FOR }H SC MACY GIFT OF Agricultural Educ.Div BIOLOGY LIBRARY G V A GENERAL PHYSIOLOGY FOR HIGH SCHOOLS BASED UPON THE NERVOUS SYSTEM BY M. L. MACY, L.B. Yy ASSISTED BY H. W. NORRIS, A.M. PROFESSOR OF BIOLOGY, IOWA COLLEGE " The physiology of the nervous system is emphatically the physiology of the future." — MICHAEL FOSTER, M.D., F.R.S. NEW YORK-:. CINCINNATI •:• CHICAGO AMERICAN BOOK COMPANY BIOLOGY LIBRARY G CJOPYBIGHT, 1900, BT M. L. MACY. MACY'S PHYSIOLOOT. E-P 7 PREFACE THE effort has been made in this book to unify the study of the parts and the functions of the human organism by the application of approved pedagogical and scientific principles. The teaching of any science proceeds logically from that which is known to that which is not known. Physiology is one of the earliest of the natural sciences to be presented for formal study in school. In respect to man's organism the one sort of knowledge absolutely original and elemental is consciousness — conscious motion and sensation. This it is that forms the most apparent difference between the two kingdoms which manifest the phenomena of life. It is characteristic of animals to possess consciousness, volition, feeling. Plants are, to all appearance, devoid of them all. Hence this study of human physiology is made to begin with that part of the body which is the organ of consciousness — the nervous system. The pupil knows that he thinks and feels and wills and moves, and he studies physiology in order to understand the appara- tus by which these wonders are accomplished. He is here given first (after a few preliminary definitions) a brief sketch of the parts com- posing the nervous system. Next he studies those physical operations into which consciousness enters as an essential quality, and becomes familiar with the organs of motion and sensation. This leads natu- rally to consideration of the provision for the sustenance of those organs — nutrition in its comprehensive sense. Finally the student comes to a more detailed examination of the mechanism for the conscious activities of the human being. Whatever may be true of philosophers, the infant begins the study of physiology at the point here suggested, and follows a method in harmony with this plan. More than one practical teacher has worked out a similar method through years of experience in the class room. By making the nervous system (the center and core of all animal life) the leading thought throughout, a unity of impression is secured, 3 674621 4 PREFACE the actual connection of every vital process with the one nervous system becomes obvious, and the emphasis is placed where it properly belongs. It is believed that this plan has advantages also for the student of general biology. It emphasizes the one grand, obvious distinction between plants and animals. To students of psychology it will likewise commend itself. Because of prevalent ignorance of the nervous system and its due predominance in the animal economy, psychologists have been forced to become physiologists in order to build across the gap, left by the ordinary manner of treatment, between physiology and psychology. Care has been taken to make no statements not in accord with established science, but no effort is made to introduce the newest conjectures. The necessary limitations of a school text-book have been kept in mind as well as the degree of mental development of those for whom the work is designed. It is believed that the instruction respecting alcoholic drinks and narcotics, while complying with the requirements of recent legislation in the various states, will be found to be based upon rational and scientific principles, and to be placed before the student in a manner to win the assent of his reason rather than to create a mere prejudice which further knowledge might overthrow. Nothing is gained by overstatement, and it is always safe to tell the simple truth, for nothing will so surely foster right living as a knowledge of the truth. The writer has had much assistance from experienced and compe- tent teachers and physicians. Dr. A. W. Alvord (M.D., University of Michigan) of Battle Creek, Michigan, has kindly revised the hygienic portions of the book. Mr. H. W. Norris, A. M., Professor of Biology in Iowa College, has read and criticised the whole of the manuscript. All of the experimental work has been prepared by him and will be found of especial value. Many of the illustrations used are such as are commonly found in schoolbooks treating the subject of physiology, but a large number have been adapted from cuts in recent advanced works, mainly those by Morris, Spalteholz, and Van Gehuchten ; while numerous other drawings expressly for this work have been made by Mr. E. W. Atherton under the direction of Professor Norris. CONTENTS PART I INTRODUCTION CHAPTER PAGE I. Matter and Cells 9 II. Tissues and Organs . . . 19 III. A General View of the Nervous System .... 27 PAET II CONSCIOUS NERVOUS OPERATIONS: MOTION AND SENSATION IV. The Framework of the Body, or the Osseous System . 37 V. The Muscular System .59 VI. The Skin as an Organ of Sensation — Touch . . 82 VII. Taste and Smell 94 VIII. The Eye and the Sense of Sight . . . . . .101 IX. The Ear and the Sense of Hearing . . . . .126 X. The Vocal Apparatus . . . .... . 136 PAET III NERVOUS OPERATIONS UNCONNECTED WITH CONSCIOUSNESS XL Blood, Lymph, and Chyle 147 XII. The Circulatory System 153 XIII. Nervous Control of the Circulation 176 XIV. Respiration . . .181 XV. Nervous Control of the Respiratory Apparatus . . . 198 5 6 CONTENTS CHAPTER PACK XVI. Food 201 XVII. The Digestive Apparatus and Nutrition .... 218 XVIII. The Ductless Glands 254 XIX. The Organs of Excretion . . 257 XX. The Heat of the Body 2CC PART IV THE NERVOUS SYSTEM. XXI. Anatomical Description 275 XXII. Functions of the Nervous System ..... 297 XXIII. Hygiene of the Nervous System 314 PAKT V THE PRESERVATION OF HEALTH XXIV. Health and Disease . . . XXV. Common Accidents and Injuries XXVI. Public Hygiene, or General Sanitation GLOSSARY INDEX 335 344 300 395 PART I INTRODUCTION It is customary to divide the study of the human body into three departments: (1) Anatomy, which is the science that describes the structure of the body; (2) Physiology, or the science of the functions, or uses, of. the various parts of the body, and (3) Hygiene, or the science of health, which treats of the care of the body and all its parts for the purpose of maintaining the whole in its best condition for usefulness and enjoyment. The term Physi- ology, as applied to a schoolbook, however] "Js; o to include all three of these lines of study., frig. 1. — The nervous system. CHAPTER I MATTER AND CELLS 1. Living and Lifeless Matter. — What matter is we are not yet able to say, but as it exists in our world it may be separated into two great divisions, — living matter and lifeless matter. So far as present knowledge goes, these two sorts of matter are wholly distinct the one from the other, and lifeless matter never becomes living matter except under the influence of matter already living. The same substances are indeed found in the two sorts of matter, and when living matter is killed, or becomes life- less, no change can be discovered in its weight. That mysterious something called life is therefore not material, and living matter may be said to be only ordinary lifeless matter existing in a different state or condition. 2. Chemical Elements. — A substance ;^}i£ch-8ajirjci;,b^ divided into two or more different kinds of matter is called a chemical element, or a simple substance. All others are called compound substances. Matter is sepa- rated into its elements by processes which affect the mole- cules or the atoms of which it is composed, that is, by chemical analysis. A molecule may be denned as the smallest particle of matter which exists alone and retains most of the proper- ties of the mass of the substance. An atom is one of the ultimate particles of which a molecule is composed The 9 10 molecules of chemical elements are composed of atoms of the same kind. Compound substances have atoms of dif- ferent kinds. There are as many khjds of atoms as of elements. Both atoms and molecules are too small to be seen even with powerful microscopes. A drop of water may be divided mechanically into many small portions, and each part will retain all the characteristics of the original drop. But when the chem- ist separates the oxygen and hydrogen which together make up the drop of water, he has no longer any matter which resembles water, but instead two kinds of gaseous matter of entirely different properties. The water has been resolved by chemical analysis into its chemical elements. Chemical elements unite in different proportions with one another to form a great variety of substances. About seventy-five elements have been isolated by chem- ists, but only a few of them are known to enter into the structure of animal bodies. These are carbon, oxygen, hydrogen, nitrogen, sulphur, phosphorus, chlorine, fluorine, r; Silicon, potja^sjufmj sodium, lithium, calcium, magnesium, 'iro'n, afi(i ntahga'tfese. As a rule these elements exist in , : *$e;Ucftty',i^ sofm?e;spri of combination with one another. '3. * Protoplasm is a name given to living matter. It is a clear, jellylike substance containing minute grains. As protoplasm, it cannot be chemically analyzed, because the process of analysis destroys its vitality so that it is no longer protoplasm, but merely dead, lifeless matter. The one essential thing about protoplasm is that it is alive; dead protoplasm is a contradiction in terms. It has been called " the physical basis of life," because without it life does not exist, and with it there is always life. But the material of which protoplasm is composed is found, when MATTER AND CELLS 11 analyzed after it has ceased to live, to be highly complex. A large part of its weight is water, while its solid portion is chiefly composed of proteids. These are substances found in many foods, white of egg being a familiar example. They contain carbon, hydrogen, oxygen, and nitrogen-. 4. The Cell (Fig. 2). — All living bodies, both plants and animals, are found to consist of cells. Cells are the ultimate units of which living be- ings are made up, just as bricks are the units of which a brick wall is com- posed. A cell is a microscopic bit of protoplasm, with or without an inclos- ing wall, having suspended within it a rounded body of denser material called Fig. 2. — Diagram of , , IT* u v • the Parts of a cel1- the nucleus. It may be living apart, or . - « nucleus. may form one of the units of an organ- & cell body or proto- ism. Plant cells have usually the cell v}*sm- J c cell wall. wall, but an animal cell may be only a naked speck of living matter. Free cells tend to assume a round shape, but under pressure they may take almost any form. 5. The cells of the human body vary in size from -g^- to -g-flVfr of an inch in diameter. All animals begin their existence as single cells, and the life of any animal is the sum of the activities of all its separate cells, while its physical structure consists of the cells themselves and the intercellular matter which they produce, together with the various lifeless substances which they deposit within themselves. 6. Essential Properties of Cells. — All living things pos- sess two properties without which they cannot exist. One of these properties is nutrition, — using the word in its broader sense to include the double process of taking 12 INTRODUCTION in material from outside and building it into the bodily structure, — that is, the making of complex chemical com- pounds out of simpler ones ; and the breaking down, or reducing to simpler forms, of compounds already formed. The former results in growth or repair of cell substance, and is the storing up of energy ; the latter is the setting free of energy, and the production of waste material to be removed as no longer valuable. Nutrition includes all the chemical changes which take place in living matter. The other essential property of living bodies is the power of reproduction, or of giving rise in some way to living beings like themselves. The first of these properties, nutrition, belongs to every individual cell, to every plant, and every animal, as neces- sary to its own continued existence. The second, repro- duction, is needful only for the continued existence of the race, and is in some cases possessed only by certain indi- viduals of the race. Single cells are, however, capable of giving rise by self-division to other cells like themselves. Life is sustained by the ceaseless exercise of the two powers of nutrition and reproduction. 7. Other Properties of Living Cells. — Certain other properties are found to exist in most cells in the body, for example, in the white corpuscles of human blood, which are clearly defined nucleated cells. These are contractility, or instability, that is, the power of changing form without the application of pressure ; irritability, or the power of vigorous action under stimulus, as, for instance, when the blood cells contract under the influence of electricity ; conductivity, or the power of pass- ing on to distant parts of the cell the influence exerted by a stimulus upon a single point; and coordination, or the capacity in all the parts to work together in definite MATTER AND CELLS 13 direction and with regulated strength to accomplish an end, as when a particle of material suitable for building up a cell is drawn in and used for that purpose. 8. Plants and Animals. — No naturalist can at the pres- ent day place his finger upon a line of separation and say: All living things upon this side are plants; all upon that side, animals. It is, indeed, easy to distinguish the higher forms of animal life from the higher forms of plant life, and the most striking difference is that the animal possesses the power of spontaneous movement, while the plant is rooted to one spot. Other distinctions appear as the two forms of life are studied. For example, both are dependent for their continued life and growth upon the food which is supplied from without themselves ; but plants (the fungi excepted) subsist mainly upon carbon dioxide, water, and mineral salts, while animals live upon water and those chemical compounds which have formed part of living bodies, that is, organic materials. Ani- mals cannot use mineral substances as food except as they are mixed with organic matter. But the simplest forms of plant and animal life cannot be distinguished with positiveness from each other. Both consist of single pro- toplasmic cells, and it is not possible to show that the proto- plasm of one is essentially different from that of the other. As animals rise in the scale of being, however, they are found to develop, as plants do not, a nervous system of ever-increasing complexity and importance. Hence man, as an animal, may be said to be distinguished from all other animals by the superiority of his nervous system ; and all the other parts of the human body may be consid- ered as created simply to minister in some way to that superior portion of the human frame which is the direct agent or instrument of the highest manifestations of life. 14 INTRODUCTION 9. The Difference between Plants and Animals in Respect to Stimulus. — Living animal cells possess the property of irritability or excitability, that is, some change in their composition results from the action of stimulus. Vegeta- ble cells also possess this property in some degree. But it is found that, as in the processes of development more and more complex forms of plant life appear, the plant does not develop special organs for the transmission of stimulus. In the animal kingdom, on the other hand, a striking dif- ference appears. In one of the lowest known representa- tives of animal life — the amoeba (Fig. 3, p. 16), which is a mere microscopic lump of naked protoplasm — each minute particle of the protoplasm appears to respond to a stimulus and to transmit it to the adjacent particles, there being no distinction of parts or functions in the single cell. But in the next higher division of ani- mals, the corals, sea anemones, etc., the rudiments of a nervous system are visible, and some division of sense organs appears. It is probable that nervous impressions are received first in but a single form, while a gradual and uninterrupted development of the senses follows as we rise in the scale. That is, one of the lower animals may be said to have but one sense, touch, or a general sen- sibility, — it receives but one kind of sense impression from influences which higher animals recognize as diverse, — while higher animals may distinguish two or more kinds of impression, and so on. It should be noticed that the common division of senses into touch, taste, smell, sight, and hearing is somewhat arbitrary, even man not being always able to discriminate, for instance, between taste and smell, while certain sensations are recognized, such as perception of temperature and of pain, which do not strictly belong to any of the " five senses " so called. MATTER AND CELLS 15 SUGGESTIONS REGARDING THE PRACTICAL WORK The amount of illustrative experimental work in physi- ology that can be done in a high school depends chiefly upon two factors : the material equipment of the school and the tact of the teacher. Vivisection doubtless has its place, but not in the pub- lic schools. Ordinary dissections sensibly performed can be made a successful part of class work in most of our high schools, but occasionally deference to public opinion will require that the dissections be performed only by the teacher, or possibly not at all. No attempt is made in this book to give detailed direc- tions for dissecting, nor for the preparation of material for study with the compound microscope. It is assumed that a teacher of advanced physiology has received some preliminary training in anatomy and microscopical meth- ods. If so, then suggestions will be far better than spe- cific directions. It is not expected that all the experiments will be per- formed by a class. When a compound microscope is not available, some of the exercises must necessarily be omit- ted. It is believed, however, that all the demonstrations, dissections, and experiments can be performed in any school of moderate equipment. A great mistake is made when much apparatus is interposed between the student of elementary science and the objects of his study. The teacher should make sure that the illustration is not sub- stituted for the idea that it is intended to explain. In some instances conditions will require that the teacher perform most of the work* of an experiment, but as far as possible the pupil should himself be responsible for each detail. 16 INTRODUCTION Fig. 3. — Amoeba in eight successive stages of movement. DEMONSTRATIONS AND EXPERIMENTS1 1. Amoeba. — The amoeba is not always easily obtained. If debris of water plants be kept in shallow dishes of water for several days, there can usually be found specimens of amoeba in the scum that forms on the surface of the water, or in the ooze that collects at the edges and bottoms of the dishes. On mount- ing some of the material on a glass slide and ex- amining with the com- pound microscope, there may be seen small, irreg- ular, transparent masses of a jellylike nature moving along very slowly with a rolling or flowing motion (Fig. 3). Attention should be given to the constantly changing form of the animal, which thus exhibits a fundamental characteristic of proto- plasm, instability. If, when an amoeba is fully extended, sending out processes, pxeudopodia, from the main part of the body, the slide be gently tapped, the animal will be seen to contract quickly into a rounded mass, showing another characteristic of protoplasm, irritability, or the capacity of response to stimulus. 2. White Blood Corpuscles. — If a drop of fresh human blood, or preferably of frog's blood, be mount- ed on a glass slide and examined with the compound microscope, among the numerous red corpuscles may be seen a few transparent ones (Fig. 4). On remaining undisturbed for some time they change in shape, Rg 4._Blood ceiiB (corpuscles) or even migrate, in a manner similar of frog. 1 Note to Teachers. — The demonstrations and experiments should precede the recitation of the lessons which they illustrate. The pupil should not be required to describe the brain, for example, until he has studied the dissected organ itself. MATTER AND CELLS 17 to that of the amoeba. Fresh blood may be obtained by pricking the finger with a sterilized needle, and by decapitating or pithing a frog. 3. Movements of Protoplasm in Plants. — The phenomena of protoplasmic movements can be observed in a variety of plants. The Stoneworts, Chara and Nitella, and the stamen hairs of the Spider- wort, Tradescantia, furnish some of the best examples. In all these the protoplasm is inclosed in a cell wall, and when observed with the com- pound microscope is seen to exhibit streaming movements and circulation of particles in Fig. 5. — Compound microscope with sim- ple warming stage (W) attached. the contents of the cell. The response of protoplasm to changes in temperature can be very easily shown by placing the slide on a warming stage upon the microscope stand as shown in Fig. 5. When the warming stage is heated, the protoplasmic movements a,re seen to increase in rapidity up to a certain point. As it cools, the movements become slower. 4. Properties of Protoplasm in Muscle. — In some animals the various tissues retain their vitality and properties for a considerable time after the death of the individual animal. The common frog fur- nishes us one of the best examples of this. If a frog's gastrocnemius muscle with sciatic nerve at- tachments (Fig. 6) be dissected out (see Figs. 7 and 8) shortly after decapitation of the animal, it will retain its properties for a considerable length of time, if kept well moistened with normal salt solution (0.75 per cent solution of common salt). If the nerve be cut with sharp' scissors a contrac- tion of the muscle occurs. Touching the nerve with a red-hot needle produces a similar contraction FLg. 6. — Nerve- muscle prepara- tion. F femur. G gastrocnemius muscle. S sciatic nerve. T tendon (tendo Achilles). MACY'S PUYS. — 2 18 INTRODUCTION in the muscle. Placing the fresh-cut end of the nerve in a saturated solution of common salt brings about a series of contractions in the muscle. The muscle also contracts when the nerve is stimulated with a weak electric cur- rent. If the nerve muscle preparation be placed on a copper plate, and the tem- perature of the latter be raised above or lowered below the normal, there will oc- cur variations in the response of the mus- cle to stimuli. (Elec- trical stimulation will be found most con- venient.) The rela- tion of muscular action to temperature will thus be striking- ly represented. These experiments with the nerve-muscle prepa- ration show that the living substance is irritable, unstable, and conductive of stimuli. M. biceps. sm M. semimembranosus. 5' Cells. — The M. gastrocnemius. st M. semitendinosus. amoeba and the white M. pyriformis. ta M. tibialis anticus. blood corpuscles al- ready studied furnish very good examples of cells which have no fixed form nor definite shapes. The red blood corpuscles of the frog are cells in which the nucleus can be easily dis- cerned by aid of the microscope (Fig. 4, p. 16). Fig. 7. Muscles of the left leg of the frog. Fig. 8 shows distribution of the sciatic nerve. ad M. adductor magnus. sc sciatic nerve. b y p pe M. peroneus. tr M. triceps. H M. rectus interims minor. CHAPTER II TISSUES AND ORGANS 10. The human body, beginning as a single cell, is gradually built up by a process of division and subdivision of that cell, so that the complete, adult man is but a mass of cells with some cementive and connective matter. It is found, however, that differences early appear in the characteristics of different cells, and these differences increase as development proceeds. A group of similar cells having a similar function is called a tissue. 11. Differentiation of Tissues. — In the lowest animals, composed of but a single cell, all the different parts of the body are essentially alike (leaving the nucleus out of consideration) and have the same functions. One part may move as well as another. All parts share in the process of nutrition, and one part responds as well as another to stimulus. But the higher animals are found to be made up of unlike parts, which minister in differ- ent ways to the life of the whole being. As the cells multiply, certain groups of cells become changed in such a manner as to adapt them to the performance of some special function, while other parts are adapted to other functions. A number of cells lying together become modified so as to make up a tissue adapted to a certain purpose. Other cells become modified in a different way to form a tissue adapted to a different 19 20 INTRODUCTION purpose, and the whole body becomes a mass of many different tissues, each having its definite and special char- acteristics and structure. This process is known as dif- ferentiation of the tissues, and is accompanied by what is called physiological division of labor. One tissue is better adapted to the performance of a certain office in the body than are others, and that special work is given it to do, so that the work of carrying on the operations of the body is divided up among the tissues. 12. Organs. — A living being is often called an organism. In order to secure the greatest efficiency in their labors, the various tissues are built up into a multitude of mecha- nisms called organs; for example, the eye, the hand, the liver. Several different kinds of tissue often enter into the structure of a single organ, and the same sort of tissue appears in many different organs. A number of organs so arranged and related to one another as to cooperate in carrying on a special process or series of processes, is called a system, — as the diges- tive system, or the nervous system. 13. Classification of Tissues. — Tissues are variously clas- sified by different authors, but one broad distinction may be noted which divides them into two great classes : (1) The tissues which have to do with the setting free of energy. These are the muscular and nervous tissues. (2) The tissues which have to do with renewing the sub- stances and restoring the power of responding to stimulus. In this second class are grouped all the remaining parts of the body, which include tissues differing widely from one another — from the solid, bony tissue of the skeleton, and the still harder enamel which covers the teeth, to the soft substance composing the brain, the elastic fat which rounds out the figure, and the fluid which we call blood. TISSUES AND ORGANS 21 14. The following table shows this classification : — 1. Tissues which have to do with liberating j Muscular energy — Master Tissues j Nervous a. Pavement 6. Cubical, Sphe- Simple Epi- K^^A-IV roidal, thelium and Co- Epithelial lumnar 2. Tissues which have to do Tissues , c. Ciliated Compound Transitkmal Columnar Ciliated with the pro- tection, sup- port, and Epithe- j Stratified hum Areolar f «• White Squamous or Scaly renewal of Fibrous .... 1 6- Yellow the Master Connective Adipose or Elas- Tissues Tissues ' Cartilage I tic Bone Blood 15. The Master Tissues. — Muscular tissue liberates energy which takes the form of motion attended by some measure of heat. But the changes in muscular tissue by which energy is liberated are guided, regulated, and adapted to the purposes of human life by means of nervous tissue; that is, the muscles are the instrument, but an instrument motionless and useless until the nerves supply the impulse which sets the muscles at work. A muscle develops energy under the action of nervous stimulus con- veyed to the muscle cell through nervous tissue. Energy is set free in nervous tissue when the nervous organ is stimulated by the influence adapted to it. The nerves carry the impulse to the muscle, and energy is liber- ated as motion and heat. Muscular and nervous tissues possess irritability, that is, they respond to stimulus, in 22 INTRODUCTION the one case by contraction, in the other by some change not yet understood, giving rise to what is called a " nervous impulse," and in the act they develop energy by the breaking down of their own substance. Unless that substance is renewed the tissue will cease to respond to stimulus, — will die. The remaining tissues of the body, therefore, are engaged in one way or another in preparing the needful food, in conveying it to these " master tissues," in taking up the waste substances produced in the evolu- tion of energy and preparing them for removal from the body, or in furnishing mechanical support to the body and its various parts. In these processes are involved all the parts which are concerned in digestion, respiration, circu- lation, and excretion. This varied and complex series of operations implies a vast array of muscular movements, and all are governed by the nervous system under the action of its varied stimuli. 16. Epithelial Tissues. — The free surfaces of the body, both within and without, are covered with a tissue called epithelium. Simple epithelium is composed of but one layer of cells, ar- ranged like flat pav- ing stones and fitted together with a very little cementing mate- rial (Fig. 9, A). The epithelium of this va- riety found lining the interior of the blood vessels, and some other surfaces which are not exposed to the outer air, is called endothelium. The cubical, spheroidal, and columnar epithelial tissues are named from the shape of their cells. In ciliated Fig. 9.— Columnar epithelium. A simple. B stratified. TISSUES AND ORGANS 23 epithelium each of the cells is surmounted by tapering, haiiiike filaments (Fig. 10). Compound epithelium is composed of more than one layer of cells (Fig. 11). Epithe- lium contains no blood vessels, but is nour- ished by lymph. It forms the external layer of the skin and the mucous membrane. 17. Connective Tissues exist in many diverse forms, but all are alike in origin, being devel- oped from the same layer of the embryo ; alike in structure, having a large amount of intercellular material ; and alike in func- tion, being devoted to supporting and connecting the master tissues. Fig. 10. — Cili- ated epithe- lium from a small bron- chial tube. A B Fig. 11. — Compound stratified epithelium. A vertical section of the skin. B lateral view of the cells. C flat side of scales like d, magnified 250 diameters, showing the nucleated cells transformed into broad scales. Areolar tissue is made up of cells, white and yellow fibers, and some intercellular matter (Fig. 12). It is found widely distributed through the body, appearing as delicate, elastic, sheathing membrane for muscles, nerves, glands, and other organs. It penetrates into the sub- stance of the organs and connects and supports their various parts. 24 INTRODUCTION White fibrous tissue is found in ligaments and tendons, the tough lining membranes of bones, brain, etc. (Fig. 13, A). It is composed mainly of bundles of strong, white fibers containing nucleated cells. Bundle of White Fibers Fig. 12. — Diagram of areolar tissue. Fig. 13. — Fibrous tissue. A white fibrous tissue. B elastic or yellow fibrous tissue. Elastic or yelloiv fibrous tissue contains yellow, elastic fibers (Fig. 13, B) bound into bundles by areolar tissue. It appears in some ligaments, and in the walls of the arteries, and in the air cells of the lungs. Adipose tissue is the fat of the body, and is found in nearly all parts, usually in connection with areolar tissue. Adipose tissue consists of protoplasmic cell walls filled with liquid fat. 18. Cartilage appears in two forms, — hyaline cartilage (Fig. 14) and fibrocartilage (Fig. 15), the first being clear and free from fibers, the second composed largely of white or yellow fibers. Cells are found in all kinds of cartilage, but there is always a proportionately large amount of inter- cellular matter which is produced by the cells. TISSUES AND ORGANS 25 Fig. 14. — Hyaline cartilage. a group of four cartilage cells, c a cell. n nucleus, m matrix. Fig. 15. — Fibrocartilage. c cartilage cells surrounded by hyaline matrix (m). / fibrous tissue. 19. Bone is more solid than the other tissues. It is penetrated throughout by minute canals, called Haver- sian canals, containing blood vessels (Fig. 16). The final Fig. 16.— Cross section of bone. a lacunae, spaces in living bone occupied by bone cells. & Haversian canal. INTRODUCTION structure of bone is fibrous, and along with the fibers are cells called bone corpuscles, while the cementing material is earthy matter. In many situa- tions, parts which are in early life composed of cartilage become after- ward replaced by bone, through the process called ossification. 20. Blood is a fluid connective tis- sue which conveys nutriment to all parts of the body (Fig. 17). It holds suspended in its liquid por- tion, or plasma, large numbers of cells called corpuscles. These are of two sorts, named from their color the red or colored corpuscles, and the white or colorless corpuscles. The latter have one or more nuclei, while the red corpuscles have none. Fig. 17. — Human blood corpuscles. R red. W white. CHAPTER III A GENERAL VIEW OF THE NERVOUS SYSTEM 21. The great distinguishing feature separating man as an animal from all other animals is, as we have seen, his possession of a nervous system of more complex and intricate structure than any other in the animal kingdom. It is now fully recognized that the nervous system is the central, unifying, coordinating element in the human organism — that for which all other parts exist and to which all are subordinate. In order, therefore, to under- stand clearly the part which each portion of the body is designed to play in the general plan, it is necessary to have some general knowledge of the nervous system, while a fuller study of its parts and their functions may be postponed to a later period. 22. Divisions of the Nervous System. — Physiologists have been accustomed to describe two great divisions of the nervous system, called the central or cerebro- spinal sys- tem, and the ganglionic or sympathetic system, the first having control of sensation and voluntary motion, the second presiding over those vital operations not under voluntary control, its nerves being in general distributed to the internal organs and the blood vessels. It has, however, long been understood that there are not two nervous systems, but one. Still, as a matter of conven- ience in description, the well-known terms are generally 27 INTRODUCTION retained, and the nervous system is treated under its twofold aspect. 23. The Cerebro-spinal System is com- posed of the brain and the spinal cord, with the nerves passing from them to the various parts of the body (see Fig. 1, p. 8, and Fig. 18). 24. The Brain fills the cavity of the skull. It consists of five principal parts : (1) the cerebrum, ; (2) the optic thalami, which are so closely united to the cerebrum as to seem to be a part of it ; (3) the optic lobes, or corpora quad- riff emina, and crura cerebri ; (4) the cerebellum, and with it the pans Varolii ; (5) the medulla oblong ata (Figs. 19 and 20). Looked at from above or from the side, the only parts of the brain that appear are the cerebrum, a part of the cerebellum, and part of the medulla oblongata. 25. Cranial Nerves. — From the un- der surface of the brain arise twelve pairs of nerves (Fig. 19), which pass through openings in the cranial bones and are distributed in a manner to be described hereafter. They are of three classes : (1) Nerves of special sensa- tion ; (2) motor nerves, that is, nerves which carry nervous impulses to the muscles and cause them to contract; Fig. 18.— Brain and and (3) mixed nerves, that is, both sen- spinal cord, ventral •> canterior) ^— sorV and motor. A GENERAL VIEW OF THE NERVOUS SYSTEM Olfactory Bulb ( to which j.s attached the Olfactory Nerve) Pituitary Body____^ Optic Ne rve Optic Chiasma _ Oculomotor Nerve-—. __ Trochlear Nerve Trigeminal Nerve- ___ Pons Faro/ft- _.__ Abducens Nerve- - _ Facial Nerve Auditory Nerve- _ Glossopharyngeal Nerve _ _ Vagus Nerve — Spinal- accessory Nerve — Hypoglossal Nerve- - Medulla Oblongata First Spinal Nerve — Cerebellum- - Spinal Cord-- Second Spinal Nerve Fig. 19. — Ventral (anterior) surface of the brain. 26. The Spinal Cord and Spinal Nerves. — The spinal cord is a column of soft nervous matter, filling the long channel in the spinal col- umn (Fig. 18). Thirty- one pairs of nerves arise from the spinal cord, each having two roots, — a posterior and an anterior root (Fig. 21). Upon the posterior root, just before it unites with the anterior root, is a ,.,,, , „ Fig. 20.— Side view of brain. little knot of nervous a cerebrum, b cerebellum. matter called a spinal c medulla obiongata. 30 INTRODUCTION ganglion. The anterior roots of spinal nerves contain what are called efferent nerve fibers, that is, fibers carry- ing impulses from the nerve center. These are sometimes called motor nerve fibers, because their stimulation usually results in motion; but the term is not strictly accurate, since other than motor im- pulses may pass over efferent nerves. Posterior roots con- Fig. 21. -Diagram of cross sec- tain afferent nerve fibers, that tion of spinal cord, showing is, fibers carrying impulses nerve roots. toward the nerve center. They P posterior root of spinal nerve. G ganglion. A anterior root. are also called sensory nerve S spinal nerve. fibers, because when they are stimulated feeling or sensation most often results, but other impulses than sensory ones may be conveyed by them. 27. The Sympathetic System consists of a chain of ganglia lying on each side and in front of the spinal column, of three main plexuses (or nerve networks) in the cavities of the chest and abdomen, of many small ganglia in all parts of the body, and of an immense number of fine nerve fibers (Fig. 22). Each ganglion of the chain is connected by nerve fibers with the one above and the one below, as well as with the spinal cord. In general, the number of pairs of ganglia corresponds to the number of vertebrae, or seg- ments of the backbone; but there are only three pairs of ganglia in the neck, and in front of the coccyx, or last segment of the backbone, there is only a single ganglion. 28. Gray and White Nervous Matter. — Two kinds of nervous matter, easily distinguished by their color, are found in the body. In the cerebrum and the cerebellum the gray matter is mainly in the surface layer, called the cortex, the deeper portions being of white matter ; while in Cervical Spinal c' Nerves Dorsal Spinal / Nerves \ Lumbar Spinal Nerves f -r^ \ Fig. 22. —Diagram of the sympathetic nervous system. G ganglion chain. Co coccygeal spinal nerve. 31 INTRODUCTION the spinal cord the reverse is the case, a central column of gray matter being surrounded by white matter (Figs. 23 and 24). The gan- glia are composed almost Fig. 24. — Cross see- wholly of tjon of spinal cord, showing arrange- graymat- ment of gray and white matter. 29. Nerve CeUs. — Gray nervous matter is made .7 ^""^ up mainly of nerve cells Fig. 23.— Cross section of the hemispheres of the brain, showing arrangement of (Fig. 25). These vary gray and white matter. much in gize and ghape> Each sends out one or more branches, or processes, one of which forms the central core of a nerve fiber and is called the axis cylinder process. A ganglion is a col- lection of nerve cells imbedded in nerve fibers. 30. Nerve Fibers. — Every nerve fiber has connection with at least one nerve cell, for its central strand, called the axis cylinder, is al- ways the axis cylinder process of a nerve cell (Fig. 26). This is the essential, indispensable part of the nerve fiber. Surrounding the axis cyl- inder there is usually a layer of white, oily mat- Fig. 25. -Nerve cells (mitral) from the ter called the medullary olfactory bulb. , , , . , £ sheath, and outside of that is a thin inclosing membrane called the neurilemma. A GENERAL VIEW OF THE NERVOUS SYSTEM 33 The last is continuous for the whole length of the fiber, but the medullary sheath is broken at short intervals by little spaces called nodes. Some fibers have only the neurilemma, and are there- fore gray in color ; for it is the medullary sheath which gives the characteristic shin- ing white appearance to nerves and nerve fibers. Those fibers possessing the sheath are called medullated nerve fibers; those without it are called nonmedullated. >_ Node of Kanvier -Neurilemma \--Nucleua DEMONSTRATIONS Neuraxon ~~ or Axis Cylinder Medullary ' 'Sheath Fig. 26. -Por- tion of a me- dullated nerve fiber. 6. The Brain. — The brain of the sheep will be found to be very satisfactory in demonstrating to a class the general structure of this portion of the cen- tral nervous system. The brain of the cat, dog, or ox may be used instead. The brain can be removed from the skull by sawing away the roof of the latter and with a scalpel cutting the attaching membranes and nerves. The brain should be prepared some days, or even weeks, before it is needed by the class, and hardened and preserved in some suitable me- dium. Strong alcohol, a 2 to 5 per cent solution of formalin (formol) in water, and a 2 to 5 per cent solution of bichromate of potash are very good hard- ening and preserving reagents. But more satisfactory is the following mixture : 95 per cent alcohol, six parts ; 2 per cent solution of formalin, four parts. When specimens are preserved in a fluid containing for- malin, they should be soaked in water a short time before using, to avoid the irritating effects of formalin vapor on the eyes, etc. Where possible, each student should be provided with one of the preserved specimens. A brain recently removed should be at hand, but it will be found to be too soft for much careful study. With care one pre- served brain may be made to suffice for an entire class. After exami- nation the specimens may be preserved for more detailed study later on. In studying the brain follow the descriptions of the general text. MACY'S PHYS. — 3 84 INTRODUCTION 7. The Spinal Cord. — Procure at a slaughterhouse a spinal cord of an ox and examine it fresh, or preserve it in one of the fluids men- tioned in the preceding section. Preserved portions can be used later in a more careful study of the structure of the cord. 8. The Sympathetic Nervous System. — If the abdominal cavity of a dog, cat, rat, or frog be opened and the viscera displaced, there may be seen on each side of the backbone a white cord with grayish enlarge- ments, ganglia. The two cords and their ganglia constitute the main chains of the sympathetic system. 9. Nerve Fibers. — Tease out with needles in water on a glass slide a small piece of a nerve. Even without the aid of a lens the nerve is seen to be composed of small, threadlike fibers. Examined with the compound microscope the fibrous structure will become more apparent. PART II CONSCIOUS NERVOUS OPERATIONS: MOTION AND SENSATION Of many of the processes which have to do with man's life he is wholly or partly unconscious. The wonderful operations of growth and development go on chiefly with- out his knowledge. The nerve cells which order and direct all the vital activities carry on their work so silently, so regularly, so skillfully — without jar or confusion — that neighboring cells may not even know that they are busy. Of other nervous activities a man is fully conscious, and without his consciousness the object of those operations is not accomplished. A large part of the nervous system and a large part of the other tissues and organs of the body have for their chief business the production of con- scious motion. Other sets of nerves, nerve cells, and special organs are employed in bringing about those experiences called sensations. These two objects are effected through what may be called conscious nervous operations : motion being the result of the action of cer- tain nervous impulses upon bones and muscles ; sensation, the result of the action of other nervous impulses upon the special organs for sensation. In order to understand these conscious nervous opera- tions it is necessary to study the skeleton and joints, the muscular system, the skin as a sense organ, the senses of taste, smell, sight, and hearing, and the apparatus for speech. 35 PECTORAL GIRDLE Fig. 27. — The skeleton. CHAPTER IV THE FRAMEWORK OF THE BODY, OR THE OSSEOUS SYSTEM 31. Functions of the Bones. — Certain parts of an animal body are more essential to its existence than others, and more important to its well-being. These portions are of especial delicacy in substance and structure, and pecul- iarly liable to injury. They therefore require protection. For this purpose the more solid substances which make up the body are arranged to inclose or shield the softer and more delicate parts. In one large group of animal forms, called invertebrates, which includes insects, mollusks, crabs, lobsters, etc., the outer portion of the body consists usually of a more or less hard and tough crust called the exoskeleton, which covers the softer parts. But the higher group of animals, called vertebrates or backboned animals, to which man belongs, possess an inner, bony framework, called the en- doskeleton, so arranged as to form a support and a defense to the more sensitive and more essential parts. In all vertebrates the skeleton consists of a somewhat firm but flexible bony column to which are attached the bones of the head, the ribs, and the pectoral and pelvic girdles, which connect respectively the upper and the lower limbs with the trunk (Fig. 27). The bones of the skeleton furnish the necessary levers and points of support for the muscles which are the organs of motion. 