beak 1-2 cm long, carinate ventrally, thickened and deeply grooved dorsally, coriaceous, covered with appressed white hae often interspersed with shorter black hairs, bilocular. Fl. June-July; . (June) July. Verea ee (mixed grass and needlegrass, or mixed grass, fescue and needlegrass), often on more or less stony ground; outcrops of sandstone, conglomerates, and limestone. — European part: V.~Ka. (Centr. and 5S. Ural). Endemic. Described from the vicinity of Verkhoturskii Factory. Type in Leningrad. 668. A. artvinensis M: Pop. in Not. Syst. Inst. Bot. Ac Sc. URSS X (1946). Perennial, suffrutescent, ca. 34 cm tall; caudices 7-8 cm long, covered with castaneous-brown bark, branched; stems of the year erect, 10-15 cm long, terete, slender, 1-2 mm thick, whitish by short appressed hairs; stipules free, triangular-oblong to lanceolate, 1-1.5 mm long, covered with appressed white hairs; leaves 4-6 cm long, the petiole 3-7 mm long, the petiole and the rachis slender, covered with appressed white hairs; leaflets 5-6 (7) pairs, oblong to linear-oblong, 5-10 mm long, 2-3 mm broad, 520 697 obtuse, covered scatteredly above and densely beneath with appressed hairs, hoary beneath; peduncles 8-10 cm long, 2-2.5 times the length of the leaves, thick, angled, erect, covered with appressed white hairs; inflorescence capitate, 5-10-flowered, rounded-oval, 3-4 cm long; bracts lanceolate, 2-4 mm long, villous with white and black hairs; calyx cylindric, 15 mm long, covered with short black hairs and more numerous white spreading hairs, the filiform teeth ca. 5 mm long, covered with black hairs; corolla violet (?); standard 24-25 mm long, the limb oblong- obovate, short-attenuate at apex and retuse, narrowed in lower 1/3, angular at base; wing-petals 22 mm long, the limb spatulate-oblong, round-tipped, the claw 1.5 times as long as the limb; keel 19 mm long, the limb semiorbicular, acute, half as long as the claw; pods sessile, obliquely ascending, slightly arcuate, 20-25 mm long, 3 mm broad, scarcely carinate ventrally, broadly and rather shallowly canaliculate dorsally, subterete, acute, covered with appressed white hairs, the slightly spreading beak 1-2.5 mm long. Fl. May; fr. June. Taluses.— Caucasus: W. Transc. Endemic. Described from a place between Artvin and Irse. Known only from this location. Type in Leningrad. 669. A. haesitabundus Lipsky.in A.H.P. XIII (1894) 290; Schmalh. Fl. I, 283.— A. xiphidium 8. distans Lipsky in Zap. Kiev ob-va estestv. XI, 2, (1888) 45.— A. Alboffianus Freyn in Bull. Herb. Boiss. II (1895) 186. Perennial, (15) 20-40 cm tall, the divisions of the buried caudex 0.5 -1 cm long, slender, ligneous; stems of the year many, (5) 10-30 cm long, erect or assurgent, rather sparsely covered with appressed white hairs; stipules triangular-lanceolate, 2-3 mm long, covered with short white or mixed white and black hairs; leaves 4-6 (8) cm long, the rachis 3-5 times as long as the rachis, both petiole and rachis slender, covered with scattered appressed white hairs; leaflets 4-6 (8) pairs, oblong or more often narrowly linear, rarely oval-oblong or oblong-oval, (0.7) 1-2 cm long, 2-3 (5) mm broad, glabrous or sparsely hairy toward the margin above, rather densely covered beneath with appressed hairs; peduncles 1.5-2 times and together with inflorescence 2-4 times as long as the leaves, sparsely covered with short appressed white hairs, finely sulcate; racemes loose, oblong, remotely flowered, 6-10 cm long; bracts ovate-lanceolate, 1-2 mm long, hyaline, white- and black-ciliolate; calyx cylindric, 7-9 mm long, densely covered with appressed black and white hairs, the lance- subulate teeth 1-2 mm long, the tube 4-6 times as long as the teeth; corolla violet; standard 17-19 mm long, the limb oblong-obovate, slightly retuse, abruptly narrowed in lower 1/4, angular at base, 5-6 times as long as the claw; wing-petals 15-16 mm long, the limb linear-oblong, slightly retuse, as long as the claw; keel 12-13 mm long, the limb triangularly obovate-oblong, obtusish, the claw 1.5 times as long as the limb; pods sessile, horizontally spreading or obliquely descending, subulate-linear, straight or slightly curved, (20) 30-40 mm long, 2-2.5 mm broad, s terminating in a subulate beak 2-3 mm long, carinate ventrally, broadly grooved dorsally, coriaceous, covered with appressed mixed white and black or rarely almost pure white or black hairs, bilocular. Fl. May-July; fr. June-August. Dry beardgrass-dominated steppes, in grass-and-wormwood associations, on clayey bluffs, limestone outcrops, pebbles, and stony mountain slopes, 521 699 at altitudes of 500-1500 (2100) m.— Caucasus: Ciscauc., Dag. Endemic. Described from the vicinity of Chir-Yurt in Daghestan. Type in Leningrad. 670. A. Reverdattoanus Sumn. in Animadvers. Herb. Univ. Tomsk. Nosaiono .(09s3) Gr Perennial, cinerescent; caudex buried, short, strong, with short divisions; aerial stems many, diffuse, 2-6 cm long, ligneous at base, densely covered with appressed white hairs; stipules free, ovate-lanceolate, obtusish, glabrate; leaves 10-12 cm long, the petiole 0.5-1 cm long; leaflets 5-8 pairs, oblong, acute, 5-7 mm long, 2-3 mm broad, appressed- hairy on both sides; peduncles about equaling the leaves; racemes up to 8 cm long; bracts linear-lanceolate, twice the length of the pedicel, hyaline, ciliate-margined; calyx cylindric, 10-14 mm long, covered with appressed black and fewer white hairs, the tube 4-5 times as long as the subulate teeth; corolla whitish-violet; standard 20-24 mm long, the limb oblong- lanceolate, retuse; wing-petals ca. 20 mm long, the limb lanceolate, retuse; keel intensely violet-tipped; ovary sessile, covered with appressed white hairs, 17-22-ovuled. Fl. June. Alpine meadows.— Centr. Asia: Tien Shan (W.: Kugart River valley). Endemic. Described from Kugart River valley. Type in Tomsk. 671. A. corniculatus M.B. Cent. pl. rar. Ross. mer. (1810) p. XLV; NiBeLbl taunsscaue. il (Lel9)"492) Edb.whl) Ross. l, Gol DC eRoercer II, 284; Bge. Astrag. geront. I, 126:II, 222; Schmalh. Fl. I, 283;—Ic. :M.B. Cent. pl. rar. Ross. mer. (1810) tab. 45. ‘ Perennial, suffrutescent, 8-20 cm tall; caudex fairly slender, buried, the divisions slender, not exceeding 2 mm in diameter, ligneous, (1) 2-3 (8) cm long, ascending, rarely erect, castaneous-brown; stems of the year (1) 3-6 (10) cm long, assurgent to erect, slender, rather sparsely covered with appressed white hairs; stipules free, ovate-lanceolate to ovate, 2-3mm long, covered with appressed white hairs; leaves 4-7 cm long, the petiole and the much longer rachis very slender, subfiliform, covered with appressed white hairs; leaflets 6-10 pairs, narrowly linear, 5-10 mm long, 0.6-1 mm broad, subobtuse, appressed-hairy on both sides, hoary; peduncles somewhat shorter to somewhat longer than the leaves, slender, sparsely covered with appressed white hairs, these sometimes interspersed beneath the inflorescence with black hairs, racemes umbelliform~-capitate, 1.5-2 cm long, 3-5 (10)-flowered; bracts ovate to ovate-lanceolate, ca. 1 mm long, somewhat shorter thari the pedicel, covered with black or mixed black and white hairs; calyx cylindric, 8-10 (12) mm long, covered with subappressed black hairs, the subulate teeth 2-3 mm long, the tube 3-4 -times as long as the teeth; corolla pinkish-violet or white; standard 17-20 mm long, the limb subrhomboidally oval, slightly and broadly emarginate, abruptly narrowed in lower 1/4, angular at base, 3 times as long as the claw; wing-petals 15-17 mm long, the limb oblong, scarcely or slightly retuse, as long as theclaw; keel 11-13 mm long, the limb broadly semiobovate, obtusish, the claw 1.5 times as long as the limb; ovary borne on a stipe ca. 0.5 mm long; pods sessile, obliquely ascending or horizontally spreading, subulate-linear, slightly curved, 20-28 mm long, 2-2.5 mm broad, carinate ventrally, scarcely canaliculate dorsally, abruptly narrowed into a straight slenderly subulate beak 1.5-2.5 cm long, coriaceous, rather sparsely covered with appressed white hairs, bilocular. Ey Wlchya heer nes 522 700 Steppes, clayey and stone-and-clay-covered slopes. — European part: M. Dnp. (S.: Baltic region)., Bes., Bl., Crim.; Caucasus: S. Transc. Endemic.— Described from the southern belt of the European part of the USSR. Type in Leningrad. Note. A.A. Grossheim reports A. corniculatus for the Northern Caucasus (Kub.-Chrn.) in the Flora of the Caucasus (Vol. II, p.320). All specimens from N. Caucasus present in the herbarium of the Botanical Institute of the Academy of Sciences belong to A. subulatus M.B. which is distinguishable from A. corniculatus M.B. by its remotely flowered, more or less elongated raceme. The herbarium contains the following specimens from the Caucasus which do not differ morphologically from the typical A. corniculatus M.B.; 1) Fl. Cauc.-Armen. Ani 1838, No. 167 (Koch), 2) Berg Bogutli in der Nahe des Alages in Armenien, (Lagowski). 672. A. subulatus MB. Fl. taur.-cauc. II (1808) 193; DC. Prodr. II, 284; Ldb. Fl. Ross. I, 631; Bge. Astrag. geront. II, 871; Fl. Yugo- Vost. V, 603; Grossh. Fl. Kavk. II, 320.—Philammos subulatus Stev in Bull. Soc. Nat. Mosc. XXIX (1856) 146.— Exs.: H.F.R. No. 868. Perennial, suffrutescent, 15-22 cm tall; caudex buried, fairly stout, the aerial divisions i-5 (9) cm long, ascending to subdecumbent, covered with thin castaneous-brown bark; stems of the year 1-5 cm long, ascending to erect, whitish by a dense coat of short appressed white hairs; stipules connate at base (the lower ones up to the middle), triangular-ovate, 1.5-2 mm long, covered with appressed black and white hairs; leaves (2.5) 3-6 cm long, the petiole 3-4 times as long as the rachis, both petiole and rachis very slender, covered with short appressed white hairs; leaflets 5-6 (7) pairs, narrowly linear to subfiliform, (4) 6-13 mm iong, 0.7-1 (1.5) mm broad, subobtuse, glabrous above, covered beneath with short appressed hairs; peduncles equaling to slightly exceeding or up to 1.5 times as long as the leaves, slender, covered with short appressed white and, beneath the inflorescence, often mixed white and fewer black hairs; racemes 2-5 (7) cm long, remotely (3) 5-6-flowered; bracts ovate, ca. 1 mm long, covered with short black and fewer white hairs; calyx cylindric, 8-10 mm long, covered with black or mixed black and few white hairs, often striped, the subulate teeth 1 (1.5) mm iong; corolla purple; standard (17) 19-23 mm long, the limb oblong-obovate, attenuate toward apex, retuse, abruptly narrowed in lower 1/4-1/3, angular at base, 5-6 times as long as the claw; wing-petals (14) 16-20 mm long, the limb linear-oblong, entire, obtuse, as long as the claw; keel (12) 14-17 mm long, the limb obovate, obtuse, somewhat shorter than the claw; ovary ona stipe ca. 1 mm long; pods borne on a stipe ca. 0.5 mm long, obliquely ascending, rarely horizontally spreading, 1.8-2.7 cm long, 1.5-2 mm broad, cylindric, with convex ventral and concave dorsal suture or scarcely carinate ventrally, scarcely canaliculate dorsally, subcylindric, attenuate into a straight beak 1-1.5 mm long, coriaceous, densely covered with short appressed or rarely ascending white hairs, incompletely bilocular. Fl. May-(June); fr. June-July (August). Steppes and stony calcareous mountain siopes.— European part: Bes., Bie Me Dip., V-=sDon, “L: /Don, wl..V-),-Dransv.; Crim, ; (Caucasus: Cisc. E. and §. Transc.; W. Siberia: U. Tceb. Endemic (?). Described from Crimea. Type in Leningrad. 923 701 673. A. ucrainicus M. Pop. et M. Klok. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, (12) 15-27 cm tall; caudex buried, the aerial divisions 0.5-1 cm long; stems of the year 1-5 cm long, slender, ascending, whitish by dense coat of appressed hairs; stipules connate at base or in lower part, triangular-ovate, acute, ca. 1 mm long, covered with white or mixed black and white hairs, adnate to petiole in lower part; leaves 3-5.5 cm long, the petiole and the mostly much longer rachis very slender, subfiliform, covered with short appressed white hairs; leaflets 5-7 pairs, narrowly linear to subfiliform, 4-10 (16) mm long, 0.5-1 (2) mm broad, subobtuse, glabrous above, appressed-hairy beneath; peduncles equaling to somewhat exceeding or rarely up to 1.5 times as long as the leaves, slender, sparsely covered with short appressed white hairs; racemes loose, 4-8 cm long, very remotely (4) 6-10-flowered; bracts ovate, ca. 1 mm long, 2/3 as long as to equaling the pedicel, covered with very short black and fewer white hairs; calyx cylindric, 8-10 mm long, covered with appressed black and fewer white hairs, the subulate teeth 1-1.5 mm long; corolla lurid; standard 18-22 mm long, the limb oblong-obovate, gradually attenuate toward apex, retuse, rather abruptly narrowed in lower ee angular at base, 4-5 times as long as the claw; wing-petals 16-20 mm long, the limb linear-oblong, entire, obtuse, as long as the claw; keel 13-16 mm long, the limb semioval, obtuse, half as long as the claw; ovary borne on a stipe ca. 1 mm long; pods essentially sessile, obliquely ascending, rarely horizontally spreading, subulately linear, slightly arcuate, 2-3.4 cm long, 1.5-2 mm broad and thick, more or less rounded dorsally and ventrally, cylindric, attenuate into a subulate beak ca. 1 mm long, coriaceous, densely covered with short appressed white hairs, partially or incompletely bilocular (with a hyaline flange along the dorsal suture). Fl. June-July; fr. August (September). Chalk outcrops, rarely in steppes.— European part: M. Dnp., V.~Do, Transv., L. Don, Bl. Endemic. Described from region of Dnepropetrovsk. Type in Leningrad. 674. A. pseudotataricus Boriss. nom. nov. A. subulatus y tataricus DC. Prodr. II (1825) 284; Ldb. Fl. Ross.I (1842) 631.— A. tataricus M.Pop. inherb., non Franch. (1883). Perennial, resembling A. ucrainicus M.Pop. et M. Klok., the distinguishing characters being the longer calyx (11-13 mm), with filiform- subulate teeth 1.5-2 mm long, and the white or violet corolla; pods straight, (2.5) 3-4 cm long. Fl. May; fr. June-July. Steppes, often saline, rarely sandy. — European part: Bl., L. Don, L.V.; Caucasus: Cisc. Endemic. Described from vicinity of Novocherkassk. Type in Leningrad. Series 9. SOGOTENSES Vass. — Herbaceous perennials; stems short; leaflets 2-3 pairs, linear; racemes very long (up to 20-30 cm ), loose, remotely flowered; corolla purple, 14 to 20 mm long; pods subulate-linear, 20-28 mm long and 2-3 mm broad. Pebbles of riverside sands in Centr. Asia. 675. A. sogotensis Lipsky in A.H.P. XXVI (1907-1909) 178. — Perennial, woody at base, 25-32 cm tall; stems 4-10 cm long, white by a dense coat of appressed hairs; stipules partially adnate to petiole, 524 triangular, 1.5-2 mm long, white-hairy; leaves 6-14 cm long, the petiole as long as the rachis, covered with appressed white hairs; leaflets 2-3 pairs, linear, 12-18 mm long, 1.5-2 mm broad, covered on both sides with appressed hairs, subobtuse: peduncles equaling to twice as long as | the leaves, rather densely covered with appressed white hairs, more or /702 less hoary, firm, subvirgate; racemes (7) 14-18 cm long, remotely 6-9-flowered; bracts ovate, more or less acuminate, 2-2.5 mm long, white-tomentose; calyx cylindric, 8-10 mm long, rather densely covered with appressed white hairs, the tube 3-4 times as long as the lance-linear teeth; corolla purple (?); standard ca. 20 mm long, the limb oval, sharply notched, twice as long as the claw; wing-petals ca. 17 mm long, the limb oblong, entire, as long as the claw; keel ca. 14 mm long, the limb broadly obovate, the claw 1.5 times as long as the limb; ovary subsessile; pods sessile, narrowly linear, 21-23 mm long, 2 mm broad, carinate ventrally, slightly canaliculate dorsally, compressed-triquetrous, slightly curved, gradually attenuate into a subulate beak 2-2.5 mm long, coriaceous, covered with scattered short appressed white hairs, bilocular. Fl. April: ir. April-May. Sands.— Centr. Asia: Balkh. (sands by the river Ili), Tien Shan. Endemic. Described from the vicinity of Sogota. Type in Leningrad. 676. A. pischtovensis Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, nearly acaulescent, 35-43 cm tall; caudex buried, very short, with many short divisions: stems of the year 3-5 (7) cm long, densely leafy, whitish-hoary by a dense coat of appressed white hairs; stipules adnate at base to petiole, broadly short-triangular, 1.5-2 mm long, short-acuminate, covered with appressed white hairs; leaves 6-8 cm long, the rachis 2-3 times as long as or rarely about equaling the petiole, both petiole and rachis covered with appressed white hairs, finally indurescent but not spinescent; leaflets 2-3 pairs, narrowly linear, 9-16 (20) mm long, 1-1.5 mm broad, densely hoary on both sides by appressed short white hairs; peduncles equaling to 1.5 times as long as the leaves, fairly slender but firm, virgate, densely covered with short appressed white hairs, hoary; racemes very long, loose, 20-32 cm long, the flowers very distant; bracts oblongly triangular-ovate, 1-1.3 mm long, short-acuminate, white-ciliolate; calyx cylindric, 10-12 mm long, densely covered with short appressed white hairs, the tube 1.5 times as long as the filiform-subulate teeth; corolla purple; standard 14-15 mm long, the limb oblong-obovate, slightly retuse, ovate in upper 2/3, abruptly narrowed below, angular at base, 4-5 times as long as the claw; wing-petals barely exceeded by the standard, the limb linear, entire, somewhat shorter than the claw; keel 12-13 mm long, the limb oblong, slightly curved, rounded-obtuse, 5-5.5 mm long, 1.5 mm broad, the claw somewhat longer than to 1.5 times as long as the limb; ovary subsessile; pods subsessile (the stipe ca. 0.5 mm long), cernuous, subulate-linear, straight, 20-27 mm long, ca. 2 mm broad and thick, broadly carinate ventrally, broadly canaliculate dorsally, 03 subtriquetrous, terminating in a straight slender subulate beak 2-3 mm long, coriaceous, densely covered with short appressed white hairs, incompletely biiocular. Stony slopes of elevated foothills and riverside pebbles in the upper part of the ephemeral vegetation zone, up to 1650 m above sea level. — Centr. | 525 704 Asia: Pam.-Al. (Darvaz region). Endemic. Described from Kyzyl-su river (from vicinity of Pishtovavillage). Type in Leningrad. Series 10. LEPTOPI Vass.— Subshrubs of the pulvinate type; leaflets 10-13 pairs, small, linear; racemes elongate, loose; corolla pale pink, relatively small (15-17 mm long). Rocks and stony slopes of W. Tien Shan. 677. A. Michaelis Boriss. nom. nov.— A. leptopus M. Pop. in Bull. Univ. As. Med. 14 (1926) 142, non Benth. (1853). Perennial, suffrutescent; caudex much divided, the divisions up to 100 in number, imparting to the plant the cushionlike aspect, covered with brownish fibrillose bark; stems erect, 5-10 cm long, slender, hoary by appressed white hairs; stipules free, small, oblong; leaves 5.5-11 cm long, the petiole very short, the slender rachis marcescent; leaflets 10-13 pairs, linear to linear-oblong, 5-8 mm long, 1-1.5 mm broad, subobtuse, green, covered on both sides with scattered hairs; peduncles many, slender, erect, firm, together with rachis 10-12 cm long, 1.5~-2 times the length of the leaves; racemes loose, 5-10 cm long, 5-8-flowered; bracts ca. 1 mm long, about equaling the pedicel, covered with appressed black hairs; calyx cylindric, ca. 10 mm long, densely covered, especially near the base, with appressed black hairs, the tube 3 times as long as the subulate teeth; corolla pale pink, 15-17 mm long; pods (young) exceeding the calyx, densely covered with appressed black hairs. Fl. July-August. Rocks and stony mountain slopes, ascending up to the subalpine zone. — Centr. Asia: W. Tien.Shan. Endemic. Described from rivers Ugam and Pskem and from Mts. Karatau. Type in Tashkent. 678. A. leptopoides M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 10-20 cm tall; caudex woody, strong, buried, the short dividions forming fairly dense mats; stems of the year 1-4 cm long, abbreviated, densely hoary appressed short white hairs; stipules adnate to petiole in lower half or one-third, hyaline, the upper ones often green herbaceous, 2-3 mm long, the free portion triangular-lanceolate, acute; leaves 4-7 cm long, the petiole about equaling the leaves, slender, hoary; leaflets 6-10 pairs, linear to lance-linear, subobtuse, 3-7 mm long, 1-2 mm broad, hoary on both sides by short appressed white hairs; peduncles slender, virgate, up to 3.5-6-15 cm long, equaling to 1.5 times as long as the leaves, covered with short white hairs; racemes loose, (4) 5-10 cm long; flowers 3-6, very distant; bracts ovate to ovate- lanceolate, whitish, ca. 1 mm long, acute, sparsely covered with short black and white hairs; pedicels ca. 1 mm long; calyx ca. 10 mm long, the tube 4 times as long as the teeth; corolla pale violet; pods sessile, erect, oblong-linear, straight, terminating in a long straight beak, together with beak 15 mm long, 2 mm broad, sharply carinate ventrally, witha broad and closed groove on the back, covered with appressed white and black hairs. Fl. June-July; fr. July. Stony and gravelly slopes.— Centr. Asia: Tien Shan (Karatau Range). Endemic. Described from Uzak-uya natural landmark. Type in Tashkent. Series 11. VIMINEI Vass.7— Shrubs or tall subshrubs (from 20-30 to 90-100 cm tall); leaflets narrow, elongate; racemes loose, elongated or short to capitate; corolla violet or lilac; pods from 10 to 20 mm long and (3.5) 4-6 mm broad, erect. Steppes and steppe~dominated mountain associations. 526 705 679. A. cornutus Pall. It. I, App. (1771) 499; Kryl. Fl. Zap. Sib. VII 1694.— A. vimineus Pall. Astr. (1800) 24; DC. Prodr. II, 286; Bge. Astrag. geront. II, 217; Boiss. Fl. or. II, 479; B.A. Fedchenko, Fl. Yugo-Vost. I, 601; Grossh. Fl. Kavk. II, 323 et auct. pl. Fl. Ross.— A. fruticosus Ldb. Fl. Ross. I (1842) 632 pp. Frutescent, 30-50 cm tall; caudices (5) 10-30 cm long, covered with thin grayish-brown peeling bark, the divisions fairly slender, erect; stems of the year (3) 5-25 cm long, sulcate, covered with whitish peeling thin smooth bark, sparsely covered with appressed white hairs; stipules free, adnate to petiole to 1/3-1/2, oblongly triangular-ovate or ovate-lanceolate or lanceolate, acuminate, 4-6 mm long, sparsely appressed-hairy, the lower ones triangular-ovate; leaves (3.5) 5-9cm long, sessile, the slender rachis sparsely covered with short appressed or ascending white hairs or glabrate; leaflets (5) 6-9 pairs, mostly narrowly linear or less often lance- linear, acute, rarely lance-elliptic and obtuse, 10-30 mm long, 1.5-3 (4) mm broad, glabrous above, covered beneath with short appressed white hairs; peduncles equaling the leaves, firm, sulcate, sparsely covered with short appressed white hairs; racemes capitate, contracted, (2)3-4cm long, 10-20-flowered, the rachis (8) 10-15 mm long; flowers subsessile; bracts linear-lanceolate, 3-5 mm long, exceeding the pedicel, covered with white and black hairs; calyx cylindric, 10-11 mm long, the linear-subulate teeth 1.5-2.5 mm long, the tube 3-4 times as long as the teeth; corolla violet- red; standard 18-20 mm long, the limb oblong-obovate, short-attenuate toward apex, scarcely retuse or rarely obtuse, abruptly narrowed in lower 1/4, slightly angular at base, 4-5 times as long as the claw; wing-petals 16-18 mm long, the limb linear-oblong, entire, obtuse, as long as the claw; keel 14-16 mm long, the limb nearly semicircular, slightly concave beneath, obtusish, the claw 1.5-2 times as long as the limb; ovary borne on a stipe ca. 0.5 mm long; pods sessile, capitately crowded, linear- oblong, straight, 10-16 mm long, 3-3.5 mm broad, abruptly contracted into an obliquely divergent straight subulate beak 2-4 mm long, rounded or scarcely carinate ventrally, flattened and scarcely canaliculate dorsally, coriaceous, villous with ascending white hairs interspersed with very few short black hairs, bilocular. Fl. (May) June-July; fr. July-August (September). Mainly in the steppe or less often in the forest-steppe region, in steppes, forest-steppe glades, and interzonal pine woods, on limestone, clayey escarpments, or chalk; ascending in the Caucasus up to 2000 m, fairly frequently in the foothill or on rocks in the mountains, less often on sandy banks.— European part: M. Dnp., Bl., L.Don., L.V.; Caucasus: Cisc., E. and S. Transc.; W. Siberia: U. Tob., Irt.; Centr. Asia: Ar.-Casp., Dzu.- Tarb. Endemic. Described from Lower Volga. Type in Leningrad. 680. A. brachylobus DC. Prodr. II (1825) 285; Bge. Astrag. geront.I (1868) 124; II (1869) 216; Astrag. turk. (1880) 289; Boiss. Fl. or. II (1872) 478; Grossh. Fl. Kavk. II (1930) 323; B.A. Fedchenko, Fl. Yugo- Vost. V (1931) 601. Suffrutescent, (35) 40-70 cm tall; caudices divided from base, (5) 10-30 cm long, up to 5 mm thick, covered with castaneous-brown longitudinally splitting bark; stems (5) 10-20 cm long, hoary by appressed white hairs, finally lignifying in lower part, covered with lustrous white thin peeling bark; stipules free, adnate at base to petiole, 3-5 mm long, acuminate, covered with subappressed white or mixed white and black hairs; 527 706 leaves 4-10 cm long, the petiole and the somewhat longer rachis slender, more or less covered with appressed white hairs; leaflets linear to lance- linear, rarely linear oblong, (10) 15-30 (40) mm long, (1) 1.5-4 mm broad, subobtuse, covered on both sides, often thinly above, with appressed hairs; peduncles equaling to 1.5 times as long as the leaves, covered with short appressed white and, beneath the inflorescence, sometimes mixed white and black hairs; racemes loose, 6-10 (14) cm long, the flowers remote, fairly numerous; bracts lanceolate to linear-lanceolate, 2-4 mm long, covered with short black or mixed black and white hairs, 2-4 times as long as the pedicel calyx cylindric, 12-15 mm long, densely villous with subappressed white and black or rarely pure white hairs, the tube (3) 4-5 times as long as the filiform-linear teeth; corolla violet; standard (20) 22-26 mm long, the limb oblong-obovate, scarcely retuse, abruptly narrowed toward base, angular or auriculate-angular at base, 4-5 times as long as the claw; wing-petals (18) 21-25 mm long, the limb linear- oblong, round-tipped, the claw 1.5 times as long as the limb; keel (15) 18-22 mm long, the limb semioval, obtusish, half as long as the claw; ovary sessile; pods spreading at a right angle, sessile, linear-oblong to oblong, 11-16 mm long, (3.5) 4mm broad, more or less compressed laterally, strongly carinate ventrally, grooved dorsally, short-acuminate into a straight subulate rigid obliquely upturned beak 2-4 mm long, coriaceous, covered with appressed or ascending white or mixed white and black hairs, bilocular or incompletely so, Fl. May-August (September); fr. June-September. Chalk or sand, rarely steppes; dry escarpments.— European part: Crim., L. Don, L.V., Transv.; Caucasus: Cisc., Dag.; W. Siberia; U. Tob., Irt.; Centr. Asia: Ar.-Casp. Endemic. Described from U. Tob. (banks of Ishim). Type in Geneva. Note. A. barbidens Freyn, described from Daghestan (in Bull. Herb. Boiss. III, 1895, 185) is probably a form of this species. 681. A. fruticosus Pall. Astrag. (1800);21; Bge. Astrag. geront.I, 124; II, 216; Krylov, Fl. Zap. Sib. VII, 1693.—A. odessanus Bess. Enum. pl. Volh. (1820) 30.— A. suffruticosus Pall. Reise I (1766) 140; DC. Astrag. 103.—Ic.: Pallas, Astrag. (1800) t. XIX. Frutescent or rarely suffrutescent, (10) 20-60 cm tall; caudices rather thick, divided, erect, (4) 15-40 cm long, covered with dark brown peeling bark; stems of the year 7-20 cm long, borne on abbreviated shoots, (1) 3-5 cm long, slender, covered with appressed white or mixed white and black hairs; stipules free, lanceolate, the lower ovate-triangular, 2-4 mm long, covered with white or mixed black and white hairs; leaves (2) 4-7 cm long, the very short petiole and the rachis sparsely covered with short appressed white or mixed white and black hairs; leaflets (5) 6-8 pairs, lance-linear or linear-lanceolate, rarely oblong or linear, mostly acute, rarely obtuse, (5) 10-20 (30) mm long, (1) 2-3 (4) mm broad, sparsely covered on both sides, though more densely and sometimes exclusively beneath, with appressed hairs; peduncles (1) 4-10 cm long, equaling to 1.5-2 times as long as the leaves, covered with white and, beneath the inflorescence or rarely throughout, black hairs, the white hairs appressed, the black ones ascending; racemes capitately umbelliform, short (the rachis 1-5 mm long), (3) 5-8-flowered, 2-2.5 cm long; bracts lanceolate, 1-2 mm long, black-hairy; calyx cylindric, (8) 10-11 mm long, tomentulose with black spreading hairs and villous with longer white hairs, the linear- 528 707 708 subulate teeth 1-2 mm long; corolla reddish-lilac; standard (18) 20-23 mm long, the limb oblong-obovate, retuse, often slightly narrowed in lower 1/3, slightly angular at base, 4 times as long as the claw; wing-petals (15) 18-20 mm long, the limb linear-oblong, retuse, as long as the claw; keel (12) 17-18 mm long, the limb semiorbicular, acutish, the claw 1.5 times as long as the limb; ovary sessile; pods sessile, erect, capitately crowded, oblong, very slightly curved, (10) 12-17 mm long, ca. 4mm broad and thick, abruptly contracted into an oblique subulate beak 1-2 (3) mm long, villous with rather short spreading soft white hairs, incompletely bilocular. Fl. (May) June; fr. July-August. ‘ Steppe slopes, pine or rarely broadleaved woods, occasionally margins of birch woods, scrub thickets, rocks, and clayey escarpments. — W. Siberia: Alt.; E. Siberia: Ang.-Say., Dau., Le.-Kol. (from north to Yakutsk and Vilyuisk districts and Kolyma river basin); Far East: Ze.-Bur. (between Khabarovsk and Blagoveshchensk on-Amur), Uss. (?), Okh. (Okhotsk). Endemic. Described from Lena River. Type in London. 682. A. iskanderi Lipsky in A.H.P. XXVI (1907—1909) 87; Fl. Tadzhik. V, 470. Frutescent, 35-90 cm tall; caudices 15-40 cm long, covered with castaneous~brown peeling bark, divided in upper part; branches of the preceding year 5-10 cm long, brownish-gray, covered with appressed white hairs; branches of the year (10) 15-30 cm long, densely covered with appressed white hairs, whitish; stipules triangular-lanceolate, 3-4 mm long, black-hairy, partly adnate at base to petiole; leaves 4-9 cm long, the petiole and rachis slender, covered with appressed white hairs; leaflets 3-5 pairs, linear-oblong to linear, rarely lance-linear, (14) 20-35 mm long, 2-4 (6) mm broad, subacute and slenderly mucronulate, covered on both sides with appressed white hairs; peduncles axillary and terminal, 8-15 cm long, 1.5-2 times as long as the leaves, firm, sulcate, covered with white and, beneath the inflorescence, white and few black hairs; racemes rather compact, ovaloid or broadly oblong, 5-7 cm long, elongating up to 10 cm and becoming loose and remotely fruited; bracts lanceolate, 5-6 mm long, villous with black and white hairs; calyx cylindric, becoming slightly inflated at maturity, finally ruptured by the pod, 17-20 mm long, shaggy- gray by spreading white and appressed black hairs, the subulate teeth 4-5 mm long, half the length of the tube; corolla violet-lilac; standard 23-27 mm long, the limb obovate, obtusish, 9-11 mm broad, gradually attenuate into and 3-4 times as long as the claw; wing~petals 22-26 mm long; the limb oval-oblong, round-tipped, the claw 1.6 times as long as the limb; keel 20-23 mm long, the limb semicircular, about half as long as the claw; ovary subsessile; pods sessile, obliquely ascending, ovoid or broadly oblong, 17-19 mm long, 5-6 mm broad, compressed laterally, slightly carinate ventrally, grooved dorsally, acute, terminating in a Subulate beak 2-3 mm long, densely villous with rather short spreading white hairs interspersed with shorter black spreading hairs, rigidly coriaceous, almost fully bilocular. Fl. May-July; fr. June-August. (Plate XLII, Figure 1). Mixed-grass steppes and juniper groves, in the intermediate mountain zone.— Centr. Asia: Pam.-Al. (Mtn. Zeravshan, northern slopes of Turkestan range). Endemic. Described from vicinity of Iskander~kulya in Zeravshan basin. Type in Leningrad. 683. A. ugamicus M. Pop. in Bull. Univ. As. Centr. XIV (1926) 142.— Exs.: H.F.A.M. No. 371. 529 709 Perennial, suffrutescent, up to 1 m tall; caudices up to 30 cm long, rather stout, they and their divisions covered with smooth castaneous- brown bark; stems of the year (2) 10-20 cm long, densely covered with appressed white hairs, grayish-white; stipules adnate to petiole to 3% 7-10 (13) mm long, herbaceous, covered with appressed white hairs, the free portion ovate or broadly oblong; leaves (4) 7-13 cm long, the petiole and the much longer rachis covered with scattered white hairs; leaflets (6) 8-11 pairs, oblong-elliptic to lanceolate, rarely oval, (7) 13-27 (30) mm long, acute, rarely round-tipped, mucronulate, covered on both sides with scattered appressed hairs; peduncles 1.5-2 times as long as or rarely equaling the leaves, firm, erect, covered with appressed white and black hairs; racemes rather compact, ovoid, many-flowered, (3) 5-7 cm long; bracts linear, 5-7 (8) mm long, shorter than to equaling the calyx-tube, black- or black-and-white ciliate; calyx cylindric, 10-13 mm long, black- pilose and white-villous with spreading hairs, the filiform-linear teeth half as long as the tube; corolla lilac; standard 19-24 mm long, the limb rhomboidally oblong-ovate, dilated at the middle, rather gradually and broadly ligulate-attenuate upward, obtusish, toward base angularly and ligulately short-attenuate, 3-4 times as long as the claw; wing-petals 15-19 mm long, the limb linear-oblong, rounded-obtuse, as long as the claw; keel 11-14 mm long, the limb semiorbicular, acutish, the claw 1.5 times as long as the limb; pods sessile, gathered in a compact ovoid head 5-12 cm long, obliquely ascending, oblong, 1.5 times the length of the calyx, 13-20 mm long, 3.5-4 mm Iroad, carinate ventrally, grooved dorsally, terminating in a slightly reflexed beak 5-6 mm long, coriaceous, villous with ascending fairly short white hairs, occasionally interspersed with few short black hairs, bilocular. Fl. (April) May-June; fr. (May) June-July. _Fine-grained or rarely rocky mountain slopes, in the loam belt of the intermediate mountain zone.— Centr. Asia: Tien Shan. Endemic. Described from W. Tien Shan (from Ugam). Type in Tashkent. 684. A. Komarovii Lipsky in A.H.P. XXVI, (1907—1909) 186 (excl. specim. ad A. dendroides Kar. et Kir. pertin. ). Perennial, suffrutescent, up to 1 m tall, the bark castaneous-brown; leaflets 6-9 pairs, oval to oblong-oval, more or less round-tipped and mucronulate, 12-30 mm long, glabrate above, covered beneath with scattered hairs, green; peduncles sparsely covered with appressed black hairs interspersed with few white ones; racemes compactly ovoid, ca. 5 mm long; bracts linear, 11-12 mm long,exceeding the calyx-tube and somewhat overtopped by the teeth; standard (24) 25-27 mm long, the limb irregularly ovate, rounded-dilated at the middle, abruptly ligulate-attenuate upward, slightly retuse, short-attenuate and angular toward base, 4-5 times as long as the claw; pods sessile, oblong to linear-oblong, 12-18 mm long, ca. 4mm broad, acute, terminating in a slightly reflexed beak 8-11 mm long, carinate ventrally, grooved dorsally, coriaceous, rather densely covered with short subappressed black hairs interspersed with isolated or scattered longer white hairs, bilocular. Fl. June-July; fr. July-August (September). Fine-grained mountain slopes in the juniper belt; not infrequently in rosaria, partially displacing juniper.— Centr. Asia: Pam.-Al. Endemic. Described from Shakhrisyabs. Type in Leningrad. 530 710 685, A. viridis Bge. Astrag. geront. I (1868) 135; II (1869) 231; Boiss. Fl. or. I, 479; Grossh. Fl. Kavk. II, 322.—A. nitens Boiss. et Buhse in Nouv. Mém. Soc. Nat. Mosc. XII (1860) 61, non Boiss. et Heldr. A low shrub, lignified to high up, 30-45 cm tall; caudices 8-15 cm long, covered with brown bark; stems of the year (6) 8-17 cm long, densely covered with appressed white hairs, silvery; stipules lanceolate, acuminate, 3-5 mm long, covered with appressed white or mixed white and black hairs; leaves 6-11 cm long, the petiole and the somewhat longer rachis covered with appressed white hairs; leaflets 5-8 pairs, lanceolate or linear-oblong to linear, 1-1.5 cm long, 1.5-4 (7) mm broad, covered above with scattered and beneath with sparse or fairly dense appressed hairs, grayish-green; peduncles subterminal, 2-3 times the length of the leaves, angled, firm, covered with appressed white hairs; inflorescence compactly capitate, ovaloid, 10-20-flowered, 4-6 cm long; bracts linear or lanceolate, 2-4 mm long, covered with black hairs; calyx ca. 15 mm long, densely villous with spreading hairs, sometimes considerably interspersed, especially on the ventral side, with black hairs, the filiform teeth 3-4 mm long; corolla pale lilac; standard 20-22 mm long, the limb oval, obtusish, scarcely retuse, 1.5 times as long as the claw; wing-petals 19-21 mm long, the limb rounded-spatulate, entire, somewhat longer than the claw; keel 15-18 mm long, the limb semicircular, acute, half the length of the claw; ovary sessile; pods sessile, erect, ovoid to broadly oblong, 15-20 mm long, 5-6 mm thick and broad, slightly arcuate, carinate ventrally, canaliculate dorsally, coriaceous, densely villous with spreading white hairs and rather sparsely tomentose with short ascending black hairs, terminating in a short recurved cuspidate beak 2.5-3 (4) mm long, bilocular. Fl. May-June; fr. July. Clayey and often stony mountain slopes.— Caucasus: S. Transc. (vicinity of Nakhichevan'). Gen. distr.: Iran (N. W. Iran), Arm.-Kurd; Turkish Armenia (region of Kars). Described from the vicinity of Nakhichevan' and N.W. Iran. Type in Leningrad. Series 12. MUGODSCHARICI Vass.— Shrubs; lower leaves trifoliate, the upper ones composed of (3) 4-5 leaflets; leaflets linear; stipules of the lower leaves connate; corolla violet; pods erect, subsessile, oblong, short, faintly hairy. — Mugodzhary. 686. A. mugodsharicus Bge. in Arb. Naturf. Ver. Riga I (1847) 236; Bge. Astrag. geront. I, 124; II, 216; Bge. Astrag. turk. 288. A small shrub, 20-40 cm tall; old ligneous branches 10-40 cm long, ascending or rarely erect, slender, covered with brown bark; stems of the year (2.5) 5-15 (20) cm long, veryslender, simple, covered with smooth sublustrous naked bark, erect; stipules 1-1.5 mm long, the lower triangular-ovate, hyaline, acute, slightly adnate to petiole, connate into a short hyaline sheath, slightly white-tomentose; leaves 3-5 cm long, the petiole 2-5 (10) mm long, slender, sparsely covered with appressed white hairs; leaflets of lower leaves in 3's linear-oblong obtuse, those of upper leaves (3) 4-5 narrowly linear to subfiliform finely pointed, (10) 20-50 mm long, 1-2 (3) mm broad, glabrous above, sparsely covered beneath with short appressed white hairs; peduncles equaling to slightly exceeding the leaves, slender, glabrous or covered with isolated appressed white hairs; racemes 2-4 cm long, rather loosely 5-12-flowered; bracts ovate, acuminate, 1-1.5 mm long, minutely tomentulose with white or mixed white 531 Galak 712 and black hairs, 2/3 as long as to equaling the pedicel; calyx cylindric, 8-11 mm long, covered with subappressed white and fewer black hairs, the lance-subulate teeth 1-2 mm long, the tube 5-6 times as long as the teeth; corolla violet; standard (16) 19-22 mm long, the limb oblong-obovate, slightly retuse, abruptly narrowed in lower 1/3, slightly angular at base, 4-5 times as long as the claw; wing-petals (14) 16-20 mm long, the limb linear-oblong, entire, as long as the claw; keel (13) 15-18 mm long, the limb almost semiorbicular, acutish, the claw (1.5) 2 times as long as the limb; ovary borne on a stipe ca. 1 mm long; pods subsessile (the stipe 0.5 mm long), obliquely ascending, linear-oblong, 9-12 mm long, ca. 3 mm broad and 2 mm thick, acute, terminating in an oblique subulate beak 1.5-2 mm long, carinate ventrally, canaliculate dorsally, slightly compressed laterally, coriaceous, rather sparsely covered with short appressed white hairs, semibilocular. Fl. May-June; fr. July-August. Steppes, mostly on bluffs, stony and gravelly slopes, and rocks. — W. Siberia: U. Tob. (Mugodzhary, Orsk district). Endemic. Described from Mugodzhary. Type in Leningrad. Series 13. GLAUCI Vass.