LULUSTRATED FLORA ILLUSTRATED FLORA OF THE PACIFIC STATES WASHINGTON, OREGON, AND CALIFORNIA IN FOUR VOLUMES VOL. I OPHIOGLOSSACEAE TO ARISTOLOCHIACEAE FERNS TO BIRTHWORTS LEROY ABRAMS VOL. it POLYGONACEAE TO KRAMERIACEAE BUCKWHEATS TO KRAMERIAS LEROY ABRAMS VOL. Ill GERANIACEAE TO SCROPHULARIACEAE GERANIUMS TO FIGWORTS LEROY ABRAMS VOL. IV BIGNONIACEAE TO COMPOSITAE BIGNONIAS TO SUNFLOWERS ROXANA STINCHFIELD FERRIS ILLUSTRATED FLORA OF THE PACIFIC STATES WASHINGTON, OREGON, AND CALIFORNIA BY LEROY ABRAMS AND ROXANA STINCHFIELD FERRIS IN FOUR VOLUMES MOL. LY. BIGNONIACEAE TO COMPOSITAE BIGNONIAS TO SUNFLOWERS BY ROXANA STINCHFIELD FERRIS STANFORD UNIVERSITY PRESS STANFORD, CALIFORNIA SHSGOURI SOTANICAL GARDEN LIBRAGY STANFORD UNIVERSITY PRESS STANFORD, CALIFORNIA Lonpon : OxFrorp UNIversIty Press © 1960 By THE BOARD OF TRUSTEES OF THE LELAND STANFORD JUNIOR UNIVERSITY PRINTED AND BOUND IN THE UNITED STATES OF AMERICA BY STANFORD UNIVERSITY PRESS Library of Congress Catalog Card Number: 23-9934 PREFACE On completion of this final volume of the [/lustrated Flora of the Pacific States, there are many to whom I wish to express my sincere thanks. I am grateful to those who have contributed the text of genera and of families. Their willingness to pre- pare this material has shortened by years the time necessary to complete this volume for publication. My thanks are also given to the many others who have helped me with advice, material, and encouragement. I wish especially to thank Sidney Fay Blake, though his name does not appear as author of any one group of the Compositae. Dr. Blake had originally planned to contribute the text of the Compositae but due to the pressure of other tasks was unable to do so. He generously made available to Dr. Abrams and later to me all the ex- tensive notes and unfinished manuscript that he had assembled toward completion of that project. This material has been most helpful not only to me but to other contributors and much material included is based on his research. I wish to thank the following botanists for their contributions: Richard William Holm for the fam- ily Valerianaceae; Charles Bixler Heiser, Jr., for the genus Helianthus; David Daniels Keck for the genus Eastwoodia, the subtribe Madiinae, and the yellow-rayed genera of the tribe Astereae; Arthur John Cronquist for the genus Erigeron and the tribe Senecioneae ; Malcolm Anthony Nobs for the genus Achillaea; George Henry Ward for the genus Artemisia; Carl William Sharsmith for the genus Antennaria ; John Thomas Howell for the tribes Cynareae and Arctotideae ; Kenton Lee Cham- bers for the genera Nothocalais and Microseris; and Quentin Jones for the genus Agoseris. Finally, I wish to express respect for and appreciation of LeRoy Abrams, to whom I owe my botanical training and who, long before his death in 1956, asked me to take the responsibility of preparing for publication this final volume of the Illustrated Flora of the Pacific States. The illustrations in the book are for the most part the work of Jeanne Russell Janish, though many of the drawings of the Compositae are made by Doris Holmes Blake. The drawings of Helianthus and Agoseris are made by other illustrators under the direction of the respective authors of the text of those genera. Elisabeth F. Allen has ably assisted in reading proof. Barbara W. Law deserves especial credit for the arduous task of assembling the comprehensive index for all four volumes, in addition to the necessary editorial work of checking references. Following the format of the preceding volumes, the description following each species heading, except in rare cases, refers to the name-bearing subspecies or variety where such segregations exist in literature. Deviations from this form are indi- cated in the text. The illustrations are reduced ,to one-half natural size except, of course, for the structural details. Any exception to this reduction is marked on the illustration to which it applies. Keys to the families of all four volumes are given in the Appendix, followed by a complete index of all scientific and common names that appear in the flora. Roxana S. FErrIs Stanford University November 1959 ' ig j Sy 3 ys ‘ ‘ a t es [> §=*3 © £ = uM 7 5 eS - ; ag V4 2 4 ag ~s LE | £ ates er m - 4 ‘ hoe ea? ‘ i + - ‘ ss « ' F i ; Ms fr ERK BESS yee pe as “ess aad, sorta fer “We (eerat Wein tie 137. 138. 139. 140. 141. 142. CONTENTS OF VOLUME IV PREFACE Vv APPENDIX 615 INDEX OF COMMON NAMES 625 INDEX OF SCIENTIFIC NAMES 653 Sympetalae (concluded) 1-613 . BIGNONIACEAE 1 143. RUBIACEAE . MARTY NIACEAE 1 144. CAPRIFOLIACEAE . OROBANCHACEAE 3 145. VALERIANACEAE . LENTIBULARIACEAE 10 146. DipsAcACEAE . ACANTHACEAE 13 147. CucURBITACEAE . PLANTAGINACEAE IS 148. CAMPANULACEAE 149. ComposITAE 98 COMMON NAMES Sympetalous Plants (concluded) BIGNONIA FAMILY 1 143. MAppER FAMILY UNICORN-PLANT FAMILY 1 144. HoNEYSUCKLE FAMILY BROOMRAPE FAMILY 3 145. VALERIAN FAMILY BLADDERWORT FAMILY 10 146. TEASEL FAMILY ACANTHUS FAMILY is 147. Gourp FAMILY PLANTAIN FAMILY is 148. BLUEBELL FAMILY 149. SUNFLOWER FAMILY 98 ap i : a : pe § j i ee . ‘3 Pia Et eli as..* nee ge a gar arses in ne witty 3 . f Ee 5 meas cae a ate ILLUSTRATED FLORA VOL. IV Family 137. BIGNONIACEAE. BIGNONIA FAMILY. Trees, shrubs, or woody vines, or some exotic species herbs. Leaves opposite or rarely alternate, epee compound or simple, estipulate. Flowers showy, terminal or axillary, u clustered. Corolla campanulate, funnelform, or tubular, 5-lobed lobed stigma. Fruit a 2-valved, sometimes woody, capsule. Seeds numerous, flat, winged, vou endyspetits sole ns broad, flat, entire or 2-lobed ; radicle straight. A family of about 100 genera and over 500 species, mainly tropical but a few in the warm temperate zones of both the es a sonthern Keuiophietes. 1. CHILOPSIS D. Don, Edinb. Phil. Journ. 9: 261. 1823. Shrub with simple, usually alternate leaves. Flowers in terminal solitary racemes. Calyx inflated, deeply 2-lipped, upper lip 3-toothed, lower 2-toothed. Corolla funn . 5- lobed and obscurely 2-lipped. Anther-bearing stamens 4, the fifth represented by a rudimentary filament. Capsule linear and terete, somewhat woody. Seeds flat, with their wieie dissected into long hairs. [Greek, meaning lip and resemblance A monotypic genus of the arid southwestern United States and eaice: ‘i eae ae (Cav.) Sweet. Desert Willow. Fig. 4945. Bignonia ? linearis Cav. Ic. 3: 35. pl. 2 796. Chilopsis saligna D. ci ae its Phil. Journ 9: 26L.. 1823. Chilopsis linearis Sweet, Hort. Brit. 2 1827. Chilopsis linearis var. arcuata Pots bis dane 3: 366. 1936. A willow-like shrub, a — several stems, 2-6 m. high. Leaves narrowly linear to linear- lanceokate, 5-15 cm. long, 2- wide, long-attenuate at both ends, [a green, glabrous, and similar on ea h idee: racemes ea bps many-flowered ; calyx mm, long, pubescent corolla ong, fragrant, white with pink lines or pink; capsule firm, 15-27 cm. 4-5 m in diameter, tapering at ds. Frequent along eee Seni Lower Sonoran Zone; d Col Deserts, California, and djacent Nevada and Ariz #3 to western Texas and south to Sp er California re the mainland of Mexico. ste ocalier: not stated. May -f ug. Sometimes cultivated as an ornamental. Family 138. MARTYNIACEAE. UNICORN-PLANT FAMILY. Herbs with mostly opposite estipulate leaves and perfect irregular flowers. Calyx 4-5-cleft or 4-5-parted, or sometimes divided to the base on the lower side and spatha- ceous. Corolla sympetalous, irregular, the tube oblique, often decurved, the limb 5- lobed and heey 2-lipped, the lobes nearly equal, upper 2 exterior in the bud. An- ther-bearing stamens 2 or 4, didynamous or the posterior pair sometimes sterile ; anthers 2-celled, fongitudinally dehiscent. Ovary superior, 1-celled, with 2 parietal placentae expanded into broad surfaces, or appearing 2—4-celled by the intrusion of the placentae or by false partitions ; ovules numerous or sometimes few, anatropous ; style slender ; stigma 2-lobed or 2-lamellate. Fruit various. Seeds often compressed : endosperm none ; cotyledons large. A family of 5 genera and about 15 species, natives of temperate and tropical regions of the western hemi- sphere. 1 F. MARTYNIACEAE Calyx of 5 free sepals; endocarp of the fruit strongly echina 1. Ibicella. Calyx 4—-5-dentate but split to the base on the lower side iar more or less ssaamminicaa 2. Proboscidea. 1. IBICELLA Van Eseltine, N.Y. State Agr. Exp. Sta. Tech. Bull. No. 149: 31. 1929 Stout, viscid-pubescent annuals and strongly scented. Leaves opposite or the upper sometimes alternate, ovate to suborbiculate. Inflorescence a dense terminal raceme. Calyx the rudiment of the fifth stamen often present. Capsule 2-valve ith a curved beak, crested above, the exocarp rather fleshy, the endocarp densely echinate Seeds compressed, rugose. [Name from the Latin, meaning seed ae of the curved horns on the fruit. ] A genus of 2 species, natives of South America. Type species, Martynia lutea ry ndl. 1. Ibicella lutea pees Van Eseltine. Yellow Unicorn-plant. Fig. 4946. Martynia lutea Lindl. Bot. Reg. 11: pl. 9 Proboscidea lutea stad in Engler & Prantl Nat. Pilanzent. 48b; 269. 1895. Ibicella lutea Van Eseltine, N.Y. State Agr. Exp. Sta h. Bull. No. 149: 34. pl. 14. 1929. Gla iat dalbelinens amas with stout creating stems 3-6 dm. long. Leaves opposite or the upper sometimes alternate, suborbicular, mostly t 10 cm . broad, incised or subcordate at base, denticulate ; er grt short: -racemose, few an ara nd r r dense; sepals distinct, about .5 cm. long, the two lower much bro ader ; shai yellow pet ‘landul r on the outside, glabrous and orange or dee eile within and often dotted w h red, about 2.5 cm. long; stamens 4 with i obvious T tidshawies ry fifth : fruit-body oblon ng- eee crested abov e, about 5 cm. long; endocarp conspicuously echinate, with short ascending spines the horn abit ‘Ghee as ben as the body. Sparingly introduced in California and mostly found in 1 cultivated areas in Sacramento and northern San Joaquin Valleys and in the central coastal area. Type locality: Brazil. June—Sep 2. PROBOSCIDEA Keller in Schmidel, Icon. (ed. Keller) 49. 1762. Stout, annual or perennial herbs, glandular or viscid-pubescent and strongly scented. Leaves opposite or the upper sometimes alternate, long-petioled, and having broad blades. i d ‘ e also sometimes above, ending in a prominent incurved beak, becoming falsely 4-celled by sae extension of the placentae ; exocarp fleshy, separating in age from the woody sculptured ocarp. Seeds numerous, tuberculate. [From the Greek, meaning snout, in allusion to oot of fruit. ] A genus about 9 species, native of central and southwestern United States, Mexico, and South America. Type species, Martynia loutsianica Mill. 7 Ata ic gona and subcylindrical, often crested both dorsally and ventrally; flowers 2 or brownish; 3-5 cm. wide. . P. althaeifolia. BTSs hay stout, more or less ovoid, crested only on the dorsal boaged flowers reddish purple ., gets dotted with purple, the throat often striped with yellow; leaves about 10 cm. wide. . P. louisianica. 4945. Chilopsis linear 4946. Ibicella lutea BROOMRAPE FAMILY 3 1. Proboscidea altheaefdlia (Benth.) Decne. Desert Unicorn-plant. Fig. 4947. Martynia altheaefolia Benth. Bot. Sulph. 37. 1844. Pro nie idea altheaefolia Decne. Ann. Sci. Nat. V. 3: 324. 1865. Martynia palmeri S. Wats, Proc. Amer. Acad. 24: 66. 1889. Spreading, viscid- teats perennial 3-4 dm. high with a long, yellow, fusiform root. Leav: 3-5 cm. broad, reniform to broadly ates . ne often broadly and shallowly 3—-5-lobed ; solicies 3-8 cm. long, densely sade siibeas acemes several-flowered, often becoming cm. long; calyx 1-1.5 cm. long; corolla 2. pa 5 cm, long, buff to brownish ye ‘sag Koy of the fruit sle nder, 5-6 cm. long, horns two to three times as long, the distal teeth of the crest on the dorsal suture often prolonged into a slender horn, the ventral suture not at all or only slightly crested. Usually in sandy soils, Lower Sonoran Zone; Colorado Desert, California, and western Arizona parward into Sinaloa and southern Lower California. Rare iar our range and limited to Imperial County, Californ Type locality: Magdalena Bay, Lower California. ite hk 2. Proboscidea louisianica ( Mill.) Thell. Common Unicorn-plant. Fig. 4948. Proboscidea jussieut Keller in Schmidel, Icon. (ed. Keller) 49. p/s. 12-13. 1762. Not a binomial, Leaflets West. Bot. 1: 80. 1934. Martynia louisiana Mill. _ Dict. ed. 8. no. 3. 1768. Martynia louisianica Mill. cit. in corrigenda. Martynia proboscidea Gloxin, ‘Obs. 14. 1785. Martynia alternifolia Lam. Encycl. 2: 112. 1786. Proboscidea loutsianica Thell. Mem. Soc. oh ci. Cherbourg IV. 38: 480. 1912. Proboscidea louisiana Woot. & Standl. Contr. U.S. Nat. Herb. 19: 602. 1915. Martynia jussieut J. T. Howell, Leaflets Wont Bot. 1: 40. Annual with prostrate or ascending branches 3-10 dm. long, slimy-viscid pubescent throughout. Leaves opposite or Aa upper ee cone deeply cuales broadly ovate to suborbicular in outline, 5-20 cm. broad, entire or sinuate; petioles 5-15 cm. long, stout, aenael s pe tsonbenns racemes oe several flo san elongated in pane calyx 1.5-2 cm. long, the lobes acutish to obtuse; corolla 3.5-5 cm. long, yellow or dull white blotched with reddish purple or sometimes purple throughout ; body of = fruit 4-6 oe coke -10) cm. long, crested on the upper side only, the horns as long or three times as long as in low moist ground, yi onoran Zones; Sacramento Val and coastal valleys south to southern Cali- fornia, eastward to the southeastern United States, and psec into Mexico. Type locality: not definitely known; probably southeastern United States or eastern Mexico. May-—Sept. Devils-claws. Family 139. OROBAN CHACEAE. BROOMRAPE FAMILY. Root-parasites without green foliage, the stems erect, simple or i pi usually yellowish or purplish. Leaves reduced to alternate appressed scales. Flowers perfect, irregular, solitary, or pees or sessile in terminal bracteate ee or spikes. Calyx free from the ovary, 4—5-toothed or 4—5-cleft, or split nearly or quite to the base on hei or both sides Corolla sympetalous, more or less irregular, the limb 2-lipped and 5-lobed, usually with a pair of bracteoles. Stamens 4, didynamous, slender, in- serted on the corolla-tube, a rudimentary fifth one sometimes present ; filaments slen- der ; anthers 2-celled, the sacs parallel and equal. Ovary superior, 1-celled with 4 @) a8) n o. oD — ‘A @. a me) i] io) 5 + < eo merous, reticulated, wrinkled or striate ; embryo minute with the cotyledons scarcely differentiated. About 14 genera and over 200 species of wide geographical distribution. Capsule 2-valved with 2 ; filaments not with hairy tuft at base. 1. Orobanche. Capsule 4-valved with 1 placenta on each coma filaments with tuft of hairs at base. 2. Boschniakia, 1. OROBANCHE [Tourn.] L. Sp. Pl. 632. 1753. Glandu a gas herbs, yellowish, purplish, or rarely white, parasitic on the roots of various plants the ie — reduced to scattered scales. Flowers yellowish or purplish, solitary on oe ache cies more com monly spicate, racemose, or agp a e, with or poarch or bilobed. he eee Pek. she g ek and vetc About 100 species, native of North and South America and the Old World. Type species, Orobanche major L. OROBANCHACEAE 4947. Proboscidea altheaefolia 4948. Proboscidea lousianica Flowers on long slender pedicels, solitary or several on the stem, without floral bracts. Stems slender, very short, bearing 1 or emma eo - 3 pedicels many times exceeding the length of the stem; ealy eichea subulate-attenuate, longer than tube. . O. uniflora occidentalis. Stems stout, each bearing 3-12 pedicels Dine or Sake than the stem; ees Sraneulas | = lanceolate, ualin gor on rte hiss the tube. oO. gg a gi ben e, corymbose, or paniculate, bey flowers sessile or pedicellate; floral beaks ee pry in oi ramosa). Calyx not parted to cg base before ae she Sak folds obso inflorescence a compact, pyramidal, terres panicle of sessile flowers; base of ‘on se rs Sat thick ned. . O, bulbosa. Palatal a, present (small in O. aia. inflorescence various, the — pedicellate, or sessile in the spicate forms; base of stem not greatly ices qe (somewhat so in 0. corymbosa). Calyx 3 rake: or -divided, Cesrcaate * native s' x short, the lobes equaling or ‘. little "surpassing the tube; oar gc elec body a con- spicuous solid maar 15- 20 mm. in dia . O. pin Calyx bic gp pcg the Sepa tom usually en, surpassing the = basal gee Ar when vident, o tra Flowers pedicellate, Ma lower ee 8-25 mm, long, sometimes shorter above; inflores- = corymbose, subracemose, or paniculate. TT or compactly corymbose; anthers woolly. Tatorheccuce mostly open-corymbose; corolla-lips about 10-14 mm. long; lobes of the upper corolla-lip acute (rounded i in var. violacea). . grayana. Inflorescence compactly corymbose; corolla-lips about 7-8 mm. bas Oe Pg ae of the upper corolla-lips obtuse or rounded. 7. O. corymbos Inflorescence subracemose or paniculate; anthers glabrous or sparsely hairy. . californica chishateabel sessile or = i = lower ones short-pedicellate; lalliciNenses Saaecelalty eee the much elongated in age. s to twice the length of the tube. BOve ue ce especially those of the lower lip, narrowly acute; sales lobes one and bt rounded; calyx-lobes twice or more ~ length BS ‘the tube. Calyx 4-toothed; introduced specie - " ramosa. Calyx parted to the base before and behind, oe divisions 2-toothed. 11. O. minor 1. Orobanche uniflora subsp. occidentalis (Greene) Abrams ex Ferris. Naked Broomrape. Fig. 49 Aphyllon uniflorum var. sea Greene, Man. Bay — 285. 1894. Aphyllon sedi Suksd. Deu t. Monatss, 18: 155. Ap inte minutum Suksd. ae Orobanche un niflora f. sedi Bec, Paansenrcch — 48. 1930. greg k, op | Bull. oad Club 60: 446. ubsp. aca Abrams ex Ferris, Contr. se Herb, 5: 99. 7958. tems simple, 0.5-2 Orobanche uniflora s' cm. long, pee anean, bearing tee ovate-oblong ong es usu- ; 8 cm. , glandu ais ubescen stly 1-flow nulat eae h lobes iciesinie sabutate he tube; rele ite or tik purple to almost white, 12-22 mm ong, ger urved, throat 3-5 mm. aly e lobes broadly ovate, subequal, scarcely sprendicay 2-3.5 mm. ine: anthers glabrous or bose bescent. BROOMRAPE FAMILY 5 asitic on various plants, principally on pasinngncees a3 n Sedum, open woods and mossy banks, Transi- aed § ower Roe bwin Montana a and British Columbia ra mie ic ge my and east to the Rocky Moun- Hema so Peas “Wooded y hills,” Zac central California. Mar nche uniflora var. edn (Heller) Achey, Bull. Torrey ye er yi 1933. (Thalesia purpurea Heller, I Bull. Porras —_ 24: 313. 1897; Aphyllon inundatum Suksd. Allg. Bot. a a ~ 27. sl Reo yg eg uniflora £. inundata Beck, Pflanze — "4201; “49, 1930; O. porphyrantha Beck, t.) Stems sometimes branche d; Sato: usually 2 , 5-15(20) em. lon ng, rather stout, slandular aie cor olla = 2 a long, pale to deep visket, somewhat streaked with yellow, tube ms mewhat cu throat constricted below, about 6-8 mm. wide above; corolla-lobes ae or bel spreading; anthers gla vale or hairy. Parasitic on vario plants ochwicae the ‘Compositae, open rocky ground; Idaho pie British Columbia —_ through Washington ana Oregon to Nevada eno in neal Sierra Nevada os Santa Cruz County in the Coa t Ranges, California. Type locality: near Lewiston, Nez Per e County, Idaho nih aching th “ Pical anectan. hi is Teil we ostly in the eastern United States, in growth habit but differing by having Sane: attenuate caly ge instead "Of triangular-acute lobes that scarcely surpass the calyx- ike, : characteristic of the eastern speci 2. Orobanche fasciculata Nutt. Clustered Broomrape. Fig. 4950. Orobanche fasciculata Nutt. Gen. 2: 59. 1818. Loxanthes deg ie nts s Raf. Neogen, 3. 1825. Anoplanthus fascicu ial hig Rep. 3: 480. 1844-45. Aphylion fantail ray ex A. Gray, i ed. 2. 281. 1856. Thalesia fasciculata gts "Mem, Torrey Club 5: 1894 Stems fleshy, mostly subterranean ory rising 2-6 cm. above ground, simple or compound with several sett branches bearing prominent scales and several to many 1-flow tpn Lather Is, laselitia- ub u mm. bi 2-3 cm. long, the tube —— its hes usually 5-6 mm. long and usually iomndar capsule Siaudie ovoid, mostly 8-10 mm. lon Parasitic on the roots ts various Sen s, Transition Zone; British Bese eg to southern California east to Michigan. In the Pacific States it is found in the Olympic, Cascade e, and Blue Mpuntains of Wee and Oregon, and in the foo Ranges and. pers Nevada in California, Typ - locality : ros Mandan, North Dakota. May-— ~Aug. A variable species from which two intergrading varieties have been get ted. banche fasciculata var. Hiei gp hataee Achey, Bull. Torrey Club 60: 449, fos 12-13. 1933. hwo lutea Parry, Amer, Nat. 8: 214. 1874; Aphyllon Fi ee sor var. alg ve Gray, Syn. Fl. N. Amer. 21: 312. 1878; Orobanche fasciculata f. lutea Beck, Pflanzenreich 251 1930:) Calyaicbes 6 qualing or al th than the tube; oe sone and sometimes tinged with rag eee corolla-tube slightly constricted at ase, corolla-lobes usually acute. asitic on grasses and other hosts; Alberta, Montan na, and Idaho south i e Pacific States to northern California, east to North and South Dakota, gute south to Chihuahua esi Son sore. ss locality: Owl Creek, Wyoming. Oroban ae fasciculata var. franciscana Achey, Bull. Torrey Club 60: 450. figs. 14-15. 1933. Pedicels 4-12; calyx-lobes somewhat longer than the tube; corolla straw-colored tinged with purple, corolla-tube not constricted, corolla- ons nded or truncate, someti beac acute. Parasitic on various hosts; ee Oregon south to San Diego ate "California. Type locality: Mount Tamalpais, Marin County, California 3. Orobanche bulbésa Beck. Chaparral Broomrape. Fig. 4951. Focsusred lrgitastg Gray, Proc. Amer. Acad. i 2h 1868. A phyllon tub m A. Gray, Bot. Calif. i: nop Orobanche bu i Beck, Bibl. Bot. 4: 83. pi. 1, rae Orobanche tuberosa Heller, Cat. N. Amer. eo a ae oe Hook. Fl. Bor. Amer. 2: 92, 1838. tems fleshy, dark purplish brown, arising from a tuber-like attachment to hos 1-2 dm. high, the stems much enlarged and sometimes bulbous at base. Scales at the base ‘roars ovate, obtuse r ar to eolate, acute, spreading, puberu- in color from yellow or brownish to purple ar so, the lobes 2 mm. or more long, acute, 4949. Orobanche uniflora 4950. Orobanche fasciculata 6 OROBANCHACEAE Parasitic on various shrubs, usually in the dry geo ae Upper Sonoran Zone; Solano County in the Coa pi and El Dorado Cyaan d in the Sie “ faa ada o San Diego County, the southern California islands and northern Lower California. Type loc lee : hi — and dry ridge of the Gavilan Mountains, in sa colk, ” California, May-— ae 4. Orobanche pinoérum Geyer ex Hook. Pine Broomrape. Fig. 4952. Orobanche pinorum Geyer ex Pest pita Journ. Bot. 3: 297. 1851. Phelipaea Sree Gray, Proc. Amer. Acad. ‘fe he 1868. Aphyllon pinetorum A ay t. Ca lif, e 585. Myzorriza Aptian m Rydb. Bull. ae Club 36: ky "1909. Ste sing from a large, tuber-like ae iy! i host, the base of the stem not greatly en- larged, ter slender simple or branched above e middle i into a few erect branches, growin stems lavender to purple becoming yello wal minutely gs Ns ee pubescent and somewha t viscid. Scales of the stem eee late and mostly acute, 4-10 mm. long; flowers many, pedicellate, in a simple or branched raceme ; pedice s 2-8m a ng, Leas ascending ; ; floral bracts are: subula te, mm. long ; calyx 7-8 mm. long, ie tomentulose, lobes subulate, about equaling the nar- i plish —2 cm. long rowly anulate 10 - A the palatal folds small and obscure, tube constricted and curved outward at the base of the limb, obscurely 2-lipped, the lobes subequal, 3-4 mm. long, heme anthers glabrous or sparsely pubes- cent. n. ings alae forests, parasitic on the roots of various coni ifers and also reported from Holodiscus, mainly Sition cs x ca Siskiyou Mountains in Bagh evi ig eee County, and in Santa Cruz County, California, and eastwa rd to Idaho —_ the Blue Mountains, Oregon. Type locality: ‘‘Top of the pon mountains near St. Joseph, Coeur d’Aleine untry, growing on the sien of Abiee tate ge Idaho. June—Aug 5. Orobanche grayana Beck. Gray’s Broomrape. Fig. 4953. Orobanche comosa Hook. FI. Bor. Amer. 2: 92. * sere 1838. Not Wallr. 1822. Apes comosum A. wed Bot. ey - 1; ad Beck, Bibl. B ee M iaptehled hese — Bull. “Torrey ob re 695. Orobanche grayana var. nelsonit Munz, op. cit. 57: 616. pi. 38, fig. 7. 1930. Stem oe high, glandular puberulent, — below, the snag Dicet k wicaite branc hed. She scales lanceolate to ovate, 5-1 . lo ong ; a sieslee nce corym ory bose-paniculate, 4-8 cm. long, the veticeti 5-25 mm. oe floral bracts tinear-stteilate, ealty alternate on the seeclinats calyx 12-16 . long, the ated lobes gth; corolla purplish with darker lines, occasionaily white, . long, - tube rather slender and palatal folds evident ; elliptic, ‘obtuse at apex, narrowed at base to a short winged petiole, the margins Sidtaally inrolled ; calyx 3-5 mm. long, the lobes obtuse, the two lower more or Vices hes corolla violet- purple, is 20 m ek including the subulate spur, | 2-lipped, the upper lip 2- lobed, the lower 3- lobed, both cal i grid capsule ovoid, 6-8 mm. long, 5-6 mm. broad. Growing o ocks sandy or tins) feslbes' a circumpolar species, extending southward on the Pacific ‘Cot t ie ng Gemui —— Washington to the Siskiyou Mountains, Grenee’ and = River, olay Se” ba i Golitornia, and across the continent to Minnesota, Michigan, and New York. Type locality: 4961. eS ee 4962. Pinguicula vulgar: 2. UTRICULARIAL. Sp. Pl. 18. 1753. Aquatic or bog herbs with horizontal, submerged, branching stems. Leaves alternate, sometimes root-like, 2-8-parted at the very base and thus often appearing as if verticillate, the divisions dichotomously or pinnately finely dissected, at least some - them bladder-bear ing. Bladders with a pair of bristles about the mouth. Flowers in racemes or som otinies solitary, the scapes naked or with a few scales basally attached or sonnetimnes replaced by a whorl of inflated floats ; pedicels from the axils of bracts, these sometimes auriculate. Calyx 2-lobed, the lobes concave, persistent. Corolla strongly 2-lipped, the palate at the base of the bd lip prominent, often 2-lobed. Anthers not lobed. apsule few- to many-seeded. Seeds more or less peltate, flat-topped, and the tee sometimes winged. [Name Latin, iiedas a little bag. ] le LENTIBULARIACEAE ; a i 300 species of wide geographic distribution in tropical and temperate regions. Type species, Utricularia vulgaris L. Pedicels fae in fruit. Lea crowded, 3-4 cm. long; corolla-spur curved but little shorter than the lower lip U. vulgaris. «pele scattered, mostly 1-2 cm. long; corolla-spur very short or sometimes almost radimentay. a erect in os salable d; corolla-spur much shorter than lower lip. 3. U. gibba. Leaves 2-4- parted; coral: spur about equaling lower lip. rollas m. broad; ultimate leaf-divisions flat. 4. U. intermedia. sae ae mm. See ultimate leaf-divisions terete. 5. U. fibrosa 1. Utricularia hier 90 L. Greater or Common Bladderwort. Fig. 4963. Utricularia vulgarts L. 8. Utricularia vulgaris var. pili os Man. ed. 5. 318. 1867. Stems immersed, usually free- floating, rather stout and ene densely va hed ae 1-2 cm. long, ofte n lobed in outline Nee divisions fringed with g irs; —4c¢ ong, 2-3-pinnately divided into filiform segments, midrib n ot gacek useally very bladdery, the bladders bout 3-4 mm. lo scapes stout in co i (3 a cm ong, flowered ; pedicels reflexed in fruit ; corolla yellow, 10-20 mm. broad, the sides of the lips reflexed and the uppe ap ese entire, barely longer than the palate, lower lip “se slightly 3-lobed; spur m. long, somewhat curved, Shunk or acutish, a than the lower lip; capsule globose. nie and aa streams, Boreal, Transition, and so mes Upper “elo Zones; British Columbia to — California and eastward across the Cee Cnenk: zy i Pe ice and Asia. Type locality : Europe. April- 2. Utricularia minor L. Lesser Bladderwort. Fig. 4964. Utricularia minor L. Sp. Pl. 18. 1753. Stems floating or creeping on the bottom in shallow water, ed slender and thread- a 1-3 dm. aise sparingly branched. Winter buds round, 1. 5-5 mm. long, leaf-divisions not fringed ; leaves about 1 cm. long, 2-4-parted, the divisions very slender but Tete, entire, of two ee the larger 25-30 mm. long and usually witho ut _ Acers, the mallae 1-3 cm. long and aring bladders; scapes almost filiform, 5-15 cm owered, and with 2-5 minute eae ale scales; pedicels Se wah 2-8 mm. long, eek in aaits calyx-lobes 1 mm. long; corolla yellow, frog. ci lip 2-4 mm. long and about half as wide; lower | ip 4-8 mm. long, palate pee! spur very short and saccate imap about 1 mm. in dia In ee allo inly 1 Zone; circumpo ee in dstraton, extending southward in North America Toke — Ae Taare Core, ecrenk, ‘also Utah, Colorado, Ohio, and Pennsylvania. Type locality: Europe. ¥ mn Utri Sie aria occidentalis A. Gray, Proc. Am Acad. 7 95.. 1883. An “cab kigptoe d known species which differs fie U. minor by the hairy- [omied eat. pia of ‘oe kB buds. Kno only from Falcon Valley, Klickitat County, Washington, the type locality. 3. Utricularia gibba L. Swollen-spurred Bladderwort. Fig. 4965. Utricularia gibba L. Sp. Pl. 18. 1753 Utricularia fornicata Le Conte, Ann. isc WY. 1: 76... 1824. Stems slender, short, creeping over moss 0 mud in shallow water, ee ene. les 1 mm. or less in poacon the leaf- divisions thread- like, pe fringed on the margin once-parted, appearing r oot-like, a henegre few, h air-like, without midribs; bladders sat” cane on leaf- divisions ; scapes s 2:5-10 ¢ igh, with short slender branches ‘at se, 1-2-flowered, the ee erect in fruit; oo pia. pete m. broad, lips rounded, the lower reflexed and somewhat longe than the thic k, blun t, Bibbous anil gveiee globose n shallow water. Ges n Zon n the Pacific States occurring in California, in the San Joaquin Valley in Sen Tonquin Coun. —. the oe gf pothole Bay area in Lake, Sonoma, and Contra Costa — and = in the northern Sierra Nevada; e wide distribution in the eastern states and also Mexico and Central Amer Type locality: Virginia. July: —Sep 4. Utricularia intermédia Hayne. Flat-leaved or Mountain Bladderwort. Fig. 4966. Utricularia int dia Hayne in Schrad. Journ. fiir de Bot. 1: 18. 1800. Stems 1-2 dm. long, creeping on the bottom in shallow er and radiating Se the base of the scape. Winter buds oval, 3-10 mm. long; the leaf- divisions frin << * ith gray hairs; leaves numerous and crowded, -ranked, sii not flaccid, 4—5-forked, the segments A near, flat, bristly- serrulate, with evident midrib ; bladders 2-5 mm. lon rne on couaiae a branches or some- i fl ee = - — bra Ww ; n fru 5 ’ long and about 1 mm. oe the | shan ie ‘slightly 3-lobed, 10-12 mm. broad, palate prominent ; a conic at base, cylindric above, acute, a little shorter than the lower lip; capsule 3 mm. in eter. w water in mountain Fy and ponds, Boreal and Transition Zones; cooler portions of the northern hem Gobcte a en Pas co ~~ ' g a) s » bal &. =] %) co o © S u~ ~ a = GQ fe) he a pair of longitudinal ridges ; stamens about coualing the: up rocky slopes and washes, Lower Sonoran Zone; western aa of re Co lorado Desert, oe. east to stern Ari = Hi and south into Lower California and western Sonora. Type locality: Cape San Lucas, Lower conten Feb.—Jun 4967. Utricularia fibr 4968. Beloperone californica Family 142. PLANTAGINACEAE. PLANTAIN FAMILY. Annual or perennial, acaulescent or short-stemmed, rarely stoloniferous herbs. Leaves basal, or in caulescent species opposite or alternate, estipulate, the venation seemingly parallel. Flowers hypogynous, small, perfect, polygamous or monoecious, bracteolate, in dense, terminal, long-scaped spikes or heads, or rarely solitary. Calyx +-parted, persistent, the lobes imbricated pale persistent. Corolla 4-lobed, scarious or membranous, usually marcescent. Stamens 4 or sometimes 2, or in one South Ameri- can genus only 1, inserted on the tube or throat of the corolla; filaments filiform ; an- PLANTAIN FAMILY 15 A ‘Siantty of 3 genera and sical 250 species of wide see distribution. 1. PLANTAGO [Tourn.] L. Sp. Pl. 112. 1753. Acaulescent or rarely leafy-stemmed herbs, the scapes arising from the axils of the alternate leaves. Calyx-lobes equal or sometimes two of them arger. Corolla salverform, the tube cylindric or constricted at the throat, the limb spreading in anthesis. Fruit a pyxis, — 2-celled, the partitions falling away with the seeds; seeds various. [The Latin ame wi genus of over 200 species of wide geographical distribution. Type species: Plantago major L. — se flowers spicate or capitate at apex of s s more or less laciniately divided (denticulate or oa entire in variety) ; spikes ne pe in bud et coronopus. Leaves entire or _ ater spikes erect in bud. Stout coarse perennials; stamens 4. em 8-16; g WE ovate, distinctly slender-petioled; caudex not woolly- bin ~ at leaf- ae SF ee Seeds 2-4(6); leaves ome linear-lanceolate, or if naxneaiee ovate these nda sg a short broad petiole; caudex usually woolly- willows at lea se Flowers ‘polygamous, of 2 kinds; the pistillate nici: lobes erect in ie: becoming condupli- d beak-like over ri — ed oo ule. . P. hirtella Seieetitena. iwc sivas oat alike, rem ets or reflexed over rnened ri ule. Capsule 6-8 mm. long, sitshie & “ie ular P. macrocarpa. besa 2-4.5 mm. long, circumscissile. Corolla pubescent without; leaves fleshy, glabrous; maritime species. 5. P. juncoides. hat Corolla glab ; leaves th Flower- bracts attenuate with alias tip, — Rocio J the flower-buds in young inflor eeds 2, conspicuously vated o: be nner surface. . P. lanceolata. saa sind ovate, wareely, as long as et aa aries in young inflorescence; seeds ually 4, flatten nner surfac 7. P. eriopoda. Slender — or if Wiech ie coarse and cond aes +" or 7% Stamens 4; corolla-lobes +: 5 mm. long. owers mostly dioecious; the pistillate corolla-lobes conduplicate, forming a beak over sould se, € : Seeds yellowish, aa face concave; sae rounded at apex. 8. P. virginica. Seeds: ‘haa red or coh apap ie face pin Ae t 9. ag uncata firma. Calyx-lobes short-acuminate at apex; seeds wine red. 10, fF. rhedosocad. Flowers perfect; corolla-lobes all spre cian or reflexed Bracts linear or subulate, = long as to much carpus the calyx, not scarious-margined Bracts 6-35 mm. long, “ait much surpassing calyx; plant darkening ying. . P. aristata. Bracts as long as or somewhat longer than calyx (lowest bracts in var. oblonga two to three times the length of calyces); plants grayish. 12. P. purshit. Bracts aa Be cg central midrib and wide scarious margins, about equaling to much shorter an ca Bracts resembling the sepals and about equaling them in length; mature seeds reddish yellow or reddish, shining 13. P. insularis Bracts geen sepaloid, — one- “half the length of the peoels? . red bw, the surface hive -_ > ctate Stamens 2; corolla minute, th mm. long. Seeds (2)3-4(6) ; scapes and leaves erect. Capsule conoidal, twice the sl of the calyx; corolla-lobes spreading or reflexed in age. 15. P. bigelovit. eta a, ovoid to et car but little longer than the calyx; corolla-lobes mostly erect e but not — 16. P. pusil See ceyk-Ad; pine and lea’ 4 aalinihouk or spreading. 17S: heterophylla Stems leafy; flowers net on axillary pedunc ce 1. Plantago corénopus L. Cut-leaved Plantain. Fig. 4969. Plantago coronopus L. Sp. Pl. 115; 1753. ual Se short- ato pa with a taproot, or in biennial or perennial plants the root- Ann crown becoming lignesc 2-2.5 cm. broad, somewhat lobed and sending out several tufts of leaves and sande Erayish-pubescent Rae cog peaed Se eaig 2.5-12 cm. long, pinnately divided into linear, ac segments, or in robust plants some Cees es bipinnatifid, the segments often remote oot — r less dec etsy cm. long including the spike; spikes slender, 2.5-9 cm "eae, dense, the Rice crowded and more or less imbricate; bracts ovate, 4 g 4970. Plantago maj 4971. Plantago hirt or ella PLANTAGINACEAE 972. DI 4973. Plantago juncoides 4974. Plantago lanceolata PLANTAIN FAMILY 17 pubescent, with br = hyaline, epee Se oh ae rages narrowed to a pect point; flowers per- fect; sepals pubescent, 3 mm. long with broad, white, scarious margins ; corolla- lobes earings reflexed ; capsule ae the calyx, re pene pga winged all hate or at least to below t iddle. Sparingly introduced along the coast, Whidby Island, Washington; Linnton (Portland) and_ Port Orford, Oregon; —— coast in California from *southern Marin County to Monterey County, and on Santa Catalina an s y,a a ug. Island, L geles County, also adventive in Solano County. Type locality: Europe. April-A nual specimens from Santa Catalina Island, Los Angeles County, are probably Plantago coronopus subsp. commutata (Guss.) Pilger, Rep. Spec. Nov. 28: 287 50. P. commutata Guss. Prod. Sic. Suppl. 46. i : ‘ ‘ 19 1832; P. parishii J. F. Macbride, Contr. Gray Herb. No. 56:61. 1918.) This Mediterranean form is character- ized by the less acuminate and scarcely spreading bracts subtending the flower and the proportionately stouter peduncle. The leaves in Pragati baa plants _ are often entire. A complete list of ph ety of this and the more common perennial form o Pflanzenreich 4°: 126-155. 1937. 2. Plantago major L. Common Plantain. Fig. 4970. Plantago ee Sp. Pi. ria. 1753: Perennial, glabrous or aailis's more or less sparsely short-pubescent. Leaves usually riage Fe Rees ovate, often subcordate, entire or coarse and irregularly : long, usually abruptly narrowed below to form a shor ae 2 on the apex of the petiole, this ecad and channeled ; scapes more or less curved, often decumbent, 8-40 cm. long including the more or less Fai piel sepals broadly ovate; corolla-lobes gre; 1-1.5 mm. long; a ule ovoid, about m. long, circumscissile near the middle, 8-18-seeded ; seeds angled, reticula able species of worldwide distribution. Occurring as a weed in Pacific States nie wide altitudinal ange an di varied habitats. A complete list of ie many eg oe may be found in p aber dss 3 4200 ; 41 56. 1937, writes by Robert Pilger. Type locality: “tim Eur ad vias.”” May—Nov 3. Plantago hirtélla var. Saige rap Leaner Pilger. Mexican Plantain. Fig. 497 penaiiaya ee dana in DE. ris 131: 726. 2: Planta irginica var a A. Gray, Bot. Calif. 2: 611. 1880 Planta | Sipe Piles, ‘Notizblatt §: 260, 19 Plantago durvillei subsp. subnuda Pil r, Pflanzenreich 4280; 234. 1937. Plantago hirtella var. galeottiana Pilger, op. cit. 257. Perennials with short thick ake clothed with yg i leaf-bases intermingled with long villous hairs, the stems erect, arcuate ase, 1.5-4 dm. long, with septate flattened hairs espe- cially near the inflorescence. Leaves thin, elliptic to rather br aie obovate, 5-20 cm. 1.5-8 cm. broad, narrowed below to a broadly winged petiole less than — the length of the leaf- blades, the veins 5-7, prominent, margin sparingly denticulate or i ire r less pilose especially on the upper surface ; inflorescence polygamous, a9 spikes densely flowered excep at base, 5-30 cm. long, ad ; acts slieent a little sho than alyx, carinate, ovate, te scarious and f ith rse cilia; sepa tuse, scarious- rgined ; corolla-lobes of pistillate flowers inrolled and erect in anthesis ; corolla-l of i- nate o rfect flowers spreading, narrowly ate, enti long; stamens white, 4; capsule oblong, abo? as long as the calyx ; seeds 3, flattened on one side. Moi und, Transition ame Sonoran Zones; in the Pacific States mostly along the coast from Grays above County, Weskincta, to San Diego and San Bernardino Counties, California; eastward to Arizona and Mexi Type locality: ‘In Mexicanis Cordil. tires Real del Monte 2500 metr. (Galeotti n. 1427).” May—Oct 4. Plantago macrocarpa Cham. & Sch. Alaska Plantain. Fig. 4972. Plantago macrocarpa Cham. & Sch. Linnaea 1: 166. 1826. Glabrous Sacra with a stout rt so oh root. pot several . any, pee the ~— thin and glabrous, broadly to narrowly lanc 7-20 c ong, wide, acu i arrow Pont the mi iddle or era to ae narrow sition. peri ahs or cisauity di blade; scapes sath with the spikes uae Sl longer to a little cgay than the leaves, glabrous below, rather sparsely puberulent just below and on the spike; spikes ~ sely > y flowered; bracts broadly oval, closely enfolding the “capsule 3 mm rather dense obtu brownish and sparsely puberulent dorsally, the whitish hyaline margin very narrow ; ceaala elliptic, pect ate ag dly so, glabrous or sparsely os on t ~ —— eae brown midrib corolla- be long, the lobes triangular-ov. 2-2.5 mm. long; stamens much ex : _ tu ellipsoidal, 68 mm. long, not eels: but iecceiacle hasten: seeds 4, black, 44.5 m long. T ition and Boreal Zones; known in the Pacific States from the Olympic Penin- ei Cicgun &f (Peck), but ranging from Sitka, Alaska, an -. the Aleutian Islands lity: Unalaska, Aleutian Islands. June—Aug Edges of swamps, Humid sula, Washington, and Lincoln County, to Vancouver Island, British Columbia. Type loca 5. Plantago juncoides Lam. Pacific Seaside Plantain. Fig. 4973. Plantage geen Lam. Encycl. 1: 342. 1783 r. juncoides A. Gray, sais, ed. 2. 268. 1856 Plastaas maritima pgs ny American authors, not Perennial, the stout taproot with a eagle or short-branched crown. Leaves Ringed to linear- tek pa slightly i ascending, attenuate at apex, aod cm. long, 2- a bad or sparsely denticulate, glabrous or sparsely puberu lent; scapes strongly eeethie s to exceeding the leaves ; spikes 4 $7 cm. Fane densely flowered, te flowers perfect ; Stats Sek. 18 PLANTAGINACEAE carinate, broadly ovate; sepals oblong-ovate, 2-3 mm. long, often ciliate on the margins at apex ; corolla-tube pubescent, the lobes Leg ipinng: 1.5-2 mm. long; stamens and styles long-exserted ; capsule 2—5-seeded, the seeds a mm long. Banks and margins of apt along the coa d Transition Zone; southern Alaska to = Francisco Bay, Rees also Patagonia. Type locality: ee | i Rie ‘del Fuego. Collected by Commerson. June—Oct. Plantago juncoides var. ocker Fernald, Rhodora 27: 100. 1925. (Plantago maritima var. californica Pilger, Pieocnrcick 4709; 187. 1937.) A a fleshy plant with caudex often branched, the leaves depressed es rosulate, pronity linear or i) oblanceolate to yet onsen aanes stout, depressed or arcuate. Blu and beaches along the seashore, Clallam County, ashington, o Mon y County, California. Type Bi Montara Point, San Mateo County, California 6. Plantago lanceolata L. English Plantain or Ribwort. Fig. 4974. Plantago lanceolata L. Sp. Pl. 113. 1753. Perennial or — _ short rootstock bearing conspicuous tufts of long brown hairs at the bases of leaves. Leaves narrowly oblong-lanceolate, mostly erect, acutish to acuminate at apex, narrowed below to the neal elongated, more or less winees ‘agile 4-40 cm. long including both blade and porte blade 5-35 mm. broad, entire or irregularly and remotely denticulate, glabrous or nearly so above, more or less pubescent Senet ea in the rather prominent, 3-5, parallel a like aoe ees rather slender, channeled, 1- igh; spikes ager short and ovoid at t, becoming elongated and cylindric i in ae 1.5- 6(10) cm. long, 5-9 m thick: flowers perfect ; Bae hyaline- margined, hice = on with an cb ai 8 bes a -acuminate on line tip, Sa ee ously surpassing the calyx; sepals with 2 a narrow gre rib and broad scarious margins, the two lower ones usually united ; corolla elabe ous, tecechish it ite tube short, the lobes ovate, spreading, sometimes ascending in fruit ; ‘filame s white; capsu e oblong g, very obtuse at apex, slightly longer than the calyx, 2-seeded ; seeds pte excavated o e fac ields, oedatle re ane myete places, frequent CHADIAN ong os ‘States and across the continent. Natural- ized pire Rubens. Highly roi: a to loath and pubescence of leaves, also as to length of mature s oe, whic ay be head ike, (The tall robust form — ig oe: — has been called P. altissima L.) A complete list 3 gurtice, sub- varieties, and forma) is to be fou n Pflanzenreich 4°; 313-327. 1937. 7. Plantago gaat Torr. Saline Plantain. Fig. 4975. Plantago — Nutt. Gen. 1: 100. 1818. (Nomen dubium.) Plantago a uata James, Lo: teak 1: 445. 1823. Not Wallr. 1820. 45, Plantago oblongifolia Decne. in A. DC. Prod. 131: 700. 1852. Plantago retrorsa Greene, Pl. Baker. 3: 32. 1901 Plantago shastensis Greene, loc. cit. Perennial ; rootstock stout, sometimes elongate, the crown more or less long-villous with rust- colored haies among the old leaf-bases. Leaves oblong-lanceolate to elliptic, acute at apex, 6-25 cm. long, 5-9-nerved, narrowed toa sae petiole about one-half the length of the blade, glabrous e athe ith “ i 8 cm oos owere " long ; bracts broadly ovate to rou — the narrow scarious margins sometimes soe vd Batis not keeled ; sepals debt sarious-marg ned, = af 5 cm. lo sie corolla-lobes ovate, often metri- cal; style much exserted, as lon age than the stamens ; capsule bro adly conical arnts cs mm. or more ise olen ger by Beeastent base of style; seeds 3-4, black, 2-2.5 m cca od soil, Ris nsitio ; Nevada and Wyoming pent o Nova Scotia and Quebec; on a ee fic Slope it is coun fro’ eens: ae ti sh Columbia ; Jordan Valley, Malhe eur County, Oregon; and eastern Siskiyou soa Ae California, is ay it was first collected by Greene and later described by him as P. shastensis. Mor re } mater be eeded to evaluate this entity. Type "era lity: “Depressed and moist situations along the Platte.” une- 4 =“ 8. Plantago virginica L. Dwarf Plantain. Fig. 4976. Plantago virginica L. Sp. Pl. 113. 1753. Plantago ere Walt, Fl. Car. 85. 1788. Plantago missouriensis Steudel, Flora 32: 409. 1849. nnual with a slender vertical taproot, eporate or conan more or less villous va septate hairs. Leaves spreading or ascending, 1. long, the blades ablences late or meals wi! 2 obovate or elliptic, entire or repand-dentate, af are wr eanel to slender or more o gined petioles ; scapes rect scending, cm. long; spikes dense or sometimes Dees A below ; flowers dioecious ; bracts. lanceolate to linear-lanceolate, 2 mm. long; calyx- lobes oblong to ovate, 2-2. 5 at with ‘prom ominent brownish midrib and broa oad, white, scarious margins ; coro olla of staminate flowe + with Eat quire: lobes, those of pistillate flowers erect and connivent in fruit ; pater ovoid or silengss oid, about equaling the calyx-lobes; seeds 2-4, rege yellow. Moist places, Babes ont tan Sonoran as. y en Texas, and a s the United States to the Atlant Coast cat sont be orthern Mexico; introduced in the Pacific States in Seetiwesterih sl and Shasta Caen. California. Type feuds: Virginia. May-July. 9, Plantago truncata subsp. firma (Kunze ex Walp.) Pilger. Chile Plantain. Fig. 4977 Plantago firma Kunze ex Walp. Nov, Act. Nat. Cur. 19: ove 1: 402. 1843. Plantago virginica var. firma Reiche, Fl. Chile 6¢: 117. Plantago truncata subsp. Sonia Pilger: Bot. Jahrb. 50: je ae Annual with a slender taproot, herbage hirsute throughout the spreading whitish hairs. Leaves PLANTAIN FAMILY 19 lanceolate, narrowed to a rather short petiole below, acute at x, 2-4 cm. long or rarely longer and 3-5 mm. wide; s ne “eye longer to shorter than the ya ths 1-2 cm. long, aS ae as d sist acute ; capsule about ee the calyx, 2-seeded; seeds elliptical, plane on the inner face, brownish. - native of Chile; sparingly introduced in Marin and Sonoma Counties, also in Butte and Calaveras Coun- ties, California. Type locality : Chile. April-June. 10. lanease eee oe Decne. Red-seeded Plantain. Fig. 4978. Plantago rhodosperma Decne. in A. DC. Prod. 131: 722. 1852. Plantago virginica var. aac ine Kuntze, Rev. Gen. Pl. 2: 532. 189 Plantago rubra A. M. Cunningham, Proc. Indiana ‘Asad 1896: 204. ree Annual or winter annual with a slender elongated r Leaves spe emehig several to many, narrowly elliptic to oblanceolate, narrowed to a winged seticke acute or obtuse at apex, commonly hairs; scapes 1 to several, longer or usually shorter nes the spikes, hirsute-pubesce et cm. long; bracts apn este shar il cute, scarious-margine ea ‘promine rape keeled ciliate e margins and kee obes 3 mm. long, ellipti ic-oblong, with broad scarious margins, cae soared petvedcot on rhe: "slightly bcaated midrib, apiculate oad hie areainae. at the apex; 4978 4975. eS ase eriopoda 4977. Plantago truncat 4976. Plantago virginica 4978. Plantago rhodosperma 20 PLANTAGINACEAE orolla- _ erect and closed over the capsule, 4-4.5 mm. ne Psa and ror at apex, suriculat * base; corolla-tube puberulent on the siege seeds ark wine red, a 35 1 wly lliptic- eee shallowly concave on inner, ee ae rounded = i. outer, faintly. na hes -punct and dry mesas, Sonoran Zones; locally distributed i a ig py ky toi from Corona Bluffs, Orange County, - San Diego; also, s outh of Tijuana to Rancho Cuevas, Lowe lifornia; east to Arizona, Texas, and adjacent ogee asin Mexico. 5 locality: “‘In Texas.’ April- i. iamicel aristata pana Bristly or Large-bracted Plantain. Fig. 4979. Plantago aristata Mic Fl. Bor. ngs +95. 1803. Plantago nuttallit . Mem. Soc. Linn. ee 470. 1828. Plantago patagonica var. aristata A. ai Man. ed. 2. 269. 1856. Dark 7 spe “ ual 1-4 dm. high, loosely villous to glabrate, yee on drying. Leaves ascending to t, linear , acuminate, 5-1. ’ i ; spikes surpassing the leave Se at with villous hairs, densely flowered, 3-10 cm. ieee Cele linear, ascend- ing, su ocr ae to seven times the length of the flower or longer at the base of the spike pals 3 mm. lon aici obtuse, villous, with a narrow scarious margin; corolla-lobes rotund- ae spreadin o ‘stam ns small, scarcely gone capsule ellipsoid, 3 mm. long; seeds 2, the nie roughened, concave on t the i ntae =i ge + : & Dry open ground, y Tran aw, Briti oe Columbia and found as an adventive in the Pacific States at Seattle, Washington gv te seh ra = Rogue River Valley, Oregon, and in Humboldt and Ventura Counties, California; exte g eastward to the eastern seaboard and south to Texas. Type locality: Illinois. May—Aug 12. Plantago pirshii Roem. & Sch. Pursh’s Plantain. Fig. 4980. Plantago purshit Roem. & Sch. Syst. Veg. 3: 120. 1818. Plantago guaphaloides Nutt. Gen. 1: 100. 1818. Plantago patagonica var. pnabhaliodes ss, Gray, Man. ed. 2. 269. 1856. nual; scapes erect or re caen slender, well exceeding the spikes, villous ‘oor epeni ing hairs and often also whit eaves narrowly linear- ere ol 1-4 m 3-12 cm. dong: ag ties craic: 2 ee em. long; bracts linear to linear- cabalate: eae the calyx-lobes metimes exceedin m be low; es hg lobes aot gi obovate, m ong, densely long-villous carol: lobes Sens S benaaty ovate, 1.5-2.5 mm. long, white with brown- ish spot at base; stamens small, equaling or slightly veel the poo capsule 3-4 mm. long, lion ads. fate , dark brown, minutely pitted. n o Minnesota, oe and Chihuahua. Much less abundant in California ead Arizona and icine with P. purshit var. oblo ga. Type locality: ‘‘In dry situations on the banks of the Missouri.’”’ Collected by Nuttall. May— ug tago purshii var. oblénga (Morris) pe sre jh & Lab. 18: 117. 1950. (Plantago picta Morris, Bull. Torrey Club 28; 118. 1901, not P. picta Colen oblonga Morris, op. cit. 119; pe Spee Morris, loc. cit. ; P. xerod a Mor ibe op. a 36: 515. eet i poe wien var. oblonga es op. cit. 55: _ 1928; P. purshii var. EE = 937. em; se, 0-5-1. and cm, Ton as in the species. Gane or sandy bovis slopes, So onoran Zones; Mojave and Colorado Deserts from San Bernardino County, California, to northern as vie California and bind Sey aid southern and central g Seer to western New Mexico; also Sonora. Type locality: Colorado Desert. Collected by Cerutt. March-April. The more — form in southeastern California and adjacent Arizona and Sonora, P. ignota Morris, repre- ariation Ee len gth of bracts. 13. Plantago insularis Eastw. Island Plantain. Fig. 4981. Plantago insularis Eastw. Proc. Calif. Acad. III. 1: 112. 1898. Not P. insularis (Godr.) Nyman ex Briquet 1901. Plantago brunnea pe Bull. Torrey Club 27: 115. 1900. Very similar to Plantago erecta in general habit, stems slender, usually spreading, canescent with oes ae se especially on the scapes below the spikes. Leaves narrowly to broadly linear- lanceolate, er va often with a few callous teeth on the margins; spikes o oblong-cyfindric, em long, 8-10 m ong, wide; bracts broadly ovate or orbicular, about equaling the ro or g ish midrib, broadly scarious-margined and _ closely resembling t yx-lobes Te conspicuously brownish at base, often brown-striped above middle; seeds oblong, ed yellow or reddish, smooth and ssy, 2-3 eeply excavated on nner face ce rh sandy ie Upper and Lower Sonoran Zones; gee ie oast of southern California from Point Contapciod, eyo Barbara County, to San Diego County and on the ilar ds off the coast; also on the western side of Lower California to the Viscaino Desert, — it intergrades with the variety. Specimens from the San Ten Valley, except for corolla-markings, resemble the variety. Type locality: San Nicolas Island, sea-shore flats,’ Ventura County, California. March— Mas. Plantago insularis var. fastigiata eet Jepson, Man, Fl. Pl. Calif. 956. 1925. (Plantago minima Ciena Proc. Indiana Acad. 1896: 2 1897, not P. ig DC.; P. fastigiata Morris, Bull, Torrey ciws 27: 116. — 2a scores. Morris, op. cit. ia: i ae gooddingii Nels. & Kenn. Muhle nberg a 3: 142. 8; P. in- sularis va a Jepson, Man. FI. Pl. Calif, sft 1925.) Conspicuously white-silky fede or ae annual; bracts orbicular to to heoety ovate, sepal-like, with greenish, usually hairy midribs and broad hyaline margins; calyx lobes faintly if at all brownish at base; seeds nee ce € species. cnish ape with the dense Rape ore habit represent PLANTAIN FAMILY 21 14. Plantago erécta Morris. California Plantain. Fig. 4982. lens oe patagonica var. californica Greene, Man. Bay Reg. 236. 1894. Not P. californica Greene, Bull. Calif. - cad. 1: 123, 1885. Plan 5 dace Morris, Bull. pages Club 27: 113. 1900. Plontavo avects Morris, op. cit a Morris, op. ae 9. Ploniésgn spetiied Morris, op. cit a8: 120. 1901. Flantago obversa Morris, op. cit. 1 Plantago hookeriana var. iitwaie Poe, op. cit. 55: 417. 1928. ual with 1 to several prs commonly erect or a ing, 7-20 cm. high, thinly to geod Sa eres pubescent with appressed ba Leaves narrowly to rather broadly linear, nar- rowed above to an acutish ape b to an elonga somewhat winged petiole, 41 cm. long, 1-4 ide, 3 d; spikes cylindric to ovoid-capitate, 1-3.5 : ¥ m. in diam eter ; bra pare: shorter than the calyx, sees usually , with a , green or middle or above the middle of the bract ; flowers ph calyx-lobes oblong, obtuse, i n, with b s i si. 5m Vv reflexed, brownish at base; seeds 2, rown, finely — halaok on inner surface. Common on grassy hillsides and flats, Transition and Upper Sonoran Zon Rogue River Valley, Oregon, southward west of the ge ee hay Nevada Divide to northwesterit. pie California. Type locality: ‘‘Abun- ant on grassy plains and hillsides,” San Francisco Bay region. March—May his species shows a great Varia ability of growth form, iiparentty: ae to habitat, and is expressed by height, number of scapes, and length of spikes 4979 ) | 1 \ VW Zz 4979. Plantago aristata 4981. Plantago insul. 4980. Plantago purshii he tint erecta ze RUBIACEAE 15. Plantago bigelovii A. eh Annual Coast Plantain. Fig. 4983. Plantago bigelovti A. Gray, Pacif. R. Rep. 4: 117. Annual, glabrous, sometimes ne on scapes below the spikes, scapes including spikes .5-20 cm. high, erect, 1 to many. Leaves erect, linear- lance olate to lin iy area —7 cm long, glabrous, the margins ciliate, rarely irregularly dentate; spikes densely flowered, 0.5-5 cm long; bracts fleshy, ovate to nearly orbicular, carinate, scarious- Fue ate about "te length o the sepals; sepals suborbicular, wit road scarious margi —2 m ong; corolla- as spreading to sere deflexed in fruit, about 0.5 mm. long or a little par eee 8 song acute; stamens 2; sule jars than the calyx, fe eae pyonecas truncate at a long, Prspnsscees foe ty : "Hittle eee the middle; seeds 4-6, only 4, dull black. pee Fae on ome pitted, winged at one end or ele so all hens in iecadiorial seeds, ellipsoid or oblong, about 1.5m m. long. marshes along wk coast and inland alkaline flats, Transition and Sonoran Zones; along the coast from British Co ntaiabis to Monterey County, California; also occurring in the Sacramento-San Joaquin Valley, Cali- fornia. Type locality: Benicia, Solano County, California. March— —July. 16. Plantago pusilla Nutt. Slender Plantain. Fig. 4984. acco pusilla Nutt. Gen. 1: 100. 1818 Plantago myosuroides Rydb. Mem. N.Y. Bot. Gard. 1: 369. 1900. lender an gon puberulent with i hairs, os filiform, 5-12 cm. high i i kes. Leaves narrowly linear, 0.5-2 m wide, enti l-nerved, long- villous base; spikes d se flowered, slender, 7c ee ney imperfectly dioecious or po someon us ; oe triangular-ovate, 2 mm. long, shorter than the sepals, rious-margined, slightly esied and not ccate at vate, 0.5 mm. or less lon not connivent in fruit; stamens 2; capsule short-ovoid, equaling or slightly exceeding the calyx, circumscissile well below the middle ; seeds 3-4, dark, finely pitted, sometimes winged at one end. Moist places, Transition and Sonoran Zones; Nate the alg rh mtiachin in Oregon and Seegpseile and east to we York, Virginia, aeons: and Texas. Type locality: ‘‘Ark ” April-Aug. nta: ngata Pursh (Fl. Amer. Sept. 729. 1814), a plant n pentlien United States and Canada, may es expected to occur in eastern Washington. sg bracts in this species or as long as A vexialee than the sepals and are strongly keeled and subsaccate at base. The spikes are usually loosely flower 17. Plantago heterophylla Nutt. Alkali Plantain. Fig. 4985. raat heterophylla Nutt. Trans. Gok Pini. Soc, 11. 5: 177. 1837. o perpusilla Decne. in A, DC. Prod. 131: 697. 1852 See californica rece ly Bull. wy Abad. 1 i238: 1885: apes several to many, slender, arcuate, ascending or spreading, 5-10 cm. long, about twice the length of the leaves, sparingly puberulent to glab preading, linear, entire or ofte th a few teeth linear lobes; spikes | ly flowered especially at the base, g; t ovate to almos icular, broadly scarious-margined, strongly carinate, usually a little shorter than the sepals ; sepals broadly oblong, obtuse, a wide scarious ‘ ng; a-lobes less t m. long, lly spreading in fruit ; stamen 2; capsul mewhat rou a , abou ic e length of the calyx-lobes, : e broadly ovoid, so t nded at apex t twice the leng (6)8-14-seeded, circumscissile below but near the middle; seeds blackish, irregularly and coarsely — — gled, scarcely concave on the face, occasional seeds slightly winged at ba in alkaline soil, Lower Sonoran Zone; Sacramento Valley, California, from Colusa County south in the — Sieawn arate to Tulare and Kern Counties and Baap fs Pre the adjacent Latin ane in cismontane ee California from Los Angeles County to San Diego Cou also Ari and Sonora eastward to brite wn and Florida. Type dir Diag yagi Collected ar sherry * kek yest “ plans ae described by Greene as P. californica usually have larger cache and often fewer seeds than the canoes plan 18. Plantago indica L. Sand Plantain. Fig. 4986. — indica L. Syst. Nat. ed. 10. 896. 1759. Waldst. & Kit. Pl. Rar. Hung. 1:51. p/. 51. 1802. Caniegte ogg nt and somewhat glandular annual with simple or much-branched ned stems 8-40 cm. high. as aves tn ia lg sometimes with shorter axillary leaves, sessile, hirsute hirsute villous, — r linear-lanceolate, 6-8 cm. ie , 2-4 mm. wide; inflorescen nce on exilingt peduncles from the jeaf-axils or umbellate at the apex of the — oe hea ds —- or subglobose ; cer bracts ovate, abruptly long-acuminate, the upper ovate or oval; calyx-lobes obovate, hyaline-margined; corolla- lobes na a —. — shorter eae the whe seeds 2, about 2.5 mm. long, reddi = brown, concave on the r fac Occurring ponte in ee abhi in ~ ree ae Seattle, Washin ati Hood River, Oregon, and scattered localities in California. Native of Asia. For complete synonymy see Pflanzenreich 4%: 418-421. 1937. Family 143. RUBIACEAE. MADDER FAMILY. Herbs, shrubs, or trees with simple, opposite or verticillate, stipulate leaves. Flowers regular and nearly symmetrical, perfect but often dimorphous. Calyx-tube MADDER FAMILY 23 adnate to the ovary, the limb various. Corolla sympetalous, 4-5-lobed, varying from rotate to campanulate or salverform or funnelform. Stamens as many as the lobes of the corolla and alternate with them, inserted on its tube or throat ; anthers linear or oblong. Ovary 1—10-celled ; style short or elongated, simple or lobed ; ovules 1 io many in each cell. Fruit a capsule, berry, drupe, drupelet, or nutlet ; seeds variou Approximately 400 genera, and 6,000 species, of wide distribution but most abundant in mei regions. Large shrub; flowers in dense, vin 3 long-peduncled heads; corolla tubular-funnelform. 1. Cephalanthus. Herbs or somewhat suffrutescent plan Flowers in pedunculate valerate heads. 2. Sherardia. Flowers not in involucrate heads. orolla pti sae Leaves opposite; flowers in a loose forked cyme. 3. Kelloggta. Leaves in whorls; Aaneess in clustered cymes, 4. Asperula Corolla rotate. 5. Galium 1. CEPHALANTHUS L. Sp. PI. 95. 1753. Shrubs or small trees with opposite or whorled, short-petioled, entire leaves and termi- nal or axillary, densely capitate, small, bracteolate, white or yellowish flowers. Calyx-tube obpyramidal with 4 obtuse lobes. Corolla tubular-funnelform with 4 short, erect or spread- ing lobes. Stamens ' inserted on the corolla-throat ; filaments very short; anthers “arte Plantago bigelovii 4985. Planatgo heterophylla ae Plantago pusilla 4986. Plantago indica 24 RUBIACEAE exserted ; stigma hat Fruit dry, obpyramidal, 1-2-seeded. Endosperm cartilaginous ; cotyledons linear-oblong. [Name Greek, meaning head-flower. About 7 species, natives . donereoan, Asia, and Africa. Type species, Cephalanthus occidentalis L. 1. Cephalanthus occidentalis L. Button-bush. Fig. 4987. Cephalanthus occidentalis L. Sp. Pl. 95. 1753. Shrub or small tree 1-8 m. high with opposite or verticillate leaves and branches, glabrous or somewhat pubescent. Leaves ee ovate or oval, acuminate or acute at apex, truncate, rounded fe) cm. long r narrowed at base, entire, 5-12 cm. long; petioles 8-10 mm. Fas peduncles 3-7 : heads siobaek 2-3 cm. broad, the receptacle pubescent; calyx enish, sessile; corolla white, 8-12 mm. long, the lobes obtuse, peg ancl W ith black; style ith giinor about twice the length of the corolla; a persisting on the fruit, the fruit obpyramidal, about 4 mm. high; seed flat- tened, eee Most soils ams and swamps, Upper and Lower Sonoran Zones; Lake and Napa Count in the California yore 7 phety and Sacraments River “peenely Shasta Counts, cant eonen the Great Vatics and the Sierra Nevada foothills to Kern County, California; east to Arizona, oe and Flo re also north- east gence n Ontario an ew Brunswick. Type locality: eastern North , but no definite locality given. June— 4 2. SHERARDIA [Dill.] L. Sp. Pl. 102. 1753. Slender, procumbent or diffuse, annual herbs with verticillate, spine-tipped leaves and small, nearly sessile, pink or blue flowers in termi an ary, involucrate heads Calyx-tube obovoid; lobes , lanceolate, persistent. Corolla funnelform, 4-5-lobed, the tube as long as bes or long or inse on the corolla-tube; fila the lobes ee n : ments slender ; ones small, oblong, exserted. Ovary 2-celled; style 2-cleft; ovules 1 in each cell. Fruit didymous, with indehiscent carpels. Seed erect. [Named in honor of Dr. William dhevkat patron - Dillenius. | A monotypic genus of the Old Worl 1. Sherardia arvénsis L. Blue Field Madder. Fig. 4988. Sherardia arvensis L. Sp. Pl. 102. 1753. Tufted — eh = patie Lees decumbent or ascending stems 6-20 cm. long, herb- age hispidulous. Leav the lower often obovate and mucronate, the pedi ee or lanceolate ae ad goes puted, sine ete on the margins, 6-10 mm. long, 2- ; flow n few-flowered, slender-peduncled, involucrate eads” Hnvelocre deeply 6-8-lobed, ang oles erie ea -poi ointed ; corolla pink or bluish, 4-5 mm. long, the lobes spreading ; fruit crowned with 4-6 lanceolate calyx- teet Lawns, gardens, and pasture land; native of ee and naturalized in the Pacific States mostly west of ba rs Mountains and the Sierra Nevada; Washington to southern California. Type locality: Europe. pri 3. KELLOGGIA Torr. Bot. Wilkes Exp. 17: 332. p. 6. 1874. Perennial herbs with att entire, stipulate leaves and small flowers in a loose fork- ing cyme terminating the simple or sparsely branched stems. Calyx-tube obovoid, somewhat flattened laterally, covere a with hee stiff bristles; teeth 4, very small, subulate-persistent. Corolla funnelform, with 4 (rarely * narrow ovate obes, valvate in th bud. Stamens 4, n ; stigmas filiform, papillose. Fruit small, oblong, dry and coriaceous, covered with hooked sheets splitting at maturity into 2 closed carpels, to the walls of which the solitary seed adheres { Named in honor of Dr. Albert Relogs: an early California botanist. ] A monotypic genus of western North Amer 1. Kelloggia galioides Torr. Kelloggia. Fig. 4989. Kelloggia galioides Torr. Bot. Wilkes Exp. 17: 332. pl. 6. 1874. Plants with woody rootstocks, glabrous or nearly so, the stems several, simple or branched, .5 dm. high. = ong, about equaling or shorter than the internodes, darkened in dried ee apelegd © afinreccens a loose, div ke wi ao ee re — sues cre eA 1.5-3 cm. long; 5s dull sink ‘or laven ong, the the length of the tube, acuminate ; fruit oblong, 4-5 mm. long, preset eed with ee ae stles. brist Dry ridges, Canadian Zone; in the Cascade Mountains of Washington south through Oregon to the Siskiyou Mourtains, California, — the Sierra Nevada south to goes Met California; ahnaite. to Idaho aay La aaa and south through Nevada to northern Arizona. Type locality: the ‘“‘Walla- Walla River.” May-—Jul _ a bh on a so -_ oe tr oO re) ae Ww — ® =) a ge - do un 4. ASPERULA L. Sp. Pl. 103. 1753. Perennial herbs with erect or ascending, 4-angled stems and small white, pink, or blue flowers in terminal or axillary, usually cymose aes alyx somewhat didymous, the limb obsolete. Ovary 2-celled; ovules 1 in each cell; style 2-cleft. Fruit globose-didymous, MADDER FAMILY 25 4987. Cephalanthus ne 4988. Sherardia arvens the selaiere aig eat Seed adherent to the pericarp; endosperm fleshy ; embryo curved. n, diminutive of asper, rough, referring to the leaves. aod se uci! natives the Old World. Type species, Asperula odorata L. 4989. Kelloggia galioides 4990. A 7 A. + 1. Asperula odorata L. Sweet Woodruff. Fig. 4990. Asperula odorata L. Sp. Pl. 103. 1753. Stems erect, slender, smooth. Leaves a. in whorls of 8, varying from 6-9, thin, eee late or oblang. lanceolate, Seate or obtuse, , l-nerved, roughish on the margins, 1-2. long, the lower smaller and often o oats: ase in several-flowered cymes on slender, pckory stg and axillary peduncles, white or pinkish, 3 mm. long ; pedicels 2-4 mm. long; fruit very hispid, about 2 oad. Frequent escape from gardens, especially in open woods; western Washington and Oregon. Native of Europe. April—July. Crucianélla angustifélia L. Sp. Pl, 108. 1753. Slender annual with linear whorled leaves and narrow, serminal and apeperiem: euilary Hi the tubular flowers subtended by 2 scarious bracts — "gatas mid- veins, the fruit glabro This European species has been collected in Tehama County, Califo 5. GALIUM* L. Sp. Pl. 105. 1753. Annual or perennial with slender 4-angled stems and branches, and opposite or appar- The following new names and combinations were ma ade in the genus bogeser by poring amay a (Brit- jets: 10: 181-192. (Oct.) 1958) after manuscript was set up bor type: Galium nuttailti var. cliftonsmithii, G. nut- talht var. ovalifolium, G. nutta ice var. tenue, andrewstt Var. Brat Bs Pisces ota var. Atty G.. cata- peeks Mie ge pre G. angustifolinm var. secehce: G. matthewst var. magnifolium, and G. munsii var. Ringst se. e also Dempster, Brittonia 11: 105-22. 1959. 26 RUBIACEAE ently phere 8 leaves (foliaceous stipules). Flowers perfect or sometimes dioecious, small, , green, yellow, , mostly in axillary or terminal cymes or panicles, the fetes usually jointed with the calyx. Calyx-tube ovoid or globose, the limb mi- nutely toothed or wanting. Corolla rotate or slightiy campanulate, 4-lobed or rarely 3-lobed, the lobes involute and inflexed in the bud, often acuminate or mucronate at ap tamens carpels maturing. Seed convex on the back and concave on the face, or spherical and hollow ; endosperm horny; embryo curved; cotyledons foliaceous. [Name Greek, milk; one of Bait G. verum, once used to curdle milk. About es of wide geographic distribution and especially well represented in the Pacific States. Type pone hey yA ah E: Plants annual. Car as = nN ag much longer than broad, curved outward on the inner face; Kier ee inconspicuous mu Carpels o tee it about as b s long, rounded or nearly straight on the inner face; plants various. Fruit with coarse or ai a hooked bristles. Lea n whorls of 4 or the upper stem-leaves opposit “Fruit ey in leaf-axils on a slender alee pedicel; uppermost stem-leaves opposite, the lower in whorls of 3—4 of unequal length. 2. G. bifolium. Fruit saaeite between leafy bracts on axillary branchlets; leaves in — of 4. . G. proliferum. Leaves i in Agia. te 6-8. F 1 mm. or less broad; plants slender, usually diffusely branched. 4. G. paristense. Fruit 2-5 mm. broad; plants coarse, with long, usually unbranched, reclining stems. 6. G. aparine. wihecich on or appesculate. inflorescence much surpassing the leav 5. G. divaricatum. Pedicels stent, aac inflorescence axillary, not or askin Fi longer than the leaves. 7. G. tricornutum, Plants perennial. Stems from slender creeping rootstocks, weak, delicate, seldom entirely erect, completely herbaceous. Fruit aap roves 3—4-parte Flowers 2.5-3.5 mm. hicad: rather numerous tak bron Scag on the upper pion mig peduncles ere EER but pedicels straight and div ate in fru Pa. Flowers less than 2.5 mm. iced: 1-3 in upper leaf- jon or on ate outs cpodicale strongly arcuate in age (1-flowered and with pedicels usually straight in vars. pusillum and pac “a pall 9. trifdum subbiflorum Fruit hispid or at least yet corolla 4- parted. Leaves in whorls of (5—) 6-8, veins Cymes a vio dy Hot axillary peduncles ; carpels covered with sie ye piaa eon belaties about as widt Cymes scala j in a diffuse panicle; carpels with short hooked bristle » scabrous, . G. asperrimum, Leaves in whorls of 4, fee: apn 3-nerved. rg G. oreganum. Stems ——— at woody at least below, erect from woody root-crown or ae aceerent gps oe climbing, or tufted. from branched oom round st as s and rootstocks; if herbaceous, plants from coarser rootstocks and Leaves in whorls of 6-8; introduced speci Flowers bright yellow; leaves narrowly linear. 13. Gee Flowers white; 1 14S GG, poi Seats peas hoe ~~ # 4; ——— frcsggvad , ort apes: dioecious , thy reokd panicle of white flowers. 15. G. boreale. Inflorescence various; if paniculate, not of soul white flow Fruit gl brous or sparsely puberulent or pubesc when waeare dry or pulpy and berry-like. Fruit dry, with short, curved or straight “i or subglabrous. Shrubby, 6-12 dm. high; inflorescence leaf . G. catalinense. y. Stems tufted, 1-2 dm. high; inflorescence much exceeding a cleaves, sub ome ge meet jepson Fruit pulpy and berry-like when ripe, glabrous (pubescent to glabrate in e. pubens and sometimes in G. californicum), Plants ak: slender rootstocks with slender creeping stems, diffuse or densely low- Leaves linear or narro crag usually much longer than the internodes and often concealing 4g tems Leaves rigid, narrowly Nee subulate, acicular. 18, G. andrewsit, Leaves firm but not rigid, 1-1.5 mm. wide, abruptly acute and cuspidate. Longest —— 9-12 mm. long; ares of northwestern a and adjacent Oregon. 19. G. ambig Longest a 4.5-6 mm. long; plants of a . oo Lucia alanine: clem: one —— to ovate-lanceolate or elliptic or Seite shorter sew the inter- s (nearly equaling them in G. murtcatum). Sos dull, copiously covered on both surfaces with spreading, ie hairs, ovate or ovate-lanceolate ie: or es shining and only on the margin in var. miguelense). . G. californ aiken MADDER FAMILY 27 Leaves shining, sparsely hispid or hirsute with curved or vA ap- — Fant on the upper surface, margins, and sometimes the dvein beneath, the lower it i — or nearly so, loonie ellip tic to naar . G. muricatum Stems rather stout from a woody cine x and erect, or aa long and dinate. tems somewhat acl: long and slender, climbing or seca ng; leaves ovate to oblong (linear-oblong in some intergrading er ae 3. G. nuttallii. Stems vik from a woody caudex or rootstock, 1.5—3 ie hig Leaves firm and yy whos — glinear- on 6 ovate-obl ong or ovate and lar e lower lea wide, marginal scaberulous inies when kamen sentttes oe Stems awe, leaves enc or nearly SO, the _leaves narrowly linear- oblong (young growth g). 24, 6. bolanderi. Stems and — mages re so in var ridum), the leaves ovate- oblong or ovate, rower on much- branched plants fay i = aienk Leaves thin, mostly oval, the large lower leaves i the m as appressed; ‘peduncles and aan Bi i. aialune inflorescence piilary. . G. sparsifior Fruit dry, conspicuously hirsute or villous, the spreading hairs stir as long as or we than the body of the fru Leaves st to “asters linear, acute. low sil rae. mete acuminate be the staminate on copttiary Pedicels: plants poly pane: t t Flowers “yellowish arose or teh waite, with acute en the staminate pedicels arkedly slender; plants dioe ae DES mm. Wak ea rey ak spreading hairs; inflorescence panicu- ate. Plants tall from a woody base, over 3 dm. tall; cismontane California. 28. G. angustifolium. Plants low, about 1-2 dm. tall, tufted; San Gabriel ioepiaies, G. gabrielense. Fruit 5-8 mm. wide including the abundant, ae. spreading hairs; inflores- cence nar i, Ba cymose clusters on short ax iad y branchlets (including subsp. puber ne. sbetecudl, Leaves lin aggimasaagl ang or octicentae and acute or lig a at the apex, or lanceolate- ac eos stem- nets closely clothing the older stems, persistent; plants low and tufted; inflorescence narrow and congested, the branches scarcely exce i ad the leaves. 36 readin Fruit including the hairs 2.5-3 mm. wide; leaves rigid and acerose. Int a, Raa sige shrubs, hispidulous- Lt asisdae a seperate si tei at eremic Lower ae -leaves not approximate, not markedly persistent; Asa ample and spr ng. . G. stellatu Plants ke. erect from a woody base; (eaten open- panicslate 32. G. matthewsii. Fruit praca ge the hairs 5-7 mm. or more wide; leaves thickish or thin but not id, sometimes apiculate but not acerose. ale branchlets as well as the fruiting —. “hye te or pendent. . G. hallii. chlets erect or widely spreading, a fruiting pedicels straight hat curving in G. hypotrichium). Inflorescene broad, many-flowered, the flowering branches yr rg rwendl eaves of midstem one-third to one-sixth the len gth of the a bran : es. Plants ie bee more or less pubescent prema with hispidulous unzit, Plants pate glabrous (hirsute in f. hirsutum). 34. G. multifiorum. ripe magaoveron narrow, few-flowered, the flowering branches erect or as- ending; leaves of midstem usually one-half to nearly equaling ength of the internodes vate to pubescence of hispidulous hairs, often scabriduiocs on mmidribe and margins of leaves; fruiting cels more or less curved. 35. G. hypotrichium. Leaves broadly elliptic to oval; pubescence of dense hirsutulous hairs (becoming nearly glabrate in a grterait: ig, fruiting, pedicils straight. cer Leaves o 1. Galium murdle All. Wall Bedstraw. Fig. 4991. cae murale All. Fl. Ped. 1: 8. pl. 77, fig. 1. 1785. minutive annual with slender stems, simple or branching from the base, mostly about 2 cm. re) roadly linear to oblanceola i 2-4 mm. long, acute high, el: ous. Leaves in whorls o <5. nd mucronulate at apex, rather sane 8 ciliate on the mar qos glabrous or sparse ely bristly- scabrous on lower surface; flowers usually 2 in each of the upper leaf- whor Is, pedicels tees PB se E a es spreading or somewhat r Laat in fruit ; corolla white, les sta 1 mm mm. long and barely 1 , the c ie polis seen "ede of yea baie becomi 11 establi va inode lant Seats ¢ Siar paca ventas: a. Native of southern paid Men hace 28 RUBIACEAE 2. Galium bifélium S. Wats. Low Mountain Bedstraw. Fig. 4992. Galium bifolium S. Wats. Bot. King Expl. 134. pi. 14. 1871. the stems erect, simple or with a few ascending branches, 3-15 cm. high, sade, sides: va on the angles. Upper leaves 2, Fa pen ong sie lower usually in a whorl of linear-lanceolate to oblong- ee Bis ay long, usually of uneven length i me ora narrowed at base, acute or ro ; ar Fe owers ‘cetieclints, usually solitary in the axils ; ae slender, 3-10 mm. lo bee yidely Sing or reflexed, usuall curve a at apex in fruit; corolla minute, white ; fruit 2.5-3.5 mm. broad, covered with slender hooked prickles Open coniferous forests and edges of aneoli Transition and Canadian Zones; sci British Peni eres Wikiklsaten. gree mgt cee southward a California in the Coast Ranges to Humbo ldt County, and County through the Sierra Nevada to the San Bernardino pei ig and east to Montana and Paisisce. “ aon, “Tn the Trinity, “Battle and Dat Humboldt Mountains, Nevada, and in the Wahsatch” Mountains, Utah. nc—. iat 3. Galium proliferum A. Gray. Desert Bedstraw. Fig. 4993. Galium virgatum var. diffusum A, Gray, Smiths. Contr. 35: 80. 1852. Galium proliferum A. Gray, op. cit. 56: 67. 1853. Galium proliferum var. subnudum Greenm. Proc. Amer. Acad. 33: 461. 1898. Annuals, the stems simple or often branched, erect or somewhat decumbent at base, 1-3. : high, sparsely scabrous on the angles or a Cotyledons pe Ra the blade about equaling the petiole, persisting; stem-leaves 1-veined, in whorls : 4, hispidulous to coaagr sees to narrowly ovate, 4-8 mm. long, short- peice on the lower stem to sessile a A internodes c sh uit; flowers white or pale yellow, about 3 mm. wide; fruit 2.5-3 mm. wide, nes iene the covering of fine som el bristles bine oo Sipe Sonoran Soe Kingston and Providence Mountains, San Bernardino County, California, —_ as n Texas and northern Mexico. Type locale: een Texas. Collected in 1851 by Wright. March— pri 4. Galium parisiénse L. Wall Bedstraw. Fig. 4994. Galium Steno Ls So. PT, 108.-1753. Galium litigt DC. in Lam. & DC. FI. Franc. ed. 3 4: 263. 1805. Ann ae we stems eet or ascending, 0.5-3.5 dm. high, very slender, much bays in aise sixes from the base, rough on the angles, the branchlets flowering from m 4-7, lin e base to mit of the. laa. Teives "ikeaty in whorls of 6, varying fro ran near linear: lanceolate, often reflexed, 4-10 mm. long, cuspidate, minutely scabrous on “yo ye cymes several-flowered, axillary and terminal on filiform peduncles, 44 mm. long, dnvirieately 'spread- eM ing; ers greenish whe minute ; fruit barely 1 mm. wide, densely sativa with slender hooked ristles. Hillsides and stream soning Upper eoeeae and Transition Zones; California Coast Ranges from Humboldt County to Santa Cruz County mera slope of the Sierra Nevada from Butte County to Tuolumne County, Cali. fornia. Native of Europe. June. 5. Galium divaricatum Lam. Lamarck’s Bedstraw. Fig. 4995. Galium divaricatum Lam. Encycl. 2: 580. 1788. Koch, S poaragte ———— var. aeaeten n. Fl. Germ. ed. 2. — 1843-45. arpum Tausch, Bot. Zeit. 18: 354 Galium jean var. ee Huds. ex Jepson, Man. FL. cong "Calif. 958. 1925, as to California plants, not G. anglicum Huds. Habit much as the preceding cgi with stems smooth or nearly so. Leaves seeoiy 3 in kph of 6, linear, cuspidate, more or less scabrous on the margins, 3-10 mm. long; inflorescence a a0 preceding ape cies but the peduncle capillary, aay see nd a Se bot Hi aes rolla white or tinged with pink; fruit granulate, devoid of hairs, less than 1 m Grassy hill slopes and fields, Upper Sonoran and Transition Zones; Willamette ver ae scat in the Coast Ranges to Marin County, Calfocnin: ahs aba Conny. Native of apace: June— 6. Galium aparine L. Cleavers or Goose Grass. Fig. 4996. Galium aparine L. Sp. Pl. 108. 1753. Galium vaillantit DC. in Lam. & DC. FI. Franc. ed. 3. ods 1805. Galium agreste a echinospermum Wallr. Sched. 59. 1 Galium aparine B minor Hook. FI. Bor. Amer. 1: 290. ae Galium spurium var. echinospermum Hayak, FI. Steierm. 2: "393. 1913. Annual, from a slender root, the stems slender, weak, ascending or usually scrambling over other bushes, 1-15 dm. high, Stories erect i n dry open places, sipiea ons hispid on the angles. Leaves in whorls of 6-8 or rarely 4, linear ‘o narrowly lanceolate or oblanceolate, coumtalaie - apex, tapering to the base, 28 ae long, Scihariety hispid on the margins and midrib; flowers 1-3-flowered cymes in the upper axils, becoming cymose-pan Seiad on welder plants ; scscolle duloes saint pentane pedicels oe 873 divarktaiety spreading, 5-20 1 mm. long; fruit 1.5-3.5 mm. broad, ered with hooked SS the body of the fruit. gre s habita y edges of thi s and open woodland, and of wide distribut mon wend Widely distributed in ihe Podae Bestes and across the continent; also Europe and Asia. hee levels - = rch—-Aug. Quite variable as to size of fruit and density of the covering of hooked bristles. Trizona MADDER FAMILY » 2 4991. Galium murale 4992. Galium bifolium 4993. Py Pe Wsfartim 30 RUBIACEAE 7. Galium tricornitum Dandy. Rough-fruited Corn Bedstraw. Fig. 4997. pomeotien een Stakes i in With. Bot. Arr, Brit. Pl. ed. 2.1: 153. 1787. (Illegitimate name.) Dandy, Watsonia 4: 47. 1957. Annual, Sa = fe decumbent or ascending, 15-45 cm. — simple or wig aa netepeseaite n the ores stg ade Eger us. snail whorls o near to narrowly eee eis 25 mm. long, wide, mucronate, p visti ly and rattle barbed on the margins and on the seiiteth « ny eat axilla ry . flowered, the i i uncles stout, about as long as or shorter tha n ve jeaves : ig erar) . stout, pr curved ‘dow ward in fr uit : corolla yan aa s and grain fields; sparingly introduced i in the Willamette Valley and in Josephine County, western Oregon, an sekteni California in the region of San Francisco Bay and a: Sierra Nevada foothills and in San Luis Obispo County; also eastern United States. Native of Europe. J nr een 8. Galium cymésum ie Pacific Bedstraw. Fig. 4998. Galium cymosum Wiegand, Bull. Torrey Club 24: 401. 18 Perennial, . stems freely branching, seas ef weak, 3-7 dm. long, more or less minutely sesthiecunt on the ngles Apna ves mostly in whorls of 6, oblanceolate to linear- oblanceolate, air at apex, nat 0. - long, 1-nerved, minutely scabrous on the margins and midve bes lower stem-leaves i ane sometimes reflexed; flowers in small c t the pa ‘of t numerous upper branches; pedicels zather short, straight, and divaricate in fruit; corolla as or ol with rose, 2-4 mm. wide, obed, the lobes po Ai ovate; fruit glabrous, each carpel globose, about 1.5 mm. in diamet meadows and sphagnum oe Humid Transition Zone; io Meera south on the ern side | sds: the Cascais Makatuns. Humboldt County, California. Type locality: ma, Pierce Ghunty, Meacham, un ne saepibhe species, perhaps a with ey at least Leteat sy resembling the eastern G. trifidum tincto- rium (L.) Torr. & ony, Fl. Lg er. 2: 22. 1841 (G. tin om. L- Sp.. Pl. 106. 1753; G. claytonts Michx. FI. Bor. Amer. AG 78. 1803; Ra nl subsp. ry a 2 city "Rhoden "it: re 1939). Some specimens are —— - like ecological Ceauke with few-flowered cymes and smaller flowers which are scarcely distinguishable rom that species - Galium trifidum var. subbiflérum Wiegand. Trifid Bedstraw. Fig. 4999. G trifid r. subbiflorum Wiegand, 3 Torrey _ 24: 399. 1897 Galium phones Wee var. submontanum Wight, Zoe 5: 53. Galium claytomii var. subbiflorum Wiegand, Rh se 12: 22. ‘1911, Galium tinctorium var. subbiflorum Fernald, op. cit. 39: 320. 1937. Perennial with slender rootstock and Se weak, erect or oe stems 1. - dm , ee sharply 4-angled, glabrous or ewe t scabrous on the angles. Lea in whorls of 5-6, so times 4, the . wer ss often exed, somewhat une linear- dhlancealate, pi ietest 7-12 m long, 0. scmserhat flaccid, usually so ee me = abrous on ri se and margin; flowers pesleeiiate e, 2 Gee gia iat n the axils Me ets r leaves or on reading, axillary branchlets in bracteate cymes o of 3 or 2, the sadivele gi ic ses about apart the leaves or a little liateer, eee deflex tae ate at least in fruit; corolla minute, about 0.5 mm. long, the lobes 3 (sometimes 4), obtuse; fruit glabrous, globose, each c carpel about 1 mm. wi % Moist ground or marshy meadows, Upper Sono and Transition Zones; British Columbia to southern Cali- io pase _ to the Rocky Mountains and across the Sorthern: United States and Canada. Type locality: Colo- bauac baartice taxon, a small plant with spreading slender branches, linear leaves in whorls of 4, and capillary, readely spreading pedicels, inhabits northeastern North America Canada, and irom Some specimens anogan County, Washingtha, resemble the species and may on further study of m aterial prove to be typical. The complex is highly varianle | an d the varieties in the 2 aig States are ee ieroke. ar wg many inter- grading nted out by many authors Galium trifidum var. ae sea Bull. BS ihe A eo 24: 400. 1897. (Galium tinctorium var. diversi- = Wight, Zoe 5:54. 1900; G. columbianum R ydb. ocky Mts. 808. 1917; G. trifidum, subsp. colum- bianum Hultén, Fl. Aleut. is "307. 1937.) Werencias ms sin rootstocks, the stems slender, diffuse and reclining, 2-6 dm. long, with creeping axillary branches sometimes a arising from the lower —— slabrous ig for the minutely retrorse-hispid ang en leaves oblong ogee to ag pete oblance Sap ong e, commonly whorls of 4-5, more or less ——— the marg and midrib, 10-25 mm. long, m. "wide: pedun bias capillary, mostly 1-flowered o ace s 2-flower red “when iebusiast, divaricately spreading often slightly an scabrous, longer than, Saualing, pe shorter than the leaves; corolla minute, about 0.5 mm. broad, 3—4-parted; fruit ene. glabrous. Moist ground o: n edg es of marshy meadows and streams, especially near the woods or Sh Transition and anadian faba ‘Aleee and tebopacte gid Island south on the Pacific Slope and Cascade Mountains of Washin — and Oreaee to the Sierra Nevada and San Bernardino Mountains, California. PDS oy locality: Sal ormad Co. Cal. er ( 1an2) Bottom ing of the rigs eg ics a jemnl ba 1661 (1893).” June—-N ome © bogs along the coast of he fcogvi d Oreg s occasionally have been callanted that suggest — tinctorium var. “abvador m Wiegand (Bull, ores | Club sy 398. 1897; G. la wide tei b a ago nd, hodora 6: 21. 1904). The lanai are ie dm. tall, ee leaves in whorls of 4, sometimes 5 below, and ja ase small, obtuse, and rather firm in texture; flowering material no pan Galium trifidum var. pusillum A. panty Man. ed. 5. 209. 1867. Perennial with slender weak-branched stems fo: neva mats, glabrous or nearly , 5-12 cm. long; leaves in whorls of 4, glabrous, narrowly aesucecuite. unequal in length, 3-8 mm. long; pedicels solitary or sometimes in 5 vain straight or somewhat curved, usually glabrous, rather slender, 2-5 mm. loam. fom as in ~ ptbst age et mountain meadows, Canadia d Hudsonian Zon Cascade Mountains in Washington and Oregon to the Sierra Sewada and San Bernardida & < Bacheray ‘Calitorsia, eastward to the Atlantic seaboard in mountainous areas. Type lity: not aes in oe original reference but said to be “in d sphagnous swamps, northward.” m trifidum v Ue e 2 t 1876). However, a specimen fe the Gray Herbarium bearing the Synoptical Flora label, collected by ‘Lemmon (no. 1217) at Webber Lake, Sierra founty,: California, and Hie ved with the description given above, was identi- fied by Gray as G. trifidum var. pusillum MADDER FAMILY 31 10. Galium aspérrimum A. Gray. Tall Rough Bedstraw. Fig. 5000. Galium asperrimum A, Gray, Mem. Amer. Acad. II. 4: 60. 1849. Galium asperrimum var. asperulum A. Gray, Bot. Calif. 1: 284. 1876. s froma ee wir rootstock, erect or eben g and jag af = supported, 3-8 dm. high, t r ; t the margins and midribs, otherwise glabrous; flowers on rather long aw pod pean mostly in terminal, many-flowered, leafy, nde hee area fakacing white, 2.5-4 mm. broad, the lobes cuminate; mature fruit 2.5 mm. broad, granulate-scabro minutely hispid ae “Siowse d prickles ; pedicels 5-10 mm. long, ciniiieee het slightly nelted just below the fru ally in moist thickets, Arid Transition Zone; Kittitas County, Washington, southward oo of ag Cascade Mountains to er California, and south to the central Sierra Nevada, Mariposa County, Califo ri ia, and east o Idaho, Ut ah, | a western Colorado, western Texas, and Si naloa, Mexico. Type Lecatien: “wet places, t nta Fe,’’ New Mexico. Yon ne~Aug. 11. Galium onan ne Fragrant Bedstraw. Fig. 5001. Galium triflorum Michx. Fl. Bor. Am Galium triflorum f, Dhidiees Lependecker, iowa Stat ‘Coll, Journ. Sci. 15: 180. 1941. Galium triflorum f. hispidum Leyendec Sweet-scented perennials, eat stemmed from a clustered root, the simple weak stems 1-4 dm. long, occasionally remotely branched, reclining or erect, and rather narrow-leaved in 4999. Galuim trifidum 5001. Galium triflorum 5000. Galium asperrimum ae RUBIACEAE exposed aig oa glabrous or somewhat barbed or hirsute on the angles. Leaves in whorls of 6, it 3-flow cyme, orked and each branch h 1-3-flowered ; peduncles axillary, slender, sur jessie the saghtine hrackeate at base of cyme; corolla greenish, Pe 3m ya the | Lie fro earn fruit mm. broad, densely hispid with rather slender, long, hose hai In woods, ae Sonoran Zone to Canadian Zone; Alaska to southern ee ao across the continent; also in Japan, the Himalaya Mountains, and Europe. Type locality: Canada. April-Aug 12. Galium oreganum Britton. Oregon Bedstraw. Fig. 5002. Galium oreganum Britton, Bull. —— Club 21: 31. 1894. Galium kamtschaticum subsp. oregan Piper, Contr. U.S. Nat. Herb. 11: 526. 1906. Perennial with very pee creeping rootstock producing slender rootlets at the nodes; stems Sneed 2-4 dm. high, e rect or decumbent at base, glabrous or slightly scabrous on the mag a sh . ang Leav whorls 4 ovate to ovate-oblong or age et ic, acute or acut = ; the lower stem-leaves smaller tg the up 1.5- thin, 3-nerved, ciliate margins and often on the nerves above, glabrous below or esis hairy, ee towar a fie apex, the leaves of the oc poe lower branchlet obtuse; inflorescence from the upper axils and terminal, the terminating, ascending, peduncle- like branches rae or less peel branched, each yea bearing 6- owers; corolla greenish yellow m. wide, s 3-nerved, bout wide, acute at apex; fruit sparsely to meld ceeied “with et tistics: about 4 mm. boon . Moist woods, Transition and Canadian Zones; Olympic Mountains and western slopes of the Cascade Moun- tains in Washing and a — in hackson and Josephine Counties, Oregon, and the . acent Siskiyou Mountain area, California (Bacigalupi). Type locality: Oregon. Collected by Howell. June— The related G. kambedhapicoe Steller differs by the oer ly Wectons es ered ert “bearing 1-4 flowers and also by the oval to orbicular leaves which are occasionally seen on specimens of G. oreganum only on axillary basal branchlets. 13. Galium vérum L. Yellow Bedstraw. Fig. 5003. Galium verum L. Sp. Pl. 107. 1753. nial, from a creeping rootstock, with a slightly woody base, the stems slender, erect or ascendicat 1.5-9 dm . high, simple or with many short axillary geaee mooth or minutely pubes- cent; herbage darkening on drying. ae es in whorls of 6-8, aa linear, 6-20 mm. long, charter than the imernodes, about wide, roughened on the 7 oie Fe margins, usually be- coming deflexed in a : flowers yatlier pene the cymes in dense narrow panicles; fruit gla- brous, about 1 mm. a Ballast and waste places at Linnton and sparingly distributed in lawns of the Willamette Valley, Oregon, and Seattle, Washington; also reported from San Mateo County, California. Type locality: Europe. July—Oct. 14. Galium molligo L. Wild Madder. Fig. 5004. Galium mollugo L. Sp. Pl. 107. 1753. nnial, glabrous throughout, the Se smooth or sometimes with spreading pubescence, difuselyt branched or erect, 3-10 dm. long. Leaves in whorls of 6, rarely 8, linear to oblanceolate, ~s g, 1 5-5 mm. wide, cuspidate at apex, sometimes slightly rough on the margins; flowers bak any-flowered, panicled cymes ; corolla white, sa with purple, 2-2.5 mm. broad; ab iio filiform, monty. diva eS fruit smooth and glabro Waste places and lawns, sparingly introduced in British Columbia, oF around Portland and in the Willamette Valley, Oregon; along the fa i Humboldt out and — meeeaty in San Mateo and Santa Clara Counties, California; also in eastern Un . Sta tive of Euro er Sept. Gal atile L. Sp. Pl. 106; 1753. . onions perennial ase peaxes ay in whorls of 6 and trailing stems, the Aowering — ecuian. Pests at leafy panicles of white flow Reported from lawns in the Wil- lamette Aes Orego: Galium dae L. Sp. Pl. ed. 2. 155. 1762. Perennial with many stems, erect or nearly so, up. to 6 dm. weld ‘leaves xe ho ap of 6-8, glabrous, teleaateag acuminate, mostly about 3-5 em. long; inflorescence a loose, inal, bracteate panicle with a scending c capillary dicels; the flowers numerous, white; ‘troit arth Ee Euro- pad plant occasionally escaping yo cultivation in Salem, Oregon, and Pullman, Washington. July—Sept. Scotch- mist or Baby’s Breath. 15. Galium oad L. Northern Bedstraw. Fig. 5005. Galium boreale L. Sp. Pl. 108. 175 Galium septentrionale Roem. & pit et 3: 253: 1818, Erect, leafy, perennial herb, smooth and glabrous, simple or branched, often strict, 1.5-8 dm high. Leaves in whorls of 4, lanceolate or sometimes =e linear, 1-nerv ed or the lower broader leaves nerves: obtuse or acute, fee 2 toe ng, 2- slit wide, margins Seinetases ciliate ; flow- ers in compact terminal cymes f ormin ny-flowered sotto he corolla wn 3-4 mm. br oad; fruit about 1 mm. broad, hispid with short. facarced al sometimes glabra Borders of meaiows and Be ep woods, mainly Canadian Zone; Alaska south to enti and Shasta Counties, California, east a s the continent; also in lit and Asia. Type locality: Europe. May—Aug. Diets A pe tui Sissi species quite variable in width of leave and pubescence of fruit. G. boreale var. hyssopifolinm 2 DC., with ibsos and G. borea edium DC., : h shor re or inc ay hairs, are ers toe that occur within our limits, as well as the species with its hon: salons. MADDER FAMILY 33 16. Galium catalinénse A. Gray. Catalina Bedstraw. Fig. 5006. Galium catalinense A. Gray, Syn. Fl. N. Amer, ed. 2. 12: 445. (Jan.) 1886. Ste oody, forming an erect, rather compact, branching shrub 4-12 dm. or more high, scars of | beat ones es E Depertces, the younger pcanctes sharply 4-angled, the were ones numerous and very leafy, sparsely and pr aia He scabrous. Leaves in whorls of 4 or merely opposite on the ‘lo chl ny “persistent, always longer than the internodes, midvein and male the lateral veins prominent, linea ong, m es ne those of the main branches 15-25 mm. long, 2-5 mm. wide, of the ultimate ‘Sranihes often only 5 mm. in length, thinly hispidulous with stout curved hairs or paige inflorescence 1 sar shy ymosely pity thesflowers on short pedicels; gig white, 3-4 mm. bro ad, the lobes ovate; “Fruit dry, glabrous or with a few hors straight hair Rocky ledges and a Upper Sonoran Zone; an insular spec gro n Santa Catalina and ice Cle- mente Islands off the coast of southern California. Type locality: Sasitn Catalina istand. ee Angeles County. arch—Aug. alium buxifélium can gia Neiee” Calif. — a: 96 cea Pe Seg, ony in — with G. catalinense ; eaves rather firm and shining, oadly oblong to obovate-oblong, 1 long broad, glabrous or sparsely hispidulous; inlorescerice oe in G, Galilataee: ‘Ta ns el or with short, Mio curved ire Rocky slopes, Santa Cruz and San Miguel Islands, Santa om thar a County, California. Type ‘localit ty: Santa Cruz Island. Perhaps only subspecifically distinct from G. catalin if; Sele jepsonii Hilend & Howell. Jepson’s Bedstraw. Fig. 5007. Galium subglabrum Jepson, Man. FI. Pl. Calif. 962. 1925. Galinm senadadt Hilend & Howell, Leaflet s West. Bot. 1.135. “1934. Stems cespitose at the ends of hes smi Reape ge rootstocks, erect or decumbent at base, 0-20 cm. high, smooth and glabrous. Leaves in whorls of 4, longer than the internodes, midvein pronibatett Daa oblong or the ‘ar el to lanceolate or lanceolate- ovate, obscurely cus- 5003 a ab th 5005. Galium boreale 5006 Pe ee te $a1i 34 RUBIACEAE pidate, setose-ciliate on the margins, pried ise combine and shining; flowers few, in the axils of the eb bract- _ leaves; inflorescence narrow, the lower ascending, bearing a bract — 1-2 flowers on ver ort pedicels ; carols pale, sere 2 mm. broad, glabrous; fruit 2.5-3 m wide, jethes thinly sitdue with asc endin ng or appressed hairs. Gravelly or rocky ridges, near the borders of the Arid Transition and Canadian Zones; San Bernardino Moun- tains, southern California. Type locality: Whitewater Basin, San Bernardino ‘Mouptaine. June—Aug. 18. Galium andréwsii A. Gray. Phlox-leaved Bedstraw. Fig. 5008. Galium andrewsti A. Gray, Proc. Amer. Acad. 6: 537. 1865. Plan pork and usually densely matted, grayish green, with branching, slender, woody roots, te stem uch bra anched ue ag ely leafy from tangled mats or tufts 4-12 cm. high, glabrous or oft sia pt scabro the a i hd Leaves persistent in whorls of 4, crowded, 10 mm. long, linear- ae rigid. ae stueely pungent, grayish green, sometimes spinulose- ciliate on the thickened margins ; yrs = flowers borne on the e upper branchlets in small clusters or singly, on capillary pedicels as Ss or somewhat surpassing as leaves ; ang flowers solitary, axillary on short pedicels; yet about 2.5 mm. wide, greenish white, e lobes acute; fruit 2-3 mm. broad, glabro en th on ert spreading or slightly id ai pedicels, bee like when ripe and blackish in ited spec Dry stony ridges, especially in eye ral yellow pine belts, Upper Sonoran and Arid Transition Zones: in the Sierra Nevada foothills and py pda ea Casat Ranges from phe County to San Diego County , and adjacent Lower California. Type locality: ‘California, Dr. Andre May-Aug. 19. Galium ambiguum Wight. Yolla Bolly Bedstraw. Fig. 5009. Galium ambiguum Wight, Zoe 5:55. 1900 Low tufted perennial from a ores woody root, with branching underground stems fro: go ude tufted, leafy, oe stems 5-10 cm. long, rarely branched, the whole plant grayish with reading hirsute pubescence. ‘Le € 5-12 mm. long, 1-2 mm. ‘wide, linear, acute, ascending, mi surpassing the inter mae and ge the stem feces on young growth, firm but no at "all rigid, midrib prominent beneath and the margins somewhat inrolled, more or less bristly- hirsute on both subtaces, short 4 aiey branchlets often sarah ent at the internodes ; inflorescence polygamo-dioecious (?), flow Reel solitary in ns eee of the upper leaves or terminal in small, short-peduncled cymes ; sctelit ¢ ish white, ut m. broad, the lobes teat sparsely — a a — Poteiss or hcieurhat hairy, blackish i in dried speci places, Arid Transition a9 aren Mountains in Jos ink oe ounty, Oregon, sie ‘o een “facodng Ccitornia also reas Flt Wi Lake County, California. Type locality: “Yolo Bolo, Yolo Co., Cal., T. S. Brandegee, Sep rch— Sept. ambiguum var. Caren lees Penta Dudley Herb. 4: fig. 955. Resembling the name beeing taxon in habit but often — Lege (ex xcept rol exposed sirtbtione), ‘the a, eg up to 16 cm. long the longer stem es branched al leaves 6-10 m long, like the species in shape and texture, dark green, shining, glabrous or with a few sanseeced. teh ciliate ‘hairs. Siskiyou Mountains of Curry and Josephine Counties, Orasen, bows to Del Norte and Si skiyou Counties and the mountains of Trinity and ot aera Counties, California. Type locality: near Gasquet, north side of Middle Fork of Smith River, Del Norte Count 20. Galium cleméntis Eastw. Santa Lucia Bedstraw. Fig. 5010. Galium clementis Eastw. Leaflets West. Bot. 1: 56. 1933. Low, tufted or densely . perennial with stems a 3.5-8 cm. long, the stems and leaves oak aioe beset with es hirsute hairs. Leaves in whorls of 4, surpassing e internodes except on young s g p of vg strongly inrolled stemlgit 3-6 m sre 1-2 mm. wide; flowers yellowish, few in the upper axils or in small, shor t-peduncled « cymes, the ee sparsely ad ee wilisiot: fruit fleshy, sotatrtat hairy, blackish in dried specim Rocky outcrops, Arid T ition Zone; it aia Cone Peak and Santa Lee Peak, Santa Lucia Moun- tains, Monterey County, California, Type locality : Santa Lucia Peak. March—May MADDER FAMILY 35 21. Galium californicum Hook. & Arn. California Bedstraw. Fig. 5011. Galium californicum Hook. & Arn. Bot oe 349. 1838. Galium mislertieegee A ages nso Nutt. ex & Gray, Fl. N. Amer. 2: 20. 1841. Galium flaccidum Greene, Pit : 34. Galium pal ear MeClitchic, eae Pi 124. 1894. Perennial with slender, branching, woody rootstock, the stems erect, forming low tufts 8-15 cm. high more diffuse and often 30 cm. high, slender, grayish-hirsute throughout. rit es in - €0 ; e lo rather abruptly apiculate-acuminate, _ mm. long, thin, hirsute with sab pron 3 hairs on both’s sur- faces or oe ons on the margins ; flowers di eerie solitary and axillar the staminate ones in twos or t short axillary pedicels ; "eproili ellowish, about 22.8 elt, broad; fruit small, fleshy: whitish- tpariatisceet igicte ies dark when yt glabrous or hairy Open woods or shaded rock Upper Sonoran and prensition Zones; a uthw n California through the Coast Ranges from Humbo dt sien rag the San Gabriel Mountains, Los Angeles Co feud M — on = pos Cruz Island, Santa SBachats County, and locally in the central Sie Racy "Nevada in hana County; ich | ommon per Santa 4 Ps County southward. Type locality : Califor, Collected by Douglas grelisdty in ‘the ‘victaily of Monterey. Aly. ccasional ie are found having the typical leaf-shape and habit of G. californicum but having the stout scabrous hairs on margin and midrib typcal re vel nuttallii instead of the punting -hirsute pubescence of G. cali- fornicum, Other specimens trond rocky ridges of San Fra o, San agg a nd) onterey Roeder sa and south- ward have the crowded and thickened e gast a var. Pek wie eh, but with the specimens mentioned above, the scabrous marginal trichomes of G. tallit. They appear to ‘be transitional between, G, call ovairom and G, nuttallti, as has been suggested by Hilend and Howell ae ts Bot. 1935). Galium cellornicuss Bd ok migpeltose. Screens) Jepson, Man. a ~ Calif. a “ 1928, (Galium miguelense pie oa Bag a zs 887.) Stems m or Scipetienes elongated; leaves crowded, oblong to 2 ta “ovate, char in tex biog lek ike or with | rie hairs on the margins: fruit white. San Miguel and Sar R ldeaie Satta Barbara. County, California. Type locality: San Miguel Tslan 5009. Galium ambigu : 5011. Gali ali 5010. Salis oem $012. Galium murciatum 36 RUBIACEAE = Galium muricatum Wight. Humboldt Bedstraw. Fig. 5012. Gali Wight, seh oi 1900. ?Galium chartaceum Wight, 1 Diffuse or a tated ‘giro perennial with at aig stems 3-12 cm. long, the stems datieon: sew shining. Leave whorls of 4, sprea py sh than on ee oe or Be aciges them in the more tufted thay 5-9 mm. long, firm re, “a rk g shining, n wly to broadly elliptic and sometimes obovate, short- aking Fo margins Papa ciliate with aici ascending hairs, the upper surface sparsely short-hispid with appressed hairs to glabrous, the lower surface glabrous or with occasional hispid hairs on the midvein; inflorescence axillary or terminal with slender pedicels, in small clusters of 2 or 3 flowers, little surpassing the leaves; corolla th cpboeny white, 1.5-2 mm. wide; fruit glabrous, black in herbarium specimens, fully mature fruit esl or ss slopes, Arid Transition Zone; Agness, Curry County, Oregon (Peck, Henderson), an Bien and Mendocino Counties, California. Type locali ity: “Westport, Mendocino County.’”’ June-July. Much Lisanti the more tufted forms of G. californicum and apparently some forms intergrading with that species. 23. Galium nuttallii A. Gray. Climbing Bedstraw. Fig. 5013. Galium suffruticosum Nutt. in Torr. & Gray, Fl. N. Amer. 2: 21. 1841. Not Hook. & Arn. 1833. Galium nuttallit A. Gray, Smiths. Contr. 35: "80. 852 Dioecious perennial with 1 or more slender stems arising from a woody root, pie below and elongate with long internodes, reclining or climbing on shrubs and rocks, 1.5-2 m length, coikpacts much branched above and leafy, the seasonal stems ge — ches bertmcceoes, more or less retrorse-scaberulous. Leaves in wher s of 4, usually rat becoming thicker in age and yp a ran eee: ees ovate: ee ng 2 broadly linear. anal . a rrowly linear-oblon n linear in s «fot ms), arely obtuse, the u urface = caaeaaiy hispid. “s gittinars he margins inroled a. age Caitty to denaely sabres with ually retrorse hairs; staminate inflor n few-flowered cymules in gl hg ie sineitate inflorescence solita from jet axils of ‘tea aie wie pedicels 2-4 m ved rolla inant i yellow, 2-2.5 mm. broa obes narrowly ovate- Eo aate:. frui pent g eter, often 5 1 aescl Gerelonhig pearly white when mature, tert tinged with lavender turning black w n dry Shrubby hillsides, Sonoran falc vate n and LS ag i Oregon, south through the Inner and Outer Coast Ranges to northern Lower California and from Siskiyou and Shasta Counties south along the Sierra Nevada foothills. Type locality: San Diego, California. MarcicAce. " [ators yades with G. iolendork are frequently found in its gabe limits and along the Sierra Nevada foothills. sulare Ferris, Contr. Dudley oe th. . figs. 2a—-c. 1955. Habit as in the thick in texture and bearing a cusp. Santa Rosa, Santa fae a d Santa Catalin Islands, California. Type cality: Santa Catlina Island, Los Angeles County. 24. Galium bolanderi A. Gray. Bolander’s Bedstraw. Fig. 5014. Galium bolanderi A. Gray, onl tal 29. Galium gabrielénse Munz & ae Old Baldy Bedstraw. Fig. 5019. Galium gabrielense Munz & Jtn. Bull. Torrey Club 51: Galium siccatum var. anotinum Jepson, Man. FI. Pl. ou 962. 1925. Dioecious sah oak gee fe a woody base, stems crowded, much branched and tufted, some- what woody at base, 5-1 high, oo pveaaaee see with a short, spreading, hirsute pubes- cence. Leaves Al wild of 4 eer 5-10 mm. lo ane, acute but not cuspidate or only obscurely so, crowded ; ariel in short in the axils e upper leaves ; Rieke a yellowish, 3 mm. broad, the lobes h ry on the pateides tr fruit including he hairs about 3 mm. wide. Dark rocky ridges and slopes, Arid Transition and Canadian Zones: mountains of sonthes oes oie in Los Angeles, aeeie and San Bernardino Counties. Intergrades with = lium angustifolium. Type locality: “‘ridges east of Ontario Peak, alt. 8400 ft.,”’ Sai Bernardino County. June-Aug MADDER FAMILY 39 30. Galium hallii Munz & Jtn. Hall’s Bedstraw. Fig. 5020. Galium hallitt Munz & Jtn. Bull. Torrey Club 49: 358. 1923. Dioecious suffrutescent perennial, woody at base, ae gprkiggd a decumbent or ascending, quadrangular stems 3-4 dm. long, leafy above, the y irsu an “eas older stems glabrate with thin, white, pefolating ere avis in whorls of > much shorter than the erodes on the lower stems, two to three times onislin above, aan green om ong, usually curved to o side corolla yellowish, about —-3 mm. broad, sparsely hirsute on outer surface; = black, slightly j juicy, 3 mm. long, densely covered with srtieg Hite white hairs about 2 mm. lon Dry flats and slopes, sn Transition Zone; cauthern Sierra Nevada, California, Kern cause, adjacent Tu- a County, oe api Mountains, and the Mount Pin southward to the San’ Gabriel and San yg oad Type locality: “Coldwater Fork of eh my Cok, iad Gabriel Mountains, in gravelly ground at 5 3700 rere pi Pa ty ” June—Aug. 31. Galium stellatum se eremicum t Hilend & hee Ehrendorfer. Desert Bedstraw. Fig. 5021 Galium stellatum var. eremicum Hilend & Howell, Leaflets West. Bot. 1: Ayres veh Galium stellatum subsp. eremicum Ehrendorfer, Contr. Dudley Herb. 5 oecious low shrub dm. high, intricately bratithied erage sometimes on young growth, fe toward the pice the stems light gray and shreddy, stramineous above with exfoliating 5019. Galium gabrielense 5017. Galium wrightii 5018. Galium angustifolium 5020. Galium 40 RUBIACEAE bark, the ultimate branches strongly 4- — the angles white, more or less granu pe and densely puberulent with hispidulous hai Leaves 4-7 mm. long, in whorls of 4, those of vigor- ous shoots longer, iced sora og to bok: lanceolate, rigid, i a Sapa rev aa acumi- nate-cuspidate, pale gra pape eo cabrous puberulence, midvein white minent, the upper reduced leaves Ecker node ; hecaetice atts muniate and Pistillate many- flowered, of crowded diffuse “panicles, leafy, the le sue some sia Abs ced in size, ey fruiting pedicels s straight usually 2-3 mm. long; corolla greenish yel 2.5-3.5 mi . broad, the lobes ovate, ci outer surface with enticed ieee eins ia Ticholing the copious Gin hairs 2.5- SD wide. Fee slopes, Sonoran Zones; common in Inyo County, California, and southern dibs and Utah one gua Pee pi flamers the Mojave and Co lorado Deserts to western Arizona and Sonor ra. Type locality: western end 0 the Sheep Hole Mewdns San alifornia. March— 32. Galium matthéwsii A. Gray. Matthews’ Bedstraw. Fig. 5022. Gali: tthewsit A. Gray, Proc. Amer, Acad. 19: 80. 1883. Suffrutescent, glabrous, much-branched perennial 2.5-3.5 dm. high, the stems rather slender and persisting more than one season, the bark whitish and shining, ppl ee in age. Leaves in whorls of 4. much shorter than the internodes, lanceolate-ovate, 4-8 mm. long, cuspidate-acute, l-nerved. rigid, bag green, the upper smaller; inflorescence ample ane open, many-flow sen the branches of the flower-clu sters — rather long, and the subtending leaves muc ie oe pedicels tem straight in fruit; coro ila greenish white, 1-2 mm. wide, the lobes mae tu acuminate, the outer surface gatarty covered sith eee “to hairs ; teal small, 2-3 mm broad "nelad ng the dense covering of short white hai slopes of desert ranges, Upper Sonoran uae en mee Zones; eastern face of the Sierra Nevada ; ae : Valley region and also the Kingston paar San Bernardino County. Type locality: “Arid district in a Co., California,” but according to label on type specimen in the Gray Herbarium collected ‘‘among sagebrush n Camp Independence. oe May-July. 33. Galium mitnzii Hilend & Howell. Munz’s Bedstraw. Fig. 5023. Galium munzti Hilend & cates —— West. Bot. 1: 135. 1934. Galium munzti var. carneum Hilend & Howell, op. cit. 136 Galium munzi £. glabrum Ehrendorer aay Dudley Herb. 5:9. 56. Ste from a branching, somewhat woody fe erect or Sree! ape ds oi 3.5(5) ae high, ase and pallid, ect reap with sprea ofine hairs or som s gla bro Leaves i whorls of 4, much shorter than the internodes, usually broadly ovate: PO 3-12 mm. ae dark green, tapering to an acute, se, or less apiculate and mucronulate apex, rather thin, and thinly hispidulous on both sides, midvein prominent, white, the 2 latefal veins short and slender or Bena) oO the —— often sparsely scaberulous ; yo sia pyramidal, ma n ered and rather ope h e hivmcciee branchlets ddivaricate fruitiie pedicels mm. long, 'straig ‘it | and aricate; corolla mm. broad, greenish or occasionally reddish, hispid exteriorly ; “truit nies the spread- ing ea 3-4.5 mm dors rocky or gravelly slopes, Upper Sonoran and Arid Transition Zones; desert ranges of Inyo County inclu diss the west face of the southern Sierra Nevada, and adjacent Nevada, south to the San Bernardino Moun- tains pe east te malthresters: Arizona and southern Utah. Type locality : “Bonanza King Mi ine, 3000 feet, east slope i the Providence Mountains, San Bernardino County, California.’’ June—Sept. ium munzit is quite variable as to leaf-shape, and especially as to pubescence in its more easterly range where f. glabrum is sometimes collected. 34. Galium multiflérum Kell. Many-flowered Bedstraw. Fig. 5024. Galium Se Kell. Proc. Calif. Acad. 2: 96. fig. 27. 1863. Galium bloomert A. Gray, Proc. Amer. Acad. 6: 538. 1865. Perennial, stems erect fro woody ock, often somewhat suffrutescent below with the epide rmal gil: extfiatng in ‘tes ai shreds, the leaf-bearing branches numerous, rather s Sy 1.5-2.5(3) dm. hi labrou p wh rarely oo scabro _ Leave es in sraeashen of x ga ihe spreading, broadly ovate to ovate-lanceo- ate. 0 mm inin r idvei he more ne € m t pistillate inflorescences Reha many-flowered, the branches sprea ing, definitely longer than he leaves, = branchlets divaricate, the fruiting pedicels 8 mm. long, div a corolla gisprow: s, greenish yellow, 2-2.5 mm. wide; fruit 4-6 mm, wide including the long, spread- air Dry rocky gee and talus slopes, mainly Arid Transition Zone; Nevada west to Modoc County, California, south to Inyo County and east to Utah. Type locality: “vicinity of Virginia City, Washoe,’ Nevada. June—Aug. Galium multiflorum f. hirsttum (A. Gray) ero nee ee Contr. Dudley Herb. 5: 10. 5956. ‘ee alium bloomeri var. hirsutum A. Gray, Bot. Calif. 1: tek 876; G. pia ar var. scabridum Jepson, Fis Calif. 963. 1925.) Differing from the preceding only Peas the presence of longish, spreading, hirsute og on eae and leaves. From the type locality south to seal x ote and a anesnt Nevada. Type locality: Sierra Valley, Ghen County, California. Collected by Lemm 35. Galium hypotrichium A. Gray. Alpine Bedstraw. Fig. 5025. Galium hypotrichium A. Gray, Proc. Amer. Acad. 6: 538. 1865. Herbaceous perennial, the slender woody rootstocks becoming much branched, the stems and MADDER FAMILY 41 branches numerous, forming low tufts or mats 5-10 cm. high, very leafy, the internodes mostly pro than the leav es, slender, gee te puberulent, with very short divergent hairs. Leaves 3-6(10- mm. long, 2-4 mm. bro ad, ather thin, ovate to ovate-lanceolate, acute, l-nerved, green but with a short, s he hu ge ‘paber ulence, ‘markedly so on the margins; inflorescence nar- / in ong, cu yellow, sometimes reddi sh, "aE lobes about 1.5 mm. long; fruit densely covered with whitish or very faint ly tawny bristles longer than sie} body Rocky alpine summits, Hudsonian and Arc fps Zones; central Sierra g's from the type wager d mne County California, south to pay Seats ‘oles it ineererees with G, hypotrichium subsp. d to the White Mountains of. perce and Nevada and to the Wassu k and Toyabe Bre of central Nevada where intergrades are found with other s subspecies of the hypotrichium ner yeep Type locality: ‘Sonora Pass, in the Sierra Revadks, alt. 8000 to 3000 feet in dry and peg places.”” July—A Bk rad taxon he hypotrichium (including all of the subspecies) is one of the more variable enits of the G. muiti- flor mplex. It is probably most closely related to G. munzii from which it differs in the narrower inflorescence rie the § frititing widiecs which. re short, tend. p be nodding. Its distribution eek a es a somewhat interme- ae postion between that of G. mu ltiflorum a G. munzii though it is found at higher elevations than either eae hypotrichium subsp. subalpinum (Hilend & Howell) Ehrendorfer, Contr. Dudley Herb. 1956. (Galinm munzit var. subalpinum Hilend & Howell, Leaflets West. Bot. 1: 136. 1934.) abit ays in ie po- precedi ing but differing hi the ovate leaves; the pubescence of stems and leaves a oi ag longer than that y trichium corse foe! pride ane At high clevations in ree Sierra Nevada, Califor from the s Bathern border of Mon ia ag unty e County; also the White Mountains in Nevada. ee locality: Sawmill Canyon, Inyo Galium hypotrichium subsp. tomentéllum Ehrendorfer, Contr. Dudley H 956. Stems about h, more tufted than subsp. bec eaeenaag or subsp. subalpinum, roy ea belly Be white-tomentulose; i in i type locality, Telescope Peak, Pana- rom Boundary Peak in the White tions of subsp. subaiolacen 5023 5023. Galium 5021. Galium stellatum munzii 5024. Galium multiflorum 5022. Galum matthewsii 42 CAPRIFOLIACEAE 36. Galium parishii Hilend & Howell. Parish’s Bedstraw. Fig. 5026. Galium multiforum parvifolium grin. Zoe 5: Galium parvifolium Jepson, Man. FI. Pl. Calif. 963. 1925, aus lige 1800. Galium parishii Hilend & Howell, Sere West. Bot. 1: 136. Perennial, with woody branching rootstocks, oo stems s slender, or pat many, tufted or matted, 5-20 cm. m. high, ascendi sid or fie rye t decumbent at base, simple with asc anid branchlets, light gray-green, scabrous on the angles and m et rou nt sh- baentcne betw Leaves in whorls of 4, often n approximate near the base ide clothing the stems, tardily dscldoce. and smaller than the more distant ones on the upper — ore acute, often broadly so, abruptly mucronate t 4 mm e, a ides ; i a , about Z— m. Sabian narrow and elongate, in small pe ha feces i. the leaf-axils on upper part of stems, the monoecious ; corolla 1-2 mm. wide, reddish, rarely greenish, sparsely puberulent exteriorly ; badly of fruit about 1 m m. broad, densely covered with spreading hairs about 1.5 mm. cote slopes and flats, abe Transition and Canadian Zones; es Antonio, San Bernardino, and San Jacinto Mountains, southern California, east to the New York Moutains, Califo ornia, — the Charleston MMacuthien; south- ern Nevada. Type locality: Bear Valley, San Bernardino Mountains. July—A 37. Galium grayanum Ehrendorfer. Gray’s Bedstraw. Fig. 5027. Galium grayanum Ehrendorfer, Contr. Pankey Herb. 5: 15. 1956. arising 2 ‘from a oody ck, stems greenish, inconspicuous sine and mt thed as Be gee leav e, < ett se rt, even puberulence. Lea m g, m. wide, a little : ess than sb “half the length = ‘the srt a to nearly evaling = 3 nerved, ‘thickish wok dark green, ovate to n Psion seem obtuse to roun ea : ; infloresc — narro wr es ae floweri ing bra comin ng, leafy, padiaits little od sae the leaves, the pall rs ie congested, the Editing peiticels ane flowers mm. broad, widtish, stten tinged with urple; lise including the copious white hairs 7-9 mm. broad. Rocky slopes, Canadian and he atin in the Coast Ranges of California from Siskiyou and Trinity Counties Me to Lake County and to the east from Butte County to Placer County. Type locality: Jonesville, Butte cai July—Aug Galium grayanum “subsp, Siaveenerps Ehrendorfer, Conte. Duds ey sabe 5: 15. 1956. Like G. grayanum subsp. papier in habit and shape of leaves but often up to 2.5 dm. in height; nearly glabrous or sparsely pubescent with somewhat longer hairs (hispidulous hairs aa present) ns nearly glabrous; inflorescence as in the preceding but fruiting pedicels often 5 mm. or more long and usually somewhat i corres downward. nag a er Lake County, Oregon; also Modoc vagy nd California, where intergrading forms, with °G. multifiorum and C. wat- e found. T ountains, Siskiyou County, California. 38. Galium watsonii (A. Gray) Heller. Watson’s Galium. Fig. 5028. ltifl var. watsoni A, Gray, sedate as a Amer. 12: 40. 1884, Galium watioals Heller, Bull. Torrey Club 2 898. Galium watsonii f. scabridum Ehrendorfer, ree Seite Herb. 5: 16. 1956. Herbaceous, somewhat cespitose perennial (sometimes woody at base), with several erect — usually 1-2.5 dm. high from a woody rootstock, the eae pie ssid throughout (long cabro airs in f. scabridum) and ssuklty brownish green in dried specimens. Le in whorls of 4, ne thin, 5-15 mm. long, 1-3 mm. wide, linear-oblong or linear-oblanceolate, the midvein prominent ; branches of the inflorescence usually not much surpassing the leaves, erect or ascend- ing, sone mm. wide, greenish yellow; fruit densely fice 3 with long spreading hairs, 6-8 mm. nee - Transition and Canadian Zones; east of the Cascade Mountains Ho Chelan County, Wash- Gal Secu subsp. cobhrulans Ehrendorfer, Contr. Dudley Herb. s: 4 1956. (Galium multiforum subsp, vaberichen Piper, Contr. U.S. Nat. Herb. 11: 527. 1906; G. sehen? var. puberulum St. John, North- west Sci. 23: 90. 1928.) Very ary fos G. watsontt subsp. watsonii in “E30 it, but the plants more or less densely perciows throughout. Dry slopes erpentine, ‘Arid Transition and nadan patos common in Chelan and Kittitas Counties, Washington. Type | locality: Ellensburg, Kittitas Counts, Family 144. CAPRIFOLIACEAE. side sik dieing FAMILY. ovary, its limb 3—5-toothed or 3-5-lobed. Corolla 5-lobed, someties 2-lipped. Sta- mens 5 or rarely 4, inserted on the corolla-tube and alternate with the lobes; anthers versatile, 2-celled, longitudinally dehiscent. Ovary inferior, t eeclieds style ‘slender ; stigma capitate or 2-5-lobed ; ovules 1 to several in each cavity. Fruit a berry, drupe, or capsule. HONEYSUCKLE FAMILY 43 ‘ on of 11 genera and about 350 species, mostly in the northern hemisphere, a few in South America and ustralia Corolla rotate or urn-shaped; flowers in compound cymes; shrubs or trees. Leaves pin eeded. ; drupe 3- 5-s 1. Sambucus. Lea ee - upe l-seeded 2. Viburnum. Corolla ‘ivcdas or rainiva neti: dittn 2-lipped. Creeping evergreen herb; flower «geminate, on Jong seit peduncles 3. Linnaea. Shrubs or woody twining vines; flower ong deaioc peduncles. Corolla campanulate, short, sit or nearly so; til shit 4. Symphoricarpus. Corolla tubular or campanulate, more or less irregular 5. Lonicera. 1. SAMBUCUS [Tourn.] L. Sp. Pl. 269. 1753. Shrubs or trees with opposite pinnate leaves, serrate or laciniate leaflets, and — white, yellowish, or pinkish flowers in compound, depressed or thyrsoid cymes. Caly tube ovoid or tur inate, 3-5-toothed or 3—-5-lobed. ‘Corolla rotate or slightly canine, regular, usually 5-lobed, the lobes imbricate or valvate. Stamens 5, inserted at the base of the corolla; filaments slender. Ovary 3-5-celled; style 3-parted; ovules 1 in each cell, pendulous. Fruit a berry-like drupe, with 3-5 one-seeded nutlets. [Latin name of the 5025. Galium hypotrichium 5027. Galium grayanum 5026. Galium parishii 5028. Galium watsonii 44 CAPRIFOLIACEAE About 20 species of wide geographical distribution, about half of them occurring in North America. Type species, Sambucus nigra L. Inflorescence a flat-topped, compound cyme; ripe berries with white evanescent bloom Many-stemmed shrubs; cymes 12-30 cm. broad; leaflets usually gradually erinainete ane gnatkedly asym- metrical at base. L, Se Usually arborescent; cymes 7-15 cm. broad; leaflets typically abruptly acuminate and pi righety to not at all asymmetrical at base. 2. S. mexicana. Inflorescence pyramidal or dome-shaped in outline; berries without bloom. Ripe berries black. 3. S. melanocarpa. Ripe berries bright red, rarely chestnut or yellow Leaflets closely serrate to the apex, more or desi pubescent beneath. . S. pubens arborescens. Leaflets coarsely and sharply serrate except at the apex, mostly glabrous beneath, E . S. microbotrys. 1. Sambucus caerulea Raf. Blue Elderberry. Fig. 5029. Sambucus caerulea (cerulea) Raf. Alsog. Amer. 48. 1838. Sambucus glauca Nutt. in Torr. & Gray, Fl. N. Amer. 2: 13. 1841. F UCU. 1872 D : bE Sambucus caerulea var. pias Schwerin, Deuts. Dendr. Ges. 1909: 37, bee 1909. Sambucus decipiens M. E. Jon gee" Univ. Mont. Biol. Ser. 15: 46. 1910. Sana ferax A. Nels. Bot. psig : 225. 1912 nentell a Heller ex ae Deuts. Rashi: Ges. 1920: 218. 1920, in synonymy. Shrub of clustered erect stems 3-7 m. high sprouting freely from the base, the panei stems usually a few centi ni in diameter, the young stems and branches brown and of laucous. Leaves 535 dm. long, the pe etioles 4-8 cm. lo ong and often eae nea (teaves of vigorous stems up to 6 dm. eit ae nd not infrequently bipinn ate) ; leaflets 5-9, green above and paler be- neath, rather thick, 6-16 cm. long, often long- sence ta Sects fascias to oblong-lanceolate taperin i base and typically s = metri dis. isappear under moist con Woods and thickets, Sanne and Canadian Zones; southern British Columbia southwa rd west and east of the Castade Siouneaies to the Siskiyou region of northw western California, and to northeastern California southward ) ierra Nevada t i ¥ unt Pi r ' o Alberta through the Sier o the mountains of San Bernardino County and the Moun os region; als rt and Montana south to northern es and New Mexico, In the Sierra Nevada and the desert ranges eastward in Nevada, many specimens are with a close velvety tomentum on the petioles and undersurfaces of the leaves. Gl abrous but similar “plants. ecu in the same localities. Type locality: ‘near Oregon Mts.” Collected it. Sambucus caerulea var. Aekehexicins Rehd. Deuts. Dendr. Ges, 1915: 228. 1915. (Sambucus neomexi- cana Wooton, Bull. q otte Club 25: 309. 1898; S. intermedia var. ne valle tcana Schwerin, Deuts. Dendr. Ges. 1909: 38, 328. 1909; S. glauca neomexicana A. Nels. in Coult. Nels. New Man. Bot. Rocky Mts. 469. 1909; S. caerulea 3. trifida Schwerin, Deuts. sgh ag 1920: 218. 1920; S. trifida Heller = gouge loc. cit. in synonymy.) Habit of S. caerulea var. caeru leaflets 3— 5(), about 3-8 cm. lon ng, m. wide, in ours oblong-lanceolate, serrate with rather small teeth, thickish, pale green; inflorescence ra Fm. canyons or among boulders, Arid Transition Zone; Mésenie Peak, Mono County, fad ‘the bidetadg go of Inyo County, California, and in Pe a Nevada, New Mexico, and Arizona and also Sonora ate Foe Ruidoso Crossing, White Mountains, New Mex 2. Sambucus mexicana Presl ex DC. Southwestern or Desert Elderberry. Fig. 5030. Sambucus mexicana Presl ex DC.. Prod. 4: 322. 1830. Sambucus velutina Dur. & Hilg. Journ. Acad. Phila. I. . 439; 1855. Sambucu Lachapade iS Var. Mexicana Salk: Silva 5: 88. pl. 893. Sambucus caerulea var. glauca Schwerin, Deuts. Dendr. Ges 1909: 37, 328. 1909, in part. raepaectvat canrales. var. vehatiaa a loc. o . inp e, Leaflets Bot. Obs. 2: 99. poeek ambucus coriacea Greene, loc. cit. ?Sambucus fimbriata Greene, ioe. ett. aerate suennren So Schwerin, or a Ges. 1920: 218. 1920. ata var. erula Schwerin, a pebatwy orbiculata var. abe [od =: cit. . arizonica Sarg, Man. Trees N. Amer. ed. 2. 885. 1922. sambaces glauca v Meds arieonticn 5 Sawe ex Jepson, Man. FI. Pl. Calif. 965. 1925. , not Nutt. hosris cueeuled var. mevicann ©: Benson, Amer. Journ. Bot. 30: 240. 1943. m. high with 1 o e trunks having furrowed bark (these 15-45 c in diameter), and also areal smaller io ieiaie from base, the smaller branches gla s omentulose. Leaflets 5-7, less often 3, oval, te-lanceolate, or oblong, abruptly acuminate, times cuspidate, and or cuneate at base, usually not —— ped Soe gases Taage petio- lulate to inf — 2-6 cm. long, coarsely or finely serrate except apex, , green aces, glabrous to densely tomentulose including the piniies and petiolules ; in- florescence a compound, at-topped cyme 5-15 cm. broad, the shiek pee yellow to creamy white ; pe fruit 5-6 mm. in diameter, dark blue or blackish, and when fu lly ripe coecese with a dense whit — eam banks, open flats, and hill slopes, mostly confined to the Sonoran Zones; Sacramento-San Joaquin HONEYSUCKLE FAMILY 45 V owe Po alt = coastal = rene valleys on either side to cismontane southern California and no Low lifornia; o Arizon w Mexico, western Texas, and Mexico. Type locality: Mexico. Aceh. oo ha “a the preceding searie. New mexicana shows much variation throughout its extensive range, and _ local are commonly found. Sambucus orbiculata Greene, unde er which Schwerin descri two varieties, 0 puberula and glabra, representing tomentose and glabrous oc is characterized by orbicular to oval leaflets that are abruptly cuspidate, and are dentate or serrate. This form with broad leaflets occurs rather frequently rd t from the San Francisco Bay region southwa o Santa Barbara County and the Channel Islands. Plants with ovate- lanceglate leaflets, often cadnates based and jig ‘pubescent to glabrous, which are considered to be typical of S. mexicana, occur just as frequently in this ar Th ioeatien at which S. velutina was originally foun 4 is s Posé eek [Poso Creek]. According to the itinerary given in the fifth volume of the report on the exploration for a Pacific railroad route, the Ocoya or Posé Creek depot camp seven miles north of the Kern River made in a flat along the stream in the lower foothills where the vegetation contrasted with “the “eis sad vat sched surface of a ape ota hills, which ba withow ae or any green vegetation.”’ This bed tomentose-leaved — of S. mexicana occurs rather mmonly n Joaquin Valley, California, and is not to be confused w the ‘nularanenn: leaved forms of S pies A ee Cs hishe? clemenbicias | in the ainccak Steere} Nevada. 3. Sambucus ey A. ay: Black Elderberry. Fig. 5031. Sambucus melanocarpa A. Gray, Proc. Amer. Acad. 19: 1883. Sambucus melanoc mmunis Schwerin, maj Deni Ges. 1909: 43, 328. 1909. Sambucus melanocarpa fiirstenbergti Schwerin, lo ne Sambucus racemosa var. melanocarpa Me Min “Ti Man, Calif, Shrubs 529, 1939. Shrub 1-3 m. high with smooth, erect, glabrous stems. Leaflets usually 5-7, dark green above Ste eta Pe, ‘ 5031. Sambucus oe 5029. Sambucus caerulea 5032. Sambucus p 5030. Sambucus mexicana 46 CAPRIFOLIACEAE and paler mice ie thin in texture, the younger leaflets —— short-pubescent becoming e, 8-15 glabrate in a . long, one-half to one- spoke as broad, rather narrowly ovate- lanceolate, asd abrantly t aie a acuminate apex, rounded or cuneate at base aid usually Shleane. the oe coarsely and rather unevenly serrate; inflorescence Satay about 5-7 cm. high, 6-10 cm. broad, pei in outline ee openly bran che d; flowers cr white, da eam on drying corolla- era oval to oblong e fruit globose, about ay in pag black, w ithout bloo lakes ae Pista mainly in the Canadia n Zone; British Columbia _— in the se ‘cn and Blue Moun- tai Ara n and Washington and occurring rarely in the Siskiyou region and the Sierra Nevada of California as potty south as Olancha Peak, Tulare County; also Idaho and el see ak ro northern Arizona and New Mex ico. Type locality: New Mexico. Collected by Fendler. June—Aug 4. Sambucus pubens var. arboréscens Torr. & Gray. Coast Red Elderberry. Fig. 5032. Sambucus pubens y arborescens Torr. & Gray, Fl. N. Amer. 2: 13. Sambucus racemosa var. arborescens Torr. & Gray ex A. Gray, Syn. FI. N, Amer. 17: 8. 1884. Sambucus callicarpa Greene, Fl. Fran. 342. 1892 moucus marit Sambucus leiosperma Leiberg, Proc. Biol. Soe. Wish. il: a Piri Sambucus racemosa var. maritima Jepson, School FI. 85. Sambucus racemosa var. callicarpa — Fl. W. Mid. Ae on" 1901. rub 2-5 m. high with smoo h bark, Pg a 5- pis! ay igic oe hig a lanceolate to oblong- ovate or sometimes oblan Eee acum 15 cm. long, ot n finely and sharply serrate to the apex, the teeth usually inc Giered,: paler beneati and in re or ape ess pu abe scent; inflorescence commonly broadly ovoid and dens e, the flowers cream- colored berries bright red or rarely chest- nut brown or yellowish, about 4 sod in niger sh ce without om. Stream banks and mud flats, ~aaee near the coast, Humid Transition and Canadi ian Zones; southern Alaska Pies Spe west of the Cascade Mountains to southern’ Oregon, where it alt to occurs in a Crater Lake r egion, and along coastal California to San “een County. Type locality : not given. April—-Jun 5. Sambucus microbétrys Rydb. Mountain Red Elderberry, Fig. 5033. oe ee Rydb. Bull. ig 2 Club 28: 503. 1901. Sam nata Greene, Leaflet t. Obs. 2: 100. 1910. pe S$ racemosa var. microbotrys ge & Peeb. Journ. Wash. Acad. 29: 492. 1939, shrub 0.5-1.5(2) m. high, Be stems and herbage glabrous throughout. Leaves 5-7-folio- late, the leaflets 6-12 cm. long, ovate to ovate-lanceolate, acute or acuminate at apex, rounded at base and equal or slightly eeetrcal sessile or short-petiolulat, coarsely serrate ot at the rather short- a tip, pale green; inflorescence ovoi road at the base and about as high, dense, the flowers cream-colored to pale yellow; fruit bright red, 4-5 mm. in diameter? out ‘aeons Rocky sinned mostly Canadian Zone; Sierra Nevada and San Bernardino Mountains, ioonak ictag east to Colorado, Utah, and Arizona. Type locality : Pikes Peak, Colored: Collected by Bessey. May-July 2. VIBURNUM [Tourn.] L. Sp. Pl. 267. 1753. Shrubs or trees with toothed or lobed leaves, and white or pink flowers in compoun cymes, the outer flowers sometimes radiate and sterile. Calyx with an ovoid or turbinate tube and short, 5-toothed limb. Corolla rotate or deeply fences Hepes regular or the —— slightly aie 5-lobed. Stamens 5. i orolla-tube; anthers ong, exserted. Ovary 1-3-celled; style short, 3-lobed or 3- -parted ; Sales solitary i in each cell, pendulous. Frui & drape, with an ovoid, glo ose, or sometimes flattened stone: endo- sperm fleshy ; cahepe minute. [The ancient Latin n About 100 species of wide pa Samia i the ‘followlhe’ about 20 other species occur in North America. Type species, Viburnum tin =. i? 2) o = is") ou ° 5 . o uit red; leaves more or less 3-lobed, sharply toothed. . edule. Fruit black; leaves crenate-dentate, not lobed. 2. V. ellipticum. 1. Viburnum édule (Michx.) Raf. High Bush-cranberry. Fig. 5034. Viburnum opulus y edule Michx. Fl. Bor. Amer. 1: 180. 1803. m paucifiorum La Pylaie ex Pi. © Griav Fl. N. fe 2: 17. 1841, Shrub 1-3 m. high; branches glabrous. Leaves 4-9 cm. long, 3-5-nerved from the base, broadly oval to orbicular or broa dly obovate, truncate or often subcordate at pom usually rather banicera involucrata (Richards.) Banks. Black Twin-berry. Fig. 5044. X yl i Ss ichards. in Frankl. 1st ae = App. 733. 1823. Loni. a Ba ae mg. Syst. 1: Panes ledebourii Eschsch. Mém. Acad. St. Pitak os ae 1826. Kell. Proc. Catt Acad. 2: 154, al 47. 1862. aon fnviotecr ata Kuntze v. Gen. Pl. 1: 274. 1891. Rcsrineatn penthang Wan. Fl. Pl. Calif. 968. 1925. Erect shrub, ee freely srgpeg bark, 1-3 m. high, with principal branches mostly erect. the branches Lins — brown or purplish, glabrous or sparse a: ae eaves lanceolate to b b nd a i -3 cm. ac rat purple, glabrous or often id gos sano cos ig and ciliate; flowers 2 in the axils of the bractlets ; corolla yellow, 12-15 mm. long, narrowly funnelform, glandular on the outer surface, the lobes nearly soi get and ened sae separate, bla ck. Mois und, Upper Sonoran Zone, and Transition Zones; southern Alaska south along the coast to Santa Barbara Coats. Caltteenia. and in the interior parts of the Pacific States: to the southern roto Nevada, ranging Engage ,te Quebec, Lake Superior, Colorado, and Arizona. Type locality: ‘wooded country from latitude 54° to north.” Canada. March- —July. 3. Lonicera conjugialis Kell. Double Honeysuckle. Fig. 5045. Lonicera conjugialis Kell. Proc. Calif. Acad. 2: 67. ae! ee 1863. Lonicera breweri A. Gray, Proc. Amer. Acad. 6: 537. 1865. ylosteon ea oth Howell, Fl. N.W. Ae: 0% 1900. ic 15 ee high young bran glabrous or ety gee arei he subsessile glands, usually soe Lea ‘mostl o € i é se — Pp esce mm. lon; . lon acts mm. long, bilabiate, the upper lip 4-toothed, fhe eects el gly defle sed. seihiee , the throa t and also the filaments and style ee ny = ite-hairy below; the pairs of ovaries joined about eo tates their length, blac red when r Margins of streams and os Canadian Zone; a and Idaho south along the Cascade and Siskiyou Mountains, and the Sierra age to Tulare Cou nty , California, and western Nevada. Type locality: ‘“Washoe,’” Nevada. Collected by Dr. J. A. Veatch. June— ay. 5 boners utahénsis S. Wats. Utah Honeysuckle. Fig. 5046. Loni: . ats. Bot. King — 133,- 1871. Lonicera sbructilal Rydb. Mem. N.Y. Bot. Gard. 1: 372. 1900. Selene kanes Howell, Fl. N.W. ose 282. 1900. Low shrub with — Specacine branches 6-15 dm. high. Leaves oblong or al e ee te or put obtu unded at apex, obtuse to blunt a se, 2-5 cm. lon , thin th sy Se in pairs on a slender peduncle 1-1.5 cm. long; bracts 2, very narrow, about 1.5 . long; corolla iowa nay 1.5-2 cm. long, white or cals yellow, fading to salmon-yello ow, the base gib- us on e and ia the lobes about — in size and shape but arranged to form a 2- tieoed corolla tim berry red, globose, about 6 mm. in diam meter Stre: anks and bogs a se ier Columbia and Olym Mountains, Washington, south to the Sigoa “Mountains = a eg Californ = Cascade "ionstalas oe Crater Lake, ay on, and - the hee Mountains, northea n Oregon; east es ‘Monta and Utah. Type locality: “Wahsat ountains, Utah, Cottonwood Cajfion; 9,000 feet altitude.” May-Jul 5. Lonicera japonica Thunb. Japanese Honeysuckle. Fig. 5047. Lonicera japonica Thunb. Fl. Japonica 89. 1784. Vigorous, half-evergreen, trailing or climbing vines with stems 2-4 m. in length. Leaves 3-8 cm. long, 1.5-3 cm. wide, ovate to ore ng, tay tage or snatiaae| aebcortatk at base, pubescent, becoming pa above; ong; unc . ee about “a —* of, or ex poe Dr the ppemrs e petiole sabeacdina the. flowers green, leafy, petiolate, shorter than the leaves ; flowers 3-4 cm. long, the tube slender, about equaling the limb, pubescent ft without with HONEYSUCKLE FAMILY 53 long hairs and glandular, white fading to dull yellow, fragrant; style and stamens exceeding the limb ; berries blac A native of Seoul, mat galivated in gardens and occasionally becoming Canpushed in the west and re- ported from Sonoma, nine Santa Barbara, and San Bernardino Counties, Califor common in eastern Un es States. May—Sep r a complete list A ‘synonyms and segregates of L. japonica see Rehder (Rep. Mo. Bot. Gard. 14: 159-162. 6. Lonicera ciliésa (Pursh) Poir. Northwest or Orange Honeysuckle. Fig. 5048. Caprifolium ciliosum Pursh, Fl. Amer. Sept. 1: 160. 1814. Lonicera ciliosa Poir. Encycl. Supp. 5: we 181 ae Caprtfolium occidentale Lindl. Bot. Reg. igh 14 1831. Lonicera ciliosa a youngit Dippel, Handb. ir ie 1889. Stems wd Mabhe branching and usually on fin ing and tw ining, often 5- 4 m. long, glabrous, younger parts . Leaves mostly ellipti casionally ov or obovate, 6—1 long the upper 1-3 an fig Menara branches connate e, e green and shee rous- mabe ‘glaucous beneath and sparsely pilose, ciliate on the rgins; flow sessile, in 1-3 terminal whorls or clust calyx-teeth short-triangular to atnangt gies lop seca reddish yellow or orange, meh Fonnel. 5046. Tt 3, tah > 5047. Lonicera japonica 5044. Lonicera involucrata 5 T = j gialis 54 CAPRIFOLIACEAE form, 2.5-3 cm. long, rather obscurely bilabiate, the lobes ite a dpi 3 ovate; stamens ai style slightly exserted, hairy below; berries red, abou n dia Open woods, often on hillsides and itv, ee Zones; Sites teed py on both sides of the Cailtinde Mountains a the aap and Trinity Mou and Butte — California, and east to Montana and Arizona. Type locality: the banks of the a M. Lewis.” 7. Lonicera etrisca Santi. Etruscan Honeysuckle. Fig. 5049. Lonicera etrusca Santi, Viaggio Montam. 1: 113. pi..1. 1795. Re gorous, a en or half-evergreen vine 2-4 m. long, ours glabrous to the inflorescence. ves 4-9 cm. long, broadly oblong-ovate to oblong or oval, obtuse, — nd not connate sd just ree the Peet glaucous and veiny beneath; inflorescence a terminal pedun- culate head or sometimes with 2 or 3 ax ime ne glandular- -stipitate at sbi shag mes Pubescent the bracts subtending the flowers orbicular, nearly as long as the ovary; flowers 3-5 cm. fed white tinged with eatatsh red, siiccler without, the tube and limb nthe equal ; fruit A European vine sparingly introduced along the coast from Douglas County to Curry County, Oregon, which cage Bae been collected in Del Norte and Humboldt Cn tes California, July—Aug. For omplete list of sroenins and segregates see Rehder (Rep. Mo. Bot. Gard. 14: 194-197. 1903). 8. Lonicera hispidula Dougl. Hairy Honeysuckle. Fig. 5050. Caprifolinm hispidulum Lindl. Bot. Reg. 21: pl. 1761. 1836. _ onice ra aro idula Dougl. ex Lindl. loc. cit., as a synonym. ica To . Amer. 2 184 Lonicera va heahtita Dougl. éx Torr. & Gray, op. cit. 8. Lonicera hispidula var. vaciNans A. ind Bae ee Acad. 8: 628. 1873. 1 y, loc Lonicera hispidula var. californica Rehd.- in Bailey, Cyclop. Hort. 3: 943. 1900. Lonicera catalinensis Millsp. Field Mus. Bot. Ser. 5: 252. 1923 Stems slender, trailing or more frequently ee freely branched, 1-3 m. long, villous- hirsute with long spreading hairs interspersed with shorter ones and also Wich stalked pe varying to watbus. Leaves ovate to ovate-oblong, 3-6 cm. long, often subcordate at base, cous beneath and more or less Sn hae to glabrate, one ie ower leres bite stipule-like fe bees at the base of the short oe, = ke gg orate fe ote glandular-pubescent, oedumpains terminal a xils, iculate; corolla 2- cm. long, reddish eet without, alee ‘within, hese a bilabiate, the ca See Sg gibbous near the base, upper lip 4-lo - d, filaments and style long-exserted, hai a hickets in open and clearings. Hu sey Transition Zone; ps atid Island pe western British Columbia south, west pe gi Csabede- schtadnn. to southern Oregon and southern Canora: also on Santa Cruz and Santa Catalina Islands. Type locality: “woods of North West America.” June— 9. Lonicera interrapta Benth. Chaparral Honeysuckle. Fig. 5051. Lonicera interrupta Benth. Pl. Hartw. 313. 1849. Lonicera hispidula var. interrupta A. Gray, Proc. Amer. Acad. 8: 628. 1873. Caprifolium interruptum Greene, FI. Fran. 347. 1892, Stem stout and woody below, becomin g widely branched — often climbing or reclining on other shrubs; young branches glabrous cae decidedly glaucous. Leav es elliptic to a ovate or icular, 2— m. on d joe r bel flor e nate at the base or Fahd distinc, ‘in others short-petioled, coriace eau, glabrous, siraes on both sides, at more soo r side; inflorescence ge ous, 3-12 ong ; calyx glabrous ; corolla 8-1 long, pars ied often with a reddish tin e, the tube hbes ely gibbous at base, shorter than che: limb, — rous without and pearls so withi in: filaments spat ily hairy; berries red. pes ridges, Upper Sonoran and Transition pete Jackson Comers: southern eh, south ei Sg Rtg Praae gE pe fg western jones of the Sierra Nevada to the San Ber rea g Mountains, south- ern California; also in Arizona. Type loca “‘Juxta fl. Carmel prope Monterey.’”? May- 10. Lonicera subspicata Hook. & i Southern Honeysuckle. Fig. 5052. Lonicera (?) subspicata Hook. & Arn. Bot. es igeit 349. 1838. ula var. subspicata A. Gray, Proc. Am gpa 8: 628. 1873. um Greene, FI. Fr. 992. rrupta var. subspicata ya FL. W. Mid. Calif. 474. 1901. Stem nee and stout ee Fe ly clambering over shrubs, see twigs light brown, ookecanel and glandular. Leaves eous, linear-oblong, 1-3 ¢ m. lon margin revolute, dark green and glabrate or witha te seattet er hairs and subacenie fiends on ihe upper side, cinereous- uberulent beneath ; aa -5 mm. long; spikes 2-4 cm. long or sometimes longer, P ith sev om - oO scen mb ed wha lobes ; filaments thinly pilose belane also the oie heer 57 1 mm. ee Geede 2.5-3 mm. long, light Riss shallow ty Dit slopes, Upper Sonoran Zone; Santa Ynez Mountains, Santa Barbara County, California. Type local- ity: iat Fee ag Collected by Douglas, probably at Santa Barbara. June—Aug. HONEYSUCKLE FAMILY 55 Lonicera subspicata var. johnsténii Keck, Bull. S. Calif. Acad 1926. (Lonicera johnstonis McMinn, III. Man. Calif. “Pe 541. 1939.) Leaves broader, varying ican cblone. ovate to broadly orbicular, whitish beneath; corolla 10- mm. long. Dry slopes, California Coast Ra anges, San Benito County, and Te. hachapi Mountains south to bia mountains of southern Lower California, Type “te ocality: Poppet Flat, Idyllwild Road, Riverside County. Southern Foran te ickle. Lonice a subspicata var. den a Rehd. Rep. Mo. Bot. Gard. 14: ode 1903, (Lonicera denudata Davids. & Moxley, Ti. S. Calif. 344. 1923. " “Siniee to the preceding variety but leaves mostly oblong- ath or cy ovate, 5- mm. broad, very ae B gee Showin cinepig | and green, _not whitish sodery Southern San Dieg ounty and adjacent Lower Califor Type locality: San Diego. San Diego Honeysuckle. 5048. Lonicera ciliosa 5052. Lonicera subspicat 56 VALERIANACEAE Family 145. VALERIANACEAE.* i eooteray ceca usually more or less zygomorphic, the tube narrow, often gibbous or spurred at the base. Stamens 1-4, epipetalous. Ovary inferior, 1-3-celled with a solitary pendulous ovule. Seed without endosperm, the embryo straight and with oblong cotyledons. A family me — 10 genera and 370 species of wide geographic distribution, but most abundant in the northern hemisph laa m3 — gibbous or spurred, white, pink, or bluish, less than 1 cm. long (except Valeriana columbiana). of plumose bristles in the fruit; native perennials with rhizomes or a ot Woleitaiin. Calyx- it on annual species. Flow cymose clusters forming . more or less flat-topped inflorescence; stem laa ao * eed: woth als, sive from Europe. Val nella, Wiese “aie clustered in vaifiahe or interrupted, spike-like inflorescences; nate e gnmual_ species. ectritis. Stamens tae. Derails ad = jong spur, usually magenta or red (rarely white), more than 1 cm. long; perennial, adve e from Eur 4. Kentranthus. 1. VALERIANA} [Tourn.] L. Sp. Pl. 31. 1753. Perennial, strong-smelling herbs from rhizomes or taproots. Leaves decussate, basal and cauline, spatulate and undivided to deeply pinnatifid. Pdlitetcnees thyrsifor rm Flowers perfect, gyn ioecious or polygamodioecious. Calyx initially involute, later u So 4, epipetalous. Ovary inferior, 3-carpellate with 1 fertile adaxial cell at maturity. dean cypsela with 3 abaxial and 3 adaxial veins. [Named for the Roman emperor alerianus. ] A genus of about — le pre in both hemispheres, chicky, _ the boreal and temperate regions but also in the aes: in the mountain , Valeriana officinalis L. Leaves mostly — — petiolate; plants from rhizomes or stolon: Leaves Het pesnlly.# stad spatulate in outline; corolla funnelform to pain dens 3-19 mm. long, the tube ° y wit ens ex ened. ete than the corolla-lobes; corolla- nade less than half the length of the tube. es ascending-ciliate, _ Alediggeee 1 lobe linear ae ic or oblanoeslaie, the lateral lobes of the a leaves in 8-1 ; adventiv bag esd uro V.. officinalis. Leaves glabrous or with cred €: pubescen Pics telatively ro robust 0 dm. tall; = least the lower cauline ot petioles. glabrous or Plants fol mapblanyd sleudiaie r, 1-6 dm. tall; cauline leaves essentially Ay a and Spe Sed ad the stem and/or the leaves sebacdlent V. capitata califor Stamens included, shorter than the corolla-lobes; corolla-lobes 3-6 mm. long about half ‘the edu of ube Wenatchee Mountains of Washi ington. . V. columbian Leaves mostly oblong in Pompe corolla subrotate to rotate, 2.0-3.5 mm. ca ge tube oot ‘‘grxiant or iaietinces gibbou ants — sa t and leafy, 3-9 dm. tis Pde sparsely to densely Pyar: sae a glabrous; northern California through Or V.. occidentalis Plants relail dy slender and less leafy, 1.5—4.5 in tall; fruits glabrous; A Sage then ape ok atic Leaves mostly Higuinte-spatulate, gradually decurrent to the clasping leaf-bases (the seri often sinestiha but more or s decurrent) ; plants from vertical, usually forked taproots. 7 Paik . V. edulis. 1. Valeriana officinalis L. Valerian or All-heal. Fig. 5053. Valeriana officinalis L. Sp. Pl. 31. 1753. _— from a short perennial rhizome, pilosulose to short-pilose toward the base, glabrescent Leaves mostly cauline, petiolate below w, sessile above, oblong to oblong-ovate, pinnate to Pinnatifd 9-36 cm. long, gl abrous to short-pilose, particularly on the veins beneath, glabrous ve, oe ira Pog eee lateral lobes 4-8 pairs, linear to oblanceolate, acute, more or less rseidoet 2.0-7.5 cm. long, 0.5-3.0 cm. pale the terminal lobe two-thirds as long but as wide as the lateral con: basal leaves 15-30 cm long ; inflorescenses thyrsiform with perfect flowers; ‘ext contributed by Richard William Holm ; Thies treatment is based largely on the monograph of F. G. Meyer, ie: Mo. Bot. Gard. 38: 377-503. 1951. VALERIAN FAMILY 57 corolla infundibuliform, 3-5 mm. long, white, the lobes half as long as the tube; stamens and ee ex shite bag uits lanceolate to ovate-oblong, 5-3.0 mm. long, glabrous or pilosulos se, tawny o rubig 14.28. REAM into gardens in the United States and Canada, and established as a garden escape in parts of Washinetsn May-July. 2. Valeriana RE Bong. Mountain Heliotrope. Fig. 5054. Valeriana sitchensis Bong. Mém d. St. Pétersb. VI. 2: 145. 1832. Valeriana hookeri Shuttlw. Flora py 450. 1837. Valeriana suksdorfii Gandoger, Bull. Soc. Bot. Fr. 65: 36. 1918. Valeriana frigidorum Gandoger, op. cit. 37. Valeriana anomala Eastw. Leaflets West. Bot. 3: 22. 1941. Stems robust from a rather s tout rhizome, 3.5-12.0 dm. tall, pert in Sgr a often wanting, “petioliite: undivided or pinnate to ger orm ovate- elliptic to obovate, 10-40 cm. long, dentate to repand or entire, the deteiend lobe 5-10 cm. long, 1.5-6.5 cm. i 8 gla brous ; scans leaves sebdenibeatiiy 2-5 pairs, pinnate to pinnatifidy 10-20 ¢ eg Lh the pied lobes 1-4 p airs, crenate to irregularly repand-dentate or es a entire, et inal lo ovate, iseubas rhombic, to psbA ve ins 5 cm. wide, see c Sihre se; in orescence thyrsiform with per- fect or gynodioecious flowers; or intondibuliformn, 4.5-7.0 long, w white to pinkish, the - usually oe than half - long as the gibbous tube; stamens pail rie exserted ; fruits ovate o oblong-ovate, 3-6 mm. long, i beige ft e, ian or purpu pcabcont Becucntiy purple- maculate. Subalpi nied a woodlan Canadian and Hudsonian Zones; Kenai anne Mie southward on the Alaska coastal mountains to east- pion and southern British Columbia castward to south-central Idaho and west- ern Montana; Washington and Oregon to os 5053. Valeriana officinalis 5054. Valeriana sitchensis 58 VALERIANACEAE pine slopes, meadows, slong creek Deon and also in coniferous woods, Boreal Zone; southern Or regon and hg Sierra Nevada of California to Tulare County, ae ie ela Nevada. Type 1 ocality : ridge south of ce Pass ,500 feet, Ne si rhe Caer California ig aleriana ee subsp. pubicarpa (Rydb.) F _ Mo. Bot. Gard. 38: 406. 1951. (Valeriana pubicorra Rydb. Bull. dy Sig Etub 36: 697. 1909; vo Poe sebeoios "Rydb. loc. cit.; V. cusickit Gandoger, Bull. Bo T. 65% 36, 8; V. utahensis Ga ndoger, op. ci ; V. maculata Eastw. Leaflets West. Bot. 3: 1941 ) This subspecies differs from the above in its uniformly puberulent stems and mostly entire and glabrous leaves. On stony sagebrush slopes or eee pines and on talus slopes in the mountains; hg Montana, western Behe pa aon central ane ho northern Utah; foray gs Oregon to the Charleston Mountains of 4. Valeriana columbiana Piper. Wenatchee Valerian, F ig. 5056. Valeriana columbiana Piper, Bot. Gaz. 22: 489, 1896. Stems leafy from stout nego 0.5-3.0 dm. tall, glabrescent above, uniformly puberulent below. Basal leaves formin gaia — petiolate, undivided, broadly ovate to ovate- oblong or sometimes suborbicu lar, etic ‘tied 1.0-3.5 cm. wide, irregularly dentate to entire or nearly so, glabrous or the veins eres s pilosulose ; cauline leaves airs, 412 cm. long, perm a mine, .9-2.0 mm. wide, elgg hese oth, sometimes purple-ma oath en to forested rocky slopes fa sag ntains, Arid Transition me rit Canadian Zones; north-central Washin oe ie a kai = Mountsidx a (so locality: ‘‘side hill above Farwell’s hone west of Wenatchee,” elan County 5. Valeriana occidentalis Heller. Western Valerian. Fig. 5057. p cheriaws wepidentalts Heller, Bull. Torrey Club 25: 269. 1898. E. Nels. Erythea 7: 166, 1899. Stems more or less leafy from the rhizomes, 4.5-9.0 dm. tall, oo = papecipon sees Basal leaves petiolate, undivided or pinnate to pin natifid, oblong to narrowly o r less spatulate, rarely suborbicular, 12-30 cm. long, essentially entire, glabrous, pte . ad fine: inal lobe of the divided leaves 2-10 cm. long, 1.3-6.0 cm. wide, short-acuminate or obtuse, the lateral lobes 1-2 pairs, soting * maller ; cee leaves 24 me s, the hewecu it short-petiolate, pinnate to pinnatifid or sometimes undivide d, 4.5-14.5 cm long, the upper toes sessile and much re- uced, the ieroial | lobe oblong- linear, ovate to obovate 6.8 c¢ ong, 1-4 on wide, acute or obtuse, the nage lobes 1-6 pairs, grading smaller ; inflorescences ths rsiform with gynodi pera owers ; ‘corolla rotate to subrotate, 3.0-3.5 mm. ong, white, the tube slightly HA, | ei stam and style ex sexi — inear- to ovate- oblong, 3-5 mm. long, 1-2 mm. wide, spatsely to si , tawny. O fornia, and eastward to ctor Nevede and Utah; also Wyo and Pabbude. Type ais near the western end of the Craig Mountain plateau, above Lake Waha, 5 pes tous county, Idaho.”’ May 6. Valeriana dioica subsp. sylvatica (Sol. ex Richards.) F. G. Mey. Wood Valerian. Fig. 5058. Valeriana sylvatica Sol. ex Richards. in Frankl. 1st joe Bot. App. 730. 1823. Valeriana wyomingensis E. Nels. Erythea 7: eg Valeriana septentrionalis Rydb. Mem. N.Y. B UP oss ee 376. 1900. Valeriana psilodes Gandoger, Bull. Soc. Bot. ay 65: 37. 1918 Valeriana dioica subsp. sylvatica F. G. Mey. Ann. Mo. Bot. Gard 38: 417. 1951. Stems more or less leafy from slender rhizomes 0 tall, ip ores ca - se a asal leaves petiolate, undivided, aris , Ovate- ania or Sieh ni ely suborbicular, 3-27 c long, essentially entire, the blades SOc m. long, 0.5-3.0 cm. wide, s voy os en acute, oe obtuse; cauline s 3-4 pairs, es lowermost short-natisinie pinnate to pinnatifid, rarely un- divided, the uppermost sessile and h re above, oblong, ovate, to ovate-oblong, 2.5-11 cm. long, glabrous, the terminal | ear- to ovate-oblong, acute rt-acu te, 1-5 long, 0.2- , the lateral lobes 1-7 pairs, grading smaller; inflorescences thyrsiform ib hes owers; coro a rotat to subrotate, 2-3 . long, white, the tube —- Mea pois ea t wooded hillsides in rocky duff-covered clay soil, or on talus slopes, Arid Transition Zone; pie Sty "northwesterly across Canada to the Rocky Mountains — southward to central Idaho and eastern Washington. Type locality: ‘‘on the Clear-water River.” May— Spa 7. Valeriana édulis Nutt. ex Torr. & Gray. Tobacco Root. Fig. 5059, Patrinia ceratophylla Hook. F\. Bor. Amer. 1: 290. 1834. Valeriana edulis Nutt. ex Torr. & Gray, Fl. N. Amer. 2: 48. 1841. Valeriana perce | A. Ne . Bull. Torrey Club 28; 232. 1901. V ach ydb. . cit. 31: 645. 1904. Valeriana ceratophy: ma tt pie western sae not H.B.K. Stems robust, 1-12 dm. tall, usually Taos, with the leaves mostly basal, from vertical, often forkes taproots, 0.5-3. fem . thick. Basal leaves linear or oblong- to obovate-spatulate, VALERIAN FAMILY 59 undivided or a to aye subacute to obtuse, 6-40 cm. long, 0.3-6.5 cm. — gradually tapering to the base, more or less repand to entire , Spreading-ciliate, pilosulose to appressed white- puberulent or fs prea or the oles often puberule nt, the lateral lobes of the divided leaves 1-4 pairs, mostly distinct or sometimes narrowly decurrent, the terminal lobe 4.5-9.0 cm. long, .9-2.0 cm. wide; cauline leaves 2-3 Gell develiped pairs, pinnate to Seat: rarely undivided, short-petiolate or sessile below , much reduced a ve, 3-22 cm. long, elliptic- to obovate-spatulate : inflorescences thyrsiform, large, with polygamodioecious flowers; corolla rotate, the perfect and staminate flowers 3.0-3.5 mm. lo ong, the sh piaeciak 0.5 mm. long, white, the tube slightly gibbous; stamens and diyte exhertet frui : poem —oe to densely hirsutulous. In moist pastures, creek bottoms, yellow pine ca asp wos, ush plains, cliffs, and subalpine parks, Arid Transition and lower Boreal Zones; ssatherh British “Colnon re wed “th United States except California to northwestern Mexico. Type locality: ‘Walla Walla, Walla Watts County, Washington. May-Sept. 2. VALERIANELLA {Tourn.] Mill. Gard. Dict. abr. ed. 4. 1754. Dichotomously branched annuals with basal leaves tufted and entire, the stem-leaves sessile and often dentate. Flowers in terminal and usually corymbosely arranged cymose clusters. Calyx-limb short or obsolete in the flower, short- tooth when present; in the fruit at the summit. Ovary 3-celled, one cell fertile, the others empty but one usually about as large as the fertile one. [Name diminutive of Valeriana.] 5055. Valeriana capitata 5057. Valeriana occidentalis 5056. Valeriana columbiana $058. Valeriana dioica 60 VALERIANACEAE About 50 species, native of the northern hemisphere, and most abundant in the Mediterranean region. Type species, Palorinne locusta L, Fruits sg in cross section, with a geosereet eosky. por on the dorsal side of the fertile oct, sd Sasol ig the sh empty cells usually not produced, separated by rrow shallow groove. V. loc Fruits square in cross section, without a ives mass on the dorsal side of the flattened fertile a erkire of the empty cells incurved, forming a broad deep groo 2. V. carinata. 1. Valerianella — (L.) Betcke. Lamb’s Lettuce. Fig. 5060. Valeriana locusta a a ol itoria L. Sp. 1753. Valerianella olitoria Poll. Hist. Pl. ote nae 30. 1776. Valerianella apne Betcke, Anim. Val. 10. 1826. Ste ee or usu suily dichotomy branched, 15-30 cm. tall, glabrous to more or less puberalent a least at the nodes, the hai = refle xed. Basal leaves spatulate to oblanceo- late, rounded or als at the pil ee oe long, 6-15 mm. wide, pate upper stem-leaves broadly linear ‘ oblong, —2. m. long, entire or sometimes sparingly toothed ; peduncles short ; cymes 1 eee ibcopibite: bracts linear to narrowly oblong; corolla bluish or pink, about mm. eee fruit 2 mm. long, about as broad as long, itr Ily compressed, finely foveolate, gla- brous, Aa Rckuid cells oe by a narrow shallow groove, the fertile cell with a thick corky mass on the dorsal side. » = 7. coe sional in waste places and cultivated fields; Washington, Oregon, and central California and across the eeiacut to the eastern and eaters states; satueatignt from Europe and sometimes cultivated as a salad plant. April-June. 5059. Valeriana edulis 5061. Valerianella carinata 5060. Valerianella locusta 5062. Plectritis congesta VALERIAN FAMILY 61 2. Valerianella carinata Loisel. Corn Salad. Fig. 5061. Valerianella carinata Loisel. Not. Pl. Fr. 149. 1810. Valerianella praecox Willk. Linnaea 30: 104. 1859. Slender erect annual, 1-2 dm. high, simple below, once to thrice dichotomously forked above, glabrous to more or less puber ulent, especially on the lower nodes and leaf-bases, the hairs some- what reflexed. Basal leaves spatulate, 1-2 cm. long, 3-5 mm. wide: upper stem-leaves linear- oblong, 1-2 cm. long, 2-3 mm. wide, ihc entire or cep and obscurely denticulate ; peduncles slender, 1 . long; cymes yor te, mm. bro ract eg . linear, 3-4 long, ciliate on the margins; corolla light ies +3 pin ; fru about 2 mm. ae and barely half as rsiont ae scent, not navopiate, the fertile cell. fattened, the margins of the empty cells incurved, form a broad dee Locally adventive on shady banks and eens Niagne and aie Ciuntiia: California, and Marion and Clackamas Counties, Creata. April-May 3. PLECTRITIS* DC. Prod. 4: 631. 