MEMOIRS ve OF THE AMERICAN ENTOMOLOGICAL SOCIETY NUMBER 10 A REVISION OF THE NORTH AMERICAN SPECIES BELONGING TO THE GENUS PEGOMYIA (DIPTERA: MUSCIDAE) BY H. C. HUCKETT PUBLISHED BY THE AMERICAN ENTOMOLOGICAL SOCIETY AT THE ACADEMY OF NATURAL SCIENCES PHILADELPHIA 1941 THE AMERICAN ENTOMOLOGICAL SOCIETY PHILADELPHIA Founded 1859 Incorporated 1862 Organized as ‘‘The Entomological Society of Philadelphia,” the present title was adopted February 11, 1867. PRESIDENT VICE-PRESIDENT Emlen P. Darlington Mark P. M. Robinson CORRESPONDING SECRETARY TREASURER RECORDING SECRETARY Ezra T. Cresson, Jr. James A. G. Rehn John W. Cadbury, 3d. FINANCE COMMITTEE PUBLICATION COMMITTEE Philip P. Calvert Carl Fenninger James A. G. Rehn Philip P. Calvert Raymond C. Casselberry H. Radcliffe Roberts Ezra T. Cresson, Jr. (Editor) Stated meetings are held in January, February, March, April, May, September, October, November and December. 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MEMOIRS OF THE AMERICAN ENTOMOLOGICAL SOCIETY NUMBER 10 - A REVISION OF THE NORTH AMERICAN SPECIES BELONGING TO THE GENUS PEGOMYIA (DIPTERA: MUSCIDAE) ee BY - H. C. HUCKETT MMLC yyy Up TOMO; LAL OX GR PUBLISHED BY THE AMERICAN ENTOMOLOGICAL SOCIETY AT THE ACADEMY OF NATURAL SCIENCES PHILADELPHIA IQ41 Si SOA a> SOPOT NE MEMOIRS OF THE AMERICAN ENTOMOLOGICAL SOCIETY NUMBER 10 A REVISION OF THE NORTH AMERICAN SPECIES BELONGING TO THE GENUS PEGOMYIA (DIPTERA: MUSCIDAE) H. C. HUCKETT * Riverhead, New York The genus Pegomyia Robineau-Desvoidy was originally pro- posed for the reception of a small group of anthomyid flies, the larvae.of which were reported as leaf-miners in certain plants. At the present, the genus stands for a large composite grouping of species, the larval habits of many of which are known to differ widely. For instance the larvae of some North American forms have been found to inhabit the burrows of rodents and land turtles, and in the case of others of wider geographical represen- tation to feed on mushrooms and to tunnel around canes of rasp- berry, whilst as leaf-miners the species are known to inhabit the foliage of an increasing variety of plants, e. g., thistles, convol- vulus, spinach, ragweed and dock. The genus is widely distributed throughout the Northern Hemisphere, and the species are not uncommon in many parts of this area. Nearly one hundred species are herein recorded as occurring in North America. The genus is not readily defined owing to the limited nature of many of the characters possessed by the species. Despite this handicap the genus serves a useful purpose in providing a common ground for the assemblage of a large number of associated groups of species. These minor segregates are frequently more easily distinguished for diagnostic purposes than the genus itself, one of MEM. AMER. ENT. SOC., IO 2 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) the most helpful clues to their recognition being the structure and character of the copulatory appendages in male. The genus, thus composed, impinges naturally at certain points on the boundaries of other genera belonging to the subfamily Anthomyiinae. Some species ascribed to such genera I have transferred to Pegomyia on the evidence provided by the common characters possessed, by the hind tibiae as denoted by the presence of two bristles on antero- dorsal and posterodorsal surfaces respectively, and in the male sex by the lack of a series of setulae on posteroventral surface. The genus may be distinguished from Emmesomyia Malloch and Taeniomyia Stein by the absence of a bristle or setulose hairs on upper border of pteropleura below the wing, and from Paregle cinerella (Fallén) by the absence of a wartlike tubercle on dorsal surface of second antennal segment. Malloch! has pointed out that there may be some pegomyian forms in North America ap- plicable to Stein’s european genus Enneastigma, but as yet no structural characters have been proposed that would warrant their transfer to that genus. SYNONYMY In dealing with the synonymy of palaearctic species occurring in North America I have in large measure restricted the citations to the period following the publication of Stein’s? first monograph on the european species of Pegomyia and of the third volume of Katalog der Palaarktischen Dipteren issued during the following year. More recent treatment of the synonymy may be found in Séguy’s‘ comprehensive work on the Muscidae of the World issued in 1937 as a fascicle of Genera Insectorum, and in Ring- dahl’s® synoptic revision of the Swedish species of Pegomyia pub- lished in 1938. 1 Malloch, J. R. A synopsis of the North American species of the genus Pegomyia Robineau-Desvoidy (Diptera, Anthomyiidae). Bull. Brooklyn Ent. Soc., Xv, pp. 121- 127, 1920. 2Stein, P. Die mir bekannten europdischen Pegomyia-Arten. Wzen. Ent. Zeitg., XXV, Pp. 47-107, 1906. $Stein, P. Katalog der Palaarktischen Dipteren. 111, pp. 700-710, 1907. 4Séguy, E. Genera Insectorum. fasc. 205, pp. 48-67, 1937: 5 Ringdahl, O. Oversikt av svenska Pegomyia-arter (Diptera: Muscidae). Entomol. Tidskr., L1X, (hft. 3-4), pp. 190-213, 1938. H. C. HUCKETT 3 EXPLANATION OF DESCRIPTIVE TERMS The distance separating the eyes is measured at narrowest part of frons: the width of parafacial is measured in profile at its greatest development near base of antennae: the height of cheek is measured below the most ventral extension of eye margin: the postgenal margin denotes the boundary between occipital and genal regions at the caudoventral angle of head: the postvertical pair of bristles is situated on the occipital region caudad of the inner pair of vertical bristles: the third antennal segment is considered yellowish proxi- mad if the yellowish area on inner surface extends distad of a point opposite base of arista; if not, the segment may be invariably considered entirely blackish. The pronotal setulae are situated on the planes of acrostical and dorsocentral bristles along the cephalic border of mesonotum opposite the occipital region of head: stigmatal bristles are situated below mesothoracic spiracle: apical setulae of scutellum are situ- ated between apical pair of bristles: the anal sclerite in male forms that part of hypopygium surrounding the anal membrane: basal sclerite is situated immediately cephalad of anal sclerite: the pre- basal sclerite is placed cephalad of basal sclerite and is in the male of many species obscured from view by the overlapping of tergum 5. The nomenclature of the wings is according to Comstock.*® ACKNOWLEDGMENTS I wish to acknowledge my indebtedness to all those entomolo- gists who have kindly permitted me to use the collections in their charge as a basis for this contribution on the pegomyian fauna of North America. Among the Institutions that have thus generous- ly helped are the Academy of Natural Sciences of Philadelphia [A. N. S. P.], United States National Museum [U. S. N. M.], Entomological Branch of the Canadian Department of Agriculture [C. N.-C.J, Illinois Natural History Survey [Ill. N. H. S.J], New g Comstock, J. H. The wings of insects. pp. 1-430, 427 figs., 1918. MEM. AMER. ENT. SOC., 10 4 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) England Museum of Natural History [N. E. M. N. H.], Field Museum of Natural History, Cornell University [C. U.], American Museum of Natural History [A. M. N. H.], Carnegie Museum and Ohio State Museum. Material has also been submitted for study from the collections of Brigham Young University, Colorado State College, Macdonald College, Quebec, Michigan State Col- lege, Oklahoma Agricultural and Mechanical College, Oregon State College, South Dakota State College, Texas Agricultural and Mechanical College, Utah Agricultural College, and from the Universities of Alberta and Wisconsin. Abroad I have had the privilege of examining North American species in the collections of the British Museum of Natural History, Zoological Museum of the University of Berlin [Z. M. U. B.], Zoological Museum of the University of Copenhagen [Z. M. U. C.], ’S Rijks Museum at Leiden and Muséum National d’ Histoire Naturelle. Reference to the source of such material in the following records has been in- cluded within brackets |], and in many instances the full name has been abbreviated to capital letters as indicated above. I also wish to express my appreciation for the loan of material from the private collections of Dr. A. L. Melander [A. L. M.], A. J. Basinger, F. S. Blanton, G. Steyskal, F. M. Snyder, J. Wilcox, and also for the help thus given by the late Dr. J. M. Aldrich, Professor J. S. Hine and Mr. C. W. Johnson. Material retained in the author’s collection is denoted by the initials [H. C. H.]. To Mr. Oscar Ringdahl I am deeply indebted for assistance and advice with regard to problems relating to the identity of pegomy- ian species common to the fauna of Sweden and North America. PEGOMYIA Robineau-Desvoidy Pegomyia Robineau-Desvoidy, Essai Myod., p. 598, 1830. Macquart, Hist. Nat. Ins., I, p. 351, 1835. Westwood, Intr. Mod. Class. Ins., Suppl., 1, p. 143, 1840. Meade, Descr. List. Brit. Anth., 1, p. 53, 1897. Stein, Berl. Ent. Zeitschr., (1897), XLII, p. 236-249, 286, 1898. Aldrich, Misc. Coll. Smithsn. Inst., XLv1, p. 558-559, 1905. Stein, Wien. Ent. Zeitg., XxXv, p. 47-107, 1906. Stein, Kat. Palaark. Dipt., 11, p. 700, 1907. Williston, Man..N. A. Dipt., 3rd ed. p. 336, 1908. Coquillett, Proc. U. S. Nat. Mus., xxxvu, p. 586, 1910. Schnabl, Hor. Soc. Ent. Ross., XXXIX, p. 105-114, 3 pls., 1910. Schnabl and Dziedzicki, Abh. K. Leop.-Carol. Deutsch. Akad. Naturforsch., xcv, nr. 2, p. 108-112, IQII. Stein, Arch. f. Naturgesch., (1913), Lxx1x, (A), hft. 8, p. 33-36, 1914. Stein, H. C. HUCKETT 5 Arch. f. Naturgesch. (1915), Lxxx1, (A), hft. 10, p. 13, 123-131, 221, 1916. Malloch, Trans. Amer. Ent. Soc., XLIV, p. 295, 300, 1918. Stein, Arch. f. Natur- gesch., (1917), Lxxxi, (A), hft. 1, p. 148, 149, 1919. Malloch, Rept. Can. - Arct. Exped., 1913-18, 11, C, p. 74C, 1919. Stein, Arch. f. Naturgesch., (1918), LXXxIV, (A), hft. 9, p. 63-73, 1920. Malloch, Bull. Brooklyn Ent. Soc., xv, p. 121-127, 1920. Séguy, Faune de France, v1, p. 148-166, 1923. Huckett, Mem. 77, N. Y. (Cornell) Agr. Exp. Station, (1923), p. 40-48, 1924. Enderlein, Fauna Deutschlands, 3rd. ed., p. 341, 1925. Karl, Tierwelt Deutschlands, XIII, pt. 3, p. 122-144, 1928. Ringdahl, Entomol. Tidskr., Liv, p. 2, 1933. Curran, Fam. Gen. N. A. Dipt., p. 393, 1934. Séguy, Gen. Insect., fasc. 205, p. 48-67, 1937. Ringdahl, Entomol. Tidskr., L1x, hft. 3-4, p. 190-213, 1938. Phoraea Robineau-Desvoidy, Essai Myod., p. 600, 1830. Zabia Robineau-Desvoidy, Essai Myod., p. 600, 1830. Chlorina Robineau-Desvoidy, Essai Myod., p. 602, 1830. Anthomyia (Pegomyia) Strobl, Verh. K. K. zool.