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NATURAL

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HISTORY SURVE1

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FIELDIANA • ZOOLOGY

Published by

CHICAGO NATURAL HISTORY MUSEUM
Volume 31 April 29, 1949 No. 28

LAND AND FRESH-WATER MOLLUSKS
FROM PERU

Fritz Haas
Curator of Lower Invertebrates

The material here reported was gathered by Mr. Jose" Maria
Schunke, a professional collector in Pucalpa. The collection proved
to be especially interesting on account of the rare species it contains
and the high number of new species. Mr. Schunke collected in
the departments of Huanuco and Loreto, in the following localities:

HuAnuco. — Divisoria, altitude about 5,000 feet; May 15, 1947,
and the months of August and September, 1947. Divisoria
is the familiar name of the Cordillera Azul, the watershed
between the Huallaga and Ucayali rivers, which is essentially
followed by the border between the departments of Huanuco
and Loreto; Mr. Schunke's locality is situated on the west
side of the crest, toward Huanuco.

Loreto. — Pucalpa (often spelled Pucallpa), on the Rio Ucayali;
February 5, 1947. Rio Neshuya, left affluent of the Ucayali,
crossed by the highway from the valley of this river to
Huanuco, in the Huanuco Valley; October 2, 1947. Conta-
mana, on the Rio Ucayali; May 20-22, 1947. Cerro Azul,
on the Rio Ucayali, near Contamana; March and April, 1947.
Metelo Yacu, near the Cerro Azul and near Contamana, an
affluent of the Ucayali; May, 1947.

For the photographs reproduced in this article I am indebted to
my friend and colleague Dr. Rainer Zangerl, Curator of Fossil Rep-
tiles, and for the drawings used for figure 54, to Miss Margaret
Bradbury, Artist in the Department of Zoology.

LIST OF SPECIES

Pleurodonte (Labyrinthus) baeri baeri Dautzenberg.

Six specimens from the Cerro Azul, May 10, 1947.

No. 626 235

THE LIBRARY OF THE

JUN 2 - 1949 Natural History Survey

iwivcrsity o.6 liUiAj- library

236 FIELDIANA: ZOOLOGY, VOLUME 31

Pleurodonte (Labyrinthus) dacostiana Preston.

Two specimens from Divisoria, the one collected between August 5
and September 10, 1947, the second on September 18, 1947. This
seems to be the first report of this species since its publication
(Preston, 1907, p. 490, fig. 3).

Solaropsis (Psadara) calstelnaudii Deville and Hupe\
One specimen from Divisoria, August 26, 1947.

Strophocheilus (Megalobulimus) maximus maximus Sowerby.
One specimen from Pucalpa, February 5, 1947.

Orthalicus (Orthalicus) gallina-sultana Chemnitz.
Two specimens from Metelo Yacu, May 10, 1947.

Bulimulus (Bulimulus) inconspicuus sp. nov. Figure 50, a.

Type. — Chicago Natural History Museum No. 30038, from
Contamana on Rio Ucayali, Department of Loreto. Collected by
Jos<§ Maria Schunke, May 20, 1947.

Diagnosis. — A medium-sized species of the typical subgenus of
Bulimulus, plain, inornate, transparent, dull horn-colored.

Comparisons. — Of all the many South American species of the
typical subgenus of Bulimulus, this new species is closest to B.
transparens Reeve, of unknown locality. It differs from that species,
however, in being smaller (17 mm.; 19 mm. in transparens) and in
having 7 whorls, whereas transparens is said to have only 6; however,
the two species resemble each other so closely that the description
of transparens, with the few features mentioned changed, might
serve for the new species also.

Description of type. — Shell subperforate, oblong-conic, thin,
rugosely striate, pellucid, dull corneous. Spire conic, slightly obtuse.
Whorls 7, a little convex, the last a little shorter than the spire,
somewhat attenuated at base. Columella arcuate. Aperture
oblique, oval; peristome simple, the right margin slightly arcuate,
columellar margin slightly reflexed above, subappressed.

