THE UNIVERSITY OF ILLINOIS LIBRARY ■581.968 42. fig-2. < % y THE GENERA OF SOUTH AFRICAN PLANTS. Digitized by the Internet Archive in 2016 with funding from University of Illinois Urbana-Champaign Alternates https://archive.org/details/generaofsouthafr00harv_0 SOUTH AFEICAN PLANTS. ARRANGED ACCORDING TO THE NATURAL SYSTEM. BY WILLIAM HENRY HARVEY, M.D., F.R.S. & L.S., M.R.I.A., AND PROFESSOR OP BOTANY IN THE UNIVERSITY OF DUBLIN, ETC. Seronts ©tttttxm. EDITED BY J. D. HOOKER, M.D., F.R.S., D.C.L. Oxon., LL.D. Cantab., etc. etc., DIRECTOB OF THE HOYAL GARDENS, KEW. CAPE TOWN: J. C. JUTA. LONDON : LONGMAN, GREEN, READER, AND DYER, 1868. TRINTEP BY J. E. TAYLOR AND CO., LITTLE QVEEN STREET, LINCOLN’S INN YIELDS, LONDON. 581-1 1*& zc & o) PREFACE BY THE EDITOR. It was the wish of my late friend, the author of the ‘ Genera of South African Plants,’ # that I should after his decease edit the materials he had prepared for a Second Edition of that work. These materials embraced descriptions of the genera of the principal families of flowering plants, except the Graminea , which were left in a very incomplete state, and the Restiacea, which were untouched ; and there were also left a few other small families to be worked up, together with the tables of the Natural Orders, and keys, etc. The Gr amine ce have been kindly completed by an intimate friend of Dr. Har- vey and myself, — General Munro, C.B. ; and the Restiacece by my friend Dr. Masters. I have also thought it right to add the genera of Perns (for the descriptions of which I am indebted to Mr. J. G. Baker) and of the allied small Orders. It was not Dr. Harvey’s intention to have (as in the First Edition) included the other Orders of Cryptogams ; of these little was known in 1838, whereas now, in 1868, owing greatly to his own exertions, especially amongst the Mosses and Alga, the number of known South African genera possibly rivals * Published in 1838, at Cape Town, where the author (then in his twenty- eighth year) held the appointment of Colonial Treasurer. 567288 (>* PREFACE. that of the Phsenogams, and to introduce descriptions of them would therefore swell this volume to inconvenient proportions. The Introduction to Botany has been taken from that at- tached to Dus. Harvey and Sokder’s ‘ Flora Capensis,’ with a few very slight additions or modifications. The Keys to the Natural Orders I have constructed to the best of my ability, and so arranged them as to harmonize as much as possible with the sequence of the Orders adopted by Dr. Harvey in the body of the work. Such being the case, I would add that it does not throughout express my own views of the affinities of the Orders as expressed by the Jus- sietjajx system ; it does so, however, to a very great extent, and this for two reasons : firstly, because Dr. Haryey pro- fessedly followed the ‘ Grenera Plantarum ’ of Mr. Bentham and myself, so far as that work had proceeded, and for the remaining Orders (after Rubiacece) he, in the main, followed a manuscript with which I supplied him, giving a rough sketch of Mr. Bentham’s and my own ideas as to the sequence, etc., of the most important alliances or cohorts of Dicoty- ledons, and of the Orders they contain. I have thought it proper to reprint with this edition so much of Dr. Haryey’ s Introduction to the First Edition as explains his motives for undertaking the work, and other matters connected with the history of the Cape Botany ^hat may prove interesting. I must refer the reader to his Pre- faces to the three volumes of the ‘ Flora Capensis ’ for such further information upon the latter subject as brings our knowledge down to the period of his decease (1866). The number of South African flowering plants supposed to be contained in European Herbaria was, in 1838, estimated by Dr. Haryey at 7860, comprised in 937 genera; this was before the exploration of the Natal district. The number is estimated in the present edition at 8777 species, contained in 1209 genera. PREFACE. 7# It only remains to add my regrets that so long a term has elapsed between the death of the gifted and amiable author and the appearance of his work ; this is chiefly due to the fact that it was upwards of a year before the MSS. were placed in my hands, and to the heavy pressure of my own official duties, which has unavoidably retarded its progress through the press. Royal Gardens , Kewy May 27th , 1868. INTRODUCTION. [The following passages from the First Edition of the 4 G-enera of Cape Plants,’ are equally applicable to this. — J. D. H.] I hate been requested by many admirers of flowers to re- commend some introductory work on Botany ; and it would at first seem that I might have taken a much easier method of satisfying their demands than by writing a book for the pur- pose. So many excellent introductions to Botany have been published, that I could not have done better than to place them in a row before inquirers, to choose from. And this I should have done, had an Introduction to Botany been all that was wanting. But I soon found, on cross-questioning, that something very different was required. One lady told me that she knew already what 44 calyx, corolla, stamens, and pistils, and all that” meant ; and another had penetrated the mystery of Monandria, Diandria, etc., and did not want to be told that over again ; what they desired was, a book in which they could discover the names of every plant that struck their fancy in rambling through the fields — in short, a Flora Capensis. Here I found myself completely at fault, for there seemed little use in recommending the Flora of Thttneerg, or the more ancient writings of Burmann, for even could they be procured — which would not be without much difficulty — they would have proved perfectly useless to my lady friends, who, not being blue-stockings, could have derived little instruction from the crabbed Latin in which they are written. Being desirous, however, to afford every assistance in my power to these would-be votaries of my favourite study, the 10* INTRODUCTION. idea of writing a Flora Capensis occurred to me ; but it re- quired only a moment’s consideration to perceive that such a work, to be useful, must not be a compilation from published sources, but must proceed from a very laborious examination of species, consume a long series of years in preparation, and require a much more extensive acquaintance with South African vegetation than I possess, or have at present the means of acquiring. And what were my poor disciples (in posse ) to do while I was thus slowly acquiring the means of meeting their wishes ? There seemed little chance of my being able to do anything to facilitate their approach to the science for many years, until the idea of the present work occurred to me, — a work which, though very far from supply- ing the place of a Flora, would at least make some approaches, towards one, and would show that, if I could not do all I wished to assist their labours, I was perfectly willing to do all that was in my power. And it struck me also, that by publishing now , thus show- ing that I was in earnest in my wish to undertake a Flora Capensis, it might be the means of introducing me to many persons interested in Botany, and living in remote districts of the country, who might, perhaps, be willing to unite with me in amassing materials from which a future Flora should pro- ceed. That there are many such I am willing to believe ; for it is hardly possible that a well-educated person can have con- tinually under his eye so many and such beautiful flowers as are scattered all over the country, without occasionally feeling an admiration of their structure, and a desire to learn some- thing of their affinities and properties. A little sympathy and encouragement are often all that are wanting to make Botanists of these. Intercourse by letter, and interchanges of specimens, foster the incipient taste, till it “ take root downward and bear fruit upward.” Botany is essentially a science of observation, and the more observers in a country to be investigated, and the more widely they are dispersed, the greater will be the chance of com- piling a perfect Flora. Every plant has its peculiar district — its “ range,” as it is technically called ; some species are more INTRODUCTION. 11# widely distributed than others, some are extremely local, and some absolutely confined to a single spot. The importance, therefore, of indiscriminate collections -of every plant of every neighbourhood, must be obvious. By this means we shall secure all the local plants, and be able to define the limits of the range of the more diffused, — a most interesting and im- portant part of Botany. And in few countries do the ranges of species present more curious results than in South Africa. The intervention of a plain, a river, a range of hills, often pro- duces a remarkable change of species ; and a comparison of the plants of any two districts a hundred miles asunder, shows even among common plants, a Flora almost entirely distinct in species. It is well known that the Ericce are, with the ex- ception of a few stragglers, confined to the South-Western districts ; the arborescent Aloes and succulent Euphorbice to the Eastern ; the Stapelice chiefly to the Northern ; the Acan- thaceae , Eubiacece , Bignoniacece , and several other small but remarkable orders to the Eastern ; that Bestiacea , which cover the Western districts, are gradually supplanted by Grasses as we approach the Eastward ; that Leucadendron argenteum is confined to Table Mountain, and the Broteacece generally are much more numerous in the Western than the Eastern dis- tricts. It would be very easy to extend these general remarks on the geographical range of our families and genera, but I rather defer a question of this sort until an extended basis of observations made in all parts of the country shall have been laid, from which a correct sketch of the geography of South African plants may be drawn. And I may take this oppor- tunity of adding, that I am most anxious to obtain information on this very important subject. But I fear that many who might, from their position, ma- terially assist the progress of Botanical Science by making observations on, and collections of, the plants of their neigh- bourhoods, lie under the erroneous supposition that because they have little or no knowledge of Systematic Botany, they are incapable of making collections or observations that can be useful to a botanist. These should recollect that the greatest botanists are at best only students ; there was a time 12* INTRODUCTION. when Linnaeus and Jussieu were as ignorant as themselves, and would have continued so had they waited for intuitive knowledge. Sharp eyes and willing hands are the grand re- quisites for a botanical collector ; and if these be once set in motion, knowledge of affinities and structure will gradually follow as operations extend. Many of the most successful collectors of plants, by whose labours in all countries the science has been so extended and enriched, have been persons ignorant of, or but slightly acquainted with, Botany at the time of their mission. There is one simple way in which all such persons may ren- der important services, and by which they may gradually ac- quire the experience which they would fain possess at start- ing ; namely, by collecting and drying specimens of the plants of their neighbourhood indiscriminately , without favour or affection, from the tall forest tree to the moss or the lichen on its trunk. From such collections only, sent from all parts of the country to a common centre, can a Flora Capensis, in any degree worthy of the subject, be prepared ; and should I be favoured with such, and with the confidence of the col- lectors, it will give me very great pleasure to promote their views in this way, by undertaking the preparation, to the best of my ability, of a Flora of South Africa. Confined as my residence necessarily is to the neighbourhood of Cape Town, I must trust to such help as I may thus receive for all plants found beyond the narrow limits of the Cape District ; and it therefore depends very much on persons attached to Botany and scattered over the country, whether I shall ever be able to undertake a Flora at all. I stand before them as a Can- didate. W. H. Harvey. OxrE Town, July 1, 1838. OUTLINES OF AN INTRODUCTION TO SYSTEMATIC BOTANY. I. Definitions. [ Taken, with slight alterations and additions , and by permission of the author, from Mr. Bentham’s admirable introduction to his ‘ Handbook of the British Flora.’] 1. A Flora of any country consists of descriptions of all the wild or native plants of that country, so drawn up and arranged that the student may easily identify any plant with the corresponding description. 2. The descriptions should he clear , concise , accurate , and characteristic, so that each may he applied to the plant it is intended for, and to no other ; they should he arranged as nearly as possible under natural divisions, so as to facilitate the comparison of each plant with that most nearly allied to it ; and when numerous they should he accompanied hy analytical tables , in which the prominent characters of the species are synoptically presented to the eye, and so contrasted and divided that the student, hy carefully comparing the peculiarities or characters of his plant with the characters laid down in the tables, may he guided with the least delay to the description belonging thereto. 3. Descriptions, to he clear and readily intelligible, should he expressed, if possible, in ordinary, well-established language ; hut, for purposes of accuracy, it is necessary not only to give a more precise, technical meaning to many terms often used vaguely in conversation, hut also to introduce purely technical words and phrases, to express parts of plants, or forms or conditions, which are of little use except to the botanist. Our object in these introductory outlines is to define and explain all technical or techni- cally limited words made use of in the Flora. 4. Mathematical accuracy, however, must not he expected. The forms assumed hy plants and by their parts are all hut infinite. Names cannot be invented for all, nor is strict accuracy in application always attainable. The parts to he described are never precisely regular, nor is the same part precisely of the same form in two individuals of the same species : the botanist’s definitions partake in this uncertainty, and his aim should he, by a few forcible words, to strike out a character applicable to average individuals of the species to be described. 11 INTRODUCTION TO BOTANY. § 1. The Plant in general. 5. Under the common term Plant botanists include every being that has vegetable life ; from the lofty forest tree to the moss upon its stem, the mouldiness on our decaying provisions, or the green scum that floats on stagnant water. 6. Every portion of a plant which has a distinct office or function to perform in the operations of vegetable life is called an Organ. 7. The organs of plants are of two kinds, the elementary and the compound. 8. Elementary organs are those ultimate parts or tissues of which the body of a compound vegetable consists, viz. cellular tissue, woody tissue, and vascular tissue. 9. Compound organs are formed by various combinations of the elemen- tary, and appear under the form of Hoot, Stem, Leaves, Flowers, Fruit. Of these the three first, whose function is to assist in the growth of the plant, are termed Organs of vegetation ; and the two last, whose office is the formation of seed, are the Organs of reproduction. 10. All these compound organs, in some shape or other, exist at some period of the life of most, if not all, flowering plants , technically called pheenogamous or phanerogamous plants ; which all bear fioxvers of more or less complex structure, and are all propagated by seeds containing a germ or embryo plantlet. Flowerless or cryptogamic plants (Ferns, Mosses, Fungi, Lichens, Seaweeds, etc.) have either very imperfect representatives of flowers, or are absolutely flowerless ; and are invariably propagated, not by seeds, but by spores , which do not contain any distinct germ or embryo. 11. The elementary organs will be described afterwards ; we shall con- sider the compound under the following heads : Root, Rootstock, Stem, Leaves, Stipules, Bracts, Inflorescence, Flower, Perianth, Disk, Pistil, Ovule, Receptacle, Fruit, Seed. § 2. The Root, 12. The primary Root, or descending axis, grows downwards from the base of the stem, divides and spreads in the earth or water, and absorbs food for the plant through the extremities of its branches. 13. Roots ordinarily produce neither buds nor leaves ; their branches, called fibres when slender and long, proceed irregularly from any part of their surface ; and they increase in length by constant small additions to their extremities. 14. Though roots proceed usually from the base of the stem or root- stock, they may be formed at the base of any bud, especially if the bud lie along the ground, or elsewhere on the stem, if this is placed in circum- stances favourable for their development. 15. Roots are fibrous, when they consist chiefly of slender fibres ; tuberous , when either the main root or its branches are thickened into short, fleshy, or woody masses called tubers ; tap-roots, when the main root descends perpendicularly, emitting only very few fibrils, as in the Carrot. § 3. The Rootstock or Rhizome. 16. The Stock of a herbaceous perennial, in its complete state, includes a small portion of the summits of the previous year’s roots, as well as of the base of the previous year’s stems. Such stocks will increase yearly so as at length to form dense tufts. They will often preserve through the INTRODUCTION TO BOTANY. Ill winter a few leaves, amongst which are placed the buds, which grow out into stems the following year, whilst the underside of the stock emits new roots from or amongst the remains of the old ones. These 'perennial stocks only differ from the permanent base of an undershrub in the shortness of the perennial part of the stems, and in their usually less woody texture. 17. In some perennials the stock consists merely of a branch, which issues in autumn from the base of the stem, either above-ground or under- ground, and produces one or more buds. This branch, or a portion of it, alone survives the winter. In the following year its buds produce the new stem and roots, whilst the rest of the plant has died away. These annual stocks , called sometimes hybernacula , offsets, or stoles , keep up the communica- tion between the annual stem and root of one year and those of the following year, thus forming altogether a perennial plant. 18. The stock, whether annual or perennial , is often entirely under- ground, or root-like. To this some botanists limit the terms rootstock or rhizome. 19. The term tuber is applied to a short, thick, succulent rootstock, as well as to a root (15) of that shape. The tuber of an orchis, by some called a knob, is an annual tuberous rootstock with one bud at the top. A potato is an annual tuberous rootstock with several buds. 20. A bulb is a subglobose or conical rootstock, formed chiefly of the fleshy bases of the preceding year, or of the undeveloped leaves of the fu- ture year, or of both ; it emits roots from its base, and a stem and foliage from its centre, and frequently forms bulblets or offsets in the axils of its scales. 21. Bulbs are, scaly, when their scales are thick, narrow, and loosely imbricated, as in the white Lily ; tunicated, when the scales are thin, broad, and closely rolled round in concentric layers, as in the Onion. 22. A corm is a fleshy, starchy, and solid rootstock, shaped like a bulb, but not scaly, though often coated with the membranous leaf-bases of a previous season ; its buds are naked, and small in comparison to the fleshy base from which they spring. The Ixias, Gladioluses, etc., afford examples of this form of rootstock. § 4. The Stem. 23. The Stem grows upwards from the root, bears buds which grow out into leafy branches, and finally produces flowers and fruit. 24. Stems are, erect, when they spring perpendicularly from the root or stock ; decumbent, or ascending, when they spread nearly horizontally at the base, and then gradually turn upwards and become erect ; procumbent , when they spread along the ground for the whole or the v greater portion of their length ; prostrate, when they lie still closer to the ground ; creeping, when they emit roots at their joints. This term is also applied to rhizomes or roots, when they spread horizontally. tufted ( ccespitose ), when short, and growing in thick, cushion-like tufts. diffuse, when spreading loosely without being strictly decumbent or procumbent. 25. Weak stems are said to twine when they support themselves by winding spirally round any object ; • and to climb when they support them- selves by their leaves, or by special clasping organs called tendrils, which are usually either imperfectly formed leafstalks or flowerstalks. Twining stems are sometimes called voluble. Sarmentose stems or branches are woody, long, and weak. 8’ b 2 IV INTRODUCTION TO BOTANY. 26. Suckers are young plants formed at the end of creeping, underground rootstocks. 27. Scions, runners , and stolons , or stoles , are names given to young plants formed at the end, or at the nodes (28) of branches or stocks, creeping wholly or partially above-ground, and sometimes to the creeping stocks themselves. 28. A node is a definite point on the stem or on a branch, at which one or more leaves are given off, and an internode is the portion of a stem com- prised between two nodes. The nodes are pervious when the pith passes continuously through them, and closed or impervious when it is interrupted by partitions, as in grasses, etc. 29. Leafbuds are small conical bodies, usually covered with scales, and found in the axils (33) of leaves of the previous season or of earlier growth ; when occurring in other positions, as they sometimes do, they are con- sidered adventitious or irregular. They contain the germs of future branches. 30. Branches (or leaves) are, opposite , when two proceed from the same node at opposite sides of the stem ; whorled or verticillate, when several proceed from the same node, arranged regularly, like the spokes of a wheel, round the stem. geminate , or in pairs, when two proceed from the same node, at the same side of the stem. ternate , in threes, when three spring from one point. fascicled, when several spring from the same or nearly the same apparent point. alternate , when one only proceeds from each node, one on one side, and the next above or below on the opposite side of the stem. decussate , when opposite, but each pair placed at right angles to the one next above or below it ; distichous , when in two ranks ; tristichous , in three, etc. scattered, when placed irregularly round the stem ; but this is often confounded with alternate. secund, when all start from or turn towards one side of the stem, like the teeth of a rake. 31. Branches are, forked , when they divide at the end into two or more equal branches ; dichotomous , when each 2-pronged fork is again divided, and this mode of division several times repeated ; trichotomous, when the forks are 3-pronged, and this repeated ; umbellate, when divided at the apex into several branches, and the central one not larger than the rest. 32. The straw-like stems of grasses and some other endogens are often called culms . § 5. The Leaves. 33. Leaves are expansions which issue laterally from the stem and branches, and usually bear a leafbud (29) in their axil, i.e. in the angle formed by the leaf and the branch. 34. An ordinary leaf consists of an expanded, usually flat blade or lamina , joined to the stem by a footstalk or petiole. The extremity of the lamina next the stem is the base , the opposite extremity the apex, and a line separating the upper and under surfaces, the margin. 35. Leaves are, sessile, when the blade rests on the stem without the intervention of a petiole. • INTRODUCTION TO BOTANY. V Lea vies are, amplexicaul , or stem-clasping, when the sessile base of the blade is not a mere point, but forms more or less of a ring, clasping the stem horizontally. perfoliate, when the base of the blade not only clasps the stem, but closes round it on the opposite side, so that the stem appears to pierce through the membrane of the leaf itself. decurrent , when the edges of the leaf are continued down the stem, so as to form raised lines, or narrow stem-borders called wings. sheathing, when the base of the blade, or of the expanded petiole, forms a vertical sheath round the stem for some distance above the node. 36. Leaves (and flowers) are called radical, when they spring directly from a rhizome or stock, or are inserted so close to the base of a stem as to appear to spring from the root or stock. Leaves are cauline, when they spring from the main portions of the stem ; rameal, when from a branch. 37. Radical leaves are rosulate , when they spread in a circle on the ground ; cauline or rameal leaves axe fascicled or tufted, when the leaves of two or more nodes are brought close together in a pencil-like tuft, by the non-development of the intemodes ; as in Aspalatlius, Asparagus, etc. 38. Leaves are, simple and entire , when the blade consists of a single piece, and the margin is nowhere indented ; simple being used as the opposite to compound, and entire as the opposite to dentate , lobed, or divided. ciliate, when bordered with straight hairs, or hair-like teeth ; cilio- late when the hairs are small. dentate, or toothed, when the margin is slightly notched at regular distances into what have been compared to teeth. Such leaves are serrate when the teeth are pointed like those of a saw ; crenate , when blunt and rounded. The diminutives serrulate, crenulate are used to express minutely serrate or minutely crenate. The hollows between the teeth are respectively called serratures and crenatures. sinuate, when the margin is bluntly indented, with broad, shallow, and irregular hollows between the projections (like the bays between the headlands of a coast) ; wavy, or undulate, when the edges of such a leaf ^re not flat, but bent up and down (like the waves of the sea). The hollows between the projections are called sinuses. lobed or cleft, when more deeply indented or divided, but so that the incisions do not reach the midrib or petiole. The teeth or sections of such leaves are called lobes. divided, when the incisions reach the midrib or petiole, but the parts so divided off, called segments , do not separate from the petiole, even when the leaf falls without tearing. compound, when divided to the midrib or petiole, and the parts so divided off, called leaflets, separate, at least on the fall of the leaf, from the petiole, as the whole leaf does from the stem, without tearing. The petiole of a compound leaf is sometimes called the common petiole (because common to all the leaflets, which often are united to it by petiolules or individual petioles) ; sometimes the rachis, a term also applied to the inflorescence (57). 39. Leaves are more or less distinctly marked by veins, which, starting from the stalk, diverge or branch as the blade widens, and spread over it in various patterns. These veins represent the woody and vascular system (170) of the leaf. The principal ones, when prominent, are often VI INTRODUCTION TO BOTANY. called ribs or nerves ; the smaller branches then retaining the names veins or veinlets. When one principal nerve runs direct from the stalk towards the apex of the leaf, it is called the midrib. When several start from the stalk, diverge slightly without branching* and converge again towards the summit, they are said to be parallel, although not mathematically so. The venation is said to he reticulate or netted, when the veins and veinlets are inosculated together like the meshes of a net; such veins are said to anastomose together. 40. The lobes, segments, or leaflets of leaves are, pinnate, when several succeed each other on each side of the midrib or common petiole (compared to the branches of a feather). A pinnately-lobed leaf is called pinnatijid. A pinnately-lobed or divided leaf is called lyrate, when the terminable lobe or segment is very much larger and broader than the lateral ones ; runcinate , when the lateral lobes are curved backwards towards the base of the leaf ; pectinate , when the lobes resemble the teeth of a comb. palmate or digitate, when several diverge from the same point ; com- pared to the fingers of a hand. A leaf with palmate lobes is called palmatijid. ternate, when three only start from the same point. A leaf with temate lobes is called trifid. A leaf with 3 leaflets, like a trefoil, is trifoliolate ; and when the common petiole is prolonged beyond • the insertion of the lateral leaflets, carrying forward the terminal one a short distance, it is pinnately trifoliolate or unijugate. pedate, when the division is at first temate, but the two outer branches are forked, the outer one of each fork again forked, and so on, and all the branches start from near the base ; vaguely compared to the foot of a bird. A leaf with pedate lobes is called pedatifid. 41. The teeth, lobes, segments, or leaflets may be again and again toothed, lobed, divided, or compounded. Some leaves are 3-4 or many times divided or compounded. In the latter case they are termed decompound. 42. The number of leaves, or of their parts, is expressed adjectively by the following numerals, derived from the Latin : — uni-, bi-, tri-, quadri-, quinque-, sex-, septein-, octo-, novem-, decern-, multi-, 1-, 2-, 3-, 4-, 5-, 6-, 7-, 8-, 9-, 10-, many-, prefixed to a termination, indicating the" part referred to. Thus, unidentate, bidentate , multidentate , mean 1-toothed, 2-toothed, manv- toothed. bifid, trifid, multifid : — 2-lobed, 3-lobed, many-lobed. miifoliolate, bifoliolate : — with one leaflet, with two leaflets, etc. unijugate , bijugate , multijugate , with one, two, or many pairs of pinnce, or leaflets of a pinnate leaf. bipinnate , tripinnate, etc., twice pinnate, thrice pinnate. biternate, triternate, twice temate, thrice temate. 43. Leaves, or their parts or any other flat organs of plants are, linear, when long and narrow, at least 4-5 times as long as broad, with two margins nearly parallel ; subulate, or awl-shaped , when long and narrow, tapering to a sharp point ; pungent , when the point is rigid and piercing. lanceolate, when three or more times as long as broad, broadest just below the middle, and tapering towards the summit ; cuneate , when broadest above the middle, blunt at the apex, and taporing to the base ; compared to an invortod wedge ; fiabelli- form or fan-shaped, is broadly cuneate and rounded at top. INTRODUCTION TO BOTANY. Vll Leaves are, deltoid , equilateral-triangular, or shaped like a Greek A : when ap- plied to the tooth of a calyx , the base of the triangle is sup- posed to rest on the calyx tube. spathulate , when the broad part near the top is short, and the nar- row, tapering part long, compared to a spatula, or flat ladle. oblong (or broadly linear) when 2-4 times as long as broad, with subparallel sides and blunt extremities ; ovate , when about twice as long as broad, broader below the middle, or shaped like the longitudinal section of an egg ; obovate is the same form reversed, the narrow end being at base. orbicular, or rotund, when approaching to a circular form. oval, or elliptical, when broadest in the middle, with curved sides, like an ellipse, or oval. transversely oblong , or oblate , when conspicuously broader than long. rhomboidal, or rhomboid, when shaped like the mathematical figure called a rhombus. falcate , shaped like a scythe, curved with nearly parallel sides and a sharp point. 44. Intermediate forms are expressed by combining two terms. Thus, a linear -lanceolate leaf is long and narrow, yet rather broader below the middle and tapering to a point ; ovato-lanceolate is broad towards the base, yet tapering towards the apex, etc. 45. The apex or summit of a leaf is, acute , or pointed, when it is sharp, or forms an acute angle. obtuse, or blunt, when it forms an acute angle or oftener is rounded. acuminate, when suddenly narrowed near the top and then pro- longed into an acumen , or projecting point, which may be acute or obtuse, linear or tapering ; cuspidate is either synonymous with acuminate, or else used to express a more exaggerated degree of acumination, with a more sudden, sharper, and more rigid point. truncate, when the end is cut off nearly square. retuse, when very obtuse or truncate, and slightly indented in the middle; emarginate or notched, when more decidedly indented at the end of the midrib. mucronate, when the midrib is produced beyond the apex in the form of a small point ; mucronulate when it projects very little. aristate, when the point is long and fine, like a bristle. 46. The base of the leaf is liable to the same variations of form as the apex, but the terms commonly used are tapering or narrowed for acute and acuminate ; rounded for obtuse ; and cordate for emarginate. But the term cordiform, cordate , or heart-shaped leaf, is restricted to an ovate and acute leaf, cordate at base, with rounded auricles ; and obcordate to an obovate, deeply emarginate or subbilobed leaf or leaflet, with rounded lobes. The word auricle is more strictly applied to the prolonged base of sessile and stem-clasping leaves. 47. If the auricles (46) are pointed, the form is said to be sagittate when the points are directed downwards, as in the head of an arrow ; hastate , when the points diverge horizontally, compared to a halbert. 48. A reniform leaf is broader than long, very obtuse at apex, slightly but broadly cordate at base, with rounded auricles, compared to a kidney. 49. In a peltate leaf, the stalk, instead of proceeding from the lower edge of the blade, is attached to the under surface, often near the lower edge, but sometimes in the very centre of the blade ; the nerves radiate from the point of attachment of the petiole, or proper base of the leaf It Vlll INTRODUCTION TO BOTANY. is structurally a cordato or reniform leaf whose auricles are confluent ; just as a perfoliate leaf (35) is the amplexicaul, in a similar condition. 50. In their consistence, leaves or other flat organs are, fleshy , when thick and soft ; succulent is generally applied in the same sense, hut implies the presence of more juice ; coriaceous , or leathery, when firm and dry, or very tough ; membranous , when thin and not stiff ; scarious , or scariose , when very thin, more or less transparent, and not green, yet rather stiff. 51. Solid leaves (or stems, fruits, seeds, or other parts of plants not flattened) are, acerose , or acicular , very slender, like needles ; setaceous , bristleform ; capillary, when very slender, like hairs ; ovoid , when egg-shaped, with the broad end downwards ; obovoid, if the broad end be upwards. globular , or spherical , round like a ball ; conical, cone-shaped, tapering upwards ; and obconical, tapering down- wards, if in both cases a cross-section shows a circle. pyramidal, when tapering upwards, obpyramidal , when tapering downwards, if in both cases a cross-section shows a triangle or polygon. fusiform, when broad in the middle and tapering to each end like a spindle, and not angular. cylindrical, when not perceptibly tapering and not angular. terete, cylindrical, with the cross-Section circular ; moniliform , and torulose, when terete and constricted at intervals. trigonous , rather bluntly 3-angled ; triquetrous, sharply 3-angled. flattened, or depressed, when vertically compressed. compressed, when laterally flattened ; and obcompressed (a bad term) sometimes used in the sense of dorsally compressed. meniscoid, shaped like a watch-glass. patelliform, saucer-shaped. 52. The mode in which unexpanded leaves are disposed in the leaf bud is called their vernation, or prcefoliation : it varies considerably, but is rarely noticed in descriptive botany. § 6. Stipules. 53. Stipules are leaf-like or scale-like appendages at the base of the leafstalk or on the node of the stem. They are often absent, when the leaf is exstipulate ; when present they are generally two, one at each side of the petiole, and they sometimes appear to protect the young leaf before it is developed. They vary extremely in size and appearance ; and are either free, i. e. separate from the petiole, or adnate, i. e. laterally attached wholly or in part to the petiole. They often afford excellent characters in distinguishing plants from each other, and ought always to be closely observed. 54. Stipellce, or secondary stipules, ar esimilar organs, sometimes found on compound leaves at the points where the leaflets are inserted. § 7. Bracts. 55. A Bractea or bract, is either the leaf from the axil of which a flower is developed, when this differs in appearance from an ordinary leaf ; or else it is any reduced leaf situated on the branches of the inflorescence (57) below the calyx. 56. When flower-stalks are branched, and have bracts at their first as well as at their second and subsequent ramifications, the formor aro called general, the latter partial bracts, or bracteoles. The terms general and INTRODUCTION TO BOTANY. IX partial are also applied to involucres (64) when similarly situated. The word bracteole is sometimes given to the uppermost bracts, when much smaller or very different from the lower ones. § 8. Inflorescence. 57. The Inflorescence of a plant is the arrangement of the flowering branches, and of the flowers upon them. An inflorescence is a flowering branch, or the flowering summit of a plant above the last stem-leaves, with its branches, bracts, and flowers. 58. A flower or an inflorescence is terminal when at the summit of a stem or leafy branch ; axillary, when in the axil of a lateral leaf. 59. A peduncle, or flowerstalk, is the stalk either of a solitary flower or of an inflorescence ; in the latter case it may he either simple or branched. A pedicel is the ultimate branchlet of an inflorescence, supporting a single flower. 60. A scape is a peduncle that proceeds from the rootstock, or from so near the base of the stem as to appear radical, provided always that it bears no leaves at all, or that the leaves are reduced to mere scales or bracts. 61. The inflorescence is centrifugal , when the terminal flower opens first, and those on the lateral branches are successively developed ; centri- petal, when the lowest flowers open first and the main stem continues to lengthen, developing fresh flowers. Both these kinds of inflorescence may be combined on the same plant ; the main branches may be centripetal, and the flowers on the lateral branches centrifugal, or vice versa. 62. An Inflorescence is, a Spike, or spicate , when the flowers are sessile along a single un- branched axis, called the rachis. Catkin is the name given to the spicate inflorescence of several trees whose flowers are reduced to scaly bracts or are very imperfect ; as in the Oak, Willow, etc. Spadix is a fleshy spike round which is rolled a single large bract, or spathe, as in the Arum plant [Richardia JEthiopica). The inflorescence of a Palm is usually a branching spadix. a Raceme, or racemose, when the flowers are borne on pedicels along a single unbranched axis, also often called the rachis. a Panicle , or paniculate , when the axis is divided into branches, each bearing two or more flowers. a Head, or capitate , when several sessile or subsessile flowers are collected into a compact, head-like cluster. The short, flat, or conical axis on which the flowers of a head, or capitulmn , are seated, is called the receptacle — a term also given to the torus or thalamium of a single flower. an Umbel, or umbellate , when several branches or pedicels spring from the same (apparent) point, i. e. from an axis reduced to a point. (It is essentially the same as a raceme with the axis sup- pressed ; or as a head, with long-stalked flowers.) An umbel is said to be simple, when each of its branches or rays supports a single flower ; compound, when each ray supports a partial umbel, or umbellule. a Corymb, or corymbose, when the branches or pedicels starting from several points on a short, but not suppressed axis, all attain nearly the same level. It is a flat-topped or fgstigiate panicle or short raceme. a Cyme , or cymose inflorescence, is a centrifugal panicle, and is fre- quently corymbose. The terminal flower opens first. The X INTRODUCTION TO BOTANY. lateral branches, successively developed, are usually forked ( dichotomous or trichotomous) . Sometimes after the first forking the branches are no longer divided, but produce a succession of pedicels on the upper side, forming apparently unilateral racemes ; but they differ from true racemes by the pedicels springing, not from the axil of the bract, but from a point opposite its insertion or above or below it. This variety, called a scorpioid cyme , is found in Drosera, the Boraginece, and many other plants ; when young the branches are frequently rolled back at the top like the tail of a scorpion, whence the name. 63. Bracts are generally placed singly under each branch of the in- florescence, and under each pedicel ; bracteoles are usually two, one on each side, on the pedicel, or close under the flower, or on the calyx itself ; but bracts are also frequently scattered along the floral branches without axillary pedicels. 64. When several bracts are collected in a whorl, or are so close together as to appear whorled, or are closely imbricated round the base of a head of flowers or an umbel, they are collectively called an Involucre. The bracts composing an involucre are variously termed, according to their appearance, leaves , leaflets , bracts , or scales, when placed close beneath the calyx, they form an epicalyx. Palece, or chaff, of the receptacle are the inner scale-like bracts of Composites, grasses, and some other plants, when of a thin, yet rigid substance, usually narrow, and pale or trans- lucent. Glumes are the bracts of sedges and grasses. § 9. The Flower. 65. A flower is a terminal bud, enclosing the organs of reproduction by seed. An unopened flower is called a flower-bud, or alabastrum ; and the period between the opening of a flower and the commencement of withering is called its anthesis. 66. The parts of a flower or floral organs are — 1st, the perianth , con- sisting either wholly of calyx , if in a single whorl ; or of calyx and corolla, if in a double whorl ; 2nd, the stamens, or fertilizing organs ; 3rd, the pistil , which contains the ovules, or germs of the future seed. 67. A complete flower is one in which the calyx, corolla, stamens, and pistil are all present ; a perfect flower is one in which all these organs, or such of them as are present, are capable of performing their several func- tions. An incomplete flower is one in which some of the floral organs are wanting ; and an imperfect flower one in which some organs are so altered as to be incapable of performing their proper functions. Imperfect organs are said to be suppressed, abortive , or rudimentary , if very much reduced in size and almost obliterated. 68. A flower is, dichlamydeous , when the perianth is in two distinct whorls. monochlamydeous, when the perianth is in one whorl, or when calyx and corolla are so consolidated as to appear in a single piece. asepalous, when there is no calyx. apetalous, when there is no corolla. naked, when there is no perianth. hermaphrodite , or bisexual, when both stamens and pistil arc prosent and perfect. male , or staminate , when stamens, but no pistil, or only an imperfect one, are present. female , or pistillate , when there is a perfect pistil, but no stamens, or only imperfect ones, present. neuter, when both stamens and pistils are imperfect or absent. INTRODUCTION TO BOTANY. XI A flower is, barren , or sterile , when it produces no seed. fertile , when it does produce seed. 69. The flowers of a plant or species are said collectively to he, unisexual , or diclinous , when the flowers are all either male or female. monoecious , when the male and female flowers are distinct, hut on the same plant. dioecious , when the male and female flowers are on distinct plants. polygamous , when male, female, and hermaphrodite flowers are variously mixed on the same plant. 70. A head of flowers, or capitulum, is heterogamous, when male, female, hermaphrodite, and neuter flowers, or any two of these, are included in it; homogamous , when all the flowers are of one kind and structure. A spike or head is androgynous , when male and female flowers are mixed in it. (See Composite, Aroidece, Cyperacece, etc.) 71. As the scales of buds are leaves undeveloped or reduced in size, shape, and consistence, and bracts are leaves likewise reduced in size and occa- sionally altered in colour ; so the parts of the flower are considered as leaves still further altered in shape, colour, and arrangement round the axis, and often more or less combined with each other. 72. To understand the arrangement of the floral parts let us take a complete flower, in which all the parts axe free from each other; definite in number, i. e. always the same in the same species ; and symmetrical or isomerous, i. e. when each whorl consists of the same number of parts. The flower of Flax ( Linum ), of Crassula, or of Oxalis, answers to this description : the two first consist of 4, the last-named of 5 whorls of altered leaves, placed immediately one within the other. 73. The Galyx forms the outer whorl. Its parts are called sepals. 74. The Corolla forms the next whorl. Its parts, called petals, usually alternate with the sepals, i. e. the centre of each petal is immediately over the interval between two sepals. 75. The Stamens form one or two or more whorls within the petals. If two, those of the outer whorl (the outer stamens) usually alternate with the petals, and are consequently opposite to the sepals ; those of the inner whorl (the inner stamens) alternate with the outer ones, and are conse- quently opposite the petals. If there is but one whorl of stamens, they most frequently alternate with the petals ; sometimes (as in Rhamneoe and Frimulacece) they are opposite the petals and alternate with the sepals. 76. The Pistil forms the innermost whorl. Its parts, called carpels, usually (when definite and isomerous) alternate with the inner row of stamens. 77. In an axillary or lateral flower (58) the upper parts of each whorl (sepals, petals, stamens, or pistil), are those that are next the main axis of the stem or branch ; the lower those that are furthest from it ; the inter- mediate are said to be lateral. 78. The number of parts in each whorl of a flower is expressed adjec- tively by the following numerals, derived from the Greek : mono-, di-, tri-, tetra-, penta-, hexa-, hepta-, octo-, ennea-, deca-, etc., poly-, 1-, 2-, 3-, 4-, 5-, 6-, 7-, 8-, 9-, 10-, many-, prefixed to a termination indicating the whorl referred to. 79. Thus, a Flower is, disepalous, trisepalous, etc., as there are 2, 3, etc., sepals. dipetalous, tripetalous , etc., as there are 2, 3, etc., petals. diandrous , triandrous , etc., as there are 2, 3, etc., stamens. digynous , trigynous, etc., as there are 2, 3, etc., separate carpels. Xll TNTKODUCTION TO BOTANY. Thus, a flower is, dimerous , trimerous , etc., if symmetrical, according as there are 2, 3, etc., parts in each whorl. 80. Flowers are unsymmetrical , or anisomerous , strictly speaking, when any one of the whorls has a different number of parts from the others ; hut when the carpels alone are reduced in number, the flower is still frequently called symmetrical, or isomerous, if the calyx, corolla, and staminal whorls have all the same number of parts. 81. Flowers are irregular when the parts of any one of the whorls are unequal in size, dissimilar in shape, or do not spread regularly round the axis at equal distances. In descriptions, it is more especially irregularity of the corolla that is referred to ; a slight inequality in other parts does not prevent the flower being classed as regular, if corolla or perianth be regular. § 10. The Perianth , and Calyx or Corolla. 82. The Calyx or outer whorl of the perianth is usually green, smaller than the corolla, and of coarser texture ; sometimes very minute, rudi- mentary, or obsolete altogether ; sometimes imperfectly whorled, or not whorled at all, or composed (as in Cactus ) of a large number of sepals , of which the outer ones pass gradually into bracts, and the inner ones into petals. 83. The Corolla or inner whorl of the perianth is usually coloured, larger than the calyx, and of a more delicate texture, and in popular lan- guage is often called the flower. Its petals, except in double flowers , are rarely indefinite in number, and the whorl more rarely broken than in the calyx. Sometimes the petals are very small, rudimentary, reduced to scales (as in Thymelece ), or absent altogether. 84. In very many cases the so-called simple Perianth is one in which the sepals and petals are nearly similar in form and texture, and present apparently a single whorl. The real nature of such a perianth may be detected by examining an unopened flower-bud, when one half of the parts will be found placed outside of the others (as in Anthericum , Ornitho- galum , Rumex , etc.), indicating an arrangement in two whorls, or calyx and corolla. Hence different authors may describe the same flower diffe- rently, either as having a single or a double perianth. 85. In the following terms the prefixes expressive of the modifications of the corolla and its petals are equally applicable to the calyx and its sepals, or to the perianth and its segments. 86. The Corolla is, monopet alous (sometimes called gamopetalous ), when the petals are united or soldered together, either entirely or at the base only, into a cup, tube, or ring. polypetalous (or dialipetalous ), when they are all separate or free from the very base. 87. When the petals are partially united in a monopetalous corolla, the lower consolidated portion of the corolla is called the tube , whatever be its shape, and the free upper portions of the petals are called the teeth, lobes , or segments , according as they are short or long in proportion to the whole length of the corolla. When the calyx or corolla enlarges after flowering it is called accrescent ; when it falls early, deciduous or caducous. 88. The flat expanded portion of a petal, corresponding to the blade of the leaf, is called its limb or lamina ; and the stalk, corresponding to the petiole, its claw. When there is no claw, the petals are sessile. 89. The cestivation of a corolla is the arrangement of the petals, or their free portions, in an unexpanded bud. It is valvate , when the edges of the petals touch, but do not overlap ; imbricate , when the edges overlap each other, at least near the top ; twisted , contorted , or convolute, when each petal INTRODUCTION TO BOTANY. Xlll obliquely overlaps the adjoining one on one side, and is overlapped by the adjoining one on the other side. In valvate aestivation, if the edges are much indexed, the aestivation is said to be induplicate ; involute , if the margins are inrolled ; and reduplicate , if the margins project outwards into salient angles ; plicate , when folded together in plaits ; crumpled , when puckered irregularly, as in the petals of a poppy. 90. In general shape the Corolla is, tubular , when the whole or the greater part forms a tube or a cylinder. campanulate , when approaching in some measure the shape of a cup or bell. urceolate, when the tube is swollen, but contracted at the top, and slightly expanded again into a narrow rim, as in many Heaths. rotate or stellate , when the petals or lobes are spread out horizontally from the base, or nearly so, like the rays of a wheel or star. hypocrateriform or salver-shaped, when the lower part is cylindrical, and the upper portion expanded horizontally. In this case the term tube is restricted to the cylindrical part, and the horizontal portion is called the limb, whether it be divided to the base or not. infundibuliform ox funnel-shaped, when the tube is cylindrical below, but gradually enlarged upwards into a subcampanulate limb, of which the lobes either stand erect or spread horizontally. The upper orifice of the tube of a monopetalous corolla is often called its mouth or throat. Irregular corollas have received various names ; some of the most im- portant are bilabiate , or two-lipped corolla, when in a four or five-lobed corolla (or calyx) the two or three upper lobes stand obviously apart, like an upper lip, from the two or three lower lobes, that form an under lip. personate, when two-lipped, and the orifice of the tube closed by a projection from the base of the upper or lower lip, called a palate ; as in Snapdragon, Nemesia , etc. ringent, when strongly two-lipped and the orifice of the tube very open. spurred, when the tube, or the lower part of the petal has a conical, hollow projection, compared to the spur of a cock; saccate, when the spur is short and round, like a little bag ; gibbous , when swollen or enlarged at one side. resupinate , or reversed, when the under lip is turned up, or appears so. The above terms are mostly applied to monopetalous corollas. Terms applied to certain forms of corolla distinctive of certain Natural Orders will be explained under the respective Orders. §11. The Stamens. 91. The stamens or fertilizing organs of a flower, though in a theoretical point of view regarded as metamorphosed leaves, are yet, except in a few cases of petal-like stamens, very different in shape and aspect from leaves, sepals, or petals. 92. Usually a stamen consists of a stalk or filament , bearing at the summit an anther, divided into two pouches or cells. These anther-cells are filled with pollen, commonly existing as minute grains, forming a yellow dust, which on the expansion of the flower is scattered abroad from an opening in each cell, called a slit or pore. The part of the anther XIV INTRODUCTION TO BOTANY. which connects the two cells is called the connective ; it is sometimes a mere line, but often variously expanded, causing the cells to separate more or less. 93. The filament is often wanting, and the anther sessile, yet Still the stamen is perfect ; but if the anther, which is the essential part, be want- ing, or does not contain pollen, the stamen is imperfect, and is then said to be barren , abortive , or rudimentary (67), according to the degree to which the imperfection is carried. Imperfect stamens are often called staminodia. 94. In unsymmetrical flowers the stamens of each whorl are sometimes reduced in number below that of the petals, even to a single one, and in several Natural Orders or genera they are multiplied indefinitely. The terms monandrous and polyandrous are restricted to flowers which have really but one stamen, or an indefinite number respectively. When several stamens are united into one piece, the flower is said to be synandrous. 95. Stamens are, monadelphous , when united by their filaments into one company or cluster. This cluster either forms a ring or tube round the pistil, or, if the pistil be wanting, occupies the centre of the flower. diadelphous , when so united into two clusters. In many Leguminosce 9 stamens are united by their filaments into a tube, slit on the upper side, and a tenth, placed in the slit, is free. In Fumariacece there are two equal groups, each consisting of 3 (or rather J, 1, ■J) stamens. triadelphous , pentadelphous , polyadelphous , when so united into 3, 5, or several clusters. syngenesious, when united by their anthers in a ring round the pistil, the filaments usually remaining free ; as in the Composites. didynamous, when (as in a bilabiate flower) there are 4 stamens in two pairs, those of one pair longer than those of the other. tetradynamous , when (in Crucifer ce) there are six, four of them longer than the others. 96. An Anther is, adnate , when continuous with the filament, the anther-cells appear- ing to lie their whole length along the upper part of the filament. versatile , when attached near their middle to the extreme point of the filament, so as to swing loosely. innate , when firmly attached by their base to the apex of the fila- ment. This is a modification of adnate. 97. Anther-cells may be parallel ; or diverging at a less or a greater angle ; or divaricate , when placed end to end, so as to form a nearly straight line. The end of each anther-cell placed nearest to the other cell is gene- rally called its apex or summit , and the other end its base ; but by some authors the sense of these terms is reversed. 98. Anthers have often on their connective, or on their cells, appen- dages termed bristles ( setee ), spurs , crests , points , glands , etc. : according to their appearance. (See Diosmeez , Ericacece , etc.) 99. Anthers have occasionally only one cell ; this may take place either by the disappearance of the septum between two very closely-placed cells, which thus become confluent ; or by the abortion or total deficiency of one cell, when the anther is said to be dimidiate , or halved. 100. Anthers at maturity will open or dehisce , to let out the pollen, either by valves , pores , or slits. The dehiscence is said to be introrse when INTRODUCTION TO BOTANY. XV the slit or opening faces the pistil ; extrorse , when the opening is towards the circumference of the flower. 101. Very peculiar structures of the anther and pollen will be described under the Orders Asclepiadece and Orchidece. § 12. The Pistil. 102. The Pistil or female system always occupies the centre of the flower and terminates the growing axis. It consists of one or more carpels , containing the germs of one or more seeds. The pistil is usually sessile ; if stalked, its stalk is called a podocarp (but this must not be confounded with the gynobasis or gynophore) (127). 