JOURNAL OF

ISSN 003S-922X

THE ROYAL SOCIETY

OF

WESTERN AUSTRALIA

VOLUME 64
PART 4

JANUARY, 1982

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REGISTERED BY AUSTRALIA POST-PUBLICATION No. WBG 0351

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ROYAL SOCIETY

OF

WESTERN AUSTRALIA

PATRON

Her Majesty the Queen

VICE-PATRON

His Excellency Rear Admiral Sir Richard Trowbridge, K.C.V.O., K.StJ., Governor of

Western Australia

COUNCIL 1981-1982.

President

.... J. F. Loneragan, B.Sc., Ph.D., F.T.S.

Vice-Presidents

A. E. Cockbain, B.Sc., Ph.D.

S. J. J. F. Davies, M.A., Ph.D.

Past President

.... J. R. dc Lactcr, B.Ed. (Hons.), B.Sc. (Hons.), Ph.D ,
F.Inst.P., F.A.I.P.

Joint Hon. Secretaries

E. D. Kabay, B.Ag.Sci. (Hons.)

J. T. Tippett, B.Sc., Ph.D.

Hon. Treasurer

... S. J. Curry, M.A.

Hon. Librarian

H. E. Balme, M.A., Grad. Dip. Lib. Stud.

Hon. Editor

.. A. E. Cockbain, B.Sc., Ph.D.

D. T. Bell, B.A., Ph.D.

S. J. Hallam, M.A.

C. F. H. Jenkins, M.B.E., M.A., M.A.I.A.S.

L. E. Koch, M.Sc., Ph.D.

M. J. Mulcahy, B.Sc. (For.), Ph.D.

L. J. Peet, B.Sc., Dip. Val., Dip. R.E.M., F.G.S., A.R.E.I.

P. E. Playford, B.Sc. (Hons.), Ph.D., M.A.I.M.M.

P. R. Wycherley, O.B.E., B.Sc., Ph.D., F.L.S.

Journal of ihc Royal Society of Western Australia, Vol. 64. Part 4, 1982. pp. 101-132.

Hair structure of some Western Australian niamnials

by A. Valenle and P. A. Woolley

Zoology l^partment. La Trobe University, Buntloora, Vic. 3083
(communicated by B. K. Bowen)

Manuscript received 19 February 7980; accepted 17 February 19H1

Abstract

The technique of identification of hair in predator scats has been used in an attempt to
locate areas in which the Dibbler, Antechinus apicalis, lives. A photographic reference
system of the diagnostic features of the structure of the hair of 15 species of mammals
indigenous to the south of Western Australia has been compiled. This has been used in
conjunction with the photographs in Brunner and Coman (1974) of the hair structure of
other species found in (he 3 regions where the scats were collected. No Dibbler remains
were found.

InlrodiicHon

The method of identification of mammalian hair
developed by Brunner and Coman (1974) has been
found useful in mammal surveys. Uncommon or
inconspicuous species which are not often registered
by conventional techniques may be detected by
analysis of hair remains in predator scats (Brunner
and Bertuch 1976, Friend 1978). It was considered
that this technique might be useful in the search for
the Dibbler, Antechinus apicalis, which is now con-
sidered to be extremely rare. The Dibbler has been
found in recent times in only two localities, Cheyne
Beach and Jerdacultup. in the south of Western Aus-
tralia. Morcombe (1967) trapped the first two speci-
mens seen for 83 years at Cheyne Beach and his
discovery, together with the finding of two Dibblers
on farms near Jerdacultup, led to further searches
being made (Woolley 1977, 1980). Trapping has
been carried out in a number of localities in the
vicinity of Cheyne Beach and lerdacuttup, and also
in the Fitzgerald River National Park which lies
within the present known range of the Dibbler. How-
ever, the only area in which Dibblers have been
trapped is the one in which they were found by
Morcombe and only 9 individuals have been captured
(Woolley 1980). Because attempts to locale other
populations of the Dibbler by conventional trapping
methods have been unsuccessful predator scats have
been collected from the three regions in the south of
Western Australia in which trapping for the Dibbler
has been carried out.

