EOL EES aN Aa canbiicasinc aS 8 el ei eesania sn Reese 2: a ies rh es Sh genet iS Ta eek ria sy ehhh SAY Cree tert aes eee pent “es Go Snax: as A earn STS Sad Oat: NATURAL HISTORY. SURVEY. STATE OF ILLINOIS Henry Horner, Governor DEPARTMENT OF REGISTRATION AND EDUCATION John J. Hallihan, Director LAKE MANAGEMENT REPORTS 3. Lincoln Lakes near Lincoln, Illinois David H. Thompson and George W. Bennett Printed by Authority of the State of Illinois NATURAL HISTORY SURVEY Theodore H. Frison, Chief Biological Notes No. 11 Urbana, Illinois August, 1939 FIG. 1.--Bathing beach at Lincoln Lakes built by the Lincoln Sand and Gravel Company. (Photo by Herbert Georg Studio.) FIG. 2.--Above, a winter view of Lincoln Lakes, showing wooded shores, FIG. 3.--Right, member of Logan County Sportsmen’s Club with largemouth bass from Sportsmen’s Lake, 1938, LAKE MANAGEMENT REPORTS 3. LINCOLN LAKES NEAR LINCOLN, ILLINOIS David H. Thompson and George W. Bennett Illinois Natural History Survey Lincoln Lakes are a series of water-filled gravel pits in the floodplain of Salt Creek, about a mile south of Lincoln, Illinois, They have been dug since 1905 by the Lincoln Sand and Gravel Company. The water area of these lakes is now about 100 acres. They are irregular in shape, and most of the wide-waters are connected by deep channels. The gravel company has developed a bathing beach, cottage sites and other recreational facilities for the general “se of the community. At the present time, Lincoln Lakes constitute about one-half of the total water area of Logan County, in central Illinois, and furnish the most convenient outdoor recreational spot for the residents of Lincoln, a city of 13,000. The water of Lincoln Lakes is clear and attractive for bathing, boating and fish= ing. The lakes are partially inundated at intervals of one to several years by flood water from Salt Creek. At such times the turbidity is increased, although the turbid condition remains for only a short period. As will be shown later, hook-and-line fish of sizes attractive to fishermen make up about 19 per cent of the total weight of fish in these lakes. Furthermore, an unusually large proportion of these fish are bass of large size. These fish do not take the hook readily, however, and fishing in Lincoln Lakes is considered poor. Early in 1938, the gravel company permitted the Logan County Sportsmen’s Club to use a small arm of water as an experimental area for the general improvement of the re~ mainder of the lakes. This arm was separated from the main lakes by a levee. Most of this report deals with work carried on in this small area, which will be called Sports-— men’s Lake, ACKNOWLEDGMENTS The Lincoln Sand and Gravel Company is interested in further developing the recreational uses of Lincoln Lakes and has given us unusually complete information. The Logan County Sportsmen’s Club, and especially those members primarily interested in fishing, have given a great deal of their time and effort in carrying out the work on Sportsmen’s Lake. Whatever benefits may arise from this work are to a great extent the result of their continued interest in the area, Many members have taken an active part in the work, but we should like to mention in particular Harry A. Gehlbach, Reginald Clark, Harold Klemm and Walter Spatz. A large part of the levee was built by the Na= tional Youth Administration under the direction of Herman Dammerman. At the time the census was made, we were helped by Dr. A. B. Taylor and Harry G. Kimple of the Univer- sity of Illinois. Dr. Donald F. Hansen, of the Natural History Survey, helped through out the census of Sportsmen’s Lake and furnished the information on the rate of growth of the white crappies of Lincoln Lakes from his manuscript on the growth of the white crappie in Illinois. The State Department of Conservation assisted with the work pre- vious to the census. Louis A. Krumholz, graduate assistant of the Natural History Survey, drew the maps and charts used in this report. DESCRIPTION OF LINCOLN LAKES Lincoln Lakes have a water area of 92.5 acres. (This does not include a new pond of about 10 acres dug since 1936.) The total shoreline is 7.3 miles. The maximum depth is reported to be 42 feet. The average depth is about 15 feet, and only 5 or 10 per cent of the water area is less than 5 feet in depth. A map of these water-filled gravel pits is shown in fig. 8. Digging has proceeded without interruption since 1905. The approximate dates of digging are shown on the map. The northwest water areas are Oldest, and those of the southeast newest. This paper is a contribution from the Section of Aquatic Biology. 