Measurements of type. — Height 17 mm., width 7.5 mm., height
of aperture 6.9 mm., width of aperture 4.2 mm.

5?0.5

'.3/

HAAS: MOLLUSKS FROM PERU

237

Notes on paratypes. — Chicago Natural History Museum No.
30039a-c. The paratypes at hand, with the same data as the type,
consist of a specimen three-fourths adult, another subadult and
fragments of an adult specimen, which, judging from the dimensions
of the aperture, must have been slightly larger than the type. The
type specimen itself is a little damaged also, as shown in figure 50, a.

Drymaeus (Drymaeus) schunkei sp. nov. Figure 50, b.

Type. — Chicago Natural History Museum No. 30040, from Cerro
Azul, on Rio Ucayali, Department of Loreto. Collected by Jose"
Maria Schunke on May 8, 1947.

Diagnosis. — A larger species of the typical subgenus of Drymaeus
Characterized by its ovate, acuminate shape, the black-brown zigzag

Fig. 50. a, Bulimulus (Bulimulus) incon-
spicuus sp. nov. C.N.H.M. No. 30038; type,
about Xl.5. b, Drymaeus (Drymaeus) schunkei
sp. nov. C.N.H.M. No. 30040; type, about
Xl.5. c, Drymaeus (Drymaeus) gibber sp. nov.
C.N.H.M. No. 30042; type, about Xl.5.

stripes of its surface, and the large aperture with its broadly ex-
panded, yellow-colored peristome.

Comparisons. — Very similar to Drymaeus (Drymaeus) aequatori-
anus E. A. Smith (1877, p. 363, pi. 39, fig. 7) from Ecuador, differing
from it mostly by the conspicuously broader expansion of the peri-
stome, which, furthermore, is yellow, not rose-colored as in aequatori-
anus. The affinities of both species, however, are so close that they
may represent geographical races only, not separate species.

Description of type. — Shell ovate acuminate, subumbilicate, thin,
substriate, and with faint spiral lines. Spire conic, apex acute, with

238 FIELDIANA: ZOOLOGY, VOLUME 31

5% whorls that are slightly convex and separated by a shallow
suture; last whorl slightly ascending in front, inflated toward the
aperture. Color of shell white with blackish brown zigzag stripes
that are also visible inside the aperture. Aperture shortly oval, a
little higher than the spire, with a wide, yellow-colored peristome
that reflexes over the umbilical chink.

Measurements of type. — Height 27.1 mm., width 14.1 mm.,
height of aperture 15.1 mm., width of aperture 11.4 mm.

Notes on paratypes. — Chicago Natural History Museum No.
30041a-c. Same data as the type. The paratypes show no traces
of variation. They correspond with the type in all characters of
shape and coloration. Their dimensions are:

mm. mm. mm.

Height of shell 27.0 26.9 26.1

Width of shell 13.6 13.2 12.8

Height of aperture 15.0 15.0 13.7

Width of aperture 10.7 10.0 10.2

Drymaeus (Drymaeus) expansus Pfeiffer.

One specimen from the Cerro Azul, May 8, 1947, which, while
otherwise quite typical, is much smaller than all the known forms
of this species. Its measurements are: Height 22.9 mm., width
14.6 mm., height of aperture 14.0 mm., width of aperture 10.5 mm.

Drymaeus (Drymaeus) inca Maxwell Smith.

Three specimens from Divisoria, collected between August 5 and
September 10, 1947. First report since original description (Maxwell
Smith, 1943, p. 61, pi. 7, fig. 10).

Drymaeus (Drymaeus) gibber sp. nov. Figures 50, c; 51, a-d.

Type— Chicago Natural History Museum No. 30042, from
Divisoria, Department of Huanuco. Altitude 5,000 feet. Collected
by Jose" Maria Schunke between August 5 and September 10, 1947.

