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MEMOIRS
OF THE
QUEENSLAND MUSEUM
VOES- Vit:
WITH NINETEEN PLATES AND SEVENTY-TWO TEXT-FIGURES.
EDITED BY THE DIRECTOR,
HEBER A. LONGMAN, F.L.S.
DATE OF ISSUE OF FINAL PART: DECEMBER -19, 1922:
BRISBANE:
ANTHONY JAMES CUMMING, GOVERNMENT PRINTER.
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CONT ESAS:
PARI:
(Issued 30th June, 1920.)
Edible Fishes of Queensland, Plates I.-IIT.
Notes on the Biology of some Queensland Flies
The Occurrence in Brisbane River of the New Zealand Amphipod,
Paracorophium excavatum
New or little-known Crane Flies in the Queensland Museum
Little Penguin in Queensland .
PART 2:
J. Douglas Ogilby
T. Harvey Johnston, M.A., D.Sc.,
and M. J. Bancroft, B.Sc.
Chas. Chilton, M.A., D.Se., &e.
Charles P. Alexander, Ph.D.
Heber A. Longman, F.L.S.
(Issued 11th February, 1921.) '°
A New Genus of Fossil Marsupials, Plates IV.—VII.
PARI 43:
Heber A. Longman, F.L.S.
(Issued 4th November, 1921.)
Australian Bees in the Queensland Museum
Catalogue of Australian Bees
‘Notes and Illustrations of Queensland Fishes, No. 2, Plates
VIIL-XI..
A New Nyctimene from South Queensland, Plate XII.
On Coleoptera mostly from Queensland, Plate XIII
PARI 4.
T. D. A. Cockerell
Henry Hacker, F.E.S.
Allan R. McCulloch, Zoologist,
Australian Museum
Heber A. Longman, F'.L.S. :
Arthur M. Lea, F.E.S.
(Issued 19th December, 1922 }
Notes and Dlustrations of Queensland Fishes, Plate XIV.
An Ichthyosaurian Skull from Queensland, Plates XV.-XVI.
Australian Bees in the Queensland Museum
On the Emergence of two tube-dwelling Homopterous Insects,
Plates XVII. and XVIII.
- Some Australian Wasps of the Genera Zoyphiwm and Arpactus
John Gould’s Notes for John Gilbert
Notes on the Fat-tailed Marsupial Mouse (Sminthopsis crassi-
caudata)
South Queensland Marsupials
Three New Queensland Fishes, Plate XIX.
A. R. MeCulloch
Heber A. Longman, F.L.S.
T. D. A. Cockerell
Henry Hacker, F.E.S.
Henry Hacker and T. D. A.
Cockerell
W. B. Alexander, M.A.
Heber A. Longman, F.L.S.
J. Douglas Ogilby
PAGE.
1-30
31-43
44-5]
52-63
64
65-80
81-98
99-163
164-178
179-151
182-240
291-294
295-296
297-300
301-304 .
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MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Parr I.
EDIBLE FISHES OF QUEENSLAND.
By J. DoucLtas OciLtay (ICHTHYOLOGIST).
PaRT XV.—SERRANIDZ (No. 1).
(Plates I to III.)
Percalates Ramsay & Ogilby, Proc. Linn. Soc. N. S. Wales, xii, 1887, p. 182; Ogilby, Edib.
Fish N. S. Wales, 1893, p. 2; Boulenger, Catal. Percif. Fish., 1895, p. 132. Type—
Lates colonorum Ginther 1863.
Body subovate and compressed. Scales moderate, adherent, concentrically
striated, more or less strongly ciliated, those of the throat and breast small.
Lateral line complete, continued on the base of the caudal fin, the tube simple
and straight, not extending to the border of the seale. Head large and partly
naked, the upper profile variable. Mouth terminal and protractile, with wide
oblique cleft, the lower jaw more or less projecting, the maxillary large and
dilated, exposed, with well developed supramaxillary. Teeth in villiform bands,
on the jaws, vomer, and palatines; tongue toothless. Nostrils of moderate size,
approximate, opening in front of the middle of the eye, and much nearer to it
than to the edge of the preorbital, the anterior valvular. Preorbital, suborbital,
and vertical border of preopercle serrated ; angle and lower border of preopercle
armed with strong teeth, which on the latter may be antrorse; opercle with two
divergent spines, the lower much the longer; postclavicle and posttemporal
usually serrated. Two dorsal fins, connected at the base, the first with viii, rarely
ix, strong pungent spines, the interspinous membrane deeply cleft; second dorsal
shorter than the first, with i 8 to 11 rays, the base concealed by a low sealy sheath.
Caudal emarginate, with 17 principal rays. Anal short, with ii 7 to 9 rays,
extending well beyond and similar to the second dorsal. Pectoral obtusely
pointed, with 12 to 16 rays, the upper middle ones the longest. Ventral longer
than the pectoral with i 5 rays, inserted a little behind the pectoral-base. Gill-
openings wide; gill-membranes separate, free from the isthmus; six branchios-
tegals; pseudobranchie present; gills four, a slit behind the fourth; gill-rakers
stout and well developed, in moderate number; pharyngeal bones large, densely
covered with small teeth. Air-bladder present, large. Stomach cecal, intestinal
canal convoluted, pyloric appendages in moderate number. Posterior processes
of the premaxillaries not extending to the frontals; parietal and supraoccipital
bones not extending to between the postfrontal processes; supraoccipital crest
strong, not produced on the frontals. Vertebre 11-+ 14 = 25.
Monotypic. The single species inhabiting the fresh waters and estuaries of
Southern and Eastern Australia, from the Gulf of Saint Vincent, S.A., to the
Pine River, 8.Q.
2 MEMOIRS OF THE QUEENSLAND MUSEUM.
Notes:—Boulenger describes and McCoy figures the scales of this fish as
being ‘‘cycloid.’’ Of the numbers which I have examined I have never seen one
to which this description could be correctly applied; the strength of the ciliation
is, however, extremely variable, and some specimens only show it weakly along the
middle of the side, where about half a dozen small teeth are alone present on the
middle of the outer edge of the scale, leaving a broad dorsal and abdominal
eyecloid band. Normally, however, the scales of the estuarine forms show distinet
ciliation, except those of the nape and a narrow, gradually contracting band
below the base of the spinous dorsal, where they are constantly cycloid. And
this variation in the scale-character of those examples which more or less
permanently reside in the brackish water of an estuary, leads naturally on to the
much coarser ciliation of the scales of those individuals which have taken on a
more or less purely fluviatile existence. This character, therefore, is manifestly
of no value as a specific factor. With regard to the emargination of the upper
profile of the head, on which Stead lays so much stress, I find that this is wholly
caused by the usually more marked convexity of the snout in the estuarine form,
which necessarily induces a hollow between the snout and the occipitonuchal
convexity. As, however, I have examined several of these fishes in which,
through the straightening of the snout-contour, the emargination was practically
eliminated, it will be understood that I can not see my way to accepting this
character as of specific value, the difference being in my opinion more apparent
than real.
PERCALATES COLONORUM (Giinther).
Lates colonorum Ginther, Ann. & Mag. Nat. Hist. (3) xi, 1863, p. 114; Castelnau, Proce.
Zool. & Accel. Soc. Vie., i, 1872, p. 43; McCoy, Prodr. Zool. Vic., dec. ii, 1878, pl. xiv;
Macleay, Proc. Linn. Soc. N. 8S. Wales, v, 1881, p. 304; Tenison-Woods, Fish & Fisher.
N. S. Wales, 1882, p. 31, pl. i; Johnston, Proc. Roy. Soc. Tas., 1882, p. 59; O’Connor,
Proce. Roy. Soc. Queensl., xii, 1897, p. 110; Zietz, Trans. Roy. Soe. S. Austr., xxvi, 1902,
p. 265.
Dules novemaculeatus Steindachner, Sitz. Akad. Wien, lili, 1886, i, p. 428, pl. i, fig. 2; id.,
ibid., lx, 1869, i, p. 674; Klunzinger, Arch. f. Nat., xxxviii, 1872, i, p. 20.
Dules reinhardtii Steindachner, ibid., lvi, 1867, 1, p. 320.
Lates similis Castelnau, ibid., p. 44; Macleay, ibid., p. 305.
Lates antarcticus Castelnau, ibid.; Macleay, ibid.
Lates victorie Castelnau, ibid., p. 45; Macleay, ibid.
Lates curtus Castelnau, Res. Fish. Austr., 1875, p. 5; Macleay, ibid., p. 306.
Lates ramsayi Macleay, ibid.
Percalates colonorum Ogilby, Edib. Fish. N. 8S. Wales, 1898, p. 2, pl. i; Boulenger, Catal.
Percif. Fish., 1895, p. 132; Stead, Fish. Austr., 1906, p. 96; id., Edib. Fish. N. S. Wales,
1908, p. 53, pl. xxii, low. fig.; id., Proe. Linn. Soc. N. S. Wales, xxxv, 1910, p. 659;
Roughley, Fish. Austr., 1916, p. 60.
Percalates flwwiatilis Stead, Proe. Linn. Soc. N. 8. Wales, xxxi, 1906, p. 261; id., Edib. Fish.
N. 8. Wales, 1908, p. 54, pl. xxii, upp. fig.; MeCulloch, Proc. Linn. Soe. N. 8. Wales,
xxxv, 1910, p. 432; Stead, Proce. Linn. Soc. N. S. Wales, xxxv, 1910, p. 658.
+ Boulenger, I believe erroneously, quotes this form as Dules reinwardti; I follow
Ginther (Zool. Ree., iv, 1867, p. 159). McCulloch, in answer to a query, kindly writes—
““ According to my slip, copied from the original reference, the name is Dules Reinhardti.’’
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 3
BASS.
Australian Bass; Perch; Estuary Perch; Fresh-water Perch.
Type localities:—Victoria (Ll. colonorum).
Neighborhood of Sydney (D. novemaculeatus).
Port Jackson (D. reinhardtii).
Gippsland Lakes (L. similis).
Melbourne Market (L. antarcticus).
Melbourne Market (L. victorie).
Richmond River, N.S.W. (L. curtus).
Waterhole at Parramatta, N.S.W. (L. ramsayr).
Rivers of New South Wales (P. fluviatilis).
Dorsal contour of body rather more arched than that of the ventral, its width
at the shoulder 1-9 to 2-5 in its depth immediately in front of the ventral fins,
which is 2-4 to 3 in its length and equal to or a little more than the length of the
head. Caudal pedunele stout, its depth 1-37 to 1-6 in its length behind the soft
dorsal and 2-4 to 2-67 in the length of the head. Head from two fifths to four
fifths deeper than wide, the upper profile varying from linear in fluviatile
examples to emarginate in those which remain permanently in estuarine waters,
the difference being solely caused by the shape of the snout, the contour of which
is more or less rounded in the latter form, thus making with the occipito-
nasal convexity an intervening fictitious emargination; depth of head 1-38 to
1-5 in its length, which is 2-55 to 3-25 in that of the body. Snout pointed,
its length 3-75 to 4-17 in that of the head. Diameter of eye from one
fourth more to one fifth less than the length of the snout and 3-33 to 4-75 in that
of the head; preorbital narrow; interorbital region wide and moderately convex,
its width equal to one half less than the eye-diameter. Maxillary extending to
below or a little beyond the middle of the eye, its length 2-33 to 2-7 in that of the
head, the width of its truncate distal extremity 1-33 to 3 in the eye-diameter.
Preorbital and suborbital finely and evenly serrated, the edge of the former
undulous or emarginate ; hinder limb of preoperele linear and subvertical, armed
with fine serree which gradually decrease in size from below and are absent or
vestigial on the upper part of the limb; angle and lower limb with a series of
strong, more or less curved spines, which are sometimes broken up distally into
two or more points and, on the latter may be arranged in groups, each individual
spine having a more or less antrorse direction; lower opercular spine two thirds
to three fifths of the eye-diameter.
Premaxillary bands of teeth broader than those of the mandible, each
separated by a symphysial hiatus, and each diminishing to a blunt point behind;
vomerine teeth in an arcuate band, of equal width throughout, as also is the
palatine band.
Seales of body more strongly ctenoid in the fluviatile than in the estuarine
form, arranged in 48 to 55 series above the lateral line, in 7 to 9/1/16 to 19
between the fifth dorsal spine and the vent. Scales of nape, a strip along the base
of the spinous dorsal, throat, and head ecyeloid. Opercles, postorbital, and
4 MEMOIRS OF THE QUEENSLAND MUSEUM.
parietal regions, and cheeks scaly, the rest of the head naked; cheek-scales much
smaller than the others, in 8 or 9 series between the eye and the angle of the
preopercle. Lateral line following the curvature of the back.
Dorsal fins with viii or ix, i 8 to 11 rays, originating slightly behind the
pectoral-base, and terminating above the fifth or sixth anal ray ; first spine short,
4.2 to 4-75 in the longest, which is usually the fourth, though the fifth sometimes
slightly exceeds it, and is 1-67 to 2-67 in the length of the head, the last as long
as or a little longer than the second, 1-9 to 2-2 in the longest, and 1-22 to 1-4 in
the spine of the second dorsal, which is 1.4 to 1-67 in the longest ray ; first, second,
or third ray longest, one sixth more to one sixth less than the longest spine, and
subequal to the basal length of the second dorsal, which is 1-17 to 1.33 in that of
the first ; first dorsal ray divided distally, the last split to the base; outer border
of soft rays gently rounded. Caudal fin emarginate, with 17 principal rays, the
lobes equal and obtusely pointed, the middle rays 1-38 to 1-67 in the upper lobe,
which is 3-55 to 3-83 in the body-length. Anal fin with i 7 to 9 rays, originating
below the third or fourth dorsal ray ; second and third spines equal or the third
a little the longest, intermediate in length to the sixth and seventh dorsal spines,
2.44 to 3-67 in the length of the head, and 1-33 to 1-5 in the first or second ray,
which is one tenth to one fourth more than the basal length and 1-8 to 2-2 in the
length of the head; outer border linear; rays similar to those of the dorsal.
‘Pectoral obtusely pointed, with 12 to 16 rays, its length 1-55 to 2 in that of the
head; fourth or fifth ray longest, extending to below the sixth or seventh dorsal
spine. Ventral bluntly pointed, the spine strong and pungent, 1-37 to 1-67 in the
length of the first and second rays, which are subequal, 1-44 to 1-7 in the
length of the head, and do not usually reach the vent.
Gill-openings extending forward to below the middle of the eye; gill-rakers.
longer than the gill-fringes, 13 to 16 on the lower branch of the anterior arch, the
longest 1-3 to 2-67 in the eye-diameter ; pyloric appendages 10.
Upper surface varying from olive-brown to deep olive-green, the head
darkest ; sides and lower surfaces shading from grayish green to grayish white,
the latter and the cheeks sometimes washed with yellow. All the fins olive-green,
except the first dorsal, which is gray or lavender; upper half of opercle, ventrals,
and anal with a conspicuous blackish blotch in the young.
Deseribed from numerous examples, measuring from 143 to 878 mm.,
collected at various localities between and including the Snowy River, N.S.W..,
and the Pine River, Q.
My ‘‘Edible Fishes and Crustaceans of New South Wales’’ being now
nnattainable, I think it advisable to republish my reasons for rejecting the
supposititious species of Castelnau and Macleay, so as to make this article as.
complete as possible ; and at the same time to bring my account of the species up:
to date by going thoroughly into the claims of Percalates fluviatilis to validity.
In the work quoted I remark :—
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 5
‘(A few words are necessary here with regard to the forms of the
Australian Perch deseribed as new by Castelnau and Macleay, the types of all of
which are missing. In 1872 the former created three new species to which he
gave the names Lates similis, L. antarcticus, and L. victoriw. To these, between
the above date and 1881, he added a fourth species, LZ. cwrtus, from the Richmond
River. In 1876, Alleyne and Macleay described, under the name Pseudolates
cavifrons, a North Australian fish, and in the following year the latter author
redescribed the same species as Lates darwimensis; these two names may be at
once dismissed from consideration as being mere synonyms of the widely dis-
tributed L. calcarifer, as an examination of the type specimens at once reveals.
Finally in 1881 Macleay described a Lates ramsayi from a single specimen taken
in a freshwater pool near Parramatta, the type of which is also missing. We
have, therefore, in the restricted genus Percalates, no less than five spurious
species, excluding the original P. colonorwm, all described from a similar and
‘somewhat limited area, on what appear to be insufficient grounds. That none of
the later writers on Australian fishes (Johnston, Catalogue of Tasmanian Fishes,
1882, McCoy, loe. cit., 1878, and Lucas, Census of Victorian Fishes, 1889) except
the author (Catalogue of New South Wales Fishes, 1885), venture an opinion on
their specific identity or otherwise, is, it must be conceded, a most unsatisfactory
state of affairs, and merits, therefore, a more extended inquiry than is usually
necessary in such cases.”’
To these must now be added Percalates fluviatilis Stead, the claims of
which have been so strenuously put forward by its author. With this form I
shall deal on a future page. Of Steindachner’s two species, both from Port
Jackson or its immediate neighbourhood, I am unable to say anything from
personal knowledge, not being in a position to consult his descriptions or figure,
but there can be no doubt as to their identity with one or other of the forms of
P. colonorum. Of the first, Dules novemaculeatus, indeed, McCulloch writes as
follows :—‘‘It appears to me that Steindachner’s figure of Dules novemaculeatus
exactly represents the slender form; and, beyond such differences as would be
caused by shrinkage due to different methods of preservation (alcohol Stein-
dachner and formalin Stead), it does not differ from Mr. Stead’s figure
published in the Edible Fishes of New South Wales.”’
My original remarks regarding Castelnau’s four species are :—
‘“‘In L. similis the characters relied on for its separation from the type
species are absurdly inadequate; these. are the shorter snout, which is ‘sensibly
less than the diameter of the eye’; with the majority of fishes the comparative
size of the eye to the head and to the snout varies with the age of the individual,
the young fish having that organ much larger proportionately than the adult.
As I have shown above the great variation existing between the comparative
measurements of the eye and the snout in twenty five specimens of indubitable
P. colonorum, ranging from Adelaide to the Richmond River, it is manifest that
the stress laid upon this character is altogether misleading, and must be regarded
as valueless. The coarser denticulations of the preopercle are also a sign of
6 MEMOIRS OF THE QUEENSLAND MUSEUM.
immaturity, and as such unreliable. In the large series which has passed
through my hands specimens have in rare instances occurred in which the teeth
of the lower preopereular limb were directed absolutely downwards; neither this
character nor that of the omission of one of the dorsal rays can, therefore, be
taken as a valid reason for separating L. similis from Giinther’s well known
species.”’
As regards the length of the snout this is, in Mr. Stead’s selected
specimen of P. fluviatilis, measuring 145 mm., one fourth less than the eye
diameter, and even in the 275 mm. example forwarded by the New South Wales
Fisheries Department, the two are of equal length, while in the former the pre-
opercular dentition is everywhere exceptionally strong, and in the latter there are
but nine soft dorsal rays, so that in these two picked specimens all the characters
“of L. similis are present. It is worth noting that in Castelnau’s five descriptions
of Lates not once does he mention the shape of the head; it is, therefore,
presumable that he attached no weight to this character.
Of Lates antarcticus Castelnau I wrote :—
“‘Tt is unnecessary to go at any length into the question of the validity of
L. antarcticus, since, with two exceptions, a comparison of Castelnau’s description
with that given above, will show that no characters are put forward, which are
not equally common to P. colonorum. These are the increased number of
branchiostegal rays, which is stated to be seven in this species as against six,
which the examination of numerous specimens has shown to be the normal number
in P. colonorum as here and elsewhere stated, and also recorded by Prof. MeCoy.
I do not, however, attach any importance to this seeming discrepancy, since it is
possible that the number is merely copied from the generic description of Lates
given by Giinther and which is correct of that genus,” as restricted to the two
species L. niloticus and L. calcarifer. It may be further pointed out that in few,
if any, of his descriptions of new genera and species does Castelnau take notice
of this important character. The second exception, namely the coloration, is not
of sufficient importance to justify the retention of LD. antarcticus as a distinet
form, and this name also must, therefore, sink into a synonym of the typical
species. Further on (see p. 7) I shall have occasion to refer to a form which,
by a casual observer, would be set down as distinct, but which, scientifically
examined, is easily seen to be no more than a local variety of the common
Australian Perch due to the greatly changed conditions under which it exists.
This form may possibly be the LD. antarcticus of Castelnau, though, if so, it is
difficult to say why he should especially designate it the ‘Sea Perch.’
“‘Of L. victorie it is needless to say more than that no rational person is
likely to believe that the substitution of a four pointed for a single pointed spine
on the opercle is of itself sufficient to constitute a species.
‘“In Castelnau’s last form ZL. cwrtus it is only necessary to point out that
no reliance can be placed on the depth of the type specimen as indicative of even
* Brit. Mus. Catal. Fish., i, 1859, p. 67.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 7
a variety. The proportion of depth to length is given by him as 1 to 3-33,
but examples from Port Stephens, Shoalhaven, Shellharbor, and Ulladulla,
examined by the writer in preparation of this article, ranged between 1 to 2-75
and 1 to 3-5. This character, therefore, having proved invalid, and there being no
other on which to rely, it follows that ZL. curtus should be merged in P.
colonorum.’’
I find all the alleged differential characters attributed to these three latter
species—L. antarcticus, L. victoriw, and L. curtus—reproduced in one or other
of the four examples forwarded to me from New South Wales, thus further
proving their worthlessness, if that were necessary.
Nothing now remains but to consider the claims of Macleay’s Lates
ramsayi, and a very cursory glance at the description suffices to show that no
mandate to specific recognition can be with justice urged on its behalf. Stress
seems to have been laid by its author on the fact of this, to him, unique example
having been provided with ten dorsal spines and seven anal rays; variations such
as these are, as has been shown above, by no means so uncommon as is generally
supposed, though the coincidence of their occurrence in the same individual is no
doubt much more rare. In no other character does Macleay’s diagnosis differ
from that of individual specimens of our common Perch, and it, therefore, with
the preceding four, must be relegated to the list of useless synonyms. The fact
of its having been taken in a land-locked waterhole* doubtless had some effect on
its external appearance and thus helped to deceive its deseriber.
‘‘Mention was made above of a variety of the common Perch which
differed greatly in outward appearance from the ordinary fish. So far these
fishes have only been sent to the Australian Museum from the pools in the Snowy
River immediately below the Falls, but without doubt other rivers, both of New
South Wales and Victoria, will, now that notice is drawn to the form, be found
to have evolved under similar circumstances a similar variety. The differences
between it and the common market fish, which present themselves at a casnal
glance, are the much more elongate habit, the proportionally longer and more
powerful fins, and, though this is a much less important characteristic, the brilliant
silvery color of this variety. A moment’s thought, however, will suggest that
these differences, however important they may at first sight appear, are only to
be expected in fishes living under conditions, which differ so greatly from those
under which the species normally exists; for, being practically forced to inhabit
rapidly running waters, subject to sudden, severe, and periodical floods, causeci
for the most part by the melting of the snow on Kosciusko and the neighboring
ranges, it is patent that their changed surroundings and conditions of life would
induce a change in the direction indicated.
““To Mr. A. M. N. Rose of Campbelltown I am indebted for the knowledge
of this well marked form, that gentleman having forwarded to the Australian
Museum at my request two specimens, the first taken at Christmas the second late
? Not in the Parramatta River as stated by Macleay (6).
8 MEMOIRS OF THE QUEENSLAND MUSEUM.
in the autumn. Neither of these fishes showed the slightest traces of spawning
though, if they breed in the river, it is ineredible that the season selected for the
deposition of its ova by a fish notoriously fond of warm sheltered spots, should
be other than one of the two seasons during which my informant forwarded them.
Mr. Rose, however, is of opinion that these fishes do not breed, at least not in the
river, and he bases his opinion on the facts, certified to me by him, and through
him to me by other residents of the district, that the examples caught vary but
little in size, the usual length being from fourteen to eighteen inches, that no
young fish have ever been observed, that no matter at what season of the year
they are captured no milt or roe has ever been found in the ovaries, that they are
always in fine condition, and finally that they infinitely excel the estuary Perch
in flavor.
‘The obvious inference is that certain individuals having made their way
into the Snowy River at its entrance into the ocean gradually work upwards and
eventually find themselves in the pool below the Falls, and being unable to
proeeced, remain there until captured or swept downward by flood to the estuary
again; these latter having, since their enforced return into calmer and warmer
waters, grown sybaritic and fat, may possibly be Castelnau’s ‘not very common’
L. antarcticus, which he deseribes as ‘silvery’ and ‘very savory.’ ”’
The sterility of those fishes, which remain permanently in the pools below .
the Falls, would be satisfactorily accounted for by the coldness of the water.
Having now cleared the way I shall proceed to discuss the status of the
latest claimant to specific honors, Mr. Stead’s Percalates fluviatilis.
Stead (4) insists on
(a). ‘‘The far more elongate habit.’’
(3) In the four New South Wales examples on which these remarks are
wholly based the extreme variation in the depth of the body to its length is 2.44
to 2-87. This is by no means excessive and shows incontestably on how slender 2
foundation this character rests. In any case the somewhat more slender habit of
the fluviatile form and its rather longer and stronger caudal peduncle may
clearly be accounted for by the fish’s permanent occupation of swiftly running
water, as compared with the more equable ebb and flow of the estuarine tideways
which P. colonorum mostly affects.
(b). ‘The non-excavate character of the upper profile of the head, whach in
P. colonorum is universally concave—often highly so.’’
This contention has been dealt with above (see p. 2). It is interesting,
nevertheless, to point out that in Boulenger’s description of the species, he, with
six specimens before him drawn almost assuredly from estuarine sources, writes
‘“upper profile of head sometimes coneave.’’ The italics are mine.
(c). ‘‘The relatively greater thickness as compared with the body-length.’’
This is a character of minor importance. The forces contributory to the
formation of a more slender type of fish (see ‘‘a’’) are also doubtless at work
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 9
here to bring about in other directions a habit of body more suitable to the
changed conditions under which it now exists. For instance in swiftly flowing
mountain torrents a fish with a cylindrical body would offer much less resistance
to the current than would one of a deeper and more compressed form, and
complies with the primary law which enacts that every natural object shall
develop along the line of least resistance; it is probable that, as in this case, a
species, the environment of which had been changed from that of 2 quiet and
comparatively sluggish existence to one of continued and strenuous exertion,
should tend to assume a shallower and broader habit as the years roll by.
(d). ‘‘The shorter head.’’
In my four specimens the comparative measurements of the length of the
head to that of the body are for P. fluviatilis 2-62 to 2-88, for P. colonorum 2-55
to 2-67. Since, therefore, they are seen to overlap in so small a series, it is evident
that little or no reliance can be placed on this character as evidence of specific
differentiation.
(e). ‘‘The much more ctenoid character of certain body-scales in all but older
specimens.”’
This claim has also been dealt with above (see p. 2).
Stead also pointed out that the two forms differed in their habits, but
surely that was inevitable in view of their vastly altered conditions of life, and
should carry no weight with it.
The only other differential character mentioned by Stead (5) is set forth
in the following terms :—‘‘The tail too is larger and altogether more powerful
than that of the Estuary Perch.’’ With this contention I have dealt under (a).
In conclusion I assert emphatically that from this re-examination I see
nothing to cause me to change my previously expressed opinion, that all the
various forms of this fish should be united under the original name—colonorum.
There is absolutely no structural character by which P. fluviatilis can be separated
from the parent stock while on the other hand every conceivable shade of
variation exists, uniting the two forms. P. fluviatilis is doubtless identical with
the fish alluded to by me (loc. cit. p. 7) as having been sent to the Australian
Museum at my request by Mr. Rose from below the Falls of the Snowy River,
and rejected by me as not even worthy of varietal rank.
None of the authors, who have so overloaded this species with useless
synonyms, seem to have taken into account that owing to diversity of environ-
ment; climatic conditions, due not only to their geographical range, but also to
the varied altitude at which they may reside; the character of their native .
streams, whether sluggish and discolored with a muddy or weedy bottom, or clear
and sparkling as it flows over a bed of rock and gravel and sand; the nature of
the geographical strata over which the stream flows; the great diversity of the
food supply ; and many other fortuitous circumstances, much greater latitude of
variation should be allowed to fluviatile fishes than to their more equably situated
10 MEMOIRS OF THE QUEENSLAND MUSEUM.
marine relatives. And the variation is apt to be greatly accentuated when the
particular fish under review leads, as in this case, both a fresh-water and a brack-
water existence.
Reproduction:—During the late autumn and early winter months the
Bass drop gradually down, from the upper reaches of the rivers and the quiet
pillabongs, where they make their home during the greater part of the year, to the
estuaries, and finally to the purer waters of the open bay off the mouths of their
home streams. Here they shed their spawn during June, July, and August, the
season being proportionally ‘later the further we proceed north. The ova are
pelagic, and the fry, such as escape their numerous enemies, quickly work their
way up stream to seek the safer sanctuary of still lagune and billabong.
; Uses, etc.:—The Royal Commission on the Fisheries of New South Wales:
(1880) reported of this species—‘‘It is a very delicious fish, but never attains a
great size and is perhaps of more value for the sport it affords to the amateur
fisherman than as an article of food.’’ While, in the light of our present-day
knowledge, much of this estimate of the Bass’ value is liable to provoke a smile,
Tenison Woods, from whose work the above quotation is taken, gives as interesting’
an account of its habits and qualities as a sporting fish as any of his successors—
or more so. He writes—‘‘The perch affords good sport to anglers. It loves quiet,
shady, and deep holes in the rivers, but when the tide is flowing it may be caught
in the stream. It is very voracious. In winter the bait is a small mullet or
herring, or better still one of the large grubs that bore into trees. In the early
spring months it will take a moth readily, either sunk or on the surface. The
artificial salmon fly is also a splendid bait for trolling at this time. When moths
are scarce a frog is a good bait at night. It must be fastened so that it can swim
or, if dead, must be played upon the water to simulate a swimming frog. No
perch can resist that bait at night. In summer grasshoppers, especially that
known as the ‘Percher,’ a red species, are good bait, but the best is a black house-
ericket or an earthworm. This is a very attractive bait, and if the perch are in a
pool, the lines are no sooner down than the bait is taken. For the rest of the year
a prawn is the best bait, that is when crickets cannot be got. The bait should be
at least four feet from the float. In landing the fish great care should be used, as
the mouth is weak and is easily torn away.’’ These simple instructions appeal
mightily to me, some of them are so deliciously Waltonian in their touch, as for
instance ‘‘if the perch are in a pool,”’ ete.
Stead contributes the following information regarding its capture for the
markets :—‘‘The Estuary Perch is a familiar object to most people in the States:
of New South Wales and Victoria at least, as a few are generally to be seen
amongst the fish in every fishmonger’s stall. Particularly is this so after heavy
freshets in our coastal rivers and during the winter months. After heavy rains
have fallen on the watershed of, say, one of our coastal rivers, there is in a short
time a superabundance of fresh water, heavily charged with silt and often
decayed vegetable matter in the tidal portions of the river; and this has the
effect of driving out to the lower and cleaner estuaries or harbour waters, most of
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 1a
the inhabitants, chief amongst which are the Estuary Perch. As there are then
many more fish in a given area than previously, the fisherman is enabled to reap
a richer harvest with his meshing-nets and his hauling-nets, The same applies to
the winter time, when the fishes come down of their own free will to the open
water for the purpose of spawning.”’
Roughley tells us that—‘‘ The flesh of the Estuary Perch is firm and tasty,
qualities which render it a valuable and highly esteemed food amongst those who
dwell in the districts watered by coastal streams. In addition to this its capture
supplies a means of enjoyment to those far removed from the great variety of
fishes to be found in the coastal waters.’’ He continues it ‘‘is rarely seen in the
markets during the summer months. When the fish congregate at the mouths of
the rivers, usually during June, July, and August, the markets are kept fairly
well supplied.”’
Mr. H. K. Anderson considers that the Bass is undeniably the premier
game fish, whether native or introduced, of the Australian rivers.
Dimensions :—Stead remarks—‘‘Though a weight of 5 lb. is attained by
this fish, examples of that size are quite uncommon; the more usual size, as seen in
the markets, varying from about 12 oz. to 2 lb.’’ Tenison Woods mentions one of
74 lb, With reference to this weight I remarked—‘‘As many persons are
sceptical of this weight it may be worth mentioning that the writer has seen a
specimen, which had been retrieved at the Hamilton Wall, Brisbane River, after
an unsatisfactory encounter with a shark. This fish was weighed in my presence,
and turned the scale at 6 lb. 12 oz., notwithstanding that a large lump had been
bitten out of its flank as cleanly as if done by a surgeon’s knife.’’4
Range:—Rivers and estuaries of Southern and Eastern Australia from
the Murray River, S8.A., to the Pine River, 8.Q. Rivers of Northern Tasmania.
PLECTROPLITES Gill.
Plectroplites Gill, Proc. Acad. Nat. Sci. Phila., 1862, p. 236 (ambiguus): typonym only;.
id., ibid., 1863, p. 286; Bleeker, Arch. Néerl. Sci. Nat., xi, 1876, p. 267.
Ctenolates Giinther, Proce. Zool. Soc. London, 1871, p. 320 (macquariensis = ambiguus) ;
Ramsay & Ogilby, Proc. Linn. Soc. N. S. Wales, xii, 1887, p. 183; Boulenger, Catal.
Percif. Fish., 1895, p. 133.
Body subovate and compressed. Scales small, adherent, finely ctenoid, the
exposed surface concentrically striated, arranged in regular series. Lateral line
complete, not continued on the caudal fin, the tubes straight, not extending to
the margin of the scale. Head large, with more or less concave upper profile and
long pointed snout, partly naked, the mucigerous system strongly developed.
Mouth terminal and protractile, with moderate, oblique cleft, the lower jaw
projecting; maxillary almost wholly exposed, only the upper edge slipping
beneath the preorbital when the mouth is shut, naked; supramaxillary present.
Jaws, vomer, and palatines armed with villiform teeth, the premaxillary and
mandibular bands interrupted at the symphysis; pterygoids and tongue smooth.
* Ogilby, Commercial Fishes and Fisheries of Queensland, p. 14, footnote 9.
p. 14,
12 MEMOIRS OF THE QUEENSLAND MUSEUM.
Nostrils subequal and approximate, situated in front of the upper half of the
eye, the anterior valvular. Eyes small and superolateral, wholly anteromediar.
Preorbital, suborbital, and subopereular bones more or less finely serrulate ;
preopercle serrated, the teeth of the lower limb antrorsely directed ; operele with
two spines, the lower the longer, and a wide naked membranous flap ; posttemporal
exposed and serrulate. One dorsal fin, with x 10 or 11 rays, the spines strong
and pungent, the spinous portion longer than the soft. Caudal rounded, with 17
rays, the upper and lower simple, the others branched. Anal fin short, with ii 7
to 9 rays, the second spine longer and much stronger than the third. Pectoral
rounded, with 15 to 17 rays, the middle upper ones the longest. Ventrals inserted
below the pectoral-base, close together, with a strong spine and 5 soft rays, the
outer of which is produced in a filament. Gull-openings wide; gill-membranes
separate, free from the isthmus; branchiostegals seven’ ; pseudobranchiz present ;
gills four, a slit behind the fourth; gill-rakers rather long and stout, in moderate
number. Air-bladder present, large and simple. Pharyngeal bones densely
armed with minute conical teeth, Stomach cxcal; intestinal canal biconvolute ;
pyloric appendages in moderate number. Premaxillary processes not extending
to the frontals, parietal and supraoccipital bones not extending to between the
postfrontal processes; supraoccipital crest strong, not produced on the frontals.
Vertebre 11 + 15 = 26.
A monotypic genus, the single species inhabiting the rivers of the eastern
half of Australia northward to about the Tropic of Capricorn. Not found in the
cismontane rivers of Victoria and New South Wales southward of the Clarence
Watershed.
PLECTROPLITES AMBIGUUS (Richardson).
Datnia ambigua Richardson, Zool. Erebus & Terror, ii, Ichth., pt. 1, 1848, p. 25, pl. xix.
Dules ambiguus Giimther, Brit. Mus. Catal. Fish., i, 1859, p: 270; Klunzinger, Sitz. Akad.
Wien, lxxx, 1880, i, p. 348.
Ctenolates macquariensis Giinther, Proce. Zool. Soc. London, 1871, p. 320, pl. Xxxiii.
Dules auratus Castelnau, Proc. Zool. & Acclim. Soc. Vic., i, 1872, p. 55.
Dules flavescens Castelnau, Res. Fish. Austr., 1875, p. 10.
Ctenolates ambiguus Giinther, Zool. Challenger, i, 1880, Shore Fish, p. 32; Macleay, Proc.
Linn. Soe. N. 8. Wales, v, 1881, p. 354; Woods, Fish & Fisher. N. 8. Wales, 1882, p. 103;
McCoy, Prodr. Zool. Vic., dec. ix, 1882, p. 15, pl. Ixxxiv; Ogilby, Catal. Fish. N. 8. Wales,
1886, p. 11; Lucas, Trans. Roy. Soc. Vic. (2) ii, 1890, p. 18; Ogilby, Edib. Fish.
N.S. Wales, 1893, p. 22, pl. v; Boulenger, Catal. Percif. Fish., i, 1895, p. 134; O’Connor,
Proc. Roy. Soc. Queensl., xii, 1897, p. 109; Zietz, Trans. Roy. Soc. S. Austr., 1902, p. 265.
Ctenolates flavescens Macleay, ibid., p. 355; Ogilby, Catal. Fish. N. S. Wales, 1886, p. 11.
Plectroplites ambiguus Bleeker, Arch. Néerl. Sci. Nat., xi, 1876, p. 267; Waite, Syn. Fish.
N.S. Wales, 1904, p. 26; Stead, Fish. Austr., 1906, p. 97; id., Edib. Fish, N. 8. Wales,
1908, p. 55, pl. xxiii; Zietz, ibid., xxxiii, 1909, p. 268; Roughley, Fish. Austr., 1916,
p- 67, pl. xvii.
*Boulenger (loc. cit.) gives the number of branchiostegal rays as ‘‘six or seven.’’
I have at different times examined quite a large number of specimens, and have invariably
found seven. This should, therefore, be taken as the normal number.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 13
YELLOWBELLY.
Golden Perch; Murray Perch; Kaakaalain (Murrumbidgee natives); Tarkee
(Lower Murray natives) ; Callop (Professional Fishermen of Victoria, fide
Anderson ) .®
Type localities:—West Australia (D. ambigua).
Macquarie River (C. macquariensis).
Riverina (D. auratus).
Murray River (D. flavescens).
Upper contour of body rising abruptly at the nape and much more arched
than that of the ventral, its width at the shoulders 1-63 to 1-88 in its depth, which
is 2-6 to 3 in its length and as long as or a little less than the length of the head.
Caudal peduncle about four ninths deeper than long, its least depth 7 to 7-75 in
the body-length. Head about one fifth deeper than wide, its width 1-67 to 2 in its
length, which is 2-55 to 2-67 in that of the body. Diameter of eye 1-5 to 2 in the
length of the snout and 6 to 8 in that of the head; depth of suborbital two thirds
to one half of the eye-diameter ; interorbital region convex, its width 5 to 6 in
the length of the head. Maxillary extending to below or a little beyond the
middle of the eye, its length 2-75 to 2-86 in that of the head, the width of its
distal extremity as much as to two thirds of the eye-diameter. Hinder border
of preopercle evenly serrated, the teeth at the angle much enlarged, those of the
lower border usually arranged in groups.
Tooth-bands of jaws broadest in front, gradually narrowing to an obtuse
point behind; vomerine teeth forming an obtusely triangular patch; palatine
bands curved, similar to those of the jaws.
Scales in 77 to 88 series above the lateral line; 12 or 13/1/33 to 37 scales
between the first dorsal spine and the ventral ridge. Cheek-scales smaller, about
half as large as those of the opercle. Lateral line forming a long curve from the
upper edge of the opercle to below the last dorsal ray. .
Dorsal fin originating a trifle behind the pectoral-base; fourth spine
hongest, 2-55 to 3-14 in the length of the head, and a little lower than the soft
portion, the base of which is clothed with minute scales, its length 1-6 to 1-9 in
that of the spinous portion. Caudal fin 4-5 to 4-88 in the body-length. Anal fin
originating below the first dorsal ray and terminating somewhat behind the last
ray ; second spine very strong, usually a little longer than the third, its length
2-83 to 3-7 in that of the head and 1-25 to 1.4 in that of the second and longest ray.
Pectoral fin 1-75 to 1-9 in the length of the head, the fifth and sixth rays the
longest, extending to below the sixth dorsal spine. Ventral fin, without the
filament, a litile shorter than the pectoral; the filament more developed in the
young, in which it reaches nearly to the vent, obsolete or nearly so in large
examples.
* Anderson writes—‘‘ Regarding aboriginal names for this fish I do not know, and believe
that most of those names used by the present aborigines are ‘pidgin English’ and quite
unreliable; hence I place no value on them. ’’
14 MEMOIRS OF THE QUEENSLAND MUSEUM.
Gill-rakers 12 or 138 on the lower branch of the anterior arch, the longest
equal to or rather less than the gill-fringes and about one ninth of the length of
the head. Pyloric ceca 11.
Upper surface of body varying from bronzy to slaty green, the sides more
or less richly golden olive, shading below to deep straw-color or even white.
Head above purple or purplish brown, the sides green with violet and orange
reflections. Dorsal spines lavender, the connecting membranes yellowish gray
dotted with blackish; soft portion tawny yellow, becoming darker towards the
margin; caudal fin purplish brown, black-dotted; anal basally golden, the distal
third deepening to a violet-brown; pectorals and ventrals golden, uniform or
dusky-dotted.’
In the Thomson River, and possibly elsewhere, there is a dark color-variety,
locally known as the ‘‘black yellowbelly,’’? in which the normal coloration is
more or less completely obscured by an overlying blackish mantle. This may
cover the entire head, body, and fins, or may take the form of numerous
small black spots, in some cases generally distributed in others arranged in
irregular groups, between which the more typical old gold coloring appears.
Structurally no difference can be detected between the two forms. For the
opportunity of examining this interesting variety I am indebted to their
collector, Miss Josephine Bancroft.
Described from nine specimens, measuring from 225 to 380 mm. Two of
these are from the Condamine at Dalby, and were presented to the Museum by
Messrs. McNaught and Williams; two are from the Upper Noosa, collected by Mr.
J. H. Stevens and myself; and the remaining five are from the Thomson River
near Longreach. Of the last three belong to the normal, two to the black, form;
all five, however, were taken from the same waterhole.
Variation:—In some respects Boulenger’s deseription differs from mine,
and notably in regard to the size of the eye, which according to him is ‘*4-5
(young) to 6 (adult) in length of head,’’ whereas I find it ‘‘6 (young). to §
(adult) ’’ in the same; similarly he gives the width of the distal extremity of the
maxillary as 1-5 to 2 in the eye-diameter as compared with the 1 to 1-5 of my
measurements. His dorsal spines also are longer than I find them, being 2 to 2-5
in the length of the head as against roughly 2-5 to 3 in mine.
Historical :—The earliest notice of this fine species occurs in the first part
of the Ichthyology of the Erebus and Terror, where Sir John Richardson
published a description and figure taken from two dried specimens, measuring
“Of the numerous freshly caught examples which I have handled, not one showed the
slightest trace of the scarlet markings mentioned by Castelnau (1), and reproduced by Tenison
Woods and Roughley in their respective accounts of this fish, while the absence of any
reference thereto by McCoy and Stead suggests that their experience coincides with mine.
Castelnau’s examples were obtained in the Melbourne market, where they were exposed for
sale after a railway journey of many miles from their Murray haunts. Consequently I have
little doubt that the scarlet patches were merely due to the extravasation of the blood in the
.affected tissues.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 15
respectively 216 and 356 mm., under the name Datnia ambigua, the specific name
being suggested by his uncertainty as to which genus, Datnia or Dules, the fish
should more properly be referred. These examples are said to have been
collected in ‘‘ Western Australia’’ by Sir George Gray, but there is apparently
some error as to the alleged locality since, so far as is known at the present day,
the species does not occur under natural conditions in any part of the great
western State, the rivers of South-Western Queensland being, so far as I can
ascertain, the limit of its range in that direction. Fifteen years after the
publication of Richardson’s description Giinther (1) removed the species to the
genus Dules, giving in the British Museum Catalogue of Fishes a very
inadequate account of it, doubtless owing to the fact that he only had the
original dried specimens to work on. After an interval of a few years he obtained,
through the agency of Mr. Gerard Krefft, a 300 mm. example from the
Macquarie River, New South Wales, preserved in liquor, and not at firsi
recognizing its identity with Richardson’s fish, proposed for it the new generic
and specific names of Ctenolates macquariensis which, having in the meanwhile
realized its true relationship, he subsequently altered to Ctenolates ambiguus,
which name remained in general use until towards the close of last century.
Gill, however, had previously recognized the necessity of separating the
Australian fish from both Datnia and Dules, and proposed the generic name
Plectroplites for its reception. This name was rejected by Dr. Boulenger in his
account of the serranoid fishes for the good and sufficient reason that at first it
appeared as a typonym only, without any indication of the characters upon
which it was based. But Boulenger unfortunately overlooked the fact that in the
following year Gill published a diagnosis of his genus which, though short, was
amply sufficient to establish the validity of the name, so that it is now generally
referred to as Plectroplites ambiguus. Count Castelnau then took a hand in
building up a synonymy for our ‘‘yellowbelly,’’ describing it under two new
names in the course of three years. In his first essay he calls it the ‘‘Murray
Golden Perch,’’ and describes it, from large specimens acquired in the
Melbourne market, under the name of Dules auratus, stating that ‘‘it appears to
be common in the Murray and other rivers of the Riverina,’”’ that it ‘‘is much
esteemed for the table,’’ and that ‘‘it often weighs five and sometimes seven
pounds.’’ He notes its similarity to the ‘‘Dules ambiguus of Richardson and
Gunther,’’ but considers that the difference of ‘‘one ray less in the anal and also
less seales on the lateral line’’ warrants him in separating it from that species.
I think, however, that most writers will agree with McCoy that ‘‘these
differences, which induced Count Castelnau.to propose a new specific name, are
unimportant.’’ His next attempt, in which he describes a 480 mm. specimen,
also from the Murray River, as Dules flavescens, is equally futile.
Reproduction:—In 1893 I wrote in regard to this phase of the life history
of the Yellowbelly—‘‘These important considerations in the economy of our
freshwater fishes will never be properly understood until a competent officer shall
be appointed by the Government to report fully on these and all other matters
16 MEMOIRS OF THE QUEENSLAND MUSEUM.
connected with the fishes and fisheries of our transmontane river systems.’’ Such
an officer has since been appointed by the Government of New South Wales, and
I am here privileged, through the courtesy of the Under Secretary, Chief
Seecretary’s Department, to record Mr. H, K. Anderson’s—the officer in question
—observations on the breeding of this fish in the waters under his control.
These are of such interest that I deem it advisable to publish Mr. Anderson’s
remarks in extenso :-—
‘‘In the spring of 1916 and 1917 I was entrusted with the conduct of
experimental hatchery operations in relation to Murray Cod and Golden Pereh,
and the following information regarding the latter species is based on the data
gathered on these occasions.
‘‘TIn the Murrumbidgee River at an altitude of between 370 and 530 feet
‘ above sea level many Golden Perch, usually large fishes from five to ten pounds
in weight, spawn during October and November in a flooded river. If the season
is late, cold and wet, with snow water coming down stream, spawning is
considerably retarded, while abnormally warm weather accelerates it.
‘‘The condition of many of the smaller fishes handled by me leads to the
belief that there is an autumnal as well as a vernal spawning, but of this I cannot
as yet speak with certainty. Examples of fishes with well developed ovaries are
captured by net fishermen throughout the whole year.
‘‘Kach spawning female, usually found without female companions, is.
apparently accompanied by a large number of ripe males. I have never yet
netted a ripe female Golden Perch without capturing at the same time, and in
the same net, from twenty to fifty ripe males.
‘‘The fishes apparently spawn at the edge of the current near the bank,
below a spot where a bend in the river leaves a projecting point with compara-
tively still eddying water behind it. It is in such places that J have caught all
the ripe and most of the nearly ripe females so far examined. As a rule the fish
were kept in captivity a few days pending complete ripeness. Many Golden
Perch, however, spawn in lagoons and outer channels of the river, where the
current is not so strong as in the main stream.
‘“‘The fertile eggs are semi-buoyant, practically of the same specific
gravity as water, and appear to be left by the fishes quite at the mercy of the
current.
‘‘The eggs are probably deposited in batches, 50,000 to 100,000 at a time.
I do not think the fish, however ripe, has power to eject all her eggs at once
because, when artificially stripping, only a limited number of eggs ean be
extruded without undue pressure, but I found that, after giving the fish a few
minutes rest (rolled in wet bagging to prevent struggling), another batch of eggs.
can be stripped, and so on until all are extruded.
‘“A five pounds Golden Perch earries from 750,000 to 1,000,000 eggs.
‘The unfertilised eggs when first stripped are of a very pale greyish.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. ay
yellow tint. They measure thirty seven to the linear inch (about -686 mm. each).
‘“Two hours after fertilisation, at an average temperature of 65° F., the
eggs are perfectly transparent under the microscope, moving about in the jars
from the bottom to the surface; when water was allowed to drip into the jars for
aération the eggs circulated in ceaseless movement all over the jars from top to
bottom ; some, probably the unfertilised eggs, floated.
‘“At two hours after fertilisation the eggs measured thirteen to the linear
inch (about 1-954 mm. each), and three hours after stripping, when absorption
of water was complete, they measured eight to the inch (3-175 mm, each).
“The egg, after full absorption of water, becomes slightly ovate and is
perfectly transparent, resembling a tiny soap bubble with a glistening oil globule
at the top.
‘“The eggs referred to above were stripped at 4-35 p.m. on 18th October,
1917. The first apparent development was visible at 10-15 a.m. on 19th October.
This appeared as a tiny foggy opaque spot at the bottom of the egg.
‘‘At 6 p.m. on the 19th October, the foggy spots in the egg had increased
in size to nearly one eighth of its capacity, but had assumed no definite shape.
At 8-50 the faint outline of the embryo was discernible. At this stage it is
colorless, semitransparent, and occupies the bottom of the egg. At 10-50 the
embryo, ineluding the opaque foggy mass round which it was curved, had
increased in size to about one fourth of the capacity of the egg.
**On the 20th October at 5 a.m. the embryo had further increased, and the
eye-spot was visible without magnification. At 12-10 p.m., under a magnification
of about fourteen diameters, movement of the embryo was plainly visible, and the
foggy mass above referred to had considerably decreased in size. At 10 p.m.
the embryo was clearly visible to the naked eye, apparently attached by the head
to the oil-globule, and its wriggling movements were distinctly seen without a
magnifying glass. At midnight the embryo was still suspended from the oil-
globule struggling vigorously in the egg. At 1am. on the 21st fully 75 per cent.
of the eggs in the jars contained living struggling embryos.
‘At 3 a.m. on the same day a cyclonic wind-storm capsized the oil-stove
used to warm the tent where the jars were kept, which with its contents was
destroyed by fire in a few moments. Several jars of eggs were carried outside the
burning tent, but as the temperature fell rapidly all were lost. I think it
probable that the eggs would have hatched out at about 10 a.m. that morning.
‘* Although I continued netting until the 16th November, and captured
many hundreds of Golden Perch, none in spawning condition were secured. The
season was exceptionally late, wet and cold, and a am of opinion that the general
spawning would not take place until December.’
Mr. Anderson supplies also the following note :—
‘‘In November, 1916, similar work was performed in almost identical
B
18 MEMOIRS OF THE QUEENSLAND MUSEUM.
conditions, and the records obtained were practically the same as above. Hggs
collected at 11 a.m. on the 24th hatched out during the afternoon of the 26th.
Owing to a sudden rise in temperature that day—minimum 50°, maximum
95°—the water in the hatching shed reached 75°, and all the fry died. The
surface of the jars and troughs was covered with oil from the eggs, and a great
number of dead fry, about the size of a comma in ordinary newspaper text, were
floating at the surface. The shells of the eggs retain their transparency, and
quickly dissolve without leaving any sediment or residue.”’
It is much to be regretted that Mr. Anderson’s two attempts to elucidate
the larval development of this fish met with unmerited failure in both cases
through causes entirely beyond his control, and it is to be hoped that the very
failure will further stimulate him to earry on future experimentai work to a
successful conclusion.
According to Mr. Glencross Smith the Yellowbelly is full-roed early in
September on the Darling Downs.
Uses, etc.:—All authorities coneur in their appreciation of the edible
qualities of this species. Stead writes—‘‘As an article of food the Golden Perch
is very well and favourably known, both in the western areas (of New South
Wales) and in the cities of Sydney and Melbourne.’’ Roughley tells us that—
‘* As an edible fish the Golden Perch ranks second only to the Murray Cod amongst
our indigenous fresh-water fishes. Its flesh is firm and tasty and is well suited to
the many methods of preparation for the table. It is deservedly popular and finds
a ready sale in the markets.’’ Glencross Smith, while acknowledging its
excellence, places it third in rank among our fresh-water food-fishes, considering
it inferior to the catfish (Zandanus tandanus) and the Murray Cod, while
Anderson places above it the Trout Cod (Oligorus mitchelli) and the Murray
Cod.2 However he continues—‘‘I do not desire to dogmatise, as I know several
men of wide experience, including Mr. Henry Dawson, late Inland Fisheries
Commissioner, who place the Golden Perch the first among our inland fishes.”’
From the little I know gastronomically of the two species I am inclined to agree
with Mr. Dawson.
From the angler’s point of view they are held in good repute, being
plucky and determined fighters when hooked. Writing of them in this connection
Mr. Smith says—‘‘ Yellowbellies lurk in weedy patches under the shade of logs
and overhanging trees, and dart out upon their food, taking it with a rush, and
are more particular about their food than Cod. They prefer frogs, shrimps,
yabbies, worms, and the white grub from the River Oak (Casuarina) ; also
crickets; but they do not care for fish bait. They are fine sporting fishes and will
take a spinning bait, and were at one time very plentiful in all our streams, but
are gradually being wiped out; dynamite, nets, and no close season being the
SIn Part II of the present volume I hope to deal with the various phases of the genus
Oligorus.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 19
ecause.’’ He has also remarked that the Yellowbelly is the most short-lived of our
fresh-water fishes after its removal from the water, invariably dying within five
minutes of its capture.