In order to identify the hair found in the predator
scats it was necessary to prepare a reference set of
photographs of the most diagnostic features of the

Figure t. — Map showing the three regions (Cheyne Beach.
Fitzgerald River National Park = F.R.N.P. and Jerda-
cuttup) in which scats were collected. Drawn from map
R2l)l, sheet til, Australia S.W. sheet 2nd cd. Division
of National Mapping. Canberra, A.C.T.

hair of mammals which might be found in the regions
in which the scats were collected. This paper reports
on the structure of the hair of some mammals from
the south of Western Australia; the mammalian prey
items, identified by reference to the structure of hair
and skeletal remains, found in the scats collected will
be reported elsewhere.

Reference photographs of hair .structure

The 3 localities (Cheyne Beach, Jerdacultup and
Fitzgerald River National Park) in which trapping
for the Dibbler has been carried out are shown in
Figure 1. A list of the indigenous and introduced
mammals which might be found in the degree squares
encompassing the trapping areas was compiled from
the following sources: Ride (1970); records of the
Western Australian Museum (compuler printout of
mammalian species recorded by one degree squares
dated 12 June 1978); information provided by Dr.
A. N. Start, National Parks Board of Western Aus-
tralia. The 38 mammals listed comprised the follow-
ing 28 indigenous and 10 introduced species: —
Antechinus apicalis*, A. flavipes leucopaster*, Ante-
cliinomys lanij^er*, Sminthopsis crassicauAata, S.
ftranulipes*, S. hirtipes^, Phasconale caiura*, Oasyurus
peofjroii*, Myrmecobius iasciatus*, hoodon ohesulus,
Tarsipes spencerae*, Cercatetus concinnus, Tricho-
surus vulpeciila, Bettouftia penicUlata* , Macropus
euftenii*, M. fuliftinosus, M. irma*, Potorous
platyops*, P. tridactylus, Setonix brachyurus*,
Tachyglossus aculeatus, Hydromys chrysogaster,
N otomys mitcbelUi, Pseudontys albodnereus, P.
occidentalis* , P. shortridgei, Rattus fuscipes, R. rattus,
Mus nwsculns, Oryctolagus cuniculus, P'elis cat us,
Canis familiaris, yulpes vulpes, Sus scrofa, Ovis arics,
Bos taunts and Equus caballiis.

The structure of the hair of 13 of the indigenous
species and of the 10 introduced species is illustrated
in Brunner and Coman (1974). Samples of hair of
the 15 indigenous species not illustrated (asterisked
in the above list) were obtained from cither museum
specimens (Western Australian Museum, WAM;
Macleay Museum New South Wales, MM) or live
animals (Murdoch University Colony MU) and a set
of photographs of the structure of the hairs of each
prepared (Figs 2-16). Hair profiles were drawn to
scale. Whole mounts, cross sections and cuticular
scale casts were prepared as described in Brunner
and Coman (1974) and photographed using a Zeiss
photomicroscope. Prints were all made to one
standard magnification (x308).

The hairs found in the scats were identified using
a photographic reference system as described in
Brunner and Coman (1974). To make identification
easier the 38 species listed above were grouped accord-
ing to various characteristics of the primary guard

101

(0—14470

Journal of the Royal Society of Western Australia. Vol. 64, Part 4, 1982.

Hair profiles:- O = over hair, G = guard hair.

O

G

mm

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 30 /jm.

C-G Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair in mid- shaft region; E, primary guard liair near base;
F, smaller guard hair in shield region; G, smaller guard
hair in mid-shaft region.

H-J Scale patterns of guard hairs.

H, shield; I, transition between shield and shaft region;

J, lower- shaft.

B

A

Figure l. —Antechinomys laniaer WAM Ml 546.
102

Journal of the Royal Society of Western Australia, Vol. 64, Part 4, 1982.