-bot. Ges. Wien, XLIn, p. 250-252, 1893. Pandellé, Rev. Ent. France, XX, p. 290, 1901. Pegomyza Karl, Tierwelt Deutschlands, x11, pt. 3, p. 121, 1928. Chortophila (Nudaria) Karl, Tierwelt Deutschlands, x11, pt. 3, p. 171, 1928. Chortophila (Nupedia) Karl, Zool. Anzeiger, LXXXVI, p. 174, 1930. GENOTYPE: Musca hyoscyami Panzer. (Designation of Coquil- lett, 1910.) The genus Pegomyia was erected by Robineau-Desvoidy (1830)? for the reception of six nominal species, the larvae of which were recognized as leaf-miners. Of these species only the first, namely Pegomyia hyoscyamt (Panzer), has retained its identity in the writings of subsequent authors. Macquart (1835) enlarged the genus to include the names of sixteen nominal species, and in so doing absorbed the four subse- quent remaining genera of Robineau-Desvoidy’s tribal segregate Pegomydae. . Westwood (1840) selected Anthomyia fulgens Meigen as type of Pegomyia, a species evidently not represented in the original series and hence may be regarded as ineligible for type designation as defined by the Code. No formal recognition of the genus is given in the larger works of Zetterstedt, Siebke, Schiner, Nehaus, Bigot or Rondani. The latter author split the pegomyian species into two groups, placing 7 Refers to literature cited in the synonymy as indicated by the year of publication. MEM. AMER. ENT. SOC., IO 6 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) in Anthomyia those with slightly developed frontal head sclerites in male and prominently extended lower calyptral’scale, and in Chortophala those with extensively molded head parts and incon- spicuous lower scale. Meade (1897) in his classification of the British Anthompidae broadened the concept of the genus to include flies of the sub- family Anthomyiinae with bare arista and yellowish legs, abdomen yellowish or darkened and calyptral scales of equal and unequal development. On the other hand Strobl (1893) and Pandellé (1901) maintained the group as a part of the genus Anthomyia.. Coquillett® in 1901 designated Musca hyoscyam1 Panzer as the genotype of Pegomyia, and later he (1910) broadened the concept of the genus to include the genera Phorbia, Egle, Chortophila of authors, and also other little known groups proposed by Robineau- Desvoidy and Lioy. Schnabl (1910) and Schnabl and Dziedzicki (1911) concluded that the genus Pegomyia contained species which could be distin- guished from those in related groups by the narrower frons in male, and that the group consisted of a number of small segregates in which the basic distinction lay in the structure of the male genitalia and copulatory appendages. The genus was divided into four subgenera, of which Pegoplata and Pegomyza were charac- terized as having plumose arista, Anthomyia with black legs, and Pegomyia with mostly yellowish legs. If Schnabl’s classification is to be adopted the name Anthomyia would necessarily supercede that of Pegomyia for the genus owing to its priority, and as a result it would become necessary to make ID UIESO.82, chang >t in the names applied to higher categories. Stein (1897) (1906) (1907) (1914) in his earlier conteibutions restricted the genus to include species with bare or nearly bare arista, yellowish basal segments of antennae, and yellowish legs; In later years he included species with plumose arista, which, in common with many others belonging to the genus, were charac- terized by the possession of two antero- and two (pose sceotee bristles on hind tibia. ° Se NT Chas Sea A 8 Coquillett, D. W. Types of anthomyid genera. Journ. N. Y. Ent. Soc., 1X, pp. g' 139-140, 1901. H. C. HUCKETT 7 Malloch (1918) (1920) following Stein in his treatment of the genus included allied species with black tibiae that had two postero- dorsal bristles on hind tibia. None of the species in the genus possessed short erect hairs on posteroventral surface of hind tibia in male. This concept of the genus included many species belong- ing to Chortophila and Phorbia of authors, Eremomyza Stein (1920), and more recently Nupedia Karl (1930). Séguy (1923) in his treatment of the French fauna utilized to a large degree the classification erected by Schnabl. The so-called Pegomyinae were restricted to three genera, namely Pegomyia, ‘Anthomyia and Calythea. The former was further subdivided into two subgenera, Pegoplata and Pegomyia. Pegomyza was regarded asa synonym of Pegoplata. Karl (1928), in classifying the German forms, more correctly, in my opinion, maintained the tribal name Anthomyini, in which both Pegomyza and Pegomyyja are given full generic rank. Antho- myia virginea Meigen is given as typical of Pegomyza, of which Pegoplata is mentioned as a synonym. Since the name vwirginea was not included by Schnabl and Dziedzicki (1911) in the original series under Pegomyza I am doubtful whether that species may be considered available for type designation unless it can be shown more conclusively that virginea is conspecific with one or more of the forms listed in Pegomyza. The group Nupedia (Nudaria preoc.) was erected by Karl (1930) for the reception of dark-legged forms as exemplified in Anthomyia dissecta Meigen. Stein (1916) had previously included such species in Chortophila. More recently Séguy (1937) in his comprehensive monograph on the genera of the family Muscidae has largely followed Stein in his citation of the characters definnig the genus Pegomyia. The groups Pegomyza and Pegoplata are contained within the meaning of the genus. The type of Pegoplata is cited as Pegomyia palpata Stein, the first name given by Schnabl and Dziedzicki (1911) in the series of species belonging to the group. Stein (1916) has drawn attention to the fact that the species was misnamed by Schnabl, the proper name for which should have been Hydrophoria palposa Stein, a species closely allied to virginea Meigen. Since palpata and palposa represent widely separated groups of species MEM. AMER. ENT. SOC., 10 8 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) within the genus the character and relationship of the group Pegoplata will depend largely on the decision concerning which species may be accepted as the valid type. Ringdahl (1938) in his review of the Swedish species has divided the genus Pegomyza into eight subgenera, namely Pegomyza with type Anthomyia praepotens Wiedemann, Pegoplata with type Anthomyia virginea Meigen, Arctopegomyia Ringdahl with type Anthomyza tunicata Zetterstedt, Chaetopegomyia Ringdahl with type Anthomyia setaria Meigen, Emmesomyia Malloch with type E. unica Malloch, Phoraea R.-D. with type Anthomyia silacea Meigen, Pegomyia R.-D. with type Musca hyoscyami Panzer and Pegomyiella Ringdahl with type Anthomyza lunatifrons Zetter- stedt. All these groups with the exception of Emmesomyia, which I regard as entitled to generic rank, and possibly Phoraea, are represented in the North American fauna as herein restricted. The true significance of such groupings in this instance however cannot be properly evaluated with any degree of accuracy for lack of the opportunity to make the necessary comparative studies. SUMMARY OF FAUNAL RECORDS One of the earliest records extant of the capture of pegomyian flies in North America is to be found in the New England collec- tions of William Harris. According to Johnson® specimens taken by Harris were submitted to Say in 1833 for identification, and among the specimens were several that had received the manu- script name Anthomyia lenis. Johnson, in studying the collection later, recognized two species among the material represented as lenis, namely Alloeostylus diaphanus (Wiedemann) and Pegomyia geniculata (Bouché). Early references to the occurrence of pegomyian species in North America are to be found among Walker’s!° published records of 1849. Such congeneric forms as Anthomyia hpsia and Eriphia flavifrons were captured in the Hudson Bay Territory, Coenosia substituta from Massachusetts. The latter species was shown by *Johnson, C. W. Diptera of the Harris Collection. Proc. Boston Soc. ‘Nat. Hist., XXXVIII, pp. 92-93, 1925. 10 Walker, F. List of the specimens of dipterous insects in the collection of the British Museum, Iv, pp. 928, 966, 971, 1849. H. C. HUCKETT 9 Stein"! to be identical with ipsta. In 1867 van der Wulp” de- scribed the pegomyian species Anthomyia tarsata from a collection of flies taken in Wisconsin. In 1878 Meade® published a synopsis of the results of his examination of a collection of North American Anthomyiidae sent to him by Osten Sacken, in which amongst others he recognized specimens that reminded him of such european species as Anthomyza gilva, A. vittigera and A. flavoscutellata of Zetterstedt. Hagen published a fuller report of the material examined by Meade when that collection had been returned by Osten Sacken. to the Museum of Comparative Zoology at Cam- ‘bridge. This later list included the species Anthomyza latitarsis Zetterstedt, a record omitted in Meade’s previous communication. Osten Sacken* did not mention the genus Pegomyza in his cata- logue of diptera from North America, preferring to retain the names of species as originally designated by the various authors. In 1898 Stein!® recorded thirteen nominal species in Pegomyia from the collections of Aldrich, Johnson and Nason. Of these forms I consider that trilineata and trivittata belong properly to Hylemyia, and setosa should be placed in Malloch’s genus Ere- momyoides. The remaining records consist of seven new native species and three european. Among the latter is included the species calyptrata (Zetterstedt), a record which in my opinion was based mistakenly on material representing another species, name- ly triseta Malloch. In 1905 Aldrich!’ following closely Stein’s records cited the names of thirteen nominal species in Pegomyia 1 Stein, P. Die Walker’schen aussereuropdischen Anthomyiden in der Sammlung des British Museum zu London. Zettschr. f. Hymen. u. Dipt., 1, (hft. 4), p. 212, 1901. 12. Van der Wulp, F. M. Eenige Noord-Americaansche Diptera. Tydschr. v. Ent., x, Pp. I5I, 1867. 13 Meade, R. H. Notes on the Anthomyiidae of North America. Ent. Month. Mag., XIV, pp. 250-252, (1877-1878), 1878. 14 Hagen, H. A. List of N. American Anthomyidae, examined by R. H. Meade, Esgq., Bradford, England. Can. Ent., xi11, pp. 43-51, 1881. 18 Osten Sacken, C. R. Catalogue of the described Diptera of North America. Smithsn. Miscel. Coll., 111, pp. 167-170, 1878. 16 Stein, P. Nordamerikanische Anthomyiden. Berl: Ent. Zeitschr., XLu, pp. 236- 249, (1897), 1898. ij W Aldrich, J. M. A catalogue of North American diptera (or two-winged flies). Smithsn. Miscel. Coll., XLV1, pp. 558-559, 1905. MEM. AMER. ENT. SOC., IO IO NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) from North America. In this list the names conformis (Fallén) and vicina Lintner are retained for species that are identical with hyoscyamt (Panzer). The species Pegomyia nitidula Coquillett I regard as allied to Chortophila sepia (Meigen), and hence does not belong to Pegomyza. There are twenty two names of North American species of Pegomyia given by Stein!’ in his analytical study of the noneuro- pean species of Anthomyiidae. Among these names are three that are included as synonyms, three that have heretofore been included in Hylemyia, and one that is doubtfully retained. Among the species recorded are Pegomyia bucculenta Coquillett which I regard as identical with Hylemyia brassicae (Bouché), Spilogaster socialis Stein as a species of Emmesomyia Malloch, and Anthomyia tarsata (van der Wulp) which I consider a valid species although listed as a synonym of lipsia (Walker). In 1920 Stein!® published his second contribution on North American Anthomyiidae, in which is presented a key to twenty seven species and one variety of Pegomyia. Seven species are described as new, of which I regard fuscinervis as identical with duplicata Malloch, and setigera as belonging to the genus Hylemyia. The european names gilva (Zetterstedt) and bivittata Stein are given for species that I consider have appeared in North American literature under the names /uteola Malloch and lativittata Malloch respectively. The european species P. squamifera Stein, which is regarded by Ringdahl as a synonym of P. rufina (Fallén), is re- corded for the first time as occurring in North America. Stein has described three new forms under Chortophila, namely glabra, latipalpis, nigroscutellata, that I have transferred to Pegomyia on the strength of characters presented by the bristling on hind tibiae. Stein, P. Die Anthomyidengattungen der Welt, analytisch bearbeitet, nebst einem kritisch-systematischen Verzeichnis aller aussereuropadischen Arten. Arch. f. Naturgesch., LXxxu, (A), hft. 1, pp. 148-149, (1917), 1919. 19 Stein, P. Nordamerikanische Anthomyiden. 2. Beitrag. Arch. f. Naturgesch., LxxxIVv, (A), hft. 9, pp. 63-74, 88-91, (1918), May 1920. H. C. HUCKETT II Malloch®* in the same year published a key to thirty nine North American species of Pegomyta, of which twenty had been recently described by him from material collected in various parts of the United States and in Alaska. The key includes thirty two native species and seven european. The species fuscofasciata Malloch is in my opinion the true winthemi of Wiedemann, and Malloch’s records of winthemi as well as those in many recent faunal lists refer to another species that I have recently renamed mallocht. Cole and Lovett?! in their catalogue of Oregon diptera refer only to three species of Pegomyia known to occur within the State. ‘Huckett” recorded twenty species in the genus from New York, including the western forms vanduzeet and acutipennis. The former had been recently studied by Frost” as a leaf-miner of dock under the name affinis Stein. Johnson: * gave a list of eighteen species of Pegomyia in his faunal records for New England, and thirteen as occurring in the Mount Desert Region, including five species which had been originally described from that area by Malloch. Leonard** in 1928 recorded twenty two species in the genus in a list of insects for New York. The faunal pattern closely follows that given by Johnson for New England. Séguy?? in The Genera Insectorum, on the Muscidae of the World lists forty four nominal species under Pegomyia that occur in North America, and one is doubtfully included. The species 20 Malloch, J. R. A synopsis of the North American species of the genus Pegomyia Robineau-Desvoidy (Diptera, Anthomyiidae). Bull. Brooklyn Ent. Soc., XV, pp. 121I- 127, 1920. 21 Cole, F. R. and A. L. Lovett. An annotated list of the Diptera. (Flies) of Oregon. Proc. Cal. Acad. Sci., X1, Pp. 313, 1921. 2 Huckett, H. C. A systematic study of the Anthomyiinae of New York, with especial reference to the male and female genitalia. N. Y. (Cornell) Agric. Exp. Station, Mem. 77, pp. 40-48, (1923), 1924. 23 Frost, S. W. Two species of Pegomyia mining the leaves of dock. Journ. Agr. Research, XV1, pp. 229-243, 1919. 24 Johnson, C. W. Fauna of New England, xv. List of the diptera or two-winged flies. Occ. Pap. Boston Soc. Nat. Hist., vil, pp. 232-233, 1925. 2 Johnson, C. W. Biological Survey of the Mount Desert Region, part I, pp. 209— 210, 1927. 26 Leonard, M. D. A list of the insects of New York, with a list of the spiders and certain other allied groups. NN. Y. (Cornell) Agr. Exp. Station, Mem. 101, pp. 840-841, (1926), 1928. 27 Séguy, E. Genera Insectorum, Fasc. 205, pp. 48-67, 1937. MEM. AMER. ENT. SOC., IO I2 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) flavipes, stlemba and virginea I have yet to recognize as occurring in this region; the species nztidula, setigera, trilineata, I consider as belonging to Hylemyia. The latter genus as treated by Séguy contains the records of such North American species as dissecta, glabra, juvenilis, latipalpis, nigroscutellata, pedestris and rubivora, all of which I have included in my treatment of the genus Pego- myta. There are eleven North American species of Pegomyza in- cluded inadvertently in the genus Pogonomyia Rondani, namely emmesia, fringilla, fuscofasciata, labradorensis, luteola, quadri- spinosa, spinigerella, subgrisea, triseta, unguiculata, and untcolor. Huckett?® has recently published the names of thirty one new or little known pegomyian species from North America, one of which is recognized as occurring also in Sweden. The present study records the occurrence of ninety seven species of Pegomyia sens.-lat. in North America, of which sixty nine are recognized as native species, twenty seven as having first been described from Europe and one from Siberia. Key to Groups 1. Arista with hairs longer than setulae on second antennal segment........... 2 Arista nearly bare or with hairs not longer than setulae on second antennal SESITIENE + & busn.cticuat ) eee ge ee oe oie ee ee eee 3 2. Apical setulae of scutellum hairlike, cruciate bristles present, processes padded with setulae on proximal half of inner border, caudal pair of ocellar bristles in female short and directed forward...................... Virginea Group Apical setulae of scutellum coarser developed than hairs on ventral surface, cruciates absent or weakly developed, processes not padded with setulae, caudal pair of ocellar bristles in female longish and directed outwards...... Bee ME eT Re oe aE 8 OG cn ok ie AOS thon we's ole Connexa Group 3. Scutellum with three pairs of strong bristles, mesopleura with a few setulae on declivity dorsad of mesothoracic spiracle, posterior notopleural bristle with afew. setulae at base sci. dnc, cca satan tee oe ee Major Group Scutellum with two pairs of strong bristles, mesopleura with no setulae on declivity dorsad of mesothoracic spiracle, posterior notopleural bristle with no:.setulde at: bases 2 20% 2.30 Rhea oi nO PEE One OES Serene 4 Huckett, H. C.. Descriptions of new North American Anthomyiidae belonging to the genus Pegomyia Rob.-Desv. (Diptera). Trans. Amer. Ent. Soc., LXV, pp. I-37, 1939. H. C. HUCKETT 1g . Costal thorn stoutly developed, costal setulae coarsely developed, at least in female, abdomen in male cylindrico-conical, hypopygium and copulatory appendages structurally similar to those of ipsia, m—cu cross vein invariably oblique and sinuate, if not, prealar bristle absent or long and robust...... 5 Costal thorn and setulae inconspicuously and finely developed, abdomen in male usually depressed, hypopygium and copulatory appendages structurally _ not similar to those of lzpsia, m-cu cross vein frequently nearly straight and erect, if oblique prealar bristle is present and shorter than length of anterior MOtOp] eurcalyb rast] ppm ee le es oases si sea ae seieenseeh eeuewe arene oer see waneuae 6 . Prealar bristle absent, second or lower posthumeral bristle absent, basal node of veins R.o+3 and R.s+5 usually with a few hairs on under surface Affinis Group Prealar bristle present, long, second or lower posthumeral bristle usually present, basal node of veins R.o+3 and R.gs bare..............--000- Lipsia Group . Apical setulae of scutellum hairlike, fifth sternum in male armed and shaped as in dissecta (fig. 61), apical anterodorsal bristle of hind tibia proportionately short, cruciate bristles present....................0..00- Dissecta Group Apical setulae of scutellum coarser developed than hairs on ventral surface, or sometimes entirely absent, fifth sternum in male not armed nor shaped as in USSCCLO PTT ee TMT TE TR oon Pack eS Sco. ae da AO cue haat ace uta rare a . Male with numerous stout bristles on prebasal sclerite of hypopygium, female with caudal sclerites of ovipositor strongly chitinized and polished, appressed corsovemtrally memes cots Se ne aie 0 hs so cl sien ts ane Rubivora Group Male with no bristles present on prebasal sclerite of hypopygium, female with caudal sclerites of ovipositor nct noticeably appressed dorsoventrally, if apparently so, not extensively nor strongly chitinized................... 