Diagnosis. — A large specimen of Drymaeus proper, characterized
by a hunchback-like swelling of the last whorl about half a whorl
from the aperture, a feature unique in the genus.

Comparisons. — This new species corresponds in almost every
feature with the Ecuadorian Drymaeus inaequalis Pfeiffer (1856,
p. 330; 1860, p. 158, pi. 42, figs. 3-5) but for the hump-like gibbosity
in the middle of the last whorl, which gives a marked asymmetrical

y

HAAS: MOLLUSKS FROM PERU

239

aspect to the shell; the color variation of D. gibber is even greater
than that of inaequalis.

Description of type. — Shell irregularly elongated, imperforate,
thin, shining. Whorls 7, rather flat, the first six regularly, the last
rapidly increasing, with a gibbosity that inflates its latter half irregu-
larly, causing it to deviate from the axis of the shell to the right;
while the back of the shell becomes very convex, its front is almost
flat. Suture well impressed, ascending a little toward the aperture.
The basic color of the surface is a pale, grayish yellow on which
slightly undulate blackish stripes appear; these stripes are wide,

Fig. 51. a-d, Drymaeus (Drymaeus) gibber sp. nov. C.N.H.M. No. 30043;
paratypes, showing variation of shape and pattern; about Xl.

as wide as the interstices, and split up in narrow stripes between
which equally wide stripes of the basic color are visible. Aperture
long, slanting forward, ovate-oblong, showing the external marking
inside, but with the blackish stripes entire, not split into narrower
ones; peristome pale yellow, outer and basal margins rather widely
expanded, columellar margin slightly expanded and reflexed and
adnate over the curved columella.

Measurements of type. — Height 38.1 mm., width 19.4 mm.,
height of aperture 19.6 mm., width of aperture 8.5 mm.

Notes on paratypes. — Chicago Natural History Museum No.
30043a-d, and No. 30044a-p. Same data as the type. The nineteen
paratypes at hand show that this species varies quite notably.
Figure 51, a-d, demonstrates the variation in shape as well as in

240 FIELDIANA: ZOOLOGY, VOLUME 31

color pattern. The shape varies from that of the type (fig. 51, a)
gradually (figs. 51, b and c) to an almost regularly convoluted
specimen (fig. 51, d) in which the gibbosity of the last whorl, though
still clearly recognizable, is much less developed. The color pattern
may be that of the type (fig. 51, c and d), or there may be narrow,
widely spaced, undulating, grayish-brown stripes (fig. 51, b), or,
lastly, the basic color may be banded by three irregularly confined
spirals, which can anastomose and on which there are minute spot-
like windows of the basic color (fig. 51, a).

Euglandina (Euglandina) sp.

One very young specimen from Contamana, May 20, 1947,
which, while permitting an unequivocal classification of the genus
and subgenus, is too minute to allow specific determination. No
Euglandina has apparently ever been reported from Peru.

Leptinaria (Leptinaria) lamellata Potiez and Michaud.
Twenty-eight specimens from Contamana, May 22, 1947.

Subulina (Subulina) yatesi Pfeiffer.

Twenty-seven specimens from Contamana, May 20, 1947.

Obeliscus (Protobeliscus) teres sp. nov. Figure 52.

Type. — Chicago Natural History Museum No. 30028, from
Divisoria, Department of Huanuco, altitude about 5,000 feet. Col-
lected by Jose- Maria Schunke, August 13-15, 1947.

Diagnosis. — A species of the subgenus Protobeliscus, characterized
by an almost cylindric shape.