H

I

103

J

^99000§BD§0§B&^§0§BOOeB^i

Journal of the Royal Society of Western Australia, Vol. 64, Part 4, 1982.

Hair profiles;- O = over hair, G = guard hair, U ^ under hair.

0 mm ^

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 40 ^m.

C-G Whole mounts of hairs.

C , primary guard hair in mid- shield region ; D, primary
guard hair in proximal shield region; E, primary guard
hair in mid- shaft region; F, primary guard hair near base;

G, under hair in proximal 1/3.

II-K Scale patterns of guard hairs.

H, shield; I, transition between shield and shaft regions;

J, shaft; K, near base.

B

A

Figure 3 . — Sminthopsis sranulipes WAM M2333.

104

Journal of the Royal Society of Western Australia, Vol. 64, Part 4, 1982.

105

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

Hair profiles;- G = guard hair, U = under hair.

0 mm 5

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 40 jum.

C-G Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair showing transition between shield and shaft; E, primary
guard hair in shaft region; F, smaller guard hair in shield
region; G, under hair in proximal 1/2.

H-J Scale patterns of guard hairs.

H, shield; I, transition between shield and shaft regions;

J, shaft.

Figure 4. —Sminfhopsis hirtipes WAM M1577.
106

usutasttianMeBigmsatii

Journal of the Royal Society of Western Australia, Vol. 64. Part 4, 1982.

Journal of the Royal Society of Western Australia. Vol. 64, Part 4, 1982.

Hair profiles:- O = over hair, G = guard hair, U = under hair.

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 80
C-G Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair in mid- shaft region; E, F, smaller guard hairs in
shield region; G, under hair in proximal 1/2.

H-K Scale patterns of guard hairs.

H, shield; I, transition between shield and shaft regions;
J, shaft; K, near base.

A

Figure 5 . — Tarsipes spencerae WAM MI5460.
108

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

( 2)— 14470

109

Journal of ihe Royal Society of Western Australia. Vol. 64. Part 4, 1982.

Hair profiles:- O = over hair, G = guard hair, U = under hair.

0 mm ^

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 45 jum.

C-G Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair in mid- shaft region; E, F, smaller guard hairs in
shield region; G, under hair in proximal 1/2.

H-L Scale patterns of guard hairs.

H, I, shield; J, transition between shield and shaft regions;
K, mid-shaft; L, near base.

A

Figure 6 . — Antechinus flavipes leitcogaster WAM M5559.

no

Journal of the Royal Society of Western Australia, Vol. 64, Part 4, 1982.

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

Hair profiles:- O = over hair, G = guard hair, U = under hair.

0 mm 10

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 65 fum.

C-H Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair in mid- shaft region; E, F, smaller guard hairs in
shield region; G, smaller guard hair in mid- shaft region;

H, under hair in proximal 1/2.

I-L Scale patterns of guard hairs.

I, mid-shield; J, lower shield; K, transition between shield
and shaft regions; L, shaft.

Figure 7 . — Aiuechintis apicalis WAM M1547I-2.

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

113

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

Hair profiles:- G = guard hair, U - under hair.

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 50 fum.

C-G Whole mounts of hairs.

G, primary guard hair in shield region; D, primai'y guard
hair in mid- shaft region; E, smaller guard hair in shield
region; F, smaller guard hair in mid- shaft region;

G, under hair in proximal 1/3.

H-K Scale patterns of guard hairs.

H, shield; 1, transition between shield and shaft regions;

J, mid-shaft; K, near base.

Journal of the Royal Society of Western Australia, Vol. 64. Pari 4, 1982.

Journal of the Royal Society of Western Australia, Vol. 64. Part 4, 1982.

Hair profiles: - O = over hair, G = guard hair, U = under hair

O

G

U

0

L

mm

10

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 80 Mm,

C-H Whole mounts of hairs.

C, primary guard hair in widest region; D, primary guard
hair in mid-shaft region; E, smaller guard hair in widest
region; F, smaller guard hair in mid- shaft region; G, smaller
guard hair near base; H, under hair in proximal 1/3.