8 . Head with parafacials, parafrontals and cheeks extensively molded, as in hyoscyami, lower portion of occiput swollen, hind femur in male with bristles forming anterodorsal series not sharply differentiated into a proximal series of finer and shorter bristles and a distal series of stouter and longer bristles, Ovipositor short, broadly conical, sclerites weak on dorsal surface and ex- tensively setulose on ventral surface, segments concolorous with abdomen. DB VOakceo ste) CPN Case GHRSST ORG CRETE Re ee eT EEC OO ROC EVP aA Hyoscyami Group Head with parafacials and parafrontals repressed and narrow, cheeks restricted and foreshortened caudad owing to the advanced position of the postgenal margin, e, g., winthemi, geniculata, lower portion of occiput not swollen, hind femur in male with bristles forming anterodorsal series much weaker and more closely set on proximal than on distal half, ovipositor with segments elongated and sclerites invariably slender and strongly chitinized, segments not concolorous with abdomen......................... Flavipes Group MEM. AMER. ENT. SOC., 10 I4 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) Virginea Group There are two species belonging to this group recorded from North America, namely palposa Stein and juvenilis Stein. They would undoubtedly fall within the concept of Pegomyza of authors. The group is delimited by a combination of the following charac- ters; Arista plumose, apical setulae of scutellum hairlike, cruciate bristles present, palpi flattish and broadened, processes of fifth abdominal sternum in male padded with setulae, basal sclerites of hypopygium polished, caudal pair of ocellar bristles in female short and directed cephalad. Connexa Group The species connexa Stein has usually been associated with juventlts on account of plumosity of arista, thoracic markings and highly shining hypopygium in male. There are however a number of characters in which the two species differ, that serve to em- phasize the artificiality of such a relationship. I am of the opinion that connexa and the european species socculata Zetterstedt belong to the same group. For diagnostic purposes both species may be described as having the arista subplumose, cruciate bristles absent, palpi slender, hypopygium and processes as in male of connexa, hypopygium highly shining, hind tibia in male with a short bristle on proximal half of posterior surface, female having discal bristles on fifth abdominal tergum and caudal pair of ocellar bristles longish and directed outward. Major Group The species belonging to this segregate are closely linked to the lipsia-group. They have been included in the North American genus Eremomyia by Stein. I am doubtful whether this genus can be maintained apart from Hylemyia, and have in this instance relegated the species with two antero- and two posterodorsal bristles on hind tibia to the genus Pegomyia sens.-lat. The group possesses the following combination of characters as found in major; Hind tibia with two antero- and two posterodorsal bristles H. C. HUCKETT 15 and a robust apical anterior bristle, scutellum with three pairs of strong bristles, wings with robust costal thorn and strongly oblique, sinuate m—cu cross vein, mesopleura with a few setulae on the declivity dorsad of mesothoracic spiracle and with the stigmatal bristles below spiracle largely devoid of accessory setulae, except in thrixia, posterior notopleural bristle with a few setulae at base, vibrissal angles armed with numerous (10-16) coarse black setulae, cruciate bristles absent, hypopygium and copulatory appendages of male as in /zpsia, female with caudal pair of ocellar bristles long and directed outwards, marginal bristles of abdomen weakly de- veloped. Species belonging to this group are major (Malloch), aincompleta (Stein), thrixia Huckett, assimilis Huckett. Lipsia Group The group contains a large number of North American species belonging to the genus Pegomyia. One of the commonest forms is P. lipsia (Walker), which may be regarded as typical of the group. These species may be defined as having the abdomen of male cylindrico-conical, with hypopygium and copulatory appendages similar to those of ipsia. The abdomen in both sexes has the bristles stoutly developed on dorsum and vestiture comparatively short and uniform, the bristles of legs are strongly developed, hind tibiae are armed on antero- and posterodorsal surfaces respectively with usually only two long bristles exclusive of the apical bristles, wings possess a robust costal thorn and an oblique and markedly sinuate m—cu cross vein, cruciate bristles absent, caudal pair of ocellar bristles longish in female, erect and directed outward. The segregate is closely allied to the major-group, differing in that the species have no setulae on mesopleural declivity dorsad of meso- thoracic spiracle nor at base of posterior notopleural bristle. The group includes the following species: lipsia (Walker), lavidiventris Huckett, chrysida Huckett, tarsata (van der Wulp), cresca Huckett, duplicata (Malloch), longimana (Pokorny), setiformis Huckett, fuscicauda Huckett, apicalis (Stein), costalis (Stein), flavicans (Stein), substriatella (Malloch), quadrispinosa (Malloch), frigida (Zetterstedt), banffi Huckett, anorufa Stein, labradorensis Malloch, partita Huckett, caduca Huckett. It is possible that the last six MEM. AMER. ENT. SOC., IO 16 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) species and the european form semirufa Ringdahl may be entitled to separate group recognition. In general the male abdomen of the latter six species is more depressed and the m-—cu cross vein is only slightly oblique and sinuate. A ffiinis Group The common North American species Pegomyia affinis Stein serves as a typical example of a small group that is known to have habits peculiarly different to those of flies belonging to other segregates within the genus. Adults of affinis may usually be found in the vicinity of rodent’s burrows.2° Malloch®* has reported having captured spinigerella in dry sandy places frequented by burrowing bees. From rearings it is known that larvae of gopheri feed on the excrement found in the burrows of land turtles, Go- pherus polyphemus.*! Characters pertaining to the abdomen and legs of the adult fly have much in common with those of lzpsia- group. In the male the abdomen is cylindrico-conical and hypo- pygial sclerites conform to the tapering outline of the caudal seg- ments, processes are simple, unarmed, with a few bristles along outer border, inner border with a few scattered setulae, marginal bristles on terga are robust on dorsum and vestiture of setulae is short and appressed, costal thorn is typically robust and costal setulae coarsely developed in female, in many species the basal node of veins R..., and R.., has a few hairs on under surface, cross veins clouded and m-cu cross vein slightly oblique and sinu- ate, abdomen not infrequently variegated or partly yellowish in color, prealar and second posthumeral bristles absent, acrostical bristles setulose, closely biserial, and third antennal segment near- ly as broad as long. Species included in this group are affinis Stein, finitima Stein, gophert Johnson, spinosissima Stein, spint- gerella Malloch. 2 Curran, C. H. Note on Pegomyia affinis Stein (Diptera, Muscidae). Can. Ent., LVIII, p. 256, 1926. 30 Malloch, J. R. Descriptions of new North American Anthomyiidae (Diptera). Trans. Amer. Ent. Soc., XLVI, p. 179, 1920. 31 Johnson, C. W. Insects of Florida. I. Diptera. Bull. Amer. Mus. Nat. Hist., XXXII, pp. 77-78, 1913. H. C. HUCKETT : 17 Dissecta Group The group is represented by nine species, of which dissecta (Meigen) is typical. It may be described as possessing the follow- ing combination of characters, which although variable may be regarded as sufficiently characteristic of the group to be of some value for diagnostic purposes: Apical setulae of scutellum hairlike, conforming in structure to the ventral hairs, palpi flattish, cruciate bristles present in both sexes, apical anterodorsal bristle of hind tibia proportionately short, m—cu cross vein erect, lower surface of costa adjacent costal cell (2C.) nearly devoid of setulae, wings in male pointed apicad, processes of fifth abdominal sternum armed and shaped as in dissecta, eyes in male closely approximated at narrowest part of frons, with a pair of parafrontal setulae present near anterior ocellus. The following species are known to occur in North America, a) tibiae black, dissecta (Meigen), pseudodissecta (Ringdahl), nigroscutellata (Stein), latipalpis (Stein), cuticornis Huckett, b) tibiae yellow, acutipennis Malloch, abnormis Stein, anabnormts Huckett, rectifrons Huckett. Rubivora Group The species associated with rubivora and dissecta have been placed in Phorbia and Chortophila respectively by many authors owing evidently to their black legs and subnude arista. It is ap- parent from an examination of North American material that there are species related to rubivora and dissecta that have yellowish tibiae, and which may be included in the respective groups on the basis of characters pertaining to the hypopygium, copulatory appendages and ovipositor. In the males of the rubivora-group the processes are subshining and lined along inner margin by a series of setulae, the prebasal sclerite is well bristled, in the female the caudal sclerites of ovipositor are shiney and strongly chitinized, being appressed dorsoventrally. The darker colored forms of the group may be further distinguished from species belonging to the dissecta-group by the proportionately greater size of third antennal segment, coarser development of apical setulae on scutellum, denser hairs on under surface of costa, absence of cruciate bristles MEM. AMER. ENT. SOC., 10 18 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) in male and their absence or weak development in female. Further in the female the caudal pair of ocellar bristles is not long and is directed