3 The gravel which is being worked.was deposited in glacial times. Four to 10 feet of topsoil must be removed to expose it. Most of the topsoil is dumped into the lakes. The first digging was done with a dragline. In 1908, this was discarded in favor of a steam dredge, which was operated until 1922, when electric equipment was installed. Digging has averaged about 3 acres a year, and future operations will extend over a period of at least 15 years. New water areas will be south of the present lakes; the company holds land as far south as Deer Creek. The water level in the lakes is the same as that of the ground water. Surface drainage from about 500 acres of farm land to the north and east flows into the lakes. Almost every year Salt Creek overflows the low land near the uam and floods Lincoln Lakes. The Lincoln Sand and Gravel Company has made many improvements for public recrea=- tion on the worked-over areas. A large public bathing beach, fig. 1, was constructed in 1930 and 1931 at a cost of $16,000. This figure does not include 300 carloads of sand and gravel which were used. There is a large modern bathhouse and a parking area. A small charge for swimming pays the lifeguards and other operating expenses and pro= vides enough extra income for some new equipment each year. Figs. 2 and 3 are from photographs taken at Lincoln Lakes. The fees for fishing and boating are as follows: Fishing—$1.00 per year (plus the state fishing license) Boating--A few rowboats rent at 25 cents per hour, or $1.00 per day Privately-owned boats may be used on the lake at the following rates: Rownoatts 2s. ese e ewe we OO por year Garloes* 65S s evel we. wselite) eon ener Our: De ANTS o, Mra, |e tie mers ie ae leone ae ee 3.00 per year MOTOR DOAESH Ss ve\cs) ee ie eae ake See ne 5.00 per year Cottage sites may be leased for 10 years (renewable) at $35.00 per year. These sites have a 50-foot lake frontage and are 100 feet long. Eleven cottages are now in use. Hunting is not allowed on this area because of the danger and the nuisance to people seeking other recreation, A watchman= “I Standard Length, Inches Orange- Spotted Warmouth Bass * Zygonectes notatus (Refinesque) 10 The census shown in table 5 includes the fish taken in both poisonings. A large number of individuals of all species was measured. The frequencies of standard lengths of all species (except largemouth bass) are compiled in table 6. The carp, buffalo, largemouth bass and channel cat were large. Most of the crappies, bluegills and other pan fish were small, and few of them were of legal size. The numbers and weights of hook-and-line fishes of legal sizes are summarized in table 7. Although fishing in Lincoln Lakes is not considered good, the proportion of legal-sized hook-and-line fish is probably rather high, since they made up 19 per cent TABLE '7.--HOOK-AND-LINE FISH OF LEGAL SIZES. Legal Length, Inches Largemouth bass White crappie . BCP ee Weta te Green sunfish Yellow bass . Sheepshead. «. Channel cat... Black bullhead. # No legal lengths specified by state law. Arbitrary sizes chosen as the minimum for table use. of the weight of all fish present in Sportsmen’s Lake. It should be noted that the _largemouth bass and the bluegill were the kinds which contributed most of the legal- sized fish. AGE, GROWTH AND SEX IN THE LARGEMOUTH BASS When Sportsmen’s Lake was poisoned the first time, detailed observations were made on each individual bass. As the number was rather large and covered a wide range of sizes, we have treated these fish in some detail. One hundred sixty-seven bass were taken in the two poisonings; 146 the first time and 21 the second (not including the newly-hatched fry). Total lengths, standard lengths and weights were taken on the first group of fish. Sex determinations were Made, and scales collected