— Subshrubs or herbaceous perennials; stems branched, commonly ascending or decumbent, rarely erect; leaflets large, elongate, from 7 to 25 mm long; racemes abbreviated, umbelliform to capitate; corolla varying in color; pods from 10 to 23 mm long and from 2.5-3 to 4mm broad, usually ascending to erect. 687. A. dealbatus Pall. Astr. (1800) 26 p.p.—A. glaucus M.B. FI. taur.-cauc. II (1808) 186; III, 448.—A. vesicarius ¢.—glaucus Beck. in Rchb. Ic. fl. Germ. XII (1909) 117 p.p. (quoad pl. e Tauria). A. vesicarius Bge. Astr. geront.I (1868) 135, II (1869) 231, non L., pups ie weal. Ast. (800)it. 23.910. 1 Perennial, suffrutescent, (20) 30-40 cm tall; caudex strong, woody, buried, 7-24 cm long, stout, branched in lower part, covered with brownish bark; stems of the year 4-10 cm long, sometimes abbreviated to 1.5-2 cm, whitish by a dense coat of appressed white hairs; stipules free, lanceolate, 3-5 mm long, covered with more or less appressed white hairs; leaves 3-6 cm long, the petiole 0.6-1.5 cm long, both petiole and rachis covered with appressed white hairs; leaflets 3-6 pairs, lanceolate or lance-elliptic to lance-oblong, 1-2.5 cm long, 3-6 mm broad, acute or rarely obtuse, covered on both sides with appressed white hairs; peduncles (1.5) 2 times the length of the leaves, 10-25 cm long, sulcate, covered with appressed white hairs; racemes capitate, 2-5 cm long, rather compactly 7-18- flowered, broadly ovaloid, often elongating in fruit up to 9 cm, oblong; bracts linear to lanceolate, 3-5 mm long, covered with white and black hairs; calyx cylindric, ca. 12 mmlong, densely covered with stiffish spreading white hairs interspersed, especially on the teeth, with very few black hairs, the teeth filiform, 3-4 mm long, the tube 2-3 times as long as the teeth; corolla whitish; standard (20) 22-24 mm long, the limb oblong- obovate, broadly short-attenuate toward apex, obtuse, scarcely broadly emarginate, auriculate-angular at base, 4 times as long as the claw; wing- petals 19-20 mm long, the limb linear-oblong, entire, the claw 1.5 times as long as the limb; keel 16-17 mm long, the limb broadly semicircular, acutish, half as long as the claw; ovary almost fully sessile; pods sessile, erect, oblong, 10-15 cm long, 3-4 mm broad, prolonged into an oblique 532 subulate beak 1-2 mm long, carinate ventrally, shallowly grooved dorsally, rigidly coriaceous, shaggy with rather short ascending white hairs, bilocular. Fl. May-June (July); fr. July-August. Steppelike and stony mountain slopes. — European part: Crimea. Endemic. Described from Crimea. Type in Leningrad. 688. A. Zingeri Korsh. in A.H.P. XI (1890) 297; Schmalh. Fl. I, 235. Frutescent, (13) 30-55 cm tall; caudex buried, strong; aerial caudices wanting or branched and 5-10 (15) cm long, covered with castaneous-brown bark; stems of the year (5) 10-25 cm long, slender, white by a dense coat of short appressed hairs, erect or rarely ascending; stipules free, triangular-ovate or lanceolate, ca. 2 mm long, covered with appressed white hairs; leaves (3.5) 5-7 cm long, the very short petiole and the rachis slender, covered with appressed white hairs; leaflets 4-6 pairs, linear- lanceolate, acute, rarely oblong or linear sometimes subobtuse, very rarely oval-oblong, 0.7-1.5 (2.5) cm long, 1-4 (10) mm broad, covered on both sides with scattered appressed white hairs or glabrous above; peduncles 1.5 times the length of the leaves, (5) 10-20 cm long, covered with subappressed white hairs; racemes capitate, closely flowered, 2-5 cm long, becoming oblong in fruit and elongating up to 10 cm; bracts lance-linear, ca. 2mm long, covered with white and sparse short black hairs; calyx cylindric, 9-11 mm long, densely subpannose with spreading white hairs very sparsely interspersed, especially on the teeth, with black hairs, the tube 4 times as long as the linear-subulate teeth; corolla white, rarely with a violet standard; standard (17) 20-23 mm long, the limb oblong- obovate, scarcely retuse, gently constricted above the lower 1/3, angularly and auriculately enlarged below, 4 times as long as the claw; wing-petals (15) 17-21 mm long, the limb oval-oblong or oblong-oval, rounded-obtuse, the claw slightly longer than to 1.5 times as long as the limb; keel (13) 15-18 mm long, the limb semiorbicular, obtusish, the claw 1.5-2 times as long as the limb; ovary sessile; pods sessile, erect, linear-oblong, (10) 15-20 mm long, ca. 3-4 mm broad, compressed laterally, rounded- carinate ventrally, scarcely canaliculate dorsally with a slightly concave suture, covered with spreading silky white hairs or villous with white and few black hairs, terminating in a short oblique subulate beak 2~3 mm long, bilocular, Fl. May-June (July); fr. June-August. Over chalk and limestone (pinewoods overlying chalk, stony calcareous steppe slopes, on chalky outcrops of ravine slopes, less frequently in sandy soil. — European part: V.-Don (right bank of the Volga, from Ul'yanovsk to Saratov, Zhiguli), L. Don (right bank of the Volga from Saratov to Kamyshin and Balashov), Transv. (Kuibyshev District, Kineshma). Endemic. Described from Kuibyshev territory (from ''Tsarev Kurgan"’). Type in Leningrad. 689. A. pycnolobus Bge. in Mém. Sav. Etr. Pétersb. II (1835) 594; Ldb. Fl. Ross. I, 630; Bge. Astrag. geront. I, 127; II, 224; Bge. Astrag. fone pests wryle Blo Zap, Sib. Vil, 1702— A.\ceratoides var? db’ Fl.; Alt. ITI, (1831) 306. Perennial, (15) 18-35 cm tall; caudex buried, with numerous spreading divisions; stems 5-20 cm long, ascending or erect, sparsely covered with appressed white hairs; stipules connate in lower part, lanceolate, long-acuminate, 4-5 mm long, covered with appressed white or sometimes mixed white and black hairs; leaves 5-10 (15) cm long, the rachis 2-4 times * 533 714 as long as the petiole, both petiole and rachis sparsely appressed-hairy; leaflets 6-9 pairs, oblong or lanceolate, (7) 10-25 mm long, (2) 3-6 (9)mm broad, acute, sparsely covered on both sides with appressed hairs; peduncles equaling to 1.5-2 times as long as the leaves, covered with appressed white, and, beneath the inflorescence, mixed white and black hairs; racemes short, compactly 7-15-flowered, 3-3.5 cm long, elongating in fruit up to 4-5 cm, covered with white and black hairs, equaling to twice as long as the pedicel; calyx cylindric, 9-11 mm long, densely covered with appressed or subappressed white and black hairs, the tube 4-5 times as long as the subulate teeth; corolla whitish; standard 20-23 mm long, the limb obovate, retuse, rather gradually attenuate in lower 1/3, scarcely angular at base, 4-5 times as long as the claw; wing-petals 18-21 mm long, the limb oblong, scarcely laterally emarginate, as long as the claw; keel 14-17 mm long, the limb almost semiorbicular, slightly concave beneath, obtuse, the claw 1.5 times as long as the limb; ovary on astipe ca. 1 mm long; pods borne on a stipe 1-2 mm long, obliquely ascending or horizontally spreading or subcernuous, linear-oblong, straight, 15-23 mm long, 4mm broad, rather strongly compressed laterally, acutish, terminating in a subulate beak 1-3 mm long, carinate ventrally, shallowly and narrowly canaliculate dorsally, densely covered with appressed white and black hairs, coriaceous, bilocular. Fl. June-July; fr. July-August. Deciduous forests, subalpine meadows, less frequently steppe-mountain slopes. — W. Siberia: Alt. (S. Altai: at Bukhtarma River, Naryn range, in region of Ust'-Kamenogorsk and others). Endemic. Described from vicinity of Zyryanovskii mine at Kurchum River. Type in Leningrad. 690. A. Ionae Palib. in Addenda XI, p. 667. Perennial, acaulescent or nearly so, ligneous, 10-14 cm tall; caudex stout, buried, short-branched, or sometimes hardly distinct aerial blackish- brown; stems short, not more than 5-6, commonly 0.5-1.5 cm long, covered with appressed white hairs; stipules 3-5 mm long, connate into a sheath, the lower to high up, the upper merely at base, covered with white and black hairs, bidentate, the teeth triangular-ovate to lanceolate, acute; leaves 6-10 cm long, the petiole 2-3 (4) cm long, both petiole and rachis covered with appressed white hairs; leaflets 5-7 pairs, lance-linear, acute, 7-15 mm long, 1-2 mm broad, hoary on both sides by appressed hairs; peduncles 2/3 as long as to equaling or rarely 1.5 times as long as the leaves, slender, sparsely covered with white and, beneath the inflorescence, white and black hairs; racemes umbellate-capitate, 2.5-3 cm long, loosely 2-5-flowered; bracts lanceolate, 1.5 mm long, white- or white-and-black- ciliate; calyx cylindric, 8-10 mm long, covered with black and fewer white hairs, the triangular-subulate teeth 2-2.5 mm long; corolla whitish with violet-tipped keel and barely violet-tinged standard; standard 20-25 mm long, the limb obovate-oblong, retuse, rather gradually attenuate in lower 1/4, angular at base, 4 times as long as the claw; wing-petals 17-21 mm long, the limb linear-oblong, rather shallowly and broadly emarginate, as long as the claw; keel 14-15 mm long, the limb broadly retuse, as long as the claw; keel 14-15 mm long, the limb broadly obovate, obtusish, the claw 1.5times as long asthe limb [sic]; ovary onastipeca.1.5mmlong; pods sessile, erect, oblong, 10-20 mm long, 2.5-3 mm broad and thick, straight, acute, terminating in a subulate beak ca. 1 mm long, rounded~carinate ventrally, broadly grooved dorsally, coriaceous, densely villous with 534 715 spreading white hairs interspersed with shorter appressed black hairs, bilocular. Fl. May-June; fr. July. Mountain slopes and bluffs.— E. Siberia: Ang.-Say. (Achinsk district). Endemic. Described from vicinity of Lake Bozh'e in Karetak Mountains and Kolba River. Type in Leningrad. 691. A. pskemensis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent; caudex stout, buried, prolonged into a very short aerial caudex, the divisions ascending or subdecumbent, covered with castaneous-brown bark, 3-5 (10) cm long; stems of the year slender, 9 cm long, covered with white appressed hairs; stipules ovate-triangular, 1-2 mm long, covered with white hairs; leaves 5-8 cm long, the petiole 1-3 cm long, the rachis 2-3 times as long, both petiole and rachis slender, covered with white appressed hairs, hoary; leaflets 5-8 pairs, linear or rarely linear-oblong, 7-12 mm long, 0.7-2.5 mm broad, obtuse or slightly retuse, covered on both sides with appressed white hairs, hoary; peduncles equaling or slightly exceeding the leaves, 5-10 cm long, slender, covered with stiff appressed white hairs; racemes compactly capitate, (3) 5-10-flowered, 2-2.5 cm long; bracts linear, ca. 2 mm long, black-hairy; calyx cylindric, 12 mm long, hirsute with short ascending black hairs and villous with longer white hairs, the teeth filiform, 3-4 mm long, covered with black hairs; corolla whitish; standard 17-18 mm long, the limb oblong-oval, retuse, twice as long as the claw; wing-petals 15-16 mm long, the limb linear-oblong, entire, the claw 1.5 times as long as the limb; keel 13-14 mm long, the limb semiorbicular, obtusish, about half as long as the claw; ovary on stipe 1 mm long; pods sessile, compactly and capitately crowded, oblong, acute, 12-20 mm long, 3-4 mm broad and thick, terminating in an oblique beak 1.5-3 mm iong, carinate ventrally, broadly and rather shallowly canaliculate dorsally, triquetrous, coriaceous, covered with subappressed white andblack hairs, either color predominating, bilocular. Fl. July; fr. August (September). Gravelly mountain slopes.— Centr. Asia: Tien Shan (W., Pskem River basin). Endemic. Described from upper reaches of Ispai gorge. Type in Leningrad. Series 14. BADAMENSES Vass.— Subshrubs, with short decumbent or ascending stems; leaflets broadly oval to oblong-obovate, very small, from 2-3 to 4-7 mm long; racemes abbreviated, umbelliform, depauperate, 1-3 (4-5)-flowered; corolla varying in color; pods 10-20 mm long and (2.5) 3-4 mm broad. Mountains of Central Asia (W. Tien Shan) and the Caucasus (Dag. ). 692. A. badamensis M. Pop. in Bull. Univ. As. Centr. fasc. 14 (1926) 143. Perennial, suffrutescent, loosely cespitose, 9-12 cm tall, the aerial ligneous divisions of the caudex prostrate, up to 3-4 cm long; stems of the year slender, up to 2-10 cm long, prostrate or ascending, hoary by appressed white hairs; stipules triangular, obtusish, 0.5-1 mm long, covered with appressed white hairs; leaves 1-2 cm long, the rachis as long as to 2-4 times as long as the petiole, both petiole and rachis covered with appressed white hairs; leaflets 4-7 pairs, oblong, 2-5 mm long, subobtuse, covered on both sides with appressed white hairs, hoary; peduncles equaling or more often 1.5-2 times the length of the leaves, 535 716 (2)3.5-4cm long, densely hoaryby appressed white hairs, these interspersed beneath the inflorescence with isolated short black hairs; racemes umbelliform, 2-3-flowered; bracts triangular-ovate, acutish, 0.5-1 mm long, covered with short white and black hairs; calyx narrowly cylindric, ca. 10 mm long, covered with appressed white and black hairs, the subulate teeth ca. 1 mm long; corolla white with a barely violet standard; standard ca. 18 mm long, the limb oblong-oval, short-attenuate toward apex, slightly retuse, scarcely narrowed in lower 1/3, twice as long as the claw; wing- petals ca. 15 mm long, the limb oblong, entire, round-tipped, the claw 1.5 times as long as the limb; keel ca. 13 mm long, the limb almost semicircular, obtuse, the claw 1.7 times as long as the limb; ovary ona stipe ca. 1.5 mm long; pods almost fully sessile, subhorizontally spreading, subulate-linear, straight, 15-20 mm long, ca. 2.5-3 mm broad, abruptly contracted into a subulate straight beak ca. 1 mm long, carinate ventrally, grooved dorsally, covered with appressed white and black hairs, coriaceous, bilocular. Fl. and fr. August. Centr. Asia: Tien Shan (W.: Ak-bashtau Mountains). Endemic. Described from vicinity of Dorofeevskaya village. Type in Tashkent. 693. A. Baranovii M. Pop. inIzv. Turk. otd. Russk. geogr. o-va XVII (1925) 30.— Exs.: H.F.A.M., fasc. XVI, No. 378. Perennial, dwarf rupestral plants, 4-8 cm tall, from a long-branched buried caudex, nearly acaulescent or producing stems up to 2 cm long covered with appressed white hairs; stipules adnate to petiole and connate to high up, hyaline, sparsely covered with short hairs, ca. 1.5 mm long, the free portion triangular; leaves 2.5-6 cm long, the petiole as long as the rachis, slender, sparsely covered with appressed white hairs; leaflets (3) 4-5 pairs, oval to rounded-oval, (3) 4-6 mm long, rounded-obtuse, hoary on both sides with appressed hairs; peduncles half as long as to nearly as long as the leaves, slender, covered with appressed white and, beneath the inflorescence, mixed white and black hairs; racemes abbreviated, capitately umbellate, loosely 4-5-flowered, 1.5-2 cm long; bracts ovate-lanceolate, ca. 4.5 mm long, covered with short black hairs; calyx cylindric, 8-10 mm long, villous with short ascending black and mostly few white hairs, the tube 2-3 times as long as the linear-subulate teeth; corolla pale purple (?); pods borne on a stipe ca. 1 mm long, lance- oblong, 10-11 mm long, 3 mm broad, short-acuminate at both ends, terminating in a slightly curved subulate beak 2 mm long, carinate ventrally grooved dorsally, villous with ascending white hairs, bilocular. Fl. July; fr. August. Rock crevices in the subalpine zone.— Centr. Asia: Tien Shan (W. : Bol'shoi Chimgan Mt.). Endemic. Described from Bol'shoi Chimgan Mt. Type in Tashkent. 694. A. fissuralis Alexeenko in Lipsky Fl. cauc. Suppl. I in Trav. Jard. Bot. Tiflis VI, fasc. I (1901) 49.— Exs.: Pl. or. exsic. No. 116. Perennial, 3-7 cm tall; caudices buried, rather slender, branched; stems of the year 1-3 (5) cm long, slender, white by short appressed hairs; stipules connate in lower part, ovate-triangular, 1-1.5 mm long, the upper oblong-ovate, covered with black hairs; leaves 1.5-3 cm long, the petiole 4 cm long, both petiole and rachis densely covered with appressed white hairs; leaflets 2-4 pairs, oval or oblong-obovate, mostly obtuse, 3-5 (7) mm long, 2-4 mm broad, silvery-sericeous on both sides; peduncles i pe Vee 3 ¢ 21 long, slender, covered with appressed white and, in upper part, black and 536 17 718 white hairs, cinereous; flowers 1-3, approximate, in an umbelliform inflorescence; bracts oblong-lanceolate, 1-2 mm long, black-hairy; calyx cylindric, 10-12 mm long, villous with black hairs interspersed on the back with white ones, the linear teeth 2-3 mm long; corolla violet; standard 21-22 mm long, the limb broadly subrhomboid-oval, broadly short-attenuate toward apex, retuse, 1.5 times as long as the claw; wing- petals 18 mm long, the limb oblong, inequilaterally emarginate at apex, as long as the claw; keel 16 mm long, the limb obtusish, subtriangular, half as long as the claw; ovary on a stipe ca. 1 mm long; pods sessile, linear, obliquely ascending, straight or slightly arcuate, 15-25 mm long, 3 mm broad, somewhat compressed laterally, carinate ventrally, rounded dorsally, acute, rather sparsely covered with appressed black and ascending longer white hairs, bilocular. Fl. and fr. June-July. Calcareous rocks and stone mounds, at altitudes of 1100-1800 m. — Caucasus- Dag. Endemic. Described from a locality between Maara and Akusha. Type in Leningrad. Series 15. FARCTI Vass.— Herbaceous perennials with developed erect stems; leaflets linear, 15-30 mm long; racemes elongated, loose; corolla white, 24-27 mm long; pods erect, 15-20 mm long and 3.5-4 mm broad, enclosed in a double-layered wall, the two layers separated by a spongy tissue. — Mountains in the south of Tadzhikistan. 695. A. satteotoichus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, 35-50 cm tall; stems 20-30 cm long, erect, slightly flexuous, simple, angled-sulcate, thinly covered with short appressed white hairs; stipules free, scarcely adnate at base to petiole, triangular-lanceolate, subulately long-acuminate, 6-7 mm long, herbaceous, covered with appressed white and black hairs; leaves 5-11 cm long, the rachis 2-4 times as long as the petiole, both petiole and rachis covered with appressed white hairs; leaflets (5) 7-10 pairs, linear, very rarely lance-linear, (1.5) 2-3 cm long, acute, sparsely covered on both sides with appressed hairs; peduncles 2/3 the length of to equaling the leaves, 7-12 cm long, angular, covered with appressed white and, in upper part, mixed white and black hairs; racemes elongated, loosely many-flowered, equaling to 1.5 times as long as the peduncle; bracts lance-linear, often subulately long- acuminate, 3-4 mm long, 3 times the length of the pedicel, covered with white and black hairs; calyx narrowly cylindric, 15-16 mm long, 2.5 mm broad, covered with appressed black and fewer white hairs, the tube 3-4 times as long as the linear-subulate teeth; corolla white; standard 24-27 mm long, the limb obovate, entire, short-attenuate and angular toward base, twice as long as the claw; wing-petals 22-24 mm long, the limb linear-oblong, entire, obtuse, about half the length of the claw; keel 20-21 mm long, the limb slightly gibbous, obtuse, half as long as the claw; ovary sessile; pods sessile, erect or obliquely ascending, oblong to lance- oblong, straight or arcuate, strongly inflated, (13) 15-20 mm long, 3.5-4 mm broad, ca. 5 mm thick, rounded or scarcely carinate ventrally, broadly grooved dorsally, acutish, terminating in a subulate beak 2-3 mm long, densely covered with appressed white hairs often interspersed on the ventral side with isolated black hairs, bilocular, the lanceolate valves splitting transversely in maturity, the two layers of the wall separated by a spongy tissue. Fl. April-May; fr. May-June. 537 71.9 Slopes of loess foothills in the ephemeral vegetation and pistachio belts. — Centr. Asia: Pam.-Al. (S.: in low mountains of Southern Tadshikistan from Kulyab district to Mt. Aruktau, right bank of river Vakhsh). Endemic. Described from Mt. Kichik-teryai eastward from Kulyab. Type in Leningrad. Series 16. PSEUDOHELMIA M. Pop. Stipules connate into a sheath; plants almost suffrutescent, ligneous at base; racemes loose, long- peduncled; corolla pink; pods sessile or more often short-stipitate, linear- oblong, deeply grooved on the back, compressed laterally, covered with appressed white hairs or glabrate. 696. A. chorgossicus Lipsky in A.H.P. XXVI (1907-1909) 257. Perennial, slightly ligneous ‘at base, ca. 20 cm tall; caudex prostrated, buried and partially aerial, branched; stems of the year strongly abbreviated, 4-5 cm long, densely covered with appressed white hairs, ascending; stipules connate to 1/2-2/3 into a sheath, 2-3 mm long, densely covered with appressed white hairs, the free portion triangular; leaves 2-4.5 cm long, the rachis equaling to 1.5-2 times as long as the petiole, the latter densely covered with appressed white hairs; leaflets 3-4 pairs, oval-triangular or lanceolate, obtuse, 5-14 mm long, 2-3 mm broad, covered on both sides with appressed hairs, hoary; peduncles 2.5-3 times the length of the leaves, 10-15 cm long, covered with appressed white hairs; racemes loose, 4-8 cm long, remotely 5-10-flowered; bracts ovate, 1.5-2 mm long, equaling the pedicel, covered with short black and white hairs; calyx cylindric, 11-12 mm long, covered with short appressed white and black hairs, the lance-linear teeth 2 mm long; corolla pink, persistent in fruit; standard ca. 25 mm long, the limb oblong-obovate, retuse, narrowed in lower 1/3, angular at base, 4 times as long as the claw; wing- petals 23 mm long, the limb obovate-oblong, scarcely retuse, somewhat shorter than the claw; keel ca. 16 mm long, obtusish; pods borne on a stipe ca. 3 mm long, linear-oblong, 13-14 mm long, ca. 3 mm broad, acute, terminating in a straight subulate beak ca. 1.5 mm long, carinate ventrally, deeply grooved dorsally, coriaceous, glabrous or covered with isolated appressed hairs, bilocular, more or less spreading. Fl. andfr. May. Stony mountain slopes.— Centr. Asia: Dzu. Tarb. (Dzhungarian Ala Tau). Endemic. Described from Khorgos river valley, near Bashkunchi. Known solely from this location. .Type in Leningrad. 697. A. kendyrlyki M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). : Perennial, almost suffrutescent, ligneous at base, 12-18 cm tall; caudex branched, buried and partially aerial; stems of the year slender, abbreviated, 1-3 cm long, ascending to erect, covered with appressed white hairs, hoary; stipules connate to high up into a sheath, 1-1.5 mm long, scarcely covered with short white hairs, the free portion triangular, acute; leaves 1-3 cm long, the petiole 0.5-1.5 cm long, slender, hoary by appressed white hairs; leaflets 3-5 pairs, small, linear to oblong-linear, subacute to subobtuse, 3-6 mm long, 0.5-1.5 mm broad, covered on both sides with appressed hairs, hoary; peduncles 4-15 cm long, covered with appressed white hairs, terete, slender; racemes 1.5-4 cm long, loosely 5-10-flowered, 3-7 cm long [sic], the flowers distinct; bracts ovate- lanceolate, 1-1.5 mm long, membranous, sparsely covered with black hairs; flowers nodding (?); calyx cylindric, 8 (10) mm long, sparsely 538 720 covered with appressed white and black hairs, the teeth ca. 0.5 mm long, black-hairy; corolla pale pink; standard 16 mm long, the limb oblong- obovate, narrowed in lower 1/3, angular at base, retuse in upper 2/3 [? |; wing-petals 11 mm long, the limb linear-oblong, entire, 1.5 times as long as the claw; pods cernuous or horizontally spreading, on a stipe ca. 2mm long, linear-oblong, straight, 6-10 mm long, 2-3 mm broad, carinate ventrally, deeply grooved dorsally, short-cuspidate, very sparsely covered with stiff appressed white hairs. Fl. May; fr. June. Mountain slopes with southern exposure.— Centr. Asia: Dzu.-Tarb. (Saur). Endemic. Described from Sary-bulak river valley eastward from Kendyrlyk. Known solely from this location. Type in Leningrad. Series 17. SOGDIANI Vass.— Herpaceous perennials or subshrubs with elongated leaflets; racemes loose, elongated, remotelyflowered; corolla white and violet; pods oblong or lanceolate, from 10 to 20 (27) mm long, 2.5-3 mm broad, reflexed.— Mountain steppes and sands in river valleys of Central Asia. 698. A. sogdianus Bge. Astrag. turk. (1880) 99. Perennial, 14-35 cm tall; stems 3-9 cm long, erect, sparsely covered with appressed white and black hairs; stipules lanceolate to linear- lanceolate, 3-6mmlong, long-acuminate, herbaceous, covered with black and white hairs; leaves (5) 8-10 (12) cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis rather sparsely covered with appressed white or mixed white and black hairs; leaflets 4-6 pairs, narrowly linear or rarely lance-linear, 10-18 mm long, 1-2 (3) mm broad, rather sparsely covered on both sides with appressed white hairs; peduncles somewhat shorter than to somewhat exceeding the leaves, 6-9 (13) cm long; racemes loose, 4-8 cm long, remotely 8-15-flowered; bracts lance-linear, 2-4 mm long; calyx cylindric, 9-11 mm long, covered with appressed white and black hairs, the tube 2-2.5 times as long as the linear-subulate teeth; corolla pinkish-violet; standard 15-19 mm long, the limb oblong-ovate, narrowed in lower 1/4, ovate upward, broadly and ligulately short-attenuate toward apex, obtusish, angular at base, 4-5 times as long as the claw; wing-petals 14-17 mm long, the limb linear- oblong, bidentate, as long as to somewhat longer than the claw; keel 11-13 mm long, the limb semicircular, the claw 1.5-2 times as long as the limb; pods borne on a very short stipe, linear-oblong, 13-18 mm long, slightly arcuate, scarcely carinate or almost rounded ventrally, slightly grooved or almost rounded dorsally, coriaceous, terminating in a slender straight beak 3-4 mm long, sparsely covered with appressed white and black hairs, bilocular. Fl. (April) May; fr. May-June. Fine-grained slopes in the witchgrass steppe belt and in the lower part of the juniper belt, in the north on rolling foothills.— Centr. Asia: Syr D., Pam.-Al. Endemic. Described from Dzhizman canyon. Type in Leningrad. 699. A. Angreni Lipsky in A,H.P. XXVI (1907-1909) 166; Fedtsch. Rast. Turk. (1915) 529.— Exs.: Herb. Fl. As. Med. No. 352. Herbaceous perennials, slightly ligneous at base, (12) 20-50 (55) cm tall, the ligneous branches 1-4 cm long; stems of the year simple, 5-15 (33) cm long, slightly and softly sulcate, whitish by a dense coat of short appressed hairs; stipules adnate to petiole in lower 1/3-1/2, triangular-lanceolate, acute, 3-4 cm long, covered with appressed white hairs; leaves (2)"3".5=6\'em 539 PLATE XLIII 1, Astragalus Fedtschenkoanus Lipsky.—2, A, Variegatus Franch,—3, A.scleroxylon Bge. | 723 724 long, the rachis 2-4 times as long as the petiole, both petiole and rachis slender, sparsely covered with appressed white hairs; leaflets 6-7 pairs lance-linear, (5) 8-20 (25) mm long, 1-3 (4) mm broad, acuminate aera on both Sides with scattered appressed white hairs, green; peduncles 2-3 times the length of the leaves, slender, covered with short appressed white hairs; racemes elongated, (7) 9-18 cm long, very loose, 6-12- flowered, the flowers and fruits very distant, the rachis (5) 8-17 cm long elongating in fruit up to 19 cm; bracts triangular-lanceolate, acute 1.5-2.5 mm long, covered with short white hairs; flowers strongly By length horizontally spreading; pedicels 1.5-2 mm long; calyx cylindric,. 19-20 mm long, covered with appressed white and fewer black hairs, the filiform- linear teeth ca.5 mm long; corolla white; standard 22-24 mm long, the limb irregularly obovate, retuse, abruptly narrowed in lower 1/2-1/3 slightly angular at base, twice as long as the claw; wing-petals 21-22 mm long, the limb oblong, round-tipped, half as long as the claw; keel20-21 mm long, the limb inequilaterally oblong-oval, obtuse, half the length of the claw; ovaryborne ona stipe ca. 1 mm long; pods sessile, horizontally spreading to subcernuous, inequilaterally lance-linear, carinate ventrally, slightly canaliculate dorsally, (10) 12-17 mm long, acute, subterete, coriaceous, densely covered with short appressed white hairs, whitish, bilocular, the slightly curved beak (3) 4-5 mm long. Fl. May; fr. June -July. Fine-grained slopes of foothills and low mountains, from the upper part of the ephemeral vegetation zone to the witchgrass steppe and Inula grandis tall grass association; in the south (Turkestan Range) in juniper groves.— Centr. Asia: Pam. Al., Syr D., T. Sh. (W.). Endemic. Described from Angren. Type in Leningrad. 700. A. karataviensis Pavl. in Bull. Soc. Nat. Moscou XLIV, 1-2 (1935) 37. Perennial, suffrutescent, strongly branched, up to 30-40 cm tall; cauline shoots many, 2-6 cm long, stout, slightly flexuous, ligneous, covered with brownish bark; stems of the year 20-25 cm long, slender, erect or assurgent, densely covered with appressed white hairs; stipules 1-2 mm long, linear lanceolate, free, white-hairy; leaves 2-3 cm long; leaflets 4-6 pairs, 5-9 mm long, 1-2 mm broad, linear-lanceolate, covered on both sides with appressed white hairs; racemes elongated, loosely 4-10-flowered; peduncles 3-5 times as long as the leaves; bracts 2-2.5 mm long, both bracts and pedicels covered with appressed white and black hairs; calyx cylindric, 6-10 mm long, covered with white appressed hairs, the linear teeth 1-2 mm long, covered with white and black hairs, the tube 4-5 times as long as the teeth; corolla pale purplish-violet, 15-18 mm long; standard 15-18 mm long, 4-5 mm broad, oval, round-tipped and slightly retuse; wing-petals elongate-oval, round-tipped; keel 10-12 mm long; ovary sessile, long-beaked, covered with appressed white hairs; pods... Fl. May. Clayey foothill steppes.— Centr. Asia: Tien Shan. (Karatau). Endemic. Described from Mt. Karatau (northern Atabaevo). Type in Moscow. 701. A. marguzaricus Lipsky in A.H.P. XXVI (1907-1909) 164. — A. seravschanicus Lipsky in A.H.P. XXVI (1907) 169, non Freyn (1904); Fl. Tadzh. V (1937) 437.— Exs.: H.F.A.M. No. 375. Perennial, 25-60 cm tall; stems 5-30 cm long, branched, ligneous at base, densely covered upward with appressed whitish hairs; stipules lance- linear, 2-4 mm long, covered with white hairs; leaves 5-11 cm long; 541 725 leaflets (1) 2-3 (4) pairs, linear to oblong or oval to obovate, (8-10) 15-25 (50) mm long, (1.5) 2-3 (5-8) mm broad, acute, pannose on both sides with appressed white hairs, canescent or greenish; racemes loose, 10-30- flowered; peduncles 12-22 cm long; bracts linear-lanceolate, 2-3 mm long, white-hairy; calyx cylindric, (12) 16-20 mm long, densely covered with appressed white hairs sometimes interspersed with black ones, the tube 2-3 times as long as the linear-subulate teeth; corollapale purple; standard (17) 20-26 mm long, oblong-obovate, obtuse or retuse; wing-petals (16) 18-22 mm long, oblong to linear-oblong; keel (14) 16-20 mm long; pods cernuous, narrowly oblong, 15-20 mm long and 2-3 mm broad, gradualiy attenuate into a beak ca. 2 mm long, applanate dorsally, faintly carinate ventrally, subtriangular in cross-section, covered with appressed white hairs, on a stipe 1-3 mm long, incompletely bilocular; seeds black- punctulate, brownish, slightly notched, 4 mm long, 2 mm broad. Fl. May- July; fr. June-August. Intermediate mountain zone, on stony and gravelly slopes and on pebbles. Centr. Asia: Syr D., Pam. Al. Endemic. Described from Lake Marguzar. Type in Leningrad. 702. A. baldschuanicus M. Pop. in Not. Syst. Herb. Hort. Petrop. IV (1923) 155; Fl. Tadzhik. V (1937) 437. Perennial, 20-45 cm tall; stems 5-20 cm long, branched, ligneous at base, rather densely covered with appressed white hairs; leaves 5-11 cm long; leaflets 1-2 (4) pairs, linear to lanceolate, acute, 10-50 mm long, covered on both sides with appressed white hairs or greenish; racemes 10-25 cm long, loosely 15-30-flowered; peduncles 10-25 cm long; bracts 1.5-2 mm long, lance-linear, white-hairy; calyx cylindric, 12-16 mm long, covered with appressed white hairs, the tube 1.5-2 times as long as the linear- subulate teeth; corolla pale purple; standard 15-17 (18) mm long, oblong- obovate; wing petals 15-17mm_ long, oblong; keel ca. 14 mm long; pods cernuous, narrowly oblong, 13-20 mm long and 2-3 mm broad, gradually attenuate into a beak 2-3 mm long, faintly carinate ventrally, applanate dorsally, subtriangular in cross-section, ona stipe 1-3 mm long, covered with appressed white hairs, nearly bilocular; seeds brownish, ca. 3 mm long and 1 mm broad. Fl. May-August. fr. June-August. Foothills and low mountains, on dry mostly clayey slopes, ascending into the intermediate mountain zone (up to 2200 m in the rosarium formation).— Centr. Asia: Pam.- Al. (S. Tadzhikistan and Uzbekistan). Endemic. Described from Tadzhikistan (Bal'dzhuan). Type in Leningrad. 703. A. Kessleri Trautv. in Bull. Soc. Nat. Mosc. XXXIII, I (1860) 496; Bge. Astrag. geront. I, 125; II, 219; Bge. Astrag. turk. 284. Perennial, nearly acaulescent, 20-27 cm tall; caudex short, buried, branched; aerial stems strongly abbreviated, densely clothed in remnants of old petioles and stipules, no more than 1 cm long; stipules adnate to petiole in lower part, triangular-oblong, 3-4 mm long, covered with ascending white hairs; leaves 4-5 cm long, the petiole equaling to 1.5 times as long as the rachis, both petiole and rachis sparsely covered with appressed white hairs, indurescent and partly persistent; leaflets 5-6 pairs, linear to lance-linear, acutish, 8-15 mm long, 1-2 mm broad, rather densely covered with appressed white hairs; peduncles 2-3 times as long as the leaves, firm, virgate, sparsely covered with appressed white hairs; racemes 5-8 (10) cm long, remotely flowered; bracts oblong-ovate, 942 1.5-2 mm long, covered with isolated very short white or mixed white and black hairs; calyx cylindric, 7-8 mm long, covered with very short appressed white and black hairs, the triangular-subulate teeth ca. 1 mm long; standard 20 mm long, the limb oblong-obovate, slightly retuse, slightly narrowed in lower 1/4, angular at base, 4 times as long as the claw; wing~petals 19 mm long, the limb linear, entire, rounded-obtuse, as long as the claw; keel 16 mm long, the limb semioval, slightly short-attenuate toward apex, obtuse, as long as the claw; ovary on a stipe ca. 1 mm long; pods borne on a stipe ca. 1 mm long, cernuous, inequilaterally oblong, 14-20 mm long, 2.5-3 mm broad, carinate ventrally, canaliculate dorsally, acute and terminating in a straight subulate beakca.1 mm long, coriaceous, glabrous or covered with isolated appressed white hairs, finely cross- nerved. Fl. May; fr. June. Mountain slopes.— Centr. Asia: Tien Shan. Gen. distr. : Dzu.- Kash. (Kulja). Described from Tien Shan (Mt. Bektaktu). Type in Leningrad. 704. A. pendulinus M. Pop. et B. Fedtsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 16-30 cm tall; caudices profusely leafy in lower part, together with branches 12-20 cm long, covered with castaneous- brown bark; stems of the year 5-10 cm long, slender, white by a dense coat of appressed white hairs; stipules connate and adnate to petiole to 1/3-1/2, 3-3.5 mm long, covered with white and black hairs, the upper ones triangular-lanceolate in their free part, acute, the lower ovate-triangular; leaves 1.5-3.5 cm long, the petiole slender, longer than the rachis, sparsely covered with appressed white hairs; leaflets 4-5 pairs, linear or rarely linear-oblong, those of lower leaves often oblong-obovate, 8-10 mm long, glabrous above, covered beneath with scattered appressed hairs; peduncles 1.5-2 times as long as the leaves, slender, covered with isolated appressed white and upward black hairs; racemes loose, subremotely few-flowered; bracts lanceolate, 2-2.5 mm long, covered with black and fewer white hairs; calyx cylindric, 8 mm long, villosulous with short ascending black and white hairs, the lance-linear teeth 2 mm long; corolla violet; standard ca. 14 mm long, the limb oblong-obovate, scarcely retuse, auriculate-angular at base, twice as long as the claw; wing-petals ca. 13 mm long, the limb linear-oblong, rounded-obtuse, somewhat shorter than the claw; keel ca. 11 mm long, semicircular, 2/3 as long as the claw; pods cernuous, inequilaterally lance-linear, 17-18 mm long, 2.5-3 mm broad, slightly falcate, gradually attenuate into a straight beak | 2-3 mm long, gradually narrowed into a slender stipe 5-6 mm long, carinate ventrally, applanate dorsally, subtriangular in cross~section, coriaceous, covered with short appressed white hairs, bilocular. Fl. April- May; fr. May. Stony mountain slopes in the witchgrass steppe belt and in the lower part of the juniper belt and needlegrass-and-fescue steppes.— Centr. Asia: Mtn. Turkm. (Centr. and E. Kopet Dagh). Described from Kopet Dagh. Type in Leningrad. 705. A. subternatus Pavl. inBull. Soc. Nat. Mosc. XLIV, 1-2 (1935) 37. Perennial, dwarf; caudex short, woody, the many divisions 1-3 cm long, stout; leaves 2-3 cm long, the petiole slender, canescent by appressed hairs; leaflets 1 or rarely 2 pairs, linear-lanceolate, 8-16 mm long, 1-1.2 mm broad, acuminate, hoary on both sides by appressed hairs; 5943 727 peduncles 3-4 times the length of the leaves, 8-10 cm long, covered with appressed white hairs; racemes elongated, loosely 1-5-flowered; pedicels 1-2 mm long, covered with appressed black and white hairs; bracts lanceolate, 0.5-1 mm long, lanceolate, hyaline, ciliate; calyx cylindric- campanulate, 5-6 mm long, glabrate except for sparse white hairs on the teeth, the tube 5-6 times as long as the lanceolate almost subulate teeth; corolla varying in color, the standard and keel purplish-violet or rarely pink, the wings whitish; standard 16-18 mm long, the limb ovate, round- tipped and scarcely retuse; wing-petals obovate-oblong, entire, round- tipped, 1.5 times as long as the claw; keel 10-12 mm long, obtuse; pods sessile, linear-oblong, 15-20 mm long, 2-3 mm broad, arcuate, terminating in a beak 1-2 mm long, glabrous, erect or horizontally spreading, triquetrous, carinate ventrally, deeply grooved dorsally, bilocular; seeds earshaped, 3-4 mm long, gray. Fl. May. Stony mountain slopes.— Centr. Asia: Tien Shan (Karatau Mountains). Endemic. Described from vicinity of Lake Koinar-kul' and Koinar-bastau natural boundary in Karatau Mts. Type in Moscow. 706. A. karabilicus M.Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 30-40 cm tall; stems 13-35 cm long, erect, terete, stout (up to 4 mm in diameter at base), finely sulcate, densely covered with appressed white hairs; stipules adnate at base to petiole, lance-linear, acuminate, 4-6 mm long, covered on the outside with appressed white hairs; leaves 6-13 cm long, the petiole and the much longer rachis firm, covered with appressed white hairs; leaflets 6-7 pairs, lanceolate or more often linear-oblong, 10-30 mm long, acute, covered above with scattered appressed hairs, green, and hoary beneath by a dense coat of appressed hairs; peduncles equaling to 1.5 times as long as the leaves, firm, virgate, densely covered with appressed white hairs, hoary, 8-11 (18) cm long; racemes 20-25 cm long, loosely many-flowered, the flowers distant; bracts ovate to ovate-lanceolate, acuminate, 3-5 mm long, covered with white and black hairs; pedicels ca. 1 mm long, white-hairy; calyx narrowly cylindric, 17-20 mm long, covered with appressed black and white hairs, striped by uneven distribution of the black hairs, the teeth filiform-linear, ca. 4mm long, the tube 3-4 times as long as the teeth; corolla whitish; standard 26-30 mm long, the limb oblong-obovate, scarcely retuse, constricted above the lower 1/3, slightly and angularly auriculate-dilated below, 1.5 times as long as the claw; wing-petals ca. 25 mm long, the limb linear-oblong, entire, round-tipped, half as long as the claw; keel 22-25 mm long, the limb semioval, obtuse, half as long as the claw; ovary sessile, covered with appressed white hairs. Fl. May. Hillsides.— Centr. Asia: Mtn. Turkm. (Badkhyz). Endemic. Described from vicinity of northern Ata~murat and Karabil'. Type in Tashkent. 707. A. psilophus Schrenk in Bull. Ac. Sc. Pétersb. X (1842) 254; Ldb. Fl. Ross. I (1842) 787. Perennial, 20-30 cm tall; caudex short, buried; stems of the year 7-10 cm long, terete, densely covered with white ascending hairs; stipules adnate to petiole to 1/3-1/2, oblongly triangular-ovate, acuminate, 5-2.5 [?