37 88 CONSCIOUS NERVOUS OPERATIONS 32. Provision for Movement in Different Classes of Ani- mals.— A distinguishing feature of the animal kingdom is the power of voluntary motion, and the ability at some period of life to move about from place to place. For this purpose the different classes and orders of animals are provided with a great variety of mechanical devices. Certain aquatic creatures propel themselves by means of pulsations in the whole body. Snakes and worms swim by the undulations of the body. The squid fills a certain cavity within itself with water and then suddenly expels it, and the force of the ejection moves the body in one direction or another, according to the direction of the current of water ejected. The jellyfish propels itself by drawing in and expelling water from its bell-shaped body. Animalcules move themselves by the rapid vibra- tions of innumerable hairlike projections. But all the higher forms of animals move by means of muscles and ligaments attached to an internal solid framework, or skeleton. 33. The Vertebrate Skeleton. — A careful study of any vertebrate skeleton discloses marvelous adaptations for accomplishing the two main objects of protection and motion. Protection to the vital parts might have been secured by means of a rigid, unyielding case, but in order to allow motion in all parts of the body also, the skele- ton is composed of many separate pieces united together by elastic tissues. The whole number of bones in the adult man is about 206, while in the child the number is yet larger, because, as a child grows older, certain bones which are at first distinct (and remain so in the lower ani- mals) grow together to form one. For convenience in study, the bony framework is divided into two parts called the axial skeleton and the appendicular skeleton. THE FRAMEWORK OF THE BODY 39 34. Axial Skeleton. — The bones of the head, neck, and trunk compose the axial skeleton. 35. The Skull. — At the upper extremity of the vertebral column appears the skull (Fig. 28), composed of (1) the cranium, the strong casket which in- closes the most pre- cious part of the animal structure, — the brain — and (2) the various bones forming the skeleton of the face. 36. The Cranium. — In the adult the cranium consists of eight bones, each composed of two firm, compact plates with a spongy layer between. The bones which form the arch Fig. 28. — The skull, of the head are closely joined to- gether by irregu- larly notched edges, the lines of union being called sutures. The cranial bones are named as follows: 1. The fron- tal bone, forming the front of the skull and of especial solid- ity and thickness, as most exposed to injury. 2. The parietal bones, a pair of thin, flat bones meeting along the top of the head. 3. The temporal bones, below the parietal 1 frontal bone. 6 occipital bone. 2 parietal bone. 7 superior maxillary 3 temporal bone. (upper jaw) bone- 4 sphenoid bone. 8 malar bone. 5 ethmoid bone. 9 lachrymal bone. 10 nasal bone. 11 inferior maxillary (lower jaw) bone. 40 CONSCIOUS NKKVnrs OPERATIONS on each side, and having large openings leading into the ear cavities. 4. The sphenoid* forming part of the floor of the brain cavity. 5. The ethmoid, forming part of the floor in front and joined to many of the facial bones. It is perforated for the passage of the olfactory nerves. 6. The occipital, a large bone at the back of the head and also a part of the floor of the skull. It is perforated by small holes through which pass nerves, and by a large opening called the foramen magnum for the passage of the spinal cord to its union with the brain. 37. The Facial Skeleton consists of fourteen bones: 7. The superior maxillary bones, or upper jaw bones, carrying the upper teeth and forming most of the hard palate. 8. The malar bones, or cheek bones. 9. The lachrymal bones, near the inner angle of the socket of the eye, and per- forated for the tear ducts. 10. The nasal bones, forming the roof of the nose. 11. The inferior maxillary or lower jaw bone. 12. The inferior turbinate bones, one in each nostril chamber. 13. The vomer, forming part of the par- tition between the nostrils. 14. The palate bones, which complete the skeleton of the hard palate. 38. The Hyoid. — A small U-shaped bone secured by long ligaments to the base of the skull and lying in the neck at the root of the tongue, is called the hyoid bone (Figs. 79, 80, and 82, pp. 136, 138). It furnishes points of attachment for many muscles. 39. Bones of the Ear. — Three minute bones in the middle ear (Figs. 74 and 75), the malleus, the incus, and the stapes, have to do with the conduction of sound. 40. The Vertebral Column. — In a man of average stature the spinal column is about twenty-eight inches in length. It consists of twenty-six separate bones (Fig. 29). The upper part, which includes uiore than half the whole THE FRAMEWORK OF THE BODY 41 length of the column, is made up of twenty- four separate bones, each called a vertebra. Of these the first or upper seven lie in the neck and are called cervical verte- brce. The next twelve bones of the spinal column are those to which the ribs are attached. They are called the thoracic or dorsal vertebrce. Next come the lum- bar vertebrce, which are larger than any other of the movable vertebrae. They sup- port no ribs, but re- ceive many large and strong muscles. Below these is seen the sacrum, composed in the infant of five vertebrae, which in the adult become one bone. To the broad spaces on its sides are attached the bones of the pelvic arch which Fig. 29. — The vertebral column as seen from left side (.4) and from behind (Li). C 1-7, D 1-12, L 1-5 cervical, dorsal, and lum- bar vertebrae. S sacrum. sp spinous process. Co coccyx. ' tr transverse process. 42 CONSCIOUS NERVOUS OPERATIONS supports the lower limbs. It lias eight openings, which communicate with the canal inclosing the spinal cord and permit the passage of spinal nerves. At the lower end of the spinal column is the coccyx, formed by the union of four very small vertebrae into one bone. It is that part of the skeleton which in the lower vertebrate animals forms the tail. 41. The Vertebrae vary somewhat in form, but are all constructed upon the same general plan (Fig. 30). There is a stout bony cylinder, called the body, or centrum. To this solid cen- trum is attached an arch, called the neural arch, which forms, with the back of the centrum, an inclosed space named the neural ring. The successive neural rings form in the spinal column a long tube in which the spinal cord may safely lie. From the back of the neural arch extends a long bony projection called the spinous process, and the successive processes, or spines (Fig. 29), extending down the backbone, give to it the name of spinal column. Six other processes project from vertebra: one on each side called transverse pro- cesses; two called anterior articular processes, extending forward ; and two called posterior articular processes, extending backward, to meet the corresponding processes of the neighboring vertebrae. These processes form, by means of the intervening cushion of cartilage and con- necting ligaments, a joint permitting a slight amount of motion. 2 Fig. 30. — A dorsal verte- bra shown in two posi- tions. 1 centrum. 2 processes. 3 neural ring. each THE FRAMEWORK OF THE BODY 43 Two shallow depressions in the forward portion of the centrum of each of the dorsal vertebrae, with correspond- ing depressions in the adjoining vertebra, form pits which receive the heads of the ribs. Similar depressions at the ends of the transverse processes of the dorsal vertebrae assist in securing the ribs to the spine (Fig. 31). Fig. 31. — Articulation of a pair of ribs to a vertebra. ce centrum of the vertebra. r rib. tr transverse process. co costal cartilage. st sternum. Between each two adjoining vertebras are elastic cush- ions of fibrocartilage, which assist in providing for motion and flexibility in the spinal column and in preventing injurious jarring of the brain and spinal cord. 42. The Atlas and Axis. — The first two cervical vertebras have certain modifications of structure for the sake of the freer motion needful in the neck. The atlas or first 44 CONSCIOUS NERVOUS OPERATIONS vertebra (Fig. 32, A) supports the skull, being articulated by two shallow hollows (a, J) with corresponding pro- jections on the occipital bone above. This permits the head to rock back and forth. The atlas has a very small body or centrum, and a large neural ring subdivided by the transverse ligament. Into the front portion of the ring projects the odontoid process, a thick bony peg arising from the axis or second cervical vertebra (Fig. 32, jB). Around the odontoid process the atlas rotates, carrying the head with it from side to side. Fig. A atlas. B atlas and axis, a, b articulations with occipital bone. c transverse ligament, o odontoid process. 43. By means of the variations in the form of the ver- tebrae, and by the four curvatures seen in the spinal column, a considerable range of movement is provided for. The vertebrae furnish strong support for the great mus- cles of the trunk and a safe channel for the spinal cord, while a firm but flexible and elastic column is secured for the support of the whole frame. 44. The Ribs. — Attached by their heads to the tho- racic vertebrae are the twelve pairs of slender curved bones called ribs (Fig. 33). The upper seven pairs are attached in front by costal cartilages to the sternum, or breastbone. The next three pairs have their costal cartilages joined each to the cartilage of the preceding rib ; while the last THE FRAMEWORK OF THE BODY 45 two pairs have their front cartilage ends unattached, and are therefore called floating ribs. All the ribs have a downward slope, their front ends being lower than the hinder ones. This per- mits of a considerable enlargement in the size of the cavity of the thorax, or chest, when, by the contraction of the muscles of the chest, the front ends of the ribs are raised. The object of this will be shown later. 45. The Sternum, or breastbone (Figs. 27 and 33), supports the forward ends of the ribs (with the excep- tion of the two lowest, or floating ribs) by means of the costal pig . 33._skeleton of thorax> showing ribs> cartilages, which give costal cartilages, sternum, and some of the more freedom of move- thoracic vertebr»- ment than would be possible were the bones solid to the end. The sternum is composed, in the adult, of three pieces, the lowest being of cartilage. 46. The Appendicular Skeleton is composed of the pectoral girdle, the pelvic girdle, and the bones of the limbs. 47. The Pectoral Girdle (Fig. 27) consists of four bones, two on each side, — the scapula and the clavicle. The scapula, or shoulder blade, is a triangular, nearly flat bone lying at the back of the shoulder and not attached 46 CONSCIOUS NERVOUS OPERATIONS directly to the spinal column. It has a shallow pit at one of the upper corners for the end of the humerus, or upper arm bone, and a projection to which the other bone of the arch, the clavicle, or collar bone, is secured. The clavicle is a round, slender bone, attached by its two ends to the scapula and the sternum. 48. The Upper Limbs contain, each, thirty bones. They are the humerus, or upper arm bone ; the radius and ulna, side by side in the lower arm ; the eight small bones of the carpus, or wrist ; the five cylindrical bones of the metacarpus, or palm of the hand ; and the phalanges, or finger bones, fourteen in number, two being in the thumb and three in each other finger. 49. The Pelvic Girdle (Fig. 27) is formed by one large spreading bone on each side, called the os innominatum, or hip bone. On the outer side is a deep socket for the head of the femur. The hip bones are made to support great weight and to resist severe shocks. They sustain the whole pressure of the trunk and of burdens carried, and also receive the force of the various movements of the lower limbs, as in running, jumping, cycling, etc. 50. The Lower Limbs are similar in structure to the upper. The femur, or thigh bone, the largest bone in the body, corresponds to the humerus ; the tibia and fibula, to the radius and ulna. In the ankle are seven tarsal bones, and in the arch of the foot five metatarsals, to which are added the fourteen phalanges, or bones of the toes. There is, besides, a bony disk, imbedded in the great liga- ment over the knee, forming a protection to the knee- joint, and called the patella, or kneepan. 51. Observe the provisions in the human skeleton for securing firmness and strength to the upright figure. It has been found that the arch is the strongest form of THE FRAMEWORK OF THE BODY 47 structure for a given amount of material. The shoulder arch of the skeleton furnishes support to the arms so strong that those limbs may be used to lift great weights and hurl them through the air, and to perform a great variety of labors. The pelvic arch and the arches of the foot are also designed to support securely the tall human figure and to carry heavy loads. 52. Table of the Bones. — 1. Skull, 28 (.1) AXIAL SKELETON Frontal Parietal Temporal Occipital Sphenoid Ethmoid Cranium, 8 Face, 14 Superior Maxillary Inferior Maxillary Palate Nasal Vomer Inferior Turbinate Lachrymal Malar Malleus Bones of Ear, 6 \ Incus Stapes 2. Hyoid, 1 3. Spinal Column, 26 4. Thorax, 25 Cervical Vertebrae Dorsal Vertebr£e Lumbar Vertebrae Sacrum Coccyx Ribs Sternum 1 2 2 1 1 1 2 1 2 2 1 2 2 2 2 2 2 1 7 12 5 1 1 24 1 48 CONSCIOUS NERVOUS OPERATIONS (/?) APPENDIC'ULAH SKKLKTOX 1 Clavicle Scapula 2 Humerus Ulna 2 Radius 2 Carpals Metacarpals Phalanges 16 '10 28 Os Innoininatum 2 Femur 2 Fibula 2 Tibia 2 Patella 2 Tarsals 14 Metatarsals 10 Phalanges 28 1. Shoulder Girdle, 4 2. Upper Extremities, 60 3. Pelvic Girdle, 2 4. Lower Extremities, 60 53. Cartilage. — In infancy a considerable part of the skeleton consists of cartilage, or gristle, which afterward becomes ossified. But there are cartilages — such as the external ear, the rings around the windpipe, and the ends of various bones — which do not ossify, and are known as permanent cartilages. Cartilage is a smooth white shining tissue of close texture, rarely containing blood vessels. It is made up, like the bones, of cells surrounded by the intercellular substance which is the product of the living cells. A thin layer of cartilage covers the surfaces of the bones which come in contact with other bones. Car- tilage also serves as padding in various parts of the body. 54. Connective Tissues of different varieties serve to com- plete the skeleton. They form the strong cords and bands and sheets called ligaments, for binding bones together, and the tendons which fasten the muscles to the bones. THE FRAMEWORK OF THE BODY 49 55. Articulations. — All unions between bones are called articulations. Some of these allow of more or less move- ment; others permit no movement at all. The bones of the skull, with the exception of the lower jaw and the minute bones belonging to the inner ear, have no motion. In most cases their union is formed by means of toothed edges which fit into each other, forming irregular lines known as serrated sutures. They lessen jar and avert injury to the brain. The different vertebrae of the spine have a very slight motion upon one another, due to the elasticity of the cartilage pads or cushions which sepa- rate them. 56. Joints, — Where two bones are articulated in such a way as to permit one bone to glide freely over the Fig. 34. — Ball and socket joint at hip. The parts are separated to show attachments of the round ligament. other, the union is called a joint. Joints are of vari- ous kinds and are adapted to various movements. MACY'S PHVS. — 4 50 CONSCIOUS NERVOUS OPERATIONS Fig. 35. — Hinge joint of the elbow. 1 hum eras. 2 ulna. The ball and socket joint seen in the shoulder and the hip has the end of one bone fitted into the hollow of another, and provides for motion in any direction (Figs. 34 and 86). The pivot joint is that in which one bone rotates round another, as in the atlas and axis joint (Fig. 32), already described, and in the rotation of the ulna on the radius at their junction with the wrist. The hinge joint permits of motion in one plane only, as in the joints of the fingers. Some hinge joints have provision for additional movements, as in the elbow (Fig. 35) and in the articulation between the lower jaw and the skull. 57. Synovial Membrane. — The broad, thin ligament sur- rounding a joint forms a closed sac. This sac is lined with the synovial membrane, which secretes a fluid whose purpose is to lubricate the joint, as oil lubricates the parts of a machine which move upon one another (Fig. 36). 58. Structure of Bone. — A living bone is tough, strong, and slightly elastic. Burned in a fire it retains its size and shape, but becomes brittle and easily crumbles to pow- der. Soaked for a few days in dilute muriatic acid, it also retains its shape, but becomes so flexible that, if one of the long bones, it may be tied in a knot. The fire destroys the 33 per cent of animal matter in the bone, leaving the calcium phosphate, calcium carbonate, and the small quan- tities of other salts which constitute the earthy or mineral portion of bone. The acid dissolves out the earthy salts and leaves the animal tissues. THE FRAMEWORK OF THE BODY 51 On examination, one of the long bones in a fresh condi- tion is seen to be covered at the two ends with smooth white articular cartilage, while the shaft is inclosed in a sheath of dense white fibrous membrane, called the peri- osteum, closely adhering to it. It is on the inner side of this membrane that the bone grows, and by it the nutri- Pelvic Bone Synovial Membrane Head of Femur -—Round Ligament - Capsular Ligament Fig. 36. — Section of hip joint. tion of the bone throughout life is assisted. If the peri- osteum is torn away, the bone dies. It covers every part of the surface of a bone not covered by the articular car- tilage, and into it the fibers of the ligaments extend, being so interwoven as to make an indistinguishable and insep- arable junction. 59. Inner Composition of Bone. — If the bone is sawn through from end to end, the shaft is found to be hollow, with the medullary cavity, as it is called, in the center, filled with yellow marrow. The walls of the shaft are of a dense, solid structure, except for a thin stratum of 52 CONSCIOUS NERVOUS OPERATIONS spongy bone around the cavity which contains the mar- row (lower end of Fig. 37). In the enlarged articular extremities of the bone, how- ever, the reverse is the case. The firm, compact part forms only a thin layer on the sur- face, the rest being a loose, spongy net- work of bony mat- ter, with the spaces filled with a soft, red substance called red marrow. Interlacing chan- nels, called the Ha- versian canals, run through the whole substance of a bone, in the densest as in the more porous parts. The perios- teum is richly sup- plied with blood ves- sels, and from them minute branches en- ter the bone itself and run along the Haversian canals. Other blood vessels reach the solid portion of the bone from within, through the medullary canal of the center of the bone, and thus nutriment is conveyed to every part. Each Haversian canal is surrounded by a set of hard bony plates, and between the plates, or lamellce, are little Fig 87. Longitudinal section of the upper end of the tibia. THE FRAMEWORK OF THE BODY 53 cavities called lacunce, connected by minute canals with one another (Fig. 16, p. 25). Within the lacunae and their canals are found little masses of protoplasm called bone cells, or bone corpuscles. These branching bone cells communicate with one another and with the blood vessels of the Haversian canals. They are the architects for building up the bony fabric. Each cell constructs walls which unite with those about them to form the solid mass. Along with the arteries of the interior bony structure very fine nerve fibers have been traced, and lymph vessels are found in connection with the blood capillaries within the substance of all bones. 60. Hygiene of Bones and Joints. — The science of hygiene has to do with all that promotes normal action of the various parts of the body and of the whole mechanism. In respect to the bones we need to consider first the con- ditions most favorable to their growth. 61. Bones of the Young. — As the skeleton grows it not only becomes larger, but also becomes changed in the struc- ture of its parts. The bones of an infant are almost wholly composed of cartilage, having the form of the completed bone, but the flexibility and elasticity of the cartilaginous tissue. They become slowly more firm and hard by a deposit of solid material furnished by the food, selected from the blood and lymph by the living cells in the carti- lage and the periosteum. As the bone is built up by the deposit of the salts of lime (chiefly calcium phosphate), which furnish nearly all the earthy part of bone, the carti- lage cells waste away, and their dead matter is carried oft' by the blood. 62. Importance of Proper Food. — It is evident that the food suitable for young children must contain lime and phosphorus in proper proportions for making bone. Milk 54 CONSCIOUS NERVOUS OPERATIONS has been found to furnish, in most digestible form, those substances and others needed by the human infant, and is for early childhood the complete and perfect food. If there is not a sufficient supply of earthy matter in the food, the bones of a child remain soft and weak, and are easily bent or deformed, as in the disease called rickets. For such conditions an abundance of suitable food, with plenty of fresh air and sunshine, supplies the cure. Alcohol and tobacco are particularly to be avoided dur- ing the time of growth, as they retard or prevent the full development of the bone cells, making the figure stunted and enfeebled. Cigarettes are especially harmful. 63. Deformity to be Guarded Against. — Children permitted to walk too early, before the bones are sufficiently hardened, may be made permanently bow-legged. They should be allowed freedom of movement and plenty of exercise, but should not be urged to walk too soon. Long-continued pressure upon the bones of children may result in deformity. Some tribes of Indians flatten the heads of their children by fastening boards upon them. Clothing should always be loose ; shoes especially should allow room for movement and growth. Care should be taken that a child should habitually assume correct positions in sitting and standing, and fre- quent changes of position are needful. The seat should not be too high to permit the feet to rest easily and squarely upon the floor, otherwise the bones of the thigh may be bent by the weight of the legs. School children should be taught to sit upright while writing or studying, lest the spine become curved and diseased. Seats and desks should be carefully adapted to the child's stature, and round shoulders — the most common deformity — should be especially guarded against THE FRAMEWORK OF THE BODY 55 In standing, the weight should be supported evenly by the two feet. One hip or one shoulder often becomes higher than the other, upsetting the firm poise of the figure by neglect of this precaution. A teacher should see that a child is not kept standing till wearied. 64. Bones of the Aged. — As the bones of the young con- tain an excess of animal matter, so those of the old have an excess of mineral substances and are consequently more brittle. The aged, therefore, need to guard espe- cially against fractures of the bones. Not only are their bones more easily broken, they are also healed with greater difficulty. In the young and healthy the vital processes are more actively carried on, and the busy bone cells go swiftly to work to repair a breakage, throwing out first around the injured parts a soft repairing material in which bony matter is afterward deposited ; but in the aged the bone cells work slowly, and a broken bone is sometimes never fully restored. 65. Broken Bones. — The two ends of a broken bone should be brought together into their correct position as soon as possible, before inflammation and swelling render this difficult. Of course a skillful surgeon should be called to " set " a broken bone ; but there may be delay, — the patient may have to be carried some distance. In such cases care should be taken to prevent injury to the surrounding parts from the fractured ends of the bone. A limb should be bound to a strip of board or even an umbrella as a temporary splint. 66. Injuries to Joints. — Dislocation of a joint stretches or breaks the ligaments and other membranes around it, producing inflammation. This renders examination and putting in place difficult, and a dislocated joint should therefore be restored to place as soon as the need- 56 CONSCIOUS NERVOUS OPERATIONS ful skill can be procured. A sprain, which is a sudden wrenching or straining of the ligaments not sufficient to dislocate the joint, is often as serious as a dislocation. Neither should be neglected or treated lightly. Inflamma- tion, if not checked, sometimes results in the destruction of the synovial fluid and the coverings of the ends of the bones in the joints, and consequently in permanent stiff- ness of the joint. Immediate and long-continued rest is imperative, and competent surgical advice should usually be secured. DEMONSTRATIONS AND EXPERIMENTS 10. The Skeleton. — For the study of the osseous system there should be accessible to the student a mounted human skeleton. In absence of this, a mounted skeleton of a cat or dog may be used. Where the school property does not include a skeleton of any kind, the enthusi- astic teacher will provide one. This can be very quickly done as fol- lows. Clean most of the flesh from the skeleton of a cat, dog, or rabbit ; boil the partly cleaned skeleton in " liquid soap," one part, and water, four parts, for forty minutes, then for thirty minutes in liquid soap and water, equal parts ; cool the skeleton in cold water ; clean with a brush and allow to dry. The liquid soap is made by dissolving 12 grams of saltpeter and 75 grams of white soap in a mix- ture of 2000 cubic centimeters of water and 150 cubic centimeters of strong ammonia. The skeleton may be studied without any attempt at mounting it. The student should follow the text of this chapter, identifying each bone as it is described. The teacher will find it profitable to have each student " demonstrate " the whole or a certain part of the skeleton, i.e. point out and name the various parts without any reference to the text and without leading questions from the teacher. 11. Cartilage. — At a meat market, bones can be procured which will show hyaline cartilage on their articular surfaces. At a slaughter- house can be obtained the windpipe, ears, costal cartilages, etc., of various animals, and the general appearance and purposes of the dif- ferent varieties of cartilage can be shown from them. To show the THE FRAMEWORK OF THE BODY 57 minute structure of cartilage, cut very thin sections with a razor from the articular surface of a fresh bone of a young animal, mount in normal salt solution, and examine with the compound microscope. 12. Structure of Bone. — Procure, at a meat market, a leg bone of some animal and compare it in appearance with a similar bone that has been exposed to the weather for months. Observe the pink color of the fresh bone, and the fibrous periosteum that covers it. Saw the two bones open lengthwise and observe the marrow cavity in each. Notice the compact shaft of each, and the cartilage on the articular surface of the fresh bone. 13. Minute Structure of Bone. — Mounted sections of bone may be procured of dealers in microscopical supplies, or the teacher may pre- pare them by sawing thin pieces from the shaft of a dry weathered bone and then filing them down till they are extremely thin. They may be mounted in water on a slide and examined with the micro- scope. But a better way is to dry the sections thoroughly, after care- fully washing them in alcohol, and then to mount them in Canada balsam that has been evaporated until it solidifies on cooling. The sections should be quickly placed in the hot balsam upon a clean slide, covered with a cover glass, and cooled to harden the balsam. 14. Composition of Bone. — Two pieces of the same fresh bone or two similar fresh bones should be obtained. Burn one piece in a fire for several hours till it turns completely white. All the animal matter has been removed. Place the other piece of bone in weak muriatic acid (10-15 per cent strength) for several days to decalcify. It becomes soft, owing to the removal of the mineral matter. Observe the brittle- ness of the burned bone, and the toughness and flexibility of the other piece. Place the burned bone in the muriatic acid, and burn the piece of decalcified bone. What is left? 15. Joints. — The various kinds of joints can be demonstrated on a skeleton. The actual movements that occur at those joints should be performed by the student in corresponding joints of his own body. 16. Dissection of a Joint. — Procure a leg joint of a sheep and show the possible movements of the bones that form the joints. Observe the tendinous attachments of muscles, also the ligaments that hold the bones together. Cut through the ligaments and open the joint cavity. Notice the synovial fluid, and the cartilage on the articular surfaces of the bones. Fig. 38.— The muscular system. 68 CHAPTER V THE MUSCULAR SYSTEM 67. Though the skeleton preserves the shape of the body, the muscles with the surrounding fat fill out the figure, giving roundness and grace of outline. Muscles and bones, with the tendons and ligaments connecting them, constitute the organs of motion and locomotion: they are the apparatus by means of which the nervous system acts when the object sought is movement; as the stomach, liver, blood vessels, kidneys, and other parts of the digestive system are the apparatus which the nerv- ous system uses for the purpose of nutrition. 68. The Muscles are the lean part of meat. They make up that part of the body which we call flesh. When a muscle is exam- ined, it is found to consist of small fibers bound to- gether in bundles (Fig. 39), each bundle being wrapped in a thin sheath of areolar tissue, called perimysium, while each minute fiber of which a bundle is composed has also its mem- branous sheath, called the sarcolemma. 59 Fig. 39. — Bundles of striated muscle cut across. ff several bundles bound together into larger bundles to make up the muscle. 60 CONSCIOUS NERVOUS OPERATIONS HUH Fig. 40. — Pieces of striated muscle fibers. Showing the spin- dle - shaped nuclei and cross-striations. At the left above is the rounded end of a fiber. 69. Classification. — The muscles are divided into two classes, usually called the voluntary and the involuntary muscles, the first being under the con- trol of the will, while the second are not. There is also a difference under the microscope between the two classes of muscles. Voluntary muscular tissue is composed of fibers which are marked by alternate dark and light stripes. They are called striated or striped muscular fibers (Fig. 40). The fibers which com- pose the involuntary muscles are, as a rule, destitute of these markings and are called plain muscular fibers (Fig. 41). Certain exceptions to the above rule should be noted. The muscles of the heart, though not under the control of the will, are made up of striped muscular fibers ; and also the muscular fibers found in the pharynx, part of the esophagus, and in the internal ear, though involuntary, have the structure of voluntary muscle fiber. On the other hand, the ciliary muscles, by which the eye is accom- modated for seeing objects at different distances, are under the control of the will, though composed wholly of plain or unstriped muscle fibers. Some striped muscles, like those of respiration and of the eyelids, are partly voluntary and partly involuntary. 70. Voluntary Muscles. — These are also called skeletal muscles, because they constitute the muscular apparatus attached to the bones. Each muscle is usually larger in Fig. 41. — Plain muscle fiber. N nucleus. THE MUSCULAR SYSTEM 61 the middle than at the ends, and the swollen middle portion is called the belly. The belly is usually unat- tached, but the two ends are secured to the bones by tendons which are continuous with the connective tissue of the muscle (Fig. 42). Between the small fibers of the bundles which make up a muscle is a little loose areolar tissue in which are distributed the blood vessels and nerves for the muscle. 71. Muscle Cells or Muscle Fibers. — It is in the microscopic threads of the muscle that the peculiar power of contraction lies. These are variable in length and thick- ness, but are said to average, in voluntary muscles, -5^ of an inch in diameter and about one inch in length. They are cylin- drical in shape, with rounded ends (Fig. 40), and as a rule do not branch. In the muscles of the face and tongue, how- ever, the muscle fibers divide into many Fi 42 _Bice branches. muscle. Each muscle fiber, or cell, consists of the 1 tendinous ends. sarcolemma and a soft, semifluid material of alternate light and dark disks, called the contractile sub- stance. Just beneath the sarcolemma are several long oval nuclei. 72. Nerve Endings in Muscle Fiber. — It is impossible to treat of muscles and their action without including some study of the other sort of irritable tissue, nervous tissue, upon which muscular action depends. The sarcolemma of each muscle fiber is pierced by a branch of a nerve fiber. The primitive sheath or neuri- 62 CONSCIOUS NERVOUS OPERATIONS Nerve Fiber lemma (the inclosing membrane of the nerve fiber) becomes continuous with the sarcolemma, and the axis cylinder of the nerve fiber branches many times, the ramifications end- ing in a flat or branched layer °^ protoplasm containing nu- clei. These terminal nerve organs are called end plates (Figs. 43 and 44). 73. How a Muscle Contracts. Fig. 43. -Nerve endings in stri- ~ A muscle contracts or be- ated muscle fibers. comes shorter in proportion to its length, without change in bulk, under the influence of stimulus. It is the office of the nerves of the muscles to carry to them their natural stimulus, but muscles also contract under the action of other stimuli; for in- stance, in consequence of a sudden blow or pinch, when heat is applied sud- denly, when certain chemi- cal substances are dropped upon them, or when an electric shock is conducted to them. In living animals the muscles are always more or less contracted. This Fig. 44. —Motor nerve endings in striated muscle fibers. is due to the nervous influence which they constantly receive. If their nerves are cut or destroyed, the mus- cles lengthen. This tension of the muscles keeps them ready for immediate action. When a nervous impulse THE MUSCULAR SYSTEM 68 reaches a muscle, the numerous branches of the nerve — one for each individual muscle cell — distribute the stimulus to all parts at practically the same instant. The end plates are situated in each muscle fiber near the middle between the two ends, and the fiber begins there to contract. The two ends draw toward each other ; the fiber becomes swollen and shorter. In the muscle as a whole contraction is simply the sum of the con- tractions of all the minute fibers which compose it. In the muscle fiber contraction appears to be some compli- cated movement of the molecules which produces a change in the appearance of the stripes. It is found that in those places where swift and rapid contraction is called for, the muscular tissue has almost invariably the striped fibers. 74. When a muscle is made to contract by a single electric shock, or by other artificial means, the movement is sudden and brief. Voluntary muscle, however, under its natural nervous stimulus, never contracts with a twitch. Its action is rather that of continued gentle vibration, called tetanus, such as follows a rapid series of electric shocks which leave no time for relaxation. The nervous stimulus comes to the muscle in a quick succession of impulses, — about twenty in a second, — so that one vibration is succeeded by a second before the first has ceased to agitate the muscle cell. 75. Internal Changes in Muscle under Stimulus. — Some of the energy set free by a contracting muscle appears as work done, weight lifted, etc., while a considerable amount becomes heat, for the temperature of muscle always rises under contraction. Certain chemical changes also ap- pear. Variable amounts of carbonic acid and lactic acid are set free, and oxygen is used up. Electric changes 64 CONSCIOUS NERVOUS OPERATIONS lasting for some time are also produced in the muscle by its contraction. These are shown by the use of a delicate galvanometer. 76. Another important effect upon itself of a muscle's contraction is what we call fatigue of the muscles, that is, a lack of readiness to respond to stimulus. This is due to the using up of the material in the muscles which was available for the production of energy, and still more to the accumulation of waste matter — the product of the activity of the muscles. Experiments have shown that it is not the muscle itself which first becomes too much fatigued for contraction. Nor is the seat of fatigue in the nerve, but in the end plate within the muscle cell. The fatigue is relieved by even a brief rest, and such relief is absolutely necessary to the health of the muscles. Even the muscles of the heart, that organ which works cease- lessly from the beginning of life to death, have a period of rest after each beat. 77. External Effects of Muscular Contraction and Relaxation. — The purpose of muscular contraction is the production of motion. The contraction and relaxation of the muscular walls of the heart keep the blood in constant movement ; the various other vital processes are also dependent upon more or less constant motion in the tissues and organs of the body, and all our outward activities are likewise the results of the shortening and lengthening of the innumer- able strands of muscular tissue. Muscular power, or the amount of force which a mus- cle can supply, varies with its health and vigor, and with its form. The thickest muscles can lift the heaviest load. Those having the longest strands can move a weight the greatest distance. Hence the human body possesses both long, slender muscles, and short, stout ones, . THE MUSCULAR SYSTEM 65 as well as those of all sizes and lengths between. Many muscles which we might at first think to be long are really short, but appear long because of the long tendons by which they are attached to the bones. Many of the muscles which move the fingers, for example, have their bellies in the forearm, and are attached to the small bones of the fingers by long, slender tendons. The force with which muscles contract is sometimes very great. A human muscle one square centimeter (.15 square inch) in section can raise a weight of 5670 grams or 200 ounces. When a muscle contracts, its two ends are drawn nearer together, and hence draw toward each other the parts to which the ends of the muscle or its tendons are attached ; the belly of the muscle becomes swollen, and in strong contraction the whole muscle becomes tense and hard. 78. Contraction is speedily followed* by relaxation. The stimulating force ceases to be supplied, and the muscle returns to a state of rest. If the hand has been raised by the contraction of the biceps muscle on the inner side of the humerus, it will fall under the action of gravitation when that contraction ceases and the muscle becomes passive. In order that the hand may be drawn down with force the action of the triceps muscle at the back of the humerus is needed. Muscles can give a powerful pull, but they cannot push. Very generally, therefore, they are arranged so that muscles which cause movement in one direction are opposed by those which cause movement in the opposite direction. 79. Dead Muscle. - - The muscles of a dead body, or muscles which have been removed from a living body, gradually undergo a marked change, which results in the stiffening known as rigor mortis. That which was translucent becomes more opaque, most of the natural MACY'S PHYS. — 5 66 CONSCIOUS NERVOUS OPERATIONS elasticity disappears, and a hard, rigid condition sets in, accompanied by more or less contraction. This is due to a coagulation of the protoplasm of the muscle cells, simi- lar to the clotting of blood. Rigor mortis passes away after a time, and the dead body becomes soft and flabby — a sign of approaching decay. 80. Plain or Involuntary Muscles. — Though the plain muscles are not under the control of the will, they still have nervous connection with the central nervous system. Most of the nerves supplying the organs having plain or umtriped muscular tissue come from the sympathetic nerv- ous system; but from every ganglion of the sympathetic chain nerve fibers communicate with the brain and spinal cord. The muscles of the blood vessels, lymphatics, glands, and other internal organs, are of unstriped struc- ture, and carry on their work without affecting conscious- ness. Their action under stimulus is similar to that of the skeletal muscles, but takes place much more slowly. 81. Plain Muscle Fibers. — Plain muscles are made up, like skeletal muscles, of bundles of fibers, and these of muscle cells. The cells, however, differ from those of the voluntary muscles. They are long, spindle-shaped fibers, having a rod-shaped nucleus in the center (Fig. 41). The nerves of plain muscle fibers do not end in end plates, but form plexuses, or networks, which ramify between and around the muscle fibers. The nerves of the heart mus- cles end as do those of the unstriped muscles. 82. Rhythmic and Peristaltic Movements of Involuntary Mus- cle. — One of the characteristics of involuntary muscles is a tendency to alternate regular periods of activity and rest. The heart is the most familiar illustration of this rhythmical tendency, but it is seen in some other organs, and especially in some of the lower animals. THE MUSCULAR SYSTEM 67 The peristaltic action of plain muscle is seen in the small intestine and in other parts of the alimentary canal. When any part of the tube is stimulated, a circular con- traction results, which slowly passes along in a wavelike manner through the length of the tube. In the diges- tive tract this movement serves to drive the food onward. 83. Involuntary muscle, as a rule, contracts more slowly than voluntary muscle. It contracts, not with a tetanus like that of voluntary muscle, but with a single, much prolonged contraction. 84. Mechanism of Movement. — The power of the muscles to change their form carries with it the power to change the positions of the bones and other parts of the body to which they are attached, and hence to change the positions of the different parts in respect to one another and to move the whole frame from place to place. When a part of the body is moved at a joint, the bone which is moved acts as a lever. A lever is a stiff bar which can be moved round a fixed point, or fulcrum. Three classes of levers are known to the science of mechanics, depending upon the position of the fulcrum with reference to the weight to be moved and the power which produces the motion. In the first class the fulcrum is between the weight and the power, as in using a crowbar to lift the edge of a stone. In the second class the ful- crum is at one end, the power at the other, and the weight between them. In the third class the fulcrum is at one end, the weight at the other, and the power is applied between the two. All three forms of levers are found in the human body, though the levers of the third class are the most numerous (Fig. 45). 85. Lever of the First Class. — The action of a lever of the first class is seen in the straightening of the bent arm. 68 CONSCIOUS NERVOUS OPERATIONS The muscle at the back of the humerus applies the power. It is attached by tendons to the scapula and to the hinder side of the humerus, while the tendon into which the lower end of the muscle narrows is inserted into the end of the ulna at the elbow, which is more than an inch above the articulation of the ulna with the humerus. By its contrac- tion the muscle pulls the upper end of the ulna upward, draw- ing down the hand, which is the weight at the lower end of the ulna, and straightening the joint. The fulcrum is at the elbow joint, between the hand and the power at the upper end of the ulna. / \ \ \ \ \ in Fig. 45. —Diagram of the foot, illustrating levers of the three classes. I tapping the toe on the floor. II rising on the toes. Ill lifting a weight with the toes. 86. Lever of the Second Class. — When the body is raised on the toes (Fig. 45, II) the action of a lever of the second class is seen. The weight is that of the whole body sup- ported by the foot at the ankle, while the power operates through the muscles of the calf of the leg at the heel, the toes acting as the fulcrum. 