G74L24G 1896 Mov. 10 1976 ye, io aE ond A Oe [Read before the Massachusetts Horticultural Society, March 21, 1896.] BY PROFESSOR F. LAMSON-SCRIBNER, B.S., CHIEF OF DIVISION OF AGROSTOLOGY, UNITED STATES DEPARTMENT OF AGRICULTURE, WASHINGTON, D.C. echo ae a le Sinss 4 * BOSTON: " Press oF ROCKWELL AND CHURCHILL. 1896. V — cP OO Vt 5 ie bee ee > ee ee ee ee _ kale GiRVAGS.SiE Ss; Coming to Boston to address you on the subject of grasses, is like carrying coals to Newcastle, for is not this the home of the widely known and justly celebrated author of Flint’s “ Grasses and Forage Plants”? But the subject is a broad one —as broad as the world is wide, and as varied as it is broad; so broad, and ' embracing so many diverse lines of investigation, all alike inter- esting, that I have found it exceedingly difficult to determine what subjects to take up, or where to draw my limitations. Good things will stand repeating, and in a multitude of counsellors there is safety. New ideas are rare indeed, but in the application of old ones may spring a happy thought of use to some one, and there always exists this possibility to encourage the speaker. “The grass faileth; there is no green thing,” is an apt expres- sion of the extremity of desolation. Where there is no grass, there are the absolute deserts. Where our best grasses abound, and where they receive the most attention, there we find our high- est civilization and greatest prosperity. Destroy the rich verdure of our pastures and meadows, and how much of the pleasure as well as the profit of the farmer’s life would be blasted. Destroy the little grass plat of the contracted yard of the citizen, and how much would the enjoyment of domestic life be narrowed. , Grasses may be considered the plebeians among the families of the vegetable kingdom. They are ubiquitous, and in all temperate regions innumerable. In their number of species they constitute one-fourth of the flowering plants of the arctic zone, \ one-twelfth of those of the temperate region, and from one- ee twelfth to one twenty-fifth of those of the tropics. In the countless myriads of individuals, particularly in the temperate zone, grasses far surpass all other orders of plants. They form the rank and file of the army of plants; but here, as in armies of : ; AWW) Lidcpaidl pone SS 4 men, it is the rank and file that does the real service. There are, however, lordly members among the grass family, for some of the tropical species vie with the tallest trees in height. The immense bamboo forests of India are forests of grasses, and to the dwellers of those regions they are as useful as are our own forests to us. They furnish material for the construction of their houses and household furniture and domestic utensils, articles of ornament, and sometimes even articles of clothing. Some of the bamboos furnish drink to the thirsty traveler ; others occasionally supply food ; and several times within historical periods have the fruits of these bamboos saved hundreds of thousands, if not millions, of people from actual starvation. So generally useful are the bamboos that their products have entered into the commerce of the world. A mere enumeration of the diverse uses of grasses would more than occupy the time which has been allotted to me here; but at the risk of wearying you, I must briefly outline the more impor- tant of these. Grains, the product of the cereal grasses, form the staple food of more than four-fifths of the human race. Wheat is a grass, and the world’s production of wheat is estimated at two billions four hundred millions of bushels. Rice is a grass, and the production of this cereal in the East exceeds one million tons, and feeds one-third of mankind. Indian corn, that king of grasses and peculiar product of America, is one of our greatest sources of income. Its cultivation now extends over ninety degrees of latitude and has been carried to all parts of the world. Oats, the most nutritious of all grain foods, barley, and rye are members of the grass family; and aside from these grains, there are a number of grasses which furnish human food, particularly to the natives of Southern Asia and the wild tribes of Africa, the value of whose product cannot be estimated. In addition to the direct usefulness of these grain-bearing grasses to man, several are used very largely to supplement the forage of our domestic animals. They have a further use also, in the production of alcoholic drinks. Nearly half of our sugar supply is derived from grasses. The world’s production of cane sugar is about three million tons. What is said here of the products of the cereal-grasses indicates only in a slight degree the great use- fulness to man of a very few members of the grass family. A larger number are scarcely less useful, although indirectly. Or These are the grasses of our meadows and pastures, which furnish us our beef and mutton, our butter and milk; which feed our horses, the noblest of our domestic animals, and still among the most useful, in spite of electric railroads, horseless carriages, and bicycles. The money value of the hay crop of the United States for the year 1894 was estimated at nearly half a billion dollars, and the value of the grasses in pastures of the Northern States may be considered equal in value to the hay produced, while in the Southern States and the great grazing regions of the West, the value of the pasturage far exceeds that of the hay crop. A conservative estimate, therefore, of the annual value of the grasses of the meadows and pastures of this country alone exceeds a billion dollars. ‘‘Before dismissing this subject of the utility of grasses in furnishing food for man and the animals he has domesticated, we shall do well to pause for a little reflection upon its relation to the industry, commerce, and wealth of nations, as well as to man’s subsistence —our dependence not only upon the cereal grasses for our staple vegetable food, but indirectly upon the forage grasses for our supplies of animal food, namely beef, mutton, venison, and dairy produce, as well as for various animal substances such as wool and hair, hides and skins, bone and horn, oil