Angelo, R. and D.E. Boufford. 2013. Atlas of the flora of New England: Fabaceae. Phytoneuron 2013-2: 1-15 + map pages 1-
21. Published 9 January 2013. ISSN 2153 733X

ATLAS OF THE FLORA OF NEW ENGLAND: FABACEAE

RAY ANGELO’ and DAVID E. BOUFFORD”
Harvard University Herbaria
22 Divinity Avenue
Cambridge, Massachusetts 02138-2020
'rangelo@oeb.harvard.edu
*david_boufford@harvard.edu

ABSTRACT

Dot maps are provided to depict the distribution at the county level of the taxa of
Magnoliophyta: Fabaceae growing outside of cultivation in the six New England states of the
northeastern United States. The maps treat 172 taxa (species, subspecies, varieties, and hybrids, but
not forms) based primarily on specimens in the major herbaria of Maine, New Hampshire, Vermont,
Massachusetts, Rhode Island, and Connecticut, with most data derived from the holdings of the New
England Botanical Club Herbarium (NEBC). Brief synonymy (to account for names used in standard
manuals and floras for the area and on herbarium specimens), habitat, chromosome information, and
common names are also provided.

KEY WORDS: flora, New England, atlas, distribution, Fabaceae

This article is the eleventh in a series (Angelo & Boufford 1996, 1998, 2000, 2007, 2010,
2011a, 2011b, 2012a, 2012b, 2012c) that presents the distributions of the vascular flora of New
England in the form of dot distribution maps at the county level (Figure 1). Seven more articles are
planned. The atlas is posted on the internet at http://neatlas.org, where it will be updated as new
information becomes available.

This project encompasses all vascular plants (lycophytes, pteridophytes and spermatophytes)
at the rank of species, subspecies, and variety growing independent of cultivation in the six New
England states. Hybrids are also included, but forms and other ranks below the level of variety are not.
The dots are based on voucher specimens primarily 4 in New England ne pans (of colleges, universities,
botanical gardens, and public museums) tions outside of cultivated
habitats. This eleventh installment includes the family Fabaceae (Magnoliophyta). Of the 172 taxa
treated, 106 are not native to the region. Future accounts will treat the distribution of additional non-
monocot angiosperms.

The habitat data are distillations from a variety of sources augmented by our own field
observations. An attempt was made to indicate habitat information as it applies to a particular taxon in
New England rather than to the entire range of the taxon. Habitat information is not provided for
hybrid taxa.

It is our hope that these articles will stimulate additional field work to supplement the
distributions portrayed in the maps. The New England Botanical Club herbarium has proven to be the
most important resource for this project. We are eager to receive information on voucher specimens in
public herbaria documenting range extensions and filling county gaps in distributions. Similarly,
because the atlas of the New England flora will be continuously updated as new information becomes
available, we are eager to receive notification of published corrections of cytological information and
new, d I counts for taxa in the New England flora.

Angelo and Boufford: Atlas of New England flora 2

MATERIALS AND METHODS
Materials and methods are as outlined in Angelo and Boufford (1996) and in a web version
(Angelo & Boufford 2011c) and are not repeated here.

TAXONOMY AND FORMAT

The taxonomy and nomenclature adopted for this work follow that of draft treatments for the
Flora of North America project, except that families, genera, and species are arranged alphabetically.
The families and their circumscription do not necessarily reflect current views on relationships or
composition. The Angiosperm Phylogeny Website (Stevens 2001 onwards) should be consulted for a
continuously updated treatment of families and their inclusive genera. Named and unnamed hybrid
taxa are placed alphabetically au ‘the end of the genus in which they occur. Unnamed hybrids combine
the names of the tically by epithet. Taxa that are not native to New England are
indicated by uppercase text. Unpublished names are not used, even if publication is pending.

Chromosome numbers are taken primarily from draft treatments for the Flora of North
America project and from Goldblatt and Johnson (1979-).

Synonymy is provided primarily with respect to names accepted in standard manuals covering
New England published from 1950 onward, including Fernald (1950), Gleason and Cronquist (1991),
and Seymour (1982) and on herbarium labels in New England herbaria. Synonyms have not been
provided where the distribution for the synonymized name does not include New England.

The following list (which includes excluded taxa) will aid readers in finding familiar names
that have been transferred to other taxa:

Cassia (in part) a Chamachrista
Cassia (in part) =o Senna
Coronilla (in part) => Securigera
Desmodium (in part) => Hylodesmum
Lespedeza (in part) => Kummerowia
Lotus (in part) => Acmispon
Trigoneila (in part) => Medicago

The following species have been reported from our area but are excluded for the reasons noted:

CYTISUS x PRAECOX Bean (C. MULTIFLORUS (L’Héritier) Sweet x ?) [no voucher
found; reported from Plymouth County, Massachusetts]

LABLAB PURPUREUS (Linnaeus) Sweet (DOLICHOS LABLAB Linnaeus) [no voucher
found; reported from Worcester County, Massachusetts]

LATHYRUS APHACA Linnaeus [no voucher found; reported from Berkshire County,
Massachusetts]

LATHYRUS PANNONICUS (Jacquin) Garcke subsp. PANNONICUS [no voucher found;
reported from Massachusetts]

LATHYRUS SATIVUS Linnaeus [no voucher found; reported from Middlesex County,
Massachusetts]

Angelo and Boufford: Atlas of New England flora 3

LESPEDEZA THUNBERGI (de Candolle) Nakai [no voucher found; reported from Plymouth
County, Massachusetts]

LOTUS TENUIS Waldstein & Kitaibel ex Willdenow [no voucher found; reported as collected
from Newport County, Rhode Island]

LUPINUS ANGUSTIFLORUS Eastwood [mistakenly listed in Maine checklist for L.
ANGUSTIFOLIUS Linnaeus]

LUPINUS ANGUSTIFOLIUS Linnaeus [no voucher found; reported from Penobscot County,
Maine]

LUPINUS NOOTKATENSIS Donn ex Sims var. NOOTKATENSIS [no voucher found;
reported from northern New England]

MEDICAGO DISCIFORMIS de Candolle [no voucher found; reported from Worcester
County, Massachusetts]

MELILOTUS ALTISSIMUS Thuillier [no voucher found; reported from Maine]

ONOBRYCHIS VICHFOLIA Scopoli [no voucher found; reported from Massachusetts and
Vermont]

SENNA CORYMBOSA (Lamarck) H.S. Irwin & Barneby [no voucher found; reported from
Massachusetts]