103. A complete pistil consists of three parts : — 1, the ovary or enlarged base, which includes a cavity or cell {loculus), containing one or more ovules (117), which are the earliest condition of the future seeds. 2, the style, a prolongation of the carpel usually proceeding from the summit, sometimes from the side of the ovary, and sup- porting 3, the stigma . This is various in appearance, sometimes a mere point to the style, sometimes a flattish cushion, sometimes a narrow line, sometimes a broad lamina ; but it always consists of loosely cellular substance, destitute of epidermis, and covered with minute protuberances, called papillce. It is through the stigmatic tissue that the fertilizing influence of the pollen is conveyed to the ovules. 104. The style is often wanting, and the stigma is then sessile, but in the perfect pistil there is always at least one ovule in the ovary, and some portion of stigmatic tissue. Without these the pistil is imperfect, and said to be barren, abortive, or rudimentary according to the degree of imper- fection. 105. The ovary being the essential part of a pistil, most of the terms relating to the number, arrangement, etc., of the carpels apply specially to the ovary. In general, the term ovary is used to designate all the carpels of a flower, especially if they are at all united. 106. The number of carpels or ovaries in a flower is frequently reduced below that of the parts of the other floral whorls, even in flowers otherwise symmetrical. In a comparatively small number of genera the carpels are more numerous than the petals, or indefinite; in these cases they are either arranged in a single whorl, or form a head or spike in the centre of the flower, as in the Buttercup and Anemone. 107. The terms monogynous , digynous, etc., poly gy nous (1, 2, or many ovaries) are vaguely used, applying sometimes to the whole pistil, some- times to the carpels alone, and sometimes to the styles or stigmas. When a more precise nomenclature is used, the flower is monocarpellary, when the pistil consists of a single, simple carpel ; hi-, tri -, etc., or polycarpellary , when the pistil consists of 2, 3, or many carpels, whether separate or united. 108. A pistil is syncarpous when the carpels are united into one com- pound ovary ; apocarpous, when the carpels are free or separate. 109. A compound ovary is, unilocular, or one-celled , when there are no partitions between the ovules, or when the partitions do not meet, so as to divide the cavity into several chambers. plurilocular, or several-celled, when completely divided into two or more cells by partitions called dissepiments {septa). These dissepiments are usually vertical, radiating from the centre or axis of the ovary to its circumference. XVI INTRODUCTION TO BOTANY. A compound ovary is, hi-, tri-} quadri-y etc., or multilocular, according to the number of the cells, or loculi, 2, 3, 4, or many. 110. In general the number of cells or dissepiments, complete or partial, or of rows of ovules, corresponds with that of the carpels, of which the ovary is composed. But sometimes each carpel is divided completely or partially into two cells, or has two rows of ovules, so that the number of carpels appears double what it really is. Sometimes, again, the carpels are so completely combined as to form a single cell, with a single ovule, although it really consists of several carpels ; but in these cases the ovary is generally described as it appears, as well as such as it is theoretically supposed to be. 111. In apocarpous ovaries the styles are usually free, each bearing its own stigma ; very rarely (as in Asclepiadece ) the larger portion of the styles or the stigmas alone are united, while the carpels remain separate. 112. Syncarpous flowers are said to have, several styles, when the styles are free from the base. one style with several branches, when the styles are connected at the base, but separate below the point where the stigmas or stigmatic surfaces commence. one simple style with several stigmas, when united up to the point where the stigmas or stigmatic surfaces commence, and then separating. one simple style , with branched, lobed, toothed, notched, or entire stigma (as the case may be) when the stigmas also are more or less united or subdivided. 113. In general the number of styles, or of branches of the style or stigmas, is the same as that of the carpels, but sometimes that number is doubled, especially in the stigmas, and sometimes the stigmas are much subdivided or penicillate , that is, divided into a tuft of hair-like branches, as in some species of Drosera. 114. An entire stigma is said to be punctiform when it appears like the mere point of the style; capitate, when globular, like the head of a pin. 115. The placenta is the part of the inside of the ovary to which ovules are attached, sometimes a mere point or line on the inner surface, often more or less thickened or raised. Placentation therefore is the indication of the part of the ovary to which ovules are attached. 116. The placentas are, apile, when the ovules are attached to the axis or centre, that is, in plurilocular ovaries, when they are attached to the inner angle of the cell ; in unilocular simple ovaries (which have almost always an excentrical style or stigma) when the ovules are attached to that side of the ovary nearest to the style ; in uni- locular compound ovaries, when the ovules are attached to a central axis, cushion, or column, rising up from the base of the cavity, and either free at the top (free central placenta), or attached also to the summit of the cavity. parietal, when the ovules are attached to the inner surface or walls of the cavity of a one-celled compound ovary. Parietal pla- centae are usually slightly thickened or raised lines; some- times broad surfaces nearly covering the inner surface of the cavity ; sometimes projecting far into the cavity and constitu- ting partial dissepiments, and even meeting in the centre, but without cohering there. In the latter case the distinction between the one-celled and several-celled ovary sometimes almost disappears. INTRODUCTION TO BOTANY. XVI 1 § 13. The Ovule. 117. The ovule is a minute body borne by the placenta (115) and destined, after fertilization, to become the seed. At first it is merely a cellular excrescence, but as it enlarges it acquires a definite form and structure, and when fully grown consists of a central mass or nucleus , en- closed in one or two bag-like coats, the outer called pr inline, the inner secundine. The nucleus is the essential part ; in it the embryo is formed after fertilization. The coats afterwards become the integuments of the seed. 118. The chalaza is that point at which the base of the nucleus is con- fluent with the coats of the ovule, and is generally discoloured in the seed. The foramen is the common aperture of the coats, opposite the apex of the enclosed nucleus, and through which the pollen is admitted in fertiliza- tion : in the seed it is called the micropyle. 119. Ovules are said to be, ortkotropous or straight (or atropous ), when the chalaza or organic base coincides with the apparent base of the ovule, and the • foramen is situated at the opposite extremity, the ovule having a rectilinear axis : as in the Nettle, Dock, Fig, etc. campylotropous or incurved, when, the base remaining the same, the axis is curved down and the foramen directed towards the base ; as in the Caryophylleee and many leguminous plants. anatropous or inverted, when the chalaza, in an ovule with rec- tilinear axis, is removed to the point most distant from the hilum, and the foramen brought close to the hilum. It is like an orthotropous ovule reversed on its cord ; the cord adhering to one side of the ovule and becoming more or less incorporated with its coats. Such an adhering cord, appearing either like a line or a ridge, is called the raphe : it connects the hilum with the chalaza. Anatropous ovules are much the commonest; good examples of distinctly marked raphe and chalaza may be found in the Orange and the Pansy. amphitropous or half-inverted, when the raphe extends but half the length of the ovule, and the chalaza and foramen, at opposite ends, are about equidistant from the hilum : as in the Mallow tribe, the Primrose, etc. § 14. The Receptacle and relative attachment of the Floral Whorls. 120. The Receptacle, or torus, is the extremity of the peduncle (above the calyx) upon which the corolla, stamens, and ovary are inserted. It is sometimes little more than a mere point, but it is often more or less lengthened, thickened, or otherwise enlarged. [The term receptacle is also extended to the summit of a branch or inflorescence, on which the flowers of a head are inserted, as in the Compositce.~\ 121. A dish, or disc, is a circular enlargement of the receptacle, usually cup-shaped, flat, or cushion-shaped ( pulvinate ), and often of a waxy or fleshy appearance. It is situated either immediately at the base of the ovary within the stamens, or between the petals and stamens, or bearing the petals or stamens or both at its edge, or quite at the extremity of the receptacle, with the ovaries arranged in a ring round it or under it. 122. The disk may be entire, toothed, lobed, or divided into a number of parts, usually as many or twice as many as the stamens or carpels. When the parts of a divided disk are separate and short, they are often called glands. 123. Nectaries are either the disk, or small deformed petals, or abortive XV111 INTRODUCTION TO BOTANY. stamens, or appendages at the base of the petals, or stamens or any small bodies within the flower which do not look like petals, stamens, or carpels. The term nectary is nearly obsolete. 124. When the disk bears the petals and stamens on its rim, it is fre- quently adherent to and confluent with the tube of the calyx : or it is ad- herent to the outside of the ovary ; or adherent both to calyx-tube and ovary. Hence arise the three following important distinctions in the relative insertions of the floral whorls : — Petals and stamens, or, as in common language, flowers, are, hypogynous (i. e. under the ovary), when they or the disk that hears them are entirely free or separate from both calyx and ovary. The ovary in this case is said to be free or superior ; the calyx free or inferior ; the petals as being inserted on the rece2)tacle. perigynous (i. e. round about the ovary), when the disk hearing the petals is quite free from the ovary, hut is more or less com- bined with the calyx-tube. The ovary is then still described as free or superior , even though the combined disk and calyx- tube may form a deep cup with the ovary lying in the bottom ; the calyx is said to he free or inferior ; the petals as being in - serted on the calyx. epigynous {i. e. upon the ovary), when the disk hearing the petals is combined both with the base of the calyx-tube and the outside of the ovary ; either closing over the ovary so as only to leave a passage for the style, or leaving more or less of the top of the ovary free, hut always adhering to it above the level of the insertion of the lowest ovule (except in a few cases when the ovules are suspended from the apex of the cell). In epigynous flowers the ovary is described as adherent or inferior ; the calyx as adherent or superior ; the petals as inserted on or above the ovary. 125. When there are no petals, the insertion of the stamens determines the difference between hypogynous, perigynous, and epigynous flowers. 126. When both petals and stamens are present, in hypogynous flowers the petals and stamens are usually separate, but sometimes they are confluent at base. In that case, if the petals are distinct from each other, and the stamens are mona- delphous, the petals are often said to he inserted on, or com- bined with , the staminal tube ; if the corolla is gamopetalous and the stamens distinct from each other, the latter are said to he inserted on the tube of the corolla. in perigynous flowers the stamens are usually inserted immediately within the petals, or alternating with them on the edge of the disk, or even on the unenlarged part of the receptacle^ in epigynous flowers, when the petals are distinct, the stamens are usually inserted as in perigynous flowers ; when the corolla is gamopetalous, the stamens are often combined at the base with the tube of the corolla, or, as it is more frequently expressed, inserted on the tube. 127. When the receptacle is distinctly elongated below the ovary, it is often called a gynobase , gynophore , or stalk of the ovary. 128. An epigynous disk is a name given either to the thickened summit of the ovary in epigynous flowers, or very rarely to a real disk or en- largement of the receptacle closing over the ovary. 129. In the relative position of any two parts of the flower, whether in the same or in different whorls, they are, connivent, when nearer together at the summit than at the base. divergent, when further apart at the summit than at the base. INTRODUCTION TO BOTANY. XIX Any two parts of the flower are, coherent, when united together, but so slightly that they can be separated with little or no laceration. Each of the two cohe- rent parts may be said to be adherent to the other, but the latter term is often used to express a closer union than mere coherence. [Some authors restrict cohesion to the connation or confluence of parts of the same whorl ; and adhesion to the connation or confluence of parts of different whorls.] connate , when so closely united that they cannot be separated with- out laceration. Each of the two connate parts, and especially that one which is considered the smaller or of the least import- ance, is said to be adnate to the other. free, when neither coherent nor connate. distinct is also used in the same sense, but is likewise applied to parts distinctly visible, or distinctly limited. § 15. The Fruit . 130. The Fruit consists of the ovary and whatever other parts of the flower persist at the time the seed is ripe, usually enlarged and altered in shape and consistence. It encloses or covers the seed or seeds till the period of maturity, when it either opens for the seed to escape, or falls to the ground with the seed. 131. Fruits are often said to be simple, when formed in a single flower ; compound (or more properly collective'), when they proceed from several flowers closely packed or combined in a head. In descriptive botany a fruit is always supposed to result from a single flower, unless the contrary be stated. In compound fruits (the fruits of several flowers) the involucre or bracts often persist and form part of the fruit, but very seldom so in simple fruits. 132. The pericarp is the portion of the fruit formed of the ovary and whatever adheres to it exclusive of and outside of the seed or seeds, ex- clusive also of the persistent receptacle, or of whatever portion of the calyx persists round the ovary without adhering to it. 133. Fruits may be divided into succulent (including fleshy, pulpy , and juicy) and dry. They are dehiscent when they open at maturity to let out the seeds ; indehiscent , when they do not open spontaneously, but fall off with the seeds. Succulent fruits are almost always indehiscent. 134. The principal succulent fruits are, the berry , in which the whole substance of the pericarp is fleshy or pulpy, with the exception of the outer skin or rind, called the epicarp. The seeds are usually immersed in the pulp. the drupe or stone-fruit, in which the pericarp, when ripe, consists of two distinct portions, an outer succulent one called the sarcocarp or mesocarp (covered by a skin or epicarp) and an inner dry en- docarp, called the putamen or stone. When there are two or more stones, they are called pyrenes. 135. The principal dry fruits are, the achene, or akene, including all one-seeded, dry and hard, inde- hiscent, seed-like small fruits, popularly called “naked seeds.” Such fruits may arise from free one-seeded carpels (as in the Buttercup) ; or from adherent or inferior carpels (as in the Composites.) the utricle, similar to the akene, but with a thin and loose mem- branous pericarp. the nut, a hard, one-celled, one-seeded fruit like an akene, but larger, and usually resulting from a plurilocular ovary, all of whose cells and ovules, save one, become obliterated in the ripe fruit ; as in the Hazel-nut, Acorn, etc. c 2 XX INTRODUCTION TO BOTANY. the samara or key-fruit , a nut or akene, having a broad wing at apex or margin (as in the Ash). All the above are indehiscent. The principal dehiscent dry fruits are, the follicle, a pod formed of a single free carpel, dehiscent, along its ventral or seed-bearing suture only (as in the Larkspur , the Asclepiadece, etc). the capsule, a pod or dehiscent fruit of any compound pistil, whether formed from an inferior or a superior ovary. The pyxis, or pyxidium, is only a capsule which opens by a circular, horizontal, nearly medial line, cutting off the upper half like a lid. 136. Peculiar names given to the fruit or parts of the fruit in Cruciferce, Leguminosce, Rosacece, Cucurbitacece, JJmbelliferce, and some other large Orders, will be explained under those Orders. 137. The dehiscence of a capsule is said to be septicidal, when the carpels separate at the line of junction; in this case the placentae are either marginal, or attached to the more or less inflexed margins, constituting the dissepiments. The dehiscence is locuUcidal, when the margins of the carpels remain joined, while the dorsal sutures split open ; in this case the placentae or dissepiments will be borne in the middle of the valve. Septi- fragal dehiscence, in which the valves fall away, leaving persistent dis- sepiments or axile placentae, may occur either in septicidal or loculicidal capsules. Circular, horizontal dehiscence (as in a pyxis) is called circum- scissile. When in a fruit, consisting of one-seeded carpels, the carpels fall away either closed or nearly closing round the seed, each segregated carpel is called a coccus . § 16. The Seed. 138. A seed is the fertilized ovule arrived at maturity. It is almost always, except in Conifers, enclosed in the pericarp. It contains, when ripe, an embryo or young plant, either filling or nearly filling the cavity, but not attached to the outer shell or skin of the seed ; or immersed in, or lying close to, a mealy, horny, oily, or fleshy substance, called the albumen or perisperm. The presence or absence of this albumen, that is, the dis- tinction between albuminous and exalbuminous seeds, is one of great im- portance. The embryo or albumen can often only be found or distin- guished when the seed is quite ripe, or sometimes only when it begins to germinate. 139. The shell of the seed consists usually of two separable coats. The outer coat, called testa, is usually the principal one, and in most cases the only one attended to in descriptions. It may be hard and crustaceous, or thin and membranous, or thin and chartaceous or papyraceous (like parch- ment or paper), or rarely succulent. It is sometimes expanded into wings, or bears a tuft of hair, cotton, or wool, called a coma. The inner coat is called tegmen. 140. The funicle (115) or stalk by which a seed is attached to the placenta, is occasionally enlarged into a membranous, pulpy, or fleshy ap- pendage, which sometimes almost closes over the seed ; this is called an aril. A strophiole or caruncle is a similar appendage, originating in the testa. Seeds having an aril are said to be arillate. 141. The hilum (115) is the scar left on the seed when it separates from the funicle. The micropyle (118) is the mark indicating the position of the foramen of the ovule. 142. The Embryo consists of the radicle or root-stem ; one or two coty- ledons or primary leaves (seed-leaves) ; and the plumule , or first leaf-bud of the young plant. In many seeds, especially when there is no albumen, these several parts are very conspicuous ; in others they are difficult to dis- INTRODUCTION TO I30TANY. XXI tinguish ; and in some cases the embryo cannot be found until the seed begins to germinate. 143. The micropyle (141) always indicates the position of the extremity of the radicle, whose direction, either as respects the fruit or the seed, it is often important to notice. The radicle is said to be superior , if pointing towards the summit of th e fruit, inferior , if pointing towards the base of the fruit. § 17. Accessory Organs. 144. Under this name are included various external parts of plants which often do not appear to act any essential part, either in the vegeta- tion or reproduction of the plant. They may be classed under four heads — Tendrils and Hooks , Thorns and Prickles , Hairs , and Glands. 145. Tendrils are either abortive petioles, or abortive peduncles, or abortive ends of branches. They are simple or branched, flexible, and coil round any object within their reach, in order to support the plant to which they belong. Hooks are similar holdfasts, but of a firmer consistence, not branched, and only hooked at the extremity. 146. Thorns and Prickles. A thorn or spine is a sharply-pointed, rigid extremity of a branch, or abortive petiole, or abortive peduncle ; it is organically connected with the woody system of a plant. A prickle is a sharply-pointed, rigid excrescence from the epidermis , or outer skin ; it is not connected with the woody system, and may occur on a branch, on the petiole, on the veins of a leaf, on the peduncle, or even on the calyx or corolla. A plant is spinous if it has thorns, aculeate if it has prickles. 147. Hairs , in the general sense, or the indument (or clothing) of a plant include all those processes .from the epidermis which have been called bristles , hairs , down, cotton , or wool. The epidermis or surface is said to be, smooth or even , when without any roughness whatever. glabrous , when without hairs of any kind; glabrescent , or glabrate , when the hairs are deciduous. striate , when marked with parallel lines, either slightly raised or merely coloured. furrowed or ribbed , when the parallel lines are more distinctly raised. The epidermis, or surface, is said to be, viscous, viscid , or glutinous, when covered with a sticky or clammy exudation. tuber culate or warted, when covered with minute, blunt, wart-like prominences. muricate, when covered with short, hard, sharp prominences. punctate, when covered with minute dots. foveolate, when covered with small pits. echinate , when the prominences are longer and sharper, almost prickly. setose, or bristly, when bearing stiff, straight hairs. glandular-setose , when the' setce, or bristles , are tipped with a minute, glandular head or drop. glochidiate , when the setae are hooked at the point. pilose , when the surface is thinly sprinkled with rather long, simple hairs. hispid, when more thickly covered with rather stiff hairs. hirsute, when the hairs are dense and not so stiff’. downy ore pubescent, when the hairs are short and soft; puberulent, when very short or minute ; velvety or velutinous, when very dense, like the pile of velvet. XXII INTRODUCTION TO BOTANY. The epidermis, or surface, is said to be, strigose, when short, stiff hairs lie close-pressed to the surface, and all in one direction ; strigillose , when such hairs are very short or small. tomentose or cottony , when the hairs are soft, short, dense, somewhat interwoven, and usually white or whitish. woolly , when the hairs are loosely intricate and long, like wool. mealy or farinose , when the hairs are very minute, intricate, and white, and come off readily, having the appearance of meal or dust. canescent , canons , or hoary , when the hairs are minute, close-pressed, and white, and not readily to be distinguished separately by the eye, but giving a general whitish hue to the epidermis. glaucous or glaucescent, when of more or less a pale bluish-green, often covered with a bloom like that on a plum or cabbage-leaf. 148. Hairs are often branched. If forked from the base, the forks spreading in opposite directions, the hairs are said to be attached by the centre. If several branches radiate horizontally, the hairs are stellate , or star-like. Stellate hairs become stellate scales when the rays are confluent at base ; and the surface is said to be scaly or lepidote. 149. The term gland is given to several different productions, and principally to the four following : — 1. Small, wart-like or shield-shaped bodies, either sessile or some- times stalked, of cellular or somewhat fleshy consistence, occa- sionally secreting a small quantity of oily or resinous matter, but more frequently dry. They are generally few in number, often definite in their position and form, and occur chiefly on the petiole or principal veins of leaves, on the branches of in- florescences, or on the stalks or principal veins of bracts, sepals, and petals. 2. Minute raised dots, usually black, red, or dark-coloured, of a resinous or oily nature, always superficial and apparently exudations from the epidermis. They are often very numerous on leaves, bracts, sepals, and green branches, and occur even on petals and stamens, more rarely on pistils. When raised on slender stalks they are called pedicellate glands , or glandular hairs, according to the thickness of the stalk. 3. Small, globular, oblong, or linear vesicles filled with oil, im- bedded in the substance of leaves, bracts, floral organs, or fruits. They are often very numerous, like transparent dots ; sometimes few and determinate in form and position. In the pericarp of Umbelliferce they are remarkably regular and con- spicuous, and take the name of vittce. 4. Lobes of the disk, or other small, fleshy excrescences within the flower, whether from the receptacle, calyx, corolla, stamens, or pistils. II. Anatomy and Physiology. (. Abridged from the writings of Professor Lindley and Professor Asa Gray.) 150. Vegetable Anatomy , or the study of the microscopical structure of the compound organs of plants, and Vegetable Physiology , or the study of the functions which each organ performs during life, are distinct and extensive branches of botany, with which the merely systematic botanist, INTRODUCTION TO BOTANY. XX111 or the student who uses a Flora for the purpose of ascertaining the names of plants, has not necessarily much concern. In this outline, therefore, we shall treat of these cognate sciences very briefly. § 1. The Elementary Organs. 151. If a very thin slice of a plant (say, of a succulent leaf or fruit) be magnified, it will be found to be made up of variously-shaped and arranged ultimate parts or elementary organs , forming a sort of honeycombed structure. The ultimate parts are called cells. 152. A cell, in its simplest state, is a closed membranous sac, formed of a substance permeable by fluids, though usually destitute of visible pores. When cells are combined, the mass is called a tissue ; but each cell is a distinct individual, separately formed and separately acting, though co- hering with the cells with which it is in contact, and partaking of the common life and action of the tissue of which it forms a part. The mem- branes separating the cells are called their walls. 153. Botanists usually distinguish the following tissues : — (a) cellular tissue , called also pulp and parenchyma, consists of roundish, oblong, cylindrical, hexagonal, or stellate thin-walled cells, and is found in every plant. All the soft part of leaves, the pith of stems, the pulp of fruits, and all young growing parts are formed of cellular tissue ; and very many crypto- gamic plants possess no other tissue. In it also are centred the most active functions of the living vegetable. It is the first tissue formed, and continues to be formed while growth continues, and when it ceases to be active, the plant dies. (5) woody tissue, or pleurenchyma, consists of long, slender cells tapering to each end, of a thicker, stronger, and much tougher substance than cellular tissue, but otherwise similarly or- ganized. It constitutes the principal part of the wood, of the fibrous inner bark, and of the nerves and veins of leaves, sepals, and petals. It is not found in the lower Orders of Cryptogams. (c) vascular tissue, or trachenchyma, consists of very thin-walled, elongated tubes, variously marked. This tissue is of two prin- cipal kinds, the spiral vessel and the duct. Spiral vessels , often called tracheae, contain highly elastic spiral fibre, usually capable of being unrolled ; they meet or overlap at the ends, and where two such vessels adhere, the intervening membrane is absorbed, * and they communicate freely. They are found round the pith in stems, and in all parts that emanate from it, especially along the nerves and veins of leaves ; very rarely they occur in the wood or bark. Ducts are tubes usually of much greater diameter and length than the spiral vessel, containing a spiral fibre in- capable of being unrolled, and often broken into imperfectly spiral bars, or rings, or dots, or disposed like the rungs of a ladder. They occur chiefly in the wood ; are abundant in the wood of Ferns ; but absent from the wood of Conifers ; their functions are not clearly ascertained. (d) luticiferous tissue , or cinenchyma , consists of uninterrupted, ana- stomosing, thick-walled tubes, which contain a peculiar fluid called latex, usually turbid ; often coloured red, white, or yellow, but often colourless. The use of this tissue is unknown. 154. Various modifications of cellular, woody, and vascular tissues are distinguished by vegetable anatomists, but need not be here enumerated. Other false elementary organs, such as air-vessels , cysts, turpentine-vessels , oil-reservoirs, etc., are all either intercellular cavities, or large cells filled with peculiar secretions. XXIV INTRODUCTION TO BOTANY. 155. A young and vitally active cell consists of the following parts : — 1, the outer wall, a permeable, transparent membrane, formed of a chemical substance called cellulose ; 2, a mucilaginous film lining the wall, and called “ the primordial utricle 3, the nucleus , or centre of cell-function or life, a soft, subgelatinous body occupying the middle of the cell, or ex- centrical; and 4, a viscid fluid, called protoplasm, filling the space be- tween the nucleus and the primordial utricle. As the cell increases in size, its contents change ; and finally, when it has attained its proper dimensions, the wall formed of cellulose alone remains as a persistent fabric, the nucleus is absorbed or dried up, and the protoplasm passes out into younger cells. 156. The principal organized contents of cells are : — (a) sap , the first product of the digestion of the inorganic food of plants : it contains the elements of vegetable growth in a dissolved condition. ( b ) sugar , of which there are two kinds, cane-sugar and grape-sugar , usually exists dissolved in the sap. It is found abundantly in growing parts, in fruits, and in germinating seeds. (c) dextrine , or vegetable mucilage, a gummy substance intermediate between sugar and starch. ( d ) starch or fecula , one of the most universal and conspicuous of cell-contents, and often so abundant in farinaceous roots and seeds as to fill the cell-cavity. It consists of minute grains, called starch-granules , which vary in size and shape, and are marked with more or less conspicuous concentric lines. Starch is unaffected by cold water, but forms a jelly with boiling water, and turns blue when tested by iodine. {e) chlorophyll , the green matter of plants, is of a resinous nature, and contains nitrogen. It is formed only under the action of sunlight, and is usually most abundant in the layers of cells immediately below the surface. (/) chromule , a name given to a similar colouring-matter when not green. (y) wax, oils , camphor , and resinous matters are common in cells ; also various mineral substances, either in an amorphous state or as microscopic crystals, when they are called raphides. These last are peculiarly abundant in the tissues of the Cacti and Rhubarb. § 2. The Epidermis and its Processes. 157. The Epidermis, or outer skin of plants, is formed of one or more layers of vertically flattened, firmly coherent, and usually empty cells, with thin and transparent, or with thick and opaque walls. It covers all parts exposed to the air, except the stigma and glands ; bfit is absent in parts submerged under water. It serves to protect the tissues from the immediate action of the air or of drought. 158. The epidermis is pierced by minute spaces between the cells, called stomates. They are oval or mouth-shaped, bordered by lips, formed of two or more elastic cells so disposed as to cause the stomate to open in a moist, and to close up in a dry state of the atmosphere. They communicate with intercellular cavities, and are obviously designed to regulate evaporation and respiration. Stomates are found abundantly on leaves, especially on the lower surface ; in succulent leaves, though abundant, they are often in an imperfectly organized condition, and seemingly inactive. 159. Hairs are minute, transparent expansions of cellular tissue, pro- ceeding from the epidermis. The hairs of roots are active absorbents ; INTRODUCTION TO BOTANY. XXV those of stems and leaves serve to protect the surface, or to control evapo- ration, and sometimes act also as absorbents. They are more frequent on plants inhabiting dry or exposed situations, or on veiy Alpine plants, which are alternately exposed, at brief intervals, to extremes of heat and cold. § 3. The Root. 1 60. Anatomically the root differs from the stem in wanting normally developed leaf-buds (29), stomates (158), and in Exogens, a distinct pith. It increases in length by constant small additions to its extremity, and thus is enabled to force its way through the soil, and to diverge when rocks or obstacles oppose its progress. 161. The functions of the root are to fix plants in or to the soil or other substance on which the plant grows, to absorb nourishment either from the soil, or in the case of aerial roots, from the air, and to transmit it rapidly to the stem. Absorption takes place through the younger fibres or the extremities of the root-branches, and through hairs which are formed on all young roots, when growing vigorously. The nutriment thus absorbed consists chiefly of carbonic acid and nitrogen, or nitrogenous compounds, dissolved in water. 162. Parasites are plants whose roots bury themselves in the cellular structure of other plants, and subsist on nourishment sucked out of the plant which they attack. Epiphytes are those whose roots attach them- selves to other plants without penetrating into their cellular tissues. § 4. The Stem. 163. Anatomically the stem consists of a cellular and a fibro-vascular system. The cellular system developes both vertically, as stems increase in length, and horizontally, as they increase in diameter. The fibro- vascular system is gradually introduced vertically, and serves to bind together and strengthen the cellular. It may be compared to the bony skeleton, the cellular to the flesh. 164. The stems of flowering plants are formed on one or other of the two following types : — the Exogenous structure , in which the woody system is deposited in annual concentric layers between a central pith and an exterior separable bark. Plants having this structure of stem are XZxogens. the Endogenous structure , in which the woody system is deposited, not in concentric rings, but in separate, small bundles or threads of woody fibre, running through the cellular system without apparent order. In such stems there is no distinct pith, nor separable bark. Plants having this structure of stem are called Endogens. 165. The stems of the lower Cryptogamia consist wholly of cellular tissue ; those of Ferns have an imperfect fibro-vascular system derived from the bases of old petioles. 166. In an exogenous stem a new layer or ring of wood is annually formed between the outermost preceding layer and the inner surface of the bark. In an endogenous , the new wood bundles are mingled with the old, or deposited toward the centre of the stem, whence they diverge to- wards the circumference in the lower part of their course. In an Exogen therefore the oldest, hardest, and most compact wood is found towards the centre of the stem ; in an Endogen towards the circumference. 167. Anatomists distinguish the following parts in an Exogenous stem: — {a) the pith, which is only active in young stems or growing XXVI INTRODUCTION TO BOTANY. branches, consists of cellular tissue, occupying the centre or longitudinal axis of the stem. (b) the medullary sheath , which surrounds the pith, abounds in spiral vessels (153, c), and is in direct connection, through its ramifications, with the leaf-buds, and the veins and nerves of leaves. (c) the wood, which lies directly on the medullary sheath, is formed of woody tissue (153, b ), through which, in most cases, ducts (153, c), variously disposed, are interspersed. A new circle of wood is annually formed, on the outside of the circle of the previous year ; the age of a stem therefore may be ascer- tained, in a large number of cases, by counting the numbers of its rings of wood : in some cases of tropical trees and ever- green trees of temperate climates, several rings of wood are formed in a year. The older and denser, comparatively sap- less wood, is called heartwood of duramen , and is often coloured ; the younger, living and incompletely formed, is the sapwood or alburnum , and is usually white. {d) the medullary r ays, which originate in the pith, traverse the wood, and terminate in the bark, and are formed of cellular tissue : they occur as vertical plates, radiating from a centre, and keep up a communication between the living portion of the interior of the stem and its outer surface. As the heartwood is formed the inner portions of the medullary rays die. In wood they are what carpenters call the silver-grain. (e) the bark, which lies outside the wood, and forms the outer layer of the stem. It is coated by the epidermis (157), and like the wood, consists of concentric layers ; namely, the corky layer, or dry, outer bark, formed of hard, compressed cells ; the cellular or green or middle bark, formed of loose, thin-walled, pulpy cells, containing chlorophyll (156, e) ; and the liber or inner bark, formed of long, tough, woody tissue, called bast-cells. The liber, like the wood, is annually deposited ; the green layer is a product of the first year only, being soon choked by the corky envelope. 168. The mineral food of plants, absorbed by the roots, passes upwards through the younger wood of the stem, mixing with previously organized matter, but not being essentially altered ; in this state it is called sap or crude sap. The crude sap, as it ascends through the stem, is attracted into the leaves, where it is exposed to the direct action of sunlight, under which influence alone can assimilation take place. As assimilated or elaborated sap, it is returned into the stem, and either used up in the processes of further growth, or deposited either in the wood, in the stock, in the peren- nial part of the stem or root, or in any other part of the plant where matter is stored up for future use. § 5. The Leaf. 169. Anatomically the leaf consists of a central fibro-vascular system or woody skeleton, derived from the woody system and medullary sheath of the stem ; a cellular system surrounding the fibro-vascular, and interwoven with it, and derived from the middle bark ; and an outer skin or epidermis, pierced by stomates. 170. The fibro-vascular system is arranged on two principal types : — (a) the exogenous, in which the nerves and veins branch irregularly and usually anastomose into a sort of network. (5) the endogenous, in which the principal nerves usually extend INTRODUCTION TO BOTANY. XXY11 unbranched from the base to the apex, and are connected by cross-bars or unbranched veins. The first of these types is generally characteristic of Exogens , the latter of Endogens ; but there are various intermediate conditions, and some Endogens have been called Dictyogens, because they have netted- veined leaves ; several Exogens also have straight- veined leaves. 171. Leaves usually extend horizontally, and have an upper and an under surface, differing in anatomical structure. In the cellular stratum of the upper surface the cells are closely set and placed vertically, with their smallest ends next the surface ; in the lower stratum the cells are more or less horizontal, more loosely placed, and very generally have large empty spaces between them. 172. Leaves are functionally the most active of the vegetable organs. In them the process of digestion or assimilation is chiefly conducted. 173. Assimilation is the name given to the process which accom- plishes these following results 1. The chemical decomposition of the oxygenated matter of the sap, and of the carbonic acid which is absorbed by the leaves ; re- sulting mainly in the liberation of pure oxygen, the fixation of carbon and the elements of water in the tissues of the plant. Thus, through the powers of life, and under the influence of solar light and heat, oxygen , hydrogen , nitrogen , and carbon , or some of these, uniting in certain proportions, become gluten, chlorophyll, gum, sugar, or starch, etc. ; and in like manner all other vegetable compounds pass from dead matter into the condition of living substance. 174. The oxygen liberated by plants during the process of assimilation passes into the air ; and as assimilation is constantly going on during sun- light, the amount of oxygen thus poured into the air by plants is enormous, and indeed, so far as we know, vegetation is the only great operation in nature which restores to the air that free oxygen gas which is consumed by animals, and in all processes of combustion is indispensable to animal life. III. Classification. 175. It has been already said (2) that descriptions of plants should be arranged, as nearly as possible, under natural divisions, so as to facilitate the comparison of each plant with those most nearly allied to it. The descriptions here alluded to are descriptions of species ; the natural divisions of the Flora refer to natural groups of species. 176. A species comprises all the individual plants which resemble each other sufficiently to make us conclude that they may all have descended from a common parent stock. These individuals may often differ from each other in striking particulars, such as colour of flower, size of leaf, etc. ; but such differences, observation teaches us, may occur in seedlings, raised from one individual. 177. When a number of individuals of a species differ from the majority in any striking particular, they constitute a variety. If the variety generally comes true from seed, it is often called a race. 178. A Variety can be propagated with certainty only by grafts, cut- tings, bulbs, or tubers, or other method which produces a new plant by the growth of buds taken from the old one. A race may very frequently, but not with certainty, be propagated by seed. 179. The known species of plants (now nearly 100,000) are far too numerous to be studied without classification. To facilitate their study, an admirable system, invented by Linnaeus, has been universally adopted, viz. one common, substantive name is given to a number of species which XXV111 INTRODUCTION TO BOTANY. resemble each other more than they do any other species ; the species so brought together are collectively called a Genus, and the common name is the generic name . Each species is further distinguished from the others of the same genus by the addition of an adjective- epithet, or specific name. Every species has thus a botanical name of two words. In Latin, the language usually used for the purpose, the first word is a substantive usually of Greek origin and Latinized, and designates the genus ; the second, usually a Latin adjective, indicates the species. 180. The genera thus formed being very numerous, they have been grouped together on similar principles — associating those which resemble each other most nearly — into Families or Natural Orders, to each of which a name has been given. This is, however, for the purpose of study or comparison. In speaking of a species it is only necessary to give the generic and specific names. The name of a Natural Order or Family, in Latin, is an adjective plural, usually formed from the name of some one typical genus, generally the best known, the first discovered, or the most marked. Thus Ranunculacece is the Order of which the Ranunculus is the typical genus ; Geraniacece the Order of the Geranium and its allies, etc. 181. The number of species included in a genus, or the number of genera in an Order, is very unequal. Some genera contain but one, others but two or three species ; in others, several hundred species are associated together. There is a similar discrepancy in the number of genera in the several Orders. 182. Orders are collected into Classes ; and where Orders contain a large number of genera, or genera a large number of species, a further subdivision is required. The names of the several groups most generally adopted are as follows, beginning with the most comprehensive or highest divisions — Classes : Genera : Subclasses or Alliances. Natural Orders : Suborders. Tribes. Subtribes. Divisions. Subdivisions. Subgenera. Sections. Subsections. Species : Varieties. 183. Classes, Orders, Genera, and their subdivisions, are called natural , when, in forming them, all resemblances and differences are taken into account, valuing them according to their importance ; artificial , when resemblances and differences in some one, or very few particulars only, are taken into account. IY. Collection, Preservation, and Examination of Plants. 184. Though plants can be most easily and satisfactorily examined when freshly gathered, yet as time will rarely admit of this being done, and as it is also desirable to compare with other plants previously observed or collected, specimens must be selected for leisurely observation at home, and for preservation for future reference. 185. A botanical Specimen should be, as much as possible, an epitome of the species which it represents, and if the species be variable, several specimens are necessary. To be perfect, it should have root , stem , leaves, fiowers (both open and in bud), and fruit (both young and mature). It is not always, however, possible to gather such complete specimens ; but the INTRODUCTION TO BOTANY. XXIX collector should aim at completeness. Fragments , such as leaves without flowers, or flowers without leaves, are of comparatively little use. 18(3. All small plants , not exceeding 15 inches in height, should be plucked up by the roots, the whole plant forming a single specimen ; or, if a many-stemmed plant, or one much branched near the base, and which if dried whole would make too dense a mass of branches or stems, it may he divided, at the origin of the branches , into several specimens. 187. If the plant to he dried he of greater length than 15 inches, and if it he thought desirable — either from the lower leaves differing from the upper, or from the branches being long and naked — to preserve the stem unbroken , the specimen may he folded in lengths of 15 inches. This is often done with Grasses , Sedges , and Ferns ; and should be done in the cases of all long-stemmed, lax -leaved herbaceous, and with tall bulbous plants, in order to preserve an indication of their habit. 188. Herbaceous plants of large size, and specimens of the branches of shrubs and trees must he broken into pieces, say 10-15 inches long, the length varying with the nature or ramifications of the plant. The object is to preserve as much of the peculiar aspect of the plant as is possible. 189. At least a dozen specimens of each plant should (when practicable) be gathered, for the purpose of exchanging with other botanists. A col- lector can scarcely have too many duplicates, especially when his explora- tions are made in a little-frequented district. Many collect much more extensively. 190. The collector should he provided with : — 1. A quantity (at his discretion) of any stout, coarse, unsized paper, of uniform dimensions, say 12 inches by 18. Old newspapers answer the purpose, and common packing-paper, whity-brown, or brown, is most excellent. Blotting-paper is much too tender and expensive. 2. A smaller quantity of very thin, unglazed paper, or chemist’s filtering -paper, for drying plants with delicate corollas (see 200). 3. Several flat, perforated boards, the size of the paper. Open wooden frames, with cross-bars, or frames of strong wire-work lattice, are better than boards, as they permit a freer evapora- tion. 4. A light portfolio of pasteboard, covered with calico, fitted up with 12-20 leaves of strong brown paper, furnished with a strap and buckle for closing, and another for slinging over the shoulders, is better and more portable than the old-fashioned collecting- box. The specimens, as gathered, are placed between the leaves, and may be crowded together, if not left too long without sorting. 5. A bag or haversack is also useful for collecting rigid-leaved or shrubby plants, that might injure those in the portfolio. 191. If the plants be gathered in dry weather, no time should be lost in placing them under pressure ; but they may be preserved for a day or two, if sprinkled with water, and enclosed in a tin box in a cool situation. Ticket the specimens, and add notes made whilst collecting. 192. On returning from the field, sort the specimens into those that are fleshy or juicy , and those that are of a drier nature, and dry them in separate bundles. If mixed together, the former are very apt to injure the latter, and to retard their drying. 193. The drying process is as follows : — Take one of the flat boards or frames, and lay three or four sheets of the drying-paper upon it. On these lay specimens, placing them as closely as they will lie without over- lapping each other. Cover the specimens with a similar layer of paper ; and on this lay other specimens ; repeating alternately a layer of paper XXX INTRODUCTION TO BOTANY. and of specimens, till you have either placed in paper all the specimens collected, or made a sufficiently thick pile. Cover the pile with one of the flat hoards, and place upon it a heavy weight, — large stones or hags of sand answer perfectly. If travelling, leather straps and buckles, drawn tightly across the bundle, are used instead of weights. 194. After the specimens have lain a day under pressure, the paper about them must be removed, and dry papers substituted ; and this process should he repeated at intervals of a day or two till the plants are perfectly dry. If many sheets of paper he placed between each layer of specimens, or if open frames he used instead of boards, the changes need not he so frequent. In changing it is not necessary to lift every specimen from the sheet on which it lies ; hut if a dry sheet he placed over the specimens, the latter, with the moist sheet, may he tilted over to the dry, and the moist sheet then removed, and this process repeated through the bundle. Much time and trouble may thus he saved. 195. On the first day of shifting a sharp look-out should he kept for caterpillars , which are apt to secrete themselves in flowers, and, if not at once removed, will quickly destroy the specimens under pressure. 196. In fine weather the bundles of specimens, weighted or strapped, may he exposed to the strongest heat of the sun ; hut as this causes a rapid extraction of moisture, in order to ensure its passing away, the plants must, on bringing in, and while still warm, he shifted into fresh papers : otherwise mouldiness and decay, and not exsiccation, will ensue. Artificial heat, not greater than 140°, may he substituted in wet weather. 197. In drying plants within the tropics, and in all damp and hot climates, frequent shifting of papers is necessary ; if neglected, the specimens will either fall to pieces, or become mouldy and rotten. 198. Fleshy fruits should he preserved in spirits ; or carefully stretched, and the seeds or hard parts dried. 199. Succulent plants, Heaths, and plants with compound leaves (such as Mimosece ), should he dipped (all hut the flowers) for a few seconds into boiling water, before being placed in the drying-papers. This will kill them, promote the drying of succulents, and prevent the Heaths, etc., from shedding their leaves. 200. Plants with delicate corollas ( Iridecc , Oxalidece, etc.), should he placed between single leaves of very thin and soft, unglazed paper (filtering or tissue-paper). In shifting to dry papers the tissue-paper is not to he removed, hut lifted with its contents to the dry layer. This will prevent the flowers from curling up or perishing. 201. Wlien the specimens are quite dry and stiff, a single sheet of paper is sufficient between each layer ; they may he placed still more closely on the sheets, hut not piled one on another ; and, finally, in sending specimens from a distance, great care must he taken to protect the bundles, by sufficient covering, from the effects of external moisture, or from the attacks of insects. 202. Having dried his plants, the student begins to form his Herba- rium, or Hortus Siccus. The first step is to assort the specimens, first into their classes ; then into subclasses , Orders , genera , and species. When this is done, he selects such specimens of such species as he wishes to retain for future reference, consigning the remainder to separate bundles labelled “ duplicates ,” and keeping them for exchange with other botanists. 203. The specimens selected, having first been examined and named, are either fastened with thin glue to pieces of stiff white or cream-coloured cartridge-paper of a uniform size, or placed loosely, or secured by cross- bands or pins, in double sheets of soft paper. In either case the species belonging to each genus are placed within a common wrapper of strong INTRODUCTION TO BOTANY. XXXI paper, and the name of the genus and Natural Order to which it belongs written on the left-hand comer near the base. 204. The most convenient size for the sheets of paper is 16 inches long by 10£ wide. 205. No more than one species should ever be fastened on one sheet of paper ; but as many specimens as will conveniently fit may be placed side by side. 206. The name, place of growth, soil, elevation above the sea, season of flowering, colour of the flower, and if a shrub or tree, the height, nature of the wood, etc. and any useful information respecting the species, and not deducible from the specimen, — should be written on an attached label or on the sheet to which the specimen is fixed. 207. The sheets of species arranged under their genera and Orders, are kept in cabinets made with compartments to suit the size of the paper used. 208. When it is required to examine or dissect flowers or fruits that have been dried, it is necessary to soften them. If the parts are very delicate, this is best done by gradually moistening them in cold water ; in most cases steeping them in boiling water or in steam, is usual, and is much quicker. Very hard fruits and seeds will require long boiling in order to dissect them easily. 209. For dissecting and examining flowers in the field, a penknife and a pocket lens of two or three powers, from half an inch to two inches focus, are sufficient. At home it is more convenient to have a mounted lens or simple microscope, with a stage holding a glass plate, upon which the flowers to be dissected may be laid, and a pair of dissecting -knives, one of which should be narrow and pointed, or a fine needle fixed in a handle ; the other should have a pointed blade, with a sharp edge, to make clean sections across the ovary. A compound microscope is rarely necessary, except in cryptogamic botany. For the simple microscope, lenses of i, a-, 14 inch focus are sufficient. 