-.] mm long, densely covered with appressed white hairs; leaves 4-10 (12) cm long, the petiole equaling to twice as long as the rachis, both 544 728 729 petiole and rachis slender, covered with short appressed white hairs; leaflets narrowly linear, subobtuse, 12-30 mm long, 1-1.5 mm broad, covered on both sides with appressed white hairs; peduncles equaling to 1.5 times as long as the leaves, covered with short appressed white hairs; racemes 6-19 cm long, loose, very remotely 8-15-flowered; bracts oblong- ovate, acuminate, 1-1.5 mm long, covered with short white hairs; calyx cylindric, 7-8 mm long, rather densely covered with appressed white hairs, the subulate teeth 1.5-2 mm long; corolla purple (?), standard 14-15 mm long, the limb oval, retuse, slightly narrowed in lower 1/4, rather gently angular at base, 4 times as long as the claw; wing-petals 13-14 mm long, the limb oblong, entire, rounded-obtuse, as long as the claw; keel 11-12 mm long, the limb broadly obovate, obtuse, the claw 1.5 times the length of the limb; pods sessile, cernuous, inequilaterally linear-oblong, straight or slightly arcuate, (10) 12-20 mm long, 2.5-3 mm broad, carinate ventrally, rounded dorsally, compressed laterally, coriaceous, covered with short appressed white hairs; bilocular. Fl. June (July); fr. July. Sands.— Centr. Asia: Balkh. (Ili River valley, Alakul' district), Dzu.- Tarb. (Emul' River in Dzhungarian Alatau). Endemic. Described from coastal Alakul'. Type in Leningrad. Series 18. SPECIOSISSIMI Vass.— Subshrubs or herbaceous perennials; leaflets elongated; stems developed; corolla yellow (sometimes yellowish- brown or yellowish~green), 20-26 mm long; pods oblong, from 15 to 20 mm long and (2.5) 3-4 mm broad. — Foothills and low mountains from Mogoltau northward to Karatau. 708. A. speciosissimus Pavl. in Bull. Soc. Nat. Mosc., sér. Biol. XLIV (1935) 39. Perennial, suffrutescent, 50-100 cm tall; caudices elongated, rarely short, the few divisions stout or slender, covered with cinereous at length brown bark; stems of the year erect, 10-20 cm long, stout, densely covered with appressed white hairs, whitish or canescent, scarcely angular; stipules herbaceous, ovate, oblong or lanceolate, 3-8 mm long, white-hairy; leaves 3-10 cm long, the petiole 3-5 mm long, both petiole and rachis canescent; leaflets 3-6 pairs, lanceolate or oblong, (8) 15-30 mm long, (3) 4-10 mm broad, acute, hoary on both sides by appressed hairs; peduncles 15-20 cm long, hoary by appressed white and black hairs; racemes short, capitate, 10-15-flowered, 2-4 cm long; bracts ovate or oblong, 5-10 mm long, villous with white or mixed white and black hairs; calyx at length slightly inflated, broadly cylindric, 17-20 mm long, covered with spreading white and black hairs, the tube 4-5 times as long as the filiform-linear teeth; corolla dingy brown; standard 22-25 mm long, the limb ovate, short-attenuate at apex, retuse; wing-petals equaling the standard, the limb oblong-ovate, entire, the claw 1.5-2 times as long as the limb; keel 22 mm long, obtuse; pods sessile, lance-linear, 17-20 mm long, 3.5-4 mm broad, carinate ventrally, broadly grooved dorsally, pannose with spreading white hairs, terminating in a curved beak 3-5 mm long, bilocular; seeds smooth, olivaceous-brown, ca. 3 mm long, subrhomboidally angular. Fl. May-June; fr. June-July. Slopes of foothills and mountains.— Centr. Asia: Tien Shan (Karatau Mountains). Endemic. Described from the vicinity of Atabaev in Karatau Mountains. Type in Moscow. Rie 545 730 709. A. tumescens M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent; divisions of the caudex up to 10 cm long, slender, ligneous, covered with brownish-cinereous bark; stems of the year 10-15 cm long, densely covered with appressed white hairs, canescent; stipules oblong-lanceolate, 3-4 mmlong, sparsely covered with white hairs; leaves 3-5 cm long, the petiole very short; leaflets 3-4 pairs, lanceolate or oblong, 10-20 mm long, 2-7 mm broad, acute, sparsely covered on both sides with appressed hairs; peduncles subterminal, 0-15* cm long, covered with appressed white hairs, canescent; inflorescence initially dense, capitate, finally becoming loose, elongating up to 10 cm and becoming remotely flowered; bracts lanceolate to oblong, 5-8 mm long, covered with appressed white and sparse black hairs; calyx cylindric, at length becoming inflated, 15-17 mm long, covered with appressed black and white hairs, the linear-subulate teeth 3-4 mm long; corolla yellowish-brown; standard 25 mm long, the limb obtuse, retuse; wing-petals entire; keel 21 mm long; pods (young) lance-linear, covered with short spreading white hairs. Fl. May. Centr. Asia: Tien Shan. Endemic. Described from Karatau (Mt. Agayuk). Type in Tashkent. 710. A. Lipschitzii Pavl. in Bull. Soc. Nat. Mosc. ser. Biol. XLIV (1935) 38. Perennial, suffrutescent or almost frutescent, 15-25 cm tall; caudices covered with blackish-~brown bark; stems densely covered with appressed white hairs, whitish-canescent; stipules ovate-triangular, scarcely adnate to petiole, 1.5-2 mm long, densely covered with appressed white hairs; leaves [?] cm long, the rachis 4-5 times as long as the petiole, bothpetiole and rachis hoary by appressed white hairs; leaflets 3-5 pairs, lanceolate to oblong-lanceolate, 5-12 mm long, 1-2 mm broad, subobtuse to short- acuminate, hoary on both sides by appressed white hairs; peduncles together with inflorescence 4-8 cm long, firm, terete, greatly exceeding the leaves; racemes short, compactly capitate, 3-7-flowered; bracts ovate-triangular, 3-5 mm long, covered with white and, at apex, black hairs; calyx cylindric, 12-15 mm long, villous with long spreading white and black or rarely pure white hairs except for admixture of black hairs on the teeth, the lanceolate-linear teeth 4-5 mm long, the tube 2-3 times as long as the teeth; corolla yellow; standard 22-25 mm long, the limb ovate, very short-acuminate toward apex. Fl. May. Centr. Asia: Tien Shan. Endemic. Described from Karatau (Tosh-Kaul). Type in Moscow. > 711. A. bossuensis M. Pop. in Sched. ad.Herb. Fl. As. Med. XV (1928) 20; A. macrotropis: var brachycarpus MPop., iinet rms tmcee H.F.A.M. No. 353. Perennial, 15-32 cm tall; stems 6-25 cm long, simple, sulcate, sparsely covered with short, appressed, white and, especially beneath the nodes, black hairs; stipules free, triangular-lanceolate, acuminate, 3-5 mm long, green, covered with white and black hairs; leaves 4-10 cm long, the petiole and the much longer rachis sparsely covered with short appressed hairs; leaflets 6-7 pairs, lanceolate to linear-lanceolate, acute, (5) 10-20 mm long, hoary-green, sparsely covered on both sides with appressed white hairs; peduncles equaling the leaves, (4) 6-9 (10) cm long, firm, sulcate, sparsely covered with appressed white and, beneath the * [Probably 10-15 cm. ] H 546 31 inflorescence, mixed white and black hairs; racemes 4-7 cm long, loose, at first closely flowered; bracts lance-triangular, acute, 1.5-2 mm long, covered with black and fewer white hairs; pedicels 1-2 mm long; calyx cylindric, 14-15 mm long, covered with appressed black and fewer white hairs, the tube 2.5-3 times as long as the filiform-linear teeth; corolla greenish-yellow; standard 22-26 mm long, the limb oblong-obovate, narrowed in lower 1/3, ovate and broadly short-ligulate upward, slightly retuse, angular at base, 4-5 times as long as the claw; wing-petals 20-22 mm long, the limb linear-oblong, retuse, somewhat shorter than to about as long as the claw; keel 16-18 mm long, the limb semicircular, acutish, about half the length of the claw; pods subsessile, obliquely ascending, linear-oblong, 12-20 mm long, 2.5-3 mm broad, scarcely falcate, acute at both ends, scarcely carinate to almost rounded ventrally, scarcely grooved or rounded dorsally, compressed laterally, coriaceous, covered with appressed white or mixed white and black hairs, mostly hoary, bilocular. Fl. (March) April-May; fr. May. Rolling foothills and slopes of low mountains. Centr. Asia: Syr D., T. Sh. (W.). Endemic. Described from Mt. Mogoltau. Type in Tashkent. Series 19. DIANTHI Vass.— Herbaceous perennials or subshrubs, with abbreviated obsolescent branched stems and small leaflets (2-7 mm long and 0.5-1.5 mm broad); racemes loosely 2-4 (5)-flowered, short- peduncled; corolla yellow; pods narrowly oblong, from 20 to 30-40 mm long and 3-5 mm broad. W. Tien Shan. 712. A. dianthus Bge. Astrag. turk. (1880) 283; B.A. Fedch. Rast. etre eouy) 5305 Consp. Hin turk. 11) (1906) 2335. hile madzhikenvi 937) WA ee ses). eH. EA. Mi No: 6. Perennial, acaulescent or nearly so, 4-12 cm tall; stems of the year not more than 1 cm long, densely covered with short appressed hairs, white; stipules triangular acute or triangular-lanceolate, acuminate, 2-3 mm long, membranous, faintly covered with short white hairs; leaves 2-5 cm long, the rachis 1.5-2 times as long as the petiole, both petiole and rachis slender, silvery by short appressed hairs; leaflets 6-8 pairs, linear to lance-linear, (2) 3-7 mm long, 0.7-1.5 mm broad, subobtuse, covered sparsely above and densely beneath with appressed white hairs, silvery; peduncles somewhat shorter than to equaling the leaves, 1.5-5 cm long, together with inflorescence 3.5-8 cm long, slender, canescent, densely covered with appressed white and, beneath the inflorescence, mixed white and black hairs; inflorescence loosely 2-3 (5)-flowered; bracts ovate to lance-ovate, acute, black-hairy, 1-1.5 (2) mm long, mostly equaling the pedicel; calyx cylindric, 12-15 mm long, covered with appressed black and ascending white hairs, the linear-subulate teeth (2.5) 3-4 mm long; corolla yellow; standard 20-24 mm long, the limb oblong-obovate, 15-16 mm long, 2-3 times as long as the claw; wing-petals 19-22 mm long, the limb oblong, ca. 7mm long; keel 18 mm long; pods sessile, linear, 3-4 cm long and 4-5 mm broad, abruptly attenuate into a straight beak 1-2 mm long, compressed laterally, thickly carinate ventrally, narrowly and deeply grooved dorsally, prominently cross-nervose, rather sparsely covered with very short appressed white hairs, bilocular. Fl. March-May; fr. April-May Foothills and low mountains — Centr. Asia: Syr D., T. Sh. Endemic. Described from Tashkent district. Type in Leningrad. 547 732 713. A. dianthoides Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent or nearly so, 3-4 cm tall, silvery-white throughout by dense appressed pubescence; stems not more than 0.5-1 cm long; stipules small, 1-2 mm long, lanceolate, acute, adnate to petiole up to the middle, appressed-hairy above; leaves 1-2 cm long, the petiole 2-5 mm long, slender; leaflets approximate, (3) 4-5 pairs, oblong- lanceolate to oblong, 2-7 mm long, ca. 1 mm broad, acute, densely hairy on both sides; peduncles stoutish, 1-1.5 or sometimes up to 3 cm long, predominantly black-hairy beneath the inflorescence; racemes 2-3-flowered; bracts small, triangular-ovate, covered with predominantly black hairs, ca. 1 mm long; pedicels ca. 2 mm long; calyx cylindric, 12-16 mm long, 3-4 mm broad, covered with appressed white and black hairs, the latter predominating on the teeth and on the nerves of the tube, the lanceolate rather thick obtusish teeth 1.5 (2) mm long; corolla yellow, sometimes drying red; standard 26-30 mm long, the oblong limb ca. 9 mm broad in upper part, retuse, graduallynarrowedtowardbase; wing-petals 25-26 mm long, the limb oblong, ca. 9 mm long, about half the length of the claw; keel 23-25 mm long, the claw 2.5 times as long as the ovate round-tipped limb; pods 20-25 mm long, ca. 3 mm broad, attenuate at both ends, terminating in a very short beak, cross-nervose, compressed laterally, grooved on one side and carinate on the other, bilocular, covered with short appressed hairs. Fl. April; fr. April-May. In gravelly serozems, on conglomerates.— Centr. Asia: Pam. Al. (foothills of northern slopes of Alai mountain range). Endemic. Described from vicinity of Mindan village. Type in Leningrad. 714. A. Knorringianus Boriss. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). ' Perennial, acaulescent or nearly so, 5-7 cm tall, grayish-green throughout by appressed pubescence; stems not more than 1.5-2 cm long, white; stipules lanceolate, acute, appressed-hairy above; leaves 2-5 cm long, the petiole 5-15 mm long; leaflets 7-9 pairs, linear to linear- lanceolate, 5-11 mm long, 0.5-1.5 mm broad, acute, covered with appressed hairs, more densely so beneath; peduncles 2-5 cm long; flowers 2-3; bracts ovate, 2 mm long, black-hairy, somewhat shorter than the pedicel; calyx long-cylindric, 20 mm long, covered with predominantly black hairs, the teeth 1-3 mm long; standard 33-36 mm long, the orbicular limb 13 mm broad, retuse, gradually narrowed into a broad claw 20 mm long; wing-petals ca. 30 mm long, the oblong limb ca. 11 mm long; keel 28 mm long, the acutish limb 9 mm long, ovary linear. Fl. March-April. Stony calcareous slopes.— Centr. Asia: Pam. Al. (Nuratau range, mountains of Dzhizak). Endemic Described from Mt. Nuratau. Type in Leningrad. 715. A. Mokeevae M.Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, dwarf, suffrutescent; caudex stout, the short divisions prostrate, angled when young, whitish, ligneous; stems of the year 1-3 cm long, rather stout, angled, whitish by appressed white hairs; stipules ovate, acute, 1-2 mm long, covered with white hairs; leaves 1.5-2.5 cm long, the petiole 3-5 mm long, both petiole and rachis slender, short, partially persistent; leaflets 3-5 pairs, linear-lanceolate or oblong, acute, 3-7 mm long, 0.5-1.5 mm broad, rigid, canescent on both sides by appressed white 837, 1117 548 733 734 hairs; flowers in 2's-4's on short firm axillary peduncles barely 4 mm long, virtually sessile; calyx 15 mm long, rather sparsely covered with appressed white and black hairs, the filiform teeth 5-6 mm long; corolla yellow (?); standard 25 mm long, retuse; wings and keel somewhat shorter than the standard; pods sessile, broadly linear, straight, ca. 20 mm long, 3-3.5 mm broad, acute, slightly compressed laterally, slightly carinate ventrally by prominent suture, rounded dorsally, hoary by appressed white hairs, bilocular. Fl. June. — Centr. Asia: T. Sh. (Karatau Mountains). Endemic. Described from Arystandy River in Chokpak Mountains. Type in Leningrad. Series 20. VIRGATI Vass.— Herbaceous perennials; stems ligneous at base; leaflets fairly long but narrow, 5-9 pairs; racemes loose, remotely flowered; corolla color ranging from purple-violet to white; pods erect, narrowly oblong, from 10 to 20 (25) mm long and 2-3 mm broad. 716. A. virgatus Pall. Astrag. (1800) 20; Bge. Astrag. geront. 1125, II 218; Ldb. Fl. Ross. I, 624 (excl. var. 8. et A. brachylobus); Bge. Astrag. turk. 282; Grossh. Fl. Kavk. II, 322.— A. varius DC. Astrag. (1802) 136.— A. ammophilus Bieb. ex Bess. in Flora XV (1832) II Beibl. Soeasulese Pally Astrag: (1800) tabi.18:— Exs.): HF R:No. 167; Herb. Normale ed. J. Dorfler No. 4237. Perennial, 30-55 cm tall; caudex very short, not more than 1-5 (rarely up to 20) cm long, ligneous at base; stems (10) 15-27 cm long, canescent by a dense coat of white appressed hairs; stipules free, adnate at base to petiole, lanceolate to lance-triangular, (2) 3-5 mm long, covered with white or mixed white and black hairs; leaves 4-8 cm long, sessile, the rachis slender, hoary by short appressed hairs; leaflets (5) 6-9 pairs, linear to lance-linear or linear-lanceolate, rarely oblong or oval-oblong, acute, rarely subobtuse, (5) 10-20 (25) mm long, 1.5-3 mm broad, appressed- hairy on both sides, more densely so beneath; peduncles equaling to 1.5 times as long as the leaves, densely covered with appressed white hairs; racemes loose, rather remotely many-flowered, 8-15 (18) cm long; bracts linear to linear-lanceolate, 2-3 mm long, covered with white or mixed white and black hairs, 3-4 times the length of the pedicel; calyx cylindric, 9-10 mm long, shaggy with appressed or subappressed or spreading white and very few black hairs (the latter sometimes confined to the teeth), the tube 4-5 times as long as the filiform-linear teeth; standard 16-20 mm long, the limb oblong-obovate, obtusish or round-tipped, slightly constricted in lower part, narrowed below, auriculate-angular at base, 4 times as long as the claw; wing-petals 15-19 mm long, the limb oblong, round-tipped, the claw 1.5-2 times as long as the limb; keel 12-17 mm long, the limb semicircular obtusish, half as long as the claw; ovary sessile; pods sessile, erect, linear-oblong, 13-19 (25) mm long, 2-2.5 (3) mm broad, 1.7-2 mm thick, more or less compressed laterally, strongiy carinate ventrally, more or less canaliculate dorsally, abruptly contracted into a straight rigid and mostly obliquely upturned beak 2-4 mm long, coriaceous, densely covered with appressed or ascending white or mixed white and black hairs, bilocular or incompletely so. Fl. May-August; fr. June-August. Needlegrass or needlegrass-and-fescue steppes, rarely meadow steppes, pine forests, sands or rarely on chalk, clayey escarpments, wastelands, at the northern distribution limit often in riverside meadows, in the steppe and semidesert regions, rarely in the desert region, penetrating northward 549 735 into the forest-steppe region; rarely occurring in wormwood deserts. — European part: M.Dnp., Bl., Bes., V.-Don (S. and E.), L. Don, Transv., ly. Ves) Caucasus: Cise Dag.) Wi. 'Siberia:@U: » Dob! »irt) (We ); Gentne Asia: Ar.-Casp. (N.). Endemic. Described from ''Southern Russia'.. Type in London. 717. A. kustanaicus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, prostrate, suffrutescent, the cauline shoots ligneous, up to 2-3 cm long; stems of the year 5-7 mm long, slender, white by a dense coat of appressed hairs; stipules linear-lanceolate, hairy; leaves 3-6 cm long; leaflets 3-5 pairs, filiform-linear, (5) 8-15 mm long and 1-1.5 mm broad, mostly conduplicate, beset on both sides with appressed hairs, more densely beneath; fruiting racemes loose, elongated up to 8-10 cm; peduncles 2-3 times as long as the leaves, green, weakly hairy; bracts ovate, 1-1.5 mm long; calyx ca. 10 mm long, cylindric, the tube 4 times the length of the short teeth; corolla purple (?), 20 mm long; standard elongate-obovate; wing-petals oblong, exceeding the keel; keel 18 mm long; pods erect, narrowly oblong, 15-20 mm long and 2-2.5 (3) mm broad, covered with short appressed hairs, attenuate into a beak 5-10 mm long, grooved dorsally, gently ribbed ventrally, more or less triangular in cross- section. Fl. May-June; fr. from June. Riversides sands.— Centr. Asia: Ar.-Casp. Endemic. Described from Ar.-Casp. (from Naurzum River — Kara-su, former Turg. district). Type in Leningrad. 718. A. macropus Bge. Astrag. geront. I (1868) 125; II (1869) 217; Astrag. turk. 282; Kryl. Fl. Zap. Sib. VI, 1696.— A. subulatus jy. uralensis Pall. Astrag. (1800) 223.—A.virgatus BP. uralensis DC. Prodr. II (1825) 285: Ldb. FI: Ross. I, 625.— Exs.: H.F.R. No. ilo. Perennial, 12-30 cmtall; caudex buried, the aerial divisions slender, 1-3 (5) cm long, ligneous; stems of the year 3-14 cm long, ascending, covered with appressed white hairs; stipules free, lanceolate, the lower ovate-lanceolate or ovate, acuminate, ca. 2 mm long, covered with appressed white hairs; leaves (4) 5-11 cm long, the rachis 2-3 times as long or rarely about as long as the petiole, both petiole and rachis slender, covered with appressed white hairs; leaflets (4) 5-6 (7) pairs, lance-linear. or linear-oblong, obtuse, 7-17 mm long, 1.5-4 (5) mm broad, scatteredly or sparsely covered on both sides or merely beneath with appressed hairs; | peduncles greatly exceeding the stems, 2-3 times the length of the leaves, 3-25 cm long, rather sparsely covered with appressed white and, beneath the inflorescence, mixed white and black hairs; racemes loose (compact in bud), 4-8 cm long, rather remotely flowered, the fruiting raceme often up to 15-18 cm long, with very remote pods; bracts ovate-lanceolate, 1,5-2 mm long, exceeding the pedicel, covered with black or mixed white and black hairs; calyx cylindric, 10-12 mm long, sublanuginose with short spreading or ascending white and black hairs, the linear-subulate teeth (1) 1.5-2 mm long, the tube 4-5 times as long as the teeth; corolla pale violet; standard 20-23 mm long, the limb oblong-oval, retuse, more or less constricted above the lower 1/4-1/5, angularly auriculate-dilated below, 4 times as long as the claw; wing-petals 18-20 mm long, the limb oval- oblong, rounded-obtuse, the claw 1.5 times as long as the limb; keel 15-17 mm long, the limb nearly semiorbicular, slightly concave beneath, 550 736 half the length of the claw; ovary ona stipe ca. 0.5 mm long; pods sessile, erect, linear-oblong, 10-15 mm long, 3 mm broad, compressed laterally, bluntly carinate ventrally, canaliculate dorsally, acutish, abruptly carinate into a subulate oblique beak 2-3 (4) mm long, coriaceous, covered with ascending white hairs thinly interspersed with minute black hairs, almost fully bilocular (unilocular at apex). Fl. May~June; fr. (June) July. Steppes. — European part: V.-Don, L. Don, Transv., L. V.; W. Siberia: U. Tob., Irt.; Centr. Asia: Ar.-Casp., Balkh. Endemic. Described from vicinity of Spasskoe in former Orenburg District. Cotype in Leningrad. 719. A. pallescens. M.B. Fl. taur. caus. III (1819) 489; DC. Prodr.II, 294; Ldb. Fl. Ross. I, 626; Schmalh. Fl. I, 284; Bge. Astrag. geront.I, 125; II, 218.—A.cinereus Bess. Enum pl. Volh. (1820) 29. Perennial, 15-35 cm tall; caudex buried, strong, the aerial divisions 1-2 (3) cm long, ligneous, brown; stems of the year (3) 5-10 (20) cm long, usually ascending, slender, densely covered with appressed white hairs; stipules free, lanceolate, 2-3 (4) mm long, acute, white-hairy on the outside; leaves 3.5-7 (10) cm long, the petiole 1-2.5 cm long, both petiole and rachis slender, covered with appressed white hairs; leaflets 5-6 pairs. linear to linear-oblong, acute or subobtuse, 7-12 mm long, 1-2 (3) mm broad, canescent on both sides by appressed hairs; peduncles twice the length of the leaves and greatly exceeding the stems, (7) 10-25 cm long, sparsely covered with appressed white and, beneath the inflorescence, often mixed white and black hairs; racemes loose, ovaloid, rarely oblong, 3-7 cm long, 5-12-flowered, often elongating in fruit up to 10-12 cm; bracts linear-lanceolate or oblong, 1-3 mm long, covered with white and black hairs; calyx cylindric, 12-13 (15) mm long, covered with ascending or subappressed white and black hairs (often arranged in stripes on the ventral side), the filiform~-linear teeth (2) 3-4 mm long, the tube (2) 2.5-3 times as long as the teeth; corolla white; standard 20-23 mm long, the limb oblong-oval, scarcely retuse, constricted above the lower 1/4, angularly auriculate-dilated below, 4 times as long as the claw; wing- petals 18-21 mm long, the limb inequilaterally lance-oblong, entire, attenuate toward apex, obtusish, the claw 1.5 times as long as the limb; keel 15-17 mm long, the limb semicircular, acutish, about half as long as the claw; ovary sessile; pods sessile, erect, linear-oblong, 11-14 mm long, 2-3 mm broad, bluntly carinate ventrally, grooved dorsally, straight, abruptly contracted into an oblique subulate beak 3-4 mm long, covered with subappressed or rarely ascending white hairs sometimes interspersed with short black hairs, bilocular. Fl. May-June; fr. July. European part: Bl., L. Don. Endemic. Described from the Black Sea region. Type in Leningrad. Series 21. PETUNNIKOVIANI Herbaceous perennials or subshrubs, caulescent, branched; leaflets oval or narrowly oblong to linear, from 10-15 to 30 mm long; racemes very loose, remotely flowered; corolla yellow; pods ovaloid or oblong, from 10 to 30 mm long. Sands of Central Asia. 720. A. Petunnikovii Litw. in Sched. ad Herb. Fl. Ross. (1905) 76; B.A. Fedch. Rast. Turk. (1915) 529.—A. Lubimenkoi B. Fedtsch. in Zhurn. Inst. Botan. AN URSR 26-27 (1938) 229.— Exs.: HFR No. 1410. 551 Perennial, suffrutescent, 20-50 cm tall, strongly branched, grayish by appressed white and fewer black hairs; stipules linear-lanceolate, 7-10 mm long; leaves 2-3 (4) pairs; leaflets linear or oblong to oval, up to 10-30 (50) mm long and 2-3 (10) mm broad, acute, whitish beneath by dense appressed hairs, more sparsely hairy above; racemes loose, elongated, 8-12-flowered; peduncles shorter or longer than the leaves; bracts lanceolate, 1.5-2 mm long; calyx cylindric, ca. 17 mm long, membranous, covered with short appressed white hairs interspersed with isolated black ones, the tube 3-4 times as long as the linear blackish-tipped teeth; corolla yellowtsh-green, ca. 20-25 mm long; standard oval-oblong, 737 8-9 mm broad, retuse; wing-petals about equaling or exceeding the keel; pods reflexed, covered with appressed white hairs, oblong, 15-30 mm long and 4-5 mm broad, acute at both ends, straight or slightly curved, ribbed ventrally, canaliculate dorsally, bilocular, each locule containing 8-12 seeds. Fl. April-May; fr. from June. Sands.— Centr. Asia: Kara Kum. Endemic. Described from Turkmenia (Farab). Type in Leningrad. 721. A. eremobius M.Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 30-40 cm tall} stems 23-30 cm long, stout, subangularly bent, sparsely covered with appressed white hairs; stipules lanceolate, covered with scattered white hairs, ca. 4mm long; leaves 11-14 cm long, the rachis 2-3 times as long as the petiole; leaflets 6-8 pairs, oval, 10-15 (20) mm long, short-cuspidate, green above with scattered appressed white hairs, hoary beneath by a dense coat of appressed white hairs; racemes loose, elongated, up to 14-15 cm long, remotely 12-15-flowered; bracts triangular-lanceolate, covered with black and white hairs, 2.5-4.5 mm long; peduncles half as long as to equaling the leaves; corolla yellow (suffused with blue or pink in dry condition). Fl. April. Sands.—Centr. Asia: Amu D. Endemic. Described from the sands of Katta-kum (between Termez and Dzhar-Kurgan stations). Type in Tashkent. 722. A. rariflorus Ldb. Ic. pl. Fl. Ross. I (1829) 21; Ldb. Fl. Alt. III (lsslyal37) dbs, Bl (Ross: 1.612; ‘Bee. wAstrag. geront. 11. tlio. Kryl; El. Zap: Sib: Vil; 1682: > te; ;iudbi2 Ic.) pl. Fl. Ross. I, tab. da. Perennial, 20-34 cm tall; stems arising at base, many, ligneous below ground level, ascending to suberect, derisely covered with appressed white hairs, firm, 3-7 cm long; stipules adnate at base to petiole, connate to 1/3-1/2, 7-8 mm long, membranous, densely covered with appressed white hairs, the free portion linear-lanceolate; leaves (7) 9-13 cm long, the petiole about as long as the rachis, both covered with appressed white hairs; leaflets 3-5 pairs, linear, 12-20 mm long, densely sericeous on both sides with appressed white hairs; peduncles axillary, 12-20 cm long, exceeding the leaves, covered with short appressed white hairs; racemes very loose, 6-13 cm long, 5-10-flowered, the flowers very distant; bracts ovate- lanceolate, 1.5-2 mm long, covered with appressed white hairs; pedicels ca. 1 mm long, white-hairy; calyx cylindric, 8-9 mm long, densely sericeous with appressed white hairs interspersed with far fewer black hairs, the tube 3-4 times as long as the linear-oblong obtusish teeth; corolla yellow; standard 20-21 mm long, the limb obovate, retuse, angularly attenuate in lower part, ca. 8 mm broad at apex, 3 times the 738 length of the claw; wing-petals 18-19 mm long, the limb linear-oblong, 552 scarcely retuse, somewhat shorter than the claw; keel 15-16 mm long, acutish,; ovary sessile, white-hairy; pods sessile, horizontally spreading to subcernuous, linear-oblong, subfalcate, 25-30mm long, 3.5-4mmbroad, compressed laterally, 1.5mmthick, 3-4 times as long as the calyx, covered with appressed white hairs, coriaceous, bilocular, many-seeded. Fl. June. Steppe meadows.—W. Siberia: Irt. (Irtysh area); Centr. Asia: Balkh. (Chingistau mountains). Endemic. Described from Mts. Chingistau. Type in Leningrad. Note. Ever since Bunge's description (1868), A. rariflorus Lab. has been placed in the section Helmia Bge. In various characters this species resembles A. Petunnikovii Litw. and A. eremobius M.Pop., and has therefore been included in the series Petunnikoviani Vas. This classification should, however, be considered as preliminary, and the same applies in fact to the inclusion of the series as such in the section Xiphidium Bge. Section 93. CYTISODES Bge. Astrag. geront. I (1868) 127.— Dwarf subshrubs or acaulescent perennials; caudices short, exposed, woody; stipules mostly connate, rarely free; leaves imparipinnate, the petiole non-persistent; flowers in short compact heads or umbelliform or rarely oblong and fairly loose racemes; calyx cylindric; bracteoles none; corolla wilting, not persistent in fruit; flowers yellow or dingy violet; pods sessile, oblong or linear or ovoid-oblong, mostly large, rigidly coriaceous, villous with spreading hairs, bilocular, the beak mostly long, rarely short, indurated, subulate, often as long as the body of the pod. — A Central Asian submontane and montane section, distributed in the peripherial parts of the Central Asian mountains from Dzungarian Ala Tauto southwest Pamir- Alai. 1. Subshrubs with short exposed branched caudices............. 23 + Herbaceous perennials, acaulescent or nearly so ............ 9. 2. Leaflets densely scabrous~shaggy with short spreading hairs. (northern foothills of W. Tien Shan —from Dzhambul to Frunze). Ry OMe cis PM Poa Med. Baye (2 eile \s 725. A. promontoriorum Gontsch. + Leaflets covered with appressed or subappressed hairs ........ 3: 3. Leaflets distant, covered with stiff subappressed hairs; pod 17-19 mm long, hairy, oblong-ovoid, the beak not exceeding 2-3 mm; calyx finally becoming somewhat dilated in lower part (mountains, Kirgiz PAN eabaaTW canal) da dies oP a epebiM iy gine, varas ( aypecsdee 729. A. Nikitinae B. Fedtsch. + Leaflets subflabellately approximate at the end of petiole ....... 4. mes, pally covered: with|black and white haird .". -dadoiter. ei a yakeael. 5. + Calyx shaggy with stiff spreading white hairs; dwarf plants, ca. 6 cm tall; leaflets obovate, 11-21 mm long, covered with appressed white hairs; peduncles obsolescent, 0.5-0.7 cm long; corolla purple (N. W. Kanata: orate SNe eee? te.) eee ve 730. A. Georgii Gontsch. 5. Calyx finally becoming somewhat dilated in lower part, ovoid-oblong (E. Tien Shan, Ili river basin). ... 726. A. Krassnovianus Gontsch. fF woaly not becoming dilated toward base =i2 07402 . 2 ei) 0. ee 6. 6. Beak of the pod ca. 5 mm long, much shorter than the body (Karatau IVicmrtATaTS) LA EN MATE Lok, PAE, 0. aoh, Tete 723. A. cytisoides Bge. + Beak of the pod long, subulate, from 2/3 to 1.5 times the length of the Lael EN PATO AOI OD. Abe Lee SNES OPIOID, va RSD SRA, oil 7 740 7. Pods densely shaggy with long stiff spreading hairs (Chu-Ili Mountains) (Ad: eh lot eee 724. A. pseudocytisoides M. Pop. + Pods shaggy or tomentose with soft ascending hairs.......... 8. 8. Pods lance-linear, 4mm broad (Zeravshan, Sarytau, West Tien Shan). WEEE. nde: toch RACRAO I Se EA En RA CA 727. A. dolichocarpus M. Pop. + Pods narrowly linear, ca. 2 mm broad (Western spurs of the Hissar Range from Kugitang to upper reaches of Kashka-darya River, northern slopes of Turkestan Range) ....... 728. A. stenocarpus Gontsch. 9. (Standand i linearsoblong yan aee te en 731. A. macropetalus C.A.M. + Standard ovate or oblong or rhomboidally obovate......... oe TE): 10. Inflorescence abbreviated, subumbellate; standard more than 30 mm lone) podsicovered wathemonochromatic hais = 275 seer a eee nie + Inflorescence a loose ovoid or short cylindric raceme; standard 18-22 mm long; pods spotted by grouping of black hairs (S.W. Pam.- ery eat cerca sp teats ig ee ae na weak 2: Waser ose Geo a Oed Ca t 735. A. Kelleri M. Pop. 11, Meatlets 6-7 pairs (Dzhunsarian Ala Tau). (05). o>: 2 25 wae ESET DREN GPR NAN Eee ON RO Pie aur eri ne eS AS 732. A. ornithorrhynchus M. Pop. +. Ieatlets (LO) Lt720 pains Ps me es 8 oge oe a ae 12. Leaflets 11-15 pairs; pods ovoid-oblong. acuminate into a short subulate beak (W. Tien Shan. Angren River basin)... ......... We gt. ENDED Tere RRS Ts MAREE ee a Ate RAR 734. A. nucleosus M. Pop. + Leaflets (10) 15-20 pairs; pods lance-linear, attenuate into a subulate beak 3-5 cm long (W. Tien Shan: from Mt. Karatau to Sary-Agacha). Ral BRE Rg cbs RA ce Oy A Lt ae mM AO 733. A. xipholobus M. Pop. Series 1. SUFFRUTESCENTES Gontsch.— Subshrubs, the woody aerial caudices strongly abbreviated but distinct; racemes rather compactly capitate. 723. A. cytisoides Bge. Astrag. geront. II (1869) 224; Bge. ex Rgl. in A.H.P. III (1874) 102; Bge. Astrag. turk. 269. Perennial, suffrutescent, 7-10 cm tall; caudex buried, woody, short, the short aerial ligneous divisions 1-5 cm long, mostly prostrate, with very short internodes, covered with lustrous brown bark, giving rise to short young sterile and fertile snoots; stipules adnate to petiole to 1/2-2/3, 6-9 mm long, membranous, white-hairy in upper part, the free portion ovate-oblong, obtusish; leaves 1-2 cm long, the petiole 3-5 (8) mm long, covered with subappressed hairs, bearing at the end 3-5 approximate leaflets; leaflets oblong-obovate, prickly short-cuspidate, cuneate-based, 5-10 (12) mm long, 3-4 (6) mm broad, rigid, hoary on both sides with a dense coat of subappressed hairs; peduncles exceeding the leaves, 1.5- 3cm long, firm, densely tomentose with subappressed white and subumbellate, rather loosely 6-10-flowered, 2.5-3 cm long, the rachis 0.5-1.5 cm long; bracts 5-8 mm long, obtusish, 3-6 mm broad, covered with spreading black and much fewer white hairs, the lower ovate, the upper ovate-oblong; calyx cylindric, 14-15 (20) mm long, densely tomentose with short spreading black hairs and villous with long hairs, the filiform~-linear teeth 4-5 mm long, the tube 2.5-3 times as long as the teeth; corolla yellow; standard 24-30 mm long, the limb obovate, short-attenuate toward apex, 3 times the length of the claw; wing-petals 22-28 mm long, the limb oblong, round- tipped, half as long as the claw; keel 20-25 mm long, the round-tipped limb half the length of the claw; ovary on a stipe 0.5-1 mm long, hairy; pods linear-oblong, subsessile, 19-25 mm long, acute, terminating in a straight beak ca. 5 mm long, compressed laterally, not grooved, 5-6 mm broad, 554 741 742 densely lanuginous~villous with spreading white hairs, bilocular, dehiscent at the tip by 2 valves and a double septum exserted from the pod as a 2- pointed projection. Fl. April; fr. May. Gravelly mountain slopes.— Centr. Asia: Tien Shan (Karatau Range). Endemic. Described from Karatau Mountains. Type in Leningrad. 724. A. pseudocytisoides M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 5-12 cm tall; caudex short woody buried, the divisions woody, smooth, stout, the lateral buried divisions slender, prostrated, 2-10 cm long, covered with brown soon blackening bark, giving rise to the annual shoots; shoots of the year prostrate, (0.5) 1-3 cm long, white by a dense coat of appressed hairs; stipules adnate to petiole up to the middle, the upper ones sometimes connate at base, 5-8 mm long, thinly membranous, covered with more or less appressed white hairs, the free portion triangular, acute; leaves 1.5-2 cm long, the petiole as long as the rachis, both slender, hoary, densely villosulous with short hairs; leaflets 1-2 pairs, approximate at the end of petiole, elongate-obovate to oblanceolate, rounded-obtuse or mucronulate, cuneate-based, hoary on both sides by a dense coat of appressed hairs, ca. 1 cm long and 2-4 mm broad; peduncles 2.5-3 cm long, rather longer than the leaves, villosulous with spreading white and upward mixed white and black hairs; racemes short, capitate, 2-10-flowered, the rachis up to 3 cm long; bracts lanceolate, acute, 5-7 mm long, densely villous with white and black hairs; calyx cylindric, 15-17 mm long, hirsute with appressed black hairs and villous with spreading white hairs, the filiform teeth 2-3.5 mm long; corolla purple (?); standard 25-30 mm long, the limb obovate upward, ligulately short- attenuate at apex and slightly emarginate, narrower inlower part with nearly parallel margins, 2-2.5 times the length of the claw; wing-petals 21-26 mm long, the limb linear-oblong, obtuse, half as long as the claw; keel 19-24 mm long, the limb rounded-gibbous beneath, straight above, obtuse, the claw 2.5 times as long as the limb; ovary on a stipe 1 mm long, densely covered with white hairs interspersed with fewer black ones; pods sessile, linear-oblong, acute, 2-3 cm long, slightly compressed laterally, subcarinate dorsally and ventrally, attenuate into a straight or more or less curved beak (1.5) 2-3 cm long, the entire pod (3) 4-6 cm long, coriaceous, densely villous with long whitish-yellow hairs, bilocular, dehiscing at the tip by separation of the septum from the valves. Fl. May-June; fr. June-August. Stony mountain slopes.— Centr. Asia: Tien Shan (Chu-Ili Mountains). Endemic. Described from Mts. Chu-Ili. Type in Leningrad. 725. A. promontoriorum Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 7-15 cm tall; caudex very short, not exceeding 0.5-2 cm, aerial, woody; stems of the year 1-3 cm long, densely covered with subappressed hairs, white; stipules adnate to petiole up to the middle, connate in lower part, 5-7 mm long, covered with appressed white hairs, the upper ones with a slight admixture of black hairs, the free portion triangular-ovate, acuminate; leaves 2-3 cm long, the petiole slender, densely covered with short spreading whitehairs; leaflets 3-4 subdistant pairs, oblanceolate or rarely oblong-obovate, broadly cuneate at base, acute and minutely mucronulate, 6-15 mm long, densely scabrous~shaggy 555 745 on both sides with short spreading hairs, hoary; peduncles equaling to 1.5 times as long as the leaves, covered with spreading white and black hairs; racemes spherical-ovoid, 3.5-5 cm long, densely 15-20-flowered, the flowers subsessile; bracts dimorphic, the lower ovate, the upper lanceolate, obtusish, 5-8 mm long, densely villous with spreading soft black and white hairs; calyx cylindric, 15-17 mm long, densely villous with spreading black and white hairs, the tube 3 times the length of the linear-subulate teeth; corolla dingy lurid; standard 25-28 mm long, the limb obovate, narrowed in lower 1/4, angular at base, ligulately short- attenuate toward apex, scarcely retuse, 3.5 times as long as the claw, 8-10 mm broad; wing-petals 21-24 mm long, the limb linear-oblong, entire, the claw 1.5 times as long as the limb; keel 19-21 mm long, the limb obtusish, the claw 2.5 times as long as the limb; ovary subsessile, white-hairy. Fl. April. Gravelly and stone-and-clay-covered foothill slopes.— Centr. Asia; Tien Shan (foothills of N. Tien Shan from Frunze district to Dzhambul district). Endemic. Described from the vicinity of Tek-Turmas near Dzhambul. Type in Leningrad. 726. A. Krassnovianus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 10-15 cm tall, the obsolescent aerial woody caudex 0.5-2 cm long; stems of the year 2-3 cm long, densely covered with appressed white hairs; stipules adnate to petiole to 1/3-1/2 and connate below, covered with appressed white hairs, finally glabrate, 4-5 mm long, the free portion triangular-ovate, obtusish; leaves 2-3 cm long, the petiole 2-4 times as long as the rachis, both petiole and rachis covered with subappressed white hairs; leaflets approximate in 5's at the end of petiole, oblanceolate or rarely oblong-obovate, cuneate at base, minutely mucronulate (7) 9-15 mm long, covered on both sides with appressed white hairs, more densely so beneath; peduncles twice the length of the leaves, villosulous with spreading white and black hairs; racemes ovoid, 3-4 cm long, densely 1-15-flowered, the flowers subsessile; bracts dimorphic, the lower oblong-ovate, the upper lanceolate, 5-8 mm long, obtusish, villous with spreading white and black hairs; calyx 16-18 mm long, villosulous with spreading white and black hairs, at length scarcely inflated at base, the tube 4 times the length of the linear-subulate teeth; corolla dingy lurid; standard ca. 32 mm long, the limb obovate, slightly attenuate toward apex, narrowed in lower 1/4, angular at base, 2.5 times as long as the claw, slightly emarginate, wing-petals ca. 27 mm long, the limb oblong, scarcely emarginate, half the length of the claw; keel ca. 25 mm long, the obtuse limb half as long as the claw. Fl. April-May. Centr. Asia: Tien Shan (E.). Endemic (known only from the vicinity of Andrakai). Described from Andrakai in Ili river valley. Type in Leningrad. 