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West, Newman imp.
M.Smith del,
J.N.Fitch lith.
CLEMATIS PARVIFLORA, A.Cunn.
PuateE 2.—CLEMATIS PARVIFLORA.
Famity RANUNCULACE.] [Genus CLEMATIS, Linn.
Clematis parviflora, A. Cunn. Precur. n. 636; Hook. f. Fl. Nov. Zel. 1, 7; Cheesem. Man. N.Z.
Fi. 4.
Clematis parviflora is one of the many species first collected by the talented
and enthusiastic Allan Cunningham, who in 1826 and again in 1838 explored a
large part of the district lying between the Bay of Islands, Hokianga, and
Whangaroa. In those days botanical exploration was a very different matter from
what it is at the present time, when no part of the Dominion is far removed from
settlement, roads, or even railways. The journeys made by the two Cunninghams,
Bidwill, Dieffenbach, Colenso, and others involved great hardships, much fatigue,
and no small amount of danger; and no estimate of the labours of these early
pioneers is complete which does not take the circumstances of the times into
consideration.
Cunningham gathered C. parviflora at Whangaroa in 1826, finding it in
“thickets on the skirts of forests,” and published it in his ‘ Precursor,” issued
during the years 1836-39. Soon afterwards it was collected in various localities
in the northern portion of the North Island by Colenso, Bidwill, and others. Since
then the typical state of the species has been found to range from the Three Kings
Islands and the North Cape to the East Cape and the northern portion of Hawke’s
Bay, but is seldom present in much quantity, and is often decidedly local. It is
perhaps more common on the Little Barrier Island than in any other locality known
to me. I am not aware that the typical form has been found in the South Island ;
but a variety with small leaves and caudate sepals was discovered at Nelson by
Mr. W. T. L. Travers many years ago, but has not since been seen. Another variety
with small trilobed leaflets, which also occurs in the North Island, has been
collected at Okarita by Mr. A. Hamilton.
C. parviflora can always be distinguished from C. fatida, which is its nearest
ally, by the small size, slender habit, smaller submembranous leaflets, narrower
silky sepals, and particularly by the broad oblong anthers, which have a minute
rounded knob at the tip of the connective. I am indebted to Mr. Hemsley for
pointing out to me that my specimens from the Little Barrier Island and
Cunningham’s type from Whangaroa both show three sorts of individuals—males,
females, and hermaphrodites. But in the last many of the anthers and carpels
appear to be sterile.
Puate 2. Clematis parviflora. The portions marked A drawn from specimens collected on the
Little Barrier Island by Miss Shakespear; that marked B from Cunningham’s type preserved in the
Kew Herbarium. Fig. 1, stamen (x6); 2. carpel from flower (x6); 3, ripe carpel—all from
Little Barrier specimens; 4 and 5, stamens from hermaphrodite flowers, Cunningham’s type
specimen.
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M.
J.
West, Newman imp.
RANUNCULUS LYALULII, Fook. f
RANUNCULUS LYALLII.
Famity RANUNCULACEZ.] [Genus RANUNCULUS, Linn.
PLATE 3.
Ranunculus Lyallii, Hook. f. Handb. N.Z. Fl. 4: Cheesem. Man. N.Z. Fl. 9.
Most travellers in the Southern Alps are acquainted with this magnificent plant,
which Sir J. D. Hooker has well styled “the monarch of the genus.” The large
deep-green peltate or saucer-shaped leaves, sometimes 12in. to 16in. in diameter,
and the tall paniculately branched flowering-stems, often bearing 15-20 large white
flowers from Lin. to 3 in. in diameter, are quite sufficient to arrest the attention of the
most casual visitor. As the name indicates, it was first discovered by Dr. Lyall,
the surgeon attached to H.M.S. “ Acheron,” which, under the command of Captain
Stokes, was employed during the years 1847-51 in a survey of the southern shores
of New Zealand. Lyall’s specimens were obtained in Milford Sound, and consisted
of leaves only. These gave no clue to the affinities of the plant, for in those days
the existence of Ranunculi with peltate leaves was not even suspected. Hence
in the “ Flora Nove Zelandie ” it was simply alluded to as “a very remarkable
possibly umbelliferous plant.” This suggestion, which proved to be wide of the
mark, was no doubt due to the similarity of the leaves to those of a gigantic
Hydrocotyle. However, in the beginning of 1861 flowering specimens were obtained
by Sir Julius von Haast and Dr. Sinclair in the Rangitata Valley, and the
systematic position of the plant definitely ascertained. Since then it has been
found to be abundant along the whole chain of the Southern Alps, from the
Spenser Mountains in the Nelson Provincial District southwards to Mount Anglem
in Stewart Island. .
Ranunculus Lyallii is usually found by the margins of mountain-streams, or
in moist hollows sheltered by subalpine scrub. Occasionally it can be seen covering
considerable areas, which in the flowering season are whitened with the abundance
of its flowers. It is most plentiful at an altitude of about 3,000 ft., but in several
places I have seen it at an elevation of nearly 5,500 ft., and it descends as low as
1,500 it. in a few localities on the western side of the Alps. In size and habit of
growth it much resembles the well-known garden plant Anemone japonica ; but it
has much bolder foliage, and more striking flowers. It is much to be regretted
that it is not better known in gardens, for although it has the reputation of being
impatient of cultivation, a knowledge of the conditions favouring its growth, with
careful management with regard to soil, situation, and supply of moisture, would
probably bring about much more favourable results.
In the “ Handbook” Sir J. D. Hooker has described an allied plant under the
name of Rk. Traversii, the distinguishing characters of which are the more deeply
crenate leaves with two incisions near the base, and the cream-coloured flowers.
No specimens have been obtained of late years, but the examination of a careful
drawing prepared from the type specimen and kindly forwarded to me by the
Director of Kew leaves no doubt in my mind that it is nothing more than a trivial
variety of R. Lyallii. Apart from this form R. Lyallii cannot be said to have any
near allies, for although R. Matthewsii and R. Buchanani have very similar white
flowers, they are much smaller in size, and the leaves are deeply lobed or even
ternately divided. Outside New Zealand the only species known with peltate
leaves are R. Cooperi, Oliv., and R. Baurii, McOwan, both natives of South Africa.
But both of these are much smaller than R. Lyallii, and have yellow flowers with
narrower petals. White-flowered Ranunculi are decidedly rare, except in the
section Batrachium, which is not represented in New Zealand; and the existence
of three such conspicuous plants as R. Lyallii, R. Matthewsii, and R. Buchanani
constitutes a remarkable feature of the New Zealand flora.
Puiate 3. Ranunculus Lyallii, drawn from specimens collected in the Mount Cook district, Canter-
bury; alt. 3,000 ft. Fig. 1, base of petal, showing nectary ; 2 and 3, stamens (x 6); 4, pistil (x 6);
5, fruit, with some of the carpels removed (x2); 6, ripe carpel (enlarged) ; 7, section of carpel
(enlarged) ; 8, embryo (enlarged).
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RANUNCULUS NIVICOLA, Zook.
PLATE 4.—RANUNCULUS NIVICOLA.
Famity RANUNCULACE. | [Genus RANUNCULUS, Linn.
Ranunculus nivicola, Hook. Ic. Plant. t. 571,572; Cheesem. Man. N.Z. Fl. 12.
This graceful buttercup is nowhere more plentiful than on Mount Egmont,
where it was discovered, in the summer of 1839, by Dr. Ernst Dieffenbach, the
naturalist to the New Zealand Company, who was the first European to ascend
the mountain. About the same time it was collected by Mr. Bidwill on Tongariro
and Ruapehu, where, however, it does not attain the same luxuriance as on Egmont.
Its abundance on the last-mentioned mountain greatly impressed me on my first
ascent, made nearly thirty years ago. After passing through the forest region
which girds the lower slopes of the mountain up to an elevation of nearly 4,000 ft., a
subalpine scrub is reached, chiefly composed of Olearia, Senecio, Coprosma, Veronica,
&c. In open sheltered places within this scrub R. nivicola flourishes in perfection,
attaining a height of 3 ft., with radical leaves often more than 6 in. across, and with
a branched flowering-stem bearing 8-15 large bright-yellow flowers 14 in. in diameter.
Higher up, on the open mountain-side, it is smaller in size, and is usually found
in the shelter of rocks, or on the sides of ravines. According to my own observations,
its upward hmit is about 5,500 ft. or slightly more. Dieffenbach states that he
collected it at an altitude of “7,000 ft., near the perpetual snow,” but this I believe
to be altogether erroneous. Mr. Buchanan, who examined the vegetation of
Egmont in 1867, gives 5,000 ft. as its limit. In 1907 I noticed it up to 5,800 ft.
on the western side of Tongariro, and many years previously Mr. Kerry Nicholls
reported to me that it occurred at an equivalent height on Ruapehu. Both on
Egmont and Ruapehu all vegetation ceases before an elevation of 7,000 ft. is
reached.
The chief home of R. nivicola is in the two localities mentioned above. TI have,
- however, seen specimens gathered by Mr. Townson on Mount Holdsworth, one of
the chief peaks of the Tararua Range, and I can entertain little doubt that it also
exists on the Kaimanawa and Ruahine Mountains. Some flowerless specimens
collected by Mr. J. H. Macmahon on Mount Stokes, Marlborough, apparently belong
to the same species, but an element of doubt will remain until the flowers have
been obtained.
The nearest ally of R. nivicola is undoubtedly R. geraniifolius. But that
species differs in its smaller size and much more slender habit, fewer leaves which
are often much divided, in the fewer and smaller flowers, and in the petals not being
conspicuously notched at the apex. I am not acquainted with any extra-New-
Zealand species which can be said to be closely allied. R. anemoneus, of the
Australian Alps, with which Bentham compared it, differs in the more finely cut
radical leaves, in the broader sessile cauline leaves, and in the white flowers.
Probably its nearest relatives will be found in some of the South American species.
Prats 4. Ranunculus nivicola, from specimens gathered on Mount Egmont, at an altitude of
4,000 ft. Fig. 1, base of petal, showing nectary (x 4); 2, stamen (x 4); 3, carpels from flower (x 4) ;
4, ripe carpel (enlarged).
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RANUNCULUS ENYSII, 7 kirk.
PLATE 5.—RANUNCULUS ENYSII.
Famity RANUNCULACE.] [Genus RANUNCULUS, Linn.
Ranunculus Enysii, 7. Kirk in Trans. N.Z. Inst. xii (1880), 394; Cheesem. Man. N.Z. Fl. 13.
Ranunculus Enysii is one of the most distinct species found in New Zealand,
and although highly variable in many of its characters can never be confounded
with any other member of the genus. It was first found, about the year 1878, by
Mr. J. D. Enys, at that time resident at Castle Hill, in the Canterbury Alps. It
fitly commemorates the services rendered to natural science in New Zealand by
its discoverer, who for many years rendered invaluable assistance to every scientific
explorer of the mountains of central Canterbury.
The range of R. Enysii is not precisely known, but it has been recorded from
many localities in the Canterbury Alps, and is particularly abundant in the upper
portion of the Waimakariri River basin, from Mount Torlesse to Bealey. In this
district it is usually found in moist sheltered places along the sides of streams or
ravines. I have not seen it at a lower elevation than 1,800 ft., and apparently
it does not ascend higher than 4,000 ft., above which its place is taken by the more
alpine Rk. Sinclairii. Outside Canterbury the only localities that I know for it are
the Taieri Hills, in eastern Otago, from whence specimens were described by
Mr. J. Buchanan under the name of R. tenuis (Trans. N.Z. Inst. xx (1888), 255,
t. 12), but which clearly represent a form of R. Enysii with more pinnately divided
leaves than usual; and Lake Harris, in western Otago, where it was gathered by
the late Mr. T. Kirk.
The affinities of R. Enysii are obscure. Mr. Kirk compared it with
R. lappaceus, and no doubt there is a superficial resemblance to highly developed
forms of that species; but it differs altogether in the glabrous habit, more deeply
divided leaves, and particularly in the turgid achene, which is never compressed
or margined, as in that species. A relationship to R. geraniifolius has also been
suggested, but the resemblances are by no means convincing.
Pate 5. Ranunculus Enysii, drawn from specimens collected by the Broken River, Canterbury
Alps, at an altitude of 2,500 ft. Fig. 1, petal, showing nectary (x 3); 2, stamen (x8); 3, carpel
from flower (x 8); 4, ripe carpel (x 8).
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M.Smith del. imp.
TNFitch lth, West, Newman imp
RANUNCULUS SERICOPHYLLUS, ook f.
PLATE 6.—RANUNCULUS SERICOPHYLLUS.
Famity RANUNCULACE4.] [Genus RANUNCULUS, Linn.
Ranunculus sericophyllus, Hook. f. Handb. N.Z. Fl. 6; Cheesem. Man. N.Z. Fl. 17.
This beautiful little plant was originally discovered by the late Sir Julius von
Haast during his adventurous explorations in the Southern Alps. His _ first
specimens were obtained in May, 1862, near the source of the Hopkins River, which,
alter uniting with the Dobson, flows into the head of Lake Ohau. In the following
year he also collected it on Mount Brewster, near the head of Lake Wanaka. Since
then it has been observed in many localities in the central portions of the Southern
Alps, from the head of the Waimakariri River southwards to the divide between
Lakes Wakatipu and Te Anau on the one side and the Sounds of south-western
Otago on the other. It is everywhere a high alpine plant, and is seldom seen below
4,000 It. altitude. In the Mount Cook district, where I have repeatedly gathered
it, it is most plentiful in sheltered hollows in which the drainage from melting snow-
fields keeps the soil perpetually moist. In such places it may be seen forming patches
several yards in diameter, and from its silky pale-green leaves and abundant bright-
yellow flowers presents a very charming and attractive appearance. It ascends
to an altitude of over 6,500 ft., and can frequently be observed blooming within
a few feet of permanent snowfields. In fact, it is often the last conspicuous plant
seen by the alpine climber when ascending the higher mountains of the Southern
Alps.
As a species R. sericophyllus is remarkably distinct from all the other New
Zealand forms. Sir J. D. Hooker has compared it with the Tasmanian and Victorian
R. Gunnianus ; but, as remarked by him, it is much more robust and silky, with
larger flowers, and more membranous leaves, the ultimate segments of which are
smaller and shorter.
PiatE 6. Ranunculus sericophyllus, drawn trom specimens obtained in the Mount Cook district,
Canterbury ; alt. 5,500 ft. Fig. 1, petal, showing the three nectariferous glands (x 2); 2 and 3,
stamens, front and back view (x 4); 4, carpel from flower (x 4); 5 and 6, ripe carpels (enlarged).
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RANUNCULUS AUCKLANDICUS, A. Gray.
PLaTE 7.—RANUNCULUS AUCKLANDICUS.
Famity RANUNCULACE. ] [Genus RANUNCULUS, Liny.
Ranunculus aucklandicus, A. Gray, Bot. U.S. Expl. Exped. i, 8; Cheesem. Man. N.Z. Fl. 22.
So far this species has only been recorded from the Auckland Islands. For
its discovery we are indebted to the United States Exploring Expedition, which,
as is well known, was engaged in geographical researches in the southern seas during
the years 1838-42. The brig “ Porpoise,” one of the vessels composing the expedi-
tion, paid a visit to the Auckland Islands when returning from a cruise to the
Antarctic regions, anchoring in Port Ross on the 7th March, 1840. A stay of three
days only was made, but during that period Dr. Holmes, the surgeon attached to
the vessel, formed a small collection of plants, among which were specimens of a
species of Ranunculus. After the return of the expedition these were described by
Asa Gray under the name of Ranunculus aucklandicus. Curiously enough, the
plant was not observed by Hooker, in Ross’s expedition, nor by the French
expedition under the command of Admiral D’Urville. Subsequent visitors, however,
have found it by no means uncommon in swampy places, especially near the sea.
I have seen specimens collected by Mr. T. Kirk in 1890, by Dr. Cockayne in 1903,
by Mr. Aston and Mr. J. S. Tennant in 1907, and again by Mr. Aston in 1909. To
the last-mentioned gentleman I am indebted for the specimens from which this
plate has been prepared, representing a somewhat luxuriant form of the species.
The nearest relative of R. aucklandicus is undoubtedly Hooker’s R. subscaposus,
from Campbell Island, which in fact differs only in its smaller size and ovate-
deltoid more acutely lobed leaves, which are usually 3-partite to the base. It is
quite possible that a leisurely examination of the two plants might result in their
union. Dr. Gray, in the ‘* Botany of the United States Exploring Expedition,”
suggests that R. aucklandicus is allied to R. pinguis. But this is clearly erroneous,
as it differs altogether in habit, in the 3-cleft and more deeply lobed much less
succulent leaves, in the appressed strigose pubescence, in the smaller flowers, and
in the altogether different achene, which wants the subulate style winged at the
base so characteristic of R. pinguis. The achene of R. aucklandicus belongs to the
same type as that of R. hirtus, but is less compressed and less evidently margined,
and is slightly villous with long scattered hairs, especially when yourg, and the
style is short and hocked.
Puate 7. Ranunculus aucklandicus, drawn from specimens collected on the Auckland Islands
by Mr. B. C. Aston. Fig. 1, flower (x 3); 2, sepal, outer face (x 5); 3, petal (x5); 4, stamen (x 5) ;
5, carpel from flower ( x 5); 6, ripe carpel (enlarged) ; 7, section of ripe carpel (enlarged) ; 8, embryo
(enlarged).
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DRIMYS TRAVERSIL, Z Hird.
PLATE 8.—DRIMYS TRAVERSII.
Famity MAGNOLIACE.] [Genus DRIMYS, Forsr.
Drimys Traversii, T. Kirk i Trans. N.Z. Inst. xxx (1898), 379; Cheesem. Man. N.Z. Fl. 30.
For the discovery of Drimys Traversii we are indebted to Mr. H. H. Travers,
who, in December, 1882, collected it in the mountains behind Collingwood, north-
west Nelson. Mr. Travers’s specimens, which were in a very imperfect condition,
were communicated to Mr. Buchanan, and were erroneously referred by him to the
genus Hymenanthera (Trans. N.Z. Inst. xv (1883), 339, t. 28). This mistake was
not rectified until 1897, when Mr. J. Dall, well known from his botanical explora-
tions in the Collingwood district, succeeded in obtaining a supply of both flowering
and fruiting examples. These were forwarded to the late Mr. Kirk, and enabled
him to refer the plant to its proper genus. Since then it has been collected by
Mr. W. Townson on Mount Rochfort and Mount Frederic, near Westport ; so that
it probably stretches along the coast ranges from West Wanganui and Karamea
to the mouth of the Buller. Its altitudinal range appears to be from 1,500 ft. to
3,000 ft.
As a species D. Traverswi is remarkably distinct. It is by far the smallest
species of the genus, sometimes barely exceeding 2 ft. in height, and its straggling
or semi-prostrate habit is unique. The small close-set leaves, with their appressed
petioles, and the reddish bark, also distinguish it at a glance from either D. axillaris
or D. colorata.
The distribution of Drimys is most puzzling. Sixteen or seventeen species are
known, of which three are found in New Zealand, four in Australia, one in Lord
Howe Island, four or five in New Caledonia, one or two in New Guinea, one of
which also extends to Borneo and the Philippine Islands. Three species (or more,
according to the different views of authors) occur in South America, stretching
from Mexico to Fuegia. Its distribution is therefore very similar to that of Nertera
and Uncinia. The present centre of the genus is undoubtedly in Australia and
New Caledonia, and an Austral-Malayan origin might have been predicted for it
but for the isolated species found in South America.