87. Lever of the Third Class. — When the body lying on the ground is raised to a sitting posture, a lever of the third class is used. The head and body are the weight, the fulcrum is at the hip joints, and the power is applied THE MUSCULAR SYSTEM 69 between the two by the muscles which pass from the front of the thigh to the hip bones. The raising of the hand by bending the elbow joint is perhaps a clearer example of the lever of the third class. The weight is at the end of the forearm, the fulcrum is at the elbow, and the power is between, at the point on the radius where is inserted the muscle which lies on the front of the humerus. 88. Coordination of Muscular Action. — Our ordinary movements involve the use of many different muscles, and very com- plicated action of levers and cords. Even simply to stand erect requires strong ten- sion of certain muscles and ligaments pulling against one another. The mus- cles on the front of the thigh contract to keep the knee from bending, while the ligaments of the joint prevent it from bending the wrong way. The muscles on the front of the leg contract to keep the body from falling backward, and those at the back contract to keep it from fall- ing forward. In the same way the trunk is balanced on the thigh bones by the muscles passing from the body to the thigh in front and back, while a particu- Fig. 46. — Diagram larly strong ligament, crossing the hip £us'°™ °^ joint from the pelvis to the thigh bone, tend to keep the keeps the extra backward weight of the body e trunk from destroying the balance of the frame. At the back of the neck are the muscles which give to the head its erect and graceful poise, while many ligaments bind it to the spinal column (Fig. 46). 70 CONSCIOUS NERVOUS OPERATIONS 89. In order to maintain all the nice adjustment and balance of muscular force constantly demanded, the brain and nerves must be continually at work. Just enough stimulus must be supplied to each set of muscles and supplied at exactly the right moment or something will at once go wrong. This harmony of muscular ac- tion and regulation of the complex relations between the hundreds of muscles in the body is called coor- dination. It is easy to show that it depends upon the nervous system. If one falls asleep or receives a blow which " stuns " the brain, the muscles are relaxed, and, unless supported, the body falls to the ground. Sudden nervous or emo- tional excitement, as surprise, grief, or fear, may cause the muscles of the heart to stop their action and the body to fall in a " faint." In some cases the effect of sight upon the brain is to destroy the power to control the muscles, as when the sight of the moving waves of lake or ocean renders one giddy. The perception of certain odors may have the same effect, and in many other ways the control of the muscles is affected by that which affects the central nervous system. The study of the brain has shown that the cerebellum is the great center for the coordination of muscular move- ment and especially of those muscular actions which have to do with maintaining the equilibrium of the body. 90. Exhaustion of Muscles. — Even when we are quite awake, and the brain is active, our muscles sometimes refuse to act. Muscular fiber cannot contract continuously for a long time. It must have periods of rest. That is the reason we require frequent changes of position, one set of muscles being thus allowed to rest while another set is called into action. If a weight be held out at arm's THE MUSCULAR SYSTEM 71 length from the shoulder, the muscles of the arm soon become exhausted and incapable of sustaining the weight. But a moment's rest restores the contractile power, and the weight may again be held out. 91 . Muscle Waste. — When a muscle contracts certain chem- ical changes take place in the substance of its cells. Some of the matter in the muscular fiber becomes oxidized, and new substances are formed which are harmful to the body if not removed. These are called waste products, and that which appears in largest quan- tity is carbon dioxide. These waste products are taken up by the blood which flows along the muscle cells and are finally removed from the body by means, mainly, of the lungs and the kidneys. If the waste matter is not removed, the effect soon appears in the cen- tral nervous system, to which the poison is carried by the blood. 92. Voluntary Movement. — Let us suppose that a man seeing an apple within reach puts forth his hand to take it (Fig. 47). In such a case the light from the apple enters the organ of sight and stimulates the nerve endings Fig. 47. — Diagram of the path of a nervous impulse which re- sults in the hand reaching to seize an object seen with the eye. G cerebrum. Cb cerebellum. M medulla oblongata. MC motor center in brain. OC optic center in brain. P pons Varolii. Q corpora quadrigemina. S spinal cord. T optic thalami. 72 CONSCIOUS NERVOUS OPERATIONS appropriated to vision. The nervous irritation is con- ducted by the optic nerve to that part of the brain con- cerned in perception by means of the eye. The nervous center is affected, and the nervous impulse is passed on, by some unknown process which we speak of as the action of the will, to the nerve fibers running down the white columns of the spinal cord. These convey the impression to the anterior horn of the gray matter of the cord, where lie the motor cells from which arise the motor roots of the spinal nerves. From the motor cells a new nervous impulse goes forth to certain muscles of the arm and hand. The muscular cells of such muscles contract, the bones are moved at the joints, and the apple is seized. This is voluntary muscular action. 93. Reflex Movement. — The voluntary muscles often act without receiving any nervous impulse from the brain, Fig. 48. — Diagram of the path of a simple nervous reflex action. and without any conscious purpose. Suppose the man in putting out his hand to take the apple is stung upon the finger by a wasp not before perceived. The end organ of sensation in the finger sends the impression of paiu THE MUSCULAR SYSTEM 73 along the afferent sensory nerve through the posterior nerve root to the cells of the spinal cord, and an impulse is at once sent forth along efferent motor fibers to the muscles of the hand and arm, which promptly jerk the hand away (Fig. 48). A great multitude of reflex actions are possible to the muscles, and one of the chief functions of the spinal cord is to act as a center of reflex action. 94. Automatic Movement. — Action of the muscles often occurs without any obvious stimulation of the nerves from without. That is, the nervous impulse may apparently arise in the nerve center itself, and efferent, or outgoing influences are not preceded by afferent, or incoming influ- ences. Such changes often, occur rhythmically — activity and rest, or diminished activity, following each other in regular alternation — as in the movement of the muscles of respiration ; and this characteristic is believed to be due to certain rhythmic changes which take place in some of the nervous material of the medulla oblongata. 95. The Muscular Sense. — It is not motor nerve fibers only that are distributed to the muscles ; the muscles receive sensory fibers also which pass to the posterior roots of the spinal nerves and convey impressions from the muscles to the spinal cord and thence to the brain. These impressions are called the muscular sense. They assist our judgments of weight, and inform the brain of the general condition of the muscles. 96. Hygiene of the Muscles. — Muscles increase in size and in strength by appropriating suitable material from the food, and by use. If ill fed and inactive, they become small and weak. If one limb is made useless from disease or injury, — as when a bone is broken, — its muscles shrink and grow soft, so that the unused limb 74 CONSCIOUS NERVOUS OP K RATIONS becomes perceptibly smaller, and, of course, weaker than its fellow. 97. Exercise is indispensable to health of muscles. Growing children need much active exercise for develop- ing and hardening the muscles, and all healthy children crave it. Nothing is better than running and walking for promoting the growth of the muscles, for developing the power of the lungs and the heart, and so for aiding both the free circulation of the blood and its purification, by means of which the nutrition of the whole body is stimu- lated. Those who exercise much in the open air (which is always best) have, as a rule, good appetites, for food is needed to repair the waste caused by the exercise. Many diseases are prevented, and some are cured, by suitable exercise. A brisk walk of several miles taken regularly every day would alone do much to keep the whole body in normal condition. If it is impossible to go out of doors for the needful amount of exercise, the indoor conditions should be as nearly as possible like those without. Fresh air and light should be freely admitted to the rooms used, additional clothing being put on when necessary. Rowing, swimming, boxing, horseback riding, climbing, sweeping, cycling, etc. are of value in strengthening the muscles of the limbs, chest, and back. A large, strong chest, wherein the lungs have plenty of room for an abun- dance of pure air, is not likely to belong to a consumptive person. It is desirable that exercise should be chosen which develops both sides of the body. Throwing a ball with one hand, if indulged in to excess or without sufficient exercise of other sorts, sometimes causes irregular devel- opment of the body, and curvature of the spine may fol- THE MUSCULAR SYSTEM 75 low. Certain kinds of spinal curvature are cured by wisely chosen exercise. 98. Exercise in the Cold. — For a healthy person nothing so well develops the whole system and hardens the con- stitution as regular, vigorous, and agreeable exercise in the cold. Hence outdoor winter sports and occupations should be encouraged. Skating, sliding, snowballing, and swimming are excellent as promoters of health. 99. Time for Exercise. — Some times are better than others for taking exercise. The morning is usually best for the severer forms, because the whole system is then refreshed and vigorous. In the evening one who has been engaged in physical toil does not need exercise, but rest ; while one whose occupation is mental labor or sed- entary business will be rested and refreshed and prepared for sound sleep by exercising judiciously in the open air after the day's work is done. When the muscles are called into use they require more blood than when at rest, that the waste which results from exercise may be repaired. This extra supply of blood is drawn from other parts of the body, and the demands of the muscles may retard the performance of other physiological functions — since the total amount of blood is practically invariable. This is why the muscles should not be vigorously exercised for an hour or two, at least, after meals, and not immediately before. Digestion requires an increased flow of bloo.d to the alimentary canal and digestive glands, and if the process is impeded by a drain of the vital fluid to other parts, harm will result. 100. Training. — The scientific development of the mus- cular system under " training " for particular purposes of sport, or in a well-equipped gymnasium, may have excel- lent results for those who are able to avail themselves of 76 CONSCIOUS NERVOUS OPERATIONS such facilities. But the number of such persons is com- paratively small, and as those who exercise in a gymna- sium are usually, and ought always to be, under the guid- ance of a qualified instructor, advice as to the use of the gymnasium apparatus is not needed here. 101. The Healthf illness of Work. — While all due stress should be laid upon the healthfulness of recreative exer- cise, it should not be forgotten that the human machine is the most skillfully designed and constructed apparatus ever made for accomplishing an immense variety of differ- ent kinds of work. The man whose daily employment brings into play his various muscles under conditions of reasonable comfort, and without overfatigue, may live a healthful life without paying any attention to the preced- ing suggestions. If that employment is carried on in the open air, and is such that he finds interest and enjoyment in it, he is still more fortunate. All the good effects of the most carefully devised systems of physical culture may be gained from a judiciously varied scheme of work, and the pleasure of being able by one's own effort to create some useful or beautiful or worthy product for the enrichment of the world may be a tonic even more health- fully stimulating than the most successful athletic con- test. The varied round of household duties, sometimes prescribed by a wise and skillful physician, has in many cases brought health to a feeble, languid, ailing woman. The effort which the idle rich man sometimes puts forth in the way of exercise, that he may secure an appetite for his dinner, would be still more promotive of health if turned to some useful purpose. The man whose business does not permit the proper activity of all the muscles must necessarily give thought and time to supply the deficiency. But to the great mass THE MUSCULAR SYSTEM 77 of mankind it is happily possible to get all needful exer- cise while doing useful work. 102. The Nervous System as involved in Muscular Exercise. — We have already learned that contraction of voluntary muscular tissue depends upon the stimulus brought to each minute muscle cell by a nerve fiber. Without such stimulus a man's limbs are motionless and the whole frame a lifeless mass. If the nervous system is enfeebled by disease or by exhaustion, the action of the muscles becomes weakened or deranged. The disease called Saint Vitus's dance, Avhich causes muscular movements beyond the con- trol of the will, is not a disease of the muscles, but of the nerves. So in other disorders which derange the action of the muscles, the real trouble is seldom with the muscu- lar tissue itself. The direct effect of muscular activity, as suggested in section 91, is to poison the nervous cen- ters. The greater the demand upon the muscle in the way of rapid and frequent contraction, the greater the con- sumption of living material and the greater the amount of poisonous, dead, waste matter which passes into the circu- lation. These waste products, if not promptly removed from the system through the excretory organs, are found to have a powerful injurious effect upon the central nerv- ous system, an effect that is soon manifest in the weak- ened action of the muscles themselves. Anything, therefore, which affects injuriously the nerv- ous system interferes with the free and easy play of the muscles. And, conversely, anything which promotes a high level of health in the nervous system is an aid to mus- cular vigor also. Exercise undertaken for the carrying out of some worthy purpose — a purpose in which the mind is deeply interested and the whole man engaged — is the most healthful exercise ; while that which is disliked 78 CONSCIOUS NERVOUS OPERATIONS or entered upon indifferently and listlessly is found to have little or no invigorating power. 103. Effect of Alcohol and Other Stimulants and Narcotics upon Muscular Action. — The most serious effects of the excessive use of alcoholic drinks, tobacco, opium, chloral, and other narcotic drugs are felt by the nervous system and will be most fully treated when we come to the special study of that part of the human organism. But it is well to notice here how those substances influence the organs of motion. No one who has ever seen a drunken man in the stage pre- ceding that of stupor can have failed to observe the uncer- tainty of his muscular movements : the shaking hand, the staggering gait, the thick, indistinct utterance. These effects are due to what is called the excessive use of alco- holic drinks, and no one doubts that in large quantities they act injuriously upon the system. Alcohol deranges the action of the muscles by its influence upon the nervous system, causing defective regulation of the supply of nerv- ous force to the several muscles. As to whether it is pos- sible to use alcohol in small amounts without impairing the perfection and vigor of muscular action, there is one very significant fact: that men in training for severe mus- cular exertion in athletic contests are strictly forbidden the use of alcohol in any form and in any quantity, whether or not they have been previously accustomed to such indulgence. As. the rules for such training are the result of long and wide experience and most careful study, it is safe to conclude that alcohol at least does not promote strength, endurance, or precision of muscular movement. It is very common for a person accustomed to a moder • ate use of alcoholic beverages to suffer from tremor of the hands, due to lack of control over the muscles, so that THE MUSCULAK SYSTEM 79 he is disabled from manual work requiring dexterity or skill. Sometimes an additional glass of liquor seems to steady the hand for a time, but tne shakiness soon returns. 104. Tobacco and other narcotics also affect muscular activity through their effect upon the nerves. All nar- cotics have as their natural, characteristic influence the paralyzing of some of the nerve centers. As medicines they may give relief from pain and so act beneficently under skillful application. Tobacco has a special effect upon the nerve centers regulating the action of the muscles of the heart, making that action irregular and less vigorous. This is particularly true of the young, and it is not very uncommon for boys addicted to excessive cigarette smoking to develop serious disease of the heart, or even to die suddenly from " heart failure." Smoking tobacco is found to interfere with work requiring fine and delicate adjustment of muscular movement, as in watch making and other delicate mechanical employments, in scientific drawing, fine penmanship, etc. It is also for- bidden to those persons in training for athletic contests, and to all pupils in many schools, as well as to soldiers in the armies of certain countries. DEMONSTRATIONS AND EXPERIMENTS 17. General Structure and Properties of Muscles. — It is important that the pupils see muscles in their natural positions and connections. For this purpose the frog is convenient, since the animal is so small that little dissection is necessary, and since entire muscles can be observed and owing to their great vitality can be made to perform their natural movements. The frog should be pithed or decapitated, and the skin removed from one of the hind legs. The muscles of the limb then stand out distinctly (Fig. 7, p. 18). The belly of a muscle can be distinguished from its tendinous ends, and the origin 80 CONSCIOUS NERVOUS OPERATIONS and the insertion can be made out. The respective share wnich each muscle takes in the movements of the limb can be shown by touch- ing the individual muscles with the electrodes of a weak galvanic battery. The change in shape of the muscle during contraction will also be very well shown. 18. Relaxation and Contraction of Muscles. — Extend one arm nearly straight from the shoulder. Then bend the arm at the elbow, draw- ing the hand up to the shoulder. With the other hand can be felt the changes in the form of the biceps muscle, as it relaxes and con- tracts. If the arm be bared the changes in form of the muscle can be seen. The tendon of the biceps at the elbow can also be seen. 19. Gross Structure of Muscle. — Obtain a piece of boiled corned beef, and dissect it with needles. The larger and smaller bundles of muscle fibers can be easily differentiated. By aid of a lens even the separate fibers can be isolated. Observe between the bundles and fibers the whitish connective tissue, in life tough and fastening the bundles and fibers together, but now softened in boiling. 20. Functional Difference between Voluntary and Involuntary Muscle. — If the abdominal cavity of the frog experimented upon in a preced- ing section be opened and the electrodes applied to the stomach and intestines, the difference between the movements of voluntary and involuntary muscle will be clearly demonstrated. 21. Fatigue of Muscle. — On applying repeated electric shocks directly to a frog's muscle, or indirectly through its nerve, the re- sponses are seen to become more and more feeble. But after a period of rest, the muscle responds as vigorously as ever. 22. Nerve Endings in Muscle. — While the tracing out of the final nerve ends in muscle must be left to the histological expert, yet the general relations of motor nerves to muscles can be very easily shown in the frog. Branches of the sciatic nerve can be traced out to the muscles of the leg (Fig. 8, p. 18), and by electrical stimulation the functional relation between nerve and muscle can be shown. 23. Rigor Mortis. — Observe in a frog, just after decapitation, that the muscles are soft and relaxed. Shortly after the tissues have com- pletely lost their vitality, the muscles will be found to be hard and contracted. The limbs which may have been bent at the joints and limp, are now straight and rigid. Death rigor or rigor mortis has set in. After some time, the muscles again become flaccid and putrefac- tion soon begins. Place fresh muscle in hot water, and observe that THE MUSCULAH SYSTEM 81 heat rigor is at once manifest. In both cases, the rigor is due to coagulation of the muscle substance. 24. Minute Structure of Voluntary or Striped Muxcle. — With needles tease out, in normal salt solution, on a glass, a small piece of skeletal muscle of a frog or other animal. Mount and examine with the com- pound microscope. It will be seen to be composed of elongate thread- like bodies, tapering (when not broken) at the ends. Some fibers will show cross markings. 25. The Minute Structure of Involuntary or Plain Muscle. — Tease out, as in the foregoing, a small piece of the outer wall of the intestine of a frog or cat. Here the fibers are seen to be spindle-shaped cells, much shorter than the striped muscle fibers. The fibers of both striped and plain muscle can be much more easily teased apart if the tissue be kept in a 20 per cent solution of nitric acid one to two days. 26. Cross Section of Muscle. Minute Structure. — If a prepared cross section of a small skeletal muscle can be obtained, the internal structure of a muscle can be very well shown by aid of the compound microscope. 27. Minute Structure of Tendon and Ligament. — Carefully tease out in normal salt solution, on a slide, a small piece of a thin tendon from the tail of a mouse. To obtain the tendon, cut off the tail from the body, and then pull out the delicate tendinous threads from the cut end. The tendon should be mounted immediately after removal, to avoid drying. A convenient method of mounting is to stretch the tendon across a slide, through a drop of normal salt solution in the center, allowing the ends of the tendon to adhere to the dry edges of the slide. On examining with the compound microscope the tendon is seen to be composed of wavy bundles of fibers. If the preparation be treated with a one per cent solution of acetic acid, the fibers will swell and disappear from view, but there will appear, between the bundles, rows of spindle-shaped cells. MACT'S PHYS CHAPTER VI THE SKIN AS AN ORGAN OF SENSATION — TOUCH 105. Functions of the Skin. — The whole body is covered with a flexible, elastic membrane of complex structure, which serves several different purposes. It envelops and protects the inner, soft parts, and especially the ends of the nerves. It is one of the three principal channels by which the waste products of the body are removed — that is, it is an organ of excretion. It regulates the temperature of the body by controlling the loss of heat through general radiation and evaporation, as well a§ by the direct action of the sweat glands in excretion. A small amount of respiration, or exchange of gases, also goes on in the skin, and it contributes by its general char- acteristics and its various modifications to the ornamen- tation of the body. 106. Structure of the Skin. — The present chapter has to do with the skin as one of the organs by means of which the nervous system is brought into direct communication with the external world, that is, as the seat of the sense of touch and of certain other allied nervous impressions. Two distinct layers are found in the skin, called the epidermis, or cuticle, and the dermis, corium, or true skin (Fig. 49). The epidermis is composed of epithelial tis- sue, or epithelium, and its cells lie in layers one above another, the outer or horny layer of cells being flat, 82 THE SKIN AS AN ORGAN OF SENSATION 83 dry, and unnucleated, or dead. The epidermis contains no blood vessels, but in the deepest layers are found minute terminations of some of the nerve fibers; and in the same layers are the fine granules called pigment which give color to the skin. This coloring matter is power- fully affected by sun and wind, causing tan and freckles. Sweat-Duct Sebaceous Gland Horny layer < Pigment layer Epidermii Dermis Fig. 49.— Diagram of section of the skin. 107. Hair and Nails are peculiarly developed forms of the cuticle. Each hair is a long filament growing obliquely from a little bulb called a papilla, lying in a hollow called the hair follicle, which reaches down below the skin into the areolar tissue beneath (Fig. 49). The part of a hair buried in the skin is called its root; the remainder, the stem, which tapers to a point. The stem is covered with scales overlapping like the scales of a fish and project- ing toward the point. When a hair is pulled out by the root, a new one will grow again so long as the papilla is uninjured. Each hair contains pigment granules which 84 CONSCIOUS NERVOUS OPERATIONS give its color. There are also cavities filled with air. These small bodies of inclosed air are very abundant in white hair, and, reflecting the / light, are the cause of its color, just as the whiteness of snow is due to the many reflections of light from the tiny bits of ice which compose it. In the nails the horny layer of the epidermis is greatly developed. Each nail has its root firmly embedded in a groove of the cuticle. The under side is fixed to the der- mis except at the end of the tinger or toe, where there is a free edge. The hair preserves the temperature of the head and is also a protection against injury. The nails protect the sensitive ends of fingers and toes and aid in their mechanical operations. Both hair and nails also help in the ornamentation of the body. 108. Mucous Membrane. — Not only is the whole outer surface of the body covered with the skin, but all the inner cavities and passages which have an external com- munication are also lined with it. These inner linings are, however, of a modified form of skin called mucous membrane, because it secretes a viscous fluid, or mucus. This membrane is thinner, redder, and more sensitive and delicate than the outer skin, but is of the same general composition ; that is, it has the two layers, the outer bloodless and insensible, the inner highly sensitive, soft, fully supplied with blood vessels, glands, etc. 109. The Dermis, or True Skin, is a close network of connective tissue fibers, forming a dense, tough, firm envelope for the body, resting upon and gradually pass- ing into the areolar tissue beneath, which is a loose network of interlacing bands and cords with meshes between. Ramifying through the derm is are nerves, THE SKIN AS AN ORGAN OF SENSATION 85 blood vessels, and lymphatic vessels, and it contains great numbers of sweat glands and oil glands. The epidermis, lacking blood vessels, does not bleed, and the horny layer, lacking nerves, has no feeling; but so fine are the networks of blood vessels and nerves in the dermis that the finest needle cannot pass between them. The whole surface of the dermis is thrown into innumer- able projections called papillce, many of them supplied with capillary blood vessels and nerve fibers. 110. Sensation. — When we become conscious of receiv- ing an impression, — that is, when we perceive that some part of our nervous system is stimulated, — we have what is called sensation. It has already been shown that nerv- ous stimulation may affect parts of the nervous system and reflex action may follow without conscious reception by the brain of any influence — that is, without sensation. In order that there may be sensation there must be (1) a stimulus, (2) a nervous end organ suited to receive the stimulus, (3) a path to the brain for the impulse excited by the stimulus, (4) a part of the brain to receive the impulse. Still another condition of a different sort seems essential to sensation, and that is an attitude of the mind which we call attention. A person absorbed in thought may look upon an object without being conscious of perceiving it, may hear music without knowing it. That is, all the con- ditions of sight and hearing may be present save the one of attention, for which we have as yet no physiologically descriptive terms. 111. General and Special Sensations. — There are various vague, indefinable feelings which are not referred to any particular portion of the body or to any external influence, and which we know as general sensibility. Sensations of fatigue, of restlessness, languor, weakness, and the like, 86 CONSCIOUS NERVOUS OPERATIONS are of this sort. It is supposed that these are associated with the ramifying, interlacing plexuses of nerve fibrils in many parts of the body, and are not due to the excitation of specially constructed nerve endings. Other sensations are more definite. We judge them to be caused by some influence acting upon particular parts of the body. What have long been known as the five senses are of this sort. But we now recognize, in addi- tion to touch, taste, smell, sight, and hearing, other sensa- tions apparently distinct from them, for which, in some cases at least, special nerve endings are provided, and spe- cial brain centers. These are sensations of temperature, of pain, of hunger, and of thirst, and the muscular sensa- tions previously mentioned (§ 95). 112. The Sense of Touch. — Touch has been defined as a sense of pressure referred to the surface of the body. It is that sort of impression upon the nervous system which gives information respecting certain properties of bodies in contact with the skin or mucous membrane. Through it we learn whether an object is hard or soft, rough or smooth, and other particulars, some of which are also given by the sense of sight. The skin is supplied with a variety of special adapta- tions which constitute it the organ of touch. 113. The Nervous Apparatus for Touch. — The thirty-one pairs of spinal nerves contain the fibers for feeling for the larger part of the body, as well as most of the motor nerve fibers supplying the muscles. It will be remem- bered that each spinal nerve arises by two roots from the spinal cord, one root containing afferent or sensory fibers, the other efferent or motor fibers. By the union of the two roots the nerve is formed, which thence contains both sorts of nerve fibers, as do its large branches. THE SKIN AS AN ORGAN OF SENSATION 87 114. In the neck, loins, and pelvis adjacent nerves interlace with one another to form a plexus, or network. In a plexus nerve fibers from two or more nerves are brought into connection in such a way that the parts of the body which receive nervous fibers from the plexus have communication with a greater number of nerve trunks and nerve centers, and may receive more complex im- pulses, than do those parts to which branches go from but a single nerve trunk. This is why these plexuses are so frequently found in connection with the nerves going to the limbs, where great complexity of motion and careful coordination are required. The nerves of the skin form plexuses in the dermis. In some parts of the body these contain fibers from spinal nerves and also from certain of the cranial nerves. From these plexuses minute nerve fibers pass to the papillse, which contain the tactile end organs. 115. Tactile End Organs. — End organs are those pecul- iarly formed nerve cells or groups of cells which receive and pass on the stimulus to which they are adapted. Tactile end organs are of several different forms. Pacinian corpuscles are found deep in the dermis, scattered along the fine nerve branches like buds on a plant. Each corpuscle consists of layers of delicate membrane within which is a single minute nerve fiber (Fig. 50). Another form of end organ, called the touch corpuscle, appears especially in FlS- 50. — Pacmian the papillae of fingers and toes, and is much smaller than the Pacinian corpuscle. The touch corpuscles are oblong masses, each containing a capsule 88 CONSCIOUS NERVOUS OPERATIONS .V Fig. 51. — Touch corpuscle in pa- pilla of the skin of the hand. C touch corpuscle. N nerve fibers winding around the corpuscle. which receives a nerve whose fibers wind round and round the capsule before entering it (Fig. 51). Still smaller end bulbs are found in the skin, made up of nerve fibers ending in corpus- cles in which the axis cylinder of the nerve terminates. All the tactile end organs are covered by the epidermis, so that the nerves themselves are not brought into actual contact with the external thing which they feel. If the cuticle were stripped off arid pressure applied to a naked nerve end- ing, there would be, not a sense of touch with ability to judge of the properties of the body causing the sensation, but in- stead a sense of pain. Certain portions of the skin are more fully supplied with end organs for touch than are others, and the epidermis there is thinner, so that the sense of touch is more deli- cate. The tip of the tongue, the skin of the face, and the ends of the fingers are most sensitive. A pair of blunt-pointed compasses applied to the end of the tongue will be distinguished as two points even when they are separated by only one twenty-fourth of an inch, while they would be felt as but one point on the finger ; and on the arm or back of the hand the two points much further apart would seem but one. 116. Sensations of Heat and Cold. — Sensations of touch arise from pressure, but through the skin we have, besides, sensations of heat and cold ; that is, we perceive changes of temperature. It is thought that experiments indicate in the skin a separate set of end organs stimulated by heat, and another set which is stimulated by cold. 117. The Muscular Sense. — Still another sensation asso- THE SKIN AS AN ORGAN OF SENSATION 89 elated with the sense of touch is that called the muscular sense (§ 95). By means of this sense we judge of the weight of a body. When we hold an object in the hand we feel its pressure upon the skin, and we also are conscious of a muscular effort to support its weight. We lift it up, move it from place to place, and by the amount of effort put forth judge of the weight of the object. In this pro- cess the muscles are involved as well as the cutaneous organs of sensation. Again, we are conscious, even with our eyes closed, of the position of the whole body. (This we shall find to be connected with a certain part of the internal ear, § 179.) We are also conscious of the position of different parts of the body in relation to one another, and when we come in contact with external objects we perceive not only the pressure from them affecting our organs of touch, but also the pressure which we exert by muscular contrac- tion upon them ; that is, the resistance to our movements which is exerted by external things. It is sometimes said that we have a "sense of effort" (or weight), a " sense of position," and a " sense of movement," but all these are included in the more general term, muscu- lar sense. 118. Pain. — It is not yet fully determined whether the sensation which we call pain is due merely to excessive stimulation of the already known sensory organs, or is a distinct sensation. Sometimes there is disease of sensory tracts which destroys sensitiveness to pain, though the sense of touch is unaffected. It may be that the nerve fibers already referred to, which, distributed everywhere through the body, constantly convey to the brain impres- sions of which we are usually hardly conscious, and which we call impressions of general sensibility, are the channels 90 CONSCIOUS NERVOUS OPERATIONS by which a sense of pain is conveyed when they are more strongly stimulated. 119. Path of a Touch Impression. — Let a touch corpuscle be stimulated by pressure, and what follows ? (Fig. 52.) The cells of the end organ communicate the impulse to the afferent nerve fibers passing from the touch corpuscles to the nerve distributed to that part of the skin. The sensory nerve fibers may run through more than one nerve plexus and through ganglionic nerve centers. There may be several breaks in the path, where the original impulse is handed over from one fiber to another, or passes from cell to cell, or from fiber Fig. 52. — Diagram of the path of a touch impression. The dotted lines show the path when the im- pulse ascends to the brain, exciting conscious- to Cell ; but the ness, and through the motor center (MC), producing voluntary motion. pulse finally reaches the spinal cord (ex- cept when a cranial nerve conveys the impulse), through a posterior nerve root, and the cells of the gray matter of the cord are stimulated. But there is yet no sensation, even though by the reflex action of the cord a motor impulse may be sent out by an THE SKIN AS AN ORGAN OF SENSATION 91 anterior root and certain muscles may be made to contract. The brain must be stimulated before there can be percep- tion of a nervous impulse. Through the nerve fibers which run from cells of the gray column into and up the white columns of the spinal cord, the nervous impression must be carried on till it affects the special nerve cells in the particular portion of the brain set apart for receiving the particular kind, of nervous stimulus from the particu- lar part of the body. Then sensation results, and through the influence of the will, carried along the efferent motor fibers, motion may be produced in the voluntary muscles, and a great variety of acts may take place in consequence of the stimulation of the sensory organs in the skin. DEMONSTRATIONS AND EXPERIMENTS 28. Simple Epithelium. — In connection with the subject of the skin, the general structure of epithelium may be profitably illus- trated. Where frogs are kept in captivity, excellent examples of simple epithelium can be obtained from the moulted skins, for examination with the microscope. 29. Ciliated Epithelium. — If the roof of a living frog's mouth be scraped with a scalpel and the debris thus obtained be mounted in normal salt solution on a slide, there can be found, on examination with the microscope, many cells which show cilia moving actively. 30. Squamous Epithelium. — Scrape the inside of the cheek with a scalpel, mount the debris in water on a slide, and examine with the microscope. Irregular flattened plates can be distinguished, singly or in groups. In most of them the position of the nucleus can be discerned. 31 . Epithelium in Section. — If possible, some prepared microscopi- cal sections of various kinds of epithelium should be studied by the class. Sections of the wall of the digestive tract, of the trachea, of blood vessels, etc., furnish excellent examples. 32. Section of the Skin. — Specially prepared sections of human skin can be purchased. Usually such sections contain hair follicles, sweat glands, and oil glands. 92 CONSCIOUS NERVOUS OPE NATIONS 33. The Epidermis. — Observe that with a needle a portion of the outer skin may be removed without pain or flow of blood. If the hands be washed in warm water and then dried, on rubbing them together briskly, portions of the dead scaly epidermis will be removed. On the palm of the hand the epidermis is seen to be thrown up in a series of curved parallel ridges. The latter are caused by the projec- tions of the papillae of the underlying dermis. On examination with a lens, large numbers of the minute openings of the sweat glands may be seen on the ridges of the epidermis. 34. Discrimination in Touch. — Find the least distance at which the points of a pair of blunt-pointed compasses can be distinguished as two points when applied to the skin of the arm. Repeat the same experiment on the back of the hand, the forehead, the finger tips, and the tip of the tongue. In this way a region of greatest sensitiveness can be distinguished. 35. Location of Touch. — Ask a person to close his eyes, touch some part of his body with a pencil, and ask him to indicate the same point with another pencil, immediately afterward. He will probably make some errors. The experiment may be made more interesting by repeat- ing the trials, taking the measure of each error and averaging the errors. By repeating this experiment on a number of persons, some very interesting results may be obtained and tabulated. 36. Aristotle's Experiment. — Cross the middle over the index finger so that the tip of the middle finger is on the thumb side of the index finger. Place between the two a marble or other small object. A sensation of two objects will result, especially if the fingers be moved. 37. Delicacy of Touch. — With small weights of pith or cork, find the least pressure that is perceptible on the skin of the arm and tips of fingers. The weights may be applied by lowering them upon the skin by means of delicate silk fibers attached to them. The surface which is applied to the skin should have the same area in all the weights, and care should be taken that the weights do not move after touching the skin. The person experimented upon should keep the eyes closed while the weights are being applied. The weights should also be applied to the forehead, temples, lips, and tongue. The main purpose of this experiment is to determine the regions of the skin most sensitive to contact. 38. Hairs as Organs of Touch. — If the weights used in the preced- ing experiment be applied to regions of the skin possessing hairs, it THE SKIN AS AN ORGAN OF SENSATION 93 will be found that a weight touching a hair may be felt, even though its contact is not perceived when it is applied directly to the skin. 39. Estimation of Weight by Sense of Pressure. — Rest the back of the hand upon some easy support, and place on the pal in a small wooden or pasteboard disk. Upon the latter place different weights. Find the least difference in weight that can be detected. A great variety of weights can be obtained by loading empty cartridges with shot to any desired extent. If the cartridges are all of the same size, then the person experimented upon can not estimate the weight by sight. Sev- eral pupils should be experimented upon, to show variation in acute- ness of pressure sense. 40. The Muscular Sense. — Modify the preceding experiment by having the weights lifted instead of simply allowing them to press on the hand. It will be found that smaller differences can be detected than by pressure alone. Demonstrate that a weight lifted slowly seems heavier than one lifted rapidly. 41. Sensations of Heat and Cold. — That there are in the skin two distinct varieties of nerve endings of the temperature sense can be very easily demonstrated by carefully stimulating any certain area of the skin with hot and cold bodies. Let a square be marked off with ink, on the forearm, and a pointed brass rod be heated and then care- fully drawn across this square in parallel lines in various directions. Here and there a sensation of heat will appear distinct from the sensa- tion of contact. The hot spots should be marked with ink dots, as they are recognized. Then in a similar way go over the square with a cold brass rod. Cold spots will occasionally appear, in almost every case distinct from the hot spots. The cold spots should be marked in ink of a different color from that of the hot spots. CHAPTER VII TASTE AND SMELL 120. The mucous membrane of the mouth contains the nerve endings which are affected by taste stimuli. The spe- cial taste organs are found chiefly in the papillce of the tongue and the palate. 121. The Tongue is a mass of striped muscular tissue, with fibers running in va- rious directions, the whole organ being covered with mucous mem- brane (Fig. 53). The papillce of the tongue are much larger than those of the cuticle and are plainly visi- ble to the naked eye. In some animals, as the dog and cat, they are very prominent Fig. 53. JT filiform papilla4. The tongue. 7 funjfiform papilla. L circumvallate papilla. and have horny spines TASTE AND SMELL 95 Fig. 54. — Section of circumvallate papilla. which give a marked roughness to the surface. On the front and sides of the tongue the papillae are generally long and slender, and are therefore called filiform papillce. Others with broad, spreading, mush- Taste B room-shaped tops are scat- tered among the filiform papillae and are called fun- giform papillce. A third variety, the largest of all, is the circumvallate papillce (Fig. 54). These last are only eight or ten in number, and are seen at the back of a man's tongue, in two rows converging to a point backward. 