I-L Scale patterns of guard hairs.

I, J, distal 1/3; K, transition between distal and proximal
regions; L, proximal 1/3.

Figure 9. — Da.synnts neoQroii WAM Ml 106.
116

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

117

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

Hair profile:- G = guard hair, U = under hair,

G
G

U

0

L

mm

10

-J

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 16 5 /uni.

C-F Whole mounts of hairs.

C, primary guard hair in widest region; D, primary guard
hair in proximal 1/3; E, primary guard hair near base;

F, under hair in proximal 1/2.

G-I Scale patterns of guard hairs.

G, distal 1/3; II, proximal 1/3; I, near base.

Figure 10 . — Myrmecohius fasciatus WAM M918.
118

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

Hair profiles:- O = over hair, G = guard hair, U = under hair.

O

G

U

0

L

mm

5

A. B Cross sections of hairs.

Maximum diameter of primary guard hairs 40 ^m.

C-G Whole mounts of hairs.

C, primary guard hair in shield region; D, primary guard
hair in mid- shaft region; E, smaller guard hair in shield
region; F, smaller guard hair in mid- shaft region,

G, under hair in proximal 1/2.

H-K Scale patterns of guard hairs,

H, shield; I, transition between shield 'and shaft regions;

J, mid- shaft; K, near base.

B

A

Figure 11. — P.seudomys occidentalis WAM MUH)93.
120

Journal of the Royal Society of Western Australia. Vol. 64. Part 4, 1982.

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

Hair profiles:- G = guard hair, U = under hair.

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 120
C-G Whole mounts of hairs.

C, primary guard hair in widest region; D, primary guard
hair in mid- shaft region; E, smaller guard hair in widest
region; F, smaller guard hair in mid- shaft region;

G, under hair in proximal 1/2.

II- K Scale patterns of guard hairs.

H, distal 1/3; I, transition between distal and proximal
regions; J, mid-shaft; K, near base.

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

123

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

Hair profiles:- G = guard hair, U = under hair.

G

G

G

U

Q mm

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 95

C-G Whole mounts of hairs.

C, primary guard hair in widest region; D. primary guard
hair in mid- shaft region; E, smaller guard hair in
widest region; F, smaller guard hair in mid- shaft region;

G, under hair in proximal 1/2.

H-J Scale patterns of guard hairs.

H, distal 1/3; I, mid- shaft; J, proximal 1/3.

Figure 13 . — Macropus eugenii MU.

124

Journal of the Royal Society of Western Australia. Vol. 64, Part 4, 1982.

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982

Hair profiles:- G = guard hair, U = under hair.

U

0 mm ^9

1 I

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 105 {jm.

C-G Whole mounts of hairs,

C, primary guard hair in widest region; D, primary guard
hair in mid- shaft region; E, smaller guard hair in widest
region; F, smaller guard hair in mid- shaft region;

G, under hair in proximal 1/2.

H-J Scale patterns of guard hairs.

H, distal 1/3; I, mid-shaft; J, near base.

Fitisure 14 . — Macropus irma MU.

126

Journal of the Royal Society of Western Australia, Vol. 64, Part 4. 1982.

Journal of the Royal Society of Western Australia. Vol. 64. Pari 4. 1982.

Hair profiles:- G = guard hair, U = under hair.

0 mm

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 120 fjm.

C-F Whole mounts of hairs.

C, primary guard hair in widest region; D, primary guard
hair in mid- shaft region; E, primary guard hair near base;

F, under hair in distal 1/3.

G-I Scale patterns of guard hairs.

G, distal 1/3; H, mid-shaft; I, proximal 1/3,

Journal of the Royal Society of Western Australia, Vol. 64, Part 4, 1982.

129

Journal of the Royal Society of Western Australia. Vol. 64. Part 4. 1982.

Hair profiles: - G = guard hair, U = under hair

U

0 mm

A, B Cross sections of hairs.

Maximum diameter of primary guard hairs 145 Mm,

C-F Whole mounts of hairs.