1830. Usually ieee annual, tage herbs with opposite, oblong to ovate leaves. In- floréscences thyrsiform, dens ely to loosely capitate or verticillate to spiciform; bracts linear, fused i a a multifid structure. Calyx reduced to a narrow ring. Corolla subcam- eer ie to narrowly infundibuliform, white to deep pink, occasionally with spots of dark red at the base of the midventral lobe, strongly bilabiate to gcse y regular, the tube spurred or merely gibbous. Stamens 3, exserted to include til inferior, 1-celled, the stigma bifid, rarely trifid. Fruit pale straw-yellow to dar -brown, vernicose to dull, usually more or less pubescent, dorsally keeled, the ventral geet with glands eo in 2 longitudinal rows near the base of the wings or as 2 prominent mammillae: wings con- nivent or spreading, often obsolete, occasionally with glands on the dorsal surface o spose those on the ventral surface. Cotyledons accumbent or incumbent. [Name from the Greek, plectron, meaning a spur. ] A genus of 3 species in the Pacific States, formerly er into many species because of the extreme varia- tion of the fruits. Type species, Il” Cierienctia congesta Lindley oe pale to medium pink, bilabilate, funnelform, spur usually less than one-third the length of the corolla r obsolete; fruit keeled, the keel rarely grooved, acutely angled (with prominent veins) or smoothly punsed (with pronounce ed vacuities between the walls of the fruit and the embryo), winged or wingless, th oes thin and thin ae pie caer sa pgp ye flexible, evenly tapered, and more or less Pic gesta. ae ger and essentially regular to light red and strongly bilabiate, tubular or somewhat fadlaitocen, spurred, e spur tem more than one-third the length of the corolla; fruit heeled: the keel often with a dorsal roove, obtusely angled, raves with visible. areal hina Boies ged or essentially wingless, the wings stiff robe often eed the +H ap Pome ype ir ight often groo Pigs cellent the trichomes stiffly clavate, cylin- drical, glandular- cylindrical or lor d ‘curly; coty ledoks incumbent, silvery green, with secondary and tertiary veins promine m~ on oraiinalinen Corolla carne ty regular, white pal le oe: ink, usually without red spots at the base of the igo lip, stout and with a stout clav. fruit with the bs ggeror ie and Lege or more or less nar- rowly cylindrical and ashes tipped, sometimes long and curly, the kee! Hobe 2 brush- like rows of hairs, the ventral surface often with a multiseriate row oe hairs along the m a Corolla strongly eed Lone to light red, usually with 2 red spots he base of the ventral lip. sender and with a ; fruit wi — the trichomes stout, Sint ical = rarely subclavate, never lon curl. "the ve gp ag with 2 brush-like rows of hairs, the ventral surface usually lac ing a es seriate row of hairs along the eae ib. 3. P. cihosa. 1. Plectritis congésta (Lindl.) DC. Pink Plectritis. Fig. 5062. Valerianell. gesta Lindl. Bot. Reg. 13: pl. 1094. 1827. Plectriti. ta DC. Prod. 4: 631. 1830. Plectritis congesta B minor H ook. Fl. Bor. Amer. 1: 291. 1833. Plectritis bt Suksd eut: Bot. Monatss. 14: 119. 1896. h. lectr istiraceme losa Gandoger, eee Soc. Bot. Fr. 65: 35. 1918. Plectritis ape Gandoger, loc Plants 1.5-6.0 dm. ta i, = Leaves obovate ne ovate, pero e below to sessile or amplexi- aule above, entire a submuc nate or inflor 3 : vertisiatt-¢ spiciform ; corolla fale pink to pink, s beau sh ate to infundibuliform, 4.5-9. : long, strongly bilabi ate "be dorsal lobes fused did extended, the ventral free and patent, the spur short to obsolescent, the tip usually conspicuously expanded ; fruit wn, 2.0-4.5 ly pubescent, the al reo gradually and evenly attenuated to a thin point, the Lena) © & = 2 g x ° mm. walls thin and readily colla sharply angled, acute, very slightly eiented dorsally, smooth, or with 1-3 prominent veins, vari- ously pube or labrous, casionally with 1 to pbs glands at the wing-roots ; ventral wall r: present bo ngitu rows n i : psed on drying; fertile cell strongly Pion deeper than broad; keel th i : broad f ally strongly spreading above; cotyledons accumbent, veins above the first order inconspicuous when expanded. *This treatment is based largely on the recently published work of Dennison Harlow Morey, Jr. 62 VALERIANACEAE Am shrubs and bushes in shady places (usually with marine influences) at low altitudes, Upper ona and Hu =a Transition aa British Columbia, Washington, Oregon, and northern California south along the coast to Monter rey and San Luis Obispo Counties. Type locality: ‘‘no orth-west coast of North America,”’ Collected congesta subsp. nitida (Heller) Morey, Contr. Dudley Her b. 5: 120. 1959. (Plectritis nitida Heller, Muhlenbergia Hy 328. 1907.) Differing from the typical subspecies in the vernicose or mare fruits Mee the keel smoothly vas ad - ae the bagi when present appressed and connivent top and bottom; walls of the fruit usually more or les e fro he embryo. Slender decumbent plants of more or or Fela wet pan poiected 3 the coastal lowland fre Monterey to Fort Bragg, California; rarely elsewhere. Type locality: Crystal prings Plectritis congesta subsp. ag ea gt (Fisch. 7 as y.) Morey, Contr. Dudley Herb. 5: 119. 19 a (Plectritis brachystemon Fisch. & Mey. Sem. Hort. Petrop. 2: 47. 1835; Betckea major Fisch. & Apn: Sci, Nat. JI... §: 189. 1836; Plectritis samolifolia Toeck, Bot. Jahrb. < 37. 1882, in part; V alerianella anomala A. Gra , Proc. Amer. Acad. ‘S35. V. aphanopter ra A. Gray, cit.: V. magna Greene, Proc. Acad. Phila. 1895: 548. 1895; Plectritis iivales "Suksd. Deutsch. Bot. ask 15: 144, Biincd i P. gibbosa Suksd. Ws denda 1: 8. 1927.) Differing from the typical subspecies in the smaller flowers (1.0-3.5 mm long), wh may lack a spur, and which have a tube 0.5 mm. long or longer, and in the jpcils tubular-funnel- form cmos. In brushy hot relatively open, pomp te “habitats scene ag ase asain to Monterey County, Cali- fornia. Type locality: ‘‘prope coloniam Ross in California,’? Sonoma Cou 2. Plectritis macrécera Torr. a art White Plectritis. Fig. 5063. Plectritis macrocera Torr. & Gray, : 50. ue ra etchleriana Suksd. Detitscti, Bot. Misetix: 15: i aay. ligera ostio latata Suksd. loc. cit. pb ie jepsontt Suksd. Erythea 6: 24. 98. Plectritis arate! Jepson, Fl. W. Mid. Calif. 475. 1901, Plectritis collina Heller, Mutter 2! 329.;,1907. lants slender, 1.0-6.5 dm. tall. Leaves obovate to ovate-lanceolate, diminishing in size upward, pe etiolate below, tag above, entire to coarsely serrate; inflorescence verticillate (some- spe 1 i aligy ornot at all expanded; fruit pale straw-yellow to red-brown, 2-4 mm. lon ore ess pubescent, the hairs stout, clavate to glandular-c ylindric, rarely obuisely donned: oe ‘erga ‘often collapsed on drying, or long and curly; keel rounded to rather angular, obtusely angled, often a dorsa : n na with a pair of equatorial pecoeget wings thick, expanded, or entially obsolete, bounded with a broad, cylindrical, usually grooved, ok ace thickening, came nt basally and apically to an equal degree and prese sora a pete t is broader than tall ; Cavlelchil incumbent, very aaa: veined to the third order en expanded ed, but not brushy habitats, Upper Sonoran and Humid Transition Zones; San Fra cisco 4 Po tase oe Citote from sea Hey o 4,000 feet, with pied pk kee sanoms to southern if cthineton and southern California. Type locality: “California.” Collected by Douglas. April—-Ma Plectritis macrocera subsp. grayii eam) Morey, Contr. Dudley Herb. Ps 120. 1959. (Aligera grayi Suksd. Deutsch. Bot. Monatss. 15: 147. 1897; A. mamillata Suksd. loc. cit.; Plectritis macroptera Rydb. il Rocky Mts. 818. 1917, not Suksd.) Differing from the typical subspecies in that the wings when expanded a thin and eth a narrow, but usually obvious marginal thickening which is only rarely or ape So igre groove a wings are connivent top and bottom to present a large circular orifice; and there is usuall median longitudinal ridge on the ventral surface of the fruit which usually bears a multiseriate row of ‘sti - clavate or cylindrical bristles. In open, moderately shaded, but not brushy habitats in piedmont and moist, low, montane peacoat north-central California northward to the Columbia River and its A ro nslveg eastward to southwestern Montana and sth hea to southern California, Nevada, and Utah. Type locality: California. 3. Plectritis cilidésa (Greene) Jepson. Long-spurred Plectritis. Fig. 5064. Valerianella oe it We Proc. Acad. Phila. 1895: 548, 1896, Aligera macroptera Suksd. Deutsch. Bot. Monatss. 15: 146. 1897. ont oe pany op. cit. 147. Not Torr. & Gray. sa Jepson, Man. FI. Pl. — O71... 1925. i: si Pst Werdenda 1: Plants slender, 1.0-5.5 dm. ta = Leaves obovate to oblong-fusiform, petiolate —— sessile above; nae wage to verticillate-spiciform ; pers deep pink, with dark s at the base of the ventral lo 5 mm. long, — and narrowly infundibuliform, bilabiat jate, the spur slender, usually ey ‘the ovary, the tip not eh ag ; fruit pale straw- ellow to brown, 34 mm. long, more or less es the hairs stout ot iff, cylindrical, rome or bluntly pointed, rarely somewhat lig eae and not readily ‘collaps ed on drying; i than dee and stretched from ridge to inner wing-margins, often wit of equatorial mammillae; wings variously expanded, rarely obsolescent, bounded with a pa ae dy "ean grooved, marginal thickening, connivent tegey # and connivent to more or less spreading above; cotyledons seabed, more or less conspicuously veined to the second and third order when expanded. In open sunny habitats, grassy hillsides, and stream margins, Upper Sonoran and lower Humid Transition Zones; central Cali fornia at higher altitudes in the interior from San Benito County north to Mendocino PSone = pons Tuolumne County po to Monterey County; rarely elsewhere. Type locality: Oakville, Napa County. pril— Plectritis ciliosa sub: nsignis (Suksd.) Morey, Contr, Dudley Herb. 5: 121. 1959. (Aligera insignis uksd. Deutsc ie Faia oe i 146. i897) A rubens Suksd. loc. cit.; A. elliformis Suksd. W. Amer. Sci. 12: 54. 1901; Plectritis davyana Jepso'! _ W. Mid. Calif. 475. 1901; ligera intermedia Suksd. Wer- VALERIAN FAMILY 63 denda 1: 44. 1927; A. glabrior Suksd. loc. cit.; A. baybuts Suksd. loc. cit.) Differing from the tyoiest subspecies oe ace hing in the smaller flowers (1.5-3.5 mm. long In hotter and Pend ey aoe pe in dappled shade in the southern part of the range; pst hdl California and — inland coastal valleys north a Cone pero ag to the Columbia Gorge, and in Arizona. Type localit rthern Lower California. Collect oe by Orcutt 4. KENTRANTHUS Neck. Elem. 1:'122. 1790. Perennial or annual, glabrous herbs with simple or pinnatifid leaves and large, termi- nal, thyrsiform inflorescences. Calyx inrolled in flow < penn ming a large plumose pappus in fruit. Corolla- tube usually with a long spur, the limb . somewhat zygomorphic. Stamen 1. Fruit somewhat compressed, Fie ad by the persistent hatase calyx. [Name from the Greek, meaning spurred flow enus of about 12 species in ee Mediterranean region. Type species, Valeriana rubra L. 1, Kentranthus ruber (L.) DC. Jupiter’s Beard or Red Valerian. Fig. 5065. Kentranthus ruber DC, in Lam. & DC. FI. Franc. ed. 3. 4: 238. 1805. Glabrous and somewhat glaucous, perennial herb to 1 m. high. Leaves simple, entire or rarely oothed, ovate: to seneshlate! to 10 cm. long, sessile; inflorescence 4 a gg a paniculiform cyme elon- gating in fruit ; calyx-limb with 5- 13 linear plumose lobes long; oo evan magenta, or white, with a prominent spur at the base, about 16 mm. long including ¢ the s A A agar plant occasionally escaped pest tiltivation: in moist areas, particularly along ey dead in California. Jan.—Dec SS Sake a EN aD = adn ee. ‘ ¢ ae SASS, a NIRK Wi MAY. Q “: 4, hat aa t\ yy We? a aN .\ ry at Ad est ~ RSS ‘i ; 1" ya a : : — a “4 ‘ 5066 \ \ — ay ‘ i : \ 5063. Plectritis macrocera 5065. Kentranthus ruber 5064. Plectritis ciliosa 5066. Dipsacus fullonum 64 DIPSACACEAE Family 146. DIPSACACEAE. TEASEL FAMILY Annual or perennial herbs with opposite or verticillate, exstipulate, simple or pin- natifid leaves. Flowers perfect, borne on an elongated or ‘globose receptacle, bracted and involucrate. Calyx-tube adnate to the ovary, the limb cup-shaped or disk-shaped, or divided into spreading bristles. Corolla epigynous, the limb 4—5-lobed. Stamens 2-4, inserted on the tube of the corolla and alternate with its lobes; filaments distinct. Ovary inferior, 1-celled ; style filiform ; stigma undivided, ter minal or lateral; ovule :; eerie Fruit an achene, its apex crowned with the persistent calyx- lobes. family of about 10 genera and 180 species, natives of the Old World Bracts of the involucre prickly-pointed, some exceeding the heads. 1. Dipsacus. Bracts of the involucre herbaceous, shorter than the heads. 2. Scabiosa. 1. DIPSACUS [Tourn.] L. Sp. Pl. 97. 1753. Tall, erect, rough- nay or prickly, biennial or perennial herbs with iat large leaves and bluish o r white flowers in dense, more or less elongated heads terminating stout pe- uncles. Bice of the he and scales of the receptacle rigid or spiny-pointed. Flow- Aeron free from or adnate to the involucre. [Greek name for the teasel n Old World genus of about 15 species. Lectotype, Dipsacus fullonum L Bracts of the receptacle hooked at apex; flowers white. 1. D. fullonum. Bracts of the receptacle straight; i lavender. 2. D. sylvestris. 1. Dipsacus fullénum L. Fuller’s Teasel. Fig. 5066. Dipsacus fullonum L. — P1::97, oe Dipsacus fullonum B sativus L. loc Dipsacus fullonum of saat 1762; os ller, 1768. Dipsacus sativus Honck. Vollst. Syst. Verz. 16. 1782. Stout biennial, 1 m r more high, with scattered short prickles on the stems, midribs of the the e we foliaceous bracts of the involucre apreane or reflexed, some shorter than the rae heads ovoid becoming cylindric, 6-10 cm. long; bracts of the receptacle with hooked tips, about equaling the flowers ; corolla white or tinged with reste 8-12 mm. long; achenes 6- 8 mm. long, strongly 4-an ngled. Waste places especially in low damp ground; frequent in central and southern California. Native of Europe. March—Oct. 2. Dipsacus sylvéstris Huds. Wild Teasel. Fig. 5067. Dipsacus sylvestris Huds. Fl. Anglica 49. 1762. Stout biennial, 1-2 m. high, with 5 ea short prickles on the stems especially below the inflorescence, ga ‘midrib of tee a es and t phage ate stg 2 Lceantacion or nearly so. L leaves obtuse, c enate or sometimes ‘pnnatifd, often 2-3 dm. long; upper stem-leaves sessile or = o 5 the uppermost slightly Fes sad acu at apex, and usually entire; foliaceous bracts of the involucre as long as the hea a oct, he ar and curved upward; bracts of the receptacle ovate, tip by a long, seat subilat, fo — d awn usually exceeding the flowers; corolla lavender, 9-12 m er ong; e 6-8 angled. Ss places, common in Was “wh ngton and Oregon and occurring in northwestern ee as far south as San Francisco County; also eastern United States. Native of the Old World. June—Sep' 2. SCABIOSA [Tourn.] L. Sp. Pl. 98. 1753. Glabrous or pubescent herbs with leaves opposite and no prickles. Bracts of the in- volucre herbaceous, distinct or slightly united at the base. Receptacle pubescent, without scales. Involucels compressed, their margins often minutely 4toothed. Calyx-limb 5-10- awned. Corolla-limb 4-5-cleft, oblique or 2-lipped. Stamens 4 or rarely 2. Stigma oblique or lateral. SL otere more or less — to the involucel, crowned with the persistent calyx- limb. [N atin, meaning scale, being a reputed remedy £ for scaly eruptions. | An Old oo genus of about 75 species. Lectotype, Scabiosa columbaria 1. Scabiosa atropurputrea L. Mourning Bride. Fig. 5068. Scabiosa atropurpurea L. Sp. Pl. 100. 1753. Stems branching, 6-10 dm. high, glabrous or sparsely and retrorsely hispidulous. Lowest GOURD FAMILY 65 leaves ee a Atti a cpg: or the uppermost coarsely serrate, or the smaller best narro and entire; peduncles 2-3 dm. long; heads 3-4 cm. broad, the flower s very dark purple varying . pale sialon i or sire corolla 10-12 mm. long ; gue of fruiting calyx 6-10 mm. Mine sca es rou cai from gardens, and naturalized in the Sacramento Valley and San Rings ped roo region; also Santa Barbara, ee Bernardino, and San Diego Counties, California. Native of Eur May-Nov. Old L ady’s Pin- Family 147. CUCURBITACEAE. GOURD FAMILY. ostly herbaceous annual or perennial vines, trailing or climbing by means of tenia with alternate petioled leaves and solitary, racemose, paniculate or cymose, monoecious or dioecious flowers. Calyx-tube adnate to the ovary, its limb usually 5- lobed. Petals usually 5, separate or united, inserted on the limb of the calyx. Stamens mostly 5, the androecia highly pal throughout the family. Ovary 1—3-celled ; style simple or lobed; ovules few or numerous. Fruit a pepo, indehiscent, or rarely dehiscent at the summit, or bursting erreoulatiy Seeds commonly flat ; endosperms none Flowers yellow, large; pistillate ao staminate flowers solitary. 1. Cucurbita. ers white or greenish, small; minate flow racemose or paniculate Fruit symmetrical, oblong, raat or = 1. a cm. long; ovary 2—4-celled. awe — roots slender ae setey 2. Echinocystis. Seeds turgid; roots very lar can tedden Fruit reas 1 cm. or less Hhteter ovary 1-celled. 4. Brandegea. 1. CUCURBITA [Tourn.] L. Sp. Pl. 1010. 1753. ough prostrate vines, rooting at the nodes, with branched tendrils, Revere lobed leaves which are often cordate at the base, and large, yellow, axillary, monoecious flowers. Calyx tube campanulate, usually 5-lobed. Corolla eenien: 5- iobed to shiek the middle, the or less united. Pistillate flowers with 1 pistil; ovary oblong, with 3-5 many-ovuled lacentae; style short, thick; stigmas 3-5, each 2-lobed, papillose; staminodia 3. Fruit large, fleshy, with a thick rind, indehiscent ; seeds horizontal, many. [Latin name for the gourd. An American genus of about 15 species, mainly tropical. Type species: Cucurbita pepo L. Leaves triangular-ovate, longer than broad, scarcely lobed and often angled at base; tendrils long-petiolate. * oetidissima, Leaves soe pingeigers lobed or divided to the base, as broad as or broader than long; sediile senate or short iolat Lome s-cleft to the base with narrowly lanceolate or linear lobes, these often sublobed toward Ag 5 em igitata. Leaves 5-lobed to tk iddl a little below, the lobes broadly lanceolate or triangular. 3. C. palmata. 5068. Scabiosa atropurpurea 66 CUCURBITACEAE 1. Cucurbita foetidissima H.B.K. Calabazilla or Mock-orange. Fig. 5069. Cucurbit apk lacanesylnin H.B.K. gs Gen. ‘s . a 1817, is Jam rlaeaei per mes in Long, Exped. 2 Cucu near tte ag Gray, een. Journ. a Fi yy 193. 1850. stout, rough, phar vrs trailing to a length of 2-5 m. eg Mags large, carrot-shaped. ate -triangular, cordate or pecans ~ base acute at Pie x, 1-2 dm. long, denticu- “at sally slightly 35. rpriar) rough abov cent beneath, on ihe in peti ioles 8-18 cm pe pede unc —5 cm. long; flowers mostly let pans 7-10 cm. long ; pepo globose, 5-10 Rp eonaang sm mooth. Wastelands, y, usually sandy soil, Upper and Lower gerade Zones; San Joaquin Valley and adjacent foothills “Of the Foi Micsae south to Lower California, Arizona, and arog also Nebraska and Missouri south to Texas and northern Mexico. Type locality: Guntiainnie: ae April-Sep 2. Cucurbita digitata A. Gray. Finger-leaved Gourd. Fig. 5070. Cucurbita digitata A. Gray, Smiths. Contr. 58: 60. 1853. ailing vine with = seated, mie i fusiform root and slender stems pustulate on the angles. Tendsits sessile or nea o, 3-5-parted, the divisions gland-tipped ; leaves 8-12 cm. broad, more green and less Si cs sive than below except on midveins, pedanely: ce lobed ‘to the base or nearly so, the lobes 4.5-10 cm. long, linear-lanceolate, sometimes attenuate at apex, each division gions near the base or bearing 1-2 distinct lobes or teeth; ottiole ribbed and scabrous, 4-7 .5 cm. long; flowers spa ringly pilose, 4-6 cm. long; staminate calyx cy indie or slightly flar below the ean subulate lobes D neicets enlarged at base of fruit; fruit depressed-globose, dark green, somewhat saa and marked with 10 narrow stripes; seeds thickened on the margin, ovate, 10-11 mm. lon In sand, pes pee washes, Lower Sonoran Zone; Riverside and Imperial Counties, patton, to New Mexico and er ghey Bebo Type locality: New Mexico. Collected by Charles Wright. July—Oct. 3. Cucurbita palmata S. Wats. Palmate-leaved Gourd or Coyote Melon. Fig. 5071. Cucurbita palmata §. Wats. Proc. Amer. — 215137, 1876. Cucurbita californica Torr. ex S. Wats. op. 138. Gray trailing vine from a deep- eel large, sometimes divided root with stems strongly ribbed and muriculate on the mieten Tendrils sessile or subsessile, weak, rid og ostly 3-divided ; leaf-blades more or less cordate at base, thick, gray, palmately 3-5-p a two Pesos lobes e up calyx 2. ong, narrowly campanulate, with short subulate lobes; fruit ater mottled, = marked with 10 faint lines, 8-9 cm. broad and long; seeds 10-14 mm. long, without a raised rid situations, Lower Sonoran Zone r Sacramento Valley southward through the San gona Valley, Californie, to northern Lower California va oun in the Mojave — Colorado mere rts to Mohave and Yuma Counties, hieleaa: Type locality: Cajon oe y, San Diego County, California. July—Se re ucur his californica Torr. ex S. Wats., which was eee Res os on the Wilkes Expedition, pre- sumably in the Sacramento Valley, is still, hpertcedy known o floral and fruit characters. As interpreted rte recent authors, it ‘earhade from C. palmata by having the teal lobe pt the leaf triangular and ioe than twice as long as and i found in the Poa abit and = sia Valleys south to San Diego County, a nia, east to the Deets Valier region and adjacent Nevada na, and in the Colorado Desert, Californi 2. ECHINOCYSTIS (Michx.) Torr. & Gray, Fl. N. Amer. 1: 542, 1840. Climbing herhs with branched tendrils and slender stems. Leaves alternate, palmately lobed or angled. Ss monoecious. aminate pg agus paniculate. Pistillate flowers usually solitary. Stamens 3; anthers more or less esced. Ovary 2-celled; les 2 in each cell. Fruit feet. pepominy dry at maturity ee deh iscing from the sum- mit. Seeds flat, dark-colored. [Name Greek, meaning hedgehog and bladder A monotypic genus of eastern and ce “te Fe ning States and adjacent Canada, occurring sbicaiticaliy in west- ern United States. Type species, Sicyos lobats Mic 1. Echinocystis lobata (Michx.) Torr. & Gray. Wild Balsam-apple. Fig. 5072. Momordica echinata Muhl. Trans. Amer. clas se 3: 180. 1793. (Nomen nudum) Sicyos lobata Michx. FI. Bor. Amer. 2: 217. Echinocystis lobata Torr. & Gray, Fi. N. — mL 542. 1840. Micrampelis lobata ah Pittonia 2: 128. 1890, Plants nearly glabrous, roots slender, fibrous, stems slender, climbing, 3-8 m. long. Leave 7-8 cm. seg inclu ang Lacinpric the blade with broad open sinus, 5-( sage 2 7-Vobed to about the middle, the lobes narrowly triangular, acute or acuminate, sparsely s rulate on the margin; staminate PA soe ag xilla narrowly paniculate, densely flowered, the corolla-lobes a spreading, slender, sthicuaie pisti tillat e flowers similar, solitary occasionally an in the — fruit about 5 cm. “ering yeni clothed with orgs ii Gaede 4, ths: netics thick. ish coa eastern United States and Canada westward to Montana but evidently escaping from cultivation and anecasiok. sporadically in the Willamette Valley, Oregon, and eastern Washington and west- ern Idaho. Type locality: Pennsylvania. Aug.—Sept. GOURD FAMILY 67 3. MARAH Kell. Proc. Calif. Acad. 1: 38. 1855. Climbing or trailing, herbaceous, monoecious vines from a deep-seated, greatly en- dry at maturity, 2—4-celled, fibrous within, dehiscing irregularly from the apical portion. Seeds turgid, often margined with a darker line; germination hypogeous. [Name Hebrew, meaning bitter. ] 5069. Cucurbita foetidissima 5071. Cucurbita palmata 5070. Cucurbita digitata 5072. Echinocystis lobata 68 CUCURBITACEAE An American genus of about 10 species, confined to western North America and adjacent Mexico. Type species, Marah muricatus Kell. Mature erg Ke et ly smooth or bearing scattered weak, fleshy spines; seeds spherical or orbicular in outline and strongly compressed. Ovary = Siclles flowers tapering to a beak; fruit ovoid, usually attenua te at th - oreganus, Ovary of pistillate flowers spherical with the beak arising abruptly from the ovary; fruit spherical tsonii. Mature fruit with crowded, slender, acicular prickles or stout spines; seeds neither mhetieat nor orbicular in outline and strongly compressed (compressed in M. juntabugonsis’. Staminate flowers aah campanulate, the lobes erect and recurved distally; fruit aes cm. long. . M. horridus. Staminate fl with rotate-sp g lobes, the tube shallowly basin-shaped; fruit 2.5— 3 cm. long. Mature fruit ovoid to oblong. Fruit 8—14- aE the seeds not over 2 cm. long, not compressed; leaves usually lobed to below the iddle. 4. M. macrocarpus. Fruit 2—4-seeded, the seeds about 3 cm. long, compressed; leaves mostly not lobed to the middle. 5. M. guadalupensis. Mature fruit globose, except for the short beak. Staminate inflorescence mostly few-flowered, the flowers bright white; leaves esa i D aera d parted to the middle. 6. M.it a ee ee many-flowered, the flowers greenish; leaves typically with ieee: broad, triangular lobes 7. M. fabaceus. 1. Marah oreganus (Torr. & Gray) Howell. Western Wild Cucumber or Coast an-root. Fig. 5073. Sicyos angulatus Hook. FI. ae Sar i ate 1833. Not L. Sicyos oreganus Torr. & Gra 1,N. Am de Aigin 1840. arah muricatus Kell. Proc. eit proves fs 1855. (Nomen confusum, not Echi tis muricatus Kell.) Remnen oregana Torr. Pacif. R. Rep. 6: Sp 1858. (Nomen nudum) h 76. . Wa Bchinocustic oregana Cogn. Mem. Acad. Sci. aa 28: 87. 1878. Marah oreganus Howell. Fl. N.W. Amer. 239. 1898. s glabrous or somewhat scabrous, reaching several meters in length. Leaves suborbicular to ke cordate, uusally scabrous above and often pubescent beneath especially along the veins, 8-20 cm aves 5-8-lobed to a mi the ‘eka sometimes dentate on the margin, basal sinus deep and usually closed, blade m exceeding the petiole in length; staminate flowers always in long slen ee racemes, inc rae ihe 20-35 1 peduncle 35 cm. long, the pedicels slender, 5-20 mm. long; corolla white, 8-15 mm. broad, the corolla-tube basin-shaped, the lanceolate-oblong lobes about equaling or a little shertae than the tube, rotately spreading and the margin sometimes recurved; pistillate pela white, the lobes rotately spreading, the pedicels 6-20 mm. long; ovar id, ering to a glabrou ak, young spines sometimes bearing soft septate hairs; fruit ovoid, glab: 47 long, rounded and sometimes —— narrowed 2A the base, attenuate and nag ielied ae ed at the apex, completely glabrous or sparsely muricate with weak fleshy spines up to 1 cm. in length; seeds 5-6, orbicular in outline, teciciele, Hattened ae rather undulate, 18-22 mm. cae 4-7 mm. thick, often rk —. s and edges of woods, mainly mid Transi tion Zone; southern British Columbia southward west Op: ye cast of bee Cascade Mountains thro “— the ae River gap. Type locality: ‘‘On the Oregon [Columbi iver 2. Marah watsonii (Cogn.) Greene. Taw Man-root. Fig. 5074. Echinocystis muricatus Kell. Proc. Calif. Acad. 1: 57. 1855. Not Maral icatus Kell. si heen muric Breen Wats. Proc. Amer. Acad. 11: 139. 1876. n DC. esi Phan. 3: 819. 1881. Marah seutioutt nif Geeenk Pysliess Bot. Obs. 2: 36. 1910. Stems slender, 2-3 m. long, seabne by simple or branched tendrils. — glabrous, paler beneath, suborbicular to reniform in outline, broader than lon ne, the blades 7-11 cm. wide, the basal sinus open, rounded, the petioles shorter than or equa ing the Deeg in eligrindg dee ply es. lobed, sometimes to within 1 cm. of the base of the bla e, the sinuses betw lobes acute or rounded ; lobes of the leaf narrowed below, widened above and 3- pon vig 4 obed, the secondary lobes sometimes angled; staminate inflorescence racemose, 4-20 cm. long, few- flower ed (2-10), the flowers campanulate, white, 4.5-6. m. long, the nee ronda: at base, much surpassing js obtuse or triangular, sp beat. sceoiilate lobes ; pistillate flowers axillary, the pedicel 2-2.5 c long, similar to the staminate flowers or a little’ larger, th ry spherical, the es t long, arising abruptly from the ovary; fruit globo. depressed-globos th or soy e€, smoo uricate with broad-based weak spines, .pale green and marked with darker veins ; seeds spherical, 11-14 mm. in diameter, not at all compressed. hruh: the di belt, U peace Zone; Shasta County south in the Inner Coast ieee Ss eS ee sang and tn ‘ ie Siar Sag S ces da hs posa County, California. Type locality: Placerville, Eldorado County. Warckh-Ace GOURD FAMILY 69 3. Marah horridus (Congdon) Dunn. Sierra Man-root. Fig. 5075. Echi tis } ida Congdon, Erythea 7: 184. 1900. Marah isovihies Dunn, Kew Bull. 1913: 151. 1913. Stems striate, glabrous, long, and branching. Leaves glabrous or scaberulous, 4-12 cm. long, the width greater than the length, the sinus broadly rounded at the base of the blade, open or closed, the petiole equaling or a Eos met hcg ae. blade this 3-7-lobed, usually 5, og to well below the middle, to a lesser deg 1 lea pi haa season, the sinuses between the lobes acute; lobes of the leaf 1-2 cm ge i t bas — bristle- uEpee ex, the margins of the lobes with 1-4 bro. ad, ceanentee beetle jis pee on each side, eh merely sinuate- toothed ; staminate inflorescence 10- g, racemose, paras ree tt loosely sich, the flowering portion shorter than the peduncle; stamin flow hite, 1-1. g, 8 long, markedly campanulate with a truncate bate, pee ay revolute, oblong, the onli els or longer that slog il xillary, tube m shallow than the ent, narrowly ell iptic, o mm. pew fruit 10-16(20) y pubesc cm. long, reg | 4-celled, ae surface Seats kc e with many long spines of unequal length, these stout or slender on the same fruit, pe baa mes pubescent with soft hairs even in age; seeds 8-10, pl 26-32 m ae ong, 15-18 mm. wide, one end slightly dine Tes ls hg other terete. Climbing over shrubbery va digger pine belt Pcie petal gad of the ponderosa a, Upper Sonoran mba —— slopes of the Sierra avade from hastens Cou n County, California. Type locality: Mari County. Collected by Congdon. March-April. 5075. Marah horridus 70 CUCURBITACEAE 4. Marah macrocarpus (Greene) Greene. Chilicothe. Fig. 5076. Echinocystis macrocarpa Greene, Bull. Calif. Acad. 1: 188. 1885. Micrampelis leptocarpa Greene, Pittonia 2: 282. 1892. arah macrocarpa Greene, cartes Bot. Obs. 2: 36. 1910. Stems reaching a few meters in length, oe ois ae or cane is scaberulous or the ne growth een with septate hairs and becoming glabra Leaves broader ee long, 5- O(11) cm. broad, the sinus obtuse, raharanty open, oats 5-7 (8)- pene Fay the middle or below, rounded between. ae Thee these 7-15 m road, broadly acute or obtuse or bn round t the pex and usually Se ninaicae, ‘oe margins entire or shallowly lobed or toothed; staminate inflorescence with big wers more or less get arranged, pubescent, paniculate or subpaniculate e e br i rs white, 7-13 and edicel 1-2 cm. long ; fruit ayn cm fae ovoid or eats deciely echinate with: cae dl broadened spines of different lengths, so ong, vestiges of = ubescence remain n body of fruit and bases of spines; oe B44) ae 13-20 mm. long, 8-10 mm. wie: Hillsides and canyons, mainly Upper Sonoran Zone; - rom Santa Barbara County south to San Diego County, California, and Lower California, including the islands off the coast, where it occurs with M., cong ies east- ward to the desert slopes of the mo Arte Ng of southern California. Type locality: Cuacamonga, San Bernardino ee California fn —April. ens from the sine off Los Angeles and San Diego Counties appear M. rocarpus var. ‘aie Pain Boeken. Madrono st 134. 1955 (M. micranthus Dunn, Kew Bull. 1913: 80. 1913), y species with small flowers and seeds 10-13 mm. long, originally described from Cedros Island, Lower California. 5. Marah eee (S. Wats.) Greene. Island Man-root. Fig. 5077. + Catalans ererers tert S. Wat c. Amer. Acad. 11: 115, 138. 1876. ‘ah Greene, Leas eek. Obs. 2: 36. 1910. Marah major ene Kew Bull. e2.451, “19135, Marah macrocarpus var. major ee. Madrofio 13: 134. 1955. Stems striate, glabrous or nearly so, reaching several meters in length. Leaves orbicular in th outline, more or less scaberulous above, the new growth spar. sely pubescent and becoming glabrate, 12-32 cm. broad, ie see obtuse or rounded, Bae y chesed | in the larger leaves, mostly 5-lobed bov b (occasionally 3- or 7-) to above or nea middle, the lobes broadly tria neuer, mucronulate, he margins usually triangular-toothed, oe sinuate-lobed, more ra nad ire; staminate inflorescence loosely flowered, pubescent, often becoming glabrate, racem S iteutate 2 cm. long, often sede late in the season; staminate flowers white, p ie tank wie out, 18-30 mm. wide, the lobes y or narrowly pe arom the pedicels 10-20 mm. long; pistilla te flowers white, equaling ie | staminate in size, pubescent without, ovary and beak p ubescent ; fruit ovoid, 5-7 cm. long, echinate with stout spines of eatin g len gths, the longest ant 1.5 cm. in length, the ‘idky of the fruit and spines for the most part pubescent ; seeds 2-4, oval and somewhat com- pressed, 28-34 mm. lon i M rc and ¢ s, Upper Sonoran Zone; islands off the coast of southern California from Santa Barbara County uth to "Guadalupe Island, Lower California; more abundant on Santa Catalina, San Clemente, and San Nicolas Islands. Type locality: Gu ada lupe Island. Fe b.—Au: 6. Marah inérmis 1 oie Dunn. Foothill Man-root. Fig. 5078. Echinocystis inermis Congdon, Zoe 5: 134 Echinocystis scabrida Eastw. Bull. Torrey cb 20: 500. 1903. Marah inermis Dunn, Kew Bull. 1913: Echinocystis fabacea var. inermis pai _ sk 2: 554. 1936. Stems striate, glabrous, long, and branching, young growth beset with soft septate hairs which persist to some extent about the inflorescence and fruit. Leaves glabrous or scaberulous, 5- 10(13) cm. broa width greater than the length, 5-7-lobed to about the kai the inc pair of lobes = deeply 5 cated the angle between the lobes typically narrow, the margin of the lobes racemose on shorter infloresc s, 11-35-flowered ; staminate flowers bright white, 1-1.5(2) cm. broad, the br sin ‘eee elis, lobes ee than the shallow, gprs gis tube, rotately spreading and plane, papillose on the inner surfaces; pistillate flowers similar what larger, the pedicel 1-3 cm. long; ovary globose, beak abru ne og attenuate, the young prickles po the beak with soft hairs; naxtuce fruit globose, 2-4-seeded but mostly 4, a short, cone-shape ak go times evident in the growing fruit, 2.5-4 cm. broad, covered with acicular prickles 5-9 mm. lon these ge AB, glabrate in age; seeds about 1.5 cm. long, oblong-ovoid, scarcely compressed. Climbing on shrubs or often prostrate on grassy slopes and in fields, Upper and Lower Sonoran Zones; lower foothills of the ne pen Nevada from Butte counts south to Kern County and the Inner Coast Ranges from Yolo County to Kern County, California. Type locality: Sherlocks, along the canyon of the Merced River, Mariposa neg? — Ma: M. fabaceus var. agrestis (Greene) Stocki and the f of the two scarcely distinguishable. ranges eo the two forms overlap. The white flowers ete usually se deeply lobed tanec jeaeniahe! it from M. fabaceus var. hg ager and t ar ° - the fruit as well as the white flowers and more deeply lobed leaves distinguish it from M. fabaceus var. fabac GOURD FAMILY 71 5076. Marah macrocarpus 5077. Marah dal 7. Marah fabaceus (Naudin) Greene. California or Valley Man-root. Fig. 5079. Megarrhiza californica Torr. Pacif. R. Rep. 6 1858. oper nudum) Echinocystis fabacea Naudin, bp n. Sci. Nat. y i 154. pl. 9. 1859. _ garrhiza californica Torr. ex S. Wats. Proc. Amer. Acad. 11: 138. 1876. arah fa ware Greene, ea Bot. Obs. 2: 36. 1910. Stems reaching several meters in er nearly glabrous to densely scabrous, pe Le mph nsely beset with soft septate hairs wh io) more or — — - ages cept about 1 18 as broa wi ith s ell as vary, the beak also sisting 4 ruptly ee the ovary ; fruit globose, with stout, stiff, usually pubescent spines 6-12 in lengtii; seeds oblong-ovoid, bordered lengthwise with a shallow groove or line, somewhat flattened at one rath Slopes and washes, Upper Sonoran Zone; Outer Coast Ranges from Northern Sonoma County to Santa Bar bara County, California; ae eported from the aged Islands. Type locality: cael ae Rawson plants caltivated t Paris, the material presumably collected from the San Francisco Bay region. Feb.— arah fabaceus var. agréstis (Greene) aa Madrofio 13: 130. 1955. ps lee fabaceus var. penal Greene, Fl. Fran. 236. ea Echinocystis fabaceus var. agrestis co Zoe 5: 133. 1901.) Leaves as in the species, mostly ara a 5 cm. broad; Gowers greenish; fruit 1.5—2. broad, globose, a spines slender and pci henge Eg mm. oe. osble “1-4 gy td usually 2. toes slopes a Inner Coast Ran and the adja- cent valley from Glenn County to San Luis Obispo County and aponreees less common in the Sie > ae a foot- ol from oe. County to Fresno County, and also on the coastal slopes in Santa Barbara County; Caio. Ie at mature fruit not setpeniele t from the species. Type locality: “‘grain field of the valley of the San pani 72 CAMPANULACEAE 4. BRANDEGEA Cogn. Proc. Calif. Acad. II. 3: 58. 1890. Perennial monoecious vines with simple tendrils. Leaves 3-5-parted or -angled. Stami- nate flowers in racemes ; pistillate flowers solitary, often axillary to the staminate racemes. Style short; stigma hemis . Fruit asymmetrical, long-beaked, tardily and irregu- larly dehiscent, l-seeded. Seed irregularly obovate, warty or echinate. [Named in honor of T. S. paisa a pioneer western botanist. | A Pp of the Sonoran Desert region. Type species, Elaterium bigelovit S. Wats. 1. Brandegea bigelovii (S. Wats.) Cogn. Brandegea. Fig. 5080. Elaterium Rap ovit S. Wats. Proc. Amer. Acad. 12: 252. 1877. Echinocystis parviflora S. Wats. op. cit. 17: 373. 1882 Cyclant sie ra monosperma Brandg. Proc. i Acad. Ul. 2: 159. 1889, Brandegea bigelovii Cogn. op. cit. 3: 58 Brandeg p Cogn. op. cit. ty Slender- inieania perennial vines of dense growth from large thick roots. Leaves Bae hastate to quadrate in outline with deep narrow sinus, 3-5-cleft or merely angled, 1-5.5 cm. lon the _ er — si _— er, sharply triangular-acute, smaller leaves often cleft nearly to the Me with n linear-acute divis wns the upper surface pair sly pustulate; petioles shorter than tbe leaves pistillate and staminate flowers similar, minute, whitish, rotate and deeply lobed, about 1.5 mm. br. oad ; fruit straw- ser 2m sg to ahr a a — Doe of fruit more or Cs wohinate about equaling the beak in benetii: ‘seeds about 5m Sandy arroyos and canyons, Lower pSniamist Zone; Colorado Desert, i am to the western parts . ee ead py and Yuma Soe, Arizona, south to Lower California and Sonora. Type locality: ‘‘the Low Colorado Valley.””, March—Apr Family 148. CAMPANULACEAE. BLUEBELL FAMILY. Herbs, shrubs, or trees with alternate, simple (rarely pinnatisect), exstipulate leaves. Flowers perfect, epigynous. Calyx 5-parted. Seay gamopetalous, regular, or zygomorphic. Stamens free or united. Style 1. y 2-10-celled, rarely 1-celled, the — on axile placentae. Fruit a capsule or ene a eiees seeds numerous ; em- bryo stra A ae a about 60 genera and 1,600 species, widely distributed in subtropical and temperate regions. Corolla regular; — on Giampente: free. Subfamily 1. CAMPANULOIDEAE. Corolla irregular; an united into a tube (anthers free in Nemacladus and Parishella). Subfamily 2. LOBELIOIDEAE. Subfamily 1. CAMPANULOIDEAE. BELLFLOWER SUBFAMILY. Ours herbs, with acrid or milky juice. Flowers racemose, spicate, paniculate, or solitary, perfect. Calyx-tube adnate to the ovary, its “ae mostly 5-lobed or 5-parted, the lobes equal or slightly unequal, valvate or imbricat e bud, commonly persist- ent. Corolla gamopetalous, regular, usually 5-lobed or aca. blue or white, inserted on the calyx-tube where that becomes free from the ovary. Stamens 5, inserted with the corolla, alternate with the corolla-lobes; anthers 2-celled, introrse, separate. Ovary usually 2—5-celled. Style 1; stigmas ae usually distinct. Fruit ac capsule or berry, usually opening by clefts or pores; seeds minute and numerous. Cap t by small valves or circular perforations below the calyx-lobes. Plants Deon Lag Pacific States) perennial (except C. exigua and C. angustiflora); earliest flowers not cleistoga- Coratia. corm Phil lobed above or to the middle, rarely a little below, the lobes triangular to oblong- lan 1. Campanula. Corolla ten vided, the lobes teal er 3. Asyneuma. Plants annual; st flowers on the lo t lei Capsule ae to narrowly cabinate jeaeieaner divided pene: the meine: 2. Triodanus. ssa ase iach nd broad, nearl 4. Heterocodon. Capsul pex of the calyx-limb within ‘ie lobes. 5. Githopsis 1. CAMPANULA [Tourn.] L. Sp. Pl. 163. 1753. Perennial or annual herbs with alternate or basal leaves. Flowers large or small, soli- tary, racemose or paniculate, regular, blue, violet or white. Calyx-tube adnate to the ovary, BLUEBELL FAMILY 73 hemispheric to turbinate, prismatic, the limb 5-lobed or -parted or rarely 3—4-parted. Corolla campanulate or rotate, more or less deeply 5-lobed. Stamens 5, free from the corolla; filaments usually dilated at base; anthers distinct. Ovary inferior, 3—5-celled; stigma 3—-5-lobed. Capsule partly or wholly inferior, crowned by the persistent calyx-lobes, opening by 3-5 small valves or perforations situated on the sides near the top, the middle, or “ ie LN: ane cumeutiye of campana, the Latin ‘Gaal for bell. of cies, mostly in the boreal and north temperate regions of both the Old and New Worlds” T306 sari ta ap latifolia L. Perennia a lenin than the corolla; reflexed lobes of the corolla nearly equaling the tube. 1. C. scouleri. ci Fase than to about the length of the corolla; lla-lot h than the tube, erect or nearly es crenate or crenate-serrate, the stems and leaves store retrorse- scare w — femme, ‘indoles or bog plants. ay alifor Leaves cotire or — das retrorse-ciliate; not weak-stemmed, swamp or bog I inear and sessile, the ‘er orbicular to ovat her oe ale a st rotundifolia. Leaves essentially ali a sais yd few apices e; plants completely glabrous; natives of northern a . C. wilkinsiana. Calyx-lobes usually toothed; plants variously pube sae, ans arene the inflorescence and leaf-bases; plants of Washington ao northward. Calyx-tube more or less pubescent with long aes: corolla 15-25 mm. long. . C. lasiocarpa. Calyx and pedicels with rather sparse bristly puberulence; corolla 10-12 mm. Jong. pipert L. Leaves and calyx-lobes enti Plants uniformly Osta ener pubescent throughou 7. C, scabrella, Plants nearly glabrous but some or all of the nis leaf-bases ~—: ae calyx-tube usu- ally somewhat puberulent, a parryi a ohasncie: Annuals. Corolla twice the length of the calyx-lobes; plants branching above the base. . C. exigua. Corolla about the length of the calyx-lobes; plants branching from the base. 10. C. angustifiora. 5078. Marah inermis 5079. Marah fabaceus 74 CAMPANULACEAE 1. Campanula scotleri Hook. ils ee or Campanula. Fig. 5081. Campanula scouleri Hook. ex A. DC. Monog. Camp. 312. Campanula scouleri a hirsutula Hook. Fl. Ni Amer. 2: = — "125. 1834. Campanula scoulert B glabra Hook. loc. c Stems from sl ‘ian erect or decumbent at base, 1-3 dm. high, slender, simple o rarely “pele below, hethiape glabrous throughout or nearly so. Leaf-blades broadly — ein .9-4.5 cm. long, sharply serrate, often remotely so, acute or acuminate apex, wed at base toa park pase 6 often as long as the blade; ‘flowers a se a terminal, usu ally few-flowered, raceme-like gees ial the p me eA Med slender a. ga su seal firactlee near the middle; yx-tube Deena ulate, 2-3 mm. long, es Patil edeliats, about twice as long, erect or spre reading corolla light ee to white, the lobes reflexed, as long as the tube, ees Sake broadly o Open woods and slopes, mainly ee Roe! ween southern Alaska southward west of the Cascade Scent to Sg aa ae: _ the Coast Range, California, and also to Sierra County in the Sierra Nevada. Type locality: Fort Vancouver e Columbia River, Waskineun, June-Aug. 2. Campanula californica (Kell.) Heller. Swamp Harebell. Fig. 5082. Wahlenbergia californica Kell. Proc. Calif. Acad. 2: 158. fig. ae 1862. Campanula linnaeifolia A. Gray, Proc. Amer. Acad. 7: 366 Campanula californica Heller, Muhlenbergia 1: 46. Perennial, from slender creeping rhizomes, with slender pepe stems simple or sparingly branched above, 1-4 dm. long, retrorsely hispid o on the angles or ribs. Leaves oblong-ovate to obovate, seen or subsessile, narrowed below to a petiolar ig paehsce es broadly and shallowly crenate-serrate abov iddle, entire elon 0.5-2 ng, the margins more or less retrorsely t e the mi — . long, hispid; flowers solitary, terminating the branches or on slender, elongated, axillary ae nin calyx-tube turbinate-campanulate, the lobes subulate, 4-5 mm. long; corolla pale blue, 10-15 m long; style inc = d, 6-7 mm. long; capsule broadly obovoid to globular, 10- ribbed with thin. walled interspace Swamps and along the coast, Humid Transition Zone; greed c¢s to =e Reyes, Marin County, California. Type locality: ‘‘coast range at some point north of San Franc June—Sep 3. Campanula ee ern L. Scotch Bluebell. Fig. 5083. Campanula rotundifolia L. Sp. Pl. 16 53. Campanula petiolata A. DC. Monog. nae 278. 18 Campanula sacajaweana M. E. Peck, Proc. Biol. a aeask: 50? 123. 1937: Perennial, with f Seceon acon perme rine arising from root-crown with slender na ms ht from offshoots, less commonly anesting on the hai stems, — i ag on petioles baie surpassing the blade, ovate to nearly orbicular, cordate to subcor t base, irregularly sinuate- entate to sinuate; lower stem-leaves short-pe beatae spatulate, eeiancosial te or lanceolate and often with a sinuate margin, merging into the ~— sini linear or linear-filiform upper leaves ; flowers in loose panicles with slender branches, sometim s solitary ; calyx-lobes narrowly oes) na cm. long, the tube much longer than the lobes; capsule nodding, spenne by valves near the base. eado ows or jTocky slopes, Transition and Boreal waraa! Alaska to the mountains of northern California and nent; also northern Europe and Asia. Type locality; Punane. uly—Sept. A highls variable and widespread oir —_ with many tnd big a gre igh-mountain eda. are low in stature, have lanceolate leaves, and are often 1-flowered. The taller plants of lower elevations are usually char- acterized by more flowers in the panicles and by * peal to linear-filiform leaves. Some large-flowered toview note the Washington, coast and in the Columbia Gorge with long, filiform, divergent calyx-lobes resemble the Alaskan form of the species. 4. Campanula aad Greene. Wilkins’ Harebell. Fig. 5084. Campanula wilkinsiana Greene, Pittonia 4 1899 Campanula baileyi Eastw. Bull. Torrey a a 525. 1902. Plants from slender creeping rootstocks, the leafy stems slender, 3-14 cm. pe glabrous throughout. Leaves sessile, 8-15 mm. long, cuneate to obovate, caesely few-toothed on the u € the ue, or a little below, the lobes iggy mee and spreading; style a the length of the corolla; capsule turbinate, about 7 m ong, the calyx-lobes shorter bot the oie and erect. In meadows and along daca: Canadian and Hudsonian Zones; Mount Shasta, Siskiyou County, aa Trinity Mountains, Trinity County, California. Type locality: head of fence Creek, Mount Shasta. Aug. and 5. Campanula lasiocarpa Cham. — Alaska Bluebell or Harebell. Fig. 5085. Campanula lasiocarpa Cham. Linnaea 4: 39. Perennial, the stems 4-18 cm. hi iva arising from a creeping der goose Leaves 2-7 cm. long, spatulate to narrowly oblanceolate, tapering to the base to a winged peti the uppe as — and bract-like, the margin laciniate-denticulate to laciniate with rather ce. ‘teeth, glabrous except for marginal white hairs along the petioles, vaca more abundant on the bracts and upper aaven: BLUEBELL FAMILY 5080. Brandegea bigelovii 5081. Campanula scouleri 5082. Campanula californica Bic a i 5083. Campanula rotundifola 5084. Campanula wilkinsi 5085. Campanula lasiocarpa 76 CAMPANULACEAE flowers usually solitary, terminating the stem ; Hagia tube broadly obconi on long, densely beset with lo nage villous hairs; calyx-lobes ras . long, becoming div gent, “laciniate, — or less ciliate on the eer corolla blue, 15-28 m mm. long, the lobes en -ovate, 5-1 10 m long ; capsule sc ca * Grassy banks and upla ‘ort Boreal Zone; northwestern Asia and Alaska south to western Alberta and British Columbia and in Snohomish County, Washington (Thompson). Type locality: Unalaska. July—Se ept. 6. Campanula piperi Howell. Olympic Harebell. F ig. 5086. Campanula piperi Howell, Fl. N.W. Amer. 1: 409. 1901. Stems clustered from a much-branched rootstock, 3-10 cm. high, leafy to the om, the leaves scarcely reduced in size, herbage glabrous throug py or more or less finely scabr bove. m. long, en ew at prod bracteate racemes yx-tube obconic, short, the lobes subulate to linear- lanceolate long, sparsely oe Gnu dteas: corolla blue, open-campanulate, ab: lon style slightly en erat to shorter than the corel: stigma- -lobes strongly talons ~ohenaties capsule subglobose Rocky crevices of cliffs, Boreal Zones; Olympic Mountains, Washington. Type locality: “On cliffs, Mount Steele, Olympic Mountains, Washington.” July— Aug. 7. Campanula scabrélla Engelm. Rough Harebell. F ig. 5087. Campanula scabrella Engelm. Bot. Gaz. 6: 237, 1881. Campanula uniflora Jepson, Man: Fi. Pi. Calif. 973. 1925. Not C. uniflora L. Plants 3-12 cm. high; stems usually tufted, arising from the much- dates underground stems, densely short- -bristly pubescent Spee out. poh, clustered at base of stems, entire, lanceolate to oblan “pean acute < pe see r ac obtuse, = 5Scm —5 mm. arted, thickish ; flowers solitary ad terminal, or i seaegy epals ans 2-8 mm. long ; corolla — oh ies oe ca “Chale! clipsoidal foe pees c 67 mm. long, opening ase the summit. ocky alpi lopes, Boreal Zones; Wenatchee Mountai Adams, belo sancti south on high peaks of “oe Mouseinn to Mount Shasta tidy Mount Eddy, akiyen County, Califor Type locality: ‘‘On bleak ed a0e- s of Scott Mountain, west of Mount Shasta.” June— oc Neh ates between this and the following taxon are Socuaiouaty found in Chelan and Kittitas Counties, ashington. 8. Campanula parryi var. idahoénsis McVaugh. Parry’s Northern Harebell. Fig. 5088. Campanula parryi var. idahoensis McVaugh, og ma Club 69: 241. 1942, ?Campanula rentoniae Senior, Rhodora 51: 302. Plants perennial from = h- iaian slender ae the sparsely leafy, glabrous, simple stems 6-35 cm. high arising from leaf-clusters at the apex of the rootstocks. Leaves sessile, lanceolate, eligntee ie eebalete. or oblanceolate, sets ; a pee, aewiag to the base, the | some- e , O- ng, upper leave sometimes bract-like, glabrous except for sparse ciliae mostly on the basal margin, these usually not present on th per leaves ; flowers ry, terminating the stem, sometimes 1-3 on axillary branchlets; calyx onic or campanulate, often a fe —- — ; calyx-lobes narrowly subulate to linear, erect, shorter than is tube, lengthening som and :often spreading in age; corolla b campanulate, 9-15 mm. long, the lobes about paar the length of the corolla; filaments with ~ expanded ciliate b bes capsule 7-11 mm. long, the 3 valvular openings near the summit of the capsule. Open rocky slopes, Boreal Zones; Montana and Idaho west to Chelan and Kittitas Counties, Washington. Type locality: near Friday’s Pass, Idaho County, Idaho. July-Aug os Campanula exigua Rattan. Chaparral Campanula. Fig. 5089. Camp: igua Rattan, Bot. Gaz. 11: 339. 1886. Annual, 7-15 c igh, simple below and oh eae branched above, or rarely branching foonuhne especially below, with spreading or somewhat reflexed hairs. Basal rom t the base, leaves obovate or oblong-obovate, ts A those of fg ater: stem Lau oblong, toothed or entire, the upper most often subulate ; flowers axillary and terminal ; calyx-lobes subulate, 4-5 mm. long, m. twice as long as the turbinate tube, glabrate ~ bel kishie ssed-hispi gra corolla a 8-12 m long, the lobes ovate or oblong-ovate, acutish; stamens mm. lon amen s dilated below the middle nd ac a oe — style about equa sine ai rim of the rae reeling; capsule somewhat urceolate, o g by 3 valves just above th € mi Roc py and . an Peay Arid Transition Zon of the Coast Ranges, mainly Inner Coast Ranges; Mount St. Helena, Mount Tamalpais, and the Senst Diablo and noun Hamilton Ranges, central Cali- fornia. "Type locality: Mount Diablo. May—June. 10. Campanula angustifléra Eastw. Eastwood’s Campanula. Fig. 5090. Campanul gustifi Eastw. Proc. Calif. Acad. III. 1: 132. Sh Ii; Le — 1898. Campanula angustifiora var. exilis J. T. Howell, Leaflets West. Annual 1, 18-25 cm. high, usually diffusely stereriore pes oe bas the upper branches erect to spreading and the stems with short hispid hairs on the angles. Leaves glabrous, sessile, 3-10 BLUEBELL FAMILY 77 m. long, narrowly to broadly ovate, coarsely and deeply dentate below, less deeply to subentire on on the upper reduced leaves; flowers solitary, lateral or terminal on the branches; pedicels 0 long; calyx 4-6 mm. lon g, the subulate aim exceeding the tube; corolla blue, tubular or slightly. erica alee: psi uous, shorter than to equaling the calyx-lobes; stamens 2 mm. repos about equaling the filaments; style heirae from the base; capsule strongly ribbed, about . long. 6m chaparral slopes, Arid Transition Zone; Marin County to Santa Cruz County and at The Pinnacles, San Benito County, California. scoot locality : paler Tamalpais, Marin County. May-June. The more slender plants with longer t the variety. 2. TRIODANUS Raf. New FI. 4:67. 1838. Annual herbs with erect or reclining stems, the branches if any basal. Leaves alternate, toothed or entire. Inflorescence spiciform, the axillary flowers 2-bracted, sessile or near] so. Early flowers small and cleistogamous, the later with a blue or purple, somes rotate corolla. Calyx-tube narrow, the lobes in the earlier flowers 3-4, in the later ones 4-5. Corolla 5-lobed or -parted, the lobes imbricated in the bud. Filaments dilated to a ciliate base; anthers separate, linear. Ovary 3-celled, or rarely 2-celled or 4-celled ; stigma usually 3-lobed. Capsule opening at the apex or near the middle. [Name Greek, meaning three unequal teeth. ] V/7 5086. Campanula piperi 5087. Campanula scabrella 78 CAMPANULACEAE A genus of 8 species, i of Europe, the Mediterranean region, and North and South America. Type species: Campanula perfoliata L. Leaves sessile, not cordate-clasping; pores near the top of the capsule. 1. T. biflor Leaves mostly cordate-clasping; pores near the middle of the capsule. 2. aE, 1. Triodanus biflora (Ruiz & Pav.) Greene. Small Venus’ Looking-glass. Fig. 5091. Campanula biflora Ruiz & Pav. Fl. Peruv. 2: 55. pl. 200. 1799. Specularia biflora Fisch. & Mey. Ind. Sem. Hort. Petrop. 1: 17. 1835. Dycmicodon californicum Nutt. hs Amer. Phil. Soc. II. 8: 256. 1843. Dycmicodon ovatum tt. loc Triodanus biflora Greene, Mite n. eee Reg. 230. (Feb.) 1894. Legouzia biflora Britt. Mem. Torrey Club 5: 309. (Oct.) 1894. Plants scabrous on the eine Knee glabrous or beseey 38 so par as stems simple or branched, ale der and often weak, 1.5-6 dm. high. Leaves ovate to oblo ong or the uppermost Beeeciate, sessile, 6-20 mm. ty entire, or te ae crenate- serrate with yes lo cae ie af teeth; earlie r flowers with 3-4 ov ate to lanceolate calyx-lobes, the ones with fnccolae e-subulate calyx -lobes ; capsule 6-10 mm. long, " Ditone Sdn "bpenida by Ride ose under th 1 Dry soils, mostly in disturbed places, Transition and Sonoran Zones; southwestern Oregon southward < Lower Californ nia and eastward across the southern United States to Virginia and Florida; also Mexico and Sou America. Type locality: Peru. April—July. 2. Triodanus perfoliata (L.) Nieuwl. Venus’ Looking-glass. Fig. 5092. Campenni es DL. Sp.P1. 169. 3. DC; One: oa ~~ 1830. Trisdawns rupestris “3 New FI. 4 Legouzta perfoliata Britt. Mem. Torrey ees 309. 1894. ehuae perfoliata Ni euwl, Am idl. Nat. 3: 192. 1914. Ste imple or bra a ai rather stiffly erect or sometimes decumbent at base and more or less spreading, hi Pic danetiatly on the angles with * reading, bristle-like hairs of uneven length, L, o suborbi s rd y cordate-clasping Rs - (e) ee) = te ct ee & 8] Cal al O83 =) a8 imes entire, short-hispid on the veins and margins; fl 3 in xils, sessile; upper flowers (the later ones) with 5 (rarely 4) rigid, elastance eolate, acuminate calyx-lobes and a rotate, lue or ite c gs d, arlier flow all ith 3-4 shorter calyx-lobes exceeding the rudimentary corolla ; capsule oblong to narrowly turbinate, 4-6 mm. long, tardily opening near the middle; eee lentic ular 9 opas and rocky ban Transition iid Upper Sonoran Zones; British Columbia southward ica Washin hatch d Oregon to pede ne fia: mboldt Chahibes: northwestern California; eastward across the c tinent. Type secehiies Virginia. April-Jul Be erin Ces Griseb. & Schenk in Wiegm. Archiv Naturgesch. 18°: 335. 1852. ennial or sometimes biennial herbs. Inflorescence an interrupted panicle or some- rotate-spreading to reflexed. Style pilose, the stigma trifid. Capsule 3-celled, dehiscing by pores which are near the apex, the base, or at the middle of the capsule wall. [Name ee ra a Greek, the application obscure. | t 40 species, mostly natives of southeastern Europe and Asia Minor but occurring also in snes per ea ng Pacific States. Type species, Asyneuma canescens Griseb. and Schenk. i, —— prenanthoides (Durand) McVaugh. California Harebell. Fig. 5093. Campanula prenanthoides Durand, Journ. Acad. foo II. 3: 93. 1855. a Campana filiflora Kell. Proc. Calif. Acad. 2: fe mpeewie pores Repet, pen aE 21239. 18 McVaugh, Dalionia a 23: 36. 1945. ae are | from creeping r ocks, 2-8 dm. high, erect or sometimes decumbent in shade forms, simple or much hee pores longitudinally ribbed or angled, glabrate above, rather sparingly Pa tig with aves som pie se sei bende oe ee 2 2 be dt -petioled, ovate- lance: boat i nce a og 5 eate ase, 1-4 cm. long, sharply ait ee ea ota serrate, Saar hispiculons pte, * cadaciate on tte veins and margins ; flowers in small clusters forming an interrupted, terminal, racemose panicle or simple, few -flowered r , the lower subtended b all ems the upper by small acuminate bracts yx cam- panulate, the re subulate, as long as or long an the tube; corolla bright blue, 10-12 mm 1 r, two to three times as long as the tube; styl ex : rved; on, e nea ‘ capsule NenGaiheric to short-oblong, subcordate at base and thin-walled between the nerves. Pee woods, Transition Zones; British Columbia south, mostly west of the Cascade Mountains, to Tulare County in the Sierra Nevada and to Monterey County in the Coast ges, California. Type locality : Nevada County, California. June—Aug. BLUEBELL FAMILY 79 4. HETEROCODON Nutt. Trans. Amer. Phil. Soc. II. 8: 255. 1843. A small delicate annual with weak slender stems and sessile or partly clasping, sub- orbicular leaves. Flowers of two forms, the earlier ones with rudimentary corollas and ate t foliaceous, much longer than the tube, 3-4 in the early cleistogamous flowers, 5 in the later open ones. Corolla blue, broadly campanulate, 5-lobed. Filaments of the saree ciliate at base. Styles as in Campanula. Capsule 3-celled, - -angled, goat at base b valve-like opening in the capsule-wall over adjacent loc See merous. Nave from two Greek words meaniiy different and bell, for the two kinds of bell-etin get flowers. } A typic g f western North America. 5092 5090. Campanula angustiflora 5092. Triodanus perfoliata : 5091. Triodanus biflora 5093. Asyneuma prenanthoides 80 CAMPANULACEAE 1. Heterocodon rarifldrum Nutt. Heterocodon. Fig. 5094. Heterocodon rariforum Nutt. Trans. Amer. Phil. Soc. IT. ed 1843. Specularia rariflorum McVaugh, Leaflets West. Bot. 3: 48. eli snes uN with almost —— to piace ies hed, almost filiform stems 1 “ dm. long, sparsely hir . Leaves sessile, cordate or gr clasping, orbicular or nearly so, m. broa pecan serrate te th br ea ne bein ra eth; ers Phage ae calyx- yal of the tess flowers riangular-ov 3-5 mm. long and jelaces ous in gy , often with a road, bristle-tipped be calyx- net broadly obpyra rar corolla of the re: pala light Kio or the lobes often da rker, pha ¢ qualing oe alyx-lobes; capsule short and a ad. Moist banks and boggy meadows, mos stly ae sition Zon British Columbia and Idaho southward through the Pacifi Re States and Nevada to the oo of San Hines: yee Dalene. Type locality: ‘‘Grassy plains of the Wahlamet and Oregon.’”’ May-—Jul 5. GITHOPSIS Nutt. Trans. Amer. Phil. Soc. II. 8: 258. 1843. Small annual herbs with angled stems and cuneate-obovate, sessile, usually inconspicu- ous leaves. Flowers all alike, solitary in the axils. Calyx with the tube 10-ribbed and the 5 lobes long, narrow and foliaceous. Corolla caibrilsie-aetaal athe 5-lobed. Stamens with short filaments, dilated at base; anthers long and narrow. Ovary 3-celled; stigma 3-lobed. Capsule clavate or obconic, strongly striate-ribbed, crowned with rigid calyx -lobes, some- what shorter to longer than the tube, opening in the summit by a circular orifice left by the falling away of the style. [Name Greek, from the generic name Githago, on account of the similarity of the calyces. ] A genus of pooper? 3 or 4 variable species, natives of western North America. Type species: Githopsis speculartoides Nut Capsule narrowly obconic to clavate, 2.5-4 mm. wide at summit; calyx-lobes as long as and usually longer in age ian the calyx-tube. olla shorter than the calyx-lobes or rarely equaling them. 1. G. specularioides. Fe half again as long as the calyx-lobes. 2. G. pulchella. Capsule but little enlarged at the summit; calyx-lobes in fruit shorter than the tube. 3. G. diffusa 1. Githopsis specularioides Nutt. Common Blue-cup. Fig. 5095. Githopsis specularioides Nutt. Trans. Amer. ris Soc. IT. 8: 258. 1843 Githopsis speculariotdes B hirsuta Nutt. loc enth. Pl. Githopsis calycina var. hirsuta Benth. loc. cit. (Nomen nudum) Githopsts latifoliuns Eastw. Proc. Calif. Acad. IV. 20: 154. 1931. Githopsis specularioides subsp. candida Ewan, Rhodora 41: 308, 1939. Annual with stems pestueeyy branched, erect or the branches ascending or sometimes some- what decumbent, simple or nearly so particularly in dwarfed plants, 3-15 cm. long, light green or grayish green (glabrate Se hirsute plants in same col naar glabrate or more or less densely hirsute and hispid with spreading or deflexed hairs, especially on the ribbed angles of the stem i ile, more or t -20 mm. long, abu to o 1 on younger plants, mostly deciduous in yi calyx-tu m. lon ng, pro omg ee ribbed longi- tudinally in wine hee rae ee mm. ape a eget calyx- lobes oat mostly as long as the tube r longer in fruit and m or less spreading; corolla blue or ste whi te 3-10 m os oe the lobes toe tha an the fies ng tube, usu ati Shoeter than the calyx-lobes or savading them Mostly in clay or adobe soil on dry slopes, often in burned areas in the chaparral, Sonoran and Transition es; western Washington (Chehalis Cou uses) and the Blue Mountains and western Oregon Putte nel through the Coast Ranges and Sierra Nevada foothilis of California to Se southern border of the state. Type locality: “‘plains of the Oregon, near the outlet of the Wahlamet.”’ April— 2. Githopsis oe gape Large-flowered Blue-cup. Fig. 5096. — pulchella racees sage 38: Githopsis specularioides glabra emai ie Fl. Pl. Calif. 974. 1925. Annual, 5-15 — soak widely — branching from the base and ay in vigorous plants as wide as high, iin erat glabrous or hirsute. gies sessile, 2-10 cm. long, lanceolate to linear-lanceolate, few-toothed, rarely pers e, the lower leaves withering at anthesis ; flowers sessile or s short-pedincled in the leaf-axils ; r eron es calyx sia mm. long fruiting calyx-tube 1 irsut inea r , glabrous or wi glabrous or hirsute, a2 CE preading in age, straight o Baics § pes ‘bhi 15-20 m ong. - opes, Upper Sonoran and Arid Transition Zones; at of the Sierra Nevada from Say Couaty 3 Merk a County, California. Type locality: iy wdhedagals Bridges 153, possibly near the Calaveras Sequoia Cisne a ward Bridges is known to have visited. May—Ju 3. Githopsis diffisa A. Gray. Southern Blue-cup. Fig. 5097. Githopsis diffusa A. Gray, Proc. Amer. Acad, 17: 221. 1882. Annual with slender stems, simple or branched above and below with ascending —— herbage glabrous except for occasional hairs. Leaves narrowly eet serrulate ; flowers BLUEBELL FAMILY 81 . Heterocodon cable 5096. Githopsis pulchella see Githopsis specularioides 5097. Githopsis diffusa tered, sessile or short-pedicellate on the branches ; flowering calyx 8-10 mm. long, the lobes on ean oe the length of the tube; fruiting calyx 9-11 m m. long, essentially linear, little edireen the 5 OB cuously r rib ; fruiting ‘calyx- -lobes Soe ascending or spreading, shorter than fie “tube n fruit; corolla “light purple,” 3-5 mm. long, shorter than the calyx-lo seabed and wanes eg Arid Transition Zone; San Gabriel Mou er _ Angeles and San Ber- nardino ‘Comnnien: California. Type locality: ees Cacamonga San Gabriel Mountai May-June. se —_— gilioides fe Apel 41: 31 w plant much branched from the cas with persistent, hirs glabrate leaves; cathe 4-5 mm. ce cinenlion: 4 the calyx. Known only from the San Gabriel Moun- tains, southern California. Type locality: forks of the San Gabriel River. Possibly a fucitant hitentulons form us Githopsis filicailis Ewan, op. Feb — lax annual with few widely spreading branches and minute flowers 3- 6 mm. long and calyces 355 long. Known ag Mission Canyon, San Diego County, California, the type locality, and from Vallecito (Orcutt), ae Californ Subfamily 2, LOBELIOIDEAE.* LoBELIA SUBFAMILY. Ours herbs with acid juice, alternate or basal, exstipulate, eee leaves and spicate or racemose, often leafy-bracted flowers. Calyx-lobes 5, equal; calyx-tube adnate to or partly free from the ovary, turbinate, ovoid or hemispheric. Conde gamopetal- ous, usually persistent on withering, often split down one side, 2-lipped, with 2 lobes * Keys and taxonomic treatment are based largely on the works of Rogers McVaugh. 82 CAMPANULACEAE on the upper lip and 3 on the lower, these often reversed by the inversion of the flower. Stamens 5, free from the corolla, or lower part of the filament sometimes adnate ; fila- celled. Style a stigtna 1—2-lobed ; ovules numerous, Fr uit a 1 “s-celled capsule. Seeds numerous, smooth or variously marked with transverse and longitudinal striae. Filaments ca or united distally; anthers distinct; flowers never blue. Flowers solitary on hig Dr pedicels borne on branches of erect stems. F; oe e stems. 2: shella, Tree ousted: “snthers che’ into a tube, two shorter, the opening of the tube thus oblique; ioe vari- Sly c Flowers pedicellate; le obconic to hemispheric or narrowly clavate. Corolla including lobes 2-10 m a a ng or lacking; more or less succulent annuals. Plants inenegyed aquatics; orl nao cewek less than 3 mm, long. 3. Howellia. corolla wich ia mm g Corolla 3. 41 mm. long; cap te-oblong, 1.5—2 mm. wide. Le. 4. Legen Corolla abo le ob , 3-4 6. peers Corolla including ie eee 50. mm. long; coarse perennials heleds L. dortmanna and L. kalmit). 7. Lobelia Le Dessningle. Flowers sessile; capsule very narrowly linear, simulating a flower-pedicel. 1. NEMACLADUS Nutt. Trans. Amer. Phil. Soc. IT. 8: 254. 1843. diffusely branched annuals with numerous slender branches. Leaves a basal, the cauline leaves minute, sessile, subtending the dichotomous branches. Inflor cence racemose. Flowers minute on proportionately long, usually capillary ‘pedicels, Feds bracteate at base. Calyx partly or entirely free, the lobes triangular to ovate. Corolla bilabi- ate, the upper lip 2-lobed or -parted, the lower 3-lobed or-parted. Filaments monadelphous at the tip; stigma capitate, 2-lobed. Capsule 2-celled, 2-valv t cx - many-seeded. [Name from two Greek words meaning thread and branch, in reference to the very slender : oad branches. ] A genus of about 12 species, natives of Oregon and California, and also adjacent Arizona, Nevada, and Mexico. Type species: Nemacladus ramosissimus Nutt. Calyx a from the a ~ ol to the base; capsule two to three times as long as the Pipe al corolla-tube eeding the Af ongiflor us. Calyx-tube saeais to aay ovary ops not peck exceeding the calyx; corolla-tube not Wick de the calyx- inhive (slightly exceeding in N. secundiflorus) Seeds ek alata’ or zigzag, ded pation ridges, separating the well- or 3 ae Corolla about equaling the lobes, not exceeding the calyx-lobes exc N. secundiflorus. Axes of the shia a to a less oppnaek gee — Em * distinctly yt than broad ature capsule ol ted a x; mature pedicels widely socesits witha tg ale (this less ret a nN ‘secundsfiorus ia leaves Lane c-ovate to elliptic, entire or irregularly crenate-dentate; plants (the la ones) toa branched; cismontane feet “California <_< eserts. 2. N. sigmoideus. Basal leaves oblong-lanceolate, oh apes crenate-dentate to saksledacihee Inner South Coast Ranges and Greenhorn Rang age yan reged included withi ns yx; distich 4. N. secundiflorus. rset 8 t ee ostly included within the calyx; distich . N. gracil ' Mature | Seer sharply pointed and a, firm at apex; mature peel ascending Rs upward only toward the N. pinnatifidu —— - the ra epoemes mreient and also the i seeds ote globose. : N. ramosissimus. Cor ube thus extremely shor Anthers 0.5-0.8 m: Branches diffuse poe spreading; pedicels mostly pj corolla-lobes ciliate; aioiave and Colorado Deserts and adjacent Nevada and Arizo rubes Branches at peice more or less stiffly ascending; corollas not, sitenes western pl 6. Nemacladus ramosissimus Nutt. Nuttall’s Nemacladus. Fig. 5103. Nemacladus ramosissimus Nutt. Trans. Amer. Phil. Soc. II. 8: 254, 1843. Nemacladus tenuissimus Greene, Bull. Calif. Acad. 1: 198. 1885. Plants 5-20 cm. high, intricately branched, the “tir and branches ascending and, including he mo : or eae secund inflorescence, straight and not at all zigzag, glabrous throughout | or pubescent below, greenish pestis brownish purple t toward the hae Basal leaves as gan ; oblanceolate, narrowed to nged etinie at base, toothed o r pinna atifid, glabrous or pode sely pilose especially on the pom cae rd the base; diets very finely capillary, Screial ing nearly horizontally but curved upward near the apex, the flowers erect; calyx campanulate in broadly conic in pole corolla 1.5-2.5 mm. long; filament-tu be 1. m. rai smooth a glabrous, usually curved pex; capsule 1.6-2.5 mm. high, broad and —— at apex; seeds about 0.5 mm. in almost nei with 10 rows ae usually 6 rounded p flower, ~ + Dry, sandy or gravelly soil, Sonoran Zones; Tassajara Hot Springs, wr 8 eine California, to northern Lowis a also inland in western Inyo and in San Bernardino Counties. Type locality: “San Diego.” May—Jul 7. Nemacladus rubéscens Greene. Desert Nemacladus. Fig. 5104. sion ane acca oeed — Bull. Calif. Acad. 1: 197. 1885. Parish, Bull. S. Calif. Acad. 2: 28. 1903 Nemacladus ibis var. rubescens Munz, Amer, Journ. Bot. 11: 245. 1924. Plants 5-20 cm. hi gh, repeatedly forked and becoming diffuse and bushy in well-developed plants, glabrous or thes ms sparsely paggdonse paw, usually shiny gray-green. 1 leaves 8-15 mm. long, usually tion ee Bg unded at apex, mostly entire, glabrous or sparsely her c ly esp ag th aoa pedice mm. long, illary, slightly a wage straight or somewhat curved upward at the tip; calyx broadly rounded in ower, mostly hemispheric in fruit; corolla with a short tube and sp ding lobes, yellow, with p rown margins on the | obe: , these = least Pose upper ciliate on the i “3 lament- tube st traight or slightly curved above, glabrous and smooth, conspicuously exserted ; seeds broadly ellipsoid, with 8-10 undulate or zigzag ice tseidiecl gee and poorly defined pits Sitwesn the ridges Sev sandy or gravelly soils, Sonoran Zones; from In a and Kern Counties, California, southwestern Nevada, and adj — Arizona south through “ Meieve and Colorado Deserts to northern Lower California. Type eee California. se ake ae forked above with ascending ake: glabrous or nearly so, dull ss Pi, brownish o rplish. Leaves 5-18 long, . broa Det broad ad ole, ar abrous or the hs and very sparsely puberule - eos els 10-15 mm. foie strongly ascending, glabro alf-i blunt a seeds elli hid, itl 10-12 1 Oe longitudinal dees sepawated ay: Pats Eapeesoed lonituinal ca Y heratige arked by about 30 fine cross-ridges. Usuall ne outcrops, Upper Sonoran and Arid Transition Zones; Inner Coast Ranges, Lake and Napa "Charticn. California, Type locality: Allen’s Springs, Lake County. May—July. 11. Nemacladus rigidus Curran. Rigid Nemacladus. Fig. 5108. Nemacladus rigidus Curran, Bull. Calif. Acad. 1: 154. 1885. Plants 4-9 cm. high, the stems saints or is from the dogs at gg glabrous or sparsely pubescent, the be cher often decumbent. s 7-9 mm. lon ey wide, elliptic, narrowed to a broad base, flowering from nie oe mee ai stems, pee as a patios 8-11 mm. long, stiff and comparatively short, soresdiie: horizontally or widely seeing: a 2- uy long, = from the base of pedicel, elliptic, usually about one-half as wide as our ; corolla 1-1. m. long, © elite or purplish, with short pide page's -1.6 mm. long, the anthers with mienihe reflexed appease capsule long, 2-2.5 mm. broad, about cage a hypanthium, — me t base; seeds prnebery with *8 10 pears ee ridges, with narrow between the r a be — d ae iti Z - theastern egon adjacent Cali- son Sand isis, oper Sogoran Tomesier evade. a He liciger Grade near Virginia City, Storey County. May-July. BLUEBELL FAMILY 87 12. Nemacladus capillaris rieagh Common Nemacladus. Fig. 5109, Nemacladus capillaris sagen Bull. Calif, pom, i: 885. Nemacladus rigidus var. capillaris Munz, Am mele oi ‘ies 11: 244. 1924, Plants 7-18 cm. high, the stems i eats or minutely steag g brownish or purplish, the branches usually several times forked, rather stiffly ascending, the axis of the raceme more or less zigzag especially in fruit. Basal leaves 5-15 m m. long, coat a ate, narrowed abruptly at base to a short petiole, entire or ahariane’ crenate, usually a Aha pedic els 8-12 mm. long, spreading or ascending, capillary a pe ya steal aight, the flower then usually horizontal and no strongly upturned, glabrous; brac long, perecely eS corolla about 1 mm. long, white, lobed almost to the base; lament ge 0.8-1.2 mm. long; calyx-tube turbinate in flower and in fruit, the lobes 0.6-1.2 mm. long, ay i or slaccane’ riangular capsule 1. mm. long, rounded at apex, about half- inferior ; seeds few, broadly ellipsoid, wit th 8-10 narrow ‘longitudinal ridges oo with rows o wey tas Dry velly slopes usually of igneous formations, mainly Arid Transition Zone; southern Oregon in Jackson Baty ‘Klamath Counties Pion cone in the Marth roe Ranges and Sierra Nevada to the no cy yay parts of the Mojave Desert, California. Type locality: “Mohave Desert.’’ Collected by Curran. June—Aug. 2. PARISHELLA A. Gray, Bot. Gaz. 7: 94. 1882. Low, diffuse, annual plants with the leaves and flowers in subcapitate tufts at the base of the Slant ¥ nd at the ends of the branches. Calyx-tube campanulate, shorter than the spatulate lobes. Corolla rotate, almost equally 5-parted. Stamens with minute a inferior part of ovary and capsule into 2 cells, the lid of the capsule without septum at maturity. Seeds pitted. [Name in honor of S. B. and W. F. Parish, botanical saliciocs in southern California. ] f the Mojave Desert. 5106. Nemacladus — 5108. Nemacladus rigidus 5107. Nemacladus m 5109. Nemacladus capillaris 88 CAMPANULACEAE 1. Parishella californica A. Gray. Parishella. Fig. 5110. Parishella californica A. Gray, Bot. Gaz. 7: 94. 1882. Low rs with 1 to several stems from the base, diffusely branched, glabrous or nearly so and easily urple. Leaves in a basal rosette or rarely the floral brac ink foliaceous, oblanc ets ais obtuse eg dene and about 5 mm. or less chee gradu 4 tapering into a margined petiole, 8-12 mm. long, or sometimes ovate and beige narrowed t erin petiole ; ies oir Raia few- to Bie cred’ corolla white, 34.5 m igh, tubular- pais e, the lobes arta about 1 m wide; seeds ellipsoid to oblong, vitted with 8-10 longitudinal 3 oa of 10-12 ach. Gravelly slopes, deserts and plains, Sonoran Zones; eastern San Luis Obis rie and in the west- ern and. central hag of i oe Mojave Desert, California. Type locality: Rabbit Rorities, tones Desert, San 3. HOWELLIA A. Gray, Proc. Amer. Acad. 15: 43. 1879. Delicate, immersed, aquatic, annual plants with flaccid stems and narrow leaves. ZA ee ers both apetalous and petal-bearing. Calyx-tube adnate throughout - ve ovary, the 5 farther than the other 2; anthers unequal, 2 of them smaller than the ee . Ovary 1-celled, with 2 parietal placentae ; ovules few. Fruit a capsule, irregularly dehiscent by the, tupture of the very thin lateral walls; seeds large, smooth. [Name in honor of Thomas Howell, a pioneer botanist of northwest America. ] A monotypic genus of northwestern United States. 1. Howellia aquatilis A. Gray. Howellia. Fig. 5111. Howellia net Gray, Proc. Amer. Acad. 15: 43. 1879. Stems flaccid and somewhat fistulous, sparingly branched, ‘7 ag Reiss Leaves narrowly linear- chaise. 2-5 cm. long, entire or w ith a few slender teeth; arlie cleistogamous in the axils of ordinary leaves, the later on panckes with more or her ke shsk ‘ene some with and others without petals; corolla 2-2.7 mm. long, whitish or pale lavender, about equa aling the 1 cal linear, acute, unequal calyx-lobes ; ovary wholly inferior, in rit with a depr essed-conic summit capped by the persistent base of the style, narrowly clavate, 8-10 mm. long; seeds 1-5 maturing, smooth and shining, 2-4 mm. long, cy lindri ric, rounded at one end, pointed at the other In stagnant ponds, mainly Humid Transition Zone; west-central Washington and norther ore to the Willamette” Valley, Oregon. Type locality: ‘Seuvies Teland. near the mouth 4 Willamette River, Multnomah ounty, Oregon. May-Aug. 4. LEGENERE McVaugh, N. Amer. Fl. 32A: 13. 1943. Annual herbs growing in moist or wet ground or the base of the plants often immersed, rooting at the nodes. Flowers loosely aie Heer both with and without corollas. Corollas of the petaliferous flowers cleft dorsa y. Stamens with the filaments and also the anthers connate; 2 of the anthers shorter than dee ce er 3. Fruit a 1-celled capsule with parietal placentae, detects at the apex. [Name an Tatas of E. L. Greene, the discoverer of the only known species. ] A monotypic genus of central California. 1. Legenere limésa (Greene) McVaugh. Legenere. Fig. 5112. Sat ia — reteset: igen 2: 81. 1890. Legenere limosa McVau: N. Amer. Fl. 32A: 13. 1943. a. so, er se or som ate decumbent and rooting at the nodes, simple or on ten with few to many lateral branchlets, the brs plant green, smooth and glabrous. Leaves the cauline entire and sessile, the lower acute and early deciduous; floral racts foliaceous, clliptic, caeeae obtuse at apex, somewhat rounded and sessile at base, 2-3 mm. wide, 6-12 mm. long; corollas 3.54 mm. long, white, the iohes about 2 mm. long, the 2 lower ones distinctly narrower than the 3 upper lobes ; capsule wholly inferior, dehiscent at apex by short, thin-walled valves ; ating se in fruit subulate to broadly deltoid, usually 4 in apetalous flowers, 5 in petaloid ones; seeds smooth and entte : . 4 pe eo oran Zones; wer acramento an an Jo Valleys, al also | ale - "Coal Mine | Fn ge yf Meng re California. Type locality: lower Sacramento Valley ‘Imira, Solano County, according to type specimens in Greene Herbarium. y. 5. DOWNINGIA Torr. Pacif. R. Rep. 4°: 116. 1857. Nomen conservandum. soft-stemmed, annual, spring-flowering herbs, erect or decumbent and some- n r eaves narrowly linear and entire or with a few narrow teeth or sometimes pinnatifid, usually not functioning at flowering time, those of the gph beans thicker and firmer, narrowly linear to broadly lanceolate or oblong. Flowers 5-merou perfect, inverted, solitary in axils of upper leaves or foliaceous bracts, Paes i Sine te, BLUEBELL FAMILY 89 stalk-like, inferior ovary resembling a stout pedicel. Calyx-lobes 5, usually with the three upper lobes longer than the other two, normally entire and linear. Corolla a varying to blue, pink, or white, usually with a white or yellow blotch at the base of the lower lip; tube entire, the limb abruptly bilabiate: the 2 lobes of the upper lip usually narrower an an th i e lower lip; 7 var usually shining, smooth or with faint striae. [Name in honor of A. J. Downing, an American horticulturalist. western North American genus of about 12 species. Type species, Clintonia elegans Dougl. —— ‘ena — or only ee incurved. lip corolla conspicuously reflexed, forming a sharp angle with the tube; corolla 7-20 mm. long, aaa much tg the ge lobes. bepais a ene white-bearded within the lower side; tip of anther-tube Pus fies and ned, bristle-like processes; seat he 2-celled. D. bicornuta, eidons: ihe ‘Slabeots within (sparsely pilose in D, ornatissima); tip of anther- jee vi or without processes, these never bristle-like and intertwined. as and capsule 2-celled, the cies attached to the longitudinal septum Ser sinuses of corolla deeply cut, the lower lip thus appearing hinged, the dorsal sinuses qually cut; corolla-tube sparsely pilose within on the lower side. 3. D. ornatissim Lateral sinuses of the corolla scarcely or not at all deeply cut; ysiog tube ei ace Filam nen shorter than corolla-tube, the anther-tube t - t least —— included ; th eg not tapering distally, the apex rounded o chee use; corolla-lobes not in Seeds Ato or lightly marked longitudinally with straight line Lobes of the og sn corolla- te marginally ciliate (under ca rey hosted corolla- lip with broad purple blotch. color, Lobes of the upper corolla-lip eciliate; lower corolla-lip with 3 purple ae . D. bella. Seeds appearing twisted, marked obliquely with fine lines on the lone € sxis 9 of he seed. spidata, spe p tube longer than the corolla-tube, the anther-tube thus as anther-tube in- lined to taper distally to the acute or slightly rounded apex; corolla-lobes in one ace, 7. D. pulchella. Ovary and capsule 1-celled, the ovules attached to ovary-wall. Dessal anthers of anther-tube pilose at apex; aia: dull, lightly marked =“ netiotioa). lines. Dorsal anthers of Bp tube omg or sparsely pilose; seeds smooth ee sogenhat intute, without longitudinal ma 11. D. yin Lower lip of corolla ascending; corolla 2-7 mm ov ding th b merely equaling them. Corolla 2-4 mm. long; ah appearing twisted, “lgkty marked with ae lines. 6. D. pusi Corolla 4-7 mm. long; seeds without mar Anther-tube aieeete Sacter weeks usually at ae uehea - Ere wasgering cad: Ova ary , the o attached to the Pp 8. D. insignis. Ovary and caeuale celled: the sia attached to the ovary-wall. 12. D. elegans. $111 5111. Howellia aquatilis 5112. Legenere limosa 5110. Parishella californica 90 CAMPANULACEAE 1. Downingia concolor Greene. Maroon-spotted Downingia. Fig. 5113. Downingia concolor Greene, Bull. Calif. Acad. 2: 153. 1886. owningta tricolor Greene, i 2: 79. 1890 con G ket Bolelia tricolor Greene, loc. c Bolelia concolor var. epreeret tsicki Fl. W. Mid. Calif. 481. 1901. Stems branched from the base, few to many or s sig See Bi ne cm. high, glabrous ex- cept for 4 the calyx-tube which i acktack minutely puberu Leaves linear, —2 mm. wide, —20 mm. heey pi tcl loosely few-flowered, ‘the ii Suan elliptic to ovate ; calyx-lobes linear-elliptic or oblanceolate, ascending , g, rly equ we often shorter ; coroll m fe brous except for the two upper lobes, these ciliate-scabrou on the margins near th | orice deep blue, the lateral sinuses more d t than lip purple except for the spot at base, this bearing 2 ridges or low, nipple-like pr ocesses ; corolla- t 3-5 mm. long, narrowly fun nelfor rm; anther-tube slightly exserted or sometimes inclu ded in the —— os glabrous or SHS pubescent on the a minutely hae at gan the 2 shorter anthers da fort horn-like, apical processes; capsule n rrowly iform, 1-2 mm. broad and 3-5 cm. long; seeds not twisted, the markings cael to tone. axis 0 oist depressions, Upper Sonoran Zone; valleys of the North ae Ranges of California from Lake Cedars ¢ to “Monterey pawns | and also in oe Sacramento Valley; collected at Deer Creek, Tulare County, by Cong- don. Type locality: wheat fiel r Suisun, Solano County. March—June. Downingia competion var. otoasd Me Va ugh, Mem. Torrey Club 191: 1941. Mature capsule mostly 12-25 mm. long, early dehiscent, the valves separated by lines of delicate ie tissue. Cuyamaca Lake, San Diego County, Californ rnia. Known only from the type locality. 2. Downingia bélla Hoover. Hoover’s Downingia. Fig. 5114. Downingia bella Hoover, Leaflets West. Bot. 2: 2. 1937. eae < to 17 cm. high, glabrous throughout or the rage ad slightly acc Fe: ae what fistulous. he eaves 5-12 mm. long, 1-1.5 mm. wide; orescence loo feacks aoe to elliptic, acute or obtuse, 7-18 mm. long, pin wide; pf -lobes aeiiae or rotately spreading, 3-6 mm. long; corolla 10-12 mm. long, enelt blue, tatceal sinuses more deeply cleft than upper sinuses and cut below the su urface of lower lip ; lobes of the upper lip lanceo- r sli lle wi e tube; lower lip with central white with a yellow t d2y ridges at base, these alternating wit 3 small purple — or the central pur t a : — - g, f lform; anther-tube ess scaberulous, the 2 lower anthers bea short, alee processes at apex; re doo linear, 3.5—5 cm. long ; pote wile ntl markin Vernal pools on alkaline plains, Sonoran Zones; Sacramento and San Joaquin ad from Colusa Soir to Talase ‘County, California. Type lscaliee.: near San Joaquin River catleea: of Modesto, Stanislaus County. April—Aug 3. Downingia ornatissima Greene. Solano Downingia. Fig. 5115. Downingia ornatissima Greene, Pittonia 2: 80, 1890. Stems simple to few-branched abové, or in more vigorous plants branched below — above, 6-20 cm. ga glabrous throughout sel ee pene See ig aciny pace ts scabrous. Lea so 16 wi “ ate shorter, Sie 9) mm. ae ‘oe or acemeaat asks Reig acti 8-13 mm. long, the tube 2-3 mm. long, narrow ne y rded in on e latera and upper sinuses about equally cleft, fon! = lip ava hinged, the margin of the sinus usually turned backward into a horn-like projection; lo of the bee corolla-lip ge eee spreading, i i i ue to t g want into a pk wer rc ve, dee whitish with a white patch bearing 2 yellow or yellowish green seo the corolla-lobes rounded or tru e base of the li i ‘ also with a sharp, horn-like process, glabrous or pubescent dorsally; capsule linear or narrowly eye 2.5-6.5 cm. long, 1-1.5 mm. thick, the pve lip tough; seeds with faa longitudinal lin ditches and is the beds of oe pools, Sonoran Zones; Butte County to Merced and _— Coun- Abo ties, California. Type locality: lower Sacramento Valley near Elmira, Solano County. April-May owningia 5 ornatiasima var. eximia tote) ee. Mem. Torrey Club 194: 24. 1941. (Downingia mirabilis J. Leaflets West. Bot. 1: 221. PES bo mirabilis var. eximia Hoover, op. cit. 2: 6. be gcd Two upper sarelig tenis minutely pubescent within ns "the apex, divergent but not curled into » Ie Moi depressions, San Joaquin Valley, California: Type locality: vicinity of Orange Cove, Fresno Cou 4. ibaa bicorniita A. Gray. Double-horned Downingia. Fig. 5116. Downingia bicornuta A. Gray, Syn. Fl. N. Amer. ed 2. 2: 395. 1886. Bolelia bicornuta esi patents 2: 127. 1890. Downingia sikota Applegate, Contr. Dudley Herb. 1: 97. pl. 6. 1929. Plant 6-30 c aeig sane times more in partially submerged plants, glabrous. Leaves linear- lanceolate, 0. 52 ¢ cm. long; sf varioon usually few; calyx-lobes widely spreading, linear to narrowly BLUEBELL FAMILY 91 without, Dey bearded within on the lower side, the lateral sinuses more de ply cleft than the upper he surface a the oa r lip, the corolla-tube broadly funnelform; upper tol, tia, ire long, ovate ai lanceolate, erect or so at divergent, the tips usually turned backward; lower lip folded or reflexed, about 10 mm. long, much wider than long, the 3 lobes truncate or rounded, mucro aie, pu urplish blue with white patch marked with yellow or greenish yellow and deep purple at base of lip, this bearing 2 prominent, nipple-like protrusions centrally and 2 less Tigo ie ones iat erally ; stamineal column included in the corolla-tu anther-tube pabernient. apically with 2 apical bristles arising from the 2 shorter anthers reflexed and pine bp iam ovary 2-celled; seeds faintly marked ater st lly t depressions, usually in adobe soil, Upper Sonoran and Ssiediton. Zones; southeastern Or gon and y et ae Tdaho and Nevada southward in California to Sierra and Merced Counties. Type locality: C ..co, Butte i, California. May—Aug. ngia bico otdsta var. picta Hoover, Leaflets West. Bot. 2: 4. gf Flowers nel ago slightly meer tan in the typical species; corolla- pos with a brownish xt $a spot on its upper side; lower lip strongly _— ve, with the 2 upper lobes lags or pale blue often tipped with darker Bio not go Rccary but directed ward sgh other so their tips cross, strongly reflexed and ha a to the tube; horn- sod enter r-processes often a oe than the secre themselves. Low = Sere Upper Sonoran Zone; San Joaquin Valley, California. Type locality: r Le Grand, Merced Coun 5. Downingia cuspidata Greene. Cuspidate Downingia. Fig. 5117. Bolelia cuspidata Greene, Erythea 3: 101. Downingia cuspidata Greene ex Jepson, Fl. W. Mid. Calif. ed. 2. 403. 1911. Pepntngts puphele's var. arcana Jepson, agoabait 1: 100. 1922. Munz & Jtn. Bul hae we Club a 300. 1924. Diéabuingie pallida sthanaé, Leaflets West. Bot. 2:1. 19 Stems simple or branched, 6-25 cm. high, see throughout or the calyx-tube sparsely and 5113. Downingia concolor 5115. 5114. Downingia bella 5116. eae tae bicornuta 92 CAMPANULACEAE minutely scabrous. Leaves narrowly linear to linear, 0.2-2 mm. wide, 3-13 mm. long; inflorescence loosely 1-20-flowered ; bracts linear to sere elliptic, 4-12 mm. long, mm. wide; calyx-lobes wae > oblanceolate, ascending, 3-10 mm. long; corolla 7-15 mm. long, glabrous, bright or pale blue o lavender, rarely white, the tube ease phe: a to almost cylindric, the dorsal and y evr’ sinuses about equally cleft, the lateral slightly cut below the surface of lower ip; upper lobes ovate, acute, slightly divergent or recurved and overlapping in age; lower li plane or ae aring 1 d wi so, with a cent more or c t yellow ridges at base, the lobes broadly ovate or oblong, rounded to retuse at apex ; upper lip diteer than the lower, purple-veined ; filament-tube glabrous, included ; Antheks tube gl abrous or sparsely ten dorsally, = minutely white-tufted at apex, the 3 shorter also React a short, horn-like ocess ; calyx-tu nape or slig tly sl in fruit, 2-4 cm. long in fruit; seeds shining, the fine face running cuencty o the ae axis of t pie Clay soils of pervs vernal pools and yn an Zones; Humboldt and Shasta Counties south t San Diego County, California. Type locality: ane Sar west of Yountville, Napa Co., Calif.” April ie eth 6. Seoul pusilla (G. Don) Torr. South American Downingia. Fig. 5118. Lobelia pusilla Poepp. ex Cham. Linnaea 8: 217. 1833. (Nomen nudum) Clintonia Crea a rate on, Gen. Hist. Pl. 3: 718. 1834. Downingia pusilla Torr. Bot. Wilkes Exp. 17: 375. 1874. hater humilis vail Pittonia 2: 226. ingta humilis Greene, Leaflets Bot. Obs. ‘2: 45. 1910. om glabrous throughout or the caly ii minutely scabrous, 2-12 cm. high. Leaves 0.5-1 mm mm. long; Rgoicciegeees 7 tow red; floral bracts elliptic lanceolate, 2-8 mm. long; calyx-lobes 3-8 mm. long, or appre , surpassing the corolla; Shee 2.5-4 mm. long, glabrous, whit “the lower lip blue-tipped, the upper and 1 about equally = = arrowly campanulate, —2 ong, pper lobes deltoid “ate eolate, about erect ; lower dl spreading, with white by centrally bearing a yellow pa ea rates the 3 thhies deltoid, g:-sta -tube 2 a he hite-apiculate, the . lon tamen-t m. 2 shorter with a blunt, ci process and a few br inten t apex; ovary 2-celled ; capsule linear, 2-3 cm. long an subulate : seeds marked with oblique oes, appearing twisted. Moist flats, ae See sparingly introduced or possibly native in Napa, pace: Stanislaus, and Merced Counties, California; also Chile and Argentina. Type locality: Chile. Nepid os 7. Downingia pulchélla (Lindl.) Torr. Flat-faced Downingia. Fig. 5119. oeeenen Lgaee ete Bot. be 22: pl. 1909, 1836. he Pacif. R. Rep. 44: 116. 1857. Bolelia eaihate Geteie: steer 2: 126. 1890. Plants up to rh = high, glabrous ae cling or the calyx-tube sometimes a a Leaves pie eal linear, 1-2 mm. wide, 4-12 mm. long; inflorescences few-flowered up to 15-20- flowered ; flor af bracts varyin from Ripe glace | to ovate, usually elliptic ; cabeetibes aealls > ‘ately pond 3-10 mm. long ; corolla 8-13 mm. long, glabrous, deep bright blue varying to p or white ; tube purple, 23 mm. lon ng, pode — oo + Uppes eg oe 2-cleft, the lobes elliptic to oblanceolate, spreading, 8 m lon ; lower lip eading in same plane as upper, the lateral sinuses somewhat de Piss Sh pote ith a cen les al white area bearing 2 yellow spots that extend down the pa Fase folds at io of lip, 3 dark purple spots afer rating with the bogs aga rel lobes oblong, acute, or mucronate; filament- tube longer than corolla-tu 4.5 m. long; tube attenuate, the anthers gla brous oe minutely ol ate-tufted apically, the 2 shocks | Pay with slender, horn-like processes ; capsule linear, 3-7 cm. long, narrowly subulate or fusiform, the walls tough in texture; seeds shining, without fine lin na Adobe or alkaline soil of vernal pools and ditches, Sonoran Zones; California from Lassen and Colusa Counties south to Merced and Monterey Counties. Type locality: California. Collected by Douglas. April—June. 8. Downingia insignis Greene. Cupped Downingia. Fig. 5120. owningia insignis Greene, Pittonia 2: 80. 1890. Bolelia insignis Greene, op. cit. 126. Stems very slender, somewhat zigzag, 1-3 dm. high, glabrous throughout - the calyx-tube sometimes somewhat scabrous. ves narrowly linear, 5-15 mm. long, 1-2 mm. wide; inflores- cence 4-20 cm. long, usually about 5-flowere d but varying to (6 scwent: floral | baiag elliptic to fe) a rin; patch, this sometimes reduced to 3 purple spots, the lip concave, not reflexed, and usually shorter than the upper lip; filament-tube glabrous; anther-tube usually strongly incurved at right angles a ees tube ; anthers and connectives whitish in color, granular-roughened throughout, the 2 shorter white-tufted at pr mi ouaey 2-celled ; ideas A 3-8 cm. long, the lateral walls tough and break- i gots = ie ak sted. al § Low fie! d depressions, am ; California in northern Sacramento Valley south to ce Cou ag cos Costa ¢ a County bg Segorng sen County and occurring Bee wie: Rem a I phase ag Vs mad he es BLUEBELL FAMILY 5117. Downingia cuspidata 5118. Downingia pusilla 5119. Downingia pulchella 5120. Downingia insignis 5121. Downingia laeta 5122. Downingia montana 94 CAMPANULACEAE 9. Downingia laéta (Greene) Greene. Great Basin Downingia. Fig. 5121. Bolelia laeta Greene, oii gee £2236. jo 3, Bolelia brachyantha Rydb. Mem. N.Y. Bot. Gard. 1: 483. 1900. Downingia laeta near Lenfleta Bot. = 2:45. 1910 ssn 2-3 dm. high, sometimes shorter, hated throughout. Leaves lanceolate to elliptic, 5 m. long; inflorescence rather loosely 1-10-flowered; floral a hie iptic or lanceolate to ovate, "7-20 mm. long; calyx-lobes elliptic, somewhat unequal, 3- m. long; oye light lie or purplish, 4-7 mm. long ; corolla- oe funnelform, 1-2 mm, long, vellow below on the lower side beneath the purple area ; upper corolla-lobes lanceolate or triangular, acute; le ip is central area white or yellow with the ache band of purple at base femeass redu ced t r 3 purple aoe, the lip spreading, its lobes aban a cute, 2-3. 5 mm. long; anther-tube little or ie at all ncurved ; anthers glabrous or ciliate on thes margins, the O shorter ones with a tuft of white hairs ; ovary fe. celled ; capsule 2-4 cm. long, the walls thin but tough ; cute slightly or not at all appearing tw poe picasa Uppe x oe an and Arid Transition Zones; southern Oregon east of the Cascade Moun- rtheastern Cali rnia eastward - Nevada, sabligeetern Wyoming, and southern Saskatchewan. he ocality: Humboldt Welle Elko County, Nevada. June—Aug 10. Downingia montana Greene. Sierra Downingia. Fig. 5122. Downingia montana Gr i ge Pittonia 2: 104. 1890. Bolelia montana Greene, op. Downingia saa: var. eee Soaks Madrono 1: 102. 1922. Plants up to 15 cm. high, _omacigit throughout se calyx-lobes sometimes minutely scabrous. Leaves ied to subulate, 1-5 mm. wide; inflorescence 1-10-flowered; floral bracts subulate to narrowly linear, similar to the leaves Sin Nightly eneaiee calyx-lobes linear- salehiate, acute, some- ‘scabr n th gin isti e — m. long, dark blue or violet, glabrous, the lateral sinuses cut below the surface of the lower lip ; corolla-ti os xs 5-5 mm. long, na arrowly funnelform and somewhat gibbous, exceeding the calyx; upper olla-lobes erect, narrowly {ingot : lower lip of corolla with central white patch, ‘the aay bluish purple with 2 proce purple folds at the angle at mouth o te new go > all of the ude, & nu anthers bear at apex, the 2 shorter ones tufted with short bristles and a horn- ie process ; capsule subulate, 2-4 cm. long; seeds not shining, marked fotleitudinally with fine line Moist mountain meadows, Arid Transition _ Canadian Zones; northern Colifipmia i Siskiyou County, a south in the Sierra Nevada to Tuolumne Coun Type locality: Lake Eleano , Tuolumne County. July— 11. Downingia yina Applegate. Cascade Downingia. Fig. 5123. Downingia yina Applegate, Contr. Dudley Herb. 1: 97. pl. 5, fig. 2. 1929. Plants small and delicate, ang gna or the calyx- ue slightly scabrous, the ae simple and 1-flowered or branched and few to several- flowered, 2.5-7 c igh. Leaves 10-30 m a long, linear-lanceolate, sale e; calyx ape shorter than the curate he: corolla 8-10 mm. lon rk blue with a yellow throat, usually t 7 mm. broad, the lateral i nee deeply cleft pre the upper, the tu narrowly tometer: upper lip with ay linear-lanceolate, acute, erect, usually approximate, about 4 mm. long an nd 1 mm. wi e; lower lip plane, Sting bluish pS the central yellowish area cerca by white except for the 2" yellow ridges at base of lip, the , ‘ 1 and 2 vary slender, 8-1 . long, 1-celled; en-column erect, included; anther-tube somewhat pubescent do rsally ; valves of the pels mit by hy aline cae Citeniink the length of a capsule and visible as iaeteased fi lin mature capsule Fates near the middle ; seed dullish or somewhat shining, without fongitudinal lines Moist r boggy meadows, Canadian Zone; Cascade Mountains of le Oregon to Siskiyou County, Cali- fornia. Type locality: Four Mile Lake, —— 7 Oregon. July-A ia yina var. major McVau te oe ee : 24, 1942. © OD eycinese ae etn tag med M “ ; 4 : invisible before splitting, with no bapressed pie on oo surface; seeds ie and without bor —, and i ype 12. Downingia élegans —— ere and meadows, Arid Transition and Canadian Zones; eastern Oregon to southwestern Mon- tana, northern Wyoming, and norther n Nevada. Fone locality: ‘‘Camas Plain, Flathead River.” Collected by Wyeth, probably in eastern Idaho. pote July. a “ereues scipcaatencys (DC.) Nutt. Narrow-leaved Mule-ears. Fig. 5139. th . Fl. Bor. Amer. 1: 313. 1834. Not Pursh, 1814. Alarconia ? pines oe ers = Ange 1836. Helanthons stage te tad op. c Tr rans. pee Phil. Soc. IT. 7: 352. 1840. Wy ethia r bus ta. ae W yethia agape a var. cpeainsti Jepson, Man. FI. Pl. Calif. 1080. 1925. Perennial herb from a eo or a vertical caudex, pilose or hirsute and somewhat resinous ; stems ascending, 1.5-3 dm. high, usually aan: iaigice — to densely pilose or sed abo petioles 2-11 cm. long (often margined above), the blades 1 rebavk ‘atconee te to "oblong long, 1-8 cm. wide, acuminate, long-cuneate at base, papery, entire to serrate, sparsely. ‘to densely hirsute or hirsutulous on both sides with spreading to subappressed hairs ; em-leaves 1 edu petioled or sessile, not clasping ; heads 1(-4), about 5-9 cm. wide ; involu road, 1. . high, i be s ceola ovate or obovate, acu subglabrous dorsally ; rays 10-18, 1.5-3.5 cm. long ; achenes puberulous at apex, about 8 mm. long; pappus a oe -denticulate = ‘lacerate crown of 's squa pape! 2 mm. long or less and 1-4 slender peep lou -_— =. ong or less, sometimes reduced to teeth. yam d oj — en Upper Sonoran = Transition — southern bp hoy tsa (Klickitat County) — gietat— om ope 9 ei te County, California, and sparingly in the Sierr: evada. ae ocality: Monterey, Califor Bs 3 th cted tad ouglas. April—Aug. Wye ustifolia var. folidsa (Congdon) H. Hall, eee felt. Pu ab. Bot. 4: 207. ger (W yethia tases ‘he (poatecliny bis sf 186. Heed Pageroar 4-5 “Mt dm. high; upper leaves ovate to lanceolat ith clasping heads 1-4, usually smaller phyllaries mostly la nee-linear or lemeaclate: Mainly in northern | California oe acent Oregon wns a 3 in rie Sierra erm fom Shasta oe to Fresno County and in the North Coast Ranges in in a and Mendocino Counties, California. Type | : “wooded slopes of the ee ranging om 3,000 to 8,000 ft.”’ . Not pelhent differentiated from W. angustifolia var. angustifolia because of existing intermediate forms but recognizable by the characters outlined above. 11. Wyethia longicatlis A. Gray. Humboldt Wyethia. Fig. 5140. Wyethia longicaulis A. Gray, Proc. Amer. Acad. 19: 4. 1883. Perennial herb, essentially g proline throughout except sometimes on involucre, resinous and yet stem es essentially s simple, ascending, about 4 dm. high. Lower leaves oblanceolate, about 20 cm. long (including the ceeroisis margined cots 3 cm. wide, obtuse, long-cuneate at base, SUNFLOWER FAMILY 105 pe sea except toward base, et ge ciliolate, rather firm, feather-veined; stem-leaves few, milar or lanceolate, 7-15 cm. long, the upper entire, all narrowed at base; heads solitary or few linesiedunclal medium-sized ; involucre broadly campanulate, 2 cm. high, subequal, the phyllaries e ss achenes about 7 mm. long, glabrous; pappus a eee cro Ridges and prairies, Transition Zones; North Coast Ranges of California in eastern Humboldt adjacent Trinity, and nort eastern Mendocino Counties. Type locality: prairies ~y ark Humboldt County. May-July. 12. Wyethia amplexicatlis Nutt. Northern Mule-ears. Fig. 5141. Espeletia amplexicau lis Nutt. Journ. Acad. Phila. 7: 38. 1834 Wyethia amplexicaulis Nutt. Trans. Amer. Phil. ee _ tf 352, 1840. Silp ? laeve Hook. Lond. Journ. Bot. 6: 244. W yethia lanceolata Howell, FI. yee Amer. 341. se W yethia emplecicouhe subsp. major Piper, Proc. ys Pie Wash. 27: 98, 1914. W yethia subsp. pices ti iver, loc. Stout eae glabrous and more or ae balsamic-resinous throughout, often vernicose, from a pepru: fusiform caudex, the stems ascending, 3-8 dm. high, essentially simple, leafy. Basal sha short petioles, these usually margined above, the blades oblong, elliptic- ‘oblong or obovat 5137. Wyethia mollis 5139. Wyethia angustifolia 5138. Wyethia helianthoides 5140. Wyethia longicaulis 106 COMPOSITAE 15-50 cm. long, acute or acuminate, cuneate to rounded at base, entire or serrate, feather-veined, rather firm; stem-leaves several, gradually reduced above, elliptic, oblong, or oblanceolate to ovate, the lower Pattee ted the upper r sessile and clasping ; heads 1-8, terminal ie axillary, peduncled, 4-10 cm. wide; involucre h .8-4 cm. high, subequal or somewhat graduate, the phyl- laries numerous or rather few pace ee to oblong or occasionally obovate, acuminate to obtuse mainly herbaceous, glabrous, not ciliate; rays ii bright yellow, 2-5 cm. long; receptacular aga ts Lede naa glabro s, 7-10 m m. lon ee a Bence crown of squamellae 2 mm. . lon 4 ao 23 = wn ~ tee Wet or dry i ag places, occasionally in woods, Arid Transition Zone; British Columbia, eastern Washington, and ine a png egon to Montana, Wyoming, Colorado, Utah, and Nevada. Type locality: about Fl athead River, Montana. y—July. Smooth Dwarf en Wyethia S eet Piper, Proc . Soc Wash 27: 98. 1914. Intermediate between W. amplexicaulis and W. heltanthoides, with both of ae i grows, and ‘evidently a hybrid between the two species. Type locality: lue Siecctgms, "Union Co ounty, Orego 2. BALSAMORHIZA Hook. ex Nutt. Trans. Amer. Phil. Soc. IT. 7: 349. 1840. Perennial herbs, scapose, subscapose, or sparingly leafy-stemmed, from a thick, ver- tical, fusiform or subcylindric, terebrinthine root and simple or branched caudex. Leaves tufted and basal, the reduced stem-leaves aticcihe or subopposite and those of the isk-flow short tube and cylindric or cylindric-campanulate throat, 5- toothed. Style- branches slen- der, hispid or hispidulous sg slender, subulate, terminal appendages. Anthers 5; aa or shortly sagittate. Achen s oblong, epappose, glabrose or pubescent, the ray-achene sag “% tri peor. fee of the disk quadrangular, usually with ei nerves. [Name from the Greek, referring to the resinous roots. A genus of abou 2 natives of southwestern Canada and northwestern United States. Type species, Heliopsis a to So hina tax ad Balsamorhiza ‘hybridize freely (Ownbey & Weber, Amer. Journ. Bot. 30: 179-187. 1943) and certain of the med entities have been proven to be pe hybrid origin: Balsamorhiza icshintwackn (Hook.) Nutt, Trans. ‘Amer. Phil. Soc. IT. 7: 349. 1840 re maga ‘4 Lig stax pen gee Hook. Fl. Bor. Amer. 1: 310. 1833), derived from = a and B. hookeri or B. sa 1, and Halsasmochine vi bénseri St. John, Fl. S. E. Was 1937, derived from B. Soloties pe y Plants with woody root and multicipital caudex; basal leaf-blades broadly deltoid or sagittate, entire or coarsely crenate or toothed, the Stem: leaves borne near (pe inflorescence. Plants densely toment t least when young. 1. B. sagittata. Plants not canescent-tomentose or ae Achenes glabrous; rays deciduous. eltoidea. B.d ‘Mian pubescent (except in var. intermedia) ; rays mostly persistent and becoming pa Panes 3. yana. Plants ah a thickened woody taproot (up to 3 cm. in diameter) and a typically simple tee Sector: basal ont Ag cet or oblong in outline, poconapd ce Pai on aa divided (except sometimes in B. serra id we rosea), the stem-leaves if present borne e base. Leaves pinnately or bipinnately divided. not no teeably pecsuns OE SEY and subcoriaceous. Outer phyllaries 2. ~ 3.5 cm. long, much surpassing the inner phyllaries and the disk, often toothed or incised distall crolepis. Outer ea hag 1-1. -5(2) cm. long, about the length of the inner piv thackes cae equaling or shorter entire. Phyllaries linear- So or lanceolat . B. hooker e, broadly efacibinks or abruptly attenuate Eat a caudate - ‘Raeves gieeniihi: eclaidey bipinnately divided or bipinnatifid with aed gay eather divisions. we . or silvery, not at all hirsute, pinnatifid, the divisions entire or shallowly incised. s 3-6 cm. long; pubescer:v< densely lanate with long, soft, tangled hai 7. Bit meana. Rays 2-3.5 cm. long; p > ?P d ly Pr d short-sericeous. - 5. B. macrolepis platylepis. Leaves crenate or sharply serrate, if at all incised not parted to the midrib, reticulate-veined and subcoriaceous. es sharply serrate; rays yellow, the width of ray about one-half the length B. serrata. Leaves crenate; rays becoming roseate, the width of ray about four-fifths o length. rosea. 1. Balsamorhiza sagittata (Pursh) Nutt. Arrow-leaved Balsamroot. Fig. 5142. Buphthalmum sagittatum Pursh, Fl. Amer. Sept. 2: 564. pat Espeletia helianthoides Nutt. Sanne: Acad. Phila. 7: 39. ceconepttes # es Nutt. Trans. Amer. Phil. Soc. 7: ue "1840. Per 2-6.5 dm. high from a deep-seated, woody taproot and amit me caudex clothed with bcc ENE the sparsely ihe Whit lomentec stems bearing 1-3 heads. Basal leaves 2-5 dm. ree up to 1.5 dm. wide, the petioles exceeding to nearly equaling * thie blades, these —— sagittate or -hastate at rath triangular or triangula titel entire, densely white-canescent o SUNFLOWER FAMILY 107 cinereous-tomentulose we gland-dotted, usually becoming more or less glabrate in age on the upper surface; stem-leaves much reduced and bract-like, spatulate or lanceolate: pik steep 1-2 cm. wide. densely eon Satie: tomentose ; outer phyllaries about 2.5 cm. long, lanceolate-acumi- nate, wy A Sc the inner ; rays 2-4 cm. long, 13-21, withering, soon deciduous ; aha 7-8 mm. long, glabro vi n cna, and — s, mostly in the Arid Transition Zone; southern British Columbia and Idaho south es eastern Washington and Oregon to northeastern California and adjacent Nevada and the eastern slope he Sierra Nevada to the Tahoe camine: Saskatchewan and Montana south to Utah and yp rng hed east to the: agg han of South Dakota. Type locality: “On dry barren hills, in the Rocky-mountains.” Collected by 2. Balsamorhiza yng Nutt. Deltoid Balsamroot. Fig. 5143. Balsamorhiza deltoidea Nutt. Trans. Amer 1. Soc. a 7: 351. 1840, Balsamorhiza glabrescens Benth. PI. Hartw. ae Perennials with the habit of rg sagittata, “ stems a dm. high, eatin 1-4 heads, sparingly leafy, green, glandular, pilose or pilosu lose. a leaves 1-5 dm. long, up to 2 dm . broad, the n age, subcoria m-leav uced, lance oe rs linear-lanceolate ; involucres the ere op one the la acl pas or as short-pilose; outer phyllaries 3-6 cm inne lavicealen - oblong i er papery in age; ace —8 mm. long, . Open rolli x pes, Transition Zones; British yg roe south on the west side nd o Dhagpe ge Mou tains i "Callioentn. ue to tae foc ony omg in the C oast Ranges and the wester of the Toes Nevada to Kern County. Type locality: Willamette Valley , Oregon. March-July. fase oe y al By than the disk; rays 2-5 cm. long, about 12-20, n > iw Sons no »e “ Bg pst) n 2 =] je 3 co bo om =} =) o +t ie] s a. 3 s 5141. Wyethia amplexicaulis 108 COMPOSITAE 3. Balsamorhiza careyana A. Gray. Carey’s Balsamroot. Fig. 5144. Balsamorhiza careyana A. ea Mem. Amer. Acad. II. 4: 81. 1849. Perennials 2-6 dm. high from a deep-seated woody taproot ae re pean caudex, clothed with Ficaas wa bases, the a leaved stems bearing 3 or more closely racemose h ds. B sal leaves 1.5—5 dm. long, o 1.5 dm. wide, the petioles nearly wie, ing to pained Rot the blades, these cordate a ae . ge ad s sinus, mostly triangular-hastate, entire or nearly so, subcoria- ceous an = Mie Ss ie glandular, and hispi dulous on bot mg ae ga hs ves few, mee poopie: becoming bracteate and sessile above; involucres 0.8-2.5 c _ br oad; r phyllarie 182 long, “ear Oa ee exceeding the inner, more or ‘Pe hirsute to elabrate Be ~3 cm. lo: z, , persisting and becoming somewhat papery ; achene s 6-7 m , hairy. Plai A eile ds, Arid Transition Zone; central Washington in Grant and L on unties south to ieuielenwa Umatilla and pore ow Counties, Orego on. Type locality: given as (dennis but collected by Spalding probably west of the Clearwater River. March-July. Balsamorhiza careyana var. eat Cronquist, Vasc. Pl. Pacif. Northw. 5: 101. 1955. Differing from fil baci yovie shyla in hav the leaves often crenate, the Paine ai Lionas of ee ep acenense enlarged, the outer ore ease, pa the — glabrous. — of the Cascad ntains from the British Sarco ge Aa border ponte to Deschutes and Wheeler Counties, Oregon. Type locality: Bee nse ga Gam Wheeler ounty, eg 4. Balsamorhiza hodkeri (Hook.) Nutt. Hooker’s Balsamroot. Fig. 5145. pape iopsis ? balsamorhiza Hook. Fl. Bor. Amer. 1: 310. 1833. a anaehiee Rente ert ies Att. Trans. Amer. Phil. Soc. II. 7: 349. 1840. Heller, Cat. N. Amer. PL 7. 1898 Perennials from thick wo oody taproot with black see bark and int unbranched caudex, stems 1-3 dm. high, scapose or with a pair of nearly basal stem-leaves, sericeo agit some longer hairs near the liteey heads. Basal leaves 10-20 cm. ola lanceolate in omliae ¢ he blade much longer than the petioles, pinnately parted to the midrib, the ices cleft into Brevi sor lobes, thinly sericeous, mgt es nsely s ase the smaller leaves, the petioles often pilose as well; involucres a .5-2.5 cm e; phyllaries 1-1.5 cm. lon g, 2- 4 mm. wide, lanceolate, attenuate at apex, the outer : phvtialies 4 sae excee ae the disk, sandiete. lanate on the margins and densely so toward the rag heen thin x. oe on the surface and appearing prea = ig rays 1.5-2.5 cm. long, about 10-16; s 4-5 mm. long, glabro ais wed Arid ees i pees ebateen Klickitat County, Washington, to Clark County. Type local- ity: Fort Vancouver, Clark County. ae I-June Intergrading with B. hookeri var. apecepeele in cme — ven oe oe Uatsauarhie. hookeri var. lanata Sharp, Ann Gar 935. Habit as in B. hookeri var. hookeri, the plants low, densely aneie throtighout a Peg ioe yay as cs hipinnately divided; i sane as in B. hookeri var. hookeri but with a longer, more dense pubescence. Know hills near Yreka, Siskiyou —— California, the ty pe locality: Bal orhiza hookeri var. Tagocéphala (Sharp) me Vasc. Pl. Pacif. Northw. 5: 103. F (Hdeaworkiaa becate var. " legocephala Sharp, Ann, Mo. . Gard. 22: 139, 1935.) Habit as in B. hookeri var. hookeri; leaves pinnately divided, the lobes cleft into a — pier broad lobes or teeth, green, hirsute or create: lous with appressed hairs pn often paneer: phyllaries seriate, the outer rarely surpassing the inner, lanceolate = aca lanceolate, more or less ee mentose es pecially at the base of the outer phyllaries. Rocky soil and ebrush, east of the Cascade Moi Washington. Type locality: Ellensburg, ee kh A highly bate taxon wlgintenbia pid with B. hookeri var. hookeri at its southern limit and w other taxa in Balsa morhiza where the ranges overlap. Balsamorhiza ‘hokest: var. neglécta Mo. Bet by cam aye Vasc. Pl. Pacif. Northw. 5: 113. 1955. (Balsa- morhiza eee var. so de esol Ann ard. omg 139. 1935.) Habit as in B. hookeri var. hookert, out 2 lowly to deeply cleft “with plant eae iy ane erasien green, with an appressed-hirsute pubescence; phyl- laries more or less se the outer seldom surpassing the inner series, typieny lanceolate to linear- lence; more or less iscastaneniace, densely so on the margins. Rocky soil, central Utah westward through northern and central Nevada to northeastern California. Ty ae ality: Truckee Pass, Wiehe County, Nevada, any aberrant forms probably of hybrid origin are foun ashoe County _. in adjacent California which may have ovate-lanceolate or ovate-attenuate, gg se glandular Slevtecrkes from which the villous-tomentose pubescence = bc lacking. Balsamorhiza hookeri var, lanata, B. macrolepis, and B. ieectolents var. platylepis occur in this os Fairey eee macrolépis Sharp. California Balsamroot. Fig. 5146. Bal. Sharp, Ann. Mo. Bot. Gard. 22: 132. 1935. Perennials ies a coarse woody root and short caudex, the several aos 2-6 dm. high, scapose or with a pair of reduced pinnate foontd ae the base, appressed-pu t below, appressed or spreading above. Basal leaves 2-4.5 dm 10 cm. wide, blade vata bees than the petioles, lanceolate in outline, pinnate, the div ate “Tanceoate, ‘usually deeply lobed, the lobes rounded or acute at apex, very thinly sericeous or appressed-t tomento: rs 2-3 cm. — abere ssed short-sericeous and aa ely glandular ; outer phyllaries 2-3.5 cm. much surpassing the j inner and the disk, oblong or prt Ba lanceolate, usually incised or toothed at on Bs. the inner lanceolate; rays about 10-15 and 2.5-3.5 c m. long; achenes 5-7 mm. long, £ » of 3¢ c & yas 28 Qa. e. 5 x -_ 3 = 2. =e = o wn Fields and r hill slopes, Upper Sonoran Zone; Butte County, California, south to Mariposa County and me the BD Coast anges from Sonoma County south ‘to Santa Clara County. Type locality: Clear Creek, Butte pent ora orhiza macrolepis var. platylepis (Sharp) Ferris. Contr. Dudley Herb. 1958. (Balsamorhiza plage Sh “Sharp, aan Mo. = . Ga + shes _— 1935.) Plants 1-3.5 pe high; aa enceihe usually more dense than B. macrol. va lepis: pi innate divisions of the leaves shallowly to not at all lobed, ed rb iecediy acute cade cheterics Shorter’ than to equaling ac disk, me outer and the second series ar or ovate, irate 3 r abruptly attenuate and often spreading, sericeous o ee sg and sometimes velutinous particularly in ‘nk o the Siskiyou Mountains area, Siskiyou aa oe phine County, Oregon, to Siskiyou ‘ounty, Califacnia. where it grows on serpentine; also Modoc Comunity Saiccnis. south to Nevada Co Cony and adjacent Washoe County, Nevada. Type locality: Marmol Station, Washoe County. Silvery SUNFLOWER FAMILY 109 6. Balsamorhiza hirstta Nutt. Hairy Balsamroot. Fig. 5147. Balsamorhiza hirsuta Nutt. Trans. Amer. Phil. Soc, II. 7: 349. 1 840, Balsamorhiza hookeri var. hirsuta A. Nels. in Coult. “& Nels. New Man. Bot. Rocky Mts. 546. 1909. rah = ith “ habit of B. hookeri, the stems 2-5 dm. hi gh, s scapose or with a pair of b : cH z =} =] F) ag ch Pe oo a a c x 3 FX) oF o zy y Z ° i} w 3 ry) ry) a oO o wn 28 wn eS - ° g3 rz) 4 o mo cm. broad, more or less hirsute or hispid and pilose- ciliate: outer phyllaries 1-2 cm. long, n surpassing the inner, ovate o ui aeaenkaas with an a ruptly FP sie hear pee te tip; Pec et ni cm. long, 10-18; achheks 3-6 mm. lo ae: glabro Rocky soil, a br oiengeriewg gd ; local in rn Oregon. oi locality: “Dry plains east of Walla-Walla, near. oy Blue "Mountain and in the Grand oa prairie.”” Probably collected at the second locality as the species is not known cae Walla Walla. Collected by Nuttall. May-July 7. Balsamorhiza incana Nutt. Woolly Balsamroot. Fig. 5148, Bal hiza t Nutt. Trans. Amer. Phil. Soc. II. 7: 350. 1840. Balsamorhiza hookeri var. incana A. Gray, Syn. Fl. N. Amer, 12: 266. 1884. Balsamorhiza floccosa Rydb. Bull. Torrey Club 27: 629. 1900. Perennials with the habit of B. hookeri, the stems 2-7 dm. high, scapose except for a pair of 5148. Balsamorhiza incana 5149. Balsamorhiza serrata 110 COMPOSITAE reduced pinnatifid leaves near the base, loosely lanate. Basal leaves 1-4.5 dm. long, 3-10 cm. wide, lanceolate to oblong in outline, the blade much exceedin ng the petiole, pinnate, the lanceolate or ov ns ly an ate division large, arsely toothed or more deeply incised, pubescence loose lanate and usually densely so; involucres m . . wide, densely ca eS with loosely spreading hairs; phyllaries 1.5-2 cm. lon outer not aang po the inner, ovate to ovate- lanceolate with an abruptly Secaaheg eee poe rays 3-5 cm. long, 13 or more; achenes 3-6 mm. long, glabrous. Meadows and grassy Arid Transition and Canadian Zones; southeastern. Montana west to south- eastern Washington” fae iow 0 central Oregon east of the Cascade Mountains. Type locality: “Rocky Moun- tains.” Collected by Nuttall. May-July. 8. Balsamorhiza serrata Nels. & Macbr. Serrated Balsamroot. Fig. 5149. Balsamorhiza serrata Nels. & Macbr. Bot. Gaz. 56: 479. 1913. Perennial from a coarse woody taproot and short simple caudex, the several stems 1-3 dm high, icone or with 2 reduced leaves near the base, sparsely pilose and usually reddish. Bas al leaves 6. cm. long, 2-10 cm. wide, the petioles shorter than the blades, lanceolate to ovate- lanceolate, the margin shallowly serrate to incised or divided often on the same Looe subtruncate or eae at base, green, coriaceous, conspicuously reticulate-veined, scabrous; involucres about 1.5-2.5 cm. i i outer not surpassing the inner; rays 2-4 cm. long, mostly 10-16; achenes 6-7 mm. long, glabrous. Dry rocky soil in open forest or sagebrush, Arid Transition Zone; dae stern Washington south to Lake — Harney Counties, Oregon, and adjacent Modoc apg , California; also northwestern Nevada. Type ieeality: r Rock Creek, Morrow County, Ovened: May-— e. s Balsamorhiza rosea Nels. & ea Rosy Balsamroot. Fig. 5150. Bal. i Nels. & Macbr. Bot. Gaz. 56: 478. 19 Baers hookeri var. rosea Sharp, Ann. Mo. Bot. “sel #2: 130. 1935; Perennial with the habit of B. serrata, the stems 0.6-3 dm nee — oe or with a pair of rofeet bract-like leaves near the base, avaicee Basal lea s 3-20 cm. long, 1-10 mm wide, the blade ‘longer than the petiole, oblong to deltoid, crenate an lobed attics to ‘the midrib ith ate divisio e or less re glandular ; psi ag oe about 2 cm. broad, densely white-pilose ; phyllaries about 1 cm. long, linear- lanceolate, acute, the outer not s surpassing the inner; rays 1-2.5 cm. long, up to 1 cm. wide, pale yellow(?) rt roseate in age; achenes 5-6 mm. long, strigose Rocky soil, Arid Transition Zone; known from three localities, Yakima eee Walla Walla si and Spokane County, Washington. Type locality : Rattlesnake Mountains, Yakima County. April-May 3. VIGUIERA H.B.K. Nov. Gen. & Sp. 4: 224. 1820. Herbs or shrubs, more or less pubescent. Leaves usually opposite at least below, ee void, pubescent or glabrous; pappus (disk-achenes) of 2 usually Se etae awns and several shorter, gd or basally united, persistent squamellae, or ah wanting. [Name in honor of L. G. A. Viguier of Montpelier, author of a work on Papav Ag t 145 species, native of North and South America. Besides the following, six others occur from entens to ly and Texas, ad another species occurs in Georgia. Trpe species, Viguiera helianthoides H.B.K. (= Viguiera dentata var. helianthoides (H.B.K.) Blake). Pappus present; achenes pubescent; ionves hash or triangular. W 4 iia dhe ee } Leaves narrowly triangular, | toothed and usually "1. Vz laciniata. Leaves erent or deltoid-ovate, serrate or entire; plant not resino Leaves densely and closely cay rather prominently iirdsia beneath, canescany pilose to subtomentose "iaore. . reticulata. Leaves somewhat veiny but not closely nor prominently reticulate odes green and tuberculate- hispidulous above. 3. V. deltoidea parishit. Pappus absent; achenes glabrous; leaves linear or lance-linear. 4. V. multiflora nevadensis. 1. Viguiera laciniata A. Gray. Laciniate Viguiera. Fig. 5151. Viguiera laciniata A. Gray in Torr. Bot. Mex. Bound. 89. 1859. Shrubby, much-branched, leafy, resinous, scabrous-hispidulous a up to 1.3 m. high, the branches sler hispidulous with mostly incurved hairs. Leaves alternate, ras RE petioled, sig with axillary fascicles, the blades narrowly triangular, 2-3.5 cm. lo eo 7-20 mm. wi ide, acu SUNFLOWER FAMILY 111 acuminate, at base shortly cuneate, laciniate-toothed and hastate-lobed toward base or rarely sub- entire firm, glandular-punctate, tr riplinerved and veiny, harshly and rather. sparsely So ea the aire especially of upper face w ith tuberculate bases ; heads 3B i in a terminal cyme or cymos panicle, 1.7-3 cm. wide; involucre 3. -seriate, slightly graduate, 5-7 mm. high, the phyllarie with ovate indurate base and a abrupt, shorter to longer, acuminate, lin aA to triangular, loo rbace tip, a epee ck rays 8-12, 0. cm. long; achenes sparsely st rigose, 3 m ; pa ay a 2 broad, somew wha t deciduous, paleaceous awns and a - short, persistent, incenate squamellae. Dry Bes and me Sonoran Zones; southw n San Diego County, California, and northern aor California. Type loc ubey: "Rancho Gensiclin. on of ean Diese, Califonse September, 1855; Schott.” April—-Nov 2. Viguiera reticulata S. Wats. Leather-leaved Viguiera. Fig. 5152. Viguiera reticulata S. Wats. Amer. Nat. 7: 301. 1873. Suffrute arent branching perennial 0.6-1.3 m ce the stem and branches slender, net appreg’sed- or incurved-pilose, ery shove Lea ed opposite or alternate, short-petioled, the blades cate: oF o pep -ovate, 2.5-8 cm. long,:1. a7 . wide, acute, broadly roun mabe gs - sub- cordate at base, entire or rarely serrate, 0 eiatinareed and strongly reticulate especially beneath, piesa Sct sia caktcdaest tose above with mostly antrorse hairs, less conspicuously so beneath with looser, more spreading ts pi gland-dotted ; he heads 1.8-2.5 cm. wide, in small, te at n a 3-seriate, graduate, about 5 mm. high, the phyllaries with ovate, indurate, ribbed body and usually shorter, narrower, oblong or erg usually appressed, herbaceous tip, cinereous- Stearn especially toward margin; rays about 8-15, about mm. long; achenes rather sely ap- ecg td oes 3 mm. long; pa Soush awns 2, Ea sity 2 mm. long or ie, about twice re Ieng a fimbriate Giadiaeting, Dry rocky slopes and canyons, Sonoran Zon desert ranges of Inyo County, California. Type locality: Telescope Peak, Panamint Mountains. Ageii- Sept. “Death Valley Golden-eye. 5150. Balsamor! 5152. Viguiera reticulata 5151. Viguiera eiertgg 5153. Viguiera deltoidea liz COMPOSITAE 3. Viguiera deltoidea var. parishii (Greene) Vasey & Rose. Parish’s Viguiera. Fig. 5153 Viguiera parishii Greene, Bull. Torrey Club 9: 15. 1882. Viguiera deltoidea var. parishii Vasey & Rose, Contr. U.S. Nat. Herb. 1:72. 1890. _ Branching shrub 0.8 m. high or less with gray-barked stems, the branches slender, densely cinereous-hispidulous, often sparsely hispid. Leaves chiefly ss a jus iene the blades deltoid-ovate, 1.5-3.5 cm. long, 0.8-2.5 cm. wide, acute or obtu unded to subcordate at base, serrate to subentire, triplinerved and y veiny, tuberculate- hispidlous a eland-dotted; pon 1-6 toward tips of stems and branches, mostly long-peduncled, 1.5-5 ¢ wide ; e 5-9 high, 2-3-seriate, somewhat gr raduate, t e ohetacies with ovate to rescey -ovate, pea te, ribbed body and usu ally longer, ong to ahetate herbaceous tip, ie aeicn Maal strig illose ; rays 0.8-1 3 mm. lon ea 8-10, .5 cm. long; achenes appressed- - appus 52) cok 3 mm. long, sada oe ie cher four times as long as the Fal ag feb ise. Mesa rocky canyons, Sonoran Zones; southern A ada south through the Mojave, pei Colorado Deserts, California, te fee Lower California, east to Arizona and se yell —— also coastal San Diego County, California. Type locality: San Luis Rey, San Diego County. April—Sep 4. Viguiera multiflora var. nevadénsis (A. Nels.) Blake. Nevada Viguiera. Fig. 5154 A. Nels. Bot. Gaz. 37: ae sie Comnotomi aiid: Reage Bull. Torrey Club 3 nevadensis Blake, Contr. pa Savy No. 54: 110. 1918. Slender, several stemmed oon from a short rootstock with stems 25-90 cm. high, s simple or branched, strigose or strigi TM eeiok opposite below or nearly throug fe on dea 2 - m. i ender- peduncled, 1.5-5 cm. wide; involucre 2-seriate, subequa 7 mm. high, the phyllaries linear- lanceolate to sig ion ate, herbaceous, acuminate or acute, densely “strigose and strigillose; rays 10-14, 8-25 mm. long; achenes 1.8-3 mm. long, glabrous, epap Mesas and canyons, Upper aa n Zone; southern Utah and a Wi . south to Inyo County, sad fornia (Inyo, Argus, and Panamint ixeaes). aie San ion ey County in the Kingston Range and Clark Mou tain and eastward laa a Type locality : Meadow Val Wash, Lincoln County, Noved da. May-— Sept Nevada Show Vigalels ciliate, Robins & Greenm.) gg Contr. Gray Herb. No 54: 918. (Gymnolomia longifolia bins. & Gree oc. Bost. Soc. Nat. Hist. 29: 92. 1899. ) An annual wae hispid. ciliate leaves = the phyl- — merely h sspid- Shed or also s pzvely hispid dorsally. This native of southwestern Uni sa States and oda: cent Mexico fog een found as an adventive plant at Santa Monica aseted 6-20 cm. in leneth nts to tal P stems usually hispid, is 1 ~ few hea . A. cusickii, oe over 1.5 m. tall; stems usually glabro head Phyllaries iieiianalt dilated near “ 3—4 mm. broad, long-attenuate and ae ap . H, californicus. Phyllaries not conspicuously dilated near base, 2-3 mm. broad, short-attenuate, pr FF gneir 8. H. nuttallit. 1, Helianthus annuus L. Common Sunflower. Fig. 5155. Helianthus annuus L. Sp. Pl. 904. 1753. ual, 0.5-4 m. tall, branched pk ig ian stems usually rough- psn Type locality: “America.” July—Oct. 5184. Bidens laevis 5185. Bidens frondosa 130 COMPOSITAE 4. Bidens vulgata ecg Western Stick-tight. Fig. 5186. Bidens vulgata Greene, Pittonia 4: 72. (July) 1 Bidens frondosa var. puberula Wiegand, Bull. akan pine 26: 408. (Aug.) 1899. Bidens vulgata var. puberula Greene, Pittonia 4: 250. 19 Annual, similar to B. frondosa but coarser aA taller ; terminal division of leaves often short- stiped; heads larger, the gs in fruit 1.3-2.5 cm thick; outer phyllaries 10-16, densely hirsute- —— oe merely pubescent or Sera vet up to 5.5 cm. long; corolla yellow; achenes broadly siceehies -cuneate, 6-12 mm. long, 2. mm. wide, usually merely antrorse-ciliolate, on Pieie ti pubescent or nearly ae aati yellowish or olive- brown ; awns 2, retrorse-hispid, 5 m ong or les Moist inser and roadsides; — southward to North Carolina and Missouri and westward to Was ee, Oregon, and in California in the | ramento Valley, prow it apparently is introduced; also adventive in Euro Type locality: not given. June—Oct. 5. Bidens pilésa L. Beggars-ticks. Fig. 5187. Bidens pilosa L. Sp. Pl. 832. 1753. Bidens californica DC. Prod. 5: 599. 1836. ender prs branched chiefly above, up to 1.3 m. high, stem striate- -angled, sparsely epi or hi rsute to arly glabrous, with long internodes. Leaves slender-petioled, pinnately 3-5-p the divisions greed -oblong to ovate or rhombic-ovate, crena iearregte to ‘incised, ee Fa long-stiped ; head -flowered, solitary or few and irregularly cymo e em a branches, 6-8 mm. thick in flower, 1-2 ick in fruit; outer phyllaries 6-8, linear or spatulate, about 3.5 mm. long, ciliate especially above, shorter than the inner; disk yellow; rays none or few, sh small, yellowish white; acher inear or linear-fusiform, 5-13 mm. long, act a A or sub- ua le outer much shorter than the inner; awns 2-4, erect ~ Duties about 2 mm. ay naked below auoedlng “hispid for about the upper half of their gue abt praia eed in waste places and cultivated ground; na of the American tropics, of — many fom segre Betray occurring - eee ia from Monterey ae psa Counties south into Arizona and Mex mye focality: “America.” March—Oct 6. Bidens béckii Torr. Water Marigold. Fig. 5188. Bidens beckii Torr. ex Spreng. Neue Entdeck. 2: 135. 1821. Megelodente beckis Greene, Fittonia 4: 271. 1901 272. Megalodonta beckti var. oregone 5 Rhee ff, Bot. Gaz. 97: 609. 1936. Megalodonta beckii var. henditsens Sherff, Amer. Journ. Bot. 25: 589. 1938, Aquatic to glabrous throughout, the stems floating o r rarely emersed, simple or few- branched above o 2 m. long or more. Submersed leaves sheers ile, 1.5-4 cm. long, 3-parted “ ‘ t oO n lee oe cad about 8, golden yellow, oblong-oval, 1-1:5 cm. long; ae linear, 1 cm. long, obcom ek. “quadrangular, striate-ribbed, gla brous, saliowish or gree rown; awns 3-6, slender-subulat —2.3 cm. long, divergen t, rigid, somewhat unequal, silonte inside, naked below w, a nse sions no “hispidulous all around at tip for about one-fourth thei r length. akes, rivers, and mar Quebec to New Jersey west to Manitoba, Minnesota, and Mis Pacific State in Washington =~ g een where perhaps introduced by aquatic birds. Type ality. prt "Scher ork, pt. garden cosmos (Cosmos bipin s Cay. Te: ¥s:10: 7p) < plant with penree divided into linear, more or bag flfform he ee and with whites pcg or crimson-rayed fl s, may be found occasionally as an adven- 13. COREOPSIS L. Sp. Pl. 907. 1753. Herbs p= 8 usually glabrous or nearly so. Leaves opposite or alternate, entire to dissect . Hes mall to medium, radiate, usually yellow, the rays sometimes with. purple- brown spot at % e disk sometimes purple-brown. Involucre hemispheric, double; a ‘Bhalaries Tscrate usually few (6-10), herbaceous, usually much shorter than the ree at base; i ore nu s, 2-seriate, equal, membranous, free, sea brown + grt es Receptacle flattish, the receptacular bracts membranous or s scarious, flat. Ray-flowers neutral or pistillate, their corollas ligulate, usually wedge- shaped, often scpihed or 3-5-lobed. Disk-flowers fertile, their corollas tubular, with sider tube and 5-toothed limb. Anthers bidentate or subentire at base. Styl e-branches tipped with a conical appendage or subtruncate. Achenes obcompressed, usually oblong or oval, tet or ohare usually ae Pappus se 2 teeth or awns ‘ never gages: barbed), te oricia peces SUNFLOWER FAMILY 131 A genus of over 100 species, mainly American but found also in Africa and the Hawaiian Islands; numerous others occur in the United States. Type species, Corvopsis lanceolata I.. Perennials; comparatively tall and s Leaves deeply bi- to ipinnatel pre d; an enaete fertile. Heads politany og fen eduncles usually 2 ee long 1. C. maritima. Heads d, the coal ante 1- dm. 2. C. gigantea. Leaves entire, sometimes iviied, a few lanceolate ee ray- pis ers sterile. 9. C. lanceolata. Annuals; slender and usually low Rays yellow — (or w vhite tipped) ; disk yellow; ray-flowers fertile. Disk- ay-achenes dissimilar; pappus of disk-achenes of 2 conspicuous awns. uter geri sea linear. nostental as anthesis; pappus-paleae of_disk- foment 2-3 mm. long; Monterey County, Califor1 outh and east to the Mojave Deser 3. C. bigelovit. Rays Ayre? pie a at arcs pappus-paleae + disk. flowers 1 mm, wd bn Bi: ‘i ee an Outer ueshaice” bedadly ovate or deltoid; pappus-paleae of disk-flowers mostly 4-5 mm. oi 5. C. calliopsidea. Disk- and ray-achenes alike; bene a small white c ves entire or with 1-2 linear pinnae, the Neal ee portion not broadened; annulus of disk-corollas earde pene, pubescent on both faces with clavellate or capitate hairs, corky wings eran ich thickened; plants of the deserts and cismontane southern California lifor cali ores . = both faces or nearly so, the corky wings thin -_ sn ae of he gs ast Ranges from Mount Hamilton Range to San Rafael Rang — Leaves pinnate -ar enna ng (sometimes entire in depauperate specimens); annulus of disk- pear glabrous or nearly so. 8. C. stillmanii. Rays purple-brown at base; disk purple-brown; ray-flowers sterile, Achenes wingless. 10. oria. Achenes winged. it: C. st ato le 1. Coreopsis maritima (Nutt.) Hook. f. Sea-dahlia. Fig. 5189. Tuckermannia maritima Nutt. Trans. Amer. Phil. Soc. II. 7: 363. 1841. Leptosyne maritima A. Gray, Proc. Amer. Acad. 7: 358. 1868. Coreopsis maritima Hook. f. Bot. Mag. 102: pl. 6241. 1876. sa gabrome throughout, the stems fleshy-herbaceous, spreading, 3-8 dm a thic eaves gee deltoid i in outline, u bi- to sriintely Wisiecte d into n . high, m. lose i 3 s 16-20, 24 cm. es; achenes narrowly inged, ane body of the arkaae and the narrow wn B d coastal bluffs and hillsides, Upper noran Zone; ego County, California, to northern “Califoraia’ | Collected by Nuttall. Son ; — Die Lower mina a and adjacent islands. Type locality: San Diego, San Diego County. March—Jun 5187. Bidens pilosa 5186. Bidens vulgata 132 COMPOSITAE 2. Coreopsis gigantéa (Kell.) H. “es Hall. Giant Coreopsis. Fig. 5190. rreciluentaag mgenren Kell. or Calif. Acad. 4: 198. H. M. Hall, Univ. Calif. Pub. Bot. 3: 142, 907. Tacheveauidia gigantea M. re Jones, Contr. West. Bot. No. 15: 74. 1929. tout, fleshy-woody, glabrous perennial 3-30 dm. high, the stem up to 1 dm. thick or ase branched ‘only above, the branches ae or neti ng, leafy only toward the ends. Leave alternate, deltoid or ovate in aa ce up to 3 dm. long, bi- to pat we goes dissected into ro seh or narrowly linear, fleshy, obtuse lobes 0. =i mm. Guu: heads 4-6 cm. wide, golden-yellow, mosely clustered toward tips of branches on Rare usually 1-1.5 dm me ie involucre essentially as in C. maritima; rays about 13, —2 cm. long; disk-corollas with puber ulous annulus tacular bracts 2-10 mm. long, linear, sok a Aa callous-thickened taiows falling tenets from the achenes ; achitties oval to oblong, 5-7 mm. long, glabrous, body of the achene and the narrow Wyias brown, epappose. Sand and sea cliffs, Upper Sonoran and ee umid Transition nes; coast of southern California from San Luis Ones, and Santa Barbara Counties to Los Angeles clase te adjacent agile (except San Clemente aa also Guadalupe Island, Lower California. ae locality: near Cuyler Harbor, San Miguel Island, Santa arbara County. Collected iy Harford. March—-May 3. Coreopsis bigelévii (A. Gray) H. M. Hall. Bigelow’s Coreopsis. Fig. 5191. Pugiopappus bigelovii A. Gray, ~— R. Rep. 4: 104. 1857. Leptosyne bigelovit A. Gray, Syn. Fl. N. Amer. 12: 300. 1884. Coreopsis bigelovii H. M. Hall, ak Calif. Pub. Bot. 3: 141. 1907. Erect, essentially glabrous, apres or subscapose annual, 3-60 cm. high, the scapose or scapi- form stems m mostly 5-10, naked or very rarely s sparsely leafy for ae -third their length. Leaves chiefly or entirely ina basal tuft, ov ate fe deltoid i in outline, 2-12 cm. long including sites 8 fleshy, e segments 2 cm. long or so in outer ski 57, linear or lance-linear, obtuse, 4— ; ; what sit , brown with narrow yellow margin; rays about 8- 0, obovate or wedge-obovate, ong; dis be orollas with puberulous annulus; receptacular bracts 4-10 mm. Jong, lance- olen or eve olate, heatine: falling with the disk-achenes; ray-achenes about 5 mm. long, ob- ovate-oblong, glabrous or = sentially so, epappose, often becoming corky-winged and cor rky-ribbed ; disk-achenes es long, about 6 mm. long, densely villous- ciliate, glabrous to Soaracle villous on the faces; pappus of 2 Shartesed. ficcatake. persistent paleae 2- 2s Ge ng. rt ikees — hillsides, Sonoran Zones; west of eed Sierra Nevada n Califor a from Fresno and Mon- oO Pients ‘of the Southern Sierra Nevada and the Tehachap bisa! da ns are rather toarke and leafy- stemmed but have tic linear outer phyllaries of C. b 4. Coreopsis hamilténii (Elmer) H. K. Sharsmith. Mount Hamilton Coreopsis. Fig. 5192. pt hamilt — Bot. Gaz. 41 906. Coreopsis hamilt H. K. Sharsmith, gee 214. 1938. Erect, ~— S ne dese annual 10-15 cm. high, scapose, sometimes branching at the base the scapes mostly 10-15. Leaves all basal or ey basal, 1-5 cm. long, spreading, bipinnate into short, linear, seve lobes ; eats 1-2 cm. broad, golden-yellow, the rays reflexed; outer phyllaries dly li ru inner . lor arrow en mm. long, meniscoidal, obovate, brown or mottled with brown, both the dy and marginal wings smooth a a acsoat wl disk-achenes 5-6 mm. ee narrowly obovate, both faces more or less Silas the ns = villous-ciliate ; pappus of 2 persistent paleae 1 mm. lo Mo Agarista calliopsidea DC. — 5: 569. 1836 Coreopsis calliopsidea A. G + Bot. Mie. Bound. 90. 1859. Leptosyne calliopsidea A. Gray. 8 Fl. N. Amer. 12: 300. 1884. : Erect, essentially ‘cobroas ana 1-4.5 dm. high, stenns 73 stout Anweiad sparsely leafy on . third to one-half Te r length and with less pronounced basal tufts of leaves. Leaves deltoid i $a te on upper t vate -) cm. long; disk- corollas with bearded a ge us; receptacular ae bot ong, broadly linear, obtuse, hyaline, falling separatély from t e aghenes; ray-achenes 5-6 mm. long, broadly oval, tan or bro winged, glabrous, noth a with a thick ae: coceriag: epappose ; disk-achenes 6-7 mm. long, SUNFLOWER FAMILY 5188. Bidens beckii 5191. Coreopsis bigelovii 5189. Coreopsis maritima 5192. Coreopsis hamiltonii 5190. Coreopsis gigantea 5193. C P P 133 134 COMPOSITAE arrowly cuneate-oblong, both faces more or less villous, the margins densely Me a pee ie ene ent, long, white cilia, the pappus of 2 linear- Suara ae persistent paleae, abou m. lon Moist hillsides and plains, Sonoran Zones; eastern slopes ner Coast Ranges, sac ae pois fornia, and the adjacent San Joaquin Valley south to the + sll Fallen beg g Barbara County, and east to the western edge of the Mojave Desert in Kern, San Bernardino, and Los Angeles Counties; also Sa augus, tn Angeles County. Type locality: California. Collected by Douglas. March-May 6. Coreopsis californica aa! H. K. Sharsmith. California Coreopsis. Fig. 5194. Leptosyne californica Nutt. Bos a a Phil. Soc. i 72.363, 1841, Leptosyne newberryt A. Gra roc, Am Sy 7: 358: 186 Coreopsis californica H. K. aa ith, Madre o 4: 217. "1938, oe douglasit of authors. Not Leptosyne islet DC. t, scapose, essentially ae annual usually 2-3.5 dm. high, the scapes about 5.0 5~ ‘et Leaves basal, erect, 2-15 cm. long, linear-filiform, entire or with a few short ee pee heads 1-3.5 cm. broad, golden-yellow; outer phyllaries 28, linear, the obtuse ex with a red callous tip, 3-8 (rarely more) mm. lor ng, te at the gibbous base; inner phyllaries 5-8, broadly so A to narrowly obovate, in age yellow and cae and constricted, the tufted apices spreadin ys 5-12, narrowly to bro dite obovate, br 16 m ong, often pale at the tip; disk- corde with a bear ded annulus, the ae ge often sterile ; ; Feceptacular bracts usually ; ep fone, line r, obtuse at apex t falling with the - nes ; achen 5-5 m. i both su corky ridge, tan or light brown, the wings of the same color, these strongly and irregularly corky- thickened, the ventral side of the wings with few to several reddish spots or blotches; pappus asm all Pla and sandy places. Sonoran Zones; southern png ranges m San arbara County to northern Lower California eastward hs ~ — Mojave and Colorado Deserts to ta Cooder. California, and western Ari- zona. Type locality: r San Diego, San Diego Pour. graced by Nuttall. March-May 7. Coreopsis fas Seats (DC.) H. M. Hall. Douglas’ Coreopsis. Fig. 5195. Lemerine dedtoiontt DC. Prod. 5: 53 is di | H. M. Hall, Univ. a Pub. Bot. .3.: ps! poe as to name only. Cvestepele stillmais var. jonesiit Sherff, Bot. Gaz. 97: 605. Scapose o or subscapose an siateal 5-25 cm. high or more, essentially glabrous throughout except for the short-pilose base of involucre, ay scapes mostl , slender, erect or as n cm. Ww e or wi m. he os He wide, golden-yellow ; outer phyllaries 4-7, linear, obtuse, 3-7 mm. lo ong, Dapillate at the gibbous base; inner phyllaries more numerous, obovate or 1, 6-10 m m. long, pe at the middle in age, the tufted apex spreading; rays about 8-10, cuneate, 0. 8-1. WE ; disk- ian with earded annulus; receptacular bracts 4-5 mm. long, linear, obtuse at le "hyaline, sige i rately ag seh eget achenes often meniscoidal, 2.5-5 m Bikey ong, the Mpe eer ie abro broad ged, the w tram i nish, slightly co ay ie Bearety thickened, unspotted on the inner tetas the body: of ‘i ‘actaaie shining: ark brown, glabrous or occasionally with a few inconspicuous papillae; pappus a small whitish cup. rocky slopes, Upper Sonoran Zone; Inner Coast Ranges, California, from Santa Clara County to Santa Dry Bitters: County and also the inner face of the southern Santa Lucia and the San Rafael Mountains. Type locality: California. Collected by Douglas. March-May (A re 8. Beka stillmanii (A. Gray) Blake. Stillman’s Coreopsis. Fig. 5196. Leptosyne stillmanii A. Gray in Durand, Journ. Acad, Phila. IT. 3: 91. Coreopsis pa Blake, Proc. Amer. Acad. 49: 342 Essentially glabrous he septa — s >the Brice and with stems occasionally branched aves 2- 10 cm. lon 3-toothed at the apex; ring of disk-corollas merely puberulous or the ‘corollas glabrous; recep- tacular bracts 5-6 mm. long, linear-lance oe obtusish, hyaline, not ae with the achenes ; 5 mm. long, obcompressed, the y of the — rk brown, rounded on the back, awns from the vat Arid slopes, often on serpentine, Sonoran Zones; foothills of the Sierra Nevada, California, from Butte County to Tulare County and those of the Inner Conse Ranges from Contra Costa County to Stanislaus County. Type locality: upper Sacramento Valley. March-April. 9. Coreopsis site ee L. Garden Coreopsis. Fig. 5197. Coreopsis lanceolata L. Sp. Pl. 908. Perennial herb 2-6 dm. om — glabrous except for the lower leaves, with several to SUNFLOWER FAMILY 135 many erect or ascending stems arising from a woody rootstock, the stems branching, leafy at or near the base, rather naked above. Lower leaves 10-15 cm. long, long-petiolate, spatulate to oblanceolate, nda nar or with 2 or 3 lateral lobes at the base, ‘usually: hirsute, the upper leaves reduced in si a i b ner 8-12 mm. 1.5-3 cm. long, obovate or cuneate, 2-3-toothed at the apex ; receptacular bracts 4-6 m ki : aches es 2.5-3 m m he body bl mm. lo: 2. long, obcompressed, the black, orbicular in outline, the wings somewhat incurved ; saciotte of 2 espe fimbriolate teeth. of the United States often planted in gardens in Bae: Pacific States and sometimes ba terry locally established (Sake Cruz Cou mney pc tapi Thomas; Santa Barbara County California, wai F. Smith); * pasa ring naturally from Michigan to Florida and to Texas and New Mexico. Type iocality: unknown. May-Jun For complete synonymy re eN, a Fl. II, 2: 18-19. 1955. 10. Coreopsis tinctéria Nutt. Calliopsis. Fig. 5198. Coreopsis tinctoria Nutt. Journ. Acad. Phila. 2: 114. 1821. arn bicolor Reichb. Ic. Pl. Cult. 2: pl. 70. 1823 ect glabrous annual 6-12 dm. high, usually single-stemmed, the stem leafy, much branched, the hae angulate. Leaves opposite, 5-10 cm. long, subsessile or short-petiolate, pinnately IA, j RS 5196 5194. Coreopsis californica emer Coreopsis still 5195. Coreopsis douglasii 97. Coreopsis parte 136 COMPOSITAE or bipinnately can into mene or linear- . etn aa the upper leaves usually un- divided; heads y, 2-3 cm. wide, more or less mbosely arranged, the peduncles 4-10 cm. ene Pg outer p hylan about 8, ig men shine. c r tri beer ain Sioa 2 mm. long; inner phyllaries ‘ r “aes -lobe oe me a8 =6 23 28 o i=" re) > 3 - a wn a a o ! ch ° wes om + n Qu 4 a o @ Qe o o Q o — canal “3 Q cS ro) ot =z my Q - 7 8.5 S96 = o = mm. ion. WwW ase; 1S or less filiform, reddish; disk-achenes of scene length, 1. mm. long, linear Pula, com- pressed, wingless, black, glabrous or nearly — pappus obsolete A native of central United States much rese’ hae atkinsoniana, occurring in gardens and occasionally becoming locally established; Washington and i i Shae Cee Stanislaus, ae, Tulare Counties, Miagaatiol ae ded Casing a Red ae 30 8, Cillected ‘by Nuttall. June— oe For ‘complete synonymy of the various mer. FI. IT. 2: 30. “Coreopsis basalis var. wrightii (A. Gray) Blake (Proc. Amer, Ac rt 51: 526. 1916) has been collec San Diego County, California. re differ rs from C. tinctorta, which it much resembles, in having the outer pat ald linear to linear-lanceolate and spreading, 5-8 mm. lon ng, and nearly equaling in len gth the ovate inner phyllaries. i ae atkinsoniana Dougl. ex Lindl. Columbia Coreopsis. Fig. 5199. Coreopsis atkinso a Dougl. ex Lindl. Bot. Reg. 16: . hikes 1830. Calliopsis saci ook. Fl. Bor. Amer. 1: 311 Annual (or bie eek aeally single- ae 0.4-1.3 m. high, glabrous throughout, ini stems lender, erect-bra d, siviate-aneulat e, rather naked abo es opposite, the low 5-16 cm. c i tely i cy is 2 cm. e€ a Oo coat 5-15 cm. long; outer phyllaries 7-8, ca friangukal, obtuse, 2-3 mm. lo ong ; inner pees 8 mm. long, ovate, brown with yellow margin; ray-flowers eet the rays about 8, wedge- vate, 1-2 cm. long, golden-yellow with purple-brown basal sp ; disk p ie -brown; ee tacular bracts linear-oblong, orange-red, with a scarious margin Pai sk- -ac cheaes narrow ly oblong, about 2.5 mm. long, glabrous, often papillose, 2. -winged, the thin’ wings half as wide as the body or less; pappus none or of 2 small teeth. River banks and lake shores, oe Zones; British Columbia south to northern Oregon (along the Colum bia River) and eastern Washington east to Idaho and Montana; also north- central United States and tetconiaed elsewhere in the United States. Type 1 ocality: Mewrice [Mensies?] Island, in the Columbia River. Collected by Douglas. May-—Sept. Probably included in mixed wildflower seed-packets with C. tinctoria and becoming locally established around dwellings elsewhere in the Pacific States. Guizdétia abyssinica (L. f.) Cass. Dict. Sci. Nat. 59: 248. 1829. An annual species which resembles Biden but has ; sn e achenes has been found growing spontaneously in San Francisco and Santa Barbara gelatin Califor 14. GALINSOGA Ruiz & Pav. Fl. Peruv. 110. pl. 24. 1794, nual herbs, erect or diffuse, more or less pubescent to glabrate. Leaves opposite, simple, thin, triplinerved, the mar rgins entire or toothed. Heads rather small, radiate, in : a me the ray-achen [ Name in honor of Mariano Martinez de Galinsoga, a Spanish botanist of the eighteenth centu A genus of about 6 species, madas of southern United States and Central and South America, Type species, Galinsoga verve ora Cav. Pappus of disk-flowers tapering to an awn; pappus of ray-flowers prese Pappus of disk- flowers truncate or rounded, the margins densely and sia fimbriate; pappus . Vis dees want- ing or vestigial. . G. parviflora. 1. Galinsoga ciliata (Raf.) Blake. Ciliate Galinsoga. Fig. 5200. Adventina ciliata Raf. New FI. Pt. 1: 67. 1836. soga parvi, 1836. alinsoga aristulata Bickn. Bull. Torrey oo 43: 270. 1916. G eG ciliata Blake, Rhodora 24: 35. Freely branching, annual herb i = dm. high, the stems, particularly the upper part, and the peduncles rather thickly beset with spreading hires and bint igen dular. Leaves deltoid- i long, the blade 2-5.5 cm ovate, the les 1-3 ¢ on | 2-5.5 -3c¢ e an see what ie on the petiole, the margin typically coarsely toot thed, ‘sparsely fiveube: involucre 2-3 mm. the phyllaries ovate ; ray-flowers 4-5, the rays white, 1-2 mm. long; kc flowers iow, but little surpassing the pappus-paleae; achenes black, 15-2 mm. long, hispidulous, obscurely angled, the disk-achenes compressed, hispidulous on the inn get rere as of ray- and disk-flowers 9-14, mere or less fimbriate- ciliate and tapering ne ste places and gardens; occurr weed in British ome and western Washington, and widespread in peated and northeastern United a ig aed adjacent Canada. Naturalized from Mexico. June—Oct. SUNFLOWER FAMILY 137 2. icnaadins parviflora Cav. Small-flowered Galinsoga. Fig. 5201. Galinsog iflora Cav. Ic. 3: 41. pl. 281. 1794. vicki -rooted, annual herb 1.5-6 dm. high with long internodes, the stems and alleges with short appres sed pubescence and some spreading glandular hairs or nearly naked. Leaves ar ly ovate to tareibatiees ovate, the petioles 5-12 mm. long, the blades 2.5-4 cm. long, .5-2.5 cm. wide, the margins crenate-denticulate, meri enti ; i : w rit y about 1 mm ng, fibrous 2-3 mm, tube villous ; disk-flowers yellow, shorter than the phen sth Sey ing; disk-achenes about 2 mm. long, black, ’ or rounded, slightly angled, sparsely hispidulous, the pappus-paleae 9-18, run babeec eat the saan finely and repsed fimbriate Waste plac nd orchards or garder curring in ‘tit Pacific States in Inyo County, California, and cis montane ral ae Ealitornix: also eaeaL yvetes and southern United States. Introduced from South Amer rica. Aug.—Feb. 15. BEBBIA Greene. Bull. Calif. Acad. 1: 179. 1885. uffrutescent ETOP AY, antearately branched, nearly leafless, strongly scented. Leaves opposite below, alternate a ar to triangular-ovate, entire or hastately lobed. Heads medium-sized, light calle ies or few pig tips of branches, discoid, many-flowered. 5198. Coreopsis tinctoria 5200. Galinsoga ciliata 5199. Coreopsis atkinsoniana §201. Galinsoga parviflor 138 COMPOSITAE Involucre hemispheric, 4-5-seriate, strongly graduate, the phyllaries ovate or oval to lanceolate or lance- linear, ke outer ai eg a the inner thinner, dry, usually whitish. i ach subentire at base with ovate terminal appendages. Style- cami recurved, with subulate i chenes s c, hispidulous appendages. Achenes slenderly obconi Spore Ngee mpressed, weakly angled, densely silky. Pappus of about 20 slender, equal, 1-seriate, eae awns longer than the he [ for Michael Schuck Bebb, 1833-1805, student of American willows. | A monotypic genus of the Sonoran region. A. Bebbia jincea (Benth.) Greene. Sweetbush. Fig. 5202. Benth. Bot. Sulph. 21. 1844. Ritbia juncea rani. Bull. Calif. Acad. 1: 180. 1885. Suffru cape Suge lecrnnag shrub 1 m. high or less; stem and hak eg oy eee slender, r lan glabrous, pale n, the stem in age whitish-barked. Leaves remote, linea ce-linear an nd entire or the sean aa ia and with one or two pairs of hastate lobes 3 cm. long or ae the upper y and squ orm; heads about 1 cm. high, ly long-peduncled, the pe- e : o (inner) lanceo- late or linear-lanceolate, ne acute or apes nate, the outer subherbaceous, SS ee i whitish or ia ong, pappus whitish, Bebbia juncea var. 4spera Greene, ‘Bull, Calif. Acad. 1: 180. 1885. (Bebbia aspera A. Nels. 273. 1904.) Plant more or less densely hispidulous with te lly tuberculate-based, sometimes deciduous te irs. et ae but atid sort to Inyo County, California, and southern Nevada and east to New Mexico ona Ana 16. EASTWOODIA* Brandg. Zoe 4: 397. pl. 30. 1894. Xerophytic shrub, somewhat glutinous. Stem white-barked, striate, glabrous, brittle. Leaves alternate, essentially linear, entire. Heads discoid, many- -flowered, yellow, solitary ofte seri Corollas funnelform , the 5 lobes long. Style-branches linear, the acuminate appendages equaling the stigmatic portion. Achenes more or less quadrangular, narrowly obpyramidal, silky-pilose especially on the ribs. Pappus of 5-6 free, persistent, linear, acute or acumi- nate, firm paleae about two-thirds as long as the corolla. [Named in honor of Alice cong hae: California — t.] 1. Eastwoodia élegans Brandg. Eastwoodia. Fig. 5203. Eastwoodia elegans Brandg. Zoe 4: 397. pl. 30. Rounded desert shrub 3-10 dm. high, erect- Pe ge the herbage minutely and sparingly hispidulos, mene econ, pallid. Leaves lin eee or the upper — 2.5 cm. long, to 3 mm. wide; heads dopeun “sb bose, 1-1.5 cm. thick; involucre age at base, hr mm. hist: arellas about 5.5 mm. long; achenes Shout 2 mm. long; ness dry —. up to 2,500 feet, Upper Sonoran com foothills on the west and south sides of San Joaquin Vater” ‘Caltfornia, from Alameda County to Santa Barbara County east to Tehachapi Mountains, Kern County. Type locality: not definitely stated. April—July. 17. MELAMPODIUM L. Sp. Pl. 921. 1753. Herbs, rarely suffrutescent, glabrous or pubescent, the stems coped dichotomously branched. Leaves opposite, entire, dentate or pinnatifid. Heads small or medium, usually solitary in the forks and terminal, heterogamous, radiate, the rays pistillate, fertile, the disk perfect, sterile; corollas yellow or white. Involucre 2-seriate; outer phyllaries 3-5, broad, herbaceous, sometimes connate at base, often accrescent; inner phyllaries as man as the ray-flowers, each closely a an ovary, coriaceous and often appendaged, deciduous with the enclosed achene at maturity. Receptacle convex, the receptacular bracts membranous, eniendine the disk-flowers. Rayscontitie few or several, 1-seriate, oblong or broader, entire or denticulate. Disk-corollas regular, tubular, 5-toothed, sterile. Anthers bluntly cordate at base, with ovate terminal appendages. Style in ray-flowers * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 139 . a in disk-flowers undivided, — hispidulous. Achenes of ray obovoid, cov- by the variously muricate and often hooded or beaked inner phyllaries which are eae duous with the achenes. foe tebe ar disk sterile, stipitiform. Pappus none. [An ancient name of black hellebore from the Greek, — trarily transferred to this genus. ] A genus of about 12 species of subtropical America but mostly Mexico; one or two introduced into the Old World. Type species, Melampodium americanum L. Li Te a nadie (Cav.) H.B.K. Melampodium. Fig. 5204. Alcina perfolia Tc. 1 ot.15- 179 Melampodiu bam ations * = K. Nov. ahr & Sp. 4: 274. 1820. Coarse annual, sparsely hispid especially toward the nodes, the peduncles puberulous, the stems about 1 m r less, dichotomously branc eaves opposite, in rat ew pairs, the internodes long; petioles broadly wing-margined, dilated at bas d usu connate; blad deltoid-ovate or rhombic-ovate, often hastately lo at , acute, serrate or my ge 5- long and wide, rough above, pidulous and gland-dotted th, green on ides, thin; heads solitary in the forks and terminal, on —— 2-11 cm. long; phyllaries 5, se or oval, mm. long, 0 —. ciliolate, united at ex e base; rays about 8-13, _ yellow, much shorter an involucre ; fruit rounded on back, m ae acats the tip, about 5 m Occurring as a bast in swampy and waste places, vicinity of Los ene ag ri Los pee ies: California; isimoducad ten om Mexico; occurring also in Guatemala. Type locality: Mex y—Nov 5202. Bebbia juncea 5204. Melampodium perfoliatum 5203. Eastwoodia elegans 5205. Sanvitalia abertii 140 COMPOSITAE 18. SANVITALIA Lam. Journ. Hist. Nat. 2: 176. pl. 33. 1792. Simple or branching, annual or perennial herbs. Leaves opposite, entire, petioled or sessile. Heads heterogamous, radiate, terminal on the branches or sessile in the upper to narrowly conical, the receptacular bracts scarious and rather firm, ge ser inet and an e pappus oo or of 1-2 slender awns or teeth. [Name in honor of a noble Italian family, Sanvita Ses ge my of about 4 species, natives of southwestern United States and Mexico. Type species, Sanvitalia pro- cumbens 1. Sanvitalia abértii A. Gray. Abert’s Sanvitalia. Fig. 5205. Sanvitalia abertit A. Gray, Mem. Amer. Acad. II. 4: 87. 1849. annual from a taproot, 1-3 dm. high, simple or with ascending opposite branches, the stems hispidulo us Lice curved asce tice hairs, usually becoming glabrate. Leaves linear- lanceolate to lanceolate, short: than the internodes, the petioles 2-12 mm. long, the blades 2.5-5 cm. long, cuneate at base and somewhat decurrent on the vietiole, the lea if “margins and captes ces hispiduus with pustulate-based hairs ; be ‘age ubequal in 2 series; receptacle narrowly cal to nearly subulate, the receptacular bracts much be pin ssing the disk- flowers; diner “6-10, the mm. long, emar a glee lobed) and many-veined ; disk-flowers about 1.5 mm. long, long, 4-angled, the angles often compressed and w — — conspicuously and irregularly warty, ‘black- ish me sometimes se ith a chalky wat ates epappo rid mesas and s Sonoran = in Sah ies x Clark Mountain, eastern San Bernardino poy (Root 4954, 4893), ea py cng in northern and central Arizo oe Pasa also Chihuahua and Sonora. Type v2 “between Bent’s Fort and Santa 6,” New Mexico. pores Subtribe AMBROSIINAE Heads heterogamous (a few unisexual staminate heads present on inflorescence of Dicoria); pistillate flowers and achenes not ccs sagpe in a bur-like or nut-like involucre. eee more or jee urgid, whee “ead phyllaries in 1 or 2 series, the inner when present not noticeably larger and not min: a accresce Anew densely wRous; rahe their lobes linear-filifor 19. rages villous; a entire, lobed, or wea se cleft. 20. scale ae and wit pectinate or toothed wings; inner phyllaries larger than the a) Eeakine ac- tcoria Heads unisexual, ek staminate borne above the pistillate; pistillate flowers and achenes in a nut-like or bar-like volucr Phyllaries ofthe the staminate heads united; fruiting involucres winged, tuberculate, or if hooked spines present Fruiting ede with conspicuous transverse wings; leaf divisi leaves linear-filif 22. Hymenoclea. Fruiting involucres without transverse wings; leaves toothed or pinnatifid, the lobes not linear-filiform. ruiting involucres unarmed or with a single row of tubercles or teeth. 23. Ambrosia. Fruiting involucres with several spines in more than 1 row. 24. Franseria, Phyllaries of the staminate heads free; fruiting involucres conspicuously covered with hooked spines or prickles. 25. Xanthium. 19. OXYTENIA Nutt. Journ. Acad. Phila. II. 1: 172. 1848. Strigillose perennial, shrubby and pth f with striate stems. Leaves alternate, petioled, pinnately divided nearly to midrib or the upper entire, the divisions linear -fili- form, revolute-margined. Heads eae aang disciform, numerous, spicate-panicled i t of the pistillate flowers reduced to a thick fleshy ring, of the perfect ones funnelform, 5-toothed, Saeed and as pilose above. Filaments free; anthers lightly te, at isj wers un , at apex slightly dilated and Achenes obovoid, convex dorsally, flattish, pecteali: l-ridged on each face, “ oc aes pappus none. SUNFLOWER FAMILY 141 [Name Greek, meaning sharp, referring to the rigid, subulate-pointed divisions of the leaves. A monotypic genus of the southwestern United States. 1. Oxytenia acerdésa Nutt. Oxytenia. Fig. 5206. Oxytenia acerosa Nutt. Journ. Acad. Phila. II]. 1: 172. 1848. Strigillose shrubby perennial 1-2 m. high, the stems conspicuously striate, greenish, leafy, erect branched above, sometimes leafless wl rush-like. Leaves petioled, 4 . long, pin- ly e€ i lobes sometimes wi 3 similar lobes, the rachis, lobes, and usually the narrowly winged petiole all strongly revolute- margined, 0.5-1.5 mm. wide, the Lists. el since: heads 3-4 mm, thick, usually not nodding achenes 2 mm. long. Dry hei and along creeks, Sonoran Zones; southwestern Color o New Mexico, southern Utah, Nevada, orthern Arizona, and Inyo a California, in the Death Valley phe Pog Type locality: ‘Rocky Mountains, a Upper California.” July—Sep 20. IVA L. Sp. Pl. 988. 