Pirate 8. Drimys Traversii, drawn from specimens collected by Mr. W. Townson on Mount
Rochfort, near Westport, at an altitude of 2,500 ft. Fig. 1, portion of under-surface of leaf, showing
indumentum; 2, flower-bud (x5); 3, hexapetalous flower (x5); 4 and 5, stamens, front and back
view (x 8); 6, pentapetalous flower (x5); 7, section of fruit, showing seeds (x 8); 8, fruit, showing
the persistent saucer-shaped calyx (x 8).
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NASTURTIUM LATISILIQUUM, Cheesem.
PLATE 9.—NASTURTIUM LATESILIQUA.
Famity CRUCIFER %.] [Genus NASTURTIUM, R. Br.
Nasturtium latesiliqua, Cheesem. in Trans. N.Z. Inst. xl (1911), 179.
Cardamine latesiliqua, Cheesem. in Trans. N.Z. Inst. xv (1885), 298; Man. N.Z. Fl. 35.
The first specimens of this fine plant were gathered by myself in the summer
of 1881, during a botanical expedition to the Mount Arthur Plateau, Nelson. It
was then abundant on limestone rocks on the northern side of Mount Arthur,
ranging from a height of 3,800 ft. to 5,500 ft., or within a short distance of the
summit of the mountain. Two years later I found it equally plentiful on the
northern slopes of Mount Owen, in the Upper Buller Valley. Since then it has
been gathered by Mr. F. G. Gibbs and others on several of the high peaks in north-
west Nelson, almost always on limestone rocks.
When publishing the species in 1883 I referred it to Cardamine, being mainly
influenced by its evident relationship to Hooker’s C. jastigiata, from which it differs
in the villous leaves, larger flowers, and much broader more turgid pods. But, as
pointed out in the Manual, both C. fastigiata and C. latesiliqua, together with the
allied C. Enysiz, differ from the typical species of Cardamine in the 2-seriate seeds.
Consequently Dr. Schulz, in his elaborate monograph of the genus published in
Engler’s ‘‘ Botanische Jahrbuchern ” for 1903 (vol. xxxii), excludes all three, but
makes no other disposition of them. But although there is no difficulty in justi-
fying the exclusion of the species from Cardamine, it is far from easy to decide
in what other genus they should be placed. Personally, I look upon it as quite
possible that the three species in question, together with the Australian C. radicata
Hook. f., will ultimately form a separate genus. But before taking a step of that
kind a careful comparison should be made with the whole of the genera constituting
the subfamily Arabide, a work which can hardly be properly carried out without
access to the large public herbaria and libraries of Europe. Partly from that reason,
and partly because the characters of Nasturtiwm, although not exactly conformable,
still approach very closely to those of the four species mentioned above, I have
decided to transfer them to that genus for the present. (See some remarks in a
paper of mine published in the ‘ Transactions of the New Zealand Institute,”
vol. xlint (1911), 178.)
Prats 9. Nasturtium latesiliqua, drawn from specimens collected by Mr. F. G. Gibbs and myself
on Mount Arthur, Nelson, at an altitude of 4,000 ft. Figs. 1 and 2, flowers (x 3); 3, petal (enlarged) ;
4, stamens and pistil (x5); 5, pistil (x8); 6, section of portion of pod, showing 2-seriate seeds
(enlarged) ; 7, section of seed, showing position of embryo (enlarged) ; 8, 9, 10, 11, 12, various embryos,
variously magnified.
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WEINMANNIA RACEMOSA, Zinn. f
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PLATE 43.—WEINMANNIA RACEMOSA.
Famtny SAXIFRAGACEAL. | [Genus WEINMANNIA, Linn.
Weinmannia racemosa, Linn. |. Suppl. 227; Cheesem. Man. N.Z. Fl. 139.
Weinmannia racemosa was first gathered by Banks and Solander in Queen
Charlotte Sound (or Totaranui, as they called it) during Cook’s first visit, and was
fully described and figured in Solander’s manuscript “ Primitize Flore Nove
Zelandie,” a work which was never published. It was again collected by the
Forsters in Cook’s second voyage, and on the return of the expedition to Europe
was published in the “Supplementum Plantarum” of the younger Linneus, no
doubt from information supplied by the Forsters, for it appeared under the same
name in George Forster’s “ Prodromus,” published a few years later. Since then
it has been gathered in many localities in the middle and southern portions
of the North Island, and throughout the South Island. It attains its northern
limit, so far as is known, in the Kauaeranga Valley, Thames, and on the cliffs
fringing the Waikato River near Hamilton. Its most southern station is in Stewart
Island, where it is plentiful, often forming the chief portion of the lowland forest.
Its altitudinal range is from sea-level to over 3,000 ft.
Weinmannia racemosa is a large forest-tree, often attaining a height of from
50 ft. to 80 ft., or even more, with a trunk 1 ft. to 4{t. im diameter. It is usually
known by its Maori names of tawhero or kamahi, the latter being principally used in
the South Island. It is very closely allied to its congener Weinmannia sylwvicola, the
towai of the Maoris and settlers, and in the middle portion of the North Island the
two are often confounded. But in W. racemosa the leaves of the mature trees are
usually larger and 1-foliolate, and the branches are nearly glabrous; whereas in
W. sylvicola the leaves are smaller, and 3-foliolate or even imparipinnate, and
the branches are more or less pubescent. The leaves of young plants are generally
pinnate in both species, and vary excessively, especially in W. sylvicola, where the
number of leaflets may range from three to ten pairs.
The tawhero is so abundant in many forest districts that the timber has been
applied to a variety of purposes, such as fencing-posts, house-blocks, tramway-
sleepers, piles, &c. The general opinion, however, is that it is not durable where
exposed to alternations of weather, although it may be serviceable where it is used
under water, or altogether imbedded in the ground. It has been occasionally
employed for inlaying and ornamental turning, and as it possesses considerable
variety of colour and grain it might well be used to a greater extent.
The distribution of the genus Weinmannia presents some remarkable points.
Over eighty well-established species are known, more than half of which are found
in South America. Another centre of the genus is in Madagascar, from whence
more than fifteen species have been described. The Pacific islands contain twelve
or thirteen species, while Australia has four. The northern limit of the genus is
in Mexico in the New World, and in the Philippine Islands in the Old.
Puate 43. Weinmannia racemosa, drawn from specimens collected in the vicinity of Nelson.
Fig. 1, a pair of flowers (x 4); 2, stamen with glands (x 8); 3, back view of stamen (x 8); 4, pistil,
with glands on the disc (x 8) ; 5, longitudinal section of pistil (x 8) ; 6, ripe fruit (x 3) ; 7, seed (x 5).
Genied Ph
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A. TILLAEA MOSCHATA, DC 1-9.
B. TILLAEA STEBERIANA, Schultz, 10-20.
PLATE 44.—TILLAAA MOSCHATA anp TILLAVA
SIEBERIANA.
Famity CRASSULACE.. | [Genus TILLAA, Linn.
Tillaa moschata, D.C. Prodr. iii, 382; Cheesem. Man. N.Z. Fl. 140.
Tillza Sieberiana, Schultz, Mant. 111, 345; Cheesem. Man. N.Z. Fl. 143.
Tillea is a cosmopolitan genus that has representatives in most parts of the world,
but I do not know any country that contains so many species as New Zealand, no
less than ten being credited to it in our floras. Of these, the most interesting is
Tillea moschata, which has a widespread distribution as a maritime plant in high
southern latitudes. It stretches along the western coasts of South America from
Chile to Cape Horn; is abundant on the Falkland Islands; is found on Kerguelen
Island and Marion Island; and in New Zealand extends from Cook Strait to the
Auckland and Campbell Islands, Antipodes Island, and Macquarie Island. It is
thus one of those species found on most of those widely separated islands which
girdle the globe between latitudes 50° and 60° S., and its distribution can therefore
be described as being more or less circumpolar in its character.
As a species T. moschata can be separated from its New Zealand allies by its
much greater size, its stems often exceeding 6in. in length, its larger oblong-
spathulate leaves, and larger flowers. It is purely a coastal plant, and is never
found far from the sea. It should be remarked that it was first observed in both
Fuegia and New Zealand by Banks and Solander during Cook’s first voyage.
Tillea Sieberiana is a very different species from the foregoing. It is smaller
in all its parts, much more erect in its mode of growth, and usually more copiously
branched. The flowers are aggregated in clusters mixed with minute leaves, and
there are no scales at the base of the carpels. Like the previous species, it was first
gathered by Banks and Solander, and is abundant through the North and South
Islands, from the Three Kings Islands and the North Cape southwards to the south
of Otago. Although common on coastal rocks, it is by no means confined to the
vicinity of the sea, but extends into many inland districts, ascending to at least
2,000 ft. above sea-level. It is also widely distributed in Australia, ranging from
Queensland to Victoria, Tasmania, South Australia, and West Australia, and has
been recorded from several localities in the interior.
Prate 44a. Tullea moschata, drawn from specimens collected on the Auckland Islands by
Mr. J. S. Tennant. Fig. 1, portion of inflorescence (x5); 2, separate flower (x 5); 3 and 4, front
and back view of stamens (x 8); 5, carpels, showing the scale at the base of each carpel (x 8) ;
6, section of carpel, showing ovules (x 8); 7, section of ripe carpel, showing seeds (x 10); 8, seed
(x 12); 9, embryo (enlarged).
Prate 448. Tillea Sieberiana, from specimens gathered at Mount Wellington, near Auckland.
Fig. 10, pair of connate leaves (x 6); 11, portion of inflorescence (x 8); 12, bract (x 10); 13, single
flower (x10); 14, sepal (x12); 15, petal (x12); 16, stamen (x14); 17, carpels (x12);
18, section of carpel, showing ovules (x 12); 19, seed (enlarged) ; 20, embryo (enlarged).
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MYRIOPHYLLUM ROBUSTUM, Hook.f
PLATE 46—MYRIOPHYLLUM ROBUSTUM.
Famiry HALORAGACE4.] [Genus MYRIOPHYLLUM, Linn.
Myriophyllum robustum, Hook. f. Handb. N.Z. Fl. 67; Cheesem. Man. N.Z. Fl. 151.
Myriophyllum robustum, which is by far the finest of the New Zealand species
of the genus, was originally discovered by Mr. Colenso in some locality in the North
Island, and has since been found in many lowland swamps or marshes from Ahipara
and Mongonui southwards to Cook Strait. It is, however, far from being generally
distributed, and is often absent from extensive areas apparently well adapted for
it. Its chief centre of distribution is in the alluvial valleys of the Northern Wairoa,
the Waikato, and the Thames Rivers; where, however, it is rapidly being destroyed
either by the draining of the swamps or by the periodical burning of the vegetation
therein. In these districts it can still be seen covering large stretches in swamps
that are deeply covered with water in winter, but dry or nearly so in summer.
Although occasionally seen in lakes or streams it is by no means either lacustrine
or fluviatile to the same extent as its allies M. elatinoides and M. variefolium. In
the South Island it appears to be rare, although it has been recorded as far south as
Awatere on the east coast, and Hokitika on the west.
M. robustum belongs to a section of the genus in which the whole of the leaves
are verticillate and pectinately pinnatifid ; and is closely allied to the northern
M. verticillatum, which ranges through a large portion of Europe, northern Asia,
and North America. It can be distinguished from M. elatinoides and M. variefolium
without the slightest difficulty by the much stouter and coarser habit, and by the
upper leaves being verticillate as well as the lower. In addition to the above, the
nuts are much larger and evidently tubercled.
Prate 46. Myriophyllum robustum, drawn from specimens collected near Te Aroha, in the Thames
Valley, and in the Waikato near Mercer. Fig. 1, hermaphrodite flower and leaf; 2, stamen ; 3, trans-
verse section of ovary; 4, male flower; 5, section of male flower; 6, female flower; 7, transverse
section of ovary ; 8, longitudinal section of ovary ; 9, fruit without tubercles ; 10, fruit with tubercles ;
11, section of ripe carpel. (All enlarged.)
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West,Newman imp.
LEPTOSPERMUM SINCLAIRI, 7 4px
PLATE 47.—LEPTOSPERMUM SINCLAIRII.
Famity MYRTACE/..] [Genus LEPTOSPERMUM, Forsr.
Leptospermum Sinclairii, 7. Kirk, Students’ Fl. 158; Cheesem. Man. N.Z. Fl. 161.
The first mention of this somewhat critical species that I am acquainted with
is in Mr. Kirk’s paper on the “ Botany of the Great Barrier Island ” (Trans. N.Z.
Inst. 1 (1869), 146), where he refers to it under the name of Leptospermum ericoides
var. pubescens, saying that it is “a prostrate or subprostrate shrub, some-
times 3ft. high, at others appressed to the rock, like an alpine plant; leaves
more or less pubescent and ciliated; flowers fragrant, produced in immense
profusion, sometimes concealing the leaves; pedicels and calyx downy. This
would make a valuable bedding plant for the culturist. It was originally observed on
the island by the late Dr. Sinclair, but I am not aware of its occurrence elsewhere.”
In 1887, and again in 1889, I observed the same plant on the Three Kings Islands,
but so far it has not been seen in any other localities. Its distinguishing
characters, such as they are, lie in the smaller size, broader and flatter silky-hoary
leaves, larger flowers, longer pedicels, and much more deeply sunk capsules. It
must be admitted, however, that the plant is very near to small states of L. ericoides,
which, as every New Zealand botanist knows, is an exceedingly variable plant
responding very readily to changes in its environment. I cannot avoid the
suspicion that the separation of our plant as a species will not be maintained.
In the Great Barrier Island L. Sinclairii principally occurs on rocky peaks,
and is usually prostrate, or nearly so. On the Three Kings Islands I observed it
mainly on declivities leading down to the cliffs on the northern side of the Great
King. In this situation it was suberect, and the leaves were slightly narrower than
in the Barrier plant. Both forms agreed in the immense profusion with which the
flowers were produced, and in their delicious fragrance.
Puate 47. Leptospermum Sinclairii, dvawn from specimens collected on the Great Barrier Island
by Mr. T. Kirk. Fig. 1, leaves; 2 and 3, flowers; 4, section of flower. with the stamens removed:
5 and 6, front and back view of anthers; 7, ripe fruit ; 8, the same with a portion of the calyx removed ;
9, longitudinal section of fruit; 10, seed. (All enlarged.)
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ver oti whale rig) jae tiba & {i : raved ' ' ‘ll ie nat by ' at hy wpel 7
Ty eee ee a ae oe os aa (ahd PARAMS or cheer larch nT gl
of wis faa gait wil “A reall 4 pee Son wil a Tt ike HH
J ‘ @ ymirthe wih epidnht | 3 F AD iis ’ ie Oe) od. id » hae Te pe )
Co ie a! Veal wee’ frase Ghyieti) iis AY ar | rh es ; bi a Ar oe, ral, eva ;
Ce a a. i, av goat pels Vhs te. eta hp $e} XRd EO ae Lal ata
ies 7 tara ta Ae fe yO af) uae 7; aeingrt shin’ Uh sliknay “Prey satin er i
Y : ‘: dat twine ol 400 ia ad ite t Len) Ljaten[o tam rut it oot anes
; cla: Shor ie ayers yl laapehyiony oer A. Mopagpal tires torte de :
i levee f' bbienleF yor ia en Lele. ot 4 ane ers ‘Tay PabL wan’, i Riya ;
7 mt ait ie ajié re ae leull, nif ale) ode (1 nh gether! “art wilh wis inGh
we rewaraiy abril ts hoary y ay find hve ety ent oe ‘oly sar ly wih ae: yale:
=e of " are iesbieash igi conten ttn wiht he Be Ws cise! ott ee Cee oe
She ’ f al a adler fol) ceil af hae Ave ite mr hie it a % ;
: re vb cenit oe
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METROSIDEROS DIFFUSA, Sm.
PLATE 48.—METROSIDEROS DIFFUSA.
Famity MYRTACE/. } [Genus METROSIDEROS, Banks.
Metrosideros diffusa, Smith vn Trans. Linn. Soc. ii (1797), 268; Hook. f. Fl. Nov. Zel. i, 67 ;
Cheesem. Man. N.Z. Fl. 164.
The New Zealand species of Metrosideros can be arranged in three categories
according to their mode of growth. In one division, represented by the northern
rata (M. robusta) alone, the plant starts its life as an epiphyte high up in the branches
of some tall forest-tree, and sends aerial roots down to the ground, which ultimately
coalesce to form an irregular stem often of enormous size. A second class comprises
species like the southern rata (M. lucida), the pohutukawa (M. tomentosa), and
one or two others, which in the ordinary manner grow into tall trees with stout
stems and spreading branches. Lastly, there are quite a number of species best
described as woody climbers, which often reach the tops of the tallest forest-trees,
but never produce an erect tree-hke trunk. To this division. the subject of this
plate belongs.
Metrosideros diffusa, when seen in full bloom, is probably the most brilliant
of the species. The flowers are a much brighter crimson than those of the northern
rata (M. robusta) and pohutukawa (M. tomentosa), and are so abundantly produced
as almost to conceal the leaves. When climbing up the trunk of a rimu or other
tall forest tree, and laden with flowers, it presents the appearance of a veritable
pillar of flame, and such specimens are often conspicuous from afar. Unfortunately,
it is not a common species, although occurring sparingly in most forests of any size
from Mongonui and Kaitaia southwards to the East Cape and Taranaki. It has
been recorded from the Tararua Range, on the authority of the late Mr. Mantell ;
but I have seen no specimens. Although of slow growth it is by no means difficult
to establish in a garden, and is well worth general cultivation.
I am not at all sure that the name diffusa has been correctly applied to the
plant figured herewith. The originator of the specific name was George Forster,
who in his “ Prodromus ” published a “ Melaleuca diffusa,” giving as a diagnosis
the words “ foliis oppositis ovatis acuminatis, pedunculis rameis cymosis.”” Now,
the term “acuminate ” cannot be correctly apphed to the leaves of our plant, in
which they are obtuse or subacute, as a glance at the plate will show. Further, as
our plant is confined to the North Island it is hard to see how it could be collected
by Forster, who never landed on any part of it. He does not mention the exact
locality m which his specimens were obtained, but there can be little doubt that
they were gathered in Queen Charlotte Sound, where most of his collecting was done.
In all probability his plant is identical with the species usually known as
M. hypericifolia. The shape of the leaves and character of the inflorescence suit
his diagnosis, and the species is known to occur in the locality, where, in fact, it
was gathered by Banks and Solander during Cook’s first voyage. I regret that I
have no opportunity of examining a set of Forster’s plants, which would probably
decide the question.
It may be remarked that, although the descriptions of M. diffusa given by
Hooker in the “ Flora” and the **‘ Handbook” clearly refer to the plant illustrated
herewith, there must have been some confusion in his mind respecting it. This is
proved by the fact that he gives Banks and Solander’s M. myrtifolia as a synonym
of M. diffusa. But Solander’s description in his ‘“ Primitiz Flores Nove Zelandize ”
undoubtedly represents M. hypericifolia, and the specimens in the set of Banksian
plants presented to the Dominion by the trustees of the British Museum are
indisputably that species.
Prats 48. Metrosideros diffusa, drawn from specimens collected at Mount Wellington, near
Auckland. Fig. 1, calyx and style (x 4); 2, longitudinal section of calyx and ovary (x 4); 3, petal
(x 4); 4 and 5, front and back view of anthers (x6); 6, transverse section of ovary (x5); 7, ripe
fruit (x 3); 8, seed (enlarged).
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METROSIDEROS PARKINSONII, Buch.
PLATE 49.—-METROSIDEROS PARKINSONII.
Famity MYRTACE/,] [GENus METROSIDEROS, Banks.