THERMOPSIS MOLLIS (Michaux) M.A. Curtis [New England herbarium specimens with this
name are misidentified THERMOPSIS VILLOSA (Walter) Fernald & B.G. Schubert]

THERMOPSIS MONTANA Nuttall var. MONTANA [Voucher specimen from Maine is
misidentified THERMOPSIS VILLOSA (Walter) Fernald & B.G. Schubert]

TRIFOLIUM CAROLINIANUM Michaux [no voucher found; reported from Chitteneden
County, Vermont]

TRIFOLIUM FRAGIFERUM Linnaeus [no voucher found; reported from Lamoille County,
Vermont]

TRIFOLIUM GLOMERATUM Linnaeus [no voucher found; reported from Worcester County,
Massachusetts]

VICIA SEPIUM Linnaeus var. MONTANA W.D.J. Koch [no voucher found; reported from
York County, Maine]
ANGIOSPERMAE (MAGNOLIOPHYTA) - ANGIOSPERMS
FABACEAE

ACMISPON AMERICANUS (Nuttall) Rydberg var. AMERICANUS—Frairie Trefoil (Figure 2). 2n =
14. Railroads, waste places. From farther west. [HOSACKIA AMERICANA (Nuttall) Piper;

Angelo and Boufford: Atlas of New England flora 4

LOTUS AMERICANUS — misapplied; L. PURSHIANUS Clements & E.G. Clements; L.
UNIFOLIOLATUS (Hooker) Bentham]

ALBIZIA JULIBRISSIN Durazzini—Silk-tree (Figure 2). 2n = 26. Waste places. From Asia.

AMORPHA FRUTICOSA Linnaeus—False Indigo (Figure 2). 27 = 40. Roadsides, thickets, waste
places. From farther south and west. [4. FRUTICOSA var. OBLONGIFOLIA E.J. Palmer]

Amphicarpaea bracteata (Linnaeus) Fernald var. bracteata—Hog-peanut (Figure 2). 2n = 40. Rich
woods, thickets. [4. monoica Elliott]

Amphicarpaea bracteata (Linnaeus) Fernald var. comosa (Linnaeus) Fernald—Pitcher’s Hog-peanut
(Figure 2). 2n =? Rich woods, thickets.

ANTHYLLIS VULNERARIA Linnaeus—Kidney-vetch (Figure 2). 2n = 12 (Europe). Roadsides, waste
places, fields. From Eurasia, northern Africa.

Apios americana Medikus—Groundnut (Figure 2). 27 = 22, 31 = 33. Riverbanks, rich, moist thickets,
primarily along streams and lakes.

ARACHIS HYPOGAEA Linnaeus—Common Peanut (Figure 2). 2n = 40. Roadsides, waste places,
fields. From South America.

Astragalus alpinus Linnaeus var. brunetianus Fernald—Northern Milk-vetch (Figure 2). 21 = ?
Calcareous river ledges and gravels.

Astragalus canadensis Linnaeus var. canadensis—(Figure 3). 2n = 16. Shores.

ASTRAGALUS CONTORTUPLICATUS Linnaeus—Hungarian Milk-vetch (Figure 3). 27 = 16. Wool
waste. From Eurasia.

Astragalus eucosmus B.L. Robinson—(Figure 3). 21 = 32. Calcareous, rocky or gravelly shores of
streams.

ASTRAGALUS GLYCYPHYLLOS Linnaeus—Liquorice Milk-vetch (Figure 3). 2n = 16. Roadsides,
waste places. From Eurasia.

Astragalus robbinsii (Oakes) A. Gray var. robbinsii—(Figure 3). 2n = ? Dry, calcareous ledges
above water. [considered to be extinct since 1894

Astragalus robbinsii (Oakes) A. Gray var. jesupii Eggleston & E. Sheldon—(Figure 3). 2n = ? Rocky
stream banks. [A. jesupii (Eggleston & E. Sheldon) Britton]

Astragalus robbinsii (Oakes) A. Gray var. minor (Hooker) Barneby—(Figure 3). 21 = ? Calcareous,
ledges, cliffs and talus. [4. blakei Eggleston]

BAPTISIA AUSTRALIS (Linnaeus) R. Brown var. AUSTRALIS—Blue False Indigo (Figure 3). 2 = ?
Roadsides, fields, thickets, shores. From farther south and west.

Angelo and Boufford: Atlas of New England flora 5

BAPTISIA LEUCOPHAEA Nuttall—(Figure 3). 2n = ? Railroads in dry sand. From farther west. [B.
LEUCOPHAEA var. GLABRESCENS Larisey,; B. BRACTEATA Muhlenberg ex Elliott var.
GLABRESCENS (Larisey) Isely,; B. BRACTEATA var. LEUCOPHAEA (Nuttall) Kartesz &
Gandhi]

Baptisia tinctoria (Linnaeus) R. Brown—Wild Indigo (Figure 4). 2n = 18. Dry fields, dry, open
woods and clearings, in sterile or sandy soil. [B. tinctoria var. crebra Fernald]

CARAGANA ARBORESCENS Lamarck—Siberian Peatree (Figure 4). 27 = 16, 20, 24. Rich thickets,
fields, roadsides, waste places. From central and eastern Asia.

Cercis canadensis Linnaeus var. canadensis—Redbud (Figure 4). 2n = 14. Rocky woods.

Chamaecrista fasciculata (Michaux) Greene var. fasciculata—Partridge-pea (Figure 4). 2n = 16.
Sandy, open fields and clearings, usually dry, roadsides. [Cassia fasciculata Michaux; C.
chamaecrista — misapplied]

Chamaecrista nictitans (Linnaeus) Moench subsp. nictitans—Wild Sensitive Plant (Figure 4). 2n =
16. Gravelly or sandy, open soil, fields, roadsides. [Cassia nictitans Linnaeus]

CICER ARIETINUM Linnaeus—Chick-pea (Figure 4). 2n = 16. Waste places. From Turkey (and
long in cultivation).

CLADRASTIS KENTUKEA (Dumont de Courset) Rudd—American Yellowwood (Figure 4). 21 = 28.
ich woods, near parks or cemeteries. From farther south and west. [C. LUTEA (F.
Michaux) K. Koch]

COLUTEA ARBORESCENS Linnaeus—Bladder-senna (Figure 4). 2n = 16 (Europe). Roadsides,
railroads, waste areas, hillsides. From Europe, northern Africa.

CORONILLA SCORPIOIDES (Linnaeus) W.D.J. Koch—Annual Scorpion Vetch (Figure 4). 2 = 12.
Roadsides. From Eurasia, northern Africa.