Comparisons. — None of the Peruvian and Ecuadorian species of
Protobeliscus bear any closer similarity to this new species, but the
Bolivian Obeliscus bacterionides Orbigny, which Pilsbry (1906, p. 250,
pi. 36, figs. 71-72) includes in Obeliscus proper, looks very much
like it. Pilsbry's copy of Orbigny's Helix bacterionides (1837,
p. 260, pi. 29, figs. 1-3) is rather inexact and misleading, whereas
the original figure shows great similarity to the new teres. Judging
from Orbigny's original figure, bacterionides has to be a member of
the subgenus Protobeliscus, in which I am placing teres also. L.
Pfeiffer (1852, p. 117, pi. 35, figs. 15-17), too, gives a figure of
what he believed to be Orbigny's bacterionides, and Pilsbry (1906,

HAAS: MOLLUSKS FROM PERU 241

p. 250) quotes this figure in the synonymy of this species; however,
I feel certain that Pfeiffer's figure represents an entirely different
species, showing a much more oval shell than bacterionides.

Description of type. — Shell long, narrow, turrite, almost cylindric,
transparent, smooth, almost lubricate, glossy, straw-yellow. Apex
obtuse. Eight and one-half whorls, suture shallow and somewhat
appressed. Aperture high and narrow, pear-shaped, with cutting
edges. Columella not truncated, slightly twisted. No umbilical
perforation or chink.

Measurements of type. — Height 21.2 mm., width 5.8 mm., height
of aperture 6.4 mm., width of aperture 4 mm.

Fig. 52. Obeliscus Fig. 53. Nenia (Co-

(Protobeliscus) teres sp. nov. lumbinia) obesa sp. nov.

C.N.H.M. No. 30028; type, C.N.H.M. No. 30025; type,

about X2. about X2.

Notes on paratype. — Chicago Natural History Museum No.
30029. Same data as the type. Height 17.9 mm., width 5.4 mm.,
height of aperture 6.1 mm., width of aperture 2.9 mm.

Nenia (Columbinia) obesa sp. nov. Figures 53, 54.

Type. — Chicago Natural History Museum No. 30025, from Cerro
Azul on Rio Ucayali, Department of Loreto. Collected by Jose"
Maria Schunke, May 9, 1947.

Diagnosis. — A species of the subgenus Columbinia, characterized
by considerable obesity.

242 FIELDIANA: ZOOLOGY, VOLUME 31

Comparisons. — While completely in accordance with Polinski's
diagnosis of his subgenus Columbinia (1925, p. 743) as far as the
internal features of lamellae and folds are concerned, this new species
differs from all the other known consubgenerics, i.e., perezi Jous-
seaume, columbiana Polinski, epistomium Kuster, and pseudepistom-
ium Bourguignat, all of which are rather slender, by its obesity. It
shares this character with Nenia reyrei Jousseaume from Ecuador
(Jousseaume, 1887, p. 172, pi. 3, fig. 9), which is still stouter, but
which apparently is very close to obesa as far as the external and
internal features of the shell are concerned. Polinski (1925, p. 744)

Fig. 54. Nenia (Columbinia) obesa sp. nov. a, Side view of the last whorl,
showing disposition of principal and lunellar folds; about Xl.5. b, Clausilium,
about X5.5; C.N.H.M. No. 30026, paratype.

had not included reyrei in his new subgenus Columbinia, most
probably because its general shape was so different from that of
the species comprised in Columbinia; a mistake or a misprint
occurring in the original description of Nenia reyrei may have
contributed to this exclusion, namely, the fact that the length of
this species is given as 13 mm., whereas it should read 23 mm.,
according to the corresponding figure on plate 3. Another species
comparable with this novelty is Nenia bryantwalkeri Pilsbry from
Huallaga, Peru (Pilsbry, 1922, p. 95, pi. 2, figs. 4, 5, 12-14). Indeed,
the similarity between the two species is great. They differ by the
length of the shell, 15 mm. in bryantwalkeri and 25.8 mm. in obesa;
by the way in which the superior lamella is joined to the spiral
lamella, as there is no bend in bryantwalkeri and a marked one in
obesa; and by the different shape of the clausilium, which does not
show any excision in obesa.