727. A. dolichocarpus M. Pop. in Tr. Turkest. Gos. univ. 4 (1922) 62; BL) \Padzhik. \V"W'937)"450.)— ks (2) Ease A IV Nov, daar Perennial, acaulescent or nearly so, 6-15 cm tall, loosely cespitose from short or fairly long divisions of the woody caudex; stipules triangular- lanceolate from a broad base, 4-6 mm long, acuminate, covered with black and white hairs; leaves 1-5 cm long, the petiole as long as the rachis, both covered with short and more or less spreading hairs; leaflets (2) 556 tle NB. AD | \ T=. kr | \ NY, 1 WagJ Ve WKS STEN Wie UT ANA i\ eH) VACHE: AUR NUE a ZYONY) VEEN ie wee NEN FOSS FA! NRT he Wy . ROK NA () 1 x i \ is os VWRABB2AL IS SS LAE | 7} ze oa = PLATE XLIV Pop.—2. A, erioceras Fisch, et Mey.—3. A. Kelleri Astragalus dolichocarpus M. M. Pop.— 4, A. stenocarpus Gontsch. 746 3-6 pairs, lanceolate or rarely oblong-obovate, (5) 8-12 mm long, mucronate, densely covered on both sides with subappressed hairs; peduncles 3-6 cm long, covered with predominantly black and fewer white hairs; racemes obovoid, 3-4 cm long, 10-12-flowered; bracts lanceolate, 4-6 mm long, acute, black-hairy; flowers subsessile; calyx cylindric, 12-17 mm long, covered with subappressed black and far fewer white hairs, striped by predominance of black hairs on the nerves, the teeth linear- subulate, 3-4 mm long, black-hairy; corolla violet; standard 20-26 mm long, the limb rounded-rhomboid, slightly retuse, gradually narrowed into a claw 10-12 mm long; wing-petals somewhat shorter than to equaling the standard, the limb oblong, scarcely retuse, the claw ca. 14 mm long; keel somewhat shorter than the wings, the limb gibbous beneath, scarcely concave above, the claw ca. 14 mm long; ovary subsessile, hairy; pods lance-linear, (17) 20-30 mm long, subsessile, carinate ventrally, carinately rounded dorsally, gradually narrowed into a subulate subarcuate beak (2.5) 3.5-5 mm long, rigidly coriaceous, densely villous with soft appressed white hairs, bilocular. Fl. May-June; fr. June-July. (Plate XLIV, Figure 1). Foothills and intermediate mountain zone.— Centr. Asia: Tien Shan (from Dzhambul district to Tashkent), Pam.-Al. (peripheral mountain zone from Sarytau in northern foothills of Turkestan Range to Yakkabag Mountains). Endemic. Described from Mt. Sarytau and from the vicinity of Tashkent. Type in Tashkent. 728. A. stenocarpus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, suffrutescent, 5-10 cm tall, loosely cespitose, from woody, mostly prostrate caudices 1-4 cm long; stems of the year obsolescent, up to 1.5 (rarely 2) cm long, concealed beneath approximate stipules; stipules adnate to petiole to 1/3-1/2, 3-4 cm long, connate in lower part, triangular, acute, covered on the outside with more or less appressed white hairs; leaves 1-1.5 cm long, the petiole as long as or longer rarely shorter, than the rachis, densely covered with appressed white hairs; leaflets (1) 2-3 pairs, approximate, oblanceolate, rarely oblanceolate-elliptic, round- tipped or acutish, mucronulate, cuneate at base, (3) 4-7 mm long, 1.5-2.5 mm broad, hoary on both sides by a rather dense coat of subappressed hairs; peduncles 1.5-3 (4) cm long, about equaling or exceeding the leaves, covered with subappressed white hairs, these interspersed beneath the inflorescence with isolated black hairs; racemes short, subumbellately capitate, fairly loose, obovoid, 2.5-3.5 cm long, 9-15-flowered, the flowers approximate, subsessile; bracts dimorphic, the lower oblong-ovate, the upper lanceolate, 2.5-4 mm long, covered with white or mixed white and black hairs; calyx cylindric, 11-12 mm long, striped by ascending black and white hairs, the linear-subulate teeth 2-3 mm long; corolla pale violet, finally turning yellow; standard 20-22mm long, the limb rhomboidally oblong-obovate, 7-8 mm broad, ligulately short-attenuate toward apex, scarcely emarginate, 2-3 times as long as the claw; wing-petals exceeded by the standard, 18-20 mm long, the limb oblong, round-tipped, about half the length of the claw; keel-tipped, about half the length of the claw; keel 17-18 mm long, exceeded by the wings, the limb obtuse, the claw 2-2.5 times as long as the limb; ovary on a stipe ca. 1 mm long, densely sericeous; pods subsessile, linear, 3-4.5 cm long, ca. 2mm broad, mostly slightly curved, gradually narrowed into a subulate beak (1.5) 2-3 cm long, coriaceous, densely covered with ascending white 598 hairs, carinate ventrally, rounded dorsally, subterete, bilocular. Fl. (May). June; fr. June-July. (Plate XLIV, Figure 4). Mountain slopes in the high-mountain zone.— Centr. Asia: Pam.-Al. (from Mt. Shakhrisyabz to Kugitang). Endemic. Described from Mull. mountain pass. Type in Leningrad. 729. A. Nikitinae B. Fedtsch. sp. nova in Addenda XI, p. 667. Perennial, suffrutescent, 7-15 cm tall; caudices buried, woody, not more than 1 cm long; stipules adnate to petiole and connate up to the middle, 7-10 mm long, densely covered with appressed hairs, the free portion triangular-ovate or oblongly triangular-ovate, acute; stems of the year 1 (3) cm long, densely white-lanuginous; leaves 2-3.5 cm long, the petiole 747 as long as the rachis, both covered with short spreading hairs; leaflets 2-3 distant pairs, obovate to oblong-obovate, broadly cuneate at base, minutely mucronulate, 7-13 mm long, rigid, hirsute sparsely or scatteredly above and densely beneath with short subappressed hairs; peduncles 1.5 times as long as the leaves, firm, covered with ascending white and black hairs; racemes densely rounded-ovoid, 3-5 cm long, 9-15-flowered, the flowers subsessile; bracts dimorphic, the lower ovate, the upper oblong- ovate to lanceolate, subobtuse, 7-10 mm long, densely covered with spreading white and black hairs; calyx cylindric, at length scarcely inflated, 15-18 mm long, densely villous with spreading white and black hairs; the tube 2.5-3 times as long as the linear-subulate teeth; corolla dingy-yellow; standard 25-27 mm long, the limb oblong-obovate, sordid at base, ligulately and broadly short~attenuate at apex, scarcely emarginate, 8-9 mm broad, 3-4 times the length of the claw; wing-petals 22-24 mm long, the limb linear-oblong, scarcely emarginate, the claw 1.5 times as long as the limb; keel 20-22 mm long, the limb obtusish, the claw 2-2.5 times the length of the limb; pods sessile, linear-oblong, 16 mm long, 4 mmbroad, carinate ventrally, convex dorsally, terminating in a slender reflexed beak 2-3 mm long, densely lanuginous~-villous with spreading white hairs, oblong-ovoid, 17-19 mm long, 10 mm broad, coriaceous, bilocular, few- seeded. Fl. April-May; fr. May. Slopes of the intermediate mountain zone.— Centr. Asia: Tien Shan (Kirgiz Alatau). Endemic. Described from Mt. Poskel'dy. Type in Leningrad. Series 2. TRIFOLIATI Gontsch.— Dwarf subshrubs with short stout aerial caudices; calyx white-shaggy; corolla purple; leaves composed of 3 approximate leaflets. 730. A. Georgii Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, dwarf, nearly acaulescent, ca. 6 cmtall; caudices many, strongly abbreviated, aerial, stout, 1.5-3 cm long, 0.5-0.7 cm thick; annual stems 0.5-1 cm long, concealed beneath stipules and leaf petioles; stipules adnate to petiole in lower part or up to the middle, 5-6 mm long, densely covered with appressed white hairs, the lower ovate, the upper oblong, acutish; leaves (1.5) 2-3 cm long, consisting of 3 approximate leaflets, the petiole covered with appressed white hairs; leaflets obovate, round-tipped, cuneate at base, 0.5-1 cm long, 11-21 mm long, (5) 6-11mm broad, rather loosely covered on both sides with appressed white hairs, 748 rigid; peduncles 0.5-0.7 cm long, white-shaggy; inflorescence capitate, 999 749 compactly spherical, 2-2.5 cm long; bracts lanceolate, 8-9 mm long, acuminate, white-ciliate on the margin and slightly white-hairy on the outside; calyx cylindric, 14-15 mm long, densely hirsute-villous with spreading white hairs, the tube 1.5 times as long as the subfiliform-linear teeth; corolla purple; standard 22 mm long, the limb oval, scarcely | emarginate, slightly constricted above the base, auriculate-angular at base, twice as long as the claw; wing-petals 20 mm long, the limb obovate-oblong, | scarcely emarginate, half the length of the claw; keel equaling the wings; | ovary borne on a very short stipe, narrowly linear, the style 3 times the length of the ovary. Fl. May. Gravelly mountain slopes.— Centr. Asia: Tien Shan. Known solely from a single site on the water shed plateau of Mardanbai-bulak in northwestern part of Mt. Karatau, whence it was described. Type in Leningrad. Series 2. MACROPETALI Gontsch. — Herbaceous perennials, acaulescent or nearly so; flowers large in short umbelliform racemes; standard linear-oblong. 731. A. macropetalus C.A.M. in Bull. Phys.- Math. Acad. Pétersb. Ill (1844) 307; Bge. Astrag. geront. II, 195; Bge. Astrag. turk. 279. Perennial, 10-20 cm tall, loosely cespitose from the short divisions of a woody buried caudex; stems obsolete or very short, not exceeding 2cm; stipules adnate to petiole up to 1/2 or 2/3, 8-9 mm long, densely covered with white hairs, the free portion triangular-oblong, acutish; leaves 7-[?]* (17) cm long, the petiole and the much longer rachis densely shaggy with short spreading white hairs; leaflets 15-18 pairs, oblong to oblong-oval, round-tipped, (5) 7-12 (15) mm long, 2-3 (5) mm broad, glabrous above, hoary beneath by a dense coat of spreading hairs; peduncles shorter than the leaves, 5-8 (11) cm long, firm, terete, finely sulcate, rather densely covered with spreading white and, beneath the inflorescence, mixed white and black hairs; racemes 5.5-6.5 cm long, loosely 6-10- flowered, the rachis 1-2 cm long; bracts oblong-ovate, acute, 2.5-3 mm long, covered with black and fewer white hairs; pedicels 3-5 mm long, covered with spreading black and white hairs; calyx cylindric, 15-20mm long, densely covered with subappressed black and white hairs, the linear teeth 2-3 mm long; corolla yellow; standard 43-47 mm long, the limb linear-oblong, round-tipped and emarginate, angular at base, about 3 times as long as the claw, 9-10 mm broad at the middle; wing-petals 42-45 mm long, the limb linear, round-tipped, ca. 3 mm broad, the claw somewhat more than 1.5 times the length of the limb; keel 40 mm long, the limb linear-oblong, slightly enlarged at apex, truncately obtuse, the claw somewhat more than 1.5 times the length of the limb; ovary hairy, ona stipe 3 mm long; pods subsessile, borne ona slender stipe ca. 2 mm long; oblong-ovoid, slightly flacate, acuminate into a straight beak ca. 2mm long, 18-20 mm long, inflated, 5 mm broad, 7-8 mm thick, flattened ventrally, shallowly and broadly grooved dorsally, rigidly nutlike- coriaceous, profusely and finely cross-wrinkled, rather sparsely villous with soft spreading white hairs, bilocular, few-seeded. Fl. April (May); fr May-June. Centr. Asia: Balkh., T. Sh. (Karatau). Endemic. Described from Attasu River. Type in Leningrad. * [Figure illegible in Russian copy. ] 560 750 Series 3. SUBUMBELLATI Gontsch.— Perennials with short buried woody caudices; flowers large, in subumbellate racemes; standard oblong- obovate or ovate. 732. A. ornithorrhynchus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, herbaceous, nearly acaulescent, from a vertical taproot and a much divided root crown; stems up to 1 cm long, stout, white- pannose; stipules free, (?) triangular-ovate, herbaceous, hairy on the outside; leaf petiole somewhat longer than the rachis, marcescent; leaflets 6-7 pairs, elliptic, acute, 5-6 mm long, hoary on both sides by spreading hairs; peduncles stout, firm, barely exceeding the leaves, 5-7 mm long, covered with spreading white hairs; racemes short, capitate, 3-10- flowered; bracts oblong, ca. 5 mm long, submembranous, covered with short and predominantly black hairs; calyx narrowly cylindric, 13-15 mm long, shaggy with long spreading ventrally black and dorsally white hairs, the filiform teeth ca. 3 mm long, much shorter than the tube; corolla yellow (?); standard 25 mm long, the limb ovate, scarcely and broadly emarginate; limb of wings bilobate, 7 mm long; keel 20-21 mm long, the limb ca. 6 mm long, obtuse, the claw 14-15 mm long; pods sessile, cylindric, subulate, nearly straight, up to 5 cm long, 2-3 times the length of the calyx, subcarinate ventrally, narrowly grooved dorsally, attenuate into a straight beak 5-8 mm long, covered with stiff spreading white hairs; seeds cylindric, 3-4 mm long, olivaceous. Fl. June. Stony slopes of bald mountains.— Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala Tau). Endemic. Described from the vicinity of Mulala station of Turk. — Sib. railway. Type in Alma Ata. Note. This species, known solely from the description of M.G. Popov, has been provisionally placed by us in the series Subumbellati. Various characters, such as shape of standard, wings, and pod, etc., display marked differences as compared with the other species of this series. 733. A. xipholobus M. Pop. in sched. ad Herb. Fl. As. Med. fasc. Pavel o28) 39: Lxs.: Herb. FY. As’”°Med. No. 374: Perennial, acaulescent, 10-22 cm tall, loosely cespitose from the divisions of a buried woody caudex; stipules green, adnate to petiole and connate up to the middle, 6-8 mm long, densely covered with short white hairs, the free portion triangular acute; leaves (4) 10-22 cm long, the petiole and the much longer rachis rather slender, densely covered and largely shaggy with soft spreading hairs; leaflets (10) 15-20 pairs, elliptic to lance-elliptic, brown, (6) 10-15 mm long, villosulous on both sides with short spreading hairs; peduncles exceeded by the leaves, 7-15 cm long, rather loosely villosulous with spreading white and, beneath the inflorescence, mixed white and black hairs; racemes short (rachis 1-2 cm long), obovoid, 5-6 cm long, loose, subumbellate, 8-15-flowered; bracts ovate-lanceolate, acute, 5-8 mm long, densely covered with black and fewer white hairs; pedicels ca. 1 mm long, covered with black and fewer white hairs; calyx long-cylindric, 21-25 mm long, densely shaggy with spreading long white and shorter black hairs, the tube 4.5-5 times the length of the narrowly linear teeth; corolladingy yellow; standard 37-45 mm. long, the limb oblong-obovate, 11-13 mm broad at apex, broadly shert- ligulate at apex, slightly emarginate, narrowed in lower 1/3, angular 561 at base, 2.5 times the length of the claw; wing-petals 35-42 mm long, the limb oblong, slightly dilated toward apex, half as long as the claw; keel 34-40 mm long, the claw 2.5 times the length of the obtuse limb; pods sessile, lance-linear, slightly curved, (3.5) 5-7 cm long, subtrigonous, 5-7 mm broad, 5-8 mm thick, carinate ventrally, canaliculate or subapplanate dorsally with a concave suture, attenuate into a slightly upturned firm subulate beak (2.5) 3-5 cm long, rigidly nutlike-coriaceous, rather sparsely villous with soft spreading white hairs, bilocular. Fl. April-May (June); fr. May-June. Fine-grained or somewhat gravelly slopes in the foothills and the lower mountain zone (up to 1100 m).— Centr. Asia: Tien Shan (foothills of W. Tien Shan, Darbaz station and Sary-Agach to Mt. Karatau, Karatau Mountains). Endemic. Described from vicinity of Sary-Agach station northwestward from Tashkent. Type in Tashkent. 734. A. nucleosus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, acaulescent, 8-12 cm tall; caudex short, buried, the divisions short, indurated, stoutish; stipules large, adnate to petiole up 751 to 1/8-1/2, ovate to oblong-lanceolate, up to 10 mm long, shaggy with appressed hairs; leaves 5-10 cm long, the petiole and the much longer rachis densely shaggy with soft more or less spreading hairs; leaflets 11-15 approximate pairs, lanceolate to elliptic-lanceolate, acute or subacute, 5-10 mm long, 2-4 mm broad, densely subpannose-villous on both sides with soft appressed hairs; peduncles 5-9 cm long, firm, angled, tomentose-villous, about equaling the leaves; racemes apparently capitate, few-flowered; bracts ovate, ca. 5 mm long, covered with black hairs; flowers large, 25-30 mm long; pods ovoid-oblong, 20-30 mm long, 10-14 mm broad and thick, strongly inflated, scarcely curved, short- beaked, rounded ventrally and dorsally, subterete, very rigid, densely subpannose-villous with soft subappressed hairs, fully bilocular, essentially indehiscent; seeds earshaped, 3-4 mm long, brown, alveolar, flat. Fl. May. Stony mountain slopes.— Centr. Asia: Tien Shan (Angren river basin). Endemic. Described from the vicinity of Ablyk village. Type in Tashkent. Series 4. RACEMOSI Gontsch.— Perennials with very short buried caudices; stipules connate; flowers medium size, in loose racemes; standard rhomboidally obovate. 735. A. Kelleri M. Pop. in Trans. of the Scient. Soc. of Turkestan, Voli (923) 36" Perennial, acaulescent, (6) 10-17 cm tall, from the very short divisions of a short stout buried caudex; stipules adnate to petiole and connate to high up, 5-10 mm long, covered with white and black hairs, densely white- lanuginous near the base, the free portion triangular~acute or acuminate; leaves (3) 4-10 cm long, the rachis slightly longer to 1.5 times as long as the petiole, both petiole and rachis hirsute and villous with spreading hairs; leaflets 8-12 pairs, broadly elliptic to.obovate, rarely suborbicular, obtuse or mucronulate, (4) 7-10 mm long, hoary on both sides with stiff ascending hairs; peduncles firm, erect, 4-8 cm long, about equaling the leaves, hirsute and villous with short spreading white and upward mixed white and black hairs; racemes ovoid or short-cylindric, 4-8 cm long, 562 752 rather loosely many- flowered; bracts linear, 4-8 mm long, acuminate, green, covered with white and black hairs; pedicels ca. 1 mm long; calyx cylindric, 9-10 mm long, covered with short appressed black hairs, these interspersed with few long white hairs, the tube 2-3 times the length of the lance-~subulate teeth; corolla purple; standard 18-22 mm long, the limb rhomboid-obovate, ligulately attenuate toward apex, more or less emarginate, 1.5 times as long as the claw; wing-petals ca. 17 mm long, the limb lance-oblong, somewhat shorter than the claw; keel 13 mm long, rounded-gibbous beneath, straight above, acute; pods on a stipe 3-5 mm long, linear-oblong, 8-18 mm long, compressed laterally, carinate ventrally, narrowly grooved dorsally, acute, coriaceous, villous with spreading white hairs and dotted by grouping of short black hairs, bilocular, the beak straight or scarcely curved, rigid, 8-13 mm long, covered with subappressed white and occasionally some black hairs. May. (Plate XLIV, Figure 3). Foothills and low mountains chiefly in association with mottled strata Centr. Asia: Pam.-Al. (peripheral western spurs; western part of Turkestan Range). — Mal'guzarskie Mountains, Kugitang Mountains, Baisun low mountain district). Endemic. Described from Mt. Kugitang, from vicinity of Baisun and from Dzhizak, Type in Tashkent. Section 94. AMMODENDRON Bge. Astrag. geront. I (1868) 128. — Shrubs or mostly subshrubs; aerial caudices more or less developed, the branches persisting for 2 or 3 years, vested with bicuspidate hairs; branches of the year terete, densely covered with appressed white hairs; stipules connate and adnate to petiole; leaves imparipinnate; leaflets 1-3 (4) pairs; racemes loose or less often somewhat compact, borne on short axillary peduncles; bracteolesnone; calyx cylindric-campanulate or subcampanulate, 4-8 mm long; corolla violet or purple; standard not more than 17 mm long; wings shorter than the standard and exceeding the keel; pods stipitate or sessile, ovoid or ovaloid to ovaloid-oblong, 5-8 mm long or rarely linear-lanceolate . up to 18 mm long, loosely textured, coriaceous, bilocular, commonly white-villous, rarely covered with appressed hairs or glabrous. A predominantly Central Asian section, distribution through Central Asia, Iran, and very rarely in Asia Minor and Arabia. Mostly psammophytes or associated with more or less sandy substrata; rarely petrophilous plants of foothills and low mountains; very rarely occurring on fine-textured mountain slopes. 1. Petioles and leaf rachises on the long branches of preceding years persistent as spines up to 5 cm long; calyx covered with black or mixed black and white hairs; leaflets oblong-linear; pods sessile (S. Pam. Pipi! TVET ViaNley) a icles aie acu 736. A. wachschi B. Fedtsch. + Petioles not persistent on any branches of the preceding year, or indurescent and persistent but not spiny. ....-......-...... 2. ZG yx Covered with) white NAIMS 2 590 3 hse 6 2 a ces 2 i 2 oe ae 13. + Calyx covered with white and black hairs................. 3. 3. Ovary and pod covered with very short appressed hairs or glabrous. 4. Tr Ovanv onde pod hirsute Ol ViNlOUs'. cs 6) alain fe totic as lances 6. 963 4. Dwarf subshrubs not exceeding 15 cm in height, with very short caudices; racemes loose, 1-4 cm long; pods sessile, inequilaterally 753 oblons*i6-8iimm! lone \(Kopet Dash badkhy aii: gk isk faa as 5. + Taller shrubs, upto 1m in height, with long caudices; racemes very short, 7-15 cm long; pods cernuous, ona slender stipe 5-6 mm long, lance-linear, 13-18 mm long, carinate ventrally and dorsally (S. W. Para Alin) ok at. Ae Oe EN MRL 737. A. Oldenburgii B. Fedtsch. 5. Standard 12 mm long, the limb rounded-oval, as long as the broad claws (Badkhiyz)). MO.p0O.. Ain ROU Ut 747. A. badghysi M. Pop. + Standard 15 mm long, the limb oblong-ovate, 1.5 times as long as the claw (Kopet Dagh: Kyzyl-Arvat)..... 748. A. arvatensis Gontsch. 6. Peduncles not developed; flowers 1-2 in the leaf axils, borne on pedicels 1.5-2 (3) mm long; dwarf subpulvinate subshrubs, 6-10 cm tall, the strong buried caudex with woody underground divisions (Bolshie Balkany, Centr. Kopet Dagh) ... 1744. A. deserti M. Pop. + Flowers in loose or fairly compact sessile or short-peduncled FACEMAE STH MIL TR EES TR LEE UT) 3 RS". ESS aS A 7. (srOviay and" podUsessSile Rin. .eliiatMeinh. Ehmine te ReIk. thal ree. c CneO Rane nee 8. +, Ovary ‘andipod istipitave Gi) 2 f REM MOL” Llama tialian=) iA eats bets) teem a3 8. Leaflets 2-3 pairs, oval-oblong, 1.5-2.5 cm long, (0.4) 0.5-1 cm broad; calyx 8-9 mm long; standard 15-16 mm long, the limb broadly rhomboid-oval, 3-4 times as long as the claw (Kopet Dagh, Kyzyl- Nee Hh Ie TEL aR Rei nea igh in 746. A. Androssovianus Gontsch. + Leaflets 1-2 pairs, oblong to linear-oblong, (8) 10-15 mm long, 2-3.5 mm broad; calyx 6-7 mm long; standard 12-13 (15) mm long, the rounded-ovate limb as long as the claw (Badkhyz, W. Kopet Dagh) . BR ee mS Caeser ee Nea Mae a ay ON RTA CNN 9) 745. A. nigrescens M. Pop. 9. Upper eee composed of 7 leaflets, the lower ones 7-9; pods borne on a stipe 1-2 mm long; racemes 2.5-5 (9) cm long, loosely 5-15- flowered (E. Transc.: W. Caspian coast). 742. A. igniarius M. Pop. + Upper leaves composed of 375, leaflets ..°) - 0.) 274-22 +=) yee 10. 10. Racemes short, compactly 2-7-flowered, the flowers approximate; pods borne on a stipe 1 mm long (E. Caspian coast: Mangyshlak, KePASMOVIOGSIC) ear lth ahr ce mare a edn Guna 743. A. turcomanicus Bge. + RaCEMeS, LOGSE se wa ca riianps oki utbin ban oe Shee Roles enced tage ate Walia dale tee cae ial 11. Standard 15-17 mm long; racemes 4-6 (8) cm long (Caspian region of Western Eurkmenial) oe sheen na CP NO ALY bing Wy, ER ee ie, Tides aap es Oa Iu ys PON ee itn BIS 741. A. transcaspicus Freyn. 12. A subpulvinate dwarf shrub, 20- 30 cm tall; caudex very short; peduncles spuriously terminal, apparently forming an extension of the annual branches; pods lance-oblong or oblong-ellipsoid, 8-9 mm long, 754 villous with spreading hairs, borne on a stipe 2-3 mm long (W. Kopet IDENel ove octal Ry eee er lc cra mnr Ba'ath Perey at oti mee teers 740. A. podolobus Boiss. + A shrub up to 80 cm tall; caudices well developed, woody; peduncles axillary; pods lance-linear, 13-18 mm long, sericeous with appressed- hairs, borne on a stipe 3-4 mm Jong (S.W. Pam.sAl,).. -.. «-..-8.: BIBT: SHA. HARARE gee) “kz, fe Neate Da aid oa dng 738. A. Vvedenskyi M. Pop. 13) Leaflets) slabrou's) above) tba Wet iy ihe Pails ie he et ae 14. +" Leafletsimone jor less hainyiiaboOves .-y-yakey/-y itis) anes celbols yi ee 15. 14. Racemes rather compact, 2.5-3 cm long; calyx 4-5 mm long, the upper ones trifoliate; leaflets obovate or oval to oval-oblong; pods 8-12 mm long (Balkh., Kyz. K., Kara K.). 754. A. paucijugus C. A. M. 964 + Racemes elongated, loose, remotely flowered, (12) 18-25 cm long; calyx 8-9 mm long; upper leaves consisting of 1 leaflet; leaflets elliptic to narrowly linear; pods 6-7 mm long (E. Kara-Kum, iS acid onaea) oa ir aaa a anhleguagl Ute ie Bap tats 755. A. longipetiolatus M. Pop. fo-aaupper leaves) consisting of! tleaflet. 10% LW. AhOl Sh Mots Sone 16. JU ppemleaves)(COnNSISting) Often leatletsy)a.. aoe) Mil es ey. 1. 16. Standard 11-13 mm long; calyx 5 mm long; pods oblong, 8-12 mm long, villous with ascending hairs, cernuous, ona stipe ca. 3 inm long (valleys of S.W. Pam.-Al.). 739. A. excedens M. Pop. et Vved. + Standard 8-10 mm long; calyx 3.5-4.5 mm long; pods oblong-ovate, 5-6 mm long, sessile, villous with spreading hairs (S. and W. Kyzyl Kum, sands of the right bank in the Amu Darya region, Kara Kum). Ree ey ener. Mur ire ae® 2 Pawan behoddaven. 756. A. unifoliolatus Bge. 17. Ovary and pod glabrous; pods borne on a slender stipe 5-6 mm long, lance-linear, 13-18 mm long, carinate dorsally and ventrally, CeEnnouS, (S- WiaWamasAl sii. joy cts ps 737. A. Oldenburgii B. Fedtsch. MOAN eanG PO vhainy, Lick co. ee abe Leh mio! os pt bt eT foie Brae 18. 18. Ovary and pod borne on a short or fairiy long stipe; leaflets mostly linear or oblong, rarely ovaloid and then petiole not indurescent and BACEIMES MOOSE iM TLSiem Mone Mk 9. WER RULER RS BUNS a SAL 19. + Ovary and pod sessile, rarely on a very short stipe and then leaflets lanceolate or elliptic; petioles strongly indurescent, arcuately Kecunved; (racemes =siem jlonp;-mostly compact), 4%; 209.)..2 2 21. 19. Upper leaves composed of 7 leaflets lower ones 7-9; pods borne on a stipe 1-2 mm long (E. Caspian coast).... 750. A. hyrcanus Pall. SOMUipper leaves*composed of 3 7o%leaflets: .. 2S. PRAY OY 2 ee 20. 755 20. Standard 17-20 mm long; pods borne on a stipe (3) 4-6 mm long (N. Aral region, Emba river, Dzhebel and western Turkmenia).... LUAU RRC TOON ie ab SoD BG 749. A. ammodendron Bge. + Standard 13-15 mm long; pods borne on a stipe 1-2 mm long (?) CSO ain A) Wit Be. OR Ss panes 760. A. surchanicus M. Pop. 21. Leaflets oval or oblong-oval or lanceolate or elliptic; standard amo, MPM LOMN Ee WIA SMe) IRE ee. LR OB a EC FA ig OTR OEE 22. + Leaflets linear or lance-linear or oblong or oblong-obovate, rarely Gbtongcovallandithen standard 11512 mm longi i. scans wwe. 23. 22. Petioles becoming lignified and persistent on the old branches, arcuately recurved, subspinescent; leaflets 2 pairs (along Syr Darya, Buona) eningrad idistrict,and\ below)ems) ods, 9. 2eiatend adi. ae. ede ea: PE URS 2. ATMA ws WINE POS 759. A. chodshenticus B. Fedtsch. + Petioles not becoming lignified; leaflets 3 (4) pairs (sands of Balkhash- Alakul' depression, Muyun-kumy)...... 752. A. cognatus C.A.M. 23. Standard 8-9 mm long; racemes many-flowered; bracts lanceolate; calyx-tube 2-3 times as long as the teeth; leaflets 1-2 pairs (Ili Pe TeV Le ya MUI A) eat tote et eM aa ate coh 763. A. iliensis Bge. + Standard 11-15 mm long, rarely 9-11 mm and then racemes few- flowered; bracts oblongly triangular-ovate or rounded-ovate, calyx- tube 4-5 times as long as the teeth, and leaflets 2-3 pairs, rarely 24. Leaflets linear to lance-linear, rarely oblong or lance-oblong... 25. + Leaflets oblong-obovate or obovate or obovate-oblong, round-tipped, cuneate at base; upper leaves composed of 3 leaflets, the lower ones of 5-7 (foothills of Kopet Dagh flatland region, southeastern Kara-kum, 565 Badkhyz, Kugitang range in S.W. Pamir-Alai, Iran) ........... AC wana we cepa te Lit. Peay ee 762. A. macrobotrys Bge. 25. Racemes 2-8 (13) cm long, loose but bearing approximate flowers and fruits .subcomipactin Aruita7egat. Oe Fee 2s, Feet 26. + Racemes short or long, with very distant flowers and fruits .... 27. 26. Standard 13-15 mm long; calyx 6-7 (8) mm long; leaflets 1 pair, sometimes the lower ones 2 pairs, linear or linear-oblong (Ili river valley, Balkhash region, Muyun-kumi). 757. A. brachypus Schrenk. + Standard 11-12 mm long; calyx 5-6 mm long; leaflets 2 (3) pairs, the upper ones 1 pair, oblong or linear-lanceolate, rarely oblong-oval (Karakalsk region and southern Kyzyl-kum, E. Kara-kum sands of Sundukli in the Amu Darya region)..... 758. A. villosissimus Bge. 756 27. Limb of standard broadly rhomboid-oval, 2-3 times the length of the miuchinarrowersclawe heats. oot ere ee oad eel eae ee 28. + Limb of standard broadly oval, as long as or 4-5times the length of the broadaclawltiwwantothit dy Amc PER. wel lt ae thee lh ee eee 29). 28. Standard 13-14 mm long (Zaisan depression and sands of Irtysh in westerm,Moneolia)ani te Grol ritikt so ted 751. A. Gebleri Fisch. + Standard 9-11 mm long (sands of Emby river valley, northern Aral region, Muyun-kumy, periphery of Tien Shan sandy strip of Balkhash- Alakultj andi Mdwriverwalley)- sey ac vie: 753. A. karakugensis Bge. 29. Calyx campanulate; caudices 1.5-3 cm long; branches of the preceding year 4-7 cm long; pods covered with short appressed white hairs, inequilaterallly, oblong). ca. 1.0)mamrlongy).. ta-i-aiec-c me ee ee att Sense gett 4 AVM SS oriihd ems 765. A. spartioides Kar. et Kir. + Calyx cylindric-campanulate; caudices 5-25 cm long; branches of the preceding year 10-25 cm long; pods villous with spreading hairs, 5-7 mm onto ys pect ek lel beget Loomer oyeetee = Grodan Pe 30. 30. Upper leaves composed of 3 leaflets; leaflets linear; standard (11-12 mm long; pods 5-6 mm long (S.W. Pam.-Al., lower Kafirnigan, BadkhyZ) petites) dt SOY Let bata. 764. A. aiwadzhi B. Fedtsch. + Upper leaves composed of 5 leaflets; leaflets linear or lance-oblong; standard 12-13 mm long; pods 7 mm long (W. Kyzyl-kum and adjacent section of Syr-Darya river valley)........-....+.-... eG. LORE. ooo o ON oes BEALS He 762. A. macrocladus Bge. Series 1. ACICULARES M. Pop.-— Petioles and leaf rachises persistent on the branches of the preceding year, finely spinescent, acicular; flowers ca. 15 mm long in short loose few-flowered racemes; calyx covered with black hairs; ovary sessile, hairy. 736. A. wachschi B. Fedtsch. in Trudy Tadzhik. bazy Akad. Nauk SSSR II (1936) 151. A strongly branched shrub up to 1 mm tall; branches stout, flexuous, covered with gray bark; branches of the current year short, not exceeding 1.5 cm, borne in the axils of spines 2-4 (5) cm long representing the petioles and rachises of preceding years; stipules connate at base, triangular, 1-1.5 mm long; leaves 1.5-2 cm long; leaflets 1-3 pairs, linear, rounded-obtuse, 8-12 mm long, 1.5-2 mm broad, covered with scattered appressed hairs; peduncles 1-3 cm long, somewhat exceeding the leaves; racemes short, loosely 3-5-flowered; bracts ovate, ca. 1 mm long, covered with scattered black hairs, barely exceeding the pedicel; calyx cylindric-campanulate, 6-7 mm long, covered with appressed black hairs, 566 “157 758 the tube 4-5 times the length of the linear-subulate teeth; corolla violet; standard ca. 15 mm long, the limb oblong-oval, slightly emarginate, angularly short™attenuate at base, 4 times as long as the claw; wing-petals equaling the standard, the linear-oblong limb as long as the claw; ovary sessile, hairy. Fl. March. Centr. Asia: Pam.- Al. (Vakhsh river valley, upper Kurgan-Tyube). Endemic. Described from Vakhsh river valley. Type in Leningrad. Series 2. XIPHOAMMODENDRON M. Pop.-— Petioles not spinescent; flowers ca. 14 mm long; racemes fairly long, loose, remotely flowered; calyx covered with white or black or mixed white and black hairs; pods on a stipe (3) 4-6 mm long, cernuous, lance-linear, compressed laterally, glabrous or appressed hairy. 737. A. Oldenburgii B. Fedtsch. in Tr. Tadzhik. bazy Akad. Nauk SSOR II (1936) 153; Fl. Tadzhik. V, 451. A strongly branched shrub up to 1 cm tall; caudices 5-7 mm thick, covered with grayish-brown bark; branches of the current year linear, (5) 13-22 cm long; stipules connate to high up, 1.5-3 mm long, white-ciliolate on the margin, the free portion triangular; leaves (1.5) 2-4 cm long, the petiole together with rachis 4-7 mm long, covered with short white hairs; leaflets 1-3 pairs, oval to oblong, (0.6) 1-2 cm long, obtusish, covered on both sides with appressed white hairs; peduncles exceeding the leaves, 5-12 cm long, densely covered with appressed white hairs; racemes about equaling the peduncles, 7-13 cm long, very loosely 10-15-flowered; bracts lanceolate, 2/3-1/2 the length of the pedicel, ca. 1 mm long, covered with short black or mixed white and black hairs; calyx cylindric-campanulate, 6-7 mm long, densely covered with appressed white or rarely black or mixed black and white hairs, the tube 5-6 times the length of the triangular or triangular-lanceolate teeth; corolla violet; standard ca. 14 mm long, the limb ovate, round-tipped or scarcely emarginate, 1.5-2 times as long as the claw; keel 9 mm long; ovary glabrous, on a stipe 1-1.5 mm long, pods cernuous, borne on a slender stipe 5-6 mm long and equaling the calyx-tube, inequilaterally lance-linear, acuminate at both ends, 13-18 mm long, 3-4 mm broad, terminating in a straight subulate beak 1.5-3 mm long, carinate dorsally and ventrally, compressed laterally, membranous, glabrous, bilocular. Fl. April-May; fr. May-June. (Plate XLV, Figure 1). On outcrops of mottled layers (red sandstone).— Centr. Asia: Pam.-Al. (southwest: foothills in the valleys of Surkhan and lower reaches of Kafirnigan rivers). Endemic. Described from the lower reaches of Kafirnigan River. Type in Leningrad. 738. A. Vvedenskyi M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). A shrub resembling A. Oldenburgii B. Fedtsch.; distinguished by the appressed-sericeous pod; calyx covered with white and black hairs; pod linear, borne on a stipe 3-4 mm long. Occupying an intermediate position between A. oldenburgii B. Fedtsch. and A. excedens M. Pop. et M. Kult. Outcrops of mottled layers.— Centr. Asia: Pam.-Al. (S.W.). Endemic. Described from Shirabad valley. Type in Tashkent. 967 759 Series 3. EXCEDENTES M. Pop.— Petioles not spinescent; flowers 11-13 mm long; racemes loose, elongated, remotely flowered; calyx covered with white hairs; pods borne ona stipe 3 mm long, cernuous, linear-oblong, appressed-sericeous. 739. A. excedens M. Pop. et M. Kult. in Not. Syst. ex Herb. Hort. Petrop. IV (1923) 157; Fl. Tadzhik. V, 453.— A. Schartusi B. Fedtsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS VII (1940) 169, £.10.— Exe.) Herbiebl. Ass) MedwNo:.364: A strongly branched shrub 50-80 cm tall; caudices short, stout, covered with pale brown longitudinally splitting bark; branches of the preceding year ligneous, 7-20 cm long, often ribbed, grayish-white by a dense coat of short appressed hairs; branches of the current year 15-25 cm long; stipules connate to high up, 1-2 mm long, ciliate-margined, bidentate, the teeth broadly triangular; leaves 3-6 (8) cm long, the petiole together with rachis 1-1.5 (2) mm long, shorter than the terminal leaflets, becoming indurated but not spinescent; leaflets 3, on upper leaves 1, lanceolate to linear, acute, (17) 20-35 (50) mm long, 1.4 (5) mm broad, densely covered on both sides with appressed hairs; peduncles equaling or slightly exceeding or rarely 1.5 times as long as the leaves, together with inflorescence much longer than the leaves; racemes (5) 8-19 cm long, mostly many-flowered, the flowers very distant; bracts ovate-lanceolate, 1-2 mm long, acute, white-ciliolate; calyx cylindric-campanulate, ca. 5 mm long, densely covered with appressed white hairs, the tube 4-5 times the length of the lanceolate acuminate teeth; corolla violet; standard 11-13 mm long, the limb ovate, slightly emarginate, 2-3 times as long as the claw; wing-petals 10-12 mm long, the limb linear-oblong, round-tipped, as long as the claw; keel 9-10 mm long; ovary on avery short stipe, hairy; pods cernuous, the 2-3 mm long stipe exceeded by the calyx-tube, linear-oblong, 8-12 mm long, 3-3.5 mm broad, inflated, obtusish at both ends, not grooved, attenuate into a beak 2 mmlong, membranous, densely villous with soft ascending white hairs, bilocular, Fl. March; fr. April-May. Desert sands, less often stony sites.— Centr. Asia: Pam.- Al. (S.W.: sands of lower reaches of Vakhsh, Kafirnigan and in Amu-Darya from lower Vakhsh to Kelifa, Shirabad valley). Endemic. Described from vicinity of Amu-Darya river and from Bishkent valley in S. Tadzhikistan. Type in Tashkent. Series 4. PODOLOBI M. Pop.— Petioles not spinescent; flowers 13-14 mm long; racemes loose, borne on spuriously terminal peduncles; calyx covered with white and black hairs; pods borne on a stipe 2-3 mm long, lance-oblong or oblong-ellipsoid, villous. 740. A. podolobus Boiss. et Hohen. in Boiss. Diagn. ser.I, 9 (1849) 49; Bge. Astrag. geront.II, 228; Boiss. Fl. or. II, 489. Perennial, subpulvinate, dwarf, suffrutescent, 20-30 cm tall; caudex very short, stout, with short divisions, covered with grayish-brown longitudinally fibrillose-splitting bark; branches of preceding year ligneous, 6-11 cm long, densely appressed-hairy, gray; branches of the current year 4-10 cm long, slender, stipules almost fully connate and adnate to petiole, bidentate, 1.5-2 mm long, covered with appressed white hairs; leaves 1.5-3 cm long, the petiole together with rachis not exceeding 2-7 mm; leaflets 1-2 pairs, on lower leaves 2-3 pairs, 10-20 mm long, 1-2 mm 568 760 broad, covered on both sides with short appressed hairs; peduncles together with inflorescence 10-20 cm long, mostly as long as the inflorescence, spuriously terminal, apparently forming an extension of the current year's branches; racemes loose, remotely 10-17-flowered; bracts ovate, ca. 1 mm long, black-ciliolate; calyx cylindric-campanulate, 6-7 mm long, covered with subappressed black and white hairs, the tube 3-4 times the length of the linear-subulate teeth; corolla violet (?); standard 13-14 mm long, the limb ovate, emarginate, twice as long as the claw; wing-petals 11-12 mm long, the limb linear-oblong, slightly emarginate, as long as the claw; keel 10-11 mm long; ovary ona stipe ca. 1 mm long, white-hairy; pods borne on a stipe 2-3 mm’ long, lance- oblong to oblong-ellipsoid, acuminate at both ends, 8-9 mm long, 3-4mm broad, terminating in a straight subulate beak ca. 2 mm long, slightly inflated, carinate ventrally, finely canaliculate dorsally, membranous, densely villous with long soft spreading hairs, bilocular. Fl. April-May; fr. May. Foothill slopes, often on marls or clays, rarely on sandy sites. —Centr. Asia: Mtn. Turkm. (W. Kopet Dagh). Gen.distr.: Iran. Described from N. Iran (vicinity of Derbend in El'burs Mountains). Type in Geneva; cotype in Leningrad. Series 5. TRANSCASPICI M. Pop.— Petioles not spinescent; flowers (13) 15-17 mm long; racemes short, loose, few-flowered; calyx covered with black and white hairs; pods borne on a stipe (1) 2-3 mm long, mostly ellipsoid-oblong, villous. 741. A. transcaspicus Freyn in Mém. Herb. Boiss. 13 (1900) 16 et in Bull. Herb. Boiss. 