1753. Annual or perennial herbs or shrubs, glabrous or pubescent, sometimes tomentose Leaves usually opposite at least below, linear to broadly ovate, entire, toothed, or dissected, sessile or petioled. Heads heterogamous, isciform, nodding, racemose, spicate, spi anicled, or scattered on the branches, sometimes ebracteate ; outer flowers (1-8 istillate. inner (3-20) perfect, ret corollas w ne . Involucre nearly hemispheric, usual double, the outer of 3-9 free or rarely connate, sometimes imbricated, mostly oval or ovate, subequal, subherbaceous phyllaries; the inner usually present, of f embranous phyllaries, opposite the outer phyllaries, each subtending d someti alf-enwrapping a pistillate flower, linear-spatulate to broadly obovate. ag va sm a basi imes n a jie parse i ft pistllae | flowers tubular or campanulate short, aimtiaes ‘tn cniecte. someti a 4-5-toothed, in some species reduced to a fleshy ring or obsolete; corollas of perfect pari ‘oiniee centile or broadly funnelform from a aka base , 5-toothed. Fila- ents free or somewhat connate ; anthers lightly connate, at m maturity isjunct and usually pitt Be | with subentire to sagittate bases and inflexed terminal appendages. Style in the apex; of the disk rudim oie Pappus none. amed because of tts sim a Rpm in smell to Ajuga iva; the name of the latter derived from an old Latin word signifying an aborti- facient. A genus of about 15 species, all North American. Type species, Iva annua L. Heads racemose, mostly solitary in the axils of reduced leaves; inner phyllaries much narrower than the ovaries (and a sero 5 corolla of pistillate flowers evident, tubular, glandular; leaves obovate to linear, entire; plants peren Phyllaries —— at least to middle; herb, from running rootstocks. 1. I. axillaris. Phyllaries free; plant gen 2: is hee saa Heads scattered on the upper anches or spicate-panicled, mostly ebracteate; inner phyllarie der than ov: aries (and achenes) and infolding them on the edges; corolla of pistillate flowers ite or wlabrods. es serrate to bipinnatifid; plants annual. Plant 15 ot high or abs epic bipinnatifid, small; corolla of pistillate flowers tubular, ge ete below, vil- lou ensis. Plant up ‘ 2 ~ high; leaves sharply serrate, large; corolla of pistillate flowers short, toblgream campanlate, glabrou: 1. Iva axillaris Pursh. Poverty Weed. Fig. 5207. Iva axillaris Pursh, Fl. Amer. Sept. 743. 1814 va axillaris 8 robustior Hook. Fl. Bor. Amer. 1: .. = aris var. kebesten ens A, Gray in Torr. noe ing SS 17: 350. 1874. Perennial herb from ree rootstock: stems several, scattered, simple or simply branc ssi c , strigose, incurved- or rar rely s spreading- pubesc ent, somewhat glandular. Leave 18-60 cm. hi s : opposite below, alternate whats obov ae to lance-elliptic, 1.54.5 cm. long, 4-15 mm. wide, obtu or acute, narrowed to the sessile or subsessile base, entire, thick, luce pubescent ii “ye the stem; heads nodding, 4-7 mm. thick, solitary in the axils of the ‘usually not much reduc leaves, Se scree involucre gamophyllous, shallowly or to middle about 5-lobed; Sistine flowers perfect, about 6-18; -signie sisneciensseic ar, 2-3 mm. long. ac troublesome weed, Transition and Sonoran Zones; Mani- toba iss os ali’ oum'to to southern pope: and Nevada, Utah, New Mexico, and Oklahoma. Type locality: “In Upper Louisiana.”” May—Sep 142 COMPOSITAE 2. Iva hayesiana A. Gray. San Diego Poverty Weed. Fig. 5208. Iva hayesiana A. Gray, Proc. Amer. Acad. 11: 78. 1876. Suffrutescent, about 1 m. high, sparsely a pee or Psion igs? oe Haagit saad on the leaves, and sess sile-glandular ; branc hes erectish. Lea opposite below, alter ve, obovate eol in m ; s tly s ea pedicelled, 3-7 mm. thick; pistillate flowers about 5, the ah up to 20; involucre of about 5 free, i enes 2 mm. long. On brackish alkaline flats, Upper Sonoran Zone; San Diego County, California, south to central Lower Cali- fornia; also on Cedros Island. Type locality: Jamul Valley ie vale of San Diego, California. April-Sept. 3. Iva nevadénsis M. E. Jones. Nevada Poverty Weed. Fig. 5209. Iva nevadensis M. E. Jones, Amer. Nat. 17: 973. 1883. Chorisiva nevadensis Rydb. N. Amer. Fl. 33:9. 1922. Diffusely br. anched, single-stemmed annual from _ taproot, 7-15 cm. high, cinereous with mostly incurved hairs and somewhat sess aH ices Leaves alternate, petioled, the blades ovate or deltoid in outline, 8-18 mm one! nately hedewstn the ultimate segments rather few, obtuse, ag ei the “ani of infloresce ce pees smaller, mostly cuneate and 3-lobed; heads —3 r in r ‘etal € a phy ae 3, free, ovate, with short, recurved-sprea ing, obtuse, herbaceous tips; acular bracts subtending the pistillate flowers broadly cuneate, villous, stg bbe Idiot the achenes ; pistillate flowers usually 3, the perfect ati sevens obovoid, planoconvex, 2 mm. long, at length crustaceous-muricate on margins a both In desert places, Lower Sono n Zone; western Nevada = ane t California from Mono County to the Death Valley region, Inyo County. Tyee locality: Hawthorne, Mineral Cone Nevada. June—Oct. 4. Iva now ltites: Nutt. Marsh-elder. Fig. 5210. Iva xanthifolia Nutt. Cyclachaena xanthifolia Prench. a ce Hort. Frankf. 4. 1836. Coarse annual up to 2 m. high, simple or branched; stem glaucescent, essentially glabrous below. cl vie and wiandali ar in the rp sesh i nce. Leaves opposite least below, slender-petioled, ades o 4-] g e, abou wid umin nate, 2.5 ~ co La 3 3 3 rt) So © Fd o bo] =a = > on o wn wn ao 3 ) na. 0.9 < ry) ce SF mh 5 o o 8 ~ oe bas | = =| +o, a ) 3 roy wm > ° 5 [ao ws a iz o 4 ao 2D a) o ° boy ° ey) ‘fe s 3 =} » o ee ey Papillose 4 in lines, ee: ut ye d in waste places, Arid oe isthe Zone; Alberta and Sasleatehesthn to Nebraska, and south hh chetcei Washington and Idaho to freee aay and pee Sickish. also as an occasional weed east to the Atlantic. he locality: Fort Mandan [North Dakota]. Au ug.—Oct 21. DICORIA Torr. & Gray ex A. Gray in Torr. Bot. Mex. Bound. 86. 1859. Coarse, much-branched, weedy, autumn annuals, aromatic; pubescence of the stems. s the upper surface, hispid hairs eaey port on ae lower epee juvenile leaves linear to ed and (in ours) continuing to enlarge ates the seeds have matured. “ei ater small, flat, with narrow, scarious, ep aecar ny bracts. Pistillate flowers 1-5, withou corollas; staminate corollas about 6-15, regular, funnelform. Stamens obtuse at ey free, the filaments united. Style-branches oer: ’ Achenes dorsoventrally ae rsa time i i nate, or pp re) bristles). [Name from the Greek, meaning twice an g. A genus of gs a Laney ag a species, natives of southwestern United States and adjacent Mexico. Type species, Dicoria . Gra Achenes 3.5-4.5 mm. long, the margin one-fourth or sore the width of the body of the achene, the teeth or toothed lobes distinct; stems with many pustulate-based, spreading, hispid helet cae — = pubescence. . D. ee ens hispidula. Achenes 4.5-6 mm. long, the margin one-half to pon ag fiat ae of the achene, ie ‘aaiead or Sidlidek — contiguous; stems with appressed pubescence g hispid ~ _ rarely completely abse Leaves oval to orbicular; margin one-half or nie aa rn pe the body o: achen: Somes Coteenso ere and adjacent Mexi pte 5 gees Leaves soon ovate; ceupha about one-third the width of the body of the achene; southern N anes and adja- t California. D. clarkiae. 5207. Iva axillaris 5208. Iva hayesiana SUNFLOWER FAMILY 5203. Iva nevadensis 5210. Iva xanthifolia 143 144 COMPOSITAE 1. Dicoria canéscens A. Gray. nent Dicoria, Fig. 5211. Dicoria canescens A. Gray in Torr. Bot. Mex. Bound. 87. pl. 30. 1859. Dicoria calliptera Rose & Standl. Contr. U.S. Nat. Herb. 16: 18. es i2. 1912. Coarse shine, 5-10 Matte oo the stems Sao eading from the base, avant much branched, see clothed with upward-curved, appt ssed hairs, these tardily deciduous, and with a few ustulate- Secor. hispid | fae Rea lea ee cm. long, reduced as rather thick, the petioles m ad i i d partially winged on the ridge, the outer often partly corky, the winged margin one-half or more the width of the body of the achene, with broad, ic lidate or bp es > fakes ually | straw-colored, thickened toward the base In sand, Lower Sonoran hase lege and Imperial Counties, California, southwestern Arizona, and north- bela Sonora. co locality; “Tn the andy desert of the Gila and of the Colorado. ” Collected by Emory. Oct.- Dicoria canescens subsp. hispidula Beck, Aliso 4: 101. 1958. (Dicoria hispidula Rydb. N. Amer. FI. = 12. 3922.) ig a D. canescens subsp. canescens and D. ‘clarkiae, Sas: pubescence of the ste m mu ch the same body < the achene. In sand, Lower Sonoran Zone; brani Mojave Desert, “california, south through t chella Valley. Type locality : Whitewater est. Riversi de County. Differing princi ally by the more Priefer ee and narrower margins of the a es. Growing with D. canescens in the Coac ella Valley. oblongifolia i N. Ace Fl. 33: 12. 1922, is said by Rydberg to grow i southern California and ices Califtrni nia. Soh s some question about the area in which the type of nia oO 7 Bale was Greg neg on almer (n numbers, many of them from the Mojave Déeiert a io The type locality, > ay cou George, Utah, if the year given on the label is correct. If it is not, the type locality could be the Moiave Desert. The achenes of the ae are about 5 mm. long, the margined lobes about 0.5 mm. oy delat double-toothed, darkened at the base, somewhat hairy or glandular- dotted. 2. Dicoria clarkiae Kennedy. Clark’s Dicoria. Fig. 5212. Dicoria clarkiae Kennedy, Muhlenbergia 4: 2. fig. 1908. Dicoria canescens subsp. clarkiae Keck, Aliso 4: 101. 1958. Coarse annuals up to 7 dm. high, stems spreading from the base and branching above, striate, poate upward-curved Sais and some longer appressed hairs, becoming scabrous in age. Upper leaves 1.5-5 cm. long, the petiole about one-half or less the length of the blade, broadly ove er 3 (20 -mar- gined; achenes 5-6(7) mm. long, the body of the achene mostly 2 mm. wide, , more often more or less oblanceolate, inner surface brown, flat, 1-nerved or ridged, — peta mina often glandular-dotted and sparsely hirsute, the outer rounded, ri ay ed, bro — — be by moe cas e or less gland-dotted and sometimes hirsute, the ma n0. more at base e achene, peg: venly or unevenly lobed ae toothed, acc anne ae ened whine eked to the body eens. In sand, Upper Sonoran Zone; act Cou Nevada, to Inyo County, Savors, and adjacent San Bernardino County. Type locality: Soda Lake, Churchill ice 3 ed i. Oct.—Nov. Seemingly merging with Dicoria fog aioha subsp. hispidula in its southern When life-cycles ae all the species of Dicoria are thoroug studied, artinelae with po possibility rm on of Dn of the nged margins of the achenes, more sae Sete change may become necessary. 22. HYMENOCLEA Torr. & Gray ex A. Gray, Mem. Amer. Acad. II. 4: 79. 184 Monoecious or subdioecious shrubs with white or straw-colored bark and greenish, striate, glabrous or sparsely hispidulous, more or less resinous branches. Le alternate, filiform or linear-filiform, entire or pinnately few-lobed with filiform np emir gree volu ike tie panicles. Stamin te heads a 5-flowered ; Gaivaucre saucer-shaped or turbi- — of —— 5-6 mos eo ov: er acre yllaries united about to middle. Receptacle few spatulate, set lar, and ciliate, membranous bracts. Pistillate involucre purse ubenaed by 1-3 usually ovate and subherbaceous or mostly scarious phyllaries, SUNFLOWER FAMILY 145 ee ae fusiform, angled below, terete and tubular-beaked above, indurated, bearing about 5-12 cuneate or oblanceolate to flabella “i horizontal, scarious wings either in a single whorl at tidale or spirally from base to middle or rarely above it. Staminate corollas funnelform, 5-toothed, somewhat glandular ; pistillate flowers Aithout corolla. Stamens 5; filaments usu- ally lightly connate to middle or nearly to apex; anthers lightly coherent, with s sagittate bases and deltoid, inflexed, terminal appendages. Style in staminate flowers undivided, dilated at apex and there fimbriate; in pistillate flowers with 2 linear, obtuse, exserted, stigmatic branches. Achene aap enclosed by the fruiting involucre, obovoid-fusiforn n, black- ish brown, aes epappose. [Name Greek, meaning enclosing membrane. Ag of cies, ot 3 ‘the southwestern United States and northern Mexico. Type species, Hy- wekoctee pene Purr: “& Gra Wings of fruiting involucre spirally arranged, 5-8 mm. wide. . H. salsola. Wings vie dei involucre in a.single whorl, or » 1 or 2 ces wings above or below ae 1-5 mm. s 5-7, flabellate or reniform-orbicular wide — Wings 7-12, mostly cuneate or obovate, 1-2 n mm. Sie. mon 1. Hymenoclea salséla Torr. & Gray. White Burrobush. Fig. 5213. Hymenoclea salsola Torr. & Gray ex A. om y, — — Acad. IT. 4: 79. 4 Hymenoclea fasciculata A. Nels. Bot. Gaz. 270. Hymenoclea fasciculata var. patula A. Sea op. cit. fs ae 1909. Shrub 1-2 m. high, erectish or spreading-branched ; branches sparsely hispidulous when young. Leaves igre deciduous ai pt from the younger branches, pica r linear-filiform, 2-6 cm. long, e, € or sintately few-lobe a) obtuse, green and often sparsely hispidulous beneath, Abauely eanescent- hispidulous and often rongly Raabe De -marg — above; heads - mixed or the staminate often above the pis a. pre ee or solitary in the aah of te —. iis te involucre about >, mm. wide, owered; nistillate involucre tu = sig siform, abou mm. long, _itagnel wg iboat 12 spirally arranged, suborbicular or flabellate, often mucronate, scarious wing mm. wide, their broad, short, stipe-like bases usually deeply sineAs teal cone- ike gee dry, 12-14 mm. in diameter when wings are opened. Desert washes and alkaline ao, Lower Sonoran Zone and lower part of Upper Sonoran Zone; San Joaquin Valley in eles County, California, and eastern Santa Barbara and Ventura Counties (Cuyama Valley: ; Mount Pinos region) southward along the pees slopes of the mountains to San Diego County and eastward across the Mojave Desert and ranges of the Death Mer J region to southern Nevada, te rn Arizona, and southern Utah. pape apeality: uplands of the Suiace River, San Bernardino County, California. Collected ag Fremont. March- July. Desert Pearl. 2. Hymenecies ere = Southern Burrobush. Fig. 5214. H lea pentalepis Rydb. N. Amer. FI. 33: Hymenoclea Ange Mee Nels. Amer. Journ a 25: 11 17. 1938. iemenecies salsola var. pentalepis L. Benson, op. cit. 30: 631. 1943. tricately cre shrub about 1 m. later weg — dioecious or essentially so, specced slender, prea andular-granular and m or less resinous, sparsely hispidulous when Leaves mostly rciceas incites filiform, entire mn nati pie lobed, 0.5-5 cm. long, about 0.5 m 5212. Dicoria clarkiae 5213. Hymenoclea salsola 146 COMPOSITAE wide, obtuse, peg and saa ye beneath, densely canescent-hispidulous above but strongly in- volute-margined so as to conceal the hairs ; ee in ies ie! ve ong a in pein vit the branches in i axils of park — leaves, the e; stamin in- volucres a a ut 7- flowered; "Bistillate pot Rae S htoce aI t subo in a single whorl or shortened spiral at the middle, these 5-7, flabellate or reniform-orbicular, with a short broad a or concave base, usually obtuse and often mucronate at apex, pinkish or straw- color wide sikien de ature, additional wings sometimes present, the whorl about 7-8 mm. in ftom Sie spread. Desert washes and alluvial fans, r Sonoran Zone; northern border of the Colorado Desert and in the vicinity of Needles, San alles oe Satiforaia. southward to the eastern side of Lower California as far south as Santa Rosalia eastward to southern Arizona and adjacent Sonora. Type locality: Pima Canyon, Pima County, Ariz — Feb.—June. The western specimens are usually more densely flowered and the flowers are a little larger than the more typical anticciad { . Arizona and Sonora. fl a eerenne les monogyra eu & Gray. Leafy Burrobush. Fig. 5215. H gy r. & Gray ex A. Gray, Mem. Amer. Acad. II. 4: 79, 1849, ae shrub wy m. high with numerous erectish branches, eee very pa at least above, branches glandular-granular. Leaves filiform, 1.5-8 cm. long, 0.3-1 m wide, entire or the lower pinnately divided, callous-pointed, green n and labrous or glan ular- pe haba eath, densely canescent-hispidulous above and strongly involute-margined, aber with axillary fascicles; hea i lusters on the b hes, formi ong, leafy, spike-like panicles, the staminate and pistillate usually intermixed; stam involuc : b ‘ 2 red; pis- staminate tillate involucre about 4 mm. long, turbinate-fusiform, glandular, bearing at or near base 2 or 3 scarious suborbicular bractlets and at middle a single whorl of 7-12 mostly cuneate or obovate, acute or obtuse, scarious wings 1-2 mm. wide, the whorl about 5 mm. in diameter when spread. Sandy washes and desert -schgeeson mo ran Zone; San Bernardino Pee gercs octane oe and eastern San Diego County seat anced eae dy = ene California and eastw. Ss; also northern Mexico. Type locality: Sp oSty the valley of the Gila,” ’ Arizona. Collected by Emor og ot: pee 23. AMBROSIA L. Sp. Pl. 987. 1753. Annual or perennial, normally monoecious, pubescent herbs. Leaves opposite - ie below, entire or usually lobed or dissected. Heads unisexual, the staminate noddi axils a e e staminate racemes, subtend d lea small herbaceous bracts. Staminate heads usually many-fl ; involucre gamophyllous, tur- ate or saucer-shaped, more or less oblique, herbaceous, the phyllaries connate nearly a nelform or campanulate, 5-toothed ; corolla wanting in pistillate flowers. Stamens 5 (rarely 4); filaments lightly connate; anther s lightly coherent, subentire at base, with deltoid terminal appendages tipped with an cies ed cusp about as long as the appendage. Style in ae flowers undivided, at apex slightly dilated and fimbriate ; in pistillate ones with 2 linear, exserted, stigmat ic branches. Achene obovoid, tightly enclosed b fruiting involucre, glabrous, epappose. reek, meaning the food of the gods, a al name of various plants, its application to these ‘weeds A genus of — 20 species, nearly all American, one or two in the Motiierreangs region. Type species, Leaves entire or palmately cleft. 1. A. trifida. Leaves pinnately lobed to bi- or si me Leaves green, coarsely lobed vided. Annual; ana thin; at least the lower usually bipinnatifid. 2. A. artemisitfolia. Perennial; leaves thickish, mostly only once- yinnatiod. 3. A. psilostachya. y silky-pilose, ae dissected, the divisi bout 1 mm, wide. 4. A. pumila. 1. Ambrosia trifida L. Giant Ragweed. Fig. 5216. Ambrosia trifida L. Sp. Pl Ambr, eames eon: Muhl, ex Willd. Sp. Pl. e 375. 1805. Ambrosia variabilis Rydb. Brittonia 1: 97. 19 Coarse annual 0 .5-6 m. high, the stems i ne or scabrous above, often ge below. cane. lacking ; - petioles 1.5-3 cm. long, more ne ee wing-margined; blades 4515 oy long, sparse ate e : and dark green ve veined ; 1 to several, many-flowered, terminal and from the upper leaf-axils ; ping os ra = pres een of staminate racemes and in leaf-axils ; staminate Aen sparsely SUNFLOWER FAMILY 147 hirsute, regular, nies short pagep pedicel, strongly 3-nerved on one side; apn pistillate invo ~ cre os mm ted with an acute beak, several-ribbed, the ribs icin in acute tubercles soil, mat and w suse in the Pacific States occurring occasionally as a — in Washing. = and California, pha wae Govinand througho ut the United States and Canada. Type locality: “Virginia, anada uly— 2. panbtpert artemisiifélia L. Annual Ragwood. Fig. 5217. Ambrosia elatior L. Sp. 987. 1753. Ambrosia artemistifolia - op. cit. 988. Ambrosia artemisiifolia diversifolia Piper, Contr. U.S. Nat. Herb. 11: 551. 1906. Ambrosia elatior var. artemistifolia Farwell, Rep. Mich. Acad. 15: 1 nching annual tn dm. high, the stem slender, scsi stri —e = incurved-pubescent, sometimes Basic ges -hirsu Tiaves opposite below, alternate above, triangular or ovate in out- = wly ihe ae petioles 1-5 cm. long, the blades 12-pinnatifd with often toothed d, ia acute divisions, rather thin, pubescent like the stem, paler and more densely ne es Sunes the uppermost leaves often linear and entire; staminate heads in terminal, often 3 mm. in diameter ut 12—32-flo i ac u the upper axils or at the base of staminate racemes, subtended by reduced leaves and a few small herbaceous bracts; fruit turgid-obovoid, 3-3.8 mm. long Ne panear ng beak, this 0.7-1.2 mm. long), pubescent : gla ndular, bearing aie apex ” body 1 -7 usually acute, short, conic a Tee Hy ey una abu ms nt weed nearly throughout the United States and southern Canada south into tro ical America, occasional in our ee (Washington to California); also naturalized in Europe. Type locality: irginia, Pik sylva * Jun t 5214. Hymenoclea pentalepis 5216. Ambrosia trifida 5215. Hymenoclea monogyra 5217. Ambrosia artemisiifolia 148 COMPOSITAE 3. Ambrosia psilostachya DC. Common Ragweed. Fig. 5218. Ambrosia psilostachya DC. Prod. 5: > mbrosia coronopifolia Torr. oe ira 1. N. Amer. 2: 291: 1842, paris sia scalifornc Rydb. mer. a 33: 1922, Ambrosia psilostachya var. pa Sonssh Blake in es. Contr. U.S. Nat. Herb. 25: 581. 1925, Per ie the stems pikomg scattered, 0.3-1 m. high, from running rootstocks, a or hirsute with = esting or spreading hairs, sally branched at least above. Leaves oppo 1 te angular in ou tline, 4-10 ¢ ong, on usually short-winged petioles, thickish, shallowly or r deeply — with fate ae acute, alten toothed or lobed, forward-pointing divisions, strigose or ascending-hirsute or -hispid a nd glandular, the aoe narrower and less divided, ofte Satie re; staminate heads 1- eds in diameter, in usually elongate an n hs nln s, about ‘1 fe) s hairs; pistillate heads solitary or eng is in the upper axils; fruit obovoid, about m. long (the short blunt beak about 0.6 mm. long), glandular, hirsute, often rugose, boat Bg or with 1-6 short, obtuse or acute, conic a around apex = mene Dry or moist places, Sono and Transition Zones; comm we in California, Arizona, and northern Mexico and in Washington, =e hoad pr eg introduced; also lilin eatin en Type locality: between San F saloon and Matamoros, Tamaulipas. Collected by Rertaier April—Oct 4. Ambrosia pumila (Nutt.) A. Gray. San Diego Ragweed. Fig. 5219. Franseria pumila Nutt. Trans. Amer. Phil. oo II. 7: 344. 1840. Hemiambrosia eterocephal ae iwet "Studi L gn. Anem. 61. 1871. Ambrosia pumila A. Gray, Proc. Amer. Acad. 205 217. 1882, ae herb 1-2.5 dm. high with running rootstocks; stems erect, canescent- ot ge at least when young, with mostly appressed hairs and sparsely hispid. Leaves mostly alternate, ovate i i 0 ] or oblong in outline, bi- or tripinnatifid, 5-10 cm. long including petiole, apn ego nd ee silky-pilose wee mo stly appressed hairs, the ultimate divisions mostly linear-oblong, obtusish, cro ss wide; staminate heads about 25-flowered, short-pediceed in bs short = the nvoluere cresaie lobed: pilose ; pistillate heads clustered in the axils of reduc AVES; frit rm is d, 2-2.5 mm. long, pubescent, longitudinally ribbed below, lgoiraly. pak arly scte Alkaline — Sonoran Zones; southern San Diego County, Califorriia, southward to central Lower California. Type locality: near San Diego. June—Sept. FRANSERIA Cav. Ic. 2: 78. 1793. Heads i wae! the staminate nodding, in n ebracteate terminal racemes, the Sota sessile 2 : ay ; th x heads many-flowered, the involucre gamophyllous, saucer-shaped or turbinate, crenate, nut-like or bur-like, bearing few to many straight or hooked, rigid s spines in 2 or more series, the beaks conic, opening obliquely, often econ 2-toothed. Staminate nies mostly funnelform, 5-toothed : corolla wanting in pistillate flowers. Stamens 5; filaments connate or lightly coherent; anthers lightly coherent, subentire at base, with ovate or deltoid, terminal appentieel often ti ped with an inflexed cusp. Style in staminate flowers ndivided, at apex slightly dilated ae ‘imbr iate ; in pistillate flowers with 2 linear, exserted, sti atic branches. Achenes usually obovoid, tightly enclosed by the fr ruiting involucre, glabrous, epappose. [Name in honor of Acetone Franser, physician and botanist of Ma- drid.] A genus of about 30 species, all American, mostly North American but a few on the western coast of South America. Type species, Franseria ambrosioides Cav. Herbs. Annual; staminate involucre with 3 of the nerves thickened and blackish rages 8 fruit armed with flattened straight spines, . F. acanthicar rpa. Perennial; nerves of the er involucre not thickened and blackish below Erect; cokes -4 mm. long, obovoid, cin pid 10-20 hooked spines 0. 8 ek -_ or less; leaves usually greenish, m eee. or less sparsely strigos cies Ba tiflora. Pramaiiea: fruit — or fusiform, 6~9 mm. long, the spines not me pu mm. long; leaves densely ca ae strigo: ostly eate or obovate, crenate-serrate. 3. F. chamissonis, takes bi. to erence ie oa cheutameais bipinnatisecta. Shrubby or suffru Leaves antalya = ee — the lower stem-leaves not more than 4 or 5 cm. long. Leaves Bh oomeas 4 canescent-strigilose, bi- to tripinnately divided into small, —: ovate or obovate divi- sions; fruit pe da and sparsely pilose. . F. dum SUNFLOWER FAMILY 149 Leaves greenish above, densely whitish- or canescent-tomentulose beneath, from crenate-serrate to pin- natifid; fruit densely villous or tomentose Lédees ene ee sinuate-toothed or pinnatifid; fruit. fusiform, 8-10 a. —_ Somer long-villous, bearing about 20 or fewer straight spines. F.e Leaves Bay type ioled, crenate-s aa rt Viruit turbinate-subglobose,, ao _ -_ long, the body densely bargiies ge tina thickly covered with mostly hooked io. . F. chenopodiifolia, Leaves o on both sides, ed stem-leaves 6—18 cm. long. Leaves merely toothed, acuminate, petiolat 7. F. ambrosioides. tae coarsely spiny-t Godied. not pai oS sessile and clasping. 8. : tlicifolia, 1. Franseria acanthicarpa Smee Coville. Annual Burweed. Fig. 5220. Ambrosia eae ibid Hook. Fl. Bor. Amer, 1: 309, Fran montana Nutt. Trans. jer Phil. Soc. IY. 5: ee 1840. Pasi peparas: Nutt. loc. cit. Franseria acanthicarpa Coville, Contr. U.S. Nat. Herb. 4: 129. 1893. Gaertneria acanthicarpa Britt. Mem. ce act 5: 332. 1894. Franseria palmeri Rydb. N. Amer. FI Weedy branching annual, 2-7 dm. hi a stem and branches strigose or hispidulous and often spreading-hispid. Leaves slender- petiole d, the blades ovate or deltoid in outline, cm. long, shallowly or decaty and usually ternately bi- i tripinnatifid, with obovate to linear, obtuse or acute divisions, hook ee and often hirsute, green ve, paler beneath, oo cinereous ; staminate rac ually panicled, the pistillate Auiers at their base and in the upper axils ; staminate slender peainsted. 2-4 mm. thick, about 15-flowered, the Passi about 6-lobed, 3 upper lobes 5218. Ambrosia psilostachya 5220. Franseria acanthicarpa 5219. er pumila 5221. Franseria confertiflora 150 COMPOSITAE larger than the ety and with the heyy thickened and blackish below; fruit very variable, 1-flowered, 1- oe ed, obovoid to fusifor. mm. long (the beak usually subulate or acicular and 2-3 mm. long), usually rugose, mc and sparsely >”: ihe or essentially sage the pd armed with 6-30 str ongly flatte ned, lan ccolate, 3 acerose-tipped, straight, i s 2-5 m long, rie —— on the same Te t reduced to min — incurved hooks Dry or ist, sandy soil, Arid Transition and “sari pe Ninna sota to prin Berg to Arizona, New Mexico, ae waarn Teng: ‘Washington s = to southern Cal not common in our ran ge . Type locality: “Banks of the Saskatchewan and Red ” Collected by Freeh vine 9 Pe Drummond. June—Dec 2. Franseria confertiflora (DC.) Rydb. Weak-leaved Burweed. Fig. 5221. Ambrosia confertifora DC. Prod. 5: 526. 1836. Franseria tenuifolia Harv. & _ ex A. Gray, Mem, Amer. Acad. II. 4: 80. 1849. 922. Franseria confertifiora Rydb. ag Sl Fl. 28. Franseria — ~— Ry jas loc. cit Franseria ine Rydb. op. cit. 30. Slender perennia al 0.2-1.5 m. high, tages simple pales the inflorescence, the stem strigose or strigillose and often spre eading- hispid or -hirsute. Leaves slender-petioled, the blades 1.5-12 cm. long, ovate in outline, interruptedly ey ts tisinnalined ai ostly linear to lanceolate, acute divisions (those of the lower leaves often broader and ob , the te egments usually use tl rminal elonga ted, all green above, paler beneath, strigose or strigillose and often hirsute, rarely canescent ; staminate racemes slender, usually 8 cm. long or less and panicled at apex of stem, the Pistillate heads at their base and in often dense clusters in the upper axils; staminate heads 2-4 m . thick, about 10-20-flowered, the wide about 6-10-lobed, some of the obes often acute; fru “oboe oid, —4 mm. long, usually 1-flow , 1-beaked, with turbinate or stipitiform base and short. conic beak, glandular and sparsely pilosulose, the body deeply alaengy 4 Fig hag “unig 0-20 uncinate- tipped, usually incurved, flattish-based, aha spines 0.8 m r les Plains, valleys, and waste places; occurring as an raion weed i am phe cop cisco, errs and Maripos Counties, California, and more commonly i in ae California and a r California; Texas rd "Cidatena oath to Mexico. Type locality: Matamoros, Tamaulipas, Mexico. May—Nov 3. Franseria nie Less. Beach-bur. Fig. 5222. Franseria chamissonis Less. Linnaea 6: 507. Franseria cuneifolia Nutt. Trans. Amer. Phil. Soe = bs ai 1840. Gaertnera chamissonis Kuntze, Rev. Gen. Pl. 1: Perennial herb, the stems numerous, up t aH 6 m. long, decumbent from a deep root, stout, mostly simple, pubes cent and often cinereo with pl Re appressed or sometimes spreading hairs. Leaf-blades ovate or oblong to cuneate or vate, cm. long, on petioles about as long, usually obtuse, cuneate at base, coarsely tichate covtate or obscurely lobed, thick, densely silky-strigose on both sides; staminate flowers in a dense Sy be raceme or spike, usually about 8 cm. long, i in t ppe i i d at s thick, about 25-50-flowered, the in wiivere shallowly about 9-toothed ; oa ovoid, 1-flowered, 1-beaked, 7-9 mm. long, glandular and pilose, covered with about 20-30 stout, rigid, danke ina — s 2.5—4 mm. long, these flattened and grooved on the upper pe 5 often with deep axilla “th sands, Humid Transition and Upper Sonoran Zones; Vancouver Island, British Colum! to oer ater yg California, and on the Santa Rastare Islands, Ventura Cou unty. Type locality: C ay mia Mg ya Franseria chamissonis subsp. Melanatiieets Hoge ) beh paps & Stockw Madrofio 4: 120. 1937. (Fran- seria chamissoni Sg bipinnatisecta Less. Lin : 508. 1831; F. bipinn Sede Nutt. Trans. Amer. Phil. Soc. y- 7: 344. 1840; F. lessingit aren & Walp. yo Swale. Nova-Ac ta, Cag 8 Leop. or rol. 19: suppl. 268. 1843; F. bipinnatifida var. dubia Eas' Proc. — Acad. III. 1 F. bipinnatifida var. insularis Reiche, Fi. Chile 4: 80. 1905; F. villogs F Rastw. ex Rydb. N. Amer. Fl. M3: a 1922.) With the habit, pubescence, and the blades 3-7 cm. site often as wide, pinnatifid to bi- or tripinnatifid, the “principal ultimate divisions mostly ova : rm, steading, shen oF slightly curved spines 2-4 mm. athe usually the lower or sometimes all more or ish flat- ened and grooved on the upper surface at “east Bag atg t A ’ toukea. jowth to eae ata er California; t of Chile. Type locality: Santa Barbara Barbara County, California. May—Nov. Hubris with r chamibboine subsp. chamissonis occur. 4. Franseria dumésa A. Gray ex Torr. White Bur-sage. Fig. 5223. pe samen A. Gray ex Torr. in Frem. Second Rep. 316. 1845. is Torr . Smiths. Contr. . 16. 1854. Seessten dumosa Kuntze, Rev. Gen. Pl. 1: 339. 1891. Shrub 2-6 dm . high, hii herked cong Rcth area the branches spinescent, young growth ill 1 densely can escent-stri rigillose. Leaves 3 ncluding the usually short but slender petiole, ong to ovate or del in outline, bi- to eriphinately ivided into usually few, small, mostly ovate or obovate, obtuse divisions, densely can t-strigillose; rac f anicles terminal, usually 5 c ng oO s, the staminate and pistillate flowers often in ixed ; - 5 thick, abo: 30-flowered, the involucre bluntly 6-9-lobed about to middle; fruit ovoid or fusiform, mm. long, 2-flowered, 2- , glandular and sparsely pilose, the beaks slenderly conic or subulate from a thickened base, the bod covered with about 25-40 orn te, rigid, flattened, acerose-tipped, straight spines 2.5-4 mm. long. and plains, Sonoran Zones; southeastern California (Colorado and Mojave Deserts) to southern southern, Arisoes., porieess hover California, and Sonora. Type locality: sandy uplands of the Mojave us SUNFLOWER FAMILY 151 5. Franseria eriocéntra A. Gray. Woolly-fruited Burbush. Fig. 5224. Franseria eriocentra A. Gray, Proc. Amer. nia 7: 355. 1868. Gaertnera eriocentra =i hrrs Rev. Gen. Pl. 1: 18 Branching shrub, 3-9 dm Peas white. bared the gs “i% ao —— ig and spreading-pilose. Leaves ovate, t angular, or oblong, 2-5.5 cm. lon wed i pose petiole or subsessile, sinuate-toot $n, to pinnatifid w nth lac iniate lobes, paittatrimlny tee gre above, densely canescent-tomentulose and sparsely -Phalear along the veins beneath; he ae in short racemes or racemiform panicles toward tips of branches and branchlets, the staminate above staminate heads 4-6 mm. in diameter, about 30-flowered, the inv ase Sem acutely about T0.- lobed i 0 to middle, the corollas pilose above ; fruit 1-flowered, rin ae m, mm. long, stipitate-glandular (especially on the spines) densely long-villous, bearing mostly near middle about 20 or fewer rigid, straigh te, minate, crowde flattish or groove Desert washes and earn, Upper Sonoran Zone; nfo eis pa eb a in eastern San Bernardino County to southern Utah aor > southern Nevada to northwestern Ariz Type : Prov ames Mountains, San Ber nardino County, California. April—July 6. Franseria epacaeage ae San Diego Burbush. Fig. 5225. Franseria chenopodiifolia Benth, Bot. Sulph. 26. Gaertnera chenopodiifolia pr tereet arom te ee ara 6: 461. 1910. Franseria lancifolia Rydb. N. Am 1, 33: Suffrutescent, leafy, slic t 0. 5m eis at length whitish-barked, branching, the branches ee cee and glutino ous, lecoaet Leaves ovate or deltoid-ovate, 23. cm. long, nearly as e, slender-petioled, acute or obtuse, broadly carinicee or round base, crenate-serrate or obecavely lobed, subcoriaceous, tomentulose but greenish above, density whitish- sorbentelons and eriocentr: Sea 5225, Franseria racist 5222. Franseria chamissonis 5224. Franseria 5223. F i 152 COMPOSITAE pitoe yeas beneath ; — simple or branched below, the pistillate heads at their base and clus red in the upper axils; staminate heads about 3-5 mm. thick, 16-30-flowere ~ the in woluere t int eel and Cae 6-10-toothed; fruit turbinate-subglobose, abou mm. lon ered, 2-3-beaked, the body densely lanate-tomentose, thickly covered with numero rigid fe a grooved, mostly uncinate-tipped spines 3-5 mm. long, the lower mostly seeks lecclale and flat- tened, the upper subulate, s apeerets: r Son uthwestern San Diego County, California, and Lower California. Type lodaiting: Y masbalecn Bay, Lower: Chtien April-May. 7. Franseria ambrosioides Cav. Ambrosia-leaved Burbush. Fig. 5226. Franseria brosioides Cav. Ic. 2: 79. 1793. Gaertner brosioides Kuntze, Rev. Gen. Pl. 1: 339. 1891. Suffrutescent perennial, 1-2 m. high, the ascending stems several from the base, leafy, mostly unbranched below the inflorescence, a andinioe. puberulent and grayish-hirsute. Le: ves 6-18 cm. long with petioles 1-4 cm. long. elongate-triangular to Gnckakie acuminate, cordate to broadly co y toot i i i —2 1-2- ed, tie be sa ew vise lon ng, spines many, spreading, slender, slightly flattened, pe nice 5 athe Seek hooked at the Sted ert slopes and sandy washes, Lower Sonoran Zon ear San Diego, San Diego County, California (where it pro gale was introduced), south to Lower California and tine southern Arizona south to Sinaloa and Durango. Type locality: Mexico. Fe .-May 8. Franseria ilicifolia A. Gray. Holly-leaved Burbush. Fig. 5227. veer tierfong te 10 Proc. Amer, eo i fst 1876. Kuntze, Rev. Gen. Pl. 1: 3 staminate racemes = ut 8 cm. long, the pistillate heads at their base and mostly solitary in the upper axils; staminate heads 7-10 mm. wi ah Rs ut 60-flowered, the involucres considerably ex- ceeding the "flow wers, ea about 14 lance-ova r lanceolate, spinescent- shine phyllaries, connate pas their length or less, the sorte Sapliate ~glandular ; fruit fusifo orm 0 gltbtes fasilor , 1-2.3 long, 1-2-beaked, usually 2-flowered, seine stipitate- “glandular, heatiee below the beaks aden rigid, oiekae ShcHeee-tisned ‘flat ish or grooved spines 4-6 mm. long. Desert washes and canyons, Lower Sonoran vie Riverside and Imperial Counties, rnia, to northern a i eee estern Arizona, Type locality : Great Canyon of the antes as riers ec Moun- ains, 25. XANTHIUM L. Sp. Pl. 987. 1753. arse monoecious annuals with branching stems. Leaves alternate, petiolate, netiate to nearly entire, or lobed. Staminate heads borne above the pistillate; in lucres 1 flowered, the phyllaries free in 1-3 series; receptacle cylindric, paleaceous. Pistillate in- i ate i Achenes linear or sometimes ovate, — Me se. [From the Greek word meaning up ee ancient name of some plant producing a dye that color. cally unstable genus of uncertain origin with pie Me about 3 species. Many species have been aie bey = “garviktion of the bur. Type species, Xanthium strumarium L. Spines lacking in the leaf-axils; leaves deltoid-ovate to subtriangulate, long-petioled. 1. X. strumarium. Conspicuous 3-forked spines at axils of leaves; leaves lanceolate, short-petioled. 2. X. spinosum. 1. Xanthium strumarium L. Cocklebur. Fig. 5228. Xanthium strumarium L. Sp. Pl. 987. 1753. Erect, ae Sed branching annual 2.5-8 dm. high with stout, striate, scabridulous stems. Leaf- blades 6-12 cm ong and usually equally as wide, the petioles as og a or longer, deltoid-ovate or - so : m margin dentate or serrate, scabrid above and below;; pistillate pectneces crowded, 1-3 in leaf-axils or on gabe par aed: sa burs about 1-1.5c¢ m. Tong, ellipsoid or fusiform, as ale brown to green- e puberulence, the pric ie ee up to 2 mm. gre straight except for the hooked ope slabrous os and puberulent and glandular at be the beaks stouter, about the length of the prickles, straight o: acura 4 ena areas; Soa in the Pacific States only from the Colorado Desert, California (Cameron Lake, Bran- degee; Fort Yuma, Pe arish), but widespread in the American tropics and in Europe. Type locality: not known. SUNFLOWER FAMILY 153 Many forms of this cosmopolitan weed are to be found and innumerable races have developed locally which have received specific ee ed the — sa involved he jog 7 hirano Rep. Spec. Baniend Magnan 20: 1-221. 1923. The synonyms listed with the following taxa are those more frequently found in Xan thins bias Hira var. pikes? énse (Mill. ) -Forr; ri pe ; piers 2: 294. 1838. ‘Kanthium cana- dense Mill. Gard. Dict. = iB 1768; X, italicum Moretti, Senha tet Giorn fis. Chim. Dec. II. 5: 1822; X. saccharatum Wallr. ot. 12: 238. 1844; X. oviforme Wallr. op ag 240; X. campestre PPodilas Pittonia 4: 99 Ae fc aro Tocsee op. cit. 63.) Burs usually 2-3(3. S) ¢ long, variable in shape and density 4 prickl es, the beaks exceeding the prickles i in —_— the prickles more ro less densely hirsute toward the base well as s glandular. ~ cosmopolitan weed g in moist valleys and flood plains throughout the Pacific States Ty pe sala not know Xan m strumarium var. glabratum (DC.) nquist, Rhodora 47: 403. 1945. eee p pepo ct B glabr =~ DC. fod: 3% 523. 1836; X. Pennsyloanicum Wallr. Beitr. Bot. 17: 236. 1844; ? X. se 1gc e weootont Cockerel Proc. Biol. Soc. Wash. a a ; X. calvum Millsp. & — Field ae Bot. Sie pl. 9.) Burs usually 2-3 s shorter, variable in shape and density of Bo i Po ostly lacking the hirsute hairs lon ke exceeding the lene sam in Bet cay ‘the gk Ty eee clas toward the base and m of X. strumarium var. — ense. A cosmopolitan weed growing in moist valleys aaa flood plains throughout the Pacific States but more common from central California south. Type locality: ‘‘in Carolina.’ 2. Xanthium spinédsum L. Spiny Clotbur. Fig. 5229. Xanthium spinosum L. Sp. Pl. 987. 1753. erect, branching annual up to 1 m. high, the leaf-axils armed with paired, papecct, ent afy si spines 3 cm. long or less, the stem whitish, strigillose or erectish pubescent. Leav ostly la henolatd or per ney we 5-12 cm lone including petiole, acuminate, cuneate at base, us zally acne fers few-lobed toward base, green above, ly canescent-strigillose beneath ; pistillate in- se volucres solitary or cnered in the axils and on pees lateral branches but not crowded, in fruit 5226. ria ambrosioides strumarium Franseria 5227. Franseria ilicifolia 5228. Xanthium 5229. Xanthium sp 154 COMPOSITAE the burs yellowish green or brownish white, oblong, about 10-14 mm. long, 6-9 mm. thick (includ- ing prickles), usually somewhat compressed, ribbed, Dyrweed pilosulose geo neg ve ah or at apex bearing a single slender straight beak 2 m s or ual beaks, the body hig red densely armed with eo re whiner begs ‘(often ‘Mackisle Sased}, ong. eq sar Jaleo 2-3 m plac ually pasture lands and around habitations. A cosmopolitan weed rather common on the Pacific Slop . aa pte throughout the United States; a native of South America but originally described from Portugal. June-Nov. Spanish Thistle. ubtribe MADIINAE* Ray-achenes obcompressed, each hidden within the obcompressed phyllary, the abruptly infolded lateral margins of which more or less overlap. Disk-flowers 6 or more. Achenes spate te and ch ansaid pappus ablong, obtuse. 26. Achyrachaena. chenes Ss not costate or s pappus when t isk- nino serie, a. heads choalig during midday. Pappus n annuals, 27. Lagophylla. Pap sab perennials. 28. H Sunt Disk- eit frie. ie open continuously. 29. Layi . Madia. Ray- Paphipe not shecmn essed or each hidden by the sp lateral margins of the Padi pliyttiry. Style of disk-flowers glabrous below the subulate branches. bias -achenes not ere ge ribbed; ray-achenes amit labrous and epap Ra dig sro oothed or -lobed, the lobes subparallel; sr without i ached glands; leaves not ag esi Pi — — Baccuieons usually laterally compressed and finely longitudinally striated, — en en- folded by the og sulcate subtending phyllary; basal leaves subentire Ray-achenes broader, not laterally compressed or striated, each only partially pees by the subtending aeriesy: basal leaves pinnatifid or toothed, rar ears uben ev ‘Hew Upper leaves’ a phyllaries aimee bye cen pit glands; ligules yellow;seach disk- flower subte’ pappus 32. Holocarpha nded by a bract Ray-ligules 3-cleft or -parted into citadel ‘ecb lobes; tack-shaped re present Ftoridang C. tenella); th narrowly linear and grast-it ike. 33. Calycadenia. Disk-achenes prominently ribbed; ray E Ppappose; papp 34, Blepharizonia, Style of disk-flowers hairy below the 2-cleft tip; pappus plumose. 35. Blepharipappus. 26. ACHYRACHAENA Schauer, Del. Sem. Hort. Vratisl. 3. 1837. ernal mesophytic annuals. Leaves opposite and clasping below, alternate above, linear. pn solitary, long-peduncled, terminating the stem and few ascending branches, heter- ccondaie flowers. Ray s l-seriate; ligules yellow, turning crimson - ceptacular bracts in a single tween ray isk, free. Achenes 10-costate, tubercu- late-scabrous ; pappus of ., scales, spreading on ripe achenes to form . "ppg ead. [Na reek, meaning —_ — in reference to the chaffy pappu 1. Achyrachaena mollis Schauer. Blow-wives. Fig. 5230. Achyrachaena mollis Schauer, Del. Sem. Hort. vehcisaee 3. 1837. Lepidostephanus le Bat Ind. Sem rt. Gotting. 1837, ex Linnaea 12: Litt. 82. 1838. Stem simple or few-branched, 1-4 pte . high, — — — Seca pi. moderately glandu- lar above. Leaves entire or remotely serrulate, o 13 cm. lon, . wide, mostly less ; heads in flow _ gt cm. cs ss fru he Ba nga >: aes 3 cm. across; phyllaries completely i investing the achen y- ? aid s about 8, the ligules senior sh 6 k-flowers _ 15-35; hence: clavate Backs about 5m m. long, those ry the ray e ib e of the di sk pappose and scabrous with yr a Rg 5 pus - 10 ‘abl ong Rinne scales, the 5" eater half as long as and alternate with the inner, he es in “4 ice uent in moist grassy fields with heavy soils, Sonor: es hea tion gine cismontane valleys from Douglas Pemaet Oregon, south to northern aie California. ; rn locality: California. April-May. i » o 8 ' = aid 27. LAGOPHYLLA Nutt. Trans. Amer. Phil. Soc. IT. 7: 390. 1841. Me esophytic or xerophytic annuals. Basal leaves serrate-dentate to subentire; cauline leaves entire, readily caducous. Involucre ee to hemispheric, the phyllaries com- pletely en nfolding the obcompressed r: ray-achenes and caducous with them. Flowers yellow, ~ opening toward evening and a in the morning. Receptacle penicillate- t centrally, its bracts in a single row between ray and disk, slightly united. Ray- Risvets 5, fertile. Disk-flowers 6, sterile. Pappus none. [Name from two Greek words * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 155 meaning hare and cat in allusion to the copious sericeous pubescence of the upper leaves of the original spec es. ] A genus of 5 species, natives of the western United States. Type species, Lagophylla ramosissima Nutt. Ligules 8-13 mm. long, conspicuous, pecan yell Ste sn abilieaniiniaas branched; involucre ucatiaiiniics chiefly Coast Ranges ay-achene esti ly cblancealate, glossy, smooth, midnerve evident; shytlaries — ayn A Sea, slender, flexuous Pig yar ‘ yilbs:0y Ray as ie aol _ mid-nerve obscure; phyllaries barbellate-ciliate; talier and stricter habit; Bat ont rat iagd val. . L. dichotom Stem rary or -patieuldnely elie bork involucre turbinate; late vernal to autumnal; Sierra Nevada a, Ligules 3—5.5 long, inconspicuous, pale lemon- Ts lt 1 we Heads eitineas or at least not densely glomer 4. L. ramosissima, Heads densely congested in compact a Soe CO bik 5231 5233 5230 yrachaena — 5232. Lagophylla dichotoma 5231. Ay tele lla min 5233. Lagophylla glandulosa 156 COMPOSITAE 1. Lagophylla minor (Keck) Keck. Lesser Hareleaf. Fig. 5231. Lagophylia dichotoma subsp. minor Keck, Madrofio 3: 16, 1935. Lagophylla minor Keck, Aliso 4: 105. 1958. Slender herb 1-3 dm. high, the ie Policing ee at least in upper half, to form an open corymbose crown, the ultimate twig y dark, strigose. Leaves linear, some tapering to bas up to 5.5 cm. long and 3 mm. wide, eg except . the basal, narrowly involute, soon deciduous strigulose-puber te st becoming hirsute-ciliate toward the heads ; inflorescence bearing few or no stalked glands, if present, not extending down the peduncles; involucre 4-5 mm. high, pilose, vi phyllaries long-ciliate on the angles with “wai almost as long as bract is wide, acuminate; ra — smooth and eeny bial a 81.3 mm. wide. poi aig slopes of the foothills, Upper Sonoran Zone; Inner North Coast Range from Glenn Cou unty o Nap: unty and on the bala cosh of reine County, California. Type locality: Pope Creek, just ae OE wane Spring, Napa County. 1—-May 2. Lagophylla dichotoma Benth. Forking Hareleaf. Fig. 5232. Lagophylia dichotoma Benth. Pl. Hartw. 317. 1849. touter than L. minor, 1.5-6 dm. high. Inflorescence bearing few to -many evident stalked glands, these extending down the peduncles involucre 4.5-5.8 mm. high, hispid-hirsute, the hairs much shorter than the width ‘of the acu e bracts ray-achenes blackish or ehtly al a gens with cellular surface, with 20-30 striae on op ig e, 1.4-1.9 mm. wide. Rather rare, in grassy places in the valleys and sea Upper ch n Zone; east side of the Central Valley from Butte County to Tulare Bose and in the Inner South Coast Ratige | from San Benito County of gaa County, California. Type locality: Sacramento Valley (probably in what is now Butte Coun ty). April 3. Lagophylla ei aduttad A. Gray. Glandular Hareleaf. Fig. 5233. Lagophylla glandulosa A. Gray, Proc. Amer. Acad. 17: 219, 1882. Stem 1-10 or 15 dm. high, simple or paniculately branched, the plant slender or bushy with neven crown; herbage danecis apprese alae and distinctly glandular. Lower leaves ete oblincselile to seid i lost before flow ing, 3-12 cm. long, 5-12 mm. wide, parities leaves bear- ing fascicles or sho st erile shoots in their axils, i lg aducous, reduc ed to bracts upward heads on short peduncles ; involucre turbinate, 5-7 mm. hi gh; ligules 8-13 m m. long, 6-9 mm. wide, pia sie lateral ay ounded, much wider ean the ai one; sie-aghnnek leat cai Hot dry slopes, Pisces nt sea Zone; common in the pee ba eaten from Shasta County to Kern County and occasional in the upper sa ma pt Valley and in Mendocino County, California. Type locality: “California, a ae Sierra Nevada from Auburn to near the Y aaa a ” July et, ophylla clainhitans subsp. ei (Greene) Keck, Aliso 4: 105. 1958. (Lagophylla serrata Greene, Bull. Lacon! Acad. 1: 280. 1886.) Inconspicuous herb 1-6 dm. high, simple or dg had ascending branches, almost glandless below the crown; leaves ae fascicles or sterile shoots in their axil e lower often persistent after time of pak perp —_ few, bes short elongated, very slender peduncles; cell ade’ 4.75-6 mm. high; Hod mE a2 Tay —3.2 mm. long. A seasonal palin growing with the species. Type locality : Grass Valley, Nevada ow ay— 4. Lagophylla ramosissima Nutt. Common Hareleaf. Fig. 5234. Lagophyile ramosissima Nutt . Trans, Amer. Phil. Soc. II. 7: 391. 1841. ell. iii Calif. Acad. 5: 1873. Legophylla hillmani A. Nels. Proc. Biol. Soc. Wash. 17: 98. 1904. Stiffly erect, 2-10(-15) dm. high, racemosely or paniculately branched above, sometimes from the , rarely simple ; herbage grayish or dull green, densely ede siege or white-sericeous, the grease: ager lice. stipitate glands confined to upper leaves and heads. Leaves as in L. glan du- losa; heads short-peduncled - subsessile, racemosely disposed along ‘the branchlets and in. small pets at _— ends ; involucre 4.4-6.7 mm. high, the lanceolate abe densely villous-ciliate the angles ; ray-achen ee eiavaie wad slightly arcuate, 2. m. lon bun pee: in open stacks in hard dry soils, Sonoran and ice eth e Sentai: Washington and western — south to San Diego County, Saliforsie, and aectherh and western Nevada. Type locality: ‘In the prairies r Walla-Walla, in Oregon’’ (possibly in what is now Washin ngton). May—Oct. 5. Lagophylla congésta Greene. Rabbit-foot. Fig. 5235. eo — Greene, Bull. Torrey Club 10: 87. Lagophylla ramosissima var. congesta Jepson, Fl. W. Mid. Calif. 539. 1901. Lagophylla ramosissima subsp. — Keck, Madrofio 3: 16. 1935. Similar to L. ramosissima except more robust, the heads densely congested in compact glo- merules 1.5-6 cm. — the slomerules intone gs terminal or in interrupted spikes or more rarely on short _— branches racemosely ged, the steens usually simple ; involucre 5-7 .5 mm. high. Hot, open slopes with L. ramosissima, wala Sonoran Zone; from Humboldt County to S soe Cruz ot and i tea Eldorado County to Stanislaus County, California. Type locality : 2 pie ont County. May- 28. HOLOZONIA Greene, Bull. Torrey Club 9: 122, 145. 1882. Perennial herb, hemicryptophyte, with elongated fleshy rootstocks. Stems erect, usually SUNFLOWER FAMILY 157 simple and densely seep below, aad branched at the nearly leafless apex, frequently ort leafy n ea s e paleaceous bristles nearly as long as the corolla. [From the Greek words for whole and girdle, from the wholly Saag ray-achenes. f Calif 1. Holozonia filipes (Hook. & Arn.) Greene. Holozonia. Fig. 5236. Hemizonia filipes Hook. & Arn. Bot. spre a Dias Lagophylia filipes A. Gray, Pacif. R. Re Holozonta filipes Greene, Bull. Sikes Ath ate fae 1882, St 0 dm. high, ee the mature herbage cinereous and villous, beset with f pany lea toward the inflorescen ower leaves ere es beagcat to subentire, 3-10 cm. long, 4-8 m ide, progressively smaller upward; involucre 3.5- igh, as broad, the linear-lanceolate pa folded) phyllaries eaniaenity hirsute ; ray-flowers 43, ae ligule 3.5-4.5 mm. long, deeply 3-cleft into linear lobes ; disk-flowers ( 28). In dry alkaline rocky Iches or beds of intermittent streams, Sonoran and Transition Zones; saan County to Waripon Couey Sha from Napa Pun nty to Marin, Santa Clar ra, and Monterey Counties, Californ Type locality: California. June—Oct 5236. Holozonia filipes 5234. Lagophylla ramosissima cae 5237. Layia acenicteniachis 5235. Lagophylla congesta 158 COMPOSITAE 29. LAYIA a & Arn. Bot. Beechey 148. 1833, nomen 1 DG. rod. 7: 294. 1838, nomen conservandum Vernal annuals with chiefly aaa ae ee or toothed to pinnatifid, narrow leaves. Heads many-flavored, on usually ed terminal peduncles, hetero S f 15 species, all of hich occur in, and 13 of which are restricted to, California. Type species, Tridax 0) "pilantiolins Hook. & Arn Bracts of receptacle subtending each disk-flower; involucre (and herbage) not glandular, the phyllaries with promi- nent p ee cr8y processes at base of hairs. Pappus ote subulate awns; disk- ni a asocely: org sip cuneate; shgllesieg hispid-ciliate or pec- ti wi ioe the folded edge, otherwise nearly ; Coast Ranges . L. chrysanthemoides. Raney of Boing tg thin paleae; disk- RCE, enna clavate; shots uniformly pepaiate: hispid reat Bracts of harsagate’ — toa dy g between ray- and disk-flowers; involucre (and er pt om the phyllaries pubescent but not a ps ve. — paleaceos, slabrons sheet for capillary hairs radiating from very base y-flow in 1 series, their ligules obovate to flabelliform, tein chines dull, glabrous or pubesc dntiesed at — ventrally; involucre hemispheric, green, the bracts not inflated, with etn tips; pappus 2-3.5 mm. long. Ligules white; ps vaio ray-achenes sericeous; leaves mostly entire. 3. L. leucopappa. ae Sarma with long white tips; anthers black; ray-achenes 2 gals or lightly sueeerent toward ea m lobed. Ree Sag Ray see ok in 2 series, their ligules oblong-spatulate, their achenes polished, pleas plump; —— broadly ur rceolate, ee Slotted, the bracts inflated, with inconspicuous ti ipsi pappus 1-2 mm. long 34 . fone. Pappus setaceous or if palaceous, plumose Ligule sited with white tip avely yellow throughout) ; pappus commonly merely scabrid, if plumose, Fs o with inner wool, of 18-30 bristles; anthers black; tips of puiinrieg. longer tha than a basal portion. Ligule Bas od or white (none in L. discoidea) but not as above (except sometimes in ra “paillardioides) : . ppus plumose, with inner woolly hairs only in L, septontrionalis and L. glandulosa; tips of phyl- tare: pk than the basal portion (except in L. heterotricha Pappus persistent; “ssigg not prominently fistulous, pubescent Kalan (glabrate in L. carnosa); disk- rigos achenes stri Ligules more >i 5 mm. “ed conspicuous (none in L. discoidea) ; self-sterile. Anthers yellow; involucre not urceolate; stems not dark-dott ed: p pappus _— Involucre ovoid, 42 mm. high; ligules golden-yellow; disk-coro —8 mm. lon ray-achenes 3.8-5.2 mm. lo ong; pappus 3.5— ru 2 mm. sy ie ‘slender Ritoeies 6-21, densely plumose, with inner wool; Lake and wea =: ounties. . L. septentr Involucre turbinate or campanulate to hemispheric, 5-9.5 mm. high; oe sham se 3-6 ong. Heads discoid; pappus paleaceous, 1-1.5 mm. long, ae ay San Benito County. . L. discoidea. Heads inane pappus setaceous, 2-5 mm. long, white. Stems hispid; basal — — or lobed, the cauline entire; ray-achenes 3-4 mm. long; 3-5 mm. long, the bristles 10(— 12), flattened, lin: ceeatercess, icant plumose, usually with i Poca: 2id wool, may 38 glandulosa. Stems pilose or ce tong basal leaves pinnatifid to oe tifid; ray-achenes 2.5-4 mm. long; pappus 2—4 mm. long, a — S pied (rarely cons, filiform, Seger plumose, without wool. Anthers black; involucre urceolate; stems dark-dotted; pappus ores 11. L. gaillardioides. Ligules 2-4 mm. long, inconspicuous; self-fertile; pappus rufous to whitish; anthers black. Ligules yellow; pappus-bristles 11-15; stem rigidly erect, 20-100 cm. tall, dark-dotted; ntly odorous; ray-flowers 8-16. wer leaves laciniate-dentate, broadly oblanceolate or oblong; —- mostly stout; San Francisco Bay to Santa Cruz 12. L. hieracioides. Lower leaves pinnatifid, if merely déstate ee ty narrower; stems mostly slender; Mount Diablo to Santa Barbara 3. L. pan piclata Ligules white; Based scale ad fli fon — rel ip sony Sinaehed. to 7 as aah gone without dots; not Pappus deciduous; oe fistulous, erent: below; POO es sericeous; teas creamy white to pale alien 15. L. heterotricha. 1. Layia chrysanthemoides (DC.) A. Gray. Smooth Layia. Fig. 5237. Oxyura ssioanrsinaioe, 8 -. — 5: 693. 1836. Hartmannia ciliata DC. op. ci Callighicte doviglan Sat tare: Bot. Beechey Suppl. 356. 1838. SUNFLOWER FAMILY 159 peel douglasii Torr. & Gray, Fl. N. Amer. 2: 396. Layia chrysanthemoides A. als Layia calliglossa var. ss ee A. Gray, Bot. ait, 1: 370. 1876. Blepharipappus douglasit var. alidechadte Greene, Man. Bay x .201. 1894, Ste mbosely branched, 1-4 dm. high. Leaves ee se 8 otherwise smooth, the basal prmak Saree a lin ear or cian blunt Shes, sai: e 6-11 mm. high; ligules yellow with white lobes, 7-15 mm. long, Bd wide; disk- flow. rs 30-100, the corolla 3-5 —_ — ray-achenes 2.7-3.9 see, long, st ron esp compr: res sed, sia nae e: peppy 2.54 m densely strigose, sometimes lacking pubescence and pappus ; pappus 9-17 tawny, old nae scabrous awns, t "s sete ones 1.5-3.5 mm. long, the Jekerveditet pba usually shorter or even reduced to rudim Moist heavy rat vn valley floors, +a per ee pay Zone; Mendocino and Glenn Counties to Monterey County, California. Type locality: Fine leagg 2 aba Layia chrysanthemoides su tima Ke ti Aliso 4: 108. 19 rostrate, even the central sudan horizontal; leaves saceuent or ssc with broad rounded Pe ig ‘ion. “of phyllaries ag rounded; late-flowering. Rare; on the oe coast of Mendocino and Sonoma Counties, California. Type ity: sou uth of Jenner, Sonoma County. 2. Layia fremontii (Torr. & Gray) A. Gray. Fremont’s Layia. Fig. 5238. Daemons ip ee, ih bf oe Bost. oe Nat. Hist. 5: 110. 1845. Layia frem A. Gra r, Acad. II. 4: 103. 1849 pdb teetin Potosi ro "Pittonia od 246. 1892. Stem 1-4 dm. high. Leaves scabrous or short-hispid, not re -_ viscid, ny basal pinnately parted; involucre 6-11 mm. high; ligules yellow, the outer half white, 9-18 mm. long, 7-12 mm. wide; disk-flowers 40-100, the corolla 3.6-5.3 mm. by Foams ray-achen s 2.5-3 1 mm. long, ical com- pressed, glabrous ; disk-achenes 2-3.5 mm. long, densely s serixeee and with a row of long capillary hairs on the areola ; pappus of 10 tawny, gio ovate-lanceolate paleae with attenuate tips, 2-4.5 mm. = Gra s, Upper Sonoran Zone; Great Valley ker apes om, Tehama County to Tulare County, California. soon locality : California. Collected by Frem rch—May 3. Layia leucopappa Keck. Comanche Layia. Fig. 5239. Layia leucopappa Keck, Madrofio 3: 17. 1935. Stem branched from base upwa ie to 5 dm. high; herbage more or less ni upper half; her bage rather anthocyanous. Leaves not very crowded, villous, the upper on SUNFLOWER FAMILY 169 what glandular, psa up to 7 cm. long and 7 mm. wide; heads paniculate to at te not con ven involuc e globose, wipers 6-7 mm. high, the phyllaries dilated but usually entirely en- closing the ro san ‘thickly beset with stout, gland-tipped hairs and viscid-puberulent ; bracts of the receptacle united only toward re ciliate ; ray-flowers 3-7 ; disk-flowers 3-6; ray- and disk-achenes Grassy slopes at low elevations, Upper Sonoran and Transition Zones; North Coast Ranges from Humboldt County to Mount Diablo, Joma Costa County, California; also Sacramento County. Type locality: probably Sapeainéate County. May. Jun 11. Madia subspicata Keck. Slender Tarweed. Fig. 5262. Madia saben Keck, Carnegie Inst. Wash. Pub. No. 564: 45. 1945. trict, slender, usually simple, or sparingly short-branching above, 1-1.5 dm. high, t herbage yelow -green , pilos e and viscid-puberulent with pices mae! stipitate glands. Leaves dihear; . long, to 3 mm. wide; heads ‘subspicat on very short ped pete overtopped by the sub- ‘esses feaven arheicte ovate, 6-7 mm. high; disk-flowers 5-15, the anthers black; ray- pret el about 3 mm. long, ane ee purple spotted, the disk-achenes similar. Gra slopes, Upper Son Sierra foothills from Butte County to Mariposa County, California. Type entity near Knights Perry. Me saa Couity. May-Jun 12. Madia gracilis (Smith) Keck. Gum-weed or Slender Tarweed. Fig. 5263. Sclerocarpus gracilis Smith in Rees, Cycl. 31 Madorella dissitifiora Nutt. bce ans. Amer, ay — Il. 7: 387. 1841. Madorell Nutt. loc. c Rs Wy 5262. Madia subspicata 5263. Madia gracilis 5260. Madia citriodora 5261. Madia anomala 170 COMPOSITAE pr serbemiiagee Cine & olay FI. a Amer. 2: 405. 1843. Madia r . & Gray, loc Madia sass var. Paddy Gray eigen Amer. Acad. 9: 189. 1874. a A. Gra Madi tiva race Lagophylla hilimani A. ee Proc. i Se: Wash. 17: 98. 1904. Madia gracilis Keck, Madrofio 5: 169. 19 Madia sativa subsp. eserchore Keck, tay: 4.1935, Stem usu ually oon simple or flexuously branching from the aon the branches not over- i-10 . hi h topping the main dm. high; herbage not as viscid as M. va, resinously fragrant. Leaves very ae gee mostly linear, sessile by a o 10 cm. long and paniculate to racemose, not congested, y bracts rarely p volucre ovoid to depressed-globose, 6-9 mm. hi phyllaries thickly beset with stout, appa sa irs, the acuminate tips short ; ray-flowers 8-12; ligules 3-8 mm. lon ng ; disk-flowers 15-35; anthers included, black; ray-achenes 2.8-5 mm. long, gibbously obovate, often mottled; di sk- a similar but straighter Open or wooded areas, poeekinice in disturbed soil, Sonoran Zones to Canadian Zone; British Columbia to eng California east to Montana and Utah; Chile and adjacent Argentina. Type lo cality: west coast of North America. April—Aug. Madia gracilis subsp. collina Keck, Aliso 4: 108. 1958. More Pith usually branching near the top, the heads spicate or glomerulate, the inflorescence abe vis Sid, the stem not g andular below; sy ha‘ pa bay eo ps ap oe involucres 8-10 mm. high, the — ies with elon mgated tips. oe ‘Sonoran Fenn, Sie oothills f Amador and Roni Counties, Califo Type locality: Vallecito, Calaveras County. yy Rio i Madia g lis subsp. pthace Keck. ‘Aliso’ 4: 108. 1958. Like the species but edeckie pre Mista ic especially on so nl and sgn rather sparsely glandular but usually so well down toward the base; leaves few, large, the upper ones rather Peeadi heads solitary or seccmone on elongated peduncles, iy ple the involucres depressed-globose, contracted are ied achenes, the phyllaries somewhat dilated, pie holding the achenes, th erect tips elongated. Humid Transition Zone; Humboldt gins California. Type cality: Buck ‘Pountai ¥ Ss, 13. Madia citrigracilis Keck. Shasta Tarweed. Fig. 5264. Madia citrigracilis Keck, Carnegie Inst. Wash. Pub. No. 564: 44. 1945, Stem sparsely branched ragite Ae the branches strict, slender, often exceeding the main stem, 2.5-5 dm. high, hispid-hirsute below, villous and v viscid-puberulent above with ae eee stiitat glands. ‘Leaves lin soured ice often Scholes. ale the lower 4-8 cm. to g, wide; heads racemose or terminating leafy peduncles ; involucre dtipvate: 6-8 mm. high; pylaries some- what hirsute and densely stipitate-glandular, villous-ciliate over the face of the achene: ; ray-flow ers 5(-14) ; disk-flowers 3-10(-30) ; ray-achenes about 4 mm. long, broadly lanceolate in cross ection. Forest openings, Arid ig ee sie Modoc, Shasta, and Lassen Counties, California. Type locality: Burney Spring, Shasta County. July— 14. Madia sativa Molina. Chile Tarweed or Coast Tarweed. Fig. 5265. Madia sativa Molina, Sagg. Chile ed. 1. 136. 1782. Stem usually st - often rigidly branched above, glandular well down toon? pats 5-10(-20) dm. past Py ace strongly odorous. Leaves ies cr a sessile by a bro eo p to a8 cm long wake. "i ads paniculate, racemo subspicate, often fox along the hace tie fe foliose: bracted ; ray-achenes ‘ae oblancgeiaie’ disk-achenes ue. the sides sometimes 1-nerved. Grasslands, roadsides, and waste places, Transition and Upper Sonoran Zones; behaving as a ruderal; along 8 coast from southern Alaska to Los Angeles County, California; Chile; Argentina. ype locality: Chile. 15. Madia capitata Nutt. Headland Tarweed. Fig. 5266. Madia capitata — _ Amer. Phil. Soc. II. 7: 386. 1841. Madia sativa var. congesta Torr. & Gray, Fl. N. Amer. 2: 404. 184. Madia sativa hes. ad Piper, Contr. U.S. Nat. Herb. 11: 576. ‘1906. Stem stout, simple or rigidly gickiolroga$ pranches above, glandular to base, gore viscid than M. sativa, 3-6 dm. igh; herbage strongly odor. Leaves sessile by a broad heads in con- gested spikes or in tetas! or fohsar slomerules, often with foliose bracts pepe the heads; ray-achenes narrower and longer than in MV. sativa, the sides a rved. Low fields and coastal headlands, Transition and pes Sonoran Zon times weedy; British Columbia south near the coast to Santa Barbara Cou Ca — Type locality: "adpaton Island, at the outlet of the Wahlamet.”’ May—Oct. but usually aisles rh M. sati 16. Madia glomerata Hook. Mountain Tarweed. Fig. 5267. Madia glom a Hook. Fl. Bor. Amer. 2: 24. Amida gracilis Natt sree a Phil. Soc. sg 7 "390. 1841. Amida hirsuta Nut wae — Piper, Bal Socrey Club 29: 222. 1902 Madia glomerata var. sa Jepson, Man. FI. Pl. Calif, 1098. 1925. Pie io. = ue ae mple or with assurgent or seine branches, 1.5-8 dm. high, vil- lous to hispid, wi pe ida nena glands above; herbage strongly and unpleasantly odorous. Leaves uaa linear, often with feat in theiz axils, the yidar- 3-9 cm. long, 2-7 mm. wide; SUNFLOWER FAMILY 171 pit cf in dense terminal glomerules of 5-30, or in more open cymes and panicles ; Neal narrowly void, 5.5-9 m pete ie ligules inconspicuous, eohanar yellow to purplish, 1.5-2.5 mm. long ; disk- Hower 1 10; aphed s 4-6 mm. long, 5-ner ved. Grassy letest: -openings, vise any aoe age Zones; southern Alaska south to the Sierra Nevada ard San Berna a o Mountains, California , and sar igh the Rocky ‘Mountains to South Dakota and New Mexico. Ty ype locality : sislan of the Saskatchewan. July 17. Madia exigua orni A. Gray. Small Tarweed. Fig. 5268. Sclerocarpus exigua Smith in Rees, Cycl. 3 Harpaecarpus madarioides Nutt. Trans. pa Phil, Soc. J1.. 7 pra 1841. Harpaecarpus exiguus A, Gray in Torr. Bot eg Bound. 101. L A Madia filipes var. pnbines os Suksd. Deuts. Bot. Monatss. 18: 97, 1900. Harpaecarpus exiguus var crocephalus Suksd. loc, cit adia exigua subsp. macrocephala Piper cae U.S. Nat. Herb. 11: 576. 1906 Harpaecarpus th ane ane Bull. . Bot. Fr. 65: 43. Harpaecarpus longipes Gandoger, loc. cit. Harpaecarpus gett» At loc. cit. Stem simple, corymbosely branched above or paniculately branched throughout, very slender, 0.5-3(-4) dm. high, en necuaiaete stipitate-glandular above; herbage aromatic. Leaves . Madia — 5266. capitata 65. Madia 5267. a glomerata v2 COMPOSITAE strigose, linear, 1-4 cm. long, 2 m r less wide; heads on filiform, pag rie often elongated peduncles in corymbose pa ag ice: pease ace peers globose, 2.5-4.8 mm. high; phyllaries early- deciduous with the ma e fruit, ‘linear do ara pair in outline Rely covere a wit prominent glands on oe often a stalks ; ; ray-flowers 5-8, the ligule 1 mm. long; ray achenes 1.8-2.8 mm. long, crescentic, stoutly beaked ; ‘di sk- hens fertile Dry apa and woodlands, Sonoran and Transition Zones; southern British Columbia and western ae tana to northern Lower California and a Ni Mecatn. Type spe Ee west coast of North America. May-—Jul 18. Madia minima (A. Gray) Keck. Hemizonella. Fig. 5269. Hemi ima A. Gra apieige hap Acad. 6: 548. 1865. Hemizonia parvula A. rein op. c scrcereeaes durandi A. Gray, lo A. Gra ic. et ay, op. cit. 9: 189. 1874. Hemizonella parvula A. Gray, loc S Harpaecarpus minimus Greene, op. cit. Hemiz peso minima var. sophia ula H. M. H all, Univ. Calif. Pub. Bot. 3: 148. 1907. Melam ghana ina West. Bot. No. 15: 156. 1929, M. iiieecalon — - E. Madia minima k, Madrofio re ty een Stems 1 to pier? from near the base, ae pei branched to form often a hemispheric plant, villous bdo, glandular-puberulent above, 2-15 cm. hi mg? Leaves often in little clusters at the nodes, otherwise sm: peeing: linear- a mig x ong ; heads solitary or in small terminai glomerules, napiform, 2-3 m hi gh; Shiai loosely appressed, becoming arcuate with the ripening achenes, rounded on ere far their glands tiny, on prominent pustulate processes; ray- nr incurved, m r less beaked. Gravelly slo; get os sont in coniferous color aig Transition and Canadian Zones; British Columbia and northern Idaho south to Diego County, California. Type locality: “pee pt near Soda Springs, alt. 8,680 feet” (Tuolumne Meadows, Yosemite National Park). May-July. 31. HEMIZONIA DC. Prod. 5: 692. 1836. Annual or perennial herbs or shrubs, poe ag glandular and aromatic, mostly fall-flowering xerophytes, very ioe S ring-flower . Basal leaves variously lobed, rarel subentire; upper leaves and bracts not A y open pit . Phyllaries half- enclosing the ray-achenes. Receptacular bracts in a single r more 0 (except in Section Hemizonia), triquetrous, the odd angle posterior, epappose, fertile. Disk-achenes usually bearing a paleaceous pappus. [Name Greek, hemi, half, and zone, girdle, the phyllaries but half-enclosing the ray-achenes. A genus of 31 species, essentially confined to California and northern Lower California. Type species, Hemi- zonia sbasste a DC. Shrubs; leaves small, crowded (Section Z onawnra). Flocs present in axils of older leaves; phy keeled; anthers black; a 1. H. clementina. Flocs absent; tees strongly keeled; anthers yellow; Santa Susana Mounta 2. H. minthornii. Herbs; leaves lar; Ray- achenes vabviously beaked; ligules yellow; inner receptacular bracts when present not adnate nor deli- quescen asa ok a —— fis se receptacular bracts confined to a row surrounding the outer disk-flowers and ited in cup; pappus of quadrate or oblong paleae (Section Deinandra). Diskachene sterile. laries not keeled; — yellow (except in H. corymbosa). Heads not glome Ray-flowers 5 ek. flowers 6; Inner Coast Ranges to southern areoras, . H. kelloggii. Ray-flowers 8 or more; disk-flowers 10 or Leav sic 2 eam ‘thro ughout; stems ae cheeky fistulous. Sor flowers 8-12; disk-flowers ot herbage pallid: Pappu obvious; bic ool = s lobed; herbage vlog “ielow liqules pale ye ; cism i Pappus none or pase oA radical leaves entire or obscurely toot herbage hispid-hirsute below; ligules deep ag 6 transmontane. Ray-flowers 18-32; disk-flowers —< herbage bright ae eae, re pinnatifid or ’ bipinnatifid: coastal. ee «se Leaves vy ciliolate, mostly entire; gives strongly fstlous pa pps Yulee one; Inner Coast Range. . H. hallia Heads aketake small; ray-flowers 5; disk-flowers 6; Mojave Desert. login — sen vasa Coast Rang Ray-fl isk-flowers 3; herbage pete and gray-green; plants low and divaricate; eatin to sou ie Monterey County. 9. H. lobbit. 8. H. mohavensis. SUNFLOWER FAMILY 173 Ray- and disk-flowers more _ ni Ray- she ‘sans flow Her! bearing” pustuiate hairs, gray- reen; plants ae intricately ments mterey County to San Luis Obispo County. 0. H. Herbage bearing simple filamentous hairs, yellow-green; plants ies. ae intri- cately branched; southern California. Heads saistclaiate, stems tall, scarcely hispid, flexuous and much branched; San Luis Obispo County to San Diego County 11. H. ramosissima. Heads in dense glomerules; stems lower, Yithis hispid above, with strict divaricate branches; coastal plain to Lower ein ae 12. H. fasciculata. Ray-flowers 8; disk-flowers 13—21; southwestern San Diego County. 13. H. conjugens. Disk- ae nied fertile. Rad eaves pinnatifid or bipinnatifid; herbage — -hirsute; ray-flowers 8-13, ligules as br og as long; cont central California to Lower California. 14. H. paniculata. sma: * ve —. — ; herbage bright green, soft- ie eo sad’ mr a by 13-20, ligules half as broad a ; rare; southern San Diego Cou H. floribw nda. } = 1 free, Leaves tipped with a mae spine or pete ean receptacular persistent ; pappus none or of very narrow paleae (Section C seaiens by Pappus none; anthers yellow Receptacular foots veneent: northern and central Califor; H, pungens. Seite bracts obtuse or more or less acute, not par southern California. H, laevis tacular bracts flesh earing Ss! nail Siowtal a if any, of mild odor fe i yellow; central Californ Anthers black; inflorescence very ime southern California. Receptacular bracts long-villous at tip; pappus-paleae 8-12, densely fimbriate at tip; herbage bearing large, eee — Leap he rank odor; anthers black. 20. H. fitchit Pappus F eoe Rec tip, not long-villous; pappus- sag 3(—5), sparsely ciliolate; herb- 18. H. parr 19. H.a usteuble. Ray achenes not — sly beaked; ligules ; inner receptacular bracts adnate, forming a cell about each disk- flower, ieiquescent ‘(eection peel pcone ia). Ligules vel dorsally veined with p pre ; S pro ges 2 time of flowering, er not crowded into 21. H. multicaulis. Spring-flowering (May— a bas: Fall- eer ay! (Aug.—Oct.), taller; leaves tai SE at time of flowering, very short, crowded a basal rosette; dens - pea throughout. 22. H. lutescens. Ligules cian eae a ag hon Heads paniculate or nal; leaves broad. ae bracts short, not overtopping the heads. e green, merely puberulent; heads scarcely glandular; esha a ae” 23. H. trac. reggae gray or silvery, pubescent; heads obviously glandular; spring- to fall-lowering: nflorescence widely paniculate, the heads scattered; herbage villous and copiously dark-glandular, or comms phyllaries 3.5—6 mm. long; mostly inland, north- forn 24. H. luzulaefolia. pero puberulent to sericeous or villous with soft hairs; lower ern and centr al Californ Inflorescence corymbosely eaieuiata: the heads usually glomerate; herbage shaggy with few glands; phyllaries 6-9 mm. long; coastal essa 3 a of a Ven sco Bay. 25. if. oi See bracts long, overtopping the heads, calyculate; herbage S dtecagte gts Men and Lake et age 26. H. calyculata. veloped side-branches, obviously Giltornia, herbage pilose; Heads epetie, pahe on the fully ower leaves narrow; basal obra pir cider: Oregon to northern H, clevelandit. 5268 174 COMPOSITAE 1. Hemizonia clementina pie Catalina Tarweed. Fig. 5270. Aiseesip —— rl ease vghaie| Erythea 7: 70. 18 Davids. & Moxley, Fl. S. uc 401. 1923, Shrub 3-8 dm. hi vga ith many erect, ascending or decumbent branches from base, densely read es, kas older “eaves deciduous ; : he rbage siahaeks hirsute, viscid. Lower leaves opposite, arrowly linear, 3-8 cm. long, 1.5-6 mm. wide, reineels tate; upper leaves alternate, entire, with pee, or lea i shoots in their ie heads ound cymes; involucre broadly cam- panulate to ve c, 5-8 mm. high; ray- -flowers 134, ie nae yellow, 4.5-6.5 mm. long; receptacular bracts in 3 mae disk. acne 18-30, t central 0 ed without bracts; ray-achenes 2-3 mm. long, emauee a the minent Pes beaks and one-half to three times as long as thick; disk-achenes or i i a of 7-10(- 2 linear- il attenuate, fimbriate, unequal paleae 1-3 mm e. In hea oils, Upp one; Anacapa, San Nicholas, and Santa Barbara Islands, Ventura County, pug San Clemente a sare vpn Islands, Los Angeles County, California, Type locality: San Clemente 2. en minthornii Jepson. Santa Susana Tarweed. Fig. 5271. thornii Jepson, Man. Fl. Pl. Calif..1092. 1925. Shrub 6-10 dm. high, 10-30 dm. wide, with up to 500 stiff, woody, ascending stems from bas herbage short- hirsute, paar fragrantly resinous. gene esa alternate, somewhat thickened, like = ai of seam tina; heads mostly solitary on long peduncles, racemosely or corymbosely paniculate ; 46 8; i : oe mm. ee: olucre 5. m. hi mm. al r receptacular bracts in about 3 series; disk-flowers 18-23, each subtende a bract; ray-achen .5-3 mm. long, smooth, the beak scarcely longer than thick; disk- rn Beate but Saat pappus of 8-12 linear, more or less fimbriate, subequal paleae 1.2-2.6 mm. long. Dry tr slopes, Upper Sonoran Zone; Santa Susana ia, Ventura County, California, the type locality. July— 3. Hemizonia kelloggii Greene. Kellogg’s Tarweed. Fig. 5272. ements aoe Greene, Bull. Torrey Club 10: 41. (Apr.) 1883. ighttti A. Gray, Proc. Amer. Acad. 19: 17. (Oct.) 1883. Sear kellogait eas Fl, eh ran. 424. Deinan ndra a wrightit Greene, loc ia wrightit var. " Rellogoti ak Man. FI. Pl. Calif. 1090. 1925. em 2-10 dm. high, corymbosely branching above or nae pes by pennchens ow ohion soft- above eae at base, hispid-hirsute and aged eerie aves narrowl long, re- —— sharp oe ed or ey tifid, m. long, 3-10 mm. bide 0 iene leaves entire; open panicle or thyrse; Ltchecre 4-5.5 mm. long, 3. mm. wide, densely stipita wie: eee tk usualy caseunre ligules 4-7 mm. long, 3.5-5.5 mm. wide; pappus of 6-12 white, linear to oblon g palea Dry nage hillsides, Upper Sonoran Zone; San Francisco Bay region south through the San Joaquin and Coast Res poked to San Diego County, California; adventive in Mendocino, Colusa, and Imperial Counties. Type locality: near Antioch, Contra Costa Cou . April- —July. 4. Hemizonia pallida Keck. Kern Tarweed. Fig. 5273. Hemizonia pallida Keck, Madrofio 3: 8. 1935. Ste m 2-8 dm. high, branching above or throughout with ascending or divergent branches, whit- ish or reddit herbage villous-hirsute and hispidulous, lightly g 55 eS Bess ildly odoro ~ Lower leaves linear xg Sehlas nceolate, epee — a cleft, 5-10 cm. long, 3-6(-10) m wide, the bes pper o oming entire ; Reta th as re 4.5-6 .5 mm. bi 5-8 mm. wide, hispid-hirsu ute and Ri sate & stipitate-glanduiar tigales 6-10 mm. long, 3-5 mm. wide pappus of 4-8 narrow distinct paleae 0.8 m r less lon Dry plains, U mee Sonoran Zone; head of “58 tioned Valley, California. Type locality: 5.3 miles north of Grapevine, Kern nty. April-May. 5. Hemizonia arida Keck. Red Rock Tarweed. Fig. 5274. Hemizonia arida Keck, Aliso 4: 109. 1958. Stem 2-4 dm. high, intricately corymbosely caer peep herbage hispid-hirsute and hirsutulous, ‘lightly glandular and mildly odor Leaves about as in pallida; heads numer- ous, cymose-paniculate ; involucre 4 mm. high, 5 mm. n wide, Bispid Sieve aw stipitate-glandular ; ligules 5-6 mm. renee 3-4 mm. wide; pappus vestigial o oe —S sandy canyon-bottom, ere aes Zone; Red Rock Pi te: Sas County, California, the type locality. ay~ ee Hees corymbésa (DC.) Torr. & Gray. Coast Tarweed. Fig. 5275. Cc Hartmannia cor DC. op. cit. 694. sa Torr. & Gray, Fl. N. Amer. 2: 398. 1843. slemizonia deculiens Nutt. Journ. Acad. Phila. II. 1: 175. 1848. SUNFLOWER FAMILY 175 emizonia balsamifera Kell. Proc. Calif. Acad. 2: 64. 1860. Zonanthemtis angustifolia a ne, a Fran. 425. 1897. Zonanthemis corymbosa Gre cit. Hemizonia vase tial var. seapaabes Jepson, Man. FI. Pl. Calif. 1091. 1925. Stem bent, 2-10 dm. long, branching from base upward or from above the middle, the branches ascending or spreading; herbage den an villous and glandular, alg and pleas- eaves linear to oblanceolate, = ply pinnatifid, 3-10 cm. lon "6, 3- 20 mm. ing entire in — plants ; heads Aaah or panicult e; involucre 5-9 mm. hig . wide; a ules 4-7 mm. long, 2.5 mm. wide ; we gs at ‘distinet, un- equal, entire or laciniate cas oi mostly 1 eae or less long, or wa sien Grassy slopes, Humid Transition Zone; along the coast, Mendocino County to Montene County, California. bic oe ig ae Collected by Douglas. May- 7 subsp. macrocéphala (Nu 109. 1958. (Hemizonia 13, 1938.) erect or decum Keck, Aliso 4: mbos ea nace Rest Phila. aL 1: 175. 1848; H. “angustifolia pom b macrocephala avarias bg aed é: 12 Stouter, denser; — densely wag shee H Ms "the = the branches, cymose, larg ray-flow 20-35; disk- flowers 40-70. ‘Coastal soiihers M — and n ner ah oo Luis Obispo Ccuation. Colforsis. age locality : “St. Stescan,”* San “Laie Obispo Count 5270. Hemizonia clementina 5272. Hemizonia kelloggii 5271. Hemizonia minthornii 5273. Hemizonia pallida 176 COMPOSITAE 7. Hemizonia halliana Keck. Hall’s Tarweed. Fig. 5276. Hemizonia halliana Keck, Madrofio 3: 12. 1935. ca —— dm a up to 16 mm. thick, a and glabrous below, corymbosely branching (only n summ o below the midd e) a becoming pilose an viscid-puberulent above hertan: cei ne ae Leaves linear- eda sessile, glabrous except for the sca abrous margin, the lower remotely short-dentate, 5-8 cm. long, mm. w * volucre 5-7 mm. high, 10 mm. wide; ray-flowers 10-14, the ligule 5.5 mm. long; disk-flowers 30-60; ray-achenes 3.54 mm. long; disk-achenes marae ened a ere omnacindes paps. Heavy adobe or serpenti Upper Sonoran Zone n the r South Coast Range, San Ben County to x hey Luis Obispo Comite, California, Tor g ene :¥ 5 ‘wiles nai oF Chaleene: coe Luis Obispo a 8. Hemizonia mohavénsis Keck. Mojave Tarweed. Fig. 5277. Hemizonia mohavensis Keck, Madrofio 3: 9. 1935. Stem 1.5-3 dm. high, subsimple or divaricately branched above; herbage very — Aken te and viscid, pleasantly odorous. Lower. leaves oblanceo a subentire, the upper lea ong- lanceolate, entire, amplexicaul, much reduced toward the expanded-co igre mbose Paindesceice: heads Sprgeeith in s Mesos at the ends of the branches; involuc —6 mm, high, 5 mm. wide; ‘ligules 5 ong; iit s of 6-8 fay uadrate, Asal or less co ieee oie 0. 5 mm. lon ong. ioe: und, Low: Upper Son ; kno ut two stations in Califor rnia, Mojave “pe fo its Pi Apa with Dee Creek, Watens eokg (the type eeahiay and Banning-Idyllwild road, Mount San Jac Riverside County. July—-S 9. Hemizonia lébbii Greene. Three-rayed Tarweed. Fig. 5278. H emizonia lobbii Greene, Bull. Torrey Club 9: 109. 1882. Hemizonia fasciculata var. lobbit A. Gray, Syn. Fl. N. Amer. BP: 310. 1884. panaeaee lobbit Greene, Fl. Fran. 425. 1897. Stem agin slender, 2-5 dm. high, rigidly branched above or throughout, the branches divaricate and again very freely bran ching toward tips; herbage pene? - SDR, hispid-hirsute, — beset vith phai s glands pores ildly odorous. poet, mall, and entire at anthesis, tho e seit ear Sa crowded, t cue iduous lower leaves TN ariatehy ghateionped or peneitisect eads very numerous, subse sie to short edn cled ; involucre mm. long, 2.5-4 mm. wide, i se sessile glands a nd some s more or te ss hispid with pustulate hats: ray- and disk-flow 3 each; pappus of 6-8 lan alate or oblong, entire or toothed paleae equaling the corolla-tu Dry ean san oye a Upper Sonoran Zone; Solano County to southern Mauter ey County, California. Type locating : “probably r Montere y” but more li kely in the Salinas Valley. May—Nov 10. Hemizonia pentactis (Keck) Keck. Salinas River Tarweed. | Fig. Saf 9. Hemizonia lobbii subsp. pentactis Keck, Madrojfio 3: 8. 1935. Hemizonia ass canes Keck, Aliso 4: 109. 1958. Having the low, onpead branched habit, grayish herbage, and pinnatisect basal leaves of ore the gee and heads m ge —— pustulate-hirsute and ‘hispid, the leaves whiter-pilose ; ray- -flow ; disk-flowers 6; wise similar to Dry “aie hills, Upper ane Dic tae southern, Monterey ‘asd San s Obispo Counties and Liens es about Stanford University, California. Type locality: San Miguel, San Re Obispo ounty, May—Oct ii, rons ramosissima Benth. Slender Tarweed. Fig. 5280. Hemizonia ramosissima Benth. Bot. Sulph. 30. 1844. Hemizonia — ta var. rbasliins A: Gray, Syn. Fl. N. Amer. 17: 310. 1884. Deinandra culata v avids. & Moxley, . S: Calif. 401,-.1923;, scicula amosissima Dav Hemizonia favcatinn athe ramosissima Keck, Madrojio 3: Stem erect, 2-10 dm. high, simple below and be ay branching above or with ascending branches from the base upward; herbage moderately hirsuite to glabrate, the glands (most fre- r le m quent on involucres) sessile, yellow. Lower leaves (most issing at anthesis) linear-oblanceolate remotely den 1 . long, 3- com Dwi: upper leav t all, and bract- like; heads pedunculate, numerous, r less remote — solitary, sometimes in twos at the mn of branches; ray- -flowers ah disk- ‘foteers & pappns s in lobbit. y fields and hillsides, Upper Son oran Zone ict Codes and the Channel Islands to Oran Camere largely replaced by the next in San Diego County. Caifooni ia, and Lower California. Type locality: on ably San Pedro, Los Angeles County, California. May—Aug. 12. Hemizonia fasciculata (DC.) Torr. & Gray. Fascicled Tarweed. Fig. 5281. cheated egies DC. Prod. 5: 693. 1836. Hartmannia glomerata Nutt. Trans. Amer. Phil. Soc. IT. 7: 391. 1841. How ionia fasciculata Torr. & Gray, Fl. N. Amer. 2: 397. 1843. A. Gray, Syn. Fl. N. Amer. 17: 309. 1884. —— fasciculata Ceceas Fl. Fran. 424. 1897. Deinandra simplex Elmer, Bot. Gaz. 39: 48. 1905. Stem 1-10 dm. high, mostly branching from above the middle, the branches rigid, sharply ascending and with Coneratively few twigs; heads subsessile in glomerules of 3 to many, the JINFLOWER FAMILY 5277. Hemizonia mohavensis 5278. Hemizonia lobbii 5279. Hemizonia pentactis 177 178 COMPOSITAE glomerules Sapient short leafy branches and sometimes a few solitary heads below; other- wise similar to ramosissima py pean plains, 7 il Sitarkh Zone; frequent in California from San Luis Obispo County to Riverside Coun se ndant thence to a California. Type locality: ‘‘Nova-California.” Collected by Douglas, probably near natn Barbara. May—Sep 13. Hemizonia cénjugens Keck. Otay Tarweed. Fig. 5282. Hemizonia conjugens Keck, Aliso 4: 109. 1958. Stem 2-5 dm. high, branching as in fasciculata ; pees ge and involucres bah hirsute and some- times Rapithane: erie | : ade inv ee bearing large, flat and s mall, capitate, sessile or sub- aii glands; heads solitary on short peduncles or edecesite in tw heated glomerules; ray- flowers 8-10; di sk-flowers 13-21. Dry mesas, rper Sonoran Zone; southwestern San Diego County, California. Type locality: river bottom near Otay. May- June 14. Hemizonia LS pei Gray. San Diego Tarweed. Fig. 5283. Hemi. paniculata A. Gray oc. Amer. Acad. 19: 17. 1883. Deinandra paniculata Davids. & a FL. S. Calif. 401. 1923. Ste 2-8 dm. high, the central shaft displaced about midway by numerous ascending branches which ramify to form a twi inflorescence; herbage moderately hispid-hirsute (especially 1] e mged to ‘amg nas sharp-toothed or pinnatifid with entire or dentate lobes, 1.5-8 cm. long, ; i pedunc ucre S mm. hi i ra) ie) - s 85 7 ma ae BS Qa 2 < os Let = . to gh, 5-7 mm. wide, the ‘ovine dottenis Sear cane cent; ray-flowers 8; disk- 8-13; disk-achenes ae pubescent, sere pil and Se clotel, the majority te. pene pale; pappus of 6-12 white, oblong, fimbriate palea Dry hills Ey. fe ae Sonoran Zone; western Riverside County to San Diego County, Paliprnia, and ~~ ern Lower cS Type locality: near Siineculs, ‘Riverside icrtitey (lectotype). May—Nov onia paniculata subsp. incréscens Hall ex Keck, Madrofio 3: af 1935. Plants erect or somewhat spnendinn “deep green, even the ailtimate twigs rather reid (never capillary); ray-flowers 8-13; disk- overs 14- 30. Abundant in fields near the coast, Upper Sonoran Zone; nterey fone to Santa Barbara Count ty and on San Rosa Island, Santa Selene Counts. California. Type lo ae 7.5 miles southwest of Arroyo Grande, San tne Obispo County. May—Nov 15. Hemizonia floribinda i Gray. Tecate Tarweed. Fig. 5284. Hemizonia floribunda A. Gray, Proc. Amer. Acad. 11: 79. 1876. Deinandra floribunda Davids. & Moxley, TL. S$. aks 401. 1923. Stem 3-10 dm. high, corymbosely branched above or throughout with strict say branches; herbage moderately pil ose and densely ek Poets -glandular throughout, mildly tag Leaves mostly entire, the lower oblanceolate, the middle cauline linear-lanceolate, 1,53 ¢ Gare 2-3.5 mm. wide; heads short-peduncled, in Sesage ely _compoun cymes o a dome Ne a glo- merate ; involucre 5-6 mm. high, 6-7.5 mm. wide, soft- chee and very g tohdu ar; ray-achen 13-20; disk-flowers about 28, their achenes all ae aaeity Pag sapens of 6-9 oblong or elliptic, closely fimbriate, rufous and flecked paleae. Dry hills and montane valleys, Upper at oran Zone; southern San Diego County, California, and adjacent be California Sei Jo locality: California, near the so uthern boundary, on the Fort Yuma road, 80 miles anal an Diego. 16. Hemizonia pingens (Hook. & ag ty _ & Gray. Common Spikeweed. Fis. ——— i sie > sd Hook. & Arn. Bot. Beechey Sy 357. 1838. orr. & Gray, Fl. N. Amer. 2: 399. 1843. Centromadia vet Greene, Man. Bay Reg. 196. 1894. Hemizonia pungens subsp. interior Keck, Madrofio 3: 14. 1935. Stem 1-12 dm. high, divergently and rigidly heat oc! oan or throughout, leafy, sparsely to copiously hirsute, not gla iy - herbage yellowish green, inodorous. Basal leaves linear- lanceolate in outline, $15 c . long, 10-40 So wide, bipinnately parted into segments; upper cauline leaves with fascic age in her axils, mostly entire, spine-tipped, stiffly ciliate and sca- brous-puberulent ; heads apheepils in upper axils and terminal; involucre hemispheric, ae mm. y bracts i i > sie disk-flowers numerous, the ligules only 2-toothed; ray-achenes angular, often rugose; > ihiak. achenes in part pea "the fertile ones more or ‘tee ‘fusiform. Dry valley floors, Upper Sonoran Zone; S n Joaquin Valley from San Joaquin County to Kern County and Selina Valley. Culitdvnins introduced in Los ‘Auieeles, San Bernardino, and San Diego Coates and in northern Oregon. Type locality: Caines ese Oct Hemizonia poneens (Gee Keck, Aliso 4: 110. 1958. (Centromadia maritima Greene, Man. Bay Reg. igs 1894.) Brant, fa fairly ion. the branches usually iia uete ~ strict; leaves and bracts not scabrous; heads lomeru! mbers, large. Salt marshes and valley land around San Francisco Bay south to nort nactnese onterey Cocky me i ast to San Joaquin and Yolo Connties: Coitermie Type locality: Alameda, has apy ch Hemizon he septen a Bone ig Aliso 4:110. 1958. Plants large, the so ogee fring noe and often oe ghee pe bracts not scabro scattered, small. Dry ground, Upper Sonoran Z Sacra- mento Valley’ and feng 3 Valley, California: aah ee in northern Oregon and Washington. icin locality : Shi alley, Siskiyou County. SUNFLOWER FAMILY 179 wy og dh Re 7 \ oye ‘Bs We » 1G XY — Mae, \ Wawa) 5 § NO ae a e7y tor, wi SX (ft ay ( NE 5280. Hemizonia ramosissi 5281. Eoranas prone 282. Q COOPER, com | = p> 5283 . Hemizonia 5284. Hemizonia gat tit eat Hemizonia pungens 180 COMPOSITAE 17. Hemizonia laévis (Keck) Keck. Smooth Spikeweed. Fig. 5286. Hemizontia pungens subsp. laevis Keck, Madrofio 3: 14. 1935. Hemizonia laevis Keck, Aliso 4: 110. Similar to pungens but of somewhat ula habit. Upper leaves and bracts — setose- ciliate, het wise very glabrous; heads small, scattered, or approximate in loo lomerul a bracts obtuse or P liphaty Prssren sometimes minutely but weakly nate not cu Dry fields, Upper Sonoran Zone; cismontane southern California away from the immediate coast from Los Angeles County south; also southwestern Kern County. Type locality: San Bernardino Valley. April-Sep 18. Hemizonia parryi Greene. Pappose Spikeweed. Fig. 5287. Hemizonia parryi Greene, Bull. oe a 9: 16. 1882. . stent seins SAP, Man. Bay . 197. 1894. var a Jepson, FLW . Mid. Calif. 532. 1901. Hemizonia p wages ns var. parryt H. M. Hall, Univ. Calif. Pub. Bot. 3: 155. 1907. Stem 17 dm. high, erect, corymbosely bran xm above, the branches rigid or lax, or rather prostrate, more or less leafy, sparsely to copiously hirsute. "Basal leaves linear- to lance-oblong, 5-20 cm. | Pa My wide, tifi i 20 c eye mm. pinnatifid > uiineees r the lower cauline more or less hirsute, thei re r less ‘spine-tiped, ae upper mostly Vegi linear-subulate, spine-tipped, wit fascicles. (incipient shoots or heads) in their axils; eee ssile or short- peduncled along the branc and terminal, 2A small clusters; peduncular thee # often exceeding the involucre; seen hhemispher ric, 5- oe mm. high, the aie glandular-pubescent, copiously ciliate, the glands subsessile; ray-flowers 9— 20, the ligules mm. long, yellow, not fading saffron, : receptacular bracts w oe eainois thickened, soft, obtuse ’ acute (not cuspidate) tip; disk- flowers or more; dis enes in part sterile; pappus 3-5 el subulate, usually fimbriate paleae about equaling the corolla, often united at base — contorted. Low, often bemcur = fields, Upper Sonoran Zone; Colusa and Napa Conti to San Mateo and Monterey Counties, "Califor . Type locality: Cali istoga Springs, Napa County oe e—Oct Hemizonia sane subsp. ridis (Greene) Keck, Madrotio 3: or aCe ntromadia rudis Greene, Man. Bay Reg. 197. 1894.) Stem erect, corymbosely or di ffusely rwlcaalice § not prostrate, herbage pale green; leaves psa! bracts scabrid-puberulent; inflorescence obscurely if at all glandular; peduncular bracts not &xce eding the involucre; heads small; ray-flowers often den en saffron. | in i. fields, a aa Sencean Zone; Great Valley of California Glenn County to Me nty. Type loc County. June—Oct Hemizonia parryi subsp. congdénil Seer & Green os K eck, Madrone 33° 5. 1935. Gtléavixonia cong- donti Robins. = Phage Eats _Gaz 1896; Centromadia congdonii C. P. Smith, Muhlenbergia 4: 73. 1908; C. pung ongdonti Je epso i oF. PF. Ca WE, 1087. 1925. Ste m erect with ascending or horizontal branches, or al "branches ® closely prostrate; herbage yellow-green, ar to copiously be stor but smooth between e long hairs; pedun r bracts exceeding the iavelaare: ray-flow not fading saffron. Low aor ege Upper Faeroe Zone; San seas Valles Pontes Costa County; lower a of San Francisco Bay, mostly Alameda or and lower Salinas Valley, Monterey County, California. Type locality: Salinas, Monterey County. June- 19. Hemizonia australis (Keck) Keck. Southern Spikeweed. Fig. 5288. ~ihetes emizonia parry: subsp. australis Keck, Madrofio 3: 15. 1935. Keck, Aliso4: 110. 1958. Similar to parryi; stem erect, the central leader.of medium length and with long and lax, divaricate poe hes, not prostrate, the twiggery above dense; herbage dark green. Leaves and bracts densely glandu ar-puberulent and villous; cm bracts not exceeding the involucre; heads small; phyllaries 4-5.5 mm. long; r Bases aig fading saffron; anthets lack. Low alkaline fields near the coast, Upper Son one gs arbara County to San Diego County, Cali- fornia, oa adjacent Lower California. Type tecuhite: soe Beach. g Roche County. June—Sept. 20. Hemizonia fitchii A. Gray. Fitch’s Spikeweed. Fig. 5289. Hemizonia fitchti A. Gray, Pacific R. Rep. 4: 109. 1857. Centromadia fitchtt Greene, Man. Bay Reg. 197. 1894. Stem 2-8 dm. high, rigid, erect, above diffusely branched; herbage we a | villous and 1g) beset with prominent stalked glands, unpleasantly heavy-scented. Leav wded, at least in the rosette, very slender, the lower pinnatisect, up to 15 cm. long, the pe oitiee, rigid, linear- su te, acerose-tipped, often with fascicles in ee — soe dB a involucroid, spreading, radiating around the head which they ele, voluc ispheric, densely villous, es — prominent glands, the phyllaries rigidly 1-nerved, with aera Ling slightly spressine the receptacular bracts soft, pointless, hairy, enfolding the disk-flower; ray-flowers 10-20, the Beet light yellow ligules bifid; pappus of 8-12 glistening-white, soft, obi sin leine prose at base and long-fimbriate above, equaling the disk-corolla. Dry hills and plains away from the immediate coast, Upper Sonoran Zone; southwestern Oregon to San L Obispo County, Ca oe Sierran foothills from Siskiyou ounty to eo County, California; Central Valles sia patie bese of toe e San Bernardino Mountains , California. Type locality: “Plains of the Sacrame nto, Cali- 21. Hemizonia multicatlis Hook. & Arn. Seaside Tarweed. Fig. 5290. Hemizonia multicaulis Hook. & Arn. Bot. — = 355. 1838. Stem scendin, ecumbent, 1-3(-5) dm. high, subsimple or openly branching above or theca: ica sel BA ar Riad not crowded into a basal rosette, the lower SUNFLOWER FAMILY 181 remotely serrulate, pean a 6-15 cm. long, 6-12 mm. wide, deep green; heads vt lege - olucre hemispheric, 5-6.5 m hi gh, the A 2 the colar tgs about equaling the body; r flower 5 8-13 3, dears, ye alton (sometimes drying green ‘ lat bluffs along the immediate coast, Sonoma and Marin Sane California. Type locality: not given, May-July. Hemizonia multicaulis yen —— Keck, Aliso 4: 110. 1958. Tot ceed citrina Greene, Man. Bay Reg. 194. 1894; H. luzulaefolia v citrina Jepson, Fl. W. Mid. Calif. 530. 1901, probably belongs in this rene Stem erect, corymbosely pstdvtert dey pied Sender: leaves grass-like, narrower, aut silvery- -sericeous. Mostly away fr “haa the immediate coast, Sonoma and Marin Counties, California. Type locality: Tiburon, Marin County. April-Jun Tae AL, te ar 5288. Hemizonia australis 5289. Hemizonia fitchii 5286. Hemizonia laevis 5287. Hemizonia parryi 182 COMPOSITAE 22. Hemizonia lutéscens (Greene) Keck. Sonoma Tarweed. Fig. 5291. Hemizonia luzulaefolia var. mee Greene, Bull. Torrey Club 9: 16. 1882. Hemizonia congesta subsp. lutescens Babe. & Hall, Univ. Calif. Pub. Bot. 13: 38. 1924. Hemizonta congesta var. hemes = Man. FI. Pl. Calif. 1089. 1925. Hemiszonia lutescens Keck, Aliso 4 Ste robust, 2-7 a high, usually much branched from the base upward; herbag dark- ni peo ay inflorescence or throughout, heavily scented; basal rosette at first eon nent. Leaves short and narrow (up to cm. long an mm. wide), often silvery-vill ith long cobwebby hairs; heads small; involucre 4.5—5 mm. high, the tips of the phyllaries shorter than on body ; ray-flowers deep yellow. Grassy valleys and hillsides away from the immediate coast, Upper Sonoran Zone; southern 5 i ions hang County rs sin and Alameda Cities: Cities, Type locality: San Pablo, Contra Costa County. July—Nov 23. Hemizonia tracyi (Babc. & Hall) Keck. Tracy’s Tarweed. Fig. 5292. Hemizonia congesta subsp. tracyi Babc. & Hall, Univ. Calif. Pub. Bot. 13: 46. 1924. Hemizonia congesta var. 6 sigh atime Man. FI. Pl. Calif. 1089. 1925 Hemizonia tracyt Keck, Aliso 958 Stem slender, 1.5-6 vv hi bi sik mple or openly few-branched above, — villous or merely densely puberulent ; herbage s oft- seen glandular ant aroun ne the Ea Leaves not crowde a basal asi grass-like, very n w, the lower up to 15 cm. lon bright ah hes ds mbo se, relatively aig piel e 5-7 mm. high, cin ts of the phyllaries shorter cs the body + eet, chenes one- “half to 3 fiths as broad as lon and forest Humid Transition Zon uthern Humboldt and nort thik n Mendocino Coun- pees Hie Poh Type lochkitey pales of t the Van Predets River’ lapouitk Buck Susie. Humboldt County. May-— 24. Hemizonia luzulaefélia DC. Hayfield Tarweed. Fig. 5293. Hemizonia peat saben, DC. Prod. 5: 692. 1836 Hemizonia sericea k. & Arn, Bot. Beechey Suppl 3 Hemizonia ano. var. fragarioides Kell. Proc. Sess pare 2: 69. 1860. i 7 rams © Hemizonia congesta subsp. Iuziulaefolia Babe. & Hall, Unite. Calif, Pub. Bot. 13: 43. 1924. Hemizonia congesta var. luzulaefolia Jepson, Man. F1. Pl. Calif. 1089, 1925. Stem slender, 2-5 dm. high, openly branching above or throughout, mostly vie es well down oward base, often also more or less villous; basal rosette none or obsc Low leaves prominent at pga oh pee “ae a 15 or more > long and to . wi ide, gieen aad chiefly eee ioe si i with cobweb-tomentum, not glan dul is rs heads sho owy; involucre 5—6 mm. high, che tips oe ‘the Shllarics shorter than the body; ray- -achenes about one- half as pre as tae Open ground, coastal valleys and foothills, Seve Sonoran Zone; Lake County to San Luis Obispo County, California. Type locality: peste April-Jun Hemizonia luzulaefo subsp. ridis (Benth.) Keck, Aliso 4: 111. 1958. (Hemizonia rudis Benth. Bot. Sulph. 31. 1844.) More poe 2-8 dm high, a ear and den sely gla ndular-viscid througho out; basal rosette h m r of the secti .5—-5 mm. high. e most widespread and abundan tion, in st polenelds and pastures peer Sonoran Zone; Sacramento ewig —— Tehama County “‘ Boas Joaquin County, Coast Range valleys fr. ake County to Santa Barbara Cou ery rare north of = noma County, and in Monterey County, California. Type locality: Santa Clara, Santa Clara County. July—Nov 25. Hemizonia congésta DC. Hayfield Tarweed. Fig. 5294. Hemizonia ets. ate DC. Prod. 5: 692. 1836. ‘¢ eve wn 5-8) 2. w eee eas = Fo} = oOo r but ina silly, saci viscid toward heads but the glands us sually obscure ; or Lo leaves usually prominent at. abthests: remotely denticulate ; heads small terminal clusters or solitary at t he e ends of the short branches and often a few watt: nerd heads; involucre 6-9 mm. high, the tips of the cay iteeles Datialine to acs peiedind the body ; iy sechieties aosily three-fifths as broa Fields near the coast, Humid Transition—Zon : Del Norte County to San Mateo County, California, but fre- quently only in Sonoma and iris Pounties Type ages: California. May—Oct. 26. Hemizonia calyculata (Babe. & Hall) Keck. Mendocino Tarweed. Fig. 5295. Hemizonia —— subsp. calyculata Babe. & Hall, Univ. Calif. Pub. Bot. 13: 42. 1924. Hemizonia c a var. calyculata Jepson, Man. F1, Pl. Calif. 1089. 1925. Hemizonia orecanee Keck, Aliso 4: 111. 1958. Stem 2-8 dm. high, divaricately branching above; herbage moderately villous and densely glandular-pubescent, the dark stalked glands abundant in pod inflorescence. Lower leaves not recent at preg Erect osiats ies cm. long, 2-5 mm. wide; heads terminating — stiff peduncles, the floral bracts crowded, ending to surpass Phe he ads ; involucre 6-12 m ~~ ae tips of the phyllaries exceeding t a body; ray-achenes one-half to three-fifths as beted Ids and open hillsides, r+ Sonoran Zone; Lake and southern Mendocino Counties, California. Type boa east of Orr’s Springs, isis Md ino County. July—Oct. SUNFLOWER FAMILY 183 ia tracyi 5293. Hemizonia luzulaefolia 5294. Hemizonia congesta 184 COMPOSITAE 27. Hemizonia clevelandii Greene. Cleveland’s Tarweed. Fig. 5296. Hemizonia puecicainl na Bull. Torrey Club 9: 10 Hemizonia congesta s clevelandti Babc. & Hall, as nen Pub. Bot. 13: 48. 1924. Stem 2-6 dm ie ie simple to divaricately br sc above or throughout, or with several cewtng branches rots base; herbage of early-flowering forms usually densely i i oe cially toward base of stem and setts and moder et inci lar above, of late-flowering for moderately villous or merely — he strongly glandu a pubescent especially ‘toward i inflorescence; basal rosette small o al leaves narrow, to 15 cm. long, to 7 m ide ; heads terminal on the branches in He tee ae pie icate- te il oA in late Fe rms ; eh we 4-7 mm. high, two- thirds as eral Dry grassy i: of the interior, Upper Sonoran Zone; southern Oregon south through both Coast Ranges to Sonoma and Napa Counties, California. Type locality: Allen Springs, Lake County, California. June—Oct. hg sa my the i iaties ie shorter than the body; ray-achenes about ong. 32. HOLOCARPHA (DC.) Greene, FI. Fran. 426. 1897. Annual herbs, mostly very glandular and aromatic. Leaves linear, the basal remotely aeuaske to dentate with slender teeth, the upper reduced to entire bracts usually bearing fascicles or peduncles in their axils, truncated at apex by a crateriform gland. Phyllaries half-enclosing the ray-achenes, bearing on back and at apex stoutly stalked pit-glands. Receptacular bracts subtending each disk-flower, pen eis the achene and corolla-tube, free, persistent. Flowers yellow, the rays 3-lobed. ee achenes ne beaked, obcom- pressed, nanan the odd angle anterior. Disk-achenes sterile or outermost fertile. Pappus none. [Greek, holos, whole, and Deameh id chaff, the cuties eee chaffy. | A genus se 4 species, all . Type species, Holocarpha macradenia Anthers black; herbage not puberulent above. wiseeaieg see 40-90; ray-flowers 8-16; heads very large, densely glomerate; low, with robust divaricate ches; rare, near the coast. 1. H. macradenia. Disk- ee Bench 25% -flowers 3-7; heads small, fusiform to subglobose, soanctan —- with Pini cad as- yivaecias: interior virgata Anthers RP: ok ium tall; interior. Herbage net densely puberu! above; inflor nee corymbose-paniculate; involucre ee the paraees bearing 5—15(-20) ens sland tipped He receptacle broad. obconic Herbage pss puberulent above; inflorescence r i eg ii subglobose, the Phares ee 25-50 very slender, gland-tippe od processes; receptacle nar H. heermannii. i: a macradénia (DC.) Greene. Santa Cruz Tarweed. Fig. 5297. C. Prod. 5: 693. 1836. Soaps sadarites Greene, Fl]. Fran. oe. 1897. Stem 1-5 dm. high, rigid, divaricately branched above the base, the few branches arising near res ether and usually again similarly br agnes rather — leafy. Basal leaves usually lost before > broadly linear, up to i2 cm. long and 8 mm. wide including the slender remote teeth; upper leaves becomin ract-like with bevedentt’ ee conspicuously bio ‘oar lar-pubescent cae dak cite the apical gland sometimes occurring only heads mostly in terminal glomerules ; sige subglobose, 5.5-8 m pie ¥ the Silas, ah about 25 stout, terete, eed -tippe eB ocesses ; ray-achenes 253 m . lon Colonial in heavy soils gras: hs ng the coast, Humid Transition . Sexik: Alameda, and Santa Cruz Counties, California; io — po teay Type locality: California. June—Oct. 2. Holocarpha ~—" (A. Gray) Keck. Virgate Tarweed. Fig. 5298. Hemizonia nett: A. Gray in Torr. t. Mex. Bound. 100. 1859. Deinandra virgata Greene, Fl. Fran. re 1897. Holoca aha é jake Keck, Aliso 4: 111. 1958. Stem 2-12 dm. high, rigid, usually simple ao with few to several ide sharply ascending aches above; herbage spa eesigty (omen bee glandula Paige lent, canescent or hispidu- lous above and st cael resinous and odor Basal leaves 6-15 cm. long, "Sie mm. wide, mostly lost before anthesis, the upper sedncee to Jareat often tinchenia: those of the peduncles ceceeded, heath-like, sp eading or recurved, the apex terete, prominently gland-tipped ; head short-peduncled, — tely = vracemosely disposed along the virgate branches; involucre m. high, the phyl- ae em, terete, ascending, okie tipped processes, Serche otis: ray- pil 2; ‘43. 5m A ant in hard- — valley and foothill soils, Upper Sonoran Zone; Central Valley from Glenn County to Fresno County, the Sierran foo ge and the Inner Coast Ranges (Lake, Napa, Santa Clara, and San Ben ae). California. Type locality: : “California, probably on the Sacramento.”’ Collected by Fremont. pore ‘Ho locarpha virgata subsp. elongata Keck, Aliso 4: 111. 1958. Stems slender, 5-12 dm. high, multibranched, the ultimate branchlets gracefully curving, elongated, the heads more scattered on less leafy peduncles up to 15 cm. long. Southw ceteue San Diego County, California. Type locality: San Diego. July—Nov. SUNFLOWER FAMILY 185 3. Holocarpha obcénica (Clausen & Keck) Keck. San Joaquin Tarweed. Fig. 5299. Hemixonts ebcontce ee & Keck, Madrofio 3: 7. 1935. k, loc. cit. Holocene obconica Keck, Aliso 4: 111. 1958. Stem 1.5-8(-12) dm. high, “teh is shaft replaced by several divaricate branches below the middle, the branches paniculately eae o forma sag nse twiggery ; herbage strongly resinous and odorous. Leaves like Greats sine ara e basal more hispid and the omiduchlar often wel spaced; heads paniculate on long s fr ‘pene ict invol 4-5 mm. hi ne i bearing ener few ahahrt psn parser 3 the termi inal one less prominent than in virgata, the surface other glabrous or minutely glandular, carele more or less hispid but never perubulent, often “fai ling ee eevee completely pe a ac - In open fields and hillsid ne; western "Prana County and Siren’ Foothills “OF —, Pe Tular Alameda iene acne Re nner South Coast Ranges from Se arcgscee County to re Counties, California. Type locality: Tesla, Holocarpha obco subsp. autu s Keck, Aliso 4: 112. 1958. Central shaft poe homie by short sterile ate fin poe with slang like ee ves and progr essively shorter from the base u rd, so that before anthesis the plants form tapering green columns; ean bap an exceptionally narrow; Renan sies branches rela- tively short. On arid plains, Upper Sonoran Zone; orth and east of Mount Diablo, Contra Costa County, Cali- fornia. Type locality: west of Beece, Sept.— 52: H alyculata 5297. Holocarpha ornare 5296. Hemizonia clevelandii 5298. Holocarpha vir 186 COMPOSITAE 4. Holocarpha heermannii (Greene) Keck. Heermann’s Tarweed. Fig. 5300. Hemizonia heermannii nee Bull. Torrey Club 9: 15. 1882 Cenanare heerucamnss Greene, Fl. Fran. 425. 1897. annit Jepson, Man. FI. Pl. Calif. 1090. 1925. Holocarpha sical Sock Aliso 4: 58. Stem 2-12 dm. high, simple rah . big ad few-branched above to diffusely branchea throughout, pilose or hispid below, cine above, with interspersed glandular pubescence ; herbage strongly resinous be = a 3 a = = 8 n a = & wn oa ba co oO ba} o > iy 3 > ee wn oO ° mh < iS) a) = a any >| a y= = es 3s B > o oO 5 n oO a 2 0 hese al tim i page areas, Mendocino County to San Mateo County and also sige some San tan “Sbispo 25 vane California. Type Beet h ocean bluffs at Dillons Beach, Marin County. pestis he freely with B. macrantha var. pauciaris- unty. Bae ria macrantha var. pauciaristata A. ae Proc r, Acad. 19: 21. 1883. Plants more or less Mage sented from heveinlal roots, Pam stems ascending, 10—20 c Ha leaty at aed base, ee leaves on the upper stems short than the internodes, mostly rounded at the apex, febbaeenae » n B. macrantha var. macrantha; rays about saa ing t ie ehy lave in Tenth we sap che shorter, scarcely emarginate; cchened a in the preceding taxa, the pappus of 1,2, or 3 awns the sometimes very short. Bluffs or coastal terraces, Curry County, Oregon, south to Sono oma Cont : rie oraias Intergrading with var. thalassophila and also with B. bakers The speci- mens from Curry County often are linear-leaved. Type localit ty: Sa aidealins County, California. Collected by Bolander and iby Prin gle. The tax. macrantha, B. bakeri, B. chrysostoma subsp. hirsutula, ion B. chrysostoma subsp. chrysostoma occur on the coastal s p from Curry County, Oregon, to San Luis Obis eComty and the Channel Islands, California. These highly variable Moe are Saas related morphologically, all havin to ~ greater or lesser degree ag Beane saceneaer marginal, ciliate hairs at least eo the Jee eed baad as well + the & that ar c z bn 1 s oe. G3 pappu i 1-5 pale brownish awns slightly isin ete at the he The ph sina aaa are subclavate in outline and somewhat com- pressed, glabrous or more or less covered with yellowish, selnted papillae. Occasional individuals also are found rio seen to be hybrids b hg sy oe another of the preceding taxa and the maritime representatives of B. minor 2. Baeria bakeri J. T. Howell. Baker’s Baeria. Fig. 5349, Beers sale T. Howell, sans s West. Bot. ue ‘ 1932. . bakeri Keck, A Mek 4:1 1958. Erect perennial from ecaies oe roots, stems erect, solitary or sometimes 2, some — Pek 5-40 c ig ore or less pu ubescent with as ecnding or appressed hairs. Hasel leaves 6-10 c . long, tin they (ne Glial nore teed usually not persisting; stem-leaves con- ae F the irae as long as or longer than the basal but reduced in length above, linear, acute at apex, up to 2 m ide, nearly glabrous except for ginal cilia; heads solitary on peduncl 6-15 ng; involucre hemispheric, the phyllaries thin, mm. long, ovate or elliptic, pubes- cent, carinate in age; rays g, elliptic, usually emarginate ex; z flowe 5 mm. long; receptacle about 3 m m. high, conic, muricate; achen mm. on narrowly su beta lavate, somewhat oie hsesaee ‘more or Pies scabrous with minute, yellowish, pointed hairs ; pappus lacking or of 1 or 2 short ool — ssy poe and swales in ecaase: « ors aks reas adjacent to the ocean, Humid Transition Zone; Mendoci nd Sonoma Counties, California. Type loc oe 6 miles south of Point Arena, Mendocino County. lnterpradine with = ee crantha the pot lt peta fheseh of different growth habit, showing to some degree characteristics of B. bakeri, such as narrow leaves or a tendency toward thickened clustered roots, 3. Baeria chryséstoma Fisch. & Mey. Coast Goldfields. Fig. 5350. Baeria chrysostoma Fisch. & Mey. Ind. Sem. Hort. Petrop. 2: se Poh oa 1836. Baeria gracilis var. aristosa A. Git. Proc. Amer. Acad. 19: 21. Baeria aristosa Howell, Fl. N.W. Amer. 354. 190 0. na mir slender, singe reddish-stemmed annual het 5 dm. high, branching from the base and above, the bran ascending and rather sparsel ay arse 0 ape eading and appressed Sens more dena - S belie the heads ves plein 1.5)2: , l- : inear, acute a pex, sparsely hirsute and hir ute-ciliat hae acatne at the ightly hiboteeg clasping base: heads solita n the branches, the peduncles 3-10 cm. lo involucres y hemispheric ; phyllaries 8-12, ovate-lanceolate or oblong-ovate, 5-9 mm. ong, rather thin, midvein not 9: eae t hat phyllaries carinate below in age, more or less hirsute; receptacle mm. high, conical, wl he is ee -10 mm. lon , rather narrowl blon sometimes roddieh:scined : ‘ak aces 2-2.5 mm. : * “ and SUNFLOWER FAMILY 215 mewhat rounded above, smooth or with minute yellowish papillae or coarse hairs; pappus completely i rah or 2-7 clear or brownish, slender, subulate awns of variable lengths, often as long as the achen Aerie and oe eal Upper Sonoran and Humid Transition Zones; Curry County, Oregon, southward mostly on the c and in the Outer Coast Ranges to Santa Barbara County, geo tm — extending into Inner Coast Manges. hone it merges with B. eeeeeottoué subsp. graci is and arch May.” other named form ra Baeria chrysostoma subsp. niveatatit prpotes Ferris, Fcgalte Dudl ey Her Es 958. (Lasthenta hirsutula aust at, Bay Reg. 206. 1894; Bato ee Greene, we tie 438. iioy j x its characteristic form 2.5-10 cm. tall, rather stout, muc sid base mea ee width of the nts sometimes greater than the height, more Beis a ret pi ee rt- “5 then h sone t and also more ne ee : str rh ed leaves 10-20 mm. long, oblong- gierulate or Kasae- erty rounded at oe Qs very rarely with 1 minu spreading lobes; phyllaries about 5 m at ng, usually broader than those of the preceding —. ettaditen parinate in age, the wa evident i oe wr out 5 mm. long, typically broadly oblong; achenes as in B. chrysostoma subsp. chrysostoma, smooth or with state coarse, yellowish "isits, epappose or with 2-4 pa or brownish, ies er, subulate awns as long as the disk-flowers. Coastal terraces and dunes Ed among rocks, from Mendocino County, California, to San ios Obispo Ton unty and the Channel Islands, Santa Barbara County. Type locality: not given, but ond to grow “‘from Marin Co. southward.” sibly interbreeding with B. chrysostoma Rovvag tiie ey to with which it grows. Forms occur frequently ave pre usly combine the characteristics of both taxa. On the coast of San Mateo County specimens having large heads and broad leaves but annual roots appear to te intergrades with the B. macrantha complex. aeria chrysostoma yaaa gracilis (DC.) Ferris, contr Dudley Herb. 5: 100. 1958. seaside gracilis DC. Prod. 5: 664. 1836; B. ten a DC., loc.cit.: #. Jong ifolia Nutt. Trans. Amer. Phil. Soc Ve 7: 380. 1841; B. pee Nutt. op. cit. 381: Ss hired Nutt. loc ae a gracilis A. Gray, Proc. Amer. Acad. se 196. 1874: palmeri A. Gray, Bot. Calif. 1: 376. _— 4 “gracti vat. - palenene K- Gray, Proc. Amer. Acad. : 21. 1883; B. clevelandti A. Gray, op. cit ae - B var tener Gray, Syn. Fl. N. Amer. is 326. 1884; palmeri var. clementina A. Gra Ff ya 2. 12; 452. 1886: B chrysostoma var. gracilis H. M. Hall, Univ. Calif. tem nee A 170. 1907; B. ichrssostoma var. graci lis £, nuda all, loc. cit.; B. chrysostoma var. gracilis . tenerrima H Hall, op. cit chrysostoma _ gracilis e ‘paleacea H. M. Hall, loc. itt B, poe s eh pide £; Perit ch H. M. Tot loc . cit.; B. chrysostoma var. gracilis f. crassa H. M. Hall. op. cit. 172; chrysostoma var. gracilis f. curta H. M. Hall, loc. cit.). Slender aaa 1—2 dm. high, in the peer 1 form strigose throughout with some spreading or ascending bee present, more conspicuously pubes: etn on the pales heads, and leaves; leaves 8—15(20 ae jones: w sually about 2 mm. wide toward the base, somewhat ciliate on the lower margins of the leaves but rarely wit h coarser cilia; involu ae hemispheric, the pia liacies e- 12, eats: wih of the usually about 5 mm. long, Bie: ly carinate in age, the midvein fale nent nbd half or more t phyllary; receptacle as i ote r two related taxa; achenes long, n aly tank ar, apacaily truncate (som times rounded in Shiphore is glabrous or minutely strigulose with whitish hairs? pappous-paleae when present 2-5, opaque, whitish (somewhat brownish in older herbarium spec ss about the length vl bos corollas, typically a small ovate-lanceolate eale attenuate into a long slender awn Bee slopes and plain on County, Ore a. south through the Sacramento and San Joaquin ts and 5 "foat ills to Ren erat Caiiforni ia and northern Lower California and eaitwardt through the aa s to central Arizona. Type locality: California. Collected by Douglas, presumably near pied 7 Goldfie Very abundant and m widespread ate “the other members of the B. omplex. Variable but characterized princip pally by nth more or less den rad strigose pubescence, the ie an gore “head and the well- defined pappus-scales tebmaanted by attenuate aw The pappus-paleae vary cqosiderebiy in different areas For a discussion of this variation see H. M. Hall (Univ. Calif, Pub. Bot. “3: 170-172. 1907). 4. Baeria leptalea (A. Gray) A. Gray. Salinas Valley Baeria. Fig. 5351. pact tests talea A. Gray, Proc. Amer. Acad. 6: 546. 1865. talea A. Gray, Syn. Fl. N. Amer. 12: 325. 1884. Tote slender, erect annual 7-15 cm. high, mostly simple-stemmed, glabrous ed pate Cai pubescent on pedunc d Jil, ‘Leaves 3-10 mm. ong, entire, filiform or nearly so; involucres campanulate to nearly turbinate; phyllaries 4-6, rathe broadly oblong, nag mm long. thin, ciiia an y pg rous, ob saeliiane 7 - e base; rays ul scurely e; rays 3-4 mm. oe ae notched at the apex; lobes of the disk-corollas narro angular ; anther-append- s 0.5 mm. or more long, filiform or et so; receptacle ss we eg spar than the phyllaries ; 5348. Baeria macrantha 216 COMPOSITAE achenes 2 mm. long, brown, glabrous or with minute hairs toward the apex; pappus of 1-4 pale or brownish bristles tapering from a somewhat broadened base, the bristles about equaling ee disk-corollas Fields Siidk slopes, Upper Sonoran Zone; of rather rare occurrence in interior Monterey ood pe suis chispo oo. Californ nia (Brewer, Brasseces. Alone. Ornduff). Type locality: Nacimiento River, Mon y Coun April. Und - adverse growing conditions depauperate specimens of B. chrysostoma subsp. gracilis have al Rowers heads and nearly subulate receptacles. The plants are strigose and have phyllaries that as strongly c ate and lack the suilitl Sees anther appendages of B, leptalea. More difficult to separate from B. leptalea are "aberrant, dwarfed, Branly, Ooi gees mens from the Inner ‘North Coast Ranges of the awned forms or. 8. sostoma subsp. chryso thope examined, however, has the diagnostic characters of subulate ese foe and subfiliform ae ta aubeadases: 5. Baeria microgléssa (DC.) Greene. Small-rayed Baeria. Fig. 5352. Burrjeee mecrourates DC. Prod. 5: 664. 1836. Greene, Man. Bay Reg. 205. 1894, 1897. a Gre Pentachaeta less Elmer, Bot. Gas: 41: 318. 1906. ak, slender-stemmed, erect or reclining annuals 5-20 cm. hi gh, the stems arising singly or several from the base, rndtty branched above, said Pa pubescent throughout with ascending i y about 2 m i hairs. Leaves thin, linear, entire, 1.5-4 cm. lon , usuall wide; peduncles 1-3 cm. long; involucres cylindric, the phyllaries 3 or 4, ahier ng or narrowly o vate and acute at apex, 6.5-8 mm. long, ciliate- margined, thin and lea fy, cot carinate in age; r ecestaalé subulate; ray- OW 3 ys mi r wanting; disk-flowe sually less than 10, u m. long, the throat broadly funnelform; achenes black, nearly 1 r; long, minutely and sparsely hispidulous; es aleae 2-4, surpassing the disk-flowers, narrowly lanceolate and attenuate into an awn, or pappus- a lacking in some flowers of the he ads. Grassy shaded uieek: or_ among rocks oulders, Sonoran Zones; Tehama ey: California, southward in the Paces Ranges to San Diego County; cae in the Death yale ah Soe in Inyo County and the western edges ye deserts from Kern County to Riverside County. Type loca : California. tad by Douglas. March- 6. Baeria débilis Greene ex A. Gray. Greene’s Baeria. Fig. 5353. Baeria debilis Greene ex A. Gray, Syn, Fl. N. Amer. 12: 325, 1884 Lax-stemmed annual, erect or Beas ng, 5-30 cm. high, the stems arising singly or several from the base, simple or with few ascending Fianches aarets soft-villous with microscopically septate ascending hairs, more Scale so below the heads. Leaves flaccid, linear, 1-6(8) cm. long, 1-5 mm. wide, entire, more rarely with 1-3 lobes; peduncles 2 ; involucre campanulate to broadly turbinate, the phyllaries 5, about 5 m ong, broadly elliptic to narrowly ovate, thin, veins not prominent in age, keeled at the extreme base; receptacle subulate, 1g about 1 mm. wid base; ri wers yellow or white, the long, narrowly pat and usually toothed; Bie Bower 15 a u ng, tube slender, throat cam- panulate; achenes narrow, mm. long, hispidulous ; pappus-paleae 2-4, white, ovate-lanceo- late and tapering to a slender awn, a little meee than the disk- Aowers: or pappus- -paleae lacking. Grassy woodlands or among boulders, Upper Sonoran Zone; lower foothills of = Sierra Behe lr and adjacent eee Mariposa County, Californias sot to Kern County. Ty ype locality: plains of Fresno County. March— 7. Baeria minor vee ) Ferris. Woolly Baeria or Goldfields. Fig. 5354. Monolopia minor DC. Prod. 6: Dichaeta tenella Nutt. Trans, ‘Amer. 8 Soc. 0), 7: 383: 1841. + 2 " ae u Baeria uliginosa A. Gray, Sin Mek — 9: 197. 1874. Baeria uliginosa var. icuciha: A. Gray, loc. ¢ Be aeria wht iginosa var. tenera A. Gray, op. fy 19: 22. 1883. een : F, 18 = ey Berindka li. he Ryd r. Fl. 34 1915. Baeria minor Ferris, ets ome Herb. ‘ eek 1955. Annuals 1-3 dm. high, erect and simple-stemmed below or diffusely branched from the base or even prostrate; stems weak and somewhat poorer ds: or firm and strictly erect in drier situa- tions; herbage very sparsely villous to dense y_lanate- villous with long, septate, often twisted hai y or narrowly li undivided portion of the mes te when mature, the papillae ring the achenes scarcely fused toward the ee “ the reac and appearing as distinct pedicels ; ligules of ray-flowers usually oblong and emarginate or toothed at the apex, mm. long; isk-flowers many, tu As ndular, throat catiinarGEahe oS spiel pie about 425 mm. long, gay eae smoot th; pappus completely lackin, y- and disk-achenes or of 2-4 awns nearly as lon ng as 'the corolla and (2)4-6 tru — feeb ee these. rorane and ges completely cleft, particularly on the ray-achenes wher y be vestigial and awns only p: In damp soil near ap ctaly Paso € areas, or is ene in fields and by roadsides, Sonoran and Transi Zones; common along the coast erage areas County to Santa Barbara County, California, and eastward a 217 5353. Baeria debilis 5354. Baeria minor 5355. Baeria platycarpha SUNFLOWER FAMILY loncca 3 5350. Baeria chrysostoma 5351. Baeria leptalea 5352. B 218 COMPOSITAE oast Ranges and Sacramento—San Joaquin Valley to the lower Sierra foothills, where it occurs from Amador fe Soe south to peed County. Type locality: California. Collected by Douglas. March—June. aeria minor subsp. maritima (A. Gray) Ferris, Contr. eed Herb. 4: 334. 1955. (Burri ielia maritima bi Gray, Proc. Amer. Acad. 7: 358. 1868; Baeria maritima A. Gray, op. cit. 9: 196. 1874.) Diffuse succulent nuals,” sparsely soft-villous as in the preceding taxon to aulatale. oa re except for the inflorescence; leaves Srcedia linear, te to sie ge i and laciniately lobed, the rachis Mi to 11 mm. wide; peduncles egg short; — ig and r rate ai in the preceding; ray -flowers few, 3-8, the ligule minute, heat t 2 mm. long, oval long, often eka apex; achenes pate and epappose, or ppd nea and bearin 3S ‘awns alter- sating mith the Pare eds. se 8 paleae. Occasional week the coast in seepage areas, from aries Columbia to Monterey Count Californ pe lo cality : Farallone Islands, Sea Prateoae Conaky Californi This taxon differs from B. eat mi eae minor in the few and minute rays and the tendency to a very short peduncles on the spring plants 8. Baeria platycarpha (A. Gray) A. Gray. Alkali Goldfields. Fig. 5355. pgs 3 aapeieagy: ei Gray in Torr. Bot. Mex. Bound. 97. 1859 Gray, Proc. Amer. Acad. 9: 196 Resins carnosa he e, Bull. Torrey Club 10: 86. 1883. Lasthenia platyc wa Greene

Hey, California, Siaidalaus Paaaty south to pew. and Tulare Counties. Type insnbity: near Lake Tulare, Palace County. Collected by Greene. March— SUNFLOWER FAMILY 221 52. BAHIA Lag. Gen. & Sp. Pl. 30. 1816. Annual or perennial, pubescent but not at all woolly, often gastos’ herbs, sepaio 4 suffrutescent at base but usually herbaceous. Leaves alternate or opposite, entire dissected. Heads terminating the branches, yellow, many- aed radiate or barely dis- coid. Involucre hemispheric. Phyllaries in lor series, subequal, herbaceous, flat, spread ing in age and sometimes reflexed. Receptacle flat. Ray-flowers when present pistillate, chenes narrowly obpyramidal, 4-angled. Pappus-paleae with a callous base and thickened midrib as long as or shorter than the paleae, or paleae completely lacking. [Named in honor of Juan Francisco Bahi, a Spanish botanist. A genus of about 15 species, natives of southwestern United States, Mexico, and western South America. Type species, Bahia ambrosioides Lag. Pappus wanting; stout biennials or perennials. 7 B. dissecta Pappus present; slender annuals. . B. neomexicana, L pa dissécta (A. Gray) Britt. Dissected Bahia. Fig. 5361. Amauria ? dissecta A. Gray, Mem. Amer. Acad. II. 4: 104. 1849. Viliadit ter eed te Gray, Smiths, Contr. 5°; 96. 1853. Bahia chrysanthemoides A. Gray, Proc. Amer. Acad. 19: 28. 1883. Te baie a Bri tt. Trans. N.Y. Acad. 8: 68. 1889. Bie nnial or short- lived Seeeitiien nate 3- 8 dm. high; stems striate, reddish, puberulent and etindelac. simple to the loosely branching inflorescence. Leaves petioled, the basal leaves with 5361. Bahia 222 COMPOSITAE longer petioles than the upper stem- ares: ——. parted into oblong or linear, blunt lobes, 2-7 cm. long, puberulent; peduncles 2-4 cm. long, enlarged below the head, den sely beset with gianitinpsd hairs ; involucre 5-6 m vochiak. pre Swics Bf: Eat glandular- notes paxtlarics trowly obovate, abruptly Ban above; rays rrowly cuneate, 3-c mm. long ; achenes Rack: striate Sal 4-angled, glabrous or as by glandular- a Seana eanectalie above, epappese Bins places along streams and roadsides, Arid Transjtion Zone; San Bernardino and Santa Rosa Moun- tains, California a, and mountains of northern Lower sce gt east — southern shee en Colorado, New Mexico, and Chihuahua. Type locality: a few miles f Mora River, New M . Aug.—Oct 2. Bahia neomexicana A. Gray. New Mexico Bahia. Fig. 5362. hi A. Gray, Mem. Amer. Acad. IT. 4: 96. 1849. Amblyopappus neo-mexicanus A, Gray in Torr. Pacif. R. Rep. 4: 106, 1857. ahi tc 883. Cephalobembix neomexicana Rydb. N. Amer. FI. 34: Slender annuals 1-2 dm. high, single- Sauda below or prancing from the base, the stems often reddish, sparsely pubescen . Rae scending or. appressed a or — te, ine ig ol ar- puberulent below the heads. Leav saeae often alternate and e m. pressed-punctate and oi hispi lous, usually tripartite into Sean: filiform viian : tes 3-flowered or more, borne on the See: of ne ranchlets on peduge 4 .5-3 cm. long; involucre obconic; phyllaries 5-8, eRe te, ok al in width, about 6 mm. long, herbaceous and often pur- plish-tinged on the margin, Rem “panctate or glandular- eercralnt and ae sparsely hirsute; ray-flowers none; disk- sie es pale yellow or whitish; achenes glandular, very narrowly ob- dbareseoee about 3-6 mm. long, sparsely strigose, Pinan white: hirsute at base with ascending hairs ; pappus-paleae 78. equa al, about 1. 26, se tg obtuse at apex, hyaline but callous- hsekevet at the base, this often bearing a ere ks Sandy washes and slopes, Arid Transition Zone; tee - oe = from Clark Mountain, eastern San Bernardino County ( (Roos 4955); Ari — =a and norther r California to Colorado, New Mexico, and Chihuahua. Type locality: Santa Fe, New M . Aug.—Oct 53. MONOLOPIA DC. Prod. 6: 74. 1837. White-lanate, erect, spring annuals simple or much branched at or above the base. Lower leaves opposite and narrowed to a petioliform base, the upper alternate, sessile, and reduced upward. Heads solitary, terminating the branches. Involucres hemispheric or i c ; bilabiate, the posterior lobe minute, the tube glandular-hispidulous. Disk-flowers yellow, few to many, hermaphrodite, fertile, 5-lobed, the lobes hairy within, the tube glandular- hispidulous. Stamens 5, the anthers rounded below, bearing apical ovate appendages a: Style-branches of ray-flowers slender, obtuse, of the disk-flowers stoutish, obtuse o sub- or quadratish, epappose. {From the Greek meaning single and husk, alluding to the bracts of the involucre. ] A genus of 4 species, all natives of California. Type species. Monolopia major DC. Phyvyll j disti t to hase Limb of ray-flowers rounded. apically oo or denticulate; disk-achenes essentially as wide as thick Bay pe strongly pete anaes plants of the Coast Ranges, Contra Costa and San Mateo ot to San 2 Os SEDO. Cou 1: cilens. Ped ts o n San — Valley and adincess Hoke 2. ee, Limb of the ray- Mowers truncate, i eat dent o. M. as Phyllaries et one-half their length into a tobesd 6 4. M. major. 1. Monolopia gracilens S a oo Monolopia. Fig. 5363. Mansiony — - Gray, Proc. Amer. Aca racilens J. F. Macbride, oe Stns ait No. 56: 49. 1918. Steins 1-4 ee high, simple below or gpa gore branching from the base, the branches of midstem divaricate es prone ‘ite. lanate and somewhat deciduous in age. s d lower leaves crowded, narrowly or broadly Rance dentate, 3-7 cm. long, tardily withering; the upper broadly langeolatt. dentate to nearl ire, acute or acuminate, 2-10 long, mostly reduce oming bract-like at the inflorescence; heads subcorymbose on divergently ascend- ing peer tes, these 2.5-12 cm. long; phyllaries rae free, Bisck: lanate, narrowly ve broadly ovate, 5-6 mm. long; ra iti reve wers ret yellow, with 7-11 greenish veins, entire shallowly emarginate at apex; disk- flowers yellow; achenes black or dark brown, essen entially Pabroat, the ray-achenes slightly convex and somewhat dorsally carinate, a little shorter than the disk- sickoces es, ee 2 m aa Sonoran and Transition Zones; central California from Contra Store Gate ny. south through the Guter Coast Ranges to San Luis Obispo County. ‘Type locality: New Almaden, SUNFLOWER FAMILY 223 2. Monolopia stricta Crum. Crum’s Monolopia. Fig. 5364. Monolopia stricta Crum, Madrojio 5: 258. 1940. Stems 1-6 dm. high, simple or much branched from the base and above, all the branches strictly ascending, white-lanate to floccose and becoming glabrate. Basal and lower — oblanceolate, entire, 3.5-8 cm. long and soon withering, upper stem-leaves atone eh to oblon lanceolate, apex obtuse or acutish, 3.5-6 cm. long, usually entire, equaling or shorter than a internodes ; heads on the lower as well as upper branches, the peduncles about 3-5 cm. long; phyl- laries about 8, free, ovate or ovate-lanceolate, usually cg pi at the apex, mm. long; ray-flowers about t 8 8-10-nerved, oblong, rounded at apex and entire or minutely oy aging 4-15 mm. long;. disk-flowers yellow; ee Hai densely grayish- Seas , 2.5 to nearly 3 m long, the ray- -achenes obcompressed, the disk-achenes not obcompre essed. lains or open slopes in the foothills, Lower Sor n Zone; Inner Coast Ran San Benito County, Cali- pes, to on * Ane. and eastward in the San g Pons Vales ps Talate Coane, x, amt ie in Lost Hills, Kern ounty. [e x 5 page as Sper Nutt. Common Monolopia. Fig. 5365. ohamestd te lanceolata Nutt. Proc. Acad. Phi 848. ‘tiie Gray, We nthe: ve 384. 1876. Ste a we high, simple or branched from the base and above, the branches diffusely edeadtug. densely wipes fauatn and becoming somewhat glabrate. Basal and lower cauline leaves 5362, Bahia neomexicana 5364. Monolopia stricta 5363. Monolopia gracilens 5365. Monolopia lanceolata 224 COMPOSITAE broadly lanceolate, obtuse at apex, 1.5-10 cm. long, soon pene go mpper stem-leaves linear to broadly lanceolate, obtuse or acute, asaalls entire, 3-1 long, stly much shorter than the internodes; heads terminating the uppe r branches, the dincegent ‘pedlincles 1-13 cm. long; phyllaries about 8, free or 2 or 3 rarely united: lanceolate to ovate- or r ombic-lanceolate, 5-11 mm ate ata xs rv aoe or broadly oblong, a at apex eon Pao age 9-21 mm. long, the pr aoa yellow Se mostly gray-strigos long, the ray-achenes obcompressed, convex ie somewhat fattened and often pubedesiakt dette: 'the disk-achenes obcompressed me carinate dorsally wld 3 rally. Open slopes eo valleys, Upper and Lower Sonoran Zones; Inner Coast Ra —— , San Joaquin County t Luis Obispo rid ged the mountains rs southern California in s Hipexeids and Los hy bae Counties tha. the —— edges San Joa a Valley from ae fomety to the Tehachapi region; also ees ene western edge of the Mjave “hee. ype locali ity: Los Ange March—May. Closely related to M. 4. Monolopia major DC. Cupped Monolopia. Fig. 5366. Monolopia major DC. Prod. 6: 74. 1837. Stems about 1-5 dm. high, simple or branched from the base and above, the branches dif- nel sa 0 densely white-lanate and becoming somewhat glabrate. Basal and lower cauline leaves oblanceolate, eae entate to heck obtuse, acute or acuminate, from about 1-10 nny one the upper leaves about 1-10 c ong, the uppermost reduced in size, the margins usually dentate, sometimes subentire ; fous “teheinalies the branches, the div i iad “Pepin i u " 8-13 Ope dv ag a Lower Sonoran Zon Bes Coast Ranges from Tehama Come rent fers to es Each and the linc eeokern ac. of az Great Valley. Type locality: California. Collected by Douglas. March-May 54. ERIOPHYLLUM* Lag. Gen. & Sp. Pl. 28. 1816. rubby or herbaceous perennials or annuals, the herbage tomentose or floccose. Leaves sooneintl alternate, poe lobed, toothed, or divided, more often entire in the annual ci annual species. Phyllaries in 1 series, firm, mostly carinate and somewhat concave on cage inner surface, permanently erect, the tip ete soften reflexed, distinct or r¥oages united at the base (more in one species). Receptacle convex or conic to nearly modified in E. mohavense), naked (with a few Cenlixie scales in one species and ee processes at- tached basally to the phyllaries in another). Ray-flowers usually few, rarely wanting, pis- tillate, fertile, the ligules yellow, oval to oblong, toothed or lobed at the apex. Disk-flowers yellow, perfect, narrowly pubescent or glabrous. Anthers obtuse at base, bearing angen ages above. Style-branches flattened, the apex obtuse or deltoid, or cuneate in E. wallac n lanosum, Achene — -nerved or -angled or 5-nerved “ some ual species, som S-s what compressed. Papp arious, nerveless, usually ppu S erose or fimbriate paleae somietiane’ slender and awit rarely epappose. [From Greek words oe wool and leaf. A genus of about 13 often variable species, all native of western North America. Type species, Eriophyllum troliifoltum Lag. or complete synonymy of the perennial species, see Constance, Univ. Calif. Pub. Bot. 18: 69-136. 1937. Perennials hat sk ti 0(15) dm. high. Heads 1.5-3 em. broad, shoe y or several o; = aie ncles Plants shrubby only at base; rays 8-20 mm. long (less in some varieties) ; of beet oi gree . E. lana Plants shrubby throughout; rays 6-8 mm. long; central Inner Coast Ranges in Calitopni ee B. jepsonit. Heads 1.5 cm. or less broad, 1 ile cl t, pedunculate, b cluste shor Ultimate ies: hein infloresserices slender; saievs 6. phd am sites = yer ee ates. onfe: eer branches bearing inflorescences stout; phyllaries 8-12, scarcely Sn sear ear es Dacitione Leaves ‘labrous above, tomentose beneath, entire or pinnately partéd into i lobes; r 4 3-5 m . E. staecha 8 hinoaea Ae shaaity tomentose on both surfaces, bipinnatifid into many lobes; rays 2 Fa long. 5; evinit. Small herbaceous annuals 2-20 cm. high (sometimes higher in E. congdonti). Pappus-paleae of unequal oa lanceolate or awn-tipped paleae alternating with short —_ paleae. ha yellow; plants of the Sierra Nevada foothi ongdonti. s white or pink-tinged; plants of the desert resicha. 9. E. coe * 'T. + “4% -s. I + +, a CL Cc S| here SUNFLOWER FAMILY 225 Pappus-paleae all alike and of equal length or pap lacking. eads re -flowered with never less than 9 or 10; rays present (except in E. pringle eceptacle baby or less dome-shaped or conical; peduncles at least 1 cm. long ‘asad shorter in ee vallacei), generally yer much longer. Rays scarcely exceeding the disk; plants of the higher Sierra Nevada. 6. E. nubigenum. Rays —— oe _ ee — of desert ranges, and desert slopes of the Sierra Ne- and t hap ae opens branched, rae or several from the base; pappus-paleae, when present, hya- lin ee ee Stems staid from the base but little branched; pappus-paleae, when present, o 10:2, wonilec? ReckoMirle ae or nearly so; heads in leafy clusters sessile or the peduncles usually ane more than ng. oc fl Hee multicaule. Ray-flowers ssi = E. pringlei. Heads few-flowered with never more than 4; rays absent. . E. mohavense. 1. Eriophyllum lanatum Robes Forbes. Common Woolly-sunflower or sce Fig. 5367. Actinella lanata Pursh, Fl. Amer. Sept. 560 Eriophyllum gas eae Doug]. ex Lindl. Bet er Pa he 1167. 1828. Eriophyllum lan Forbes, Hort. Wobur Bahia liacanhera DC. Prod. 5: 2 Loosely tomentose perennial, erect or decumbent from a ares base, the dans stems (1)2-6 dm. high, the tomentum tending to be de ep in age. Leav 2-6 cm. long, p nently white- woolly bel labrate or glabrous above, variable in size at outline, the oe sp pas or te nat corymbose on ped s cm. long; involucre hemispheric, 8-10 mm. high, 10-12 mm. broad; phyllaries ge loosely fidcsese: or glabr bi fire to pate lanceolate, strongly tag distinct to oe base but with overlapping mar y:flowers 8-12, yellow Dis iB . long; disk-flow S34 mm. sed with a qinmalie any pi : shee enes buy aioe sbioaaeh in outline, narrowe as ard th angled, glabrous, the pappus- “paleae 4-10, denatly obtuse and erose, sometimes Bier taaccolate aie alee. not exceeding 2 m = NORE In thickets and open dry places, Transition ommon west of the Cascade Mountains from British Co- lumbia to southern Oregon a adjacent Del Réce Ca ie my California, and on the groans saa in ~ seatoome of the Columbia River in southeastern hae egg iB and northeastern Oregon eastward to western Mont. Type locality: banks of the Ki eaekie® as —Jul a 8 NE Se lanatum var. s hilicaotoes (DC.) Jepson, Man. FI. Pl. Calif. gs bio (Bahia ie aga DC: Prod; a : boric wits aren ar’ Pittonia 3: 1S5.01897 3 2 mag fy tomo * . Mid. Calif. 524. 5 01; £. 9 Elmer, Bot. Gaz ; E. cusickit Eastw. ex Sera e 93. 1915, as a synonym; ss ‘achilaxesdes var. By ie ~ Howell, Leaflets West. Bot. 3: 126. 194) Stems ve A Pp mes r ¢ ong; hig ys 6-9 mm. long lacking in var. pet lia: achenes short, 2.5-3 mm. lone, ‘cuneate-oblong, hispidulous or vislobronset pappus present. vay pt in Oregon from Eid and Harney Counties to rahi tc alifornia and nee tie more commonly in the Calif rnia Coast reap as far south as the Santa Cruz Mountains, and in east- BS SD: though occurring fhe? commonly m M nis 5366 5367. Eriophyllum lanatum 5366. Monolopia major 226 COMPOSITAE Eriophyllum - lanatum var. eo (A. Gray) Jepson, FI. Mid. Calif. 524. 1901. (Bahia lanata Benth. Pl. Hartw. 317. 1849, not B. lanata DC. pers falda seliormeus Kell. Proc. Calif. Acad. 1: 56. 1855; B. lanata var. grandiflora ee Gray’ Bot. Calitsc ts 3 tophyllum caespitosum var. grandiflorum f om a wo Ne cst y “be o a ely so : or lanceolate or laciniately toothed and sometimes pinnatifid; heads large, solitary at the ends o e stems (rarely Ss dm. oa ar side of the Sacramento Valley as rhe south as Mariposa County. Type locality: Sacramento Valley, Callforsie. Collected by Hartw Eriophyllum ar. arachnoideum (Fisch. & Avé.-Lall.) Jepson, Man. Fl. Pl. Calif. 1119. 1925. Aa cre pe rolifoium Lag or & Sp. Pi. 28. fod identity uncertain; Bahta arachnoidea Fisch. & Avé- . Sem ; 9: 63. 1842; B. h. Bot. Sulph. 30. Lf a Rages var. brachypoda % ree ok Cann Ls one 1876; Eriophyllum caeloneban var. latifo lium A, Gra . Amer. Acad. 19: 26. a: in part.) Stems 3-6 dm. high, branching from a woody base and usually aaubere usually densely leafy e 0: ty" y =. Soe 3 =. & ao g- aa zs with 3-5 incised teeth or lobes; heads on eek es 3-10 cm. long; involucres 8- m. high; s 8-10 mm. long achenes 3—4 mm. long, turbinate, glabrous; pappus reduced to a crown of erose Secth, Principally in the folleeas4 belt, Del i ad Pa pig De icansigt aaa i south to Monterey County. Type locality: California, presumably from the area aroun Eriophyllum lanatum var. créceum (Greene) Jepson, Man. Fl. =. palit, it. 1925. LBrigptaiem caespi- tosum var. ini: haere A. Gray, Soa Amer. Acad. 19: 26. 1883, in eum Greene, Erythea abe 1895.) Ste -5-6 dm. high, simple, leafy thro ughout to the ih 2 te ata et leaves pity ienide green, lightly oceose to ih i age ore coarsely abel or aver i eds ra cng or few at ends of the un ng -— peduncles 3-8 cm. long; involucres 5-8 mm. hi eh; s 8-1 achenes as in the preceding form. In the Arid Tvansition Zone on the western face of Pa y BBiag Nevada, California, from Butte County to Tela County. Type locality: ‘Amador and Calaveras County hills uch resembling E. Janatum var. arach- noideum _ differing principally in the vesture of the eva and to a = degree in the gece Eriophyllum lanatum var. hallii Papeete Proc. Nat. Acad. Sci. 20: 411. 1934. Plants 3—4 dm. hig with siiber Saale stems arising from the leafy geld leaves agonety floccose on both satige Sones tadieed or pinnatifid; heads solitary or few at the ends of the branches slender peduncles 5-12 long; involucres 8-12 mm. high; rays 10-13 mm. long; tube of disk, corollas io differing in this aes from all the other related eee: ene 4-5 mm. long, narrowly oblong, glabrous or anew hat pubescent; pappus present. ro me om the Palo of ro Te e) i eas County, Cali ae Ie type localit aie Tio lan Mer neatu ) Jepson, Man. FI. Calif. 1118. (Bahia Kell. gf ag vane Abad oe 49. a73- "B. infeoriflte A. Gray, as Bere ‘ sai (in Pah 5 1876, nip integrifoti DC. 1836; grein ny chy ysanthum yd 915; E. bolan Rydb. Ol yk raria Ry db. t. 93.) Plants 2-4 dm. hi ae fs go vt wage spreading oo the onl ans se, tomen tome or Rocka iarcaurkour: tenes are tcothed or pode Nas so above; need solitary on the branches, the peduncles 5-10 cm. long; involucres 8-10 mm. high; rays "8-10 a long; achenes 3-4 mm. long, — pre labrous sie hairy fs ppus evident but etn sonal In the Sierra Nevada Cal ifornia, from southeastern Lass Pian 8 to Placer County and in adjacent Washoe Sonate, Nevada. Tron Seaatiay : re wiles ngs patent var. integrifélium (Hook.) Smiley, Univ. Calif. Pub. Bot. 9: 378. 1921. (Tricho- phyllum + zgrifoli m Hook. Fl. Bor. Amer. 1: 316. 1833; T. multiforum Nutt. Jonrt. Acad. Phila. 7: 35. 1834; Bahie recite Hook = Arn. Bot. Beechey 353, 1840; E riophyllum caespitosum var. integrifolium A Gray, roc. A d : og 3, in p E. caespitosum var. leucophyllum A. Gray, ioe cit., no ta leuco- phylla DC. 1836; E. w t A. Gray, loc. cit.; E. lutescens Rydb Amer. Fl. 34: 87. 1915; E. monoense Rydb. loc. cit,; E. ssichotordue Rydb. op. cit. 89; E. nevadense Gandoger, Bull. Soc. Bot. Fr. 65: 40. 1918.) Stems 1-2 dm eee: many, erect or decumbent from a woody b r short caudex, the ge persistently tomentose, ca or floccose; er lasues entire or 3—5-toothed or -Iobed at the apex, the stem-leaves in cised or phisatind: sie: into 3 divisions; heads solitary or few on pedu 3-10 ; involucres 6-8 m high 6-10 long henes mostly hairy, clavate, the pappus present but extremely variable in_ this In the ific Northwest east of the Cascade Mountains in Oregon and Washington to ¢ tana and Wyoming; in California it occurs bee ed pr besa counti t rd in the Sierra Nevada, along the crest and on the eastern face, as far south as Tulare and Inyo Counties and also on the adjacent higher ranges in Ss aa Type locality: sources. be the "Columbia cen Eri lum lanatum var. obovatum (Greene) H. M. Hall, Univ. Calif. Pub. Bot. 186. 1907. (Erio- PATE rigs cocipitosum var. integrifolium A. Gray, Proc. Amer. Acad. 19: 26. 1883, in part; ai obovatum Greene, Erythea 1895; E. brachylepis Rydb. N. Amer. Fl. 34: 88. 1915.) Stems few to several, 2-4 dm. high, i i otal facts Us at the woody base, herbage persistently tomentose throughout; leaves entire or coarsely serrate toward the apex, narrowly or broadly oblanceolate, the lower leaves bsg heads mostly solitary on peduncles 3-10 cm. long; involucre 7-12 mm. high, campanulate; care * . long; achenes 2-3 mm. long, cuneate-oblong, glabrous; pappus present. In a tn Sierra Nevad d Greenhorn Range in Tulare County and adjacent a County and in the San > Mwantatnen og eather ‘Caltoriia. Type locality: San are be no Mou Eriophyllum footie var. lencecth teen Howell) Jepson, ig Ra Pl. Calif. 1118. 1925. (Eriophyllum lauciolabens "Howell, Fl. N.W. Amer. 355. 19 ixfordit Eastw ‘Calif. Acad. TV. 20: 158. 1931.) Stems several, 2-4 high. persistently woolly, Soak: = Mt entire or ae serrate, S waalic 4 m otk surfaces: heads solitary to om he peduncles stout and somewhat cee Reto ned —— involucres a . high, m more or less hemispheric; rays 7-10 mm. long; achenes hairy No Si akiy iyou_region of Curry, Josephine and Tackson Counties, Oregon, south to Aap and “Hambotat “Counties, ‘California, Type local- ag Bi: kso a County. Oregon. One of the ears ‘distinc and less variable taxa om comple. en for 2. Eriophyllum jepsénii Greene. Jepson’s Eriophyllum. Fig. 5368. Eriophyllum jepsonii rons Pittonia 2: 165. 1891. Plants woody to well above the base, 5-8 dm. high, the stems closely ang Faian pia the internodes long. "Leaves alternate, 3-6 cm. long, floccose and glabrate above, pinnatifid into 5-7 se : branches puberulent or hispid tube aboot the length of the funnelform throat; a achenes n arrowly clavate, -angled, more or less app pocngpea de pappus-paleae unequal, the narrowly anapdiate paleae aitogsic. with the Sbloag erose wooded slopes or edges of ras chanatral Upper Sonoran Zone; Motint Diablo, Contra Costa County, Sank the Mount Hi Hamilton eerie San Benito County, California. Type locality: between Arroyo Mocho rat y ce, Valle. Alameda Coun’ unty. April-— et ey h markedly different in its Sta ae this taxon shows close affinities with E. gg? Ae a var. confertifiorum and especially with E. ertiflorum var. laxiflorum. In the southern part of the Mount Hamilton SUNFLOWER FAMILY 227 manatee aero Clara County, and in San cme po eee intermediates are to be found in which the inflorescence nsely flowered and the peduncles are shor yllum tanacetiflbrum Greene, neti * 21. 1889. (Eriophyllum confertiforum var. tanacetifolium Teeunnn 4784 “a Pl. ae whip oi ue ie FO: shrubby, 3-6 dm. high, with erect, pita simple, leafy stems, tomentum dense annos Pathan nt. like those of E. jepsonit but m wy orter; inflorescence of close corymbose Aber wit ith 2 —10 Ress. bee! bag sessile or with peduncles to 2. Se long; involucre densely tomentose, 5—7 mm. high, ea aggre the Bg rab overlapping, ovate, broadly pe to eben cariaate at the — “_ sist Pi lacking, es AF rs long, ‘glandu lar-hairy, tube roat nar- wly achenes TAs - ti wey to i ar iiabcoan pappus-paleae of avandia “icaath. Foot- hills oe ee — "Nevadin California, in Re and Mariposa Counties. ype locality: between Sheep Ranch and Murphy’s, Calaveras Count Eriophyllum latilébum Redb. N. Amer. FI. 1914. Plants perennial, stems leafy, 3-4 dm. high; leaves Gens ic to obovate in outline, deeply 3-lobed, a divisions often toothed or lobed, prin abors, tommentone heneath; heads 10 or more in loose clusters on a gana 2.5 em. lon = involuc ~— a high, 7-10 mm. broad; phyllaries 6-8, ovate and somewhat overlapping, ra 8 igh ia entire dala oblo ong; tubes - disk. and ray-flowers glandular; achenes 3 mm. long, pi iat or very aparsely hispelcious. the pappus- agg of texted laciniate. Ay very restricted endemic found in San Mateo County in the hills west of San Mate where it ¢ on slopes covered with mixed California live oak, buckeye, and shrubs. 3. Eriophyllum confertiflorum (DC.) A. Gray. Yellow Yarrow. Fig. 5369. Bahia confertiflora DC. Prod. 5: 657. 1836 Eriophyllum csnteriiel ds. Gray, Prod. Amer. Acad. 19: Eriophyllum confertifiorum var. discotdeum Greene, Man. Bay ag tenis 207. 1894, Eriophyllum cheiranthoides Rydb. N. Amer. Fl. 34: 95. 1915. Erioph a biternatum Rydb. op. cit. < ir i i Frio ph vifiie 4 crucigerum Rydb. loc. cit. bhai plants, the tomentum more or less a ee Sieg stems ge below and branch- ing, the upper stems erect and slender, 2-6 dm. high. Leaves 1-4 cm. long, the upper surface raat ag psa Se , once- or twice-pinnatifid into lends Saaer lobes (s papers. a earlier i imes cleft ; s in clusters at the ends ranches 3- : —- broadly elliptic, carinate, strongly overlapping; piace mm. long, few, metimes lacking, the rays oval or gis J isk-flowers 2-3 mm. long, glandular-hirsute or nuheralen t, the tube shorter than the narrowly campanulate throat; achenes linear-clavate, Poms hisnididous or sparsely pr peas: -paleae about 1 mm. long, few to several, = open chaparral and rocky places, mostly Upper Sonoran Zone; Outer and Inner Coast Ranges from Mendo- cino and Tehama Countijes south to southern Pky Sues and northern Lower Califoriia, and in the foothills of the Sierra Nevada from Calaveras County to Tu unty and the mountain ranges of southern California. Type locality : be te Collected by Deans: aoc rig, is As indie ntl ~ ens anee Madd Calif. Pub. Bot. 18: 107-109. 1937), this is a widespread and polymorphic complex in which individuals are seemingly, susceptible to oe modification and the growth forms show i ties have bee sed on such distinct characters as discoid heads range of here condidensd to be svnonyms of the typical form. Two o sect Pid rar show a gar re dis- aww bonne with ~ seonog ae sae a oblong, usually ob- d s long as the divided portion, the internodes shorter than the leaves; heads solitary on the ends ucres 5-6 mm. h scurely toothe apex, mm ; disk-flow ut achenes narrowly obov aid: black, more or les appre see hairy, renee pacer te aie but reed on the flattened saviiees epappose or with ryeatigiat pappus-paleae. On open wooded slopes, Upper Sonoran Zone; slopes of the Sierra Nevada, California, from Butte County to Kern County; also occurring rarely in the Santa Lucia Mountains, Monterey County. Type Incality; “Cala- veras. ™ Collected by Heermann. Aberrant forms occur in the gocthera part of the range. March—-May 5380. Pseudobahia bahiaefolia 378. Eri inglei 5378. Eriophyllum pringlei sennren eae 5379. Eriophyllum mohavense 234 COMPOSITAE 56. SYNTRICHOPAPPUS A. Gray in Torr. Pacif. R. Rep. 4: 106. 1857. Low, usually diffuse annuals. Leaves alternate, sometimes opposite below, entire 3-lobed at apex, narrowed to a petioliform base. Heads many, terminating the beutichtees, short-peduncled. Phyllaries of the involucre few, partly enclosing the ray-achenes, in 1 i J e series, erect or a di n anthesis, spreading after achenes ed. R acle flat, naked. Ray-flowers pistillate, fertile, as many as the phyllaries, the ligules yellow or white and pinkish-tinged, oval and toothed at apex. Disk-flowers perfect, fertile, yellow, tube much shorter than the trumpet-shaped throat sat nly lobes. ue ers obtuse - Bios, ap- pendaged at apex. Style-branches flattened, Achenes narrowly obpyramidal or aie, 5-angled. Pappus none or the paleae aisected into many bar sol cicorhas united at base into a ring. [From the Greek meaning united, hair, and pa A genus of 2 species, natives of the deserts in California, sition Utah, Nevada, a far erie Arizona, Type species, ay whivickotabene fremontit A. Gray. Ligules of the ray-flowers yellow; pappus of numerous bristles united at the base. Ae Ligules of the ray-flowers white or pinkish; epappo as 1. Syntrichopappus freméntii A. Gray. Yellow Syntrichopappus or Fremont’s Xerasid. Fig. 5383. Syntrichopappus fremontii A. Gray in Torr. Pacif. R. Rep. 4: 106. pl. 15. 1857. Loosely pian ann uals 3-10 cm. high, diffusely much branched from the base, except in depauperate specimens, the Boner ions ascending. Leaves 5-20 mm. long, linear- spatulate to spat tiifate. 3- tpgibe ris apex or less often entire, narrowed to 2 Pega estes base; heads many, terminating t bean chlets, ~ peduncles ae involucre . high, narrowly campanulate to nearly cae ric; phyllar s 5, oblong, and thin at buns hand the mid- saltoe strongly concave seid cahanee ers 5, ‘ee: thesis — fe a mm. long, 5-7- — disk-corollas 20 or more, "golden yellow, about 2. es 3 mm. lo toes angled, ob- pyramidal, hirsute we asc cending hairs, the paieae or femcnd: are 35-40 white barbellate bristles united at the base and uous ina r Sandy desert plains ot slopes, a Zones; bea Valley region, Inyo County, south i e Mojave we eer to Los Angeles, pees an nd San Bernardino Counties, California, east to southern Nevada, Utah, ‘ northwestern Arizona. Type ocality : “somewhere between the Roc ky Mountains and the Sierra Nevada.” Collected i ere mont. April’ May 2. Syntrichopappus lemmonii A. Gray. Pink-rayed Syntrichopappus or Lemmon’s Xerasid. Fig. 5384. Actinolepis lemmonii A. Gray, Proc. Amer. Acad. ste 101. 1880. popelgeane te lemmonit A. Cree op. cit. 19: 20. 1883. ‘a lemmonii Cockerell, Muhlenbergia 3: 9. 1907. an floccose to glabrate annuals 2-8 cm. a with reddish gaa et iy mostly branch- ing above the base with asce ending | - ct branches. Leaves ong, linear or linear- spatulate, obtuse ; heads se everal = wai peduncles 8-10 mm. ey jnvolucre 4-5 mm. high, narrowly campanulate; phyllar so G oblong, scarious-margined, the midsection con- cave, the ap acute and thin Tanully spre ading i #4 age; neues of ray-flower s 6-8, white or pinkish above with darker veins conspicuous g, 3-veined ; disk- flowers pale yellow; achenes se pias ut 2 mm. long, attiet thinly atrighae on nearly glabrous, oa Sandy slopes, 1 mostly lower eek of Upper Sonoran Zone; western edge of the ee. sert in ‘Saini Los Angeles, and San B "Califo rnia. Type locality: Mojave Desert. April-May 57. RIGIOPAPPUS A. Gray, Proc. Amer. Acad. 6: 548. 1865. Slender, more or less hirsutulous annuals, simple or branched above, _ lateral branches usually surpassing the ma in axis. Leaves aiding. linea e lowe ually deciduous. eads many-flower oT inati ranch laries narrowly lanceolate, firm, subequal, in 2 series, iia narrowly hyaline- -margine ad. partially clasping the outer achenes. Receptacle flat. ay-flowers pistillate, the ligules hort. Disk-flowers perfect, the tube shorter than the Ania fe throat, the teeth very gant Anthers obtuse at base, subentire. Style-branches short, with subulate hairy tips. Achen linear, abruptly narrowed and truncate at apex, somewhat compressed, finely roa ei rugose and strigose with broad-tipped hairs. Pappus of 3-5 firm, strigulose, awn-like eens [From the Greek meaning 2 a alg and pappus. } A monotypic genus of western United Stat 1. Rigiopappus elena A. mea Rigiopappus. Fig. 5385. Rigropappus leptocladus A. Gray, Proc. Am cad. 6: 548. 18 heed leptoctadses as lonoiaristatus 8. hen Syn. se N. oe 12: 339. 1884. giopapp N. Amer. Fl. 34: 64. Gray-green annual 1-3 dm. high. Leaves ire linear, mostly erect, 1-3 cm. long; h jacranatio ng the branches; involucres 4-7 mm. high; phyllaries hirsutulous ; ; ray-flowers aS SUNFLOWER FAMILY 235 pale yellow, — bie ar with purple, 1.5-2 mm. long, as short as or shorter than the pappus- pale ae; disk-flowers 5-35, shorter than the pappus- pi the teeth of the disk-corollas minute, 2-3- tooth oh acPabes brown, about 4 mm. high, the pappus-paleae as long as to half as long as the Pen: ssland or with sagebru Upper Sonoran and Gogg sara Zones; central b Seach and sauna siiaieds honed ' Utah and Nevata ya in whe Pacific State mountains of Los Angeles County, California. Type locality: The Dalles of the Columbia River. Sst Me ig Pome April-June. 58. TRICHOPTILIUM A. Gray in Torr. Bot. Mex. Bound. 97. 1859. ow floccose plants much branes from the base, annuals or persisting more than one season. Leaves alternate or the lowest subopposite, broadly oblanceolate or spatulate, the margins incised-dentate, narrowed below to a long petiolar base. Heads on long, slender, naked peduncles much surpassing the leaves. Phyllaries in 2 nearly equal series. Recep- tacle eae: k ers perfect, fertile, the ray-flowers wanting. Corolla-tube short, the throat longer, expan fing to the spreading lobes. Anthers oblong-lanceolate, minutely sagittate at base. Style-branches linear, subtruncate at apex. Achenes obpyrami- dal, the surface white ith minute papillae, clothed with ascending hirsute hairs. e Pappus-paleae broad, minutely scaberulous and dissected above into many bristles of un- equal length which are as long as or longer than the length of the scale. [From the Greek Nga meaning hair and feather, in reference to the pappus-paleae. onotypic genus of southwestern United States. 5384 5382. Pseudobahia heermannii 5384. Syntrichopappus lemmonii 5383. Syntrichopappus fremontii 5385. Rigiopappus pene 236 COMPOSITAE 1. ee eee A. Gray. Yellow-head. Fig. 5386. Lgehyrctes incisa rte Gray, Mem, r. Acad..II. 5: 322. 1854. Gray in ig Bet Se yee 97. 1859. t herbs branched from the base, more or less densely floccose-villous iroughout 0d ase except ae gots ncles, the stems 0.5-2 clustered toward the sl : i cres 6-7 mm. high, hemispheric; phyllaries lanceolate to oblanceolate, glandular as well as floccose-villous ; peduncles 8- . long, slender, h, so h a = lar; flowers 35-80, the corollas 4 mm. long, puberulen t and somewhat glandular ; achenes 3 m long, sharply truncate at apex, 5- ib the pappus- Sale as long as or longer than the abhetie hyaline, stramineous or slightly brownish. Gravelly soil, mesas and canyons, sis Sonoran Zone; southern a, eastern San Bernardino County, mcr fei southward through the —— ere to wental Lower Cilitoonis and a to western Arizona. locali ity: ‘On the Californian deser e Rio Colorado.”? Feb.-May; Oct.— 50. CHAENACTIS DC. Prod. 5: 659. 1836. Annual, biennial, or perennial herbs sometimes suffrutescent as base. Leaves basal, persisting in some perennial forms, and alternate above, subentire to more or less pinnately : : si species. Flowers white, yellow, or pinkish, perfect, regular; tube very short; throat nar- rowly funnelform to almost tubular, the short spreading limb 5-cleft and usually densely margined, mostly conspicuous, sometimes vestigial, the pappus-paleae of the margina flowers markedly shorter than those of the disk. [From the Greek words meaning to gape and ray, in reference to the marginal flowers A genus of about 25 species, all somewhat contorted, not flat, rachis about 1 mm. broad; heads 2 or 3, whitish, 12-14 mm. high, terminating the leafy stems, the peduncles short; phyllaries cap m. long, obtuse or r spatula ate; — lar- fap oe aad sparsely nate: coro olla puberulent with occasional glands ; achenes 6-7 mm. long, strigose; peraeae Spalesé 10-46, itacal: 2-5 mm. Tong sbiGae or linear and rounded at the a i en serpentine slopes, Can and Hudso Tilews ; Wenatchee Mountains in Chelan and Kittitas Céiattee: fA oy si osha = locality : omen af Beverly "Creek, Kittitas County. June—July. 8. Chasers douglasii (Hook.) Hook. & Arn. Hoary Chaenactis. Fig. 5394. Hy papp a! ii Hook. Fl. Bor. Amer. 1: 316. 