Metrosideros Parkinsonii, Buch. in Trans. N.Z. Inst. xv (1883), 339, t. 28, f. 2; Cheesem.
Man. N.Z. Fl. 163.
Metrosideros Parkinsonii has by tar the most restricted range of any of the
New Zealand species, being confined to the north-west portion of the South Island,
stretching from Collingwood southwards to Mount Rochfort, the lower portion
of the Buller Valley, and the northern temination of the Paparoa Range. It
descends almost to sea-level in the Buller Valley, and ascends to nearly 3,000 ft.
on Mount Rochfort. According to Mr. Townson, it is at times a straggling shrub,
at others a small tree from 20 ft. to 25 ft. in height. Its brilliant crimson flowers
render it a very conspicuous object during the flowering season.
According to Mr. Kirk, it was originally discovered by Mr. W. S. Hayward on
the mountains behind Collingwood, but it was first made known to botanical science
through specimens gathered in the same district by Mr. H. H. Travers, and
communicated by him to Mr. Buchanan, who described and figured it in the
“ Transactions of the New Zealand Institute’ for 1882. It has since been found
in several localities in north-west Nelson by Mr. Dall, by Mr. Kingsley in the Buller
Valley, and by Mr. Townson on Mount Rochfort, Mount Frederic, and on the spurs
of the Paparoa Range. Iam indebted to the last-named gentleman for the specimens
from which this plate has been prepared.
The distribution of Metrosideros in New Zealand is mainly northern, although
one species (M. lucida) advances as far south as the Auckland Islands, where it
constitutes the principal tree. In the South Island the species are mainly found
on the western side of the Island. This is well seen in the neighbourhood of Westport,
where no less than six species occur on the seaward side of the Mount Rochfort
Range, their names being M. florida, M. lucida, M. hypericifolia, M. robusta,
M. scandens, and the subject of this plate.
Prats 49. Metrosideros Parkinsonii, drawn from specimens collected by Mr. W. Townson on
Mount Rochfort, near Westport. Fig. 1, flower-bud, not yet fully expanded ; 2, calyx and style ;
3, longitudinal section of calyx-tube and ovary; 4, petal; 5 and 6, front and back view of stamens:
7, transverse section of ovary; 8, ripe fruit ; 9, cartilaginous endocarp ; 10, seed. (All enlarged.)
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METROSIDEROS, ALBIFLORA, Solander
PLATE 50.—METROSIDEROS ALBIFLORA.
Famity MYRTACE| [Genus METROSIDEROS, Banks.
Metrosideros albiflora, Sol. ex Gaertn. Fruct. i, 172. t. 34, f. 11; Cheesem. Man. N.Z. FI. 164.
One of the most handsome species of the genus, at once recognized by the large
broad leaves and conspicuous panicles of white flowers. For its discovery we have
to go back to the year 1769, when it was collected by Banks and Solander at
Mercury Bay during Cook’s first voyage, a locality where it is still abundant in
those remnants of forest which have escaped the hands of the sawmiller. Solander
gave a full description of it in his unpublished “ Primitie Flore Nove Zelandiz,”’
accompanied by an excellent plate. It does not seem to have been again noticed
until 1826, when Allan Cunningham collected it at Whangaroa, unfortunately making
the mistake of referring it to Forster’s M. diffusa. Shortly afterwards it was
obtamed by Mr. Edgerly at Hokianga, and by others in various localities in the
northern half of the North Island. The most southern locality that I am personally
acquainted with is in the Bay of Plenty. Mr Kirk, in his “ Notes on the Botany of
the East Cape District ” (Trans. N.Z. Inst. xxix (1897), 520), records it from thence,
on the authority of Banks and Solander ; but, I think, erroneously, for the only locality
given by Solander is that of “ Opuragi,” or Mercury Bay. On the west coast of
the North Island I do not know of any station to the south of the Kaipara Harbour.
Its northern limit appears to be on the Maungataniwha Ranges near Kaitaia. Its
altitudinal range is from 250 ft. to 2,800 ft.
The distribution of Metrosideros albiflora, both latitudinal and altitudinal, is
very similar to that of the kauri (Agathis australis). But it is far more local, and
is consequently absent from many localities where the kauri is abundant.
Nevertheless, it cannot be denied that the Metrosideros has a special predilection
for kauri forests, and is more frequent therein than in ordinary forest. In other
words, it belongs to the kauri association, an assemblage of species which on the
whole always appears where the kauri is predominant. Common members of this
association are Astelia trinervia, Gahnia zanthocarpa, Alseuosmia macrophylla,
Dracophyllum latifolium, Senecio Kirkii, Lomaria Fraseri and L. discolor, and others
which cannot be particularized here.
Metrosideros is more highly developed in New Zealand than in any other
country, and assumes a much more diversified form and habit of growth. No other
country has climbing species, and no other country possesses a species which builds
a lofty trunk for itself by the coalescence of aerial roots. It has been suggested
that the climbing species have arisen on account of the presence of a moist atmosphere
and equable climate. But this theory does not explam why the Polynesian and
Malayan species, placed in a climate even more humid and equable, have not
developed in a similar direction.
Prate 50. Metrosideros albiflora, drawn from specimens collected by Mr. A. Gordon at Rua-
tangata, Whangarei. Fig. 1, flower-bud (x4); 2, petal (x4); 3 and 4, front and back view of
anthers (x 4); 5, upper part of style and stigma (x 4); 6, section of ovary (x 4) ; 7, ripe fruit (x 4) ;
8, the same, showing dehiscence by three valves (x4); 9, abortive seed (x5); 10, seed (x 6);
11, embryo (x 8).
Plate $1.
EPILOBIUM PALLIDIFLORUM, Soland.
PLATE 51.—EPILOBIUM PALLIDIFLORUM.
FAMILY ONAGRACE/. | [GENUS KPILOBIUM, LINN.
Epilobium pallidiflorum, Sol. ex A. Cunn. Precur. n. 550; Cheesem. Man. N.Z. Fl. 173.
The genus Hpilobium is remarkably well represented in New Zealand, about
thirty species being known. In whatever country the genus may exist, its species
for the most part present an extraordinary amount of variation. In the Northern
Hemisphere this has resulted in the creation of a multiplicity of so-called
“species ” differing only in shght and inconstant characters, and so closely linked
to one another as to make their discrimination a matter of great difficulty, even
those experts who have studied the genus being far from agreement. In New
Zealand the species are equally variable, but so far they have been treated on
fairly conservative lines, and but few attempts have been made to follow the
practice of the northern botanists. In the subject of the present plate, however,
we have a species possessing such strongly marked characters that there can be
no difficulty in separating it from any of its allies.
Epilobium pallidiflorum was one of the many plants observed during Cook’s
first visit to New Zealand in 1769. It was originally collected by Banks and
Solander in Mercury Bay (or Opuragi, as Cook called it), and was described by
Solander in his manuscripts under the name it now bears. The species, however,
was not actually published until 1839, when it appeared in Allan Cunningham’s
“ Precursor.’ Since its original discovery it has been found to have a wide range
in marshy places, along the banks of rivers, &c., and is now known to be plentiful
in such localities from the North Cape to Foveaux Strait. Its southern limit is
in Stewart Island, where it has been recently detected by Dr. Cockayne. It is
purely a lowland plant, and I have not seen it at a greater elevation than 1,500 ft.
It also occurs in Victoria and South Australia, and is abundant in Tasmania.
As already remarked, Epdobium pallidiflorum is a remarkably distinct species.
It can be recognized at a glance by its great size—I have seen specimens quite 4 ft.
in height—by its long acute leaves, and by its large white flowers. It is a great
ornament in the swamps of the Lower Waikato district, where it grows in the
utmost luxuriance along the banks of the streams which flow through the swamps,
mixed with such plants as Phormiwm, Calystegia sepium, and various sedges.
Prate 51. Epilobium pallidiflorwm, drawn from specimens collected at St. John’s Lake, near
Auckland. Fig. 1, flower (x 2); 2, flower with ovary and bract (x24); 3 and 4, anthers, front
and back view (x5); 5, stigma (x5); 6, ripe capsule (x 2); 7, seed (x 8).
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EPILOBIUM ROTUNDIFOLIUM, Forst.
PLATE 52.—EPILOBIUM ROTUNDIFOLIUM.
Famity ONAGRACE/.|] [Genus EPILOBIUM, Linn.
Epilobium rotundifolium, Forst. Prodr. n. 161; Cheesem. Man. N.Z. Fl. 179.
As in the case of the preceding species, for the discovery of this plant we have
to go back as far as the time of Cook’s first voyage, when it was collected by Banks
and Solander in Queen Charlotte Sound. Although fully described in Solander’s
manuscript “ Primitiz Flore Novee Zelandie” the species was never published
by him; and Forster, who collected it in the same locality durmg Cook’s second
voyage, selected the name it still bears when describing it in his ‘* Prodromus.”
His diagnosis, like all those given in the “ Prodromus,” is very short and
insufficient ; but a more complete description, taken from his manuscripts, appeared
in A. Richard’s excellent ‘‘ Flore de la Nouvelle Zélande.” Since then it has been
observed by almost all those who have interested themselves with New Zealand
plants, and is now known to be distributed in moist shaded localities throughout
almost the whole of the Dominion, from the North Cape to Stewart Island, and
from sea-level to nearly 3,000 ft. altitude.
E. rotundifolium is found in a great variety of situations. It is often seen
creeping over bare ground in somewhat open forest, especially where the soil is
naturally damp ; it is common on moist shaded banks down which water trickles,
and it is quite at home in mossy places by the side of waterfalls or on the shelving
sides of streams. Of late years it has taken possession of the sides of many railway
cuttings, where shaded from the sun and where the soil is sufficiently damp.
As a species EF. rotundifolium is perhaps most closely allied to E. linneoides.
But it is larger, and has a more erect habit of growth; the upper leaves are
alternate, and the inflorescence is terminal. In the “ Handbook” Sir J. D. Hooker
grouped it with the plants now known as E£. chlorefoliwum and E. imsulare, the
specific distinctness of which is now generally recognized. The first can always
be distinguished by the much more erect and firmer habit, and by the ovate or
ovate-cordate nearly sessile leaves; while the latter has weaker and more
elongated stems, with more distant sessile leaves, and the seeds are perfectly smooth.
It has also been compared with EH. alsinoides, but in reality there is little
relationship between the two plants. E. alsinoides is much smaller, the leaves are
smaller, paler, and more uniform in shape, the flowers are smaller and on longer
peduncles, and the capsules are evenly covered with a close and fine pubescence.
Puate 52. Epilobvum rotundifolium, drawn from specimens collected near Mercer, in the Waikato
district. Fig. 1, flower, with its subtending leaf or bract (x3); 2, flower (x3); 3, petal (x 6) ;
4 and 5, front and back view of anthers (x6); 6, stigma (x6); 7, ripe capsule (x 2); 8, seed
(enlarged).
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EPILOBIUM BREVIPES, Hook.f
PLATE 53.—EPILOBIUM BREVIPES.
Famity ONAGRACE. | [GENus EPILOBIUM, Liny.
Epilobium brevipes, Hook. /. Fl. Nov. Zel. ii. 328 ; Cheesem. Man. N.Z. Fl. 182.
Epilobium brevipes was originally collected in the Kaikoura Mountains
by Mr. Macdonald, a shepherd in the employ of Sir David Monro. Although
far from being a common plant, I have seen specimens from a considerable number
of localities in Marlborough and North Canterbury, and consequently can hardly
agree with the statement made by Mr. T. Kirk in his “ Students’ Flora,” where
he says that it is “of extreme rarity.” Mr. Kirk himself has recorded it from the
Upper Awatere; the Rev. F. R, Spencer has gathered it on Taylor’s Pass; and
Dr. Cockayne on Mount Fyfie. Mr. H. J. Matthews has sent me specimens from
the Hanmer district; Dr. Cockayne found it not uncommon in the gorge of the
Conway River; while Mr. J. D. Enys, Mr. T. Kirk, and myself have all collected
it on Mount Torlesse. Quite recently Dr. Cockayne has recorded its presence in
the gorge of the Broken River. It is usually, if not invariably, found on the faces
of limestone cliffs or precipices, and must be considered a purely rupestral species.
When once known EF. brevipes cannot be easily confounded with any other
species. The hard and woody base, the perfectly glabrous densely leaty stems,
the coriaceous smooth and shining elliptic leaves, and the glabrous capsules, which
are borne on very short peduncles which hardly elongate in fruit, are well-marked
characters, the significance of which cannot be overlooked. Probably its nearest
ally is EB. crassum, which is a much smaller more creeping plant, with longer more
obtuse leaves with the broadest part above the middle, larger flowers, and much
longer peduncles. Coriaceous states of BE. glabellum have been mistaken for it,
but the two species differ so completely in habit, foliage, inflorescence, and fruit
that if ordinary care is taken they ought never to be confused.
Prater 53. Epilobium brevipes, drawn from specimens collected by Dr. Cockayne in the gorge
of the Conway River, and by myself on Mount Torlesse. Fig. 1, tip of leaf (enlarged) ; 2 and 3, flowers
(x 4); 4 and 5, front and back view of anthers (x 6); 6, stigma (x 6); 7, ripe capsule (x 3): 8, seed
(enlarged).
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J.N.Fitch lith.
EPILOBIUM MELANOCGAULON, Zook.
Puare 54.—EPILOBIUM MELANOCAULON.
Famiry ONAGRACE. | [Genus EPILOBIUM, Lin.
Epilobium melanocaulon, Hook. Ic. Plant. t. 813 ; Cheesem. Man. N.Z. Fl. 183.
Epilobium melanocaulon was discovered at about the same time by Mr. Colenso
in the Ruahine Range and by Mr. Bidwill in the Nelson Provincial District. So
far as the North Island is concerned, I have seen no specimens except those
collected by Mr. Colenso; but in the South Island it has been found to be
generally distributed in suitable localities from Nelson to Foveaux Strait. It is
essentially one of those plants most abundant on the great stretches of shingle which
fill the beds of many of the larger rivers which drain the Southern Alps. These
banks of shingle may be covered in heavy floods, but ordinarily are quite bare, and
support a very peculiar vegetation, such as several species of Raoulia, Helichrysum
depressum, Muhlenbeckia axillaris, Veronica Bidwilla, and many others, among
which the subject of this plate is always conspicuous.
In its typical state H. melanocaulon is perhaps as easily recognized as any
species of the genus. Its erect sparingly branched rigid stems, springing in clumps
from a hard and woody rootstock, and remarkable for their black or purplish-black
colour, are most characteristic, as are the close-set and deeply-toothed leaves.
The small flowers, and glabrous capsules on very short peduncles, are also excellent
marks of the species. Its nearest ally is doubtless my FH. rostratum, which has the
same habit, but is much smaller, paler in colour, and covered with a uniform grey
pubescence. The capsules are markedly different, being abruptly narrowed into
a short beak at the tip. I do not know that it has any other close ally. In the
“Handbook” Sir J. D. Hooker remarks that “‘ states of it are difficult to separate
from E. confertifolium, glabellum, and brevipes.” But the first and last of these
are really altogether different in habit, foliage, and inflorescence; and as for
E. glabellum, it can generally be distinguished without difficulty by the much
more developed inflorescence, which gives an entirely different aspect to the plant.
The leaves are also much larger and broader, and not so deeply toothed.
In the Manual I have reduced Professor Haussknecht’s £. polyclonum to the
position of a variety of H. melanocaulon. It chiefly differs in the more slender
branched stems, which are bifariously pubescent, and in the more distant less deeply
toothed leaves ; but these characters are barely sufficient to warrant its separation
as a species.
PratE 54. Epilobium melanocaulon, drawn from specimens collected on the shingle of the Cass
River, Lake Tekapo. Fig. 1, flower, with its subtending leaf or bract (x 3); 2, petal and stamen
(x5); 3, back view of anther (x5); 4, stigma (x5); 5, 11pe capsule (x 2); 6, seed (x 8).
Lary, op taliiili
fpaleas viens Poa
ae soy ak bE . ak
al a arise Nas, | ab Hy - oT ; ‘ eo
i ‘ 3, Wad ees’. wll ‘ } .o is ‘a ee
- se mt itr it eral tai ae Ma 4 viet en : - ened “
4 ‘ ihe’ ol hw ite vu Ne % ‘al atyeliteipA, «i Ao cia! J , ated i
ae ne if ape id mide ceed ALi Gn ST id . oy i S am on
. {ees i, o * ale Rk Mal bine ag lait asy? eats ie] “" Pb Nd
gl 7 Pad
=
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y
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savadedls vital & alsa bia) be ose is tion eb! enn Ald adore mes eh ise
hime eu! teh Jalal varia ‘ ; ih, Sb dy aa: raat falzoobaas ‘
ae raga? ai Jive 4) Aerie tii) rene ‘2 sie % \
; iif os ate
; . ; ; vs
\
a ott bu wlenide nie ag ii Os Bhai mS woe {coll
i isan Hs Wear ee Le “) vrais 2 20%
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Plate a5.
West, Newman imp.
M.Smith del
J.N.Fitch lith.
EPILOBIUM GLABELLUM, forse.
PLATE 55.—EPILOBIUM GLABELLUM.
Famiry ONAGRACEA:. | [Genus EPILOBIUM, Linn.
Epilobium glabellum, Forst. Prodr. n. 160; Cheesem. Man. N.Z. Fl. 185.
The subject of this plate was first gathered by Banks and Solander in Queen
Charlotte Sound in 1769, and was again collected in the same locality by Forster
during Cook’s second voyage in 1773. Forster published the species in his
* Prodromus ” under the very appropriate name of H. glabellum, which appears
to have been taken from Solander’s manuscripts, but his diagnosis was much too short
and scanty for a difficult genus like Hpilobiwm ; and succeeding writers have not
always properly understood his plant. According to Professor Haussknecht, whose
splendid monograph of the genus is usually followed by botanists, this was the case
with Allan Cunningham and A. Richard, both authors mistaking a form of
E. junceum tor Forster’s glabellum. The matter i still further complicated by the
wide limits which Sir J. D. Hooker assigned to EF. glabellum. He included in it,
in part if not altogether, the Austrahan plants now distinguished as 1. sarmentaceum
and EH. erosum, the South American #, Lechleri and EF. australe, and in addition
took in several New Zealand forms now considered to belong to other species. It
is consequently by no means surprising that the species has been much misunder-
stood.
The characters which separate H. glabellum from the allied species are the
numerous erect stems, which are either glabrous or provided with 2 or 4 faint
pubescent lines decurrent from the petioles; the perfectly glabrous remotely
sinuate-denticulate oblong or ovate-oblong leaves, which are often reddish or
purple, and frequently shinmg; and by the lax terminal inflorescence, with
moderately large flowers, the capsules being glabrous and borne on short peduncles
which are seldom longer than the leaves. The average or typical state of the
species, as characterized above, is a very common plant throughout the greater
part of the Dominion, especially in mountain districts in the South Island, and
can be recognized without the slightest difficulty. But, unfortunately, it runs into
numerous varieties the systematic position of which is very difficult to define. One
of the most conspicuous of these is the plant separated by Haussknecht as a distinct
species under the name of #. erubescens. It has more rigid stems, with crowded
and more erect leaves; the flowers are more numerous, and the capsules shorter
and nearly sessile. It is certainly a well-marked variety, but is connected with
the type by too many intermediate forms to be retained as a separate species.
I have not myself seen EH. glabellum to the north of the Waikato River,
and look upon it as decidedly rare northwards of the East Cape, although common
enough further south. Haussknecht, however, states that he has examined
specimens from the Bay of Islands, “between Waitangi and Kerikeri.” It is
somewhat remarkable that it has not yet been found in Stewart Island, especially
as Dr. Cockayne has recorded the very closely allied EF. nove-zelandie from thence.