cephalad, and the fifth tergum has comparatively stronger discal and marginal bristles than in females of dissecta-group. The paler forms of the rubtvora-group exhibit certain affinities to triseta and related forms in hyoscyami-group, from which they differ basically on account of abdominal characters as defined above. The species are as follows: a) tibiae black, rubivora (Coquillett), b) tibiae yellowish, glabra (Stein), cedrica Huckett. To these may also be added the european forms Chortophila dentiens Pandellé and C. rubricola Enderlein. Hyoscyami Group This is a large rather heterogeneous group consisting mostly of species known to mine the leaves of vegetation, e. g., hyoscyami, vanduzeet, bicolor, nigritarsis, albimargo. ‘These five typical species are characterized by having the parafrontals and parafacials full and prominent in profile, cheeks invariably broadly maintained caudad, ventral half of occiput swollen, arista subnude, cruciate bristles absent, abdomen in male slightly depressed, basal sclerite of hypopygium proportionately robust, giving the abdomen a truncated appearance, anal sclerite (ninth tergum) broadly round- ed when viewed from above, gonostyli (inferior forceps) short, broad, deeply fissured and incised, processes weakly bristled, ovi- positor short, broadly conical, dorsal sclerites weakly chitinized and ventral sclerites minutely setulose, segments concolorous with abdomen, anterodorsal bristles of hind femur in male not sharply differentiated into a proximal series of finer and shorter bristles and a distal series of stouter and longer bristles, nor is the series of posterior setulae on hind femur so extensively developed apicad as in the flavipes-group, wings with costal thorn and setulae weakly developed, m—cu cross vein semierect and nearly straight, caudal pair of ocellar bristles short and directed cephalad in female. Hyoscyamt Subgroup.—Processes frequently shiny and bare apicad, antennae proportionately short, face and oral margin con- stricted, ventral region of occiput swollen, prealar bristle weak or absent, ventral bristle of caudal pair of sternopleural bristles fre- H. C. HUCKETT 19 quently short in female, mesonotum not quadrivittate in male. Species include hyoscyami (Panzer), carduorum Huckett, striata Stein, 7adicta Huckett, atlantis Huckett, convergens Huckett, ruficeps Stein, rufescens Stein, minuta Malloch, cognata Stein, vicaria Huckett, marginata Huckett. Vanduzeer Subgroupb.—Processes with a tuftlike series of setulae at base and at apex on inner margin, widely spaced apart at base, arista swollen at base, prealar bristle longer than in hyoscyami- or albtmargo-subgroups, ventral half of occiput swollen, ventral bristle of caudal pair of sternopleural bristles moderately long in female. ‘Species include, vanduzeet Malloch, haemorrhoa (Fallén), and the european form versicolor (Fallén). Albimargo Subgroup.—Processes bare distad, with fine setulae restricted to basal area, apex of inner margin produced, antero- dorsal series of bristles and posterior series of setulae on hind femur of male more markedly differentiated and extended respectively than in allied subgroups, presutural acrostical bristles well de- veloped, mesonotum in male with four linear markings when viewed from behind, situated in the planes between acrostical and dorsocentral, and dorsocentral and intraalar series of bristles re- spectively, arista not swollen at base, abdomen more distinctly depressed in male. Species include flavifrons (Walker), sitiens Huckett, albimargo (Pandellé) and the european form esuriens (Meigen). Bicolor Subgroup.—Processes restricted, bare and rounded distad, with setulae confined to basal region and in a sparse series along inner margin, hind femur stoutly bristled, presutural acros- tical bristles weakly developed, in two adjacent series, in female the ventral sternopleural bristle of caudal pair is well developed and frontal vitta proportionately narrower than in female of allied subgroups. Species include bicolor (Wiedemann), jacobi Malloch, triseta Malloch, setarza (Meigen), variegata Huckett. Nigritarsts Subgroup.—cClosely allied to bicolor-subgroup, dif- fering in that the processes are broad laminate plates with a dense series of setae along inner margin. Species include nigritarsis (Zetterstedt) and rumictfoliae nom. nov. (p. 81). MEM. AMER. ENT. SOC., IO 20 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) Flavipes Group This group consists of three smaller segregates which may be distinguished from one another by the structure and bristling of the processes in male sex, as represented in gilva (Zetterstedt), geniculata (Bouché) and intersecta (Meigen) respectively. The group characters are most evident in the male, where they may be described as follows: Abdomen depressed, with lateral margins subparallel and terga clothed with longish erect setulae, abdominal vitta if present composed of lineal sections, hind femur with bristles forming anterodorsal series much weaker and more closely set on proximal than on distal half, posterior surface with median series of fine setulae extending distad to preapical bristle, head with parafrontals, parafacials and cheeks reduced to slender and narrow proportions, eyes nearly contiguous at narrowest part of frons, oral margin between vibrissae not constricted, ventral region of occiput not buccate nor swollen, antennae moderate to large, wings pointed apicad, costa slender and with setulae and thorns weakly developed, m—cu cross vein slightly to moderately oblique, Ovipositor with sclerites invariably strongly chitinized and seg- ments elongated. _ Flavipes Subgroup.—Hypopygium considerably enlarged by the angular broadening of the ninth tergum caudad (ventrad), par- ticularly in the region of gonostyli and cerci, fifth sternum com- posed of a large basal plate from which the processes extend as slender pliant lamellae, the latter bear a series of prominent bristles which often appear flaccid, the series extends from base to near apex, otherwise the fifth sternum is largely devoid of bristles, processes are markedly constricted at junction with basal plate through the intrusion of the outer conjunctival membrane, in many of the species the caudal sclerites of ovipositor are rigid and compressed as if for piercing, and marginal bristles of fifth tergum in female are robustly developed laterad. Species are gélva (Zetterstedt), pilosa Stein, tenera (Zetterstedt), incisiva Stein, rufipes (Fallén), vittigera (Zetterstedt). Geniculata Subgroup.—Hypopygium and processes inconspicu- ous, the latter clothed with numerous short bristles throughout, H. C. HUCKETT 21 acrosticals in two widely separated series, antennae large. Species are geniculata (Bouché), univittata (von Roser), unicolor Stein, winthemt (Meigen), mallocht Huckett, rufina (Fallén), flavipalpis (Zetterstedt), corrupta Huckett, pollinosa Ringdahl, longicornis Huckett. Intersecta Subgroup.—Abdomen in male tapering caudad, hypo- pygium subshining, ninth tergum angular and protruded, armed dorsad (cephalad) with a few weak bristles, basal sclerite polished, processes horney, bare and laminate, cruciate bristles present in both sexes. Species are intersecta (Meigen), unguzculata Malloch, tacta Huckett, solitaria Stein. The following species have not been associated with any group, alticola Huckett, analis Schnabl, caesia Stein, littoralis Malloch, lunatifrons (Zetterstedt), marginata Huckett, palpata Stein, tinc- tusquama Huckett. Key to Species of Pegomyia Males I. Legs entirely black, at most knees yellowish tinged..................... 2 Legs at least partly yellowish, occasionally tibiae fuscous...............18 2. Mid tibia with a mid ventral or anteroventral bristle.................... 3 Mid tibia with no mid ventral or anteroventral bristle................... 5 3. Antennae separated by a prominent rounded facial elevation, basal hairs on outer margin of calyptrae blackish tinged, apical setulae of scutellum hair- like as those on ventral surface, propleura with a few hairs near center... . BLESS B20 Bec a gh Bea EE Ee ore rR Se crene Maeee cere eter ea Paregle aestiva (Meigen) Antennae not separated by a prominent rounded facial elevation, basal hairs on outer margin of calyptrae not blackish tinged, apical setulae of scutellum coarser developed than hairs on ventral surface, propleura bare........ 4 4. Prealar bristle short, second antennal segment with a small wartlike tubercle on dorsal surface, oral margin protruded cephalad, hind femur with no bristles on posteroventral surface.............. Paregle cinerella (Fallén) Prealar bristle long, second antennal segment with no wartlike tubercle on dorsal surface, oral margin not protruded cephalad, hind femur with bristles on posteroventral surface......................-. analis Schnabl . Arista plumose, processes with mattlike pad of setulae on inner border, lower calyptral scale extensively protruded beyond margin of upper, prealar bristle not shorter than length of anterior notopleural bristle. palposa (Stein) On MEM. AMER. ENT. SOC., 10 22 NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) Arista short pubescent or bare, processes with no mattlike pad of setulae along inner border, lower calyptral scale not extensively protruded beyond margin of upper scale, prealar bristle shorter than anterior notopleural [oy g (15 (remem i aunte Ata ser eee Oe ING Mceerete Ote MELE DRS Gaal Dis oko oo 0.0 6 6 6. Hypopygium shining black, processes devoid of bristles and are extended as thin chitinous plates (as in wnguiculata) which are prolonged at apex into an accessory digitlike appendage (fig. 55)............ miersecta (Meigen) Hypopygium not polished black, processes with a few setulae and not term- inating in an accessory appendage. ...............0.0.0ceepsesesaees 7 7. Head with inner pair of vertical bristles erect and bristlelike, frontal vitta ribbandlike throughout its length to ocellar callosity, velvety black, cruciate setae longish, hind tibia with apical posterodorsal bristle well developed, as long as apical dorsal bristle.............. littoralis Malloch Head with inner pair of vertical bristles setulose, cruciates weak or absent, frontal vitta largely constricted or obliterated caudad by parafrontals, hind tibia seldom with apical posterodorsal bristle as long as apical dorsal bristle 8 8. Apical setulae of scutellum hairlike as those on ventral surface, hind tibia with apical anterodorsal bristle short, not longer than height of hind metatarsus, processes like those of dissecta (fig. 61)... 