122. Taste Buds. — In the walls of the circumvallate and in some, at least, of the fungiform papillae the end organs for taste have been found. These consist of a number of overlapping epithelial cells, like the leaves of a bud (Fig. 55). The innermost core of the bud is a number of slender, closely packed cells terminating in fine, stiff spikes which project at the surface of the bud. These are the Jc Iff* 00 • A isolated taste bud, from taste ce^s and are tne essential part whose upper free end of the taste buds. Around these project the ends of the ,, ., ,., taste cells. CQLls the nerve filaments from cer- B supporting or protect- tajn branches of the ninth pair of ing cell. ,0.1 7 7 C taste cell. cranial nerves (the glossopharyn- geal) end in brushlike expansions. Branches from the fifth cranial nerve (the triyeminaT) are also distributed to the tongue, and are believed by many physiologists to be concerned in taste. Taste Cells upporting Cells 96 CONSCIOUS NERVOUS OPERATIONS Taste buds are affected so as to distinguish different tastes only when the substances submitted to them are in solution. The effect is increased by friction between the mouth and the tongue. 123. Classification of Tastes. — Tastes are of four sorts : (1) sweets, which are best appreciated by the tip of the tongue ; (2) sour, or acid tastes, perceived best by the side of the tongue ; (3) bitter tastes, most affecting the back of the tongue ; (4) salts. It is believed that separate taste buds are provided, one sort being stimulated only by bitter substances, one by sweet, one by sour, and one by salt. Some substances taste sweet at the tip of the tongue and bitter at the back of it, because they are able to stimulate two sorts of taste buds, but one kind of buds recognizes only a sweet taste, the other only a bitter taste. 124. Flavors. — We are accustomed to say and to think that we taste a great variety of flavors in our food ; but physiologists tell us that we really taste only the four flavors mentioned above, while others are recognized by the sense of smell. This may be tested by holding the nostrils closed by the fingers while different kinds of food are eaten. An onion will not taste different from a potato, though one would be known from the other by its texture. 125. The Sense of Smell. — The organ for receiving im- pressions from the minute particles called odors is the mu- cous membrane lining the upper part of the nasal cavity. 126. The Olfactory Nerves are the first pair of cranial nerves. They spring from the olfactory lobes, which are prolongations of the hemispheres of the brain (Fig. 19, p. 29) and extend forward from the base of the cere- brum. Branches of the nerves of smell are distributed from the olfactory bulbs, in which the olfactory lobes end, TASTE AND SMELL 97 to the mucous membrane of the nasal passages (Fig. 56), where the fine filaments of the nerve end in delicate rod - shaped cells crowded in among the columnar cells of the epithelium lining the nasal passages (Fig. 57). Fig. 56. - Section of nose, showing outer wall Flf • 57 ~ Cells from ol- of right nasal cavity factory membrane, a, 6, c, d interior of nose. -# epithelial cell. K olfactory bulb, below which are seen the nerve W nerve cell, fibers spreading out in the mucous membrane. 127. An Olfactory Impression. — Odors are usually car- ried to the membrane of the nose by means of the atmos- phere, but they must be dissolved or suspended in liquid before they can affect the end organs for smell. Hence the glands of the olfactory membrane lining the nasal passages, whose secretions keep the surface always moist, are important subsidiary organs. As the air is brought to the membrane in ordinary breathing, one is able to perceive various odors when moderately strong. By what is called " sniffing " the air is drawn into the upper as well as the lower nasal chambers, and more of the MACY'S PHYS. — 7 98 CONSCIOUS NERVOUS OPERATIONS odoriferous particles reach the olfactory cells. Thus one is able to examine more fully the odors of the air, and several different " smells " may be sometimes distin- guishable at once. Usually the odors reach both nostrils at the same time and two impulses are conducted along the two olfactory nerves ; they are, however, fused into one sensation. If different odors are brought at one time to the olfactory cells of the two nasal passages, one sensation sometimes destroys the other ; sometimes first one and then the other odor is perceived ; in any case there is but one sensation. 128. Other afferent impulses than those of smell may arise in the nasal membrane. A very pungent substance, such as ammonia, causes sensation distinct from smell, sensation which is found to belong both to parts of the nasal membrane on which the olfactory nerves ramify and also to other portions. A very small quantity of odoriferous material is suffi- cient to excite the sensation of smell. A very minute particle of musk, for instance, will fill a large room with its odor, and that for an indefinitely long time. The end organs of the olfactory neryes are soon ex- hausted, and sensation dies out. We soon cease to notice the odors in a room, though we may have thought them overpowering on entering. Many animals are much more liberally endowed with the power of detecting and discrim- inating odors than is man. 129. Path of an Olfactory Impulse. — The olfactory cells in the epithelium of the nasal passages send a process to the surface of the mucous membrane, and another inward (Fig. 57). The latter process of each cell ends in fine spreading fibrils which mingle with similar brush- like fibrils from a deeper layer of nerve cells in the olfac- TASTE AND SMELL 99 tory lobe. These deeper cells send axis cylinder processes up and along the olfactory tract to different centers in the gray matter of the cerebrum. These centers lie in the temporal lobe, back of the eye (Fig. 58). The olfactory nerves have a path exceptionally direct from their external or peripheral end to the brain center in which they rise. Just what is the connection between this fact and the other often observed fact that stronger mental asso- ciations cling about sensations of smell than about almost any other external impressions, it is not easy to say. Strong reflex nervous action often results from excessive stimulation of the olfactory nerves, as when a person faints in consequence of inhal- ing certain odors. 130. Effects of Alcohol upon Taste and Smell. — The ha- bitual use of drinks containing alcohol, of tobacco, and of very strongly flavored foods is found to dull the sense of taste, and by alcohol, at least, the olfactories are rendered less acute. Fig. 58. — Diagram of the path of an olfactory impulse. The impulse passes from the olfactory cells of the nose to the olfactory bulb (01. B} , and thence to the olfactory center (01. G) on the inner side of the temporal lobe of the cerebrum. DEMONSTRATIONS AND EXPERIMENTS 42. The Tongue. — By the aid of a hand mirror the pupil can easily distinguish the filiform and fungiform papillae, on his own tongue. The circumvallate papillae lie so far back that it will be found more convenient to demonstrate them on the tongue of one of the domestic animals (dead). 100 CONSCIOUS NERVOUS OPERATIONS 43. Varieties of Tastes. — Wipe the tongue dry and place on its tip a crystal of sugar. It is not tasted until it dissolves. Place a crystal of sugar on the tip and another on the back of the tongue. The sweet taste is more evident at the tip. Repeat the process, using a strong solution of quinine sulphate dissolved in water by aid of a little sulphuric acid. The bitter taste is most pronounced on the back of the tongue. In a similar way determine where acids and salts are tasted, using a 1 per cent solution of acetic acid and a 10 per cent solution of common salt respectively. 44. Organs of Smell. — The turbiuated processes of the human ethmoid and maxillary bones, or those of some of the domestic animals, should be accessible to the pupil for examination, as they show how a great deal of surface in a small space is provided for the olfactory membrane. The teacher should perform such dissections upon the head of some one of the domestic animals as to show the olfactory epithelium, the exterior nasal passages, and the posterior passages opening into the pharynx. 45. Combination of Taste with Smell. — Close the nostrils, shut the eyes, and try to distinguish by taste alone between an apple, a potato, and an onion. Chew a grain of roasted coffee and notice how nearly tasteless it becomes when the nostrils are closed. 46. Fatigue of Smell. — For several minutes smell continuously of a piece of camphor gum, breathing in through the nose and out through the mouth. The intensity of the smell becomes much lessened. But if some other odoriferous substance, as clove oil, is brought near the nostrils, it will be found that the fatigue is only for the odor of camphor. CHAPTER THE EYE AND THE SENSE OF SIGHT 131. By means of touch, taste, and smell the brain per- ceives external objects through actual contact between particles of matter from the objects perceived and the human organism. But we need to be able to acquire knowledge of the properties of objects at a distance from ourselves. The sense of smell does indeed bring to us limited information respecting some classes of objects at no great distance; but it is by means of the eye and the ear that we gain our most valuable knowledge of the uni- verse, and through these that we enjoy the most refined and elevated of all our pleasures. 132. Vision. — When rays of light fall upon a nervous apparatus so made as to be affected by that stimulus, and the impulse is carried by a nerve to the nerve center for vision, there results the sensation of sight. Some animals possess a simple arrangement for vision, consisting of only three parts. Certain modified parts of the epidermis are stimulated by the light, nerve fibers carry the impulse to the nerve center, and light is perceived. Man, however, is provided with organs of vision of elaborate and com- plex structure. 133. Light. — All space is believed to be filled with an extremely thin, perfectly elastic medium called ether, in which atoms, molecules, and masses of matter are immersed 101 102 CONSCIOUS NERVOUS OPERATIONS as fishes are immersed in the sea. Of this ether little- is known, but it is supposed to transmit energy by waves. The energy resident in vibrations of the ether is called radiant energy, and it receives special names according to its special, manifestations. For example, when it raises the : temperature of objects^ which receive it, it is called radiant K|{ ATK'NS edges, in little grooves, are minute glands from which an oily secretion flows to the free edges of the lids and prevents their adhesion when closed. Above the orbits the thick ridges set with hairs, called eyebrows, also serve to shield and shade the eye. In a depression in the upper and outer part of the orbit lies the lach- rymal gland, with ducts opening on the inner surface of the upper lid. It secretes a wa- tery fluid designed to lubricate the surface of the eyeball. When stimulated by the irritation of the mucous membrane of the eye, the nose, or the mouth, or by strong mental emotion, the lachrymal fluid becomes excessive, and is called tears. Canals or ducts placed at the inner angle of the eye carry off the ordinary supply of lachrymal fluid to the nasal passages. Branches from the fifth (trigeminal) cranial nerve supply this gland and send fibers also to the eyelids and to inner portions of the eyeball. 151. The Eye as an Optical Instrument. — As has been said, the eye is like a photographer's camera obscura, the various parts of which all have to do with the produc- tion of distinct images of external objects upon the back portion of the box ; that is, in the camera upon the ground glass screen, in the eye upon the nervous membrane called Fig. 67. —Front view of the right eye, showing the position of the lachrymal apparatus. G lachrymal gland. CC upper and lower lachrymal ducts. B naso-lachrymal duct. THE EYE AND THE SENSE OF SIGHT 113 the retina (Fig. 68). In the camera a glass lens serves to bring the rays of light to a focus upon the screen. In Jig. 68.— Formation of an image on the retina. the eye the cornea and the crystalline lens accomplish the same object. 152. Accommodation (Fig. 69). — The camera is pro- vided with an apparatus for changing the distance of the lens from the sensitive plate or screen, so that light from objects at different distances may be focused. The eye is likewise supplied by means of what is called accommodation, which is the power of the eye to adjust itself to objects at different distances. This power is due primarily to the varia ble shape of the crystalline lens. In the camera, the lens and the screen which receives the image are moved nearer to- gether or farther apart in order to change the focus. In the eye the same object is gained by changing the convexity of the lens itself. This is accomplished by the contraction and relaxation of the ciliary muscle, which lies just beyond the outer margin of the iris in the front part IOCY'S PHYS. — 8 Fig. 69. — Dia- gram show- ing how the lens changes its form. 114 CONSCIOUS NERVOUS OPERATIONS of the clioroid coat. It consists of plain muscle fibers, whose nervous supply comes from the third cranial nerve. The ciliary muscle is attached to the ciliary processes, and these to a membrane called the suspensory ligament. This ligament is secured also to the circumference of the lens in such a way that when the eye is at rest it is in a state of tension, which causes it to pull upon and slightly flatten the lens. This keeps the eye at the focus neces- sary for seeing clearly objects at a distance of perhaps twenty or twenty-five feet. When the eye is directed to a nearer object, the fibers of the ciliary muscle contract, thereby drawing forward the ciliary processes, and thus lessening the tension on the suspensory ligament, and the elasticity of the lens causes it to push forward, or become more convex on its front surface. This shortens the focal distance, that is, causes the rays of light to converge more rapidly. The cornea alone is capable of forming distinct images, and the chief function of the crystalline lens seems to be that of accommodation. 153. Function of the Iris. — The iris (Fig. 63) is an adjust- able curtain, designed, by narrowing the pupil, to cut off a portion of the light which might render the image con- fused, and to prevent too strong a light from entering the interior of the eye. This change in the size of the pupil is effected by the contraction and relaxation of its mus- cular tissue, under control of the ciliary nerves. 154. Inversion of Images (Figs. 60 and 68). — As in a camera, so in the eye the image formed is inverted. The rays of light cross in being brought to a focus, so that the picture of a man, a house, or a tree on the retina is upside down, and also much smaller than the object itself. The reason why we do not see objects inverted and re- duced in size is because it is not the picture on the retina THE EYE AND THE SENSE OF SIGHT 115 that we see, but the object itself. Sensation is not in the eye, but in the brain, or in the mind acting through the brain. It is only by study and research that we learn the fact of the inverted image in the eye, and meantime we are accustomed to supplement our visual impressions by the use of our muscles and our organs of touch. The hand interprets the impression on the eye, and we learn to see objects in their true positions. We judge of their positions by the direction from which the light comes to the eye, and of their size by a variety of experiences which complete the impression given by sight. The figure on the retina has little or nothing to do with those judgments. 155. Seeing with Two Eyes. — Two images of one object are formed on the two retinas, and two optic nerves and tracts convey the impression 'to the two opposite sides of the brain. Why, then, do we not see two objects ? Here again we must remember that our perceptions are never simple, due to the action of a single organ and an isolated set of nervous connections. Probably in every act of perception the nervous system acts as a whole through the intricate interlacing of nervous fibers and the close connections of the cells in the various nerve centers. An impression upon one set of end organs is supplemented and corrected by a great number of familiar perceptions of diverse sorts brought before the mind by memory, so that the resulting judgment is an act too intricate and complex to be disentangled. We come to think of the object as we know it from all these combined impressions, and not from a single one of them. Perception is the result of association and experience combined with the physical processes involved. 156. Advantages of Two Eyes. — For perfect vision the retinal images must be formed upon corresponding portions 116 CONSCIOUS NERVOUS OPERATIONS of the two retinas. The two pictures are not, however, identical. The right eye will see more of one side of an object than will the left, and the left eye will see more of the opposite side. This enables us to form more accurate judgments of form and distance than would be possible with only one eye. Then, too, one eye may be wholly destroyed and a person may still retain distinct vision. 157. Duration of Sight Sensation. — The impression made upon the retina by a flash of light remains for about one eighth of a second, so that if flashes of light follow one another at a shorter interval than that they appear as one continuous impression. Children make a circle of fire by whirling rapidly a lighted stick, and the spokes of a swiftly revolving wheel appear continuous. If one looks at the sun or other bright object and then closes the eyes, he will continue for an instant to see the object. These delusive appearances are due to the fact that the nervous impressions made by light upon the end organs in the ret- ina remain after the removal of the rays which excite them. 158. Fatigue of the Retina. — While the retina is extremely sensitive it is also easily fatigued. If one looks steadily for a time at a bright object and then turns the eye away, he will still see the outline of the bright object, but it will be dark. This is because that part of the retina upon which the light fell from the bright body has become wea- ried and no longer responds to the stimulus of light. If the body looked at is of a bright yellow color, the figure seen when the eye is turned away will be blue, because the retina is no longer able to respond to the stimulus of yellow rays, but is affected by the rays of the complemen- tary color. 159. Defects of Vision. — It is very common to see per- sons wearing lenses, or " glasses," to correct what is called THE EYE AND THE SENSE OF SIGHT 117 a shortsightedness," or "longsightedness." In short- sighted persons the rays which in normal eyes come to a focus exactly upon the retina meet at a point in front of it, so that no distinct image is formed. The eye is too long from front to back, and the difficulty must be cor- rected by using lenses which will carry the focus back to the surface of the retina (Fig. 70). Fig. 70. — Diagram showing position of retina. In natural sight (5). In far sight (£). In near sight (C7). Those who habitually use their eyes for seeing only objects near at hand are apt to become shortsighted. It has been found that children who have grown up in crowded parts of smoky London, with little opportunity for looking off long distances, are very often nearsighted. On the other hand, sailors and others accustomed to use the eye constantly for distant vision grow longsighted. Their eyes become shortened from front to back, so that the focus for ordinary vision is beyond the retina. As persons grow old the eye usually becomes flattened on this axis, and glasses are needed to converge the rays of light more rapidly. 160. Squinting, etc. — The muscles of the two eyes act simultaneously so that the visual images are formed upon corresponding parts of the two retinas. But if the internal rectus muscle of one eye is paralyzed, or for some reason 118 CONSCIOUS NERVOUS OPERATIONS the external rectus is too short, that eye will turn outward and defective vision will result. If the external rectus is paralyzed, the person will be cross-eyed, or squint-eyed. Paralysis of either of the nerves distributed to the muscles of the eye will result in abnormal action of those muscles. 161. Astigmatism. — Another defect, known as astigma- tism, is due to the irregular curvature of the cornea or the lens or both. The eye may be more convex on one merid- ian than on others, so that rays of light falling upon one part of the cornea (which is most frequently affected) are brought to a focus at a different spot from the rays which pass through other parts. Thus the image on the retina is indistinct and the vision is blurred. 162. Color Blindness. — Some persons are unable to dis- tinguish certain colors from certain other colors. They are said to be color blind. Usually they differ from persons of normal sight in their inability to distinguish red from green. Sometimes only one eye is color blind, the other being normal. The reasons for these phenom- ena belong to the abstruse subject of color sensation, which is beyond the scope of this work. The power to distinguish colors accurately is of great importance to those engaged in certain occupations, — for example, to those employed upon railways, who are re- quired to undergo examinations which test their eyes in that particular. 163. Training of the Eye. — The eyes of the young may be easily trained by practice under a teacher's guidance to see quickly and accurately, and to judge correctly as to the size of objects, distances, etc. Such training is of great value in all circumstances of life. It is indeed more strictly a training of the mind than of the eye, but niay be fitly mentioned in this connection. THE EYE AND THE SENSE OF SIGHT 119 164. Care of the Eye. — More than some other organs, the eye depends for its healthy condition upon the gen- eral health of the system. If that is impaired, the eye is often weakened and liable to disease. The children of the poor, who are ill-nourished and inadequately clothed, oftener suffer from sore eyes and defective vision than do those in better circumstances. Living in filthy sur- roundings or rubbing the eyes with dirty hands often provokes diseases of the eyes and eyelids. Some forms of general disease — measles, diphtheria, scarlet fever, for ex- ample— are apt to leave the eyes for some time in a sensitive condition and in need of special care. Many eyes, even those of young children, are abnormal in re- spect to focalization, and the defect is often unsuspected until a child has endured much inconvenience or even suffering. Hence it is well for the eyes of every child to be examined by a competent oculist, and to have any defects corrected by suitable glasses. Children in school often suffer from severe headaches and appear dull at their studies simply because of easily remedied defects of vision. 165. Very strong light should never be allowed to enter the eye directly. When reading, sewing, writing, etc., one should sit so that the light will fall upon the work from the left side without shining into the eyes. But one should not read or write with direct sunlight falling upon the paper. Lamps should be provided with shades to shield the eyes, and the light should be steady, for a flickering light is exceedingly trying to the eye. Too faint a light also strains the eye. One should not read or work by twilight, or by any light too dim to per- mit the book or work to be clearly seen at about eighteen inches from the eyes. Eyes may be made nearsighted 120 CONSCIOUS NERVOUS OPERATIONS by carelessly acquiring a habit of holding books, etc., nearer than is necessary. Reading while in a moving railway train or carriage is bad for the eyes, as the motion necessitates constant adjustment of the eyes to varying distance, and the power of accommodation is overstrained. It is well when using the eyes closely to raise them often and look off to a distance, or to close them for a moment of rest. Warm or tepid water is better than cold water for bath- ing the eyes. A compress wet in very hot water and laid over the eyes a few minutes at a time, several times a day, will cure slight inflammation, or relieve the weariness of the eyes after close application. 166. Effects of Drinks containing Alcohol upon the Eye. — Through its influence upon the nerves and the muscles, the continued and too free use of alcohol renders the eye unsteady and its adjustment uncertain ; the small blood vessels become dilated, and the eyes are blood-shot and often inflamed. The optic nerve is frequently affected, causing dimness of vision, and specific diseases of parts of the eye may result, such as cataract and disorders of the retina. The confirmed inebriate is the victim of dis- eased conditions in which the sight becomes untrust- worthy. He sees horrible visions, frightful, venomous creatures appear to threaten him, and he is haunted by specters. Under his imaginary suffering he may become a raving maniac, and repeated attacks of the disease are likely to prove fatal. DEMONSTRATIONS AND EXPERIMENTS 47. Dissection of the Eye. — The eye of the sheep or of the ox should be studied. It may be examined fresh or after preservation THE EYE AND THE SENSE OF SIGHT 121 in one of the fluids used in hardening and preserving the brain (Ex. 6). First, the six muscles of the eyeball should be noticed and identified. To examine the internal structures an equatorial incision should be made about halfway between the cornea and the back part of the eyeball, thus dividing the eye into two parts. The structures thus made visible can be identified by reference to the corresponding parts of the text. If the teacher has any knowledge of histological methods, some excellent sections for study with the microscope can be prepared by hardening the eye of a rat or frog in Perenyi's fluid for two or three days, following with alcohols of increasing strength, infiltrating and embedding in celloidin, and sectioning on a micro- tome. Sections made horizontally through the entire eyeball, and properly stained, show not only the different coats and inclosed structures in situ, but the different layers of the retina, the entrance of the optic nerve, or blind spot, and the yellow spot. Similar sec- tions can be purchased. 48. Refract 'ng Media. — Refraction of light should be demonstrated by means of lenses of various forms. Especial attention should be given to the formation of images by convex lenses. The office of the lens of the eye can be shown by removing it from the eye of a recently killed animal and allowing the direct rays of the sun to be focused by it. 49. Inversion of the Image on the Retina. — This can be very easily shown by cutting away the posterior part of the sclerotic coat of a fresh ox eye, leaving the retina intact. Then on turning the cornea toward some bright object, as a candle flame, an inverted image of the object may be seen shining through the retina. The image appears much clearer if the eye is placed, cornea forward, in a tube of blackened paper. 50. Model for demonstrating the Optical Properties of the Eye. — At little labor and expense the teacher or pupil can construct simple appa- ratus that will illustrate many of the optical features of the eye. Pre- pare an oblong box from twelve to eighteen inches long, open on one side, and blackened within (Fig. 71). One end should be perforated in the center by an opening one half to one inch in diameter, to represent the pupil of the eye. A watch crystal can be fastened over the open- ing, outside, to represent the cornea. The amount of light admitted through the opening in the box can be regulated by means of paper diaphragms with different-sized perforations. Inside the box a reading 122 CONSCIOUS NERVOUS OPERATIONS glass or other biconvex lens can be arranged on a support, so as to focus the rays of light admitted through the pupil upon a movable screen at the back part of the box. A serviceable ground glass screen can be made by rubbing a piece of ordinary window glass with emery powder and water. The box is left open on one side to permit observation and manipulation ; but while experimenting the observer will find it neces- sary to exclude all light, except that which enters through the end of the box. This can be done by covering the box and the head of the ob- server with a black cloth. While with this apparatus one can illustrate Diap \ ragm (Ir J /s >•> V i Screen (Retina) i» Fig. 71. — Apparatus for illustrating the optical properties of the eye. most of the characteristic optical features of the eye, yet obviously it shows nothing regarding the functions of the aqueous and vitreous humors. It must be borne in mind, also, that in the eye accommo- dation for objects of different distances is brought about by changes in the form of the lens ; here in the model by changes in the position of the lens. Shortsightedness and longsightedness can be very easily illustrated ; also the means of correcting them by spectacles. Astig- matism can also be shown by holding in front of the artificial cornea a bottle with sides of unequal curvature, filled with water. 51. Accommodation. — Hold up the forefinger six or eight inches from the eye. Close one eye and look at the finger ; it appears dis- tinct, while objects across the room seem blurred. Look at these latter ; they become sharply outlined, but the outline of the finger becomes indistinct. Notice that in accommodating for the near object there is a feeling of effort. Cease looking at anything in par- ticular, and allow the eyes to come to rest. They will be found to be accommodated for distant vision. 52. Ask a person to accommodate his eye for distant objects. Then look at his eye from the side, while he adapts his vision to a near THE EYE AND THE SENSE OF SIGHT 123 object without moving the eyeball; the pupil and iris next the observer will be seen to move forward, owing to the increased curva- ture of the anterior surface of the lens. 53. Movements of the Iris. — In the preceding experiment the diame- ter of the pupil was smaller when the eye was accommodated for near objects, but dilated on changing to distant vision. Close one eye, and by aid of a mirror observe the size of the pupil of the other eye. Then open the closed eye ; the pupil of the othar eye contracts. Cover, with the hands, another person's eyes. On suddenly removing the hands, the pupils are seen to contract. 54. Astigmatism. — Close one eye and look at the radiating lines in Fig. 72. Notice which lines, if any, appear with the greatest black- ness and distinctness. Try the other eye ; do the two eyes agree ? Look at the concentric circles of Fig. 72. In what portion of the figure, if Fig. 72. any, do the lines appear clearest ? The teacher should obtain, from an optician, one of the charts commonly used in preliminary tests for astigmatism. With this he can detect some of the more pronounced cases of this optical defect among his pupils. It may happen, how- ever, that very great defects cause little disturbance, since if the two eyes are astigmatic on different axes, one eye may correct the defect of the other. 55. Near sight and Farsight. — These are common defects, and their causes should be illustrated by use of the apparatus described in Ex. 50. If the teacher will perform some of the simpler tests for optical defects of the eye (and almost any local optician will loan the necessary apparatus, and give instructions), he may not only inter- 124 CONSCIOUS NERVOUS OPERATIONS est and instruct his pupils, but confer lasting benefits upon some of them in showing them their defects, and the means of correcting them. 56. The Blind Spot. — Close the left eye and with the right look intently at the cross in the following diagram (Fig. 73), holding the book about fifteen inches in front of the eye. Both cross and circle Fig. 73. — Diagram to demonstrate the existence of the blind spot. are seen. Gradually bring the book nearer the eye; at a certain distance the circle disappears because its image falls upon the entrance of the optic nerve. Bring the book still nearer; the circle reappears. 57. Field of Acute Vision. — Look at a printed page without moving the eyes and observe how few words can be seen distinctly. The diameter of this field of distinct view will probably be found to be about one and one-half inches. Wherever the image of an object falls outside the yellow spot, it is seen indistinctly. 58. Binocular Vision. — Hold before each eye a blackened tube of pasteboard. Two distinct fields will be seen on looking through the tubes. Cause the tubes to converge at their free ends, and the two fields will finally fuse into one. This position of the two tubes repre- sents approximately the normal convergence of the two optical axes. Converge the tubes still more ; the two fields reappear, but they are crossed. Look through the tubes at near and at far objects. It will be found that, in order to have a single field of vision, the tubes must be converged more for the near objects. 59. Close one eye, and, looking steadily ahead, note how much is comprised in the field of view. On opening the eye the field is con- siderably enlarged. 60. Holding the forefinger six or eight inches in front of the nose, look at a distant object, as a tree. The forefinger appears double. Now accommodate the eyes for the finger; the tree appears double. An explanation of this can be deduced from Ex. 58. 61. Movements of the Eye. — Close one eye, and, holding the finger tip on the lid, feel the movements of that eye as the other eye looks about in various directions. THE EYE AND THE SENSE OF SIGHT 125 62. Duration of Sight Sensations. — Cause a small wheel to revolve rapidly ; the spokes no longer appear distinct, but seem to be thinned out and fused together into a semitrausparent membrane. Spin a top composed of an angular piece of card fastened to a suitable axis. It appears circular instead of angular. 63. Look at a bright light for a moment. Then close the eyes; the image persists for a short time. 64. Fatigue of the Retina. After Images. — Look steadily, for one or two minutes, at a window, and then at a plain light-colored wall. An image in which the light parts of the window are dark and the dark parts are light will now be seen. 65. Complementary Colors. — Look steadily at a piece of red paper and then at a light background ; a light green after image will be seen. Repeat, using different colored papers. From dealers in kin- dergarten supplies can be obtained packages of colored papers suitable for this and other experiments on color. 66. Color Blindness. — The teacher frequently finds pupils unable to name colors correctly. This may be due to color blindness, but in most cases is caused by defective training. To make any accurate tests of color blindness, the teacher should procure a set of Holm- gren's test worsteds (price $2.50) and directions for experimenting. CHAPTER IX THE EAR AND THE SENSE OF HEARING 167. The ear is the mechanism by which we hear. It has three main divisions, called the external, the middle, and the internal ear (Fig. 74). Endolymphatie Duct ,'Saccule ' "estibular Passage ^Tympanic Passage Eustachian Tube Fenestra Rotunda Fig. 74. — Diagram of the ear, showing relationship of its parts. 168. The External Ear. — The parts of the external ear are formed to collect and conduct waves of sound to the inner portions of the auditory apparatus. They are the pinna, the external meatus, and the membrane of the tympa- num. The pinna is a sheet of elastic cartilage covered with skin, so folded as to direct the waves of sound through the cartilaginous tube of the meatus, which is continuous with it, to the membrane of the tympanum, or eardrum, which closes the end of the passage. Along 126 THE EAR AND THE SENSE OF HEARING 127 the tube of the meatus are placed the wax glands with their ducts. 169. The Middle Ear, or Tympanum, is an irregular cavity in the temporal bone, lined with mucous membrane which is supplied with small glands. By means of the Eusta- chian tube, which passes from the lower back part to the pharynx, or upper cavity of the throat, the tympanum communicates with the external air, and so equalizes the pressure upon the two sides of the membrane of the tym- panum which separates the middle from the external ear. Opposite the membrane of the tympanum are two smaller openings into the chamber of the inner ear, called the round window (fenestra rotunda^), and the oval win- dow (fenestra ovalis^). These are closed, however, by thin membranes. Three small bones, called the auditory ossicles, form a chain across the middle ear (Figs. 74 and 75). The malleus, or hammer, is attached by one end to the membrane of the tvmpanum. J ' Fig. 75. — The ossicles of the ear. while the other end articu- lates with the incus, or anvil bone. The incus articulates with the third bone, the stapes, or stirrup bone, at the top of its arch. The foot plate of the stapes fits into the open- ing on the inner side of the tympanum called the oval window, and is attached to the membrane which closes it. 170. The Internal Ear, or Labyrinth, is the essential part of the organ of hearing, the others being merely con- ductors of sound waves. The labyrinth is an irregular chamber in the rocky part of the temporal bone (Figs. 74 128 CONSCIOUS NERVOUS OPERATIONS Posterior Canal A ntertor Canal Fenestra Ovalia Fenestra Rotunda and 76). Within it lies a closed membranous sac called the membranous labyrinth, which follows the windings of the bony cavity, and whose parts receive names corre- sponding to the names given to the parts of the cavity. The central part of the labyrinth is called the vestibule ; it is about one eighth of an inch in diameter. In its walls are the round and oval Avin- dows already mentioned. The membranous vestibule is composed of two bags, called the utricle and the saccule, connected by a roundabout passage. 171. From the utricle arise three semicircular canals, lying in the bony passages of the same name. One of these is horizontal when a person stands up- right; the others are vertical but at right angles to each other. Two of these canals are united at one end, so that there are but five openings from the canals into the vesti- bule. Each canal has a swelling at one end called the ampulla. At those swellings fibers from the eighth cranial (auditory) nerve pass from the bony wall through the membranes of the canals, firmly attaching one to the other. For the rest of their course the membranous tubes are free, or only loosely fastened by bands, of connective tissue to the bony walls. 172. The Cochlea is the third and most complex division of the inner ear. It has much the appearance of a snail shell of two and a half coils. A bony tube is coiled spirally round a central tapering pillar of bone. Into this tube Fig. 76. — The bony labyrinth of the right ear. THE EAR AND THE SENSE OF HEARING 129 projects from the central pillar a thin shelf of bone partly dividing the tube into two parts. 173. The Membranous Cochlea. — From the edge of the bony shelf of the cochlea two membranes reach to the opposite wall of the cochlea and divide the cavity into three spiral tubes (Fig. 77). One of these, the vestibular Fig. 77.— Diagram of a cross section of a single coil of the cochlea. Showing the structures that compose the organ of Corti. passage (scala vestibuliy, communicates with the vesti- bule. A second, the tympanic passage (scala tympani), has its base against the membrane of the round window. At the apex of the coil of the cochlea the vestibular and tympanic passages communicate with each other (Fig. 74). The middle passage, the membranous cochlea, or cochlcar MACY'S PHYS. — 9 130 CONSCIOUS NERVOUS OPERATIONS canal, is closed at its apex, but opens near its base into the saccule. In the cochlear canal is situated the most deli- cate part of the ear, the organ of Corti. 174. Fluids of the Labyrinth. — The bony canal of the inner ear is lined with a fine membrane which secretes a thin fluid, filling all the spaces of the chamber. The closed sac of the membranous labyrinth also secretes a fluid similar in composition, but containing less solid matter. When the membrane of the tympanum is thrown into vibration, the movement is communicated to the fluid filling the bony labyrinth, and then through the thin wall of the membranous labyrinth to the fluid inclosed, and so to the terminations of the auditory nerve. 175. The Auditory Nerve and the End Organs for Hearing. — The eighth cranial nerve arises by two roots in certain nerve centers of the medulla oblongata. Its two divi- sions enter the labyrinth between the base of the cochlea and the vestibule. One division, having several branches, goes to the vestibule and semicircular canals ; the other passes up through a channel in the bony axis of the coch- lea, giving forth fibers on its way to the bony shelf described above. These pass through, or come into rela- tion with the spiral ganglion and reach the organ of Corti. 176. The Organ of Corti (Fig. 77), within the mem- branous cochlea, is understood to contain the end organs for the discrimination of degrees, variations, and qualities of sound. It is composed of the rods of Corti with adja- cent hair cells and supporting parts. The rods are pillar- like cells attached by an expanded foot, or base, to one of the membranes of the cochlea and ending in a swelling called the head. The pillars are arranged in pairs, of which there are from three thousand to five thousand, separated at their THE EAR AND THE SENSE OF HEARING 131 bases but leaning toward each other to form an arched roof or tunnel. Toward the apex of the cochlea the rods increase in length but are more widely separated at the base, so that the tunnel becomes lower and wider. Against the rods lean other cells called hair cells, which end in many long, haiiiike processes. Between the hair cells lie certain elongated supporting cells. The nerve fibers end in fine branches between and around the hair cells. 177. Path of an Auditory Impression. — Sound waves pass through the air and fall upon the membrane of the tym- panum. In the middle ear they travel partly through air and partly through solid bodies — membranes and bones, — and in the inner ear through fluids and mem- branes. Vibrations of the membrane of the tympanum are u damped " by the ossicles of the middle ear, which also receive and pass on the auditory tremors to the mem- brane closing the oval window. These bones are so closely bound together that they vibrate as if they were one, the very slight amount of play at the articulations serving to prevent jar and fracture. From the middle ear vibrations pass to the inner ear through the attachment of the stapes to the membrane of the oval window. Movement of that membrane sets up motion in the fluid filling the cavity. That, however, would not be possible (since the fluid is inelastic and incompressible) Avere not a vent provided at the round window. When the stapes pushes in the membrane of the oval window, that of the round window bulges out- ward, and the action agitates the whole body of the fluid which fills the bony labyrinth. But the vibrations in the fluid are also communicated to the walls of the membra- nous labyrinth which it bathes, and the fluid which the latter contains is thereby set in motion. 132 CONSCIOUS NERVOUS OPERATIONS Within the swollen ends of the semicircular canals, and upon the walls of the utricle and the saccule, are project- ing ridges composed of especially modified cells of the lining, between which are spindle-shaped auditory cells from which project auditory hairs into the fluid. At- tached to these thickened disks, or ridges, are minute hard particles, called otoliths, which serve to increase the effect of the vibrations. The auditory vibrations in the fluid and in the membranous walls of the labyrinth reach the auditory hairs and give, according to the opinions of some authors, the sensation of sound, or mere noise. A branch of the vestibular division of the auditory nerve is distrib- uted to the semicircular canals, and when its ends are affected by the vibrations in the fluids of the labyrinth there result, as is now believed, sensations other than perception of sound. 178. The same vibrations pass at the same time up the channel of the cochlea from below, affecting on their way the walls of the membranous cochlea, and throwing into vibration the fluid which they inclose. By the vibrations of the fluid and the membrane the nerve endings in the organ of Corti are acted upon in such a way as to give rise to auditory impulses, resulting in perception of sound quality — musical notes, harmony, etc. The cochlea alone is now regarded as concerned with hearing, other parts of the inner ear with equilibrium, etc. Sound waves may also reach the auditory cells by transmission through the bones of the head, as when one- hears the ticking of a watch held between the teeth. The auditory stimulus passes from the auditory cells by the minute nerve fibers to that branch of the eighth cranial, or auditory nerve, which passes through the coch- lea and into the medulla obloiigata (Fig. 78), whence THE EAR AND THE SENSE OF HEARING certain fibers have been traced to the corpora quadri- gemina, while others convey the impression to the gray matter of certain convolutions of the temporal lobe of the cerebrum. 