Crotalaria sagittalis Linnaeus var. sagittalis—Common Rattlebox (Figure 5). 2n = 32. Sandy or
gravelly, often dry, open soil, shores, waste areas, railroads, roadsides.

CYTISUS SCOPARIUS (Linnaeus) Link var. SCOPARIUS—Scot’s Broom (Figure 5). 27 = 46, 48.
Sandy fields, openings and roadsides, dunes, beaches. From Europe.

DALEA CANDIDA Willdenow var. CANDIDA—White Prairie-clover (Figure 5). 2n = 14. Railroads.
From farther west.

DALEA LEPORINA (Aiton) Bullock—Foxtail Prairie-clover (Figure 5). 2n = 14. Rich, open soil,
waste places. From farther west, Mexico, Central and South America.

DALEA PURPUREA Ventenat var. PURPUREA—Purple Prairie-clover (Figure 5). 27 = 14. Sandy
waste areas. From farther west.

Desmodium canadense (Linnaeus) de Candolle—Showy Tick-trefoil (Figure 5). 2n = 22. Thickets,
fields, open woods, meadows, riverbanks, sandy, open soil, roadsides, railroads, waste places.

Angelo and Boufford: Atlas of New England flora 6
Desmodium canescens (Linnaeus) de Candolle—(Figure 5). 2n = ? Dry, sandy woods, fields, thickets,
roadsides and waste areas.

Desmodium ciliare (Muhlenberg ex Willdenow) de Candolle var. ciliare—(Figure 5). 2n = ? Dry,
often sandy, woods and clearings.

Desmodium cuspidatum (Muhlenberg ex Willdenow) de Candolle ex G. Don var. cuspidatum—(Figure
5). 2n =? Rocky, rich, open woods.

Desmodium glabellum (Michaux) de Candolle—(Figure 6). 2n = 22. Dry, sandy woods, clearings,
roadsides. [D. dillenii Darlington — in part, ambiguous name]

Desmodium marilandicum (Linnaeus) de Candolle—(Figure 6). 2n = ? Dry, open, rocky or sandy
woods, fields, roadsides.

Desmodium obtusum (Muhlenberg ex Willdenow) de Candolle—(Figure 6). 21 = ? Dry, sandy or
rocky, open woods. [D. rigidum (Elliott) de Candolle]

D li icul (Linnaeus) de Candolle var. paniculatum—(Figure 6). 2n = 22. Dry, open,
often rocky or sandy woods, dry, woodland borders, thickets, clearings, fields, roadsides.

Desmodium perplexum B.G. Schubert—(Figure 6). 2n = ? Dry, sandy woods, dry, woodland borders,
roadsides. [D. dillenii Darlington — in part, ambiguous name]

D li lium de Candolle—(Figure 6). 2n = 22. Dr, often rocky, woods, clearings,
fields, woodland margins, fields, roadsides.
D li lifolium Torrey & A. Gray—(Figure 6). 2n = ? Dry, sandy soils, roadsides,
railroads.
— Desmodium hybrid —

Desmodium * humifusum (Muhlenberg ex Bigelow) Beck—(Figure 6). [D. paniculatum (Linnaeus)
de Candolle var. paniculatum * D. rotundifolium de Candolle]

GALEGA OFFICINALIS Linnaeus—Professor-weed (Figure 6). 21 = 16. Roadsides, waste areas,
wet, salt meadow borders. From Eurasia, northern Africa.

GENISTA TINCTORIA Linnacus—Dyer’s Greenweed (Figure 7). 27 = 48. Dry fields, woodland
margins, roadsides, waste places, usually in dry, sterile, sandy or rocky soil. From Eurasia.

GLEDITSIA TRIACANTHOS Linnaeus—Honey-locust (Figure 7). 2 = 28. Roadsides, riverbanks,
shores, moist woods, waste areas. From farther west and south.

GLYCINE MAX (Linnaeus) Merrill—Soybean (Figure 7). 2n = 40. Fields, railroads, roadsides. From
eastern Asia.

GLYCYRRHIZA LEPIDOTA Pursh—Wild Licorice (Figure 7). 27 = 16. Waste places, fields. From
farther west.

Angelo and Boufford: Atlas of New England flora 7

GYMNOCLADUS DIOICUS (Linnaeus) K. Koch—Kentucky Coffeetree (Figure 7). 27 = 28. Waste
places, woodland margins, dry hillsides. From farther west and south.

Hedysarum alpinum Linnacus—(Figure 7). 2n = 14. Calcareous river-shores, ledges. [H. alpinum
var. americanum Michaux]

Hylodesmum glutinosum (Muhlenberg ex Willdenow) H. Ohashi & R.R. Mill—(Figure 7). 2n = ?
Rich, dry or rocky woods. [Desmodium glutinosum (Muhlenberg ex Willdenow) Alph. Wood;
D. acuminatum (Michaux) de Candolle]

Hylodesmum nudiflorum (Linnaeus) H. Ohashi & R.R. Mill—(Figure 7). 2n = ? Rich, dry or rocky
woods. [Desmodium nudiflorum (Linnaeus) de Candolle]

KUMMEROWIA STRIATA (Thunberg) Schindler—Japanese Clover (Figure 7). 2n = 22. Sandy
roadsides, gravel pits. From eastern Asia. [Lespedeza striata (Thunberg) Hooker & Arnott]

LABURNUM ANAGYROIDES Medikus—Golden Chain-tree (Figure 8). 2n = 42, 50. Woodland
margins. From Europe.

— Laburnum hybrid —

LABURNUM * WATERERI (Wettstein) Dippel—(Figure 8). [L. ALPINUM (Miller) J. Presl * L.
ANAGYROIDES Medikus]

Lathyrus japonicus Willdenow var. maritimus (Linnaeus) Kartesz & Gandhi—Beach Pea (Figure 8).
2n = 14. Sea beaches, gravelly sea shores, sandy or gravelly shores of Lake Champlain. [L.
Japonicus var. glaber (Seringe) Fernald; L. maritimus Bigelow var. maritimus; L. maritimus
var. glaber (Seringe) Eames]

Lathyrus japonicus Willdenow var. pellitus Fernald—(Figure 8). 2n = 14. Sea beaches, gravelly sea
shores, sandy or gravelly shores of Lake Champlain. [L. maritimus Bigelow var. pellitus
(Fernald) Gleason]

LATHYRUS LATIFOLIUS Linnaeus—Everlasting Pea (Figure 8). 27 = 14. Roadsides, waste places,
fields, thickets. From Europe, northern Africa.