Anderson remarks—‘‘ The food of the Golden Perch consists principally of
small fishes, including its own species, yabbies, prawns, and insect larve. It
prefers live food. The most effective baits for the capture of this fish are small
live fishes, yabbies, prawns, and artificial spinners. The Aborigines on the
Murray seem to prefer the yabbie. The Golden Perch will not, I think, rise to the
surface for an artificial fly, but a salmon fly skilfully worked through the water,
particularly if fitted with a spinning head, will tempt them. From the point of
view of sport I consider the Golden Perch second among the fishes of the Murray
‘System.’’
Stead informs us that ‘‘it is pre-eminently what might be ealled a ‘still-
water’ fish, being very abundant in lagoons and billabongs. It is also a ‘mid-
water’ fish; that is, it swims at some distance from the bottom usually. It is a
fish of the plains rather than one of the mountains.’’ Further on he says—
‘‘Though taken by hook and line, the usual method of capture is by means of a
short meshing-net (gill-net) which is set at night across the billabong or lagoon.”’
And, quotes the evidence of a professional fisherman, as telling him ‘‘that one of
this species will blunder straight into a meshing-net, apparently without seeing
it; while a Murray Cod under the same circumstances would swim up to within
a Short distance of the net, and then quickly take fright and double back again.’’
Dimensions :—In large rivers, such as the Murray and its more important
tributaries, the Yellowbelly grows to a considerable size. The largest weighed by
Mr. Anderson was a female, taken in Yanco Creek, which turned the scale at
14 Ib. 2 oz. This, however, seems to be quite an unusual size, most writers being
content to credit it with a maximum weight varying between 7 and 10 lb.; while
eecording to Roughley the average of the fishes sent to the Sydney Market is
from 4 to 5 Ib. In the Queensland rivers, with their lesser volume of water, it
does not as a rule attain to such a size as in more southern streams, nevertheless
specimens approximating to the 10 lb. limit are occasionally captured, and I have
before me now a photograph, given to me by Mr. Glencross Smith, of an example
taken in Oakey Creek, which weighed 9 lb. 12 oz.
Range:—All the rivers of the Murray System; cismontane rivers of
Eastern Australia from the Clarence to the Dawson; rivers of South-Western
Queensland—Barcoo, Landsborough, Cooper, and Diamantina (Anderson) ;
Thomson River, common (Miss Bancroft). Dr. T. Baneroft writes—‘‘The
Yellowbelly does not occur in the Burnett River, but I caught a few by netting
an the Upper Dawson.”’
29 MEMOIRS OF THE QUEENSLAND MUSEUM.
Part XI.—LUTIANIDZ (No. 2).
LUTIANUS NEMATOPHORUS Bleeker.
(Plate I.)
Mesoprion nematophorws Bleeker, Act. Soc. Sci. Indo-Neerl., viii, 1860, Celebes, p. 56; id.,.
Vers]. Akad. Amst., xii, 1861, p. 46.
Lutjanus nematophorus Bleeker, Verh. Akad. Amst., xiii, 1873, Revis. Lutjan., p. 12; id., Atlas-
Ichth., viii, 1876, p. 47, pl. celxxxv, fig. 3.
THREADFINNED SEA PERCH.
Type locality :—Celebes.
Body deeply subovate and strongly compressed, the dorsal and ventral
contours symmetrical, its width 2-55 in its depth, which is 2-55 in its length to the
-root of the caudal and one eighth more than the length of the head. Caudal
peduncle three eighths longer than deep, its least depth 2-5 in the length of the
head. Head two ninths longer than deep, its upper profile behind the snout and
that of the nape gently and evenly rounded, its width 2-25 in its length, which is
2.85 in that of the body. Snout long and pointed, with linear moderately
acclivous profile, its length 2-33 in that of the head. Eye small, its diameter 2-37
in the length of the snout, 5-44 in that of the head, and 1-5 in its distance from
the end of the maxillary groove. Interorbital region convex, its width one eighth
more than the eye-diameter and 4-87 in the length of the head. Jaws subequal;
maxillary extending to below the anterior border of the eye, its length 2-5 in that
of the head, the width of its subtruneate distal extremity 1-3 in the eye-diameter.
Preopercle with a slight emargination above the angle, in and beyond which it is.
feebly serrulate ; opercle without spine, the lobe obtusely pointed; post-temporal
feebly serrated.
Scales in 57 series above the lateral line, disposed somewhat obliquely to:
the dorsal profile, in 51 horizontal series below the line; 8/1/22 scales between the
first dorsal spine and the ventral ridge. Head naked, except the opercle, sub-
opercle, a patch from seven to nine scales wide on the cheek posteriorly, a triseria!
postorbital band, and a short biserial temporal band. Lateral line with 54 pore-
bearing scales, the tubes branched. Soft dorsal and anal fins each with a well
developed scaly sheath, the membranes naked ; accessory ventral scale small and
blunt, 1-5 in the eye-diameter.
Dorsal fin with x 17 rays, originating above the lower angle of the
pectoral-base, the spinous portion one fifth longer than the soft; spines iow,
slender but pungent, increasing in length to the fifth, beyond which they deerease
to the ninth; last spine a little longer than the fifth and 3-44 in the length of the
head; fourth to seventh rays produced and filiform, extending to or beyond the
extremity of the caudal; beyond these the rays gradually decrease, the last being’
about equal to the fifth spine. Caudal emarginate, the middle rays 1-5 in the
upper lobe, which is 3-67 in the body-length. Anal fin with iii 9 rays, originating
below the third dorsal ray; spines short and stout, the third longest, 4-14 in the
length of the head; soft portion with acutely pointed outline, the anterior rays
produced, the fourth the longest, extending beyond the base of the caudal, length
\
b
’
*
1
at
r
‘
r
Weal apa”
THE QUEENSLAND MUSEUM.
MEMOIRS OF
PLATE I.
QUEENSLAND FISHES.
‘IoyooT_ SAYOHAUOLVWAN
SONVILO'T
‘yap ‘aywnig syphyg
Face page 21.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 21
of soft portion 2-37 in its height, 1-75 in that of the dorsal fin, and 2-55 in that of
the head. Pectoral with 16 rays, its length 1-22 in that of the head; fifth ray
longest, extending to below the first dorsal ray. Ventral inserted below the end
of the pectoral-base ; spine rather weak but much longer than the longest dorsal
spine, 1-75 in the outer rays, which are subequal, 1-13 in the length of the
pectoral, and reach almost to the vent.
Gill-rakers short and stout, 10 on the lower branch of the anterior arch,
the longest 1-8 in the eye-diameter.
Roseate, the lower surfaces lighter and obscurely tinged with yellow; body
with ten narrow, more or less wavy, longitudinal blue bands; the upper pair
meet across the nape; the second pair extend upon the occiput, where they are
broken up into irregular spots; the third reaches to above the post-temporai,
beyond which it is similarly broken up ; the fourth is continued forward to the eye
and, though less clearly, along the upper eyelid and across the nostrils, meeting
near the end of the snout; the fifth ceases at the opercle; these five pairs termi-
nate at various points along the base of the dorsal fin; the sixth reaches to the
eye and is continued on, the snout as a series of small blue spots, which meet
across the tip of the snout; the seventh extends to the opercle; the eighth to the
middle of the preorbital; the two lower to the pectoral, in front of which are
some irregular spots and bars; a narrow blue ecross-bar between the eyes. All
the fins pinkish, the caudal with a narrow dark terminal band.
Described from a fine specimen, 353 mm. long, trawled in Hervey Bay,
and presented to the Queensland Museum by the Department of Fisheries
through Captain Hoult of the ‘‘Bar-ea-Mul.’’
The species was only known previously from two young examples,
measuring 82 and 86 mm., recorded by Bleeker from Singapore and Celebes. Its
rediscovery on the South Queensland coast is, therefore, of more than ordinary
interest.
Part XVI.—OPISTHOGNATHID& (No. 1).
Opisthognathide Jordan & Evermann, Fish. North & Mid. America, pt. 3, 1898, p. 2279;
Regan, Ann. & Mag. Nat. Hist. (8). xii, July 1913, p. 138.
THE JAWFISHES OR SMILERS.
Body elliptical to elongate, moderately compressed, more or less completely
covered with small, cycloid scales. Lateral line single, incomplete, running close
to and parallel with the dorsal contour, ceasing below the anterior dorsal rays.
Head large and naked, with strongly curved anterior profile, its upper surface
smooth. Mouth terminal, horizontal or nearly so, protractile, the cleft usually
very wide; jaws normally formed; maxillary broad and exposed, with supple-
mental bone. Jaws with bands of villiform or ecardiform teeth; vomer usually
with a few teeth; palatines and tongue toothless. Two nostrils on each side.
Eyes very large, lateral, anterior. Opercles unarmed. Dorsal fin continuous, the
spinous and soft portions of nearly equal length, the spines slender and flexible,
22 MEMOIRS OF THE QUEENSLAND MUSEUM.
passing gradually into soft rays. Caudal rounded or lanceolate, with 14 or 15
principal rays, the outer ray above and below simple. Anal fin long and low,
with two weak spines. Pectorals short and fan-shaped. Ventrals inserted in
advance of the pectorals, close together, each of a spine and five rays, and without
accessory scale. Gill-openings wide; gill-membranes partially united, free from
the isthmus; six branchiostegals; pseudobranchie present; gills four, a slit.
behind the fourth; gill-rakers long and slender. Air-bladder present, small. No
pyloric eewea. A narrow subocular shelf. Skull narrow between and expanded
behind the orbits; postorbital part evenly convex above; occipital crest only on
the posterior face of the skull, which is long and oblique; no parietal crest;
exoccipital condyles separate; prootics forming a roof for the myodome; basi-
sphenoid present. Foramen in hypercoracoid; radials rather broad and flat, two.
. on the hypercoracoid and two on the hypocoracoid. Vertebre 29 (10 x 19)°®;
precaudals with parapophyses from the fifth; three ribs sessile, five on para-
pophyses; epipleurals on ribs.
The Opisthognathide, though weak in point of numbers, forms a most
interesting family of trachiniform percoids; it consists of fishes of small or
moderate size, inhabiting rocky and coralline bottoms within the tropical and
temperate zones. Its distribution is peculiar, no species having so far been
recorded from the Mediterranean nor the Eastern Atlantic, nor from any of the
Pacific Islands, nor the West Coast of the Americas, except the Gulf of California,
where an isolated colony, comprised of five species, exists. The extreme limits of
their polar range lie between lat. 40° N., where Gnathypops hopkinsi has been
taken off Misaki, Japan, and lat. 34° S., where Merogymnus jacksomensis is
found. These fishes are everywhere of rare occurrence, many of the species being
only known from the single example described. The ‘‘Smilers,’’ to give them
the local vernacular name bestowed upon them, so a friend tells me, ‘‘because of
their fine open countenanee,’’ are essentially carnivorous and rock-loving fishes,
delighting in boulder-strewn shoals and coral reefs at a moderate depth. Hence
we find that in America as here many of the species are only known from off-
shore snapper grounds and similar localities, where they are occasionally taken
by hand-line in company with more valuable fishes or even from the latter when
captured disgorging them. The species vary greatly in coloration, some of them,
such as my Merogymnus eximius, being arrayed in a livery of most gorgeous
splendor, while others, as my Gnathypops inornata, are soberly clad in uniform
brown. One of our species, G. maculata, is said by the pearl-fishers to scoop holes.
in the sand among the sea-fan forests of our tropic seas, from which it sallies.
forth to pounce upon a passing prey, returning again to its lair after each
_ excursion. Regarding this habit Mr. W. H. Longley, from personal observation
when ‘‘equipped with a diving hood in the unknown world of coral labyrinths at
the bottom of the sea,’’ gives some interesting information, which may advan-
tageously be reproduced here.*° He writes:—‘‘Gnathypops aurifrons prepares.
° Vertebre 27 to 34, fide Jordan & Evermann, ut supea.
* American Museum Journal, xviii, February 1918, p. 81.
KDIBLE FISHES OF QUEENSLAND.—OGILBY. 23
its own shelter in sandy places where the substratum is sufficiently compact to
make successful tunneling possible. Jaws and gaping mouth are its only
entrenching tools, but meet its every need. It is found not uncommonly upon
the open reef in little colonies, the formation of which is probably due in part. to
the discontinuous occurrence of suitable bottom rather than to the social instincis
of the fishes themselves. During the day, if undisturbed, Gnathypops may be
regularly observed resting nearly motionless in a semivertical position above its
burrow. When alarmed it retreats into its hole tail foremost and conceals itself
until the disturbance outside has ceased. Then it reappears cautiously, its beady
black eyes being so situated that it is able to sweep the horizon with minimum
exposure. If nothing happens to renew its alarm, it mounts a little farther until
its ventral fins are free, rests for a moment in the mouth of its burrow, and finally
rises easily and gracefully to its original position. Still another of these fishes
(as yet unidentified) shows a different variation of the tubiculous instinet. It
lives in holes, quite possibly worm-tubes, which it discovers ready formed in
pieces of dead coral upon the bottom, Its most striking structural feature is the
immense dorsal fin, which when raised seems nearly as high as the fish is long.
Its most interesting habit is that of protruding its body for about half its length
from the chamber it occupies, and then elevating and depressing its great fin
rapidly as if it were wigwagging in piscine code. This impression is heightened
when two individuals separated by no great distance stand erect and repeat the
performance in alternation.’’ Nothing is known as to the breeding habits of these
fishes, nor have I ever seen an example with ripe spawn, though all those which 1
have handled were fully adult. That the young are never found along the fore-
shores nor in the débris of the seine net seems to demonstrate the demersal
character of the ova. No data are available as to their edible qualities, unless it
be that of an acquaintance, who took one home and had it fried; he told me it
was ‘‘as good as any other rockfish.’’ “On account of their scarcity and the
difficulty of obtaining them all the species are greatly in request for museum
collections, and every example should, therefore, be carefully preserved and
forwarded, with as little delay as possible, to the nearest scientific institution. Six
genera and thirty species are now recognized. As mentioned by Waite it has
been suggested in some quarters that Gnathypops may merely be the female of
Opisthognathus, but the fact that the latter genus is quite unknown in Australian
waters sufficiently contravenes any such theory.
Appended is a key to the opisthognathoid genera. Those which are
printed in italics have not so far been found in Australian seas, but, with the
exception of the West Indian Lonchopisthus there is no insuperable reason why
the other genera should not occur here.
a’. Maxillary about as long as the head, produced behind ina flexible lamina i. Opisthognathus.”
a’. Maxillary much shorter than the head, but extending well beyond the eye, its distal extremity
truncate.
“Cuvier & Valenciennes, Hist. Nat. Poiss., xi, 1836, p. 498. Type O. sonneratui =
O. nigromarginatus Riippell.
24 MEMOIRS OF THE QUEENSLAND MUSEUM.
b*. Caudal fin rounded.
c. Trunk anteriorly naked oe = ev 45 + i% ii. Merogymnus.
c. Trunk everywhere scaly.
ad. Dorsal spines all simple ste 5 ; ae ey ye iii. Gnathypops.
d*. Anterior dorsal spines transversely forked Ee ie sf iv. Stalia.”
6°. Caudal lanceolate .. = 3. ar 5k ate ve .. Vv. Lonchopisthus.*
a*, Maxillary not extending beyond the eye .. Fae 5p: a fe vi. Owstonia.™*
i. MEROGYMNUS Ogilby.
Merogymnus Ogilby, Proc. Roy. Soc. Queensl., xxi, 1908, p. 18 (eximius); MeCulloch, Ree.
West Austr. Mus., i, 1914, p. 216.
Body elliptical or elongate-elliptical, the greater part of the trunk naked.
Maxillary much shorter than the head, its distal extremity much dilated. Jaws
with bands of villiform teeth, the outer row scarcely larger than the others.
Caudal fin rounded. Gill-rakers in rather large number, long, and slender.
Eastern and South-Eastern Coasts of Australia. Two species.
a’, Seales very small, covering about half of the sides and belly .. ess 1. eximius.
@. Seales larger, but narrowly entering the trunk; belly naked .. Se 2. jacksoniensis.
1. MEROGYMNUS EXIMIUS (Ogilby).
(Plate IT.)
Merogymnus eximius Ogilby, Proc. Roy. Soc. Queensl., xxi, 1908, p. 18.
' HARLEQUIN SMILER.
Type locality :—Snapper Banks off Moreton Bay, S.Q.
Body elliptical ; its width at the shoulder 1-7 to 1-83 in its depth, which is
3-62 to 3-75 in its length and 1-25 to 1-33 in the length of the head. Caudal
peduncle four fifths deeper than long, its least depth 2-55 to 2-7 in the length of
the head. Head a little deeper than wide, the fronto-occipital profile evenly
rounded, its width 1-4 to 1-5 in its length, which is 2-83 to 2-9 in that of the body.
Snout short, with linear or gently rounded, vertical profile, its length 1-75 to 1-8
in the eye-diameter, which is 2-83 to 3-2 in the length of the head. Interorbital
region narrow and feebly concave, its width 3-43 to 3-75 in the eye-diameter.
Maxillary extending to about one diameter of the eye behind the eye, its length
1-4 to 1-5 in that of the head, the width of its distal extremity about two ninths
of its length.
Seales extending forwards on the trunk to below the middle of the
* Jordan & Snyder, Proc. U. S. Nat. Mus., xxiv, 1902, p. 495. Type S. histrio from
the Japanese coast.
* Gill, Proc. Acad. Nat. Sci. Phila., 1862, p. 241. Type L. micrognathus from Cuban
waters.
“Tanaka, Journ. Coll. Sci. Tokyo, xxiii, 1908, p. 47. Type O. totomiensis from the
seas of Japan.
Pose 4 ie.
Fhe,
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QUEENSLAND MUSEUM.
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Face page 25.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 25
appressed pectoral, anteriorly distant and deeply imbedded, becoming more
crowded and finally imbricate on the tail; belly scaly. Lateral line composed of
about 80 short tubes, terminating below the 5th or 6th dorsal ray.
Dorsal fin with xi 13 rays, originating above the tip of the maxillary;
outer border of the fin linear, the spines and rays gradually increasing in length
to the 9th to 11th rays, which are 1-63 to 1-75 in the length of the head. Caudal
fin long, 3-9 to 4-3 in the body-length. Anal with i 12 or 13 rays, originating
below the 1st dorsal ray, its longest ray 1-7 to 1-83 in the length of the head.
Pectoral rounded, with 19 rays, the middle the longest, 1-88 to 2 in the head.
Ventral produced, about one third longer than the pectoral, the 2nd and 3rd rays
subequal, reaching to or nearly to the vent.
Gill-rakers long and slender, with the inner edge feebly spinulose, 16 + 26
on the anterior arch, the longest 2-16 in the eye-diameter.
Golden or golden brown above; sides with two series of large, round or
oval, golden spots, separated by broad blue interlacing bands; abdominal region
-and extremity of tail violet, with splashes of greenish gold. Head lilaceous, with
irregular violet spots and bars; a deep blue blotch, prolonged upwards as an
uneven band on the opercle; branchiostegal region blackish. Outer half of
spinous dorsal dark olive-green, narrowly bordered above with purple, below with
pale blue; the lower band is continued to the end of the soft dorsal, the outer half
of which is pale olive-green, with part of the membrane blue, as also is the base.
Anal blue, with a median and a basal series of golden spots. Caudal rays olive-
green or purple, the interradial membrane blue. Pectoral pale yellowish brown,
the base with one or two vertical blue bars. Ventral bluish black.
Described from five specimens, 224 to 305 mm. in total length, in the
collections of the Queensland Museum and the Amateur Fishermen’s Association.
These were taken on the Snapper Grounds off Moreton Bay, where it. does not
appear to be uncommon, and which is still the only recorded habitat of the
species. It is said to be good eating.
Our illustration is taken from a Queensland Museum example. Reg. No.
I. 17/2840.
2. MEROGYMNUS JACKSONIENSIS (Macleay).
‘Opisthognathus jacksoniensis Macleay, Proce. Linn. Soc. N. 8. Wales, v, 1881, p. 570; O’Connor,
Proce. Roy. Soc. Queensl., iv, 1888, p. 43.
Gnathypops jacksoniensis Waite, Rec. Austr. Mus., v, 1904, p. 240, pl. xxvi, fig. 2.
Merogymnus jacksoniensis McCulloch, Rec. West Austr. Mus.,
SOUTHERN SMILER.
Type locality :—Port Jackson, N.S.W.
Body elongate-elliptical, its width at the shoulder 1-6 to 1-75 in its depth,
which is 4-33 to 4-5 in its length and 1-4 to 1-55 in the length of the head.
Caudal peduncle three fourths deeper than long, its least depth 3-44 to 3-6 in the
length of the head. Head a little deeper than wide, the fronto-occipital profile
26 MEMOIRS OF THE QUEENSLAND MUSEUM.
linear or gently rounded and moderately acclivous, its width 1-5 to 1-67 in its
length, which is 3-12 to 3-33 in that of the body. Snout short, with feebly rounded
subvertical profile, its length 1-5 to 1-63 in the eye-diameter, which is 3-25 to 3-5.
in the length of the head. Interorbital region narrow and concave, its width 3-33
to 3-6 in the eye-diameter. Maxillary extending to about one diameter of the eye
behind the eye, its length 1-6 to 1-67 in that of the head, the width of its distal
extremity 3-16 to 3-5 in its length.
Seales extending forwards along the middle of the trunk to above the vent
only, the naked area extending backwards above and below in a gradually
contracting band to about the last third of the vertical fins; only those scales on
or near the caudal peduncle truly imbricate; belly naked. Lateral line termi-
nating below the 12th or 18th dorsal ray.
Dorsal fin with x 17 rays, originating well behind the tip of the maxillary ;
outer border of spinous dorsal rounded in front, linear behind, the spines
gradually increasing in length to the last, but much lower than the soft rays, the
12th or 13th of which is the longest, 2 to 2-33 in the length of the head. Caudal
fin short, 5-16 to 5-67 in the body-length. Anal with 1 16 rays, originating below
the 2nd or 3rd dorsal ray, the longest ray corresponding to and as long as those
of the dorsal. Pectoral rounded, with 20 rays, its length 1-83 to 2 in that of the
head. Ventral as long as or a little longer than the pectoral, the 2nd ray some-
what produced, but not nearly reaching to the vent.
Gill-rakers long and moderately slender, with the inner edge spinulose,
§ + 19 or 20 on the anterior arch, the longest 2-1 in the eye-diameter.
Ground-color varying from rich light brown to lemon yellow, the upper
surface and sides of the trunk broadly reticulated with chocolate brown, a wavy
median band, which is sometimes black, being always present; throat, breast, and
belly immaculate; tail with a few irregular brown spots and wavy lines, each
scale with a brown central dot. Upper surface and sides of head profusely
dotted and pencilled with brown; inner posterior edges of maxillary and
mandible black; distal extremity of maxillary and a broad stripe on the
mandible white. Dorsal fin dark purplish brown, with a basal series of lighter
spots; soft portion with an additional submarginal row of similar spots and two
oblique bands posteriorly. Anal rich brown, with a light basal band. Caudal
with three transverse rows of light spots. Pectoral pale yellow, profusely
powdered with brown, Outer portion of ventrals purplish black.
Described from four specimens, 212 to 264 mm. in total length, in the
collection of the Queensland Museum; they were taken by hook on the Snapper
Grounds off Moreton Bay, where, however, it appears to be scarce.
Originally described from Port Jackson in 1881, its range was extended,
eight years later, to the Queensland Coast, through a specimen obtained in
Moreton Bay by the late Mr. D. O’Connor; these two localities still form the:
extreme limits of its recorded distribution.
MEMOIRS OF THE QUEENSLAND MUSEUM.
Puate III.
QUEENSLAND FISHES.
‘(Avapvyy pue oudo[py) VIVTOOVIN SdOdAHLYNY)
‘jep ‘ayumi syrinud
Face page 27.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 27
Waite is certainly incorrect in stating that ‘‘the premaxilla is much
shorter (than the maxilla), and does not extend to below the hinder margin of
the eye.’’ As a fact it is as long as the maxillary, but its distal third is slender
and toothless, composed of flexible cartilage. Nor in those which I have examined
has the bony interorbital bridge ever been so narrow as described by him.?®
The name ‘‘leopard fish’’ applied to this species by Waite’® and Stead’ is
inappropriate, as it rightly belongs to Plectropoma maculatum Bloch, the
Holocentrus leopardus of Lacépéde.
i. GNATHYPOPS Gill.
Gnathypops Gill, Proce. Acad. Nat. Sci. Phila., 1862, p. 241 (mazillosa); Jordan & Evermann,
Fish. North & Mid. Amer., pt. 3, 1898, p. 2283; McCulloch, Rec. West Austr. Mus., i,
1914, p. 216.
Differs from Merogymnus in having the trunk wholly covered with scales,
the outer row of teeth enlarged, and the gill-rakers fewer, shorter, and stouter.
The most prolific genus of the family, containing 20 species, ranging from
the Persian Gulf through the Indian and Malayan seas northward to Japan and
eastward to New Guinea and East Australia. Crossing the Pacific it reappears
on the coasts of Lower California and Western Mexico, reaching thence to the
shores of the Southern United States, the West Indies, and Brazil. Three species
oceur in Australian waters, and have been thus classified by MeCulloch—
a’. Body and fins light, with numerous small, dark brown spots .. af .. 1. maculata.
a*, Body and fins dark, sometimes with irregular, scattered darker spots . . ..- 2. inornata.®
a’, Head with small, body with large, incomplete, brown rings; dorsal, anal, and caudal fins
with oblique bars, the former with a large, black spot anteriorly 3. darwiniensis.™
8. GNATHYPOPS MACULATA (Alleyne & Macleay).
(Plate ITI.)
Opisthognathus maculatus Alleyne & Macleay, Proc. Linn. Soe. N. 8. Wales, i, 1877, p. 280,.
pl. ix, fig. 3.
Batrachus punctatulus Ramsay, Proc. Linn. Soc. N. S. Wales, viii, 1883, p. 177.
Gnathypops maculatus McCulloch, Ree. West Austr. Mus., i, 1914, p. 216.
SPECKLED SMILER.
Type localities:—Palm Isles, N.Q. (O. maculatus).
Torres Strait, N.Q. (B. punctatulus).
Body elliptical, its width at the shoulder 1-84 in its depth, which is 3-1 in
its length and 1-2 in the length of the head. Caudal peduncle one and a half time
Rec. Austr. Mus., v, p. 240.
18 Synop. Fish. N. 8. Wales, p. 30, No, 231.
Fish, Austr., p. 108.
% Ramsay & Ogilby, Proc. Linn. Soc. N. S. Wales, xii, 1887, p. 561; Derby, N. W.
Australia. The species has been beautifully figured by McCulloch, Rec. West Austr. Mus., 1,
| WSye-o:0-<5
° ” Macleay, Proc. Linn. Soc. N. 8. Wales, ii, 1878, p. 355, pl. ix, fig. 3; Port Darwin, ND.
28 MEMOIRS OF THE QUEENSLAND MUSEUM.
deeper than long, its least depth one third of the length of the head. Head a
little wider than deeper, the fronto-occipital profile linear and strongly acclivous,
that of the nape rounded, its width 1-33 in its length, which is 2-55 in that of the
body. Snout short, with linear, subvertical profile, its length 1-25 in the eye-
diameter, which is 3-67 in the length of the head. Interorbital region narrow and
concave, its width one third of the eye-diameter. Maxillary extending rather
more than an eye-diameter behind the eye, its length 1-5 in that of the head, the
width of its distal extremity 3-9 in its length.
Seales everywhere imbricate, disposed in regular series over the whole
body, becoming somewhat larger posteriorly. Lateral line terminating below the
9th dorsal ray.
Dorsal fin with xii 16 rays, originating well behind the tip of the
maxillary; outer border of spinous dorsal rounded in front, linear behind, the
spines gradually increasing in length to the last, but lower than the soft rays,
the 8th, 9th, and 10th of which are the longest, a little more than half the length
of the head. Caudal fin of moderate length, 4-6 in the length of the body. Anal
with ii 14 rays, originating below the 3rd dorsal ray, the rays increasing in length
to the 4th, which is 2-37 in the length of the head and as long as the succeeding
seven, beyond which they decrease. Pectoral rounded, with 21 rays, all except the
upper and lower pair about equal in length, and 2-25 in the length of the head.
Ventral longer than the pectoral, none of the rays produced.
Gill-rakers 4-+-12 and some tubercles, rather stout and cultriform, the
longest 2-3 in the eye-diameter,
Pale rufous brown, scarcely lighter below, everywhere spotted with dark
blue or black, the spots largest on the upper part of the trunk and tail, smallest
on the head; a large blackish or smoky brown blotch beneath the appressed
pectoral. Spinous dorsal sparsely, pectorals profusely spotted; the other fins
immaculate.
Described from a fine specimen, measuring 346 mm. in total length,
collected at Dobo, Aru Islands, by Mr. John Colclough, who presented it to the
Amateur Fishermen’s Association of Queensland, by whom it was kindly lent for
the purposes of this review.
This fine species was first described from a small example, 178 mm. long,
obtained at the Palm Isles, N.Q., during the cruise of the ‘‘Chevert.’’ We next
hear of it from Torres Strait, through a specimen forwarded to the Australian
Museum by Mr. Cousens, and described as a Batrachus by the late Dr. E, Pierson
Ramsay. A few years later Saville Kent, when a guest on board H.M.S.
‘‘Myrmidon,”’ collected one at Port Darwin, N.T., as recorded by de Vis. From
that time I find no record of the species until 1906, when Mr. Banfield sent me a
320 mm. example from Dunk Island, while in the following year the specimen
described and figured came into my hands. It is said to be good eating.
EDIBLE FISHES OF QUEENSLAND.—OGILBY. 29.
LIisT OF THE OPISTHOGNATHOID FISHES OF THE INDIAN AND
WESTERN PACIFIC OCEANS.
. OPISTHOGNATHUS Cuvier & Valenciennes, Hist. Nat. Poiss., xi, 1836,,
p. 498 (sonneratu = mgromarginatus).
1. nigromarginatus Ruppell, Atlas Fisch. Roth. Meer., 1828, p. 114.
Distribution :—Red Sea: and Mozambique Channel through the Seas of
India and Malaysia to Celebes and Batchian.
Dimensions :—To 180 mm..
Figure :—Day, Fish. India, pl. lvii, fig. 5.
i. MEROGYMNUS Ogilby; see p. 24.
. eximius Ogilby ; see p. 24.
bo
. jacksomensis Macleay ; see p. 25.
fe OOo
. yoms Jordan & Thompson, Proc. U. 8. Nat. Mus., xlvi, 1918, p. 65. Only
known from the type.
Distribution :—lyo, Shikoku, Japan.
Dimensions :—74 mm.
Figure :—Jordan & Thompson, ibid., text-fig. 1.
iil. GNATHYPOPS Gill; see p. 27.
. darwiniensis Macleay, Proc. Linn. Soc. N. 8. Wales, ii, 1878, p. 355.
Distribution :—Port Darwin, N.T. Only known from the type.
Figure :—Macleay, ibid., pl. ix., fig. 3.
6. dendritica Jordan & Richardson, Bull. U. 8. Bur. Fisher., xxvii, 1908, p. 261.
Only known from the type.
Distribution :—Cuyo Island, Philippine Archipelago.
Dimensions :—133 mm.
Figure :—Jordan & Richardson, ibid., text-fig. 9.
This species should be carefully compared with the preceding.
On
7. evermanm Jordan & Snyder, Proc. U. 8. Nat. Mus., xxiv, 1902, p. 493..
Only known from the typical specimens.
Distribution :—Wakanoura, Japan.
Dimensions :—80 mm.
Figure :—Jordan & Snyder, ibid., text-fig. 6.
8. hopkinst Jordan & Snyder, ibid., p. 492.
Distribution :—Misaki and Sagami, Hondo, Japan.
Dimensions :—To 80 mm.
Figure :—Jordan & Snyder, ibid., text-fig. 5.
30 MEMOIRS OF THE QUEENSLAND MUSEUM.
9. inornata Ramsay & Ogilby, Proc. Linn. Soe. N. 8. Wales, xii, 1887, p. 561.
Distribution:—Coast of North-Western Australia (Derby and Port
Headland).
Dimensions :—To 485 mm.
Figure :—McCulloch, Ree. West Austr. Mus., i, pl. xxx.
macrolepis Peters, Mon. Akad. Berlin, 1866, p. 520.
Distribution :—Bangkok, Siam, to the Philippines.
Dimensions :—To 100 mm.
Unfigured.
11. maculata Alleyne & Macleay ; see p. 27.
—
10.
12. muscatensis Boulenger, Proc. Zool. Soc., 1887, p. 662.
Distribution :—Persian Gulf at Maskat, Arabia.
Dimensions :—To 305 mm.
Figure :—Boulenger, ibid., pl. liv, fig. 1.
13. papuensis Bleeker, Versl. Akad. Amst. (2) 11, 1868, p. 33.
Distribution :—Waigiu.
Unfigured.
14. rosenbergii Bleeker, Nat. Tijds. Nederl. Ind., xii, 1857, p. 220.
Distribution :—East Coast of India to Nias.
Dimensions :—To 120 mm.
Figure :—Day, Fish. India, pl. lviii, fig. 5.
15. solorensis Bleeker, ibid., v, 1853, p. 81.
Distribution :—Goram, Amboina, Solor.
Dimensions: To 83 mm.
Unfigured.
16. versluysi Weber, Siboga Fische, 1913, p. 261. Only known from the type.
Distribution :—Roma Island.
Dimensions :—136 mm.
Figure :—Weber, ibid., text-fig. 65.
iv. STALIX Jordan & Snyder, Proc. U. S. Nat. Mus., xxiv, 1902, p. 495 (histrio).
17. histrio Jordan & Snyder, ibid. Only known from the type.
Distribution :—Bay of Nagasaki, Japan.
Dimensions :—62 mm.
Figure :—Jordan & Snyder, ibid., text-fig. 7.
v. OWSTONIA Tanaka, Journ. Coll. Sci. Tokyo, xxiii, 1908, p. 47 (totomiensis).
18. totomiensis ‘Tanaka, ibid.
Distribution :—Off the Province of Totomi, Japan. Only known from the
type. :
Dimensions :—500 mm.
Figure :—Tanaka, Fish. Japan, pl. viii, fig. 26.
NOTES ON’ QUEENSLAND FLIES.—JOHNSTON AND BANCROFT. 31
NOTES..ON THE BIOLOGY.,.OF. SOME
QUEENSLAND FLIES.
By ProFessor T. HARVEY Jounston, M.A., D.Sc., Hon. ZooLocist, QUEENSLAND
Museum; AND M. J. Bancroft, B.Sc., WALTER AND ELizA HALL FELLOW IN
Economic BroLocy, UNIVERSITY, BRISBANE.
(With 48 Text-figures.)
During our experimental work with Diptera as possible transmitting
agents of certain worm parasites of horses and stock! we used a number of
species of flies found associated with these animals, some of them being either
insufficiently described or apparently undescribed. We have already dealt
with a few of these flies (Johnston and Bancroft, 1919) and now propose to
give an account of certain others which, as far as available literature allows
us to judge, belong to undescribed species of Musca.?
We desire to express our appreciation of the assistance given by Dr. E. W.
Ferguson, Health Department, Sydney; Mr. W. W. Froggatt, Government
Entomologist, Sydney; Dr. A, B. Walkom, Linnean Society, Sydney; and
Mr. W. A. Rainbow, Australian Museum, Sydney. Typical specimens have
been deposited in the Queensland Museum, Brisbane, and in the Australian
Museum, Sydney.
MUSCA TERRZ-REGINZ n. sp.
(Text-figs. 1, 2, 9, 10, 15, 17, 18, 27, 28, 31, 37, 38, 41, 42, 43, 44, 45.)
This is a rather small fly about 4-5 mm. in length, which has been taken
occasionally on stock in the Hidsvold district and is referred to in our previous
paper (1919, p. 182) as Musca sp. indet.
MALE.
Head (fig. 9)—One is struck with the relatively enormous size of the
eyes which occupy more than three quarters of the front of the head. They are
separated from one another by a narrow frontal stripe bounded by numerous
1 Johnston and Bancroft, The Life History of Habronema in relation to Musca domestica
and native flies in Queensland, P.R.S. Q’land, 1920.
Johnston and Bancroft, Experiments with certain Diptera as possible transmitters of
bovine Onchocerciasis, P.R.S. Q’land, 1920.
? Unfortunately for us Professor M. Bezzi’s excellent key deals only with African and
Mediterranean species of Musca (Miodarii Superiori raeccolti dal Sign C. W. Howard nell
Africa australe orientale—Boll. Lab. zool. gen. agr. d. R. Scuola Sup. d’Agrie. Portici, 6,
1912, Keys pp. 85-88).
32 MEMOIRS OF THE QUEENSLAND MUSEUM.
rather short chete. The ocellar triangle is not very prominent and bears.
about four pairs of short ocellar bristles. Behind the triangle there is
on each side a prominent vertical bristle, with a shorter one adjacent to it.
The facial region is practically triangular in outline. The silvery parafrontals
do not extend upwards between the eyes except as a thin black line on either
side between the frontal stripe and each eye. The characters of the silvery-
grey antenne are shewn in fig. 31.
Text-figs. 1-4-—Camera lucida drawings to show dorsal colouration. All figs. drawn:
to same scale. Fig. 1, Musca terre-regine male; 2, M. terre-regine female; 3, M. hilli male;
4, M. hilli female.
Thorax (fig. 1)—When viewed with the light falling from the front
the thorax is black and shiny, but when viewed from the opposite quarter the
general effect is dark greyish.* There are four longitudinal stripes, the two
outer extending backwards to the sides of the scutellum whilst the two inner
extend only to the middle of the scutum. The central silver-grey stripe is
ratuer broader and in individuals of either sex may extend much further
forward than the others. The black stripes of each side fuse anteriorly. The
posterior half of the midregion of the scutum is rather darker than the other
portions. On the anterior part of the scutellum there is a prominent black
marking followed by a well-defined smoky patch extending to the hind margin
of the scutellum. The chetotaxy is shewn in fig. 15.
* The ‘accompanying descriptions of the thorax and abdomen of each species are based
on specimens with the head towards the observer and with the source of light opposite the:
observer,
NOTES ON QUEENSLAND FLIES.—JOHNSTON AND BANCROFT. 33
Wings.—The veins are yellowish brown. The venation is shewn in fig. 17.
The squame is pale yellow.
Legs.—The tarsus of the third leg is figured in fig. 27.
Abdomen.—The abdomen is pale yellow with a median dorsal band, the
intensity of whose colouration is interrupted in places so that there appears a
succession of light-brown and dark-brown markings. The colouration extends
to the anterior half of the first abdominal segment? where it is rather dark
brown. Laterally on each segment there is a patch of darker yellow (fig. 1).
FEMALE.
Head.—The main external difference between the male and female is in
the size of the eyes which in the female are separated (where they most closely
approximate) by about a third of the width of the head (fig. 10), whereas the
minimum distance is about one twenty-fifth in the male. The silvery para-
frontals extend upwards between the eyes as a fairly broad band separated by
Text-figs. 5-8—Camera lucida outlines of face views drawn to same scale to show
relationship of eyes, frons, ete. Fig. 5, M. australis male; 6, M. australis female; 7,
M. vetustissima male; 8, M. vetustissima female; 9, M. terre-regine male; 10, M. terre-regine
female; 11, M. hilli male; 12, M. hilli female. Lettering as in preceding figures—ant.,
antenna; e., eye; fr., frons or frontal stripe; fr.b., frontal bristles; fc., facialia; 0.b., ocellar
bristle; o.t., ocellar triangle; p.fc., parafacialia; p.fr., parafrontalia; v.b., vertical bristles.
the very shallow smoky-coloured frontal stripe. There are two prominent
vertical bristles on either side behind the eyes. The ocellar triangle does not
project and the ocellar bristles are short.
*The terms Ist to 4th are applied to thé obvious abdominal segments—i.e., the reduced
first segment is not taken into account as a separate one but is considered as a part of the
second with which it is fused, the two together being dealt with as the first segment.
Cc
34 MEMOIRS OF THE QUEENSLAND MUSEUM.
Thorax.—The thorax is similar to that of the male (fig. 2). The form
of the last tarsus is shown in fig. 28.
Wings—As in the case of M. hilli there are slight differences in the
sexes in regard to the wing, particularly in the region of the first posterior
cell and the posterior cross-vein (fig. 18).
Abdomen (fig. 2).—The general colour is yellow with a brown median
dorsal stripe, rather broad on the anterior portion of the first segment and
more or less interrupted in the middle portion, but on the posterior region of
that segment the colouration makes its reappearance extending on to the
succeeding segment and widening to form a large dark-brown blotch of more
or less triangulsr outline. A similar patch of colour occurs on the succeeding
segment and is continued on to the fourth. Faint shimmering whitish patches
occur laterally towards the posterior end of the abdomen. The under surface
is yellow.
Text-fig. 13, Outlines of thorax (dorsal) to show chetotaxy; M: australis; 14
M. vetustissima; 15, M. terre-regine; 16, M. hilli. Lettering—a., acrostichal (inner dorso-
central); d.c., dorsocentral; hu., humeral; i.a., intra-alar; npl., notopleural; p.a., postalar;
ph: posthumeral; pr., presutural; p.s.d.c., presutural dorsocentral; s.a., supra-alar; sc.,
scutellar.
2
Breeding Habits—The eggs are laid in cowdung or horsedung, each egg
measuring about 1 mm. in length by -25 mm. in breadth, being very similar to
that of the house-fly. In less than twenty-four hours the eggs hatch, the larve
being in the first instar (fig. 42). Twenty-four hours later the first moult
occurs; the anterior spiracles each with six or seven processes appear (fig. 45),
while the posterior spiracles are in the form of two almost straight slits (fig. 43).
The second instar also lasts about twenty-four hours. In the third instar the
posterior spiracles take the form of three sinuous slits surrounded by a black
chitinous D-shaped ring (fig. 44). The mature larve are creamy white and
measure 9 to 10 mm. in length by 1-5 mm. in breadth. They leave the dung
NOTES ON QUEENSLAND FLIES —JOHNSTON AND BANCROFT. 35
and pupate in damp sand under laboratory conditions. The puparium is
reddish brown and measures 4 to 5 mm. by 1-5 to 2mm. The larval stage lasts
from 5 to 7 days, and the pupal from 7 to 10 days, the total thus being from 12 to
17 days during midsummer (Eidsvold, Burnett River), which is similar to that
-of the house-fly.
In bred flies the males were usually slightly in excess of the females,
about 51 per cent. being males and 49 per cent. females. When bred flies of both
‘sexes were kept together in a small cage and fed on raisins, honey, &¢., copulation
was observed to take place in from 5 to 9 days after emergence (midsummer
observations), while the females were ready to oviposit about five days later.
The female genitalia (fig. 41) closely resemble those of the house-fly,
-except that the number of ovarian follicles is less, the maximum number
detected in each ovary being thirty-five.
Parasites —M. terre-regine was found to be parasitised by a flagellate,
Herpetomonas (probably H. musce-domestice Burnett), and by three larval
nematodes (Habronema musce Carter, H. megastoma Rud, and AGOMSRE ETE
muscarum Jnstn. and Baner.).
The following observation may be of interest as it allows one to compare
‘the various periods taken by the three flies M. terre-regine, M. vetustissima,
and M. domestica to pass through their larval stages in horsedung, since all
‘were under similar conditions, being bred in material from the same source and
collected at the same time (November, 1919, Eidsvold) :-—
— Musca terre-regine. M. vetustissima. M. domestica.
Eggs deposited November 13th .. B 3 .. | 14th “ .. | 14th
Hatched as first Sens November 14th ~, at recog) LLL 8 er) both
Second instar .. .. | November 15th ay .- | 15th and 16th .. 146th:
Thirdinstar .. .. | November 16th, 17th Bi 17th ~ ay eleabe
Mature larva .. .. | November 18th, 19th, and 20th’ 18th and 19th .. | 20th and 21st
Larval stages .. .. | 5 to 7 days oy . | 4 to 5 days .. | 6 to 7 days
Emergence—
25 AGE Aer: es LS 2 males, 2 females
26.11.9 Ed 2 males, 5 females
Dr lls) 2 males, 3 females ae -.. |9 males, 2 females
28.11.9 Few—both sexes at i zt a
29.11.9 Few—both sexes a Aas A A few
SORE SOK oS. .. | 1 male, several females .. ie Aa A few
Pupal stage .. i 8 to 1l days ae or 7. | 7 to 9 days .. | 9 to 10 days
‘Total period for larval 13 to 18 days re Age -» | L1 to.l4 days .:.| 15 to l7 days
stages (egg to imago)
approximately
MUSCA HILLI n. sp.
(Text-figs. 3, 4, 11, 12, 16, 19, 20, 29, 30, 32, 39, 40, 46, 47, 48.)
This is an outdoor species which is occasionally found on horses and
cattle in Hidsvold and also in Brisbane. We have also seen it at Tweed Heads,
N. S. Wales (March, 1920). The average length of full-grown specimens of
either sex is about 6 mm.
36 MEMOIRS OF THE QUEENSLAND MUSEUM.
MALE.
Head.—The eyes occupy the greater part of the front view and are
contiguous for a considerable distance (fig. 11), separated only by a very narrow
frontal stripe. The ocellar triangle is elongate and narrow, projects prominently,.
and bears about four pairs of ocellar bristles. The parafrontals are wide and.
silvery. The frontal stripe is narrow and brownish.
The antenne are smoky in colour. The form of the arista is shewn in.
fig. 32. There are two prominent bristles, also a shorter third one, on the second
joint of the antenna.
Thorax (fig. 3).—When lighted from the anterior end the general effect is.
black and shiny, but when viewed from the opposite quarter four distinct stripes.
are to be seen separated by narrow greyish zones. The outer stripes extend
from the anterior portion of the thorax to the end of the scutum and on to
the anterior corners of the scutellum. The two inner stripes extend from the
anterior end of the prescutum to about the midregion of the seutum where
they merge into a rather smoky colouration occupying the middle and posterior
20
| am
Text-fig. 17, Wings drawn to same scale; M. terre-regine male; 18, M. terre-regine
female; 19, M. hilli male; 20, H. hilli female.
part of the seutum between the longitudinal stripes. The four stripes are of
approximately equal width and are each rather narrower than the central
silvery stripe. The two dark stripes of each side are joined up by a black
band anteriorly, somewhat as in M. ferguson. The posterior end of the seutellum
is smoky, this colouration becoming intensified so that at the anterior end the
scutellum is practically black. The chetotaxy is indicated in fig. 16. It might
be mentioned that the scutellum is very hairy, the sete being particularly well
developed and the macrochietz quite long,
Wings.—The wings are clear, the veins yellowish brown. The venation.
NOTES ON QUEENSLAND FLIES—JOHNSTON AND BANCROFT. 37
4s indicated in fig. 19. To the naked eye the squame appears opaque and some-
what pearly. ;
Legs—The characters of the tarsi of the third leg are shewn in fig. 29.
Abdomen (fig. 3).—The fly can be very readily distinguished from
M. vetustissima and M. ferguson by the pale-yellow abdomen in both sexes.
There is dorsally a median narrow brownish stripe, interrupted in the middle
of the first abdominal segment. It is more pronounced in some individuals
than in others, being occasionally reduced to a brownish blotch on the posterior
edge of each segment. Whitish reflections are visible posteriorly and postero-
laterally. The abdomen is well provided with long chetz. The ventral surface
is pale yellow with brownish colouration in the vicinity of the genital aperture.
FEMALE.
Head (fig. 12).—The head of the female differs from that of the male
in the following particulars :—The eyes are widely separated, the distance between
them, where they approach most nearly, being about a third of the total width
25
23 ote
o>
ees a7
ian
28
\y/
Text-fig. 21, Tarsi of third leg—final segments only—all drawn to same seale,
M. australis male; 22, M. australis female; 23, M. vetustissima male; 24, M. vetustissima
female; 25, M. demestica male (for comparison); 26, M. domestica female; 27, M. terre-
regine male; 28, M. terre-regine female; 29, M. hilli male; 30, M. hilli female.
of the head, whereas in the case of the male it is about one thirtieth. There is
a broad shallow dusky frontal stripe bounded by strongly ineurving fronto-
orbital bristles. On each side of the stripe is a fairly wide parafrontal which,
as in the case of the male, is silvery.
Thorax (fig. 4)—The female thorax has much the same characters as
the male.
Wing.—The wing (fig. 20) differs from that of the male in the shape of
the first posterior cell (fifth radial) and also in the form of the elbow of the
fourth longitudinal vein (M. 1 + 2) and its relation to the posterior cross-vein
(mediocubital).
38 MEMOIRS OF THE QUEENSLAND MUSEUM.
Abdomen (fig. 4) —When viewed from certain points the abdomen appears.
to possess shimmering white patches, this being of course due to the arrange-
ment of the tiny hairs. Ventrally the colour is pale yellow, there being only a
faint brownish tint around the genital opening.
Breeding Habits—The breeding habits of this fly resemble those of the
preceding species, the small white eggs, which measure approximately the same
as those of M. terrw regine, being laid in cowdung or horsedung. The successive
instars occupy about the same periods. The posterior spiracles of the second.
and third instars are quite typical (figs. 46, 47). The anterior spiracles contain
from 4 to 7 processes (fig. 48). The mature larva measures from 10 to 12 mm.
The larval stage occupies from 5 to 6 days, the pupal 6 to 9 days—a total of 11
to 15 days (January and February, Eidsvold and Brisbane). The puparium
is red-brown to deep chocolate and measures about 6 mm. in length by 2-7 mm.
in breadth. The female genitalia are very like those of the preceding species:
(fig. 41), the maximum number of follicles observed in each ovary being, how-
ever, slightly larger, viz., about 40.
s
oe
—— ae
Smm
Text-fig. 31, Ariste of females, WZ. terre-regine ; 32, M. hilli.
Text-fig. 32, Palps (all to same scale) drawn from mounted specimens, M. australis
male (fig. 33) and female (34); M. vetustissima male (fig. 35) and female (fig. 36) ;
M. terre-regine male (fig. 37) and female (fig. 38); M. hilli male (fig. 39) and female:
(fig. 40).
The fly can be readily distinguished from Musca pumila by the coloura-
tion of its abdomen (Macquart, Dipt. Exot. Suppl. 3, 1847, p. 58). It appears to:
be more closely related to Musca minor (Macquart, l.c. Suppl. 4, 1850, p. 253):
NOTES ON QUEENSLAND FLIES—JOHNSTON AND BANCROFT. 39
Text-fig. 41, Genitalia of female M. terre-regine. Lettering: ac.g., accessory gland;
€.g.g., conglobate gland; n.c., nurse cells; o., ovum; ad., oviduct; od.c., common oviduct;
0.f., ovarian follicles; ov., ovary; sp., spermatheca,
40 MEMOIRS OF THE QUEENSLAND MUSEUM.
Text-figs. 42-44, Larval spiracles, all drawn to same scale. Posterior spiracles of
M. terre-regine ; first instar (42), second instar (43), third imstar (44).
Text-fig. 45, Anterior spiracle of third instar of VW. terre-regine.
Text-figs. 46-48, Spiracles of larva of M. hilli: fig. 46, posterior spiracle of second
instar; 47, posterior spiracle of third instar; 48, anterior spiracle of third instar. a.sp., anterior
spiracular process, surrounded by a clear envelope (c.e.).
from Tasmania, but in view of the brevity of the description, particularly in
regard to the thoracic and abdominal markings, we do not feel justified in
identifying our form with his. We have pleasure in associating with this new
species the name of Mr. G. F. Hill, Entomologist to the Australian Institute of
Tropical Medicine, Townsville, North Queensland, who has published important
papers dealing with Australian flies.
NOTES ON QUEENSLAND FLIES.—JOHNSTON AND BANCROFT. 41
Parasites —M. hilli readily becomes parasitised by .the nematodes
Habronema musce and H. megastoma,
EUMUSCA VETUSTISSIMA (Walker).
(Text-figs. 7, 8, 14, 23, 24, 35, 36.)
Townsend (1915) stated that the type of the genus Musca—so designated
by Latreille—is MW. vomitoria L., a blowfly which was transferred to Calliphora;
and that the latter generic name fell into synonymy, a new name being required
for the species usually placed under Musca. He accordingly erected the genus
Promusca (p. 484) with M. domestica L. as its genotype. He had previously
(1911, p. 170) separated off certain species making M. corvina Fabr. the type of
Eumusca.
The characters of Musca vetustissima, which have already been described
by us (J. and B., 1919), agree with most of those given by Townsend (1915,
p. 435) but differ in certain important respects—e.g., (1) the eggs, though
macrotype, are not stalked; (2) the puparium is red-brown like that of the
house-fly; and (3) copulatory vesicles are present in the female. The species
thus occupies a position between Promusca and Eumusca, more nearly approach-
ing the latter genus of which it may perhaps be regarded as an atypical
member, otherwise a new genus would need to be erected for its reception.
In our previous account we did not refer to the chetotaxy which is now
figured (fig. 14). There are two weak dorsocentrals on either side in front
of the suture. The presutural and posthumeral are rather strongly developed.
Postsuturally there are four dorsocentrals, only the last being strong. Three
pairs of scutellar macrochetx are present, two pairs being well developed. The
vertical bristles behind the eyes are arranged as in Musca domestica.
The palp of the two sexes are shown in figs. 35 and 36 and the tarsus of
the third leg in figs. 23 and 24.
In our earlier account (1919, fig. 5) we omitted to indicate in our figure
that the fourth abdominal segment (7.c. true fifth) of the male possessed a dark
colouration.
Dr. Cumpston, Director of the Federal Quarantine Service, informed us
that he had observed the common Western Australian bush-fly (JZ. vetustissima)
copulating thirty-six hours after emergence. He used a rather large cage
measuring about 6 by 6 by 8 feet for his observations.