179. Function of the Vestibule and Semicircular Canals. — Much research has in recent years been directed to the part played by these portions of the inner ear, but no investigator has yet reached a conclusion which is accepted in all its details by all others. It is, however, gener- ally believed that these parts Fig. 78.— Diagram of the path have little, if any, direct con- cern with the sense of hearing and discriminating sounds. The nerve branches distributed here arise in the brain from a root of the auditory nerve dif- ferent from that which sends nerves to the cochlea. The cerebellum, from which some of its fibers come, is well known to be the great center for coordination of muscular movement, and experiments seem to indicate that what is called the vestibular branch of the auditory nerve, which ends in the vestibule and semicircular canals, conveys to the brain impressions of position and of movement in space which have to do with the sense of equilibrium. 180. Hearing with Two Ears. — The two organs and two nerves of hearing convey to the brain, not two sensations, but one. By means of two ears we are able to some extent to determine the locality from which sounds come, of an auditory impulse. The impulse passes from the auditory cells in the ear to the cen- ter of hearing (Au C) in the tem- poral lobe of the cerebrum. Some fibers pass to the anterior corpora quadrigemina (Q). 134 CONSCIOUS NERVOUS OPERATIONS but our judgment as to position in space of sound-pro- ducing bodies, and of directions of sound, are indirect and not always reliable. 181. Differences in Sound Perception. — The ears of differ- ent persons vary greatly in power to distinguish differ- ences in sound and in pitch, or the number of vibrations in a given time. All are limited in the perception of high-pitched notes. The ear may become wearied in respect to sound of a particular pitch, much as the eye is soon fatigued in respect to a particular color. 182. Care of the Ear. — A cold which causes inflamma- tion of the throat often affects the lining membrane of the Eustachian tube and that of the middle ear, causing temporary partial deafness. If the cause often recurs, one or both ears may be permanently impaired. Some- times repeated attacks of inflammation in the ear — as from abscesses — result in perforation of the membrane of the tympanum and great injury to the hearing. Generally the wax which is secreted in the external canal of the ear needs no attention, and should not be picked out. Occasionally, however, it accumulates and hardens upon the membrane of the tympanum so as to interfere with its vibrations and impair the hearing. In such a case it should be removed by a surgeon. Warm or tepid water should be used for washing the ears — never very cold water. Before going to baths in cold water or in salt water the ears should be filled with soft absorbent cotton. A sudden very loud noise, as from a gun or cannon, has been known to rupture the eardrum, and a sudden shout close to a child's ear has been known to make it deaf. Insects sometimes crawl into the canal of the ear, in spite of the wax and the hairs there which usually THE EAR AND THE SENSE OF HEARING 135 prevent such accidents. They do not often do harm, and may be removed by dropping warm water into the ear. DEMONSTRATIONS AND EXPERIMENTS 67. Dissection of the Ear. — The external and the middle ear can be fery easily studied by making a dissection of the head of a cat or other domestic animal. Remove the lower jaw, expose and open the temporal bulla. The latter is in many animals a conspicuous rounded protuberance near the articulation of the jaw. The tympanic cavity with the contained ossicles, etc., can then be studied. The internal ear cannot be easily examined because of its smallness and well-nigh in- accessible situation in the rocky portion of the temporal bone. The bone which contains it can be dissected out, and the general outlines of the cochlear region discerned. Or, this portion of the skull may be treated with weak solution of muriatic acid for some days, after which the parts may be partly dissected out. 68. Effect of Varying A ir Tension in the Tympanum. — While listening to a ticking watch, close both nose and mouth, and expel as much air as possible from the lungs, thus forcing the air through the Eustachian tube into the middle ear. The ticking sounds fainter. Or under like conditions inhale as much as possible. The result is as before. 69. Judgment of the Direction of Sounds. — Let a pupil, seated, keep his eyes closed. Clink together two coins at varying distances and directions from his head, and require him to indicate the direction of the sound. Observe that while he rarely fails to distinguish between right and left, he often errs in respect to other directions. Have him hold his hands vertically one in front of each ear, and see if his judg- ment of direction is thereby altered. Close one of his ears with absorbent cotton, and try the effect upon his location of sounds. 70. A uditory Fatigue. — Strike a tuning fork, press the stem down upon the crown of the head, and hold it there until the sound dies away. Then remove it, and after a short interval replace it. The sound will be heard again, but very faintly. CHAPTER X THE VOCAL APPARATUS 183. The Larynx, which contains the vocal cords, is the special voice organ. It is a chamber made up of cartilages, membranes, and muscles. Four cartilages compose the framework. The thyroid is the largest, and forms a prominent ridge in front (called " Ad- am's apple ") with broad, flat sheets at the sides, ending in prolonged angles, above and below (Figs. 79 and 80). It does not meet at the back of the lar- ynx. The cricoid car- tilage, on the other hand, is a complete Fig. 79. -Front (ven- • ,1 i i hpino- Fig. 80. -Side view tral) view of larynx. of larynx. much wider than the front. On top of the broad hinder portion of the cricoid are the small, triangular arytenoid cartilages, which form with the cricoid a true joint, having synovial membrane and ligaments (Fig. 81). Besides these four principal cartilages, which are of hyaline, or nonfibrous cartilage, there are five others of 136 Trachea Trachea THE VOCAL APPARATUS 13T Fig. 81. larynx, view. Cartilages of back (dorsal) yellow fibrocartilage. They are the epiglottis, which is attached to the upper front part of the larynx and forms a lid to the chamber ; the cartilages of Santorini, little horn- shaped projections perched on top of the arytenoids ; and the carti- lages of Weisberg, which are still smaller bits of cartilage lying in folds of the mucous membrane in the sides of the upper membranous lining of the larynx. A sheet of membrane connects the thyroid cartilage with the hyoid bone at the root of the tongue (Figs. 79 and 80). The thyroid is also joined to the cricoid by joints with synovial membrane, and the lower projections, or horns, of the thyroid clasp the cricoid closely, though permitting movement between them. The cricoid is at- tached by membrane to the upper cartilaginous ring of the trachea, or windpipe. These parts, with the many small muscles and ligaments attached, form the vocal apparatus. 184. The larynx is flattened behind, where it closely adheres to the esophagus (Fig. 82). The esophagus is the muscular tube which conveys food and drink from the mouth to the stomach. Its enlarged upper portion is called the pharynx, and lies back of the cavity of the mouth. The larynx lies below and in front of the pharynx. In ordinary respiration the epiglottis stands nearly erect, leaving open the glottis^ or passage into the larynx. In the process of swallowing, the epiglottis is pressed backward and downward, closing the glottis and permitting the food to slide down the esophagus instead of dropping into the windpipe. 138 CONSCIOUS NERVOUS OPERATIONS 185. The Vocal Cords. — The larynx is smoothly lined with mucous membrane except where it narrows at the glottis. Here at the base of the epiglottis are seen, first, on each side of the lining membrane, ridges called false vocal cords, which are not concerned in speech (Fig. 82). They play the chief part in closure of the glottis during expiration. A little below them are the true vocal cords* Opening of Kustachian tube Soft palate Epiglottis Glottis Esophagus Larynx — Hani palate Tongue Hyoid bone False vocal cords . • True vocal cords Thyroid carti- lage carti- lage Fig. 82. —Vertical section of the head and neck. thickened bands of elastic tissue running in the mucous membrane from the front angle of the thyroid cartilage backward to their attachment at the base of the arytenoid cartilages. Between the true and the false vocal cords is, on each side, a recess called the ventricle. The true vocal cords have fine, smooth edges, and are shining white in color. 186. The Glottis. — In ordinary respiration the glottis is a triangular opening with its apex in front. In deep or THE VOCAL APPARATUS 139 labored breathing it widens during inspiration and narrows during expiration. 187. Vocalization. — There may be a sort of speech with- out the action of the larynx or the vocal cords, when the muscles of respiration, the lips, and what are called the resonating cavities above the larynx are alone brought into play. This produces whispering. But for vocalization the vocal cords must be brought very near together, made tense and parallel, and a current of air must be forced swiftly through the narrow slit, throwing them into rapid vibration. This is effected by means of a complicated arrangement of muscles and ligaments attached to the various cartilages. The two inner angles of the arytenoid cartilages are drawn together by the contraction of certain muscles, while others contract to stretch the vocal cords. The thyroid and cricoid cartilages move upon each other to assist in the process. The muscles of the thorax and the abdomen are also brought into special action, and the column of air in the air chamber composed of the trachea and the bronchial system is thrown into vibration. Sound is a result of the whole process. 188. The Resonating Cavities (Fig. 82). — The pharynx, the mouth, and the nasal chambers are resonating cavities, which, by very slight changes in form and size, are able to bring into special prominence different parts of the general tone produced in the larynx, and so modify the resulting sound. 189. Speech is the enunciation of articulate sounds to express thought, and is the result of the action of the vol- untary muscles by which the vibrations produced in the larynx are modified in the resonating cavities. The fac- ulty of speech is a distinctive gift of man, and is possessed by no other animal. It is a natural gift, but its use is the 140 CONSCIOUS NERVOUS OPERATIONS result of training, and skill is acquired only by long years of practice. The complex and greatly varied action required in speech may take place with great rapidity, and may be continued for hours without exhaustion, as in the case of an accomplished singer or public speaker. 190. Vowels and Consonants. — Voice becomes speech through the modifying action of the lips, tongue, throat, etc. Those sounds of the spoken alphabet which require the more open mouth, the more resonant and more pro- longed tone, are called vowels; those which are uttered with the closer position, and are less prolonged and less resonant, are called consonants. Compare the position of the parts of the mouth in enunciating the a in far with that in sounding the b in cab, for example. Between vowels and consonants there is no absolute division. Sounds represented by some of the letters are more open in some words or syllables than in others. Thus, I and n are sometimes vowels and sometimes consonants. 191. Quality of Voice. — Voices are spoken of as "soft," " harsh," " rasping," " rough,'? " sweet," " low," " gentle," etc. An agreeable voice is a most attractive characteris- tic and a most desirable possession. As the vocal appa- ratus is wholly under the control of the will, and as its use is chiefly a matter of imitation, it is of great importance that while the habit of speech is forming the young should be associated with those whose vocal habits are agreeable and refined, and that the attention of children should be early directed to the cultivation of -soft and pleasant tones of voice. In this respect Americans are especially negli- gent, and "the American voice" has become a byword and a reproach in Europe. 192. Musical Sounds. — Sounds produced by regular vibra- tions are musical. Irregular vibrations result in noise. THE VOCAL APPARATUS 141 No sharp line of separation can, however, be drawn. The sounds of ordinary speech are due to regular vibrations, and are hence musical. 193. Pitch depends upon the rapidity of the vibrations, and that varies with the length of the cords and with their tension. In women the vocal cords are shorter than in men, and the voice is an octave higher in pitch. Pitch due to tension of the cords is a matter of voluntary con- trol within the range of a voice. Loudness depends upon the force of expiration. Stammering is due to lack of coordination in the muscles of speech. 194. Nervous Mechanism of the Larynx. — A certain area in the left hemisphere of the cerebrum is recognized as the nervous center in which impulses resulting in speech origi- nate. From the cells of this center nerve fibers run to other cells in the wall of the fourth ventricle of the brain, to the medulla oblongata, and on into the spinal cord. By means of these communicating fibers the center for speech is brought into connection with other groups of nerve cells, from which arise the various nerves which are con- cerned in vocalization. These are very numerous, for the muscles of the face, the tongue, the thorax, and the abdo- men, as well as those of the larynx, are called into action in speaking, singing, etc. Those nerves distributed to the muscles of the larynx are branches of the vagus, or pneu- mogastric, which is the tenth cranial nerve and rises from the medulla oblongata. Suppose there arises in a man's mind a thought which he desires to express in audible speech. He remembers the sounds of the words which will serve his purpose, and impulses arise in that part of the speech center called the auditory word center, from which they pass to the motor center. Thence the nerves of the various parts of the 142 CONSCIOUS NERVOUS OPERATIONS vocal apparatus distribute the impulse to the necessary muscles. In reading aloud it is the end organs for vision, the eyes, which are first stimulated (Fig. 83); then the impulse is carried to the visual center in the brain, from which nervous influences travel by connecting nerve fibers to the auditory word center. There, as before, the sound of the words is revived and the impulse follows the path previ- ously described. 195. — When a person writes from dictation, another course is followed (Fig. 84). The auditory end organs of the inner ear are first stimulated; the impression travels by the auditory nerves to the audi- tory word center, then across to the visual word center, reviving there the appearance of the words. Impulses pass thence to the motor centers and by motor nerves to the various muscles of the arm and hand involved in writing. If nerve fibers which connect the two word centers, visual and auditory, arc diseased, nei- Fig. 83.— Diagram of path of nervous impulses in reading aloud. Au C hearing center. OC sight center. Sp C speech center in left hemi- sphere. V, VII, X, XI, XII, Sp If nerves supplying motor fibers to the speech organs: lips, tongue, chest, etc. ther reading aloud nor writing from dictation is possible. Vocalization may also be the result of reflex nervous action, as when an involuntary scream follows sudden fright. THE VOCAL APPARATUS 148 196. Dumbness is most frequently due to deafness. The auditory word center has never been stimulated. In recent years it has been found possible to stimulate the speech center through its connection with the visual center. Chil- dren born deaf are taught to imitate the movements of the mouth, tongue, and throat of one who speaks, and speech results from these remembered movements acting upon the speech center. 197. . Care of Throat and Voice. — If the delicate lining mem- brane of the larynx becomes in- flamed through " taking cold " or from exposure to dust or irritating gases or from over- use or strain of the vocal cords, the voice is injured and its use may become painful or impossible. These causes should be avoided. When a daily cold bath is not practi- cable, a dash of cold water over the neck on rising in the morning will prove a tonic for the throat and help to avert colds, hoarseness, and sore throat. Breathing should be through the nose, and not through the mouth, and children should be trained in infancy to sleep with the mouth closed. In going from the warm air of the house into the cold outer air in winter the precaution of closing the mouth is especially to be observed. Pass Fig. 84. —Diagram of the path of the nervous impulses in writing from dictation. Au C center of hearing. MC motor center. OC sight center. 144 CONSCIOUS NERVOUS OPERATIONS ing through the winding nasal canal, the air becomes tempered before reaching the sensitive membranes of throat and lungs; also much of the dust always present in the atmosphere is caught by the moist mucous lining of the nostrils, and the deeper air passages are protected. It is not necessary, in order to avoid cold, that the throat should be thickly swathed in wraps of wool and fur. Too much clothing about the neck causes excess of perspiration there and makes the parts weak and tender. 198. Special Training of the voice for singing or public speaking should not be begun by either boys or girls before the age of sixteen or seventeen, and should always be attended with judgment and care against overstrain. 199. Alcohol and Tobacco as affecting the Vocal Organs. — As the perfect control of the voice depends upon the healthy condition of all the muscles connected with the vocal apparatus, and upon the accurate adjustment of nervous force to their varying needs, anything which affects those muscles or the nerves affects also the voice. Alcohol and tobacco do affect both. The mucous mem- brane of the larynx is often much inflamed by tobacco smoking, and especially by the use of cigarettes. The inflammation may extend through the Eustachian tubes, impairing the hearing, and into the bronchial tubes, caus- ing an annoying cough. A disease known as "smoker's sore throat " may result. Alcoholic beverages irritate the throat and are often forbidden to those cultivating the voice for singing. DEMONSTRATION 71. Dissection of the Larynx. — At a slaughterhouse can be obtained a trachea of an ox with the larynx intact and a portion of the esoph- agus ; the hyoid bone may be present also. With this material the principal topics of this chapter can be illustrated. PART III NERVOUS OPERATIONS UNCONNECTED WITH CONSCIOUSNESS Those nervous operations of which man is necessarily conscious and which are directly concerned in his useful- ness and happiness, cannot continue to minister to his higher nature without the assistance of another set of actions of which he is in health almost wholly uncon- scious. The complicated mechanisms for producing sen- sation and voluntary motion are constantly worn away at every point, and every tissue must- be as constantly re- newed. Through the action, at every moment of life, of nerves and nerve cells whose office it is to preside over what are called the vital processes, the body is kept in condition for the exercise of its conscious -powers. Those vital processes are included under the general term nutri tion, that is, the growth, waste, and repair of tissue. Nutrition is effected by means of the circulation of the blood, digestion (including absorption and assimilation), respiration, and excretion. Because these operations are wholly dependent upon nervous influences, and because they go on through life without necessary connection with consciousness, they are grouped here as unconscious nervous operations. 145 Fig. 85. — The chief blood vessels. 146 CHAPTER XI BLOOD, LYMPH, AND CHYLE 200. Function of the Blood. — It will be remembered that one of the essential properties of the living cell is its power to incorporate into its own substance matter from outside itself, a process which results in growth or in repairing waste ; while another property, involved in this, is the power to break down by oxidation, that is, to resolve portions of its own body into simpler chemical substances, thus producing waste matter. In order that this double process may go on continually (as it must while the cell is living), oxygen and oxidizable substances, that is, food, must be brought to each cell, and provision must be made for removing the waste products. The blood is the medium for accomplishing this. 201. The Blood as a Tissue. — Blood is classed, for valid reasons, among the connective tissues. While it does not furnish support to the body or its parts in the same sense as do more solid connective tissues, — such as bone or cartilage, — it does support the whole body by conveying nutriment to every part. It is like other connective tissues also in that the cells make up a comparatively small portion of its substance, the* intercellular material being largely in excess. And, finally, it is formed in the development of the embryo from the same layer as^ are the other connective tissues. MACY'S PHYS. — 10 147 148 UNCONSCIOUS NERVOUS OPERATIONS 202. Structure of the Blood. — Under the microscope the blood is seen to consist of a nearly colorless fluid, the plasma, in which float cells of two sorts, called from their color the red and the white (or colorless) corpuscles (Fig. 17, p. 26). Ordinarily there are several hundreds of red corpuscles to one of the white corpuscles, and to them the color of the blood is due. 203. The Quantity of Blood in the human body is esti- mated at about one thirteenth the body's total weight, or, in a person of average size, about one and a quarter gallons. Any deficiency of blood in the body (as from hemorrhage), is soon supplied by the passage of water from the tissues to the blood by means of the lymph (§ 209), and an excess is removed by the transfer of water to the tissues, and by the secretion of the kidneys. Thus the quantity of blood in the system is practically invariable. 204. The Red Corpuscles are unnucleated cells, all of nearly the same size — about -$^$ of an inch in diameter, and one fourth of that in thickness. They are round and flat, but slightly thicker at the edge than in the middle. Being flexible and elastic they are bent out of shape as they are crowded together in the current, but resume their usual form when the pressure is removed. They have a close, colorless, spongy framework, — the stroma, — while by far the larger part of their substance is a red coloring matter in the meshes of the stroma, called hemo- globin. This is the useful part of the corpuscle, the stroma apparently having only the office of holding the hemoglobin in convenient shape. 205. The Formation of the Red Corpuscles is found to take place in the red marrow of the bones. The peculiar tissue called red marrow is richly supplied with blood vessels BLOOD, LYMPH, AND CHYLE 149 having very thin walls. Within these vessels are found colored nucleated cells, some of which become changed in the marrow into unnucleated red disks which are swept into the blood current. It is the important function of the red corpuscles to take in oxygen from the air which reaches the blood in the lungs, and carry it to the other tissues of the body. The red corpuscle lives in the blood for an unknown time. When it dies, a new one takes its place. The spleen also is believed to aid in the manu- facture of both the red corpuscles and the white, but we have little positive knowledge upon the subject. 206. The White Corpuscles are mostly larger than the red, being generally about ^Vo .°^ an ^ncn *n diameter, though some are smaller than the red. Some are globular masses of granular protoplasm without cell walls and hav- ing one or more nuclei. Others are of irregular and con- stantly changing shape, less granular than the first and with several nuclei. Many writers have remarked the striking likeness of these corpuscles to the one-celled animalcule, the amoeba. They have the same power as the amoeba to change their shape spontaneously, sending out processes from various parts of their circumference; and they are able to take in and digest the bacilli which are sometimes found in the blood, as the amo3ba digests food. There are found in the blood some other small bodies, whose nature and purpose are unknown. 207. Chemical Composition of Blood. — The blood is alka- line, owing to the presence of small quantities of alkaline salts. It contains chlorides, phosphates, and carbonates of sodium and potassium, and smaller quantities of cal- cium and magnesium. 208. Clotting of the Blood. — The blood in the blood ves- sels is perfectly fluid, but if drawn out and allowed to 150 UNCONSCIOUS NERVOUS OPERATIONS stand for a few minutes, it becomes a firm mass of jelly. After an hour or more, a yellow fluid, called serum, begins to ooze from the clot, which shrinks in size. The clotting is caused by the formation, in the liquid blood, of a close network of fine fibrils, called fibrin, in which the corpus- cles of both kinds are entangled, while serum is the plasma of the blood, minus an element in its composition called fibrinogen, which changes into the solid fibrin in the coag- ulation. It is thought that when the blood leaves the blood vessel, or in some way comes in contact with foreign matter, a portion of the white corpuscles are broken up, and thus is set free a peculiar substance called fibrin fer- ment. It is this which acts upon the fibrinogen, and causes it to become fibrin. The fibrin may be gotten out from a quantity of freshly drawn blood by quickly stir- ring or whipping it with a bunch of twigs. The tiny white threads cling to the sticks, and by washing in water may be freed from the few entangled corpuscles which remain, leaving the fibrin pure. This power which the blood has to clot is of great value, since by its means small breakages in or injuries to the innumerable tubes conveying the blood throughout the system are quickly stopped, and the serious hemor- rhage which would otherwise result is quickly checked, while the ruptured wall of the blood vessel is given time to heal. 209. Lymph. — It is by the blood that nutriment is car- ried to every part of the body; but the blood is always inclosed within the walls of the tubes called blood vessels, and, as blood, does not come in contact with the cells of the tissues. In the capillaries, which are the finest ramifications of the blood vessels, some of the plasma passes from the BLOOD, LYMPH, AND CHYLE 151 blood through the walls of the vessels into the spaces between those walls and the substance of the tissues around. This fluid is called lymph, and is that which nourishes the tissue elements. It is clear, nearly trans- parent, and contains more water with less solid matter than the plasma of the blood. White corpuscles similar to those in the blood are found in it, and, like the blood, it coagulates by the formation of fibrin. Like the blood, also, lymph is conveyed from the tissues in tubes, the lymphatics or lymphatic vessels, which finally join the great blood vessels, and so return to the blood the substances drawn from it in the capillaries. 210. Lacteals and Chyle. — The lymphatics of the small intestine are called lacteals ; after a meal containing fat, they convey, instead of clear lymph, a milky fluid which is called chyle, and is a product of digestion. EXPERIMENTS 72. Blood Corpuscles. — Prick the finger with a sterilized needle, mount the drop of blood thus obtained, and examine it with both low and high powers of the compound microscope. There will be seen large numbers of round bodies of a faint red tint — the red cor- puscles. These are seen to be small disks, and appear dumb-bell shaped when viewed on edge, owing to their being thinner in the center than on the edges. Occasionally among the red corpuscles may be seen slightly larger, transparent, sometimes irregular, bodies — the white or colorless corpuscles. If watched for some time, they will probably show slight changes in shape. 73. Clotting of Blood. — At a slaughterhouse fresh blood can be obtained. If it be stirred vigorously immediately after being drawn from the blood vessels, the fibrin can be separated from the blood serum and corpuscles. Blood allowed to stand after being drawn shows a firm clot. Both "whipped" blood and fibrin and the clotted blood should be examined by the pupils. The clotting of blood may be prevented by adding to it, as it is 152 UNCONSCIOUS NERVOUS OPERATIONS drawn from the blood vessels, about one fourth its volume of a satu- rated solution of sulphate of magnesia. This " salted " blood may be kept in a cool place for several days without clotting. It may be made to clot by diluting it with five to ten times its volume of water. A more satisfactory method of preventing the clotting of blood, the writer has found, consists in adding oxalate of potash in the propor- tion of one part of a 5 per cent solution of oxalate of potash to twenty-five parts of blood. The oxalate solution of the requisite amount should be placed in a vessel and the blood be allowed to flow into and mix with it. The mixing should be made thorough by vigorous shaking. To produce a clot, add a few drops of a 2 per cent solution of calcium chloride to some of the oxalate-blood. The potas- sium oxalate prevents clotting by precipitating the calcium salts nec- essary to coagulation. The addition of calcium chloride restores the calcium and renders clotting possible. 74. The Minute Structure of the Fibrin Framework. — To a drop of fresh blood on a slide add two drops of normal salt solution. Put on a cover glass and set aside an hour or so to clot. Add 50 per cent alco- hol at the edge of the cover glass (to wash out the corpuscles and harden the fibrin). Observe with the microscope the network of fibrin fibrils. Care must be taken not to move the cover glass during the preparation for examination. CHAPTER XII THE CIRCULATORY SYSTEM 211. The apparatus distributing the blood throughout the body and keeping it constantly in motion is composed of the heart, the great central pump; arteries, tubes to carry blood from the heart ; veins, which are tubes carrying blood to the heart ; and capillaries, a network of small tubes connecting arteries and veins. These constitute the vascular system (Fig. 85, p. 146). In addition to them and forming a part of the complete circulatory system are the lymphatics and lac- teals, sometimes called the lymph vascular system. They are tubes having walls thinner than those of the blood vessels, running from periphery to center, conve}dng lymph and chyle. 212. The Heart is a hollow, cone- rp. The heart is partly covered shaped muscle, inclosed in a mem- by the lungs, hut its true out- branous sac called the pericardium, line is shown by a dotted liue' lying in the thorax between the right and left lungs (Fig. 86). Its base is directed backward and upward, while its 153 Fig. 86.— Front view of the viscera in their natural relations. 154 UNCONSCIOUS NERVOUS OPERATIONS apex points downward ai}d forward a little to the left of the sternum, or breastbone. The heart is divided longitu- dinally irito two divisions wholly separate from each other. Each of these is divided again into two chambers which have free communication. The chambers at the base Left Pulmonary Artery ..Left Auricle Fig. 87.— Front view of heart. Coronary arteries and veins are injected, arteries red, veins blue. of the cone are called the right and left auricles, while the other two are called right and left ventricles (Figs. 87 and 88). Lying upon the outside of the auricles are two flat earlike structures which are called the right and left appendi'ces. 213. The Eight Auricle lies on the right side of the upper part of the lieart. Its walls are thin, and are pierced by openings for the two great veins, — the superior vena cava. THE CIRCULATORY SYSTEM 155 entering from above, and the inferior vena cava, entering from below. These two veins bring to the heart the blood from all parts of the body, except from the lungs and the heart itself. Close beside the inferior vena cava the coro- nary vein opens into the right auricle. This brings to Right Pulmonary Artery Left Pulmonary Artery" Left Pulmonary Left Auricle. fij?/ /Supertor* Vena Cava Right Pulmonary .--'•%'' Veins iferior Vena Cava Fig. 88.— Heart seen from behind. Coronary arteries and veins are injected, arteries red, veins blue. the auricle the blood from the capillaries of the heart itself, which like all other organs of the body is supplied with blood vessels and lymphatics for its own nutrition. 214. The Left Auricle lies at the left and back of the upper part of the heart. Its walls are slightly thicker than those of the right auricle. It receives four pulmo- nary veins, two from each lung. The openings have no valves. 156 UNCONSCIOUS NERVOUS OPERATIONS 215. The Right Ventricle occupies the principal part of the forward surface of the heart, but is not a part of the apex. The walls are thicker than those of the auricles and less smooth. From the upper side of the cavity of the ventricle a large opening leads to the great pulmonary artery, while a still larger opening admits the blood from Right Auricula-Ventricular Aperture\ J^eft Auricula-Ventricular \ Aperture Aorta Pulmonary Artery •Semilunar Valves .Flaps of Tricuspid Valve Tendinous - Cords '--^Papillary Muscles 89.— Eight ventricle, cut open to show the structures that regulate the action of the tricuspid valve. the right auricle (Fig. 89). The tricuspid valves prevent the return of blood from the ventricle to the auricle. The three main divisions of the valve and three smaller ones are all triangular in shape, and are attached by their bases to a tendinous ring surrounding the opening. The thin- ner edges of the valves are flaps of transparent membrane THE CIRCULATORY SYSTEM 157 hanging downward and held by .slender white tendinous cords attached to little projecting columns in the walls of the ventricle (papillary muscles). These cords and mus- cles keep the valves from being pressed back into the auricle further than is necessary to close the communica- tion. Blood can thus flow from the auricle into the ven- tricle, but not from the ventricle to the auricle. The passage into the pulmonary artery is guarded by three folds, or pockets, in the lining membrane, called semilunar valves, which have their free edges turned upward, so that when the blood is forced upward into the artery, the valves lie flat against its walls, while if the blood should begin to flow backward, the little pockets would at once be filled, their edges crowded together in the center of the tube, and the opening closed. 216. The Left Ventricle occupies the chief part of the hinder surface of the heart, and includes the apex (Figs. 87 and 88). It opens at its upper side from the left auricle and into the aorta, the great artery whose branches bear the blood to the general system. The walls are much thicker than in any other part of the heart because greater force is required here to send the blood to the most dis- tant parts of the body. The mitral or bicuspid valves guard the opening into the left auricle. They are similar to the tricuspids, except that they have only two main divisions instead of three. Tendinous cords and papillary muscles hold their edges in place as in the case of the tricuspids. A strong fibrous ring surrounds the end of the aorta, and within its mouth are three semilunar valves, thicker and stronger than those of the pulmonary artery. Their action is like that of the other semilunar valves. 217. All the cavities of the heart are lined with a 158 UNCONSCIOUS NERVOUS OPERATIONS smooth, shining membrane, the endocardium* which also covers the valves, and is continuous with the lining of the veins and arteries. 218. Cardiac Muscle. — As stated in the chapter on Mus- cles, the muscular iibers of the heart form a class by them- selves, being striped but involuntary (Fig. 90). The fibers lie side by side, but send off at short intervals branches which unite them. The muscular fibers, moreover, are arranged in the wall of the heart in bundles in such a way that in contracting they draw the two sides of the walls of the chambers together until they meet. The muscle fibers of the walls of the auricles are distinct from those 90 —TWO °^ ^e vehicles, so that thev contract sepa- cardiacmus- rately, as we shall see. Each fiber, or muscle cell, contains one nucleus. Cardiac muscle fiber appears, to a large extent, to origi- nate its own contraction, and is not so entirely as is a skel- etal muscle fiber a mere instrument of a motor nerve fiber. The action of cardiac muscle under stimulus is not stronger or weaker in proportion to the strength of the stimulus, as is the case with skeletal muscles. A weak electric shock, if it causes any beat at all in the heart muscles, causes as strong a beat as does a strong stimulus. 219. Arteries are the vessels which convey the blood from the ventricles of the heart. The smallest of them have a few plain muscular fibers wrapped round the tube outside the endothelium. As the arteries grow larger, the number of muscle fibers increases till they form a definite muscular coat with a little connective tissue. In the largest arteries the walls consist of three layers (Fig. 91) : (a) the inner coat, consisting of endothelium with a THE CIRCULATORY SYSTEM 159 thin elastic layer on its outer side; (5) a muscular and elastic coat; (; the water issues in jets corre- THE CIRCULATORY SYSTEM 175 spending to the strokes of the pump. While working the pump press lightly with the finger on the rubber tubing of the arterial side ; a distinct pulsation is felt with each stroke. Repeat the same on the venous side. 82. Osmosis. — Prepare a dialyzer by tying a thin animal membrane (sausage skins, to be obtained of a butcher, furnish excellent mem- branes for osmosis) over one end of a small lamp chimney. Partly fill the dialyzer with a strong solution of sugar and place it in a larger vessel of pure water, so that the liquids in the two vessels are at the same level. In a short time the contents of the dialyzer begin to rise, owing to the greater flow toward the denser liquid. It will also be found that the water in the outer vessel becomes sweet as osmosis goes on. If the membrane is allowed to become thoroughly dry after being tied on the dialyzer, osmosis goes on more rapidly. M N Fig. 98. — Apparatus for illustrating the circulatory system. A bulb syringe, by which water is forced through the apparatus. B rubber tubing packed with bits of sponge to represent capillaries. C rubber tube connecting arterial and venous regions. When C is closed with the pinchcock the liquid must all pass through tubes B. D small tube ending in capillary point and closed with a pinchcock. When open it allows arterial " spurting " to be demonstrated. E tube which shows venous flow. It ends in a tube of small caliber in order to produce a venous pressure. M, N manometers consisting of glass tubing bent in a U-shape and partly filled with mercury. They show the pressure applied to the arterial and venous sides respectively. CHAPTER XIII NERVOUS CONTROL OF THE CIRCULATION 240. Functions of the Nerves of the Circulatory System. — When any special activity is required of one of the organs of the body, an increased flow of blood is needful to that part. There is not blood enough in the body to enable all the muscles, all the organs of digestion, the brain, and the organs of respiration, etc., to work in full activity at the same time. There must therefore be some method of regulating the activities of the different parts and organs, so that some may rest while others work. Then, too, some arrangement must exist for correlating the action of the heart and the blood vessels, so that the steady flow of nourishing blood may be kept up in all the capillaries, with more powerful pressure applied when and where it is needed, and not at the wrong place and the wrong time. As everywhere else, we find that in the circulatory system the nerves furnish the controlling, coordinating, and regulating force. 241. Nerves of the Heart (Fig. 99). — Three sources of nervous control of the heart are usually mentioned. They are the cardiac or heart branches of the tenth and eleventh pairs of cranial nerves, — the vagus or pneumoyastric and the spinal accessory; the cardiac branches of the sympa- thetic^ from the ganglia of the neck ; and what have NERVOUS CONTROL OF THE CIRCULATION 177 been called tlie intrinsic nerves of the heart, which were formerly treated as independent of the other two sets of nerve fibers. The intrin- sic nerves are now known, however, to be merely the terminations of the other nerves in the heart wall, while the rhythmic beat of the ventricles, which is found to continue for some time even after the heart is removed from the body, is believed to be a peculiar property of the heart muscle itself. 242. Path of the Nervous Impulse. — • The roots of the vagus and the spinal acces- sory nerves rise near together /, kgp^ in the gray matter of the medulla oblongata. A branch Fig. 99. — Diagram illustrating the from the spinal accessory soon joins the vagus, and supplies some of its motor or efferent fibers. Fibers from the sympathetic nerves of the neck also join the va- gus. These have been traced back into the spinal cord. The cardiac nerves, there- fore, are all connected with the central nervous system, and, since all parts of the nervous control of the heart. A cardiac accelerator center. a accelerator fibers connected with accelerator center through sym- pathetic system (Sy), spinal nerves (Fis lowered. Replace a and remove b ; DF is raised. Observe that when DF is raised from position IT, the dis- tance between DF and CE is increased, unless DF is raised too far. 86. To show the Moisture in Respired A ir. — Breathe upon a cool plate of glass; some of the vapor in the breath will be condensed upon the glass. To show that the presence of moisture is due to respiration, blow upon the glass writh bellows. 87. To show that A ir is made Warmer in Respiration. — Notice the reading of a thermometer, then breathe upon its bulb. 88. Some Properties of the Constituent Gases of A ir. — The teacher should prepare oxygen, nitrogen, and carbon dioxide by methods given in ordinary text-books of chemistry. Place a smoldering splinter of wood in a vessel of oxygen ; the stick bursts into flame. Place the flaming splinter in a vessel of carbon dioxide or nitrogen ; the flame is at once extinguished. Place a blazing splinter in an empty jar, i.e. containing only air; after burning some time it goes out. Before the last trace of fire disappears, transfer the splinter to a jar of oxygen; it burns actively again. Into a clean, empty jar put a little limewater, prepared by dissolving lime in water. Cover the jar and shake it well; if any considerable amount of carbon dioxide is present, a white precipitate will form in the limewater. Now place a burning splinter in the jar and allow it to burn out. Then shake RESPIRATION 197 the jar ; the presence of carbon dioxide will be very evident. Repeat the latter operation after allowing splinters to burn out in a jar of oxygen. From these experiments it becomes evident that oxygen is necessary in ordinary combustion, and that carbon dioxide is a product of combustion. 89. To show that Respiration increases the Amount of Carbon Diox- ide in the Air. — By means of a glass tube blow through lime water; the white precipitate of carbonate of lime shows the presence of car- bon dioxide in the breath. To show that most of this is a product of respiration, pass some of the ordinary air of the room through lime water ; the white precipitate is much smaller in amount. 90. To illustrate a Difference between "Arterial" and "Venous" Blood. — Obtain some fresh ox blood at a slaughterhouse and "whip" it to remove the fibrin and prevent clotting. By means of a carbon dioxide generator, pass carbon dioxide gas through the blood; the blood becomes darker. Now pass air through it, thus supplying oxygen ; it becomes more scarlet in color. CHAPTER XV NERVOUS CONTROL OF THE RESPIRATORY APPARATUS 276. The movements of respiration may go on in ordi- nary quiet breathing without consciousness and without volition, but they are also, in a measure, under voluntary control — not wholly so, for it is impossible to commit suicide by holding the breath. 