Lathyrus ochroleucus Hooker—Pale Vetchling (Figure 8). 27 = 14. Dry or rocky banks and bluffs.

LATHYRUS ODORATUS Linnaeus—Sweet Pea (Figure 8). 2n = 14. Waste places. From southern
Italy.

Lathyrus palustris Linnaeus—Marsh Vetchling (Figure 8). 2n = 14, 42. Riverbanks, salt marshes,
shores, meadows, swamps, damp thickets, beach bluffs. [Z. palustris var. linearifolius
Seringe, L. palustris var. macranthus (T.G. White) Fernald, L. palustris var. myrtifolius
(Muhlenberg ex Willdenow) A. Gray; L. palustris var. pilosus (Chamisso) Ledebour]

LATHYRUS PRATENSIS Linnaeus—Yellow Vetchling (Figure 8). 2n = 9, 14, 16, 28, 42. Fields,
roadsides, railroads, waste places, meadows, shores. From Eurasia, northern Africa.

Angelo and Boufford: Atlas of New England flora &

LATHYRUS SYLVESTRIS Linnaeus—F lat Pea (Figure 9). 27 = 14. Field borders, thickets, roadsides,
waste places. From Eurasia, northern Africa.

LATHYRUS TUBEROSUS Linnaeus—Earthnut Pea (Figure 9). 27 = 14, Fields, meadows, roadsides,
railroads. From Eurasia.

LENS CULINARIS Medikus—Lentil (Figure 9). 2n = 14, 21, 26. Roadsides, waste places. Origin
unknown (long in cultivation).

Lespedeza angustifolia (Pursh) Elliott—(Figure 9). 2n = 20. Dry, open, sandy soil, sandy, pond
shores.

LESPEDEZA BICOLOR Turezaninow—Shrubby Bush-clover (Figure 9). 2n = 18, 22, 42. Waste
places, disturbed, open, sandy or gravelly soil, roadsides. From eastern Asia.

Lespedeza capitata Michaux—(Figure 9). 2n = 20. Dry, open, sandy soil, roadsides, railroads, fields,
dry, rocky or sandy woods, waste places. [Z. capitata var. stenophylla Bissell & Fernald; L.

capitata var. velutina Fernald; L. capitata var. vulgaris Torrey & A. Gray]

LESPEDEZA CUNEATA (Dumont de Courset) G. Don—Chinese Bush-clover (Figure 9). 2n = 20.
Dry roadsides, fields, waste places. From eastern Asia, Australia.

LESPEDEZA CYRTOBOTRYA Miquel—(Figure 9). 2n = 22. Fields. From eastern Asia.

Lespedeza frutescens (Linnaeus) Hornemann—(Figure 9). 27 = 20. Dry, rocky woods. [Z. violacea —
misapplied]

Lespedeza hirta (Linnaeus) Hornemann subsp. hirta—(Figure 10). 2n = 20. Dry, open woods and
other dry soils, often sandy or rocky.

Lespedeza procumbens Michaux—Trailing Bush-clover (Figure 10). 2n = 20. Dry, sandy or rocky,
open woods and clearings.

Lespedeza repens (Linnaeus) W.P.C. Barton—Creeping Bush-clover (Figure 10). 27 = 20. Sandy or
rocky, open woods, dry ledges, dry, rocky fields.

Lespedeza stuevei Nuttall—(Figure 10). 2n = 20. Dry woods, dry, sandy, rocky or sterile, open soil,
roadsides.

Lespedeza violacea (Linnaeus) Persoon—(Figure 10). 2n = 20. Dry, open, often rocky woods, dry,
open, rocky soil, thickets, roadsides. [L. intermedia — misapplied; see L. frutescens for

traditional, misapplied use of name “ZL. violacea’’ (Reveal & Barrie 1991]

Lespedeza virginica (Linnaeus) Britton—Slender Bush-clover (Figure 10). 2n = 20. Dry, open,
usually sandy or rocky soil, dry, open, often rocky woods, fields, roadsides.

— Lespedeza hybrids —

Angelo and Boufford: Atlas of New England flora 9

Lespedeza * acuticarpa Mackenzie & Bush (pro species)—(Figure 10). [Z. frutescens (Linnaeus)
Hornemann x L. virginica (Linnaeus) Britton; L. violacea — misapplied x L. virginica
(Linnaeus) Britton]

Lespedeza angustifolia (Pursh) Elliott x L. capitata Michaux—(Figure 10).

Lespedeza * brittonii E.P. Bicknell (pro species)—(Figure 10). [Z. procumbens Michaux x L.
virginica (Linnaeus) Britton; L. procumbens var. elliptica S.F. Blake]

Lespedeza capitata Michaux x L. violacea (Linnaeus) Persoon—(Figure 11). [Z. capitata Michaux x
L. intermedia — misapplied]

Lespedeza frutescens (Linnaeus) Hornemann x L. procumbens Michaux—(Figure 11). [L. violacea —
misapplied x L. procumbens Michaux]

Lespedeza frutescens (Linnaeus) Hornemann x L. violacea (Linnaeus) Persoon—(Figure 11). [L.
violacea — misapplied x L. intermedia — misapplied]

Lespedeza hirta (Linnaeus) Hornemann subsp. hirta x L. procumbens Michaux—(Figure 11).
Lespedeza hirta (Linnaeus) Hornemann subsp. hirta x L. stuevei Nuttall—(Figure 11).

Lespedeza hirta (Linnaeus) Hornemann subsp. hirta x L. violacea (Linnaeus) Persoon—(Figure 11).
[L. hirta (Linnaeus) Hornemann subsp. hirta x L. intermedia — misapplied

Lespedeza hirta (Linnaeus) Hornemann subsp. hirta x L. virginica (Linnaeus) Britton—(Figure 11).

Lespedeza * longifolia de Candolle (pro species)—(Figure 11). [L. capitata Michaux x L. hirta
(Linnaeus) Hornemann subsp. hirta;, L. hirta var. dissimulans Fernald]

Lespedeza * neglecta Mackenzie & Bush (pro species)—(Figure 11). [L. stuevei Nuttall x L.
virginica (Linnaeus) Britton]

Lespedeza * nuttallii Darlington (pro species)—(Figure 12). [L. frutescens (Linnaeus) Hornemann x
L. hirta (Linnaeus) Hornemann subsp. hirta, L. violacea — misapplied x L. hirta subsp. hirta]

Lespedeza procumbens Michaux * L. stuevei Nuttall—(Figure 12).