It should perhaps be stated that EF. glabellum is confined to New Zealand, the
Australian and South American plants associated with it by Hooker and other
writers being now regarded as distinct.
Puate 55. Epilobiwm glabellum, drawn from specimens collected in the Hooker Valley, Mount
Cook district, at an altitude of 3,000 ft. Figs. 1 and 2, flowers (x 2); 3 and 4, front and back view
of anthers (x 4); 5, stigma (x 4); 6, ripe capsule (x 2); 7, two seeds (enlarged).
ak hs Se ——
a8 F thi
it te ciara
aol Nis. iM
1 arta eo
solide iy ) =e
“aed vara Aten Lisi
sift & APL im
iietanings | sot rabae
sig’ dialer ches?
dyiw™ lit Wesnsi ree
‘pelrtnintaay PTD i BA eat bes! OP Pa) a a bee! inne ranean
ads = ah: hs rin’ War unt 1 os : ~ a La dh egyl ie?
Dia os at paint wren? Pripiag pedir i’ ' Fi " a uy he 4
es, Snipa. Wa wet, gif) “no aPleelely) wot ueeree? eh" L Lit iligerare (8
ret wer Hi eitwenstrot, mina ed TAG ds ive. odd. J VOTE Slay +00 ie
vs, ~ ee “onitth ue iP Acie ay » gee, GUE FY eh), dei RN HSMN,
: glean wal, bo oy yan Hindle ves or Print il play ak bball ott Ate
ceithity Priyy 10 0 nol ‘] po ae) rh ne) UE ty ee bale’ wy aes
F watt tent en aoe LAs: ae uh. Ee a ie
“Aatieniad at tut vielen berth taten THE, G Vint pithy: dBeAt tif 1:5 ed
wn) vas & ab imathdsr ml yee We) peal ‘ pays peti aN pete ees
el 3 beds Thee ali wn tpn Cone lis nie fr Pes
ast Aba au) healt wl? vo | Ara vf i Kile re vari si iA Beet
Na a ha in 9 mee) ott ica Poa neal ies) ‘wait 7 sey: be
28 guid ont le! inte) (iil paval lek eto i wise 2
Pf & Snijato! at mies Tint! Awol 1 " ‘ {a5
wriakt inte ean ott Sighs 4 ra
wh? vielen » PY a tid rie , a went ly. 4
hy ele, % houht Pn ge [pat disshaglan nt
ay “wie Ratt agra i “fl te
Plate 56.
West, Newman imp.
FUCHSIA PROCUMBENS, 2. Cunn.
PLATE 56.—FUCHSIA PROCUMBENS.
Famity ONAGRACEZ.] [Genus FUCHSIA, Linn.
Fuchsia procumbens, R. Cunn. ex A. Cunn. Precur. n. 534; Cheesem. Man. N.Z. Fl. 187.
Quite apart from the interest which from the point of view of geographical
distribution must always attach to the presence in New Zealand of three species
of the otherwise purely South American genus Fuchsia, there are certain biological
facts concerning the New Zealand species which deserve more searching examination
than has yet been given to them. I allude to the curious heterostyled flowers
which all the species produce. Mr. T. Kirk, who was the first to investigate the
matter with any detail, showed that in F. excorticata and F. Colensoi there are three
forms of flower, each with a style of different length, which can be distinguished
as the long-styled, the mid-styled, and the short-styled forms; and that these
differences in the length of the styles are correlated with differences in the
length of the stamens, the long-styled form having the shortest stamens, and the
short-styled form having the longest. The remaining species, F. procumbens, the
subject of this plate, possesses the three kinds of flower differmg in the length of
the style, but the stamens are the same length in each case.
The discoverer of F. procumbens was Richard Cunningham, who in 1834
collected it at Matauri Bay, a little indentation on the coast-line between Whangaroa
and the Bay of Islands, and situated almost directly opposite the Cavallos Islands.
It was first described by his brother, Allan Cunningham, in the well-known
“ Precursor,” and a few years later was figured by Sir W. J. Hooker in the
“Icones Plantarum” (t. 421) from specimens collected in the same locality by
Mr. Colenso. From an examination of this plate, and from Cunningham’s
description, it is evident that the plant was the one now recognized as the long-
styled form, the large globose stigma being far-exserted beyond the flower. For
many years no further information was obtained, and no one supposed that other
forms of the species existed. In December, 1867, however, Mr. T. Kirk gathered
a Fuchsia in Tryphena Harbour, Great Barrier Island, that differed from
Cunningham’s plant in the shorter broader flowers, and especially in the short style,
which was entirely included in the flower, the stigma being much below the level
of the anthers. These differences were so noticeable that Sir J. D. Hooker treated
the plant as a distinct species, and figured it in the “ Icones Plantarum ” (t. 1084)
under the name of Fuchsia Kirkii. But he also acknowledged that in habit and
foliage the two plants were identical, and that at first he was inclined to regard
the differences as sexual. A few years later another form of the species found
its way into cultivation, in which the style was of intermediate length, the stigma
being on the same level as the anthers. This discovery proved that F. procumbens
agreed with F'. excorticata in possessing three distinct forms of flower, but differed
(as has been stated above) in the stamens being of the same length in each form.
Since the publication of Mr. Kirk’s paper on “ Heterostyled Trimorphic
Flowers in the New Zealand Fuchsias ” (Trans. N.Z. Inst. xxv (1893), 261)
much information has been obtained respecting F. procumbens, and some of
his conclusions require modification. In the first place, the species is not nearly so
rare as he supposed, as will be seen from the following list of localities : Cabbage
Bay, J. Adams! (short-styled form) ; Tryphena Harbour and Mine Bay, Great
Barrier Island, T. Kirk! A. J. Osborne! L. T. Griffin! (short-styled form) ;
Whangarei Heads and coast near Ngunguru, T. F. C. (short-styled form) ;
Whangaruru Harbour, T. Kirk (short-styled form); Matauri Bay, R. Cunningham
2
(long-styled form); Kawerua, Hokianga, Dr. Cockayne (form not stated) ;
Ahipara, T. F. C. (long-styled form) ; Waihi, fifteen miles north of Ahipara, T. F. C.
(mid-styled and short-styled forms); coast near Houhora and near Cape Maria,
T. F. C. (form not recorded) ; between the North Cape and Parengarenga, abundant,
T. F. C. (mid-styled and short-styled forms). From the above it will be noticed
that Mr. Kirk’s belief that the different forms never grow intermixed is not correct.
At Ahipara all three forms occur within fifteen miles of one another, if not nearer ;
and two of them grow side by side at Waihi. At the same time, it is most remark-
able that one form alone—the short-styled—has yet been found to the south of the
Bay of Islands.
Mr. Kirk also states that fruit is seldom seen on the mid-styled and _ short-
styled form in cultivation, and never in the wild state. But I have cultivated the
mid-styled form (and no other) in my garden for more than twenty years, and
every year fruit has been produced in comparative abundance. I have seen as
many as forty ripe berries at one time on a single plant. And I understand that
the experience of other cultivators has been the same. It is therefore clear that,
so far as the mid-styled form is concerned, it is not necessary for the production
of fruit that the flowers should be fertilized by pollen from one of the other forms.
I much regret that neither of the two other forms has been in cultivation in
Auckland for some years past, and that consequently I am unable to say whether
they also ripen their fruit in gardens. As regards the production of fruit in the
wild state, all my visits to the localities where the plant exists have been too early
in the season for me to obtain it. But Mr. A. J. Osborne informs me that the
short-styled form ripens fruit freely at Tryphena Harbour, and I am assured that
fruit is regularly observed on all the forms in the North Cape district.
It is much to be desired that some horticulturist would cultivate the three
forms of F. procumbens and ascertain what results would follow if they were inter-
crossed. It must be admitted that the fact that the mid-styled form is fertile with
its own pollen, and that the stamens are of the same length in all the forms, is
opposed to the idea that the case is one of pure heterostylism as usually under-
stood. We require to know if there are differences in the pollen of the three forms ;
if some of the unions are fertile and others infertile—in short, whether the forms
are specially adapted for reciprocal fertilization. Until such proof is obtained
it is perhaps reasonable to suppose that the long-styled form, with its larger
stigma and somewhat smaller anthers, is simply a variety advancing in the direc-
tion of becoming a female plant; while the short-styled variety, with its smaller
stigma and more abundant pollen, is in a similar way tending in the direction of
becoming purely male.
PuatE 56. Fuchsia procumbens. A, mid-styled form, drawn from specimens cultivated at Auck-
land; B, short-styled form, from specimens collected at Tryphena Harbour, Great Barrier Island,
by Mr. A. J. Osborne; C, from specimens gathered in Ahipara Bay, North Cape district. Fig. 1,
section of flower (mid-styled form); 2 and 3, front and back view of anthers from same; 4, section
of ovary ; 5, section of ripe fruit; 6, seed; 7, embryo; 8, section of short-styled form; 9, section
of long-styled form (the stigma is usually larger and further exserted than represented by the artist).
(All the figures enlarged.)
Plate 57.
West, Newman imp.
M. Smith del.
JN Pitch lith.
TETRAGONIA TRIGYNA, Banks & Sol.
PLATE 57.—-TETRAGONIA TRIGYNA.
Famity FICOIDE.] [Genus TETRAGONIA, Linn.
Tetragonia trigyna, Banks & Sol. ex Hook. f. Fl. Nov. Zel. i, 77; Cheesem. Man. N.Z.
Fl. 192.
Tetragonia trigyna was first discovered by Banks and Solander in Queen
Charlotte Sound in 1769. It was not only excellently described by Solander in
his manuscript “ Primitiz Flores Nove Zelandie ” under the name it now bears,
but a faithful drawing was also prepared for the unpublished Banksian plates,
making it quite easy to be sure of the identity of Solander’s plant. Unfortunately,
succeeding explorers confused it with the closely allied 7. expansa, an easy thing
to do when working on dried specimens, which in a genus possessing the thick and
fleshy habit of Tetragonia lose many characters which are obvious enough in a fresh
state. Thus neither A. Richard, A. Cunningham, nor Raoul mention more than
one Tetragonia as found in New Zealand. Sir J. D. Hooker, in his “ Flora Nove
Zelandiz,” also places both species under 7. expansa, but he quotes Solander’s name
of T. trigyna as well, this being the first occasion that it was actually published.
About this time specimens and information respecting the species were forwarded
to Kew by Dr. Sinclair and Mr. Colenso, and it was definitely published by Sir J. D.
Hooker in the ** Handbook” in 1864.
The range of 7. trigyna within the Dominion is now fairly well ascertained.
It occurs in rocky or sandy places on many parts of the coast-line, from the
Kermadec Islands and the North Cape to Stewart Island, and is also found on the
Chatham Islands. Although common in many localities it is comparatively rare
in others, and notwithstanding its wide distribution is by no means generally
spread. Comparing it with 7. expansa, it is quite obvious that no one familiar
with the two plants in a living state would ever confuse them. T. trigyna differs
markedly in its trailing or almost climbing habit, in its smaller and broader more
obtuse leaves, and particularly in the globose bright-red succulent fruit, which is
not at all horned, which is always the case in 7. expansa.
It is doubtful whether 7. trigyna is not identical with the Australian
T. implexicoma, Miq.; and, if so, Miquel’s name will take precedence. It was
published in 1844; whereas 7. trigyna cannot possibly date earler than the
publication of the “Flora Nove Zelandiz” in 1853; Solander’s manuscript
** Primitie,” unfortunately, not counting as “effective publication.” According
to descriptions, however, the Australian plant has longer and more acute leaves,
and longer peduncles. Until a careful comparison can be made it is better to
treat the two plants as distinct.
Puate 57. Tetragonia trigyna, drawn from specimens collected on the Little Barrier Island and
Cuvier Island. Figs. 1 and 2, flowers; 3, calyx laid open; 4 and 5, front and back view of anthers ;
6, longitudinal section of ovary ; 7, transverse section of ovary ; 8, fruit; 9, section of fruit ; 10, seed ;
11, embryo. (All enlarged.)
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Plate 58.
West, Newman imp.
M. Smith del.
JN. Fitch lith.
HYDROCOTYLE ELONGATA, A.Cunr.
PLATE 58.—HYDROCOTYLE ELONGATA.
Famiry UMBELLIFER.. | Genus HYDROCOTYLE, Linn.
Hydrocotyle elongata, 4. Cunn. Precur. n. 495; Cheesem. Man. N.Z. Fl. 195.
The cosmopolitan genus Hydrocotyle is well represented in New Zealand, nine
or possibly ten species being found therein. Of these, by far the finest is the
subject of this plate, which was first discovered by Richard Cunningham in
the year 1834 near the well-known Kerikeri Falls, in the Bay of Islands district:
the favourite botanizing-ground of the Cunninghams, Colenso, Hooker, and others.
Subsequent collectors have found it to be fairly plentiful in moist shaded
localities from the North Cape to the south of Otago. It is mainly a lowland
plant, and I have no notes of its occurrence at a greater altitude than about
2,000 it.
H. elongata is not closely allied to any other of the New Zealand species.
Its distinguishing characters are the large size, softly pilose habit, deeply lobed
leaves, long peduncles, and very conspicuous slender pedicels to the flowers.
In some points of habit and appearance H. dissecta approaches it, but is easily
separated, by the smaller size, more deeply lobed leaves, many-flowered umbels,
and the sessile flowers. H. americana, H. pterocarpa, and H. nove-zelandia@
have shallow lobes to the leaves, which are almost glabrous, and the flowers are
nearly sessile in the umbels. I am not acquainted with any foreign species
which can be considered closely allied.
_ Puarr 58. Hydrocotyle elongata, drawn from specimens collected in the Whangarei district.
Figs. 1 and 2, flowers (x 8); 3 and 4, front and back view of anthers (x 8); 5, ovary (x 8); 6, ripe
fruit (x 5); 7, a single mericarp, seen from the commissure (x 5); 8, section of fruit (x 5).
Flate 59.
West,Newman mp.
.N.Fitch lith.
M. Smith del.
J.
AZORELLA HAASTII, Benth 2 Hook f
PLATE 59.—AZORELULA HAASTII.
Famity UMBELLIFER/..] [Genus AZORELLA, Lam.
Azorella Haastii, Benth. & Hook. f. ex T. Kirk, Students’ Fl. 192: Cheesem. Man. N.Z.
Fl. 201.
For the first discovery of this fine plant we are indebted to Sir Julius von
Haast, who in May, 1862, collected it in the upper part of the Hopkins River,
which, after uniting with the Dobson, flows into the head of Lake Ohau. Later
on he gathered it in the valley of the Hunter, above Lake Hawea, and subsequently
in several localities in the central chain of the Southern Alps. In 1864 Sir J. D.
Hooker published it in the first part of the ‘‘ Handbook of the New Zealand Flora,”
very appropriately using the name of its enthusiastic discoverer, who in the years
between 1860 and 1870 did more for the botanical exploration of the mountains
of the South Island than any other person. Since that period it has been proved
to be fairly plentiful in most of the mountain districts of the South Island, from
Nelson to Otago. Its altitudinal range is from 2,0001t. to 5,000 ft. There are
specimens in Mr. Kirk’s herbarium collected on the Ruahine Range by Mr. A.
Hamilton, but I have seen no other examples from the North Island.
The nearest ally of A. Haast is undoubtedly the Auckland Island
A. reniformis, which in fact only differs in its smaller size, entire stipules, few-
flowered umbels, and fruit always longer than its pedicel. But reduced states of
A. Haastii come very near to A. reniformis, and in fact have been mistaken
for it both by myself and the late Mr. Kirk (see ‘Students’ Flora,” p. 192).
I cannot doubt that both plants are recent offshoots from a common parent.
The New Zealand species of Azorella were originally placed in the genus Pozoa
by Sir J. D. Hooker. In the “Genera Plantarum” Pozoa was merged with
Azorella ; but the New Zealand species (with the exception of A. Selago, which
is a typical Azorella) were placed in Hooker’s subgenus Schizeilema, which was
originally proposed in the “Flora Antarctica” for A. reniformis. In a recent
number of Engler’s “ Botanische Jahrbucher” (vol. xl, p. 573) Dr. Karl Domin
proposes to constitute a separate genus of Schizeilema, pointing out that it is
not only separated by a marked difference in habit, but by other characters of
importance. He would include in it not only the New Zealand forms, but also two
species from the extreme south of South America, and the Australian Azorella
Muelleri, Benth. (Schizeilema fragosum, Domin). There is much to be said in
favour of Domin’s views; and if the present plate had not been already struck
off when his memoir reached New Zealand I should have adopted his name of
Schizelema Haastii.
It may be stated that in Domin’s memoir a variety of A. Haasti is stated
to possess “blue petals.” This I believe to be incorrect so far as the fresh state
is concerned, but I have frequently noticed the petals to take on a blueish tinge
after drying.
Prare 59. Azorella Haastii, drawn from specimens collected in the Hooker Valley, Mount Cook
district, at an altitude of 3,000 ft. Fig. 1, sheathing-base of petiole, showing the ciliate margins
(enlarged) ; 2 and 3, flowers (x 8); 4a single petal (x 10); 5 and 6, front and back view of anthers
(x 10); 7, fruit (x 6); 8, a single mericarp of the fruit (x 6); 9, section of fruit (x 6).
i Tv t iit mh yeule Dede’ 9 A. -
PAG. re’ Ge bal ae dy ayia! as oats af aps
miele 58 og He si wi wvenf ge polyem oo We : Ag
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i ; ‘oy
Plate 60.
CHP 29
Recess fi: West,Newman imp.
AZORELLA ROUGHII, Benth 2 Hook f
Sr
PLATE 60.—AZORELLA ROUGHII.
Famity UMBELLIFER 4. } [Genus AZORELLA, Lam.
Azorella Roughii, Benth. & Hook. f. ex T. Kirk, Students’ Fl. 192; Cheesem. Man. N.Z. Fl.
201.
Azorella Rough was one of the discoveries of Captain D. Rough, who
between the years 1858 and 1862 gave a considerable amount of attention to
the mountain - flora of the Nelson Provincial District. His specimens were
obtained on Dun Mountain, near Nelson—an excellent collecting-ground, and one
on which many interesting plants were noticed for the first time, as, for instance,
Stellaria Roughw, Lobelia “‘Roughii, Myosotis Monroi, Pimelea Suteri, &c. In 1862
the species were published in the ** Handbook” by Sir J. D. Hooker under the
name it now bears. Its range is much more limited than that of A. Haastii, and
so far I have not seen specimens from the south of the Spenser Mountains. — It is
abundant in many parts of the Dun Mountain Range, extending southwards along
the Raglan Mountains to the Wairau Gorge, where ‘T collected it many years ago.
Still further south it has been gathered by Mr. T. Kirk and myself on several of
the mountains flanking the Waiau and Clarence Valleys. Its north-eastern limit
appears to be on Mount Stokes, from whence I have seen specimens obtained by
Mr. Macmahon. Mr. Gibbs and myself have gathered it on Mount Arthur and
other peaks in north-western Nelson; and I “have seen specimens collected by
Mr. Townson in some locality near Westport, although it is not mentioned in his
list of Westport plants printed in the ‘ Transactions of the New Zealand Institute ”
(vol. xxxix, p. 380).