277. The Respiratory Center and Nerves (Fig. 107). — A certain restricted area in the medulla oblongata is recog- nized as the respiratory center, and there are believed to be other such centers lower down in the spinal cord. Nerv- ous impulses pass from the center down the spinal cord, and thence by the anterior roots of many of the spinal nerves to the plexuses which those nerves form. By com- municating branches from these plexuses connection is made with the spinal ganglia of the sympathetic system, and with the tenth and eleventh cranial nerves. From these various sources motor fibers pass on to the numerous muscles concerned in respiration. That which supplies the diaphragm is the phrenic nerve, which is traced back to the three or four upper pairs of spinal nerves. If the spinal cord be divided below the fourth pair of spinal nerves, the diaphragm will continue to act, but the intercostal muscles will be paralyzed. If the cord be cut just below the medulla oblongata, all respiratory movement of the chest ceases ; and if that small portion of the medulla 198 NERVOUS CONTROL OF RESPIRATORY APPARATUS 199 oblongata known as the respiratory center be removed, no further respiratory movements will take place, and death immediately follows. On the other hand, the whole of the brain for- ward of the medulla may be removed, and breathing will not stop. 278". The Expiratory Center. — It is now un- derstood that the cen- ter of respiration in the medulla is in real- ity double, — that there is one center for inspir- atory movement and another beside it for expiratory movement. In ordinary quiet breathing the first only is excited, expiration taking place by the re- laxation of the muscles contracted in inspira- tion. But in violent or forced expiration the internal intercostal and abdominal muscles are brought into active use, and the nervous influ- ence stimulating them to action comes from the expiratory center. Fig. 107. — Diagram of the nervous con- trol of the respiratory organs. ab motor spinal nerves going to muscles of the abdominal walls. E expiratory center. ex sensory nerve fibers from the larynx ex- citing the expiratory center. ex' sensory fibers from the lungs exciting the expiratory center. ext motor spinal nerve fibers passing to external intercostal muscles. / inspiratory center. in sensory fibers from larynx inhibiting the inspiratory center. in' sensory fibers from the lungs that excite the inspiratory center. int motor spinal nerve fibers passing to in- ternal intercostal muscles, motor fibers of phrenic nerve going to diaphragm. ph 200 UNCONSCIOUS NERVOUS OPERATIONS 279. Reflex Action of Respiratory Center. — Though the action of the breathing center in the medulla oblongata is shown by experiment to originate efferent nervous impulses independent of irritation conveyed to it by sensory fibers, it is also largely under reflex control. For instance, when the mucous membrane of the air passages is irritated by foreign substances, a sudden sneeze or cough results. A sudden dash of cold water upon the skin causes a quick, long inspiration. So it appears that in normal respiration the movements are not reflex, while reflex movements are also possible. 280. The Normal Excitation of the Respiratory Center is due, more than to any other cause, to the relative amounts of oxygen and carbon dioxide in the blood which reaches it. If the blood contains too little oxygen and too much carbon dioxide, the center is stimulated, and the resulting respira- tory movements tend to correct the evil. Afferent nervous influences brought to the center along the .vagus nerves also excite its action. The main trunk of this tenth nerve contains fibers which excite inspiration, and other fibers which inhibit, or check, inspiratory move- ments. It is supposed that the expansion of the air cells in the lungs, where many of the vagus fibers end, produces impulses along those fibers which result in inhibitory or expiratory impulses from the center in the medulla ; while, on the other hand, collapse of the air cells excites along other fibers contrary impulses which result in inspiration. Thus the action of the lungs becomes that of a self-regulat- ing pump. Other afferent impulses along branches of the vagus, or from other nerve fibers connected with it, may also affect respiration. CHAPTER XVI FOOD 281. Losses of the Body. — One effect of respiration is to reduce the weight of the body. A man gives off from the lungs, in the course of twenty-four hours, about eight ounces of carbon and about half a pint of water, which are taken from the tissues of the body as worn-out mate- rials. There are also other sources of loss to the body in the processes of excretion, by which the waste or used-up matter of the body is cast off. 282. Sources of Restoration. — These losses must, of course, be made good, or the body will waste away. Through the lungs we get oxygen only, while we need hydrogen, nitrogen, carbon, and other chemical elements besides, which enter into the composition of our bodies. These come to us in the food which we eat, and are pre- pared for use within the body by the process called digestion. Food is that which, taken into the alimentary canal, sup- plies material for the growth and repair of tissue, for the generation of force, or for the regulation of force. 283. Nutrition. — Digestion is but one portion of a com- plicated series of processes, called nutrition, which result in the growth and repair of the constantly wasting tissues. In its broad sense nutrition includes respiration, which supplies the oxygen needed; digestion, or the conversion 201 202 UNCONSCIOUS NERVOUS OPERATIONS of food by the chemical action of the digestive juices into soluble substances ready to be acted upon by the absorb- ents; absorption, or the passage of selected elements from the digested food through the walls of the blood vessels and lymphatics (lacteals); assimilation, or the conversion of the new particles of food into the substance of the tissues of the body; and the breaking down of the sub- stance of the tissues to form waste. The building-up processes constitute what is called anabolism, the break- ing down of tissue Jcatabolism, while the whole double operation receives the scientific name of metabolism. We are now to study anabolism. 284. Object of Nutrition. — The various processes of nu- trition take place in order to supply the human machine with material in suitable form for the production of energy. The oxidizable substances are distributed to the tissues and are there submitted to the action of the oxygen in the blood. Combustion then takes place, that is, the substances in the tissue cells are burned. The complex chemical compounds there found are decomposed, and new and simpler compounds are formed, which being of no further use to the body are removed from it by mechanisms prepared for the purpose. 285. Forms of Energy in the Body. — The force set free in the tissues by the combustion of their particles takes many forms within the body, but as it leaves the body it appears in two forms only : as heat or as work done by the muscles. Just as matter may be changed from one form to another, but cannot be destroyed, so energy, or force, may be changed from one form to another, but can- not be destroyed. The heat driven off in the combustion of coal may be changed througli the agency of steam into the motion of machinery, which may in turn be con- FOOD 203 verted into electrical force or light, or again become heat. A mechanic's blow upon an anvil converts muscular force into motion, heat, and light. Any form of energy must result from some other form of energy. It cannot come from nothing. 286. Conservation of Energy and Correlation of Forces. — These facts, in works upon the science of physics, are stated as the laws of conservation of energy and of the correlation of forces. The human body, like all other matter, is subject to these laws. As the power for a large proportion of our machines comes from the heat of combustion, — that is, the union of oxygen with other matter, — so does the energy of the body come mainly from the same chemical action, and the products of the slow combustion of food which takes place within the body are much the same as they would be if the food were burned in a furnace by swift combustion. Those products are carbon dioxide, water, and a nitrogen- containing substance which is discharged from the body as urea, besides certain salts not oxidizable which would appear in the furnace as ashes. 287. Food Elements. — The valuable parts of the matter which we call food enter in varying proportions into many different articles which we eat. Chemists divide these elements, which are essential to the maintenance of the body in health, into five classes, as follows : — 1. Proteids (Nitrogenous foods) 8 F^0^^68 [ (NonnitrogenoiiB) 4. Water ) 5. Salts f Inorganic Milk and eggs are examples of food containing all these materials in proportions suitable for young, grow- 204 UNCONSCIOUS NERVOUS OPERATIONS ing animals, and they are therefore spoken of as " perfect foods." 288. Characteristics of a Healthful Diet. — (1) A health- ful diet must contain all five chemical classes of food elements in due proportion. (2) It must be adapted to the climate, to the age of the individual, and to his mode of life, that is, to the kind and amount of work which he does. (3) Not only must the different kinds and amounts of necessary food appear in the diet, but they must appear in digestible form. 289. Classification of Foods. — With reference to the use made of them, foods are divided into three classes : (1) tissue builders ; (2) force generators ; (3) force regu- lators. The proteids constitute the first class, though we are not certain but that they may be sometimes oxi- dized, yielding force without having entered into the composition of living cells. In the second class are placed the oxidizable substances, — the fats and carbohydrates. They may also assist in forming tissue to some extent. The third class includes the inorganic compounds,— water, salts, etc., — and certain food accessories. 290. Proteids are the most important substances which en- ter into the animal organism, being absolutely essential to the phenomena of life. Plants are the great manufacturers of proteids ; no process has yet been discovered for making them artificially. Proteids are complex chemical com- pounds and are characteristically represented by the casein of milk and the albumin of egg. All are com- posed of carbon, hydrogen, nitrogen, and oxygen in various proportions, with a trace of sulphur. Proteids are the only group of foods which contain nitrogen, and whose chief if not sole purpose is the building of tissue. Animals can live without carbohydrates and fats, but not FOOD 205 without proteids. The chief proteids are the gluten of all cereals, peas, potatoes, beans, and lentils; the albu- min in white of egg, milk, and blood; globulin from yolk of egg and blood; myosin found in lean meat; casein in milk and cheese; fibrin in clotted blood. Though certain vegetables (as beans and peas) contain more proteid than does meat, they furnish it in a less digestible form, that is, a considerable part of it passes from the body unaf- fected by the digestive processes, and the proteid of these vegetables is therefore less valuable as food. Since only the proteids contain nitrogen, they are called nitrogenous foods ; and carbohydrates and fats are called nonnitrogenous. A healthy, well-fed animal is found to lose by excretion as much nitrogen daily as is supplied in his food. If the food contains an insufficient amount of nitrogen, the quan- tity excreted is greater than that received, and the tissues waste away. 291. Fats are found in butter, milk, cheese, and meat, in some of the grains, and in oils. They contain carbon, hydrogen, and oxygen. They are oxidized in the body, and furnish energy and heat. 292. The Carbohydrates, like the fats, contain carbon, hydrogen, and oxygen; but the proportion of oxygen is larger than in fats. The carbohydrates include (1) starch, which is found in cereals, vegetables, nuts, etc. ; (2) sugars of different kinds, — grape sugar, cane sugar, malt sugar, and milk sugar, besides sugar manufactured from starch; and (3) cellulose, found in fruits, cereals, and all vegetables. Carbohydrates are very rapidly oxidized in the body, pro- ducing heat, and it is they that are used up in the setting free of energy when the muscles are vigorously exercised; for it is found that the nitrogenous waste does not increase MACY'S PHYS. — 13 206 UNCONSCIOUS NERVOUS OPERATIONS in proportion to the increase of muscular effort. Carbo- hydrates cannot alone form tissue, but under some circum- stances may be used for that purpose with other elements. 293. Water, composed of oxygen and hydrogen, is found in all foods, and has a variety of uses in connection with nutrition, partly as a solvent for various elements in the food, and partly as promoting osmosis, and as an aid to the varied changes which take place in the tissues. Water uncombined with food, introduced into the stomach as in drinking, is all absorbed directly into the blood, of which it forms about 80 per cent. 294. Salts. — The food elements that are salts are chiefly the chlorides, phosphates, and carbonates of sodium and potassium, and to a less extent those of calcium and mag- nesium, with salts of iron and of some of the organic acids. Common salt (chloride of sodium) appears in all animal bodies, and to some extent in plants also. It helps to dis- solve certain of the albumins of the body, promotes the flow of the digestive fluids, and aids digestion in various ways. About half an ounce each day is needed with the food. That these saline matters are essential to health, is proved by experiments upon animals. When they are eliminated so far as possible from the diet, the central nervous sys- tem soon suffers, and paralysis results, besides general derangement of nutrition. When an animal body is burned, the various salts which entered into its composi- tion appear in the ashes which are left, while the other sub- stances have been changed into gases. 295. Food Accessories. — These are the various drinks — alcohol in different forms, tea, coffee, cocoa, etc. ; and condi- ments — mustard, pepper, ginger, and other spices, and a variety of other flavors added, not for their food value (though of that they may have a small amount), but to give FOOD 207 a pleasant taste which may assist digestion, and to stimulate the secretion of digestive juices. Used to excess, how- ever, all these drinks injuriously affect the nerves, and the condiments may irritate the sensitive lining membrane of the alimentary canal, besides impairing the delicacy of the sense of taste. 296. Food Values. — The nutritive value of a diet lies chiefly in the amount of nitrogen and carbon derived from it. A man of moderately active life will give off, mainly from the lungs as carbon dioxide, from eight to nine ounces of carbon each day. If he is engaged in severe muscular effort, he will give off much more carbon. The amount of nitrogen passing from the body (chiefly as urea) during a day is from .47 to .56 ounce, and with the increase of muscular activity it does not increase to nearly so great a degree as does the carbon. In order to repair the daily waste of the tissues, the proportions of carbon and nitrogen contained in the food should be the same as in the excretions, viz. about 16.6 to 1. While the proteids contain carbon, they contain only about 3.5 parts of carbon to 1 of nitrogen, hence other groups of food elements must be depended upon to supply the necessary carbon. The oxygen contained in food, being already combined with other elements, cannot be used in oxidation, so that from the lungs alone comes the needful supply of that Since the slow combustion within the body sets free the same amount of energy as does the rapid combustion of the same substances, by burning a quantity of food equal to that which a man eats in a day and measuring the heat given off, the amount of energy which that food can sup- ply may be estimated. In that way it has been shown 208 UNCONSCIOUS NERVOUS OPERATIONS that the energy from the fats is about equal to that from proteids and carbohydrates together. It has been esti- mated by a high authority that a healthful diet contains from three and a half to four and a half times as much of nonnitrogenous as of the nitrogenous foods. 297. The following table, from Landois and Stirling's Physiology, gives approximately the relative amounts of nitrogenous and nonnitrogenous elements in common articles of food, and shows that, next to human milk, wheat flour has most nearly the right proportion of the two elements. Beef and other kinds of flesh have too much proteid and should be eaten with potatoes or rice, which supply the nonnitrogenous matter needed to make the food complete. Vegetables contain too little nitro- gen to be used alone as food. Nitrogenous Nonnitrogenous Veal 10 to 1 Rabbit's flesh . . . . ~ 10 to 2 Beef . . . . . '.10 to 17 Beans . . . . . . 10 to 22 Peas . .. . .'j . . 10 to 23 Mutton . r / .. . 10 to 27 Pork . . ' * . 10 to 30 Cow's milk . ... . 10 to 30 Human milk . . . . 10 to 37 Wheat flour ... . . 10 to 46 Oatmeal . . . . 10 to 50 Rye meal . . ;' . . 10 to 57 Barley . . . < . . 10 to 57 Potatoes * . . . " 10 to 115 Rice ....... 10 to 123 Buckwheat . . „ . > 10 to 130 298. Variation in the Amount of Food Required. — When the body is at rest the amount of waste is much less than it is when the muscles are engaged in active labor. Then FOOD 209 the muscular tissue is rapidly broken down under the strong contraction constantly called for, and the circula- tion becomes swifter to supply material to rebuild the decomposed cells. Respiration, too, must be quickened to furnish a sufficient quantity of oxygen to arterialize the blood flowing faster to the lungs, loaded with the products of chemical changes in the tissue cells. Nerv- ous tissue also is worn away by the constant demands upon it for conveying impulses to the many muscles engaged and in coordinating all their related actions. If the body is exposed to a low temperature, a still larger demand is made for food to supply the greater loss of heat. During the period of growth a quantity of food is needed in excess of the waste products to furnish mate- rial for enlarging and strengthening all parts of the body. 299. Undigested Food. — Some (an average of about one tenth) of the food taken into the stomach, and especially a part of our vegetable food, seems to play no part in sup- plying nutrient material, but passes through the alimen- tary canal to be expelled from the body unchanged. It serves, however, as an aid to digestion by giving an increase of bulk to the food, and so assists the action of the digestive organs. 300. Cooking is the application of heat in one way or another to articles used as food. Most of our diet comes to the table after being submitted to this process, which renders it more wholesome and more palatable. By cook- ing, which lessens the cohesion of particles, the amount of work required of the digestive organs is reduced, and the chemical effects of heat prepare the various elements in the food to receive more readily the action of the diges- tive juices. Cooking also develops, especially in animal 210 UNCONSCIOUS NERVOUS OPERATIONS foods, the agreeable flavors which stimulate not only the appetite, but also the secretion of the digestive juices. Another very important effect of the application of heat in the cooking of food is the destruction of many of the germs of disease which are sometimes introduced into the human system with food. Certain diseases of animals whose flesh is eaten by man may thus be communicated to him if the meat is not first thoroughly cooked. Dis- ease is often conveyed also by means of the water supplied to a town or a dwelling, and in all cases where there is reason for suspicion respecting the purity of the water used for drinking, it should be boiled for at least half an hour before using. This sterilizes it. Ice should never be mixed with drinking water, on account of its impuri- ties. Sterilized water may be cooled for drinking pur- poses by inclosing it in tight cans of glass or metal and placing the cans in contact with the ice. Much of our food comes to the table badly prepared, cooked too mucli or too little, or in an unhealthful man- ner. The importance of good cooking cannot be overesti- mated; the lack of it is a fruitful source of the widely prevalent disease, dyspepsia, from which result innumer- able physical and mental ills. In recent years much attention has been paid to scientific cookery, and it is now taught in many schools. Only a few hints can be given here. 301. Cooking of Meats. — Fresh meat which is to be boiled should be put at first into water which is already boiling briskly. After fifteen minutes the kettle should be placed where, for the rest of the time, the water will only bubble slightly. By applying strong heat at first, the albumin of the surface of the meat is hardened and forms a close coating which retains the nutritive juices Avithin. FOOD 211 When soup or broth is to be made, this process is reversed — the object being to extract the soluble portion of the meat from the fiber. The meat is cut into small pieces and the bones are cracked. All is then placed in cold water without salt, and heated slowly to a tempera- ture just below the boiling point. It should be kept as hot as possible, without actually boiling, for from six to eight hours. Salt meat may be placed in cold water and gradually heated. Corned beef requires boiling for about five hours. Roasting and baking are done before an open fire or in an oven. The meat should first be browned by exposure to a very high temperature, in order to preserve the juices, and the heat should then be reduced. In broiling and frying the same principle applies. Frying is regarded as the least healthful of the various ways of preparing food, because the fat which coats the surface is supposed to be indigestible. Anything cooked by frying should be quickly and wholly immersed in fat so hot that the surface browns at once and further absorption of fat is prevented. Then the heat should be lowered. By stewing and braising, meats may be economically cooked at a moderate temperature, but the process re- quires several hours. In stewing, the temperature should not quite reach the boiling point. In braising, which is done in a closely covered pan in an oven, a higher degree of heat is applied. By these two methods the tougher and cheaper cuts of meat may be made entirely tender, nutritious, and appetizing. 302. Fish. — Principles similar to those suggested above apply to the cooking of fish. 212 UNCONSCIOUS NERVOUS OPERATIONS 303. Eggs. — Eggs are made most digestible by placing them (in their shells) in cool water, applying heat, and removing them from the water as soon as it boils. Or, they may be put into water already boiling, the vessel being covered and at once removed to where the water will keep warm but will not boil. They will be " done " in from ten to fifteen minutes, according to the weather and the amount of water used. In eggs cooked in this way, the albumen, which by boiling becomes hard and difficult of digestion, remains soft, creamy, and nutritious, while the yolk is partly solidified and rendered more palatable. 304. Cooking of Vegetables. — Fresh vegetables should be placed in boiling water only long enough to soften the fiber and cause the starch granules to burst. Too much cooking injures them. Dried seeds require longer boiling, and it is often well to soak them in water for several hours before cooking. 305. Bread is our most valuable food. It is made from the various grains, also from the flour of certain nuts and rootstocks. Salt, water, and yeast, in proper propor- tions, and sometimes a small quantity of sugar to hasten fermentation, are added to the flour or meal, and the mass is allowed to become " light " by the fermentation of the yeast before baking. Numerous chemical changes take place in the process. The water dissolves the gluten and sugar of the flour and swells the starch granules. If the dough is at a temperature of 100° F., fermentation at once sets in ; some of the starch becomes sugar ; sugar is converted into alcohol and carbonic acid ; the gas formed expands in little bubbles, which are surrounded by walls of sticky gluten, and "raises" the bread. If the fer- mentation is allowed to continue too long, a new chemical FOOD 213 product, called acetic acid, appears, rendering the bread sour and unwholesome. A few things are indispensable to the making of good bread. A good quality of flour must be used and good yeast (Miss Parloa considers the " compressed yeast " sold in the shops and bakeries the best procurable). The dough should have at first a temperature of 100° F., which should later be reduced to 70°. The mass must be kneaded sufficiently to distribute the yeast evenly throughout, and again after it has " risen " to break the bubbles of gas and force it to permeate every part, that the loaf may be rendered light and spongy by innumera- ble fine pores. After the final kneading and shaping into loaves, they should be left to rise to about twice their original size before baking. The oven should be heated to about 400°, but for the last half of the baking the temperature should be reduced to 300°. In baking, the process of fermentation is checked as soon as the loaf is raised to a temperature of 212°. The alcohol is vaporized and driven off, the starch granules burst, and by the transformation of starch into sugar and dextrine the delightful sweetness of good wheat bread is developed. Very large loaves are undesirable, as acetic acid may be formed in their interior after they are placed in the oven. Small loaves are better than large ones, also because they have a larger proportion of crust, which is the sweetest and most wholesome part of the loaf. Bread made of whole-wheat flour is especially valuable for children, because it contains more of the elements which are needed for making teeth and bone than does white flour. The same is true of what is called Graham flour, but the coarse bran which such flour contains is to some persons unwholesome. 214 UNCONSCIOUS NERVOUS OPERATIONS EXPERIMENTS Food Elements. — Some very interesting and easily performed experiments may be made upon substances common in ordinary food. 91. Tests for Proleids. — To one fourth of a test tube of dilute white of egg1 (a proteid) add a few drops of strong nitric acid and boil. Cool and add a little ammonia; the yellow color changes to orange. To show that the orange color is due to the presence of a proteid, repeat the experiment, using pure water instead of white of egg. Another test is as follows : To one fourth of a test tube of a 15 per cent solution of caustic soda add two or three drops of a 1 per cent solution of copper sulphate. Shake the mixture, and after warming add a little white of egg or other solution of proteid ; the blue color becomes violet. To show that these color changes are tests for pro- teids, repeat the above experiments, using a solution of sugar and the starch solution mentioned below, instead of the white of egg. 92. Coagulation of Albumin by Heat. — Boil dilute white of egg in a test tube; it does not clot. Then add, drop by drop, dilute acetic acid (2 per cent) ; a precipitate of coagulated albumin finally sepa- rates. If the white of egg is not diluted, it coagulates, of course, on boiling. 93. Solubility of Starch. — Stir some starch into cold water and observe that it does not dissolve. Boil the mixture ; solution occurs, but the liquid remains somewhat cloudy. 94. Test for Starch. — To some of the starch solution prepared above, add a drop of iodine solution. The blue color resulting is the characteristic reaction of iodine with starch. Dilute the starch solu- tion and repeat the experiment. The dilution may be much pro- longed before the blue color fails to appear on adding iodine. To show that the blue color is related to the presence of starch, add iodine to pure water or any liquid from which starch is known to be absent. 95. Microscopical Examination of Starch. — Scrape the fresh cut -sur- face of a potato, and mount in water on a slide some of the material thus obtained. Examine with the compound microscope. Numerous 1 To prepare dilute white of egg, beat up the white of an egg with about twenty volumes of water, filter through muslin, and pour off gently to remove air bubbles present. FOOD .-••- 215 small starch granules will be seen. Each granule shows concentric markings. Add iodine solution ; the granules turn blue or blue black. Examine other kinds of starch granules, of corn, oats, rice, etc. 96. Test for Sugar. — To one fourth test tube of a weak solution of glucose, add an equal amount of a 15 per cent solution of caustic soda. Shake, and after adding a few drops of a 1 per cent solution of copper sulphate, boil for a few minutes. The liquid changes from blue to yellow, or, if much sugar is present, to brown in color, and a precipitate is formed. To show that the change in color is due to the sugar present, repeat the preceding, using pure water, or any solution known not to contain sugar, instead of the sugar solution. This test is known as Trommer's test. It is not a test for cane sugar, nor does it distinguish between glucose, malt sugar, and milk sugar. 97. Fats. — Put some melted butter, or olive oil, into a test tube one fourth full of water. The fat rises to the top. Shake well ; a whitish mixture, or emulsion, is the result, but the oil and water quickly separate. To the contents of the test tube, add an equal amount of 1 per cent solution of carbonate of soda (an alkali), and shake. The emulsion with the alkali lasts much longer than that with water alone. Add a drop or two of oleic acid (a fatty acid) to the mixture, and shake well; the emulsion lasts longer than before. It will be learned later that the fats are only slightly dissolved in digestion, but are chiefly emulsified. 98. Examine with the microscope some of the emulsion prepared above. The fat will be found to be broken up into innumerable fine particles. 99. Shake some olive oil with dilute white of egg in a test tube ; an emulsion results. 100. To one fourth test tube of water, add a few drops of oleic acid, and shake. The oil rises to the surface as in the preceding- experiments. To the contents of the test tube add carbonate of soda as before. A white precipitate of soap is formed. An alkali and a fatty acid form soap. During digestion some of the fat is broken down into glycerin and fatty acids. The latter unite with alkalis in the intestine to form sol- uble soap. 101. Flour. — Boil a little flour in water and test with iodine for starch. 102. Place a little flour in from five to ten times its bulk of water 216 UNCONSCIOUS NERVOUS OPERATIONS in a flask and allow it to stand several hours, shaking it occasion- ally. Filter and test the nitrate for proteids, as in Ex. 91, and for sugar as in Ex. 96. 103. Shake some flour with ether in a test tube and allow it to stand for an hour or two, keeping the tube tightly corked and shak- ing it occasionally. Filter off the ether and place some of it on a clean glass surface and allow it to evaporate. A greasy residue remains, showing that the flour contained fat, some of which was dissolved out by the ether. 104. Gluten. — Moisten flour with water till it forms a tenacious dough. Tie it in muslin cloth and knead it in a vessel of water till all the starch is separated. There remains in the cloth a sticky, elas- tic mass of gluten, consisting of the insoluble albumins, some of the ash, and the fats. Draw out some of the gluten into threads, and notice its tenacity. 105. Milk. — Examine with the microscope a drop of fresh milk. It is seen to be an emulsion of oil globules floating in a liquid. 106. Warm some milk in a flask, and add a few drops of acetic acid. The mass clots and separates into a solid curd, consisting of casein and fat, and a liquid, the whey. Caseinogen is the chief proteid of milk. In curdling it is changed to casein. 107. Dilute milk with ten times its volume of water and slowly add dilute acetic acid. As long as the liquid remains alkaline, or neutral, — as can be tested with litmus paper, — no visible change occurs, but on adding more and more of the acid there is finally formed a precipi- tate of casein which, as in the preceding experiments, carries with it most of the fat. 108. Filter the curd from the whey obtained in one of the preced- ing experiments, and test the filtrate for sugar (milk sugar) and pro- teids (see Exs. 96 and 91). 109. Test, with litmus paper, some perfectly fresh milk. It will be found to be neutral, or alkaline. Allow it to stand in a warm place till it becomes sour and curdles. It will be found to be acid in reac- tion. In the souring of milk the milk sugar changes to lactic acid, and curdling is finally produced by the acid, as was illustrated in preceding experiments. 110. Lean Meat. — Mince finely some perfectly fresh muscles of a cat, dog, or rabbit. Place them in a large jar of water and stir. Tn about a quarter of an hour filter through muslin and place the muscle FOOD 217 in a fresh jar of water. Test the nitrate for proteids (serum albumin) as in Ex. 91. Repeat the washing till the nitrate gives no test of pro- teids. An hour or two of washing will suffice. Then squeeze out the water from the minced muscle, grind up the latter with clean sand, and add ten times its volume of a 10 per cent solution of common salt. Stir occasionally, and after an hour or more filter through mus- lin. Add some of the nitrate, drop by drop, to a large vessel of pure water ; there is formed a milky precipitate of myosin, the chief proteid of muscle. In the living muscle the myosin exists as the soluble myosinogen. Thus in muscle there are seen to be two kinds of pro- teids : one, serum albumin, soluble in water, the other, myosin, in- soluble in water, but soluble in weak salt solution. CHAPTER XVII THE DIGESTIVE APPARATUS AND NUTRITION 306. The apparatus for digestion consists of the alimen- tary canal with its appendages. This is a long, irregular tube having a continuous lining of raucous membrane. It comprises the mouth, pharynx, esophagus, stomach, and large and small intestines. Numerous glands along its length furnish the digestive juices. 307. The Mouth (Fig. 108) is the chamber which re- ceives the food through the opening and closing lips. The soft palate at the back forms a curtain over the opening at the back, while the hard palate of the roof, the soft muscular walls of the cheeks, and the large muscle of the tongue, together with the teeth, all assist in the process of mastication to which the food is first subjected. As in the skin, many minute papillce are found in the mucous membrane, containing networks of nerves and blood vessels. Some of these contain taste corpuscles. Some have organs for touch. 308. The Teeth are the special organs for cutting and grinding the food. Two sets" of teeth are provided. The first, which appear in infancy and are only twenty in number, are called temporary or milk teeth. They fall out after a few years, to be replaced by the permanent teeth, thirty -two in number. The four front teeth on each jaw are called incisors. 218 THE DIGESTIVE APPARATUS AND NUTRITION 219 Next to them come the canines, one on each side, then the two bicuspids, or premolars, and next to them three molars on each side. The third pair of molars on each jaw are called the wisdom teeth, and they sometimes fail to appear. Opening of Kustachiantube Soft palate Epiglottis Glottis Esophagus Larynx ~~ Hard palate — Tongue Hyoid bone False vocal cords • True vocal cords Thyroid carti- lage --"--==»•, Cricoid carti- lage Fig. 108. —Vertical section of the head and neck. The teeth of different animals are adapted 'in form and structure to the food upon which they subsist. Carnivo- rous animals are provided with strong, sharp teeth for seizing and tearing flesh, while the teeth of herbivorous animals are broader and relatively shorter, with wide- ridged surfaces for grinding grains and plant fiber. Man, as requiring both animal and vegetable food, is provided with teeth of both sorts. 309. Structure of a Tooth (Fig. 109). — A tooth has three parts, — crown, neck, androote. The crown is the part which projects beyond the gum, and is covered with the firm, 220 UNCONSCIOUS NERVOUS OPERATIONS solid layer of enamel, the hardest tissue found in the body. The neck is the narrowed portion just below the crown, surrounded by the gum. The root (one or more) is the re- mainder of the tooth, which is secured to the socket of the jaw by means of the perios- teum, through which it derives nourishment. A tooth is com- posed of a hard, close material called dentine, much like bone, but with less animal matter. This is molded round the cen- tral pulp cavity, which contains the tooth pulp — a mass of loose connective tissue, blood vessels, nerves, and cells of different shapes, one sort of which builds the dentine. A layer of true bone, called tooth cement, sur- rounds the dentine of that part of a tooth embedded in the gum, as enamel caps the crown. 310. The Tonsils. — Between the arches of the soft palate lies on each side a soft rounded body covered with mucous membrane and containing many small glands which se- crete mucus. They are called the tonsils (Fig. 108). Their use is unknown except that they furnish some pro- tection to the larynx and pharynx. Being sometimes permanently enlarged and subject to frequent inflamma- tion, they are occasionally removed by the surgeon with apparent advantage to the patient. 311. The Pharynx (Fig. 108) lies behind the soft palate. Fig. 109. - Diagram of the structure of a tooth. THE DIGESTIVE APPARATUS AND NUTRITION 221 It has muscular walls, mostly of voluntary fibers, which contract upon the food to push it into the esophagus, or may force it back into the mouth if desired. Seven passages open into this cavity: the mouth, the two nasal passages, the two Eustachian tubes, the larynx, and the esophagus. 312. The Esophagus connects the pharynx with the stom- ach (Figs. 108 and 110). It is a muscular tube lying along the spinal column behind the trachea. The muscu- lar coat of the wall of the esophagus contains an external layer of fibers running lengthwise, while the second layer is of circular fibers. They are mainly involuntary and supplied by the vagus nerve, with fibers from sympathetic ganglia also. 313. The Stomach lies in the cavity of the abdomen immediately below the diaphragm (Fig. 86, p. 153). It is a large sac, formed by the dilation of the alimentary canal (Fig. 110), and its walls have the three coats of the rest of that canal, — the inner mucous coat, the middle connective tissue coat, and the external muscular coat, with a fourth coat in addition, the peritoneum, which forms the lining of the abdominal cavity and is reflected back over the organs (or most of them) which it contains. The peritoneum adheres to the walls of the abdominal cavity, folds of it surround the blood vessels running to the stomach, and a large pouch of the same forms a sling for the stomach. From the lower side of the stomach another large fold of the peritoneum, called the great omentum, spreads over the rest of the abdomen. After middle life it often holds a large accumulation of fat. The muscular coat of the stomach is composed of layers of unstriped muscular fibers. Its mucous lining is inelas- tic, and as it lines the organ smoothly when it is distended, MACY'S PHYS. — 14 222 UNCONSCIOUS NERVOUS OPERATIONS it must lie in folds when the stomach is empty and shrunken (Fig. 110). The blood supply of this mem- brane is very large during digestion, and its appearance is then much redder than at other times. The two open- Fig. 110. — Stomach and duodenum. The anterior walls are cut away to show the folds of the mucous membranes rugne of stomach, and valvulae conniventes of intestine. ings of the stomach are both upon the upper side. That near the left end, the cardiac orifice, admits the food from the esophagus; that on the right, the pyloric orifice, con- nects with the small intestine. The latter is surrounded THE DIGESTIVE APPARATUS AND NUTRITION 223 by a thick ring of circular muscular fiber, forming a sphincter muscle, which keeps the opening closed except when food is ready to pass from the stomach. 314. Nervous Supply of the Stomach. — Nerve fibers from many centers reach the different coats, glands, and blood vessels of the stomach and those of the large and small intestines. They are gathered in large tangles of nervous matter, called plexuses, which contain branches from the vagus nerves, from many of the spinal nerves, and from the ganglia of the sympathetic chain. The great solar plexus, or epigastric plexus, is placed at the pit of the stomach ( Fig. 22, p. 31) ; the hypogastric plexus lies before the last of the lumbar vertebrae, and divides into two parts, one lying on each side of the rectum. These plexuses give rise to innumerable branches which control the complicated processes of digestion, the precise path of each different sort of nervous influence not having been yet made out. It is easy to see, however, that this close nervous connection of the digestive organs with all parts of the system implies important relations between them. Vagus nerve fibers appear to stimulate the peristaltic or wavelike movement of the stomach and bowels, which by the progressive narrowing of the tube forces on its contents, while the sympathetic fibers are inhibitory and bring the movements to an end. It is believed that the walls of these organs possess some power of spontaneous action such as appears in the walls of the heart. 315. The Small Intestine ( Fig. Ill ) is a tube with many curves, about twenty feet in length, two inches in diameter at its upper end and somewhat smaller in its lower por- tion. Its coats are the same as those of the stomach, but the peritoneum, a fold of .which forms the outer coat, does not entirely surround the tube, but runs off to form a sup- 224 UNCONSCIOUS NERVOUS OPERATIONS porting membrane for the intestine, called the mesentery. This is gathered up and connected with the wall of the abdominal cavity near the spinal column. The blood vessels and nerves pass through the mesen- tery to reach the intestine, and the absorbents also pass through it from the intestine. The two layers of muscular fibers, one circular and one longitudinal, in the intestinal wall, are of the unstriped, involuntary kind. 316. Nerves of the Small Intes- tine. — Between the muscular layers is a plexus of nerve fibers W^] lnany ganglia ;i11(l i]l tin; layer beneath the mucous mem- brane of the lining is another nerve plexus, also gangliated. 317. The Mucous Membrane of the Small Intestine has a very important part in the function of digestion, and is of peculiar structure. Like the lining mem- brane of the stomach, it has an inner layer of columnar epithe- lium, but unlike that of the stomach the lining of the intes- tine is for a large part of its length laid in folds which do not disappear when the canal is are called valvulce conniventes (Fig. 110). They lie around the inner surface of the Fig. 111.— The stomach and intestines. 1 stomach. 2 duodenum. 3 small intestine. 4 termination of the ileum. 5 caecum. <5 vermiform appendix. 7 ascending colon. 8 transverse colon. 9 descending colon. 10 sigmoid flexure of the colon. 11 rectum. 12 spleen. 13 anus. distended. These folds THE DIGESTIVE APPARATUS AND NUTRITION 225 intestine, each separate valvula reaching about one half or two thirds the way round. Their function is to fur- nish a large amount of secreting and absorbing surface. 318. The Villi. — Still further to increase this surface, the valvulse are covered with villi. These are minute projections from one fiftieth to one eighth of an inch Fig. 112. —Diagrammatic representation of a small area of the mucous membrane of the small intestine. 1 cellular structure of the epithelium, or outer layer. 2 a vein. 8 fibrous layer. 4 villi covered with epithelium. 5 a villus in section, showing its covering of epithelium, and its blood vessels and lymphatics. 6 a villus partly uncovered. 7 a villus stripped of its epithelium. 8 lymphatics or lacteals. 9 orifices of the glands opening be- tween the villi. 10, 11, 12 glands. 13 capillaries surrounding the orifice of a gland. in length, and are found only in the small intestine (Fig. 112). They give to its mucous lining a peculiar velvety appearance. Each vittus is covered with columnar epithe- lium resting on a fine membrane. Within are blood vessels 226 UNCONSCIOUS NERVOUS OPERATIONS (two or more arteries, a dense network of capillaries, and one or two veins), connective tissue fibers, and a single lymphatic or lacteal ves- sel, which may be looped or branched (Fig. 113). Between blood vessels and lymphatics is a very thin layer of fine muscu- lar fibers, which help to propel the chyle along the lacteals. Fine nerve fibers are also found. In the villi the digested food passes through the cells of the thin wall into blood vessels and lacteals. 