for ages. Observations made on bass from the second poison= ing included only the number, the total weight, 45 pounds, and the sizes of a few indi- Vidual fish. For many years fish investigators have used standard length measurements (tip of Snout to end of backbone) in preference to total length measurements. Total length asurements in inches, on the other hand, are better understood by laymen and are, therefore, more useful in fish management. Test measurements of standard lengths made by three workers on a single fish were ound to vary considerably because of a difference of opinion about the exact location of the end of the backbone. On the other hand, total length measurements varied less. Total lengths are most uniform when measured from the tip of the snout, with the mouth tlosed, to the tip of the tail. The tail should be extended, i.e., not spread. The usual objection to the use of total length is that part of the tail fin may be broken ff. However, scarcely one fish in a hundred under natural conditions shows injury to mertail fin, RE Standard and total lengths were compared on all measured bass taken from Sports— men’s Lake to learn whether the ratio of total length to standard length is constant. The ratio was found to change as shown in table 8. It may be seen that the body grows in length more rapidly than the tail.* It was possible to determine the ages of all but two of the bass taken following the first poisoning. The numbers of bass of known ages are shown diagramatically by age groups in the shaded areas of fig. 5. The two bass of undetermined ages from the first poisoning and 21 bass from the second poisoning were arbitrarily assigned to age groups on the basis of size and are shown in unshaded areas in fig. 5. Spawning and survival must have been more successful in some years than in others. As may be expected, the greatest numbers of bass were one- and two-year-olds. But, there were more fish in the five- and six-year groups than in the three- and four-year groups. This fact may indicate an abundance cycle with dominant broods appearing every three or four years. Eschmeyer** presents evidence of dominant broods among the bass / TABLE 8.-=RATIO OF TOTAL LENGTHS TO STANDARD . LENGTHS IN BASS OF DIFFERENT SIZES. ' Standard Ratio of Length, Total Length to Inches Standard Length which he poisoned in Howe Lake, Michigan. In his case the first summer and fourth sum- mer fish were much more numerous than bass of other ages. Age determinations were made from the scales, distances between ‘annuli were meas— ured and the length of each fish at the end of each year of life was calculated in the usual manner. Table 9 shows the average of these calculated lengths for each year’s | spawn. Table 10 shows the average yearly length increases of the fish shown in table 9. Curves showing the sum of the length increases and the average lengths of these bass are given in fig. 6. The sum of the length increases (solid line) gives, in most respects, a more nearly true picture of the growth history of these bass. Average length curves are subject to greater irregularities. For example, the nine-year old fish average slightly shorter than the eight-year old fish. This does not mean that bass shrink between their eighth and ninth years but that the eight-year old average includes two unusually large fish which are not represented in the nine-year average. a No significant differences in growth rates of the two sexes could be found. : Scales and measurements from nine largemouth bass taken by anglers from Lincoln Lakes during 1937 and 1938 show a rate of growth similar to that of the poisoned bass of Sportsmen’s Lake. * Bennett found no change in the ratio of total to standard length in smallmouth bass when the former was measured from the tip of the snout to a line across the end of the tail spread in the natural position. Bennett, G. W. 1938. Growth of the Small- mouth Black Bass, Micropterus dolomieu Lacépede, in Wisconsin Waters. Copeia, No. 4, pp. 157-70. #* Eschmeyer, R. W. 1938, The Significance of Fish Population Studies in Lake Manage- ment. Trans. Third N. Am. Wildlife Conf., pp. 458-68. 12 OF ~ FISH NUMBER INCHES TOTAL LENGTH, | 2 3 a 2S 6 FA 8 9 AGE IN YEARS FIG. 5.