210. Many species vary within limits which it is very difficult to express in words. It may also happen that the specimen gathered may present some occasional or accidental anomalies peculiar to that single one, or to a very few individuals of the species. Hence the difficulty constantly ex- perienced by the young student, and not seldom by the more expert botanist, of recognizing the various forms of a species by the technical characters assigned to it in a Flora. It may be useful, therefore, to point out a few of the anomalies likely to be met with, and we may divide them into two classes, as follows : — 1. Aberrations from the ordinary type or appearance of a species , for which some general cause may be assigned. A bright light and open situation, particularly at considerable ele- vations or in high latitudes, without too much wet or drought, tends to increase the size and heighten the colour of flowers, in proportion to the stature and foliage of the plant. Shade, on the contrary, especially if accompanied by richness of soil and sufficient moisture, tends to increase the foliage and draw up the stem, but to diminish the number, size, and colour of the flowers. A hot climate and dry situation tend to increase the hairs, prickles, and other productions of the epidermis, to shorten and stiffen the branches, rendering thorny plants yet more spinous. Mois- ture in a rich soil has a contrary effect. The neighbourhood of the sea, or a saline soil or atmosphere, im- parts a thicker and more succulent consistence to the foliage and almost every part of the plant, and appears not un- XXX11 INTRODUCTION TO BOTANY. frequently to enable plants usually annual to live through the winter. Flowers in a maritime variety are often much fewer, hut not smaller. The luxuriance of plants growing isolated in a rich soil, and the dwarf, stunted character of those crowded in poor soil are well known. It is also well known how gradually the specimens of a species become stunted as we advance into the cold, damp regions of the summits of high mountain-ranges, or into high northern latitudes ; and yet it is very frequently for want of attention to these circumstances that numbers of false species have been added to enumerations and Floras. Luxuriance en- tails not only increase of size of the whole plant or of particular parts, but increase of number of branches, or leaves, or leaflets of a compound leaf ; or it may diminish the hairiness of the plant or induce thorns to grow out into branches, etc. Capsules which, while growing, lie upon or close to the ground, will often become larger, more succulent, and less readily dehiscent, than those which are not so exposed to the moisture of the soil. Herbs eaten down by sheep or cattle, or crushed underfoot, or burnt over, or otherwise checked in their growth, or trees or shrubs cut down to the ground, if then exposed to favourable circumstances of soil and climate, will send up luxuriant side- shoots, often so different in the form of their leaves, in their ramification and inflorescence, as to be scarcely recognizable for the same species. Annuals which have germinated in spring and flowered without check, will often be very different in aspect from individuals of the same species, which having germinated later, are stopped by summer droughts or the approach of winter, and only flower the following season upon a second growth. The latter have often been mistaken for perennials. Hybrids, or crosses between two species, come under the category of anomalies from a known cause. Frequent as they are in gardens, where they are artificially produced, they are probably rare in nature. Absolute proof of the origin of a plant found wild is of course impossible ; but it is pretty generally agreed that the following particulars must always coexist in a wild hybrid. It partakes of the characters of its two parents ; it is to be found isolated or almost isolated, in places where the two parents are abundant ; if there are two or three, they will generally be dissimilar from each other, one partaking mor ■ of one parent, another of the other ; it seldom ripens good seed ; it will never be found where one of the parents grows alone. Where two supposed species grow together, intermixed with numerous intermediates bearing good seed, and passing more or less gradually from the one to the other, it may generally be concluded that the whole are varieties of one species. The beginner, however, must be very cautious not to set down a specimen as intermediate between two species, because it appears to be so in some, even the most striking characters, such as stature and foliage. Extreme varieties of one species are connected together by transitions in all their characters, but these transitions are not all observable in the same speci- men. The observation of a single intermediate is, therefore, of little value, unless it be one link in a long series of intermediate forms, and, when met with, should lead to the search for other connecting links. INTRODUCTION TO BOTANY. XXX111 2. Accidental aberrations from the ordinary type , that is, those of which the cause is unknown. These require the more attention, as they may sometimes lead the beginner far astray in his search for the genus, whilst the aberrations above reduced more or less to general laws, affect chiefly the distinction of species. Almost all species with coloured flowers are liable to occur occa- sionally with white flowers. Many may be found, even in a wild state, with double flowers, that is, with a multiplication of petals. Plants which have usually conspicuous petals will occasionally appear without petals, either to the flowers produced at par- ticular seasons, or to all the flowers of particular plants ; or the petals may he reduced to narrow slips, or variously cut. Flowers usually very irregular (81) may, on certain individuals, lose'more or less of their irregularity. Spurs may disappear, or he produced on all, instead of only one of the petals. One part may he occasionally added to or subtracted from the usual number of parts in each floral whorl, more especially in regular, polypetalous flowers. The relative adhesion of the floral whorls may vary ; hypogynous stamens appearing in flowers usually with perigynous, and free or half-free ovaries in flowers usually with adherent. Plants usually monoecious or dioecious may become occasionally hermaphrodite, or hermaphrodite plants may produce occa- sionally unisexual flowers by the abortion of the stamens or of the pistils. Leaves alternate where they are usually opposite ; cut or divided where usually entire ; variegated or spotted where usually of one colour, or the reverse, — must also he classed among those accidental aberrations which the botanist must always he on his guard against mistaking for specific distinctions. d XXXIV INDEX TO TERMS, ETC. [The numbers refer to the paragraphs in the foregoing Introduction .] Aberrations, 210. Abortive, 67, 93,- 104. Accessory organs, 144. Accrescent, 87. Acerose, 51. Achene, 135. Acicular, 51. Aculeate, 146. Acumen, 45. Acuminate, 45. Acute, 45. Adherent, 124, 129. Adhesion, 129. Adnate, 53, 96, 129. Adventitious, 29. Estivation, 89. Akene, 135. Alabastrum, 65. Albumen, 138. Albuminous, 138. Alburnum, 167. Alternate, 30, 74. Amphitropous, 119. Amplexicaul, 35. Amyloid, 156. Analytical tables, 2. Anastomose, 39. Anatropous, 119. Androgynous, 70. Anisomerous, 80. Annual, 17, 18. Anther, 92, 96. Anthesis, 65. Apetalous, 68. Apex, 34, 45, 97. Apocarpous, 108. Aril, arillus, 140. Aristate, 45. Artificial, 183. Ascending, 24. Asepalous, 68. Assimilation, 168, 171. Atropous, 119. Auricle, 46, 47. Axil, 29, 33. Axile, 116. Axillary, 58. Axis, 109. Bark, 164, 167. Barren, 68, 93, 104. Base, 34, 97. Berry, 134. Bi- (2 in composition). Bicarpellary, 107. Bidentate, 42. Bifid, 42. Bifoliate, 42. Bijugate, 42. Bilabiate, 90. Bilocular, 109. Bipinnate, 42. Bisexual, 68. Biternate, 42. Blade, 34. Bract, 55, 64. Bracteole, 64, 56. Branch, 30, 112. Bristles, Bristly, 98, 147. Bud, 185. Bulb, 20. Bulblet, 20. < Caducous, 87. Csespitose, 24. Calyx, 43, 66, 73, 82. Campanulate, 90. Campy lotropous, 119. Canescent, 147. Canous, 147. Capillary, 51. Capitate, 62, 114. Capitulum, 62, 70. Capsule, 135. Carpel, 76, 102. Caruncule, 140. Catkin, 62. Cauline, 36. Cells (elementary), 151, 152. Cells (of anthers), 92. Cells (of the ovary), 103. Cellular tissue, 8, 153. Cellulose, 155, 173. Centrifugal, 61. Centripetal, 61. Chalaza, 117. Character, 2. Chartaceous, 139. Chlorophyll, 156, 173. Chromule, 156. Ciliate, 38. Ciliolate, 38. Circumscissile, 137. Class, 182, 202. Claw, 88. Cleft, 38. Climbing stem, 25. Coats, 139. Coccus, 137. Coherent, 129. Collection of specimens, 131. Coma, 139. Common petiole, 38. Complete, 67, 72. Compound, 62, 197, 131. Compressed, 51. Confluent, 99. Conical, 51. Connate, 129. Connective, 92. Connivent, 129. Contorted, 89. Convolute, 89. Cordate, 46. Cordiform, 46. Coriaceous, 50. Corm, 22. INDEX TO TERMS, XXXV Corolla, 66, 74, 82, 83. Corymb, 62. Corymbose, 62. Cotton, 147. Cotton^, 147. Cbtyledons, 142. Creeping, 24. Crenate, 38. Crenature, 38. Crenulate, 38. Crestsr 98. Crude sap, 168. Crumpled, 89. Crustaceous, 139. Cryptogamous plants, 10. Culm, 38. Cuneate, 43. Cuspidate, 45. Cylindrical, 51. Cyme, 62. Cymose, 62. Cysts, 154. Deca- or Decern- (10 in composition). Deciduous, 87. Decompound, 41. Decumbent, 24. Decurrent, 35. Decussate, 30. Definite, 72. Dehisce, 100. Dehiscence, 100. Dehiscent, 133. Deltoid, 43. Dentate, 38. Depressed, 51. Dextrine, 156. Di- (2 in composition). Diadelphous, 95. Dialipetalous, 86. Diandrous, 79. Dichlamydeous, 68. Dichotomous, 31, 62. Diclinous, 67. Didynamous, 95. Diffuse, 24. Digitate, 40. Digynous, 79, 107. Dimerous, 79. Dimidiate, 99. Dioecious, 69. Dipetalous, 79. Disepalous, 79. Disk, 121. Dissepiment, 109. Dissolved, 156. Distichous, 30. Divaricated, 97. Diverging, 97, 129. Divided, 38, 122. Divisions, 182. Double flowers, 83. Down, 147. Downy, 147. Drupe, 134. Dry, 133. Ducts, 153, 167. Duramen, 167. Echinate, 147. Elaborated sap, 168. Elementary organs, 7, 15 1 . Elliptical, 43. Emarginate, 45. Embryo, 14, 117, 138, 142. Endocarp, 134. Endogen, 164, 170. Endogenous, 164, 166. Ennea- (9 in composi- tion). Entire, 38, 122. Epicalyx, 64. Epicarp, 134. Epidermis, 146, 157. Epigynous, 124, 126. Epigynous disk, 128. Epiphyte, 162; Erect, 24. Even, 147. Exalbuminous, 138. Exogen, 164, 166, 170. Exogenous, 164, 166. Exstipulate, 53. Extrorse, 100. Families, 179. Fascicled, 30, 37. Fastigiate, 62. Female, 68. Fertile, 68. Fibre, 13. Fibrous roots, 15. Filaments, 92, 93. Flabelliform, 43. Fleshy, 50, 133, 192. Floral whorls, 120. Flower, 10, 65, 68, 79, 84, 185. Flowering plants, 10. Follicle, 135. Foramen, 117. Forked, 31. Foveolate, 147. Free, 53, 72, 80, 124, 129. Fruit, 130, 143, 185. Function, 6. Funiculus, 140. Funnel-shaped, 90. Furrowed, 147. Fusiform, 51. G-amopetalous, 86. Genus, genera, 182, 202, 203. Gibbose, 90. Glabrate, 147. Glabrescent, 147. Glabrous, 147. Glands, 98, 144, 149. Glandular-setose, 147. Glaucous, 147. Globular, 51. Glochidiate, 147. Glume, 64. Gluten, 173. Glutinous, 147. Gum, 173. Gynobasis, 102, 127. Gynophore, 102, 127. Hairs, 144, 147, 159. Hastate, 47. Head, 62. Hepta- (7 in composi- tion). Herbarium, 202. Hermaphrodite, 68. Heterogamous, 70. Hexa- (6 in composi- tion). Hilum, 141. Hirsute, 147. Hispid, 147. Hoary, 147. Homogamous, 70. Hooks, 144, 145. Hybemaculum, 17. Hybrids, 210. Hypocrateriform, 90. Hypogynous, 124, 126. Imbricate, ,89. Imperfect, 67. Impervious, 28. Incomplete, 67. Incurved, 119. Indehiscent, *123. Indumentum, 147. Induplicate, 89. (I 2 xxxvi Inferior, 124, 143. Inflorescence, 55, 57. Infundibuliform, 90. Innate, 196. Insertion, 124, 126, Intemode, 28. Introrse, 100. Involucre, 64. Involute, 89. Irregular, 81, 90. Isomerous, 72. Knob, 19. Lamina, 88, 34. Lanceolate, 43. Lateral, 77. Latex, 153. Leaf, 33, 64. Leafbud, 29. Leaflet, 38, 64. Leaves, 33, 64, 169, 185. Lepidote, 148. Limb, 88, 90. Linear, 43. Linear-lanceolate, 44. Lobe, 38, 87. Lobed, 38, 122. Loculicidal, 137. Loculus, 103, 109. Lower, 77. Lyrate, 40. Male, 68. Margin, 34. Mealy, 147. Medullary rays, 167. Medullary sheath, 167. Membranous, 50, 139. Meniscoid, 51. Mesocarp, 134. Micropyle, 117, 141. Midrib, 39. Monadelphous, 95. Monandrous, 94. Moniliform, 51. Mono- (1 in composition). Monocarpellary, 107. Monochlamydeous, 68. Monoecious, 69. Monogynous, 107. Monopetalous, 86. Mouth, 90. Mucronate, 45. Mucronulate, 45. Multi- (many in compo- sition).. INDEX TO TERMS. Muricate, 147. Naked, 68. Natural, 2, 183. Natural Order, 179, 182, 203. Nectary, 123. Nerve, 39. Netted, 39. Neuter, 68. Node, 27, 28. Novem- (9 in composi- tion). Nucleus, 117, 155. Nut, 135. Obcompressed, 51. Obconical, 51. Obcordate, 46. Oblate, 43. Oblong, 43. Obovate, 43. Obovoid, 51. Obpyramidal, 51. Obtuse, 45. Oct- or Octo- (8 in com- position). Offset, 17, 20. Opposite, 30. Orbicular, 43. Order, 202. Organs, 6,* 7, 9. Orthotropous, 119. Oval, 43. Ovary, 103, 105. Ovate, 43. Ovoid, 51. Ovule, 66, 117, 203. Palate, 90. P&lea, palese, 64. Palmate, 40. Palmatifid, 40. Panicle, 62. Paniculate, 62. Papillae, 103. Papyraceous, 139. Parallel, 39, 97* Parenchyma, 153. Parietal, 117. Patelliform, 51. Pectinate, 40. Pedate, 40. Pedatifid, 40. Pedicel, 59. Peduncle, 59. Peltate, 49. Penicillate, 113. Penta- (5 in composi- tion). Perfoliate, 35. Perennial, 16, 18. Perianth, 66, 82. Pericarp, 132. Perigynous, 124, 126. Perisperm, 138. Personate, 90. Pervious, 28. Petal, 74. Petiole, 34. Petiolule, 38. Phaenogamous Plants, 10. Phanerogamous Plants, 10. Pilose, 147. Pinnate, 40. Pinnately - trifoliolate, 40. Pinnatifid, 40. Pistil, 66, 76, 102. Pistillate, 68. Pith, 167, 164. Placenta, 115. Placentation, 115. Plant, 5. Plicate, 89. Plumule, 142. Plurilocular, 109. Podocarp, 102. Pollen, 92. Poly- (many-, in compo- sition). Polyandrous, 94. Polygamous, 69. Polygynous, 107. Polypetalous, 86. Pore, 92, 100. Praefoliation, 52. Preservation of speci- mens, 182, etc. Prickles, 116, 140, 144. Primine, 117. Primordial utricle, 155. Procumbent, 24. Prostrate, 24. Protoplasm, 155. Puberulent, 147. Pubescent, 147. Pulvinate, 121. Punctate, 147. Punctiform, 114. Pungent, 43. Putamen, 134. Pyramidal, 51. NDEX TO TERMS. xxxvii Pyrenes, 134. Pyxis or Pyxidium, 135, 137. Quadri- (4 in composition) . Quinque- (5 in composi- tion) . Race, 177, 178. Raceme, 62. Racemose, 62. Rachis, 62, 38. Radical, 36. Radicle, 142. Rameal, 36. Raphe, 119. Raphides, 156. Ray, 62. Receptacle, 62, 120, 124. Reduplicate, 89. Reniform,- 48. Resupinate, 90. Reticulate, 39. Retuse, 45. Rhizome, 18. Rhomboid, 43. Ribs, 39. Ribbed, 147. Ringent, 90. Root, 12, 160, 185. Rootstock, 18. Rosulate, 37. Rotate, 90. Rotund, round, 43. Rudimentary, 67, 93, 104. Runcinate, 40. Runner, 27. Saccate, 90. Sagittate, 47. Salver-shaped, 90. Samara, 135. Sap, 156, 168. Sapwood, 167. Sarcocarp, 134. Sarmentose, 25. Scale, 64, 148. Scaly, 21, 148. Scape, 60. Scarious (or scariose), 50. Scattered, 30. Scion, 27. Scorpioid, 62. Section, 38, 182. Secund, 30. Secundine, 117. Seed, 10, 138. Segment, 38, 87. Sepal, 72. Septem- (7 in composi- tion). Septum, septa, 109. Septicidal, 137. Serrate, serrulate, 38. Sessile, 35, 88. Septifragal, 137. Serrature, 38. Setae, 98, 147. Setaceous, 51. Setose, 147. Sex - (6 in composition). Sheathing, 35. Simple, 62, 38, 131. Sinuate, sinus, 38. Smooth, 147, Spadix, 62. Spatha, 62. Spatulate, 43. Species, 175, 176, 182, 202, 205. Specimen, 184, 185, 205. Spherical, 51. Spicate, 62. Spike, 62. Spine, spinous, 146. Spiral vessels, 153. Spore, 10. Spurred, 90, 98. Stamens, 66, 75, 91, 94. Staminate, 68. Staminodia, 93. Starch, 156. Stellate, 90, 148. Stem, 23, 163, 185. Stem-clasping, 35. Sterile, 68. Stigma, 104. Stipellse, 54. Stipule, 53. Stock, 16. Stole, or stolon, 17, 27. Stomates, 158. Stonefruit, 134. Striate, 147. Strigulose, 147. Strigose, 147. Strophiole, 140. Style, 103, 104. Subclass, 182, 202. Suhgenus, 182. Suborder, 182. Subsection, 182. I Subtribe, 182. Subulate, 43. Succulent, 33, 50. Sucker, 26. Superior, 124, 143. Symmetrical, 72. Synandrous, 94. Syncarpous, 108. Syngenesious, 95. Tapering, 46. Taproot, 15. Teeth, 87. Tegmen, 139. Tendril, 25, 144, 145. Terete, 51. Temate, 30, 40. Tetra - (4 in composition). Tetradynamous, 95. Testa, 139. Thoms, 144, 146. Throat, 90. Tomentose, 147. Toothed, 38, 122. Torulose, 51. Torus, 62, 120. Tracheae, 153. Tri- (3 in composition). Tribe, 182. Trichotomous, 31, 62. Trifid, 40, 42. Trigonous, 51. Triquetrous, 51. Tristichous, 30. Truncate, 45. Trunk. Tube, 87, 90. Tuber, 15, 19. Tuberculate, 147. Tuberous, 15. Tubular, 90. Tunicated, 21. Twine, 25. Twisted, 89. Two-lipped, 90. Typical, 179. Umbel, 62. Umbellate, 31, 62. Undulate, 38. Uni- (1 in composition). Unijugate, 40, 42. Unilocular, 109. Unisexual, 69. Unsymmetrical, 80. Upper, 77, 171. Urceolate, 90. Utricle, 135. XXXV111 INDEX TO TERMS. Valvate, 89. Valves, 100. Variety, 122, 177, 178. Vascular tissues, 8, 153. Vegetable Anatomy, 150. Vegeatble Physiology, 150. Vein, 39. Velutinous, 147. Ventral, 135. Vernation, 52. Versatile, 96. Verticillate, 30. Viscid, viscous, 147. Vittae or vittas, 149. Voluble, 25. Warted, 147. Wavy, 38. Whorled, 30. Wing, 35, 139. Woody tissue, 8 Wool, 147. Woolly, 147. , 153. XXXIX TABLE OF THE CLASSES, OEDEES, ETC., OF THE SOUTH AFEICAN FLOEA. In the following pages I have endeavoured to group the Natural Orders of Sooth African plants in such a manner as to afford some indication of the principles according to which they have been brought into the sequence adopted in this work by Dr. Harvey. This sequence is in the main that pro- posed by Jussieu, and carried out by De Candolle, and most subsequent authorities. In so far as the limitation and order of the Classes and Subclasses and of the Cohorts and Orders of Monocotyledons and Acotyledons is concerned, it is no doubt a very natural system; but this is not so with the Orders of Angiospermous Dicotyledons, the arrangement of which is very artificial. The principle upon which De Candolle arranged the latter Orders involved two assumptions : one, that plants with their floral whorls complete, and each whorl regular and composed of separate parts (as Polypetalece Thalamiflorce) , were more highly organized than those with fewer floral whorls, and these irregular, and their constituent parts combined (as in Monopetalece , etc.) ; — the other that the presence of but one whorl in the perianth, or of no perianth, indicated that such Orders should be kept apart from the rest. Advanced know- ledge has, however, carried conviction to many minds, that Dicotyledonous plants with combined organs are really more highly organized than those with these parts free ; that irre- gularity of flower prevails in the highest organized groups, and that the majority of the Orders with reduced floral enve- lopes are really members of other Orders whose prevailing features are of a complex and high type. The fact is, that the Dicotyledonous Orders cannot be ar- ranged in a linear series, — but as descriptions and arranged collections of them must follow a linear series, the Candollean is adopted for its facility, and because none better (though several others as good) has been proposed. It further pos- sesses this advantage, that most of the Orders of the highest xl TABLE OF THE CLASSES, ORDERS, ETC., types (as Leguminosce , Composite and other Monopetalce) stand towards the middle of the series, and the Orders of a lower type occupy the ends ; thus the complete-flowered plants with much separated organs are those with which the series begins, and the incomplete-flowered Orders (some of which have no recognized affinity with higher ones) occupy the end of the series. The extent to which this system is artificial is best illus- trated by a few examples, thus : — 10, Bixacece, passes into 58, Passiflorece, various genera ac- tually uniting the tw~o. 16 and 17, Caryophyllece , passes into 62, Ficoidece , 99, Phyto- laccece , 101, Amaranthacece , and 101, Paronychiece. 22, Malvacece , passes into 109, Fuphorbiacece. 34, Olacinece , passes into 65, Cornea , 66, Lorantliacece , and 108, Santalacece ; and many other cases might be quoted of Orders removed to three far distant groups by one or two characters alone, and these very inconstant ones. If it is asked, why then not bring all together ? The auswer is, if we do, — 1, we should lose all means of finding the locality of any genus, without hunting through every group ; and 2, we must often then intercalate between two Orders that are most closely re- lated, another Order containing a vast number of plants not so much related to either Order as these two Orders are to one another. Tor instance, if we bring Euphorbiacece in between Malvacece and Tiliacece , we break up the character of the class, subclass, group, series, and cohort under which Malvacece and Tiliacece are classed, and thrust between these a vast host of Euphorbiaceous genera that are not so nearly related to Mal- vacece as Tiliacece are. It must then be clearly understood, that the Natural Orders established throughout the vegetable kingdom are in the main perfectly natural groups, but that the so-called Natural sys- tem of plants is by no means a natural one, in respect of the sequence of the Dicotyledonous Orders, and that this is be- cause they cannot be arranged naturally in a linear series. To classify these Orders arbitrary characters have been sought and used, which bring a larger proportion of them into proper position and sequence, but remove others very far from their proper places. — J. D. Hooker. CONSPECTUS OF THE CLASSES, COHOETS, ETC. Class I. DICOTYLEDONES. — Stem, when perennial, with pith concentric layers of wood and bark. Leaves usually with branched and netted venation. Perianth usually of 4 or 5 OF THE SOUTH AFRICAN FLORA. xli parts in a whorl, or multiples of 4 or 5. Embryo with 2 co- tyledons. In germination the radicle lengthens and branches. Subclass I. ANGtO&PERMEiE. Order 1-116. Group I. Polypetalese. Series 1. Thalamiflora:. Cohort 1. Ranales .... . . . Orders 1-4 99 2. Parietales . . . ... „ 5-12 99 3. Polygaline® . ... „ 13-14 99 4. Cary ophy lline® . ... „ 15-18 99 5. Guttiferales . . ... „ 19-21 99 6. Malvales . . . ... „ 22-24 Series 2. Disciflora. Cohort 7. Geraniales . . 8. Olacinales . . . . . „ 34-35 99 9. Celastrales . . . . . „ 36-38 99 10. Sapindales . . . . . „ 39-40 Series 3. Calyciflora. Cohort 11. Rosales . . . . . . Orders 41-49 99 12. Myrtales . . . . . „ 50-55 99 13. Passiflorales . . . „ 56-60 99 14. Ficoidales . . . . . „ 61-62 99 15. Umbellales . . . . . „ 63-66 Group II. Monopetalese. Series 1. Ovary inferior. Cohort 16. Rubiales Orders 67-68 99 17. Compositales . * * • 99 69-70 99 18. Campanales 71 Series 2. Ovary superior. Cohort 19. Ericales . . . . . . Orders i 72 99 20. Jasminales . . . . „ 73-74 99 21. Primulales . . . . . „ 75-76 99 22. Plantaginales . • * • 99 77 99 23. Sapotales . . • 99 78-79 99 24. Gentianales • • • 80-83 99 25. Convolvulales . • • • 33 84-86 99 26. Personales . . 87-92 99 27. Verbenales . . • * • 99 93-96 Group III. IMIonochlamydeEe. Cohort 28. Nyctaginales . . . Orders 97 „ 29. Chenopo'diales . . . . „ 98-102 ,, 30. Pen®ales '. . • • • „ 103-104 „ 31. Laurales . . . . . „ 105-107 „ 32. Santalales . . . . . „ 108 „ 33. Euphorbiales . . . . „ 109 „ 34. Urticales . . . . . „ 110 xlii TABLE OF THE CLASSES, ORDERS, ETC., Group IV. Achlamydese. Cohort 35. Amentales Orders 111-113 „ 36. Rhizanthales .... „ 114 ,, 37. Piperales ,, 115 Anomalous Order of Dicotyledons. 116. Podostemaceae. Subclass II. GYMN OSPERMEJE. Orders 117-119. Class II. MONO COTYLEDONES.— Stem, when peren- nial, without separable bark, wood, and pith. Veins of the leaf usually parallel, and if netted the veinlets are transverse and parallel. Perianth, when present, 3-6-merous. Stamens 3-6. Embryo with one cotyledon ; the plumule lies in a cavity at its side. Radicle rarely elongating and branching. Group I. Petaloidese. Subclass I. Ovary inferior. Cohort 1. Hydrales Orders 120 „ 2. Amomales „ 121-122 „ 3. Iridales „ 123-127 Subclass II. Ovary superior. Cohort 4. Alisraales „ 5. Arales „ 6. Palmales „ 7. Liliales „ 8. Commelynales . . . „ 9. Restiales . . . . . Orders 128-129 „ 130-131 „ 132 „ 133-136 „ 137-138 „ 139-140 Group II. Glumacese. Cohort 10. Glumales Orders 141-142 Class III. ACOTYLEDONES, or CRYPTO Gr AMiE. — Plants cellular or vascular, without true stamens, pistil, or ovules. Organs of fructification usually very minute, giving origin to microscopic spores, by which the species are propa- gated. Spores germinating by a prothallium, or by micro- scopic threads. Subclass I. ACROGENS. Cohort 1. Filicales Orders 143-146 „ 2. Muscales „ 147-151 Subclass II. THALLOGENS. Orders 152-154. OF TilE SOUTH AFRICAN FLORA. xliii CONSPECTUS OE THE OBDEES. Class I. DICOTYLEDONES. Subclass I. Angiospermeas. — Ovules enclosed in an ovary, and seeds in a seed-vessel. Group I. POLYPETALEiE. Perianth consisting of both calyx and corolla, the latter of distinct petals. (The exceptions are very numerous ; for in all the large Orders, genera occur that want calyx or corolla, or both, and in some the petals cohere or are united into a monopetalous corolla.) Series I. Thalamiflorse. — Sepals distinct, usually herbaceous, inferior. Stamens inserted usually immediately under the ovary, rarely on a lobed or expanded disk, free from the calyx, very rarely on the base of the petals. Ovary very rarely inferior. Cohort I. Ranales. — Stamens very numerous (few, and opposite the petals in Menispermacece) . Carpels distinct or immersed in a fleshy torus in Nymphceacece. Embryo small, in fleshy or mealy albumen. — Herbs, rarely shrubs or trees. Order 1. JR anunculacece (p. 1). All herbaceous but Clematis. ” 3 usually 3- or 6-merous. „ 4. Nymphceacece (p. 4). All aquatics. Cohort II. Parietales. — Stamens definite or indefinite. Ovary 1-celled, with parietal placentation, rarely (as in Crucifer a) divided into 2 cells by a membranous expansion of the placenta. Order 5. PapaveraceafaS). ) These orfer8 are usuaI1 united> „ 6. Fumounaceee (p. 5). ) J „ 7. Cruciferce (p. 6). Herbs, all with alternate exstipulate leaves. „ 8. Capparidece (p. 11). „ 9. Resedacece (p. 13). „ 10. Bixacea (p. 13). Has equal affinity with Cohort 13, Passifiorales. „ 11. Violariece (p. 17). ,, 12. JDroseracece (p. 17). More closely allied to 44, Saxifragece. Cohort III. PoLYGALiNEiE. — Flowers regular or irregular. Sepals and petals each 5, rarely 3 or 4. Stamens 5 or 10. Ovary 2 -celled, usually of 2 carpels ; ovules numerous, horizontal or pendulous and solitary. Albumen fleshy, rarely absent. Order 13. Poly galea (p. 18). Flowers very irregular. „ 14. Pittosporece (p. 19). Flowers regular. Cohort IV. Caryophyllineje. — Flowers regular. Sepals 2-5, free, rarely united. Petals as many, connate at the base in some Portulacece. Sta- mens as many as the petals, alternate with them. Ovary 1-celled ; ovules attached to a free basal placenta. Embryo usually curved round mealy albumen. — Chiefly herbs. Order 15. FranJceniacece (p. 20). Embryo straight. „ 16. Caryophyllaceee (p. 20). xliv TABLE OF THE CLASSES, ORDERS, ETC., Order 17. Portulacece (p. 23). Ovary inferior in Portulaca. „ 18. Tamariscinecp, (p. 25). Albumen 0 in Tamarix. Cohort V. Guttiferales. — Flowers regular. Sepals and petals 4 or 5, the former imbricate. Stamens usually indefinite. Ovary 3-many- celled, ovules attached to the inner angles of the cells. — Leaves usually opposite and exstipulate. Order 19. Elatinacece (p. 25). „ 20. Hypericinea (p. 25). „ 21. Guttiferce (p. 26). Cohort YI. Malvales. — Flowers regular. Sepals or calyx-lobes 4-5, val- vate. Petals 4-5, twisted in bud. Stamens usually very numerous, free or monadelphous. Ovary 3-many-celled, or of 3 or many free carpels ; ovules attached to the inner angles of the cells. Leaves stipulate. Order 22. Malvaceae (p. 27). Stamens united to base of connate petals. Anthers 1-celled, opening outwards. „ 23. S 'terculiacece (p. 30). Anthers 2-celled, opening outwards. Flowers 5-7-merous, unisexual and apetalous in Sterculia. „ 24. Tiliacece (p. 32) . Inner bark very tough, often used as cordage. Disk sometimes conspicuous. Series II. Disciflorese. — Sepals distinct, rarely connate, imbricate, rarely valvate, or adnate with the ovary. Disk usually very evident ; ex- panded into a torus or a ring, or conspicuous lobes or glands. Petals inserted at the base of the disk. Stamens usually definite, inserted at the base of or upon or within or between the lobes of the disk. Ovary free, or immersed in the disk, rarely inferior. Cohort VII. GrERANlALES. — Flowers regular or irregular. Sepals and pe- tals 3-5. Disk annular, within the stamens, or of glands alternating with the petals. Stamens usually definite. Carpels free or combined ; ovules 1-2, attached to the inner angle of the cells or carpels, pendulous, with the raphe towards the axis. Order 25. yy yy yy 26. 27. 28. 29. 30. jj 31. 32. 33. Linece (p. 34). Disk 0 in Erythroxylon. Malpighiacece (p. 35). Calyx-lobes often with glands at back. Zygophylleae, (p. 36). Petals twisted, 0 in Augea and Seetzenia. Geraniacece (p. 38). Flowers often irregular. Disk 0 in Im- patiens. Rutacece (p. 41). Leaves with pellucid dots. Disk and petals 0 in Empleurum. Ochnaceae (p. 46). Sepals persistent. Disk enlarged after flowering. Rurseracece { p. 47). Flowers usually polygamous. Disk usually cup-shaped. Meliacece (p. 47). Stamens usually monadelphous. Chailletiacece (p. 49). Petals 2-lobed. Cohort VIII. Olacinales. — Flowers regular, hermaphrodite or unisexual. Calyx small. Petals, if present, often valvate. Disk cup-shaped or glandular. Stamens definite. Ovary 1-celled, with 1-3 ovules pendu- lous from a central placenta, or 2- or more celled, with 1-2 ovules pen- dulous from the top of each cell ; ovules with the raphe turned away from the axis. — Leaves simple, usually exstipulate. OF THE SOUTH AFRICAN FLORA. xlv Order 34. Olacinece (p. 49). Disk annular or 4-5-lobed. „ 35. Ilicinece (p. 50). Disk 0. Cohort IX. Celastrales. — Flowers regular or irregular, always small and usually hermaphrodite. Sepals and petals 4-5. Disk cushion-like and adnate to the base of. the calyx. Stamens definite on the margin or base of the disk. Ovary 2- or more celled, free or immersed in the disk ; ovules 1-2, erect from the base of the cells, with the raphe next the axis, rarely 3-6, attached to the inner angle of the cells. — Leaves simple, except in Ampelidece. Order 36. Celastrinece (p. 51) . Flowers small. Ovary sometimes sunk in disk. Ovules 6-8 in Cathastrum and Putterlichia. „ 37. Rhcimnece (p. 55). Ovary often inferior or adhering to the calyx. Petals often absent, always minute. „ 38. Amjpelidece (p. 57). Petals valvate, often cohering at the tips. Cohort X. Sapindales. — Flowers regular or irregular, usually unisexual. Disk adnate to the base of the calyx. Stamens definite or indefinite, in- serted upon or within the disk. Ovary 1- or more celled ; ovules 1-2 in each cell, erect from its base, or pendulous from a basal funicle. — Leaves often compound. Order 39. Saypindacece (p. 58). Petals usually 1 fewer than the calyx-lobes, or 0, often with a claw at the base. Stamens often 8, and disk often unilateral. „ 40. Jnacardiacece (p. 62). Petals 0 or as many as calyx-lobes. Stamens usually twice as many. Series III. Calycifiorse. — Sepals connate into a more or less distinct tube, which is free or adnate to the ovary. Disk usually indistinguishable from the calyx-tube. Petals as many as the sepals, inserted with the sta- mens on the calyx-tube or on the base of its lobes. Ovary very often more or less inferior. Cohort XI. Fosales. — Flowers regular or irregular, usually 5-merous and hermaphrodite. Stamens definite or indefinite. Carpels 1 or more, free, rarely united more or less ; styles usually distinct, or separable. — Leaves simple or compound. Order 41. Connaracece (p. 65). „ 42. Leguminosce, (p. 65). Stamens indefinite in Mimosece. „ 43. Rosacece (p. 93.) Petals 0 in Alchemilla , Poterium , and Cliffortia. „ 44. Saxifrages (p. 97). Disk large in Brexia. „ 45. Crassulacece (p. 99). Perigynous scales opposite the carpels in many species. „ 46. Hamamelides (p. 102). Flowers unisexual and apetalous in Tricliocladus. Grubbia should perhaps be referred to Santa - ' lacece. „ 47. Bruniaces (p. 103). Petals connate in Lonchosioma. „ 48. Haloragece (p. 106). Flowers very imperfect in all the species. „ 49. Balanophores (p. 107). Flowers extremely imperfect. The Order has been placed next to Santalacece recently by Dr. Eichler, of Munich. Cohort XII. Myrtales. — Flowers usually both regular and hermaphrodite. Stamens definite or indefinite, 4-5-merous. Disk 0 or covering the ovary. xlvi TABLE OE THE CLASSES, ORDERS, ETC., Ovary inferior ; style very rarely divided. — Leaves simple, and usually entire. Order 50. J Rhizophoreoe (p. 108). Ovary free in Weihea. „ 51. Combretacece (p. 109). Flowers polygamous and apetalous in Terminalia. „ 52. Myrtacece (p. 110). „ 53. Melastomacece (p. 112). Anthers often of two forms and colours in each flower. „ 54. Lyihrariece (p. 114). Heteropyxis has stamens opposite the petals, and pellucid dotted leaves as in Myrsinece. „ 55. Onagrarieee (p. 116). Montinia has unisexual flowers and parietal placentas. Cohort XIII. Passiflorales. — Flowers regular, unisexual or hermaphro- dite. Calyx free or connate with the ovary. Stamens various. Ovary 1- celled, with parietal placentae. Order 56. Turner acece, (p. 119). „ 57. Loasacece (p. 119). Ovary 3-celled in the only Cape genus, with cells 1-ovuled. „ 58. Passiflorece (p. 120). „ 59. Cucurbitacece (p. 122). „ 60. Begoniacece (p. 128). Ovary 2-3-celled. Cohort XIV. Ficoidales. — Flowers hermaphrodite, usually regular. Calyx free or adnate to the ovary. Stamens numerous, rarely few. Ovary 1- celled, with parietal placentae, or 2- or more celled, with basilar ovules. Embryo usually curved. — Leaves quite entire or 0. Order 61. Cactece (p. 129). Succulent, leafless, spinous plants. „ 62. Ficoidece (p. 129). Petals often 0. Stamens sometimes hypo- gynous. (Closely related to Caryophyllece and Phytolacceee.) Cohort XV. Umbellales. — Flowers small. Calyx adnate to the ovary. Stamens as many as the petals. Disk crowning the ovary. Ovaryin- ferior, 1-2- or more celled ; styles usually free ; ovules 1 in each cell, rarely 2, pendulous. Ripe carpels indehiscent. Embryo minute, in copious albumen. — Leaves often compound. Order 63. TJmbelliferce (p. 135). „ 64. Araliacece (p. 146). „ 65. Corneee (p. 147). „ 66. Loranthacece (p. 148). Corolla monopetalous in Loranihus. Flowers unisexual in Viscwm. (This Order should be placed next to Santalacece.) Group II. MON OPETALEiE. Perianth consisting of both calyx and corolla j the petals of the latter combined. Series I. Ovary inferior. Cohort XVI. Rubiales. — Stamens inserted on the corolla-tube. Ovary 2- or more celled ; style usually simple or 2-fid at the apex only ; ovules 1 or more in each cell. — Leaves very rarely toothed or cut. Order 67. Rubiacece (p. 148). „ 68. Valerianece (p. 157). Fruit 1-celled in Valeriana. or THE SOUTH AFRICAN FLORA. xlvii Cohort XVII. Compositales. — Stamens inserted on the corolla- tube. Ovary 1-celled ; style simple or 2-fid at the apex ; ovules solitary. — Leaves simple or compound. Order 69. Dipsacece (p. 158). Corolla imbricate. Stamens free. „ 70. Composites (p. 158). Corolla valvate. Anthers united. Cohort XVIII. Campanales. — Stamens usually epigynous. Ovary 2-10- celled ; style simple ; ovules usually numerous. — Herbs, rarely shrubs j leaves simple. Order 71. Lobeliacece (p. 209). Petals sometimes free in CypMa. Ovules solitary in each cell in Sccevola . Stamens on the corolla in Phigiophyllum. Series II. Ovary superior. Cohort XIX. Ericales. — Flowers regular, hermaphrodite. Stamens as many as corolla-lobes, hypogynous or epigynous, rarely on the corolla, all equal. Disk usually evident. Ovary 3-more-celled ; style and stigma simple ; ovules numerous on the inner angle of the cells. — Usually shrubs or trees ; leaves alternate, exstipulate, simple. Order 72. Ericaceae (p. 215). Cohort XX. Jasmin ales. — Flowers often unisexual, regular. Petals some- times free. Stamens 2 or more, inserted on the corolla, alternate with its lobes. Disk 0. Ovary 1-2-celled ; style simple or 2-fid ; ovules 1-2 in each cell. — Shrubs or trees ; leaves opposite, simple or compound. Order 73. Jasminece (p. 219). „ 74. Salvador acece (p. 220). Petals free. Cohort XXI. Primtjlales. — Flowers usually hermaphrodite, regular. Petals very rarely free. Stamens 4-5, all opposite the corolla-lobes, small. Ovary 1-celled, with free central placenta. — Herbs shrubs or trees ; leaves simple, alternate, exstipulate. Order 75. Myrsinece (p. 220). Ovary inferior in Mcesa. Petals free in Embelia. „ 76. Primulacece (p. 221). Ovary half inferior in Samolus. (Order 94. Plumbaginece , should come here.) Cohort XXII. Plantagunales. — Flowers hermaphrodite, regular. Corolla persistent. Stamens alternate with the corolla-lobes, filaments long and pendulous ; anthers versatile. Ovary 2-4-celled ; style simple ; stigma filiform, hispid. Fruit dehiscing transversely. — Herbs. Order 77. Plantaginece (p. 223). Cohort XXIII. Sapotales. — Flowers often hermaphrodite, regular. Co- rolla 4-24-lobed. Stamens (with alternating staminodes) solitary or in fascicles opposite the corolla-lobes. Ovary 2-12-celledj cells 1-2-ovuled. Stigma as many lobed as cells. — Trees or shrubs. Order 78. Sapotacece (p. 223) . ,, 79. Ebenacece (p. 224). Cohort XXIV. Gentianales. — Flowers hermaphrodite, regular. Corolla 4-5-lobed. Stamens 4-5, alternate with the corolla-lobes. Ovary 2- xlviii TABLE OF THE CLASSES, ORDERS, ETC., celled, with usually numerous ovules in each cell ; stigma simple or 2- lobed. Fruit a capsule or berry. — Leaves opposite, usually without stipules and quite entire. Order 80. Asclepiadece (p. 226). Pollen collected in masses like those of Qrchidece. „ 81. Apocynece (p. 244). „ 82. Loganiacece (p. 248). Leaves often stipulate. „ 83. Gentianece (p. 250). Leaves alternate in tribe Meny anthem. Cohort XXY. Convolvulales. — Flowers hermaphrodite, regular. Corolla 4-10- usually 5-parted ; lobes plaited in aestivation. Stamens as many as and alternate with corolla-lobes. Ovary on a disk, 2-4-celled ; ovules definite or indefinite ; stigma simple or 2-lobed. — Leaves almost always alternate. Order 84. Convolvulacece (p. 253). Carpels separate in Falkia and Di- chondra. „ ? 85. Hydrophyllaceoe (p. 256). „ 86. Solanaceee (p. 256). Cohort XXVI. Personales. — Flowers hermaphrodite, irregular. Corolla often 2-lipped. Stamens usually fewer than the corolla-lobes, and un- equal in length. Ovary on a disk, 2-celled (rarely 1-celled), many-ovuled ; style usually slender ; stigma simple or 2-fid. — Leaves almost always opposite, exstipulate. Order 87. Scrophulariacece (p. 259). „ 88. Lentibularinece (p. 273) Ovary 1-celled, with free central placenta. „ 89. Orobaneheee (p. 274). Leaves 0, or reduced to alternate scales. „ 90. Bignoniaceee (p. 274). ,, 91. Gesneriacem (p. 276). Leaves often alternate. „ 92. Acanthacece (p. 279). Cohort XXVII. VerbenalRS.— Flowers irregular, rarely regular, herma- phrodite. Corolla usually unequally 2-lipped. Stamens usually fewer than the corolla-lobes, often unequal in length. Ovary 2-4-celled ; cells 1-ovuled ; style usually filiform ; stigma simple or 2-lobed. Fruit a 2-4- celled berry or drupe of 4 little nuts. — Leaves opposite or alternate. Order 93. Verbenacece (p. 287). „ 94.# Plumbagineee (p. 295). Flowers regular. Corolla often of 5 petals, with as many opposite stamens. Ovary 1-celled, with 3-5 styles. „ 95. Poraginece (p. 296). „ 96. Labiates (p. 301). Group III. MONOCHLAMYDEiE. Perianth consisting of one series, there being no distinct mss., n. sp.) species, dispersed through the colony. 5. RANUNCULUS, Hall. Sepals 3-5, green or yellowish, imbricate, falling. Petals 5-10, flat, yellow or white, with a minute fleshy scale or pit near the base on the inside. Carpels many, tufted, 1-seeded, dry in fruit, pointed or beaked. — FI. Cap. i. p. 5. Herbs. Stems weak, leafy. Leaves stalked, deeply cut, lobed or mul- tifid in our species. — 6 South African species, dispersed: 5 with yellow flowers are terrestrial ; 1 {FL. aquatilis ), with white flowers, grows in ponds and rivers. Order II. ANONACEiE. Plowers bisexual. Sepals 3, valvate. Petals 6, in two rows. Stamens many, hypogynous ; filaments thickened up- wards ; anthers fixed. Carpels several, separate or cohering. — Trees or shrubs. Leaves simple, entire, alternate, without stipules. Plowers leathery, lateral, on short stalks. Carpels separate, on short pedicels. Petals imbricate. Ovules many 1. Uvaria. Petals valvate. Ovules 1-2, erect 2. Popowia. Carpels confluent into a many-seeded fruit 3. Anona. 1. UVARIA, Linn. Petals 6, imbricate in two rows, plano-convex. Stamens very many, compressed, with a prolonged connective. Torus little-raised, hairy. Carpels oblong, furrowed on the inner face ; style continuous ; ovules many, in two rows. Berries many- or 1-seeded. — FI. Cap. i. p. 8. II. ANONACEiE. 3 1 South African species : U. Caffra , E. Mey., from Natal ; a climbing shrub, 'with laurel-like leaves, 2-5 inches long, l-2£ inches wide. Flowers solitary, lateral, nodding. Petals downy. Berry as large as a cherry. 2. POPOWIA, Endl. Petals 6, valvate in two rows, short, broadly ovate, concave, not spreading. Stamens many, wedge-shaped, with a pro- longed connective. Torus little raised. Carpels 5 or many ; style capitate ; ovules 1-2, erect. Berries globose or egg- shaped, stalked. — Benth. and Book . Gen. Bl. i. p. 25. Guat- teria, FI. Cap. i. p. 9. P. Caffra , H. and S. ( Cuatteria Caffra, Sond. !), from Natal, is a climb- ing shrub, 10-20 feet high, hanging in festoons from the branches of trees. Leaves oblong, 3-4 inches long, pale or livid beneath. Flowers 2-4 toge- ther, on short stalks, cream-coloured. 3. ANONA, Linn. Petals mostly 6, valvate in two rows, the outer fleshy, con- cave, scarcely spreading, inner smaller, rarely wanting. Sta- mens many, with a prolonged ovate connective. Torus hemi- spherical. Carpels many, mostly connate ; style oblong ; ovules solitary. Berries fleshy, obtuse, confluent into a many- celled fruit. — Benth. and Hook. Gen. PI. i. p. 27. A. Senegalensis , Pers., a shrub 6-12 feet high, is found near Natal. Leaves ovate, feather-nerved and netted-veined beneath, pubescent. Fruit edible, 1^-2 inches diameter, ‘‘well-flavoured” (W. T. Gerrard). Order III. MENISPERMACEiE. Flowers minute, unisexual, green. Stamens definite (few), mostly monadelphous ; filaments short ; anthers fixed. Car- pels 1-3, separate ; ovules solitary. Fruit of fleshy, oblique, very small drupes. — Slender climbing half-shrubs, with alter- nate, simple, netted-veined leaves. Flowers in axillary cymes racemes or umbels. Female flowers with 3-5 petals and 3-5 sepals ; style 3-fid 1. Stephania. Female flowers with 1 petal and 1 sepal ; style 3-fid . 2. Cissampelos. Female flowers with 2 petals and 2 sepals ; style none . 3. Antizoma. 1. STEPHANIA, Lour. Flowers dioecious. Male : Sepals 6-10, in two rows. Pe- tals 3-5, shorter than the sepals, obovate, rather fleshy. Sta- minal column peltate at the apex ; anthers sessile, in a mar- ginal ring. Female : Sepals 3-5. Petals as in the male flower. Carpel 1 ; style 3-parted. Drupe compressed, dor- 4 III. MENISPERMACEiE. sally tubercled, hollowed on both sides. — BentJi. and Hook. Gen. PI. p. 37. Homocnemia, FI. Cap. i. p. 10. A single species, Homocnemia Meyeriana , Miers, from Natal. A vine- like twiner, with peltate, ovate-orbicular, many-nerved, pubescent leaves. Flowers in umbels. 2. CISSAMPELOS, Linn. Flowers dioecious. Male : Sepals 4, separate. Corolla cup-shaped, nearly entire, shorter than the sepals (composed of 4 confluent petals). Staminal column peltate at the apex ; anthers 4-12, sessile in a marginal ring. Female : Sepal 1, anterior. Petal 1 (or 2 confluent) in front of the sepal, clasp- ing the ovary. Carpel 1 ; style 3-fid. Drupe kidney-shaped ; nut compressed and wrinkled at edges. — FI. Cap. i. p. 10. Mostly twining, slender, shrubby plants. Leaves simple, petioled, en- tire, ovate reniform or cordate, often peltate. Male flower in cymes ; female racemose, densely tufted in the axils of leafy bracts. — 3 Cape spe- cies, 2 of which are confined to the Eastern district and Natal. 3. ANTIZOMA, Miers. Flowers dioecious. Male flower as in Cissampelos. Female : Sepals 2, opposite, very concave, fleshy, slightly imbricate in bud. Petals 2, opposite the sepals, minute, scale-like, orbi- cular, fleshy, hyp ogy nous. Carpel 1, ob ovate ; style none ; stigma obsolete or bluntly 2-lobed. Fruit unknown. — FI. Cap. i. p. 11. Endemic. Suberect or twining shrubs. Leaves linear or lanceolate, en- tire, opaque, leathery, on very short petioles ; the petiole armed at base with a dorsal spur-like spine. — 5 reputed species (probably over-estimated), from the Northern and North-Eastern frontiers. Order IY. NYMPHiEACEiE. Flowers bisexual, large and showy. Petals numerous, in several rows. Stamens many ; filaments flat ; anthers adnate. Carpels numerous, sunk in a fleshy torus, and thus concreted into a many-celled ovary. — “ Water-lilies.” Leaves on long stalks, cordate or peltate. 1. NYMPH1EA, Linn. Sepals 4, at the base of the fleshy torus, in which the carpels are sunk. Petals and stamens numerous, in several rows, co- vering the sides of the torus. Ovary many-celled ; stigmas sessile, radiating. Berry leather-coated, irregularly bursting ; seeds many, lodged in pulp. Water-plants, with submerged prostrate rootstocks, throwing up leaves IY. NYMPHiEACEiE. 5 and flower3. Leaves on long, terete petioles, cordate. Flowers on simple peduncles. — 1 South African species, N. stellata , with blue or rarely white, sweet-scented flowers, standing out of the water. It is found in rivers and ponds throughout South Africa. Order Y. PAPAVERACE.®. Blowers bisexual. Sepals 2-3, deciduous. Petals 4-6, equal, spreading, crumpled in the bud. Stamens many ; fila- ments slender ; anthers adnate. Ovary 1-celled, with 2 or se- veral parietal placentae ; ovules many. Capsule many-seeded. - — The Poppy family. Herbs, with coloured, narcotic juices. 1. PAP AVER, Linn. Ovary globose or obovoid, crowned with 4-20 radiating, li- near, sessile stigmas. Capsule oblong, dry, opening by small pores under the stigmas ; placentae projecting into the cavity, and dividing it into several incomplete chambers. Seeds very numerous. — FI. Cap. i. p. 15. 1 South African species, P. aculeatum , Th. ; the Wild Poppy. Radical leaves many, sinuous or pinnatifid, setose and bristly ; stem-leaves sessile, very hispid. Flowers scarlet-orange, on long peduncles. — Native of the Nor- thern and Eastern districts. P. horridum , DC., from Australia, is a syno- nym of this species. Order VI. EXJMARIACEiE. Plowers bisexual. Sepals 2, minute, scale-like. Petals 4, unequal, connivent in pairs, one or both of the outer pair spurred or saccate at base. Stamens 6, diadelphous, 3 in each parcel. — Grlabrous, often glaucous, twining herbs, with much- divided leaves, and small, purple white or yellow, racemose flowers. Fruit many-seeded, capsular. Capsule bladder. like, subglobose 1. Cysticapnos. Capsule lanceolate, compressed 2. Corydalis. Fruit 1 -seeded, indehiscent. Fruit flattened, orbicular 3. DiscocapnoS. Fruit subglobose, not flattened 4. Ftjmaria. 1. CYSTICAPNOS, Boerh. Capsule 2-valved, bladdery ; epicarp inflated, spongy within ; endocarp delicately membranous, supported by slender fila- ments in the centre of the cavity, and. bearing at the margins of the valves many-seeded placentae. Seeds compressed, beaked. — FI. Cap. i. p. 16. 6 VI. EUMARIACEiE. Endemic. C. Africana , the only species, is frequent near Capetown; also at Stellenbosch and Swellendam. Its bladdery capsules are 1 inch in diameter, pointed, and splitting into two boat-like valves. 2. CORYDALIS, DC. Capsule pod-shaped, compressed, 1-celled, 2-valved ; valves separating from a persistent, placentiferous replum (or frame) . Seeds lenticular, beaked. — FI. Cap. i. p. 16. 3 South African species, all annuals, with climbing stems and multifid leaves. 3. DISCOCAPNOS, Ch. and Schl. Fruit (a utricle) orbicular, flattened, membranous, with a marginal wing, indehiscent, 1-seeded. Seed lenticular, beaked. — FI. Cap. i. p. 18 ; Thes. Cap. t. 10. Endemic. D. Mundtii, the only species, is found near Capetown, etc. It is like the common “ Fumitory,” but differs in its flattened fruit. 4. FUMARIA, Linn. Fruit subglobose (not flattened), dry, indehiscent, 1-seeded. F. officinalis , L., the common “Fumitory,” is a weed of cultivation, in- troduced from Europe. Grows in waste places and old gardens. Order VII. CRUCIFERS. Flowers bisexual. Sepals 4, deciduous. Petals 4, clawed, mostly equal, spreading in the form of a cross. Stamens 6, 4 long and 2 short ; anthers versatile. Ovary 2-celled, with pa- rietal placentae. Fruit a long pod ( siliqua ) or a short one (silicula). Seeds without albumen. — Herbs or shrublets, with alternate leaves and no stipules. Flowers in racemes, white yellow purple or sky-blue. — Many garden vegetables, such as Cabbage, Turnips, Mustard, Cress, Radishes, Horseradish etc., and garden flowers, such as Stocks, Wallflowers, etc., belong to this Order. A. Siliqttosa:. Fruit-pod several times as long as broad. * Pod with flat or flattish valves. Calyx equal at base (not spurred). Seeds with flat, accumbent cotyledons (0 = ). Pod linear-elongate, narrow ; valves not elastic 4. Arabis. Pod acute at each end ; valves bursting with elasticity 5. Cardamine. Seeds with long, twice-folded cotyledons (0 1| || ||). Pod linear or beaded 13. Heliophila. Calyx 2-spurred at base 12. Chamira. ** Pod 2-edged or 4-angled, with keeled valves . . 3. Barbarea. VII. CRUCIFERS. 7 *** Pod with convex, round-backed valves. Pod dehiscent when ripe. Seeds with flat accumbent cotyledons (0 = ). Sepals elongate, erect, saccate at base . 1. Matthiola. Sepals spreading, equal at base ... 2. Nasturtium. Seeds with flat incumbent cotyledons (0 ||) 7. Sisymbrium. Seeds with broad cotyledons folded over the radicle (0 >>) 11. Brassica. Pod indehiscent ; seeds as in Heliophila . 14. Carponema. B. Siliculos.®. Fruit-pod short, few- or 1-seeded, less than thrice as long as broad. * Pod dehiscent. Pod with flattish valves and a broad septum. Cotyledons accumbent (0 = ) . . . . 6. Alyssum. Cotyledons elongate, twice folded on the radicle (0 || || ||) 13. Heliophila. Pod with keeled valves and a narrow septum. Seeds 1 in each cell 9. Lepidium. Seeds several in each cell 10. Capsella. ** Pod indehiscent ; seeds solitary. Pod deeply 2-lobed or 2-parted. Petals minute, shorter than the sepals . 8. Senebiera. Petals much longer than the sepals . .15. Brachycarpea. Pod orbicular or ovate. Pod ovate, beaked, with convex, wrinkled valves 16. Cycloptychis. Pod orbicular, with flat valves . . .17. Palmstruckia. 1. MATTHIOLA, E. Br. Sepals erect, the 2 lateral saccate at base. Pod subterete, elongate, with round-backed valves. Stigma thickened, its lobes erect, connivent. Seeds compressed, 1-seriate; cotyledons accumbent. — FI. Cap. i. p. 20. 1 South African species : M. torulosa, a rigid, branching, tomentose herb, with stellate pubescence ; radicle-leaves subsinuate. Flower pur- plish.— Northern districts and frontier. 2. NASTURTIUM, E. Br. Sepals spreading, equal at base. Pod nearly cylindrical, long or short, with round-backed valves. Styles short or longish ; stigma 2-lobed. Seeds small, turgid, 2 -seriate ; coty- ledons accumbent. — FI. Cap. i. p. 21. Water or marsh plants. — 2 South African species : N. officinale (common Watercress), with white flower ; N. fluviatile, E. M., with yellow flower ; the latter an Eastern district plant. 3. BARBAREA, E. Br. Sepals erect, equal at base. Pod 4-sided, 2-edged, the valves keeled at back, awnless at the apex. Stigma capitate. Seeds not margined, 1-seriate. — FI. Cap. i. p. 22. 8 VII. CRUCIFERS. 1 species : B. pracox, R. Br., a roadside weed, introduced from Europe. Stem branching ; lobes lyrate-pinnatifid ; flower small, bright yellow. 4. ARABIS, Linn. Sepals erect or spreading, equal, or the lateral saccate at base. Pod linear, compressed, with flat valves. Seeds in a single or double row .—FI. Cap. i. p. 22. Also Turritis, L. ; FI. Cap. 1. c. 2 South African species, found on high mountains of the Eastern district. Flowers white. 5. CARDAMINE, Linn. Sepals erect or spreading, equal at base. Pod linear or sub- lanceolate, mostly acute at each end ; valves flat, nerveless, opening with elasticity. Seeds ovate, not margined, 1-seriate. — FI. Cap. i. p. 23. 1 South African species : C. Africana , found in damp woods throughout the colony. Lobes on long petioles, 3 -partite ; the leaflets petiolate, ovate- acuminate, toothed. Flowers small, white. 6. ALYSSUM, Linn. Sepals erect, equal at base. Petals entire. Stamens toothed or entire. Pod orbicular or elliptical, the valves flat or convex in the centre. Seeds 1-4 in each cell, compressed, sometimes margined ; cotyledons accumbent. — FI. Cap. i. p. 23. Small annuals or perennials, covered with whitish stellate pubescence. Leaves entire. Flowers white or yellow.— 2 South African species. 7. SISYMBRIUM, All. Sepals spreading, equal at base. Filament toothless. Pod subterete, sessile ; valves convex, usually 3-nerved. Seeds several, ovate or oblong ; cotyledons incumbent or oblique. — FI. Cap. i. p. 24. Roadside weeds or weed-like plants, glabrous or hispid. Leaves either simple, pinnatisect or decompound. Flowers mostly yellow, rarely white. — 8 reputed South African species. 8. SENEBIERA, DC. Sepals spreading. Petals very small. Stamens 2-4-6. Pod 2-parted, subcompressed, indehiscent ; valves subglo- bose, ridged or crested. Seeds solitary. — FI. Cap. i. p. 27. Littoral or roadside weeds, widely dispersed over the globe, decumbent or prostrate. Leaves entire or mostly pinnatifid. Flowers very minute, in short racemes opposite the leaves. Smell very pungent. — 4 South African species. VII. CRUCIFERiE. 9 9. LEPIDIUM, Linn. Sepals equal at base. Pod ovate or subcordate ; tbe valves keeled; cells 1-seeded. Seeds 3-cornered or compressed. — FI. Cap. i. p. 28. The garden Cress (Z. sativum) is the type of this widely-dispersed genus. Sonder enumerates 10 species in FI. Cap. These are weed-like plants, with entire or variously and irregularly-cut leaves and small flowers in terminal elongating racemes. 10. CAPSELLA, Yent. Sepals flattisb, equal at base. Pod triangular or wedge- sbaped, the valves boat- shaped, wingless ; cells many-seeded. — FI. Cap. i. p. 31. 1 species, Shepherd’ s-purse, a common weed throughout the temperate zones ; introduced from Europe. 11. BRASSICA, Linn. Sepals erect or spreading, the lateral often saccate at base. Pilament toothless. Pod long, subterete, with a beak (some- times indehiscent and 1-seeded) ; the valves convex, 1-3- nerved, the lateral nerves often flexuous ; septum membranous or spongy. Seeds 1-seriate, subglobose or oblong; cotyle- dons broad, conduplicate (0 >>). — FI. Cap. i. p. 3. Also Sina- pis, Koch; FI. Cap. i . p. 32. Cabbage, Turnip, Mustard, etc. Mostly biennials, natives of the tempe- rate zones. Lower leaves petioled, lyrate or pinnatifid ; cauline sessile, subentire. Racemes elongate ; flowers yellow. — 4 South African species. 12. CHAMIRA, Thunb. Calyx 2-spurred at base. Pod substipitate, oblong, com- pressed, with a subulate beak. Seeds compressed, immargi- nate ; cotyledons elongate, twice folded on the radicle. — FI. Cap. i. p. 32. Endemic. C. cornuta , the only species, is a weakly, glabrous annual. Lower leaves opposite, reniform-cordate, 2 inches long, 3-4 inches wide ; upper alternate deeply cordate, the uppermost acuminate. Flowers small, white. — About Simon’s Town and near Saldanha Bay. 13. HELIOPHILA, Linn. Sepals equal at base. Pilaments of lateral stamens simple, or with a tooth near the base. Pod sessile or stipitate, dehis- cent, long or short, compressed, linear, oblong or lanceolate, the margins straight or undulate ; valves l-3-nerved,flattish, mostly membranous ; septum hyaline ; style short or long ; stigma 2-lobed. Seeds many or few, sometimes solitary, 1-2-seriate ; 10 VII. CRUCIFER JE. cotyledons twice folded on the radicle. — FI. Cap. i. p. 35 ; Ikes. Cap. t. 166. A large woolly South African genus of upwards of sixty species. These are distributed under the following sections : — 1. Leptormtts. Pods linear, moniliform ; the beads oval. Herbs. 2. Ormisctjs. Pods linear, moniliform ; beads globose. Herbs. 3. Selenocarpe.®. Pods oval or suborbicular. Herbs. 4. Orthoselis. Pods linear, with straight or straightish margins. Herbs or shrublets. 5. Pachystylium. Pods linear ; style short and thick. Pubescent half-shrubs. 6. Lanceolaria. Pods lanceolate. Glabrous shrublets and half-shrubs. 14. CARPONEMA, Sond. Pod sessile, indehiscent, linear, terete, tapering to each end, somewhat constricted between the seeds, with a very thin septum, 2-celled, one cell smaller and empty, the other several- seeded ; valves hardened, nerveless ; style conical. Seeds in a single row, oblong, terete, immarginate, separated by cross- partitions ; cotyledons twice folded on the radicle. — FI. Cap. i. p. 35. Endemic. 1 species, C. filiforme, Sond., a glabrous or pilose annual, 12-18 inches high. Leaves linear, the lower long ; racemes long ; flowers blue or purple ; pods pendulous, 1 inch long. — Found on the Cape flats. 15. Bit ACHY C ARPiE A, DC. Sepals equal at base. Pod indehiscent, deeply 2-lobed, with a very narrow septum ; valve very convex, netted and ridged ; cells 1-seeded. Seeds with long, linear cotyledons, spirally rolled on the radicle. — FI. Cap. i. p. 33. An endemic genus of 2 species, natives of the West and North-West districts. Glabrous, half-shrubby perennials, with oblong or linear entire leaves ; long leafless racemes, and handsome yellow or purple flower. 16. CYCLOPTYCHIS, E. Mey. Pod hard-shelled, indehiscent, orbicular-ovate, beaked ; sep- tum orbicular ; valves somewhat convex, with raised ridges, radiating from a prominent keeled centre ; seeds solitary in each centre ; cotyledons long, linear, twice folded on the ra- dicle.— FI. Cap. i. p. 34 ; Thes. Cap. t. 59. An endemic genus of 2 species, natives of the Western districts. They resemble Brachycarpcea in aspect, but differ in fruit. 17. PALMSTRU CKI A, Sond. Pod sessile, orbicular, very much flattened, indehiscent, 1- locular, 1-seeded. Seeds orbicular, compressed, with a mem- branous marginal wing ; cotyledons linear, incumbent, twice- folded. VII. CRUCIEERiE. 11 Endemic. A glabrous, erect herb, with remote, linear-filiform leaves. Flowers unknown. Fruiting racemes long ; ripe pods 7-8 lines long and wide, rounded at top, with a very short style ; valves flattish, nerveless. — This plant was found by Thunberg, about 1772, on the Onderste Bogge- veld ; no more recent collector has met with it ! Order YIII. CAPPARIDE^]. Flowers bisexual. Sepals 4. Petals 4-8 or more, clawed, often unequal, sometimes none. Stamens 4, 6, or many, not tetradynamous. Ovary 1-locular, with 2 parietal placentae. Fruit a dry capsule or fleshy berry. Seeds without albumen ; embryo curved or spirally rolled. — Trees shrubs or herbs, with alternate simple or compound leaves. Pubescence often glandular and fetid. Tribe 1. Cleome.®. Fruit a dry, dehiscent, pod-like capsule. Annual or perennial herbs ; leaves simple or compound. (Gen. 1-2.) Torus small, depressed or inconspicuous 1. Cleome. Torus tapering into a stalk-like gynophore .... 