As a species A. Roughii comes very close to A. Haastw, of which it has the
size, habit, and inflorescence. But it can always be readily separated by the
3-5-foliolate or -partite leaves. A. pallida is also allied, but is much smaller and
more slender, and the leaves are much more membranous.
ee
Prate 60. Azorella Roughii, drawn from specimens collected by Mr. F. G. Gibbs on Ben Nevis,
at an altitude of 4,500 ft., and by myself on Mount Arthur. Fig. 1, sheathing-base of petiole, showing
the stipuliform margins ; 2 and 3, flowers (x8); 4, petal (x 10); 5 and 6, front and back view of
anthers (x 10); 7 and 8, fruit (x 6); 9, section of fruit (x 6).
.
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Flate 61.
West, Newman imp.
J.N.Fitch lith.
ACIPHYLLA HOOKERI, 7 Kirk.
PLATE 61.—ACIPHYLLA HOOKERI.
Famity UMBELLIFER.] [Genus ACIPHYLLA, Forst.
Aciphylla Hookeri, T. Kirk, Students’ Fl. 209; Cheesem. Man. N.Z. Fl. 210.
The subject of this plate is one of the most remarkable species of a very
remarkable genus. It was first discovered by Mr. J. Dall, well known for his
botanical explorations in the Collingwood district, in the high mountainous country
near the source of the Heaphy River, which, rising on the western side of the
watershed behind Collingwood, flows in a south-westerly direction until it enters
the sea no great distance to the north of Karamea. In all probability it
extends southwards on the summits of the coastal ranges; for Mr. Townson,
when investigating the vegetation of the vicinity of Westport, found it to be
not uncommon on the Brunner and Paparoa Mountains, at an elevation of
from 2,500 ft. to 4,000 ft. It is specially plentiful on Mount Faraday and
Mount Buckland, from whence Mr. Townson obtained the specimens used in
the preparation of this plate.
A. Hookeri is a very distinct species, quite unlike any other. It can be
recognized at a glance by the short flat squarrose ultimate segments of the
leaf, which are flat or concave above, but keeled beneath, and which give the
tips of the leaves a curious trifid appearance quite foreign to the rest of the
genus. Probably its nearest relative is A. Lyallii; but that species has much
longer and narrower leaf-segments, which are never squarrose, as in A. Hookeri.
In addition to that, the whole aspect of the two plants is different. 4. Hookeri
should therefore be regarded as an unusually isolated form of the genus, restricted
to a limited district in the north-west portion of the South Island. Its
geographical area almost exactly corresponds with that of Drimys Traversii,
Metrosideros Parkinsoni, and Gentiana Spenceri, three equally distinct and
remarkable plants.
Prate 61. Aciphylla Hookeri, drawn from specimens collected on Mount Buckland, near West-
port, at an altitude of 4,000 ft. Fig. 1, tip of leaflet (enlarged); 2, flower (x 8); 3, the same with
the petals removed (x 10); 4 and 5, front and back view of anthers (x 10); 6, fruit (x5); 7, single
carpel (x 5); 8, transverse section of carpel (x 5).
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Plate 62.
West, Newman imp.
M.Smith del.
J.N.Fitch lith.
Cheesem.
ACIPHYLLA TOWNSONI,
PLATE 62.—ACIPHYLLA TOWNSONI.
Famity UMBELLIFER.] [Genus ACIPHYLLA, Forst.
Aciphylla Townsoni, Cheesem. Man. N.Z. Fl. 1138.
This is a comparatively recent addition to the New Zealand flora, having
been first gathered by Mr. Townson, in the autumn of 1901, on Mount Buckland,
the northern termination of the Paparoa Range, and situated only a short distance
to the south of the mouth of the Buller River. Since then Mr. Townson has
collected it on Mount Kelvin, Mount Faraday, and some others of the peaks of
the Paparoa Range, usually between the altitudes of 3,000 ft. and 4,500 ft. He
has also met with it on the Lyell Mountains, to the north of the Buller Valley;
but, curiously enough, has never been able to find it on Mount Rochfort and the
other coastal peaks immediately to the north. The species was published by
myself in the Appendix to the Manual, and I had much pleasure in associating
Mr. Townson’s name with it. For many years he had diligently investigated the
vegetation of the Westport district, making numerous interesting discoveries,
and obtaining much novel information respecting the geographical range of the
species.
As a species A. Townsoni falls naturally into the neighbourhood of A. Lyallit,
and is not very far removed from the variety crenulata of that species. But it is
amply distinct in the much more slender and more flaccid habit, and in the
extremely narrow leaf-segments, which do not exceed sj M. In diameter, and in
some forms are even narrower. No other species of Aciphylla, in fact, has
such narrow leaves. The flowers and fruit appear to be of the same type as
A, Lyalliv.
PuavE 62. dAciphylla Townsoni, drawn from specimens collected on Mount Buckland, near West-
port, at an altitude of 4,000 ft. Fig. 1, portion of leaf (enlarged) ; 2, male flower, with bracts (x 10) ;
3, flower, with petals and stamens removed (x 12); 4, petal (x 10); 5 and 6, front and back view
of anthers (x 12); 7, fruit (x 8); 8, 9, and 10, sections of fruit, showing variation in the number
of the ribs (x 8).
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Plate 63.
AGIPHYLLA MUNROI, Zook. f ~
PuarE 63._-ACIPHYLLA MONROI.
Famity UMBELLIFER. | [Genus ACIPHYLLA, Forsr.
Aciphylla Monroi, Hook. f. Fl. Nov. Zel. ii, 330 ; Cheesem. Man. N.Z. Fl. 212.
One of the earliest explorers of the vegetation of the north-eastern portions
of the South Island was Sir David Monro, who for many years occupied a
prominent position in the parliamentary life of the colony, ultimately holding
the position of Speaker of the House of Representatives. How much he did for
botany in New Zealand can be roughly estimated from the following list of
prominent species dedicated to him by Sir J. D. Hooker or other botanists:
Ranunculus Monroi, Carmichelia Monroi, Aciphylla Monroi, Celmisia Monroi,
Raoulia Monroi, Senecio Monroi, Myosotis Monroi, and Euphrasia Monroi.
And these names represent only a small portion of the species which he added
to the flora of his adopted country at various times between the years 185)
and 1870.
Aciphylla Monroi, the species figured on this plate, was discovered in the
year 1853 on the “summit of Macrae’s Run, alt. 4,500 ft.” I am not quite
certain of the exact position of this locality, but believe it to be in the Awatere
Valley, in which a tract of pastoral land was held by a Mr. Macrae, many
portions of which are known to have been frequently botanized over by Monro.
The species was first published in a list of addenda printed at the close of the
second volume of the “ Flora Novee Zelandie.” In the ‘* Handbook” Sir J. D.
Hooker gives several additional localities for the plant, and many others have
been recorded by subsequent observers. But I have long considered it highly
doubtful whether the whole of these habitats have been correctly assigned to
A. Monrov; and Mr. W. B. Hemsley, who has kindly examined the whole of
the specimens referred to the species in the Kew Herbarium, is of the same
opinion. Under these circumstances, I have decided to figure Monro’s original
specimens, preserved at Kew, so that workers in the Dominion may, at any rate,
be sure of the type of the species.
From the above it will be readily understood that it is difficult to indicate
the precise range of A. Monroi, several of the recorded localities being referable
to forms which in my opinion should be excluded from the species, while in
other cases I have had no opportunity of examining specimens. It appears to
be most plentiful on the mountains of Marlborough and Nelson, but I have
seen examples in several stations in Canterbury, and have reason to believe
that it also occurs in Otago.
Puate 63. Aciphylla Monroi, drawn from specimens collected on “ Macrae’s Run” by Sir David
Monro. Fig. 1, tip of leaflet; 2, bracts; 3, flower; 4, the same with the petals removed ; 5, petal ;
6 and 7, front and back view of anthers. (All enlarged.)
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Flate 64.
West, Newman imp-
M. Smith del
J.N-Fitch hth.
COXELLA DIEFFENBACHI, Cheesem & Hemsl.
PLATE 64.—COXELLA DIEFFENBACHII.
Famity UMBELLIFER 4. ] [Genus COXELLA, CHEESEM.
Coxella Dieffenbachii, Cheesem. in Trans. N.Z. Inst. xliv (1912), 160.
Ligusticum Dieffenbachii, Hook. /. Handb. N.Z. Fl. 729.
Aciphylla Dieffenbachii, 7. Kirk, Students’ Fl. 211.
The remarkable plant figured on this plate was discovered by Mr. H. H.
Travers in 1863 during his first visit to the Chatham Islands. The collections
formed on that occasion were placed in the hands of the late Baron Mueller, and
were worked out in his little book entitled ‘“‘The Vegetation of the Chatham
Islands,” published in 1864. Our plant was described and figured, but evidently
from imperfect material, under the name of Gingidiwm Dieffenbachii, the specific
name being intended to commemorate the services to botanical science of
Dr. Ernst Dieffenbach, who was the first to examine the botany of the group.
According to Mueller, the plant was gathered “in damp places under cliffs at
the seashore”?; but he gives no information as to whether it was abundant
or rare. I understand, however, that it was observed in small quantity during
Mr. H. H. Travers’s second visit in 1871, although I have seen no specimens
collected at that time. No further information was obtained until about 1895,
when Mr. F. A. D. Cox forwarded flowering and fruiting specimens to Mr. Kirk.
These formed the basis of the description given in his “ Students’ Flora” and of
mine printed in the Manual. Since then, through the kind action of Mr. F. A. D.
Cox and Captain Dorrien-Smith, I have received a much better series of specimens,
from which the accompanying plate has been prepared.
Previous writers have held very diverse views as to the systematic position of
our plant. As already stated, Mueller associated it with Forster’s Gingidium ;
Hooker transferred it to Ligusticum; while in the “Index Kewensis” it was
placed in Angelica. Mr. Kirk disposed of it in Aciphylla, making the remark that
“it will probably form the type of a new genus.” This latter view I believe to
be correct. It differs from the typical Ligustica in the flattened and conspicuously
winged fruit, one carpel being 3-winged and the other 2-winged, or very rarely both
carpels are 3-winged. The vitte are unusually large, and are either 1 or 2 in the
interspaces, with 2 or 3 on the commisural face. From Angelica it is separated
by the equally winged fruit, by the smaller number of wings (or ribs), and by the
number being unequal in the two carpels. It has much of the habit of Aciphylla,
although the leaves and the bracts are never spinescent ; but differs in the flattened
and winged carpels, and in the smaller number of wings (or ribs) to each carpel,
to say nothing of the much larger vitte. Believing that it is best treated as a
separate genus, Mr. Hemsley and myself have described it under the name of
Coxella. This will commemorate the services to botanical science of Mr. F. A. D.
Cox, of Whangamarino, Chatham Islands. During a lengthened residence in this
remote outlier of the Dominion Mr. Cox has regularly and consistently collected
specimens of the flora of the islands, communicating them to most New Zealand
botanists, together with much valuable information. It is largely through his
assistance in supplying material that our present knowledge of the Chatham Islands
florula is in such a satisfactory position.
At the present time Cozella Dieffenbachii is exceedingly rare, if not tending
towards extinction, and is apparently confined to a few stations on the precipituous
slopes of cliffs in the southern portion of the islands. In an interesting paper
by Captain Dorrien-Smith, entitled “ An Attempt to introduce Olearia semidentata
2
into the British Islands,” printed in the Kew Bulletin for 1910 (pp. 120-126), and
which contains much information of value respecting the vegetation of the Chatham
Islands, Captain Dorrien-Smith gives an account of a visit to two of the few
localities known for Cozella, accompanying his remarks with an _ excellent
photograph of the plant in its natural habitat. He suggests that its present
scarcity is due to the depredations of cattle, which eat it readily wherever
they can get at it. A parallel case exists in Myosotedium nobile, which once
fringed the sea-coast for miles, but which now, through the combined attacks
of sheep and pigs, hardly exists in the wild state, although, fortunately, well
established in cultivation.
Puate 64. Cozella Dieffenbachii, drawn from specimens collected for Mr. F. A. D. Cox at Te Tuku,
Chatham Islands. A, male plant; B, female; C, fruit. Fig. 1, bracts (enlarged); 2, male flower
(x7); 8, same with petals and stamens removed (x7); 4, petal (x7); 5 and 6, front and back
view of anthers (x 8); 7, female flower (x7); 8, the same with the petals removed, showing the
sterile stamens (x 7); 9, the same with the stamens removed also (x7); 10, petal (x10); 11, fruit
(x 2); 12 and 13, transverse sections of fruit (x 7).
Plate 65.
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M. Smith del.
J.N-Fitch lith.
LIGUSTICUM HAASTII, & Muedd.
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PuatTeE 65.—LIGUSTICUM HAASTII.
Famity UMBELLIFER. ] [Genus LIGUSTICUM, Linn.
Ligusticum Haastii, 7. Muell. ex Hook. f. Handb. N.Z. Fl. 95; Cheesem. Man. N.Z. Fl. 217.
Ligusticum Haastii is another prominent alpine plant discovered by Sir Julius
von Haast during his many explorations in the Southern Alps. So far as I can
ascertain, his first specimens were collected in the summer of 1861, on the mountains
flanking the Rangitata and Ashburton Valleys. He appears to have forwarded
some of them to Baron Mueller, of Melbourne, who suggested the name under which,
a few years later, the species was described by Sir J. D. Hooker. Subsequent
investigators have found that it is widely distributed in the central portions of
the Southern Alps, particularly in the districts subject to the large western rainfall,
where it is a prominent constituent of the subalpine meadows. It is most plentiful
at an altitude of about 3,000 ft., but descends as low as 1,500 ft. in several localities,
and I have seen it as high as 5,000 ft. Its northern limit appears to be on Mount
Stokes, near Pelorus Sound ; but as a rule it is rare and local in the north-eastern
portion of the South Island. In the Nelson Provincial District I have not myself
seen it further north than the mountains between the Clarence and the Waiau ; but
on the western side of the Island it is more frequently seen—in fact, Mr. Townson
informs me that it is a most abundant plant on all the coastal mountains as far as
Mokihinui, and possibly farther north.
L. Haastii is related on the one side to LZ. Lyall, and on the other to
L. brevistyle, but is abundantly distinct from both. From the first it can be
separated at a glance by the much more finely cut membranous leaves, the
segments of which are bristle- pointed. The latter differs in its much smaller size,
more slender habit, sparingly divided leaves, and very short styles.
In another publication (** The Subantarctic Islands of New Zealand,” vol. ii,
p- 408) I have mentioned that in the “Genera Plantarum” Mr. Bentham has
transferred to Aciphylla all the southern species referred by Hooker to Ligusticum,
and recently several botanists have followed his lead. No doubt the floral and
carpological characters of the two groups are very similar; but, on the other
hand, the habit of the true Aczphylla, with their rigid and coriaceous spinous
leaves, and paniculate inflorescence protected by spinous bracts, is most distinctive.
I feel sure that Mr. Bentham would never have made the proposal if he had been
familiar with Aciphylla in a living state. In any case, the two groups would have
to be separated as subgenera, which is practically the same arrangement under
another name without its conveniences. If the southern plants placed by Hooker
in Ligusticum are to be excluded from that genus (and it may be remarked that
the numerous vitte of the typical Ligustica constitute a marked difference), then
Hooker’s genus Anisotome should be revived for the reception of the greater portion
of them.
Prare 65. Ligusticum Haastii, leaf, and the inflorescence from male and female plants, drawn
from specimens collected in the Mount Cook district, Canterbury, at an altitude of 4,000 ft. Fig. 1,
male flower, with the petals and stamens removed (x6); 2, male flower, complete (x 6); 3, petal
(x6); 4, female flower (x 6); 5, the same with the petals removed (x 6); 6, fruit (x5); 7, trans-
verse section of same (x 5).
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Plate 66.
LIGUSTICUM CARNOSULUM, Hook.f
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PLATE 66.—LIGUSTICUM CARNOSULUM.
Famity UMBELLIFER. ] [Genus LIGUSTICUM, Liny.
Ligusticum carnosulum, Hook. /. Handb. N.Z. Fl. 96; Cheesem. Man. N.Z. Fl. 219.
On most of the higher mountains of New Zealand immense slopes of dry
rocky debris occur, which are locally known as “shingle-slips.” In not a few
localities these cover the sides of the mountains from base to summit, and
present an aspect almost indescribably bare, barren, and desolate, apparently
devoid of all vegetation. But on closer examination it is seen that they
support a peculiar flora, composed of plants of very diverse relationships—
buttercups, chickweeds, clove-worts, umbellifers, composites, figworts, grasses,
and many others. Most of these are of small size and low stature, while all
have long and often tortuous roots. Some are succulent, others are covered
with a felted mass of hairs, and nearly all are a peculiar gray-green, closely
matching the colour of the shingle. From this assemblage of highly specialized
plants it would be difficult to select one more singular than the subject of this
plate.
Ligusticum carnosulum was first collected by Sir Julius von Haast on
shingle-slopes on Mount Torlesse in 1861, and has been observed in the same
locality by many other botanists. It is, however, by no means a common
plant, and is fast becoming scarcer through the combined attacks of sheep and
rabbits. The most northern station known to me is on the mountains flanking
the Wairau Valley, Nelson, where I gathered it many years ago. Mr. Kirk has
collected it on shingle-slopes on Mount Captain, in the Amuri district ; and
during a hasty visit paid by myself to Lake Tennyson some years ago I noticed
it in some quantity. As far back as 1864 or 1865 Mr. J. D. Enys collected it
on the Black Range and other mountains of the Middle Waimakariri basin,
where it has since been observed by myself and others. It was gathered by
Mr. Armstrong in the Rangitata district, where it has also been noticed by
Dr. Cockayne and Mr. Laing; but I know of no locality farther south, although
I have specially searched for the plant on the mountains near Lake Tekapo,
also in the Mount Cook district and other localities in southern Canterbury.
Its altitudinal range appears to be from 3,000 ft. to 6,000 ft.
The nearest ally of L. carnosulum is undoubtedly my L. diversifolium
(Manual, p. 1139). This has the same habit and foliage, and the flowers and fruit
are almost indistinguishable. But the latter plant differs widely in the involucral
leaves, which are linear, entire, and much shorter than the umbel; whereas (as
will be seen from the plate) they are ternately multifid and much longer than
the umbel in ZL. carnosulum. It may be remarked that L. carnosulum and
L. diversifolium differ widely from all the other Ligustica and Aciphylla@ hitherto
found in New Zealand in the fruit, which is not at all winged, but simply
furnished with low rounded ridges (see figs. 5, 6, and 7). When the southern
species of Umbellifere are systematically worked up I anticipate that these two
species will find a home in another genus.
It has been pointed out by Diels and others that ZL. carnosulum has been
largely modified to suit the peculiar nature of its environment. The long and
fleshy tortuous rootstock, with its stringy roots, is plainly adapted to suit the
never-ending combat the plant has to wage with the moving shingle. The
leaves are finely divided, so as to present as little surface as possible to the
2
rain of shingle-dust which in high winds must often fall upon them, while the
divisions themselves are unusually elastic, so that Diels has compared them to
tiny indiarubber tubes. The protection afforded to the flowers and ripening
fruit by the involucral leaves within which they are enclosed is another adaptation
of considerable importance.
Pirate 66. Ligusticwn carnosulum, drawn from specimens collected on the Black Range, Middle
Waimakariri, Canterbury Alps, at an altitude of 4,000ft. Fig. 1, inflorescence and involucral
leaves (x 2); 2, male and female flowers (x 5); 3, petals (x 8); 4, female flower, with petals removed
(x 8); 5, fruit (x 5); 6 and 7, transverse sections of a single carpel (x 8).
=
Plate 67.
West, Newman imp.
M. Smith del.
JN, Fitch lith.