VIVIPAROMUSCA FERGUSONI J. and B.
(Text-figs. 5, 6, 13, 21, 22, 33, 34.)
This species was described last year (J. and B.) as Musca ferguson.
If Townsend’s genus Viviparomusca (1915, p. 435) be recognised then the rela-
tionships of this Northern Australian fly are rather with it than with Promusca
42 MEMOIRS OF THE QUEENSLAND MUSEUM.
(Musca of authors). It differs, however, in several important points, viz—(1)
the larva is carried in the uterus to the second stage, not the third; (2) eopula-
tory vesicles are present in addition to the accessory glands in the female. In
regard to (1) we might point out that the two posterior spiracular slits of this:
instar are not straight but sinuous as they are in those of the third instar where
there are three. Then again in the type species of the genus M. bezzew Patton,
‘a larva is deposited, though in which stage mention is not made by Patton and
Cragg (1913, p. 23). In M. fergusont the larva is deposited while in an eggshell
which at once bursts liberating it.
This fly appears to be very hairy on account of the presence of well-
developed macrochetz on the dorsal part of the thorax. Many of the seutellar
bristles are very strong and are practically macrochete. The arrangement is:
shown in fig. 13. The vertical pairs between the eyes are strong. The ocellar
bristles are weakly developed in both sexes, especially in the male where they
are practically absent.
The form of the tarsi and claws of the third leg in the two sexes is shown
in figs. 21 and 22. The palps bear very strong sete in both sexes, especially in.
the male (figs. 33, 34).
OTHER FLIES.
In our earher paper (1919, p. 182) we referred to the presence of certain
other flies in the Eidsvold district. The cobalt-blue species therein indicated as:
Lasiopyrellia is a Pseudopyrellia, according to Williston’s Key to the North
American genera (1908), while the very small fly which was regarded as being
probably a Pseudopyrellia falls within the genus Pyrellia. It was mentioned
that both of these metallic flies frequented cowdung for oviposition. So also:
does a rather handsome large blowfly which Mr. W. W. Froggatt has identified
for us as Stenopterina gigas Macq. A tiny species of Sepsis is very commonly
found frequenting similar material and breeds in it. Patton and Cragg (19136,.
p. 310) refer to its presence in India.
Muscina stabulans has been bred out of rotting potatoes received at
Eidsvold from elsewhere and no doubt the fly is occasionally to be met with
in the township.
Amongst the Anthomyide occurring there one might mention that
Phaoma personata Walker, which at fivst sight somewhat resembles a large
greyish or bluish long-winged house-fly, is to be found in the vicinity of rotting
fruit and has been bred out from that material.2 Mr. Froggatt (1907, p. 311)
who kindly identified the species and also the one next mentioned, informed us
that it occurred commonly in fruit-shops in Sydney. Sapromyza fuscicorms:
Macq., a fairly large fly (Sapromyzide) with a striped thorax and a dark head,
has been bred from horsedung (Hidsvold).
°See also W. B. Gurney. Fruit-flies and other insects attacking cultivated and wild!
fruits in N. 8. Wales—Farmers’ Bulletin, No. 55, July, 1912, Dept. Agric. N. S. Wales, p. 29;
and in Agric. Gaz. N. 8. Wales, January 1912 (pp. 75-80).
NOTES ON QUEENSLAND FLIES—JOHNSTON AND- BANCROFT, 43.
LITERATURE QUOTED.
1907. Froceart, W. W. Australian Insects, Sydney, 1907.
1919. JounNstTon, T. H., and Bancrorr, M. J. The life histories of Musca australis and
M. vetustissima, P.R.S., Q’land, 31, 1919, pp. 181-203.
1913. Parron, W. S., and Crace, F. W. Hematophagous species of the genus Musca. Ind.
Jour. Med. Res. 1 (1), 1913, pp. 11-25.
1913b. Parton, W. S., and Crace, F. W. A textbook of Medical Entomology.
1911. Townsend, C. H. Review of work by Pantel and Portschinsky on reproductive and
early stage characters of Muscoid flies. Proc, Ent. Soc. Washington, 13, 1911,
pp. 151-170.
1915. TownsEnD, C. H. Correction of the misuse of the generic name Musca, with a
description of two new genera. Jour. Wash. Acad. Sci. 5 (12), 1915, pp. 433-6.
1908. Wituiiston, 8S. W. Manual of North American Diptera, edit. 3, London.
44 MEMOIRS OF THE QUEENSLAND MUSEUM.
THE OCCURRENCE IN BRISBANE RIVER OF
THE NEW ZEALAND AMPHIPOD, PARA-
COROPHIUM EXCAVATUM (G. M. THOM-
SON).
By CuHas. Cuitton, M.A., D.Sc., M.B., C.M., LL.D., F.L.S., C.M-Z.S) iene eiieer
Roy. Soc. N.S.W., PROFESSOR OF BIOLOGY, CANTERBURY COLLEGE, NEW ZEALAND.
(With Text-figures Nos. I to XIX.)
Tow Arps the end of 1918, I received from Professor T. Harvey Johnston,
of the University of Queensland, and honorary Zoologist to the Queensland
Museum, a few Crustacea from Brisbane River, sent chiefly because of the
boring Isopod, Sphwroma terebrans Bate, which was doing considerable destruc-
tion in timber immersed in the water. Among the specimens, however, were
numerous examples of a small amphipod evidently belonging to Corophium
or some allied genus. The male of this species first attracted my attention because
of the character of the second gnathopod and the possession of a lobe on the
inner side of the end of the penultimate joint of the peduncle of the lower
antenna. Later on when I came to examine the females, which differed in the
structure of the second gnathopod and in having no lobe on the antenna, I was
struck by their resemblance to the deseriptions of Paracorophium excavatum
(G. M. Thomson), an amphipod found in brackish waters of New Zealand. In
that species no sexual differences had hitherto been described. However, I found
on closely examining specimens in my eollection that there were some males
among them having the characters of the second gnathopod and the lobe on the
lower antenna quite similar to those of the Brisbane specimens, and careful
comparison shows conclusively that the Brisbane specimens belong to the same
species as the New Zealand. I take this opportunity of giving a fuller account
of the species than has hitherto been published and of describing the differences
between the male and the female.
Paracorophium excavatum was described by Mr. G. M. Thomson in 1884,
under the name Corophium excavatum, from specimens obtained in Brighton
Creek, near Dunedin, the water of which he described as being salt. A little later
I received some specimens from Napier, and in January, 1890, I collected a
number at Brighton itself at a time when the water in the estuary was nearly
fresh or only slightly brackish; in 1894 I obtained a few specimens from Nelson,
also in brackish water. In 1902 Messrs. Lucas! and Hodgkin obtained specimens
* Keith Lucas, A Bathymetrical Survey of the Lakes of New Zealand, Geographical
Journal for May. and June, 1904.
PARACOROPHIUM EXCAVATUM.—CHILTON. 45
from Lake Roto-iti (fresh water) in Auckland which I afterwards identified as:
belonging to the same species. The Brisbane specimens now to be described
also come from brackish water, and the occurrence of this species, which is
apparently confined to brackish and fresh waters, in Australia as well as in
New Zealand, is of some importance in connection with the general question of
the geographical distribution of New Zealand and Australian Amphipoda.*
In 1899 Stebbing established the genus Paracorophiwm for the reception.
of the species now under consideration, which is the only one of the genus at.
present known.
PARACOROPHIUM Stebbing, 1899.
Paracorophium Stebbing, 1899, Ann. Nat. Hist., ser. 7, vol. 3, p. 350; 1906, Das Tierreich>
Amphipoda, p. 663.
In general appearance and in most of the appendages Paracorophium
comes close to Corophium, the chief points of difference being—(1) The
mandible has the palp well developed and three-jointed, (2) the second
gnathopod of the male is markedly different from that of the female and in
both sexes has the merus produced into a scoop-like process different from that.
in Corophium, (3) the third uropoda are two-branched.
PARACOROPHIUM EXCAVATUM (G. M. Thomson).
Corophium excavatum G. M. Thomson, 1884, Trans. N. Z. Inst., vol. 16, p. 236, pl. 12, fig. 1 to 8).
Paracorophium excavatum, Stebbing, 1906, Das Tierreich Amphipoda, p. 664.
Paracorophium excavatum, Chilton, 1906, P.Z.S., p. 704.
As this is the only species of the genus it is unnecessary to give a separate-
specific diagnosis. Thomson’s original description and figures, on which that.
in Das Tierreich Amphipoda appears to be based, apply to an immature male..
A detailed description is given below.
Size—About 4 mm. in length.
Colouwr.—Greyish.
Localities—New Zealand—Brighton, Napier, Nelson (brackish water),.
and Lake Roto-iti (fresh water) ; Australia—Brisbane River (brackish).
Antenna 1 (fig. 1) has the second joint of the peduncle longer than the
first but much more slender; the third about half the length of the second;
* P.Z.S. 1906, p. 704.
* Several of the Corophiide appear to be able to live in brackish or fresh water. Speaking
of the whole family Stebbing says (1906, p. 662), ‘‘ Marine, but extending into brackish or even.
almost fresh water’’; and of Corophiwm crassicorne Bruz. he says, ‘‘Found in Norfolk in
almost fresh water.’’ C. volutator is recorded as ‘‘forming tubular galleries in the mud of
tidal swamps.’’ I have specimens of this species obtained for me by Mrs. Sexton, of Plymouth,
labelled ‘‘Mouth of Issel, near Kampen. Quite fresh water’’; and Mr. Robert Gurney,
speaking of this species under the name ‘‘C. grossipes (Linn.),’’ says ‘‘it seems to thrive-
well in fresh water’’ (Trans. Norfolk and Norwich Naturalists’ Soe., vol. viii, p. 435, 1907).
Later on he recorded it as found in the Oued Tindja at its outflow from Lake Garaa Achkel.
in Tunisia (Jour. Roy. Mier. Soc. 1909, p. 283).
46 MEMOIRS OF THE QUEENSLAND MUSEUM.
flagellum shorter than the peduncle, containing about 10 joints. This appendage
appears to be the same in the male and in the female.
sabes stale bag \
Fig. 1—First antenna of male (Brisbane specimen).
Fig. 2.—Second antenna of male (Brisbane specimen).
Antenna 2 appears to vary considerably according to age and development,
becoming stouter, especially at the base, in older specimens. In a fully developed
male (fig. 2) the first three basal joints of the peduncle are short and very broad ;
the fourth joint is fully as long as all the preceding though much more slender,
and is produced at the apex on the inner side into an oval lobe which when
fully developed reaches nearly halfway along the fifth joint of the peduncle; the
fifth is only half as long as the preceding; the flagellum is shorter than the last
joint of the peduncle and consists of about 5 or 6 joints. The lobe at the end
of the fourth joint of the pedunele varies in-size in different specimens according
to their development; in one specimen from Brighton, the second gnathopod of
which is shown in figure 10a, the antennal lobe is quite short, the basal joints of
the peduncle are not so broad and the fourth joint scarcely so long as in more
mature specimens (fig. 2a). In a specimen from Lake Roto-iti which, judging
by the second gnathopod, is an undeveloped male, the second antenna (fig. 20)
x
Fig. 2a.—Second antenna of male, immature (Brighton specimen).
Fig. 2b.—Second antenna of male, immature (Roto-iti specimen).
is very stout, the fourth joint being almost as stout as the preceding but without
any trace of the apical lobe, while the fifth joint is larger in proportion than in
the mature males from Brighton and from Brisbane. In some of the specimens
the proximal joints of the flagellum appear to be partially fused though the line
of junction is clearly visible; like the peduncle they bear a few fine sete. In the
females the second antenna remains much more slender and without the lobe on
the fourth joint. .
PARACOROPHIUM EXCAVATUM.—CHILTON 47
The antennal lobe being found only in the male probably has some sexua!
function, but being flat on the inner side it perhaps also facilitates the holding
together of the right and left antennae as the animal swims backwards.
The mandible (fig. 3) is of normal structure with large molar tubercle,
the cutting edge consisting in the left mandible (fig. 3) of 3 or 4 teeth, the
accessory process being similar and of approximately the same size; the spine
row contains about 5 spines. In the right mandible (fig. 4) the accessory process
is narrower. The palp is large, second joint the longest with a small tuft of
sete at the distal end, third joint widens distally and bears numerous long sete
at the apex with a small tuft at a little distance from the apex which is obliquely
truncate.
The first maailla (fig. 5) has the inner lobe very small and delicate,
triangular, and bearing a minute setule at the end. The outer lobe ends in about
7 to 9 dentate spines; the palp has the first joint slightly swollen, the second
joint widens distally and bears at the apex 7 or 8 short setules.
- Fig. 3.—Left mandible, male (Roto-iti specimen).
Fig. 4.—Cutting edge of right mandible, male (Roto-iti specimen).
Fig. 5.—First maxilla, male (Roto-iti specimen).
Fig. 6.—Second maxilla, male (Roto-iti specimen).
The second maxilla (fig. 6) has the inner lobe shorter and broader thai
the outer and is fringed along its inner margin as well as at the rounded apex,
in the outer lobe the long sete are mainly confined to the apex.
The maxillipeds (fig. 7) have the inner lobe bearing at the apex 4 long
setz and several shorter ones, the outer lobe is somewhat narrow and reaches
nearly to the end of the second joint of the palp, the inner margin is thickly
fringed with sete. In the palp the second joint is much the longest; the propod
is oval, produced at the outer apex slightly beyond the base of the finger; the
finger is about half as long as the propod; all the joints bear long setules as
shown in the figure.
48 MEMOIRS OF THE QUEENSLAND MUSEUM.
The first gnathopod (figs. 8 and 9) has the side plate large and subtri-
angular, projecting forwards, its anterior angle rounded and bearing a fringe of
long sete which extends also along the lower margin; the basal joint widens
considerably distally and bears a few long sete towards the apex; the ischium
N
Fig. 7.—Maxilliped, male (Roto-iti specimen).
Fig. 8.—First gnathopod, male (Brisbane specimen).
Fig. 9.—First gnathopod, female (Brisbane specimen).
and merus are both short, their posterior margins bearing long sete, those on.
the ischium reaching nearly to the middle of the carpus; the carpus is greatly
elongated, being narrowly oblong in shape, its posterior margin densely fringed
with a double row of long sete, a few scattered ones being also found on the
anterior margin; the propod is much shorter than the carpus, about the same
width throughout, its palm nearly transverse, not defined, being closely over-
lapped by the dactyl; near the base of the propod is an oblique row or tuft of
about 6 or 7 long sete. This appendage seems to be the same in both sexes.
The second gnathopod in the fully developed male (fig. 10) is much
stouter than in the female, the side plate is oblong with the inferior angles.
rounded, its inferior margin with a few long sete; the merus is produced into a
long scoop-like process reaching considerably beyond the end of the carpus which,
together with the basal portion of the propod, appears to fit into the hollow of
the scoop, one margin of which is fringed with very long sete most of which
19 1G A
Fig. 10.—Second gnathopod, male (Brisbane specimen).
sig. 10a.—Second gnathopod, male, immature (Brighton specimen).
PARACOROPHIUM EXCAVATUM.—CHILTON. 49
reach quite to the end of the propod, the other (inner) margin being apparently
free from sete ; the carpus is short, triangular in shape, widening greatly distally,
its posterior border bears long hairs and there is an oblique row on the outer
surface ; the propod is oblong-oval, as wide at the base as the carpus, but becoming
slightly wider distally; both anterior and posterior margins are convex, the
posterior one being produced into a long slender curved tooth, separated by a
deep depression from a similar but shorter tooth arising from the middle of the
palm ; the finger is short, not reaching quite to the end of the palm.
In the female the second gnathopod (fig. 11) has the scoop-like process
on the merus as in the male, but differs greatly in the carpus and propod, both
of which are much longer and more slender, as shown in the figure; the carpus
is longer than the propod and has the posterior margin fringed with two widely
separated rows of long sete; the propod is slender and ends simply, being
almost entirely without palm. In some cases the end of the propod is narrower
than is shown in figure 11.
103
Fig. 10b.—Second gnathopod, male, immature (Roto-iti specimen).
Fig. 11—Second gnathopod, female (Brisbane specimen).
In the young male the second gnathopod appears to be at first similar
to that of the female, the adult male characters being gradually developed.
Figure 10b shows an intermediate stage in which the carpus and propod, though
broader than in the female, are still somewhat narrow; the propod has the
posterior margin produced into a tooth only half the length it afterwards
becomes, while the tooth in the centre of the palm is hardly yet apparent; the
daetyl reaches slightly beyond the end of the palm in this case. This figure is
drawn from a Lake Roto-iti specimen. I have a similar one mounted from
Brighton (fig. 10a) in which the defining tooth of the palm is still shorter and
the finger overlaps the palm as it does in the female.
The first (fig. 12) and second pereopoda are of normal shape and do not
eall for special description.
In the third perwopod (fig. 18) of fully developed animals the basal joint
is very large and much produced posteriorly, the posterior margin being fringed
with long sete. The carpus bears at its postero-distal angle 4 strong curved
D
50 MEMOIRS OF THE QUEENSLAND MUSEUM.
spines with a similar tuft of 3 placed more proximally. Similar curved spines
are found also on the propod and appear quite similar to those in several species
of Corophium. In more immature speéimens the basal joint is much narrower
Fig. 12.—First pereopod, male (Brisbane specimen).
Fig. 13.—Third perropod, male (Brisbane specimen).
Fig. 13a.—Third pereopod, male, immature (Roto-iti specimen).
and only slightly produced posteriorly. That of the Roto-iti specimen whose
second gnathopod is shown in figure 10b is represented in figure 13a. The third
pereopod of an ovigerous female from Brisbane River is represented in fig. 14,
which shows that it is very nearly the same as in the adult male.
ot te 4.
14 15 z=
MGs es
Fig. 14.—Third perropod, female (Brisbane specimen).
Fig. 15—Fourth perropod, male (Brisbane specimen).
Fig. 16—Fifth pereopod, male (Brisbane specimen).
The fourth perwopod (fig. 15) is longer than the third, and has the basai
joint broad but less produced posteriorly than in the third, its posterior margin
fringed with long spinules; strong curved spines are found on the carpus and
propod as in the third pereopod. :
PARACOROPHIUM EXCAVATUM.—CHILTON. 51
The fifth perwopod (fig. 16) is longer than the fourth, the basal joint is
somewhat narrower, the merus, carpus and propod longer; the various joints bear
stout sete and some long hairs, but not the special groups of curved spines that
are present in the third and fourth.
The first wropod (fig. 17 and 17a) has the peduncle considerably longer
than the rami, its lower margin being produced between them into a flattened
process, which when fully developed reaches nearly halfway to the end of the
rami; this is shown in side-view in figure 17; in figure 17a it is shown as seen
from above with a broadish base and the extremity forming a narrow vertical
knife-edge; numerous short spines are present on the upper margins of the
peduncle and of the rami.
The second uropod (fig. 18) is shorter, the outer ramus shorter than the
inner, both rami and the peduncle bearing a few short stout spines on the upper
margin.
The third wropod (fig. 19) has the outer ramus about twice as long as
the inner, which bears a sharp spine on its inner margin and three long hairs
at the apex; similar long hairs are found at the end of the outer ramus and at
the end of the peduncle.
ae ees) pees,
17 iF pease Fe
17 Cer
1A —z
5S eee
ne aaa
18 Tr alia =e i
Fig. 17.—First uropod, male (Roto-iti specimen).
Fig. 17a.—First uropod, female (Roto-iti specimen).
Fig. 18.—Second uropod, female (Roto-iti specimen).
Fig. 19.—Third uropod and telson, female (Roto-iti specimen).
The telson (see fig. 19) has the posterior margin somewhat produced to
a blunt triangular point, a pair of spinules being situated at each side at some
distance from the extremity.
The specimens from Lake Roto-iti are larger than those from Brighton
and Brisbane River and, as stated above, they differ in a few points, e.g. in the
second antenne of the male. It is possible that they may constitute a distinct
variety, but I have not found among them a male as fully developed as those
examined from the other localities, and in the meantime I consider the differences
as being due to immaturity. In any case the differences are slight and do not
seem to be of specific importance.
52 MEMOIRS OF THE QUEENSLAND MUSEUM.
NEW OR LITTLE-KNOWN CRANE-FLIES IN
THE QUEENSLAND MUSEUM
(TIPULIDA, DIPTERA).
By CHARLES P. ALEXANDER, PH.D., URBANA, ILLINOIS, U.S.A.
TuHrouGH the kindness of the Director, Mr. Heber A. Longman, the writer
has been enabled to study the collections of crane-flics made in the vicinity of Bris-
bane by the Entomologist of the Queensland Museum, Mr. Henry Hacker. These
collections, though not extensive, include many species of such interest that a
special report is deemed necessary.
The types, which are preserved in alcohol, are returned to the Queensland
Museum. :
At this time, it is well to designate the genotypes of the numerous new
genera of Australian crane-flies, proposed in 1889 and 1890 by Skuse :—
Triphana Skuse (subgenus of Geranomyia); Proc. Linn. Soe. New
South Wales, vol. 4 (ser. 2), p. 777; 1889. Two species. Type,
Geranomyia (Triphana) lutulenta Skuse, the first species, by
present designation.
T etraphana Skuse (subgenus of Geranomyia) ; l.c., pp. 780, 781. One
species. Type, Geranomyia (Tetraphana) fusca Skuse, by
monotypy.
Leiponeura Skuse (subgenus of Gonomyia) ; 1.c., pp. 795, 796. Two
species. Type, Gonomyia (Letponeura) skusei Alexander (gracilis:
Skuse, preoccupied) ; the first species by designation of Alexander,
Proc. U.S. Nat. Mus., vol. 44, p. 503; 1913.
Amphineurus Skuse (subgenus of Ormosia) ; Le., pp. 800, 801. Two
species. Type, Ormosia (Amphineurus) wmbratica Skuse, the
first species, by present designation.
Tastocera Skuse; l.c., pp. 815, 816. Two species. Type, Tasiocera
tenuicormis Skuse, the first species, by present designation.
Rhabdomastix Skuse; l.c¢., pp. 828, 829. One species. Type, Rhabdo-
mastiz osten-sackent Skuse, by monotypy.
Lechria Skuse; l.c., pp. 830, 831. One species. Type, Lechria.
singularis Skuse, by monotypy.
NEW OR LITTLE-KNOWN CRANE-FLIES—ALEXANDER. 53
Clytocosmus Skuse; Proe. Linn. Soc. New South Wales, vol. 5 (ser. 2),
pp. 74-76; 1890. One species. Type, Clytocosmus helmsi Skuse,
by monotypy.
Platyphasia Skuse; l.c., pp. 84, 85. One species. Type, Platyphasia
princeps Skuse, by monotypy.
Plusiomyia Skuse; l|.c., pp. 86, 87. Five species. Type, Plusiomyia
olliffi Skuse, the second species, by present designation.
Habromastix Skuse ; 1.c., pp. 93, 94. Three species. Type, Habromastix
cinerascens Skuse, the first species, by present designation.
Phymatopsis Skuse; l.c., pp. 97, 98. One species. Type, Phymatopsis
ngrirostris Skuse, by monotypy.
Acracantha Skuse; l.c., pp. 109-111. Three species. Type, Acracantha
sydneyensis Skuse, the first species, by present designation.
Ischnotoma Skuse; l.c., pp. 114, 115. Three species. Type, Ischnotoma
serricornis Macquart, the first species, by present designation.
FAMILY TIPULIDA.
SuspramMity LIMNOBIIN AS.
TrisE LIMNOBIINI.
Genus LIMNOBIA Meigen.
1818. Syst. Beschreib., vol. 1, p. 116.
Limnobia bidentata Skuse.
1889, Limnobia bidentata Skuse; Proc. Linnean Soc. New South Wales, vol. 4 (ser. 2), pp.
782, 783.
The following records are included in the material at hand :—Brisbane,
April 7, 1914 (H. Hacker). Caloundra, Sept. 30—Oct. 28, 1913 (H. Hacker).
Sandgate, Jan. 24, 1914 (H. Hacker).
Skuse describes the radial sector as being only one-third longer than the
deflection of R4+5; in the present material the deflection is proportionately
longer but there are no other apparent differences.
Trine ANTOCHINI.
Genus RHAMPHIDIA Meigen.
1830. Syst. Beschreib., vol. 6, p. 281.
Rhamphidia communis Skuse. ;
1889. Rhamphidia communis Skuse; Proc. Linnean Soc. New South Wales, vol. 4 (ser. 2), pp.
788, 789. j
A few specimens in poor condition, from the following locality and dates :—
Brisbane, July 30, 1914; September 16, 1918 (H. Hacker).
54 MEMOIRS OF THE QUEENSLAND MUSEUM.
Trine ERIOPTERINI.
GeNus GNOPHOMYIA Osten Sacken.
1859. Proc. Acad. Nat. Sci. Philadelphia, p. 223.
Gnophomyia fascipennis (Thomson).
1869. Limnobia fascipennis Thomson; Eugenies Resa, Dipt., p. 447; 2.
1887. Gnophomyia cordialis Osten Sacken; Berliner Ent. Zeitschr., vol. 31, pt. 2, p. 199; ¢ .
There are numerous specimens of this very interesting fly from the
following stations:—Brisbane, October 10-17, 1916 (H. Hacker); several.
Sunnybank, near Brisbane, October 27, 1916 (H. Hacker). Caloundra, October
28, 1913 (H. Hacker). Stradbroke Island (H. Hacker).
The differences between the sexes of this fly are remarkable and quite
unequalled in any other member of the genus.
GeNuS CONOSIA van der Wulp.
1880. Tijdschr. v. Ent., vol. 23, p. 159, Pl. 10, figs. 5-7.
Conosia irrorata (Wiedemann).
1828. Limnobia irrorata Wiedemann; Aussereur. Zweifl. Ins., vol. 1, p. 574.
Several specimens from the following station and dates:—Brisbane, March
23, 1916; April 23, 1916; December 12, 1915 (H. Hacker).
This species is presumably of wider distribution than any other crane-fly,
ranging from Australia to South Africa and northward to Japan.
TrinE LIMNOPHILINI.
Genus LECHRIA Skuse.
1889. Proc. Linnean Soc. New South Wales, vol. 4 (ser. 2), pp. 830, 831.
LECHRIA SUBLA-VIS sp. n.
Male.——Length, 6 mm.; wing, 6-6 mm.
Female. —Length, about 6-2 mm.; wing, 7-3 mm.
Generally similar to L. rufithorax Alexander (North Queensland),
differing as follows :—
Rostrum, palpi, and antenne brown; flagellar segments of the latter
suboval to subeylindrieal.
Mesonotum reddish brown, the three usual prescutal stripes ‘entirely
confluent, the lateral stripes continued back onto the lobes of the scutum.
Halteres pale. Wings with the veins light brown, not so distinct as in rufithoraz ;
the costal cell is scarcely darker than the remainder of the wing. Venation:
Rs considerably longer, the section before r—m being about equal to the section
of M1+2 before r-m; the section of Rs beyond 7—m longer than this crossvein ;
Sc ends opposite or before the end of Rs.
NEW OR LITTLE-KNOWN CRANE-FLIES.—ALEXANDER. 55:
Male hypopygium similar to L. rufithorar, but the shape of the pleural
appendages different; outer pleural appendage slender, gradually tapering to
the acute, curved point, with scarcely any denticles on the inner face before
the tip as in rufithorax ; inner pleural appendage more slender and more strongly
arcuated.
Habitat: Queensland.
Holotype, ¢ , Caloundra, September 30, 1913 (H. Hacker).
Allotopotype, @ .
Type in the collection of the Queensland Museum.
The genotype of Lechria, L. singularis Skuse, differs from all the other
species of the genus, as known, in that the radial crossvein connects with R2+3
at its fork, in the other species connecting with R2 some distance beyond the fork:
Skuse interpreted the genus as having the vein RZ ending in vein R2+3 and
the crossvein r lacking. The writer would interpret the venation as having the
extreme tip of RZ atrophied and the radial crossvein present but oblique in
position and simulating a section of vein R7. These characters are found in many
Cylindrotomine, and it may be that Lechria belongs in the neighbourhood of this
subfamily rather than in the tribe Limnophilini where it is now placed.
Genus GYNOPLISTIA Westwood.
1835. London and Edinburgh Phil. Mag., vol. 6, p. 280.
Gynoplistia viridis Westwood.
1835. Gynoplistia viridis Westwood, London and Edinburgh Phil. Mag., vol. 6.
One male from Sunnybank, near Brisbane, October 27, 1914. One female
from Brisbane, October 17, 1916 (H. Hacker).
Suspramity TIPULINA.
TriBeE DOLICHOPEZINI.
Genus DOLICHOPEZA Curtis.
1825. British Entomology, vol. 2, p. 62.
DOLICHOPEZA BREVIFURCA QUEENSLANDICA subsp. n.
Mesonotum reddish brown, the prescutum with three darker brown stripes;
pleura yellow; legs light brown; wings pale brown, the stigma darker brown ;
an obliterative area before and beyond the stigma and across the inner end of
cell 1st M2; abdomen yellow, ringed with brown.
Male—Length, 9-9-5 mm.; wing, 9-4-10 mm.; antenna, 4 mm.
Female.—Length, 9-4 mm.; wing, 9-3-10-3 mm.
Frontal prolongation of the head short, yellow; palpi pale brown, the
56 MEMOIRS OF THE QUEENSLAND MUSEUM.
long terminal segment whitish. Antenne of the male moderate in length, light
brown. Head brownish grey, especially on the vertex and occiput; front and
gene more yellowish.
Mesonotal prescutum reddish brown, with three darker brown stripes;
scutal lobes marked with this same colour; remainder of the mesonotum yellow.
Pleura yellowish. Halteres pale brownish yellow, the base of the knob darker.
Legs with the coxe and trochanters yellow; remainder of the legs light brown.
Wings with a faint brownish tinge; cell Sc pale brown except at the distal end;
an obliterative area before and beyond the stigma and another across the
proximal end of. cell 7st M2; stigma large, subrectangular, brown; an indistinct
brown cloud at the basal deflection of Cu1; veins pale brown, the tip of Sc and
R below it yellowish. Venation: Rs transverse, about equal to r—m; cell M1
rather short; vein M1 about equal to M@1+-2; petiole of cell M2 from one-half
to two-thirds of M1+2; basal deflection of Cui about its own length before
the fork of M.
Abdomen yellowish, ringed with dark brown; on the second segment at
about midlength, on the other segments appearing as a basal ring and a narrow
ring on the posterior margin of the preceding tergite. Male hypopygium with
the ninth tergite bidentate as in the Australian species of the genus; outer
pleural appendages relatively short, about twice the length of the small, flattened,
more complicated inner appendage; the proximal face of the outer appendage
is indistinctly bidentate. Female ovipositor with the valves acicular.
Habitat: Queensland.
Holotype, d , Oxley, near Brisbane, September 4, 1914 (H. Hacker).
Allotopotype, @.
Paratopotype,d ; paratypes, 5 2’s, Brisbane, May 23, 1916 (H. Hacker) ;
a few dry fragments, Brisbane, September 18, 1914 (H. Hacker).
Type in the collection of the Queensland Museum.
The present form would seem to be a variety of Dolichopeza brevifurca
Skuse, although the pleural appendages of the male hypopygium are very much
smaller than in any other member of the subgenus Apevlesis that I have seen,
agreeing more nearly with the normal type of the genus. The variety may be
told from the typical form by the darker wings and the structure of the
hypopygium.
TrisE TIPULINI.
Genus PTILOGYNA Westwood.
1835. Zoolog. Journ., vol. 5, pp. 448, 449.
Ptilogyna ramicornis (Walker).
1835, Tipula ramicornis Walker; Ent. Mag., vol. 2, p. 469.
A few specimens of this beautiful crane-fly, from the following station :—
Caloundra, September 28—October 28, 1913 (H. Hacker).
NEW OR LITTLE-KNOWN CRANE-FLIES.—ALEXANDER. 57
GENUS LEPTOTARSUS Guérin.
1838. Voyage de la Coquille, Dipt., p. 286.
Leptotarsus (Leptotarsus) macquarti Guérin.
1838. Leptotarsus macquarti Guérin; Voyage de la Coquille, Dipt., p. 286, pl. 20, fig. 1.
1848. Tipula tricincta Walker; List Diptera Brit. Mus., vol. 1, p. 73.
One female, Brisbane, February 12, 1914 (H. Hacker).
Suscenus PSEUDOLEPTOTARSUS subgen. n.
Antenne with nine segments. Terminal segment of the palpus short.
Wings with cell M7 lacking, there being but two branches of media attaining the
wing-margin.
Type of the subgenus: Leptotarsus (Pseudoleptotarsus) liponeura sp. n.
(Australia).
The presence of but two branches of media is almost unique in the subfamily
Tipuline, the only other case known to the writer being the genus Jdiotipula
Alexander (Natal).
LEPTOTARSUS (PSEUDOLEPTOTARSUS) LIPONEURA sp. n.
Antennez 9-segmented ; last segment of the palpus short; general coloura-
tion yellow, the mesonotal prescutum with four brown stripes; mesepisternum
and mesosternum plumbeous brown; wings pale brown, sparsely variegated with
yellow; cell M1 lacking; abdominal tergites yellow, the basal four segments
ringed caudally with black.
Female.—Length, about 13-13-5 mm.; wing, 13-5-14-6 mm.
Frontal prolongation of the head yellow; nasus broad; palpi brown, the
terminal segment darker, shorter than the second and third taken together.
Antenne with but nine apparent segments, the first segment yellow, the remainder
of the organ brown; first scapal segment long and slender; second segment sub-
elobular ; first flagellar segment oval, the second to fourth produced into a basal
pedicel, the terminal three segments nearly cylindrical; the last segment is longer
than the penultimate and may be formed by the union of two small segments.
Head obscure yellow, very broad behind the eyes.
Pronotum brown. Mesonotal prascutum obscure yellow with four brown
‘stripes, the intermediate pair separated only by a capillary line of the ground-
colour, entirely confiuent behind; a subtriangular dark-brown area at the humeral
angle; remainder of the mesonotum lght yellow, each scutal lobe with a
conspicuous dark-brown spot at the cephalic lateral angle. Pleura obscure
yellow, the mesepisternum and mesosternum plumbeous brown, the latter with
a yellow spot immediately cephalo-dorsad of the mesocoxa. Halteres pale, the
knobs brown. Legs with the coxe obscure yellow, the anterior coxz more
infuseated ; trochanters dull yellow; femora light brown, gradually darkening
to the tips; remainder of the legs dark brown. Wings pale brown, sparsely
variegated with yellowish; cells C and Sc brown; conspicuous brown seams
58 MEMOIRS OF THE QUEENSLAND MUSEUM.
along the cord and longitudinal veins beyond the cord; the yellow areas are
restricted to the basal and anal cells, the first spot basal in position, occupying
the inner end of cell M and near the base of 7st A but interrupted in between;
the second area lies across cells R and M immediately before the origin of Rs;
a similar pale area across the bases of cells R3 and R5; veins dark brown.
Venation: Cell Mi lacking; m—cu punctiform, situated at about midlength of
eéll 1st M2.
Abdominal tergites yellow, the basal two segments brighter, each with
a low triangular black band across the caudal margins, the apex of the triangle
being directed cephalad; segments 3 and 4 are narrowly margined caudally with
brownish black; remainder of the tergites yellowish; sternites yellow, segments
4 to 8 obscure brownish. Ovipositor with the valves very blunt and fleshy as
in the genus.
Habitat: Queensland.
Holotype, 2, Brisbane (H. Hacker).
Paratopotype, 9, February 12, 1914.
Type in the collection of the Queensland Museum.
Genus HABROMASTIX Skuse.
1890. Proc. Linnean Soc. New South Wales, vol. 5 (ser. 2), pp. 93, 94.
There have been three Australian and six Ethiopian species of this genus:
so far made known. In the present collection, two species are included, neither
of which agrees with any of the hitherto known forms.
HABROMASTIX PARALLELA sp. n.
Frontal prolongation of the head elongate, ght brown; mesonotal
prescutum pale brownish yellow with four darker brown stripes; mesonotal
postnotum dark brown posteriorly, including a conspicuous mark in front of the
halteres; halteres pale at their apices; legs yellowish, the femora and tibize
tipped with dark brown; wings hyaline, the basal cells cross-banded with grey,
in the cell behind vein Cu appearing as four dark-brown spots.
Female —Length, 23 mm.; wing, 19-3 mm. Hind leg, femur, 13 mm.;.
tibia, 15-8 mm.; tarsus, 35 mm.
Frontal prolongation of the head very long and slender, light brown,
with a narrow, slightly darker lateral line; palpi light brown. Antenne with
the basal segments light yellow, the flagellum broken beyond the basal segment,
this latter elongate. Head dark brown; eyes large and protuberant, very
narrowly separated beneath, more widely separated above.
Mesonotal prescutum pale brownish yellow with four darker brown stripes ;
remainder of the mesonotum yellow, only the posterior half of the postnotum
dark brown, this including a conspicuous area on the lateral sclerites of the
postnotum in front of the halteres. Pleura yellow, a conspicuous brown area
on the mesosternum, ventral portions of the mesosternum, and on the outer faces
NEW OR LITTLE-KNOWN CRANE-FLIES.—ALEXANDER. 59°
of the fore cox. Halteres long and slender, brown, the apices of the knobs pale.
Legs with the coxe brown, the fore coxe darker; trochanters brownish; femora
pale brown, the tips broadly dark brown; tibixwe yellow, the tips narrowly dark
brown; tarsi yellowish. Wings hyaline, variegated with dark brown and light
grey; the costal cell is brown with three conspicuous hyaline spaces, one just
beyond the h crossvein, the last at the end of the cell; cell Sc hyaline; the grey
areas appear as about four bands that traverse the wings, being continuations.
of the brown costal areas; in the space immediately behind vein Cw these four
bands become dark brown and very conspicuous; the hyaline bands between
these grey areas are about one-half the width of the latter; indistinct whitish
areas in the base of cell #2, bases of cells M1, 2nd M2, and M4 and in eell
ist M2; stigma conspicuous, dark brown; a narrow brown seam along the cord;
veins dark brown, paler in the hyaline areas. Venation: Sc2 ending just before
midlength of k2+3; Rs short, scarcely longer than cell ist M2 and shorter
than the basal deflection of Cul; R3 about one-half longer than R2+3, running
parallel to h4+-5 except at the extreme tip; cell 2nd R1 narrow; inner ends
of cells R3, R5 and 1st M2 in alignment; petiole of cell Mi less than one-half
this cell; m and the deflection of M3+4 subequal; m—cu obliterated by the
punctiform contact of Cul and M3.
Abdominal tergites yellow, the first brown medially; segments 2 to 5
with an oblique dark-brown dash on either side, these marks converging behind,
becoming more approximated on the posterior segments, on segments 6 to 8-
appearing as confluent median dashes; a triangular brown mark at the anterior
lateral angle of tergites 3 to 8 and at midlength of tergite 2; sternites yellowish.
Valves of the ovipositor long and straight, the tergal valves longer.
Habitat: Queensland.
Holotype, 2 , Brisbane (H. Hacker).
Type in the collection of the Queensland Museum.
Habromastiz parallela is the largest species so far made known. It is.
most closely related to H. remota (Walker) in the long petiole of cell M1 and
other characters but is readily told by the pattern of the body and wings and
the venational details.
HABROMASTIX TERRA-REGING sp. n.
Antenne of the male about one-half longer than the body; legs yellow,,.
the tips of the femora and tibie dark brown, remainder of the tarsi dark brown;
wings light grey, sparsely variegated with brown and subhyaline; petiole of
cell M1 very short to lacking; abdomen obscure yellow, the tergites with two
brown sublateral stripes.
Male.—Length, 11-5-12-5 mm.; wing, 14-15-4 mm.; antenna, 17-5-17-8 mm.
Generally similar to H. parallela.
Frontal prolongation of the head yellow, with a narrow brown lateral
line; palpi brown. Antenne of the male considerably longer than the body, the:
‘60 SEHMOIRS OF THE QUEENSLAND MUSEUM.
basal four segments yellowish, thence gradually passing into brown. .Head with
a rather rather small brown area on vertex, the remainder of the. vertex and
occiput pale buffy.
Mesonotal praescutum brownish yellow, with four brown stripes, the
_ intermediate pair indistinct in front and strongly narrowed behind; postnotum
darkened posteriorly but with no conspicuous mark on the lateral sclerites as
in parallela. Pleura obscure yellow, the mesepisternum and mesosternum dark
coloured, with a conspicuous stripe of the ground-colour across the dorsal portions
of the latter. Legs with the femora yellow, the tips broadly dark brown; tibiz
light brown, narrowly yellowish basally; tarsi dark brown. Wings light grey,
sparsely variegated with brown and subhyaline; cells C and Sc brown, the
former with three small yellowish areas, two being basal in position, the last
apical; two small areas in the basal cells, one subbasal, the other subapical; a
conspicuous whitish area beyond the cord, extending without noticeable inter-
ruption from the inner end of cell R2 across cells R3, R5, 1st M2 into the base
of M4; the brown area in the space behind vein Cw is almost continuous, but
interrupted by small pale spots at midlength and near the end. Venation: Sc
ending slightly beyond the origin of R2+3; Rs short, rather strongly arcuated
at origin; R2+-3 rather short, about equal to the basal deflection of Cui; inner
end of cell ist M2 slightly more proximad than cell R3 or R5; petiole of cell
M1 very short to entirely lacking, shorter than m; cell 1st M2 comparatively
large; 2nd Anal vein comparatively short and straight.
Abdominal tergites obscure yellow with two sublateral brown stripes, the
yellow dorso-median stripe thus formed being most distinct on segments 1 to 6,
segments 7 to 9 being almost entirely dark brown; these brown marks on the
individual segments oblique, with the posterior ends directed proximad, the
posterior lateral angles of the tergites pale; basal sternites light yellow, the fifth
to seventh sternites dark brown, especially across the posterior margins.
Habitat: Queensland.
Holotype, ¢ , Brisbane, May 23, 1916 (H. Hacker).
Paratopotypes, 6 ¢’s.
Type in the collection of the Queensland Museum.
Habromastiz terre-regine belongs to the group of H. cinerascens Skuse
and H. ornatipes Skuse. It is the closest to the latter but the deseription of the
details of the wing-pattern of ornatipes is very different. It is readily told
from H. parallela by the short petiole of cell M1, the wing-pattern, and other
characters.
Genus MACROMASTIX Osten Sacken.
1886. Berliner Ent. Zeitschr., vol. 30, pt. 2, pp. 185-187.
MACROMASTIX FLAVOPYGIALIS sp. n.
Antenne short; frontal prolongation of the head very long and slender,
dark brown; head dark, the vertical tubercle yellow; mesonotal prescutum
NEW OR LITTLE-KNOWN CRANE-FLIES.—ALEXANDER. 61
with four dark-brown stripes; scutellum and postnotum obscure yellow; wings
pale brown, the costal margin dark brown; abdomen dark brown, the base and
hypopygium yellow.
Male——Length, 12-3-13 mm.; wing, 13-15-5 mm.
Frontal prolongation of the head very long and slender, nearly twice the
length of the head, brownish black; palpi dark brown. Antenne short; scape
yellow, flagellum dark brown, the base of the first segment obscure yellow; first
flagellar segment not elongated; last segment shortest. Head dark-coloured,
the low but conspicuous vertical tubercle yellow; front obscure yellow.
Mesonotal prescutum greyish brown with four darker brown stripes;
scutum brown, the scutal lobes dark brown; scutellum and postnotum obscure
yellow, the latter narrowly darkened posteriorly. Pleura obscure brownish
yellow, the mesepisternum and mesosternum marked with darker. Halteres
brownish yellow, the knobs darker. Legs with the fore coxe dark brown, the
remaining coxe yellow, dark brown basally; trochanters yellow; femora yellow,
the apical half passing into brown; tibie brownish yellow, tarsi brown. Wings.
pale brown, the costal and subcostal cells dark brown; stigma pale; veins brown.
Venation: Rs longer than R2+-3; petiole of ceil M1 shorter than m, sometimes
only half its length; m—cu punctiform; cell 2nd A rather narrow.
Abdomen dark brown, the caudal margins of the segments indistinctly
paler; segment 1, the base of segment 2, and the hypopygium conspicuously
light yellow. Male hypopygium with the pleurites moderately long and slender,
with one complicated pleural appendage, the upper lobe of which is produced into
a long, slightly curved beak, the crest with several small blackened spicules ;
lower lobe pale, flattened, oval, the apex rounded.
Habitat: Queensland.
Holotype, ¢, Brisbane, May 23, 1916 (H. Hacker).
Paratopotype, d .
Type in the collection of the Queensland Museum.
Macromastiz flavopygialis may readily be told from all the described
Australian species by the very long, slender, dark-coloured frontal prolongation
of the head and the conspicuous colouration of the abdomen.
MACROMASTIX TORTILIS sp. n.
Antenneze short; general colouration light yellow, the prescutal stripes
slightly more reddish; wings greyish yellow, the costal and subcostal cells rich
brownish yellow; abdomen yellow, trilineate with brown, segments 7 and 8
chestnut brown; ninth tergite of the twisted male hypopygium large and tumid.
Male. —Length, about 10-12 mm.; wing, 12-14-7 mm.
Female.—Length, about 10-11 mm.; wing, 11-55-12-5 mm.
62 MEMOIRS OF THE QUEENSLAND MUSEUM.
Frontal prolongation of head short and stout, yellowish; palpi brown,
the last segment about equal to the basal three taken together. Antenne shorter
than the palpi, the scape yellow, the flagellum dark brown, flagellar segments
beyond the third slender, cylindrical. Head light yellow; vertical tubercle low.
Mesonotal preescutum light yellow, with the three usual stripes more
reddish yellow ; remainder of the mesonotum pale yellow; an indistin¢et, capillary,
median brown line. Pleura yellow. Halteres pale brown, the knobs darker.
Legs with the cox and trochanters light yellow; femora yellow, the tips
narrowly and indistinctly infuscated; tibie and tarsi brown. Wings greyish
yellow, the costal and subcostal cells rich brownish yellow; stigma yellowish
brown; veins brown. Venation: Rs scarcely longer than k2+-3; h3 long, parallel
to R4+5 except at the tip; petiole of cell M1 variable in length, from as long
as m to entirely lacking, cell 1/7 being short-petiolate to sessile ; m—cw punctiform.
Abdomen yellow, the tergites with three brown stripes, the median stripe
very broad and conspicuous; segments 7 and 8 chestnut brown; hypopygium
yellow, only the ninth tergite brown; sternites yellow. Male hypopygium with
the ninth segment twisted around nearly one-half so that the ninth tergite
occupies a ventro-lateral position; this latter is large and tumid, with a broad
U-shaped median notch, the margins very thick, the conspicuous lateral lobes
thus formed stout-triangular. The pleural appendages are two in number, the
dorsal one (nearest the tergite though actually ventral in position) a small
blackened blade with the inner edge conspicuously serrate ; the ventral appendage
is larger, pale, flattened, gradually dilated distally, the apical margin set with
about fourteen blackened spicules of which the four inner are isolated from one
another, the remaining one confluent in a slightly curved row. Ovipositor with
the valves short and fleshy as in the genus.
Habitat: Queensland.
IIolotype, ¢ , Brisbane, July 1, 1913 (H. Hacker).
Allotopotype, 2.
Paratopotypes, 18 ¢9 ’s, July 1 (1913) and May 23 (1916)
Type in the collection of the Queensland Museum.
Macromastiz tortilis is readily told from the other described species of the
genus by its light-yellow colour, the trilineate abdomen, and especially the large,
twisted male hypopygium.
MACROMASTIX HACKERI sp. n.
Antenne short; general colouration yellow; wings brownish yellow, the
costal margin more saturated; stigma darker brown; cell 2nd A very narrow;
male hypopygium semi-inverted, the ninth tergite pale; ninth sterno-pleurite
with a pencil of about seven powerful bristles near the dorso-caudal angle.
Male.—Length, about 11 mm.; wing, 12-5 mm.
Generally similar to M. tortilis.
NEW OR LITTLE-KNOWN @GRANE-FLIES.—ALEXANDER. 63
Frontal prolongation of the head short and stout, obscure yellow; palpi
brown. Antenne short, pale brownish yellow, the distal flagellar segments long-
cylindrical. Head yellow.
Mesonotum and pleura dull yellowish, the prescutum without distinct
stripes. Halteres pale brown. Legs with the coxe and trochanters dull yellow,
the remainder of the legs pale brown, the tips of the femora narrowly darkened.
Wings with a strong brownish yellow tinge, the costal and subcostal cells more
saturated; stigma conspicuous, darker brown. Venation: Basal deflection of
Cul at about one-third the length of cell 1st M2; cell 2nd A very narrow.
Abdomen obscure yellow, the lateral margins of the tergites narrowly and
indistinctly dark brown; the dorso-median stripe of tortilis is lacking; segments
7 and 8 brown; hypopygium entirely pale yellow, including the ninth tergite.
Male hypopygium of the semi-inverted type of MW. tortilis. Ninth tergite not so
tumid as in tortilis, with a broad and deep U-shaped notch, the dorsal surface
provided with numerous stout sete, the free ends of the tergite produced caudad
and slightly inward into a subspatulate, flattened lobe. Ninth sterno-pleurite
with a large, square median notch; near the dorso-caudal angle with a pencil of
about seven powerful bristles that are directed dorsad and caudad.
Habitat: Queensland.
Holotype, ¢ , Brisbane (H. Hacker).
Type in the collection of the Queensland Museum.
This interesting species is named in honour of the collector, Mr. Henry
Hacker, Entomologist of the Queensland Museum. It is closest to M. tortilis but
is readily separated by the diagnostic characters listed above.
64 MEMOIRS OF THE QUEENSLAND MUSEUM.
LITTLE PENGUIN “IN QUEENSEAN@
By HeBER A. LONGMAN, F.L.S. (DIRECTOR).
TuHrRouGH the kindness of Mrs. Fagan, ‘‘Greenmount,’’ Tweed Heads, a
specimen of the Little Penguin, Ludyptula nuinor nove-hollandiew (Stephens),
captured at Coolangatta beach near the southern border of this State, has been
added to the Queensland Museum collections. Mrs. Fagan records that this
specimen was picked up alive on the 15th May of this year; it refused to eat
anything and died within about twenty-four hours after capture.
The occurrence of. this specimen within the Queensland border is a matter
for surprise. According to A. J. North,! Cabbage-tree Island, near the entrance to
Port Stephens (about S. lat. 32° 40’) was the most northerly record (breeding).
Coolangatta is north of Point Danger, about S. lat. 28° 7’. The Little Penguin
just succeeded in crossing our boundaries.
Apparently this bird requires trinomial designation. Mathews refers
to the variations shown in specimens from West Australia, Tasmania, South
Australia, and New South Wales, but finds ‘‘that the white tail of the Australian
form renders it separable from the New Zealand bird, which is also constantly
darker.’’? He utilises Stephens’s term nove-hollandie for Australian birds as
a subspecies of H. minor, subsequently restricting this to New South Wales forms.
and noting other subspecies. Alexander and Brooke Nicholls, however, after a
valuable study of a large number of specimens, came to the conclusion that all
the Australian birds (including H. wndina) should be classified as Eudyptula
minor nove-hollandw.*
In view of the scantiness of available material for comparison no extended
description is made. Our specimen has the characteristic blue-grey colouring,.
with a beautiful sheen, on the upper surface; the inner margins of the flippers
are whitish, as is also the apical third of the tail; the fore-neck and lower surface
are white. Following the body curves of the mounted specimen the bird is
460 mm. in length. The bill is 38 mm. in length; depth 15 mm.
From the excellent skin made and forwarded by Mrs. Fagan, our taxider-
mist (Mr. M. J. Colelough) has mounted the bird, which makes a very attractive
addition to the case of penguins on exhibition.
* Nests and Birds Austr. & Tas., iv, 1914, p. 392.
? Birds of Australia, i, pt. 5, 1911, p. 285.
3<¢The Emu,’’ xviii, 1918, pp. 50-57.
[MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Part I, 1920.]
By Authority: ANTIIONY JAMES CUMMING, Government Printer, Brisbane.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Parr Il.
(ISSUED FEBRUARY 11, 1921.)
A NEW GENUS OF FOSSIL MARSUPIALS.
By Heper A. LonemMan, F.L.S.,
Director of the Queensland Museum.
(Plates IV-VIL.)
In 1915, when describing a giant turtle from the Queensland Cretaceous
formations, the writer ventured to forecast that, when our areas were better
known, novelties rivalling the grotesque monsters of other lands would be
exhumed. No new vertebrate material has yet been received from these
Cretaceous sources, but remains from the Post-Tertiary deposits on the
Darling Downs, which form the subject of this paper, exhibit a large marsupial
with remarkable cranial contours. In life this mammal must have been bizarre
as a monster in an artist’s realm of phantasy. Here is a member of. the
Nototherium group with a skull the maximum width of which exceeds the
maximum length by 46 mm.
This extreme brachycephalous condition is mainly the result of masseteric
processes or large inferior lateral extensions of the anterior part of the jugal,
which flare widely outwards on each side of the head, almost at right angles
to the sagittal plane, at the junction of the infratemporal bar with the zygomatic
processes of the maxilla. For reasons to be subsequently set out, it is suggested
that this unusual development of the zygomatic arches was associated with the
presence of large cheek-pouches.
Diagnosis or New Genus.—The extraordinary development of the
inferior lateral processes of the anterior part of the zygomata and the
architecture of the very prominent suborbital platform, which acts as a buttress,
demand generic recognition. These characters are also associated with a
subtriangular upper premolar (dealt with in detail later).
EURYZYGOMA, genus new.
De Vis associated with the mandibles of his Nototheriwm dunense (1887,
p. 1065, and 1888, pp. 111-116),* two cranial fragments (Nos. 12622 and 12618)
which are obviously of the type of our new material, and which also came
*In this paper references are noted in the manner suggested in a circular recently
issued by the Committee of the British Association on Zoological Bibliography.
E
66 MEMOIRS OF THE QUEENSLAND MUSEUM.
from the Darling Downs, being heautotypes. These specimens, however, were
too incomplete to suggest to De Vis the immense accessory processes. Sufficient
of the anterior. zvygoma root is present to show the special characters of
Euryzygoma. It would therefore be unwise to give our material a new specific
name. In the circumstances, it is necessary to use De Vis’s specific name in
conjunction with the new genus.