Description of type. — Shell entire, not decollate, fusiform, widest
at the sixth whorl. Color Cinnamon to Verona Brown. Seven

HAAS: MOLLUSKS FROM PERU 243

and one-half whorls, a little convex, the last becoming free and
descending more rapidly. Apex blunt. Suture not very distinct.
Sculpture of oblique, low, but distinct, straight or slightly arcuate
riblets, of which there are 5 or 6 in 1 mm. on the penultimate whorl,
while on the neck they become much more crowded. Neck rounded.
Aperture slightly protracted, irregularly ovate, with strongly ex-
panded, continuous, flesh-colored peristome. Superior lamella some-
what protruding above the peristome, strong, vertical, channelled
on the left side; inferior lamella not reaching to the margin of the
peristome, rather low, almost horizontal in a frontal view. Principal
fold dorsal, about half a whorl long, lunella subdorsal, strongly
arched, free from the principal fold (fig. 54, a).

Measurements of type. — Height 25.2 mm., width 6.7 mm., height
of aperture 6.1 mm., width of aperture 5.3 mm.

Notes on paratypes. — Chicago Natural History Museum No.
30026 and No. 30027a-m. Same data as the type. Paratype 30026,
which is partly broken, enables me to describe the interior features
of this species. The superior lamella joins the spiral one in a decided
bend to the left, and the inferior one, which is low in the aperture,
is rapidly becoming high in the interior and extends about half a
whorl deep. The clausilium, whose shape can be better explained
by a figure (54, b) than by a verbal description, is thicker on the
distal end and does not show any excision. The remaining paratypes,
No. 30027, vary from 22.2 mm. to 25.1 mm. in height, and from
6.1 mm. to 6.8 mm. in width; one specimen, however, has the out-
standing width of 6.9 mm. with a height of only 24.9 mm.

Nenia (Incania) jelskii Polinski.

Four specimens from the Cerro Azul, May 16, 1947.

Nenia (Incania) trigonostoma O. Boettger.

Five specimens from Divisoria, between August 13 and 15, 1947.

Happia (Drepanostomella) ammoniformis Orbigny.
One young specimen from Contamana, May 22, 1947.

Systrophia (Systrophia) affinis Pilsbry. Figure 55.

This species will be discussed, under comparisons, with the follow-
ing one.

Twenty specimens from Contamana, May 20-22, 1947.

244 FIELDIANA: ZOOLOGY, VOLUME 31

Systrophia (Systrophia) obvoluta sp. nov. Figure 56.

Type. — Chicago Natural History Museum No. 30030, from
Contamana, Department of Loreto. Collected by Jose1 Maria
Schunke, May 20-22, 1947.

Diagnosis. — A small species of the typical subgenus of Systrophia,
characterized by its most densely convoluted whorls, by a rather
narrow umbilicus, and by a relatively great height of shell.

Comparisons. — Since all the species of Systrophia proper exhibit
basically the identical shell features and since their distinction rests

Fig. 55. Systrophia (Systrophia) affinis Pilsbry. C.N.H.M. No. 30032; shown
from side, above and below, about Xl.25.

almost entirely upon the combination of relative differences in width,
height, number of whorls, etc., it is difficult to compare this new
species to any individual congeneric species. However, since a

Fig. 56. Systrophia (Systrophia) obvoluta sp. nov. C.N.H.M. No. 30030;
type, shown from side, above and below, about Xl.75.

second kind of the same genus was found together with obvoluta,
and since this other species has never been figured before and has
been mentioned but once in the literature, I compare our novelty
to it; the species I have in mind is Systrophia (Systrophia) affinis,
described by Pilsbry (1900, p. 388) as Polygyratia affinis. This
species, which is figured here (fig. 55) for the first time, measures
up to 12.7 mm. in diameter, has a flat, even slightly sunken spire,
a rather wide umbilicus, and S}4 whorls, whereas obvoluta has a
diameter measuring up to 9.2 mm., a flat but somewhat raised spire,
a narrow umbilicus and 9 whorls. The height of both species being
equally about 4 mm., the relative height of the shell of obvoluta is
considerably greater.