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PLATE 67.—LIGUSTICUM PILIFERUM.
Famity UMBELLIFER 4. | [Genus LIGUSTICUM, Linn
Ligusticum piliferum, Hook. /. Handb. N.Z. Fl. 96; Cheesem. Man. N.Z. Fl. 220.
Lagusticum piliferum was first discovered by Sir Julius von Haast in 1861
on Mount Torlesse, well known to the residents of Christchurch as the nearest
prominent outlier of the Southern Alps. As it attains a height of nearly
6,500 {t., and stretches from the Waimakariri River to Porter’s Pass, it is high
enough and massive enough to present many diversified plant-stations of rock,
shingle-slope, and subalpine meadow; and hence no small number of alpine
plants first became known from specimens obtained from such a convenient and
accessible collecting-ground. In the case of L. piliferum, it was soon found in
other localities. Haast obtained it in several additional stations in the Southern
Alps; Hector and Buchanan gathered it in the Lake district of Otago; Travers
in the Hurunui Valley and other localities in northern Canterbury. Since then
it has been found by Mr. Macmahon on Mount Stokes, near Pelorus Sound, and
by myself on the Mount Arthur Range, in north-west Nelson, these two stations
constituting its northern limit. Roughly speaking, it can be said to be an
abundant mountain - plane from thence to the south-west of Otago, chiefly
between the elevations of 2,500 ft. and 5,000 ft.
L. piliferum is a very distinet species, and I do not know any plant that can
be said to be closely allied. Its stout fleshy stems, bold foliage, and large umbels
of white flowers make it a very conspicuous and showy plant, and it is a great
ornament to the rocky banks and ledges on which it delights to grow.
Pirate 67. Ligusticum piliferum, drawn from specimens collected on Mount Arthur, Nelson,
at an elevation of 4,500 ft. Fig. 1, flower (x 7); 2 and 3, the same with the petals and stamens
removed (x7); 4, petal (x7); 5 and 6, front and back view of anthers (x7); 7, fruit (x 12);
8, transverse section of fruit (x 12).
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Plate 68.
Wee 4
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M. Smith del.
J.N.Fitch lit
ANGELICA GINGIDIUM, ook. f.
West, Newman imp.
PLATE 68.—ANGELICA GINGIDIUM.
Famity UMBELLIFER.] [Genus ANGELICA, Linn.
Angelica Gingidium, Hook. /. Handb. N.Z. Fl. 97: Cheesem. Man. N.Z. Fl. 222.
Ligusticum Gingidium, Vorst. Prodr. n. 140.
This well-known plant was one of those discovered during Cook’s first
voyage, having been collected by Banks and Solander in Queen Charlotte
Sound in January, 1770. Dr. Solander described it in his ‘ Primitize Flore
Nove Zelandize”’ under the name of Ligusticum anisatum, and an excellent
drawing was prepared for the Banksian plates. But, as already explained in
this work, Solander’s names were never actually published, and consequently
have no standing in systematic botany. It was again collected by the Forsters
during Cook’s second voyage, and was published in their ‘‘ Characteres Generum
Plantarum” (p. 41, t. 21), with an excessively crude and imperfect plate, as
the type of a new genus under the name of Gingidiwm montanum. George
Forster, in the later ‘* Prodromus ” (p. 22), abandoned Gingidium as a separate
genus, and referred the plant to Ligusticwm as L. Gingidium. Hooker, in the
“Flora Nove Zelandix ” (i, 89) included it in his genus Anisotome, but in the
“Handbook” (p. 97) transferred it to Angelica, where it has since remained.
From the above it is obvious that montanum is the oldest specific name, and
a purist in nomenclature would undoubtedly call the plant Angelica montana.
But I think it can be reasonably objected that the authors of the first specific
name, not long alter its publication, themselves proposed the substitution of
the second name, and that at the present time nothing but confusion could
result from the change.
A. Gingidium has a wide distribution. In the North Island, however, it
is far from common, attaining its northern limit, so far as I am aware, at
Marikopa, a little to the south of Kawhia, where it was observed a few years
ago by Mr. EK. Philips Turner. In the South Island it is (or rather was) a
very common plant, stretching from Nelson to the south of Otago, and with an
altitudinal range stretching from sea-level to quite 4,000 ft. It is one of the
tew New Zealand species that show a marked preference tor calcareous soils,
although by no means restricted to such localities. It is greedily sought after
by cattle and sheep, with the natural result that it is now comparatively scarce
in districts where it was once abundant. The whole plant is strong-scented
and aromatic, and is generally known in country districts in the South Island
by the name of “aniseed,” from the resemblance of its odour to that of the
well-known Anise. It may be remarked that Solander’s specific name of
anisatum was applied for the same reason. He says “‘semina ut et tota planta
aromatica sapore Foeniculi dulcis, odore seminum Anisi.”
Prare 68. Angelica Gingidium, drawn from specimens gathered by Mr. F. G. Gibbs, in the Maitai
Valley, Nelson, and in the Mount Cook district by myself. Fig. 1, male flower (x 8) ; 2, male flower,
with the petals and stamens removed, showing the calyx and rudimentary ovary (x 8); 3, female
flower (x 8); 4, the same with the petals removed (x8); 5, petal (x 8); 6, fruit (x6); 7, trans-
verse section of fruit (x 12).
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Plate 69.
West, Newman imp.
M. Smith del
J.N.Fitch lith.
ANGELICA ROSAEFOLIA, Zook.
PLATE 69.—ANGELICA ROSAZFOLIA.
Famity UMBELLIFER. | [Genus ANGELICA, Linn.
Angelica rosefolia, Hook. Ic. Plant. t. 581; Cheesem. Man. N.Z. Fl. 224.
For the discovery of this curious plant we have to go back as far as October,
1769, the time of Cook’s first visit to New Zealand. It was then collected by
Banks and Solander on or near the site of the present town of Gisborne, and
a few days later was observed at “ Tigadu”’ (Anaura Bay) and at Tolaga Bay.
It was also gathered by them at Opuragi (Mercury Bay), and on Motuarohia
Island, in the Bay of Islands. An excellent description was written by Solander
for his manuscript “ Primitive Flore Nove Zelandizw” under the name _ of
Ligusticum aromaticum, and a very faithful drawing was prepared for the
Banksian plates. Notwithstanding all this, Solander’s name and drawing both
remained unpublished, and the plant was not mentioned by any of the earlier
writers on the New Zealand flora, as Forster, A. Richard, and Allan Cunningham ;
in fact, it was not until seventy-five years had elapsed that it again came
under the notice of the botanical world.
Some time before 1843 specimens were obtained in some locality in the
North Island by Mr. Colenso, and forwarded to Sir W. J. Hooker at Kew. Soon
after, the plant was also gathered in the East Cape district by Dr. Sinclair,
and from these two sources it was figured and described in the “ Icones
Plantarum ” (t. 581) under its present name. Sir J. D. Hooker, in the “ Flora
Nove Zelandie ” transferred it to his genus Anisotome, but in the ‘** Handbook”
he replaced it in Angelica. In neither publication does he mention Banks and
Solander as the first discoverers—in fact, through some curious misconception
he applied Solander’s name of Ligusticum aromaticum to the mountain-plant
which is now well known under that appellation. It is hard to see how the
mistake can have arisen, for neither Solander’s description nor plate will suit
the present LZ. aromaticum, and there is no proof that he ever collected it.
Fortunately, however, as Solander’s name was never actually published, its use
for a different species is admissible; so that there is no occasion to disturb
the existing nomenclature.
Angelica rosefolia is mainly a coastal plant, and I have not myself seen it
any great distance from the sea. Its northern limit is on the Three Kings
Islands, where I observed it in 1887 and again in 1889. From thence it is
not uncommon southwards as far as Hawke’s Bay on the east coast and Raglan
on the west. Inland it has been recorded from the Ruahine Range by
Mr. Harding, and the Upper Rangitikei by Mr. Buchanan. I have not seen
specimens from the South Island, but it has been recorded from Akaroa (Raoul,
fide Hooker) and from Canterbury, without precise locality, by Travers and
Armstrong.
This species and the allied A. geniculatus are anomalous in the Umbellifera
from their scrambling or subscandent stems. The leaflets are also provided
with a pair of minute stipelle at the base: one on the upper surface, the
other below—a very unusual character in the family.
Pirate 69. Angelica rosefolia, drawn from specimens collected on the Little Barrier Island by
Miss Shakespear. Fig. 1, pedicel (enlarged) ; 2, base of a pair of leaves, showing the minute stipelle
(enlarged) ; 3, male flower (x 10); 4, the same with the petals and stamens removed, showing the
calyx and the rudimentary ovary (x 10); 5, petal (x 12); 6, female flower (x 10); 7, the same
with the petals removed (x 10); 8, petal from female flower (x 12); 9 and 10, fruit (x 8); 11, trans-
verse section of fruit (x 14).
ined,
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Plate 70.
G@ ee EN
fa
Be
West Newman imp.
M. Smith del.
J.N.Fitch lith.
STILBOCARPA LYALLII, Azmsér
PLATE 70.—STILBOCARPA LYALLII ann STILBOCARPA
POLARIS.
Famity ARALIACE:.] [Genus STILBOCARPA, A. Gray.
Stilbocarpa Lyallii, Armstr. in Trans. N.Z. Inst. xiii (1881), 336.
Aralia Lyallii, 7. Kirk in Trans. N.Z. Inst. xvii (1885), 295.
Stilbocarpa polaris, A. Gray, Bot. U.S. Expl. Exped. 714; Cheesem. Man. N.Z. Fl. 227.
From the point of view of geographical distribution the genus Stilbocarpa
is one of the most interesting found in the New Zealand area. It has no near
allies; and although in technical characters it approaches Aralia on the one
side, and the herbaceous species of Panax on the other, the affinity which it
possesses with either is obscure and inconclusive. At the same time, these
affinities appear to be sufficient to preclude the possibility of the genus having
developed anywhere close to its present area of distribution. Consequently, the
problem of what its ancestry has been, from what lands it has wandered, and
how it arrived in its present habitat, become questions of some moment; and
although answers cannot be provided at the present time, we may hope that
patient inquiry and observation may yet supply them.
The discovery of Stelbocarpa dates back to 1840, when S. polaris was collected
on the Auckland Islands successively by Dr. Holmes, of the American Exploring
Expedition; by Admiral D’Urville, the commandant of the French ships ‘ L’ Astro-
labe’ and ‘La Zélée’; and by Sir J. D. Hooker, who, as is well known, accom-
panied the Antarctic expedition of Sir James Clark Ross. It was made known to
the scientific world in 1843, under the name of Aralia polaris, through a beautiful
plate issued in advance of the ‘“ Botany of the ‘ Astrolabe’ and ‘ Zélée.’”
Through a succession of mishaps, however, the letterpress for this and other
plates did not appear until ten years later. In the meantime, it was figured
by Hooker in the “Icones Plantarum”; and his description given in the
“Flora Antarctica,” published in 1844, was the first good account to appear.
In the “Flora Nove Zelandiz,” issued in 1853, Hooker expressed the opinion
that the plant was “certainly of a different genus from Aralia proper, but in
the present confused state of the natural order it is not expedient to separate
it in this local flora on grounds which on extensive examination of the order
may prove not to be the legitimate ones.” Further on he proposed the
sectional name of Stilbocarpa, in allusion to the shining fruit. Dr. Asa Gray,
writing in 1854 in the “ Botany of the United States Exploring Expedition ”
(vol. i, p. 714), definitely established Stibocarpa as a distinct genus, pointing out
that the acetabuliform fruit and styles fewer than the petals must exclude the
plant from Aralia.
All visitors to the Auckland and Campbell Islands agree in stating that
S. polaris is an abundant plant, ranging from sea-level to a considerable height
on the hills. In 1880 Professor J. H. Scott collected it on Macquarie Island,
remarking that it “‘is found all over the island growing in large patches ” ;
and Mr. A. Hamilton also obtained it in his later visit of 1894. In 1890
Mr. T. Kirk recorded it as plentiful on Antipodes Island. The physiognomy of
the plant has been. so often described that no detailed account need be given
here. As Hooker has well remarked, it is “‘one of the most handsome and
singular of the vegetable productions in the group of islands it inhabits, which
certainly contains a greater proportion of large and beautiful plants, relatively
to the whole vegetation, than any country with which I am acquainted.”
Stilbocarpa Lyallii was originally discovered in Stewart Island by Dr. Lyall
in the year 1848. Both in the Flora and the ‘“ Handbook” Sir J. D. Hooker
included it in his circumscription of S. polaris, although in the last-mentioned
publication he said that ‘“‘Lyall’s Southern Island specimen wants the long
bristles, and may belong to another species.” In 1881 Mr. J. B. Armstrong,
2
who had cultivated Lyall’s plant side by side with S. polaris in the Christchurch
Botanical Gardens, described it as a distinct species, relying principally on the
upper surface of the leaves being nearly-glabrous and devoid of the long and
stout bristly hairs so conspicuous in 8. polaris. In 1885 Mr. T. Kirk, who had
good opportunities of studying S. Lyallii in a wild state in Stewart Island,
pointed out that it departed from the characters of Stilbocarpa in the 2-celled
ovary and the globose fruit not hollowed at the apex. He therefore considered
that it should be transferred to Avalia, and this view was maintained in the
“ Students’ Flora.” In 1898 Dr. Harms, in “ Die Naturlichen Pflanzenfamilien,”
restored it to Stilbocarpa, placing it in a separate section, to which he applied
the name of Kirkophytum. At the present time this view is generally accepted
by botanists who are familiar with the plant.
Stilbocarpa Lyallii, according to Kirk and Cockayne, is fairly abundant as a
coastal plant in Stewart Island and on the small islands in Foveaux Strait.
Mr. Kirk has also recorded it from Coal Island, in Preservation Inlet; but as it
was “identified from the deck of a passing steamer” a slight element of doubt
attaches to this locality. While very similar to S. polaris in habit and appearance,
S. Lyallic differs markedly in possessing strongly arcuate stolons sometimes
almost 3ft. in length. According to Mr. Kirk, “these at first are suberect,
but gradually become inclined or arched until the terminal bud comes in contact
with the ground, when roots are given off, and a new plant is speedily developed.”
In this way a single plant soon forms a large patch covering several square yards.
The differences in the glabrous under-surface of the leaves and the 2-celled ovary
have already been mentioned.
A third species of Stilbocarpa is the plant discovered on the Snares by
Mr. Kirk in 1890, and described by him under the name S. Lyalli, var. robusta.
He states that it is ‘the most striking herbaceous plant on the island,” and that
“it is sometimes 3 ft. high or more, with noble orbicular reniform leaves over 2 ft.
in diameter. It differs from the typical form in wanting the remarkable stolons
of that plant, in the petioles being very stout, flat on the upper surface and concave
beneath, giving a plano-convex section, and in their being nearly solid imstead
of terete, thin-walled, and fistulose. The flowers also, although forming equally
large masses with the type, are individually smaller, and invariably of a dull pale-
yellow hue, never lurid; but there is no structural difference, although it must
be admitted that at first sight the plant appears to depart widely from the type.”
Notwithstanding the opinion thus expressed by its discoverer, it appears to me
that the plant fully deserves the rank of a species. This is also the opinion of
Dr. Cockayne, who has had opportunities of examining it in its native home, and
who has proposed for it the name of S. robusta.
The following synopsis of the species of Stidlbocarpa may be of service :—
A. Stiupocarpa. Ovary 3-4-celled. Fruit hollowed at the apex.
Stolons not developed. Upper surface of leaves more or less clothed with stout bristly
hairs. Flowers yellowish-green with a purplish centre D6 5c .. L. S. polaris.
B. KirkopHytum. Ovary 2-celled. Fruit globose, not hollowed at the apex.
Stolons not developed. Upper surface of leaves glabrous or nearly so. Flowers
yellowish - Be a at ae S ah .. 2. 8. robusta.
Arcuate stolons strongly developed. Upper surface of leaves glabrous or nearly so.
Flowers reddish-purple .. Ra Se be on ay .. 3 S. Lyalla.
Prats 70. ‘The leaf that of Stilbocarpa Lyallii ; the inflorescence and dissections from S. polaris.
Specimens from plants cultivated in the garden of the late Mr. H. J. Matthews, Dunedin. (Nore.—lIt
was intended to figure S. Lyallit alone, but through an accidental mistake the inflorescence of S. polaris
was sent to the artist.) Fig. 1, flower-bud (x 5); 2, male flower (x 5); 3 and 4, anthers, front and
back view (x5); 5, female flower (x5); 6, longitudinal section of same (x5); 7, fruit, nearly
ripe (x 5); 8, longitudinal section of ripe fruit (x 5); 9, seed (x 8); 10, longitudinal section of same.
Plate 71.
cia
Weiss ee
ASS ee : = 2 A
\ ‘ | = a } WE : eS AY
\ : \S Je VW) ROSS
M.Smith del. West, Newman imp.
J.N. Fitch lith.
PANAX LINEARE, Hook.f.
PLATE 71.—PANAX LINEARE.
Famity ARALIACE/..] [Genus PANAX, Linn.
Panax lineare, Hook. {. Fl. Nov. Zel. i, 93; Cheesem. Man. N.Z. Fl. 228.
For the discovery of Panax lineare we are indebted to Dr. Lyall, the surgeon-
naturalist to H.M.S. ‘ Acheron,” who in 1848 collected it in Chalky Bay, on
the south-west coast of Otago. His specimens—only two in number—were
forwarded to Sir J. D. Hooker, and were described by him in the “ Flora
Nove Zelandiz ” (vol. i, p. 93); but as Lyall only obtained the female flowers,
and did not notice ier very different foliage of the young plants, the account
given was necessarily imperfect. In 1863 Sir James Hector and Mr. Buchanan
sathered it in Dusky Sound; but it was not until 1876, when it was found by
Mr. J. D. Enys and Mr. T. Kirk in the Upper Waimakariri, Bealey Gorge, and
adjacent districts, that flowers of both sexes were obtained, and a series of
specimens collected showing the remarkable changes the foliage passes through
during the growth and development of the plant. In 1881 I observed it in
great abundance on the Mount Arthur Plateau, Nelson, and in the upper part
of the Takaka Valley. Since then it has been observed in many of the sub-
alpine forests along the central chain of the Southern Alps, especially on the
western side, the most southern locality known to me being Preservation Inlet.
It reaches an altitude of over 4,000 ft. in north-west Nelson, but descends to
1,500 ft. or lower on the coastal ranges near Westport. In the Sounds of the
south-west coast of Otago it probably comes down to sea-level.
A reference to the “plate will show that the foliage of the young: seedling is
very narrow-linear, and from 2in. to 3 in. in length by about +in. in breadth.
As the plant increases in size the leaves also increase in length, so that a
young tree 4{t. or 5{t. in height frequently has leaves 10in. in length by #in.
in width. In older stages the leaves gradually become shorter, so that in
flowering examples they are frequently less than 2in. long, with a breadth of
from din. to fin. The leaf-change through which the species passes thus resembles
that of Pseudopanax crassifolium var. unifoliolatum, but is not so conspicuous,
and the leaves of the young plant are never deflexed, as they usually are in
one of the stages of P. crassifoliwm.
But for the long recurved styles, and ovary usually (but not always) with
less than 5 cells, P. lineare might be referred to Pseudopanax, with the simple-
leaved species of which it certainly has some affinity. Pseudopanax crassifolium,
P. ferox, and P. chathamicum, together with Panax lineare, would form a very
natural group, considered from the point of view of the leaf-changes which take
place between the seedling and adult plant.