EURYZYGOMA_ DUNENSE.
MarertAL.—This consists of a cranium which was received in over
seventy pieces. The maxilla and premaxilla were practically the only parts
which were intact. Fortunately the bones were in excellent condition, not
being decomposed, and as many of the fractures were obviously 1nade in
bringing the remains to light, the zygomatic arches, with their inferior lateral
processes, and the main portion of the superior cranial contours have been
united without any doubt as to their real position. The fitting together of the
smaller pieces, especially in the basioccipito-sphenoidal region, however, demanded
infinite patience. Practically no reconstruction was necessary, the actual bones
themselves supplying the natural contours, and it has been deemed inadvisable
to fill in the comparatively unimportant missing parts. The full molar series
is present on each side, but the incisors have been lost post mortem.
The specimen was probably a fully mature male. Many of the cranial
sutures are ankylosed to extinction. Reg. No. F 1327.
Locauiry.—The cranium was discovered in sandy soil at a depth of
about 70 feet at Brigalow, Darling Downs, Queensland, when a well was being
sunk on the property of Mr. G. A. F. Kleidon, who subsequently donated the
fragments to the Queensland Museum. On behalf of this institution it is my
pleasant duty heartily to thank Mr. Kleidon, and also Mr. Wilson who
forwarded the pieces. It is to be hoped that this handsome donation will be
supplemented later by remains from this district demonstrating an association
of bones.
DESCRIPTION.—The maximum length of the cranium ‘‘between uprights’’
(condyles to gnathion) is 634 mm.; the maximum breadth across the zygomatic
processes is 680 mm.; the maximum height (between parallels from the bregma
and the inferior border of the zygomatic processes) is 343. The calvarium is
dwarfed by the extraordinary development of the zygomatic arches and
processes. The breadth of the occiput (calculated from the fairly complete left
moiety) is approximately 290 mm. The occipital region, which is somewhat
concave, with a median vertical ridge, slopes forward at an angle of 45° to
the plane of the bony palate. Posteriorly the lambdoid crest is broadly convex.
From the lambdoid erest, the superior contour of the calvarium along the
MEMOIRS OF THE QUEENSLAND MUSEUM.
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A NEW GENUS OF FOSSIL MARSUPIALS.—LONGMAN.
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sagittal suture to the fronto-nasal depression is practically a straight line,
which is parallel with the plane of the bony palate. There is no parietal
platform, and the cranial walls, when losing their curves of contact with the
occiput, slope in straight lines to their fractures. In the mid-parietal region
these walls form an angle of 50°. The lower moieties of the lateral walls are
missing.
On the dorsal surface the sutures between the parietals and frontals are
obscured. In this area the sagittal crest bifurcates, as in Phascolarctus; the
frontals rapidly widen and when in line with the inferior zygomatic processes.
they attain a breadth of 148 mm. This breadth is apparently continued
(contours incomplete) through the course of the fronto-nasal depression. The
concavity of this depression is wide, the sides and posterior wall sloping sharply
to the main frontal platform. The course of the fronto-nasal sutures cannot
be traced with accuracy. The terminal areas of the frontal bones are slightly
rugose on the slope of the depression, and there are a few very small nutrient
foramina. A small horn or frontal boss may have been attached here (cf.
Scott & Lord, 1920). ;
Nasals.—From the frontal depression the nasals reach almost to a point
above the gnathion. Except for a slight convexity, which is double in the
anterior region where the median suture is open, they are surprisingly flat on
their upper surface, which projects forward at an angle of about 18° to the plane
of the bony palate. Their course does not appear to be curved, or to terminate
in a deflected obtuse apex. The left-hand moiety.is perfect anteriorly, and is
88 mm. in width. A right-hand fragment is sufficient to give fair evidence of
symmetry, and the approximate width would thus be 176 mm. There is no
evidence whatever of the presence of bony studs, of which Scott and Lord have
made interesting studies in Tasmanian material (NV. mitchelli), or of attachments
for a horn, as first suggested by Macleay in his original description of
‘‘Zygomaturus trilobus’’ in 1857 (quoted by Owen, 1859, p. 169).
The lateral portions of the naso-maxillo-premaxillary region are, unfortu-
nately, too incomplete to be correctly allocated, and it has been thought
inadvisable to restore the missing parts. It is evident that the anterior portions
of the nasals projected clear of lateral supports for a distance of at least 75 mm.
Fragments of the sides, and especially one piece exhibiting the maxillary-nasal
suture, show the great strength of the bones supporting the main nasal arch. On
the inner median surface of the nasals, near the suture with the frontals, may be
seen two large sub-contiguous channels, 11 mm. wide and 5 mm. deep, and these
extend towards the anterior extremity where they are merged in the general
concavity of the inner tables.
68 MEMOIRS OF THE QUEENSLAND MUSEUM.
Zygomatic Arches—These are fairly symmetrical. Part of the ventral
border of the left arch is missing, and the right inferior lateral process is
incomplete in its inner contours, but fortunately the opposite bones are in
excellent condition. The dorsal or superior border of the squamosal element of
the arch is at first shortly concave when leaving the occipital region, being then
slightly and evenly convex until the downward sweep of the orbit is reached.
The inferior or jugal border is much more robust, attaining a thickness of 56 mm.
where the jugal approaches the glenoid cavity. In this respect the cranium
resembles the relationship of these parts in Phascolarctus, the posterior extension
of the jugal being a primitive condition.
The squamosal element appears to be produced anteriorly to a greater
~ extent than is to be found in either Phascolarctus, Phascolomys, or Macropus, and
sends a V-shaped strip of bone as far down as the mid-region of the orbit. The
line of demarcation between the squamosal and jugal bones can be readily traced
throughout the arch. The height of the arch opposite from the fronto-parietal
suture is 130 mm., the squamosal element here having the greater share. The
walls are not vertical, but sweep outward as they gain depth; in fact, a section
taken through the arches in the mid-parietal region would show, with the
contours of the occiput behind, a slightly flattened semicircle, the radius of which
would be about 8 inches. The maximum width of the zygomata, apart from the
outstanding processes, is 415 mm.
W. K. Parker (1886) pointed out that an inordinately large squamosal is
characteristic of the Marsupialia. In the arch of Euryzygoma this bone
reaches its maximum development. It is to be regretted that the squamous plate
is missing and that its extent on the walls of the cerebral chamber cannot be
traced. In the Marsupialia this plate reaches its maximum in Wynyardia
bassiana, (Baldwin Spencer, 1900, p. 779).
Owing to the incompleteness of the cranial walls the width of the temporal
fossa cannot be accurately estimated, but in the mid-parietal region it was
evidently about 65 mm., widening somewhat anteriorly. The facial portion is
elongated, as may be seen from Plate V, and the width averages 77 mm. The
orbits are low, but not to the same extent as in V. mitchelli, being 90 mm. above
the alveolar margins of the molars. Although the outer margins of the orbits are
more posteriorly placed than is the case in NV. mitchelli, there would have been
little scope for lateral vision, and Euryzygoma evidently went through its world
viewing only the region immediately in front.
A slight prominence on each side of the orbit, situated at the origin of the
suture between the jugal and the squamosal, indicates rudimentary postorbital
processes. Part of the inner margin of the orbits and the lachrymals are missing.
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[Face Page 69.]
A NEW GENUS OF FOSSIL MARSUPIALS.—LONGMAN. 69
INFERIOR LATERAL PRocEsses.—Across the processes the extreme breadth
is 680 mm. On the left-hand side the maximum transverse diameter of the
inferior lateral process is 231 mm. The vertical height of the process.in its
lateral extension beyond the arch is 121 mm. The maximum thickness is 47 mm.
The extreme height of the combined elements, from the superior border of the
squamosal to the inferior border of the vast processes (taken from parallels)
is no less than 285 mm. When the cranium is resting on the inferior borders
of the processes the molar series are 64 mm. above the surface. This downward
development is surpassed by the inferior processes of Mylodon, but in that.
mammal there are no lateral extensions.
The processes flare outward almost at right angles to the sagittal plane,
as may be seen from Plate V. They are not, however, straight, but shghtly
eurved, the lateral margins being in advance. There are prominent rugose areas
on the superior border of these processes, just at the origin of their lateral
extensions, these being doubtless for the attachment of parts of the masseteric
muscles.
The origin of the zygomatic process of the maxilla is primarily a vertical
plate which abuts on practically the whole of the vertical face of the maxilla,
commencing parallel with the anterior lobe of the third molar, and being about
30 mm. in thickness; this is supported above by the horizontal orbital plate
14 mm. thick, which is produced anteriorly at right angles to the vertical
constituent, forming a triangular platform which is continuous behind with the
superior border of the maxilla: this horizontal orbital platform unites the
vertical plate with the strong convex bar which forms the lower border of the
orbit, and through the inner root of which the large infra-orbital foramen passes.
The lateral extension of this bar forms a strong supporting buttress for the
accessory processes. With the exception of the suborbital bar, the whole pier
is composed of maxillary elements, but the sutures at the orbit cannot positively
be traced, owing to fractures. The whole architecture of this part of the skull
is quite unlike the piers of the zygomata in Macleay’s cast, and is also absolutely
distinct from other cranial fragments in the Queensland Museum attributed to
N. victorie. The deep rectangular suborbital recess is a marked feature.
Of the extreme width of the accessory process, the maxilla forms less than
one third. The zygomatico-maxillary sutures on the anterior surface are
produced laterally to a far greater extent than are those on the posterior surface,
giving the maxillary element, as illustrated in Plate V, a greater proportion
than really exists. It may here be noted, as W. K. Gregory (1910, p. 221) has ~
pointed out, that many of the cranial bones of marsupials are very oblique in
their areas of contact. :
Mazilla and Molars.—The bony palate is well preserved and there are no
signs of fenestrations. The floor is slightly concave transversely, with a median
70 MEMOIRS OF THE QUEENSLAND MUSEUM.
ridge. At the border of the mesopterygoid fossa there is a prominent transverse
ridge, through which a foramen runs on each external corner.
depth of the palate below the alveolar margins is 7 mm.
The average
The molar series are in symmetrical ares, the lobes of the teeth being
obliquely set. Fortunately all the teeth are in situ.
Premolar: The premolar is subtriangular: length 20 mm., breadth 18 mm.
There is a single oblique transverse lobe, on which a wide tract of dentine with
a posterior loop is exposed. This has evidently been worn from a single central
cusp with a median posterior depression. There is a narrow posterior talon,
the lingual border of which is continuous on the imner side with an anterior
talon on the contracted front angle of the tooth. This premolar agrees well
“with that described by De Vis (1888, p. 115) as typical of his VN. dunense and
also illustrated in his accompanying Plate. Both teeth are well preserved in
our specimen, and no special differences are to be noted between them. The
relations of the premolar with those of other Nototheres are commented on
elsewhere.
Molars: The bilophodont molar series is well worn, a tract of dentine
8 mm. wide being exposed on the hind lobe of m*. From m+ to the front lobe ~
of m® the series gradually inereases in size. The posterior lobe of m* is less
wide than the anterior. M* is anteriorly slightly less wide than the corresponding
lobe of m®, and the hind lobe is markedly narrowed. Anterior and posterior
talons are well developed throughout, except in the first true molar, where
demarecations are lost in the surfaces of wear. The anterior talons on the second,
third, and fourth molars are more developed on the lingual surface, whereas the
posterior talons are more prominent labially, this being in consonance with the
oblique setting of the lobes. On the lingual side the valleys between the lobes
are bounded and partly closed by a tubercle arising from cingular processes,
but these are not so marked on the outer side.
Dimensions. mm,
Antero-posterior diameter of molar series with pm'*, right 172
Antero-posterior diameter of molar series with pm*, left 169
Width of m’, anterior lobe 28
Width of m*, anterior lobe 35
Width of m*, anterior lobe 36.5
Width of m*, anterior lobe . ae = 35
Palate breadth between outer corners of pm* Be ar 102
Palate breadth between outer corners of m’*, front lobe 158
Palate breadth between outer corners of m*, hind lobe 152
Palate breadth between inner corners of pm’, front lobe 64
Palate breadth between inner corners of m*, front lobe 85
Palate breadth between inner corners of m*, front lobe 90
Diastema Se ie 22 eo 112
Palatal length,* palation to gnathion 390
*Oldfield Thomas, Nomenclature of Measurements, Proce. Biol. Soc. Wash.,
1905.
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MEMOIRS OF THE QUEENSLAND MUSEUM.
Vout. VII, Puare VI.
EuURYZYGOMA DUNENSE; ventral view.
Photo.—H. W. Mobsby.
A NEW GENUS OF FOSSIL MARSUPIALS.— LONGMAN. (al
The maxilla extends anteriorly on the floor of the palate for a distance
of 110 mm. beyond the premolars. The palate is here convex with a median
groove. The lateral sutures between the maxilla and the premaxilla cannot be
satisfactorily traced.
Premaxilla.—Euryzygoma dunense is relatively longer in the facial region
than is the cast of the cranium described by Macleay. The gnathion is 187 mm.
from the anterior borders of the premolars. The diastema is 112 mm. In
section the premaxilla is subquadrate. Owing to fractures its union with the
nasals is missing on both sides. On its palatal surface it is concave in the
region between the two posterior incisors, where there are three small foramina.
The premaxilla is thickened anteriorly and its superior margin terminates in
a raised oval boss, which suggests an attachment for cartilage. At the alveoli
of the large first incisors the width is 74 mm., and the height is 101 mm. The
incisors have been lost, post mortem, but the dimensions and disposition of their
cavities give valuable evidence of their nature. The large anterior pair were
evidently strongly curved in their downward course, and were subeylindrical
and only shghtly diverging. At the antero-inferior border the alveoli are
separated by a distance of 19 mm. A loose incisor, 23 < 29 in diameters, from
another specimen, fits fairly well into either cavity. On the anterior surface of
the premaxilla there are extensive exposed alveolar surfaces. The alveolus of
the second incisor, which is contiguous with that of the first on the labial border,
is approximately 13 x 11. The third incisor alveolus is placed on the labial
border about 15 mm. behind that of the second, to which it is subequal. It is
evident from its contours that the third incisors were obliquely set and
procumbent in position.
For purposes of description the premaxilla has been treated as a single
and not as a paired bone.
Glenoid Fossa.—Although the postglenoid process is very incomplete, the
actual fossa is well preserved on the right-hand side. This is 102 mm. in length,
whilst the actual concavity, apart from its shelving anterior border, is 15 mm.
Some idea of the articular surface of the mandibular condyle may be gauged
from these dimensions. The plane of the fossa is set at an angle of 85° to the
median line of the cranium, the condyle thus being scarcely oblique in its setting.
Laterally the posterior extension of the jugal forms part of the shelving anterior
border of the fossa.
Basal Bones—The basioceipital and basisphenoid have been greatly
fractured, about fifteen segments forming the contours which have been put
together. The combined bones slope upwards at an angle of 18° from the plane
of the bony palate, being 12° less than the angle noted by Owen for the cast of
lo
MEMOIRS OF THE QUEENSLAND MUSEUM.
Nototherium.. From the palation to the basion the distance is 220 mm. The
aperture of the posterior nares at the palation is 53 mm. deep and about the
same in width. The posterior borders of the mesopterygoid fossa are incomplete,
but there was a depth of at least 65 mm. The walls of the fossa expand somewhat
in their course from the maxillary region. In section the fossa is U-shaped.
The external walls are marked with very prominent tubereules for muscle
attachments. On each side a large entocarotid foramen is to be seen, which
perforates the floor of the basisphenoid and enters the inner tables between
the Gasserian grooves, its course having been cleared on the left-hand side.
The sutures between the basioccipital and basisphenoid on the inferior
surface can be traced at a distance of 83 mm. from the basion. The inferior
surface of the former bone is deeply concave, with a strong longitudinal ridge.
In its advance from the foramen magnum the bone somewhat narrows in breadth.
The contours of the foramen magnum are not quite complete. The
transverse section is oval, with diameters (approximately) of 65 X 35. Both
condyles are present, but only the right can be conjoined with its natural
surface. Its diameters are: Maximum length 78 mm.; extreme breadth 42 mm.
Unfortunately the lateral elements of the occiput are largely missing.
The prominent condylar foramina are paired, being subequal, and
externally the openings are separated by a distance of 7 mm. It has been
suggested that the accessory foramen is homologous with those of the Creodonta,
Interiorly the basioccipital and basisphenoid form a level platform about
65 mm. in width, but the supero-lateral elements are missing. On the external
border of the former may be seen, at the fractures, part of a large groove which
probably corresponds with the inner border of the jugular foramen.
There are prominent grooves, slightly diverging anteriorly, for the
accommodation of the Gasserian ganglion; from the grooves posteriorly the
position of the foramen rotundum is barely indicated on each side. Anteriorly
the basal bones are incomplete, and the whole ethmoid region is missing.
In the mature cranium of Phascolomys the cranial walls are greatly
thickened in the supra-occipital region, and large sinuses are present which are
not to be found or are but slightly developed in Macropus and Phascolarctus.
Our fossil resembles the wombats (Phascolomys) in this respect. The walls of
the cranium are surprisingly thick in this area and large sinuses are present.
The internal tables of the cranium, in so far as they are available for
description, exhibit irregularities in the region of the sagittal suture. There
is an irregular median ridge, which is much distorted in the mid-parietal
region, probably associated with a sinus. The crania of many marsupials are
much thickened in the sagittal region. Anteriorly there is evidence of large
frontal sinuses.
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MEMOIRS OF THE QUEENSLAND MUSEUM.
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[Face Page 73.]
A NEW GENUS OF FOSSIL MARSUPIALS.—LONGMAN. 73
The cerebral chamber was evidently small and elongated. The brain was
of the elongated type, having such reptilian affinities as may be assumed from
a comparison of the cranium of the marsupial Thylacinus with that of the dingo.
Judging from such measurements as can be taken, the brain of Euryzygoma
appears to have been relatively smaller than those of the native bear, kangaroos,
and wombats. The parietal walls meet in transverse section at an angle of 50°,
and the superior areas of the brain were thus much constricted laterally.
Cheek-Pouches.—Owen (1877, p. 259) refers to the maximum develop-
ment of the zygomatic arches in the Nototheres, and records the necessity for
increased attachment areas for the premasseter muscles. In Euryzygoma the
masticatory muscles were doubtless strongly developed, but it seems to the
writer that the bony architecture of the inferior lateral processes is too massive
to be entirely accounted for in this way. Much of the area is also so smooth
that the extensive origin of masticatory muscles is not suggested. The develop-
ment of large cheek-pouches may therefore be assumed in association with these
huge processes. There can be no direct evidence bearing on this point, but it is
interesting to note that the Pocket Gophers, or Geomyide, of North America
as illustrated by C. H. Merriam (1895) show a great development of the
zygomata, although there are no inferior lateral processes. The cheek-pouches
of these rodents are characteristic.
It would undoubtedly have been of advantage for this gigantic herbivorous
marsupial to be able to obtain large masses of food, probably from swamp
vegetation, some of which could be temporarily stored in cheek-pouches, and
digested at leisure. It may even be suggested that the presence of the large
crocodile, Palliumnarchus pollens De Vis (whose remains are found in the same
deposits) made a hurried meal sometimes a necessity for Euryzygoma.
In his study of the Geomyide, C. H. Merriam (1895) has pointed out
that with advancing age there is a lateral development of the zygomatic arches.
Probably the inferior lateral processes of Huryzygoma are relatively more
prominent in adult than in juvenile forms. A study of present-day marsupial
erania, however, shows that no great change takes place with age.
It is of great interest to note that in the Native Bear (Phascolarctus) and
the Wombats (Phascolomys) small cheek-pouches have been recorded (Forbes,
1881, p. 182).
Had the cranium of Nototherium nmitchelli not been already known, the
special processes of Huryzygoma would have appeared to be still more
remarkable. The evolution of the latter from true Nototheriums can be realised,
however, and another instance is afforded of the astounding variety to be found
amongst the Marsupialia.
74 MEMOIRS OF THE QUEENSLAND MUSEUM,
Euryzygoma may be defined as a specialised member of the Nototherian
group, and, to use Gregory’s terms, its large processes were a _ cenotelic
character related to special food-habits.
Mandibles—Unfortunately there is no mandible directly associated with
the cranium under review. De Vis (1887, p. 1065) described three examples,
one (No. 5489) being illustrated as the type. His diagnostic characters are
here reproduced :—
‘‘Tusk-like incisors well developed; premolar small, sub-triangular,
unilobate; inlet of dental canal much behind postmolar angle and much above
alveolar level; exterobasal ridges of molars imterrupted: talons anterior and
posterior well developed: molars gradually enlarged: inner symphysial curve
opposite hinder lob of m?.”’
A full description of the three examples follows in the original paper.
In the following year, De Vis (1888) illustrated the upper subtriangular
premolar of ‘‘Nototherium dunense,’’ previously referred to in this paper as
a heautotype.
Text-fig. 1.—EuryzyGOMA DUNENSE. Reconstruction of cranium ; one-fifth natural size.
Drawn by O. W. Tiegs.
Taxonomic.—In classifying the Marsupialia of to-day the premolar is
recognised as the most important tooth, and that tooth alone is sufficient to
establish the identity of many species. The evidence of divergent premolars is
thus not to be lightly put on one side. Lydekker (1889) thought that the large
A NEW GENUS OF FOSSIL MARSUPIALS.—LONGMUAN.
Sa |
complex premolar on the right side of Macleay’s ‘‘Zygomaturus trilobus’’ was
an abnormality or was incorrectly associated with the cranium. Seott (1915,
p. 10) also quotes a remark from him which shows that Lydekker did not
consider this oval tooth as characteristic of Nototheriwm. But Seott’s illus-
trations and descriptions show that. this large tooth is associated with the
specimen described by him as Nototherium tasmanicum and with N. mitchelli.
Glauert (1912) also figured (Plate VI, fig. 10) a West Australian tooth of a
somewhat similar character. Amongst our specimens are three loose teeth of
this type and an additional one is in situ in a maxilla with the rest of the molar
series. It should also be noted that one of the teeth figured as the type of
Procharus celer by De Vis is of interest here, as pointed out by the writer when
dealing with this material (Longman, 1916, p. 86).
In response to my request, Mr. Charles Hedley, Australian Museum, has
kindly forwarded a note on the premolars of Macleay’s ‘‘Zygomaturus trilobus,”’
as observed by Messrs. Thorpe & Troughton, of the Museum staff. This states
that both teeth are firmly 7m situ, the right one being nearly complete, but that
“on the left side is badly fractured.’’
This clearly shows that the large oval premolar is not an abnormality,
but a definite characteristic of Macleay’s ‘‘Zygomaturus trilobus.’’ It certainly
follows that all crania with triangular premolars are generically distinct.
Although Nototherium inerme, first described by Owen (1845, p. 231) as
indicating a distinct genus ‘‘by a total absence of incisors,’’ has page precedence
(a single page!) over N. mitchelli, the rules of international zoological nomen-
clature give latitude for the selection of the second species as the type.
Nototherium mitchelli is certainly the ‘‘best deseribed, or best figured, or best
known’’ of the three species recorded by Owen from mandibular types.
Notwithstanding the discrepancies between the views of Owen, Lydekker, and
De Vis as to the premolars of Nototherium, Lydekker’s action in taking
N. mitchelli as the typical species should be confirmed, and that species is
hereby selected as the type of Nototherium, sensu stricto. The cranium described
by Macleay thus becomes a heautotype, and Simoprosobus De Vis (1907) is a
synonym. ~~
(Nx ay é ( EA A YY A
ad
Fig. 1—SyYNODUS HOULTI sp. nov. Holotype, 203 mm. long.
Fig. 2.—SyNopus SIMILIS sp. nov. Holotype, 185 mm. long.
A. R. McCulloch, del.
Face page 165.
QUEENSLAND FISHES.—McCULLOCH. 165
gg. Atlantic Ocean species .. ~ we 26 ic synodus and a‘lanticus.
ff. L. lat. 68.
i. 10 anal rays; light-coloured with dark cross-bars .. = .. dermatogenys.
ii. 8 anal rays; black with dark cross-bars .. Se a * lacertinus,
ee. 34 scales between lateral line and dorsal fin.
j. 11 dorsal and 10 anal rays: snout slightly longer than broad; eye about
44 in the head =: oe A ae Fe a ee katanus.
jj. 13 dorsal and 8-9 anal rays.
k. Snout as broad as long; dorsal and caudal fins spotted, gill-membranes
plain oe Pe = ME oh SE Be a indicus.
g; dorsal and caudal fins plain, upper pari of gill-
membranes with black blotches a. a a se similis.
kk. Snout broader than long ;
ce. Anal fin with 11 or more rays .. acu'us, altipinnis, saurus, scituliceps, jenkinsi,
fetens, and lucioceps.
The characters of S. erythreus Klunginger, from the Red Sea, are
unknown to me.
SYNODUS JAPONICUS Houttuyn.
Cobitis japonica Houttuyn, Verh. Holl. Mat. Har!em., xx, 1782, p. 450.
Salmo variegatus Lacepéde, Hist. Nat. Poiss., v, 1803, p. 157.
Saurus varius Ginther, Brit. Mus. Cat. Fish., v, 1864, p. 395—part.
Synodus japonicus Jordan & Herre, Proc..U.S. Nat. Mus., xxxil, 1907, p. 517—synonymy.
A single specimen 131 mm. long, from Murray Island, Torres Strait, enables
“me to add this species to the Australian list. Nine others from Lord Howe
Island and one from Amboyna agree in having the postoral portion of the cheek
naked as in S. dermatogenys Fowler, but have eight or nine anal rays instead of
ten as in that species. The two species are evidently very similar.
SYNODUS HOULTI sp. nov.
(Plate VIII, fig. 1.)
Br.dd; Dal2: A.9: P 135 V083.C. 19. Blatt. 59 =, tr. 44/7- Depth ofthe
body before the ventrals less than its breadth, and 6-5 in the length to the hypural
joint; head 3-3 in the same. Eye 1-8 in the snout, and 8-1 in the head;
interorbital width 1-1 in the eye. Third dorsal ray 2-4, and pectoral fin 3 in
the head.
Body subeylindrieal, a little broader than deep, tapering backwards. Snout
pointed, its width at the base distinctly greater than its length; jaws subequal.
Nostrils close together, nearer the eye than the end of the snout; the opening
of each is quite small, and the first has a posterior cutaneous lobule. Eye with
very narrow adipose membranes anteriorly and posteriorly. Interorbital space
concave. Cranium with some radiating ridges behind the eye and across the
occiput. Mouth oblique, the premaxilla extending about two eye-diameters
beyond the eye. Preopercular margin rounded, with a membranous border
bearing mucigerous canals. Opereculum unarmed, with a membranous border.
166 MEMOIRS OF THE QUEENSLAND MUSEUM.
A single row of fixed, spaced, and compressed teeth along the outer edge of
each premaxillary bone is covered by the lips; directly inside this row is another
of longer, more numerous, and depressible teeth which are exposed when the
mouth is closed. Mandibular teeth similar, in three rows, the outer row smallest
and fixed, the other two depressible and the innermost largest. The symphyses
of both jaws are toothless, but there are one or two larger teeth on each side of
that of the mandible. A long band of depressible teeth on each palatine bone,
arranged in four rows; the innermost teeth are largest, and they increase in
size forwards, and are enlarged anteriorly. Tongue covered with large
depressible teeth anteriorly, and a band of smaller ones posteriorly.
Body covered with cycloid scales, which have broad membranaceous borders.
There are about sixteen rows between the occiput and the dorsal fin. Lateral
line straight from the shoulder to the caudal pedunele; its scales are not raised,
and are scarcely differentiated from the others. Five or six rows of scales on
the cheeks, and a few on the upper’ portion of the operculum. ‘Some rather
elongate scales are present both above and below the base of the pectoral and
above that of the ventral; an enlarged pinnate scale on the base of each caudal
lobe.
Origin of dorsal fin slightly nearer the adipose dorsal than the tip of the
snout; the third and longest ray just reaches the base of the last when adpressed ;
the two anterior rays are simple, and the last is double. Adipose dorsal finlet
inserted above the middle of the anal. All the anal rays are simple, and the
last is double. Pectoral fin short, not nearly reaching the vertical of the first
dorsal ray. Ventrals inserted about midway between the verticals of the pectoral
and dorsal origins; the inner and outer rays are simple and the others are bifid;
the sixth ray is longest, and reaches about half its distance from the anal origin,
and to the vertical of the last dorsal ray. Caudal forked.
Colour-markings.—Greyish on tle back, the sides and lower surfaces white.
Some very ill-defined cross-bars on the back, the most prominent of which is at
the base of the tail, and some darker markings above the lateral line. Upper
surface of the head with vermiculating grey lines which extend onto the lips
near the end of the snout. Fins without markings.
Described and figured from the holotype, 203 mm. long from the snout to
the end of the middle caudal rays.
Locality.—Near the Capricorn Group, Queensland, 25-30 fathoms.
This species differs from SN. similis in having a much smaller eye, 43 instead
of 34 supralateral scales, and small instead of large nostrils. It also lacks the
dark marking on the upper portion of the gillmembranes. It is nearer
S. japonicus Houttayn, but has a much smaller eye and less rugose cranium,
and the characteristic dark body markings and spots on the fins of that species
are wanting in S. houlti.
_—
QUEENSLAND FISHES.—McCULLOCH. 167
SYNODUS SIMILIS sp. nov.
(Plate VIII, fig. 2.)
Br. 15; D.13; A.9; P.13; V.8; C. 19; L. lat. 58; L. tr. 33/6. Depth of the
body before the ventrals slightly less than its breadth, and about one-seventh of
the length to the hypural joint; head 3-5 in the same. Eye 1-2 in the snout, and
5-1 in the head; interorbital width 1-3 in the eye. Third dorsal ray 1-8, third
anal ray 3-1, and pectoral fin 2-1 in the head.
Body subeylindrical, a little broader than deep. _Snout pointed, its width
at the base distinctly greater than its length; jaws subequal. Nostrils close
together, nearer the eye than the end of the snout; each has a rather large
opening, and the first has a cutaneous lobe posteriorly. Eye with narrow adipose
membranes anteriorly and posteriorly. Interorbital space a little concave.
Cranium with some radiating ridges behind the eye and across the occiput.
Mouth oblique, premaxilla extending about two-thirds of an eye-diameter beyond
the eye. Preopercular margin rounded, without a free edge, with numerous
mucigerous canals extending from it onto the operculum; operculum unarmed,
with a broad membranaceous border.
A single row of fixed, spaced, and compressed teeth along the outer edge
of each premaxillary bone is covered by the lips; directly inside this row is
another of more numerous, longer, and depressible teeth which are exposed when
the mouth is closed. Mandibular teeth similar, in three rows, the outer row
smallest and fixed, the other two depressible and the innermost largest. The
symphyses of both jaws are toothless, but there are a few enlarged teeth on each
side of that of the mandible. A long band of depressible teeth on each palatine
arranged. in four rows; the innermost teeth are largest and they increase in
size forwards and ‘are enlarged anteriorly. Lingual teeth depressible in about
five rows anteriorly, the outermost of which are largest; these are followed by
a band of small teeth.
Body covered with cycloid scales which have broad membranaceous
borders. There are about fifteen between the oceiput and the dorsal fin. Lateral
line straight from the shoulder to the caudal pedunele; its scales are not keeled,
and scareely differentiated from the others. About five transverse rows of
eycloid scales on the cheek, and two on the upper portion of the operculum. A
few rather elongate scales both above and below the base of the pectoral fin, and
above that of the ventral; an enlarged pennate scale cn the base of each caudal
lobe.
Origin of the dorsal fin almost midway between the end of the snout and
the adipose dorsal; the third and longest ray reaches a little beyond the base of
the last when adpressed. The two anterior dorsal rays are simple, and the last
is double. Adipose dorsal inserted above the middle of the anal fin. The latter
is short and composed of simple rays; the last is double. Pectoral fin short, not
quite reaching the vertical of the first dorsal ray. Ventrals inserted between
the verticals of the pectoral and dorsal origins; the inner and outer rays are
168 MEMOIRS OF THE QUEENSLAND MUSEUM.
simple, and the others are bifid; the sixth ray is longest and reaches almost half
its distance from the base of the last anal ray, and well beyond that of the
dorsal. Caudal forked. '
Colour-marking.—Greyish on the back, the sides and lower surface white.
Ill-defined grey markings form indefinite cross-bars and about eight imperfect
rings on the lateral line. Grey lines form narrow interrupted rows along each
series of scales on the upper half. Head with grey spots on the opereles, and
the gill-membranes are ornamented with two large black spots on each side
above the operculum. Fins without markings.
Deseribed and figured from the holotype, 185 mm. long from the snout to
the end of the middle caudal rays.
This species is apparently very similar to S. indicus Day, but its vertical
fins are without markings, and the gill-membranes are ornamented with a black
blotch which is not present in the Indian species. According to Day’s deserip-
tion the ventral fins of N. andicus are shorter than in S. similis, the snout is as
long as broad instead of broader than long, and the adipose membranes around
the eye of S. similis are said to be wanting in NS. indicus. 8S. similis differs from
kaianus Giimther in the number of dorsal and anal fin-rays, and in the
proportions of the snout.
Locality—Near the Capricorn Group, Queensland, 25-30 fathoms.
Famity APOGONIDA.
APOGON BREVICAUDATUS Weber.
Apogon brevicaudatus Weber, Notes Leyden Museum, xxxi, 2, 1909, p. 158, and Siboga Rept.,
Fische, lvii, 1913, p. 232. pl. viii. fig. 3.
Amia berthe Ogilby, Ann. Qld. Mus. x, 1911, p. 47, pl. v, fig. 1, and Mem, Qld. Mus. ree (Be
p. 50.
A fine example 112 mm. long, agrees in all details with Weber’s figure of
this species, its markings being even better defined than in his illustration. A
comparison of it with a smaller example of A. berthw, which was received by the
Australian Museum from Mr. J. D. Ogilby, proves the two to be synonymous;
the vertical bands shown in Ogilby’s figure disappear with age.
Localities —Capricorn Group, 25-30 fathoms; coll. Queensland State
trawler.
Wide Bay, Queensland; Australian Museum, exch. J. D. Ogilby.
Famity POMACENTRID.
The following key to the genera of this family is submitted as tentative
only. It is largely a compilation derived from various smaller keys published
by Giinther, Jordan and Evermann, and others.
a. Teeth fixed, extending along greater portion of free edge of each jaw.
b. Teeth conical or villiform, not compressed.
c. Preorbital with a strong spine directed backwards. 40 a sys PREMNAS.
cc. Preorbital without a strong spine.
QUEENSLAND FISHES.—McCULLOCH. 169
d. Atleast some of the opercles serrated.
e. All the opercles serrated .. ie a 38 = : .. AMPHIPRION.
ee. Only the preoperculum and sometimes the preorbital pe
f. 12-13 dorsal spines.
g. Less than 30 transverse series of scales... aie .. Dascy3uus.
gg. More than 30 transverse series of scales .. ve ore .. Leprpozyeus.
# 17 dorsal spines ae .- .s +. .. ACANTHOCHROMIS.
dd. None of the opercles ser eaap
h. Lateral line not developed on the tail .. ae we 2. CHROMIS.
hh. Lateral line almost complete .. us sic AF st AZURINA.
bb. Teeth more or less compressed.
:. Lips greatly thickened, fimbriate, and curled back over the snout .. CHEILOPRION.
vi. Lips normal.
j. Preoperculum serrated.
k, Spinous portions of dorsal and anal scaly like the soft portions; suboperculum
and interoperculum entire .. st aa af “ee POMACENTRUS,
1. Teeth biserial, truncate; soft dorsal short, often elevated; caudal deeply
forked wae oe ae os ae ae Subg. PoMacENTRUS.
. Teeth rounded ; preorbital notched; caudallunate Subg. PsrupOPOMACENTRUS,
ho
3. Teeth uniserial; snout and lower jaw scaly; spinous dorsal with membrane
incised and lobed .. a ss oe -.. Subg. PARAPOMACENTRUS.
4. Snout and lower jaw naked ... ay 3 .. Subg. AMBLYPOMACENTRUS,
5. Snout scaly, lower jaw naked; teeth uniserial; membrane of spimous dorsal not
notched an age ce - he er Subg. EUPOMACENTRUS,
6. As above, but membrane of spinous Seven deeply notched : a
Subg. BRACHYPOMACENTRUS,
kk, Spinous portions of dorsal and anal naked; suboperculum and interoperculum
serrated ae Sia ar fs = be srg sa ae, - DAWA?
jj. Preoperculum smooth.
1. 30 or less transverse series of scales.
m. Suborbitals adnate to the cheeks.
n. Suborbitals completely covered by scales He oF .. NEXIDARIUS
nn. Suborbital scales not well formed we ts ae: fe NEXILOSUS,
mm. Suborbitals not adnate to the cheeks.
o. Teeth emarginate, in a single row 5 sae at . .GLYPHISODON,
1. Preorbital very narrow, snout rather acute; 12 anal rays ay,
Subg. GLyPHIsODON.
2. Preorbital broad, snout blunt, 10 anal rays .. Subg. Eucuistopwts.
3. Lower pharyngeals quadrate ae -. Subg. HemMicLyPHIDODON.
4. Scales above lateral line in 1-2 rows .. Subg. AMBLYGLYPHIDODON.
oo. Teeth not emarginate.
p. Teeth in two rows .. as ee oe ae .. CHRYSIPTERA,
pp. Teethinone row .. oe oe at ag .. HYPSYPOPS.
il, More than 30 transverse series of scales oe A a pee .. PARMA.
aa. Teeth uniserial, movable, those of lower jaw confined to front portion.
q. Vertical fins not elevated, caudal simply lunate .. als ade ie AZURELLA.
qq. Vertical fins much elevated, caudal lobssfaleate .. ae ie MICROSPATHODON,
=
~!
=)
MEMOIRS OF THE QUEENSLAND MUSEUM.
Genus DAYA Bleeker.
Daya Bleeker, Verh. Holl. Mij. Haarlem, 1877, p. 71 (Pomacentrus jerdoni Day). Id. Weber,
Siboga Fische, lvii, 1913, p. 343.
This genus differs from Pomacentrus in having no scaly sheaths at the
bases of the spinous dorsal and anal fins, and the membrane between the spines
naked instead of sealy. The edges of the subopereulum and interoperculum are
finely serrated like the preoperculum, and the suborbital bones are small and
inconspicuous; operculum with two spines. The anterior teeth of each jaw are
enlarged, and, though compressed, have more or less rounded points; the lateral
teeth small. Snout and mandible naked. Caudal fin forked. Scales in about
thirty rows. Membrane of spinous dorsal scarcely incised and without distinct
lobes.
DAYA JERDONI Day.
(Plate IX, fig. 1.)
Pomacentrus jerdoni Day, Proc. Zool. Soc. Lond. 1873, p. 237; and Fish. India, 1877, p. 383
pl. Ixxx, fig. 7.
Daya jerdoni Weber, Fische Siboga Exped., lvii, 1913, p. 344.
Pomacentrus dolii Macleay, Proc. Linn. Soc. N.S.Wales vi, 1881, p. 65, pl.i, fig. 1.
D. xiii/13; A.ii/14; P.18; V.i/5; C.15; L. lat. 19; 30 scales between the
origin of the lateral line and the hypural joint, 5 between the origin of the
dorsal and the lateral line, and 10 more to the origin of the anal.
Depth before the ventrals 2-6 in the length to the hypural joint; head 3-5
in the same. Eye 3-4 in the head and a little longer than the snout, which is
4.2 in the head; interorbital width a little greater than the length of the eye,
3-1 in the head. Sixth dorsal spine 2-1 in the head, longer than the last, but
‘much shorter than the ninth ray; pectoral 1-2 in the head.
3ody longer than is usual in Pomacentrids, with the upper and lower
profiles equally curved from the obtusely conical snout. Suborbital bones small
and inconspicuous, the first wider than the others and hinged with the preorbital ;
free edge of the preorbital notched and feebly serrated. Hinder edge of the
preoperculum serrated, as is that of the suboperculum, and the interopereculum
also has a few serrations. Operculum armed with two flat spines, the lower of
which is the larger. Each jaw with about six large flattened teeth in a row
anteriorly, and a row of much smaller ones on each side; palate toothless.
Nostril a simple opening on each side.
Scales ctenoid with narrow ciliated borders. _They extend forward on the
upper surface of the head to before the middle of the eye, and cover the whole
head with the exception of the snout, lips, and mandible. There is an enlarged
scale at the base of each ventral fin, and two others are present between those
fins. Seales cover the bases of the pectoral fins, and extend up between the
dorsal, anal, and caudal rays, but the spinous portions of the dorsal and anal
are naked. Lateral line formed by a broad canal, with two or three pores
opening on each scale; the scales of the median series on the caudal pedunele
also bear minute pores.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, PLaTE IX.
psgh® 34
elds
y
Fig. 1—DaAya sERDONI Day. A specimen 108 mm. long.
Fig. 2.—DAYA JERDONI, var. FUSCA, var. nov. Holotype of variety, 99 mm. long.
A. R. McCulloch, del.
Face page 170.
QUEENSLAND FISHES.—McCULLOCH. 171
Median dorsal spines highest, the others decreasing slightly in length
backwards; the rays increase again to the ninth, which is longer than the longest
spine. The margin of the dorsal fin is slightly sinuous between the spines, and
obtusely pointed posteriorly. Anal similar in form to the soft dorsal. Pectoral
rounded, the fifth upper ray longest. Ventrals rounded, without produced rays,
and reaching backwards to the vent. Caudal forked.
Colour.—Yellow, with blue spots, the fins darker. The body is much
lighter than the fins after preservation, and the scales have olivaceous markings
which combine to form the pattern illustrated in the accompanying figure. Two
dark stripes cross the preorbital, one extending under the eye, and the cheeks
und opercles are marked with dark spots and short lines. A large blackish spot
at the upper base of the pectoral. Vertical fins slate-coloured in formaline;
the dorsal and anal have narrow white borders followed by a very narrow
darker line, and there is a blackish spot between the anal spines; a dark horizontal
line extends along the soft portion of each. Caudal with irregular rows of grey
spots. Pectorals and ventrals whitish.
Described and figured from a specimen 108 mm. long, which was taken
by the Queensland State trawler around the Capriccrn Group, Queensland, in
25-3C fathoms. Two others taken with it exhibit some little variation in the
degree of colour-marking on the fins, and one bears indications of blue spots
on each scale; the extreme outer rays of the caudal! fin may also be light-coloured.
Synonymy—An examination of the two typical specimens of Pomacentrus
dolii Macleay, in the Macleay Museum, proves them to be similar in all details
to the example deseribed above. They were found in a large Dolium shell in
Port Jackson, and were doubtless mere stragglers southward from the warmer
waters of Queensland.
DAYA JERDONI, var. FUSCA, var. nov.
(Plate IX, fig. 2.
A series of twenty-one specimens, 59-104 mm. long, which were trawled
by the F.L.V. *‘Endeavour’’ in Queensland waters, includes several specimens
which cannot be separated from those described above; also others which differ
somewhat in both form and colour, and others which are intermediate between
the two. In a specimen 99 mm. long, which is figured on the accompanying
plate, the depth at the ventrals is 2-4 in the length to the hypural joint. The
body is brown lke the vertical fins, and cach seale bears a basal darker (blue)
spot. The dark lines on the dorsal and anal are wanting in this specimen, but
are indicated in another somewhat larger example. The outer rays of the caudal
fin ave whitish like the margins of the dorsal and anal.
Although the two forms illustrated differ in their general appearance,
they are evidently specifically identical, since other specimens are intermediate
between them, and can be as readily assigned to the one as the other.
172 MEMOIRS OF THE QUEENSLAND MUSEUM.
Localities—Twenty miles N.N.E. of Double Island Point, Queensland;
3U fathoms.
Thirteen miles 8.E. of Cape Capricorn, Queensland; 13 fathoms.
Four to five miles 8.E. of Bustard Head Light, Queensland; 11-16 fathoms.
Ten miles N.W. of Bustard Head Light, Queensland; 14-17 fathoms.
Three to seven miles N.W. of Hervey Bay fairway buoy, Queensland;
9-11 fathoms.
Twelve miles N.E. of Bowen, Queensland; 19-25 fathoms.
Port Jackson, New South Wales: types of P. dolii Macleay.
GENUS GLYPHISODON Lacepéde.
GLYPHISODON PALMERI Ogilby.
(Plate X, fig. 1.)
Glyphisodon palmeri Ogilby, Mem. Qld. Mus., ii, 1913, p. 87, pl. xxii, fig. 2.
As the original figure of this species is imperfect in several details, I have
figured another specimen, 135 mm. long from the snout to the end of the middle
caudal rays, which is well preserved though it has lost many of its scales.
Fourteen specimens, 42-135 mm. long, show that the disposition of the transverse
bands and the other characters of the species are very constant, and indicate that
G. palmeri is a valid species.
Localities —This species is evidently not uncommon on the Queensland
Coast. I have speared it among coral on the surface of the reef at Masthead
Tsland, while the Queensland State trawler has taken it in 25-30 fathoms near
the Capricorn Group. It has also been taken in a hauling net on a beach at
Cape Bedford and in a reef pool near-by on Two Isles.
Capricorn Group, surface to 30 fathoms. Caloundra. Cape Bedford. Two
Isles, off Cape Bedford. Torres Strait. Sweers Island, Gulf of Carpentaria.
Famity SCORPANIDA.
Genus PARACENTROPOGON Bleeker.
? Hypodytes Gistel, Naturg. des Theirreichs, viii, 1848, Genotype uncertain, vide Jordan, Gen.
Fish., ii, 1919, p. 235.
Paracentropogon Bleeker, Versl. Akad. Amsterdam (2), ix, 1876, p. 297 (Apistus longispinis.
Cuv. & Val.).
Daia Ogilby, Proc. Roy. Soc. Qld. xviii, 1903, p. 9 (Centropogon indicus Day).
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, PuaTE X.
Fig. 1.—GLYPHISODON PALMERI Ogilby. A specimen 135 mm. long.
Fig. 2.—PARACENTROPOGON VESPA Ogilby. Holotype?, 87 mm. long.
A. R. McCulloch, del.
Face page 172.
~~
—
QUEENSLAND FISHES.—McCULLOCH. 173
PARACENTROPOGON VESPA Ogilby.
(Plate X, fig. 2.)
Paracentropogon vespa Ogilby, New Fish. Qld. Coast, 1910, p. 116.
Br. 7; D. xiv/8; A.iii/5; P.10; V.1/4; C.18; 22 tubular pores. on the
lateral line between its origin and the hypural joint, and 1 on the base of the
caudal.
Depth before the ventral fins almost 3 in the length to the hypural joint ;
head, to the end of the opercular lobe, 2-7 in the same. Eye 3-7 in the head, and
slightly longer than its distance from the end of the snout. Interorbital space
much less than the length of the snout, 1-5 in the eye. Third dorsal and anal
spines equal in length to the anterior dorsal and anal rays, and 1-6 in the head.
Pectoral fin 1-1 in the head.
Profile very oblique from the snout to the origin of the dorsal fin, with
a depression above the premaxillary processes.. Interorbital space with two bony
ridges on the upper end of which are two short tentacles; behind these is a
median ridge before the first dorsal spine. Supracrbital ridges very prominent,
each with a short tentacle. Another ridge extends backward from the upper
margin of the orbit, and there are two short ridges between the eye and the
origin of the lateral line. Mandible projecting very slightly beyond the pre-
maxillaries. Mouth oblique, maxilla extending backward to below the anterior
third of the eye. Anterior nostril situated just behind the middle of the snout,
with a short dermal tentacle; posterior nostril preceded by a large open pore,
and placed just in front of the eye. Preorbital with two spines directed back-
ward, the anterior short, the posterior large and reaching to below the hinder
portion of the eye. Suborbital bone flat and unarmed. Preoperculum with
five spines, the uppermost of which extends well beyond, the middle of the
operculum; the others are small and scarcely project beyond the rounded
preopercular edge. Operculum with two diverging ridges which end in obtuse
spines; its posterior portion forms a narrowly rounded lobe. Minute pores open
on the preopercular margin between each spine, while others are placed sym-
metrically on the head and neck above and behind the eye, and on the cheek,
preorbital, and mandible. A broad band of minute teeth on each of the pre-
maxillaries and one on each side of the mandible; a A-shaped patch of similar
teeth on the vomer, and a patch on the end of each palatine bone. In the specimen
described and figured there is a rounded patch of teeth attached to the palatal
membranes in the angle of the vomerine patch, but this is not developed in other
specimens. Pseudobranchize well developed; no slit behind the last gill-arch.
The back is highest at the base of the fifth dorsal spine, behind which it *
descends evenly to the caudal pedunele. Head, anterior portion of the back.
pectoral region, and abdomen naked, but imperfect scales are present beneath
the skin on the greater portion of the sides and caudal peduncle. Lateral line
extending in an oblique line from the shoulder to the middle of the caudal
pedunele; it forms a complete canal from which minute tubules open obliquely
upwards.
174 MEMOIRS OF THE QUEENSLAND MUSEUM.
First dorsal spine inserted over the middle of the eye; the first five spines
are widely separated though connected by membrane; the third is longest, is
but little longer than the last, and is as long as the anterior dorsal and anal
rays. Soft dorsal rounded, formed of branched rays, of which the last is broadly
united with the caudal peduncle by membrane. Anal originating below the
twelfth dorsal spine, its third spine as long as that of the dorsal; soft portion
rounded, the last ray united ‘with the caudal peduncle. Pectoral formed of
branched rays, the median of which are longest and almost reach the vertical
of the anal origin. Ventrals inserted behind the bases of the pectorals, each
with a strong spine and four branched rays, the longest of which reach the vent.
Caudal rounded, with ten branched rays.
Colour-marking—Light brown after preservation, variegated with darker
brown marblings; these are not well defined in the specimen described so they
have been copied from another taken with it and of the same size in the
accompanying figure. A darker patch is present on the dorsal fin between the
fifth and eighth spines, and the whole of the fin is marked with oblique wavy
cross-bands. A light spot is situated above the lateral line over the posterior
half of the pectorals. Pectorals and caudal with darker ecross-bars, the basal
portion of the latter whitish. Ventrals blackish with a light patch near the
middle of the spine and anterior ray.
Deseribed and figured from a specimen in the ‘‘ Endeayour’’ collection,
87 mm. long, of which both the length and data differ from those given by
Ogilby. No specimen in the collection agrees exactly with his description, which
was prepared under unfavourable conditions on board the ‘‘Endeavour,”’ and
it is not improbable that this specimen is the actual type of the species.
Affinities.—This species is very similar to P. longispinis, with a small
and bleached example of which I have been able to compare it, though its third
dorsal spine is proportionately shorter. In all other structural details the two
appear to be similar, but colour differences may be found which will maintain
them as separate species.
P. vespa differs from P. rubripinnis in having much finer teeth, lower
anterior dorsal spines, better developed scales, and in colouration. The black
dorsal spot is largely restricted to the dorsal fin instead of descending onto the
back, and the ventrals are dark instead of light m colour.
Localities—Platypus Bay, Queensland; 7-9 fathoms. ‘‘ Endeavyour’’
collection, two specimens.
Rat Island, Port Curtis, Queensland; coll. A. R. McCulloch. One specimen
with abnormal dorsal spines, but exhibiting all the other characters shown in the
figure.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, PLATE XI.
Fig. 1.—Lr10craAniuM scorpio Ogilby. Holotype, 75 mm. long.
Fig. 2.—Erosa ErOsA Cuvier & Valenciennes. Holotype of E. mrmEA Ogilby, 80 mm. long.
A. R. McCulloch, del.
Face page 175.
Ou
QUEENSLAND FISHES.—McCULLOCH. 17
Genus LIOCRANIUM Ogilby.
Liocranium Ogilby, Proce. Roy. Soe. Qld., xviii, 1903. p. 24 (Orthotype L. prepositum Ogilby).
Jd, McCulloch, Biol. Res. Endeavour, iv, 4, 1916, p. 195.
a. Diameter of eye much greater than width of interorbital space; gill-rakers of first arch
short, thick, and rounded .. Hs : preposiium.
aa, Diameter of eye much less than width of ‘interdibitar space ; “a: Zo of first arch
long, slender, and flattened ae ae ys Ale ae ve ar scorpio.
LIOCRANIUM SCORPIO Ogilby.
(Plate XI, fig 1.)
Paracentropogon scorpio Ogilby, New Fish. Qld. Coast, 1910, p. 115,
Br.7; D. xiv/6; A. 11/7; P.6/8; V.i/4; C.13; 22 to 24 tubules on the
lateral line between the shoulder and the Higteal joint.
Depth before the ventrals 3 in the length to the hypural joint; head, to
the end of the opercular lobe, 2-6 in the same. Eye 4-3 in the head, slightly
shorter than the snout, which is 3-5 in the head. Interorbital width greater than
the length of the snout, 3-1 in the head. Third dorsal spine and third anal spine
2-4 in the head, and a little shorter than the fourteenth dorsal spine. Pectoral
1-1. caudal 1-2 in the head.
Snout with a prominent hump caused by the posterior processes of the
premaxillaries ; interorbital space broad and slightly convex, with a median bony
ridge which is bifurcate anteriorly. The back is highest at the bases of the
fourth and fifth dorsal spines, and thence descends evenly to the caudal peduncle.
Mandible projecting beyond the premaxillaries when the mouth is closed. Mouth
oblique; maxilla broad, its hinder margin oblique, the upper posterior angle
reaching a trifle beyond the vertical of the hinder margin of the eye. Eye with
prominent bony margins. Nostrils in the posterior half of the snout, separated
by a narrow interspace, the anterior with a dermal lobe. Preorbital with two
spines directed backward, the posterior large and reaching beyond the middle
of the eye. Suborbital ridge unarmed. Preopereulum with five spines, the upper-
most of which is largest; the others decrease in size downwards, and the lowest
is minute. Operculum a weak flat bone without ridges or spines, and terminated
by a pointed lobe. Minute pores open cn the preopercular margin between each
spine, while others are placed symmetrically on the head and neck above and
behind the eye, on the preorbital and mandible. A band of villiform teeth in
each jaw which is interrupted at the symphysis; vomer with a curved patch of
similar teeth, palatines toothless. No slit behind the last gill-arch; about fifteen
gill-rakers on the lower limb of the first gill-arch, the posterior of which are
slender and flattened and about half as long as alte eye.