I

HAAS: MOLLUSKS FROM PERU 245

Description of type. — Shell planorboid, flat or somewhat raised in
the middle, with a rather narrow, funnel-shaped umbilicus below,
pale yellow, glossy, faintly and distantly striatulate and showing
more or less distinctly some spiral lines. Whorls 9, excessively
densely coiled, the last about three times as wide, but four times as
wide near the aperture, as the preceding, peripherally rounded,
suddenly widening and slightly descending in front of the aperture,
somewhat flattened there on its upper part. Aperture oblique,
higher than wide, semilunar, peristome thin and slightly reflected
all around, but mostly at the base.

Measurements of type. — Diameter 9.2 mm., height 4.0 mm.,
diameter of umbilicus 2.9 mm.

Notes on paratypes. — Chicago Natural History Museum No.
30031a-k. Same data as the type. Variation of size, in the material
at hand, ranges from 9.2 and 8.7 mm. in width and from 3.5 and
4.0 mm. in height. In two specimens, each measuring 9.2 mm. in
diameter, the height is 3.7 and 4.0 mm., respectively.

Systrophia (Systrophia) retinella sp. nov. Figure 57.

Type. — Chicago Natural History Museum No. 30035, from
Contamana, Department of Loreto. Collected by Jose" Maria
Schunke, May 22, 1947.

Diagnosis. — A species of Systrophia proper with 7J/£ regularly
increasing, not tightly coiled up whorls and a rather narrow, deep
umbilicus.

Comparisons. — As pointed out above, all systrophias have so
much in common in the shape of their shells that it is hard to find
one relative closer than another to a given species. This case, how-
ever, has its alterum comparationis in S. inca Crawford (1939, p. 324,
pi. 20, figs. 20-23) from Pisacc in the Urubamba Valley, Peru. This
species is smaller, has a more elevated spire, a greater number of
whorls, and a much wider umbilicus. Both species belong to that
group of the typical subgenus of Systrophia that is not planorboid,
that has whorls not closely convoluted, and that, on behalf of these
characters, looks very much like the Retinella of the family Zonitidae
of the Northern Hemisphere.

Description of type. — Shell with very low spire, with the apex
raised above the upper surface, thin, transparent, pale grayish yellow.
Whorls 5 }/2, rounded, the first 43^ regularly, the last one faster
increasing, being almost three times wider at the aperture than the

246 FIELDIANA: ZOOLOGY, VOLUME 31

preceding; the first two whorls smooth, the following ones with
irregular transverse striae and faint spiral lines. Suture distinct.
Umbilicus deep and narrow. Aperture as wide as high, circular,
with simple, unexpanded lips.

Measurements of type. — Diameter 7.1 mm., height 3.3 mm.,
diameter of umbilicus 2.0 mm.

Notes on paratypes. — Chicago Natural History Museum No.
30036a-c. Same data as the type. The three paratypes are some-
what smaller than the type, 6.7 mm. to 7.0 mm., and their spire is

Fig. 57. Systrophia (Systrophia) retinella sp. nov. C.N.H.M. No. 30035;
type, shown from side, above and below, about X2.

a trifle higher than that of the type, though the entire height of the
shell does not exceed that of the type.

Systrophia (Systrophia) stenogyra Pfeiffer.

Eighteen specimens from the Cerro Azul, April 12, 1947.

Systrophia (Systrophia) helicycloides Orbigny.
Nine specimens from the Cerro Azul, May 12, 1947.

Systrophia (Systrophia) altivaga Crawford.

One specimen from Divisoria, August 26, 1947; with 12.5 mm.
diameter slightly larger than Crawford's specimens, which measured
910 mm. in diameter.

Systrophia (Systrophia) angigyra sp. nov. Figure 58.

Type.— Chicago Natural History Museum No. 30034, from
Divisoria, Department of Huanuco, altitude about 5,000 feet.
Collected by Jose- Maria Schunke, August 26, 1947.