2. Gynandbopsis. Tribe 2. C apparel. Fruit a fleshy, indehiscent berry. Shrubs or trees. (Gen. 3-6.) Calyx-tube obconical, persistent ; limb of 4 deci- duous valvate lobes ; torus elongate, filiform. Berry ovoid, 1-celled, few- or many-seeded ... 3. Niebtjhria. Berry cylindrical, torulose, transversely many- ceLLed, each cell l-3eeded 4. M®rtja. Calyx of 4 deciduous separate sepals, not united into a tube at base. Torus long, slender, filiform. Stamens 4-8 . . 5. Cad aba. Torus short, hemispherical. Stamens few or many 6. Capparis. Tribe 1. Cleome^:. (G-en. 1-2.) 1. CLEOME, Linn. Calyx 4-toothed or 4-parted, persistent or deciduous. Pe- tals equal or unequal, sessile or clawed, entire ; convolute imbricate or open in the bud. Torus short or depressed, sometimes produced behind into a process or gland. Stamens 4-6-8 or more, either all perfect or some sterile, very generally unequal and decimate, the filaments often thickened under the apex. Ovary sessile or stipitate ; ovules many ; style none or elongate. Capsule mostly elongate, sometimes inflated, 1- celled, with thin valves. — FI. Cap. i. p. 55. Also Polanisia, Fajin. FI. Cap. i.p. 56. Dianthera, Kl. ; FI. Cap. i. p. 57 ; Thes. Cap. t. 136. Tetratelia, Sond. ; FI. Cap. i. p. 58. A large tropical and subtropical genu9 of herbs or half-shrubs, mostly strongly scented. Leaves simple or compound. The following subgenera are represented in the South African flora : — 1. Eucleome. Stamens 6, fertile. (2 species.) 12 VIII. CAPPABIDEiE. 2. Polanisia. Stamens 8-12, fertile. (2 species.) 3. Dianthera. Stamens 4-10 ; 2-8 short and sterile ; 2 anterior alone fertile, very long, declinate. (3 species.) 4. Tetratelia. Stamens 8 ; 4 sterile, short ; 4 fertile, elongate. (1 species.) All the Cape species are natives of the Northern or North-Eastern fron- tier, or of the regions beyond the Northern bonndary. 2. GYKAJNDROPSIS, DC. Sepals 4, short, spreading. Petals 4, clawed. Stamens 6, inserted on the summit of a long stalk-like torus ; filaments subequal ; anthers 2-celled, fertile. Ovary stipitate ; ovules many ; stigma subsessile. Capsule pod-like, 1-celled, 2-valved, many-seeded. Seeds rugose. — FI. Cap. i. p. 55. Annuals, chiefly tropical. G. 'pentaphylla is found on the Asbestos Mountains and Magalisberg; it is a native of the West Indies, but now naturalized throughout the tropics generally. Teibe 2. Cappaeea:. (Gen. 3-6.) 3. NIEBUHRIA, DC. Calyx with a funnel-shaped, persistent tube, and a 4-parted, deciduous limb ; lobes valvate in aestivation. Petals 0 or very small. Torus filiform, long or shorter, cylindrical. Sta- mens many, on the apex of the torus ; filaments filiform. Ovary on a long stipe, 1-celled ; ovules many or few. Berry ovoid or subglobose, 1-celled, many- or few-seeded. — Niebuhria, FI. Cap. i. p. 60 ; and Boscia, FI. Cap. 1. c. ; Thes. Cap. t. 134. Small trees or shrubs, glabrous or pubescent. Leaves 3-foliate or simple. Flowers terminal or axillary, solitary or corymbose. — 5 South African species, of which 3 have 3-foliate, and 2 simple leaves. 4 are na- tives of the Eastern district, or Natal ; 1 ( Boscia angustifolia , H.) of Namaqualand. 4. MJERUA, Eorsk. Calyx with a funnel-shaped, persistent tube, and a 4-parted, deciduous limb ; lobes valvate. Petals 0, or 4 in the throat of the calyx. Torus filiform, elongate, cylindrical. Stamens many, inserted in the middle of the torus ; filaments free or connate at base. Ovary on a long stipe ; ovules many ; stig- ma subsessile. Berry cylindrical, torulose, transversely many- celled ; cells 1-seeded. Trees, chiefly tropical, often glaucous. Leaves simple, quite entire. Flowers axillary or terminal. — M. Angolensis has been recently found in the Natal country by Mr. Gerrard ( Gerr . and M‘K. ! 1148). 5. CADABA, Forsk. Sepals 4, separate, unequal, deciduous, concave, the two outer valvate in the bud and covering over the 2 inner. Petals Till. CAPPARIDEiE. 13 (2-4 or) none. Stamens 4-8, fertile, inserted on the apex of a long, filiform torus, which has a hood- or bottle-shaped ap- pendage at its base, on the upper side. Ovary stipitate, ob- long, 1-locular ; ovules many ; stigma sessile. Berry cylin- drical, fleshy and indehiscent, or leathery, splitting irregularly. — FI. Cap. i. p. 59 ; and Schepperia, FI. Cap. 1. c. ; Th.es. Cap. t. 60 and t. 135. Shrubs, glabrous or glandular, sometimes leafless. Leaves simple or 3- foliolate. Flowers axillary, solitary or racemose. — 2 South African species : C.juncea (Schepperia juncea, DC.), a native of dry plains in the Northern, North-Western, and Eastern districts ; and C. Natalensis , Sd., found in Natal and British Caffraria. 6. CAPPARIS, Linn. Sepals 4-5, free or connate at base, valvate or imbricate. Petals 4, rarely more, imbricated. Torus small, hemispherical. Stamens mostly numerous, sometimes 8, on the torus; filaments free. Ovary stipitate ; ovules many ; stigmas sessile. Berry on a long stipe, globose or cylindrical, many- or few-seeded. Seeds lying in pulp ; embryo spirally rolled. — FI. Cap. i.p. 61. Trees or shrubs, often climbing or scrambling, unarmed or spiny. Leaves simple. — 9 Cape species, all Eastern or from Natal. Order IX. RESEDACEAD. Blowers bisexual, small, green or white. Sepals several, persistent. Petals minute. Stamens 3 or several. Ovary 1 -celled, open at the summit, with 3-4 parietal placentae. Bruit a gaping capsule. Seeds without albumen ; embryo curved. — Small herbs or suffrutices, with alternate, simple or cut leaves. “ Mignonette” belongs to this Order. 1. OLIGOMERIS, Cambess. Calyx 4-5-parted, the segments sometimes unequal. Petals 2, alternate with the upper sepals, flat, simple (not lobed), without appendage, separate or connate at base. Torus obso- lete. Stamens 3-8, hypogynous ; filaments subulate, flat, united at base into a cup, persistent ; anthers deciduous. Ovary 1-celled, 4-angled, with 4 conical points ; placentae 4, parietal ; ovules many. Capsule membranous, inflated, open at the summit, 4-horned. — FI. Cap. i. p. 64. Small, slightly fleshy perennials ; leaves very narrow, undivided ; flowers minute, white, in terminal, bracteate spikes. — 4 species, found on dry hills in various parts of the colony. Order X. BIXACEiE, Endl. Blowers small, often unisexual, regular. Sepals often 14 X. BIXACEiE. united at base, persistent; sometimes partly adnate to the ovary. Petals sometimes wanting, rarely large. Stamens de- finite or indefinite, hypogynous or rarely perigynous. Ovary 1-locular, with parietal placentae. Emit either a fleshy berry, or a pulpy, dehiscent capsule. Seed albuminous ; embryo straight, central. — Trees or shrubs, with alternate, simple leaves ; often spiny. Suborder 1. Bixinea:. Ovary quite free. Stamens bypogynous. (Gren. 1-7.) Flowers bisexual or polygamous. Style columnar or short. Sepals deciduous. Petals 10-12, much larger than the sepals 1. Oncoba. Sepals 4-5, persistent. Petals 4-5, small, deciduous 2. Rawsonia. Calyx persistent, deeply 10-12-fid in 2 rows ; petals none 3. Scolopia. Flowers unisexual, dioecious. Stigma subsessile. Ovule solitary 4. Xylosma ? Flowers unisexual, dioecious. Styles as many as the carpels. Anthers opening by slits. Petals 0. Calyx 6-10-fid in a double row ; capsule .... 5. Trimeria. Calyx 5-7-fid in a single row ; berry 6. Dovyalis. Anthers opening by pores. Petals 5, with a scale at base 7. Kiggelaria. Suborder 2. Homalinea. Flowers bisexual. Ovary half-inferior. Stamens perigynous. (Gren. 8.) Calyx with a conical, adherent tube, and a 10-30-fid limb ; the segments in two rows 8. Homxlium. Suborder 1. Bixineas. (Gren. 1-7.) 1. ONCOBA, Forsk. Flowers polygamous. Sepals 3-5, deciduous, imbricated. Petals 6-12, much larger than the sepals, obovate, clawed, im- bricate. Stamens many, inserted in several rows on a fleshy torus; filaments filiform; anthers linear or oblong, erect, opening by slits. Ovary with 3-10 parietal placentae ; style simple ; stigma radiate ; ovules many. Berry leathery, pulpy within ; seeds numerous. — FI. Cap. i. p. 66. Trees or shrubs, glabrous or pubescent, sometimes spinous. Leaves simple, entire or serrulate. Flowers terminal, large, white or yellow. — 3 South African species ; 2 found at Natal, 1 at Delagoa Bay. 2. RAWSONIA, Harv. and Sond. Flowers perfect or polygamous. Sepals 4-5, very unequal, concave, imbricate, persistent. Petals 4-5, unequal and con- cave like the sepals and not longer, deciduous. Petaloid scales (staminodia) opposite the petals and longer, each with a 2-lobed, fleshy gland at base. Stamens many, in several X. BIXACE.E. 15 rows, the inner bypogynous, the outer attached to the base of the petaloid scales ; anthers sagittate, erect. Ovary on a con- vex torus, with 4-5 parietal placentas; ovules many;" stigma subsessile, 4-5-parted. Fruit . . . ? — FI. Cap. i. p. 67. R. lucida is a Natal shrub, with glossy, exstipulate, serrate leaves, and axillary, subcapitate spikes of (yellow ?) flowers. 3. SCOLOPIA, Schreb. Flowers bisexual. Calyx persistent, with a short, conical tube, and a 10-12 -parted limb ; the segments in two rows, the inner smaller. Torus fleshy, filling the calyx-tube, its margin, opposite the bases of the outer calyx- segments cut into nume- rous, glandular lobules. Stamens very many, in several rows within the margin of the disk, slightly perigynous ; filaments capillary ; anthers 2-celled, acuminate or horned, splitting. Ovary free, sessile, with 2 indexed placentae ; ovules few ; style filiform ; stigma subcapitate: Berry fleshy, few-seeded. — Phoberos, Lour. ; FI. Cap. i. p. 67. Shrubs or small trees, often spiny. — 3 species found in the Eastern dis- trict ; others recently at Natal. 4. XYLOSMA, Eorst. Flowers dioecious. Male : Sepals 4-5, scale-like, often ciliate, imbricate. Petals 0. Stamens many, often sur- rounded by a glandular disk ; anthers short, versatile, splitting. Female : Ovary on an annular disk ; placentae parietal, 1 (or 2-3-6) ; ovule solitary (or few) ; style none (or entire or divided) ; stigma dilated. Berry indehiscent, small, l-(or few-) seeded. — Thes. Cap. Trees, often spiny. Leaves toothed, rarely entire. Flowers shortly ra- cemose or tufted in the axils of the leaves, minute. X. monospora , Harv., recently found near Natal by Mr. Cooper. 5. TRIMERJA, Harv. Flowers dioecious. Male : Calyx 6-10-parted in a double row, the inner larger. Disk bearing marginal glands opposite each of the outer lobes of the calyx. Petals 0. Stamens 9-10, subperigynous, inserted in parcels of 3-4, alternating with the glands of the disk. Female : Calyx as in the male, but without glands. Ovary free, sessile, with 3 parietal pla- centae ; styles 3, short, persistent; ovules 1-2 on each placenta. Capsule dry, 3-valved, 1-3-seeded. — FI. Cap. i. p. 68. Shrubs or trees. Leaves alternate, 3- or many-nerved, toothed. Flowers minute, spiked or panicled. — 2 species, from Eastern frontier and Natal. 16 X. BIXACEiE. 6. DOVYALIS, E. Mey. Flowers dioecious. Male : Sepals 5-8, tomentose, scarcely imbricate. Petals none. Stamens 12-20 or many, inserted on a fleshy, glandular disk ; filaments filiform ; anthers didymous, short. Female : Ovary seated on a lobed, fleshy disk, with 2-6 or more, strongly inflexed placentae ; ovules solitary or few, pendulous ; styles as many as the placentae, spreading. Berry indehiscent, pulpy within, few-seeded. Seeds glabrous or woolly. — FI. Cap. i. p. 69. Also Aberia, FI. Cap. i. p. 70, and ii. ( Suppl .) p. 584. Shrubs or small trees, often spiny. Leaves simple, entire or toothed, alternate. Flowers axillary, small ; the males tufted, females solitary. There are two sections or subgeneray viz. : — 1. Dovyalis, E. M. Seeds glabrous. (2 species.) 2. Abeeia, Hochst. Seeds woolly. (4 species.) In the ‘ Flora ’ I retained Aberia as distinct from Dovyalis , but the recent discovery of two new species with intermediate characters, has induced me to unite them. One of these, the “ Kei apple” ( Aberia Caffra, Hook. f. and Harv.), has 6 (or probably more) styles and placentae. There is abso- lutely no difference whatever in habit between the species of the two sub- genera. 7. KIGGELABIA, Linn. Flowers dioecious. Calyx 5-parted, deciduous ; sepals val- vate in bud. Petals 5, imbricate, leathery, each with a fleshy gland at its base inside. Male : Stamens 10, crowded in the base of the calyx ; filaments short ; anthers hard and dry, 2- celled, opening by terminal pores. Female : Ovary sessile, with 2-5 parietal placentae ; styles 2-5, short ; ovules many. Capsule globose, pubescent, leathery, several-seeded, bursting imperfectly into 2-5 valves, — FI. Cap. i. p. 71. Endemic shrubs or small trees. Leaves simple, entire or denticulate. Male flowers in axillary cymes ; female solitary. — 3 species. Subokdek 2. Homaline^:.* (Gen. 8.) 8. HOMALIUM, Jacq. Flowers bisexual. Calyx persistent, with a conical tube and multipartite (10-30-cleft) limb ; segments in two rows, the inner ones largest. A gland opposite the base of each of the outer segments. Stamens perigynous, alternating with the glands singly or in parcels of 2 or 3 ; filaments filiform ; anthers didymous, opening longitudinally. Ovary half-infe- rior, 1-celled, with 3-5 parietal placentae ; styles 3-5, subulate, * This Suborder is usually regarded as a distinct Order, and placed in Calycijlorce. I retain it here on account of the very intimate relationship between Domalium, Scolopia, and Trimeria. X. BTXACEJE. 17 divergent ; ovules few, pendulous. — FI. Cap. i. p. 72 (Black- wellia). Shrubs or small trees, mostly tropical. Leaves toothed or entire, sim- ple. Flowers in axillary or terminal spikes, racemes or panicles. — 1 or perhaps 2 species found at Natal. Order XI. VIOLARIE.®. Blowers bisexual. Sepals 5, persistent, separate. Petals 5, unequal, the lower one spurred at base. Stamens 5 ; fila- ments broad and flat ; anthers adnate, conniving round the stigma. Ovary 1-locular, with 3 parietal placentas. Capsule 3-valved, bearing the seeds in the middle of each valve. — Herbs or half- shrubby plants, with alternate, simple, entire or cut, stipuled leaves. Blowers axillary, solitary or variously arranged. “ Yiolet ” and “ Pansy.” Sepals eared at base 1. Viola. Sepals not eared at base 2. Ioniditjm. 1. VIOLA, Linn. Sepals 5, nearly equal, prolonged at base into ear-like lobes. Petals 5, unequal, the under one ( labellum ) spurred or saccate at base. — FI. Cap. i. p. 73 ; Thes. Cap. t. 46. Herbs or small under, hrubs. — 2 South African species, with linear or linear-lanceolate leaves, and subulate stipules. Flowers violet-blue. Na- tives of the South-Western districts. V. arvensis , Murr., a weed of culti- vation, is naturalized throughout the colony. 2. IONTIDIUM, Vent. Sepals 5, unequal, not prolonged at base. Petals 5, very unequal, the under one ( labellum ) much larger than the rest, with a dilated and concave claw, shortly spurred or saccate at base. — FI. Cap. i.p. 74 ; Thes. Cap. t. 47. Herbs or undershrubs. Leaves alternate or opposite, serrate or entire, stipulate. Flowers axillary or in terminal racemes.— 4 South African spe- cies, natives of the Eastern district and Natal. Order XII. DROSERACEAD. Blowers bisexual, regular. Sepals 5, persistent. Petals 5, equal. Stamens 5, on slender filaments. Ovary 1-celled, with 3-5 parietal placentas, or 3-celled ; styles 1-3-5, often forked or multifid. Capsule dry, splitting. — Herbs or under- shrubs, covered with viscidly glandular hairs. “ Sundew or Catch-fly.” * Styles 3-5, 2-fid or 2-partite. Ovary 1-celled .... 1. Drosera. Style 1, simple, stigma capitate. Ovary 3-celled ... 2. Roridula. c 18 XII. DltoSERACEiE. 1. DROSERA, Linn. Calyx 5-parted. Petals 5, obovate. Stamens 5 ; anthers adnate, opening by slits. Ovary 1-celled, with 3-5 parietal placentas ; ovules many ; styles 3-5, 2-fid or 2-partite, the branches undivided or multifid. Capsule membranous, 3-5- valved, many-seeded. — FI. Cap. i. p. 75 ; Thes. Cap. t. 26. Stemless or caulescent herbs. Leaves scattered or rosulate, bearing on the upper surface and margin glandular hairs, stipulate. Flowers in scor- pioid cymes or secundly racemose, rarely solitary ; rosy purple or white, handsome. Petals very delicate, soon withering. — 8 South African species, dispersed through the colony. 2. RORIDULA, Linn. Calyx 5-parted. Petals 5, oval or oblong. Stamens 5 ; anthers adnate, opening by pores. Ovary 3-celled ; ovules soli- tary or in pairs, pendulous from the summit ; style simple ; stigma capitate. Capsule 3-celled, 3-valved ; seeds solitary. — FI. Cap. i. p. 79. Shrublets, very viscid and gland-hairy. Leaves linear-lanceolate, acu- minate, entire or pinnatifid, 2-3 inches long, lg— 2 lines wide. Flowers spiked or racemose. — 2 species, natives of the Western district. Order XIII. POLYGALEiE. Plowers bisexual, irregular. Sepals 5, unequal, the two lateral often coloured like petals. Petals 3-5 ; the two lateral small ; the lower (keel) large, enclosing the stamens and ovary, and often crested in front. Stamens 8, monadelphous ; an- thers opening by pores. Ovary 2-celled ; ovules solitary, pendulous. Fruit a capsule or drupe. — Small shrubs, half- shrubs, or herbs. Leaves simple, entire, without stipules. Flowers racemose or spiked. Sepals very unequal, the two lateral wing-like. Fruit a 2-celled, oblong or obcordate capsule ... 1. Polygala. Fruit a fleshy, ovoid drupe 2. Mundtia. Fruit 1-celled, dry, indehiscent, winged at the apex . 4. Securidaca. Sepals subequal, similar ; fruit an oblong capsule ... 3. Muraltia. 1. POLYGALA, Linn. Sepals 5 ; the two lateral much larger than the rest, wing- like and coloured. Petals 3-5, united at base and attached to the staminal tube ; the lower boat-shaped, usually crested below the apex ; the lateral small, simple or 2-fid ; upper fre- quently wanting. Stamens 8, united into a split tube, and hidden within the lower petal ; anthers fixed, 1-celled, opening by terminal pores. Style bent upwards ; stigma oblique. XITT. POLYGALEiE. 19 Capsule 2-celled, membranous, compressed, dehiscing at the margins ; seeds mostly pubescent. — FI. Cap. i. p. 80. A cosmopolitan genus. — About 40 South African species (several recently discovered at Natal, undescribed), dispersed. Shrubs, half-shrubs, perennial or annual herbs. Juice very bitter, tonic. 2. MUNDTIA, Kunth. Sepals, petals, and stamens as in Polygala. Style com- pressed, thickened upwards, 2-lobed, the posterior lobe hori- zontal, the anterior vertical. Fruit a fleshy, ovoid, 1-2 -seeded drupe. — FI. Cap. i. p. 95. M. spinosa, a spinous, much-branched, rigid shrub, with oblong, ob- ovate, cuneate or linear, thick, glabrous leaves, and red or white flowers, is common throughout the colony. The succulent fruit is eaten by birds and children. 3. MURALTIA, Neck. Sepals 5, dry and membranous, subequal, the two lateral somewhat longer than the rest. Petals and stamens as in Polygala. Capsule membranous, compressed, with 4 horns or tubercles at its upper angles ; very rarely hornless. — FI. Cap. i.p. 95. An endemic genus of upwards of 50 species. They are small but rigid shrubs or half- shrubs, with tufted or rarely scattered, rigid, mostly pungent- mucronate, narrow, entire leaves. Flowers axillary, solitary, subsessile or pedicellate, bright purple, 3-bract ed at base. 4. SECURIDACA, Linn. Calyx and stamens as in Polygala. Two lateral petals attached to the base of the staminal tube, separate from the keel, erect, connivent; keel about of equal length, concave, helmet-shaped, erect or 3-lobed ; upper petals rudimentary or 0. Anthers 2-celled. Ovary 1-celled, 1-ovuled. Fruit samaroid, indehiscent, hard or woody at base and often crested, produced at the apex into a long or shortish wing. — FI. Cap. ii. p. 585. Shrubs or shrublets, often climbing ; natives of America, Africa, and Asia. Leaves mostly entire and 2-glanded. Racemes or panicles terminal or axillary. S. oblongifolia, also a native of Abyssinia, occurs at Algoa Bay. Order XIY. PITTOSPORE^l. Flowers perfect, regular or suboblique. Sepals 5, separate, imbricate. Petals 5, hypogynous, imbricate, longer than the sepals, generally with an erect claw and spreading limb. Stamens 5, hypogynous, free, alternate with the petals ; an- thers versatile. Torus small, conical. Ovary 2- (rarely 3-5-) c 2 20 XTY. PITTOSPOREiE. celled ; style single, terminal ; ovules many, axile. Fruit capsular, or fleshy and pulpy. — Trees or shrubs, commonly glabrous. Leaves alternate, entire or toothed. Flowers white blue yellow greenish or red, conspicuous. 1. PITTOSPORUM, Banks. Sepals separate or connate at base. Petals 5, their claws erect, connivent ; limbs spreading. Filaments subulate. Ovary sessile, imperfectly 2-5-celled ; style short ; stigma capitate. Capsules subglobose or obovate, with leathery, thick, septife- rous valves. Seeds lying in viscid resin, smooth. — FI. Cap. i. p. 443. A considerable genus, widely dispersed, chiefly in the southern hemi- sphere. P. viridiflorum , Sims, the only Cape species, is a glabrous shrub, common in the Eastern district, Caffraria, and extending to Natal. Leaves obovate, tapery at base, very entire, leathery, shining ; panicle subumbel- late, terminal. Petals yellow-green. Order XY. FEANKENIACEJE. Flowers bisexual, regular. Calyx tubular, 4-5-toothed, ribbed, persistent, hardening after flowering. Petals 4-5, with long claws, deciduous. Stamens mostly 6. Ovary 1- celled, with 3-5 parietal placentas ; ovules many. Capsule enclosed in the calyx, many-seeded ; seeds albuminous, with an axile embryo. — Small, perennial, or half-shrubby plants, with crowded, small, narrow leaves. Flowers purple or white. 1. FRANKENIA, Linn. The only genus of the Order. — FI. Cap. i. p. 114. 3 species found in South Africa, natives of salt-marshes or seashores. Order XYI. CARYOPHYLLEiE * Flowers bisexual, regular. Calyx 4-5-cleft or parted, persistent, imbricate. Petals 4-5, entire or 2-fid, or minute or 0. Stamens 4-10, inserted with the petals on a hypogy- nous or slightly perigynous ring ; filaments filiform ; an- thers 2-celled. Torus often minute (in some Silenea elon- gate, stipe-like). Ovary free, 1-celled, or rarely at base 2-5- * The following genera, included in this Order in ‘ Flora Capensis,' (following the views of Bentham and Hooker) are removed as follows : — Corrigiola, Herniaria , Pollichia, to Paronychie.® ; Orygia , Glinus, Mollugo , Pharnaceum , Hypertelis , Codanthum , Psammatropha, Adenogramme , and Polpoda, to Ficoide^:. XVI. CART OPHYLLEiE . 21 celled ; styles 2-5, spreading, or more or less combined, stigmatose on tbe upper surface ; ovules 2 or many, on cords rising from tbe bottom of tbe ovarian cavity, free or united in a central column. Capsules membranous or crustaceous, opening by apical teeth or splitting into valves, rarely sub- indehiscent or opening transversely. Seeds reniform, subcom- pressed ; embryo curved round the albumen or excentric. — Herbs or half-shrubby plants ; branches mostly swollen at the nodes. Leaves opposite, quite entire, exstipulate or with mem- branous stipules. Flowers in cymes panicles or clusters. Tribe 1. Sileneje. Calyx tubular, 4-5-toothed. Petals and stamens hypogynous, borne, with the ovary, on a stipe-like “ gynophore,” rarely sessile. Petals clawed. Styles separate to the base. Stipules 0. (Gen. 1-3.) Styles 2. Calyx bracteate at base ....... 1. Dianthtjs. Styles 3. Calyx nude at base, 5-toothed 2. Selene. Styles 5. Calyx nude at base, with leafy lobes ... 3. Agrostemma. Tribe 2. Alsinea:. Sepals 4-5, separate to the base, or nearly so. Petals and stamens hypogynous. Petals not clawed. Styles separate to the base. Stipules 0 or membranous. (Gen. 4-7.) Stipules none ; petals 2-fid. Styles 3 4. Stellaria. Styles 4-5 5. Cerastium. Stipules membranous ; petals entire. Styles 5. (Petals white.) 6. Spergttla. Styles 3. (Petals pink or purple.) 7. Spergularia. Tribe 3. Polycarpea:. Calyx as in Alsinece ; also the petals, but com- monly much smaller or obsolete. Style simple at base, 3-fid or rarely 2-fid above. Stamens 5 or fewer. Stipules membranous. (Gen. 8-10.) Petals deeply 2-fid or 4-fid .8. Drymaria. Petals entire. Sepals membranous, not keeled .9. Polycarp^a. Sepals membranous-edged, sharply keeled . . . . 10. Polycarpon. Tribe 1. Silenea. (G-en. 1-3.) 1. DIANTHUS, Linn. Calyx tubular, 5-toothed, with 2 or more close-lying bracts at base. Petals 5, with long claws, often toothed or lacerate. Stamens 10. Styles 2, filiform. Capsules cylindrical or ob- long, opening by 4 teeth. — FI. Cap. i. p. 122. Herbs or small half-shrubs. Leaves generally grass-like. Flowers ter- minal, solitary or in cymes or panicles. — The garden Pink and Carnation belong to this genus. 9 reputed South African species, several of them depending on very slight characters, are dispersed through the colony. 2. SILENE, Linn. Calyx tubular, 5 -toothed, ebracteate at base. Petals 5, on 22 XYI. CAltYOPHYLLE^E. long claws, the limb entire, 2-fid or multifid. Stamens 10. Ovary partially 3-5-celled at base, 1-locular above. Styles 3. Capsules opening by 6 teeth. — FI. Cap. i. p. 125. Mostly herbaceous, often annual, dispersed over the globe. Leaves broad or narrow, frequently pubescent or viscidly hairy. — 13 Cape species. 3. AGROSTEMMA, Linn. Calyx tubular, leathery, with 5 long, leafy, spreading lobes. Petals 5, clawed, not coronate. Stamens 10. Styles 5. Capsules opening by 5 teeth. — FI. Cap. i. p. 129. A. Githago, Linn., the “ Corn-cockle,” a weed of cornfields, introduced from Europe. Tribe 2. Alsike^. (Gren. 4-7.) 4. STELLARIA, Linn. Calyx 4-5-parted. Petals 2-fid or 2-parted. Stamens 8-10, rarely fewer. Styles 3. Capsules opening by 6 teeth. — FI. Cap. i. p. 130. S. media , Till, (common Chickweed), a cosmopolitan weed, occurs in cultivated ground. 5. CERASTIUM, Linn. Sepals and 2-fid petals 4-5. Stamens 8-10. Styles 4-5. Capsules opening by 8-10 teeth. — FI. Cap. i. p. 130. Weeds and weed -like plants, mostly hairy, often clammy. Sonder de- scribes 4 species, one of which is a weed of cultivation. 6. SPERGULA, Linn. Calyx 5-parted ; sepals herbaceous, membrane-edged. Pe- tals 5, ovate, entire, subhypogynous. Stamens 5-10, onasub- hypogynous ring. Styles 5. Capsules 5-valved. — FI. Cap. i. p. 135. Small herbs of temperate climates. S. arvensis, Linn., is a weed in neg- lected fields. 7. SPERGULARIA, Pers. Calyx 5-parted ; sepals herbaceous, obtuse. Petals 5, en- tire, subhypogynous. Stamens 10, rarely 5-3, on a subhypo- gynous ring. Styles 3. Capsules 3-valved. — FI. Cap. i.p. 134. Lepigonum, Fries . Small cosmopolitan herbs, found by the seashore and in salt soil, through- out the colony. Flowers purplish or rosy, pretty. Leaves awl-shaped, fleshy, with membranous stipules. Tribe 3. Polycarpe^:. (Gren. 8-10.) 8. DRYMARIA, Willd. Calyx 5-parted. Petals 5, deeply 2-4-lobed or parted. XVI. CARYOPIIYLLKiE. 23 Stamens 5 or fewer, subhypogynous. Ovary many-ovuled. Style filiform, 3-fid above. Capsules membranous, 3-valved, many-seeded. — FI. Cap. i. p. 135. JD. cordata, W.. a common South American plant, was found at the Hanglip by Mundt. It has cordate or roundish leaves, bristle-like stipules, and small white flowers in forked, much-branched cymes. 9. POLYCARP JEA, Linn. Calyx 5-parted ; sepals more or less membranous, concave, nerveless, not keeled, entire, pointless. Petals 5, subhypogy- nous, entire or 2-dentate. Stamens 5, hypogynous. Ovary many-ovuled; style 3-fid. Capsules 3-valved, many-seeded. — FI. Cap. i. p. 133. Small, much-branched herbs, common to the tropical and subtropical regions of both hemispheres. Leaves opposite or pseudo-verticillate, linear, oval, or spathulate. Stipules shining, silvery, copious. Flowers in tufts or corymbose cymes. — P. corymbosa , Lam., a widely-dispersed species, occurs at Port Natal. 10. POLYCARPON, Loeffl. Calyx 5-parted ; sepals herbaceous, membrane-edged, com- pressed, strongly keeled and mucronate. Petals 5, entire or emarginate. Stamens 3-5. Ovary many-ovuled. Style 3-fid. Capsules 3-valved, many-seeded. — FI. Cap. i. p. 133. Small herbs. P. tetrwphyllum , a decumbent, forked, much-branched annual, with obovate, opposite or 4-nate leaves, panicled flowers, short emarginate white petals, and silvery stipules, is a common weed by road- sides and in waste places throughout the colony. Order XVII. PORTULACEiE. Plowers regular, perfect. Sepals fewer than the petals, commonly 2, rarely 5, free or adnate to the ovary, strongly imbricate. Petals 4-5, rarely many, hypogynous or rarely pe- rigynous, free or connate, imbricate, withering early. Stamens inserted with the petals, mostly definite. Ovary free or half- inferior, 1-celled ; style 2-3-fid ; ovules 2 or many, on long, slender cords rising from the base, and either free or united in a central column. Capsules membranous or crustaceous, splitting into as many valves as the style branches. Seed as in Caryoyhyllece. — Herbs or shrubs, mostly succulent and gla- brous. Leaves alternate or opposite, entire, often fleshy. Capsules half-inferior, opening transversely, many- seeded 1. POETTJLACA. Capsules 3-valved, splitting longitudinally. Seeds winged, numerous 2. Anacampseros. Seeds wingless, numerous 3. Talinum. Fruit 3-winged, indehiscent, 1-seeded 4. Portulacaria. 24 XVII. POltTULACEJE. 1. PORTULACA, Linn. Sepals 2, united at base into a tube attached to the ovary, free above, deciduous. Petals 4-6, free or slightly connate at base. Stamens 8 or many. Ovary half-inferior, many-ovuled ; style deeply 3-8-fid. Capsule membranous, half-inferior, splitting across (circumscissile) about the middle. — FI. Cap. ii . p. 381. Small, fleshy, widely-dispersed herbs. P. oleracea is a common weed in gardens throughout the colony. P. quadrifida and P. pilosa are rare. 2. ANACAMPSEROS, Sims. Sepals 2, at length deciduous. Petals 5, very fugacious. Stamens 15-20 or more, hypogynous ; filaments distinct, ad- hering to them. Ovary free, many-ovuled ; style filiform, 3-fid at the apex. Capsules conical, 1-celled, 3-valved, the valves often 2-partite, and then apparently 6-valved ; seeds many-winged. — FI. Cap. ii. p. 382. A South African genus of 9 species, natives of dry plains to the north and north-east. Yery dwarf plants. Leaves roundish or ovate, or lanceolate, fleshy, sometimes very minute. Stipules membranous, larger than the leaves, much split. Flowers sessile, involucred or on long peduncles, opening only in sunshine. 3. TALINUM, Juss. Sepals 2) ovate, separate, deciduous. Petals 5, ephemeral. Stamens 10-30, inserted with the petals, and often adnate to them. Style filiform, 3-fid at the apex. Capsules 3-valved, 1-celled, many-seeded. Seeds wingless, many. — FI. Cap. ii.^?. 385. Chiefly Asiatic and American. T. Caffrvm, E. and Z., the only South African species, grows in Caffraria. It is a glabrous perennial, £-1 foot high. Leaves oblong-linear or linear-lanceolate, with revolute margins. Flowers solitary, in the axils of the upper leaves. 4. PORTULACARIA, Jacq. Sepals 2, persistent, membranous. Petals 5, persistent, equal, obovate, hypogynous. Stamens 5-7, inserted with the petals ; anthers short, often imperfect. Ovary ovate, 3-angled ; style 0 ; stigmas 3, spreading. Fruits 3-quetrous, 3-winged, indehiscent, 1-seeded. — FI. Cap. ii. p. 385. Glabrous, fleshy shrubs or small trees, natives of South Africa. Leaves opposite, roundish, flat, deciduous. Peduncles opposite ; pedicels 1-flowered, in threes. Flowers very small, rosy. “ Speckboom.” — 2 species : P. Afra, Jacq., grows in the Karroo and in the Eastern district; P. dAamaquensis , Sond., in Namaqualand. Order XVIII. TAMARISCINEiE. Mowers bisexual, minute, regular, 4-5-parted, spiked or ra- cemose. Stamens 4-5, united at base into a ring. Ovary 1- celled, with parietal placentas ; ovules many ; styles 3. — Shrubs, with minute, scale-like, crowded or imbricate leaves. 1. TAMARIX, Linn. Sepals 4-5, unequal, imbricated. Petals 4-5, hypogynous, equal. Stamens 4-10 on the glandular margin of a fleshy, hy- pogynous ring ; filaments separate at base. Styles 3. Cap- sules 3- rarely 2-4-valved, many-seeded ; seeds with a terminal tuft of silky hairs. — FI. Cap. i. p. 119. Small shrubs, growing in deserts or in salt soils. Leaves very minute, often reduced to mere scales. Flowers pink or white, conspicuous by their abundance, in terminal spiked racemes.— T. articulata , Vahl, is found in Namaqualand. Order XIX. ELATINACEiE. Plowers bisexual, , regular. Sepals 2-5, separate. Petals 2-5. Stamens as many or twice as many. Ovary 2-5-celled ; ovules many; styles 2-5. Capsule splitting into 2-5 valves. Seeds without albumen. — Small herbs or half-shrubs, growing in wet places. Leaves opposite, entire or toothed, with inter- petiolar membranous stipules. Flowers axillary. 1. BERG-IA, Linn. Sepals 5. Petals 5, hypogynous. Stamens 10, those oppo- site the petals shortest, and occasionally wanting. Ovary 5- celled ; styles 5 ; ovules many. Capsule 5-celled, 5-valved, many-seeded. — FI. Cap. i. p. 115 ; Thes. Cap. t. 24, 133. 4 South African species, dispersed through the colony. Order XX. HYPERXCINEjE. Flowers bisexual, regular. Sepals 5, persistent, imbricate. Petals 5, unequal-sided, spirally twisted in the bud, and often black-dotted at margin. Stamens numerous, united in 3-5 parcels ; anthers 2-celled. Ovary imperfectly 3-5-celled ; styles 3-5, spreading ; ovules many. Fruit a dry or fleshy cap- sule, 1-5-celled. Seeds without albumen. — Shrubs or herbs, with resinous juice. Leaves opposite, very entire, mostly pel- lucid-dotted, simple, exstipulate. Flowers showy, but not fragrant. 26 XX. IlYPERICINEiE. 1. HYPERICUM, Linn. Calyx 5-parted ; sepals equal, or 2 outer largest. Petals 5. Stamens many, in 3-5 parcels. Ovary sessile, 1- or imper- fectly 3-5-celled. Capsule commonly splitting. — FI. Cap. i. p. 117. A cosmopolitan genus, much more numerous in the northern hemisphere. There are 2 South African species, small herbs, with opposite, sessile, dotted leaves, and yellow cymose flowers. A third species ( H . humifusum), proba- bly of European origin, occurs among Drege’s Cape plants ; no habitat given. Order XXI. GUTTIFEBJE. Flowers regular, commonly dioecious or polygamous. Sepals 2-6, rarely more, very strongly imbricate, or decussate in pairs. Petals as many as sepals, hypogynous, strongly imbricate or twisted. — Male : Stamens mostly many, hypogynous ; fila- ments mostly short and thick, free or variously connate ; an- thers thick, mostly opening outwards. Ovary rudimentary or more or less formed. — Female : Staminodia surrounding ovary, mostly definite or fewer than in the male. Ovary sessile on a flat or fleshy torus, 2- or many-celled ; stigmas as many as the ovarian cells, sessile or subsessile, radiating or connate in a peltate disk, rarely on a long style ; ovules 1 or many axile. Fruit mostly fleshy, or with a leathery coat, pulpy within, rarely dehiscent. Seeds thick, without albumen. — Mostly tropical trees and shrubs, with resinous juice, and opposite, quite entire, simple, exstipulate leaves. 1. GARCINIA, Linn. Flowers polygamous. Sepals 4, decussate. Petals 4, im- bricate.— Male : Stamens many, free or connate in an entire or 4-lobed mass, or tetradelphous ; anthers sessile or on fila- ments, erect, 2-celled, opening by slits or pores, rarely 4- celled. — Female and hermaphrodite : Staminodia 8 or many, few or variously combined. Ovary 2-12-celled ; stigma broadly peltate, entire or variously lobed ; ovules solitary, erect. Berry coated. Seeds enclosed in pulp. Tropical trees, with yellow juice; some of them yield the gamboges of commerce. Leaves leathery. Flowers terminal or axillary. — A species 10-12 feet high, with cream-coloured flowers, and beautiful dark green, glossy, acute leaves, which (if new) may be called G. Gerrardi, has recently been found in the Umvote district, Natal, by Mr. Gerrard (n. 1181). 27 Order XXII. MALVACEAE. Flowers bisexual, regular, mostly conspicuous. Calyx 5-fid, with valvate aestivation, mostly involucelled at base. Petals 5, strongly twisted in bud, withering and persistent, attached to the base of the staminal column. Stamens many, united in a tube, concealing the ovary and styles ; anthers reniform,l-celled. Carpels 3-5 or many, free or combined in a several-celled ovary. Fruit dry or fleshy. Seeds with a curved embryo, plaited cotyledons, and little or no albumen. — Leaves alter- nate, simple, palmately nerved, often lobed, stipuled. Tribe 1. Malvea:. Staminal column bearing anthers at or to the very summit. Carpels separable at maturity. Stigmas as many as the carpels. (Gen. 1-6.) Styles filiform, stigmatose along the upper side ; calyx furnished with an involucel. Involucel of 6-9 bracteoles 1. Althea. Involucel of 3 bracteoles 2. Malva. Styles capitellate, the stigmatic tissue confined to the apex. Calyx with an involucel. Ovules solitary. Involucel 3-bracted 3. Malvastrum. Ovules 2-3 in each carpel. Involucel 3-leaved or 3-fid 4. Sph^ralcea. Calyx nude at base. Ovules solitary 5. Sida. Ovules 3 or more in each carpel 6. Abutilon. Tribe 2. Urenea:. Staminal column bearing anthers on its outer surface, the apex with none, protruding, truncate or 5-toothed. Carpels separable at maturity. Stigmas twice as many as the carpels (10), capitellate at the summit. (Gen. 7.) Involucel 5-20-leaved 7. Pavonia. Tribe 3. Hibiscea:. Staminal column bearing anthers on its outer sur- face, the apex nude, protruding, truncate or 5-toothed. Carpels united into a several-celled capsule. Stigmas as many as the carpels (3-5), capitate. (Gen. 8-10.) Ovary 3-celled ; styles 3 ; bracteoles 3 or many ... 8. Fugosia. Ovary 5-celled. Involucral bracteoles 5 or many, separate .... 9. Hibiscus. Involucral bracteoles united in a toothed cup . . .10. Paritium. Tribe 1. Marvel. (Gren. 1-6.) 1. ALTHJEA, Linn. Involucel of 6-9 bracteoles, united at base. Staminal co- lumn antheriferous at the summit. Ovary of many carpels ; ovules solitary ; styles as many as the carpels, stigmatose 28 XXII. MALVACEJE. along the inner face. Fruit of many reniform, dry, 1-seeded, separable carpels. — FI. Gap. i. p. 158. Annuals or perennials, natives chiefly of the northern hemisphere. A. Ludwigii, Linn., a hispid, much-branched annual, with nearly glabrous, deeply 5-lobed leaves, and axillary white flowers, occurs at the Orange River. A. Burchellii , DC., an endemic species, is unknown to me. 2. MALVA, Linn. Involucel 3-bracted, persistent. Staminal column bearing anthers at the multifid summit. Ovary of many carpels ; styles as many as the carpels, stigmatose along the inner face ; ovules solitary. Fruit of many dry, 1-seeded, hard- shelled, indehiscent, separable carpels ; seed filling the cavity. — FI. Cap. i . p. 159. Herbs, natives of temperate climates. M. parvijlora, Linn., a prostrate procumbent plant, with long-petioled, reniform, bluntly 5-7-lobed leaves, and small, clustered, subsessile, axillary flowers (a European weed), occurs on roadsides and in waste places. 3. MALVASTRUM, A. Gray. Involucel 3-bracted or wanting. Styles capitellate. Other characters as in Malva. — FI. Cap. i. p. 159. Herbs or small shrubs, known from Malva by the capitate stigmas. Leaves lobed, stipuled. Flowers axillary or racemose. About 15 South African species, dispersed. 4. SPHJERALCEA, St. Hil. Involucel of 3 (or 2) narrow bracteoles, or monophyllous and 3-fid. Staminal column and stigmas as in Malvastrum. Ovules 2-3 in each carpel. Fruit of many compressed, 2- valved, dehiscing, 2-3-seeded carpels, separating at maturity from a central torus. — FI. Cap. i. p. 165. Also Sphseroma, Harv. 1. c. p. 166. Herbs or shrubs, with the habit of Malvastrum. — 4 South African species, natives of the Western and Northern districts ; 2 have 3-leaved and 2 have 3-fid involucels, the latter constituting the subgenus Sphceroma. 5. SIDA, Linn. Involucel 0. Staminal column antheriferous at its mul- tifid summit. Ovary of 5-15 carpels ; styles 5-15 ; stigmas capitellate ; ovule solitary, pendulous. Fruit of 5-15 1-seeded carpels, dehiscent at the summit and at length separating ; seed 3-cornered, suspended. — FI. Cap. i . p. 166. An immense tropical or subtropical genus, very various in aspect. 5 spe- cies in the Eastern district and at Natal, of which 2 are endemic, the others subtropical weeds. — Leaves entire or lobed, often penninerved. Flowers small, yellow or white. XXII. MALYACEJE. 29 6. ABUTILON, Tourn. Ovules 3 or several in each carpel. Fruit of numerous membranous, 3- or more-seeded carpels, opening by the ven- tral suture and subpersistent. Other characters as in Sida. — FI. Cap. i. p. 168. 2 (or more) species in the Eastern districts of Natal. Herbs or shrubs, often with very soft, velvety, cordate, subentire leaves. Tribe 2. Urexe^;. (Gen. 7.) 7. PAVONTIA, Cav. Involucel 5-15-leaved, persistent. Staminal column naked and 5-toothed at the apex, bearing stamens on its outer sur- face. Ovary of 5 carpels, united round a central axis ; styles confluent below, 10-cleft above ; stigmas 10, capitate. Fruit of 5 indehiscent, 1-seeded carpels. — FI. Cap. i. p. 169. Shrubs or herbs, chiefly tropical or subtropical, variable in habit. — 2 spe- cies from the Eastern district and 4 from Natal. Tribe 3. HiBiscEiE. (Gen. 8-10.) 8. FUGOSIA, Juss. Involucels 3 or several, often small or deciduous, sometimes tooth-like. Calyx 5-fid. Staminal column naked and 5- toothed at the apex, bearing stamens on its outer surface. Ovary 3-4-celled, cells 3- or many-ovuled ; style club-shaped at the summit, 3-4-furrowed, or divided in 3-4 short erect lobes. Capsule 3-4-valved. — FI. Cap. ii. p. 587. Shrubs or half-shrubs, chiefly subtropical, with the habit of Hibiscus. Leaves entire or lobed, rarely parted. Flowers mostly yellow. Calyx and petals often black-dotted. — 2 South African species : F. Gerrardi , found by Mr. Gerrard near Ladismith, Natal ; F. triphylla , in Damaraland, by Mrs. Kolbe. 9. HIBISCUS, Linn. Involucel 5-20-leaved. Petals expanded. Staminal column 5-toothed at the apex, bearing stamens on its outer surface. Ovary 5-celled ; style 5-cleft ; stigmas 5, capitate ; ovules nu- merous. Capsule 5-celled, 5-valved, loculicidal, enclosed in the persistent calyx. — FI. Cap. i. p. 170. A large and varied tropical and subtropical genus, usually with large, handsome, bright-coloured flowers. Leaves entire or lobed, sometimes multipartite, often stellate-pubescent. — 25 species, many of them endemic, mostly natives of the Eastern district and Natal. H. JSthiopicus, Linn., is found throughout the colony ; and H. Trionum , Linn., an equally dispersed (annual) weed of cultivation. 30 XXIT. MALVACEAE. 10. PARITIUM, A. Juss. Involucel monophyllous, 10-12-toothed or cleft. Petals and staminal column as in Hibiscus. Ovary 5-celled, each cell partially divided into two by a spurious parietal dissepiment ; ovules numerous. Capsule 5-celled, the cells imperfectly 2- locular. — FI. Cap. i. p. 177. Tropical trees, mostly growing near the sea. Leaves cordate, entire or lobed ; stipules broad, ovate. Flowers yellow, with a dark centre. — P. tiliaceum occurs at Natal, on the coast. Order XXIII. STERCULIACEiE. Plowers regular, hermaphrodite or unisexual. Calyx mono- phyllous, mostly persistent, 5-(rarely 3-4-7- )fid, valvate in bud. Petals 5, free or attached to the staminal tube, twisted, or 0. Stamens monadelphous, either merely united at base into a ring, or combined in a longer or shorter tube ; sta- minodia often alternating with stamens ; anthers 2-celled, opening outwards. Ovary free, of 1-2-5 carpels, more or less united; style entire or splitting into as many branches as there are carpels ; ovules 2 or many. Fruit dry, capsular or rarely indehiscent. — A large and varied Order, known from Malvaceae by its 2-celled anthers. Leaves alternate, stipulate, simple or digitate. Inflorescence axillary or terminal, race- mose or cymoso-paniculate, or reduced to solitary flowers. Petals very frequently persistent, withering. Tribe 1. Sterculie.®:. Flowers unisexual. Calyx mostly coloured. Petals 0. Anthers 5-15, on the apex of a long or short column. Ripe carpels free, sessile or stipitate 1. Sterculia. Tribe 2. Dombeye.®. Flowers bisexual. Petals often persistent, flat. Stamens (in the Cape genera) united at base into a cup, alternating singly or in parcels of 2-3, with 5 strap-3haped staminodia. (Gen. 2-3.) Bracts deciduous. Fertile stamens 2-3 2. Dombeya. Bracts persistent. Fertile stamens 5 5. Melhania. Tribe 3. Hermannl®. Flowers bisexual. Stamens 5, connate at base ; no staminodia. (Gen. 4-6.) Ovary 1-celled, unequal-sided ; style lateral 4. Waltheria. Ovary 5-celled, many-seeded ; style central. Filaments broadly linear, oblong or obovate .... 5. Hermannia. Filaments abruptly dilated in the middle 6. Mahernia. Tribe 1. Sterculieje. 1. STERCULIA, Linn. Flowers unisexual, without petals. Calyx coloured, campa- nulate or tubular, 5-7-cleft or parted, the segments leathery. XXIII. STERCULIACEJ3. 31 — Male : Staminal column shorter or longer than the calyx, solid, bearing anthers at the 5-10- toothed apex ; anthers ad- nate, 2-celled, opening outwards. — Female : Staminal column adnate to the carpophore, the abortive anthers at the base of the ovary. Ovary stipitate, of 5 connivent or partially connate carpels ; styles more or less united ; ovules many. Follicles 5 or fewer, leathery or ligneous, few- or many-seeded. — FI. Cap. i. p. 178; Thes. Cap. t. 3. Trees, chiefly subtropical. S. Alexandria Harv., our only species, was found near Uitenhage by Dr. Alexander Prior. Tribe 2. Dombeye^e. (G-en. 2-3.) 2. DOMBEYA, Cav. Involucre 3 -leaved, unilateral, deciduous, sometimes 0. Calyx 5-parted, at length reflexed, persistent. Petals 5, obovate, unequal-sided, convolute in bud, flat when expanded, persistent. Stamens united at base, 5 imperfect (stammodia), strap-shaped or filiform, 10-15 fertile, antheriferous, alternat- ing with the sterile in parcels of 2-3 ; anthers oblong, erect. Ovary sessile, 3-5-celled ; style 3-5-cleft, with revolute arms, stigmatose above ; ovules 2-4 in each cell. Capsules leathery, 3-5-celled. — FI. Cap. i. p. 220 ; Thes. Cap. t. 89, 137-138. Subtropical shrubs and small trees, chiefly African. Leaves cordate or lobed, many-nerved. Flowers rosy or white, in umbels or corymbs, the petals somewhat enlarged after flowering, and drying to a papery consist- ence. 6 South African species, natives of the Eastern district, Caflraria, and Natal. 3. MELHANIA, Forsk. Involucre 3-leaved, persistent. Calyx 5-parted. Petals 5, obovate, unequal- sided, convolute in bud, erect. Stamens united at base, 5 sterile, strap-shaped, 5 alternate fertile, shorter ; anthers sagittate, erect. Ovary 5-celled ; style 5-fid at the summit; ovules numerous. Capsules 5-celled. — FI. Cap. i . p. 221. Small, softly hairy, half-ligneous or herbaceous plants. Leaves ovate or oblong. Peduncles axillary, 1-2-flowered. — 4 South African species, natives of the Eastern district and Natal. Tribe 3. Hermannie^j. (Gren. 4-6.) 4. WALTHERIA, Linn. Calyx 5-cleft, 10-nerved, with or without a 3-leaved lateral involucel. Petals 5, oblong, with slender claws. Stamens 5, opposite the petals ; filaments united into a tube at base ; 82 XXTIT. STERCULIACEjE. anthers oblong, erect, 2-celled. Ovary oblique, 1-celled, 2- ovuled ; style lateral. Capsules 1-celled, 2-valved, 1-seeded. — FI. Cap. i. p. 180. Herbs or shrubs, chiefly tropical. W. Indica , Linn., a widely-diffused species, occurs at Magalisberg. 5. HERMANNIA, Linn. Calyx 5-cleft, often inflated. Petals 5, with hollow claws, spirally twisted in bud. Stamens 5, opposite the petals ; fila- ments connate at base, broad, flat, oblong or obovate ; anthers erect, taper-pointed. Ovary shortly stipitate, 5-celled ; styles coalescing, separable. Capsules coriaceous, 5-celled, 5-valved, many-seeded, simple or crested at the summit. — FI. Cap. i. p. 180. Small shrubs or undershrubs, almost all South African. Pubescence stellate, woolly, velvety, glandular or scanty. Leaves entire, toothed or pinnatifid, often plaited. Peduncles axillary or subterminal, pseudo- racemose. Flowers yellow or orange, rarely creamy or white, often sweet- scented. — Upwards of 70 Cape species, dispersed. 6. MAHERNIA, Linn. Pil aments of the stamens suddenly dilated and mostly tu- berculated in the middle or cruciform. Other characters as in Hermannia. — FI. Cap. i. p. 207. Small plants, with the habit of Hermannia. Peduncles mostly 2-flowered, terminal or opposite the leaves ; pedicels slender, bracteolate at base. Flower nodding, red orange yellow or violet. — 33 (or more) South African species, dispersed. Order XXIV. TILIACEJE. Like SterculiacecB , but the stamens not connate or tubular at base, and usually numerous ; anthers 2-celled, opening in- wards.— Trees shrubs or herbs, with mostly stellate pubes- cence, and stipulate simple leaves. Flowers 4-parted ; outer stamens beaded, without anthers 1. Sparmannia. Flowers 5-parted. Fruit fleshy, of 1 -4 hard-shelled drupes 2. Grewia. Fruit a globose or oblong capsule, covered with hooked or straight, sharp and rigid bristles 3. Trifmfetta. Fruit a pod-like capsule, 2-5-valved, splitting, many- seeded 4. Corchortjs. 1. SPARMANNIA, Th. Sepals 4, lanceolate, pointless, deciduous. Petals 4, ob- ovate, spreading horizontally. Stamens very many, the outer XXIY. TILTACEJE. 33 sterile, bearded, ihe inner fertile, with nodose filaments. Ovary sessile, 4-celled ; ovules many ; style single ; stigma 5-toothed. Capsules globose or oblong, 4-celled, 4-valved, loculicidal, covered with rigid bristles. — FI. Cap. i. p. 223. Shrubs or small trees, with pubescent, long-petioled, 5-7-angled or lobed leaves, and umbellate white flowers. — 2 South African species : S. Africana , from the South-Eastern district, and S. palmata , from Caffraria and NataL 2. GEEWIA, Linn. Sepals 5, linear, leathery, coloured within, deciduous. Pe- tals 5, each with a nectariferous gland or pit at base, inserted at the base of a short columnar torus which supports the stamens and ovary. Stamens many ; filaments filiform ; an- thers roundish. Ovary 2-4-eelled ; style simple. Drupe 2-4- lobed, containing 2-4 hard-shelled, 1-2-celled, 1-2-seeded nuts. — FI. Cap. i.p. 224. Trees or shrubs. Leaves undivided, entire or serrulate, 3-7-nerved at base, often pale beneath. Pubescence stellate. Flower purple or yellow, in cy mules or solitary. Petals often shorter than the sepals. — 11 South African species, dispersed. 3. TRIUMFETTA, Linn. Sepals 5, linear, membranous, coloured, hooded at the point and dorsally mucronate, deciduous. Petals 5, without glands, inserted at the base of a short columnar torus, which supports the stamens and ovary. Stamens definite, 5-30 ; filaments thread-like ; anthers roundish. Ovary 2-5-celled, the cells divided by a false parietal vertical septum ; ovules in pairs ; style filiform ; stigma 2-5-lobed. Capsules subglobose, co- vered with straight or hooked prickles, 2-5-celled ; cells 1-2- seeded. — FI. Cap. i. p. 227 ; Thes. Cap . t. 52. Shrubs half-shrubs or herbs, common in tropical countries. Leaves variable in shape in the same species, entire or lobed, serrate, many-nerved, often with glands on the serratures. Flower small, yellow or orange, soli- tary or clustered. — 4 species occur in the Natal country 4. CORCHORUS, Linn. Sepals 4-5, ovate or lanceolate, unequal, deciduous. Petals 4-5, obovate, clawed, hypogynous. Stamens many, mostly indefinite. Ovary sessile or shortly stipitate, 2-5-celled ; ovules many ; style short ; stigmas 2-5. Capsules pod-like or roundish, 2-5-celled, 2-5-valved, loculicidal, many-seeded. Herbs or half-shrubs, chiefly tropical. Leaves alternate, serrate, the teeth sometimes bristle-pointed. Flowers yellow, axillary or opposite leaves. — 4 species, found at Natal and on the North-Eastern frontiers of the Cape colony. I) Order XXY. LINE.®. Flowers bisexual, regular. Sepals 5, rarely 4, free or united at base, imbricate. Petals as many, mostly twisted. Stamens as many (or twice or thrice as many) as petals,. connate in a ring at base ; anthers 2-eelled. Glands 5, entire or 2-fid, adnate to the staminal tube, sometimes obsolete. Ovary free, 3-5- celled ; ovules 1-2 in each cell, pendulous ; styles 3-5, distinct or more or less united ; stigma terminal. Fruit capsular or fleshy. Seeds with or without albumen. — Herbs shrubs or trees, mostly glabrous. Leaves alternate or rarely opposite, simple, entire or crenato-serrulate. Stipules present or ab- sent. Flowers racemose or cymose, or clustered or solitary, axillary. Petals blue yellow or white, rarely red, mostly very fugitive. Tribe 1. Line.®. Petals twisted, fugitive. Stamens as many as petals. Capsule opening through the septa. No stipules. Herbs or undershrubs 1. Linum. Tribe 2. Erythroxyle^e. Petals imbricate, each with a plaited scale at base inside. Stamens twice as many as the petals. Drupe fleshy, in- dehiscent. Shrubs 2. Erythroxylon. 1. LINUM, Linn. Sepals 5, entire. Petals 5, twisted, fugitive. Stamens 5, perfect, alternating with as many tooth-like abortive filaments. Styles 5, rarely 3, separate or connate below ; stigmas capitel- late or linear. Capsules spuriously 10-celled, 10-seeded ; seeds without albumen. — FI. Cap. i. p. 308. Small undershrubs or herbs, found throughout the temperate zone. Leaves alternate, quite entire, sessile, without stipules. Flowers yellow. — 4 Cape species, dispersed. 2. ERYTHROXYLON, Linn. Sepals 5-6, connate at base or free. Petals as many, im- bricate, deciduous, furnished at base on the inside with a plaited, mostly 2-lobed scale. Stamens 10-12, united at base into a short glandless or 10-glanded tube, often a little pro- longed beyond the insertion of the filaments. Ovary 3-, rarely 4-celled ; ovules 1-2 ; styles 3-4, distinct or more or less united at the apex, capitate or clavate. Drupe 1-celled, 1- seeded ; seed albuminous. — FI. Cap. i. p. 233. Shrubs or small trees, mostly glabrous. Leaves alternate, entire, pe- tioled. Stipules within the petiole. Flowers small, whitish, axillary. — 3 species, found near Natal. Order XXVI. MALPIGHIACEiE. Flowers bisexual, regular. Calyx 5-parted, imbricate, some or all of the segments 2-glanded at back. Petals 5, clawed, spreading. Disk expanded or often inconspicuous. Stamens 10 (5 sometimes abortive), connate at base. Ovary of 3 or 2 imperfectly united carpels, 3-2-lobed ; ovules solitary. Albu- men 0. — Shrubs, erect or climbing, with mostly opposite, sim- ple, mostly entire leaves, and corymbose or racemose, red or yellow flowers. Pubescence silky. Carpels each with a single dorsal wing 1. Acridocaepus. Carpels each 2- or several-winged. Styles 3. Petals fringed 2. Triaspis. Style 1. Petals sagittate-ovate 3. Tristellateia. 1. ACRIDOCARPUS, Guill. and Perr. Calyx 5-fid, minutely glandular or glandless. Petals un- equal, clawed, glabrous, subentire. Stamens 10, all perfect ; filaments distinct, short, rigid ; anthers large, cordato-lanceo- late, glabrous. Ovary 3-celled, hairy, each lobe with a dorsal wing ; styles 2, very long, divergent, filiform, flexuous, acute, coiled up in aestivation. Samaras 1-2, on an oblong recep- tacle expanding above into a straight or oblique wing, which is thickened along its upper margin. — FI. Cap. i. p. 231 ; Thes. Cap. t. 19. Trees or shrubs, erect or climbing. Leaves alternate or opposite, entire, glabrous or silky. Racemes or corymbs terminal or lateral. Flower yel- low.— 3 species in the Natal country. 2. TRIASPIS, Burch. Calyx short, 5-parted, without glands. Petals longer than the calyx, clawed, fringed. Stamens 10, fertile, unequal, con- nate at base and adnate to the stipe of the ovary. Carpels 3, each expanded into a wing at each side, and united into a 3-lobed, 6-winged, shortly stipitate ovary ; styles 3, glabrous, elongate, flat, acute. Samaras 3 or fewer, winged at the mar- gin, the wing shield-like, sometimes interrupted at the apex, commonly dorsally crested in the middle. — FI. Cap. i. p. 232. Mostly climbers. T. Jiypericoides, Bell., an erect shrub 3-4 feet high, with opposite, linear, glabrous leaves and rosy flower, was found by Bur- chell at Kosi Font, lat. 27° 52'. 3. TRISTELLATEIA, Thouars. Calyx 5-parted, with minute glands or none. Petals clawed, keeled externally, glabrous, sagittate- ovate. Stamens 10, fertile, connate at base, those opposite the petals longer. XXYT. MALPI GHIACEAS. 30 Ovary 3-lobed, the lobes many-crested dorsally ; style 1 (the other 2 reduced to papillae), slender, elongate. Samaras 3, many-winged ; the wings narrow, elongate, stellately-patent. — FI. Cap. ii. p. 591. Climbers, with opposite or quaternate leaves, the petioles often 2-glanded. Racemes terminal and lateral. Flowers yellow. — F. Madagascariensis, Poir., was gathered at Delagoa Bay by Commander Owen. Oedee XXYII. ZYGOPHYLLE^]. Flowers bisexual, regular or irregular. Calyx 4-5-parted. Petals 4-5, clawed, twisted, rarely 0. Stamens 8-10 ; fila- ments frequently furnished with a scale at base. Ovary sessile or on a short gynophore, furrowed or winged, 5-celled ; ovules axile, 2 or several ; style single, terminal, rarely 5 separate styles. Fruit capsular or fleshy. — Herbs or shrubs. Leaves opposite, mostly compound, stipulate. Flowers soli- tary, axillary or terminal, yellow or white, rarely red. Petals 4-5. Fruit thorny and tubercled, indeliiscent. Leaves pinnate 1. Tkibttlus. Fruit capsular, 4-5-angled, dehiscent. Filaments simple at base. Anthers linear. Ovary densely silky .... 2. Sisyndite. Anthers cordate. Ovary glabrous .... 4. Fagonia. Filaments each with an entire or bifid scale at base 5. Zygophyllum. Petals 0. Styles short, filiform. Stamens 10. Leaves simple, fleshy 3. Augea. Styles 5. Stamens 5. Leaves 3-foliate .... 6. Seetzenia. Tribe 1. Tribtjleaj. Seeds without albumen. (Gen. 1-3.) 