Skin almost naked, only rudimentary scales appearing on the sides of the
body and caudal peduncle. Lateral line extending obliquely from the shoulder
to the middle of the caudal peduncle, and terminating on the base of the tail;
it forms a complete canal from which small tubules extend obliquely upwards and
backwards.
176 MEMOIRS OF THE QUEENSLAND MUSEUM.
First dorsal spine inserted over the anterior third of the eye; the first
four spines are widely separated but connected by membrane, and the third is
but little shorter than the last though distinctly longer than those immediately
before and after it. The first dorsal ray is as long as the last spine, and is
unbranched; the four following rays are bifurcate, while the last is simple and
joined by membrane to the caudal peduncle. Anal fin originating below the
thirteenth dorsal spine; its spines increase in length backward, and the third
is about as long as the last dorsal spine but distinctly shorter than the succeeding
rays. Soft portion of the anal rounded; its two outer rays are simple and the
others bifurcate, and the last is united with the caudal peduncle by membrane.
Pectoral rounded, its middle rays longest and reaching to the vertical of the anal
origin ; five of the upper rays are bifurcate, and the eight lower ones are simple
and thickened. Ventrals inserted behind the bases of the pectorals, each with
a strong spine and four weak rays, of which three are bifurcate; the longest
does not nearly reach the vent. Caudal rounded, with eleven branched rays.
Colour.—Light brown changing to whitish below, with a paler stripe along
the back at the base of the dorsal spines; some very indefinite darker .cross-bands
are apparent on the back which become more distinct on the caudal peduncle.
Some irregularly placed dark-brown spots with lighter edges are scattered over
the neck and back above the lateral line, and several lighter ones occur on the
scapular region. Upper surface and sides of head closely covered with rounded
brown spots with lighter edges, some larger ones of which are situated above the
operculum and one at the postero-inferior angle of the eye. Lips erossed by
pale-brown narrow bands. Spinous dorsal irregularly marbled with brown, and
crossed by oblique darker-brown bands which become narrower and more
numerous on the soft portion, and coalesce so as to leave only a series of white
spots on the rays. Anal with a few iight-brown cross-bars on the rays. Pectoral
and caudal fins barred with dark brown. Ventrals whitish.
Described and figured from the holotype of the species, which is 75 mm.
long from the snout to the end of the tail. It differs in several small details
from Ogilby’s description owing to the fact that the latter was written on board
the ‘‘ Endeavour,’’ where the conditions rendered accuracy impossible.
Variation—A second specimen 79 mm. long differs in having the third
dorsal spine distinctly longer than the last; the cross-bands on the back are more
distinct, and the spots of the head extend on to the lower portion of the sides of
the body.
Head almost entirely concealed from above; deeply coneave, highly
polished and with small punctures between eyes. Antenne with third joint
thinner and slightly longer than second and fourth, sixth to tenth slightly
transverse. Prothorax about thrice as wide as long, sides strongly rounded and
“mereasing in width to base; surface uneven and with fairly large, shallow,
crowded punctures. Elytra rather wide, sides feebly dilated, each with three
discal coste, of which the two inner ones are long and almost as stout as the
sutural thickening, the other is finer, submarginal, almost as distant from the
second as the second is from the suture, commences ‘near the shoulder (this eon-
siderably thickened and slightly paler than the adjacent surface), and terminates
before the others; punctures dense, sharply defined and somewhat smaller than
on prothorax. Abdomen with first white segment slightly more than half the
length of the preceding one, the following segment considerably longer, with its
tip produced in middle. Legs thin, but rather short. Length, 6-5-7 mm.
Hab.—Queensland: National Park in November (H. Hacker).—Type,
C/2290 in Queensland Museum; cotype, J. 12282 in South Australian Museum.
The interocular space and the abdomen (except the two apical segments)
are of an intense black, the other black parts have a more or less brownish tinge;
the pale margins of the prothorax are very distinct, at the base from each side
the elevated part curves round and is directed obliquely inwards for a short
distance, but the base between has also a pale marginal strip. Structurally it
is close to A. brevis and A. olivier’: Mr. Hacker obtained twelve specimens, all
males.
ATYPHELLA SCINTILLANS Oll.
Mr. H. J. Carter sent for examination a pair of this species,
taken in cop. on Bunya Mountain. The female is almost entirely
pale (the antennx and elytra are slightly infuscated only), the
abdomen has a mottled appearance, as if the phosphorescent
material had been irregularly scattered through it (but no doubt
would look different on living specimens). The prothorax is
shightly larger than on the male, and has the sides and apex
conjoined to form an almost perfect semicircle, but the base is
somewhat sinuous, the elytra (each of which has two diseal costa) |
: are very short, being less than their combined width; they leave
eee | 2 six abdominal segments exposed and most of the preceding one.
Fic. 1—Atyphella
seintillans Ol).
COLEOPTERA—LEA. 199
TELEPHORUS VARIIVENTRIS sp. nov.
$ Flavous; antennx, palpi, most of elytra, knees, tibie, and tarsi black.
Densely clothed with short pubescence, golden on the pale parts, blackish on the
dark parts.
Head rather wide; with minute punctures. Antenne rather long,
moderately stout, joints after the fourth slightly decreasing in width, and slightly
inereasing in length. Prothoraz strongly transverse, disc uneven; with very
minute punctures. Elytra rather long, almost parallel-sided ; with crowded
rugose punctures, smaller about base than elsewhere, and with remnants of
feeble coste#. Apical segment of abdomen deeply notched. Legs moderately
long. Length (¢@ ), 7-5-10 mm.
Q Differs in having the head smaller, with shorter antenne, rather more
of the elytra pale, metasternum and abdomen (except apical segment, which is
not notched) black or infuscated, and more of femora dark.
Hab.—Queensland: South Johnstone River (H. W. Brown); Malanda
(Dr. E. Mjoberg).—Type, J. 12207 in South Australian Museum; cotype, C/2291
in Queensland Museum; others in Stockholm Museum.
In my table? would be associated with 7. rubriceps and T. rufiventris ; the
many species allied to 7. mastersi differ from it as follows:—T. mastersi and
T. froggatti have base of elytra and part of head black or blackish; 7. mossman
and 7. macrops have elytra entirely pale; T. palmerstoni has thicker and shorter
antenne, and base of elytra dark; 7. rufiventris has much less of elytra dark;
and T. rubriceps has basal joint of antenne distinctive in the male. On the
male the base of the elytra, for a distance about equal to the length of the
prothorax, is pale, but the sides are narrowly pale almost to the middle, or even
beyond it, and the suture is very narrowly pale also for a short distance.
TELEPHORUS ATRICORNIS sp. nov.
S Black and flavous. With rather sparse, ashen pubescence, and fairly
numerous, suberect, dark hairs.
Head wide and gently convex between eyes, strongly narrowed to base;
with dense minute punctures. Antenne moderately long, fairly stout, but
‘ becoming rather thin towards apex, third joint about half the length of fourth.
Prothorax slightly longer than wide, dise somewhat uneven, sides and suture
thickened. Elytra long, thin, and parallel-sided to near apex, sides and suture
thickened, a fairly distinct discal costa to beyond the middle; with crowded,
rugose punctures. Legs rather long and thin. Length (9d), 7-13 mm.
Q Differs in being rather more robust, head narrower across eyes and
wider at base, much less of muzzle pale, and legs slightly shorter.
Hab.—Queensland: Mount Tambourine (A. M. Lea). New South ‘Wales
(H. J. Carter) ; Dorrigo (W. Heron) ; Tweed River (R. Helms).—Type, I. 9240
in South Australian Museum; cotype, C/2292 in Queensland Museum.
2Lea, Trans. Ent. Soc. Lond., 1909, p. 113.
200 MEMOIRS OF THE QUEENSLAND MUSEUM.
At first glance apparently belonging to T. imperialis, with which it would
be associated in my table, but antenna decidedly shorter, thicker, and hairier,
the ninth and tenth joints no paler than the adjacent ones, apical joint of palpi
smaller, prothorax longer, elytra with a distinct costa on each, and the black
part at the base rounded posteriorly, instead of truncated. There are numerous
specimens of both species before me, and the differences noted are constant. On
both species the antenne are slightly thicker on the male than on the female.
On the male the flavous parts are much of the muzzle, a narrow space at apex
of prothorax, and more of its base (the basal portion advanced on the sides),
elytra (except for a small space at base and another at apex), parts of sterna,
much of under surface of abdomen, trochanters, and parts of coxe. On the
male the extreme base of the head is about half the width across the eyes, on the
female it is about two-thirds.
HETEROMASTIX PUSILLUS Boh.
Five specimens from Mount Tambourine, and one from the Queensland
National Park, probably represent another variety of this species; they differ
from Sydney (the type locality) ones, in being smaller (2-2-75 mm.), and in
having the legs flavous, except that the femora on most of them are partly
infuseated. Another, from Brisbane, in the Queensland Museum, measuring
2 mm., has much of the prothorax infuscated.
HETEROMASTIX PALLIPES Lea.
A specimen from the National Park, in the Queensland Museum, appears
to belong to this species, but is slightly larger (4 mm.) than the type, the apical
joint of its antenn:e is slightly stouter, and the two basal joints are dark on the
upper surface.
5 i WV i) E r G if 1 J K
\ \ \ 2 f \
Fic. 2.—Tips of antenne of species of Heteromastiz. A, B, C, spinicornis Lea; D, E,
melanocephalus Lea; F, G, castor Lea; H, I, pollex Lea; J, K, scutellaris Lea.
HETEROMASTIX SPINICORNIS sp. nov.
¢ Black, basal two-thirds of elytra flavous. Densely clothed with dark
pubescence, becoming golden on pale portion of elytra.
Head with dense and minute punctures. Antenne moderately long, first
COLEOPTERA.—LEA. 201
joint scarcely longer than third and at apex no wider, second about half the
length of third, third-ninth subequa! in length and apical width, tenth distinctly
wider and with a small spine at inner apex, eleventh longer than ninth and
tenth combined, base irregularly compressed. Prothorax more than twice as
wide as long, distinctly margined throughout. Elytra parallel-sided to near
apex; with crowded and small rugulose punctures, and vague remnants of coste.
Length, 6-5 mm.
Hab.—Queensland: Mount Tambourine (H. J. Carter).—Type, J. 11861
in South Australian Museum.
In my table would be associated with H. luridicollis, but distinguished
from that, as from all other species having the apical joints of antennz distorted,
by the black prothorax and bicolorous elytra. The eleventh joint of antenne
appears to alter in shape with the point of view; from one direction it appears
to have a subbasal notch and to be narrower than the tenth joint; on its under
surface it has a short longitudinal groove which appears to be continued on to
the tenth; the spine on the latter joint, although small, is quite distinct from
above.
Var.—Another male, from Mount Tambourine (taken by Mr. Hacker and
in the Queensland Museum), has the prothorax entirely pale, the black part
of its elytra is slightly advanced near the suture, and decreased towards the
sides; as a result it appears to have an almost circular outline. A female (taken
by Mr. Carter), also with flavous prothorax, differs from the male in the usual
particulars of antenne, legs, and abdomen.
HETEROMASTIX MELANOCEPHALUS sp. nov.
¢ Black; prothorax, basal two-thirds of elytra, tibie, parts of femora
and of tarsi, and three basal joints of antennex flavous. With moderately dense
pubescence, varying in colour with the derm.
Head gently convex between eyes (these rather small and prominent),
slightly depressed in front. Antennze moderately long, second joint very short,
tenth and eleventh distorted. Prothorax almost twice as wide as long, margined
throughout, the lateral margins somewhat thickened near apex. Elytra almost
parallel-sided to near apex; with dense and small, rugulose punctures. Length
($2), 44-5 mm.
Q Differs in having the head smaller with shorter and somewhat thinner
antennex, tenth joint similar to the ninth, and their combined length slightly
less than the eleventh, only about one-fourth of the elytra black, legs somewhat.
shorter, and abdomen not notched.
Hab.—Queensland : Bribie Island in August (H. Hacker and A. M. Lea). —
Type, I. 11862 in South Australian Museum; cotype, C/2293 in Queensland
Museum. ;
In my table would be placed with H. luridicollis, from the description of
which it differs in being smaller, head entirely black, legs not entirely pale, and
202 MEMOIRS OF THE QUEENSLAND MUSEUM.
in the bright flavous colour of its elytra, of which less of their tips are black,
The tenth joint of the antenne of the male is shghtly longer than the ninth, and
more produeed on one side (and curved on it) than on the other; the eleventh
is about as long as the ninth and tenth combined, closely applied to the latter
and with a narrow notch on one side near the base, the notch invisible from most
directions. Mr. Hacker and I obtained eleven specimens, but only one male.
HETEROMASTIX CASTOR sp. nov.
¢ Black; prothorax, parts of under surface of head, two basal joints of
antennae, tibix, and parts of femora flavous; tarsi and part of second joint of
antenne infuseated. With short, ashen pubescence.
Head with a shallow interocular depression. Antenne rather long and
not very thin, tenth and eleventh joints closely applied together. Prothorax
about twice as wide as long, sides slightly dilated near apex, but scarcely thickened.
Elytra almost parallel-sided to near apex; with dense and small subrugulose
punctures. Length, 3-75-4 mm. ;
Hab— Queensland: Brisbane (H. Hacker); Glen Lamington (Dr. E.
Mjoberg) —Type, C/2294 in Queensland Museum.
The tenth and eleventh joints are somewhat distorted but they are so
closely applied together that from some directions they appear to be simple; the
tenth, however, has a slight apical notch, in which is received a basal process
from the eleventh; it is wider than the ninth, shghtly longer on one side, and
more noticeably on the other; the eleventh is somewhat dilated at the base, and
the process received into the tip of the tenth may be regarded as a remnant of
a spine; its longest side is about twice the length of the tenth. So little of the
muzzle is pale (scarcely more than the labrum) that the species might be regarded
as belonging to ee of my table,® and there associated with H. gagaticeps, which
has the apical joints very different; but, regarding the muzzle as pale, it would
be associated with A. imitator, which is a smaller and more fragile species, with
thinner antenne, the eleventh joint of which is much thinner than the tenth.
HETEROMASTIX POLLUX sp. nov.
6 Black; prothorax, two basal joints of antennse, and knees flavous. With
short pubescence.
Head with two faint interocular impressions. Antenne rather long and
not very thin, two apical joints somewhat distorted. Prothorax and elytra as
described in preceding species. Length ( ¢@), 3-5-4 mm.
Q Differs in having the head smaller, antenne shorter, thinner, and
simple, and abdomen not notched.
Hab.— Queensland: Brisbane (H. Hacker). Type, C/2295 in Queensland
Museum ; cotype, 7. 12260 in South Australian Museum.
3Lea, Trans. Ent. Soc. Lcnd., 1909, p. 130.
COLEOPTERA.—LEA. 203
The black scutellum associates this species in my table with H. gagaticeps,
which is a larger species, with two apical joints of antenne somewhat different :
to the preceding species it is closer, but the tenth joint of antenne is different,
and the knees are the only pale parts of the legs; the following species has the
seutellum and middle tibize pale, and the apical joints are not quite the same.
The tenth joint on one side is distinctly ineurved, with its tip shghtly produced
beyond the base of the eleventh; on one side it is partly excavated for the recep-
tion of the base of that joint; the eleventh is slightly wider than the tenth, but
from some directions appears to be of the same width.
HETEROMASTIX MINOR sp. nov.
| ¢ Black; prothorax, scutellum, mesosternum, legs (tarsi slightly imfus-
cated), and two basal joints of antenne flavous. With short, ashen pubescence.
Head with a shallow interocular impression. Antennz not very long, two
apical joints somewhat distorted. Prothorax and elytra as described in H. castor.
Length, 3 mm.
Hab.—Queensland: Buderim Mountain (H. Hacker).—Type (unique),
C/2296 in Queensland Museum.
The eleventh joint of the antenne is slightly longer than in H. decipiens,
‘and is not quite simple; the tenth is much shorter, but at first glance the type
appears to be a small specimen of that species. It is shghtly smaller and with
antennx somewhat similar to those of H. castor and H. pollux, but the scutellum
is pale; this character would associate the species, in my table, with
H. pallipes, which has much longer antennz, with the terminal joints different.
The tenth joint is somewhat similar to the ninth, but is slightly wider at apex,
and a little lopsided; the eleventh also is lopsided and somewhat dilated at base,
but the two joints are so closely applied together that it is diffieult to see their
junctional parts.
HETEROMASTIX SCUTELLARIS sp. nov.
6 Black; prothorax, scutellum, mentum, mesosternum, legs (tarsi infus-
eated), and under surface of three basal joints of antenne flavous. With very
short pubescence.
Head with two faint longitudinal impressions in front, terminating pos-
teriorly in two faint interocular ones. Antenne long, two apical joints distorted.
Prothorax and elytra as described in H. castor. Length (dQ ), 3-75-4-5 mm.
Q Differs in having the head smaller, with shorter and simple antenne,
and abdomen not notched.
Hab.—Queensland: Mount Tambourine (A. M. Lea).—Type, J. 11867 in
South Australian Museum ; cotype, C/2297 in Queensland Museum.
The tenth joint of the antenne of the male is about the length of the
ninth, but longer on one side than on the other; the eleventh is constricted
204 MEMOIRS OF THE QUEENSLAND MUSEUM.
somewhat nearer the base than apex, and one side of the base is produced into a
short spine on to the shorter side of the tenth; the two joints, however, are so
closely applied together that it is difficult to see their junctional parts. In my
table would be associated with H. pallipes, but from the position in which the
hasal spine of the eleventh joint is visible, on that species, the apical portion of
‘he joint seems set at a tangent, very different from that of the present species;
H. frater has a pale scutellum, but its muzzle is also pale; H. minor is more
fragile, with thinner antennie, of which the two basal joints are entirely pale;
H. major is a larger species, with legs mostly dark; H. castor has the scutellum
black, legs mostly black, and two basal joints of antenne entirely pale; all these
species also differ, inter se, in the eleventh joint of antenne.
HETEROMASTIX TIBIALIS sp. nov.
¢ Black and fiavous. With rather dense, suberect pubescence.
Head rather large, with a shallow interocular depression. Eyes large and
prominent. Antenne long and thin, ninth joint slightly longer than tenth, and
shorter than eleventh. Prothorax about twice as wide as long, margined through-
out; with distinct, submarginal punctures. Elytra almost parallel-sided to near
apex; with dense and sharply defined punctures of moderate size, becoming
very small on tips. Front tibie dilated to apex and notched there, basal joint of
front tarsi strongly inecurved on one side. Length ( 69), 5-5-5 mm.
Q Differs in having the head smaller, with smaller and less prominent
eyes, antenne shorter, subapical segment of abdomen not notched in middle,
front tarsi only slightly thickened at apex and not notched, and basal joint of
front tarsi symmetrical.
Hab.—New South Wales: Dorrigo (W. Heron); Tweed River (A. M.
Lea).—Type, J. 11870 in South Australian Museum; cotype, C/2298 in Queens-
land Museum.
The black parts are the head (except part of its under surface), antenne,
palpi, metasternum, and abdomen; the apical sixth of elytra, apical half of
tibiz, and the tarsi, are more or less deeply infusecated or blackish. In the male
the antenne almost extend to the tips of the elytra; the notch, at the tip of its
front tibiz, has proceeding backwards from it a narrow groove on each side to
about the apical third (as if the tibia had been split), the tibia itself is some-
what produced on one side of apex. The general appearance is somewhat as in
some forms of H. luridicollis, but the antenne of both sexes are simple.
A male from Queensland (Gympie) appears to belong to this species,
but has the flavous parts brighter, almost the whole of the apical half of elytra
black, tibie infuscated only at tips, the front ones somewhat thicker at apex,
not longitudinally impressed near apex, but quite as strongly notched there,
and the basal joint of front tarsi longer and more strongly curved.
COLEOPTERA.—LEA. 205
HETEROMASTIX PUNCTICORNIS sp. nov.
6 Deep black; head, prothorax, two basal joints of antenne, and front
coxe bright reddish flavous; front knees feebly diluted with red. With rather
dense and short pubescence.
Head with a faint impression near each eye. Antennz moderately long
and rather thin, third to tenth joints each with a puncture on a small polished
space near apex on the upper surface. Prothorax about twice as wide as long,
margins fairly wide at base and sides, but feeble across apex, sides slightly wider
near apex than at base, but not thickened; with distinct, submarginal punctures.
Elytra almost parallel-sided to near apex; with dense and sharply defined
punctures of moderate size. Length (¢Q), 4-4-5 mm.
Q Differs in having the head smaller, with less prominent eyes, antenne
shorter, without polished punctate spaces, legs shorter, and in the abdomen.
Hab.— Queensland: Bribie Island in August (H. Hacker).—Type, 0/2299
in Queensland Museum ; cotype, I. 12255 in South Australian Museum.
In my table it would be associated with H. geniculatus, from which it
differs in being smaller, middle and hind knees and scutellum black, antennze
much shorter, &¢.; from H. compar it differs in being more robust, with darker
legs and scutellum, antenne slightly stouter, and many of the joints with a
polished punctate space. Structurally it is closest to H. mgripes, and in appear-
ance it is much like the female of H. bryanti. The shining spots on the antennz
from some directions look like granules.
HETEROMASTIX TRICOLOR sp. nov.
6 Head, prothorax, scutellum, two basal joints of antennx, parts of palpi
and front legs (tarsi and tips of tibiw infuscated) flavous; rest of legs, mesoster-
num, metasternum, and abdomen black; elytra deep purple. With rather dense
pubescence.
Head gently convex, with two feeble impressions in front. Antenne
moderately long and fairly stout, third joint slightly shorter than fourth, eleventh
thinner and conspicuously longer than tenth. Prothorax about twice as wide as
long, sides and base with fairly wide margins, front margin very short. EHlytra
parallel-sided to near-apex; with crowded and sharply defined but rather small
punctures, becoming smaller at base and apex. Length ( ¢ 9), 4-5-6 mm.
Q Differs in having the head smaller, with less prominent eyes, antenne
shorter and thinner, and abdomen not notched.
Hab.—Queensland: Mackay (Blackburn’s and French’s collections from
R. E. Turner) ; Mapleton in October, and Brisbane in April (H. Hacker). New
South Wales: Galston (D. Dumbrell) ; Sydney (A. M. Lea).—Type, J. 12258
in South Australian Museum; cotypes, C/2300 in Queensland Museum, and in
National Museum.
A rather wide species; on several specimens the middle and hind knees,
206 MEMOIRS OF THE QUEENSLAND MUSEUM.
the middle tibiwe, and part of the third joint of antenne are pale; on one female
the scutellum is rather dark. In my table would be placed with H. anticus
and #H. geniculatus, from each of which it is distinguished by its greater width
and purple elytra; in the latter species the antennx are also decidedly longer and
thinner; the former species is also considerably smaller, with elytra wider pos-
teriorly, and antenne of male entirely pale.
HETEROMASTIX TARSALIS sp. nov.
¢ Black; prothorax, scutellum, parts of under surface of head,
mesosternum, and legs flavous; tarsi and sometimes tips of tibia infuseated.
With moderately dense pubescence.
Head with slight interocular impressions. Eyes rather large and
prominent. Antenne long and rather thin. Prothorax not twice as wide as
long, margined throughout; with submarginal punctures. Hlytra moderately
wide, almost parallel-sided to near apex, with dense and sharply defined but
rather small punctures. Front tibie moderately dilated to, and notched on
one side of apex; basal joint of front tarsi strongly curved on one side. Length,
4-5-25 mm. :
Hab Queensland: Mount Tambourine in December and January
(H. Hacker and A. M. Lea) ; National Park (Hacker).—Type, I. 11875 in South
Australian Museum; cotype, C/2302 in Queensland Museum.
The front legs approach those of H. tibialis, but the noteh at the apex
of the tibizw is less pronounced, and the incurvature of the basal joint of tarsi
is less; the elytra are also entirely black. In my table would be placed at F,
from all the species of which it is distinguished by its front legs and pale
seutellum; in addition H. victoriensis is a narrower species, with much darker
legs; H. pauxillus has longer and thinner antenne and black legs; and H. simplex
has shorter antenne and darker legs. There are seven males before me, but
I have been unable to identify the female amongst the many unidentified ones
under examination.
Var.—A male from the Blue Mountains (in Dr. E. W. Ferguson’s collec-
tion) structurally agrees with the type, but has the femora and tibiw (except
the knees) infuscated.
HETEROMASTIX PUSILLIOR sp. nov.
6 Black; prothorax, two basal joints of antenne, parts of under surface
of head, and knees flavous. With short pubescence.
Head with two feeble interocular impressions. Antenne long and rather
thin, third to eleventh joints subequal. Prothorax about twice as wide as long,
margined throughout, lateral margins slightly increasing in width to near apex,
where they are slightly thickened; with submarginal punctures. Elytra long,
thin, and parallel-sided to near apex; with dense and rather small but sharply
defined punctures, becoming smaller at base and apex. Length, 2 mm.
COLEOPTERA.—LEA. 207
Hab.—Queensland: National Park in December (H. Hacker).—Type
(unique), C/2301 in Queensland Museum.
The sides of the prothorax are dilated and thickened anteriorly, but not
abruptly as in H. pusillus and allied species, so that in my table it would be
associated with H. victoriensis, H. pauxillus, and H. simplex, from each of which
it is distinguished by its minute size. The tenth joint of the antenue is slightly
thicker than the ninth, but the difference is very slight, and not noticeable from
several directions.
LAIUS FLAVONOTATUS Lea.
Specimens of this curious little species were taken by Mr. Hacker on
mangroves, at Sandgate, in September.
BALANOPHORUS SCAPULATUS Fairm.
Several specimens, sexes, of this species from the Queensland National
Park, and some females from the Richmond River and Dorrigo (New South
Wales), have most of the head of a deep shining black, the tarsi and apical
~ parts of middle and hind tibiz more or less infuscated.
CARPHURUS LONGUS Lea.
C. atricapillis Lea, var.
Numerous specimens, all females, from the Cairns district, convince me
that C. atricapillis should be regarded as one of many varieties of C. longus, and
as the latter name was the first used it must be recognised as the typical one,
although by no means the commonest. Starting with it the various forms before
me, represented by four or more specimens, may be thus noted (the clothing and
apical half of antenne, which are dark on all the forms, not here taken into
consideration ) :—
Form 1, 2 .—Typieal longus. Entirely pale.
Form 2, 2 .—lLike 1, except that half or more of the hind femora are
deeply infuseated, or black.
Form 3, 9.—Var. atricapillis. Like 1, except that part of the elytra is
black or infuscated; the dark part sometimes continued along the suture for a
short distance towards the base, and sometimes encroached upon by the suture;
it occupies from one-fourth to three-fourths of the elytra, usually about one-
third; occasionally the tip of the abdomen is dark.
Form 4,9.—Like 3, but with a conspicuous black or infuscate, slightly
curved fascia, connecting the eyes; tip of abdomen always black.
Form 5,9 .—Head with a fascia as in 3, elytra black, with a wide but
somewhat irregular median flavous fascia, metasternum, tip of abdomen and
parts of hind legs black, middle legs sometimes partly dark, and some of the
other segments of abdomen infuscated in parts.
208 MEMOIRS OF THE QUEENSLAND MUSEUM.
There is a single specimen with the elytra almost entirely dark, the hind
femora and cox and part of the metasternum dark, and the interocular fascia
distinct; another has the head entirely pale, the elytra entirely dark, and parts
of the middle and hind legs and of the metasternum dark.
So many females of this species have been before me that I think it almost
certain that I have seen males, but not associated them with the species. If the
male is really before me it may be C. elegans, of which only the male is known,
and which has remarkable front tarsi.
CARPHURUS PISONIZ sp. nov.
d Flavous; two spots near base of head (sometimes conjoined) and
-metasternum black, five -to seven apical jomts of antennz infuscated. With
sparse white pubescence and straggling black hairs.
Head rather wide and irregularly impressed between eyes, with two
oblique median elevations; with irregular punctures, becoming crowded near
eyes. Antenne moderately long and scarcely serrated. Prothorax slightly
longer than greatest width (near apex), a shallow open depression near base;
with a few scattered punctures. Elytra almost twice the length of prothorax,
and much wider at base, each side near apex with a deep semicircular notch,
the anterior end marked by a subtriangular portion of the elytron, the posterior
by a long acute spine directed forwards and outwards; with dense and sharply
defined punctures, suddenly terminated near apex. Basal joint of front tarsi
with a small black comb. Length, 3-4 mm.
Hab.—Northern Queensland (Blackburn’s collection) ; Cairns district, in
abundance on sticky seeds of Pisonia brunomana (F. P. Dodd).—Type, I. 11938
in South Australian Museum; cotype, C/2341 in Queensland Museum.
Allied to C. cristatifrons, and with similar elytral armature, but the head
bimaculate and the elytra not; the head from the side appears evenly convex,
whereas on cristatifrons the strong ridges, abruptly terminated in front, give a
very different appearance ; its head also has much larger and deeper impressions
than the present one. The impressions between the eyes are not very deep;
there are two oblique elevations between them, and these, with a feeble Jongi-
tudinal interocular elevation, appear to form a V (or feeble Y); each fork of
the YV touches one of the black spots. The spines on the elytra are broken on
many of the specimens, and when about half of each is left the notch from some
directions appears as an almost circular hole; the part beyond the spines is
sometimes of the same colour as the rest of the elytra, but is usually of a lemon-
yellow colour, and is impunctate. The abdomen is of a somewhat redder tone
than the other pale parts. Under the microscope the tarsal comb is seen to
consist of ten or eleven teeth. More than one hundred specimens were removed
from the seeds, but all are males.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Puate XIII.
CARPHURUS PISONIZ Lea. Upper figure, anterior leg; lower figure, elytron.
Photos.—H. Hacker. Face page 208.
COLEOPTERA—LEA. 209
CARPHURUS PURPUREIPENNIS sp. nov.
¢ Flavous or reddish flavous; elytra deep purple; mesosternum, meta-
sternum, cox, most of femora, and six or seven apical joints of antenne, black
or blackish. With rather long, blackish hairs, rather dense in places; elytra
with short, whitish pubescence.
Head rather large, irregularly impressed between eyes, a rather wide and
deep impression behind them, a hairy longitudinal ridge crossing its middle, an
acute oblique ridge on each side of middle between eyes, a curved and slightly
elevated line in front; base punctate and transversely strigose. Antenne long,
moderately stout, most of the joints serrated. Prothorar shghtly longer than
greatest width, a wide shallow subbasal depression; with a few scattered punc-
tures. Hlytra more than twice the length of prothorax; each side at basal third
with a triangular pale-tipped projection; punctures dense and sharply defined,
becoming sparser and smaller on tips. Basal joint of front tarsi with a black
inner comb. Length (dQ ), 6-8 mm.
Q Differs in having the head smaller, with feeble depressions only, with-
out ridges, and less hairy, apex of prothorax less produced, sides of elytra
unarmed, legs and antennw shorter, and front tarsi simple.
Hab.—Northern Queensland (Blackburn’s collection) ; Cairns (E. Allen
and F. P. Dodd); Yarrabah and Mount Bellenden-Ker (Dr. E. Mjoberg).—
Type, J.12233 in South Australian Museum; cotype, C/2303 in Queensland
Museum.
Alhed to C. armipennis, but readily distinguished by the two acute ridges
on head; these, when viewed from behind, appear as two conical tubercles. On
several specimens the suture near the base is obscurely reddish; the base of the
prothorax is sometimes of a different shade of colour from its other parts. On
the middle of each elytron of the male, in a line with the lateral teeth, there is a
space like a small scar; it appears at a glance to be accidental, but is alike on
the six males under examination.
CARPHURUS MACULICOLLIS sp. nov.
S$ Black and flavous, or reddish flavous. With short whitish pubescence
and long dark hairs.
Head rather large and unevenly convex, a rather long, shallow, oblique
depression each side in front; with unevenly distributed punctures; base trans-
versely strigose. Antenne moderately long, fourth to tenth joints serrated.
Prothorax distinctly longer than wide, widest at about apical fourth, apex not
much wider than base, a wide shallow depression near base; punctures fairly
numerous on sides, sparse elsewhere. Hlytra more than twice the length of
prothorax, sides rather suddenly dilated near base, apex about one-third wider
than base; with dense and small, but rather sharply defined punctures, becoming
minute on tips. Basal joint of front tarsi with an inner comb from base to apex.
Length (¢Q ), 5-7 mm.
te)
210 MEMOIRS OF THE QUEENSLAND MUSEUM.
Q Differs in having the head smaller, with shallower depressions and
sparser hairs, eyes much smaller and less prominent, antenne shorter, legs
shorter and front tarsi combless.
Hab.—Queensland: Blackall Range in October (F. E. Wilson) ; Brisbane
(HI. J. Carter) ; National Park (H. Hacker) ; Mount Tambourine (A. M. Lea).—
Type, 7.12254 in South Australian Museum; cotype, (/2304 in Queensland
Museum.
The eyes of the male are larger and more prominent than usual, their
combined width being almost equal to the space between them; on the female
their combined width is scarcely half that of the intervening space; on the head
of the male the dark hairs form two or three feeble fascicles. The species is close
to C. pallidipennis, but the male has the head wider, with larger eyes, impressions
different, and small black fascicles, elytra with smaller punctures, and a dark
spot on each side near the scutellum; in my table the specimens with immaculate
elytra would be associated with CU. pallidipennis, but the spotted ones not with
C. marginiventris, as the spots are too short and the elytral punctures finer; the
heads of the males are also differently sculptured. On typical males the pale
parts are the head (except for a transverse black mark near the base—on some
specimens owing to the projection of the prothorax it appears to be at the base’
itself), prothorax (except for a large black spot on each side not quite reaching
the base), elytra (except for a rounded black spot on each side of the scutellum),
scutellum, prosternum, mesosternumn, tips and sides of abdominal segments,
three to five basal joints of antenne, trochanters, knees, and most of tibiz and
tarsi. The typical females are coloured much as the males except that the black
part of the head is larger, and that the outer apical angles of the elytra are
infuscated, the basal spots are sometimes more extended. On most specimens,
of both sexes, there is a dark streak, almost the entire length of the upper edge
of the hind tibiz.
Var. A.—Seven females, from Mount Tambourine, differ from typical
females in having the head entirely pale (on one of them there is, however, a
very small medio-basal spot), elytra entirely pale, and more of the antenne
dark; they differ from females of pallidipennis in being larger, and with smaller
elytral punctures; it would, however, be unsafe to identify specimens of either
species from females only.
Var. B.—A female, from Mount Tambourine, has the head entirely pale,
as also the elytra; except for a slight infuscation on each side near the apex,
most of its abdomen is pale.
Var. C.—A male, in the Queensland Museum, has the prothorax and
elytra entirely pale; it seems fairly close to the description of C. xanthochrous
and (. tachyporordes, but its scutellum (as on all the other specimens before me)
is pale instead of black.
COLEOPTERA.—LEA. 211
NECCARPHURUS ANGUSTIBASIS sp. nov.
© Black and Iighly polished; muzzle, basal half of antenne, and extreme
base of prothorax favous.
Head deeply impressed between eyes; inter-antennary space elevated and
subtubereulate. Antenne rather long and thin, none of the joints transverse.
Prothorax longer than wide, apex more than twice the width of base, sides
strongly rounded and narrowed from apex to near base, and then subparallel to
base, which is feebly bilobed, a deep, transverse, open, subbasal depression.
Elytra slightly wider than widest part of prothorax, in parts slightly undu-
lating; almost impunctate. Basal joint of front tarsi lopsided, with an inner
comb. Length ( ¢Q ), 2-5-3 mm.
Q Differs in having the head smaller, without transverse impression,
inter-antennary space very feebly elevated, antennze thinner, and front tarsi
simple.
Hab.— Queensland: Cairns district (F. P. Dodd and A. M. Lea) ; Innis-
fail—tType, J. 9182 in South Australian Museum; cotype, C/2305 in Queensland
Museum.
On several specimens the sides of the elytra near the base are obscurely
diluted with red, on one female parts of the legs are obscurely reddish; from
four to six apical joints of the antenne are more or less deeply infuscated. One
of the specimens was attached to a sticky seed of Pisonia brunomana. In general
appearance the species is close to \V. sobrinus, but the head of the male is
differently excavated, and the inter-antennary elevations differ as follows :—
sobrinus. angustibasis.
From directly in front.
They appear to terminate on an The median one is posterior to the
even line posteriorly, each being others and on a lower level.
separately rounded there.
From behind.
The head appears to have two The head appears to have a median
small tubercles.. flat-topped convexity.
From each side.
The head appears to have a The head does not appear to have
tubercle before each eye, one behind it a tubercle behind the eye and at its
and one at its middle, the front one middle.
being more conspicuous than the
others, which disappear when viewed
from a shghtly lower elevation.*
4 Lea, Trans. Ent. Soc. Lond., 1909, fig. 6.
212 MEMOIRS OF THE QUEENSLAND MUSEUM.
HELCOGASTER PUNCTIPENNIS Lea.
A male of this species, from Cairns, has only five joints of each antenna
dark; by a printer’s error® the female was described originally as having the
head ‘‘absolutely’’ bifoveate in front. instead of ‘‘obsoletely’’; two other females
have the fover rather distinct, and one of these has the inner apices of the elytra
obscurely testaceous.
HELCOGASTER VARIUS Lea, var. FLAVOPICTUS var. nov.
Four specimens, two of each sex, from Bribie Island, appear to represent
another variety of this species; they are rather larger than usual, up to 5-5 mm.,
and have the pale portion of the elytra larger; their dark parts are a narrow
triangle about the scutellum, and the tips for about one-fourth their length at
the suture, and sides (but more between) ; the pale parts of the legs are brighter
and more extended than in other described varieties. On the males the black
patch at the base of the head is terminated before the sides; on the females there
is a narrow irregular reddish line near each eye, and on one of them the two are
transversely connected.
HELCOGASTER INSIGNICORNIS sp. nov.
¢ Black and flavous. Upper surface with sparse, dark, erect hairs.
Head wide, with a large excavation behind muzzle. Antenne with first
joint thick, with a fovea near apex; second short, third to fifth rather wide, the
following ones rather thin. Prothorax about as long as the greatest width, sides
dilated from base to apex, near base with a large depression closed behind and
at the sides, but shallowly connected, towards each side, with a shallow, latero-
apical depression. Elytra rather long, with a few inconspicuous punctures.
Basal joint of front tarsi with a small black comb. Length (¢@Q ), 2-25-3 mm.
Q Differs in having the head narrower, with eyes slightly smaller,
excavation replaced by a rather shallow depression, that is notched posteriorly
in middle; antennx shorter, basal joint much smaller and non-foveate, third and
fifth joints smaller; prothorax with subbasal depression not, or scarcely, con-
nected with latero-apical ones (and these often scarcely defined) ; and legs shorter,
with front tarsi combless.
Hab.—Queensland: Mount Tambourine (A. M. Lea).—Type, J. 11911 in
South Australian Museum ; cotype, C/2306 in Queensland Museum.
Allied to H. foveicornis, but basal joint of antennze of male smaller and of
different shape, elytra with a pale basal zone and, at most, only three apical
segments of abdomen entirely black. The female is much like females of H.
tuberculifrons, H. simpliciceps, H. maculiceps, and others having bicolorous
elytra, but the male is at once distinctive. On the male the black parts are the
apical three-fourths of elytra and the tip (one or two segments) of abdomen; the
®Lea, Trans. Ent. Soc. Lond., 1909, p. 225.
COLEOPTERA.—LEA. 213
antenne have from three ‘to five basal joints and the tip of the eleventh pale, the
others being deeply infuscated; the head and prothorax are of a slightly redder
tone than the other pale parts. The excavation on its head occupies about one-
third of the width across eyes, its bottom is somewhat irregular and hind margin
semicircular ; only the five apical joints of its antenne are seen to be longer than
wide from most directions, but from several the two preceding joints seem also to
be longer than wide. On two males the metasternum is dark; on all the females
it is dark, and sometimes the mesosternum as well. On many of the females the
head, except at base, and upper portion of the first joint of antenne, are more or
less deeply infuseated, on some only the muzzle is infusecated, on nine specimens
the head is entirely pale; on several the apical joint of antenne is entirely dark;
the dark segments of the abdomen vary in number from one to three. There are
some fine punctures and strigosities behind the eyes, but they could be easily over-
looked ; there are no sharply defined punctures on the elytra.
One male (also from Mount Tambourine) differs from the others in having
the excavation on the head larger, and at bottom with a distinct longitudinal ridge
(on the others the bottom of the excavation is obscured by a mealy substance),
and the antenne entirely pale, although the basal joints are paler than the apical
ones.
HELCOGASTER HACKERI sp. nov.
6 Black; head, prothorax, and most of antenne and of legs flavous.
Sparsely clothed with white pubescence, and with a few dark hairs.
Head wide, with a large, deep, interocular excavation. Antenne rather
long, moderately serrated, apical joint almost as long as two preceding combined.
Prothorax distinctly transverse, apex wider than base, near base with a rather
wide and deep, closed depression. Elytra moderately long; with rather dense
and minute rugulose punctures. Basal joint of front farsi with a black imner
comb. Length (¢ 9), 2-25-3-5 mm.
2 Differs in being larger, head smaller, without tubercles or excavation,
with a shallow depression each side in front, and a shallow median line, pune-
tures sparser and more sharply defined, and black, except that the muzzle is
obscurely reddish; the antenne are thinner and much darker, prothorax with
only the base and sides pale, and narrower across apex, abdomen larger and
wider, legs with hardly more than the knees pale, and front tarsi simple.
Hab.— Queensland: Bribie Island in August (H. Hacker and A. M. Lea) ;
Brisbane in October (Hacker).—Type, J. 17905 in South Australian Museum ;
eotype, C/2307 in Queensland Museum.
_ One of the most interesting beetles occurring on the island. It is close to
H, foveiceps, to which the specimens at first glance appear to belong, but on that
species the sub-basal fovew of the head leave a medio-basal space, which projects
subtriangularly forwards, the projection itself longitudinally grooved; on the
present species the place of the projection is taken by a rather large depression,
214 MEMOIRS OF THE QUEENSLAND MUSEUM.
which appears on some specimens to extend to the base itself, although it does
not do so; from the inter-antennary space a wide and uneven elevation projects
subtriangularly backwards into the excavation, and from directly behind the sides
of this elevation appear like two minute tubercles (on foveiceps a smaller median
one may also be seen). I have repeatedly compared the heads of the males from
many points of view, and cannot satisfy myself that the two forms belong to
but one species, despite the close similarity of all parts but the head, and the
curiously coloured antenne; the females of the two species are practically indis-
tinguishable. The middle of the base of the head cf the male is usually black,
but, as that part is normally concealed, the head appears to be entirely pale; the
antenn are usually entirely pale, but with the middle joits slightly darker than
the others, but sometimes much darker; the tarsi and middle and hind tibie
are usually distinctly infuscated; on the female the tip of the antenne is some-
times black, but is usually obscurely paler than the preceding joints. The head
of the male is densely punctate, but the punctures are not sharply defined, its
prothorax also has fairly numerous punctures towards the sides; on the female
those of the prothorax are more distinct but scarcely larger.
HELCOGASTER TRIFOVEICEPS sp. nov.
¢ Flavous; scutellum, abdomen (except tips of segments on upper
surface), mesosternum, metasternum, apical half of antenne, and parts of legs,
black or infuscated. With sparse, white pubescence ; denser on head and abdomen
than elsewhere.
Head wide, with a rather deep, transverse, interocular depression, its
posterior margin trisinuate; front with an obtuse elevation having two small
tubercles posteriorly. Antenne moderately long. Prothorar distinetly trans-
verse, base much narrower than apex, near base with a large, deep, closed,
transverse depression ; punctures rather dense on sides, sparse elsewhere. Elytra
rather short, dilated posteriorly ; with sparse, small, rugulose punctures. Basal
joint of front tarsi with a small black comb at inner apex. Length, 2-5-3 mm.
Hab.—Queensland: Dalby (Mrs. F. H. Hobler).—Type, 7. 12124 in South
Australian Museum ; cotype, C/2508 in Queensland Museum.
The sculpture of the head, which, with the prothorax and elytra, are
entirely pale, readily distinguishes this species from all previously named ones.
At least four of the basal joints of antennz are flavous, the fifth and sixth are
also sometimes scarcely darker ; the cox and bases of femora are black, the tibiz
are usually slightly infuscated in the middle. The head is opaque, owing to
dense punctures; these are individually so small as to be scarcely traceable, and
in addition are partly concealed by clothing ; from some directions the interoeular
space appears to have three small fovex connected by a curved line; from directly
in front the median one is seen to be much larger than the others; from behind
the two small tubercles on the interocular elevation are quite distinct.
COLEOPTERA.—LEA. : ; 215
HELCOGASTER TRISINUATUS sp. nov.
¢ Flavous; apical three-fifths of elytra, tip of abdomen, and six or seven
apical joints of antenne black, metasternum infuscated. A few dark hairs
seattered about.
Head with a deep interocular excavation, its posterior end trisinuate, a
large obtuse tubercle in front; with sparse and small punctures, becoming denser
about base. Antenne rather long. Prothoraxr about as long as the greatest width,
base decidedly narrower than apex, a large, rather deep, transverse, closed, sub-
basal depression. Elytra moderately long, almost parallel-sided; with fairly
numerous and small but (for the genus) rather sharply defined punctures. Tip
of abdomen with two small processes. Basal joint of front tarsi with a small
black comb. Length (¢@), 2-5-3-75 mm,
@ Differs in having the head smaller, a shallow fovea representing the
middle of the excavation, and a feeble depression on each side, its sides; frontal
tubercle much smaller and more obtuse, antennz shorter and thinner, elytra less
parallel-sided, abdomen with the tip simple, and front tarsi combless.
Hab.— Queensland: Cairns distriet (A. M. Lea).—Type, J. 11949 in South
Australian Museum; cotype, C/3209 in Queensland Museum.
In my table® would be placed with F, from all the species of which it is
distinguished by its pale head. At first glance it appears to belong to H. puncti-
penns, but the sculpture of the head of the male is different, and the elytral
punctures are smaller; from the male of H, seminigripennis it is distinguished
by the large cephalic excavation, with its base conspicuously trisinuate, the
median sinus wider than the others. Six males and twelve females were obtained.
DASYTES SUBELLIPTICUS sp. nov.
Black, parts of antenne and of legs flavous. With rather dense, depressed,
ashen pubescence.
Head with small and rather dense punctures; with two feeble depressions
in front. Antenne slightly passing base of prothorax, most of the joints trans-
verse. Prothorax widely transverse, base and sides finely margined; with very
small punctures. Elytra at base scarcely wider than prothorax, sides slightly
dilated to beyond the middle; with dense and small, but rather sharply defined
punctures. Length, 1-75-2 mm.
Hab.— Queensland: Bribie Island (H. Hacker and A. M. Lea).—Type,
I, 12285 in South Australian Museum ; cotype, C/2310 in Queensland Museum.
A minute elliptical species, much like D. ellipticus on a greatly reduced
seale; in my table,’ on account of the bicoloured legs, it would be associated with
D. bourgeoisi (now D. julest), but it is much smaller and wider in proportien; in
_®Lea, Trans. Ent. Soe. Lond., 1909, p. 215.
7 Lea, Trans. Ent. Soc. Lond.. 1909, p. 240.
216 MEMOIRS OF THE QUEENSLAND MUSEUM.
size it is about equal to D. corticarioides. The femora are usually deeply infus-
cated, except at base and apex; on some specimens the front legs are almost
entirely pale; the apical half, or less, of the antenne is infuscated; on some
specimens vague remnants of a subbasal depression may be seen on the pronotum,
but from most of them even these are absent.
Famity TENEBRIONID.
PALORUS EUTERMIPHILUS sp. nov.
Bright castaneous. Upper surface glabrous, under surface almost so.
Head moderately wide, with rather dense punctures. Clypeus with smaller
punctures than on rest of head, its hind suture semicircular. Eyes small, without
canthi, extreme sides only visible from above but distinct from below. Antennz
scarcely longer than their distance apart, parallel-sided except near base, third.
to tenth joints distinctly transverse, the eleventh almost as long as wide.
Prothorax moderately transverse, sides rounded, distinctly dilated to near apex,
and finely margined, hind angles rectangular; punctures slightly larger but
otherwise as on head. Scutellum widely transverse. Elytra opaque, parallel-
sided to near apex, base wider than base of prothorax, but less than its greatest
width; with deep strie containing rather shallow punctures, interstices acutely
costate almost throughout. Under surface with dense punctures on prosternum,
mesosternum, and sides of metasternum, much sparser and smaller elsewhere.
Legs short. Length, 2-75-3 mm..
Hab.— Queensland: Townsville, twelve specimens from termites’ nest,
Eutermes sp. (G. F. Hill, No. 1033).—Type, J. 11588 in South Australian
Museum ; cotype, C/2311 in Queensland Museum. ;
This species should perhaps have been regarded as the type of a new
genus, but I am averse from proposing new genera for inquilines except on very
strong grounds. The entire absence of a club to the antenne and the eyes not
encroached upon by canthi seem to exclude it from Tribolium, to the species of
which it bears a strong general resemblance. The antenne and eyes, except that
the latter are smaller, with their edges just visible from above, are much as in
several species of Palorus. The dilated front of prothorax, and opaque elytra,
with acute coste, are very distinctive amongst the allied genera. The colour is
an almost uniform and rather pale castaneous, the antenne are slightly darker
than the head, but the terminal joint is slightly paler.
Famity MELANDRYID.
PAROMARTEON MUTABILE Blackb.
I have previously* commented upon this species, but as there are now
before me many other specimens, including several sharply defined and more
or less constant varieties, it appears desirable to name some of them. The sexes
8 Lea, Trans. Roy. Soc. S. Aust., 1917, p. 168.
COLEOPTERA.—LEA. 217
may be readily distinguished by the abdomen; in the male the middle of the
apical segment is gently incurved, in the female that segment is larger, and the
tip is evenly rounded. The under surface, except the prosternum, appears to be
always deep. black.
Var. nigripenne var. nov.—F rom Victoria (Dividing Range), New South
Wales (Sydney and Dorrigo), and Queensland (Brisbane), there are nine
specimens with the elytra entirely black or blackish, and the head usually with
the basal half black.
Var. apicale var. nov.—Twenty-one specimens, from Brisbane and Bribie
Island, have the apical two-fifths of elytra and the basal half of head deep black ;
the scutellum varies from flavous to deeply infuscated ; on some of the males the
hind femora are infusecated in the middle.
Var. parvum var. nov.—F our specimens, from Bribie Island, are close to
the preceding variety, but are smaller (3 mm. only), and have the apical half of
elytra black, but the head and scutellum entirely pale.
Var. fasciatum var. nov.—F ive specimens, from Bribie Island, are very
small (3-3-5 mm.), and their elytra have two black fascix : a complete narrow one
at the apical two-fifths, and an interrupted one at the apical fifth; the hind and
middle femora are partly black; in the males the head is almost entirely dark, in
the females it is entirely pale.
Famiry MORDELLID®.
MORDELLA BRIBIENSIS sp. nov.
Black ; base of antennz and parts of front legs obscurely diluted with red.
Clothed with black and greyish-white pubescence.
Comparatively short. Scutellum semicireular. Pygidium rather short, its
apical portion almost parallel-sided and then truneated. Spurs to hind tibiz
unequal. Length, 3-4 mm.
Hab.—Queensland: Bribie Island (H. Hacker and A. M. Lea).—Type,
I. 12132 in South Australian Museum; cotype, C/2312 in Queensland Museum.
The pale pubescence is uniform on the head, and so placed on the prothorax
as to distinctly define three black spots: a large median one and a smaller one on
each side; on the elytra it forms a fairly conspicuous narrow basal edging, but
elsewhere the pale hairs are scattered thinly and do not form spots; from some
specimens, except at the base, they are absent; from the metasternum the pale
pubescence is almost absent, and it is absent from a large spot on each side of
four basal segments of abdomen. About five joints of the antenne are transverse.
The external sexual differences are feebly defined ; the male has the apical portion
of the pygidium more parallel-sided, and the front tarsi slightly wider, although
thin. In some respects the species is close to some forms of M. baldiensis, and it
218 MEMOIRS OF THE QUEENSLAND MUSEUM.
belongs to the same group,’ but the pale pubescence of the prothorax bounds
three sharply defined dark spots, and the pygidium is differently shaped; from
the form of M. nigrans, with somewhat similar prothoracic markings, it is distinet
by the pygidium, its narrow portion being only about half as long as on that
species.
Famity CANTHARID.
HORIA CEPHALOTES Oliv., Ent., iii, p. 5, pl. 1, fig. 3.
A specimen labelled *‘ Johnstone Riv.,’’ in Miskin’s writing, is a male of
this species; remarkable for its wide flat head, and conspicuous jaws. The
Queensland locality, however, needs confirmation, as the species is a well-known
Javanese insect.
Faminy (ZK DEMERID_.
PSEUDOLYCUS HHMORRHOIDALIS Fab., var. MARGINATUS Guer.
Form 5.—Three specimens from the Queensland National Park differ from
the preceding four forms of this variety in having the sides of the prothorax
entirely pale, the dark discal portion is wider at the base than the apex, and
the elytra are blackish for a short distance from the base—near the suture on
two of them there is a pale spot on each side of the base of the head.
MORPHOLYCUS COSTIPENNIS Lea.
A female of this species, from the Queensland National Park, has the red
of the prothorax reduced to € small spot on each side of the base. :
COPIDITA MARITIMA Lea. .
A specimen, from Bribie Island, and another, from Stradbroke Island,
have similar elytra to those of a specimen commented upon as in Mr. Carter’s
collection, except that there is a slight infuscation on each side of the scutellum ;
on the Bribie Island one the dark prothoracic markings are conjoined, on the
other they are not conjoined across the middle. Another, from Bribie Island
(a small male), has the cephalic spot larger than usual, and the derm of both
seutellum and elytra entirely black.
COPIDITA TENUICOLLIS sp. nov.