319. The Large Intestine (Fig. 111).— The small intestine passes into the large intestine, the ileo- ccecal valve at the junc- tion preventing any re- Fig. 113. — Diagram of the essential parts of a villus a epithelium winch takes up food and transports it to the tubes within. b an artery, c capillaries. d a lactea^ flux of the contents from the large to the small tube. The large intestine has three parts, called the caecum, the colon, and the rectum. The caecum is a large sac on the right side, which receives the contents of the small intestine. Attached to the lower side of the cjecum is the small vermiform appendix, which has no known function, but is regarded as a " survival " from a previous type of animal structure. Continuous with the ciecum is the colon, having ascend in; j, transverse, and THE DIGESTIVE APPARATUS AND NUTRITION 227 descending parts. The rectum is the final portion of the alimentary canal ; it opens externally at the anus. The walls of the large intestine are like those of the small intestine, except that the valvulse conniventes and the villi are wanting. In the caecum and colon the longitudi- nal muscular fibers are for the most part collected in three bands, which, being shorter, from end to end, than the other coats, draw up the intestinal wall into puckers, or folds. The muscular coat of the rectum is much thicker than elsewhere, and at the anus is a strong band called the internal sphincter muscle. 320. Secretion. — All the living cells of the body are engaged in taking from the blood certain substances suit- able for their own special purposes, and returning to the blood those particles of matter which have fulfilled their mission and are no longer of use. Every cell requires oxygen, and oxidation is now understood to occur within all the cells. But certain cells or groups of cells take up also other substances from the blood, and manufacture within themselves a new product having a special function. This process is called secretion, and the or- gans of secretion are called glands. 321. Glands. — The sim- plest glands are merely minute tubes lined with cu- bical cells (Fig. 114). Sometimes these tubes branch at the inner end, all the branches being lined with the secret- Fig. 114. — Structure of glands. 1 simple pit, surrounded by capillaries. 2 flask-shaped gland, with short duct. 3, 4 more complex glands, with longer ducts. 228 UNCONSCIOUS NERVOUS OPERATIONS ing cells, and all uniting in a single tube, or duct, through which the secretions pass. Frequently a large number of branches from a secreting tube are grouped together in clusters to form an organ of considerable size, as the liver or the pancreas. The secreting surface is always composed of living cells, and the processes carried on in them are similar to those occurring in other cells. That is, secretion involves building up, or growth, and breaking down, or waste, of the cell substance, along with other changes. Under the microscope the cells of the glands are seen to contain a nucleus and many granules. These granules are products of the cell itself. When the secreting process is going on, water and other substances pass through the cells from the blood, and at the same time the granules are dissolved in the water and pass out along the secretory duct. Secre- tion is a manufacturing process, and not merely a filter- ing out of certain substances from the blood. In each gland the chief or specific constituents of its peculiar juice are formed in the cell and not simply extracted from the blood. The mucous membrane of the whole of the alimentary canal is largely made up of glands. 322. Salivary Glands (Fig. 115). — There are three pairs of salivary glands. Those lying in front of each ear are the parotid glands; those under the lower jaw on each side, the submaxillary glands; and those under the tongue, the sublingual glands. They are large glands whose ducts pour their watery secretions into the mouth. 323. Nervous Action upon the Salivary Glands. — Ordina- rily the nervous action affecting the salivary glands is reflex. The organs of taste are stimulated by food, or the sight or odor of food stimulates the optic or olfactory THE DIGESTIVE APPARATUS AND NUTRITION 229 nerves, the nervous center in the brain sends impulses to the special center in the medulla oblongata from which efferent secretory im- pulses are reflected, and they pass along the fibers of a branch of the seventh cranial nerve, which probably contains fibers from the ninth nerve, with which it communi- cates. The impulse finally reaches the cells of the submaxil- lary and sublingual glands, and a flow of saliva results. Even the thought of food, by stimulating a center in the brain, may produce nervous impulses having the same effect. Nerve branches from the sympathetic system also carry stimulus to these glands. For the parotid glands the chief secretory nerve fibers arise in the glossopharyngeal nerves (ninth cranial). 324. Action of Saliva. — A part of the digestion of food takes place in the mouth. Saliva is mixed with the food by mastication, and serves to moisten the mass and lubri- cate it for swallowing. It also causes a chemical action, due to the presence of its active principle, ptyalin, which affects the starch in food, converting it into malt sugar. If one chews slowly a few grains of wheat, he will notice that the paste becomes sweet. This is because some of the starch in the wheat is changed by the saliva into sugar. Fig. 115. The salivary glands of the right side. 230 UNCONSCIOUS NERVOUS OPERATIONS 325. Ferments. — The ptyalin belongs to a kind of sub- stances called ferments. Of these there are two classes, — the organized and the unorganized. The organized ferments are living vegetable organisms. Yeast is one of them, the vinegar plant is another, and the various kinds of bacteria, some of which cause disease, are other examples. The unorganized ferments are chemical sub- stances resulting from the activity of living cells, and capable of effecting certain changes in particular sub- stances. The ptyalin is one of these. The ferments are the agents for effecting most of the changes which the food undergoes in digestion. 326. Gastric Juice. — The epithelium of the lining of the stomach consists of a single layer of cells, and the mucous membrane is almost entirely composed of simple tubelike glands closely packed together. When food reaches the stomach, more blood is sent into the dilated blood vessels, and the glands make from the blood a colorless fluid called gastric juice, which flows into the cavity of the stomach. Chemical analysis shows that gastric juice contains, besides water, a small amount of salts, a little free hydro- chloric acid, and two of the ferments, called pepsin and rennin. Rennin is that element in the gastric juice which causes milk to curdle. The use of rennet (which is ob- tained from the stomach of a calf) in the making of cheese depends upon the presence of this ferment. Pepsin is the ferment which is able to change proteids into soluble form, and to make diffusible such as are not already so. The ferments of the gastric juice act only in the presence of an acid, and to assist their action seems to be the func- tion of the hydrochloric acid. . 327. The Food in the Stomach. — The effect of the gastric juice upon the food in the stomach is to make the whole THE DIGESTIVE APPARATUS AND NUTRITION 231 mass acid. This destroys the ptyalin, and no more starch is converted into sugar. As the saliva acts only upon starch, so the gastric juice acts only upon the proteids. By the muscular movements in the walls of the stomach the food is moved from side to side and thoroughly mixed with the gastric juice, and the mass becomes semifluid. The fats and carbohydrates remain unchanged, except as affected by the warmth of the stomach and by the removal of the proteids which are dissolved out of the mass. 328. Digestion in the Small Intestine. — The food by re maining in the stomach from one hour to three or four, is converted into what is called chyme. The sphincter muscle of the pyloric orifice relaxes at intervals, and the chyme is then passed on into the small intestine by the contractions of the wall of the stomach. Here it soon encounters two other juices, the bile and the pancreatic juice, *>y which it is still further changed. 329. The Pancreas is a long, slender gland, enlarged at its right end, lying back of the stomach and along its greater curvature, and supported by the mesentery (Fig. 94, p. 163). It is pinkish yellow in color, and resembles the salivary glands in structure. A duct runs from one end to the other, joins the common bile duct from the liver, and passes with it obliquely through the wall of the small intestine (Fig. 110). 330. The Pancreatic Juice. — The pancreas has reflex nervous connection with the stomach, and as soon as food enters the latter, secretion begins in the pancreas, and the secreted fluid accumulates in the small intestine. The pancreatic juice is a clear, somewhat viscid, alkaline fluid, containing many different substances, the most im- portant being the ferments. Of these there are four. One, similar to pepsin, but able to act only in an 232 UNCONSCIOUS NERVOUS OPERATIONS alkaline medium, affects the proteids more rapidly and more powerfully than pepsin, and so digests those which were not sufficiently changed by the gastric juice. Another ferment in the pancreatic juice acts like the saliva upon starch, converting it into malt sugar, but its action is far more powerful. This ferment is not present in the pancreatic juice of infants, and they are therefore unable to digest starchy foods properly. The pancreatic juice has two different effects upon the fats, they having hitherto been unchanged. The first effect is to separate them into exceedingly small particles, which can pass through the walls of the intestine, that is, the juice forms an emulsion with the fat. The second effect is a chemical decomposition of fat into fatty acid and glycerin by the action of the third of the pancreatic ferments. The acids set free unite with the alkaline sub- stances present to form soaps. • The fourth ferment possesses the power of curdling milk, as does rennin, though its action is not identical with that of rennin. It is able to act upon any particles of milk which have by any possibility escaped the influ- ence of the gastric juice. 331. The Secretory Nerves of the Pancreas have been found to be fibers of the vagus, or tenth cranial nerve, which, as already mentioned, are stimulated by efferent impulses ex- cited in the brain by afferent impulses from the stomach. 332. Other Functions of the Pancreas. — In addition to its office in connection with digestion, experiments have proved that this gland has some further influence upon the general condition of the body ; but what that influ- ence is, is as yet unknown. 333. The Liver, which weighs from fifty to sixty-four ounces, is the largest gland in the body. It lies chiefly THE DIGESTIVE APPARATUS AND NUTRITION 233 on the right side, immediately under the arch of the dia- phragm (Fig. 86, p. 153). The peritoneum, without entirely covering it, adheres closely to it and attaches it to the diaphragm and other parts. By a deep fissure it is separated into right and left lobes (Fig. 116). The tissue elements of the liver are the hepatic cells, and all the other parts contribute to their support, protection, Fig. 116. — The liver seen from below. and nutrition. The hepatic cells are grouped together in little masses to form lobules, which are permeated by the blood capillaries from the portal vein and the hepatic artery. From these capillaries the blood passes to a small vein which unites with those from other lobules to form the hepatic veins, which empty into the inferior vena cava. 334. Blood Supply of the Liver. — The liver has an im- mense supply of blood. That which is brought by the 234 UNCONSCIOUS NERVOUS OPERATIONS portal vein has first circulated through the stomach, the spleen, and the intestines (Fig. 95, p. 165), and has been deprived of most of its oxygen. The supply of arterial blood through the hepatic artery is relatively small. 335. Functions of the Liver. — One of the functions of the liver is to store up sugar, or, rather, to form from the sugar brought by the portal vein a substance called gly- cogen, which is readily converted again into sugar. The protoplasm of the hepatic cells possesses the power of manufacturing glycogen from carbohydrates and of stor- ing it up. This is effected by means of a ferment found in the liver ; but what is the part played by glycogen in the animal economy is not fully determined. Glycogen is turned into sugar again and supplied by the liver to the blood as it is needed. There is always sugar in the blood, but more than a small amount seems undesirable. There is much evidence that another function of the liver is to act upon the nitrogenous foods in some manner resulting in the production of the waste substance, urea, which is eliminated by means of the kidneys. A third and the most familiar office of the liver is the formation of bile. This is stored in the gall bladder, lying between the two lobes on the under side of the gland (Fig. 116). Still another function of the liver is to protect the system from the action of poisons produced by the processes of digestion or by defective digestion. These are arrested or transformed when they reach the liver. 336. The Bile is secreted from the portal blood as a yel- lowish, reddish brown, or green fluid, according to the pre- ponderance of different coloring matters which are formed by the liver out of the hemoglobin of the red blood cor- puscles. It is alkaline and is thought to have antiseptic properties. Bile is a poison, and in one person enough is THE DIGESTIVE APPARATUS AND NUTRITION 235 secreted in a day to kill three men ; but it is neutralized in the intestines, in the liver, in the tissues, and in the blood. Upon the food with which it mixes in the small intestine bile seems to have little effect, and its function there is supposed to be mainly to assist the pancreatic juice in neutralizing the acid in the chyme, and in its further action. Secretion of the bile goes on continu- ally, but it passes into the intestine only when digestion is proceeding. At other times it is carried by a side branch of the bile duct to the gall bladder and there stored. About two pints of bile is secreted daily. 337. The Intestinal Juice is a secretion of the mucous membrane of the small intestine itself, and contains a ferment which changes cane and malt sugars into glucose, besides having possibly further action upon the food. 338. Bacterial Action. — There have been found to exist in the alimentary canal certain microorganisms which modify the digestive processes. Bacteria are especially numerous in the intestines. The antiseptic gastric juice destroys bacteria, or neutralizes their action in the stomach; but some escape into the intestines, where they multiply. Their presence is shown to be, under normal conditions, beneficial, though certain forms of bacteria produce disease. 339. The Peristaltic Movements of the Intestines. — By suc- cessive contractions and relaxations of the muscular fibers in the two muscular layers of the intestinal wall, a wave- like motion passes along the whole length of the organ and forces the contents, from the constricted to the relaxed portion, slowly on into the large intestine, bringing the different substances in contact with the absorbing cells in the mucous membrane. 340. Function of the Large Intestine. — Most of the food which can be used in the body is absorbed in the stomach 236 UNCONSCIOUS NERVOUS OPERATIONS and small intestine, so that what is left to pass into the large intestine is indigestible matter and the remains of the juices. The tubular glands in the mucous membrane here absorb what is left of nutrient material and a con- siderable amount of water, which is carried by the small veins to the portal vein. The residuum, becoming more solid as the water is absorbed, passes into the rectum and is discharged as faces. The chief alteration in the con- tents of the large intestine is due to the action of micro- organisms which cause chemical changes, giving rise to organic acids. 341. Absorption. — When the food has been digested it is ready to be absorbed. It cannot be used for the sup- port of the body until it reaches the blood, and there are two ways by which it may reach the blood. First, it may be absorbed directly into the blood. The mucous lining of the alimentary canal is richly supplied with absorbing cells. Little of the food, however, is taken up in the mouth and esophagus, because of the thickness of the epithelium in those cavities, and because the food passes quickly through them. In the stomach, and especially in the small intestine, the absorbents are very numerous, and a large amount of digested food passes directly into the blood stream by the capillaries through the columnar cells of their epithelium ; while the walls of the large intestine also absorb food, but to a less extent. This direct pas- sage into the blood is now regarded as the more important of the two channels of absorption. The living cells, while allowing water and soluble salts to pass through them unchanged, effect certain changes in the organic food materials while in contact with them. The carbohydrates and proteids are mainly absorbed into the blood in this immediate way. THE DIGESTIVE APPARATUS AND NUTRITION 342. The second agent of absorption is the lymph cells in the tissues of the walls of the alimentary canal. It is chiefly by the lymphatics that the fat of the food gets into the circulation. Two changes are effected in the fats by the digestive juices in the small intestine to prepare them for absorption by the lymphatic vessels, which are in the intestines called lacteals, because the presence of fat ren- ders the fluid they contain milky. First, by the mixture of the bile and pancreatic juice with the food an emulsion is formed ; that is, the fat is broken up into minute particles, which float in the liquid, giving it a milky appearance. Secondly, a chemical union takes place between some of the acid fats and the alkaline secretions of the digestive organs, — a process called saponification, which is the making of soap. The digested food in its milky form is called chyle, and this is collected in larger and larger tubes, and finally poured by the great thoracic duct into the blood of the jugular vein. Being then carried to the right auricle of the heart, it passes to the right ven- tricle and thence to the lungs, in whose capillaries it meets with the oxygen from the air, and having returned to the left side of the heart is distributed throughout the system. 343. The Lymphatic Vessels are delicate tubes which drain the spaces between the cells of the tissues, gradu- ally uniting to form the main lymphatic vessels, of which the thoracic duct is the largest. These vessels are sup- plied with numerous valves, like those of the veins, to prevent the reflow of their contents, and the opening of the thoracic duct is guarded by a valve. The flow of the fluid is kept up by the pressure in the capillaries, which is greater than is the pressure in the veins into which it is emptied, and by the movements of the body, which MACY'S PHYS. — 15 238 UNCONSCIOUS NERVOUS OPERATIONS constantly cause pressure upon the tissues and so force the lymph onward in the vessels, the valves preventing any return. 344. Assimilation. — Though the food has been masti- cated, digested, and absorbed, it has not yet nourished the body. Still another process is needful before the new material becomes part of the continually wasting tissues. That process is called assimilation, and, though we cannot pretend to understand it, it may be described as the action of the living cells in choosing, appropriat- ing, and building into their own substance the suitable elements in the food-laden fluid which comes to them from the alimentary canal and from the lungs. Correlative to the process of assimilation is the destruc- tive process by which the cells, by combustion and other chemical changes, break up and send out as waste the substances of their structure, to be expelled from the body as excretions. 345. Hunger and Thirst. — We associate our feelings of thirst with a dry ness of the mucous membrane of the mouth and throat, and we say our throats are " parched " when we are very thirsty. But under ordinary circum- stances the feeling of thirst arises from a general con- dition of the system, in which the throat shares, due to a lack of water in the blood, or rather in the lymph. Thirst may be temporarily relieved by moistening the mucous membrane of the soft palate. Hence follows the inference that the afferent nervous impulses originate there, and are caused by a too great removal of water from the lymph of the investing membrane. Hunger is referred in our consciousness to the par- ticular locality of the stomach, and that organ seems to us to be empty when we are hungry. Indigestible mate- THE DIGESTIVE APPARATUS AND NUTRITION 239 rial introduced into the stomach may for a time relieve the hunger, as will a very small quantity of food. The special sensation of hunger appears to be connected with the state of the lining membrane of the stomach, while it must be ascribed in a more general sense to a deficiency of nutrient matter in the blood. Hunger may be alle- viated by the introduction of soluble food into the circula- tion, through the rectum, or through the absorbents of the skin, but the relief comes more slowly thus than through the stomach. The nervous path of hunger sensations has not been made out. The vagus is regarded as the sensory nerve of the stomach, but it is said that both vagus nerves may be cut and the sensation of hunger be unaffected. The brain centers for thirst and hunger are believed to be in the occipital lobes of the cortex, but they have not been definitely located. 346. Some Practical Points connected with Nutrition. — In order that the first of the digestive operations may be properly performed, it is necessary to have a good set of teeth and to chew thoroughly the food taken into the mouth. The intensely hard enamel covering the exposed portion of the teeth is a full protection to them against all dangers .under proper conditions of life, and under such conditions the teeth would last while life lasts. That this is true is shown by the fact that nature makes no provision for restoring or improving the enamel after it is once formed. Here alone the special cells, whose office it is to form the peculiar substance, entirely disap- pear when their work is once completed. In all the other tissues these formative cells remain to continue the nutri- tion and repair of the tissues. But the tooth enamel, whose growth, except in the wisdom teeth, is complete 240 UNCONSCIOUS NERVOUS OPERATIONS when a child is ten or eleven years old, cannot be renewed or improved after that time. It is therefore of the utmost importance that young children should be fed upon food which will build up perfect teeth. Milk should be largely relied upon for the first three years, the diet to be varied during the third and after years in accordance with sug- gestions given in the chapter on Food. Great care should be taken to guard children against attacks of what are known as " infantile diseases," — measles, whooping cough, etc., which sometimes suddenly arrest or disturb the gen- eral nutrition, and especially that of the teeth, so that the enamel becomes rough and irregular, and the teeth are exposed to early decay. Another point should receive special attention. It is observed that young children who live a life of excessive nervous activity, with over- stimulation of the brain, are particularly liable to defect- ive development of the enamel of the teeth. This is one among many reasons which make imperative a quiet, regu- lar life for children, without excitement and without undue mental activity. 347. But even perfect teeth may be injured by certain bacteria, which multiply in the decaying particles of food allowed to remain in the mouth. These minute organisms form a corrosive acid which destroys the enamel and breaks down the tooth substance. If the teeth are perfect and are always kept perfectly clean, they will not decay. They should be thoroughly brushed — the upper teeth downward, the lower ones upward — after each meal, and a thread of soft untwisted silk floss or fine strips of rubber should be drawn back and forth between the teeth to cleanse those parts which a brush cannot reach. In brushing the teeth a powder or liquid should be used which contains some safe germicide, — which is a substance destructive to THE DIGESTIVE APPARATUS AND NUTRITION 241 the microorganisms mentioned above, — and the mouth should be well rinsed with a solution of the same. 348. It is a mistake to suppose that a child should be supplied with soft, pulpy food. Just as soon as the first set of teeth are in place, he should have a fare which will require vigorous mastication. He should not be allowed to reject bread crusts and eat only the soft crumb, nor should his bread be always soaked in milk or gravy. Plenty of hard " chewing " is not only good for the teeth; it also promotes the flow of the saliva necessary to diges- tion and aids in the development of the jaws, and so helps to provide room for the second set. The teeth should be under the care of a competent den- tist, who by yearly or seiniy early examination and repair will be able to forestall and prevent the inroads of decay. 349. While a due action of the mechanism of mastica- tion is to be sought, that overactivity which results from the habit of chewing tobacco or gum is to be avoided. While a sense of propriety and good taste should alone be suffi- cient to condemn such a habit, there are hygienic rea- sons for its avoidance. The constant stimulation of the salivary glands leads finally to their weakness and defect- ive action, thus laying a foundation for general derange- ment of digestion. Many dentists also regard it as directly injurious to the teeth. 350. Food is not ready for the action of the gastric juice until it has been finely divided by the teeth and all portions well moistened with saliva. Slow and thorough mastication is therefore necessary to perfect digestion. Too rapid eating not only shows bad manners, but also is exceedingly bad for the health. 351. The temperature of our food should not be so hot as to stimulate unduly the glands of the mucous mem- 242 UNCONSCIOUS NERVOUS OPERATIONS brane of mouth and stomach, nor so cold as to retard the digestive processes, which normally require a temperature of about 100° F. (38° C. ). Ice water should never be drunk, both because of the impurities usually found in the ice, and because its coldness is injurious to the stom- ach. Very large quantities of any liquid taken with the food may dilute the gastric juice so much as to delay digestion and weaken the organs. 352. It is well to establish and adhere to regular hours for meals. The intervals between meals should be long enough to permit the digestive organs to rest between their periods of activity, and fresh food should not be taken into the stomach to mix with that partly digested ; that is, "eating between meals" is to be avoided. A habit of continually nibbling at dainties is extremely per- nicious, and may give rise to serious and perhaps incur- able disease. 353. A considerable variety in diet is wholesome, but as a rule one should adhere to the simpler and more easily digested kinds of food. A person in health is scarcely conscious of possessing a stomach, but injudicious indul- gence may so disorder the natural processes that they will be constantly attended with discomfort or suffering. 354. It is impossible to prescribe definite rules for the quantity of food to be taken daily. A strictly natural appetite is undoubtedly a safe guide ; but appetite is so often and so early perverted that it is seldom reliable. Food enough must be taken to supply the daily waste of tissues. Continuous loss of weight is usually a seriously unfavorable symptom. During the natural period of growth the amount of food must be sufficient to supply also what is needed for the full development of the body. One living a life of physical activity requires, as a rule, THE DIGESTIVE APPARATUS AND NUTRITION 243 more food than one engaged in sedentary occupations. Brain workers, however, need a varied and generous diet, and along with it gre"at care should be taken to secure sufficient outdoor exercise. More food is called for in winter than in summer, and more of the carbohydrates to supply the demand for additional heat. Those who work vigorously in the open air, and especially in cold climates, often consume prodigious quantities of fats without injury to digestion. With the coming of old age the vital proc- esses in general are carried on more slowly ; digestion and especially the power of assimilation are enfeebled. Less energy is called for as the activities are lessened, and less food is then required, with longer intervals between meals. Foods rich in proteids are less needful and should be diminished in quantity, while those which yield a large amount of heat should be substituted. 355. What is Alcohol? — All organic bodies are subject to decay ; the complex compounds of which they are composed are broken up into simpler ones, and that which was living, organized matter becomes lifeless and inorganic. This destruction of organic tissue is due under ordinary circumstances to the process called fer- mentation in some one or more forms. This is the growth and rapid multiplication of minute organisms, of which yeast is the most familiar example. When the decom- position of organic matter takes place under certain con- ditions and reaches a certain stage, it is called putrefaction. This is always attended by the multiplication of the low forms of life known as bacteria, and by the production of poisonous and ill-smelling gases. Another form of fer- mentation is that which occurs in the juices of fruits, grains, and vegetables which contain sugar; and is called vinous fermentation. In this form of decomposition the 244 UNCONSCIOUS NERVOUS OPERATIONS fungus known as the yeast plant is the active agent in producing the changes which occur. The sugar of the fruit or plant must be in solution, and the germs of the yeast must in some way be introduced. Alcohol is one of the products of vinous fermentation. It is composed, like sugar, of carbon, hydrogen, and oxy- gen, but in proportions different from their proportions in sugar. The drinks which contain alcohol differ widely in flavor according to their sources, and also vary in the amount of alcohol which appears in them. In cider and some kinds of beer the proportion may be as low as 2 per cent. As alcohol is a very volatile fluid, it may be readily separated from the other substances in the fermented liquor by the process called distillation. This is the driv- ing off of the alcohol in the form of vapor by the applica- tion of heat, and its recondensation, by cooling, to liquid again. In this way is obtained the strong alcohol which is mixed with various coloring and flavoring matters to form the spirituous liquors of commerce. Whether a drink contains the 2 per cent of alcohol found in cider, or the 50, or more, per cent found in whisky, or the 90 per cent of " cologne spirits," the alcohol is in all cases identical in its nature and properties. 356. Properties of Alcohol. — Pure spirit, or "absolute alcohol," is a colorless, volatile liquid with a strong affin- ity for water, which it rapidly absorbs from the atmos- phere or from any other substance containing water with which it comes in contact. Alcohol burns readily in the open air, that is, it is quickly oxidized and changed in its chemical composition. It is a powerful solvent, dissolv- ing many substances not soluble in water. Though itself the product of fermentation, it is a preventive of putre- faction.— that is, it preserves animal tissue from decay7 THE DIGESTIVE APPARATUS AND NUTRITION 245 — and introduced in sufficient amount into a liquid in a state of vinous fermentation it destroys the power of the yeast plant to multiply, while in smaller quantities it retards the growth of the living cells in direct proportion to its amount. When by the decomposition of sugar in vinous fermentation the amount of alcohol produced has reached 14 per cent, no further growth of the yeast takes place. That proportion of alcohol destroys the vitality of the living cells. It is thus useful as an antiseptic. Brought in contact with food elements outside the body, alcohol is found to harden them by abstracting the water which they contain, and to coagulate the albumin, which is thus rendered insoluble. 357. Is Alcohol a Food ? — Alcohol contributes nothing to the formation of tissue, and cannot, therefore, be classed in the first division of foods, according to the definition given in § 289. As it is now proved that when taken into the stomach in dilute form and in small quantities it may be fully oxidized, producing energy, it must be reckoned in the second class of foods, as a force generator. For its stimulating effect it must also be included among the force regulators. It acts upon the digestive glands, causing them to pour out their special products more rapidly, and so seems sometimes to assist digestion. But, although alcoholic drinks in very small amounts are found to come, strictly speaking, under the definition of food, in that they may and do develop or regulate force, they possess at the same time properties so peculiar and so dangerous that it is wise to exclude them wholly from our dietary, and use them, if at all, only under the advice of a skillful physician in case of illness. In cer- tain abnormal conditions of the system, when ordinary food cannot be digested, it has sometimes been found that 246 UNCONSCIOUS NERVOUS OPERATIONS an alcoholic drink, not requiring digestion, will supply the necessary energy to sustain life until the diseased organs have time to regain the power to assimilate better food. But to the healthy body there is no need of such a whip and spur, and the stimulus of alcohol upon the secre- tions of the digestive tract, frequently applied, is likely to result in overstimulation of the organs, and consequent weakness, with a long train of evils to follow. As a food, alcohol is of little value compared with other substances. It is more expensive than almost anything else that is ever used as food, and cannot by itself sustain life ; for, while it does generate a certain amount of energy, the body is really feeding upon the stored-up proteids, and the cost of the few spoonfuls of whisky or brandy, or the quart of beer which may perhaps be drunk without immediate bad effects, would buy of wholesome bread and meat enough to produce in the body many times the amount of normal force which the alcohol imparts. 358. Alcohol as a Poison. — The beneficent use of alcoholic drink seems to be wholly confined to its application as a medicine to diseased conditions of the system, and with that sort of use we have here nothing to do. It has been demonstrated that a healthy man may consume drink, in twenty-four hours, which contains from two to two and a half ounces of alcohol without apparent injury, when all circumstances are as favorable as possible for the perfect action of all the bodily organs. But it by no means fol- lows that it would be equally safe for a man in the varying and uncertain conditions of ordinary life to incur the risk of disturbing the nice balance of the physical adjustment upon which vigorous health depends by introducing into his organism an element which may, and more likely than not will, disorder the action of some one or more of the THE DIGESTIVE APPARATUS AND NUTRITION 247 delicate organs. Let him take just a little more than the exact amount which can be at once oxidized in the blood or other tissue, and it is carried on through the system as alcohol, to work its characteristic effects. In the stomach the alcohol may harden the albumin of the food, and so prevent its complete digestion. If strong enough, it may attack the albumin in the cells of the lining of the stomach itself. This is the reason for the well-known fact that alcoholic drinks do less harm if taken after a meal than if taken on an empty stomach. A large quantity of strong drink taken at once seems to paralyze the nerves control- ling the absorbents in the stomach, and often results in sudden death. Passing into the circulation, alcohol, as we have already seen in respect to yeast, acts directly upon the vitality of the living cells, hindering their growth and, when strong enough, wholly destroying their vital power. Any excess beyond the amount which can be oxidized at once interferes with normal cell activity, and works vari- ous physiological evils, as pointed out in different chapters of this book. All these are the actions of a poison. 359. In the stomach alcoholic fluids of all sorts increase very greatly the flow of gastric juice, and it would appear that this stimulation might assist digestion. But since the alcohol is found to disappear wholly from the alimen- tary tract within half an hour, this direct influence upon the secretion of gastric and other digestive juices can be but slight. Moreover, excessive or abnormal stimulation of any organ results ultimately in the weakening of its functional power. In the healthy animal wholesome food supplies all the stimulus needed by the various digestive organs. In the intestines alcohol is rapidly absorbed into the blood. By dilution with the juices of the mouth and the 248 UNCONSCIOUS NERVOUS OPERATIONS stomach its power for direct injury has already been re- duced; but if the work of the stomach has been imper- fectly done because of its presence, more labor remains to the intestines, and that may disorder the whole system. 360. The portal vein carries the alcohol absorbed from the stomach and the intestine directly to the liver. Here its evil effects in developing disease are earliest and often- est apparent. Alcohol in the liver, as in other places, greedily uses up the oxygen needed for the necessary vital operations. In that way it prevents the normal action of the hepatic cells, for not enough oxygen remains for them to do their work perfectly. Now the liver, as the largest organ in the body, has a correspondingly important part to play in the vital processes, and any interference with its functions is extremely serious. It is well known to physi- cians that a drunkard's liver presents a greatly modified appearance, which is seen to a less degree in the liver of a moderate drinker. Alcohol does not build useful tissue, but instead it promotes the abnormal deposit of fat cells, causing what is called " fatty degeneration." This is often seen in the liver of alcohol users and causes enlargement of the organ; the connective tissue, also, is sometimes inflamed and hardened. Because of the imperfect action of the liver, poisons which should have been neutralized are allowed to circulate through the system and undermine its health. So it appears that throughout the digestive tract alco- hol is liable to work harm to the organs more or less serious in proportion to the frequency of its use and its amount. 361 . Other Sources of Danger from Alcoholic Drinks. — Many of the drinks containing alcohol contain also a mixture of nutritious substances, such as unfermented sugar and THE DIGESTIVE APPARATUS AND NUTRITION 249 other foods found in the vegetables used in the manu- facture ; though no one drinks those beverages for the sake of the trace of nourishing matter in them. But there are formed along with the alcohol in fermentation other products, some of which are known to be directly poisonous, while others are at least probably injurious. Fusel oil and various ethers are among these substances. Another source of danger is found in the frequent adul- teration of alcoholic beverages. The great expense attend- ing their manufacture leads to the use of inferior materials, impure sugars, defective fruits, etc., and to the addition of poisonous coloring and flavoring matters. In some cases even so virulent a poison as strychnine has been found. These adulterations still further men- ace the health of the users of such drinks. 362. The Most Dangerous Quality of Alcohol. — Bad as it is to suffer from enfeebled physical conditions, it is yet worse to become weak in mind and unstable in moral character. No one will deny that such weakness and instability are results of the excessive use of alcoholic liquors. But no man ever lived who deliberately determined to make himself a drunkard, when first beginning to taste the exhilaration of the moderate use of such' beverages. Every one thinks he will confine himself to the small quantity which he believes will do him no harm. How is it, then, that in all the great cities of the world certain streets are by night full of the sounds of crazy drunken revelry; that prisons are crowded with criminals made such by involuntary acts when "in liquor"; that myriads of human beings tumble every year into drunkards' graves, dying as the beast dieth, all the beauty and dignity of life wrecked long before, all hope, all possibility of rescue long since abandoned ? 250 UNCONSCIOUS NERVOUS OPERATIONS We class alcohol among the foods because it comes within our definition of food, but in a higher degree than any other substance used for food, it possesses a peculiar power which is not characteristic of food. It is the power of developing a progressive craving, an uncontrollable appetite for itself, which is never satisfied, and which leads the wine bibber to long ever for more and stronger wine, or whisky, or gin, or brandy. It is easy to say, " I will never drink any more alcohol in a day than the two ounces and a half which science has proved can be wholly oxidized in the body, yielding force and conserving to the extent of its own service the physical powers." The man who speaks thus may indeed be able so to regulate his actions, but millions of his fellow-men have not been able to do so. Men do not rightly estimate the full force of the insidious power of alcohol to create an ever-grow- ing appetite which demands ever more alcohol for its satisfaction. Bread and meat and milk and fruit, which build the tissues and supply the forces for vigorous and worthy life, do not create an abnormal appetite for them- selves. Sometimes a man or a woman indulges in the excessive use of tea or coffee, and may possibly experi- ence something of the unhealthy craving for those bev- erages which the drunkard has for his liquor. But harmful as the effect of such indulgence is upon the physical system, it does not so undermine the mental and moral health as does the alcoholic habit. 363. It is at least perfectly safe to avoid wholly the use of alcoholic beverages. One who does so is certain to escape the frightful danger of acquiring that overmastering appe- tite for alcohol, to satisfy which he might become willing to commit murder or arson, or any other crime, and for whose indulgence he may be led to ruin all his hopes of happi- THE DIGESTIVE APPARATUS AND NUTRITION 251 ness for this world and for the world to come, and crush out all joy from the lives of those dear to him. What is the wise course for a being endowed with reason ? DEMONSTRATIONS AND EXPERIMENTS 111. A General Dissection of the Digestive Organs can be performed on the body of a rat, cat, dog, or rabbit. In this dissection other viscera besides the digestive organs should be examined. Just how much of the dissection is to be done by pupils, if any at all, must be left to the judgment of the teacher. Whether the actual class work takes on the nature of a dissection or merely of a demonstration, the attempt should be made to examine, as far as possible, every organ described in the text. To show the villi, cut out a piece of the wall of the small intestine, and after gently washing it examine the inner surface with a hand lens. Teeth of various animals can be obtained to show the arrangement in the jaws and the general structure and materials of a tooth. 112. Minute Structure of Digestive Organs. — Some prepared micro- scopical sections of various parts of the digestive tract will aid the pupil greatly in understanding the structure and properties of the alimentary organs. Very instructive are sections of the wall of the esophagus; of the stomach, showing the three muscular coats; of the small intestine ; and sections of a salivary gland and of the liver. Tissues of any of the domestic animals can be used. Experiments in Digestion. — It should be borne in mind that diges- tion carried on in test tubes is not normal, and that these experi- ments in digestion are merely illustrative. 