0... cece ce eee eens 9 Apical setulae of scutellum coarser developed than ventral hairs, hind tibia with apical anterodorsal bristle invariably slightly longer than height of hind metatarsus, processes not like those of dissecfa.................. 13 g. Cheek as high as width of third antennal segment, margin of eye scarcely reaching a level below the vibrissae, ventral border of cheek with a coarse series of setulae and slender bristles...............6...00-0 000 e eeu 10 Cheek narrower than width of third antennal segment, margin of eye extend- ing ventrad of level with oral vibrissae, cheeks with a single series of bristles along ventral" border e 2c ee ee ne Be eee eee ares II 10. Cross vein m-cu oblique, ultimate section of M.1+2 shorter than 1.5 times length of penultimate section, anterior sternopleural bristle as robust in development as ventral bristle of caudal pair, hind femur with bristles on distal half of anteroventral surface longer than those on proximal half...... Bee eee ye eat erst th tus otiranih scsi dean 6 acute O.did-e8c cuticornis Huckett Cross vein m-cu erect, ultimate section of M.142 fully as long as 1.5 times length of penultimate section, anterior sternopleural bristle slightly finer developed than ventral bristle of caudal pair, hind femur with a series of uniformly long bristles throughout anteroventral and proximal half of posteroventral surfaces...................008- pseudodissecta (Ringdahl) 11. Anterior sternopleural bristle finer developed than ventral bristle of caudal pair, occipital setulae on ventrocentral plane adjacent oral margin stiff and 1G 14. 15. 16. 17h H. C. HUCKETT 23 spinulose, postsutural area of mesonotum when viewed from behind with three broad vittae which are continued to base of scutellum............ ee C/A PEON Per au Si CIE EN SRY AIRS geet Dor pee 3 dissecta (Meigen) Anterior sternopleural bristle as robust as ventral bristle of caudal pair, occipital setulae on ventrocentral plane slender, postsutural area of meso- notum with lateral vittae at least obscure or obliterated before reaching asexolescutellluimaegy. teste eases ACs cates Riis 6% wis te RA Se NL Se 12 . Stigmatal bristles below mesothoracic spiracle with five or six setulae at base, mesonotum brownish black, under surface of costa bordering costal cell (QC) WAR Fehon g plows ceaccon a rote Ree ea eee latipalpis (Stein) Stigmatal bristles seldom with more than one or two setulae at base, meso- notum grayish black, under surface of costa bordering costal cell has a few SCCUlACH RE a oA Se AUR ae awa he nigroscutellata (Stein) Prebasal sclerite of hypopygium prominent and bristled, eyes separated by a distance not less than that between posterior ocelli inclusive.......... 14 Prebasal sclerite concealed and bare, eyes seldom separated by a distance as great as that between posterior ocelli................00 cece ee eeeee 15 Parafacials narrower than breadth of third antennal segment, processes truncated at apex, inner margin fringed with a continuous series of setulae F ih 9 0:0 :4.4 ONO ASN HES BLO LO OE ETRE ET ae ae rubivora (Coquillett) Parafacials fully as broad as width of third antennal segment, processes _ tapering apicad, inner margin with a partial series of setulae, becoming discontinueddbasadhrnarnccaaemine sacs eosin aa tat weeint glabra (Stein) Mesonotum, viewed from behind, with four short postsutural stripes, situated respectively between the series of acrostical and dorsocentral, dorsccentral andsintradlambnristlesm ene een en ore aee ae ae flavifrons (Walker) Mesonotal stripes, if present, situated between the series of acrostical bristles orintthe planesofthesmacrochaetae...- ..--.2-05-0c0asssse soe ss oe 16 Processes widely separated at base by the basal plate, tufted with setulae at base and at apex as in vanduzeez (fig. 63), cerci broadly yellowish........ & 5 0 Slot op a oto HRS Reals CHAT Nero a SE ee Macc ne haemorrhoa (Zetterstedt) Processes nearly contiguous basad, apical region not tufted with setulae, Cencignotibroad|lyyellowishtysa-acy ci. oie ee oes = orice ree As ane = tne 17 Middle pair of presutural acrostical bristles closer to one another than to their respective series of dorsocentral bristles, with no setulae between the series, calyptrae with brownish hairs and margin, processes sparsely SEU OSE a eee yn teehee ee ese AI, ces nee nematese fumipennis Huckett Series of presutural acrostical bristles not closer to one another than to their respective series of dorsocentral bristles, with setulae between the series, calyptrae with pale hairs and margin, processes with a dense series of marginal setulae proximad......................06. longicornis Huckett MEM. AMER. ENT. SOC., 10 24 18. 19. 20. 21. 23% 24. 25, NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) All femora largely and uniformly infuscated, sometimes mid and hind pairs slightly ‘paler than:fore* pair: 2¢ 052) Hage S eet eee eee eos 19 All femora largely yellowish, or first pair more or less infuscated, second and third pairs largely yellowish or at most tinged superficially............ 48 Prealar bristle as long as or longer than anterior notopleural bristle... ... 20 Prealar bristle shorter than anterior notopleural bristle, or absent........ 26 Posterior notopleural bristle with a few setulae at base, mesopleura with a few hairs on declivity dorsad of mesothoracic spiracle, hind tibia with apical anterior bristle strongly developed, vibrissal area with numerous (10-16) coarse: black setulae. 35 .:.ietse yee ra op hao eco es ee 21 Posterior notopleural bristle with no setulae at base, mesopleura devoid of hairs on declivity dorsad of mesothoracic spiracle, hind tibia with apical anterior bristle weakly developed or absent, vibrissal area with a few slender setulae wo Sac. We auc aigrta ee iets re ree om ts rs a ene 24 Second antennal segment extensively reddish yellow........ major (Malloch) Second antennal segment extensively blackish......................... 22 . Mid femur with no bristle on distal half of anteroventral surface, mid and hind femora broadly reddish suffused throughout distal third, eyes separated by a distance not greater than maximum width of frontal vitta cephalad EM de rt, Ama ce es Pee eA Stns oy SO ts GSC a Eas Oe oS assimilis Huckett Mid femur with a bristle on distal half of anteroventral surface, mid and hind femora narrowly reddish distad, eyes separated by a distance slightly greater than maximum width of frontal vitta cephalad............... 23 Parafrontals with a few setulae opposite ocellar callosity, mid and hind femora with short posteroventral bristles, which are not longer than maxi- mum breadth of femur, abdominal sterna 3 and 4 with sparsely arranged bristles, hypopygium not notably large, abdomen tapering caudad........ Bee er ee ees Care etic .o hn Hod cha ore ee wae aro obo OS thrixia Huckett Parafrontals with no setulae opposite ocellar callosity, mid and hind femora with many posteroventral bristles longer than maximum breadth of femur, abdominal sterna 3 and 4 with bristles densely arranged on caudal half, cephalic half being bare, hypopygium large, abdomen well maintained CAUdAG ais sh Santina ee ee ee eee incompleta (Stein) Mid femur with a bristle on distal half of anteroventral surface, fore tibia with apical posterodorsal bristle robust, eyes separated by a distance equal to width of third antennal segment..................... apicalis (Stein) Mid femur with no bristle on distal half of anteroventral surface, fore tibia with apical posterodorsal bristle setulose or weakly developed, eyes sepa- rated by a distance less than breadth of third antennal segment... ....25 Abdomen conical, hypopygium black and shining, processes like those of lipsia (fig. 78), m-—cu cross vein oblique and sinuate, presutural acrostical 26. 27. 28. 29. 30. Bite H. C. HUCKETT 25 bristles in two well separated series, with setulae between the series. ... 5 fb 23-3! GR sttcs omar os Ro Oona) OEE nOT eo cree ec ROR ecme tone peer enter tae fuscicauda Huckett Abdomen depressed, hypopygium pruinescent and concolorous with terga, processes like those of vanduzeer (fig. 63), mci cross vein erect, presutural acrostical bristles in two closely adjacent series, with no setulae between SERIES ECs ree sete bl den eu a aa haemorrhoa (Zetterstedt) Costal thorn robust and prominent, cross veins usually clouded, basal node of veins R.o+3 and R.s+5 usually with a few hairs on under surface.... ..27 Costal thorn short and inconspicuous, cross veins usually clear, base of veins Ro+3 and R.g+5 bare on under surface... ........0 0.0. e eee eee eee 28 Third antennal segment yellowish proximad, apical posterodorsal bristle of fore and hind tibiae robust, mid femur with a strong bristle on distal half of anteroventral surface, costal setulae strongly developed, as long as postocularsetulaers . seeeiciowsiwiask Sos oe A ose Mae spinigerella Malloch Third antennal segment blackish, apical posterodorsal bristle of fore and hind tibiae invariably setulose or weakly developed, mid femur with no bristle on distal half of anteroventral surface, costal setulae in no prominently SUOMPwe SEH CS Paar tem i apeta ste ccna eho. aoe eos acrehon eee oie eos afinis Stein Parafacials in profile narrow, at base of antennae not broader than one fifth diameter of eye measured at a point immediately caudad, frontal vitta restricted, shortest distance between vibrissa and eye margin much shorter . than length of third antennal segment, eye margin extending caudoventrad to almost coincide with postgenal margin, thereby restricting the genal sclerite caudad, occipital region not swollen, e. g., geniculata (fig. 94). . .29 Parafacials in profile and from above prominent and extensive, at base of antennae as wide as one fourth diameter of eye measured at a point im- mediately caudad, frontal vitta extensive, shortest distance between vibrissa and eye margin nearly equal to length of third antennal segment, cheeks well maintained caudad, extending distinctly caudad of the lower margin of eye, e. g., hyoscyamt (fig. 93)... 