Lespedeza * simulata Mackenzie & Bush (pro species)—(Figure 12). [L. capitata Michaux x L.
virginica (Linnaeus) Britton]

Lespedeza stuevei Nuttall x L. violacea (Linnaeus) Persoon—(Figure 12). [L. stuevei Nuttall x L.
intermedia — misapplied]

LOTUS CORNICULATUS Linnaeus—Bird’s-foot Trefoil (Figure 12). 2n = 12, 24, 28, 36.
Roadsides, fields, waste places, meadows. From Eurasia, Africa.

Lupinus perennis Linnaeus var. perennis—Wild Lupine (Figure 12). 2n = 48, 96. Dry, sandy or
gravelly soil, roadsides, railroads, dry, open woods, clearings, fields.

Angelo and Boufford: Atlas of New England flora 10

Lupinus perennis Linnaeus var. occidentalis 8. Watson—{Figure 12). 2n = ? Dry, sandy or gravelly
soil, roadsides, railroads, dry, open woods, clearings, fields.

LUPINUS POLYPHYLLUS Lindley subsp. POLYPHYLLUS—Garden Lupine (Figure 12). 2n = 48,
96. Roadsides, fields. From farther west.

MEDICAGO ARABICA (Linnaeus) Hudson—Spotted Medick (Figure 12). 21 = 16. Waste places.
From Eurasia, northern Africa.

MEDICAGO LACINIATA Miller—(Figure 13). 2n = 16. Wool waste, waste places. From south
central and southwestern Asia, Africa. [TRIGONELLA LACINIATA — misapplied]

MEDICAGO LUPULINA Linnaeus—Black Medick (Figure 13). 21 = 16, 32. Roadsides, waste
places, fields. From Eurasia, northern Africa. [AZ LUPULINA var. GLANDULOSA
Neilreich]

MEDICAGO MINIMA (Linnaeus) Bartalini—Bur Medick (Figure 13). 2n = 16. Wool waste, waste
places, fields, dry, open soil. From Eurasia, northern Africa. [M4 MINIMA var. COMPACTA
Neyraut; M@. MINIMA var. LONGISETA de Candolle]

MEDICAGO MONANTHA (C.A. Meyer) Trautvetter—(Figure 13). 2n =? Waste places. From south
central and southwestern Asia.

MEDICAGO POLYMORPHA Linnaeus—Toothed Medick (Figure 13). 2n = 14, 16. Wool waste,
waste places. From Eurasia, northern Africa. [M4. POLYMORPHA var. VULGARIS
superfluous name]

MEDICAGO PRAECOX de Candolle—Mediterranean Medick (Figure 13). 2n = 14, 16. Wool waste.
From northern and eastern Mediterranean.

MEDICAGO RIGIDULA (Linnaeus) Allioni—Tifton Medick (Figure 13). 2n = 14, 16. Wool waste.
From Eurasia, northern Africa. [Af AGRESTIS Tenore]

MEDICAGO SATIVA Linnaeus subsp. SATIVA—Alfalfa (Figure 13). 21 = 16, 32. Roadsides, fields,
waste places, meadows. From southwestern Asia.

ee SATIVA Linnaeus subsp. FALCATA (Linnaeus) Arcangeli—Yellow Alfalfa (Figure 13).
16, 32. Roadsides, waste places, wool waste. From Eurasia, northern Africa.
[MEDICAGO FALCATA Linnaeus]

MEDICAGO SATIVA Linnaeus subsp. VARIA (Martyn) Arcangeli—Bastard Medick (Figure 14). 21 =
? Roadsides, fields, waste places. From Eurasia. [M@. SATIVA subsp. SATIVA * M. SATIVA
subsp. FALCATA (Linnaeus) Arcangeli; Mf. SYLVESTRIS Fries]

MELILOTUS ALBUS Medikus—White Sweet-clover (Figure 14). 27 = 16. Roadsides, waste places,
fields, rich woods. From farther Eurasia.

MELILOTUS INDICUS (Linnaeus) Allioni—(Figure 14). 27 = 16, 18. Waste places, roadsides, fields,
riverbanks. From Eurasia, northern Africa.

Angelo and Boufford: Atlas of New England flora. 1]

MELILOTUS OFFICINALIS (Linnaeus) Lamarck—Yellow Sweet-clover (Figure 14). 21 = 16.
Roadsides, waste places, fields, railroads. From Eurasia.

ORNITHOPUS SATIVUS Brotero subsp. SATIVUS—Serradella (Figure 14). 2n = 14. Sandy, open,
disturbed soil. From southwestern Europe, northern Africa.

Oxytropis campestris (Linnaeus) de Candolle var. johannensis Fernald—(Figure 14). 2n = 32, 48.
Rocky or gravelly river shores. [O. johannensis (Fernald) Fernald]

PHASEOLUS COCCINEUS Linnaeus—Scarlet Runner (Figure 14). 2n = 22. Roadsides, waste
places. From Mexico, Central America. [P. MULTIFLORUS Lamarck]

Phaseolus polystachios (Linnaeus) Britton, Sterns & Poggenburg var. polystachios—Wild Bean
(Figure 14). 2m = 22. Rocky woods, dry, wooded ledges, roadsides. [P. polystachios var.
aquilonius Fernald]

PHASEOLUS VULGARIS Linnaeus—Common Bean (Figure 14). 27 = 22. Waste places, roadsides.
From Mexico, Central and South America. [P. VULGARIS var. HUMILIS Alefeld]

PISUM SATIVUM Linnaeus var. SATITVUM—Garden Pea (Figure 15). 27 = 14. Roadsides, waste
places, fields, railroads. From Eurasia, northern Africa.

PUERARIA MONTANA (Loureiro) Merrill var. LOBATA (Willdenow) Maesen & S. M. Almeida ex
Sanjappa & Predeep—Kudzu (Figure 15). 2n = 22, 24. Roadsides, waste places. From
eastern Asia, southwestern Pacific. [P. LOBATA (Willdenow) Ohwi]

ROBINIA HISPIDA Linnaeus var. HISPIDA—Bristly Locust (Figure 15). 2n = 30. Roadsides, fields,
sandy hills, rocky woodland margins, waste places. From farther south.

ROBINIA HISPIDA Linnaeus var. FERTILIS (Ashe) R.T. Clausen—(Figure 15). 2 = 20. Roadsides.
From farther south.

ROBINIA PSEUDOACACIA Linnaeus—Black Locust (Figure 15). 2n = 20. Roadsides, fields,
thickets, woodland margins, waste places. From farther west and south.