C, primary guard hair in widest region; D, primary guard
hair in mid-shaft region; E, smaller guard hair in mid-shaft
region; F, under hair in proximal 1/2,

G-J Scale patterns of guard hairs.

G, H, distal 1/3; I, mid- shaft; J, near base.

A

Figure 16 . — Setonix hrachyurus MU.

130

Journal of the Royal Society of Western Australia, Vol. 64, Part 4. 1982.

131

Journal of the Royal Society of Western Australia, Vol. 64. Part 4. 1982.

hairs (Table 1). In addition to the photographs hair
samples from most of the species were available so
that direct comparisons could be made if necessary.

Discus.sion

The species to which a sample of unknown hair
belongs can be identified by comparison with the
photographs of the structure of hairs in a reference
collection. Some species are very easily identified
because the hairs have very obvious distinguishing
characters while others lack such obvious characters
and may show only small differences from related
forms.

Among the species illustrated examples with
obvious distinguishing characters include Antechintts
apicaliSt Myr/necohius fasciatus, Tursipcs speiicerae.

Table 1

A groupinn of terrestrial mammals from the south of Western
Australia based on the structure of the primary guard hairs

Group 1

Group 2

Sub-group (a)

Sub-group (b)

Sub-group (c)

Sub-group (d)

Group 3

Hairs predominantly circular in cross section.
Dasyuridac •Antechinomys laniger

•Sminthopsis granulipes
•Sminthopxis hirtipes
Tarsipedidae *Tarsipes spencerae

Burramyidac Cercatetus concirtntts

Hairs predominantly oval in cross section.
Hairs with medulla much reduced or absent.

Bovidac Ovis aries

Sutdac Sus scrofa

Maximum diameter of guard hairs 45um,
with a distinct constriction before shield,
Dasyuridac *Antechinus fiavipes

leucogaster
Sminthosfsis crassi-
caudata

Sminthopsis niurina

Maximum diameter of guard hairs greater
than 45um. with a distinct constriction
before shield.

Dasyuridac *Antechinus apicalis

•Phascogale calura
•Oasyurus geoffroii
•Myrmecobius fasciatus
Maximum diameter of guard hairs greater
than 45um, no constriction before shield.

Bovidae
Canidae

Equidae

Felidae

Hairs lenticular
section.
Phalangeridae

Bus taurus
Canis famitiaris
yutpes lulpes
Equus cabatlus
Fells catus

(double convex) in cross

Group 4

Sub-group (a)

Sub-group (b)

Sub-group (c)

Sub-group (d)

Trichosurus vulpecula

Hairs predominantly oblong in cross section.
Medulla absent.

Tachyglossidac Tachyglossus aculeatus
Maximum diameter of guard hairs 45um,
with a distinct constriction before shield.
Muridae Notomys mitchelUi

Pseudomys albocinereus
*Pseudomys oeddentaUs
Maximum diameter of guard hairs greater
than 45um. with a distinct constriction
before shield.

Muridae Hydromys chrysogaster

Maximum diameter of guard hairs greater
than 45um. with no constriction before
shield.

Macropodidac *Beitongia penicillata
•Macropus eugenii
Macropus fuliginosus
• M acropus irma
•Potorous plafyops
Potorous tridactylus
•Setonix brachyurus

Hairs predominantly reniform (concave-con-
vex) in cross section.

Hairs with divided medulla.

Peramelidae isoodon obesulus

Hairs with bilobed or large medulla.
Muridae Mm musciilm

Pseudomys shortridgei
Rattns fuscipes
Rattus rattus

Hairs predominantly dumb-bell shaped in
cross section.

Leporidae Oryctolagits cuniculus

Species illustrated in Figures 2-16, remainder illustrated in
Brunner and Coman (1974).