1834. pe tis d te) za Bot. Beechey 354. 1840. carphus ps [Re tt. s. Amer. Phil. Soc. IT. 7: peat os suksdorfit Ae 2 Cane Dudley Herb. 3: 108. — an es Sea 7. 1940. igh, stems densely or sparsely tomentose, also iennial or perennial herbs am ee bas a ee below the Stout, b 1.5-6 dm ane glandular below the inflorescence, single or few biform Ge aifhresciice. Basal eaves 5-1 . long, 2-3-p stem-leaves reduced ad me dissected, all thick in texture, m or less euentnee rulandnine: Sal es “alin ate lobes obtus con d and so hat Pevolite: the leaf thus not flat; heads large, 50-70-flo , white r pinkish; phyllaries 11-17 m g, the outermost shorter, narrowly oblanceolate or oblong, granular-gl lar, more or less woolly with soft, slender, jointed hair also me coarse straight hairs on the back and margin and also o e peduncle; ee ee hairy on the outer surface, the tube glandular ; achenes 6-8 mm. long, densely hirsute; pappus-paleae unequal, 10- » a near or oblong about — — the length of the corol oc andy, sagebrush a , Arid Transition Zone; Yakima County, Washington, south to Jef- ferson “County S Ordena Type loeality: se Celilo Falls on the Columbia River. June—July. Douglas” Pin- cushio at axon which is part of an extremely variable and widespread complex which p tege from the avail- ric seach data, is quite pees ” Beers conditions; consequently ecological as well as regional forms o be found. In eastern Was eastern Origen plants occur that are intermediate in their character- pps betwee C. douglasit var. git Gad C. douglasti var. “sialaetlonh a These forms that cygnetatars might be considered od be C. douglasii var. achilleaefolia have more flow ead and some coarse ight hairs on the phyling ie: All of the taxa of the douglasti complex have to a greater or less degree a adi ah ory 2 leaf-character. The leaves do not — a plane Ps ae but rather a contorted one, due partly to the orientation of the nn nd a tendency to become revolute. This is specially noticeable in the voloced forms pe higher eleva- tions. Aig enactis douglasii var. achilleaefélia (Hook. & Arn.) Roc ts. 577. 1909, (Chacnactis appceci ole Hook. & Arn. Bot. Beechey ah leafolio Nutt. Trans. Am Phil. Soc 376. 1841; Chaenactis imbric ‘a Greene, 1912; C. bracteata Gs ene, op. cit. Tine? é. cheilanthoides Cresits op. 7 . t m . i stud ab co A. Nels in Coult. & Nels. New Man. Bot 1840; Macrocarphus achil. Leaflets Bot. Obs. h; 5 te landular-puberulent as well loosely canescent; corollas, F tanes s, and pappus- pale ae as in C. douglasti douglasit, but the pappus- py especially of plants of the southern ex- pies Ot the range, usually sechatk - Bs 20g the length of the nggriges Plains and dry slo opes, British Columbia o Montana and south to northern Ari and southward east of the Caeade Decarhaing in Washington and Sane to northeastern acs rnia; on a Sierra Nevada principally on the eastern face and the desert ranges as far south as Inyo Count As here identified ae penilleasiete is the mon segregate of the douglasii complex and m uded under the Also peiecane hy with named ee from nigher elevations as oe riants are incl a rebase of lower eto scarcely evident; corolla white, 5-5.5 mm. long, puberulent without, the marginal flowers narrowly funnelform, the lobes este than those of the disk; achenes 5-5.5 m or lon ng with iechdine hairs; pappus-paleae 8-10, eislae tierce, the rex about 2 mm., the Puhortest about 0.5 mm. long. Clayey or greed slopes, Upper Sonoran ie! southwest Tdaho and eastern Oregon in Baker and Malheur Counties. Type locality: sandy hills of the Malheur, Baker County. April-June. 10. Chaenactis névii A. Gray. John Day Chaenactis. Fig. 5396. Chaenactis nevii A. Gray, Proc. Amer. Acad. 19: 30. 1883. Glandular- eae annual often red-stemmed, 10-25 cm. high, simple or branching from the base and a midstem with ascending branches,.glan aeai- puberulent, — ly so below the heads and otichtly athena! villous. Lower leaves at flowering time 4.5 cm. long or less RON 5395. Chaenactis cusickii 242 COMPOSITAE the petiolar base, pinnately divided nearly to the midrib with 3-5 widely spreading, linear, obtuse oe these 5-10 mm. long; upper leaves reduced above in length an nd often linear on unbranched soscir te the outer flowers regular but larger than the inner, the peduncles 1-6 cm. long; invo- lucres 6-7 mm. high; phy aries linear lanc poset F agngeel puberalent with occasional cobwebby hairs, thin, the midrib becoming somewhat prominent in age; achenes black, about 5 mm. long, thickly beset with shine cherie pint. sirentiog hairs nace s of 10 vestigial paleae. Barren dry slopes in heavy clay A Arid Transition Zone; Wheeler and Grant Counties, Oregon. Type hoe or shen ee an or 2 the col co ae oo g data, as the species sien te be limited to the John Day aen 11. Chaenactis artemisiaefolia (Harv. & Gray ex A. Gray) A. Gray. Artemisia-leaved Chaenactis. Fig. 5397. h Harv. & Gray ex A. Gray, Mem. Amer. Acad. II. 4: 98. 1849. Cwasunetin ek Aw, tes grlss Acad. 10: 74. 1874. Coarse, single-stemmed annual 2.5- 10(20) dm. high, often reddish, leafy below, the leaves reduced yo absent in the open cymose-paniculate inflorescence; stems usually mealy-pubescent below, glandular-hirsute above. Leaves satiolae: $-15(20) ¢ m. long, ovals mealy-pubescent, the flat blade ovate in outline, about two-thirds as wide, bi- Printers finely divided into small, oblong or linear iba the petioles Spin or more the length of the blade; heads whitish- or pinkish-flowered, hemispheric, on curved, ascending, glandular- sags peduncles : cm. long; phyllaries 5-9 mm. oe fiseae doreuniate acute, glandular-hirsute; corollas about 5 mm. long, e well-defined tube ee ee vee Rigsiiae satis throat glabrous or nearly so, marginal ere. like the pple eas ac long, linear-clavate and somewhat flattened, the outer somewhat curved, black and _ieroscopcaly oie glabrous or nearly so; pappus- -paleae lacking, "f present np trendaon ntary and so: er common on mountain slopes, often on n disturbed areas, Upper Sonoran Zone; coastal be: evil Santa Barbara County to ip Diego aig pete paehern Lower California. Type locality: Cali- fornia. Collected by Coulter. April—Jul 12. Chaenactis macrantha D. C. Eaton. Large-flowered Chaenactis. Fig. 5398. Chaenactis AELTS C. Eaton, Bot. King Expl. 171. pi. 18. figs. 1-5. 1871. Annual, 7-25 cm. high, stems rather stout, ae senegal from the mee and branctene, ine PS "thinty floccose-tomentulose, often bec ecoming more or less glabra Leaves 1.54 c long, reduced above, pinnatifid to bipinnatifid, the blade ovate in route the se bo base beaaee than or at least Sagat the blade; heads on peduncles 1. cm. long, ‘white-flower ed, the flowers — at night, — united ‘by day; involucre broadly turbinate, the se tg of unequal 4 in and idvei iti i nce ¥ pes ffe t the disk; achenes about 6 mm. g, strigos e; psa iand ee ae in 12 agg , the four inner linear- oblong, 5. mm. mips the four "Blait outer ones about 1 mm. lon On n dese washes, Sonoran and Arid Shanes aid sf! Eon stern Idaho and southeastern Oregon Pos siooeda., get Utah, and northwestern Ari and the ateave Desert in San Bernardino County, aconcg io a locality: “Pahute Mounta ains, Nye Cor pedo Nevada.”’ Collected by Watson. May-June. ince Soe Chaenactis xantiana A. Gray. Xantus’ Chaenactis. Fig. 5399. Ch ti A. Gray, Proc. Ame cad. 6: ise 1865. Chaenactis xrantiana var. rerio A. pcg gd i: 5-3. - dm. high, stems rather toes reddish, the leaves branching at or a little rather | oadly obtu tose tip; corollas age mm. long, nearly tubular, lobes of marginal carallag but little Caer ‘han those of the disk ; achenes black, 7-8.5 mm. long, strigose ; pace: pappus-paleae 4, lanceolate, pl ing the corollas, outer pappus s-paleae usually 4, about 1.5 mm. long, broad and rounded at the Sandy flats and open desert slopes, Upper Son n Zone; pide: and Malheur Po 2 Oregon, ge through western Nevada =< adjacent California po pete ic Face Arizona and in the San Joaquin Valley, pane ony from eastern uis i and western Merced Counties south to the inner northern ranges ad southern California and a western ae Seaast. Trove locality: Fort Tejon, Kern County. April-Jun Desert Chaenactis; Xantus’ Pincushion. 14. Chaenactis tanacetifdlia A. Gray. Inner Coast Range Chaenactis. Fig. 5400. Chaenactis Sper ge A. Gray, Proc. Amer. Acad. 6: 545. 1865. Ann 6-15 cm. high, stems few to several from and near the base, ascending, often curved, little or ot at all branched shows lightly floccose, more densely so at the nodes and below the he es basal or near the , tending to remain at flowering, few above, 1-5 cm. long, about 5-10 mm. ah fleshy, dag to glabrate, the pinnae crowded and often contorted, redivided with short, often minute lobes; heads yell ow-flowered, terminating the stems and branches, borne on SUNFLOWER FAMILY . Chaenactis nevii 0. . Chaenactis glabriuscula 243 244 COMPOSITAE peduncles 2-7 cm. long, the involucre Hog turbinate; phyllaries SEmaige , the midvein mo prominent in age, 6-10 m oe ng, up to 2 wide, ms ae race 4 near- Tioaceae broadly — more or less ay oe ually beco naa na achenes abou g, rsely hi rsute ; inner pappus one-half to Pe “fourths the length - the connir: sb often ieee iy so, Mai oy outer series 4, abou £0. 5 mm. long, ovate or rounded. By en — slopes and Fe oe rans serpentine, Upper Sages Zone; northern Inner Coa nges in Sonoma, Lake, Napa, and Yolo Counties, the Hamilton Range and inner slope of the Sierra heal Range in Sante Clara County, south through the Inner Coast Ranges of ‘Ses Benite and Monterey Coun- ties, California. ite Le age near Clear decay Lake ener: April-June. Serpentine Chaenactis. Inter- nge. ading with C. tanacetifolia var. gracilenta in its northern r nactis Seaarethatia” var. aero (Greene) Stockwell, Contr. Dudley Herb. 3: 124. 1940. (Chae- ctis practiakis Greene, Fl. Fran. 447. 1897; C. glabriuse _ Res Big ere tert — Meg e* Calif. 1124. 1925, as to northern plants only; C. g ula var. gracilen 8.) min arian’ soon wit hering, and leaves in which the pinnate Sahagea are — remote; pappus as in the “getrican’: but ane outer series often incomplete and much shorter. Open pla Yolo, Lake, and Napa Counties, Califor Typ locality : ory ase of Napa County. Intermediate sil ol tanacetifolia var. tanacetifolia and C. Glalitesteta var. heterocarpha eo wpsteces glabriiscula DC. Common Yellow Chaenactis. Fig. 5401. g DC. Pred.:5 + 659. 1836, Annual, 1.5-4 dm. high, more or less leafy, Pera stemmed and branching above with saree Beiacticn 5 sometimes also branched from the base, the herbage thinly floccose, often be- coming glabrate. Basal leaves soon see ara the stem tents es 3-5 cm. long, longer than the inter- nodes, pinnately divided to midrib into flat, remote, linear lobes 2-10 m ong, the uppermost leaves more reduced and often entire; Race Bh res bro adly turbinate or ‘habehetipels ulate, — ngly floccose except at the base, obscurely if at all oviiada yh ananlar* ply llaries 6.5-10 long, up to 2 mm. wide, mostly linear-lanceolate, broadly acute to obtuse at 3 e€ rh ierbaccons. ‘rather thin in age; marginal flowers conspicuously enlarged ; achenes about ong, appressed-hispid, dark or some with pal iz eas ; pappus-paleae equal - Soa me one- sie to three-fourths the length of the corolla, usually broadly lanceolate and sometimes o Open 4 wooded hill slopes and valleys, Upper Sonoran Zone; Monterey te a ee Counties, Cali- fornia, south through the Coast Ranges to Los Angeles and western San Bernardino Counties. Type locality : California. Collected by pos as. March—June. intergrade freely areas of contact. Those of the mountains of southern California a, where C. glabriuscula ar. Siebeiwecs. sig denudata, and var. curta occur, are particularly confusing. Key to Varieties sau tnt — not concentrated at the base; heads borne on branches of the typically single-stemmed plants. er series of pappus-paleae 1—4; plants of northern borders of the Sacramento Valley Cc; jaboied coke heterocarpha. rd. Outer series g (sometimes vestigial); plants of central California southwa Heads teigees: ‘etiae 8-11 mm. long Seer one-half or over the len oe ‘of the — often obtuse. C. glabriuscula eee 5 Pal early equaling the corollas, always ac C. glabriuscula megacephala. Heads siscathe hy llaries 5-7 mm. lon C. glabriuscula esl Leaves many, basal or nearly basal; beads oe on few to several nearly naked stems. Heads (including ihe e corollas) about 15 mm. high; plants about 2 dm. high. C. glabriuscula denudata. Heads i ~ feroflan) 10-11 mm. high; plants 1-1.5 dm. high. C. glabriuscula lanosa. Cha _—_ cula oca4rpha (Torr. & Gray ex A. Gray) H. M. Hall, Univ. Calif. Pub. Bot. : 190. 1907, ( “pur heiprocae aye ' “Tor rr. & Gray ex A. Geay: Mem. Kiaes. Acad. Il. 4: 98. 1849.) Habit Tike that ae & Bose ks var. glabriuscula but usually more woolly: Speed some achenes of the heads often with pale areas; inner sane s-paleae broadly lanceolate, nearly as the disk-corollas, also having an bah Rcties eed 1—4 minute paleae. Northern borders of the Sacramento sony California, in Glenn and Tehama Counties and from Butte County south to Mariposa County. Type locality: Sacramento Valley. Collected by ge Se Hist Soe. 7: var. megacéphala A. Gray in Torr. Pacif. 1857, not A. Gray, Proc. Bost. Nat. t. Soc. 7: 146. 1861. beige mac es var. ctr < Bi Sexckeril, "Contr. Dudley Herb. 3: 123. 0.) Distinguished from glabriuscula var. glabriuscula only by a rge heads with usually larger marginal oe, and broad any eae “28 mor *§ robust habit, by floccose hairs the involucre which are a flecked babs — — Bd a oe nearly sauaies ~~ disk- Gade (vestiges of outer paleae rarely f achenes nC. uscula var. glabriuscula, Com- . Com monly bet sto in the lows er foothills f the Gare Pi oa gy California, Pee Am: eet County south to the Bb eg Mountains, Kern County; also ao te Mba ig 3b slopes of the — Eg be Valley, where it merges completely with c: glabriuscula var. glabriuscula. Type locality: Knights F , Stanislaus County. Weakly separable from the name-bearing taxon by the long paleae eis larger size of held i Chacnactis ee var. landsa (DC.) H. M. Hall, Univ. Calif. Bot. 3:192. 1907. (Chaenactis a be : 659. 1836.) we oad eo 8—-1.5(2) dm. high, densely be! Loma throughout, with several to a mart from the base; s basal or nearly basal, linear h few linear divisions or entire; heads smaller ‘than C. glabriuscula var. A rakeeing on long gubatnive eg "face Cane Ranges, California, from Mon and San Henin t Counties aoe to San Luis Obispo and northeastern Santa Barbara Counties. Type iocaltes California. Collected by Dougla Chaenactis cake var. ascdiix (Nutt.) Munz, Man. S. Calif. 567. 1935. (Chaenactis denudata Nutt. Proc. Acad. Phila. IT. 4: 21. 1848.) b rag like C. glabriuscula var. lanosa in habit but the plants taller, leaves more divided, pam larger, and stems more evidently branched. Southern coastal San Luis Obispo She tl California, south to Los Minitles County rl eastward to San Gorgonio Pass, Riverside County. Type locality Los — Collected by Nuttall. carpha var. curta A. Gray, Syn. ai N. Amer. ed. 2. 12: 452. 18 reo? Greene ex Rydb Amer, FI. 34: 68. 1914; C. glabriuscula var. aurea seen ee Contr. Dudley Nerd. 3: 131. 1940.) Plants no's da high, simple or E gpealt bra sa or ig shorter maller sg esis ame-bear shag taxon; pappus-paleae oval to broadly obl all Soock sh than the — oe mi of the four sn long: Greenhorn Mountains, Kern County. Cakiorna, to iar sheg "Co ounty and south to northern San ven Coane, Type locality: upper part of the Santa Clara Valley, Ventura County. In its southern range intergrading with C. tenuifolia. SUNFLOWER FAMILY 245 a. remem tenuifodlia Nutt. San ati Chaenactis. Fig. 5402. ifolia A. ats e f Chaenactis glabriuscula var. tenuifolia H. M. Hall, Univ. "Calif. Ei Bot. 3: 191. 1907. Chaenactis glabriuscula var. filifolia Tepeok. Man. FI. Pl. Calif. 1124. 1925, as ‘‘ name only. Annuals, (1.5)2-6 dm. high, stems am and wiry, reddish at least at a base, cnares branch- ing above the base with ‘ascending branches, the herbage sparsely floccose to glabra Leaves —6(8), about 0.5-2 de, pinnate or often bipinnate, di ided to t cided in lees ‘divis ons of unequal length, the uppermost leaves reduced, pinnately parted and mostly much the inter a. Rosine mostly many, rather small, on slender peduncles ; involucres mm. high, hemispher any- flowere , the gore hagysage not greatly enlarg mh aa faealei snag wly linear- tte Pi and a about 1 mm. wide, not conspicuously herbace and bec rather firm in age, noticeably eigen Bs csi a metimes — sO, slightly. if at all woo colly: ‘achenes mm. or more long, s Sait pappus caitiaae: narrowly oblong or sometimes acute, one-half t two-thirds as long as the corollas Don places in the aac, i sandy canyons, Sonoran Zones; Orange and San Diego —— east to western Imperial County; ete ped a "County where it teterecaaes with C. glabriuscula var. curta. Type locality: gmp Diego, Californ May-Jul cae tenuifolia var. insatins na Greene, W. Ame oe Dap ce, 33 1887. (Chaenactis cof epee Parish, wyeenS: 92. 1898; C. glabriuscula var. ptt ass H. M. Univ. eat, Pub. Bot. 3: 192. 1907.) Annuals, -3 igh, Bs ms and branches more divaricate and po a C. tenuifolia var. tener es leaves more or oe suabalent, the ultimate lobes rather broad and obtuse; involucres hemispheric, 8— dy high; phyllaries and pedunc ~* as ig conspicuously glandular than the preceding taxon. An ecotype of d c beaches: and Pit of the San Diego oe med coastline and that of adjacent Lower California; merging with ‘C. i ifolia var, tenutfolia away ala the oce: Some plants fro coastal Los Angeles County (Ballona; Redondo) have been oe which have the coarse glandular pubescence of this taxon but the habit and leaves of C. tenuifolia var. tenuifolia 17. Chaenactis freméntii A. Gray. Fremont Pincushion or Chaenactis. Fig. 5403. Chaenactis fremontii A. Gray, Proc. Amer. Acad. 19: 30. 1883. Annuals, 10-30 cm. high, stems often reddish, usually rather stout, 1 to several from the base branched above, the thea nches accendine. glabrous though sometimes spa arsely and loosely ‘inate ar cm. imb; oO abou paleae 4, occasio nally 6, as long as the corollas, fear laceeonates nae of the i eration usually with 1 long and 3 short obtuse scales desert, mostly aeses Sonoran Zone; pire western Nevada and the adjacent Death Beco region, California, southward through the eis and Colorado Dcasts to Tenerial and eastern San Diego aeseties. and eastward to Arizona; also in the San Joaquin Valley in western Kern County. Type locality: California. Collected by Fremont on his iat pais hog March- Was 18. Chaenactis stevioides Hook. & Arn. Broad-flowered Chaenactis. Fig. 5404. Chaenactis stevioides Hook. & Arn. Bot. RRs 353. 1839. ¢ ‘pit 8 floribunda Greene, Pittonia 3: 168. 1897. Annual, 10-15(25) cm. high, sin oe stemmed or several-stemmed from the base, branching above the middle, the fete many, ade h the branches, the herbage finely grayish floccose- 5403 5402. Chaenactis tenuifolia 5403. Chaenactis fremontii 246 COMPOSITAE rage ane phegvenng more or less Satente: the peduncles and involucres gorse -hispidulous well a mentose. Leaves 1-3.5 c ong, the upper leaves rather few, ing or shorter = wba the Boas a asharely div ‘ded to o midrib with divergent — of pie length, 5-15 mm. long, these o often Sey ene with short lobes; involucres subhem “og eric, borne on Satitidies 2-5 cm. lon ng 5 phyllaries 5-7 m ong, ‘bee obtuse t B treadly acute at apex, firm, the midvein ev vident, bec —. ‘Gore so in age: arginal flowers moderately enlarged nearly regular, creamy white Fagen “low s 4.5-6 mm. lo bar nearly tubular, crea ite, sometimes tinged with rose; achen £6 in m. long, strigose pappus-paleae "4, Sulit elanceslate somewhat unequal, the laineat Pov Pe the disk-flow Open, rocky or sandy deserts, poate venues and Sonoran Zones; southeastern Oregon, southern Idaho, ee pore and > ag Colorado sou uth to New Mexico, Arizona, and Seaman: genainen areas of veg hs) ae fornia r California and also pobre in + a southern ae Joaquin Valley, California. Type lit Idaho. Cctleeted by Tolmie. March— Has nah Pincushio ctis stevioides var. brachyp a (A. Gr ay) H. M. Hall, Univ. Calif. Pub. Bot. 3: 194. (Chacnuctis brachypappa A. Gray, tae ta Acad. 8: 390. 1872. aan often larger than the preceding; pappus of the disk-flowers as short as those of the marginal ones. Occurring with C. stev toides var. stevioides ae the type. ming in the Pahranagat noe Vales. central Nevada, gs the desert regions of California 19. Chaenactis carphoclinia A. Gray. Pebble Pincushion. Fig. 5405. Chnanectis RR a se Mex. Bound. 94. 1859. t. Gaz. 47: 434. 1909. Ch ti. ? i Tease qiatkes 25951929; Slender, wiry-stemmed annuals, 1 .5-3 ae high, leafy, much-branched above the base, irregu- larly dichotomous, the heads many, terminating pe spreading branches, gar ool puberulen t throughout and often with glandular Pe a “hai irs on peduncles and involucres. Leaves 1-6 c long, petiolate, the upper leaves reduced, fe ica goo teate, pinnate to bipinnati fd. po lobes remote and narrowly linear to filiform : heads white- or pinkish- flowered on slender pedun cles 2-6 cm. long; involucres subcampanulate, the phyllaries 7-10 mm. long, firm, linear, caudate-attenuate with a pale to reddish, setaceous tip, this more noticeable on the inner phyllaries; receptacle with few to several persisted rigid, hilects. hocks as long as the phyllaries ; corollas Pa mm. long, narrowly funn stot the é obes of the marginal flowers but little enlarged; achen ut 4 mm. long, with a sider "hair ; pappus-paleae of the disk-flowers broadly lanceolate, aisat one-half ie pat of ‘the seared "Copa of the marginal flowers truncate, less than one- -half the length of the c Sandy and gravelly, desert flats, Lower Sonoran Zone; southern Nevada and the Peale alley hen bing Cali- fornia, uk coor the sare and See Deserts to Lower California and western Arizona to Sonora. Type locality: Fort a, Ari March—May. ni s carphoclinia ‘ai Wied uata (A. Gray) M. E. Jones, Proc. Calif. Acad. II. 5: 699. 1895. (Chaenactis attenuata A. Gray, Proc. Amer. Acad. 10: 73. 1874.) Differs from C. carphoclinia var. carphoclinia by having all — pappus-paleae short and truncate and heads often, but not always, fewer-flowered and more narrow; receptacular bra eo vie ely lacking. Growing with the species throughout its range. Type locality: Ehren her rg, fain ‘Coes y, Arizon Chaenactis latifolia "Stockwell, Con tw Herb. 3: 128. pl. figs. 4-6. 1940. Somewhat leafy- stemmed, succulent annual, 15-35 cm. hi wh “ S 7 cm. long, the winged peta wikia the Bae, the blade with a broad rachis and i oa tae “livided with some econ bipinnatifid; heads about high, the } tat cream- chatty phyllaries a ute, hispid, glandular; corollas emadad near the + nibs about 5 —- long, the poppe s-paleae 4, lanceolate, three- fourths the length of the caress Manton San Diego County, California, and n n Lower Joanne Type locality: yey San aa ego Cou Chacnactis nevi stisiat Contr. Dudley Herb. 129. ia "79 0. Wiry- stemmed, The, descriptions of the two preceding taxa have been ada apted from the original descriptions, as no material nh been found hs that Rovers sc en cenrorms * to the original diagnoses. Perhaps they are hybrids derived from some “of ie severa ring in 60. OROCHAENACTIS Coville, Contr. U.S. Nat. Herb. 4: 134. pl. 10. 1893. Slender annual with long internodes, c Saag branching above. Leaves alternate or saree below, linear. Heads discoid, hie lustered in the ieaf-aciis, sessile or ° t Es oS io 3 < °° . Lan | fa) ° a yh ie*] = fe n =e 3 Q ot nm) a ro =o ~ i i9'] 77) 3 — n om as o — 3 ° peg & pp o =) 3 jy QO wn a ° > = o nes cla striate, granular-glandular ; he pappus-paleae thin, hyaline, narrowly S tulate, lacerate- dissected on the margins and apex. [From the Greek word meaining mountain, and the genus Chaenactis.] A monotypic gen 1. Orochaenactis ef Seorang (A. — 8 gs Orochaenactis. Fig. 5406. Chaenactis thysanocarpha A. Gra: a Amer. Acad. 19: 30. 188 Bahia palmeri S. Wats. op. cit. poe Orochaenactis thysanocarpha Covi me diese U. S. Nat. Herb. 4: 134, 1893. Slender, reddish-stemmed annual 5-35 cm. high, simple below and branching above or branch- ing from the base, more or less granular-glandular t throughout and bearing sparse, loose, cobwebby SUNFLOWER FAMILY 247 hairs about the nodes, leaves, and heads. Leaves 1.5-3 cm. long; phyllaries 4 or 5, herbaceous, 5 mm. long, Pee and rounded at the e apex, densely glandular ; corollas 3.5-4 mm. long, stadute® without ; achenes about 3.5 mm. long, the surface dull with darker spots ; pappus-paleae Bi epee he eas as leet as the achen y flats and Sore upper Canadian pe Hudsonian Zones; southern Sierra Nevada in Inyo tyre and kite pehre California. Type locality: southern Sierra Nevada. Collected by Rothrock. July— ‘Aug 61. PALAFOXIA Lag. Gen. & Sp. Nov. 26. 1816. Erect, branching, more or less pubescent, annual or perennial herbs, in some species arising from a woody base. Leaves alternate, thick, entire. Heads corymbose or paniculate, discoid or sometimes radiate, the flowers in shades of pink. pest tees turbinate to nearly : : ‘i aceou ceptacle small, flat, and naked. Ray-flowers when present deeply lobed. Disk-flowers with short tube, the 5 lobes longer or shorter than the throat. Anthers obtuse at base, entire or obpyramidal, 4-angled, pubescent. Pappus-paleae 4-12, subequal, with strong midribs, those of the aacueal flowers often shorter. [Name in honor of José Palafox, a Spanish general. | A genus = i ys 10 or 12 species, natives of southern United States and adjacent Mexico. Type species, Ageratum linea ‘S Roar aM by Be . Oxodhine ss lacssdisesigbn 5407. Palafoxia linearis 5404. Chaenactis stevioides 5405. Chaenactis carphoclinia 248 COMPOSITAE 1. Aen linearis (Cav.) Lag. Spanish Needle. Fig. 5407. Ageratum lineare Cav. Ic. 3: 3. pl. 205. oa Stevia linearis Willd. : 1774. 180 Stevia sig ana HE Schlecht. in Willd. aa Pl. Suppl. 57. 1813, as a synonym. Palafo-xia linearis . Gen. & Sp. Nov. 26. 1816 Paleolaris — ass. Bull. Soc. Philom. 1818: 47. 1818. Hispid or scabrous annuals, glandular above, the herbage dark, 2-7 omy high, branched above the iy Leaves sparsely 0 nescent, linear to linear- lanceolate, narrowed to a sho chthan —_— 1-nerved, cm. long; s few, loos sty corymbose on glan dular shi ncles, discoid, flow pinkish ; voluct e 15-18 le He, 10-20- flowered ; phyllaries 7-13, bay baceous, neat acute, hispid and glandular, camewhat eeled at a corollas perfect, 7-10 mm. long, the lobes shorter than the throat, — styles lng see a e achene dark, 4-angled, vitor Siriiely strigose, the body of the achen = ee ong as hiatrar pappus-paleae subequal, usually 4, spreading at haterity, a little ister on half the len eth of the a siete! "lice alate par strong midrib bordered by a corsa a gile “Nealaac a: margin, Slik outer rete sometimes with ne paleae unequal. Sandy washes and flats, i Sonoran Zone; Inyo County, California, south to the Colorado Desert where Lee teers and adjacent Lower California, east to Arizona and northern aaa Type locality: Mexico. Palafoxia linearis var. Gisvotés M. E. Jones, Contr. West. Bot. No. 18:79. 1933. (Palafoxia linearis arentcola A. Nels. Amer. Journ. Bot. 23: 265. 1936.) Plants becoming shrubby, 1—2 m. tall, the herbage green = scabrous; leaves lanceola ie wairowad to a short petiole, strongly 3-nerved; phyllaries 15-20 mm. long, eglandu- ; achenes ‘about the same length or longer, the pa pus-paleae about 8, four about = the length of the Calif 62. AMBLYOPAPPUS Hook. & Arn. Hook. Journ. Bot. 3: 321. 1841. Ere ees: Sadie 3 linear, the lower pinnate wi ew linear lobes. In florescence many n dark, 4-angled, narrowed below, the pappus of obtuse, scarious, often colored paleae. m the Greek words meaning blunt and pappus + A monotypic genus. 1. Amblyopappus pusillus Hook. & Arn. Amblyopappus. Fig. 5408. Ambisababeas tpt Hook. & Arn. Hook. Journ. Bot. 3: 321. 1841. Aromia tenutfoli Trans. Amer. Phil. Soc. II. 7: 396. 1841. Infantea chilensis a. in = FI. Chil. 4: 259, pi. 48. 1849 Plants yellowish, erect, 14 dm. high, stems eae simple or branched below, much branched above, the branches ascen citime. Leaves toe entire, 1-3 cm. long, the lower somewhat longer and often pinnately 3-5-parted ; heads many, yellowish; ‘iwelicte 3 mm. high, the phyllaries 4-6, thin; marginal a isk-flowers minute, about twice the length of the pappus; long, pa achenes roughened, shi ae sparsely cov ered with coarse hairs, these mostly on the angles ; pappus-paleae 8-12, striate, shining, 0.5 mm. lon ng. On ocean bluffs and by sa hes. Sonoran Zones; coast of San Luis Obispo County and on the Channel Islands, California, seatiieard. m pe California; also Pern and Chile. Type locality: Coquimbo, Chile. April- 63. BLENNOSPERMA Less. Syn. Comp. 267. 1832. Annual, nearly glabrous ws: branching from the base. Leaves alternate, pinnately parted, the lobes remote, or owest rarely entire. Heads terminating the branches, many-flowered, on base of the gee somewhat enlarged. cig + ay flat. a disk- faa dinsiied. Achenes obovoid, obscurely 6-10-angled, the surface papillate, pentane mucilaginous when wet; epappose. [From the Greek words meaning mucus and see A genus of 3 species, natives of California and Chile. Type species, Blennosperma chilensis Less. Lower leaves 8-10-lobed, the lobes short; branches of the stigma yellow. 1. B. nanum. Lower leaves entire or 3-lobed, the lobes long; branches of the stigma red. 2. B. bakeri. 1 ieee oe gala nanum (Hook.) Blake. Common Blennosperma. Fig. 5409. Chrysanth nanum Hook. Fl. Bor. Amer. 1: 320. 1833. ei Coniothel clifornc De Prod, 5: 531. 18 To . N. Amer. 2: 272. 1842. i. nanum Blake, Pia Biol. Sex: Wash, ai 144. 1926. SUNFLOWER FAMILY 249 Herb 8-20 cm. high, Les rather succulent stems diffusely branched from ve base and above, rarely simple, glabrous except for scattered hairs on the peduncles and base of the involucre and tips of the phyllaries. Lower leaves 3-5.5 cm. long, pinnately parted into 8-10 short ried the pper much s er, with 3-6 shor r mm, lo re i with ddish brown , curved o i achenes are shed; ray-flowers rounded at apex, purplish brown on outer face, yellow within ; disk- i henes Wet places on hill slopes a in cultivated fields and vernal pools, Upper Sonoran Zone; Mendo Lake Counties, California, to San Luis Obispo County, — Butte and Glenn Counties south ra Tulare Coma; less common in southern wer aaa os where it occurs in aes and San Diego ponies. Type locality: “North-West coast of Am ”” proba nes San Francisco or ine rey. Collected by re es. Feb.—April. Blennosperma r. robistum J. T. How ef, Le oie West. Bot. 5: 10 ibe 8. Leaves as in B. nanum var. nanum Dae, sini seals ucre 7 mm. sie the tet ovate- rubetes achenes 3-4.5 mm. long, some achenes wi ak the oe illate coat. Sandy soil, Point Reyes peninsula, Marin County, California. Type locality: McClure Beach, Marin County 2. Blennosperma bakeri Heiser. Baker’s Blennosperma. Fig. 5410. Blennosperma bakeri Heiser, Madrofo 9: 103, 1947. Rather succulent herbs up to 30 cm. high, haber om the base and mre ng gecn spi for scattered hairs on the peduncles fs bases of the involucres. Lower leaves 10-15 c : linear, entire or 2-3-lobed with linear lobes ee m. lo Naess the upper leaves shorter, 3-5- lobed he lobes about 1 mm. wide; involucre 8-9 mm. long, the free phyllaries 6-8 mm. long, reddish brown at the tip, scarcely spreading after achenes are ney stigmas of ray- tag red: ee 3-4 m long, 4—-6-angled. Tn met pools, Upper Sonoran Zone; known only from the type locality. Type locality: Sonoma, Sonoma County, California. March-April. . Amblyopappus ee 5410. Blennosperma bakeri 5409. Blennosperma nani 5411. Pectis papposa 250 COMPOSITAE 64. PECTIS L. Syst. Nat. ed. 10. 1221. 1759. Low, branching, aromatic, annual or perennial herbs. Leaves opposite, glandular-dot- ted, entire, usually with a few marginal setae. Inflorescence cymose, the heads radiate. Involucre various ved ours turbinate), the phyllaries 3-12 in 1 series and carinate below, often glandular-dotted. Receptacle naked. Ray-flowers perfect, usually equaling the bracts in number, the ligules yellow or tinged with red or purple. Disk-flowers perfect, few, with short tube and funnelform throat, the limb 5-lobed and spreading. Anthers what angled. Pappus of scales, awns, or bristles, sometimes lacking. [From the Greek word meaning to comb, referring to the setose margins of the leaves A genus of about 70 species, natives of Mexico and South America and Smdbeuaues United States. Type species, Packs ciliaris L. 1. Pectis papposa A. Gray. Chinch-weed. Fig. 5411. Pectis papposa Harv. & Gray ex A. Gray, Mem. Amer. Acad. II. 4: 62. 1849. Dichotomously much-branched annual 1-2.5 dm. high with yellowish green herbage. Leaves narrowly linear with 2-5 an ae, setae n near the base, 1-6 cm. long ; heads yellow-flowered in leafy cymes, the peduncles 1-3 cm. lon hyllaries ee , 7-9, linear ; involucre keeled and gib- us at base, obtuse and scarious ray-flowers 7-9, the ligules m i isk- flowers 10-15; achenes black, 1 near-clavate, sparsely strigulose, 4-5 mm. long; pappus of disk- flowers of 12- 20 short plumose biciatlon these sometimes r se pase toa crown, that of the ray- Hoders a short crown of united scales sometimes cotta into an needs and gravelly soil of the des Lower Sonoran Zone; Dea ie ley region, Inyo County, California, and eastern Mojave Desert south hitack a f 5 Hing Desert to 5 pte California, east to Utah and New Mexico and south to adjacent Mexico. Type locality: California. Collected by Coulter. June—Oct 65. NICOLLETIA A. Gray ex Torr. in Frem. Second Rep. 315. 1845. Perennial gebsae herbs with slender rootstocks, and stems corymbosely few- branched above. Leaves alternate, pienately’ D arted. Heads large, terminating the branches. In- volucres tu thin fete, the phyllaries in “¢ series with a single gland at the tip, subtended by a tuft of hairs at the apex. Achenes narrowly pte fc 0iie hirsute. Pappus in 2 series the inner of 5 lanceolate, awn-tipped paleae; the er of numerous et bristles. Name in honor of Nic ollet, an early American Ente A genus of 3 species, natives of southwestern United =e es Mexico. Type species, Nicolletia occt- Rentatte® ‘A. Gray. 1. Nicolletia occidentalis A. Gray. ee Fig. 5412. Nicolletia occidentalis A. Gray ex Torr. in Frem. Second Rep. 316. Stout, glaucous, ill-sme ne: — als with erect 8 stems 2-6 dm. high, these pi arising ssn deep-seated root-c . Leave oe 3-5 cm. long, pectinately pinnatifid, with a mm Ye the — ming fewer, about on ey “third as bee ee the principal phyllaries ; rays pinkish, 4-5 mm. long nes strigose-hispidulous, 8 mm. long, the awn-tipped pappus-paleae equaling the achenes, the beatles a little short Sandy soil, Lower Sonoran saa western edges of the Mojave and Colorado Deserts, California, from San Bernardino County to San gad County. Type locality: banks of the Mojave River, San Bernardino County. Hole-in-the-sand Tagetes — L.. Sp. ee at, Rid Annual, 3-8 dm. high, with conspicuous glands; leaflets lanceolate- serrate ads in a congested c volucres usec rays villous: minute. This species ada ee ted at Riverside, Riverside County, Caltieinis.. in 1921 and apparently has not become established as a we 66. DYSSODIA Cav. Descr. 202. 1802. Annual or perennial, strong-scented herbs often with a woody. caudex or shrubs, the herbage with conspicuous, translucent, oil glands. Leaves opposite or alternate, entire u an is) c 5 i a. ce ay ® 3 : ° Les | ot apex. Receptacle naked or fimbrillate. ssn when present pistillate, fertile. apse hermaphrodite, fertile, the throat trumpet-shaped, scarcely ace from the tube, lobes 5. Anthers obtuse at base. Style- tian teh slender, truncate or sometimes appendag ed. er chenes narrowly obconic; ea paleae many, tipped with 13 bristles or ai into many bristles. [From the Greek word meaning an evil smell.] SUNFLOWER FAMILY 251 A genus of about 40 species, natives of peereeets United States and Mexico. Type species, Tagetes papposa Vek (= Dyssodia glandulosa Cav. Des 202. 1802.) Involucres ary ae mm. high; stems sparsely leafy, the leaves alternate, much shorter to almost equaling the inter Phyllaries teen ly — leaves 3—5-parted into narrow = p nae 8B i fetegarceas Phyllaries long-attenuate; leaves simple, eae den 2. D. cooper Involucres “Ye aie m. high; ont densely leafy, the leaves cel and mostly longer than the pipette 3. D. thurberi. 1. Dyssodia porophylloides A. Gray. San Felipe Dyssodia. Fig. 5413. Dyssodia porophylloides A. Gray, Mem. Amer. Acad. II. 5: 322. 1854. Lebetina porophylloides A. Nels. Bot. Gaz. 47: 435. 1909. Ch ma porophylloides Rydb. N. Amer. Fl. 34: 166. 1915. Ill-smelling low shrub or shrubby aap et glabrous throughout, 3-6 dm. high, with numer- ous striate branching stems arising from the woody base. Leaves mostly alternate, much shorter than the internodes, 1-2 cm. long, thick, "planatess the lower Gatrowed to a Detioliform | Saas yedtisl into 3-5 ape to lanceolate, entire or incised divisions, the upper often entire or merely incised ; peduncles 3-8 c g; involucres turbinate, 12-15 mm. long, subtended by attenuate piece beacticts ‘ieirce tne: : phyllar ries 14-20, not oe abruptly ccute: terminal gland present and usually i tbe lateral ones ; rays are ge-yellow, about 5 mm. long, sometimes la cking ; disk-flowers orange or deep yellow Tabs ut 8mm. lon ng; ac keke ap pre em 3. mm. long; pappus-paleae 10-12, longer than the achene, dissects d into Dane bristles velly or cng slopes ria alluvial fans, Lower Sonoran Zone; along the Colorado River in San proinong od Coun te ag Ca liforn and the Colorado Desert in Rivera ie: iad San Diego Counties, and northern Lower Cal fornia, east to fe: a Arizona and Sonora. Type locality: San Felipe, San Diego County. March—June. 5412. Nicolletia occidentalis 5413. Leite 3 lloides ve Dyssodia coope’ 5. Dyssodia eeurterd 252 COMPOSITAE 2. Dyssodia codperi A. Gray. Cooper’s Dyssodia. Fig. 5414. Dysodia coopert A. Gray, Proc. Amer. Acad. 9: 201. 1874. Lebetina cooperi A. Nels. Bot. Gaz. 47: 435. 1909. Clomenocoma cooperi Rydb. N. Amer. Fl. 34: 166 1915. Clomenocoma laciniata Rydb. loc. cit. Tll-smelling sh rubby perennial 34.5 dm. high with numerous striate, a scabridous or seabridulos stems bein ing leafy ascending branches arising from a woody base. Leaves alternate, about i od 2 cm. lo re) gla 10 mm. long; disk-corollas about as long as the ; achenes striate, 6-7 m m. high, oe te maturity, pappus-paleae 10-15, mos ay lorie? tan’ the achene, dissected into ial bri ocky slopes and gravelly alluvial fans, Lower Sonoran Zone uthe Nevada and the sia Death Va — region, California, and Mohave County, Arizona, to the eiuth oon 5 oa Se of the Mojave Desert, San oo Nanne eas eat he Type locality: on the eastern side of the Providence Mountai ns, California. uly, 3. Dyssodia thirberi (A. Gray) Robinson. Thurber’s Dyssodia. Fig. 5415. Hymenantherum tenuifolium var. A. Gray, Smiths. Contr. 5: 93. 1853 Hymenantherum thurberi A. Gray, Proc. Amer. Acad. 19: 41. 18 Dysodia _— A. Nels. Bot. Gas, 47: 435. 9. Dysodtia thurberi A. Nels. op. cit. 436, oe Dyssodia Durheed Robinson, pA Amer. Acad. 49: 508. "1913. Pubescent perennial 1-2 dm. high with — slender, ee densely leafy, branching stems arising directly from the perennial root or from a woody caudex. Leaves opposite, sessile, rigid, sparsely beset vith glands, 9-16 mm. Sie cincatet pad: nearly to the zaidrib into 3-7 acicular, eichuel ie divisions, the cous thus appearing fascicled; heads many, terminating the branchlets, the slender peduncles ong ; involucres turbinate-campanulate, 4-5 mm. high, the calyculate bractlets few, short, the 2 se series Oi phyllaries united to near the apex, coriaceous, thinner above, the outer linear series ciliate along the free margin, ~ series port pens small mAbs glands; rays yellow, oblong to oval, about 3 mm. long ; disk-flowers about m. long ; achenes 2-3 mm. long sparsely hispidulous wt gtaae bas as about 10, about eat tee hea] in Menctt the paleae ali - or ravelly or rocky slopes, often growing on limestone, Sonoran Zone Ss; southern Nevada and eastern San Bernardino County, California, eastward through Arizona to western Texas; ’also adjacent Mexico. Type locality: near El — Texas. _ —Sept. pa a ent.) Hitche. Trans. Acad. St. Louis 5: 50 a” 1891. (Tagetes papposa Vent. Descr. Fok Cels “rg 1801; Dyssodia ge tata Cav. Descr. 202. 1802: Boebera tn oes ye rd Willd. Sp. a he 2425. ieee 2 papposa Rydb. ex Britt. Man. 1012. 1901.) Hieapnted, muc eh tre net, leafy annual, the leaves mostly opposite, pinn atifid or Bipineatiéd with linear eeession ns; i involucres subtended by near bractle' ets, these and the phyllaries bearing a few conspicuous, linear or elliptic glands; achenes pubescent, the pappus-paleae dissected into bristles. Roa pace: Recrge' sate, and cultivated areas; collected by Roos (5056) at Loma Linda , San Bernardino Coots, fi nia, not = et well established; widespread as a weed from Illinois west to Montana and rizona. Type ie rn Tilinoi 67. POROPHYLLUM [Vaill.] Adans. Fam. Pl. 2: 122. 1763. Annual or perennial herbs or often low shrubs. Leaves simple, alternate or opposite with marginal oil glands, these sometimes present on the surface of the leaves. Heads discoid, solitary on the branches. Involucres cylindric or campanulate, the phyllaries in 1 series, 5-9, oblong, equal, bearing oil te nds. Flowers perfect, fertile, purplish or yellow. Throat of the corolla funtncltg orm, longer or shorter than the tube, the lobes reflexed, often irregularly cleft. Anthers rounded at hale. Style-branches conspicuous, slender, hirsutu- ous, the apices subulate. Achenes slender, striate. Pappus of many scabrous or hirsutulous hues [From the Greek meaning pore and leaf, OF ses to the translucent oil glands. ] A genus of about species, nati f th sities ode Prone Re ves of southwestern United States, Mexico, and South America. Type 1. Porophyllum gracile Benth. Odora. Fig. 5416. phytilum gracile Benth. Bot. Sulph. 29. 1844 Porophyllum junciforme apg — Bot, Che 2: 354. 191%. Porophyllum vaseyi Greene Porophyllum caesium ao 155. Bushy perennial, woody at base, 2 dm. high, with many slender, erect, rush-like branches; herbage dark green, often pu shetty glaucous, with a gig oat me reeable odor ie the scattered oil glands. Leaves few, li linea ar-filiform, entire, 1-5 cm. g; involuc wly campanulate, 10-15 mm. long; phyllaries 5, often tinged with purple, oil a. a Tinear- one obtuse, the hyaline margin often pinkish, r Sierra = the back and somewhat gibbous at the base; corollas 7-8 mm. long, purplish white with purple lines, throat funnelform = ace cxocedine the tube, puberulent ; —_— ee — . long, ‘taal “orton the pappus shea 6 6 mm. long, of coarse, hispidulous, often SUNFLOWER FAMILY 250 Stony desert slopes, Lower Sonoran Zone; sme pee goers County (Clark ag ge preonrd California, pesos theaeh pe sage Pag i and the mounta f San and Orange Counties to Lower California, eastward t ern , Texas, and Son ma sl ie i er Bo Bay, Lower California. March- aoe Tribe 3. ASTEREAE* Heads all Pelaci perfect disk- sista commonly radiate, not dioe be fee Ray only pieeineh wanting; disk- Pate ae Pappus ot pala, scales, or flat wns. Leaves usually rat e pe more than 5 mm. wide; achenes glabrous; Ba cart a firm deciduous Leaves ae mostly much less than 5 mm. wide; achenes hairy; pappus of 10 or more persistent members. pi Leaves filiform to narrowly oblanceolate; pappus of 10-12 sordid, oblong, free scales; phyllaries not prominently § scarious-margine ed. . Gutierrezi Leaxen: bade wg in Oey pappus of 15-40 bee, — narrow paleae; ahs ities broad, an. ‘o unded, with p ent, scarious, erose mar ads lomerste ‘disk: ae ers 4-7; plant spinescen 70. Amphipappus aoe solitary but cciecicaa approximate; ix flowers mostly 20 or more; plant not spinescent. . Acamptopappus. Papp f capill secs a) sometimes with some short outer scales as well. a Pappus "ak disk: flowers double, the inner series of capillary Male the outer much shorter. R ay-flowers with copious pappus and pubescent achenes. rysopsis Ray-flowers epappose or the pappus eae sa to a Cg crown and with essentially ahs achenes 73. Heterotheca. Pappus of a ge i: Ie generally unequal but not divided into two lengths. Disk- ower fer eh sauna never rhizomatous. eads nodding in bud; slender, usually diffuse annuals with slender or filiform stems; shytlaries thin with prominent scarious margin. 74, PELTON EE Heads erect or at least not nodding in bud; ostly perennial herbs o ia i heads usually few and yo soak fs “large, if ponte and ag nicled thon opappus Renan shr ae . Haplo Plants oe numer roots arising fro a zome or caudex, eset a taproot; aot peo ay ad and very PO, we herbace olidago. Disk- ee sterile. Plant a nnual; phyllaries not in vertical ranks, with reeuered ae sits a prominent land. 76. Plant a tufted perennial herb; phyllaries in vertical ranks, ih 6 erede tip not bearing a prominent gland. 78. Petradoria. Ray-flowers none. Plants are: Tie | in more or = eee vertical ranks. yi ND BF nl eeretticay wi llaries not in vertita 75. Ha blo wr Aguas Plant a diffusely ch, 92. Les. Ray-flowers ai pink, purple, or blue, sometimes very inconspicuous, occasionally pooh ae —. yellow, white, pink, or reddish purple. Pappus none; receptacle comnse i pants scapose 80. Bellis. Pappus present; habit various. Pi pil nnien few to numerous, in 1 or more series, usually with conspicuous rays; disk-flowers few to generally numerous Pappus of a ee or scales, sometimes with additional hapigee Di y leafy panicles. pene . Boltonia. Disk-achenes not winged; flowers mostly solitary. Pappus of iti bristles | alternating with short pes or of a scarious cup and bristle nnuals 2. Monopftilon. Pappus of “numerous rigid narrow awns, or awns ae neal or scales only; ours mostly plants. 83. Townsendia. Pappus of mostly capillary bristles. Ray- baled present, sometimes very inconspicuous ‘ i firm bristles; phyllaries with MS scarious margins; small an wit — whi ite or pexplaia? : 74. C hactopappa bellitore. Pappus usually of dents bristles, in Erigeron very rarely as few as 5 a plants perennial. Achenes beaked; phyllaries decid t 1 with — branches. 84. acyina. Achenes not beaked; phyllaries persistent. us in ray-flowers obsolete or none; leaves mostly laciniate-pinnatifid; slender ee annual. . Psilactis. Pappus present in both ray- and disk-flow Phyllaries ent obviously pecan? and imbricated in 3 or more series. St vlebranches of disk-flowers not tipped with a prominent tuft of : w hairs; ray-flowers fertile; pappus not reddish. isa usually rhizomatous or =~ fibrous roots, if an- nuals then leaves pacts So entire * Key by David Daniels Keck. 254 COMPOSITAE eager Barer pricy 4 chartaceous and white at base, tip green and pyre) lax; Pied: few, short and oc . Sericocarpus gies if indurated not conspicuous and white at base; ink or yer ed several to many, if pal or vestigial not on plants from a woody caudex r stout ome eS ae of sth Sadhana pad as to subulate, phyllaries aay od an even scarious Appendages of style-bra oe siphon or oblong, obtu tte desert perennial with bristle- aan sabiisteie & 8 Annuals, biennials, or t-lived pace im with a taproot; leaves pan Bs Fay entire or usually spinulose-den- tate to pinnatifid or dissected. . Machaeranther Style- es of disk- -flowers baru tufted with rigid vase hairs; ; pappus reddis pi orethrogyn Phyllaries only slightly or not at all graduated, in 1 or 2 Magda? Bed branches of disk- foxes with lanceolate and acu or ith triangular and obtuse, appendages. 91. Eri Ray-flowers absent, but the marginal disk-corollas sometimes enla st Ph resemble rays. Mar, wpa corollas enlarged, becoming palmately 5- gel 2 appt usually eceul & to . Lessingia. ia pinat coeds not enlarged, regular; pappus not biGirietah or reddish. pisteiate te ery numerous, several-seriate, tubular-filiform, rayless or the anes ray Sgt the pappus. . Con Heads unisexual, Pvt dioecious; pappus of male flowers of clavellate bristles. : 94. Baccharis. 68. GRINDELIA* Willd. Ges. Naturf. Fr. Berl. Mag. 1: 260. 1807. Annual, biennial, or usually perennial herbs, mostly with a taproot, rarely suffrutescent t e€ re ss resinous particularly on the involucre. Leaves alternate, punctate, sually serrate and sessile, often clasping. Heads medium to large, yellow, usually radiate, usually solitary at branch-tips. Involucre multiseriate, imbricate, the phyllaries thickish, c A gular, few-ang wled, glabrous? pappus of 2-8 stiff, “bftea urved, deciduous, co paleaceous awns. med for Professor David Hivenntices Grindel, esc iaae af sear of Dorpat and Rig: genus of Boag % species, of western North and South America. Type species, Grindelia inuloides Willd. Tips of p spreading, some gradually ply reflexed. Fruticose; salt tonne e eas Francisco Bay. 1. G. humilis. Herbaceous, m woody at caudex. Branches qrotree or —— a bovat t ecotey 1. . G, stricta. eaves no t interior 3. G. integrtfolia. Involucre pubesc cent a Ranges, central California. 4. G. hirsutula. Involucre glabrous; cenkeed Califor Stem fs 8 dm. hi gh, erect, strict branching above; interior apr . G. procera. Stems 3-8 dm. high, ascending, openly branched; San Francisco. ae 6. G. maritima. Tips of phyllaries 1 f iddl d outer ones) sharply reflexed or Jooped. Coa — raccural 1 plants 1 dad +. T i +. - a | +1 p ‘ate. north of San Francisco Bay. i hi 2. G. stricta and subspecies. Leaves acutely or obtusely pointed at tip; mostly south of San Francisco Bay. 7. G. latifolia, 'y below; nearly erect. 2. G. stricta blaket. hideas an nonsucculent plants; stems more or less erect. es ate; rare introduction. 8. G. squarrosa. Leaves sharply toothed or entire, not callous-serrulate. neice mostly 1 cm. in diameter; ligules 5-8 mm. long; San Diego ax: poe gaye it. Involucres 12-25 mm. in diameter; ligules (when present) mostly 8-15 m mn, ips of phyllaries spreading ~ a (usually some a. refed) sce Cin looped ; Lape or bemere s * ‘Cae poe aed Meade relatively small, the involucre 7-10 1 mm. high, 9-15 _ aging northern California Heads noc pet involucre 8-15 mm. high, 12-25 mm. thick; southern Calif Latenn. '* Text contributed by David Daniels Keck. SUNFLOWER FAMILY 255 1. Grindelia humilis are - Arn. Marsh Grindelia. Fig. 5417. Grindelia humilis Hook & Arn. Bot. Beechey 1 Grindelia robusta var. eg lia A. sgl Bot ies 304. 1876. Grindelia cunetfolia of most authors, no’ Frutescent, to 1 gis . high, stout, #6. woody stems giving rise to herbaceous shoots of the season, the branches subcorymbose, glabrous to villous, few-headed. Leaves eeee re acta resi mate s, remotely serrulate, cuneate-oblanceolate to lance-oblong, 2-8 cm. long, 5-1 ro to a sessile base or amplexicaul ; heads 3-5 cm. across ; phyllaries of the inv ere Gaatiatacteest, with short, flat, appressed or slightly reflexed tip, or tip more subulate and strongly reflexed; rays 16-34, ~ 18 mm. lon ng. Sal marshes, Hu map Trawttion Zone; San Francisco, San Pablo, and seve Bays, California. Type locality: * Califor Principally Aug.—Oct. but found in flower throughout the yea Grindelia sataiacs gra om Bay Reg. LN Bg (Coindees ria kao bo panto Jepson, Fl. a Mid. es lif. the Re Stems herbaceous re the wn, stout, freely b glabrous, 5-15 dm. high Leaves subcor not resinous. motely pirate. serrulate, or pool an one hanya or oblanceolate, strongly ‘clasping, the "awe mostly u ti o 8 cm. long and 1 m. wide, those near the heads much reduced; heads 3- pcisccn baghe: resinous, the senna green tip the phyllaries erect or. spreading, the outer ones str s 20-35, 10-12 mm. long. Salt Pate del Tower Nouceun Zone; Suisun Bay, Solano County, Ca Vinci As all’ protiability this is a stabilized hybrid population of the parentage G. humilis and G. camporum, Type locality: Suisun marsh. July—Sept. wea ots x AS slit XS : > Ne At yee Ms \ / a Porophyllum gracile 5418. Grindelia stricta : . Grindelia humilis 5419. Grindelia integrifolia 256 COMPOSITAE 2. Grindelia stricta DC. Pacific Grindelia. Fig. 5418. Grindelia stricta DC. Prod. 7: 278. 1838. Grindelia oregana A. Gray, Syn. Fl. N. Amer. ie nes 1884. Grindelia a Saath Pittonia 2: 18.18 Grindelia lanata Greene, op. cit. 290. ee Z. seeders saad See dg Greene, = cit. 3: 297. 1898. wilkesiana Diner in Piper & Beattie, Fl. Northw. Coast 363. 1915. Grindeli d. i Pip r, Proc. Biol. Soc. Wash. 31: 77. 1918 Grindelia stricta var. anderson beta Ann. Mo. Bot. Gard. 21: 559. 1934. Grin de. lia stricta var. aestuarina Steyermark, op. cit. 560. ark. o cl 1 aia lla J . Grindelia stricta var. hendersonii Steyermark, op. cit. 562. Gorindelis pogtepata. pearson op. cit. rophylia Cxthduin, Vasc. Pl. Pacif. Northw. 5: 207. 1955. Herbaceous sel to 0.5 dm. high; stems simple or corymbosely branching above, decum- bent or ascending, 3-8 dm. long, glabrous to rslobate x or ps Hen de villous and sometimes bres Leaves peas the basal oblanceolate, acute to rounded, entire to serrulate toward a apex, 10-25c long, 14 ¢ ~~ narrowed to a margined petiole, the cauline broadly oblong to oblanceolate, le or acu eats e or slightly serrulate, 3-12 cm. long, amplexicaul; heads oft ten leafy-bracted staid ucre 8-14 mm. high, 12-20 m m. wide, the linear- “snicentate clrllacieg pi (occasionally the ou ere eli mn the slender, nerine. herbaceous tip erect or gradually r ane rays 10-35 20 Coastal dunes and salt marshes, Humid Transition Zone; southern Alaska and Vancouver Island to Lane eos, Oregon. Type locality: Mulgrave (Yakutat Bay), Alaska, but quite possibly from farther south. June- Grindelia stricta subsp. — (Jepson) Keck, Aliso 4: 102. 1958. (Grindelia venulosa Jepson, Man. Pie 2 a. 1021. 1925; G. cola Steyermark, Ann. Mo. Bot. Gard, 21% on 1934; G. stricta bs procum- bok ns Steyermark, op. cit. 9: C. pean st var, pachyphylla Steyermark, op. 596.) Procumbent to decum- bent, he pons usually whitish or yellowish; leaves often shorter, broader, and ‘fleshier, rounded at te: phyl- laries with prominently heyrangy r looped tips. Coastal salt ee and seaside bluffs, Coos spa ce Oregon, to San Francisco Bay; rare in Roecisres cheated California. Type locality: Big Flat, Humboldt Cou Grindelia stricta asa blakei (Steyermark) Keck, Aliso a "102: 1958. é Grimeriie blakei nee Ann, -— tages ca tna — hat 1934.) Bh trcinaerst the stout red-brown stems o season big mei wig be wood high; leaves or wine pope usually os heads to 5. v. Pay across, the tap a ties A Bay, California. Ton locality: Ex i gi ee —Sept. "Detpite its woody axis and essentially erect branches, a Fas nt is doubtless more Ke ga govare. ~ forms - G. stricta from the Puget Sound area than to G. humilis fe) mimics 3. Grindelia integrifélia DC. Puget Sound Grindelia. Fig. 5419. Grindelia integrifolia ee Prod. 5: 315. 1836. Grindene vir pera } Bint . Trans. Amer. Phil. Soc. vs BAR 1840. virgata Torr. & Gray, pie Amer. 2: 248. 1842. Grindelia coltlan: ein y, FL. S. Brit. Columbia ats Grindelia integrifolia var. aestivalis Henry, loc cit. Grindelia adil 4 sage gubaeenes mand loc. ¢ Grindelia ke ee Gard. 21: 564. 1934. ence os ah several erect or chine stems from the ligneous caudex, 2-8 dm. high, gla- rate or somewhat villous. Leaves membraneous, the basal tufted, oblan ceolate, rounded, moder- ately serrate or more commonly entire, up to 35 cm. long and 4 cm. cate: the cauline lanceolate or oblan ie usually acute, sessile or even auriculate-clasping ; heads salbate to u igo neck n a corymb, cm. across; involucre somewhat gl uti nage the slender green oxi of the phyl- laries teaming loose or spreading or gradually recurving ; rays 10-35, mostly 8-15 mm. or ae mm ong ar rshes and wet meadows west of the Cascade Mountains, Humid Transition weg British Columbia : acai ee Puget Trough, to the head of the Willamette Valley, Oregon. Type 1 ocali ity: Williamette Valley. June—Sept 4. Grindelia hirsitula Hook. & Arn. Hirsute Grindelia. Fig. 5420. Grindelia hirsutula Hook. & Arn. Bot. Beechey 147. 1833. Grindelia hirsutula var. brevisquama Steye sg Ann. Mo. Bot. Gard. 21: 572. 1934. p.c lees ¥. Grindelia hirsutula ge subintegra Steyermark Grindelia pacifica oA: Bull. Torrey oat 9: . 1882. . ennial 3-8 dm. high; stems erect, slender, simple or commonly ssadrarraggse U1 s chartaceous, gray-green, the basal oblanceolate or spatulate, obtuse, ens serrate = oF ant ce etimes merely ~ crenate, tapering to a narrowly margined petiole, 1 ong, 3'5 5 cm. wide, the cauline much reduced in size upw. —: from oblanceolate and sessile he below = narrowly isan. patie bear t-like and clasping above; heads 2.5-4 or 5.5 cm. ross ; involucre more or less pubescent, the gtaleeerinte shevincies erect or nearly so, not caudate ; rays 20-35, 1 San long. Arid slopes, Humid Transition Zone; Coast Ranges of central California from Napa County to Monterey County; also — a ounty. Type locality: California. April—July. a Grindelia a subsp. rubricatlis (DC.) Keck, Aliso 4: 102. 1958. (Grindelia hav Prod. 5: 316. Bros te paicne t Greene, Pittonia 2: 50, 1892: G. a var. patens Jepson, Fl. W. Mid. Calif. 554. 1901.) Stems usua lly becoming dues villous above; peduncles prominently bracteate, the uppermos SUNFLOWER FAMILY 257 linear br: rowded, retrorse, usually lanate, equaling or surpassing the disk; outer phyllaries often spreading with eoirnel ‘he: iiccohtlc gically well marked but sympatric with the typical subspecies; also in the Sierran foothills of Fresno County. Tose locality: Berke ley 5. Grindelia si Greene. Tall Grindelia. Fig. 5421. Grindelia procera Greene, Grindelia camporum var. porviftora fg at ‘Ai. Mo. Bot. Gard. 21: 534. 1934. Herbaceous perennial 8-18 dm — stems erect, pallid, unbranched below, rather strictly much-branched above, bearing numerous heads. Leaves chartaceous, dark ~— ae bally:< cauline, oblong-obovate to ie. oblong obtuse, clesping, shallowly toothed, "3-7 cm. lon up to2c ide; heads 2-3.5 cm. across; idan: slightly resinous, mostly under = mm. fe the shock green or ace eg erect or slightly saat at tip; rays 21-45, 8-10 mm. lon Deep soils, Upper er nehig gene: bottom lands of the San Joaquin eee Sacramento County to Kern County, California; also rin County. Type locality: “the lower San Joaquin.”’ July—Oct. 6. Grindelia maritima (Greene) Steyermark. San Francisco Grindelia. Fig. 5422. Greens Sabonennits var. maritima Greene, Pittonia 2: 289. 1892. Steyermark, Ann. Mo, Bot. Gard. 21: 576. 1934. Stems erect or ascending from a ligneous caudex, 3-8 dm. high, slender, glabrous or rarely villous, icuaie branching. Leaves thickish, ne pita the basal tufted, narrowly oblanceo- rindelia 5423. Grindelia eat 258 COMPOSITAE late, up to 18 cm. long, tapering to slender petioles, the cauline oblong B ee ge 3-7 cm. long, 8-15 mm. wide, clasping; heads terminal on the slender bran cross; involucre more or iss gla ig ate the phyllaries with gine erect, aeen gn eR short tips; rays 30-40, 10-13 mm. lon Ocean bluffs and open hillsides Humid Transition Zone; San Francisco, California. Type locality: Point Lobos, San Francisco, Aug.—Sep Z. — latifolia ~ Coastal Gum-plant. Fig. 5423. se latifolia f. Acad. 5: 36. 18 Grindelia robu vial var. gible a Man. FI. Pl. cae. Eset 1925. Grindelia puritans var. latifolia Steyermark, Ann. Mo. Bot. Gard. 21: 227, 1934. Very succulent and leafy, cebe pam perenni * oy dm. high, the stout, decumbent or ascending nches monocephalous or topped by a close cluster of 2 or 3 large heads, the heads often par- bra tially enveloped by su “ae see leaves. castes pr vincinaily cauline, thick, irregularly serrate to regu- larly and Sea dentate, scabrociliate, lance-ovate to broadly a long, ange. 1 to subcordate, rounded at apex, 3-8 cm. long, 1.5-4 cm. wide; heads 3-5 cm. acr milky- ous, the outermost phllaries fliaceous, their cath tips ae oni those of many ee ones Potton rays 30-45 ng. Salt marshes and dunes, Upper Sonoran Zone; Watsonville coast, Santa Cruz Courty, god Surf to Point Maye eek ne Santa Barbara Islands, California. ge 8 mins Santa Rosa Island. May-—Se Gri: a latifolia subsp. platyphylla (Greene) K o 4: 102, 1958. “pclae ag ieee var. platy- ye seg Pittonia 22 S89. TR92 °C. rubricaulis ag “plaixpigil Steyermark, Ann. Mo. Bot. Gard. 21: 227. rag decry var. permixta Steyer mark, op. Neither so succulent nor a grade we mony simple or re nched, mbent or ascending, hori gabe ah poe radiating from the crow & pla ants up to 6 dm. high sds “y m. in diameter, thes te -resinous taal sometimes subtended yy severai smal ‘les y bracts a never enveloped ss these. Beaches and low ground near the coast oom common species from Mar County to Santa Barbara County, Calitocnia, Type locality: Monte rey. Jun oo 8. Grindelia squarrésa oe Dunal. Resin-weed. Fig. 5424. onia squarrosa Pursh, Fl. Amer. Sept. 559. 18 Grindelia squarrosa Dunal, Mem. Mus. Paris 5: v4 1819. ect biennial or short-lived perennial 2-10 dm. high, openly branched above and bearing many heads. “eave ee rly callous-serrulate, sometimes sharply toothed or even e entire, mostly oblong, 2-5 ¢ e upper clasping; heads 2-3 cm. —— sated resinous, the green tips of the piyllaries strongly rolled back; rays 25-40, 7-15 m ds and waste places, per Sonoran and po A ransition Zon sparingly introduced in eastern Waiktaxien? gg go and te. Ga ne a chiefly east of the pea Ease *bivide to the Great Plains. Type locality: banks of the Missouri. 9. Grindelia hallii Steyermark. San Diego Grindelia. Fig. 5425. Grindelia hallii Steyermark, Ann. Mo. Bot. Gard. 21: 229. 1934. _ Several pong oe stems from the woody crown, 3-6 dm. high, corymbosely branch- ing above. Lea sharply and regularly serrate, subcoriaceous, prominently resinous- punctate, he rai i in a per — rosettes, Clee pie 5-7 cm. long, the cauline oblong, much smaller ; heads numerous, terminating the cha s, 2-3 — “hs oot resinous, the green tips of the outer phyllaries sidngl¥ tec urved o ids ked ; ra m. lon ats and grassy mesas id id 1 Transition Bare ope a Saas San Dae Coauké: California. Type locali ity: Cuyamaca Lake. Jul 10. Grindelia campérum Greene. Great Valley Grindelia. Fig. 5426. Seg sani oat var. rigida A. Gray, Bot. Calif. 1: 304. 1876. Grindel mporum Greene, Man. vor Reg. 171. 1894, Grindelia robuste _ de Jeu, F W. Mid. vouge 554. 19 interioris “tne Man. FI. Pl. Calif. pet 1925. Grindeli davyi Stey ermark, Ann ie. Bot. Gard. 21: 536. 1934. eee —a var. E eccos toris Steyermark, op. cit. 538. ect herbaceous stems from the su oe —" 5-12 roa a simple or openly branched ous or nearly so. Leaves glabrous or scabrous, rarely e hairy, subcoriaceous, very sth gird os rows ici slices to ica oblanceolate, the cauline 2-8 cm. long, 7-15 mm. wide ; hea ie nating the branches, 2.5-4 c cross, strongly and translucently resinous, the green xe of tx itie Cadesestite elongated iets ahsely recurved or hooked ; 18-35, 8 8-15 mm. lon Dry banks, rocky oid and plains, low alkaline ground, Upper Sonoran Zone; the common Grindelia_ of the pad tL th ieee ogg and adjacent foothills — to northern Los Angeles Counts and west to San Francisco Bay. east of Mo unt Diablo Ran May-Oct. 11. Grindelia nana Nutt. Idaho Resin-weed. Fig. 5427. Grindelia nana Nutt. Trans. Amer. Phil, eee Il. 7: 314. 1840. ‘indelia nana var er. ban oo. loc. c: pa redenete nana var. altissim ermark, Ann. Mo. Bot. Gard. 21: 544. 1934. Grind. a var. celine an pda op. cit. 545. Ghacue perennial with erect, usually branching stems, 2-8 dm. high. Leaves spinulose-toothed SUNFLOWER FAMILY 259 to ep she lower oblanceolate, up to 15 cm. long and 3 cm. ide. fapering to slender petioles, abe upper smaller and clasping ; heads 2-3 cm. across; involucr: aes vesy resinous, the gre tips of gr phyllaries strongly rolled back; rays — 12-25, 15 m Open ground, dry hillsides, roadsides, and subalkaline flats, U Sonoran oF Ax ging roast Zones; wit "Caliseraia, eastern Oregon and Washington lochaais phat gf mse to _T petbiety Type locality: pr rob- ably northeastern Or Beit . June—Oct Grindelia nan sales colu a Piper, Contr. U.S. Nat. Herb. 11: 556. 1906. (Grindelia discoidea Nutt. Trans. mare Phil, ges ate, 5. 1840, not Hoo k. & Arn. 1836; G. nana var. discoidea A. Gray, Syn. Fl. N. Amer, 17: 119. 1884.) Heads discoid. Common in dry or moist pi in central Washington, where it viens roe the typical subspecies, west to Portland; northern Idaho. Type locality: “‘on the banks of the Oregon.” ug June- 12. epee robista Nutt. Big Grindelia. Fig. 5428. Grindelia robusta Nutt. ans. ~— Phil. Soc. IT. 7: 314. 1840. Grindelia cuneifolia yi op. cit. 315. dasa pea fe = ctits che var. ive is rmark, igew Mo. Bot. Gard. 21: 227. 1934. usta Steyermark, loc. cit. isp siekchom ¥ var. ss stale! Steyerm ae op. cit. 228, a. Stems few, erect from a subligneous crown, stout, usually corymbosely branching above glabro i ‘5 12 dm. high. aves sharply toothed to fin ely and remotely serrate or often entire, the basal abinteolais up to 18 cm. long (including the maiteised petiole) and 3 cm. wide, the cauline 4) NVE ta, Sa SS TAINS 5424, Grindelia squarrosa 5426. Grindeli porum 5425. Grindelia hallii 5427. Grindelia nana 260 COMPOSITAE reduced, ovate-lanceolate to linear-oblong, broadly clasping ; heads 3-5 cm. across, —- aed and translucently resinous, the long green tips of the phyllaries rolled back in a loop; r = 8-15 mm. long. Dry eee and banks, also borders of salt marshes and shores, Upper Sonoran Zone; not far inland, Santa Barbara County, California, to northern Lower California. Tyee ‘locality: San Pedro. Mar ch—Sept. Sia cuit var. bractedsa (J. T. Howell) Keck, Aliso 4: 102. 1958. le nae Bids te a Daal eo ogg Beste 2: 22. 1931; G. “Shieleaalis var. bracteosa Steyermark, Ann. Mo. Peas rd. : 227. 1934.) Hea scoid. Dry hills and eee Ana Up . se Zone; away from the snc Shere ag tt; tone tae perni4 Hil, Los Angeles cued: to e Hills, n Diego County, California. Type ontitr! Santa Ana Cany May-Jul 69. GUTIERREZIA* Lag. Gen. & Sp. Pl. 30. 1816. Perennial herb or subshrub, glutinous, glabrous to hirtellous. Leaves alternate, entire, filiform to narrowly oblanceolate, usually punctate-glandular. Heads very small, radiate, yellow, numerous, scattered or crowded in cymes or panicles. Involucre cylindric to turbi- nate-subglobose, the imbricated phyllaries coriaceous, appressed, whitish. Receptacle fo- veate, sometimes hairy ay-florets pistillat fertile. Disk-florets perfect, sometimes sterile. Achenes obovoid or aS ong, paeeeent: pappus of 10-12 oblong, unequal, free scales, shorter on the ray-achen | Named for the Spanish nobleman, Pedro Gutierrez. ] foli es genus of about 25 species, ese = western North and South America. Type species, Gutierrezia lineari- olia Lag. Heads flaring, more than 4‘ flowered. , 3-5 mm. thick; ray-florets 5-10; disk-florets 6-16; cismontane. 1. G. californica. Involucre narrowly turbinate, 2—3 mm. thick; ray- and disk-florets each 3-6; transmontane. an - sa Heads cylindric, about 1 mm. thick, 2—4-flowered. . G. seerniia 1. Gutierrezia californica cases Torr. & Gray. San Joaquin Matchweed. Fig. 5429. Brachyris californica DC. Prod. 5: 313. 1836. Gutierrezia californica Torr. & Gray, Fl. N. Amer. 2: 193. 1842. = manoeeeyean oe ee um Greene, Man. Ba Pye ihe tenes Abrams, Bull. pad cist 34: Gutierrezia californica var. bracts ata H. M. Hall, sa: ee a Bot. 3: 36. 1907. Subshrub 3-6 dm. high, nearly as to densely poms the branches stiff es ay or metimes sping and spreading, very twiggy above. Lea preading to deflexed, up to 5 cm. long, ostly 1 mm. wide; heads usually solitary at ends of hratich bas. arel¥ crow es oe oie ag curb ae -O oveid or campanulate, 5-7 mm. high, 3-5 mm thick, the usually broad blunt sdvyltisics with definite green tips; ray- -florets 5-10; disk-florets 6— Dry hills and plains, Sonoran Zones; Marin County, California, the Sana South foam Range from San Fra Bove a Bay to sone San ar, pe: and Seemontane pat oi California east to Arizona and Chinaahds. Type locality: California. May tergrading with G. sarothrae in southern California. 2. Gutierrezia saréthrae — Britt. & Rusby. Common Matchweed or keweed. Fig. 5430. semeas estar Pursh, Fl. —_— Ce 2: 540. 1814. 63. 1818. Gutierrezia e eT ; Fi, . 1842. Gutierrezia sarothrae Britt. & Rusby, Trans. N.Y. Acad. 7: 10. 1887. sy eereete PETERS. Gree Pittonia 4:56. 1899, in large part. ricina » Rep. Nov. Sp. 7: 195. 1909. Gutierrezia ionensis Lunell, Amer Midi. Mat. 2: ng 1912. Xanthocephalum — Shinners, Field & Lab. 18: 29. 1950. above. Leaves ‘punctate, a5. ce on 1m. ide; in ii escence flat-topped, the numerous resinous heads scattered or usually mall glomerules ; involucre narrowly turbinate, 45 mm high, 2-3 mm. thick, the a bi faith lea peat thic eed herbaceous tips; ray- and disk- florets 3-8 each, the ligule about 3 mm. long; achenes subsericeous- bevery pappus of ray about 0.7 mm. long. of disk about 1.5 m m. long. plains and mountainsides, a Zones; southeastern Washington and eastern Oregon south along the Califo Mojave Desert ‘and San Diego County; more abundant eastward to the Great Plains, from Saskatchewan to northern Mexico. Type locality: plains of the Missouri River. —Oct. 3. Gutierrezia microcéphala (DC.) A. Gray. Small-headed Matchweed. Fig. 5431. elie dara DC. Prod. 5: 313. 1836. Not Hook. 1837. Gutterr. microcephala A. Gray, Mem. Amer. Acad. II. 4: 74. 1849. pasttctaed euthamiae var. eatiroetihel A Gray, Syn. Fl. N. Amer. 12: 115. 1884. * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 261 WIZ Wi. Sit Se i Pie ‘ly Ohare CA She 3 Y doo ae BES 5428. Grindelia robusta 5430. Gutierrezia sarothrae 5429. Gutierrezia californica 5431. Gutierrezia microcep. Xanthocephalum — Greene, ve e 282. 1892. Gutierrezia lucida Greene, Fl. Fran Gutierrezia glomerella biwe ene, tig €: cs u Gutierrezia sarothrae var. microcephala L. Benson, keer: Journ. ~~ 30: 631. 1943. Xanthocephalum microcephalum tae Field & Lab. 18: 29 Many-stemmed, 3-6 dm. high, seb resinous, cnt glabrous, si slender stems striate- angled, much-branched above. Leaves 2-5 cm. long, 0. e 2m wae frvaee 2 — ed; in — rescence cymose-paniculate, the see heads about 3 mm. hi si wide: eect" terminal glomerules of 2-5 or a few solitary ; phyllaries yellowish white eith yellow t tips, the teealitie rei prominent, the inner row of only 2 a as long as the disk-florets ; ray- and disk-florets 1-2 each, the ligule up to 2.5 mm. lon ay-achenes fertile, appressed- ios, their pappus of about 8 oblong paleae 0.8 mm. Pa Sh okcoee of about 12 paleae 1 mm. lon desert, Sonoran Zones; Mojave Desert from White ee a0 oe nty, to Palmdale and Little + pepe pic i ocpietccigess . California; east to Colorado, Texas, and northern Mexico. Type locality: Saltillo, ug 70. AMPHIPAPPUS* Torr. & Gray, Bost. Journ. Nat. Hist. 5: 107. 1845. Low shrub with divaricate spinescent branchlets. Leaves alternate, small, entire, oval to elliptic or obovate, short-petioled. Heads radiate, few-flowered, small, glomerate at tips * Text contributed by David Daniels Keck. 262 COMPOSITAE of branchlets. Involucre obovoid, — 3-seriate, strongly graduate, pale, the 7-12 broad rounded phyllaries thin, with scarious erose margins. Receptacle fimbrillate. Ray-florets 1-2, pale yellow, small, pistillate, fertile. Disk-florets 4-7, perfect, sterile. Anthers nar- ) 5 tortuous, hispidulous, irregularly basally united, white paleae of unequal width, ae the corolla. [Name Greek; amphi, both (kinds of), RE 5 pappos, pappus. | A monotypic genus. 1. Amphipappus fremontii Torr. & Gray. Chaff-bush. Fig. 5432. Amphipappus fremontii Torr. & Gray, Bost. Journ. Nat. Hist. 5: 108. 1845. Amphiachyris fremontit A. Gray, Proc. Amer. Acad. 8: 633. 1873. Much-branched, white-barked shrub 3-6 dm. high with yellow- -green cast, glabrous throughout, slightly ee te around heads. Leaves 5-12 mm. long, 2-4 mm. wide, obtuse or acute, l-nerved; hea , the phyllaries closely app pressed. pen igh ss ve an a atiitine flats, Lower Sonoran Zone; Death Valley region, California, from Inyo and Argus Ranges to Funeral Mountains and Renestep Valley, Inyo and San Bernardino Counties; east to Nevada. Type locality: m the M ohave abies a Apri 1—-May Am bhis ipappus fremonti bsp. spindsus tA. s.) Keck, Aliso 4: 102. 1958. aig, “eae: ga fremontit var. spinosa A. Nels. Bot. Gas. ‘47: ert 1909; , Amphipappus spinosa A. Nels. Pine Journ. Bot. 21: 579, 1934; tit var. spinosus C. L. T, op. 30: 483. 3.) Herbage densely scabrohispidulous. Similar . fremon sites, eastern Mojave Desert, California, and farthee south in ee Bernardino County; east to Utah and Arizona. Type locality: Moapa, Neva ada. 71. ACAMPTOPAPPUS* A. Gray, Proc. Amer. Acad. 8: 634. 1873. Low, much-branched, desert shrubs with white bark. Leaves alternate, small, usually spatulate or oblanceolate, entire, l-nerved. Heads yellow, radiate or discoid, subglobose, — Achenes subturbinate, short, densely villous. Pappus persistent, of about very flattened paleae and bristles of different whee the broader ones usually somewhat dist ae [ Name Greek, meaning stiff pappu s of 2 known species. Type species, Aplopappus Te licwislis Harv. & Gray. Heads discoid, small, gathered in small cymes. ae ea Heads radiate, large, solitary at tips of branches. ACS 1. Acamptopappus sphaerocéphalus (Harv. & Gray) A. Gray. Goldenhead. Fig. 5433 Aplopappus sphaerocephalus Harv. & Gray in A. Gray, Mem. seg aero Il. 4: 76. 1849. Acamptopappus sphaerocephalus A. Gray, Proc. Amer. Acad. 8: Round-topped shrub 2-9 dm. high, corymbosely MEE often iat twiggy, glabrous fa gsi or rages hispidulous o nm leaf-margins. Leaves linear et — ate, 5-20 mm. long, m. wide, obtuse to acute, eucronulate firm, sessile, gray-gre spears " subglobos e, 7- 10 mum. hig beste mostly liats but approximate at tips ‘of cymosely caaianl branchlets ; involucre -6 oe igh. Open Lower Sonoran Zone; eastern Mojave Desert and western border of Colorado Desert, Inyo Kot ee. to oe "nae *Comuty, California, east to Nevada, Utah, and Arizona. Type locality: ‘‘California.” pri Aca ampt opappus sphaerocephalus var. hirtéllus Blake, Journ. Wash. Acad. 19: 270. 1929. Stems ms leaves peneely herteflows. Western borders . eer Dessert to Lone Pine, Inyo County, California; wester Nevada. Type locality: Lone Pine. May-Jun 2. Acamptopappus shéckleyi A. Gray. Shockley Goldenhead. Fig. 5434. Acamptopappus shockleyi A. Gray, Proc. Amer. Acad. 17: 208. 1882. Rounded shrub 1.5—5 dm. high, poor ge pean the spoken: 6 finely hirtellous or hispidulous. —_ ge pm oblanceolate, oa . SHlipt 15 mm. long, u mucronulate; heads glo pati wide, solitary a adr or nearly naked Pebiaces: ; favolacre 8-11 mm. Eich; ra 8-13, the oblong. lieu le about 1 bso toes Desert and washes, Lower Sonoran Zone; White Mountains, Inyo County, to Clark Mountain, San Bernardino County, California; “Southern Nevada. Type locality: near Candelaria, Nevada. April-June. * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 263 72. CHRYSOPSIS* (Nutt.) Ell. Bot. S.C. & Ga. 2: 333. 1824. Low, oaatrhag ete annual), pubescent herbs, sometimes suffrutescent. Leaves alternate, usually entire. Heads radiate or discoid, yellow, medium -sized, jemtilhattng the stems and branches. Involucre campanulate to hemispheric; phyll merous, n and imbricated. Receptacle low-convex, foveol ay-flowers pistillate, fertile; ligules narrow isk-flowers perfect, fertile, slender. Style-branches flattened, th endages hairy, elongated, much longer than the sti tic porti blo r obovate, less flattened, often twisted. Pappus usually o the of r- sh, the inner ous capillary bristles, the outer (when present) of short linear scales (or bristles). [Name Greek, meaning golden aspect, from the color of the heads. A genus of about 20 species, all native in temperate North America. Type species, Chrysopsis mariana (L.) Ell. Heads radiate; outer ree seat ar sr 1. C. villosa, Heads discoid; outer pappu ne or indistinct. orolla filiform; pappus leciuiien the involucres by most of its length; mostly below 3,000 feet. 2. C. oregona. Corolla funnelform; pappus oe the involucres by less than half its length; above 4,500 74 . brewer. 5432. Amphipappus fremontii 5433. Acamptopappus 2 eseaeeseatia 5434. Acamptopappus — kleyi 5435. Chrysopsis villos 1. Chrysopsis villésa (Pursh) Nutt. ex DC. Hairy Golden-aster. Fig. 5435. . 1836. Diplogon villosum Kuntze, Rev. Gen. Pl. 1: 334. 1891. Gray-green perennial herb with several erect stems from an often more or less woody base * Text contributed by David Daniels Keck. 264 COMPOSITAE ; meer a taproot, 1-5 dm. high, the herba ntly strigose and often spmiey int giandalas, Lea yeas ee -spatulate, e Hedy 1-5 cm. long, ree cathe spe smaller, firmer, and m sessile re the ea -de 1 ones ; heads paniculate or cymose in a shor ti ote: sae Nuokets 7- stg Fs See | eS Soantete niredions. rays 10-16, Beconane revolute from the tip; sehcnes oblong- -ovate, villous; outer pappus evident. Open, well-drained slopes, often in sand, Upper Sonoran and Transition Zones; both sides of the Cascade Mountains hinge southern Beta Columbia to southern Oregon, reported from the ‘northern Sierra Nevada in Ca rape nia east to the Great Plains. Type locality: “On the banks of the Missouri.” June—Sep rysopsis gers var. sessiliflora (Nutt.) A. Gray, Syn. Fl. N. Amer. Fei 123. 1884. i han 9 sessilior Nutt. Trans. Amer. Phil. Soc. II. 7: 317. 1840; C. californica Elm Bot. AR = Heterotheca sessilifiora Shinners, Field & Lab. 19: 71. 1951.) Herbage mostly eG yish, vil ti pee Fides "leaves oblong or spatulate, crowded, 1-2 pe long; heads mostly pirates and wrt Misongs Tolinats Geacereten outer pappus evident, squamellate. Brushy isand- flats, washes, etc., Upper Sonoran Zone; usually near the coast from Santa Barbara, Ca + fornia, to norther: wer California; reported north to Mendocino Soniey, California. : “St. Barb ara, Upper Californie July—Sept. mpeopais villosa var. bolanderi (A. Gray) A. Grae ex Jep Man. Fl. Pl. Calif. 1036. 1925. (Chrysop- sis bolande ert A. Gray, Proc. Amer. Acad. 6: 543. 1865; C. sessiliflord var. bolanderi = ve ray, Bot. —_ £22 309, ate hirsute and m or less de nsely beset with sessile glands; leaves oblanceolate or spatulate, rounded, mucronate, the eat =Hrcnc he 2-4 cm. long, 5-10 mm. wide; heads large, mostly solitary, foliose-bracteate; involucre 10-15 ps high ees a en acuminate, villous and glandular; outer pappus of very slender scales. Gra slop unes on ear the coast, Humid Transition Zone; Mendocino County to Prat Francisco Bay, California. “Tye esc iis of Oakland, near San Francisco. % June— eu Chryso po casbie (Greene) H. M. Hall, Univ. Calif Bot 3: 42. 1907. (Chrysopsis fastigiata ence Pitona a 7396. 1898.) Sows 3-10 dm sick. ‘ee Zs ES cliheaached, usually closely beset with — —— linear- oblong to be pa obtuse or acute, mucronate aging i sinc tomentose — dorsally, 1-2 0 cm. age the margins often c Segre heads s mall, 1-3 a ae stiff ere a Counties and from Ventura Count y to Orange County ‘Tyne loc a Fhe vicinity of San Bernardino, San Bernar- Vv. Chrysopsis villosa var, echioides (Benth.) A. Gray, 3 See 12; 123. 1884. I 6 echiot- bt Benth. he t. Sulph. 25. 18442, Cc. paste ae var. echioide. es "Bok: Calif. 22 309.- 1876: rotheca hioides Shinners, Field & Lab. 19: 71, 1951.) Stems sated 3-8 dm. high, virgate, simple or (gotten fhraceende: Wires crowded, ae ca aa! hirsute- igre nt, firm, ‘t the u perils, 2 _ nee sessile; heads Saag numerous, paniculate to cymose yllari numerous, sle nder, hispid-hirsute pappus. of short narrow squame Dry sandy soils eg sag ag Upper Sonceun oti Coast Ranges ‘of “California mostly away from the Sevenedinte: oat from Sonoma and Solano Counties to San Dieg: o County, and also adjacent Lower California. age : Bodegas [B paces Bay], cenbenn Loa — —Nov Chrysopsi i var. hispida (Hook af A. Gra Bot. King Expl. 164. 1871. (Diplo- pappus Tespudus Hook. Fl. Bor. Amer. 2: 22. 1834: Ciecpet hispida DC. Prod. 7}: 279: is GC. Race teos Greene, Erythea 2: 95. 1894 re. hirsuta Gre Pittonia 3: 296. 1898.) Stems slender, virgate, 2-4 hi herbage spa arsely or densely spreading- hr somewhat glandular; leaves eo oblanceolate or ‘Spatulte, 1-4 cm. long, mm. wide, often hirsute-ciliate; eats solitary to numerou olde sid aereue re 6— + 3 ai phyllaries minutely glandular and sparsely hispid: outer pappus of very squamella Dy ocky ground, Upper Sonoran and Arid Peansition Zones: Saskatchewan to British re aia var a tha aT cetade Mocuintna south to the Sierra Nevada and Colorado Desert of California (rare), east to Arizona and the Roc = fete tains. Type locality: Carlton House Fort, Saskatchewan. April—Nov sopsis villosa var. camphorata (Eastw.) Jepson, Man. FI. Pl. Calif. 1036, _- Piet: -sag'sd cam- sre Eiee, Zoe 5: 81. 1900.) Stems slender, from a slender branching rootstock, ne herbage ee id —— nsely hirsute and conspi icuously glandular-pubescent, poate "ed ves lance- oblong. eads usually many, cymose-paniculate, rather a, usually “Subtended by small faeesn bracts: sistiaeen aparvely hispid and ciliate or merely ciliolate from the fine dissection of the narrow hyaline margin , prominently ¢ andular. Open ane slopes and wooded banks, Humi d Teeneition Zone; Santa Clara County to San Benito cor ha Counties, California. Type locality: near Glenwood, Santa im Mountains, Santa Cruz County. July—O a ie ae oregona (Nutt.) A. Gray. Oregon Golden-aster. Fig. 5436. 4 9 Nutt. Trans. Amer. Phil. Soc. II. 7: 321. 1840. Ch Psi: A. Gray, Proc. Amer. Acad. 6: 543. 1865. sw wreeinn Kuntze, a Gen. Pl. 1: 334. 1891. Heterotheca oregona Shinners, Field & Lab. 19: 71. 1951. Stems clustered from a woody base, much branched, erect, 3-6 dm. high, the ee glabrate or sparsely hirsute, rather sau | Eggs - eaves = ceolate to elliptic-ovate, sessile, ascending, I-nerved, 24 cm. | 0(15) mm. wide; heads corymbosely pan culate, discoid ; Re clncr re 9-12 igh; ; phyllaries ‘-seriate vaaanite saint glandular-atomifer s but scarcely ae ae with white hyaline margin, acuminate; corolla nearly filiform; outer pap ig aioe and obsc Sandy or gravelly stream banks, Humid Transition Zone; Washington chiefly west of the Cascade M tains wens to =e tg California. ype locality: “On the sand and gravel bars of the Oregon (Cstumbial and its acs June—Sept og sis —— var. ridis (Greene) Jepson, Fl. W. Mid. Calif. 558. 1901. (Chrysopsis . are. e, 1 eg. 174. 1894.) Stems 3-8 dm. high, sing bee Lae Raghnenadl aa. hispid -hirsute and m r less one ican lnrguhapeceiate oes dly oblong or oblan “a or ‘spreading, * 5 em long; involucre 8-1 hi ate, saacatar and som tt at his spi ae mage e. war 9 eid Transition Zone; California in the ibd ano cae Nevada Sed mine c eee) Real North Coast anges from aa ha and boldt Counties to Sa Francisco Bay. Type loc m banks pa Co.” July—Oct. hrysopsis oregona var. compacta Keck, pe on 102, prokg eve p ended ie 1.5-4 ree high, with pene — —— the woody base herbage gray-green, apeoris hispid- or lives cancepent fe" glandular-atomifer- e hair: prominent pustulate sir age leaves typienily small (1-2 cm.), densely crowded, spreading, once se og Taig couard tow ard the ends of the br aches: involucre 8- 10 mae: high, glandular but usually sparingly hairy. Dry sandy pla Upper Soaren Zone; Central vailee of California from Tehama County to Fresno County. Type ieeneys "Newsille: Tehama County. June—Oct psis oregona var. scabérrima A. Gray, Syn. Fl. N. Am 124. 1884. we rather openly ening: the heads commonly corymbosely paniculate on slender Pritieha: leaves in part of plant becoming bract-like; herbage visc salons gd ne scabrous from the pustulate ber of ike gy Baenihion hairs. Sandy roadsides and dry ta Uppe oran Zone; California in the South Coast Ranges and Sierran foothills (Tulare County). Type loca lity: eae County. Oct. SUNFLOWER FAMILY 265 3. Chrysopsis bréweri A. Gray. Brewer’s Golden-aster. Fig. 5437. Chrysopsis brewerit A. Gray, Proc. gre Acad. 6: 542. 1865. Chrysopsis wrightiit A. Gray, Syn. Fl. N. Amer. 12: 445. 1884. Heterotheca breweri Shidneda. aor. & Lab. 19: 71. 1951. t stems from a woody caudex, 2-8 dm. high, simple or more commonl ate 3-nerved from base; heads solitary at the ends of the slender le afy br anches, broadly campanulate, is 1 mm. hi iscoid ; involucre 7-1 igh, 2—3-seriate ; phyvere s lance-linear he lance- oblong, acuminate ; corollas slender-funnelform; outer pappus of short rit bri stles or no Open rocky slopes and coniferous forests, Canadian and Hudsonian ear Sierra Nevada from ae Cicaky to ulate Caaas and on San Gorgonio Peak, eet adjacent Nevada. Type locality: ‘‘Nea Sonora Pass and same 8 Pass, in the Sierra Nevada.” July-S Chr sis ver var. multibracteata Jepson, nies re Pi; ’ Calif. (3037. 1925. ee, “er ba Eastw. Bot’ Gaz [: 291. 1906.) Herbage sparingly arachnoid-villous; phyllaries somewhat thic owly — to eebeclaity 4 acute or acuminate, sometimes purple-margined, 4- 5 -seriate. Dry rocky irl Canedinn ne; ware aay a bg inity, and Tehama Counties, California. Type locality: Sisson (Mt. Shasta City), Siskiyou 5436. Chrysopsi 5437. Chrysopsis breweri 73. HETEROTHECA* Cass. Bull. Soc. Philom. 137. 1817. Coarse erect herbs with yellow-flowered heads disposed i in terminal corymbose panicles. Leaves alternate. Involucre hemispheric; phyllaries narrow, closely imbricate in bt eral series, appressed. Ray-florets 1-seriate, fertile; disk-fl many, ferti i ene triangular-compressed ; pappus none or caducous. Disk-achenes cuneiform ; pa ouble, the copious inner bristles Selig long, the outer setose, short. ame Grek: meaning es mn ig: (or ovary), from the unlike achenes of ray and disk ore species in the esis United States and Mexico. Type species, Heterotheca lamarckit dea” = "H. pa ep (Lam.) Britt. & Rusby). Upper leaves narrowed to a sessile base; heads relatively large; involucre 7-10 mm. high, gy pede ree hes? + } + aan : : lu 6-8 mm. high, glandular- Lg ae and also a aoe sis ee. ep Seana tmeraetimanrny A. subaxillaris. 1. Heterotheca grandifléra Nutt. Telegraph Weed. Fig. 5438. Nutt. Trans. Amer. Phil. te II. 7: 315. 1840. Heterotheca forbade ise Bot. Sulph. 24. 184 Annual or biennial, the stout stem simple below, 5-20 dm. high, hirsute, the ample inflorescence peo is Set nt and heavy-scented. Leaves thickish, villous, ovate to pues lobes at bas te, —6 cm. long, evo, serrate, the lower eats, commonly with a cag of stipule- sn . = at base; heads medium large; involucre 7-9 mm. high; ray-florets 25-3 35, the corolla é. meh wide, revolute from the tip, the tube hairy; disk-florets 50-65, very sender, the iubbe sper ranches scarcely exserted; pappus brick-red, the outer series goo : ee ee southern aliro’ ia n ] San 5 oe re A Lag ying sn ir este labs s; also southern heiaanes Type locality: Santa Barbara. March—Jan * Text contributed by David Daniels Keck. 266 COMPOSITAE 2. Heterotheca subaxillaris (Lam.) Britt. & Rusby. Camphor Weed. Fig. 5439. Inula eras aed rages Encycl. Meth. 3: 259. 1789. Inula scabra Pursh, Fl. Amer, Se Ene 2: 531. Chrysopsis si posh t. Gen, 2: 1818. Heterotheca lamarckii Cass. “AS sing Nat. 212-331. 123), Heterotheca scabra DC. Prod. 5: 317. 1836. Heterotheca subazillaris Britt. & Rusby, Trans, N.Y. Acad. 7: 10. 1887. Annual or biennial, the eae pane ge stem simple below or openly branching, ee sit me hispid-hirsute, glandular above. rather coa orb eto glandular, ov aoe to lance-oblon serrate-dentate or subentire, the lower spetiolate, the r subcordate- clasp Heiay relatively small, numerous; involucre 6- 8 m igh, g eadelee and: somewhat canescent; ’ ray-florets about 20-28 ; ganar 40-60 : pappus fa sy the r serie. ‘sually conspicu Sandy des and ditches, Sonoran Page easternmost Mojave Desert in eof: intepenert at San Gabriel, Cotitonins common eastward to Florida and Delaware. Type locality: Carolina. "Aug.— —Oct 74. CHAETOPAPPA* DC. Prod. 5: 301. 1836. Ours low, very slender annuals with simple or diffusely branched stems and alternate, entire, chiefly linear leaves. Heads small, few- to many-flowered, terminating very slender peduncles, radiate, disciform, or discoid all flowers potentially fertile, yellow, white, or eddish ‘volu cent. Pappus of 3 to many fragile slender bristles, sometimes dilated and more or less joined at very base, or wanting. [Name Greek, meaning bristle-pappus. ] A genus of about 15 species of the southwestern United States and Mexico. Type species, Chaetanthera asteroides Nutt Ray-fl Pp t, conspicuous; involucre broadly hemispheric. Ligul Id llow ; southern California jipenkeese pubéscent. 1. C. lyonii. Involucre glabrous. Phyllaries in strongly satis alg scsi to the short-caudate tip. 2. hyllaries in subequal se obtuse, nulat ess itis. Ligules white ed ne — naeee Franc npg ae region. 4. C. bellidiflora. Ray-fl sent; involucre tu rbin: Stem simple or with few tet ‘iancien: disk-corollas ind at throat, contracted at sola oc -. ext Stem diffusely branched; disk-corollas narrowly linear. iC. ak 1. Chaetopappa spe (A. Gray) Keck. Lyon’s Chaetopappa. Fig. 5440. Pentachaeta lyonii A. Gray, Syn. Fl. N. Amer. 12: 445. 1884. Chaetopappa lyonti Keck, yesh ei 958. Stem 1-5 dm linear or Stl ne it ce aoe . long ; involucre about m. high, the ptr pares: mpbesa:. lance-linear, acuminate, n zane scarious-margined ; fay Mo ioe abou 0; -flowers about 00 ; achenes of ray tor disk similar, d rown, oicaty one with short hairs; ; paps of 10-12 ve ery fragile, filiform bristles, flared at ver y base and forming a rudimentary c In clayey soil of grassland areas, Upper Sonoran Zone; coastal.part of Los Angeles County, Rovio’ and Santa Catalina Island, o cases. ppd Type locality: San Pedro, near Palos Verdes Mountain. March-April. 2. Chaetopappa atirea (Nutt.) Keck. Golden Chaetopappa. Fig. 5441. nceeareeigicra aurea ns Trans. Amer. Phil. Soc. II. 7: 336. 1840. eck, Aliso 4: 102. Usual y — branched, 8-30 cm. high, entirely gatey, or the leaf-margins paul ciliate. Leaves > ar linear, the lower 1-3.5 cm. lon o 2mm. wide, the upper m me pies hi i obtuse a 1, ean a ually 10-40, the ligule 5-12 mm. long ; disk-flowers usually —— achenes abdist on m. long, sparsely and finely short-strigose ; pappus of 5(-7) scabrous filiform i rm bristles + Anse diickened toward apex, flared at very base and often face about equaling the c Dy open ound and grassy ar oe up to 5,000 feet, mostly wha Sonoran Zone; cismontane southern Cali- fornia from pices anantes County to Diego, San Diego County ype locality: San Diego. April—July. * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 5438. Heterotheca grahdiflora 5441. Chaetopappa aurea 5439, Heterotheca subaxillari 5442. Chaetopappa fragilis 5440, Chaetopappa lyonii . Chaetopappa bellidiflora 267 268 COMPOSITAE 3: hn aL slg. Shee (Brandg.) Keck. Fragile Chaetopappa. Fig. 5442. Pentachaeta fragilis Bra: n H. M. Hall, Univ. Calif. Pub. Bot. 6: 170. Chaetopappa fragilis Spe “Aliso 4: 102. 1958. slender, wiry, diffusely Pies glabrous except for the hirsutulous leaf-margins, the lon al 8-15 mm. lon branche 4-10 cm. long. Leaves spatulate to linear-oblanceolate, the bas wide, obtuse, the caulin 8 ide; volucre a high, the Sia ilaries subequal, ee as ee sea * agierire scarious-margined, lacerate-ciliate at apex; ray- flowers a 0, achenes sparsely pilose 20 very fragile, . the en ; papp filiform rivties ies por leery econ apex, not at all dilated at base, slightly shorter than the disk-corolla. assy, foothill slopes, Upper Sonoran Zone; southern Sierra Nevada and Greenhorn Mountains, Kern County ane ge Taner South Coast Range, San Luis Obispo County, California. Type locality: Havilah, Kern County. ay— 4. Chaetopappa bellidifiora Shey Keck. White-rayed Chaetopappa. Fig. 5443, Pentachaeta bellidifilora Greene, a: Calif. spa 1: 86. 1885. Chaetopappa bellidiflora Keck, Aliso 4: 102. Stem 6-15 cm. = simple or with a erect branches, = ea or nearly so. Leaves narrowly linear, 8-35 mm. lon mm. wide; heads 1-1.7 cm. wide; involucre 3-4 mm. high, the sid laries "subequal, after aeepints oblong, short-acuminate or apiculate 5 a the truncate, icfoe. ciliate ADE, ere scaious-margined A cla us on ly; ray- —16, white or purplish- noes e ligule 5 mm. long; disk-flowers numero ee the ample trae not contracted at hee ERICH yairate, rarely gla ai ; pappus of 5 ee vely firm, scabrous bristles, sce dilated at pe shoei than the disk- cordila: obtezatonally lac Open, dry, rocky slopes, Humid Transition Zone; Marin, San Mateo, and Prise ia Counties, California; also seporied from Monterey County. Type locality: Corte Madera, Marin County. March—May. 5. Chaetopappa éxilis (A. Gray) Keck. Meager Chaetopappa. Fig. 5444. Aphantochaeta exilis A. cae Pacif. R. Rep. 4: 100. Pl. 11. Fig. A. 1857. oe ilis A. Gray, Proc. Amer. Acad. 8: 633. 1873. Pentachaeta exilis var. shen Gihawe Gray, Bot. — 1: 305. 1876. Pentachaeta aphantochaeta Greene, Bot. Gaz. 8: 256. Pentachaeta es ar. grayi Jepson, Man. FI. Pl. ody ae 1925. s K , Aliso 4: 103. m 318 on bine simple or with few erect branches, often purplish, sparsely pubescent. mm. long, ciliate-pubescent ba cle white- Villoa: Ratio the head involu , glabrous, often purplish, the phyllaries subequal, , broad, oblong, wea she: bristle- inge at the obtuse or t ate, mo 1 iate apex, moderately s a gined; pistillate flo 0-5, be a nd x Les rt filiform tube; disk- fl s 4-8, purplish, distinctly widened at throat, contracted a nes brown, moderately illous or rarely glabrous; pappus of 3-5 sl oe aig ee cine: os “dilated at base, or these Vv ro reduced to triangular scales, or often entirely wa Grassy slopes of valleys one foothills, Sonoran and ae Zones; from Placer and Mendocino Counties to Mariposa and Monterey Counties, California. Type locality: Napa Valley, Napa County. April-May 5444. Chaetopappa exilis 5445. Chaetopappa alsinoides SUNFLOWER FAMILY 269 6. Chaetopappa alsinoides (Greene) Kéck. Tiny Chaetopappa. Fig. 5445. Pentachaeta exilis var. discoidea A. Gray, Bot. Calif. a Foes 1876, in part. Pentachaeta alsinoides Greene, Bull. WES sate 1882. Chaetopappa alsinoides Keck, ‘Aliso 4: pK i 8. sie ies branched, 3-12 cm. high and wide, somewhat villous. Leaves narrowly linear or fili- form, 1 mm. wide or less ; bende t tiny, not strictly solitary; involucre 2.6-3 mm. high, glabrous or ual, few (6-7), oblon 1 se aaa sli light ly thicker, reddish-tinged disk-flowers, these only erfect anthers ; achenes brownish, lightly to moderately villous; nine seni | cf 3 ahr ea bristles Pesan bb exceeding bias orets, AS at all dilated at base oothill slopes, Upper Sonoran Zone; gant near : ie coast, Napa and M Counties t Santa rey ow and Tular . Courties. golfer nia. Very inconspicuous peppers mT agp a ore ps gi i he locality: Berkeley, Alameda County. April-— g 75. HAPLOPAPPUS* Cass. Dict. Sci. Nat. 56: 168. 1828. or shrubs, very varied in habit, often glandular. Leaves alternate, entire to bi- stsuattee often thickish, sometimes glandular- -punctate. Heads radiate or disc oid, large cymose i it gr TOW ur alveolate. Ray- 7 sa Distillate, rarely sterile; disk-florets fertile, their style-branches ovate to subulate. Achenes terete or angled, linear-fusiform to turbin ate, glabrous to silky- pilose. Pappus of numerous ca Oi an , subequal or graduate bristles, usually persistent. [Name Greek, meaning simple eae pus. genus of neh ade? 150 ppcies, all Be chiefly of the western United States, Mexico, and Chile. Type species, Haplopapp Achenes turbinate, 2-3 mm. long; leaves dentate to bipinnatifid, the teeth spinulose-tipped or bristle-tipped. (Section Diepuarodo mn.) Annual; herbage, including involucre, hirsute with strigose hairs, the phyllaries pores minutely glandular- berulent. . H. gracilts. Perennials; involucres not strigose. Tufted ae seve sa slender ff: Ss very narrow; heads radiate. Phyllaries ccaaiieailt ene Phen Te and scabrous; sbaee 25 em. Jon "2. H. gooddingis. Phyllaries beset with granular glands; leaves 1—2 cm. Jon . H. junceus. Rigidly branched shrub; leaves mostly oval, — dentate; ree disc . H. brickelliotdes. Achenes nearly prismatic, subcylindric, or fusiform, 3 mm more long; sae various aa if toothed these not as above Perennial herbs with shoots of the season arising from prominent basal rosettes of leaves surmounting a deep fusiform taproot. (Section Pyrrocoma.) Heads tea ml ~ involucre mostly 1.5—3 cm. high, the phyllaries mostly 3-8 mm. wide, the rays incon- eng see 4 cm. wide; involucre 14-20 mm. high; widesp 5: carthamoides. Basal piel 5- ib cg wide; involucre about 25 mm. high; sist very robust; upper en =< = Snake any —. iain, ic raat mostly 5-15 mm. high, the phyllaries less than 3 mm. wide, the rays gee Oe i nflor CoCenice, eads ald or cymose, or if Z Plants stipitate-glandular as well as hirsute or villous. . A. hirtus. Plants not cove ndul He lly solitary, terminating long peduncles. , thin, acres graduate; achenes “sah peer . H. unifiorus. Phyllari only toward tip, firm, evidently graduate; achenes ee us. ¢ 7 9. H. apargioides Heads corymbose or c 1 pai rarely solitary; phyllaries = toward base, green- tipped; ge seric . H. lanceolatus. Heads usually numerous, spica Plants poor strongly if at ae Sader phyllaries coriaceous and tet ee Secon ee to Fignes vernicose from sessile glands; phyllaries coriaceous-herbaceous, obscurely igraduate acu- 4S. Shrubs or penetra. Pp fted, usually mat- forming, with much-branched caudex; heads mostly solitary at ends of oseal eit several in H. whitneyt). Stems relatively leafy; on glandular-puberulent; plants o: f high mountains. (Section Tonestus.) hyllaries lanceolate; echoues glabrous or sparsely pilose. - 13. H. lyallis. Leaves toothed. Achenes ses ig leaves sharply serrate; involucre campanulate-oblong ; gad atea linear- lanceolate; stems to 50 cm. high. 14. H. whitneyi. Achenes eet pubescent; leaves saliently dentate in outer half; involucre hemispheric; stems to 15 cm. high. * Text contributed by David Daniels Keck. 270 COMPOSITAE Involucre 7.5-10 mm. high, 12-15 mm. wide (pressed), shorter than the disk-florets, pies outer go doterlony meee oblong to oblong-obovate, ces ee? vel Peet disk- oro aie 6-7 m sure Involucre Jt 18 m 0-30 mm. wide fpresse@) « sine hd the ‘ai his , the Ghsflaries ee or lance-oblong; or ees disk- corolla, 9-10 eho ‘0 ong. wit. peir Stems fants. (Sect Steno) herbage scarcely if at all glandular; “ethos entire; sit dined pla t Leaves soft, floccose-tomentose 17. H. lanuginosus. Tons rigid, glabrou ne scabrous pag: Leaves linear, 7—1 m. long, 1 m ig wa wide. 18. H. stenophylius. Leaves fener PSA to spain IR 10-60 mm. long, 1.5-7 mm. wide. 19. H. acaulis. Plants not at all cespitose or mat-forming, but sometimes forming low cians ers Appen ee of s tyle-branches at least twice as long as the stigmatic port Plants low, vik icately branched shrubs mostly under 3 dm. high. (Se came Macronema.) wigs closely white-tomentose; heads discoid; leaves densely glandular — macronema. glandular, if rarely loosely tomentose then heads radia Seeds relatively large, 20-45-flowered; involucre broadly campanlate, — 2- ee e phyllaries not obviously ciliate or scarious-mar. gined; lea undu i - 5% jer ned; herbage densely stipitate-glandular rutt Heads smaller and narrower; involucre more hea 2-seriate; nanan ae eek very rarely stipitate-glandular. fenton villous-ciliate; leaves without axi ~— — cles. volucre narrowly campanulate, 3—6-ser the enti phyllaries not squar- rose-tipped; heads radi now aghat pein ce ponte Spencenes” 1.5-3.5 cm. Peta 3-7 mm. yg = A rae or 22: Leaves Baga linear, 2-6 cm. long, 0.5-3 mm. Gack ead ieietal or erous in “4 raceme ve aurea panicle. omer Involucre cylindric, 5—8-seriate, the outer phyllar x sqarone tipped heads discoid, mostly 5- flowered, solitary oi few, openly ¢) bap oo 0 ts. en — (obscurely ciliate in H. gilmanit) ; ‘cece often with axillary Pong as spatulate, spaced; phyllaries resinous, the outer squarrose; flowers white (or pale yellow); disk-florets 15-18; Death Valley region. Leaves cea linear or filiform, crowded; phyllaries not pedkivnds. Out se dove ys often squarrose; flowers white (or pale yellow); disk- florets 10-15, their narrow lobes 1-2 mm. — Washington and Outer phyllaries erect; flowers yellow; oe — ae au 0.5 ; Great Basin. 27. H. nanus Plant an herbaceous perennial pty a ce base, mostly 3-6 dm. high. — Hesperodoria. ) . A. halli Appenda ges O of s b h or sho ti Involucre hemispheric, 10-18 1 mm. spare se tlt i -punetate; leaves entire. (Section Stenotopsis. ae 29. H. linearifolius. Involucre turbin subcylindric. Disk-c corolla ab abruptly dilated from narrow tube to much wider throat; heads discoid. (Section Phyllaries with green but thin tips; stems brownish; coastal. 30. H. venetus. Phyllaries with thickened subepidermal resin-pocket near tip, not green; stems age 31. H. acradeniu ~~" only slightly ampliate upw Heads large, the 8-15 mm . high manera tightly imbricate, mostly 6—8-seriate; herbage ithout distinc ves resin-pits. (Section Hazardia.) e tomen snare an, lec € not to; cuaaetvans ws inland. 33.20 Hs Heads ‘onal, , the 3-8 mm. ‘igh Se se imbricate, Pe Stig pis “with ‘cam: Ray-florets resent: leaves filiform; heads more or less paniculate. ter phyllaries more or less caudate-t Leaves 10-35 mm. long, with gente fauldles § in ea axils; achenes pilose. me I ‘pin nifolius. Leaves 4-12 mm. — scarcely exceeding the salle, fascicles, ericoid; achenes glabro' . H. ericoides. Outer phyllaries acute to ove use. Heads solitary or few; invol p , 7-8 mm. pe. 36. eastwoodiae. Heads many; involucre turbinate, 5—6.5 mm. high. 37. H. palmeri. Ray-florets reduced or wanting (see — laricifolius). - soar: filiform to cha less than 3 mm eads solitary or racemose- CRISS. ienid: cer ee County south, . propinquus. Heads c: cymose. onde 0. Wich hiss long; broad rounded shrubs seldom more than 1 serts, Herbage glabrous; leaves subterete, without persistent fascicles in the old axils; ray-florets 3-11; io flowering. fae o Far laricifolius. SUNFLOWER FAMILY 271 Herbage hairy; leaves Sst, with persistent fascicles in the old axils; ray—florets 0-2; spring-flowering 0. eo cooperi. Leaves a = omg a arborescent shrubs mostly 1-3 m. high; cis- 41, H. arborescens. hl lat 3 oa 3 to oDoVate, 2—- 10 mm. wide. Heads 9-12-flowered, discoid; erect shrub 2-5 m, high; leaves 2-6 cm. long ey tapering to base and apex. 42. H. parishii. Heads Regs flowered, sometimes radiate; spreading cnr montty less a ka leaves cuneate, 43, H. cuneatus. 1. Haplopappus gracilis oe A. aig Annual Bristleweed. Fig. 5446. Distr wronties a fags Acad. Phila. II. 1: ay, Mem. Amer. ke bs 4: “3 1849, As tev d ieteria : Kuntze e, ee v. Gen. Pl. 1: 315. wg Brice carpal suger pig Erythea 2: 109. 18 Nels. Bot. Gaz. 37: fae 1 4, Méchaeranihare sine Shinners, Field & Lab. 18: 41. 1950. Annual herb 6-35 cm. high, yond divaricately branching throughout, hirsute, usually strigose. Leaves numerous, lerleéte the lowe a ceolate, pinnatifid or bipinnatifid, 1. 5-3 cm. 1 , mm. wide, the upper linear, appres reduce ; — aos gn to serrulate, euch tooth or lobe and apex tipped with a prominent white bristle; heads cymose or solitary; inv — re hemi- spheric, 6-7 mm. high, 8-12 mm. wide; phyllaries finiea Aguas: wel imbricated, green with hyaline ee cinereous or strigose ‘aad ainanely: se ular- puberulent, with adureabed bristle- tip; ray-florets 16-28, the ligules 7-12 mm. long; pappus of numerous tawny unequal bristles slightly dilated below Sandy or rocky flat eae slopes, Lower Sonoran Zone; eastern Mojave Desert from Clark Mountain to Provi- dence Mashtaloe Ban Reroadetae County, California, east to Colorado, Texas, and Mexico. Type locality; near Santa Fe, New Mexico. April-June. 2. Haplopappus gooddingii (A. Nels.) Munz & Jtn. Goodding’s Bristleweed. re 5447.: pescerhraare. gooddingti A. Nels. Bot. Gaz. 37: 266. aplopappus qqoaamoe gone & Jtn. Bull. peeik cua 49: 44, Pe t i Blake, Contr. U.S. Nat. Herb. a 543. 1925. Haplopappus spinulosus ome: gooddingii H. “4 Hall, Carnegie Inst. Wash. Publ. No. 389: 75. 1928. Taprooted perennial with ie stiffly erect or ascending, slender stems 2-6 dm. high, slabrate to harshly pra lar-puberulen metimes also canescent. Leaves scattered, lanate ventrally, = doraatn eeeniee, ee ge and remote lobes linear and bristle-tipped, 2-5 cm. toe. the upper becoming entire and much reduced; heads terminating long branches ; involucre depressed- paisa esineong 9 mm. hig 8 mm wide ; pera pea Tanceolat ate, well imbricated, Pe. prominently slandular-puberutent and “scabrous , with short apica a bristle; ray-florets 30-45, t ligules 6-16 mm. lon of numero 5 ed snceval brist Rocky mesas, ‘aie sae pe cliffs, ph: Sonoran Zone; eastern sie reaps 9 “ae to southern Nevada and northeastern Arizona. Type locality: ‘‘The Pockets,” southern Nevada. 5446. Haplopappus gracilis 5447. Haplopappus gooddingii 272 COMPOSITAE 3. Haplopappus jianceus Greene. Rush-like Bristleweed. Fig. 5448. Haplopappus junceus Greene, Bull. Calif. Sr pe 1: 190. 1885. Eriocarpum junceum Gr ae e, Eryt ee ee 894 Sideranthus juncens Da s. & M ns 3 Calif atts 1923. Machaeranthera juncea ermal ean & Lab. 18: 1950. Stems baat 4-10 dm. high, Aad ty t at base, slender, branching, spari ingly strigose, slightly glandular near the ‘heads. Leaves chiefly linear, pinnat tifid or serrate with bristle-tipped t ; : , slender y- bracted branches or in open cymes; involucre hemispheric, 5-8 mm. high, the closely imbricated phyllaries linear, covered with granular glands, bristle- e-tipped ray-florets 15-25, the ligules 5-6 mm. long; pappus of numerous tawny unequal bristles Dry brushy hillsides, Lower Sonoran Zone; southern San Diego County, California; northwestern Mexico. Type locality: San Diego County. June—Oct. 4. Haplopappus brickellioides Blake. Brickellia-like Goldenweed. Fig. 5449. Haplopappus brickellioides Blake, Proc. Biol. Soc. Wash, 35: 173. 1922. _ Rigidly branched shrub 25 cm. or more high, the older stems white-barked and more or less pilose, the branches yellowish, densely pilosulose, some hairs thickened and tipped with yellow glands. Leaves oval, ellie or obovate-cuneate, 1-3.5 cm. long, 5-25 mm. wide, acute, spinescent- tipped, dentate with 1-4 pairs of eric teeth, bes triplinerved, the midnerve prominent, pilose, and yellow-glandular shenda discoid, about 12- flow red, rather small, sessile or subsessile in ones to threes toward tips 0 of leafy shine i oy oti 8 ucre sa mm. high, the phylla ries 4-5-seriate, lanceo- late, l-ribbed, hispidulous and glandular, the tip greenish; eae oblong, hispidulous; pappus sparse. Rocky canyons, Lower Sonoran Zone; Death Valley region, California, and adjacent Nevada. Type locality: Ash Meadows (or Sheep Mountain), Nevada. April—Sept. 5. Haplopappus carthamoides (Hook.) A. Gray. Columbia Pyrrocoma. Fig. 5450 Pyrrocoma carthamoides Hook. Fl. Bor. Amer. 1: 307. pl. 107. 1833. Haplopappus carthamoides A. Gray, Proc. Acad. Phila. 1863: 65. 1864. c oid 17. 1 Hoorebekia carthamoides Piper, Contr. U.S. ‘i * ‘Herb 11: 559. 1906. pis eae rersnemoreys emis pee H. M. 1, Carnegie ee Svan Publ. No. 389: 103. 1928. acusacainee ae Hall, op. c aoe few, stout, erect or ascending from a thick mer 1-6 dm. high, glabiare to canescent- villous, few-leaved. Basal leaves tufted, 1-2(-4) dm. long, 1-4 cm. wide, oblanceolate to ova al, fe) miform inflor Ce; large f involucre hemispheric, 15-20 m vege often cubtcadie by a few por sae the anyllarics coriaceous-herbaceous, lance-oblong, hte e to acuminate, easpeeate, the margin obviously scarious and usually ciliate, entire to erose or seercieeek te -toothed ; ray-florets 0-30, ae seldom exc necting the pappus; achenes glabrous; pappus stiff, sordid. Prairies and open hillsides, Upper Sonoran and Arid Transition Zones; Columbia River Gorge, Washington- Oregon, north and east to northern Washington and western Montana. Type locality: ‘‘North West Coast of America.”” June—Aug. ry a carthamoides subsp. cusicki (A. Gray) H. M. Hall, Carnegie Inst. Wash. Publ. No. 389; 104. oes. ( ras much-branch initand caudex, 5-10 cm. high, aingg 4 clothed at base with the t leaves, the whole mat m up to several dm. ves linear-oblanceolate to piri pre iy erect, entire, chase to pave ty acuminate, caiapiate tipped, veiny, pale green, densely Pisa to glabrous except for t scabrid margin, 1-6 cm. long, 1.5-7 mm. eee: "involucr e hemispheric, 7-10 mm. high; Statics SUNFLOWER FAMILY 5460. Haplopappus 5463. Haplopappus stenophyllus 5461. Haplopappus peirsonii 53464. Haplopappus acaulis 5462. Haplopappus lanuginosus 5465. Haplopappus macronema 279 280 COMPOSITAE 2-3-seriate, broad, acute = acuminate, pallid; ray-florets 6-10, the ligules 6-10 mm. long; achenes densely sericeous to = us Dry ridges and pla KA Transition and Canadian Zones; central Oregon and eastern California south to Saye re east to ‘ae ‘Sie Mountains and Saskatchewan. Type locality: ‘“‘Little Goddin River,’’ Idaho, May- 20. Eewepapps macronéma A. Gray. Discoid Macronema. Fig. 5465. M id acy Trans. Amer. Phil. Soc. II. 7: tes 1841, Haplop Gray, Proc. Amer. ye 6: 542. 1865. Aster macronema Kuntze, on Gen Pl. 1: 318. 189 M. E. Jones, Proc. Calif. Acad. a. 5: Hablopappas discoideus Hall & Hall, Yosemite Fl. 246. Liha es qonls 1836. Undershrub 1-4 dm. high with numerous erect stems of the season from a multibranched woody base, the twigs masked by a white Sac sings m. Leaves numerous, oblong or oblanceolate, sessile, entire or undulate-margined, 1-3 c mm. wide, ats poy cand stipitate-glandu- lar; heads discoid, yellow, usually solitary a branch- ove — several racemose, turbi or campanulate, 10-26-flowered; involucre 11-15 m pi la af a puberulent, the phyl- nate laries subequal, few-ranked, the outer fcaaed and isn herbaceous the inner acuminate to attenuate, thin, dry; ac chenes appressed-pilose; pappus browni Rocky, mostly open slopes at a3 elevations, rime? Alpine Zone; a Oregon to oc Cou Califoraix, poe in the Sierra Nevada from Nevada County to: Tulare County, yr egy bei eae to Utah "Colorado, and Wyoming. Type locality: ‘“‘Banks of ous’ Re eal Cara ther streams of the Ore July- 21. Haplopappus suffruticésus (Nutt.) A. Gray. Big-head Macronema. Fig. 5466. Macronema suffruticosum Nutt. Trans. Amer. Phil. Soc. ors — 1840. Haplopappus suffruticosus A. Gray, Proc. ee Acad. 6: 865. as er suffruti kcoewe ws Kuntee, Rev. Gen. Pl. 1: dor Macbr. ah — 62 19 Haplopappus suffruticosus subsp. tenuis H. M. Hall, Na ae Wash. cde Ne 389: 190. 1928. Haplopappus suffruticosus var. tenu als McMinn, Ill. Man. Calif. Shrubs 573. 19 Low com aac subsh un up to 2 or even 4 dm. high with ‘ead ae nova -glandular, fragrant herbage, 1 very numerous on the brittle twigs, linear- oblanceolate to spatulate-oblong, 1-3 cm. lon mie mm. wide, entire, usually crisped and with peas fascicles ; eads 1-4 at aes Wye , mostly solitary, broa dly ca campanulate ; involucre with several outer foliaceous oblong bracts often ones n the 2-seriate. rh artaceous, lanceolate, acuminate, obscurely Cae al = aaa -_> lie sg mm. high, je Agena ony -florets s 3-6, showy ; disk-florets 18-40; villou what flattened; pappus mineou fe geen slopes and ridges, wee n one ne; agen wa Mount , Orego ast across northern Nevada and central cog to northwestern b Loe tiie and Montana and eerie by Cali seat Soy in the high Sierra Nevada and the White —_ ains. Type locality: banks of the Malade, a tributary of the Columbia, near the Blue Mountains. July—Sept. 22. Haplopappus greénei A. Gray. Greene’s Macronema. Fig. 5467. Haplopappus oS Gray, eige Amer. Acad. 16: 80. 1880. Haplopappus mollis A. Gray, loc. ¢ Haplopappus —. var. is ks aos a8 Fl. N. Amer. 12: 135. 1884. aoe greenet Kun’ ee . Gen. Pl. 1: 318. 1891. greenei Greene dese: 73. 1894. Mecronemas molle Greene, loc. c Hoorebekia greenei mollis “og Chati: U.S. Nat. Herb. os 561. 1906. Macro ema Se Nels. & Macbr. ge Gaz. 62: 150. 1916. molle Jepson, Man. FI. Pl. igi 925. Nisadabeus greenet iting mollis H. M. Hall, Carnegie Inst. oe Publ. No. 389: 195. 1928. va send ae 1-3 dm. high, glabrate to more or less densely neg ao aden to more or less s from punctate or sessile (very rarely stalked) g Lea very numerous, a le ag 1.5-3.5 cm. long, 3-7 mm. wide, plane or rarely fae Siched: heads clustered at ends of the twigs; a i 8-12 mm. high, subtended by a few leafy bracts; phyllaries seriate, subequal to somewhat imbricate, the o with her erie ligulate = three to four pues as long as the body, the inner with pro ne shorter caudate tips or acuminate or anes the body ceniienmly save cs sek wv itlces cea ay Pa 1-7, showy; disk. -florets 6-20. Roc Si flats and B sparsely 6 pe ion y slopes, Canadian and Hud an Zones; Cascade Mountains of Washington on the east slopes in Oregon, ree? a onbaton rel F ciotecasiers Oregon to Idaho; in California in —- as North Coast Maton genes eee County and also in Modoc County. Type locality: Scott Mou is, Siskiyou County, California. j cies 23. Haplopappus bloémeri A. Gray. Bloomer’s Macronema. Fig. 5468. Haplopappus bloomeri A. Gray, Proc. Amer. Acad. 6: 541. 1865. Haplopappus bloomeri var. angustatus A. Gray, op. cit. 7: 354, 1868. Ericameria erecta a fal asi Natiirf. Ges. Halle 15: 326. 1882. Haplopappus bloomeri et Greene, Pittonia 2: 17. 1889. Aster bloomeri Kuntze, “Rev. Ge Gen: Pit: = 1891. SUNFLOWER FAMILY 281 Chrysothamnus soaps 5 Greene, facie 3: 115. 1895. Chrysothamnus Greene, N. Amer. Fauna 16: 166. 1899, Macronema ese Nels, & Macbr. Bot. Gaz. 62: 148. 1916. 49 Ericamer ie Ooemar ri J. Fe Ma siheide. Contr. Gray Her « 56 Haplopapp angustatus H. M. Hall, ‘Carnexie pan oe Publ. No, 389: 198, 1928. Haplopappus proomers subsp. pledge: M. Hall, o 99: . pubescens Heniicrens, ibechee 322-27: 1936; i sage onto shrub sabia. mi tall with woody trunk up to 3 cm. thick, 1.5-4(-8) dm. high, r less to n glabrate metimes mentose, eglandular to pruinose-glandular or occasionally glutinous an sessile clave Apes umerous, nearly filiform to narrowly oblanceolate, mostly —6 mm. wide, plane or rarely twisted; heads in small terminal cl rs or com- monly more numerous in subrac sé spi or panicles, narrowly c nulate; involucre 7-12 mm. high; phyllaries 3—-6-seriate, clearly pecan Bap acer te to oblong, stramineous, the outer with caudate herbaceous ti IDs, at least ee er prominently scarious-margined, villous- ciliate; ray-florets mostly 1-5, wanting from s Seats. not very showy; disk-florets 4-13. andy or rocky soils, aa in iia i iN ge? Transition and Canadian pie add pegshesn British Columbia east of the Cascade Divide south through central Washington and Oregon to “ie uthern Sierra Nevada, California, and western Nevada. Type locality: Mount Davidson, Nevada. July-Oc 24. Haplopappus ophitidis (]. ee” Keck. Serpentine Macronema. Haplopappus bloomeri var. ophitidis - Howell, ae West. Bot. 6: 85. 1950. Haplopappus ophitidis Keck, Aliso 4: 1958. t-like undershrub up to , 20 em. high, the stout woody trunk (up to 12 mm. thick) soon Seitesecty 3 on compactly branched, black-barked, the herbage resinous from sessile yellow glands. ° a: bee ws 1 wy 8 SR rat gee: a et « SPs = .* ~ ps ph ey 5466. Haplopappus suffruticosus 5467. Haplopapp 5468. Haplopappus bloomeri 5469. Haplopappus ophitidis 282 COMPOSITAE eaves numerous on the very slender stems, narrowly linear, pores falcate-recurved, sulcate, wide; very sparsely arachnoid-ciliate, 5-15 mm. long, 0.5- ~ , eads disc sane terminating leafy twigs, solitary or in small cymes, cylindric at anthesis ; gn a 2-1 high; phyllaries 5-8-seriate, strongly imbricate, the outermost linear- nee olate, hataceone, male inner Sonate ide h oblong with wide hyaline margin, ety te to the base of the lanceolate to deltoid, herbaceous, Srickeneds often oe Sp, Sn nous, somewhat ciliate; florets mostly 5 ), light yellow. Serpentine soil, a esaage t of Mount Tedoc, northwestern Tehama County, California, the type locality and Pe Big per Teaoes ug. 25. Haplopappus gilmanii Blake. Gilman’s Macronema. Fig. 5470. Haplopappus gilmanii Blake, Proc. Biol. Soc. Wash. 52: 97. 1939. Low, rounded, intricately branched, aromatic shru b 2-3.5 dm. high, the old stems pallid like the new, the very slender twigs oe ed S en, adleate ‘ad furs Leaves numerous, often with tu 12 axillary fascicles, vernicose, spatu long, 2-3 mm. wide, Shai conduplicate and recurved at tip; heads solitary or eaiiae t as nch- ane patron campanulate; involucre 7- igh, resinous; phyllarie soa ate, clearly imbricate, the outer uate lakseolte often with thickened, Spenae ike. green, squarrose or flexed tip, the inner oblong, chartaceous, h with prominent, hyalin age Bit hte. Saye te appressed ; ray-florets 4-6, white; disk-florets 15-18, bee achenes silky-sericeous; pap ochroleu igh limestone ridges and daar ey Trade moat gd and ae Mountains, Inyo County, Cali- Fa i a, Seay ality: summit of Telescope Peak, Panamint Range. Aug.—Sep 26. Haplopappus resinésus (Nutt.) A. Gray. Columbia Macronema. Fig. 5471. Ericameria resinosa Nutt. Trans. Amer. Phil. Soc. IT. a 319. 1840. Haplopappus resinosus ray, Bot. Calif. 1: 313 ster resinosus Kuntze, Rev. Gen. Pl. 1: 31 891. Coyne ss aaa How ell, FL. N.W. pris 303. 1900. blopapp andoger, be Soc. Bot. Fr. 38. 1918. Haplopappus hamatus Gandoger, loc. c Renngeoeind ptewmpri Fe Gin aromatic shrub — dm. high, the old wood very black, new ery sle eae. resinous. Leaves numerous, often with axillary fasicles, Set eoun, resinous, filiform to — y linear, involute-thickened or conduplica mS recurv ee at tip, mostly less than 15 mm. long; heads solitary o r loosely cymose, nar rom in aig e 6-8 mm. high, glabrous: phylla ries +s. seriate, a aah da throughout or with s p, the outer often arrose; ray-florets 0-7, white or pale yellow (as also the base Pek tet 10-15, their rae obes 1-2 can long. Rocky plains and banks, mostly on basalt, Upper Sonoran and Arid Transition Zones; central Washington from the ni = Cascade Mountains _ northern and eastern Oregon and adjacent Idaho. Type locality: shelving rocks on the Blue Mountains of Oregon. July—Sept. 27. a nanus (Nutt.) D. C. Eaton. Rubber Weed. Fig. 5472. meria nana Nut Fe 0 Amer. Phil. Soc. IT. 7: 319. 1840. Erica Haplopappus nanus C. Eaton, Bot. King Expl. 159. 1871. Chrysoma nana Greene, Brythea 3 3: 10. 1895. size of H. resinosus. mie very aoe linear-spatulate to linear or involu and linear-filifo , subu ia fell mucronate, obscurely if at all punctate but resinous, often with bud-like axillary fascicles; heads in small eafy cymes, yellow; involucre turbinat 4 jiate what herbaceous, the tips obtuse to sharply acute or even aristate- caudate, not squ ray- ie (0-)2-7; disk-florets 5-10, their lobes about 0.5 mm. long; pappus stramineous, ge a as bx | goes rocky plains, cliffs, and crevices, Sonoran Zones; rare in Mono and Inyo Counties, California; common chron the Gre Great ig tat hs Utah and the Snake Plains of Idaho. Type locality: “Blue Mountains of Oregon, ” but ore 28. Haplopappus hallii A. Gray. Hesperodoria. Fig. 5473. Henlow: hallti A. Gray, Proc. — Acad. ot 389. 1872. i Kuntze, Rev. Gen. Pl. 1: 316. 189 Henhsvodoria belltt Glade, Lantictd Wik Oe. i: in 1906. Hoorebekia hallii Piper, Doute: U.S. Nat. Herb. 11: 560. 1906. Suffrutescent, 3-6 dm. high, the woody base branched, the virgate stems usually simple below the inflorescence, smooth or hirtellous. Leaves oblanceolate, sessile or the lowest petiolate, io mm. wi i wha’ fe a Dry, grassy or openly wooded slopes, Transition Zone; overlooking the pi Rives Gorge, Washington and Oregon, south to Calapooia Mountains, Oregon. Type locali ity: blufis at The Dalles, Oregon. Aug.—Oct. SUNFLOWER FAMILY 283 5470. Haplopappus gilmanii 5472. Haplopappus nanus 5471. Haplopappus resinosus 5473. Haplopappus hallii 29. Haplopappus linearifdlius DC. Stenotopsis. Fig. 5474. Haplopappus linearifolius DC. Prod. 5: 347. 1836. Stenotus linearifolins Torr. & Gray, Fl. N. Amer. 2: 238. 1842. ifolius Ku ster linearifoli ntze, gp te en. Pl. 1: 318. 1891. Haplopappus interior Coville, Biol. os pes 7: 65. 1892. Stenotus interior Greene, sikae * 72 Haplopappus linearifolius var. interior M. pa nes, Proc. ae Acad. IT. 5: 697. 1895. 19 Stenotopsis linearifolia Rydb. Bull. Torrey Club 272-617. Stenotopsis interior Rydb. loc. cit. Stenotus linearifolius var. interior H. M. Hall, Univ. Calif. Pub. Bot 3: 49, 1907. Toe: linearifolia var. interior J. F. Macbride, Contr. Gray Herb. No. 49: 59. 1917 Haplop s linearifolius subsp. interior H. M. Hall, Carnegie Inst. Wash. Publ. No. 389: 158. 1928. Mu ee branched shrub 4-15 dm - high, essenti cated glabrous but usually puberulent at summit of peduncles, the twigs fastigiate, very leafy, resino s, glandular-punctate. Leaves nearly linear, narrowed toward base, cm. long, ‘2. mm. ae entire, at or becoming subterete, some- times fasciculate; heads numerous, solitary on nearly naked peduncles; involucre ri high, the phyllaries 2-3-seriate, scarcely graduate, lance-oblong linear, acum . re, beset with granular glands, with greenish center and —_— e-ciliate, scarious oie Ga hares 13-18, ligules 8-15 mm. long; achenes silky-pilose; pappus white, deciduo Roe or sandy soils of mountainsides and deserts, Arid ae i Sonoran pala i AR Buttes ate “County, California, south _— ugh the Inner So _ ie —— to —— cismontane southern Cali- fernie nee ees r California east across ie Mojave Desert o County, California, southern Utah, a est- ern eteg Ty vie locality: Caifornia pete -~-May pala modest variation in head size and Teaf — has led to persistent attempts to recognize a desert var. futevine. The reduction in organ size in more arid a , however, seems to be merely environmental modi- fication rather than menahicnliv eenned eat ation. 284 COMPOSITAE 30. Haplopappus yen (H.B.K.) Blake subsp. vernonioides (Nutt.) H. M. all. Coastal Isocoma. Fig. 5475. it Hook. & A a Ragone? 351. 1838. isan vernoniotdes Nutt. Trans. hes il. Soc. II. 7: 320. 1840. sie are appus menziesit Torr. & Gray, FL. . Amer. 2: 242. 1842. . Gray, Proc. Amer. Acad. 8: 638. Vas Tevemme veneta var. vernonioides Jepson, Fl. W. Mid. Calif. . 1901. aad ocoma pclae nie ia G eaflets Bot. Obs. 1: 171. ene, loc. mai Isocoma villosa Greene, op. cit. 172 Haplopappus venetus maniees We cuoninides 11. M. Hall, Carnegie Inst. Wash. Publ. No. 389: 224, 1928. Shrub 4-12 dm. high with erect, ascending or decumbent s from a branched suffrutescent base, usually simple is He the inflorescence, somewhat ier Soa nearly Ter heone to pilose or tomentose, very leafy. Leaves aeae _ oblanceolate or spatulate-oblong, I 2-8 m. wide, spinulose- derate throughout or only near apex, sometimes even lobed or essentially entire, usually with axillary fascicles; niin discoid, turbinate, 15 fl Pe seominet usually compact cymes; involucre 5-7 mm. high, strongly graduate, t phyllaries ong, obtuse or acute, firm, pale, with short, usually granulous-greenish, appressed cate achenes oie pappus brownish. Coastal valleys, in sandy, often subsaline places, Upper Sonoran Zone; San Francisco to San Diego, Cali- fornia, and islands. Type locality: marshes near the sea, Santa Barbara. Rost —Dec Exe god variable in habit, pubescence, shape and cut of leaves, FP “involucre, Rh H. venetus grows in cen eieal Mexico; all other forms of the species ae in California and L r California. The more noteworthy Haplopa venetus subsp. oxyphyllus (Greene) H. M. Hall, Peseta Inst. Wash. Publ. No. 389: 225. 1928. (lsocoma oxyphylla Greene, Leaflets Bot. Obs. 1: 171. 1906; Haplopappus venetus var. oxyphy — Munz, Man. S. Calif. 523. 1935.) Robu “ eek 1~2 m. high, loosely villous to glabrate; leave oblanceolate to narrowly spatulate, acute or acuminate, ent 3-5 c m. long; hse openly paniculate. els Be bo bh ag pre Boog ties in outhern =e Diego County and ai J (ca Hidenia. Type locality: Jamul Valley b of San pn Haplopappus venetus subsp. furfuraceus (Greene) po - Hall, Carnegie “ae Wash. wo ik 389: 226. ei » (Bigelovia B bad elke Greene, Bull. Calif. a p Es 883i Isocoma decumbens epaerse Leaflets Bot. 2: vencta var. decumbens Jep Man. rH Calif. 1029. 1925; Haplopappus plage i var. einbeus. cet Man 3) Calif. B22, 19353 H. Sa ueiat Var. he rfuraceus Munz, op. cit. 523.) Stems slender, decumbent or curved, 3-5 aa One; leyye. crowded, narrow, small, few- ley 1s or entire, with eaiinent fascicles in the axils, mostly glabro metim: oolly; heads loos: ely c cymose, not in large glomerules. Dry sandy vt southern San Diego Couts: California, i praed California, and the poe lle panda. Type locality: not kno aplopappus venetus var. sedoides (Greene) unz, Man. S: Calif. - 1935. (Bigelovia veneta var. sedoides Greene, Bull. Calif. ame 2: 400. 1887; Isocoma sedoides Greene, Leaflets Bot. Obs. 1:172. 1906; I. latifolia Greene, loc. cit.; J. eta var. sedoides Jepson, Man. FI. Pl. Calif, $029. 1925.) Prostrate, almost glabrous, stout; leaves poe pie fe erate: de sspmai heads large, glomerate. Santa Cruz and Santa Rosa Islands, Calif Type locality: Santa Cru Haplopappus venetus var. argutus eke} e905 Aliso 4: 103. 1958. (Jsocoma arguta Greene, Man. Ba “aS aa 1894; I. veneta var. arguta Jepson, FI. W. Mid. Calif. 500. 4701. ) Low bush with erect stems 4 dm. high; ‘leaves pinnately cleft into acute lobes or only saliently but pungently denate. North and south _ Carquines Straits, Solano and Contra Costa Counties, California. Type focally: subsaline plains east of the Vac ounta 31. Haplopappus acradénius (Greene) Blake. Desert Isocoma. Fig. 5476. Bigelovia acradenia Greene, Bull. Torrey Club 10: 126. 1883. Aster seen Kuntze, Rev. Gen. Pl. 1: pe _ Tsocoma acradenia Greene, aothogaet 241%. te eae 5 iaiciesesee acraden aH. M. Hall, on ‘Calif. P ub. Bot. 3: 64. 1907. Blake, Contr. U.S. Nat. Herb. 25: 546. 1925. {Binion to H. venetus, low shrub with numerous erect, more woody, aes as barked, striate, glabrous stems 3-10 dm. high. pag linear- spatulate to oblong, 1~4 cm. long, 1-5 mm. wide, thick, entire, mostly mucronate and glabrous, minutely impressed-punctate, riety in teen 2 fascicles; heads in smaller cymes, i sessile, 6-13-flowered; involucre 5-6.5 m igh; oe mi pie wcnalaais = subepi dermal resin- eee near ie rounded or Bere tip, the ry narrow argin fimbrillate, amathitiacen or ails is -ower Sonoran Zone; southwestern Mojave Desert (common in Antelope easier) and Mojave Desert Avs N to — Valley, California; rare in southern Nevada and western Arizona. Type lo ity: ve Haplopappus acradenius subsp. ereméphilus (Greene) H. M. Hall, Carnegie Inst. Wash. Pu = = 389: 233. 1928." CFrocoma eremophila. reene, — Bot.) Obs, 1 171;: 3906; + pee tiaenacingd aevadeni ar. ere- ~ ophilus Munz, Cahf.. 523. vs Leaves denticulate to dentate or even saliently ve efi some ntire, 2-5 cm. Fi any as a mm, wide, scabrid-hirtellous: heads 15—25-flowered; involucre 6-8 mm. high. Sortie owienar Mojave Desert and a ee the Colorado Desert, California, extendin g slightly into Arizona and Lower California. Type locality: southwestern part of the Colsenio Desert Haplopappus acradenius subsp. Shaler Bana 20 (Greene) H. M. Hall, Carnesic Inst. Wash. Publ. No. 389: 233. 928. vibe bracteosa Greene eager Bot. Obs. 1: 170. rane Haplopappus acradeninus var. bracteosis McMinn, Ill. Man. Calif. are "574. 939.) Leaves mostly denticulate, the lowermost 2-3 cm. long but the great majority only 6-10(-15) lon gtr or reflexed; heads A aa eenwenegs involucre 6-9 mm. high. Both s f the San Joaquin Viney, California. Type locality: Tulare Cou oe: epmantaacaae: canus (A. Gray) Blake. Island Hazardia. Fig. 5477. Diplostephium can A. Gray, Proc. Amer. Acad. 11: 75. 1876. Corethrogyne de ik Greene, Ahi Torrey Club 10: 41. 1883. Cor ethrog. ogyn g cane Gp eene, Bull. Calif. pais 22s. a ie onia 1; 29. Hazardia detonsa Gredae: ‘ae cit. SUNFLOWER FAMILY 285 Hazardia s a Greene, op. Haplopa Pee canits Blake, Cont tr. a S. Nat. Herb. 24: 86. 1922. si raskae Eastw c. Calif. Acad. IV. py 156, 1931; Openly branched ae 6-25 dm. high, sini i tely to densely lanate- eo isohce coree the atone onal eet a above. Leaves obovat oblanceolate, jcinagi pe rea sic ov cm. long wide, entire ‘to sharply ai thick; heads numerous, cymes, pe depiciilate or sessile; involucre broadly turbinate, 10- 13 m m. high, the eipiiacicn “nul seriate, strongly dp jens appressed, oblong, acute, tomentose, or the i nner tomentose only a or gla brous; ray-flor 6-14, iconspicuous, not exceeding disk, like the 20~54 disk- rphind turning from yellow = ath ish vation nes nerved, pilose; pappus tawny or brown. Dry rocky slopes, Upper peer Zone; Santa Rosa, Santa (4 and S an te Islands, California; Guiadahape "Este nd, Lower Califor Type locality: Gaadslene. Island. June 33. Haplopappus squarrésus Hook. & Arn. Common Hazardia. Fig. 5478. Haplopappus ds ng k. & Arn. Bot. vee 146. 1833. Hasardia squ a Greene, Erythea 2: 112. 18 Mbattleesbera shrub, woody at ‘ee i 10 dm. high, glabrate to ered phage or, iti pat above, pilose. Leaves many, oblong to cuneate-obovate, obtuse, clas t base, 1.5-4 cm. long, 1-2 cm. wide, sharply serrate throughout with m ul Feat re o ctate a ronulate teeth, es rom can glands, glabrous or pubescent on midrib beneath; heads discoid, 15-30-flowered, spicate or race- mosely er involucre turbinate, 11-15 mm. high; phy llaries 8-10- seriate, strongly graduate, 5474. Haplopappus linearifolius 5476. Haplopappus acradenius 5475. Haplopappus venetus us canus 5477. Haplopappus 286 COMPOSITAE prominently glandular-scurfy at the green, rg to acute, usually squarrose tips; achenes glabrous or sparsely pilose; pappus yellow-taw Coastal bluffs and montane canyons and ridges, often in chapar Upper Sonoran Zone; Monterey County to oe — County, California. Type locality: probably near "Maiteres Aug.—Oct. seg ppus —— rrosus subs wh phone pag (DC.) eyo — 4: aye 1958. (Pyrrocoma grindelioides pe. t S: 350; 1836; as _orindetioi ntze, Rev 316. 1891. } ag Bn hairy, the stems arsed tomentulose 1 — the heads, rik tg seine ie a He yon often somewhat heads sma iller ; involucre oe eS . high; hyllaries i cinereous on both faces oF the ee hin neat glandular only marginally i Pe an pappus red-brown. Cismontane southern California from Santa Barbara to San Diego and in Lower Californie. Type locality: “‘California. ot July—Oct. Haplopappus squarrosus subsp. obtisus (Greene) H. M. Hall, Regge once Inst. Wash. Publ. No. 389; 253. 1928. (Hazardia obtusa Greene, Fl. Fran. 375. 1897; H. squarrosa var. obtusa pres, Man. FI. Pl. Calif, 1030. 1925; Haplopappus squarrosus var. obtusus McMinn, Ill. Man. Calif. Shrubs 571 9.) Heads relatively large, 18-25- flowered; involucre ssi ge rate oc 15 mm. high; sible ied” broad, very blunt, mucro- nate, the resinous-granular, pallid, glabrous tips appressed. Can in the mountains west of Tejon oy Sept No Socnty. south iecomat Ventare County to Nordhoff Peak. Cali ternias 5 pay locality: San Emigdio Cany ept plopappus squarrosus subsp. stenolépis Hall, Carnegie Inst. Wash. Publ. No. 389: 253. 1928. (Hoplopappus squarrosus scr stenolepis Bo inn, hs Man. Calif. Shrubs 571. 1939.) Usually eerie 4 base, very branched, forming dense shrubs to 1 a ema nd 2 m. oss, = twigs scabrid or hirsutulous rela- tively small; heads 4-8- flowered, densely Bevin e; Pease ucre on narrowly turbinate, 13-17 mm. high; gern Sab 6—7-seriate, regularly but loosely Saicicane’ eh acuminate, at length spreading but not a eae ose, sparingly = with — glands on pe small herbaceous portion near tip, somewhat viscid but glabro bus! pappus red- own. Serpentine areas in Inner South Coast Range, Fresno and sseperetey Counties, California to Cuyama River, Santa Barbara County. Type locality: Parkfield Grade. Sept.— 34. Haplopappus pinifélius A. Gray. Pine-bush. Fig. 5479. See ee Sa iseselan Gray, Proc. Ne popes 8: 636. 1873. Aster pityp s Kuntze, Rev. Gen _ Pe 1891. Chrysoma i stale Greene, Erythea 3 Ericameria pinifolia H. M. Hall, Ciin Pu tk Bot. 3 54. 1907. Haplopappus illinitus Eastw. Proc. Calif. Acad. ITV. 20: 155. 1931. rub 6-25 dm. high, the main stems ecaks like, fastigiately branched, resinous, glandu- i -3. lar , sometimes slightly pilose. Leaves linear-filiform, g, mu €, - terete, with shorter leaves fascicled in the axils; hea litary in the spring form, large, terminat i igs, ofte sed by subtending leaves, in the a aller, in short racemes or ic mose-clustered at tip branches; involucre of the turbinate, mm. high, overlapped by leafy bracts; phyllaries loosely -seriate, lance ae rent oblong, the outer often more or less ca ate-tipped, oe inner shorter-tipped or tips green, otherwise pale with scarious ciliate marg e costa sometimes dL iagetglat. thickened above; ray-florets 5-10 (15-30 in vernal heads) ; ak “docks 12-18; achenes sparsely pilose sie buff or reddish. Cis e dry slopes and washes, Upper Sonoran Zone; — southern California from northern Los Angeles County to shathern San Diego County. Type locality: r Los Angeles. April-July; Sept.—Jan. 35. Haplopappus ericoides (Less.) Hook. & Arn. Mock Heather. Fig. 5480. Diplopappus ericoides Less. Linnaea 6: 117. 1831. Haplopappus ericot ides Hook. & Arn. Bot. Raging! 146. 1833. Ericameria microphylla Nutt. Trans. Am . Soc. II. 7: 319. 1840. tag ericina Runtse, Rev. Gen. Pl. 1: an 91 rysoma Greene, Erythea 3: 11. Ericameria ericoides Jepson, Fl. W. Mid. ies sab: 1901. d compact shrub with dense crown, 3-8(-15) dm. high, the a ore apeosely rare uieetinr with the — sparsely pilosulose and somewhat resinous. Leaves i Hesen’ te) i ch e, ea : m. ait laries loosely LSE eek villous- ciliate, the eater ote ate-lanceolate with short-caudate, greenish tip, the inner broadly oblong, acute, not herbace ise bass costa thickted : hive into a S eiiform gland; ray-florets 2-6; disk- ye 8-14; achenes glabro Sand d on and — Bhs Upper Sonoran aie Point Reyes, Marin omen to Los Angeles County, Cotiforuie; reported f i 5 ik Island, Type localiey' Cab tone. Aug aplopappus a subsp. blakei C. B. Occ. Papers Rancho Santa re Le wae id. 3: 87: ah Achenes moderately sericeous. og hills away fr oie i immediate coast, Be 2 1,500 Led altitude “Uppe noran and Transitio: o nee oat ruz Mountains, Santa Cruz and Monterey Counties, Santa Maria Y hives Valley, San Luis Obispo and ooree Barbara Counties, and near Saute Wachee: ‘California. Tyee locality: near a oo School, Sant Cruz County. Sept.-Nov. The leaves are usually but not always longer in the southern material 36. Haplopappus eastwoddiae H. M. Hall. Eastwood’s Ericameria. Fig. 5481. Chrysoma fasciculata Eastw. Bull. Torrey hee wa c2is: a itera tc fercwatey. HS Macbride Contr. Gray Herb. No. 56: 36. 1918. = Hall, Poet lad Wa om Publ. No. fa 258. 1928, " Rtowk dense, ek branched shrub 5-10 dm. -— glabrous, very resinous, a punctate, densely leafy. Leaves linear, 8-20 mm. long, e€ or —~ what flatten cee cronate, nearly all with axillary fascicles of smaller lensel: heads so solitary o Tally several in Fcereaitial cymes of close raceme ulat 78m high; shotiasins about 5-seriate, strongly graduate, scarious aad ‘pale yellow, t the outer "Grade lankoolene: sharply acute, the inner oblong, SUNFLOWER FAMILY 287 obtusish, none sae at ey tong wy on the narrow hyaline margin, the costa somewhat glandular- thickened abov ar -florets 1-6; disk-florets 18-22; achenes densely ailky-D ilose coastal s. wa we S, mt oran ie enite y and Carmel] Bays, California. Type hese near ‘oan J Se ad 37. Haplopappus palmeri A. Gray. Palmer’s Ericameria. Fig. 5482. Ha plopappus ut Gray, Proc. ep _ 11: 74, 1876. Aster nevinii Kuntze, Rev. Gen. Pl. Be Chrysoma palmeri qed Erythe n§ ty Ericameria palmeri H. M. Hall, “sab Calif. Pub. Bot. 3: 53. 1907. Stout shrub 1-4 m. hi 1 glandular-punctate, very lea Mak Leaves filiform, 1.5—-4 aif long, subterete, faucieuts te heads many, in thyrsoid panicles; involucre “ta binsite, 5-6.5 mm. high; phyllaries 30-40, loo ely 4-5- seriate, linear or a so, blunt, glabrous or the outer glandular: atomiferous, ciliolate at tip, the hyaline margin n the costa thickened for most of its a into a linear-oblong gland; ray-florets 4-10; disk-f florets 8~20; achenes moderately sericeo Dry mesas, Sonoran Zones; sites Ele: San Diego b> iad Ss hae to northwestern Lower California. Type elit Tecate cae Lower California. Sept._Nov ppus palm ss pas hylépis H. M. Bas il, Ca arnegie Inst. Was By bl. No. 389: 267. eaploteene palmert wae * picket s Munz, Man. S. Calif. 522. 1935. Sianter "5-15 dm. hi gh; leaves mostly 8-12 mm. long; involucre FE ai ce ake 6-7 mm, hake phyllaries anes prea e “thicker, “tSeriate, soared lanceolate to —— ne, ~~ bullate costal gland in apical half only; ray-florets 1-4; disk-florets 5-10; achenes densely seric Rat Hite on clayey plains, Upper Sonoran Zone; densbatine California trea southeastern eee paseo ing fo aoe desert borders, Diverside County; Santa Catalina Island. Type locality: summit of Box Springs Grade near Riverside. Dec 5478. Haplopappus squarrosus 5480. Haplopapp icoid 5479. Haplopappus pinifolius 5481. Haplopappus eastwoodiae 288 COMPOSITAE 38. Haplopappus propinquus Blake. Boundary Ericameria. Fig. 5483. Bigelovia brachylepis A. Gray, Bot. ot 1: 614. 1876. Aster brachylepis Kuntze, Rev. Gen 317. 1891, Chrysoma brachylepis Greene, batt ne : « 1895, Ericameria brachylepis H. M. Hall, Univ. Calif. me Bot. Haplopappus propinquus Blake, Contr. U.S. Nat. (kN 23: 1490. 1926. Shrub 1-2 m. high, saees epee Sara ey more or less resinous, glandular- prea Leaves crowded, linear- -filifor ,12¢ .5-1 mm. wide, flattish or subterete, often nate, sharply ascending, wit ciliary faces heads disenid 9-14-flowered, yellow, ernie or racemose; involucre turbinate, 4.5-5.5 mm. high; phyllaries 3—4-seriate, strongly graduate, ovate to linear-oblong, the outer grading into "ihe scaly eos of the pecnacies, the oo promi- nasithy glandular-thickened throughout its Sige achenes densely villou Desert slopes, Upper Sonoran Zon n San Diego Cou nty, oe ornia, and ca Lower California. Type locality: ‘‘Larkens’ Station, 80 niles ~ nee iaeth ot San Diego.”’ —Dec 39, Heplepepas laricifolius A. Gray. Turpentine-bush. Fig. 5484. H icifolius A. Gray, Smiths. Contr. 5°: 80. 1853. Aster rarflin Kuntze, Rev. Gen. Pl. 1: 318. 1891. Greene, Erythea 3: 11. 1895. a cis Saricitola Sinners, Field & Lab. 18: 27. 1950. Compact, fastigiately pags ee broadly rounded shrub t “yo Heh: resinous, prominently sieeiresied. pin ctate, glabro Leaves linear, 2 cm. lon wide, usually subterete, onate, ometimes with ek maller ones in axillary (ie ; heads in ‘naa eal cymes ; a -acu oO a ; ph and ciliolate at tip, rather firm, the costa thickened for most of its length into an olive-brown gland; ray-florets 3-11; disk- florets 10-16, much exceeding the in Scere. Rae densely pilose. Rocky desert oe and mesas, Sonoran Zon pein eastern ears Desert from Clark Mountain to H coor Mountain; east to western Teas aad ier ent Mexico. Type locality: Guadalupe Pass, New Mexico. ept 3 40. Haplopappus codperi (A. Gray) H. M. Hall. Goldenbush. Fig. 5485. Bigelovia cooperi A. Reed dia Amer. Acad. 8: 640. 1873. Gray, op. cit. 19: 1. 1883 dies baker’ Kuntze, Rex Gen. BU 333337..1891. Aster monactis Kuntze, o t. 318, Ericameria monactis MeChatshie, 2 egg 2: 124. 1894. 1895. Chrysoma cooperi Greene, op. c Acamptopappus nirocephaas ao Ao ones, — ng Bot. No. 7: 30. 1898. Chrysothamnus corymbosus Elmer, Bot. Gaz 905. Tumionella monactis . ne, peed Bot. Obs. 4 ee a priscomisoae oopert vi H. M. ere ble Ca lif Pub. B Low seed. Scns branc aa coe 2 cay dm hi gh, very woody, the bark oe peo the herbage glutinous, loosely pilosulose. Leaves oe 6-15 mm long, up to 1.5 mm. wide, wi th Nain pwctate, diminutive fascicles in the lower axils that persist after the fall of the principal ie seas prominently peduncled in small, rounded, terminal cymes; involucre narrowly campanulate, 4-5 mm. ig i aaarics 2-3-seriate, few (9-15), the outer ovate, acute, the inner i roa en ‘oblong, obtu or less puberulent, the ning of the costa ae oak ; ray-florets 0-2; disk- fost ',7(-11), much exceeding teh achenes silky-pilos mmon in rocky desert basins ne mesas, Lower Sonoran Zone; California across ~ Mojave Desert from . County to an roa! Valley and Little San Bernardino Mountains; rare in interior ee shes ornia; Pasa ste t Nevada. Type locality: eastern slope of Providence te cantains. southeastern Cali- ornia are ~ Bie igh arboréscens (A. Gray) H. M. Hall. Golden Fleece. Fig. 5486. b A. Gray in Torr. Bot. Mex. Bound. 79. 1859. b A. Gray, Proc. Amer. Acad. 8: 640. 1873. tis gece US Kuntze, Rev. Gen. Pl. 1: 315. 1891 Ericameria arborescens Greene, Man. Bay ah 175. 1894, Chrysoma arborescens Greene, Erythea 3: 95. Haplopappus arborescens H. M. Hall, pels odes Pub. Bot, 7: 273. 1919. Stout erect shrub 0.6-3 m. high, fastigiately a glabrous, resinous, prominently glandular-punctate. Leaves narrowly linear to filiform, 3-6 cm. long, up to 2 mm. w wide, thick, unded ter cymes or cymose egg peduncles mm, yrs bearing scales like the pesestre involucre turbinate, 4-5 mm. high; phyllaries 4-seriate, graduate, lanceolate to linear, acum to acute, thin and chaffy except for ee peadiles Gockre! costa; achenes turgid, Sates 5-angled, less than 2 mm. long, softly ous; p ry fragile. Dry foothill slopes, in chaparral, mostly eekly 4 000 Pi He up to 9 AB eet in Fresno County, Upper Sonoran — anit braesse § Aga reds nae at ed pid Nevada — a xa County to Wiinre County Outer Coast Ranges Oregon li o Vent , California. Type : “California.”” Aug.—Nov SUNFLOWER FAMILY - RO 5482. Haplopappus palmeri . Haplopapp peri 5483, Haplopappus propinquus $486. Haplopappus arborescens 5484. Haplopappus laricifolius 5487. Haplopappus parishii 289 290 COMPOSITAE 42. Haplopappus parishii (Greene) Blake. Parish’s Ericameria. Fig. 5487. Bigelovia parishit Greene, Bull. Linch gg Club 9: 62. 1882. Aster parishii Kuntze, Rev. Gen. Pl. 1: 318. 1891. Chrysoma parishii Greene, Kiyibes 3: as 1895. Ericameria parishii H. M. Hall, Univ. Calif. Pub. Bot. 3: 55. 1907. Haplopappus parishii Blake, Contr. U.S. Nat. Herb. 23: qe 1926. Erect aoe 2-5 m. high, glabrous, resinous, densely glandular-punctate, the stems trunk-like t base, very leafy above. Leaves linear-o blanceo late to lance- se ptic, tapering to base, acute use, 2-6 cm. 1 2-fl or , 2-6 cm. long, 3-10 mm. wide, flat, coriaceous; heads , 9-12- ered, in com- pact rounded cymes, the short pedun cles bracteate with small ‘scales Fovokigte turbinate, abo mm. high ; phy aries 4-seriate, lanceolate | to lance-oblong, a to ac uminate, whitish, _ fra Outwash fans and dry hillsides in chaparral, not common, Upper Sonoran Zone; south slopes of San Gabriel _ Beco ve, pera bag teers = California, eat to Lower California. Type loc ality: Waterman Canyon, San ernardino Mou 43. Haplopappus cuneatus A. Gray. Cuneate-leaved Ericameria. Fig. 5488. eres oe A. Gray, Proc. Amer. Acad. 8: 635. 1873. Bigelovia spathulat sa ig op. cit. 11: 74. 1876. Aster cuneatus Kentes , Rev. Gen. Pl. 1: 317. 1891. Bricomeria eaten oty MeCiatee, Erythea 2: 124. 1894. Chrysoma Gre bead pence ae snide sthalite Priest ie oc, Chrysoma merriami Eastw. Bull. Torrey Clab bed 215: Ericameria cuneata (var.) ébatlodaia £L. M. Hall, Univ. ati Pub. Bot. 3: 52. 1907. Haplopappus cuneatus spathulatus Blake, ie U. S. Nat. Herb. 23: 1489. Low eggs ob epiatt -5(-12) dm. high, much branched, glabrous, balsamic-resinous, glan du- lar-punctate. Leaves crowded, deep green, cuneate to suborbicular- obovate, entire, often undulate, so at the obtuse or broadly rounded or often retuse apex, -< g, 3-1 0 mm. wide, coriace eads actly c pane: involucre turbinate, 5-7 mm. hi gh; siesthisies igakciase, regularly ‘imbricate, Tihear- oblong to lance-ovate, the costa gla ndular-thickened above, the outer passing into minute, ovate, thick scales of the p ily ray-flor 1-5 (in northern Sierra evada) o} Yate wanting; disk-florets 16-28; rae henes densely pt edey -pilose. iffs and rocky slopes, Upper Sonoran and Arid Transition Zones; both slopes of the Sierra Nevada from Plumas County to Tular : oe ty and ranges toil the Mojave ona Colorado Deserts from southern Rec nog County to Ventura and ‘Seg Diego Counties, and eastern San Luis Obis —s County: to the San cone ky Mou ne to Nevada, Arizona, and Lower California. Type locality: Annual herb from a taproot. Stem erect, cymose-paniculately branching above, the heads terminating the branchiets. Herbage heavy-se scented from a harsh glandular pubes- cence. Leaves essentially entire, prominently reticulate-veiny. Flowers yellow, tinged wit red. Involucre cylindroturbinate, 5—6-seriate, the phyllaries 35-50, corneous, linear-attenu- ate, glandular-atomiferous, the spreading or recurved tips of the outer ones bearing a prominent gland. Ray-florets fertile, 5-14, the irate Oe ng Mendy circinate, the achene 3-angled: disk-florets sterile, 9-25, the corolla constricted at base of throat, its style- abst ecualing the achenes. he: for San Benito County, California, where it occurs. A monotypic genus of California. 1. Benitoa occidentalis (H. M. Hall) Keck. Benitoa. Fig. 5489. ol Inst. Publ. No. 389: 214. 1928. Dentist seribanselie Keck, iis’ Wik Bot Pa guy a lan —_ a. rast rlandular pubescence, the leaves often additionally ahewhat fear hirsute, sien on anthocyan bas wien mature. Leaves linear-lanceolate, obtuse or acute, narrow wed to a + clasp base, ane cm. oak. 6-15 sats wide, becoming bract-like and apiculate in upper half of eels ee olucre 8-10 mm. high, 3-5 mm. wide ; ray-achenes olive, brown-maculate, finely sericeous, Hot, Fay. exposed, serpentine eae Upper Sonoran Zone; Diablo Range in San Bena, Monterey, and , California. Type locality: east side of Parkfield Grade, Fresno County. June—Nov * Text contributed by David Daniels Keck. SUNFLOWER FAMILY 291 77. SOLIDAGO* L. Sp. Pl. 878. 1753. Perennial herbs with leafy, usually simple stems arising from rhizomes or a ca he Leaves alternate, entire or toot eads radiate, yellow (in ours), small, campandlat to subcylindric, panicled, racem ose, or cymose. Involucre few-seriate, graduate sub- equal, the phyllaries usually with obscurely herbaceous tips. Receptacle usually iveclnte, ay-florets small, fertile. Disk-florets perfect, fertile, their anthers subentire at base, their style-branches with mostly lanceolate appendages. Achenes short, pubescent (in ours), usually few-nerved. Pappus setose, nes Rae whitish. [Name Latin, meaning to make whole, from its reputed medicinal value. | A genus of about 100 species, chiefly Kauees American, a few also in ome America and Eurasia; the genus is best Necchog ed in the eastern United States. Type species, Solidago virgaurea L. Plants — -~— ny Padi, a rhizomes; stems rather equably leafy, the lowest leaves not prominently differ m the r cauline and at length deciduous (except in S. missouriensis). ‘ase pen, rahe lance- ees Sulleeeedee ample, copiously bracteate, the heads in terminal cymose clusters; ray-florets 15-25. 1. S$. occidentalis. Leaves ee ee inflorescence more compact, not interrupted, the heads not glomerate in cymes; Stems puberulent at least above the middle. Leaves densely puberulent on both faces, the middle and upper usually ante, and entire. . S. californica. Leaves puberulous chiefly on nerves beneath or chiefly glabrous, the middle aa upper lanceolate, usually sharply toothed. Involucre evidently imbricate; inflorescence a compact thyrse to a broad a ee oem 5- ae: ame long. Involucre obscurely imbricate; inflorescence smaller, short and compact; ay” —_ “hen Stems seas below the inflorescen a Ste Nias ane at nd n ge oe “uniformly leafy throughout, glacuous, 5-20 dm. “et Spa al — lan- Stems with vali gt aga etaec. not glaucous, 2-5 dm. high; upper leaves Be linear ve allie, ourtensts. rhizomes or a branched oust stems with lower and basal Dae poe much Plants with — shor Secas larger = upper reduced sessile ones; inflorescen tgs ut recurving branches. Plants - "i mid- elevations, usually more =— 4 dm. hig In = aot “tag nous; involucr: 5-5 mm. oe leaves entire (rarely remotely serrulate in ei ti Panicle subracemose, few-headed; st 1 Californi 7. S. guiradonis. Pinicic usually oblon aind — dense; stems rather mont Phyllaries acute or acuminate; rays mostly 8 Californi 8. S. confinis. Phyllaries obtusish; coal mostly et Sigs Haske: 9. S. spectabilis. Inflorescence glutinous; involucre 5-6 mm te; coastal 10. S. spathulata. Plants of high altitudes, mostly less than 3 dm. high. pulttces us; phyllaries obtuse. Leav 11. S, decumbens. Laci ciliate at least toward petiole; phyllaries acute to acuminate. 12. S. multiradiata, * Text tributed by David Daniels Keck. 292 COMPOSITAE 1. Solidago occidentalis (Nutt.) Torr. & a Western Goldenrod. Fig. 5490. Euthamia occidentalis Nutt. Trans. Amer. Phil. Soc. II. 7: 326. 1840 Solidago occidentalis Torr. & Gray, Fl. N. Amer. 2: 226. 1842 Euthamia linearifolia Gandoger, ape best Bot. Fr. 65: 41. 1918. Euthamia age ey Gandoger, loc tout, from creeping eri che much branched, 6-20 dm. high, glabrous throughout, often withinons a s lance-linear ssile, , 3-5-n 4-10 long, 3- wide, the margin often scabrid, glandular-punctate; inflorescence ample, leafy-bracteate, inter- rupted-elongate or rounded, eads in 1 cymose clusters; in re 4 i 1- es firm, lance- ae to lance-linear, acute; ray-florets 15-25, 1 .5-2.5 mm. long ; disk-florets 14; achenes pilos Mo ist ground at oa altitudes, Bere ates Shot and Transition Zones; British Columbia - ee eee east to Alberta, N Linton and Texas. Type locality: ‘‘Banks of the Ore egon and Wahlamet, and L in the Rocky Mountains.” July—Nov Solidago ae emer’ F var, major (Michx.) Fernald, Rhodora 46: 330. 1944. (Solidago ec ga ig var. jrssagath Michx. FI. ty a 2: 116. 1803.) Leave s lance-li near, seven to eleven times as long as wide, oo t rgins scabrous-hirtellous; inflorescence compact, flat-topped, the heads mostly sessile in Bixters of 2-5 a tins a ‘the branchlets; ae 4-5 mm. high, the So mostly broader, blunter, and often oe obvigusly green-tipped aac Ba S. eaaret. Rare in our (Aberdeen, Washington) ; British Colum o New foundland, Vir re, ars hocality: "Take St. John, Quebec. Typical J. pe see care in the aeceer dt Ba United cae J cinth 2. Solidago californica Nutt. California Goldenrod, Fig. 5491. Solidago californica Nutt. Trans. Sloe Phil. Soc. II. 7: 328. 1840. Solidago californica var. nevadensts A. Gray, age Cali. 1: 319. 1876. t i icus Kunt tems fr eep omes, dm Basal and lower cauline leaves spatulate to obov ate or “igen obtuse to acute, seeniate to base, rena Sit ded =. ee 1-3.5 cm. wide sually m r narrowly oblong, g, sharply acute to sien puberulent or glabrous ; ray-florets 8-13; disk- axe 4-11; jpn ees Tispidu Coun m dry or moist fields, clearings, and forest openings, Upper Sonoran and Transition Zones; western Orca awash the Coast Ranges and the western flank of the Sierra Nevada to San Diego County, mont fornia; also Inyo County. Type locality: Santa Barbara. July—Oct 3. Solidago canadénsis L. subsp. elongata (Nutt.) Keck. Meadow Goldenrod. Fig. 5492. Solidago elongata Nutt. Trans. Amer. Phil. Soc. II. 7: 327. 1840. Aster elongatus Kuntze, Rev. G Pi-¥e Hse 1891 id uri ipe Pp Solidago lepida var. siones Fer ora 17: 9. —— lepida var. caurina M. E ‘k, Man. Pl. Oren 7. 1941 Solidago canadensis a a Mao ey Aliso 4: 10 es from creeping rhizomes, 3-10(15) dm. gh, puberulent or pilosulose up toward (and always including) the inflorescence or pened tan densely leafy. Leaves nearly uniform, lan ceolate , 12 om: ae 1-2 cm. wi aperi and apex n or oblong-lanceolate, de, t ng to b , triplinerved, “shart serrate tire, contig mat ined, from essentially glabrous to one rid-puberulent on faces; panicle 5- = é dene. usually rhombic — broad or he the Mower branches not obviously recurved n ee “heads secund; involucre 3.5-5 m . hig Ss llaries Hae, linear- ace ray- se aera oats A 13, little phassesne ha the diole paneee hispidulou Meadows and moist openings in woods, from sea-level to 7,500 feet. Transition Zone; southern : itish Colum bia to snare coastal California and in the Sierra Nevada to Tulare County, east to the Reged Mountains; also Lower California. Type locality: ‘‘Wappatoo Island and the plains of the Oregon.”” July—Oct, ‘A variable species with rather rly marked regional subspecies and covering most of the United States and Canada o New foundland. Typical canadensts is northeastern. ng ay canadensis subsp. salebrésa (Piper) mers Aliso 4: 104. 1958. Sener serotina var. salebrosa Piper in Piper & Beattie, Fl. Paisase ag 485. 1901; S. Nekeastns var. salebros. ones, Bull. Univ. Mont. Lan Ser. hey 1910; S. salebrosa Rydb. Fl. Rocky Mts. 870. 1917; s. ceantie var. salebrosa Friesn. Butler 4: 196. 1940.) Similar to subsp. elongata, differing in having large broad panicles, with ob- viously te rec mere ine gRSOR tae lower branches bearing secund heads. Much ‘lens common in the Pacific States than the preceding; eastern Orepon and Washington to the Rocky Mountains. Type locality: Pullman, Whitman County, Washington. 4. So tua lépida DC. Alaskan Goldenrod. Fig. 5493. Solidago lepida DC. Prod. 5: 339. eulge Solidago lepida var. cmhieone DC. loc A pies a tze , Rev. Gen. Pl. 1: Hay Solidag Var. vokactuahe Cronquist, Ven Pl. Pacif. Northw. 5: 305. 1955. Stems mostly 9s dm. high. Leaves elliptic-lanceolate, usually sharply serrate-dentate and scarcely reduced up to the inflorescence ; panicle short and compact, not at all secund; involucre not very im since a ces ob cae aries more than half as long as the inner. Otherwise similar to S. canadensis s ubsp. elon Moe ground, Humid Teanaton Zone; Saddle Mountain, Clatsop County, Oregon; Vancouver Island north to southern Alaska. Type loc : Nootka or Multgrave (Yakutat Bay). July—Sept. SUNFLOWER FAMILY e : ss 5491. Solidago californica 5492. Solidag densi 294 COMPOSITAE 5. Solidago gigantéa Ait. Smooth Goldenrod. Fig. 5494. Solidago gigantea Ait. sain Kew. 3: 211. 1789. Solidago serotina Ait. loc. cit. Not Retz. 1781. Solidago serotina var. ousotes A. Gray, Proc. Amer. Acad. ai Na 1882. Aster latissimifolius var. serotinus Kuntze, Rev. Gen. Pl. 1: 1891. Solidago gigantea var. leiophylla Fernald, Rhodora 41: 457. es o S. canadensis; stems stout, 6-20 dm. high, glabrous and often glaucous below the inflorescence the latter pilosulose. Leaves lanceolate to elliptic-oblong, =" ong, mm. wide, acuminate, sharply serrate above the entire cuneate base, ppc ipeibeabd cata essentially glabrous on both sing panicle large, pyram midal, the heads secund on t ed branches ; involucre igh, the phyliaries rather firm and often obtuse; ray- aBiiets 9-16, dis- ti ad surpassing the disk ; achenes hispidulo ets and meadows at low elevations, Arid ea sition Zone; southern ig | — to heel (mostly east g the Cascade Movgiains) east to Te eaniiint.. Cer a) and Texas. Type locality: North America (the type a garden plant). July—Sep 6. Solidago missouriénsis Nutt. Missouri Goldenrod. Fig. 5495. Solidago is Nutt. Journ. Acad. Phila. 7: 32. 1834. Solid tol: A. Gray, Syn. Fl. N, Amer. 17: 151. 1884. er missouriensis Kuntze, o% Gen. Pl. 1: 318. 1891. pn tolmieanus Kuntz a Solidago missouriensis v l c ist, Vasc. Pl. Pacif. Northw. 5: 307. 1955. _ Stems from creeping ae < or clustered from a simple or branched caudex, 2-5(-9) d high, essentially glabrous below the parses peerwient inflorescence. Basal leaves ae anceolate, 5~20 cm. 5-20 m th bo obl ng, wide, entire or toothed above, tapering into margined peers. scabrid-ciliolate, often lost pee the paling secpely t educed above, the upper mostly linea subulate, entire; panicle usually oblong or rhombic, with erectish branches, sometimes pyr. ae, with spreading branches, the heads then Secune ; involucre 3-5 mm. high, the phyllaries en broadly lanceolate, obtuse to acutish, rather firm, the oe gland prominent ; ray-flor 8-13), u soatly distinctly cecodiien the disk; achenes hispidul Rat a open places, U po Bon age and Transition Zones; British e PRa to yale Oregon east to Guitarto, ae exas, and A ona. A’common and variable species from which a number of varieties have been segregated. Type locality: son "the upper branches of the Missouri and in Arkansas.”’ July—Sept. 7. Solidago guiradénis A. Gray. Guirado’s Goldenrod. Fig. 5496. Solidago guiradonis A. Gray, Proc. Amer. Acad 6: 543. 1865. Stems slender, erect, fro woody rhizome, 8-10 dm. high, Regents. < throughout. Basal leaves lanceolate, tapering to File beticke. 12. 15 cm. long, 5-10 m wide, the cauline elongate but reduced, linear, above becoming linear-subulate bracts, all gntice: whi tte ee nder, sub- racemose, few-headed, not secund, 10-20 cm. long; i nvolucre about 4-5 mm. high, t e phyllaries in broad; ray-florets 8-10, little cacenda the disk: disk- aah 10-12; lrg e f — i 5 * ance- achenes pu Moist ie Upper Sonoran Zone; California, rare, in the Coast Ranges from Santa Clara County south- So and in the southernmost Sierra Nevada. Type locality: base of San Carlos Peak, San Benito County. Sept.- 8. Solidago confinis A. irk gee Goldenrod. Fig. 5497. Sol 0 confints A. Gray, Proc. Amer. Acad. 17 188 perks confints f. luxurians H. M. Hall, Univ. ne a Bot. 3: 46. 1907. Solidago confinis var. luxurians Jepson, Man. Fl. Calif. 1035. 1925. Stems usually stout, terminating rather short woody rhizomes or sometimes clustered on a short caudex, 3-14 dm hi gh, glabrous throughout. Leaves thick, pale green, entire, glabrous or th b id, the basal spatula ceolate to ol apering t y den: ong. a, igh, he dhyiiarics | ate slig' htly ckedeate. linear-lanceolate, acute or acuminate, rather firm EPS Tas s 6-10, scarcely surpassing the 11-21 disk-florets; achenes sparsely idsas to canescen Dry or moist banks, Upper Sonoran and Transition Zones; Ventura County to San Diego County, gaat to the gf Han edge, mostly in the mountains. Type locality: “southern borders of California.’ July—Oct 9. Pee spectabilis (D, C. Eaton) A. Gray. Basin Goldenrod. Fig. 5498. Solid: var. spectabilis D. C. Eaton, Bot. Pog a 184, 1871. Solidene ok Gray, Proc. Amer. Acad. 17: Stems rather stout, terminating a rat see a woody rhizome or caudex, 4-13 dm. high, glabrous throughout or becoming somewhat hispidulous within the inflorescence. Leaves — or rarely remotely ate, the basal oblanceolate, acute or obtuse, © a lon: ing’ clasping petiole, including the peace 28 cm. long, 1.3-4 cm. wide, the upper ‘tae neat lanceolate and often much reduced toward the inflorescence, scabrid-c Sickete,. eka ae noni a = panicle usually oblong and very reac abethy less than 10 cm. long; involucre about 4 mm. high; SUNFLOWER FAMILY 295 phyllaries linear- ebtong, obtusish; ray-florets 11-15, little exceeding the 15-22 disk-florets ; achenes puberulent Alkaline meadows or bogs, Sonoran and Arid Transition Zones; southeastern premn south to Death Valley, California, pote to Utah. Type ieeality: mountains of western Nevada. July—Sept. 10. Solidago spathulata DC. Dune Goldenrod. Fig. 5499. Solidago Fame ers DC. Prod. 5: 339. 1836. Homopap + nee ethopets Nutt. Trans. Amer. Phil. Soc. II. 332. 1840, Solid Torr. & Gray, Fl. N. Amer, 2: 202. Aster candoliés Kuntze, i Gen. Pl. 1: 315, 1891. Stems stout, from a caudex woody rhizome, 2-6 dm. high, usually glabrous throughout, glutinous pearl) above. Basal. leaves hioatty obovate to spatulate-oblan ite he mos oy blunt or rounded, crenate- Maier ta apering t o the ee the cauline similar but uced, the eel most acute and subse ~ he sat ina simple Or ompound, sometimes racemi aie thyrs e628 c long; involucre ws igh, phyllaries 9 oblong, yald blunt; ray-florets 7-9, pelseety exceeding the 10-16 ‘disk- oreks Laisizn den agg pu Sandy coastal hills and dunes, Humid Transition eas ; Coo: i Coe ee Oregon, to Monterey, California. Type locality: Monterey, California, erroneously given as i" pod ng . * Collected dig Haenke. May—Oct Solidago spathulata subsp, glutinédsa (Nutt.) Keck, Aliso ie Jot 1958. {5 ofons contertifora DC. Prod. 5: 339. 1836, not Nutt. 1834; S. glutinosa Nutt. Trans. “eg tga bie S. vespertina Piper in Piper & Beattie, Fl. Northw. Coast 365. 1915.) Stems 8 dm je or les sie ge eit ng glutinous and apuredy hispidulous within the capac hice basal leaves nacrowly phoriars to narrowly Bt ee ie) eae ao w oe fa) fw = w ae a ue QO o | (e} ot QO te Oo w nal = 1 ate 5 < ° S. @m-< c Hp, ‘he inner oblong, acuminate-tipped, t rrow hyaline margin somewhat erose; corollas -8 aie Mast ng, the lobes about 2 mm. Tae achenes pilose, glandular saeviek pappus copious, ssbb the co Dry, alkal Sak or silty soils, Sonoran Zones; Owens Valley and — Valley region, California (rare), east to the Salt 5 aes He Desert, Utah. Type locality: Wells, Nevada. Aug.—Oct 4. Chrysothamnus viscidiflérus (Hook.) Nutt. Sticky-leaved Rabbit-brush. Fig. 5506 Crinitaria viscidifiora Hook. F1. Bor. Amer. 2: 24. 1834. elovia viscidifiora DC. Tres Y ie eno 1838. hrysothamnus viscidiflorus Nutt. Trans. Amer. Phil. Soc. II. 7: 324, 1840. Sscidi Linosyris ora Torr. &G N —. 2:2 elovia douglasit A. Gray, Proc. A : 645. 1873. PvE Su douglasit var. tortifolia A. Gray, si ci ca er viscidifiorus Kuntze, R n. 18 idi, ev. " ide Sasa viscidiflorus var. tortifolins Greene, ‘Erythes 3:96. 1895. Chrysothamnus tortifolius Gre 18 Chr ysothamans douglas Cea & iat gi sariay Mt. Fis. 266. 1914. Ri fastigiately branched, white-barked shrub usually less than 1 m. high, the plo brittle. Leaves linear or i re er olate, flat or twisted, erect, spreading, or reflexe 2-5 c 1 long, 2-5 mm. wide, 1-3-n d, glabrou oar sometimes scabrous -ciliolate, viscid, sometimes with punctate e glands ventrally ; ends about 5-flowered, in term phere broad cymes; involucre 5-7 mm. high; phyllaries ihear'ctdote to lanceolate, obtuse i a purse monly mucronate) to acute, not keeled, strongly graduate but in o| re cal rank chartaceous; achenes + ie villous ; pus brownish white. A highly sat com ies Dry plains and hillsides, at low or moderate elevations, Upper Sonoran and Arid lg cong ~ age east of the Cascade-Sierran axis from soabieeret British Columbia to southern Castor. east to New Mexico and ogre tana; this CO haa subspecies most frequent in Pood by in in Rocky Mountain Type locality: Garren: Slain of t Columbia, from the Great Falls to the Mountains.” Collected by Douglas. tae rae A fk of several te x intergrading subspecies of overlapping distri Rosen Key to Subspecies Leaves glabrous (sometimes viscid or with punctate glands ventrally) at least on the faces, the margin sometimes scabrociliate. Shrubs mostly 4-12 dm. high (shorter in some wide-l re wide. Leaves Ante to linear-lanceolate, tian to base and apex, 2-5 mm. wite: 4a we aaneks acute acumi- widespread. su ubsp. Ciciditorss Pop Seoniity elliptic to elliptic-oblong, 6-12 mm. wide, 3—5-nerved, -obtuse but mucronate; shrubs often only 2-4 dm. high; nearly limited to northeastern Nevada. subsp. latifolius. SUNFLOWER FAMILY 299 hrubs mostly 1-3.5 dm. high; leaves linear or linear-filiform, 0. 5-2 mm. wide. subsp. pumilus. a Sh Leaves more or less densely puberulen Leaves mostly 2.5—6 mm. wide, 3—5-nerved. subsp. a Leaves 1-2 mm. wide, 1-nerved. subsp. pub : Chrysothamnus viscidiflorus subsp. aay 3 {D.:G. —— Hall & Clem. Carnegie Inst. a Publ. No. 326: 184. 1923. (Einosyris “viscidiftor ra Jaane D. Heaton, Bot. King Expl. 157. 1871; Bigelonia 646. Sars red seothhtonni viscidiflorus var. latifolius +152. 1906.) Mountains of northeastern douglasti var. gt hey i and Clements but doubtfully a mem- Greene, Erythea 3: 895: Revege bg en adjacent Sadako. Re eport C. latifolins , Riyadh, the rorrey Club d from northeastern California Be Hall Chry amnus viscidiflorus subsp. gan elescag, Hall & Clem. Carnegie Inst. Wash. Publ. No. : r. Phil. Soc. II. 7: 323. 1840; Pawongy Jeed rrees oe haaag . 5&: 80. age ig Sie douglasit sag _stenophylla. A. Gray, Proc Aca sien 1873; Kf do ay @ var. ae 5 A. Gray, Syn bh ee | 884; Caruahannee apne iat Se Gree Ery- idiflo rus var. stonophsllus ee M. Hail, Univ Calif. Pub. Bot tons on 1907; res gin pot A var. horus subsp. stenopiyllus Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 183. 923 ee tlus Jep Pi. ig 1031. 1925.) Dry, sagebrush-covered tn and ridges, often in alkaline Pwils, from a0 core mn m Washin gton to Inyo County and the San Bernardino eastern Montana and Uta are re the southern Rocky Mountains and so n Great Basin. Type locality: ‘fon the ee of Lewis’ River and the Rocky Mountain A at Chrysothamnus viscidiflorus subsp. Nentesthige (N -ya 8 Hall & Sages eee Inst. Wash. Publ. No. 326 181. 1923. (che ysothamnus lanceolatus Nutt.-_ Trans, Phil. Soc 324. 1840; j, Linosyris leubiolea Torr. & Gray, Fl. N. Amer. 2: 233. 1842; meee S lanceolata A. fam Prot Amer, Acad, 8: = 1873; A baci a : Chr sdthamnus visete Me us var. nceolatus douglast Greene, Ery Washington (frequent) ‘and ea cality: ocky Pessoa ag wowada ah ces of the amnus vis pada hy subsp. Sabian ( one Hall & Clem. Carnegie 4 Wash. Publ. 3 | 19 ( othamnus pumilus euthamoides Nutt. Trans. Amer. Fae Soc. II. 7: 323. 1840, a . Eaton, Bot, King Expl. 1 i 89 herala Gray, aa oar Eee 8: C. humilis Greene, Pitton Publ. No. 326: 182. 1923: c * iseldifiorns v var. humilis Jepson, loc. cit.) Sag favtuah eastern Oregon mgr: ‘California to the Sa Bernardino Sie — path pe og Utah and Yellowstone Park. Type 5502. Petradoria pumila 5503. Chrysothamnus paniculatus 5504. Chrysothamnus teretifolius 5505. Chrysothamnus albidus 300 COMPOSITAE 5. Chrysothamnus axillaris Keck. Inyo Rabbit-brush. Fig. 5507. Chrysoth illaris Keck, Aliso 4: 104. 1958. rounded shrub 6 dm. high, intri icotely branched, =~ branches reese white-barked, , “i sharply defined vertical ranks, spree: at maturity, broadly linear, sharply acute or apiculate, the costa glandular-thickened above, the small herbaceous tip ciliolate ; corolla about 5 mm. long, post lobes 1-1.2 mm. long; style- anpentages longer than the stigmatic portion; achenes sericeous ; pappus tawny, not very long or copious Desert slopes in granitic sand, Upper Sonoran Zone; eastern Inyo County, California, and adjacent Nevada. Type mena head of Lay Springs Watley peers County. Sept.—Oct ated to C. albidus (M. E. Seger Greene om which it differs in having t o few lea and these not at all poreney puncta and in the smaller involucres and florets. 2 ae pila Rag more Gatnwai related to C. greenet (A. Gray) Girton subsp. filifolius (Rydb.) Hall & a nt known only from eastern Nevada and eastward, weak has plane rather than tightly involute leave: 6. Chrysothamnus depréssus Nutt. Long-flowered Rabbit-brush. Fig. 5508. Chrysothanenns see tok Nutt. Journ. Acad. ong 1E.4-2,171;.. 1847. rr. in Sit 61 Bigelovia depressa i. pS Pree: Riau, rn e 643. 1873. ion eyes with many erect herbaceous stems from a much-branched, spreading, woody crown, 1-3 d igh, cinereous with a dense scabrid puberulence. Leaves oblanceolate or m maces in com nal cymes, 5- . phyllaries onal ele -seria , in 5 pr ly defied vertical ranks, lan minate, drawn to a soft o, strongly keeled, oe outer metas ceous and minutely scttesiand “the 3 inner Shee scarious, wi with es line margins; achenes glabrate; pappus s brownish white Dry canyons, Sonoran Zones; occasional on mountains of e n Mojave Desert, San Bernardino Coun California, east to southern Colora 'do and New Mexico. Type toate: ~ ey ieee as the type see tn - labelled; not “‘in the Sierra of Upper California,” as stated by Nuttall. Aug.—Oct 7. Chrysothamnus gramineus H. M. Hall. Charleston Rabbit-brush. Fig. 5509. 4 gramineus H. M. Hall, Muhlenbergia 2: 342. 1916. med from a branched roe caudex, 2.5-6 dm. high, light green, essentially gl one dicsodout the striate-angled stems si imple or erect-branched above. Leaves equally nes nar in: distributed, the larger ones wly linear-lanceolate, 3-7 cm g, ; ate, oa i , coriaceous ; incase mostly subsessile in small ee clusters, 4—5-flo ; pale ; involucre cylind 0-13 high; phyllaries 4-6-seriate, strongly ate, ob scurely pec ah ranked, stramineous, oblong, 1-nerved, abruptly otepiiate at the truncate or retuse, ciliolate apex; corolla abou As «i m. long; penile linear, ellipse about 6 mm. long; pappus sone stramineous, 8-9 m Rocky wooded slopes, 7, fig 9 pe Arid ae Zone; Inyo and Panam ntains, Inyo County, California: Chasiestan Mountains, Nevada. Type locality: head of Lee Canyon, Saco ovutatnn, July—Aug. 8. Chrysothamnus parryi (A. Gray) Greene. Parry's Rabbit-brush. Fig. 5510. Eteonyrit re = — Proc. Acad. Phila. 1863: 66. 1863. 1 y, Proc. Amer. Acad. Cc. he-ysothomuns fewer Grccae. Hivtues S.213. Shrub up to 5 dm. high, the nu choy Sie — Lesage erect or spreading, densely clothed with a white, gray, or greenish yellow tomentum eafy. pa narrowly linear to sie 1-8 cm. long, 0.5-8 mm. wide, —. ed: heads in ep ike ae racemes or caceaiicien! icles, te) = nn Mountainsides and flats, Arid Transition Zone; hater to New Mexico and California. July—Sep Variable in habit, pubescence, foliage, inflorescen Ba ae of head. The typical form, ranging from Wyoming and Cohanaile to eastern Nevada, has lin cag Bo 3-nerved leaves 1.5—3 mm. wide. Sh aahat flowered heads, and 10-15 er in oe — ranks. The 5 dalicniae subspecies occur in ou Ch: thamnus parryi p. bolanderi (A. Gray & Hall & Clem. Carnegie Inst. Wash. "Pu ae Ro 326: 199. 1923:. € — fe heletaeche “Coy y, rc Amer. ‘kent : 354. 1868; sane Leder at Aa Gray, op. cit. 2: 641. 1873; oe * landeri Kuntze, = Gen, Pl-1: 317. 1891; Chr ysotham nus bolanderi ig + eyes io 85234. 1895; M ; nderi Greene, Leaflets Bot. Obs. 1: 81. 1904; C. hrysothinaans parr oy andert epson oo Calif.. 1033. 1925 .) Leaves 3-4 cm. long, 4-5 mm. wide, green, viscidlandulars heads us flowered, in short, sometimes branched racemes; involucre 9-10 mm. high, the phyllaries about 11-15, in pee: ranks. Known -— from the type locality, Mono Pass, Sierra Nevada, California, at 9,000—10, ay feet siehade: Arctic Alpine = us parryi subsp. latior Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 199. 1923. (Chry- siahelenet ey emia var. latior Aepaen,