¢ Flavous; three irregular lines on prothorax, elytra (except suture and
extreme sides), knees, parts of tarsi and two apical joints of antennx, black or
infuscated. Densely clothed with short, ashen pubescence.
Head rather long, gently convex between eyes, with rather dense and
sharply defined punctures, becoming crowded at base, and smaller in front;
jaws notched at apex. Eyes large and coarsely faceted. Antenne long and thin,
most of the joints cylindrical, eleventh semi-double. Prothorax much longer than
wide, sides slightly dilated near apex, base narrowly margined; punctures
® Lea, Trans. Roy. Soc. 8S. Aust., 1917,.p. 217.
ie
COLEOPTERA.—LEA. 219
crowded and larger than on head. Elytra much wider than prothorax, parallel-
sided to near apex; each with four discal coste; punctures small and crowded.
Legs long and thin; tibiw bispinose at apex; claws each with an obtuse basal
appendix. Length, 11 mm.
Hab.—Queensland: Stradbroke Island in December (H. Hacker ).—Type
(unique), C/2313 in Queensland Museum.
The elytra are much as in C. macleayi, and the prothorax also has three
dark markings, but the prothorax itself is decidedly longer and thinner, with
much stronger punctures, head immaculate between eyes, and these decidedly
larger, and the antenne entirely pale. On the prothorax the dark part on each
side is continuous from base to apex, and widest at the apical third, the median
line is thinner, shorter and irregular; the pale sutural portion is shghtly wider
than the scutellum at the base, and becomes very narrow posteriorly.
COPIDITA NIGRIPENNIS sp. nov.
6 Flavous, elytra blackish with a vague bluish gloss, parts of tarsi and
three spots on prothorax slightly infuseated. Rather densely clothed with ashen
pubescence, more conspicuous on elytra than elsewhere,
Head rather long, with dense sharply defined punctures, becoming smaller
in front; jaws notched at tips. Antenne long and thin, second to sixth joints
cylindrical (the following ones missing). Prothorax long and thin, sides some-
what dilated near apex, base margined and bisinuate; punctures crowded and
slightly larger than on head. Elytra much wider than prothorax, parallel-sided
to near apex, each with four discal costa, of which the third is scarcely traceable ;
with crowded and mostly rugose punctures, but many sharply defined. Legs
long but not very thin; tibiw unispinose at apex; claws thin, each with a small
basal appendix. Length, 10 mm.
Hab.—Queensland: Bribie Island (H. Hacker).—Type (unique), C/2314
in Queensland Museum.
In some respects close to C. mira, but prothorax longer and less dilated
in front, and palpi normal; from the preceding species it differs in having the
legs thicker, with only parts of the tarsi infuseated, the tibiz unispinose, and less
of elytra pale; the three faint infuscations on the prothorax are across the
apical third; the suture is very obscurely and narrowly diluted with red.
DOHRNIA SEMIFLAVA sp. nov.
2 Black and flavous. Head and prothorax rather sparsely clothed, else-
where with short, dense pubescence.
Head gently convex between eyes, shallowly depressed in front; with
dense and rather small but sharply defined punctures; jaws notched at apex.
Eyes rather long and finely faceted. Antenne long and thin, the joints
cylindrical. Prothorar about as long as the greatest width (near apex), a
220 MEMOIRS OF THE QUEENSLAND MUSEUM.
narrowly impressed line (dilated in middle) across base, and a shallow depression
across apex; punctures as sharply defined as on head, and slightly larger.
Elytra much wider than prothorax, parallel-sided to near apex, each with four
diseal coste; punctures crowded and usually sharply defined, but some
transversely confluent. Length, 6-7 mm.
Hab.—Queensland: Brisbane (H. Hacker); Glen Lamington (Dr. E.
Mjoberg).—Type, C/2315 in Queensland Museum; cotypes, J. 12241 in South
Australian Museum, and in Stockholm Museum.
Referred to Dohrnia on account of the eyes. The flavous parts of the
type are the head between the muzzle and eyes, base of six first joints of antenne,
and under surface of the first two, palpi (except the tips), prothorax (both
surfaces), scutellum, basal half of elytra, and legs (except tips of middle and of
“hind tibizw, and the middle and hind tarsi, which are infuscated). The specimen
in the South Australian Museum is like the type, except that the hind femora
are also infuscated; the other cotype has the legs (except the front cox) and
antenne entirely dark, and with slightly less of the elytra pale. On all three
specimens the dark parts usually have a metallic gloss; only two of the discal
coste are at all distinct on each elytron.
Faminry CURCULIONID-t.
EUOPS TUBERCULATUS sp. nov.
2 Black, in most parts with a sight purplish gloss.
Head with sparse and irregularly distributed punctures. Eyes large,
almost touching in middle. Rostrum rather short, dilated and finely serrated
at apex, and narrowed to base. Antenne short; club stout, about the length of
six preceding joints combined. Prothorax about as long as basal width, narrowed
to apex, a distinct bisinuate punctate line near base, a shallow transverse impres-
sion on each side at apical third, a rather large but obtuse tubercle in front of
each; sides with distinct punctures, upper surface almost impunetate. Elytra
not much longer than wide, much wider than prothorax ; with somewhat irregular
rows of fairly large punctures; third interstice at basal third with an obtuse
tubercle, between it and base shallowly depressed, sixth interstice with a small
obtuse tubercle at summit of apical slope; each shoulder with an acute conical
tubercle, projecting outwards. Abdomen irregularly punctate and _ strigose,
four basal segments each with a short, hairy, double stripe across middle;
pygidium with rather large punctures on most of its surface. Femora stout;
front tibize bisinuate on lower surface. Length, 3 mm.
Hab.—Queensland: National Park in December (H. Hacker) —Type
(unique), C/2316 in Queensland Museum.
Readily distinguished from all other Australian species by the tubereulate
upper surface, with armed shoulders.
COLEOPTERA.—LEA, 221
Famiry CHRYSOMELID_E.
CHALCOLAMPRA TENUIS Lea.
Mr. Hacker has taken several specimens of this species in the National.
Park, near Brisbane, and these have the elytral markings not in the form of
four abbreviated fasciw connected with the suture only, but the three apical ones
are connected at their outer edge as well, so that there are two pale elliptical
spots on each side of the suture (one postmedian and one subapical) ; on one
specimen the two basal fascix are free externally, and on another the three
apical fasciw are not quite connected externally, so that the spots are not entirely
enclosed.
DITROPIDUS DAVISI Saund.
This species is abundant in South Australia and is very variable in size
and markings; the length ranges 2-25-3-75 mm.; the width, apart from sex,
also varies, the smaller specimens being decidedly narrower than the larger ones.
The male is narrower and less robust than the female, its abdomen is smaller,
nonfoveate, incurved to the middle, and with the tips of the pygidium slightly
produced forwards; the seriate punctures on the elytra are really rather small,
but owing to ‘‘waterlogging’’ appear to be decidedly large; their true sizes may
“be seen from an oblique direction. All the forms have the intercoxal process
of the prosternum semicireularly emarginate posteriorly, with the hind angles
sharply produced; the legs and parts of the under surface are also variable, but,
disregarding these, some of the forms before me are as follows :—
Typical—On this form (a rather rare one) the prothorax is immaculate,
the head is dark at the base, and the elytra have the base, apex, and suture dark,
so that each elytron has a large pale spot, but each spot is sometimes. greatly
reduced in size and sharpness. Hab.—New South Wales and South Australia.
Var. A.—Like the typical form, except that the head is entirely pale; on
one specimen the elytral spots are greatly reduced in size and brightness. Hab—
South Australia (Ooldea and Port Lincoln).
Var. B.—Like the typical form, except that it is not quite so wide, and
tiiat the prothorax has a transverse black or blackish fascia extending almost to
its sides; the fascia varies from about one-third the length of the segment to
covering its entire surface except for narrow edgings, occasionally it actually
touches parts of the sides; the elytral spots on such specimens are usually greatly
reduced in size, and the black part of the head is greatly extended; on one
specimen of it the prothoracic fascia appears as three semi-detached spots; on
another the fascia appears as a shght and rather narrow infuscation. Hab—
Scuth Australia. ;
Var. C.—Like the preceding variety, except that the prothoracic fascia
is broken up into two spots. Hab.—South Australia (Adelaide and Lucindale).
Var. D.—Prothorax entirely dark, head dark except for parts of muzzle;
222 MEMOIRS OF THE QUEENSLAND MUSEUM.
a fairly large but ill-defined pale spot on each elytron; one female of the variety
is still attached to a male of B; another female has a part of the scutellar lobe
pale; one male from Gladstone (Queensland) of the variety agrees perfectly
in outlines with the typical male, but has distinctly coarser punctures on both
prothorax and elytra; its elytral spots are very obscurely defined, and the legs
and prosternum are almost entirely dark. Hab.—Queensland, New South Wales,
South Austraha.
Var. E.—Head and prothorax coloured as on the typical form, but elytra
with the red extended to cover most of the surface, excluding a rather narrow
but somewhat zigzag strip at the base; the suture is also very narrowly dark,
but near the apex the dark part is dilated to form an oval spot; the apex,
however, 1s entirely pale. Hab.—Queensland (Charters Towers).
Var. F.—Head and prothorax entirely pale reddish flavous, except that
the base of the latter is very narrowly black; elytra flavous, the base and suture
yery narrowly black. a narrow part of the apex black, but the black part slightly
advanced along the suture and sides, so as to be strongly bisinuate on its immer
edee. On this variety, except for the claws, the legs are entirely pale. Hab.—-
South Australia ; Moonta).
There are other varietal forms before me, but I have not considered it
advisable to attach letters to those of which I have seen but one specimen. It
is probable that several published names will have to be treated as synonyms of
the species.
DITROPIDUS IGNITUS sp. nov.
¢ Brilliant coppery red, in places coppery green, under surface and legs
black, with a bluish gloss; labrum and basal half of antenne reddish. Under
surfaee and legs with rather sparse pubescence.
Head with sharply defined punctures of medium size; median line distinct.
Eyes separated about the length of two basal joints of antenne. Prothorax
at base more than twice as wide as the median length, sides strongly rounded ;
with dense, sharply defined, and not very small punctures. Elytra not much
longer than the basal width, which is almost twice that of the apical; with rows
of not very large punctures, on the sides set in deep striz; interstices with small
and fairly numerous punctures. Legs moderately stout, front ones slightly
longer than hind ones. Length (¢@ ), 2-5-3 mm.
Q Differs in being shightly more robust, eyes about one-third more distant
from each other, elytra less narrowed posteriorly, legs somewhat thinner, the
front ones no longer than the hind ones, and in the abdomen.
Hab.—Queensland: Cairns (E. Allen); Bowen (Aug. Simson).—Type,
T. 10925 in South Australian Museum; cotype, C/2317 in Queensland Museum.
A beautiful, briefly oblong-elliptic species, in general appearance close to
D. venustus and fairly close to D. costatus, but distinguished from both by the
non-strigose sides of prothorax; the punctures there are not even confluent, and
COLEOPTERA.—LEA. 223
are mostly slightly smaller than those in the middle. The eyes of the male are
more widely separated than in the male of D. dorie. From some directions the
head appears coppery red, from others coppery green; from some the prothorax
appears coppery green throughout, but usually only the sides appear to be of
that colour; the scutellum and tips of elytra, the latter varying with the point
of view, are also coppery green; the pygidium similarly varies from coppery
green to coppery red. In some lights mest of the upper surface of some
specimens appears purple. The seriate punctures on the elytra are about as
long as those on the prothorax, but narrower.
Vars.—One male has the head, prothorax, and pygidium purple, the
prothorax in some lights with a bluish gloss, its seutellum and elytra are bright
coppery green, except the tips of the latter, which are purple, its tibie are
obscurely diluted with red in parts; a female mounted with it has the head and
prothorax coppery green, the muzzle of the former and the sides of the latter
purplish in some lights, its elytra are deep blue, with the sides and punctures
purplish, the purple from some directions appearing to cover almost the whole
surface, its pygidium is blue and purple. The only specimén from Bowen, in
the Museum, is a male, and has the whole of the upper surface and pygidium
deep blue, altering to purple; from some directions the prothoracic specimens
have a coppery glitter (as on D. striatipennis) ; its head is slightly pubescent.
The bluish specimens differ from the description of D. cwrulescens in the colour
of the clypeus and legs, and in the punctures of the prothorax.
DITROPIDUS SOLITUS sp. nov.
§ Black with a bronzy gloss, basal half of antenne reddish, the apical
half infuseated or black, labrum and basal half of front femora obscurely
reddish. Head, under surface, and legs with sparse pubescence.
Head with dense punctures at base and on clypeus. Eyes large and close
together. Prothorar as wide at apex as along the middle, with fairly dense and
rather small but sharply defined punctures in middle, becoming larger but not
confluent on sides. Hlytra subquadrate; with rows of rather large punctures,
becoming larger’and set in deep striz on the sides; interstices with very minute
punetures. Front legs slightly longer than hind ones. Length (24), 2-2-25 mm.
2 Differs in being more robust, elytra less narrowed posteriorly, and with
smaller seriate punctures, front legs no longer than hind ones, and abdomen
foveate.
Hab.—South Australia: Mount Lofty (S. H. Curnow, A. H. Elston, and
J. G. O. Tepper); New Mecklenburg and Adelaide (Tepper); Moonta and
Kilkerran (Blackburn’s collection No. 1318); Parachilna (Natural History
Expedition, 1917); Quorn (Elston). Victoria: Dividing Range (Blackburn).
New South Wales: Sydney (Dr. E. W. Ferguson and A. M. Lea); Forest Reefs
(Lea).—Type, J. 10846 in South Australian Museum; cotype, (/2318 in Queens-
land Museum.
224 MEMOIRS OF THE QUEENSLAND MUSEUM.
A feebly metallic species, with eyes close together, those of the male
being separated slightly less than the length of the basal joint of antenne, in
the female about equal to that of the two basal joints; on many specimens the
bronzy gloss is hardly in evidence, on some the prothorax has a distinet coppery
eloss; the front legs are often entirely dark, and occasionally the labrum is
conspicuously red. The punctures at the apex of the prothorax are sometimes
almost as coarse as those on the sides; the metasternum is shining, and with
sparse, sharply defined punctures in the middle, but the sides appear shagreened,
owing to the dense and somewhat asperate punctures there. It is close to
D. quadratipennis, but is smaller, less metallic, with smaller punctures and inter-
ocular space not quite the same; also about the size of D. odewahm, but with
coarser punctures, darker legs, and eyes much closer together.
DITROPIDUS TROPICUS =p. nov.
¢ Black, basal half of antennz and sides of labrum reddish. Head,
under surface, and legs with sparse white pubescence.
Head with rather dense partially concealed punctures, median line rather
distinct. Eyes rather close together. Prothorax not twice as wide as the median
length, sides strongly rounded; with dense and sharply defined but not very
large punctures, becoming larger but not confluent on sides. Elytra subquadrate ;
with series of rather large punctures, on the sides set in deep striz, the interstices
between which are costiform posteriorly, the other interstices impuncate or almost
so. Front legs scarcely longer than hind ones. (Length ( 69), 2-3 mm.
Q Differs in being more robust, elytra less narrowed posteriorly, and
with smaller punctures, legs shghtly shorter and thinner, and abdomen foveate.
Hab.—North-west Australia (Blackburn’s collection); Roebuek Bay
(H. H. D. Griffith, his No. 3304, and C. French). Queensland: Thursday
Island (G. E. Bryant) ; Cairns (E. Allen) ; Bowen (Aug. Simson, his No. 88).—
Type, J. 10903 in South Austrahan Museum; cotype, C/2319 in Queensland
Museum.
A feebly metallic species, that appears to occur in abundance at Roebuck
Bay ; the upper surface usually has a vague bluish gloss, the prothorax, especially
in the females, occasionally has a faint coppery one. The distance between the
eyes of the male is slightly more than the length of the basal joint of antenne,
in the female it is about one half more; the female is usually larger than the
male. It is very close to D. solitus, but the eyes are not quite so close together,
sex for sex, the eyes of the male being about as far apart as those of the female
of that species, the prothoracic punctures are somewhat different, and those of
the metasternum are larger and more sharply defined on the sides; in general
appearance it is like D. striatopunctatus but the sides of the prothorax are
nonstrigose ; from D. lobicollis it differs in being smaller, eyes of male slightly
closer together, and prothoracie punetures more sharply defined in the middle;
COLEOPTERA.—LEA. 225
it is also smaller than D. quadratipennis, much less metallic, and prothoracic
punctures differ; D. pygidialis has much smaller punctures on head and
prothorax, and eyes more widely separated.
DITROPIDUS VICARIUS :p. nov.
Black, upper surface bronzy or ecoppery bronze, basal half of antennze
; ] _ A ,
obscurely reddish. Glabrous.
Head shagreened, and with fairly dense but feeble punctures ; median line
vaguely impressed. Eyes rather widely separated. Prothorax about thrice as
wide as the median length, sides strongly rounded; punctures fairly dense and
rather sharply. defined but small, becoming still smaller on sides. EHlytra rather
short ; with rows of very small punctures, becoming larger and set in moderately
deep striz on the sides. Length (¢@ ), 1-5-2 mm.
Q Differs in the usual particulars of the eyes, legs, and abdomen.
Hab.—North Queensland (Blackburn’s collection) ; Cairns (EK, Allen).
New South Wales: National Park and Ourimbah (G. E. Bryant); Sydney
(A. M. Lea).—Type, I. 10865 in South Australian Museum; cotype, C/2320 in
(Queensland Museum.
Slightly narrower and more metallic than D. rotundiformis, prothoracic
punctures decidedly smaller and those of sterna different; the intercoxal process
of the prosternum has a few punctures in front, that of the mesosternum has
a distinct transverse row, but the middle of the metasternum is impunctate.
The prothorax could hardly be regarded as shagreened, although at first glance
it appears to be so; on some of the specimens, from New South Wales the pale
joints of the ems are almost flavous.
DITROPIDUS VAGANS sp. nov.
d Black, sometimes with a slight bronzy gloss, basal half of antennz
flavous, the other infuscated, front legs partly or entirely pale, labrum and
tarsi more or less obscurely diluted with red. Glabrous.
Head shagreened and with very minute punctures, median line scarcely
traceable. Prothorax shagreened and with minute punctures. Scutellwm narrow
and distinct. Length ( dQ ), 1-25-1-5 mm.
Q Differs in the usual particulars of the eyes, legs, and abdomen.
Hab.—Northern Territory: Darwin, on Acacia flowers (G. F. Hill, No.
371). Queensland: Cairns, Charters Towers (Blackburn’s collection) ; Brisbane
(E. M. Hockings). New South Wales: Blue Mountains (Blackburn) ; Wentworth
Falls (Simson’s collection) ; Sydney (Dr. E. W. Ferguson and A. M. Lea);
Galston, Como, and Windsor (Lea). South Australia: Port Lincoln (Blackburn
and Lea); Murray Bridge (Lea); Quorn (A. H. Elston).—Type, 7. 10875 in
South Australian Museum; cotype, C/2321 in Queensland Museum.
P
226 MEMOIRS OF THE QUEENSLAND MUSEUM.
The outlines, eyes, and punctures of sterna are as described in the preeed-
ing species, but it is less metallic, the prothorax as well as the head is shagreened,
and the seriate punctures on the elytra are different; they are small, narrow,
and so close together that the elytra might fairly be regarded as striated
throughout; on the sides, however, the strize are deep and well-defined as on
most species of the genus. In general appearance it is somewhat like large
specimens of D. punctulum, but is more oblong, scutellum narrower and more
distinct, prothorax less opaque, although shagreened, and with more distinct
punctures, and parts of front legs pale; these are sometimes entirely flavous,
or at least decidedly paler than the others; occasionally the knees are infuseated
and sometimes the femora are entireiv dark; on one specimen from Cairns all
the tibie are pale. The punctures on the prothorax, although minute, are -
sufficiently distinct on close examination, but on the head they are almost
invisible.
Var. DUBIUS ver. nov.
Some specimens (sexes) are structurally so close to this species that I
have not ventured to give them more than a varietal name. They differ in being
slightly more rounded, prothorax with scarcely visible punctures, and polished
but becoming subopaque on sides; the legs are all black, or at least very obscure.
Hab.—New South Wales: Sydney (W. du Boulay and G. E. Bryant) ;
Illawarra (Bryant) ; Hornsby (C. Gibbons).
DITROPIDUS BREVICOLLIS sp. nov.
d Black; head, antenne (club infuscated), palpi, and legs more or less
flavous. Glabrous.
Head subopaque and with searcely visible punctures, median line very
feeble. Eyes close together. Prothorar more than thrice as wide as the median
length, sides strongly rounded; punctures minute. Elytra about as long as the
basal width; with rows of distinct punctures, becoming smaller posteriorly, and
on the sides set in deep striw. Length (¢ 9), 1-75-2 mm.
2 Differs in being more robust, infuscation of head extended to cover
clypeus, elytra less narrowed posteriorly, and abdomen foveate.
Hab—North Queensland (Blackburn’s collection) ; Cairns distriet (A. M.
Lea).—Type, I. 10866 in South Austrahan Museum; cotype, C/2322 in Queens-
land Museum.
The eyes are closer together than in any of the preceding small species;
the distanee between them in the male is slightly iess than the length of the
basal joint of antenne, in the female it is slightly more. From D. vicarius it
differs also in being non-metallic, prothoracic punctures smaller and legs paler:
from D. tranguillus in having the head opaque, and lateral striew of elytra
deeper. The legs are often entirely pale, but frequently the hind femora, and
sometimes the middle ones as well, are deeply infuscated. Some specimens,
COLEOPTERA—LEA. 227
ea
from Cairns, differ in having scarcely visible prothoracic punctures, and legs
more brightly flavous. A female, from Mount Tambourine, possibly belongs
to the species, but differs from normal females in having slightly larger pune-
tures on prothorax (although still small), and legs black, with the tarsi brown.
Var. ?—Twelve specimens (Northern Queensland and Bundaberg, Black-
burn’s collection; Cairns district, F. P. Dodd; and Kuranda, H. Hacker) are
so extremely close in general appearance to this species (they even differ sexually
in the colour of the head), that it seems undesirable to name them as distinct,
but they certainly have the eyes more distant, those of the male being as widely
separated as in the female of the typical form, and those of the female about
one-third more than in its female; placing specimens side by side, the differences
are at once apparent. One male has the prothorax reddish, with its middle
infuseated.
DITROPIDUS OPACICEPS sp. nov.
S$ Black; clypeus, labrum, basal half of antennx, palpi, and parts of
legs, more or less obscurely flavous or reddish. Glabrous.
Head shagreened and with very minute punctures; median line feeble.
Eyes rather widely separated. Prothorax about thrice as wide as the median
length, sides strongly rounded; punctures sparse and minute. Elytra subquad-
rate; with rows of fairly distinct punctures, becoming smaller posteriorly and
on the sides set in fairly deep striew. Length, (¢ 9), 1-75-2 mm.
Q Differs in being more robust, clypeus darker than labrum, and in the
usual particulars of the eyes, legs, and abdomen.
Hab.—New South Wales: Sydney, Galston, Como. Queensland: Mount
Tambourine (A. M. Lea); Dalby (Mrs. F. H. Hobler); Bribie Island (H.
Hacker and Lea)—Type, /. 10930 in South Australian Museum; cotype, C/2323
in Queensland Museum.
The outlines and general appearance are almost as in the preceding
species, but the prothorax is almost—on some specimens quite—impunctate, and
the distance between the eyes, sex for sex, is about twice as great; the prothorax
and shagreened head readily distinguish the species from D. tranquillus. The
legs are sometimes entirely pale, but usually the femora, or at least the hind
ones, are deeply infuscated; the head could hardly be regarded as reddish, but
it is not of the deep shining black of the prothorax. The prosternum and meta-
sternum are fairly densely punctate in the middle, the metasternum is shining
and sparsely punctate there. The distance between the eyes of the male is
about equal to the length of the five basal joints of antenne; in the female it is
about one-fourth more. The only specimen from Mount Tambourine has darker
legs than usual, and the middle of its labrum is infuscated.
GELOPTERA TETRASPILOTA Lea.
A specimen of this species, from the Queensland National Park, has an
infuscate spot on the suture between the two large median spots, and these are
almost connected with the sides, —
228 MEMOIRS OF THE QUEENSLAND MUSEUM.
EDUSA DECEMLINEATA sp. nov.
2 Dull coppery bronze, elytra somewhat purplish, under surface more
shining; labrum, basal half of antennz, basal joints of palpi, and parts of legs
reddish. Densely clothed with short, ashen or white pubescence, on the elytra
forming ten distinct lines.
Head shagreened and with minute punctures, distinct only on front of
clypeus, median line feebly impressed and confined to basal half. Antenne
long and thin. Prothorax shagreened and indistinctly punctate. Hlytra densely
and finely granulate-punctate, with larger, but not confluent, punctures scattered
about, and forming geminate rows. Femora stout, front pair acutely dentate.
Length, 5-25-5-75 mm.
Hab.—New South Wales: Dorrigo (Dr. R. J. Tillyard).—Type, C/2324
in Queensland Museum; cotype, J. 11990 in South Australian Museum.
There are ten well-defined lines of pale pubescence on the elytra, each line
bounded by geminate rows of punetures; the tip of the clypeus and a small
space near each antenna are metallic green. To associate the species with others
in my table!® a new section would be required, as it could hardly be placed in F,
as the elytra are conspicuously striped, and as they are without longer hairs it
could not be referred to FF.
COLASPOIDES FASCICULATA ¢p. nov.
S Testaceous with a conspicuous brassy-green gloss; under surface,
antenne, palpi, and legs paler and not metallic; tips of seventh, eighth, and
eleventh joints of antenne infuscated.
Head with fairly dense but unevenly distributed punctures; with a shallow
median line. Third joint of antenne distinctly shorter than fourth. Prothorax
about twice as wide as long, sides evenly rounded, all angles dentate; middle with
rather sparse punctures, somewhat larger than those on head, but becoming
larger and denser on sides. Scutellum impunctate. Elytra rather elongate,
parallel-sided to beyond middle, with rather dense and fairly large punctures
about base, more crowded and transversely confluent behind shoulders, much
smaller and seriately arranged posteriorly. Abdomen shallowly depressed, with
long straggling hairs, fourth segment carinated along middle, fifth irregularly
impressed. Legs rather long, front femora stout and acutely dentate; basal
joint of front and of middle tarsi large and slightly concave on lower surface ;
hind tibiw rather thin and longer than the others, gently emarginate on lower
surface near apex, a loose fascicle before the emargination, tip with a long curved
faseicle ; basal joint of hind tarsi shorter than the rest combined. Length ( ¢ @),
7-7-5 mm.
Q Differs in being less elongate, abdomen simple, legs shorter, hind tibie
simple, and basal joint of front and middle tarsi shorter, and much thinner.
10 Lea, Trans. Roy. Soc. 8. Aust., 1915, p. 193.
COLEOPTERA.—LEA. 229
Hab.— Queensland: Blackall Range—Type, male and female, C/2325 in
Queensland Museum; cotype, male, in South Australian Museum.
With the general outlines and appearance of C. tarsalis and C. picticornis,
with which it would be associated in my table,*' but readily distinguished from
those and all other described species by the shape and clothing of the hind
tibiz of the male; the third-sixth joints of antenne of the male have some
rather long hairs on the under surface; the tips of the antennz of the female
are missing, and only its eighth joint is infuscated.
MACROGONUS QUADRIVITTATUS Jzec.
Of sixteen specimens of this species before me five have the elytra
purple or blue, except that the margins and suture are narrowly flavous; on the
others the blue (sometimes almost black) part of each elytron is divided into
two by a flavous vitta, the vitta narrow and hardly passing the middle on some
specimens, wider and almost touching the apex on others; on twelve the basal
spot of the head is divided by a narrow longitudinal vitta, on the others the spot
is divided into four parts, each of which is hardly more than a stain.
MACROGONUS BIFOVEICOLLIS sp. nov.
6 Flavous, elytra dark blue or greenish blue, tarsi coppery green, second
to fifth joints of antenne metallic blue or coppery green, the following ones
opaque purple, tips of mandibles blackish.
Head with a fairly deep interocular impression, connected with the base
by a distinct median line; punctures sparse, irregular, and mostly small. Pro-
thorax about twice as wide as long, each side with a large obtuse median tooth,
front and hind angles slightly armed, between median and hind teeth a deep
notch; with a large, somewhat transverse, deep fovea on each side of middle;
punctures sparse and rather small, but sharply defined. Scutellum triangular,
slightly longer than wide. Elytra much wider than base of prothorax; each
with four irregular fovea: two on the sides behind the shoulder, one halfway
between the front one of these and the suture, the other slightly nearer the
second marginal fovea than the suture; with regular rows of rather small
punctures. Length (¢@ ), 10-5-11-5 mm.
Q Differs in having the prothcrax smaller and Jess transverse, sides
gently rounded and unarmed in middle, notch near hind angles less pronounced,
dise nonfoveate, and with slightly larger punctures; elytra with lateral fovee
not traceable, and the others smaller, antenne and legs slightly shorter and
thinner, and knees and most of tibize dark.
Hab.— Queensland: Mount Tambourine (H. Hacker, Nos. 741 and 742,
R. Illidge, and A. M. Lea).—Type, J. 4771 in South Australian Museum ; cotype,
C/2326 in Queensland Museum.
11 Lea, Trans. Roy. Soc. 8. Aust., 1915, p. 279.
230 MEMOIRS OF THE QUEENSLAND MUSEUM.
Of four males before me three have most of the abdomen infuscated,
but on the other, and on four females, it is no darker than the rest of the under
surface. The medio-lateral tooth on the prothorax of the male is somewhat
larger but less acute than on M. quadrivittatus; the male is unmistakably a
WVacrogonus, but I am unable to point out how the female may be distinguished
from Macrolema.
Famitry EROTYLID 2.
‘ LANGURIA ALBERTISI Har.
L. vulgaris Har.
L. vandepolli Fowler.
L. australis Macl.
: The species of Languria so abundant in the Cairns district (it oceurs also
at Bowen, Cooktown, and Melville Island, and there are specimens in the South
Australian Museum from the Madang district of New Guinea, and from Aru)
has the head and prothorax red, and elytra black, with a bluish or purplish gloss;
the antennz and legs are black and the mesosternum, metasternum, and abdomen
are black or deeply infuscated; this is the most common form, and has been
named vulgaris, vandepolli, and australis. On 28 specimens before me the under
surface is red, and the legs and antenne partly red, agreeing with the form
named albertisi; the differences are probably due to immaturity, as the two forms
occur freely together, and there are others before me in which the normally
black parts are infuscated in varying degrees. The eighth joint of the antenne
is shghtly larger than the seventh, but distinctly smaller than the ninth; on the
males being produced slightly to one side, it might fairly be regarded as part of
the club, but on the females it evidently could not be so regarded; in albertisi
the club was described as three-jointed, and in vulgaris as four-jointed, but J
believe these names to belong to but one species. Blackburn has already noted
vandepolli as a synonym of vulgaris, and australis has now to be noted as another.
The size ranges 2-4-5 mm.
In Wytsman’s Genera Insectorum, Fase. 78, in which the Langurides are
dealt with by Fowler, albertist was referred to Stenodastus, vulgaris to Caio-
languria, vandepolli to Anadastus, and australis was overlooked. Of the other
species recorded from Australia LZ. militaris has the suture reddish (of 121
specimens of albertist before me not one has the suture reddish), and DL. picea
has the head and prothorax nigropiceous (several specimens of albertisi have the
head infuseated at base).
EPISCAPHULA OPACA Crotch.
A remarkably distinct species, the curved red mark on each shoulder is
sometimes entire, but is usually broken up into two parts; on one specimen the
markings are reduced to a dull spot on the shoulder, and another near apex of
each elytron, the spots so dark that, to the naked eye, the upper surface appears
entirely black. Specimens before me are all from Queensland: Cairns, Port
Curtis, Bowen, and Bluff.
COLEOPTERA.—LEA. 231
EPISCAPHULA BIFASCIATA Macl.
Macleay described the elytra of this species as ‘‘finely striate-punctate’’ ;
on the flavous fasciz (these become reddish with age) the series of punctures,
through ‘‘waterlogging,’’ sometimes appear as fairly distinct, but they are really
so small that where not waterlogged they are scarcely visible. There is a speci-
men of the species, from Aru, in the South Australian Museum, with the pro-
thoracic spots smaller than usual, but in other respects agreeing well with typical!
specimens from Cairns.
EPISCAPHULA AUSTRALIS Boi.
E. froggatti Mael., var.
On the typical and common form of E. australis the elytra, as described
by Boisduval, have nine black spots: three basal, three antemedian, two post-
median, and one apical; but in a fairly common variety named froggatti by
Macleay, and figured by Kuhnt’? the antemedian spots are combined to form a
zigzag fascia. The series of punctures on some specimens appear to be large
and close together, but this is due to ‘‘ waterlogging’’; on examining such speci-
mens from the sides the punetures will be seen to be small and distant, although
fairly sharply defined. The species occurs in Northern Territory (including
Melville Island), Queensland, New South Wales, Victoria, and Tasmania; and
there are also many specimens of the variety froggatti, in the South Austrahan
Museum, from the Madang district of New Guinea.
EPISCAPHULA BREVICORNIS Blackb.
I am unable to distinguish this species structurally from the preceding,
but the prothoracie and elytral markings are more extended, and on a some-
what different plan. On the prothorax the markings may consist of-a large
semi-double mediobasal blotch, and an isolated spot on each side, or they may
be all conjoined; on the elytra the black markings may be continuous almost to
each side, or with a projection from the red portion on each elytron, so that a
large black square or oblong is isolated on each shoulder, the two large black
subapical spots may be completely isolated, or conjoined across the suture.
EPISCAPHULA FLAVOFASCIATA sp. nov.
Black, with flavous or reddish-flavous markings.
Head with rather dense but very small punctures. Antenne with third
joint twice as long as fourth. Prothorar about twice as wide as long, sides
oblique, front angles acute and each with a puncture, a vague basal depression
on each side of middle; with smal! punctures much as on head, and a few of
larger size scattered about. Elytra with minute punctures; sutural striew distinct
only on apical third. Abdomen with inconspicuous coxal lines. Length, 7-10 mm.
12 Kuhnt in Wytsman’s Genera Insectorum, Fasc, 88, pl. iv, fig. 11.
232 MEMOIRS OF THE QUEENSLAND MUSEUM.
Hab.—Queensland: Brisbane (R. Illidge); Mapleton; National Park
(H. Hacker). New South Wales: Dorrigo (R. J. Tillyard and W. Heron) ;
Richmond River (A. M. Lea).—Type, J. 11767 in South Australian Museum;
cotype, (/2327 in Queensland Museum.
The markings are slightly variable, but on the types the prothorax is
pale, with three large spots occupying the basal half, except for oblique Imes
between them and the sides, the median spot is more than twice the size of the
others; the elytra have an irregular pale fascia at the basal third, touching
neither suture nor sides, and another at the apical third, not quite touching the
suture, continued around the sides and apex till the two parts are interrupted
at the suture (on some specimens the fascia has a narrow extension on each side
of the suture, so that two large black spots are isolated); the prosternum is
flavous, and part of the abdomen obscurely diluted with red; parts of the legs
are also obscurely reddish. Some of the specimens have the dark parts hardly
more than castaneous, and on one such specimen the lateral spots of the pro-
thorax appear as feeble infuscations. The punctures at the base of the head
are larger than those in front, but are usually concealed by the overlappmg
prothorax; the elytra from some directions appear to have feeble rows of
minute punctures, but from most directions the punctures are almost or quite
invisible. At first glance the species appears close to EL. foveicollis, but the head
has much smaller punctures, the larger ones of the prothorax are mostly basal,
and certainly not congested in the front angles, the outer spots are basal instead
of median, the black basal marking of the elytra is continuous from side to side,
instead of twice interrupted to the base, and the apical markings are different.
It is also close to FE. bifasciata, but with three basal dark spots on the prothorax
instead of two, and the front angles less acute. In general appearance some of
the specimens look like large ones of EH. brevicornis, but may be at once
distinguished by the longer third joint of antenne.
EPISCAPHULA INCLUSA sp. nov.
Black, upper surface blackish purple or blackish blue; a large red mark
on each elytron, commencing at the base, and almost touching the scutellum and
shoulder, curved round so as almost to touch the side of the basal third, and then
directed to the suture, which it almost touches just beyond the middle; parts of
under surface and of tarsi reddish.
Head with fairly numerous, sharply defined, but not very large punctures ;
clypeal suture distinct but not deeply impressed. Third joint of antenne twice
the length of fourth. Prothorar at base more than twice as wide as the median
length, sides diminishing in width to apex, front angles produced but not very
acute, submarginal line deep from base to apex; punctures sparser and mostly
smaller than on head, a few slightly larger ones in a feeble depression on each side
of base. Elytra slightly dilated from shoulders to basal fourth; with regular
rows of distinct, but not very large punctures, the interstices with much smaller
ones. Coxal lines of abdomen well defined almost to apex. Length, 6-8 mm.
COLEOPTERA.—LEA. 233
Hab—Queensland (National Museum); Cairns (E. Allen and A. P.
Dodd) ; Mapleton (H. Hacker). Type, J. 11774 in South Australian Museum ;
cotypes, C/2328 in Queensland and National Museums.
On several of the specimens the head and shoulders have a slight mefallic-
ereen gloss; to the naked eye the red markings of the elytra appear to be almost
circular, they almost completely enclose a dark space about the size of the
prothorax; on most of them the abdomen is paler than the rest of the under
surface, and on several each segment, except the basal one, is darker at its extreme
base and apex, so that the middle appears obscurely fasciate ; on several the large
apical joimt of the palpi is conspicuously reddish. The species is wider than
usual, and the elytral markings are very different from those of any previously
named Australian one.
THALLIS PERPLEXA Blackh.
Numerous specimens from Cairns, .Cooktown, Coen River, and Darnley
Island, and one from Manumba in the Madang district of New Guinea, agree
with the description of this species; which is possibly also 7. bizonata, but that
species was described as having the prothorax ‘‘very finely punctate’’ and the
elytra as ‘‘very faintly striate-punctate’’; on the specimens before me the
prothoracic punctures are of moderate size and sharply defined, and the series
of punctures on the elytra are larger than usual, and sharply defined even to
the apex.
THALLIS MACLEAYI Blackb.
Readily distinguished from several somewhat similarly coloured species
by the wide prothoracic margins, the spots close to the suture vary somewhat
in intensity of colour; it occurs in Queensland (Brisbane and Bowen), Northern
Territory (Darwin), and North-west Australia (Derby and Port George IV).
THALLIS INSUETA Crotch.
The four large spots, on the elytra of this species, vary somewhat in size
and intensity, but are always conspicuous. The species occurs in Queensland,
New South Wales, Victoria, Tasmania, and South Australia.
THALLIS AUSTRALIZ sp. nov.
Dark castaneous brown, elytra with two reddish fasciw. Densely clothed
with dark pubescence, becoming almost golden on the fascizw, in addition with
numerous sub-erect hairs; under surface and legs with almost white pubescence.
Head sub-opaque, and with crowded but sharply defined punctures.
Antenne rather short, second to eighth joints sub-equal. Prothorax not twice as
wide as long, sides gently rounded and slightly uneven; punctures much as on
head. Elytra no wider than widest part of prothorax, parallel-sided to near
apex; with regular rows of fairly large punctures, the interstices with numerous
234. MEMOIRS OF THE QUEENSLAND MUSEUM.
small ones. Prosternum with rather coarse punctures on sides, across. middle
some of them transversely confluent, interecoxal process small and almost parallel-
sided. Abdomen with coxal lines obscured by clothing. Length, 4-5 mm.
Hab.— Queensland: Cunnamulla (H. Hardeastle) ; Bowen (Aug. Simson,
No. 554) ; Dalby (Mrs. F. H. Hobler). New South Wales: Narromine (Dr. E. W.
Ferguson) ; Condobolin (H. J. Carter from — Halligan) ; Belltrees (8. Jackson) ;
Albury (A. M. Lea). South Australia: Adelaide. North-west Australia: Derby
(W. D. Dodd).—Type, J. 72006 in South Australian Museum; cotype, C/2329
in Queensland Museum.
Similar in size to and densely pubescent like 7. erichsoni, but without a
small red spot on each side of the apex, and hence the pale markings of the elytra
in two series only (constant on twenty-eight specimens), the dark median fascia
considerably wider, and not zigzagged, and the punctures somewhat stronger.
Of the pale fasciew the first occupies the basal two-fifths of the elytra, except for
a large subquadrate patch adjacent to the scutellum (the spot is sometimes rather
feebly infuseated), the second is at the apical third, the part on each elytron is
convex on its anterior margin, concave on its posterior and narrowed to the
suture; the under surface and legs are somewhat paler than the head and _
prothorax. Two of the specimens from Derby have the dark parts black; on a
few of the Queensland ones they are almost black.
THALLIS MELANCHOLICA sp. nov.
Black; legs, antennz, and palpi more or less reddish; abdomen obscurely
diluted with red. Rather densely clothed with sub-erect, rusty pubescence.
Head with dense and fairly large punctures. Antenne with fourth joint
about two-thirds the length of third, and slightly longer than fifth. Prothorax
about once and two-thirds as wide as long, apex truncated in middle, and notched
near each side, sides slightly and irregularly serrated, a rather deep lne near
each side; punctures shghtly larger than on head, becoming smaller and denser
-in front. Elytra parallel-sided to near apex, with regular rows of fairly large
punctures, becoming smaller posteriorly; interstices with numerous distinet but
rather small punctures. Intercoxal process of prosternum moderately wide,
obtusely pointed. Abdomen with coxal lines traceable to beyond middle of basal
segment. Inner edge of front tibae minutely serrated. Length, 7-9 mm.
JTab.— Queensland: Mount Tambourine (A. M. Lea). New South Wales:
Blue Mountains (Dr. E. W. Ferguson) ; Mount Wilson, Eden (H. J. Carter) ;
Galston (D. Dumbrell and Lea); Sydney (J. J. Walker and Lea). Tasmania
(Aug. Simson, No. 3825); Hobart (Lea) —Type, J. 12009 in South Australian
Museum; cotype, C/2330 in Queensland Museum.
Although the derm of the majority of the specimens is of a shining black, it
appears, to the naked eye, more or less dark rusty brown, owing to the pubescence ;
some of the specimens, however, are really rusty brown with the abdomen paler,
COLEOPTERA.—LEA. 235
but this is probably due to immaturity. The prosternum and outlines are much
as in 7. insueta and T. venustula. One of Mr. Carter’s specimens was labelled
as from West Australia.
EUXESTUS VULNERATUS sp. nov.
Black, a large basal patch on elytra red, muzzle and under surface
castaneous, legs and antenne flavous. Upper surface with sparse, sub-erect
pubescence.
Head evenly convex; with small but sharply defined punctures; a shallow
depression on each side of clypeal suture. Antenne short, club large, slightly
wider than long. Prothorax at base about thrice as wide as the median length,
base bisinuate, apex evenly incurved to middle, margins very narrow, punctures
as on head. Elytra with outlines continuous with those of prothorax, widest at
about basal third; with rows of small punctures. Abdomen with coxal lines
traceable to near apex of basal segment. Length, 1-75-2 mm.
Hab. —Queensland : Little Mulgrave River (H. Hacker); Cairns (C. J.
Wild) —Type, J. 12015 in South Australian Museum; cotype, C /2331 in Queens-
land Museum.
A briefly elliptic, strongly convex species, very distinct by a large blood-
red patch on the elytra; it covers part of the base (leaving a rather narrow
strip of black on each side), then extends rather narrowly to each side, from there
its margin extends obliquely towards the suture, and then is truncated across
the suture itself, at the middle (on some specimens its posterior margin is
rounded) ; the tips of the elytra are usually obscurely diluted with red, and the
dark parts sometimes have a coppery gloss. On one specimen the large patch
is flavous, and the apex of the elytra is conspicuously pale. The clothing of the
upper surface appears to be easily abraded, as several of the specimens are now
almost glabrous. ‘he elytral punctures are all small, but are fairly distinet on
the paler parts.
EUXESTUS BIVULNERATUS sp. nov.
Black, a large blood-red spot on each shoulder, legs, antenne, palpi, and
elytral epipleure reddish flavous. Glabrous. Length, 2-2-25 mm.
Hab.— Queensland : Mount Tambourine (A. M. Lea).—Type, 7. 11784 in
South Australian Museum; cotype, C/2342 in Queensland Museum.
Structurally extremely close to the preceding species, but slightly larger,
elytral markings, which are of the same blood-red colour, extending from each
side to about one-third from the suture, so as to resemble an interrupted fascia ;
elypeal impressions deeper and punctures smaller, especially on the prothorax
and elytra. The dark parts of the upper surface, on ten specimens, are deep
polished black, but on another they have a slight coppery gloss; the extreme
tips of the elytra are sometimes obscurely diluted with red. The coxal lines are
distinct, and enclose a plate on each side, but these are without punctures.
236 MEMOIRS OF THE QUEENSLAND MUSEUM.
Var.—A specimen, from Wollongong, possibly belongs to this species, but
has the elytral markings smaller and much less distinct, appearing as a rather
obscure spot on each shoulder.
EUXESTUS PARKI Woll.
Black; head, tips of prothorax, elytra, and under surface (except meta-
sternum) more or less obscurely castaneous; legs, antenne, and palpi paler.
Glabrous.
Head with rather dense and small, but sharply defined punctures; a small
fovea on each side of clypeus. Antenne short; club large, slightly wider than
long. Prothorax not thrice as wide as the median length, base feebly bisinuate,
apex almost straight, margins very narrow; punctures rather less dense, but
“otherwise as on head. Elytra with outlines continuous with those of prothorax ;
with feeble rows of small punctures; interstices with punctures as on prothorax.
Abdomen with coxal lines inconspicuous. Length, 1-75-2 mm.
Hab.—Queensland: Mulgrave River (H. Hacker) ; Cairns (A. M. Lea).
More evenly elliptic and slightly less convex than either of the preceding
species, and about one-fourth narrower in proportion; at first glance it resembles
several species of Paracymus and other small Hydrophilide. On_ several
specimens the elytra aré obscurely diluted with red about the extreme tips, but
on a few the red almost covers the apical third, although not sharply limited.
A specimen from New Ireland (Edgar R. Waite) is a trifle larger than
Queensland specimens, but I can find no other distinctions.
DIPLOCGELUS MAXIMUS sp. nov.
Blackish brown; legs, antennz, and palpi obscurely reddish. Moderately
clothed with short, depressed pubescence, becoming denser and paler on under
surface; upper surface, in addition, with moderately long, erect, reddish sete.
Head with fairly numerous moderate and small punctures; a deeply
impressed transverse line near base. Antenne rather short; club three-jointed,
apical joint about once and one-half the length of ninth or tenth. Prothorax
about twice as wide as long, sides evenly rounded with margin thickened, hind
angles-rectangular, front angles acute and produeed, rest of apex straight, each
side with three impressed lines, the outer continuous and within the thickened
margin, the next continuous, the other shallow in front, interrupted in middle,
and deep and wide at base, near base with a deeply impressed sinuous line, with
a short median projection; punctures sharply defined and somewhat sparser
than on head, except on margins, where they are crowded. EHlytra almost
parallel-sided to near apex; with rows of fairly large, sub-oblong punctures,
becoming smaller and rounded towards suture and posteriorly; interstices each
with a row of distinct punctures, and with smaller ones scattered about. Length,
8-9 mm.
Hab.—Queensland: Cairns district (H. Hacker and A. M. Lea).—Type,
I. 11787 in South Australian Museum ; cotype, C/2076 in Queensland Museum.
COLEOPTERA.—LEA. 237
Close to D. leai, but distinctly larger, less hairy, and as a result the pune-
tures appear more distinct and the derm shinier, with the median line of the
prothorax not traceable throughout, but represented by a short impression joing
the basal line. The sete on the elytra, when viewed from behind or in front,
are seen to be in regular lines, rows of longer ones on the interstices alternating
with somewhat shorter ones set in the seriate punctures.
DIPLOCGELUS SERICEUS sp. nov.
Blackish; legs, antenne, and palpi obscurely reddish. Densely clothed
with short, depressed, brownish or greyish ‘sericeous pubescence; upper surface,
in addition, with dense, fairly long, erect, reddish seti.
Head: with fairly large but partially concealed punctures, a transverse
impressed line at base. Antenne slightly passing base of prothorax; club three-
jointed, apical joint almost as long as the two preceding combined. Prothorax
about twice as wide as long, sides strongly rounded, front angles produced, rest
of apex straight, with three deep somewhat curved lines on each side, a deeper
and somewhat sinuous line across base; punctures minute, and more or less
concealed. Elytra with sides gently rounded, widest at about basal third; with
- rows of fairly large, sub-oblong punctures, becoming much smaller and rounded
lor Al A=
near suture; interstices with minute punctures. Length, 7-7-75 mm.
Hab.—Queensland: Coen River (W. D. Dodd); Cairns district (A. M.
Lea). —Type, 7. 11788 in South Australian Museum ; cotype, C/2075 in Queens-
land Museum.
About the size and shape of D. leai, but with very different clothing; the
elytra have a curious mottled appearance, owing to numerous sericeous-looking
patches, which alter their positions with the point of view; the arrangement of
the elytral sete is much as on the preceding species, but the pubescence is
different, and the punctures are much smaller; from the prothorax, owing to
the denser pubescence, they appear at first to be absent; the transverse basal
line of the prothorax is also without a median projection.
Vars. ?—Two other specimens, from Cairns, probably belong to this
species ; they have the sete on the upper surface much denser than on the typical
form, scareely half their length, and not at all seriate in arrangement; the
pubescence has a mottled appearance, but on one of them is even denser than on
the types, and on the other sparser, with the result that on one the derm appears
to be sub-opaque, and on the other more polished.
DIPLOCELUS DILATATICOLLIS ¢p. nov.
Dull reddish brown or castaneous, appendages slightly paler. Densely
clothed with short, depressed pubescence, denser and paler on under surface
than on upper, the latter in addition with lines of short, semi-erect sete.
Head with dense and rather small puncturés; a shallow depression on
each side of clypeus. Antennz short; club three-jointed. .Prothorax more than
238 MEMOIRS OF THE QUEENSLAND MUSEUM.
thrice as wide as long, base and apex subequal, front angles slightly produced,
sides strongly and evenly rounded; with ten longitudinal elevations, of which
those near the sides are fairly distinct and continuous, the median ones less
distinct, and almost disappearing near base; punctures dense and small. Elytra
parallel-sided to near apex; with rows of fairly large punctures, becoming
smaller towards suture; interstices with small punctures. Length, 3-3-25 mm.
Hab.— Queensland: Mount Tambourine (C. J. Wild and A. M. Lea).—
Type, J.11791 in South Australian Museum; cotype, C/2078 in Queensland
Museum,
The sides of the prothorax are dilated so that their greatest width is
slightly, but distinctly, more than that of the elytra, a character that at once
distinguishes the species from D. decemlineatus, and from D. fasciatus (on some
specimens of the latter the elytral markings are very feeble); from D. latus,
which has somewhat similar prothoracic sides, it is distinguished by the much
smaller punctures of the entire upper surface. On several specimens there is a
slight infuscation about the middle of the elytra.
Faminry ENDOMYCHIDA.
IDIOPHYES BREVIS Blackb.
The type and only specimen of this species known to Blackburn is now
in the British Museum, but numerous specimens before me appear to belong to
the species, which at first glance seems to be a minute Stenotarsus (except for its
shorter and entirely pale antenne it resembles 8. pisonie in miniature). In the
generic diagnosis Blackburn stated ‘‘ prosternum inter coxas sat angustum, postice
vix productum,’’ but later ‘‘I cannot satisfy myself as to whether its prosternum
projects slightly or not at all clear of the front coxe.’’ Examining unset
specimens it is difficult to see the end of the intercoxal process clearly, but on
removing the prothorax from the hind body the intercoxal process appears
rather acute, produced beyond the coxe, with a notch in the mesoternum for
its reception. Arrow states'® that it belongs to the genus Exrysma, of which Csiki
in the Catalogue of Endomychidw records species only from America; as the
genus was first recorded from Central America by Gorham,'* and two of the
species figured’ are very different in appearance from the Australian ones,
I prefer to retain the name /diophyes for the latter.
Hab.— Queensland: Brisbane (many specimens from wattle blossoms in
July); Mulgrave River. New South Wales: Glenfield (many specimens from a
nest of termites,. Coptotermes sp.) ; Forest Reefs. South Australia: Adelaide
(one specimen from a nest of the same species of Coptotermes) ; Kangaroo
Island. Tasmania: Kempton.
13 Arrow, Trans. Ent. Soc. Lond., 1920, p. 3.
14 Gorham; Biol. Cent. Amer., Col., vii, p. 145.
15 Lc., pl. vii, figs. 14 and 15 (EZ, orbicularis and E. ? tenuicornis).
COLEOPTERA—LEA, 239
STENOTARSUS PISONIZ sp. nov.
Reddish or castaneo-flavous, antenne partly black or blackish. Moderately
densely clothed with pale upright hair.
Head almost impunctate, with a shallow interocular impression. Antennze
short; club rather stout. Prothorax widely transverse, sides strongly rounded,
apex much narrower than base and semicircularly emarginate, sublateral striz
deep, dilated at base; almost impunctate. Elytra at base slightly wider than
prothorax, sides gently rounded; with regular rows of fairly large punctures, in
feeble striz; interstices with small punctures. Length, 3-5-4-5 mm.
Hab.—Queensland: Cairns district (F. P. Dodd, C. J. Wild, and A. M.
Lea) ; Little Mulgrave and Coen Rivers (H. Hacker).—Type, J. 12024 in South
Australian Museum; cotype, C/2336 in Queensland Museum.
Structurally close to S. arithmeticus, but entirely pale except for the
antenne ; of these the three terminal joints appear to be always black, the four
or five preceding ones vary from blackish to hardly darker than the basal ones.
Some of the specimens are paler and some more densely clothed than others,
probably due to better preservation. Many of the specimens from Mr. Dodd
were trapped by sticky seeds of Pisonia brunoniana.
STENOTARSUS PICTICOLLIS sp. nov.