2 sér.V (1905) 1018. Perennial, suffrutescent, 30-80 cm tall; caudices stout, angular, 7-15 cm long, covered with brownish-gray longitudinally splitting bark; branches of the preceding year ligneous, (10) 15-43 cm long, fairly stout, angled-ribbed, whitish-gray, very densely covered with short appressed hairs; branches of the current year 7-20 cm long, except for rarely occurring brachyblastic forms; stipules adnate in lower part to petiole and connate to high up, acutely bidentate, 3-5 mm long, subhyaline, covered with short appressed white hairs; leaves (2) 3-4 (8) cm long, the petiole together with rachis 1.5-3 (5) cm long, densely covered with short appressed white hairs; leaflets 2 pairs, narrowly linear, (1) 1.5-2.5 (4) cm long, 1-2 mm broad, densely covered on both sides with appressed white hairs; peduncles exceeded by the leaves, 1.5-2.5 cm long, densely covered with short appressed white hairs often interspersed with isolated black hairs, together with inflorescence 5-9 (11) cm long; racemes loose, 4-6 (8) cm long, remotely 5-12-flowered; bracts ovate, 1-1.3 mm long, black- or black-and-white-ciliolate; pedicels equaling to 1.5 times as long as the bracts; calyx cylindric-campanulate, 7-8 mm long, densely covered with ascending black and fewer white hairs, the tube 2.5 times the length of the linear-subulate teeth; corolla violet (?); standard 15-17 mm long, the limb rounded-ovate, emarginate, 1.5 times as long as the claw; wing- petals 13-15 mm long, the limb oblong, slightly enlarged toward apex, as long as the claw; keel 11-11.5 mm long; ovary on a stipe ca. 1 mm long, white-hairy; stipe of pod 1/2-2/3 the length of the calyx-tube, 2-3 mm long; pods oblong-ellipsoid or ellipsoid-oblong, 6-10 mm long, 2.5-3 mm broad, acute at both ends, terminating in a curved subulate beak ca. 2 mm 969 761 long, membrabous, densely villous with long spreading white hairs, pilocular. Fl. April-May; fr. (April) May. Sandy and argillaceous~sites in the desert region; rarely on slopes of foothills and mountains.— Centr. Asia: Ar.-Casp. (Mangyshlak peninsula), Kara K. (vicinity of Krasnovodsk, Molla-kor, Akhcha-kuima, Dzhebel, Cheleken peninsula, Mtn. Turkm. (foothills of W. Kopet-dagh). Endemic. Described from the vicinity of Krasnovodsk. Type in Geneva. 742. A. igniarius M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946).— A. hyrcanus Boiss. Fl. or. II (1872) 488, non Pall. ; Grossh. El. Kavk. Ul) 320. Exs. 2 Herbs ie Cauc.- No. lag: Perennial, suffrutescent, (15) 30-55 cm tall; caudices scarcely developed or short, stout, 2-10 (20) cm long, covered with grayish- brown bark, often prostrate; branches of the preceding year ligneous, mostly short; 3-12 cm, rarely up to 34 cm long, densely covered with short appressed hairs, grayish-brown; branches of the current year 8-35 cm long; stipules at base connate and adnate to petiole, ovate- triangular, acute, 3-4 mm long; leaves (3) 4-7 (10) cm long, the petiole together with rachis 2.5-5 (8) cm long, densely covered with short appressed hairs, whitish, indurescent and persistent but not spinescent; leaflets 3-4 pairs, linear, 8-13 (23) mm long, obtusish, densely coverd on both sides with appressed white hairs; peduncles mostly exceeded by the leaves, 0.5-2 (3) cm long, densely covered with appressed white hairs, together with inflorecence (2.5) 4-10 cm long, often equaling the leaves; racemes loose, 2.5-5 (9) cm long, remotely 5-15-flowered; bracts ovate, ca. 1mm long, acute, covered with short white and black hairs, somewhat exceeded by the pedicel; calyx cylindric-campanulate, 6-9 mm long, densely covered with ascending black and white hairs, the tube 3-4 times the length of the linear-subulate teeth; corolla violet (?); standard 13-17 mm long, the limb ovate, scarcely emarginate, slightly longer than to 1.5 times the length of the claw; wing-petals 12-16 mm long, the limb linear-oblong, as long as or somewhat longer than the claw; keel 11-14 mm long; ovary hairy, on a stipe not exceeding 1 mm; pods borne on a stipe 1-2 mm long, inequilaterally ellipsoid-oblong, acute, obtusish at base, 9-12 mm long, 3.5-5 mm broad, carinate ventrally, scarcely canaliculate dorsally, somewhat inflated, rigidly coriaceous, densely villosulous with soft spreading hairs, bilocular, the curved subulate beak ca. 2 mm long. Fl. May; fr. (May) June. Sands.— Caucasus: E. Transc. Endemic. Described from Apsheron peninsula. Type in Leningrad. Series 6. TURCOMANICI M. Pop.— Petioles not spinescent; flowers 14-20 mm long; racemes short, compactly few-flowered or peduncle obsolescent and flowers 1-2 in the leaf axils; calyx covered with black and white hairs; pods subsessile or borne on a stipe up to 1 mm long, oblong, villous. 743. A. turcomanicus Bge. Astrag. geront. II (1869) 227; Astrag. turk., 295. Perennial, suffrutescent, prostrate, 15-30 (35) cm tall; caudices short, rugose, brown, mostly closely ribbed, up to 2 cm in diameter, covered with brownish-gray longitudinally splitting bark, with short divisions; branches of the preceding year ligneous, (1) 2-10 cm long, gray, densely 570 762 763 appressed-hairy; branches of the current year (1) 6-18 (30) cm long; stipules 1-1.5 mm long, yellowish, covered with short appressed white hairs; leaves 1-2 cm long, the petiole together with rachis (2) 4-10 (15) mm long, densely covered with appressed white hairs; leaflets 1 (2) pairs, linear, 1-2.7 cm long, 1-3 mm broad, obtuse, covered on both sides with appressed white hairs; peduncles 3-20 (30) mm long, covered with appressed white hairs often interspersed with scattered longer black hairs, together with inflorescence 3-5 cm long; racemes short, compact, 2-7- flowered, the flowers approximate; bracts ovate, acute, ca. 1 mm long, equaling the pedicel, covered with short white and black hairs;. calyx cylindric-campanulate, 6-8 mm long, densely villous with short ascending or spreading white and black hairs, the tube 4-5 times the length of the lance-linear or triangular-lanceolate teeth; corolla purple (?); standard 14-20 mm long, the limb broadly ovate, scarcely emarginate, somewhat shorter than the claw; keel 11-15 mm long; ovary on a very short stipe, white-hairy; pods borne on a slender stipe 1 mm long, ellipsoid-oblong, acute at apex, obtusish at base, 6-8 mm long, ca.3 mm broad, terminating . in a curved subulate beak 2.5-3 mm long, carinate ventrally, rounded or carinately rounded dorsally, rigidly membranous, densely villous with long soft spreading white hairs, bilocular. Fl. May (June); fr. (May) June. In gravel-and-clay, clay, and often solonchak deserts (the latter containing Nanophytonand Anabasis); chalky mountains, limestone fissures, or rarely consolidated sands.—Centr. Asia: Ar. Casp. (Mangyshlak peninsula), Kara K. (Mts. in vicinity of Krasnovodsk). Endemic. Described from the Airakli foothills, on the eastern coast of the Caspian Sea. Type in Leningrad. 744. A. deserti M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, dwarf, pulvinate, 6-10 cm tall; caudex strong, buried, woody, the aerial divisions ligneous, covered with gray bark; branches of the current year 1.5-3 (4) cm long, slender; stipules adnate to petiole and connate throughout their length, ca. 2 mm long, white-hyaline, sparsely white-ciliate on the margin; leaves 1-1.3 cm long, the petiole very short, covered with appressed white hairs; leaflets narrowly linear, 7-10 mm long, hoary on both sides with a dense coat of short appressed hairs; flowers 1-2 on densely black-hirsutulous pedicels 1.5-2 (3) mm long; peduncles wanting; bracts ovate-triangular, covered with black and white hairs, half as long as to equaling the pedicel; calyx campanulate, ca. 7mm long, covered with appressed black and fewer white hairs, the linear-subulate teeth 1-2 mm long; corolla violet (?); standard ca. 15 mm long, the limb ovate, scarcely emarginate, as long as the broad claw; wing-petals ca. 13 mm long, the limb oblong, slightly enlarged at apex, rounded-obtuse, about as long as the claw; keel ca. 12 mm long; ovary on avery short stipe, hairy; pods subsessile, oblong, ca. 7 mm long, ca. 3 mm broad, not grooved, terminating in a curved subulate beak (2) 3-5 mm long, thinly coriaceous, densely villous with spreading white hairs, bilocular. Fl. April; fr. May-June. Gravelly mountain slopes.— Centr. Asia: Mtn. Turkm. (Mt. Dushak in Kopet Dagh and Bolshie Balkhany Mountains). Endemic. Described from Mt. Bolshie Balkhany. Type in Leningrad. 571 Series 7. NIGRICANTES M. Pop.— Petioles not spinescent; flowers 12-13 (15) mm long; racemes loose, elongated; calyx covered with black and white hairs; pods sessile, oblong, villous. 745. A. nigrescens M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 35-60 cm tall; caudices short, covered with grayish~brown bark, the divisions mostly virgate; branches of the preceding year ligneous, 15-35 cm long, densely covered short appressed hairs, white, short in brachyblastic forms; branches of the current year 5-20 cm long; stipules 1.5-2.5 mm long, the lower connate to high up, the upper up to the middle, the free portion triangular; leaves 1.5-2 and, on dolichoblasts, 5 cm long, the short petiole and the rachis covered with appressed white hairs; leaflets 1 (2) pairs, oblong to linear-oblong, obtuse or round-tipped, sometimes slightly enlarged toward apex, (8) 10-15 mm long, 2-3.5 mm broad, rigid, covered on both sides with short appressed white hairs; peduncles equaling to twice the length of the leaves, covered with appressed white hairs, together with inflorescence 5-6 cm long on the brachyblasts and 12-17 cm on the dolichoblasts; racemes loose, on the brachyblasts, few-flowered and 3-5 cm long, on the dolichoblasts many- flowered and 10-15 cm long, the flowers distant; bracts ovate, ca.1 mm long, covered with short black and white hairs; calyx campanulate, 6-7 mm long, covered with white and black hairs; the lance-subulate teeth 1-1.5mm long; corolla lilac; standard 12-13 (15) mm long, the limb rounded-ovate, slightly emarginate, as long as the claw; wing-petals 11-12 mm long, the limb oblong, round-tipped, as long as the claw; keel 10 mm long; ovary sessile, hairy; pods sessile, oblong (together with vesture ovaloid), 6-7 mm long, not grooved, terminating in a beak ca. 1 mm long, spreading, the lower ones cernuous, thinly coriaceous, densely villous with soft spreading white hairs, bilocular. Fl. April; fr. May. Sands or rarely (in W. Kopet Dagh) on rocky and clayey slopes of loess hills in the foothills. — Centr. Asia: Kara K., Mtn. Turk. (W. Kopet Dagh). Endemic. Described from Turkmenia. Type in Leningrad. Series 8. MACRANTHI Gontsch.— Petioles not spinescent; flowers 15-16 mm long; racemes loose, elongated; calyx covered with black and white hairs; pods sessile, ovaloid. 746. A. Androssovianus Gontsch. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). 7164 Perennial, suffrutescent, 28-30 cm tall; caudices short; branches of preceding year ligneous, 10-18 cm long, often ascending; branches of the current year 15-20 cm long; stipules adnate to petiole and connate at base, triangular, acute, 3-4 mm long; leaves 4.5-8 cm long, the petiole together with rachis 2.5-5 cm long, covered with appressed white hairs; leaflets 2-3 pairs, oval-oblong, 1.5-2.5 cm long, (0.4) 0.5-1.0 cmbroad, round- tipped, green on both sides with scattered appressed hairs; racemes sessile, elongated, loose, 18-20 cm long, remotely many-flowered; bracts ovate-lanceolate, 2-3 mm long, acute, white- and black-ciliate, somewhat exceeding the pedicel; calyx broadly cylindric-campanulate, 8-9 mm long, densely covered with ascending white and black hairs, the tube 2-2.5 times the length of the lance-subulate teeth; corolla violet; standard 15-16 mm long, the limb broadly rhomboid-oval, emarginate, 7-8.5 mmbroad, 972 765 3-4 times as long as the claw; wing- petals 14-15 mm long, the limb oblong, slightly enlarged toward apex, obtuse and scarcely emarginate, as long as the claw; keel 10-11 mm long, the limb obtuse, about as long as the claw; ovary sessile, hairy; pods (imperfectly mature) sessile, ovaloid, densely villous with soft spreading white hairs, cernuous. May. Pebbles of mountain streams.— Centr. Asia: Mtn. Turkm. (mountains of Kyzyl-Arvat). Endemic. Described from the vicinity of Kyzyl-Arvat. Type in Leningrad. Series 9. HUMILES Gontsch.— Petioles not spinescent; flowers 12-15 mm long; racemes short, loose, few-flowered; calyx covered with black and white hairs; pods sessile, inequilaterally lance-oblong or oblong, glabrous or covered with very short appressed hairs; dwarf subshrubs, 14-15 cm tall. 747. A. badghysi M. Pop. in Not. Syst. ex Herb. Hort. Petrop. IV, (1923) 157. Perennial, suffrutescent, dwarf. ca. 15 cm tall; caudex short, stout, with short divisions, covered with longitudinally splitting bark; branches of the preceding year 2-3 cm long, whitish-gray, densely covered with appressed white hairs; branches of the current year 5-7 cm long; stipules almost fully adnate to petiole and connate, 2-2.5 mm long, pale yellow, sparsely covered with short appressed white hairs; leaves 1.3-2 cm long, the petiole together with rachis 0.2-1 cm long, densely covered with very short appressed hairs; leaflets 1-2 pairs, lance-oblong, 6-8 mm long, 1.5-2 (2.5) mm broad, appressed-hairy on both sides; peduncles barely exceeding the leaves, covered with short appressed white hairs; racemes loose, 3-4 cm long, 5-10-flowered; bracts ovate, 1 mm long, sparsely covered with short black hairs; calyx cylindric-campanulate, 6 mm long, covered with subappressed white and black hairs, the tube 3 times the length of the linear-subulate teeth; corolla violet (? ); standard 12 mm long, the limb rounded-oval, scarcely emarginate, as long as the claw; wing- petals 11 mm long, the limb oblong, scarcely emarginate, as long as the claw; keel 9 mm long; ovary glabrous or covered with short appressed hairs, sessile; pods sessile, inequilaterally lance-oblong, 6-7 mm long, 2-2.5 mm broad, acuminate into a slightly reflexed subulate beak 2 mm long, strongly carinate ventrally, rounded dorsally, thinly coriaceous, glabrous or covered with very short appressed hairs, bilocular. Fl. March- April. Centr. Asia: Mtn. Turkm. (Badkhyz). Endemic. Described from the vicinity of Lake Er-oilan-duz. Type in Tashkent. 748. A. arvatensis Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, suffrutescent; dwarf, ca. 14cm tall, strongly branched; caudices short; branches of the preceding year up to 9cm long; branches of the current year 4-9 cm long; stipules 1-1.5 mm long, sparsely covered with short white hairs; leaves 1-1.5 cm long, the petiole together with rachis 3-6 mm [sic] long, densely covered with appressed white hairs; leaflets in 3's, obovate-oblong, 5-10 mm long, 1.5-3 mm broad, obtuse, fairly rigid, covered sparsely above and densely beneath with short appressed hairs; peduncles exceeded by the leaves, 3-8 mm long; racemes 1-2 cm long, 2-6-flowered; bracts ovate, 1 mm long, covered with short 573 766 white and black hairs; calyx cylindric-campanulate, 5 mm long, covered with short appressed white and black hairs, the tube 4 times the length of the lance-subulate teeth; standard 15 mm long, the limb oblong-ovate, scarcely emarginate, 1.5 times as long as the claw; wing-petals 10.5 mm long, the limb oblong, as long as the claw; keel ca. 10 mm long; ovary sessile, covered with very short hairs; pods sessile, unequilaterally oblong, obtusish, ca. 8 mm long, ca. 3 mm broad, terminating in a straight subulate beak ca, 1mm long, carinate ventrally, scarcely canaliculate dorsally, thinly coriaceous, covered with short appressed white hairs, bilocular. Fl. April. Centr. Asia: Mtn. Turkm. (vicinity of Kyzyl-Arvat). Endemic. Described from the vicinity of Kyzyl-Arvat. Type in Leningrad. Series 10. HYRCANI M. Pop.— Petioles not spinescent; flowers i5-20 mm long; racemes loose, short, few-flowered; calyx covered with white hairs; limb of standard ovate, somewhat longer than the claw; pods on a stipe 1-4 mm long, ellipsoid or ovaloid-cordate. 749. A. ammodendron Bge. in Arb. Naturf. Ver. Riga I (1848) 233; Bge. Astrag. geront. II, 228; Bge. Astrag. turk. 296.— A. hyrcanus Kar. in Bull. Soc. Nat. Mosc. XII (1839) 152, non Pall. Frutescent from stout caudices up to 1 cm in diameter covered with grayish-brown bark; branches of the preceding years 10-35 cm long, finely sulcate, densely appressed-hairy, grayish-white; shoots of the current year stout dolichoblasts up to 25-30 cm long, and slender white brachyblasts to 5 cm long; stipules adnate to petiole in lower part or up to the middle, connate to high up, 3-5 mm long, faintly covered with short appressed white hairs, the free portion broadly triangular acute; leaves (3) 4-7 (9) cm long, the rachis 2-3 times the length of the petiole, the petiole together with rachis (2) 4-6 (7) cm long, covered with short appressed white hairs, indurescent and persistent, not spinescent; leaflets 2-3 pairs, narrowly linear, acute, (1) 2-4 cm long, (1.5) 2-3 mm broad, covered on both sides with short appressed white hairs, hoary; peduncles 1.2-3 cm long, exceeded by the leaves, together with inflorescence 4-7 (15) em long, covered with short appressed white hairs; racemes loose, the flowers distant; bracts lanceolate, acuminate, 1.5-2 mm long, covered with short white hairs; calyx campanulate, 7-9 mm long, densely villous with short more or less spreading white hairs, the tube 2-3 times the length of the linear-subulate teeth; corolla purple-violet (?); standard 17-20 mm long, the limb ovate, emarginate, 3 times the length of the claw; wing- petals 15-18 mm long, the limb oblong, slightly enlarged toward apex, somewhat longer than the claw; keel 10-11 mm long; ovary on a stipe 1-1.5 mm long, hairy; pods borne on a stipe (3) 4-6 mm long, mostly somewhat exceeding the calyx-tube, ovaloid-oblong, obtusish, 7-11 mm long, 3-5 mm broad, slightly carinate ventrally, scarcely canaliculate or almost rounded dorsally, rigidly membranous, villous with soft spreading white hairs, bilocular. Fl. May-June; fr. June (Plate XLV, Figure 2). Exposed sand dunes, often partly consolidated sandhills or rarely flat sands.— Centr. Asia: Ar.-Casp. (northern Aral region, Emby river valley, easterly part of southern rebuilt Ust'-Urta). Kara K. (vicinity of Dzebel). Endemic. Described from Airakli foothills, from Irgiz River and from Syr-Darya River near Malibash. Type in Paris. 574 PLATE XLV 1. Astragalus Oldenburgili B. Fedtsch.—2. A. ammodendron Bge.— 3. A. paucijugus Gc A.M. 769 770 750. A. hyrcanus Pall. Astrag. (1800) 25; Bge. Astrag. geront. II, 225: db. FI) Ross: I; 610.—A. "karakugpensi's Grossn HI: Kavkall (1930)"3204 non Be | Ic.: "Palla ews 22° Perennial, suffrutescent, 20-50 (75) cmtall; caudices woody, very short, usually not exceeding 5 cm; branches of the preceding year ligneous, 4-17 (35) cm long, covered with short appressed hairs, grayish-brown; branches of the current year 8-25 (65) cm long; stipules adnate to petiole and connate up to the middle, 2-3 mm long, covered with appressed white hairs, the free portion triangular, acute; covered with appressed white hairs; leaves 4-9 cm long, the petiole (2) 3-8 cm long, white by short appressed hairs; leaflets 3-4 pairs, narrowly linear, 5-15 (20) mm long, 1-2 mm broad, covered sparsely above and densely beneath with appressed hairs; peduncles 1-3 cm long, mostly exceeded by the leaves, together with inflorescence (6) 7-17 cm long, covered with short appressed white hairs; racemes loose, 5-10 cm long, remotely 8-15-flowered; bracts broadly lanceolate, 2-2.5 mm long, white-hairy, equaling or somewhat exceeding the pedicel; calyx cylindric-campanulate, 8-9mm long, densely covered with appressed white hairs, the tube 2-2.5 times the length of the linear-subulate teeth; corolla violet (?); standard 15-18 mm long, the limb ovate, scarcely emarginate, somewhat longer than the broad claw; wing- petals 13-16 mm long, the limb linear-oblong, round-tipped, barely shorter than the claw; keel 11-13 mm long; pods on a stipe 1-2 mm long, ellipsoid- oblong, acute, obtusish at base, 8-11 mm long, 3-4.5 mm broad, carinate ventrally, narrowly applanate dorsally, terminating ina slightly curved subulate beak 2-3 mm long, rigidly membranous, densely villous with soft spreading white hairs, bilocular. Fl. May-August (September); fr. June- September. Coastal (coast of the Caspian Sea) or rarely lacustrine sands.—Caucasus: Dag., E. Transc. Endemic. Described from the vicinity of Derbent. Type in Leningrad. Series 11. SEPTENTRIONALES Gontsch.— Petioles not spinescent; leaflets 2-3 pairs; flowers 9-15 mm long; racemes mostly short, remotely flowered; calyx covered with white hairs; pods sessile, ovaloid or rarely oblong-ovaloid, villous. 751. A. Gebleri Fisch. ex Bong. Verz. Pfl. Saissang-Nor (1841) 24; Bge. Astrag. geront. II, 225; Bge. Astrag. turk. 295; Kryl. Fl. Zap. Sib. vil, 1702. Perennial, suffrutescent, 30-40 cm tall; caudices short, covered with grayish-brown bark; branches of the preceding year ligneous, 5-15 cm long, stout, densely covered with short appressed hairs, silvery-white; branches of the year 7-15 cm long; stipules adnate at base to petiole and connate to high up, 3-5 mm long, obtusely bidentate; leaves (3.5) 4.5-10 cm long, the petiole together with rachis (3) 4-9 cm long, densely covered with short appressed white hairs; leaflets 2 (3) pairs, the lowermost 1 pair, linear-lanceolate, acute, 1.5-3 cm long, 0.2-0.5 cm broad, densely covered on both sides with appressed silvery hairs; peduncles 1-3 cm long, exceeded by the leaves, together with inflorescence about equaling the leaves, 4-10 cm long, covered with appressed white hairs; racemes loose, (2) 3-4 cm long, remotely 4-6-flowered; bracts ovate, ca. 1 mm long, somewhat exceeded by the pedicel; calyx cylindric-campanulate, 5-5.5mm long, densely covered with appressed white hairs, the tube 2-2.5 times the length of the linear teeth; corolla pale purple; standard 13-14mm long, 576 the limb broadly rhomboid-oval, 5.5-6 mm broad, deeply emarginate, twice the length of the claw; wing-petals 12-13 mm long, the limb linear- oblong, obtuse, somewhat longer than the claw; keel 8-10 mm long; ovary sessile, hairy; pods sessile, ovaloid, obtuse, terminating in a subulate beak ca. 2 mm long, inflated, not grooved, slightly carinate ventrally, rounded dorsally, 5-7 mm long, 3.5-5 mm broad, densely villous with soft spreading white hairs, thinly coriaceous, bilocular. Fl. andfr. June. Sand hills.— Centr. Asia: Balkh. (in Zaisan depression, on the Irtysh River above the mouth of Kurchum River and on the Chernyi Irtysh River). Gen. distr.: Dzu.-Kashg. (the Chernyi Irtysh). Described from the sands of Irtysh, above mouth of the Kurchum. Type in Leningrad. | 752. A. cognatus C.A.M. in Schrenk, Enum. Pl. nov. I (1841) 81; { Edbwwiieihoss. 1.611: Boe. Astrap. turk. 295). } Perennial, suffrutescent, 35-50 cm tall; caudices obsolescent, rarely attaining 3 cm in length; branches of preceding years absent, or ligneous upto 10cm long; branches of the year many, prostrate, 20-40 cm long; stipules almost fully connate, adnate at base or in lower part to petiole, 2-3 mm long; leaves 4-10 cm long, the petiole (3) 4-9 cm long, densely covered with appressed white hairs; leaflets 3 (4) pairs, oval to oblong- oval, round-tipped, rarely subacute, (3) 5-15 (20) mm long, 3-8 (12) mm broad, rather densely appressed-hairy on both sides; peduncles half as long as to equaling the leaves, covered with short appressed white hairs; _ together with inflorescence 5-11 cm long; racemes loose, remotely few- flowered, 4-10 cm long; bracts ovate to oblong-ovate, 1-2 mm long, | densely covered with appressed white hairs; calyx cylindric-campanulate, 6-8 mm long, densely covered with appressed white hairs, the tube 3-5 times the length of the linear-subulate teeth; corolla pale violet; standard 13-15 mm long, the limb rounded-ovate, slightly emarginate, as long as the rather narrow claw; wing-petals 11-13 mm long, the limb oblong, obtuse, Somewhat shorter than to as long as the claw; keel 10-12 mm long; ovary sessile, villous; pods sessile, ovaloid to oblong-ovaloid, 6-8 mm long, 3.5-4 mm broad, obtuse, terminating in a short subulate beak not exceeding 1 mm, inflated, carinate ventrally, rounded or somewhat concave dorsally, thinly coriaceous, densely villosulous with spreading white hairs, bilocular. Fl. June-July (August); fr. July-August. Sands.— Centr. Asia: Balkh. (Balkhash sand dunes to medium course | of River Ili, Muyun-kumy). Gen. distr. : Kulja. Described from sands of the Balkhash region. Type in Leningrad. | 753. A. karakugensis Bge. in Arb. Naturf. Ver. Riga I (1848) 232; ‘71 Bge. Astrag. geront. II, 226; Bge. Astrag. turk. 291. Perennial, suffrutescent, strongly branched; caudex not more than 7 cm long, often obsolescent, not infrequently largely buried; branches of preceding year 6-30 cm long, densely covered with short appressed hairs, grayish-white branches of the year 7-25 cm long, slender, subsulcate; stipules adnate to petiole in lower part (the lower ones up to the middle), connate, 4-5 mm long, triangular-lanceolate, acute; leaves on spurs 2-3 cm, on vegetative shoots 7-10 cm long, the petiole together with rachis ca. 1cm long on spurs and 5-8 cm long on vegetative shoots, densely covered with appressed white hairs; leaflets 2 (3) pairs, rarely (mostly on shoots) 1 pair, linear, acute, (1.3) 2-4 cm long, silvery-white on both sides by a dense coat of short appressed hairs; peduncles 2-4 cm (on shoots O77 772 0.4-1) cm long, covered with appressed white hairs, together with inflorescence 5-20 cm long (on shoots ca. 2 cm); racemes loose, 5-15 cm remotely few-flowered; bracts rounded-ovate, ca. 1 mm long, somewhat exceeded by the pedicel, covered with white hairs; calyx campanulate- cylindric, 4-5 mm long, densely covered with short appressed white hairs, the tube 4-5 times the length of the linear~subulate teeth; standard 9-11 mm long; wing-petals 8-10 mm long, the limb linear-oblong, obtuse, as long as the claw; keel 7-9 mm long, the limb obtuse, the claw 1.5 times as long as the limb; ovary sessile, hairy; pods sessile, ovaloid, 5-6 mm long, 3-3.5 mm broad, inflated, not grooved, slightly carinate ventrally, rounded dorsally, obtusish, terminating in a subulate beak 1 (1.5) mm long, densely villosulous with soft white hairs, thinly coriaceous, almost fully bilocular. Fl. April-July; fr. May-July. Partly consolidated sand dunes and sand hills. Centr. Asia: Ar.-Casp. (from Taisugan sands and flatlands of Emba River to Aral Kara-kum, northwest extremity of Muyun-kum sands Balkh. (Mugyun-kumi, sand hills of Ili River along northern border of Tien Shan to foothills of Dzhungarian Ala Tau). Endemic. Described from Irgiz, vicinity of Lake Kara-kug and Kara kum sands. Type in Paris. Series 12. ARBORESCENTES M. Pop.— Fairly large shrubs, with well-developed caudices; upper leaves trifoliate or consisting of single leaflets; petioles not spinescent, up to 12 cm long; leaflets glabrous above; flowers 12-15 mm long; racemes short and fairly loose or strongly elongated and remotely flowered; calyx covered with white hairs or glabrate; pods sessile or subsessile, ovoid or oblong, villous. 754. A. paucijugus C.A.M. in Bull. Phys. Math. Ac. Petrop. 2 (1844) 2; Bge. Astrag. geront. II, 225; Bge. Astrag. turk. 294. — A. arborescens Bge. in Arb. Naturf. Ver. Riga I (1848) 229. — Exs. : H.F.R. No. 661. A large shrub with woody caudices; growth branches of the preceding year ligneous, 25-55 cm long, stout, slightly angled and sulcate,, densely covered with appressed white hairs; older branches stout, ribbed, gray; branches of the current year 2-7 cm, vegetative ones 25 cm long; stipules almost fully connate at base into a sheath, adnate to petiole, 1.5-2 mm long; leaves (3) 4-17 cm long, the petiole persistent of preceding year's extension branches, spreading, together with rachis 5-15 cm long, 1-1.5 (4) cm in brachyblastic specimens, covered with short appressed white hairs; leaflets 2-3 pairs (1-2 pairs on axillary branches), oval to oval- oblong or obovate, 1.5-3 cm long, 6-12 mm broad, obtuse, glabrous above, rather sparsely appressed-hairy beneath, greenish; peduncles on axillary branches 1.5-4 (5) cm long, covered with short appressed white hairs; racemes short, rather compactly 4-10-flowered, ovoid, 2.5-3 cm long; bracts ovate, 1-1.5 mm long, densely white-ciliolate; calyx campanulate, 4-5 mm long, glabrate except for appressed white hairs toward apex chiefly on the teeth, the triangular teeth ca. 1 mm long; corolla lilac (?); standard 12-15 mm long, the limb broadly ovate, slightly emarginate, 3 times as long as the claw; wing-petals 11-14 mm long, the limb linear- oblong, round-tipped, 1.5 times as long as the claw; keel 8-9 mm long; ovary on a very short stipe not exceeding 0.5-0.7 mm, white-hairy; pods subsessile, gathered in a compact raceme, ovoid, 8-12 mm long, mucronulate or obtuse, slightly carinate ventrally, convex dorsally, 578 773 somewhat compressed laterally, thinly coriaceous, villous with long spreading white hairs, bilocular. Fl. April-May; fr. May. (Plate XLV, Figure 3). On sandhills often in Haloxylon associations. — Centr. Asia: Kyz. K. (S.W. Kyzyl Kum), Balkh. Endemic. Described from lake Balkhash. Type in Leningrad. 755. A. longipetiolatus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, 50-80 cm tall; caudices not exceeding 15 cm, covered with grayish-brown longitudinally splitting bark; branches of the preceding year 15-30 cm long, hoary by a dense coat of appressed white hairs; branches of the current year (7) 10-20 cm long; stipules adnate at base to petiole and connate to high up, 3-4 mm long; upper leaves 3-5 cm, the lower ones 7-11 cm long, the petiole 0.5-2 and 4-9 cm long, respectively, covered with appressed white hairs or glabrous; leaflets 1 and 5-7 on upper and lower leaves respectively, elliptic to lanceolate or narrowly linear, 10-30 mm long, 3-12 mm broad, acute or obtuse, glabrous above, covered beneath with appressed white hairs; peduncles equaling the leaves, together with inflorescence (15) 20-27 cm long or rarely inflorescence subsessile; racemes loose, remotely many-flowered, (12) 18-25 cm long; bracts lanceolate, 2-3 mm long, glabrate or white-ciliate; calyx cylindric-campanulate, 8-9 mm long, covered with appressed white hairs or glabrate, the linear-subulate teeth half the length of the tube; corolla pale violet (?); standard 12-13 mm long, the limb rounded-oval, scarcely emarginate, as long as the broad claw; wing-petals 11-12 mm long, the limb oblong, rounded-obtuse, as long as the claw; keel 9-10 mm long; ovary sessile; pods sessile, 6-7 mm long, oblong, 2-5 mm broad, acute, terminating in astraightbeak ca. 1 mm long, inflated, not grooved, thinly coriaceous, densely villous with spreading white hairs, bilocular. Fl. April-May; fr. (May) June-July. More or less consolidated sands.— Centr. Asia: Kara K. (E.), Mtn. Turkm. (Badkhyz, foothills of Kopet Dagh plain). Endemic. Described from E. Turkmenia. Type in Tashkent. Series 13. MICRANTHI M. Pop.— Petioles not spinescent, very short; upper leaves consisting of a single leaflet; leaflets hairy above; flowers 8-10 mm long; racemes loose, elongated, the flowers very distant; calyx covered with white hairs; pods sessile, oblong-ovoid, villous. 756. A. unifoliatus Bge. in Arb. Naturf. Ver. Riga I (1848) 231; Bge. Astrag. geront. II, 226; Bge. Astrag. turk., 291; Boiss. Fl. or. II, 487.— A. confirmans Freyn in Mém. Herb. Boiss. No. 13 (1900) 17. Perennial, suffrutescent; caudex short, woody; shoots of the current year prostrate, 13-22 cm long; stipules adnate to petiole to high up and connate at base, 2-2.5 mm long, the free portion broadly short-triangular ; leaves 2-6 cm long, the upper and lower ones composed of 1 and 3-5 leaflets, respectively, the petiole rigid, persistent but not spinescent, 0.5-1.5 (2.5) cm long; leaflets elliptic to lanceolate, subacute, 1-5 cm long, 3-7 mm broad, hoary on both sides by a dense coat of short appressed hairs; peduncles 1.5-3 times the length of the leaves, together with inflorescence (7) 13-30 cm long, white by a dense coat of appressed hairs; racemes (6) 10-22 cm long, many-flowered; flowers distant, obliquely 579 774 775 ascending, finally nodding; bracts lanceolate, 1.5-2 cm long, somewhat exceeding the pedicel; calyx campanulate, 3.5-4.5 mm long, densely covered with short appressed white hairs; the tube 3-4times the length of the lance-linear teeth; corolla red (?); standard 8-10 mm long, the limb rounded-oval, scarcely emarginate, about equaling to 1.3 times as long as the very broad claw; wing-petals 8-8.5 mm long, the limb oval-oblong, round-tipped, as long as the claw; keel 7-7.5 mm long; ovary sessile, white-hairy; pods sessile, oblong-ovoid, 5-6 mm long, terminating ina straight beak ca. 1 mm long, convex dorsally and ventrally, villous with soft spreading white hairs, thinly coriaceous, nearly bilocular, 1-seeded. Fl. May; fr. May (June). Sands.— Centr. Asia: Kara K., Kyz. K. (Karakalpak Kyzyl-kum, southern Kyzyl-kum), Amu D. (sands of Sundukli sandhills on right bank of Amu-Daryariver). Endemic. Described from the sands of Kyzyl-kum. Type in Paris. Series 14. VILLOSISSIMI M. Pop.— Petioles not spinescent or more or less indurescent and persistent as spines, but not subulately acuminate, up to 2.5 cm long; flowers 11-15 mm long; racemes short, rather compact, the flowers subapproximate, the pods more closely set; calyx covered with white hairs; pods sessile, ovaloid-oblong or oblong-ovoid, villous. 757. A. brachypus Schrenk, Enum. Pl. Nov. I (1841) 79; Bge. Astrag. geront: 1l;-22”7> “Astrag- turk: 2927) "Uidbe "hl. Hoss: "610i, A. Holodendron Bge. Astrag. geront. (1868) 227; Astrag. turk, 293. Perennial, suffrutescent, 60-90 cm tall; caudices (10) 15-40 (50) cm long, stout, angled, covered with grayish-brown longitudinally splitting bark; branches of the preceding year 10-30 cm long, in brachyblastic specimens 3-5 cm, those of the current year 10-40 cm and 5-7 cm long, respectively; stipules almost fully adnate to petiole and connate, 2-3 mm long; leaves (1.5) 2-4 cm long, the petiole together with rachis 2-6 mm long, covered with short appressed white hairs; leaflets 1 pair, the lower ones sometimes 2 pairs, linear, 15-35 (50) mm long, often 10-15 mm in brachyblastic specimens, (1.5) 2-3 (4) mm broad, subobtuse, hoary on both sides by a dense coat of appressed hairs; peduncles equaling or slightly exceeding the leaves, together with inflorescence 5-10 (17) cm long; racemes loose but rather closely 6-many-flowered, 3-8 (13) cmlong; bracts lanceolate, 1-2 mm long, equaling the pedicel; faintly white-hairy; calyx cylindric-campanulate, 6-7 (8) mm long, densely covered with short ascending hairs, the tube 4-5 times the length of the lance-subulate teeth; corolla purple; standard 13-15 mm long, the limb broadly rounded-ovate, slightly emarginate, as long as the claw; wing-petals (10) 11-12 mm long, the limb oblong, obtuse, as long as the claw; keel (8) 10-12 mm long; ovary sessile, hairy; pods sessile, ovaloid-oblong, 5-7 (8) mm long, 3 (4) mm broad, carinate ventrally, rounded-convex dorsally, obtusish, terminating in a slightly curved subulate beak ca. 1 mm long, thinly coriaceous, densely villous with soft spreading white hairs, bilocular. Fl. May-June; fr. June-July. Sands, often sandhills and argillaceous-sandy sites, often in Haloxylon associations.—Centr. Asia: Ar.-Casp. (northwestern extremity of Muyun-kum, lower reaches of Sary-su River), Balkh. (Muyun-kum, Balkhash, Ili river valley). Endemic. Described from Balkhash sands. Type in Leningrad. 837, 1117 580 776 758. A. villosissimus Bge. Astrag. geront. II (1869) 347; Bge. Astrag. turk. 294. A branched shrub up to 70 cm tall, vested with white hairs; ligneous branches elongated, stout, covered with fibrillose bark; stems of the year very short (brachyblasts) or elongated, (dolichoblasts), ranging from 6 to 23 cm in length; stipules adnate to petiole and connate up to 1/22/3107. almost fully, 3-3.5 mm long, the free portion triangular, subacuminate; leaves 1.5-3 cm long, the petiole 0.4-2 (2.5) cm long, not spinescent; leaflets 2 (3) pairs, the upper ones 1 pair, oblong to linear~lanceolate, 6-15 mm long, 2-3 (3.5) mm broad, the terminal shorter than. the lateral ones, subobtuse, scabrous and grayish on both sides by appressed hairs; peduncles mostly equaling the leaves, together with inflorescence 2.5-8 cm long; racemes 2-6 cm long, loose (2) 3-8-flowered; bracts ovate- lanceolate, 1-2 mm long, equaling the pedicel, faintly covered with short hairs, yellowish, glabrous at base; calyx 5-6 mm long, covered with subappressed white hairs, the tube 3-4 times the length of the lance- subulate teeth; corolla reddish; standard 11-12 mm long, the limb rounded- ovate, scarcely emarginate, as long as the claw; wing-petals 10-11 mm long, the limb oblong, rounded-obtuse; keel 9-10 mm long; ovary borne on a very short stipe, white-hairy; pods rupturing the calyx, sessile, oblong-ovoid, 7-9 mm long, acutish, terminating in a curved beak 1.5-2mm long, slightly carinate dorsally and ventrally, rigidly membranous, densely villous with soft spreading white hairs, bilocular. Fl. April-May; fr. May. Sands, often in Haloxylon associations.— Centr. Asia: Kyz. K. (Karakalpak and south Kyzyl-kum), Amu D. (sands of Sundukli on right bank of Amu-Darya River). Endemic. Described from Kyzyl-kum. Type in Leningrad. 759. A. chodshenticus B. Fedtsch. in Tr. Tadzhik. bazy Ak. Nauk SSSR II (1936) 153; Fl. Tadzhik. V, 453. A strongly branched shrub, the stout caudices and their divisions covered with pale brownish-gray longitudinally splitting bark; woody old dolichoblasts and branches of the current year slightly flexuous; stipules connate in lower part, 1-2 mm long, the free portion triangular; leaves 1-3.5 cm long, the rachis covered with appressed white hairs, becoming lignified after the shedding of leaflets and transformed into rigid subarcuately recurved spinelike structures (1) 1.5-2.5 cm long; leaflets 2 pairs, lanceolate to elliptic, 8-15 mm long, (2) 3-7 mm broad, round- tipped, rigid, appressed-hairy on both sides, more densely so beneath; peduncles 1-2 cm long, covered with short appressed white hairs; racemes 1-2 cm long, at first dense, finally rather loose but closely fruited; bracts lance-ovate, 1-1.5 mm long, covered with short white hairs, calyx cylindric-campanulate, ca. 7 mm long, covered with appressed white hairs, the tube 4-5 times the length of the subulate teeth; corolla violet; standard ca. 1.5 mmlong, the limb oblong-elliptic, round-tipped, gradually narrowed into the claw; wing-petals somewhat exceeded by the standard, the claw 1.5 times the length of the oblong limb; keel shorter than the wings; pods borne ona stipe 1-2 mm long, ovoid with hairs, oblong without them, 8-10 mm long, terminating in a beak 1.5-2 mm long, membranous, densely villous with spreading tubercle-based white hairs, slightly carinate ventrally, rounded dorsally, unilocular. Fl. April; fr. May. Sands. — Centr. Asia: Syr D. (from Begovat village and upward). Endemic. Described from the vicinity of Leninabad. Type in Leningrad. 581 777 Series 15. LAXIFLORI Gontsch.— Petioles not spinescent, short; leaflets 1-2 pairs; flowers 8-14 mm long; racemes loose, elongated, remotely flowered; calyx covered with white hairs; pods sessile, ovaloid or oblong-ovaloid or oblong-ovoid, villous. 760. A. surchanensis M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, suffrutescent, up to 70 cm tall; caudex stout, covered with grayish longitudinally splitting bark, the fairly stout divisions up to 6 cm long; younger lignified branches short or fairly long, tightly angled, cinereous, densely hairy; floriferous branches of the current year 12-20 cm long, terete; stipules almost fully adnate to petiole and connate, ca. 2 mm long, short-bidentate; leaves 3.5-6 cm long, the petiole densely covered with appressed white hairs, persistent, obtuse, those of lower and upper leaves 0.5-2 and 1-2 cm long, respectively; leaflets of lower and middle leaves 2 pairs, those of upper leaves 1 pair, lance-linear to linear, covered on both sides, sparsely so above, with appressed white hairs, hoary, 1-4 cm long, 1-3 mmbroad, acute or rarely subobtuse, exceeding the leaves; peduncles 7-10 cm long, densely covered with appressed white hairs; racemes loosely 10-24-flowered, 7-15 cm long; bracts ovate- lanceolate, ca. 2 mm long, sparsely white-ciliolate; calyx campanulate, 5-7 mm long, covered with appressed white hairs, the teeth lanceolate, acuminate, 1.5-2 mm long; corolla purple; standard 13-14 mm long, the limb rounded-oval, round-tipped, 3 times the length of the claw; wing- petals 11-12 mm long, the limb linear-oblong, as long as the claw; keel ca. 10 mm long; ovary on a stipe ca. 1 mm long, white-hairy; pods borne on a stipe ca. 1 mm long, 7-8 mm long, acute, densely covered with long spreading hairs. Fl. April; fr. May. Sands.