and tallow, used for textile and other manufactures (notably woolen fabrics and leather), or for domestic purposes — the large proportion of the world’s inhabitants engaged in agricult- ural (chiefly cereal cultivation) and pastoral pursuits, in some countries from seventy to ninety per cent of the adult male pop- ulation — the vast internal and foreign trade connected with the distribution of agricultural products by land and sea — the numerous and important industries concerned in operating upon one or other form of this produce in order to prepare it for con- sumption ; and lastly, the enormous capital employed in all these industrial activities, and the consequent accumulation of wealth. It is only when we take a comprehensive survey, such as here indicated, that we are able to form some conception of the tran- scendent importance of the Graminee.’’' There are a number of minor uses to which many species of grasses have been applied; probably the most important is the material they furnish for paper-making. Several of our native 1 William Hutchinson in ‘‘ Handbook of Grarses,’’ 1895. — species furnish a good fibre for this purpose, but the grass which has been used most largely in. the manufacture of paper is the Esparto grass of the Mediterranean region. The quantity of this grass annually imported into England at present amounts to over two hundred thousand tons, valued at three-quarters of a million pounds sterling. Some grasses are used in the manufact- ure of cordage, or hats, or of matting; others make thatch ; some are employed in medicine ; others yield perfumery. Among the natural uses of grasses may be mentioned that of binding - drifting sands and the protection of our coasts and river banks from the action of the tides or floods, and their use in protecting the soils of our fields and meadows by the covering which their turf affords. They extract from the earth and the air elements which they transform into substances that serve as food, and in doing this they help to purify the air we breathe. Contrary to the general idea, there exists among grasses a remarkable diversity of form. So varied is this that botanists have already defined nearly four thousand distinet species. This diversity appears throughout all the organs of the grass. In some the roots are simply fibrous, and the plants grow in tufts or bunches, as Sheep’s Fescue and Orchard grass; others have what we call creeping roots, and it 1s among these that we should look for the best turf-forming species. Some have stems less than an inch in height and appear like mosses covering the soil and rocks; others attain the height of our tallest forest trees. Some have leaves as fine as the finest thread; in others the leaves are those of the ideal blade of grass, while others again have leaves ike those of palms, or leaves as short and as broad and as round as those of the well-known smilax. To explain the details and the varieties existing among the flowers of grasses would be wearisome. That grasses have flowers is an idea rarely entertained by any except botanists, and I have frequently heard the remark, “I did not know that grasses had flowers.” They do, however, although their special characters may differ from those of other plants; and provision exists here, as it does throughout nearly all the tribes of vegetation which bear flowers, for securing cross-fertilization. The flowers of grasses are inconspicuous and secrete no nectar. They are not, therefore, attractive to insects, which play so important a part in the process of cross-fertilization. The pollen of grass flowers is ~I dry, light, and powdery, and easily blown about by the wind, and cross-fertilization among grasses is effected by this agency. If the variety in the external form of grasses is wonderfully great, their internal structure is scarcely less so, and the histo- logical studies of grass stems, leaves, and fruits are exceedingly interesting. Intricate problems in mechanics are exhibited in the structure of the slender cylinders which constitute the grass stem, and which, in many cases, possess a strength most surprising. The amount of mechanical tissue entering into the structure of the stem which holds a heavy head of wheat is insignificant, but the disposition of the various elements of this tissue gives it the strength necessary to perform its proper functions. Again, the leaves of grasses, which many think so much alike that they would consider the expression “as like as two blades of grass” as forcible as the more common phrase “as like as two peas,” exhibit a di- verse and marvelous interior structure. No more delicate trac- ings or beautiful designs of lace work can be imagined than are presented by these same grass leaves when viewed under the microscope. The designer might well study these tissues, for in them he would find many new figures and combinations of lines, the beauty of which he could not hope to excel, and the repro- duction of which could not fail to receive the highest admiration. In their internal structure as in their outward contour, the leaves of grasses present such definite characters that these may be used to distinguish species. A minute transverse section of the leaf of Kentucky Blue grass, which one could barely see with the naked eye, would, under the microscope, present characters which at once distinguish it from all other grasses. They are totally different from those exhibited by a leaf of Orchard grass, and these again are wholly unlike those of Meadow Fescue. There are certain cells in the leaf tissue, running from the base to the summit, which are larger and have thinner walls than the surrounding cells. These special cells readily absorb or give up moisture, and because of this property they exercise the mechani- eal function exhibited in the expansion or opening out of the leaves, or their contraction and rolling together. The provisions which nature has made for the distribution of grasses 1S an interesting subject, and one worthy of passing notice. In many cases the seeds are covered with delicate chaff- like scales, or are furnished with winged or feathery appendages, ~~. ae — a