Reddish flavous, three spots on prothorax conjoined at base; scutellum,
two large transverse spots on elytra, most of antenne, and femora black. Length,
3-25-3-0 mm.
Hab.—Queensland: Brisbane (H. Hacker and R. Illidge).—Type, I. 12025
in South Australian Museum; cotype, C/2337 in Queensland Museum.
Structurally close to S. arithmeticus, but head and two spots on prothorax
pale, the blotch on each elytron before the middle not 3-shaped, and without
apical spots. It is shghtly narrower than the preceding species, but the sculpture
and clothing are as noted for it. The dark parts of the prothorax are a large
equilateral triangle, and a spot about two-thirds of its size on each side, thus
leaving a large pale subtriangular spot on each side of apex; on the elytra the
spots are of somewhat irregular shape, and appear like a fascia widely interrupted
in middle, and not touching the sides; three or four of the basal joints of antennz
are obscurely diluted with red, and the tibie and coxe are usually deeply
infuseated,
STENOTARSUS QUINQUENOTATUS sp. nev.
Red or reddish flavous; a subtriangular spot on prothorax, scutellum,
four spots on elytra, antennz (basal joints obscurely diluted with red), femora,
and tibizw black. Length, 3-3-25 mm.
Hab.— Queensland (Blackburn’s collection). New South Wales: Gosford
(H. J. Carter) ; Ourimbah (Dr. E. W. Ferguson) ; Wollongong, Sydney (A. M.
Lea).—Type, J. 120.26 in South Australian Museum; cotype, C/2338 in Queens-
land Museum.
240 MEMOIRS OF THE QUEENSLAND MUSEUM,
Structurally very close to the preceding species, and with similar clothing,
but elytral punctures rather less regular, the prothorax with but one dark spot,
and the elytra with four placed transversely across the middle, of the latter
the inner spots are oblong-elliptic and somewhat larger than the outer ones,
which are circular.
Famitry CORYLOPHIDA.
APHANOCEPHALUS BIMACULATUS sp. nov.
Black; elytra with two large, round, red spots before the middle; antennzx
(except club), palpi, and tarsi reddish. Upper surface shining and almost
glabrous, under finely pubescent.
Head with dense small punctures. Antenne moderately long, first joint
stout, ninth forming a one-jointed club. Prothorax at base about four times as
wide as the median length, sides strongly rounded and finely margined, apex
gently ineurved to middle and about half the width of base; with fairly dense,
minute punctures. Elytra with outlines continuous with those of prothorax,
and with somewhat stronger margins; with fairly dense and sharply defined
punctures, mostly rather small. Length, 2-2-25 mm.
Hab.— Queensland: Bowen (Aug. Simson’s No. 1556); Mackay (R. E.
Turner).—Type, 7.11800 in South Australian Museum; cotype, C/2339 in
Queensland Museum.
An oblong-elliptic, deep-black species, with two conspicuous red spots on
elytra; of the nine specimens before me the elytral spots are alike on all, except
for a slight variation in size and sharpness of definition. The punctures on the
elytral epipleure, sides of metasternum, and basal segment of abdomen are denser
and somewhat stronger than on the elytra.
APHANOCEPHALUS QUADRINOTATUS sp. nov.
Black; tarsi and four spots on elytra reddish; antenne (except club),
palpi, parts of tibie, and elytral epipleure obscurely reddish. Moderately
clothed with pale, subdepressed pubescence. Length, 1-75 mm.
Hab.— Queensland: Mount Tambourine, and Cairns district (A. M. Lea).
—Type, 7.11799 in South Austrahan Museum; cotype, C/2340 in Queensland
Museum.
The shape and margins are as in the preceding species, but the size is
smaller, punctures on elytra somewhat larger, those on the basal segment of
abdomen and sides of metasternum denser but smaller than on elytra, and those
on the elytral epipleure still smaller. The first spot on each elytron is larger
than the other, curved like an inverted comma, commences near the shoulder,
and terminates in the middle about one-fourth from the suture; the second one
varies considerably in size and shape, and is placed at the apical third; on one
specimen the spots on each elytron are conjoined, on another they are conjoined
on the right elytron (so as to appear like an S) but are free on the left.
[MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Part III, 1921.]
By Authority: ANTHONY JAMES CUMMING, Government Printer, Brisbane.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Parr IV.
(ISSUED DECEMBER 19, 1922.)
NOTES AND ILLUSTRATIONS OF QUEENSLAND
BISEEES,=No. 3. |
By ALLAN R. McCuLLocH, ZooLocist, AUSTRALIAN MUSEUM.
(By permission of the Trustees of the Australian Museum.)
(Plate XIV, fig. 1.)
A SMALL -collection of fishes from Queensland waters has been submitted
to me for examination by the Director of the Queensland Museum. Most of these
were obtained by the trawler “ Bar-ea-mul” during her investigations on the
Queensland coast. They include several species not previously recognised from
Australian waters.
Famity CLUPEIDA.
ILISHA HCVENNI Bleeker.
Pellona haevenii Bleeker, Verh. Bat. Gen. xxiv, 1852, Haring. p. 21. Jd. Weber and Beaufort,
Fish. Indo-Aust. Arch. ii, 1913, p. 86, fig. 29.
Ilisha hevenii Bleeker, Atlas Ichth. vi, 1872, p. 117, pl. cclxix, fig. 2.
A specimen from Queensland, 163 mm. long, is the first of the genus to be
recognised from Australian waters. It is similar to a smaller specimen from
Amboyna, which was part of Dr. Bleeker’s collection, but differs from his figure in
having colourless fins and a dark ill-defined shoulder-spot.
Locality Between Cairns and Rockhampton, Queensland.
Famity MENIDA.
MENE MACULATA Bloch & Schneider.
Mene maculata (Bloch & Schneider) Valenciennes, Regne Anim. Illustr. Poiss., 1843, p. 139, pl.
Ixii, fig. 2. Id. Day, Fish India, 1876, p. 249, pl. 53, fig. 5.
Two specimens, 163 and 172 mm. long, were trawled between Cairns and
Rockhampton, Queensland. This monotypic family has not been previously
recognised from Australian waters.*
*Editor’s Note.—Day’s figure of Mene maculata has been reproduced from “ Fishes of
India,” pl. 53, to illustrate this new Australian record (Plate xiv, fig. 2).
Q
24? MEMOIRS OF THE QUEENSLAND MUSEUM.
Famity SPARIDA.
Gexus ARGYROPS Swainson.
Argyrops Swainson; Nat. Hist. Class. Fish, &c., ii, 1839, p. 221 (spinifer).
The similarity of Argyrops and Pagrosomus Gill has been noted by Jordan
and Thompson (Proc. U.S. Nat. Mus. xli, 1912, p. 575). Having compared the
genotypes of both genera, I find that they can only be separated by the following
characters :—
a. Two rows of large teeth on each side of the upper jaw, and an inner row of much smaller ones.
b. Dorsal spines filamentous; depth of the body greater than half its length to the hypural
joint oe 3% sis 2 aie : : Argyrops
bb. Dorsal spines not produced ; depth of the body less than half its length to the hypural
joint or se ahs on 45 ae a rs jie .. Pagrosomus.
ARGYROPS SPINIFER Forskal.
Sparus spinifer Forskal, Descr. Anim., 1775, p. 32. Jd. Bleeker, Atlas Ichth. viii, 1877, p. 109,
pl. 213, fig. 3.
Pagrus spinifer Day, Fish. India, 1875, p. 138, pl. xxxiii, fig. 5. Id. Kent, Gt. Barrier Reef, 1893,
pp. 285 and 369, pl. xliv, fig. 2.
A specimen, 144 mm. long to the hypural joint, forwarded for identification
by the Director of the Queensland Museum, differs from a smaller Indian example
only in minor details. The second to the sixth dorsal spines are filamentous, the
former reaching well beyond the hypural joint. Its general colour appears to have
been pink with somewhat darker crossbars.
Locality.—The species has been recorded from Port Denison by Kent. This
specimen was secured at Peel Island, Moreton Bay, Queensland.
Famity CHATODONTID.
HOLACANTHUS, CHZTODONIOPLUS, CONSPICILLATUS Waite.
Holacanthus conspicillatus Waite, Rec. Austr. Mus. iii, 7, 1900, p. 203, pl. xxxv.
Ogilby’s record of H. conspicillatus from Queensland (Mem. Qld. Mus. iii,
1915, p. 114) was based upon specimens properly referable to H. personifer
McCulloch. He regarded the two as synonymous, but a comparison of five specimens
of personifer with three of conspicillatus shows no intermediate forms, although
the former varies considerably in its colour-marking.
A fine example, 205 mm. long, agrees in all details with Waite’s types from
Lord Howe Island. It was obtained near the Capricorn Group, Queensland.
Famity HEPATIDA.
ZANCLUS CANESCENS Linne.
Zanclus canescens and Z. cornutus (Linn) Bleeker, Atlas Ichth. ix, 1878, pp. 77-78, pl. 366, figs.
ESE -
Zanclus cornutus Macleay, Proc. Linn. Soc. N. 8. Wales v, 1881, p. 548. Jd. Stead, Proc. Linn.
Soc. N. S. Wales xxxiv, 1909, p. 274.
QUEENSLAND FISHES.—McCULLOCH. ; 243;
Localities—This widely distributed species was included in Macleay’s
Catalogue of Australian Fishes on the strength of an old collection specimen sup-
posed to have been collected in Australian waters. ahve, fies 1.)
? Tetrodon cinctus Richardson, Zool. Samarang, Fishes, 1848, p. 20.
Canthigaster cinctus Jordan and Evermann, Bull U.S. Fish Comm. xxiii i, 1905, p. 433, fig. 189.
A beautifully preserved specimen, 131 mm. long, agrees with Jordan and
Evermann’s figure quoted above, and differs from C. valentint in the disposition
of its dark cross-bands. It is from near the Capricorn Group, Queensland.
CANTHIGASTER VALENTINI Bleeker.
Tropidichthys valentini Bleeker, Nat. Tijd. Ned. Ind. iv, 1853, p. 130.
Canthogaster valentyni Bleeker, Atlas Ichth. v, 1865, p. 80, pl. ecvili, fig. 1.
Canthigaster valentini McCulloch, Proc. Linn. Soc. N. 8. Wales xxxvi-3, 1911, p. 423.
Two specimens, collected at Murray Island, Torres Strait, were recorded
by McCulloch.
CANTHIGASTER BENNETT! Bleeker.
Tetrodon ocellatus Bennett, Fish Ceylon, 1828-30, pl. xxi (name preoccupied).
Tropidichthys bennetti Bleeker, Nat. Tijd. Ned. Ind. vi, 1854, p. 504.
Canthogaster ocellatus Bleeker, Atlas Ichth. v, 1865, p. 80, pl. cexiv, fig. 5.
Canthigaster bennetti McCulloch, Proc. Linn. Soc. N. 8. Wales xxxvi, 1911, p. 305.
Several specimens from Murray Island, Torres Strait, were recorded by
McCulloch.
CANTHIGASTER CALLISTERNUS Ogilby.
Tetrodon callisternus Ogilby, Mem. Aust. Mus. ii, 1889, p. 74, pl. ii, fig. 5.
Eumycterias callisternus Ogilby, Mem. Qld. Mus. i, 1912, p. 62.
Recorded from Southport, Queensland, by Ogilby. Several specimens of
various sizes are in the Australian Museum from Lord Howe Island.
946 MEMOIRS OF THE QUEENSLAND MUSEUM.
AN ICHTHYOSAURIAN SKULL FROM
QUEENSLAND.
By HeBerR A. Loncman, F.L.S., DiREcToR, QUEENSLAND MUSEUM. .
(Plates XV and XVI and Text-figures 1 and 2.)
The remains which are the subject of this paper were found at Galah Creek,
about twelve miles from Hughenden, in the Rolling Downs formation (Lower
Cretaceous) of Western Queensland, and were collected, forwarded, and kindly
donated to the Queensland Museum by Mr. 8. Dunn and Mr. William Elliott in
May, 1914. It is my pleasant duty heartily to thank these gentlemen for their
enthusiastic work in securing this large and valuable specimen for our collections.
MatTerRIAL.—As will be seen from the profile view, illustrated in Plate XV.,
this large skull is in six pieces. The extreme end of the rostrum is missing, but,
judging from the structure of the anterior part preserved, only a small portion
would be needed to complete the skull. Gilmore’ has pointed out how frequently
the extreme anterior segment is missing in Ichthyosaurs, and how fractures are
caused by the cracking of specimens when enclosed in an elongate concretionary
mass.
The skull is massive, with a maximum length (mandibular) of 1,026 mm.,
and a maximum width (articular area of mandible) of 395 mm. It is evident that
great changes have taken place since it came to rest. As a result of tremendous
vertical pressure, the whole of the teeth, with the exception of broken roots, have
been forced from the continuous dental grooves, characteristic of Ichthyosaurus,
and the premaxillary and mandibular rami are now in juxtaposition. Fortunately,
many of the teeth have been preserved, mostly as fragments, on the lateral and
lower surfaces of the jaw. In the posterior part of the skull there are still greater
evidences of changes under intense pressure. On the left-hand side the orbit has
been crushed down and its original contours are not distinguishable. As a result
of this lateral torsion, the mandible has been somewhat displaced to the right. The
supratemporal fossz are preserved in fairly symmetrical condition. Great difficulty
has been experienced in studying some of the component parts. The distortion of the
skull is accompanied by a very close investiture of the remains by a fine hard lime-
stone matrix, which in places is almost indistinguishable from the actual fossil.
The matrix involving Cratochelone berneyi,? described by the author in 1915 from
the same district, was very similar in texture. This investing material evidently
penetrated the skull after the decay of cartilage, cementing the disrupted elements
together.
*C. W. Gilmore, Mem. Carnegie Mus., Pitts., II, 1905, p. 80.
* H. A. Longman, Mem. Qld. Mus., III, 1915, p. 25.
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On the upper part of the skull the elongate premaxillz can be traced back
for a distance of 615 mm. to the narial openings. Between these two bones in the
superior surface is a well-marked symphysial groove. The nasals are exposed from
beneath the premaxille at about the anterior third of the length of the skull, and
at first are on a lower plane than the hemispherical superior borders of the diverging
premaxille, forming a triangular recess. Further back there is a secondary
triangular depression, the borders of which are parallel to the anterior recess, but
this is entirely internasal. Lateral divisions of the nasals extend outwards beyond
the frontals towards the superior border of the orbits. Sutures with post frontals
cannot be traced.
The external narial openings can be seen on both sides, but they are distorted.
A semicircular raised border is present behind the openings. Incomplete maxille
are present, but the sutures between them and the lachrymal bones and the jugals
cannot be positively traced on either side.
In the region of the frontals a remarkable rectangular raised process was
present in the undisturbed fossil. On careful development this proved to be mainly
matrix closely investing a troughlike depression, with raised lateral borders, as
may be seen in Plate XV. At first this was thought to be a veritable raised bony
border surrounding the pineal foramen, and suggesting an unusual development of
“the third eye,” but the true foramen is apparently situated in a more posterior
position. This closely adpressed structure consists of two parallel bars, thinly
joined anteriorly ; the bars are 90 mm..in length and are symmetrically disposed
at a distance of 10 mm. from the median line. If this structure is actually in situ,
which seems unlikely, it would demand generic recognition. The frontal bones
evidently do not extend far beyond the area of this structure. In view of the partial
disorganization of the specimen, possibly associated with an attack from other
predaceous or scavenging monsters of the period, it is suggested that this curious
structure represents an inverted cranial element. It cannot, however, be allocated
with any of the bones, the contours of which are so clearly demonstrated by Sollas’
classic sections,* and possibly represents hyobranchial elements. And it is, of course,
possible that further material will demonstrate characters which will warrant the
establishment of a new genus fcr this large Australian Ichthyosaur. The prominent
ridges, which are present in the parietal region and on the nasal bones, appear
to be distinctive features. From the occipital border of the parietal region a convex
median ridge extends anteriorly, and this is accentuated by the presence on each
side of elongate valleys, the lateral sides of which curve upwards to form the
borders of the supratemporal fosse. At the anterior termination of the median
ridge there is a cavity which could not be traced into the internal tables of the
skull, but which probably represents a disrupted pineal foramen. This is nearly
in line with the anterior borders of the supratemporal fossz, and is thus in the
usual position for the foramen.
3 W. J. Sollas, Phil. Trans. Roy. Soc., B, 208, 1916, pp. 66-126.
248 MEMOIRS OF THE QUEENSLAND MUSEUM.
In the temporal region the anterior horn of the squamosal extends to the
middle of the large oval fosse, articulating with the postorbital. The fosse are
fairly symmetrical, approximately 120 mm. in length, with a breadth of 75 mm.
On the right-hand side the orbit is well preserved, except for its posterior
border. The cavity has been largely set free from the cement-like matrix, which here
contained molluscan fragments. It was carefully excavated in the hope that the
characteristic sclerotic plates, possibly driven inwards, might be exposed, but
these have entirely disappeared. It is evident that the orbit was the characteristic
oval. Its vertical diameter at the periphery is 110 mm. Beneath the orbit, portions
of the jugal can be seen, but the full extent of the zygomatic arch with its sutures
cannot be outlined. The jugal appears first as a raised process near the midline of
the anterior border of the orbit and then curves down to form its lower edge.
_ A large supratemporal bone is present on the left-hand side, and its superior
margin junctions with the lateral border of the squamoso-postorbital arcade. Much
controversy has taken place over the “‘ additional temporal bone,” as 8. W. Williston:
called it in the Ichthyosaurs, and the author has followed the nomenclature of
Lydekker,4 Sollas (loc. cit.), Gilmore, and Andrews in calling it supratemporal.
Williston considered the inner bone of this ‘‘ Diapsid”’ group the tabular and the
outer the squamosal,> but in view of Watson’s demonstrations® it is surely better
to reserve the name ‘“ squamosal” for the more constant element. Perhaps the latero- —
temporal or “ sclerodermal plate,” as Owen called it, is really a separated division
of the quadratojugal.
The quadratojugal is present, and its posterior portion is visible in the
occipital region, where it forms the inferolateral border of the vacuity presented by
the curved shaft of the quadrate. Its sutures with the supratemporal are obscure.
Bast-OccrprraL.—The stout symmetrically convex condyle extends back-
wards beyond the pterygoids for a distance of 33 mm.; the tranverse diameter is.
74 mm., vertical diameter 64.
Basis CraNit.—The suture between the basioccipital and the basisphenoid
can be traced, giving a length of 60 mm. to the former bone. The basisphenoid
is about 75 mm. in length, and forms with the posterior element a rectangular
rostrum for the support of the pterygoids. The basisphenoid has a visible width
of about 60, whilst the basioccipital is about 40 mm. Near the posterior margin
of the basisphenoid the opening of the single canal for the carotic arteries can be
clearly seen; this foramen is circular and has a diameter of 10 mm. In the median
line of the interpterygoid vacuities the splint-like parasphenoid may be seen, but this
has been only partially freed from the matrix, compared with which it is very friable.
This bone evidently increases in thickness towards its upper surface and is trian-
gular in section. It can be traced anteriorly for a distance of 220 mm., where it is:
4R. Lydekker, Catal. Foss. Rept. Brit. Mus., Part 11, 1889, p. 3.
°>§. W. Williston, Phylogeny and Classification of Reptiles (Journ. Geo XXYV), 1917,
p. 416.
6D. M. S. Watson, Ann. Mag. Nat. Hist. (8) XIV, 1914, pp. 84-95.
249
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250 MEMOIRS OF THE QUEENSLAND MUSEUM.
lost in matrix. The pterygoids provide the greater part of the base of the posterior
moiety of the skull. On the left-hand side the pterygoid is in juxtaposition with
and somewhat overlaps, owing to displacement, the basioccipital and basisphencid.
The right pterygoid has been tilted below the plane of the flanges presented by the
axial bones. From th2 basioccipital to the lateral border the pterygoid attains a
maximum breadth of 110 mm.
Owing to the presence of superimposed hyoid rods, and a brecciated mass
of broken teeth and matrix containing associated fossils, the central portion of the
lower surface of the fossil is obscured. The extent of the interpterygoid vacuities
cannot be gauged, but, judging from the converging inner margins of the bones,
the contours in this area are similar to the skull of Ichthyosaurus longifrons as
figured by Owen.? The palatine elements appear to be displaced and are not visible
in the same plane.
Posterior REGIoN.—The contours of the superior border of the occipital
region, formed by the parietals and processes from the squamosals, are quite
continuous, being convex in the median area and then sioping to lateral concavities.
Viewed from above, the postero-external borders of the squamosal are seen to
curve symsnetrically backwards, and, although the occiput is somewhat disrupted,
the contours are quite elegant. The inner process of the squamosal unites in an
oblique suture with the lateral arm of the parietal near the median line of the
supratemporal fossa.
A large quadrate is present on each side, but owing to mandibular pressure
these bones have been forced somewhat out of position.
Above the region of the foramen magnum, only small fragments are visible
in the matrix of elements which correspond to the superior occipitalia, which have
apparently been forced inwards. Possibly these are paired extensions of the
supraoccipital which form part of the lateral borders of the foramen magnum.
The opisthotics are in place on each side, and junction with the basioccipital,
the ~ stapes,” and the squamosal.
SUGGESTIONS FOR A RE-INTERPRETATION OF THE SO-CALLED STAPES—
Next to the quadrate, the largest bone in the occipital region is the element
called ‘‘ stapes’’ by Sollas and Andrews. This acts as a strong lateral buttress of
the basioccipital, and lies above the posterior flange of the pterygoid. It has an
expanded facet for junction with the basioccipital, with an adjoining superior
surface for association with the opisthotic. Cope,’ who was the first to name this
bone, did so with diffidence, and figured it as distinctly separated from the basi-
occipital, whereas modern authors rightly show it as a buttress supporting the
rostrum of the condyle. Owen? named it the paroccipital, but apparently only
® Cope, Proc. Amer. Assn. Ad. Sc., 1871, p. 199, fig. 2.
® Owen, Mon. loc. c't., p. 94.
AN ICHTHYOSAURIAN SKULL FROM QUEENSLAND.—LONGMAN. 251
‘dealt with two pairs of occipital elements below the supraoccipital. C.W. Andrews
notes that this bone (stapes) “‘ seems to have lost its auditory function,”!° and it
is obvious that this so-called stapes cannot be associated with the fenestra ovalis,
as Cope supposed. The stapes is ususlly regarded as the homologue of the hyomandi-
bulare of fishes, and a large stapes is recorded for the Cotylosauria. Case figures the
stapes of Dimetrodon,! a bone which in this and allied Permian reptiles is regarded
by Broom!? as the tympanic. The writer is unable to find, however, a parallei in
literature to the interpretation of this buttress bone of the Ichthyosaurs as a stapes.
The columella auris of modern reptiles, the proximal end of which is
presumably homologous with the stapes, is always placed antero-laterally to the
basioccipital, and is quite distinct in position from this buttress bone.
The writer suggests that these lower lateral elements in the occipital region,
the so-called stapes, should be interpreted as inferior divisions of the exoccipitals.
That the upper elements are true exoccipitals seems to be demonstrated by the
position of the foramen for the post-auditory nerves, as clearly shown in Andrews’s
illustrations (Joc. cit.), and also by their relations to the foramen magnum. The
unusual extension of the intermediate lateral occipitalia, the opisthotics, to the
basioccipital, to which they also act as buttress bones, has probably brought ge
a division of the exoccipitals into upper and lower portions.
These lower lateral elements may thus be interpretated as inferior divisions
of the exoccipitals. This change in nomenclature, giving the exoccipitals a ventral
extension, appears to be generally supported by the position of the occipital elements
in the Permian Tetrapoda studied by von Huene'? and by R. Broom", and. in the
Stegocephalia illustrated by C. Wiman.? It is in consonance with the general
arrangement of the bones in modern reptiles, where the exoccipitals are usually
the lower lateral elements in juxtaposition with the basioccipital, the opisthotics
uniting with them antero-superiorly in adult life (distinct in Chelonians) ; these
relationships of the two elements are shown by Parker's studies of the development |
of the skull in the snake and the lizard.'6 It is not at variance with Howes and
Swinnerton’s interpretation of the development of the skull of Sphenodon.’ It agrees
also’with the positions given by Kingsley in his diagram of the schematic vertebrate
skull.!8 Huxley wrote that but for its large size he would have regarded the
adjoining bone; now generally aocepved as the opisthotic, as es stapes.!?
POW SEW - Restless. Marine Rept. Oxford Clay, Brit. Mus., 1910, p. 11.
11. C. Case, Bull. Amer. Mus. Nat. Hist., XXVIII, 1910, p. 190. -
12, R. Broom, Bull. Amer. Mus. Nat. Hist., XXVIII, 1910, p. 223.
13 yon Huene, Bull. Amer. Mus. Nat. Hist., XXXII, 1913, pp. 315-386.
14R. Broom, Bull. Amer. Mus. Nat. -Hist. XX XII, 1913, p. 563, etc.
15 ©, Wiman, Bull. Geol. Inst. Upsala, XIII, 1915, Pt. 1.
16 W. K. Parker, Phil. Trans. Roy. Soc., Vols. 169 and 170, 1878-79.
17 Howes and Swinnerton, Trans. Zool. Soc., XVI, 1901.
18 Kingsley, Outlines Comp. Anat. Vert., 2nd edit., p. 74.
19 Huxley, Anatomy of Vertebrated Animals, 1871, p. 211.
252 MEMOIRS OF THE QUEENSLAND MUSEUM.
The opisthotic or paroccipital very rarely appears to meet the _ basi-
occipital below the exoccipitals in other reptiles, although the relations between
these elements are variable. The writer has diffidence in using terms that are
not accepted by leading authorities, but the occipital region of the Ichthyosaurs
provides material for several interpretations, and the use of exoccipital for the
lower element seems to solve the difficulty of a most anomalous “ stapes.”
In the modern cetaceans the stapes is frequently reduced to a small conical
plug, and, judging from analogy, the auditory functions of the Ichthyosaurs would
not have been greatly utilised. Possibly the real stapes is the “long slender
process ’’ demonstrated in Section 494 of the very fine series in Sollas’ great work
(loc. cit.).
The stapes is often missing in fossils. The elaborate studies by D. M. S..
Watson of the position of the fenestra ovalis in Therapsids, Seymouria, etc.
(P.Z.S., 1914, and 1919) have an important bearing here, but the posterior
aspect of the occipitalia in our specimen presents no evidence on this point.
The massive architecture of the occipital region was evidently associated
with the attachment of powerful nuchal muscles. Perhaps a specialist will one
day. work out details of the probable musculature of the Ichthyosaurs on similar
lines to the recent studies by Gregory and Camp on Cynognathus.*°
Lower Jaw.—On the left hand side the lower jaw is practically complete,.
except for the missing anterior segment and a small portion of the angulare. The
dentary is very elongated and is no less than 875 mm. in maximum length. Parallel
with the alveolar border, and situated about 20 mm. below it, is a groove which
is shallow anteriorly, but then deepens, giving the characteristic conjoined gun-
barrel effect of the Ichthyosaurian rostrum. The posterior process of the dentary,.
which overlaps the surangulare, runs back to below the mid-region of the supra-
’ temporal fossa. Here the semi-spherical contours (in section) of the upper rod, or:
gun-barrel-like process, sink into the same plane as the surangulare and angulare.
The left angulare is not quite complete at its posterior end, and here its:
outer contours have been disturbed. It is a longer and more massive bone than the
surangulare, but just at the termination of the dentary the two posterior elements
are of equal depth. In this region the arrangement of the bones is very similar to.
the outer view given by C. W. Andrews for Ophthalmosaurus in Text-fig. 20 (loc. cit.),
except that the angulare is distinctly extended to form the posterior portion of the
mandible. Strong depressor muscles were evidently attached here, working with
short leverage in association with the powerful levators placed in front of the
articulation. Sollas points out that the levator muscles originating in the temporal
region, inserted on the lower jaw, acting as levers of the third order, were “‘admirably
adapted for snapping ; and the Ichthyosaurus, from all that we know of it, must have
obtained its food by seizing fish ‘ upon the wing.’ ”
*0 Gregory and Camp, Bull. Amer. Mus. Nat. Hist. XX XVIII, 1918, pp. 447-563.
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. V1I, Puate XVI.
Fig. 1—Skull of ICHTHYOSAURUS AUSTRALIS. Posterior view. Approximately one-third
natural size.
Fig. 2.—Anterior section of restrum, ICHTHYOSAURUS AUSTRALIS, showing disrupted teeth.
Approximately one-half natural size.
AN ICHTHYOSAURIAN SKULL FROM QUEENSLAND.—LONGMAN, 2
vt
The articular is present and evidently rests in its original position on the
angular, with its outer lateral surface adjoining the surangulare. Only the post-
articular area has been freed from the matrix, and this is obliquely set on the
angular, with its upper surface turned towards the supraoccipital region,
The splenials first appear on the lower surface of the mandible at a distance
of one-fifth of the length of the skull, this being greatly in advance of the divergence
of the premaxillze to expose the nasals in the upper jaw. They are visible for a
considerable distance on the inferolateral margin. The relative proportions of
these elements are shown in Text-fig. 1. The condition of the fossil does not present
evidence for the description of the inner side of the mandible.
Text-figure 2.—Section showing premaxillary and dentary elements, with teeth fragments,
in the polished surface of anterior segment of rostrum, Ichthyosaurus australis (natural size).
The internal structure of the anterior end of the rostrum has been demons-
tratedjby polishing, and is shown in Text-fig. 2. The alveolar groove has a well-
developed inner wall with an expanded margin. On each side of the lower jaw the
section exposes a foramen leading from the nutrient groove to the vascular canal.
This section is entirely from premaxillary and dentary elements.
TrETH.— Unfortunately there are no complete teeth in situ but, as previously
ated, remains have been preserved adhering to external surfaces. Some of these
254 MEMOIRS OF THE QUEENSLAND MUSEUM.
which were cemented to the lateral surface of the rostrum are shown in Plate XVI..
Between the mandibular rami about the mid-region of the skull there is a brecciated
mass formed mainly of broken teeth, although containing other fossils. The teeth
are subeylindrical, and there are no anterior or posterior carine. They are about
55 mm. in length, with a maximum diameter of 18 mm. After counting the number
of grooves on exposed contours, it is considered that a perfect specimen would have
about thirty longitudinal strie. The teeth are evenly conical, and there is no-
marked tumidity at the alveolar border. Judging from the numerous fragments,
they were fairly isodont, but some of the roots are decidedly compressed. They
are somewhat larger than those in the type specimen in this Museum of
Ichthyosaurus marathonensis, Etheridge fil. It is estimated that about forty teeth
were present in each alveolar groove.
On the posterior third of the lower jaw, and cemented thereto, may be seen
remains of the stout, cylindrical “ hyoid rods ”’ (to use Sollas’ term). These are 15
mm. in diameter, and attain a length of 250 mm. They also overlap the lateral
surface on the left-hand side, as may be seen in Plate XV.
Unfortunately no other remains were found in association with the skull, and
although this Museum contains several specimens of vertebre, a paddle, and a few
other fragments from Western Queensland Cretaceous formations, it would be
unwise to describe these in conjunction with the present material.
CLASSIFICATION.—The Ichthyosauria have been divided into four families :—
Mixosauride, Ichthyosauride, Baptanodontide, and Shastasauride, the last-named
being added by Merriam in 1902.21. The diagnostic characters of the majority of
recent genera are not founded on cranial characters. It is evident, however, that
our fossil has no affinity, generically, with the Ophthalmosaurus-Baptanodon group
(Jurassic), which is characterised by an enormous eye and a very reduced dentition.
Neither is it related to Mizxosaurus (Triassic), in which the teeth are not set in a
continuous groove. In the occipital and mandibular regions and in the basis cranii
the elements in the Queensland skull approximate to the Ichthyosaurus type, but
apparently unique characteristics are present on the superior surface of the skull.
Two species of Ichthyosaurus have been described from fragments obtained
in the Cretaceous deposits of Western Queensland. In 1867, McCoy briefly described
Ichthyosaurus australis and in 1869 supplied some additional details. His material
consisted of two portions of the skull, teeth, some dorsal vertebre, paddles, and
a humerus, radius, and ulna, and his short description was not illustrated. In 1888,
Etheridge fil. described Ichthyosaurus marathonensis™ from a maxillary and dentary
section ten inches in length, which is now in the Queensland Museum. In 1892
21 Merriam, Bull. Dep. Geol., Univ. Calif., Vol. IIT, 1902, p. 87.
22 McCoy, Ann. Mag. Nat. Hist., XIX, 1867, p. 355; XX, p. 196..
28 McCoy, Trans. & Proc. Roy. Soc. Vic. IX, 1869, pp. 77-78.
*4 Etheridge, Pr. Linn. Soc. N.S.W., III, 1888, p. 405, Pl. VIL.
Ll
AN ICHTHYOSAURIAN SKULL FROM QUEENSLAND.—LONGMAN. 258
)
Etheridge gave details of these species with actual localities and records of other
fragments, and queried the identity of the two species.2?> In 1897 he made a further
note on precaudal vertebra as being Ichthyosaurus australis.26 Direct comparison
between the type fragment of I. marathonensis and the present skull shows too
great a similarity, so far as this region is concerned, to warrant the introduction of
a new specific name; indeed the type fragment, except for the slightly smaller
teeth, could almost be placed as a section of the rostrum of our fairly complete
skuJl without doing violence to its contours.
Unfortunately, McCoy’s material was too inadequately described to permit
Etheridge to make a satisfactory comparison. The teeth were compared with those
of I. campylodon, and the antero-posterior diameter of the orbit was given as five
and a-half inches. Chapman figured a cranial fragment and a paddle of the type
material in his useful handbook.?? McCoy’s species has priority, and in the present
state of our knowledge it is deemed inadvisable to ignore it in connection with the
Queensland. skull, with which its sectional dimensions fairly agree. Our specimen
has therefore been catalogued as Leh'hyosaurus australis McCoy.
I. marathonensis is regarded as a synonym. ‘There is evidence, however,
of another species occurring in Queensland, distinguished by smaller teeth, but
this awaits further material before elucidation.
The relationships of the Ichthyosaurs have ‘been the subject of recent
research by such prominent authorities as Sollas, Andrews, Williston, Merriam,
van Huene, and Broom, and partly centre around the homologies of the temporal
elements. It is now generally agreed that the Ichthyosaurs are descended from the
primitive Permian Carboniferous Cotylosaurs. This primitive reptilian group exhibits
ordinal characters which, in the words of W. K. Gregory, are “ structurally ancestral
to all the higher vertebrates.”*8 This fascinating problem, with its wealth of litera-
ture illustrating very numerous forms, is too intricate to be dealt with here. From
the standpoint of popular interest it may be mentioned, however, that an outstanding
characteristic of most Ichthyosaurs, an elongated rostrum, is a common feature in
very diverse vertebrates. Among the large present-day reptiles the prolonged
snout of the gavials is remarkable. There are commonplace examples amongst
birds and fishes. The Ziphoid Cetaceans, such as species of Mesoplodon, are
mammalian types which rival the Ichthyosaurs in the extreme development of the
anterior elements. This elongated rostrum is evidently a characteristic associated
with special feeding habits, and is a demonstration of the mobility of vertebrate
structure. Fortunately the specialised forms in these groups are not isolated, in
this respect, but can now be studied in conjunction with allied genera and families
> Jack & Etheridge, Geology and Paleontology of Queensland, 1892, pp. 505-598.
°6 Etheridge, Rec. Austr. Mus., III, 1897, pp. 66-68.
27 Chapman, Australasian Fossils, 1914, fig. 133, p. 277.
*8 W. K. Gregory, ‘‘ Origin and Evolution of Human Dentition,’’ Jr. Dent. Res., II, 1920,
p. 100.
256 MEMOIRS OF THE QUEENSLAND MUSEUM.
which are more conformable to the usual type of skull. The various developments
of the rostrum in vertebrate crania provide abundant material to illustrate
remarkable processes of evolution.
Marrix.—Professor H. C. Richards, Hon. Petrologist and Mineralogist on
our staff, kindly contributes the following note on the matrix :—** Stone-coloured,
fine-grained, homogeneous calcareous mudstone. There is sufficient calcareous
material to cause rapid effervescence with dilute hydrochloric acid, and a residual
mass of cream-coloured clay material is left behind.”
AssocIATED Fossrts.—An almost-complete shark’s tooth is present in the
brecciated mass between the mandibular rami, and is probably an anterior tooth of
Lamna appendiculata Agassiz.
Several specimens of the common bivalve, Aucella hughendenensis Etheridge
were taken from the matrix, including fairly complete valves from the orbits. Two
specimens of a small gasteropod were found, and Mr. W. H. Bryan, M.Sc., Hon.
Palzontologist on our staff, has identified these as T'wrritelia (?) microlinea, a rare
shell which was named by Etheridge in 1920.9
LIST OF PLATES.
Plate XV.
Figure 1.—Skull of Ichthyosaurus australis; superior view. Approximeteh—one-third
rratureal—size.
Figure 2.—Skull of Ichthyosaurus australis; lateral view. Approximately one-seventh
natural size.
Plate XVI.
Figure 1.—Skull of Ichthyosaurus australis; posterior view.
Figure 2.—Anterior section of rostrum, showing disrupted teeth. Approximately one-
half natural size.
29 R. Etheridge, Publ. No. 269, Qld. Geol. Sur., 1920, p. 13.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL.,
bo
Ol
~]
AUSTRALIAN BEES IN THE QUEENSLAND
MUSEUM.
By T. D. A. CocKERELL, UNIVERSITY OF COLORADO.
PARASPHECODES Smith.
This genus, which is very closely allied to Halictus, consists of a large number
of species, nearly all apparently with very restricted distribution. They may be
oligotropic on particular flowers, but at present we know hardly anything of their
babits. Smith described a number of species, giving them names which are anagrams
of Halictus. I have examined the types of all of these, but to my surprise I fail to
find them in the various Australian collections coming to hand in recent years. The
genus is especially south-eastern, with no less than twenty-one species described
from Tasmania, and nineteen from Victoria, one of the latter extending to Tasmania.
Two of the Victoria species go north to Queensland, which has eight species and
a subspecies of a Victoria one not known elsewhere. From South Australia five
species have been described, but from West Australia only two, and N.W. Australia
one. Two are only known from N.S. Wales. The series now before me from the
Queensland Museum (to which the types will be returned) adds a number of species,
separable from one another as follows :—
Clypeus partly yellow; males .. % st ts a fe ee ne ey Il
Clypeus all black; females a =e “ .- s :- -: -- -. 4.
1. Abdomen entirely black; mesothorax glaucous... 3 ar .. cervicalis Cockerell.
Abdomen partly or wholly red Fe 5 ae se Sr a ee Ae 2
2. Abdomen wholly chestnut-red ; tubercles entirely black .. ts ee ao 5, SOUSID ASE
Abdomen partly black .. +e 5 Sc af se a rie ee it 3.
3. Tegule lght ferruginous sf Ate ok ~ =i xs ae longmani n. sp.
Tegule black a ae an ae “A ea ane cs .. subrussatus n. sp.
4, Abdomen entirely black bes Ay ee Se ee a xa ae = 5.
Abdomen at least partly red .. Deis Pf ae ate ee ae Sue a 6.
5. Area of metathorax short, distinctly plicate, with a prominent raised rim .. cervicalis Cockerell.
Area of metathorax long, feebly sculptured, subemarginate at apex, without a raised
Tahan)” Ee a: se zi parameloenus . sp.
6. Flagellum red beneath .. Se se = as Sa as ae lichatinus n. sp.
Flagellum wholly black .. 5 a ay AA tic te et zo os ile
7. Larger; fourth abdominal segment entirely black ae ifs .. hirtiventris n. sp.
Smaller; fourth abdominal segment red e as te oe Ps annexus Nn. sp.
R
258 MEMOIES OF THE QUEENSLAND MUSEUM.
Parasphecodes cervicalis- Cockerell.
Described from the ©, collected in Tasmania. In December, 1919, Mr. Hacker
took both sexes in the National Park, Queensland, where he also found the Tasmanian
Callomelitta littleri. The specimens of 9 P. cervicalis seem to d'ffer slightly from
the Tasmanian form, and when Tasmanian males are available may possibly be
separable, but at present I am quite unable to state any valid distinctions. The
occurrence of supposed Tasmanian endemics at various points on the mainland
begins to carry the conviction that the Tasmanian bee-fauna does not represent:
a high degree of differentiation since the separation of the island, but rather the
survival of a series of old species, some of which are now extinct on the mainland,
while others are restricted to limited areas.
; The male of P. cervicalis is new. It resembles the female, but is more slender,
while the clypeus is chrome-yellow except for a large quadrate black mark on each
side above. Labrum, mandibles, and antenne black, the flagellum not strongly
moniliform. There is black hair at the apex of the abdomen; the second ventral
segment has a median elevation, without any tuft of hair. The abdomen is entirely
black, the margins of the segments concolorous. There is a close general resemblance
to P. nigritus Meyer, from Tasmania.
Parasphecodes solis n. sp.
Male. Length nearly 10 mm., slender, head and thorax black, abdomen
entirely chestnut-red, suffused with dusky beyond the third segment, but with no
black segments or dark markings; hair of head and thorax dull white, abundant
on front and sides of face ; some dark hair at end of abdomen ; labrum and mandibles
black or nearly ; clypeus with an extremely broad pale yellow band, emitting a
cuneiform extension upward; scape short, flagellum very long, strongly monili-
form, the joints of the basal half marked with red beneath ; mesothorax entirely
dull, with a granular appearance ; scutellum bigibbous, dull, and granular, without
special tomentum; area of metathorax distinctly defined, angulate posteriorly, covered
with very coarse wavy rug, between which it is distinctly shining; tegule dull red ;
wings dusky reddish, stigma dark reddish, nervures fuscous ; second submarginal
cell higher than long, receiving recurrent nervure very near its end ; femora black,
red at extreme apex; tibiee red, the anterior pair clear and bright, the hind ones
dusky ; anterior tarsi dusky red, the others black or nearly ; abdomen moderately
shining, with excessively fine punctures, close and uniform on first two segments, —
on first segment extending to the margin; no prominence on under side of second
segment.
Sunnybank, 19-11-13 (H. Hacker). In Meyer’s table and in my table of
Tasmanian species, this runs to P. perustus Ckll., but is larger, with darker antenne, |
&c. It resembles perustus in having the third submarginal cell much broader above
than second.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL, 259
Parasphecodes longmani n. sp.
Male. Length about or nearly 8 mm, rather slender, head and thorax black.
abdomen dusky-red, with a broad. black stain on middle of first two segments, and
segments beyond the third wholly black; hair of head and thorax thin, brownish
on face, front, vertex, and thoracic dorsum, pure white on cheeks and mesopleura ;
head broad, orbits strongly converging below; broad band on clypeus, with a.
cuneiform upward extension, labrum and greater part of mandibles yellow ; antennz
black, the flagellum moniliform, not especially long (length somewhat over 4 mm.) ;
front entirely dull, but a shining space in front of middle ocellus ; mesothorax and
scutellum dull, slightly glistening on disc, very minutely and densely punctured,
scutellum bigibbous, without special tomentum ; area of metathorax poorly defined,
with fine vermiform ruge; tegule bright yellowish-ferruginous ; wings hyaline,
slightly greyish ; stigma dark reddish, nervures fuscous ; second submarginal cell
nearly square; first recurrent nervure entering base of third submarginal, as in
P. sulthica Smith ; femora black, with red knees ; anterior and middle tibiz and all
the tarsi red; hind tibize black with broadly red base and a little red at apex ;
abdomen above with very scanty short hair, venter with much white hair, second
ventral segment simple.
Caloundra, 20-1-16 (H. Hacker). Rather like P. minimus Meyer, but larger,
with dark antennze, &c.. The tubercles are red. Dedicated to the Director of the
Queensland Museum.
Parasphecodes subrussatus n. sp.
Male. Length about or a little over 8 mm., not very slender, the abdomen
broad at base, almost parallel-sided but slightly widening to the fourth segment ;
head and thorax black, with rather long dull white hair, faintly brownish dorsally ;
first abdominal segment black, broadly red apically, with a further red extension
on sides; second segment red, with a transverse median black mark ; third red,
with a black spot at each extreme side toward base ; fourth and following segments
black, but the fourth red beneath, with a black median stain; at least lower half
of clypeus shining lemon yellow, with a small median extension; labrum black ;
mandibles black, dark-reddish apically ; flagellum long, clear ferruginous beneath ;
mesothorax and scutellum dullish, somewhat shining on disc, extremely finely
punctured, scutellum bigibbous; area of metathorax poorly defined, with fine ruge ;
tegule black ; tubercles black ; wings hyaline, slightly greyish, stigma and nervures
ferruginous ; second submarginal cell much higher than long, receiving recurrent
nervure well before its end; legs black; abdomen shining, the red colour bright ;.
second ventral segment simple.
Kosciusko, 26-1-14 (A. J. Turner). Very distinct by the marking of the
abdomen, combined with the black tegule and legs.
260 MEMOIRS OF THE QUEENSLAND MUSEUM.
Parasphecodes paramelzenus n. sp.
Female. Looks like P. cervicalis, with which I had mixed it, but it is a little
smaller, with anterior wing about 7 mm. ; face narrower, clypeus without a distinct
median groove ; mesothorax entirely dull black, not at all glaucous, with excessively
dense and minute punctures; scutellum densely and minutely punctured ; area of
metathorax long, with no raised rim, its surface obscurely rugose, except apically ;
wings paler and strongly reddish; first recurrent meeting second intercubitus ;
outer side of third submarginal cell without a double curve ; first two abdominal
segments very finely, closely, and regularly punctured, the depressed apical part
of second segment, and first to extreme apex, punctured ; second ventral segment
hairy, but without a distinct prominence. The apical part of abdomen has short
black hair.
National Park, Queensland, Dec., 1919 (H. Hacker). P. atronitens Ckll. is very
closely allied, but distinguished by the colour of wings and sculpture of abdomen.
Paresphecodes lichatinus n. sp.
Female. Length about or nearly 10 mm., anterior wing nearly 8; robust,
head and thorax black ; abdomen with the first three segments deep chestnut-red,
the first with a very broad blackened area across the middle, the third with the hind
margin blackened ; fourth and following segments black ; hair of head and thorax
dull whitish, thin on thorax above ; clypeus shining, with sparse strong punctures ;
flagellum red beneath and at extreme tip, the colour almost brick red ; mesothorax
and scutellum densely and distinctly punctured, but glistening between the punctures ;
scutellum slightly bituberculate ; area of metathorax with coarse irregular ruge,
more or less broken up, and a sharp hind margin ; tegule dark red ; wings reddish,
strongly dusky in marginal cell, stigma and nervures piceous, stigma unusually
narrow ; second submarginal cell large, about square, about as broad above as third,
receiving first recurrent nervure almost at apex; legs black, with coarse dark hair
on outer side of hind tibia and tarsi; abdomen shining, finely punctured, apical
part with black hair dorsally.
Ararat, Victoria (H. W. Davey). Very close to P. lichatus Smith, from Western
Australia, but with darker wings, and I think separable.
Parasphecodes hirtiventris n. sp.
Female. Length about 10 mm., robust, head and thorax black, their pubescence
fulvous dorsally ; abdomen with the first three segments dark red, the others black ;
first segment with a large T-shaped black mark, third with a transverse dusky cloud
in middle, and hind margin darkened ; an even narrow line of white cilia overlapping
extreme base of third segment; tongue cuneiform, short and broad; clypeus
shining, with scattered punctures ; antenne entirely black, scape very long ; meso-
thorax and scutellum dull, extremely densely and minutely punctured ; area of
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL, 261
metathorax long, shining, the basal half obscurely sculptured; extreme sides of
metathorax densely hairy; mesopleura very hairy; tegule rufopiceous; wings
dusky, darker at apex, stigma and nervures rather dilute sepia ; outer recurrent and
intercubitus weakened ; second submarginal cell broad below, narrowed above, the
recurrent nervure joining its extreme apex ; legs black, hind legs with pale hair, that
on tibize shining silvery, hind basitarsi with a bright red tuft at apex; abdomen
shining, first two segments with extremely minute punctures; venter extremely
hairy.
Ebor, 2-1-14 (A. J. Turner). In Meyer’s table runs to P. insignis Meyer, from
Tasmania, but the sculpture is very different. It is also related to P. arciferus Ckll.
and P. leptospermi Ckll. P. arciferus differs at once by the area of metathorax and
colour of pubescence, and the same may be said of leptospermi.
Parasphecodes annexus n. sp.
Female. Length a little over 7 mm., robust, head and thorax black, abdomen
broad, chestnut-red, first segment with a very broadly triangular black area, and
hind margins of all the segments strongly dusky ; hair of head and thorax thin, slightly
ochreous dorsally, white on cheeks and pleura; clypeus shining, with sparse
punctures, and no median groove; antenne black; mesothorax and scutellum
shining, with small punctures, well separated on disc; area of metathorax semi-
lunar with no raised rim, shining, finely rugose, subplicate basally ; tegule dark-
reddish, anterior margin hyaline ; wings hyaline ; stigma and nervures dark-brown,
outer recurrent and intercubitus extremely weak; second submarginal cell about
square, about as broad above as third ; first recurrent nervure joining base of third
submarginal, as in P. sulthica Sm., legs black, hair on hind legs pale, on hind tibiz
silvery, with a green stripe posteriorly ; hind basitarsi with a bright red apical
brush ; abdomen without conspicuous punctures, first two segments excessively
minutely punctured ; venter hairy.
Adaminaby, N.S.W., 19-10-18 (4. J. Turner). In many ways similar to:
P. melbournensis Ckll., but much smaller.
HALICTUS Latr.
The following table separates a series of species in which the mesothorax
is metallic, green or blue. Up to the present, thirty-two species of this type have
been described from Australia (including Tasmania). Some additions are given
below :—
Males ss 6 5 Ea in oe St he a i Ie I
Females a Ke oe Ess es ae +2 Ss a iy: ae 3.
1. Clypeus largely yellow, mandibles yellow .. SE me a5 .. purnongensis Ckll.
Clypeus without yellow .. tee fe ae ee oe a be bet 3 2.
2. Hind tibie light red
, a 45 be ru Se Ne olivinus n. sp.
Hind tibie black, red at extreme ends. (Caloundra, 20-1-16) .. ne kesteveni Ckll-
262 MEMOIRS OF THE QUEENSLAND MUSEUM.
3. Abdomen metallic % a3 i . a ha Ar e oe sit 4.
Abdomen not metallic .. 42 ¥ Bs Fs ee i a bye on 5.
4, Abdomen bright steel-blue. (Brisbane, 12-2-18) .. ae Bi .. caloundrensis Ckll.
Abdomen greenish. (Brisbane, 12-2-18) ae ae ¥: oe 4 dampieri Ckll.
5. Abdomen red, head black ot ots ee oy ae th a os ot 6.
Abdomen black or dark-brownish .. oh ne ate aa ae re a he
6. Smaller; mesothorax shining = zit “4 sé 3 se erythrurus Ckll.
Larger; mesothorax dullish, with larger scattered punctures. (Oxley, Brisbane,
Sept., 1914; Brisbane, 4-9-14, 6-10-14, 26-9-16) .. brisbanensis Ckll.
7. Head metallic .. oy: a cid a $c a pe es Ea ae 8.
Head black b a x5 ae aes ae Bs we ar ee! a 9.
8. Stigma very dark. (Brisbane, 12-2-18, National Park, Q., Dec., 1919) .. urbanus Smith.
Stigma reddish ; scutellum shining, steel-blue, contrasting with the green mesothorax
urbanus var. lomatic n. var.
9. Mesothorax green, dullish St $d <2 we Se Fe .. limatiformis n. sp.
Mesothorax blue. . ats oe By Me Ac oe 3f Bic 5 a6 10.
10. Mesothorax steel-blue, highly polished; stigma dark .. La .. humiliformis n. sp.
Mesothorax not polished; stigma pallid .. ae oe ve 1. mesocyaneus n. sp.
The Queensland specimens were all collected by Mr. Hacker.
Halictus purnongensis Cockerell.
The range is greatly extended by a male from Brisbane, 6-10-14 (Hacker).
It represents a variation with the hind margins of the abdominal segments dark.
Halictus erythrurus Cockerell.
Brisbane, 3-10-16, a variation with tegule darker than usual.
Halictus olivinus n. sp.
Male. Length about or nearly 5 mm., slender, head and thorax shining dark
olive green, the metathorax steel-blue, contrasting ; labrum and mandibles dark-
reddish ; head unusually broad, eyes strongly converging below, clypeus polished
and shining, front dull, shining along orbits ; scape black ; flagellum long and slender,
ferruginous beneath; mesothorax shining but not polished, scutellum highly
polished ; area of metathorax large, with weak strie, apical margin thickened ;
posterior truncation without sharp lateral edges ; hair of head and thorax extremely
scanty, white; tegule pale tes‘aceous; wings hyaline, faintly dusky; stigma
piceous ; marginal nervure dark, the other nervures mainly pallid, the outer recurrent
and intercubitus hardly visible ; first submarginal cell much larger than the other
two combined ; second submarginal very narrow, third almost as narrow as second ;
first recurrent nervure meeting second intercubitus; femora rufopiceous, knees,
tibize, and tarsi pale ferruginous ; abdomen dark brown, brightly polished, venter
with erect hair.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL. 263
Brisbane, 3-3-14 (H. Hacker). I am at a loss to associate this with any known
female. Among the described males it has some resemblance to H. dampieri Ckll.,
but in that species the area of metathorax is dull and quite different ; in the present
insect it is concave and shining. It is easily known from H. hackeriellus Ckll. by the
red tibie.
Halictus urbanus var. lomatiz n. var.
Female. Mesothorax peacock-green ; scutellum bright blue, highly polished,
contrasting ; legs rufotestaceous; stigma rather dilute reddish-brown ; flagellum
bright ferruginous beneath. Outer recurrent nervure and intercubitus hardly visible.
Sunnybank, Brisbane, on flowers of Lomatia, 13-5-12. (H. Hacker.) I had
taken this for a distinct species, on account of the shining blue scutellum and paler
stigma, but microscopic examination shows that the hind spur, sculpture of area of
metathorax and scutellum, &¢., are precisely as in H. urbanus.