a ie complete synonymy is not given; only those names applied to Pacific States entities are listed. Plants perennial. St s monocephalous, subscapose, a short caudex terminating a wee, ag root; alpine or subareiine plants with basal tufts ot if grese like leaves. . OREOSTEMMA, Stems usually with s sae to man uf ne , not arising from a deep conical root. Heads solitary at tips of s epee nches; low-tufted plants with narrow, a. Benoa cuspidate leaves; ones soubie the ont brite shorter than the in NTHE, Heads several or numerous, if ra spice not with teaves as S ihcies pappus pond in a single row. Plant t wont toward bas aa ucous and glabrous, paniculately mach branched, with discoid ine else herbaceous, glabrous, Grate temmed, often spiny, and ap Bs esi a a, naked, Plants otherwise. Phyllaries oblong to ovate, rarely 1 nee-linear, chartaceous with Bcd scarious margin and often herbaceous tip, more or teas keeled by the strong midne = apy Nee Phyllaries Poin inear (varying to ppt Ad = ee below and i eo or the outer herbaceous anbipe Se withou ong midne Bias, pes! leaves a often toothed or loon Sar and ait at all oe omits sessile, grass-like lea Plants (in ours) wit th sessile, grass-like leaves at base and above, glabrous cnlaek for the glandular inflorescence VI. OrtTHOMERIS. Plants annual. Phutt ry 4. tly graduate; rays in one series, evident d r ing the pappus. VIT. Oxytrirotivum. Phyllaries subequal or slightly graduate; rays if present in more than one series, very inconspicuous and equa! aling or shorter than the mature pappus. "VIII. Conyzopsis. - naar gouge . : a Pee | 3 Aunt Involucre (at least on the margins of phyllaries) and usuall zg Le apres lancemate to chownte; heads menetly. few, comparatively jarge. din: Be “ovate, eat clasping; pterie ies graduate, the Leaves sharply serrate, - mostly lance- ly ; oe on Leaves entire, at least the lower obovate, with cons spicuously sping. a phyllaries but little gra ad ante, the indurate base shorter than the cuplreasone tip a : pra Leaves thin Leaves linear or Hise naiestite, 6 mm. wide or less; heads small, numerous. is aa delet T LK 2 3.4. ee | dela: nat eg seam with purple tips or margins, oblong to fapecolats, usually strongly graduate; leaves obovate to elli iptic -oblon; g, firm, usu ually rough beneath or on nasi ides n a nearly caked ~— or cymose panicle. . A. radulinus. Leaves sharpl y" Leaves entire to obtusely serrate; heads 1 to few, in axils of leafy ee . sibericus meri: Phyllaries not purple-tipped nor margined (except in pfaludicola and some ‘forass of porkesitatag ‘weilatle in shape; leaves not rough beneath (except A. greatet at). Heads Iam 4 numerous, in an open, near] arger leaves 2-4 cm. wide. very pale green; 5: lens more cise essentially glabrous. . A. laevis geyeri. 312 COMPOSITAE Stem and leaves not glaucescent; leaves thin; achenes pubescent 8. A. ee atat Heads, when numerous, in leafy-bracted Sack or nearly naked cymes or cymose panic Heads mostly not small, not racemosely arrange san ft the tb mp eck rays Goaanny blue to purple; phyllaries not ie nincronaint e, the pute 3 appressed or spre posh leaves obovate to postin usually more eae 2 cm. nig e, “apt conspicuously clasping bases. eads crowded toward tips of stem and branches, usually short-pedicelled or subsessile; involucre 6- ‘3 mm. high, rarely Teaiy: bracted, the a ee s asap pilosulose. Heads solitary or few at tips of stem and branches ei er Ea . A. foliaceus. pene © — lanceolate or linear, 0.5—2 cm. wide, slightly if at oe clasping. Plants of cold bogs; heads aalitiees at pia of widely peter te ents or ne aap paludic Plants not of cold bogs; heads not oe. at pig of widely divergent branches or branchlets. Inflorescence a leafy panicle; heads y (sometimes few in A. subspicatus). ches of au and ae cles aan caeals os ag with small leaves; involucre usually try Sitghtly: graduate (more so in A. hesperius). Leaves mostly linear or linear-lanceolate, entire, gee cas rarely serrate. Coarse plants 1-1.5 m. high; pubescence of s s and branchlets in Pee urrent lines from the igs hance: “involucre, rarely if at all with pest ual subtending bract 10. A. hesperius. Plants usually less than : m. high: pubes: cap em uniform, or if in lines at least uniform under dhe heads je neatly glabrate. eaton Leaves Poagenicnt cidanceerate or lanceolate, even the = middle one ones mostly serrate xo: tu —_ of the: cane’ and en cisaebindenty ibtlaeis with small aves; involucre strongly graduate. A ooo: open, sometimes widely Greceent. namie: involucre 7-10 mm. haa ge of er : : 1 4 hil NSIS. Inflorescence a close panicle or raceme; involucre 4-6 mm. high. He rate Mapai ebbing pubescent rah ee hairs; plants of uthern mgs 19. A. bernardinus. Socbace essentially areas haem y scabrous, on the leaf-margins) ; plants of western Washin and Ore Inflorescence essentially a naked cyme or cymose He oy head usually few (often ameny in A. adscenden ax +. raduate, in 3, sometimes 4, s ries. Stem-leaves mostly linear, not reduced i in size on the upper stems; branches of the 16. A. sanamens: Upper stem-leaves fe ws 8 reduced in size; the cymose panicle open with spreading bra 17. A. occidentalis intermedius. Involucre scarcely cha the phyllaries in 2 series. ne 17. Heads — and very numerous, racemosely — arranged on the Reger spreading = a rays white (r y purp ple ) 7P Plants sit well-developed creeping rhizomes; disk 6-7 mm. high, the rays 22-30. . A. falcatus crassulus. Plants with stems clustered from a short rhizome or caudex; disk 4-5 ny eect the rays 12-20. . A. pans II. EucepHALus. e pi ly pale beneath with a more or less dense tomentum. Rays 0-4; inner phyllaries ovate, obtuse, canescent. . A. brickellioides. ay 3 inner phyllarie es lances, gla ndu ook 25. A, ledophyllus. Leaves not conspicuously pale glabrous t pub t, or sparsely pilose. eads discoid or occasi sana lly ga 2-3 Inflorescence corymbiform with daa branchlets; peduncles glabrous or somewhat F7 Sane beneath the involucre. 23. A. siskiyou Tadeene ake, an elongated panicle with short branchlets; peduncles eagerer ar an somewhat hickite. . vialis. Heads radiate with 6-18 r Involucr Fok agua ae phylaries closely imbricate and appressed in age; leaves scabrous-puberulent on surface s and margin 26. A. perelegans. —— — hemispheric, or campanulate, the phyllaries more or less loosely imbricate and spreading Inflorescence c orymbiform , with several to many heads; plants robust, 6-15 dm. tall. es glaucescent, soit eight or more times as long as wi me, the margins obscurely serrulate. pee A, REMCeTC Ene: Leaves not glaucescent, three to six times as long as wide, tl 28. A. pip Ay Heads a rarely 3 or 4; plants 1.5-3.5(5) dm. tall. from a woody caudex; phyllaries lanceolate-acuminate ; acgr of the ane Mountains. paucicapitatus Plants from a creeping woody -eipiaae phyllaries oblong-ovate to ovate; jeri of the vicinity of Mount Jefferson, Oregon 30. A. gormanii. II. TANTHE. Leaves linear, usually 2-3 cm. long, not definitely callous-margined; plant green. ; 31. A. stenomer Leaves chiefly elliptic or linear-oblong, usually under 1.5 em. long, with whitish earns margin; oiant subcinereous. . A, scopulorum. SUNFLOWER FAMILY 313 + ee Stems and involucres more or less ndular Stems poe involucres more or + Silos wells: serbian see Pe cm, tall page gerne caries. in subsp. ander- it). at el Stem nd glabrous; plants 30-70 cm. tall. ae A.e Stems aivotsbede more or oan i -scaberulous ee pirate LeucosyRis. Plant essentially herbaceous, not glaucous, often with spines in or above the leaf- ge th = radiate, white; achenes glabrous. 36. spin Plant shctsbiey. glaucous, not spiny; heads discoid, yellowish; achenes pubescent. % 37, A. intricatus. VI. OrtTHoMERIs. Pappus simple; leaves grass-like; a single species. 38. A. pauciflorus, VII. Oxyrripotium, Syprte glabrous; leaves —* or lance-linear, usually entire; heads numerous, panicled; rays pink, little- serted; a single specie 39. A. exilis. VIII. Conyzopsis. Phyllaries mostly oblanceolate or spatulate, obtuse; rays about 2 mm. long. 40. A. frondosus. Phyllaries linear, acute; rays wanting or vestigial. 41. A, brachyactis. 1. Aster conspicuus — spires Aster. Fig. 5526. Aster pre er Lindl. in Hook. Fl. Bor. Amer. 2: 7. Aster macdougali Coult. & Fisher, Bot. Gaz. 18: 301. ioe anise k elongate, woody, stoloniferous; stems usually solitary, stout, simple below the in- florescence, usually 30-60 cm. high, leafy, densely glandular, pee ya Si n the at ea sparsely hi bov. pilose or hirsute. Leaves ie ores o oval or obovate- he | r 8-17 ng, 2. wide, acute, sessile and usually clasping, sharply bois except tow base, y firm, veiny, scabro yates ; heads f r numero mids in a usually rounded cy or cymose panicle; involucre broadly campanulate, ica parted e, about 6-seriate, 7-10 mm. high, the places Fistor ohlee to lance- = chien g or oblong, iceaaly. "grandular, strongly ciliate, with vole, in ot , l-ribbed base and usually s Cou often spreading, acute to acuminate, her- feos tip ey shoe sometimes entirely herbaceous; rays about 20-30, violet, 1-1.5 cm lo ae ae ieaced pubesce Forests and open woods, Arid ee, and Canadian Zones; British Columbia south through eastern Wash- ington to northeastern Oregon om east to nag pectin Wyoming, eat South Dakota. Type locality: “Carlton o the Rocky Mountains.” July—Sep 2. Aster integrifélius Nutt. Entire-leaved Aster. Fig. 5527. Aster integrifolius Nutt. Trans. Amer. Phil. Soc. II. 7: ap 1840. Aster amplexifolius Rydb. Mem. N.Y. Bot. Gard: 1: 391. 1900. Stems several, stout, ascending from sake woody, fibrous-rooted rootstock, 2-7 d igh, often sat PN from densely erieede pilube to nearly glabrous below, d ensely ‘tipitate- glandular illous above. Lowest Neaves obovate to spsacric pasa ine blade a nate at i 5-18 cm. lo i each end, narro ; ed, v ge eu tially glabrous to short-pi ; lower stem-le bovate o gp cote "teste pity strongly clasping, the middle and fanae raid much smaller, elliptic “8 lance-linear ; heads few to several, 5526. Aster conspicuus 5527. Aster intergrifolius 314 COMPOSITAE 2-4.5 cm. wide, mostly solitary at tips of peduncles, in an oblong false raceme or narrow panicle; involucre broa dly campanulate, pa 3-seriate, little graduate, the phyllaries arts linear-oblong, with short indurate base and a r acuminate, erect or spreading, herbaceous tip o outer wholly i densely g aera ten deep sets tinged; rays about 10-18, ite « or gale about .1 cm. long; achenes Be rt-pilos a sede granite slopes se oo. of meadows, Canadian and Hudsonian Zones; ains of sion pry htm ononcgeeg —-! of begginbs ig south to Tulare County in the Sierra Pinte ‘Calitardin: eastward to Montana and Colorado. Type locality: the vicinity of Thornberg’s Pass, Blaine County, Idaho. July—Sept 3. Aster modéstus Lindl. Great Northern Aster. Fig. 5528. Aster unalaschkensis B? major Hook. Bor. Amer. 2: 7. 1834. Aster modestus Lindl. in Hook. op. cit Aster sayianus Nutt. Trans. ange a Soc. II. 7: 294. 1840. Aster mutatus Torr. & Gray, Fl. N. Amer. 2: 142. 18 Aster majus Porter, Mem. Torrey Club 5: 325. 1894. Stems usually solitary from a slender stoloniferous rootstock, ri very leafy, 0.3-1 m. high, simple below the inflorescence, densely stipitate-glandular above or thro ughout, and usually sparsely or heats villous, sometimes glabrous below. Leaves nearly anitncas linear-lanceolate to oO 2: cm. lon 3 cm. wi sile or laspi sm above, g heads L cs ey cymosely a ged, 2-3 cm. wide; involucre hemispheric c, 2-3-seriate, subequal, ut 7 mm. high, the phyllaries ‘ances or linear- lanceolate, acuminate, spreadin g tipped, ne gd stipitate-glandnlar, often purplish, the outer herbaceous throu ghout, the inner po ae gin ow ; rays about 20-30, purple or violet, about 1 cm. long ; achenes strongly about 5- ribbed. Ha NS pubescent. Poe a woods and stream banks, Arid Transition and Canadian Zones; Alaska to southwestern Oregon east Minnesota, se estern Ontario. Type locality: “Mount tain "woods, at the mouth of the Smoking platy ens 36°. ” July—Sep 4, one radulinus A. Gray. Rough-leaved Aster. Fig. 5529. Aster Pree al Gray, Proc. Amer. Acad. 8: 388. 1872, Aster yt Porter pose eae Cha 172 37: ol te 1890. Aster . iasit iA. Nels. Univ. Wyo. Publ. Sci. 1: 924. Stems solitary or several sae a sae - icisidboes rootstock, 15-60 cm. high, usually simple below the inflorescence, ascending or erect, from rather densely 'spreading-hirsute to inc curved- otal or strigillose, sometimes cn below, not glandular ; leaves reduced to entire bracts e, the lowest obovate to oval, obtuse to acute, narrowed to short, ciliate, margined petioles, the i 4- sonnei and middle ones obovate to oval or tances oblon ng, 4-12.5 cm. long, 1-7 cm. wide, acute, sessile, not clasping, sharply serrate or d ntate except on the cuneate base, firm, scab ous-pube t be- eath, scabrous or smooth above ; heads 1.5-2.5 cm. wide, several to numerous in a flat or rounded cyme or cymose bag a the peduncles densely hirsutulous or hirsute- pilose ; Gestine cre 5—7-seriate, gronely graduate, 6-9 mm. high, the phyllaries chia to (inner) linear, usually acute, hirsutulous base an n back, ae with pale indurate short Pi sahie'y = very rarely spreading, her- fakes ‘tip, at least the inner often purple-t s 10-15, pale violet to white, —11 mm. , th ok ia ed forms ecking farsle’ a in ‘he phyllaries; achenes sparsely hirsutulous ; pappus ie ie slopes in galery and mountains, mostly Transition Zone, Vancouver Island and central Was! ington ‘eos wbrtg ~ (west of the Cascade Mountains and Ochoco Mountains on the eastern bea ” San ha’ Obispo Co eanky and to Mariposa Couey in the Sierra Nevada, California; also from Santa Cruz Island, Santa Barbara Conuier, Type jcalty: Beck July—Oc The form described as Aster aioxii lt to some degree robust forms of A. sibiricus var. meritus in habit but the 1 ranges and tats of the two do not overlap. 5. Aster sibiricus var. méritus (A. Nels.) Raup. Arctic Aster. Fig. 5530. Aster sibiricus A. Gray, Syn, FI. N. eon§ 12: 176. 1884, Not A. sibiricus L. 1753 Nels. Ga 1904. Aster bakerensis St. John, in St. J oa yi gi ig Mazama 11: 93. 1929. Aster richardsoniti var. meritus Raup, Contr. Arnold Arb, 6: 204. 1934. Aster sibiricus var. meritus Raup. Sargentia 6: 240. Stems 1 or several, often tufted, ascending or spreading to suberect, from slender stoloniferous rootstocks, 3-30 cm. high, apie or little branched, usually purple, pilosulose with usually loosely ascending hairs. 5 ph sya adually or abruptly reduced above, obovate to elli ptic or the upper m. wid r herbaceous throughout), at least the inner with conspicuous purple tip margins; rays about — — to violet, 7-12 mm. long; achenes sparsely Fansite ppaitanrn. about 8-ribbed; deeper brown than in rer radulinus, usually purplish-tinged. SUNFLOWER FAMILY 315 Open rocky slopes, reaching 3,000 meters elevation, Hudsonian and Arctic-Alpine Zones; South Dakota _and Wyoming to eee A and British Columbia; known in our range from the northern Cascade Mountains in hg ington, and the Wallowa Mountains, eastern Oregon. Type locality: Yellowstone National Park. July—Aug 6. Aster campéstris Nutt. Western Meadow Aster. Fig. 5531. Aster campestris Nutt. Trans. Amer. Phil. Soc. II. 7: 293. 1840. Stems usually several, ascending from slender s ioe ls rootstocks, 1-5 dm. high, slender, simple or much branched, pet insets stipitate-glandular at least above and usually more or feed strigilose, often _Blabr elow. m-leaves rl ice, i to li spatulate, 2-5 em. long, 2.6-8 m ae obtuse to acuminate, sessile and often ptt eping, entire, firm pale ree ciahinste wind glabrous to hispidulous, 1-nerv popu sd a pair of weak lateral veins ; branch-leaves much reduced, linear; heads 1 to ma ide. a "4 usu: ay narrow cyme or panicle ; involucre about sap dcie subequal o r slightly grad die 5 m. high, the phyl- laries linear to lane eae acute or acuminate, appressed or slightly Sertaticle, | Bendel Bcd ded with whitish base a orter or ‘longer pkey ous tip, the outer often herbaceous throughout ; ky. nd s rays about 20-30, violet or hp idl 5-8 m ; achenes sil Meadows and dry open slopes, often in si soil, Aad Transition Zone; Montana and Wyoming to paar a Columbia and eastern Washington and Oregon. Type locality: ‘‘along the plains of Lewis’ River.” Aster campestris var. blodmeri A. Gray, Syn. Fl. N. Amer. 12: 178. 1884. (A. bloomert A. Gray, Proc. Amer. Acad. 6: 539, 1865; A. nanneria ery eh: bebe al coos U.S. Nat. Herb. 11: 572. 1906; nats gp og Peck, Proc. Biol. Soc. Wash. igh o less; stem glandular least above involucres glandular ras in the rete ay Tae readin MBs spreading- ivsuan: This nether poo cane defined variety ‘icuianan A. campestris var. pd trenton oan Klickitat adios Washington, southward ype of ce Cascade Mountains) cha the eastern side of the Sierra Nevada and ott Be hess in the region of Lake Tahoe. Inter- grading for are to be found particularly in the northern par its range. Type locality: “thigh slopes of Mount Khaciddens, near Virginia City, Nev: ea er modestus 5530. Aster sibiricus 5531. Aster campestris 5528. Ast 5529. Aster radulinua 316 COMPOSITAE 7. Aster laevis var. geyeri A. Gray. Geyer’s Aster. Fig. 5532. Aster laevis var. geyert A. Gray, Syn. Fl. N. Amer. 12: 183. 1884. Aster brevibracteatus Rydb. Mem. N.Y. Bot. Gard. 1: 392. 1900. Aster geyeri Howell, Fl W. Amer. 1: 308. 1900 Aster pickettiana Suksd. Werdenda 1: 42, 1927. 2 Stems erect from ; stoloniferous rootstock, 0.3-1 m. high, simple or branched, glabrous and glaucescent throm ughout or the peduncles and branchlets very rarely (also the st ) pubesc n em) pubescent i lines leaves us scale doa ate, obtus se, Bersuanssg deciduous; lower stem-leaves oblanceolate to obovate, m. g, 1 e, et nar oa gined, clasping, petioliform se, entire or serrate, thick, pale green, sss rgined; middle stem-leaves linear-lanceolate t ovate or lanceolate, inekg and cordate-clasping, the upper reduced, mostly lanceolate, entire, those of the branchlets eaey educed, usu ually subulate and somewhat clasping; heads few to many, 1.3-2.5 cm sy ag mple corymbiform panicle; involucre about 5-seriate, strongly graduate “oy mm. aoe ee phyilaries ar erect, with white, indurate, l-nerved base and shorterx, rhombic o r lance-rhombic, her ous tip, cilio late; rays about 2 , blue oy ie about 7-10 mm. long; ae renes 4—5-nerved, pen © tally glabrous, the Pais usually reddis ually in moist soil, Arid Transition Zone; South Dakota to Colorado, eastern Washington, and Alberta. EA Be Socnhiy: “Valleys of the Northern Rocky Mountains to Idaho, south to Wyoming, &c.’’ Collected by Geyer. une— vpistineuished from the more eastern A. laevis L. only by the narrower and longer herbaceous tips of the phyllaries 8. Aster greatai Parish. Greata’s Aster. Fig, 5533. Aster greatai Parish, Bull. S. Calif. Acad. 1: 15. fg. 2. 1902. Stems 5-10 dm. high from a creeping rootstock, leafy, sparsely coarse- 7 sat alee gla- brous ho" reddish below. Leaves longer than the internodes, the lower elliptic, na petiolar base, deciduous at flowering time, the stem-leaves sessile, clasping to suba ulate. et ally subtending a small leafy bract, es ng- bores to din’ rough above, hispidulous-pilosulose — — — ae somewhat e above; usually bearing numerous heads 2-2. wide n bra oe a ao phy aries. Shiefly peas witht ually tees rhombic, uminate, sometimes loose, green tip, ciliolate, sometimes Hebidilous on back; rays 25-40, Tight he 5-10 mm. long; gor cit ubesc stk maces canyons, Upper Sonoran and Transition Zones; the San Gabriel Mountains, Los Angeles = San Bernardino Counties, southern California. Type locality: Eaton Canyon, San Gabriel cea eee California June—Oct 9. Aster jéssicae Piper. Pullman Aster. Fig. 5534. Aster jessicae Piper, Erythea 6: aes Aster latahensis Henderson, Contr. U.S. Nat. nace §: 201. 1899. Aster mollis Rydb. Bull. Torrey on 28: 22. 190 Stems erect from a stoloniferous rootstock, 0.6-1.5 m. high, densely cinereous-pilose with spreading to loosely ascending hairs, the numerous erect anlaan floriferous toward apex. Leaves aang ~ elliptic 0 r the upper oblong, 5-15 cm. long, 1.5-3 cm. wide, acuminate or acute, the ower winged clasping base, the others sessile and clasping by slightly or not iatilenieke pee entire of crenulate, often wavy-margined, firm, roughish-pubescent, or the hairs 1.5-2.5¢ beneath soft; hea m. wide, c — tar tips of branches and hier , usually short- peduncled or subsessile ; involc cre sometim r 2 equal o tiger eafy ‘bracts at bases sei fig iate, 6-10 mm. high, subequal ae ie io tly ‘graduate, we area linear to line or somewhat uehcedite. densely pilosulose, with whitish 1-ribbed base and usually shorter, erect, hi "i hacen tip, @ sf the outer wholly herbaceous ; rays about 20-28, violet or purple, 8-10 m ong ; achenes pubesc River sig Arid Eiger Zone; southeastern Washington and adjacent Idaho. Type locality: Pullman, Washington. .-Sept 10. Aster peepentee A. Gray. Marsh Aster. Fig. 5535. Aster Sarasa: of ag Not DC. 1836. Aster hesperi namie ae Fe 17: 192. 1884. Aster ensatut pote: hice 4: Aster foliaceus var. leit tlie. eins Fl. Pl. Calif. 1047. 1925. Stem 1-2 m. high with numerous ascending branches, pubescent in a above, very leafy. Leaves lance-linear, rathe? thick with midvein i below, 6-13 cm g, 3-15 mm. wide, attenuate, clasping, entire or the lower leaves serrate, chart ae ak ned; he ads Sait 2 cm. wide, eee very numerous, in rather narrow or spreading panicles ; involucre distinctly graduate m: m. high, sometimes subtended by a few hispid-ciliate, chiefly herbaceous, linear cts; the phyllaries linear, acuminate, ciliolate, — or loose, the hyaline margin of the inner phyllaries ssieidite to the tip or nearly so; rays about 25, white or purple, about 8 mm. long. Marshy meadows and along streams and i n ditches rt Son nd Transition Zones; Alberta and North Dakota south to Texas and cn to a hg og Hh of the be til poate ee from Mono County south and throughout southern California. Type locality: not definitely stated. Aug.—Oct. SN MZ SS 5532. Aster laevis 5533. Aster greatai 5534. Aster jessicae SUNFLOWER FAMILY $535. 5536. 5537. Aster hesperius Aster paludicola Aster foliaceus 317 318 COMPOSITAE 11. Aster paludicola Piper. Western Bog Aster. Fig. 5536, Aster paludicola Piper, Contr. U.S. Nat. Herb. 16: 210. 1913 Stem very slender, 1.5-8 dm. high or less, from slender rootstocks, usually pubescent in lines, with few or many branches ere. Leaves narrowly linear, 3-15 cm. long, 2-6 mm. wide, sessile, subelasping sane rpuern -ma ites more abundant at base of stems, reduced above; heads few h many, a avide. mose, solitary at tips of branches and divaricate branchlets evens bear appressed, Tanceolate feats bracts ; involucre about 5~7 mm. high, definitely graduate, the phyllaries erect, , acute o r acuminate or the outer sometimes ig) ee oblong and obtusish, the base whitish, the chin green tae lanceolate or pisabet ia lanceolate and usually fi Ss ea and -tipped; rays about 22-38, white, sic se a violet, or purple, about 8 mm. long; achenes bedi or pubescen Bogs, Can Sis u Mountains of satel Oregon and northern California. Type locality: Eight Dollar otis Tialptine jel Oregon. July—Se 12. Aster foliaceus Lindl. ex DC. Leafy Aster. Fig. 5537. Aster foliaceus Lindl. ex. DC. Prod. §: 228. 1836. Plants 20-50 dm. tall from a creeping épaay pac the stems (in ours) goon ng aig below. Lower 6s aves Samak te, 12-20 cm. long, 16-24 mm. wide, narrowed to a peti olar bas argins caure, ciliate-appressed, the stem- leaves sessile and conspicuously claspilte ¢ ; inflore soni -itieetty nocephalous, so as s subcym aring 4-6 heads, these up to 3.5 cm. wide, usually pubes- —s below the involucre ; iva | ection 9-12 mm hah: the a and inner subeq ual, additional foliaceous large bracts sometimes present, glabrous on the back, ciliate-margined, the outer en- larged and Pa nah oblong, obtuse, or broadly acute at the apex; rays 10-17 mm. i purple; enc’ glabrous or sparsely pubescent Damp places, Boreal Zone; Alaska ‘iciieaite British Columbia and Alberta Pegs: to the Olympic and Cascade Mountains, foliaceus: im the I to Montana. Type ag A Unalaska. July—Se large sense breaks up i many recognizable related entities although at points of contact these forms intergrade freely. In addition to ate extreme variability within the group itself, py veg ——e "3 pr esis ony ios A, nddakcatias and also with A. occidentalis. A ca to the varieties of 4. folt sis below based upon that of Arthur Cronquist (Amer. Mid]. Nat. 29: 429-468. 1943). Outer esac tane de ee broadly lanceolate to ovate, obtuse, or broadly ac nd onocephalo the was times bearing as many as 6 heads); Olympic and ‘thas Mountains of Weal Be: aor ioward var. foltaceus. Stems with sm heads in a cymose inflorescence. Stem-leaves Paya. Fei Lac dame plants of stream banks and meadows with the lower stem-leaves usually pers Phyllaries bal a om igad and acute or acuminate, the foliaceous ones if present linear or oer or yall long-po1 var. ee broader = ish more blunt, foliaceous ones if present mostly —,. “ oon to t iT} Stem- ive: ce Conapbedidaty late-clasping; plants of drier habitats with the tier ove “src mostly deciduous. var. canbyi. eT ar or narrowly lanceolate, markedly ac Oute Plants 1. 5-2. 5 dm. tall, often samen 9 (sometimes bearing 4—6 heads) ; in Caine habitats. var. apricus. Plants usually over 5 dm. tall; heads few to several in r le; not in alpine habitats ar. parryi. Aster foliaceus var. lyallii (A. Gray) Cronquist, Amer. Midl. Nat. 29: ip saatckes var. lyalliti_ A. Gray, Syn. Fl. N. Amer. 12: 195, 1884; A. hendersonii RacieP Bull, geet “Club, re 27%. tees A. eriocaulis Rydb. op. cit. 37: 14 Hee | kootenayi Nels br. Bot. Gaz. 56: 4 Much re- sembl v. cusickit but differing principally by the characters indicated in the key Subehdee mountains of eastern ho on nd, less commonly, adjacent British Columbia, Washi northeastern Oregon. ype ity € ootenai and Pend Oreille Lake, Idaho, Collected by Lya i “gi . 3 a | A. Gray, Proc. Amer. Acad. 16: 99. 1880.) Plants 6-10 dm. high; leaves rather thin, the lower leaves not oad edly exceeding the broadly clasping stem-leaves in length; outer Sho tiaries green and foliaceous. Southern British Columbia mostly east of the Cascade Mountains, ane southeastern Pg osc nal ay northeastern Oregon east to Montana. Type locality: small subalpine streams in the ntains 0 fe mn County, Ore hates foliaceus var. canbyi A, Gray, Syn. Fi. N. hee. 19: 193; 884, bast Ee fo Wace var. burkei A. Gray, loc. cit.; A. majusculus Greene, Fupeie. 4 : 215. 10005 A. tweedy i ie Bull. f pinot Cub 31: 655. 1904; <4. phyllodes Rydb. op. cit. 37: eee 910.) yee ut 5 dm. high; leaves rather thick, the lower not t much enlarged, and usually deciduous; the outer phy ilart ies much enlar ead poe usually obtuse. Usu ally growing W drier habitats than the last, eastern Sais and Oregon east to , pice and south to New Musing. Type locality: Whit River, Colorado. Collected by A pag eo var. oe A. Gray, loc. cit. (Aster apricus Rydb. Mem. N.Y. = a. 1: sigh 1900; A. incer 37: 20. ada a A 5 dm. or less high, cespitose, pubescent above, rootstoc ks Seanchedle head. “tending to be sol metimes subcymose. Alpine situations, southern eoBritish Columbia to nor’ alifornia ast ‘t ia st pone Co lorado. ‘heed locality: Union Pass, Color. é oh Eaton) A. Gray, Syn, F Amer. 17: 193. 1884. es SPE s var. parryi D, C. Eaton, Bot. ing Exai. 139, a7}; 4. Samplisimus Greene, Proc. Acad. Phila. 1895: 550. 1896; A. frondeus Greene, op. ci ye 1 egg ius G Pittonia 4: Spa A. ciliomarginatus Rydb. Mem: N.Y. Bot. Gard, 1: 392. i500, A, diabolicus Pipe: Bull. ‘Torrey Chub 28 645. 1902; A. vaccinus Piper, op. cit. pica ndivagear about 5 dm. high with ese usually persistent, lower leave and few stem- leaves; inflorescence ymbose, of few to several heads. In damp places, northern Wa: shington ‘aig pygmmonl ae to northern Cali- fornia and the central Sierra Nevada and east i 0 Colorado and New Mex: m. Type locality: ocky Mountains of Colorado. Collect ed by Parr 13. Aster subspicatus Nees. Douglas’ Aster. Fig. 5538. aw subspicatus Nees, Gen. & a Aster. 74. 1832. er douglasti Lindl. in Hook. Fl. Bor. Amer. 2: 11. 1834. a Sm oregonum Nutt. este. Amer. Phil Soc. IT. 7: 296. 1840. Not A. oreganus of authors. SUNFLOWER FAMILY 319 Aster elmeri Piper, Bull. Torrey Club 29: 645. 1902. Not A. elmeri Greene, 1891. Aster wattii Piper, loc. cit. Aster eto Re us Sheldon, op. = 30: 310, 1903 Aster ok anus Piper, Proc. Biol. Soc. Wash, 29: 101. 1916. Aster pio hl oe "Hear s, Otava Nat: Shs Aster grayt Suksd. Uieritnds ks 1927 Slender, about 1 m. aaa or less, with erect or ascending branches; stem asc in lines. Leaves rough-margined, practically glabrous, the lowest oblanceolate, narrowed to a winged petiolar base, serrate, the middle ones lanceolate, 43 cm. long, 1-2 cm. wie: slightty or not clasping, usually serrate above the middle; inflorescence cymose-panicled, heads ma 5 . high, graduate to subequal, loosely imbricate, the phyllaries pred e or the outer oa go with green, obtuse or acute, often loose, herbaceous tips; 20- "30, violet, about 1 cm. lon In moist places Hig? ie re a and stream banks, mostly in the Humid Transition Zone; coastal Alaska < Fa ao 8 south, ~ par west the Cascade Mountains, in hee erg, ee and Oregon and along ome n California from De i Norte Epon mz o Mo sini Coun’ ty; also occ rring less abunda: i east of the = Sacatatns. ral gi tet and e ag Ome Type locality: ton cit in insulis Multgrave,”’ Atike: une—Oct. ai — —— and apparently intergrading with other species such as A. foliosus and _ chilensis rs cis A, grayt Suksd., from Skamania and Klickitat ta ant Washington, and adjacent Oregon cca py soft pubescence; in the plants adjacent to the seashore ‘the outer phyllaries are petits oon spatulate with loose herbaceous tips. 14. Aster eatonii Sig Gray) Howell. Eaton’s Aster. Fig. 5539. Aster foliaceus var. eatoni A. Gray, Syn Zen guar: 12; 194. 1884. Aster eatoniit Howell, Fl. N.W. Amer. ae Aster oreganus of authors, not Nut Stem 1 m. high or less, escen usually in lines below the inflorescence, with numerous, erect or ascending branches. Leav beret! lance-linear or linear, rather thin, with the midvein prominent beneath, 5-13 cm. long, m. wide, acuminate, se essile with a so mewhat truncate base, usually entire, rough-mar ned, olten rough above; inact : 3c cm. wile. very numerous n an oblong te i n re qu 1 ate, 5-9 mm. igl aceous bracts, the hg or te thin, “3 anceolate or spatulate to (inner) linear, acute, often ciliolate, glabrous on back, the her ous tips of the sid oes longer than the pale base rad spreading ; rays about 20-35, Tavender or violet, about Moist ground along streams, mostly Arid Transitio = ee sou aoe British cles and Alberta Loe to California mostly east of the Cascade Tixatainn pre east to Idaho, Montan Wyoming south to Nevada and northern New Mexico; in Contes in the mountains a ee Siskiyou re: ils a a ity Hoare and ae unty south to Inyo Count ty. Type locality: Virginia Mountains, Nevada. Collected by Watson. July—Sept. 15. Ke hallii A. ~: Hall’s Aster. Fig. 5540. Aster hallii A. Gray, Syn. Fl. N. Am Aster Rieciae Sheldon, ce. Torrey Club 30: . 1903. A ster chilensis subsp. hallii Cronquist, Amer. Midl, Nat. 29: 462. 1943. Slender, ers 1 m, high or = = meal erect branches; stem sparsely pubescent all around, at least above ote, linear, 5-7 ¢ ong, 4-8 mm. wide, sessile, entire, rough-margined, depkity, elena abate. glabrous beneath ; seas small, 2 cm. wide, subracemose to cymose toward at ee fate AR " 5539, Aster eatonii 320 COMPOSITAE tips of the more or less leafy-bracted, ascending branches, forming a narrow gees sear r le about 4-6 m ou involucre more or less graduate, m. high, the phyllaries narrow-oblon the usually spatolate, erect, obtuse to neue ciliolate, glabrous on back, with whitish “cide or oval to subrhombic or oblong green tip, the outermost sometimes herbaceous and equaling the inner; rays about 25, white to violet, 5-7 m ie All — il or meadows, Transition Zones; Okanogan County, Washington, south to Wasco County, Oregon, and west of the Cascade dd he ins from the lower Pes River Valley, British Columbia, south to the Wil- lamette Valley, Oregon. Type locality: Oregon. July—-A 16. Aster adscéndens Lindl. Long-leaved Aster. Fig. 5541. Aster adscendens Lindl. in Hook. Fl. Bor. Amer. 2: 8. 1834. Stems slender, 0.3-0.6 m. high, pubescent all around or only in lines eight neeeciae: branched bove. Lower leaves narrowly oblanceolate, tapering to the petioliform base others baa to linear, all usually entire, often half-clasping, rough-margined, glabrou re ore ess pubes cent, thickish; heads about 2.5 c ide, few to many, in a near el or op r oa involucre 4-7 mm. high, usually strongly graduate, erect, the phyllaries linear or linear-oblong, the outermost posed spatulate, obtuse to acute or the inner acuminate, ciliolate, glabrous or Sodeucen t on back; s 22-35, violet or purple, about 8 mm. lon Moist or dry soil in various orn ad aah ie Pern and Canadian Zones; mihi Columbia and southeastern Wessnston, southward chiefly east of the Cascade Mo ee stele and the Sierr a Ne vada to the pag Bernardino and San Gabriel Mountains of southern California; Saska an south to Ut ae and Colorado. the west face of the mountains from Jackson County, Oregon, ‘to fae ‘County and southern California, it is represented by a densely leafy form —- a rather close cyme and glabrous or pubescent herbage. Type locality: “‘Banks of the Sas res wan.” July—Sept cage complex. aged Sc aphal up into ill-defined forms. All have strongly graduated involucres and a combine: "thd naked inflorescence with ascending branches but are highly variable as to calventaie ‘nk leaf-texture and ra abit 17. Aster occidentalis (Nutt.) Torr. & Gray. Western Mountain Aster. Fig. 5542. spathulatus Lindl. ex DC. Prod. 5: 231. 1836. Not sf Fy eaenscee Lag. 1832. o Tipo. site occidentale Nutt. Trans. Amer. Phil. Soc. II. 7: 1840. Aster o€ cidentalis Fo et Fl. N. Amer. 2: 164, ah Aster ] tit r. & Gray, op. cit. 503. 1843, ster durbrowi Eas S Gat Acad. ITI. 2: 292. 1902. Aster misellus Heaps oh “Biol . Soc. Wash. 33: 105. 1920. Stems usually 0.5 m. high or ine slender, arising a creeping rhizomes, oe dei below, usually pubescent in lines or all around above, at least toward tips of the rather few erectish branches. Lower leaves usually persisting: Shiances late fiieny-Obhanceolate ip g to petiolif ciliate base, entire or serrulate. gh-margined, thickish; middl pper leaves linear-lance ; de, entire, scarcely clasping ~ : : u in a nearly naked cym r cymose panicle ; involucre about m. high, not at all to slightly gradu- ate, the der he chiefly linear, acute or somewhat Gotusish, “Appitieed, ‘iliolate; rays about 30, violet or purple, 6~9 mm. long. Mountain aasilow s oa river thickets, Boreal Zone; Sie ee Lege = Tulare County in the Sierra Nevada, California, east to Colorado and Idaho. Type loc : “marg y ponds in the Rocky pe tains.”” ‘July- ept. ter occidentalis var. yosemitanus a oo. peas Amer. Midl. Nat. 29: 467. 1943. (Aster adscendens Mogh yosemitanus A. Gray, Syn. Fl. N. Am : 191. 1884; A. arg Greene, Leaflets —. Obs. Es 200. 6.) Stems slender, lealy to a phe Karol ‘leaves alike, thin, 2-3 cm. long, linear nag and gradually’ chien ha pede heads solitary or several, 1. g ‘2 em. broad, the —. linear. Reena Hy wig oosely emir 3 she adows, southern Oregon to the Siskiyou region, California, and south ingeg Nevada to Tula e Cou Type locality : Vernal Falls, Yosemite Valley. A variable aor aL defined “for Specimens are to - ‘fous which are evidently ae age with A. adscendens and also ental: ha ster occidentalis var. parishii (A. Gray) F. Madrofio 15: nds vinlit Oe er fremonti var. parishi A. Gray, Syn. Fl. N. Amer. 12: 192. 1884.) *Pisats heat 3 dm. or more tall, the ons leafy, the leaves aay clasping, — aang ag and reduced upward, a —_ leaves oblanceolate: or a shtirt cymose panicle; involucres 5-8 high, phyllaries sr sh i near, loosely imbricate, acute, Mou nv meadows of the southern California iltihes Phin to the San Pedro Martir in Epc California. Type Joca’ tag a Valley, San Bernardino beep a Very close to A. occidentalis var. occidentalis of the Sierra Nevada. Evidently aaa Re with the next Ast cidentalis var. delectabilis (H. M. Hall) Ferris, Madrofio 15: 128. 1959. re tone delectabilis H. M. Hall, Tints, Calif. Pub. Bot. 3: 82. 1907.) Plants 1.5—4 dm. high; stems iat the I pode narrowly cing 12-20 em. . long, the leaf-bases definitely daiplag? heads solita ary or few, the involucres large, 810 high, oes ‘phil laries linear, acu cute, loosely _imbricated. Mou ntain me i Al WS, southern Sierra Nevada i south to ae oars pee and gon Bern o Mountai pod. & the San Pedro Martir in Lower Californ Type ee Mi ll Creek, San Bernardino 3 Mou aka Calif ‘we Aster A. : Fi. N. Am u Contr. U. is. Nat. Herb. Pl ‘210. 1913.) Plants 3-5 dm. high, herbage “een to ieaacke diner eous per ulent inf 0-1 . lon n deci po the Fagont es ut muc Kickistet Comaties south along the eastern fac eo al = —— e ¢ and Plumas County, Californie. Occasional s; s from the Sie ge ada as far south as A fen ee pear te be this form. Type icalie: Falcon ¥ Valley, Klickitat County, Washington. Collected by Suksdor Pucther'’ studies will probably Fn that this sagged hould be removed from A. occidentalis from which it differs most markedly in its strong y predaet ge eer 18. Aster chilénsis Nees. Common California Aster. Fig. 5543. Aster chilensis Nees, Gen. & Sp. cai 123. 1832. Aster menziesii Lindl. in eik. Fl. Bor. Amer. 2: 12. 1834. SUNFLOWER FAMILY 321 Aster chamissonis A. Gra “i shea Exp. 341 74, py militaris ciioe Deu. Acad. Phila. 1895: 550. 1896. Erect, 0.5-1 m. high, pascal branched above; stem pubescent all around varying to nearly Bieoat takes leaves often he ere narrowe ee oo > somewhat eer rR! base, 12 cm. long 3.5 cm. wide, the middle ones mostly lanceolate, 4-9 c g, 0.5-2 cm. wi ntire or serrate: rough-margined, aften rough above, essentially elabrows| henéatii: inflorescence racemose -paniculate, sometimes a simple raceme; he ae usually 2-2.5 cm. wide, borne on leafy -bracteate branches; involucre 5-7 mm. h, 4-5- seriste ahe. yee llaries chats and clo eae graduate (or the outer rarely herbaceous and Shatin the inner in intermediate fo rsh: ised — hi oblong, sometimes slightly broadened hares with whitish, l-nerved base and shorter, somew rhombic, obtuse to acute, mucronulate, narrowly pale-margined, hi oy pees tip, ditotate elabious on back; rays about 20-30, violet mace or white, 6-12 mm. lon Usually in moist soil, Humid Tiswaitien and ober Soule. aie: Sed the coast from northern Oregon uth to Santa Bar aie County and Santa Rosa Island, California. Type locality: California, erroneously pub- lished as Chile. June~ er chilensis sabe sp. . chilensis as identified by the earlier botanists from the collections of Haenke, Chamiss Mewvices and Hinds is the coastal form described above in which the rp stem- leaves are oblong- spatulate ‘es lanceolate with the pete ne entire or rire cg ntate, sessile to somewhat clasping, and ~~ ubescence of the stems and to a lesser degree the leaves varying from rather coarsely canescent to almost glabrous. It socaka on the Oregon coast sine etl to Santa Barbers, California, eaiieniy more abundant from Humboldt County south to Monterey a along the coast = 0 pebnee — Ranges than at the extremes of the range. The occurrence of Aster subspicatus, where its mor therly coastal distribution coincides with that of cesaragr may account for the ease ence of the \coublesede Yor whith are evidently hybrids. Other segregates of this variable taxon are —— locally but intergrading forms are frequently found from the Oregon border to Tulare C ; Califor Aster chilensis var. The tus (Greene) Jepson, Man. Pl. Calif. 1047. 1925. (Aster lentus Gree Man Bay Reg. 180. 1894.) Plants I m. or more in height, nena or nearly so; leaves linear-lanceolate; leinceisknee ample, widely branching, the branches conspicuously leafy-bracted, the flowers large. Suisun marshes, Solano occidentalis 5540. Aster hallii 5542. Aster 5541. Aster adscendens 5543. Aster chilensis 322 COMPOSITAE County, California, the type locality ; also, specimens approaching this form have been collected in salt marshes along the coast in Humboldt and M ster chilensis var. sonoménsis fiteciens} loses, loc. cit. (Aster sonomensis Greene, Man. Bay Reg. 180. 1894.) Plants 3 3.5 dm. high, acd eri rather vg sens ion leaves oblong, the stem-leaves reduced, lanceo- late; inflorescence a cymose, leafy-bract anicle, the heads and solitary at the ends of the branches. Marshes at northern nd of San Wosmciaen i. Colitemta, Sonoma pect Napa Counties, and in similar situations at the southern end in Santa oul our ae peal: “open plains of the Sonoma Valley. Aster chile fee loc. Plants 1 high, glabr arly s y leafy, the leaves eal: acetate or gt ape danceotate, ity spreading; inflorescence evith Welety divacscate. pat Bee atl the leafy bracts ‘somewhat a than so nit leaves, the heads few. Along watercourses, a eg of er and Tuolumne Counties, and in Sac and San Joaquin Counties, California. Type ocality: lower Sac paces os a ae er chilensis var. invenistus (Greene) peieatt, Ae cit. (Aster jacentstes Greene, Man. Bay Reg. 179. 1894)" "Plants stout, Pa to 6 _ high, herbage scabro short-hirsute; lea anceolate, the lower stem-leaves ——— tulat -~7 cm. lo ong; infloveacente deena ewes ed in an rant pr panicle. Described by approximate this form bat vary in pubesc scence from n arly g jabrous to short-hirsute and have few to many heads in the inflorescenc hav te gee coll goto d at various loca lities in the Taner Coast Ranges from Siskiyou County south to Sonoma Connty, and the Sierra Nevada foothills as far south as Tulare County. 19. Aster bernardinus H. M. Hall. San Bernardino Aster. Fig. 5544. Aster bernardinus H. M. Hall, Univ. Calif. Pub. Bot. 3: 79. 1907 syeet deserticola J. F. Macbride, Contr. Gray Herb. No. 56: 36. 1918. Erect perennial 3-10 dm. high, densely cinereous throughout, rarely somewhat glabrate, stems 1 to seseral from a woody root, densely leafy, the internodes short, simple below, the leaves early- deciduous, branched above. Leaves 3 cm. long, mm. wide, sessile, Meas to ‘linear-lanceolate ; onl * Boe tl short-branched, leafy- bracted racemes or an elong: ated narrow pan icle with the heads congested on the branchlets; involucres 5-6 igh; phyllaries strongly celeaee. pihorins otters to point glabrate, phish n-tipped, with the argins white-chartaceous on ciliate, the outermos engi oe _ or rounded, the inner scutes rays 30-35, purple, 6-10 m long ; achenes canescent, the ondid.! oa me and drainage aie os er Sonoran Zone; Los Angeles and San Bernardino Counties south San a “ awoag OB rma , reariig Re a ge _ ‘the oss cat rt | in Tes cen and San Beroardins Counties. Type 1 vic Ber iy dane safiatgians Parish, Bot. Baz. ny ps "1904, Slendet rigid, sh ict te 3 Na erennial about high; Jealiens below at flow we i igh os the branchlets of the in Hp ae pio clothed with linear acute eaflets about 1 cm. long and longe internodes; branchlets widely divaricate, . long, the heads wolitar +7; soaseidines 2: Hy the tips of Aa: ice, phyllaries strongly graduate, ‘eoalll? aoe aiins rgined below; pappus copious. Known only from the type Sag 5 5336) collected in dam p meadows, anent 900 feet altitude, San Bernardino Valley, San Bernardino County, California, collected October 37573903. 20. Aster falcatus var. crassulus (Rydb.) Cronquist. Little Gray Aster. Fig. 5545. Aster adsurgens Greene, Pittonia 4: 216. Aster ihe as Rydb. Bull. Torrey — - 304. eo Aster ssulus Blak . one USN b. 25: 560. 1925: Aster falcatus var. crassulus Pc Ball. ices a 74: 144. 1947. Stems usually 0.5 m. high or less with well-de stats — jo es, often much branched at resse Leaves linear, 3-6 cm. lon mm. wide, firm, en vitae: cuspidu rity ile. subclasping, usually hes much reduced ads 1-1.5 r fe) high, cealigite: the phsares Pla cblong i seat 6 r the outer usually spatulate, with a strongly indurate, whitish base and a rhombic (or e inner paves io Be thick-herbaceous, mucronulate, 0 acute, tees loose ds hispidous: = usually pubescent dorsally; rays 5 22-30, white or ard Volek ti 3g 5-8 mm. long. Dry open places, Arid Bo iice Zone; eastern Washington and Oregon east to Saskatchewan and Colorado south to Utah, bere and New Mexico. ey locality: La Veta, Colorado. Aug.—Oct. 21. Aster pansus (Blake) Cronquist. Heath-like Aster. Fig. 5546. — rag var. pansus Blake, Rhodora vag et Me ster onquist, Leaflets West. Bot. 6: 45. Pesci about 0.6 m. high, i saloedt ew from a short rhizome or woody caudex; stems clustered, strigose or in ours mostly with spreading-hirsute or hispidulous hairs. Leaves deciduous below, linear, 2-7 cm. long, 1-6 mm. wide, obtus caine bag vi apiculate, erg rigid, rough-margined, strigillose, those of the ececher crowded, abruptly reduced; heads small, usually nume d racemosely arranged on the arching branches on sn a minutely spreading-bracted branchlets, the disk 4-6 mm. high; involucre about 4 mm. high, graduate, the outermost phyllaries usually squar- rose and — the others mostly linear, with indurate whitish base ior abrupt, more or less tip allo wn —_— — Tr , olten ' mucr. omlate; rays 12-20, iess commonly si api = rs mm. ig achenes pu * Usually in dry open places, Arid Transition ritish Columbia south to eastern Washi ale ortheastern Oregon and ener poet 0 s Calerada ay Wines. Type locality: Ellens- burg, frat tosgalg Aug.—Oct. Aster columbianus U.S. Herb. 210. 1913. (Aster multiflorus var. columbianus Blake, Rhodora 30: 227, nd ) en rake been eacoued by Cronguist that this rarely occurring, violet- rayed i —- of hybrid origin, having been derived from A. compestris and A. pansus. Type locality: Waitsburg, Walla Wa’ Ma SUNFLOWER FAMILY 323 22. Aster brickellioides Greene. Brickellbush Aster or Rayless Leafy Aster. Fig. 5547 Sericocarpus tomentellus eg Sere 1: 283. 1889. Not Aster tomentellus Hook. & Arn. 1833. Aster brickellioides Greene, op. = rage Fabien on Greene op. cit Eas oj Celis a Bi po 157.3931. eevetth 6-9 dm a me WE tomentose or tomentulose, glabrescent, leafy, sometimes much branched. Leaves mostly spreading or deflexed, oval or elliptic ao to linear-oblong, scm. ng, 1-2 cm. se ki acute, the broader leaves pedacire apiculate, roun at z e sessile ye soni sometimes narrowly hue dered A subcoriaceous, glabrous or ata: rcbawe canescent~ or cinereous-tomentose and sometimes glabrescent in age, seit Ceaetia’y pi of the Sanches greatly reduced, mostly linear-subulate or linear-lanceolate; heads numerous, cymose-clustered at tips of branches in a usually elongate-oblong scnpuang involucre turbinate, about 6-seriate, strongly graduate, 7-9 mm. high, the phyllaries linear or ovate to Py eetagtans or oblong, acute or the inner obtusish, with indurate-chartaceous, crt margined base, the short poishige easy tips of the outer spreading, the tips of the inner erect, not herbaceous, pilose- ciliate at least above, i dorsally, sometimes glabrescent, the inner sometimes purplis Taye : light violet, ut 6-8 mm. long; a es compre see, 4 sparsely pilose to essentially glabrous; pappus tawny, the inner pappus- -bristles somewhat fattened apicatly, the outermost short, setul Dry ridges and rocky slopes. Arid Transition Zone; poche, ahr ir southwestern jem and adjacent California. Type locality: near Waldo, Josephine County, Oregon. July~Oct. 5544 5546. Aster pi 5547 . Aster bernardinus 5545. Aster falcatus . Aster it ER 324 COMPOSITAE 23. Aster siskiyouénsis Nels. & Macbr. Siskiyou Rayless Aster. Fig. 5548. Aster brickellioides var. glabratus Greene, Pittonia 2: 17. 1889. Eucephalus glabratus Greene, op. cit. 3: 56. rigiveg Ast. ak SURE TOMENAE NE. & Machbr. Bot. Gaz. 56: 477. 1913. E Eastw. Proc. Calif. Ac ads EV. 2025157.. 1931. Aster glabratus Blake ex M. E. Peck, Man. Pl. Oregon 726. 1941. Not Kuntze, 1891. ms 3-6 dm. high, simple, reddish below, arising from a woody, creeping, branched rhizome, sistally ne brous or somewhat g a especially beneath the involucres. Lower leaves reduced and scale-like, the others nearly u m, somewhat smaller toward the inflorescence, 3-6 cm. long, ovate-lanceolate to oblong, natgols eee glabrous and so metimes roughish- -puberulent beneath, occasionally thinly pilosulose in intergrading forms; heads few to several in a leafy- bracted, corymbifo panicle, the pedicels slender and spreading, the involucre campanulate, 4-5-seriate, and strongly graduate, 8-9 mm. high ; outermost a ries loose, lanceolate, and green, the inner broader, indurated, pale ie the tip greenish, the margin. scarious, ciliate a t least above, rarely tinged with purple, glabrou on the back Ne einetinied minutely 7 landular ; rays absent, rarely 1 or 2; achenes siiesintt uakiae: pappus tawny. Dry forested slopes, Arid Transition and lower Canadian Zones; Siskiyou Mountain n area, Josephine and Jackson Counties, Orezon, ss ta (sto Siskiyou County, California. Type locality: ‘“‘toward the summits of the Siskiyou Mounta Ore g.—Sept. e marked sree Ds ° “his species saeeeste. bee hybridization may oceur between A. siskiyouensis and the other species of the Eucephalu 24. Aster vialis (Bradshaw) Blake. Wayside Aster. Fig. 5549. Eucephalus vialis Bradshaw, a 20: 122. 1921. Aster vialis Blake, Rhodora 30: 228. 1928. Sericocarpus sipei Henderson, Madrofio 2: 105. 1933. Stems about 1.2 m high, simple below the inflorescence, leafy, densely stipitate-glandular especially above and sparsely pilose. Lowest leaves minute, the others gradually reduced above, ipti i cm. long, 0.8-2.3 cm. wide, acute or subacu at este eee tire or the larger rarely with a few shar rp teeth, rather firm, green, pe obscurely "etindula above beneath dull, tipitate- “glandular, eh sparsely pilose, 1-nerved, co with a pai basal v - heads discoid, 1-1.2 cm. hi gh, several to numerous, cymose rot in a narrow, eine. leafy soir ie the peduncles short ; in estate carbine to peodbee abet and sometimes the upper stem jeneely | glandular, - not or scarcely floccose tinesiuers and peduncle just below the head glandular only in C. fil poo turbinate (broadly so in C. filaginifolia bevnevdina): phyllaries squarr y the involucre and upper peduncle glandular; plants of San harcardina Valley. eres C. filaginifolia bernardina. inflorescence sisndalar; plants not a restricted. A ut 1] A + Stem slender; i hern seacoast aginifolia virg Stem 7 inflorescence glomerate or a short-branched raceme or paviles os 9 . inland moun- tai C. filaginifolia glomer I ity } ne emispheric; phy] i ly or not at all Squarrose. Erect plants from a woody root, the infl many-flowered ; rag California coast. C. filaginifolia rigida. Decumbent suffruticose plants, the inflorescence few-flowered; Channel Islands. . filagi inifolia robusta. Corethrogyne filaginifolia var. hamiltonensis Keck (Aliso 4: 104. 1958), a Plant hg - habit and inflores- cence of C. filaginifolia var. filaginifolia, has involucres obconic, 7.5—11 — igh, iate, the phyllaries appressed, damnmetat tomentose at patios and recess if at all glandular. snd on me western side of the Mount Hamilton Range in Santa —- and San Benito Counties and poi Logout with the posh ase" exten- rowel C, filaginifolia var. filaginifolia, and was described from material collected on Mount Hamilton (Hall 8 a Coreothrogyne fila: prifctie: var. rigida A. Gray, Syn. Fl. N. Amer, 1°: 170, “ow as to es (?) Corethro- ane viscidula Greene, Fl. Fran. 378. a ?C. viscidula var. greenet Jens, 3) C Ries ape oy Sal rigt 6. eaberain | ; C. filag ir. UES 2.5-5 dm. high with stems stouter than C - laginifolie var. filaginifolia, peal or decuml nt = te pais oe ss udin * narrowly or broadly. pblaieeaiaie. acu and usually toothed on the upper third; heads abe! saa several; pesdetaied 5-6(7) mm. high, hemispheric = passer turbinate, densely glandular (except in intermediate forms) and often snitele gtananlec the phyllaries — a, 3-4(5)- perieta: not squarrose. Sand “hills and bluffs g the n and open hill Pant spre 30 to asi “Alam a County south to Monterey County, California. below and Cawiaie ae ove, with an et , many-flowered kere: scence. Corethrogyne ‘laine var. rigida differs from C. filaginifolia var. glom with hick ‘t som ae Hosa been identified, co hone eres a panulate to hemispheric involucres, Ce ohcics which are eddies 4 S cme ~ : ore ope inflorescence with lon pig x Petite ulate branches. Corethrogyne riscidula var. erocuri Jepson an Specimens ann aided by Geckae s C. viscidula differ considera abi from C. filagin rpbdiet var. rigida, Further investigation may prove that these plants ge have large ean with distin nctly ie ric involucres and rather broad Pade represent . pop taxon. The large heads and the habitat in which they gr “os suggest ata epic with the rarely collected . califor aa Co orethrogyne filaginifolia var. robista Greene, gy 1: 89. 1887. she stout, ascending, densely c s s co 8 upper zi 2 of the same shape; inflorescence loosely aniculate or corymbiform; heads rather few, about as broad as high in pressed specimens; phiyllaries ‘glandular with a broad green tip. tf sector San — Santa Rosa, and oe Cruz I = a anta Barbara = y, and apparently. banat t Be al, oe not in the typical form, along the coast of Sa sob ispo sad: Santa Bar oot mage es ene A Sun iguel Island. Ha Fein some characteristics in common C. filaginifolia var. rigida but differing from that taxon in rowth form. : Corethrogyne Soccoss Greene (Leaflets Bot. Obs. 2: 25. 10), described from material collected by Alice Eastwood in September at Elwood near — Ph reg is aa in “habit, oh slender stems and filiform, glandular, pedunculiform aeonetee. The hs oH es are — —— or oblong and somewhat toothed. It is possibly a hybrid between the above and laginifolia var. virgata. ak hro. ae Mineiier ris kor Hod cit. 26); aeacribed from materia d by Braunton in ay Redondo, Los Angeles County, and also collected by Abrams and Braunton “g Daye t del Rey, has white-tom linear a ee yf Sasi the habit of C: * fleginifolia var synnogy i.e aed in older pl ants, and gine ei large hemispheric nvolucre s which resemble those filag nifolic var. Cirniladaa ‘fila aginifolia var. latifélia H. M. ties Unie. Cal Pub. eer 3: 70. aig — woody stems, the leaves and stems and sk wali tomentose ep not at all sep ‘leaves 1-4 cm wide, sessile, mostly oblong, sometimes narrowed at t e. somewhat toothed toured the r ot apex: heads mener See, few, ad . at the tips and not glandular. Occa cat along the coast of Santa gy ee ar an entura Coumates, Californ and apparently ered freely nn the Caan forms in he same area. Type locality: Oxnard, polar Coun h fil ifoli virgata (Benth.) A. Gray. Bot. Calif. 1: 321. 1876. (Corethrogyne virgata ne rethrogyne rary eg Crane nica q virgata Kuntze, Rev. Gen. Pl. 1: 330. 1891; C. lavandulacea 342 COMPOSITAE Greene, Leaflets Bot. Obs. a: 27. 1910.) Tall, slender-stemmed plants 6-10 dm. high, widely paniculate-branched above; heads many, borne short, divaricate, pedunculate branchlets; lower stems loosely floccose, rather leafy below, the rng he: ne ay. lacking on the upper stem and inflorescence, these short-stipitate-glandular; lower leaves as in elgg vinifolia ook Sapna the upper much, reduced, bracteate; involucre small, turbinate, the phyllaries closely , squarrose, cartilaginous, marked at the tip with a well-defined, green or brownish green spot. Oc hal ti yi os "near ihe coast from Santa Barabara County, California, to coastal northern Lower California. Type locality: San Pedro, Los Angeles County. Collected by Hinds. Intermediate forms between t iserch — and other forms of the C. filaginifolia complex are not infrequently tound in cismontane aarlace Califor Corethrogyne poymrase var. a aN Canby (Bull. S. Calif. Acad. 26: 14. 1927), a local form seemingly related to the above C. filagin ge Lane var. Seas te found growing in the vicini ay Bed ghar — ae County. The. sl = pide Me ted are stout, have a many-flowered inflorescence, and h volucres which a 6-8-seriate. Lorttiveg ine scabra Greene (Leaflets aie Obs. 2: 25. 1910), another local species gain by Hasse in Lael at Los Angeles, is sae temmed, leafy to the few-flowered inflorescence, and has green and scabrous erbage. Corethrogyne filaginifolia var. —— a H. M. Hall, Univ. Calif. Pub. Bot. 3:72. 1907. bpd "4 Seeudtule —— Leaflets Bot. Obs. 26219105 .C, — osa Greene, loc. cit.; filaginifolia var. pinetorum I, M. Johnston, Bull. S. Calif. Acad. 18: a1. 1919; C. fill aginifolia var. brevicula Canby, op. cit. 26; 12. 1927; °C. " plagiad a2 hea rigida A. Gray, in part; C. filaginifotia bso g irl la Gray, of authors, not Torr. & Gray.) Stems A dm. high, stout or more der, r divaricately ascending, persistently leafy at and near the Se scent base a 0.5-3.5 ¢ = eats, png Hel ly white-tomentose; heads ase or yor so in the upper leaf. walle or few oT. in one er brite menos or guetta ie involucres 7-9 mm. high, turbinate; cat Upper ac Type locality: Oak ie Yucaipe Ranch, near Redlands, San Bernardino County. Intermediate forms bet this | taxon and C. filigintfolia var. virgata de> rn folia.va found in southern California. n the ic agin Th of C. filagintfolia var. tener ap shoneh. short in stature, aaa pone di ost pacity saan eo taller plants - the north, Consthrogyne filaginifolt SE gmig ses: perhaps an ecological form of higher peti oy has few heads and a more naked in Sdortieence as ad tes s from the southern Sierra aig tee to Fong tow r California border Corethrogyne filaginifolia var. senucaias (Abrams) H. M. Hall, Univ. Calif. Pub. Bot. 3: 71. 1907. “(Co sellrtane virgata var. bernardina Abrams, Fl. Los Ang. 401. Toee Erect plants 6-8 dm. high, with h heads rather few on metodo: branches; stem ee? white- tomentose eons sme for the glandular involucres and the upper part of the peduncles; ap 5-7 Pore dne the phyllaries squarrose. Consett on ay phe ns especially in the San Bereardinc Valley. Type locali ee tone, San Bernardino County. The densely white-tomentose stems and somewhat larger heads suggest Sisal with C. sessilis. 91. ERIGERON* L. Sp. Pl. 863. 1753. Annual, biennial, or perennial herbs with alternate (or sometimes all basal) leaves. Heads solitary to numerous, hemispheric to turbinate, usually radiate, the few to usually more or less numerous pistillate fect: usually bearing evident and often narrow rays, : ~e : s na ing from herbaceous and equal to scarcely herbaceous and evidently imbricate, the loss of herbaceousness either emgplaes triple: bog their length or more prominent toward the tip. 1 Receptacle flat or a little c k Disk-flowers more or less numerous, generally yellow ; some species with pelos pistillate flowers between disk-flower the ray- flowers. Anthers entire or nearly so at the base. Style-branches flatten , with introrsely s marginal stigmatic lines and short (up to 0.5 mm.), externally minutely hair , lanceolate and acute to more often broadly triangular and obtuse appendages, or the appendage rarely (E. annuus) obsolete. Achenes 2- to man ny-nerved ; pappus of capillary and often fragile bristles, with or without a s Bork —_ series of minute bristles or scales. [Name e Greek, meaning early old man, or old man in the spring, presumably referring to the early flower- ing and fruiting of many of the ani s.] A genus of — 200 species, of North and South America, Euro i Asia, nearly all of temperate or boreal regions, eas in tropical America. Type species, niflorus L. Pistillate corollas very numerous, filiform, with very narrow, short, erect rays, these sometimes not exceeding the disk (inner pistillate corollas some times — bs VY. TRIMORPHAEA Pistillate corollas few to numerous (or absent), the tube generally einen rays adi devel- oped and spreading or sometimes eee a rahenae but not short, narrow, and e Pappus of the ray- and disk-flowers unlike, that of the disk-flowers composed by bristles and short outer setae, that of the rave labetts lacking the bristles; weedy, mostly annual plant TV. PHALACRO Pappus of the ray- and disk-flowers alike, of bristles, sometimes also with outer setae or scales; iy mostly —— nnial, a few species biennial or casually annual, seldom weedy except for E. philadelphicus and ergens. Internodes very and short; leaves linear or narrowly oblong, te a uniform from the base to aa the { ron Gf: the plant. the basal ones, if present, not markedly larg venacpet cauline ones; hyllaries markedly imbric TIL. PyYcNOPHYLLUM. Internodes an excessively numerous nor pond very short; leaves var reed shaped, “sometimes linear, but then the basal on usly lar; than the cauline ones; phyllar Aan or imbrica Achenes 4-8(10) cosas tae and foothill plants with the shyitaricy’ reeacatl? imbriate io the leaves si silvery-strigose. Il. Wyomin * Text contributed by Arthur John Cronquist. SUNFLOWER FAMILY 343 Achenes in most species pe persed few species that sometimes or regularty have more pegged nerves occur in m Woutland: or maritime habitats, have the oe ——— < early so, and are not at all sivicy primes, . EvERIGERON Plants maritime and submaritime; stems — curved-ascending; heads large, hemispheric, the disk 14-35 mm. wide, the disk-corollas 4.7~ 7 mm. Ion Heads radiate, the rays about 9-15 mm. long; cauline leaves ample in reer specimens; oo and California. Dy glaucus. Heads discoid, the flowers all tubular and perfect; cauline leaves narrow, mi tly lance-linear so narrowly oblong, or the lower somewhat oblanceolate; Humboldt and Mendocino Counties, pra ies 9; Plants cei Panam or submaritime | (save perhaps occasionally in E. sanctarum?); tah various tot mostly except in Cauline leaves ample, usually lasicnolate or broader, entire or toothed, never trilobed; phyllaries equal or subequal; achenes 2—7-nerve Seu at least when well developed: tall and erect (up to nearly 1 m, tall), somewhat Aster-like; achenes 2-7-nérved. Rays ee ely broad, y 2-4 wide hes mm, in E, coulteri, which ee the hairs of the volucre with Bora jo thy near the base); pappus simple or near Hairs - = ine lounee black accel rays mostly 2-4 mm. sc as whlté or colored; airy or gla Aubelbes 4—7-nerved; ledivee glabrous or occasionally hairy; involucre ‘mostly merely glandu- lar “oh sometimes hirsute or with the bracts more or less ciliate-margined but not hirsuite below and glandu a abov ve; disk-corollas mostly 4-6 m a ioe naa head) blades usually t to the petiole; cauline "tt various but rarel a thin and proc “clasping; rays beeen ny 6 sometimes white; widaeprend, t moderate to high elevations in the moun . E. peregrinus Basal leaf-blades abruptly contracted to the petiole; cauline teesak thin, conspicuously clasping; rays white; south side of Columbia River cm ee in Or eenen panne. 2—4-nerved; leaves hirsute; involucre hirsute below, glandular an, a t corollas ostly 3-4 mm. aaa: Olympic, Oregon Cascade, and Klamath ahh eaioes.. ceae. Hairs of ey involucre with black cross-walls near the base; rays wide, white; leaves hai achenes 2-nerved; mountains of northeastern etal Pe California’ and eastward. mm. wide or less; hairs of the lab gg if seenrnh: ‘witheas ‘black cross- Rays poly about walls; pappus, Pe Ta in E. phila delphi cus, mostly 1 Rays sad excessively numerous, commonly about 75-150, about 1 mm. wide; disk-corollas mostly 4-5 mm. long; achenes 2-4- nerved; true perennia als. Leaves mea or nearly so except for the ciliate gna stem Lago ag — the in- florescence or merely bearing a few scattered hairs; involucr ive: few or no long airs; widespread. . hai Leaves, stem, and involucre more or less long-hairy; Washington and ‘anaee : i: es Seren Rays very numerous, commonly 150-350 or more about 0.2-0.6 wide; di corollas mostly 2.5~3. ca mm, oat achenes 2. neecedi gv Be or dhs rere perennial; widespread, somewhat weedy. 7. E. philadelphicus. Plants low and often haete ae or ascending 0.5-3 dm. tall, scarcely Aster-like; achenes 2-nerved. e or nearly so; pappus-bristles straight. nd stem glabrous or feiatinihe above) somewhat pa sometimes with a few x eet ry a ag glutinous to glandular, sometimes with a few long hairs; Oregon to ern Calif th as distributed, the middle cauline about as e as the lower, — upper Leaves ‘con ara ably distrib Del Norte County, California, peek id adjacent Or aha Leaves inequably distributed, the basal ones obviously larger ae the few and wien es reduced cauline ones. urple; disk-corollas usually more or less puberulent at least below th Rays po middle 7 nvolucre ae glandular = not at all hairy; stem ordinarily white tabroes except directly under the ana where ep cnaceel Siskiyou Mountains of northwestern California cal sue t Orego 12. EB. cervinus. te. R all hite; disk-corollas essentially glabrous; —— usually with a few 1 "bal ro ordinarily with a few hairs or glands o = es Cascade and Klamath pec s of Oregon but more northern or eastern than . cervinus, . &. cascadensis, Leaves, involucre, and usually the stem with obvious long hairs as oe as stipitate-glandular; rays blue or purple; Cascade Mountains of Washington. 1“. re git. led for Basal leaves coarsely toothed or incised; pappus-bristles characteristically twisted and cur at least the upper half; Columbia River Gorge in Oregon and Washington. 15. E. oreganus. 11 developed, commonly linear or oblan ceolate, sometimes linear-oblon ee narrowly lance oblong (sometimes yi ag in E. caespitosus, which rete Ss an a, imbricate i rer ong in E. basalticus, which has the leaves ev mig tri prin at those of : eens OSHS, ane a ol other species, while narrow, somett not muc aller than the aca oe ones; gp Ses rved. ’ mally 2-n Leaves, or many of them, trilobed or two to four times ; Stem fat, leafy, the cauline leaves more or ite gre trilobed; ier ag oe Stem scapose or subscapose; the cauline leaves if present few and much reduced, atl 5 Sader and entire. * The perhaps artificial section oer consisting of short-lived, mostly biennial species, is here for convenience submerged in Exeriger 344 COMPOSITAE Caudex divided into several or many long, slender, rhizome- re branches; leaves mostly merely + Sa se lobes short, iced, rounded, and obtuse; Sierra Nevada, Wallowa Moun- tai and eastward (absent from the Cascade Mounta ay, . vagus Caudex stout, occasionally branching, but the branches mostly relatively cigte 2 age “aieialty short, not slender ane peat (caudex pico alvit that of E. vagus in so lants from the ascade Mou of Washington) ; leaves bd or more often two to four times eee stly with walatively slender lobed or idespre 18. E. compositus Leaves all entire or heat os so, not at all trilobed. Involucre woolly eltisihe with sy pene a alpine and subalpine plants with solitary heads, always r: adi ate; Oregon and Washingto Rays yellow; Cascade Mountains of Wastinetac and northward. 19, E. aureus. Rays si or pink to white, not yel Rays about 50-125, pink or ei aceenaaten Oregon and eastward. 20. E. simplex. Rays about 25-50, white; northwestern Washington. E. flett senieree variously hairy or glandular = Lanna but not woolly-villous; ssa ae to disciform r discoid; habit and distribution ibnacanee of the —- widely seogneritog™ etimes scanty (or lecgely. ae ying in three species; of these, E. multiceps has the pubescence Hetuictey sprea under eagle . es perianus and E. engelmanniit have conspicuous, coarse, et Rog hairs on the ns or petioles of at iast the lower leaves). Alpine _ s ne plants with solitars: or rarely 2 meade. always radiate; California. with several more or less leaves wpe ie or glandular as well as hirsute; phyllaries pena equal; taproot poorly r scarcely developed. aes petiolaris. Stem os om or glandular; phyllaries more or less ao = well de- velop 23. E. clo ke yi Plants scapose or very nea Basal leaves loon toe or linear-oblanceolate, up to 2.5 wide, ~— tape ing to oe he ole; rays blue or purple; disk corollas - oe 5.3 mm 1 E. py on gm Basal Lod Mates subrotund to broadly oblanceolate, rather abruptly contracted ud rter than the petiole, — etimes as much as 8 m eaoct ee Ms Pwidtcs disk-corollas 2.7—-3.4 me long; oth eae cant ei i a. 25, E. unctalis. Plants chiefly of the valleys, foothills, and moderate elevations in the dédntains, the Meth species E. aphanactis ascending to rather high Geevatieke in southern Cali- chenes peittt perenaas the hairs completely covering the surface of the achenes and sometimes obscuring the outer pappus; heads radiate, with Srak or purple to deep blue or Wisle rays, or rarely disciform, with the pistillate flowers rayless; pie or = southern British Columbia and ns. 27. E. poliospermus. Achenes sparsely or paar hairy, the surface ex posed ed mane the hairs, the outer pappus if any not obscured by the hairs; rays vario Rays blue or purple to ok or white, or wanting, but not stews Heads recat radia ‘appus simple or aay so; disk-corollas 5-6 mm. long; Santa Bar- hark and San Luis Obi ispo Counties, agate ' . sanctarum. Pappus he Seni tee — disk-corollas cae 3-5 mm. Seay more Sie cpeoet ors ead except in E. multiceps 3-5 mm. long; ys and pappus various. Reshma of the st L li J aiges ys mo: ahite, turning pink or pinkis porn ess lively hear with short broad fas and heavy limb, he lower part tary; near nake River ce eastern Oregon, 3 southeastern, Bal 5 ngton, and adja- t Idaho. dispartpilus. Rays aero 0.7-1.5 mm. a ored or occa col ally white; aa calles relatively slender, with slender, comparatively and narrow limb, the lower part of So hich “a peo ngly in- durated; heads 1 or usually more; — read. 29. fe “ii umilus tse pappusristes about 12-20; 5, dskecorollas ab _ sis Inner pappus-bristles capri diate corolag oa +2 3 mm. long; rays less wide; pubescence of the stem ae pore the heads; pubes- cence of the Paid wholly appressed; southern Sierra Neva 31. E. multiceps. Hieeninis gtisk-comatins 2-3 mm. long; rays about 75-150, 0.5 mm. wide; inner gern s-brii as ig lee widespread. SUNFLOWER FAMILY 345 Heads disciform, the fig en iris present but essentially rayless. Bie gos southern Calif pappus about “ 12 Aaiite bristles; gen encmery 2~3 mm. divergen ong. 32.28. Inner fon about 15-20 firm pore lish corollas 4-5 mm. best babe Oute bese P ‘ adaak scales; stem often leafy, subnaked iefly in forms from southern California; southeastern Or fegeai to southern California ae eastward. te: erate Outer Lia setose and o or ng; stem naked; rthern Nevada fe Caliterain. pin As Be sr Washington. 35. E. chrysopsidis. Rays yellow Pu iemcenie of the stem spreading throughout; leaves narrowly oblance- olate, str. a Hes or slightly Bibles scarcely Seenene; stem sub- naked, the leaves all or nearly all in a basal cluster; extreme southeastern Washington to northern California and Nevada. hoe & stdts, Pubescence of the stem mostly appressed, at least above; leaves linear, fle eXUOUS; stem ee e. less leafy, at least below the middle; t scade unta . E. piperianus, Basal leaves more or less strongly triple-nerved. Involucre more or less spre rps. hirsute as well as | ons —_— t 10-25, or none; cautnab rn Califor . &. lass TInvolucre canescent with fine vatitte hairs (sometimes sae wed ‘Siikeaticaes also glandular, but i ao color ae about 30-100, Basal leaves ac phyl only slightly or obscurely thickened on the back; eae cider pi ish en She biak: pot of oe -- scade Moun- tains from Oregon to southern British Columbia and eastward. ee cor i OSUS. Basal Sete usually rounded or obtuse at the tip; yin es evidently thick- ned on the bac ik stem rarely purplish at ing e hase: eacenns eastern espitos phomty sinc a. = stem and leaves more or less closely appressed or weer never easels fends ieee: the flowers all tubular and perfect; widespread. 37. E. bloomeri. Heads evidently radiate. ase of the stem conspicuously enlarged, shining, and somewhat indurated, straw- colored or purplis Rays yellow; pa nptis-heteties 10-20: widespread. 38. E. linearis. Rays sin purple, or pink; barigna -bristles mostly 20-50. Lea ges: > e or less; involucre 3.5-5 mm. high: disk 6-11 mm. > pappus- bristles 20- 30; a os . Hanser og Modoc, n x peels n Cou s, Califor tulus. Leaves oblanceolate, a pony 5-5 m ; invo ree ce 5. 5 9 mm nee — 13-18 mm. wie Bah Jpristles | “rosy 30-40, sometimes n County to Fresno "County, Califor: 40. 7: barbellulatus. Base * lg stem not conspicuously enlarged, shining, and indurated (alpine specimens E. peregrinus would be sought here, except for the 4—7-nerved achenes). ges narrrow. linear or rather ngage So aaa the blade if distin- guishable tapering very gradually to the Basal leaves with their bases neither eee nor of different texture from the blade = leaves linear or linear- filiform; stem more = geod sal toward the base than above; southern Briti = Colum t erra County, Cilifacnia and eastward. E. filtfoli eM Basal =i de aan their bases co: aca SS . en whaseel, shilicntoms or rated, of different texture — the blades; leaves linear or more cho oblanceolate; stem not more densely hairy toward the base than above. Plants neither ee a nor scapose; basal leaves mostly triple- BE, flexruosus, Disk- lores mostly 5-7 m long; s ein Aare aes mostly 0.3- m. a lak! pappus . coaree and copious, the inner of 25-40 bristles; . involucre 7.5-10 m am gh; eastern C: ne to Disk- corllas ae 25-5 long; style-appendages vi 1-0.25 ong; pappus edletivety scanty “gas fragile, the inner of atts bristles dees mii 3.5-8 mm. high. or te pos mostly abruptly rete ed and smaller than the : ee ones; rays ches white; east of the Cascade and a Nevada summits excent in northern California. . E, eatonii. caine a only gradually ‘cies, rays mostly colored; iefly west of the Cascade summits. ae leaves more or less evidently triple-nerved; heads few or —- and romana or less naked- -pe- ee ns Ps ray ostly 20-5053 volucre hairy but poneey. pancslag.” tad in Californ nia forms mostly 6-8 mm. high; western vor and northern Cali- ep . E. decumbens. Basal — only obscurely if “se ‘all t triple-nerved; heads ral or rather numerous on leaty branches: rays eaouty 12-20; involucre gl — as well ba Sairy, 4-5 mm. high; Trinity aor ‘aliforn E. fle -TUOSUS. 346 COMPOSITAE Plants har alige Ara age scapose; leaves not triple- fro ibis Rese Moun- , California, and eastward. 49, E. compact Basal pny Mees sae roadi y oblanceolate or jae broader, ‘the blade defined, usually aah or less cheugits contracted to the pet Mca sree a ; involucral bracts subequal, finely glandular, a ‘hairy; Sis- kiyou Mountains re aetineestern California am Sg pen Fe Orego Leaves strigose, sometimes seein! so; involucral mes iaitanie imbricate, Lang Guieading: -hairy as well as Asset as ie — and south- rn Cascade Mountains pie att II. Wrom Achenes mostly 4-nerved, rarely 6-nerved; basal leaves often — hg flowering time, not forming a conspicuous persistent tuft; San Bernardino County, California, and e: Outer ar of evident narrow scales; stem silvery sine eed i. San a ge een re- 50. E. par Outer pappus = inconspicuous setae; stem merely gray-green except near the ie Providence, Mountains and e ard. . E. utahenst. iinet mostly e ins. nerved; basal leaves tufted and persistent, the cauline ones reduce: Inyo aa. Cali- ornia, and eastward. . BE. argentatus. III. PycnorpHyL_uM Heads risen radiate Root-crown nce subterranean, giving rise ender rhizomatous stems rarely more than 1.5 mm. thic ie’ hich become aerial stems; — pe By ir aaeihee, usually with solitary or few heads; Poca "gener- ally branched into equal parts n sone ground-level; pubescence of the stem except in one reduced and slender variety spreading or retr Stem finely glandular, as well as sissois with relatively long spreading hairs; ay County, California. . E. aequifolius. Stem not glandular, though the hair: a little viscid; pubescence of a praei relatively short or wanting; Shasta County to Biverade pal California, and eastward. Root-crown superficial or nearly so; base of stem, if rhiz —— either very short or soltioals a. or both; plants variously geey or stout, with go to generally several or many heads; stem except in var. confinis generally not branching near the ground- Nk g oe pad potter pubescence of the stem varying from apres to sirigdee Age spreading-hirtellous or even spreading- -hirsute ; rag a Oren o Lower Californ E. foliosus. Heads 7 the Pie s all tubular and perfec Stem and besa aiaicseaials she ese or spreading-villosulous, occasionally rather sparsely 6 gine +30 eaves generally 2-4 cm. long; outer pappus obscure or wan nting; Klamath re gion of siaaleck Cali- fornia t Oregon and in the Coast Ranges of California. "'s. E., petrophilus. Larger leaves generally 1-2 cm. long; outer pappus evident; Sierra Nevada el te ee Pass. miser Middle and upper part of stem not spreading-villous, varying from glabrous to glandular ond sometimes asc oer Sp ndiygine or he ree d-hairy ; leaves not | ‘spreading-villous, though sometimes otherwise 38. E. inornatus V. PHALACROLOMA Foliage ample; plants mostly 6-15 dm. tall; pubescence of the stem long and spreading. eae Foliage sparse; plants mostly 3-7 dm. tall; pubescence various. 60. E. Canoe: V. TRrimMORPHAEA Rayless pistillate flowers wanting; inflorescence racemiform, the peduncles erect or money so or the head solitary. . E. lonchophyllus. Rayless pistillate flowers present between the ra fe - disk-flowers; inflorescence ase. the peduncles arcuate or obliquely ascending or the heat soli 62. E. acris. 1. Erigeron peregrinus (Pursh) Greene. Wandering Daisy. Fig. 5582. Aster peregrinus Pursh, Fl. Amer. Sept. 2: 556. 1814. Erigeron peregrinus Greene, Pittonia 3: 166. 1897 Fibrous- mee perennial from a short rhizome or short stout caudex, up to 7 dm. tall, amply leafy or in salt f orms subscapose. Basal and lower cauline leaves well developed, with linear- eolate to broadly oblanceolate or spatulate blade tapering to the petiole, the other rs rae tly sessile and often slightly clasping; heads solitary or few, the di sie 10-25 mm. wi vo- lucre oN mm. Byles the phyllaries yl attenuate, loose, mostly rather herbaceous aadaaiie, (June) July—Aug. nomenclaturally typical phase does not occur in our range. The foregoing description is drawn to cover all forms of the species. Key To SuBSPECIES AND icasewise Phyllaries villous on the back or sometimes merely ciliate on the margins and glutinous on the back, not at all glandular; rays commonly rather pale or even white: herbage meee soft-pubescent, the stem usually gros villous, the peduncular hairs rather loose; leaves often penal coastal and Cascade Mountains from tral Washington northward, not i to the interior ra subsp. n peregrin TInUs. é SUNFLOWER FAMILY 347 Upper ae leaves reduced and distant; northern Washington to Unalaska. r. dawsonit, Upper ax ine Joncas sates ample or closely set; chiefly from the Alaska panhandle northward, occasional h Colum r. peregrinus. Phyllaries sechety vrrcalatien « on ie back, ciliate on ser margins; rays ordinarily white; foliage crown § bogs along the southern fringe of the — ee yo a ains in Washington r. thompson Phyllaries densely glandular on the back, ra a ee long hairs; rays donamonty rich rose-purple or 5 cane her i bage (except in var. hirsutus) Diag ie peat except for the closely villous peduncles; leaves ordinarily en entire; i "interior mountains of British Columbia and Alberta southward to Washi mate nm, California, Utah, d : ew Mexico. ubsp. callianthemus. aac hirsute on both surfaces; peduncles spreading-hirsute; central and southern Sierra Nevada of Cali- fornia extending into adjacent Nevada. ee sly Leaves geneous on ein Moa rfaces, sometimes with a few hairs along anny midvein, or ‘rabaly short-villous; peduncles with ssed or as acon. biter euvieae or rai hai Reduced aie oan gee than 2 dm. tall, with a sample, eis bree ae there leaves and very ich smaller cauline leaves, often subse: ey of the subsp wn the Sierra Bevade. caposus, Larger, mostly subalpine or merely montane plants, up to 7 dm. tall (smaller in ial phases of var. angustifolius but then with narrow, acute, basal got Leaves Bae ier the basal oblanceolate or na the cauline linear or lanceolate; Sierra Nevada f California and adjacent athe pieading aatiwask less copsmamaped is southern pene Colt mbia pl tfolius. Leaves more ample ee basal oblanceolate | or broader, the saviine mostly “tease “and ot greatly re- duced; t for the Sierra Neva callianth emus. Erigeron peregrinus Caen, Seite tink var. dawsdnii Greene, Pittonia ra 166. 1897. Type locality: Queen Charlotte Islands, British Columbia. Erigeron — nus subsp regrinus var. thompsonii (Blake) Cronquist, Brittonia 6: 1947, (Briger ae war 2? Blake or Diiunpsee, Rhodora 34: 238, 1932.) Type locality: Lake Quinault, Ph Harbor ounty, Wa Erigeron peregrinus —, callianthemus, (Greene) Cronquist, Rhodora 45: 264. 1943, — callianthemus. (Erigeron membranaceus Gre Pittonia 3: 294. 1898: E. callianthemus Greene, Leaflets Bot. Obs. 2: 197, 1912; E. peregrinus a callianthemus var. callianthemus £. membranaceus Cronquist, Brittonia 6: 145, 1947, a form yaad be n, broad, strongly clasping leaves; E. peregrinus subsp. callianthemus var, callianthemus f. sub- villosus Cro t, op. cit. 146, a form with shortly villous leaves; E. percgrinus subsp. callianthemus var. ca ‘alli. a net f. dentate Cronquist, loc. cit., a form with toothed leav es.) Type locality: Petaail, Albany County, n peregrinus subsp, callianthemus var. angustifolius (A. Gray) Cro mae ri Brittonia 6: 147, (Aster eiigiacee var. angustifolins A. Gray, Bot. Calif. 1: 325. 1876; Eri rigeron salsuginosus var. angu. stifolius A. Gray, Proc. Amer. Acad. 16: 93. ens Ee Bas yar bbe ry Ry db. Bull. aoe, Crab. 24: 295. 1897; E. hespero callis Greene, Leaflets Bot. Obs. sh 200. fis us Greene, op. cit E. regalis Greene, op. cit. 205.) Type locality: S i Erigeron Pisa teg subsp. callianthemus var. hirsitus Cronquist, Brittonia 6: 147, 1947. Type locality: Lake Kiger ss semite National Park, California Eri nm peregrinus subsp. relilsethedius * var. scaposus (Torr. & Gray) sewer Pencrnonnty 6: 146. 1947. (Aster gracilis Nutt. Trans. Amer. Phil. Soc. II. 7: 291. ioe; A, salsuginosus v osus lag & Gray, FI. N. Bs 503. 1843; Erigeron salsuginosus, vee. glacta Gray, Syn. Fl. ym pr on 12; 209. 1884; E. glacia Nels Bot. Gaz. ae 207. 1904; E. suksdorfit Crem: Leaflets Pe Obs. 2: 203. 1912; E. ciliolatus 2. ib Se howéllii A. Gray. Howell’s Daisy. Fig. 5583. r. howellii A. Gray, Proc. Amer. Acad. 16: 93. 1880. ve howethe 1c i Syn. Fl. N. Amer. 17: 209. 1884. Pgs pspeged perennial from a short rhizome, 2-5 dm. tall, marty oe noes under t es entire or irregularly few-toothed, thin, glabrous, the lowermost ones with clliptic or pri Pike 2 blade 2-8 cm. long and 1.5-5 cm. wide, abruptly eer my rs “the 2-12 cm. petiole ; 348 COMPOSITAE middle cauline leaves ample, ovate to cordate, strongly clasping ; upper leaves similar but smaller ; heads solitary, the disk 12-20 mm. wide; phyllaries loose, equal, acuminate or attenuate, glandular, seeaptlesa oe, Hage rays 30-50, white, 13-25 mm. long, 2-4 mm. wide; | pubigass mm, long, m ring than in FE. peregrinus ; eclecueemaies acute, 0.3-0.4 mm. long; achenes mostly aieneewanicdity 5-nerved; pappus of 20-30 bristles. Moist, often rocky places, Hamid: Si nsition Zone; south side of the Columbia Gorge, Oregon. Type locality: Cascade Mountains, Oregon. April—Jun s eigersp aliceae Howell. Eastwood’s Daisy. Fig. 5584. Erigeron aliceae Howell, F poe Piaget 317. 1900. Eerie ie Howell loc 9 Mg eee Bot. Obs. 2: 210, 1912. Fibrous-rooted perennial from a rather short rhizome or woody caudex, 3-8 dm. fall amply. leafy. Leaves hirsute on both sides, entire or coarsely toothed, the lowermost ones up to 20 ¢ long (including the petiole) and 3.5 cm. wide, the middle ones sessile, narrowly lanceolate a ae or ovate ; heads 1 to several, the disk mostly 2-20 1 mm. wide; phyllaries se vaeie 2 attenuate, subequal, conspicuously white-hirsute on the lower one- -fourth 8 three-fourths, glandular thence a the tip; rays as ce 1 mm. long and 2-3 mm. wide, white to pink- acute disk-corollas mostly 3-4 m g; ne appe enianes 2 Santis: about 0. 25 ‘mm. long; achenes 2-nerved or sometimes Ae — pica of 20-30 bristles Moist or fairly dry soil in uae or open places in the mountains, Canadian and Hudsonian Mount Hood ooa tag a Mountains, and‘in ~ oe Mountains = Oregon and seen “California; epi fi the Olympic Mountains, Washington. Type locality: top of Siskiyou Mountains near Waldo, Oregon. 4. Erigeron coulteri Porter. Coulter’s Daisy. Fig. 5585. Erigeron coulteri Porter in Port. & Coult. Fl. Colo. 61. 1874. Erigeron frondeus Greene, Fi. Fran. 387. 1897. Erigeron leucanthemoides Greene, Leaflets Bot. Obs. 2: 211. 1912. Fibrous-rooted perennial from a slender rhizome or branching caudex; stems 1-6 dm. tall, usually amply leafy, spreading-hirsute at least above. Leaves hirsute, at least the lower bites generally toothed, the middle cauline ones mostly broadly lanceolate to oblong or ovate, tending ‘s sping at the base, up to about . long and 3 , larger or smaller t he per- sistent or deciduous, mostly more petiolat e, low r ones; heads 1-4, the involucre 7-10 mm. high, its nes Bae phyllaries equal, attenuate, villous hirsute, Be) hairs with black cross- ee — the base; ray aye Sah white, 9-24 mm. long, 1.2-1.7 mm. wide; disk-corollas 3 ong ; eri itil a ore or les soo 0. 2-0. "38 mm. ees inner pappus of 20-25 bristles, the outer obscure or sanjay Se achenes 2-nerved. Meadows and stream Sant rite at rather high elevations in the mountains, Borea] Zone; norther and ee an ntana to northeastern Oregon southward irregularly to the southern Cascade Mageen Me. Sierra Nevada in California, the Wasatch Mountains vA _ ah, and the mountains of Colorado and northern New Mexico. Tata locality: Weston’s Pass, Colorado. July-A 5. Erigeron specidésus (Lindl.) DC. Showy Daisy. Fig. 5586. Stenactis noreeAd Lindl. Bot. — ao: on arr 1833. Erig DC Prod. 5: More or less fibrous-root se peren iad from a woody caudex, the stems clustered, 1. 5-8 tall, pei ready, naeraded giabr rous below the inflorescence. Leaves entir e, glabrous o an except toe the commonly ciliate margins, often triple-nerved, the lower oblanceolate or seatelate, petiolate, mostly deciduous, the others becoming sessile but fairly ample, the uppermost ones mostly lance olate ; heads 1-13, the involucre 6-9 mm. high, glandula ay gee ommonly with a very few long airs; rays 65-150, blue or rarely white, 9-18 mm. long, a m. wide; disk-corollas 4-5 mm. ong; style-appendages acute, 0.15-0.2 m m, long pappus of “30-30 bristles and ‘some ae outer setae ; achenes 2-ne rved or occasionally . -nerved. pen in woe ooded are, mostly in the foothills and at moderate elevations in the mountains, Tisaation Steg pee aeg Bri ish Columbia, Washington, northwestern Oregon, nor Be lao, and _ north- estern Mon bide: Deahts: “California” ; probably actually in northern regon so n Washington I une—Aug, A few of = resemble the otherwise more eastern and southern var. agg Pata: leg (Nutt.) Cronquist the Rocky Mountains and eastern Great Basin region, characterized by its broader (m ~ oe } upper leaves that tend to be less ae ‘ciliate: likewise, occasional specimens throughout the veneer re acranthus re- semble typical E. speciosus. 7 6. Erigeron subtrinérvis Rydb. var. conspicuus (Rydb.) Cronquist. ie ee Daisy. Fig. 5587 Eee COnSPIEN RE | Rydb. Mem. N.Y ies 1: 400. 1900. Greene. Leatiets Bot. "ae 2: 215. 192. Princes sROEriNEr eis subsp. conspicuus Cronquist, Bull. Torrey Club 70: 271. 1943. Erigeron var. conspicuns Cronquist, Vasc. Pl. Pacif. Northw. 5: 193. 1955. Closely related to E. specie s and not always wasiried ee tinct, but more pubescent, the s leaves, os especially the seni ie evidently pubescent with long sp reading tale the ae of the leaves ir “ ee chiefly to ig mara on larger Open woodlands, usually in drier places than E. speciosus, Canadian and et Transition Zones; southern SUNFLOWER FAMILY 349 British Columbia and Alberta to Washington, central Idaho, western Montana, and northwestern Wyoming. Type locality: srg Creek, Montana. (June) July-Aug. Typical E. egediond rvis, characterized by its algal and denser pubescence, occurs from Wyoming, Utah, d Naw Mesicltc estern Nebraska and South Dakot 7. Erigeron philadélphicus L. Philadelphia Daisy. Fig. 5588. Saale philadelphicus Ev Sp. Pl. 863. 1753. n purpureus Ait. Hort. Kew: 3: 186. sie: apis purpureus var. sttnnat Nutt. Trans. Amer. Phil. Soc. II. 7: 307. 1840. ssenia philadelphica L 1, Amer. Midl. Nat. 5:59. 191 é. Erigeron phi ladelphicus £. ce Farwell, op. cit. 11: 70. "1928. Biennial, or short-lived perennial or eek apparent, che mostly fibrous-rooted, commonly 2-7 dm. tall; herbage long-spreading-hair subglabr gue Basal leaves mostly ob- eect toothed 0 r lobed to Senaetinbes Sotre. geliota dls re cm. long and 3 cm. wide; cauline leaves becoming sessile mostly ample and more or less clasping, Aaa so in smaller specimens; head 1 to baie’ invo om ucre 6 mm. high, the phyllaries subequal, light greenish or brownish, the brov idvein more or les Sina with flattened hairs, or nearly glabrous, the broad hyaline mar- a pectslanetty wit ptr: disk Py Lb m. Pee rays 150-400, deep pink to white, 0.2-0.6 mm. wide, 5-10 mm. long; disk-corolla 2.8. 3. ong; pile appendages obtuse or acutish, about 0.1 mm. long; pappus of 20-30 fcaatie bistles acta 2-nerved. In a wide variety of habitats. most commonly in moist, often also ie turbed soil; throughout the United States and most of Canada. She locality: Canada. Mostly May fils or sometimes until fall. Highly variable in size and luxuriance, according to the habitat. 5584. Erig: 5586. Erigeron speciosus : 5585. Erigeron coulteri 5587. Erigeron subtrinervis alice: 350 COMPOSITAE 8. Erigeron glaicus Ker-Gawl. Seaside Daisy. Fig. 5589. Erigeron glaucus Ker-Gawl, Bot. <2 1: fas bass 1815. Stenactis glauca Nees, Gen. & ai thbeer: ii eaed W oodvillea calendulacea DC. . 5: 318. 1836 Erigeron hispidus Nutt. Trans ‘mes Phil. Soc. II. 7: 310. 1840. Erigeron maritimus Nutt. loc. ?Erigeron squarrosus Lindl. art Won: 27: 44. 1841. Somewhat succulent, maritime perennial from a stout rhizome or branched caudex, highly variable according to exposure, 5-50 cm. high, the stem curved at the base, more or less spreading- i at gla mm. long, narrow, with short lobes about 0.5 m m. long ; style- appendages obtuse, a uit mm. long ; iio usually . —— bristles, Cae eat with some inconspicuous, short, outer. setae; pie oe mostly 4( ved. Sea bluffs and oat | cae s along the coast, under the influence of “ water, Canadian and Humid Transi- tion Zones; Clatsop County, Oregon, southward to San Luis Obispo Cow and the Santa Rertecs a Cali- fornia. Type locality: iathalty thought to be South America; dectilcas Salty California. April-A 9, Erigeron sapplex A. Gray. Supple Daisy. Fig. 5590. Erigeron supplex A. Gray, Proc. Amer. Acad. 7: 353. 1868. Submaritime perennial with a short branched caudex which sometimes surmounts a taproot, 15-40 cm. tall, the stems decumbent at the “aan sparsely to moderately pubescent with long. spread- L ing hairs and sometimes obscurely glandular. Leaves entire, cHiatenaareite , otherwise sub- glabrous to mod villous-hirsute, relatively numerous and crowded, scarcely tufted at the se, the lower ones oblanceolate, petiolate, up t lo d7 mm. wide, those above similar b ‘ a subnaked peduncle 2-10 cm. long, the disk 14-20 mm. wide, very broad in shape; involucre 7-11 mm. high, sparsely to densely villous-hirsute with long, sprea ading, white hairs, alo viscid or finely glandular, the phyllaries subequal, green, wi ith paler thin margins and thin, attenuate or acura tip; pistillate flowers wanting ; disk-corollas bright pion be ow, 4. en lon e lobes ut 1 mm.; style-appendages obtuse, 0.2-0.25 mm. long; pappus of 17-30 firm jon, ‘with a i d. Near the seacoast, Humid Transition Zone; aig no and Humboldt Counties, California. Type locality: near Mende no, Mendocino County. Muactly May-July 10. Erigeron sanctarum S. Wats. Saints’ Daisy. Fig. 5591. Erigeron sanctarum S. Wats. Proc. Amer. Acad. 24: 83. 1889. Perennial, apparently from a slender a which may perhaps connect to a se bios root-crown, 5-35 cm. tall, the stem ascending or erect from a mostly curved base, sp y pubes- cent, especially above, with spreading or ir i hairs. Leaves entire, shortly hispid. * Shire on at least the upper surface and generally aie ciliate-margined, the basal ones 5 cm. long, 3-10 eolate e i i upward, i na nake 2-10 cm. long; disk 12-17 mm. broad; involucre 6-9 mm. high, evi idently spreading-hirsute, the poy ies oT narrow, attenuate, green with bia brown midvein; rays 45-90, blue, 7-13 . long, 1 9 mm. wide; disk-corollas ong ; style-appendages o obtuse or acutish, 0. 1-0.25 mm es pappus of 18-25 beistles: seanetibes with some short and inconspicuous outer setae; achenes 2-nerved. Hills near the seacoast, Upper Sonoran Zone; Santa Barbara and San Luis Obispo Counties, California. Type locality: Santa Inez Mountains eae ‘Santa Barbara. March-June. 11. Erigeron delicatus a Del Norte Daisy. Fig. 5592. Erigeron delicatus Cronquist, Brittonia 6: 216. Fibrous-rooted perennial from a ae rhizome or slender caudex, glabrous below, finely stipi- ai ate etka & above, the stems 2-3 dm. tall, slender, arising singly or few together Leaves entire, rather numerous and eauably ist distributed the middle cauline about as eee as the ‘ier. the uppe gradually reduced, the low with gee cegre obtuse, or rounded blade 1-3 cm. long and 4-8 mm. wide tapering to t pte arent petiole, the others promee sels less petiolate or sessile ; head ry or short, SGemiiietandaiar = ncles ; 13 mm. wide; involucre 5-6 mm high, the phyllaries glandular or glabrous, subequal, gree Siidh strarainencs, the tips acute or acumi- nate, sometimes purplish ; rays about 40, blue, 7-10 mm. long, 1.5-2 m m. wide ; disk-corollas about 3.5 mm. long, — based, slightly flaring above the middle; style-a rosctte acutish, 0.2 mm. ong; pappus of about 15 slender bristles and some inconspicuous, short, slender setae; achenes 2-nerved. At lower elevations, nas mcutiee: 5 iatg — Ear gg Eg Spee! jot lata po California, and adjacent Orego ype SUNFLOWER FAMILY 351 YW f My aH, \ "4 iN i ; A NH 5588. Erigeron philadelphicus 5589. Erigeron glaucus 5590. Erigeron supplex 352 COMPOSITAE 12. Erigeron cervinus Greene. Siskiyou Daisy. Fig. 5593. Erigeron cervinus Greene, Pittonia 3: 163. 1897. rous-rooted perennial from a branching caudex, 1-3 dm. tall, the herbage glabrous except immediately under the heads, where usually fine . a lar, Leaves entire, the basal ones broadly oblanceolate to elliptic or obovate, up to 12 cm. g (petiole iacadday nd 15 mm. wide, usually —4, mm. high; plaries subequal, ei raceous purple- ig He the outer sagen the er acuminate; rays 20-45, blue or sometimes pink-purp “Sa 10 mm. long, 1.3-2.7 mm: wide; “disk. corollas 3.5-5 mm long, ae limb so somewhat puhereleet ee the era Zt le- sara iarts obtuse or acutish 0.2-0. 25 m. lon pus of about 12-15 bristles, with 4 few short outer setae; achenes 2-ne gL ea nd open pla Canadian Zone; Siskiyou Pid region of northwestern Cations ad adja- cent Oregon. ‘Pree locality : De: Creek Mou ntains, Josephine County, Oregon. June-early Aug 13. Erigeron cascadénsis Heller. Cascade Daisy. Fig. 5594. Erigeron spatulifolius Howell, Fl. N.W. Amer. cok —— Not E. spathulifolius Rydb. 1899. Erigeron cascadensis Heller, Muhlenbergia 1: 6. Erigeron pentane Greene, Leaflets Bot. Obs. ;. co 1912. Perennial from a stout branched caudex which may surmount a short taproot, 5-15 c m. tall, the Site glabrous or with a few scattered glands and/or spreading hairs. Basal leaves ce. a cm. long, 5-1 i nous green with purplish acuminate or attenuate tip; rays 30 50, mia white, mm. long, 1.2-2. mm. wide ; disk-corollas essentially glabrous, 7A. 4 mm. long; style-a ppendages acute, 0.2- mm. ctor Be pappus double, the inner of 15-20 firm bristles, ‘the outer of short narrow scales ; achenes rved. 2-n ah rocky places, moderate to high elevations in the mountains, Canadian and Hudsonian Zones; Cas- cade and TCclapnole Nokia ins, Satin + , from about the 43rd to the 45th carallel” Type locality: Pansy Camp, Cascade Mountains. June-Aug two 14. Erigeron leibérgii EP Leiberg’s Daisy. Fig. 5595. Erigeron leibergii Piper, Bull. Torrey Club 28: 41. Erigeron chelanensis St. John, Sacral Stud. St. ne ce 13.107. 1929; rennial from a stout branched caudex which may surmount a taproot, 7-25 cm. tall; sac and invol soe sparsely to eoderaee villous-hirsute with flattened, flexuous hai irs and more 0 r less eaelia rtanduler Basal leaves tpn se ance ae to obovate or elliptic, sometimes ie nerved, entire o i wide; cau al, i to l developed, ovate to oblong or eo aclate: “etita tire or up to e disk 7-14 mm. wide; involucre 5-8 mm. high, the phyllaries subequal, thin, loose, green, the attenuate or acuminate tip often purplish; rays 20-45, mostly blue or pink, rarely white, 5-12 mm. long, 1. =. 2.0 mm. wide; Ge oncetas 3.0-4.3 mm. long; style-appendages acute or peutish, tak stly 0.25 mm. long; pappus of 12-16 bristles, sometimes with a few short, slender, Sito etae ; sed pe " -nerved. Cli a and rocky places at moderate elevations, Boreal Zone; ata a and Wenatchee Moatain of Okanogan, Chelan, and Kittitas Counties, grin tivetony Type lo cality: Mount Stuart, Chelan County. June-Aug 15. Erigeron oreganus A. Gray. Gorge Daisy. Fig. 5596. Erigeron oeeren & Gray, Proc. Amer. ee 19: 2, 1882, ial with wintieh villas: stems lax, 5-15 cm. lon ng. Basal leaves pee ods spatu ate to obovate, coarsely toothed or incised, up to 9 cm. long and 2s em. wide; cauline leaves well developed, broadly lanceolate to elliptic or ovate, up to 4 cm. em: and 1 cm. wide; San ds 1 to several in a leafy inflorescence, the - sk 9-13 mm. wide; Seale e 5-7 mm. high, glandular and viscid-villous, the ica we loose, ual, thin, green, attenuate rays mostly 30-60, bluish to more often pink or white, on disk-coro llas 3.4-4.7 m pappus simple, of 15-20 bristles racbich are sii ceiscristically urled and twisted for at “least the upper half. Moist shady cliffs and ledges, Humid Transition Zone; Caentis River Gorge, most frequently collected on the Oregon side. Type locality: Columbia River, Oregon. June—Sep 16. Erigeron basalticus Hoover. Basalt Daisy. Fig. 5597. Erigeron basalticus Hoover, Leaflets West. Bot. 4: 40. 1944. Stem ral fro ial taproot, sprawling or pendent, branched, leafy especially near the tip; eethert spreading-hirsut and finely gla rahul Leaves cuneate to obovate, up to about 4 cm. long and 1.5 cm. wide, more or less de a and often irregularly trilobed, the lobes broad, ium slightly lobed amity heads terminating the branches, the disk 8-12 mm. wide; involucre 5-6 mm. high, densely glandular and sometimes sparsely lon g-hairy, the phyllaries subequal or slightly imbricate, —_— the slender acuminate i ps commonly purplish; rays 25--30, pink or pink-purple, 5-7 mm. long and 1.5 mm. wide; disk-corollas 3-4 mm. long; s Poh faotionatersai obtuse, 0.1-0.15 mm. long; pappus of 10-15 bristles, wh ionic taconipien uk Sime outer setae; achenes 2-nerved. SUNFLOWER FAMILY 353 crevices in rocky canyons at low elevations, Arid Transition Zone; Yakima County, Washington. Type A vi at Creek ana ticity. Yakima County. May. —Oct 17. Erigeron vagus Payson. Loose Daisy. Fig. 5598. Erigeron vagus Payson, Univ. Wyo. Pub. Sci. 1: 179. 1926, Perennial from a diffuse, oe st pede caudex which sometimes connects to - we te taproot; herbage and involucre more or less evidently spreading-hirsute and glandular. Lea crowded on the short aerial stems, meetly 3-lo sak with short, broad, r oo nded lobes ; head solitary, borne on short scapiform peduncle up to 5 cm. long; disk 8-16 mm. wide; involucre 5-7 m high, the phyllaries subequal, purple at least —— the tips; ray 5 25, 35, white or sink 4- long, 1-2 mm. wide; Aisle beitatias 3.0-3.8 mm. long; style- candies’ acute, 0.2-0.25 mm. ace pappus of about 20 bristles; achenes Alte or Shifting talus slopes at high altitudes in the mountains, Ar-_tic-Alpine Zone; 180 stage cand pa, ang agg: Nevada, California (Sierra Ne mses - gic ortheastern Oregon (Wallowa Mountains). Type locality maski, LaSal Mountains, Utah. July-A 5594 v3 F wy == a = XS e Woe Yor : : oY, a4? hry igh we Vl \ : VE oe Erigeron hme nsis 5596. Erig sane 95. Erigeron leibergii 5597. Erigeron basalticus 354 COMPOSITAE 18. Erigeron compdsitus Buea Cut-leaved Daisy. Fig. 5599. Erigeron compositus Pursh, Fl. Amer. Sept. 2: Cineraria lewisit Richards. in Frankl. 1st tert Bot pane 1823. Erigeron compositus var. grandiflorus Hook. or. Amer. 17. 1834. Erigeron compositus var. submontanus M. E. stabs Torreya a 56. 1928. Perennial from a taproot and branching caudex, 1-2.5 dm. tall, the herbage densely glandular and pe cated haxsute to subglabrous. Basal lea aves Panam dissected, the better- Pua eloped ones mostly three to four times ternate, often irregular ly so, with 1 relatively peer an ind linear divisions ; Ww ved. Sandy river banks at low icra Arid Transition Zone; Washington, iene sa ae Idaho. Type locality: “‘banks of the sts robably r the present ou ae” Tashi April—May. Erigeron compositus var. pests atus Se kanint Cat. Can 1884. ri multifidus Rydb. Mem. N.Y. Bot.’Gard. 1: 402. 1900; E. — var. “ac "Ryab. Te yes : E. multifidus var. incertus A. Nels. Bot. Gaz. 30: 198. 1900; E. compositus var. s A. Nels. in Coult. & Nels. New Man. ee ocky Mts. 529. 1909; E. compositus var. nudus A. Nels. loc, srl 4% compositus var. spinal on & Payson, Contr, Gra ay. No. 49: 75. 1917; E. compositus var. petraeus Macbride & P . 76.) Best- serene leaves n the seg com- a in typ ‘ positus ; plants averse mega Bare in typica compos tam re rage aoa slandulse and Abed in the aac oO subg ye A high siemakianar pi ra to Green land southward in the cordillera rf California (Sierra Nevada), northern Ari- gone Colorado, and South Dakota; also in Quehes Type locality: South Kootanie Pass, 49th parallel. June— ug Erigeron seeipeetens var. discoid Gray, Amer. Journ. Sci. IT, 33: 237. 1862. (Erigeron trifidus Hook. FI. Bor. =. tid 1834; E. ‘edatus Nutt. Trans. Amer. Phil. Soc. IT. 7: 308. 1840; E. compositus var. trt shees A. Gra Sings dl Acad. : 90. cls. in E. ee var. discoideus Rvdh. Mem. N.Y. Bot. ° 3 oo 4 o 4 w Al a s Mo. S . + = a < > a ee ae Rx. s ae 6 sl Sy oS. 29 . ° = a ae Q baal Pe a ° a by pe g s} S a =e ~ =* i 4 Py “ co other varieties and sometimes Re ted Esra Me scien mbt: Total distribution sibel og re in var. glabratus, but in pts range rare and confined to the highest elevations. Type locality: alpine ridges lying east of Middle Park, Colorado. June—Aug 19. Erigeron aiireus Greene. Alpine Yellow Daisy. Fig. 5600. Aperere. hers ei A. Gray, Syn. Fl. N. Amer. 12: 132. 1884. Not Erigeron brandegei A. Gray, 1884. Aster brandegei Kuntze, Rev. Gen. PI. 1: 317. 1891. Erigeron aureus Greene, Pittonia 2: 169. 1891. Stenotus brandegei Howell, Fl. N.W. Amer. 300. 1900. Fibrous-rooted to weakly taprooted perennial from 2 branching caudex, 2-15 cm. tall, the herbage pte heacent with short, appressed or loose hairs. Basal leaves petiolate, with ‘elliotic to obovate or subrotund, mostly broadly rounded or eitae blade up to 13 mm. wide; cauline leaves few and reduced; head solitary, the disk 7-16 mm. wide; involucre 5-8 mm. high, its bracts loose, equal, herbaceous, sometimes anthocyanic, sparsely to densely bg gs By einen the hairs sometimes with purple cross-walls ; rays mostly 25-70, yellow, 6-9 mm. long, 1.4-2. 5 mm. wide; disk-corollas 3.6-4.9 mm. long; style-appendages acute, 0. 3-0.5 m rigs HE pappus of 10-20 bristles and some sie outer setae or narrow scales; achenes 2- ue A ¥. ocky es at high te eet in 7a mountains, Arctic-Alpine Zone; southern Alberta, southern Brit tish LB Sicniies rie “the Case: ge s of Washington. Type locality: “Mountains of Washington Terr., in the Yakima district.” J ta 20. Erigeron simplex Greene. Alpine Daisy. Fig. 5601. Erigcron simplex Greene, Fl. Fran. 387. 1897. Fibrous-rooted perennial with few or solitary, more or less viscid- villous, pee erect stems 2-20 em. tall arising from a simple or moderately branched caudex. Basal leaves oblanceolate or spatulate, up to 8 cm. long and 13 mm. wide; Canin ne leaves pets and eae head solitary, the dis mm. wide; involucre 5-8 mm. high, eteray to densely white-woolly-villous with flattened multicellular bani the Fe tine thi n, equal, green or often rig sh often ae of the outer loose or reflex rays 50-125, blue, pink, a rarely white, 7-11 mm. long, 1— wide ; disk-corollas 3-3.6 mm. long; Ge sheciiages acute, about 0.2 mm. long; pappus of 10-18 caulk lary fal Afr eg and some short, outer, setose scales ; achenes ree rved. slopes and dry ibhciiies at hich altitudes in the oe Arctic-Alpine Zone; northeastern Oregon (Wallowa Frye to Montana, northeastern Nevada, northern Arizona, and New Mexico. Type locality: or 21. Erigeron fléttii G. N. Jones. Olympic Daisy. Fig. 5602. Erigeron flettii G. N. Jones, Univ. Wash. Pub. Biol. 5: 244, 1936. less fibrous-rooted perennial from a stout branched caudex, 5-15 igs tall, od Saad sparsely spreading Bind a = largely basal, spatulate or oblanceolate, up to 5 esr and 12 mm. wide, glabrou t hirsute, marginally ciliate; cauline leaves tow an head solitary, the disk mostly “0-15 mm. wide; involucre 6-8 mm. high, sparsely to o eatty long-villous and sometimes also viscid, the phyllaries shou equal, thin, green, purplish above; rays st SUNFLOWER FAMILY 356 COMPOSITAE mostly 25-50, white, ay mm. long, 1.5-2.5 mm. wide; disk-corollas 3.5-4.5 mm. long; style- appendages acute, about 0.25 mm. long; pappus of 15- 20 capillary bristles and numerous stork outer setae ; achenes cpus Cliffs and other rocky places, at ie altitudes, Arctic-Alpine Zone; Olympic Mountains of Washington. Type locality : Olympic Mountainé. July- 22. Erigeron petiolaris Greene. Sierra Daisy. Fig. 5603. Erigeron petiolaris Greene, Leaflets Bot. Obs. 2: 205. 1912. Erigeron algidus Jepson, Man. FI. Pl. Calif. 1052. 1925. Fibrous-rooted to weakly tapr peed pipe from a branching caudex, 2-30 cm. tall, te stem loosely soteatiine hiesut te and also viscid or glandular. Leaves shortly spreading-hairy or occa- sionally glabrate, the basal ones ve iauceataee io elliptic or obovate, up to about 7 cm. lo * and 12 mm. wide, eat or rounded at the apex, the blade commonly rather abruptly contracted Lt the petiole ; cauline leaves few, reduced, linear ; head solitary, the disk 8-18 m e; e 5-8 mm. high, glandular and more or less hirsute or villous-hirsute with spre eading ties the Tohitiacies about equal, green with purple margins and tips, or purplish throughout; ra seg 125, blue, rose- purple, or pink, 5-13 mm. long, 1.2-2.6 mm. wide; disk-corollas 3.04.8 m inn tyle- pci ages acute 0.15-0.3 mm. long; pappus of ad a ) coarse but rather fragile th py with or it a few obscure outer setae ; achenes 2-ne ne : places and meadows near and above Pe eae Zone; Sierra Nevada from Mount ee Nevada, to Mount Whitney, California. Type locality: Alta Meadows, Tulare’ County, eee July—Aug 23. Erigeron cloékeyi Cronquist. Clokey’s Daisy. Fig. 5604. Erigeron clokeyi Cronquist, Brittonia 6: 214. 1947. Perennial with a stout taproot and usually branched caudex ; stems lax, spreading or ye 3-20 cm. long, moderately or densely terse with short spreading ats not glandular a few uniformly pubescent with short sp a ie hairs, sometimes er hairs along the pr Siri basal leaves pri oblanceolate, up to about 8 cm. long d 6 10) oe wide, the blade sometimes slightly triple-nerved at cauline te ssn ous, mo linear o eg - mee head solitary or occasionally emispherical to turbinate, the disk 8-11 wile: nvoluc e 4-7 m m. high, the ofeilaries a eis thickened and somewhat imbricate, tees ae senecialie on the midrib more or less Seakg epee ; rays 20-50, blue or pink, 6-10 mm - lon g, 1-2.2 mm. wide; disk- -corollas 2.9 m. long; style appendages blur AMY es Aaah rocky places at moderate to more often high elevations in the mountains, often in dry meadows or flats, osm ge Zone; ir wi “ie east slope of the Sierra Nevada from Mono County to Inyo ppt eg a —_ ard to Nye and Clark Counties, Nevada, and Beaver County, Utah. Type locality: Lee Canyon, Charleston iasisintes, "Clark County, Nevada. July— Aug. 24. Erigeron pygmaéus (A. Gray) Greene. Dwarf Daisy. Fig. 5605. Ererron senadensss var. legos nade Pe Proc. Amer. Acad. 8: 649. 1873. Greene, FI. Fran. 1897 Palvinate-cespitose perennial eee a stout taproot and a stout branched caudex ; stem u es 6 cm. high, more or less pubescent with short spreading hairs and usually also viscid or ine glarninler: Leaves ve a hirtellous or coarsely strigose with — ens or spreading hairs, the basal ones oblanceolate or linear-oblanceolate, up to 2.5 cm. lon 2.5 mm. wide; t or occasion a re i e dis : more or hirsute ; ig cane oh pave subequal or somewhat imbricate, deep blackish Bgsl-2 or with green body and purple “thi ¢ s 15-35, blue or purple, 4-7 mm. long, 1.1-1.8 mm. wide; dis k-corollas -5.3 m tig Se ng; s Loe appendages acute, 0.3-0.5 m m. long; pappus of 18-35 firm bristles, with a few are cate d short setae; achenes 2-nerved. Rocky places at high elevations in the mountains, Arctic-Alpine Zone; Sierra Nevada from ome Rose, Nevada, to Mount. Whitney, California. Type locality: Mono Pass, Mono County, California. July—Aug ad 25. Erigeron uncialis Blake. Cliff Daisy. Fig. 5606. Erigeron uncialis Blake, Proc. Biol. Soc. Wash. 47: 173. 1934. Perennial with a taproot and branching caudex; stem 1-5 cm. high, naked, hirsute or villous under the head, with spreading faite: ee all heal, ag se or hirsute- strigose with loose or aneens ed hairs, sometimes slabrate on one or both surfaces, up to abo m. lon and 8 mm. wide, the blade subrotund to broadly oblanceolate, codes to pe ute at the apex, epee —— contracted ig and usually much shorter than the petiole; head solitary, the disk 11 mm. “ae ; involuc mm. long, finely and sometimes obscurely glandu lar, and more or less vil- ous fds ance a toward the base ; phyllaries Gate or Sor pe ‘clightly imbricate, thin, green or with green midrib, the margins often purplish; rays 15-40, r light rose, 4-6 mm. long, 0.7-1.4 mm. wide ; disk-corollas 2.7-3.4 mm. long ; style- grasa tones! acutish, 0.15—-0.2 mm. long; pappus o of 1 13-22 firm bri stles, with some inconspicuous short setae; achenes 2-nerv d. onion ig lower Boreal Zone; mountains o f San oy nardino and Inyo Counties, California. Type locality : Clark Mountain, eastern San unty. June— The var. ‘conjugans Blake. with the hairs of the etales and lower part of the stem mostly appressed or ascending, occurs in Clark and Nye Counties, Nevada. SUNFLOWER FAMILY 357 26. Erigeron téner A. Gray. Slender Daisy. Fig. 5607. Erigeron Ha ge var. tenerus A. Gray, Bot. spots = i 1876. Erigeron tener A. Gray, Proc. Amer. Ac oc 16 Erigeron hclondis Sad Bot. Gaz. 41 1: 291, dae Perennial with a taproot and branched caudex, slender, lax, 3-15 cm. tall, the herbage moder- ately o ap fel strigose. Basal leaves petiolate, with oblanceolate to obovate or rhombic, mostly blade d 2- 1 acute or eae a ~25 mm. long an mm. wide; cauline leaves few and linear ; heads 1-3, the in r 5-5 high, finely glandular and usually shortly spreading-hairy ; phyllaries im- icate, firm, somewhat thickened especially the outer, mostly greenish bro ati d k idrib, the inner with thin, often purpli argins; rays 15- ue or sometimes mm. long, 1-1.7 mm. wide; see eeptns 2.7-4.1 mm. long; inner ‘pappus of 15-30 Eriatice the outer obscure or pai 2 — ved. Roc d dry rocky soil, eal Zone; mountains of California (Cascade Mountains and Sierra Newsday” any eaten le to won eee and cont) vehi rare in our range. Type locality: Silver Mountain, near Ebbett Pass, Alpine County, California. June-A 27. Erigeron Soaks Acape A. Gray. Hairy-seeded Daisy. Fig. 5608. Erigeron poliospermus A. Gra. A 10. 1884. Erigeron poliospermus £. dlctorma: aay Antonis’ 6: 194. 1947. A form with the pistillate flowers eligu- inte Per secntah with a taproot and short branched caudex, the several stems seldom over 15 cm. tall; herbage and involucre evidently spreading-hairy and glan idular, more hairy than glandular, the 5603. Erigeron petiolaris 5605. Erigeron pygmaeus 5604. Erigeron clokeyi 5606. Erigeron uncialis 358 COMPOSITAE aos gripe more glandular than hairy. Basal ivi linear-oblanceolate to spatulate, up to 8 cm. long m. wide, the cauline ones generally more or less reduced ; heads hemispheric, mostly soli- tary, reg ‘i sk 9-20 mm. wide; involucre 3-9 m mm. high, the phyllaries equal or nearly so, green, maar with el prion midrib ssc narrow scarious margins; rays 15-45, pink or purple to deep violet, 5-14 mm. long, 1.3-3.6 mm. wide; disk-corollas (3.5)4-5.5(6.5) mm. long; inner us of 20-30 bristles ps 2- sea very "densely long-silky, the hairs sometimes obscuring ig aereas outer pappu Dry places in i plains ~ foothills, often among sagebrush, Arid Transition Zone; Washington from the Cascade Mountains eastward, ending into ma ete Idaho and southernmost British Columbia, weg soutawark to northern Harney Cuaaty, Giese, Type locality: Umatilla, Umatilla County, Oregon. pril-Jun Erigeron poliospermus var. céreus Cronquist, Brittonia 6: 194. 1947. Herbage and involucre an nsely glan dular and only sparsely or moderately hairy; plants often branched and several-headed and then with fairly well developed cauline leaves. Kittitas cay and adjacent parts of Grant and Chelan Counties, Washington. Typ locality: Rock Island, Kittitas County 28. Erigeron disparipilus Cronquist. Snake River Daisy. Fig. 5609. Erigeron disparipilus aca Brittonia 6: 194. 1947. nnial with a t and short branched caudex; stems 3-12 cm. high, with spreading, Per proo very ccasal hairs. Co at allina ~ cluster, finely hirsute, tineat or linear-oblanceolate, Pp 1 igh, spreading-hirsute and t head solitary, the inv m often glandular; phyllaries subequal or slightl bricate, acuminate, green, with o a arke drib etime narrow scar < s rays 3 100), white, or piokich in age, 5-10 a 2 ne .5-2.3 mm. wide; disk-corollas 3.5-4.3 mm. lo ong, coarser and less indu- rate than in E. pumilus; pappus of 15-28 capillary bristles and some inconspicuous outer Pee achenes 2- nerved father ensely pa hai s in the foothills and at moderate elevations in the ntains, Arid Transition Zone; extreme la Sethe. een | Wartoctas and iene dh <3 or sor Idaho Hapa waa in be vicinity = the _ River to Owyhee County. Type locality: 14 miles west of Clarkston, Asotin County, Washington. May—Jul 29. Erigeron pumilus Nutt. subsp. intermédius Cronquist. Hairy Daisy. Fig. 561 Erigeron hispidissimus Piper, Contr. U.S. Nat. Herb. 11: 565. 1906. Erigeron pumilus subsp. in dermgilias Cronquist, ais ia 6: 180. 1947, Perennial (sometimes short-lived) with a eatorn at 2 yes or branched caudex; stems c tered, mostly 2-5 dm. tall, vey opus, only e than 1.5 mm. thic at the base poe aring 5 to many heads; herbage Sronsty” spreading hineute Sa sometimes a little Senge. Leaves oblanceolate, oe to 8 cm. cit an mm. wide, cauline as well as basal; heads mostly seni ic, the disk mostly 7-15 mm. sigan involucre 4-7 mm. high, spreading-hairy aot very row, a n, wi rib cad sometimes very tied scario gins; rays mostly 50-100, blue, pink, or white, 6-15 mm. long, mostly 0.7-1.5 mm. wide; diel: a 3.5-5 mm. long, slender, the limb pale and indurated below, the indurated portion. often finely oshesuleans pappus of 13-20 ee pit some sho y or vi og —, = the foothills, valleys, and plains, often among sagebrush, Arid Transition canes east of the Cascade Mountains fro: ashington to northwestern Montana, southward to northern Orego: an: 2 te ore com- monly to 2 pn ‘California and adjacent Nevada. Type locality : near Lewiston, Idaho. ies — rigeron pumilus subsp. intermedius var. gracilior Cronquist, Brittonia 6: 180. gal A tg oa smaller and more slender than in typical subsp. intermedius, 0.5—-3 dm. tall, the larger stems seldom hick at the base and rarely with as many as 5 heads. Central and southern Idaho and adjacent anise ie: en atria and vost ek ag northeastern California, and adjacent Nevada. Type locality: United States Sheep Farm, unty, I Erigeron pumilus subsp. concinnoides oi bongs eg eos G: 181, 1947. mS ea concinna Hook. & Arn. Bot. taro, > 350. 1838; Erigeron concinnus Torr. & Gray, Fl. N. ened 7 Prag E. concinnus var. eremicus Jepson, Man. Fl.. Pl. Calif. 1057. 192 5.) Routes imilar to " gra n habit or a get bene sebost: rays nearly always colored; indurated , tata of the disk sneer pest hard ubeeuiaut: inner pu of 7-12(15) — bristles. the outer of a neeerety broad ware Dry n (but nga desert) hrs in the foothills and at moderate elevation he mountains; sou wth n California in Inyo and San Bernardino Counties; also peer fas € Idaho and wenea cisteline pear to Bearers New Mucies Fae southern Neves a. Type locality : Kyle Canyon, Charleston Mountains, Clark County, Nevada. 30. Erigeron engelmannii A. Nels. var. davisii Cronquist. Engelmann’s Daisy. i it: Fig. Erigeron en =" davisit Cronquist, Leaflets tating Bot 167. Erigeron engelmanni var. davisii Cronquist, Vasc. Pl. Pac ee ee sie ies: Erec Sih ee with a taproot rite shortly aca caudex, 10-30 cm. tall, ay sonarrel Basal leaves Gient-ohlangtalie up t cm. long and 4 mm. wide, with some coarse, ing. marginal hairs toward the base; Sealine leaves linear and reduced; heads solitary or ie ot gv rm peduncles ; involucre (5)6-8 igh, spreading-hairy and usually finely glandular, the phyllaries subequal, narrow, alg usual y with bro rib and pale margins an corollas 3.3-4.5 mm. long; ra s 35-55, mostly white, 8-14 mm. long, 4, 1-2: 2' mi oe Meeeh pappus yt De goad bristles stg sid inconspicuous outer setae or narrow scales ; ecicnes -nerved, iry. Spa meadows, and open hillsides in the foothills and at moderate elevations in the mountains, Arid Tra sition wis bad western Idaho Nags a Idaho, aod =o Erect Bit cone Type | ocality : Whitebird peony gern Comets Idaho. mg) 7 eegee cal E. engelmannit i ll parts and occurs in Wyoming, Colorado, Utah, and southeast SUNFLOWER FAMILY 5607. Erigeron tener 5610. Erigeron pumil b: 5608, Erigeron poliospermus 5611. Erigeron engelmannii 5609. Erigeron disparipilus 5612. Erigeron multiceps 359 360 COMPOSITAE 31. Erigeron milticeps Greene. Kern River Daisy. Fig. 5612. Erigeron esticee Greene, Pittonia 2: 167. 1891. Perennial from a stout ps ee short branched caudex ; stems slender, branched, up to 2 dm. tall, soe with a sed hairs except under the heads where the hairs are spreading. Leaves appressed-hairy, t he “teat Bbianteciste to obovate, up to 4 cm. sot he an shorter than eg petiole ; cauline leaves linear or the lower oblanceolate ; heads several, the inv e = m. hi finely glandular and moderately or sparsely hirsute with short simeading hairs ; olla es atigneee with slightly darker midrib and sometimes paler ome the outer a little shorter nae evidently narrower than the relatively broad, thin, inner ones; rays about 75, lilac or hile 5-7 mm. long, 0.6 mm. wide ; disk-corollas about 2.5 mm. long, the limb white and strongly pert near the base, flaring a above ; style -appendages less than 0.1 mm. long; pappus double, the inner of 5-8 slender bristles, the outer of evident and sometimes more or less tinited “short sable: lichines Gravelly spots near river banks, mostly Arid Transition Zone; north fork of as Kern River, Tulare County, California. Type locality: near north fork of the Kern River, near Kernvi lle. June 32. Erigeron divérgens Torr. & Gray. Diffuse Daisy. Fig. 5613. ron divaricatus Nutt. Trans. Amer. Phil. Soc. IT. 7: 311. 1840. Not Michx. 1803 1 Taprooted biennial or pees lived seecrcuiak 1 7 dm. ‘teal freely branched, # herbage covered with sp is ot = ostly well under 1 mm. long. Basal edi petiolate, with oblanceolate or wana blade u Som oun usually eel ‘cauline leaves numerous, smaller, often linear ; heads mostly n ous, the involucre 4-5 mm. hi gh, glandular aad spreading-hairy ; phyllaries light yellow- conte with broad brown sidveli, the outer a little shorter and evidently narrower than the eels broad, ge -margined, inner ones; rays 75-150, blue, pink, or white, 5- 10 mm. long, 0.5-1.2 mm. wide, or rarely much reduced ; disk-corollas 2-3 mm. long, the limb white and strongly indurated toward ihe base ; style-appendages less than 0.1 m m. long; pappus double, the inner of 5-12 y fragile bristles, the outer of conspicuous, re, shor scales; achenes 2-nerved ed or sometimes pe: ely 4-nerved, the nerves usually inconspicuou Dry and waste places in the valleys, aearsagaed and lower mountains, Se uaitis in sandy soils, Upper Sonoran and Arid Transition Zones; southern British Columbia and Montana to Lower California, Texas, and Okla- homa. Type locality “Rocky Mountains pg plains of Oregon.’”’ April—July, sometimes continuing until fall. 33. Erigeron calvus Coville. Inyo Daisy. Fig. 5614. Erigeron calvus Coville, Proc. Biol. Soc. Wash, 7: 69. 1892. Taprooted biennial promt 1 dm. tall, branching widely from the base, the herbage hirsute with long spreading hairs and the stems finely glandular especially under the heads. Basal leaves nu- merous, with oblong to shsate blade 1-1.5 cm. long tapering to a ectibte twice as long: se leaves reduced, oblanceolate or spatulate; he a terminating the eae dis wide ; involucre abou mm. high, coarsely spreading-hirsute and finely andular “phyllaries moderately imbricate, es greenish, sometimes with chaffy margins stillate flow pba but the ligules wanting or weap ges and much shorter than the > disk- oe 4- ong; st vlesappendae 0.15-0.25 mm. long; velpows i coed oor ‘of 15-20 firm sandat ‘Basties the outer of several relatively (eae firm setae; achenes 2-nerved. pper Sonoran Shy Inyo — untains, California. Type locality: foot # — Shae about 4 miles north of Keeler, Inyo County. tay 34. _ aphanactis (A. Gray) Greene. Basin Rayless Daisy. Fig. 5615. Erigeron nn r. aphanactis A. Gray, Proc. Amer. Acad. 6: 540. 1865. Erigeron Siok tis ip tes , FL. Fran. 389, 1897, Perennial with a taproo’ ve branching caudex, mostly 1-3 dm. tall, the henge densely spreading-hairy. Basal wows linear-oblanceolate to spatulate, long-petiolate, up to 8 cm. lon and 6 ide, persistent or deciduous, the cauline similar but nearly sessile, or linear and re- ; heads several or solitary, yellow, sometim rning brownish in age; inv 6 mm high, glandular and spreading-hirsute, the phyllaries subequal or slightly imbricate, slender, acu- minate, green reenish brown, with thick idrib; pistillate flowers present, tubul and eligulate or sometimes bearing inconspicuous ligules s et pany: the disk; disk-corollas 2.8-4. 1 the limb commonly with a firm, whitish, often pu ent portion below the middle; mm. long, style-appendages 0.1-0.25 mm. long ; pappus double, shed inner of 7-20 bristles, the outer of evident, sometimes narrow scales; achenes 2-nerv Mostly in hot dry places in the foothills and deherte s, Arid Transition Zone; southwestern Idaho and so uth- to Colorado, Arizona, and southern California, largely or was. races om fae, ce a rigeron aphanactis var. congéstus (Greene) Cronquist, Brittonia 6: 177. 1947. (Erigeron iauaies Greene, Leaflets Bot. Obs. 2: 218. 1912.) Plants about 1 dm. ta a hed less, essentially scapose; corolla-lobes and often the upper part of the limb turn ning reddish or purplish at in age. Dry places at moderate to high elevations im ne San Bernardino Mountains = Snes Type ow Peg Gold Hill, Bear Valley, San Bernar- dino Mountains, San Bernardino County. Jun SUNFLOWER FAMILY 361 353 Erect chrysopsidis A. Gray. Golden Daisy. Fig. 5616. Chrysopsts hirtella DC. Prod. 5: 237. 1836. Not Erigeron hirtellus ite 1836. Erigeron = aoe var. ages A. — te oc. Amer ey ad. 16: 1880. Erigeron chrysopsidis A. Gray, Syn, FI. er. 12: 2 1884. Erigeron curz ore Piper, Bull. Tairey meat 27: 396, ey as to type only. Perennial from a taproot and ns meer 2 caudex, 6-16 cm. tall; a spreading-hairy or the pubescence of the leaves occasionally partly appressed. rfpeese all or n arly all in a basal cluster : : : a sw (2)3-9 cm. long =F -3 mm. wide; head solitary, hemisphe i : ucr : m. high, s pase reading-hairy and sometimes slightly glandular, the phyllaries sub- qual, ow, het or gree ish at bast 4 Rowe the midri en Wi li margins, gradu ate Soupinate - pistillate rca a ne the rays bright yellow, . long - disk-corollas mo ong; pappus o L 15-25 rel. bristles, com- and 1-2.5 mm. wide stly ¢ perth with a fw short, tendo: incon ga baler setae; — Dry open places, often sagebrush, exten to a litt r 4, As n the ponderosa pine a ‘arid Transition a east of the ‘gate ot natn in ‘athata Creme fae ‘Grant (and Crook?) Counties a ward, barely extending into southeastern Washington. Type locality: Columbia River. May June (Jul eireck chrysopsidis var, brevifélius Piper, Bull. Torrey Club as 395. 1900. Smaller in all respects r 985 pol asi = Sah ee i = 6(9) cm, tall; basal pri 1- oO ¢ . long: involucre 4-5 mm. high; disk wide; dis rollas 3-4 mm. long; pu ubescence of leaves often more or less appressed. Wallow a Moun- Bs or Oba pat Ronen altitudes : typical E. chrysopsi sidis. Type locality: Wallowa Mountains. Erigeron eharecpsidls subsp, austiniae (Greene) Cronquist, Brittonia 6: 196, 1947. (Erigeron austiniae Greene, Erythea 100. 1895.) Pistillate gin ers present but hae tars wanting or at and inconspicuous; other- wise resem mahi total E, chrysopsidis or smaller and more peed Be Harn ney, and Malheur Counties, Oregon, southward to the vicinity of Mount Tassen California, yer eastward to Twin Falls County, Idaho, and Elka County, Nevada. Type locality: Davis Creek, Modoc County, California. 5616. Erigeron chrysopsi 362 COMPOSITAE 36. Erigeron piperianus Cronquist. Piper’s Daisy. Fig. 5617. Erigeron curvifolius Piper, Bull. Torrey Club 27: note 1900, pro max. part., excl.-type. Erigeron piperianus gt a Brittonia 6: 197. 19 nd basal, linear or ax and usually curv pt lon d1.5 de, hispid ciliat gins, pile ially elow, a appre ssed- hai he e, the lower with con- spicuously enlarged, wh ph iadure’ d base; heads ni or few, small, t i mm. wide; in m gh, spreading-hirsute with long white hair , the phyllaries subequal or imbri- ate, Loin or greenish, with darker thickened midrib, acuminate at the tip; rays 0, yellow ong, .8 mm. wide; disk-corollas mostly 2.8-4.2 mm. long; pappus of 15-25 ceisie and often some short outer setae ; achenes 2- nerved. open places, often ng sagebrush, Arid Transition Zone; Columbia River plains of Washington from Douglas Pe Ay to Benton Somete Type Secular: cneide oe Soap Lake in the Grand Co ulee, Grant County, Wash- In 37. Erigeron blodmeri A. Gray. Bloomer’s Daisy. Fig. 5618. Erigeron bloomeri A. Gray, Proc. Amer. Acad. 6: 540. 1865. Bipeon filifolius ieamar K: Nels. Bot. Gaz. 54: 413. 1912. Perennial from a taproot and much-br anched caudex, 5-15 cm. tall, the ef ncaa finely white- strigose. ee ves all or nearly all in a basal cluster, linear, 2-7 cm. long, nf wide; hea solitary, turbinate to subhemispheric, the disk wide; involuc ee 5-1 ches: “high, ptt to villous especially toward the base, — phyllaries gens acute, firm, imbricate, usually green pistillate flowers wanting ; disk-corollas 4.5- mm. long; style- appendage’ acute, mm. long ; pappus-bristles 25-40, gen alee unequal, sometimes with a few inconspicuous outer setae; achenes 2-nerved, glabrou s below, short-hairy abov ee rock laces in the mountains a fothils, Arid Transition and Canadian Zones; Kittitas adie: Washington, to Siskiyou (Marble Mountains) and Nevada —- Be cnet eastward to Lemhi and Cassin Counties, Idaho — Elko and Lander Counties, Nevada. Type 1 y: near Virginia City, Storey County, Nevada. June— Erigeron Seon var. nudatus (A. Gray) Cronquist, ear 6: 199. 1947, Cees nudatus A. Gray, Proc. Amer. Acad. 20: 297. 1885.) Stem and leaves glabrous or finely and sparsely strigose; involucre glabrous or very nearly so; style-appendages acutish, 0.2-0.3 mm ene Ccrpeatins areas in the mith region of southwestern Oregon and adjacent California. Type | ocality Boden 1 a page? Se Specimens of Erigeron bloomeri var. pibens Keck, Aliso 4: 105. — gooey ‘con ee Aaa ani E omeri var. nudatus in being soft- pilose throughout with -cpharerheg hairs. In the Marble Mountains, Siskiyou oe Cali- fornia. Type locality: King’s Castle, Marble Mountain an Ea gerap linearis (Hook.) Piper. Desert Yellow Daisy. Fig. 5619. Diplop p li Hook. Fl. Bor. Amer. 2: 21. 1834. Erigeron m peuteplie ane Gray, Proc. Amer. = or 16: 89. 1880. eset luteus A. Nels. Bull. Torrey oe 2: 1900. Erigeron linearis Siar Wonk 2. OS t. Herb. aa 567. 1906. Brive eron yakimensis A. Nels. Amer. ate, Bot. 23: 268. 1936. Perennial from a stout taproot and branched caudex, 5-30 cm. tall; Beare finely gray strigose ; tases of stems and of basal leaves conspicuously i indurated and Ye) oes enlarged, li rl , stramineous to sometimes purplish. Tove inear or nea 9 cm. long, 0 mm. wide sal and cauline or nearly ; heads solitary or few, the disk most! ide; in- volucre 4-7 m h, strigose-villous and etimes finely glandular, the phyllaries subequa imbricate, green or g ineous, ont. th da As idrib, rm, kened on the e inner sometimes purple-tipped ; or 5, yel 1 mm. long, mm. wide; disk-corollas 3.5-5.3 mm. long; pappus of 10-20 oo i sad some short, "often narrow, outer scales ; achenes 2-nerved, short-hai Dry often rocky soil from reste plains and foothills to moderate elevations in the mountains, often with sage- brush, Arid ‘Franshion Zone ilern British Columbia through eastern Washington and Oregon to Yosemite National Park, A pS ogg To eastward to Park County, Montana, Yellowstone National Park, Wyoming, and Elko - ounty, Nevad col ity: ‘‘near the ‘Priest’s Rapid’ of the Columbia, and also on Lewis and Clarke’s River.’ May Je uly (Aug.) 39. Erigeron elegantulus Greene. Volcanic Daisy. Fig. 5620. eee sagen tains Me rem Erythea 3: 65. 1895. gantulus J. T. Howell, Leaflets West. Bot. 1: 205. 1936. Pecenall stants a taproot and much-branched caudex, 3-15 cm. tall; herbage poy or mod- erately gray-strigose ; bases of stems and basal leaves ped cartes Bre b mooth, shining, somewhat narrowly linear, one y 1.5-6 cm. long and so 0.5-1 mm a sk ; involucre 3.5-5 mm. high, stri — -_ glutinous ; anc arcni imbricate, the short outer ones narrow and rather firm, the inner ones broader, with scarious and usually erose margins, acute or abruptly acuminate ; rays 15-25, py aesy or pink to ciennee white mm. long 8 wide ; disk-corollas 3.4-5.0 mm. long; pappus o sle ‘a bristles, eae ath a few slender, outer, sho above t ut. Open, often rocky ses in the plains and foothills, canna in basaltic or volcanic recks, Arid Transition and Canadian Zones; er, Union, and Klamath Counties, Oregon, to ray, Siskiyou, cg "Lassen Counties, California. Type locality : Dixie Valley, Lassen County, California. June—Jul SUNFLOWER FAMILY 363 5617. Erigeron piperianus 5619. Erigeron linearis 20. Erig rT, 7 g a | 5618. Erigeron bloomeri 40. Erigeron barbellulatus Greene. Shining Daisy. Fig. 5621. Erigeron barbellulatus Greene, Erythea 3: 65. 1895. id volucre 5 m. and somewhat glutinous ; Sander subequal or slightly imbricate, long-acuminate, usually a litile thickened on the en: ys 15-35, blue or purple to white, 7-12 mm. long, 1.5-3 mm. wide; disk-corollas 4.4-7.0 m eo et pappus of (50) slender bristles, neg or raza with a few slender, pa 4 short Baie - achenes 2-nerved, sparsely strigose, or glabrous Rocky places at moderate to fairly high elevations in the mountains, Canadian and Hrudsonian Zones; Lassen County to Fresno County, California. Type locality: Silver Lake, Exanea ‘County. May-Aung. 41. tgs ast filifolius Nutt. Thread-leaved Daisy. Fig. 5622. filif r. Amer. 2: 21. 1834. Chr reeds canescens DC. Pr i: ee "ta. 1836. Erigeron filifolius Nutt. Trans, Amer. Phil. Soc. IT. 7: 308. 1840. Perennial ed : Re and branching woody cau 1-5 dm. tall; herbage finely on the stem densel t the base. Leaves linear or aes ah 1-8 cm. long, grit in di wide, well distributed along the stem but the basal ones peas nly longer o 4 heads usually several, the disk 5-1 mm, wide ; ra e 4-6 mm. high, beaut Sy villoas-str- r finely glandular or both; phyllaries firm, som arg thidkeied-Ga e back, subequa pabticae ouatty greenish, often ‘with darker midrib ; pitti 15-50(75), blue = Gk or ‘white, i "7g ik 364 COMPOSITAE . long, 1-2 mm. wide; disk-corollas 2.54.4 mm. long; style-appendages 0.1-0. 15 m m. long; pappus Pg 20-30 slender white bri atles, porate ee a few very slender and i ieditspicnous short, r setae; achenes 2-nerved, hairy or subglabro pits y, vig sandy ary in the valleys i foothills, pate with sagebrush, Arid pare ye Zone; south- ern British Columbia and extreme northwestern Montana through eastern Washington and Oregon to Sierra County, California, and adjacent Nevada eastward through the Snake River plains to Rexburg, Idaho, and thence southward to Logan, Utah. Type locality: “‘In the Rocky Mountain range, in Oregon. ”” May-July. Erigeron filifolius var. robtistior M. Peck, Proc. Biol. Soc. “aby pe ot, Rie 4: a stout; head ire solitary, the disk 12-18 mm. wide; involucre 5-7 mm. high s 50-1 6-1 long; disk: corollas 3.7—5.5 mm. long; style- Sieiadnices. 0.15-0.3 mm. long. Kittitas ‘Couets, Waskicctom "southward to Wasco and Gilliam Counties, Oregon. Type locality: Columbia River near Rowena, Wasco County, Ore 42. ee hate lassenianus Greene. Lassen Daisy. Fig. 5623. Greene, Fl. Fran. 389. 1897. piesatte lassenianus var. deci ciens Cronquist, Brittonia 6: 171. 1947. A discoid form. lender perennial with an evide ident slender taproot and a simple or subsimple crown or sh ‘ ge fin r m. g. Basa te, taper- ing gradually to the slender petiole, up to cm, long and 5 mm. bog cauline leaves several, uced, linear, cocaine mele peeved heads usually several (up to 20), occasionally soli- pate the disk 5-11 m wide; e 5-6 m m. high, glandular and spreading abe te; wre llaries scarcely to eitdenity sdibrieate: a or greenish, sien oe -margined, acuminate; rays 10-25 blue or pink to sometimes white, mm. —2.0 mm. wide, or sbedsionalty¥ wanting; disk- machen 3-4 mm. long; pappus of 15-25 ireoile bristles ead a few short outer setae; achenes 2- nerved. Dry places at moderate elevations, Arid Transition Zone; Butte and Plumas Counties to Eldorado County, California. Type locality: Mount Dyer, Plumas County. June-Aug. 43. Erigeron corymbésus Nutt. Foothill Daisy. Fig. 5624. Erigeron corymbosus Nutt. Trans. Amer. Phil. Soc. II. 7: 308. 1840. 1 with a taproot and not much-branched caudex, 1-5 dm. tall, ec: or teed u y : gradually or strongly reduced; heads 1-16, the disk 7-13 mm. wide; involucre ie canescently villous-hirsute with short subappressed hairs; phsliniaes saraeetak 3 mbric only — or rays 35 thickened on the back, green or tan, with darker midrib and purplish, ‘vals i i m. 1 tips; r. —65, deep e or occasionally pink, 7-13 m ong, 1.2-2 mm. w Serolies (3.0 )4.0- mm. long; style-appendages ate, joceey. .2-0.25 mm. long; pappus of 20-. m bite and some evident, short, outer setae rrow scales; achenes 2-nerved. Ope lly dry places, often among sagebrush, Arid i ansition Zone; scathien British Columbia to cen- tral Fete pe gece lg A sete east of the Cascade Mountains eastward to western Montana and western Wyoming. Type locality: ‘‘Rocky Mountains, towards the Oregon,”” June—July. 44. soci as caespitésus Nutt. Gray Daisy. Fig. 5625. ook. Fl. ~— ape 2: 23.: 4834. ee epappus pronifor Hook. loc Eriger espitosus Nutt. Trans. ‘Amer. Phil. Soc.t1. 7+ 307. ‘on cane. sceas Tor r. & Gray, Fl. N. Amer. 2: 179. 1841. te Hd & Arn Erigeron caespitosus var. aaldiieias Torr. & Gray, loc. cit. — pretditirice he 1834. Erigeron subcanescens Rydb. Bull, Torrey Club 24: 294. Perennial with a stout taproot and stout, esa branched caudex, sometimes also creeping below ground; stems curved at the base, 5-30 cm. high; herbage densely pubescent with short spreading hairs. Basal leaves triple-nerved ( Ghscitihy so in depauperate apetinens): oblanceolate or spatulate, usually rounded or obtuse at the tip, up to 12 cm. long a mm. wide; cauline ova ong to li : ic hack baceous ; rays 30-100, blue, pink < or white, 5-15 al: "Be g, 1-2. Sih mm. aide iek-covollds 3.2-4.4 mm. long; style- appendages 0.1 aw ee mm. long; neat of 15-25 firm bristles and some evi- dent, a setose scales ; achenes ape Dry, open, often rocky pre from sa “oti fe ae elevations in the mountains, Arid Transition and lower Tiwaat Zones; — —— and Yukon through British Columbia and extreme eastern Wash- ington to. Idaho. , Uta ah, Arizo ew Mex mei posinkoet to Saskatchewan, North Dakota, a nd Ne braska. Type locality: ‘ ‘summits of dry hills in a Rocky Mountain range, on the Colorado of the West.” Ju 4. er nevadincola Blake. Nevada Daisy. Fig. 5626. A. Gray, Proc. Amer. Acad. 8: — sgt = Wedd. 1857. maa nevadincola Blake, ear: Biol. Soc. Wash. 35: Coarse perennial with a strong taproot and en: or slightly branched caudex ; eats usually Sorsicheet at a 10-30 cm. high; herbage strigose or short-hirsute with app pressed or ascending hairs. 1 triplesnerved, ae “ sometimes : Fie tapering gradually to the rile up to o e; cauline leaves few and a ruptly reduced ; a d soli- 5623 ~ PAS AS ae 4 = aS Zé \ nia: “fr Vi VA af SUNFLOWER FAMILY 5621. Erigeron barbellulatus 5622. Erigeron filifolius 5623. Erigeron lassenianus 5624. Erigeron 5625. Erigeron caespitosus 5626. Erigeron nevadincola 365 366 COMPOSITAE villous-hirsute, scarcely or not at all sareemgt pe kage subequal, broad, dark green, usually mew bas somewhat indurate and heownish at the _ the tips scarious, merely acute; rays 20-40, white or blue, 5-11 Frances ong, 1. mm. wide; is lisk- corollas mostly 5-7 mm. long; style- ape acute, 0.3-0.4 mm. lo ong; pappus of 25-40 rather coarse, firm bristles and a few slender and 1n- conspicuous, short, outer setae; achenes 2-nerved, ovientie hairy above, usually glabrous or gla brate below open shorn in the foothills and at moderate ag pte in the mountains, mostly Arid Transition Zone; Pints *“Comaty to Mono County, California (east of the Sie crest), eastward to Elko and Nye Counties, Ne- vada. Type locality: near Virginia City, Nevada. May—Aug. 46. ae eatonii A. Gray var. villésus Cronquist. Eaton’s Daisy. Fig. 5627. Greene, Pittonia 3: 293. 1898. Ee aoe on iueikons Howell, eS N.W. Amer. 319. 1900. atari eatoni subsp. villosus Cronquis' w ‘Beitcaia 6: 172. 1947, Erigeron eatont var. shiteiod Zicdauist, Vasc. Pl. Pacif. Ninth. Se 37599985; Peren nial with a taproot and crown or short, slightly branched caudex, 5-30 cm. tall, the stem oer ecumbent at the mostly reddish purple base; herbage strigose to vile. hirsute with “apprenid or closely ascending hairs Hej leaves well developed and conspicuous, narrow, acute, tapers tre os to the petiole, up t o 15 cm. long and 10 mm. wide, triple-nerved (occa- sionally linear and 1-n in depauperate see): Cauline leaves several, more or less one codices as hice Tina or a s 7, m r less naked-pedunculate, the disk 8-15 wide ; involucre 5- ele oe ae eyiscaly se cihevillbine: iraute repel” Te ward the base; phyilaries subequal, green, oft sai purple-tipped and ee cy with a narrow da - midvein; rays 20-50, mostly white, ate light blue, 5-10 mm. long, on wide ; disk-corollas mostly g; style-appendages obtuse to acuti a 0.1-0.2 m ai ng; pappus of 15-25 fragile 5 mm. lon bristles with a fe ew very fine and inconspicuous, short, outer setae ; a mostly 2-nerved, shortly villous-hirsut te. Open place the mountains and foothills, mostly Canadian Zone; east (and occasionally just west) of the Coane: Moeniial summit in Oregon, extending northward to the We natchee Mountains of Washinavon and east- ve through the Blue ircxmtaine region fapiuding « extreme southeastern ees tcn) tot = ntains of central aho (as far ea: ‘ : nt n bri ern, occurring from southwestern Montana to Célor ado and Arizona. Erigeron eatonii A. Gray var. ai ae ne Cronquist, Vase. Pl. Pacif. Northw. 5: 175. 1955. (?Erigeron sonnei Greene, Pittonia 1: 218. plantagineus Greene, op. cit. 3: 292. 1898; E. eatont subsp. plantagineus Cronquist, Brittonia 6: 73. eR ) Pubescence of the leav ves, stems, and a achenes averaging s} coa wok Bas PPE Se i i i e 47. Erigeron decimbens Nutt. Willamette Daisy. Fig. 5628. Erigeron decumbens Nutt. Trans. Amer. Phil. Soc. II. 7: 309. 1840. Perennial with a taproot and crown or short, slightly branched caudex, mostly 1.5-5 dm. ee the stems decumbent at the mostly reddish purple base; herbage strigose. Basal leaves and som r most of the only gradually reduced cau ae ones triple- -nerved, the sal ones up to 25 cm. eae k neul de ; involucre 3.5-6 mm. high, more or less hirsute and usually obscurely visct phyllaries slightly or scarcely cp tale bi or dark pilin to brownish; rays 20-50, blue or a 6-12 mm. lon gt —2 mm. wide; disk-c as 2.5-4.0(4.5) mm. long; style- appendages oper acutish, 0.1-0 ccarettig pappus of ie fragile ‘pristles, occasionally with a few very fine a oe ie outer setae; achenes 2-nerved, finely hairy. places, Humid Transition Bese: Willamette River drainage of western Oregon. Type locality: “Rocky Mousisins, towards the ty rt); involuere commonly greener than the gorttods varieties; rays gen erally blue, averaging bourse numerous "hy oh typical E. breweri. on and moderate elevations in the moun- tains; western Nevada and eastern and southern California. Type locality: Hawthorne, Big Indian Canyon, Mineral County, Nevada. Erigeron breweri A. Gray var. jacinteus (H. M. Hall) Cronquist, Brittonia 6: 284. 1947. (Erigeron jacinteus H. M. Hall, Univ. Calif. Pub. Bot. 1: 127. 1902.) Prostrate or ascending plant; stems oe 1 dm. long or less; n pubescence averaging longer and finer than in the other pasitiane leaves not ov bo 2 mm. long, Boat ve = Bs Nowa pee solitary, at large for the species; phyllaries often purplish especially a she: ireine and tips; light-colored. Talus slopes and rocky ridges at bn elevations in the San into, San Antonio, San Gabr iel, and San Bernardino Mountains, California. Type locality: near Tauquitz, ea Ja acinto Mountains, River- side — California. nm breweri A. Gray var. élmeri (Greene) Jepson, Man. Fl. Pl. Calif. 1056. 1925. (Aster elmeri Gicoet Gittonia 2: 170. a Erioeras elmert Greene, Fl. Fran. 393. 1897.) Very osseous prostrate or as- cending plant not ove Boag es: high; herbage subglabrous or sparsely appressed-pubesce By dd relatively distant, not over 15 m mes a reek longer than the internodes; heads solitary or few, naalt + r the specie involucre giendas : tan ‘with oe without a waliignrones: subapical. green spot; rays mostly pink or lavender, es = darker Cliffs rock crevices at high altitudes in the mountains: from Yosemite Na- : onal Par i pee oia National Park. 4 Cuiifornin Type locality: Drea Canyon of the Tuolumne River, Cali- ornia. 55. Erigeron folidsus Nutt. Leafy Daisy. Fig. 5636. igeron foliosus Nutt. Trans. Amer. Phil. Soc. IT. 2 309. 1840. aah douglasti Torr. & Gray, Fl. N. sagt “# 177. 1841. Erigeron mariposanus Congdon, Erythea 7: 30. rigeron striatus Greene, Bull. S. Calif. Utes iA Erigeron foliosus f. quist, Britt aa a 1947. A broad-leaved form. Perennial sith a stout taproot and superficial or nearly superficial root-crown ; stems erect or nearly so, often a little woody at base, 1-10 dm. tall, usually waits nched at base except in var. confinis and p. Nota var gf Seton the base S aesne som atat rhizomatous but if so then pani very apenie hig rey y stout and more than 1.5 m a tack. or both;-herbage varying from t rather. r ve mm, | n s ite to acute, 0.05-0.z mm. long; pappus do: uble. “the inner of 20-33 sordid barbellate a the outer of a few inconspicuous setae; achenes 2-nerved, sparsely hairy or sometimes Dry, often rocky places or in ve d, mostly Transition Zones; sovth m through California to Lower California. Type locality: near Santa Barbara, Santa Barbara County, tonto April-July. The foregoing description is p Sis Src = cover all the varieties of the speci SUNFLOWER FAMILY 371 Key To VARIETIES Plant 25 cm. high or usually less; leaves very crowded, the internodes of the middle part * the — Seeraning 3 mm. long or usually less; chiefly i in the Klamath region. r. confint. Plant eau over 25 cm. high; leaves less crowded, the internodes often averaging more t wae " mm. Stem eipbrows to more or less strigose or hirsute-pubescent with appressed hairs, or the hairs near te base of stem spreading; leaves glabrous to variously hairy Hind: ego to narrowly oblong, the larger — mostly over 1.5 mm. wide or if occasionally less than .5 mm. wide then flat and without subconical hai Leaves did often also the stem more or ss ‘aes subtouical hairs often present; chiefly west of the Sierra Nevada and sited of San mien var. foli iosus. Leaves and stem subglabrous only s y strigose; year gene hairs _— “ag soe Bi a of San Francisco and a sang ational ad northward to Ore r, har’ Leaves suneae filiform or narrowly linear, 1.5 mm. wide or less, pail ‘Gthes ‘gaeaa ot some 0 nal hairs enlarged toward the base and subconical; Fresno and Monterey: Pacaties a tenophyllus. Stem = leaves deauits and finely acticin or hirtellous with distinctly spreading hai s glabro ap osha bubtr ules i ith fine curly hairs; sand dunes near the seacoast in Santa Barbara and San s Obisp tie r. blochmaniae. pee more or Kyi bey kee hoi lous with short, generally straight paren ee and dry moun- tains of the interior of southern California, extending into the Sierra Neva “ip var. covi Erigeron foliosus var. hartwégii (Greene) Jepson, Man. Fl. Pl. Calif. ee 1925. (Erigeron Lenten Penne i age: Sr 2hr agi erly wi nee 9 ornia thro _ the Sierra Nevada and adjacent easterly 62. Erigeron acris L. var. asteroides (Andrz. ex Bess.) DC. Northern Daisy. Fig. 5643 Erigeron bah aiepale Muell. Fl. Dan. p/. 874. 1782. Erigeron asteroides Andrz. ex — Enum. Pl. Voth. $3; 1822; iY ot Erigeron acris var. phe cena rere orges Fl. 1: 562. 1861. Erigeron — var. kamtschaticus Hara, Rhodora 41: 389. 1939. Biennial or Mo a with a short, simple or slightly branched caudex, 3-8 dm. tall; herbage subgla i preading-hirsute, becom ming evidently glandular in the inflorescence. = sal Cees mostly eGle ABs onc, up to 15 cm. long and 12 mm. wide, seldom much tufted ; cauline eaves most * . * * . m. —. sometimes wi ith a few inconspicuous, short, outer setae; ocky pla n the mountains, Boreal ‘ naar Color, Fo sctnan — a rh Sh agp De rience anata ora to northern Oregon, northern acris var. débilis A aie Syn. Fl. N. Amer. 12: 220. 1884 nn Eri rigeron nivalis Nutt. Trans. Amer. Phil. Soc, 11.7: 311. iBs0: E. jucundus Greene, Pittonia 3: 165. 1897 E. debilis Rydb. M N. ° y' em. Bot. Gard. 1: 408. 1900.) Plants 0.2-3 dm tall, the stem often curved at the base; leaves relatively wider than SUNFLOWER FAMILY 375 nil qi we : \ } 5641. Erigeron strigosus 5642. cet a snerageeacaltaet 5643. Erige snorum in var. asteroides, the basal ones generally Custesed, Jad the cauline ones often much reduced; gece few or solitary; rays generally pinkish, a little hash gl than asteroides (up to 4.5 mm. long an .55 mm. wide) and more evidently surpassing the pappus; ye ‘pistillate yc often few. Alaska to northern "California (Mount 'S a and ie ase Comren. Type locality: Woodruff’s Falls west of Upper Marias Pass, Rocky 0 92. Lesa Cham. Linnaea 4: 203. 