Diagnosis. — A small species of the typical subgenus of Systrophia,
characterized by the low, almost flat spire, the close convolution of
the S1/^ whorls, and the deep, narrow umbilicus.
I

HAAS: MOLLUSKS FROM PERU 247

Comparisons. — This novelty belongs to the same group within
Systrophia proper, of which the species affinis Pilsbry and obvoluta
sp. nov. (mentioned above) are members. S. angigyra is very close
to both, but differs from them in the still closer convolution of the
whorls and in its small size.

Description of type. — Shell transparent, waxy, pale yellow, almost
planorboid, with scarcely raised spire. Whorls 83^, exceedingly
closely convoluted, separated by a rather deep suture and showing
a dense striation on the upper side while the lower one is smooth.
Last whorl very convex on its upper half, less so on the lower one,

^fc it&k i&fc

Fig. 58. Systrophia (Systrophia) angigyra sp. nov. C.N.H.M. No. 30034;
type, shown from side, above and below, about X2.

suddenly descending somewhat and widening one-fourth of one
whorl before the aperture, with a bump-like swelling that gives the
shell an asymmetrical appearance. Umbilicus narrow, deep, almost
cylindrical, showing a very deep, almost furrow-like suture between
the whorls. Aperture oblique, crescent-shaped, a little bit protracted
in the middle, higher than wide; margin thin, somewhat expanded
all around.

Measurements of type. — Diameter 6 mm., height 2.9 mm., diameter
of umbilicus 1.9 mm.

Systrophia (Systrophiella) peruviana Preston.

Six specimens from Divisoria, August 26, 1947. Apparently this
species has never been found again since its original description
(Preston, 1907, p. 490, fig. 1).

Austroselenites (subgenus?) variegatus sp. nov. Figure 59.

Type. — Chicago Natural History Museum No. 30037, from Cerro
Azul, on Rio Ucayali, Department of Loreto. Collected by Jose*
Maria Schunke, April 20, 1947.

Diagnosis. — A rather solid, scarcely transparent, obtusely conical
shell of 5% whorls with an angulate periphery and a wide, funnel-
shaped umbilicus, with a simple peristome of the aperture and with

248 FIELDIANA: ZOOLOGY, VOLUME 31

the surface covered by a conchinic layer showing alternating, closely
set brownish and yellowish transverse stripes.

Comparisons. — No similar shell seems to be known. The hitherto
known species of the haplotrematid genus Austroselenites are all
much larger. The allocation of this new species to Austroselenites
can, hence, be only tentative. Its real nature will most probably
be known only when its anatomy has been studied; unfortunately
my only specimen does not contain the soft parts.

Description of type. — Shell of medium size, rather solid, almost
opaque, smoothish, depressedly conical above and rather flat below,

Fig. 59. Austroselenites variegatus sp. nov. C.N.H.M. No. 30037; type,
shown from side, above and below, about X2.

with an angle at the periphery. Whorls h%, regularly increasing,
very little convex, separated by a shallow suture. The conchinic
layer on the surface has a pattern of alternating brownish and yellow-
ish transverse stripes of which there are about three to a millimeter
on the last whorl and which are less conspicuous on the under side.
Under side of shell rather flat, hollowed by a wide, funicular um-
bilicus. Aperture oblique, simple, almost triangular, narrow above,
wide and full below.

Measurements of type. — Diameter 10.3 mm., height 5.5 mm.,
diameter of umbilicus 3.9 mm.

Helicina (Helicina) bourguignatiana Ancey.

Five specimens from the Cerro Azul, April 30, 1947. This species
is represented by two phases, a small one (diameter 8.2 mm.) and a
big one (diameter 9.9 mm.).

Helicina (Oxyrhombus) laus ucayalensis A. J. Wagner.

Ten specimens from the Cerro Azul, April 30, 1947. First report
of this subspecies after its description (A. J. Wagner, 1910, p. 290).

HAAS: MOLLUSKS FROM PERU 249

Aperostoma (Aperostoma) depressum Da Costa.