--Shaded areas show numbers of largemouth bass of different ages taken from Sportsmen’s Lake following first poisoning. Unshaded areas represent numbers of bass arbitrarily assigned to age groups on basis of size: 2 from first poisoning, 21 from second. | 2 3 a 5 6 if 8 9 AGE IN YEARS FIG. 6.--Rate of growth of largemouth bass in Sportsmen’s Lake. Solid line, sum of length increases; broken line, average lengths. 13 | ever | | 9s*6T_ [ user | 96°FT | LTS°ST soseelout yysueT Jo mm Tae eseIOAY VL Ris FSORNMNAMN AH) ie] NOONoODr MW hi 1k T. g T 4 0 ra poseloay usta JO eJTT JO reer Iequmnn p ita} ci *@S6T ‘ANNC-AVW SOMIVI S,;NOWSLYOdS WOU NOIVL SSVA HLNOWHOUVT 0 SHHONI NI SASVHYONI HLONYI GALVINOTVO--°OT WIaVL eseIOAY FPL 82H mks TE 48 00 Le peseloAy Ustwd Jo BIeeX UT 83 Jeqmunn "@S6T ‘ANOP-AVW ‘SDIVI S.NWSLYOdS WOUd NODIVL SSVA HLNOWHOUT 4O SHHONI NI SHLDONYT TVLOL (&LVINOTVO--"6 WIdVL peumeds Jeez 14 All bass of four years and older were of legal length (i.e., 10 inches or more). Eight out of 10 three-year-olds were of legal length, as well as 5 of the 47 two-year- olds. The largest yearlings were 9 inches long. Ages, length frequencies and sexual development of these 144 bass are shown in table 11. It may be seen that sexual maturity is correlated more closely with length than with age. All fish over 10$ inches long were sexually mature; all below 10 inches were immature. Examination showed that males and females become mature at about the same size. Of the 56 sexually-mature specimens examined, 19 were females and 37 were males. TABLE 11.--NUMBERS OF LARGEMOUTH BASS (TAKEN FROM SPORTSMEN’ S LAKE, MAY-JUNE, 1938) OF DIFFERENT LENGTHS AND AGES, SEXUALLY MATURE AND IMMATURE. (SEXUAL MATURITY DETERMINED BY DISSECTION. ) Total Age in Years Length, Inches Sexually immature 2 3 2 wwe « DPrH. May 20 apparently bisected the time of annulus formation in the five younger age groups. Sixty of these bass had newly-formed annual rings, while 66 had not formed their 1938 rings. None of the 18 fish six or more years old had formed new rings. ’ 15 These data are summarized in table 12. May 20 is about a month earlier than the mean date determined by Hansen for annulus formation in the white crappie.#* TABLE 12.--NUMBERS OF LARGEMOUTH BASS WITH AND WITHOUT NEW ANNULI ON MAY 20, 1938, SPORTSMEN’S LAKE. Number Without New Annulus Formed Number With New Amnulus Formed il 2 3 4a 5 6 Ui 8 9 There is a tendency for the larger bass of an age group to form their annual ring before smaller bass of the same age. This may be seen in table 13. Some of the bass which escaped the first poisoning in Sportsmen’s Lake spawned within the next week or two; large numbers of inch-long fry were killed by the second poisoning, June 14. This seems to indicate a close correlation between the time of annual ring formation and the time of spawning. TABLE 13.--NUMBERS OF BASS OF DIFFERENT LENGTHS WITH AND WITHOUT NEWLY-FORMED ANNULI, MAY 20, 1938, SPORTSMEN’S LAKE. One-Year Old Bass Two-Year Old Bass Standard aati Ring Ring Ring Ring Formed Not Formed Formed Not Formed HPWAWAOA ONE - Fish are often described as plump or thin, but investigators have devised a more accurate method of measuring condition by dividing the weight of the fish by the cube * Hansen, Donald F., 1937. The Date of Annual Ring Formation in the Scales of the Whi Crappie. Trans. Am, Fish. Soc., 66:227-36. 16 of its length. It is most convenient to divide the weight in pounds by the cube of the total length in inches. In order to avoid long decimals, the result is multiplied by 10,000; in other words, the decimal point is moved four places to the right. For ex- ample, a 10-inch fish weighing one-half pound will have an index of condition of oe en? or 5.0. A 20-inch fish weighing 4 pounds will have an index of condition of pipes Tien 1OSO00 7 > or 5.0. Thus it may be seen that they are 20 hin se 20" i, x 20° in? equally plump. This figure can be referred to as the index of condition, or abbreviated as C. Other workers have devised similar methods for comparing the plumpness of fish of different sizes, but most of them have used lengths in centimeters and weights in grams; or have used standard lengths, or some linear measurement other than total length. One of the most common measures is calculated from standard lengths in millimeters and weights in grams. This is called the coefficient of condition, or K. This is trouble- some to the average fisherman because he usually measures fish in inches and weighs them in pounds. The index of condition, C, was calculated for each of the bass taken from Sports-— men’s Lake which were weighed and measured. Table 14 shows that C becomes progressively greater in longer fish. This indicates that large bass are stockier than small bass. TABLE 14.