1829. ‘ nds. nate, entire, toothed, or pinnatifid, those of the branches sessile and bractiform above. Heads solitary or clustered at the tips of stems and branches, in some species spicate in the axils of leaves or sessile among the basal leaves, homogamous, but the outer flowers often enlarged Pe palmately 5- cleft and reflexed. Involucres cylindric, turbinate, or cam- panulate, the phyllaries graduate in several series, usually much appressed. Corollas laven- der, pink, yellow, or rarely white. Anthers ro ounded at base, bearing a terminal appendage a . Style-branches with subtruncate or triangular, hairy appendages, these often bear- ng a terminal cusp. Achenes turbinate to obconic, silky-pilose ; ee many bristles r awns, sometimes connate in phalanges, or r uced to slender palea tan, br ish, or reddi sh. [Name in honor of the Lessings, a German faintly of iisaéte and authors a * Based largely on the earlier published work of John Thomas Howell and later discussions with him. 376 COMPOSITAE _A genus of 11 or 12 species, except for one species which extends into Arizona, limited to California. Type species, Lessingia germanorum Cham. Plants depressed or dwarfs; tips of inner phyllaries white and cartilaginous. 5. L. nana. Plants not depressed or Seaita: tips of inner ys fog herbaceous or scario: All corollas yellow (heads wit —_ orollas purplish or white in L. Ruauless involucres campanulate or, if turbinate, broad in relatio aden gth. Style- sean aa subulate, “Het ing a long cusp, nearly or quite as long as the stigmatic portion; plants of Mojave Desert (occurring A in S Means Pinos and Cuyama Valley regions). z Style- apreear less than half the length of the stigmatic porti t (longer in oi) forms of L. glandulifera); plants not of desert ene ppabcng Lt iden var. tomen- to s few, — to os ers San Brancisco Count ty. 1. L. germanorum. Pee the glands if f ned ; widely distribut Phyllaries 18a Sd dake, Geoke 4- Svinte: slender spring-flowering enudie 2. L. tenuis Phyllaries closely imbricate, about 6-seriate; often stout summer-flowering ann nile Sk. old ulifer. Corollas pink, lavendar, or sometimes winte. never yellow; involucres turbinate to cylindro- bis tO long ulosa). T * relation to Lb he a sti: in L. ramulosa var. ram axils , rarely glomerate on axillary arn t ore rarely glomerate. Basal leaves relatively few and withering at or hitare anthesis; oe a relatively slender at base; Phyllaries woolly. tomentose (see | also L. micradenia senchontins) punctate pignds only pres- nt, lar stipitat . L. leptoclada. Phollaies not adie tomentene fexcept . Zz. percents eee | punctate and larger stalked ands present, either sparse or abundan oe glands abundant, evident; wides a species 7. L. nemaclada. Punctate glands very few and inconspicuous; scvpentiie areas in Marin, San Mateo, and Santa Clara Counties. 8. L. micradenia. Basal evita many and tending to persist through anthesis; stems thickened at base; punctate glands ng. Sine cauline leaves with tack-shaped glands; outer corollas only slightly enlarged and not palmately spreading. Heads 6-10 mm. broad; pappus-bristles many, nearly free 10. L. ramulo. Heads 3-5 mm. road pappus of few bristle-tipped awns or or completely paleaceous, Ag 10, MW PD Upper coe — lacking tack-shaped glands; outer corollas much Sail and Gadeadicty eadin 11. L. hololeuca 1. Lessingia germanoérum Cham. San Francisco Lessingia. Fig. 5644. Lessingia germanorum Cham. Linnaea 4: 203. 1829, 1-3 dm. high, rather slender, —— branched from the base, procumbent, rarely erect, = herbage and — ae = sel ish-tomentose, glandless, glabrescent, and dark- d in g, obla anc Cede acute or obtuse, tapering to a petioliform base ; pice Meas saciter: jet pnt d or pinnatisect, the uppermost subentire, bracteate ; h 38-flowered, solitary on the pede divergent branchlets ; bnokic 5-7 m m. high, campanulate, the phyllaries about 6-seriate, loosely imbricate with recurved tips, gat tl bove, > rous ; corollas deep lemon-yellow with a brownish or purplish band at t the t hroat, the outer nda sty anches ae eg wae with or without a short > papp bri stles about 30, equal, essentially free to the bas Coas' wp ae Humid Transition Zone; San Francisco County, the type seni Aug.—Nov. 2. eee ténuis (A. Gray) Coville. Spring Lessingia. Fig. 5645. ar. tenuis A. Gray, Bot. Calif. % me 1876 Lessingia tenuis Coville, Contr. U.S. Nat. Herb. 4: 1893 pa Lessingia heterochroma H. M. Hall, Univ. Calif. Pub. Bot. 3: 67. 1904. Lessingia tenuis var. jaredit Jepson, Man. FI. Pl. Calif. 1041. 1925. Lessingia germanorum var. tenuis J. T. Howell, Univ. Calif. “Pub. Bot. 16: 16. 1929. Annual, 3-15 cm. high, diffusely branched from the base, the stems and the very slender often reddish divergent branches — loosely ‘a below when young, olde er pla nts glabrous. 1 leaves ulate, narrowed to a petiole, not persisiting, entire or shallowly eeply fant regularly innately parted, tomentose on both faces; stem-leaves 0.3-1.5 cm. long, s sessile, oblanceo late to obovate, entire (some of the plants of the t Ham Ran with irr nee -leaves), mostly persistently tomentose on both sides but sometimes e, also glandular; heads solitary lender branchlets, these nak ith few d stem-leaves, 1 ow gat 4- anu loosely 4-seriate, the phyllaries m ored, suffus purple; appendages of the style-branches ut a cusp; appus-brstles eo mostly free to the base, claret ¢ o 5-8 awns in forms described a Dry slopes, open or in chaparral, Unper ‘ moran ma Inner Coast Ranges, Santa Clara and Stanislaus Counties, ue to the Mount Pinos’ regi Ventura County. Type locality: Piru Creek, Ventura County. April-July. SUNFLOWER FAMILY 377 3. Lessingia glandulifera A. alge Valley Lessingia. Fig. 5646. L ia glandulifera A. Gray, Proc. Amer. Acad. 1 882, L gt var. elo I T. Howell, Mls ae Pub. Bot. 16: 19. 1929, [ ngia var. tenutpes J. T. Howell, op. cit. 20. Lessingia var eaters: T. Howell, op. cit. 22. Annual, (1)2-7.5 dm. high, more or less slender to rather stout, branched below the middle with stiffly spreading ssaine: these much branched above, the herbage in younger plants loosely tomentose below often up to the “aeaees beet inflorescence but becoming glabrate in age, very eaves 2-6 cm. ar and strongly scented. Lowest | ong, vate or oblanceolate, pinnate to pinnately lobed (rarely bipinnatifid), narrowed to a petioliform base; st leave ller, sessile, ] te, en thed, the uppermost bract-like and rather densely beset h ender floweri ranches, usually with tack-shaped marginal gland e r ome- times glandular ; heads 17-33- ered; involucre 5 h, broadly turbinate, the phyllaries closely imbricate, about 6-seriate, gland also with tack-sha gla yellow with ular t pe a pil tds plish band at the throat, the outer palmately cleft; ese i of style-branches deltoid to arrowly tri uss el with or without a short cusp; pappus-bristles 30 or more, i to the base. Grasslands, Son Zon San Faves 5": County in the Central Pater and the adjac Coast Ranges to cients citing California and L r California; le the foothill area of the Sierra Nevada from Pere Coun ais southward. Type locality: not defnit ie stated. y—Nov singia glandulifera var. pectisata. iucene) “Tensom 7. . Pl. Calif. 1041. 1925. iLersingie pecti- nata cist Proc. Acad. Phila. 1895: 548. 6; a var. pectinata J. T. Howell, Univ. Calif. Pub. Bot. 16:18. 1929.) Plants very diffuse ia Mod nched from the base, glabrous or soon gla Soise. more or less glandular throughout; canine leaves pectinate- vinnalifd with cuspidate segment; shy lave with con- spicuous glands, the tips squarrose. Dunes near the coast, Monterey ay south to northern Santa Barbara pena dha locality: Monterey. ngia glandulifera var. tomentosa (Greene) Ferris, Contr. Dudley Herb. 5: 101. 1958. (Lessingia choy pannel Leaflets Bot. Obs, 2: 32. Phe L. germanorum var. tomentosa J. T. ity Univ. Calif. Pub. eo 16: 24. 1929.) Plants depre ssed, ely tomentose throughout except the involuc these 5-6 mm. high, with large marginal glands. Gecdear part of the Colorado Desert, San Diego County. Collected by reutt. 4, pes lemmonii A. eo oe Lessingia. Fig. 5647. oe lemmonit A. eae roc. Amer. Acad. 2 essingta | eee. T. sha Gan hoe Pub. Bot. 16: 25. 1929. Annual, 6-30 cm. high, branching from the base when young with few freee branches, oe much Laas d above in age, the branches je pe os when you soon green and finely glandular-puberulous. Lower leaves 3-3.5 cm. long, spatulate or 5 ovate ; stem- out 1 gi ered; lucres 5-6 mm. h close imbricate, obtuse or acute, glandular-puberulent and bearing large margina lands; corollas yellow, w aioe a purplish band at the throat, the outer pa alin ately sigh aed appendages of the style-branches . 7- mittee m. long, slender-subulate ; pappus-bristles mostly fre abundan Dry sa we Zones; northwestern Arizona west through San neniee County, eS and pr in te ee ae is the Mount Pinos region. Ventura and Los Angeles Counties; occurring sparingly in eastern Santa Barbara County. Serpe locality: Ashfork, Yavapai County, 5 eto Jun 1e—Sept. ia ermal var. ramulosissima (A. Nels.) Ferris. Contr. Dudley Herb. 5: 101. 1958. (Les- Fi ma Nels. Univ. ate Publ. Sci. 1:.138. 1926; L. noel var. ramulosissima J. T. deci "Unie, Calif. Pub Bot. 16: 1929.) Like L. Jemmonii var. lem in stature and branching but older plants more intricately ed “herbage including phyllaries more uae ats differs in having narrower 5645. Lessingia tenuis 5646. Lessingia glandulifera 378 COMPOSITAE involucres and more bracteate primase branches. Owens ada Inyo County, California, and in the north- western Mojave Desert. Type locality: Hinckley, San Bernardino County. Lessingia lemmonii var. peirsonii (J. T. Howell) Ferris, Contr. Dudley Herb. 5: 101. 1958. (Les- singia germanorum var. peirsonu J. T. Howell, > basil Calif. Pub. Bot. 16:26. 1929.) Like L. lemmonii var. lemmonit in growth habit, pode in hav sd dig a oe matted tomentum wie is only partly deciduous in age and by persistently woo olly i ucres. Wes Mojave Desert and the adj cent mountains of Kern and north- ern Los Angeles Counties. Te locality: Kings Canyon, Liebre ome Los Angeles County. 5. Lessingia nana A. Gray. Dwarf Lessingia. Fig. 5648. Lessingia nana A. Gray in Benth. Pl. Hartw. 315. 1849. cage gingra sane onl caulescens A. Gray, Syn. Fl. N. aoe 12; 163. 1884. 885. reen e, Bull. Calif. jemi ) ae 3 Dw at plants, stemless or nearly so, 1 = high or with a few erect or decumbent, some- times prancing stems 2.5-10 cm. high, sabi e g white-lanate throughout with long tangled hela the leaves becoming elabrate i in age. Basal leaves rosulate, up to 3.5 cm. long, petiolate, linear- oblanceolate, acute and spinescent-tipped, entire or waver iemeis thed, pene eined, the stem-leaves similar, atl apc te, about Ace m. ; heads hatches at the base or at ends of branches, 10-15-flowered; invo ucres 7 high, as e outer fies vee ge woolly, lanceolate, mucronate, green-tipped, the in scans: sc us- ene taceous, with a stiffly erec cartilaginous, awned tip; corollas rose- “aiorel or ole ‘all essentially regular ; Wie: branches with or without a sho pe p; achenes 2-3 mm. aioli: pappus-bristles showy, rose-colored o rufous, = twice as — - — chenes. Open ces plains and slopes, Sonoran Zones; Sacramento Valley and adjacent eastern slopes from Tehaden Score seat thecual on “Bae —_ Valley — i Rel sag ohare in Kern County; also na- tividad, Monterey County (Abbott). Type ality: near S ento. Collected by Pickering. June—Oct 6. Lessingia virgata A. Gray. Virgate Lessingia. Fig. 5649. —— virgata A. Gray in Benth. Pl. Hartw. 315. 1849 Lessingia subspicata Greene, Leaflets Bot. Obs. 2: 29. 1910. en rather stout annual, 4-6 dm. high, Tag te at and also above the base with imes present with heads in small glomerules), the h bage densel lly-tomentose, becom- ing glabrate. Lowest lea oblanceolate, acute, soon w , the line 1 0 long, the uppermost scarcely reduced, usually longer than the inter , appressed, obl vate to ovate; ; involucres 5-7 mm. high, indro-turbinate, the phyllaries -seriat ly imbricate, glandular, loosely woolly below, and pur ; corollas lav e, close : der, the outer palmately cleft ; appendages of the style a amen with a short es : sie subulate cusp; pappus-bristles 27-32, about pier aling the achene, tending to be united at the bas __ Dry plains vagy Pry io OP ono aap the ie r Sonoran Zone; Sete ae Co ho ount Hamilton Range (H. Sharsmith) and the foothills of the Lessingia nemaclada var. albi a (East 9. ke Univ. Calif. Pub. Bot. 16:29. 1929. ‘tee sia aibtteos 4 Eastw. Bull. Tae Club $2: 217, 1905; hc 2 yon a om var. albiflora Jepson, Man Cali aes: Be .) A slender er ase with slender divaricate branches and - ads 6—8-flowered aa white ok a he lobes suffused with pale lavender. Inner Coast Ranges, southern Monterey ro sents to the Tehachapi iacmrsine and the southern Sierra foothills in ae County. 8. Lessingia micradénia — Tamalpais Lessingia. Fig. 5651. Lessingia micradenia Greene, Leaflets Bot. Obs. —r ramulosa var. micradenia J. T. Howell, ana po Pub. Bot. 16: 39. 1929. er annual, 34 dm. high, stems simple below, paniculately branched from about mid- SUNFLOWER FAMILY 379 stem with divaricately spreading branches, Sarees | dafaghre: but bearing tack-shaped glands below the heads. Basal leaves 3-4 cm. long, wed t petioliform base, irregularly toothed, soon deciduous; upper leaves progressively pied ae lat oblanc olate to linear-lanceolate, mostly entire, bearing an apical cusp, loosely et tata ok flowered: few bract-like gis ae ei ; heads 5-10-flowered ; uppe at pita glands, the outer more densely glandular ; coeottes lavender, ‘he’ outer port. Re scarcely palmately spreading; appendages of the style-branches evidently cusped; pappus united into 5 paleaceous awns. Thin gravelly —_ mostly = serpentine, Upper Sonoran Zone; hills of Marin County, especially on Mount Wagitlenie: the type locality. Se This taxon ke Pt taxa are very limited in distribution. They appear to be akin to the L. nema- clada carer Lessingia micradenia var. glabrata (Keck) Ferris, Contr. Dudley Herb. 5:.101. 1958. (Lessingia ramu- losa var. glabrata Keck, Aliso 4: Boe ioe Like L. micradenia var. micradenia in habit and leaf-shape, the — on sys er pl co ak heads 3-5-flowered; involucre shout 4 ee uppermost bract-like leaves persistently tom mm. long, cylindro- eer ape pe lari ies wi ith Vv y few small stipitate glands, cpl more abundant, larger bse sessile or short- gpa ne large vitae pes as present; pit an m7 tles more or _ ted below. On pe ig inner slo “of the Coast Range, Santa Clara County. Type locality: eat of L Gat atos on road to Imad ar Lessingia virg 5650. Lessingia 5647. Lessingia lemmonii 5648. Lessingia nana pita 380 COMPOSITAE Lessingia micradenia var. arachnaldes. (Greene) Ferris, Contr. Dudley Herb, 5: 101. 1958. (Lessingia arachnoideu tape Me rey iy Bot. ag 1910; LE. phololeuca var. arachnoidea J. T. Howell, Univ. Calif. Pub. Bot ; L. leptocla eee chnoidea Blake, Journ. Wash. Acad. : 271. 1929.) Habit and leaf-sha ape o bh Pe (oe cic vent “oleate am gs at woolly below. the heads; peer nyt bract-like leaves Reames tomentose on the ag oe surface; heads Boe od wered; even cre 5-8 mm. high, about 5- pee e; phyl- laries sely imbricate, acute, loosely woolly, very sparsely bese Sipe sessile or "hort at alked, large ands; outer pee tA palmately cleft; meeps br istles ork fre 5 ar un nied} ets. On serpentine, vicinity of Crystal Springs Lakes, San Mateo County, the type locality. Perhaps specifically — = goss basis of the abundance of wool on the involucres and the somewhat lar rger, more abund y flowered hea 9. bensings leptoclada A. Gray. Sierra Lessingia. Fig. 5652. ia leptocl A. Gray, Proc. ates veges 7: 351. 1868. Lessingia po var. tenuis A. Gray Erect slender annual, 3-8 dm Se except depauperate forms, commonly simple at the base and ieakelttte about the midstem with v. ry slender divergent branches, sometimes basally branched, woolly when young, becoming glabrate. Basa! leaves 0. cm. long, oblanceolate or spatulate, mostly narrowed to a petioliform base, entire or irregularly and few-toothed, woolly ; stem-leaves 2.5—4 cm. long, sessile, Ape hey fewer and much reduced below the pale. siandular. punctate; heads solitary at the ends of the branches, sometimes in clusters of 2 or more, rarely somewhat spike-like, (6)12-22-flowered; involucres 5-10 we po gh, sl Stony turbinate, closely imbricate; phyllaries 5-7(8)-seriate, the ag nich mostly tide | punctate glands sparse; corollas lavender to ee purple, the eae he ate; appendages of the style- branches triangular, with or without a pappus- aa 1840, typically distinct to the base, mor rel omewhat ig “ahh united ee g ngs in forest, mostly Arid Transition sas Gilts Nevada, Eldorado County to Kern County, Type teaches: Yoserin Valley, iertrcee County. July- Oct. 10. aces. ramuldsa A. Gray. Sonoma Lessingia. Fig. 5653. Lessingia ramulosa A. Gray in Benth. Pl. Hartw. ae ate Lessingia bicolor Greene, Leads Bot. Obs. 2: 28. Annuals, 2.5-4.5 dm. high from a aa tomentose base, the main stem erect and divari- cately inet ed, also beseetane from the base, finely and rather LS are aa ee. Basal and lowest leaves 3-7 c i Soy mostly oblong- oblanceolate, some n: wed t ioliform se, margin irregularly de pate to nearly entire, loosely tomentose, perrating : seiet per bract- like leaves progressively smaller, ovate, sessil, and often clasping, densely and persistently woolly r he lower glabrate, more or less densely b ith tack ed heads solitary at the ends of t nches, a wered ; invol ide, broadly turbinat 0 c late; phyllaries about eriate, rather loosely imbricate, thin, he inner oer aie oid stipitate-glandul with so larger tack-sh nds; corollas rose-colored o e pink, er eg scarcely enlarged and not palmately cleft; appendages te) ou ed c of the style- piasckes very short and not cusped; pappus- bristles 25-37, mostly free at the base Clay soil on slopes or in valleys, Upper pie Seal North Coast Ranges, Mendocino County to Sonoma County. Type bean near Sonoma, — ese Aug ich Lessingia ram var. adenéphora (Gre — Syn. FI. Amer. ed. 2. 446. 1886. Pec singia adenophora “unr Bull. alt. gon 190. 188 : Visas smaller, ‘stems en ‘hickened at the en- tose base ee basal and lower leaves mostly pene; stem-leaves ovate, woolly, margin densely lly ‘with tack-shaped glands; heads t cto 2 with fewer flow the involucres 3-5 m mm. a aaa: pappus seeseg aber or — ted into 5 bristle ipped sin n the Inner Poast eens in Lake, Napa, and Colusa Counties. Type localit r Epperson’s, Lake Coun 11. Lessingia hololetca Greene. Woolly-headed Lessingia. Fig. 5654. ee ——. Greene, hd ae: n. 377. 1897. , Lea’ ies Obs. 2: 27. 1910. Lessingia pid syed Green, 4 loleuca var Man. FI. Pl. Calif. 1040. 1925. ect or asc ae annual, 0.5-3 dm. high. high from a thickened tomentose base, the stems le w or branching from the b: rather stout and densely tomentose, the branches more slender, stiffl ending and tardily glabrescent. Lowest leaves 1. ong, y as (8) vate oblanceolate, acute, sessile or narrowed to a petioliform base, persisting at anthesis, entire o arly toothed; e aes reduced, Bathe e to oblongish, sessile, becoming bract- like below the heads, densely tomentose and beco athe hose in age; heads mostly s pita tary on the ends of the branches, sometimes eeu 3 in she lent ils, ‘the flowers 13-18, p: inkish or lavender ; involucres 10-12 mm. long, turbinate ; phyllaries S-6-seriate, spreading, lanc Lacey sharply oe: glandless, the outer woolly, the inner glabrous or nearly inged apically with purple; m margin flowers palmately cleft pel enlarged; style eke ne with Aram appendages ; pappus- Dbtlsti¢s 40-55, up to 8 mm. long, not united at the - Grasslands on hills ‘ane in valleys, Upper one; Yolo, Napa, and Sonoma Counties south to San Mateo County. Type locality: San Rafael, Marin per Fone Nov. 93. CONYZAL. Sp. Pl. 861. 1753; emend. Less. Syn. Compos. 203. 1832. No dum omen conservan nnual or perennial herbs. Leaves alternate, pinnatifid or bipinnatifid to subentire. Heads small, usually numerous, disciform, whitish or minutely radiate with white or purple SUNFLOWER FAMILY 381 rays. Involucre (in ours) about 2-seriate, subequal, of narrow subherbaceous phyllaries Pecertoete: flat, naked. Pistillate flowers numerous, several-seriate, their corollas tubu lar- flowers. ew, their pre ta (in n ours) with a slender tube and slen eae na en a e, ce penda Style-branches hispidulous above with short, ovate, hispidulous appe dages. Achenes small, oblong, compressed, the pappus (in ours) o rather ee and fragile, unequal, capil- lary bris - aero Greek, applied to some kind of fleaban Ag of about 50 species, most of which are natives of tropical or kote regions in both hemispheres. Type aclad, Soman chilenas Spreng. All leaves cuneate-obovate, sharply and —— toothed. 1 Leaves linear or linear-lanceolate, entire or the lower sometimes lacin Involucres densely <2 ae 5-8 mm. "high Pistillate flowers without igules. . C. coulteri. 2. C. bonariensis. . C. canadensis. a 5654. Lessingia EEE 382 COMPOSITAE 1. Conyza coulteri A. Gray. Coulter’s Conyza. Fig. 5655 Conyza coult tert A. Gray, Proc. Amer. Acad. 7: 355. 1868. Conyzella coulteri Greene, Fl. Fra Esche askie coulteri Rydb. Bull. eehotns Club 33: 154, 1906. Annual, usually 1-stemmed, 0.2-1 m. high, usually simple below the much-branched in- orescence stipitate-glandular and spreading-pilose aoe de many-celled hairs throughout (including to 10 cm. long volucres), very leafy. Basal leaves cuneate-obovate . long, sharply toothed, acute, rian ng to the ne eine petiole; stem-leaves din gg oblong, 1-6.5 cm. long, 2-20 mm. wide, sessi z somewhat clasping, sharply serrate to coarsely and doubly serrate, with a id- b; heads small, 4—5 h in fruit, very erous in a vi pyramidal panicle; in 2 igh, about 2-seriate, subequal, the phyllaries linear, te, ee center and whitish margin, the outer m herbaceous, the subscarious; pistillate flowers very numerous, their corollas whitish, about -_ as long as the style and pappus; perfect flowers 5-10; pistillate achenes elliptic-oblong, 0.7 mm. long, hispidulous, their pappus fragile, rather sparse, white, 3 mm. long; disk-achenes saiuhedivas with glanduliform Valley flats and plains, atten in alkaline soil, Langa ay aot goceerine as a weed, Up pper r Son Zones; central Calibicnia in Santa Clara, Santa Cruz, and Am r Counties south, tooheta ~~ Pt oe a to San Diego County and Lower Californ nia, east to Colorado pee "Teens and south to Mexico, Type locality: not definitely stated. Collected rated Coulter. April- Oct 2. Conyza bonariénsis (L.) Cronquist. South American Conyza. Fig. 5656. Erigeron bonari as SpePl. 863.1753: Erigeron cr rispus P Mém. Acad. Toul. 3: 318. 1788. Erigeron undulatus wou Meth. 598. 1794 Erigeron linifolium bak on Be od Be . 1955, 1804. Conyza ambigua DC. Franc. 6: 468. 1815. — ign aal eg ba Acad. St. sesh i 235. 1856: Il . Fran. 386. Conyza b is Cronquist, Bull. asta i, 70: 632. 1943, denial usually 5 dm. high or less, subsimple or with erect oe si branches as long as or longer than the ma stem and leaves oe Sse strigose or incurved-pubescent and also hir- sute — hispid. Leaves linear to lance-linear, or the lower sla ceolate, mostly 8 cm. long or less, wide or less, the lower coe laciniate-toothed, uF upper re; heads trad bin or pi isicscesy 5-6 mm. high, racemo: in a narrow pan Tesh similar t at of C. cana- ensis, 4 mm. high, the ohoteaiics ; Seauely hirsut aes aie middle; pis Sette corollas very ons sm whitish, filiform, slightly dilated and acutely 3- Boothed at the often purplish apex, at maturity much te =. the style, equalling the pappus; perfect pte bes ; achenes Siveaeaioes. ot rved; pappus simple, brownish white, rufescent in old s Waste places near aly ag adventive at Portland, Oregon, sat occurring more ae ia central Cali- fornia and the Bay Region to southern Cal ifornia, east to Florida and New Jersey, south to South America; intro- duced from South Am es, Type locality: Buenos mee June—Au Conyza floribinda H.B.K. Nov. Gen. & 4 1820. (Conve albida Willd. ex Spreng. Syst. 3: 514. ing Erigeron for eeenne t pone ee. Bull, ibe “Fr. 12:81. 1865. . Lateral branches of the Scavecaleane not surpassing the $; pubescence Se C. bonariensis; heads somewhat smaller than that species. Adventive at Portiond. ches Type locality: near Quito, Ecuador. Conyza bilbaoana Remy ok Gay, Fl. Chil. 4: 76. 1849. Differs from C. canadensis in having obliquely sg, — flowers in the hea mo ie in being perennial - at least persisting more than one season. Adventive n San Francisco Count ne Aemiecg italiey : South America. Cone 3 oe =| ~ wn, n be 3. Conyza canadénsis (L.) Cronquist. Horseweed. Fig. 5657. rigcron canadensis L, Sp. . pe uS Eri Conyzella canaden. mie d. St. Pétersb. VIT. 14: 51. 1869. id dg in conadens dhe Wrown, Til FL 3: 391. 1898. Con , Bull. Torrey Club 70: 632. 1943. Marsea anadenst. Badillo, Bol. Soc. Venez. Cienc. Nat. 10: 256. 1946. Annual, strict, . a high sd less, simple below, with many erectish branches above, leafy into the i em and leaves (es specially - ’ margin) hispid or hirsute. Leaves numerous, paged to a ig et or the lower oblanceolate, 2-10 cm. long, usually 2-8 mm. wide, entire r the lower ser eather-vei 3-4.5 fe a pi central glandular area and narr a auteacions margin; rays numerous ef not more than , white, short and inconspicuous, er aah, a the disk by 1 mm. or less ; achenes hirsutulous ; pappus ’simple, dirty-whitish. Cultivated and waste places, nearly throughout temperate Bort, America south to South America; naturalized in Europe. Type jain “Canada, Virginia.” Jun stent. Hogwee Cen ensis var. egy nll (A, Gray) Cronquist, — gus Club 74: 150. 1947. (Erigeron cana- densis var poe vrcanes A. Gray, . Nat goon = ver 850.) Differs from C. canadensis var. canadensis in having the stems glabrous or ee so. Mor aie area than the typical form but having much the same distribution. Type locality: Texas. 94. BACCHARIS L. Sp. Pl. 860. 1753. _ Dioecious shrubs, rarely only suffrutescent. Leaves alternate, entire or toothed, usually thick and firm. Heads discoid, small or medium, usually numerous and many-flower red. SUNFLOWER FAMILY 383 Involucre few- to several-seriate, of usually thickened, dryish phyllaries. a flat, alea naked, very rarely conical or paleaceous. Pistillate corollas tubular-filiform, shorter than the style; hermaphrodite (functionally staminate) flowers with tubular, Yee 5-toothed corollas. Act s of the llate flowers narrow, -ribbed, of the staminate flo . Styles of sta pidulous. [Named for Bacchus, without evident spbiicnoe ” An American ae of about 300 species, the great majority of the species in South America. Type species, Stems and b hes distinctly puberulous or — Stem finely puberulous; leaves entire, linear or squamifor 1. B. brachyphylla. ssmines? ag idly pilage; $e hes mary and oes toothed, nates oblong or elliptic-oblong. 2. B. plummerae. or glutinous. bh aay fastigiately much branched, with very numerous erect, broom-like branches, usually leafless at flower- ng t meas of the pistillate flowers —_ about 3 mm. long, not surpassing the style; receptacle so omewhat chaffy ; larger leaves obova’ 3. B. sergiloides. Pappus of the pit mayer in fais 6-11 mm. long, much surpassing the sabe receptacle naked; larger s lin . B. sarothroides. igh not ot fasiiatly ear: branched B. emoryi), not broomlike, persi eae leafy. Le neate to oblong, 1-nerved or with a pair of ‘ak lateral veins; heads partly or wholly in sessile ae Fei neulate ae! glomerules. Inflorescence sparsely leafy, i bl b1 lat te-obl 5 7 s.2 leafy, its leaves oval, obovate, or broadly cun 6. B. pilular Leaves eens lanceolate to quate, ually rather strongly Siolteaares: hii in small or large eis tips of stems or bra s. ati essentially eds ie sa leaves ovate to lanceolate; phyllaries thin; fete ae a ae ee Stems ih tsk leaves chiefly linear or linear-lanceolate; phyllaries thick and firm; ia paral glabro: Heads eanieled at apex of s 8... on Heads mostly in panicles = esi of short, lateral, flowering b h 9, B.v 1. Baccharis brachyphylla A. Gray. Short-leaved Baccharis. Fig. 5658. Baccharis brachyphylla A. Gray, Smiths. Contr. 5: 83. 1853. uffrutescent, intricately bran ched shrub less than 1 m. high, spreading-puberulous or hir- tellous throughout, not ig 2 cet diffusely a onled: slender; branches sulcate-striate. Larger lin 1.8 cm. long or les i e whitish ari heads 4-5 mm. high, their involucres uti but : fertile achenes about mm. long, 4-5-nerved, sparsely puberulous, their pappus rerio eae. about 7 mm. long, rather copious, — —— Lower Sonoran Zone; San Diego and southern Riverside Counties, California, east to Ari- og south to Sonora. Type said “Between Conde’s Camp and the Chiricahui Mountains, in ston oat. * “Collected by Wright. Aug—Nov 384 COMPOSITAE 2. Baccharis plammerae A. Gray. Plummer Baccharis. Fig. 5659. Baccharis pl A. Gray, Proc. Amer. Acad. 15: 48. 1879. Herbaceous from a woody base, | m. high or less, usually little branched below the inflores- cence; stem striate, sordid crisped -pilose. Leaves linear to elliptic-oblong or narrow-oblong, 1.5-4.5 cm. long, 3-12 m wide, a obtuse, narrowed at base, sessile, sharply and closely serrate, 3-nerved, glabro whee ex seine ae costa settee crisped- eed beneath ; heads loosely or closely panicled; pist siHlate heads 1-1.2 cm. high i in frit, their involucres 6-8 mm. high, gradu- ate, the phyllaries linear or rice lanceolate, acute or acuminate, eg narrow or ieeat crisped- pilose, green center and cili subscarious marsh; saminat hea high, their in- A is aoe but ssrebo verti achenes 3 mm. long soso i puberulous, 4-5- rved, their pappus becom rufous, rather pare 5 mm. | Saieaeten ravines, Upper pe Zone; coast of Santa Barbara and L ngeles Counties, California, and Santa Cruz Island. Type locality: Glen Loch’ ravine, near Santa Rotace: Caltorere Aug.—Oct 3. Baccharis sergiloides A. Gray. Squaw Waterweed or Desert Baccharis. i Baccharis sergiloides A. Gray in Torr. Bot. Mex. Bound. 83. 1859. hii shrub, 1-2 m. high, the stem and branches green and strongly siete eo the nchlets numerous, Font tigiate, broom-like, casetially leafless or sometimes (especially the sterile ne leafy. Larger leaves obovate, 1- ong, 3-8( wide, pare prs with 1 or 2 teeth on the largest leaves, obtuse, api a bigt va te to the sessile base, thick, with aed peers and a pair of weaker nerves arising from PS the leaves of the branchlets mostly tgs form; heads eerste Hes densely panic ae hoa ag nt eg pistillate heads 4- hi ve in fruit, their involucres 3 mm. high, about 5- eats: strongly graduate, the thin « ale or lance-ovate, obtuse to peer Phe with Beni center — whit cea mara tr; sta sree heads 3.5-4 mm. high, with similar involucre ; n both ome vies onic, bearing few to many linear paleaceous bracts ; pistillate pas eg with 5-6 enter” teeth; fertile Sehanel eSout .5 mm. long, glabrous, 10-ribbed, their pappys sparse, 2.5-3 mm. long, sei shorter than ie =A Nag satan bristles of staminate sleigh ated and wbathollat: ata gE creek nee 8 nd can pom r Sonoran Zone; the Death atic region, Inyo County, and the Mojave a rado Des Californ me east oe southwestern, "Utah and Arizona, and south to sn Type locality: not decanters ee but from the Colorado Desert region in Arizona or California. April—Jun 4. Baccharis sarothroides A. Gray. Broom Baccharis or Hierba del Pasmo. Fig. 5661 tne conrce sarothroides A. Gray, Proc. Amer. Acad, 17: 211. 1882. Eastw. Proc. Calif. Acad. IV. 20: 155. 1931 Erect cpa la shrub . high with very ties fastigiate, broom-like branches, nearly leafless ; stem bro heehee ridg ed, “the branchlets green, strongly ingrne angled. Leaves linear, as m. long or less, ihe mm. dong 1-nerved, those of the branchlets squamiform ; heads numerous, ostiy solitary at tips of short or long, naked, 4-angled branchlets, scceveaiae or. racemose-panicu- latch rranged ; pistillate heads 1-1.7 cm. high in fruit, their involucres oblong, (3)6-8 mm. high, 6-seri te (the inner — s similar, 2.5-4. achenes glabrous, 10-striate, 2-2.5 mm. long, ’ their appus copious, silky, rufidulous, one, greatly exceeding the styles ; pappus-bristles of the staminate flowers dilated but not bearded at tip. hes A desert places, often in alkaline soil, Sonoran Zones; San Diego County, California, and south o Lower California, cai to southwestern New Mexico and south to Sinaloa. Type locality: San Diego County, California. Feb.—Sept. Desert Broo 5. Baccharis émoryi A. Gray. Emory Baccharis. Fig. 5662. Baccharis emoryi A. Gray in Torr. Bot. Mex. Bound. 83. 1859. us, somewhat resinous-granular, evergreen shrub 1-4 m. high, the brane? erect, 4 a atk in age somewhat angled, like the stem usually persistently lea arger eaves cuneate to lance- Mcbiong m. long (includi i n rule 1 sely tips of branches, the whole forming usually broad and pyramidal veins a ipistillate tise 1; "2-1. em. high, their involucres 5-8 mm. high, about 6-seriate, strongly graduate, the phyllaries ovate outer) innerm = whiti ith or obtuse, with more conspicuous greenish spot, the inner scarcely elo ongate ; Tate « chenes glabrous, about 10-nerved, about 1.5 mm. long, set pappus sli _. rufescent, silky, 1 cm. long; pappus- reese of staminate flowers with lacerate-bearded, somewhat dilated ti ips. Along w Sonoran Zones; Kern County ‘ia econ. California, south to morrhere Lower hi fornia caeard 1 to the te Colorad a Desert, d th Nevada, Secality Safieg the Gils Rivet, Artoous, Cilicciel by Kec Senge, Utah and Arizonpto weetern.Texss, SUNFLOWER FAMILY 5660. Baccharis sergiloides ‘ is sarothroides 5662. Baccharis emoryi 5657. Cony densis 5658. Baccharis brachyphylla 5661 5659. Baccharis plummerae 385 386 COMPOSITAE 6. Baccharis pilularis DC. Dwarf Chaparral-broom. Fig. 5663. Baccharis pilularis DC. Prod. 5: 407. 1836. Matted and widely spreading shrub 12-30 cm. high, the stems prostrate or nearly so, glabrous, resinous- ey stems brown-barked, sige the br i striate-angled, very leafy through- lat 8) i i out. Leaves obovate-cuneate or aneyh = 1-4 (ra cm. long (including petiole, this 8 m ea or wanting), 0.4—-1 ke 3 cm. es or sometimes acute, cuneate at base, repand- serrate above t ntire oe e or the up aimaee thick, 1-nerved and with obscure or evident pair of lateral veins, somew ieatee ved d ; erous, in sessile o Monculte axillary glomerul also spicate or clustered at tips of bran p tillate heads in fruit 8- m. high, their involucres 4-5 graduate, the phyllaries ovate- e (outer) to linear (inne st), indurate, whitish, with subapical greenish s and narro liol arious gin; staminate heads 4-5 high, wes invol w, Ciliolate, scarious margi : high ; fertile achenes glabrous, 10-ribbed, about 1.3 mm. long, their pappus ve sear, rufidu- lous, 6-10 mm. long; pappus-bristles of hag seats nowy with dilated faceaate Hill slopes, ee and bluffs along th mid Tra: n Zone; Sonoma County to aed, Sur, Mon- ~ County, Cal nia. Type locality Caltfoenia. ees by stciar Feb. —-Aug. — ris eae var. consanguinea (DC.) K untze, Rev. Gen. Pl. 1: 319. 1st (Baccharis consanguinea DC. Prod. 5: 408. 1836; B. pilularis subsp. ose C. B. Wolf, Occ. Papers Rancho Santa Ana Bot. Gard. : ty 5.) h-br. anched evergre h Not: 21, n shrub 2— igh, differing from B. pilularis var. dintarte only in habit and of ch more common occurrence than that ei Hills and aul of the Outer i Ranges from illamook County, Oregon, south to Sage — Diego County, California, and the isl ast of southern California; also occurring loc foothills of the central Suore Nevada from "Nerata County to Tuolumne County. Type locality: Calitorsta. "Enea by Douglas. Chaparral Broom or Coyote-brush. ae paceeeis douglasii = Salt Marsh or Douglas’ Baccharis. Fig. 5664. rod. §: 400. 18 erbaceous — tially to bas a m. high or less, subsimple below the inflorescence or erect- Sak hed, glabrous, resi sc soeateeie the stem and branches green, striate. Leaves ovate to ie i in, with narrow whitish arious margins, sometimes purplish above; staminate heads about 5 mm. high, ow phyllaries mostly oblong or ge oblong; receptacle br vga’ spocwal: fertile sakes abou mm. long, hirtel- lous, about 5-nerved, their pappus rather sparse, 3-5 mm. long, dull whitish or at length rufidulous, the bristles hspidulous about equaling et ieee pappus-bristles of staminate flowers hispidulous, slightly widened apically, Watercourses ba pre often in salt marshes, Humid Transition and U per Sonoran Zones; Humboldt County to — Diego County in the Coast Ranges and occurring less commonly in the Great Valley and = Sierra Nevada foothills in Amador County southward to western San Hernariinl Saas AM California; also i southern Curry County, Oregon. Type locality: California. Collected by Douglas. July—Dec 8. Baccharis glutinésa Pers. Water-wally or Seep-willow. Fig. 5665. accharis glutinosa hee Syn. Pl. 2: 425. 1807. -epeites coerulescens DC. Prod. 5: 402. 1836. oody at least below, 1-3 m. high, forming thickets, we stems simple plow Shoe niga’ ranched, vey : aly, elabrous, glutinous, stria . Leaves lance-linear to w_lanceolat or linear, 5-15 , 0.5-2.3 cm. wide, acu tape cine into the short sctiele, nee mm. te or less), firm ae ser trate to ne greets fisted, bright green, punctate ; heads numer- us in small ‘or sometimes rather large cymes or cymose panicles at apex of stem and principa ranches ; pistillate heads 6-8 mm. high in fruit, their invo ere 5 mm. high, about 4-seriate, ob “3 above; stam : gy ee ile hears Are di ay 4-5-nerved, about 0.8 mm. long, the pappus er . slightly exceeding the style; piodea-brlaties of staminate Gomis ‘agidalans, aahity dilated at apex. and along watercourses, Sonoran Zones; Fresno County t rn County in the San Joaquin Moist places Valley, ay reas bp south to Lower California; eastward in the Colorado ee" = aie Deserts to Colorado and Texas and south to Mexico; also South America. Type locality: South America. May—Oct. Sticky Baccharis. 9. > atone sents DC. Mule Fat. Fig. 5666. Baccharis viminea DC, Prod. 5: 400. branches eee glutinosa but more Foes, up to 4 m. high, with secrral andl short, lateral, — i es more usually entire, rarely denticulate, less gluti ;h eads in small cym or cy See clic: at tips of the numerous lateral branches as well a agi Aa sal _—: pstllat involucre 4-5 mm. high, about 5-seriate, graduate, the oh tlaries Seat to (in rmost) 1 0 acute or subacumina ate, or purplish eal the outer often with ‘distinctly inet center ; Staminat heads similar; gmc and pappu milar Borders of streams and cany = aia nae said Butte Counties and in the Sacramento Valley and adjacent Coast Rang ant ype . "Nevada foothills: pte 43 pas pee onais. California, the Channel Islands, and northern Nee California; occurring les we and southwestern Utah. Type locality: California. Collected by Deakins. poe i €. SUNFLOWER FAMILY 387 Tribe 4. ANTHEMIDEAE All flowers of the heads with corailas, * ee or = os ones ecco eceptacle chaffy at least Rays in cultivated variety of Antheonis | ble: annual or perennial herbs. 95. . present (abs ds solitary at the tips of the branches; rays elon rae Heads closely corymbiform; rays short and broad. 96. Achill Rays absent; subshrubs. Rad Sontotina sii siadoté naked eet soa baits in Tanacetum potentillioides; scaly in Art i Pappus (in ours) p Flowering heads yellow; pappus- pulese not awn-tipped. 98. Hymenopappus. sh; tipped. 99. Hymenothrix. Flowering heads white or nen rpli papf Pappus absent or a short crow ceptacle flat or pe lesz convex. Rays ¢ cenepicnen, white or yellow (pistillate flowers lacking and thus rayless in sta ithemum balsamita). 100. ps santhem ai Saeed or Load small in proportion to the head. 101. nacetum, 102. aisivindeia mispheric or conical. sin spiciform, racemose, or sresgopiptee inflorescences. 105. Artemisia. Hea Marginal flowers only of the head Heads peduncled; marginal ‘tae ers scdueatie on the receptacle; style deciduous, only the inconspicuous base remaining. 104, Cotula, Heads sessile; marginal flowers sessile; style persisting as a conspicuous stout spine. 103. Soltva. 5663. Baccharis pilularis . Baccharis glutinosa 5664. Baccharis douglasii ome Baccharis viminea 388 COMPOSITAE 95. ANTHEMIS L. Sp. Pl. 893. 1753. Annual or perennial, aromatic herbs, usually with leafy stems. Leaves alternate, la- ciniately incised to pinnately or bipinnately parted. Heads solitary at the ends of the a ; phyllaries many, subequal or imbricated, dry and scarious. R le convex, hemispheric or conical, receptacular bracts sent at lea the summit, hyali y rtilagi- nous, slender or broader, sometimes enclosing the achene wers pistillate, fertile sterile. Disk- ers ct, tube cylindric or compressed, throat narrow m campanulate, lobes short ah cucete truncate or obtuse, times penicillate n thers entire at base, ovate, acute, obtuse at apex. Achenes ellipsoid, terete or obcom- Sabai ribbed, Sa Ay smooth or ihercutate Pappus wanting or a short crown. [Ancient e of the cham eo genus of about a species, natives of Europe, Asia. and Africa. Type species, Anthemis nobilis L. Annuals; rays whit igsites cs dee subulate or awn-tipped, firm Bracts of the receptacle present only toward = middle; — of the achenes tuberculate. 1. A. cotula. ceptacle bracteate throughout; ribs of t 2. A. arvensis. Rece ties bracts oblong, hyaline 3. A. fuscata. ple rays yellow 4. A. tinctoria. 1. Anthemis cotula L. Mayweed. Fig. 5667. Anthemis cotula L. Sp. Pl. 894. 1753. Anthemis foetida La FL. F 2: 164. Ta Maruta vulgaris Bluff & Fingerh. Comp. Ee a 2: 392.1825. Maruta c a:6:-135— 1837. Matricaria pubalcedi Schultz-Bip. Bonplandia 8: 369. 1860 Ill-scented, leafy, erect annual 1-6 dm. high, usually much branched. Leaves 2-6 cm. lon sparsely hairy, bi- or tr ‘pinnatifid with linear divisions; heads wer numerous, Peart esiocor at maturity, solitary at the ends of the branches, the peduncles x m. long; phyllaries loosely short-villous, with a ish or gree ish, thickened wi el and ine margins; receptacle conic, the receptacular bracts a nl boue the middle, slender, subulate, firm, about m ray-flowers sterile, the rays 6-11 mm. long, oblong; disk-flowers pte funnelform ; achenes rea 5 mm. long, ni Shealng anid. with about 10 tuberculate ribs; pappu sah introduced European weed very common in disturbed areas and waste places in the Pacific States; also common throughout the United States. May-—Sept. Dog Fennel. Stinking ae mile. 2. Anthemis arvénsis L. Field Chamomile. Fig. 5668. Anthemis arvensis L. Sp. Pl. 894. 1753. Leafy, more or less br a villous-puberulent annuals 1-6 dm. high, often decumbent at se, not ill-scented. Leaves 3-5 cm. long, bipinnatifid, equaling or shorter than the inter- nodes ; heads alanis at the ssids of the branches, the geen ae Ni 4-11 cm. long, phyllaries villous- t ntose, narrowly hyaline-margined ; receptacle low-conic, chaffy throughout, the rece ptac cular es t wn-tipped —13 mm. long; dis achenes 1. mm. long, quadrangular ied peace Sice adiate, aggregated i at to round-topped, corymbifor: a Pe few, 3-8, broadly ovate to orb mets - ba te white, Leo AH: pink ; ade. flow 15-40, 0 the ae same color as the 1 rays A highly sonics ic ¢ bore: bluffs to arctic pete niptne comditiona: tetraploid, n=18 (=A. lanulosa Nutt.), and the he: ccapiotd, Noo orealis Bong pes sind Key to Varieties Stems stout, 10-20 dm sm cespitose fie om a pow. ‘tire a gray, villous - giga Plants rhizo erate wit eis orming a hekivals at the base of the stem; herbage dark green, uberulent to sparsely v r. puber ——— generally fe than = amy. me me the t inal te t te-] late, densely congested. var. litoralis. ar. arentcola. Hevhawe dark green, marnals villous. Herbage gray, ti sely isin? hiccenra Leaf ot fleshy, Herhaxe gray, Sik tow Margins of phyllaries tight brown to black; roots fibrous from slender rhizomes. ar. me of phyllaries ee brown to straw-colored; nearly cespitose, with short stout Auadien and deep central ta r. lanulosa. Hertogs assay green, moverately omer g to black; stems tawny-villous; lenterements acicular. r. borealis. Margins of involucral bracts pale greenish white to straw-color Leaf-segments s oblanceolate, all segments tending to be Aue in the same plane. var. millefolium. T € = ae. aeeere generally ae mm. high. var. pac 9 mm. high. Achillea millefolium var. analdaa ¢ (Nutt.) Piper in Pibee & Beattie, + vais Bee Reg 196. _1901. (Achillea tomentosa Pursh, Fl. Amer. Sept. 563. 1814, not A. ntosa L. 1753 ; A. lanulosa Nutt. Journ. eae Shona 7: 36. 1834; "A. millefolium lanulosa Piper, Mazam Pe ref 97; 1901: a mien. ge part lanu- losa Pi r, Contr. U.S. Nat. Herb. 11: 584. 1906; A. lanulosa a typica Keck > Car ash. Publ. No. 520; 99 me) Stems 3-5 dm. high, arising from short stout rhizomes sa cespitose) herb e gray, ; leav es em. i im chillea millefolium var. ects (Rydb ) Garrett, Spri Fl. Wasatch Reg. 101. Ts. lanulosa alpicola Rydb. Mem. N.Y. Bot. Gard. 1: 426. esr dk sehen Greene, "Leake Bot. one e Vas. ies A, areal’ Bon yes Bull, Torrey Club as 157 : : , Man. FI. - fusca G. N age sp. alpic t ash. Publ. No. 520: 300. 1940.) Stems seers and erect, 0.5-3 dm. high, arising from long slender r roots shallow, fibrous, distributed ove whole rhizome system; herbage gray, white-villous; procs 5-1 * Text contributed by Malcolm Anthony Nobs. SUNFLOWER FAMILY 391 long, linear-lanceolate, finely dissected into linear-lanceolate segments, prominently spine-tipped; heads few, 5-7 mm. high, in dense, flat- sopped corymbs, the margins of phyll to black, yee va Zone, = Asie Alpine Zone; Alaska southward to Inyo and T tere Counties, California, eastward t e Rocky Type locality: Teton F orest Reserve, Week ing. July—Oct. n=18. Achillea millefolium var. ae (Rydb.) c. N. Jones, Univ. Wash. Publ. Biol. 5: 250. 1936. (Achillea pacifica Rydb. N. gee’ l. 34: age ara? gue jeg 3-10 dm. high from long slender ee: roots fibrous from the entire re ome system; herbage n, moderately ye sontne rm nce pes 15-35 cm. long, dissected into ig ieweds spine- tipoed’ Reins oe, ye many, round- to saad eet “per raga —— hyaline = hier straw color. Interior area, Arid Be Humid ” Transition Zones; eget nee British Columbia, Washington, and Oregon along the ee — of the Cascade Mountains an rm fin aes of Humboldt Con unty, California, extending eastward along the western slopes of se Sictra Meuse. south to San Diego County. Type locality: Grenada, Siskiyou Coabty. California. vent n=18. A variable taxon containing many forms, some indistinguishable from form of var. cali- “Achillea millefolium var. pubérula (Rydb.) Nobs in Ferris, Contr. Dudley Herb. 102. 1958. (Achillea te han Rydb. N. Amer. Fl. 34: 223. 1916.) ae robust, 10-18 dm. high, eas, arising from long ender rhizomes; roots fibrous, concentrated into a dense but gg rye ~ base % Brass seme; herbage dark green, po sely villous to puberulent; leaves Nass lancetiane: Benes 4 sected into linear, spine-tipped segments; upper surface nearly glabrous; heads 5-7 tr % = Sm end: ai veera corymbs, the ag ac eos eee st Salt cand ee sarshen, amid Transition one; ‘San Francisco tay region n Marin, Solano, and Contra Costa Cou , California. Type locality: Suisun Marsh, Solano County. July— ug. += Achillea poatyetinns var. litoralis Ehrendorfer ex Nobs in Ferris, Contr. Dudley Herb. 5: 101. 1958. Stems stout 4 dm. high, ascending to dec siya arising from well-developed rhizomes; roots essentially fibrous from di rhizomes but eres strong central taproot; herba . bev green, sparsely igre through- out; leaves abundant, congested, onal: iseresiaa. primary pinnae borne at right angles to the Dawes ultimate segments highly congested oa very fleshy, tapering into es wa callous spine-tips; heads ¥, 4-6 mm. high, densel ne ted into broad, roun os spre corymbs, the eg rf fp south of ee white Ex. posed coastal bluffs, Curry and oos Counties, Ore: Type locality: - bluffs = geen County. May—Sept. n=18. A maritime form of we tetrapiold closely allied to va fl? Ros ta meer eres in- ahgin a habit. Achil lefolium var. califérnica Potent) Jepson, Man. Fl. Pl. Calif. 1137, 1925. (Achillea cali- ere Pe Bull. Torrey Club 26: 369. 1899; millefolium f. californica H. M. Hall, ber seer Pub. 3:211. 1907; A. borealis subsp. californica Ket, Carnegie Inst. Wash. Publ. No. 9. 1940; : icra var. californica J. T. Howell, Leaflets West Bot. 5: 107. 8.) Stems stout, S12 tne ao" aris- ing from thick, well-developed dieses root-s ns stem a well-d peeoees : tapeent secondary fibrous roots from the rhizomes; herbage green, occasionally grayish, moderately villous; Cag gel ge long, linear-lanceolate, finely dissected into — linear, spine-tipped segments; heads mum fOBR , up to high, in many-branched, Scio loose corymbs, the phyllary-margins light brown. Coastal regions, Upper Seatinene aad Humid Transition Zones ays Harbor County, Washington, south to Lower Californi _—_ ee babes 4 the foothills of the scsi Ate gentise gralifecnia (Eldorado County). Type locality: seat ast at Sant sabel, San Diego County are = illea saioesida var. arenicola (Heller) Nobs in Ferris, Contr. Dudley Herb. 5: 102. 1958. (Achillea crenicole Sagan: eps st 9 me ~ 1904: A. mill efolium var. maritima aay —, Fi. Pl. Calif. 1137. 1925; A, bor, subsp. a arnegie me Wash. Publ. No. 520: 299. 0.) Stems stout, 1-6 igh, epoch | Say nyo ecu sabe iste ys "8 yas well- ss oped rhizomes; root-system m ialy fibrous from the rhizomes but retaining J ‘strong central taproot; herbage Mes te nsely white-villous throughouts leaves oblanceolate, 10— long, the primary pinnae at right angles to wre rachis, secondary and tertia i densely crowded, fie ones to oblanceolate, fleshy, the tips mucronate; heads 5—6 mm. high, densely packed i ae und-topped corymbs, the ph ei aa argins light brown, Coastal dunes and sea bluffs; Del ca Co piece pet to Monterey County, Californ ype ma from a pee vo ee pide pf te ‘horticultural form of C. sal wat Ramond (Bull. Soc. Philom. 2: 140. 1800), has _— reported as an esca gg oa a Cascade iaciaine in Washingioa and Oregon; also Bolinas Lagoon, Mari fe ite poe — L. Corn Chrysanthemum. Fig. 5676. Chrysanthemum segetum L. Sp. tad ceca > ¥ laciniatum Gilib. oc gts ida 218. cha 82. , Baier. Fl. 2: 406. Pyrethrum eticiun Moench, Meth. 597. 1794. Erect, mostly se annual from a fibrous root, 3-5 dm. high, branched from the base or single-stemmed, branching above. Lower leaves and those of the midstem 3.5-6 cm. long, sessile, obovate to broadly spatula in outline, pingatid mostly on the upper half of the leaf, the divisions 3.5 sai not reaching t the mic a upper leaves 1. cm. og sessile, clasping, incised or irregularly toothed or macised t the Buf heads solitary at the ends of the branches, 12-15 mm. broad not including the rays, ‘cad e peduncles 2-10 ay i phyllaries usually 3-seriate Ss long, broadly vate, rm, owish, rounded, the hyali cee widest at om long, tun orm, acute; ray-achenes about 2 mm. ong, somewhat piensa Pecos SUNFLOWER FAMILY 395 the body of the achene ribbed; disk- achenes pale brown or straw-colored, subcyclindric, conspicu- faaaky and evenly 10- —— pappus An attractive introduced European she ro ae Pee s and fields near the coast; Del Norte County to Santa Bar- bara Sak California, ‘de common in its northern range. Type locality: Skane, Sweden. May-July. Corn- marig 3. Chrysanthemum coronarium L. Garland Chrysanthemum. Fig. 5677. be ee sspears coronarium L. Sp. Pl. 890. tcaria a Desr. in Lam. Encycl. 3: 737 Pyrethrum breviradiatum Ledeb. Mém. Acad. St. siatieai Hi 577.” 1015. Erect annuals 3-5 dm. high from a fibrous root, eras or slightly pubescent mostly on the leaves, single-stemmed or branched from the base, the stems branching above with ascending i size, obovate in ine, ow, 1 mm. lipt tube slender, with funnelform throat and acute lobes; ray-achene ut ong, more less triquetrous, the angles bearing wings broad and truncate above, avenue iow: disk-achenes more or less 4-angled but aay COMNEEST one a angle with a wing about 1 m m. broa d and trun- cate at the apex, attenua the opposite angle somewhat w bow iy other two ridged, the space between the inet ond eiane ‘glandu lar-tuberculate and ribbed; us non A native of the Mediterranean region, —— and fields; Marin County, Aiectia: aah to San Diego County; also in Alameda County. May-July aa eb 5675. PR OIE seen —— 5676. themum seget 396 COMPOSITAE 4, a eg. Senne oie parthénium (L.) Bernh. Feverfew. Fig. 5678. S 0. Matricaria odorata Lam. FI. Franc. 2: 135. 1778. Chrysanthemum parthenium Bernh. Syst. Verz. Erf. 145. 1800. Pyrethru a Pees: um J. E. Smith, FI. aes Pitt 1800-4. Matricaria vulgaris S. F. Gray, Nat. Arr 7 Pl2 454. 1821, Much- ais leafy, aromatic ponte 2.5-10 dm. high, sparsely puberulent to glabrous, simple-stemmed or branched sa e base sok rom a taproot or stout caudex, branching paniculately above into a corymbose inflorescence. Leaves petioled, broadly ovate or ovate-oblong in outline 4-10 cm eae the petiole one- sthird or less thee length of the Bias. the leaves somewhat reduced above, bipinnat ely or pinnately parted, the segments acute or ro unded ; s many, abou 18 mm. wide including the white rays ; pylon aries 2- or — eriate, coriaceous and keel ed, lanceolate, en ee E ro ed t u , many more in double forms, 4— 3 mm eee oblong ; disk-flowers 2 mm. or more long, narrowly funnelform, somewhat glandular below and thickened at the ose: aehetes subterete, 8-10- oan: pappus lacking ora anaue § own. of Europe g arden. often escaping; widely distributed but not common throughout the Pacific ‘St “ik es; also in sonar "tnd ves Poe States. Forms with double flowers as well as the usual single- flowered forms are often met with as adventives. May—Aug 5. Chrysanthemum balsamita (L.) Baillon. Costmary. Fig. 5679. Tanacetum balsamita L. ee Pl. 845. 1753 Balsamita major Desf. Act. Soc. Hist. Nat. Paris 1: 3. 1792. 4. Pyrethru — var. tanacetoides Boiss. se ie 3: 346. 1875. Chrysanthemum balsamita Baillon, eer Pi.8: 1882. Chrysanthemum balsamita var. tanacetoides nek ex W. Miller in Bailey, Cyclop. Hort. 313. 1900. Balsamita balsamita Rydb. N. Amer. “ 34: 238. 1916. Coarse perennial 6-12 dm. high with 1 to several erect leafy stems arising from a stout caudex the herbage mint-scented, silvery- Sstrigtse throughout, becoming more or less glabrate especially below. Basal ant many, oblong or elliptic with a pare apex, Setiate d, the blade 15-30 cm. tong, about 3- . broad, crenate-margined, the — a little shorter than the blade; stem- ea p i white ipped with a conspicuous, erose, hyaline lobe; disk- Sooes glabrous, about 2 mm. long, tubular-funnelform, em to the 5 short lobes; achenes about 2 m tae with about 10 ibs’ pappus a small cro oe native of western Asia often grown in gard and occasionally becoming established locally in the ce: States; Whitman County, Washington, and in Je ae Hoo and Josephine Counties, Oregon. Aug.—Sept. Mint 101. TANACETUM [Tourn.] L. Sp. Pl. 843. 1753. Aromatic, stout, erect, annual or perennial herbs from a rootstock, or low, cespitose, woody-based h erbs, or toate subshrubs. Leaves alternate, entire to 1-3-pinnatifid. In- florescence corymbiform or capitate or sometimes solitary at the ends of the branches. o vo i i acute to ¢ fe) tuse. ‘ee of the fears or marginal ones exserted, the branches short, " i¢ ; : obtuse or truncate and niinutely penicillate at the tip, those of ‘the disk-flowers included or shortly exsert enes commonly gerade ——— led or $viied, truncate at apex. appus none or coroniform. [Origin of the na bscure. A genus of about 5 0 species, natives of the northern samen Type species, Tanacetum vulgare L. Heads numerous, 20-200, 5-10 mm. heuad: herbage glabrous or nearly so; introduced weed. > vx ing Heads oo 6-20, about 12-18 mm. broad; herba ill t = a flowers iene a ray; —. densely villous-tomentose. 2; TT. camphoratum. - 3. T. douglasit. ae sender herbe from a thickened candex,’s or subshrubs. 3—S-cleft; receptacle nake: a 4. T. canum. — bi- or tripinnatifid; p hai 5. T. potentilloides. 1. Tanacetum vulgare L. Tansy. Fig. 5680. Nom. 825 SUNFLOWER FAMILY 397 Coarse, aromatic, glabrous or sparsely hairy perennials pp dm. ni from a stout rhizome, the mew stems densely leafy to the compound, many- red, corymbiform inflorescence he unctate, 1-2.5 dm. i i cised or toothed; heads many, yellow ; involucre broadly hemispheric, about mm. wide, a 4 mm. high; phyllaries imbricate in about 3 series, carinate, firm except for the scarious margin and tip, the outermost lanceolate, the inner oblong, obtuse; marginal flowers glandular, with a 3-lobed limb; disk- eee: shortly i bapa = = ese er sparsely glandular-dotted, 5- antec, about 1 mm. lon ery , toothed c A common weed in ee capt pat western cate and occurring el commonly in California along the coast as tar south as vga cpa and in Plumas County; also widespread throughout the United States. Naturalized from Europe. July— 2. Tanacetum camphoratum Less. Dune Tansy. Fig. 5681. nprennr i pempnorann Less. Linnaea 6: 521. 1831. ss. Syn. Comp. 260. 1832. fy es ae DC. Prod. 6: 84. 1837. Tanacetum elegans Decne. Fl. Serres 12: 19. 1857. Stout aromatic perennial 3-5 dm. high from a branched creeping rhizome with striate, erect or Sadie leafy stems, loosely villous-tomentose throughout, densely so on new growth and 5679. Chrysanthemum balsamita 81. 5680. Tanacetum vulgare 5682. Tanacetum douglasii 398 COMPOSITAE somewhat deciduous in age on stems and upper surfaces of leaves. spac thick, 7-25 cm. long, and t mm. ers sat pn short terminal ate with 3 triangular lobes; disk-flowers tubu sn ith 5 triangular seer —— s about 3 mm. long, tr uncate, 5-angled, aeutiees dotted : = a somewhat lobed c Humid Transition al basis and San Francisco Counties, California. Type locality: San Fran- cisco. nf o° ete by Chamisso. June-Sept Closely related to the following species. 3. Tanacetum douglasii DC. Northern Dune Tansy. Fig. 5682. Tanacetum douglasitti DC. Prod. 6: 128. 1837. Stout aromatic perennial 2-6 dm. high from a branched creeping rhizome, with stout, erect or ssonndie ne. leafy, sometimes reddish stems, thinly and — villous-tomentose to glabrate. seve rather thick, 0 cm. long, 2-5 cm. wide, with an expanded clasping base, a tri- natifid, more rarely bipinnatifid, ~~ a of the idan not strongly revolute, 3-5 mm. long, divided to the a ok the segments n t at all a ds ogg deeply redivided into apes or obtuse, mucro e lobes; icc Pek bs bara the simple or branched stems, the peduncles of My sete heads 1-2 cm. long; heads 8-18, y flier “flowana the mature heads 10-15 mm. br: d;p h yllar : ies mm. long, trelate to oblong, ? Files except for the scarious margin and tip; ecient pistillate flowers few, 5 mm. long, with a short but distinct, emarginate ray —. 3 mm. long, the throat ‘eshulsd sages ate; disk- “a gh about 4 mm. long, 5-toothed; achenes 3 o 4 mm. long, truncate, 5-angled, glandular- dotted; pappus a somewhat lobed crown Dunes along ne soem Humid Transition Zone; British Me at: saan south to Humboldt Coranty: California. Type locality: wester America. _Collected by ees. Sony Some of the. collections from southern Oregon and northern Ca Siiee a have less divided leaves more sug- gestive of T. camphoratum than T. douglas sit, but ar : placed — ieccuee of the aS of a definite oaks on the pistillate flowers and the lack of copious villous tomentu 4. Tanacetum canum D. C. Eaton. Gray Tansy. Fig. 5683. Tanacetum canum D. C. Eaton, Bot. King ~~ SEN pl. 29, figs. 8-14. 1871. BY sulambieies cana Heller, Mchlenbetete teas S Subshrub fro: ee root, 1.5-3 = hi gh, markedly branched and woody below with many erect leafy stems. tele 12 mm. lon ng, sessile, oblance 7“ 2 gees or cleft above into 3-5 divi- sions ; heads E rayless, shore Rediunctell in t site nal red metimes solitary, the flowers pale loa invo cgi a 3 agri a 3 mm. high, alia’ wider Heh high; phyliaries ovate, canescent, sc ined ; ot 2 ‘owen vlatiaat: disk-flowers about 2 m long, subcylindric, "acdslarictied witet the short lobes villous ; achenes Nedkcneous’ eran. 1.5 mm. long, 10-ribbed ; pappus none. Cl ae and talus slopes, Hudsonian and A ic-Alpine Zones; ranges of Harney and Malheur Counties, Ore- =. southward through the ran ges of western gh jaan in California pocarring, in the pee Nevada from Mono unty to Inyo and Tulare —— and in the Inyo and Panamint Mountains. Type locality: eastern Humboldt iewiseiac Nevada. July—Aug 5. Tanacetum potentilloides A. Gray. Cinquefoil Tansy. Fig. 5684. Artemisia potentilloides A. Gray, Proc. Acad. 6: 551. 1865. Tanacetum potentilloides A. Gray, op. cit. 9: 204. 1874. Sphaeromeria — illoides Heller, Muhlenbergia 1: 7. 1900. Vesicarpa potentilloides Rydb. N. Amer. Fl. 34: 242. 1916. Si hie very-canescent herbs, the decumbent or —— geunee leafy stems 1-3 dm. long, arising from a stout, sometimes branched caudex. Basal lea clustered, 5-9 mm. long, petiolate, the lades bi- or ‘tripinnately dissected into narrow lobes, én Gir tel sheila: few, resembling the | but less divided, mostly shorter than the tecnueee heads rayles, yello Yee several to numerous, short-peduncled, clustered at the ends of the stems and sheiers - involuc mm. broad ; phyllaries 3-4 mm. long, ovate to broadly ss hairy excent t for the br oat scarious »8 rgi covered. with pilose hairs ; achenes- obovoid, sc Rapier ‘swe elling ‘whee moist alkaline meadows, Arid Transition Zone; Wheeler Count y, Oregon, er ‘ie eastern California to iene County pre adjacent” Nevada. Type locality: Carson City, Newads June—Aug 102. MATRICARIA [Tourn.] L. Sp. Pl. 890. 1753. Annual or perennial, erect or decumbent herbs, often aromatic. Leaves alternate, bi- or tripinnatifid into linear rb Inflorescence corymbiform or the heads solitary on the sands of the branches. $ many- -flowered, sometimes radiate, the rays white. oe lucre saucer shaped, the phyla in 2 or 3 series, not much imbricated, dry with scari margins. Receptacle ed, spheric or long-conical. Marginal Stotlas cistillate, with or without white rays. Biiecarnlien yellow or green, perfect and fertile, tubular, SUNFLOWER FAMILY 399 with a gradually aio somewhat inflated throat, the lobes 4 or 5, ovate or lanceolate. Anthers entire or nearly so at base. pie branches flattened, ea minutely pencillate. Achenes glabrous, vai netticat: 3-5-ribbed or -nerved on one side, usually smooth on the other. Pappus core [From he Latin word mending emi so named for its supposed medical virtues. A genus of about 40 Moai natives of the northern hemisphere and also South Africa. Type species, Matri- caria chamomilla L. th on the outer face, thus lacking glands at its apex. 1, M. chamomiila. R tacle d , d; mat h g the outer face, this bearing 2 rounded glands at its apex. 2. M. maritima, Heads bearing white ray-flowers. R a 1 7 + te, neon discoid. appus-crown minute, entire, the achenes with 2 epee ara balay game the fone tes the home ns 6 ig Pappus- crown evident, with 2 short lobes or feath these bearing oblanceolate or ene glands sitting ae of the achene; matu e heads usually 10-11 mm. high. . M. occidentalis. 1. Matricaria age L. Sweet False-chamomile. Fig. 5685. Matricaria chamomiila L. Sp. Pl. 891. 17 — veolens ~ Fl, Suec. ed. 2. ee 1755. Leucanthemum chamaemelum Lam. FI. Franc. 2: ag 1778. pi ae Paton Gilib.. Fi. Lihue: Po 1781-8 Chamomilla vulgaris S. F. Gray, Nat. Arr. Brit. rm 2: 454. 1821. Matricaria obliqua Dulac, Fl. Hautes- Pye 505. 1867 Chamomilla chamomille Rydb. N. Amer, FI. 34: 231. 1916. Sweet-scented, piahrous, rather slender annuals usually much branched above, the stems 2-8 dm. high, striate. Leaves 2-6 c ong, bipinnatifid into linear-filiform divisions about as lo ong as or a little longer ean the internodes heads ppascdss on the upper branches ; involucres about 3 mm. high, 6-10 mm. broad; phyllaries subequal, m ostly oblong, broadly scarious, obtuse or rounded att apex; fa La 10-20, reflexed in age, 4-10 mm. gis: disk-flowers many, yellow, about ong ; receptacle ut 4 mm. high, narrowly dome-shaped; achene s asymmetrical, smooth except . 5 narrow, wie. like ribs ; pene none or sometimes a short c js A native onal around gardens and waste places st the Pacific States and occurring more of Eur scomeiatie § in eastern United a tate: also introduced in Arizona. May—Jun 2 epigic ty maritima L. False-chamomile. Fig. 5686. Matricaria pene ark: Sp. Pt S9h: 2755. Matricaria inodor Fi. vied ed, 2. 297. 1755. cane or pce leafy, coarse, ae -stemmed herbs 1.5-6 dm. high, glabrous or nearly so, branching above, not aromatic. Leaves 2-7 cm. long, much longer than the internodes, s . natifid, an leaf-segments 1.5-5 mm. by mostly linear-filiform ; ; heads iate, pedunculate; in- volucre 4-5 mm. high, about 11-14 mm. broad; phyllaries subequal, mostly oblong, rst at ee. with a broad scarious margin and central, dark greenish or brownish area; rays 12-29, owly oblong, 6-13 . long ; disk-flowers about . long, yellow, wit short acute 1 ; receptacle —8 . high, dot shaped chenes about 2 mm. long, metrical, with 3 y ergs roun em gi ribs od ee inner face, the ber brown and rugose, the o sai ttened, brown and and bearing 2 glands at ee apex ; pappus a minute denticalate’e ts ‘iibed of pura: introduced along oi Phas seaboard and oe savory eee the coast in Wash- ington and along the lower Columbia River in northern Oregon. July—Aug 400 COMPOSITAE 3. Matricaria matricarioides (Less.) Porter. Pineapple Weed. Fig. 5687. esse ey apne ishngssas Fl. Amer, Sept. 520. 1814. Not Matricaria suaveolens L. Less. Linnaea 6: 210. 1831. Matricaria discoidea DC. Prod. 6: 50. 1837. Matricari tri ides Porter, Men. Torrey Club 5: 341. 1894. Pineapple-scented, nearly glabrous annuals 1-3 dm. high, usually much branched from the base, the leafy striate stems divaricate or asce ane. < aves 1-5 cm. long, bipinnatifid with short, linear, acute divisions, longer than the ede qualling ony heads rayless rather stout peduncles, 0.5-1.5 cm. long; iain ede about 3 igh; phyllaries subequal, broadly , with scarious mar entral, brownish or greeni : k-fl r - long, yellowish green, 4 ; receptacle gh, narrowly dome-shaped to conical; achenes 1 , asymmetrical, usually ae sya’ oe on the inner face and having 2 marginal nerves, the outer fac th and usually ro , the margins bearing linear, brown, glandular lines ne eke or completely the scemtly of the body of the achene and not extending nto the minute pappus A common sO weed of waste gh ane cultivated exons throughout the Jig ne States; probably a native oF Voestera North — and ong hl sewhere in the United States and Europe. Type locality : Unalaska. Collected by Cham Rayless Cham en 4. Matricaria occidentalis Greene. Valley Pineapple Weed. Fig. 5688. Matricaria occidentalis Greene, Bull. Calif. Acad. 2: 150. 1886. Chamomiilla occidentalis Rydb. N. Amer. Fl. 34: 232. 1916. Rather stout (rarely slender), glabrous, not strongly scented () annuals 1.5—5. = dm. high, the single-stemmed or branching from the base, stems normally erect, branched above the middle with erect or ascending branches. Leaves 1-6 cm. long Baca with aes eee ere divi- sions, somewhat longer than the internodes or equaling them; hea ess, few to several on stout peduncles 1-3 cm. long; Sichese 4(3) mm, high; phyllaries sancaa broadly oblong to oval, with scarious margins and a central, reenish or brownish area; disk-flowers tig 1.25 mm. long, asi iel green, 4-toothed ; ‘receptacle about 8 mm. high beet in depauperate specimens, dome-shap: ometimes nar rrowly so phibe about 1 mm. long, asymmetrical, with usually 2 distinct, wing- like, whitish nerves on the i ng r face, 2 marginal aie are topped by the elliptic or oblanceolate glands extending to the crown, and 1 rather indistinct nerve on the slightly rounded outer face; pappus-crown a A short bist distinct lobes, each bearing brown glands that obits d slightly on the body of the In vernal pools and fields in prin pesca’ — top Joaquin Valleys and in valleys of ml Inner Coast Ranges ~~ Pag a weed in other Pacey Por ead County, Oregon, and Eureka, Humboldt Cou California, and in n California. Type locality: not definitely prin but collected by Greene in Contra vai and Solano Coun peti California. April-May. Much resembling M. oteicavisites but differing in the distinctive achene characters and growth habit. 103. SOLIVA Ruiz & Pav. FI. Peruv. Prod. 113. pl. 24. 1794. Small, diffusely branched annuals. Leaves alternate, petioled, bi- or sepinnately parted. Heads few- t to: many-flowered, —> e in the leaf-axils. Phyllaries thin, free, rotate, s sub- equal, in 2 series. Receptacle convex or low-conic, ee d. Ma al flower le heads either broad w, margina wings produced above (in ours) into cuspidate tips that are free from the persistent, spine-like style. [Name in honor of Salvador Soliva, a Span- ish physician. A genus of 9 species, natives of temperate South America. Type species, Soliva sessilis Ruiz & Pav. Wings of the achenes en BS on Wings of the achenes ea ceioinaly lobed near the base. ica Seroshiotid Soliva — Ruiz & Pav. Common Soliva. Fig. 5689.. Soliva sessilis Ruiz & Pav. Fl. Peruv. Prod. 113. ne 24. 1794. Gymnostyles chilensis cane Syst. 3: 500. 182 Soliva daucifolia Nutt. Trans. Amer. Phil. Soc. a: sf 403. 1841. Soliva microloma Phil. Anal. Univ. Chile 27: 331. 1865. Fibrous-rooted annual, stems few to s aca from the base, 3-20 cm. long, 2 ager oes tomously bravakins, the branches Sane thinly villous throughout. Leaves 1-4 c on brs Be dissected into narrow oblanceolate lobes, t the blade orbicular to oval in outline, the : 1 flowered ; phyllaries 2-2. a enw. broadly ovate and abruptly acute, thin, green above and hyaline below : Serta ip vere fertile, margi inal in 2 or 3 series; central flowers 2-4. sterile, subeylindric ; body of the achenes 2-2.5 or more long, — in — planoconvex, ispidulous on surfaces, sometimes becom glabrate, the wing rved, ver narrow to nearly 1 mm. wide at its ele width, ip be Sa and meat ces. sometimes 401 SUNFLOWER FAMILY 4 T lp 5685. Matricaria chamomilla 5686. Matricaria maritima { | Bag qe gag Rao EES qaq= = San £38 mow w 5687. Matricaria matricarioides 402 COMPOSITAE narrowed at the base but never notched, ence goo? ge the body of the achene with incurved cuspidate lobes much shorter than the indurated s native of Chile, well_established as a weed along - coast and adjacent ranges; Lincoln County, bi, southward in Cali uae rnia to Santa Barbara County; also adventive in lawns in southern California. Type locality Chile. April—Jun Soliva Deicivone was originally described by Nuttall from specimens collected at Santa Barbara in which the Sah: of the achenes are quite narrow. a width of the wing is quite variable and specimens with wide and with narrow wings are found in the same colo: vA Paps cat A (Juss.) Less. South American Soliva. Fig. 5690. yles pt Juss. Ann. Mus. Paris 4: 262. pl. 61. fig. 3. 1804. R. I oir. Sexy 6: 127. 1804. Soliva aad tle al Syn. Comp. 268. 1832. Soliva bare ma DC. Prod. 6: 143. 1837. Soliva se hen of authors, not Ruiz & Pav. Habit, height, leaves, and inflorescence as in S. sessilis. Body of the achene about 3 mm. long or less not including the indurated style, obovate in outline, planoconvex, hispidulous on both faces, the wings rin or incurved, each about 1 mm. wide at its greatest width, glabrous and somewhat cartilaginous, becoming thinner on a extreme edge, deeply notched near the bi se and extended above the body of the achene into cuspidate-tipped lobes that are shorter than the in- durated style. Occasional along roadsides and waste places; a South American herb introduced in the Sierra Nevada foot- ; mo elie ary yo es County to Tuolumne County y; also adventive in Australia and New Zealand. y 104. COTULA [Tourn.] L. Sp. Pl. 891. 1753. w, diffuse or creeping, strong-scented, annual or perennial herbs. Leaves oseloat a persisting on the receptacle, compressed, margined or winged. Pappus a short crown or A genus of about 50 species y on distribution in the tania and subtropical regions of the world. Type species, Cotula coronopifolia Leaves pinnately cleft or entire, with a sheathing base, glabrous. Pigs, ele olia. Leaves pinnately or bipinnately dissected, without a sheathing base, more or less villous. ram Sg s a coronopifélia L. Brass Buttons. Fig. 5691. Cotula coronopifolia L. Sp. P' 1753. Lancisia coronoptfolia Rydb. 5 Ray Fl, 34: 286. 1916. Rather ene glabrous ihe ge branching from the base, decumbent ane sh Oee root- ing from the s, 5-30 cm. long. Leaves sessile with a sheathing, ponent ous base, linear, lanceolate, or sem entire, Soothe. or with narrow lobes, 1-6 cm. long; = si bright af ellow, solitary from the upper leaf-axils, on peduncles 1-5 cm. long; eats flat-hemispher 5-11 mm. broad; phyllaries in 3 subequal series, oblong-lanceolate or ‘elliptic ; Faces srieal b istillate flowers a single series, corolla lacking ; disk-flowers tubular ; achenes or marginal flowers shorter than he pedicel, conspic cuously winged and e — ate above eal below, the checies of the disk-flowers ead epee vite ate, not winged; pappus none. ommon in tidal flats set _ coast o eae nland in wet places; British Columbia south to northern Lower ear: also on the Atlantic Gonet and a ak elsewhere especially in the southern hemisphere. A native of South Africa. ies ie 2 Cotula australis (Sieber) Hook f. Australian Cotula. Fig. 5692. Anacy stralts Sieber ex Spreng. Syst. 3: 497. 1826. Low slender annual 3-20 cm. high, diffusely peonching from the base, sparsely ie through- out. Leaves w ah 4 a petiolar base, pinnately or bipinnately dissected into linear or lanceolate, acute divisions 2-4 cm. long; heads te minal on the < hance pale yellow, on peduncles "23 ¢ m. long; vn pace places and about Se widespread oat iin i Se righ At and Cali- fornia; also eect from Utah and Maine. Introduced from Australia com —March. SUNFLOWER FAMILY 403 5692 5691. Cotula coronopifolia 5692. Cotula australis 105. ARTEMISIA* L. Sp. Pl. 845. 1753. Annual, biennial, and perennial herbs or shrubs, usually aromatic, the roots fibrous, or with taproot; perennial herbs often with rootstock or ca dex. Leaves entire to finely divided. Inflorescence paniculiform, sometimes reduced and raceme-like or spiciform. Phyllaries dry, imbricate, at least the inner ones s arious or with scarious margins. Heads small, discoid or hediocs Ray-florets pistillate and fertile or wanting; corolla tubular and usually tapered upward with 2, 3, or 4 teeth; style 2-cleft, more or less ex- serted, the somewhat flattened branches usually pines and rounded, or truncate and entire, or erose at the tip. Disk-florets perfect, fertile or the pistil sometimes sterile by an abortive ovary; corolla campanulate or unnelform, regular and 5-toothed; style usually eft i es elli ngled or -ribbed, or with numerous faint striae, usually glabrous or resinous-granu a te rarely hairy. Receptacle flat to hemispheric, naked or with many long hairs, with pale in only 1 species. [Name Greek, from ancient name of mugwort, in memory of Artemisia, Queen i solus. A circumboreal genus of a species, depending upon interpretation. Type species, Artemisia vulgaris L. Pistillate marginal florets present (may be absent in some heads of A. bigelovit). Disk-floret te per fees | and robe? Pp on Abrotanum). Plants re spas s a little Actes at the bas Plants perennial from rhizome audex, saaitibies taprooted. ave: concentrated ecally. pees and r du educed upward. Pubescence of leaves loosely villous or wanting; "thats — 2-5 dm. high. : A. norvegica saxatilis. Pubescence of leaves sericeous; plants mostly under 2 (carey - 3) dm. hig a hikecta or oe chiefly caulin L — divided, at least some of them ss some of the —— again ooth . A. michauriana sy et to bipinnatid with entire lobes, these usually mh broader han above. nvolucre higher than broad or height and breadth about eq Leaves ng cm. or less in width (rarely to 1.5 cm x agi of lobes when present; plants rarely over 1 m. tall. Plants Rael rata at base, tending to be ae well-devel- indleya Plants Herbaceous to a hs never taprooted, fret . reas by creeping rhiz 3. A. Indovictan Leaves Sym tall - =e cm. exclusive of lobes when scat pet often over 1 late, more or less tomentose. 4. a Sanglaens: Invol lindri narrow-ovoid, g : 4 suksdorfii. Involucre broader than high. as ane unalascheensts. * Text it i by George Henry Ward. 404 COMPOSITAE Pee 1 biennial, taprooted; 1 ially glabrous. 9. A. biennis. Sen gear Il shrub, 5-25 dm. high; leaves ternately to bipinnately divided into narrow linear lobes. 10. A. californica. Low spreading shrub, 2—4 dm. high; leaves narrowly cuneate, ——- : 11. A. bigelovii. Receptacl d with long hairs (Section Absint Divisione Sly leaves oblong or Hoon aie iia segments ——— 1.5-4 mm. ee: plants su 4-12 dm. tall. » A re Divisions fe =. linear, ultimate seg ts about 1 mm. wide; plants sual 1-4 dm. ‘ai A, frigida. Disk- i perfect but pistil infertile, the ovary os (Section Dracunculus), Plan Sesiien eet achenes essentially glabro ns ineuety entire, occasionally to 3- sane 14. A. dracunculus, ete finely divided int A — or linear-obl late lobes Pubescence wk t 1 silky, f 1 bescent or glabrate. Heads ‘relatively small, the ates usuaily 2-3 mm, tall ne, paniculiform, less condensed; ters to 10 dm. in height. 15. A. mpestris pacifica Heads posits ge involucre pues 3-4 mm. tall; Gy ercresnee pb often Soinifeen or nearly so; ges 1—4 dm. in height. fA. campestris borealis. tueawey loosely s villous, very dense over all parts of the plant yeh re 2 Ee coast f Cali sone and in so uthern Oregon. . A. pycnocephal: ‘ A, spihersbeae Plants shrubby wid spinescent; Pistillate marginal ee eR. all florets perfect al fertile (Section Seriphidium). ecepta ubs. naked; plants woody shr a ones nc or Looe — 3-7 teeth at the apex, or sometimes linear, entire and truncate unded (a few lea of A. ockii may be entire and acute). Moke poten ro comtcieg ony pre to large, 3-12 —— per head; bbe mg ad ~~ or saukewerieg shoots, or if such leaves present — uncate or rounded; leaves deeply divided only o: montane forms of A. arbus Leaves mostly t thre or more times as long as eh 1.5-5 ¢ long, entire or 3—7-toothed at apex, lin aired cuneate or with divergent 5eke ‘lobes; habit variable. Achenes glandular. -granuliferous, r arely sparsely short-villous; ; cuneate, 3—5-dentate metimes linear aie entire Pith a rounded or truncate apa 18. A. tridentata tridentata. Achenes arachnoid srerteritions, rarely less pubescent; ves ai with an obscurely notched, truncate or rounded apex, or with Giveteesn ‘obes at apex. 7 ae ee a. Leaves mostly as long or three times Pig long as wide, 0 long (spri ‘Ss sometimes longer ger), broadly cuneate or flabelliform, 3—5- ace 6 — “lobed at apex; | sie spreading arb Plants often root-sprouting; heads large, 8-20 — per head; often with many leaves entire re acute, leaves on often peony | ie: x eee y parted; found at Se clevations | in the Sier Nevada and San Bernardino Mou . A, rothrockit naa Lae 7 or dbianbeola te, entire prey an acute or acuminate Tae} or leaves deeply divided 030 e linear or li beapobtaneedlate lobes. be narrowly oblanceolate, jnoety. seve or ait a few irregular - or shes (some leaves may gth in some plants). » Asif Leaves ‘eauels divided into s or acess narrow lobes, entire linear leaves dai may he prone: At least the upper g their gh — Heads all exceeded by their subtending bracts; leaves deciduou “ A.r were _ — all or most of the florets subtended by bracts; plants ‘with sorubby ate ant long herba- A. pal 1. Artemisia norvégica subsp. saxatilis (Bess.) Hall & Clem. Mountain Sagewort. Fig. 569 Artemia arctica Less. Linnaea 6: 213. 1831. var. saxatilis Bess. in Hook. Fl. Bor. Amer. 1: 324. 1833. Artemisi r. pacifica A. Gray, Syn. Fl, N ry 12: 371. 1884, Artemisia sabicoke Rydb. Bull. Torrey Club 32: 128. 1905, Artemisia norvegica subsp. saxatilis Hall & Clem. Carnegie Inst. Wash. Pub. No. 326: 58. 1923. Artemisia norvegica var. saxatilis Jepson, Man. Fl. Pl. Calif. 1141. 19 25. Tufted perennial from a branching caudex, 1.5-6 dm . tall; plant loosely villous to essentia labrous. Basal leaves Souiteat, petiolate, the broad blade 2- 10 cm. lo ong, pinnately dis cr tata ed rrow and : i he ile ; narrowly rge; h moderately woolly-villous, 4-6 mm. high, the phyllaries dark-margined; ray-flor 12; disk- florets 30-80, fertile, the corolla Ao a at least near the base; achenes seach olinicic glabrous or granu iferous : recepta ae glabro Talus slopes, ack oute and o woods, Boreal Zone; Cascade and Olympic Mountains of Washington southward in the moutains oe ‘the ili Sierra Novel da, in the Rocky Mountains south to a north- ward through Canada and Alaska to eastern Siberia. Type locality: Rocky Mountains. July—Sep * Artemisia trifurcata Steph. ex Spreng. Three-forked Sagewort. Fig. 5694. rtemisia trifurcata nip ex Spreng. Syst. 3: 488. 1826. Artemis ribbons Nuov. Mem. Soc. Nat. Mosc. 3: 74. 1834. rtemisia tacomensis Fee ab. a Amer. FI. 34: 362. 1916. eee norvegica Pas heterophylla Hall & Clem. Carnegie Inst. Wash. Publ. No. 326:59. 1923. Artemisia trifurcata subsp. tacomensis Hultén, Fl. Alaska & Yukon 1577. 1950. SUNFLOWER FAMILY 405 oi : d usually yaa so Basal leaves ee natinlates: the blade "0. 5-3 cm. long, tr ri fid or pa alm beng i i more w and rather blun Vv ty a Te 7 6.7 j= *) ° ad o o> aw i: a eal Me ae oe SS one co - ro) ° jo) +7 oes ‘Sp w k=, yy & Se of ° jan o =] wn o. I 25 Pieaeehirants at the summit ; enes nearly ey lind. Cnaesel g-hairy ; receptacle glabrous. Open rocky ledges and talus poset at high elevations, Boreal ate pra diecgt nd Cascade be snnigs of Washington to aieikn. Menchurid, and Kurile Islands. Type locality: Soongaria, Scnchuris. July—Sep 5693. Artemisia norvegica 5694. Artemisia trifurcata XH Raate: ludoviciana Nutt. Western Mugwort. Fig. 5695. Artemisia Iudoviciana Nutt. Gen. 2: 143. 1818. grist gnaphalodes Mar loc. cit misia Intous ciana var. gnapha rs orr. & Gray, FI. fc Reng 2: 420. 1843. var. gnaphalodes sates Rev. Gen. Pl. Artemisia ute ris var. ludoviciana Kuntze, loc. cit. rtemisia vulgaris subsp. Iudoviciana Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 76. 1923. Artemisia vulgaris subsp. gnaphalodes Hall & Clem. op cit. 77 Artemi a ubs: eck, op. cit. No. 520: 330. 1940 Perennial herb, 3-10 dm. tall; stems herbaceous to base, slender to moderately gmp or nu ; g nsely white-tomentose on both sides or loosely floccose t and glabrate above, linear to lanceolate, oblanceolate or elliptic, pear or few-toothed or “tol sey espe- cially near the apex; scence paniculiform, us ll ther elongate and Seg the heads often in glomerules ; involucre ovoid to campanulate, 2-3 mm. wide, mm. hig ually densely tomentose, occasionally glabrate ; ray-florets 5-12; disk-florets Fil fertile ; gp a ellipsoid, mento: rate glabrous, sometimes resinous-granuliferous ; receptacle abcnes Lowlands and mountains up to middle elevations, Arid Transition Fat ne pper aaa Zones; Washington south to California and north and bias? to Alberta, Indiana, and Arkansas; rare west of the Cascade Mountains and —— Nevada. Type locality: e banks of the Mississippi River, near St. Louis. July—Sept. misia ludoviciana subs pe (Woot.) Keck, Proc. Calif. Acad. IV. 25: 446. 1946. carters microcephale ale ot. Sue ares Club 25: 455. 1898, not Hillebr. ri A. albula Woot. Contr. U.S. Herb, 16: 191 i. mostly 1-2 cm. long, lanceolate, en n both s invol mm. hig densely touseteaees ray-florets 8-11; disk-florets 8-13. Mountains of southern ‘California east to New Mexico and souther Bilge cshors south to northern Lower California, Sonora, and a huahua. Type locality: Organ Mountains, Dona aon County, New Mexico. May—Oct. Artemisia 1 ici bsp. candicans (Rydb.) Keck, Proc. Calif. —_ Ty. ig 447. 1946. (Artemisia ludoviciana var. latiloba Nutt. penne. A Amer. Phil. Soc. II. 7: 400. 1841; 4. candicans Rydb. Bull Torrey Club 24: 296, 1 A. latiloba Rydb. N.Y. Bot. Gard. 1: 429, 1900; a piaiyphotia Rydb. N. America 34: Psst 6; A. vulgaris subsp. cendicons Hall & Clem. Carnegie Inst.” ash eee 73. _v gar ie M. E. Peck, Man. Pl. Oregon 766. 1941.) Principal peg S40 ) long, more or less parted. po divi ded, often deeply pin mpatifid with some of the lobes again seme’ or oe Age on r w .5- fs es i Attemiais ludoviciana piles incémpta (Nutt.) Keck, Carnegie Inst. Wash. Publ. No. 520: 327. 1940. a gar erg Nutt. Trans. Amer. Phil. Soc. II. 7: 400. oo; A discolor var. incompta A. Gray, Syn. _ N. me ea: ‘A. atomifera Piper, Contr. U.S. erb. 11: 588. 1906; A. ludovtictana var. Bh vee E. “Jones, B Univ. Mont. Biol. Ser. 15: 48. 1310: es vulgaris subsp. michauriana var, tn- a St. John, esearch —S St. Coll. Wash. 1: 106. 1339.) Stems 3-9 dm. tall; principal leaves 2-8 cm 406 COMPOSITAE long re or less parted into linear or lanceolate, forwa rd-projecting lobes, some of these again toothed or lobed, uony glabrate — and white- gemma 944 aioe but sometimes tomentose throughout and rarely essentially Sect ar nei ; involucre 3-3.5 mm. high, the shy ltiteies sericeous-tomentose to glabrate and shiny ‘ide = dis ig. 30 or rarely penn Mountains of southern and eastern Cate rnia to eastern PS a Ae pin central Utah, giaen at somewhat higher sieeations than Iudoviciana subsp. candicans when erorine in the same area. Type locality: ‘“‘Thornberg’s Pass, és probably Custer County, Idah iri 8 sia vulgaris L. Sp. Pl. 848. 1753. ta ecipe Aso otaigigey herb; stems 5-15 dm. tall, erect, simple or Ab ke d poy neoay in small clumps from “ = llow leaves ‘sm ooth, green, ‘mek glabrous or nearly so above, densely white-tomentose beneath, the eel pane pos ates. 5-10 cm. iong, 3- i; m. wide, obo sa or erage Sey ancl in outline, cleft into shequat lobes of hha the eh oo is the lar. and which are again cleft oothed; ewer gonns paniculiform, leafy pro or occasionally reduced ant compact; favelner’ cavalry gros 3.5-4 Binh, more or “meg hiectase 6-12 pistillate — -florets, 8-18 perfect and fertile fies encness: Mate it be, sein =8,;,9. pe rae hp ths established in eastern United States ~ pte adjacent Cana da, but in our area ‘cnaak only from near ballast dumps at our ae ee ports. Type locality: urope ws be bs saig douglasiana Bess. Douglas’ Mugwort. Fig. 5696. Artemisi i Bess. in Hook. Fl. Bor. Amer. 1: 323. 1833 Artemi. alifornica Bess. Linnaea 15:91. 1841. aes heterophylla Nee Trans. Amer. Phil. a II. 7: 400. 1841, in part. Not Bess. 1834. var. douglasiana D, C, Eat en King by 183,~1871; pire kennedys A. Nels. Proc. Biol. Soc. Wash. 1 175. is subsp. heterophylla Hall & Clem. eit te t, Wash Publ. No. 326: 76. 1923. 925. 4rtemisi var, heterophylla Jepson, Man. FI. PI. Calif, ser Artemisia onepaeis subsp. douglasiana St. John, Fl. S.E. Wash. Perennial herb, 5-15 dm. tall (sometimes taller) ; stems herbaceous to base, simple or with erect to dicee ee bran eg Leaves lanceolate to elliptic and entire, or obla nceolate to obovate and coarsely few-lobed or -toothed sliag the apex, 7-15 cm. long, 1-5 cm. wide exclusive of salient lobes, gradually reduced in inflorescence, all sparsely tomentulose to cabeane an ce, gray-toment ; inflorescence paniculiform, lea open or dense and elongated, he heads erect or n nodding g: involucre Campanula ate, mm. igh, 2-3 mm. wide, the phy aries more or shinee Ag Zones; Washington south to northern Lower California; rare east of the C an axis except near Lake Tahoe and rarely to 6,000 feet elevation. Type locality : “North-West jee pet gg toe —Oct. 5. Artemisia suksdorfii Piper. neat or Coastal Mugwort. Fig. 5697. a heterophylla Nutt. Trans. Amer. seis Soc. II. 7: 400. 1841, in part. Not Bess. 1834. Artemisia vulgaris var. si Suksd. Deut h. Bot Mona 18: 98. 1900. pcaenets hee dorfii Piper, Bull. Torrey Cha me Artemisia vulgaris subsp. Borelis Hall & Cle gees Ane Wash. Publ. No. 326: 76. 1923. rennial herb 6-15 dm. tall; stems stout, herbaceous mewhat aun ruieserat at base. Leaves oh with m habe narrow heads; involucre terete or nearly ovoid, 3-4 mm. high, the p hiyllaries fis ligh abrate, yellow-green and a ing ; ray-florets 37: disk-florets 2-8, fertile; op eae Sines paket: receptacle glabrous. ants Coastal a roarigers and Humid Transition Zones; Vancouver Island pacthwand along the coast and some- what Pans along some of hes re to Sonoma ged California, not ascending to over 300 feet elevation. Type locality: omer sea-beaches near Fairhaven, Whatcom County, Washi wat a 6. Artemisia tilésii subsp. unalaschcénsis (Bess.) Hultén. Aleutian Mugwort. es 6 pina tiaberted var. vulgatissima Bess. in Hook. Bor. a 1: 322, 1833. ? lgaris var. americana basi Ltaitaes 1s: 105. Artemisia tilesii var oe Bess. o Artemisia tilesii var. r. & Gray, Fl. N. Amer. 2: 422, : 1843; ? Artemisia obtusa i oy er. Fl. 34: 274 yd : Artemisia tilesti var. elatior f. pubescens G. N. Jon Aleks. Wash. Publ. Biol. 5: 254. 1936. Artemisia — hawt unalaschcensis aise Fl. pele Isl. 327. 1937. ? Artemisia ana var. americana Fernald, Rhodor iti ae 1945, in part. Artemisia phi a elatior Hultén, Pg Soma & Yukon 1950, joatalee ©. Coke dm. tall; isaed her os o base, floccose when young sa soon glabrate. Leaves long | lin r-lanceolate to ches ndlate ovate or s in outline, the narrowest sometimes entir Snare ew d or -lo t ply pinnately te subp Imately , some of the nts again toothed or parted, tom e y tomentu- lose above, those of the oo “sare or nearly so; inflorescence paniculiform, more or less narrowed and spike tke gconiby ceeding the eavex the heads erect or noddin ng; involucre cam- panulate or tu te, 3- a gh, mm. wide, the ee purplish or sometimes green, glabrate or tomentulose ei tav- flores 'B-15: disk-florets 20-40, fertile; achenes Patiineniit: glabrous ; brous. Open, rocky or Pavctly sites, Boreal and Transition Zones; northern Oregon east to Montana and north -Hudson’s Bay, southern Alaska, the Aleutian Islands, and Japan. Type locality: island of Unalaska. Fuly Sept, SUNFLOWER FAMILY 407 cal A. tilesii is more northerly and has a shor more compact inflorescence which is usually over topped 23 the leaves. In parts <7 our area A. tilesti vor al unalaschcensis seems to intergrade to some extent ith A. Iudoviciana subsp. candicans and with A. douglasiana. 7. Artemisia lindleyana Bess. Artemisia lindleyana Bess. in Hook. FI. Artemisia lindleyana var. is ifolia Bess. ‘a Columbia River Mugwort. Fig. 5699. Bor. Amer. 1: 322, 1833. it Artemisia lindleyana var. subdentat Besa: in Hook. loc. cit Artemisia lindleyana var. corono Be n Hook. loc. cit. Artemisia pumila . Trans. Amer. Phil. Soc. II. 7: 399. 1841, Not Link, 1822. Artemisia arachnoidea Sheldon, Bull. Torrey Club 30: 310. 1903. is ibergii Rydb. N. Artemisia vulgaris var. lindleyana Jepson, Man. FI. Pl. Calif. 1142. Stems several, 2-7 dm. tall, suffrutescent at the base. Leaves linear-oblanceolate, rarely wider, 2-5 cm. long, mostly under 1 cm. wide, entire or with a few teeth o rrow lobes, hite-tomentose beneath, u sually green and glabrate i inflorescence paniculif ort, narrow, anc spike-like; involucre campanulate, 3 mm. high, 2-3 mm. wide, the phyllaries usually lightly iB 4 Chess, Carntale Inst. Wash. Publ. No. 326: 79. 1923. 1925. Ne. pe a UA kedortii 5695. Artemisia ludoviciana 569’ 5696. Artemisia douglasi 697. 5698. Artemisia tilesii 408 COMPOSITAE eee: ray-florets 5-9; disk-florets 10-30, fertile; achenes ellipsoid, glabrous; receptacle glabro oe res of streams below the high-water level, Transition Zones; mostly along the Columbia River and its hie sepa in Washington alt cnn sporadically to western Montana and southern British Columbia. Type ocality: ‘“‘North-West coast of Am July—Sept. ne sia Bdge otti +4 Bess oa Hook. Fi. Bor. Amer. 1: 1833. Differs from A, lindleyana py e fra d filiform chee tas than 1 mm. wie, green, the ‘Gane closely revolute. This is the for Saal found in the poo pars a the Co lumbia nie Tah Plants with characters intermediate between this and 4. dleyana are also com- this area, and it is probably at best only a variety of that species. Type locality: bi West America.”’ 8. Artemisia ceurigcanaai Bess. Michaux’s Mugwort. Fig. 5700. Artemisia michauxiana Bess. in Hook. + Bor. Amer. 1: 324. 1833. Artemise discolor r Dougl. ex Bess. Balt oe Nat. Mosc. 9: 46. 1836. subsp. discolor Hall & Cle em. Carnegie Inst. pie Publ. No, 326: 74, 1923. Artemisia teidears ris var. discolor Jepson, Man. FI. Pl. Calif. 1141. 25: — —- — machonstone St. - hn, — Stud. st aa: Wash. 1: 106. 1929. Artem ulgar var. typica St. John, loc. Artemisia discolor var. ‘and i A gece: Rhodora 32: 27. po dulifer E. Peck, Man. Pl. Oregon 766. 1941, Pe cnet ne & fe dm. a sometimes taller; stems several from a woody caudex. Leaves eee beneath at least me n young, genera ally glabrate and green above, fier crowded, 2-5 cm. long, bipinnatifid with secondary lobes again toothed, the lobes linear, widely spreading u ead mm. e, the phyllaries se or Se atag ts iene ose; ray-florets 9-12; ye ia 15-38, fertile ; “achenes ellipsoid, glabrous ; receptacle a aneo sites in the mountains, cei 8 ae fos a ae Mou ns of Washington north to British Columbia, Alberta, and northern Utah, also spr adios ily, in soulanitare. pind California, and Nevada. Type ocality: ee Kettle Falls and sources of the Cobsunibis. ” May—Aug. This s es, at least in the southern part of its range, seems to intergrade somewhat with A. Iudoviciana subsp. secseta. 9. Artemisia biénnis Willd. Biennial Sagewort. Fig. 5701. Artemisia biennis Willd. Phytog. 11. 1794. annual or biennial herb, 3-30 dm. tall ; stems simple below, erect, striate, glabrou € seg » © us, broader, the outer phyllaries narrow an e inner ones with only a green nian 2 ind broad scarious margin; ray-florets 6-25; disk- aes 15-40, fertile ; sain ellipsoid, glabrous; recep- rous. waste places and stream banks, Upper Sonoran and Transition Zones; introduced occasionally throughout our area and eastwar - AS. weed; native in British Columbia and the northern Rocky Mountains. Type locality: erroneously given as “New Zealand.” Aug.—Oct isia 4nnua 4 apr rooted Sania’: 3-30 dm. tall; stem simple below, erec flexuous, striate, glabrous. Leaves petiolate, 2-10 c¢ long, glabrous, pinnatifid to se coe 8 the segments lanceolate or acing ludigeeecente broad and open, selon, Pang - id the heads pe led, =e involucre glabrous, 1-2 mm. highs slightly broader, outer phyllaries row and green, the | er ones with oa “ a green midrib and a broad ir us =e in; ray-florets 5-10; disk- pad a0, Fig) bs anes nartowly turbi- nate, glabrous; receptacle gla’ oraoe. = 9. Fields sid waste pistes) nati asia, naturali gel Sal ae and astern United States, rare and orohahty not established in our area, Ty oe nv re Siberia. 1 Sage- wort se Artemisia californica sy Coast Sagebrush or Old Man. Fig. 5702. Art 7 lifornica Less. Linnaea 6: 523. 18 Artemisia fisheriana Bess. Nuov. Mem. Soc , Nat pres 33 21. 1834. Artemisia Gikchiane eS Bess. op. cit. Arteta poneee Nutt. Trans. Amer. Phil. oe Il. 7: 397. 1841. id. utt. op. cit. 399. Chrossostephium foliosum Rydb. N. ee ne 34: 243. 1916. Chrossostephium californicum Rydb. Rounded sh sem ‘ab higher ; stems ac ected the older with brown fibrous bark, the twigs erect, stout, striate, and — red witha nearly white, dense canes- ce. Leaves sessile, terna ates Bsa ately, or sometimes bipnnately divided into long narrow segments 0.5-1 mm. wide, minutely vag sae cinereous or canescent, with strongly revolute margins, usually some borne in pic n the axils of others, fascicled and upper leaves. reduced mm : s a thick back and broad scarious margin, canescent ; ae -florets 6-10: pied florets 15-30, fertile; achenes ee Ween, wie minute oeacinn crown, resinous-granuli ferous ; receptacle es- gis / glabro Exposed uae Upper Sonoran and Arid Transition Zones; Coast Ranges from north of San Francis Bay south to northern L, wer California and on adjacent ora extending eastward in southern California pf edge of Colorado Desert. Ory ype locality: California. Aug.-Oct — californica var. insularis (Rydb.) Munz, Man. S. Calif. 575. 1935. (Crossostephium insulare Rydb. N. er. FI. 34: 244, 1916.) Differs from A. californica var. californica in that the leaves are little or SUNFLOWER FAMILY 409 not at all revolute, the lobes therefor mm. in has P lobes also generally fewer; ray-florets to 15, disk- aaiag! = 40. San Clemente Island, ge nge Cou n Nicolas ee sy Ventura County, California. cality: ‘“‘along Pot’s Trail, San Clemente tinea. oi laecied by Blanche Trask. 11. Arfeaisis — A. Gray. ict Sagebrush. Fig. 5703. Artemisia bigelo . Gray in Torr. . R. Rep. 4: 110 Artemisia aplephils Wack & Standl. "ene US oNat Mati. ys 193, 1913, ergreen shrub, 2-4 dm. tall; stems many, spreading below and sometimes rooting when i paid brown, sheathing or becoming shredded on the larger in Ghatact en the soil, the bark branches, twigs densel scent, striate. Leaves of receecGe shoots sessile A a narrow base or short-petioled, narrowly cuneate, 1-2 cm. long, 2-5 mm. wide, -toot e truncate pex or sometit entire; leaves of the inflorescence mostly entire ear a re sae te, or gtd obtuse; floweri — , erect or the ti oping, nar culiform, den he heads several on h short recurved Biche involucre turbinate, wae ' ; broad, ay” inner phy ae ae as lon wee all densely tomentose, the ma rgin s scarious ray-florets 0-2, usually th listing’ ae bre t in heads of i p08 plant; disk-florets 13 us three Em fer tlle: aes ellipsoid, glabrous ; eas cle glabrou a 3 me; occurs in California at the east end a < paeure Desert, to Colorado in western Texas. Aug.—Oct Rocky soils, Ups r Sonoran Zon and Texas. Type locality: ger the upper Canadian” 5699. Artemisia lindleyana 5701. Artemisia biennis 5700. Artemisia michauxiana 5702. Artemisia californica 410 COMPOSITAE 12. Artemisia absinthium L. Absinthe or Wormwood. Fig. 5704. Artemisia absinthium L. Sp. Pl. 848. 1753. ial herb sometimes slightly woody at base, 4-12 dm. tall; myer stems clustered, — ‘below striate, sericeous or glabrate. Lower leaves weak Bc tiolate ee e 3-8 cm. long, Vv profuse, leafy ; involucre hemisphe -3 mm. high, finely ana densely an s, the o phyl- laries linear, the inner broadly elliptic with broad scarious margins; ray- 5 9-20; disk-florets 50, ferti Se Spe nearly cylindric, narrowed at the base, pies ie de recendacle thickly beset with long hai =, 9: Fields pe waste places, Arid Transition Zone; introduced in eastern Oregon and Washington eastward across the continent; native to Europe. July—Sept. 13. Artemisia frigida Willd. Prairie Sagewort. Fig. 5705. Artemisia frigida Willd. Sp. Pl. 3: 1838. 1804. Absinthium frigidum Bess. Bull. Soc. Nat. Mose. 1: 251. 1829, erennial, mat-forming herb often woody at base and even somewhat shrubby, 1-4 dm. high, fragrant. Leaves small and numerous, crowded at the base and well distributed along the year sericeous-tomentose, twice or thrice ternately divided into linear or linear-filiform lobes 1 m or less wide, usually with a pair of trifid, stipule-like divisions at the base, petiolate or the sear ones subsessile, the blade 5-12 mm. long ; inflorescence paniculiform or much reduced and racemi- form on depauperate plants, the heads sessile ie shone epee’, nodding ; involucre h Sag ene 2-3 mm. high, the phyllaries loosely tomen ray-florets 10-17; am Horets 25-50, fertile; achenes pS Saba glabrous ; peeptacte thickly” beset with long hai tik n places in plains and foothills, Arid Transition and Boreal as northeastern Washington to Ariz ne, Wisegnsin, Alaska, aaa Siboiay oes Fiabe i of the high plains. Type locality: Davuria, eastern Siberia. July—Sept. 14. Artemisia dracinculus L. Dragon Sagewort or Tarragon. Fig. 5706. Artemisia dracunculus L. Sp. Pl. se — esha A annie Pall. Ps Willd. Sp. P 1804. neuloi Pursh, FI. Siok Sar. 742. 1814. Arecenieie precenentes var. glauca Bess. in Hook. Fl. Bor. Amer. 1: 326. 1833. a A. Nels. Bull. Torrey — 27: 73.1906; Artemisia ar guage ge typica — & Clem. maberee Inst, Wash. Publ. No. 326: 115. 1923. y u op. cit. Artemisia dracunculoides var. glauca soils, Man. S. Calif. re 1935. Perciibiad herb, 5-15 dm. tall, sponely ecor rous to nearly inodorous; stems several but not crowded, erect, simple to — mi cen Leaves linear to somewhat lanceolate, mostly 3-8 mm. lon mm e, gla s or Sea Licteally ville: oubertilest and glabrate, entire or a few of them saat inflorescence ne m, leafy-bracted and pss ascending brake the heads gen- erally no re hemisphe sok, 2-3 mm. high, 2-4 mm. wide; ray-florets 6-30 ; disk-florets ees the oistil esate acbeae glabrous, tps of the disk-£ florets abortive ; receptacle glabrous s ne et Open places, plains to moderate elevations in the mountains, Arid Tra on Zone; eastern isabel south to Lower California, to Texas, Illinois, Yukon, and Eerania. Type a Siberia. July—Oct 15. Artemisia campéstris subsp. pee (Nutt.) Hall & Clem. Pacific Sagewort. Pe } . douglasiana Bess. in Hook. Fl. Bor. Amer. 1: 325. 1833. rtemisia desertorum var. scouferiana Bess. in Hook. loc. cit. Artemisia pacifica Nutt. Trans. Amer. Phil. Soc. II. 7: 410. 1841. Artemisia scouleriana Rydb. Bull. Torrey Club 33: 157, 1906. rtemisia camporum Rydb. N. Amer. tye 34: 254. Artemisia campestris var. pacifica M. E. Peck, Man. PI. Oreanet 768. 1941 rtemisia campestris var. scouleriana ~ eaeing 7 sales s West. Bot. 7: 20. 1953. Perennial herb, 3-10 dm. high, with somewhat oe hi acing ce e; stems several to numerous, ti very leafy below, somewhat leafy to ie inflore: es bipinnately divided into nar- w linear iecenn about 1 mm. wide, edgpaion or ie ak “glabrate, 4-12 cm. long including the sah reduced upward; infloresc gs culiform, usually with numerous closely a oa branches, the ye horizontal or erect, n Pon nly w en se al ; involucre hemispheric, 2-3 ei about 2. m. wide, the sieflatics es greenish and glabrous or sparingly tomentulose; a florets 8-20; disk- florets — the pistil sterile; achenes glabrous, tioae of the disk- florets sco — cle glabro ces, usually in sandy soil, Canadian and Transition Zon d Washington to Yukon, hia Rive Nebraska ene Mexico, Type 1 peels y: “shores of the Pacific ° ake yen! of the Oregon [Colum- rtemisia campestris subsp. borealis (Pall.) Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 122. 1923." (Artemisia phased Pall. Reise 3: 755. 1776; A. spithamea Pursh, Fl. Amer. Sept. 522. 1814; A. bore- is var. purshii Bess r. Amer. 1: 326. art) A. borealis var. wormskioldit Bess, in Hook. = cit. 327: rd hongalte var. besseri Torr. & Gray, Fl. N. Amer. 2: 417. 1843; A. borealis var. spithamea SUNELOWSS FAMILY 411 & Gray, loc. cit.; A. bargelie subsp. wormskioldii Pip Contr, U.S. Nat. Herb. 11: 587. 1906; A. ripi- co. bony Ry db. N. Amer. FI. 256. 1916; A. comer Reed pee BE Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 123.1923 a — ris var. —— Pens E. Peck, Man. Pl. Oregon 768. 1941; A. borealis € wormskioldit nee ‘Ls nadien 71 4: A, campestris var worauniatae Cronquist, Leaflets West. Bot 1950.) Miriwwapioe gc wm 1-4 hig oe often with . ——— base; stems crowded, erect r twice tern ely divided into or oe pea leafy below, less so up into the oe ality ea o ¢c mostly linear or linear-filiform lobes or sometimes pinnately divas, sot tig 4 sericeous or villous, the upper surface sometimes glabrate, 2-10 cm. ong, 0.7-4 cm. wide, reduced upward, the uppermost often entire; in- florescence spiciform, recemiform or narrowly paniculiform, ‘the tas is erect to nodding; involucre hemispheric, 3—4 mm. tall, 3.5—-5 mm. broad, the phyliaries brownish or yellowish green with a brownish median line, glabrous to densely villous; ray-florets 10-25; disk-florets 15-30; achenes glabrous, ese of the disk-florets abortive receptacle glabrous. Open places in sandy soil, along the Columbia River between Oregon and tc hewn oan “2% in the Cascade Mountains of Washin es to northern Montana and northward, aid in Siberia. Type loc Siberia Art sia campestris ape age (Michx.) Hall & Clem. Carnegie Inst. Was Pubr No, 326: 122. 1923. (artemisia se Mic Bor. Amer. 2: 129. = A, ptr ee Bin Wat Proc mer. Acad. 25: 133. 1890.) Root koscapere 5 stem plea single; leaves jin o r plants m ostly canescent gs es young, glabrate; otherwise pret like subsp. pac A few plants of this are gs und in "the Sr t Sound and Columbia Gorge areas; common from PRoeky Mountains eastward. Type locality: sandy banks of ath Missoari River. The plants here included in A. —— Ti er eg to a number of different races. The e ntire group is 0 ood and i A campes: - come established North America only in a eastern pets ur western plants are mostly s of two taxa here referred to as A. campestris subsp. borealis and A. campestris subsp. pacifica, which have been variously treated as sectiek: “subspecies, and varieties, a even split into a number of species. imi ll i m ment of a sete: rg? po Uy The form with gate as Peete is separa some as a separate ee (A. spithamea Pursh, A. bore alis var, purshit Ege: Ber. ig nly “spithamen (Posey all m.), ee to be ogra ifica’ characters. he Sentara River ‘form also hag been cacitel erate with more ju on, od lea ecological grounds—as 4 . ripico ole Ryd oe A. borealis var. wormskioldti Bess., or A. campe on oy wo mebioldit (Bess.) Cronquis igi emisia dracunculus 412 COMPOSITAE i nagiicnmen pycnocéphala (Less.) DC. Beach Sagewort. Fig. 5708. Oli. Less. ee 524. 1831. Artemisia phi stiahding DC. paige 6: 83 Artemisia pachystachya he 0 é rt Artemisia pycnostachya Nut rire “A r. Phil. Soc. II. 7: 401. 1841. rtemisia campestris ey PERE "Hall & Clem. > tenes Inst. Wash. Publ. No. 326: 123. 1923. Artemisia campestris var. pycnocephala M. E. Peck, Man. Pl. Oregon 768. 1941. Perennial herb, 2-6 dm. high; stems several or numerous on a stout woody caudex, erect, or ascending near the base, net! leafy up to the inflorescence. Leaves twice or thrice pinnately di- vided into narrow, linear or linear-spatulate divisions, densely “ana permanently silky- Sh stor ae cm. long, reduced upward inflorescence paniculiform but very dense and spike-like ith sile and erec Ce i pheric, 54. mm. high. oot 3 as broad, the vabptie Hes ee. villous ; ray-florets 8-15; disk-florets 12-25, the pistil sterile ; oye aabeylndric, glabrous, t of the ‘di sk- ea. abortive ; receptacle glabrous. n= 9. n beaches, Humid atone Zone; from central Oregon eouek to Point Sur, California. Type San locality : Cattenie Aug.—Sept 17. Artemisia spinéscens D. C. Eaton. Bud Sage or Spring Sagebrush. Fig. 5709. nna desertorum Nutt. Trans. Amer. Phil. Soc. II. 7: 417. 1841. Not Artemisia desertorum Spreng. 1826 Artemisia spinescens D. C. Eaton, Bot. King Expl. 180. 1871. Rounded shrub, 0.5-5 dm. tall, args — aoe and much ee from the base, mer ta a spiny by the itis eae of the of prev Set inflorescences. Leaves up to about 2 cm. long including the petiole, a a 3- a 5. Jobed , the ‘lohes Dace cleft into linear- spatulate ra oa esedll villous, reduc ed and less divided upwa ; inflorescence racemiform, 1-5 . lon me leafy-br. ih sometimes Sodeten to a single head; involucre -broadly turbinate, 2-3.5 ide, oe scarcely unequal phyllaries thick and herbaceous with narrow seatigue margins, pt villous; ray- s 2-8, the corolla long-hairy; disk-florets 5-15, the cor long-hairy, the pistil stile style undivided with an expanded penicillate summit enes olla -hai ellipsoid, densely arachnoid-hairy, those of the disk-florets abortive and essentially absent; re- acle glabrous lains and hills, Sonoran Zones; southeastern Oregon and eastern California north of the San Bernardino Mountains to South westeea Ricgiisna and New —— ee locality: ‘‘Rocky Mountain plains, in arid deserts, towards the north sources of the Platte.’ Apri Jun 18. Artemisia tridentata Nutt. . 3am Fig. 5710. Artemisia tridentata Nutt. Trans. Amer. Phil. Soc. IT. 7: 398 41. pidemencs tridentata var. rt ha A Gray, Proc. Amer. site} 19: 49. 1883. temisi ta Rydb. . Am 1, 34: 283, Artemisia vaseyana Rydb. Artemisia tridentata subsp. — ‘Hall & Clem. Carnegie Inst. Wash. Publ. No. 326: 136. 1923. Strong-scented evergreen shrub, rounded or somewhat spreading and flattened on top, rarely root-sprouting, go — sually 3- 30) ‘ten, high, with a single short trunk or with few ascending branches from th older stems geen with ee ie or light brown bark, younger stems covered Fc : se nse tomentum, striated. Leaves of vegetative shoots ante or short- s 1-4 cm. long o ey sate —1. sigs wi e mostly th tim ong of the Hatertiae shoots the same or more often entire and linear to tactolate. rounded t ; inflorescence well exceeding to barely exceeding the leafy shooth narrowly to broadly panieuliform, bi e or sometimes drooping ; involucre .8 mm. ‘igh 2-2. 8 mm. wide, the outer phyllaries short, orbicular-ovate, canescent, the inner elliptic, what Bo igh fee so, obtuse, ei cent or sometimes nearly glabrous; ray-florets absent ; disk- Horcts usually 3- ras many as 12 in some high-elevation forms) ; achen a ee alah turbinate, resinous- eater van rarely sparsely short- us ; receptacle glabrous. a re On dry plains and hills and apne ap into the mountains to timberline mostly on rocky open ground, Upper Sonoran Zone to Hudsonian Zon hy Poe sn and palifors ia mostly east be the err axis into the Coast a of southern California to northern Lower California and to the w n edge of the reat Plains. Type ef the Oregon [Columbia], “a eoty tSnake) River "July—Oct. A om eh polymorphic sin . wk = eat v vartahion and numerous ecological races, some of which may be int ti Sag acs, if s affies aa correlation of ecologic and morphologic euatacters’ an be discovered. To ate this ae Sy Senate aban. ge Sioa hes Gray) Hall & Clem. Sespenie Inst. Wash. Publ. No. 326: 1923. gg nds goons A. Gray, mer. Acad. 17: ae one tridentata var, pene Jepson, Man. Fil. Pl. Calif. 1140. 1925.) Rounde z go Moe ne shrub, usually 10-20 dm. Ae ine leaves mostly mond or narrowly spatulate, entire or shallowly notched or toothed at the apex ee frequen ntly with 3 short tees t lobes, rarely more deeply divided or pinnately 5-lobed; achenes glandular ps. reray Short villous to. “arachnoit hairy; otherwise much the same as A. tridentata subsp. tridentata. n= 18. Western Mojave Des of California, oan toward the Pacific Coast through the Santa Clara Valley. Type loca ae “Newhall, Los Angeles 19. Artemisia arbiscula Nutt. Dwarf Sagebrush. Fig. 5711. Artemisia arbuscula Nutt. Trans. Amer. Phil. Soc. II. 7: 0 ys Artemisia tridentata subsp. arbuscula Hall & Clem. Carnegie Inst gta No. 326: 138. 1923. Artemisia tridentata var. arbuscula McMinn, Ill. Man. Pion Sig sta esi SUNFLOWER FAMILY 413 Low, spreading, evergreen shrub, 1-4 dm. high, not root-sprouting, odor strong to mild, bark light bro wn to nearly black, aap on older aspen twigs densely canescent, often becoming edd nearly glabrous in late summer, light greenish yellow or reddish brown, striate d. Leaves of vege- tative ose roadly dese i cs flabelliform, 0.5- 5. em. long, 0.3-1 cm. wide, length one and one-half to three — (usually about twice) "ae width, usually 3-5-toothed but sometimes deeply divided i 3-5 v lobes o ely leav rec ent; leav f flowering shoots tr a few es ent Ww SC eiuceidoapeciak, chs entire to cuneate with a truncate, 3-toothed apex, or sometimes more deeply divided with 3 narrow lobes; inflorescence spiciform to nar bel paniculiform, when ewe ig branches few and erect; involucre narrowly to i. y campanulate A —3 mm. — aan the earn canescent or the innermost sometimes near iy glabro ; ray-flor abse nt; disk. rets or sometimes fewer in Se 1 wale ie fertile ; po Nai celine. pia iainng resinous— pate ct aienink cle glabrous. n = 9, 1 Usually on rocky or gravelly sit nd Sonoran raid bed ransition, and Canadian Zones; arid plains and hills and rocky openi i e mo ia from Mono County northward, eastern Creme. and isolated stations in the bichiirmtnabil ks, porecel of Washington kod the Siskiyou Mountains and Coast R ears of California eastward | to southwest Mon and northwestern Colorado. Type locality: ‘‘arid plai Lewi oo i Ronan Aug.—Oct. Art sia arbus subsp. ndva (A. Nels.) G. Ward, Con Dudley Herb. 4: 183. 1953. (Artemisia nova A. "Nels. Bull. 5 poten Club 27: Fring ie A. tridentata pel nova Hall & Clem, Carnegie Inst. Wash. saat No. 326: 137. 1923; A. triden var. nova McMinn, Il. ag Pewee Shrubs 608. 1939; A. arbuscula var. nova Cronquist, Vasc. Pl. Pacif. “Nort w. ie: 58. 1955 .) Differs m A. arbuscula subsp. arbuscula prin- us or rarely with a i r p shiny; there is als so a tenden ney, sacliculisls to the east of our range, toward leaves of a ar| greenish color and very glandular; 8 hog also average somewhat smaller ie. in the typical subspecies 1 East of the Sierra Nevada from Mono County, California, southward in the San Bernardino car dane Desert tas to northern New Mexico and southern Montana. Type Goality: Medicine Bow, Carbon County, Wyoming 5707. Artemisia campestris 5709. Artemisia 5 5708. Arte: inescens misia pycnocephala 5710. Arte: ke iden 414 COMPOSITAE 20. oeneigy tripartita Rydb. Cutleaf Sagebrush. Fig. 5712. Artemisia trifida Nutt. Trans. Amer. Phil. Soc. II. 7: 398. a Not Turez. 1832 Artemisia 'tripartita mae Mem. N.Y. Bot. Gard. 1: 432. Artemisia tridentata ahs trifida Hall & Clem. Carne . os . Wash. Publ. Mo,.326:_137:; 1923; Artemisia tridentata trifida McMinn, Ill. Men. Calif. Snrace 608. 1939 A low ad evergreen shrub, 2-8 dm. high with a shiek central trunk or with numerous br anches : ssbstbudiver from the base, frequently root- cbircastiags mildly aromatic, bark light brown or grayish, shredding on older branches ; twigs canescent. Leaves of vegetative shoots 0.5-3 cm. long o r rarely onger, canescent, sorely ‘divided into 3 linear or narrowly linear-lanceolate lobes i i mselves be 3-cleft or som i i meng g paniculiform; involucre campanu age no nig -3 mm. wide, the phyllaries canes- cen ci uter ones broadly ovate: sometimes with a w herbaceous tip, the inner ones oblong ray- “hon: absent ; disk-florets 4-7 (r: ane ee, fertile ehenes taste or nearly cylindrical resinous- beam uliferous ; sation glabrous. n= 9, 18. Dry hills and ohana Upper Sonoran and aie Transition Zones; eastern Washington and no rtheastern Oregon, through noe yout Ida = Wg Wyoming, northern Utah, and western Mostiie: Type locality: ‘‘Plains of 21. Artemisia cana Pursh. Hoary Sagebrush. Fig. 5713. Artemisia cana Pursh, FI. ign nary 521. 1814. Artemisia columbiensis Nutt ~at 142. 1818. Low, rounded, evergreen (at least in our area) shrub, 4-9 = rarely to 15) dm. high, often root- audio: freely branching, somewhat fastigiate, mill aromatic; twigs striate, canescent ; older a hes covered with a brown fibrous bark. Leaves of “vegetative shoots cm. long, 0.1-0.7 cm. wide, linear or linear - obla iiiiate, — amie oh ute or acuminate or sometimes with 1-2 ietdentas teeth or lobes, silvery-canescent, sometimes becoming wieciduboas with age; leaves of the inflorescence similar, reduced above ; in erences Bi Sa or i reduced, in the extreme aay aan volucre campanulate, 2.3-4.5 mm. wide, 3.5-5 mm. high, the outer ee or ovate as an acute or acuminate tip and eniely cancscent: or only sparsely ner phyllaries elliptic to ae nea ate with a broad scarious margin, canescent ee ne sar iG se 8 us; ray-florets absent ; disk-florets 8-20, rarely fewer, fertile; achenes cylindric- tur ee: granuliferous : receptacle glabrous. » = =. 3 Mostly on low ground, Upper Sonoran, Transitio: nadia southea n Oregon and eastern California south to Mono County east to Saskatchew am Tease, =i Caloskdo; ncaa foi ouster sites t gw n A. tridentata and also more tolerant of alkali. Type locality: bluffs of the Missouri River. Collected by Lewis. Artemisia cana subsp. dolbadent (A; crs y) G. Ward, Contr. Dudley Herb. 4: 192. 1953. (Artemis: neg A. — Proc. Am 19; 50. 1883; A, “ipudentat or. bolandcri Hall & Clem. D oveait eg ash. Publ. No. 326: 139. Ts23 — WideWiels var. bolar di ot Mags pis Calif. Shr . - . vn She ub 2-6 high, much branched, often root-spro vatiae neg 1— long, 0. 1-0 m. linear or bean: oblanceolate and entire or sometimes dilated at the ip oa divided into 3 “narrow = Mes apees. canescent; involuc A i sely canescent; disk-florets 8-16. This neces differs from A. cana suhep. cana mainly in its sm = ke e and usually somewhat more ee lea = 9. Limited t a the area around Mono Lake, Mono Coanty, California, Type locality: Mono . Me fe a alige County ail Sierra Nevada. 22. Artemisia rothréckii A. Gray. Timberline Sagebrush. Fig. 5714. Artemisia rothrockii A. Gray, Bot. Calif. 1: 618. Artemisia tridentata subsp. rothrockit Hall & Cle Cocueats tout, Wash. Publ. No. 326: 139. 1923. Artemisia tridentata var. rothrockii McMinn, Ill. "Man. Calif. Shrubs 608. 1939 Low, spreading, evergreen shrub, 2-6 dm. high, often sage? “sprouting, mildly aromatic, bark light gray or straw-colored to dark grayish brown, fibrous o older stems ; twigs striated, “re — .5-5 with a dense tomentum at least when young. Leaves of vereiative chooks cm. long, 5 cm. wide, broadly cuneate or flabelliform, 3-toothed ti ceolate or oblanceolate and entire, 1 escent, someti becoming glabrate or viscidulous with age; leaves of flowering shoots similar or all lanceolate and ntire; in ence s m to narrowly paniculiform ; involucre campanulate, m, wide, 4-5.5 high, the nites phyllaries ovate with a acute or acuminate rea and canescent or only sparsely tomentose, the in = phyllaries elliptic to nar- rowly obovate and usually sparsely tomentose labrous; ray-florets absent; disk-florets nee to glabro: oe wae: fertile: achenes cylindric-turbinate, granuliferous ; poche: glabrous. 2 | y sites or sometimes in moist pockets at lower Ts Boreal Zone; mostly at high elevations in the Bowed Nevada and the White and San Ferns eee Mountains of California. Type locality: Monache Meadows, Tulare County, Sierra Nevada. Aug.—Sep' 23. Artemisia rigida (Nutt.) A. Gray. Scabland Sagebrush. Fig. 5715. Ariomisis trifida or rigida Nutt. ap Amer. Phil. rao II. 7: 398. 1841 A. Gray, Proc. Amer. Acad. 19: 49. 18 Low, spreading, ie ah ub, 1-4 dm ot probably never root-sprouting, pungen ntly aro- matic ; ce thick and rigid, the older covered with dark gray, shredding bark, the twigs striated and covered with a dense tomentum at least when Leaves of ucts shoots silvery- — . lon i i ie: into SUNFLOWER FAMILY 415 the base; involucre Papas pt 4-5 mm. high, 2.5-3.5 mm. wide, canescent, the outer phyllaries Piso orbicular and acute, hes inner elliptic to spatulate and obtuse; ray- nite ets absent; disk- br cart 18. s 5-16, fertile; cencien somewhat prismatic, glabrous ; ee acle glabro sont eep rocky hillsides and rocky ab mer ic pper Sonoran Zon ral and duibishive Washin ton, att Boot. oe , and western Idaho. Type locality: ‘‘plains of pon gy tare ] River, in the Rocky a ta 24. Artemisia a A. Gray. San Diego Sagewort. Fig. 5716. onbeeuinte nsenitie: Gray, ae r. Acad. 11: 76. 1876. Imert Rydb. ae Fl. 34: 285. 1916. Shrubby at the base a long, apogee wand-like stems 10-30 dm. high, in clusters up to 10 dm. ac bane casemate the | woody stems covered with a grayish yellow bark, herbaceous ems green or yellowish below, reddis ve, piinnt striated, glabrous or minutely puberulent. Lea cm. long, pinna parted into 3-9 linear or linear-la eolate | , some entire, linear or linear-lanceolate, marg sely revolute, glabrous or min puberule bove, d ly tomentose bene h; inflo errens broadly paniculiform, the eads sessile or on peduncle long; involucre hemisphe c or campanulate, 2.5-4 mm. high, mm. aoe sparingly pubescent or glabrous, the inner phyllaries te na ahd than the o ; ray- -florets absen nt; disk-florets 12-30, nearly Mae smatic, franuliferous acle chaffy, each Hae subtended by an elliptic obtuse bract, or center florets without beatles me eo. soc and moist banks, Upper Sonoran Zone; southwestern San Diego County, California, and north- western Lower California. Type locality: Jamul Valley, San Diego County. July—Aug. 5711. Artemisia arbuscula 5713. Artemisia cana 5712. Artemisia tripartita 5714. Artemisia rothrockii 416 COMPOSITAE Tribe 5. ease eae se * mipatar ae least those of the disk) yellow or nged with purplish (some spp. of Luina merely yellowish) ; ps ts with all va vehaie al pra a disk. el fertile = lipeci divided styles; in- co ral bracts f 4 or more; pappus well developed at least in the disk-flow cee leaves dei (or sometimes some of the reduced uppermost ones ofieets: perennial herbs with the style of Senecio. 108. Arnica. Cauline leaves all or nearly all alternate (or the leaves all basal) ; styles and habit divers Pappus erideatyy plumose; style-branches with elongate, slender, hairy appendages; perennial herbs, discoid or wi —— radiate. 109. Raitllardeila. Pappus oa at ins ose; splebranches with deltoid or eer appendages, or a St alin be = penile, truncate o arly so Involu te) oa , and essentially exappendiculat £1 on ries of equal, t-connivent bracts, lh ing with some smaller, 2 Sed oe ones; plants ‘of pea habit, but not depressed and dichoto mously ranched. Heads Faametea = less often discoid, the pistillate flowers if present bara (aemab some- very shortly so); herbs or shrubs. Heads peta the 2 or more rows of marginal flowers stilt, nie Subteks filiform, peer corolla; herbs. 15. Erechtites Involucre of 2 distinct, subequal series of bracts, the outer rather ey ours Wearekont: sub- dichotom: ae bran chat: -— forming plants; heads discoid. 113. Psa athyrotes. Style-branches not ote rooataiag See in some spp. of Tetradymia), with or without well- ds merely except in Crocidium. Heads radiate; style. oo with a flat, deltoid, externally short- hairy ba itis SP diminu- tive Cro Heads isco: thos Bh teat nearly or quite exappendiculate, ae direkt? cabal and are the _— c lines extending nearly or quite to the blunt or acute tip; perennial oa ie sue with 1 narrow leaves (less than 5 mm. wide), these sometimes reduced to scales, or Involucral heats conspicuously imbricate in several series — among our genera of the Senecioneae in this respect). 118. Lepidospartum. Involucral bracts uniseriate or nearly so. ae lucral bracts only 4-6. 117. Tetradymia. ucral bracts more numerous, commonly about 16. Peucephyllum. Herbs (sometimes si endy chip OM with broader leaves, at least i” gee peeved ones r more Flowers white, sometimes +h AD inged with pink or eee = i ee or orange; plants with Bee disk. Rogice and undi vided & or nearly undivided styles, ps in Dimere Involural — and flow more than s not plumose; erect perennial herbs Pappus o merous capillary bist heads unlike, the plants shes ame cious. 112, Petasites Papens none; menee alike, th 106. Adenocaulon. 2 or sometimes 3; pappus Denedi ade annuals 107. Dimeresia. 106. er LON Hook. Bot. Misc. 1: 19. 1830. gto acle naked. Anthers se ie strongly sagittate but Pappus none. i n, , and kaulon, stem. _A genus of 4 s western tan America, one in eastern Asia, one in Guatemala, and one in Chile. Type species, gy pote tng bicelr Hook. = Adenocaulon — Hook. Trail-plant. Fig. 5717. — aulon bicolor Hook. ene: eS a 2S aulon paves Nui oo rans, Amer. Si Sc ate ia yas Adeaccoeles bicolor Bi coe Torr. & Gray, FI. s-rooted, slender perennial up to nearly is sle tall. Le pn mostly near the base, lon Borie reich and thin, deltoid-ovate to cordate or subreniform, m. wide, essen ntia ie glabr — — closely white-woolly lobed ; luc 1840. N. Amer. 2: 94. 1841. rnia, thence southward in "the Slerce Revids and in the ‘Const Ranges to Santa Cruz County; a i Juan de Fuca, and about Fort Vancouver on the C so in ear ae. Type locality: Straits of olumbia Rivet June-Sep 107. Nee seatalara A. Gray, Syn. Fl. N. Amer. ed. 2. 12: 448. 1886 scoid, 2-3-flowered, the flowers tubular and perfect. Involucre of 2 or times 3 herhaccas bracts, iin 3 at the base, each broadly rounded on the back and evely as contributed by Arthur John Cron ; In the Senecioneae, the term Mawphacrat sag is used instead of ‘‘phyllaries.” SUNFLOWER FAMILY 417 embracing a flov Anthers sagittate but scarcely caudate. Style-branches flattened, slightly broader shes ard, papillate-puberulent over the outer surface, the stigmatic lines ventromarginal, extending completely around the rounded apex. Achenes glabrous, stri- ate; pappus about 20 coarse, sparsely long-plumose bristles, united at the base and deciduous in a ring. Compact little annuals with entire leaves. [Name from the Greek dimeres, we two parts or members A f northwestern United States. r & ‘ Hea tron A. Gray. Doublet. Fig. 5718. Dimeresia howellit A. Gra . Fl. N. Amer. oad . 12: 449, 1886. Ereminula howe diaeGikene, ee 248. 1892 aely acaulescent, cushion-like, anweet annuals somewhat arachnoid at the base, more aonigreaed 1 up s. Leaves spatulate to elliptic or ovate, often punctate, up to 1 cm. wide and 3 cm. long tele en as petiolar pet aggregated around the | or sev ten compact Muee of sees ra in- volucre about 4-6 m igh ; shapes white to pinkish or purplis eee se or ae) an the foothills and at moderate ab eae in ae mountains, sometimes on serpen- d Transition Zone: oe Bones Baker County, Oregon, south to northwestern Nevada and Modoc and oe cation California. Type locality: Steen Mountains, Oregon. May —Aug 5715. Artemisia rigida 5717. eA a wy 5716. Artemisia palmeri 5718. Dimeresia ho 418 COMPOSITAE 108. ARNICA L. Sp. Pl. 884. 1753. Heads radiate or discoid, the rays when present pistillate and fertile, yellow or orange, relatively few and broad. Involucral bracts herba aceous, more or less evidently biseriate but ubeq nd connivent. Receptacle convex or nearly flat, naked. Disk- ers f or perennial herbs from a rhizome or caudex. Leaves sim nple, opposite, or the reduced upper-. most.ones occasionally alternate. Heads rather large, turbinate to per eit ae to rather numerous. [Name of uncertain duccaten perhaps a corruption of Pta A genus of about 30 species, of circumboreal distribution but most highly developed in western North sengy o ask species, Arnica ~~ ontana pecies—A. alpina, A. chamissonis, and A girder Aegati have been reported to s and the hess ont i of these, at rate may well prove to be argely or wholly apomictic. Some ane "the roan: in specific delimitation suggest the existence of a polyploid- Pekan Me complex, but this remains to be demonstrated. Heads characteristically radiate Seero ioc rayless forms of some of these species occur, chiefly in company with the normal radiate plants). Cauline —— relatively numerous and well developed, mostly 5-12 p nvolucral bracts obtuse or merely acutish, bearing | a tuft of dain hairs at or just within the tip; apn es elongate and oe naked, t 1. A. chamissonis. Involucral ig Vo more or tes shar pooh the tip not markedly more hairy than the y. ntire or ne s densely —— d (often in very large clones), the rhizome com- nly pi Aaa Tats a branching cau 2. A —— Leaves more or less toothed; tufted, the rhizomes mostly more elongat J. As Saiieeicondls. Cauline leaves few, mostly 2-4(5) pairs, not including those, if ids of the basal cluste Rays short, mostly 7-15 mm. long; lower petioles and lower part of the stem n densely and ae ipa pul ubescent with long, loosely spreading hairs; veer heads nodding; n Ca pei Rays a mostly 1.5-3 cm. long or occasionally a little shorter in some ousits ee he not have conspicuous long hairs; heads erect; widespread. Pues Aa 3 el gl umose, more or less tawny; rhizomes freely rooting, often shortened into Heads broad, mostly subhemispheric; cauline leaves variable in shape; ge ee Riccio the best a. . A, mollis. Heads narrow turbinate; cauline leaves relatively broad, mosty adc or oid to br sati er tanag a broadly lance- elliptic, the middle ones ign ta we ie the Sheree a olia, Pappus merely barbellate, or becoming weakly subplumose in some species =< Taste, nearly naked rhizomes, generally white or nearly so Leaf-blades relatively narrow, mostly (2. 5)3- 10 times as long as wide; basal leaves often TF rcdbeni —— but not always so; rhizomes short and freely rooting except sometimes in Heads lative large, hemispheric or: nearly so, with mostly 10-23 (about 13 or about 21 hag Mt lower aogag leaves generally petiolate; foothills and moderate wend tions in the mountai Old leaf sme with ve nse tufts of long brown wool in the axils; _ disk-corollas with some eading glandless hairs as well as commonly 7. A. fulgen Old wee age without axillary tufts or the hairs few and white; disk- eels stipi- wapecane generally not otherwise hairy. 8. sororia Heads ee ,» with mostly 7-10 (about 8) rays; inter cauline leaves ede to be sessile; - d 9; Aa page nee Leaf- — Bain! std broad, the lar ones mostly “8 2.5(3) times as long wide; basal S sometimes persistent rug pat Seneety tufted; PDE Mee in “forms of A. lati- anton elongate and subnaked, sometimes subapically branched and more scaly. Achenes generally glabrous below or glabrous throughout; basal leaves (those on separate short shoots) seldom cordate when present, the cauline ones even less Pireate uently so; leaves generally more or less toothed; sapeleers wit tn few 24 no long hairs. Pappus BS i Lay sere Feng 1 to several; plants w gs oce ane on not so Sidhew A pyre rine from June to Aue 10. A. latifolia. Pappus eure subplumose; ages strictly solitary; ~— res Bi” ~, Cag tnd in and near the Klamath region, flowering mostly in April a y (June). 1; A pl mi : Achenes mostly short-hairy (or glandular) a, or = to the base; leaves and in- volucre various but mos’ ag he not prese g the foregoing combina tion of characters. ppteeeal bracts obtuse or ey eee ae had cha a ge tuft of hairs within; s ( 3)5-7 : laces OG thin y tetutcas. ovate iy orate die ieee not cordate; sa ge of the Sierra Nevada and Klamat ie. a zasteit pe bracts m or less sharply acute, mostly without any very promin ~ tuft pg taire: heads commonly 1-3, or sometimes more in Senta oi ares leaves either glandular (rather than tomentulose) or cordate; iano speci: Involucre with geen or no long hairs; leaves entire or nearly so, the low br — rolendel to oe at the base; lateral setae of the pa) pus stles more prominent in either A. Jat atifolia or A. cordifolia; ponicinek in hy Sierra Ne mak: extending budmdan te less commonly, to | sagan . A. nevadensis. Involucre to usually copiously periited. p with Sk white hairs, es cally ¢ onc the base; — toothed t etimes entire, the basal and enerally also the lower cauline ones tiuncauasty more or less strongly cor- SUNFLOWER FAMILY 419 dat cely so in reduced alpine forms); common in Oregon and Washington. grant Bee? southward, less commonly, to the pata ts une! of the Sierra Nevada. 14. A. cordifolia sents Sherecteuiprsy” disedid gh oy corollas occasionally a ag abe truly radiate). ast the lowermost lea more or less petiolate, the petiole sometimes (4. ee sage — heads idiies: avilndeas bracts narrow, rants and slenderly acute Aye 'g d Was ington, i Sarkyi: “ Heads erect; involucral bracts broader, oe acutish or hb acute. Lowermost leaves with mostly ovate subcordate or re igs blade more or less abruptly con- tracted to the mass 8 or at: wing i petiole, rely spatulate and wing- teen achenes usually tm not always hairy as well o “stipitate: a rasiotor: a aren on to southern California, ‘nig ih the Klamath reg . A, discot lLowermost ili more or atulate, usually with Ngcatis winged, often poor ssi ‘petiole: achenes stipitate- ih g Rtg region only. A. spathulat Leaves all broadly sessile, numerous, rather small; local species of northern Ke neas and cau regon Leaves tuotbied: reticulate-veiny; achenes hairy, not glandular. 17. A. venos Leaves entire, merely 3—5-nerved; achenes stipitate-glandular. 18. A. viscosa 1. Arnica chamissénis subsp. chamissonis var. intérior Maguire. Meadow rnica. Fig. 5719 Arnica chamissonis subsp. chamissonis var. interior Maguire, Madrofio 6: 154. 1942. Perennial from long, nearly naked rhizomes; stems solitary, 2-10 dm. tall; herbage mor less villous puberent to illo: Nikite, saa ng glandular jeg scid above. Cauline leaves Sanity , not m reduc par. lanceolate to slacia. sessile or the lowermost usually cm. long, 1-4. wide; heads generally eo ra) g ~ | ro) eh o a ® cfr —e S. FX} co o oO re So: o 33 Z5 e< Sg — oO a ° a | wn 2&8 a5 oS e of rie involucre with ver y prominent c walls; rays commonly about 13, usually pale, about 1.5 cm. long; achenes ae hairy and Diahdalie to palaces pappus tawny, subplumose. ows and other wet places, Boreal and Transition Zones; Alaska and Ay 3 age = oo ol earl ngto ypical A. chamissonis, a maritime phase of coastal —_ a and ihe Aleutian “Islands, has larger and mostly fewer (typically 1-3) heads, with the lower leaves hardly i = rnica chamissonis subsp. folidsa (Nutt.) Maguire, se a nin 0. “4999. (Arnica foliosa Nutt. repe Amer. Phil. Soc. IT. 7: 407. 1841.) Le a rl aad entire — teiety denticulate; hairs at the base of the volucre less prominently ave pappus eous ~ Ds itish, merely barbellate, Br grocw ge Boor. iss otherwise as in var. inter of the typical eaideaies: uthern Mac Kenzie o Onta: uth t athern Cali- fornia and northern New Me xico. Type locality: Hg ithe alluvial flats o f the Celaada af the West. eartioe. pols near anid oe of the lake Timpanago s [Salt Lake].”’ This is a ag tract subspecies, composed of eral varieti Those occurring in our sane may be keyed as follows ee bracts very blunt ‘ relatively broad; herbage often but not always rather —T emi: n Bernardino Mountai r. bernardin uaa bracts a little narrower ie more poi ete conspicuously silvery- pee. = ee an mae of very wet places (typically tod Bead shallow ater). Herbage less densely hairy, scarcely silvery; moderately dry or usually merely moist habitats len. ‘sometimes n habitats as extreme as those characteristic of var. incana. r. folio. 5719. Arnica chamissonis 5720. Arnica longifolia 420 COMPOSITAE Arnica chamissonis subsp. foliosa var. ereanenn (Greene) Maguire, Amer. Midl. Nat. 37: 140. ome bernardina bag Pittonia 4: 170. a es var. bernardina Jepson, Man. Fl. Pl. Calif, ins, es — nar: Mountai ains, California. Shy ge ocality: Bear Valley. chamissonis << foliosa var. incana Gray) re Fl. Alaska & Yukon a A 1950. Pel4s-1 re a Greene Cc the s end of th i oe ag Beiutains of St thern Wa line gton, ex xtending eastward occasionally fate pets so Utah and south- western Idaho and apparently sometimes northward to southern Yukon. Type locality: Sierra Valley, California. Arnica chamissonis subsp. foliosa var. foliosa. Range of the subspecies except the mountains of southern aiicons. pet hosaadinggan ie m of A. chamissonis subsp. foliosa, which approaches As parryi var. i in its some- me Ror iavonticral bracts ape, in its more or less reduced cauline leaves, has been = pene a he taaii of a few collections Lhe the Sierra Nevada as var. jepsoniana Marcie, Aare. Midl. Nat 37: 140. 1947. The real biological nature of these s sche iid remains to be determined. 2. Arnica longifolia subsp. aon ead Maguire. Seep- as ee Fig. 5 Piper, Proc. Biol. Soc 06. 1920. Arnica louse ibe weirichei anges eda 4: i 1943. Plants densely tufted, often in very large clones, with many smaller, re leafy stems in addition to the floriferous one s, these commonly 3-6 dm. tall, leafy, the rhiz commonly short- ned into a branching caudex. These mostly 5-7 pairs, one or more of the ss pairs connate- sheathing, with reduced blade, the others sessile or shortly connate-petiolate, not much reduced upward, narrowly lanceolate or lance-elliptic, pal tapering to the acute or acuminate tip, or tly 5 a 2 ide; no we entire or sometimes slightly toothed, mos 4 -12 cm. long and 1-2 cm. wid well-developed basal leaves produced, either on stem or on separa sie short shoots ; herbage more or less scabrid- puberulent, at least upward, and usually a “ttle viscid ; heads setaral to rather numerous, cam- panulate, sometimes rather narrowly so; involucre mostly 7-10 mm. high, the bracts sharply acute or acuminate, glandular-puberulent and generally with a few intermingled, longer, conspicuously septate hairs, especially below ; rays mostly 8-13, 1-2 cm. long; achenes subglabrous or glandular and hairy; ore or less tawny, barely subplu umose Il-drained soil (or rocks : —_ oes Peg: pectag? are salons oist cliffs and river banks, at modera o hig 1 ether and oe ‘Alber ay gm peaitee: Mount Rainer National Park, Washington. July— ee 58 seit amplexicaulis Nutt. Streambank Arnica. Fig. 5721. Arni. bl. lis Nutt. Trans. Amer. — Soc. IT. 7: 408. 1841. ica macounii Greene, hing 160. 190 Arniea abe oakte Sal Rydb. Mem. N.Y. Bot. re 1: 434. 1900. eene, Ottawa Nat. 15 or 1902 ria ciliate Rydb. N. ene Fl. 34: 351. 1927. Arnica macounii var. aspera G. N. Jones, Oa Wash. Publ. Biol. 5: 256. 1936. Perennial from sittick coarse, freely ing rhizomes, mostly 3-8 dm. tall, more or less glandu- lar and hairy, especially upward, or s ate ial beds leafy. Le eaves all caine mostly 5-12 pairs, the lowermost often reduc oe e other: ers elf developed and generally no h redu rd, nar- rowly lance-elliptic to ecg € or ovate, sania or the lowe seth hota: etiolate, more or less toothed, only cm. long and 1-3.5(4) cm. — heads generally several, campanulate ; involucre mostly 9-15 mm. high, its bracts sharply acute or acuminate; rays 8-14, pale yellow, 1-2 ~ long; achenes spaniels hirsuite and sometimes Yeadisla ar; pappus tawny, subplumose. am banks moist woods, Canadian and Transition Zones; Aiaake to western Montana, northeastern sl — to the southern Sierra Nevada. Type in locality: “On the nie oof the Wahlamet, at the * me i keke var. piperi St. er & Warren in St. John, Proc. Biol. Soc. Ph a 36. 1931. (A, Rrtica Amer. Fl. 34: 349. 1927.) Leaves very broad, up to 6 or reput edly 8 wide; in- volucre a genes hairy cad less glandular; otherwise much as in typical A. Seniesa but ‘ess variable. Columbia River Gorge in Oregon and Washi ington i Type "Pocatiog Cape Horn, Washingt 4. Arnica méllis Hook. Cordilleran Arnica. Fig. 5722. Arece mollis Hook. Fl. Bor. Amer. 1: 331, 1834. : e, Pittonia 4: 36. 1899, rnica rt levis Claene. ~ cit. 163. 1900. — calle teak Greene, op. cit. 165. Arnica mollis var. keahicvina Smiley, Univ. Calif. Pub, Bot. 9: 386. 1921, rennial from freely rooting rhizomes, which ma ortened into a loosely branched caudex, 2-6 dm. tall, variously puberulent to long-hairy Bp tag Cauline Lene poe ee, a lower commonly the largest, all sessile, or the lower often s petiolate, variously elliptic, or lanceolate to oblanceolate or bovate, irregularly denticulate (or nl a "Vita loped, petiolate, basal leaves often enh i Sg © few or se FS gorge fg ca ulate or broader, me rs times as s e; involuc ‘Doe places at aidan to hie cage in oe mountains, Canadian and Hud ian Zones; Alberta and British Combi to southern Colorado, southern Utah, and the sout ae faa of the: Sie erra Nevada in Cali- e higher mountains of. Washington and Oregon but absent from the California Coast Ranges, SUNFLOWER FAMILY 421 and from the Oregon Coast Ranges south of Clatsop County. Type locality: ‘“‘Alpine rivulets of the Rocky Mountains Teaser, ”* June—Sept. 5. Arnica diversifélia Greene. Lawless Arnica. Fig. 5723. Arnica latifolia var. viscidula A. Gray, Syn. Fl. N. Amer. 1°: 381. 1884. Arnica diversifolia Greene, Pittonia 4: 171. Perennial from freely rooting rhizomes; stems solitary on in loose tufts, 1.5-4 dm. tall, disaular -onberulent to subglabrous. Cauline leaves mostly 3-4 pairs, sessile or at least ong lower generally shortly Ne cing ie ovate or lies to elliptic or ila t: lat irregularly toothed, the i commo e largest, wit s cm. long an se developed, Nappa basal leaves seh produced ; be ads generally graeereg narrow w, more or less turbi- the disk-flowers much fewer than in characteristic 4. mollis ; involucre 10-14 ai high, its bracts wien “A acuminate, shortly stipitate-glandular inroughont-4 sometimes sparsely long- pest oo rays commonly about 8 or about 13, 1.5-2 cm. long; athe stramineous to tawny, um i jen at moderate to high elevations in the mountains, Canadian and Hudso onian Zones; Alaska to Mon tans “Utah, and the northern meres Nevada in California; not common, type locality. Eagle Creek (Wal- lowa] Nisan tains, Cream . July—Sep venient nam a complex series of ga hybrids and hybrid progeny, invobeing ai vei or a. pst Ho on ‘the one chan d, and A. cor Anabey lia or A. latifolia on the other. The pollen is irregular and often very scanty, and apomixis has been report 5721. Arnica amplexicaulis 5722. Arnica mollis 422 COMPOSITAE 6. Arnica parryi A. Gray. Nodding Arnica. Fig. 5724. Arnt tifoli ge eradiata A. Gray, Proc. —~ Paso 1: 68. 1863. Arnica parryi A. Gra nC. C. Parry: rene sp: 1874. picove att Heller, Cn N. Amer. Pl. 7. ae Perennial ak freely rooting to ay naked rhizomes; stems mostly solitary, 2-6 dm. tall, often somewhat woolly-villous toward the base, becoming glandu lar at least above. Cauline ek mostly 2-4 pa eachake reduc art upward, the low ermost petiolate, with lanceolate or lance-ovate blade 5-20 cm. long and 1.5-6 cm. wide; well-dev coped basal leaves similar to the lower cauline ones often pro acti 3 on separate Hort shoots ne: nerally several, nodding in bud, later more or less erect, campanulate, sometimes narro 0, 5 Oftinatily discoid, the marginal corollas some- times ampliate or nee ee badites hivalecre mostly 10-14 m m. hi gh, its rather narrow bracts sharply acute or acu achenes glabrous to sanontar or wie pappus tawny, strongly bar- bellate to weakly subohia w , drier me and moist slopes, in the foothills and at moderate elevations in the mountains, y Canadia ng Pee otk ertt Ribera ings Py Lee: Columbia to scone sg eho Wane southern Oregon, rarely poss ie hes into northern tie comm the Cascade and Olymp c¢ Mounta of Wa shi iaaien but seldom collected i in Orego: ocality: Clear acc Colorado. July—Aug Arn var. sOnnei ireeee) Cronquist in Ferris, Poe, Dudley Herb. 5: 102. 1958. (A. sonnet Greene, Pittonia 3: 104. 1896; vee | rete var. esa Je ete — n. Fl. Pl. Calif. 1157. 1925; A. parryt subsp. sonnei Maguire, boat 4: 482. 3.) Lower part of the and lower petioles more consistently and more densely and conspicuously woolly-v file ous han an the typical subspecies; beats cndinasehy pr: cana the rays Os mostly 7-15 mm. ee. Sierra Niveas of ee et and aia t Nevada, exte ng © rthward, rarely, the Cascade Mountains of southern Oregon. e locality: near Tru ckee Rives ents 7. Arnica fulgens Pursh. Hillside Arnica. Fig. 5725. Arnica fulgens Pursh, Fl. Amer. Sept. _ haere Arn sca montane ET mrese Nutt. Gen. 2: rn Rydb Bull. Torrey Nes ‘a: ee 1897, Per ennial from short, freely rooting, densely scaly rhizomes, the basal leaves and persistent old leaf. bases with oo axillary tufts of long brown wool ; stem stout, 2-6 dm. tall, stipitate- glandular and alten also hairy, more densely so upward. Leaves 3-5-nerved entire “ nearly sO, a to) u ro ie 1m. isk-corollas with some spreading white hairs as well as aaecly: Siendaiad “achenes densely hairy; pappus whitish or stramineous, barbs llate. shia chiefly in the foothills and at moderate elevations in the mountains, Arid qipeaision and Cana- dian Zonas: analog fy British Columbia to Ceci tchewan south to Co lorade, northern Nevada, northeastern California. Type locality: ‘‘On the banks of the Missouri.’”? May—Jul 8. Arnica sordéria Greene. Twin Arnica. Fig. 5726. Arnica sororia Greene, Ottawa Nat. 23: 213. 1910, Similar to te a Jwinens _ averaging more ses a ane less scaly, more slender, and en more elo Leaves averaging narrower and ‘few ool nearly or fi absent from the axils of a Toeat oe white when prese nea Sinvolucra pay narrower ess hairy, gen- erally broadest at or near the base, tapering then Bo more acute tip; isk: corollas stipitate- glandular, not otherwise hates pappus white or ee Open, often rather dry places in the foothills and at ee elevations in the mountains, Arid Transition Zone; southern ng peg nag and Alberta to Wyoming, northern Utah, and northeast epoca t annes nearly to Donner Pas e Sierra Nevada). Type locality: between the Kettle and Columbia Rivers, Cascade British Columbia. tay «5 Bug 9. Arnica rydbérgii Greene. Subalpine Arnica. Fig. 5727. Arnica rydbergii Greene, Pittonia 4; 36. 1899. Arnica aurantiaca Greene, Torreya 1: 42. 1901. Arnica sulcata Rydb, N. ‘aie . Fl. 34: 344. 1927. late or spatulate, 3-5-nerv: 7 1 a de frequently pie d on e short shoots; cauline leaves mostl pairs, sessile, or the lower shortly and broadly wing-petiolate, oblanceolate or spatulate to la late o metimes a little broader, entire or nearly so, 3-5-nerved, mostly aggregated on the lower part of the stem, 3-10 cm. long a wide, or the r smaller and distant; heads ry inate-campanulate ; involucre t ciliate or scaberulous on the margins, moderately acute; rays mostly about 8, 1-2 cm. long, t ip minutely toothed or entire; achenes densely ot ice sed eaten the upper hairs the es Ae barbellate. open slopes, mostly at high elevations in the mountains, Hudsonian Zone; British Co- lumbia and Thee ‘e Ps vides northern Utah, eastern — and (rarely) northern California (Scott Moun- tains). c= locality: Little Belt Pass, Montana. July—A SUNFLOWER FAMILY 423 10. Arnica latifélia Bong. Mountain Arnica. Fig. 5728. Arnica latifolia Bong. Mém. Acad. St, Pétersb. VI. 2: 147. 1832. 1 enziesit Hook. Fl. Bor. Amer. 1: 331. 1834. ry ; R Arnica glabrata Rydb. N. Amer. Fl. 34: 335. 1927 7 ee hb. Ryd 1 1 scaly at the apex; stems mostly solitary or few together, 1-6 dm. tall. Broad (occasionally cor- 424 COMPOSITAE subhemispheric ; involucre mostly 10-18 mm. high, more or less glandular and sometimes with a te Bear at achenes generally Feu: 2 least toward the base, often glandular and/or shortly y above; pappus white, barbel the oist woods, meadows, and sg open the seldom at very high elevations, mostly pac een ae Hudson nian Zones; Alaska Be opoting to Seveariigg Utah, and northern California (southward occasionally to Eldorado County). Type locality: island of Sitka. Jur e-Aug. a. ietifolia var. gracilis (Rydb.) Cronquist, Vasc. Pl. Pacif. Northw. 5:51. 1955. (Arnica gracilis Rydb. Bull. Ltd oo 24: 297. 1897; A. betonicacfolia Greene, eran ha 163" 1900; A. betonicacfoha var. gracilis M. E. Bull. Univ. Mont. Biol. Ser. 15: 48. 1910.) —— yee plants (ie 3d tall) with sev- eral slender dg a ising from the een which is commonly shorten d into oosely arias wi scaly caudex. Leaves seldom over 2.5 cm. wide; heads mos sph 3—9 (occasionally soutaee: mateo small, the involucre ts m. high. Rocky places at moderate or usually high elevations in the mountains; Alberta and British Colum ee asrogton. ig sa resend central Idaho, northern Utah, and Colorado. Type locality: Spanish Sieg a 11. Arnica cérnua Howell. Serpentine Arnica. Fig. 5729. nica cernua Howell, Fl. N.W. Aimer. ie 1900. ie chandleri Rydb. N. Amer. Fl. 34: 339. 1927. pete) from elongate, nearly naked rhizomes; stems solitary or few together, 1-3 dm. tall. Broad e or less ovate-cordate), long-petiolate, basal leaves sometimes produced on separ oe short eee cauline leaves mostly pairs, glabrous or scabrous-puberulent, mostly petiolate and lance-ovate to obovate or even subcordate, the lower ecsanilty the larger, up to about 5 cm. n c sc argined ; achenes evidently hir bite above, glabrous below; p appus pie. with 7s ay eral etna of the bristles better developed than in A. latifolia and os cordi ifolta, less developed n A. mollis and A. amplexicaulis. Serpentine slopes, Arid Transition Zone; pares County, Sinan! 4 wee in pe kiyou, Humboldt, and Del Norte Counties, California. Type locality: near Waldo, Josephine County, Oregon. Apa May (June). = Arnica tomentélla Greene. Recondite Arnica. Fig. 5730. Arnica t t G , Pittonia 4: 155. 1900. rennial ws well- igi d, long, nearly naked rhizomes ; stems solitary, 2-5 d all, elandular and hai eaves apparently Be caine: mostly 3-4 pairs, thinly Cnehii. Riansene the ogee developed ones petiolate, with ovate to ovate- cae toothed or sometimes cell blade about 2.5-7 cm. long; heads (3)5-7, cam panate-h emispheric ; involucre mostly 10-13 m m. high, ue bracis scoais or obtuse, fen. papers ent and ciliat e, and bearing a tuft of longer hairs rnally near the e tigte ays 12-20 mm. long; pene s short- bed throughout, and very sparsely stipitate-eandular pappus white to stramineous, Sunk ly ba bel llate or shortly subplumose. Open slopes or open doing bel mberline in the mountains, Canadian Zone; Josephine County, Oregon, oe adjace oe gece Coun at feria: reappearing in ~ Sierra Mevada Sons the! Srrickes River Tulare County, California. ge gy tl Middle Tule River, California, folk ——— referred to A. tomentella seem to combine the ieatares of A. cordifolia and A. chamissonis subsp. ones 13. Arnica nevadénsis A. Gray. Sierra Arnica. Fig. 5731. 4rni. densis A. Gray, Proc. Amer. Acad. 19: 55. 1883. Perennial — long, nearly naked rhizomes, these often apically branched and more scaly ; r few loosely clustered together, 1-3 dm. tall; herbage more or less strongly r e ober 3-8 cm. lon wide. ope or occasionally denticulate; long-petiolate basal oe, similar to igi oe the kag part of the stem, often produced on separate short shoots ; heads 1-3, broadly turbinate to c mpanu ulate ; etabon mostly 10-18 mm. high, the bracts more or less sharply acute, densely short stipitate-glandular, and sometimes with some longer hairs below ani these less conspicuous than in A. cordifolia: rays 6 about 12- reap r 25 mm. long; achenes mostly un form es glandular or chore or both; pus white stramineous, strongly barbellate a shortly subplumose, the lateral setae of the ‘bristles better developed than in A. cordifolia and A. Tattolte ang: developed than in 4. anoilis and A. ample-ricaulis, Open rocky slopes at high a Bits in the mountains, Canadian and ads sonian Zones; petonss in the Sierra iene of California and adjacent Nevada, northward les: — nly to the Cascade og Olympic Moun- tains of Washington, Type bog ity: Lassen Peak, Californ Tal In the aforestated area A. nevadensis appears to be al characterized and readily ls from et mest relative A. cordifolia. In the Rocky Mountains and Great Basin, how ye — alp see extreme . _ rs folia var. pumila ( Rydb.) Maguire are barely if at all to be distin valid; m Sie ke nevadensis, ‘fro om — se nbd chiefly in distribution and in the slightly shorter barbels of ee Fete bilities. these pla t the closest approach of A. cordifolia to A. nevadensis, have been treated as A. cordifoha ne dost Pe (Rydh, Maguire 14. Arnica cordifélia asta Heart-leaved Arnica. Fig. 5732. Arnica cordifolia Hook. Fl. Bor. Amer. 1: 331. mer. Phil, te < 7: 408. 1841. Arnica austiniae Rydb. N, Nader. Fl. 34: 340. 1927. Arnica cordifolia var. macrophylla Maguire, Amer. Midl. Nat. 37: 137. 1947, Perennial from long, nearly naked rhizomes, these often apically branched and more scaly; stems solitary: or a loosely aad ‘egethox bioety 2-6 dm. tall, glandular-puberulent to more ES a S 3 a 35 = + 2& oy Z S pas lar a p 3 D SUNFLOWER FAMILY 425 often loosely white-hairy. Long-petiolate, cordate, basal leaves commonly produced on separate short shoots; cauline leaves mostly 2-4 pairs, the lower obviously the larger, iis Bre, with ply 4+ and 3-9 cm. wide, more or less te blade ( ee ees dee sh: commonly 4-12 cm. lon entire or more often to othe s l- roots At up to 7 broadly tu paar! to lanceuhinlate: involucre Torey 13-20 mm Sigh sparse ty 6 co pina agin with fone: i Rap rie rahe especially toward the base and generally glandular as well; s 1.5-3 cm. long, mo 0-15, o rarely wanting in individual p eee achenes mostly andtorialy: shore bigiry | or a he fe both) : pappus white or whitish, barbellat Mostly in woodlands, from the Fa mi to moderate or sometimes rather high elevations in the mountains (where . pumece into the var. pumila), Arid Transition Zone to Canadian Zone; Alaska and Yukon to geome Dakota rthern New Mexico, ce To Tuan ge ane California to the southern end of the Sierra Nevada re west of th th 33 ascade summits in Ore ashington; also in northern Michigan. Type aiily “Alpine woods of the Rocky Mountains, on the os fey aie ” pre —June (Sept.). tmnica cordifolia var. pumila (Rydb.) Maguire, Madrofio 6: aor 1942. (A. tig oe Mem. Bot. Gard. 1: 433. 1900; A. humilis Rydb. N. Amer. Fl. 34: 341. 1927; A. cordifolia r. humilis Magui Amer. Mid]. Nat, 37: 138. A dwarf ecotype mostly less than 2 pik ‘tall with amet more often utite, only slightly or ie cordate leaves commo nly 2—5 cm. long, with more glandular achenes, and often with less panty re ee. sagt aloe re. ee and aakatolak eerone nearly or quite throughout the range of the oo. om in a where is mostly replaced by A. nevadensis (quod vide). Type locality: Gray’s ea olor 5729. Arnica cern 5731. Arnica nevadensis 5730. Arnica ‘encase 5732. Arnica cordifolia 426 COMPOSITAE 15. Arnica discoidea Benth. Rayless Arnica. Fig. 5733. Arnica discoidea Benth. Pl. Hartw. 319. 1849. Perennial from long, nearly naked ioe sg stems mostly solitary, 3-6 dm. tall, glandular- puberalent and often also more or less long-hairy: Leaves sparsely to rather copiously long-hairy n sides, and often also glandular, rather esate toothe * A Mi subentire, the basa ones “cicero te, with narrowly ovate to deltoid or even subcordate blade up to about 8 and 3. wid i cm. long cm. e, kate oadly wing-petiolate ps with ill- iat blade almost as in Al. Se ath ulata ; oaltiae leaves ral pairs, evidently and -prpercalyely reduced, mostly gin ral with broad w met ee ot pi oy ill- in ed blade; only branched above from near the nie producing several or rather many font res ane: 30) h pees the wer sder up leaves often alternate? heads turbinate- Sepanulee to subhemispheric, ‘Alsen (the heraenl corollas earels ermal) involucre 9-13 mm, high, glandular and SDFEAA ELUNE, the bracts obtuse or somewhat acute, carrying their width well above the middle or nearly to the tip; achenes glandular see natty throughout, or glabrate below; pappus rather strongly barbellate, white or stramineous. Open woodlands, Upper Sonoran_and d Transition Zones the Outer Coast Ranges of California sig oie ae ocino County to eo by he uis Onin “sone (the Pp eeptiroreny pi of Jepson). Type locality: woods near On arin California. June— - nica — oe eradiata (A. Gray) = ge Vas 1. Pacif. Northw. 5:49. 1955. a parvi- flora Gray, Proc. Amer. Ac ai 7 363. 1868; Aha as. er hadi A, Gray, Syn. Fl. N. Amer. 1%: 381. 1884; my orasi Heller, Mublenbergia 1: a, 1900; A, falconaria Greene, Ottawa Nat. 23: 215. 1910; - tustckti Rydb. N. Am 927, in part.) Similar to typical A. discoidea but the middle and lower cauline i i i to ovate rarely through the Cascade Mountains to the east end of the — River Gorge in Oregon and Washington, where again common. Type locality: Hood River, Oregon. ay— — discoidea var. eave Ph wags Cronquist in Ferris, el ee Herb. pi 1958. (A. alata Rydb. N pore Fl. eras ; A. sanhedrensis Rydb. loc. cit.; A. parvt afore ubeb, alata Maguire, apd oa Maes 4: 455, 1 ey Similar to nae eradiata but ph the basal leaves genera whe dist mon cordate and th : e vhich | e sha fro inary fo A. cordifolia in being rayless. Moderate and lower ovations } in the Sierra Nevadi, carientrd lead commonly to the Klamath region, Bind common in the Coast ee just ainsi of San Francisco wey and oceasionally load northward in the Inner Coast Ranges of California. Type 1 ity: Yosemite, California. June-Aug 16. Arnica spathulata Greene. Klamath Arnica. Fig. 5734. Arnica spathulata Greene, — 3: 103. 1896. Arnica cusickii Rydb. N. Amer. Fl. 34: 343, in part. 1927. Perennial from br: aed nearly naked rhizomes, the stems ——~ solitary, mostly 1.5-4 dm. tall; herbage glandular and spreading-hairy. Leaves coarsely toothed or occasionally sub- entire, the basa yes more or less spatulate, with a broad, winged, ecole base and poorly defined blade, the blade and petiole together up to t 10 cm. long and 3.5 cm. wide; cauline leaves pairs, sessile and d-bas r the lowermost similar to the basal ones; stems com- monly branched above and producing several (up to about 20) heads, these turbinate-campanulate, discoid ; involucre ears mm. high, glandular and hairy-spreading like the herbage, more coarsely and conspicuously so than in A. discoidea, the bracts merely acutish, carrying their width well beyond the iiaie: vachenes stpitate sn ne ta throughout, generally not otherwise hairy; pappus barbellate, white or nearly Open woods, aida sd wall up in the mountains, mostly Arid Transition Zone; in the Klamath region of soutumesert Oregon and adjacent Californ nia. Type locality: Glendale, Oregon. rend July. Arnica spathulata subsp. ctdek tes (Rydb.) Lr hadi Brittonia 4: 458. 1943, (Arnica eastwoodiae Rydb. N. Amer. Fl, 34: 343. 1927.) More slender and less ently hairy than the typical form of ~ —— and more inclined to branch (sometimes even from near the bacek afd commonly with the stem or leaves or both more or less suffused with red (anthocyanin): petioles sometimes less winged. Foothill and seacoast ht es in Curry and Josephine Counties, Oregon, and Del eg eam vt semen < ward econ ak ag common aly 17. Arnica venésa H. M. Hall. Veiny Arnica. Fig. 5735. Arnica venosa H. M. Hall, Univ. Calif. Pub. Bot. 6: 174. 1915. Habitally similar to 4. apKee. usually ke branched, — less ys vas camgyat pubescent and less glandular. Leaves larger (up Abi and 3c mer, evidently reticulate- veiny (at feat apres as well as fen sang sharply and ‘saclay ape toothe d; heads a little larger, terminating ranches; disk pin! 9 y cae y long-hairy below, more sparsely so above, not glandular ; achenes chortle ae mo glandu ress Hot dry slopes in the foothills, Arid Transition Zone; Shasta County (Salt Creek and Iron Mountain), Cali- fornia. Type locality: Salt Creek, Shatte County. May-June. . r 18. Arnica viscésa A. Gray. Shasta Arnica. Fig. 5736. Arnica viscosa A. Gray, Proc. Amer. Acad. 13: 374. 1878. eer sere penecaiaee Greene Erythea 8: 48. 1895. Jepson, Man. FI. Pl. Calif. 1037. 1925. Picea! with numerous cee branched stems arising from a branching caudex, 2-5 d tall; herbage copiously pubescent with hairs of varying length, many or most of which are aad SUNFLOWER FAMILY 427 Pane, ays Nt f Fs at (4h Nt | 5733. Arnica discoidea 5735. Arnica venosa 5734. Arnica spathulata 5736. Arnica viscosa tipped, or the shorter hairs merely viscid. Leaves numerous and equably distributed along the stems, the lowermost ones reduced and bract-like, the others nearly all alike, sessile, elliptic or oblong-ovate to oblong-obovate, wel 2-5.5 cm. long an cm. wide, entire, evidently o a weakly Pes ees not obviously veiny; no separate Gadat leaves produced ; heads ree tip rather nume short-pedunculate, discoid ; involucre 8-11 mm. high, glandular-hairy like the Herbage + corottne i aea acted sw 5 -stipitate- seeagrn’ below, glabrous above ; achenes uniformly long- stipitate-glandular ; pappus white or creamy, str ngly barbellate. Rocky places eg ee Hudsonian ti Mok Shasta, California, and Mount Mazama (Crater Lake), Oregon. Type locality: Mount Shasta. Aug 109. RAILLARDELLA Benth. & Hook. Gen. Pl. 442. 