Group 5

Sub-group (a)
Sub-group (b)

Group 6

Setonix brachyurus and Antechinomys laniger. A.
apicalis displays a globular arrangement of the
medulla which is very distinctive in cross section.
The hairs of both M. fasciatus and T. spencerae can
be easily recognised by the appearance of the medulla
in whole mounts and cross sections. S, brachyurus
has thick, long hair which displays a very distinctive
scale pattern along the proximal half of the hair.
Many of the guard hairs of A. laniger show an un-
common profile, with constrictions at several points
along the length of the hair.

Some of the marsupials illustrated are difficult to
distinguish from closely related forms illustrated in
Brunner and Coman (1974). These include: —
Antcchinus fiavipes leucogaster. the western form
of A. fiavipes; Sminthopsis hirtipes. which appears
to differ from S. crassicaudata only in the width of
the primary guard hairs and Potorous platyops and
P. tridactylus, in which there are only subtle
differences in the appearance of the medulla, best
appreciated by examining hairs rather than photo-
graphs. Among the murids, hair from Pseudomys
occidentolis differs little from other species of
Pseudomys illustrated in Brunner and Coman (1974),
P. shortridgei being the exception.

The grouping of species in Table 1 shows some
discrepancies with the grouping in Brunner and
Coman (1974). We have placed Sminthopsis crassi-
caudata in Group 2 (hairs predominantly oval in
cross section) and not in Group 1 (hairs predomin-
antly circular in cross section) on the basis of the
illustration in Brunner and Coman which shows
mainly oval hairs, and on the examination of refer-
ence hairs. Macropus fuliginosus and Potorous
tridactylus have been placed in Group 4 (hairs pre-
dominantly oblong in cross section) whereas Brunner
and Coman place them in Group 2 (hairs predo-
minantly oval in cross section). The difficulty in this
case appears to lie in the rather subjective interpre-
tation of the difference between oval and oblong
sections.

The primary aim in preparing this reference collec-
tion of photographs was to provide a method for
identifying the hair of the Dibbler, Antechinus
apicalis. The distinctive character of the hair of this
species makes it unlikely that any samples of it would
be misidentified. and none was found in any of the
predator scats examined.

Acknowledgements. — Wc wish lo lhank Mr. H. Brunner
(Keith Turnbull Research Insiilute) for instruction in the tech-
niques involved in the identification of hair and Dr. D. J.
Kitchener (Western Australian Museum). Dr. P. Stanbury
(Macleay Museum) and Dr. M. B. Rcnfrec (Murdoch Uni-
versity) for hair samples.

References

Brunner. H. and Bertuch. 1. (1976). — The broad-toothed rat
still in Sherbrooke Forest. A successful search
for Mastacomys fuscus Thomas. Viet. Nat., 93:
55-56.

Brunner, H. and Coman. B. (1974). — "The Identification of
Mammalian Hair". Inkata Press, Melbourne.
Friend. G. R. (1978). — A comparison of predator scat analysis
wi(h conventional tcchniquc.s in a mammal survey
of contrasting habitats in Gippsland. Victoria.
Aust. Wild!. Res.. 5: 75-83.

Morcombe, M. K. (1967). — The rediscovery after 83 years of
the Dibbler Antechinus apicalis (Marsupialia,
Dasyuridae). W. Aust. Nat., 10: 103-111.

Ride. W. D. L. (1970). — "A Guide to the Native Mammals of
Australia" . Oxford University Press, Melbourne.
Woolley, P. (1977).— In search of the Dibbler, Antechinus
apicalis (Marsupialia: Dasyuridac). J. Roy. Soc.
West Aust.. 59: 111-117.

Woolley, P. (1980). — Further searches for the Dibbler, Ante-
chintts apicalis (Marsupialia: Dasyuridae). J. Roy.
Soc. West Aust., 63; 47-52.

132

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Journal

of the

Royal Society of Western Australia

Volume 64

1982

Part 4

Contents

Hair structure of some Western Australian mammals. By A. Valente and
P. A. Woolley (communicated by B. K. Bowen)

Page

101

Editor: A. E. Cockbain
Journal Manager: J. Backhouse

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12 months after publication of part 4 of each volume.

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