--AVERAGE INDEX OF CONDITION, C, OF LARGEMOUTH BASS OF DIFFERENT SIZES, SPORTSMEN’S LAKE, MAY 20, 1938, Index of Condition, (S Qn > - OMrHNN SHRaDA It may be interesting in later work to use this bass population for comparison with the bass populations of other waters. The relation of standard length to total length has been smoothed of its minor irregularities by fitting a curve. In like manner the relation between length and weight has been smoothed. Table 15 shows corresponding lengths, weights and values of C and K, read or computed from these smoothed curves. TABLE 15.--CORRESPONDING LENGTHS, WEIGHTS, INDICES OF CONDITION (C) AND COEFFICIENTS OF CONDITION (K) OF LARGEMOUTH BASS FROM SPORTSMEN’S LAKE, MAY 20, 1938. Standard Weient Total Standard Length, Ents Length,| Length, Inches Inches Inches OODIATP EW BRPAASGRS e YNHONNNNNWND Pea ene o Ss . tn iS DoH fo e CS SaAaesres OPrA Ono t & RYE OSS . noon OPP OMNMHHH ooo) DWP O BRANNHASS PPP PPR RRO DiyO PC Oo RACHA ERA AO On eoese so eS © Aonaannnon e YOVNWNWMNWNNWNWD DW © CRaNSSase a 5 6 ti 8 9 10 ial 12 C1 17 LIMNOLOGICAL OBSERVATIONS On June 2, 1938, temperatures were taken at two stations in the deeper water (Sta- tions A and B) and at the head of one of the larger fingers (Station C) of Sportsmen’s Lake. These temperatures are shown in table 16. TABLE 16.--TEMPERATURES IN DEGREES FAHRENHEIT IN SPORTSMEN’S LAKE, JUNE 2, 1938. Station A Station B Station C Temperature Temperature Temperature 0) 1 2 3 a 5 6 7 8 9 59.5, bottom 47.5, bottom Later in the summer, on August 12, temperatures and oxygen samples were taken at one station in the deep, open water of Lincoln Lakes, as well as in Sportsmen’s Lake. The first location was about 200 feet west of the boatyard. The station in Sports- men’s Lake was the same as Station B on June 2. The temperatures and dissolved oxygen concentrations found are shown in table 17. TABLE 17.--TEMPERATURES IN DEGREES FAHRENHEIT AND DISSOLVED OXYGEN CONCENTRATIONS IN PARTS PER MILLION IN SPORTSMEN’S LAKE AND OTHER PARTS OF LINCOLN LAKES, AUGUST 12, 1938. Lincoln Lakes Sportsmen’s Lake Dissolved Oxygen PO MH O 5 6 Li 8 9 @) it a=) Continued on next page) 18 TABLE 1'7.-—-TEMPERATURES IN DEGREES FAHRENHEIT AND DISSOLVED OXYGEN CONCENTRATIONS IN PARTS PER MILLION IN SPORTSMEN’S LAKE AND OTHER PARTS OF LINCOLN LAKES, AUGUST lé, 1938--Continued. Lincoln Lakes Sportsmen’s Lake Depth, Feet Temperature Dissolved Temperature Dissolved Oxygen Oxygen 0.0, bottom The two series of temperatures taken in Sportsmen’s Lake, June 2 and August 12, are diagramed in fig. 7. The common kinds of fish found in artificial lakes in Illinois--largemouth bass, white crappie, black crappie and bluegills--require two to three parts per million SURFACE 4 a ie so ir oo ape uJ ra) 16 BOTTOM ° TEMPERATURE, F. FIG. 7.--Water temperatures in Sportsmen’s Lake, June 2 and August 12, 1938. 19 of dissolved oxygen. In Lincoln Lakes, in summer, they cannot be expected to live below a depth of 15 feet. Bullheads, however, do not require such a high concentra- tion of oxygen as the other kinds of fish of Lincoln Lakes. Both the black bullhead and the yellow bullhead can tolerate an oxygen concentration as low as one-half part per million. TABLE 18.--CHEMICAL ANALYSES OF WATER FROM SPORTSMEN’S LAKE. April 30, 1938, Nov. 16, 1958, Determination Parts Per Million | Parts Per Million Iron PaLGEmeds a. «-chtare eh Leu eure 0.08 trace METRE! 6 ct emte olen On Oe gitteds (SiO)... eee see 4.0 1.5 Calcium (ca) a eee, laa 49.5 37 el Maenesivm (Me) . 26% eel 23.5 24.2 Potassium: (KS). < sci comrente 0.9 4.4 gultate (SOgye: « : . . ; gies se : Bore Dtecrw eer " GE BEE ES ees NARA ee ne See ee Ee od es ae