Fourteen specimens from the Cerro Azul, April 20, 1947. Apero-
stoma (A.) peruense and Aperostoma (A.) leai, both described by
Bartsch and Morrison (1942, p. 245, pi. 35, figs. 10-12 and p. 246,
pi. 35, figs. 17-19, respectively), seem to be identical with Neocyclotus
depressus Da Costa (1906, p. 9, pi. 1, figs. 14-16), apparently not
found again since the time it was described.

Ampullarius (Limnopomus) columellaris Gould.

One specimen from Contamana, May 18, 1947, and five from
Metelo Yacu, April 30 and May 14, 1947, respectively.

Mycetopoda soleniformis Orbigny.

Three specimens from the Neshuya River, October 2, 1947.

Mycetopodella falcata Higgins.

One specimen from the Neshuya River, October 2, 1947.

Anodontites (Anodontites) crispatus crispatus Bruguiere.

Seven specimens from Metelo Yacu, May 2, 12 and 13, 1947,
respectively.

REFERENCES

Bartsch, Paul and Morrison, J. E.

1942. The cyclophorid mollusks of the mainland of America. Bull. U. S. Nat.
Mus., 181, pp. 142-293, pis. 19-42.

Crawford, G. I.

1939. Report on the terrestrial mollusca collected by the Percy Sladen Expedi-
tion to Lake Titicaca, 1937. Proc. Mai. Soc. London, 23, pp. 318-332, figs.
28-30, pis. 19-20.

Da Costa, S. I.

1906. Descriptions of new species of Drymaeus, Amphicyclotus, and Neocyclotus,
from South and Central America. Proc. Mai. Soc. London, 7, pp. 7-10, pi. 1.

Jousseaume, F.

1887. Mollusques nouveaux de la republique de l'Equateur. Bull. Soc. Zool.
France, 12, pp. 165-186, pi. 3.

Orbigny, Alcide d'

1835-45. Voyage dans l'Amerique Meridionale (le Bresil, La Republique
Orientale de l'Uruguay, etc. . . .) execute pendant les annees 1826, 1827, . . .
et 1833), 5, pt. 3, Mollusques, 758 pp., 85 pis.

250 FIELDIANA: ZOOLOGY, VOLUME 31

Pfeiffer, Louis

1845-55. Die Gattungen Bulimus, Partula, Achatinella, Achatina and Azeca.
In Abbildungen nach der Natur, mit Beschreibungen. Begonnen von Dr.
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Chemnitz, Syst. Conch. Cab., I, 13, 395 pp., 70 pis.

1856. Descriptions of fifty-eight new species of Helicea from the collection of
H. Cuming, Esq. Proc. Zool. Soc. London, 1856, pp. 324-336.

1860. Novitates Conchologicae. Series Prima. Mollusca Extramarina, 2,
pp. 139-303, pis. 37-72.

Pilsbry, Henry Augustus

1900. New South American land shells. Proc. Acad. Nat. Sci. Philadelphia,

1900, pp. 385-394, pis. 11-12.
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12-17.
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No. 8. Proc. Acad. Nat. Sci. Philadelphia, 84, pp. 387-402, 5 figs., pis. 27-28.

POLINSKI, WLADYSLAW

1925. Sur un nouveau sous-genre et deux nouvelles especes de Clausiliides
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Preston, H. B.

1907. Descriptions of new land and freshwater shells from Central and South
America. Ann. Mag. Nat. Hist., (7), 20, pp. 490-498, 20 figs.

Smith, Edgar A.

1877. Description of new species of South-American Helicidae in the British
Museum. Proc. Zool. Soc. London, 1877, pp. 361-365, pi. 39.

Smith, Maxwell

1943. New species of Cerion, Nenia, and Drymaeus. Nautilus, 57, pp. 59-62,
pi. 7, figs. 7-10.

Wagner, A. J.

1907-11. Die Familie Helicinidae. Martini-Chemnitz, Syst. Conch. Cab., I,
18, (2), 391 pp., 70 pis.