1. TRIBULUS, Tourn. Calyx 5-parted, deciduous or persistent, imbricated. Pe- tals 5, spreading, longer than the calyx. Stamens 10; fila- ments subulate, the 5 opposite the calyx-lobes with a gland externally at base ; anthers cordate, introrse. Ovary sessile, in a short 10-lobed cup, hairy, 5-celled ; ovules 3-4 in each cell ; style short or 0, rarely filiform ; stigma large, 5-angled. Fruit depressed, 5 -angled, of 5 indehiscent, dorsally tubercu- lated, thorny or winged, spuriously plurilocular carpels, each loculus 1-seeded. Seeds without albumen. — FI. Cap. i. p. 352. Weeds and weed-like, diffuse or prostrate herbs. Leaves opposite, one usually much smaller than the other, abruptly pinnate ; leaflets in several pairs. Flowers yellow or white. — 2 species : one a common weed through- out the country, the other found at Springbokkell by Zeylier. XXVII. ZYGOPIIYLLEiE. 37 2. SISYNDITE, E, Mey. Calyx 5-parted, the lobes somewhat imbricate. Petals 5, at first short and truncate, at length oblong, longer than the calyx. Stamens 10, hypogynous ; filaments subulate, glabrous, equalling the calyx ; anthers linear, versatile, 2-celled, longi- tudinally slitting. Hypogynous scales 5, circling the ovary opposite the sepals. Ovary sessile, very hairy, 5-angled, 5- celled ; ovules solitary, erect ; style filiform, hairy, thickened into a clavate, 5-furrowed stigma. Fruit capsular, 5-lobed, the carpels compressed, ovate, acute, cross-furrowed, in all parts clothed with long golden-yellow hairs, at length separat- ing and opening by the ventral sutures. Seed compressed, erect ; testa membranous ; embryo without albumen ; cotyle- dons thick; radicle short, superior. — FI. Cap. i. p. 354 ; Thes. Cap. t. 120. S. spartea, E. M., is a broorp-like bush, found between Natvoet and the G-ariep, 2-3-chotomous and quite glabrous. Leaves opposite, minutely stipulate, pinnate, the petiole terete and like a twig, elongate ; leaflets mi- nute, subopposite, distant, in few pairs, Flowers large and handsome, axillary. 3. AUGEA, Th. Calyx 5-cleft, persistent, valvate. Petals 0. Hypogynous disk cup-shaped, membranous, 10-toothed, with subulate-seta- ceous teeth. Stamens 10, inserted between the teeth of the disk ; filaments very short, broad, trifid, the medial segment bearing an anther; anther fixed below the middle, oblong. Petaloid scales linear, white, bifid, outside the stamens and opposite them. Ovary angular, glabrous, 10-celled ; ovules pen- dulous ; style short, filiform ; stigma simple. Capsules 10- angled, 10-valved. Seeds solitary, without albumen. — FI. Cap. i. p. 355. A. Capensis, Th., is an annual, glabrous, fleshy, Karroo-land herb, with the aspect of Mesembryardhemum. Leaves connate, terete, obtuse, flattish above. Stipules short. Flowers axillary, solitary or 2-3 together. Tribe 2. Ztgophylle^e. Seeds albuminous. (Gen. 4-6.) 4. FAGONIA, Tourn. Calyx 5-parted, deciduous. Petals 5, clawed, longer than the calyx. Stamens 10, hypogynous, equal; filaments fili- form, naked at base, erect ; anthers cordate. Ovary sharply 5-angled, 5-celled ; style 5-anglecf, continuous with the ovary ; stigma acute. Capsules pyramidal, 5-sided, of 5 cocci, which fall away when ripe from a persistent axis. — FI. Cap. i. Add. p. 21*. Small herbs. F. Cretica , Linn., a procumbent, much-branched, glabrous XXVII. ZYU0141Y LLEyE. 38 or glandularly-pubescent plant, with 3-foliolate leaves and lilac-purple flowers with bright orange stamens, has been found in Namaqualand by Mr. Wyley. 5. ZYGOPHYLLUM, Linn. Calyx 4-5-parted, persistent or deciduous, imbricate. Pe- tals 4-5, clawed, twisted, imbricate. Stamens 8-10 ; filaments subulate, with an entire bifid or tripartite scale at base. Disk fleshy, 8-10-angled. Ovary 4 -5-angled or lobed, 4-5-celled ; ovules 2 or more ; style furrowed, continuous with the ovary ; stigma minute. Capsules 4-5-angled or winged, 4-5-celled, few- or several-seeded. — FI. Cap. i. p. 355. Small shrubs or half-shrubs. Leaves fleshy or membranous, simple or bifoliolate ; stipules membranous or spinous. Flowers solitary, white or yellow. — 25 Cape species. 6. SEETZENXA, E. Br. Calyx 5-parted, valvate. Petals 0. Stamens 5, opposite the calyx-segments; filaments subulate, naked; anthers introrse, 2-celled, subglobose-didymous. Ovary oblong, 5 -celled ; ovules solitary ; styles 5, terete, reflexed ; stigmas capitate. Cap- sules ovoid, 5-coccous, the cocci separating from a central 5-angled axis. — FI. Cap. i . p. 365. A woolly or glabrous, Clanwilliam, etc., half-shrub, with jointed branches, trifoliolate leaves, intrapetiolar stipules, and axillary, 1-flowered peduncles. Order XXVIII. GrERANIACEiE. Blowers bisexual, regular or irregular. Sepals 5, rarely fewer, mostly separate and imbricate, the upper ones some- times spurred. Petals 5 or fewer or 0, imbricate or twisted in bud. Torus commonly with 5 glands alternating with the petals, usually prolonged through the centre of the ovary. Stamens 5-10-15 or fewer by abortion ; filaments mostly connate at base, rarely free ; anthers versatile, 2-celled. Ovary 3-5-lobed, 3-5-eelled, rarely 2-celled ; carpels united to the axis below, above either prolonged into a beak, continued into a style, or beakless, with free or partially united styles, or crowned with sessile stigmas. Ovules 1, 2 or many, axile. Fruit rarely indehiscent. Seeds with or without albumen ; embryo straight or curved, often green. — Herbs or shrubs of various aspects. Tribe 1. Oxalide.®. Flowers regular. Petals convolute. Stamens 10, connate at base. Ovary beakless. Styles 5, separate ; stigmas capitate. Capsules many-seeded. Leaves compound, without stipules. Usually trifoliolate herbs 1. Oxalis. XXVIII. UEItANIACEJE. 39 Tribe 2. Geraniea:. Flowers regular or irregular. Sepals unequal, strongly imbricate. Stamens 5-7-10-15, monadelphous or polyadelphous. Ovary beaked, the beak prolonged into a style ; stigmas 5, linear. Carpels l-seeded, tailed. Leaves mostly .simple, variously cut or multifid, stipulate. Flowers regular. Stamens 15, in parcels of 3 each 2. Monsonia. Stamens 15, monadelphous 3. Sarcocaulon. Stamens 10, monadelphous 4. Geranium. Stamens 5, monadelphous 5. Erodium. Flowers irregular. Stamens 7 or fewer. Petals |, or the lower absent ; back sepal tubular at base, the tube connate with the pedicel 6. Pelargonium. Tribe 3. Balsamineje. Flowers irregular. Sepals coloured, very un- equal, the posterior spurred, two anterior very small or none. Petals hy- pogynous. Stamens 5, short. Ovary beakless ; stigma sessile. Capsules bursting with elasticity. Herbs 7. Impatiens. Tribe 1. Oxalidea;. 1. OXALIS, Linn. Sepals 5, free or united at base. Petals 5, convolute, their claws conniving into a funnel-shaped tube. Stamens 10, con- nate at base, 5 alternate shorter. Ovary 5-lobed, 5-celled, beakless ; ovules few or many ; styles 5, filiform ; stigmas capitate or pencilled. Capsules deeply 5-lobed, globose or oblong ; seeds 1 or several, albuminous. — FI. Cap. i. p. 313. A cosmopolitan genus, particularly abundant in the southern hemisphere. Leaves compound ; leaflets rarely 1 or 2, commonly 3, sometimes many and digitate. Flowers red purple white yellow or streaked. — There are (at least) 108 Cape species, all herbaceous, and most of them bulb-rooted ; they blossom in the winter and early spring months. Tribe 2. Gteraniea:. 2. MON SONIA, Linn. f. Sepals equal at base, mucronate. Petals spreading equally, longer than the calyx. Stamens 15, connate at base and spreading in 5 parcels, each of 3 stamens, whose filaments co- here for half their length. Ovary 5-lobed, beaked; ovules so- litary.— FI. Cap. i.p. 254. Annual or perennial, herbaceous or half-shrubby plants, with slender stems. Leaves simple, subentire, toothed, or deeply-lobed or cut. Pe- duncles 1-2, or umbellately several-flowered. — 8 Cape species, dispersed. 3. SARCOCAULON, DC. Sepals equal at base, mucronate. Petals spreading equally. Stamens 15, connate at base ; filaments subulate, not coher- ing in parcels. — FI, Cap. i. p. 256. 40 xxvni. geraniacejE. Divaricately-branched, fleshy or succulent, rigid shrubs, armed with spines formed from persistent and hardened petioles. Leaves obovate or obcordate, entire or crenate. Peduncles 1 -flowered. — 3 species: in the Northern, North-Western, and North-Eastern districts. 4. GERANIUM, Linn. Sepals equal at base. Petals spreading equally. Stamens 10, all perfect, the alternate longer. Glands at the base of the longer stamens. — FI. Cap. i. p. 257. A cosmopolitan genus. — 5 Cape species, all slightly suffruticose. Slender perennials, with palmately-lobcd or cut and multifid, long-petioled leaves. Peduncles 1-2-flowered, opposite the leaves or in the forks of the branches. 5. ERODIUM, L’Her. Sepals equal at base. Petals spreading equally. Stamens 5, perfect, bearing anthers ; 5 sterile, subulate or obsolete. Glands at the base of the sterile stamens. — FI. Cap. i. p. 259. Herbs, rarely undershrubs. Leaves simple, either pinnatipartite, pinna- tifid, cut or digitately parted. Peduncles mostly umbellately several- flowered. — 5 Cape species, of which 2 are endemip, 3 probably of European origin. 6. PELARGONIUM, L’Her. Calyx 5-parted, the upper segment produced at base into a slender nectariferous tube, which is decurrent along the pedicel and actuate to it. Petals 5, rarely but 4 or 2, more or less unequal. Filaments 10, unequal, monadelphous ; 2-7 bearing anthers, the rest abortive. A large genus, chiefly South African. — -About 163 Cape species, distri- buted under the following sections : — 1. Hoarea. Stemless, with tuberous roots. Petals 5 or 4. 2. Seymoitria. Stemless, with tuberous roots. Petals 2. 3. Polyactium. Caulescent, with tuberous roots. Leaves lobed or pinnatipartite. Umbels many-flowered. Petals subeqiial, obovate, entire or multifid. 4. Otidia.. Stem succulent and knobby. Leaves fleshy, pinnately or bipinnately cut. Petals subequal, the upper eared at base. 5. Ligtjlaria. Stem either succulent or slender and branching. Leaves rarely entire, mostly much cut or pinnatisect. Petals rather unequal, spa- thulate, the upper tapering at base. 6. Jenkinsonia. Shrubby or succulent. Leaves palmately-nerved or lobed. Two upper petals on long claws, very much larger than the lower. Stamens 7. 7. Myrrhidium. Slender, suffruticose or annual. Leaves pinnatisect. Petals 4 (rarely 5), two upper largest. Calyx segments membranous, strongly ribbed and mucronate or taper-pointed. 8. Peristera. Herbaceous, diffuse, annual or perennial. Leaves lobed or pinnatifid. Flowers minute. Petals scarcely longer than the calyx. (Habit of Geranium or Erodium.) 9. Campylia. Stem short, subsimple. Leaves on long petioles, undi- vided, entire or toothed. Stipules membranous. Flowers on long pedicels. XXVIII. GERANIACEAJ. 41 Two upper petals broadly obovate, three lower narrow. Fertile sta- mens 5, two of the sterile ones recurved. 10. Dibrachya. Much branched, with weak jointed stems. Leaves peltate or cordate-lobed, fleshy. Petals obovate. Stamens 7, the two upper very short. 11. Eumorpha. Slender, suffruticose or herbaceous. Leaves on long petioles, palmately 5-7 -nerved, reniform, lobed or palmatifid. Petals un- equal, the 2 upper broad. Stamens 7. 12. G-laucofhyllum. Shrubby. Leaves fleshy, simply or ternately compound, the lamina articulated to the petiole. Stamens 7. 13. Ciconium. Shrubby, with fleshy branches. Leaves either obovate or cordate-reniform, palmately many -nerved, undivided. Petals all of one colour, scarlet pink or white. Stamens 7, 2 upper very short. 14. Cortusina. Caudex short, thick, and fleshy ; branches (if any) slender and half-herbaceous. Leaves reniform or cordate, lobulate, on long petioles. Petals subequal, 2 upper broader. Stamens 6-7. 15. Pelarguum. Much-branched shrubs or undershrubs, not fleshy. Leaves entire or lobed (never pinnatipartite). Stipules free. Inflorescence frequently panicled, the partial peduncles umbellate. 2 upper petals longer and broader than the lower. Stamens 7. Tribe 3>. Balsamine^:. 7. IMPATIEHS, Linn. Blowers irregular. Sepals 3, rarely 5, coloured, imbricate, the two lateral flat, the two anterior, when present, small, the hinder one very large, produced at base into a hollow spur. Petals 3, the anterior concave, the lateral 2-fid. Glands 0. Stamens 5 ; filaments short, flat ; anthers conniving round the pistil. Ovary oblong, 5-celled ; stigma sessile, 5-toothed or lobed ; ovules many. Capsules 5-valved, the valves bursting with elasticity and falling off. Seed exalbuminous. — FI. Cap. i. p. 312. A large genus, chiefly from tropical Asia. — 1 Cape species (perhaps 2 ?), found in the Eastern district and at Natal. Order XXIX. RUTACEiE. Blowers bisexual (or rarely unisexual), mostly regular. Sepals 4-5, imbricate. Petals 4-5. Stamens inserted at the base or on the margin of the torus, as many or twice as many as the petals, rarely fewer or more numerous, free or rarely united into a tube. Disk between the stamens and ovary, annular or expanded. Carpels 4-5, rarely more or fewer, united into a 2-5-lobed, or solid ovary ; styles either free or united into one, long or short ; ovules 2 in each cell. Bruit various, capsular or berried ; seeds with or without albumen. —Trees or shrubs, rarely herbs, gland-dotted and strongly scented. Habit various. 42 XXIX. EUTACEiE. Leaves simple. Flowers bisexual, 5-parted. Ovary stipitate. Fruit a rough, 5-celled capsule 1. Calodendron. Ovary sessile. Fruit of 3-5 separate cocci. Staminodia 0. Petals clawed, bearded on the limb. Style short ; stigma capitate .... 2. EtrcH astis. Style long, much protruded 3. Macrostylis. Petals sessile, not bearded 4. Diosma. Staminodia 5, one between each stamen. Style short ; stigma capitate. Petals channelled, the staminodia infolded in the channel . . 5. Coleonema. Petals flat ; staminodia free. Petals clawed, the claw bearded ... 6. Acmadenia. Petals subsessile, broad, nude ... 7. Adenandra. Style as long as petals ; stigma simple. Petals sessile. Flowers axillary . . .8. Barosma. Petals clawed. Flowers terminal . . 9. Agathosma. Flowers unisexual, 4-parted ; carpels solitary. Petals 0. Leaves lanceolate, dotted .... 10. Emplettrum. Petals 4. Leaves acicular, not pellucid, dotted . 11. Empleurldium. Leaves trifoliolate 13. Toddalia. Leaves abruptly pinnate ; ovary lobed 12. Zanthoxylon. Leaves unequally pinnate ; ovary solid 14. Clausena. Teibe 1. Diosme.®. 1. CALODENDEON, Tb. Calyx short, 5-parted, deciduous. Petals 5, oblongo-lanceo- late, much, longer than the calyx. Stamens 10, inserted under a short, tubular disk, 5 fertile, 5 alternate (staminodia) sterile and petaloid. Ovary stipitate, shortly 5-lobed ; style filiform, elongate. Capsules stipitate, ligneous, roughly tu- bercled, 5-angled, 5-celled, septicidally 5-valved ; seeds 2 in each cell, angular. — FI. Cap. i. p. 371. C. Capense , Th., is a noble tree, a native of the Eastern district and Natal. Leaves decussate, petioled, ovate or ovato-lanceolate, acute or ob- tuse, pellucid-dotted, evergreen, 4-5 inches long. Flowers in terminal panicles ; petals white, with purple, glandular spots. The “ Wilde Kasta- nien ” of colonists. 2. EUCHJETIS, Bartl. and Wendl. Calyx 5-parted. Petals broadly clawed, oblongo-lanceolate, with a transverse beard. Stamens 5, fertile, shorter than the calyx ; anthers rounded, with an apical gland ; staminodia 0. Ovary deeply sunk in the cup-shaped, 5-lobed disk, 5-lobed ; style short ; stigma capitate. Fruit of 5 cocci, shortly horned at the summit. — FI. Cap. i. p. 371. Small, slender shrubs, with scattered, rarely opposite, lanceolate, keeled XXIX. BUTACE.fi. 43 leaves, and terminal, capitate or glomerate flowers. — 5 species from the Western and Midland districts. 3. MACROSTYLIS, B. and W. Calyx 5-parted. Petals clawed, spathulate, bearded in the middle. Stamens 5, fertile, exserted ; staminodia 0 ; anthers roundish, with a minute, apical gland. Disk closing over the ovary, perforated by the style. Ovary deeply 3-5-lobed; style lengthening after flowering, slender at base ; stigma ob- tuse. Cocci 3-5, horned. — FI. Cap. i. p. 441. Small bushes, with alternate or opposite, short, nerve-keeled leaves, pellucid-dotted along the margin and nerve. Flowers small, white or rosy, subumbellate at the end of the twigs. — 8 species, natives of the Western district. 4. DIOSMA, Linn. Calyx 5-parted. Petals sessile, obovate, longer than the calyx, nude (not bearded). Stamens 5, fertile, shorter than the petals ; staminodia 0 ; anthers roundish, with a sessile, apical gland. Disk fleshy, 5-lobed, cup-like. Ovary sunk in the disk, small, deeply 5-lobed ; style short ; stigma capitate. Cocci 5, longer than the calyx, rough, horned at the summit. — FI. Cap. i. p. 373. Small shrubs, with alternate or opposite, linear-acute, channelled, serru- late or ciliate, gland-dotted leaves ; and white or reddish, terminal, sub- solitary or corymbose flowers. — 11 species, all (except D. vulgaris , which is everywhere) found in the Western district. 5. COLEONEMA, B. and W. Calyx 5-parted. Petals obovate, twice as long as the calyx, tapering at base into a channelled claw. Stamens 5, fertile, equalling the sepals ; anthers roundish, tipped with a sessile gland ; staminodia 5, filiform, nude, enclosed within the channels of the petals. Disk cup-like, crenate. Ovary deeply 5-lobed ; style short ; stigma capitate. Cocci 5, compressed, rough, shortly horned at the summit. — FI. Cap. i. p. 377. Shrubs with scattered, linear leaves. Flowers axillary, solitary, white or purple. — 4 species, 3 of which are chiefly Western. 6. ACMADENIA, B. and W. Calyx 5-parted. Petals clawed, the claw bearded within (except in A. psilopetala) . Stamens 5, fertile, equalling the claws of the petals ; anthers ovate or oblong, tipped with an erect, sessile, conical gland ; staminodia filiform, short or ob- solete. Disk cup-like, entire or 5-crenate. Ovary 4-5-lobed, sunk in the disk ; style short ; stigma capitate. Cocci 4-5, compressed, horned at the apex. — FI. Cap. i . p. 379. 44 XXIX. llUTACEiE. Small shrubs, with imbricate (rarely scattered), linear, oblong or round- ish leaves. Flowers terminal, either solitary or few together, rarely in many-flowered heads. — 14 species, chiefly in the South-Eastern district. 7. ADENANDRA, Willd. Calyx 5-parted. Petals broadly obovate, with very short claws, nude. Stamens 5, fertile, shorter than the calyx ; an- thers oblong, erect, tipped with a stalked, spoon-shaped or globose, at length reflexed gland ; staminodia also tipped with a gland. Disk cup-like, 5-10-crenate. Ovary sunk in the disk, 5-lobed; style short ; stigma capitate, 5-crenate. Cocci obtuse or horned, glandularly-muricate above. — FI. Cap. i. p. 384. Yirgate or much-branched shrubs, with scattered, rarely opposite, pellucid-dotted leaves, and terminal, sessile or pedicellate flowers, wThich are larger and handsomer than in the allied genera. — 21 species, natives of the Western and South-Eastern districts. 8, BAROSMA, Willd. Calyx 5-cleft or parted. Petals much longer than the calyx, oblong, subsessile. Stamens 5, fertile, longer than thepetaloid or filiform staminodia, which alternate with them ; anthers ovoid, glandless or with a minute apical gland. Disk cup-like, entire or lobed. Ovary 5-lobed ; style long, filiform ; stigma simple. Cocci eared at apex, gland-xlotted. — FI. Cap. i. p. 392. Small shrubs, with mostly opposite leaves, gland-serrated. Flowers on axillary twigs, solitary or tufted ; very rarely subumbellate and terminal. — 15 species, among which is B. crenulata , Hook., the true “ Buku,” though others are indiscriminately collected for it, particularly B. serratifolia , W. 9. AGATHOSMA, Willd. Calyx 5-parted, rather unequal. Petals longer than the calyx, clawed. Stamens 5, fertile, alternating with as many filiform or petaloid staminodia ; anthers subglobose. Disk cup-like, crenulate or lobed. Ovary 2-4-lobed; style long, filiform ; stigma simple. Cocci mostly 3, horned. — FI. Cap. i. p. 399. A large genus of small shrubs, with alternate, rarely opposite leaves. Flowers at the ends of the branches, capitate or umbellate; in one species axillary. Petals white red or lilac-purple. — 100 (or more) species, dis- persed. 10. EMPLEURUM, Soland. Plowers monoecious. Calyx 4-cleft. Petals and disk 0. Stamens 4 ; anthers large, 4-sided, the cells divergent and prolonged beyond the sessile, apical gland. Ovary of 1, rarely 2 carpels ; style short ; stigma simple. Fruit lanceolate, compressed, tapering upwards into a beak. — FI. Cap. i. p. 441. XXTX. RUTACE/E. 45 E. serrulatum , Ait., is a 2-3 feet high shrub, with close-set, lanceolate, 1-2 inches long, gland-serrate leaves. Male and female flowers on different branches, axillary. It is found as far east as TJitenhage, at least. 11. EMPLEURIDIUM, Sond. Flowers dioecious. Calyx 4-parted, persistent ; the sepals acute, imbricate. Petals 4, deciduous, sessile, ovate-rotund, inserted under the edges of a fleshy, 4-lobed disk. — Male : Stamens 4, on the margin of the disk ; filaments subulate, short ; anthers didymous, glandless. An abortive ovary. — Female : Ovary (not seen) . Capsules oblong, follicular, open- ing at the side, and tipped with a short, persistent style ; seed solitary. — FI. Cap. i. p. 442 ; Thes. Cap. t. 77. E. juniperinum , S. and H., is a small undershrub, found by Ecklon, near Caledon. It is not gland-dotted in any part. Leaves scattered, acicular, 6-12 lines long, 4 line wide, scabrous on the margin and keeL Flowers axillary, very minute. Tribe 2. Zanthoxylea. 12. ZANTHOXYLON, Linn. Flowers polygamous. Calyx 4- (3-5-) parted, small. Petals hypogynous, as many as the calyx lobes, imbricate. — Male : Stamens as many as petals, alternating with tbem ; filaments free, subulate. A rudimentary ovary. — Female : Stamens 0 or abortive Carpels 1-5 on a fleshy disk, separate or sub- coherent ; ovules 2 in each carpel ; Styles terminal, cylindri- cal, short or long ; stigma capitate. Capsules leathery, 1-5, sessile or stipitate, 2-valved, 1-2-seeded; seeds black and shining. — FI. Cap. p. 445. Trees and shrubs of both hemispheres, armed with very large and strong thorns on the branches and stem, and often with prickles on the petioles and leaves. Leaves in our species abruptly pinnate, dotted. Flowers small, panicled. — 3 Cape species, all Eastern. Tribe 3. Toddaliea. 13. TODDALIA, Juss. Flowers polygamous. Calyx short, 2-5-toothed, lobed or parted. Petals 2-5, imbricate or valvate. Torus inconspicu- ous or slightly elongate. — Male : Stamens 2, 4, 5 (or 8, those opposite the petals abortive), inserted at the base of the torus ; filaments subulate ; anthers oblong. A rudimentary, simple, or 4-lobed ovary. — Female : Ovary ovoid, oblong or globose, sessile or substipitate, 2-7 -celled, very rarely 1-celled ; style short or 0 ; ovules 2. Fruit leathery or fleshy, dotted, sub- globose, 2-7-celled. — FI. Cap. i. p. 446. Also Yepris, Comm., FI. Cap. i. p. 447. XXIX. RUTACEJE. 16 Shrubs, unarmed or aculeate. Leaves alternate, 3-foliolate, dotted. Flowers small, in cymes or panicles. — 3 species, 1 from the Eastern district, 2 irom Natal. Tribe 4. Attrantieas. 14. CLAUSENA, Burm. Calyx 4-5-lobed or parted. Petals 4-5, free, mostly deli- cate, elliptical or roundish, imbricate. Stamens 8-10, free, the alternate shorter ; filaments dilated at or below the middle, and often concave, subulate above ; anthers short. Disk stipe-like. Ovary 4-5-(rarely 2-3-)celled, stipitate ; style mostly distinct, at length deciduous ; stigmas obtuse, entire or lobed ; ovules 2, collateral or superposed. Berry ovoid, oblong or globose, 2-5-celled (or abortively 1-celled), few- or 1-seeded. Seed with a membranous coat, and no albumen. — FI. Cap. i. p. 444 ( under Myaris, Fr.). Trees chiefly Asiatic. Leaves imparipinnate ; leaflets membranous, en- tire or crenulate, sometimes oblique. Panicles terminal or axillary ; flowers small, white. C. inrsqualis, Oliv., our only species, is common in the Eastern district and at Natal. Order XXX. OCHNACEiE. Blowers bisexual, regular. Sepals 4-5-6, rigid, persistent, imbricate. Petals as many, rarely twice as many, deciduous, sessile. Torus enlarged after flowering, never annual or glan- dular. Stamens definite or indefinite ; filaments short ; an- thers long, erect, opening by pores or slits, hard and dry. Ovary 2-10-lobed (or elongate, 1-10-celled) ; style simple, subulate. Bruit of 2 or more drupes (or capsular).— Trees or shrubs, with watery (not resinous) juice. Leaves alternate, stipulate, glabrous, coriaceous, shining, mostly serrulate. 1. OCHNA, Schreb. Sepals 5, coloured, persistent, imbricate. Petals 5-10, ob- ovate or oblong, deciduous. Torus thick, lobed or elevated in the centre. Stamens many ; anthers linear, basifixed, opening by short or long, terminal pores. Ovary deeply 3-10-lobed ; styles connate, or partly free, central. Drupes 3-10, or fewer, sessile on the enlarged torus. — FI. Cap. i. p. 448. Chiefly tropical trees or shrubs. Flowers yellow ; the calyx after flowering red or vinous-purple, brightening as the fruit advances. Leaves simple, shining, serrulate or subentire. — 3 species, all natives of the Eastern district and Natal. Order XXXI. BURSERACEiE. Flowers perfect or polygamous. Calyx 3-5-fid or parted, imbricate or valvate. Petals 3-5, deciduous. Disk annular or cup-like, free or adnate to the calyx tube. Stamens mostly twice as many as petals, on the margin or at base of the disk ; anthers subglobose or oblong, versatile. Ovary 2-5-celled, often with a short style; ovules 1-2 in each cell. Fruit drupaceous, indehiscent, 2-5-celled. — Trees or shrubs, balsami- ferous or oily. Leaves 3-foliolate or pinnate, rarely opposite, without stipules ; leaflets very rarely pellucid- dotted. Flowers small, racemose or panicled. Flowers on very short, 1-flowered peduncles . . 1. Balsamodendron. Flowers many, in a long peduncled panicle ... 2. Protium. 1. BALSAMODENDRON, Kth. Flowers polygamous. Calyx urqeolate or tubular, 4-toothed, persistent. Petals 4, suberect, linear-oblong, induplicate- valvate. Disk erect, cup -like. Stamens 8, on the margin of the disk, erect, free ; the alternate shorter. Ovary girt by the disk, 2-3-celled, tapering into a short style ; stigma 4-lobed. Drupe ovoid or globose, 1-3-celled. — FI. Cap. i. p. 526. Trees or shrubs. Leaves unequally pinnate ; leaflets 3-5, sessile, dot- less. Flowers on very short (1-2 lines long) peduncles, solitary or tufted. B. Cajpense , Sd. ; on the North-Western frontier, near the Gariep. 2. PROTIUM, W. and A. Calyx small, cup-like, 4-6-fid or toothed, valvate. Petals 4-6, erect or spreading, linear-oblong, valvate. Disk urceolate, covering the calyx-tube, margin free. Stamens 8-12, inserted under the margin of the disk, unequal, erect, free. Ovary girt at base by the small disk, ovate, 2-4-celled, tapering into a short style ; stigma 3-4-lobed. Drupe fleshy, globose, 1-4- celled. — FI. Cap. ii. p. 592. Small, halsamineous trees. Leaves toward the end of the twigs, 3-folio- late or imparipinnate ; leaflets in few pairs, entire or denticulate. Panicles on long peduncles, diffusely branched. Flowers small. — Readily known from Balsamodendron by its inflorescence. P. Africanum, H., found near Durban, Natal, by Gerrard and M'Ken. Order XXXII. MELIACEiE. Flowers regular, mostly perfect. Calyx short, 4-5-fid or parted, imbricate. Petals 4-5, longer than the calyx, twisted or imbricate, sometimes connate and valvate. Stamens 8-10 (rarely 5-16-20) ; filaments inserted outside a fleshy disk, 48 XXXIT. MELIACE7E. more or less united in a tube, which is entire or toothed at the apex ; anthers sessile or subsessile on the staminal tube, included or exserted, erect, 2-celled. Disk various. Ovary free, 3-5-celled ; style simple ; stigma peltate. Ovules 2, collateral. Fruit a capsule, drupe or berry. — Trees or shrubs. Leaves alternate, exstipulate, mostly pinnatipartite, in Tur- roea simple. Leaves simple. Petals and staminal tube both very long 1. Turba:a. Leaves simply pinnate. Fruit a dry, splitting capsule 2. Tbichilia. Fruit a leathery, nearly juiceless, indehiscent, 2-5-celled berry 3. Ekebeegua. Leaves doubly pinnate. Fruit a drupe 4. Melia. 1. TURRiEA, Linn. Calyx cup-shaped, 5-toothed. Petals 5, very long, strap- shaped, convolute in bud. Stamens 10, connate in a long tube, 10-toothed at the summit ; anthers sessile between the teeth. Ovary sessile, 5-10-20-celled ; style 1 ; stigma thickened. Capsules 5-celled, cells 2-1-seeded, the valves septiferous. Seeds compressed. — FI. Cap. i. p. 244. Shrubs, with alternate, simple leaves. Flowers on short twigs, solitary or tufted. — 2 South African species, natives of the Eastern frontier and Natal. 2. TRICHILIA, Linn. Calyx short, 4-5-toothed or cleft. Petals 4-5, erect or spreading, imbricate. Staminal tube 8-10-cleft or 8-10-parted, rarely entire, the segments linear, entire or 2-toothed, bearing anthers between or on the teeth ; anthers erect, exserted. Disk annular. Ovary sunk in the disk, 2-3-celled ; style long or short ; stigma 2-3-lobed. Capsules coriaceous, 2-3-celled, 2-3-valved. — FI. Cap. i. p. 246 ; Thes. Cap. t. 76. Trees or shrubs, chiefly American. Leaves pinnate. Flowers in axillary panicles. — 3 South African species, all found near Natal. 3. EKEBERGIA, Sparm. Calyx short, 4-5 -fid ; the lobes obtuse, imbricate. Petals 4-5, scarcely longer than the calyx, elliptical or oblong, imbri- cate in bud. Stamens 10, united in a short, campanulate, 10-toothed tube ; the teeth bearing anthers. Ovary 4-5-celled, girt by an annular disk ; style short, thick ; stigma obsoletely lobed. Berry dry, leathery, globose, 4-5-celled, 1-5-seeded. Seeds arillate. — FI. Cap. i. p. 246. E. Capensis , Sp., the only species, is a handsome, ash-like tree, native of the Eastern district, Caflraria, and Natal. XXXTI. MELIACE2E. 49 4. MELIA, Linn. Calyx small, 5-fid. Petals 5, linear- oblong, spreading, con- volute in bud. Stamens 10, the filaments connate into a 20- toothed tube, the anthers sessile within the throat of the tube. Ovary on a raised torus, 5-celled ; style filiform ; stigma 5-angled. Drupe with a 5-furrowed and 5-celled bony stone. — FI. Cap. i. p. 245. M. AzedaracJi, Linn., the “ Cape Lilac,” or “ Pride of China,” is cultivated throughout the colony, and partly naturalized. Order XXXIII. CHAILLETIACEAD. Elowers bisexual or unisexual. Sepals 5, free or connate, coriaceous, imbricate. Petals 5, inserted at the base of the ca- lyx, and rather longer, free and equal or connate and unequal, with broad claw and narrow 2-fid limb, tipped by an indexed process, which is connate with the margins of the lobes of the petals. Stamens 5, inserted with the petals, alternating with the lobes of the disk or with hypogynous glands ; anthers ob- long, 2-celled. Hypogynous glands 5, scale-like. Ovary free, 2-3-celled ; ovules in pairs, pendulous ; styles 2-3, free or partly united. Pruit drupaceous. — Trees or shrubs, chiefly tropical. 1. CHAILLETIA, DC. Calyx 5-parted. Petals separate, broad-clawed, deeply 2- parted or 2-fid. Stamens 5, equal ; anthers oblong, the con- nective thickened. Hypogynous glands 5, opposite the petals, distinct or united in a 5-lobed disk. Ovary subglobose ; styles 1-3, free or connate, short or long and slender. Drupe leathery, dry, 1-2-celled. — FI. Cap. i. p. 450. Chiefly tropical. C. cymosa, Hk. Ic. t. 591, our only South African spe- cies, is a very dwarf, little branched, leafy, villous shrub. Leaves 3-4 inches long, 6-10 lines wide, alternate, crowded, narrow-oblong, obtuse, glabrous and netted- veined. Cymes shorter than the leaves. Found at Aapjes river, by Burke and Zeyher. Order XXXIY. OLACINE^]. Plowers regular, perfect or unisexual. Calyx small, 4-5- toothed, lobed or parted. Petals 4-5, free or connate in a monopetalous corolla, valvate or minutely subimbricate. Sta- mens 4-10, inserted with the petals and often more or less adnate to them ; filaments mostly free ; anthers 2-celled. Disk annular or 4-5-divided. Ovary free, 1-celled (or spuri- E 50 XXXI Y. OLACINEJE. ously 3-5-celled) ; ovules 2-4, rarely 1, pendulous from the summit of a thread-like, free, central placenta, or attached to the sides of the ovary, or of the false septa ; style simple. Fruit in the unaltered or enlarged calyx, 1-celled, 1-seeded. Seed with much albumen. Petals 4-5, hairy within. Stamens 8-10 1. Ximenia. Petals 4-5, glabrous. Stamens 4-5 2. Apodytes. Corolla rotate (monopetalous), bearing the stamens . . 3. Cassinopsis. 1. XIMENIA, Linn. Calyx small, 4-5-toothed, unchanged in fruit. Petals 4-5, hypogynous, valvate, narrow, bearded within. Stamens twice as many ; anthers linear, erect, opening by opposite slits. Ovary 3-celled at base; style simple; ovules 3, linear, pen- dulous from a central placenta, which is free at summit, or at- tached to the wall of the cavity. Drupe fleshy. — FI. Cap. i. p. 235 ; Thes. Cap. t. 126. X. Capra , Sond., our only South African species, grows at Magalisberg and Natal. 2. APODYTES, E. Mey. Calyx small, 4-5-toothed, unchanged in fruit. Petals 4-5, hypogynous, free, valvate, glabrous. Stamens 4-5, alternate with the petals and slightly attached to them at base ; fila- ments thickish ; anthers oblong or linear, dorsally affixed. Ovary 1-celled, often thickened on one side at the apex ; style oblique or excentric ; ovules 2, superposed, pendulous from an adherent placenta. Drupe fleshy, oblique, compressed, with a protuberance on one side. — FI. Cap. i. p. 235. Trees or shrubs. A. dimidiata , E. M., the only Cape species, occurs chiefly in the Eastern district, Caffraria, and Natal. It turns blackish in drying. Leaves ovate-oblong, glossy above, very entire. Flowers minute, in terminal, much-branched panicles. 3. CASSINOPSIS, Sond. Calyx 5-fid. Corolla rotate, 5-fid, the segments oblong, slightly imbricate in bud. Stamens 5, inserted at the base or in the throat of the corolla, alternating with its lobes ; fila- ments subulate ; anthers oblong-cordate, 2-celled. Disk 0. Ovary sessile, 1-celled, 2-1-ovuled ; ovules superposed, pendu- lous from the apex of the cavity. Drupe nearly dry, ovato- globose. Seed inverted, compressed ; embryo minute, in the apex of copious albumen. — FI. Cap. i.p. 473 ; Thes. Cap. t. 168. Shrubs or small trees, with opposite, entire or toothed, petioled leaves, and axillary cymes of minute flowers. — 2 South African species : C. Capensis, Sond., found throughout the Eastern district and in Caffraria ; C. tinifolia, H., found in Zululand. 51 Order XXXV. ILICINE.®. Flowers regular, bisexual. Calyx 3-6-parted, imbricate. Petals 4-5, rarely more, free or combined in a rotate corolla, bypogynous, deciduous, imbricate. Stamens as many as the petals, free or attached to the base of the corolla ; filaments subulate ; anthers introrse. Disk 0. Ovary free, 3-5- or many-celled ; style 0 or terminal ; ovules 1-2, pendulous. Fruit a fleshy drupe, containing 3-18 bony, 1-seeded cells. — Trees or shrubs, mostly evergreen, with shining leaves. 1. ILEX, Linn. Flowers perfect. Calyx small, 4-6-toothed, persistent. Corolla rotate, 4-6-parted, the segments obtuse, imbricated. Stamens alternate with the lobes of the corolla ; filaments su- bulate ; anthers introrse, erect. Ovary sessile, 4-6-celled ; ovules 1-2, pendulous ; stigmas 4-6, sessile. Drupe fleshy, subglobose, crowned with the stigmas. — FI. Cap. i. p. 473. Evergreen shrubs or trees, widely dispersed. I. Capensis, our only spe- cies, found throughout the colony, is a large shrub or small tree, with ob- long or lanceolate, shining, entire leaves, and axillary, fascicled, white flowers. Order XXXVI. CELASTEINEiE. Flowers mostly bisexual. Calyx small, 4-5-lobed or parted, imbricate, persistent. Petals 4-5, short, spreading, sessile under the margin of the disk, imbricate. Stamens 3-5 (rarely 2-10), inserted at the base, on the margin, on the surface, or on the lobes of the disk ; filaments mostly short ; anthers 2- celled. Disk conspicuous, convex or expanded or lobed. Ovary sessile on or partly immersed in the disk, 3-5- (rarely 1-) celled ; style short, simple or 2-3-fid ; ovules commonly 2, mostly erect, rarely pendulous. Fruit a capsule drupe or berry, or winged ( [samara ) ; seeds with or without albumen. — Trees and shrubs, often spinous. Leaves opposite or alternate, mostly leathery, simple. Flowers axillary, small. Tribe 1. Celastre2E. Stamens inserted on or beneath the margin of a conspicuous, fleshy disk. Anthers introrse. Fruit capsular, dehiscent. Leaves alternate. Ovules about 6 in each ovarian cell .... 1. Putterlichia. Ovules 2 in each ovarian cell. Yalves of capsule not winged 2. Gtmnosporia. Valves of capsule dorsally winged .... 3. Pterocelastrtjs. E 2 52 XXXVI. CELASTRINE^E. Leaves opposite. Ovary 3-celled ; ovules in pairs, erect . . . Ovary 1-celled ; ovules 6-8, parietal .... Fruit a fleshy, indehiscent drupe. Ovules solitary, erect. Leaves opposite . . . Ovules in pairs, pendulous. Leaves opposite . . Ovules in pairs, erect. Stamens 4. Ovary 2-celled. Leaves opposite Stamens 5. Ovary 3-celled. Leaves opposite or alternate 4. Catha. 5. Cathastrum. 6. Hartogia. 7. Matjrocenia. 8. Latjridia. 9. Elaiodendron. Tribe 2. Hippocrates. Stamens 3 (rarely 2-4-5), inserted much within the margin of the disk ; filaments flat ; anthers extrorse. Fruit a 1- or several-seeded berry. Seeds wingless. 10. Salacia. Tribe 1. Celastre^:. (G-en. 1-9.) 1. PUTTERLICHIA, Endl. Calyx flat, 4-5 -parted. Petals 4-5, spreading. Stamens 4-5, spreading, inserted under tlie margin of the disk ; fila- ments subulate ; anthers subglobose. Disk thick, hemispheri- cal, ribbed. Ovary half-sunk in the disk, 3-5-angled, 3-5- celled ; style 3-5-angled ; stigma 3-5-lobed ; ovules 6 in each cell, 2-seriate. Capsules obtusely 3-angled, 3-celled, loculici- dally 3-valved ; cells 3-6-seeded. Seeds with a fleshy arillus, albuminous. — Celastrus, Sect. 1, in FI. Cap. i. p. 453. Glabrous, spiny shrubs, with alternate or tufted, obovate leaves, and axillary, diffusely -branched cymes of white flowers. — There are 2 species, one of them dispersed, the other Eastern. 2. GYMNOSPORIA, W. and A. Calyx 4-5-fid or parted. Petals 4-5, sessile, spreading. Stamens as many, inserted on or under the margin of the disk. Disk widely spread, 4-5-lobed or crenate. Ovary very gene- rally with a broad base confluent with the disk, 3 -angled or pyramidal, 2-3-celled ; style short ; stigmas 3 ; ovules in pairs, erect. Capsules obovoid or subglobose, 3-angled or globose, 2-3-celled, 1-4-seeded. Seeds with or without an aril, albu- minous.— Celastrus, # Sect. 2, FI. Cap. i. p. 454. Also Scyto- phyllum, F. and Z. (which has dehiscent fruit), FI. Cap. i. p. 471. A large genus of shrubs, often spinous. Leaves alternate, entire or toothed, sometimes pubescent. Flowers in axillary cymes or tufts. — Up- wards of 20 South African species, dispersed. * The true Celastrus , Linn., differs in having an ovary seated on , but not immersed in, the disk, and by other characters. Its species, about 18 in number, are chiefly Asiatic ; a few American and Australian, and one from Madagascar. They are mostly climbing shrubs, without spines. XXXVI. CELASTRINEiE. 53 3. PTEROCELASTRUS, Meisn. Ploral characters nearly as in Gymnosporia. Capsules carti- laginous, 3-6-winged, 1-3-celled, loculicidally 3-valved, slowly dehiscing, the valves septiferous in the middle ; cells 1-2- seeded. Seeds with a thin, membranous arillus, albuminous. — FI. Cap. \.p. 461. South African shrubs, with alternate, leathery, quite entire leaves and axillary, small, white, cymose or tufted flowers. — 6 species, dispersed. 4. CAT HA, Porsk. Calyx 5-lobed, small. Petals 5, erecto-patent. Stamens 5, on the margin of the disk ; filaments subulate ; anthers 2- parted. Disk thin, with an undulate margin. Ovary ovoid, immersed in the disk, free, 3-celled ; style short, thick ; stigmas 3 ; ovules in pairs, erect. Capsules linear- oblong, 3-celled, 3- valved, 1-3-seeded. “ Ripe seeds winged.” — Methyscophyllum, FI. Cap. i. p. 463. C. edulis, Forsk. ( = Methyscophyllum, glaucum E. and Z.), the “ Bosjes- man’s-the” of the colonists, grows in North and South Africa and in Arabia. The leaves, chewed to excess, are intoxicating. Leaves opposite, glaucous, lanceolate, acuminate, repando-serrate, with revolute margins, netted- veined. Peduncles axillary, dichotomous, short. 5. CATHASTRUM, Turcz. Calyx 5-parted, the lobes rounded, fimbriate. Petals 5, obovate, ciliolate, revolute. Stamens 5, inserted under the margin of the disk ; filaments thickish, recurved ; anthers sub- cordate, affixed to the broadish apex of the filaments. Disk thin, obtusely 5-angled. Ovary sessile on the disk, narrowed at base, oblique, free, 1-celled ; style short, stigma thick, uni- lateral, peltate ; ovules 4-8, in two rows, ascending. — FI. Cap. i.p. 526. C. Capensis , Turcz., the only species, is a glabrous shrub, with opposite, petioled, oblong or linear-oblong, leathery, obtuse, undulate, entire leaves, and short, axillary cymes of small flowers. The fruit is unknown. It in- habits woods in Uitenhage. 6. HARTOGIA, Thnnb. Calyx 4-5-fid. Petals 4-5, spreading. Stamens 4-5, be- tween the lobes of the disk. Disk annular, 4-5-lobed. Ovary sessile, scarcely confluent with the disk, pyramidal, 2-3 -celled, tapering into a thick style ; stigma obtuse ; ovules solitary in each cell, erect. Emit ovoid, dry, indehiscent. Seed without albumen. — FI. Cap. i. p. 464. H. Capensis , Th., the only species, is a shrub with opposite, leathery glaucous leaves, with revolute, serrulate margins. Cymes axillary ; flowers small, white. Found in the Western and middle districts. 5 4 XXXYI. CELASTRINES. 7. MATJROCENIA, Linn. Calyx minute, 5-parted. Petals 5, longer than the calyx. Stamens 5, under the margin of the disk, longer than the petals ; filaments filiform ; anthers broadly oblong. Disk cup- like, sinuate, 5-lobed. Ovary sessile on the disk, not confluent with it, ovoid, 2-3-celled ; stigmas sessile, 2-3-lobed ; ovules in pairs, pendulous. Drupe ovoid, fleshy, 1-3-celled. Seed albuminous. — FI. Cap. i. p. 465. Cassine, Linn., BentJi. and Hook.f. Gen. PI. i. p. 363. M. Capensis , Sond., the only species, is a glabrous shrub, with 4-angled twigs, opposite, leathery, quite entire, glossy leaves, and axillary short, cy mules of small white flowers. It is frequent in the Western districts. 8. LAURIDIA, E. and Z. Calyx 4-parted, the lobes strongly imbricate. Petals 4, ovate-oblong, revolute, with an uneven margin, imbricate. Stamens 4, on the margin of the disk ; filaments broad-based, flattish, subulate ; anthers broadly oblong. Disk adnate to the calyx-tube, the limb thin, obscurely lobed. Ovary subimmersed in the disk, 2-celled ; style very short ; stigma 2-lobed. Drupe rather dry, 2-celled, 1-2-seeded, with a crustaceous stone. — FI. Cap. i. p. 468. L. reticulata, E. and Z., the only species, is a glabrous, trichotomous shrub, with opposite, very entire, or sparingly toothed, netted-veined leaves, and axillary, paniculate-racemose, small flowers. It grows in the Eastern district. — As a genus, Lauridia scarcely differs from Elceodendron. 9. ELCEODENDRON, Jacq. f. Elowers sometimes polygamous. Calyx 4-5-parted. Petals 4-5, spreading. Stamens 4-5, under the margin of the disk ; filaments short, subulate ; anthers subglobose. Disk thick, expanded, 4-5 sinuate-angled or lobed. Ovary pyramidal, confluent with the disk, mostly 3-angled, 3-celled, rarely 2-5- celled ; style very short ; stigma 2-5-lobed ; ovules in pairs, erect. Drupe dry or pulpy, 1-3-celled. Seed albuminous. — FI. Cap. i. p. 467. Also Cassine, S. and H.,p. 465, and Mys- troxylon, F. and Z., 1. c.,p. 469. A considerable genus, of which there are about 18 Cape species, dispersed. Leaves opposite or alternate, entire or toothed, glabrous or pubescent, leathery, mostly evergreen. Peduncles axillary ; flowers small. Tribe 2. Hippocrates. 10. SALACIA, Linn. Calyx small, 5-parted. Petals 5, spreading, imbricate. Stamens 3 (very rarely 2 or 4), inserted on the inner margin XXXYI. CELASTRINEJ3. of the disk, close to the ovary, free or connate with the ovary ; filaments flattened, recurved ; anthers mostly extrorse, 2-1- celled. Disk thick, flat or conical, sinuate. Ovary immersed in the disk, 3-celled, tapering into a short or longer style ; stigma simple or 3-lobed ; ovules 2, 4, or more, axile. Fruit berried, 1-3-celled ; cells 1-4-seeded. Seed exalbuminous. — FI. Cap. i. p. 230. Trees or shrubs, often climbing. — 1 South African species, found near Natal, with alternate leaves. Flowers in axillary tufts. Order XXXYII. RHAMNE^]. Flowers perfect, regular. Calyx-limb 4-5-fid, the lobes acute, valvate in bud. Petals 4, 5 or 0, inserted in the throat of the calyx, usually small, hood- shaped or flat. Sta- mens 4-5, inserted with the petals and opposite them. Disk perigynous (rarely 0), either thick, filling the calyx-tube, or annular, or cup-like, simple or lobed. Ovary sessile, free or more or less adnate to the calyx-tube ; 3- rarely 2-4-celled ; style erect, simple or cleft; ovules mostly solitary, erect. Fruit fleshy or capsular. Seeds mostly albuminous. — Trees or shrubs, often spiny, sometimes climbing. Leaves simple, al- ternate or opposite, often 3-5-nerved, or narrow-linear, 1- nerved. Flowers small, in axillary cymes or terminal, capitate. Ovary quite free (not adhering to the calyx-tube). Ovary sunk in and confluent with the flat, 5-angled disk . 1. Zizyphtjs. Ovary not sunk in or confluent with the disk. Disk thin, lining the calyx-tube ; styles 3-4 2. Khamnus. Disk fleshy, filling up the calyx-tube, but not confluent with the ovary ; style simple 3. Scutia. Ovary more or less adhering to the calyx-tube ; fruit inferior or half-in- ferior. An erect shrub, with expanded, penninerved, serrate leaves ; flowers panicled, glabrous 4. Noltea. A climbing shrub, with expanded, penninerved, entire leaves ; flowers in axillary cymes, glabrous 5. Helinus. Small erect shrubs, with crowded, often linear, entire leaves, and flowers in heads or spikes, rarely panicled ; calyx hairy 6. Phylica. 1. ZIZYPHTJS, Juss. Calyx 5-fid, the tube broadly obconic, the lobes ovate, spreading, keeled within. Petals 5, rarely 0, hood- shaped. Disk flat, 5-angled, the margin free. Stamens 5, exserted. Ovary immersed in the disk and confluent with it at base, 2- rarely 3-4-celled ; styles 2-3, free or connate, divergent. Drupe fleshy, globose or oblong, 1-3-celled. — FI. Cap. i. p. 475. 56 XXXVII. RHAMNE.E. Shrubs or trees, often trailing, mostly armed with hooked prickles. Leaves alternate, petiolate, entire or crenate, 3-5-nerved. Cymes axillary, few-flowered. — 3 South African species, from the Northern and Eastern districts. 2. RHAMNUS, Linn. Calyx 4 -5-fid ; tube urceolate, tbe lobes triangular, erect or spreading, keeled witbin. Petals 4-5 or 0, hooded or flat. Stamens 4-5, with very short filaments. Disk clothing the whole calyx-tube, the margin thin. Ovary free, ovoid, hidden in the calyx-tube, 3-4-celled, tapering into a 3-4-lobed style. Drupe fleshy, oblong or globose, girt by the persistent base of calyx ; stone 2-4-celled.— FI. Cap. i. p. 476. Shrubs or small trees. Leaves opposite, penninerved, entire or toothed. Flowers axillary, racemose or cymose. — 2 South African species, natives of the Eastern district and Natal. 3. SCUTIA, Comm. Calyx 5-fid, with a hemispherical or turbinate tube, the lobes ovate, thickened at the point, deciduous. Petals 5, clawed, erect, flat or hooded. Stamens 5. Disk filling up the tube of the calyx, the margin free, undulate. Ovary ovoid or glo- bose, hidden in the disk, but not confluent with it, free, 2-4- celled, tapering into a short simple or 2-4-fid style. Drupe obovoid or subglobose, dry or sparingly fleshy, girt by the cup-like base of the calyx ; stone 2-4-celled. — FI. Cap. i. p. 477. GTlabrous, unarmed or spiny shrubs, often with angular twigs. Leaves approaching in pairs, but not strictly opposite, petioled, coriaceous, penni- nerved. Flowers in axillary tufts or umbels. — S. Commersoni , Br., the only South African species, is common in woods, from Swellendam to Natal. 4. NOLTEA, Echb. Plowers polygamous. Calyx 5-fid, the tube turbinate ; lobes ovate, erect or recurved. Petals 5, cucullate, sessile. Sta- mens 5, equalling the petals. Disk thin, lining the calyx- tube, the margin inconspicuous. Ovary half-inferior, 3-lobed at summit, 3-celled, tapering into a 3-angled style. Drupe dry, obovoid, girt below the middle by the persistent and ad- herent calyx-tube, 3-lobed, the lobes dorsally keeled. — FI. Cap. i. p. 478. N. Africana , Rchb., the only species, grows wild in the Eastern district and at Natal ; it is commonly cultivated throughout the colony. Leaves alternate, oblongo- lanceolate, serrated, penninerved, obtuse. Flowers small, white, in terminal or axillary panicles. 5. HELINTJS, E. Mey. Calyx-tube obconical, adhering to the ovary ; limb 5-parted, XXXVII. RHAMNEiE. 57 spreading, deciduous. Petals 5, hooded, inserted on the mar- gin of the disk. Stamens 5, as long as petals. Disk epigy- nous, flattish, filling the calyx-tube. Ovary 3-cel led ; style 3-fid. Fruit inferior, coriaceous, obovate-globose, areolate at summit, 3-coccous ; cocci crustaceous, dehiscing within. — FI. Cap. i.p. 478. Climbing, tendrilled shrubs, with slender, angular branches, and alter- nate, entire, cordate leaves. Flowers umbelled. H. ovata , E. Mey., grows on the Eastern frontier and at Natal. 6. PHYLICA, Linn. Calyx-tube obconic, urceolate or cylindrical, adherent to the ovary, its limb 5-fid or parted ; lobes hairy outside, mostly persistent. Petals wanting or bristle-shaped, or hooded. Sta- mens 5, short. Disk epigynous, and filling up the calyx-tube, distinct or inconspicuous. Ovary inferior, 3 -celled ; style short, rarely elongate, 3-fid. Fruit inferior, globose or ovoid, areolate, smooth or tomentose, with a leathery outer coat, 3- coccous within ; cocci at length opening, on the inner face. — FI. Cap. i. p. 479. A large genus, chiefly South African. Small, much-branched shrubs, with alternate, crowded, entire, linear, lanceolate or rarely ovate leaves. Flowers in terminal, bracteate spikes or heads, rarely pedicellate, and either solitary or panicled. — 58 South African species, dispersed. Order XXXYIII. AMPELIDE.®. Flowers regular, perfect or unisexual. Calyx small, entire or 4-5-toothed or lobed. Petals 4-5, separate or cohering, valvate in bud. Stamens 4-5, opposite the petals, inserted at the base or between the lobes of the disk ; filaments subulate ; anthers introrse, 2-celled. Disk various. Ovary very com- monly sunk in the disk, 2-6-celled ; cells 1-2-ovuled ; style single or 0. Fruit a berry. Seeds erect, with very hard, bony coats, and abundant fleshy albumen. — Mostly climbing or trail- ing shrubs, with knobbed or jointed stems. Leaves petioled, simple or compound. Flowers small, mostly green. 1. VITIS3 Linn. Calyx cup-like, 4-5-toothed. Petals 4-5, separate or co- hering in a cap. Stamens 4-5. Ovary ovoid or sub-4-fid, 2-celled (very rarely 2-4-celled) ; style filiform or 0 ; ovules in pairs. Berry 1-2-celled ; cells 1-2-seeded. — FI. Cap. i. p. 248. Also Cissus, Linn. ; FI. Cap. i. p. 249 ; T/ies. Cap. t. 65. Cirrhose, mostly climbing or scrambling shrubs. Leaves simple or com- pound, rarely 2-pinnate ; leaflets entire, serrate or cut. Stipules membra- 58 XXXYIII. AMPELIDEiE. * nous or 0. Peduncles opposite the leaves. Flowers small, cymose, panicled or spiked. The Grape Vine is the type of this genus. — There are about 18 South African species, all but V. Capensis , which is dispersed, natives of the Eastern district and Natal. Order XXXIX. SAPINDACEAS. Flowers regular or irregular, frequently polygamous. Se- pals 4-5, free or connate, often unequal, imbricate, rarely val- vate. Petals 0 or .3-5, imbricate. Disk various, rarely defi- cient, often unilateral. Stamens 8, rarely 5-10 (very rarely 2^-12 or many), mostly hypogynous, inserted either within the disk, sometimes unilateral, straight or declined, or rarely round the base of the disk. Ovary entire or lobed, mostly 3- celled, or 1-4-celled ; style simple or divided, terminal ; ovules 1-2, rarely more in each cell, ascending. Fruit capsular or indehiscent, often pulpy within. Seeds rarely (in Melianthece) albuminous. — Trees and shrubs, rarely half-herbaceous. Leaves alternate, mostly pinnate, rarely simple. Fruit membranous, inflated, 3-4- lobed and celled. Fruit 3 -lobed, the lobes dorsally winged. Calyx 4-parted. Herbaceous, tendril-bearing climbers 1. Cardiospermttm. Calyx 5-parted. An erect, rigid shrub ... 2. Erythrophysa. Fruit 4-lobed, the lobes sharply angled or winged. Calyx 4-parted. Petals equal. Stamens 8, monadelphous 8. Aitonia. Calyx 5-parted, unequal. Petals unequal. Sta- mens 4, 2 long and 2 short 9. Melianthus. Fruit fleshy or leathery. Carpels 2-3-4, not com- pressed or winged. Sepals and petals 4. Ovary 2-lobed 3. Schmidelia. Sepals and petals 5. Ovary 3-4- celled. Stamens 8-10. Ovary 3-celled. Ovary lobed. Fruit 3-coccous 5. SapinduS. # Ovary undivided. Fruit drupaceous ... 4. Hippobromus. Stamens 5. Ovary bluntly 4-kngled, 4-celled . 10. Bersama. Fruit strongly compressed or winged, not inflated. Petals 3. Stamens 5-8. Capsule 2-4-winged . 6. Dodonaia. Petals 4. Stamens 4. Capsule oblong, com- pressed, 2-lobed at the apex. Seeds winged . 7. Pt^eroxylon. Tribe 1. Sapende^:. (G-en. 1-5.) 1. CARDIOSPERMUM, Linn. Flowers irregular, polygamo-dioecious. Sepals 4, concave, imbricate, the 2 outer small. Petals 4, in opposite pairs, the 2 larger with a large scale, 2 smaller with a small crested scale at base. Disk unilateral, undulate, swelling into 2 glands opposite the lower petals. Stamens 8, excentric. Ovary ses- XXXIX. SAPINDACEiE. 59 sile or stipitate, 3-celled ; style short, 3-fid ; ovules solitary. Capsule 3-lobed ; lobes inflated, membranous, veiny, loculici- dally opening. — FI. Cap. i. p. 237. Mach-branched, half-herbaceous climbers. Leaves 2-ternate or decom- pound, the common petiole bearing tendrils. Flowers in axillary racemes or panicles. — C. Halicacaba, Linn., a common tropical weed, is found at Natal. 2. EEYTHROPHYSA, E. Mey. Flowers perfect, irregular. Calyx campanulate, oblique, coloured, 5-lobed, the lobes obtuse, unequal. Petals 4 (the place of the fifth vacant), inserted under the margin of a fleshy, cup-like disk, on long, linear, pilose claws ; limb oblong, ob- tuse, hooded at base, and furnished with a short, petaloid, toothed and crested, but beardless scale. Stamens 8, ascend- ing, inserted together under a fleshy gland, on that side of the flower where the fifth petal is deficient ; filaments hairy ; an- thers oblong, 2-celled. Ovary shortly stipitate, 3-angled, tapering into a short 3-angled style, 3-celled ; ovules in pairs, one above the other. Eruit inflated, of 3 membranous, dor- sally-winged, valveless carpels, connate by their inner faces. Seed solitary, globose, exalbuininous. — FI. Cap. i. p. 237. JE. undulata , E. Mey., the only species, is a rigid, glabrous shrub, with imparipinnate leaves on winged petioles, and racemose red flowers. It grows in Namaqualand. 3. SCHMIDELIA, Linn. Flowers irregular, polygamo-dioecious. Sepals 4, in oppo- site pairs, hooded, membranous, imbricate, the outer small. Petals small or 0, glabrous or bearded. Disk unilateral, either lobed or swelling in glands opposite the petals. Stamens more or less excentric, short. Ovary excentric, 2-celled, compressed or 2-parted ; style 2-3-lobed or partite ; ovules solitary. Fruit of 1-2 ovoid or globose, leathery or fleshy car- pels.— FI. Cap. i. p. 238. Trees or shrubs, chiefly tropical. Leaves rarely 1-, commonly 3-foliate. Flowers small, in axillary racemes. — 5 South African species, in the Eastern district and at Natal. 4. HIPPOBROMTJS, E. and Z. Flowers regular, polygamous. Sepals 5, persistent, rounded, concave, unequal, imbricate. Petals 5, obovate, glabrous, ciliate-fringed, without scale. Disk annular. Stamens 8, central, exserted ; filaments glabrous. Ovary subglobose, 3- celled, silky ; style short ; stigma 3-fid ; ovules mostly solitary. Fruit globose, leathery, 3-celled, indehiscent. — FI. Cap. i. p. 241. 60 XXXIX. SAPINDACEiE. //. alata , E. and Z., the only species, is a resiniferous tree, with abruptly pinnate leaves, the common petiole winged . Panicles axillary, short ; flower reddish, velvety. It is common in woods in the Eastern district and at Natal. 5. SAPINDUS, Linn. Flowers polygamous. Calyx 5-parted or of 5 imbricating sepals. Petals 4, 5, 6, naked or bearded on the base inside, or having a scale above the claw. Stamens 8-10, rarely more, inside an annular disk. Ovary 2-4-lobed, 2-4-celled ; style 1 ; stigmas 3. Fruit fleshy or leathery, of 3 (or 1-2) oblong or globose, indehiscent cocci. — FI. Cap. i. p. 240. Trees, with pinnate or (rarely) simple, leathery leaves. Flowers in axillary racemes or terminal panicles. — 3 Cape species, found in the Eastern district and at Natal. Tribe 2. Dodoxea:. (G-en. 6-8.) 6. DODONEA, Linn. Flowers deciduous. Sepals 2-5, imbricate or valvate. Pe- tals 0. .Disk in the male obsolete, in female small. Stamens 5-8, central ; filaments very short ; anthers linear- oblong, bluntly 4-angled. Ovary sessile, 3-6-angled, 3-6-celled ; style 3-6-fid ; ovules in pairs. Capsules membranous or leathery, 2-6-lob ed, the lobes dorsally winged. — FI. Cap. i. p. 241. A large genus, chiefly Australian. — 2 South African species, one of them dispersed, the other at Natal. Leaves simple, obovate-oblong, lanceolate or linear-lanceolate, glabrous, often gummy. Flowers racemose, green. 7. PTEROXYLON, E. and Z. Flowers polvgamo-dioecious. Sepals 4, short, obtuse. Petals 4, at first erect, slightly imbricate, then spreading, con- cave. Disk annular, 5-crenate. Stamens 4, at the base of the disk ; filaments glabrous ; anthers oblong. Ovary compressed, obcordate, 2-celled ; ovules solitary ; styles 2 or connate in 1 ; stigma capitate. Capsules compressed, 2-lobed at apex, cor- date at base, 2-celled, 2-seeded, at length 4-parted. Seeds compressed, with a membranous wing. — FI. Cap. i. p. 242 ; Thes. Cap. t. 17. P. utile, E. and Z., is a tree (Sneezewood) with pinnate leaves, the leaf- lets unequal-sided, in 5-8 pairs. Racemes panicled, axillary, shorter than the leaves. Woods in the Eastern district. 