0.0... c cece ee eee eee ee 36 Apical setulae of scutellum hairlike, abdominal terga 3 to 5 with trace of fuscous markings along cephalic border in addition to dorsocentral stripe, sternum 5 with processes like those of dissecta..................4.0-. 30 Apical setulae of scutellum coarser developed than hairs on ventral surface, or absent, abdominal terga 3 to 5 with no trace of fuscous markings along cephalic border, processes not as in dissecta, with a fringe of fine slender setulae along: inner margine sec soma: sens eo ores aetmeelae nice es 31 Ralpirentirelyablackish ese enee errno oeecneteence acuttipenmis Malloch Palpilargely: yellow: Ae 1))tele tees che decade be Sea ere anabnormis Huckett Hind femur with a series of ten or more black spinulose setulae on postero- ventral surface, copulatory appendages and hypopygium like those of TALVGR SAGO) eae ete cere see ey AN tn St Ale rufipes (Zetterstedt) MEM. AMER. ENT. SOC., IO 26 Bo. 33- 34. 35. 36. 38. NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) Hind femur not armed with a series of spinulose setulae on posteroventral surface, copulatory appendages and hypopygium not like those of gilva. .32 Abdomen slender, terga clothed throughout with slender erect bristles, marginal series inconspicuous, costa bounding cells Sc. and R. becoming notably much broader than the attenuated section bounding cell 2 C..... she ha, Sieg a RENIN eet ke es lunatifrons (Zetterstedt) Abdomen of normal robust proportions, terga with a conspicuous marginal series of stout bristles, costal vein of normal and uniform breadth along cells: Seq, sad bay swe cea RRS Ge ee Sn ere ee Be Abdomen subfuscous, reddish with blackish incisures along caudal margin of terga, scutellum obscurely reddish testaceous apicad, wings brownish tinged! throughouts.4 4.6.2 eee ee eee corrupta Huckett Abdomen and mesonotum concolorous, entirely grayish black, scutellum entirely blackish, wings at most faintly smoky....................... 34 Third antennal segment about three times as long as second segment, apex reaching a level with oral margin, series of presutural acrostical bristles separated by a distance fully equal to that between the respective series of acrostical and dorsocentral bristles, processes sharply tapering apicad.... cul Bilge A a ne en a SN, cee longicornis Huckett Third antennal segment shorter than three times length of second segment, apex not reaching a level with oral margin, series of presutural acrostical bristles closer together than distance separating the respective series of acrostical and dcrsocentral bristles, processes not sharply tapering on S61) #71 WOE] Gera at Me GG Gaba ais autre wlan niga acia'a ecnotol 35 Mesonotum viewed from behind with four postsutural stripes, situated re- spectively between the series of acrostical and dorsocentral bristles, and between the dorsocentral and intraalar bristles, processes short, widely spaced apart at base, apical region extended slightly at inner margin.... ee tee ca Peedi ene ce oe bicia. A tadbavo for o.oo aces flavifrons (Walker) Mesonotum with no such postsutural stripes, processes longish, twice as long as average width, approximated basad, apical region not produced at inner META ys Ys ok PD ak eee ear aninotata Huckett Prealar bristle absent, ym cross vein slightly infuscated, abdominal terga 4 and 5 with transverse series of discal bristles, proboscis slender cognata Stein Prealar bristle present except in minula, r—m cross vein clear, abdominal terga 4 and 5 with no transverse series of discal bristles.............. 37 . Third antennal segment yellowish at base.................---.ee000ee 38 Third antennal segment entirely blackish....................--..+-5-- 39 Third antennal segment with transverse pale area at base on inner side, presenting the appearance of an overlapping scale, processes with a dense series of setulae along inner border at base...............- ruficeps Stein Third antennal segment with no marked indications of pale scalelike pattern at base on inner side, processes with no dense series of setulae along inner | L030 (2 ee Rea tnee nr Meee EIP EYED as eeSrenee ep Acute Bs i Luo ath Oner che ied wii rufescens Stein 39. 40. 4I. 42. 43. 44. 45. 46. H. C, HUCKETT 27 Palpisyellowishvorineddishs..). 050.4 225.4. bao ds setae cea eee i aahew es 40 Palpi fuscous, at least at apex. .............0..00005 cee Bate aed oer PERC 41 Tarsi yellow, second antennal segment entirely yellow, mesonotum with no markings, processes with no tuft of setulae at apex or at base, prealar priStlevalySeritepew a ccetey i ic eres eyes dou dio sucyieuceusect eee Reem cReNewoNer minuta Malloch Tarsi blackish, second antennal segment tinged with fuscous, mesonotum with pronounced vittae along planes of dorsocentral bristles, processes with tuftlike series of setulae at apex and at base, prealar bristle present...... 5.6/6 0: Oo BiG. O10 GG OF. 0. POL ono CLI act ot tee IPE Pn ee nee ce eters vanduzeet Malloch Second antennal segment entirely yellowish, vibrissae separated by a distance less than height of cheek, processes grayish drab with a thickened polished strip along distal half of inner margin................ convergens Huckett Second antennal segment infuscated or brownish, vibrissae separated by a distance greater than height of cheek..................0 0000 cece eeeee 42 Fore, mid and hind femora largely yellowish, with trace of fuscous permeating the integument, coxae reddish, processes devoid of bristles, with only a few sparse setulae, cerci and styli black and polished............ striata Stein Fore, mid and hind femora decidedly and for the greater part infuscated or blackish, processes with numerous bristles and setae, coxae blackish... .43 Mesonotum viewed from behind with four postsutural vittae, situated re- spectively between planes of acrostical and dorsocentral, dorsocentral and intraalar series of bristles, scutellum with a dark fascia along base...... 0.6.0.0 6 Godiva che c {Ol Rae ATEee SC aee OR ree a a A et ee enters flavifrons (Walker) Mesonotum with no such interserial stripes, if present they lie dorsocentral or along the planes of dorsocentral and intraalar series of bristles, scutellum with no dark fascia across base........... 000. e ee cece ete ees 44 Hind tibia with apical posterodorsal bristle strongly developed, mid tibia with a weak mid anterior bristle near anterodorsal bristle, parafrontals bristled for entire length, abdomen conical, processes as in bicolor (fig. 90) 0:65 100, GS Bea OSL ORCC Eee FTE a ERE MERC eM RAR re eM Pace jacobt Malloch Hind tibia with apical posterodorsal bristle slender and weakly developed, mid tibia with no weak anterior bristle, parafrontals not bristled for entire length, abdomen depressed, processes not like those of bicolor.......... 45 Processes broadly produced throughout, widely truncated at apex, distal half and inner border yellowish, polished and bare, mid and hind femora broadly yellowish on apical region, extending to include preapical bristles, hind tibia with four or more anterodorsal bristles............. indicta Huckett Processes not truncated at apex but tapering distad, mid and hind femora narrowly yellowish at apex, hind tibia with not more than three antero- dorsallbnistlesivs peisisrs foreach ae aise ane isu eye ate eee sins Sick nce 46 Parafacials at base of antennae fully as wide as height of cheek, eyes separated by a distance fully as great as that between posterior ocelli inclusive, pre- basal sclerite with numerous bristles...................... glabra (Stein) MEM. AMER. ENT. SOC., 10 28 47. 48. 49. 50. SI. 52. 54. NORTH AMERICAN GENUS PEGOMYIA (DIPTERA: MUSCIDAE) Parafacials not wider than height of cheek, eyes separated by a distance less than that between posterior ocelli inclusive, prebasal sclerite bare...... 47 Processes with a tuftlike series of setulae at apex as in vanduzeer (fig. 63), proboscis polished and slender, postocular setulae fine and long, mesonotum inconspicuously striped....................-5. haemorrhoa (Zetterstedt) Processes devoid of setulae at apex, proboscis pollinose and short, postocular setulae short and coarse, mesonotum with a conspicuous brownish dorso- Central! vata dh: sheesh We ere ee ne atlanis Huckett Prealar bristle long and robust, longer than or as long as anterior noto- pleural bristle, if of about equal length abdomen black............... 49 Prealar bristle absent or shorter than length of anterior notopleural bristle, if of about equal length abdomen reddish yellow..................... 71 Arista with hairs longer than setulae on second antennal segment, sparsely distributed casa decdncine sco a eee 50 Arista with hairs scarcely longer than setulae on second antennal segment, pubescenitittrent vidal oh ae Rinse cater. SERS ee ee 51 Processes with a matt of short stiff setulae on proximal half of inner border, arista plumose, longest hairs as long as width of third antennal segment, palpi spatulate, fore femora at base with a conspicuous well defined brownish area on anterior (inner) surface, hind tibia with no posterior bristle on proximal half syd R ie c the ee ean eo ea ee rte juvenilis (Stein) Processes with no mattlike series of stiff setulae on proximal half of inner border, arista subplumose, longest hairs not as long as width of third antennal segment, palpi slender, fore femora at base of anterior (inner) surface with no such well defined marking, hind tibia with a posterior bristlejonsproximaluhalieeeee sree eer n reer rerere connexa Stein Abdomen largely reddish yellow, or if mcre or less infuscated or blackened, is not concolorous with thorax and the humeral callosities are also reddish 52 Abdomen and mesonotum, including humeral callosities, grayish black, CONCO] GROWS hs ise sP eee Se es ee 56 Thorax largely reddish to yellowish, hind tibia with a short bristle on proximal half of posterior surface, mid femur with a bristle on distal half of antero- ventral surface.wia oS COSSUOBUED so ccccccccscvcvvcve 119 GRESCaNS ant eek ene ee 86 CULICOMIS wee ars eS 54 diaphanus, Alloeostylus.......... 