113. Salivary Digestion. — To a test tube about half full of starch solution * add a little saliva and place the tube where the temperature can be kept at about 37° C. (98° F.). In a few minutes the starch solution becomes clear, and while at first it gave the characteristic reaction with iodine it now no longer turns blue, but if Trommer's test (Ex. 96) be applied, turns yellow, showing presence of sugar. To 1 Rub a gram of laundry starch into a paste with a little cold water. Then add a hundred cubic centimeters of boiling water, and boil for a few minutes. Cool before using. 252 UNCONSCIOUS NERVOUS OPERATIONS be sure that the sugar is a product of digestion, Trommer's test should be applied to the solution before saliva is added, and also to dilute saliva. Prepare two other test tubes in a similar way, but boil the contents of one, and place the other on ice, or in a very cool place. From this conclusions may be drawn regarding the relation of temperature to the activity of the ferment of saliva. The saliva of some persons has little or no digestive effect, hence this experiment will occasionally fail. 114. Gastric Digestion. — Fill three test tubes about half full of artificial gastric juice,1 and three other test tubes with (1) water, (2) water containing a little powdered pepsin, and (3) a ^ per cent muriatic acid solution, respectively. Place in each test tube a few shreds of fibrin. Fibrin is used because it is a solid proteid, and the progress of its digestion can be followed with the eye, without making special tests. Boiled white of egg may be used, but it digests more slowly. Boil one test tube containing artificial gastric juice, place a second on ice, and set away the other test tubes in a warm (37° C.) place. In a short time the fibrin in the tube of gastric juice kept in the warm place is seen to be much swollen, and gradually it disappears iii solution. Compare the test tube with the others. What effect has temperature on gastric digestion? Is the presence of pepsin necessary? \Does pepsin alone (in water) digest the fibrin? 115. Action of Gastric Juice on Milk. — To a test tube about half full of fresh milk, add a little artificial gastric juice that has been neutralized by the addition of dilute carbonate of soda. Keep at a temperature of about 37° C. (98° F.). In a short time the milk curdles. In previous experiments on milk, curdling was produced by acids; here, since the gastric juice was neutralized, it is due to some other cause. To the test tube add a little dilute muriatic acid to acidulate the contents, and keep it in the warm place for several hours. The casein is finally digested in the presence of acid, forming a straw- colored fluid. 116. Action of Rennet on Milk. — To some fresh milk en a test tube add a little commercial extract of rennet, and keep at a tempera- 1 Add a little powdered pepsin (to be obtained at a druggist's) to a ^a per cent solution of muriatic (hydrochloric) acid. THE DIGESTIVE APPARATUS AND NUTRITION 253 ture of about 37° C. The milk curdles in a few minutes. In the previous experiment the milk was curdled by the renuin ferment in the artificial gastric juice. 117. Action of Pancreatic Juice on Starch. — Repeat Ex. 113, using, instead of saliva, artificial pancreatic juice.1 118. Action of Pancreatic Juice on Proteids. — Repeat Ex. 114, osing artificial pancreatic juice instead of gastric juice, and carbonate of soda solution instead of muriatic acid. 119. The Emulsifying Effect of Pancreatic Juice. — Rub together, in a mortar, some olive oil, or cod-liver oil, with pieces of fresh pan- creas. An emulsion results. Shake together in a test tube some olive oil and a little artificial pancreatic juice, as used in preceding experiments. An emulsion occurs as before. Boil some artificial pancreatic juice to destroy the ferment. It still forms an emulsion with oil. In the experiments on fats (Exs. 97-100) it was seen that an alkali, or a soluble proteid, forms an emulsion with fats. Natural pancreatic juice contains both alkali and proteids. Hence, even when boiled, pancreatic juice emulsifies fats. 120. Bile. — Obtain bile at a slaughterhouse. Observe its color. Test with litmus paper. It is neutral or alkaline if fresh. 121. Action of Bile in Fats. — Shake some olive oil in a test tube, with five times its bulk of bile. Make a similar mixture of olive oil and water, and observe in which case the emulsion lasts longer. Shake up bile with olive oil, to which a little oleic a'cid is added. The emulsion lasts longer than before. 122. Action of Bile in Filtration and Absorption. — Into each of two small funnels of exactly the same size, put a filter paper. Moisten one with water and the other with bile. Pour into both equal amounts of almond oil, and after covering to prevent evaporation, set aside twelve to fourteen hours. The oil passes through the filter moistened with bile, but scarcely at all through the other. 1 Add a little powdered pancreatine to a 1 per cent solution of car- bonate of soda. Commercial pancreatine commonly contains both the starch-digesting ferment, qmylopsin, and the proteid-digesting ferment, trypsin. MACY'S pnvs. — 16 CHAPTER XVIII THE DUCTLESS GLANDS 364. The ductless glands are organs whose functions are not yet well understood. Unlike other glands, they do not form a definite secretion poured forth by means of ducts. Some of what we call " true glands " have been shown to send into. the lymph and blood, in addition to the secretion passing through their ducts, material of great importance to the healthy working of the body. This is true of the liver and of the pancreas, though these addi- tional functions are only partly understood. It is probable that the ductless glands have similar offices in the economy of the system. 365. The Spleen (Figs. 94 and 95, pp. 163 and 165). — The largest of these peculiar glands is the spleen, a dark purplish body of variable size and spongy texture, lying on the left side of the abdominal cavity, just below the stom- ach. In the meshes of the tissue of the spleen is a soft sub- stance called spleen pulp. This consists largely of red blood corpuscles and colorless cells, some of which are like the white blood corpuscles. 366. The Blood Supply of the Spleen. — The splenic artery, a branch from the aorta, carries an abundant supply of blood to the gland (Fig. 94), and the smallest branches of the artery open directly into the spleen pulp. This is the only place in the body where the blood comes 264 THE DUCTLESS GLANDS 255 into actual contact with the cells and fibers of a tissue. The veins of the spleen unite to form the splenic vein, which carries the blood into the portal vein and so to the liver. 367. Functions of the Spleen. — Little is yet positively known as to the functions of the spleen. That the organ has some connection with digestion is shown by its enlarge- ment as soon as gastric digestion is completed. It has also some close relation to the liver, and it is understood to be engaged, like the lymphatic glands, in the manufac- ture of white corpuscles. In some animals it forms the colored corpuscles likewise, but we do not know whether that is true of the human spleen. It is thought by some that the spleen is the organ where the red corpuscles which are worn out undergo disintegration, their coloring matter being carried to the liver and there used to form the coloring matter of the bile. 368. The Thyroid Gland is a body whose two lobes lie on the sides of the trachea (Fig. 117). The disease called goiter is an enlargement and alteration of struc- ture of the thyroid, and the effects sometimes ex- tend to the impairment of muscular and nervous activity and to a semi- idiotic condition of mind, resulting in death if the whole gland is affected. The diseased condition Fig. 117. — Thyroid and thymus glands may be relieved or cured of an infant- by grafting a portion of the thyroid gland from an animal under the skin of the afflicted one, or by adding — Thyroid Gland ... Carotid A. -- Internal Jugular V. Trachea ~ ~ Suspensory Ligament -\--Innominate V. Thymus Gland — -Lung 256 UNCONSCIOUS NERVOUS OPERATIONS Svpraren a I Cortex -Medulla Pyramids new thyroid tissue to the food, or even by subcutaneous injection of the juice of a healthy gland. It is concluded that the gland either forms, or helps to form, some sub- stance needful to health, or has some place in the destruc- tion of deleterious substances in the system, but positive knowledge in respect to it is lacking. 369. The Thymus Gland lies in the thorax beneath the sternum (Fig. 117). It is a small organ, weighing only about half an ounce at birth. < .1111 After growing with the body until the second year it gradu- ally shrinks away, and before the age of sixteen usually disappears. Its use is quite unknown. The thymus of calves and lambs forms an article of food called sweet- bread. 370. The Suprarenal Capsules (Fig. 118) are small organs Fig. 11 8. -^Longitudinal section of resting on the upper portion kidney. of the kidneys. They are supplied with an abundance of blood vessels and nerve fibers and nerve cells. It has been found that their removal from animals is invariably and quickly fatal. All that can yet be said as to the function of these bodies is that they appear to form something which is essential to the healthy tone of the muscles. — Ureter CHAPTER XIX THE ORGANS OF EXCRETION 371. We have already learned that a double vital proc- ess is continually carried on by the living cells in the tissues of the body. One side of this vital activity is the taking up, from the blood, of oxygen and the nutrient material which the blood receives from the food in the alimentary canal, and it results in growth and repair. The other side of cell activity is the oxidation, or decom- position by burning in the tissues, of worn-out matter, and its return to the blood to be expelled from the body. This removal of waste matter from the blood is called ex- cretion. The waste material from the tissues leaves the body under three principal forms, — as carbon dioxide, water, and urea. The lungs, as we have seen, not only supply oxygen to the blood, but also give off' daily a large amount of water and carbon dioxide. Two other organs also have the function of excretion. They are the skin and the kidneys. The skin gives off water and certain salts; the kidneys remove urea and other nitrogenous waste, along with a large amount of water. 372. The Skin as an Excretory Organ. — We have studied the skin as an organ of sensation, and have learned some- thing of its structure and its use as a sense organ and as a protector of the more delicate parts. Now we are to study it as a remover of waste. 257 258 UNCONSCIOUS NERVOUS OPERATIONS 373. Structure of the Skin. — It will be remembered that the skin is composed of two layers: the epidermis, com- posed of many layers of cells; and the dermis, or true skin, in which are found the papillae, blood vessels, and nerves, with the end organs for touch and glands of differ- ent sorts (Fig. 49, p. 83). 374. Perspiration. — The excretion of the skin is called perspiration, or sweat, and consists of water, a little dis- solved salt, and some fat. When the perspiration is evapo- rated from the skin as fast as it is secreted, it is called insensible perspiration; but if the quantity is larger, so that it collects upon the surface, we call it sensible perspiration. When the amount of sweat produced is scanty, it is acid in chemical composition; but when the discharge is profuse, it is alkaline. This difference is understood to be due to the mixture of the products of the sebaceous glands with those of the sweat glands. The former are extremely minute glands pouring their secretions into the hair follicles. Their product is acid and fatty, and is constant, or nearly so, in quantity, while that of the sweat glands is alkaline and variable in amount. 375. The Sweat Glands are very abundant over the whole skin. They consist of coiled tubes lying deep in the der- mis, and the duct of each reaches the surface by a cork- screwlike channel. 376. The Nerves controlling the Sweat Glands are of two or three different sets. Those affecting the blood circulation, vasomotor, diminish the secretion of sweat by narrowing the size of the blood vessels, and increase it by dilating them. The special secretory fibers, when stimulated, cause production of perspiration. Still other nerve fibers supply the plain muscle fibers of the glands and regulate the expulsion of the fluid. All these nerve fibers are THE ORGANS OF EXCRETION 259 found in the same nerve trunks. There are subsidi- ary nervous centers for this secretion in different parts of the spinal cord; but the chief center controlling the others is the medulla oblongata, and the nerve fibers for the sweat glands run in the nerve trunks supplying the different parts of the body. For example, the sciatic nerve, supplying the muscles of the leg, carries secretory fibers to the swe'at glands of the leg. 377. Functions of Perspiration. — By means of the sweat glands waste water taken from the blood continually passes into the air. The amount varies greatly, but may be said to average about one and a half pints daily. A very little carbonic acid and solid matter are found in the sweat, along with the fat from the sebaceous glands, and a mere trace of urea. Besides the removal of waste matter, another important function belongs to the skin in connection with perspi- ration. It is well known that as water passes from the liquid to the gaseous state a large amount of heat becomes latent, and this heat is supplied by adjacent bodies. Hence it is clear that by the evaporation of the perspi- ration the surface of the body is cooled, and the sweat thus becomes a regulator of the temperature of the body, the amount of evaporation depending upon the state of the body and of the surrounding air. The amount of sweat secreted is also affected by nerv- ous impulses from the emotional centers in the brain. Fear, for example, sometimes causes profuse sweating. Strong muscular activity, developing heat and stimulating the circulation, increases perspiration. A low temperature in the surrounding air constricts the blood vessels of the skin, and so diminishes the production of sweat and pre- vents loss of heat from the surface. 260 UNCONSCIOUS NERVOUS OPERATIONS 378. The Kidneys (Fig. 118) are bean-shaped organs, lying on each side of the lumbar portion of the backbone. They are dark red in color, about four inches long, two and a half broad, and about one inch in thickness. At the center of the concave, or inner edge, of each kid- ney the arteries enter and the veins leave the organ. The arteries are branches of the aorta ; the veins empty into the inferior vena cava. From the same portion of each kidney passes another tube, the ureter, which con- veys the urine secreted by the kidneys to the sac called the bladder, in the lower part of the abdomen, for storage. The ureters pass obliquely through the wall of the bladder, so that return of the contents of the latter is prevented. 379. Structure of the Kidneys. — By dividing a kidney lengthwise through the middle, two distinct parts may be seen : an outer, granular portion, called the cortex, lying next to the inclosing capsule, and an inner medullary por- tion (Figs. 118 and 120). The latter consists of a number of conical parts, called pyramids, with their bases toward the cortex. In the cortical portion the tiny uriniferou* tubules commence around tufts of blood capillaries {glo- Fig 11 9. -The kidneys and bladder viewed from behind. A aorta, from which the renal arter- ies extend to the kidneys. V inferior vena cava, from which the renal veins extend to the kidneys. u ureters. THE ORGANS OF EXCRETION meruli) and are gathered by a complicated arrangement into larger divisions, and finally empty into the enlarged upper portion of the ureter. From the little bunches of capillaries spreads throughout the cortex a fine network of capillary tubes, which gather into veins and pour the blood into the renal vein. 380. Nervous Supply of the Kidneys. — The kidneys receive nerves from the renal plexus upon each side. This is com- posed of both white and gray nerve fibers and of nerve cells. They come from many sources, but mainly from the sympa- thetic system by way of the Fie 12°- -Diagram of a longi- . J ~/ J . , tudinal section of a kidney, solar plexus. The renal plexus a renal artery. c capillaries. g glomerulus. t uriniferous tubule. v renal vein. has thus indirect connection with the vagus and with other nerves distributed to the inter- nal organs. These nerves seem to have only vasomotor functions. As yet, we are unable to trace the special secretory nerves of the kidneys. 381. Functions of the Kidneys. — The food which we eat, after rendering to the tissues of the body its proper serv- ice, is converted into the waste products water, carbon dioxide, small quantities of salts, and urea (or some sub- stance closely allied). The first two excretions result from decomposition of the carbohydrates and fats, while from the proteids come certain salts and nitrogen. These last are excreted almost wholly by the kidneys, along with a large quantity of water and a very little carbon 262 UNCONSCIOUS NERVOUS OPERATIONS dioxide. To the kidneys, then, belongs the important function of removing from the body the waste product of nitrogenous food in all the living tissues. 382. Urea. — The final result of the changes which nitrogenous foods undergo is urea, and its chief ultimate source is the most abundant tissue — muscular tissue. There is, however, no urea in the muscles. Other steps in its production lie between the muscles and the kidneys. The spleen, lymphatic glands, and other glands have to do with its formation, but the final and most distinctive changes appear to occur in the liver. If the urea is not removed from the system, it destroys life. The quantity daily secreted by the human organism is about five hundred grains, which is normally 2 per cent of the total excre- tion from the kidneys. 383. Relation between the Kidneys and the Skin. — The relation between the skin and the kidneys is such that if one channel of excretion becomes clogged, extra labor appears to be thrown upon the other. If perspiration is checked, there is, along with diminished activity of the sweat glands, constriction of the blood vessels of the sur- face, followed by dilation of those of the viscera, which permits an increased flow of blood in the internal organs, including the kidneys. The secretion of the kidneys, there- fore, becomes more abundant. When by warmth or exer- cise the sweat glands are stimulated, the reverse is the case : the vessels of the skin are dilated, while those of the abdo- men are constricted, and the renal secretion becomes scanty. 384. The Kidneys and the Alimentary Canal. — Still more important seems to be the connection between the amount of the kidney secretion and the water absorbed by the walls of the alimentary canal. When a large quantity of water is drunk, it passes directly into the circulation, and, THE ORGANS OF EXCRETION 263 though the general blood pressure is not raised, it does appear to affect directly the action of the kidneys and to increase the excretion. 385. The Kidneys and the Nervous System. — The state of the central nervous system greatly affects the activity of the kidneys. This may be by the passage of emotional impulses, originating in the brain, along vasodilator fibers to the kidneys. The blood vessels being thus dilated, the activity of the glands would be stimulated. Very large quantities of almost pure water are sometimes thus elimi- nated under emotional excitement. 386. Excretion by the Alimentary Canal. — Portions of the food taken into the stomach are unfit to enter into the structure of the tissues, and are expelled unchanged through the intestines. A small amount of true excretion also takes place there, by which used-up matter is removed. 387. Autointoxication, or Self-poisoning. — Even under normal conditions man's organism is " a receptacle and a laboratory of poisons." They are taken in with the food in the form of those minute bodies which cause putrefac- tion. The process of katabolism (§ 283) is a manufacture of poisons, and many of the secretions, such as the saliva and the bile, are poisonous. A large number of poisonous substances are formed in the intestines, and several forms of deadly poison have been discovered in the urine. All of the poisons made by the tissues, and some of those manufactured in the digestive tube, are poured into the blood, so that even normal blood contains poisons, though an excessive amount of the same poisons threatens health and life. Many diseases are now understood to result from the self-poisoning due to disordered nutrition, along with the opportunities for infection from specific germs which are always present; and reabsorption of excrementitious 264 UNCONSCIOUS NERVOUS OPERATIONS matter once separated from the blood for rejection is a danger to be guarded against. 388. Attention has already been called to some of the ways* by which man is protected from the injurious prod- ucts of his own organism (§§ 335, 336, 338). The gastric juice contains more than enough hydrochloric acid to prevent all fermentation in the stomach ; but in the intestines its action is neutralized by the alkalis of the intestinal juices. In the intestines a variety of poisons are found, and when digestion is disordered the num- ber and quantity may be dangerously increased. Some of these are excreted, while others are absorbed into the blood. The liver is the great defense against the poisons in the blood, many of which are caught by that gland, and either transformed or passed on to be removed from the body by means of the kidneys. By the lungs enough carbon dioxide is removed from the body every day to poison a man to death many times over ; but other injurious matters are found in expired air, especially in the case of persons suffering from defective nutrition. The skin also plays its part in the elimination of poisons, and the peculiar odor of the perspiration in certain abnor- mal conditions is a guide to the physician as to the internal state of the system. All these facts show the immense importance of keep- ing constantly open and in healthy condition the various channels of excretion. 389. Influence of Alcohol upon Excretion. — If the waste substances constantly formed in the body are not promptly removed, they tend to poison the system. When the or- ganism is at a high level of health, the breaking down of tissue by oxidation, which produces waste, goes on rapidly and vigorously. When this is retarded, as we have seen THE ORGANS OF EXCRETION 266 it to be when alcohol is introduced into the circulation and uses up the oxygen which should be applied to the oxidation of food, then the weight may increase, but it is by the retention of poisonous matter which ought to be removed. No other one cause creates so much disease of the kidneys as does the use of alcohol. Imperfect oxida- tion of food develops poisons which the kidneys are overtaxed to remove. This may be caused by eating too much, or by eating unwholesome food, or too much of certain kinds of food, as sugar especially; or it may be caused by alcohol. " Fatty degeneration of the kidneys " is a frequent result of the use of alcoholic drinks. The cells of the tissues become so altered, also, that they fail to act normally by removing only the poisonous sub- stances, and they allow the valuable elements in the blood to be drained off with the waste. This is seen in the serious disease called " Bright's disease " in which the albumin which is necessary to health is excreted by the kidneys. DEMONSTRATIONS 123. Dissection of the Kidney. — Procure a kidney of a sheep or of a pig. As much as possible of the ureter should remain attached. The kidney is seen to be inclosed in a capsule. Remove the latter, and notice the shape of the kidney and the enlarged attachment of the ureter. Split the kidney lengthwise parallel to the broad surface, and observe on the outside of the section a layer, the cortical layer, differing in color from the more internal, medullary, portions. Notice the projections, pyramids, of the medullary portion into the sinus. The latter is a cavity in the concave side of the organ, continuous with the cavity of the ureter. 124. Minute Structure of the Kidney. — For this some prepared sec- tions will be needed: (1) section showing general structure of cortex and medulla, with uriniferous tubules plainly demonstrated ; (2) sec- tion showing blood vessels injected ; (3) a cross section of the ureter. CHAPTER XX THE HEAT OF THE BODY 390. Inanimate bodies tend constantly to assume the temperature of the air, water, or other objects near them. An object which has been heated gives out heat to sur- rounding objects until all are of the same temperature. There is, as we say, a tendency to equilibrium in respect to temperature. 391. Animal Heat. — Warm-blooded animals (birds and mammals) maintain within their bodies, summer and win- ter and indoors and out, with slight variations, the same degree of heat, and are independent of their surroundings in that respect. 392. Temperature of the Body. — In order that the vital processes necessary to human health and comfort may go on under the most favorable circumstances, it is necessary that the body should maintain a temperature of from 98° to 99° F. (from 36.6° to 37.2° C.). If it rises much above or sinks much below this, it is an important indication of abnormal condition in some part of the system. We may be exposed to extreme heat and to severe cold without any marked change in the bodily heat. The skin, it is true, being in contact with external" objects, is usually cooler than other parts. Some of the internal organs have in health a temperature several degrees higher than the gen- eral average, and any special activity of an organ develops 266 THE HEAT OF THE BODY 267 a local excess of heat. A group of muscles, for instance, by their contraction, produce heat. A gland, by the act of secretion, does the same. But the tide of blood, flow- ing swiftly through the system and bathing every part, tends to equalize the heat throughout. 393. The Sources of Animal Heat are of two sorts, — direct and indirect. The great source of heat is the combustion of food, that is, the oxidation which takes place in the living cells in all parts of the body. This is equal to the amount of heat which would be given off by the burning in the open air of the same quantity of food which is con- sumed in the body. The muscles and the glands are the parts in which the greatest amount of oxidation takes place. Some little heat is also received by the body in hot foods and drinks. Heat is produced indirectly by the transformation of other forms of energy. Friction of one part upon another — as of the blood along the walls of the blood vessels — becomes heat. All mechanical work, all nervous activity, and the slight manifestations of electricity within the body liberate heat. 394. Regulation of Temperature is accomplished in two ways; viz. by variation in the loss of heat and by varia- tion in its production. Variation in loss of heat. Everything which leaves the body carries away a portion of its heat. The expired air, the perspiration, the excretions, are sources of loss ; and radiation and conduction of heat from the surface are con- tinually going on, as well as evaporation from the skin and the lungs. The skin is the chief regulator of loss. By clothing the body a portion of the loss by radiation is pre- vented, and we have already seen (§ 376) how the secre- tion of sweat, and hence the cooling of the surface by 268 UNCONSCIOUS NERVOUS OPERATIONS evaporation, is affected by the vasomotor and nervous mechanisms of the skin. Variation in production of heat. The processes of diges- tion are attended by the setting free of heat. The temperature rises after a meal, while a marked condition in starvation is the fall of the bodily temperature. Mus- cular contraction always going on develops heat, and the more active the muscles are, the greater is the amount of heat produced. There is also evidence, from numerous experiments, of direct nervous control over the production of heat. Afferent impulses from the skin or other organs reach the central nervous system and some restricted " heat center " not yet anatomically made out in the central part of the brain. The exact path of these impulses is not yet ascer- tained. The stimulation of the heat center, wherever it may be placed, gives rise to efferent impulses by which activity in the tissues is increased and heat is produced. 395. Clothing. — Though the body is able to endure a large amount of exposure to heat and cold without injury, yet the mechanism for heat production may be overtaxed, as well as the digestive, the muscular, or the nervous system. Any such overtaxing interferes with the other functions of the body. Excessive exposure to cold, or insufficient clothing, forces the body to use an undue amount of energy in manufacturing heat, and other parts of the vital economy suffer ; growth in the young is inter- fered with, and mental and muscular effort become diffi- cult. The clothing worn should therefore be such as will assist in preserving the natural' temperature of the body, and the amount and the material will vary with climate and season, as well as with the age, habits, and health of the wearer. THE HEAT OF THE BODY 269 While clothing is designed to prevent too rapid radia- tion of heat from the surface, it should still permit the evaporation of perspiration, for an accumulation of mois- ture uf)on the skin may expose one to dangerous chills. 396. Fabrics of wool have been found to possess more fully than any other materials the qualities desired for clothing. They are so light and porous as to admit of sufficiently free evaporation and the circulation of air ; and as wool is a bad conductor of heat, it retains the heat of the body, while it holds in its meshes a considerable quantity of air, which is also a nonconductor of heat. In variable climates, such as that of our Northern states, it is wise, at least for those in delicate health or especially sensitive to changes of temperature, to wear wool next the skin at all seasons. The gentle friction of woolen garments against the skin tends to prevent clogging of the pores, to promote even circulation, and in general to keep the surface in a healthy condition. 397. Silk is less valuable than wool in preserving heat and permitting evaporation, though better than cotton oi- lmen. It may be worn next the body when wool causes irritation of the skin. 398. Linen is less useful than other materials for the innermost garments, as it quickly becomes saturated with moisture. Cotton, being more porous, answers the purpose better. In respect to all these materials the weaving of the fabric has much to do with its value for clothing. Closely woven cloth of hard-twisted threads should not be chosen for underwear, but rather that of soft, loosely twisted fibers, loosely woven. 399. Fur is indispensable in the coldest climates, as it retains better than anything else the bodily heat. But it prevents the evaporation of perspiration, and should not MACY'S PHYS. — 17 270 UNCONSCIOUS NERVOUS OPERATIONS be worn in moderate weather ; nor should fur garments be retained indoors — as when sitting in an assembly room. The practice of muffling the throat in wraps of fur should be avoided, as liable to render the larynx unnecessarily sensitive to cold, and to cause the evil it is intended to prevent. 400. Waterproof wraps, which prevent the escape of per- spiration, should be worn only as a protection from rain or snow. 401. Light-colored clothing reflects the rays of heat and absorbs but little warmth, while dark colors absorb heat and reflect little. Hence the common custom of wearing light-colored garments in summer and dark ones in winter is founded in reason. 402. Frequent changes of clothing, especially of that which touches the skin, are needful. The pores of a fabric soon become filled with the poisonous waste mat- ters secreted by the skin. The clothing worn by day should never be worn during the night also. If it is to be resumed in the morning, it should be exposed to free circulation of air during the night. Night clothes and all bedding should be carefully sunned and aired each day. 403. It is important that all clothing and bedding should be thoroughly dry. Damp clothes do not retain the natural heat of the body, but rapidly conduct it away, leaving the surface chilled and the system exposed to attacks of disease. Especially should shoes and stockings which have become wet be removed as soon as possible. Many a serious or fatal illness -has resulted from neglect of this precaution. Damp bedding is especially dangerous to health, as the relaxed condition of the body during sleep renders it an easy prey to every cause of disease. THE HEAT OF THE BODY 271 404. Care should be exercised in changing from the warm clothing of winter to the thinner garments needed in the spring. It is most prudent to make changes first in the outer clothing, retaining the warm inner garments until the mild weather is well established. Often in the Northern states it is necessary to return to heavy winter wraps after a season of high temperature. In the early autumn, too, when cold, damp evenings and nights follow hot, sunny days, judicious attention to clothing will often ward off the intestinal and febrile diseases prevalent in that season. It is frequently wise to change the underclothing with the approach of night, putting on the warm wool which was perhaps intolerable at midday. Many physicians advise as a safeguard the wearing of a broad woolen bandage over the abdomen both by day and by night. 405. The Bodily Heat as affected by Alcohol. — The paralyz- ing effect of the use of alcoholic drinks, upon the muscles in the walls of the minute blood vessels, has been men- tioned in connection with the muscles, the circulation, and respiration. It should be referred to also in connec- tion with the subject of this chapter. Because alcohol is quickly oxidized, and because heat is produced in the process, it was long believed to be of value in maintaining the heat of the body. A different view now prevails as the result of much observation and experiment. Travelers in Arctic regions and others exposed to intense cold agree that those who use no alcohol whatever are far better able to resist the cold than are those who indulge in it. Physiologists show by careful experiments that though the temperature of the body rises during digestion of food, it is lowered for some hours when alcohol is taken. The flush which is felt 272 UNCONSCIOUS NERVOUS OPERATIONS upon the skin after a drink of wine or spirits is due in part to an increase of heat in the body, but also to the paralyzing effect of the alcohol upon the capillary walls, allowing them to dilate, and so permitting more of the warm blood of the interior of the body to reach the sur- face. There it is cooled by radiation, and the general tem- perature is lowered. PART IV THE NERVOUS SYSTEM All our previous study has had to do, directly or in- directly, with the nervous system, with its methods of action, its instruments, its nutrition, its arrangement and functions. Now we are to examine its wonderful and delicate mechanism more in detail, to inquire further into the functions which the various parts fulfill, and into the manner of life which study and experience have shown to be most conducive to the preservation of the most impor- tant part of the human organism in health and efficiency. Even the limited amount of knowledge upon this great subject which is within the reach of young students may be made valuable in enforcing the necessity of hygienic living, and also as a basis for a ready appreciation of that larger revelation respecting the nervous system which science will from time to time unfold. In presenting this fuller description of the nervous system some repetition of the statements given in Chap- ter III is found necessary. S78 Fig. 121.— The nervous system. 274 CHAPTER XXI ANATOMICAL DESCRIPTION 406. Composition of the Nervous System. — The nervous system is made up of nerve centers, nerves, and peripheral end organs. These, though really constituting a single system for the whole of man's organism, are commonly described in two separate groups, or systems: the cerebro- spinal or central and the sympathetic or ganglionic. The nerve centers are the brain and spinal cord (often called the cerebro -spinal axis), and little knots of nervous matter found in different parts of the body, along the course of the nerves, called ganglia. All nerve fibers arise in the nerve centers. 407. Nervous Elements. — We have learned that nerv- ous tissue exists in two forms : gray matter, which is almost wholly composed of nerve cells, and white matter, which is almost wholly composed of nerve fibers. Nerve fibers and nerve cells constitute the nervous elements. 408. The Nerve Cell. — Nerve cells are microscopic, irreg- ular bits of protoplasm like other cells. Each contains a large nucleus, within which is a nucleolus, and usually each cell sends off one or more fine branches, or proc- esses (Fig. 122). Sometimes these are so numerous as to give the cell a stellate appearance. Nerve cells are found only in the central nervous system, in the 275 276 THE NERVOUS SYSTEM ganglia, and in the peripheral terminations of certain nerve fibers. Fig. 122. — Nerve cells from the spinal cord. A nerve cell with all its processes. B body of cell, showing nucleus (N). 409. The Axis Cylinder, or Neuraxon. — One of the proc- esses of the nerve cell — and as a rule only one — becomes what is called the axis cylinder of a nerve fiber. It is a protoplasmic thread continuous with the substance of the cell and usually inclosed within a sheath. Axis cylinder processes give off, usually at right angles, fine side branches which ramify in the adjacent nerve substance (Fig. 123), and the final ending of the axis cylinder itself is in many minute divisions. Many or most of the nerve cells have other processes which do not become axis cylinders, but end in fine twiglike divisions in the gray matter around them. 410. The Nerve Fiber (Fig. 124).— The essential part of every nerve fiber is the central protoplasmic core, which is always the axis cylinder process of a nerve cell. There is hence, after all, only one fundamental form of nervous matter, viz. the protoplasm of the nerve cell. ANATOMICAL DESCRIPTION 277 The central thread of a nerve fiber may have two inclos- ing sheaths. A layer of white, oily matter immediately surrounding the axis cylinder is called the medullary sheath, or sometimes the white sub- stance of Schwann. Out- side of this is a coat of thin, elastic membrane called the primitive sheath, or neurilemma. The lat- ter covers its fiber from end to end, but the med- ullary sheath is broken at frequent intervals, and between the breaks the microscope shows along the course of the axis lit- tle nuclei buried in min- ute masses of protoplasm. Fig. 123. — Nerve In some nerve fibers cells (pyramidal) the medullary sheath is of the cortex of the . cerebrum. wanting, leaving only the Fig. 124. — Por- Ax axis cylinder proo neurilemma inclosing the duSated^erve axis cylinder. These are fiber. called nonmedullated nerve fibers. The medullary sheath is brilliant, shining white in color, and gives to the nerve its characteristic white appearance. The nonmedullated fibers, therefore, are gray. Nerve fibers may be very short or they may be many feet in length. Every nerve fiber originates in a nerve cell, but there are several ways in which it may end, as has already been shown. 411. A Nerve is a bundle of nerve fibers bound together l)_Nodeof Ranvier •Neurilemma (--Nucleus Neuraxon " 'or Axis Cylinder Medullary ' 'Sheath 278 THE NERVOUS SYSTEM by a little connective tissue in which run blood vessels and lymphatics, the whole inclosed in a sheath called the perineurium. The nerves from the central system are composed chiefly of medullated fibers, with which are bound up a few nonmedullated ones, while the nerves of the sympathetic system are made up almost wholly of the gray nonmedullated fibers. 412. A Ganglion is a little group of nerve cells forming a nervous center. As a rule (to which the spinal ganglia form an exception), the nerve fibers running from the cen- tral system to a ganglion are medullated, while those passing from the ganglion toward the periphery are non- medullated and also more numerous. 413. Neuroglia. — The fibers and cells of both the gray and the white nervous matter are supported by a tissue called neuroglia (Fig. 125), which is composed of ex- tremely fine fibers and cells. It differs chemically and in origin from the connective tissues, though like them in function. 414. The Nerve Unit or Neu- ron. — A nerve cell, with its two sorts of processes, con- stitutes a nerve unit^ or neuron (Fig. 126). That is, the whole nervous system is built up of an indefinite but enormous number of these units, supported by neuroglia and connective tissue. A nerve cell sends off one process (rarely more), which is pro- longed into the axis cylinder, or neuraxon* while the numerous other branches, called dendrons or dendrites, almost immediately break up into fine twigs or brushlike Fig. 125. — Neuroglia cells. ANATOMICAL DESCRIPTION 279 £--', Dendrites V™CteH Body --------- Neuraxon ....... Medulla \ ------ Node of Ranvier --- -Neurilemma divisions. The dendrons are thought by some to be con- cerned only in absorbing nutriment for the nerve cell, while other investigators believe them to have also some part to play in the conduction of nerve impulses. Each neuron is anatomically independent of every other. There is no continu- ous path from one nerve cell to another. The fine branches from one cell mingle and interlace with those of another cell, but do not become connected with them to form a continuous channel, any more than do the interlacing branches of two trees standing side by side form com- municating channels for the passage of the sap of one to the other. Nervous influences do indeed pass from one nerve unit to another by some method not yet understood, perhaps by a process similar to that of electrical induction; but the old idea of an uninterrupted channel for the passage of a nervous impulse from center to periphery and from periphery to center is now abandoned. There may be several breaks in the course of transmission, as there often are in the sending of a telegraphic message. The two sorts of branches also preserve their identity from beginning to end of each minutest filament. There is no real network of nerve fibers in the nervous system. 415. The Spinal Cord (Fig. 18, p. 28) is a column of nervous matter from fifteen to eighteen inches in length in the adult, from the foramen magnum, through which Nerve-ends Fig. 126. —Diagram of a neuron or nerve unit. 280 THE NERVOUS SYSTEM Posterior Fissure its fibers pass into the brain, to the fine gray filament which forms its termination in the lumbar vertebrae. A cross section shows it to be composed of white fibrous matter, surrounding a central core of gray cellular matter (Fig. 127). The gray matter presents in section a rough outline of the letter H. In the middle of the cross bar, or isthmus, connecting the two sides of the let- ter, is a minute chan- nel which extends the whole length of the cord and on into the brain. The ends of the letter H which point forward are club-shaped and are known as the anterior horns of the gray mat- ter of the cord, while those pointing back- ward are pointed and are called posterior horns. The last cut through the white matter nearly to the surface of the cord. The white matter of the cord is composed of medullated fibers, running for the most part longitudinally, together with fibrous connective tissue and neuroglia. These fibers are arranged in several different strands, or bundles, and their areas have been carefully mapped out and named. In each half of the cord is an anterior, a lateral, and a pos- terior column, named from their positions and separated from one another by the shallow depressions seen in the Anterior Fissure Fig. 127. — Diagram of a cross section of the spinal cord, showing the divisions of the white and gray matter. The right half shows the threefold division of the white matter commonly described ; the left half shows the physiological divisions as they are at present understood. ANATOMICAL DESCRIPTION 281 surface of the cord. The gray matter consists largely of nerve cells with many branching processes, but there are also many delicate nerve fibers, together with the support- ing neuroglia. The cord is divided into symmetrical halves by fissures before and behind, the posterior fissure being the deeper, and the anterior wider and more distinct. Neither of the fissures cuts quite through the white matter to the gray in the center. 416. Spinal Nerves. — From the grooves nearest the ante- rior fissure spring, by many fine rootlets, the anterior roots Fig. 128. — Diagrammatic cross section of the spinal cord, showing the origin and the chief divisions of a spinal nerve. A anterior column of white matter. D posterior branch of nerve trunk. Ga ganglion on the posterior root of the nerve, containing the cells from which arise most of the fibers of the sensory root (