ROBINIA VISCOSA Ventenat var. VISCOSA—Clammy Locust (Figure 15). 21 = 20. Roadsides,
fields, waste places, thickets, often in sandy soil. From farther south.

ROBINIA VISCOSA Ventenat var. HARTWIGI (Koehne) Ashe—(Figure 15). 2n = ? Roadsides.
From farther south. [R. HARTWIGH Koehne]

—Robinia hybrid—

ROBINIA x MARGARETTA Ashe (pro species)—(Figure 15). [R. HISPIDA Linnaeus var. HISPIDA
x R. PSEUDOACACIA Linnaeus]

SCORPIURUS MURICATUS Linnaeus—Caterpillar-plant (Figure 15). 2n = 28. Wool waste. From
the Mediterranean.

Angelo and Boufford: Atlas of New England flora 12

SECURIGERA VARIA (Linnaeus) Lassen—Crown-vetch (Figure 16). 27 = 16, 24. Roadsides, fields,
waste places. From Eurasia. [CORONILLA VARIA Linnaeus]

Senna hebecarpa (Fernald) H.S. Irwin & Barneby—Wild Senna (Figure 16). 2n = ? Streambanks,
thickets, roadsides, fields. [C. hebecarpa Fernald]

SENNA OBTUSIFOLIA (Linnaeus) H.S. Irwin & Barneby—Java-bean (Figure 16). 2n = 28. Waste
places. From farther south.

SESBANIA HERBACEA (Miller) McVaugh—(Figure 16). 2n = 12. Waste places. From farther
south. [S. EXALTATA (Rafinesque) Cory]

Strophostyles helvola (Linnaeus) Elliott—Trailing Wild Bean (Figure 16). 27 = 22. Sandy or gravelly
soil, usually near coast, shores, marsh margins, railroads, fields, roadsides.

STROPHOSTYLES LEIOSPERMA (Torrey & A. Gray) Piper—(Figure 16). 2n =? Old gravel or sand
pits. From farther west.

Strophostyles umbellata (Muhlenberg ex Willdenow) Britton—Perennial Wild Bean (Figure 16). 21 =
22. Beaches.

Tephrosia virginiana (Linnaeus) Persoon—Goat’s-rue (Figure 16). 21 = 22. Dry, sandy, clearings
and open woods, ledges, fields, roadsides, railroads.

THERMOPSIS VILLOSA (Walter) Fernald & B.G. Schubert—(Figure 16). 2n = ? Roadsides, fields.
From farther south.

TRIFOLIUM ARVENSE Linnaeus—Rabbit’s-foot Clover (Figure 17). 2n = 14. Dry roadsides, fields,
waste places. From Eurasia, northern Africa.

TRIFOLIUM AUREUM Pollich—Yellow Clover (Figure 17). 21 = 14, 16. Roadsides, dry fields,
waste places. From Eurasia. [7. AGRARIUM Linnaeus]

TRIFOLIUM CAMPESTRE Schreber—Low Hop Clover (Figure 17). 2n = 14, 16. Roadsides, dry
fields, waste places. From Eurasia, northern Africa. [7. PROCUMBENS Linnaeus]

TRIFOLIUM DALMATICUM Visiani—(Figure 17). 2n = 10. Waste places. From southeastern
urope.

TRIFOLIUM DICHOTOMUM Hooker & Arnott—(Figure 17). 2n = ? Wool waste. From farther
west.

TRIFOLIUM DUBIUM Sibthorp—Small Hop Clover (Figure 17). 2n = 16, 28, 32. Dry roadsides and
fields, waste places. From Eurasia, northern Africa.

TRIFOLIUM ECHINATUM M. Bieberstein—Hedgehog Clover (Figure 17). 27 = 16. Wool waste.
From Eurasia, northern Africa.

Angelo and Boufford: Atlas of New England flora 13

TRIFOLIUM HYBRIDUM Linnaeus—Alsike Clover (Figure 17). 21 = 16, 32. Roadsides, fields,
meadows, clearings. From Eurasia, northern Africa. [7. HYBRIDUM var. ELEGANS (Savi)
Boissier]

TRIFOLIUM INCARNATUM Linnaeus—Crimson Clover (Figure 17). 2n = 14. Fields, waste places,
roadsides. From Eurasia. [7. INCARNATUM var. ELATIUS Gibelli & Belli]

TRIFOLIUM MEDIUM Linnaeus—Zigzag Clover (Figure 18). 2n = 48, 80. Oak woods, fields,
roadsides, hillsides. From Eurasia.

TRIFOLIUM OLIVACEUM Greene—(Figure 18). 2n =? Wool waste. From farther west.

TRIFOLIUM PRATENSE Linnaeus—Red Clover (Figure 18). 2n = 14. Fields, meadows, waste
places, roadsides, clearings. From Eurasia, northern Africa. [7. PRATENSE var. SATIVUM
Crépin]

TRIFOLIUM PURPUREUM Loiseleur-Deslongchamps—Purple Clover (Figure 18). 27 = 14. Wool
waste. From Eurasia, northern Africa.

TRIFOLIUM REPENS Linnaeus—White Clover (Figure 18). 2n = 16, 28, 30, 32. Meadows, waste
places, fields, roadsides. From Eurasia, northern Africa.

TRIFOLIUM RESUPINATUM Linnaeus—Persian Clover (Figure 18). 2n = 16. Waste areas. From
Eurasia, northern Africa.

TRIFOLIUM STRIATUM Linnaeus—Knotted Clover (Figure 18). 27 = 14, 16. Sandy fields. From
Eurasia, northern Africa.

TRIFOLIUM SUBTERRANEUM Linnaeus—(Figure 18). 2n = 16. Wool waste. From Eurasia,
northern Africa.

TRIFOLIUM TOMENTOSUM Linnaeus—(Figure 18). 27 = 16. Wool waste. From Eurasia, northern
Africa.

TRIGONELLA CORNICULATA (Linnaeus) Linnaeus—(Figure 19). 2n = 16. Ballast waste. From
Eurasia.

ULEX EUROPAEUS Linnaeus—Common Gorse (Figure 19). 2n = 32, 64, 96. Rocky headland turf,
dry slopes, pine woods, usually in sandy soil. From Europe.

ULEX MINOR Roth—Dwarf Gorse (Figure 19), 27 = 32. Sand plains. From Europe. [U. NANUS
TF. Forster ex Symons]

VICIA CRACCA Linnaeus—Cow Vetch (Figure 19). 27 = 12, 14, 21, 22, 24, 28. Fields, meadows,
roadsides, waste areas, thickets, shores. From Eurasia.