Pirate 71. Panax lineare, adult in the flowering stage, together with examples showing the changes
of leaf-form between the seedling and the mature plant ; the whole of the caveat ae collected
by Mx. F. G. Gibbs in the Cobb Valley, north-west Nelson. Fig. 1, tip of leaf (x 4); 2, flower-bud,
showing 4 petals (x 4); 3, male flower, showing 5 petals (x 4); 4 and 5, back and front view
of stamens (x 6); 6, female flower (x 4); 7, female flower, with two of the siotals removed, showing
the styles (x 6); 8, ete ee section of ovary (x 6); 9, female flower, with three styles, the petals
removed (x 6).
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Plate 72.
West, Newman imp.
PANAX ANOMALUM, 4ook.
PLATE 72.—PANAX ANOMALUM.
Famiry ARALIACE4.] [Genus PANAX, Linn.
Panax anomalum, Hook. in Lond. Journ. Bot. i (1843), 422; Cheesem. Man. N.Z. Fl. 230.
The typical form of this curious plant is one of the many interesting
discoveries made by Mr. Colenso during the early part of his long and active
residence in New Zealand. He first observed it in swampy forests by the
Waikato River in the year 1842, and specimens collected by him in that
locality were figured and described by Sir W. J. Hooker in the following year.
Since then it has been noted in many places between Kaitaia, in the extreme
north, and the valley of the Waikato. The var. microphyllum, which is the
form selected for illustration here, has a much more extensive and more southern
distribution. It was first collected by Mr. Bidwill in some locality in the
Nelson Provincial District, but is now known to range from the Middle Waikato
to Southland, with an outlying locality in the Rakiahua Valley, Stewart Island.
It occurs from sea-level to an altitude of 2,500 ft. or more. It differs from
the type chiefly in the smaller size, smaller and often narrower leaves, and more
glabrous habit.
Panax anomalum, as it was well named by Sir W. J. Hooker, is quite
unlike any other member of the genus. Its small size, setose branchlets, and
few-flowered umbels, often reduced to a single flower, are very remarkable
characters; but, in addition, its widely divaricating and often interlaced or
zigzag branches give it a most peculiar aspect, altogether different from that of
any of its allies, but very similar, so far as superficial characters are concerned,
to a considerable number of plants belonging to widely separated families, such
as Melicytus micranthus, Melicope simplex, Myrsine divaricata, Paratrophis hetero-
phylla, several Coprosme, and young states of Elwocarpus Hookerianus and
Pennantia corymbosa. Most of these agree so closely in the mode of ramifica-
tion, in the size, shape, and even colour of the leaf, and in the production of
small and usually unisexual flowers, that it is almost impossible to distinguish
them at a little distance; while dried specimens of several of them have been
confounded by excellent botanists. To my mind, all attempts that have been
made to explain why a number of plants of diverse relationships should have
assumed a form so closely similar that they can hardly be distinguished without
minute examination, or, in short, why they should apparently mimic a common
standard of growth and habit, have fallen far short of solving the difficulties of
the case. No sufficient reason has also been.given for the curious fact that so
many of these plants grow intermixed in large numbers, in some cases forming
special associations of their own, and that their chief habitat is in alluvial
soils or along the banks of rivers.
In the ‘ Pflanzenfamilien” all the New Zealand species of Panax are
referred to Miquel’s genus Nothopanax, a course which has been followed by
Dr. Cockayne. As explained in the Manual, I retain the species in Panax for
the present because the classification of the family is admittedly in an unsettled
condition, and because many changes are sure to follow when it is thoroughly
worked up. In all probability the New Zealand species at present included in
Panax (or Nothopanax) will ultimately find their way into more than one genus.
For instance, I can hardly suppose that the botanists of the future will retain
P. anomalum and P. arboreum in the same genus.
Pirate 72. Panax anomalum var. microphyllum, drawn from specimens collected at Norsewood,
Hawke’s Bay, and Foxhill, Nelson, adult and seedling. Fig. 1, unifohate leaf from mature plant (x 4) ;
2, fully expanded male flower and bud (x 8); 3, female flower, with the petals removed (x 8); 4 and 5,
front and back view of anthers (x 10); 6, transverse section of ovary (x 10); 7, ripe fruit (x 4);
8, longitudinal section of same, showing seeds (x 4); 9, transverse section of same (x 4); 10, embryo
(x 12).
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iM ie Pi\reiWNhOONK *- org comes uA hereviaie BAO bie orweiqall: larov sie: GPA
in wie Yo hy pie am Haga MOS Ci LL aon) aint &
ft : ’ *woltowlerny oY a ane. ti A a ala: a Misia ney eas i.
GP, pp Mniy uinitab a4 tf deeceanait me At dr dnd: (Erawak lncaeenian “VieveD ae Hecate .
a Pa Del iad, ivett be Were 7 statewgc Lath Liles | wat nt GEL) a ae woul)
_ anal ov al rede: Det Le Na Pati ovis ‘y stelg att: dyallones VO iTgnrrcdiayon.
avy FavilS biota yay { diiitnlo Serbeths ty. ald sala. D.tni & iw niekgey © uted
ne 5 tihink ys Re af itunes Vode tile pullinia viaie sep Ge odvalel
ps itived “ehp ve ape MIs Wad Vem. ally re cle neh RD muy
vtiitcrale sarivlon 1 onde Aahisilat PG) SUE lke OFM Ty Me Inala
7 “eh a) Aopars eet’? am hd Pi (yeu che (items SLO ea (re qsigiaey, gh
ao ae ane enue if Tilt ney Tat Ui Lbezhaiaitt “a DA i ak
7 wine hi a) fetid aids orate Yee hag .owy. que) Wi eatin Ja is he
| 7 ; > » ; ZA eiiariy ‘4. wala vt, ee allie
a
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a . 7S polly) sind: veal lithe A eh 43 i, alney w foup tl Ot ‘fpretatry
i owe i“ amuse aiid olaiet ‘tia 10 ttylalyge od sagatyd aul
* peer AA sy ‘elhedsytlin net raat if cathasl> NN) wqstetennl SURGE uly
dywnode Wy adh oalht of 4th ote aoe Yoin wba Tite Jonagebnon
us wouivnt Inset th seleeye Diet Wit. pit Ethel gh vty ie. yt yu ) eh
Seti!
pQM hay oe cally oe vwinti aw. F mangle ¥y, pe ay 4 a
> Sues Hin otitul ee Ai: ae! be . sale
RAS. 1,
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heb 48 ¢ pirvener cology ear ate spp ee Adal aesatoek ie sat revi Oldie Be
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eer (OMT Pete, 1) eerie wees iar hl DS |
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Plate 78.
CSI YK
West, Newman imp.
M. Smith del.
J.N.Fitch lith.
MERYTA SINCLAIRII, Seem.
PLATE 73.—MERYTA SINCLAIRIT.
(THE PUKA.)
Famiry ARALIACEA.] [Genus MERYTA, Forst.
Meryta Sinclairii, Seem. in Bonplandia, x (1862), 295; Cheesem. Man. N.Z. Fl. 232.
Meryta Sinclair, or puka, as it is generally called, is one of the most
remarkable of the New Zealand trees, and is certainly one of the rarest. It
was first observed by Mr. Colenso, who was shown by the Maoris a single tree
growing in a Native cultivation at Paparaumu, Whangaruru Harbour, which
they informed him had been brought from the Poor Knights Islands. It was
surrounded by a high fence, and was strictly tapu, Mr. Colenso not being
permitted access to it, or allowed to remove specimens. Major W. G. Mair
was the next to visit the locality, and succeeded in obtaining specimens of the
foliage, some of which were given to Dr. Sinclair. Later on Mr. Robert Mair
obtained ripe fruit, which was also forwarded to Dr. Sinclair. This material,
imperfect as it was, formed the basis of the descriptions given by Sir J. D.
Hooker in the “ Flora Nove Zelandiz,” and in the later published “ Handbook.”
About 1862 the tree was cut down by the Maoris, who resented the attempts
made by Europeans to obtain specimens from it; but shortly afterwards a
Mr. G. Henson discovered it in an indigenous condition on the Morotiri or
Chickens Islands. In 1869 Captain F. W. Hutton and Mr. T. Kirk made a
special visit to that locality, with the result of finding eight or nine plants.
A few years later I visited the islands and saw thirteen old plants and a few
seedlings. Mr. Reischek, who paid several visits to the Chickens while pursuing
his ornithological researches, and who was familiar with the whole group,
informed me that not more than thirty trees existed on the islands. He
observed, however, a solitary specimen on the north side of the adjacent Hen
Island. Mr. Sandager, who for several years was lightkeeper on the Fanal, or
Mokohinou Islands, states that it is not found thereon; and its reported
presence on the Poor Knights Islands, which has always rested on Maori
authority alone, has been rendered very improbable by the fact that it was
not observed during Dr. Cockayne’s recent visit (see Trans. N.Z. Inst. xxxvili
(1906), 353).
For many years the Chickens locality was the only one known. In 1889,
however, I discovered it in great abundance on the Western King, one of the
Three Kings Group, situated about thirty-eight miles to the north-west of Cape
Maria van Diemen. In this locality it constituted the greater portion of the
light bush which clothes the northern slopes of the island. Among the shrubby
plants associated with it were Myoporum letum, Melicope ternata, Leptospermum
scoparium and L. ericoides, Paratrophis Smaithii, and Cordyline australis. The
undergrowth was composed of Piper excelsum var. major, Hymenanthera nove-
zelandie, Asplenium lucidum, Pteris comans, and Davallia Tasmani, a plant-
association which cannot be matched in any other locality. During the same
visit I observed from the deck of the steamer that it was equally plentiful
on the top of the Eastern King; but as we failed to find any practicable way
of ascending the cliffs which everywhere surround the island I can say nothing
about the associated vegetation. Since then no fresh stations have been
discovered, and we may fairly conclude that the Hen and Chickens and Three
Kings Islands are the only localities for the species.
Zz
In a native state Meryta Sinclairw is a much-branched small tree from
12ft. to 20ft. in height, rarely more. It was introduced into cultivation by the
late Mr. Justice Gillies and Mr. G. B. Owen about 1870, and is now com-
paratively common in gardens in the vicinity of Auckland. It is easy of culti-
vation, perfectly hardy in most parts of the North Island, will bear exposure
to the strongest winds, and in good soil makes very rapid growth. A tree
planted in my own garden attained a height of 20ft. in less than ten years,
with a spreading crown of branches 16 ft. in diameter, and with a trunk 24 in.
in circumference at the base. Many of the leaves (including the petiole) were
quite 2 ft. 6in. in length.
New Zealand is the southern limit of Meryta, which is otherwise a purely
Melanesian and Polynesian genus. Its main home appears to be in New
Caledonia, from whence no less than eleven species have been described. Two
are known from Norfolk Island, one from Samoa and the Tongan Islands,
one from Rarotonga, two or three from the Society Islands, one from the
Caroline Islands, and one from New Guinea.
Puate 73. Meryta Sinclairii, drawn from specimens cultivated at Remuera, Auckland, showing
leaf and male and female inflorescence. Fig. 1, male flowers (x 4); 2 and 3, front and back view
of anthers; 4, female flowers (x 4); 5 and 6, transverse and longitudinal sections of ovary (x 4) ;
7 and 8, longitudinal and transverse sections of fruit (x 3); 9, longitudinal section of seed (x 6) ;
10, embryo (x 10).
Plate (4.
West, Newman imp-
- Smith del.
.N-Fitch lith.
PSEUDOPANAX DISCOLOR, farms.
PLATE 74.—PSEUDOPANAX DISCOLOR.
Famity ARALIACE. | [Genus PSEUDOPANAX, C. Kocu.
Pseudopanax discolor, Harms in Pflanzeur, t. ili, abt. 8, p. 46; Cheesem. Man. N.Z. FI. 233.
For the discovery of this fine plant we arefindebted to Mr. T. Kirk, who
in November, 1867, collected it on the Great Barrier Island, where it is
exceedingly abundant, ranging from sea-level to the tops of the highest hills ;
altitude 2,400 ft. It was referred by Mr. Kirk to Panax Sinelairii, which is in
reality a very different species, and was mentioned under that name in_ his
“Botany of the Great Barrier Island,” published in the first volume of the
“ Transactions of the New Zealand Institute” (p. 144); but in 1870 he corrected
the mistake, and described it as a new species under the name of Panar
discolor. In the meantime he had also observed the plant on the Little Barrier
Island; at Whangaroa (which appears to be its northern limit); at Omaha ;
and on the dividing range of the Cape Colville Peninsula, where it extends from
Cape Colville as far south as Tairua. In all these localities it is abundant ;
but no additional stations have been recorded of late years.
Although originally placed in the genus Panax by Kirk, and retained in it
in his “ Students’ Flora,” the plant really falls into Koch’s genus Pseudopanaz,
which differs from Panax in the 5-celled ovary and 5 styles, the latter being
very short and connate into a minute cone or column. It was therefore
correctly transferred to that genus by Harms in Engler and Prantl’s “* Pflanzen-
familien.” At the same time, it may be mentioned that Pseudopanaxr as at
present constituted contains two very different sets of species—one comprising
P. discolor and the allied P. Lessonii and P. Gilliesii; the other including the
species with strongly marked heteromorphous foliage, as P. crassifolium, P. ferox,
and P. chathamicum. These two divisions appear to me to deserve the rank of
distinct genera.
Pseudopanax discolor forms a handsome shrub 6 ft. to 15 ft. high, and is remark-
able for its peculiar bronzy or yellow-green foliage. Its nearest ally, as remarked
above, is undoubtedly P. Lessonii ; but that species is larger and much more
robust, with a very different habit of growth, and has dark-green coriaceous foliage,
the leaflets being entire or sinuate-serrate, and almost veinless. In P. discolor
the leaflets are thin, distinctly veined, and sharply and coarsely serrate. It
does well in cultivation, but is of slow growth.
Pirate 74. Pseudopanax discolor, drawn from specimens collected on the Little Barrier Island
by Miss Shakespear and myself. Fig. 1, flower-bud (x 4); 2, male flower (x 4); 3, male flower, with
petals and stamens removed (x 4); 4, stamen (x 6); 5, female flower (x 4); 6, the same with the
petals, &c., removed (x 6); 7, section of ovary (x 6); 8, ripe fruit (x 3).
A Sere AM +i
alt ats ye. ae :
Lh A asir gla
ny “ghd Pa
isi? ope bTetl er
lt Pratl! Lae we lis
; ni, arts oe i eo =
‘“ a “1 pcre tes) pagal tam io :
by began atin fre bea alae bf i
ty ut ore ae Dero ntns ile ® ahapy |y ~
+ 7
at ae ie
Plate 75.
E
‘
nt
A
3
=
PSEUDOPANAX CHATHAMICUM, 7 Airké.
PSEUDOPANAX CHATHAMICUM.
Famity ARALIACE.] [Genus PSEUDOPANAX, C. Kocu.
PuatE 75.
Pseudopanax ¢chathamicum, 7’. Kirk, Students’ Fl. 223: Cheesem. Man. N.Z. Fl. 236.
Pseudopanax chathamicum was one of the first plants obtained on the Chatham
Islands, for it was among those collected by Dieffenbach, the naturalist to the New
Zealand Company, who visited the group in 1840. His specimens were probably
imperfect, for otherwise Sir J. D. Hooker, who examined them while engaged in
the preparation of the “ Flora Nove Zelandix,’ would never have referred them
to the common New Zealand P. crassifolium. It was also collected by: Mar, Es H.
Travers, who explored the group in 1863; but apparently a fragment only of the
plant was brought back. In Mr. Travers’s second visit, made in 1871, a more
complete suite of specimens was obtained; for Mr. Buchanan, who prepared a
report on Mr. Travers’s collection which was published in the Trans. N.Z. Inst.
(vol. vii, p. 333), explicitly says that the Chatham Islands plant differed from the
New Zealand form in the leaves of the young plant never being deflexed at any
period of growth. In 1891 Mr. J. D. Enys paid a hurried visit to the group, and
on his return gave a series of specimens of the plant to Mr. Kirk and myself, at
the same time stating his conviction that it was specifically distinct from
P. crassifolium. Shortly afterwards Mr. F. A. D. Cox succeeded in supplying
Mr. Kirk with a fairly complete set of specimens, in all stages of growth, and
the plant was accordingly described in the ‘Students’ Flora” under its present
name,
As already stated, the chief differences between P. chathamicum and
P. crassifolium lie in the foliage of the young plants. Both P. crassifolium and
its near ally P. feroxr have a juvenile stage in which the leaves are very long,
narrow-linear, and conspicuously deflexed. As the plant approaches maturity
these linear deflexed leaves gradually become shorter, broader, and more erect ;
eventually passing into the comparatively short linear-oblong leaves of the adult.
In P. chathamicum, on the contrary, the leaves of the juvenile stage are
never much longer than those of the adult, nor much narrower, and are
never deflexed. The leaves of the mature plant are also larger and broader
than in P. crassifolium, and the fruit is much larger. These differences are
quite sufficient to prove the distinct nature of the species.
Pseudopanax chathamicum appears to be generally distributed in the wooded
portions of the Chatham Islands. Dr. Cockayne, in his paper on the “ Plant
Covering of the Chatham Islands” (Trans. N.Z. Inst. xxxiv (1902), 243 et seq.)
mentions it as an abundant factor in the ordinary lowland forest, in the forest
of the tableland, and in the somewhat different association found on limestone
soils. In the lowland forest he classes it as seventh in the order of most
frequent occurrence, the more abundant. plants being (1) Corynocarpus laevigata,
(2) Olearia Traversii, (3) Coprosma chathamica, (4) Hymenanthera chathamica,
(5) Myrsine chathamica, and (6) Corokia macrocarpa. On calcareous soils the
proportion of frequency is much higher (if I correctly interpret his remarks),
the Pseudopanax taking the third place.
For an account of the early development of the seedlings of P. chathamicum,
together with some remarks on its life-history, reference should be made to
Dr. Cockayne’s paper in Trans. N.Z. Inst. xxxii (1900), 88.
PLATE 75. Pseudopanax chathamicum, drawn from specimens supplied by Mr. F. A. D. Cox, ot
Whangamarino, Chatham Islands. Fig. 1, male flower (x4); 2 and 3, front and back view
of anthers (x8); 4 and 5, female (or hermaphrodite ?) flowers (x 4); 6 and 7, longitudinal and
transverse sections of ovary (x 6); 8, longitudinal section of fruit (x 4); 9, embryo (x 12).
Th
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AN Wabe te
ina ¢
od ~~ i j
) | i ‘ ' pkg la lh
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shen! Des ; ~ tj : ] ‘a ou "
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sa . rie } Des ii! ule Ali wor rahi meyte ia the ry aallithy is
wy ' ch 1 Maly rp! AwmitGe hh wi bale ieenint mand hi
7 radi ok ht | a ; i
et ere ag deuth, dele are! re 46 it Tp Mie
si 7 ® aa seat
; iat a why Tihihi at iy Sor .S; eu fii jtdgz
6) ow b & ih i ‘ oe
a ® rAvadats ath eye n ° TY a qt (Vy "
hh, htop winds) Dye
Vile - oer ayy din: 7%) i» VW i
Tl Wy wor gi) mmo em gay gn Yo mip ry yy my
spies nist vi! = (hath Ge? moreiy an
ov (ee ug valirws ls eh La itugline “Ab er ree. a
wn or@arahiag al th hel, PR pert hae aa
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gt Geis
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= . _
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ness rn
rt. =
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Plate 76.
West, Newman imp.
M. Smith del.
J.N.Fitch ith.
COROKIA BUDDLEOIDES, A. Cunn.
PuatE 76.—COROKIA BUDDLEOIDES.