1873. Heads — or sometimes with more or less well-dev eloped, fertile, yellow rays. In- volucral a ut equal, subherbaceous, uniseriate or nearly so. Receptacle naked, flat r truncate or nearly so at the base. Style-branches flattened, exter- nally einiawlobe. with introrsely marginal Seats lines and an elongate, slender, exter- nally and internally hispidulous appendage. Pappus of more or less numerous, cc haits plumose bristles that are somewhat flattened toward the base, or of narrow and elongate, bristle like plumose scales. Achenes linear, somewhat megressed. rian ae ater Pere 428 COMPOSITAE nials with simple, entire or sparingly toothed, aa ie or basal leaves (or the reduced lowermost leaves sometimes opposite), and solitary or few, yellow or orange-colored heads. | Named for its similarity to Raillardia, of the Hawsiian Islands. | A genus of 5 species, native to the mountains of California (and adjacent Nevada) and Oregon. Type species, "Reillerdia argentea A, Gray. Stems iamgaa leafy = ss cath — hed; 1l-developed basal leaves present Le sa or obtus e lowermost ones apposite, reduced and somewhat scale-like, more or less te-sheathin Sat. “. poo ; heads ordinarily with a few short rays; Inner North — perms 52 of Ca Leaves Saar aay all alternate, none sheathing; heads strictly discoid; southern Sierra Nevada 2 1rit, Stems mets nd monocephalous, scapose or subscapose; leaves chiefly or more often entirely basal. Leaves green, either glabrous or glandular, not at all tomentose. "He ads bout 20 or more (probably com pesly peng 21) involucral bracts and 6-10 well- Sot hy ka rays; leaves strictly eons’ tea! vicinity of Mount Eddy, California 3. R. pri a Heads mostly with about about 13 involucral maticg discoid or soandtines with 1-3(5)_ irregu- onal Setnaes rk leaves pie to densely and sometimes in smoneaieisile hai well; Cascade thern Te of central Oregon south through the Sierra Rae and on io e pia, apos Leaves gray, silky-tomentose; Cascade Mountain tral Oregon south through the Sierra Nevada. 5. ee argentea. 1. Raillardella scabrida Eastw. Leafy Raillardella. Fig. 5737. Raillardella scabrida Eastw. Bull. Torrey Club 32: 216. 1905. Raillardiopsis scabrida Rydb. N. Amer. FI. 34: 320. 1927. Stems several from a stout woody root and branching caudex, often curved at the base, 1-5 d tall, chaanealy branched and bearing several heads ; herbage e hirsute and stipitate- glandular, cists hirsute below, more glandular above. Leaves rather numerous and well distributed along the stems, principally alternate, small, och or nearly so, more or less obtuse, mostly 1-3 cm. long and 1. mm. wide, the lowermost ones reduced and more or less ae opposite, connate-sheathing ; heads naked-aeramculats at the fa of the branches, discoid o e commonly at least some of them with 2 or 3 short broad rays up to about 6 mm. long; cavaraere o 12m — coarsely long- ent eid the broad bracts mostly about 8 or 10; rays epappose, their achenes glabrous or nis hairy ; disk-flowers wit ‘a a papus of about 15 or more elongate, plumose awn-scales, shorter he He ubescent linear achen ye en, stony places, Canadian Sain in the Inner Coast Ranges of Lake, Mendocino, and ie ao mg Counties, Californie. glovatioe up to 7,500 feet. Type locality: Snow Mountains, Lake County. July— ae " Raltigrets muirii A. Gray. Muir’s Raillardella. Fig. 5738. Raillardella muirii A. Gray, Bot. cag 1: 618. 1876. Raillardiopsis muirti Rydb. N. Amer. Fl. 34: 320. 1927. Habitally similar to R. s se ida, Leaves acute, apparently all alternate, none sheathing ; pond age less hairy and less glan dar heads age the involucre with about 13 bracts ; pappus- bers _ = t 10, longer than achen d local in open sak a et aa agg oie Canadian and Hudsonian Zones; in the southern Rar Sierra Tvada; ely Type locality: in the Sierra Neva da, supposedly near Yosemite, perhaps actually farther south. July—Aug. ee ape ptinglei Greene. Showy Raillardella. Fig. 5739. Greene, Torrey Club Mt 17. 1882, Raillardella ‘scaposn var. aaa ieee Man. FI, Pl. Calif. 1146. 1925. Scapose perennial with a stout kc rhizome or caudex. Leaves oblanceolate or linear- ho 310 cm. long, 6-8 mm. wide, wholly glabrous, firm, some of them remotely and shallowly ars e : — $ n R. scaposa ; involucr high, shorter than the disk bracts relatively numerous, rep y 20-30, coarsely stipitate-glandular and more or less long-ciliate as ll; rays well developed, about m. long, d rifid, salmon or brick-colored; pappu Stream banks and w Hudsonian "par in the vicinity of Mount Eddy, northern California. Type locality : mountains west of Moone ir thang Aug.-Sep 4, Raillardella scapésa A. Gray. Green-leaved Raillardella. Fig. 5740. Raillardia scaposa A. Gray, Proc. Amer. Acad. 6: 551. 1865 Raillardella scaposa * Gray, Bot. Calif. 1: 417. 1876. on saree ite se central Oregon (the Sine Sieteres sallel Crreeee the Sier ore evada and on San Gorgonio Peak in the San Bernardino Mountains. Type locality: Sonora Pass, California. July—Sept. 110. SENECIO L. Sp. Pl. 866. 1753. Heads radiate or sometimes discoid, the rays — is fertile, yellow to orange o occasionally reddish, purple, or white or w Inv ral bracts herbaceous or i r ially equal, e r entire to minutely sagittate. Style-branches more or less flattened, truncate, penicillate, with introrsely marginal stigmatic lines extending to the tip or ate short- appendiculate. Achenes subterete, 5-10-nerved ; pappus of numerous, usually white, entire or barbellulate, or divided leaves and solitary to numerous, mostly small to medium-sized, cylindric to cam- panulate or hemispheric heads. [Name from the Latin senex, an old man, probably ferring to the white pappus or hoary pubescence of some species. | One of the largest genera of plants, containing probably well over 1,000 species, of very wide geographic distribution, ro 100 in North America north of Mexic - Bit ype species, Senecio vulgaris L. Man the species are a related, often, indee ith only about the ‘aap ree of distinctness more mmo aye hee ih! ted with varieties. This is epectatly true o the species prse treated under the groups Aurei, Lobake and Tomentost, which collective form a mike complex bec few no clear-cut specific lines. The ei. Pa ps indicated in the key in large part natural and correspon some degree to the ns of the species-groups 0 “ Rydberg: “or purposes of this erentieast i. are, however, considered to be groups of convenience without taxonomic s Senecio élegans 3 ay 869. 1673, % ee species with lyrate or - geeeacaias leaves and purple rays, is adventive in sandy aut. ut San Francisco. It is native to South Afric Stems erect or ee never —— or climbing; leaves various but scarcely ivy-like; heads various. Plants perennial (or in S. jacobaea sometimes merely bienn Leavy -. —, distributed along the stems, only Seis or ‘ol reduced upwards; no tuft of basal Leaves to skis times Bdsrinsmr ps - Scena gment: id a than linear; espa lived, all woody, VIII. JacosBagEa Touvedl venir. to toothed or pinnatifd « or in the woody insular species, S. /yonti, often ‘isleianiide specie: Leaves either na rrowly linear or entire or pide pinnatifid (or in S. lyonii often bipin- -pocomy 28 ‘with reer i mostly elongate segments; plants taprooted, with numerous stems endi ate tacone o. y below oe UFFRUTICOSI. coer evident ATE than linear, merely tocthan or entire (or ae me pepnatiionnte in S. clarkianus) ; herbs, fibrous-rooted except for S. fremontit. II. Trr Leaves basally disposed, the basal or lower cauline pact well developed, often tufted, thet cui evi- dently and progressively reduced (or the stem scapose). Plants with the numerous "yr ae — arising hese a ver age — short- nage crown withou (except goss nae pe a Ste sulus) ae more elongate cau udex or rhizo eaves entire od ate, or a integerrimus occasionally Petes As sublobed; “herbage crisp- hairy to pS gp eC i a nee gaa or tom . COLUMBIA Plants taprooted, or more often fibrou a from ascending or horisontl, simple or often ca og caudex phe a creeping rhizome; ne Boceine variously fl or tomentose to al iry. Leaves all sharply dentate or gta age sg to , soon (in forms of S. sphaero- cephalu: >, all ye never crenate, lobed, nnatifid; bracteoles br to = Pegi dl well dev as are also Ro least’ the lowe: a leaves; _ y tom Fest villous, at wane when young. . Lv sat Ara Leaves otherwise (some or all of them crenate, serrate, lobed, or pinn lait or all entire in ‘ in vince which differ in other respects from the foregoing group; habit and vesture Breen lea entire or toothed to sometimes pinnately irregularly few-lobed, ordinarily panies pinnatifid, lyrate, nor at all palmately lobed. Herbage more = less tomentose at flowering time, sometimes only thinly or rather ¥ SO; Species a a dry habitats at various altitudes. : V. ToMeEnNTOsI. SUNFLOWER FAMILY 431 viesleaiee 8 b pcp from the first, or sometimes lightly Soceeee en when young — by Prati time except often for a little cae icaese tomentum the base and in the axils; species mostly of moist o ane habitats. acan EI. Basal leaves either pinnately dissected, deeply pinnatifid, lyrate- geenenns or tending to be shallowly a, —— os the latter case the lobes usually tending to bias again toothed a lade r less cordate; heads paca or — y. (For S. ionophyllus cli ds i pedline 4 ie e.) VII. Los Plants annual or winter-annua IX: poset Stems ota mt heduae petiolane; cordate, ivy-like; heads small, discoid; ee species of coastal Californ . MIKANIOIDEAE . SUFFRUTICOSI. Involucral bracts mostly about 21, — only 13; usually many of the leaves with some long lateral segments. Heads = sachin: small, the invo e 5-7 mm. high; leaves more dissected; bracteoles short and inconspicu- uch less than half * eae as the inv volucre; plants bearing dense tufts _ ersistent wool in the ps Bi : alee a the coast of southern repices . lyonii, Heads relatively large, the involucre wit t 7-11 “hi gh; leaves less dissected; list relatively and conspicuots some at them rally - jehut halt as long as the involucre; plants reat ford axillary wool; continental a qaactar r, from central California southward . S. douglasii. Involucral bracts mostly about 8 or about 13; bracteoles short and inconspicuous; leaves easils entire, sometimes a few of them with lateral segments Plants about 6-12 dm. tall, pond strongly woody, not dying back to the base each year; leaves very narrow, commonly about 1(2) m wide or less; Santa Barbara and San Luis a po Cou ee California. mania Plants mostly 2.5—8 dm, tall, often somewhat woody below but oon wy vip Nay to the base ale year; leaves a little wider, the Bia ones commonly 2 mm Sa ar Be deol 1 Zam, White Moantaini and e ard. . S. spar . TRIANGULARES, Plants ibis often freely branched; leaves about 4 cm. Jon Lea ostly obovate to spatulate or broadly chance, up to about 2 cm. wide; plants eventually tap- ed, although the branches of the caudex y become elongate and rhizome- sera She a a ag in the ains. 5.5, fre sequins Ss cHaneeelale to linear-oblong, 5 mm. wide or less; perennial from a system of aa pr rhizomes, ithout a taproot; mountains of Mono Camke: California. attersonensis. Plants taller, mols 2-15 dm. tall or more, the stems generally simple below the tailadeiaiecines leaf-blades mostly ng. nires or niet of the leaves irregularly pinnatilobate or incised; leaves neither Marto’ nor regularly serrate; Sierra Nevada. . clarkianus. Leaves src Hag dentate to serrate or occasionally subentire, never sinustilobate or incised; widespread specie Leaves, except sometimes for the reduced upper ones, petiolate, or tapering to a narrow petioliform At ead the lower te ves triangular with i) Itoid t beordate b . S. triangularis. Leaves all fy 6a to the b t 1 . S. serra Some or all of the pete "Ses leaves ‘sessile with ee more or less clasping base; northeast Bracesem and stwa oi , oe Crass III. CoLuMBIANI. Herbage loosely crisp-villous or arachnoid-villous with ev i enae Pepoeneeppizeed hairs, much of the pubescence sooner or later deciduous; widespread, mostly in well-drai Heads seoells eudinte (discoid in var. vaseyt of Washi — a northern Oregon), = a eed or e flow: widespread. 1 St Heads discoid or ae mes with 1 or 2 short rays and with about 15-20 flowers; California. 12. S. aronicoides Biers pigeons? glabrous from the first, sometimes with a very little arachnoid tomentum in the Uidlaccenncs. seldom as many as a dozen, apparently always radiate; plants of dry to fr ggesince moist habi- ta tat : northeastern Oregon and eastward. 1 sulus. ae more numerous, seldom less than a dozen in well-developed plants, often very eke: radiate or often oo plants art of wet - pep and other wet places. "Robust, more or less glaucous plants 0 dm. tall; leaves generally entire, occasionally seremreerty oothed ; ge eso widespread, tolerant ee alk 14. S. hydrophilus Less sauie scarcely glaucous plants 3-10 dm. eit; Mekeved generally ot, cag ag rarely sihendive: plants intolerant of alkali. S. foetidus. Vv GENTES. Heads discoid; San Luis ae County to San Bernardino County, California. 15. S. astephanus. Heads radiate; more northern deine species. Heads about 3-30, aa medium-sized (involucre 3.5-12 mm. high), erect or in S. elmeri sometimes slightly nodding. — bracts not dark-tip epet rays cone wie mostly 1.5—2 cm. long; oa 9-12 mm. high; plants 3-7 dm. tall; Cah . S. layneae. Involucral re usually with satan to prom cniinent, black or brownish pies or the plants otherwise syne Vv Plants aS single-stemmed from a short shionsae: thinly tomentulose when young, later often or less glabrate. Leetatal bracts with minute, black or brownish tips; species not boreal, in our range not extending as far ee as Washington Leaves slightly denticulate or entire; plants mostly 3-8 dm. ath involucral bracts mostly about 21, acs nly oh achenes hispidulous or sometim Asia pipet Oregon, north- eastern Nevada, oak no; ard. —— ocephalus. stly salient] sieing pete a Base Boe A sacepl paeagen roe leas abcat 23, piss as man ; achenes glabrous; Sierra ‘Nevada sn White Mountains, Cali forn’ os 3 etn bracts with very conspicu us black bier commonly about s rarely iy up to 21 i ee Beodes extending south in seo seme to the Olympic Mountains a dugens. 432 COMPOSITAE Plants with several stems from a well-developed branching caudex, subtomentosely rp eoiitgees il tchee Mountains of central W ashington no ges aca uthern ritish Columbia. 19. S. elm Heads solitary or 2, large shuablos gly 11-17 mm. high), nodding; Olympic Mositeion of Wasbiuetia: (The Californian, flame-flowered S. greenet might mistakenly be sought here except for the erect heads.) 20. S. neowebsteri, V. TomeENTOSI. Heads several or rather many Leaves nearly all tufted et the base, the c ne ones few and abruptly very much reduced ms ap- pearing subs ; basal leaves Prone ack toothed ri occasionally gases mountains of: pce es alifornia. . S. bernardinus, Leaves not all tufted at the base, the cauline ones tho ugh evidently smaller than pe basal relatively better developed, and the stem not appearing aaiae capose; basal “ig ves entire to siteatele lobe. not teemarly toothed; more northern plan sometimes irregularly aad abana relatively thin or even obscure; robust plants, 2 2-7 dm. tall, _ relatively long and ender leaves; west of the Cas cade Mountains in Oregon and Washington. 21. - macounit. Tomentum relatively dense though sometimes partly deciduous; smaller plants, mostly 1—3(4) dm Era oo various; plants occurring in and more especially east of the Cascade Mou mones and Sierra Nevada, extending farther west only in the Klamath region of f southern ¢ Oreg co solitary or sometimes 2-6. eads large, the involucr 14 mm. high, the - ar ngd 1.5-3 cm. wide. z lowers pellon:; mountains ots southern Calif ed-o 24. (3 yikes al ; North Coa sh ef aagoaieoe 25 Heads smaller, the sient ucre er ‘ mm. high, ‘hd disk 7-15 mm. Gencre don a branching caudex; plants about ee 3) dm. tall or i 2 Pit Se Nevada and east- ard. . S. werneriaefolius. Stems solitary from a short, simple or tenia caudex; heads notably long- ‘pedunctlate; cae about dm, tall; Josephine County, Oreg . S. hesper: Heads solitary or occasionally 2 or 3, ordina I rily radiate (but d pach the hockh of our area). ut oid forms of S. ate ws pee are net uncommon nd -Seaghoalggey and _ a n s “he 2 of the Tomentosi might be sought here are evidently tomentose at the top of the peduncle and base of the involucre Plants with 1 or more, r less sai Sena cauline leaves, scarcely scapose, ‘ea less than 1 dm. tall; rhizome short. a. very slender; widespread in wet alpine and subalpine meadows. . S. subnudus Roots oe meeet and ripe pt drier, more ro ig and exposed alpine habitats; boreal species, pi Fa south ange t ntral va shington eS Plants patie’ aie capose, resedifolius. ss than 1 dm. tall, the asia merely bearing 1 or - minute peace rhizome elongate, Siesae pee Watiown Mountains of Ore’ 30. S. por Heads ordinarily several, sometimes only 1 or 2 in the mostly “discoid species, S. pauciflorus. Head character istically areca (rare radiate forms occur, chiefly in company with the normal discoid nts). Heads 1-6, rarely 12, orange or reddish; “ heelys thick and firm, crenate; alpine and sub- alpine plants; boreal species, extending south into the hi gh Sierra Nevada. 31. S. paucifilor Heads more numerous, mostly 6-40, yellow; leaves relatively thin and lax, serrate to cine -pinnatifid; stream banks and moist woods; northern species, in our range rarely extending — of north- ern Washington. 32. S. indec Heads esgelvaitai geen 5 radiate (rare discoid forms of some of these species occur, chiefly in company with the normal radiate plants). Basal oe: or some of them, cordate or pinnatifid, at least toward the eir Basal leaves not cordate or Sere Heads relatively large, County, Califecnia. Heads —— the Iaith an serial, sharply toothed; cauline leaves a: ps eagon ses; leaves t 33. S. pseudaur , though sometimes .subrotund. involucre 9-12 mm. high, rays mostly ae ee i Eldorado lay —9 mm. high, the rays mostly 5-12 mm. long. vag slong relatively Soe up t o about 7 dm. gr mostly with rather numer- nto leaves all entire, or care ety some of the cauline leaves too —_ plants a on serpettica in Caiiinenia: 36. S. clevelan Plants pers or not at all glaucous, commonly but not always smaller and fewer- cle than A ie orig eee fome or all of ine most leaves generally toothed to pinnatifid; widespread plants y not abiting serpentin Lente prageohban thin and lax, a basal ones mostly elliptic or ee crenate or ser- o subentire ae Ae eo Leaves Felatively thi “ and fi he basal ones mostly silage to und, coarsely nate to shallowly bate. wavy, or subentire. 55°, rutin, VII. Los. Plants fibrous-rooted from a simple or branched poate or — rhizome; herbage glabrous or very Piaets robust, le: nearly afy-stemmed, commonly pias tall; leaves all pinnatifid, the lower lyrate; ~ag ‘elatively ~~ pe ai the gf cebnin mostly 7— ns high and es ‘ ——— County, California o San ispo County and inland to cha Tehachapi Mount. ewer ae Plants goat, atone. 1 in one or more (us nts commonly ~ 0.5-2(4) naked; Olym ore) Aes from the alee more northern species. dm. tall; basal leaves mostly lyrate-pinnatifid; stem eae nearly c Mountains, and in the Cascade a A near hing 8. S. flettii. Plants Pcaaeee: ian 1-6 dm. tall; basal leaves more often shallowly palmately lobulate, sometimes yrate. sechipcageo ns glabrou: es relatively thin and lax; Cascade region from southern Washington Ore regon and ges less commonly, in the Willamette — _— oe —— ~ seit si hack from the ocea ie usually provided mi few to numerous coarse, conspicuously porn oe as ie rarely labrous; leaves rela aueay thick and firm; along the coast from ~~ — of og Columbia iver to northern Californ Plants with a more or less well-devel ices which may be arneied by a “aaa rag ronal herbage generally thinly tomentulose, varying to sometimes essentially glabrous. SUNFLOWER FAMILY 433 Heads ab (ab = ag ere mostly 8-12 mm. high, the dry disk-corollas mostly 7-10 mm. long, the 3.5-6 ; heads ls 3-30; species of northern cismontane reo doe 42. S. eurycephalus. Heads for, a the involucre mostly 4-8(9) mm. hich, the dry disk-corollas mostly 4.5— 7. cae long, the es 2-3.5(4) mm. long; heads sein tim ore numerous; transmontane species, extending west ‘aout thern and northeastern California. 43. S. muitilobatu aa VIII. Jacopakae. A single species in our range. 44. S. jacobaea. IX. ANNUvI. Rays = ta ae and conspicuou 45. S. californicus. Rays v short and inconspicuous, Bra ee well developed, pea ol oa tipped; rays wanting; involucral bracts Stenlpa bit 21; intro- uced weed. “ Bertones. few and inconspicuous, eH mesic -tipped, or wanting; rays present but ute Tad en ae n S. mohavensts sometimes wanting; involucral bracts commonly about 3 a about 13. Lavalustal bracts mostly about 1 eaves coarsely dentate, eee clasping by a broad base; herbage glabrous; native ci species. 47. ’S. mohavens Leaves more or less pinn natifid, not notably expanded at the base; ae more or less ie. pube i introduced weed. . S. sylvaticu. Involucral bracts mostly about 8; nae es glabrous or nearly so; native species of the So otk Coast Ranges of California and adjacent islands. 49. S. aphanactis, , uted X. MIKANIOIDEAE, : A single species in our range, 50. S. mtkanioides. 1. Senecio lyénii A. Gray. Island Butterweed. Fig. 5742. Senecio lyontt A. Gray, Syn. Fl. N. Amer. ed. 2. 1°: 454. 1886. Taprooted perennial with numerous woody stems arising from the base, commonly 3-12 dm tall and eine back only part way to the ground each year; stems and leaves thinly tomentulose e si noe Rte! soon as aa or subglabrate except for the ie guest dense tufts o i 4-12 cm. lo i nh per. the Vv = (especially the larger ones) bip nati Baie er blunt, narrow, and often s rags more or less linear segments ; heads vera in.a6 ins inflorescence, the dis 15 mm. wide; eae 5-7 mm. high, i principal aoe: “mostly about 21; bracteoles Sonkpiinodd few. short; rays about 1 cm. long or les 5 Ca Ocean bluffs, pista sit pee aaa of the coast of southern California and adjacent Lower California. Type locality; San Clemente Island. March- ~Jun 2. Senecio douglasii DC. Shrubby Butterweed. Fig. 5743. psong douglasit DC. Prod. 6: ne 1837. necto regiomontanus DC. loc. pee douglasti var. tularensis “aan Aliso 4: 99. 1958. Taprooted perennial with numerous woody stems arising from the base, commonly 0.5~2 m tall he ame back only part way to the ground each year; leaves, young twigs, and often also the involu ucres to mentulose at first, often eventually (rarely early) more or less glabrate. Leaves ng the st ear, 4-l2¢ rm inflor ti the di voluc 1m heh: the Abcipal trace usually about 21, rarely only 13; bracteole eel degcloped: eodiel: _ generally some o 7 i rage half as long as the involucre r longer ; rays showy, about 1-1.5 cm. lon cibiek chs d plains, are ties cismontane Paes! from the lower slopes of the Sie Dr sag 4 te ff bate ap ges ett to Nay a and San Mateo Counties, south t Aged ee of ep aah Califorsis and adjacent Lower California; passing into t — € var. monoensis on the desert or basin side of its range in ae ren ceptors Type locality: sutania. ‘a sii var. monoénsis (Greene) teen. a Fl. Pl. Calif. 1149. 1925. (Senecio monoensis Green tee t et Obs. 1: 221. 1906.) os glabrous or ry rarely a Ted pele ira Da ya a little ai zz | sanniies’ more slender, and less shrubby than the typical form of the species, often dying back t ae ound each year. Leaves aver. aging a bs ttle — dissected and “Gractevtes averaging a little nthe: than in Fon douglasti. Southern Great Basin from southw n Utah to the White Mountains of Mono County, California, ‘and south- ward into northern tealieg and Tooeee cattordia: Mexico. Type locality: White Mountains. March-May, and sometimes again in the fall. 3. Senecio blochmaniae Greene. Blochman’s Butterweed. Fig. 5744. Senecio blochmaniae Greene, Erythea 1: 7. 1893. Perennial with numerous woody stems arising from the base, 4-12 dm. tall, apparently not dying back to the as each year; herbage glabrous or with. inconspi uous evanes to s numerous, well distributed along the stems, linear-filiform, entire, commonly 3-8 cm. long and about 84 * less, the lower ones dryi pais on ee branches ; eads several or rather ifor yeas agin the wide ; in- volucre 7-10 mm high, the principal ‘bracts mostly about 13; bra Sele aa and Bron. incon- spicuous ; rays abou t 1 cm. long or less; achenes aly canescent. and dunes and flood Sethi near me ane EE cide Sonoran Zone; Santa Barbara and San Luis Obispo Coun- ties, California: Type locality: Santa Maria River, San Louis Obispo "County. June—Oct. 434 COMPOSITAE 4. Senecio spartidides Torr. & Gray. Narrow-leaved Butterweed. Fig. 5745. Senecio spartioides Torr. & Gray, Fl. N. Amer. 2: 438. Senecio serra var. sanctus H. M. Hall, Univ. oe Pub. Bot. 3: 230. 1907. Taprooted perennial with numerous stems arising from the base, pone 2-8 dm. tall, somewhat woody below but commonly dying back to the ground each year; herbage glabrous. Leaves numerous, well distr inated gprs the stems, linear and entire coamite a few with some lateral segments), the lowermost reduced, the others long a 5 mm wie ea erous in a paniculate-corymbiform inflorescence, the disk mostly 5-10 mm volucre 5-11 mm. high, the principal bracts about 13 or sometimes only 8; pencenss ssl et few, 9 short; rays 7-15 mm. long; achenes canescent or imei glabro y open slopes, Arid Transition and Canadian Zones; southern California (San Bernardino Mountains) and northern Lower a niaer® PB = Sierra San Rabe eer north and east to New Mexico, western Nebraska, and Wyo . Typel ity: Sw ater River . i iy- Aug rise Cake var. peace shah . Holmgren ex Cronquist in Ferris, Contr. Dudley Herb. 5: 102. 1958. Herbage oo 2 ope im Seti short, crisped, sega hairs and obscurely ee rn - Pon dag, —_ brous; otherwise similar to low, small-headed plants the typical variety (plants 3 dm. tall; ucre 5—7 Seg hi oy Upper nitittades 4 in California in Moab County "meth to the White 5 potash and the adiaceet mountains or Mineral County, Nevada. Type locality: head of Crooked Creek, White Mountains, Mono County, California. 5. Senecio fremontii Torr. & aes Dwarf Mountain Butterweed. Fig. 5746. Senecio fremontit Torr. & Gray, Fl. N. Am : 445. 1843. Senecio ductoris Piper, Contr. U.S. Nat. ae ii 601. 1906. Glabrous perennial from a taproot and branching caudex (caudical branches — elongate and rhizome-like), freely te from the decumbent base, commonly about 1-1.5 dm. tall or t t someti longer and sprawling with ascending branches. pee thickish, om oO ar 1 adly o hare: heads oars Ce the Ra usually shortly os aa Seles State involucre about a igh. the principal beens mostly about 13; fea vincles few, usually short and bro oce nally narrower “ more elongate; rays 6-10 mm. long; achenes glabrous or ecacboaile es Teeciiesicis ues Wrdeie. “Tepe elite: Wind River Moun Waele ng. July Saat Replaced in _ southern Rocky Mountains and_o Y ihe vanterh rim of the yet Basin by the well-marked pa (Greene) Cronquist, Vasc. Pl. Pacif. Northw. 5: 290. 1955 (S. blitotdes Greene, Pittonia 4: 123. nec io | ee var, occidentalis A. Gray, Bot. Calif. 1: 618. 1876. (Senecio occidentalis Greene, Pit- ° Erospicuoualy poanes ueree pons of shires & nd adjacent Nevada and in the San Bernardino egerierred aaa A seis s from Mount Rose much resemble . fremontit. Type locality: Mamit Whitney and vicinity. J ug. 6. Senecio pattersonénsis Hoover. Mono Butterweed. Fig. 5747. menees nevolate Hoover, Leaflets West. Bot. 3: 256. 1943. Not Kirk, 1899. oover, op. cit. 5: 1947 Res Bees a small form of S. hidlsie but narrower-leaved and lacking a taproot; plant tic snttal from a branching eyuslens of wary rhiz gd about 1 dm. tall or less. Leaves sMisnicebbate to oe up to about 3.5 cm. long and 5 mm. wide. Open y piace s at very high elevations in me mountains, Hudsonian and Arctic-Alpine Zones; Sweetwater Mountains: ian rae Nevada, in Mono County, California. Type locality: Mount Patterson, Mono Comite u 7. Senecio clarkianus A. Gray. Clark’s Butterweed. Fig. 5748. Senecio clarkianus A. Gray, Proc. Amer. Acad. 7: 362. 1868. Fibrous-rooted, apparently single-stemm ed perennial about 6-12 dm. tall, the stem and_some- times also the leaves obscurely to evidently crisp-pubescent with multicellular hairs as in S. inte- isc eaves numerous, — par ianed along the stem, up to 25 cm. long and 6 cm. wide, grad upwards, the pering to a petiolar base, the cilddle and upper sessile ; some or all of the leaves pinnately fied 6 or lobed, mostly Bieticauert 4 so, with pointed segments, the others sharply and remotely dentate; heads several in corymbiform inflorescence, the terminal pe- duncle shortened and surpas: ssed by the share: each about 7— m. high, the bracts about 1 or sometimes only 13, not black-tipped ; bracteoles well developed, loose, and elongate; rays about 8 or 13, 1 cm. long or more; achenes glabro: Meadows and other moist place: € County, California. Type locality: sy nian, Zone in te Matias Mintice Gice Cahttiie Aub Sepe. SUNFLOWER FAMILY lasii 5746. Senecio fremontii i 5747. Senecio pattersonensis . 7 ei aniie 5745. Senecio spartioides 435 436 COMPOSITAE = Senecio eA 3 Hook. Arrowhead Butterweed. Fig. 5749. Ss h gularis Hook. Fl. B eres 1: 332. 1834 Si id. DC. Prod. ‘ 428. 1837. rapping subvestitus Howell, Erythea 3: fe “Ws, . hanseni Greene, op. cit. 124. enecio inioonopotick Greene, Pittonia 3: 306. 1896. one salions Rydb. Bull. Torrey Club 24: 298. 1897. var. subvestitus Greenm. Monog. Gatt. Senecio 1: 25. 1901. gularis var. trichophyllus St. John & Hardin, Mazama 11: 95. 1929. is var. trigonophyllus M. E. Peck, Man. Pl. paptinne 780. 1941. Sev ch stemmed, often rather coarse and lush, fibrous-rooted perennial, mostly 3-15 dm tall, niabecats By obscurely agin evidently) vi Pon ‘puber ulent. Leaves numerous, neither tufted at the base no ee, very strongly reduced upwards, the lower broadly or n narrowly gular to asrnar ia ate o pac ia ar-cor ea "father ne -petiolate, the upper with a on Setioles r becoming sessile, efit relatively narrower and less e ae or scarcely triangular ; leaf-blades only tty ¢ o n cm. w ge thed; heads few or rather numerous in a short, flat-topped Sndlove scence; involucre 7-10 mm. high, its principal bracts about 13 or sometimes only 8, often black-tipped ; bracteoles few, narrow and cei somewhat elon- gate; rays mostly about 8, sometimes only 5, 7-13 mm. long; achenes glabrou Stream banks and other moist places in the mountains, Canadi ian and Hudsonian Zonkst Alaska and southern Yukon to ‘Scskateiewnn, northern Sioa ‘Mexice, and California; in all the higher mountains of our range, pero pad "oe iam also at lower elevations. Type locality : “Moist Prairies among the Rocky Mountains.’’ [Canada] ept Senecio triangularis var. angustifélius G. N. Jones, Univ. Wash. Pub, Biol. 5: 257. 1936. A rather small, slender form with relatively narrow leaves, the upper strongly reduced and oe pe often — ae lower- most ones triangular or subtriangular. Sphagnum bogs at low elevations near the t from Vanco sland to southern Oregon. Type ocality: Raft River, pity Fe paeahad County, Waskicin Senecio gibbénsii Greene, Pittonia 2: an. a Rags hologically similar to normal, fairly robust S. angularis and seemingly not to be separated from it ering markedly from that species in its heart oa worthy of further investigation. Salt marshes at the Hirai a the Euloestia River 3 in both Oregon and Washi 9. Senecio sérra Hook. Tall Butterweed. Fig. 5750. Senecio serra Hook. Fl. Bor. Amer. 1: 333. 1834. Senecio andinus Nutt. Trans. Amer. Phil. Soc. II. 7: 409. 1841. Ss io | lat orr. & Gray. Fl. N. Amer. zs 440. 1843. Senecio serra var. integriusculus A. Gray, nord 1, N. Amer. 17: 287. 1884. Senecio serra Seer f andinus Rydb. Mem. N.Y. Bot. Gard. ts ie 00. Senecio serra [subsp.] Janceolatus Piper, Canty. U.S. Nat. Herb. 11: 601. 1906. Senecio millikeni stds Bot. Gaz. 41: 293. 1906 Senecio serra var. altior Jepson, Man. FI. Pl. Calif. 1150. 1925. t, fibrous-rooted perennial 5-15 or 20 dm sie hte or ser ah ye hte especially pate a base; stems clustered. Leaves cont cas not tufted at the base uch reduced upwards, the lower mostly oblanceolate, short-petiolate ta deciduous, r ae 2 er = more nc petiole or petiolifo co i cm. wwiile d or occa- sionally subentire ; = numerous on slender — narrow, almost cylindric, the disk commonly only m. wide; involucre 6-8 mm. high, its bracts commonly about 8 or about 13, often Madi tisond ions oles few, a Si nae oon rather elongate; rays few, commonly about 5 or 8, 5-8 mm. long; — es glabro reat s and og open place the foot sila ons at moderate elevations in the mountains, in favorable aoa tat: base of the Caande Mountains ef shington, south t crag an nd wester n Wyom ng, Utah, moe nigel Cali n W s n 10. Senecio crassulus A. Gray. Mountain Meadow Butterweed. Fig. 5751. Senecio crassulus A. Gray, Proc. Amer. Acad. 19: 54. 1883. Senecio crassulus var. cusickii (Piper) Greenm. ex M. E. Peck, Man. Pl. Oregon 780. 1941, without Latin diag- nosis, Glabrous, fibrous-rooted perennial a7 dm. tall from a short, erect or ascending caudex, this ” branched and bearing several stems. Leaves thickish, entire to sharply dentate, the basal nes sorietiee larger than ose below, sometimes more or less strongly uced ; heads several (rarely s eee: & corsmbitorm, usually open inflorescence, the terminal head little if at all overto so. gt m. wide; re abou mm. high, s achenes In dry hell moist soil in open woods, meadows, and other open places from the foothills to uti high elevations: + in ae mountains, yea Fa = ig Hudsonian Zones; Sone Montana to nenthensperse Oregon south to South Dakota, New Mexico, and Utah. Type locality: mountains of Colorado. June-Aug SUNFLOWER FAMILY 437 11. Senecio integérrimus var. exaltatus a7 ) Cronquist. Single-stemmed Butte Se 5752 enecio exaltatus Nutt. Trans. Amer. Phil. Soc, II. 7: 41. Senecio ingens var. sealant D, C. Eaton Ba King Mer tng 1871. Greene, Pittonia 3: 17 a — densatus Greene, ie cit. 298, 18 S io atriapiculatus Rydb. M ee Ei SO 442. 1900. ra ne oidcous Senec yd Senecio integerrimus var. Croke Crotraile: Leafiets West. Bot. 6: 48. 1950. Stout, fibrous- po perennial from short, erect crown; stems solitary, 2-7 dm. tall; ace. j Sanh to arachnoid-villous with ae tose rie when young, generally mor or le labrate in age. Leaves uke to pnariarks dentate, the ba nes petiolate, the blade and petiole commonly cm. long a - wide; cauline leaves progressively red up : ing sessile; heads several or rather numerous in an sted inflorescence, the peduncle of the terminal head thickened and much shorter than the Peper involucre abou mm. high, he principal bracts commonly about 21, sometimes only 1 ortly but evidently “black: -tipped ; t bracteoles few, ee ly narrow and rather elongate; rays 6- iG mm. long, yellow; achenes gla- brous or es Moderately dry to vadier moist open places and open woods, from the valleys to near timberline, ordinarily in well- Samed — Arid pcan a Canadian Zones; seathirar Alberts and eon Serta ra to Colorado, Nevada, oe California; in nge found chiefly east of the Cascade — Sierra Nevada s but occasional nth mette Valles and re the ian ath region of southwestern Oregon and nort fo rac “Ca aliforai a; on pated ned arly or ce to the coast. Type locate: “plains of Oregon, near cee outlet of the Wahlamet.” May~Ju Typical S. integerrimus is a plant of the Great Plains. In our range, several additional varieties are 2 as Yolre: 5749. Senecio triangularis eg om mecio serra } oe Senecio integerrimus 438 COMPOSITAE Senecio ietgmeseinews var. gcheolenee. (A. G may), Cronquist, Leaflets West. Bot. 6:48. 1950. (Senecio cordatus Nutt. Trans. Amer. Phil. Soc : 411. 1841; Senecio ines var. ochroleucus A. Gray, Syn. Fl. N, Amer. 17: 388. 1884; S. evxaltatus pelle S| Se i Bing Con U.S. Nat. Herb. 11: 600. 1906.) Basal leaves tending to have deltoid or subcordate blade though sometimes “i narrow as in typical var. exaltatus (which with deltoid o ~ r ad as in v. xaltatu Commonly in slightly more A habitats than var. exaltatus, ee in woodlands; western ntana, across north- ern Idaho to the Vascahe Mountains of Washin 4 , south to n n Oregon and reputedly to foarn California. Type locality: Columbia River, Klickitat County, Washin gton. $s io integerrimus var. major (A. Gis) Cronquist, sing i oa 1958. (Senecio eurycephalus var. major A. Gray in — Pacif. R. Rep. 4: 111. rie S. men ndocine Proc. Amer. Ac = 72 362; pri a: whippleanus A. Gray, Syn. FI. i Sar ip : 384. 1884; es "condensin s Davy, ae day 3: 117. ne : se r ‘ Bot, ~ : Wa foi § Pl. Calif. 1152. 1925.) Leaves subentire to toothed or even ilsiuie ded: hheads ‘mostly ged and. ag vely large, the involucre 7-12 mm. high, its bracts twit bate or sometimes somewhat purplish tips; “ee cm. long; otherwise as in var. exaltatus. On the west slope of the Sierra Nevada and in othe Nerth Cone’ Baie of Cali- fornia; also in the White Mountains in Nevada. Type locality: ‘near Murphy’s [Calaveras County], Califor nia.” Senecio integerrimus ov vaseyi (Greenm.) Cronquist, Vasc. Pl. Pacif. Northw. 5: 293. 1955. (Sen vaseyt serena. I n Piper, Con US. Nat Herb. 11: 600. 106. Heads not very numerous; principal involueral bracts co: monly. B Baia oT. evidently black. -tipped; rays wanting; ovis as in var. eraltatus. Ey and Wenatchee 1 region of central Washington; also in the Wallowa alcaictathe of northeastern Oregon pe Aiealiiy: ashington be Senecio aronicoides DC. California Butterweed. Fig. 5753. icoid rod. 6: 426. 1837. var. sini ulus A. Gray, Pacif. R. Rep. 4: 111. 1857. 7 e, Pittonia 2: 166. 1891 Senecio leptolepis Gib: F 1 Fran. 468. 189 8 :, Dens. Stout, fibrous-rooted perennial from a very short, erect crown; stems solitary, 3-9 dm. tall; herbage srachnelt aitiecs with pes loose hairs at least when young. Leaves highly variable, subentire to i often toothed or even De erred incised-lobulate, the basal and lower cauline ones 8-30 ide, the long (petiole gear eds d 2-12 cm. wi blade often deltoid or subcordate, often hate hi narrower; ¢ auline leaves prabvectvely. reduced upward, becoming sessile; heads few to more often — umerous in a oose or compact sa ole the terminal peduncle shorter than the other vale e 4-8 m m. high, with mostly about 8 or about 13 bracts, <— with pale aw “ary sees in, in op in the foothills and lower mountains, Upper Sonoran and Arid Transition Foote Bride rim of the —— Valley of pag sc Behing the vicinity of Mount Shasta south to the San Franc o Bas m5 region and thence along the western the Sierra Nevada to Fresno County. Type icality? California. April—July. 13. Senecio foétidus Howell. Sweet-marsh Butterweed. Fig. 5754. Senecio foetidus Howell, Fl. N.W. Amer. 377. 1900 labrous but scarcely aplenty hada Lae perennial from a very short erect crown, 3-1 tall, Pi stems solitary or more commonly clustered. Leaves thickish and somewhat succulent, pa Saat sharply dentate, the basal od lowermost cauline ones petiolate, with mostly elliptic or broadly oblanceolate blade 6-25 cm. long and 2-7 cm. wi e; middle and upper leaves rea strongly and progressively reduced, becoming sessile; heads more or less numerous (rarely as few as 8) in a congested inflorescence; involucre 6-9 mm. high, the bracts nt ie about C13 ometimes only 8, minutely black-tipped; bracteoles few, narrow and rather elongate; rays race typically ut 5, = o about 8 mm. long or more often wanti enes glabrous. Wet meadows in the mountains and foothills, Arid cranes sas ashington east of the Cascade Moun- tains aad adjacent southern British Columbia east to western south to no Eethcketern ete Sei northern Nevada, and central and southwestern Idaho. Type locality: aires Vatiay: Washington. May-July. ng foetidus var. hydro ophiloides (Rydb.) T. Rog ge ahaa ex Cronquist in Ferris. Cate Dudley Herb. 5: 102,:. 1958. (Senecio h SRN ses Rydb. Mem el ie << 1: 441. 1900.) Single-stemmed; leaves thi rn or sometimes thin; head an open, corymbito ans inflor the terminal one short-pedunculate and Mes others; rays melt 5 or 8, 5-10 mm. long; "otherwiae ‘ike typical >. foetidus. Wet meadows ong moist sti d foo thills, western ) Mcatase to northeastern Washington and northeastern Ore: Type pee hes Forest, Nez Perce County, Idaho. 5 Senecio hydréphilus Nutt. Alkali-marsh Butterweed. Fig. 5755. y drophilus Nutt. Trans. Amer. Phil. Soc. II. 7: 411. 1841. be phil: Yr. pactficu. ens Greene, P Pittonia 1: 220. 1888 Senecio paitires Rydb. Fi. Rocky Mts. 917. Glabrous, > oat et or less glaucous, stout, hollow-stemmed, fibrous-rooted perennial from a short most ? erect cro 20 dm. tall, the gang often sa shin clustered. Leaves thick mewhat succulent, commonly entir early Imes more evidently callous-too al or lowermost cauline one long-petiolate, tae large, mostly narrowly elliptic blade c long and 2-5 cm. ‘ven larger ; middle and upper leaves few, strongl and progressively reduced, coming ine ti usually rather numerous r ; in- io 5-8 mm. high, rather narrow, its bracts common y about 8, sometimes 13, often black- ; bracteoles few, narrow, and short; rays few, about 4-8 mm. long or often wanting ; Fi ae andsp rous. py places in the valleys and foothills, tolerant of salt and alkali, Upper Sonoran and Transition Zones; southern British pe to the San Francisco Ba: region, east i ) Montana, Bo uth Dakota, and Colorado. Type locality: ““Ham’s Fork of the Colorado of the West yoming ].”’ y—July SUNFLOWER FAMILY 439 Ae: iia astéphanus Greene. San Luis Obispo Butterweed. Fig. 5756. Greene, Pittonia 1: 174. 1888. Seeecks ditcetaiains Pe ted Erythea 2: 85. 1894. apparently a ae ned perennial about 6 dm. tall ag probably often taller, con- ather thinly tomentulose nearly throughout. Leaves sha rply dentate, the lower- (including the artis “ge a nd 7 cm. wide; ee and upper ed, becoming sessile, numerous ; hea veral in an fee large for the gta “tiseoid, yellow ; sek glabrous. : San Luis Obispo County to San pesuerdien County, California; in- Luis Obispo County. May-Fen Coarse, piped a os he largest, up to m. long oe duc ery compact inflorescence, rat sigh mountains, Upper Sonoran Zon shreds known. Ty pe locality: cane of San 5755. Senecio hydrophilus 5756. Senecio astephanus 5753. S i icoi 5754. Senecio foetidus 440 COMPOSITAE 16. Senecio sphaerocéphalus Greene. Mountain-marsh Butterweed. Fig. 5757. Senecio sphaerocephalus Greene, Pittonia 3: 106. 1896. Fibrous-rooted perennial from ort, thick, horizontal or sometimes ascending rhizome 3-8 dm. tall, thinly tomentulose and eric eventually (rarely early) glabrate; stems mostly aris- ing singly from the rhizome. Leaves slightly denticulate or entire, the lowermost ones oblanceolate ade petiole 5- cm, long and cm. and upper leaves few, strongly and progressively reduced, becoming sessile; heads about 3-25 in a usually rather compact cyme, the terminal seacticle. often eee d and thi ckened ; involucre mm. high, its br rani commonly about 21, rarely only 13, bearing a minute, brownish or black tip; bracteoles few, n w, shor nhac siccel elongate; disk 8-15 mm. wide; rays 6-10 mm. long; achenes usually biajihaious but in our forms glabro Wet meadows in and near the ws awa Canadian oe. Montana to northeastern Oregon (Wallowa Moun- tains) south to Colorado and northea: n Nev. Type loca lity: Deeth, Humboldt River, Nevada. past Aug. Sen s Howell, FI. N. fy a 7. 1900. Similar to S. sphaerocephalus, to which it perhaps should ve Fy in teat saa glabrate and with elongate, relatively narrow, lower leaves. Type loealits : Lake Labish near Salem, Oregon, where it has been collecend pase times. June. 17. Senecio scorzonélla Greene. Sierra Butterweed. Fig. 5758. Senecio ella Greene, Pittonia 3: 90. 1896. Senecio willed Cobe e, Fl. Fran. 469. 1897. Senecio covillet var. Sock Jepson, Man. FI. Pl. Calif. 1152. 1925. Stems mostly arising singly from a short, thick, — ntal or ascending rhizome, 2-5 dm. tall, rarely taller; herbage thinly tomentulose when ng, later often more or less. glabrate. asa u shor strongly an ane progressively reduced, beco sile; heads about 10-30 in a compact cyme, the aes on ne ona short thick ee ny ayeriance ed by the others; involucre 3.5 mm. the m sigue ely black-tipped bracts commonly about 13, sometimes 8 or 21; bracte oles few So ‘ten fairly well developed ; rays scarcely 1 cm. long or sometimes wanting ; achenes glabrous. Meadows and moist slopes below timberline, ae and Hudsonian Zones; Sierra Nevada and adjacent southern alt — renin from Mount Lassen to Tulare a and in the White Mountains of Inyo County, California. Type lo y: - northeastern California.” July—Aug 18. Senecio lugens Richards. Black- ees Butterweed. Fig. 5759. Senecio lugens Richards. in Frankl. 1st Journ. Bot. App. 747. 1823 Fibrous-rooted ccarargn with the stems mostly abe 2 singly from a short, thick, ascending or fhorisontal rhizome, 1-5 dm. tall; hates thinly tomentulose at first, generally subglabrate by owering time. Lewes ost aed nely (or eT acon gvinein coarsely) but not closely callous- toothed, well-developed, mostly 5-20 cm. long and 8-30 mm wide, the oblanceolate to sometimes elliptic or nearly obovate blade pot a to the petiole = netiolita orm base; cauline leaves few, strongly and es reduc _upwards, eet aie sile; heads mostly 3-21 in a compact c t te i i = e others ucre 5-8 mm. high, the bracts very conspicuously black-tipped, commonly about 13, rarely 21; bracteoles chee enti — developed, evidently black- tipped: disk 8-13 mm. wide; rays 7-15 mm. long; achenes gla nee grassy alpine or arctic slopes, and rich northern woods, Boreal Zone; Alaska and Yukon south to af Olymate Mountains of Washington and eotie to northern Wyoming. Not known from a Baga ork Cascade Mountains. Type locality: Bloody Fall, on the Coppermine River, Yukon Territor ry. July— 19. Senecio élmeri Piper. Elmer’s Butterweed. Fig. 5760. Senecio elmeri Piper, Erythea 7: 173. 1899, __ Perenn ial from a well-dev eloped hranching caudex, 1-3 dm. tall, aig Sage arachnoid- villous at first, only thinly so (or even Rabcate) at flowering time. Basal or lower cauline leaves the largest, the basal sometimes tufte n separate short shoots, pss ly eae broadly m. lon i 3 cm. wide, sharply ards an n ; heads several in a corymbiform inflorescence, the peduncles often flexuous so that the heads may be a little nodding ; involucre 7-12 mm. high, bracts mostly about 13, villosulose at the often dark or blackish wad bracteoles narrow but well developed; disk 8-15 mm. wide; rays 8-16 mm ous. d other rocky places, Hudsonian and Arctic-Alpine Zones; at alpine stations in the Cascade and i ‘Wenatchee. setts of cane and northern Washington, extending into southern British Columbia. Type locality: “North Fork of Bridge Creek, Okanogan County, Wash.” July—Aug. 20. Senecio neowébsteri Blake. Olympic Butterweed. Fig. 5761. Senecio websteri Greenm. Bot. Gaz. 53: 511. par Pigg hs Hook. 1846. Senecio neowebsteri Blake, Leaflets West. Bot Piha, from coher Mode veloped, SORES caudex or short rhizome, 0.5-2 dm. tall, r less arachnoid-villous at first and often also somewhat t glandular, later more or less wish. Basal or lower eauliie leaves the hatcet the basal sometimes tufted on separate short SUNFLOWER FAMILY 441 shoots, petiole the blade pune obiapenate 2, subernnes up to about 7 cm. long and 4 c wide, dentate Es by ens la cauline leaves few, r less eh ngly reduced or the middle Bais sometimes sla s those he low: beads solitary (rarely 2), mostly nodding, large, the involuc its 7 tots. high the ens mo .5-2.5 cm. wide; Savele ucral bracts commonly about cay villosutow at the usually pal bracteote Ww ell de) cloped and cee more than half as ] the ety ah aes ire w, 10 15 mm. long; achenes glabro fit slopes, Hudsonian and scree Alpine gi il ae Hissia of uk a Type locality: Mount Angeles, Clallam (pas Aug.—Sep 5757. Senecio sphaerocephalus 5759. Senecio lugens 5758. Senecio scorzonella 5760. Senecio elmeri 442 COMPOSITAE 21. Senecio macounii Greene. Puget Butterweed. Fig. 5762. wanacie festigtaas: Nutt. Suse A _, Soc. IT. 7: 410. 1841. Not Schweinf. 1824. er it Senecio spatuliformis Heller, Bal ‘Tore ip 26: rar 1899. Sen ene, Lea s Bot. Obs. 909. eas ilar to S. canus but ge er and more — on 2-7 dm. tall, more thinly tomentose i tomentum sometimes obscure by flowering time), and with consistently narrow leaves, the basal ones commonly merely ibintera inte: heads getiaicis larger, the involucre up to 1c m. high and dry n places, Upper Sonoran and Humid Transition Zones; west of the nen — tains from Vancouver ‘aend to seethara Oregon, mostly in the Puget trough. Type locality: Mou nson V cei Island. June-July. 22. Senecio — es Woolly Butterweed. Fig. 5763. Senecio canus Hook. Fl. Bor. Am 333 eae ie ewrerer Nutt. Trans. oe mer Phil. a ts: 7: 412, 1841. , Bull. Torrey Club 8: a Briges Senecio ligulifolius Greene, Leaflets Bot. Obs. 909. enecto canus [var.] purshianus A. Nels. in sath es 9 nase a Bot. Rocky Mts. 581. 1909. Senect 10 howellia var. Sette gee Bot. Gaz. 48: 148. 19 Sene ecio ciesignams is Greenm. ty o. Bot. Gard. 1: 286. 1914. Y p Greenm te S i bolus £ i saben ace eae, op. cit. 269. Several-stemmed perennial from a more or less branching caudex, often with an evident — taproot, 1-3( m. tall, more or less strongly white-tomentose, often less so in age, the upper surfaces of the leaves sometimes glabrate. Basal lowermost cau ie leaves more or less tufted, m narrowly anceolate Bs broadly elliptic or ovate, the blade mostly 14 5 > wide, entire metimes irregularly subpinnately ort or elongate petiole; middl pper leaves few, often iairecky oothed or subpinnatifid, strongly and pro- gressively reduced, becoming + gears but the stem generally not appearing a aed — several ; involucre 4-8 mm. high, the bracts about 13 or about 2; bracteoles wanting or and Pp 1 inconspicuous; disk 6-13 mm. aes rays mostly 6-13 mm. long or rarely wanting; scheits glabrous. Dry, n, often rocky places, from the a —_ foothills to timberline or above, Arid Transition and Boreal Zones; southern British co ageana to Saskate sete! outh to California, Colorado, and Nebraska; in our range occurring east of the Cas summits in tnahe P ge) norteade "‘Origte. eens rine: - near the coast in the Klamath region o ot et ta Oregon and fsa along the se aig Mountains and Sierra Nevada to Tulare County, California. Type locality: banks of the Saskatchewan Riv May-—Aug 23. Senecio bernardinus Greene. San Bernardino Butterweed. Fig. 5764. Senecio bernardinus Greene apt aze i 298. 1898. Senecio inapiaieee var. eeisirdinn H. M. Hall, Univ. Calif. Pub. Bot. 3: 232. 1907. ial with 1 ae erect stems fro audex, 1-3 dm. tall, rather thinly tomentulose or in Ry ‘eee. Sp hot ong at the base, nerioiete the elliptic to obovate or subrotund blade 0.5-2.5 cm. long and 5-15 m wae e, more or less toothed (often regularly s o) or subentire; ae eaves few nc very m mah uced, po stems somewhat scapiform sheaths abou t 3-20; involucre 5-8 mm. high, the bracts abcak 21 or about 13; bracteoles inconspicuous or eater: disk seen 8-14 mm. wide; s 6-10 mm. lo ici’ achenes glabrous o r hispidulou Dry open places, ie Transition and Canadian Zones; San Bernardino Mountains, rece Type locality: Bear Valley, San Bernardino Mountains. June—Aug 24. Senecio ionéphyllus Greene. Tehachapi Butterweed. Fig. 5765. mentee fonophylius — Pe ange 20. 1899, t. Gaz. ge 462. Seperee ionophyilus v var. sind ane H. M. Hall, Hoa Calif. Pub. Bot. 3: 232. 1907. S var. sparsilobatus Greenm. Ann. Mo. Bot. Gard. 5: 46. 1918. S. io ionophyll: wine enm. loc. cit PAYVUUS Var. intr Seve eral- Se ence a ial from a branching caudex, 1.5—4 dm. tall, sir tomentulose or metimes a elatwede: Biel id: lowermost cauline leaves with obovate o rotund or flabel- late, coarsely blunt-toothe d, sometimes more or less lyrate blade up to scat "3 acm. nll and , peti th m ; involucre about 8-10 mm. high, the principal bracts about 21 or sometimes only 13; bracteoles few but sometimes well ror: or none ust di ~ necb Po asout 8-10 mm. long; rays about 8-14 mm. long; achenes 5-6.5 g, glabro Dry open slopes in the Saints tes icici and Canadian Zones; mountains of southern a from Tehachapi aoatherard Type locality: near the summit of the mountains south on Tehachapi. June—Jul 25. Senecio gréenei A. Gray. Flame Butterweed. Fig. 5766. Senecio greenei A. Gray, Proc. Amer. Acad. 10:75. 1874. Perennial from a branching, rhizome-like caudex, 1-3 dm. tall, thinly floccose-tomentose, eventually more or hese glabrate. Basal and lower cauline leaves well developed, the basal often SUNFLOWER FAMILY 443 444 COMPOSITAE tufted on separate short shoots, petiolate, the blade subrotund or a ovate, up to about 6 c long and broad, more or less dentate; middle and eae cauline leaves few and much relied: heads 1-3, naked-pedunculate, erect, relatively very large, the involucre 10-14 mm. high, o much shorter than the broad (1.5-3 cm.) disk, the bracts es 13 or about og, Bocieotts few or none; rays 1-3 cm. long, flame or red-orange like the disk; achenes glabro Brushy slopes, commonly on serpentine, Upper Sonoran and Transition Ronen: Coan Ranges of northern California. Type locality: near the Geysers, Lake County, California. May—Jun 26. Senecio werneriaefolius A. Gray. Alpine Rock Butterweed. Fig. 5767. ayeecte aureus ‘gph bbe Gray, Proc. Acad. Phila. 1863: 68. n, Mo, t. Ga 18 1863. rd. 5:56, 1918. Not oy muirit L. Bolus, 1915. Senecio speculicola J. T. Howell, an West. Bot. 4: 64. Lax, several-stemmed perennial from a loosely ae ing caudex, up to 1.5 dm. tall, thinly Sicine and often Se citaally glabrate. Leaves tufted at the base, oblanceolate to Reon ie or occasionally broader, tapering to the petiole or petiolar — entire or nea he whol abou i ry, open, often rocky places at high elevations in the ntains, Arctic-Alp Zones; oe hk Nevada east to northern New Mexico, Colorado, western South Dakota, siad ‘western’ Montana m2 locality: et the Rocky iscntains.. in oerery ery, fat. 39°—41°." Jin The bbs: aba plants of S. riaefolius aie ye ap oncaran d ~ stem ge ieee gr eads solitary or sometimes 2, the disk €, ye voluc 8m , the bracts about 21 or rarely only 13, ae long ; ach poe and ciiiee wet meadows, Hudsonian and Arctic-Alpine Zones; Washington to Montana and south- River yoke aan ne south to California (throughout the Sierra Nevada) and ‘Wyoming. Type locality: Columbia er. y-S purplish toward the tp ap: biacteples few, uate and rather elongate, or none; rays about 7-14 m 29. Senecio resedifélius Less. Dwarf Arctic Butterweed. Fig. 5770. Senecio resedifolius Less. Linnaea 6: 243. 1831. Sene, oto Tyailts Klatt, Ann. = Hofm. Wien 9: 365. 1894. Not Hook. f. 1853. Sen eene, Pittonia 4: 110, 1900. i reenm. hae . Mo. . Gard. 3: 101. 1916. _ bs ¥ om a branched caudex or short, rather thick rhizome, about 0.5-2 dm. tall, essen- tiathy pi opined or occasionally on a little persistent tomentum at the base and in the axils of the leaves. Leaves thickish, the basal ones mostly arbicidatcbvatet we eniform ven o| e, blunt- toothed or subentire, the b ptoa it 2.5 cm. long and wide, generally abruptly contracted to the petiole; cauline leaves reduced upwards ‘becomi g sessile, in the larger pl less pinnatifid ; xsi hse arse ted 2, the disk a -18 mm. wide, yel more often orange or olucre h, the bracts a _ 21 S rarely only 13, dactish = feast usally heccleles. averaging shorter and broader than in S. s us; rays variously dev: times as much as 14 mm. long and 4 mm. wide or - Wetasionally wanting ; ; Exposed situations at alpine or occasionally subalpine stations oe the h mountains, mostly upper Lata ar: pre Reape and Alaska to Ri frei nee south to central ice ington, sg ana, and northwestern yoming. Type locality: St. Lawrence Island n the Bering Sea. July—S. SUNFLOWER FAMILY 445 5769. Senecio subnudus 5770. Senecio resedifolius 5767. Senecio osetia 5768. Senecio hesperiu 30. Senecio pérteri Greene. Porter’s Butterweed. Fig. 5771. layed renifolius Porter in Port. & Coult. “ag Colo. 83. 1874. Not Schultz-Bip. 1845. Senecio porteri Greene, Pittonia ie 186. 189 fiecuce ooted perennial from a “wel developed, eens tae seroma creeping rhizome, gla- brous throughout, “essentially scapose, t cm. peduncle naked or with 1 or 2 minute bracts. Leaves SBA he thick, mostly eras crenate- oblate blade a to 2.5 cm. wide; heads solitary, fairly large the sone saben? involucre about . high. site ocky s (? talus es) in the ains, Arctic- Alpine Zone; Wallow a Mountains of northeastern Oregon (w ace re a only “otha a ici collection fe ith the collector’s note “very little seen’’) and in Colorado. Teel locality: White House Mountains, Colorado. Our plant appears to be identical with that ty claesiie: 446 COMPOSITAE 31. Senecio pauciflédrus Pursh. Rayless Alpine Butterweed. Fig. 5772. we lo paucifi Pursh, Fl. Amer. Sept. 529. 1814. Fi br rous-rooted perennial from a penile or slightly ee cane 1.54 dm. tall, glabrous or lightly floccose-tomentose when young. Leaves thickish, s what succulent in life, the basal ones goncvices gh 8 Bc to subr ottttid, abruptly paaiarked to Nthe truncate or shallowly creat base, crenate, petiolate; cauline leave § reduc ed and gee coming sessile, bluntly toothed or m less pinnatitd with blunt johes : heads m mane 2-6, rarely 12, orange or reddish, discoid or pane ith s - involucre 6-8 m «ey its bracts ee ut 21 or about 13, generally suffused with reddis h purple, at — aboue the "middle: bracteoles inconspicuous or wanting; disk 1-1.5 cm. wide ; achenes glabro Alpine and tae a Seated and moist cliffs, Boreal Zone; Alaska and Yukon to Labrador, south to Quebec, and to northern Vrouing, nor teeny na’ and northern Washington, and throughout the Sierra Nevada of California. Type locality: Labrad July—Aug. eplanty rs California eh to be more robust than the northern, more typical forms of the species, with euinehak as re numerous heads, and more often neonuee i cease forms. Many of the California specimens are wholly characteristic of S. pauciforus, howev and n nomic segregation seems possible. Not known to me on 52. Senecio indécorus Greene. Rayless Mountain Butterweed. Fig. 5773. Greene, Fl. Fran. 470. (Aug Ps | 1897 Ss io tdah Rydb, Bull. i — “ihe 183. 1900. Senecio p ifl p. fallax Greenm. in Piper, Contr. okey Nat. Herb. 11: 597. 1906. Senecio p ifl f fallax Fernald, ehndars ah: 225. 19 Fibrous-rooted perennial from a simple or oe Liner caudex, 3-8 dm. tall, glabrous or lightly floccose-tomentose when young. Leaves tom sigied thin, not succulent, the basal ones eeatts elliptic or broadly ovate, tapering or subtruncate at the 4 Se se, serrate sometimes in- cised, petiolate; cauline leaves sharply incised- pinnati, ‘the iad irregularly a gain 1 SOAS, reduced and becoming sessile upwards; heads mostly 6-40, , discoid or rarely with shor rays; involucre mostly 7-10 m in, tech i ts bracts 1 mostly about Sf sometimes on ay a8 often sbi tipped: bracteoles short and i os slg 6-1 wide; achenes glabro Moist woodlands, stream banks, swales, and a ii go Alaska and Yukon to Aaa ae n Washing- ea a Wyoming, and a pparently in northern California. Type locality : Pine Creek, Lassen Conner. "Calitoeata. uly—Aug It is ‘possible that the ty ype of S. indecorus Greene is merely an aberrant form of S. pseudaureus or a hybrid males S. pseudaureus and Ee pet and ere he proper name for the species here ‘Aoectibed:i is S. idahoensts ydb. 33. Senecio pseudatreus Rydb. ~~ Butterweed. Fig. 5774. Ss io pseud Rydb. Bull. Torrey Club 24: 298. (June .4 io p ifl var. jucundulus Jepson, Man. FI. Pl. aE Mert 1925. Fibediss rooted perenni a from a short, horizontal or ascending rhizome or caudex, 3-7 dm tall, lightly floccose-tomentose at first, soon essentially glabrous. Leaves relatively thin, not suc culent, the basal on eg aeniclote, the blade serrate, tending to be cordate, subcordate, or ‘rut cate at the base; caging leaves few and progressively reduced oe b ing sessile, more or less laciniate-pinnatifid at least toward their bases; heads several or disk about 8-13 mm. wide; involucre 5-8 mm. high, the bracts about 13 0 agra 2b hemeenees inconspicuous or tia rays a ets 6-10 mm. long; achenes glabrous. Stream banks, meadows, and moist woodlands in aed. near the aang “pu mostly vind aoe Zone; Hite res Columbia to Caskethewen om to tes atte ornia in the southern Sierra Nevada and i w Mexico. Type locali e Belt Mountains, Montan ug. tpt ar. pseudaureus, as Genarih rnc occurs in our range. The var. pr esr f Greene) Greenm., with nore eenee aged leaves, replaces the var. pseudaureus in the southern Rocky Mountains and on the northern reat Plains 34. Senecio paupérculus var. thomsoniénsis eloraaey Boiv. Canadian Butte 5775 rweed. Fig Senects mnalinoman ste. Greenm. are Nat. 25: 115. 1911. . thomsoniensis Greenm. op. cit. 116 Sensei flavovirens var. thomsonien. Li Gee eenm. Ann. Mo. Bot Aap 3: 169. 1916. enecio pestle var. thomsoniensis Boivy. Nat. Can. 75: 214. 1948, ook rooted perennial with a rather short, pe or slightly branched caudex, occasionally with some ng short, slender alin, mostly 2-7 dm. ta ll, lightly floccose-tomentose when young, soon glabrate except. frequen aati t the base and in the leaf-axils. Basal leaves petiolate, the blade oblanceolate * elliptic or occasionally suborbicular, crenate or serrate to subentire ; cauline leaves ors or less pinnatifid, the lower sometimes larger than the basal, the others conspicuously re- se and be ing sessile, all relatively thin and not at all succulent; heads eral, the disk 7-12 mm. wide ; involucre 9 mm. high, the bracts about 13 or about 21; bracteoles 1n- conspicuous 0: re rays about 5-10 mm. long or very rarely wanting; achenes glabrous or poniunm hispidulou am banks, swamps, m meadows, moist woods, and ist cliff: tl the foothill d valleys, sometimes at sone elevations in the mountains, Transition and Canned ‘a ‘ n Zones; louthern Natale tevatathorn Oregon and western Montana. Type locality: North Thompson River, Deitch Columbia. Ma The species as a whole ranges from Labeaaor te 0 southern Yukon, entitk to Virginia, New Mexico, and northern \* \ it vill! SUNFLOWER FAMILY tl I | Senecio porteri . Senecio paucifl Senecio pauperculus Senecio cymbalarioides 447 448 COMPOSITAE ens 35. Senecio cymbalaridides Nutt. — Mountain Butterweed. Fig. 5776. Senecio cymbalarioides Nutt. Trans. Amer. Phil. Soc ~ es 412, 384k, ecio onreey var. es realis Torr. & Gray, Fl. N. pod es 442. 1843. Ss , Pittonia 3: 88. 1896. Kea acss adewist Howell, Fl. NW. Amer. 379. 1900. Not S. adamsii Cheesm. 1896. Sealed preteomes Piper, Contr. U.S. Nat. Herb. 11: 598. 1906. Greene, Leaflets Bot. Obs. 2 190) Senecio sabsdorfis G nm. Bot. Gaz. 53: 511. 1912 Sen eS ymbelarioides var. borealis Greenm. Ann. NO Bot. Gard. 3: 177. 1916. brous-rooted perennial from a caudex or short rhizome, 1-5 dm. tall, glabrous or lightly floccose-tomentose when young. Leaves thickish, somewhat succulent in life, the basal ones long- petiolate, with mostly elliptic to subrotund tund-obo blade, wavy or coarsely Eendte to shallow ulate or entire; cauline leaves few and reduced, becomin acaaile Farms somewhat pinnatilobate, at least to es their bases; heads several or rather many, the 4 mm. wide; involucre 5-7 mm. high, the bracts about 13 or about 21, rarely only 8; brartéales inconspicuous ; rays Se mm. long; Siok glabrous. ands and fairly moist to moderately dry open places in the mountains, Arid Transition and Bor eal Zones; pire oes Yukor" British oes, and by eb =n to pe aha and New Mexico; common in “the paket mountains of Orego: n and Washington, less frequent in the Sierra Nevada. Type locality: ‘In Oregon.” A font characteristic of alkaline meadows in southeastern Oregon and northern California has been ria = 5. eer i Aivigg but appears to be morphologically indistinguishable from many of the eee 36. Senecio clevelandii Greene. Serpentine Butterweed. Fig. 5777. Senecio clevelandti Greene, Bull. Torrey Club 10: 87. 1883. Perennial with 1 or several stems from a — cau ie ex, 3-7 dm. tall, glabrous and strongly glaucous 7 oe Leaves thick and firm cely veiny, all entire, ‘the lowermost 1-2 dm long and 5- wide, with finlpiociestite or “clinic "inde equaling or more often = than : s se about 13 or it 2b: oe hies small but evident ; rays small, about 5-7 mm. long; achenes Moist places, generally on serpentine, Upper Sonoran Zone; North Coast Ranges of California, especially in Lake C cunt Tp locality : Indian aint Lake County. June_J uly. Sen elandii var. heter6phyllus Hoover, Leaflets West. Bot. 2: 132. Be . Robust plants resembling ia s. “clevela ndtt but with some me the upper cauline leaves more or less pinnatilobate, especia ae toward - hase. serpentine in the — of the Sierra Nevada in Tuolumne County, California. Type locality: ne Chinese bye ly Tuolumne Cour 37. Senecio layneae Greene. Layne’s Butterweed. Fig. 5778. Senecio layneae Greene, Bull. Torrey Club 10: 87. 1883. Senecio fastigiatus var. layneae A. Gray, Syn. Fl. N. Amer. 17: 390. 1884. Perennial with 1 or several stems from a well-developed caudex, 3-7 dm. tall, slightly tomentu- lose when young, soon essentially glabrous. Leaves rather thick and firm, scarcely veiny, dark green above, sometimes paler and glaucous oe at entire or with irregularly scattered, sharp teeth or some of the cauline ones with a Uae arrow lobes; lowermost aap well developed and more or less Sateen cm. ao wide, the oblanceolate or aehosha e elliptic blade tapering to a petiole often as lon segs alles | eae es progressively reduced and less petiolate upward, the u upper idten slightly ecpahded - the seperti or subsessile base; heads relatively large, 5-19 in an open inflorescence, the central one commonly overtopped by the lateral ones; dis 1-2 cm. wide; ifoluc re 9-12 m m. high, the bracts conse about 2 or sometimes only 13, relatively broad and a n some a them partly connate; pest few but well developed = often almost like the Pets or none; rays few, commonly 5 or 8, showy, nities mostly 1.5-2 cm.: longs achenes glabrous, 4-5 mm. long. y banks in the Bias O douglasii belt; foothills of the Sierra Nevada, Eldorado County, Cc aitettion May 38. Senecio fléttii Wiegand. Flett’s Butterweed. Fig. 5779. Senecio flettti Wiegand, Bull. Torrey Club 26: 137. 1899. na ennial fro finde tee branching, rhizomatous caudex, 0. 5-2(4) dm. tall, glabrous ohoat: or with: a litt icuous, persistent, tawny, floccose tomentum at the base and in the: leaf-axils. Basal foes wel developed and ersistent, up t a ie ¢ m. long (including the well-developed petiole) and =7 hogou mostly pinnatifid or ivpate-qnnminid the broad segments again toothed or lobulate, vary casionally simple, cordate, and merely palmately lobulate as in S. bolndcH and S. harfordi; cau leaves few and much reduced, with mostly narrower segments than the basal leaves, tem erage ne: : ral in compact cymose cluster ; Eivcluces 5-9 mm - high, gla piace the bracts about 13 or sometimes only 8; bracteoles — ext well Sager or wa sapeorss rays lt mm. long; pa glabrous. SUNFLOWER FAMILY 449 39. Senecio harférdii Greenm. Cascade Butterweed. Fig. 5780. Senecio harfordit Greenm. in Piper, Contr. U.S. Nat. Herb. 11: 597. 1906 similar to S. bolanderi but with somewhat thinner, = more evidently lobed leaves and bok Bess nate heads, the glabrous ‘alba sean 4-6 mm. high. Moist rocky woods and banks, Humid Tran sition Zone; Cascade region phe southern Washington (Skamania County) to southern Oregon, cepecistly in and oe ie a: Columbia eens ; also, less coramaly, in the Willamette Valley and Oregon Coast Ran ell back from the ocean. Type locality: Cascade Mountain s, Oregon, presumably n the Columbia Gorge. May a a RA 40, Senecio bolanderi A. Gray. Seacoast Butterweed. Fig. 5781. Senecio bolanderi A. Gray, Proc. Amer. Acad. 7: 362. 1868. Perennial from a gen ching A chia mostly 1-6 dm. tall; herbage glabrous or with a little inconspicuous, persistent, tawny, floccose tomentum at the base and in the leaf-axils. Basal me well developed and persistent rather ick and firm, petiolate, the blade commonly c orda te, ng and wide, gen i e lo : olucre 5-8 igh, the about 13 to about 21, habe neatly with few s loose, conspicuously multicellular hairs and often with a few well- developed, iiss gros Hy rays 6-12 mm. long; achenes glabrous. Bluffs, woodlands, and beaches, Humid Transition Zone; along the coast from the saprawt of the Columbia River to northern Calitoenta (ikeetiing County). Type locality : near Sadccins City, California. June-July. dii 5779. Senecio flettii a777. 5 io clevel 5778. Senecio layneae 5780. Senecio harfordii 450 COMPOSITAE 41. Senecio bréweri Davy. Brewer's Butterweed. Fig. 5782. senere abdontte eAUBee claw a: E16, var. ractus Greenm. pay Mo. Bot. Gard. 4: 31. 1917. Leafy-ste — gies with solitary stems from a short c —— mostly 4-8 dm. tall; herbage eoeiainiiy glabrous or with a little si istent tomentum aut e axils. Leaves thin, all pinnati the large ower ones “generally lyrate, with enlarged, ro terminal seg- t to 10 c ong and 8 r ir rge n e stem thus e€ i several or rather many in a corymbiform noi ping relatively large; the glabrous involucre mostly 7-11 mm. high, “i ecadl | bracts about 21 or sometimes only 13; i baarnples few and small or wanting; disk 1-1.5 cm. wide; rays el 15 mm ge achenes glabro Open or lightly wooded slopes in the valleys and foothills, Upper Sonoran same? matey 7 wate eo from Contra Costa County to Son Luis Obispo. ‘Counts and in the Tehachapi and Greenh n Mountains, Ker County. Type locality: Alameda County, California. April-Jun 42. Senecio eurycéphalus Torr. & Gray. Cut-leaved Butterweed. Fig. 5783. Senecio eurycephalus Torr. & Gray ex A. Gray, Mem. oe Acad. IT. 4: 109. 1849 Senecio austiniae Greene, Bull. Calif. Acad. 1: 93. sets sa asenng var. austiniae Jepson, Man ee m Cali ‘ae 1925. oset J. T. Howell, Leaflets West. Bot. 3: Pecaiiat with several or stems arising aa a taproot and Lateageets woody caudex, 2.5-6 dm tall ; soe aeat fae thinly tomentulose and often eventually more or less glabrate, or prasica Oy early- glabra asal or lower cauline leaves well pita ee pinnatifid or pinnately dissected to merely Sate with sharply toothed segments or rarely merely laciniate-toothed, iddle € cauli ced and becomi n corollas a — 7-10 mm. long; rays 1-2 cm. long; achenes 3.5-6.5 Peas long, ue teal Dry s, Upper Sonoran and Transition consi fasshers Cascade Mountains in Shas Butte, and Mo soot Atdel “California, south through the Inner Coast Ranges wag ie Bay. ape Mlecality: Cali- fornia. April-Jun 43. Senecio multilobatus Torr. & Gray. Basin a eos Fig. 5784. Senecio neigh Torr. & Gray ex A. Gray, Mem. Amer. Acad. II. 4: 109. 184 s Gr 95; enecto lync eene, Erythea 3: 22. 1 Senecio meleoaii [var.] wintahensis A. Nels. “ee Torrey eee 26: 484. 1899. Senecio wintahensis Greenm. Monog. Gatt. ol: 24. Senecio stygius Fr es — Bot. 2-2 ee Senecio prolivus Gr . Ann. Mo. aes . 914. Senecio souibligtiics +i var. i oe oe aty gre) Fi. - ae 1154, 1925, Perennial, sometimes rather a ort-lived, with 1 or several stems arising from a short oot which is esol surmounted by a short F ckod caudex, 1-5 dm. tall; herbage thinly oie nd often eventually glabrous or sg glabrous from the first sal leaves well developed, pin- natifid ften lyrate-pinnatifid, commonly with toothed segments, up t u cm. lo’ wide; ca leaves more less reduced but still generally fairly well developed, o ore dissected than those below and less often lyrat ds 1 nly rather numerous, 1 medium-si he disk com ‘ ; involucri igh, the bracts about 13 or rai 21; bracteoles small and inconspicuous or w ; dry Sag rhe) about 4.5-7.5 m ong; rays 5-10 mm. long or occasionally wanti ‘ie vachenes 3.5 m. long, glabrous or "hispida lous. Dry o . aces in the deserts and sokole. sometimes extending to high elevations in the drier southern mountains, Arid Transition a Sonoran outhwestern Wyoming to western Texas and ba Mexico, west t gon, northeastern Cotten Nevada, and os — south of the Sierra Nevada Type ape sea ‘on the Uintah | River | in the interior of California.” April—Jul 44. Senecio jacobaea L. Tansy Ragwort. Fig. 5785. Senecio jacobaca L. Sp. Pl. 870. 1753. Biennial or rather short-lived perennial with a poorly developed to evident taproot; stems solitary or 22% ral, erect, simple up to the inflorescence, 2-10 dm. tall; pubescence thinly floccose- tomentose but evanescent and generally nearly or quite wanting 2 "flower ing time, except fre- quently in the I scence. em ves equably gion garee mostly 2-3 _— pinnatifid, about 4-20 cm. long and 2-6 cm. wide, the lower petiolate and often deciduous, upper get sessile ; heads several or rather nu merous in a short ee sloresteaae the ‘dis k about 7-10 mm. wide; involucre beige 2 tes cake lade ie its bracts about 13, over 1 mm. wide, generally dark-tipped ; ng tne narrow rather well developed ; i co mmonly about oe mostly 4-10 mm. long; ae of the d tisk pita minutely Ske ca. those of the rays glabro nm pastures and other sete hs situations; native of Europe, now unions in rug of the United States and Bee god bee becomin ape He = seed west of the hae Sa Mo Rin n Oregon and Washington Poisonous to livestock. northern California, ee SUNFLOWER FAMILY . Senecio bolanderi . Senecio breweri 2 hal . Senecio multilobatus . Senecio jacobaea a 4 34, 4 . Hen 451 452 COMPOSITAE 45. Senecio califérnicus DC. California Butterweed. Fig. 5786. Aen oh dedwad DC. Prod. 6: 426. 1837. larior DC. loc. cit. Senec coronopus Nutt. Trans. Amer. Phil. Soc. II. 7: 413. 1841. Gre e, Bull. Calif. es ‘ee be @ Senecio calitaicuiias var, a haces Greenm. Ann. Mo. Bot. Gard. 2: 590. 1915. Taprooted annual, glabrous or 5 cies seh a 0.5-5 dm. tall, simple or branched. Leaves senile (or the ‘lower etic ate) and more or less clasping, subentire or more often co oreriatid toothed to pinnatifid or even subpinnatifid, Somnethnies broad-based as in S. mohavensis, sometim narrower and sass aed Hears S. aphanactis, 1-7 cm. long and up to 3.5 cm. wide; heads several in i itary in sm i i nflor ce 513. mm. wide; involuc m. high, the principal | bracts about 21 or sometimes only 13, usually minutely black-tipped; rays ell developed an nspicuous, aeenonly about 1 cm. long; achenes strigillose-canescent. Sandy and Pa oils, i n Zones; sand dunes along 2 ee of California from Monterey County south to nort tbern ae ower California, geet in dry, open, often sandy places farther inland (east to Kern and Tulare Counties). Type locality: California, Feb. —May 46. Senecio vulgaris L. Common Groundsel or Old Man in the Spring. Fig. 5787 Senecio enue Sp. Pl. 867. 1753. mple or strongly branched annual or winter annual with a more or less evident taproot; stem i die. tall, leafy throughout ; herbage sparsely crisp-hairy or subglabrous. Leaves coarsely and 0 i a 2-10 cm. long and “9 wide 5-8 mm. high, the principal bracts gat ghost 21, often black-tipped ; ieaetouten anes but well developed, evidently black-tipped; pap very copious, equaling or generally surpassing the corollas ; achenes strigillose- hirtelions thiefty along the a oa Seen in a ee soil and wa places; native oot the Old ioe rd but now widely distributed throughout most te the temperate zone; pe ale wast of the Cas ade Mountains and Sierra Nevada than elsewhere in our ea. Type tocatty: Barone. Flowering nearly the year gee un 47. Senecio mohavénsis A. Gray. Mohave Groundsel. Fig. 5788. S i h sis A. Gray, Syn. Fl. N. Amer. 12: 446, 1884. Taprooted annual, 1.54 dm. tall, freely branched, the herbage wholly glabrous. Leaves sessile (or the toa chaos! subpetiolate) aad cordate-clasping, ovate to broadly oblong, 2-6 cm. long, 1-4 . wi i oothed ; nde ed : mm. hig r few and small, not evidently black-tipped; pistillate flow with very much pa eet and incon- spicuous be or wanting; pappus copious; achenes still: -canescent. Desert washes and flats, Sonoran Zon nes; Panamint Mountains of California south to western Arizona and nicl. peeeary Type locality: Mojave region, California. April-May. 48. Senecio sylvaticus L. Wood Groundsel. Fig. 5789. Senecio sylvaticus L. Sp. Pl. 868. 1753. Annual weed with a more or less evident taproot ; stem about 1.5-8 dm tall, g ee ubescent sp up to the inflorescence, leafy throughout ; herbage sparsely or moderate ly pu loose hairs, scarcely or not at all glan ular. Leaves all more or less pinnatifid and irregularly toothed, commonly 2-12 cm. long - m. wide; heads s or nume t 0 ide ; involucre about 5-7 mm. high, the rather narrow, 1—2-nerved bracts mostly abou 13; bracteoles inconspicuous or wanting, not blac -tip ra y much uced incon- spicuous, less than 2 mm. long; pappus very copious, equaling or surpassing the slender disk- coro 5 ak achenes strigillose-canescen t. taps in disturbed soil and waste places; native of E urope, introduced in parts of the United States and Can n our region lene confined to the area west of the Coen — in Oregon and Washington and weer re ae Great Valley in California. Type locality: Europe. June +9. Senecio aphanactis Greene. California Groundsel. Fig. 5790. Senecio aphanactis Greene, Pittonia 1: 220, 1888. Taprooted slender annual, simple or branched, 0.5-2. f dm. tall, slightly serge in the in- florescence, otherwise essentially glabrous. Leaves sessile, small, 1-4 cm. long and 1-12 m. wide, oO. oothed or (espe he rved; bracteo g or incon ack- ; ced and inconspicuous, barely or scarcely nO the papp ; gpa: copious, shout eq lice ee slender disk-corollas : ichinies seraste ate Bose-catesce ag: - ) —— places, Upper Sonoran Zone; in the California cs Ranges and on adjacent jr Esc San Abe y to northern Lower Pe oe Type locality: Mare Island, San Praciien Bay. Feb.—March. SUNFLOWER FAMILY 453 5789. Senecio sylvaticus 790. e - ph sui eiae 5787. Senecio vulgaris 5788. S i h 50. Senecio mikanioides Otto. German Ivy. Fig. 5791. Senecio mikanioides Otto, Allg. Gartenz. 10: 168. 1842, nomen subnudum; ex st Ps op. cit. 13: 42. 1845. Glabrous perennial se Mion agit mbin a m. or more. Leaf-bla thin, somewh y-like, cordate, sharply lo apr ip ec Oe cm. ae and about as wide ; sctiotes about as long ‘lad s or pa those of the lar aves ‘A only 2 stipule-like appendages at the base; heads borne in small, condensed, pedunculate, corymbiform usters, small and few-flowered, the disk only abou ide; involucre 3— gh I bout 8; rays wanting; achenes glabrous shorter than the ek the principal bracts commorly a Along streams an il * near the coast of California from of South ARICA. gi gn lity: horticultural specimens. mistakenly thoug origin. Jan.—March. Al Bala and Marin Counties southward; sere ht to have been of probable Mexic 454 COMPOSITAE 111. CROCIDIUM Hook. FI. Bor. Amer. 1: 335. 1834. Heads radiate, the rays pistillate and often fertile, yellow. Involucre a single series of rather broad, herbaceo ous, equal bracts. Receptacle poe conic, sia Disk-flowers n basal, entire or few-toothed leaves and ne er small, long-pedunculate heads. [Name a diminutive derived from the Greek croce, loose thread or wool, referring to the persistent axillary tomentum. ] single species, of doubtful affinities. 1. Crocidium multicaule Hook. Spring Gold. Fig. 5792. Crocidium persian Hook. Fl. Bor. Amer. 1: per 1834. Crocidium eli el e St. John, Torreya 28: 74. 1928 cate, eenerall several-stemmed ue up to 1.5 or rarely 3 dm. a bearing loose tufts ra : c naked-pedunculate, solitary at the ends of the simple unbranched stems; rays 5-13, typically 8, 4-10 mm. long, fertile or sterile, Peet ali subtended by the thin and. m embranous involucr al bracts, these 3-7 mm. long; disk ‘about 1 cm. wide r les and plains, cliff ledges, and other dry open ehia at lh Na Worcs pense and Transition sons the 9 ale trough from southern Weneoucer Island to southern Oregon, extending eastward along and near the Columbia River to the oor of the Blue Mountains in Walla Walla Coane, Washington, ch oe County, ; n t i ton, and 112. PETASITES [Tourn.] Mill. Gard. Dict. abr. ed. 4. 1754. Heads radiate or discoid, subdioecious, the flowers in the female heads all or nearly all pistillate | and fertile, with or without rays, thos ose in the male heads chiefly or entirely her- aph divided or nearly so. Achenes linear, 5—10-ri ; pappus of numerous capillary sph elongating in fruit, that of the sterile flowers more or less reduced. More or less w tomentose or woolly perennial herbs See large basal leaves, merely biactedte stems (the ae apoio | and several or numerous medium-sized, purple, white, or rarely yellowish hea 1 ame from the Greek Reais. a road brim ne ae Selene to the large basal re s of about 12 species, native to the cooler parts of the northern hemisphere. Type species, Tussilago pene ay ie Petasites hybridus (L.) Gaertn., Mey. & Scherb.) Leaves evidently lobed, varying to rire few-toothed with 5-15 teeth on each side; chiefly in ~ escae region and westward (and in the California Coast Ranges). gidus. Leaves varying from merely a little wavy and Siow ion to more commonly consicwouy, Fes =e 20-45 teeth on each side; northeastern Washington northward and eastward. . P. sagittatus 1. Petasites frigidus (L.) Fries. Sweet Coltsfoot. Fig. 5793. Tussilago frigida L. Sp. a 865. 1753. Tusstlago corymbosa R. Br. Chior. Melv. 21. 1823. Nardosmia frigida Hook. FL. mi ioe 1: 307. 1833. Nardosmia corymbosa Hook. Petasites frigidus Fries, eat Veg. s acel d. 182. 1845. Nardosmia frigida var. corymbosa sa Herder, Bull. Soc. es atch Nat. Mosc, 38: 372. 1865. Petasites corymbosa Rydb. Bull. Torrey Club 37: Petasites warrenti St. — meeps Sacha St: nas Wash, ‘ 109, 1929. Petasite tig us var. ionaaiing se Ate iens 48: 123. 1946. Per asal leaves expanding with or shortly after the flowers, bier geet anon or or glbrate ab above loosely cl gaarst ew beneath, sometimes eventually glabrate, of various sizes up to 1. 5 or rarely 2 dm. wide, coarsely few-toothed or very shallowly pinnipal y Siig: a em erec s "1-5 dm. tall, with @ approximately or imbricate parellel- veined bracts most mostly 2.5~6 cm the upper yr and more pth heads several or rather numerous in a corymbiform or SUNFLOWER FAMILY 455 racemiform inflorescence, campanulate ; involucre about 5-9 mm. high; flowers whitish, the pistil- late with short rays. Typically in tundra and moist subalpine meadows and bogs, Boreal Zone; circumboreal, extending south into the high mountains of southern British Columbia and, in a form approaching var. nivalis, to the Wenatchee i ntains of Lapland. Mountains of Washington (where found at low elevations). Type loc April—June. Petasites srigitvs var. nivalis (Greene) Cronquist, Leaflets ae Bot. (? 30. 1953. (Pet nivalis apr "Pittonia 18. 1889; P. vitifolia Greene, Leaflets Bot. Obs. 1: 180. 1906; P. hyperboreus Rydb. N. Am 34: 312. 27.) Leaves Lars wey poe — and veined, seldom more than 2 dm. wide, seldom spidastbe, wider than long, the saangced seldom (though s metimes) extendi re th half way to the base; short, slender, callous teeth few or wanting. Mostly: in Comate and Alaska extending southward at high altitudes to th ympic ° ns of Washington and the Cascade Mountains of northern Oregon, and to northern Mi ntai nesota. Type locality: Mount Rainier, Washington. July—Aug. Petasites frigidus var. palmatus (Ait) Cronquist, pay 48: 124. 1946. | oe nd — Ait. Hort. ein 3: 188. 1789; Nardosmia palmata Hook. FI. Amer. 1: 308. 1833; N. —— utt. Trans. Amer. Phi oer Hog: — 1841; N. Pigida var. palmata Peden Bull. Soc. Impér. Nat ose. “st 372. 1865; Potasites plan Gray, Bot. Calif, 1: 40 SH xe ciosa Piper, Ma 2:97. 1901.) Leaves pal mately lobed card veined, | very large (u up t 4 dm. wide), tending to be broader than long, the lob monly extendi ord bs least half way to the seo wees much deeper; short, slender, callous teeth rather freely pro- duced in additi to ike larger = Stream banks, boggy ground, and moist woods, Foun: the lowlands to moderate cleveisnds in the mounta r he sa oc tal in Canada bu : a southern than the other ben gee ae Yukon, and sarang Ti south to Secu hanieiar: Michig and Minnesota; in our area occur a a e Cascade Mountains in D Wasbieeter and Oregon mie Peskendi ing south in the California Coast Rane pig ral erey County. Type locality: Newfoundland. Feb.-May 5793, Petasites frigidus 5794. Petasites sagittatus 5791. Senecio mikanioides 5792. Crocidium multicaule 456 COMPOSITAE 2. Petasites sagittatus (Banks) A. Gray. Arrowhead Coltsfoot. Fig. 5794. Tussilago sagittata Banks ex Pursh, Fl. Amer. Sept. 531. 1814. Nardosmia sagittata Hook. Fl. Bor. Amer. 1: 307. 1833. Petasites sagittatus A. Gray, Bot. Calif. . 407. 1876. Similar ar to ug Ue ds but the basal leaves merely dentate, with 20-45 wae on Nai side or even, especially i mall ce chet te: pinnipalmately veined, cordate or more commonly sagit- tate, sometimes re he 30 cm. long and 25 cm. wide; bracts of on as areal tie aan and narrower, more often with abortive blades Wet places, Boreal Zone; Alaska to Pabvatoe south to northeastern eqninate: in Okanogan County, north- ern Idaho, Montana, and Colorado. Type locality: Hudson he April-June 113. PSATHYROTES A. Gray, Smiths. Contr. 5°: 100. 1853. Heads discoid, the flowers all tubular and perfect, yellow, often turning purple in age. Involucral bracts ‘biseriate, the outer more he rbaceous and often a little shorter than the inn e€ c . Cc < tened, truncate or nearly so, minutely penicillate. Achenes hairy, mo ss turbinate, subterete or angled; pappus of relatively few and short, rather firm, capillary bristles. Strongly odorous annuals or ac cprrp ae with alternate (or all basal), petiolate, small but relatively broad leaves. [Nan poe transliteration of the Greek word for brittleness, reteerene to the pe stems oer bra A ge 4s native to the deserts is drier mountains of southwestern United States and adjacent es lig =— spe 2 ig Balbostsiis annua Nu Psathyrotes, Peucephyllum and so AG small southwestern genera have customarily been excluded from the Heliantheae (Helenieae) because of “th eir capillary pappus, and referred to the Senecioneae instead, Their relationship to the main bulk of the Senecioneae is doubtful, however, and further investigation of the affinities oO ese genera is well warranted. Outer sayernerss bracts relatively broad, more = ee oblong-obovate, the expanded green tip commonly 1.5-3 mm. wide; plants shortly = as well as 1. P. ramosissima. Outer pivblaeeat bracts relatively narrow, n elady ae commonly a little constricted at ~ Kit the — the slightly expanded green tip cpeneally 0.4-1.3 mm. wide; plants scurfy-pubescent, vidently woolly. 2 nnua, 1. Psathyrotes ramosissima (Torr.) A. Gray. Velvet Rosettes. Fig. 5795. etradymia ramosissima Torr. in Emory, —_ re wie 145. 1848. Psathyrotes ramosissima A, Gray, Proc. Am cad. 7: 363. 1868. Winter annual, or in any case te ah er and subdichotomously much branched, the stems up to about 15(20) cm. long; herbage scurfy-pubescent and eviden tly short- woolly as : es b uc hi aling or a little shite thas the i inner, relatively broad, more or less oblong-obovate, the ex- peuded green tip oe only 1.5-3 mm. wide and often turned back; corollas long- hairy above, yellow Ww, piten turning since A in age; achenes dehenny long-hairy ; pappus ulvous Dry, open, often sandy places in the deserts and desert mountains, Sonoran Zone es; Colorado and Mojave Viasiete of souther n California south to Lower California, east to southwestern Utah, western Arizona, and northern Sonora; rarely ex sterie as far north as Reno, Nevada. Type locality: “Hills bordering the Gila” [River, Ari- zona]. March— May, and sometimes annie in the fall or winter. Turtleback. 2. Psathyrotes annua (Nutt.) A. Gray. Mealy Rosettes. Fig. 5796. Bulbostylis annua Nutt. Journ. Acad. Phila. II. s 179. 1848. Psathyrotes annua A. Guay. Smiths. Contr. 5°: 100. 1853. Annual or winter annual, resembling : ramosissima, differing chiefly in the characters given in the key Dry n, often sandy or alkaline places, Upper Sonoran Zone; typically at somewhat higher elevations 0 farther jie than P. ramosissima, apparently rare in bir saadenrt southwestern Idaho to southern Utah, a oh Nevada, northwestern Arizona, and the Mojave "Sond n California, and reputedly to Sonora and Low i fornia. Type locality: ‘Rocky Mountains, near a Fe” but pease oe 9 actually taken farther west. May 114. LUINA Benth. in Hook. Ic. Pl. III. 2: 35. 1876. Heads discoid, the Pion all perfect and fertile, yellow or yellowish. Involucre a single series of rather firm, equal, sca arcely herbaceous to subherbaceous bracts. Receptacle nak thers Gorin to an entire or minutely sagittate base. Style-branches flattened, externally merely papillate or papillate-puberulent, with broad, introrsely marginal, stig- matic lines and a thickened, very short and blunt, papillate appendage. Achenes promi- nently several-nerved; pappus of numerous capillary bristles. Perennials with simple, entire to deeply cleft, ahetars leaves. [Name an anagram of /nula. The genus consists of the following 4 habitally very different but thel llied , all natives of the Pacific States and British Columbia. T Type species, a fae eyneleace Benth rece SUNFLOWER FAMILY 457 ma” Wy Ih My) 5795. Psathyrotes ramosissima 5796. Psathyrotes annua Leaves palmately cleft; heads large, the disk 12-40 mm. wide, the flowers numerous (more than ~ . L. nardosmia. Leaves Phage ot slightly toothed; heads smaller, the disk mostly about 1 cm. wide or less, the Pa less than 30 ad. Inflorescence cy a Peet icce or “> apace leaves white-tomentose on the lower surface; stem rather equably ; basal leaves wantin roa mee s petiolate, narrowly elliptic or agg ey 7-13 cm. long (petiole included), five to eleven mes as long as wide; involucre 8-10 mm, high, Bryon of a ge em, 18) pied aoa a t 15-29- (averaging shear D1-) flow eral: Grant Cou Ore Leaves all —, rather broadly elliptic or ping 2 wy eae a - ad one-half to ead aad? one- whalf times as long as wide; involucre 5-8 m pbigh, _ composed af pre: t 8-10 bracts fas na about gal averaging 13-) flowered; Cascade Mo sietiiath hypoleu pinot = gate, thyrsoid-racemiform; leaves gla ice Mas peeks 1 ana lowermost wee ones large and persistent, the middle and upper ones phar ciel a “reduced; involucral bracts mostly about 5 or 6 and - poi about 5 in each head. 4. L. stricta. 1. Luina nardésmia (A. Gray) Cronquist. Cut-leaf Luina. Fig. 5797. Cacalia nardosmia A, Gray, Proc. Amer. Acad. 7: 361. 1868. Adenostyles nardosmia A. Gray, op. cit. 8: 631 Cacaliopsis na mia A. Gray, op. cit. 19: 50. mia Cronquist, Vasc. Pl. Pacif. ae S$: 257. 1958; ood m tall, leafy chiefly toward the base, the middle a cauline leaves strongly reduced. Principal leaves Lote ek tiolate, with a broad, palmately cleft blade up t de, the lobes seldom extending beyond the le, commonly coarsely toothed r again cleft, the teeth or segment tly rather bl aves thinly tomentose e en young, ge irkee soon glabrate, more persistently but still rather thinly gray-tomentulose a heads yeloe. above 1 aa in a cory biform inflorescence, r ely large, the Foch get su an re sometim n serpentine, Upper Sonoran and_ Transition Zones; Klamath area of south- Gelere: tacuad and plrreanions ‘Califor rnia and southward rir in ine Inner Coast Ranges to Sonoma County. Type locality: near the Geysers in Sonoma County , California. April-Jun ina nardosmia var. glabrata (Piper) Crtuaiil ist, Va ase. “PI. Pacif. Nonk w. 5: 257. 1955. (Cacaliopsis nardosmia [var.] ¢ rabiaia rae Bull. i hig — 29: 222. 1902; C. nardosmia [subsp.] skate Piper, Contr. U.S. Nat. Herb. a 594. 19 . glabrata Rydb. N. Neg :. Fl. 34: 316. 1927.) Plants taller and more robust, . hi : m early eath; heads smaller and more numerous, mostly 5-18, the inflorescence — imes becoming somewhat elongate; involucre mostly 10-17 mm. high; disk 12-30 mm. wide. Meadows and open woods afone the summit _ the east side of the Cascade Mountains in Washington and page geal abst oF “the Columbia River in Ore , perhaps irregu- larly southward in the Cascade Meccntass of Ore ess commonly in the Klamath reeion ge the adjacent southern Willamette Valley. T ype fresie de Klickitat vised Washington. May—Jul A well- cea geo- graphical race but not fully Stine 2. Luinaserpentina Cronquist. Colonial Luina. Fig. 5798. Luina serpentina Cronquist, Vasc. Pl. Pacif. Northw. 5: 257. 1955. Perennial from a stout, branching, woody base, which may be wholly — and rooting or may have branches arising as much as 2 dm. fro ae ground, forming clones eral ae across; stems of the season numerous, ieee & white-tomentose 3-5 bee = tall. gti iy oats equably leafy. Leaves densely tomentose beneath, t inly so above, lar eola te or narrowly SHictic entire, acute, the middle and lower ones tapering oy an often ill- defined Salas base 1-3 cm. long, 458 COMPOSITAE the upper more or less sessile ; leaves five to eleven times as long as wide (petiole included), those near or shortly below the middle yen —— 7-13 cm. long and 1-2 cm. wide, the upper and lower gradually reduced; basal leaves - ing; heads ale in a pear Pelee ibe'y. seelpeee re: rather b ight Zz llow at first later ¢ dull the disk well surpassing t volucre, about wide; involucre white ape ise m. m. high, composed of 10-17 (chiaine 13) so : Rawes 15-29 abana: about 21) in ee head. Stee mtine slopes, Arid Transition Zone; Grant County, Oregon. Type locality: Fields Creek, 17 miles piiibeest of Dasville. Grant County. July. 3. Luina hypoléuca Benth. Little-leaf Luina. Fig. 5799. Luina a Benth. in Hook. Ic. Pl. III. 2: 36. pl. 1139. 1876. Luina hypoleuca var. eae Gray, Proc. Amer. Acad. 9: 206. 1874. Luina eet AST Rydb. N. Amer. FI. 34: 316. Perennial from a stout, branched, woody caudex, which sometimes apparently surmounts a taproot ; stems se flowering st a ier aceous, erect, poate ely leafy, 1-2-headed, 3-11 cm. high. sal and s eaves narrowed downward into their sessile base by the recurved margins of oe blade, Por by the impressed and concealed midvein, spatulate or cuneate-spatulate, 4-12 mm. long, 2-4.5 mm. wide, obtuse or rounded but appearing emarginate or even bifurcate by reflexing een an acumina : nat involucre shorter a broader, the inner phyllaries with more conspicuous, dilated, obtuse, erose, white tips, the innermost linear ; ret ie parsely puberulous or glabrous; bristles of staminate pappus s with n sehr reece if at Ao dilated t Dry ridges of the Coast rid ie Eibied Zone; Josephine and Curry Counties, Oregon, south t Del Norte and Humboldt ‘pane Ser iterets. Type locality: near Waldo, Josephine County, Oregof. eta Ph 17. Antennaria dimérpha (Nutt.) Torr. & Gray. Low Everlasting. Fig. 5849. Gnaphalium dimorphum Nutt. Trans. Amer. a Soc. II. 7: 405. 1841. peaatoner ay dimorpha Torr. & Gray, Fl. N. Amer. 2: 431 < Antennaria latisquama Piper, Bull. wenses Club pe 41. 1901, Not Greene, 1905. Began, cespitose, low perennial from a compac tly pa eetnital caudex, forming small mats, or subsericeous-tom sie 99 “throughout except on involucre, with ere ct basal leaves and erec 2 few lewtied. 1-headed stems 14(-10) cm. high or heads ee ie in he ‘bot leafy tuft. Basal 1 8-25(-40) m n bien base, ~5 mm. wi obtu cauline leaves similar, a few often reduced ones at base of i volucre ; pistillate tate Sindee turbinate, 10-15(-18) mm. high, thinly oolly at bas c, the ee uter phyllar s ovate or ovate-lanceolate, w with brownish center and acute to in € or whitish, ome to acute or subacuminate, scarious or hyaline tips; pistillate achenes with p sistent cr of pappus and with body finely puberulous, the hairs bifurcate when pee Bl ef pula of staminate pappus obscurely if at all thickened upward, barbellate and entire or some of em forking abov Dry, open bar and stony slopes, usually Arid Transition Zone; British Columbia and Alberta south through eastern Washington and eastern Oregon to California (Sitivan eal Glenn Sheypirg Modoc County + sg throug the Sierra Nevada to Alpine — nty but recurring also in Tulare County; east er Sierra hase da Mono and oii DA vec gpa sped in the Transverse Ranges of southern ae in Ven ere s Angeles, i Sen Bernardino C es), east to Cotoratio. Nebraska, Wyoming, and Montana. Type hie bet On the Black “Hits of the Platte. Se 57 pid Shek —July. id 5850. Antennaria flagellaris SUNFLOWER FAMILY 485 18. Antennaria flagellaris (A. Gray) A. Gray. Flagellate Everlasting. Fig. 5850. Antennaria dimorpha var. gener Gray in Torr. Bot. bel og oo 17: 366. 1874. Antennaria flagellaris A. Gray, Proc. Amer. Acad. 17: 212. ants in small tufts or solitary, from a slender caudex, 1-3. 5 cm. high, slender, few-leaved, 1-headed, the base of the plant emitting filiform, purplish, at first erectish and later spreading o prostrate stolons, these naked, u m. | and bearing a terminal tuft of small, subulate, ovate- d leaves enclosing a propagative bud by which the plant seemingly propagates chiefly a bienni Leaves rather closely and fabsbctcoonsaly gray-to tose, the stolons glabrate; lowermost leaves oblong or obovate-oblong, 6 long, . wide, obtuse, similar to those in the center of the -buds; inne leaves and cauline leaves narrowly linear- spatulate, 1 m. g including the peti liform base, 1- m wide: acutish, a few smaller involucre; pistillate involucre subcylindric to narrowly campanulate, 7—12(-13) broader, 4-7 mm. high, the phyllaries rane blackish brown, acum e btus pistillate yo it papillate with nonbifurcate hairs, the ring of pappus deciduous ; he ake of staminate pappus ba ha upward, secs ner slightly widened or some of them forking above D line kad’ res BA i Sonoran Zone to Canadian Zone; eastern Washington south es ccatcal and northea gg ol o and northern Wyoming. aype locality: “Between Spipen [Naches] River aa the ris fork of the et ae there bial (April-) May—Jul 124. ANAPHALIS DC. Prod. 6: 271. 1837. ntose perennial herbs with running branching rootstocks; stems equably leafy, fted. Leaves alternate, the lowest scale-like, the others lanceolate to linear, in ours) numerous, in usually small, rounded, cymose panicles, mostly short pedicelled, disciform me plants composed only of bristles stently thickened at tip, less so in hermaphrodite flowers in pistillate head as thers caudate at base, the tails acuminate, those of adinnie anthers connate. Style of heriiaphroaite eters either undivided or 2-branc , the branches linear, trun- cate, ee in pistillate flowers the va linear, obtuse, smooth, [Said to be an ancient Greek name of a plant allied to Gnaphali A genus of about 30 species, all Asiatic, a single one occurring in North America. Type species, Anaphalis nubigena DC. 1. Anaphalis margaritacea (L.) A. Gray. Pearly Everlasting. Fig. 5851. Gnaphalt argarit L. Sp. Pl. 850. 1753. t R. Br. Trans. Linn. Soc. 12: 818. Antennaria earperitns a var. ace Gray, agg Acad ‘ila 1863: 67. 1863. Anaphalis margaritacea A. oc, Amer. Acad. 8 1873. Anapha lism argaritacea var. su sbalpine A. Gray, Syn. FI oN Amer. 17: 233. 1884. entalis orn Fl. Fran. 399. 18 g. Bot. pany 12:7; om ) 1906. 906. Anaphalis sierrae Heller, Muhlenbergia 1: Fm “tTely) rennial, from slender running rootstocks, the stems usually solitary and simple, 20-80 c 1 high, ray- -tome se Leaves linear ue lance-linear or the lowest oblance oie. 5-15 cm. ne 2-20 m id e and more or less clseying: ere larger triplinerved, earl or s 2 uo sisbwatk and fab above a nd persistently ‘tomentose below heads n nded cymose panicle; involucre subg lo 5-7 m igh, y at base; shellac ‘with capes ovate and aiane papery, watkserhite tips, the hone sometimes y bigest ulo rown; achenes finely hispidu tates south ie ugh Washington and Oregon to Jgiter mene Monterey County and in the Sier evada to re County and in the San Be ino Mountains, San Bernardino County; also from Alaska east to eee and south to Peincsfewsin and to Kansas and the ky Mountain = Great Basin sige as far south as exico se nd chad rthern Arizona. Native in eastern Asia and introdu in Europe. Type locality: PY hemg ad i haneriak septentrionali, tug agnbot seen July—Oct. Var cabin. in foliage, Pata us mainly by vias fallowing ill-defined varieties: var. revoluta Suksd., balpi: ow and with rather few broad in >, glabrate above; va A. Gray, Bong oy ae pp nes Tig with the oe leaves (mostly bho ieves narrowly linear, 2-5 mm 1 these woolly above along the coa eavi found mostly in the mountains; an 3- 20 1 mm. wide) quickly pe BY ae and bright green above, found more commonly 486 COMPOSITAE 125. STYLOCLINE Nutt. Trans. Amer. Phil. Soc. II. 7: 338. 1841. Low, slender, erect or diffuse, usually much-branched, woolly ae a small, nar- row, entire, alternate leaves and small, Surered, leafy- bracted, ovoid to obose, disci- ao : orbicular, plane, hyaline tip or on this glabrous except for the pier Cintee: corol- : . : : ; A er flowers hispidulous, truncate or subtruncate. Achenes of pistillate Sats slender] obovoid or elliptic, smooth, glabrous, few-nerved, slightly compressed or obcompressed, epappose, bearing the corolla at the symmetrical summit. [Name Greek, meaning column- bed, from the form of the receptacle. ] A genus of 7 species, six from western and southwestern United States and adjacent Mexico and one from Afghanistan. Type species, Stylocline gnaphalioides Nutt. Receptacular bracts surrounding the Mapibiate flowers ineospicwous, hyaline, oblong or linear, neither qociiete nor stellately spreading in age; stam e flowers with 2-5 deciduous pappus- Siiatiea (lacking in S. psi carphoides). Marginal bract ing th istill fl b lly hyaline-winged tl gl their eA naphal lioides se ahaa > bracts enclosing the pistillate flowers with a terminal hyaline appendage, hee sasha not estas ged or with a narrow wing much shorter than the body of the bract. Hesiine —— of the outer row of pistillate bracts horizontally inflexed; plants of the San Francisco _ Bay re 2. S. amphibola. H f all the pistillate bracts erect; plants of the Great Basin and southwestern deserts. Plants. loosely: ieaante fro ‘om the toa open; leaves subtending the erences. heads attenuate at apex and u: onspicuously cusped. . §. micropoides. Plants aes mrt i from the as dense; leaves subtending the clustered het merely acute and inconspicuously if at all cuspe . S. psilocarphoides. Receptacular bracts surrounding the pamients a. conspicuous, rigidly inat rig te, prec TN spreading in age; staminate flowers epappos . S. filiginea. j 1. Stylocline gnaphalioides Nutt. Everlasting Stylocline or Nest-straw. . Fig. 5852. Stylocline gnaphalioides Nutt. Trans. Amer 1h: Soe TF: se sing 1841. ea: arizonica Coville, Proc. Biol. Soc. eagle 2.79. to diffuse, gray-wooly annual 15 cm. a or less, ride siad much-branched from base or wn a eo RZ : 0 a: wri spreading wing, the whole suborbicular-ovate, obtuse: staminate flowers w ascending, somewhat thickened bristles ; achenes Bendee straight on inner ‘Side: convex on outer somewhat compressed laterally. Dry plains and open

B ter trongly age, scarious only at the extreme tip; por wb of the pistillate flowers epappose; receptacle flat- tish, the shai bracts surrounding the fertile and few sterile hermaphrodite flowers longer than the outer bracts, scarious and less woolly, glabrous above, the achenes papillate and bearing a pappus of capillary bristles Adventive from Europe, inpaning common in overgrazed areas; southeastern British Columbia and adjacent Washington to Idaho and Montana. July—Aug. 127. PSILOCARPHUS Nutt. Trans. Amer. Phil. Soc. IT. 7: 340. 1841. wv oconse-wholly annuals, rp ae except in depauperate specimens, with narrow, and s lobed, bluntly muricate. Pistillate flowers numerous (about 20-50), closely imbricate, each loosely enclosed by and deciduous with its subtending receptacular bract, this obliquely 5859. Filago arizonica 5861. Filago californica 5860. Filago depressa 5862. Filago arv 492 COMPOSITAE bovoid in outline, somewhat compressed, saccate, reticulate, with sides meeting in the center, woolly, bearing below the rounded tip on inner side a small, ovate, horizontally tro m sd f ; es p ou receptacular bracts; corollas with slender tube and funnelform or funnelform-campanulate throat, 4—5-toot “ ovaries abortive, epappose. Anthers Sagittate-auriculate, with short cute auricles, those of ceibe anthers connate. Style-branches in hermaphrodite flowers slender, obtuse, hispidulo Name Greek, meaning naked chaff ; erongty explained by ae all as meaning deuier chaff, in allusion to the membranous pales a of about 5 species. In addition to the following, one occurs in Chile. Type species, Psilocarphus PRE Ta Ne utt., as to ya North American plants. Bracts i etapie a achenes comparatively thickly and loosely —— pov ag ~— : _ long; tomentum of whole comparatively long and loose; heads larger, 3- five Nes ™ ually — doce prostrate; leaves mostly 5-15 mm, long; ach 1. P. brevissimus. nts more or less erect; leaves mostly 12-25 mm. long; achenes a ARG ‘shine or elliptic-oblong. ‘atio Bracts sont the eres a igre Phage thinly or Sercitd woolly, about 2 mm. long; ienaidieien of meee plant mparatively close; heads smaller, about 3—5 mm. thick, usually numerous. Plants closely subsericeous ie: leaves Sale 10-22 mm. long, linear or linear- a ype bag ellipsoid-cylindric ES OF Plants — thinly grayish-woolly; leaves 4-10 mm. long, spatulate, oblong (ovate elipticovate. in the iety) ; achenes obovoid-ellipsoid. . P. tenellus. - Psilocarphus brevissimus Nutt. Round or Dwarf Woolly-heads. Fig. 5863. Psilocarphus Palomas Nutt. Trans. eee oe re a. 7: 340. <—s Sarees globiferus Nutt. loc. cit. Bert. ex DC. cy wate depicts Reany i in Gay, Fl. chil en 110. 1849; Atlas, Se ‘ “i856 A. Gray, Syn. Fl. N. Amer. ed. 2. 12: P il ph var. lied Seosees, FL W. Mid. ee see 1901. Dwarf, tals whitish-woolly annual, simple and 1-headed or with es ae branches up Leaves s spatulate to lanceolate, 5-15 mm. long, 1.5-3 m wide, pee babi : i m an he he . is oping more, their enclosing brac — mm } Ong woolly, bearing on inner “~ near the top an ovate, = rizontally inflexed or ‘ofte scarious pe ell about 0.5 mm. long; hermaphrodite flowers about 6-11; achenes subeyiieaee terete, 1.3-2 mm. long. Dri vie beds of v pools and moist ‘elke Arid Transition and Sonoran Zones; teil common in eastern Wishes on, scthees “Idaho, and western Montana and in eastern and southern Oregon —— through California to second Me: ee California and n eer o Loe California; Fal Chile and Argent Type locality: ‘Plains "s the Ore; Se a] River, in tiendaiad tracts.”” April-Jul Petit “hgpomcal brevissimus var tifldrus Cronquist, Research Sead, oar Coll. Wash. 18: 80. 1950. Plants tending to be er the eaves more prs co linear-oblong; woolly pubescen the heads rather sparse and close; wiatiione: eeics of of the att many, about 100. In Py same habitat as tee eeeetion ag Conots Alameda and Santa Clara Counties, California. Type locality: dried vernal pools, Suisun, Sol 2. Psilocarphus elatior A. Gray. Tall Woolly-heads. Fig. 5864. a oreanens var. elatior A. Gray, Proc. Amer. Acad. ef — 1873. A. Gray, Syn Fl. N. Amer. ed. 2: 12: 448 -woolly annual 15 ¢ cm. high or less, erect or sometimes diffuse, with few branches, de- ; olaints sometimes 1 cm. high. Leaves oblanceolate or the upper linear or linear-oblong, ua : . wide, acute or obtuse, c arcely narrowed to base ; heads solitary or clustered in the forks and at tips of stem and branches, mm. thick, usually about half as long as the involucrating leaves ; pistillate —— about 40-50 above , 2.6-3 mm. lon or more, the receptacular sctape ee loosely woolly especially above ng, bearing on the inner side below the tip an ovate, horizontal, introrse, scarious appen ndag: £0. 5-1 mm. long; staminate flowers , their coniiad usually reddish- -tipped; achenes fubelindcie slightly com- pressed, a mm. long. Fields and moist places, Tran m Zone; Vancouver Island, om ia. the jeues along the Pacific Slope to southern Oregon where it nao age sr ak less common in easter ashin, and adjacent Idaho. Type locality: [Portland], Oregon. Collected by Hall. May—Aug 3. Psilocarphus oregénus Nutt. Oregon Woolly-heads. Fig. 5865. Psilocarph g Nutt. Trans. Amer. Phil. Soc. IT. 7: 341. 1841. er, much-branched annual at first erectish, becom g diffuse and forming mats up to 25 cm. ac oss, closely sericeous-woolly. Leaves Pik Sor og spatulate, mostly 8— m , 1 de, gradually narrowed int tiolif base, obtuse or acutish, callotis-apiculate ; head mostly solitary in the forks and at tips of stem and bra es, 3-5 mm. thick, much surpassed by the involucrating leaves; pistillat. rs much as in P. tenellus, - a i h head, what larger, at maturity — 2-2.4 mm. long, the scarious dage rather t t app smaller, often borne near the middle of the inner face of the bract, achay hor peeetelts introrse or deflexed ; achenes more slender, pikeesd-cohaiac terete, 0.8-1. 2 mm. lon SUNFLOWER FAMILY 493 Dried beds of vernal pools and dry open places, Arid bp icay ae Sonoran Zones; eastern rat sitaieade Idaho, and eastern Oregon south in the valleys eh the Inner Coast Ranges and the cou s of the Sacramento and lower San Siiceor vad Valleys, California; also dg ad ‘an of occurring locally in Santa neaae and San Diego Counties and i n Lower California. Toei ocality: In inundated places ‘‘near the Sn [Columbia River] one outlet ‘of tie Wahlamet [Willamette 5 Soa: ree robably collected in eastern Ore according to Cronquist (Research Stud. St. Coll. Wash. 18: 85. fy pai 4 ‘Psilocarphus tenéllus Nutt. et contigs -heads. Fig. 5866. 18 Psil Nutt. Trans. Amer. Phil. Soc. II. 7: 34 Slender, Puke branched, puch floccose- a annual at first often — soon diffusely spreadin ot mats 5-30 cm. wide, the filiform sntesnages usually lon r than the leav ves. Leaves pol Se Wai oblanceolate, or ahs or som metimes nearly li : mak long wide, eee apeiatc, _sradually narrowed ” hedds Be solitary in th gs lon umpbacked, rather thinly woolly chiefly above, produced on inner below rn fem top into an ovate, horizontal, la ynen scarious appendage about . long; hermaphrodite Sacre 5-6 usually reddish above ; achenes obovoid- ee 0. 6-1. 2 _. long. Dried beds of vernal pools and dry open Fae aces, gyri <8 ao Zon in is d locali- ties, Vancouver I Pte a Br oe Columbia, ashingt Ore and w oe Tdahe ber geen ben in poe Maley Oregon and the Coast Ranges be! Caters dates Catalina Island) and northern Lower Cali- Cato also abu iat a ee n the Sacramento and San Joaquin Valleys, California. Type locality: near Santa Barbara, ali rte Ma ocarphus eaaifns var. ténuis (Eastw.) Cronquist, Research Stud. St. Coll. Wash. 18; 88. 1950, (Poilocarphs ses 7 ion Bot. Gaz. 41: 292. ed iffers from P, gers iagh var. tenellus in having the in- volucrating | a less extent those of the s aga 2 sy cil si Peake adly oblon racts aver aging’. saaller chan the name-bearing varie Be: ng; ieee | 5863. Psilocarphus brevissimus 5865. Psilocarphus oregonus 5864. Psilocarphus elatior 5866. Psilocarphus tenellus 494 COMPOSITAE 128. MICROPUS L. Sp. Pl. 927. 1753. w, floccose-woolly annuals with alternate, narrow, entire leaves and small, disciform, l-seriate or pin ag te, our species), enclose in oe as Be tetas, at u usually bifid, with slender branches. [Nam eek, is iar small foot, in allusion perhaps to its likeness on a small scale to Leontopodivon, lion’s-foo enus of about 5 species os Europe, Asia, and North America. ea species, Micropus supinus L. oe Micropus californicus Fisch. & Mey. Slender Cottonweed. Fig. 5867. cropus californicus Fisch. & Mey. Ind. Sem. Hort. Petrop. 1835: 42. 1835. Micropus erent Nutt. Trans. Amer. Phil. Soc. II. 7: 339. 1841. subvestitus A. Gray, a Calif. 12.335. 1876, Guabhaloder pats iaielp Cicene: Mie Bay Reg. Slender, erect, gray-woolly annual etion m. high, simple or erect-branched, leafy. —— nearly uniform, line ear to ogi dee olate or sivowly ate, 0.5- 2. 5 cm. lon , 0.7- wide, acute or acuminate, callous-pointed, sho rtly decurrent, enti ; heads in ba dense, ‘crite and terminal clusters, at maturity about 3 mm. high an ; ot ; involucre of about 8 green- s with short appressed w n rat y mm. lon bearing at apex of straight inner side and ‘thus below the rounded top an erectish beak about 1-1.5 mm. long, flattened and scarious for about its upper half; hermaphrodite flowers ee re sed pose, not Fameigsal by fi udheular bracts; achenes obovoid, greenish, about 1 und, Upper Sonoran Zone; ici County, Willamette ~ southwestern — wae Dry (west s the ony “Nevada) to northern Lower California. Type locality: probably Fort Ross, Sonoma County, California. April—Jun 129. EVAX Gaertn. Fruct. 2: 393. pl. 165. 1791. Small, erect, diffusely branched or acaulescent, more or less woolly annual (some Euro- ves entire, alternate along the stem or clustered beneath the 5 n a wn ae fl s prod a slender column. Pistillate flowers fertile, several to , in series on the outer part of the receptacle, each subtneded by a plane or slightly cave, scarious or chartaceous, rsistent receptacular ct. Hermaphrodite flowe rile (in e corolla fili form ; hermaphrodite corolla tubular, the limb somewhat expanded nd 4-5-den- tate. Achenes smooth or minutely papillose, more or less obcompressed; pappus absent. [Name of an Arabian chief. ] A genus of 12 or 14 species, natives of both hemispheres. Type species, Evaxr umbellata Gaertn. ndaspodigt stoned (rarely 2) in leaf-axils or at ends of branches; upper stem-leaves 4—25 mm. a5 Plan ect; heads markedly longer than broad, mostly in lea aitts . E. sparsifiora. roasts tip heads nearly as broad as long, terminal on the bekndies (axillary heads rarely & igen! acaulis. Flower-heads clustered (6-20) at apex of stem; upper stem-leaves 4.5-8.5 cm. long. * E. caulescens. L. ss sparsiflora (A. Gray) Jepson. Erect Evax. Fig. 5868. rsifilora A, Gray, Syn. Fl. N. Amer. 12; 229, 1884. anit linlicderax var. Fee Gray, loc, cit. Hesperevax sparsifiora Greene, Fl. Fran. 402. 1897. Hesperevax brevifolia Greene, loc. cit. Evax sparsifiora Jepson, FI. W. Mid. Calif. 549. 1901. Slender, erect or ascending annual 3-10 cm. high, — or usually branched at base, thinly floccose-woolly. Leaves Gate 5-25 mm. long, 1-6 mm. wide, the blade obovate or suborbi cular, ice rs sessile, solitary in the leaf-axils, not whorled-involucrate, or sometimes 2 or 3 at ti em subtended by 3-4 leaves; receptacle with convex > eee villous base bearing the pistillate ats SUNFLOWER FAMILY 495 and their bracts, prolonged in the center into a slender cylindric oo (about 1.5 mm. high about 5 receptacular bracts suttindice the inner rare pistillate flowers about t bas e outer o e 3-4 with reddish tips and without ovary, s surrounded by a circle herbaceous-coriaceous a bracts about 1. mentose on inner face, glabrous on outer ; achenes prone T) obcompressed, s ry and hillsides, Upper Aon any and Transition Zones; ae gevisncion in eat ry and Josephine Counties south to = Lui Chen, , California; alee in the foothills of nen and a sean Patlarsie. Type penlity: a cifically a ol Be ba bly from the collections made by Bre the central saneeal counties ot "Cali — April-Ju Some dwarfed spec to represent 'the variety brevtfolia as originally defined by A. Gray, appear e ibercactinte se teomany By cpabeliadina nd E. acaulis. The plants are erect but the heads tend to concentrate the apex of the shortened stem, and fa to the thortnae d a the leaves subtending th 1 ng agp cones Boy concentrated toward the apex of the stem. The leaves are usually narrower than the typical leaves sparst 5867. Micropus californicus 5868. Evax sparsiflora 2. Evax acailis (Kell.) Greene. Dwarf Evax. Fig. 5869. Stylocline acaule Kell. Proc. Calif. Acad. 7: 112. 1877. — aca — Gress ote ae 8: 257, 1883; as to name, not as to type. ax sv . Fl. N . 12: 229. 1884. Hesperevax acaulis Greene, FI. Pion: 402. 1897. e sacle bracts, reflex Dwarf woolly ae on 1 cm. high, acaulescent ba branching from the base, the branches ing, for more vigor plants. _Leaves rather densely woolly, re. n a concealing the ee of the ed bracts Trending dis stile flowers; achenes yellowish, obovoid, about 1 mm. high. Dry fields and vernal pools, Sonora - Sacramento and San Joaquin Valleys and foothills of ‘si ig hea Hier California, _ olusa Fa i i Santa Clara Type ered one from Placer County to Fresno County. ocality: Fresno, Fresno County. Collected by Eisen. Ao 3. Evax cauléscens (Benth.) A. Gray. Involucrate Evax. Fig. 5870. Psilocarphus caulescens Benth. Pl. Hartw. 319. Evax —— ens A. Hesperev vax involucrata Greene, Man. Bay Reg. 185. (es 849 aie Tore Pacif. R. Rep. 4: 101. pl. 10. 1857. x caulescens A. Gray, Proc. Amer. Acad. 7: 356. 1868. 1. N. Amer. 12: 229. 1884. Hesperevax humilis Greene, F1. Fran. 401. 1897 Caules ing beanies chiefly from the base, the cent or acaulescent, ies capo te annual 8 cm. high or less, ges or with ascend- tem when present leafy, erect or baer, wi ecumbent eads glomerate at pain rged a x of ens and Shcbea nd cons Secale sie eg involucra tem-leaves similar to the iolecvaine leaves but smaller ; aes a Niaiag the heads asain 496 COMPOSITAE spatulate, 4.5-8.5 cm. long, 5-17 mm. wide, the blade obovate to arp negn sree pte or acutish, callous- apiculate, tapering into a petioliform base ally two t imes ong, midvein scarcely costate, this dilated and indurate at base, brownish and smooth on the inner surface ; heads subcylindric eae villous at base, 8-20 (less in se eae? sere Wes rowded in terminal sinha ies 7- thick; details of structure eriey i: ly as in E. spars fons but pr (clgeatision of —— ied and thicker ; pistillate flowers few, with subtending obovate receptac vaya bracts, these concealed in the glomerules y the teak mathe: spreading receptacular bracts circling the wiih oe flowers, these with reddish tips; achenes smooth, sseonasiale obovoid, 2. Sin sa Drying are a roe Aine “Sonoran ee Butte County, Heaps south in the Sacramento Valley to n Joaquin nty i ent Inner Coast nt m Napa Cooks to Alameda County. ee locality : ey note dos i Valley. “Collected ty Hactwes: ‘April vax multicaulis DC. Pro 6. hey gear erat the ‘southwest has sia retested in various floras as growing in the Mojave ioe bien dhee Seas ders of California but no specimens have been found confirming its occurrence. An ro ast ete specimen eotected by Caden is apparently the nae of the presence of this name in the floras Tribe 8. EUPATORIEAE Achenes 5-ribbed. Pappus, at evs n part, of awns or Fue Involuc e 2-seriate, of subeq seed subherbaceous phyllar 130. Trichocoronis. Tavebiers imbricated in sev eal series, of dry and lngitinaly striped phyllaries. Achenes obi § fusiform, distinctly narrowed at apex; arnuals; leaves linear. 131. Malperia. Ache ou bi nase peiare matic or linear, not sania toward the apex; weniercess or shrubby plants; not line 132. Ho Aaneesivheg Pappus of auisied ‘bristi es 0 se 133. Eupatorium. Achenes 10-ribbed; involucral bracts striately nerved. 134. Brickellia. 130. TRICHOCORONIS A. Gray, Mem. Amer. Acad. II. 4:65. 1849. Low herbs of wet places. Stems weak, usually ascending or sometimes floating. Leaves cgi or alternate, sessile, serrate or 3- lobed, thin. Heads small, discoid, cymose-panicled, solitary. Involucre 2-seriate, subequal, of subherbaceous phyllaries. Receptacle rounded hi ighly convex, naked. Corollas with slender tube abruptly dilated into the throat, 5-toothed. Achenes prismatic, 5-angled. Pappus a minute setulose crown or of 5 short, weak awns with srernanee lacerate squamellae. Anthers subentire at base. Style-branches linear, the linear, see uispidulous appendages equaling the stigmatic portion. [Name G reek, meaning hai -apex, referring to the pappus & Peg genus of 3 species, oe the southwestern United States ‘sid Mexico. Type species, Ageratum wrightii Torr. 1. Trichocoronis wrightii sane & Gray) A. Gray. Trichocoronis. Fi sa 1 Ageratum wrightti Torr. & Gray, Proc. Amer. Acad. 1: 1848. Trichocoronis wrightii A. Gray, Mem. Amer. ae ad. IT. * ys 1849, Trichocoronis riparia Greene Besilies 1: 42. + Annual, oe or branched from base, the stems ascending to wseaincgpe 11-35 cm. long, villous ‘with lax many-celled hairs. Leaves below the ingorencence opposite, oblong or rhombic- oblong, 1-2.5 cm. long, obtuse, sessile and clasping, bluntly serrate, villous on the nerves s beneath ; heads tiny, choot tm m.h igh, cymose-panicled ; tiiohiere essentially glabrous ; corollas SuEpISh, the teeth white; a enna hispidulous above, mm. long; pappus- prtaties 5, weak, about 0.4 m long, sfcitine with minute lacerate squammelae ihc in wet ground along the Sacramento and pe Joaquin Rivers, California, and at Cienega near erside Same: also in Texas where it is native. Type locality: “‘Prope flumen Colorado Fexas: Coheceel by Wright. April—Sept. & 2 131. MALPERIA S. Wats. Proc. Amer, Acad. 24: 54. 1889. Slender = slightly pubescent, freely branched. Lowest leaves opposite, the others alternate, linear, entire, sessile. Heads discoid, rather small, numerous, loosely cymose- icled. “ota about 5-seriate, strongly graduate, the phyllaries chiefly ES to ; : : ; : 0 oblong to obovate, erose, scarious squamellae. Anthers entire at base. Style-branches linear-clavate, elongate. [Anagram formed from the name of the collector, Edward almer.] A monotypic genus of California and adjacent Mexico. 1, Malperia ténuis - Wats. oma or Brown Turbans. Fig. 5872. Malperia tenuis S. Wats. Proc. Amer. Acad. 24: 54, 1889. Hofmeisteria tenuis 1. M. Johnston, ite Calif. Acad. IV. 12: 1188. 1924. SUNFLOWER FAMILY 497 Plants tr about 25 cm. high; nye minutely and sparsely strigillose. Leaves linear or linear-lanceolate, 1. = cm. lo i 1-3 mm. wide, nag ed Soot -tipped, e ntire, revolute-margined, 1-nerved, puber hod : heads n arly cy lindr ric, 1 cm. high; phyllaries scien | puberulous ; corol- las ochroleucous or w whit ish ; achenes hispidulous on the ike 3.5 mm. long; pappus- -bristles 6 mm long, ihe squamellae 0.5 mm. lon In desert washes, Lower ponerey 4 pe! Split Mountain and Carriso Mountain in the Colorado Desert, Im- pe County, "California, to Lower California and Sonora. Type locality: near Los Angeles Bay, Lower ’Cali- 132. HOFMEISTERIA Walp. Rep. 6: 106. 1846-47. nts suffrutescent or shrubby, branching. Leaves alternate or opposite, usually fleshy, Erbe to pinnatisect, the petioles usually much longer than the blades. Heads discoid, OW , outer gee with short Harbaccors tips. saree flat, naked. Corollas all —— 5869. Evax acaulis rich 5872. Malperia tenuis 498 COMPOSITAE longer than achenes, of 2-15 bristles, sometimes chaffy-dilated at base, alee ee with scarious scales, these rae eanee dissected into bristles. Anthers subentire at bas tyle- branches linear- clavate, e te. [Named for Wilhelm Friedrich Benedict Holes of Leipzig, eat 1877, batatical morphologis and physiologist. } Ag of about 6 species, of the southwestern United States and northwestern Mexico. Type species, A BR ee Bent h. 1. Hofmeisteria pluriséta A. Gray. Arrow-leaf. Fig. 5873. fermaistere pluriseta A. Gray. in Torr. Pacif. R. Rep. 4: 97, pl. 9, 1857. ; cosa A. Nels. Bot. Gaz. 37: 263 Slender shrub, up to 80 cm. high wich tangled branches, ae pubescent, the bark white. Leaves opposite below, ‘ibvhale above ; oe 2-6 cm. lon ; blades lanceolate to ovate, acute, 0m h fe entire or usually sharply toothed, 2- long ; eads nt in pedunculate cymes or cymose panicles, rarely solitary, about 9 m se high Hie glandular-puberulous, the phyllaries dry, often purplis h-tinged, striate, at least the r with acute to acuminate, mucronate, spreading, herbaceous tips ; corolla white ; aaa ae, hispidulous ; pappus of about 12 slender bristles, sometimes paleaceous-dilated t foward base, alternating with about as many shorter bristles or Hnear-atent ate scarious squamellae n desert places, Lower Sonoran ee arene. and Utah to Arizona, southeastern California, and Lower Calloraie. Type locality : Bill Williams Fork, Ari . Oct.-May 133. EUPATORIUM L. Sp. Pl. 836. 1753. s or shrubs of varied habit. Leaves usually opposite at least tes commonly 1 petioled and toothed. Heads discoid, many-flowered, never yellow. Involucre 2- —many-~ seriate, equal or gradu as or usually dryish phyllaries. 3 oes cle flat or sapioeoe nak Corollas all sobiler: 5-toot chenes usually cylindric, 5-ribbed. Pappus of nume rous, l-seriate, rigid, capillary eines Stamens entire at base. Style-branches elongate, blunt, bas clavellate. [Named after Eupator Mithridates, King of Pontus, who is said to have the plant in medicine. } a very large genus of at least 500 species, chiefly of the —- bbe boi Besides the following, numer- ous species occur in the Untied States. Type species, Eupatorium bi Plants not conspicuously if 11 ; heads 8-16-flowered, the achenes glandular or sata Si: Leave: s chiefly eee: fowers crimson to pink. . E. occidentale. Leaves opposite; flowers white. : E. herbaceum. Plants conspicuously glandular; heads 20—30-flowered, th h glab 3. E. adenophorum. 1. Eupatorium occidentale Hook. Western Eupatorium. Fig. 5874. Eupatorium occidentale Hook. Fl. Bor. Amer. 1: 305. 1833. Kyrstenia occidentalis Greene, Leaflets Bot. Obs. 1:9. 1903. Many- ce oa perennial from a woody 25-80 cm. — the nee re finely in- curved-puberulous. Leaves chiefly alternate; petiole s 3-15 mm ; blades ovate, 1.8-6 cm. long usually acute, at ase € : slightly puberulous beneath ; heads Bf flowered, about 8 mm. high, cr owded in small cymose panicles terminating stem and branches; involucre 2-seriate, grey 3-4 mm. high, the phyllaries linear, acute or acuminate, puberulous ; corollas crimson, pale purple or white ; achenes 3 mm. long, sessile-glan dular. cam bank and among rocks, Transition and Canadian Zones; Idaho to Utah, western Nevada_and Was shington 20 south . the mountains of northern Calituriits and in the Sierra Nevada to Tulare County. Type ete. “On the low hills between the north and south branch of Lewis and Clarke’s River,’’ northern Idaho. 2. Eupatorium herbaceum (A. eked, Greene. Desert Eupatorium. Fig. 587 ? herbaceum A. Gray, ae Contr. 5°: 74. 1853. Eupator ncciidentals var. arizonicum A. Gray, Syn. Fl. N. Amer. 12: 101. 1884. Enpetorivs herbaceum Greene, a atl 1901 Eupatorium arizonicum Greene, ai Opposite-leaved perene, ial, we dm. high, from a woody root, herbaceous or somewhat suf- frutescent at base, the pee ¢ minutely scaberulous, especially on the upper part. Leaves glabrous or somewhat poets neath, 1-4.5 cm. long, the petiole 4-11 mm. long, the blade rather firm, veins cereatnieat on older leaves, deltoid to -g hdelggat- ves acute at apex, truncate or subcordate at , the margin crenate om at the de inflorescence of short-pedunculate d Aghary terminal or terminating the upper branchlets "heeds -16-flowered, white, about 5 m igh, the peduncles es aenichy caberulous ; phyllaries Bd Lip ual in length, ‘coeiaiete: more or less — 3.5-4 mm. lo sped ewes, hispidu lou the Be Canyon wana a asi on desert mountains, Upper Sa: moran Zone; Colorado and Utah south to New Me fe ar Ppt ede aad tee the New York haniiatns ahowg 28 and Clark Mountain : tee s), San Bernardino County, California. Type locality: Santa Cruz, Sonora. Collecte y Wright. June— SUNFLOWER FAMILY 499 3. Eupatorium adenophorum Spreng. Sticky Eupatorium. Fig. 5876. smepteriows glandulosum H.B.K. Nov. Gen. & a 4: 122. pl. 346. 1820. Not E. glandulosum Michx. 1803. 826. Eupatorium pasadenense Parish, Zoe 5: 75 nace opposite-leaved perennial with glandular puberulent stems 3-10 = . high. Leaf- blades 3-7 cm. long, slightly longer than the petiole, oh deltoid, iy acute at apex, promi- nently 3-veined at the base and truncate or somewhat cuneate at the base, the leaf- Boa wa crenate- dentate from the widest portion of the “pect to ihe nol Siaonceinn ce of compact cymes on dichotomous peduncles longer than the leaves; heads 20-30-flowered, white, about 5 mm. long: the i ava Teachers glandular ; phyllaries about 4 mm. long, lanceolate, phe AG puberulent ; achenes glabrous. Moist shrubby slopes, frequently escaping from cultivation but not abundant in the Coast Ranges from Marin and Alameda bapa pee adh Angeles County, California; also introduced in the Hawaiian Islands. Type local- pril Eupatorium maculatum bi pg (A. Gray) Breitung, ay Bly “ase 61:98. 1947. (Eupatorium —- Gray, Syn. Fl. N. Am : 96. 1884.) Tall perennial up to 20 dm. high, differing from E. maculatum in the sharply serrate, more ion ct densely covered beneath with “Aik neceotind. curly hairs. Within our range, yore he al only in swamps at Sumas, Whatcom County, Washington, but occurring in British Columbia south to Utah and eastward. Type locality: Fort Collins, Colorado. Joe-pye Weed. 5875 AV te 5873. Hofmeisteria pluriseta 5875, Eupatorium herba 5874. Eupatori idental 5876. Eupatorium ecaelicies 500 COMPOSITAE 134. BRICKELLIA Ell. Bot. S. C. & Ga. 2: 290. 1823. erbs or shrubs. Leaves opposite or alternate, sessile or petioled. Heads discoid, 3- oa a small to large, solitary to panicled, white, o saree or purplish. ‘Tne volucre several-seriate, strongly graduate, the phyllaries ay, conspicuously striate, the outer herbaceous-tipped. Receptacle flattish, naked. Corollas all me tubular, 5-toothed. Achenes cylindric-prismatic, 10-ribbed. Pappus of numerous ce pao: & ristles, pert white. Anthers rounded at base Style-branches linear-clavat Nam 2 ea oO Brickell, naturalist and physician, a contemporary of i uhtenteee and “Elio A genus of 93 species, dl = of the Americas. Besides those here described. others occur in southwestern and southeastern United Stat Type species, Brickellia cordifolia El). Heads 3—7-flowered, very small Leaves linear or nade linear-lanceolate, 1-10 mm. wide. 1. B. longifolka. Leaves — Lea entire or afi —* — glabrous. 2. B. multiflora. tana ves serrate; phyllaries puberulous a gs 3. B. knappiana. Heads 8— Coie Sereret ‘eaeeatly. sade ium or ae Leaves mostly tt ‘a mm. long or less; intricately branched sonic . B. frutescens. — usually dt , larger, sometimes elliptic and entire or suben Leaves white-tomentose; ied large (2 cm. high or more), solitary, naked- psc AS 5. B. incana. Leaves green, or else — much smaller and on leafy pedun Leaves firmly coriaceous, spinescent-toothed; heads poo naked-pedun cled. Outer sivaA ee broadly ovate-oblong; peduncles glabrate to finely glandular. a 6. B. atractyloides. Outer phyllaries ovate-lanceolate; peduncles with dense, short, spreading pubescence. rie B. arguta. Leaves — ony coriaceous ‘Ror spinescent toothed; heads clustered or leafy-peduncled Outer Harie te tips. 8. B. grandiflora. Outer pecans: S not v tt te tips ads ocean clustered in gp axillar 1 Petioles 5-20 mm. long; leaf-blades 1—5 cm. Le not cinerous-puberulous. So. B. rs i Petioles 1-3 mm. long; leaf-blades 3—14 mm. long, usually cinereous-puberulou 10:..B. eriave een. Heads otherwise. Stem and leaves ig -lanat 11. B. nevinii. Stem and a ee rary Heads Skate. bracted. ; eaves a ally distinctly toothed; he ady numerous, solitary, or in clusters uf 7 at tips of divergent bracteate branches Stem and pedicels pilose with wide- apis gland- tipped hairs. B. microphylla. Stem and pedicels lanulose with mostly eglandular hairs 13 Leaves mostly elliptic to oval, entire; stems reap Be or penta hana: heads 1 to many, cymo osely arranged HW a 4 1 rN * ih ecal p y leafy-bracted at base. “14, = ee 15 1. Brickellia ee S. Wats. Willow-leaved Brickellia. Fig. 5877. . Amer. Nat. 7: 3 1873. — Shae ae rae Rev. Gen. Pl. * 328. 1891, Much branched shrub, the stem glabrous, viscid, white-barked. Leaves alternate, the blades linear or narrowly linear-lanceolate, 2-13 cm. long, mm. wide, pe entire or obs sae denticulate, gland-dotted, short-petioled; heads abaek: 8 mm. high, 3-5-flowered, in sma sin shor - peduncled, axillary and terminal, umbelliform cymules, forming a ‘lea fy thyr or involucr high, the phyllarie: € 10-12, ovate to linear- aa obtuse, dry, Dies eiteia: corollas Thine achenes hispidulou Rare on canyon ae and in washes, Sonoran eam southern Utah to northern Arizona and Nevada, and Inyo County, California. Type locality: southern Nev . Au ug.—Oct. 2. Brickellia multifldra Kell. Gum-leaved Brickellia. Fig. 5878. Brickellia multifiora Kell. Proc. Calif. a 49. 1877. Coleosanthus multiflorus Kuntze, Rey. Gen. Pl. 1: 328. 1891. Branching shrub 5-9 apy ce more a less —s. the stem white, baler ag Leaves ogg os blades pre eeabee the per narrowly lanceolate, 3.5-8.7 cm. long, 1.2-2.8 ¢ €, cuminate, entire or obscurely Slentic slate, SL oh Shacks lous, es ee dot ted; phyllaries " Seeat i 20; heads ic nflorescence much as in B. longifolia, the leaves tanec the flowering branchlets short ¥ ommon yon slopes and in gravelly or sandy washes, Sonoran Zones; southern Sierra Nevada, California, mostly on the eastern slopes Sitwnet in the eis ranges to adjacent Nevada. Type locality: Kings Canyon, She Nevada, Fresno County, California. Aug.—Oct SUNFLOWER FAMILY 501 3. Brickellia sities Drew. Knapp’s Brickellia. Fig. 5879. Brickellia knappiana Drew, Pittonia 1: 260. Coleosanthus saat Greene, "Exythea 1: fe 1893. Branching shrub, 0.4-2.4 m. high, viscid, somewhat ee, gi stem white. Leaves alternate fue shades broadly iscealate to ovate, cm. , 1- m. wide, usually some- serrate, gland-dotted; heads 7 mm. hi 7: Acuered, re - very leafy panicle ; phyllaries m igh, pabe alos especially tow ard apex ; achenes hispidulous. e, along streams, Sonoran Zones; Pana t Mountains, i ne County, and San Bernardino County. Type PE ag near the Mojave River, San Bo aaetine: C plo Califor 4. Brickellia frutéscens A. Gray. Shrubby Brickellia or Brickellbush. Fig. 5880. , abana A. Gray, Proc. Amer. Acad. 17: 207. 1882. Coleosanthus frutescens Kuntze, Rev. Gen. Pl. 1: 328. 1891. matic, intricately seh eid low shrub with spinescent branches and whitish Se the Asi and tacts griseous-tomentellous, sometimes scabrid. Primary leaves (rarely seen) up to 4 long, 1 r , remotely capettanesieath ulate, the others eerie oval, or perry , 3-14 long, obtuse, entire, sessile or short-petioled; heads 1.3-1. sity flowered, solit at the tips of rather short, amar bracteate peduncles ; involuc e 9-12 mm. high, the setblaada chiefly linear, obtuse or acutish, ous-puberulous, rise uter with ak sometimes squarrose herbaceous tips; corollas purple; lente hi ia lou In desert places ag among rocks, Lower Sonoran Zon edge mountains in California southward on the P nage pd amare ig ee, ‘of th ne San Pedro Martir and Esmeralda County Citoane?), Nevada. Type locality: not definitely stated. April—Oct f the Colorado ey yo ay, adjacent r California, 5879. Brickellia knappiana 5880. Brickellia frutescens 5877. Brickellia longifolia 5878. Brickellia multiflora 502 COMPOSITAE 5. Brickellia incana A. Gray. Woolly Brickellia or Brickellbush. Fig. 5881. Brickellia incana A. Gray, Proc. Amer. os 7: 350. 1868. Coleosanthus incanus ae Rev, Gen. Pl. 1: 328. 1891. Much branched shrub, 3-6 dm. high, white- pes on Ae yes Fone ak ane young branches, the stem glabrate, white-barked. Leaves alternate, ovate, 1-2 cm. long, ac t os sessile, serrulate to entire, glabrate at length; heads mostly ie at tips one stem pe bra 2-2. cm. high, about 60- ‘ eee. Breck yee involucre 1.7-2.2 cm. high, the Siehees ie ovate to linear, obtuse to acuminate, ait ulate; corollas ago whitish or ochroleucous ; achenes densely a dls hispidu lou shes and canyon bottoms in the a ert, Sonoran Zones; midi pndag California, in the Death Valley south through oe. Mo} ave Danesh to Riverside County; also s vada and western Arizona. Type = nc ha eid Mountains, San Bernardino County, California Mas 6. eee atractyloides A. Gray. Spear-leaved Brickellia. Fig. 5882. Brickell tyloides A. Gray, Proc. Amer. Acad. 8: 290. 1870. s Kuntze, Rev. Gen. Pl. 1: 328. 1891. Coleosanthus webilesak: Nels. Bot. Gaz. 37: 262. 1904. Low, leafy, branching shrubs with white- barked, rather stiff branches and flexuous es branchlets. Leaves a ternate, short-petioled or subsessile, the blades 1-2.5 cm. lon ng, ovate narrowly acute to acuminate at the spinescent apex, spinescent-toothed on the margin, écHaviinas i ] i icu ; heads a i bright green and gla , conspicuously reticulate d t 50 er several, solitary on the s of the —- ts, 13-16 ; peduncles naked, glabrate or finely glandular below the heads; the o r phyllaries broadly ovate- cies , resembling t eaves in texture, 14 loose, entire; the dite er "phyllines finder. acuminate, stramineous ; corolla white or yellowish; achenes hispidu lou Rocky hillsides ae slopes, sepa Upper Sonoran Zone; southern Nevada and Utah to eastern Inyo and San Bernartinn Counties, Californ and Yuma County, Arizona. Type locality: “Utah, near the Rio Colorado.” Collected by Palmer. March—Sep 7. Brickellia eu Robinson. copes Spear-leaved Brickellia. Fig. 5883. Brickellia arguta Robinson, Mem. Gra — Herb. 1: 17; mbar ‘atractyloides w we hells n, Man i i Cali, 1015. 1925, Blak a ieiticy Gants U.S. Nat. Herb. 23: 1483. 1926. Low, branchin pros closely resembling B. atractyloides in general appearance, densely spreading- bixtellous with mostly gland-tipped ie n the leaves, involucre, and young bra sche the stem glabrate, white-bar ae eaves alternate, tsa the blades ovate, 1-2 a : Mh : : e veins om flowered, solitary at tips of branches, the naked e densely glandular- puberulen nt with spreading hairs; involucre 1-1.3 cm. high, the Gilteriioat” phyllaries usually rigid-herbaceous, more or less puberulent with spreading hairs ercenialy at the base, lanceolate to ovate, rather loose, entire or bist se with 1 <2 s mall teeth ; the inner phyllaries linear or lance-linear, acuminate ; corol- idu mr s. On desert slopes and washes, Lower Sonoran Zone; western ey a ox: Colorado Deserts, California, from the Death Valley fasion in Inyo Co ounty ebathe % northern Lower Californ Type locality: not stated. May—June. Intergrading for ms are to be found on the eastern limits of Gisteitution which show strongly the close rela- = ~ — S$ species with B. atractyloides Bri arguta var. odontélepis Taine (Mem. Gray Hes 917) differs from B. arguta var. arguta in 4 pen phyllarie petted are broad 7 conspicuously oi ek Valley tsb 8 ole ideaty, Cali- season Eoin: 2aar, 37 31). ‘ied southern Colorado Desert, Teter County, cing to the San Felipe Desert inn eastern Lower California. Type locality: gf ie Desert. Collected by Ore 8. Brickellia grandiflora Nutt. Large-flowered Brickellia. F ig. 5884. Eupatorium? grandifiorum Hook. Fl. Bor. Amer. 2: 26. 1834. Brickellia grandiflora Nutt. Trans. Amer. Phil. Soc. II. 7: 287. dhgasian erent fons var. —— ris A. Gray, Proc. Amer. ia 7. er 1882. Kuntze, Rev. Gen. Pl. 1: 328 a ueaakie practiipes Greene, Pita rs ma? Herbaceous perennial 2-7 dm. high, from long, eh maski. fusiform roots, the stem simple or branched, incurved- -puberu lous. Leaves chiefly alternate; petioles 1-7 cm. long; blades g scab flowered, nodding or erect, mbelliform clusters at tips stem and branches; involucre 8-11 mm. high, the outer phsllaries with eennnty BA ee tips, the inner linear, acute; corollas whitish; achenes hispidulou: In canyons and o: n cliffs, Tra i Zon eon linha and er ce e Nebraska, Missouri, and Arkansas south to New Mesiba, Arizona, the Sierra Mevula we hp jouecegeed Ryo yp wer California. Ty Pompe “On the low hills between the north and south branches ms ,” northern Idaho. July 9. Brickellia californica A. — California Brickellia or Brickellbush. Fig. 5885. Bulbostylis californica Torr. & Gray ae on 1841. Brickellia californica A. Gray, Mem. Amer. Acad. II. 4: 64. 1849. SUNFLOWER FAMILY 503 Brickelli htit A. Gray, Smiths. Contr. i" 72::.1853. Coleosanthus pc ace See fet Rev. Gen. Pl. 1: 328. gate Coleosanthus albicaulis Rydb. Bull. Torrey ‘Clab 31: 646. 190 Brickellia californica var.? jepsonii Robinson, Mem. Gray 4. 227k 2917. Branching shrub 5-10 dm. high, the stem tomentellous to scabrid-puberulous, whitish ac hoeoag 1 Leaves chiefly alternate; petioles 5-20 mm. long; blades usually deltoid-ova eri —5 cm. long, obtuse or acute, truncate to subcordate at base, coarsely crenate or crenate-serrate, firm, cr eer puberulous, gland-dotted beneath, reticulate; he -10 mm. high, 8- ¥ flowered poets clus- tered in axillary and terminal cymules, forming leafy thyrses ; involucre a the phyllaries dry, obtuse to acutish, ovate to linear; corollas aenroleuccie posal a corelisir: achenes hispidulous. Dry and streambeds, Sonoran Zones; poeernine baton? Creat southward to Lower California and éaatened tee aa desert ranges to Colorado and New adjacent Texas, and Chihuahua. Type locality: California. July—Oct. 5881. Brickellia incana 5883. Brickellia arguta 5882. Brickellia atractyloides 5884. Brickellia grandiflora 504 COMPOSITAE 10. Brickellia desert6rum Coville. Desert Brickellia or Brickellbush. Fig. 58 sabeceaiae ay deanehases Coville, Proc. Biol. Soc. 2368; Coville, Contr. U.S. Nat. Herb. 4: Brickellia californica var. desertorum Parish ex " M. Hall, sate ge Pub. Bot:3: 33.1907, uch branched shrub 1 m. high, the stem and branches slender, white-barked, whitish- tomentellous glabrate. Leaves opposite or alternate petioles sr mm. long; blades ovate, m. long, obtuse, Egg i or subtruncate t base, crenate-serrate, saat ae eos ke or gree ere heads 7-9 mm. high, 8-12-flowered, mall axilla ary and terminal glomerules; in- volucre abo ut 7 mm. high, the phyllaries ovate rth ‘aa obtuse to auc cy seer sin achenes ou ocky sts opes of desert ranges, Sonoran Zones; western Riverside County, California, eastward to southern Heras and anethweatern shane open Type locality: between Banning and Seven Palms on the Southern Pacific railroad, California. Nov.—March. 11. Brickellia nevinii A. Gray. Nevin’s Brickellia. Fig. 5887. Brickellia nevinit A. Gray, Proc. Amer. oar 20: 297. 1885. Coleosanthus nevinti Bees Cat. N. Amer. Pl. 8. 1898. Suffrutescent, oe the stems ae branching, gray-lanate, glabrescent. Leaves alternate; petioles 21 mm. long or less; blades ovate, 5-16 mm. ong, acute or obtuse Pa es ow gray. ish lanate, eat glabrescent ; heads 1.2-1.5 cm. high, .. ee ‘flow red, 1-3 a s of usually minutely leafy ec wana age 8-12 “hi r less accel sl the phyllaries ovate or lanceolate to linear, e to acuminate, the sate “with spreading tips; corollas whitish or purplish; panna ietty hispidulow In dry places, ices Sonoran Zone; coastal slope i ‘the mountains of southern California from western Kern County to Ventura and Los Angeles Counties. Type locality: near Newhall, California. May—Oct 12. Brickellia microphylla A. Gray. Little-leaved Brickellia or Brickellbush. Fig. 5888. Bulbost phyll. Pet Trans. Amer. Phil. Soc. II. 7: 286. 1840. Brickellia wuicveia A. Gray, Smiths. Contr. 35: 85. 1852. Brickellia cedrosensis a SS , Bull. Torrey Club 10: 86. 1883. Calesinatines microphyllus Katies Rev. Gen. Pl. 1: 328. 1891. Shrubby, m. high, much branched, glandular-pilose on stem and leaves, the bark whitish or grayish. tas aves ae rnate, _—- petioles 1-5 mm. long, — broadly ovate, 1-2 cm. long, eads 1-1.4 ¢ acute or obtuse, coarsely few-toothed t Zepto entire, veiny; h m. high, about 22- flowered, in groups of 1-7 at tips mat the divergent, minutel y 3 fy branches; invoiuere about 1 cm. high, the a ovate to linear, ee ac acuminate, the r with squarrose herbaceous tips; corollas ochr oleucous or purplish; Se sel i oeaioiow: ahoees rae m. long. In rocky gos tikes Sonoran and Transition zones eastern Oregon in he Ag 9 ee oienned to Idaho and “eects ied . rags - re ee and in California on the eastern face of the Sierra Nevada from Sierra County to Tulare C and i hite and Inyo Secntarie mit y San Gabriel F pecbartl Los Angeles County; also on Cedros isa ‘Cen Calton Type locality: Blue Mountains of Oregon. Aug.—Sept. Foliage said to be pineapple-scented. 13. Brickellia watsénii Robinson. Sweet Brickellia or Brickellbush. Fig. 5889. Brickellia watsonit Robinson, Mem. Gray Herb. 1: 42. ep 19. 1917. Coleosanthus watsonti Rydb. Fl. Rocky Mts. 843. 1917 Similar spreading, Sand: tipped hairs. Leaves not peianttior y veiny; heads 9- ms mm. high, about 18-flowered ; involuc € 7-8 m pk eh; cachenes hispidulous, about 3.5 mm. lon San not stated. Aug.— Brickellia a. (A. Gray) A. Nels. in Sgt & Nels. New Man. Bot. Rocky Mts. 909, in synonymy as ae sili micr eehyle < var. scabra A. Gray, Proc. Amer. Acad. 11: 74. 1876.) + itoring ate = _ prec ving the stems and peduncles with a short, glandular, ba tg an pubes pected in pag Inyo County, California, as it has been collected in adjacent Esmeralda and Nye Ccuniies. ev 14. Brickellia oblongifélia Nutt. Narrow-leaved Brickellia. Fig. 5890. Brickellia oblongifolia Nutt. Trans. Amer. Phil. Soc. II. 7: 288. 1840. Coleosant master ea Kuntze, Rev. Gen. Pl. 1; 328. 1891. Suffrutesc t base, 1-5 dm. high, the stems numerous, erect, whitish, hispidulous and glandular, Sucaiti Seats below the rence Leaves alternate, elliptic to oval, —4 cm. lon acute or apiculate to obtuse, subsessile, entire or rarely geen ing pubescent like neg ‘stem ; heads 1 to ex 0 b .4-1.8 cm. high, 40-50 » “"g cymosely arranged at f stem anch oe, 50- owered ; involucre about 1.5 cm. high, the ae lance-oblong to ; linear, acute to —. glandular-puberulous, sein green and white; rollas purplish or ect atric achene pot, etime with a sie eglandular 3 In sandy or gravelly anes Sonoran and Transition Zones; British Columbia to Oregon, —— and Utah. Type locality: gravelly Sesion th g the Willam =. June— SUNFLOWER FAMILY 5885. Brickellia californica 5886. Brickellia desertorum 5887. Brickellia nevinii 5888. Brickellia microphylla 5889. Brickellia watsonii 5890. Brickellia oblongifolia 506 COMPOSITAE Brickellia Narotnay tories var. riggs (D. C. Eaton) Robinson, Mem. Gray Fas rb. i: 1917. Apr tea linifolia D. C. Eaton, Bot. King Ex : 137. pl. 15, figs. 1-6. 1871; B. mohavensis A. re Syn N. 12: 104. 1884; Coleosanthus vlinifolins Kuntze, Rev. Gen. Pl. 1: 328. 1891; C. -gblonoifolins ‘inifolins oo Eaton) Blake, Constr: U.S. erb. 25: 534. 1925.) ‘Achenes —— hispidulous, In more e desert places, Upper cs Si Zone; Colorado to New Mex rthern Ari Ms gh ey th . Death Valley region of Inyo County, California, ‘south to eastern San gee nl Danese "Wee lohan American Fork, Jordan Valley, Utah. 15. Brickellia gréenei A. Gray. Greene’s Brickellia. Fig. 5891. Brickellia greeneit A. Gray, Proc. Amer. Acad. 58. et Coleosanthus greenei Rontek Rev. Gen. Pl. 1: 189 Stems numerous from a woody —— erect, 2.5-4.5 dm. high, simple or corymbosely branched above, spreading-pilose with short and long gland- pga pony throughout, the herbage with a strong sweetish odor. Leaves Fase petioles 4 mm. lon ; blades ovate, 1-3 cm long, ie pride serrate to entire, glan Sau “barn wy igh, 60-flowered, solitary a sofs and branches, involucrate-bracted by the uppermost leaves; proper phyl- laries nears alee i acuminate, essentially glabrous; corollas ochroleucous ; ad Ba hispidu- lous above velly ri rocky mountain sides, Transition Zone; Josephine County, Oregon, south through the Si ake iyou face on to Ghee = and caleert Humboldt Counties, Californta. Type locality: south branch of Scott River, California. July—Aug Tribe 9. CYNAREAE* Heads 1-flowered, numerous, aggregated into dense, a capitate clusters. 135. Echinops. Heads few- to many-flowered, not forming spherical cluste Rea of achenes (‘“‘hilum’’) pase pappus Lsehingie , deciduous (or the inner series of bristles more or less persistent in Saussurea) ; s homogam e flowers alike and perfect Leaves neither spinose nor ae "obed, the — — thistle-like; pappus- Acistles free at least in the outer series ig linear or linear-lanceolate, the outer and middle tipped with an uncinate py Pah Sala ose, 136. Phyllaties aa ovate to lanceolate, acute, unarmed; receptacle naked or ees TH pe ait Leaves usually spinose and paenly a to pinnatifid, the plants pore thistle-like; pappus- aie united at the base into a low r Receptacle deeply tess the si setae; bordered, not densely setose. 139. Onopordum. 2 aa Ganeckt tulose, not plu i: hekees mottled with Sih along eae veins; iighytipeies with a we et pinnately spinose appendage. . Silybum. Leaves not white-mottled; phyllaries entire, gradually narrowed to a spinose apex 143. Carduus. esd Hin plumos Leaves very farkee Sicieda large, the phyllaries numerous, broad, imbricate; —— fleshy. 141. ynar Leaves usually small or medium; heads small to large, the phyllaries usually narrower, frequently sprea ting: prs te not fleshy. 142. Cirsium caaamagr oe: cal of achenes lateral; pappus, when present, consisting of free bristles or paleae, setsutent or Phyllaries with a foliaceous blade, the Liles hee and leaves usually spinose or sotenioee Comets 2 heads thick. 140 t Phyllaries not foligceous, either F spinose or bhai heads ee rie sea — the outermost flowers sterile, or s the head homogamous; achenes terete o mpresse Achenes compressed, ak or rugulose. not = toothed at apex; pappus eset or lacking, not in series of 10; leaves not spinose or bristly. 144, Voumatie Achenes terete, 20- Pibed. 10-toothed at apex; pappus in 2 series of 10 bristles med pot eplanione. 14 135. ECHINOPS L. Sp. Pl. 814. 1753. Annual, biennial, or perennial, arachnoid-tomentose herbs with more or less spiny leaves. Leaves pinnately divided with the larger divisions again once- or twice-lobed or -parted, the uppe ile by road clasping base, the lower with blades narrowly de- current along a petiole-like base. Heads 1- ered, numerous, aggreg t m a com- pact, spherical, itate inflorescence that is terminal on le-like stems, the heads deciduous at maturity. Phyllaries very numerous in many series attac to an elongate, stipe-like receptacle, the lowest phyllaries bristle-like le-like, chartaceous, pass! gradually into broader, herbaceous-tipped p ie the mi ppermost phyllaries attenuate-spinose, pectinate-fimbriate with margina istles. Corolla deeply 5-parted, dages. St tyle hairy at the base of the branches. Achenes elongate, little-com pressed, hairy, remaining attached to the receptacle and falling eload by the husk- like involucre. Pappus paleaceous, forming a low crown on the top of the achene, the paleae * Text of Cynareac contributed hy John Thomas Howell. SUNFLOWER FAMILY 507 hairy or barbellate- oes coe or united to form a more or less erose membrane. [From the Greek, meaning urchin e-] An Old World genus 7 stad 70 (or 12 ms ording to Lemée) species, most common in the eastern Medi- terranean region. Type species, Echinops siatincebelin se 1. Echinops ruthénicus M. Bieb. Ruthenian Globe Thistle. Fig. 5892. Echinops ruthenicus M. Bieb. Fl. Taur. Caucas. 3: 597. 1819. Echinops ritro var. ruliicns Halacsy, Consp. Fl. Graecae 2: 91. 1902. erennial herb with erect stem 1-1.5 m. tall, simple below and few- gorse be =. — clothed with arachnoid tomentum that ma more or less deciduous especi the or ate a weak or strong, spinose tip, the lower leaves with a ‘narrowly rdered_ petiole-like “es he middle sessile with a more or less clasping base, the er leaves — ba ler, lanceolate to ovate- pias re plat divide d or serrate-lobed ; infloresce ence 3-4 cm. in diameter, the heads in flower 1.5-2 cm. long, the lowest br ate like phylla ries white and aiaaing the middle and iritier tat shvTlaties narrowly lanceolate with elongate attenuate tips, the outside lavender-tinged, scabrous-roughened on exposed parts, not junta the inside brownish-tinged; corolla-lobes lavender rope Baeaee ene, 7 mm. long; filaments short, violet, anther-tube conspicuous, 6 mm. long; a e oblongish, a little constricted — mr middle, 6 mm. long, thinly covered with Soeeniaed a igece nding, tog a tb mie d D is Lg -paleae united to form a shallowly lobed, membranous c n 1 mm. lon e lobes barbell lat Establi aie in weedy vice in Whit man Dae Washington. Introuce from pegtein Aug.-Se Ech atu ratzka, pede Zool. Bot. Ges. Wie 17. 1858. A plant collected in 1954 in Cowlitz County, Pay abhingsoet) are ars to be = ogi Enwobeaa: uaa thistle. It may be distinguished from E. ruthenicus =. Bieb. by the silvery wp ese t ray-green middle and gy ok phyllaries, esd the fewer sordid, bristle-like phyllaries and by the whitish pons reg Perhaps only a fu vei e from cultivat Echin sphaerocéphalus L. Sp. Pl. 814. 1753. Differing from E. henicus and E. comitatus in the glandular pubescence on She exposed hacks of vo middle and epee ohipilaries, in the glandular hairs pro- tru pine § from e arachnoid tomentum on the stems and lower leaf-surfaces, and in the denser me -scabridous pubescence on es upper leaf-surfaces. Locally vataldished 3 in Modoc co unty, California, east of Tulelake, Siskiyou | panes aetna in Europe and Asia. 136. ARCTIUM L. Sp. Pl. 816. 1753. Large biennial herbs with scene stems and leaves. Leaves reac ovate, ony cordate, long-petiolate. Bee s homogamous, many-flowered, medium-sized, subcorym- bosely or subracemosely ar wa on 4 to long oi eae Phyllari ies very sora Lae in many series, narrowly ia tl tapering into a usually uncinate spine, the outer and middle phyllaries reflexed or spreading above the appressed base. Receptacle coarsely setose. Achenes compressed with a basal hilum, the sides tending to be spongy-rugulose or 5891. Brickellia greenei 508 COMPOSITAE -ribbed. ee bristles in several series, free, a little unequal, strongly scabrous-setulose, deciduo s Limeiat “ name derived from the Greek word for bear, perhaps in reference to the eat involuc ; A or of sis " species found chiefly in Eurasia; becoming naturalized elsewhere. Type species, Arctium appa L Heads mostly 2.5 cm. broad or less, gi Maange pint along the branches; inner phyllaries usually purplish- 2. tng the a serrulate with very fine, ardly pointing, oan er teeth; pappus rhe jatios. short Heads most] cm. broad or more, subcorymbosely arranged at the ends of stems and bra nchlets; inner thy llecies bi is fea margins ciliolate with small reflexed or outwardly pointing Sich pappus-bristles to Sa r 6 mm. lon 2. A. lappa. 1. Arctium minus a Bernh. Common Burdock. Fig. 5893. Lappa minor Hill, Veg. Syst. 4: Arctium minus Bernh. Syst. iy re wn 1800. Stems to 1 or 1.5 m. tall, the branches Sabet eey erect. Basal leaves ovate to round-ovate, deeply cc awk the nPah “blades small, Seg cond-year blades becoming as much as 6 dm. long d mu maller bcor uneate a ubglobose, 1.5-3.5 cm. wide, sessile or pedun cul: ate, | racemose or subracemose along the branch- ine, the inner : _Sp the lower margins of the middle phyllaries ane ihe upper ee gins of the inner phyllaries peetrmed serrulate, the teeth strongly ascending, ie apenasies sometimes a little pile i Ses my or peg pe Se hones setees ally pink or plish; achenes bias -cuneate, trunc an Re te = lightly gonads atisdled or -ribbed, pappus buff, te satis frietles. hoetiy . 52. oS a Fields, roadsides, and waste Ley in towns, occasional but widespread, sometimes locally abundant; Wash- — - central. sEea ate Se bs inary a; east to the Atlantic. Naturalized from Europe. July—Oct ~ rts ants are her: ted as A. minus, one with heads about 2 cm — and the Sune with heads abo: Ba tend 5 The. daiter Soni may be the one called A. nemorosum in eastern United States, but with us the ties appear to intergrade completely. 2. Arctium lappa L. Great Burdock. Fig. 5894. Arctium lappa L. Sp. Pl. 816. 1753. Lappa major Gaertn. Fruct. 2: 379. 1791. erb 1.5-3.5 m. tall. Leaves ovate, errs cordate, the basal leaves of the flowering abrate e at bas Ss : ; orym bosely arranged at the ends of the main branches or on short lateral branches; gees series hd or rarely Despuee west sabrous the outer and middle lenceilite. ee base appressed, the u part spreading or reflexed, tapering to the slender uncinate spine, the inner phyllaries Mia chartaceous, wekcatine or caisson ihe € spine straight or uncinate, the aia a the inner phyl- i e of the outer and middle phyllaries narrowly scarious, minutely ciliolate with flexed, gl 2 irs; i horte he ph u 1 , purplish red; achenes oblong-oblanceolate or somewhat cuneiform, 6-7 mm long, bro or grayish brown, sacl smooth or spongy-wrinkled or -ribbed, pappus ivory- white or buff, the longest bristles 5-6 m long. Occasional weed of seal roadsides, and waste ground; west the Cascade Mountains in Wash- mates and in the Coast Ranges in California suiath to southern "Coie Sritich Columbia; central and east- ern United States and Canada. Native of Eurasia. June—Aug 137. SAUSSUREA DC. Ann. Mus. Paris 16: 156, 196. pls. 10-13. 1810. No men conservandum ae unarmed herbs with alternate, ene re toothed, or pinnatifid leaves. Heads m r large, solitary or corymbosely clustered, homogamous, the flowers all perfect. Ss ory Weuedae cylindr rc-campanlate, or turbinate, the phyllaries numerous in many series, appressed or squarr cute. Receptacle plane or convex, nake or bearing chaffy scales or isties 4 Cordilas elas 5-lobed, bluish or purplis sh. Achenes oblong, more or less angled, hilum basal. Pappus-bristes i in 1 or 2 series, plumose, united at the base and generally decidu ous, an outer, shorter, dissimilar series present or lack- ing. [ Named for Horace Benedict de Saussure Be his son, Theodore, ai known natural- ists of Geneva, Switzerland.] A genus of about 130 species, m mostly ‘h montane, occurring chiefly in Asia, with several species in Europe and North America and one in Roan. high 2 species, Serratula alpina Willd. SUNFLOWER FAMILY 509 : y Spi. Ge $893. Arctium minus $834. Arctium lappa 1. Saussurea americana D. C. Eaton. American Sawwort. Fig. 5895. Sa D. C. Eaton, Bot. Gaz 283. 1881. Saussurea ne var. cordata Kurtz, Bot.’ Jahrb. 19: 354. 1894. Plants with simple, erect, leafy stems from the wn of a short stout rootstock, sparse puberuent = roniehtalee and a little stacitoler, pantioulatty on the re side of the leaves ; eae 3-10 ) dm. tall, terete, mostly unwinged or sometimes the leaf-bases a little decurrent. Leaves thin herbaceous, sieges, Roney to oblongish or lanceolate, saliently dentate, the lower nuate and acute at apex, 5-1 cylindric or oblong-turbinate, about 2 cm. long; phyllaries numerous in many series, closely appressed, acute, sparsely arachnoid- or ig tee Detar i coriaceous, the outer and middle broadly triangular-ovate or te, the inner lanceolate, somewhat membranous on the margin, exposed parts of the phyllaries frequently tinged with dark brown or purplish black; receptacle or fimbrillate ; s lavender i re mooth ; pappus to 7 mm. lon inner series more = eaceous an e or less connate at the baie about long, pa ig light bro ee ereoreulls persistent. ae st ped wy or sa ihe pes in the mountains, Canadian and Hudsonian Zones; Olympic and Cascade Mountains, Washington, south a a" east to the Blue and Siskiyou Mountains, Oregon, and ~. = Sera Moun- enetig ‘Cal chores: "pait to tdahe and Montana; astern Alaska and perhaps adjacent British Columbia. Type loca lity: “mountains of Union Co., Oregon.”’ Collecte by Cusick in 1877. July—Sept. 2 dm. tall o r less with | narrow ‘elliptic ‘leaves that are scarcely reduced upward and frequently surpass the densely comecunchiseced heads. Mountain slopes, Hadeonies or Arctic-Alpine Zones; reported from Mount Benson, Vancouver Island; Rocky Mountains ta British Columbia and Alberta. Type locality: “elevated parts of the Rocky Mountains.” Collected by Drummond. 138. SILYBUM [Vaill.] Adans. Fam. Pl. 2: 116. 1763. Annual or biennial pies with vy! stems a alternate i ag leaves. Heads peduncu- late, medium or large, many-flowered, homogamous. Phyllaries numerous in many series, the outer and middle Sah ed a tiff spreading, eee folinccocs appendage. Receptacle fleshy, setose. Achenes compressed, nearly smooth, hilum basal. Pappus-bristles in sev- eral series, setulose, united at the base into a ring, aa hsainas. [From the Greek name of a thistle that was used for food. ] of 2 species indigenous to the Mediterranean region. Type species, Carduus marianus L. 510 COMPOSITAE 1. oe dene marianum (L.) Gaertn. Milk Thistle. Fig. 5896. rduus marianus L, Sp. Pl. 823. 