VICIA ERVILIA (Linnaeus) Willdenow—Bitter Vetch (Figure 19). 2n = 14. Ballast waste. From
Eurasia, northern Africa.

Angelo and Boufford: Atlas of New England flora 14.

VICIA FABA Linnaeus—Broad Bean (Figure 19). 2n = 12, 14. Fields, waste places. Origin unknown
(long in cultivation).

VICIA GRANDIFLORA Scopoli—(Figure 19), 27 = 14, Fields. From Eurasia.

VICIA HIRSUTA (Linnaeus) Gray—Tiny Vetch (Figure 19). 2n = 12, 14. Waste places, roadsides,
fields. From Eurasia, Africa.

VICIA LATHYROIDES Linnaeus—Spring Vetch (Figure 19). 27 = 10, 12. Roadsides, fields, usually
sandy. From Eurasia, northern Africa.

VICIA PANNONICA Crantz—Hungarian Vetch (Figure 20). 2n = 12, 14. Roadsides. From Eurasia.

VICIA SATIVA Linnaeus var. SATIVA—Common Vetch (Figure 20). 2n = 10, 12. Fields, roadsides,
waste places. Origin unknown.

VICIA SATIVA Linnaeus subsp. NJGRA (Linnaeus) Ehrhart—Smaller Common Vetch (Figure 20). 2
= 12, 14. Waste places, railroads, roadsides, fields, meadows, sea beaches, shores. From
Europe, Asia, Africa. [/& ANGUSTIFOLIA Linnaeus var. ANGUSTIFOLIA; V.
ANGUSTIFOLIA var. SEGETALIS (Thuillier) W.D.J. Koch; ANGUSTIFOLIA var.
UNCINATA (Desvaux) Rouy]

VICIA SEPIUM Linnaeus var. SEPIUM—Bush Vetch (Figure 20). 2 = 12, 14. Roadsides, railroads,
fields. From Europe, Asia.

VICIA TETRASPERMA (Linnaeus) Schreber—Sparrow Vetch (Figure 20). 2n = 14. Fields, railroads,
roadsides, waste places. From Europe, Asia, northern Africa. [V. TETRASPERMA var.
TENUISSIMA (Druce) Druce]

VICIA VILLOSA Roth var. VILLOSA— Winter Vetch (Figure 20). 27 = 14, 28. Fields, roadsides,
waste places. From Eurasia, northern Africa.

VICIA VILLOSA Roth var. GLABRESCENS W.D.J. Koch—(Figure 20). 2 = 14, 28. Fields,
roadsides, waste places. From Eurasia, northern Africa. [V/. VILLOSA subsp. VARIA (Host)
Corbier; % DASCYCARPA Tenore]

WISTERIA FLORIBUNDA (Willdenow) de Candolle—Japanese Wisteria (Figure 20). 2n = 16.
Woodland margins, fields, thickets. From Japan, Korea.

WISTERIA FRUTESCENS (Linnaeus) Poiret—American Wisteria (Figure 20). 27 = 16. Woodland
margins, roadsides. From farther south. [W”. MACROSTACHYA (Torrey & A. Gray) Nuttall
ex B.L. Robinson & Fernald]

WISTERIA SINENSIS (Sims) de Candolle—Chinese Wisteria (Figure 21). 2n = 16, 32. Woods,
woodland margins, roadsides. From China.

ACKNOWLEDGMENTS
We thank the curators and directors of the herbaria of the New England Botanical Club, the
Harvard University Herbaria, the University of Massachusetts, and the University of Vermont for

Angelo and Boufford: Atlas of New England flora 15

allowing access to their collections. For the University of Maine and University of Connecticut
herbaria we used their exceptional online databases of specimens. We are grateful also to Karen
Searcy for facilitating access to the herbarium and to the notebooks of Harry E. Ahles at the University
of Massachusetts (Amherst) and for kindly answering a request for information after our visit. James
Hinds generously checked information on voucher specimens at the University of Maine (Orono). The
following persons also checked certain records for us at their respective institutions: Janet Sullivan,
Robert Capers, Patrick Sweeney, and Craig D. Layne. John T. Kartesz and Misako Nishino
generously provided the latest draft version of the Floristic Synthesis of North America, which was
consulted for reports of occurrence and the sources of such reports. Erika Sonder kindly assisted with
reviewing specimens at the Harvard University Herbaria and the University of Vermont. We thank
Kanchi Gandhi for nomenclatural advice.

LITERATURE CITED & CONSULTED
(general references listed in our previous articles are not repeated here)

Angelo, R. 1994. A computer method for producing dot distribution maps. Rhodora 96: 190-194.

Angelo, R. and D.E. Boufford. 1996. Atlas of the flora of New England: Pteridophytes and
gymnosperms. Rhodora 98: 1-79.

Angelo, R. and D.E. Boufford. 1998. Atlas of the flora of New England: Poaceae. Rhodora 100:
101-233.

Angelo, R. and D.E. Boufford. 2000. Atlas of the flora of New England: Monocots except Poaceae
and Cyperaceae. Rhodora 102: 1-119.

Angelo, R. and D.E. Boufford. 2007. Atlas of the flora of New England: Cyperaceae. Rhodora 109:
237-360.

Angelo, R. and D.E. Boufford. 2010. Atlas of the flora of New England: Magnoliidae &
Hamamelidae. Rhodora 112: 244-326.

Angelo, R. and D.E. Boufford. 2011a. Atlas of the flora of New England: Salicaceae to Brassicaceae.
Phytoneuron 2011-12: 1-12 + figs. 1-53 (maps).

Angelo, R. and D.E. Boufford. 2011b. Atlas of the flora of New England: Paeoniaceae to Ericaceae.
Phytoneuron 2011-53: 1-13 + figs. 1-18 (maps).

Angelo, R. and D.E. Boufford. 2011c. Atlas of the flora of New England: Pteridophytes and

Gymnosperms: Introduction. <http://neatlas.org/Intro-Pterid&Gym.htm|>.

Angelo, R. and D.E. Boufford. 2012a. Atlas of the flora of New England: Caryophyllidae. Rhodora
113: 419-513.

Angelo, R. and D.E. Boufford. 2012b. my of the flora of New England: Ast Phyt

012-34: 1-39 + figs. 1-56 (ma

Angelo, R. and D.E. Boufford. 2012c. Pails of the flora of New England: Rosaceae. Phytoneuron
2012-81: 1-25 + figs. 1-31 (maps).