Famiry CORNACE4. | [Genus COROKIA, A. Cunn.
Corokia buddleoides, A. Cunn. Precur. n. 579: Cheesem. Man. N.Z. Fl. 237.
The subject of this plate was first collected by that enthusiastic botanist
and explorer Allan Cunningham in the Bay of Islands district in the year 1826.
Recognizing that it constituted a new genus, Cunningham gave it the name of
Corokia, which is an adaptation of its Maori name korokio, or more correctly
korokio-taranga; and it was published under that title in his “ Precursor,”
issued in the years 1838-39. Cunningham attributed to his plant dicecious
flowers and white petals, both of which statements are erroneous, the flowers being
always perfect, and their colour yellow. The plant was collected by Mr. Colenso
shortly after his arrival in the colony, and an excellent drawing, prepared from
specimens forwarded by him, was published in the “Icones Plantarum” in the
year 1843. Since then it has been observed by all botanists who have examined
the vegetation of the northern portion of the North Island; in fact, it is common
in forests from Kaitaia and Mangonui southwards to the Bay of Plenty, and
probably to the East Cape, the most southern locality from whence I have seen
specimens being inland from Opotiki. It is most plentiful at a little elevation
above the sea, and was noted by Mr. Kirk on the summit of Mount Tarawera, prior
to the eruption of 1886, at an altitude of 3,400 ft.
Corokia buddleotdes is a handsome much-branched slender shrub 10 {t. or 12 ft.
in height. The pale-green leaves, with their silvery-white under-surface, and the
bright-yellow starlike flowers, render it an attractive plant, and it is somewhat
curious that it has not found its way into gardens in the northern portion of the
Dominion. It is easy of cultivation, of quick growth, and comes into bloom while
comparatively young. It might well be planted in the place of many far less
suitable exotics.
Prare 76. Corokia buddleoides, drawn from specimens collected on the Waitakerei Ranges, near
Auckland. Fig. 1, unexpanded flower (x 3); 2, expanded flower (x 3); 3, peltate hair (enlarged) ;
4, section of calyx and ovary (x 3); 5, portion of corolla laid open, showing the scales at the base of
the petals (x 5); 6 and 7, front and back view of anther (x 8); 8, scale (x 10); 9, ripe fruit (x 2) ;
10, longitudinal section of fruit (x 2).
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jet). oan) ii whl we bde wit Ault 7
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(AE) vi tyh ond bation ing
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TW Oe eho gts Grell nod lf Vip ate sh ‘elite, Ala retains gly
hic. ein tu ovilt, oe ua ree Ne ai
feata yal, 1h, conga e ai BY wih Oe em nat ofall Heh age: uw ddetiqg
tHulfieen’® ‘ileiiiew, 14 hed id jeune Nese dig)) on) Rit are mitt node
Swe Hehe | NUM we gin et tas OE Ags id eee ae Ad orodA
ore J, ME ti ahdle ny ‘Vee Teh rey wid cal
Neh Gehl hp ijnelbipalitodn', 2 a) eeeetO & NEL ak
ony The bgkutecvohet il Pty yerrin aed 9 a wel af What yal niyiod uw
Pe give a Bipreters, (moe oe See Nine? @ lang tidy hi .
: vii iy Sale lads vale alr vviny (mw ve Gta hier) seek awh Tt iit? ol recy
wih ane me ermine A ogaagy Te Vena bi Gt nil secivt fag /
ays! yet vite, rf Beige ee iy 61 | ie obo erage
euhiss oneliye
tegen i. a Byres eee een ae alana -4 yay
sire ey et Baling © fra) ut aa his A ; pr
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eet te Se " 1 pe ARA fa ret hina a ca Pie
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eihis vale datvngtgied’ bh) soir aved
Plate 77.
West, Newman. imp.
M. Smith del.
JN Fitch Lith
GRISELINIA LUCIDA, Forst.
PuatTE 77.—GRISELINIA LUCIDA.
Famity CORNACE. | [Genus GRISELINIA, Forsvt.
Griselinia lucida, Forst. Prody. n. 401; Cheesem. Man. N.Z. Fl. 238
The first specimens of this handsome tree were obtained in 1769 by Banks
and Solander at Tolaga Bay during Cook’s first voyage. Later on they also
gathered it at “‘Opuragi”’ (Mercury Bay) and “* Totaranui”’ (Queen Charlotte
Sound). Dr. Solander, in his manuscript “ Primitie Flore Nove Zelandiz,”’
recognized that it constituted a new genus, and proposed the name of Lissophyllum
lucidum ; but as his names were never actually published they have no standing
in systematic botany. The two Forsters, who also collected the plant in Queen
Charlotte Sound during Cook’s second voyage, published it under the name of
Scopolia lucida ; but, as this proved to be invalidated by the earlier Scopolia of
Jacquin, a genus of Solanacea, George Forster substituted Griselinia for it, a
name which has received general acceptance.
Griselinna lucida has a wide range, being found in lowland forests from the
North Cape to the Bluff, but is much more plentiful in the North Island than
in the South, where its place is largely taken by the alhed G. littoralis. It is
usually epiphytic or rupestral, and is often found growing amongst the immense
masses of Astelia and other epiphytes which clothe the spreading branches of
the rata (Metrosideros robusta). In such situations it does not attain any great
height, but its leaves are often unusually large, sometimes exceeding 8 in. in
length. It reaches its greatest stature when growing among or on rocks, or on
well-drained rocky soil, “where specimens can be found 25 ft. in height or even
more, with a trunk 18 in. to 24 in. in diameter. The finest rupestral specimens I
have observed were growing among huge masses of crystalline limestone at
Whangarei. It is a common plant on the basaltic lava-streams of the Auckland
Isthmus, and is particularly frequent on Rangitoto Island, where flowering
specimens can be seen of all sizes, from 3 ft. upwards. I cannot agree, however,
with Mr. Kirk in considering that it shows any unusual luxuriance of foliage in
that locality, or that the flowers are “of deeper and brighter tints than are
produced under ordinary conditions” (see Trans. N.Z. Inst. xi (1879), 450).
Griselinia lucida is one of the New Zealand trees which ought to be commonly
cultivated in gardens and shrubberies. It is easy of cultivation, and will grow
in any ordinary soil; it stands exposure to the strongest winds, and is of rapid
growth, while its pale yellow- -green smooth and glossy folage is most attractive.
These remarks also apply to the allied G. littoralis, which reaches a greater size,
but has smaller foliage and a more compact habit.
Puate 77. Griselima lucida, male, female, and fruit, drawn from specimens gathered on the
lava-streams at Mount Wellington, near Auckland. Fig. 1, flower-bud of male flower (x 6); 2, fully
expanded male flower (x 6); 3 and 4, front and back view of anthers (x 8); 5, female flowers (x 6) ;
6, section of same, showing the pendulous ovule (x6): 7, transverse section of ovary (x 6) ;
8, longitudinal section of fruit (x 4); 9, embryo (x 8).
,e~ ay a a aa a ’ —— 7
7
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f 1 cu 4 ron. 6 Th) ee fail oe howe in
5 Stee ti Lj witeiary f 4 ) Qreay sane tera a
a 2 elnanisy iy ‘Ty a ee Ae ct Ti a vy “A (ltd -
: > mpi dageged,! 6: Sanka 910 Bre ee Ay, are yp ap teetlig hag
: uti eta ea Mad vast 1 Mat sem ee” “veg ao enho@e Loo oon that NR
enut! &. bop oid 3 Wal els " way Ah al e770 etht sellin - sh: ot
vale, (Gtle atly iby umile Jang! tewele: is od eieliifoy ome,
bid Siddall warlhih sit ad letetods TUNE le Cena allay sy fue ee ae
Pw aust ea tity bid wiltiettd. avon ) ich ai pital fog MW ~
UTTER wor ty ‘ : bly J mice ieinsaerderr, 2000s wes 7
THB OMe. Srey wy, i ' alae noi a age we wid noe AN ’
ee ee ee if “ads ny, vind. pin nt t ifs] bat
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oi het Taal es ‘ slvadtmoll eee? Utils doen ins i (he) eevee Gee
od sewilabeye S . (2) a) abdten | A) ae bof 5 (a *\ fyi oni tga ty on
i = > (aes | Chane
4 me aw
Plate 108.
M. Smith del. . 2
JN Fitch lith. ' West, Newman imp.
COTULA PYRETHRIFOLIA, Hook f
PLATE 108.—COTULA PYRETHRIFOLIA.
Famity COMPOSIT/. | [Genus COTULA, Tourn.
Cotula pyrethrifolia, Hook. . Handb. N.Z. Fl. 142; Cheesem. Man. N.Z. Fl. 357.
Cotula pyrethrifoia is a very different species to the one figured in
Plate 106. The almost glabrous habit, the decumbent or prostrate stems, the
deeply pinnatifid leaves with their few linear lobes, and the large strictly
unisexual heads, mark it off from all the other species. Its only near ally is my
C. linearifolia, which also has unisexual heads together with a very similar
habit, but which differs in the much smaller size and entire leaves. All the
other species of the section with unisexual heads have very different foliage
and ebracteate scapes.
C. pyrethrifolia was originally discovered by Dr. Sinclair and Sir Julius von
Haast in the Rangitata Valley; but was quickly found to have an extensive
distribution in mountain districts in the South Island, both on the eastern
and western sides of the watershed. I have seen specimens from almost all
the important mountain districts, from the Mount Arthur Range, in north-west
Nelson, and Mount Stokes, in northern Marlborough, to Lake Wanaka, which
is the most southern locality of which I have any knowledge. Quite lately
it has been gathered by Mr. Petrie and Mr. Aston on Mount Hector, one of
the higher peaks of the Tararua Range, in the North Island. It is most
plentiful at an altitude of from 3,000 ft. to 4,000 ft., but I have observed it
at fully 6,000 ft. at the Wairau Gorge, and it descends to 2,500 ft., or perhaps
lower, in eastern Canterbury.
C. pyrethrifolia is usually found in sheltered places among rocks, or on
rocky slopes, when these are moderately stable. The finest specimens I have
ever seen were gathered by Mr. J. Adams and myself in the deep rocky
gorge just below the terminal face of the Waimakariri Glacier, at an altitude
of about 3,500 ft., where it was remarkably plentiful. Like many species of
the genus, it is highly aromatic, especially when bruised.
Puate 108. Cotula pyrethrifolia, drawn from specimens collected by the Broken River, Canter-
bury, at an altitude of 3,500 ft. Fig. 1, section of receptacle, with the involucral bracts surrounding
it (x 4); 2, flower-head (x 4); 3, female floret (x 10); 4, anthers from the same (x 15); 5, style-
branches from the same (x 15); 6 and 7, inner or sterile florets (x 10); 8, ripe achene, with the
corolla still attached (x 8).
Plate 109.
S)
=A)
M. Smith del.
West, Newman imp.
J.N.Fitch lith.
PUSILLA, Hook f 9-14.
A. ABROTANELLA LINEARIS, Bergg. 1-8.
BE
PLATE 109.—ABROTANELLA LINEARIS anp
ABROTANELLA PUSILLA.
Fammy COMPOSIT A. | [Genus ABROTANELLA, Cass.
Abrotanella linearis, Berggr. in Minnesk. Fisiog. Sallsk. Lund. viii (1877), 14, t. 3, f. 28-38 ;
Cheesem. Man. N.Z. Fl. 361.
Abrotanella pusilla, Hook. f. Handb. N.Z. Fl. 139; Cheesem. Man. N.Z. Fl. 362.
Abrotanella is one of those genera often called antarctic ; not on account of
inhabiting the Antarctic Continent, which in point of fact possesses only a single
phanerogam, but because their species are mainly found on the land masses which
approach most nearly to the Antarctic regions. Out of the fourteen species which
compose the genus, seven are known from the New Zealand area, ranging from the
Ruahine Range to the Auckland and Campbell Islands; three are found on the
Australian or Tasmanian mountains ; and three more occur either in Fuegia or on
the Falkland Islands. In addition to the above, a single isolated species has been
described from the Island of Rodriguez, to the eastward of Mauritius, being the only
representative of the genus found within the tropics.
A, linearis was one cf the discoveries made by Dr. 8. Berggren, of Lund, Sweden,
who in the years 1874 and 1875 travelled through the greater part of New Zealand
for the purpose of studying its vegetation. His specimens were collected on Kelly’s
Hill, on the western side of the Otira Valley, Westland, and were fully described
and figured by himself in a memoir issued under the auspices of the University of
Lund. A few years later it was collected by Mr. T. Kirk and myself in many localities
in the Provincial Districts of Canterbury and Nelson. It is now known to extend
from the Mount Arthur Range, in the north-west of Nelson, throughout the whole
length of the South Island, reappearing in Stewart Island, to the south of Foveaux
Strait. As a rule, it is most plentiful on the western side of the watershed of the
Alps. It ascends to quite 4,500 ft., and in the northern part of the South Island
is seldom seen below 2,500 ft. In Stewart Island it descends to sea-level. It is
usually found in damp sheltered places in subalpine meadows, or in open places
amongst scrub, and is very variable in stature and in the length of its leaves.
A. pusilla was collected in 1845 by Mr. Colenso in his first journey to the Ruahine
Range, one of the most adventurous expeditions made by Europeans during the
first explorations of the interior of the Dominion. His specimens, which were
gathered in “snowy places” on the summit of the range, were forwarded to Sir
J. D. Hocker at Kew, and were described by him in the ‘“‘ Flora Nove Zelandiz ”
under the name of Trineuwron pusillum. A few years later Professor Asa Gray showed
that Trineuron and the allied groups, to which Hooker had given the names of
Ceratella and Scleroleima, were not really separable from Abrotanella ; and in the
“Handbook” Sir J. D. Hooker reduced all three to that genus.
For more than sixty years Colenso’s plant was not gathered by any other botanist.
In the summer of 1907, however, Mr. Petrie and Mr. Aston made an ascent of Mount
Hector, the culminating-point of the southern portion of the Tararua Range, and
were fortunate enough to obtain a fine series of specimens of it, some of which have
been used in the preparation of this plate. Mr. Petrie remarks (Trans. N.Z. Inst.
xl (1908), 295), “In the more level spots considerable areas of shallow, haltf-peaty,
half-swampy soil are met with, and here the vegetation is very different. The most
abundant and most characteristic plant is Abrotanella pusilla. . . . It is a very
slender, low, densely matted, moss-like plant, with an inflorescence that barely
exceeds the leaves, carpeting quite a large proportion of all wet and boggy ground.
With it occur Astelia linearis (in fine fruit), Carpha alpina, Centrolepis viridis,
Plate 110.
West, Newman imp.
M.Smith del
J.N.Fitch lith.
ERECHTITES GLABRESCENS, Z Atrk.
PLATE 110.—ERECHTITES GLABRESCENS.
Famity COMPOSIT. | [GENUS ERECH TITES, Rar.
Erechtites glabrescens, T. Kirk in Trans. N.Z. Inst. ix (1878), 550; Cheesem. Man. N.Z.
Erechtites glabrescens was discovered by Mr. W. T. L. Travers in the Wairau
Valley, Nelson, prior to 1872 ; but was not described as a distinct species until 1876,
when Mr. T. Kirk conferred upon it the name it now bears. In the meantime it had
been observed by Mr. Kirk himself at Lake Rotoiti, Nelson, and by Mr. G. M. Thomson
in several localities in Otago. Further inquiry has shown that it has a tolerably
wide range, usually in mountain-forests or in scrub. In the North Island it has
been gathered by Mr. Petrie in the Upper Rangitikei, and by Dr. C ockayne and
myself on the central volcanic plateau; and it will probably yet be discovered in
the mountain - woods flanking the Ruahine and Tararua Ranges. In the South
Island it is found in most mountain-forests from Nelson to Foveaux Strait. It
attains its southern limit in Stewart Island, where it descends to sea-level, as it
also does in the south of Otago. In the North Island and in the Nelson mountains
it ascends to quite 4,500 ft.
L. glabrescens can, be distinguished without much difficulty from the other
species found in New Zealand. Its habit is more slender, and it is almost entirely
glabrous. The leaves are more membrancus, and vary excessively in size and in the
extent to which they are cut or divided. Occasionally all the leaves are coarsely
and irregularly sinuate-dentate, but more generally the lower ones are pinnate
with a large terminal leaflet and with few or many smaller lateral ones. The under-
surface is frequently of a pucplish colour. The heads are almost or entirely glabrous,
the involucral bracts bemg greenish with pale margins. The achenes are longer
than in any other New Zealand species, and are quite glabrous.
Prate 110. Erechtites glabrescens, drawn from specimens collected in the Mount Cook district,
at an altitude of 2,500 ft. Fig. 1, flower-head (x 4); 2, section of receptacle (x 4); 3, outer or female
floret (x 6); 4, inner or hermaphrodite floret (x 6); 5, pappus-hair (x 10); 6, anthers (6
;
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Plate 118.
M. Smith del.
J.N.Fitch lith.
West, Newman. imp.
SCAEVOLA GRACILIS, ook. f
PLATE 118.—SCAISVOLA GRACILIS.
Famity GOODENIACE/.. | [Genus SCANVOLA, Liv.
Scevola gracilis, Hook. /. in Journ. Linn. Soc. i (1857), 129; Cheesem. Man. N.Z. Fl. 395.
F)
Distant over six hundred miles from New Zealand, and surrounded by an
immensely deep ocean, lie the Kermadec Islands, a chain of four widely separated
islands stretching in a south-west to north-east direction. All are of small size ;
the largest, known by the name of Sunday Island, being under seven miles in length
by less than five in breadth. Their situation, rather more than half-way between
New Zealand and the Tongan Islands, gives special importance to them in any
inquiries respecting the origin of the New Zealand flora, and I am pleased to be
able to figure three or four of their endemic plants in this work.
Scavola gracilis, the subject of this plate, was first collected on Sunday Island
by Messrs. Milne and McGillivray, of H.M.S. “ Herald,” which, under the command
of Captain Denham, visited the group in 1854. In August, 1887, I accompanied
an expedition to the Kermadec Islands for the purpose of annexing the group to
the Colony of New Zealand, and formed a tolerably complete collection of its flora,
finding S. gracilis fairly plentiful in rocky places both on Sunday Island and
Macaulay Island. It was also gathered on Sunday Island by Miss Shakespear in
1904; and in 1907 by Mr. R. B. Oliver, who made a stay of nine months on the
island investigating its fauna and flora.
S. gracilis forms a procumbent undershrub from 2 ft. to 3 ft. or even 4 ft. high,
with long straggling branches more or less densely clothed with silky hairs. The
flowers are small, white with a yellow centre, sweet-scented, and are produced
abundantly throughout the year. The fruit is white, succulent, about 3 in. long.
Although usually found in rocky places, both on seacliffs and inland, it is by
no means confined to such. Mr. Oliver, in his paper on “ The Vegetation’ of the
Kermadec Islands” (Trans. N.Z. Inst. xli (1909), 172), records it, im addition, on
sand-dunes, gravel-flats, land-slips, &c. So far it has not found its way into
cultivation, but it would probably do well anywhere in the northern part of the
North Island, and from its free-flowering habit might prove a welcome addition to
our gardens.
Puate 118. Scevola gracilis, drawn from specimens collected on the Kermadec Islands. Fig. 1,
flower with the corolla removed (x 4); 2, corolla laid open, showing the glandular hairs on the inside
of the tube (x3); 3, two of the glandular hairs (x 15); 4 and 5, front and back view of anthers
(x 8); 6, longitudinal section of ovary (x5); 7, indusium, with one side cut away, showing the
2-lobed stigma (x 5).
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