— Centr. Asia: Pam.-Al. (Shirabad valley, Amu-Darya)river valley above Termez, below Kafirnigan). Endemic. Described from Bunitau. Type in Leningrad. 761. A. macrobotrys Bge. Astrag. geront. II (1869) 226; Boiss. Fl. or. II, 487.—Solenotus gypsaceus Nevski in Acta Inst. Bot. Acad. Sc. URSS. ser.II, fasc. IV (1937) 352. A shrub up to 70 cm tall; caudex branched, up to 20 cm long; branches of the current year 15-25 cm long; stipules almost fully adnate to petiole and connate, 2-3 mm long, the free portion triangular; leaves 1.5-7 cm long, the petiole half as long as to equaling the axis, both petiole and rachis firm, sparsely covered with appressed white hairs, partially persistent; but not spinescent; leaflets of lower leaves 2-3 pairs, oblong-obovate or obovate-oblong, round-tipped, cuneate at base, 10-17 mm long, covered sparsely above and densely beneath with appressed hairs, or glabrous above, rather thick-textured; peduncles equaling or slightly exceeding the leaves, covered with short appressed white hairs; racemes 7-15 cm long, the distant flowers obliquely ascending; bracts ovate-lanceolate, 1.5-2 mm long, somewhat shorter than to equaling the pedicel, white-ciliate; calyx campanulate, 5-7 mm long, covered with short appressed white hairs, the tube 3 times the length of the linear~ subulate teeth; corolla reddish; standard 12-13 mm long, the limb oval or rounded-oval, scarcely emarginate, as long as the claw; wing-petals 11.5-12 mm long, the limb oblong, slightly enlarged toward apex, entire or scarcely emarginate, as long as the claw; keel 11 mm long; ovary subsessile, white-hairy; pods 582 778 subsessile, oblong-ovaloid, ca. 7 mm long, carinate ventrally, rounded or subapplanate dorsally, rigidly membranous, densely villous with soft spreading white hairs, bilocular, 2-seeded. Fl. April-May (June); fr. May- June. ; Sands in the desert region, more rarely clay-and-gravel-covered foothills. — Centr. Asia: Kara K. (eastern part), Mtn. Turkm., Pam.-Al. (Kugitang Range). Gen. distr.: Iran. Described from E. Iran. Type in Leningrad. 762. A. macrocladus Bge. Astrag. turk. (1880) 293. Perennial, suffrutescent, 50-65 cm tall; caudices mostly 5-8 (20) cm long, covered with grayish-brown longitudinally splitting bark; ligneous branches of the preceding year 10-20 cm long (3-5 cm in brachyblastic forms), gray by a dense coat of appressed hairs; branches of the current year 5-15 cm long; stipules adnate in lower part to petiole and connate at base, 2-2.5 mm long; leaves 3-6 cm long, the petiole together with rachis (1.5) 2-4 (5) cm long, commonly overtopping the terminal leaflets; leaflets 2 pairs, linear-oblong to lance-oblong, 10-20 (40) mm long, 3-3.5 (7) mm broad, covered sparsely on both sides or rather densely beneath, with appressed hairs; peduncles mostly equaling the leaves, covered with appressed white hairs, together with inflorescence (10) 12-20 cm long; racemes (8) 10-15 cm long, many-flowered, the flowers distant; bracts lanceolate, 2mm long, sparsely white-ciliate; calyx cylindric-campanulate, 5-6 mm long, densely covered with appressed white hairs, the tube 3-4 times the length of the lance-subulate teeth; corolla purple (?); standard 12-13 mm long, the limb rounded-oval, scarcely emarginate, as long as the broad claw; wing~petals 11-12 mm long, the limb oblong, slightly enlarged toward apex, as long as the claw; keel 9-10 mm long; ovary borne on a very short stipe, villous; pods sessile, oblong, acutish, ca. 7 mm long, ca. 9mm broad, carinate ventrally, rounded dorsally, rigidly membranous, densely villous with spreading white hairs, bilocular. Fl. April-May; fr. May. Sand dunes or consolidated and more or less flat sands, rarely in gritty loams.— Centr. Asia: Kyz. Kum (W.), Syr D. (the part of Syr-Darya river valley adjacent toKyzylKum). Endemic. Described from W. Kyzyl Kum (near Chardary). Type in Leningrad. 763. A. iliensis Bge. in Bull. Soc. Nat. Mosc. XXXIX, 2 (1866) 20; Bge. Astrag. geront. II, 227; Bge. Astrag. turk. 292. Perennial of medium height; caudices fairly stout, covered withgrayish- brown bark; ligneous branches of the preceding year terete, grayish-white by a dense coat of appressed hairs; shoots of the current year 7-16 cm long; stipules connate into a sheath nearly all the way up, short-bidentate, 2-3 mm long; leaves 3-5 cm long, the petiole together with rachis 5-11 cm long, covered with short appressed white hairs; leaflets 1-2 pairs, linear, 2-3 cm long, 2-3 (4) mm broad, subobtuse, densely covered on both sides with short appressed hairs; peduncles half as long as to equaling the leaves, densely covered with appressed white hairs; racemes (5) 6-12 cm long, many-flowered, the flowers very distant; bracts lanceolate, 1.5-2 mm long, covered with short white hairs, 1.5-2 times as long as the pedicel; calyx campanulate, ca. 4mm long, densely covered with short appressed hairs, the tube 2-3 times the length of the lanceolate, acuminate teeth; corolla purple; standard 8-9 mm long, the limb ovate, scarcelyemarginate, 583 779 780 the broad claw 1.5-2 times as long as the limb; wing-petals 7-8 mm long, the limb oblong, round-tipped, as long as the claw; keel 6-7 mm long; pods sessile, oblong-ovoid, 4-5 mm long, slightly compressed laterally, not grooved, acuminate intoa beak ca. 1 mm long, rigidly membranous, densely villous with soft spreading white hairs, almost fully bilocular, mostly 1-seeded. Fl. April-May; fr. May-June. Sand dunes.— Centr. Asia: Balkh. (Ili river valley). Gen.. distr. : Kulja. Described from Ili river valley. Type in Leningrad. 764. A. aiwadzhi B. Fedtsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS VIII, f.10 (1940) 166. Perennials, suffrutescent, 55-65 cm tall; caudices 15-25 cm long, stout, covered with brownish~gray longitudinally splitting bark; ligneous branches of the preceding year up to 25 cm long, grayish, densely covered with appressed white hairs; branches of the current year 20-55 cm long; stipules almost fully connate, 2-2.5 mm long, bidentate; leaves 3.5-5 cm long, the petiole together with rachis 1.5-4 cm long, rather densely covered with short appressed white hairs, indurescent and persistent; leaflets of lower leaves 1 pair, linear, 1.5-3 cm long, 2-3 mm broad, acute, densely covered on both sides with appressed white hairs; peduncles 1.5-3 cm long, equaling or rarely exceeding the leaves, up to 7 cm long, covered with appressed white hairs, together with inflorescence 13-30 (35) cm long; racemes loose, many-flowered, 10-20 mm long, the flowers very distant; bracts lanceolate, ca. 2 mm long, white-ciliolate, equaling or slightly exceeding the pedicel; calyx cylindric-campanulate, 6 mm long, densely covered with appressed white hairs, the tube 3-4 times the length of the lance-linear teeth; corolla reddish (?); standard 11-12 mm long, the limb rounded-oval, scarcely emarginate, as long as the very broad claw; wing-petals 10-11 mm long, the limb oblong, scarcely emarginate, somewhat shorter than the claw; keel 9-10 mm long; ovary sessile, densely covered with white hairs; pods sessile, oblong-ovaloid, 5-6 mm long, terminating in a beak ca. 1 mm long, not grooved, rigidly membranous, partially bilocular, densely villous with soft white hairs. Fl. May-June; fr. June-July. Consolidated sands in submontane hills; also sands and pebbles of riverside terraces.— Centr. Asia: Mtn. Turkm. (Badkhyz), Pam.-Al. (S.W.). Endemic. Described from vicinity of Aibadzha in lower reaches of Kafirnigan River. Type in Leningrad. Series 16. SPARTIOIDEI Gontsch.— Subshrubs with very short caudices; calyx campanulate, white-hairy; limb of standard rounded-oval; pods sessile, inequilaterally oblong, densely covered with sho1t appressed hairs. 765. A. spartioides Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 330; Bge. Astrag. geront. I, 125; II, 219, p.p. (excl. syn. A. psilopus C.A.M.); Bge. Astrag. turk. 285, p.p.— Exs.: H.F.R. No. 1409. Perennial, suffrutescent, 25-40 cm tall; caudex stout, 1.5-3 cm long, covered with castaneous-brown longitudinally splitting bark, the divisions short; ligneous branches of the preceding year terete, whitish-gray by a dense coat of white tomentum, 4-7 cm long; stems of the current year many, simple, erect, 10-20 cm long, white by a dense coat of appressed hairs; stipules adnate to petiole up to 1/2-1/3, not connate, ovate-triangular, acuminate, 2-3 mm long, densely covered with appressed white hairs; 584 leaves 4-10 (11) cm long, the petiole 1.5-2 times as long as the rachis, both petiole and rachis slender, covered with appressed white hairs; leaflets 2-3 pairs, narrowly linear, subobtuse, (12) 15-30 mm long, 1-1.5 mm broad, covered on both sides with short appressed white hairs; peduncles equaling or somewhat exceeding the leaves, slender, covered with appressed white hairs; racemes 10-15 cm long, remotely 7-16-flowered; bracts oblongly triangular-ovate, 1-1.5 mm long, covered with very short white hairs; calyx campanulate, 5 mm long, covered with short appressed white hairs, the subulate-lanceolate teeth 0.8-1 mm long; corolla purple (?); standard 10-11 mm long, the limb rounded-oval, scarcely emarginate, angularly attenuate at base, 4-5 times as long as the claw; wing-petals 9.5-10.5 mm long, the limb oblong, entire, obtuse, somewhat longer than the claw; keel 8 mm long, the limb semiorbicular, acutish, as long as the claw; ovary sessile; pods sessile, cernuous, inequilaterally oblong, ca. 10 mm long, ca. 2 mm broad, compressed laterally, carinate and convex ventrally, rounded dorsally, straight, terminating in a subulate beak 1 mm long, coriaceous, densely covered with short appressed white hairs. Fl. April-May. Sands.— Centr. Asia: Balkh. (between source of the Chingil'dy and the Leps and along the Leps River). Endemic. Described from the areabetween the sources of Chingil'dy and Leps. Type in Leningrad. Section 95. BULIMIOIDES Bge. Astrag. geront. I (1868) 130.— Strongly branched subspherical spiny shrubs, vested with bicuspidate appressed white hairs; branches densely studded with long spines formed by old leaf petioles; stipules adnate to petiole to high up, not connate; leaf petioles and rachises spinescent, persistent as spines on the old branches; flowers 1-2 inthe leaf axils, short-pediceled; bracteoles none; calyx cylindric-campanulate; corolla small, pink; pods cernuous, subsessile, linear-oblong or rarely lance-oblong, small, triangular, carinate ventrally, flat dorsally, bilocular or nearly so, few-seeded. An ancient monotypic section. Deserts in the northern part of Central Asia. 766. A. unijugus Bge. Astrag. geront. II (1868) 228; Bge. Astrag. turk. 297.— A. oligophyllus Schrenk in Bull. PhysMath. Acad. Petersb. Il (1844) 197, non Boiss. (1843); Fisch. in Bull. Soc. Nat. Mosc. XXVI (1853) 420. A small shrub, (11) 15-23 (32) cm tall, spherically pulvinate, profusely short-branched; shoots of the year short, densely clothed in leaf petioles, white by a dense coat of appressed hairs; stipules adnate to petiole up to 1/2-2/3, 3-4mm long, covered with appressed white hairs, the free portion narrowly lanceolate or linear from a broadly triangular base; leaves (3) 4-6 cm iong, the petiole rigid, spinescent, sparsely covered with short appressed white hairs, persistent on the branches as firm spines; leaflets 1 pair inserted on the upper half of the petiole and an early deciduous terminal leaflet, narrowly linear, acute, sparsely covered with appressed white hairs; pedicels 1-1.5 mm leng, densely white-hairy; bracts lanceolate, 1.5-2 mm long, white hairy; calyx 7-8 mm long, densely covered with appressed white hairs, the linear -filiform teeth 2 mm long; standard 10-12 mm long, the limb obovate, rarely oblong-obovate, round-tipped and slightly emarginate, 4 times as long as the broad claw; wing-petals equaling the standard, the limb oblong, scarcely emarginate, as long as the claw; 585 782 keel 9-10 mm long, the limb obtuse, the claw 1.5 times as long as the limb; ovary borne on a stipe ca. 0.5 mm long, faintly covered with short white hairs; pods 6-9 mm long, 2-2.5 mm broad, terminating in a straight beak 0.5-1 mm long, at first glabrate, finally quite glabrous, cross-nervose. Fl. May; fr. June (July). Clayey and sandy deserts and takyrs; also gritty or sandy loams. — Centr. Asia: Ar.-Casp. (N.E.), Balkh. Endemic. Described from vicinity of lake Balkhash and from Golodnoi steppe north of Chu River. Type in Leningrad. Subgenus VIII. EPIGLOTTIS Boiss. Fl. or II (1872) 205.— Annuals vested with bicuspidate hairs attached at their middle. Section 96. BUCERAS DC. Prod. II (1825) 290, pro parte; Bge. Astrag. geront. I (1869) 12.— Annuals, vested with appressed bicuspidate white or mixed white and black hairs; stipules adnate to petiole and connate at base; leaves imparipinnate, the petioles marcescent; racemes dense, subcapitate, somewhat elongating after flowering, borne on axillary peduncles; bracteoles none; calyx campanulate; wing-petals shorter than the standard and longer than the keel; pods sessile, linear, hairy, more or less curved, grooved dorsally, cartilaginous-coriaceous, bilocular, many~-seeded. A Mediterranean and Near Asian section, distributed through the Canary Islands, South Europe, North Africa, Syria, Arabia, Mesopotamia, Armenia, Iran, W. Kopet Dagh, Caucasus, Crimea, and the lower reaches of the Volga. 767. A. hamosus L. Sp. pl. (1753) 758; DC. Astrag. 155; Bge. Astrag. geront. If} 13; Boiss. El’-or. Il, 2385 ‘Grossh. &l” Kavko I 20a A. brachyceras Ldb. Ind. Hort. Dorpat. (1822) 3; Bge. Astrag. geront. If "13 Boiss. Fl. or. Il, "238;"Grossh. Fl-"Kavk- ie" 20’08> A. hamosus £$ brachyceras Ldb. Fl. Ross. I (1842) 638. — Ankylotus hamosus Stev. in Bull. Soc. Nat. Mosc. XXIX (1856) 148. Annual, (5) 10-55 cm tall; stems erect or ascending or prostrate; (5) 10-45 cm long, appressed-hairy; stipules connate and adnate to petiole at base, 4-9 mm long, white-ciliate on the margin, the free portion triangular- ovate, acuminate; leaves (3) 5-12-cm long, the very short petiole and the rachis covered with short appressed hairs; leaflets (5) 7-10 pairs, oblong - to oblong-obovate, retuse, (3) 5-15 mm long, glabrous above, appressed- _ hairy beneath; peduncles half as long as to equaling the leaves, covered with appressed white hairs; racemes densely capitate, 5-10-flowered, loose in fruit, 2-3 cm long; bracts linear-subulate, somewhat exceeding the calyx-tube but surpassed by the teeth, 3-4 mm long, white-hairy, 4.5-5 mm long; calyx campanulate, covered with white and black or pure black or rarely pure white hairs, the subulate teeth as long as the tube; corolla pale violet or nearly white; standard 7.5-8 mm long, the limb ovate, scarcely emarginate, 3-4 times as long as the claw; wing-petals 5-6 mm long, the limb linear-oblong, twice the length of the claw; keel 4.5-5 mm long, the limb broad, obtuse, as long as the claw; pods sessile, linear, 2-4 cm long, 2.5-3 mm broad, subterete, scarcely canaliculate dorsally, pendulous and arcuately upturned, the upper geniculation longer than the 586 783 lower one, acuminate, covered with scattered appressed hairs, almost indehiscent, bilocular, many-seeded. Fl. April-May; fr. May-June. Dry stony mountain slopes, in steppes and sands. — European part: L.V. (Astrakhan), Crimea; Caucasus: Cisc., Daves) andiSualranseafaa als y Centr. Asia: Mtn. Turkm. (W. Kopet Dagh). Gen. distr.: Med., Bal.-As. Min. (Balkans), Arm.-Kurd., Iran (Iran). Described from the South of France. Type in London. Subgenus IX. CALYCOCYSTIS Bge. Astrag. geront. I (1868) 133.— Rather low shrubs, or acaulescent or more or less caulescent perennials, vested with bicuspidate hairs; racemes loose or more often compactly capitate, borne on rather long peduncles; calyx at first cylindric, becoming strongly bladdery-inflated or rarely slightly inflated; corolla purple, pink, violet, or yellow: pods sessile or nearly so, mostly small, included in the bladdery calyx or rarely rupturing the slightly inflated calyx and in that case equaling or slightly exceeding it, ovaloid or ovoid or lanceolate or oblong, coriaceous, bilocular, rarely incompletely so or unilocular. Section 97. CYSTODES Bge. Astrag. geront.I (1868) 133.— Subshrubs with short caudices; stems of the year developed; stipules not connate, adnate at base to petiole; inflorescence rather loose, short, spherical or ovoid, long-peduncled; bracteoles none; calyx at first cylindric, at length oblong-ovaloid inflated, finally ruptured by the pod; corolla pale yellow or violet; pods sessile or nearly so, rupturing the pod, erect, oblong, carinate ventrally, grooved dorsally, compressed laterally, rigidly coriaceous, densely covered with stiff spreading hairs, bilocular, rarely semibilocular. A section widely distributed through Spain, South of France and Austria, to the foothills of southern Altai; within the confines of the USSR mostly in the steppe, semidesert, and forest-steppe zones. Xerophytes associated chiefly with native strata, limestone and chalk; less frequently in zonal steppe associations. 1. Bracts ovate, acuminate (steppes and semideserts of Kazakhstan). Pere Ta.” One. Sete. om, Beha: Gabe, Ae 768. A. medius Schrenk. eo wacissianceolateyor linearglanceolaters: Qh). oi. «ice -. tem) cn) oat 25 Leaflets (5) 6-8 pairs; standard violet or rarely corolla whitish; stems 1-5 cm long; inflorescence spherical or subspherical (S. W. part of European USSR, to the west of Dnieper, Kharkov).............. POEM Vs Tyee oieeort a Seite Ma Sees ENED. ee. 770. A. vesicarius L. + Leaflets 3-4 (7) pairs; corolla pale yellow; stems 5-10 cm long; inflorescence ovoid (south of European SSSR, to the east of Dnieper, chiefly on the Volga and Don watershed, eastward to Orsk and Sauciward to. Giscaucasia)iat. ik fn. cacpes. lke: 769. A. albicaulis DC. is) 768. A. medius Schrenk in Bull. phys.-math. Acad. Pétersb. II (1844) 196; Bge. Astrag. geront. I, 134; II, 231; Bge. Astrag. turk. 308; Kryl. Fl. Zap. Sib. VII, 1703. — A. albicaulis Ldb. Fl. Ross. I (1842) 628; non DC. Perennial, suffrutescent, (18) 25-50 cm tall; stems of the preceding year ligneous at base, (5) 10-20 cm long, covered with yellowish-brown 587 bark; stems of the current year (2) 6-17 cm long, densely covered with appressed white hairs; stipules adnate at base to petiole, ovate, 2-3 mm long, rather sparsely white-hairy; leaves (1.5) 2.5-4 cm long, the rachis 2-4 times the length of the petiole, both petiole and rachis densely covered with appressed white hairs; leaflets 2-4 pairs, oblong-oval to oblong, obtuse, rarely linear-oblong, acuminate, (5) 9-20 mm long, appressed- hairy on both sides; peduncles 4-11 cm long, 2-2.5 times the length of the leaves, covered with appressed white hairs; inflorescence ovoid, oblong, 3-7 cm long, rather loosely 5-15 (20)-flowered, elongating in fruit up to 9-12 cm and becoming oblong; bracts ovate, acuminate, (2.5) 4-5 mm long, covered with white and black or almost pure black hairs; calyx at first cylindric, becoming oblong-ovaloid inflated in fruit, 12-14mm long, covered 784 with ascending or appressed white and, mainly on the teeth, some black hairs, the tube 3-4 times the length of the lanceolate teeth; corolla pale yellow, sometimes with a pale violet standard; standard 22-26 mm long, the limb oblong-obovate, auriculately angular-dilated in lower 1/3, 3-4 times as long as the claw; wing-petals 21-24 mm long, the limb oblong, more or less enlarged at apex, entire, the claw 1.5 times as long as the limb; keel 17-20 mm long, acutish; pods virtually sessile, erect, inequilaterally oblong, 10-18 mm long, 4-5 mm broad, terminating in a straight subulate beak 2-3 mm long, carinate ventrally, rather shallowly grooved dorsally, compressed laterally, coriaceous, densely covered with stiff spreading white hairs, semibilocular or incompletely bilocular. Fl. May; fr. June. Steppes, on chalk.— W. Siberia: U. Tob. (S.), Irt. (S.); Centr. Asia: Ar.-Casp. (N., southward to Emba), Balkh. (N., southward to Ayaguz). Described from Mt. Ulutau and from Kara-Kingir River and Ters-~Akkan. Type in Leningrad. Note. The form growing on chalk (var. cretaceus M. Pop.) differs somewhat from the steppe form. It is distinguished by the following characters: the plant whitish-gray by dense vesture; calyx covered with short spreading mainly white hairs; racemes loose; leaflets linear-oblong; bracts 2-3 mm long. 769. A. albicaulis DC. Astrag. (1802) 166; DC. Prodr. II, 289; Lab. Bl Rossi, \6283}, Bee: ,Astrag:)geront) 1) 134; e0l 23h) Boisisheil: sore stl 495; Schmalh. Fl. I, 285; Grossh. Fl. Kavk. II, 319.—A. dealbatus Pall. Astrag. (1800) 26, p.p. tab. 23, £.2; Willd. Spec. pl. III, 1263; M: By Fl. taux.-cauc. JI, 186 (exel) jsyninGmels)) jley) DO. WAsifageaiabr ZirS EVER ONO (Gil. Perennial, suffrutescent; branches of the preceding year ligneous at base, gray; stems of the current year 5-10 cm long, ascending or erect, slender, terete, white by a dense coat of short appressed white hairs; stipules 2-3 mm long, the lower ovate, the upper lanceolate, covered with appressed white or mixed white and black hairs; leaves 2-6 cm long, the petiole very short, slender; leaflets 3-4 (7) pairs, oblong or oval, acute, rarely obtuse, (7) 10-20 mm long, 3-5 (6) mm broad, hoary on both sides by appressed hairs; peduncles 10-15 cm long, hoary; racemes capitate, ovoid, rather loose, 2-10 cm long; bracts lance-linear, 3-5 mm long, covered with white or mixed white andblackhairs; calyxca.15mm, sometimes not more than 10 mm long covered with subappressed white and black hairs, the latter usually arrangedin stripes, the filiform teeth 3-5 mm long, the tube 588 2-3 times the length of the teeth, somewhat enlarged in fruit; corolla pale yellow; standard 18-22 mm long, the limb obovate, emarginate, auriculate- angular at base; wing-petals 17-21 mm long, the limb oblong, entire at 785 apex; keel acute, 13-17 mm long, the claw 1.5 times as long as the limb; pods sessile, erect, oblong, 10-15 mm long, 3-4 mm broad, carinate ventrally, rather shallowly grooved dorsally, obliquely cuspidate, coriaceous, densely pubescent-villous with spreading hairs, bilocular. F1. May-July (August); fr. June-August. On chalk, in chalky pine woods, rarely on limestone. — European part: V.-Don (S.), L. Don, L.V. (Astrakhan); Caucasus: Cisc. (Stavropol' plateau); W. Siberia: U. Tob. (Orsk, Guberlya Mountains). Endemic. Described from the Volga-Don watershed. Type in Paris. 770. A. vesicarius L. Sp. pl. (1753) 760; DC. Astrag. 91; Bge. Astrag. Benoni) tao; It, Zateschmalhiar). 1, 285, 8A. ‘ao libndus AW. etuk., Ic. pl. rar. Hung. I (1802) 39, tab. 40.— A. mixtus M.B. Cent. pl. rar. Ross. I (1810) tab. 26 et in Fl. taur.-cauc. III, 488.—Solenotus vesicarius Stev. in Bull. Soc. Nat. Mose. XXIX (1856) 144.— Exs.: Puno exS.. NO. oie Perennial, suffrutescent, (10) 15-35 cm tall; branches of the preceding year ligneous along 1-7 cm from base; stems of the current year erect or more often ascending, 1-5 cm long, white by a dense coat of short appressed hairs, simple or rarely branched; stipules lanceolate, acute, free, white-hairy, 2-3 mm long; leaves 4-6 cm long, short-petioled; leaflets (5) 6-9 (10) pairs, oblong obtuse or rarely elliptic acute, 7-12 mm long, 2-3 mm broad, canescent on both sides by a dense coat of appressed white hairs; peduncles (6) 10-20 cm long, 2-3 times the length of the leaves; inflorescence capitate, subspherical or rounded-ovoid, 3-5 cm long, rather loose, not elongating in fruit, 3-10-flowered; bracts lanceolate, 3-5 mm long, 2-3 times the length of the pedicel, covered with ascending black hairs; calyx cylindric, ca. 10 mm long in fruit, distinctly ovoid- enlarged and 11-12 mm long, subpannose with ascending white and black hairs, the tube 2-6 times the length of the triangular or subulate teeth; corolla violet, rarely whitish; standard 17-23 mm long, the limb oblong- obovate, emarginate, narrowed in lower 1/4, angular at base, 4 times as long as the claw; wing-petals 16-20 mm long, the limb linear-oblong, entire, somewhat shorter than the claw; keel 14-17 mm long, acute; pods rupturing the calyx, sessile, erect, oblong, 8-13 mm long, 5 mm broad, slightly carinate ventrally, rather shallowly grooved or rounded dorsally, obliquely cuspidate, coriaceous, covered with spreading white hairs, semibilocular. Fl. May-July; fr. July-August. A variety, var. albidus (W. et K. Beck), occurring in the USSR, has a violet standard and whitish wings and keel, or a uniformly whitish corolla (f. pallidiflorus M. Pop.). Fl. June-July; fr. July-August. Gullies, slopes of river valleys, and stony mountain slopes, mostly on 796 limestone outcrops. — European part: M. Dnp. (S. W.) V.-Don (vicinity of Khar'kov), Bes., Bl., (W.), Crim. Gen. distr.: Centr. Eur., W. Medit., Bal.-As. Min. (Balkans). Described from the South of France. Type in London. Section 98. MACROCYSTODES M. Pop. sect. nova in Addenda XI, 662. (incl. sect. Bucharica B. Fedtsch.).— Coarse perennials, vested with bicuspidate hairs always appressed on stems and leaves; 589 789 stems of the year well developed, up to 25 cm long, or strongly abbreviated and densely clothed in approximate leaves, obsolescent; stipules more or less adnate to petiole, free or connate to high up; racemes loose, oblong, strongly elongating in fruit, borne on long axillary peduncles; corolla yellow or whitish or pale yellow, glabrous; calyx at first cylindric, becoming inflated oblong or oval, finally ruptured by the pod; pods sessile or short~stipitate, coriaceous, villous with long white or yellow hairs, bilocular, few-seeded. A central Asian endemic section, distributed in W. Tien Shan and Pamir- Alai. 1. Distinctly caulescent plants, the stem firm; Stipules freer: ane. eee Zhe + Nearly acaulescent plants, the stems strongly abbreviated, clothed in stipules and approximate leaf petioles; lower stipules connate; stems Rarely prancned andothemm sTEmG ety cn car lie el 3. 2. Leaflets (3) 4-8 pairs, the terminal equaling or slightly exceeding the lateral ones; pods semiovaloid to oblong-semiovaloid, 12-18 mm long (Fergana and Alai ranges, Vakhsh)........ 771. A. rhacodes Bge. + Leaflets 1-2 pairs, the terminal larger than the lateral ones; pods linear-oblong, (15) 20-25 mm long (Babatag Range)... ..-...- 77-2 oprainsatleaned rectal, (amore eric ry tokens cal nase rere 772. A. pseudorhacodes Gontsch. 3. Apetricolous plant, 9-15cm tall; stems many, slender, 2-6 cm long; leaflets thin; corolla pale violet........ 773. A. Abolinii M. Pop. + Plants 20-50 cm tall; stems obsolescent, stout, clothed in approximate leaf petioles and stipules; corolla yellow.................. 4. 4. Leaves trifoliate, the single pair of lateral leaflets at a distance from the large terminal leaflet, or simple, the single large leaflet 7-10 cm long; stipules densely covered with appressed hairs (interior Pam.~ Al.: Gazimailik, Karshitau, Babitag and Chul'bair Mountains)...... LY ESA REECR CGAY ANTS YUL AA rete eee mans Tee ante 776. A. bucharicus Rgl. + Leaflets 3-7 (8) pairs; stipules glabrous or nearly so.......... 5. 5. Pods villous with white hairs (Guzar, Kyzyl-Dar'ya river basin, western spurs of Hissar Range). 774. A. pseudomegalomerus M. Pop. + Pods villous with yellow hairs (western spurs of Zeravshan Range). OT thar, eset QeRL ER GeO) ea oI MaNS ace ic Soir ines eect) Rosh Mor 775. A. chrysomallus Bge. Series 1. CAULESCENTES Gontsch. — Stems distinct, firm, the internodes distant; stipules free; corolla yellow; pods borne on a stipe 1-3 mm long. 771. A. rhacodes Bge. Astrag. turk. (1880) 308.—A. jassiensis B. Fedtsch. in A.H. P. XXIV (1905) 236. Perennial, ligneous at base, 23-40 cm tall; stems of the year (7) 10-20 (32) cm long, hoary by appressed hairs; stipules partially adnate to petiole at base, triangular-lanceolate, 3-6 mm long, membranous, glabrate; leaves 7-12 (17) cm long, the petiole 8-17 (20) mm long, both petiole and rachis densely scabrous by appressed white hairs; leaflets (3) 4-8 pairs, oblong-obovate or rounded-obovate, (6) 12-20 (25) mm long, acute or round-tipped, mucronulate, slightly narrowed or cuneate at base, rather thick-textured, scabrous on both sides by appressed white hairs; peduncles equaling to twice the length of the leaves, 5-16 cm long; racemes loose, oblong, (5) 7-14 cm long, (12) 15-25-flowered; bracts linear- 590 [WE in oo 4s ee nc othr wereceancs, Sea: Mocacan aA So re tee Ra MM Oakes 798. A. flavidus M. Pop. + Dwarf plants, 5-18 cm tall; stems 2-6 cm long; peduncles 1-5 cm long; leaflets obovate to oblong-obovate or elliptic, rarely lanceolate. EET ee Ler aR A 7 ee NEE a Se OES AT ery coe 13} 13. Leaflets (1) 2-3 pairs, obovate to oblong-obovate (W. Pam.-Al. — Kashka-darya river basin)......... 795. A. subtrijugus M. Pop. + Leaflets 3-4 pairs, elliptic or lanceolate to lance-oblong (W. Pam.-Al.- Kashka-dakya river basin). 30 2" 2 797. A. ambigens M. Pop. 14. Nearly acaulescent (stems not more than 0.5 mm [sic] long) plants 3-6 cm tall; leaflets obovate to rounded-obovate, rarely elliptic, small, 4-7 mm long (W. Pam.-Al. — high mountains of the Kugitang AME) Rete shen TS) cere 796. A. aemulans (Nevski) Gontsch. + Plants 10-40 cm tall; stems 4-10 cm long; leaflets linear-oblong or narrowly lanceolate or oblong-oval, larger, 10-50 mm long (foothills OIG CUAtTeRITOUNtAINOUSTOEIL) o9-s0s = =) Ss eal mie Mmeclen wae ote kB 15. Leaflets oblong-oval, 15-23 mm broad, subobtuse or more often THOWIAGC USOC) 5: Gkeaaswes ‘c+ Ns aaeews ees 792. A. tanchassii Gontsch. + Leaflets narrowly lanceolate or linear-oblong, 2-4 mm broad, acute. 5a cy URNS Lat RIS inne: lama og’, -=% Aegina nomiaritetae Soa. camaaimes oy! 16. 16. Leaflets hispid; peduncles 2-4 (5) cm long (N. Pam.-Al. — foothills ie AVMaa, TEE WG NERS) Jona eal sen cy Oy. AP alc cielo 793. A. melanocomus M. Pop. + Leaflets velutinous-villous beneath with soft hairs; peduncles 5-10 305 (15) cm long (W. Pam.-Al.— Khodzha gurgarata Mountains) ....... TT ee ae eve ere, PS Me roc ie ner 794. A. pedunculosus M. Pop. 17. Stems pannose with stiff spreading hairs; dwarf plants 6-9 (12) cm tall, stems strongly abbreviated, 0.5-2 (3) cm long (high mountains of Central Tien Shan and Pamir). . 805. A. breviscapus B. Fedtsch. + Stems white by appressed or subappressed hairs; plants (8) 10-20 (40) cm tall; stems (2) 4-12 (35) cm long (lowland wastelands and FeTTo ar nae ie a aah eels RCO eMC raion UPN REE aire Ingo 7.65 3.75 18. 806 18. Peduncles 0.3-0.7 (2) cm long; pod truncate at summit, oblong, 5-9mm long, incompletely bilocular (desert plains from Aral Sea to Balkhash LESION) i...) HOO ARLE A ROT REM Sein ah 1 804. A. chaetodon Bge. + Peduncles 2-5 (8) cm long; pods 11-16 mm long, linear or oblong or linear-lanceolate (foothills of eastern Tien Shan and Dzhungarian Ala EDU) ccc ATE NOME soy deer: Sie aemitis ga gaits Tena Ste uN toni tes Renta, SA) elicit ieee 19. 19. Pod lance-oblong, gradually acuminate into a subulate beak 3-4 mm long; leaflets elliptic (Dzhungarian Alatau) ................. spit PARES. oA AG BRITE Ge ORR NP a 802. A. saccocalyx Schrenk. + Pod linear-oblong, obliquely truncate at the summit; leaflets oblong- elliptic or lanceolate (vicinity of Dzharkent — foothills of Ili river walle, Kiabja)icieh 202)... s.d eMiler WLeRn tO) stk 803. A. suidunensis Bge. Series 1. STENOCYSTES M. Pop.— Plants vested with ascending or spreading bicuspidate hairs; calyx cylindric, not inflated in fruit or slightly so, finally ruptured by the pod; corolla dingy yellow, rarely (in A. nigrimontanus) brownish-green; pod oblong or lance-ellipsoid (W. Pamir-Alai, W. Tien Shan). 786. A. stenocystis Bge. Astrag. turk. (1880) 301.— Exs.: H.Fl. As. Med. No. 396. Perennial, 13-30 cm tall; stems 3.5-15 cm long, suberect, sulcate, pubescent with stiff spreading white hairs and hirsute at the nodes with few long black hairs; stipules triangular-lanceolate, subulate-tipped, 10-17mm long, greenish, covered with white and black hairs; leaves 5-13 cm long, the petiole and the much longer rachis covered with stiff spreading hairs; leaflets oblong-elliptic to lanceolate, (12) 15-25 (35) mm long, 5-7 (9) mm broad, cuneate in outline, acute or rarely subobtuse, mostly subulately mucronulate, covered sparsely above and rather densely beneath with stiff spreading hairs; peduncles 3.5-6 (10) cm long, sulcate, densely hispid- villous with spreading hairs and pubescent; inflorescence commonly narrowly cylindric, rarely oblong-ovoid, 4.5-9 (13) cm long, many- flowered; bracts lance-linear to linear, subulate-tipped, 12-17 mm long, covered with white and black hairs; calyx 18-22 mm long, intially narrowly cylindric, at length scarcely inflated, oblong, covered with long stiff spreading white hairs interspersed on the teeth and onthe tube with scattered black hairs, the linear-subulate teeth 6-9 mm long, surpassed by the standard; corolla dingy yellow; standard 22-24 mm long, the limb oblong- obovate, obtusish, narrowed and slightly constricted below the middle, scarcely auriculate-angular toward base, longer than the claw; wing-petals ; 20-22 mm long, the limb oblong, round-tipped, half the length of the claw; keel 18-21 mm long, the limb rounded-obtuse, half the length of the claw; ovary subsessile, white-sericeous; pods virtually sessile, inequilaterally oblong or lance-oblong 12-14 (17) mm long, compressed laterally, carinate ventrally, grooved dorsally, terminating in an uncinate beak 3-4 mm long, densely villous with soft spreading white hairs, bilocular, 4-6-seeded; seeds angular-reniform, 2.5-3 mm long, smooth, grayish-brown. FI. April-May; fr. May-June. Stone-covered fine-grained slopes of foothills in the semidesert and witchgrass steppe regions. — Centr. Asia: Pam.-Al. (western spurs of Zeravshan and Turkestan Ranges). Endemic. Described from Dzhizman Type in Leningrad. 604 wy | q 4 | é y 809 787. A. excelsior M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, (20) 30-50 cm tall; stems (8) 15-30 cm long, fairly stout, densely lanuginous-villous with white or yellowish hairs and pubescent; stipules lance-linear from a subtriangular base, long-acuminate, 10-15 mm long, covered with stiif white hairs; leaves (5) 8-15 mm long, the petiole 0.5-1.5 cm long, both petiole and rachis rather densely pubescent and villous with spreading white hairs; leaflets 6-9 pairs, oblong or lance- elliptic to lanceolate, rarely elliptic, 1.3 cm long, 0.5-0.9 mm broad, subacute, glabrate above, rather sparsely covered beneath with stiff white hairs; peduncles (3) 5-9 cm long, terminal and axillary, firm, densely lanuginose-pubescent and villous with spreading white hairs and often with black hairs beneath the inflorescence; inflorescence capitate, cylindric, 5-10 cm long, 2.5-3 cm broad, many-flowered; bracts linear, 10-15 mm long, covered with white and shorter black or rarely pure white hairs; calyx cylindric, slightly inflated in fruit and ruptured by the pod, 22 mm long, densely hispid-villous with spreading white hairs, the linear-subulate teeth 7-9 mm long, covered with white hairs interspersed with fewer short black ones, the black hairs occasionally present on the tube but fewer than on the teeth; corolla dingy yellow; standard 22-24 mm long, the limb oblong-obovate, round-tipped, narrowed in lower 1/3, angular at base, twice as long as the claw; keel 20-22 mm long, the limb acutish, the claw 2-2.5 times the length of the limb; pods sessile, lance-oblong to lanceolate, (9) 10-13 mm long, acute, terminating in an uncinate beak (2) 3-4 mm long, compressed laterally, carinate ventrally, rounded or scarcely canaliculate dorsally, densely villous with soft spreading hairs, fully bilocular or nearly so, few-seeded; seeds ovaloid, 2-2.5 mm long, smooth, yellowish, spotted blackish-red. Fl. May-June (July); fr. June-July. (Plate XLVIII, Figure 1). Elevated foothill slopes in the witchgrass steppe belt. — Centr. Asia: T. Sh. (southwestern slopes of Fergana Range extending into Fergana valley). Endemic. Described from Dzhalal-Abad district. Type in Tashkent. 788. A. nigrimontanus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, (5) 8-12 (15) cm tall, rather loosely cespitose; caudex short, woody, buried, branched; stems diffuse, firm, 1-2 cm long, hispid-lanate with spreading white hairs; stipules triangular-lanceolate, acuminate, 7-9 mm long, covered with spreading white and fewer black hairs; leaves 3-5 cm long, the rachis 2-3 times as long as the petiole, both petiole and rachis rather sparsely pubescent and villous with stiff spreading white hairs; leaflets 5-7 pairs, narrowly linear to lance-linear, rarely oblanceolate, 1-3 cm long, 1.5-4 mm broad, hispid scatteredly above and densely beneath with spreading hairs, hoary; peduncles 1-4 cm long, densely lanuginous-pubescent and villous with spreading hairs; inflorescence capitate, compact, ovoid or short-cylindric, 5-8 cm long, 3-3.5 cm broad, many-flowered; bracts linear, acuminate, 10-11 mm long, covered with spreading long white and fewer short black hairs; calyx 18-21 mm long, slightly inflated in fruit, obliquely gibbous at base, 4-5 mm broad, ruptured by the pod, densely villous with white hairs interspersed with few short black ones, the teeth 6-10 mm long, equaling the standard, covered with spreading black and white hairs; corolla brownish-green; standard 20-24 mm long, the limb oblong-obovate, round-tipped, strongly narrowed in lower 1/2-1/3, angular at base, 2-2.5 times the length of the claw; 606 810 811 wing-petals 19-23 mm long, barely exceeded by the standard, the limb oblong, round-tipped, the claw 2-3 times the length of the limb; keel 18-21 mm long, somewhat shorter than the wings, the limb obtuse, half as long as the claw; pods sessile, lance-elliptic, ca. 11 mm long, ca. 4mm broad, compressed laterally, acute, terminating in an uncinate beak Ga. 5 mm long, carinate ventrally, almost rounded dorsally, densely villous with soft more or less spreading white hairs, bilocular, few-seeded. Fl. May-June; fr. June-July. Gravelly, stony, or fine-grained gravelly sites on submontane plains and mountain slopes; often in wormwood associations. — Centr.. Asia: T. Sh. (central and southeastern part of Karatau Range and vicinity of Mt. Dzhambul). Described from Su-Alma plateau in central Karatau. Type in Tashkent. 789. A. chaeturus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 15-30 cm tall, vested with pure white hairs; stems several, 8-18 cm long, firm, angled, densely lanuginous-pubescent and villous with spreading hairs; stipules triangular-lanceolate to triangularly linear- lanceolate, 12-15 mm long, hairy; leaves 9-11 cm long, the petiole much shorter than the rachis, 1.5-2 cm long, pubescent and villous with spreading hairs; leaflets 4-6 pairs, mostly oblong-oblanceolate, obtuse or mucronate, 2-3 cm long, 0.4-0.5 cm broad, hispid rather scatteredly above and densely beneath with fairly short hairs; peduncles 3-4 cm long, firm, sulcate, densely lanuginous~pubescent and villous with spreading hairs; inflorescence compact, cylindric, 7-10 cm long, ca. 3 cm broad, many-flowered; bracts linear, acuminate, 13-17 mm long, covered with rather stiff white hairs; calyx (20) 25-28 mm long, softly villous, the teeth ca. (9) 10-12 mm long, equaling the standard; corolla dingy yellow; standard 24-26 mm long, the limb oblong-obovate, emarginate, narrowed in lower 1/3, scarcely auriculate-angular at base, about twice the length of the claw; wing-petals 20-22 mm long, the limb oblong, round-tipped, half the length of the claw; keel 19-21 mm long, about equaling the wings, the limb obtuse, the claw 2.5 times as long as the limb; ovary sessile, densely white-villous; pods (immature) sessile, lance-oblong, acute, terminating in an uncinate beak, white-villous. Fl. June; fr. June-July. Mounds.— Centr. Asia: T.Sh. (Karatau Range: Kel'temashat, Chaen River). Endemic. Described from Kel'temashat. Type in Tashkent. 790. A. rumpens W. Meff. in Fl. Tadzhik. V (1937) 678.