8. AITONIA, Linn. f. Calyx deeply 4-parted, slightly imbricate, deciduous. Pe- tals 4, much longer than the calyx, erect, oblong, sessile, con- volute-imbricate: Stamens 8, hypogynous, monadelphous, exserted ; filaments decimate, subulate, fiat, their dilated bases XXXIX. SAPINDACEiE. G 1 confluent. Disk cup-like, crenulate, fleshy, within the sta- mens. Ovary sessile, 4-lobed, 4-celled ; ovules in pairs ; style filiform, exserted. Capsules inflated, membranous, deeply 4- lobed, sharply 4-angled, 4-celled.' — FI. Cap. i. p. 243. A. Capensis, Linn, f., the only species, is a shrub with narrow linear evergreen leaves, purple flowers and bladdery capsules, found in the Uiten- hage, Albany, etc., districts. Teibe 3. Meliantheaj. (Gen. 9-10.) 9. MELIANTHUS, Linn. Flowers perfect. Calyx compressed, oblique at base, sub- saccate, 5-parted, segments very unequal, imbricate. Petals 5, excentric, the fifth very minute or 0, decimate, subperigy- nous, narrow, long-clawed, tomentose in the middle. Disk thickened, unilateral, covering the gibbous base of the calyx, secreting honey. Stamens 4, hypogynous, inserted within the disk, didynamous, and curved. Ovary oblong, 4-lobed, 4- celled ; style central, incurved, filiform, 4-toothed at apex ; ovules 2-4 in each cell. Capsules membranous, deeply 4- lobed, 4-celled; cells 1-seeded. Seeds albuminous. — FI. Cap. i. p. 367. Glabrous or hoary, often glaucous, often strong-scented. Leaves alter- nate, stipuled, pinnate ; leaflets sharply-toothed or entire. Racemes axil- lary and terminal. — 5 species, two of which are dispersed, three North- Western. 10. BERSAMA, Presen. Flowers perfect or polygamo-dioecious. Sepals 5, subequal, free, or two more or less connate, imbricate. Petals 5, un- equal, clawed, the front one larger, the claws nude or glandu- lar at apex, imbricate. Disk unilateral, semi-annular or sub- complete, raised. Stamens 4, central, inserted with the disk, either all connate or 2 free. Ovary oblong, terete, 4-5-celled ; style elongate, curved ; ovules solitary, erect. Capsules glo- bose or oblong, coriaceous or ligneous, 4-5-celled, loculicidally 4-5-valved, valves septiferous. Seed arillate, albuminous. — Natalia, Hochst. ; FI. Cap. i. p. 369. Trees and shrubs. Leaves alternate, imparipinnate ; leaflets entire or serrulate. Racemes terminal, lateral, simple j flowers small, white. — 2 South African species, both found near Natal. Genes of enceetain Affinity. GREYIA, Hook, and Harv. Calyx 5-parted, persistent, its segments obtuse, subequal, imbricate. Petals 5, oblong, sessile, deciduous, coriaceous, imbricate. Stamens hypogynous, in two rows, those of the outer row (staminodia) 10, without anthers, their very short G2 XXXIX. SAP1NDACE-®. filaments adnate to the inner face of a fleshy, cup-like, 10- crenate disk, and each crowned with a peltate gland ; those of the inner row 10, fertile, free, alternating with the staminodia, much exserted ; filaments subulate, declined ; anthers termi- nal, short, didymous, opening at the sides ; pollen oval, very soon shed. Ovary free, deeply 5-furrowed, formed of 5 in- duplicate-valvate carpels, 1-celled, tapering into a subulate, exserted, decimate style ; stigma minutely 5-toothed ; ovules on sutural placentas, very numerous. Capsules deeply 5-lobed, of 5 follicular, papery carpels, slightly cohering at the su- tures. Seeds minute, with membranous testa, and a small straight embryo, in the base of copious, fleshy albumen. — FI. Cap. ii. p. 309. 6r. Sutherlandi, Thes. Cap. t. 1, is a middle-sized tree or large shrub, with alternate, simple, exstipulate leaves. Petioles clasping at base. Leaves subrotund, cordate at base, 2-4 inches diameter, multilobulate and crenate, glabrous. Eacemes terminal, densely many-flowered ; flowers crimson, very showy. Grows near Natal. — By Dr. Hooker this genus is regarded as a genuine Sapindacea, of the tribe Melianthece. Though I allow that there are many points of agreement in the general structure of the flower, yet I am of opinion that the 1 -celled ovary, parietal placentas, indefinite and very numerous ovules, copiously albuminous seeds, and minute , straight embryo are all important characters at variance with Sapindacece. Order XL. ANACA&DIACEiE. Flowers either complete, polygamous or unisexual. Calyx 3- 7-fid or parted, sometimes enlarged after flowering, rarely irregularly torn. Petals 3-7, rarely 0, free, sometimes en- larged after flowering. Disk annular (rarely stipitate). Sta- mens mostly twice as many as petals, rarely only as many or numerous, inserted at the base of the disk. Ovary in the female mostly ovoid, 1-celled ; in Spondieee 2-5-celled, or rarely of several separate carpels ; in the male, 3-4 empty separate or confluent carpels ; styles 1-3. Ovules solitary, pendulous, either from the side of the ovary or from a slender cord rising from its base. Fruit mostly drupaceous, 1-5-celled. Seed exalbuminous. — Trees or shrubs, with caustic, balsamic or gummy juice. Leaves alternate, simple or compound, without stipules. Tribe 1. Anacaediea:. Ovary 1-celled, with one ovule (sometimes with 3-4 empty carpels). Stamens 8. Sepals and petals 4 1. Odina. Stamens 4-5. Calyx unchanged in fruit. Drupe compressed, wingless 2. Khtts. Drupe much compressed, sharply edged. Style 1 3. Botryceras. * XL. ANACARDIACEiE. 63 Fruit compressed, oblique, broadly winged. Styles 3 4. Smodinguttm. Calyx in fruit enlarged and coloured 5. Loxostylis. Tribe 2. Spondie.®. Ovary 2-5-celled; ovules 1 in each cell. Male flower spiked. Stamens 12-15 6. Sclerocarya. Male flower panicled. Stamens 8-9 7. Harpephyllum. Tkibe 1. A^acakdiea:. (G-en. 1-4.) 1. ODINA, Roxb. Mowers poly gamo- dioecious. Calyx 4-5-lobed, persistent, tbe lobes imbricate. Petals 4-5, patent or reflexed. Disk annular, 4- 5 crenate. — Male : Stamens 8-10, under the disk. Barren ovary 4-5-parted. — Pemale : Ovary sessile, free, oblong, 1- celled ; styles 8-4. Drupe compressed, oblong-reniform, crowned with the persistent styles. — FI. Cap. i. p. 503. Large trees, Indian and African. Leaves usually pinnate ; leaflets oppo- site, subsessile, entire, pale beneath. Racemes terminal, tufted. Flowers small. — 2 South African species, both from Magalisberg. 2. RHUS, Linn. Blowers polygamo- dioecious. Calyx small, 5-6-parted, per- sistent. Petals 5-6, oblong or ovate, spreading. Stamens 5- 6. Ovary subglobose, 1-celled (abortive in the male flower); styles 3, free or connate. Drupe nearly dry or slightly fleshy, 1-celled, containing a bony, 1-seeded nucleus. — FI. Cap. i. p. 504. A large and varied cosmopolitan genus. The leaves in the Cape species are either 3-foliate or simple. Flowers panicled, small, green. — Perhaps 60 (several undescribed newly-discovered at Natal) South African species, dispersed. 3. BOTRYCERAS, Willd. Blowers dioecious. — Male: Calyx 4-5-cleft nearly to the base, spreading ; lobes oblong, imbricate. Petals 4-5, lanceo- late, reflexed. Stamens 4-5, around the base of the fleshy, annular disk. Ovary 0. — Bemale : Calyx and corolla per- sistent, as in the male, but the petals are oblong, obtuse. Ovary ovate, compressed, 1-celled ; style 1, thick, oblique ; stigmas 3, bristly. Bruit with a membranous pericarp, com- pressed, winged at the margin, rugose, tipped with the persis- tent style. B. laurinum , Willd., the only species, is a large, glabrous and resinous shrub, with simple, elliptic-oblong, penninerved, serrate leaves. 4. SMODINGIUM, E. Mey. Blowers polygamous. Calyx 5-cleft, persistent. Petals 5, oblong, deciduous. Stamens 5 ; filaments subulate. Ovary free, sessile, 1-celled; styles 3. Bruit compressed, the margin 64 XL. ANACAItDIACEiE. winged, obliquely-oblong, on both sides multivittate ; the vittse flexuous, parallel. — FI. Cap. i. p. 523. 8. argutum is a glabrous shrub, with long-stalked, 3-foliolate leaves ; leaflets coarsely serrate, lanceolate, penninerved. Flowers minute, in ter- minal panicles. Found near Natal. 5. LOXOSTYLIS, Spreng. Flowers polygamous, Calyx 5-parted, segments imbricate, membranous, lanceolate ; in the female flower enlarged after flowering, leafy. Petals 5, lanceolate, imbricate. Disk of 5 2-fid, perigynous glands. Stamens 5, between the glands of the disk; the filaments unequal. Ovary (in the female) ob- lique, compressed, 1-celled ; styles 3, lateral, unequal ; stigmas capitate. Drupe small, oblique, compressed, hidden within the large leafy calyx. — FI. Cap. i. p. 524. L. alata , Spreng., the only species, is a small, glabrous tree. Leaves imparipinnate ; leaflets opposite, subsessile, lanceolate, entire, the common petiole winged. Flowers in terminal panicles. — Eastern district and Natal. Tribe 2. Spohdie^. (Glen. 6-7.) 6. SCLEROCARYA, Hochst. Flowers polygamo-dioecious ; males spiked. Sepals 4, co- loured, suborbicular, imbricate. Petals 4, oblong, obtuse, spreading, reflexed, imbricate. Disk depressed, entire. Sta- mens in the males 12-15 ; in the female fewer, some abortive. Ovary subglobose, 2-3-celled ; styles 2-3, short, thick, distant ; stigmas peltate. Drupe rather fleshy, with a hard, woody nucleus, 2-3 celled. — FI. Cap. i. p. 524. African trees or shrubs. S. Caffra , Sond., our only species, has impari- pinnate leaves ; leaflets 5-13, ovate or elliptical, shortly cuspidate, with long petiolules. Male spikes 2-4, terminal. — Magalisberg and Natal. 7. HARPEPHYLLUM, Bernh. Flowers dioecious ; males panicled. — Males : 4-5-fid, the lobes obtuse, imbricate. Petals 4-5-fid, longer than the calyx, imbricate. Disk crenate. Stamens 8-9, beneath the margin of the disk. Ovary rudimentary, 4-lobed. — Females : Flowers unknown. Drupe obovate, smooth, with a 2-celled, bony stone. — FI. Cap. i. p. 525 ; Thes. Cap. t. 125. H. Caffrum , Bernh., the only species, is a glabrous tree, found in the Eastern district and Kaffraria. Leaves crowded at the ends of the twigs, imparipinnate; leaflets sessile, falcate-lanceolate, entire, unequal-sided. Flowers white, in terminal panicles ; the females unknown. 65 Order XLI. CONNARACE.®. Flowers mostly bisexual, regular or subirregular. Calyx 5-fid or 4~5-parted, often persistent. Petals 5, perigynous, in- serted in the bottom of the calyx, sessile or clawed, mostly imbricate. Stamens 5-10, inserted with the petals ; filaments free or slightly connate. Ovarian carpels 5 (rarely fewer), se- parate, either all fertile or some abortive, the fertile 1-celled, with 2 collateral ascending ovules ; styles terminal, as many as the carpels. Ripe carpels mostly solitary, follicular, leathery, rarely indehiscent. Seed solitary, with or without albumen; radicle remote from the hilum. — Trees or shrubs, chiefly tropical. 1. CNESTIS, Juss. Calyx 5-cleft, valvate. Petals 5, shorter than the calyx, glabrous. Stamens 10 ; filaments free ; anthers at length re- curved. Carpels 5, sessile ; styles short ; stigmas capitate. Capsules 1-2, oblong, reniform or cylindrical, and curved or undulate, velvety without, within clothed with rigid and stinging hairs. Seed without aril, albuminous. — FI. Cap. i. p. 527. Shrubs, often climbing. Leaves alternate, imparipinnate ; leaflets coria- ceous, entire. Flowers in racemose panicles. — C. Natalensis , PI. and Sd., is our only species. Order XLII. LEGUMXNOS.®. Calyx free, 5-toothed, cleft or parted, equal or unequal, the odd segment in front. Petals 5 (some or all occasionally wanting), usually unequal, inserted in the base of the calyx. Stamens perigynous or hypogynous, definite or indefinite, va- riously combined. Ovary of one carpel, with 1, 2 or many ovules, attached sometimes by long cords to the ventral suture ; style proceeding from the upper margin (i. e. continuous with the ventral suture) ; stigma simple. Pruit a legume or a lomentum, rarely drupaceous. Seeds usually exalbuminous. — A vast Order, very varied in habit, but naturally divisible into 3 Suborders, as follows : — 1. Papilionace.®:. Corolla papilionaceous ; petals 5, imbricated in aesti- vation, the upper petal ( vexillum , standard) exterior, folding over the 2 lateral petals (alee, or wings), which fold over the 2 anterior (or carina, or keel). Stamens 10, all perfect, either diadelphous (9 united and 1 free) monadelphous or free. 2. Gesalpinie.®:. Corolla irregular or subregular, not papilionaceous ; petals imbricating, the 2 anterior folding over the 2 lateral, which enclose the upper petal. Stamens 10 or fewer, free or monadelphous. 3. Mimoseas. Flowers minute, in dense heads or sjhkes. Corolla re- s' GO XLII. LEGTJMINOSiE. gular, its petals free or united in a tube, valvate in aestivation. Stamens definite or indefinite. Suborder 1. Papilionacese. Tribe 1. Podalyrie.®. Stamens free. Legume 2-valve, continuous* Shrubs, with simple or palmate-compound leaves. Legume compressed ; leaves sessile, 3-foliolate ; flowers yellow 1. Cyclopia. Legume turgid, woolly ; leaves petioled, simple ; flowers purple 2. Podalyria. Tribe 2. Liparie®. Stamens diadelphous (except in Ccelidivm and Walpersia). Legume 2-valve, continuous. Ovary 1- or several-ovuled. — Shrubs, with simple, exstipulate leaves. Stamens diadelphous. Flowers yellow. Lowest calyx-segment very large, petaloid . . 3. Liparia. Lowest calyx- segment not longer than the rest. Corolla conspicuous. Ovary several-ovuled . 4. Priestleya. Corolla inconspicuous. Ovary 1-ovuled . . 6. Lathriogyne. Flowers purple or purplish, the keel darker . . 5. Amphithalea. Stamens united at base into a short tube, or nearly free. Flowers purplish; leaves sessile, with indexed margins 7. Ccelidittm. Flowers yellow ; leaves petioled, with reflexed margins 8. Walpersia. Tribe 3. G-eniste®. Stamens completely monadelphous. Ovary 2- or several-ovuled. — Shrubs half-shrubs or herbs, with simple or palmately com- pounded leaves. Leaves simple, without stipules. Legume compressed. Calyx equally 5-fid, lobes pungent. Yexil villous 9. Borbonia. Calyx with the lowest segment very narrow. Standard glabrous 10. Bapnia. Legume turgid. Keel subtruncate, shorter than the alee . . .11. Ettchlora. Keel sharply rostrate 12. Crotalaria. Leaves palmately 3-5-7-foliolate (in some Lebeckia and in Lotononis monophylla , 1-foliolate, hence apparently simple). Legume very turgid. Keel sharply beaked ; flowers racemose 12. Crotalaria. Legume not turgid (or but slightly so). Lateral calyx segments connate in pairs ; front segment separate and narrower. Standard erect, straight, concave ; keel and short style straight 13. Pleiospora. Standard reflexed ; keel indexed, long style sharply-bent. Legume straight or falcate 14. Lotononis. Legume repeatedly folded and twisted from side to side 15. Listia. XLTI. LEGUMINOSiE. G7 Calyx distinctly 2-lipped, the upper lip 2-fid, lower 3-fid. Legume linear, compressed. Calyx deeply divided. Corolla conspicuous. Standard longer than the keel j legume silky 16. Argyrolobium. Standard shorter than the keel ; legume torulose 17. Dichilus. Calyx shortly 2-lipped. Corolla small. Le- gumes glandular or viscid-pubescent . - 18. Melolobium. Calyx bell-shaped, hollow or intruse at base, shortly 5-fid. Standard much longer than the keel. A large shrub 19. Hypocalyptus. Standard much shorter than the keel. A slender undershrub 20. Loddigesia. Calyx oblique, 5-toothed or 5-fid. Stipules 0. Legume linear, flat, terete or turgid, several- seeded 21. Lebeckia. Legume ovate or oblong, winged on upper side, 1-seeded 22. Yiborgia. Calyx bell-shaped, acute at base, 5-toothed or * 5-fid. Leaves petioled, 3-foliolate, mostly with stipules 23. Buchenrcedera. Leaves sessile ; leaflets as if tufted leaves, without stipules 24. Aspalathus. Tribe 4. Psoralie^:. Stamens diadelphous. Ovary 1-ovuled. — Leaves never stipelled, variously compound, commonly sprinkled with resinous, glandular dots. The only genus 25. Psoralea. Tribe 5. Trifolie.®. Stamens diadelphous. Ovary 2- or several- ovuled. — Small herbs. Stem erect or trailing, but not climbing. Leaves palmately or pinnately'3-foliolate, rarely 5-foliolate, never stipelled. Keel very acute. Legume cylindrical, many-seeded 26. Lotus. Keel obtuse. Keel adnate to the wings. Legume hidden in the calyx 27. Tripolium. Keel free. Legume longer than calyx. Legume oval or oblong, 1-4-seeded. Flowers racemose 28. Melilotus. Legume linear, curved, many-seeded. Flowers subumbellate 29. Trigonella. Legume much-incurved or spirally twisted, often bordered with spinous teeth .... 30. Medicago. Tribe 6. Indigofere^:. Stamens diadelphous ; anthers apiculate ! Ovary 2- or many-ovuled. — Stems not twining. Leaves variously com- pound, rarely 1-foliolate. Flowers racemose. Pubescence often strigose. Standard reflexed ; keel with a spur at each side ; flowers red purple or white 31. Indigofera. Tribe 7. Oalege.®. Stamens monadelphous or diadelphous. Ovary 2- or several- ovuled. Legume 2-valved, 1-celled. — Stem not twining. Leaves pinnate, very rarely 1-foliolate, sometimes stipelled. Flowers racemose. Legume coriaceous or rigid, not membranous. F 2 08 XLII. LEGUMINOSiE. Flowers purple pink or white. Small shrubs undershrubs or herbs. Legume coriaceous, compressed, linear 32. Tephrosia. Trees. Legume hard woody with thick mar- gins, elliptic or lanceolate, few-seeded, slowly opening 33. Millettia. Flowers yellow. Legume very long, slender, nodose 34. Sesbania. Legume membranous, pellucid, compressed or bladdery. Leaves pinnate. Flowers racemose. Standard shorter than the acute keel . . . .35. Sutherland ia. Standard longer than the obtuse keel . . . 36. Lessertia. Leaves simple. Flowers axillary, minute . . . 37. Sylitra. Tribe 8. Astragale^e. Stamens diadelphous. Legume completely or incompletely longitudinally 2-celled, by the introflection of one of the sutures. — Stem not twining. Leaves pinnate. Legume with its lower {carinal) suture introflexed . 38. Astragalus. Tribe 9. Hedysareje. Legume more or less completely jointed, usually separating at maturity into indehiscent, 1-seeded fragments, sometimes re- duced to a single joint. — Leaves variously compound or simple. Leaflets 2-4, from the apex of a common petiole {/pellucid- dotted) 39. Zornia. Leaves pinnate ; leaflets in 2 or many pairs. Stamens 10, in two 5-androus sets. Legume jointed 40. iEsCHYNOMENE. Stamens monadelphous. Pod subterraneous, in- dehiscent 41. Arachis. Leaves pinnately 3-foliolate. Calyx tube very long ; corolla inserted in its throat 42. Stylosanthus. Calyx short, 2-lipped ; corolla inserted in its bottom. Legume many-jointed, spontaneously break- ing up 43. Desmodium. Legume imperfectly jointed, not spontaneously separating 44. Anarthrosyne. Leaves simple or 1-foliolate. Unarmed undershrubs or herbs. Legume separating into 2 or more joints . . 45. AlySICARPUS. Legume 1-seeded, indehiscent ; flowers axillary. Leaves obcordate ; stipules free 46. Requienia. Leaves cordate or lanceolate ; stipules adnate 47. Hallia. Spinous shrubs. Legume irregularly constricted, indehiscent 48. Alhagi. Tribe 10. ViciEjE. Stamens diadelphous, or monadelphous above the base. — Herbs, with abruptly pinnate leaves, the common petiole prolonged into a tendril or excurrent point. Style with a tuft of hairs below the stigma . . .49. Vicia. Tribe 11. Phaseole.®. Stamens diadelphous, or monadelphous above the base. Legume 2-valved, 1-celled. — Stem frequently climbing, prostrate or diffuse. Leaves usually pinnately-3-foliolate, sometimes 1-foliolate (in A brus multijugate), stipellate. XLII. LEGUMINOSiE. 69 Leaves 3-foliolate. Ovary more than 2-ovuled. Calyx tubular, obliquely truncate, entire . . .50. Dumasia. Calyx 4-5-lobed or toothed, or cleft down one side. Standard oblong, incumbent, much longer than the wings and keel. Stamens exserted . . 53. Erythrina. Standard spreading, not much, if at all longer than the wings and keel. Stamens included. Standard with 2 prominent, callous ridges on the vaulted claw, within. Calyx subequally 4-5-fid. Style channelled, with a hooked point and oblique stigma. 55. ViGNA. Calyx 2-lipped, upper lip very large, of 2 rounded lobes, lower small 54. Canavalia. Calyx 2-lipped, upper lip short, 2-fid, lower 3-fid 56. Dolichos. Standard without callous ridges on the claw. Stamens monadelphous. Flowers minute 51. Teramnus. Stamens diadelphous. Calyx 4-fid. Flowers small . . . .52. Galactia. Calyx 5-fid. Flowers conspicuous. Plant viscidly hairy 57. Fagelia. Leaves 3-foliolate (1-foliolate or pinnate). Ovary 2-ovuled. Ovary glabrous or pubescent. Seeds globose- reniform, with a short scar, and subcentral seed cord 58. Rhynchosia. Ovary hirsute. Seeds oblong, with a linear scar, and a seed cord affixed near its end (excentric) 59. Eriosema. Leaves abruptly pinnate, in many pairs. Ovary many-ovuled. Seeds globose, scarlet and black 60. Abrtjs. Tribe 12. Dalbergiea:. Stamens monadelphous or variously combined. Legume either completely indehiscent, or rarely splitting eventually into rigid, ligneous valves. — Stem woody arborescent or shrubby, sometimes climbing. Leaves pinnate, with few or many leaflets (rarely 1-foliolate). Leaflets opposite 61. Lonchocarptjs. Leaflets alternate. Anthers versatile. Legume orbicular . . . .62. Pterocarptts. Anthers terminal. Legume oblong or linear . . 63. Dalbergia. Tribe 13. Sophorea:. Stamens free. — Stem woody arborescent or shrubby. Leaves pinnate in many pairs, rarely 1-foliolate. Leaves pinnate. Keel blunt, straightish ; legume torulose . . . 64. Sophora. Keel sharply rostrate ; legume compressed ; flowers purple 65. Virgilia. Keel incurved, blunt ; legume flattened, sharp edged ; flowers yellow 66. Calpgrnia. Leaves 1-foliolate. Corolla very open .... * 67. Bracteolaria. Suborder 2. Csesalpinieae. Leaves simply pinnate. Stamens 10 ; anthers splitting lengthwise. 70 XLII. LEGUMINOSiG. Ovary sessile. Legume linear, very long, many- seeded 68. Parkinsonia. Ovary stipitate. Legume broadly oblong, few- seeded 74. Schotia. Stamens (fewer than 10 perfect) opening by ter- minal pores 73. Cassia. Leaves 2-pinnate. Half-herbaceous plants ; all parts sprinkled with black dots 70. Melanosticta. Arborescent. Legume covered with sharp prickles .... 69. Guilandina. Legume unarmed. Flowers pedicelled. Filaments as long as petals, hairy below i 71. Peltophorum. Flowers sessile. Filament and style very short 72. Bttrkea. Leaves simple, 2-lobed (formed of 2 confluent leaflets) 75. Bauhinia. Suborder 3. Mimoseae. Tribe 1. Eumimose^s. Stamens definite (10) ; pollen powdery. Flowers uniform, spiked, sessile. Legume breaking into joints 76. Entada. Flowers uniform, racemose, pedicelled. Legume continuous 77. Elephantorhiza. Flowers of two kinds in the spike ; the upper fer- tile, the lower neuter, with long, thread-like, barren filaments 78. Dichrostachys. Flowers uniform, capitate, sessile. Legume 1-seeded, semiorbicular 79. Xerocladia. Tribe 2. Acaciea:. Stamens indefinite ; pollen in small masses. Corolla small, tubular ; stamens free 80. Acacia. Corolla funnel-shaped ; stamens shortly tubular at base 81. Albizzia. Corolla narrow funnel-shaped ; stamens connate in a long, exserted tube 82. Zygia. Suborder 1. Papilionacese. (Glen. 1-67.) Tribe 1. Podalyrieal (Glen. 1-2.) 1. CYCLOPIA, Vent. Calyx subequally 5-cleft, with the base indented. Petals subequal ; standard roundish, plaited at base, with a short, re- curved claw ; wings oblong, with a cross fold ; keel incurved, bluntly beaked. Stamens separate, or slightly connate at base ; filaments dilated. Ovary glabrous, several-ovuled. Le- gume oblong, compressed, 1-locular, 2-valved, coriaceous. Seeds strophiolate. — FI. Cap. ii. p. 6. South African shrubs. Leaves sessile, palmately 3-foliolatc ; leaflets narrow, linear or lanceolate, rarely ovate, glabrous or pubescent, often with XLII. LEGUMINOSiE. 71 revolute margins. Stipules 0. Peduncles axillary, 1-flowered, 2-bracted at base; flowers bright yellow. — 9 species, chiefly Western and South Western ; only 1 Eastern. 2. PODALYRIA, Lam. Calyx widely campanulate, subequally 5 -cleft, with the base indented. Standard ample, rounded-emarginate, with a short, recurved claw ; wings obovate, oblique, rather shorter than the standard, longer than the broad, obtuse keel. Stamens separate, or slightly connate at base. Ovary sessile, hairy, many-ovuled. Legume turgid, leathery, villous. — FI. Cap. ii . p. 9. South African, silky or silvery shrubs. Leaves simple, alternate, ex- panded. Stipules subulate, deciduous. Peduncles 1-2 or rarely 3-4- flowered. Bracts solitary, falling off before the opening of the flowers. Flowers purple rosy or bluish- white. — 17 species, chiefly Western and South-Western. Tribe 2. Liparie^]. (G-en. 3-8.) 3. LIPARIA, Linn. Calyx indented at base, with a short tube, 5-lobed ; the 4 upper lobes lanceolate, acute, the lowest very large and broad, petaloid. Corolla glabrous ; standard oval-oblong ; the wings oblong, one infolding the other in the bud ; keel straight, acute, narrow. Stamens diadelphous. Ovary sessile, few-ovuled. — FI. Cap. ii. p. 14. South African shrubs. Leaves alternate, simple, lanceolate, rigid, pun- gent, exstipulate. Flowers bright yellow, in terminal heads. — 4 species, all Western. 4. PRIESTLEYA, DC. Calyx subequally 5-cleft, the lowest lobe equal to the rest, or scarcely longer. Corolla glabrous ; standard roundish, shortly clawed; wings obtuse, subfalcate; keel incurved, ’without late- ral processes. Stamens diadelphous. Ovary sessile, several- ovuled. Legume piano-compressed, 4-6-seeded. — FI. Cap. ii. p. 15. South African shrubs. Leaves alternate, simple, exstipulate. Flowers yellow, in terminal heads or racemes, or axillary. — 15 species, very few Eastern. 5. AMPHITHALEA, E. and Z. Calyx subequally 5-cleft. Standard roundish, shortly clawed, reflexed ; wings oblong ; keel straightish, obtuse, spurred on each side. Stamens diadelphous. Ovary 1-4-ovuled. Legume ovate, 1-2-seeded, rarely oblong, 3-4-seeded. — FI. Cap. ii .p. 21. ; Thes. Cap. 1. 184. Small, generally heath-like, South African shrubs. Leaves alternate, sim- XLII. LEGUMINOSiE. 72 pie, entire, sessilo, exstipulate, often with revolute margins. Flowers purple or rosy, with the keel dark-tinted, axillary and subsessile or crowded in a leafy spike. — 9 species, all but 3 either Western or South-Western. 6. LATHBIOGYNE, E. and Z. Calyx, ovary, and legume, as in the 1-ovuled species of Am- phitJialea. Corolla scarcely longer than the calyx \ the keel incurved, beaked. — FI. Cap. ii. p. 593. Only 1 species, L. jparvifolia, E. and Z., found in Hott. Holl. and Zwarteberg Mountains. A small, twiggy shrub, 12-18 inches high. Leaves 4 lines long, lanceolate, flat, silky and silvery. Flowers 2-4, capitate, ter- minal, yellow, almost hidden in the very hairy calyx. 7. CCELIDIUM, Vogel. Calyx nearly equally 5-fid. Standard obovate, shortly clawed, reflexed ; wings oblong ; keel oblong, straight, obtuse, bluntly spurred at each side. Stamens monadelphous, the tube often very short. Ovary 1-ovulate. Legume ovate, 1-seeded. — FI. Cap. ii. p. 24. Small, much-branched South African shrubs or undershrubs. Leaves simple, entire, sessile, exstipulate, with the margin mostly inflexed, closely pubescent on the upper, either glabrous or silky on the lower. — 8 species, chiefly South-Western. 8. WALPERSIA, Harv. Calyx campanulate, 5-cleft ; the two upper lobes broader than the 3 lower. Petals nearly of equal length, all attached to the base of the stamina! tube ; standard ovate, with a small callosity at the top of the claw ; wings oblong, eared at base ; keel subincurved, bluntly spurred at each side. Stamens shortly monadelphous, 5 longer. Ovary 2-ovuled ; style subu- late. Legume P — FI. Cap. ii. p. 26. W. burtonioides, H., the only species, was found by Zeyher at Glassen- bosch. A small shrub. Leaves petioled, linear, with reflexed margins. Flowers axillary, yellow. Calyx 2-bracted at base. Ovary sessile, silky, with a long style. A very rare, little known plant. Tribe 3. Geniste^:. (Gen. 9-24.) 9. BORBONIA, Linn. Calyx acute at base, equally 5-cleft, the segments pungent. Standard hairy, emarginate ; keel obtuse. Stamens 10, mona- delphous, with a split tube. Ovary 2- or several-ovuled ; style filiform ; stigma capitate. Legume linear, compressed, longer than the calyx, several-seeded (rarely 1-2-seeded). — FI. Cap. ii. p. 27. South African shrubs or undershrubs. Leaves alternate, simple, very rigid, XLII. LEGUMINOS^!. 73 many-nerved, sessile or amplexicaul, exstipulate. Flowers yellow, axillary or terminal, scattered or racemose. — 13 species, chiefly Western and South- Western. 10. RAFNIA, Thunb. Calyx unequally 5-fid, the lowest segment narrowest. Co- rolla glabrous; standard roundish; keel incurved, either sharply beaked or obliquely truncate. Stamens 10, monadelphous. Ovary sessile or stipitate, many-ovuled ; stigma capitate. Legume lanceolate or linear, the upper suture sharp or some- what winged. — FI. Gap. ii. p. 31. Glabrous and frequently glaucous South African shrubs and half-shrubs. Leaves simple, very entire, alternate, exstipulate. Flowers yellow. — 22 species, very few Eastern. 11. EUCHLORA, E. and Z. Calyx deeply 5-cleft, the lowest segment much narrower than the rest. Corolla glabrous ; standard long-clawed, roundish, reflexed ; wings obtuse, longer than the subtruncate keel. Sta- mens monadelphous, with a slit tube. Ovary few-ovuled, hairy ; style glabrous. Legume swollen, ovate, few-seeded. — FI. Cap. ii. p. 38. E. serpens , E. and Z., the only species, is a small, prostrate undershrub. Stem, branches, and leaves hairy. Leaves lanceolate, sessile. Peduncles terminal. Flowers small, purplish, in a dense subcap itate spike. Western districts. 12. CROTALARIA, Linn. Calyx sub-2-labiate, the upper lips 2-fid, the lower 3-fid. Standard large, cordate ; keel falcate-acuminate. Stamens mo- nadelphous. Ovary 2- or many-ovuled; style long, sharply-bent, often laterally pubescent. Legume turgid, with very convex valves, sessile or stipitate, few- or many-seeded. — FI. Cap. ii. p. 39. A large genus of both hemispheres, mostly tropical. Leaves either simple or palmately 3-5-7-folioled, with or without stipules. Flowers ra- cemose or subsolitary, yellow, rarely purple. — 24 South African species, dispersed. 13. PLEIOSPOJRA, Harv. Calyx ovoid, 5-fid, 4 upper segments approaching in lateral pairs, the lowest narrowest. Standard straight, vaulted ; wings patent ; keel straight. Stamens monadelphous, with a split tube. Ovary sessile, tapering into a subulate, straight style ; ovules numerous ; stigma simple. Legume ? — FI. Cap. ii. p. 47. P. cajanifolia , H., the only species, is a shrub, with the aspect of a Pso- ralea. Leaves 3-foliolate ; leaflets broadly lanceolate, silky. Stipules seta- ceous. Peduncles terminal and axillary, subcorymbose, each bearing glo- bose or oblong, spicate heads of flowers. Magalisberg and Crocodile river. 74 XLII. LEGUMINOSiE. 14. LOTONONIS, DC. Calyx subequally 5-fid, the lowest segment narrower than the rest, and unconnected with them, the four upper approach- ing in pairs, and more or less connate into 2 2-fid lobes, rarely quite separate, and then all the segments subequal. Standard obcordate or oblong, commonly pubescent; keel obtuse or acute. Stamens monadelphous. Ovary many-ovuled. Le- gume oblong or linear, more or less compressed, many-seeded. — FI. Cap. ii .p. 47. A large genus, chiefly South African (a few European and Asiatic). Leaves very generally 3-foliolate, rarely 5-foliolate, in one case 1-foliolate. Flowers various, racemose umbelled capitate or solitary, yellow, rarely purple. 15. LISTIA, E. Mey. Calyx 3-fid, the front segment subulate, the lateral broader, each 2-dentate. Keel obtuse, longer than the standard and wings. Stamens monadelphous. Legume linear, compressed, many-seeded, repeatedly folded and twisted from side to side. — FI. Cap. ii. p. 66. L. heteropJiylla , E. Mey., the only species, is a small procumbent plant, nearly glabrous. Leaves 3-foliolate ; leaflets narrow-cuneate or lanceolate- oblong. Peduncles 1-2 inches long, bearing 6-8 subumbellate yellow flo wers . — E astern . 16. ARGYROLOBIUM, E. and Z. Calyx campanulate, deeply cleft, 2-lipped, the upper lip 2- fid, lower 3-fid or 3-toothed. Standard ample, longer than the keel. Stamens monadelphous. Ovary many-ovuled. Legume linear, compressed, silky, not glandular, many-seeded. — FI. Cap. ii. p. 67. Small shrubs, undershrubs or herbs, generally silky or silvery, some South European and Asiatic. Leaves 3-foliolate, stipuled. Flowers yellow, peduncled or pedicelled, solitary racemose or subumbelled. — 30 Cape spe- cies, dispersed. 17. DICHILUS, DC. Calyx as in Argyrolobium. Standard oblong, shorter than the blunt keel. Stamens monadelphous. Ovary many-ovuled; stigma minute. Legume linear, compressed, subtorulose. — FI. Cap. ii. p. 77. Erect or diffuse, subglabrous half-shrubs. Leaves 3-foliolate. Peduncles 1-2-3-flowered ; flowers yellow. — 3 species, all South African and Eastern. 18. MELOLOBIUM, E. and Z. Calyx tubular, shortly or deeply 2-lippcd, the upper lip 2- partite, lower 3-fid or 3-toothed. Corolla not much longer XLII. LEGUMINOSiE. 75 than the calyx ; standard oblong ; keel blunt. Stamens mona- delphous. Ovary several-ovuled. Legume linear, compressed, torulose, glandular or hairy. — j FI. Cap. ii. p. 77. Small South African shrubs or half- shrubs, more or less -viscid or glan- dular, variously pubescent. Leaves petioled, stipuled, 3-foliolate. Flowers yellow, in spikes or spicate racemes, with 3 bracts under each flower. — About 11 species, dispersed. 19. HYPOCALYPTUS, Thunb. Calyx widely bell-shaped, shortly 5-toothed, hollowed at base. Standard roundish, reflexed, longer than the wings and keel. Stamens 10, monadelphous. Ovary lanceolate, many-ovuled. Legume linear, flat, the upper suture thickened, many-seeded. — FI. Cap. ii. p. 81. H. obcordatus , the only species, is a glabrous, densely leafy shrub, with palmately 3-foliolate, stipulate leaves and purple flowers. — North-Western. 20. LODDIGES1A, Sims. Calyx of Hypocalyptus. Standard much shorter than the wings and keel. Legume ovato-lanceolate, acute at each end, flat, the upper suture thickened, few-seeded. — FI. Cap. ii .p. 82. L. oxalidifolia , the only species, is a small, erect or diffuse, glabrous un- dershrubs, with palmately 3-foliolate leaves, and small purple and white flowers, in short terminal racemes. Standard and wings white ; keel dark purple at the point. — Caledon and Swellendam. 21. LEBECKIA, Thunb. Calyx obliquely bell-shaped, shortly 5-toothed, with rounded interspaces, rarely 5-cleft. Keel obtuse or subrostrate, longer than the wings, and usually than the standard. Stamens mona- delphous. Ovary linear, sessile or stipitate, many-ovuled. Le- gume linear, either flat subcompressed terete or turgid. — FI. Cap. ii. p. 82. South African shrubs or undershrubs, very diverse in habit. — 24 species, dispersed, but chiefly Western. They are arranged under five sections, as follows : — I. Stiza. Legumes flat. Rigid, spiny shrubs, with 1-foliolate leaves. (3 species.) 2. Phyllodiastrttm. Legume flat. Glabrous and glaucous, unarmed lialf-shrubs and herbs, with filiform leaves. (4 species.) 3. Eulebeckia. Legume narrow linear, terete or subterete. Glabrous and glaucous half-shrubs, with filiform leaves. (4 species.) 4. Calobota. Legume terete or turgid. Keel obtuse, longer than wings. Shrubs or half-shrubs, pubescent or canescent, with simple or 3-foliolate, flat leaves. (10 species.) 5. Viborgioides. Legume terete or turgid. Keel subrostrate, scarcely longer than wings. Rigid, unarmed shrubs, with subsessile, 3-foliolate leaves. (3 species.) 76 XLII. LEGUMINOSiE. 22. VIBORGIA, Thunb. Calyx oblique, shortly 5-toothed. All the petals with long, slender claws ; standard ovate ; keel incurved or rostrate, longer than the others. Stamens monadelphous. Ovary stipitate, few-ovuled. Legume stipitate, ovate or rarely oblong, inde- hiscent, winged on the upper suture, sharp and thin along the lower. — FI. Cap. ii. p. 90. Rigid, slender, sometimes spiny, South African shrubs, with palmately 3-foliolate leaves, and yellow, racemose flowers. — 7 species, natives of the Western and Northern districts 23. BUCHENRGEDERA, E. and Z. Calyx bell-shaped, subequally 5-fid. Petals villous, on long claws ; keel roundish, short and blunt. Stamens monadelphous. Ovary 8-10-ovuled. Legume obliquely ovate, somewhat tur- gid, 1-3-seeded. — FI. Cap. ii. p. 92. Densely silky or silvery small shrubs or half-shrubs, with petioled and sti- puled, 3-foliolate leaves, and white or purple flowers. — 8 species, all Eastern and beyond the Eastern frontier. Easily known from Asjpalathus by the petioles and stipules. 24. ASPALATHUS, Linn. Calyx bell-shaped, subequally 5-toothed or 5-cleft, or the 2 upper lobes shorter and broader. Standard short-clawed, erect, keeled at back, spreading ; keel incurved or rarely straight. Stamens monadelphous, with a split tube. Ovary 2-4-8- or rarely many-ovuled ; style glabrous, incurved. Legume ob- liquely ovate or sublanceolate, subcompressed, acute, 1- or few- seeded. — FI. Cap. ii. p. 94. A large and most natural South African genus of shrubs and half-shrubs, with heath-like or furze-like, rarely flat and broadish, sessile, entire leaves, without stipules. — About 150 species, arranged under 12 sections (fully de- scribed in FI. Cap.), which may be recognized by the following Key : — Flowers sessile or short-stalked, solitary or spiked, capitate Or racemose. Leaves flat (broad or narrow) . Leaves glabrous or roughly hairy (not silky) . 1. CephalanthtE. Leaves silky or very softly and closely hairy . 2. Serice^:. Leaves terete or 3-sided, linear or subulate. Claws of keel and wings attached to the staminal tube 3. Synpetal^:. Claws of keel and wings quite free from the sta- minal tube. Legume obliquely ovate, short. (Flowers small) 4. Leptanthje. Legume villous, turgid, ovate or lanceolate, reflexed. (Flowers sessile, lateral, mediocre) 5. Laterales. Legume linear-lanceolate, many-seeded . . 6. MacrocarPjE. XLII. LEOUMINOSiE. Legume thick, villous, obliquely lanceolate, several-seeded. Flowers lateral or subterminal, 1-2 together 7. GrafdieloR/E. Flowers terminal, subcapitate 8. Pachycarpa:. Legume glabrous or silky, obliquely lanceolate. Leaves fleshy. Flowers mediocre or large, subsessile. Legume glabrous .... 9. Carfosa:. Leaves fleshy. Flowers small, sessile. Le- gume glabrous or silky 10. PiFGUES. Leaves not fleshy. Flowers terminal, soli- tary, in pairs or racemose 11. Termifales. Flowers 1 or few, at the end of a long filiform peduncle 12. Pedufculares. These shrubs are dispersed over South Africa, chiefly in dry, stony or sandy places. Tribe 4. Psoralie^i. (Gren. 25.) 25. PSORALEA, Linn. Calyx campanulate, unequally 5-lobed, the lowest lobe longer and usually broader than the rest. Standard broad, with reflexed sides ; keel shorter, dark-coloured. Stamens diadel- phous. Ovary sessile, 1-ovuled ; style slender. Legume con- cealed in the calyx, 1-seeded, indehiscent. — FI. Cap. ii. p. 143. A large, widely-dispersed, tropical and subtropical genus of shrubs half- shrubs or herbs, in most cases copiously sprinkled with resinous black or pellucid dots, and strongly resin-scented. Leaves pinnate or 3-foliolate, rarely 1-foliolate. Stipules free or attached to the petiole. — 41 South African species, dispersed. Tribe 5. Trieolie^. (G-en. 26-30.) 26. LOTUS, Linn. Calyx bell-shaped, 5-cleft or 5-toothed. Standard roundish, spreading, recurved, equalling the prominent, connivent wings ; keel ascending, narrow, rostrate. Stamens diadelphous. Style ascending, subulate. Legume linear, terete or subcompressed, many-seeded, 1-celled, or having septa between the seeds, when ripe splitting into 2 valves. — FI. Cap. ii. p. 157. A nearly cosmopolitan genus of herbs or half- shrubs, chiefly from tem- perate zones. Leaves 3-foliolate. Stipules in pairs or connate, free, large resembling the leaflets. Peduncles umbellately 2- or many-flowered, with leafy bracts under the flowers. — 1 South African species, from Natal and Transvaal ; 3 other doubtful species. 27. TRIFOLIUM, Linn. Calyx bell-shaped or tubular, unequally 4-cleft or toothed (sometimes inflated after flowering). Corolla persistent, the standard longer than the wings and keel. Stamens diadelphous. Legume minute, 1-4-seeded, enclosed in the calyx, indehiscent. — FI. Cap. ii. p. 158. 78 XLII. LEGUMINOSiE. Herbs, dispersed ovor the globe : “ Trefoil,” “ Clover,” etc. — There are 7 species found in South Africa, of which 3 are endemic, the rest probably introduced from Europe. 28. MELILOTUS, Tournef. Calyx bell-shaped, subequally 5-toothed. Corolla deciduous ; standard and wings longer than the obtuse keel. Stamens diadelphous. Legume oval or oblong, 1-4-seeded, longer than the unaltered calyx, indehiscent. — FI. Cap. ii. p. 161. Annuals or biennials, strongly scented, of Northern origin. Leaves pinnately 3-foliolate, the leaflets sharply and coarsely toothed. Flowers small, yellow or whitish, in racemes. M. parvijiora , Desv., is a weed near cultivation. 29. TRIGONELLA, Linn. Calyx bell-shaped, 5-toothed. Corolla deciduous ; standard and wings spreading; keel obtuse. Stamens diadelphous. Le- gume linear or oblong-linear, compressed or terete, acuminate, many-seeded. — FI. Cap. ii. p. 161. Strongly-scented herbs, chiefly from the northern hemisphere. Leaves pinnately 3-foliolate. Flowers racemose, umbelled or subsolitary. T. ha- mosa, Linn., is found in the Eastern district. 80. MEDICAGO, Linn. Calyx campanulate, subequally 5-toothed. Standard longer than the wings and the blunt keel. Stamens diadelphous. Stigma capitate. Legume 1- or many-seeded, spirally twisted or falcate. — FI. Cap. ii. p. 162. Herbs, abundant in Central and Southern Europe and Middle Asia, na- turalized in various countries. Leaves pinnately 3-foliolate, very rarely im- paripinnate. Flowers minute, yellow or purple. — 4 species, naturalized at the Cape. Tribe 6. Ikdigoeerejg. (Gen. 31.) 81. INDIGOEERA, Linn. Calyx small, bell- shaped, 5-fid or 5-toothed. Standard round- ish, reflexed ; keel with a spur or prominence at each side, near the base. Stamens diadelphous ; the connective of the anthers apiculate ! ovary 2- or several-ovuled. Legume linear, terete, compressed or flattened, 1- or several-seeded, mostly with septa between the seeds. — FI. Cap. ii. p. 163. A vast genus, found in all hot countries. Shrubs undershrubs or an- nuals. Leaves imparipinnate or digitate, 3- or many-folioled, rarely 1- foliolate or suppressed. Hairs commonly rigid, fixed by a middle point, and set in subparallel lines ; some are softly hairy or villous, a few glabrous. Flowers purple rosy or white, mostly racemos’e. — About 120 South African species (several new, as yet undescribed), dispersed. XLII. LEGUMINOSiE. 79 Teibe 7. G-ALEGEiE. (Gen. 32-37.) 32. TEPHROSIA, Pers. Calyx ebracteolate, bell- shaped, subequally 5-toothed or cleft. Standard suborbicular, large, spreading, silky or villous externally ; wings adhering to the keel. Stamens monadelphous or diadelphous. Ovary many-ovuled ; style filiform, glabrous or bearded. Legume linear, compressed, coriaceous, straight or curved, sessile or stipitate, continuous or with partitions between the seeds ; seeds compressed. — FI. Cap. ii. p. 203. A considerable genus of hot countries. Shrubs half-shrub3 or herbs, with imparipinnate, rarely digitate or 1-foliolato leaves and free stipules. Flowers racemose, red purple or white. — 21 South African species, chiefly from the Eastern district and frontier, Caflraria, and Natal. 33. MILLETTIA, W. and A. Calyx urceolate, bluntly toothed. Standard recurved, emar- ginate, longer than the wings, which are longer than the keel. Stamens imperfectly monadelphous, the upper stamen free at base. Legume elliptical or lanceolate, few-seeded, hard and woody, with thickened margins, tardily splitting open. — FI. Cap. ii. p. 211. Trees of subtropical Africa and Asia. Leaves pinnate ; leaflets opposite, stipelled. Racemes or panicles axillary or terminal. Flowers purple or reddish. — 2 South African species, both from Natal. 34. SESBANIA, Pers. Calyx 2-bracteolate, cup-shaped, subequally toothed or cleft. Petals subequal in length ; standard roundish, complicate, crested on the claw or nude ; wings oblong ; keel long- clawed, ascending, sharply eared or toothed at base. Stamens diadel- phous, the tube wide and eared at base. Legume very long, slender, compressed or cylindrical, with thickened sutures, constricted between the seeds, and divided by cross septa into many 1-seeded loculi. Seeds cylindrical, oblong. — FI. Cap. ii. p. 212. Tropical shrubs or tall herbs. S. aculeata, a tall, mostly glabrous and glaucous shrub-like annual, several feet high, grows in the Natal country. Leaves abruptly pinnate, inultijugate ; leaflets linear, obtuse, mucronulate. Racemes short ; flowers yellow. Pod 1 foot long, not 2 lines wide. 35. SUTHEELANDIA, B. Br. Calyx bell-shaped, 5-toothed. Standard oblong, shorter than the oblong, boat- shaped keel, its sides reflexed; wings very short. Stamens diadelphous. Ovary stipitate, many-ovuled ; style bearded along the upper side and in front below the terminal 80 XLII. LEGUMINOSjE. stigma. Legume papery, inflated, many-seeded, indehiscent. Seeds reniform. — FI. Cap. ii. p. 212. S.fruteseens , the only species, is a shrub, very variable in pubescence, mostly hoary or canous. Leaves imparipinnate, multijugate. Flowers handsome, scarlet or bright red, in axillary racemes. On dry hills through- out the colony. 36. LESSERTIA, DC. Calyx bell-shaped, shortly and subequally 5-toothed. Stand- ard obovate, emarginate, expanded, longer than the blunt keel. Stamens diadelphous. Ovary substipitate, several-ovuled ; style filiform, ascending, bearded in front below the terminal stigma. Legume scarious, compressed or inflated, unequal- sided or linear, at length opening at the apex, several-seeded. — FI. Cap. ii. p. 213. South African half-shrubs or herb3, mostly canescent. Leaves impari- pinnate, multijugate; leaflets often alternate, rarely glabrous. Flowers pink crimson or purple, rarely white, in axillary racemes. — About 30 African species, dispersed. 37. SYLITRA, E. M. Calyx sub-2-labiate, 5-fid. Keel erect, round- pointed, shorter than the subequal standard and attached wings. Stamens mo- nadelphous. Ovary 4-ovuled; style glabrous, the stigma capitel- late. Legume scarious, indehiscent, compressed, much broader than the seeds. — FI. Cap. ii. p. 224 ; Thes. Cap. t. 78. S. bifiora is a rod-like perennial, with 1-foliolate leaves, and small axillary flowers. Found near the Gramke river. Teibe 8. Asteagale.®. 38. ASTRAGALUS, Linn. Calyx tubular or bell-shaped, 5-toothed. Standard equalling or exceeding the wings ; keel obtuse. Stamens diadelphous. Ovary many-ovuled. Legume (variable in form) incompletely or completely divided longitudinally into 2 cells by the intro- flection of the carinal (dorsal) suture. — FI. Cap. ii. p. 224 ; Thes. Cap. t, 82. An immense genus, but chiefly of the northern hemisphere. A. Burlcea- nus, our only South African species, is a glabrous and glaucous annual, with large, leafy stipules, pinnate leaves, and small, slender, racemose flowers. Found at Magalisberg. Teibe 9. Hedysaeejs. (Gen. 39-48.) 39. ZORNIA, Gmel. Calyx 2-labiate, the upper lip obtuse, emarginate, the lower 3-fid. Corolla inserted in the base of the calyx ; standard roundish, with reflexed sides ; wings oblong ; keel of lunate XLTI. LEGFMTNOSJE. 81 petals cohering in the middle. Stamens monadelphous, the alternate anthers small. Legume sessile, compressed, 3-6- jointed, the joints roundish, often hispid. — FI. Cap. ii.p. 225. Tropical and subtropical herbs or undershrubs. Leaves digitate, of 2-4 pellucid-dotted leaflets. Stipules broad, rigid, peltate. Bracts similar but larger, enclosing the small flowers. — Z. tetraphylla , a widely-distributed plant, occurs in the Eastern district and at Natal. 40. 2ESCHYNOMENE, Linn. Calyx 2-bracteate at base, more or less 2-lipped or 2-parted, the upper lip entire or 2-fid, the lower either entire 3-fid or 3-toothed. Standard roundish or oblong, simple at base ; wings oblong, equalling or exceeding the incurved keel. Stamens 10, in 2 equal parcels, each of 5. Ovary stipitate. Legume sti- pitate, compressed, exserted, transversely jointed ; joints se- veral.— FI. Cap. n.p. 225. Subtropical herbs or shrubs. Leaves pinnate, with many or few pairs of leaflets, stipuled. Peduncles racemose or 1-flowered, axillary, rarely terminal. Flowers yellow or reddish. — 3 species, all from Natal. 41. ARACHIS, Linn. Flowers polygamous. — Male: Calyx-tube very long and slender (resembling a flower stalk) ; limb 2-partite, the upper lip 4-toothed, the lower slender, entire. Corolla inserted in the throat of the calyx ; standard roundish ; wings oblong, free ; keel incurved, rostrate. Stamens monadelphous, inserted with the petals. Ovary concealed in the base of the calyx- tube, subsessile, 2-3-ovuled, abortive. — Female : Calyx, corolla, and stamens 0 ! Ovary on a quickly elongating, rigid, re- flexed (pedicel-like) torus, stipitate, 1-celled, with 2-3 anatro- pous ovules ; style very short ; stigma dilated. Legume (buried underground) oblong, thick, netted, indehiscent, sub- torulose, 2-3-seeded. Embryo straight, with thick cotyledons. — FI. Cap. ii. p. 226. A. Jiypogcea (the Earth Nut) is found near Natal, introduced from tro- pical America. Stems diffuse or trailing. Leaves abruptly 2-jugate ; leaf- lets obovate, obtuse. Flowers solitary, axillary. 42. STYLOSANTHES, Linn. Flowers polygamous. — Male : Calyx-tube very long and slender ; limb deeply 2-lipped, the upper lip 4-fid, the lower long, entire. Corolla inserted in the throat of the calyx ; standard roundish ; wings oblong, free ; keel incurved, rostrate, shorter than the wings. Stamens monadelphous, with a split tube. Ovary sessile, in the base of the calyx-tube, commonly abortive ; style filiform, very long. — Female : Calyx, corolla, a 82 XLTT. LEGTJMINOSiE. and stamen 0. Ovary subsessile, erect, 2-ovuled ; style short, hooked. Legume sessile, mostly 2-jointed ; joints compressed, the lower often sterile, the upper 1-seeded, separating. — FI. Cap. ii. p. 227. Weed-like herbs or undershrubs, frequently viscid-pubescent, common in hot countries. Leaves pinnately 3-folioled. Flowers in dense terminal or axillary spikes, or solitary or in pairs. — S. setosa , our only species, grows at the Aapjes river. 43. DESMODIUM, DC. Calyx 5-parted or deeply 2-lipped, the upper lip 2-fid, lower 3-fid. Standard roundish ; wings oblong, longer than the straight, obtuse keel. Stamens diadelphous. Ovary sessile, many-ovuled. Legume several-join ted, the joints compressed, 1-seeded, membranous or rigid, separating at maturity. Seeds compressed, reniform. — FI. Cap. ii. p. 227. Herbs or half-shrubs, common in warm countries. Leaves pinnately 3-folioled or 1-folioled. Racemes terminal, slender or dense ; flowers small, purple or white. — 5 South African species, natives of Caffraria and Natal. 44. ANARTHROSYNE, E. Mey. Same as Desmodium , but — Legume compressed, linear-sub- falcate, imperfectly jointed, but not spontaneously separating into 1-seeded fragments. — FI. Cap. ii. p. 229. Tropical and subtropical plants, with the habit of Desmodium. A. robusta , E. M., a tall, robust plant, 3 feet high, softly tomentose, is found near Natal. 45. ALYSICARPUS, Neck. Calyx persistent, glumaceous, deeply 4-parted, the upper segment emarginate or 2-fid. Corolla papilionaceous, small, scarcely longer than the calyx. Stamens diadelphous. Le- gume terete or subcompressed, several-jointed, the joints equal-sided, separating. — FI. Cap. ii. p. 230. Small weed-like plants of hot countries. Leaves 1-folioled, 2-stipuled ; stipules and bracts dry. Flowers racemose, pedicelled, in pairs, purple, inconspicuous. — 2 South African species (endemic), found in Natal and Zululand. 46. REQUIENIA, DC. Calyx bell-shaped, 5 -fid, the segments acute, the lowest longest. Standard obovate ; keel obtuse, 2-petalous. Stamens monadelphous, the tube cleft above. Ovary sessile, 1-ovuled ; style short, incurved. Legume oval, compressed, mucronate, 1-seeded. — FI. Cap. ii. p. 230. Tomentose-canescent half-shrubs of doubtful affinity. Leaves alternate, 1-folioled ; leaflets obcordate, closely penninerved, mucronate. Stipules free. Flowers very small, subsessile, solitary or clustered. — It. spheero- sperma, DC., our only species, is found in Transvaal. XLIT. LEGUMINOS/E. 83 47. HALLIA, Thunb. Calyx subequally 5-fid. Standard ovate ; wings oblong, longer than tbe obtuse keel. Stamens completely monadel- pbous. Ovary substipitate, 1-ovuled. Legume compressed, membranous, 1-seeded. — FI. Cap. ii. p. 231. Small, ascending or trailing slender plants, all South African. Leaves alternate, simple, very entire, often black-dotted, 2-stipuled. Flowers axil- lary, solitary, small, purple. — 6 species, dispersed. 48. ALHAGI, Tournef. Calyx shortly 5-tootbed. Standard obovate, complicate ; wings oblong ; keel straight, obtuse. Stamens diadelphous. Ovary several-ovuled ; style filiform. Legume stipitate, ligne- ous, terete, few-seeded, irregularly constricted here and there, but not jointed, indehiscent. — FI. Cap. ii. p. 233. Undershrubs, natives of the deserts of Northern Africa and Central Asia. Leaves simple. Peduncles axillary, spinous ; flowers few, red. — Manna is collected from these plants in the East. A. Maurorum is said to grow in the Karroo ; but requires evidence to establish it. Tribe 10. Yicie^). (Gen. 49.) 49. VICIA, Linn. Calyx bell-shaped, subequally 5-cleft or toothed. Corolla much exserted ; standard expanded. Stamens diadelphous. Ovary subsessile ; the style bent upwards at a right angle, with a tuft of hairs under the stigma. Legume compressed or turgid, 2- or many-seeded. Seeds subglobose, wdth an oval or linear scar. — FI. Cap. ii. p. 233. Annual or perennial, climbing herbs, natives of the northern temperate zone ; naturalized in South Africa. Leaves abruptly pinnate, the com- mon petiole produced into a tendril. Peduncles axillary, 1-2- or many- flowered. Flowers blue purple yellow or white. — V. sativa and V. atro- purpurea , both introduced. Tribe 11. Phaseole^. (Gen. 50-60.) 50. DUMASIA, DC. Calyx cylindrical, obliquely truncate, entire, 2-bracteolate at base. Claws of the petals equalling the calyx ; limb of the standard cordate-oval ; keel obtuse. Stamens diadelphous. Ovary few-ovuled ; style filiform at base and apex, dilated be- yond the middle. Legume tapering at base, 2-valved, com- pressed, few-seeded, contracted between the seeds. — FI. Cap. ii. p. 234. Twining undershrubs, common in tropical Asia. Leaves pinnately-3-folic G 2 84 XLTI. LEQUMINOSJE. late. Racemes axillary. Easily known by its curious calyx. — D. villosa , DC., grows at the Knysna. 51. TERAMNUS, Sw. Calyx tubular-campanulate, 4-5-fid . Standard obovate, with a longish claw ; wings narrow-oblong, oblique ; keel shorter, oblique, obtuse. Stamens monadelphous, the alternate ros- trate. Ovary sessile, with a short, thick style and capitate stigma. Legume linear, many-seeded, hook-pointed, septate within. — FI. Cap. ii. p. 234. Slender, twining, tropical plants. Leaves pinnately 3-foliolate ; leaflets stipellate, the terminal remote. Flowers minute, oh slender, axillary peduncles, in pairs, in tufts or interruptedly racemose.— T. labialis , a common tropical plant, grows in Caffraria and at Natal. 52. GALACTIA, P. Br. Calyx 2-bracteate at base, 4-fid, the segments acute, nearly equal. Standard ovate or suborbicular, spreading or reflexed ; wings oblong, shorter than the subincurved keel. Stamens diadelphous. Ovary several-ovuled, subsessile ; style filiform, incurved, glabrous ; stigma small. Legume linear, compressed, with cellular partitions between the seeds, several-seeded. — FI. Cap. ii. p. 235. Voluble or prostrate herbs or half-shrubs. Leaves pinnately 3-foliolate ; leaflets stipelled. Racemes axillary, few-flowered. Flowers small. — G. tenuiflora, W. and A., a common coast-plant in hot countries, occurs at Natal. 53. ERYTHRINA, Linn. Calyx either truncate or 2-labiate, or cleft on one side and spathaceous. Standard ovate-oblong, without basal ears or calli, incumbent, very much longer than the wings and the 2-petalous keel. Stamens straight, exserted, diadelphous or incompletely monadelphous. Ovary stipitate, many-ovuled ; style straight, glabrous, with a lateral stigma. Legume indehiscent, com- pressed between the seeds, tipped with the hardened style. Seeds oval, with a linear scar. — FI. Cap. ii. p. 236 ; Tfies. Cap. t. 61, 62. Trees or shrubs, natives of warm countries. Stem and leaves often prickly. Leaves pinnately 3-folioled, the terminal leaflets remote. Stipels glandular. Flowers racemose, large and handsome, scarlet or red. Seeds commonly red and black.— At least 5 (some new undescribed) South African species, natives of the Eastern district, Caffraria and Natal. 54. CANA VALIA, DC. Calyx 2-labiate, the upper lip very large, truncate, emargi- nate or 2-fid, with broadly rounded lobes ; lower small, sub- XLII. LEOUMINOSvE. 85 entire or 3-fid. Standard ample, suborbicular, rigid at back, 2-callous within, with a short claw ; wings oblong, eared at base ; keel equalling the wings or longer, shorter than the standard, incurved. Disk sheathing. Stamens monadelphous or im- perfectly diadelphous. Ovary linear, multi-ovulate ; style incurved, glabrous, with a terminal stigma. Legume com- pressed, subfalcate, with partitions between the seeds. Seeds compressed, with a linear scar. — FI. Cap. ii. p. 238. Climbing or prostrate, tropical or subtropical herbs or undershrubs. Leaves pinnately 3-foliolate, the terminal leaflets subdistant. Stipules small; stipels minute or 0. Racemes axillary, subspicate; flowers solitary .or in pairs, rosy purple or white. — 2 species found near Natal, but neither endemic. 