8 dissecta was asada nce ae eee 55 duplicatans 7 on cee soe 94 | Sed KSeraem tees aaa ero t one Hea sone d che 6 (See also tunica) EMIMESIAzs ac: seen eae Oe 79 ephippium, Anthomyia......... 112 Eremomyia (see apicalis) AMAIA 6 oe 6 ee eae ee 115 Hawa CANS 2c; cayceare eat oeah ree ee 84 favatronS a... eae eee Ne ee 71 Mavipalpistycs.tm.a eee eee 109 SLOUUDOS som cn or ee 98 flavoscutellata, Anthomyia....... 9 Pa gidaie.s pace Aa ee 90 SUI OUG eae ee ee ee ee ee 71 Ul SENS ices cg Pe EO 5 TUVTMOSNWVIS oo gosacnaaccas eaten: 58 fuscicaudayc necator te ore oe 64 PLUS CUCU US Monica: oe pe eee 94 fuscofasciata, 4),.j4 4050. 8 acaeeos 102 genticulataiy, «ce seen ee 112 GilvViae ators Seema ee era 104 Slab rane seo Lee ee 61 SOPH ETA asa sare Merce eeew Ree a 115 haemorrhoadeeeene errr 58 Hydrophoria (see orientalis) 130 Hylemyia (see artica, brassicae) hy OSCY AMAL. 25.0: ineieyehe codes aoe eee 126 PCLERICAL. en jancnace eee ee 58 TMICISIVASS = acanccds cae eee 106 INCOMPletal. +s. onan eee Re 62 INGICtA... ci: dee ee ee ee ee 77 INteKSeCta. ac oe jase 52, 50 JaACODL.oe a4 eee ee eee ae 76 juventlisy S..7..0a as oe eee Wal labrad orensisne eter 80 latipalpis.2c aoe eae 56 LGHIGESIS. 22: ee ee 102 lativiligia eo eee ene 90 lenisayAntthomiyialeee ine 8 lipsia.. 2k cor ome eo e 87 littoraliss2.:: cad ary eo eee 53 lividiventriss ere coe eee 80 longiconnlisaenenee eee ee 70 longimana=... eee ye 95 lunatifronss A sersene omc oie 69 Luteolas 33 none ee 105 IMA OL sc yei steerer eer 61 mialllO@ht) cae eae See Oe Oe IOI Mavginata <2) eos severed are ere 120 MANU a i ee 69 TTI Cal -< cera eae eee oe 74 neglecta, Phorbia............... 52 MUSTACEDS kin ee oe 100 MIGUItALSIS:: o/s ciara eee ete 83 nigroscutellata................. 57 mutet hula ee ak een ee 10, 12 INUGGIIG geste Coen 5 INGDUDEDIO 8s ee) cod ee 5 ODN OTIS his Ante eee 108 ODSCURIOI Ry sade hoecenen ee 98, 107 Qnvenialisi AG neces yee eee 51 DOL Rrary, erica aia Romine Ae 105 Palpatamerrnciorte cet c oes 51, 123 PalpOsaly pence eae 51 DAR Gltaye sees wearageek eee en eee 80 pedestrisy ty se eee 119 Pegoplata'y... 430 cen eee 6 pertusats pS ee oe eee 116 PROVES Lh oe on oe 5 Phorbiainc. tats ote eee 6 (See also neglecta) pildsar.is Aso: Gets ee 98 POllin@sa® .A 7. ashe eee ees eee 110 DOU COND a Nae ee ee 121 praepotens, Anthomyia........... 8 pseudodissecta................. 54 Quadnispinosay.. 4540-4 eee gI quintilis, Chortophila........... 52 FECtUTONS 52 soins eo aie) Wee 108 RUDIVOLaz aca seca aes ee 60 TUleScenSiea cc sn acre ee 74 TUMICE DS). fs, oo ey ee WB Huda, Se aes ee ee 100 is1D0 Oh OS urna” cane Meta ease Ativan) 2 119 rumicifoliae (Huckett).......... 81 sepia, Chortophila.............. 10 sepiellay Maciviicne sin Saran ate 53 Setanlar ae ant natant eee 121 silacea, Anthomyia.............. 8 SILIEHS ha phere aie ete 123 SCHLORMIS A> <2 eam tee ee 93 SOLE an sauce ee eee II, I2 SCtOSal Ree. yt erent eee 10 SIOSSONGCrm is ene ee ee ee 57 socialis, Spilogaster............. 10 Solitaria. “hen csc ees ee Oe ee 96 Spilogaster (see socialis) spimigrellai ne oisc ue es eee 65 spinosissima.................. 114 SCULCINA CL mara eee 100 Stlembainc5 Scheer oe ee 12 SUHlatA ce; crsamus ts cet cance eee 76 SUD ENUSE Dae ee ROL eee eee gI substituta, Anthomyia........... 87 substituta, Coenosia............. 9 Sulbstiiatte] aaa ene 84 INDEX nan CATSatay wees aca ewioite ieeeares 85 UNICOLOG Meters eee ee te eee owe 100 PEMERA vee Miro aeioci nad Deca 98, 107 univittata........ LL Ee III Chrixiawetesee eerie eee cee os 62 {OUNCHIG|UNT a onan conagesasaoa: 122 VIROL WASE ss oop cosas ceraaccuecy 75 transversalis@a ssn aes once 58 Walle aitalrn cm t at ee rane 120 RUNMOEN a aacvocanccoceosuge 10, 12 WACAT I 27 aren cg ce een 122 GhiSC Laer treee tes cae eam ee 118 CUR oy no 06000 bed 08RD osl08 66, 127 thivittatal ne cen ccis ne cee Awe 10 virginea, Anthomyia.......... 7, 12 tundrica, Egle.................. 55 VALE C2 eee 103 tunicata, Anthomyia............. 8 : : WaAmthemias. son au was eee IOI, 102 unguiculataseene same aaa oer 97 LEDUC IIA aeRO 5 MEM. AMER. ENT. SOC., 10 JIL AS) PVA Dorsal or caudal aspect of male copulatory appendages Fig. 1.—Pegomyia palposa (Stein) Fig. 2.—Pegomyia intersecta (Meigen) Fig. 3.—Pegomyia unguiculata Malloch Fig. 4.—Pegomyia littoralis Malloch Fig. 5.—Pegomyia nigroscutellata (Stein) Fig. 6.—Pegomyia pseudodissecta (Ringdahl) Fig. 7.—Pegomyia glabra (Stein) Fig. 8.—Pegomyia incompleta (Stein) Fig. 9.—Pegomyia acutipennis Malloch Fig. 10.—Pegomyia major (Malloch) PuaTE I. Mem. AMER. ENT. Soc., No. Io. HUCKETT—PEGOMYIA PLATE II Dorsal or caudal aspect of male copulatory appendages. Fig. 11.—Pegomyia quadrispinosa (Malloch) Fig. 12—Pegomyia substriatella (Malloch) Fig. 13.—Pegomyia apicalis (Stein) Fig. 14.—Pegomyia duplicata (Malloch) Fig. 15.—Pegomyia frigida (Zetterstedt) Fig. 16.—Pegomyia tarsata (van der Wulp) Fig. 17.—Pegomyia longimana (Pokorny) Fig. 18.—Pegomyia spinigerella Malloch Fig. 19.—Pegomyia gopheri Johnson PLATE II. Mem. Amemr. ENT. Soc., No. 10. PEGOMYIA sO 0 PLATE III Dorsal or caudal aspect of male copulatory appendages. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. 20.—Pegomyia flavifrons (Walker) 21.—Pegomyia cognata Stein 22.—Pegomyia ruficeps Stein 23.—Pegomyia setaria (Meigen) 24.—Pegomyia triseta Malloch 25.—Pegomyia flavipalpis (Zetterstedt) 26.—Pegomyia univittata (von Roser) 27.—Pegomyia unicolor Stein 28.—Pegomyia incisiva Stein 29.—Pegomyia pilosa Stein PLATE III. Mem. AMER. ENT. Soc., No. 10. HUCKETT—PEGOMYIA PLATE IV Lateral aspect of male copulatory appendages. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. 30.—Pegomyia palposa (Stein) 31.—Pegomyia intersecta (Meigen) 32.—Pegomyia unguiculata Malloch 33.—Pegomyia littoralis Malloch 34.—Pegomyia pseudodissecta (Ringdahl) 35.—Pegomyia glabra (Stein) 36.—Pegomyia nigroscutellata (Stein) 37.—Pegomyia incompleta (Stein) 38.—Pegomyia acutipennis Malloch 39.—Pegomyia major (Malloch) Mem. AMER. ENT. Soc., No. Io. PLATE IV. HUCKETT—PEGOMYIA PLATE V Lateral aspect of male copulatory appendages. Fig. 40.—Pegomyia quadrispinosa (Malloch) Fig. 41.—Pegomyia apicalis (Stein) Fig. 42.—Pegomyia duplicata (Malloch) Fig. 43.—Pegomyia tarsata (van der Wulp) Fig. 44.—Pegomyia longimana (Pokorny) Fig. 45.—Pegomyia substriatella (Malloch) Mem. AMER. Ent. Soc., No. 10. PLATE V. HUCKETT—PEGOMYIA PLATE VI Lateral aspect of male copulatory appendages. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. 46.—Pegomyia frigida (Zetterstedt) 47.—Pegomyia spinigerella Malloch 48.—Pegomyia gopher Johnson 49.—Pegomyia pilosa Stein 50.—Pegomyia incisiva Stein 51.—Pegomyia unicolor Stein 52.—Pegomyia flavipalpis (Zetterstedt) 53.—Pegomyia univittata (von Roser) PLATE VI. Mem. AMER. Ent. Soc., No. Io. HUCKETT—PEGOMYIA PLATE VII Ventral aspect of fifth abdominal sternum in male. Fig. 54.—Pegomyia palposa (Stein) Fig. 55.—Pegomyia intersecta (Meigen) Fig. 56.—Pegomyia ungurculata Malloch Fig. 57.—Pegomyiu littoralis Malloch Fig. 58.—Pegomyia glabra (Stein) Fig. 59.—Pegomyia acutipennis Malloch Fig. 60.—Pegomyia nigroscutellata (Stein) Fig. 61.—Pegomyia dissecta (Meigen) Fig. 62.—Pegomyia pseudodissecta (Ringdahl) Fig. 63.—Pegomyia vanduzeei Malloch Fig. 64.—Pegomyia incisiva Stein Fig. 65.—Pegomyia pilosa Stein Fig. 66.—Pegomyia gilva (Zetterstedt) Fig. 67.—Pegomyia unicolor Stein Fig. 68.—Pegomyia flavipalpis (Zetterstedt) Fig. 69.—Pegomyia umivittata (von Roser) Mem. Ame_er. Ent. Soc., No. Io. PLATE VII. HUCKETT—PEGOMYIA PLATE VIII Ventral aspect of fifth abdominal sternum in male. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. 70.—Pegomyia incompleta (Stein) 71.—Pegomyia major (Malloch) 72.—Pegomyia quadrispinosa (Malloch) 73.—Pegomyia subsiriatella (Malloch) 74.—Pegomyia apicalis (Stein) 75.—Pegomyia duplicata (Malloch) 76.—Pegomyia tarsata (van der Wulp) 77.—Pegomyia longimana (Pokorny) 78.— Pegomyia lipsia (Walker) 79.—Pegomyia frigida (Zetterstedt) 80.—Pegomyia gopheri Johnson 81.—Pegomyia spinigerella Malloch Mem. Amer. Ent. Soc., No. 10. Piate VIII. HUCKETT—PEGOMYIA PLATE IX Lateral aspect of male copulatory appendages. Fig. 82.—Pegomyia cognata Stein Fig. 83.—Pegomyia flavifrons (Walker) Fig. 84.—Pegomyia ruficeps Stein Fig. 85.—Pegomyia setaria (Meigen) Fig. 86.—Pegomyia triseta Malloch Ventral aspect of fifth abdominal sternum in male. Fig. 87.—Pegomyia flavifrons (Walker) Fig. 88.—Pegomyia ruficeps Stein Fig. 89.—Pegomyia cognata Stein Fig. 90.—Pegomyia bicolor (Wiedemann) Fig. 91.—Pegomyia triseta Malloch Fig. 92.—Pegomyia setaria (Meigen) Lateral aspect of male head. Fig. 93.—Pegomyia hyoscyami (Panzer) Fig. 94.—Pegomyia geniculata (Bouché) Mem. Ame_r. ENT. Soc., No. Io. PLATE [X. HUCKETT—PEGOMYIA te MEMOIRS OF THE AMERICAN ENTOMOLOGICAL SOCIETY An irregular serial, containing monographic papers $y students authority in their respective subjects. No. 1. The Cresson Types of HaeeOnteRe By Ezra T. Cresson, 1916, I41 pp. No. 2. The Blattidae of North America, North of the Mexica Boundary. (Orthoptera.) By Morgan Hebard, 1917. 284 PP 10 pls. No. 3. A Venational Study of the Suborder Zygoptera, with Ke for the Identification of Genera. (Odonata.) By Philip A. Mur 1919. 78 pp., 20 pls. aay z No. 4. The Blattidae of Panama. (Orthoptera.) By Morgan mie Hebard. 1920. 148 pp., 6 pls. ee. ¥ Sciences of Philadelphia other than those of Ezra T. Cresson. Ezra T, Cresson, 1928. 90 pp. No. 6. Revision of the Rhipiphoridae of North America. (Cole pacers 3: optera.) By Ezekiel Rivnay. 1929. 68 pp., 4 pls. a No. 7.. A Revision of the Dipterous Family Rhagionidae (Lepti- - dae) in the United States and Canada. By Mortimer D. Leonard. £930:)' 182) pp., 3 pls. ei No. 8.. The Eumastacinae of Southern Mexico and Central Amer- rt ica. (Orthoptera.) By James A. G, Rehn and John W. H. Rehn, © 1934. 84 pp., 6 pls. No. 9. The Generic Names of the Sphecoid Wasps and their Type — Species. (Hymenoptera.) By V.S.L. Pate. 1937. 103 pp.