— Cryptor- rhynchus propinquus Nevskii in Acta Inst. Bot. Ac. Se. URSS, Serena a(l93n)a2549— Ione Fl ahadzhik, Vite 03. Perennial, 7-22 cm tall, shaggy with short gray hairs; stems several, assurgent to suberect, 2-7 cm long, pannose-lanate (pubescent and to a less extent villous) with spreading white hairs; stipules lanceolate from a broad base, subulately acuminate, 5-15 mm long, membranously herbaceous, covered with white and black hairs; leaves 4-9 cm long; leaflets (2) 3-6 (7) pairs, obovate to oblong-obovate, sometimes subelliptic, 9-25 mm long, 4.5-12 (15) mm broad, cuneate at base, round-tipped or rarely obtusish, terminating in a short point, unequal (the terminal leaflet larger than the lateral ones), subvelutinous on both sides with short spreading white hairs; peduncles (1.2) 2.5-5 (8) cm long, lanuginous- pannose with white hairs more or less interspersed, especially below the 607 812 inflorescence, withblack hairs; inflorescence cylindric to oblong-ovoid, many- flowered, (2.2) 5-9 (14) cmlong; bracts 5-10mm long, linear-lanceolate to lanceolate, acuminate, covered with white and black hairs; flowering calyx cylindric, 16-22 mm long, villous with white hairs interspersed on the teeth and often also onthe tube withblack hairs, the linear-subulate teeth 5-9 mm long; corolla dingy yellow, slightly suffused with violet; standard 21-24 mm long, the limb oblong-obovate, slightly constricted below the middle, 16-18 mm long, round-tipped or obtusish; wing-petals 19-22 mm long, the limb oblong to oval-oblong, round-tipped, 6.5-8 mm long; keel 16.5-20mm long, the claw 1.5-2 times the length of the obtusish limb; pods sessile or nearly so, ruptured by the pod, oblong or lanceolate, acuminate, 9-15 mm long, 3-3.5 mm broad, terminating in an uncinate beak 3-5 mm long, not grooved, subterete or slightly flattened toward the ventral suture, tomentose-villous to subpannose, bilocular, 4-6-seeded; seeds angular-reniform, greenish- brown, smooth, 3-3.2 mm long. Fl. May-June; fr. June-July. Juniper groves, small-leaved maple woods, and scrub, mostly on stony slopes.— Centr. Asia: Pam.-Al. (mountains of southwest Tadzhikistan and Uzbekistan, mountainous part of Zeravshan River). Endemic. Described from Mt. Teryai in Southern Tadzhikistan. Type in Leningrad. Series 2 ADPRESSEPILOSI Gontsch.— Plants vested with bipartite appressed hairs, except for the spreading-villous calyx; calyx asymmetrically inflated, unilaterally saccate-gibbous, ruptured by the lance-oblong pod; corolla dingy yellow (high mountains of West Tien-Shan, and West Pamir-Alai). 791. A. aschuturi B. Fedtsch. in A.H.P. XXIV (1905) 238. Perennial, 4-6 cm tall; stems many, 1-2 (4) cm long, slender, angled, glabrate or covered with appressed white hairs; stipules free or slightly connate, ovate-lanceolate, acute, 4-10 mm long, herbaceous, covered with ascending white or black hairs; leaves 2-3 (5) cm long, the petiole covered with appressed white hairs, the rachis 1.5-3 times the length of the petiole, leaflets 3-6 pairs, ovate or elliptic, acute, 3-7 mm long, sometimes elongate-lanceolate or linear, up to 25 mm long and 1-2 mm broad, grayish on both sides by appressed hairs; peduncles exceeded by the leaves, 1-2 (3) cm long, thicker than the petiole, firm, covered with appressed or ascending white hairs, these interspersed in upper part with black hairs; inflorescence compactly capitate, spherical or ovoid, 3 cm long, 2-3 cm broad , 10-15-flowered; bracts oblong or lanceolate, acute, the outer 9-15 mm, the inner 7-10 mm long, herbaceous, pubescent with black hairs interspersed with some spreading white hairs; calyx unilaterally gibbous below, gradually attenuate toward the orifice, subovoid, 17-20 mm long, villous with soft spreading hairs, the subulate teeth as long as the tube, about reaching the apex of standard or slightly surpassed by it, covered with short black hairs interspersed with some white ones; corolla bright or dingy yellow; standard 18-21 mm long, the limb oval, 8-9 mm broad at the middle, auriculate-angular at base, 2-2.5 (4) times the length of the claw; wing-petals 16-18 mm long, exceeded by the standard, the limb oblong, emarginate, somewhat shorter than the claw; keel shorter than the wings, 15-15.5 mm long, the limb gibbous beneath, slightly concave above, acutish, the claw 1.5 times as long as the limb; pods sessile, inclosed in the slightly inflated ovoid early-ruptured calyx, lance-oblong, 608 up to 10 mm long and 3 mm broad, compressed laterally, acute, covered with short spreading white hairs, bilocular; seeds ear-shaped, 3 mm long, smooth, brown. Fl. July-August; fr. August-September. Stony mountain sites in the alpine and subalpine zones, occasionally descending into the juniper belt.— Centr. Asia: T. Sh. (Talasskii Ala tau). Endemic. Described from Ashutur river valley in Talasskii Ala tau. Type in Leningrad. Series 3. DOLICHOCARPI Gontsch.— Plants vested with ascending or spreading bicuspidate hairs; calyx asymmetrically inflated, unilaterally saccate-gibbous; corolla dull yellow; pod inclosed in the calyx, lance- oblong (N. and W. Pamir-Alai). 792. A. tanchassii Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. URSS IX (1946). Perennial, ca. 37 cm tall; stem slender, firm, ca. 22 cm long, densely velutinous with short yellowish hairs interspersed with isolated longer spreading hairs; stipules partly adnate at base to petiole, ovate-triangular, 813 acuminate, 11-12 mm long, villous with ascending hairs; leaves 9-11 cm long, the rachis 2.5-3 times the length of the petiole, both petiole and rachis pubescent with ascending hairs; leaflets 3-4 pairs, oblong-oval, subobtuse or more often round-tipped, broadly cuneate at base, (2.5) 4-5.5 cm long, 1.5-2.3 cm broad, sparsely covered on both sides with short subappressed hairs, green, prominently nerved beneath; peduncles stout, 7-10 cm long, densely velutinous with short hairs and sparsely hirsute with spreading hairs, the white hairs interspersed beneath the inflorescence with short black hairs; inflorescence compact, cylindric, 6-11 cm long, 2-2.5 cm broad; bracts linear-lanceolate, villous with short black and long spreading white hairs, ca. 6 mm long; flowers subsessile; calyx soon becoming inflated, unilaterally gibbous-saccate, 15-19 mm long, densely villous with soft spreading white hairs, the teeth and the upper part of the tube covered with black hairs, the linear-subulate teeth 4-7 mm long, somewhat surpassed by the standard; standard dingy yellow, 21-22 mm long, the limb broadly obovate, short acuminate toward apex, slightly emarginate, narrowed downward above the lower 1/4 and angled-auriculate below, 1.5-2 times as long as the claw; wing-petals 21 mm long, the limb oblong, round- tipped, half the length of the claw; keel 20 mm long, the claw 2.5 times the length of the obtuse limb; ovary sessile, densely lanate-villous. Fl. June. Gypseous outcrops and taluses of red-colored strata.— Centr. Asia: Pam.-Al. (former Bukhara territory). Known solely from a single location — Canyon of Central course of Tankhas River near Urt~kishlak. Endemic. Described from Tankhas River. Type in Leningrad. 793. A. melanocomus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 15-20 (25) cm tall, covered with stiff spreading hairs; stems (3) 5-10 cm tall, stout, firm, densely lanate-villous with stiff white hairs, these interspersed below the nodes with long black hairs; stipules oblong- triangular or triangular-lanceolate, acuminate, 8-10 mm long, covered with stiff black and white hairs; leaflets 5-8 pairs, linear-oblong to narrowly lanceolate, (1 0) 15-24 mm long, 4 mm broad, glabrate above, densely hispid-villous beneath; peduncles 2-4 (5) cm long, stout, densely 609 814 815 lanate-villous with spreading short black and long white hairs; inflorescence cylindric, 7-11 cm long, many-flowered; bracts lance-linear, 10-13 mm long, acuminate, covered with long white and chiefly short stiff black hairs; calyx at length strongly inflated, subspherical-ovaloid, strongly narrowed toward orifice, 18-20 (22) mm long, lanate-villous with stiff mainly rather long white hairs interspersed with shorter black ones, the latter predominant on the teeth; calyx-teeth linear-subulate, 6-10 mm long, more or less exceeded by the standard; corolla brightly dull-yellow; standard 20-22 mm long, the limb obovate, 8-9 mm broad, round-tipped, strongly narrowed and slightly constricted in lower 1/3-1/4, auriculate-angular at base, 1.5-2 times as long as the claw; wing-petals barely exceeded by the standard, the claw 2.5 times as long as the oblong round-tipped limb; keel somewhat shorter than the wings, the claw 3-3.5 times the length of the obtuse limb; ovary subsessile, densely lanate-villous with spreading hairs. Fl. May; Tey AT Stony and stone-covered fine-grained slopes of foothills. — Centr. Asia: Pam.-Al. (foothills of Alai and Turkestan Ranges along the southern border of Fergana valley). Endemic. Described from Shakhimardan river basin. Type in Leningrad. 794. A. pedunculosus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 10-40 cm tall; stems erect, 4-17 cm long, rather slender, angled, densely pubescent with spreading white hairs and more sparsely villous with black hairs, these more numerous below the nodes and isolated elsewhere; leaves 5-8 (12) cm long, the 1-3 cm long petiole and the longer rachis rather sparsely pubescent and villous with spreading hairs; leaflets 4-5 (6) pairs, narrowly lanceolate to linear-oblong, 1-3 cm long, 2-4 mm broad, acute, villous above with scattered hairs, densely subvelutinous~ villous beneath with soft and somewhat spreading hairs, green or canescent~ green; peduncles densely pubescent and villous with spreading white hairs interspersed with long black hairs, the latter predominant beneath the inflorescence and few elsewhere; inflorescence cylindric, (4) 5-10 cm long, 2.5-3 cm broad, many-flowered; bracts linear-lanceolate, finely pointed, 7-9 (13) mm long, covered with spreading white and fewer black hairs; calyx obliquely gibbous at base, finally 6-7 mm broad, gradually narrowed toward the orifice, 20 mm long, densely sericeous-villous with long soft white hairs interspersed with short black hairs, the latter fairly numerous on the teeth but concealed elsewhere, the teeth linear-subulate, 7 mm long, somewhat surpassed by standard (by 1-4 mm); corolla pale dull yellow; standard 21-24 mm long, the limb oblong-obovate, round-tipped, 6.5-7 mm broad in the broadest upper part, narrowed and scarcely constricted below the upper third, auriculately angular-dilated in lower 1/3, slightly longer than to twice as long as the claw; wing-petals 20-21 mm long, the limb oblong, round-tipped, half as long as the claw; keel equaling the wings, the claw 2.5 times the length of the semiorbicular acutish limb; ovary subsessile, densely villous with soft white hairs; pods (imperfectly mature) sessile, inclosed in the calyx, lance-oblong, ca. 9mm long, somewhat compressed laterally, carinate ventrally, almost rounded dorsally, uncinately short-beaked, densely villous with spreading white hairs, almost fully bilocular, few-seeded. Fl. June-July. 3 In juniper groves on fine-grained slopes of the intermediate mountain zone.— Centr. Asia: Pam.-Al. (Khadzha-gurgur-ata mountains, Surkhan 610 816 river basin). Endemic. Described from Mt. Khodzha-gurgur-ata. Type in Tashkent. 795. A. subtrijugus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, diffuse, 10-18 mm [?] tall, velutinous-villous with whitish or yellowish hairs; stems 3-6 cm long, stout, firm, densely villous-lanate; stipules lance-linear, finely long-acuminate, 10-15 mm long, hispid-villous; leaves 5-8 cm long, the petiole 1.5-2 cm long, mostly shorter than to as long as the rachis, both petiole and rachis densely lanate-villous with spreading hairs; leaflets (1) 2-3 pairs, large, 1.5-3 (4) cm long, 0.8-2 cm broad, obovate to oblong-obovate, obtuse, rigid, velutinous-villous rather sparsely above and densely beneath; inflorescence cylindric, (5) 7-10 cm long, ca. 2.5 cm broad, many-flowered; bracts lanceolate, narrowly linear, acuminate, ca. 5mm long, white-villous; calyx strongly gibbous at base, gradually narrowed toward the orifice, 19-20 mm long, densely pannose- villous with soft white hairs, the teeth 4-6 mm long, linear-subulate, about reaching the apex of standard; corolla dingy yellow; standard ca. 21 mm long, the limb obovate, scarcely emarginate, 7 mm broad in broadest upper part, narrowed in lower 1/3, angular at base, 1.5-2.5 times as long as the claw; wing-petals barely shorter than to equaling the standard, the limb linear-oblong, round-tipped, the claw 2-3 times the length of the limb; ovary sessile, densely villous with spreading white hairs, Fl. June. Mountain slopes.— Centr. Asia: Pam.-Al. (W., mountains in Yakkabag district). Endemic. Described from Mt. Ishak-Maidan. Type in Tashkent. 796. A. aemulans (Nevski)Gontsch. comb. nova—Cryptorrhynchus aemulans Nevskii in Acta Inst. Bot. Ac. Sc. URSS, ser.I, 4 (1937) 255. Perennial, acaulescent (stems not more than 0.5 cm long), 3-6 cm tall; stipules triangular to lance-triangular, acuminate, 3-4 mm long, herbaceous, white-hairy; leaves 2-5 cm long, the 0.5-1 cm long petiole and the rachis rather slender, pubescent with spreading white hairs and to less extent villous; leaflets 5-7 pairs, obovate to rounded-obovate, rarely elliptic, 4-7 mm long, 2.5-6 mm broad, obtuse or minutely mucronulate, hoary villous-pubescent rather sparsely above and densely beneath with short spreading hairs; peduncles 0.5-2 cm long, densely pubescent and villous with spreading white hairs interspersed beneath the inflorescence with few long black hairs; inflorescence oblong-ovoid to subspherical, 2.5-4 cm long, 2.2-2.5 cm broad, 10-17-flowered; bracts linear, 5-7 mm long, densely covered with spreading white and shorter black hairs; calyx obliquely gibbous at base, finally somewhat inflated, gradually and rather gently narrowed toward the orifice, 15-17 mm long, densely villous with long soft ascendihg hairs, the teeth filiform, surpassed by the standard, covered with white hairs interspersed with short black ones, the tube 1.5 times the length of the teeth; corolla yellow or dingy purplish-yellow; standard 20-21 mm long, the limb obovate, 7-9 mm broad in broadest upper part, abruptly narrowed in lower 1/3, angular at base, 3 times as long as the claw; wing-petals about reaching the apex of standard, the limb oblong, round-tipped, about half the length of the claw; keel exceeded by the wings, ca. 17 mm long, the limb almost suborbicular, obtusish, half as long as the claw; ovary almost fully sessile, densely white-villous. Fl. (June) July. 611 817 Stony sites in the subalpine woodless mountain belt.— Centr. Asia: Pam.- Al. (Kugitang Mountains). Endemic. Described from Kugitang Range, above villages of Khodzha-i-fil' and Markushi. Type in Leningrad. Series 4. CYRTOBASES M. Pop.— Plants vested with ascending or spreading bicuspidate hairs; calyx asymmetrically inflated and saccate- gibbous in lower part; flowers rather dingy yellow; pods inclosed in the calyx, small, ovaloid or ellipsoid or ovoid (W. Tien Shan, W. Pamir-Alai). 797. A. ambigens M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 5-10 cm tall, pubescent-villous to subvelutinous; stems 2-5 cm long, lanate-villous and pubescent; stipules triangular-lanceolate, acuminate, 7-10 mm long, white-lanate; leaves 5-9 cm long, the petiole as long as or more often shorter than the rachis, 1.5-2.6 cm long, pubescent with spreading white hairs, and, to a somewhat lesser extent, villous; leaflets 3-4 pairs, commonly elliptic, 1-2 cm long, 5-8 mm broad, acute, or often narrower and elongate sublinear to 5 cm long and 3-4 mm broad, velutinous-pubescent on both sides with more or less spreading hairs, more densely so beneath; peduncles 1-5 cm long, densely lanate-villous with long soft spreading white hairs; racemes capitate, compact, ovoid or cylindric, 4-8 cm long, many~-flowered; bracts linear-lanceolate, finely long-acuminate, 9-10 mm long, white-hairy; calyx 17-18 mm long, strongly and obliquely enlarged at base, gradually narrowed toward the orifice, densely villous with soft spreading white hairs, the teeth linear-subulate, 5-7 mm long, somewhat surpassed by the corolla; corolla pale dingy yellow; standard 19-20 mm long, the limb oblong-obovate, round-tipped,6.5-7 mm broad in broadest upper part slightly narrowed in lower third, angular at base, 2-2.5 times as long as the claw; wing-petals equaling the standard, the claw about 2.5 times as long as the oblong round-tipped limb; keel 18-19 mm long, the limb almost suborbicular, obtuse, half the length of the claw; ovary borne on a very short stipe, densely white-villous; pods (imperfectly mature) subsessile, ellipsoid or lanceolate, ca. 8 mm long, carinate ventrally, flattened-rounded dorsally, compressed laterally, terminating in a slender hooked beak ca. 5 mm long, densely villous with spreading white hairs, bilocular, few-seeded. Fl. June; fr. July. In the region of mottled strata, on gravelly fine-grained slopes, over outcrops of red sandstone and its detritus; in the witchgrass steppe and juniper belts. — Centr. Asia: Pam.-Al. (Kashka-darya river basin and Kyzyl-darya and Tankhas-darya tributaries). Endemic. Described from j Yakkabag-darya river valley. Type in Tashkent. 798. A. flavidus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, ca. 25 cmtall; stems ca. 14 cm long, densely pubescent~- villous with short yellowish hairs; stipules triangular-lanceolate, acuminate, 8-9 mm long, yellowish-villous; leaves 6-8 cm long, the rachis 2-4 times the length of the petiole, this densely pubescent with soft spreading hairs and villous; leaves 6-8cm long; leaflets 4-5 pairs, oblong-oval, 2-3 cm long, 0.6-1 cm broad, rounded-obtuse, rarely mucronate, pubescent rather scatteredly above and densely beneath; peduncles ca. 7 mm long, densely villous and pubescent to subpannose with soft spreading white hairs; racemes capitate, compact, ovaloid, 3-5 cm long, many-flowered; bracts 837, 1117 612 818 linear-lanceolate, acuminate, 8-9 mm long, villous with soft white hairs; calyx obliquely gibbous at base, ca. 0.5 mm broad, strongly narrowed toward the orifice, 18-19 mm long, densely villous with soft pure white hairs, the linear-subulate teeth ca. 7 mm long, about reaching the apex of standard; corolla pale dingy yellow; standard ca. 19 mm long, the limb oblong-obovate, round-tipped, ca. 6 mm broad in broadest upper part, narrowed in lower third, auriculate-angular at base, twice the length of the claw; wing-petals equaling the standard, the claw slightly more than twice the length of the linear-oblong round-tipped limb; keel ca. 18 mm long, the claw 2.5times as long as the semiorbicular obtusish limb; pods sessile, lance-ellipsoid, ca. 8 mm long, compressed laterally, carinate ventrally, rounded dorsally, terminating in a slender reflexed spinescent beak ca. 5-6 mmlong, densely villous with spreading white hairs, almost fully bilocular, few-seeded. Fl. June; fr. July. Outcrops of red sandstone.— Centr. Asia: Pam.-Al. (Babatag Mountains). Endemic. Described from the vicinity of Dzhida-bulak village in Babatag Mountains. Type in Tashkent. 799. A. polyzygus M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, ca. 20 cm tall; stems (3) 5-9 cm long, stout, very densely pannose-villous with spreading hairs; stipules ovate, acuminate, 8-15 mm long, green, sparsely white-villous; leaves (6) 9-14 cm long, the 1-3 cm long petiole and the rachis lanuginous~-pubescent with spreading white hairs and villous; leaflets 11-15 pairs, oval-orbicular, (0.7) 1-1.5 cm long and broad or rounded-obovate, obtuse or mucronulate, rigid, reticulately veined above, velutinous with gray pubescence, notably so beneath, (0.7) 1.5 cm long and broad; peduncles 1-4 cm long, lanate-villous with long spreading white hairs; racemes cylindric, 8-12 cm long, 2.5 cm thick, many-flowered; calyx 20 mm long, obliquely gibbous at base, in fading gradually narrowed toward the orifice, pannose-villous with subpannose pure white hairs, the linear-subulate teeth ca. 7 mm long, reaching the apex of standard or nearly so; corolla bright dingy yellow; standard 20-22 mm long, the limb broad, subrhomboid-obovate, 7.5-8 mm broad in broadest upper part, attenuate toward apex, obtuse, narrowed and slightly constricted in lower 1/4, auriculate-angular below, 1.5 times the length of the claw; wing-petals equaling the standard, the limb linear- oblong, round-tipped, about half the length of the claw; keel 18-20 mm long, the limb almost suborbicular, obtusish, the claw 2.5 times as long as the limb; ovary sessile, densely villous with soft spreading white hairs; pods sessile, lance-ellipsoid, ca. 9mm long, compressed laterally, carinate ventrally, rounded-convex dorsally, terminating in a hooked beak 4 mm long, coriaceous, densely villous with soft spreading white hairs, bilocular, few-seeded. Fl. May-June; fr. June. Gravelly slopes of low mountains and foothills; also gravelly desert plains. — Centr. Asia: Pam.-Al. (Zerabulak Mountains). Endemic. Described from vicinity of Tym village. Type in Tashkent. 800. A. Zerabulaki M. Pop. in Not. Syst. Inst. Bot. Ac. Sc. URSS X (1946). Perennial, 6-7 cm tall; stems obsolescent, 1-2 cm long, stout, firm, densely villous with long soft white hairs; stipules triangular to oblong- triangular, acute, 5-8 mm long, rigid, densely lanate-villous with spreading 613 819 white hairs; leaves 5-8 cm long, the petiole shorter than the rachis, 820 1-2.5 cm long, fairly slender, densely villous with rather long spreading white hairs; leaflets 4-8 pairs, orbicular to oval-orbicular, obtuse or minutely mucronulate, 5-15 mm long, rigid, velutinous-villous sparsely above and densely beneath, hoary, prominently veined above; inflorescence sessile or nearly so, oblong-ovoid, ca. 5.5 cm long, many-flowered; bracts lanceolate, acuminate, 4-5 mm long, white-hairy; calyx obliquely enlarged at base, gradually narrowed toward the orifice, ca. 20 mm long, densely villous with soft spreading white hairs, the teeth linear-subulate, 7-8 mm long, somewhat surpassed by the standard; standard ca. 22 mm long, the limb oblong-obovate, slightly emarginate, orbicularly enlarged in upper 2/3, ca. 8 mm broad, slightly constricted below, auriculate and angularly enlarged in lower 1/3, 2.5 times as long as the claw; wing-petals equaling the standard, the limb linear-oblong, round-tipped, half as long as the claw; keel ca. 20 mm long, the limb almost suborbicular, obtuse, the claw more than twice the length of the limb; ovary sessile, densely villous with spreading white hairs. Fl. May-June. Stony mountain slopes and their feet.— Centr. Asia: Pam.-Al. (Zerabulak Mountains in West Pamir-Alai). Endemic. Described from Zerabulak Mountains. Type in Tashkent. 801. A. cyrtobasis Bge. ex Boiss. FI. or. II (1872) 497; Astrag. turk. 302% Exsh ont hie Ast Med Nor. saG2h He HR NOM 2ioM- Perennial, 7-15 (18) cm tall; stems 2-5 (8) cm long, densely woolly- pubescent and villous with white spreading hairs; stipules lanceolate, acuminate, 7-8 mm long, more or less covered with white hairs; leaves 5-7 (8) cm long, the petiole and the much longer rachis covered with spreading white hairs; leaflets 6-8 (10) pairs, lanceolate to oblong-obovate, subobtuse to subacute, mucronulate, 11-15 (18) mm long, 4-5.5 (7) mm broad, glabrate above, densely villosulous beneath with spreading hairs; peduncles 0.5-1.5 cm long, stout, densely wooly-pubescent and villous with spreading white hairs; inflorescence oblong-ovoid, cylindric in fruit, 5-12 cm long, many-flowered; bracts linear-subulate, 5-8 mm long, white- hairy; calyx cylindric, becoming inflated in fruit in lower part, obliquely- gibbous, 22-24 mm long, 6-7 mm broad at base, gradually and rather strongly narrowed toward the orifice, densely villous with spreading white hairs, the linear-subulate teeth (5) 7-10 mm long, somewhat exceeded by or equaling to rarely surpassing the corolla; corolla dingy yellow; standard 21-24 mm long, the limb oblong-obovate, obtusish, slightly constricted below the middle, slightly auriculate-angular at base, longer than the claw; wing-petals equaling the standard, the limb oblong to oblong- elliptic, round-tipped, 6-8 mm long; keel 20-21 mm long, somewhat shorter than the wings, the curved obtusish limb 6-7 mm long, 2.9 mm broad; ovary sessile or nearly so, white-shaggy; pods sessile, ovaloid to ovoid, 6-8 mm long, compressed laterally, carinate ventrally, almost rounded dorsally, terminating in a hooked beak ca. 4mm long, densely villous with spreading white hairs, bilocular, mostly 2- or rarely 4- seeded; seeds angular-reniform, 3.5-4mm long, smooth, light brown or greenish. Bl Mayolones ir June = July. Stony and gravelly mountain slopes.— Centr. Asia: T. Sh. (Mogoltau mountains), Pam.-Al. (Aksu river basin on northern slope of Turkestan. Range). Endemic. Described from Mogoltau Mountains. Type in Leningrad. 614 821 Series 5. VESICARII Gontsch.— Plants vested with ascending or spreading bicuspidate hairs except for the appressed-hairy stems; calyx symmetrically ovoid-inflated, not ruptured by the pod; flowers pinkish- violet fading yellow; pods lanceolate or oblong, inclosed in the calyx (desert plains of northern Central Asia, foothills of eastern Tien Shan and Dzhungarian-Ala tau and high mountains of Central Tien Shan and Pamir). 802. A. saccocalyx Schrenk. ex Fisch. et Mey. Enum. Pl. Nov. I (1841) 83; Bge. Astrag. geront. II, 234; Astrag. turk. 308. Perennial, 8-11 cm tall; root stout, woody; caudices strongly abbreviated, buried, woody, 0.5-2 cm long; stems 2-5 cm long, angled, densely appressed-hairy, white; stipules ovate-triangular, 2-3 mm long, acutish, densely covered with appressed white hairs; leaves (3) 5-7.5cm long, the petiole and the much longer rachis rather slender, hoary by appressed white hairs; leaflets 5-6 pairs, elliptic, subacute, (5) 7-11 mm long, (3) 4-6 mm broad, hispid-villous sparsely above and densely beneath with subappressed hairs; peduncles 2.5-3 mm long, covered with subappressed white hairs interspersed beneath the inflorescence with isolated black hairs; racemes fairly loose, 4-5 cm long, few-flowered; bracts lanceolate, 6-8 mm long, acutish, covered with black and white hairs; flowers 1-1.5 mm long; fruiting calyx saccate, ovoid, not ruptured by the pod, 19-20 mm long, thinly membranous, sparsely hispid-villous with spreading white hairs, the teeth linear-subulate, 5-7 mm long, covered with white and fewer black hairs; standard 26-27 mm long, the limb oblong- obovate, slightly narrowed upward and obtusish, 17-18 mm long, gradually narrowed into the claw; wing-petals exceeded by the standard, 23-24 mm long, the claw 2.5 times the length of the oblong limb; keel somewhat exceeded by the wings, 22-23 mm long, the claw 3 times as long as the obtuse limb; pods sessile, lance-linear, acute, 12-16 mm long, 5} 5}, 3) jestaal broad, gradually attenuate into a straight beak 3-4 mm long, compressed laterally, carinate ventrally, grooved dorsally, sparsely hispid-villous with spreading white hairs, bilocular (the septum not quite reaching the beak), few-seeded; seeds angularly short-reniform, 2-2.5 mm long, smooth, dark brown. Fr. June. Dry foothill slopes.— Centr. Asia: Dzu.-Tarb. (Labassy Mountains of the Dzhungarian Ala tau system). Endemic. Described from Labassy Mts. Type in Leningrad. 803. A. suidunensis Bge. in A.H.P. VIII (1880) 378. Perennial, (5) 10-17 cm tall, from a much divided root crown and buried woody caudex 1-3 cm long; stems several, (2) 4-12 cm long, angled, fairly slender, densely covered with appressed or subappressed hairs, white; stipules triangular, acuminate, 3-4 mm long, covered with white hairs; leaves (3) 5-8 cm long, the petiole and the much longer rachis slender, hoary by ascending white hairs; leaflets (4) 5-9 pairs, 7-13 mm long, ; 3-5 (6) mm broad, oblong-elliptic to lanceolate, acute-or rarely subobtuse, fairly rigid, hoary sparsely above and densely beneath by ascending hairs; peduncles 2-5 (8) cm long, sparsely covered with spreading or ascending white hairs interspersed beneath the inflorescence with isolated black hairs; racemes rather loose, broadly ovoid to subspherical, 4-5 cm long, 3-4 mm [sic] broad, many-flowered; bracts lanceolate, acuminate, 5-6 mm long, covered with spreading white and black hairs; pedicels ca. 1 mm long; fruiting calyx ovoid-bladdery, not ruptured by the pod, 18-20 mm long, 615 822 sparsely villous with spreading white hairs, these sometimes interspersed on the tube with isolated (rarely profuse) black hairs, the teeth linear- subulate, 5-7 mm long, covered with white and black hairs; corolla pinkish- violet (?); standard 21-26 mm long, the limb oblong-obovate, slightly attenuate toward apex, obtusish, angular at base, 12-15 mm long; wing- petals equaling the standard, the limb oblong, round-tipped, the claw 2.573 times the length of the limb; keel 19-24 mm long, the claw 3-4 times the length of the obtuse limb; ovary on a very short stipe, white-hairy; pods subsessile, linear-oblong, obliquely truncate, 11-13 mm long, 2.5-3 mm broad, terminating in a beak 0.5-1 mm long, compressed laterally, carinate ventrally, grooved dorsally, thinly coriaceous. mostly rather sparsely villous with spreading white hairs, bilocular (the septum lacking only at the tip), few-seeded Fl. June. Stony substrates.— Centr. Asia: Balkh. (Ili river valley). Gen. distr. ;: Kulja. Described from the vicinity of Suidun. Type in Leningrad. 804. A. chaetodon Bge. in Mém. Sav. Etr. Acad. Pétersb. VII (1851) 272; Bge. Astrag. geront. Il, 234; Bge. Astrag. turk. 301. Perennial, (5) 10-20 (40) cm tall; stems several, (0.5) 2-12 (35) cm long, densely subappressed-lanate, white, angled; stipules triangular to lance- triangular, acutish, 3-5 mm long, white-hairy; leaves (2) 3-7 cm long, the petiole and the much longer rachis rather slender, densely hispid by ascending hairs; leaflets (5) 6-9 pairs, lanceolate to elliptic, 4-8 (15) mm long, 2-4.5 mmbroad, acute, rarely round-tipped, lanate sparsely above and densely beneath with spreading or ascending hairs; peduncles 0.3-0.7 (2) cm long, pubescent with spreading hairs and lanate; inflorescence compact, subsessile, ca. 3 cm broad, 12-20 (25)-flowered; bracts lanceolate, 4-6 (7) mm long, acute, green, covered with black and white hairs; flowers subsessile; fruiting calyx ovoid, (10) 14-17 mm long, rather densely villous with spreading white hairs, bearded-wooly with white hairs on the inside between the teeth; calyx-teeth linear-subulate, (3) 4-5 mm long, covered with spreading white and fewer black hairs; corolla pale pinkish-violet to yellowish; standard 20-25 (28) mm long, the limb oblong-obovate, slightly attenuate toward apex, obtusish, short-attenuate below, angular at base, as long as to 1.5 times as long as the claw; wing- petals 19-24 (26) mm long, somewhat surpassed by the standard, the limb oval-oblong, round-tipped, half the length of the claw; keel equaling the wings, the claw 3 times the length of the obtuse limb; ovary sessile, white-lanate; pods sessile, oblong, inclosed in the calyx, 5-9 mm long, 3-4.5 mm broad, truncate, compressed laterally, carinate ventrally, grooved dorsally, terminating in a reflexed beak 1-1.5 mm long, villous with spreading hairs, few-seeded, partially bilocular (unilocular in upper part); seeds ovaloid, 2.5-3 mm long, smooth, brown. Fl. May; fr. (May) June. (Plate XLIX, Figure 2). Clayey solonetz or solonchak, rarely in sandy or gravelly soils, in the wormwood desert region. — Centr. Asia: Ar.-Casp. (E.), Balkh. Endemic. Described from the deserts between Syr-Darya and Kuvan-Darya rivers. Type in Paris. 805. A. breviscapus B. Fedtsch. in A.H.P. XXIV (1905) 234. — A. oophorus Freyn in Bull. Herb. Boiss. sér. V (1905) 1019. 616 823 824 Perennial, nearly acaulescent, 6-9 (12) cm tall, from a strong woody root; stems several, 0.5-2 (3) cm long, scabrously pannose by a dense coat of short spreading white hairs; stipules triangular to triangular - lanceolate, acuminate, 5-6 mm long, green, covered with Spreading white hairs; leaves 6-11 cm long, the rachis as long as to rarely twice the length of the petiole, both petiole and rachis rather densely hispid-villous with short spreading hairs; leaflets 3-6 pairs, elliptic or obovate or oblong - obovate, subobtuse or rarely subacute, terminating in a short point, (5) 7-15 (18) mm long, (3.5) 4.5-8 mm broad, villous-lanate and canescent on both sides with ascending hairs, more densely so beneath; peduncles 0.6-2.2 cm long, firm, densely lanate with short spreading white hairs, these interspersed above the inflorescence with very few black hairs; inflorescence compact, ovoid to oblong-ovoid, rarely oblong, often subspherical in the bud, 3.5-6 (8) cmlong, 3.5 (4) cmbroad, many~-flowered; bracts lanceolate, acuminate, 7-12 mm long, green, mostly hyaline- margined, covered with spreading white and black or rarely pure white hairs; fruiting calyx ovoid, 17-22 mm long, hirsute-villous with spreading white hairs, these interspersed on the teeth and rarely also on the tube with black hairs; calyx-teeth narrowly subulate, 5-8 (9) mm long; corolla pinkish-violet, drying yellow; standard (17) 20-24 (27) mm long, the limb oblong-obovate, narrowed and slightly constricted below the middle, slightly auriculate-angular at base, 2-4 times the length of the claw; wing- petals (14) 20-23 (27) mm long, the limb oblong to oblong-oval, the claw 1.5-3 times as long as the limb; ovary subsessile or borne on a stipe 0.5-1 mm long, hairy; pods subsessile, oblong, 8-12 mm long, 3.5-4 mm broad, compressed laterally, subtriangular in cross section, carinate ventrally, deeply grooved dorsally, obliquely truncate and terminating in a reflexed beak 2.5-3 mm long, lanate with long spreading white hairs, subbilocular; seeds angularly reniform, 2.5-3.5 mm long, smooth, brown or grayish-brown. Fl. May-June. (July); fr. June-July (August). Stony and gravelly slopes in the high-~mountain zone.— Centr. Asia: T. Sh. (Centr. and W.), Pam.-Al. (Pamir, Shugnan, upper reaches of Zeravshan). Endemic. Described from Central Tien Shan. Type in Leningrad. Section 101) HYPis OP HILL US Bee. Astrag. ceront. (1868) 137 = Herbaceous perennials, vested with bipartite hairs; stems of the year well-developed; stipules connate to high up, not adnate to petiole; inflorescence capitate, spherical (often subumbellate), few-flowered, borne on a long peduncle; bracteoles none; calyx initially cylindric, becoming inflated, bladdery ovaloid or ovoid in fruit, not ruptured by the inflated pod; corolla violet; wings bilobate at apex; pods borne on a very short stipe, small, oblong to ovaloid-oblong, carinate ventrally, canaliculate dorsally, compressed laterally, thinly coriaceous, covered with spreading hairs, subbilocular or partially bilocular.— A section of Soviet Central Asia and the interior of the Asian continent. 1. Fruiting calyx 10-11 (12) mm long, covered almost exclusively with short appressed black hairs, these interspersed with few longer ascending white hairs; standard 15-17 (18) mm long; bracts ovate, Aes OT On esp. erteve: RN eee BMY. ete 806. A. nivalis Kar. et Kir. 617 825 + Fruiting calyx (13) 14-17 mm long, covered with appressed white or short black hairs and villous with spreading white hairs; standard QA 23 | rein, LO veh 9 8 Ls Re NE eer Is et eee See are et 2. 2. Calyx covered with subappressed long white hairs rarely interspersed with short black hairs; bracts covered with white hairs .......... SERPS aE Sa aS ea ate fea tone: 808. A. Nathaliae W. Meff. + Calyx covered with short appressed black hairs and villous with spreading white hairs; bracts covered with black or mixed black and whiteshair's (ei: wees weet BAP eas 807. A. orthanthoides Boriss. 806. A. nivalis Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 341; Bge. Astrag. geront. I, 137; Il, 234; Bge. Astrag. turk. 301, ex parte:> Exs.: HFR No. 1862; H.F.A.M. No. 392. Perennial, 8-25 cm tall, loosely cespitose; stems (4) 8-15 cm long, mostly ascending, often decumbent, covered with short appressed white hairs; stipules connate to 1/2-2/3, 3-6 mm long, covered with appressed white and black hairs, the free portion triangular, acute or acuminate; leaves 2-5 cm long, the short petiole and the rachis covered with short appressed white hairs; leaflets 5-8 pairs, oval to rounded-oval, rarely oblong-oval, round-tipped, (1) 2-5 mm long, covered onboth sides with appressed hairs, canescent-green; peduncles equaling to 1.5 times as long as the leaves, covered with appressed white and beneath the inflorescence mixed white and black hairs; inflorescence spherical, few-flowered, 2.5-3 cm long; bracts oval, 2-3 mm long, covered with black hairs often interspersed with few white ones; calyx cylindric and 8-10 mm long in flower, spherical-ovaloid and 10-11 (12) mm long in fruit, rather densely covered with short appressed or subappressed black hairs interspersed with few longer subappressed white hairs; calyx-teeth trianguar-subulate, ca.. 1mm long; corolla violet; standard 15-17 (18) mm long, the limb oblong-obovate, emarginate, narrowed in lower 1/3-1/4, angular at base, 3-4 times as long as the claw; wing-petals 15-16 (17) mm long, the limb oblong, slightly enlarged toward apex, bilobate, the claw 1.7-2 times as long as the claw; keel 14-16 mm long, obtuse; pods borne on a stipe ca. 0.5-0.7 mm long, ovaloid-oblong, obtuse, 5-6 mm long, 3.5 mm broad, carinate ventrally, canaliculate dorsally, thinly coriaceous, covered with spreading white and black hairs, incompletely bilocular, the reflexed beak 0.6-1 mm long. Fl. July-August; fr. August. Stony slopes and riverside pebbles in the high-mountain zone. — Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala tau, Tarbagatai). T. Sh. (Centr. ; and N. westwardto Talasskii Ala tau). Gen. distr.: Kulja. Described from Dzhungarian Ala tau (fromthe upper reaches of Sarkan River). Type in Moscow; cotype in Leningrad. 807. A. orthanthoides Boriss, nom. n.— A. ortanthus Freyn in Bull. Herb. Boiss., sér. 2, V (1905) 1018, non Gray (1864). Perennial, resembling A. nivalis Kar. et Kir., from which it differs chiefly in the size and vesture of the calyx; bracts oblong-ovate to oblong, (3) 4-5 (6) mm long, covered with black and white or pure black hairs; calyx cylindric and (9) 10-13 mm long in flower, spherical-ovoid and (13) 14-17 mm long in fruit, hirsutulous with appressed black hairs and rather densely villous with spreading white hairs; calyx-teeth subulate, ca. 2 mm long; corolla 21-23 mm long; pods 7-8 mm long, borne ona stipe 1.5 mm 618 826 long, hirsute-villous with spreading white hairs often interspersed with fewer black ones. Fl. July-August; fr. August-September. More or less stony mountain slopes and river valleys inthe high-mountain zone.— Centr. Asia: Pam.-Al. (E.), T. Sh. (westward from Talasskii Ala tau to Fergana Range in southern part of Centr. Tien Shan). Endemic. Described from Pamir, Lake Yashil'-kul and Kara-su River). Type in Geneva; cotype in Leningrad. 808. A. Nathaliae W. Meff. in Fl. Tadzhik. V (1937)678.—Ic.:I.c.t. 52. Perennial, 12-18 (32) cm tall, loosely cespitose; floriferous stems more or less branched, (2.5) 6.5-12 (20) cm long, bearing 1-3 inflorescences; stipules connate to high up, 3-4 mm long, covered with short white hairs, the free portion triangular; leaves 2.5-6 cm long; leaflets 5-7 pairs, elliptic to oblong-obovate, 4-8 mm long, 2.5-4 (5) mm broad, round-tipped, hoary on both sides by a dense coat of short appressed white hairs; peduncles 4-5 (8) cm long, covered withappressed white hairs; inflorescence spherical or rarely ovoid, 5-10-flowered; bracts broadly ovate, covered with white hairs sometimes interspersed with few black ones; calyx cylindric and ca. 12 mm long in flower, spherical-ovoid and 14-15.5 mm long in fruit, densely covered withappressed white hairs, these often interspersed with few shorter black hairs; calyx-teeth triangular-subulate, 1-1.5 mm long, densely white-pannose on the inside; corolla pale violet; standard 21-23 mm long, the limb oblong-obovate, emarginate, narrowed in lower third, angular at base, 4-5 times the length of the claw; wing- petals 18-21 mm long, the limb oblong, bilobate, somewhat shorter than the claw; keel 16-17 mm long, obtuse; pods borne on a very short stipe, oblong to ovaloid-oblong, 8.5-9 mm long, 3-3.5 mm broad, carinate ventrally, complanate dorsally, obtuse, terminating in a reflexed beak 1.5-2 mm long, covered with spreading white hairs; partially bilocular Hl June-July; ir. July-August. Centr. Asia: Pam.