55. VI GST A, Savi. Calyx 2-bracteate at base, bell-shaped, 4-5-fid (the upper lobes separate or connate), the lowest lobe longest. Standard ample, spreading, with an arched and vaulted claw, and 2 cal- lous ridges at base within ; wings oblong, produced at base or eared on the claw ; keel not twisted, indexed or rostrate. Stamens diadelphous or monadelphous. Disk sheathing. Ovary linear, several-ovuled ; style compressed and channelled on one side, incurved ; stigma hooked, oblique. Legume terete or compressed, subfalcate, subtorulose, with cellular partitions between the seeds. Seeds subreniform, with a small strophiole. — FI. Cap. ii. p. 239. Twining or erect, undershrubs or herbs, natives of warm countries. Leaves pinnately 3-folioled. Flowers on long peduncles, floriferous at summit, or racemose. The pods of many are eaten as “French Beans V. Catjang is often cultivated for its pods. — At least 8 species in the Eastern district and Natal, of which 6 are endemic. 56. DOLICHOS, Linn. Calyx bell-shaped, 2-lipped, the upper lip 2-fid or subentire, lower 3-fid. Standard spreading or incumbent, equalling the keel, with 2-4-eallous ridges within ; wings oblong ; keel fal- cate or incurved (or nearly straight), neither twisted nor bent to one side. Stamens diadelphous. Ovary substipitate, several- ovuled ; style channelled or terete ; stigma capitate. Legume compressed, straight or falcate, 2- or several-seeded, with cellular partitions between the seeds. — FI. Cap. ii. p. 242. Twining or prostrate plants of warm countries. Leaves pinnately 3- folioled or rarely 5-folioled, stipellate. Flowers racemose subcorymbose or solitary, 2-bracteolate, red purple blue or white.— 11 species, of which 10 are endemic, chiefly Eastern. 57. FAGELIA, Neck. Calyx 5-cleft beyond the middle, the segments linear, acute, 8G X LIT. LEGUMINOSVE. straight, the 2 uppermost somewhat connate. Standard re- flexed; keel very obtuse, longer than the wings. Stamens diadelphous. Ovary sessile, several- ovuled ; style subulate, glabrous ; stigma obtuse. Legume turgid, about 6-seeded, constricted between the seeds, 2-valved. Seeds ovate, stro- phiolate, with a linear hilum. — FI. Cap. ii. p. 247. A twining, strong-smelling, viscidly hairy shrnb. Leaves pinnately 3- folioled, the terminal leaflet remote. Racemes axillary ; flowers yellow, the keel tipped with purple. — Common in the Western districts. 58. RHYNCHOSIA, Lour. Calyx bell-shaped, mostly oblique, 4-5-fid, the 2 upper lobes more or less united, the lowest longest. Petals nearly of equal length or the wings shorter ; standard obovate or orbicular, mostly with 2 minute, inflexed ears at base, naked or 2-callous within ; wings narrow, eared at base ; keel broader, incurved, obtuse or subrostrate. Stamens diadelphous ; the vexillary filament quite free, mostly knee-bent. Ovary subsessile, with 2 ovules ; style incurved beyond the middle, quite glabrous and mostly thickened above, filiform and often haiiy at base. Legume compressed, oblique or falcate, rarely septate within. Seeds 1-2, compressed with a lateral short or oblong scar, and a subcentral seed-cord. — FI. Cap. ii. p. 247. Climbing or prostrate, rarely erect, herbs or undershrubs, mostly sprinkled with resinous dots, natives of warm countries. Leaves commonly pinnately 3-foliolate, rarely 1-foliolate or pinnate, or 2-3-pinnate, plurijugate. Pe- duncles mostly racemose, rarely umbelled or 1-flowered. Flowers yellow, streaked with brown, rarely purple. — About 30 South African species (some new undescribed), dispersed, but chiefly Eastern. 59. ERIOSEMA, DC. Calyx bell-shaped, 5-fid, the upper lobes sometimes connate. Petals subequal ; stan,dard obovate or oblong, with inflexed ears at base ; wings narrow, longer or shorter than the wider, incurved, obtuse keel. Stamens diadelphous. Ovary sessile, very hairy, 2-ovuled ; style filiform, quite glabrous above the middle, incurved and often thickened upwards ; stigma small or capitate. Legume compressed, obliquely orbicular-rhom- boid or broadly oblong, hairy. Seeds 2-1, compressed, oblong, obliquely transverse, the seed-cord fixed at one end of a linear scar. — FI. Cap. ii. p. 258. Erect or prostrate, rarely twining, herbs or undershrubs of warm countries. Foliage and inflorescence of Rhynchosia , to which this genus is closely allied, and from which it is known by the obliquely transverse seeds and the excentrical seed-cords ; a character easily seen in the unripe fruit. — At least 9 Cape'-species, all Eastern or from Natal or Transvaal. XLII. LEGtJMINOSiE. 87 60. ABRUS, Linn. Calyx bell-shaped, shortly 4-fid or 4-toothed, the upper lobe entire or 2-fid. Standard ovate, about as long as the subfalcate keel. Stamens 9, monadelphous, in a split tube ; no vexillary stamen ! Ovary several-ovuled ; style short, incurved, gla- brous. Legume oblong, compressed, 4-6-seeded, with parti- tions between the seeds. — FI. Cap. ii. p. 262. Diffuse or climbing, slender, woody plants, chiefly of tropical Asia, but naturalized in the tropics generally. Leaves abruptly pinnate, multij ugate. Flowers racemose, orange. Seeds oblong, red, with a black spot round the hilum ; sometimes used as beads. The roots may be used as a substitute for liquorice. — The common A. precatorius, Linn., grows at Natal; also A. Icevigatus , Em., an endemic species. Tribe 12. Dalbergie^. (Gren. 61-63.) 61. LONCHOCARPITS, H. B. K. Calyx truncate, or shortly 4-5-toothed. Wings slightly co- hering above the claw of the keel, whose petals slightly cohere at back, above. Vexillary stamen quite free at base, above Connate with the rest into a complete tube ; anthers versatile. Ovary 2- or several-ovuled. Legume flat, oblong or elongate, membranous coriaceous or ligneous, indehiscent, with a ter- minal style ; the sutures not winged. Seeds if many, distant, compressed ; radicle indexed. — FI. Cap. ii. p. 263. Trees or climbing shrubs. Leaves alternate, imparipinnate ; leaflets op- posite ; stipels few or 0. Racemes simple or panicled ; flowers violet pur- ple or white, but not yellow. — L. Philenoptera , Bth., also a native of Abyssinia, occurs near Lake Ngarai. 62. PTEROCARPUS, Linn. Calyx turbinate- campanulate, acute at base, oftener incurved, 5-toothed, sub -2 -labiate. Petals of the keel at back, near the apex, shortly connate or nearly free. Stamens 10, sometimes monadelphous with a split tube, sometimes equally diadelphous, And sometimes 1 only free ; anthers versatile. Ovary 6-8- ovuled. Legume compressed, indehiscent, orbicular or ovate, more or less oblique or falcate, with a lateral or rarely terminal style, bearing seeds in the middle, more or less indurated or thickened, with a surrounding membranous ring or sharp ridge, sometimes almost completely attenuate-coriaceous or mem- branous. Seeds 1-3, separated by hard partitions. — FI. Cap. ii. p. 264. Unarmed, tropical and subtropical trees. Leaves imparipinnate ; leaflets alternate or irregularly opposite. Racemes simple or panicled ; flowers yellow, rarely whitish, with violet shades ; petals glabrous. — P. sericeus , Bth., our only species, grows by the Ilex and Aapjes rivers. 88 xlh. LEoinvirNosiE. 63. DALBERGIA, Linn. Calyx campanulate, 5-toothed, the upper teeth broader, the lowest longest. Petals of the keel keeled above at the apex. Stamens 10, monadelphous with a split tube, or 9, the vexillary wanting ; or equally diadelphous ; anthers small, erect, didy- mous, shortly opening at the apex, or rarely longitudinally splitting. Ovary stipitate, few-ovuled. Legume oblong or linear, rarely falcate, flat, thin, indehiscent, either 1-seeded or distantly few-seeded, slightly hardened and often netted at the seed, the margins neither thickened nor winged. — FI. Cap. ii. p. 264. Trees or climbing shrubs of warm countries. Leaves imparipinnate, the leaflets exstipelled and mostly alternate. Inflorescence dichotomous, or ir- regular. Flowers small, purple violet or white. — 3 species in Caffraria and at Natal. Tribe 13. Sophore^;. (G-en. 64-67.) 64. SOPHORA, Linn. Calyx widely bell-shaped, obliquely truncate, obsoletely or shortly 5-toothed. Petals of equal length ; standard obovate or roundish, erect or spreading ; wings oblong, clawed, eared at base ; keel obtuse, straight, its petals imbricating and connate in the middle, free above. Stamens 10, free, glabrous. Ovary subsessile, manv-ovuled ; style slightly curved, glabrous. Le- gume constricted at intervals, indehiscent, wingless, several- seeded. — FI. Cap. ii. p. 265. Trees shrubs or herbs, chiefly tropical. Leaves imparipinnate. Flowers racemose, white blue or yellow. — S. nitens , Bth., our only species, is a densely silky-silvery shrub, as yet only found by T. Williamson, near Natal. 65. VIRGILIA, Lam. Calyx widely bell-shaped, shortly 2-lipped, the upper lip 2-fid, lower 3-fid. Standard orbicular, strongly reflexed ; wings oblong ; keel incurved, rostrate. Stamens 10, free. Ovary ses- sile, villous, several -ovuled ; style glabrous. Legume coria- ceous, compressed, tomentose, many-seeded, stuffed between the seeds, the sutures very obtuse. — FI. Cap. ii . p. 266. V. Capensis , the only species, is a tree (Wilde Keureboom). Leaves pinnate, in 6-10 pairs, exstipulate ; leaflets linear-oblong, glossy above, to- mentose beneath. Flowers rosy-purple, in many-flowered racemes. Found along riversides, throughout the colony. 66. CALPURNIA, E. Mey. Calyx widely bell-shaped, shortly 5-fid, the 2 upper lobes semiconnate. Standard erect ; wings oblong ; keel incurved, XLTI. LEUUMINOSjE. 89 obtuse, 2-fid. Stamens 10, free or connate at base, persistent. Ovary stipitate, several-ovuled. Legume membranous, com- pressed, glabrescent, netted, few- or several-seeded, somewhat winged along the ventral suture, the valves cohering between the seeds. — FI. Cap. ii . p. 266. African trees or shrubs. Leaves imparipinnate, multijugate. Racemes axillary and terminal, the peduncle often panicled. Flowers yellow. — At least 6 Cape species, all Eastern. 67. BRACTEOLARIA, Hochst. Calyx 2-bracteolate, deeply 2-lipped, reflexed. Corolla ex- panded; standard ample, suborbicular ; wings spreading widely; petals of the keel shortly connate in the middle, spurred at base. Stamens 10, free, glabrous, exserted. Ovary sessile, villous, few-ovuled ; style short, reflexed ; stigma simple. Le- gume unknown. — FI. Cap. ii. p. 268 ; Thes. Cap. t. 20. B. racemosa , Hochst., the only species, grows near Natal. Leaves simple, ovate-lanceolate, acute, glabrous. Flowers white (?), in axillary ra- cemes. Another species is found in North Africa. Suborder 2. Csesalpiniese. (Gen. 68-75.) 68. PARKINSONIA, Linn. Calyx coloured, with a short urceolate tube, and 5-parted, subequal, deciduous limb. Petals 5, in the throat of the calyx, ovate, flat, the upper one with a long claw. Stamens 10, free, declined ; filaments villous at base. Ovary sessile, many- ovuled; style subulate, ascending; stigma simple. Legume very long, acuminate at each end, compressed between the seeds, 2-valved, many-seeded. — FI. Cap. ii. p. 269. Tropical and subtropical shrubs, armed with spines. Leaves pinnate, multijugate; leaflets small. Flowers racemose, yellow. — P. Africana , Sd., our only species, grows in Namaqualand. 69. GUILANDINA, Juss. Calyx with a short, urceolate tube, and subequally 5-parted limb. Petals 5, in the throat of the calyx, sessile, nearly equal. Stamens 10, free ; filaments villous at base. Ovary stipitate, several-ovuled ; style short ; stigma simple. Legume ovate, ventricose, compressed, 2-valved, 1-2-seeded, covered with straight prickles. — FI. Cap. ii. p. 269. Tropical trees and shrubs, the stem and petioles armed with hooked prickles. Leaves abruptly 2-pinnate. Flowers in spicate racemes ; bracts long. — G. Bonduc , Ait., a native of India and Arabia, occurs on the coast, near Natal. 70. MELANOSTICTA, DC. Calyx deeply 5-parted, segments deciduous, the lowest 90 XLII. LEGUMINOSiE. largest. Petals 5, in the throat of the calyx ; 4 lower ob- ovate-oblong, subequal ; upper one shorter and broader, with indexed edges. Stamens 10, free, ascending, equalling the petals ; anthers short. Ovary sessile, ovate-oblong, 4-ovuled ; style short, straight. Legume compressed, oblong, setose. — j FI. Cap. ii. p. 270 ; Tkes. Cap. t. 2. Small plants, the stem, petioles, leaflets, peduncles, calyx, ovary, and legumes, sprinkled with black, resinous dots. Leaves 2-pinnate, stipulate and stipellate. Flowers racemose. — 2 species, from Transvaal and Zulu- land. 71. PELTOPHOBLJM, Yog. Calyx-tube turbinate ; limb 5-parted, deciduous, the seg- ments oblong, reflexed. Petals 5, obovate, curled at the edge, clawed. Stamens 10, inserted with the petals, free; filaments equalling the petals, hairy at base, indexed in bud ; anthers versatile, slitting longitudinally. Ovary sessile, com- pressed, few-ovuled ; style filiform ; stigma peltate, depressed in the centre. Legume broadly oblong, much compressed, unarmed, acute at each end, 1-2-seeded. Seeds oblong, with a subterminal hilum and straight embryo. — FI. Cap. ii. p. 270. Trees or shrubs. Leaves abruptly pinnate. Flowers racemose, yellow. — P. Africanum , Sd., our only species, is found near Crocodile river and at Magalisberg. 72. BUKKEA, Hook. Calyx 5-parted, the segments equal, imbricate. Petals 5, subequal, spreading. Stamens 10 ; filaments very short, the alternate slightly longer ; anthers oblong, equal, tipped with a deciduous gland. Ovary subsessile, 2-ovuled; style very short ; stigma obliquely peltate, concave, with a wavy margin. Legume piano-compressed, oblique, narrowed at base, stipitate, thinly coriaceous, indehiscent. Seeds ovate -orbicular ; embryo straight. — FI. Cap. ii. p. 271. B. Africana , Hook., the only species, is a shrub or small tree, with ab- ruptly 2-pinnate leaves, and axillary, many-flowered racemes. Grows at Magalisberg. 73. CASSIA, Linn. Calyx 5-parted nearly to the base, more or less unequal. Petals 5, clawed, more or less unequal. Stamens 10, the 3 upper commonly sterile (sometimes wanting) ; fertile anthers opening by 2 terminal pores or short clefts. Ovary sessile or stipitate, multiovulate ; style filiform ; stigma simple. Le- gume terete or compressed, linear, many-seeded. — FI. Cap. ii. p. 271. A vast tropical genus, much diversified. Leaves abruptly pinnate, often XLII. LEGUHINOSiE. 01 having glands on the petioles. Flowers yellow or orange. — About 4 South African species, all Eastern or from Natal. 74. SCHOTIA, Jacq. Calyx-tube obconical ; limb 4-parted, the segments oval, ob- tuse, imbricating, deciduous. Petals 6, in the throat of the calyx, nearly equal. Stamens 10, more or less connate at base ; fila- ments free above, the alternate shorter ; anthers ovate, longi- tudinally slitting. Ovary stipitate, ovate, several-ovuied ; style filiform, elongate ; stigma capitate or simple. Legume coriaceous, oblong, compressed, the upper margin or both margins winged. Seeds 1-6, either with the scar naked or having a large, fleshy, cup-like aril. — FI. Cap. ii. p. 273. South African trees or shrub. Leaves pinnate ; leaflets leathery, entire. Flowers panicled, crimson pink or flesh-coloured. — 3 species, from the Eastern district and Natal. 75. BAUHINIA, Plum. Calyx-tube cylindrical or bell-shaped ; limb 5-parted, deci- duous or persistent, its segments separate or cohering in a reflexed, strap-shaped lobe. Petals 5, clawed, subequal, va- riously inserted. Stamens 10, monadelphous or free, exserted, either all fertile or several (5-7-9) sterile ; filaments filiform ; anthers incumbent, slitting. — FI. Cap. ii. p. 275. Trees or climbing shrubs, chiefly tropical. Leaves formed of 2 partially connate or nearly confluent leaflets, resembling a 2-lobed leaf. Flowers racemose. — At least 4 South African species, all but one Eastern. Suborder 3. Mimosese. (Gen. 76-82.) Tribe 1. Eumimose^i. 76. ENT AD A, Linn. Flowers sessile or shortly pedicelled. Calyx bell-shaped, shortly 5-toothed. Petals 5, free or nearly so. Stamens 10, anthers gland-bearing. Legume linear, piano-compressed, margined with thickened, persistent sutures, the valves trans- versely jointed, separating into 1-seeded, indehiscent fragments. — FI. Cap. ii. p. 276. Shrubs, mostly climbing. Leaves 2-pinnate, the terminal pair often changed to tendrils. Spikes of flowers slender, solitary or panicled. — 3 species, all Eastern. One of them, E. scandens, Benth. (the Sword-bean), grows also in the East Indies. 77. ELEPHANTORHIZA, Benth. Flowers pedicelled. Calyx short, 5-toothed. Petals lan- ceolate, at length free. Stamens 10 ; anthers tipped with a deciduous, stalked gland. Legume straight, compressed, lea- 92 XLII. LEGUMINOSiE. thery, the sutures remaining closed, but the long, persistent, rigid valve separating (as in a siliqua), without transverse septa, and not pulpy within. — FI. Cap. ii. p. 277. Glabrous undershrubs, with large fleshy roots (Elandsboontjes). Leaves 2-pinnate, multijugate. Flowers densely spicato-racemose. — 2 species, both Eastern. 78. DICHROSTACHYS, DC. Flowers of two kinds in the spike ; the uppermost flowers hermaphrodite and sessile, as in Fntada, the lower neuter, with calyx and corolla as in the perfect, 10 long, slender fila- ments, without anthers, and a rudiment of an ovary. Legume linear, twisted, compressed, membranaceo-coriaceous, or some- what fleshy, 1-celled, without pulp, indehiscent or the valves breaking irregularly from the sutures. — FI. Cap. ii. p. 278. African and Asiatic shrubs, sometimes spiny. Leaves 2-pinnate. Spikes of flowers peduncled, nodding, solitary or in pairs. — 2 South African spe- cies, from Natal and to the north-east of that region. 79. XEBOCLADIA, Harv. Flowers capitate, sessile. Calyx 5-parted to the base. Pe- tals 5, free. Stamens 10 ; filaments free, the 5 alternate shorter ; anthers with a very minute, sessile gland. Legume sessile, semiorbicular, piano-compressed, 1 -seeded, indehiscent, the carinal suture arched and wing-bordered. Seed flattened ; embryo straight. — FI. Cap. ii .p. 278. X. Zeyheri , H., from Namaqualand, is a small, dry, and very rigid bush, with pale bark, spinous stipules ; distant, 2-pinnate, deciduous leaves ; and subsessile heads of flowers. Tribe 2. Acacie^e. 80. ACACIA, Willd. Flowers frequently polygamous. Sepals 3-5, either con- nate in a bell-shaped cup or free. Petals as many, more or less united in a monopetalous corolla, rarely at length free. Stamens numerous (mostly more than 50), free or connate at base, rarely (in male flowers) collected in a central column. Legume various, mostly dry. — FI. Cap. ii. p. 279. Trees or shrubs of warm countries. Leaves 2-pinnate. Stipular or axillary spines often present ; prickles in many. Flowers minute, yellow, in heads or spikes. — About 20 South African species (some undescribed), chiefly Northern, Eastern, and from Natal. 81. ALBIZZIA, Duraz. Flowers mostly bisexual. Calyx bell-shaped or tubular, 5- toothed. Corolla monopetalous, funnel-shaped. Stamens in- XLII. LEGUMINOSiE. 03 definite, often numerous, united at base into a tube. Le- gume flat, dry, membranous or papery, with their margins either dehiscent or indehiscent. — FI. Cap. ii. p. 284. TJ narmed trees or shrubs. Leaves 2-pinnate. Flowers in heads or spikes, with long, white or rosy, rarely purple, bundles of stamens. — 2 South African species : one from Delagoa Bay, the other from Lake Ngami. 82. ZYGIA, P. Br. Calyx tubular, 5-toothed. Corolla between funnel-shaped and tubular, shortly 5-lobed. Stamens very many, connate in a tube much longer than the corolla, spirally twisted in the bud, free at the summit only. — FI. Cap. ii. p. 284. Shrubs or trees, with the foliage of Albizzia , from which genus this dif- fers in the long staminal tube. Z. fastigiata, E. Mey., our only species, grows near Natal. Order XLIII. ROSACEA. Calyx free or adnate with the ovary, open or closed, its limb mostly regular, 3-4-5-parted (or of twice as many parts, in two rows). Petals 3-5 or 0, inserted in the throat of the calyx. Stamens inserted with the petals, indefinite or rarely definite ; filaments filiform, free. Ovary of one or more, often many separate carpels (except in Grielum ) ; carpels 1-, 2- or many-ovuled. Styles 1 to each carpel, terminal or lateral. Fruit various. Seeds without albumen. — A large and varied Order, chiefly of the north temperate zone. Leaves alternate, pinnate or digitate, or cleft or parted, rarely simple. Stipules mostly attached to the base of the petiole. Tribe 1. Chrysobalane.®. Calyx tubular or bell-shaped. Carpel soli- tary, 1-2-celled, mostly attached to one side of the calyx-tube ; ovules 1-2, erect. Style lateral or basal. Fruit a drupe. 1. Parinarium. (A dwarf shrub, with oblong, obtuse, simple leaves, white-woolly beneath ; flowers small, corymbose.) Tribe 2. Dryadeas. Calyx open. Carpels many, 1-ovuled, free, crowded on a convex or columnar receptacle. * Shrubs ; calyx h-parted ; fruit juicy. 2. Rubus. (The Bramble or Blackberry.) ** Herbs ; calyx 10-cleft in tivo rows ; fruit dry. 3. Potentilla. Receptacle conical. Carpels without tails. 4. Geum. Receptacle columnar. Carpels with long, twisted tails. Tribe 3. Sangtjisorbe^:. Calyx-tube turbinate or pitcher-shaped, con- tracted in the throat. Carpels 1-4, 1-ovuled, free, concealed within the persistent calyx-tube. * Flowers with petals , yellow. 5. Leucosidea. Calyx-tube unarmed, limb 10-parted, in two rows. 6. Agrimonia. Calyx-tube armed with hooked bristles ; limb 5-parted. XLTIT. ROSACE®. 94 ** Floivers without petals ; bisexual. 7. Accena. Calyx-tube armed with hooked prickles ; limb 4-5-lobed. 8. Alchemilla. Calyx-tube unarmed ; limb 8-lobed, in two rows. *** Flowers without petals , unisexual or polygamous. 9. Poterium. Herbaceous. Flowers in dense terminal spikes ; calyx 4-fid. 10. Cliffortia. Small shrubs or half-shrubs. Flowers axillary, sessile, 3-fid. Tribe 4?. Nettrade®. Flowers bisexual. Calyx-tube concrete with the ovary. Petals 5, convolute. Stamens 10. Carpels 5-10, united in a 5-10-. celled capsule. 11. Grielum. Herbs with many-cleft, lioary leaves, and large, yellow flowers. Tribe 1. Chrysobalane®. (G-en. 1.) 1. PARINARIUM, Juss. Mowers bisexual. Calyx-tube long or short, subequal or unequal-sided ; limb 5-parted, subequal, imbricate. Petals 5, rarely 4, sessile or clawed, deciduous. Stamens 10 or many, shortly connate at base or united in a lateral parcel, all per- fect or some barren. Ovary adnate to the side of the calyx- tube, exserted, 2-celled (or partly so) ; ovules solitary, erect ; style basal, filiform, hairy. Drupe ovoid or globose, with fibrous or pulpy flesh, and a bony, 1-seeded nut. — FI. Cap. ii. p. 596. P. Capense , our only species, is a very dwarf bush, the young parts clothed with foxy hairs. Leaves 2-4 inches long, oblong, obtuse, glabrous above, white-woolly beneath. Peduncles not much branched. Grows on the Aapjes river. Tribe 2. Dryade®. (Gen. 2-4.) 2. RUBUS, Linn. Calyx-tube open, short ; limb 5-parted, imbricate. Petals 5, crumpled, deciduous. Stamens indefinite, perigynous. Car- pels indefinite, on a convex-conical receptacle, 1-ovuled ; styles subterminal, filiform. Pruit of many little drupes, heaped to- gether on the dry, hardened receptacle. — FI. Cap. ii. p. 286. The Raspberry and Bramble. Leaves pinnate or digitate. Flowers mostly panicled. — 5 Cape species, dispersed. 3. POTENTILLA, Linn. Calyx-tube short, concave, open ; limb 8-10-parted in two rows, the inner segments valvate in bud. Petals 4-5, deci- duous. Stamens indefinite. Carpels indefinite, on a convex receptacle, 1-ovuled ; styles lateral, stigmas simple. Pruit of dry achenes, sessile, on a dry, hairy receptacle, not tailed. — FI. Cap. ii. p. 228. XLIII. ROSACEiE. 05 A large genus in the northern hemisphere. — P. supina, Linn., a common European species, occurs by the Orange river, near its mouth. Leaves pinnate-parted, the upper 3-par ted ; leaflets sharply and deeply toothed. Flowers yellow. 4. G-EUM, Linn. Calyx-tube short, open ; limb 10-parted in two rows, the inner segment valvate in bud. Petals 5, deciduous. Stamens indefinite. Carpels indefinite, on a columnar receptacle, 1- ovuled; styles terminal, inflexed or sharply bent; stigmas simple. Achenes on a long receptacle, tailed with the har- dened, awn-like, hooked or curved styles. — FI. Cap. ii. p. 289 ; Thes. Cap. t. 18. A considerable and chiefly northern genus. — G. Capense, our only species, grows in the Eastern district and Caffraria. Its leaves are chiefly radical and lyrate-pinnatisect, the terminal lobe very large. Flowers laxly pa- nicled or solitary, large, handsome, yellow. Tribe 3. Saxguisorbeje. (Gen. 5-10.) 5. LEUCOSIDEA, E. and Z. Elowers bisexual, complete. Calyx-tube obconic, con- stricted in the throat, with an annular perigynous disk ; limb 10-parted, in 2 rows, persistent, outer lobes short, ovate, inner lanceolate, acuminate, valvate in bud. Petals 5, ob ovate, de- ciduous. Stamens 10-12, inserted on the disk. Carpels 2-3, enclosed in the calyx-tube ; styles exserted, filiform, terminal ; stigmas hook-pointed. Utricles membranous, enclosed in the hardened calyx-tube. — FI. Cap. ii. p. 289. P. sericea is a densely leafy shrub, the “Dwa-Dwa” of the natives, who use it as an astringent medicine. Leaves pinnate-parted, with 2-3 pairs of pinnae ; pinnae ovate-oblong, sharply toothed, dark green above, white and silky beneath. Flowers racemose, greenish -yellow. — Eastern frontier and Caffraria. 6. AGRIMONIA, Linn. Calyx naked at base, the tube turbinate, armed with many hooked bristles constricted at the throat, with an annular disk ; limb 5-parted, the lobes imbricate, at length closing in. Petals 5, deciduous. Stamens 12-20. Carpels 2, 1-ovuled, enclosed in the calyx-tube ; styles terminal, exserted. Achenes 1-2, enclosed in the hardened and densely hook-bristled calyx- tube. — FI. Cap. ii. p. 290. A. Pupatoria , Linn., a European, Asiatic and North American species, occurs in Caffraria and on the east frontier. Leaves interruptedly pinnate ; leaflets coarsely-toothed. Flowers in terminal, spiked racemes, yellow. 7. ACiENA, Yahl. Elowers bisexual. Calyx-tube oblong, constricted in the 90 XLITT. ROSACEA!. throat, bristly or smooth, compressed, 3-4-5-angled, the angles armed with hooked bristles ; limb 4-, rarely 3-5 -parted, per- sistent. Petals 0. Stamens 2-5. Carpels 1-2, enclosed in the calyx-tube, 1-ovulate ; styles terminal, short ; stigma pen- cilled. Achenes hidden in the hardened, hook-bristled calyx- tube. — FI. Cap. ii. p. 290. Herbs or underehrubs of temperate climates. Leaves pinnate-parted, the segments toothed or cut. Flowers small, green, in spikes or globose heads. — 2 Cape species, both Western. 8. ALCHEMILLA, Tournef. Plowers bisexual. Calyx-tube urceolate, constricted in the throat, with an annular disk, unarmed ; limb 8-parted, in 2 rows, the outer lobes small, with imbricate aestivation. Pe- tals 0. Stamens 1-4, exserted. Carpels 1-4, in the base of the calyx-tube, substipitate, 1-ovuled ; styles basal, filiform • stigmas capitellate. Achenes 1-2, in the calyx-tube. — FI. Cap. ii. p. 291. Small, hairy herbs of the temperate zones. Leaves alternate, fan-shaped or reniform, lobed or deeply parted or crenate. Flowers minute, green, in corymbs or tufts. — 2 Cape species, dispersed. 9. POTERJUM, Linn. Plowers spiked, polygamous or unisexual, the females in the upper part of the spike. Calyx-tube turbinate, unarmed, con- stricted in the throat with an annular disk ; limb 4-parted, imbricate. Petals 0. Stamens 20-30. Carpels 2-3, en- closed in the calyx-tube, 1-ovuled ; styles terminal, exserted ; stigma pencilled. Achenes concealed in the hardened or fleshy, 4-angled calyx-tube. — FI. Cap. ii. p. 292. P. Sanguisorba , Linn., introduced from Europe, grows near Simon’s Town. Leaves chiefly radical, of many cut leaflets. Stems branched, laxly leafy ; spikes globose, terminal, very dense. 10. CLIFFOKTIA, Linn. Plowers dioecious. Calyx-tube urceolate, unarmed; limb 3-parted (rarely 4-parted). Petals 0. — Male: Stamens in- definite, 8-20-30-40 ; filaments very slender, much exserted. - — Pemale : Carpels 2, enclosed in the calyx- tube, 1-ovuled ; style lateral ; stigmas long, feathery. Achenes 1-2, mem- brane-skinned, enclosed in the hardened and variously-sculp- tured, rarely fleshy, calyx-tube. — FI. Cap. ii. p. 292 ; Thes. Cap. t. 95. A South African genus, of about 40 small, leafy shrubs or undershrubs. Leaves properly digitately 3-foliolate, often seemingly simple or l-foliolat,e, either from the confluence of the 3 leaflets into one, or from the lateral XLTTT. ROSACEiE. 97 being minute or abortive ; rarely 2-foliolate, the medial leaflets disappearing. Stipules attached to the petiole. Flowers axillary, small and green, sub- sessile — C. strobolifera, Linn., is very common from Capetown to Natal ; the great majority of the other species scarcely extend further east than Swellendam. Tribe 4. Neurade.®. (G-en. 11.) 11. GRIELCJM, Linn. Calyx-tube short, at length concrete with the ovary ; limb 5-lobed ; lobes nearly valvate in the bud. Petals 5, inserted in the throat of the calyx, large, obovate, convolute in bud. Stamens 10, inserted with the petals. Carpels 5-10, in the base of the calyx, confluent with the calyx- tube and with each other, 1-ovuled ; styles 5-10, filiform, short ; stigmas capitate. Capsules depressed, 5-10-celled, the cells at length opening in the axis, 1-seeded. — FI. Cap. ii. p. 304. South African herbs, growing in sandy places and in salt ground. Leaves alternate, hoary, pinnately decompound, with narrow segments. Flowers large, yellow. Order XLIY. SAXIFRAGES. Calyx 5-cleft (rarely 3-10-cleft), regular, adnate to the ovary or free ; limb mostly persistent, sometimes enlarged in fruit. Petals as many as the calyx-lobes, Rarely wanting, sometimes cleft or lacerate. Stamens inserted with the petals in the throat of the calyx, as many as the petals and alternat- ing, or twice as many, rarely fewer or more numerous. Ovary inferior or more or less superior, of 2 (rarely 3-5) carpels, more or less cohering by their inner faces ; ovules commonly many ; styles distinct, or more or less confluent. Fruit cap- sular, splitting at maturity through the centre. Seeds almost always albuminous. — A large and heterogeneous Order, to which it is nearly impossible to affix natural limits. Tribe 1. Saxifrages. Herbaceous or half-shrubby plants with exsti- pulate leaves. 1. Vahlia. Ovary inferior ; styles 2, spreading. Leaves opposite, simple, linear. Tribe 2. Cunonies. Shrubs or trees, with opposite leaves and interpe- tiolar stipules. 2. Cunonia. Flowers racemose. Petals entire. Leaves pinnate. 3. Platylophus. Flowers panicled. Petals 3-fid. Leaves 3-foliolate. Tribe 3. Escallonies. Trees and shrubs, with alternate, simple, exsti- pulate leaves. Stamens as many as the petals. Ovary inferior. 4. Choristylis. Flowers 5-fid, minute and green, in axillary panicles. Leaves ovate, serrate, strongly-nerved, and veiny. H 98 X LTV. SAXIFRAGES. Tribe 4?. Bkexiea:. Shrubs, with alternate, exstipulate leaves. Sta- mens as many as the petals. Ovary superior, 5-angled, 5-celled. Seeds without albumen. 5. Brexia. Flowers 5-parted, sepals and petals coriaceous, imbricated. Oavry surrounded by a 5-lobed, imbricate disk. Leaves oblong or obovate, entire. Tribe 1. Saxifrages. 1. VAHLIA, Th. Calyx-tube adhering to the ovary ; limb 5-parted, persistent, valvate in bud. Petals 5, spreading, entire, epigynous. Sta- mens 5. Ovary inferior, 1-celled, with 2 many-ovuled pla- centas pendulous from the summit of the cavity ; styles 2, spreading ; stigmas capitate. Capsules membranous, opening between the styles. Seeds minute, very many. — FI. Cap. ii. p. 306. V. Capensis , Th., our only species, is a much-branched, lialf-woody plant, 6 inches to 2 feet long, more or less hairy or smooth ; leaves linear or lanceolate ; peduncles 2-flowered, shorter than the leaves. — Found in the West and North-West districts, and Namaqualand. Tribe 2. Cunoxie^. (Gren. 2-3.) 2. CUNOmA, Linn. Calyx free, 5-parted, deciduous. Petals 5, oblong, entire. Stamens 10. Ovary free, 2-celled, conical, with many-ovuled, sutural placentas ; styles 2, diverging ; stigmas simple. Cap- sules conical, 2-horned, 2-celled, separating from base to apex, from a free, placentiferous column. Seeds many, compressed, with a narrow membranous wing. — FI. Cap. ii. p. 306. C. Capensis , Linn., the only species, is a large shrub or small tree, com- mon throughout the colony, glabrous in all parts. Leaves pinnate ; pinnae in 2-4 pairs, lanceolate, sharply serrate. Stipules broadly ovate, deciduous. Racemes axillary, opposite, very many-flowered ; flowers white. 3. PLATYLOPHUS, Don. Calyx free, 4-(rarely 5-)parted, persisteni, valvate in bud. Petals 4-5, 3-fid. Stamens 8-10, on the outer edge of a fleshy, perigynous disk. Ovary free, 2-celled ; ovules 2 in each cell, collateral, pendulous ; styles 2 ; stigmas simple. Cap- sule turgid at base, compressed above, membranous, 2-celled, at length splitting ; cells 1-seeded. — FI. Cap. ii. p. 307. P. trifoliatus, Don, the only species, is a tree 40-50 feet high, glabrous in all parts, extending at least as far east as Uitenbage, more frequent in the west. Leaves long-petioled, pinnately 3-foliolate ; leaflets lanceolate, mi- nutely toothed. Panicles axillary, much-branched ; flowers small, white, almost always 4-parted. XLTY. SAXIFRAGES. 99 Tribe 8. Escallontes. 4. CHORISTYLIS, Harv. Elowers polygamous. Calyx-tube obconic, adnate with the ovary ; limb 5-cleft, persistent. Petals 5, inserted within the margin of the calyx-tube, longer than the lobes, sessile, entire, persistent, valvate in bud. Stamens 5, alternate with the petals and inserted with them ; filaments short ; anthers ovate, 2- celled, slitting. Ovary 2-celled, many-ovuled, with axile pla- centas ; styles 2, short, at first connate, then widely diverging ; stigma capitate. Capsule more than half-inferior, its co- nical, acuminate apex girt by the persistent calyx-limb and petals, opening through the styles. — FI. Cap. ii. p. 308 ; Thes. Cap. t. 123. C. rhamnoides, Harv., the only species, is a leafy slirub, extending from the eastern frontier to Natal. Leaves alternate, ovate, serrate, ribbed and nerved. Flowers small and green, in axillary panicles. Tribe 4 P. Brexies. 5. BREXIA, Thouars. Calyx free, 5-cleft, persistent, with short, acute, leathery segments, imbricate in bud. Petals 5, inserted outside the margin of a perigynous ring, leathery, oblong, obtuse, imbri- cate in bud. Stamens 5, alternate with the petals and in- serted with them; filaments subulate ; anthers oblong, erect, basifixed. Annular disk thick, attached to the base of the ovary, with 5 fimbriate lobes. Ovary superior, 5-angled, 5-celled ; ovules many, on axile placentas ; style short ; stigma 5-lobed. Seeds without albumen ; embryo almond-like. — FI. Cap. ii. p. 597. B. Madagascariensis, Lindl., was collected at Delagoa Bay by Forbes. A glabrous and subglaucous shrub, with oblong or obovate entire leaves, with revolute margins. Flowers green, in imperfect umbels. Order XLV. CRASSULACEJE. Calyx free, 4-7-cleft or parted, imbricate. Petals inserted in the bottom of the calyx, as many as its lobes, regular, free or connate in an imperfectly tubular corolla, imbricate. Sta- mens inserted with the petals, as many or twice as many. Ovary (in the Cape genera) of 4-7 carpels, nearly apocarpous ; styles terminal, subulate, free. Fruit of 4-7 follicles. Seeds albuminous. — Succulent plants, with fleshy, entire, rarely crenate or pinnate leaves. Flowers in Cymes or solitary, showy or minute. ii 2 100 XLY. CRASSULACEvE. Tribe 1. Isostemones. Stamens as many as the petals. Sepals and petals four. Ovules 1 in each carpel 1. Helophytttm. Ovules several in each carpel 2. Bulliarda. Sepals and petals 5 (or rarely 6-9) . Carpels, each with a horn-like crest at the back of the styles 3. Dinacria. Carpels not crested, tapering upwards. Corolla subgamopetalous, salver-shaped. Calyx bell-shaped, 5-toothed. Tube of corolla not longer than the calyx 4. Grammanthes. Calyx deeply 5-cleft. Tube or corolla longer than the calyx. Anthers subsessile, in throat 6. Rochea. Corolla of 5-9 separate or slightly connate petals, spreading 5. Crassula. Tribe 2. Diplostemones. Stamens twice as many as the petals. Calyx 5-parted. Corolla tubular, 5-lobed .... 7. Cotyledon. Calyx 4-parted, sessile, narrow. Corolla 4-lobed . . 8. Ealanchoe. Calyx inflated, shortly 4-lobed. Corolla 4-lobed . . 9. Bryophyllum. 1. HELOPHYTUM, E. and Z. Calyx 4-fid or 4-toothed. Petals 4, roundish or obovate, spreading. Stamens 4, shorter than the petals. Scales cu- neate, truncate. Carpels 4 ; ovules solitary ; styles short. — FI. Cap.ii.p. 328. Small marsh or water-plants, with weak, erect or floating, simple or slightly- branched stems. Leaves opposite, linear, spatliulate or obovate. Flowers axillary, solitary or in cymes, white. — 2 species, 1 of which is Eastern, the other diffused. 2. BULLIARDA, DC. Character as Helophytum, except, carpels several- ovuled ; follicles several-seeded. — FI. Cap. ii. p. 329. Small, mostly annual, 2-3-chotomous plants, growing in moist ground. Flowers small, white, terminal, and axillary. — 5 Cape species, dispersed. 3. DHSTACRIA, Harv. Calyx deeply 5-cleft. Petals 5, slightly connate at base, with broad, erect claws and spreading or recurved limbs. Stamens 5, shorter than the petals. Scales narrow-cuneate, truncate. Carpels 5, several-ovuled, each with a short dorsal horn at the summit behind the style ; styles short, subulate. Eollicles several- seeded. — FI. Cap. ii. p. 330. H. Jiliformis, Harv., the only species, is a small, 3-chotomous annual, na- tive of the Western district. Leaves obovate or oblong, blunt, fleshy. Corymbs dense, terminal ; flowers small, white. XLV. (JltASSULACJiLE. 101 4. GRAMMANTHES, DC. Calyx bell-shaped, semi-5-fid. Corolla gamopetalous, the tube as long as the calyx ; limb 5-6-lobed, spreading. Sta- mens 5-6, attached to the corolla-tube, shortly exserted. Carpels 5-6, many-ovuled, with subulate styles. Scales very minute or obsolete. Follicles many-seeded. — FI. Cap. ii. p. 331. A small, 2-chotomous, glabrous and glaucous annual, very variable in size of all parts ; found in sandy soil throughout the Western districts. Stems rigid, wiry ; leaves in distant pairs, fleshy, oblong ovate or sublinear. Flowers panicled, few or many, or solitary, terminal !, orange yellow or creamy-white, each petal commonly (not always) having a darker mark shaped like the letter V. 5. CRASSULA, Linn. Calyx 5-parted or deeply 5-cleft, rarely 6-9-parted, spread- ing or erect. Petals 5 (rarely 6-9), free or connate below, spreading or erect, or erect with recurved points, ovate ob- ovate oblong or panduriform or lanceolate, either simple at the apex or mucronulate or gland-tipped. Stamens 5 (rarely 6-9), shorter than the petals. Scales various. Follicles several-seeded. — FI. Cap. ii. p. 332. A large genus of succulent shrubs undershrubs or herbs, very variable in habit and size. Leaves opposite, mostly connate at base, broad or narrow, flat or round, more or less fleshy, glabrous pubescent or scaly, often carti- lagineo-ciliate. Flowers white, red, rarely yellow, mostly of small size, solitary cymose or capitate. — About 100 species (probably more), dispersed, chiefly in dry ground. 6. ROCHEA, DC. Calyx 5-parted or deeply 5-cleft. Corolla more or less per- fectly gamopetalous, salver-shaped, its tube longer than the calyx ; limb 5-parted, spreading. Stamens 5, adnate to the claws of the petals ; anthers subsessile, in the throat of the tube. Scales very small. Carpels 5, several-ovuled ; styles subulate or clavate. Follicles many-seeded. — FI. Cap. ii. p. 368. Shrubby or half-shrubby succulents. Leaves connate or sheathing at base, fringed with small cilia. Flowers handsome, crimson rosy white or pale yellow ; some very sweetly scented. — 4 species, all Western. 7. COTYLEDON, Linn. Calyx 5-parted, much shorter than the tube of corolla. Corolla gamopetalous, with an ovate or linear, 5-angled tube, and a spreading or reflexed and revolute, 5-parted limb, spirally twisted in bud. Stamens 10, attached to the base of the corolla-tube, exserted or subincluded. Scales oval. Car- pels 5, many-ovuled ; styles subulate. — FI. Cap. ii. p. 370. 102 XLY. CRASSTJ LACEiE. Shrubby half-slirubby or herbaceous succulents. Leaves entire, oppo- site or scattered. Flowers showy, panicled or racemose, peduncled. — 23 (perhaps more) Cape species, dispersed. 8. KALANCHOE, Adans. Calyx 4-parted, sepals small, acute. Corolla monopetalous, salver-shaped, with an urceolate tube and a 4-parted, spread- ing limb. Stamens 8, attached to the base of the corolla. Scales 4, linear or oblong. Carpels 4, many-ovnled, with su- bulate styles. Follicles many-seeded. — FI. Cap. ii. p. 378. Succulent undershrubs, with opposite, toothed, entire or pinnatifid, fleshy leaves. Flowers panicled, yellow red or cream- colour. — 6 Cape species (perhaps more), dispersed, chiefly Eastern. 9. BRYOPHYLLUM, Salisb. Calyx inflated, 4-cleft nearly to the middle, valvate in bud. Other characters as in Kalanchoe. — FI. Cap. ii. p. 380. Succulent undershrubs. Leaves opposite, fleshy, petioled, imparipinnate or 1-foliolate ; pinnules crenate. Flowers panicled, yellow changing to red. B. tubiflorum , H., a little-known species, grows at Delagoa Bay. Others, probably, may be discovered at Natal or in Zululand. Order XLYI. HAMAMELIDEiE. Trees or shrubs, with simple, petioled, penninerved, entire or toothed leaves. Stipules minute, deciduous. Flowers small. Floral characters nearly as in Saccifragacece ; but the anthers are erect, 2-celled, each cell opening by a lateral, in- trorse valve, or splitting at the sides. Seeds solitary, albumi- nous. 1. Trichocladus. Flowers unisexual, spiked. Petals 5, long, linear. Stamens 5. 2. Grubbia. Flowers bisexual, capitate. Petals 4, ovate, hairy. Sta- mens 8. 1. TRICHOCLADUS, Pers. Flowers polygamous, monoecious or dioecious ; female flowers without petals. Calyx 5-cleft, adnate to the base of the ovary, persistent, valvate in bud. Petals 5, linear-clavate, much longer than the calyx, with revolute margins, valvate in bud. Stamens 5, alternate with the petals ; filaments short, thick- ened in the middle ; anthers erect, adnate, dehiscing by valves. Styles 2, spreading. Capsules didymous, 2-celled, endocarp separating from the sarcocarp. Seeds solitary, pendulous. — FI. Cap. ii. y. 324. South African shrubs, with opposite or alternate, ovate or oblong leaves, XLYI. KAMAMELIDEiE. 103 and densely hairy or pubescent twigs and branches. Flowers white, in terminal spikes.— 2 species, both Eastern and South-Eastern. 2. GRUBBIA, Berg. Blowers perfect, capitate, in a 2-leaved involucre. Calyx adnate, its limb abortive, truncate. Petals 4, epigynous, de- ciduous, ovate, hairy outside, valvate in bud. Stamens 8, the 4 alternating with the petals rather longer, all slightly attached to the bases of the petals ; anthers 2-celled, minute, roundish, opening by valves. Ovary inferior, crowned with a fleshy disk, when young ( fide Dene.) 2-celled, with a pendulous ovule in each cell ; afterwards 1-celled (the septum breaking up), with an ovule pendulous fronra central, columnar placenta. Nuts laterally connate, 1-seeded, crowned by the disk and style. — FI. Cap. ii. p. 325. Small, much-branched South African shrubs. Leaves opposite, with re- volute margins, exstipulate. Flowers 3 or many, soldered together iu bracteated heads. — 4 species, 1 only going so far east as Uitenhage. Order XLYII. BBUNIACEJE. Blowers bisexual, regular, small or minute, sessile, spiked or capitate, rarely solitary, axillary. Bloral characters nearly as in Saxifragacece , but the ovules are definite, either solitary or in pairs, very rarely 10. — Small South African shrubs, with needle-shaped or heath-like, rarely ovate, crowded, small, very entire, sessile or subsessile leaves, with a discoloured or withered tip ( ustulate ). Stipules 0. Fruit 1-seeded, mostly indehiscent. Ovary 1-celled. Style 1 1. Berzelia. Ovary 2-celled. Style 1. Flowers axillary 2. Tittmannia. Style 1. Flowers solitary, terminal 3. Thamnea. Styles 2. Flowers capitate or panicled .... 4. Brunia. Fruit dicoccous, or a splitting capsule. Ovary 2-celled, 2-ovuled. Flowers capitate. Styles 2. Petals free, or slightly cohering at base 5. Berardia. Style 1. Petals free 6. Staavia. Ovary 2-celled, 4-ovuled. Flowers in leafy spikes. Flowers monopetalous, 5-lobed 7. Lonchostoma. Flowers 5-petaled. Anthers apiculate .... 8. Linconia. Ovary 3-celled, 6-ovuled. Style trigonous. Flowers in spike-like, terminal heads, red 9. Audouinia. 1. BERZELIA, Brongn. Calyx adnate ; lobes 5, rarely 4, unequal, gibbous. Petals 5, rarely 4, free. Stamens 5-4, longer than the petals. Ovary 104 XLVII. B11UNIACEJ2. half-inferior, oblique, 1-celled, 1-ovuled ; style simple, with a subeonical stigma. Fruit indehiscent, gibbous. — FI. Cap. ii. p. 310. Heath-like shrubs, with short, 3-angled, imbricate or spreading leaves. Flowers in globose, terminal heads, white. — 8 species, dispersed.* 2. TITTMANNIA, Brongn. Calyx-tube spherical, wrinkled and glandular outside, ad- nate ; limb 5-cleft, with scarious, erect segments. Petals with the claws 2-keeled inside ; limb roundish-ovate, spreading. Ovary inferior, spherical, 2-celled, with a membranous septum, free at the edges ; cells 2-ovuled ; style simple, conical ; stigma 2-dentate. — FI. Cap. ii. p. 312. A small shrub, with subumbellate branches. Leaves linear, subcylindrical, wrinkled, incurved, erect, imbricate. Flowers solitary in the axils of the upper leaves. — Native of Tulbagh and Stellenbosch districts. 3. THAMNEA, Brongn. Calyx adhering to the ovary at base, free above, with 5 lan- ceolate, smooth, scarious, imbricate segments. Petals with 2- keeled claws, and an ovate, spreading limb. Stamens included. Ovary inferior, crowned by a fleshy disk, imperfectly 2-celled, 4- or 8-ovuled; style simple ; stigma entire. — FI. Cap. ii. p. 324. T. uniflora , Soland., the only species, f is a small shrub, with filiform, erect, fastigiate branches. Leaves very small, rhomboid, bluntly keeled, close- pressed, spirally inserted, the upper ones a little longer than the rest, form- ing an involucre to the terminal white flower. — A specimen of this, found by Masson, exists in the British Museum ; no recent collector has found it. 4. BBUNIA, Linn. Calyx half-adnate. Petals ovate or spathulate. Ovary half- inferior, 2-celled ; ovules 1-2 in each cell ; styles 2, diverging. Fruit indehiscent, rarely septicidal-dehiscent, 1 -seeded. — FI. Cap. ii. p. 313. Small shrubs, erect or diffuse ; in some the leaves are small, linear or subulate, and the flowers in heads ; in others (Sect. Beckea) the leaves are larger, often expanded, ovate cordate or lanceolate, and the flowers in panicles. — 10 species, chiefly Western, 2 or 3 going east to Uitenhage. 5. BERARDIA, Brongn., ex parte. Calyx adhering to the ovary at base, free at apex, 5 -cleft. Petals 5, free or cohering at base. Stamens 5 ; anthers with- * A species, discovered by Burchell, and not included in FI. Cap., is figured in Hook. Ic. PI. t. 1014. — J. T). H. f Two additional species, discovered by Burchell, are figured in Hook. Ic. PI. t. 1011-13. All are Western. — J. D. H. XLVIJ. BRUNIACEJ5. 105 out any apical appendage. Ovary 2-celled ; cells 1-ovuled ; styles 2. Fruit dicoccous. — FI. Cap. ii. p. 318. Small, fastigiate slirubs. Leaves small, subulate or rhomboid, keeled, close-pressed, covering the stem on all sides. Flowers in terminal heads. — 11 species, all west of Swellendam. 6. STAAVIA, Th. Calyx adnate to the base of the ovary, free above, with 5 subulate lobes. Petals free. Ovary half-inferior, 2-celled ; cells 1-ovuled ; style simple ; stigma 2-lobed. Fruit dicoccous. — FI. Cap. ii. p. 321. Small shrubs, with linear, spreading leaves. Flowers in terminal, flat- topped heads, involucrated by numerous, whitish floral leaves. — 6 species, all from the Western districts. 7. LONCHOSTOMA, Wikstr. Calyx attached to the half-inferior ovary, 5-cleft. Corolla monopetalous, tubular, 5-cleft ; segments clawed, spreading. Stamens 5, very short, inserted in the mouth of the corolla ; anthers oblong, 2-celled, cells divergent at base. Ovary pu- bescent, 2-celled ; cells 2-ovuled ; styles 2 or 1. Fruit splitting from the base, 2-4-valved ; seeds 4, netted. — FI. Cap. ii. p. 316. Erect shrubs. Leaves imbricate, ovate or lanceolate, leathery. Flowers in terminal, leafy spikes, rosy. — 3 species, all Western. 8. LINCONIA, Linn. Calyx adhering to the ovary, with a 5-cleft limb ; lobes short, membranous, smooth. Petals oblong, convolute, enclos- ing the stamens ; anther-cells divergent at base, their connec- tive with a conical gland at tip. Ovary half-inferior, 2-celled ; cells 2-ovuled ; styles 2. Fruit dicoccous. — FI. Cap. ii. p. 317. Erect, fastigiate shrubs. Leaves spreading or loosely imbricate, linear oblong or ovate-oblong, leathery. Flowers in terminal, leafy spikes, white. — 3 species, dispersed. 9. ATJDOUINIA, Brongn. Calyx adhering to the ovary, 5-cleft ; lobes large, imbricate. Petals with a long, 2-keeled claw, and a spreading, roundish limb. Stamens included. Ovary half-inferior, 3-celled ; cells 2-ovuled ; style 3-angular ; stigmas 3, short. — FI. Cap. ii. p. 323. A. capitata, Brongn., the only species, grows on the mountains round Simonstown, etc. Leaves spirally inserted, linear, imbricate, 3-angled, scabrous. Flowers crimson, in oblong spikes or heads. 106 Order XLVIII. HALORAGEiE. Flowers minute, uni- or bisexual. Calyx-tube adnate ; limb 2-3-4-tootbed or entire. Petals 2-3-4 (or 0), epigynous, with valvate or imbricate aestivation, deciduous. Stamens as many or twice as many as the petals, and inserted with them, ' rarely fewer. Ovary inferior, 1-3-4- celled ; ovules solitary or in fours, pendulous ; style, if any, separate, very short ; stig- mas villous or feathery. Pruit nut-like or fleshy, 1-4-celled. Seeds pendulous ; embryo in the axis of fleshy albumen. Leaves on long petioles, all radical. Peduncles scape-like, bearing a compound spike of minute flowers 1. Gunnera. Leaves sessile, entire or toothed. Flowers axillary. — A creeping or prostrate herb 2. Serpicttla. Leaves whorled, pectinate-parted. Flowers in ter- minal spikes. — Water plants 3. Myriophyllum. 1. GUNNER A, Linn. Flowers unisexual or bisexual. Calyx-tube terete or an- gular ; limb 2-3-lobed. Petals 2 or 0, coriaceous, deciduous. Stamens 1-2, opposite the petals ; anthers 2-celled. Ovary 1-celled, 1-ovuled ; stigmas 2, long, simple, papillose. Fruit succulent, with a bony endocarp. Albumen copious ; embryo very minute. — FI. Cap. ii. p. 571. Nearly stemless herbs, with long-petioled, reniform, many-nerved, sub- radical leaves. Scapes tall, ending in a compound spike or thyrsus of many minute flowers ; the female spikelets occupying the lower half of the spike. — G. perpensa, our only species, grows in wet ditches throughout the colony. 2. SERPICULA, Linn. Flowers unisexual. — Male : Calyx minute, 4-toothed. Pe- tals 4, oblong, concave. Stamens 4 or 8. — Female : Calyx- tube 4-angled, 8-ribbed ; limb 4-toothed. Petals and stamens 0. Ovary 1-celled ; ovules 4, pendulous ; stigmas 4, long and feathery. Fruit 8-ribbed, fleshy, with a bony endocarp, 1- seeded. — FI. Cap. ii. p. 572. S. repens, Linn., our only species, grows in moist places throughout the colony. Stems decumbent, creeping. Leaves alternate or the lower oppo- site, sessile, lanceolate or oval, entire or toothed, ‘often reddish. Stamens 4. 3. MYRIOPHYLLUM, Linn. Flowers unisexual, rarely bisexual. — Male : Calyx 4-parted. Petals 4, ovate, caducous. Stamens 4-6-8. — Female : Calyx- limb 4-parted. Petals 0 or very minute. Stamens abortive or 0. Ovary 4-celled ; stigmas 4, long, compressed, papulose XLYIII. HALORAGEJE. 107 on the inner face. Fruit 4-lobed, of 4 nut-like carpels.— FI. Cap. ii. p. 572. Water plants, with opposite or whorled leaves, the submerged ones pec- tinate-parted. Flowers in terminal leafy spikes, or axillary. — M. spicatum, Linn., a widely-distributed species, occurs in several South African rivers. Order XLIX. BALANOPHOREiE. Blowers unisexual, in dense heads spikes or panicles. — Male : Perianth 3- parted, valvate in bud, or 0. Stamens 1-3, opposite the segments of the perianth. — Female : Perianth epigynous 3-lobed or obsolete. Ovary inferior, 1-celled ; ovule solitary, pendulous ; styles filiform, 1-2 or*0. Fruit dry or suc- culent, indehiscent, the seed filling the cavity. Embryo very minute, in fleshy or friable albumen. — Fleshy, fungous-like root-parasites of hot countries. Leaves represented by fleshy, coloured scales. Flowers either bracteate, involucred, or naked. Flowers monoecious, in dense bracteate spikes . . 1. Mystropetalon. Flowers dioecious, the males panicled ; females con- crete, in globose heads 2. Sarcophyte. 1. MYSTROPETALON, Harv. Flowers monoecious, in dense bracteate spikes. — Male : Pe- rianth 3-parted, 2-labiate, the segments with long claws and spathulate limbs, valvate in bud, the 2 posterior connate. Stamens 2, opposite to and inserted on the posterior segments, conniving ; anthers 2-celled, extrorse, opening longitudinally ; pollen cubical. — Female : Perianth epigynous, tubular, 3- toothed. Ovary seated in a cup-like, fleshy disk or recep- tacle, 1-ovuled ; style filiform, exserted, deciduous ; stigma capitate, 3-lobed. Fruit subtended by the unchanged recep- tacle, with a thin, juicy epicarp, and a crustaceous endocarp, 1-seeded. Embryo very minute, in the base of easily-friable albumen. — FI. Cap. ii. p. 573. Stem simple, fleshy, densely imbricated with linear-spat hulate scales, ending in a dense spike of flowers. Female flowers in the lower, males in the upper part of the spike. Bracts 3 under each flower, 1 anterior, 2 lateral. Bracts orange. Flowers bright red. — 2 species : M. Polemanni , from How- hoek Pass ; M. Thomii, from Caledon Baths. 2. SARCOPHYTE, Sparm. Flowers dioecious. — Male flower panicled ; perianth 3-lobed, valvate in bud. Stamens 3, opposite the lobes ; filaments free ; anthers multilocular. — Female flowers in globose heads, densely crowded. Perianth 0. Ovary seated on a subglobose common 108 XLIX. JJA.LAXOP1LOREA. receptacle, becoming concrete, unilocular ; stigma sessile, pel- tate. Syncarpium (compound fruit) globose, berry-like. S. sanguined, Sparm., the only species, grows on the roots of Ekebergia Capensis and Acacia Caffra , in Uitenhage and Albany. Stems 9-10 inches high, an inch or more in diameter, dull flesh-coloured or reddish, branching, imbricated with scales below, ending in a panicled inflorescence. Smell offensive, like that of rotten fish. Male flowers purplish. Fruit like a mulberry. Order L. RHIZQPHOREiE. Flowers bisexual, regular. Calyx-tube adnate to the ovary or nearly free ; limb of 4-12 valvate lobes. Petals as many as the calyx-lobes, alternate with them, inserted below them, often cut or jagged. Stamens as many, twice as many or more ; filaments separate ; anthers erect. Ovary more or less inferior, rarely quite superior, 2- or more celled; ovules in pairs, or few in each cell, pendulous ; style simple. Fruit 1- or few-seeded. Seeds with or without albumen. — Trees or shrubs, with opposite leaves and axillary flowers, either mari- time or submaritime. Calyx-tube adnate to the ovary. Petals entire emargi- nate or 2-fid. Seed germinating within the fleshy, indehiscent fruit, exalbuminous. Calyx 4-fid. Petals 4 1. Rhizophora. Calyx 5-12-fid. Petals 5-12, 2-fid 2. Beugttikea. Calyx-tube free. Petals 5, multifid. Capsule 3-celled . 3. Weihea. 1. RHIZOPHORA, Linn. Calyx- tube obovate, adhering to the ovary ; limb of 4 oblong, persistent segments. Petals 4, oblong, coriaceous, entire or emarginate, conduplicate, with a double row of long, woolly, marginal hairs. Stamens 8 ; anthers nearly sessile, linear-ob- long. Ovary 2-celled ; ovules 2 in each cell ; style conical, short, 2-furrowed ; stigma 2-toothed. Fruit ovate or oblong, girt near the base with the persistent calyx-lobes, at length perforated at the apex by the germinating embryo. Seed ex- albuminous. — FI. Cap. ii. p. 518. Trees, growing on muddy seashores in hot countries, with quite entire, leathery leaves, — R. mucronata, Lam. (Mangrove), occurs near Natal. 2. BRUGUIERA, Lam. Calyx-tube turbinate, adhering to the ovary ; limb of 5-12 (or 13) persistent segments. Petals as many, oblong, 2-fid, coriaceous, conduplicate, each embracing 2 stamens, woolly on the margin. Stamens twice as many as the petals, inserted in pairs opposite to them ; filaments unequal, half as long as the l. RmzopnoRE.E. 109 petals ; anthers linear or sagittate. Ovary 2-4-celled ; ovules 2 in each cell ; style nearly as long as the stamens ; stigma 2-4-toothed. Fruit crowned by the calyx-lobes, at length perforated by the germinating embryo. Seed exalbuminous. — FI. Cap. ii. p. 514. Trees and shrubs of muddy seashores, in warm countries. — B. gymno- rltiza , Lam., grows near Natal. 3. WEIHEA, Spr. Calyx free, deeply 4-5-parted, the lobes valval e. Petals 4-5, inserted within the calyx-tube, wedge-shaped, palmately multifid-lacerate. Stamens 20-30, inserted below the petals, 1-seriate ; filaments subulate ; anthers oblong, erect, 2-celled. Ovary free, sessile, globose, 3-celled ; ovules 2 in each cell, pendulous ; style simple, persistent ; stigma 3-toothed. Berry dry, girt by the persistent calyx, 3-4-celled, at length dehis- cent. Seeds solitary in each cell, albuminous. — Spr. Syst. ii. 594. Bichaeia, Thouars. Anstrutheria, Gard. in Calc. Journ. iv. p. 344. t. 4 ; Walp. Ann. ii. p. 173. African and Asiatic trees or shrubs, not strictly littoral, but growing near the sea. Leaves opposite, with interpetiolar, deciduous stipules, and axil- lary flowers. Mr. Gerrard has recently sent from Natal a species of this genus, nearly allied to W. Madagascariensis , if it be not the same ; its leaves are ovate-lanceolate, acuminate, denticulate, and flowers axillary, shortly pedicelled, the pedicels 2-bracted at base. Calyx-lobes silky outside ; stamens about 20. Order LI. COMBRETACEjE. Flowers either unisexual or bisexual. Calyx-tube adnate to the ovary, limb 4-5-lobed. Petals inserted at the summit of the calyx-tube. Stamens as many or twice as many as the petals. Ovary inferior, 1-celled ; ovules few, pendulous ; style filiform. Fruit a winged or ribbed dry drupe. — Trees or shrubs, with simple, entire, exstipulate leaves. Flowers in spikes racemes or heads. Flowers without petals 1 . Termin alia. Flowers with 4-5 petals. Calyx short, 4-toothed. Petals 4. Stamens 8 . . 