Fernald, M.L. 1950. Gray's Manual of Botany (ed. 8). American Book Company, New York.

Gleason, H.A. and A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and
Adjacent Canada 2nd ed. The New York Botanical Garden, Bronx, New York.

Goldblatt, P. and D.E. Johnson (eds.). 1979-. Index to Plant Chromosome Numbers (IPCN).
Missouri Botanical Garden, St. Louis. <http://www.tropico: [Py /IPCN>.

Reveal, JL. and F.R. Barrie. 1991. On the identity of Hedysarum violaceum Linnaeus (Fabaceae).
Phytologia 71: 456-461.

Seymour, F.C. 1982. The Flora of New England, 2nd ed. Phytologia Mem. V. Plainfield, New
Jersey.

Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website, version 9, June 2008 [and more or less
continuously updated since]. <http://www.mobot.org/MOBOT /research/APweb/>

HAMPSHIRE cee

VERMONT

MT. DESERT ISL.

SAGADAHOC

MAINE

° NANTUCKET
BLOCKISL.

CONNECTICUT RHODE ISLAND
Figure 1. Key map for counties of the New England states (and Mt. Desert
Island, Maine; Block Island, Rhode Island; arbitrary divisions of larger Maine
counties and of Coés County, New Hampshire).

ACMISPON AMERICANUS ALBIZIA JULIBRISSIN AMORPHA FRUTICOSA
var. AMERICANUS

Amphicarpaea bracteata ANTHYLLIS VULNERARIA
var. bracteata var. comosa

ARACHIS HYPOGAEA

Figure 2. Distribution maps.

BAPTISIA AUSTRALIS BAPTISIA LEUCOPHAEA
var. AUSTRALIS

Figure 3. Distribution maps.

Chamaecrista fasciculata
var. fasciculata

CLADRASTIS KENTUKEA COLUTEA ARBORESCENS CORONILEA SCORPIOIDES

Figure 4. Distribution maps.

Crotalaria sagittalis CYTISUS SCOPARIUS DALEA CANDIDA
var. sagittalis var. SCOPARIUS var. CANDIDA

DALEA LEPORINA DALEA PURPUREA
var. PURPUREA

Desmodium cuspidatum
var. cuspidatum

Figure 5. Distribution maps.

Desmodium glabellum Desmodium marilandicum Desmodium obtusum

Desmodium perplexum Desmodium rotundifolium

var. paniculatum

Desmodium sessilifolium Desmodium X humifusum GALEGA OFFICINALIS

Figure 6. Distribution maps.

GENISTA TINCTORIA GLEDITSIA TRIACANTHOS GLYCINE MAX

GLYCYRRHIZA LEPIDOTA GYMNOCIA DUS DIOICUS

Hylodesmum glutinosum Hylodesmum nudiflorum KUMMEROWIA STRIATA

Figure 7. Distribution maps.

LABURNUM ANAGYROIDES LABURNUM X WATERERI

var. pellitus

LATHYRUS ODORATUS Lathyrus palustris LATHYRUS PRATENSIS

Figure 8. Distribution maps.

LATHYRUS SYLVESTRIS LATHYRUS TUBEROSUS LENS CULINARIS

LESPEDEZA CUNEATA LESPEDEZA CYRTOBOTRYA

Figure 9. Distribution maps.

Figure 10. Distribution maps.

Lespedeza hirta subsp. hirta Lespedeza hirta subsp. hirta
XL. stuevei . violacea

XL. procumbens XL. viol

Lespedeza hirta subsp. hirta Lespedeza X longifolia
X L. virginic

Figure 11. Distribution maps.

LUPINUS POLYPHYLLUS MEDICAGO ARABICA
subsp. POLYPHYLLUS

Figure 12. Distribution maps.

MEDICAGO LACINIATA MEDICAGO LUPULINA MEDICAGO MINIMA

MEDICAGO MONANTHA MEDICAGO POLYMORPHA MEDICAGO PRAECOX

MEDICAGO RIGIDULA MEDICAGO SATIVA MEDICAGO SATIVA
subsp. SATIVA subsp. FALCATA

Figure 13. Distribution maps.

MEDICAGO SATIVA MELILOTUS ALBUS MELILOTUS INDICUS
subsp. VARIA

MELILOTUS OFFICINALIS ORNITHOPUS SATIVUS Oxytropis campestris
subsp. SATIVUS var. johannensis

PHASEOLUS COCCINEUS Phaseolus polystachios PHASEOLUS VULGARIS
var. polystachios

Figure 14. Distribution maps.

PUERARIA MONTANA ROBINIA HISPIDA
var. SATIVUM var. LOBATA var. HISPEDA

ROBINIA HISPIDA

var. FERTILIS var. VISCOSA

ROBINIA VISCOSA ROBINIA X MARGARETTA SCORPIURUS MURICATUS
var. HARTWIGH

Figure 15. Distribution maps.

SESBANIA HERBACEA Strophostyles helvola STROPHOSTYLES LEIOSPERMA

Strophostyles umbellata THERMOPSIS VILLOSA

Figure 16. Distribution maps.

TRIFOLIUM ARVENSE TRIFOLIUM AUREUM TRIFOLIUM CAMPESTRE

TRIFOLIUM DALMATICUM TRIFOLIUM DICHOTOMUM TRIFOLIUM DUBIUM

TRIFOLIUM ECHINATUM TRIFOLIUM HYBRIDUM TRIFOLIUM INCARNATUM

Figure 17. Distribution maps.

TRIFOLIUM MEDIUM TRIFOLIUM OLIVACEUM TRIFOLIUM PRATENSE

TRIFOLIUM PURPUREUM TRIFOLIUM REPENS TRIFOLIUM RESUPINATUM

TRIFOLIUM STRIATUM TRIFOLIUM SUBTERRANEUM TRIFOLIUM TOMENTOSUM

Figure 18. Distribution maps.

TRIGONELLA CORNICULATA ULEX EUROPAEUS ULEX MINOR

VICIA GRANDIFLORA VICIA HIRSUTA VICIA LATHYROIDES

Figure 19. Distribution maps.

VICIA SATIVA
var. SATIVA subsp. NIGRA

VICIA SEPIUM VICIA TETRASPERMA
var. SEPIUM var. VILLOSA

VICIA VILLOSA WISTERIA FLORIBUNDA WISTERIA FRUTESCENS
var. GLABRESCENS

Figure 20. Distribution maps.

WISTERIA SINENSIS

Figure 21. Distribution map.