Halictus limatiformis n. sp.
Female. Length about 5 mm., black, with the mesothorax only (not even
scutellum) dark bluish-green ; pubescence dull white, abundant and curled on under
side of abdomen; apical part of mandibles very dark-reddish ; front dull, orbital
margins shining; antenne black; mesothorax shining but not highly polished,
with scattered punctures; scutellum shining; area of metathorax granular and
dull, feebly plicate, with a shining margin; tegule dark castaneous ; wings dusky
hyaline ; stigma large and very dark, nervures brown, outer recurrent and inter-
cubitus almost obsolete ; first recurrent nervure ending just before second inter-
cubitus ; third submarginal cell short, little longer than second ; legs black, anterior
knees red; abdomen shining black, without bands. Under the microscope the
mesothorax is seen to be minutely tessellated, with widely scattered distinct
punctures. Area of metathorax entirely without a sharp rim. Hind spur pectinate
with small teeth.
National Park, Queensland, Dec., 1919 (H. Hacker). Very close to the
Tasmanian H. limatus Smith, but that has the area of metathorax more distinctly
‘striate, and pale testaceous tegule. It may be only a race of limatus.
Halictus humiliformis n. sp.
Female. Closely resembling H. limatiformis, but differing thus :—Mesothorax
‘shining blue, with a polished surface; area of metathorax with fine but regular
and distinct plice or striz (as in H. limatus, from which it differs by the shining
mesothorax). By the small shining mesothorax it agrees with H. humilis Smith,
but from that it differs by the black legs, with anterior knees red, and the entirely
black antenne. The hind spur is not pectinate; the scutellum is very sparsely
punctured, and the surface of the mesothorax, between the punctures, shows only
traces of tessellation.
Kbor, N.S.W., 30-12-15 (A. J. Turner).
4
264 M#tMOIRS OF THE QUEENSLAND MUSEUM.
Halictus mesocyaneus n. sp.
Female. Length about 5 mm., head and thorax black except the mesothorax,
which is deep blue, somewhat shining, but not polished ; legs black (anterior knees
not red); abdomen dark brown, suffused with dark-reddish, the hind margins of
the segments paler: venter with long curled hair. Mandibles reddened apically ;
flagellum obscurely brownish beneath; greater part of front somewhat shining ;
scutellum shining; area of metathorax with delicate radiating striz, and a shining
rim ; tegule reddish testaceous ; wings yellowish hyaline, stigma large, pale-reddish ;
abdomen broad, shining. The area of metathorax is microscopically sculptured
between the raised lines or plice. The mesothorax is microscopically tessellate,
with widely scattered extremely weak punctures.
Bribie Island, 1-4-18 (Hacker). Smaller than H. inclinans Smith, and easily
séparated by the polished abdomen and black scutellum.
Halictus bicingulatus Smith.
Mr. Hacker has collected this as follows :—Males, Bribie I., 2-11-13, 2-11-15;
females, Brisbane, 18-9-14. Sunnybank, Brisbane, at flowers of Leptospermum, 9-9-12
Kelvin Grove, 14-9-12, entering a hole in a clay bank.
Nomia moerens Smith.
National Park, Q., Dec., 1919 (Hacker).
Anthophora lilacina Cockerell.
Misprinted “Jilacine’”’ in Mem. Qu. Mus., VII, p. 84, but given correctly im
Hacker’s Catalogue.
GNATHOPROSOPIS Perkins.
Gnathoprosopis amiculiformis (Cockerell).
Prosopis amiculiformis must go in Gnathoprosopis. A female was taken at.
Brisbane, 17-10-16 (Hacker).
Gnathoprosopis bituberculata (Smith).
Males were collected by F. M. Littler in Tasmania ; at Launceston, 7-11-15;
and St. Helens, 14-1-16. The species ‘is new to Tasmania.
Gnathoprosopis amicula (Smith).
Females, National Park, Q., Dec., 1919 (Hacker); a form with rather dusky
wings. The chrome-yellow face-marks and much narrower clypeus readily distinguish.
this from G. hackeri.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM:.—COCKERELL., 265
Gnathoprosopis simpliciventris n. sp. (? hackeri var.).
Male. Agrees with G. hackeri, except that the under side of the abdomen
has a pair of low inconspicuous elevations instead of large dentiform processes.
Brisbane, 6-10-14 (Hacker). This was taken on the same day as a male hacker,
and it seems nearly certain that it is a form of that species, in spite of the great
difference and the prominences of the abdominal venter. A final decision can only
be reached by observations in the field.
Gnathoprosopis amiculina n. sp.
Female (Type). Resembles G. amicula, but is somewhat less robust, with
the anterior and middle tibie entirely black, and the hind ones narrowly cream-
colour at base. The second submarginal cell is shorter, the scape is entirely black
(though the flagellum is ferruginous beneath), and the lateral face-marks are more
pointed above.
Male. Described as male of amicula, Mem. Queensl. Mus., V, p. 198.
The type is from Brisbane, 17-10-16 (Hacker). This is the Brisbane so-called
amicula ; renewed study, with more material, convinces me that it must be separated.
Gnathoprosopis hackeri Cockerell.
Female. Length about 7 mm., robust, considerably larger than amicula or
amiculina ; lemon-yellow lateral face-marks and swollen orange collar and tubercles
as in amicula, but the orange of the prothorax is broadly interrupted in middle, and
the lateral face-marks are distinctly narrower, obliquely truncate above. Scape black,
flagellum ferruginous beneath ; mandibles as usual in the genus; mesothorax and
scutellum finely punctured, but the punctures are distinctly visible under a lens ;
anterior tibie rufous on inner face; hind tibiz with only a small basal cream-
coloured spot; tegule black ; second submarginal cell large and broad, first recurrent
nervure meeting the intercubitus; abdomen with excessively fine punctures, the
basal segment shining.
Brisbane, 8-10-18 (Hacker). I give a description, as no adequate account of
this sex has appeared. .
PROSOPIS Fabricius.
Prosopis cyaneomicans nigrescens Cockerell.
Buibie Island (Hacker).
Prosopis eburniella Cockerell.
Female. Brisbane, 24-9-18 (Hacker).
266 MEMOIRS OF THE QUEENSLAND MUSEUM.
Prosopis coronata Cockerell.
Male. Variety with supraclypeal mark small, tranverse, entirely red.
Brisbane, 6-10-14 (Hacker).
Prosopis kelvini Cockerell.
Brisbane, 8-9-18, two males (Hacker).
Prosopis leptospermi n. sp.
Female. Running in my M.S. key to Gnathoprosopis hackeri, but entirely
different in colour of tubercles and legs, and other characters. It could also be run
near amicula, but is quite different. Length about 7-5 mm., robust, black, with dull
pale-yellow markings, consisting of cuneiform lateral face-marks (ending obtusely
above, away from orbital margin), interrupted band on prothorax above (not
approaching tubercles), and tubercles ; legs black, the anterior tibize with an elongate
red patch in front ; mandibles sulcate, not of the broad Gnathoprosopis type ; clypeus
entirely dull, with a fine median raised line ; scape long black ; flagellum unusually
short, very obscure reddish beneath ; front dull; cheeks beneath with long white
hair; mesothorax and scutellum dull, extremely finely punctured ; postscutellum
large; basal area of metathorax short; tegule black; wings dusky, with black
stigma and nervures; second submarginal cell very broad, receiving recurrent
nervures some distance from base and apex ; more of marginal nervure is on second
submarginal cell than on first; legs ordinary, spurs pale ferruginous; abdomen
dull without evident punctures, first two segments laterally with very thin white
hair; fifth ventral segment with a fringe of short hair, appearing pale reddish in
some lights.
Sunnybank, Brisbane, at Leptospermum flowers, 17-9-14 (Hacker).
Prosopis crassifemorata n. sp.
Male. Length about or hardly 6 mm., black, with the face below antenne,
and the greatly swollen upper border of prothorax (interrupted in middle), connected
with and including tubercles, bright chrome-yellow ; orbits converging below, face
dull, clypeus very long, no supraclypeal mark ; scape rather short and thick, pale at
base and apex; flagellum dull pale red beneath; front dull; mesothorax and
scutellum somewhat shining, very minutely punctured ; metathorax rounded, not
sharply truncate, with only a very narrow basal rugose band ; tegulz partly pallid ;
wings hyaline, very faintly dusky, stigma dark brown; second submarginal cell
broad, receiving recurrent nervures near base and apex ; legs black, anterior tibiz
pale red in front ; hind femora very stout, covered beneath with dense white felt-
like hair ; hind tibize swollen ; abdomen shining, with excessively minute punctures ;
venter simple.
Sunnybank, Brisbane, 19-11-13 (Hacker). A peculiar little species, best
recognised by the hind femora. It runs to 36 in my table.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL. 267
Prosopis xanthopsyche n. sp.
Male. Length about 4:5 mm. ; black, with the face below antennz (the surface
dull), bright chrome-yellow, including a quadrate supraclypeal mark ; lateral face-
marks broadly truncate above, but notched ; mandibles (except tips) and labrum
bright yellow; face rather broad; scape ordinary, with a yellow line in front ;
flagellum long, pale dull reddish beneath ; upper border of prothorax (not reaching
tubercles) and tubercles yellow; mesothorax and scutellum dull, not evidently
punctured (the compound microscope shows minute well separated punctures on
a tessellated surface) ; metathorax rounded, distinctly glistening above, the extreme
base rugose and dull; tegule small and dark; wings dusky hyaline, stigma dark
brown ; recurrent nervures joining first and second submarginal cells near ends ;
second submarginal narrowed above; anterior knees, tibize (except a large mark
behind) and tarsi, middle tibiz at base and apex, and hind tibize broadly at base
all yellow ; middle basitarsi and basal half of hind ones yellowish-white ; abdomen
‘dullish, with a satiny lustre, venter simple.
Brisbane, 24-9-18 (Hacker). The specific name is fanciful, given because
the yellow area of face, viewed upside down, has the outline of a moth with wings
directed backward. In my table this runs to 36, but is unlike any of the species
falling in that vicinity.
Prosopis amatula n. sp.
Male. Length about 3-5 mm., black, slender, with the face below antennz
and narrow upward extensions along orbits, the very short (heart-shaped) scape,
second antennal joint (short and transverse) and the tubercles very pale yellow ;
face rather broad, polished; flagellum long, very pale yellowish-red beneath ;
labrum and mandibles entirely pale yellow; mesothorax and scutellum shining,
but minutely sculptured all over; area of metathorax broad, ill-defined, micro-
scopically cancellate ; wings hyaline, stigma sepia; first recurrent nervure joining
first submarginal cell near end, second meeting outer intercubitus ; legs light yellow ;
hind femora black, yellow at base and apex; hind tibiz with apical half dark ;
abdomen slender, claviform, second segment pale red apically.
Caloundra, Q., 20-1-16 (H. Hacker). Runs to P. primulipicta Ckll. in my table,
but is very distinct. It may also be compared with P. asinalla Ckll., but that is
larger, with longer clypeus and supraclypeal area, &c. P. bacillaria Ckll. differs at
once by the slender scape and dull mesothorax. P. minuscula Ckll. is readily
distinguished by the scape and absence of yellow supraclypeal area. P. pulchripes
Ckll. has a slender scape, larger (deeper) stigma, &e. Under the compound microscope
the second antennal joint of P. amatula appears circular, about as broad as long.
Prosopis brevior Cockerell.
Male. One from Caloundra, 20-1-16, taken with P. amatula. One Oxley,
Brisbane, 17-9-14 (Hacker).
268 ' MEMOIRS OF THE QUEENSLAND MUSEUM.
Prosopis melanocephala n. sp.
Female. Length about 5 mm., head and thorax black, without light markings,
but flagellum bright chestnut-red; legs black, the tarsi more or less reddish ;
abdomen shining chestnut-red, with a black patch on middle of first segment, and
the other segments more or less suffused with dusky medially, the basal part of third
and fourth segments slightly purplish ; tegule black ; wings hyaline, stigma black ;
second submarginal cell broad, receiving recurrent nervures very near base and
apex; clypeus broad; face and front striato-punctate ; a slender groove (visible
under compound microscope) up middle of front; mesothorax dull and closely
punctured ; base of metathorax coarsely rugose.
Type from Darra, 14-10-13 (Hacker). Also from Brisbane, 14-11-12 (Hacker).
A very distinct species, running out at 7 in my table.
Frosopis ancorata var. subconstricta n. var.
Male. Length about 5 mm., face-marks cream-colour; firs) abdominal
segment dark red. Very close to P. constricta Ckll., with the same face-markings,
antenne, &c., but abdomen black with the first segment dark red (bright chestnut-
red at sides of base) and second with apical margin narrowly reddened ; tubercles.
black with a slight red edge ; wings brownish.
On flowers of Malaleuca preissiana, Brisbane (Hacker). P. ancorata appears
distinct from P. constricta Ckll., but the variety now described is intermediate, and
it will probably prove that all three are forms of a single species.
Prosopis scintillans n. sp.
Male. Length hardly 3-5 mm., slender, clypeus, labrum, mandibles, and
narrow bands along orbits (ending very slenderly about halfway up front) all white ;
scape and second antennal joint pale yellowish beneath, the rest of antennz pale
red beneath ; head round seen from in front; mesothorax and scutellum dullish ;
the compound microscope shows vertex, mesothorax, and scutellum all with distinct
well separated punctures on a tessellated surface ; metathorax rounded, the broad
base with microscopic transverse lineolation, more or less joined to form tessellation,
but with no trace of plice or ruge ; tegule dark brown, wings hyaline, faintly dusky,
iridescent ; stigma dark brown; second submarginal cell small, recurrent nervures
meeting the intercubitals ; upper border of prothorax dark, but tubercles apically
yellow ; anterior knees and tibie very pale-reddish, the tibiz with a faint dusky
cloud; middle and hind tibiz white or reddish-white at base and apex; all the
tarsi light; abdomen shining reddish-black, the venter simple. There is a very
small supraclypeal mark.
Brisbane, 26-9-16 (Hacker). Goes to 34 in my table, and appears to be close:
to P. scintilla Ckll., but with white face-markings. It may possibly be a subspecies.
of Scintilla.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL. 269
The above species of Prosopis and Gnathoprosopis, none of which have the
sscutellum or postscutellum yellow or orange, may be separated by the following
table :—
Clypeus black (females) ye ac 5 aie x aA e ae Dees
Clypeus light (males) .. ve ve we me As Se Be ie ae tle
1. Face entirely black, abdomen blue .. a ae bis cyaneomicans nigrescens Ckll.
Face entirely black, abdomen red .. ve ae LM = .. melanocephala Ckll.
Face with light lateral marks an es “ic ae ae os “th ae 2.
2. Markings white .. she ae eke a =e be - ors eburniella Ckll.
Markings yellow ee we = a ni a s*s a ee Be 3.
3. Light colour of collar not continuous with that of tubercles .. ie leptospermi CkIl.
Yellow or orange of collar continues with that of tubercles .. 7 os 2% 4.
4. More than basal half of hind femora pale orange ei: (Gnathoprosopis) amicula Sm.
Less than basal half of hind femora pale, the colour yellowish-white. . Pa: 7. Ds
5. Clypeus narrow ; first recurrent nervure meeting intercubitus (Gnathoprosopis) amiculina Ckll.
Clypeus broad .. ts ale ce = ae 5s ie ate cue Bi 6.
6. Larger; first recurrent nervure meeting intercubitus He (Gnathoprosopis) hackeri Ckll.
Smaller ; first recurrent nervure falling short of intercubitus 5 ot “ih
(Gnathoprosopis) amiculiformis Ckll.
7. Antenne entirely bright ferruginous ae A: a ae dle coronata Ckll.
Antenne not so coloured At a Ve ais or By? Ee aie ae 8.
8. Scape circular, with a yellow mark in front a ar ae ae kelvint Ckll.
Scape oval, entirely black ae Si = vs (Gnathoprosopis) bituberculata Smith.
Scape ordinary, or (in P. amatula) very short .. aa 2 ce oe * 2:
9. Face-marks bright orange ; orange of collar joining that of tubercles ; hind legs with
tegument entirely black .. .. crassifemorata Ckll.
Face-marks pale orange ; yellow of collar separated from that of tubercles ; hind legs
partly yellow Se Ke .. «anthopsyche Ckll.
Face-marks pale yellow or white, not orange Se a aa si: os ae 10.
10. Collar bright orange, continuous with tubercles ; venter with low tubercles awe
(Gnathoprosopis) simpliciventris Ckll.
Collar not orange; small species by or ake =, a Hale
1l. Scape very short, light yellow; middle tibize entirely yellow .. See anatula Ckll.
Scape otherwise ; middle tibiz at least largely dark iy8 a ae me ee 12.
12. A cuneiform black mark at each side of clypeus ae ancorata subconstricta Ckll.
No such marks on clypeus .. oe or of Se vac ¥: Se Se 13.
V3.
Larger ; face-marks pale yellow or creamy-white ; supraclypeal mark well developed
brevior Ckll.
Smaller ; face-marks clear white ; supraclypeal mark represented by a minute trans-
verse mark just above clypeus, hardly noticeable ar scintillans Ckll.
BINGHAMIELLA Cockerell.
Binghamiella antipodes (Smith)
Bright, V. (H. W. Davey).
£76 MEMOIRS OF THE QUEENSLAND MUSEUM.
EURYGLOSSA Smith.
Euryglossa furcifera Cockerell.
Female. Brisbane, 15-2-16 (Hacker).
Euryglossa calliopsiformis Cockerell.
Female. Brisbane, 10-10-16, and 8-10-18 (Hacker); Logan Road, Brisbane,,
at Leptospermum.
Male. Sunnybank, on flowers of Jacksonia scoparia R. Br., unusually small,
about 4:3 mm. long, with second submarginal cell shorter, and its upper apical angle
more acute. Another male, Brisbane, 10-10-16 (Hacker), has the band on cheeks.
broader, a triangular yellow mark on the postscutellum, and the second submarginal
cell more as in the female, with the first recurrent joining it far from base. Possibly
the Jacksonia specimen should be separated, but at present I conclude that we have-
a single very variable species. The scutellum is dark with a variable yellow band on
hind margin, and the axille have yellow spots.
Euryglossa terminata Smith.
I saw the type in the Saunders collection at Oxford, and noted that it was a.
large species, wings dilute fuliginous; fifth abdominal segment red, with golden.
hair. Smith’s measurement, “‘ length 4 lines,” is evidently erroneous. It therefore
appears practically certain that 4. hematura Ckll. is identical with terminata.
Euryglossa depressa Smith.
A female from Portland, Victoria (H. W. Davey), is evidently the true depressa.
of Smith, as is shown by the broad subtriangular facial fovez, which are described
by Smith. Th> tarsi are dark (hind tarsal joints red at ends), not rufo-piceous as.
described by Smith, but the description otherwise agrees very well. The pale
yellowish fulvous hair on occiput is as indicated by Smith. I noted at Oxford that.
E. depressa had clear hyaline wings and very broad abdomen; in the presen’
specimen the abdomen is very broad (4 mm. wide), and the wings, which are 6-7 mm.
long, are hyaline tinged with brown, the stigma ferruginous. The disc of mesothorax
and scutellum are sparsely punctured, the punctures on the scutellum small ;
flagellum very obscure brown beneath, bright red at extreme tip. The mesothorax
shows a broad concavity mesad of each wing, traversed by the parapsidal groove.
My E£. depressa sparsa appears to be correctly considered a variety of this,
but the spesies I had as H. depressa Sm., from Victoria, is distinct, and may be
described as follows :—
Euryglossa polysticta sp. n.
Female. Length 8-5-9-5 mm.; similar to H. depressa, but smaller and with.
narrower abdomen ; facial fovez linear ; flagellum dark, not red at tip ; mesothorax
anteriorly with very numerous minute punctures, and scutellum quite closely
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL, 271
punctured ; wings slightly (sometimes conspicuously) brownish, stigma very dark ;
hair of occiput very pale yellow ; abdomen dull green, hind margins of segments black ;
legs black, the tarsi ferruginous apically. Very close to #. subsericea Ckll., from
Mackay, Queensland, but larger, with longer second submarginal cell, and much
broader face.
Type from Dandenong, Victoria, Nov., 1902 (7. Kershaw). Also occurs at
Croydon.
Euryglossa hypoleuca Cockerell.
Mr. Hacker has discovered the female, and sends specimens from Caloundra,
20-1-16. This female resembles that of HL. albocuneata Ckll., but is less robust, with
the clypeus narrower, the cuneiform lateral face-marks longer, the lower margin
of clypeus, mandibles (except tips), and labrum bright ferruginous, the antenne
(including scape) ferruginous on outer side. The knees, tarsi, and anterior tibie in
front are red, but the hind tibiz are white basally. The face is concave. The
abdomen is without distinct banding, but the hind margins of the segments are
narrowly pallid.
Euryglossa politifrons sp. n.
Female. Length nearly 6 mm., with very broad abdomen; head _ black,
with the polished, bare, hardly punctured clypeus ferruginous ; supraclypeal area
highly polished, dark reddish ; front shining, with a median groove ; scape red, with
the apex intense black ; labrum and basal half of mandibles reddish ; mesothorax
and scutellum bare, ferruginous, polished, with minute very sparse punctures ; pro-
thorax red, dusky below at sides; pleura black; axilla much lighter red than
scutellum ; postscutellum and upper part of metathorax red ; tegule ferruginous ;
wings hyaline, stigma and nervures dusky reddish ; basal nervure falling far short
of nervulus; second submarginal cell receiving first recurrent nervure far from
base, the second almost or quite at apex ; legs pale ferruginous, anterior and middle
femora suffused with dusky above ; abdomen ferruginous, with subquadrate black
marks at extreme sides of the segments, apex dusky reddish.
Emerald, 1916 (#. Allen). Allied to EF. frenchii Ckll., but distinguished by
the red legs and clypeus, as well as the small size.
Euryglossa occipitalis sp. n.
Female. Length about 10 mm., very robust; head and thorax black,
abdomen very broad, ferruginous, black (with black hair) at tip, and the segments
with inconspicuous biundulate dusky (somewhat bluish) bands, failing sublaterally ;
legs very dark brown, small joints of hind tarsi clear red. Face very broad, shining,
clypeus sparsely punctured ; mandibles with an orange patch on outer side, before
apex, and just before this a reddish area ; antennz dark, flagellum reddish beneath
subapically ; occiput with bright orange-fulvous hair; mesothorax and scutellum
polished, sparsely punctured ; tegulz reddish ; wings hyaline, slightly dusky, stigma
272 MEMOIRS OF THE QUEENSLAND MUSEUM.
dark reddish, nervures fuscous ; second submarginal cell long, receiving recurrent
nervures some distance from base and apex; abdomen very sparsely and feebly
punctured ; venter light yellowish-ferruginous.
Portland, Victoria (H. W. Davey). Closely related to EH. victorie Ckll., but
without metallic colours on head and thorax, and abdomen with much more red.
Euryglossa longicornis sp. n.
Male. Length about 9 mm., not very robust, black, the abdomen with a
satiny lustre, and the hind margins of the segments obscurely brownish ; apex of
abdomen pointed. Head thick, face broad, eyes not converging below; scape
robust, shining black; flagellum very long, with the first five or six joints dull:
ferruginous beneath ; third antennal joint short, but not so short as second ; face,
labrum, and mandibles black ; clypeus and supraclypeal area shining, with strong
moderately close punctures ; hair of cheeks white, of front and occiput tinged with
yellowish ; mesothorax truncate anteriorly, very closely punctured but shining ;
scutellum with stronger rather sparse punctures; tegule dark reddish; wings
brownish, s igma and nervures dark brown ; basal falling short of nervulus ; second
submarginal cell receiving first recurrent nervure some distance from base, but
second recurrent meeting outer intercubitus; knees, anterior tibiz, middle and
hind tibiz, except broad dusky clouds, and all the tarsi, rather deep red, the colour
of the anterior tibize bright ; abdomen with no distinct bands.
Kelvin Grove, Brisbane, 27-11-11 (Hacker). A distinct species, which I
cannot associate with any described female. It must be rare at Brisbane, as the
specimen taken in 1911 remains unique.
Euryglossa walkeriana Cockerell.
Female. Brisbane, 12-9-16 (Hacker). Previously known only by the unique
type, collected at Launceston, Tasmania. It seems to be a little smaller than the type,
and the hind margin of the first abdominal segment is red.
Euryglossa neglectula subsp. mica Cockerell.
Female. Brisbane, 20-9-16 (Hacker). Male. Brisbane, 3-10-16 (Hacker).
Typical neglectula was described from ‘‘ Australia ” from an old specimen in F, Smith’s
collection.
Euryglossa reginee Cockerell.
Male. Brisbane (Hacker); Ebor, N.S.W., 1-1-16, a variation with the
flagellum dark, reddened only at base beneath.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL., 273
Euryglossa nubifera sp. n.
Male. Length about 6 mm., black, the head and thorax with rather abundant
dull white hair, and no light markings ; face broad but orbits converging below ;
clypeus dullish, with long pale hair, not conspicuously punctured ; mandibles dark
red apically; front dull; facial fovee linear; antenne long and rather thick,
flagellum obscure reddish beneath; mesothorax and scutellum dullish, without
evident punctures (the compound microscope shows a minutely tessellate surface) ;
tegule dark, with broad reddish margin posteriorly ; wings hyaline, faintly dusky,
beautifully iridescent, with a large fuliginous apical cloud ; stigma pale reddish with
dark margin, nervures fuscous ; second submarginal cell large, receiving recurrent
nervures not very far from base and apex ; femora black with red knees ; tibize and
tarsi bright ferruginous, the hind ones strongly infuscated posteriorly ; first three
abdominal segments dull black, with the broad hind margins (that on first reddish)
shining ; rest of abdomen shining; apex with a red spoon-shaped plate; venter
red, the second and third segments with dark transverse bands.
Coolangatta, Queensland, 7-9-13 (Dr. A. J. Turner). Easily recognised by the
clouded apices of anterior wings.
A female from Coolangatta, of the same date, was set apart as an entirely
different species, but on closer inspection it is evidently nubifera. It is nearly 8 mm.
long, much more robust than the male, and the strongly dusky (brownish) wings have
the apical cloud indistinct though perceptible. The outer intercubitus has an angle
at the middle, from which proceeds an appendicular vein, and there is another
similar short appendicular vein near its lower end. These are only slightly indicated
in the male. The legs are coloured as in the male, except that the hind tibiz are black.
and their tarsi largely black. The face is broad, not hairy, the clypeus with very
few weak punctures. Flagellum ferruginous beneath except at base ; mandibles with
only a faint suggestion of red near apex; facial fovee linear. The abdomen has the
first two segments dull, except the margins, the others shining. The mesothorax
and scutellum are dullish, without distinct oma There is black hair on the
abdomen just below the apex.
Euryglossa jucunda Smith.
Female. Ebor, N.S.W., 1-1-16. Length about 7 mm.; sides of face with
much white hair ; tegule dark reddish-brown ; stigma same colour as tegule ; only
mall joints of tarsi red. This is what I have determined as jucunda, but it is possible
that actual comparison with the type from Western Australia would show that
the form from New South Wales and Queensland is separable. It is also possible,
judging from Smith’s account, that the original jucwnda was a mixture of two
species.
S
274 MEMOIRS OF THE QUEENSLAND MUSEUM.
in
Abdomen red _.. re it i: bie i aie 52 s% ae a He
Abdomen not red a aP aie ye ate = Pe a 2 ete 2.
1. Mesothorax red .. ae ‘ = % if igs ie .. E. politifrons Ck.
Mesothorax black; wings hyaline .. ae Be 35 =a .. . occipitalis Ckll.
2. Mesothorax dark green. . of = 5 eS sf ie Sh .. E. walkeriana Cklil.
Mesothorax yellow ae ao Se ae as a A .. . furcifera Ckll.
Mesothorax black, marked with yellow; clypeus yellow as a ae a2 ae
Mesothorax entirely black .. = Hf: 3 as a 3 4 = 5:
3. Scape dark ; clypeus with two dark spots ; a yellow spot on front . calliopsiformis Ckll. 2
Scap «yellow, at least in front oh ae oe = Ze ae Bi os 4.
4. Axille spotted with yellow .. be Be ‘% x E. calliopsiformis Ckll. ¢
-Axille entirely black .. Te 24 “fe ae ey i .. P. humeralis Ckll.
5. Face with light markings xP -¢ st ae te oh re 53 A 6.
Face entirely black ae Ss bs Pe a se 278 or se we Ti
6. Legs mainly black; region of mouth red .. 50 i 45 .. E. hypoleuca Ckll.
Legs yellow; cheeks black .. is = 3. 3h s .. P. aurantipes Ckll.
7. Males; antenne long and slender .. ae 5e a2 pe ae a a 8.
Females ; antennz shorter... aa we me Aan aie A ae 0 32
8. Large species, about 9 mm. long... ss Sis aie oe .. E. longicornis Ckll.
Much smaller... ss “i as bi. ane 2% Se ae sa 3 9_
9. Flagellum light red beneath .. os ve Ye Se oe i oe cts 10.
Flagellum dark .. a - = * ei aes 7 _ ff A coin
10. Apex of abdomen red .. ae ws esd ares or Re a P. barbata Ckll.
Apex of abdomen not red_.. a ie Sic st 7: a E. regine Ckll..
11. Wings with a conspicuous apical cloud oe 58 ot se .. . nubifera Ckll.
Wings without an apical cloud je bts ae St: a #3 ae as 12.
12. Larger; hind tibie largely red te eh oe ane 5 E. regince Ckll., var.
Smaller; hind tibize without red... $2 56 Ks .. E. neglectula mica Cxu.
13. Large and robust ; anterior wing about 7 mm. long; abdomen greenish 4. depressa Smith..
Much smaller... Bc a Sa. x -{? ie Se Rae fe 14.
14. Abdomen metallic ; only small joints of tarsi red 2 = .. E£. jucunda Smith.
Abdomen not metallic ; tarsi largely red or nearly all red Ae si Si 28 15..
15. Larger; middle tibiz red in front .. ae ute che ae .. E. nubifera Ck
Smaller; middle tibiz black .. ee ae Ke . .. E. neglectula mica Ckll-
The following key will facilitate the separation of the species of Huryglossa
the present collection. I have also added the species of Pachyprosopis :—
PACHYPROSOPIS Perkins.
Pachyprosopis aurantires Cockerell.
Brisbane, 3-10-12 (Hacker).
Pachyprosopis humeralis Cockerell.
Oxley, Brisbane, 24-9-14 (Hacker).
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL.
bo
~I
oy
Pachyprosopis barbata Cockerell.
Male. Sunnybank, Brisbane, 19-11-13 (Hacker); Tambourine, 23-10-12.
CERATINIDA.
NEOCERATINA Perkins.
Neoceratina australensis Perkins.
Glen Innes, N.S.W., April 28, 1916. One female. The mesothorax and’
scutellum are dark-green. Previously known from Queensland.
EXONEURA Smith.
Exoneura insularis Cockerell.
Stradbroke Island, five females, Sept. 17, 1915 (Hacker).
Exoneura bicolor Smith.
Armidale, N.S.W., 5-2-15, two females ; Stanthorpe, Q., 6-11-14, one female.
Exoneura nitida n. sp.
Female. Length about 6 mm.; black, with the margins of the fovrth and.
fifth abdominal segments very narrowly and often hardly perceptibly reddened ;.
eyes rather pale purplish-brown ; face and front highly polished and shining ; face:
broad, orbits slightly converging below ; clypeus with a very broad creamy-white
band, broadest above where it occupies the whole width of the clypeus; flagellum
obscurely brownish beneath ; thorax and first three segments of abdomen smooth
and shining ; tubercles cream-colour ; tegulz black ; wings dilute brown, stigma and
nervures very dark ; under side of thorax with short pale hair ; hind tibie and tarsi
with much black hair.
Male. Length about 6mm.; similar to this sex of H. aterrima, but larger,.
head broader, eyes larger, reddish-brown ; white (creamy-white) area on face broader ;.
wings browner. The face is bare, except for a very scanty short pale pubescence.
Stradbroke Island, Q., Sept. 17, 1915, four females, 1 male (Hacker). Very
close to H. aterrima (CkIL.). but larger, with brownish wings and black legs. Also.
near EH. botanica Ckll., but differing in the legs and other characters.
Exoneura robusta n. sp.
Female (Type). Length about 6-5 mm. ; robust, black, the very broad abdomen
shining dark chestnut-red ; with the first segment (except apical margins laterally),
nearly all of second, and a transverse arched band on third, black ; face very broad,
orbits not converging ; clypeus with a broad cream-coloured band, which in the
type specimen is urn-shaped and pointed below, but in others variable, hooked at.
276 MEMOIRS OF THE QUEENSLAND MUSEUM.
sides above, and reaching clypeal margin below the sides irregular; mandibles
dark reddish in middle; antennze dark; mesothorax dullish, not highly polished ;
tubercles black, with a fringe of grey hair ; tegulz black, reddish posteriorly ; wings
strongly reddened; stigma dusky red, margined with darker; nervures fuscous ;
venter of thorax with shining white hair; hair on hind tibiz and tarsi reddened
anteriorly.
Male. Length about 6-5 mm.; more slender, with the usual male characters.
Face narrow, with long black hair, and no light markings; stigma narrower and
darker; tibiz apically, and all the tarsi red; abdomen darker, even the apical
segments suffused with black. This is possibly a different species, but probably
belongs here.
National Park, Q., Dec., 1919, four females, one male (Hacker). Close (2) to
E-hamulata Ckll., with the same broad face, but easily distinguished by the much
darker legs, only partially and very obscurely reddened, if at all. The abdomen
also is much darker. The male is remarkable for the entirely black face.
Exoneura baculifera n. sp.
Female. Length somewhat over 6 mm., but smaller than H. robusta ; black,
robust, with the broad abdomen dark chestnut-red, the first two segments mainly
black, and a transverse dusky cloud on third; knees, anterior and middle tibiz
apically, and their tarsi, rather dusky-red ; clypeus with a narrow rod-like pale
yellow stripe, often subobsolete ; orbits converging below ; antenne dark ; tegule
piceous, reddest behind ; wings strongly brownish, or reddish, stigma and nervures
dusky ferruginous; tubercles dark; hind tibiz and tarsi with much black hair,
shining reddish at apex of tibiz.
National Park, Q., Dec., 1919, five females (Hacker). Allied to E. angophore
obliterata Ckll., but easily separated by the dark legs and abdomen. It is to obliterata
what.robusta is to hamulata. |
Exoneura excavata n. sp.
Female. Length about 7-5 mm. ; black, with the abdomen dark chestnut-red,
the first segment with a broad blackish suffusion on disc; anterior knees, tibiz
apically, and all the tarsi dark red; in certain lights the. middle and hind tibiz
appear to be bright red above, but this is due to remarkable coppery hair; the
hair on hind tarsi is also red, subappressed ; face broad, excavated and basin-like,
shining (though the clypeus is minutely punctured), wholly black ; labrum convex,
dark red; mandibles reddish ; antenne entirely dark ; mesothorax and scutellum
polished ; tubercles black; hair on under side of thorax pale reddish; tegule
piceous ; wings strongly reddish ; stigma dusky-red with darker margin, nervures
dusky-reddish ; apical part of abdomen with short red hair.
National Park, Q., Dec., 1919, one female (Hacker). A remarkable species,
easily known by the black excavated face.
a |
sj
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL., 2
Exoneura diversipes n. sp.
Male. Length about 7 mm.; black, the knees, tibie at extreme apex, and
middle and anterior tarsi pale reddish, hind tarsi dark reddish; eyes very large.,.
converging below ; face with a very broad reversed T of greenish-white, including
all of clypeus except a narrow stripe down each side; lateral marks represented
by short slender lines running up from the ends of the arms to the T ; labrum greenish-
white ; face with long black hair ; antennz entirely black ; hair of head and thorax
above long, brownish-black, pleura with blackish hair, under side of thorax with
white ; tegule black; wings hyaline, stigma and nervures clear ferruginous ; all
the femora, and anterior tibiw, and tarsi slender, hind tibiz slender basally and
broad apically, their basitarsi long and very thick.
National Park, Q., Dec., 1919, three males (Hacker). Very distinct from all
described males, and I know of no female to which it could be assigned.
Exoneura rhodoptera n. sp.
Female. Length about 6 mm. ; very robust, black, with the abdomen marked
with dark red, at sides of first and second segments, a pair of hook-shaped marks on
second segment, a transverse band (weak or broken sublaterally) on third, base of
fourth, and an indistinct transverse band on fifth ; face broad, orbits parallel; clypeus
with a broad median cream-coloured bar, irregular along the margins, and emitting at
the upper end very long hook-like extensions, the whole upper margin of clypeus
being narrowly pale ; on each side of the lower end of the pale bar is a reddish area ;
no lateral face-marks ; labrum black ; flagellum obscure reddish beneath ; tegulee
reddish ; wings strongly reddish-fuliginous, stigma and nervures dark-reddish ;
tubercles black ; femora with a pale red stripe above, not reaching base ; anterior
_and middle tibiz and tarsi dark red; hind legs darker.
Stradbroke Island, Q., Sept. 17, 1915, one female (Hacker). Allied to HE.
hamulata Ckll., but with very much darker abdomen, legs, and wings.
Exoneura perpensa n. sp.
Male. Length about 6 mm. ; head and thorax black, with long hair, black on
head, dull white on thorax, very faintly yellowish dorsally ; head transverse ; eyes
very large and convex, face narrower, in middle hardly so broad as an eye ; clypeus
and labrum greenish-white, the light facial area like a reversed wineglass with an
extremely thick stem ; no lateral marks ; antenne black ; mesothorax and scutellum
not highly polished ; tegule dark; wings hyaline, faintly reddish, pale clear red
at base ; stigma narrow, ferruginous ; nervures ferruginous ; basal nervure strongly
arched, falling far short of nervulus; femora and anterior and middle tibiz and
tarsi slender ; hind tibiz claviform, their basitarsi long and thick ; knees, anterior
tibiz except basal half behind, hind tibiz in front and apically behind, hind tibiz
at apex in front and anterior and middle tarsi all light ferruginous ; abdomen parallel-
sided, broad at base (slender basally in bicolor), first segment black, second black
278 MEMOIRS OF THE QUEENSLAND MUSEUM.
with dull red apical margin, third with broader red margin, fourth red clouded with
dusky, fifth and sixth more strongly clouded; venter clear red except base and
apex.
Armidale, N.S.W., 5-2-15 (Dr. A. J. Turner). This is structurally allied to
E. diversipes. 1 cannot refer it to any described species.
Exoneura abstrusa n. sp.
Male. Length about 6-5 mm.; head, thorax, and abdomen black ; knees,
anterior, and middle tibiz, and anterior tarsi reddened, the anterior tibiz clear light
ferruginous, with a large elongate black mark basally on outer side, middle tibize
much more obscurely coloured, with a larger black mark ; head and thorax with
rather long thin pale hair, tinged with brownish dorsally ; eyes very large ; clypeus
(except a small spot on each side of middle), labrum, and linear lateral face-marks
clear ivory-white, the face-marks diverging from the clypeus at the lateral spots,
and ending very acutely some distance below level of upper margin of clypeus,
which is straight ; antenne black ; tegule very dark-brown ; wings hyaline, faintly
dusky, stigma and nervures dull ferruginous ; anterior and middle tibize and tarsi
slender, but their femora stout ; hind femora rather robust, with much white hair
beneath; hind tibiz claviform, very broad apically, hind basitarsi very thick ;
abdominal venter reddish.
Brisbane, Q., 8-2-16 (Hacker). One male. Very distinct among known males,
and apparently not to be associated with any described female.
The males of this genus show remarkable differences. In such species as
E. nitida the general appearance is more like that of a female ; the face is not hairy,
the eyes are not remarkable, and the legs present no unusual features. In contrast.
with this, EZ. diversipes has a narrow face with much long erect black hair, enormous
eyes converging below, and greatly modified legs with slender femora. Such insects
seem so far apart as to be hardly congeneric. H. abstrusa, however, is intermediate,
and the females seem all to be strictly of one genus.
The above species of Hxoneura may be separated by the following table. The
distinction between those with red and black abdomen becomes obscure in certain
members of this series, so in order to avoid any chance of error, I have repeated
them under both categories :—
Males Ae
Females : 5.
1. Abdomen at faa pokes red .. 2.
Abdomen black : 3.
2. Middle of face creamy-white .. ore Sec ae sa Ye a perpensa Ckll.
Face black ol, ar ui Ao Ac fc if She ag robusta Ckll.
3. Anterior tibiz mainly pale red; clypeus and narrow lateral marks white abstrusa Ckll.
Anterior tibize dark ae ae as Ms a ae *: ae ir ive 4.
AUSTRALIAN BEES IN THE QUEENSLAND MUSEUM.—COCKERELL, 279
4, Face with long black hair... ae a ae a4 a sg diversipes Ckll.
Face without long hair Nis as a Ses ne oh ae nitida Cll.
5. Abdomen bright red, with thin short orange hair on apical part .. =e bicolor Smith.
Abdomen dusky or dark red, or partly reddened a Par Ri rae as 6.
Abdomen black, or slightly reddened eS SE ane ie ha es ne 10.
6. Face concave, depressed, entirely black .. ME & eis ae excavata Ckll.
Face not entirely black; if nearly all black, not concave vis Be es He ae
7. Face with lateral cream-coloured spots ; abdomen very broad, reddish .. insularis Ckkll.
Face without lateral spots .. re oie ve chs ae ae of ye 8.
8. Clypeal mark broad, or with lateral hook-like extensions at upper end ae ae 9.
Clypeal mark a simple narrow stripe, not always well defined ne sis baculifera Ckll.
9. Legs largely reddened ; hook-like extensions of clypeal mark very long . . rhodoptera Ckll.
Legs black, at most knees reddish, or tibize and tarsi obscurely reddened ; face-marks
variable ag sh : fs robusta Ckll.
10. Face with light lateral spots; flagellum pallid beneath ae a insularis Ckll.
Face without light lateral spots ae at oy aie Be Gh af a hig
11. Face-mark a narrow subobsolete stripe == + ahs ae os baculifera Ckll.
Face-mark broad, or with lateral extensions above ax <¢ aa “if ae 12,
12. Anterior and middle femora with a red stripe above .. a os rhodoptera Ckll.
Anterior and middle femora without such a stripe ste Ae Sh ae nie 13:
13. Tubercles entirely black, densely fringed with grey hair Ne So robusta Ckll.
Tubercles largely light; legs black .. a oe a Fe a nitida Ckll.
MELIPONID:.
TRIGONA Jurine.
Trigona cassize Cockerell.
Workers from Brisbane, 8-2-16 (Hacker), and Caioundra, 20-1-16. (Hacker).
One of the Brisbane specimens has a dark red abdomen, but it is certainly conspecific,
and possibly not fully matured.
Trigona leeviceps Smith.
Many years ago I recorded T'rrigona canifrons Smith from Adelaide R.,
Australia, basing my identification on comparison with specimens from Ceylon,
received from Mr. E. E. Green as canifrons. Unfortunately the Ceylon specimens
were really 7’. Jeviceps Sm., and not canifrons at all. A specimen now before me,
from Gordonvale, N.Q., June, 1918, collected in the scrub by Edmund Jarvis, is
identical with the so-called canifrons of Australia. I am quite unable to clearly
separate it from Jeviceps, though it is rather slender, with the abdomen dark
sepia-brown, becoming black apically. Possibly a larger series would indicate a
distinct form, but I am inclined to think that we have the Indian le@viceps probably
accidentally introduced into Australia. Bingham states that it builds in crevices in
the walls of houses, so it seems quite possible that a nest might be carried with some
kind of merchandise. The red antenne distinguish if from the allied and certainly
native T. carbonaria Smith.
280 MEMOIRS OF THE QUEENSLAND MUSEUM.
ON THE EMERGENCE OF TWO TUBE-DWELLING
HOMOPTEROUS INSECTS.
By Henry Hacker, F.E.S.
(Plates XVII and XVIII).
PECTINARIOPHYES PECTINARIA Kirk.
A good account of the calcareous tubes made by this and an allied species,
together with the curious nymphs that inhabit them, was given by F. Ratte,! in 1884 ;
but the method of emergence was very briefly mentioned by this author. Kirkaldy?
in 1906, placed these insects in the subfamily Cercopine and erected two new genera.
Polychetophyes and Pectinariophyes for their accommodation. Subsequently? he
placed them in the subfamily Macherotine.
These calcareous’ tubes have been frequently observed by me, attached to -
small Eucalyptus saplings, and during the last two years I have included them.
amongst other studies in the life histories of Australian Homoptera. As far as I
am aware the final ecdysis of these insects is not paralleled by that known of any
other group. All the tubes seen, however, do not contain living nymphs, as those
deserted in previous seasons, owing to their durable nature, remain on the twigs for
an indefinite time. The empty tubes have a bleached appearance in comparison
with those that are occupied. The occupied tubes contain a clear liquid, and in
it the nymphs live submerged with their heads downwards. Their suctorial apparatus.
is inserted through a longitudinal slit on the inner side of the tube into the twig to
which the tube is attached.
The first indication that the insect is about to emerge is the appearance of
small bubbles at the mouth of the tube. This occurs in early spring, generally in
the evening or at night. Viewed through a lens at this stage, the posterior end of the
nymph is seen continually moving from side to side; this end protrudes for about.
a second, evidently to obtain a supply of air, and then retracts, after which fresh
bubbles are blown; this renewing of the air supply takes place at intervals. The
operation Gontinues for about an hour, by which time a large mass of froth has been
produced, covering the mouth of the tube and hanging over the side. When about
to emerge, the nymph forces its way to the top of the tube, protruding its posterior
end first until the legs have reached the lip. It then swings itself over, and with
the Lead now upward it climbs down the outer side of the tube until it is merged
1 Pree. Linn. Soc. N.S.W., Vol. LX, p. 1164, 1884.
* Bull. Haw. Sugar Plant. Exp. Sta. i, pp. 384-386, 1906.
° Bull. Haw. Sugar Plant. Exp. Sta. xii, p. 10, 1913.
EMERGENCE OF TWO TUBE-DWELLING HOMOPTEROUS INSECTS. 281
into the froth which has accumulated on the lower side. There is constant move-
ment inside the mass of froth caused by the insect getting out of its nymphal skin.
The froth now gradually subsides; all movement has ceased, and the newly-
emerged insect is seen clinging to the empty nymph skin, which in turn is clinging
to the side of the tube.
The pale-yellow wingless insect remains quite motionless. After a short
interval tiny tegmina and wingbuds begin to appear. These expand rapidly, and
in half an hour from the time they were first seen are fully developed. While this
growth develops, and for some time after, the wings hang down perpendicularly ;
they are then suddenly flexed once or twice, and closed to their normal roof-like
position. At this stage, though rather soft, the insect is able to walk and jump,
but if left undisturbed it will remain quietly on the twig until the next day.
The time occupied by the metamorphosis of this insect is about an hour,
and the total period from the first appearance of the froth, about two hours. The
capture of adult specimens by me, in September and January, proves that there are
two broods a year.
POLYCHATOPHYES SERPULIDIA Kirk.
This insect occurs in the Brisbane district, but more sparingly than the
previous species. The tubes differ from that species in their larger size, darker
colour, and the transverse serrated lines, which give them a coarser texture. They
are invariably attached to the twigs for their entire length, while in the previously
mentioned species, the upper part of the tube is always bent out away from the
twig. This causes a slight difference in the method of exit between the two species.
When the froth is coming from the tube it often runs down the twig instead of the
tube, dependent on the angle at which the twig is inclined. The emerging nymph
following the froth consequently often clings to the twig instead of the tube to
undergo its metamorphosis. When this occurs, the insect immediately after
emergence generally walks a little way up the twig and there stops while its wings
develop. In other respects its method of emergence is similar to that of
Pectinariophyes pectinaria, and the time occupied is about the same.
F. Ratte remarks :—‘‘ In the dry parts of the interior it is probable that the
water contained in these shells is resorted to for drinking by the ants so numerous
in Australia, as if it was a speciality among the small homopterous insects to provide
during their life for the Formicide.” All the evidence which I have acquired in
the coastal district is against the above opinion; no ants have been seen drinking
the liquid, nor were any noticed in the vicinity of the tubes. It is well known that
many Coccide, Membracide, &c., excrete liquids which are eagerly sought by ants.
In return the Homoptera receive certain services from the ants, so the benefit is
reciprocal ; but in the case of these soft tube-dwelling nymphs it would be of no
service to be robbed of their protective liquid, without which they would speedily
perish.
282 MEMOIRS OF THE QUEENSLAND: MUSEUM.
REMARKS ON THE ILLUSTRATIONS.
(Plates XVII and XVIII.)
Pectinariophyes pectinaria Kirk.—Figs. 1, 2, 3, and the upper insect in fig. 4.
Polychetophyes serpulidia Kirk.—Figs. 5, 6, 7, 8, and the lower insect in fig. 4.
Figs. 1 and 5 show the first appearance of the nymphs of both species, while forcing
their way, posterior ends first, out of their tubes. The next stage is shown at fig. 6,
where the nymph is clinging to the twig, while its skin is about to split longitudinally
up the back. In figs. 2 and 7, the upper portion of the insects are free, while they
are supported by their lower parts which are still enclosed in the nymphal skins ;
the wings are seen beginning to develop. Figs. 3 and 8 show the fully developed
insects resting in their characteristic positions. In fig. 8, a circular anal plate can be
seen on the empty nymph skin; it is peculiar to this species, and is used by the
nymph as an operculum to close the mouth of its tube.
UEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Prare XVIL.
x3 diam. FitQ tps XS4 dean.
EMERGENCE OF TUBE-DWELLING HomoprTERA, Hacker.
Face page 282.
Se
a
MEMOIRS OF THE QUEENSLAND MUSEUM, Vou. VII, Puate XVIII.
X3 diam,
X3 dian. X3 diam.
EMERGENCE OF TUBE-DWELLING HomoptErRA, Hacker.
Face page 282,
SOME AUSTRALIAN WASPS.—HACKER AND COCKERELL. 283
SOME AUSTRALIAN WASPS OF THE GENERA
ZOYPHIUM AND ARPACTUS.
By HENRY HAcKER AND T. D. A. COCKERELL.
Famity CRABRONID.
Supramity PARANYSSONIN. >
Genus ZOPYHIUM Kohl.
The genus Zoyphium Kohl, based on a single species as recently as 1893,
‘is now found to include a large number of Australian forms, presenting excellent
specific characters. The following table separates those at present known, but it
cannot be doubted that many others will be found when collectors in various parts
of Australia interest themselves in these small wasps :—
Females ate ae Bt we as ie ais ae a te ae 1.
Males oi sie Ee Ef es 3. ss ae a3 = ate 15.
Two cubital cells in anterior wings .. ke ts a ie Sac frontale Turner.
Three cubital cells “he Et ate ve eve he a a ai: ar ils
1. First abdominal segment with a tooth or tubercle on each side ne 6 Le 2.
First segment without such teeth .. oe “ie ake Sy Se ye se a
2. Clypeus with two small teeth on each side se oh sericeum Kohl.
Clypeus with one tooth on each side; first esate! enter a campanulate, not
truncate, with a pair of basal keels, and a small tubercle on each side
not far from base .. . : Be oo flavofasciatum Turner.
33. Abdomen black in middle and pallid at either ela its basal segment very short and
broad, abruptly truncate ; clypeus without lateral teeth dipteroides Turner.
Abdomen red in middle ze : : St ee 4.
Abdomen black, or with hind margins of dopants more or ame fad es che ve
4. First abdominal segment black, with red hind margin; pronotum black; clypeus
eream-colour .. Bf SE sg = ie -. > ornatum n. sp.
First abdominal segment red .. af 2 a <- aie a ae 38 5.
5. Upper margin of pronotum yellow, scutellum and postscutellum mainly red .. collare n. sp.
Pronotum black; scutellum and postscutellum without red .. os BS a: 6.
Mesonotum dark blue .. ies sie ae ate Bes cri erythrosoma Turner.
Mesonotum black ies : am oe : a3 a splendidum n. sp.
‘7. Pronotum with an Barend cream-coloured paid above ; mesonotum faintly
brassy, very finely and closely punctured ate rele funebre Turner.
Pronotum with upper margin wholly dark =f Ae a ae a ere 8.
‘8. Antenne red or pale brownish Be se om a Le Ef Ye oa ae
Antennz wholly or mainly black .. a Ete a ¥ a. ae af 10.
9. Legs bright yellow oe a oF Sic at ae Se tie affine n. sp.
Legs pale ferruginous brown .. v6 oA ar oa dye 4e Kohli Turner.*
* In his key, Turner says that 9 kohlii has no clypeal teeth, but his deseriptions indicate
a single short tooth on each side.
284 MEMOIRS OF THE QUEENSLAND MUSEUM.
10. Pronotum strongly emarginate in middle above .. Fc Ag: se Se 46 We
Pronotum not emarginate ate ane a ae ye Ac ae zs aie 12.
11. Tibiz entirely red 34 os + St te are in emarginatum n. sp-
Tibie not red .. A: va he Sc Ge be af oi nigrum n. sp.
12. Legs mainly black, not red or yellow aie i sts a a argyreum Nn. sp.
Legs red 4H 2 ate s ahs ase as oe oe . ae 13.
13. Wings strongly dusky .. ee a6 ae ah ae As .. fuscipenne n. sp..
Wings clear hyaline... oe oe oe sla Bx i a oe ae 14..
14. Face with golden hair .. ve fe =e 33 52 se Se crassicorne Ckll.
Face without golden hair ae i: ye 40 he Rie ..
DO SUPE ay APPS A ATI SE ABE ATLA SAAT OC
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“WITH NINETEEN PLATES AND Sas
_SEVENTY-TWO_ TEXT-FIGURES. re aa
”
Or Oty a RSE AINE ele oe howe gear Gack
x DATE OF ISSUE OF: ‘FINAL. PART:
_ DECEMBER: 19, 1922.
"EDITED BY THE DIRECTOR, HEBER A. LONGMAN, F.LS: |
j ¥ oy By Bt , ’ : : ‘ ;
Why tet “ “and prince m Aentratiac ” a Shite cit
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Notes end, ‘iysratione of |
Lit SE
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