-Al. (Kara-Shur, Alai valley). Endemic. Described from Kara-Shur. Type in Leningrad. Section 102. CYSTICALYX Bge. Astrag. geront. I (1868) 133. — Branched frutescent plants, vested with bipartite hairs; caudices and stems of the year well-developed; stipules adnate at base to petiole, free or rarely the lower ones connate at base; inflorescence compactly capitate or subumbellate, borne on a fairly long peduncle; bracteoles none; calyx cylindric in thebud, at length bladdery-inflated, inclosing the pod; corolla yellow, the standard sometimes suffused with violet; pods sessile, oblong, carinate ventrally, canaliculate dorsally, covered with spreading hairs, coriaceous, bilocular or rarely almost unilocular. A distinctive Central Asian section, distributed from S. Altai and W. Mongolia (Irtysh region).to W. Pamir-Alai and Kulja. 1. Fruiting calyx 15-18 mm long; standard 20-25 mm long, rarely shorter (A. inflatus DC.); limb of standard obovate or oblong-obovate.. 2. + Fruiting calyx 8-13 (14) mm long, rarely up to 20 mm long, rarely up to 20 mm long and then limb of standard oblong.............. di 2. Standard 18-19 mm long; leaflets narrowly linear, 3 or more rarely 4 amos VAIHEND Te dey ahs Is ct Seto Mn cent does Loko 813. A. inflatus DC. Standard 20=25. mim) longus). ays deeds cae nem anew, hee melee 3. 619 3. Peduncles (excluding the inflorescence) equaling or somewhat exceeding: thetleaviesicy (tae iirhen Menem eetsn Monatsh, fac toa os Cente a 4. +=) sPeduneles#is>2\time'ss asvlongiastthesleavesi-yayy Wiis) eels eee ene 5. 4. Calyx covered with appressed white and black hairs............ Seen ein ve Oe CAPER CRIME 0 ccc 810. A. Veresczaginii Kryl. et Sumn. + Calyx villous with spreading ferruginous and fewer blackhairs .... sails highs: aan PIne slot diel «eB teh aur aah ab hegre ie ES ea A 809. A. xanthotrichus Ldb. 54 Leaflets, 456) pa ntis iy oy co Svcs sure ceca der R a hatte tl Sands ng ell ae 6. eat eES Rl. 9 Dames.) Wseu ena soialen Ah imanie 814. A. semideserti Gontsch. 6. Branches of the current year densely pubescent with appressed white hairs; leaflets linear to linear-lanceolate (E. T. Sh., Kulja)...2.. SEL eb Mein dec Be aashes! Gre eictien ie gM WAM aie cea 812. A. leucocladus Bge. + Branches of the current year densely covered with appressed white and fewer black hairs; leaflets lanceolate or oblong-elliptic, rarely elliptic omoval (Sauna. | erst cey ssn 811. A. melanocladus Lipsky. 7. Leaflets elliptic or oblong-elliptic or oblong, rarely lanceolate. . . 8. + wiueallers Linearmor lance sine ary oa mn Nee) se ae HAP 827 8. Limb of standard obovate or oblong-obovate, narrowed in lower part. epg att quit TUscal de ta Nemes Oh GM CU EI ORI Mace Wee aL Re at ae oF + Limb of standard oblong, slightly narrowed in upper part........ Sinbad esteneie A eprint M eh SolaahselDah fas 821. A. dendroides Kar. et Kir. 9) Branches of the current year cOvereduwit i Whitey Mellinci.) cients. issietE LG + Branches of the current year covered with white hairs and black hairs. LOM es same Mute acy tok ea ata Men cabal (cima Teg eels? ectha on eideye tics ka be en en a ea en a 10. Leaflets 4-5 pairs, oblong-elliptic; standard 19-20 mm long; peduncles 1.5-2 times as long as the leaves; pods covered with white hairs (DZhungardanyAlaytau)) jee cee, cet: 815. A. cysticalyx Ldb. + Leaflets 5-10 pairs, oblong or lanceolate; standard 15-16 mm long; _peduncles equaling to rarely 1.5 times as long as the leaves; pods covered with black and very few white hairs (Altai, Saur)........ Sees. uke Poe ase A Su seb aa va) Mpeg an aig Ee key ot 818. A. scleropodius Ldb. 11. Calyx covered with appressed black and fewer white hairs; pods covered with short black hairs sometimes interspersed with few white onés| (Altaa))) 2i...gneree ers Al (ak oe 817. A. Majevskianus Kryl. + Calyx villous with spreading white hairs and pilose with very short subappressed black ones; pods hispid-villous with white hairs and minutely black-hirsutulous .... 816. A. intermedius Kar. et Kir. 12. Standard 13-14 mm long, barely surpassed by the calyx, the limb pandurate (Dzhungarian Alatau)...... 822. A. kopalensis Lipsky. + Standard 16-20 mm long, greatly exceeding the calyx, the limb oval- oblong jor oblong st oviateyiiyau.. del base anid ost aol ogvcd fie a ut) ole 13. 13. Leaflets (10) 12-20 mm long; standard 17-20 mm long, the limb oval- oblong; branches of the current year covered with white and fewer black hairs\(DMienShan) Nie. Mera nt 0 819. A. tyttocarpus Gontsch. + Leaflets (16) 22-35 mm long; standard ca. 16 mm long, the limb oblong-ovate; branches of the current year covered with white hairs (Allad Valle yi Pie ye UN vo eehy Witte Pe NT. gale o 820. A. Woldemari Juz. Series 1. MACRANTHI Gontsch.— Fruiting calyx 15-18 mm long; standard 20-25 mm long, the limb obovate or oblong-obovate, narrowed in lower third, more or less angular at base; inflorescence subspherical, few-flowered, often subumbellate. 620 828 i831 809. A. xanthotrichus Ldb. Fl. Alt. III (1831) 324; Ldb. Fl. Ross. I, 642; Bge. Astrag. geront. I, 136; Il, 233; Bge. Astrag. turk. 311; Kryl. Fl. Zap. Sibel Ovso~ le: Tudb. Ic. pl. El. Ross. Ill, taba 96. Frutescent, 50-90 cm tall; caudices up to 25 cm long, covered with castaneous-brown bark; ligneous branches of the preceding year 7-10 cm long, densely covered with appressed white hairs, gray; branches of the current year 6-13 cm long, densely covered with appressed white or yellowish hairs; stipules ovate-lanceolate, acuminate, 7-8 mm long, covered with appressed white and black hairs; leaves 4-7 cm long, the petiole 7-12 mm long; leaflets 3-4 pairs, oblong-elliptic, rarely oblong- obovate, 2-3.5 cm long, 7-10 (12) mm broad, covered scatteredly above and sparsely beneath with appressed hairs; peduncles equaling or slightly exceeding the leaves, covered with scattered appressed white hairs; inflorescence subspherical, 3-5 cm in diameter, few-flowered; bracts lanceolate, 4-7 mm long, black-hairy; fruiting calyx ovoid, 15-18 mm long, hirsute-villous with spreading ferruginous and fewer black hairs; the calyx tube 2.5-3 times the length of the lance-linear teeth; corolla yellow; standard 20-26 mm long, 8-10 mm broad, the limb oblong-obovate, narrowed in lower third, slightly auriculate-angular at base, 3 times as long as the claw; wing-petals somewhat exceeded by the standard, the limb linear-oblong, slightly emarginate, 2-2.2 mm broad, the claw 1.5 times the length of the limb; keel 17-23 mm long, obtuse; pods sessile, oblong, 12-14 mm long, ca. 3.5 mm broad, carinate ventrally, scarcely canaliculate dorsally, abruptly attenuate into a beak 1-1.5 mm long, coriaceous, densely villous with spreading white and golden hairs, these interspersed at the summit and on the beak (sometimes also on the ventral side) with short black hairs, bilocular. Fl. June; fr. July. (Plate XLIX, Figure 1). Mountain slopes.— W. Siberia: Alt. (S.W.). Endemic. Described from Kurchum Mountains. Type in Leningrad. 810. A. Veresczaginii Kryl. et Sumn. in Animadvers. Syst. ex Herb. Unive ctomsk. No. 1-2 (1933); Kryl. Fl: Zap. Sib. VII}, 1706: Frutescent, up to 1 m tall; caudices 15-40 cmlong, covered with lustrous brown bark; ligneous branches of the preceding year densely pubescent with appressed white hairs; branches of the current year 7-13 cm long, whitish by dense appressed white pubescence; stipules adnate at base to petiole, lanceolate to linear-lanceolate, acuminate, 6-10 mm long, sparsely covered with appressed white hairs; leaves 4-6 cm long, the petiole 4-6 mm long; leaflets 3-4 (5) pairs, lanceolate to oblong-elliptic and acute or oblong-obovate and round-tipped, 1.5-2.5 (3.5) cm long, 0.4-1 (1.3) cm broad, covered beneath or on both sides with scattered appressed hairs, green; peduncles exceeding the leaves, 4-7 (10) cm long, sparsely covered with white and beneath the inflorescence mixed white and black hairs; inflorescence subspherical, ca. 4 cm in diameter, few-flowered; bracts lanceolate, acuminate, 3-3.5 mm long, covered with white and black hairs; fruiting calyx ovaloid, 1.5 mm long, rather sparsely covered with white and black appressed hairs, the tube 3-5 times as long as the linear~subulate teeth; corolla yellow; standard 26-30 mm long, the limb oblong, 6-7 mm broad, broadly emarginate, slightly constricted above the lower third, 3-4 times the length of the claw; wing-petals 24-28 mm long, the limb oblong, slightly emarginate, the claw 1.5-1.8 times as long as the limb; keel 22-25 mm long, obtuse; ovary sessile; pods sessile, oblong, 12-15mm long, 3-4mmbroad, carinate ventrally, slightly canaliculate dorsally, 621 PLATE XLVIII 1. Astragalus excelsior M. Pop.—2. A. Chaetodon Bge. 622 832 acuminate to a slightly reflexed straight subulate beak 2-3 mm long, coriaceous, densely hirsute-villous with spreading white hairs, these interspersed on the ventral side with short spreading black hairs, bilocular. Fl. May-June; fr. June-July. Exposed, often gravelly mountain slopes, often in scrub thickets. — W. Siberia: Alt. (S.W.). Endemic. Described from S. Altai (vicinity of Malokrasnoyar and Naryn Ranges). Type in Tomsk. 811. A. melanocladus Lipsky in A.H.P. XXVI (1910) 272. Frutescent, 50-75 cm tall; caudices 15-20 cm long, covered with castaneous brown longitudinally splitting bark; ligneous branches of the preceding year 10-17 cm long, whitish-gray, densely covered with appressed white hairs; branches of the current year (5) 10-35 cm long, densely covered with white hairs interspersed with scattered black ones, whitish- gray; stipules oblongly triangular-ovate to lanceolate, adnate in lower part to petiole, covered with appressed black and fewer white hairs; leaves 5-8 cm long, subsessile, the rachis covered with appressed white hairs; leaflets 4-6 pairs, lanceolate to oblong-elliptic, rarely elliptic or oval, acute, rarely round-tipped, 9-18 (25) mm long, covered scatteredly above and sparsely beneath with appressed hairs; peduncles 2-2.5 times the length of the leaves, 13-15 cm long, rather densely covered with white and toward the summit with mixed white and black hairs; inflorescence spherical to ovoid-spherical, 4-5 cm long, few-flowered; bracts lanceolate, acuminate, 5-7 mm long, covered with black or mixed black and fewer white hairs; calyx covered with short ascending white and black hairs, initially cylindric, 11-12 mm long, the tube twice the length of the teeth; fruiting calyx ovoid, 15-18 mm long, the tube at most 1.5 times as long as the linear-subulate teeth; corolla yellow; standard 20-21 mm long, the limb obovate, 9.5-10 mm broad, narrowed in lower third, emarginate, auriculate-angular at base, 3-4 times as long as the claw; keel 15-16 mm long, obtuse; ovary sessile; pods sessile, oblong, ca. 12 mm long, ca. 3.5 mm broad, abruptly attenuate into a beak 2-3 mm long, carinate ventrally, slightly canaliculate dorsally, coriaceous, villous with short entangled ascending white hairs and puberulent with shorter spreading black hairs, bilocular. Fl. June-August; fr. August. Scrub and grassy mountain slopes, up to the subalpine belt. ; Centr. Asia: Dzu.-Tarb. (Saur.). Endemic. Described from the Kenderlyk and Aby river watershed. Type in Leningrad. 812. A. leucocladus Bge. in Bull. Soc. Nat. Mosc. XXXIX, 3 (1866) 27; Bge. Astrag. geront. I, 136; II, 233; Bge. Astrag. turk. 312. Frutescent, 30-45 cm tall; caudices rather short, covered with castaneous-brown bark; branches of the current year 9-16 cm long, white by dense appressed pubescence; stipules triangular-lanceolate, 4-7 mm long, covered with appressed white and black hairs; leaves 4-6 cm long, sessile, the rachis covered with appressed white hairs; leaflets 4-6 pairs, linear to linear-lanceolate, acute, 13-22 mm long, 3-4 mm broad, covered scatteredly above and sparsely beneath with appressed hairs; peduncles 2-3 times the length of the leaves, 11-15 cm long, covered with appressed white or mixed white and few black hairs; inflorescence compact, subspherical, few-flowered, 4-5 cm long; bracts lanceolate, acuminate, 4-5 mm long, covered with white and black hairs; calyx soon becoming inflated, 15 mm long, rather sparsely villous with long spreading white 623 833 hairs, and pilose with short spreading black hairs; calyx-tube 3 times as long as the filiform teeth; corolla yellow; standard 20-25 mm long, the limb oblong-obovate, emarginate, slightly constricted above the lower third, angular-auriculate at base, twice as long as the claw; wing-petals 19-24 mm long, the limb linear-oblong, slightly emarginate, the claw 1.5 times as long as the limb; keel 16-20 mm long, obtusish; ovary sessile; pods (immature) sessile, oblong, densely hirsute-villous with spreading white hairs and slightly pilose with short black hairs. Fl. June. Mountain slopes.— Centr. Asia: T. Sh. (E.). Gen. distr. : Kulja. Described from the watershed betweenrivers Assaand Merke. Type in Moscow (?). Series 2. MELANOTRICHI Gontsch. — Fruiting calyx 17-18 mm long; standard 18-19 mm long, the limb oblong-obovate, narrowed in lower third; inflorescence spherical-ovoid, few-flowered. 813. A. inflatus DC. Astrag. (1802) 190.—A. melanotrichus Ldb. Fl. Alt. III (1831) 323; Ej. Fl. Ross. I, 643; Bge. Astrag. geront. I, 136; ll, 2333), Bee. wAstrag. tunes tls Kayla bls) Zap. StbeqVils 61009 oc. A. anthylloides Pall. Astr. (1800) 15, tab. 13, non Lam.—Ic.: Ldb. Ic. pl. MieaRessy Il; stab.2 94: Frutescent, 20-40 cm tall; caudices covered with grayish-brown bark; branches of the current year 2-5 cm long, white by a dense coat of appressed white hairs; stipules ovate-lanceolate, acuminate, 5-6 mm long, adnate at base to petiole; leaves 4-7 cm long, the petiole 3-10 mm long; leaflets 2-3 or rarely 4 pairs, linear, acute, 3-4 (5) cm long, 2-4 mm broad, the appressed hairs sparse beneath, isolated above; peduncles 1.5 times the length of the leaves, 6-10 cm long, sparsely covered with appressed white hairs; inflorescence spherical-ovoid, 3-4 cm long (6 cm long in fruit), few-flowered; bracts lanceolate, 4-6 mm long, black-hairy; calyx initially cylindric, ca. 12 mm long, the tube 5-6 times the length of the teeth; fruiting calyx ovaloid-inflated, 17-18 mm long, the tube 3 times the length of the subulate-linear teeth, the appressed white hairs interspersed on the teeth with black ones; corolla yelow; standard 18-19mm long, the limb oblong-obovate, emarginate, narrowed in lower third, angular at base, 2-3 times the length of the claw; wing-petals 17-18 mm long, the limb linear-oblong, entire, the claw 1.5 times as long as the limb; keel 16-17 mm long; ovary sessile; pods sessile, oblong, 9-10 mm long, carinate ventrally, slightly complanate dorsally, compressed laterally, gradually attenuate into a straight subulate beak 2-4 mm long, coriaceous, covered with stiff spreading white hairs, bilocular. Fl. June; fr. July. Mountain slopes.— W. Siberia: Alt. (S.W.). Endemic. Described from the ''Kirghiz Steppes'' according to collections of Sievers and Schangin. Type in Geneva. Series 3. SEMIDESERTI Gontsch. — Fruiting calyx 15 mm long; standard 22-23 mm long, the limb broadly obovate, narrowed in lower third; inflorescence ovoid or oblong, rather many~-flowered; pods unilocular. 814. A. semideserti Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. IX URSS (1946). Frutescent, up to 60cm tall; caudices up to 15 cm long, covered with brown bark; ligneous branches of the preceding year 2—5 cm long, densely 624 834 pubescent with appressed white hairs; branches of the current year 4-10cm long, white by very densely appressed pubescence; stipules adnate at base to petiole, lance-triangular, 2.5-3 mm long, covered with white and black hairs; leaves (3) 5-8 cm long, the petiole and the longer rachis canescent by very short appressed white hairs: leaflets 7-9 pairs, linear, subobtuse, (5) 9-15 mm long, 2-2.5 mm broad, canescent on both sides by appressed hairs; peduncles 1.5-2 times the length of the leaves, 7-12 cm long, covered with appressed white and beneath the inflorescence mixed white and black hairs; inflorescence compact, oblong or ovoid, 5-13 cm long; bracts triangular-lanceolate, 3-4 mm long, lanate with spreading white and black hairs; fruiting calyx ovoid, Slightly constricted at the orifice, 15 mm long, densely villous with spreading white hairs interspersed with few very short black hairs; calyx-tube 3-4 times as long as the linear-subulate teeth; corolla yellow; standard 22-23 mm long, the limb broadly obovate, scarcely emarginate, ca. 11 mm broad at apex and 5-6 mm in the lower narrowed apart, slightly angular at base, 3 times as long as the claw; wing- petals 22-23 mm long, the limb entire, half as long as the claw; keel 20mm long, acute; pods sessile, oblong, broadly and Shallowly canaliculate dorsally, terminating in a subulate beak ca. 3 mm long, coriaceous, densely lanate with spreading white hairs, unilocular. Fl. May; fr. June. Foothills and the ephemeroid-wormwood semidesert belt. — Centr. Asia: Pam.Al. (foothills of the northern slopes of Turkestan Range). Endemic. Described from the vicinity of Ak-tyube village in Nauskii district and known only from this locality. Type in Leningrad. Series 4. MICRANTHI Gontsch. — Fruiting calyx 9-13 (14)mm long; standard 13-19 (20) mm long, the limb obovate to oblong-obovate, narrowed in lower 1/3; inflorescence ovoid or oblong, rather many-flowered. 815. A. cysticalyx Ldb. Fl. Ross. I (1842) 643; Bge. Astrag. geront. I, 136; II, 234; Bge. Astrag. turk. 313.— A. physocalyx Kar. et Kir. in Bull. Soc. Nat. Mosc. XIV (1841) 409, non Fisch. (1837). Frutescent, up to 60 cm tall; caudices short, branched, covered with castaneous-brown bark; branches of the current year 7-17 cm long, covered with scattered appressed white hairs; stipules oblongly triangular - ovate, acuminate, 7-9 mm long, sparsely covered with appressed white hairs; leaves 7-9 cm long, sessile, the rachis sparsely covered with appressed white or mixed white and black hairs; leaflets 4-5 pairs, oblong- elliptic, acute and mucronulate, on the lower leaves sometimes oblong- obovate round-tipped and mucronulate, (13) 25-35 mm long, glabrous above, covered beneath with scattered appressed hairs; peduncles 10-15 (22) cm long, 1.5-2 times the length of the leaves, covered with scattered appressed white hairs; inflorescence ovoid, (3.5) 4-6 cm long, 3.5cm broad, many- flowered; bracts narrowly linear, 7-9 mm long, covered with spreading black hairs; calyx initially cylindric, ca. 12 mm long, the tube 1.5 times the length of the teeth; fruiting calyx spherical-ovoid, 13-14 mm long, villous with white and on the teeth mixed black and white hairs; calyx-teeth linear-subulate, half as long as the tube; corolla dingy yellow; standard 19-20 mm long, the limb oblong-obovate, emarginate, narrowed in lower third, angular at base, twice the length of the claw; wing-petals equaling the standard, limb oblong, slightly emarginate, about half the length of the claw; keel 15-16 mm long, obtusish; ovary sessile; pods sessile, oblong, ca. 10 mm long, densely villous with spreading white hairs. Fl. May-June. 625 835 836 Mountain forests.— Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala tau, Tarbagatai). Endemic. Described from Ul'dzhar in Tarbagatai. Type in Leningrad. 816. A. intermedius Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 340; Bge. Astrag. geront. I, 136; II, 234; Bge. Astrag. turk. 313. Frutescent, 35-70 cm tall; caudices 10-30 cm long, covered with castaneous-brown lustrous bark; branches of the current year 6-16 cm long, densely covered with appressed black and white hairs, gray; stipules triangular -ovate to oblong-ovate, acuminate, 6-8 mm long, covered with black and fewer white hairs; leaves 4-7 cm long, sessile, the rachis covered with appressed white and black hairs; leaflets (4) 5-7 (8) pairs, oblong-elliptic, rarely lanceolate, short-acuminate, (8) 10-20 (28) mm long, glabrous above, sparsely appressed-hairy beneath; peduncles 2-3 times the length of the leaves, finely sulcate, covered with appressed white and black hairs; inflorescence oblong, 10-15 cm long, many-flowered; bracts narrowly lanceolate, acuminate, 4-6 mm long, densely covered with black and white hairs; calyx early inflated, ovaloid, 9-12 mm long, villous with spreading white hairs and pilose with short subappressed black hairs, calyx-tube 3-4 times as long as the linear-subulate teeth; corolla dingy yellow; standard 15-16 mm long, the limb oblong-obovate, scarcely emarginate, slightly constricted above the lower third, angular-auriculate and enlarged below, subpandurate, 2-3 times as long as the claw; wing- petals 14-15 mm long, the limb linear-oblong, slightly emarginate, the claw 1.5-1.8 times the length of the limb; ovary sessile; pods tightly inclosed in the calyx and only slightly rupturing it at dehiscence, sessile, oblong, 6-8 mm long, 3 mm broad, abruptly attenuate into the slightly reflexed subulate 1.5-2 mm long beak, carinate ventrally, slightly canaliculate dorsally, compressed laterally, coriaceous, hispid-villous with spreading white hairs and minutely pilulose with ascending black hairs, bilocular. Fl. June (July); fr. (June) July-August. Open (crab apple) woods, scrub, and stepped mountain slopes. — Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala tau). Endemic. Described from watershed of Baksan and Sarkan rivers. Type in Leningrad. 817. A. Majevskianus Kryl. in Animadv. Syst. ex Herb. Univ. Tomsk. 3 (1932) 1; Kryl. Fl. Zap. Sib. VII, 1707.—Ic.: Animadyv. 3 (1932) 5. Frutescent, 50-100 cm tall; caudices up to 37 cm long, covered with castaneous-brown longitudinally splitting bark; branches of the current year up to 15 cm long, covered with appressed white and black hairs; stipules adnate in lower part to petiole, lanceolate, acuminate, 3-5 mm long, covered with appressed black and fewer white hairs; leaves 3-7 cm, the petiole 3-7 mm long; leaflets 5-8 pairs, elliptic or oblong-elliptic, rarely oblong, round-tipped or minutely mucronulate, 7-20 mm long, 3-10 mm broad, covered on both sides or merely beneath with scattered appressed hairs; peduncles together with inflorescence exceeding the leaves, 8-17 cm long, rather sparsely covered with appressed white and fewer black hairs; inflorescence ovaloid or oblong, 4-5 cm long, 2.5-3 cm broad, finally elongating up to 7-10 cm; fruiting calyx ovaloid, 10-13 (14) mm long, densely covered with appressed black and fewer white hairs, the tube 3-4 times the length of the filiform teeth; corolla pale yellow; standard 16-18 mm long, the limb oblong-obovate, emarginate, narrowed in lower third, angular at base, 2.5 times as long as the claw; wing-petals 15-17mm 626 837 long, the limb linear-oblong, scarcely emarginate, half the length of the claw; keel 14-16 mm long, acute; ovary sessile; pods sessile, oblong, ca. 8 mm long, ca. 2mm broad, carinate ventrally, canaliculate dorsally, abruptly attenuate into a subulate beak ca. 1 mm long, coriaceous, covered with short ascending black hairs often interspersed with few white ones, bilocular. Fl. July; fr. (July) August. Scrub thickets and mountain slopes.— W. Siberia: Alt. (Naryn Range). Endemic. Described from the vicinity of Katon-Karagai. Type in Tomsk. 818. A. scleropodius Ldb. Fl. Alt. III (1831) 326; Ldb. Fl. Ross. I, 642; Bge. Astrag. geront. I, 136; II, 263; Bge. Astrag. turk. 312; Kryl. Fl. Zap. Sib. VII, 1705. Frutescent, branched, 30-50 cm tall; caudices 15-20 cm long, covered with lustrous brown bark; branches of the current year 5-25 cm long, slender, pale brown, sparsely covered with short appressed white hairs; stipules lanceolate, adnate at base to petiole, 6-8 mm long, covered with appressed black hairs; leaves 2.5-6 cm long, the petiole 0.5-1 (2) cm long; leaflets 5-10 pairs, oblong, obtuse or lanceolate short-acuminate, 10-28 mm long, (3) 4-7 mm broad, glabrous or glabrate above, green, covered beneath with scattered appressed hairs; peduncles equaling to rarely 1.5 times as long as the leaves, sparsely covered with short appressed white and black hairs; inflorescence oblong, secund, 4-8 cm long and 1.5-2 cm broad, many-flowered; bracts lance-linear, acuminate, 5-7 mm long, covered with black or mixed white and black hairs; fruiting calyx ovoid to ovaloid, ca. 13 mmlong, covered with appressed black and fewer white hairs; calyx tube 3-4 times as long as the setaceous teeth; corolla yellow; standard 15-16 mm long, the limb obovate, emarginate, narrowed in lower 1/3, angular at base, 3 times the length of the claw; wing-petals 14-15 mm long, the limb oblong, entire, half as long as the claw; keel 13-14 mm long, acutish; ovary sessile; pods sessile, oblong- ovaloid or ovaloid-oblong, canaliculate dorsally, abruptly attenuate into a subulate slightly reflexed beak 1-1.5 mm long, coriaceous, hispid-villous with spreading black and very sparse white hairs, bilocular. Fl. May- June; fr. June-July. More or less stony foothill slopes.— W. Siberia: Alt. (S.W.); Centr. Asia: Dzu.-Tarb. (Saur). Endemic. Described from the vicinity of Baty on Irtysh River. Type in Leningrad. 819. A. tyttocarpus Gontsch. in Not. Syst. ex Herb. Inst. Bot. Ac. Sc. IX URSS (1946). Frutescent, 25-47 cm tall; caudices 8-20 cm long, covered with castaneous-brown bark; branches of the preceding year up to 10 cm long, covered with gray densely appressed-pubescent bark; branches of the current year (3) 5-15 cm long, hoary, densely pubescent-hirsute with white hairs interspersed with black hairs, these isolated or scattered, more numerous at the nodes; stipules adnate in lower part to petiole and the lower ones connate, 4-5 mm long, covered with appressed black and fewer white hairs; leaves 4-7 cm long, the rachis covered with short appressed white and fewer black hairs; leaflets 5-8 pairs, linear, (10) 12-20 mm long, 2-4 mm broad, covered scatteredly above and sparsely beneath with appressed hairs; peduncles 1.5-2 times the length of the leaves, covered with appressed white and beneath the inflorescence mixed white and black, hairs; inflorescence dense, oblong, (4) 6-10 cm long, 2.5-3 cm broad, 627 838 many-flowered; bracts linear-lanceolate, 4-5 mm long; fruiting calyx oblong-ovoid, 8-10 mm long, sparsely villous with ascending white and black hairs; calyx-teeth linear-subulate, 2-2.5 mmlong; corolla dingy yellow; standard 17-20 mm long, the limb oval-oblong, slightly constricted below the middle, scarcely emarginate, yellowish [?] at base, 2-3 times as long as the claw; wing-petals 15-16 mm long, the linear-oblong entire limb half the length of the claw; keel 14-15 mm long, obtuse; ovary sessile; pods sessile, oblong-ovate, ca. 5mm long, ca. 2.5mm broad, carinate ventrally, scarcely canaliculate dorsally, obtusish, terminating in a beak 1-1.5 mm long, coriaceous, covered with short subappressed white and black hairs, bilocular. Fl. June; fr. July. Grassy mountain slopes in the juniper belt. — Centr. Asia: T. Sh. (Kirgiz Ala tau, Fergana Range near Gul'cha). Endemic. Described from Chulek~ Kainda ravine at upper reaches of Bol'shoi Kebin River. Type in Leningrad. 820. A. Woldemari Juz. spec. nova in Addenda XI, p. 668. Frutescent, ca. 40 cm tall; caudices ca. 15 cm long, covered with castaneous-brown longitudinally splitting bark; branches of the current year 11-20 cm long, slender, whitish by dense appressed pubescence; stipules triangular-lanceolate, 5-6 mm long, acuminate, covered with appressed black hairs; leaves 6-7 cm long, sessile; leaflets 5-6 pairs, lance-linear, (16) 22-35 mm long, 3-4 mm broad, covered scatteredly above and sparsely beneath with appressed hairs, acute and mucronulate; peduncles twice the length of the leaves, covered with short appressed white and beneath the inflorescence mixed white and black hairs; inflorescence ovoid, ca. 5 cm long, rather few-flowered; bracts lanceolate, acuminate, 2-2.5 mm long, black-hairy; fruiting calyx oblong-ovoid, ca. 11mm long, sparsely villous with spreading white hairs and pilose with short subappressed black hairs; calyx-tube 4-5 times as long as the linear- subulate teeth; corolla dingy yellow; standard ca. 16 mm long, the limb oblong-ovate, scarcely emarginate, angular-auriculate in lower half, 2-2.5 times the length of the claw; wing-petals equaling the standard, the limb lance-oblong, entire, half as long as the claw; keel 14 mm long; ovary sessile, densely villous with spreading white hairs. Fl. July. Mountain slopes.— Centr. Asia: Pam.-Al. (Alai valley). Endemic. Described from Kal'ta-bulak river valley near depression of Kyzyl-su river. Type in Leningrad. Series 5. DENDROIDES Gontsch.— Fruiting calyx 10-13 mm long; standard 17-25 mm long, the limb oblong, slightly attenuate toward apex; inflorescence oblong, many-flowered. 821. A. dendroides Kar. et Kir. in Bull. Soc. Nat. Mosc. XV (1842) 339; Bge. Astrag. geront. I, 135; II, 239; Bge. Astrag. turk. 309.— A. Komarovii Lipsky in A.H.P. XXVI (1907) 186, ex parte. Frutescent. up to 1 m tall, with grayish bark; young branches covered with short appressed hairs; stipules adnate in lower ‘part to petiole, 5-7 (10) mm long, lanceolate, subulately acuminately, hairy; leaves 3-7 cm long, the petiole and the much longer rachis covered with appressed white hairs; leaflets 4-5 pairs, oval or elliptic, obtusish or round-tipped, mucronulate, (5) 8-17 mm long, covered with isolated hairs or glabrate above, covered beneath with scattered appressed hairs; peduncles 3-14 cm long, covered with appressed black and white hairs; racemes compact, 628 839 cylindric, 3-7 cm long; bracts linear-subulate, 4-6 mm long, rather sparsely black-hairy; calyx cylindric, covered with white and black hairs, becoming ovaloid, inflated, 10-13 mm long, constricted at the orifice; calyx-teeth linear-subulate, 2-2.5 mm long; corolla greenish-yellow; standard 17-25 mm long, the limb oblong, slightly attenuate upward, obtusish and slightly emarginate, broadly cuneate at base, twice as long as the claw; wing-petals (6) -8 mm long, the limb oblong, emarginate, the claw ca. 10 mm long; keel 14-16 mm long, acutish; pods about equaling to somewhat exceeding the calyx, oblong, 9-11 mm long, compressed laterally, coriaceous, carinate ventrally, narrowly complanate.dorsally, densely lanate with short black and long [?] tubercle-borne hairs, bilocular, few-seeded, rupturing the calyx in dehiscence. Fl. June-July; fr. July. (Plate XLIX, Figure 3). Steppes mountain slopes, in scrub and juniper groves.— Centr. Asia: Dzu.-Tarb. (Dzhungarian Ala tau), T. Sh., Pam.-Al. Endemic. Described from Dzhungarian Ala tau. Type in Moscow; cotype in Leningrad. Series 6. PANDURIFLORI Gontsch.— Fruiting calyx 11-13 mm long; standard 13-19 mm long, barely exceeding the calyx, the limb pandurate; inflorescence oblong, cylindric, many-flowered. 822. A. kopalensis Lipsky in A.H.P. XXXVII (1924) 69. Frutescent, 40-70 cm tall; caudices 8-20 cm long, covered with castaneous-brown bark; stems of year (11) 20-42 cm long, roughly pubescent with short appressed white hairs; stipules adnate at base to petiole, triangular-ovate or oblongly triangular-ovate, acuminate, 5-7 mm long, green, sparsely covered with appressed white hairs to glabrate; leaves 7-10 cm long, subsessile; leaflets lance-linear, (5) 6-7 pairs, acute, (1.5) 2-5 cm long, 2-4 mm broad, green, glabrous above, covered beneath with scattered appressed hairs; peduncles 10-22 cm long, 1.5-2 times the length of the leaves, covered with scattered appressed white hairs; inflorescence very compact, cylindric, 6-9 cm long, ca. 2.5 cm broad; bracts narrowly linear, acuminate, 6-8 mm long, densely villous with spreading white and fewer black hairs; calyx initially short-cylindric, ca. 9mm long, the tube 1.5 times the length of the teeth; fruiting calyx ovaloid-inflated, 11-13 mm long, barely exceeded by the corolla, villosulous with spreading white and, on the teeth, mixed white and black hairs, the subulate teeth half as long as the tube; corolla dingy yellow; standard 13-14 mm long, the limb pandurate, scarcely emarginate, 1.5 times as long as the claw; wing-petals equaling the standard, the limb linear-oblong, entire, about half as long as the claw; keel somewhat shorter than the wings, acute; ovary sessile, villous with ascending white hairs; pods small, inclosed in the calyx. Fl. July. (Plate XLIX, Figure 1). Meadows and valleys of mountain streams.— Centr. Asia: Dzu.~Tarb. (Dzhungarian Ala tau). Endemic. Described from Muratkinai ravine in vicinity of Kopal. Type in Leningrad. Section 103. LAGUROPSIS Bge. Astrag. geront. I (1868) 137 (incl. sect. Sphaerocystis Bge., l.c. 138); N. Basil. in Not. Syst. ex Herb. H.B.P., fase. III, 27-30 (1922) 103-120.— Perennial, acaulescent or nearly so, vested with bipartite hairs; stipules slightly adnate to petiole and sometimes connate at base; inflorescence compactly capitate, ovoid, or 629 840 841 oblong, borne on a fairly long peduncle; bracteoles none; calyx initially cylindric, at length bladdery-inflated, ovaloid to subspherical; flowers violet or pink or yellow, rarely white; pods sessile or subsessile, small, inclosed in and not rupturing or rarely rupturing the calyx, pubescent with appressed or spreading hairs, coriaceous, bilocular or unilocular. Species of this section are chiefly distributed in Central Asia, penetrating eastward to Dauria in N. Mongolia, southward to N. Iran, and westward to the Caucasus. wt ¢ I. Key to Species in Flowering Condition Corolla violet or purple or pink, rarely yellowish with a purple keel. Beata’ OR) ee tig al RR She UAT eicHelly stacy et ASR Vern eESE RAEI ae AG Gols io. 0 0 2. Corolla yellow... 2 stha ssyeee: © Ae PO Gb ad cud Rot 26 tr 12. Calyen hirsute) withyspreadine hairs om pvillOusives.)| 4 20-i.9 |) eee ee an Calyseovered withwappressed hairsi- 3 ma: == 4 2) -) ori cme ides Calyx-teeth reaching the apex of standard or nearly so; plants 5-10 cm tall, vested with white hairs (residual mountains of S. E. Kyzyl-kum). LB) Sr Sa ach UL S Line ol ae Bh eal eens ap ae 840. A. kisylkumi Boriss. Corolla 1.5-2 times as long as the calyx; plants 10-25 cm tall, vested at least on the calyx with white and black or rarely pure white hairs. SE ei US ere ae ale) he Rib eu EN cn gh Te a WEIMAR Welds | Gs yo~’ o -o- 4. Leaflets oblong-obovate; standard 20-25 mm long; bracts ovate or oblonga that Ot eee ee seth ety Bite ta 825. A. Krasnovii M. Pop. Leaflets elliptic or lanceolate or linear-lanceolate; standard 15-17 mm long; "orn L6-22/ mm andithen bracts: linear Ai V.s 0. >We eee 5. Peduncies; shorter than) to rarelyrequaling the’leaves) se)0en a 6. Peduncles twice the length of the leaves (Karatau and Nuratau Mountains) 4) 2 Ry Bk oe Fit 2 BO OR 841. A. pulcher Eug. Kor. Leaflets linear-lanceolate or lance-linear; bracts small, lanceolate, 2-3 mm long; standard 15-16 mm long (Karatau and Nuratau Mountains). teva be heoll, BARE aks OOD 5 BET ahs Ole 842. A. inaequalifolius N. Basil. Leaflets elliptic or lanceolate; bracts linear, 5-7 mm long; standard 18-22 mm long (E. Altai, S. Transcaucasia, Dauriya, N. Mongolia). SURO t RALSTON, ole MG Oe Lyne Sa ee. 834) 6A.) Lagunoides i eauls Calys: cover ediexchusively withiwhiteshairse: © i . vale emer: emer 8. Calyx covered with white and black hairs, the latter sometimes confivied? to the teethiy- keri Pi aie.go . Sibeeteee Shoei. eared eee $). Standard 18-22 mm long, the limb oblong-obovate; wing-petals slightly emarginate (sands of Balkh.)......... 839. A. sabuletorum Ldb. Standard ca. 13 mm long, the limb oval, wing-petals entire at apex (Pam.-Al.; Kugitang Range). 846. A. subschachimardanus M. Pop. Leaflets linear to lance-linear; peduncles equaling to 1.5 times as long as the leaves (Alai Range in Shakhimardan river basin).......... eee are eee ane D Merete eae SRE Mey eve ene Ie 845. A. schachimardanus Basil. Leaflets elliptic, or obovate or lanceolate, rarely linear-lanceolate and then peduncles 2-3 times as long asthe leaves ..........-. 10. . Limb of standard oblong or oblong-obovate........-+...+....-. algal Limb of standard broadly oval (northern slopes of Alai Range, Isfairam DEUS) oeeadipi tae dh catced he a ete al ie ik 844. A. violaceus N. Basil. 630 842 ule 14. 15. 16 19. 20. 22. Standard 17-20 mm long; peduncles equaling or slightly shorter than the leaves; bracts ovate-lanceolate (Altai)... 826. A. dilutus Bge. Standard 14-17 mm long; peduncles 2-4 times as long as the leaves; bracts linear-lanceolate (Mugodzhary, Ulutau, Arganaty Mountains, Tien Shan from Chu-Ili Mountains and Kirgiz Ala tau to Karatau and Tashkent Alatau)....... 843. A. Schrenkianus Fisch. et Mey. Calysqcovered withiappressed: hains *. as .u) se 2h) aepiateaee bee ei 13 Calyx villous with spreading hairs, rarely hirsute with ascending HU Sa reat ROS RG tae i taers Nato ke CTS CLASES, FARRIS dehae patdbetts, 20. Leaflets obovate, 3-4 (5) pairs (Caucasus). 836. A. calycinus M.B. Leaflets elliptic or lanceolate, rarely obovate-elliptic and then (6) foblepairs (Altai, Centr-“Acgia) +, 2.7) 208 Taw Bs RI eSe cake. 14. Calyx covered with black and white hairs, the black hairs sometimes COnmInNed tO the teeth LY i chen. sos es chee coy SBee men eens get ain ee, Pa, Me 15. Calyx often covered with white hairs (Kugitang Range) .......... 5 a a RCE RR les aeedinntol ecapte aie cre le 849. A. plumbeus Nevski. Flowers small; calyx 8-10 mm long, with black hairs confined to the orifice; standard ca. 14 mm long, the limb oblong-oval; plants HO oem, talla(Moroltau Wlountainis)ii.i0. % Gect< saps mae ties bis ede Goa PE ra gh ch i). Pio Sy gtbas Cyne be cue os 848. A. pseudonobilis M. Pop. Flowers large; keel 13-17 mm long, rarely 9-10 mm and then the limb of standard oblong-obovate and black hairs scattered all over the CAIs ts HO Remaster: PCr MC ee nt rae ts ieRNMeL Sa Tero 8S 6 2) )9 cE ae 16. Limb of standard elliptic, constricted in lower part; standard 20- 22 mm long; limb of wings bilobate at apex (S. Altai)........... ei FB ee toes seu Bimsoks, Mapua Cuterty Seat 838. A. zaissanensis Sumn. Limb of standard oblong-obovate, narrowed in lower 1/3, angular at base, rarely suboblong-oval and then standard 17-18 mm long and Wing petals Slightly emarginate -.-.. 5+. ...-.-+.5...25°%.. he Fruiting calyx 15-17 mm long, covered with white (very rarely mixed white and black) hairs interspersed on the teeth with black hairs ... RA hr ac ct a leareg ha Mas esse 236; Kryly’ Fly Zap! Sib: Vil ei0S jase: HR Noli sio7- Perennial, acaulescent, 3-10 cm tall, cespitulose; caudex short, stout, buried, with short divisions; stipules ovate-lanceolate, the lower ones triangular, 3-4 mm long, densely covered with subappressed hairs; leaves 2-7 cm long, the rachis 1.5-2 times as long as the petiole, both petiole and rachis appressed-hairy; leaflets 4-6 pairs, elliptic to oblong-elliptic or obovate, 5-12 mm long, densely covered on both sides with appressed hairs, whitish-gray; peduncles somewhat shorter thanto equaling the leaves, together with inflorescence equaling to somewhat exceeding the leaves, covered with appressed white hairs; inflorescence ovoid to subspherical, 2.5-3.5 cm long; bracts ovate-lanceolate, barely exceeding the pedicel, 3-3.5 mm long, covered with subappressed white hairs; calyx initially cylindric, 9-10 mm long; fruiting calyx oblong-ovaloid, 10-11 mm long, covered with appressed black and white hairs, the tube 4-8 times the length of the subulate teeth; corolla pale purple or yellowish with a purple keel; standard 17-18 mm long, the limb oblong-obovate, emarginate, narrowed in lower 1/5-1/4, angular at base, 3 times as long as the claw; wing-petals 16-19 mm long, the limb emarginate or almost entire, barely shorter than the claw; keel 13-16 mm long; pods inclosed in and rather tightly enveloped by the calyx, subsessile, ovaloid to lance-oblong, acutish, 8-9 mm long, 3-3.5 mm broad, carinate ventrally, scarcely complanate dorsally, coriaceous, hispid-villous with spreading white hairs, bilocular. Fl. June-July (August); fr. July-August. Desert-steppe river valleys, in more or less gravelly or saline soils, on pebbles, rarely on stony mountain slopes, — W. Siberia: Alt. (S.E. Dis Gen. distr.: Mong. (N.W.). Described from Tuktugem river valley in Chui river basin. Type in Leningrad. 637 851 Series 3. SPHAEROCYSTES M. Pop.— Pod inclosed in the calyx; flowers yellow. 827.