2. Combretum. Calyx short, 5-lobed. Petals 5. Stamens 10 . . . 3. Poivrea. Calyx with a very long, slender tube, 5-toothed. Pe- tals 5. Stamens 10 4. Quisqualis. Tribe 1. Termihalie^j. 1. TEBMINTALIA, Linn. Flowers often polygamous. Calyx-limb deciduous, bell- shaped, 5-cleft ; lobes acute. Petals 0. Stamens 10, in a 110 TJ. COMBRETACEjE. double row, longer than the calyx. Ovary 2-3-ovuled. Style filiform, acutish. Drupe not crowned by the calyx, usually dry, indehiscent, 1-seeded. — FI. Cap. ii. p. 508. Trees or shrubs. T. sericea, Burch., our only species, has oblong, silky leaves crowded round the ends of the branches, and flowers in spikes shorter than the leaves. — Aapjes river. 2. COMBRETUM, Linn. Calyx funnel-shaped ; tube as short as the ovary or longer ; limb bell-shaped, 4-lobed, deciduous. Petals 4, small, in- serted between the lobes of the calyx. Stamens 8, in 2 rows, exserted. Ovary 2-5-ovuled; style exserted, acute. Fruit 4-winged, 1-celled, 1-seeded, indehiscent. Seed pendulous. — FI. Cap. ii .p. 508. Trees or shrubs. Leaves often opposite. Spikes terminal or axillary, some- times capitate ; flowers small, greenish. — 10 species, all Eastern or from Natal. 3. POIVREA, Comm. Limb of the calyx funnel-shaped, 5-lobed, deciduous. Petals 5. Stamens 10, protruded. Ovary 2-3-ovuled; style fili- form, protruded, acute. — FI. Cap. ii. p. 512. P. bracteosa , Hochst., our only species, is a Natal shrub, 8-10 feet high, with ovate or ovate-oblong glabrous leaves, and nodding, bracteate spikes of reddish flowers. 4. QUISQUALIS, Rumph. Tube of the calyx slender, produced much beyond the ovary ; limb 5-lobed. Petals 5, oblong or roundish, obtuse, longer than the calyx-teeth, imbricate. Stamens 10, inserted within the throat of the calyx, those opposite the petals longest. Ovary 4-ovuled ; style filiform, exserted, its base adhering to the calyx-tube. Drupe dry, 5-furrowed and 5-ribbed, 1-seeded. Seed pendulous. — FI. Cap. ii. p. 512 ; Thes. Cap. t. 130. Q. parvijlora , Gerr., our only species, discovered by Mr. Gerrard in the Natal country, has opposite, oval-oblong, acute, thinly pubescent or glabrate leaves and terminal spikes of slender flowers. Petals small, shortly oblong. Anthers subsessile, in 2 widely-separated rows. Order LII. MYRTACE^l. Flowers bisexual. Calyx-tube adhering to the ovary ; limb 4-5-parted or obsoletely lobed. Petals 4-5. Stamens indefi- nite, inserted with the petals on the fleshy margin of the calyx-tube. Ovary inferior, 2- or several-celled, with few or many ovules ; style filiform ; stigma simple. Fruit either a succulent berry or a dry capsule. Seeds without albumen. — Trees or shrubs. Leaves mostly opposite, quite entire, penni- LTT. MYRTACEJE. Ill nerved, with an intramarginal vein, almost always pellucid - dotted and aromatic or resin-scented, exstipulate. Flowers solitary or in cymes panicles or heads. — The Order is very abundant in South America and Australia, much less so in Asia, and very scantily represented in Africa. Tribe 1. Leptospermea:. Fruit dry, capsular, many-celled, dehiscent. Stamens much-exserted, free. Capsules 2-3-celled . 1. Metrosideros. Tribe 2. Myrte.e. Fruit a fleshy berry. Leaves dotted. Calyx-limb almost entire or repand. Petals concrete into a calyptra 2. Syzygium. Calyx-limb more or less deeply 4-5-parted. Petals separate. Calyx-tube turbinate. Petals 5. Flowers in ter- minal cymes 3. Acmena. Calyx-tube globose. Petals 4 (very rarely 5). Pe- duncles axillary 4. Eugenia. Tribes 3. Barringtoniea:. Fruit berried or dry, valveless. Leaves without pellucid dots. Petals 4. Stamens in many rows, connate at base . 5. Barringtonia. Tribe 1. LEPTOSPERMEiE. 1. METROSIDEROS, R. Br. Calyx-tube adhering to the ovary, not angular ; limb 5-cleft. Stamens 20-30, free, very long, exserted. Style filiform. Capsules 2-3-celled; cells many-seeded. Seeds wingless. — FI. Cap. ii. p. 521. Trees or shrubs. — M. angustifolia, Sm., our only species, is common by riversides in many parts of the colony. Its leaves are linear-lanceolate ; peduncles axillary, umbellate. Flowers yellowish. Tribe 2. Myrtejl 2. SYZYGIUM, Gsertn. Calyx-tube obovate ; limb nearly entire or repandly-lobed. Petals 4-5, roundish, joined into a cap (or calyptra) and falling off either in that state from the calyx, or immediately after expansion. Stamens numerous, distinct. Ovary 2-celled, with few ovules in each cell ; style simple. Berry 1-eelled, 1- or few-seeded. Seed globose ; cotyledons large, fleshy, nearly hemispherical ; radicle small, inserted between the co- tyledons below the middle, and concealed by them. — FI. Cap. ii. p. 521. Trees or shrubs. Leaves opposite, quite entire, glabrous. — 8. cor datum > Hochst., our only species, is a Natal tree, 30-40 ft. high, with subsessile, elliptic-cordate, leathery leaves, veiny and paler beneath. Cymes terminal, many-flowered. 112 LIT. MTltTACEJF,. 3. ACMENA, DC. Calyx-tube turbinate ; limb shortly bell-shaped, either sub- entire or more or less deeply 5-lobed, involute in bud. Petals 5, small, separate. Stamens numerous, distinct. Ovary 3- or 2-celled ; cells many-ovuled ; style simple, long or short. Berry globose or oval, 1-seeded. Australian or Indian shrubs. — A. Gerrardi , Harv. mss., lately found in Zululand by Mr. Gerrard, is nearly allied to A . Zeylanica. It is arborescent, with ovate-oblong, acuminate, closely-veined leaves, and terminal, 3-cho- tomous-multifid cymes ; calyx limb deeply 5-lobed ; ovary 2-celled. Calyx- tube nearly globose ; limb divided down to the ovary into 4, rarely 5 segments. Petals 4 or rarely 5. Stamens numerous, distinct. Ovary “ 11 1 rarely 2-celled. Seeds 1-2, large, roundish ; cotyledons very thick and confluent ; radicle very small. — FI. Cap. ii . p. 521. Trees or shrubs. Leaves opposite, pellucid-dotted. Peduncles axillary or terminal, 1- or several-flowered. — About 8 species (some undescribed), all Eastern or from Natal. Calyx- tube ovate ; limb 2-3-4-parted ; lobes ovate, obtuse, concave, persistent. Petals 4, coriaceous, attached to the ring at the base of the stamens. Stamens numerous, in several rows ; filaments filiform, long, distinct, combined at the base into a short ring, all bearing anthers. Ovary 2-4-celled, crowned by an urceolus sheathing the base of the style ; ovules 2-6 in each cell ; style simple. Bruit fleshy or corky, more or less 4-angled, crowned by the limb of the calyx, 1-celled. Seed solitary ; embryo fleshy, not separable into cotyledons and radicle, formed of 2 concentric, combined layers. — FI. Cap. ii. p. 523. Tropical trees, growing near the sea. — B. racemosa , Roxb., occurs near Natal. Blowers bisexual, regular. Calyx-tube enclosing the ovary, either quite adnate to it, or attached by its ribs or angles, leaving interspaces, or rarely quite free ; limb with valvate or imbricate aestivation. Petals on the summit of the calyx-tube, as many as its lobes, imbricate in bud. Stamens inserted with 4. EUGENIA, Linn. nearly globose, crowned Tribe 3. Barrihgtonie^:. 5. BARRIN GTONIA, Borst. Order LIII. MELASTOMACE.®. . LIII. MELASTOMACE.E. 113 the petals, either once or usually twice as many, those oppo- site the petals sometimes abortive ; filaments inflexed in bud ; anthers 2-celled, opening by terminal pores, or rarely splitting. Ovary usually plurilocular (rarely unilocular) ; ovules nume- rous ; style simple ; stigma undivided. Fruit a capsule or berry. Seeds without albumen. — Leaves opposite, entire, usually 3- or more ribbed, without pellucid dots, exstipulate. * Undershrubs, with 3-5-ribbed, rough leaves. Calyx -limb persistent, with accessory teeth ; anthers with a club-shaped appendage at base 1. Argyrella. Calyx-limb deciduous ; tube covered with bristly scales ; anthers with a 2-fid or 2-parted appendage at base 2. Dissotis. *'* Trees and shrubs with 1-nerved, glabrous leaves. Calyx-tube ovoid ; stamens 8 ; ovary 1-celled ... 3. Memecylon. Calyx-tube cylindrical ; stamens 5 ; ovary 5-celled . 4. Olinia. 1. ARGYRELLA, Naud. Calyx-tube bell-shaped ; limb of 5 ovate acute lobes, equal- ling the tube, twisted imbricate in aestivation, alternating with as many subulate accessory teeth (not stellato- setose at apex). Petals obovate. Stamens 10, unequal ; anthers linear-subu- late, opening by a single pore, recurved, undulate along the inner side, the 5 larger with the connective very long, pro- duced beyond the insertion of the filament into a clavate appendage ; the 5 smaller with a very short 2-dentate connec- tive, confluent with the apex of the filament. Ovary 5-celled, half-inferior, the upper half free, 5-lobed, tomentose ; style filiform. Capsules contained in the persistent calyx. Seeds cochleate, striate. — Nand. in Ann. Sc. Nat., Ser. 3. xiii. p. 300. Osbeckia, ex parte, FI. Cap. ii .p. 518. A. canescens ( Osbeckia canescens, E. Mey. ! ; Graham in Bot. Mag. ! t. 3790 ; O. TJmlaasiana , Hochst.) is an erect, virgate branching undershrub, found near Natal. All parts pale, tomentose with very minute, stellate hairs. Leaves sessile or subsessile, oblong-lanceolate, 5-nerved, netted- veined beneath. Flowers purple, handsome, corymboso-paniculate. 2. DISSOTIS, Benth. Calyx-tube ovoid, covered with palmately-cut scales, ad- hering by its ribs to the ovary or at length free ; limb 4-5- parted, deciduous, many-bristled at the apex. Petals obovate, ample. Stamens 10, unequal ; anthers linear- subulate, open- ing by a single pore, sub recurved, the 5 larger with the con- nective very long, produced beyond the insertion of the fila- ment into a deeply 2-lobed or 2-parted appendage ; the 5 smaller with a short connective, equally 2-lobed at base. Ovary crowned with bristles, 1 4-5-celled ; style equal or i Ill LIII. MELASTOMACEiE. slightly thickened upwards. Capsules included in the urceo- late calyx-tube, 5-celled. Seeds cochleate. — Benth. in Niger Flora , p. 346. Osbeckia, FI. Cap. ex parte , ii. p. 518. To this genus belong Osbeckia eximia, Sond., and O.phceotricha, ITochst., botli found near Natal ; the former has 5-merous, panicled flowers ; the latter 4-merous, capitate. 3. MEMECYLON, Linn. Calyx-tube hemispherical or subglobose, adnate to the ovary; limb very short or cup-like, obsoletely 4-toothed or subentire. Petals 4, oval. Stamens 8, mostly longer than the petals ; filaments filiform ; anthers short, with a thick connective, forming a conical spur at base, opening at base by a short slit. Ovary inferior, 1-celled ; ovules on a central placenta; style filiform; stigma simple. Perry crowned with the limb of calyx, frequently 1-seeded. Seeds with convolute cotyledons. — Endl. Gen. n. 6269; Benth. Hongkong Flora, p. 117. Trees or shrubs, chiefly Asiatic. Twigs 4- sided, with tumid nodes. Leaves opposite, impunctate, 1-ribbed, coriaceous, quite entire. Flowers axillary, tufted or subcapitate. — A species (as yet undescribed) of this genus has recently been found, by Mr. Gerrard, near Natal. 4. OLmiA, Th. Calyx tubular, adhering to the ovary ; limb of 5, rarely 4, minute teeth. Petals 5-4. Scales 5-4, minute, obovate, al- ternating with the petals. Stamens 5-4 ; filaments very short, adnate to the calyx below the scales ; anthers subglobose, 2- celled, introrse, with a thick connective. Ovary inferior, 5-4-celled ; cells 3-ovuled ; style subulate ; stigma obtuse. Fruit drupaceous, subglobose, truncate, 3-4-cellcd ; cells mostly 1-seeded. Seeds with a spirally rolled embryo.— FI. Cap. ii. p. 519. O. cymosa , Thunb., is a glabrous shrub or tree, with 4-siaed twigs. Leaves opposite, petioled, coriaceous, green, shiny above, impunctate, 1- ribbed and penninerved, quite entire, varying in shape from obovate obtuse to ovato-lanceolate acuminate. Cymes terminal or axillary, densely many- flowered. Flowers white. — Found in kloofs, throughout the colony, and beyond the Eastern frontier. Order LIY. LYTHRARIEiE. Flowers bisexual, rarely polygamous. Calyx free, persis- tent, tubular or bell-shaped, 4-12-toothed, often ribbed. Pe- tals on the summit of the calyx-tube, deciduous. Stamens usually inserted at or below the middle of the calyx-tube, as many or twice as many as the petals ; rarely inserted at the summit of the tube and opposite the petals. Ovary free, 2-6- LIV. LYTHRARIEiE. 115 celled, with several ovules in each cell ; style single, filiform. Capsules enclosed in the base of the persistent calyx-tube, or girt with the calyx. Seeds without albumen. — Herbs or shrubs, with simple, entire, exstipulate leaves. Herbaceous plants. Stamens 2. A minute, subsimple plant ; leaves subu- late 1. SUFFRENIA. Stamens 4 or more. Calyx bell-shaped. Capsules longer than the calyx 2. Ammannia. Calyx tubular, cylindrical. Ovary 2-celled ... 3. Lythrum. Calyx bell-shaped. Ovary 4-celled. Capsules hid- den in the calyx-tube 4. Nesa:a. • Shrubs or trees. Stamens 5, opposite the petals, and inserted with them ; leaves pellucid-dotted, aromatic ; flowers small, panicled 5. Heteropyxis. Tribe 1. Lythrarie^). (G-en. 1-4.) 1. SUFFRENIA, Bellard. Calyx bell-shaped, 4-toothed, with 4 alternate, exterior, very minute or obsolete alternating teeth, the inner teeth ovate, acute, erect. Petals 4 (or OF), very minute, fugacious. Stamens 2, inserted within the calyx-tube, included ; filaments short ; anthers subglobose. Ovary sessile, 2-celled ; ovules numerous ; style filiform ; stigma capitate. Capsules oblong, girt by the calyx, 2-valved. — Thes. Cap. t. 189. Small marsh plants. — S. Capensis , H., our only species, was found by Mr. Cooper, on the Draakensberg, Orange Free State. Stems 1-2 inches high, tufted, subsimple. Leaves opposite, linear- subulate, 2-toothed. Flowers axillary, £ line long. Petals either 0 or not seen by me. 2. AMMANNIA, Linn. Calyx 2-bracteolate, bell-shaped, 4-14-nerved, with 4-7 erect, flat teeth, and frequently as many spreading minute or obsolete alternating teeth. Petals 4-7. Stamens as many or twice as many as the petals. Ovary 2-5-celled ; style simple, persistent ; stigma capitate. Capsules girt by the calyx, ovato-globose, delicately membranous, either splitting across or opening by valves. — FI. Cap. ii. p. 515. Small, much-branched herbs, growing in wet soil. Leaves opposite, quite entire. Flowers small, axillary, sessile or pedicelled, usually pink or red. — 2 South African species, both Eastern. 3. LYTHRUM, Linn. Calyx bracteated, cylindrical, ribbed or striate, with 4-6 erect broad teeth, and as many spreading, smaller, alternating teeth. Petals 4-6. Stamens inserted in the middle or at the base of the calyx-tube, twice as many as the petals or rarely i 2 116 LIT. LYTIIRARIEvE . only as many. Ovary 2-celled ; style filiform ; stigma capi- tate. Capsule oblong, enclosed in the calyx, 2-celled, many- seeded. — FI. Cap. ii. p. 516. Herbs, occasionally half-shrubby, with opposite alternate or whorled, entire leaves. Flowers axillary, purple or purplish, often handsome ; petals quickly withering.— 3 species, of which 2 are Eastern and endemic, the third nearly cosmopolitan. 4. NESJEA, Comm. Calyx hemispherical, bracted or bractless ; ribbed or striate, with 4-6 erect, triangular inner teeth, and as many smaller, narrow, or horn-like, spreading outer teeth.. Petals 4-6. Sta- mens 8-12, nearly equal. Ovary sessile, globose, 4-celled. Capsules globose or subglobose, covered by the calyx, many- seeded. — FI. Cap. ii. p. 517. Herbs, with lanceolate or oblong, nearly sessile, obtuse or acute, entire leaves. Peduncles axillary, longish, 3-flowered or capitate, and many- flowered at the summit. — N. floribunda, Sd., our only species, grows near Natal. Flowers capitate. Tribe 2. Heteropyxidea:. (G-en. 5.) 5. HETEROPYXIS, Harv. Plowers polygamous. — Male : Calyx cup-shaped, with 5 erect, triangular lobes. Petals 5, ovate, inserted in the throat of the calyx, subsessile, pellucid-dotted. Disk, lining the calyx-tube, thin. Stamens 5, inserted with the petals and opposite to them ! ; filaments subulate ; anthers 2-celled, versa- tile. Abortive ovary 2- rarely 3-eelled, many-ovuled ; style short ; stigma obtuse. — Pemale : Calyx 10-nerved, with 5 erect, triangular lobes, and as many minute, alternating denticles. Petals and stamens as in , but the 2-celled anthers abortive. Ovary 2-celled ; ovules many, on axile placentas ; style fili- form, much-exserted ; stigma capitate. Capsules oblong, 2- celled, girt by the persistent calyx. Seeds (unripe only seen). — Thes. Cap. t. 128. An aromatic tree or large shrub, found near Natal, with alternate, short- petioled, lanceolate, pellucid-dotted, penninerved leaves, and dull white fragrant flowers in terminal or axillary panicles. When I published this genus in the ‘ Thesaurus ’ I was only cognizant of the male flowers, in which, however, the ovary, ovules and style, though abortive, were so completely organized that the flower passed as bisexual. Recently I have received from my zealous and most obliging friend Mr. Gerrard, the true female flower and half-ripe fruit. These afford an additional calycine character, which completely reconciles Heteropyxis to Lythrariece. Order LY. ONAGRARIE^. Flowers bisexual, rarely dioecious. Calyx-tube adnate with LV. ONAGRARIEiE. 117 the ovary, its limb 4-5-lobed, valvate or open in bud. Petals on the margin of the calyx-tube or 0. Stamens as many or twice as many as the calyx-lobes, and opposite them. Ovary inferior, 2-4-5-celled ; style mostly filiform (rarely splitting) ; stigma 2-^4-5-lobed. Fruit a capsule or berry. Seeds exalbu- minous or nearly so. — A considerable Order, chiefly of tempe- rate climates. Leaves opposite or alternate, exstipulate, simple, entire, or variously lobed or cut. Flowers either axil- lary or in racemes or spikes, often showy. The well-known garden-flowers (Enothera and Fuchsia belong to this Order. Tribe 1. Jussieu. Flowers bisexual. Petals spirally twisted in bud (sometimes 0). Ovary 4-5-celled ; ovules numerous ; style filiform. Fruit capsular. Seeds many, not winged. Limb of the calyx persistent ; capsule septicidal. Stamens 8-10 1. Jusslea. Stamens 4-5 2. Ludwigia. Limb of the calyx deciduous ; capsule loculicidal. Calyx-tube much produced beyond the ovary. Seeds nude 3. (Enothera. Calyx-tube not longer than the ovary. Seeds with a silky tuft 4. Epilobium. Tribe 2. Montinie.®. Flowers dioecious. Petals imbricate. Ovary imperfectly 2-celled ; ovules numerous ; style splitting through the middle. Fruit capsular, 2-valved. Seeds broadly winged. A glabrous, small shrub, with alternate, simple leaves . 5. Montinia. Tribe 3. Trapes. Flowers bisexual. Petals imbricate. Ovary 2-celled ; ovules solitary ; style filiform. Fruit a hard, 1-seeded nut. Water plants, with rhomboid, long-petioled, floating leaves, and spinous or homed nuts 6. Trapa. Tribe 1. Jussieje. (Gen. 1-4.) 1. JUSSIiEA, Linn. Calyx-tube not produced beyond the ovary ; the limb 4-5- parted, persistent. Petals 4-5. Stamens 8-10. Stigma ca- pitate. Capsule 4-5-celled, crowned by the calyx-lobes and opening longitudinally between the ribs. Seeds numerous, small, naked. — FI. Cap. ii. p. 504. A considerable genus, chiefly of tropical America. — 2 species, one of them endemic, occur at Natal. Leaves alternate, entire. Flowers axillary, soli- tary, yellow. 2. LUDWIGIA, Linn. Characters as in Fussicea, except stamens 4-5. Petals want- ing in L. palustris. — FI. Cap. ii. p. 504. 2 species, neither endemic, found in the Eastern district and Natal : L. palustris (without petals) and L. jussieeoides (with petals). 118 LV. ONAGRARIE./E. 8. (ENOTHERA, Linn. Calyx-tube much produced beyond the ovary, deciduous ; limb 4-parted. Petals 4, obcordate. Stamens 8. Stigma 4- lobed or capitate. Capsule various in form and texture, 4- celled, 4-valved, many-seeded. — FI. Cap. ii. p. 505. A large genus, of American origin. — 2 species (“ Evening Primroses ”) are naturalized near Capetown. 4. EPILOBIUM, Linn. Calyx-tube not produced b'eyond the ovary ; limb deeply 4-lobed or 4-parted, deciduous. Petals 4, obovate or obcor- cordate. Stamens 8. Stigma clavate or 4-lobed. Capsule linear, 4-sided, 4-celled, 4-valved, loculicidal. Seeds with a tuft of silky hairs at one end. — FI. Cap. ii. p. 506. Herbs of the temperate zones. Leaves alternate or opposite, entire or serrulate ; flowers axillary or in terminal bracteate spikes, purple or rosy, rarely yellowish. — 3 species, of which one is said to be endemic. Tribe 2. Moutinie^. (G-en. 5.) 5. MONTINIA, Linn. Flowers dioecious. Calyx-tube wholly adnate to the ovary ; limb short, persistent, 4-toothed, with open aestivation. Pe- tals 4, ovate, epigynous, imbricate, deciduous. Disk (in the male) fleshy, 4-angled. Stamens 4 ; filaments short ; anthers adnate, slitting lengthwise ; pollen trigonous. Ovary inferior, imperfectly 2-celled, with very prominent, fleshy, multiovulate placentas, filling up the greater part of the cavity ; style single, short ; stigma large, deeply 2-lobed. Capsule ligneous, crowned by the style (now become 2-parted) and the calyx- limb, 2-celled, splitting through the centre. Seeds 4-6 in each cell, compressed, with a broad, membranous, marginal wing. — FI. Cap. ii. p. 307. M. acris, Linn., the only species, is a glabrous, somewhat glaucous shrub, common all over the colony. Leaves varying from oblong to lanceolate or linear, entire, margined, 1-nerved, veinless. Flowers small, white, the males in terminal corymbs, the females mostly solitary. Tribe 3. Trapes. (Gen. 6.) 6. TRAP A, Linn. Calyx-tube adnate to the lower part of the ovary ; the limb half-superior, 4-parted, valvate in bud, then spreading, persis- tent, with spinous lobes. Petals 4, inserted under a fleshy, epigynous disk, imbricate in bud, the margins plaited and wavy, spreading, deciduous. Stamens 4, inserted with the LY. ONAGRARIEiE. 119 petals. Ovary half-inferior, 2-celled ; ovule 1 in each cell ; style cylindrical, simple ; stigma obtuse. Fruit a somewhat horny nut, 2-4-spined from the enlarged, spinous calyx-lobes, often with supplementary spines or prominences on the disk, 1-celled, 1- seeded. Seed without albumen. Water plants, floating on lakes and ponds. — T. bispinosa, Foxb., occurs at Natal, where its singularly-formed fruits, which are full of farinaceous substance, are eaten by the Zulu Kaffirs. The submerged leaves are opposite, pinnatisect, with very narrow lobes, the floating crowded in a rosulate cluster, petioled, rhomboid, toothed, the petioles swollen and hollow in the middle, forming floats ; flowers axillary, solitary. The black-brown oval nut is crowned with 2 straight barbed horns ; and its either side, in the spe- cimens I have received, is curiously furnished with prominences arranged like the features of a diabolical face ; a long, hooked nose, a pair of peering eyes, and a pursed-up mouth. Specimens sent to me by Mr. Sanderson, of Natal. Order LVI. TUKNERACEjE. Flowers bisexual. Calyx tubular, free, 5-cleft. Petals 5, inserted in the tube or at its summit, twisted in bud, deci- duous. Stamens 5, alternating with the petals. Ovary 1- celled, with 3 parietal placentas ; styles 3, separate or partially connate ; stigmas fimbriate. Fruit capsular, 3-valved ; valves placentiferous. Seeds albuminous. — A small Order; scarcely differing from Passifloreae. 1. TURNER A, Plume. Calyx coloured, tubular-funnel-shaped, more or less deeply 5-parted. Petals inserted in the throat of the calyx. Sta- mens 5 ; style 3-fid at the apex ; stigma multifid. Capsule ovate or oblong, 3-valved. — FI. Cap. ii. p. 599 ; Thes. Cap. t. 140. Undershrubs or shrubs, chiefly American.— T. Capensis, H., our only species, is a very dwarf, hairy plant, with crowded, lanceolate, serrate leaves, and axillary, small, white flowers. It grows near the Aapjes river. Order LVLL LOASACEiE. Flowers bisexual, regular. Calyx adnate to the ovary; limb 5-lobed. Petals epigynous, 10, in 2 rows, those of the outer row concave. Stamens indefinite, in many parcels. Ovary 3-celled. — An Order almost exclusively American, chiefly near the west coasts of North and South America. 1. KISSENIA, R. Br. Calyx- tube 10-ribbed ; limb 5-parted, the lobes equal, en- 120 IjVII. LOASACEiE. larged in fruit, persistent. Petals 10, deciduous, inserted at the summit of the calyx-tube, 5 outer roundish, concave, 5 inner smaller, ligulate, angularly bent. Stamens indefinite, those of the outer row barren, with cordate bases. Ovary turbinate, 3-celled ; cells 1-ovuled ; styles 3, erect, subconnatc. — FI. Cap. ii. p. 502. The only species is K. spathulata, Br. ( Fissenia Capensis , Endl. ; Thes. Cap. t. 98), a robust, bristly undershrub, with alternate, petioled, 5-7- lobed, toothed leaves, and rather large yellow flowers, disposed in scorpioid, terminal, bracteate cymes. It is a native of Namaqualand, the mouth of the Oariep, etc., and extends thence northwards through tropical Africa to Arabia. Order LYIII. PASSIFLOREjE. Plowers bisexual or unisexual. Calyx monophyllous, free, tubular or rotate ; limb 3-4-5 -cleft or parted. Petals as many as the calycine lobes, often herbaceous on the outside, con- tinuous with the apex of the calyx-tube, or inserted much within the tube, persistent, separate or (rarely) united in a bell-shaped corolla. Corona various, exterior to the fertile stamens. Stamens as many or twice as many as the caly- cine lobes, free or monadelphous ; anthers versatile or fixed. Ovary mostly stipitate, 1-celled ; ovules many or few, on pa- rietal placentas ; styles as many as the placentas ; stigmas thickened. Fruit a berry or capsule. Seeds on long cords, mostly arillate, with a furrowed and ridged seed-coat, albumi- nous.— Climbing (rarely erect) shrubs or herbs. Flowers bisexual. Calyx 3-parted. Petals 3, half-herbaceous. Co- rona double, the outer fringed 1. Teyphostemma. Flowers monoecious or dioecious. Dioecious. Corolla 0 or polypetalous. Calyx 5-fid. Disk with 5 conspicuous glands . 2. Modecca. Calyx 5-parted. Disk without glands ... 3. Ophiocaulon. Monoecious. Calyx 3-5-parted. Corolla monope- talous, 3-5-lobed, herbaceous. A vine-like climber. Capsule slender, pod-like 4. CeeatiosicyoS. A suberect, low herb. Capsule ovoid ... 5. Achaeia. 1. TEYPHOSTEMMA, Harv. Flowers hermaphrodite. Tube of the perianth short, obco- nical ; limb 6-parted, in 2 rows, the 3 inner segments unequal, 2 of them larger, herbaceous, and white-edged, the third linear and petaloid. Corona perigynous, annular, double, the outer fringed, the inner entire or crenulate, bearing the stamens. Stamens 5, within the rim of the inner corona; filaments subulate ; anthers erect, sagittate, 2-celled. Ovary LVIII. PASSIFLOltEiE. 121 subsessile, 1-celled ; ovules few, on 3-4 parietal placentas ; styles 3-4, filiform ; stigmas capitate. Capsule shortly stipi- tate, membranous, 3-4-valved, few-seeded. Seeds pendulous, with a membranous aril, areolate-corrugate. — FI. Cap. ii. p. 499 ; Thes. Cap. t. 51. T. Sandersoni, H., the only species, grows near Natal. Stems subsimple, 4 inches to 4 feet high, the taller somewhat climbing. Leaves alternate, sessile or subsessile, 2-stipuled, ovate or ovato-lanceolate, distantly toothed, netted-veined. Flowers 2-5 lines diameter, greenish, in short axillary ra- cemes. 2. MODECCA, Lam. Flowers dioecious. Calyx tubular-conical, bell-shaped, 4-5- lobed. Corolla of 4-5, ovate oblong or linear, sometimes fim- briate petals, smaller than the calycine lobes, and inserted either at the summit of the calyx-tube or far within it. — Male : Stamens 4-5, inserted in the bottom of the calyx and opposite its lobes ; filaments subulate, connate in a ring at base ; anthers introrse, 2-celled, erect. A rudiment of an ovary. Grlands 4-5, opposite the petals. — Female : Abortive stamens 5, subulate, girding the ovary. Ovary stipitate or subsessile, 1-celled ; ovules numerous, on 3 parietal placentas ; style short or 0 ; stigma dilated or fimbriate. Capsule thinly fleshy (leathery when dry), subglobose, 3-valved, many-seeded. Seeds arillate, corrugate. — FI. Cap. ii. p. 499 (species 1 and 2) ; Thes. Cap. t. 12. Mostly vine-like climbers, often with tendrils. Leaves alternate, un- divided or lobed, the petioles mostly 2-glanded at the apex. Stipules ob- solete or none. Peduncles axillary ; flowers small, greenish. — About 6 (some undescribed) South African species, all from Natal or Zululand. 3. OPHIOCAULOH, Hook. f. Flowers dioecious. Calyx-tube very short ; lobes spread- ing, marked with black lines. Petals 5, subserrate. — Male : Stamens 5, almost hypogynous ; filaments very short ; an- thers narrow-linear. G-lands of disk 0 or very obscure. — Fe- male : Abortive stamens 5, subulate. Ovary sessile, 1-celled ; stigmas 3, flabellate ; ovules numerous, on 3 parietal placentas. Capsule sessile, coriaceous, 3-valved, many-seeded. An African genus of climbing shrubs, with smooth, snake-like stems, full of gum. Leaves alternate, green, glaucous beneath, entire or lobed. Ten- drils simple or branched. Flowers small, greenish, in long, peduncled, axillary cymes. — 1 species, Modecca ? gummifera , Harv. FI. Cap. ii. 501, a native of the Eastern districts. 4. CERATIOSICYOS, Nees. Flowers monoecious. — Male : Calyx-tube short ; limb of 122 LYIII. PAS3IFL011E2E. 4-5 slender, subulate lobes. Corolla continuous with the calyx-tube, bell-shaped, 4-5-lobed. Stamens 4-5, inserted in the base of the calyx, free, alternate with the lobes of the co- rolla, dilated upwards ; anthers adnate to a club-shaped con- nective, the cells slightly separated, introrse. Grlands as many as the stamens, and alternating with them, oblong, fleshy. — Female : Calyx-lobes obsolete. Corolla as in the male. Grlands as in the male, but smaller. Ovary stipitate, unilocu- lar ; ovules numerous, on 4-5 parietal placentas ; stigmas 4-5, subsessile, channelled, 2-lobed. Capsule pod-like, slender, 4-5-valved, several-seeded. — FI. Cap. i\.p. 501. C. EcJclonii , Necs, the only species, is a slender, nearly glabrous climber, growing in the Eastern district and at Natal. Leaves palmately 5-7-lobed, membranous. Flowers axillary, greenish, the males in racemes, the fe- males solitary. 5. ACHARXA, Th. Flowers monoecious. — Male : Calyx 3-4-parted. Corolla bell-shaped, 3- (rarely 4-)lobed, herbaceous, continuous with the short calyx-tube. Stamens adnate to the tube of the co- rolla for more than half their length, as many as the lobes and alternate with them ; filaments dilated upwards, subexserted ; anthers adnate to a broad, 2-lobed connective, the cells sepa- rated, introrse. Grlands 3-4, fleshy, in the base of the perianth, alternating with the stamens. — Female: Calyx and corolla as in the male, but enlarged in fruit, persistent. Ovary subses- sile, with 3 glands at base, 1-celled ; ovules few, on 3-4 pa- rietal placentas ; style 3-4-cleft ; stigmas 3-4, channelled, 2- lobed. Capsule shortly stipitate, ovoid, membranous, 3-4- valved, few-seeded. Seeds pendulous, with a small arillus. — FI. Cap. ii. p. 501. A. tragioides , Th., the only species, is a small, thinly-pubescent, branch- ing herb, growing in shady places of Uitenhage and Albany. Leaves alter- nate, petioled, 3-lobed, and cut. Flowers small, green, axillary. Order LIX. CXJCUEBXTACE^]. Flowers unisexual. Calyx 5-lobed, adnate. Corolla mostly monop etalous, 5-lobed or 5 -parted, continuous with the sum- mit of the calyx-tube. Stamens inserted within or at the mouth of the calyx-tube, 5-3-2, either free or with the anther- cells monadelphous, or the anthers and filaments variously co- hering ; anthers usually linear, adnate, extrorse, cells straight curved flexuous or conduplicate. Ovary inferior, usually with prominent, parietal placentas, often meeting in the centre, sometimes 2-3-celled ; ovules many, usually horizontal ; styles LIX. CUCURBITACEiE. 123 united or distinct ; stigmas 1-3, entire or 2-lobed or parted. Fruit a succulent or dry berry or gourd. Seeds lying in pulp or corky or fibrous substance, exalbuminous. — Herbs, with prostrate or climbing, very long stems, and simple or branched tendrils. Leaves alternate, petioled, exstipulate, palminerved, often multifid or lobed, sometimes 3-5-foliolate. Tribe 1. Cucurbitea:. Ovules horizontal. Stamens usually three. Fruit decaying or bursting irregularly, rarely 3-valved. Seeds not winged. § 1. Cucumerinecs. Anther-cells flexuous or conduplicate (rarely straight in Cucumis and Momordica). Corolla rotate or bell-shaped ; limb divided wholly or nearly to the base. Calyx-tube short. Connective of anthers prolonged beyond the cells 1. Cucumis. Connective not prolonged. Calyx-tube without internal scales. Ten- drils usually branched 2. Citrullus. Calyx-tube with 2 scales between the sta- mens. Tendrils simple 3. Momordica. Calyx-tube long (at least in the female). Fruit a few-seeded berry 4. Trochomeria. Gourd dry and fibrous within ; seeds flat . . 5. Luffa. Gourd fleshy ; seeds with tumid border . . 6. Lagexaria. Corolla bell-shaped, 5-lobed to or above the middle, rarely below it. Tendrils simple. Fruit a berry 7. Cephalandra. Tendrils forked or branched. Fruit a gourd . . 8. Cucurbita. § 2. Melothriea. Anther-cells straight. Fruit a berry. (See Cucumis and Momordica in Tribe 1.) Anthers 3, all 2-celled. Seeds flat 9. Zehxeria. Anthers 3, 2 of them 2-celled, the other 1-celled. Female flowers with 3 staminodia. Style with a tumid annulus at base. Seeds tumid . . . 10. Rhynchocarpa. Female flower with 3 staminodia. Style without disk at base. Seeds tumid 11. Pisosperma. Female flower without staminodia. Style with a cup-shaped disk at base. Seeds compressed . 12. Mukia. Tribe 2. Zanoniea:. Ovules and seeds pendulous. Stamens 4 , anthers 1-celled, straight, cohering in pairs. Fruit opening at the apex. Seeds with a broad wing 13. Gerrardanthus. Tribe 1. Ctjcfrbite^:. 1. CUCUMIS, Linn. Flowers monoecious or dioecious. — Male : Calyx bell-shaped, 5-fid. Corolla spreading, its limb 5-parted. Stamens 3, in- serted at the base of the corolla, one 1-celled ; anthers flexuous, rarely straight, terminated by a papillose, lobed prolongation of the connective. — Female : Calyx and corolla as in the male ; m LIX. CUCURBITAGE^. stigmas 3, thick. Fruit a gourd, 3- or spuriously 6-celled, many-seeded. Seeds oval, compressed, not margined. — FI. Cap. ii. p. 494. Annuals or perennials, scabrous, with succulent stems, rarely wanting tendrils, and angular or deeply-lobed leaves. Flowers axillary, solitary or tufted, yellow. — 9 ascertained Cape species, several others from Natal, of which incomplete specimens have as yet only reached us. 2. CITRULLUS, Schrad. Flowers monoecious. — Male : Calyx bell-shaped, deeply 5-fid. Corolla 5-parted, flattish. Stamens 3, inserted at the base of the corolla, 2 bilocular, deeply parted, the third unilocular ; connective without any terminal appendage ; anther-cells flexuous. — Female: calyx and corolla as in the male. Ovary with 3 prominent placentas, villous or smooth ; style 3-fid ; stigmas 3, thick. Fruit a globose, rarely oblong, 3- or 6- celled, many-seeded gourd. Seeds oval, compressed, with ob- tuse margins. — FI. Cap. ii. p. 492. C. vulgaris, a prostrate plant, with deeply 3-5-lobed leaves, forked ten- drils, and axillary yellow flowers, occurs on sandy flats in many places. It is the “Kaffir Water-melon” and “ Bitter Apple ” of the colonists, and a wild variety of the common European and Asiatic Water-melon. 3. MOMOKDICA, Linn. Flowers monoecious or dioecious. — 'Male : Panicled and ebrac- teate or bracteate or solitary, with the peduncle bearing a large, sessile, clasping bract. Calyx 5-cleft, with a yery short tube, closed at the base with 2 or 3 horizontal or incurved scales. Corolla 5-parted to the base, much longer than the calyx. Stamens 3, one 1-celled ; anther-cells flexuous or con- duplicate, rarely straight, free or connate. Ovary with 3 pla- centas ; style simple ; stigmas 3. Glourd fleshy, not fibrous, prickly, often bursting when ripe, with or without elasticity. Seeds compressed or tumid, enveloped in a fleshy pulp. — FI. Cap. ii. p. 491. Annual or perennial climbers, with petiolate, lobed or compound leaves, simple, rarely 2-fid tendrils and yellow or white flowers. — 4 ascertained Cape species, dispersed, and some imperfectly known, undesci’ibed. 4. TROCHOMERIA, Hook. f. Flowers monoecious or dioecious. Calyx with a cylindrical or funnel-shaped tube, and a 5-parted limb. Corolla spread- ing, 5-parted to the base; the lobes sometimes very long. Stamens 3 ; filaments inserted within the calyx-tube ; anthers subexserted or included, connivent ; cells conduplicate, 2 bilo- cular, the third unilocular. Female perianth as in the male. LIX. CUCURBITACEiE. 125 Ovary 3-celled ; ovules few ; style cylindrical ; stigma fleshy, 3-lobed. Fruit a berry ? — Thes. Cap. t. 9(5, 182, 183. Perennial climbers, with simple tendrils and more or less deeply-lobed leaves. Several have a large ciliate or pectinate bract at the base of the peduncle, others want it. — To this new genus are referred the following species of c Flora Capensis’ : — Zehneria Garcini ( Trochomeria Hoolceri , H.), Z. pectinata, Z. Wyleyana , Z. macrocarpa, Z. debilis , and Lagenaria ? sa- gittate/,. The true Zehneria Garcini of Stocks is different from the species so named in * Flora Capensis.’ 5. LUFFA, Tournef. Flowers monoecious, rarely dioecious. Calyx-tube in the male bell-shaped or turbinate, in the female oblong-clavate ; limb 5-toothed. Petals 5, somewhat deciduous. — Male : Sta- mens 3, exserted, 2 bilocular, deeply 2-parted, the third uni- locular ; anther-cells very flexuous, marginal on a very broad connective.— Female : Stamens abortive. Style 3-cleft ; stig- mas reniform or 2-parted. Gourd at length dry and fibrous within, usually opening by the falling of a terminal lid, some- times indehiscent. Seeds broadly oval, flattened. — FI. Cap. ii. p. 490. Climbers, with angular stems, branching tendrils, palmate-lobed leaves and large, yellow flowers. Male flower racemose ; female solitary. — 2 Cape species, both Eastern. 6. LAGENARIA, Ser. Flowers monoecious. Calyx bell-shaped ; segments broadish. Corolla (white) 5-parted ; petals obovate, springing from within the calycine rim. — Male : Stamens 3, included ; anthers subsessile ; cells very flexuous. — Female : Style scarcely any ; stigmas 3, subsessile, thick, 2-lobed. Gourd hard, dry, inde- hiscent. Seeds numerous, obovate, compressed, with a tumid border. — FI. Cap. ii. p. 489. Climbing, softly-pubeseent annuals. Tendrils 2-fid. Leaves cordate, nearly entire; flowers axillary. Fruit often large, pear-shaped or clavate. — L. vulgaris occurs spontaneously in gardens of Kaffraria. 7. CEPHALANDEA, Schrad. Flowers dioecious. — Male : Calyx short, campanulate, 5- toothed. Corolla bell-shaped, semi-5-fid, veiny. Filaments 3, inserted at the mouth of the calyx, free, but the connectives connate, united into a globose, antheriferous head; anthers distinct, 2 of them 2-celled, the other 1-celled ; cells flexuous. — Female : Calyx and corolla as in the male. Ovary oblong, with 3 placentas ; style with a thick, lobed stigma. Fruit a many-seeded, smooth berry. Seeds compressed, obliquely subattenuated at base. — FI. Cap. ii. p. 492. 126 LIX. CUCURBITACEiE. Four ascertained species, some others imperfectly known, all Eastern and from beyond the Eastern frontier. Herbaceous, climbing perennials, with 5-lobed leaves, simple tendrils, yellow flowers, and purple fruit. 8. CUCURBITA, Linn. Flowers monoecious. — Male : Calyx short, bell-shaped, 5-fid. Corolla bell-shaped, 5-fid. Stamens 3 ; filaments 4 ; anthers one 1-celled and two 2-celled, very sinuous. — Female : Calyx and corolla of male. Sterile stamens. Ovary 3-5-celled, many- ovuled ; style 3-fid ; stigmas thickened, 2-lobed. Grourd many- seeded. Seeds ovate, compressed, with a tumid margin. Annuals, with branched tendrils and cordate or 3-5-lobed leaves. — To this belong the universally-cultivated Gourds and Pumpkins of gardeners. 9. ZEHNERXA, Endl. Flowers monoecious or dioecious. — Male : Calyx hell-shaped, 5-toothed. Corolla spreading, 5-parted. Stamens 3 (some- times 4), inserted deeply within the calyx-tube, separate ; an- thers all 2-celled, on a broad fringed connective. — Female : Calyx and corolla as in the male. Staminodia 3, club-shaped. Style inserted in a tumid, 3-lobed epigynous disk, 3-fid ; stigmas 3, fleshy. Fruit a few-seeded, subglobose berry, with a tough skin. Seeds flattish. — FI. Cap. ii. p. 485 (in part). Perennial climbers, with simple tendrils and cordate, angular or lobed leaves. Flowers small, white ; the males peduncled, racemose j female in subsessile imperfect umbels or tufts. — 5 Cape species, dispersed. 10. RHYN CHOC ARP A, Schrad. Flowers monoecious or dioecious. — Male : Calyx bell-shaped, 5-fid. Corolla spreading, 5-parted. Stamens 3 (rarely 4), in- serted in the tube of the calyx ; filaments short ; anthers free or connivent, two 2-celled, the other 1-celled; cells oblong, straight, the connective rarely crested. — Female : Calyx and corolla as in the male. Staminodia 3. Style inserted in a tumid, epigynous disk. Fruit a sharply-beaked or taper- pointed, few-seeded, 3-celled berry. Seeds tumid. — FI. Cap. ii. p. 483 (Coniandra). Tuberous-rooted, climbing herbs, with simple tendrils, palmate or digi- tate-parted, rarely reniform, entire leaves, and small, greenish flowers. — 7 species, dispersed. 11. PISOSPERMA, Sond. Flowers monoecious, aggregated on radical, leafless branches, the males racemose, the female subsolitary. — Male : Calyx- tube bell-shaped ; lobes 5, lanceolate. Petals 5, oblong. Sta- mens 3 ; filaments short ; two anthers 2-celled, the other 1- celled ; cells linear, straight. — Female : Calyx and corolla as LIX. CUCURBITACE2E. 127 in the male. Staminodia 3. Style 1, without any epigynous disk ; stigma thick, lobed. Fruit a small berry with a tough coat, subglobose, apiculate, 3-celled, (3-12-seeded. Seed tumid. — FI. Cap. ii. p. 498. A tuberous-rooted perennial. The flowers are borne on very short, radical branches close to the ground. They are pale yellow, striped with green ; the males numerous, females few. When the little fruit is almost ripe, the root sends out long, twining, leafy stems, bearing small, simple tendrils. Leaves pedatifid, with linear lobes ; the middle lobe very long. — P. Capense , Sond., grows beyond the Eastern frontier. 12. Mina A, Arn. Flowers monoecious. — Male : Calyx campanulate, 5-toothed. Corolla 5-parted ; lobes obtuse. Stamens 3, inserted in the tube of the calyx ; filaments free ; two anthers 2-celled, the other 1-celled ; cells linear, straight, cohering ; connective produced into a conical point. — Female : Calyx and corolla as in male. Ovary ovoid, setulose, with 2-3 placentas ; style short, insertedin an annular, fleshy, epigynous disk. Stami- nodia 0. Fruit a small, smooth or bristly, globose, few-seeded berry. Seeds oblong-oval, subcompressed, surrounded by a broad or narrow rim, scrobiculate. — FI. Cap. ii. p. 488. M. scabrella, Arn., a common annual plant of tropical Asia, occurs near Natal. Stem climbing, with simple tendrils, and angled or lobed hairy leaves. Flowers small, yellow ; the males tufted, females solitary or ag- gregated. Tribe 2. Zanonie^. 13. GEERARDANTHUS, Harv. Flowers dioecious? — Male: Calyx small, rotate, 5-lobed. Corolla rotate, 5-parted to the base ; lobes linear-oblong, fleshy, 2 rather larger than the others. Stamens 4, and 1 sterile ; filaments separate ; anthers 1-celled, linear, cohering in pairs.— i-Female : Calyx and corolla as in the male. Ovary long, tapering at base, 3-cornered, with thick placentas and several pendulous ovules on each placenta. Fruit a long, funnel-shaped, nearly dry, at length membranous gourd, 3- valved at top. Seeds many, flat, with a crustaceous testa, and long membranous wing. Gr. megarhiza, Dene, and Harv., the only species, grows in the Natal colony. It has a large, placentiform, tuberous root, lying on the surface of the soil, 3-4 feet diameter, 1-2 feet thick, slightly acrid and bitter, and used, as Mr. Gerrard informs me, “ by the Kaffirs for various medicinal purposes ; among others, they give it to heifers, after the first calf, to in- crease the quantity of milk ; but it appears to deteriorate the quality, for, it is said, that cows give no butter during the time they are taking the root.” Stem woody, tuberculated, very long, rising to the tops of the highest trees, the branches hanging down in graceful, pendulous wreaths. All parts of 128 LIX. CUCURBTTACEiE. stem and foliage are glabrous. Branchlets slender, angular. Tendrils long, 2-fid. Leaves petioled, cordate, quite entire, 5-7-nerved at base, membra- nous, mucronate. Flowers small ; the males on long pedicels, in subum- bellate fascicles. — This very remarkable plant is named in honour of W. T. Gerrard, Esq., who collected it in May, 1862, on the Nonoti, and to whom I am indebted for specimens of male and female flowers and ripe seeds. For several years Mr. Gerrard* has devoted himself, with an ardour which repeated attacks of fever and repeated thwartings from the unruly conduct and superstition of the natives have not abated, to the exploration of the natural history of Zululand ; and the pages of ‘ Flora Capensis ’ and of * Thesaurus Capensis,’ so far as published, bear ample witness to his success in botanical discovery. He has also, in conjunction with Mr. M'Ken, most generously contributed nearly 2000 species of dried plants to the Dublin University Herbarium. I have, therefore, peculiar pleasure in inscribing this genus to his honour. As a genus it will stand next to Alsomitra , Wall., but it differs in habit and foliage, as well as in several characters of flowers and fruit. My friend Professor Decaisne, who has succeeded in raising young plants at the Jardin des Plantes, joins me in the specific name se- lected. The seedlings, after forming a few leaves, have already acquired tubers as large as a pigeon’s egg. Oedee LX. BEGONIACEiE. Flowers monoecious. — Male : Perianth 4-leaved, coloured. Stamens many ; anthers adnate, with a thickened connective. — Female : Perianth with a 3-winged tube, adnate to the ovary, and a 4-9-parted, coloured limb. Ovary inferior, 3-celled, with many axile ovules ; stigmas 3, subsessile, fleshy. Cap- sules 3-winged, membranous, opening by slits ; seeds nume- rous, without albumen. — Herbs, with swollen nodes, juicy stems and unequal-sided leaves, with membranous stipules. 1. BEGONIA, Linn. Capsules opening by arched or longitudinal slits along the face of the cells. Placentas from the inner angle of the ovarian cells. — FI. Gap. ii. p. 480. A large genus, chiefly American and Asiatic. Leaves alternate, petioled, palmate-nerved, subentire, toothed or lobed, unequal-sided, frequently semicordate at base, often scaly and brightly coloured on the lower surface. Flowers cymose, mostly pink or rosy. — About 5 South African species, all Eastern and from Natal. Order LXI. CACTE.E. Flowers bisexual. Perianth many-leaved, imbricating, its tube adnate to the ovary. Stamens indefinite. Ovary inferior, * Since the above was written, Mr. Gerrard left Natal upon a scientific expedition into the interior of Madagascar, where he died of fever in 1866. —J. I). 11. LXI. CACTEAI. 129 1-celled, with several parietal placentas ; style filiform, termi- nal ; stigmas as many as the placentas. Fruit succulent. Seeds mostly without albumen. — Succulent, mostly leafless plants, almost all of American origin. 1. KHIPSALIS, Graertn. Tube of the perianth not produced beyond the ovary ; limb rotate, of 12-18 short, scale-like lobes, the outer resembling sepals, the inner petaloid. Stamens many, about equalling the lobes of perianth. Style filiform ; stigmas 3-6, radiating. Berry globose, smooth, often crowned with the dried-up limb of the perianth. Seeds numerous, exalbuminous. — FI. Cap. ii. p. 479. Slender succulents, growing on trees, with whip-like (or leaf-like, often jointed) stems, and small, lateral flowers. — R. cassytha, or some closely-allied species, grows in Caflraria and at Natal ; branches slender, green, terete, pendulous, remotely scaly. Order LXII. EICOIDE^l. Flowers regular, bisexual. Calyx gamosepalous, 4-5-8-cleft, adnate or free, mostly coloured within. Petals either very numerous or, more usually, 0. Stamens perigynous or sub- hypogynous, definite or indefinite, free. Ovary inferior or superior, 2-5-20-celled or of 2-5 separable cocci; ovules many or few or solitary, usually on long cords, attached to the base or inner angle of the cavity ; stigmas as many as the carpels. Fruit capsular or nut-like. Embryo curved round a central, mealy albumen. — Herbaceous or suflruticose plants, with more or less fleshy, mostly quite entire leaves, with or without stipules. Flowers either showy or minute. Tribe 1. Mesembeyanthe.®. Petals very numerous, narrow. Stamens many. Ovary inferior, 5- or many-celled. Capsules 5-20-celled, opening by as many valves. — Leaves sessile, mostly opposite, without stipules. Herbs or shrubs, with fleshy leaves and bright flowers 1. Mesembeyanthemum. Tribe 2. Tetbagoniea:. Petals 0. Ovary inferior. Drupe 1-9-celled, indehiscent, often winged. — No stipules. Herbs or shrubs, with flat, alternate leaves . 2. Teteagonia. Tribe 3. Aizoidea:. Petals 0. Stamens definite or indefinite, mostly perigynous. Ovary superior. Capsules loculicidal. — No stipules. Stamens perigynous. Stamens about 20, in the base of the calyx- tube. Ovary 5-celled ; ovules 2-many in each cell 3. Aizoon. K 130 LXII. FICOIDEtE. Stamens 10-40, in parcels on the top of the funnel-shaped calyx-tube. Ovary 2-celled ; cells 1-ovuled 4. Acrosanthes. Stamens 8-10, alternating in pairs with the lobes of the rotate calyx. Ovary 2-5- celled ; cells 1-ovuled 5. Galenia. Stamens hypogynous. Stamens 5. Ovary 3-celled ; ovules solitary 6. Plinthus. Tribe 4. Molltjgine.®. Petals mostly 0. Stamens 5, 10 or many, mostly hypogynous. Ovary superior. Capsule loculicidal. — Leaves usually stipulate. Ovary 3-5-celled ; each cell with several ovules. Calyx 5-parted nearly to the base. Sepals unequal, cuspidate. Petals 15-20, shorter than the calyx. Seeds arilled at base 7. Orygia. Sepals subequal, ovate. Seeds not arilled. Stigmas linear. Stipules obsolete . . 8. Mollugo. Stigmas obovate, fleshy. Stipules sca- rious, lacerate, conspicuous .... 9. Pharnaceum:. Calyx 5-fid to the middle or above it . . . 10. C(ELANTHUM. Ovary 1-2-3- or 5-celled; cells 1-ovuled. Calyx 5-parted, segments quite entire. Ovary 3-5-celled 11. Psammatropha. Ovary 1-celled 12. Adenogramma. Calyx 4-parted, segments fimbriato-lacerate 13. Polpoda. Tribe 5. Giesekieze. Petals 5 or 0. Stamens 5 or many, hypogynous. Ovary superior. Fruit of 2-5 separable nuts. Petals 0. Ovary of 3-5 carpels. Nuts warted or crested 14. Giesekia. Petals 3-5 (rarely wanting). Ovary of 2 plano- convex carpels. Carpels flat, with a marginal wing . . . .15. Semonvillea. Carpels hemispherical, w'ingless, dorsally pitted 16. Limeum. Tribe 6. Petals 0. Stamens 5 or many, mostly perigynous. Ovary superior, 1-5-celled ; cells 1- or many-ovuled. Capsule circumscissile. Stamens perigynous ; ovary 2 -3 -celled. Flowers sessile, axillary 17. Trianthema. 1. MESEMBRYANTHEMUM, Linn. Calyx 5-, rarely 2-8-lobed, its tube attached to tbe ovary, tbe lobes unequal, usually leaf-like. Petals very numerous, linear, in one or many rows, united at the base. Stamens in- numerable, in many rows, united at base. Ovary 4-20, but usually 5-celled ; stigmas 4-20, usually 5. Capsules 5-20- celled, opening like a star at the summit. Seeds very many. — FI. Gap. ii. p. 387. A vast genus, chiefly South African, natives of dry sands, Karroo plains, etc. ; few to the east of the Great Fish river. Leaves mostly opposite, LXII. FICOIDEiE. 131 thick and fleshy. Flowers mostly terminal, white red or yellow, some dazzlingly brilliant, opening mostly in bright sunshine. — About 300 species. 2. TETRAGONIA, Linn. Calyx 4-cleft, rarely 3-5-cleft, its tube adhering to the 4-5- angled ovary ; lobes coloured within. Petals wanting. Stamens variable in number, sometimes definite, sometimes fascicled ; anthers oblong or linear. Ovary inferior, 3-9-celled, by abor- tion 1-2-celled ; ovules solitary ; styles as many as the ovarian cells, very short. Drupe or nut winged or horned, indehiscent, 1- 9-celled. Seeds solitary. — FI. Cap. ii. p. 460. • Herbs or undershrubs, with alternate, flat, fleshy, undivided, usually quite entire leaves, and axillary, sessile or stalked, green flowers. — 23 Cape species, dispersed. 3. AIZOON, Linn. Calyx 5-parted, coloured within. Petals 0. Stamens about 20, in the bottom of the calyx, and disposed in 3-5 tufts. Ovary free, 5-angled, 5-celled ; cells with 2 or many ovules ; styles or stigmas 5. Capsules 5-celled, opening at the top, like a star ; cells 2-10-seeded. — FI. Cap. ii. p. 469. Small herbs or undershrubs. Leaves opposite or alternate, fleshy, quite entire. Flowers in the axils of the leaves or forks of the stem, mostly sessile. — 7 species, dispersed. 4. ACROSANTHES, E. and Z. Calyx 5-parted, coloured within. Petals 0. Stamens 10- 40, 2-seriate, in parcels on the top of the funnel-shaped calyx- tube, the outer longer ; filaments capillary ; anthers linear. Ovary free, 2-celled ; ovules solitary ; stigmas 2, filiform. Cap- sules subglobose, enclosed in the persistent calyx, 1-celled, 2- valved. Seeds 1-2, fixed to the base. — FI. Cap. ii. p. 473. Decumbent, forked, glabrous undershrubs. Leaves subconnate, opposite or in fours. Flowers axillary or in the forks, solitary, peduncled. — 4 species, all Western. 5. GALENIA, Linn. Calyx 4- or 5-parted, coloured within. Petals 0. Stamens 8-10, in 4 or 5 sets, or 1 or a pair alternating with the calyx- lobes ; anthers versatile, didymous ; cells turgid. Ovary free, 2- 5-celled, or by abortion 1-celled ; cells 1-ovulate ; styles 2-5. Capsules 2-5-celled, or by abortion 1-celled, 3-5-fur- rowed or 2-edged, dehiscent, or, if unilocular, indehiscent. Seeds solitary, shining, striate, fixed to long cords rising from the base of the cell.— FI. Cap. ii. p. 473. Herbaceous or half-shrubby, much-branched plants, with alternate or opposite, fleshy, quite entire leaves, and minute, axillary, mostly sessile flowers. — 18 species, dispersed. K 2 132 LXTI. FICOIDE^. 6. PLINTHUS, Fenzl. Calyx tubular ; limb 5-parted, lobes erect, subequal, co- loured witbin. Petals 0. Stamens 5, inserted in tbe base of tbe calyx, alternate with the lobes, hypogynous ; filaments exserted. Ovary 3-celled ; ovules solitary, pendulous ; style 3-parted. Capsules ovoid, densely papillose, not depressed, 3-celled, loculicidal, 3-valved ; cells 1-seeded. Seeds shining, striate. — FI. Cap. ii . p. 479. JP. crypto carpus, Fenzl, found by Drege in the Nieuweveld, is a small, prostrate sh?ub. Leaves very minute, ovate-trigonous, opposite and alter- nate. Flowers hidden in the axils of the leaves, minute. 7. OBYGIA, Forsk. Calyx 5-parted; segments cuspidate, of unequal length. Petals 15-30, spathulate-linear or oval, shorter than the calyx, at the base confluent into a fleshy cup. Stamens 12-40, in the bottom of the calyx, some free, some united at base ; filaments subulate-triangular ; anthers versatile, oblong. Ovary globose, 5-celled, many-ovuled ; stigmas 5. Capsules dry, roundish, 5-angled, 5-furrowed, 5-celled, loculicidally 5-valved. Seeds many, on ascending cords, reniform, black, concentrically furrowed, arilled at the scar. — FI. Cap. i. p. 136. O. decumbens, the only species, is a glabrous, glaucous, diffuse plant, with angular stem and branches ; fleshy, orbicular obovate or elliptical, entire or retuse, muticous or mucronate, alternate, petioled leaves, and cymose flowers. It grows in the Eastern and North-Eastern districts, and is also found in Arabia and the East Indies. 8. MOLLUGO, Linn. Calyx 5-parted ; segments ovate. Petals 0 or numerous, shorter than the calyx, very narrow, 2-3-forked or setaceous. Stamens 3-5-20, in the base of the calyx, hypogynous, free or united in parcels ; filaments subulate ; anthers oblong or glo- bose. Ovary ovate, 3-5-celled, many-ovuled; stigmas 3-5, linear. Capsules membranous, 3-5-angled, loculicidally 3-5- celled. Seeds numerous, on ascending cords, reniform, smooth or tubercled. — G-linus, FI. Cap. i. p. 136 ; and Mollugo, FI. Cap. i. p. 137. Annual or perennial, procumbent, glabrous or stellately-liairy plants, with alternate or whorled, entire or denticulate leaves, and small umbellate or clustered flowers. — 3 Cape species, dispersed. 9. PHAENACEUM, Linn. Calyx 5-parted ; sepals very obtuse, coloured (white) within and at the margin. Petals 0. Stamens 3-5 or 12-16 in two rows, the 5 outer alternating with the sepals; or 20-30, in LXII. 1'ICOlDEJi. 133 3-5 parcels, connate at base, alternating with a few free sta- mens. Disk 3-5-fid or 0. Stigmas 3-5, obovate, fleshy, coloured or white, rarely terete. Capsules membranous, 3-5- angled, 3-5-celled, 3-5-valved, loculicidal ; cells many-seeded. Seeds lenticular, smooth, shining. — Pharnaceum, FI. Cap. i. p. 138 ; and Hypertelis, FI. Cap. i. p. 144. Small, slender, half-shrubby plants or herbs, with setaceous, filiform or linear, rarely lanceolate leaves ; the cauline leaves alternate, the upper crowded in dense, brush-like tufts. Stipules conspicuous, either fimbriate or entire. Flowers in cymes or pseudo-umbels. — About 20 species, dis- persed. 10. CCELANTHUM, E. Mey. Calyx funnel-shaped or bell-shaped, semi-5-fid; the segments petaloid, obtuse. Petals 0. Stamens 5, inserted between the calyx-lobes, perigynous ; filaments short ; anthers sagittate, erect, longer than the filaments. Disk 0. Stigmas 3, round- ish, fleshy. Capsules oblong, 3-angled, 3-celled, many-seeded, loculicidally 3-valved ; valves septiferous. Seeds globose, subcompressed. — FI. Cap. i. p. 14