1753. Silybum marianum — Fruct. 2: 378.. 1791. p e le road, clasping, shortly ge ab base, shallowly or deéply lobed, apinogl iiectare most leaves much reduced with strongly spiny basal lobes and attenuate tip appen i en enuat id tipped, equa ey or a little longer than the owers, the inner phyllaries peace coriaceous, bearin a reduced appendage or narrowe pod. an acute Resim a : rs numerous, purplish, rarely ong, tely and dotted with buff an nd dark bro n; pa ppus- briatige ivory-white, a little unequal, 1-2 cm. long, the outer setiform, the inner hesated and a little paleaceous, the top of the pappus-ring bearing fine erect hairs to 2 mm. | i Occasional in waste grou d along roads west_of the Cascade Mountains in Washington and Oregon; a common and wide pptiad: ees Y B eee euewbions in gia — of the Sierra Nevada; Vancouver Island; occasional east to the Atlantic; South America; Australia. Native of the Mediterranean region. ‘Apri: Aug 139. ONOPORDUM [Vaill.] L. Sp. Pl. 827. 1753. Tall herbs with erect stems and alternate, simple, spiny leaves, the cauline leaves de- current and the stems conspicuously spiny-winge . Heads homogamous, large, mostly soli- tary at the ends of the branches. Involucre broad, the phyllaries numerous in many series, entire, narrowed into a stiff spine. Receptacle fleshy, not setose, alveolate, the pits mem- branous-bordered. Ach me compressed, pubescent, transversely rugulose, hilum basal. Pappus setose, the bristles slender, scab ous, in sever series, united at the base into a low ring, the whole deciduous. [Name from me Greek, meaning donkey flatulence, the plants believed to cause a flatulent state in don bout 20 species, native in Europe, Asia, and northern rem Type species, Onopordum acanthium L. Onopordum acanthium L. Scotch Thistle. Fig. 5897. Onopordum acanthinm L. Sp. Pl. 827. 1753. Plants biennial, sometimes short and slender in depauperate specimens but Ae rgiatt very port sd 2 m, tall or more, openly, fastigiately and virgately branched, greenish or canescent ith a close arachnoid tom entum ; stems broadly winged by the secorvert leaf-bases, ray wings i i Lowest | achenes narrowly obovate, 5-6 mm. somewhat essed, transversely rugulose, m less pubescent especially below the m ‘dae. brownish Fite pale ‘below, almost black at the top ; pappus of numerous slender bristles, about 8 mm. lon Fields and roadsides, becoming commen in the cue . aes beg souerect® Washington to cen tral and eastern Oregon, and Modoc, Passe, and Lake Bs ig: mas Californ western Idaho and Nanaimo, British Columbia; widespread in the United States, Native pe and is oe ka gre ept. 140. CAETHAMYS een L. Sp. Pl. 830. 1753. Ann ial, sinielte spinescent or spiny leaves. Heads medium to larg: e, homogamous, many-flower solitary at the ends of leafy branches. Phyllaries fee se} 32, in many series, the outer and middle foliaceous with a spreading, spi ny, coriaceous blade, the inner phyllaries seg and spine-tipped or bearing a denticulate or pinnately parted, spinose appendage. Receptacle coarsely setose. Achenes turgidly setae hilum obliquely lateral. Pappus none or the paleaceous bristles numerous in many series, unequal, free, persistent, setulose, occa- nt rudimentary. [Name probably derived from the Arabic name for the safflower ant. i. A Bers: of about 20 species occurring chiefly in the Mediterranean region. Type species, Carthamus tinc- orius SUNFLOWER FAMILY 511 Leaves and outer phyllaries pinnately parted or lobed, Yate spinose; stems and leaves more or less glandular- pu oo scent ee arachnoid, especially near the inflorescence; corollas yellow, the upper part of the tube eolate-in Sends broadly ovate, the phyllaries _— or a little longer than the flower . C. lanatus. Heads narrowly ovate or oblongish, t er phyllaries attenuate and sac hones than the flower ws eticus. Leaves and outer phyllaries sag -serrate or sometimes entire; stems and leaves glabrous; corollas orange- -red, the tube slender throughou 3. C. tinctorius. 1. Carthamus lanatus L. Distaff Thistle. Fig. 5898. Carthamus lanatus L. Sp. Pl. 830. 1753. Plants annual or perhaps sometimes biennial with slender elongate taproots, leafy, stiffly erect, few-branched above the middle; stems 1-10 dm. tall, pubescent below with sparse jointed hairs, 5895. Saussurea americana ci) hh, s 5896. Y 5897. Onopordum acanthium att ¥ Vai We 512 COMPOSITAE pubescent and sag ea a Bd fons above, as well as a little glandular, terete and wingless. Lowest leaves herbace to7c g, obla nceolate, petiolate or sessile, pinnately parted si acute or i n rgins bearing a one arp spino er ; ng, sessile by a broad bas stiffly coriaceous, promi- nently veined, viscidulou bed pinnately lobed salient denta ate, ‘te lobes and teeth rigidly spinose ; heads solitary at the e of the r bra large, about 3 cm. long, o i volucres arachnoid-pubescent ri viscidu ae: “hai ae the 9 ‘and middle phyllaries foliose, shorter or a little longer than the flowers, ditaietely spreading or squarrose, pinnately lobed or dentate rigidly spinose, — inner phyllaries s erect, entire, coriaceous except for ee scarious margin and appen e appendage with a terminal spine and lateral sitiles sides merely denticulate ; flo a many, the throat inflated ; era an t as broad ale brown or buff, mottled or speckled with dark brown or black, the sides irregularly horizontally pitt wer rim of the pits and the tr a ge 2 of the fig ad ia dentate or denticulate, or the longitudinal angles of the achene produce ard into ac ppus no h chenes, p and persistent on the i inner, aohich yay the pileae in several series, unequal, se shorter outer ones emarginate, the inner longest ones acute, the innermost sometimes abbrevi- ted. Dry open hills and along roads; locally common in central California and around San Francisco Bay; reported from southern California. Native of the Oid World. April—Oct. 2. Carthamus baéticus (Boiss. & Reut.) Nyman. Smooth Distaff Thistle. Fig. 5899 Kentrophyllum baeticum Boiss. & Reut. Pugil. ae - ov. 65. 1852. Carthamus baeticus Nyman, Conspec. FI. Eur. 878. Kentrophyllum lanatum Bi netics Battandier, F 1 lage 508. 1888--90. Carthamus nitidus of C Similar to C. lanatus but the stems whitish and more shiny. Leaves frequently longer and narrower, attenuate at the tip into a stiff slender spine; heads more sle eupl oblong-ovate, and fewer-flowered; outer and middle phyllaries divaricately spreading, the tips et od attenuate and i h lon t ers inne spinose, x r, denticulate or the phyllaries entire and attenuate; the outer epappose schieess darker and more evenly and regularly rugulose, the 338 pappus- ener achenes with smoother, less denticulate sides Opes or brushy hills and roadsides in central and southern California; foothills of the Sierra Nevada in evade ~ Tuolumne Counties, and local in S ; an Joaquin County; cesasiontl from San Luis Obispo County to San Diego County. Naturalized from the acitecvinaee region. Jun It is no S$ easy to bi gol ay ein from C. llistar & soy some European botanists have wees as a variety. Cytological — Ai ri, however, have howe that 2n = 44 in C. Janatus and 2n= 3. Carthamus tinctérius L. Safflower. Fig. 5900. Carthamus tinctorius L. Sp. Pl. 830. 1753. Plant annual with erect stems narrowly branching above the middle, leafy, glabrous; stems generally 4-6 dm. tall, smooth, terete, whitish or straw-colored. Leaves coriaceous, spinulose- a s le and upper ca o a pine ivory-white, nearly smooth, subobovate, about 6 mm. long and 4 mm. wide, generally without pappus or with a few short rudimentary paleae, ee the pappus well ‘developed with paleae numer- ous i seenerp — buff, unequal i in length, m stly shorter than the achen Occasional fugitive from vation on Seaddshtcs and Feasied the edge of fel, probably oe Bagh eres, gala” mento Valley to re "kieine Conde , California. Of widespread occurrence wa of the Old an New Worlds wherever the plant is cultivated; originally deacktad from , al Sane-Oek, 141 CYNARA [Vaill.] L. Sp. Pl. 827. 1753. Robust perenay << with large basal leaves and stout erect stems. Heads large, pedunculate, many-flow omogamous. Involucre globose, the phyllaries imbricate in many aig aneeeite sad: 4 tase suliriesdly coriaceous, narrowed into a stout terminal ae (or rarely the phyllaries rounded and innocuous ). Peeteiacie fleshy, long-setose. perfect. Achenes a little turgid, glabrous and smooth, hilum basal, apical i a n med. Pappu paleaceous, united at the base and deciduous. [Name derived from the Greek word for dog early applied to a kind of artichoke. ] a nat genus of 11 species, natives of the Mediterranean region and Canary Islands. Type species, Cynara car- SUNFLOWER FAMILY 513 1. Cynara cardinculus L. Cardoon or Thistle Artichoke. Fig. 5901. Cynara cardunculus L. Sp. Pl. 827. 1753. Plants perennial with broad oe rosettes of large basal leaves and erect thick stems, the og and lower sides of the mares gray or white with an arachnoid tomentum; stems striate, tall. Basal leaves oblong or ohn d aneeolate to over 1 m. long cad about. half as pre ifpairiati- se divided to the didureeat tly winged rachis, the cegiiietitl irregularly parted o ded into nar- w, lobed or toothed laciniae, the lobes and teeth iene ie — t, stiff, yellow seine see fee and pies above, tomentose below, middle and upper cauline leaves much smaller, sessile, a little de- current, those in the inflorescence reduced to line tes or oblong. spin e- sheathed bracts to 5 cm. long; ] cm. long, solitary ends of sho i ubglabrous, sides os ing or reflexed, the puede and upper appressed-imbricate, the middle narrowed into a short, stout, rigid spine, the uppermost bearing a rounded or subtruncate, denticulate, shortly spiny appendage ; poh bluish n purple: achenes 6-7 mm. long, oblongish, light brown mottled with brown and black; s 2-3 c¢ g, ivory-colored. S Oiecabtaiidtly escaping from cultivation; central to southern coastal California, sometimes becoming a pant noxious we — as on Nae hills at the north end of San bein Bay; Lower California; southern South America. Nati of the diterranean region and Canary Islands. June-July. Cynara sabi dusl . pik Pl. 728. 1753. (Cynara carduncilus subsp. _scolymus Hegi, Fl. Mitt. Europa ~ 924. 1929.) Differing from Cc; carduacnios in the spineless leaves and in the blunt or acute, spineless or arly spineless phyllaries. nie ci weg cultivated in Pg California where it is an occasional escape. Probably : pow fe of C. cardunculus. Globe or Garden Art 5900 5898. pease lanatus 5899. Carthamus baeticus f 4 ©5900. aces tinctorius 5901. Cynara cardunculus 514 COMPOSITAE 142. CIRSIUM Sis * Adans. Fam, Pl. 2: 116. 1763; emend. DC. Prod. 6: 634. 1837. Annual, biennial, or perennial, more or less arachnoid-tomentose herbs with posias x or stout taproots, the stems of the perennial species arising from the caudex-like wn of the taproot or from buds produced on widely spreading, horizontal, rhizome- lik roots. Leaves alternate, generally spiny-lobed or -divided, sessile by a petio ole-like or amplexicaul base, the base sometimes more or less decurrent along the stem as a spiny wing. a ge hairy. Flowers perfect or — wr paree and the plants nearly or quite dioecious ; corollas whitish, yellowish, rose, or red and tubular with a slender tube and 5-cleft limb. Achenes narrowly obovate or Sites, sapeee ot smooth, the hilum basal. Pappus of numerous plumose bristles (or sometimes a few merely barbellulate), these united at the base and deciduous in a ring. [Name Eons the Greek, referring to the use of thistles as a remedy for swollen veins. ] A genus of about 250 species, in the northern hemisphere. Type species, Carduus heterophyllus L. For authorship of generic name, see ag —— 8: 278. 195 e of the reasons for difficulties e ered in treating our native species of Cirsium is ae natural lines = a become blurred, if not all bat Cbiiterated dad . Phat 93 hy —— zation. Not only does hybridiza- r be pals closely related series} rs between species which wo distantly related i sie onde Ea oS prin a Cc. ee rcetoeik ele d08 C. brevifolium and C. utahense, cy: Dur rse 0 thistle over i in “field and herbarium, _Sybrigs (suspected or otherwise) have becin tute seal sting in hae ington, Or — and mut ornia her 99 the following entities: andrewsiit and rcetorum, ee cwsit sat remotifolium Cc . mendocinum Petr: eeeialbion and foliosum, brevifolium tad undulatum, brevifoli and utahense, ae Sr rnicum and cymosum, “calsfornicum var. pierce and foliosum a quercetorum _ eek wean cum f. parishii paok). callilepis and querc m (cf. C. amblylepis Petrak), callilepis and remotifolium, callilepis var. pseudocarlinoides and cymosum, call epis var. pseudocarlinoides and hallit, canovirens and undulatum, canovirens an F eaphense cf. “C. subniveum” of eraeed ee _ cymosum, cymosum and pastoris (?), cymosum and proteanum, cymosum and remotifolium, douglastt and quer torum, douglasii var. canescens and cymosum, foliosum and undulatum, fontinale and quetcrtos yeh, yes ei and proteanum, occidentale and quercetorum, pastoris and proteanum, proteanum and remotifolium. In the — wing account of the genus Cirsium on the Pacific Coast, only ree a ™ the or immediately pertinent to the accepted name has been given. For further synonymy a many other data, see Franz Petrak, cea Pines mcg Arten der Gattung Cirsium (em Bot, Centralbl 35: 223- 567. region Plants introduced ag ged Old World, either dioecious with unisexual flowers in small heads or monoecious perfect flowe medium-sized. heads, if monoecious the oo on of the leaves scabrous or Sis Plants dioecious REISS heads commonly numerous, small, 1.5—-2.5 cm. long; leaves pet scabrous-hispid . arve Siuac maroon biennial; heads medium-sized, mostly 3-4 cm. long; leaves see ‘ispid on the upper Pl. e New eho monoecious; upper surface of leaves not scabrous-his eae abe pectinate on the margins (rarely the outer piataries Pearle bi in C. callilepis, C. doug- lasit, C. hookerian ly and C. nidulum; uppermost spiny-margined leaves sometimes closely subtend the head = simulate phyllaries Back of at least some of the oliitlecies iene a mare! or spot or ridge. cence of stems and | oid. Flowers dat i i aay red or ia mine, straight in the head; soncecten perennial from the tra Nevada ea ssi 14. C. nidulum., vida es lavender, pink, or pial, the outermost in the head spreading. lants perennial ep new rosettes arising from horizontal roots; heads medium-sized o large, (2 or)3— long; plants “oF . hills and forest borders, chiefly of Oregon and Washi 1 tend ee and eastward). Stems mostly 1-2 m. tall; heads 3-4 cm. - ng, somewhat constricted above the sub- lobose hase; oraee om phyllaries 1-2 mm. long; south: centres Oregon and adjacent Californi B.C: ch Stems mostly 1 m. tall or less; spines on phyllaries mostly 3- ‘“ mm, .. northern Oregon northward and eastward, Flowers pink or ila rarely white; leaves wine, yg and below; heads 4—5 cm. lon undulatum. Flowers elon secdia: leaves mostly bicolored, green above Se ll (2 or)3-4.5 4, evifonw ft if sometimes perennial the ne . rosettes co. from acoene on the, cigeoer: — leaves sometimes markedly decurrent — medium-sized, 2-3.5 cm. long; plants of wet ine chiefly in Califor Plants a arachnoid or 2 labrescent, the upper side of the leaves elaros or nearly plants of central Calif 7. C. hydrophilum Plants pi aa scarcely glabrescent, the upper side of the Sen with per- sistent arachnoid tomentum; plants of desert ranges oa — m Plants mo the —_. like s small or avense. of stems and leaves crispy-puberulent as well as arachnoid (the crispy-puberulent t hairs are monbea ular. translucent, shining, and often iridescent; sometimes they are quite scant and sometimes a: ered by a thick coating of arachnoid tomentum). Phyllaries toed & re except for the erect or spreading chartaceous tip (i.e., all o —— all mech the herbaceous part of the metiare at Pct A nga os spot on phyl- Spine at bie Sat ok bares widely spreading, mostly 3-5 mm. long; taprooted, monocarpic a a ennial plants of wet places from southern Oregon yap Raat to central Californ SUNFLOWER FAMILY 515 Spine at tip of phyllaries erect or somewhat spreading, mostly 2-3 mm. long; taprooted pare ennial of dry slopes east of the Cascade eC. eaK0 ee ‘and northern egon ca Phyllaries not clos sely imbricate and e rapa ely spreadin bove the psa Mag = fe rae Bi pat ecg i °C. yore riica im ag gocher : wipe dilar spot, frequently inconspicuous or lacking on many phyllaries in a head. pomeeo tipped by a short spine mostly 1-3 mm. long. nts _ Perennial, the n new — -rosettes arising from rag sm aa — or from the f the taproot. . C. cymosu t Plants monocarpic, winter ssiree or biennials with ma 11. C. californicum into an elongate slender spine mostly 3-10 mm. long; plauts gener- oca pale: biennials, rarely perennials, with taproots. Heads often aeaee than long, hemispheric; phyllaries we spines mostly 5-10 mm. ee elt he oute . C. neomexicanum Heads — ose; ass laries with spines mostly 3-7 mm. long, the outer Sissies spreading or ascending 13. C. utahense. Back of phyllaries without a ae lar spot or ridge. Corolla-throat gradua lly wed downward into the tube, the corolla without a distinct ion between an throat and the. tube; flowers usually straight in the head, pinkish, hea oa or reddish (or white in albinos); pubescence arachnoid or aganpecne villous-arachnoid. PRerigret Log pe seeenctt more nk ive saree ent; perennials stout oy ; southern vada e ard to des 13. tar ke - — Phyllares, thickly aaa persistently t arachnoid, or in C. pro m the tomentum mes thin and the Dhyllaries nl ccenedtt ical snniaia "oe “biennials, rarely rane s southern Oreg western Nevada and southern erect Heads often as bro and r broader; phyllaries generally str poag! ost acicular above a: Vanpressed. ae Seunonak pes a fine, grayish white aoe corolla 2.5-3.5 . long, scarcely exceeding. the involucre; rer ian core at of central Cali- i 15 ct Heads generally longer than broad; phyllaries generally curved down ae outward or castro linear to ovatish but a acicular; corollas to 4 cm, rg generally con- soit usly exceeding the involucr Planta, thinly t ly arachnoid- Aomentose, eseey ae -tomentose; phyllaries often subglabrate; widespread and common in California hills and mountains from Ventura and Kern Counties Gerthersed. 6. C. protcanum ny po ide! edge with dense, felt-like siebsiies phyllaries whits: arachnoid- — cept for the glabrous yellowish spine; southwestern Oregon to al Californ, and adjacent Nevada. 17..€. ” es Cocoa shenet wrosid or Shea distinctly contracted at the base, the throat and t ne are nerally distinct; Besse: aight in oot aap vol on the outermost Beds h spreading, whitis elspa pink, sometimes re bescence, at least on the consisting of crispy-puberu ied ‘sdnciannid reir an ae y-puberulent hai Aecaem rn a el yee ellular, coreelitan S visc ree fea sometimes abundant, lacking i in C. moha in margins lac corr or yer (nbyiaries mre wees a little catendol in intermediates latiaen c callilepis and between C. foltosum and C. hookerianum). Plants embiaad: mostly biennial with taproots; bccn ea in eters] North America, mostly near the C. 6. C. folios Plants Peres) with cee Pegs roots; California Coast Hole and Oraie chiefly west f the Cascade Mountain Heads medium-sized or 524 COMPOSITAE 1d Cirsium neomexicanum A. Gray. New Mexico Thistle. Fig. 5913. A. Gray, Smiths. Contr. 56: 101. 1853. Plants biennial or short-lived perennial from a stout taproot, generally monocarpic; se “8 (or 20) dm. tall, stout ur ra aber, rah elongate slender branches, white- sgpeneae pitabaee a little crisp- Seat nt, the tom m becoming thin and floccose on the lower part. Basal leaves forming a dense, many-leaved salieis that probably develops through more thes one season, narrowly oblanceolate, ie earliest shallowly sinuate-lobed or deltoid-lobed, becoming more deeply daend up to 3 dm. long and 6 cm. wide, the i pig sions 2- or 3-lobed with the lobes terminating in a stoutish vetlow spine, hes biides' narrowed t short, more or less on ccigar ied piny-bordered, petiole-like base, tomentose on both sides but greener ro subglabrescent above; lower cauline leaves similar to the basal leaves but eo r and becoming sessile ie < cpr decurrent base, he stem someti spiny-winged near] ihe next node; upper leaves becoming much smalle € Ss imes -win yt and bract-like, very spiny but scarcely decurrent; heads solitary, oe ly “ he Nes: of er slender branches, usually broader than long and hemispheric, 3-6 cm. long, wide volucre ay nom to broadly bowl-shaped, more or less ir A tom aati e or ‘the ileiets tending to be i i sometimes e cose and d ; phyllaries linear-lanceol ap- pressed, the elongate tips of the outer phyllaries cacy d, h phyllaries divergently spreading or loosely ascending, straight or the uppe es sigmoid-curved, the base chartaceous, the tips herbaceous and often too snail rie Alou the middle, sometimes glandular, tipped by a strong yellow spine, the in phyllaries flat, attenuate into a slender chartaceous spinul flowers white, pink, or inveader ‘the Sererciniit generally cutacatdy. Dekadiim: the corolla- be ~2 cm. long, the throat 5-8 mm. a i aij lobes 8-10 mm. long; achene purplish bro wks elliptic- oblong or -obovate, 5 mm. long, 2-2. wide; pappus 1.5-2.5 cm. long, white, sometimes with dark-colored processes are sides of cee near the bas Dry, gravelly or rocky slopes and canyons, Lower and 98 Sonoran Zones; mountains in the rai = of the, id staee Desert. California, at to Colorado and New Mexico. Type locality: “Side of the Org tains,”” New Mexico. April- Sept. 13. Cirsium utahénse Petrak. 56 a Sees Thistle. Fig. 5914. ? Carduus nevadensis Greene, Pittonia 3: 26. 1896. Not Cirsium nevadense Willk. 1859 ? Cirsium humboldtense Rydb. Fi. Rocky Mts. 1007, 1068. 1917 este eas eee Petrak, — Bot. reps ie 470. 1917. M. E. k, Madrofi Plants with a long, ee or stout tape the —— Labia woins ce through two to four years, monocarpic or sometimes perennial with n osettes arisin the branched, caudex-like crown of the taproot; stems erect, 1 esi tall. see or Sightly tne yee arachnoid-tomentose and crisp-puberulent. Earliest — ellie o oblanceolate, alm diately denticulate and with numerous small spines fringing the 2 Sarl sp posite dents estar n d natel wi e ‘ Ss, closely t solitary at the ends of longer or shorter, leafless or leafy-bracteate branchlets, subglobose, 3.5-5 em. long; involucre campanulate, conspicuously aca tomentose or the tomentum tending to WwW \wif yy LY = Nie ~~ “Wy ie yh f SW ey — NS 5913. Cirsium neomexicanum : 5914. Cirsium utahense | SUNFLOWER FAMILY 525 come somewhat floccose; phyllaries appressed at the base, loosely ascending or wer spread- ith s atin Pia and niin lacerate ; flowers whitish, evender tinged, or pinkish, the corolla- tube 1 m. long, a throat 9-10 m m. long, the oe 7 mm. long; achenes oblong- oblanceolate, poate h, 71 mm. lon payee whi tish, 2-3 cm. lon Gravelly and sto, Wibod, meadowy flats, ie open a usas woodland, Sonoran and Boreal Zones; mostly east of the Cascade-Sierran crest from Panay gg orgs to middle ws rage eastward to the Rocky pay po s and southward to Utah and perhaps to Arizo Type locality: Silver Reef, Washington County, Utah. ay— Plants from the east side of the Sierra Nevada in California and Nevada are quite variable in habit, pubes- cence, and flowers. In all, however, the niga: to sage pense 3 phyllaries are ngs to one ithe middle and are attenuate into a free tip ending in a slender, stiff, spreading or ascending spine. se plants may be the same as C. humboldtense Rydb. rduus nevadensis Greene; type locality, West 0, Og Si ceeurics, Nevada), and they may prove to be specifically distinct {rome C. utahense. t astern O C. wallowense M. Peck), the phyllaries are scarcely ever glandular, whereas in th m from southern Utah, the png line or ridge is present and conspicuous. In other parts of its range (as in eastern California), C. utahense may b riable his character. In northeastern Diesée, where a —— intergrades with C. Seevtietiam and with C. canovirens, the intermediates seem to be the — of ~ acon Car Fl. Fran. 479. 1897. (Carduus vaduleins var. nev aden Comms , Proc. d. Phila. oo ne omy ee Plant to about 3 m. tall; stem stout, thinly floccose-arachnoid; S green and gla- brate on the upper side, white-arachnoid on the lower side; heads pose about 2.5 cm. lalee Bhyllaries appressed except at the slender-spinose tip, closely imbricate in 6 or 7 rows, not glandular, the outer lanceola vate- lanceolate, the inner linear-lanceolate; flowers Lc aa Pegi plant, definitely known only from Greene och gr in 1883 near Truckee, Nevada County, California, may be a small-headed form of " Cirsi um utahense Petrak, s that —. ot along the east side of the Sierra Nevada; or ~ Truckee thistle may repveuin a hybrid a ivative between the Utah thistle and Cirsium meee sit var. canescen Cirsium pies Rydb. FL. Rocky Mts (100. 1068). 1917 [1918]. Plants from northeastern Oregon that are related to : Rccprtsd but with pate beds ve been referred to this species Cronatist, Re — Pi. Pact: Book hw. 5: 4955), ai tee “ this coe es we have seen from Wallowa County Cee, coelie 4 plants ie ah mse (i.e., C, llowense for fod srk hele ig ome ater Gi rens, peel it seems Weahenie that the ‘ pfctl x) plant: . may have b tion Gecvecn a thos species. Type locality of C. subniveum is Jackson’s Hole, Wyomin ae 14. Cirsium nidulum (M. E. Jones) Petrak. Nidulous Thistle. Fig. 5915. Cnicus nidulus M. E. Jones, Proc. Calif. Acad. IT. 5: 705. 1895. Cirsium nidulum Petrak, Beih. Bot. Centralbl. 35: 553. 1917. s perennial with long woody taproot that may be once if several times branched at the top to add ce a leafy crown and 1 or 2 erect toate seis the base of the stems closely invested with the dried scaly bases of eee leites stems 2.5-10 dm. tall, aiimple or few-branched above, asal leaves few cm. lon ong, the g, 2-2.5 mm. wide, strongly of the lobes i Bag light to dark wore ;pappus W whi — - cm ee pra cade ine morain no: Zones ngrong Sot gt na fem desert ca pei ~ tees ranges ot eastern Calitareia, caateraed mec “Utah ee Arizona. Type locality : n red alkaline sand, along the 3 athe. of the Pahria River, at Pahria [Paria],” Kane County, Utah. June—Oct. 15. Cirsium occidentale (Nutt.) Jepson. Cobweb Thistle. Fig. 5916. Cardu ee Nutt. Trans. Amer. Phil. Soc. = 7: 418. 1841 Cirsium c ulteri Harv. & Gray in A. Gray, Mem. Am read: 1g OS 110. 1849. cr sium occidentale Jepson, Fi. W. pre Calif. 509. 1901 coulteri Jepson, Man. FI. Pl. Calif. 1167. 1925. annuals, —— a slender ng ingheb taproot; stems erect, leafy, usually rtas7c Biennials or winter an — - tall but sometimes a s sho’ . or as tall as 1.5 m., thinly to thickly arachnoid- or entose, rarely glabrescent, Sine temocals stata a ulent. Earliest mes oblanceo- late-elipte, sparsely spinose-dentate ; rosette-leaves numerous, at first sinuately lo 1. g, becoming 3 dm. long and 0.8 dm. wide, meaty oblanceolate, rather deeply lobed, the lobes 215 526 COMPOSITAE elow into a spine-bordered, ‘pet iole-like base ; basal leaves and also ne peng thinly atacand, . thickly white lust on the side, ar id-fl se, or arachnoi p i e cauline oblongish or lanceolate, shallowly to deeply spiny-lobed, heccrnie: Rte sessile with lo or shorter, decurrent, ings the point of attachment st leaves quite reduced but scarcely bracteate, conspicuously spiny ; heads solitary at the ends of more or less leafy stems, medium-sized to 1 ,3-5.cm. | broad ; involucres campanu- te to broadly hemispheric, the numerous phyllaries interconnected b b : grayish white, filmy tomentum, the tomentum tending to b occose in age; phyllaries acicular, linear or linear-lanceolate, th g downward-poin f the base, the middle phyllaries spreading horizontally or ascending fr rt appressed base he spreading part thickish te, attenuate into the short, slender but stiffi e, the inner phyllaries flattened towar e tip, sharply acute or weakly spine-tipped, spinulose-ciliolate and a oO ward, not pa co = o 6 c = wn co oO a a > ao i =A ae oy i = 3 n (md | ist) = 08 > Coa “= a “ go =) ) 35 + ze je S fe) less curvin rd an glandular ; flowers purplish red or rarely white, scarcely ex ceeding the in valiire: corollas ee of 3.5 cm. long, the tube 13-18 mm. long, gradually widening es the throat, the lobes 7-1 lone. bags af pidaggting linear-filiform due to Sirol trig of margins; achenes oblong to ieee cun oe . long, 2-2.6 mm. wide, brownish, somewhat shiny ; pappus white or sordid, 33 ¢ i ong. Claye 7 andy, or gravelly soil of coastal dunes, grassy mesas, and i pan washes, Upper Sonoran and Humid Transition roses: Séncink County, California, oe ard to cismonta a Giualar southern California. Type locality: Santa Barbara, Santa Barbara County ich may also be ise: ‘Pe, locality of fe coulteri). April—July. cobweb Rye is aia? kable and distinctive in its typical form but in places where its rene e approaches or cece’ that of C. proteanum a seeming genetic blending results that oun account for numerous intermediate 16. Cirsium ghia tami T. Howell. Red or Venus Thistle. Fig. 5917. Carduus venustus Greene, Proc. Acad. io beset 359. 1893. Not Cirsium venustum Porta, 1909. Cirsium occidentale var. venustum Jeps FI. Aa var 3167... F925. Cirsium proteanum J. T. Howell, betes rea "Bot. 9 ong a and reflexed, typically the outer ones short, ovate-lanceolate, more or less reflexed, the le ones elongate-lanc coate, bp ap sed carinate-thickened, sovebenty ipceadlana above the. appressed ee so 2 ihe ro and more or less introrse ost erect, flattened, chartac cabrous-ciliola te, alle cent he innermost shortly Te Gases an Fo tees -arachnoid- teiriontoee persisten ntly pubescent or glabrescent, none glandular ; siabak usually crimson-red but varying to ny urplish red ite, strai a 18- an m. long, the lobes 10-12 mm. long; achenes poe ahiea! obovate, 6-7 mm. long, 3 mm. wi bee aS, i base ; pappus akc about 2 om . lon Clay: dahity. or rocky tt Diss grasslan br an or open woodland, widespread and common in the drier nip ore Guscnteias, Up: Son Ge Svat rcstd’ = Canadian Zones; common in the Coast Ranges and_ occasional n the Sierra Nevada, aitienin Shasta County s uth to Kern and Ventura Counties; perhaps local in western middle Ne evada. Type locality: ‘hills of the aaee Coast Range of California, from Vacaville southward. le (Cirsium pastoris J. T. —— Snowy Thistle. Fig. 5918. Card Greene, Proc. Acad. Phila. 1892: 893. Cirsi: d i te mum J. F. Macbride, Cont. Cog Herb. No. 53: 22. 1918. Cirsium candidissim ns Gack s. & Moxley, FI. S. Calif 1923. Not Dammer, 1898. Cirsium pastoris J. T. Hone ele . Midl. Nat. 30: sty Plant biennial wit h a stout taproot, the stems and — white-lanate with a close, felt-like Il e t tire a lobed. ae ent base, the leaves on the flowering branches almost bracteate; heads rather few solitary on longer or shorter, peduncle- like stems, oblongish to hemispheric, ecm Tg lesee and SUNFLOWER FAMILY 527 4-6 cm. long, rarely (in forms intermediate to C. proteanum) more numerous, smaller, subglobose, 2-3 cm. long; involucre bowl-shaped = campanulate, era glabrous spines, the tome entum sake n etween phyllaries iioatly linear to n arrowly lanceolate or the lowest sometim axel pag soasehy spreading or pn) st rec ved a uch as 4.5 ¢ the the d iddle arcuate- divari icate to ascendi m. m. long, the ends of the lobes copitelate: shickerieil below the acute tip; achenes brown, oblong- cuneate, 7-8 oy g bea and about lon 3 mm. wide; pappus white, te slopes and flats, Transition Zone; southwestern Oregon southward in the Coast Ranges to Mendo- ne Coun. hetotadgels mBior and in the Sierra Nevada to the Lake Tahoe region in California and Nevada. ocality: ‘‘ext nort 2-3 cm Type California.” June—Oct 5915. Cirsium nidulum 5918. Cirsium Cee 528 COMPOSITAE 18. Cirsium peer naa Blake. Surf Thistle. Fig. 5919. Carduus maritimus Elmer, Bot. Gaz 905. Cosine auiyacost Petrak. Beih. = prey 35: 288. a Not Makino, 1910, lum Blake, Journ. Wash. Acad. 21: 336. Bi peeks or perennial, leafy herb with stout la stems and leaves succulent-fleshy, every- where covered by a white, persistent, close, suede-like tomentum, the stems erect, 0.1- 1 m. ong, simple or “much branched from the base, giving the plant a rounded We shy appearance.” Earliest hae ne 9p obtuse, entire or u ndulate, unarmed except for small spinescent mucro at apex or i a : 1 b aA fe ed into a rac like base, the blade pinnately. par rted pad rounded, seeente < or broly elliptic Seenient s, the segments entire or undulate or shallowly lobed, tippe ed by a short stiff s and bearing several similar spines on the margins ; lower cauline leaves longish i WwW se that dev amplexicaul auricles ; uppermost leaves closely clustered below the heads; heads solitary or 2 to on short branches in rather loose terminal a rs, ee campanulate, 3-4 cm. long; involucre 2.5-3.5 cm. long, densely arachnoid-tomentos ; phyllaries numerous, strongly ascending, loosely imbricate or more or less ape ~: ci of all pics: the inner bearitig several stiff, spread- ing, straight or slightly curved spines and the apices ending in a stout yellowish spine, the outer and middle ihechue ae tence bike or linear, the inner phyllaries thinner but ending in a stiffish spine, the sides chartaceous-margined and ciliate-spinulose, the spinules sometimes more or less crested ; cor 10 achen rowly cuneate or oblongish, 5-7 mm. long, compressed or plumply turgid, ‘with lateeal pte a evident, chestnut-brown or light t brown; pappus buff A remarkable endemic known only from maritime fees r Sonoran Zone; eg Luis Obispo and Santa Raitace “Countica, California. Type locality: Surf, Santa Sstee ‘Soanty. April- Jul 19. Cirsium “Cine naa Cronquist. Indian Thistle. Fig. 5920. Cirsium edule of authors in large part, no Cirsium brevistylum Cronquist, Leaflets = 7226. 1953. Erect leafy biennial al rarely caring me herb with stout or slender, fusiform taproot ; sinter a nearly simple or beco robust, and much branche ’ with fastigiately amen i 5 ae erminal or racem arise a ote e stems; volucre campanulate, conspicuously arachnoid-tomentose; phyllaries numerous, loosely imbricate in many ate AA the outer and middle fanstotate to line songl attenuate and spinose- cates the inner elongate-attenuate, acute or subspinose, flattened and somewhat twist — te, sometimes a eines “tiie eee ye reroll: urplish rose, harety whi te, 2-2:5 ¢ g, g, the throat 3.5-6 , the lobes 2-4 sat lon eed minutely cucullate- capitellate a pose tip; achenes elliptic or blo slightly cu a ‘5 mm. lon ng, t Se ckish, brown; pappus buff or light brown, about 2 cm. long, the bri et Hig a tt poi eer at the Mo One Ss in woods and brush-or on meadow sas and in forest clearings, Upper Sonoran, Transition, and Lower ene Zones: "British Columbia to southern California in the Coast Ranges and Cascade Moun- — east to Montana and Idaho. Type locality: near Montesano, Grays Harbor County, Woshinetax. April- 20. Cirsium édule Nutt. Edible Thistle. Fig. 5921. Cirsiten edule Nutt. Trans. tsk Phil. Soc. i A 420. 1841. , Ottawa Nat. 16: 902. Biennial or perennial monocarpic ie with suena taproot ; stems erect, widely and openly branched above, frequently stout and hollow, up to 2 m. tall, more or less purplish-tinged, thinly arachnoid-pubesc r glabrescent. Early a te rather Sgr ee eae ate, 0.3-1.5 d deeply _ nt a below ; av flowering season becoming . long and Hsieh g pinnately divided, the ‘divisions lobed or again divided into oblong or deltoid, spine-ti ipped ‘lobe , the = ps narrowed below to a spinose- margined, petiole-like base, weakly arachn te ore sien Habeas above and — Soe teed and i — Narr Vv te, spreading tip, the inner phyllaries flattened, weakly s ianee ciliate- pa ame sane y bright — purple or lavender-rose, more or less me tutcly spreading, the tube about 1 cm. long, the thro SUNFLOWER FAMILY 529 about 8 mm. long, the lobes 4-10 mm. long; achenes cuneate- se bg flattened, 4-6 mm. long, dark cn brown; pappus buff, about 1.5 c gar! ¥e a little s Moist deep soil of woods and meadows or a gh, sandy rocky co eee Canadian, and Hudsonian Zon nes; southern British Columbia and WV pehinge a es = Cas cade Wountsine and Coast Ranges. Type ogee “plains of Ore. caer and the Blue Mountains; co e young posers eripped of their bark, ar monly eaten raw by aah aborigines, her nee a arueweak pledsant sweetish taste.” June—Aug. 21. Cirsium hallii Se eyy x E. Jones. Hall Thistle. Fig. 5922. Cnicus hallti A. Gray, Proc. Amer. Aca = 56: Cirsium hallit M. E. oes Bull. Univ. pes oy a No. 15: 47. 1910. Plan g perce opal short-lived, first blooming the second or ay year, thereafter aebstontoy at the mit of the taproot a several-branched crown that may produce in oe plants as many as 5 or 7 flowering shoots at ne time, green and subg uerouE the stems and leave crisply puberulent and sometimes thinly arachnoid, or the lower Fates of the leaves eravibhe rachnoi s i h i arachnoid; stems erect, — 1, virgate and branching near the top, or bushy and branching fr e. Earliest leaves broadly to narrowly anceolate, merely dentate-spinose o shallowly undulate-lobed, or pinnately parted ; a rather few, to m. long and 1.2 e, oblanceolate, pinnately ivided nearly to the midvein, the divisions crowded or distant, triangular or quadrangular, again lobed or oar with the toed tipped by short weak spines, the hs isn y nid) ALT, ‘ 5919. Cirsium rhothophilum $921. Cirsium edule 5920. Cirsium brevistylum 5922. Cirsium hallii n= IQ - 530 COMPOSITAE margin of the lobes and of the rachis-like border along the midrib spinulose-ciliate, the blade eos sae toa rgunina Bi gt petiole-like base ; lower cauline leaves similar to the rosette- leaves but m € spi ny a the base developing rounded, semi-amplex iat spiny, shortly decurrent savictes upper ap s reduced, sessile by a broadly clasping base, shallowly to deeply erm i i i i t uterm nsed plants ie heads subcongested and leafy-bracteate, parte tan to subglobose, s the head s Gattis deicdinmaneed phyllaries; heads solitary or clustered at the ends of elongate branches or in more conde L d .5 cm. long, in cha ten te bra nched plants the heads no odding after a nthesis ; involucre owly i nd e along the margin flowers lavender-pink to purplish or whitish, more or less doh a ok in the ing the sek ap Ba 10 mm. long, the throat sven peer fa above the t —6 m obes 3-6 m long ; ngewe s 5 mm. long, 2 mm. wide, narrowly obovate, Seem E pappus a little sordid, Prise 1.5.6 ere Deep m of meadows, forest borders, and maritime slopes, Humid Transition and — Zones; Oregon asey FY i. Cascade Milstein: oan ed in the Coast Ranges, from the Columbia River south to Coos Bay. Type locality: Salem, Marion Sete April—Aug. Some of the coast . ds nts are remin sean he C. andrewsii and some of the montane plants resemble C. edule, but all are referred to C. hallit pending further study. 22. Cirsium andréwsii (A. Gray) Jepson. Franciscan Thistle. Fig. 5923. Cnicus andrewsii A. Gray, Proc. Amer. Acad. 10: 45. 1874. Coleone enekioline Greene, Pittonia 1: 70. 1887. i Jepson, Fl. W. Mid. Calif. 506. 1901. Plant monocarpic, biennial or perhaps the rosette sometimes So. through two years; stems erect, 0. m. tall, in robust plants up to in diameter and succulent, usually widely much- 1 te coro aig td purple, more or _ — in the heads, the tube mm. long, the throat 4— long, the lobes 5-7 m madi _ ne cuneate- oblong, flattened: 4-5 mm. long, ety paneer Recncie: pappus buff, aboot A 3 cm Coastal slopes and arroyos, frequently in wet o hy g alo ms around seepages, Humid ‘Trenaites Zone; central cag Sided - per a Lu Monterey County. In San Mateo County, plants of this variety form a raristie ei toa iad exhibit csguees ee. CG. Sar C. callilepis, and ‘tesibiy C. fontinale. Type locality: northwest of San Bruno, San Mateo 25. Cirsium loncholépis Petrak. Gracious Thistle. Fig. 5926. Cirsium loncholepis Petrak, Beih. Bot. Centralbl. 35: 375. 1917. Plants with a stout or slender taproot, annual or perhaps sometimes living more than one eason, generally stout and succulent, subglabrous or tea 8 crisp-puberulent and arachnoid ; stems erect, hollow, subsimple, 0.3-1 m. tall, vascular-striate, commonly reddish-tinged. Leaves id and more or le lent or subglabrate below; basal leaves unknown; er ine leaves oblanceolate, to 3.5 dm. long and 1 dm. wide, ivided nearly to the midri : into several-lobed segments, the lobes tipped y short slender spines, narrowed below into an elongate, spiny-margined, petiole-like base, the base extending a short distance downward as ie ing; upper cauline leaves much more e mp o ndric, 3-4 cm. long, s or densely clustered in twos to fours at the top of the nearly simple stems; severe glabrous, the Senge saga a a Satna ometimes a little scabrous and the margin freq uently ro ine near the margi spinescent, chartaceous, a little twisted; flowers whitish tinged with pur er corollas rise long, the tube very slender, 13-17 mm. long, the throat 5-8 mm. long, the lobes . long; achenes compressed or a little turgid, obovate-oblong, 3-4 mm. Pec 1.5 mm. wide, veteanh brown mottled w own us ligh 5 cm. ig Wet soil of coastal hills and flats, often og sare with willow owin the n with marsh plants, Upper — —— San Luis Obispo and Santa Barbara Cacgaien: ‘Ccltornis. ag focality: near La Graciosa, Santa Barbara Cou June-Aug. This rare thistle which was ce known until it was studied recently in the field > Clifton F. "Sete is closely related to the C. foliosum com 26. Cirsium folidsum (Hook.) DC. Leafy or Dwarf Thistle. Fig. 5927. Carduus besieta Hook. Fl. Bor. Amer. 1: 303. 1833. Cirsium foliosum DC. Prod. 6: 654. _ Cirsium Scmawide Torr. & Gray, Fl. N. Amer. 2: 459. 1843. ts Cirsium oo anum K. — um. in Just: Bot. mec 291: 566. 1903. Cirsi K = Toe. cit. Carduus ecuue Eastw. ex x Heller, Mublenbersia 2: 160. (Nomen nudum.) pe irsiu ssipodenanrmeaee i su bsp. lanatum Petrak, Beih, B. Ske .. 354. 1917. anatum var. oregonense oe rak, op. ¢ irstum i eaiienaaa sillenk latisquamum Petrak, op. cit. 358. nes drummondii subsp. vexans Petrak, op. cit. ey q et. var. citrinum Petrak, op. cit. 363. SUNFLOWER FAMILY 533 Plants — genres or short-lived perennials with a thick taproot, eee or with stems to 1 m. tall; often stout, more ter strongly striate, and hollow ; pubesc sparse to ee risp, elonga u s os obose, 3-5 cm. lon c bres the phyllaries = closely to rather loosely imbricate, the backs glabrous but often more or less scabrous, he margins a little crisp- -puberulent; ——— ni tom mentor, scabrous-ciiolat, or smooth, riangul Vv hort r la a lavender-pink oe rose, not curving outward, the corolla 2-3 cm. long, the tu —15 mm. long, the throat 5-10 mm. long, the lobes 4-10 mm. long; achenes light brown, 5-7 mm. long, 2-2.5 mm. wide, oblongish, a little widened upward; pappus pale brownish buff, 2-3 cm. lon ng. y or wet places in mountain meadows, sagebrush-covered valleys, open rancor and subalpine slopes, aistiien Canadian, and Hudsonian Zones; 7 Aaa the Cascade-Sierran re —_ to the east in. Washi ngton, Or fase and Ca ei i ya pe —— in the — Ranges; witearaad in the ntains and high & ae kon Rocky Mow tains. Type locality: reraiics vat the Rocky Moun aan ay—Sept. The lea e y thistles constitute a Cniyaiorpide postaine in ge of critical study before they can be satisfac- torily aligned on either specific or subspecific levels. 27. Cirsium ipesasi satin ae: Hooker Thistle. Fig. 5928. Carduus nae the oe Fl. Bor. Amer. 1: 302. Cirsium hookerianum Nutt. Trans. Amer. Phil. se = 7: 418. 1841. Pr et a short-lived perennial with a oot ; stems erect, 0.6-1.5(2) m. tall, nearly simple or narrowly branched above, thinly a yickmonk neater and sparsely crisp- -pubesce nt. Basal 5926. Cirsium loncholepis 5927. Cirsium foliosum * 534 COMPOSITAE leaves narrowly oblanceolate, to 3 dm. long or bk - os iis iv wide, pinnately partes or divided, the divisions mostly 2- ee with deltoid or ova ish s , the segm mg pig pped with stiffish yellow spines, the margin sparsely ss el tas. ‘oe below Bo ered, petiole-like base; cauline leaves similar nae the leaves eer ming shorte neat — ae sessile upward, the base ‘narrowly spiny-auriculate and more or less decurrent as narrow, spinose-pectinate hc below the point of attachment; the upper so leaves much reduced, very spiny, linear or eon ast the ——- st closely subtending the heads, the basal and cauline leaves thinly gray- a li : “ _ mt alon ei bresce ve sparse pir pu iebaicran: hea osely or subspicately arranged along the stem from the middle or above, becoming aut ‘clustered a ge top m 5, plant and oe occasionally at the ends of abbreviated lateral branches, subg ie g; involucre shortly campanulate, arachnoid and crisply puberulent “arachn oi :phyliaries fotser, loose, the outer and middle wide ly spreading and somewhat upw sin -cu above € appresse base, ovate-lanceolate, ending in a short, stiff, yellow spine, the inner nea phan into a slender, twisted, chartaceous, spinose tip, the margins citiclase and ane dous, the. outer phyllaries sometimes laterally spinescent, inter- grading in this character with the u permost bract-like, spiny leaves; flowers whitish or yellow- i in did ading in the heads; a € e ish, becom ordid-tan, somewhat spreading in the heads; corollas 2- on tub 2 . long, the t t 6-8 mm. long, th s 5-6 mm. long; achenes oblongish, a little nar- rowed downward, strongly flattened, 5-6 mm. long, 2 mm. wide; panes Moist slopes and flats of mountains, thriving in aoe and along roads, Canadian and Boreal Zones rthern Wa ibiaston vena Me the Concate. Mountains north to British aay reece? and Alberta Beare: to abe Rocky yi Gaisstngaty Rl ya or ty: BF A Ait oe of the Rocky Mountains.” July—Au ug. Acco g to Cro t_(Vasi Pe mee Northw. 955), specimens Bans of the Canadian border ‘‘vary i he vais ecti jou rs C. fol 28. Cirsium péckii Henderson. Steen Mountain Thistle. Fig. 5929. Cirsium peckti Henderson, Madrofio 5: 97. 1939. lowish green, si _ below, branching above the middle, the stem stout, fistulous, ae in dried specimens brownish or shining and golden, sparsely loose- — with elongate, shining, translu sate multicellular hairs, nd ogg below. Basal leaves unknown cauline ‘es narrowly oblanceolate or elliptic-oblanceolate, becom oblong- Sait eolate abov e, to 2.5 dm. long and 5 cm. wide, parted or divided into as deltoid lobed segments, the lobes tipped by set or Vygis yellow spines; aves eaves narrowed below to a narro wly bordered, e ate scatiehion pubes nt on the upper aad lower sides es oo se ara 8 and veins or the : s cm. long. isphe! ge ! eous and flattene e sie p straight, pale lavender, the corolla 2-3 cm. long, the tube 7-12 mm. long, the throat abou long, the lobes 7-8 mm. long; achen es light brown, Sapnagete a little whleaed upward, 6-7 mm long, ie mm. wide; pappus a little pe d, 2 cm. lon Moist soil and dry stream banks in sagebrush tam Tales Sonoran Zone; eastern ~— of the Steen and Pueblo Mountains, er County, Oregon. Type locality: Alvord Ranch, east base of Steen Mountains, Harney ppt June—Jul Cir: raetériens i. Macbride, Contr. Gray Herb. No. 53: 1918. Stems very leafy, arectnctd- faunaiiteas with a few pol (re Po shining, Nec Song hairs intermixed: cauline leaves oblong-elliptic, Dai ey into narrowly deltoid or linear, strongly spin eel pubese ent above get apace wl mt hai white-tomen ath i heads spi ole etree ag eo stem in the xia of the uppermost cat ine poem and Po mage subtended by —— reduced, bract- ike anon large, 5.5-7 cm. hi aly cs Hie invo mire rather den gray nulate-bow1- shaped; phyllaries loosely ascending, strai ak the outer an middle oie attenuate pe toa on tiff 5) spine, the atten the argin, The. inner flatten and somewhat curving and Svisied, eats scabridous below the slender. spi fer é tip; corolla whitish,” 3-3.5 em. long, the tube 1.5 cm. long, the throat 8 mm. long, the lobes 1 cm. long; pappus about m. long, light brown oe buff, copiously plumose from the base to the scarcely dilatate, barbellula te tip. Known nly from two collection made at Palo Alto, Santa Clara County, California, by J. W. Congdon in July, in 1897 and in 1901. 29. Cirsium callilépis (Greene) agattes Fringed-bract Thistle. Fig. 5930. Carduus callilepis ea eee Proc. Acad. Phila. 1892: 358 rae irsium callilepis Jepso: rip Mid. Calif. 507. 1901. var lilepis Jepson, Man. FI. Pl. Calif. 1164. 1925. Perennial herbs, aerial shoots arising from the see or branched, woody n of a slender or stout root, the new shoots developing from seeds om buds on the sits Rv re ading roots ; stems erect, generally rather slender, few-branched ate nt sely leafy, 2 dm. tall, a ; : 2 i or rather strongly striate-angled, crispy-puberulent with elongate, davies tlular, shining hai mi rachnoid. Earliest Be» small, subentire, prickly-ciliate, narrowly elliptic, the abre hnoi belo to 4 dm. long eep dm ns den weak, the blade narrowed to a more or less spiny- cere ‘petiole vtike base, this base mid- rib both abo below crispy-puberulent or arachnoid ; lowest cauline leaves senitar but ae a pom as —— auriculate base, the middle cauline facto ons semi-amplexicaul with a enlarged, rounded, spiny, shortly rrent base; uppermost leaves much reduced fee posi tikes ich subglobose or broadly papeniiet. 2-4 cm. long, solitary or clustered on elongate. SUNFLOWER FAMILY 535 slender, somewhat spreading, Ms ehacaith net jade eel Hh ey subglabrous or thinly arachnoid and hoe arly glabrescent ; phyllaries numerous in many series, a emaing appressed and well imbricate or (in var. pseu udocarlinoides) Jrequent y souskty prea deena less graduated, the outer and middle pli llaries narrowly. obov or Sblanbeolatesohovate Sith roundish or ovate, widely margined, r bee ine een- t ps dee dae - lower margins eS a ine cea att rulat mb the inner 0 bent oblong-lanceo- : i ul riilute-laterate 6 on the margins and is atpernaati tig a wea twit or curved, spine-like point ; flowers ee a > light brownish, corolla 2-2.5 cm. lon ng, t the 1 megs — ot 4-7 mm. long, the tube abou ; achenes rod rowly oblanceolate or nearly elliptic, 7 mm. long, thers brown; vada Tight beeen, sit 5-2 cm. long, the erie scarcely enlarged at the rr Grassy places on the edge of brush or woods, Humid Tra —_ on Beef eg fart skew Canal sada just north of a Erancisco Bay. Type echt: “Western Cali for and ‘ Mt. ais and northward” (FI. Fran. ral m remotifolium var. odontolépis vadage pe, Bot. Centralbl. 35: 298. 1917. Tips of the ae asennad % oe recurved. Known only from the hi yo collected on Mount Tamalpais, Marin County ‘irsium callilepis var. or onénse it 5. well, Leaflets West. Bot. a8 10. a (Cirsium Pattee Beih. pwn “Genial 35: 300. 1917.) ‘Phy llaries numer in ey series, subappressed, conspicuous “ool scarious or ceous, Foo uter shortly spinose at ‘he ema filamin cordate tip, the margins Picly fre riat te taewente oe to t = baee € the middle ee Race | HP gee dee with. ae et and expanded, gestions, Manette tips, the inner s light rose enanter, ovate-de a nginx! acu — or inulose; flowers pink. n forests of central Oregon, Thich in the Cascade hecntald Type | locality : mp Polk, Crook County. uly ut Cirsium callilepis var. potent (Petrak) J. well, loc. cit. (Cirsium remotifolium subsp. pseudocarlinoides Petrak, Beih. gad Pra ire 35: 297. Hots -Phyllaries oblance ry or oblong to oblong- linear, mostly loosely ascending, Paes r phyllaries entire and gradually narrowed to the spiny tip or frequently the upper part chartaceous and spars fe spinose-dentate the tip of the a sag scarious- or Tharencende: expanded, the margin deeply gaa irregularly lac bate Bes briate, the in phylla with scarious, curved sed twisted tips gs are serrulate- bape rong and spinulote- attenuate. rass' and and die of valleys and mou tains, mostly the Humid Transition Zone or Canadian Zone; from central California north to carthwestern Oregon. Type for ality: Mount "Tanialp ais Marin Count _ Wasmann Journ. Biol. lke € ; On Mount Tamalpais, ae hale been found that are perhaps perived from a cross with C. quercetorum. At Willits, Mendoc no Cou , a plant with the inner or ola glandular may ae a pba with C. cymosum. Cir mbly pa ak, Beih. Bot. Centralbl. 35:.312. 1917. simi lar to llilepis var. pseudo- carlinoides poh sath Oiytinsies appress sed and imbricate and with the , -tips less Goatiy scarious-margined ; a pres sus beet - tween C. callilepis var. pseudocarlinoides and C. etorum. Type locality: Mount Tamal- pais, Marin pot 30. Cirsium acanthodontum Blake. Klamath Thistle. Fig. 5931. Cirsium acanthodontum Blake, Contr. Gray Herb. No. 53: 28. 1918. pai sitions eset oped ee . Biol. Soc. Wash. 32: 43. 1919 rivulare Nels, Man. FI. PI. Calif. 1164. 1925. ee hort “he aerial paige arising singly or igi from the simple or somewhat che od n of the stout or slender taproot, cee owns developing from seedlings or produced along elongate te hneeate spreading roots; stems ie cag oe aoe. 0.5-1.1 m. tall (or dwarfed and Jess than 1 dm. tall), simple or few-bran ched abov more robust a Baki gan 536 COMPOSITAE crisp-pubescent or glabrescent. Earliest leaves small, pained elliptic- et entire or sparsely spinulose-denticulate, the leaves of the basal rosette usually few, varyin one sbi m — to deeply pinnately divided, narrowly to broadly chistes Le 54 bits subentire leaves 1.5—3 cm. broad, the lobed or Pap leaves 3-10 cm. broad, the lobes and divisions short and deltoid to elongate and again lobed or divided, the lo bes _ eee in slender wea spines, the leaf-margin more or less as al edt tow the narrowed, pctinke fie base; lower cauline leaves similar to the mee pews but the ae blade-like part extending t ec a h u ing, the inner tenis sia si inear-oblong, the sides smooth or spinulose- eee, the tip more or less chartaceous-scarious and lacerate, triangular-acute or sometimes flabellate, the median line of the inner silanes eiaiidndar ; corollas slgeite tapings: 2 me the Bead, rosy-purple or lavender- the 5m ipti Moist deep soil of woods and meadow borders, or on rocky cuts and exposed maritime slopes, Humid Transi- tion Zane: Coast Ranges from Curry County, ae south to oe County, California. Type locality: “dry rocky soil, 6.4 km. north of Agness, Curry Co.,’’ Oregon. June—Sept. 31. Cirsium remotifdlium (Hook.) DC. Remote-leaved Thistle. Fig. 5932. Corduns remotsfolius Hook. Fl. Bor. Amer. 1: 302. 1833. Prod. &: 655. 1837. Cirsium stenolepidum Nutt. Trans. Amer. Phil. Soc. II. 7: 419. 1841. Biennial or short-lived Bohtipas: ¥ her se with a rather stout taproot (or perhaps with ne rosettes pacts a izome-like ts) ; stems erect, 0.3-1.5 m. tall, remotely iy rather closely cee terete but somewhat longitudinally pray purplish- tinged above poopeers with oe S ae ate, multicellular, alliage sed hairs, also thinly arachno id a cely the base. Basal leaves an es na long and 1 dm. wide, narrowly elliptic, deeply ete 8 d or parted, the segments br val obongish and genera ally 1-lobed on the outer side, the segments an lobes shortly and weakly s tipped, the margin spa a spinulose ciate, chen. arachion tomentose below with ainin a Sanltcetinies baits along midrib and veins, gla brate above with scattered, abenicig 1 adie lular hairs, the blade narrowed graduall a a od su a te phe ae a € at the ends . short, Lagat or sparsely leafy-bracted, peduncle-like stems or rarely some 0 the nvolucre oi ho i e gi L below the tip, the inner phyllaries attenuate into a thin, slender, somewhat twisted, spinescent tip, the margins spinulose-ciliate, or the fe a little expanded and Grate. fringed; flowers —— sordid, the corollas 2 cm. long, the mm. long, the lobes a ual, mm. long; achen dark or light brown speckled with aoe eee sh brown, scarcely sh ing, 4-5 mm. long, reams cuneate ; Dappite 2 cm. long, sordid-whitish, the bar bellulate tips oor exp ande d. Ran Type locality: ee = Pie valley of ie Columbia.” i? har “Almost cvecywherd this this 4 et nee to r i n. ybrid-suspect between C. pacciee Sec and C. pons hes been collected on Sexton Mountain, Josephine yanigg =r It resembles C. semchihann but it is more tomentose and many of the phyllaries are glandular ong the midr Cirs: mendocinum whol Beih. Bot: Centralbl. 35: 339. 1917. Sim o C. remotifolium but the leaves much a spiny, the involucres more copiously arachnoid, and the flowers ara ve urplish due either = = corolla or colored anther- nr a ese characters are reminiscent of C. andrewsii. rushy and wooded places on the coastal pind of Mendocino County, California, the type locality 82. ——— andersonii (A. Gray) Petrak. Anderson’s Thistle. Fig. 5933. C: A. Gray, Proc. Amer. Acad. 10: 44. 1874. Cirsium andersonii Petrak, Bot. Tidsskr. 31: 68. 1911. Perennial sete with widely ee Sagi ——— roots from which flowering plants with stout taproots, woody and so ee - — ces, and erect stems develop; stems usually 4-7 dm retry rarel ees crane eal ai eS = much as 10 dm. tall, sparsely crisp puberulent and arachnoid-tomentose or pron atte Paws purplish. Leaves of the first season few, small, obl olate, obtuse or subacute, spin dentate some undulate and spiny the margin, in s (or perh after several years?) developing a large rosette from which the flowering sh or s 3 m. long and m. wide, oblong- gro osulate leave: 0:3: oblanceolate, pinnately divided with the divisions again lobed, rather sparsely and weakly spiny particularly on the acute tips of the lobes, the bl nd narrowed below to a petiole- like base; caul- SUNFLOWER FAMILY 537 ine leaves frequently rather sparse, oblanceolate or oblongish, to 2 dm. long, pinnately lobed o divided into 1-3-parted, ovatish lobes or divisions or in depauperate plants the cauline sts m and ¢g . palate, G. 5)4-5(6 long, closely subtende y fe ew or several reduced leaves with spiny marg r rarely the subeoridine feives foliaceous ; favéliere thinly floccose-tomentose or glabres- ene 35 cm. long; phyllaries numerous, appressed or somewhat spreading, the outer narrowly ovate to lanceolate, the middle aaeh tant late, both the outer and nde: phyllaries tipped by sp ir h atten e mutico r spinescent tips, the sree: part of the inner phyllaries — scabrous-pubescent and pupal: eokotias purplish rose, 3-4.5 cm. long, the lobes about 1 cm. long, the throat 1-1.6 cm. long achenes strongly flattened, oblong-cuneate, 6-7 mm. long, Cowh: pappus sordid-white, 2.5-4 long. n gers forests and dry meadows, Arid Transition Zone to Hudsonian Zone, 2,700-10,000 feet; One mountains of a California mt Boe urd ihrouah the Sierra Nevada st sage ees southwestern Tdaho and western Nevad Type locality: “Sierra Nevada, California, and adjacent part of Nevada.’’ June—Oct = — ee ee —— ——— ————— —S y Za ——— SS i eed : anutet Willaty Or 7 oh Sae ay vi a , sol ah 5930. Cirsium — 5932. Cirsium remotifolium 5931. Cirsium acan 933. Cirsi dersonii Ca) 538 COMPOSITAE 33. Cirsium campylon H. K. Sharsmith. Hamilton Thistle. Fig. 5934. Cirsium campylon H. K. Sharsmith, Madrofio 5: 85. 1939. Stout, succulent, perennial herb with a thick woody root-crown; stems erect, thick, hollow, 0.3-2.1 m. tall, greenish or sometimes purplish-tinged, arachnoid- tom entose and also sparsely glandular-puberulent, the to mentum mostly deciduous. Earliest leaves small, thin, oblanceolate, undulate-lobed and spinulose- sa st subgla brous; later basal leaves forming a small or large, up to7d 2 i ose rosette lan n dm parted or divided into broad ae peasiente, the lobes ending ina aig bogs: spine and ne a i with thin ar id tom sisting and sometim to an elongate, more or less spiny-bordered petiole; cauline leaves smaller, oblanceolate or more fre requently oblongish or lanceolate, the petiole-like base spiny-lobed to the bottom or the cauline leaves bro adly sessile with semi-amplexicaul base, the basal lobes more or less decurrent and strongly spiny ; uppermost leaves becoming reduced and bract-like but not a sphtending the heads; it ng solitary or wiser eee Samed at the ends of ea ayes eads or cluster nodding, the heads broader than long, 2.5-3.5 cm. long an aati coe prose or more or less ged ith ~ aoe ped buberulent tnct not arachnoid, age outer strongly recurved, narrowly ovate, e ply acute o tip; flowers sordid-white or somewhat pinkish-tinged, becoming sordid-brownish in a e, the corolla about 2 cm. long, the tube 5-6 mm. long, the throat about 10 mm. long, the lobes more or less unequal, 4-5 mm. long; achenes oblong-cune ate, 4-5 mm. long, light to eek brown; pappus rather sparse, light brownish, i oi 1.be Restricted to the Mount Hamilton Range a the South Coast Ranges of California in Stanislaus and Santa Clara a e middle. nutans. SUNFLOWER FAMILY 539 Plants annual; heads erect, densely clustered, rarely solitary, cylindric, 1.5-2.5 cm. long, 1-1.5 cm. wide, rather ew w-flowered; phyllaries spreading slightly near the tip. a Phyllaries more or less scarious-margined, the tips glabrous and smooth except on the ap ape margin. richom uiflorus. oe . C. pycenocephalus. 2. rina 0° ea rian arena -margined, the tips roughened on the margin and back by sient caus a 1. Carduus nitans L. Musk Thistle. Fig. 5936 Carduus nutans L. Sp. Pl. 821. 1753. ants pinay thinly t poneitiees or glabrous elongate branches abov s; stems =p Sr the base, the lower leafy win 1.5 (or 2) m. tall, simple or Po mre 3 winged, the peduncle g& 5934. Cirsium campylon 5935. Cirsium fontinale . Carduus nutans 5937. Carduus tenuiflorus 40 540 COMPOSITAE bract-like, the basal leaves short-petiolate, the cauline sessile ane decurrent in the spiny wings; heads soli tary. modding. | oe mo ostly 3-5 cm. long and 4-8 cm. wide; involucre hemispheric, th ate, scarcely spinose, purplish; flowers purple; achenes obovate, 3-4 mm. long, rina wr rinkled, bie ish; pappus- -bristles numerous, whitish, 1.5-2.5 cm. long. Rar d local weed along roads and in grassy places; Whatcom and wii tman Counties, Washin on ballast, ‘Linnton, pyre mah County, beer Walnut, Los Angeles Courity: California; occasional ene ia to British — a (Alexis Creek) a nd eastward to the Atl antic; Argentina. Native of the Old World. May-Sept. Carduus acant catia ~ as. ‘Pl. on. pobieae Plant biennial, the erect — spinose-w eo leave brescent, petrol margined; hea late or hemispheric, solitary or few-clustered, 1.5-2. os, eae and octane mar tie 7 phyllaries ae or linear- lanceolate, the outer and middle at “spreading or machiaal. short- spino orolla reddish purple, nearly 2 c ong; achene oblong- eevee Pass ng, brown, ob- arciy ieneliadinatty nerved and finely transversely rugulose; pappus whitish, cy. : ng. Ne ee Lake Waha, Nez Perce County, western Idaho; occasional east to the Atlantic; Argentina. Native a Europ 2. a tenuiflorus Curtis. Slender-flowered Thistle. Fig. 5937. Carduus tenuiflorus Curtis, Fl. Lond. Fasc. 6, pl. 55. 1790-98. Carduus neglectus of pen authors, in part. Not Tenore. Plants annual, more or less tomentose, the upper side of the leaves glabrescent; stems atstt, 3-20 dm. tall, aetowly to broadly spiny-winged, simple or fastigiately few-branched from the base. Leaves Aaa ey to elliptic or obovate, to 1.5 dm. long an cm. wide, the banat shortly petioled and sometimes only gis lly EA Re apantine i and decurrent from the n J i pa d in e gt al and paw ak eure: at the ends of st and branche oP fre sy more than 5 in a cluster ; phyl- laries numerous in several series, tay Se lo oosely imbricate, the outer and middle ovate to lanceo- r sordid-whitish, 1-1. 5 cm ssy slopes, open ey and r Coast Ranges from_ notte County, Oregon, to central paeriavi, seuainete in the Sierra Nevada foothills — poe Pees toR ; Texas. Native ‘of Europe. April—Aug in rm he corolla f as the c bia rece sched oblong, 4-5 mm. long, brownish, usually finely 10-13- pie tenet . long. 3. Carduus pycnocéphalus L. Italian Thistle. Fig. 5938. Carduus pycnocephalus L. Sp. Pl. ed. 21151. 1763. Plants annual, gray-green to white-tomentose, the upper side of the leaves glabrescent, scabrid ; stems erect, 1.5-10 (or 20) dm. tall, simple or few- branched, rather narrowly spiny-winged, the wing-spines Svaticaie toctnmtal: often ong cae rigid. Leaves oblanceolate or lanceolate to oblongish and rarely ovate, to 15 cm. lon —_ cm. wide, pinnately parted into spiny-toothed or -lo segments, the terminal spine of the segments and s the most prominent and rigid, the basal and lowest cauline leaves short- seticlate. the middle cauline leaves narrowly cuneate, the upper So heads cylindric, 2-2.5 cm. long, gene rally 2-5-clustered at the ends of stems and branches, or metimes solitary ; chyllties eatin in several series, rather loosely imbri- n scab rt ic midrib, the inner phyllaries Snes, less rigid and nearly innocuous; flowers reddish purple, the corolla-lobes about three times as long as the corolla-throat ; achenes oblong, buff or pale brown, shining, 6 mm. ppu 2 cm. long. On grassy flats and slopes and along ro: oads; widespread and common in the San Francisco Bay area, Cali- fornia; local in Lane County, ore n, and in Eldorado and San Diego Counties, California. Native of the Mediterranean region. April—June 144. CRNTOURRA L. Sp. Pl. 909. 1753. s graduated, the innermost frequently shortened and connivent, or the innermost occasionally longest. [An ancient Greek name, signifying the centaurs’ plant, of obscure application] A genus of about 400 species, found largely in the Mediterranean r but occ rides also in North a South America and Australia. All of ours are introduced. Type species, 'aaeares gone ide m L. SUNFLOWER FAMILY 541 Flowers purple, blue, pink, or white; stems more or less angled but not winged. Pangea numerous, _subplumose, deciduous, the cc puagwraeel the y's as es to 1 * waar flowers per- fect; phy pectinately fringed . C. repens. ig depres if Leese spree frequently paleaceous, : mm. Aisle or Seba: eed de in head Sterile, r Phyllaries with a dria’ or short terminal spin Heads 2-3 ong; involu ar spines snes and eosinengga —3 cm. long; fl pink to p Paseul & nneingt the paleaceous bristles 1-2 mm. lon 2. C. sber: Pappus none. pe ka pore Heads 1-1.5 cm. nee, oe seen aeres involucral spines short and oe oy a long; flowers white or rarely Phyllaries not spine- Nanad or, if middle termi a pond division of the appendage is viata ‘ai ieaiat, then the spinule shorter Pt ‘ie lateral “Grice Leaves pinnately to bipinnately parted or divided; pice pink; plants biennial or perennial. 5. C. maculosa. Leaves entire or with a few salient lobes ~ = flowers blue or purple, rarely white or pink. Plan s annual; leaf-blades ig 08 ear to narrowly oblanceolate, 1 cm. wide or less; Letina well developed, to aps OF Plants perennial; leaves pcan or elliptic, few-lobed ra ries to 3 cm. wide; “papeus generally poorly developed or none, if present 1 mm. long or les: piacere - middle phyllaries broad and rounded, scarious or lacerate, not pectinate, vering the base of ip phy llaries; sterile marginal flowers coletee aor none - por re sietted by small si A ope Appendages of middle plas pestinate fringed; pappus usually p Z Involucre as wide endage of _ egg A dagge bea bre completely covering the base of the pet Doe the "ie ciniae se se, mostly once to three times the width of the undivided body of the phyllary Appendages id phyllaries light to dark beolir, the laciniae about as long as the undivided body PF the phyllary; sterile marginal watt Pool a. ey enla lla Appieisdglics of snstlariae blackish, the laciniae two three times o width of the body of the phyllary; sterile marginal Pethestts lacking. 9. C. nigra joudece cylindric, longer than wide; appendage of middle poe he ke gall, not en- rely covering the base of the jy bee sterile ma ou enlarged; oapiiik reduced t Apnaiivts scales Flowers yellow, ~ outermost eure stems. win the an pte eal fase phyllaries weaning salina -tipped. Heads about 1.5 cm. long; spines Fy ie "olay acts s 0.5-1 cm. long, slender, purplish- wa Kt i nish- ti cok Fiat on the ap 11. C. mel Heads about 2 cm. long; spines on oo cage phyllaries 1-2 cm. long, stout, yellow, terete. 12. C. solstitialis. 1. Centaurea répens L. Russian Knapweed or Turkestan Thistle. Fig. 5939. Centaurea repens L. we lLede2. ng 3: Centaurea picris Pall. ex Wil ld. Sp. Pi. 3: 2302. 1804. Acroptilon picris Boiss. Fl. Orien. 3: er: 1875. Plants perennial with widely anrending: deep-seated rootstocks from which the aerial chee arise; stems erect, openly branched, leafy, terete, wingless, 2-10 dm. ta ll. Leaves thinly tomen- tulose or glabrate, pale green or canescent, scabrous-margined, the basal leaves sacle shlong” oblanceolate, to 8 cm. long and 2 .5 cm. wi = acute, deeply and seeularty pinnately lobed or divided, the i nt again lobed or only serrate, acute, petiole shorter than the blade, the lower cauline leave what ener pinnately ied 6 r divided or saliently serrate, the upper leaves narrowly oxigtag: 13 cm. long, 2-7 mm. wide, maceiwed to a sessile base, abruptly acute and 5938. Carduus pycnocephalus AW 542 COMPOSITAE ucronate, entire or serrate; heads solitary at the ends of the branches or sometimes a few in loot cluster, ovate, 1.5- cm. igh; involucres thinly tomentulose or glabrous, the outer aia middle phyllaries with a rounded herbaceous part capped by a broad fe aline appendage, oy ap- pendage obtuse or acute, thinly pilose and ciliate, the inner phylaries becoming narrower and onger, oblong or lanceolate, the appendage Taite and acu acuminat if the in pe i nd penic ; m a raps oblongish, somewhat compressed or obcompressed, ivory-w ite; p pappus- -bristles o 1 cm. long, numerous, early deciduous, the outer shorter in more slender, the inner longer and fnanouiie a Tittle paleaceou at sien all more or less plumos Cultivat ed fields, pastures, and roadsides, widespread and locally common at lower elevations and in warmer drier parts; Washington so sithern, Caton British Columbia; east to the central United States; Argentina. Native in athers Russia and central Asia. May—Oct 2. Centaurea ibérica Trev. Iberian Star-thistle. Fig. 5940. Centaurea iberica Trev. in Spreng. Syst. 3: 406. 1826. Plants biennial _ a — 0 me stems —_ 5-10 dm. high or more, much branched and bushy from below, terete or a little angled. Leaves pu ebenuil ent or thinly tomentulose, the upper somewhat eaberisiss: the ois 1 ives Teautite ebladetotate o 4.5 dm. long, once or twice deeply lobed or divided, the segments oblong to elliptic, obtuse or oa mucronulate-serrulate or entire, i auline le & ivi -lo heads sessile or. shortly pedunculate, ovate or sialeccan et a cm. 008 involucres glabrous, the phyllaries nerveless, very stiff-coriaceous, pale with n white-scarious margins, the outer phyllaries small and nearly oe eiie the middle | phyllaries sibddnced into a rigid, spre pe ing, i gant spine 1-2.5 cm. long, the spine channeled on the upper side and bearing 1 3 pai of small spines near ‘the base, the co phyllaries sachet tipped by a broad, “denticula te or lacerate, sratioas appendage; flow Achner or purplish, the outer not enlarged; achene oblong, mm. long; pappus ‘res sent, ae ‘brist es flattened and in several series, the outer short, the inner longer and narrower and about half the length of the achene An uncommon weed, occurring my abe in o Sear Santa Barbara, and Sie Diego Counties, California. Native of southern Europe and western Asia. July—Sep 3. Centaurea calcitrapa L. Purple Star-thistle. Fig. 5941. Centaurea “yep L. Sp. Fi. 917: 1752. ’ € . oer: about 1 dm. Ion cm. wide, aig or twice- < ae age rip ee oblongish, n shor he saline leaves short-petiolate or sessile, decurrent, danoky. peace divided into oblong- linear segments, the uppermost leaves sessile, easire-or ate or serrate-lobed, the apex and teeth m or ious margins, the outer phyllaries with a short rt spine, the middle —— melger ig ed into a stout, spreading, stramineous spine 1.5-3 cm. long, the Si a channele r side, bearing 2 or 3 pairs of short, stiff, lateral spines at the sole the inn , asa. Raat q small spine or a spineless sido ee Med owers purple, the outer ears enlarged; achenes subobovate, re ease m. long; pappus C, and roadsides, common middle western California; Humboldt County to aug Clara gear Capaetare iheniins g, Washington; = ballast, Linnton, Oregon; Nanaimo, Vancouver Island; eastern United States. Native in the 5, World from southern Sweden to North Africa and central Russia. ay—Nov. umboldt_and Riv alifornia, a variant of C. calcitrapa is found in which the beads are spines more slen : and shorter, and the inner achenes (which are fertile) pappus-bearing.- itrapoides Ait. j ‘ C261. 1 feo" 'P Plants annual (on perennial?), giebeoss oe thinly nach a “an bove; stems to 2 m. _ fog ; lower cauline L.. lyrate-pinnatifid, cm. long and 10 cm. wide, sessile hi i tly # Gea hase, the vg ae og irregularly epic all lobed or parted, the nf 98 ong; t ermost ir lave outer sterile corollas much oe gh achenes buff, 3-3.5 mm. lon s of graduated white pee to ees r long, none on outer sterile achenes. Locally established in central aa gra California; San put: anta Barbara, and in Los Angeles ited hg near Watts and Whittier. Native of North Africa. May ; ; ; * 1837.) nual; s and pedeneics ic ia p _lower leaves innately divided a parted, eet og ut : s phyllaries appresse ie epermn tae “event cry ovate, Nena narrowed be a yh vr Ped or reflexed spin e, the some rather erent eh yellow iu, 2-5 mm. jong, the base of the spine and margin of ~ ited came. the outer sterile and radiately much enlarged; achenes poe t “¢ 5m _ ong, phen 5 sh br rown, finely pitte ween the longitudinal ribs, the ise of oe achene and margin of the aitecbenent- scar smooth, bony, ivory; Aang +08 : bristles paleaceous, graduated i in rng to 3 mm. long, pale brownish. An established garden escape in Sa sj ie California, since 1904. Native in the western part of the Mediterranean region. April—Jun urea moschata T.. Sp. PL. 909. 1753. (Amberboa forage DC. Prod, 6: 560. 1837.) Plants poms ; y rounded Dn witha wie of 100 spreading, appendage; ee f, thinly oil purplish, or whitish, the outer a a much enlarged; achenes nearly smooth, brownish blac thinly mm. long; pigpiee bitten paleaceous, the innermost the hengeat: and 3 mm. lo f garden yg sometimes becoming established in waste ground; native in southwestern ‘acie. lao Se Sweet Sultan. SUNFLOWER FAMILY 543 4. Centaurea diffiisa Lam. Tumble Knapweed. Fig. 5942. Centaurea diffusa Lam. Encycl. 1: 675. 1783. lants biennial with an ny ih taproot, asi Sie or pale green, pubescent or thinly floccose- stenoses becoming glabrate, scabrous; stems erect, (1 or) dm. tall, angled but not winged, ranched near or abov he branc ne poeta and intr nese Basal leaves rosulate, i i d wide i ster] : sn or bla ckis h “tetany striate with several conspicuous or faint, pale or ivory lines; ee none or on the inner achenes represented by white paleaceous scales less than 1 mm . lon Fields, roadsides, and waste ground, locally common; Washington and northern fea east of the Cascade Mountains; Tdaho and southern British sey occasional in central and eastern United States. Native in southeastern ican and western Asia. June—Sep \ WE gs Wi tre A 5940. Centaurea ibe: 5941. Centaurea ak . Centaurea diffusa 5943. Centaurea maculosa ~ 544 COMPOSITAE Centaurea virgata var. squarrésa Boiss. Fl, Orien. bi Ae 1875. (Centaurea squarrosa Willd. Sp. Pl. 3: 2319. 1804.) Plants Liab ont — Sane from the crown of a stout taproot; stems about 5 dm. tall, branch: ing widely above, scabro low S pinnate or eee ia se wie, mg SERRE, long- petioled, the middl and upper leaves sessile, pin mene or pita “iy Sa revolute; heads small, -3 cm. long; involucres vat - about 7 mm. = tote green and preted Pet a little violet-tinged, the middie whollatieg pri ab fringed with short spines cen the sides below the terminal squarrose spine which is 1—2(or 3) mm. long, the innermost phyl- _—— eb agai or nea arly > the Pd narrowed and hyaline; flowers pinkish or lavender, the eee ones sterile and omewhat enlarged; achen -5-3.5 mm. long; pappus- -bristles slender white, to 2.5 mm. long. Occurring locally n County, Califo ornia, and in adjacent gore co? bey aeae Utah. Native of middle eastern Europe to petry Asia pe eae, July—Aug. In typical C. vir the terminal spine of the involucral bracts is shorter or even lackin 5. Centaurea maculésa Lam. Spotted Knapweed. Fig: 5943. Centaurea maculosa Lam. Encycl. 1: 669. 1783. Biennial or short-lived perennial with 1 to several stems arising from the top of a stout tap- root, ane or pale green, arachnoid-tomentulose or glabrate, scabrous; stems erect, 2-8 (or 15) dm. tal , angled but not winged, branching near the middle or above, the branches not very numer- us a e low oO s pink, rarely white, = ; yo 3m g, bro blacki sath several faint pale pi gig lines ; pappus present, i Sa iy salad cant bristles tae in length, the longest 2-3 mm. lon Fields, —— and waste ground; 1 Oregon east of the Cascade Mountains north through weste post ome va Washington to British edt hin Tdaho and Montana; widespread in central and eastern United States and Canada. Native in Europe and western Siberia. June-—Oct Cask taurea cineraria L. tek . 912. 1753. Plants perennial, bushy, stems and 1 s white- tomentose or pale green. All | ves except the youngest an a ‘uppermost bipinnate or bipinnately pets ded, the oe narrowly oblongish ne) obtuse, the uppermost leaves seg eit divided or set Sb He rather large, 2.5 cn ome. teers 8 arranged; Sng apg d seb 2 vate, about 1.5 cm. long, the phyllaries pale green except for the hea ac or ig sh margin and a pa pressed, the aseue margins oechinate: Seiliite, the terminal seta stiff and subspin shorter than the Metered” aoige and. less than 1 mm. long, squarrose, the innermost ohyE laries with a deneculate scarious tip; flowers purple, t terile outer corollas enlar ed; achenes 3 mm. long; pap} cf Pa ua numerous whitish bristles 4 mm. long. el ally spontaneous near eardenk in central and southern Calif. Native of southern Europe and North Africa, ay: Dusty Miller. Cen entaurea salmantica L. Sp. Pl. 918. 53. (Microlonchus salmanticu 1837.) Plants perennial. Lower leaves pinnate, ‘the lateral ean ets few and rather remote, i and ru uncinately pinnatifid oe aot arany Bose! Ra ularly caine, scabrid-pubescent, the upper leaves Bs sm alles than the dower, pinnatifid ; heads 2— "2. m. long; involucre globose-conic, strongly constricted at the top, 1-2 cm. long, the shyliarics amet , triangular. rete: subglabrous, pale yellow-green, blacki hs at the PR aes by a Thaet spinose mucro about 0. Poy long or ag t i ; flowers op h or whit atkenes about 3. 5 mm. long, light brown with vertically arranged: _ tran dark seal areolae, the name of the achene and ial = the smooth, bony, and yellowish; pappus- Lt gourys LS weep aca . long except the inner: ngle, a pie acuminate bristle ahh ch is 3.5 mm. long. dere. Son soma fone Calitornia. wre Sg it has not — ee collected; dooryard weed, Jerome, shading Pgaidsb of the Mediterranean region. April-Sept. Escobi 6. Centaurea cyanus L. Cornflower or Bachelor’s Button. Fig. 5944. Centaurea cyanus L, Sp. Pl, 911. Centaurea cyanus var. denudata Suskd. Schiele vs 43. 1927, lants eres "ong occose- secoentnne usually thinly so, rarely almost re : little —— 3 stems erect, 1-7 dm. tall, simple or more commonly openly branched, the stems and bran slender, terete or s conenitiae ed, not winged. Basal and lower cauline sg ee e linear, prominently pinnately few-lobed or lyrate-pinnatifid, short-petioled, to 1 ng = 1 cm. wide, the and upper cauline leaves narrowly oblanceolate or linear, sessile, dentate entire, sharply acute or acuminate, the 1 side arachnoid-to se, a ve inly branches, or € or glabrate, the margin revolute; heads solitary at the ends of slender, pedun 1 oss ; involucre ovate or oblong-ovate, thinly tomentulese, t the cm. long, about 3 cm. acr phy laries coriaceous, spineless, the outer and middle duet with t Sues margins, the margin pectinately y parted into sharp tr a = lobes and forming a i serrate-parted appendage at the tip, the innermost phyllaries oblong, entire except at the m r les serrate-lacerate tip; flowers blue varying t thi nae 54 rple, the outermost sterile flowers with much elongated, palmately i mi corollas, the ray-like limb about : e the funnelform throat ; achenes 4 mm. | ye ad ete nin brown or buff, thinly pubescent above, densely so at the very ba ppu ait 4 mm. long, con i 4 of numerous, ood graduated, brown bristles, nar- rowy oblong, Saree aaa a little pie brushy hills and valleys as well as in fields and along roads; widespread and naturalized in Wash scaeen’ ie ring and ph rage aa i fornia, occasional southward to southern Californie as an escape from m cultivation; deer Colemabie? Fike & tlantic coast. Native in the Mediterranean region; introduced in many parts SUNFLOWER FAMILY 545 7. Centaurea jacea L. Brown Knapweed. Fig. 5945. Centaurea jacea L. Sp. Pl. 914. 1753 oe the stems eae or Ferree from a woody root- sagdviainn pale green, thinly mostly 5-10 dm and pubescent, becoming scp: sabe stems erect, veniane near the mid le or atoee, the Srenthes rather r few and ate. -B: and lower cag = s oe sparse, oblanceolate or aural elliptic, the blade - — — oe and 3 cm. wide, entire or coarsely few-lobed, pubescent above below, scabrous s, long- ine salts. Pa betiole slender, nearly as long as the blade or longer, the adie! cantina: eves lanceolate, oblon oblanceolate, entire, serrulate or shallowly lobed, Sia the uppermost leaves similar ae smalle hig a solitary at the ends of the branches; invo i i b the phy tip, entire to e; flowers rose to purple, rarely white, the outermost corollas enla nspicuous ; parece oblongish, 3-3.5 mm. long, light brown with a few mega lennttoeeal lines ; pappus none or represented by a few very short scales less than 0.5 mm. lon Se be Scene = nh idee and along roads; Washington and Oregon, chiefly west of the Cascade Mountains; east to the Atlantic. Naturalized from Europe. j une—Aug. a ue Ui REN M ws y 5944. Centaurea cyanus 5946. PORRRE e selina 5945. Centaurea jacea 5947. Centaurea nigra 546 COMPOSITAE = Centaurea preen Thuill. Protean Knapweed. Fig. 5946. Cent Thuill. Fl. Paris “nk 2.4 1799, yee nigra var. radiata DC. Fl. Fra ae ak 1815. lants similar to C. jacea; ‘ieoloo broadly elliptic or subglobose, about 1.5 cm. long and as wide or wider, the appendag es - the 1 Renan — o dark brown, roundish with broad, pectinately frin nge ed margins, the laciniae filifor Deas anaiadsees more or less setulose, ut as lon ad i the width of the essere of the Hote hue the appendages completely covering the base of ox le phyllaries; marginal flowers with radiately vp orollas, sterile; pappus mere peseesentl by a few or numerous unequal paleae, mostly 0.5 mm. long or less, rarely obso * por and roadsides, widespread and common particularly at lower elevations west of the Cascade Mountain Washington to northwestern California: British Columbia; eastern United States. Cultivated around Raters, may account for i “Pie — yaa it is hercrees specific recognition because in our area it is by far the most 0 e Y= 2 S This plant, introduced. from Europe, is generally regarded as a hybrid of C, age and C, nigra, a fact which t ‘ & 9. Centaurea nigra L. Black Knapweed. Fig. 5947. Centaurea nigra L. Sp. Pl. 911. Centaurea jacea var. nigra Briq. in Heal. Fl. Mitt.-Eur. 6: 954. 1929. similar to C. jacea; involucre roundish-cupshaped, about 1.5 cm. long and as wide as a - Occa oo in weedy places and in fields; i ea and northwestern Oregon; adventive in central Cali- fornia; caters United States. Introduced from Europe. J canes pt. 10. Copetete nigréscens Willd. Short-fringed Knapweed. Fig. 5948. Centaurea pratbeage oto d. Sp. Pl. 3: 2288. 1804. Centaurea dubia Suter, Fl. Helvet. 2: 202. 1802. Not Gmelin. Plants ari to C. jacea; cauline leaves lanceolate, oe Sate at base, sessile; involucre broadly oblong or cylindric, longer than broad, about 1.5 cm. lon , the appendages of t the phy 1- laries smaller, not covering the greenish or straw-colored Sosea. ‘triangular, pectinate-fringed, h t i ile margina the segments a the width of the undivided part of the phyllaries; sterile mar | flowers generally present and radiately aia pappus reduced to sma ne. Widespread but rather rare, ruderal - font ral; Washington and northern Oregon; Idaho; eastern United States. Introduced from Europe. June—Sep 11. Centaurea meliténsis L. Napa Thistle or Tocalote. Fig. 5949. t itensis L. Sp. Pl. 917. 1783. and leaves sessile and decurrent, narrowly oblong or oblong-oblanceolate, up to 5 cm. long a e, margin entire or gen eeatly: more or less dentate, the u ec leaves small but not ut 1.5 cm. s te) ong triangular, LHe gs ap) eons e; flowers yellow, numerous, the outermost with filiform lobes and inconspicuous ; ac 75 mm. lo ong, light es or buff, ‘all ith pappus ; pappus- bristles to : mm. long, slen E sof British Columbia, Washington, northern Oregon, and Lower Cali fornia; occasional east to the pec 2d coast; Hawaiian otra South America; South Africa; Australia. Native in the Mediterranean region. April-Sept. 12. Centaurea solstitialis L. Barnaby’s Thistle or Yellow Star-thistle. Fig. 5950. Centaurea solstitialis L. Sp. Pl. 917. 1753. Plants annual, es arachnoid-tomentose and scabrid, generally 3-10 dm. tall; stems erect, openly branched from near the base, the branches stiff and somewhat sal leave winged by the de- ti Ss! id. 6 t shorter than the blade, the cauline i“ sessile and decurren t, line near i it. nae Me as muc 5 _as 10 cm. long and 5 mm. wide near the base but much johcces and even scale-like on the uppermost SUNFLOWER FAMILY 547 branchlets, entire, frequently undulate; heads solitary at the ends of the branchlets, ovate, about 2 cm. long; involucres thinly to mentulose, early glabrescent, the phyllaries nerveless, coriaceous, the lowest phyllaries tipped by short, weak, gees tely disposed spines, the middle phyllaries bear- ing as stiff, erete, yellow, divergent, ter retical spine 1-2 cm. long and 1 or 2 pairs of ver shor lateral spines at the base of the terminal spine, ft innermost phyllaries expande snc nf are denticulate or lacerate, obovate appendage, spineless or nearly so; flowers yellow the outermost with filiform lobes and inconspicuous; achenes 2.5-3 mm. rte his ating: either ‘blackish brown or light brown and mottled, some of the outer without pet or with poorly developed pappus het inner with numerous, fine, white pappus-bristles to 3 mm. long. hodiisldes, fields, and waste ground about ect very oo and Cainaia from southern Oregon to central California at lower betel ng less common rare in Lower California, southern California, northern Oregon, and souther aly eastern Washington; peame g te and local in nahe central and eastern United States. Native in southern ie rope and the Mediterranean region. June—Nov Centaurea sashptes Willd. Enum. Hort. Ber. 930. _ 1808. bh dmg og of California authors. Not L.) Stems conspicuously ler a Lower leaves coarsely _pinnately parted or divi By. or the lower and upper leaves ‘ p pe ie zontal - tinged, Lage gel ong the central gee of the middle phyllaries 2-2.5 cm. long, the innermost phyllaries nar- rowed toa y acute or slightly spinose tip; flowers sige achenes 5 mm. long with numerous, strongly gradu- ated oe pes) fs nity bristles to 1 ps long. Locally stablished near Folsom, Sacramento Geuake, alifornia. Nativ: one western part of the eee ee region Was —Jun pra ea eriéphor So. (P1916. 1753. Stems urea esrect win Lower leaves pinnately parted or divided, the apeebaed entire or dentate, abruptly acute or mucronate; own veer "eek, abou em, high; in- volucre conspicuously loosely baal sie He entose, about 2 cm. long, broadly ovate or rou undish, "constricted. at the top, the appendages on the phyllaries spreading, yellowish or brownish, pinnately spinose, the spine of the — phyllaries 1-2 cm. long, the lateral spines short and stout, 2-4 on each side, the innermost phyllaries i a coriaceous acute tip; flowers yellow; pe sei compressed, 4-5 mm. long; pappus of strongly graduated pal and bristles, rose to purple and dark brown, 5-8 mm. long. Alon o ranietcies mayan Park, Los Pigs a Cali. fornia; perhaps not persisting. Native of the western. part of the. Iectitcrraness re May-Jun 145. CNICUS [Tourn.] L. Sp. Pl. 826. 1753. Annual herb with leafy stems and alternate, parted lobed or dentate, spinulose leaves. Heads medium-sized, sessile or terminating the branches, heterogamous with the outermost rf OQ. “=, Sj — et) 4 =) = 2. p > a ay o 3 o wn co - i) FA eat i) = o i ° fo or * nd 3 ° co = oO fn ‘2 ae | 7g Cc eeply 3-parted c ; hilum lateral, large, deeply excavated. Pappus pee in 2 unlike series of 10 bristles eac e outer alternate with the inner and with the teeth on the achene-rim, subrigidly setose, subterete ; the inner much shorter, vabewid and a little paleaceous-flattened. [Name derived trom) the Greek and applied to the safflower in Latin. native of the Mediterranean region and adjacent parts of Asia. 1. Cnicus benedictus L. Blessed Thistle. Fig. 5951. Cnicus benedictus L. Sp. Pl. 826. 1753. Plants with erect or spreading, leafy stems, sparsely arachnoid-tomentose and villous as well as punctate-glandular; primary stem sometimes lacking or very short with elongate sprawling 5949. Centaurea melitensis — 548 COMPOSITAE branches pala from the basal or lower cauline leaf-axils, or She primary stem erect and u branched for uch as 2 dm. and branching above, somew what ngled, not winged. sal are subrosulate, eae me the lower cauline leaves obla nr te or cblongish, pinnate ely lobed to pin- nately parted, narrowed to a longer or shorter petiole, the middle and u auline leaves sessile by a broad, decurrent base, oblong, up to 25 = lon ene 6 cm. wae ae pen much smaller, sharply spinulose-dentate to pinnately parted, bright eee and vi clap wlan pec les the upper- most ereres (or subbasal leaves if fear is s vee wie clustered ee the and exceeding it in = math eads narro aie to broadly ovate, 2-4 cm. long; phases SE AY Oe or ap- sed, i in several series, the outermost i ovate, abruptly tipped by an cae pig slender se the middle Hk Sig ovate phyllaries bearing a spreading, oes apinnes appendage to about 2 cm. long, the lateral spines in about 5-7 pairs, the appendage in the innermost elongate hile much reduced; flowers yellow, the pala na sterile ones inconspicuous; achenes oblong-obovate, about 8 mm. long, grayish brown, scarcely compressed, slightly coristricied at the truncate apex below the dentate rim; pappus br sank tinged, the outer bristles about 10 mm. long, de coca face a thes roughened, the inner bristles 2-4(5) mm. long, bearing a few ee disposed processe: alon ides ds, iecapeias and roadsides, flourishing in rich cultivated soils, widely al alae but not yore. common; arr! gp gu south to southern California; occasional eastward to the Atlan Introduced in many parts of the world from the Mediterranean region. April- Sept. Tribe 10. ARCTOTIDEAE* 146. GAZANIA Gaertn. Fruct. 2: 451. 1791. Nomen conservandum. lants herbaceous, mostly perennial, low and spreading, with subrosulate radical leaves av cent. Re cle Seo or convex, shallowly honeycombed. Flowers hots yellow or orange, the ra Saris inged or marked with blue, purple, or brown, the disk-flowers sometimes dark. Achene dhave turbinate, long- villous. Pappus-bristles in 2 series, narrowly acumi- nate, thin, s a inconspicuous and concealed by the long copious hairs of the achene. [Probably in honor of Theodore of Gaza who translated the botanical works of Aristotle r South African genus of about 24 species. Type species, Cinistle rigens L. 1. Gazania longiscapa DC. Treasure Flower. Fig. 5952. Gazania longiscapa DC. Prod. 6: 513. 1837. Scapose pe equaling or exceeding the leaves. Leaves entire or tii ately divided sey ee oblanceolate = ole 1- m. elliptic, acute, attenuate at base into a nabos petiole, the blade and peti ee ong, the up side smooth and glabrou s, the lower side (except the midrib) densely lanate-tomentose, the revo- lute margin scabrous Gliste ; involucre about eas long, the cup eas 36 phyllary-lobes attenuate-acute or spine escent, the outer np te bes scabrous-ciliate ; heads 3-6 cm. across, the flowers bright, cic in shadow and in late st Siicitoe pappus- -bristles 3-4 mm. long, hidden by the hairs of the achene which are shoe twice as long. Feb. “Mas g from cultivation in southern California but perhaps not persisting. Introduced from South Africa. e Arctodtis L. Sp. Pl. 922. 1753. Acaulescent or caulescent herbs or shrubs. Leaves alternate, unarmed; he ads radiate, the ray-flowers pistillate and fertile, the disk-flowers perfect; payers in several serie distinct; re- ceptacle deeply pitted, fimbrillate; achenes winged dorsally, the wings appearing to form 2 pits, the base of o chene forked with a long tuft of hairs; pappus in 2 series, the outer pong scarious, paleaceous, be gee e€ inner series someti = ate! all. * hadifolia var. grandis (Thunb. ) Less. Synop. Gen. Comp. 26. 1 (A. grandis Thunb. . Pl. Cap. 706. 1820.) Annual tomentose herb with erect stems and undulate, dentate 0 lo leaves; te ee very eonoak: the outer herbaceous, end elliptic, the middle and inner elongate- ‘oblong, scarious or pe out apt ed; achenes plumply obovate, 2 sea ‘age for the buff, incu Shae dentate, lateral margin f the d rsal pits, the basal hairs Tight 1 Br od co 4 sgn outer pappus-pa hyaline, 3 mm four, “tg inner aus consisting Of rudimentary bristles. Y Octasiount ‘escape from cultivation in southern California probably — persisting. Native of the Cape region, South Africa. Bérkheya Ehrh. Beitr. 3: 137. 1788. South African herbs o} ascneogel = more or less spiny, or spin “ai dentate leaves; heads discoid es. radiate; phyllaries in several series, Thai r less united by thei Snares thes ; a cup, the free ends spine-tipped; ; receptacle mingle,” B. heterop or von a pitted, ‘more oF om ‘ess a + Pe ae: ae 1 — (Stobaca cterophylia Thunb. . Pl. Cap. 622. ye Thistle-like he ~ wit th bine 2 divided aoe ZB heads discoid; 4 ries divaricate, Aho on margin and at apex; achenes enclosed within receptacle- pits; PRDhive — inka. -25—-0.3 mm. long, obtuse or subtruncate, denticulate, On ballast near Portland, Oregon. Nativ of Sout enidium Linnaea 6: 91. 1831. Acaulescent or rese eeabiiey the closely related Arctotis, i Sevine | in glabrous achenes that are Goony epappose, aot PE il ie ae gts t Prag oy (Jacq.) Stapf, ou Ma ey: pl. ai 1928. (Ar — Fryers F pen Hort. Schoenbr. 21, pl. 166. 1 . wyleyvt Harvey 1. Ca 463. ae aa Annual her ith thin cobwebb Seeker lower lea — more or less onibine ae S oeee i ne leaves idea tox with an auriculate base, all lanceolate ee lanceolate or oblongish and ete pinnately lobed; — sig dont at the ends of branches; phyllaries or 1c, the outer linear with a slender spreading herbaceous tip Rett ate-spatulate —e Z rounded scarious ap- pendage; rays orange with dark purplish me achenes s hae th led; pappus pres = a minute own or lneki ing. Occasionally escaping from cultivation in southern gay we dasg Nee of South Ate * Text of Arctotideae contributed by John Thomas Howell. SUNFLOWER FAMILY 549 Tribe 11. MUTISEAE sedans several- to many-flowered; leaves alike. Shr flowers yellow. 147. a ais. Her ete : flowers pink or white. 148. Involucres 1. flowered; stem-leaves linear-lanceolate, floral leaves ovate. 149. By ae ae 147. TRIXIS P. Br. Nat. Hist. Jamaica 312. 1756. (Hyponym) ; Crantz, Inst. Herb. 1: 329. 1766. Shrubs or rarely herbs with alternate ren as or dentate leaves. Inflorescence cymose or paniculate. Involucres 2-seriate, 9-12- red, the outer phyllaries few and usually shorter than the inner. Flowers perfect, mi oat "Cos olla bilabiate, the outer lip erect, 2-cleft, the inner spreading, 3-toothed or subentire. Anthers with tail-like appendages. tyle- e inne branches flattened and truncate at apex. Achenes subcyclindric, the scabrous pappus- -bristles copious, white to tawny, arising from the a bie disk-like summit of the achene. [From the Greek meaning threefold, king e 3-cleft corollas. ] A genus of about 35 species, natives of isan tad United States, Mexico, and Central and South America. Type species, Trixis suffrutescens P. Br. i 5950. Centaurea solstitialis 5952. Gazania longiscapa 5951. Cnicus benedictus 5953. Trixis californica 550 COMPOSITAE 1. Trixis californica Kell. California Trixis. Fig. 5953. Tents celiforasce Kell. ge Calif. Acad. 2: 182. sd 53, 1862. Bot. Calif. 2: 459. Trizis aapartifolia var. ee Gray, Syn. - N. Amer. 17: 410. 1884. uch bran sasag shrub about 0.5-1 m. high, leafy . to the heads, lower stems gray, upper aa chlets striate, tan, glandular- puberulent. Leaves alternate, sessile or subsessile, hen late to ovate- “te late, 6-15(20) mm. broad, 2-8 cm. felis sparingly appressed-puberulent on both surfaces, densely glandular especially beneath, midvein prominent below, the margins entire i on the older leaves; i i g yellow, about 1 ong; getienns dark, lon ps eyelivideic and narrowed above, densely widelay hesiaelous the pappus copious, straw w-color ie Rocky slopes and canyons wall Lower Sonoran Zone; ea n San Bernardino County and south through the Colorado Desert, California o Lo wer California sae ‘east ‘to gehen Texas and northern Mexico. Type locality: Cedros I Island, Lower Californie Feb.—June and Oct.— 148. PEREZIA Lag. Amen. Nat. 1:31. 1811. Perennial herbs or rarely sriseonti ‘aoe (in ours) erect and leafy, arising from a more or less woody caudex more or less covered with felty, rust-colored woo eaves alternate, mostly sessile. Inflorescence single and terminal or of few to many heads in panicles, or : lu s lavender, or white. Corollas bilabiate, the outer lip 3-dentate, the inner wit free or strongly recurved segments. Anthers with tail-like appendages. Style- Seguches flattened and truncate at apex. Achenes linear-cylindric or fusiform, the pappus copious, of scabrous, white to OS eeuny ° or brownish bristles. [Name in honor of Lorenzo Perez, a pharmacist of the sixteenth centu genus of wet 75 species, natives of southwestern United aratee, Mexico, and Central America and of western South America. Type species, Vocdivicns magellanicum L, 1. Perezia microcéphala (DC.) A. Gray. Sacapellote. Fig. 5954. Acourtia microcephala DC. es: 7: 66. 1838. Pavess microcephala A. Gra miths. Contr. 35: 127. 1852. Perezia sericophylla Millsp. & oe Field Mus. Bot. Ser. 5: 297. 1923. (Incidental mention) Stout erect perennials 6-14 dm. high, the striate stems densely leafy and glandular-scabrid. eaves thins ovate to elliptic-oblong, a m. long, sessile with a Ss d base, the larger cordate- claspi gin spinose-denticulate, chartaceo ous i a what ——— -ve , glandular-scabrid or rulent on both surfaces; infloresc rat d, m ined i ence r ose-pan aralate easy bre cted ; heads about 10-12 mm. high, 10-20-flowered; phyllaries oblong- pe ie nceolate, mucronulate, glandu lar-puberulent, loosely imbricated in about 3 series; corollas lavender- dele white; achenes glandular- Poberutent, the pappus white. Common on dry slopes, Upper Sonoran Zone; San s Obispo County and the Channel Islands south t San Diego County, California, and siiacoar pated, eattornia. Type locality: California. Collected by Donpiar June— ey aa A /, \ i, i) Vg i Wi Z Le Ih SS G “iy 5955 5955. Hecastocleis shockleyi SUNFLOWER FAMILY 551 149. HECASTOCLEIS A. Gray, Proc. Amer. Acad. 17: 220. 1882. Shrubs with cauline leaves wai appearing singly on younger shoots, later develop- ing axillary fascicles of smaller leaves in the axils. Involucres in terminal clusters, sub- tended by whorls o oval or Aare reticulate, stramineous, floral leaves, each involucre 1-flowered with narrowly lanceolate, spinose-tipped phyllaries. Corolla whitish, tubular, deeply divided into 5 linear aprendings ‘lobes firm in texture and widely spreading in anthesis. Stamens fete in texture with a glabrous appendage. Style purplish red, surpassing the anthers, stigma emarginate. Achene glabrous in age, the pappus s coroniform and laciniate- fare coriaceous. [Name Greek, referring to the separate enclosure of each flower in its involucre A monotypic genus, native in the desert regions of southwestern Nevada and the Death Valley region of adjacent California. 1. Hecastocleis shockleyi A. Gray. Prickle-leaf. Fig. 5955. Hecastocleis shockleyi A. Gray, Proc. Amer. Acad. 17: 221. 1882. out, divaricately branched shrub 5-7 dm. high, with rigid leafy oe bark of the older stems tate gray or blackish, somewhat shreddy, that of the younger stems stramineous, shining the ultimate ifasnhlete tomentose. Cauline leaves ahetuate sessile, fascicled and shorter on the older leafy stems, coriaceous, linear-lanceolate, strongly spine-tipped, the margins with few sle spi young es fi so mm, long, reticulate-veined, becoming stramineous and papery ; involucres 1-flow ered, narrowly cylindric, he rs et in 4-5 series, appressed, cuspidate-acuminate, loosely woolly on the margin ; achenes sparsely tomentose when young, glabrous in age, subcyclindric, somewhat narrower at bas e, the covering thinner in texture than the coroniform lecinicie’g us. Desert mountains, Upper Sonoran Zone; western Esmeralda catia ineral Counties to Clark Nevada, westward to the Death Valley region and the In la Mountains, cong" om Type ring Fat Gasdels “at og Tt eralda County, Nevada, altitude 6, ee shies Said by Shockley to have been collected at Silver Peak, Esmeralda County, instead of the locality given by Gray in the selina description. ton a uly. Tribe 12. CICHORIEAE. Achenes widvent paps fxith short, crown in variety of Eee’ aves basal, t r bract-like; eg pecies. Dese ee : flowers white or pinkish. 163. Atrichoseris. Derchatily of ot Sierra = fies ada; flowers Arig 154. penance Leaves happen eae and cauline; Siteteced species. 176. Lapsa Achenes with p Pappus ast or = completely eo or plumose s aceous, 0 aw above the calananioele base (capillary bristles in Nothocalais alpestris, Seer is eat A Phyllaries enclosing the outer achenes; pappus of the outer achenes aifcrian Bla of the inner. 'ypnois. Phyllaries not cae the outer achencs; peeess of _ — — outer ace ali - Pappus of 2 or 3 series of u 150. Cici : Pappus- cr ina ress series, surmounted by a smooth, scabrous or Ae awn (pappus ls fw ms of M onsets. Tig lee shades of yellow ith hentia ot erect; outer and inner phyllaries equal or nea rl —— Sy nearly so ae Annual or ah Lihat with heads nodding in bud; outer aay n rip so tg iortér than the in atiapegt bos -conspiewousl plumose and sometimes paleaceous-dilated at is extreme besie or with wien arte sa and slender or short and stout beaks (outer achenes truncate in Hypochoeris . ees we r papers paleaceous in Leontodon leysseri). ed into a short bea 169. Leontodon. Achenes rie a long slender beak. Phyllari 2 series, unlike, the outer broadly ovate, subcordate; pubescence coarse- tee i . Picris. Phyllaries in a series or loosely imbricated, alike; plants gisheous: (except Hypochoeris). Stems scapos 166. Hypochoeris. Stems leafy. Leaves grass-like; involucr lat ctlets. 164. Tragopogon L pinnatifid; icauers with calyculate bractlets. 158. nehiesguia, Achenes truncate. Involucre ly imbricated, the pt im ies rounded and broadly scar Anisocome. —_ not or but little suave subtended by calyculate ces the phyllaries nar- owly or not at all meets ined. us of stout awns g d the b. 155. Chaetadelpha. Pappus of slender p se bristles. Ligules yellow; bristles in more than 1 series, unequal. 165. Scorzonera. : gules pale or acts pink; bristles in 1 series, equal. 157. Stephanomeria. Li ris! Pappus-bristles capillary, either smooth or scabrous, ee barbellate. Achenes flattened (obscurely so in ‘Soiche 2. 552 COMPOSITAE erate not beaked. 170. Sonchus. Achenes beaked. 171. Lactuca. Achenes me flattened, either angulate or terete. Ligules pink or purplish. Heads nodding before and after anthesis; plants of coastal Oregon and b fee ngto: eads anthesis; a of the dese 156. Lygodesmia. Shewien shades of yellow, sometimes rae r cream-colo oe fick in Malacothrix blairii)., Lea ll basal; heads solitary on scapose peduncle ew truncate; Fens tees Eaictiens 151. Nothocalais ne 152. Microseris, Achenes beaked or mer ae tapering at the summit in some species of Agoseris; aap bristles not akipy Achenes -ri scien muricate abov 175. Taraxacum. Achenes 10-15-ri ibbed or -nerved, not inuiode: muricate above. 153. Agoseris. Stems ei ; heads not on scapose peduncles. Achenes minutely to strongly rugulose or tuberculate between the angles Dep ressed branching annuals with crustaceous-margined ee achenes abruptly beaked. 160. Glyptopleura. Erect, rena glandular annuals without crustaceous-margined leaves; achenes taper- Nis beak, 161. Caius. Achenes st the angles. baat early deciduous, 1—8 bristles remaining in a few speci 159: Mclecithets, cages: persistent (deciduous at maturity in some species of Crepis). Phyllaries not at all thickened; pappus sordid or Le gidrrens rarely white P Hicvactumt. Phyllaries somewhat thickened at base or on midrib; pappus sett; white. 174. Crepis. 150. CICHORIUM [Tourn.] L. Sp. Pl. 813. 1753. Herbs with erect branching stems, alternate and basal leaves, and large heads of usually ncle blue flowers, pedu or in sessile clusters along the br Involucral bracts in 2 series, herbaceous, the inner erect btending or partly enclosing the outer achenes Receptacle flat, naked or slightly fimbrillate. Ligules pes and 5-toothed at the apex Anthers sagittate at the base tyle-branches slender, o s 5-angled or 5 ribbed, truncate, beakless. Pappus of 2-3 series of short bEfat scales. [From the Arabic name. | An Old World genus of about 8 species. Type species, Cichorium intybus L. 1. Cichorium intybus L. Chicory. Fig. 5956. Cichorium intybus L. Sp. Pl. 813. 1753. Perennial from a long deep taproot, the stems sparsely hispid, stiff, branched, 3-10 dm e high. Basal leaves spreading on the ground, runcinate-pinnatifid, spatu late in outline, $20 e — t base; any, Tro on the nearly naked or —_ branches ; “Mowers br ight blue or rarely white; involucres 9-15 m high; achenes 2-3 mm. long, 5-angled, truncate above, the pappus reduced to a minute fringed n. Roadsides and waste places, i in all the Pacific States, and more or less generally throughout the United States and Canada. x substitute for coffee is sometimes made $e he the roots. March—July. Scélymus hispanicus L. Sp. Pl. 813. 1753. At e-like herb 3-5 dm. high, more or less arachnoid-pubescent or sometimes oe gees leaves alter ate, 4-8 cm. hog pmo it or pinnatifid, the lobes strongly spinescent, midrib broadened below the confluent ribs from the lobes, and decurrent on the stem; heads in the axils of the smaller upper leaves; ~inwolueral erssa _— nbpe “et flowers heir 4 chaffy bracts more or less enclosing the beakless achenes. — of the Med an r and sparingly adventive at Los Gatos, Santa Clara County, California. Golden 151 NOTHOCALAIS* _ a) Greene, Bull. Calif. Acad. 2: 54. 1886. spherical, the outer phyllaries only slightly shorter than the innermost, glabrous to ciliate e) e-villous along the midline. Receptacle naked. Corollas all ee, yellow, well exceeding the involucre. Achenes columnar-fusiform. th or brous above, not beaked, the upper portion often vacant but not said differentiated from the body of - Text contributed by Kenton Lee Chambers. SUNFLOWER FAMILY 553 the fruit. Pappus of 10 to many silvery bristles, some or all of which are gradually pressed and widened downward im a narrowly paleaceous basal portion (or the Sristies essentially capillary in N. Rds tris). [From the Greek word meaning spurious, plus Calais, a synonym of Microseris.] 2 Gas genus of 4 species native to ater and western North America. Type species, Microseris troximoides *“ Nothocalais appea o be most nnd allied to Microseris and to Agoseris, resembling each of these genera in one or another of its. fd morphological features. The distinctive combination of characteristics displayed by its species, however, marks it as a neta group separate from either related gen Pappus of 30-50 capillary bristles; leaves usually toothed or pinnatifid. N. alpestris. Pappus of 10-30 elongate narrow paleae tapering into short awns; leaves entire or undulate- margined . N. troximoides. 1. Nothocalais alpéstris (A. Gray) Chambers. Alpine Lake-agoseris. Fig. 5957. Troximon alpestre A. Gray, Proc. Amer. Acad. 19: 70. 1883. Trox imon barbell Gieeh e ex A. Gray, Syn. Fl. N. Amer. 12: 437. 1884. Agoseris alpestris Gre ere 177. 1891 Agoseris edntinicia Green, lo oc. Microseris alpestris Q. Jone myst onquist, Vasc. Pl. Pacif. apa 5:-267.1955, Nothocalais alpestris Cheohee Contr. Dudley Herb. 5: 66. Glabrous or puberulent, perennial herb with an stniae: blackish, simple or multicipital, underground caudex and deep taproot. Leaves all basal, 3-20 cm. long, linear to oblanceolate or 8 S ay entire or more often remotel thed or pinnatifid with linear to broadly deltoid lobes s 1 to several ee the basal age erect, simple, 3-30 cm. t rous above, sometimes bracteate near the base; heads erect, 10- to many-flowered ; cna a res 10-20 mm. high, broadly narrowly c sshiarenlats, strictly hieuve the outer pyila gs ee te or ovate-lanceolate, attenuate, lightly shorter than the narrower inner ones, fin vn a ted with ink ate dorsally ; achenes br rown, about 10-ribbed, 5-10 mm. long, raicale, or the -3 m what nar- rower ge lightly scabrous ; pappus 6-10 mm. long, the 30-50 se recites Teristics r tarbelisiate or rarely subplumose. Open meadows and g lly slopes, Boreal Zones; Cascade Mountains from Mount Rainier, W. shington, io Gitkiyou remntchy California, and in the Sierra Nevada near Lake Tahoe. Type locality: Mount Adams, ash- ington. June—Aug. 2. Nothocalais oxime (A. Gray) Greene. False-agoseris. Fig. 5958. Microseris siaspornawee Gray, Proce. r. Acad. 9: 211. 1874. Nothocalais troxim s Gr reine; re bi Acad. 2:55. 1886. Nothocalais re reene, op. cit. 54. Scorzonella troximoides Jepson, Gas. FI. Pl. Calif. 994. 1925. Glabrous or oe villous, a al herb with a stout, blackish, underground caudex and long, often rope-like taproot. Leaves in a bas al cluster, ‘rather sti tiff and apn ar linear to linear- te) i ire 1 s nutely m n un 1 to several, Lge oe cm. tall, simple or only bracteate below, glabrous or + whine villous toward the e apex ; heads erect, 10- to mana flowered; involucres 14-25(30) mm. high, broadly or narrowly mpan nulate, the ehribasies narrowly or broadly lanceolate, attenuate, glabrous or ciliate and iia: villous along the midline, misien dotted or lined with purple on the midline, the outer 5957 5956. Cichorium kane 5957. Nothocalais 554 COMPOSITAE a more than three-fourths as long as the inner ones; ~~ straw- colored or tan, 8-1 long, about 10- aig slender, columnar-fu yee rm, scabrous towar e apex : POERUS-Pante 10-30, silvery 1 m. long, rather unequal, each ea in: pete Romans d from a short nal awn into a si narrowly palaceous portion about 0. on wide near the tas a=339 erie slopes po flats in hl often rocky soil, Arid Transition Zon s and canis east of th Cascade Range from British Columbia to Sierra County, California: east through ere give g? jd ie’ Snake River Plain to southwestern ne eo northern Utah. Type locality: “hills on the Clea River, Oregon (now Idaho Tee. ).” April-June. ast of the Pacific States, evpscisily 3 in Idaho, N. troxrimoides tends to vary in the direction of N. nigrescens (Henders.) Heller, often acquiring the broader, plane, — leaves and glabrous, diffusely speckled outer phyl- laries of that species. Nothocalais sine nia itself Bi n wet meadows in the Rocky nen of PB oi Montana, and Wyoming; it may oc our the high mountains of northeastern Orego 152. MICROSERIS* D. Don, Phil. Mag. 11: 388. 1832. Caulescent or acaulescent herbs with milky juice, the plants perennial, with 1 or mo eshy taproots, or annual. Leaves entire to variously toothed or pinnatifid, i eat or iat apace de (villous in section Calocalais). Heads many-flowered, erect or noddi in bud, bo singly on naked or remotely bracteate, glabrous or scurfy, scape-like pe- e Corollas al sie tieh elongate and showy or short and Ww or white. Achenes columnar to truncate- Hansa not beaked (or short-beaked in section Calocalais), about 10-ribbed, smooth o scabrous, the outermost often densely white- 5-6 ; : 5 a ° =) mn 2 a : 4 Le fe oy : ° 5 ry i=} or a8 MS 2 So ' ceenicuinte to plumose. [Name Greek, meaning small c i of 16 species, occurring principally in western North America but with a bs sg a in Scie and on ey ete id eal y New Zealand. Type species, es nto pygmaea D. Don. Chrom numbers, deter- pe A or verified by the contributor, are given where kno Plants perennial; fl 1 exceeding the involucre; pappus-part Pappu ‘ion ee Srownick — bristles; plants acaulescent, the scapes naked ah rag 08 paconches: (subgenus par m) ee bert 6-30 ee or tawny, awn-tipped bra plants caulescent, s Ast wey aa leafy above ple and only bracteate. (subgenus ae lla ss Paces pasts 15-30, bright white, the awns soft, ros ans rie tet Btn the awns rather stiff, karen par sirate to barbéliuiate or “ sebclentose then the pap Pappurpaleze ‘ 10 mm, long, fimbriate at the apex about the base of the awn; awns subplumose, tawny. 6. M. sylvati ca. Seasgpen ee usually less than 5 mm. long, tapering into the awn or emarginate; awns minutely culate to barbellulate, wire tawny Pe phyllaries ovate- ‘peg to Laondly ovate or circular, acute or cuspidate, usually gla- iata brous dorsally, all over 2.5 mm. broad. 3. M. laciniata lacin Outer age s linear to si eedaie acute to attenuate, often scurfy- Hicher lent dor- dag *'e ldetlent on each head not over 2.5 mm. broad. Involuge cy ee or narrowly campanulate, 15—25- aia g 5 ag hon —_ high; Siskiyou owellii anviners. narrowly one Rapedx campanulate, 25-—75- Moa 10-25 mm. high; Coastal re d Cali Pergee ruler - 5-1. re mm. long, deltoid or ovate; sg achenes brown; awns white . M. laciniata leptosepala. Peitie ‘Palene ee — long, lanceolate; inner ee @ pale straw-colored or dul white; awns taw 5; . paludosa Plants Pormenvedh florets inconspicuous, sanatine or penis exceeding the involucre; pappus-parts 2. 5. (subgenus is) Heads erect before anthesis; achenes attenuate into a short slender beak; pappus silvery, deciduous. (section Calocalats) 7. M. linearifolia. Heads inclined or ve pee anthesis; achenes truncate or slightly narrower toward the apex; pappus silvery or sordid, p Pappus-paleae fiber adie tee: bifid or minutely lacerate at the apex; plants frequently caulescent. (section Brachycarpa) Achenes columnar, + the apex, gray or straw-colored to brown or bluish may or if blackish then equaling a aig in length. 8. M. heterocarpa. Achenes Bina k= Fa ag not flared at the apex, blackish, about ty ae ~ Tength of the paleae. Pappus-paleae various but if linear- owe sed then entire at the apex, Heiaaty pice into the awn; plants acau bee ent. (section Microseris) 3.0 mm. long or less anes Kg argu to chenaieal; the flared apex as broad as or broader than the body of I , th t 1 ed. ei leae over 2.0 mm, long, the margins strongly incurv: a. i lotphied puasearees Paleae 2.0 mm. long or less, flat or the margins lightly per . M. elegans. hex ributed by by Kenton Lee Chambers. (For complete synonymy of the annual species, see Contr. Dudley Herb. 4 a: 283-306. 1955. ) SUNFLOWER FAMILY 555 Achenes truncate-fusiform, the pig narrower Sie the wae of the fruit. Paleae conspicuous, at least 1.0 m . M. bigelo Pabiex Leeititicite seca: tecliekly. 0. Pie mm, long. ” M. houatiadt tenella, Achenes over 3.0 mm. long. Paleae averaging 1.0 mm. long or Palea Shaan pe eae the midrib ay stout, tapering, joirwions ee to one- * ift h of the maximum palea w 11. M. acum Paleae circular to 3a oe. the stout or slender midrib techie less than or la clad one-fifth of the maximum palea Paleae arcuate only at Nee agen, aia and flat rey the midrib linear, abruptly thickened at the . M. bigelovii. Paleae — lonicue pe margins ica < convolute, the midrib linear tapering He the bas Palens Bac, sper y 5, minutely scabrous, am, ae lightly smoky, the margins sapere the ved. . M, campestris. Paleae 23 or fewer, villous to rine ig ‘lee, clear or chalky to tawny or blackish, the margins strong Paleae eg iy srg tool to rea a ge" age of the. ‘promot achenes a mm, shorter has the rs Pe or of equaling or exceeding it then the latter more than 4.5 mm, long. 10. M. douglasti douglasit 10. M. douglasii tenella. Paleae 1 Paleae mostly less than 1.0 mm. long. = 6 ontiittl tery, + \\\ ony Nothocalais troximoides 959. Micros eris borealis cna: 556 COMPOSITAE 1. Microseris borealis (Bong.) Sch.-Bip. Apargidium. Fig. 5959. Apargia borealis Bong. Mém. Acad. * Pétersb. VI. Math. Phys. Nat. 2: 146. 1832. Leontodon boreale DC. Prod. 7: gon ares ium & boreale Torr. & Gra 3 Ais er. 2: us 1843, Sch. -Bip. athe — 310 866, Scorzonella borealis Greene, Pittonia 2: Glabrous, acaulescent, perennial herb with numerous fleshy roots from a short rootstock. Leaves up to 30 cm. long, linear- vchlaneanlane acute to acuminate, ent or remotely denticulate ; scapes slender, naked and unbranched, erect or curved at t se, 1S m. tall, sometimes lig = urfy above; heads - ed, nodd before anthesis; involucr 8 m hig panulate, the inner phyllaries Se the outer phyllaries shorter, 1 ane imbricate, linear or lan ne serrate, ove ro c ee le black-tomentose; florets yellow achenes bro ong, nar, atten the : a panels s 5-10 mm. he a ete Browitieh harbelialate awns GHEY slighty thickened 2 the bas rshy meadows and sphagnum bogs, Hu Transition and_Boreal soni along the — Seer rion PERS wag te British Columbia and south at pen or iri in the Cascade Mountains of Wa shing and Ore to Humboldt County, California. Type locality: Sitka, Alaska. Collected by Mertens. June— yf tre 2. Microseris nutans ( Hook.) Sch.-Bip. Nodding Scorzonella. Fig. 5960. Sg }7. t Hook. Lond. Journ. Bot. ah 1847. Crepis nutans Geyer ex Hook. loc. cit., as as Ptilophora nutans A. Gray ——— Amer. Ac af "IL on “143. 1849. A. 0. Ptilophora major B rape A. Grad , loc Calais nutans A. pee in ta Pacif, R. ae #2113. -1857< 1 Calais nutans var. latifolia D. C. Eaton, Bot. Key ear 197. $1871, Stephanomeria (?) waite ae Proc. Calif. Acad. 5: 39. 1873. Calais gracitiloba K llogg, o Ptilocalais nutans Green ne, Bull, “ali Acad. 2: 54. 1886. Pesecaiote major Greene, loc Leg gee ae | ureen Ptilocalais ce Oster Mahiskces te 4% ge cros ged r Nels. & Macbr. Bot. Gaz 1916 var. ase Tee Man. FI. Pl. tobe en 1925, Glabrous 1 few, erect or curved at t A base, sle nder, 10-70 cm. or usually branched and leafy abo ve. flaves it spat ; r t anulat i outer phyllaries unequal, shorter and nearly area st eas core r often scurfy-puberu- lent, aged = —. Bdegrs the smallest ones on each head m. br a G r less; florets yellow; achenes g —8.0 mm. long, columnar or eetainea fusiforia: pappus 5-10 m fa bright whe ‘the i. 15-30, oblong or nk 1-3 mm. long, obtuse or somewhat fim- riate — ses slender distinctly plum Range a and conte in the Sier > Nevada and North Coast Range of California east to Wyoming and Colorado. Typ € loc: “Dry sunny loamy ge Porte ok of Spokan and Coeur d’Aleine cncusthita Kicil —Aug 3. Microseris laciniata (Hook.) Sch.-Bip. Cut-leaved Scorzonella. Fig. 5961. —— mardgnets Hook. Ae a Amer. 1: 300. 1833. beg onema ? laciniatum Hoo! 01. Scscaecelh pe as. Trans sone Phil. Soc. II. 7: 426. 1841. a Nutt. loc s Gar in ice Pacif. R. Rep. 4: 113. 1857. Microseris laciniata Sch.-Bip. Pollichia 22-24: 309. 1866 Calais glauca A. Gray, Proc. Amer. Acad. 7: 364. 1868. 7 procera A ci Pe SF 8 r ~~ — fe procera A. ae op. cit. 9: 209. 1874. ris proc ray, op. cit. 19: 64. 1883. Scorsnalla shoot ye Greene, a Calif. Acad. 2: 50. 1886. Scorzonella procera Greene, ] aac ella pratense Greene ts. oe Sl. Drew, Bull. Torrey Club 16: 152. 1889, a maxima Bioletti, Erythea 1: 69. 18! Scorzonella procera var. pratensis Jepson, Man. Fl. Pl. Calif. 994, 1925, labr taproot, the stem goed : rect, aie nder to very stout, 15-1 120 cm. tall, rather agecione'd leafy or bracteate “then Rs: above o Lp Sanat hs only near the “sg or uw c all indi- viduals. Leaves chiefly basal, up . 50 cm. long, a: to broadly oblanceolate, attenuate to SUNFLOWER FAMILY 557 obtuse, entire to few-toothed bis laciniate-pinnatifid ; heads 30-250-flowered, meni nodding befo . high i one-third as long as the innermost phyllaries; florets yellow achenes bro columnar, attenuate toward the base; pappus 7-15 mm. long, the — ae 6-10, “Gel toid to lanceolate 0.5-4.0 — a mm. long, each tapering itis a white tawny, minutely spiculate to ines een aw eadows and ad cits ssy hillsid es, Upper sam and Transition Zones; Pacific Slope mostly west of the Cas ae ies tain 5 tine the vicinity of uget Sound, "Wa shin renee outh to Sonoma County, California, and Mg teouse the Klamath region to Lake Cane pet 4 and porta Nac teens County, California; dager not on the ee coa: Type locality: ‘“‘Dry plains of the Columbia, from the Rocky Mountains to the an.’ Collecte ee by Douglas. a il-July. Toward the ~ enateyer amit t of the range, individuals of this subspecies may possess smaller heads and in- volucral phyllaries superficially resembling those found in the generally coastal subspecies /eptosepala, below. icroseris laciniata subsp. leptosépala (Nutt.) K. Chambers, Contr. Dudley Herb. 5: 61. paced (Scor- zonella yeti Nutt. Trans. Amer. Phil. Soc. IT. 7: 426. 1841; Calais are es A. Gray, Proc. Amer. Acad. a: 8; Microseris leptosepala A. Gray, op. cit. 9: 209. 1874; M. bolanderi A. Gray, op. cit. 19: 64. 1883; Scorz ue bolanderi bp pe Bull, Gt, Acad. 2: aa 1886; S. laciniata con bolanderi Jepson, Man. FI. Pl. Calif. 994, 5; S. leachiana M. E. Peck, Proc. Biol. Soc. Wash. 47: 188. 1934.) Involucre narro t ererar areas, Humid Transition Zone; plains of the lower i ette Seats Sryeen, and south “a the coa a o Mendocino County, California, Stender inland on the lower Klam and Trinity Rivers and probably else- wince on the western. drainage of Bsa Coast Ranges. Type coshiee: poo the pute of the Wahlamet, ” that ig, the mouth of the Willamette Rive Phi Microseris howéllii A. Gray. Howell’s Scorzonella. Fig. 5962, Mi h ht aie Gray, Proc. Amer. Acad. 20: 300. 1885, emend. Syn. Fl. N. Amer. 12; suppl. 454. 1886. Scorzonella h LI: , Bull. Calif. Acad. 2: 52. 1886. Glabrous or Puenly se urfy-puberulent nial herb with a fleshy fusiform taproot, the stem slender, erect, usually aie 15-50 cm. tall "sparingly pete koh pm below the middle or some- times unbranched and only bracteate below. Lea iefly basal, up to cm. long, linear to narrowly lanceolate, attenuate, entire or Taciniate? perne F with slender lobes; heads 15—25- flowered, nodding before anthesis ; involucres 8-18 mm. high, cylindrical to narrowly campanulate, i er, un , ne yceu- es on each head only 0.5-1.5 mm. broad as less than one-third as long as the innermost phyl- i n u r i oe bro oy m. base ; pappus 6-12 mm. long, the paleae 6-10, lanc colate, Slates. mm. long (or infrequently only 1 mm. long), — tapers into a white, minutely spiculate Moi ocky fl frequently in serpentine te aig Zones; Siskiyou Mountains of southern Jose- phine ej echeacs C untiek. Oneabks and adjacent California. Type locality: Waldo, Josephine County, Oregon. April-July. ig axon is ne ee rather similar to M. laciniata subsp. leptosepala, but its habitat and typically much gear pappus-paleae ap o be distinctive. otf , cet 10 5961 of 5961. Microseris laciniata 5962. Microseris howellii 558 COMPOSITAE 5. Microseris paludésa (Greene) J. T. Howell. Marsh Scorzonella. Fig. 5963. Microserts sylvatica var. stillm ages NAD y, Proc. Amer. Acad. 9: 208. 1874. Scorsonella paludosa Greene aL aad. Bone. Scorzonella sittin var. wtillmands Jenson, Man. a PE, Calif. 994, 1925, ntegrifolia Jepson, loc ee plows f T Ho well, Leaflets West Bor. 5: 108. 1948. or seaciget: Reagan sks: a nd pi Seachente | ralaaieds lepidota Heller, Muhlenbergia 2: 14 Microserts montana H. M. Hall, Univ. ‘Calif. are Hs 3: 252. 1907. Glabrous or seury. puberulent, perennial herb with a slender elongate taproot, the stems few, erect or curved at the base, slender or stout, 15-75 cm. tall, leafy and branched gp the base or : i 3 q om es y above. Leaves iefly te) cm. long, na rowly elliptic to broadly oblanceolate, attenuate above, cay ar; Soaks mostly 6-8 nim. long; pappus mostly iGge mm. long; com- mon » Widesprea . A, aurantiaca. Some trichomes onetigd head gland-tipped; beaks commonly 8-10 mm. long; pappus a — hing this nitanteema i in eae age Sierra Nevada. chee A. Beak of mature vewdenns more than twice as long as the body, or sometimes shorter in ma oe pe Heads pa od ely lar fe, rarely ity than a a mm. high in fruit; — 11-25 Sti at mm. long; s 8-18 (mostly 10-17) mm. oe coastal hills and inland Achenes s abruptly beaked from a truncate apex; leaves initorady retrorsely lobed; pp lh agg mm. ; pappus 14-18 mm. lon Ar Achenes Aba ng gradually into the beak; leaves irregularly lobed; ligules 3- 6 mm. «Sty (r arely longer) ; pappus 8-12 (rarely ae mm. long. . A. grandiflora. Heads eens small, less than 25 mm. high in fruit; beaks 4-9 (rarely —12) mm. ge pappus 5-9 mm. lon, ong; plants of mar itime and coastal hills. 6. A. apargiotdes. Plants annual; purr fF involucre. 7. A. heterophylla. 1. Agoseris glaiica (Pursh) Raf. Short-beaked Agoseris. Fig. 5973. Troximon glaucum Pursh, Fl. Amer. Sept. 2: 505. 1814 Agoseris glauca Raf. Herb. Raf. 39, 1833. Macrorhynchus glaucus D. C. Eaton, Bot. King Expl. 204. 1871. * Text contributed by Quentin Jones. SUNFLOWER FAMILY 563 A goserts | issima Greene, Leaflets Bot. Obs. 2: 122. 1911. Apos seris isomeris Greene, op. cit. 123. Agoseris potent Greene, op. cit. 125. Slender perennials, glabrate, mostly 20-45 cm. tall, from a long heavy taproot. Leaves numer ous, hess: Slane eolate to narrowly oblanc a pa sme nate, entire or rarely somewhat denticulate, glaucous and pe brous or sparsely te n the mar gins and midrib below, ascending an falcate to ext begar uch s sect than t : ze several scapes ; scapes slender, glabrous ; heads usually narrowly campan HAddec florets yellow, in fruit 21-22 mm, high, 15-20(-25) mm. wide; mi pe aceite tapering gradually from the base, frequently purple-spotted, glabrous, the — series Sipe ore shorter ; ligules mostly 12-15 mm. long; anther-tubes 3.5-5.2 (m ostly 4.0-4.8 mm: ong; achenes mostly 6-7 mm. long, the beak 1.5-6.0 mm. long; pappus commonly 10-12 mm. ae wt sesh mountain meadows, Arid Transition Zone; east of the Cascade Mountains in Washington and regon, in northeastern California, ana eastward beyond our range. Type locality: ‘‘On the banks of the Missouri.” mp oer Agos glauca var. dasycéphala (Torr. & Gray) Ae ge Man. Fl. Pl. Calif. 1005. 1925. (Ammogeton scorgoneracfolis Schrad. Ind. Sem. Hort. Gotting. 1 Troximon ae B dasyce ported Torr. & Gray, Fl. N. Amer. 843; Macrorhynchus Fag var. dasycephalns a: Coat Bot. ng Expl. 205. 1871; Agoseris scorz rk ah ahs nein mostly 6-9 mm. long ; cies: tubes 1. a cipiens ong ; achenes mostly lon monly hispid dictate. tapering Bradvally. into wie beak 0 umid on one side distally and abruptly tapering; beak mostly 6-8(3-11) mm. long; ake. Gee or sordid, 9-14 mm. lon cabatenis to “age elevations in cig mountains ge ae North eae tore mostly Boreal gee ee Peat iny south to Tulare + Seri Bact nia, and southern Colorado, with disjuncts in Quebec. Type focal iy! “Alpine Prairies of the ky Mountains.”’ Co Nec ted by Drummond, Jun = he Only the typical variety, described above, occurs in oe range 4. Agoseris retrérsa (Benth.) hg Spear-leaved Agoseris. Fig. 5976. Macrerhanchss ores Benth. ee Hartw. 320. Kell. Proc. Calif. segs Y 47. 1873. Troximon retrorsum A, "Ope sen Amer. Acad. 9: 216. 1874. Agoseris retrorsa Greene, Phionis 2: 278: Stout perennials, mostly 2.5-4.5 dm. tall, from a well-developed taproot. Leaves numerou 1.2-3.5 dm ee pom at uniformly laciniate except for the long-acu igen entire aid the lobes more or les rorse, all callous-tipped, usually canescent toward the base; 1-2 n 1S stout scapes, g¢ r wi 3 ccose-pu , a t at eae : heads uniformly campanulate, 3-6 cm. high in fruit; phyllaries bout 3 series, the outer strik- ingly shorter in fruit, the inner about equaling the # floret w, becomi ots ci anthesis, Sis partic Ts Li ia ri aie usually 6-10 mm. long; anther-tubes 2.5-4 dark yellow ; achenes 5-7 mm. lor r less truncate af the apex ; beak smooth, igo pen 18-21) mm. ican eats 14 i8 ‘ei red A cate Radecki upper Sonoran Zone; the Cascade and Blue Mountains of Oregon and Washington, south to the Coast Ranges and the Sier rra Nevada, California, to the Angeles Ranges of southern California. Type locality : “Tn montibus Sacramento,”’ California. Collect ted by Hartweg. April—July. 5. Agoseris Brangeine (Nutt.) Greene. Large-flowered Agoseris. Fig. 5977. Stylopappus grandifiorus Nutt. Trans. Amer. Phil. Soc, IT. 7: 432. 1841. Macrorhynchus aan Torr. & Gray, Fl. N. Amer. 2: 492. 1843. Troximon grenciperans A. Heo Proc. oa Acad. 9: 216. 1874. Troximon Green ittonia 1: 174. 1888 Troximon plebeit i Pesta Ses cit. 2: 79. 1890. prnites arandifore Greene, op. cit. 178. 1891, e, loc, cit. pe essiaa eer = Grower: Eryth 1893. Tro } obt ibe sie ce Deutsch. Bot. Monatss. 18: 98. 1900. dif be ake dia Jepson, Fl. W. Mid. Calif. 500. 1901. goseris cinerea Grace ne, beaks s Bot. Obs. 2: cite eee Agate sek be = db. Bull. Torrey Club 38: r. plebeia Wittr. Publ. ica riae Biol. Sta. 6: 253. 1928. Pe Penny hk Bie 2.5-6.0 04 tall, from a multicipital caudex. Leaves polymorphic, ranging from linear an panenare to broadly spatulate and laciniate, the lobes irregularly disposed, gla- brous to canescent, usually at least pokecoee on the midribs ; scapes coarse, usually paaaincuiasty 5975. Agoseris aurantiaca 566 COMPOSITAE * ribbed distally, tomentose below, glabrous or scattered- ont above, —— canescent apices —4.0c¢ gh, 2:3 the head; heads lar. arge, broadly campanulate to hemispheric, 2.2 ohi 2.3-6.0 cm. wide in fruit; phyllarie s in 4— < series, the outer eeapleumgsty cilit e with white €-opaque, never sindidas or aL oage t trichem : florets yellow, drying purplish, a than the involucre; ligules 3-6 mm. lon nther cabes €) .0-2.5 mm. long — spindle-shaped, tapering gradually into the long filiform beak: pappus 12 fedid aay te . long Gra ba and bon val Wi ad in the lowlands a aig aaa in the mountains, Upper Sonoran and Transition Zone anc sland, British Columbia to southern California. Type locality : “High plains of e Wahlamet. m ‘Collected t ‘ay Nutt May-Jul 6. Agoseris apargioides (Less.) Greene. Seaside Agoseris. Fig. 5978. Troximon apargioides Less. Linnaea 6: 501. 1831. ot pee resis Hook. Fl. Bor. Amer. 1: 296. 1833. haus it Hook. & Arn. Bot. Beechey 145. 1833. Ma salen nchus Snsetearss Hook. & Arn. op. cit. 361. 1840. Taraxacum hirsutum Torr. & Gray, Fl, N. Amer. 2: 494. 1843. 849. rorhynchus h Macrorhynchus harfordti Kell. Proc. Calif. Acad. S247; 1873; pe elses humile A. Gra y, Proc. Amer. Acad. 1 2c YS883. Agoseris apargioides Crecne: Pittonia e277) ‘Aguastti hirsuta Greene, loc. cit. Agoseris humilis ants, Rev. Gen. Pl. 1: 304. 1891. Perennials, mostly 1.2-3.5 dm. tall, with a long cag sa taproot. Leaves mostly more than 9 cm. long, ascending, apni hl gues linear, eer ate or narrowly shisncoote: usually arise the lobes Sodlccts rarely entire o rely dentate, glabrous or sparsely pube rulent to tomentos low and on the 2 ied petioles e. pieicd usually several, ey exceeding the leaves, glabrous or scattered-pubescent, usually villous bene 2 the hea ; hea s broadly campanulate to hemispheric, in fruit 1.3-2.3 cm. high, 1.1-2.0 cm. bro phyllaries in 3 or 4 series, those of the outer 2 or 3 n series usually ecthaety commonly ciliate and Prather Soccly villous on the outer surface, trichomes in part glandular and transluc ent or frequently with scattered, glandular or white-opaque trichomes ; —6.0 lon, florets alice. ete hea ene sian: ligules 3.5 mm. long; anther-tubes 1. mm long ; achenes 5— . long ; beak fine, glabrous or hispidulous, mostly 4-9 mm. long; pappus niiialy ‘basbefinlats. 59 mm. lo g. Grassy hillsides, Henid Trans i tae near the coast from ewe mba! County south to Santa Barbara County, California. Type locality: Cattacse. Collected by Chamisso. May—Jun Agoseris apargioides subsp. maritima (Shelton) Q. Jones ex pail sage Maa. Pl. Pacif. Northw. 5: 1955, cer oseris maritima Sheldon, Bull. Tor rrey Nelo a0: 310. 1903.) Leaves mostly 4-10 ecm. long, “usualy reclined, tosis oblanceolate or spatulate, the tips obtuse, irregular i and remotely lobed, or more co peed rly merely dentate en trichomes of head eglandular and white- -opaque; anther-tubes 1.5-2.5 mm. long; wcheae mostly over 4.5 mm. long. Coastal sand dunes, beaches, and maritime a his ecesonnyy County, cs AP inane south 4 Monterey. County, California. Type locality: Clatsop Beach, Cla County, Oregon. Collected by ay— Aaosetis apargioides var. eastwoddiae (Fedde) Munz, Aliso 4: 100. 19 fe (Agoseris hiking sind Legs pa Fi foa? Segal 311; 808. 1904.) A related form which differs dic the preceding in having translucent trichomes on > heads; outer lig a8 8-16 mm. long; anther-tubes 3.5—4.5 mm. long; cchents mostly less that 4.5 mm. long. y from Humboldt County, California, to erttidton ee County. 7. Agoseris heterophylla (Nutt.) Greene. Annual Agoseris. Fig. 5979. Mecerh inch us: apie! ‘ares Trans. gees Phil. Sec. IT. 7: 430:° 1841. er K ymapl. cit., in Ciypiopiéues californ od ng op. ¢ le 431. Macrorhynchus Lettoteines Mtg ones be N. Amer. 2: 493. 1843. Tikitaes hetevephs as im Greene, Bull. T y Club 10: = 1883. Troximon phy var. aioe Gotne loc pee n heterophyll . kymapleura Gre reene, Joc. — hetero, oph hylla mnie Pittonia 2: 178. 18 pres heterophylla var. cryptopleura — “ae ct 73; ymapleura Greene, loc Agoserts major “Japenes ay Eat loc. cit. Troximon heterophyllum var. Se sient emecee ‘eens? Bot. Monatss. 18: 98. 1900. uksd. loc. on heteroph Agoseris heterophylla var. glabra atatcet 9 N. Ww. oa » 2: 402. °1901, A ts het. Pie sy subsp. normalis Piper, oo U. S._ Nat. Herb, 11: 544, 1906, Agoseris heterophylla californica Pi ae Agoseris Ratevbehvile en eda a Piper, loc. Troximon heterophyllum £, kymapleurum 3 M. pon Univ. Calif. Pub. Bot. 3: 278. 1907. Troximon ste 4 ae te H. M. 5a 1 hag: cit. hyllum var. californicum f. crenulatum H. M. oy Agee nee 279. Troximon erties var. californicum f. ae H. M. Hall, Agoseris heterophylla var. saliforn nica Jepson, Man. FI. Pl. Calif. oe toe, Ag heterophylla var. rot tags Jepson, loc. cit. loc, cit. prevnties heterophylia var. turgida Jepson, Slender erect annuals, 3-45 (mostly 6-20) cm. tall, sometimes weakly caulescent. sit s ascending, y Pbidaeeciaie. Senticy late, occasionally ne or lanceolate, entire or laciniate ; scapes frequently several, glabrou with a sparse pubescence, becoming glandu iar pubescent beneath the head; heads na Saree baateoe acs to ieciteoher: 3 in fruit eel 1.4-1.7 cm. high, SUNFLOWER FAMILY 567 1.7-2.8 cm. in diameter ; phyllaries commonly in 2 length-classes, more or less villous with septate A eam some = digo bra hae h and gland-tipped; florets yellow, frequently pinkish on drying ; ——_ 2.5-6.0 n or Fe lants from central California) 6-15 mm. long; aber ties be: 0 ke? or (plants Meat central California) 2.0-3.9 mm. long; outer achenes polymorphic, usually spindle shaped re ly 10-ribbed, tawny, sometimes the ribs abnormally developed, straight . sinuate, glabrous or h nat whitish es purplish, or the pericarp inflated to the obliteration of oeiieg ert capillary, anally 6-10 mm. long long. Dry requently weedy situations, ma er cilia Zone; mostly at lower elevations from British Co- lumbia on el! Lower California, be fg locality: “‘The plaing me Oregon.” Collected by Nuttall. April-— July. Only the typical subspecies, described above, occurs in our 154. PHALACROSERIS A. Gray, Proc. Amer. Acad. 7: 364. 1868. Glabrous perennial ay with a tuft of basal leaves and 1 to several 1-headed scapes. Involucre campanulate, 12-16 equal, narrowly lanceolate phyllaries. Flowers white. Achenes short-o ong, sles urely 4- peed slightly incurved. Pappus none. [Name from the Greek word meaning bald. headed, and seris, the Greek name for chicory. ] A monotypic genu 5979. Agoseris heterophylla 5977. Agoseris grandiflora 5980. Phalacroseris bolanderi 5978. Agoseris apargioides 568 COMPOSITAE He Sa ewegele bolanderi A. Gray. Bolander’s Dandelion. Fig. 5980. PI | A. Gray, Proc. Amer. Acad. 7: 364. 1868. Acaulescent: with a stout perennial root, glabrous throughout. Leaves narrowly ——- linear-lanceolate, or some linear-oblanceolate, “iain e on the simple or branched, woody root-cro slightly aecaicer 8-20 ¢ ng; scape 10-25 cm. 1 2 owered; involucre a m. lo out 1 cm. ar diameter ; phyllaries linear-lanceolate, 6-7 mm. long; ligules yellow, well peep achenes ‘somewhat 4-angled, truncate at both ends, about 3 mm. lon Wet meadows, Arid Transition and Canadian rene — ont of the Sieers Nevada in Mariposa and Madera Counties. Type locality : ““Westfall’s Meadows, above Yosemite Valley, alt. 8,000 feet.” June—Aug. Phalacr — boland ar. coronata H. M. H all, "Bot. nth ng 393. suit. " Rhanels resembling the typical ecies in kg habit bu “3 ivheues with a short crown. Western slopes of the gly Bold lair from a —— amas to Fresno County, California. Type locality: Pine Ridge, Sierra Nevada, alt. 5,4 eet, Fresno County. 155. CHAETADELPHA A. Gray, Proc. Amer. Acad. 9: 218. 1874. Perennial herbs with a deep-seated rootstock, many-branched from the base and dicho- tomously branching above. Leaves alternate, linear or linear- lanceolate. Heads 5-flowered, solitary at the ends of the branchlets. Phyllaries 5, keeled at the base, membranous, sub- tended by 5c estat bractlets. Receptacle naked. Achenes 5-angled. Pappus persistent, the bristles 5, rigid, branched from the base and also about one-half their length into slender a ra the ulti cae aes oT and surpassing the involucre. [From the Greek =~ meaning bristle and s he western = of the Great Basin, 1. Chaetadelpha wheéleri A. Gray. Chaetadelpha. Fig. 5981. Ew Chaetadelph heelert A. Gray, Proc. Amer. Acad. 9: 218. 18 Glabrous peer bags 4 branching above, 1-3 dm. high. eg i glaucous, 2-5 cm. long, es h; phyllar Feds igh, the subbending calyculus of 5 bractlets 2-3 mm. long; flowers ordid-white or ak : 2 ial £10 mm. long, the 5 pappus- Srtsties cen. as long as or ae ea the achen Desert valley Satcas Upper Sonoran Zone; southern Malheur County, Oregon, south through western donc a hg adjacent Inyo County, California. Type locality: ‘Southern Nevada.” Collected by Wheeler. 156. LYGODESMIA D. Don, Edinb. New Phil. Journ. 1829: 311. 1829 Perennial or annual, glabrous or pubescent phe: branched, and in 1 species spinescent. Leaves linear and entire or the lower and basal ones broader and sometimes pinnatifid, the uppermost sometimes rice to ee a werk pink or purple, solitary, and erect at the ends of the branches or sometimes racemose. Involucre cylindric, the phyllaries 5-8, equal, at the base. Style-branches slerder. Achenes beakless, few-ribbed, cylindric or nearly so or angled and linear, the pappus of eilary ‘beictles. [Name from Greek words meaning pliant twig and bundle, from the numerous branches. ] mS a genus of about 6 species, natives of western and Bic North America. Type species, Prenanthes juncea urs Perennial, with rigid spine-tipped b hes; pappus-bristles tawny, 6-10 mm, Jong. 1, L. spinosa, Annual, the very slender branches not spine-tipped; pappus-bristles white, 2-2.5 mm. long. 2. L. exigua. 1. Lygodesmia spindsa Nutt. Thorny Skeleton-plant or Spiny Lygodesmia. Fig. 5982. Lygodesmia spinosa Nutt. Trans. Amer. Phil. Soc. II. 7: 444. 1841. Stems several from a branching woody root-crown, with oo tufts of wool at base, om hie 5- fice rose-colored, wet exserted; achenes mm. long, d an be Hy lightly. fie Haha toward the apex ; pappus tawny, 6 mm. Jobe short-plumo ore. Dry plains and = Treg 8 slopes, y Upper Sonoran Mon and British a “rr to porthern Popes Sao } occu g in the Pacific, ‘Secs in ag tay oo otk Malheur Counties, rere nia i. | o east of the at id Nevada to Es my Gabriel (Swartout Canyo pie a Ber- nardino GBalaetn Lake) Mountains. Type locality: ‘In the y Mountain plains towards California.” June— 2 Lygodesmia exigua A. Gray. Annual Lygodesmia or Egbertia. Fig. 5983. th. A. Gray, Smiths. Contr. 5¢: 105. 1853. Sendvsnid octane A. Gray, Proc. Amer. Acad. 9: 217. 1874. Stephanomeria minima M. E. eg Conte. West. Bot. No. 17: 31. 1930. SUNFLOWER FAMILY 569 Slender, diffusely much-branched annual, glabrous or the stems sparsely eens eco tent elow, the branchlets slender and divergent. ower leaves oblanceolate or spatulate, 5-20 m (9) Oo olte ong; flowers gules abou . long, rather deeply lobed, ae e ae whitish; achenes 3-4 mm. long, oak ie Lspeanay he iscsi pete rather rigid, bright white. ane ledges et eter aoe a Zones; Moja sah) ste Colorado sia. southern California east to a = Texa: as. e loc : El Paso, Texas. "Apr 1M mia jincea UP urs an D. ake. Edinb. New Phil. BR 1829: 311. pec (Prenanthes agers ea Purs' FI. Pressey Fees eae “I8i4 ) iE, ose umes ee be denser sharia a om a deep-seated, creeping root and Gearine no sera tufts of w y be ted in rn Washington, as it occurs in British Columbia oa Idaho and Montana, but no cee Wires haere b ay n. 157. STEPHANOMERIA Nutt. Trans. Amer. Phil. Soc. II. 7: 427. 1841. Nomen conservandum Annual or perennial, Riche _osmeton — often glaucous herbs, sometimes woody at the base, wit th erect, simple or branched s Leaves alternate o r basal, entire or runcinate- pinnatifid, those of the ‘cantine often ses and scale-like. Hea ids subsessile or pedunculate in panicles or solitary at the ends of the branches. Involucre cylindric or oblong, the phyl- 5983. Lygodesmia exigua 5984. Stephanomeria cichoriacea 5981. Chaetadelpha wheeleri 5982. Lygodesmia spinosa 570 COMPOSITAE laries few, mostly in 1 series, equal, scarious- margined, slightly united at base, subtended by several calyculate bractlets, thin and withering in age. Flowers pink, flesh-colored or in the morning i A western North American genus of about 15 species. Type species, hee minor (Hook.) Nutt. Phyllaries 10-15 mm. long; leaves (3 .5— m. long) present on midstems at flowering ti Receptacle deeply pitted, cea ae sabe sessile or subsessile on elongated naked branches of ae lc il cte, acea. Receptacle naked; heads terminating the branchlets, corymbosely or racemosely arranged. Leaves thin, entire or saliently toothed. : 7: lactucina. Leaves thick and usually callous-margined, runcinately pinnatifid. S. parryi, Fugilaries 6-9 feare) y 19). mm es longs, leaves mostly bract-like on midstems at flowering time or ae minutely rugulose. Pappus-t Ainiged plumose to posik base or nearly so; plants mostly herbaceous at the base, the stems with ending branche 4. S. tenutfolia. Pappus>ristles sionlttacts hae hens one-fourth - more of their length and long-plumose above; plants y below and divaricately hranching abov Plan iat bro; 4 5: oe ext Plan OF Geuaety't omentulose. . S.cin Annuals or biennials; mature fertile achenes more or less tuberculate or rugose between 2 teleteodinal ribs. Papp gare ss mnate below in groups of 3 or 4, plumose to pois of attachment or poy 0 hirsutulous e loees one-half or one-third; sepa peduneulate (shortly so in robust forms of S. paniculata aint a Se . rape S. exigua and S. v ta). ps a stout; pedun —1.5 cm. ete nat aiding: clavate, a little onacnlaee. minutely his- pi ulous, 7 i 2 grooved between the angles 5 Ore paniculata. Stems slender; peduncles 1-4 cm. long; achenes linear, ghsbrowa; angled, and erooves evident. exigua, Pappus-bristles free to the base and deciduous, ne ronng just above the base leaving a minute crown, plumose throughout; heads typically subse: 9. S. virgata. 1. Stephanomeria cichoriacea A. Gray, sane leaved Stephanomeria. Fig. 5984. Stephanomeria cichoriacea A. Gray, Proc. Amer. Acad. 6: 552. 18 Perennial from a long deep taproot and somewhat ae the stems erect, simple or virgately branched, 6-15 dm. high, stout, glaucous, glabrous or commonly more or less woolly-pubescent especially when young. Basal leaves spreading on the ground, ubeinate: fniAAned, spatulate in outline, 8-16 mm. long, narrowed into Sigrenl stem-leaves sessile and clasping or auricled, the upper smaller, lanceolate or oblong, entir mmonly irregularly and saliently toothed; heads numerous, mostly on short ramets: Dedintcn” along the stems, mostly -flowered ; corolla pink, r. arely white ; schene s smooth, striate; pappus sordid, ahaa throughout, 10-12 mm. long. Dry washes and slopes, ae Sonoran a d Tra ansition Zones; southern Monterey and San Benito 1 angggone and de - Pass south to the souther = open of the San Rernsidinn hagintaina and the Santa Ana Mounta S. z Island, California. Tyee locality: Fort Tejon, Tehachapi Mountains. Aug.—Dec. Tejon Milk tae. 2, ee oe A. Gray. Large-flowered Stephanomeria. Fig. 5985. Stephanomeria lactucina A. Gray, Proc. Amer. Acad. 6: 552. 1865. Pillorie fateh Greene, Pianta e ays: 1 Dp a! with slender, deep-seated, creeping rootstock, the stems abit singly, preoching near the base with the slender branches spreading, or strictly erect with ascending bra cm. ‘hi gh, glabrous or ee y puberulent. Leaves linear, 2. cm. 3 ag —-7 mm. ae more 0 + ag attenuate at apex, entire or more or less sparingly runcinate-denticulate; heads terminating a or escteate slender branches and someyhat co imes somewhat race- socal involucres narrowly campanulate, the inner phyllaries linear-at m. long, the few outer a 5 mm. long, about twice the length the calyculate bractlets, pale green or rose-ting a gt usually 7 or 8, the ligules well exserted, rose-colo or og." -purple ; achenes 5-6 mm. long, ribbed, glabrous, light brown pappus-brist tles 9-10 mm. lon abo ut 20, white, chnsone ¢ deurticut, slightly broadened and somew vhat nena in Hair oO r 6. Sandy flats and open pine forests, Arid Transition and _ n Zones stern slope of the pele Moun- tains from southwestern Jackson County and Deschutes Cou mts, Omnia gon, msec aa gh the Cascade Moun- tains se Sierra Nevada to Mari sa County, California; Pin os the Siskiyou Mountains i Mount Sanhedrin in the North Coast —_ Califor ornia. Type locality: “Dry hill near Big-tree Road in the Sierra Nevada, alt. 6,000 Pal July—Aug a, A scape’ sag parryi A. Gray. Parry’s Stephanomeria. Fig. 5986. — parryi A. Gray, Proc. Amer. Acad. sa 61. 1883 ta par iaierinss Contr. *O. S. Nat. Herb. os 9 Perea 8 ae glaucous-green and ge tan throughout, the stems rather stout, erect, 15-30 cm. high, widely branching from near the base, the branches ascending or mor ommon nly di- varicately pinta ng. oe linear-lanceolate, rather thick and firm, deeply runcinately_pin- natifid, toothed; the margin somewhat calloused, anes or sparsely "pubescent beneath, 2.5-4 SUNFLOWER FAMILY 571 cm. long; heads terminating the ages = the short, lateral, bracteate branchlets, 10-14- wered ; inner phyllaries 10-12 m ong, scarious-margined; corolla pale pink to white, well exs sa ; — Sy ristles 7-8 mm. oe ag, long-plumose above, barbe ipa te .at pws te widened, deciduous in groups of 2 or 3; achenes i 9 se een ow rugulose ribs. Sandy and sae slopes, pee Firma a Zone; Inyo and Panamint teier Inyo County, and western Mojave i ahary California, east to ae n Arizona and southern Utah. Type ‘locality : ‘near St. George, S: Utah,” May_— July. Parry’s Rocke 4. Stephanomeria tenuifolia (Torr.) H. M. Hall. Narrow-leaved Stephanomeria. Fig. 5987. Prenanthes ? tenuifolia Torr. Ann. Lyc. N.Y. 2: 210. 1827. ih tenuifolia Ra f. Atl. Journ. 145. 183 L r Ho k Bor. ue i: 295. pl. 103, fig. A. 1833. Phil. Soc. IT. 7: 427. 1841. ¥ Nutt. Trans. Am Ptiloria filifolia Greene, scree a 1, ‘* 898. Stephanomeria tenuifolia H, M. Hall. Univ. Calif. Pub. Bot. 3: 256. 1907. s herbaceous, pale green and glabrous, 2-5 dm ii erect from a perennial root, with several to flexuose, ascendin ng branches. Lo ves narrow, runcinate-pinnatifid, about 5 cm. long, the upper leaves aaar, Prosi si somewhat shorter; heads terminal, ae j eae \ a y 436) as 5987. Stephanomeria t 5988. Stephanomeria jaaatitiers 5985. Stephanomeria lactucina 5986. Stephanomeria parryi 572 COMPOSITAE ered ; involucre 8-10 mm. long, the phyllaries eal 5; achenes 5-angled and grooved, the pappus- Sristics 15-25, white or sordid, plumose throughou Plains and jon mountain slopes, Upper Son and Transition Zones; southeastern British Columbia south to Modoc Cou California; Bg Moola ae pa Cclncalee. mene northern Abicsna, Type locality: ‘“‘near the Rocky Mountains,”’ “Collected by EP ames. June-Aug. A form differing from = name-bearing taxon ey having divergent branches and shorter stature has bee tg in the La ken Tahoe regi n Nevada and California ohed also at Sonora Pass, Tuolumne Caasity. and = H ats 5. ia The name Pislona Gewgns was applied to this entity by Greene in C. F. Baker, West Amer. PI. 1]: 19, 190. meria tenuifolia var. myrioclada (D. C. Eaton) Rrongisint, Leaflets West Bot. 1950. (Stepha momerte myrioclada D. C. Eaton. Bot. King Expl. 5: as pl. 20, tsi AP TST 1s Aa ngs tenfolig myrio- clada Blake, Contr. U.S. Nat. Herb. 25: 623. 1925; Stephanomeria paucifior a var. myrtioclada Mun S. Calif, 589, 601. 1935.) Plants 1-3 dm. high of densely crowded, slend ems arising from a woo nape pont ‘linear, filiform; involucre 5—6 mm. high; phyllaries 3-5; achenes as in oe poe Arid Transition ee Canadian Zones in the desert ranges of Nevada and Wyoming westward to the eastern fers arg of be Cascade Mounta Oregon south to the southern Sierra Nevada, California. Type locality: ‘‘Thousand Spring and ‘Groce Creek Wileré: [Elko County] Nevada; 6—6,500 feet elevation.” 5. Stephanomeria paucifléra (Torr.) A. Nels. Few-flowered Stephanomeria or Wire Lettuce. Fig. 5988. Prenanthes (?) pauciflora Torr. Ann. Lyc. N.Y. 2: 210. 1827. ig nee peuciiane Bat Atl. Journ. 145. 1832. rythea 1: 224. Steph A. Nels. in Coult. & Nels s. New Man. Bot. Rocky Mts. 588. 1909. Stephanomeria — Eastw. Titles West. Bot. 2: 55. 1937 Perennial, woody at the base, the several stems divaricately and intricately branched and more or fest rigid, ipieting rounded, glaucous and glabrous bushes 3-5 dm. high. Lower leaves runcinate-pinnatifid with narrow segments, the upper entire and spreading, with tufts of wool , or reduced ales, these also sometimes ; he olitary, terminal on the nd also scattered along the branchlets on short bracteate pedu , dt red; in- volucre 7-8(10) . high, Be Ss ; ri s grooved between the 5 ribs and rather incon- spicuously transversely r €; pappus what tawny, the bristles rather firm ie ee from the base for about ne Honrth their ek and long-plumose above, deciduous in g Sandy or ss goats desert slopes, Sonoran Zones; western Kern County and the “Mojave and fate iauite: Pb ag eastward through Nevada sa a > Kansas and south to northern Son and Texas. Type locality: “near the Rocky Mountains.” Collected by E. P. James. April—Oct. Desert Straw 6. Stephanomeria cinérea (Blake) Blake. Gray Stephanomeria. Fig. 5989. Sena cinerea Blake, Proc. Biol. Soc. Wash. 35: 177. 1922. a eria rene var. tae eer ae Fl. Pl. sei 998. 1925. fr; iy A tshit Angee Aue: 42 100. ie vie renni ial, sie at base and divaricately much-branched above, densely cinereous-tomentose throughout, 3-4 dm. high. ae es of the midstem linear-lanceolate, acuminate, runcinate-toothed and ed to above; ds pagel at the tips of the branchlets or on sh peduncles, out 5-flowered ; phyllaries 5, 7-8 h, subtended by calyculate bractlets ; achenes 5-angled, whitish, finely ie daha oi rugulose when mature ; shee Mong ete out 14, somewhat tawny, his- pidulous at the extreme base, plumose above and deciduous in a r Hig Gravelly soil, a Sonoran Zone; Death Valley region in ten County, Slag steer and adjacent Nevada Ene Mg “Lancaster, Los Angeles County, California, in the Mojave Desert. Type locality: ‘‘Pahrump Valley, Nev: 0-915 meters.” Tone) ‘uly. F “Mich pinot th age western desert fore of S. paucifora and differing principally in the cinerous tomentum of the stems, Z. Steppe erin acne a Nutt. Stiff-branched Stephanomeria. Fig. 5990. Steph i Nutt. Tra gi sep Phil. Soc. If. 7: 428. 1841. Ptiloria paniculata me Pittonia = 1890. Stephanomeria oregonensis Gandoger, Pal Soc. Bot. Fr. 65: 53. 1918. Stephanomeria suksdorfii Gandoger, loc stems usually a rather octet B 5 tiie | eee above, 3-6 dm. high, gla- e som p us t thro leaves narrowly oblance wing eae’ or decticuiste: the otis narrowly linear, eae red ced and bract-like in the inflorescence; heads on short divergent bractlets th for narrow or sometimes more widely branching panicles, 5—7-flowered; phyllaries 5, ‘tine ear, 5-7 mm. long, Edbtesicled by caly i ight pink; achenes parey 5-angled, the intervals tuberculate; pappus-bristles 1 , united in groups of 2 or 3, white or more commonly tawny, rather short- plumose above and becoming di — near the base Dry hillsides and plains, especially in open sagebrush, Arid Transition and Upper Sonoran Zones; Wash- Soe east of a Cascade Mountains cad Idaho southward to eastern rene and northern California. Type “On the Rocky Mountain pinins, towards the Colorado.”” July—Aug 8. gecdeaaces exigua Nutt. Small Stephanomeria. Fig. 5991. Stephanomeria exigua ns. Amer. ae Soc. IT. 7: 428. 1841. Ptiloria exigua Greene, on toa 132. 18 Annual or rarely biennial, 1-5 a high, commonly fastigiately branched from the base in SUNFLOWER FAMILY 573 older or more vigorous plants, occasionally nearly Baia with paniculately ascending branches ; stems usually slender, pale bluish green, slabrows with a few minute glandular hairs. Lower and basal leaves narrowly oblong, coarsely to bed or runcinate-pinna atifid, 5 cm. or less long, the upper feaves small and bract-like; heads at the tips of the branches, more rarely clustered on : cr mm, c angles, the intervals with 2 long itudinal rows of tubercles; hcp Aca 8-18, white, plumose above, the lower third naked, ere aking off toward the base, more or less adhering in groups of 4 or 5 by the widened bases, these often with minute marginal bristles. Brushy slopes and in forests, Upper Sonoran and Transition nes; Wyoming, Idaho, and eastern Oregon south to New Mexico and California on the Gunes face of the signe Nesaie: aad the mountains of southern California; also on the western face of kat ape ra Nevada in Kern County and near the coast in southern Cali- fornia and Lower California where sever 1 diverse forms ben to be found, — of which appear to be a ego —o S. exigua, S. exigua var. “deane 1, and S. virgata. In som es -bri Aaa gg tend = shatter readily, leaving an iseulay crown on the sehenes formed by the brcudeaed and pasate as nited nonplumose bases of t he iy les. These forms have aie interpreted by authors as S. exigua var. coronaria tisccene) Jepson ‘Tian: i, . Calif. 998. 1925.) by t are different from S. coronaria aed a plant of the c aren = Spee of CAtfornia which r s part o he s. virgata complex. Type loc + On: the os ky Moun plains, towards the Colorado.”” July—O Siesickcaueie aries var. pentachaéta (D. C. Fate, H. M. Hall, Univ. Calif. Pub. Bot. 3: 260. 1907. rag rte Mg eta D. C. Eaton, Bot. King Expl. 199. pl. 20, fas . 8-10. 1871; Ptiloria pentachaeta tto: Loar Lee ° mh co ° - 3 w ae i-" a < ES Ss a of as o = w a ° e, base but deciduous i 2 oe ups, plum Tate on the’ ‘upper malt bea eg discal at ‘the base fee usually with a few minute teeth. Dry desert regions, otis rney and Malheu , southward east of the ak ipa + Pipi SOO corer’? = ae 5989. Stephanomeria cinerea 5990. Stephanomeria paniculat: ia Stephanomeria exigua 992. Stetphanomeria virgata 574 COMPOSITAE Cascade Mountains and the Sierra Nevada to Inyo County and the Genexts of southern California and adjacent Lower California; also Nevada, southwestern Utah. and lacing estern Arizona. Occurring with S. erigua var exrigua, ba hacthy coe “Truckee and Humboldt Valleys,’’ Neva St na exigua var. déanei J. F. Macbride, yey Grey Herb. No :22. 1918. (Ptiloria exigua var. deane Fr. Machriie Foss Davids. ea smog y; is hn Calif.: 355.1923.) Sle Mii ‘annual, divas ivete-penigciees with bracteate _ pecunaies rsisting e than season and becoming intricately branched; m or less glandular e lally in the infor leiconet: yeah pee + in the oe intervals of the achenes stro cate grooved and little or ot at all tuberculate; pappus- — as in 9 species, often breaking off i ievepudarty gee the base. Hill slopes, Sag Diego County, California, and mountains of yer Lower California; naerar ese ve the north with the slender nonglandular form of Ky exigua i I a the achenes are slightly rugose and strongly grooved, Tene locality : Sweetwater Valley, San Diego Cou 4 Stephanomeria virgata Benth. Virgate or Tall Stephanomeria. Fig. 5992. St ta Benth. Bot. Sulph. 32. 1844. paiement ehaie Nutt. Journ. Acad. Phila. II. 1: 173. 1848. Stephanomeria coronaria Greene, Bull. Calif. Acad. 1: 194. 1885. tomentos 886 en eet, Ptiloria canescens Gr iid Pittonia 2+13%, eign pleurocarpa Gree e;, loc cae Munz, Aliso 4: 100. 1958. aceuat Seite.” erect with rigid stems of reaiely branching above, 3-10 dm. high or rarely dm. ; erbage u sually Fe ie throughout, in some forms tomentose. Lower leaves oblong o spatulate, often sete or shallowly pgs soon deciduous, the upper ones linear and mite ea solita i icle s with = without an evident ates: meen strongly rugose to nearly smooth; pappus-bristles white, ong, copious, a mber, fragile, the plumose hairs usually a little shorter ehvlaerd | the base, completely deciduous o or sp ‘base es of the bristles in some forms remaining on the achene as an extremely short, mn crow Dry hills and alice Transition | and Sonoran Zones; southwestern rogue southward to northern Lower California. ng oat San Pedro, Los Angeles Counts: California. Bi ae : Future field studie ay prove that some of the synonyms listed under thi * variable taxon mn may hav taxonomic status. ieerent aa are sometimes locally recogniz able “but intergrading forms are cocitentiy ge be found even in the same re ; The photograph of the ty *, S. virgata var. virgata from San Pedro shows a plant wit th hy Be irgate- eerste inflorescence in which the hemi are subsessile nag fascicled along the stems, a form which is found rather com- monly northward along the coast. The enes are angled, slightly incurved, and rugose- tuberdclate, es usually dark or mottled. Apparently they are > tne not grooved on the faces between the an ngles as this character is not ag ed in the careful original FSi ew The of S. elata from Santa Barbara has not bee As interpreted from the original description it tideres A "ibe ca virgata complex ba t differs ola the sia form in the ‘ nae ore te in which each of the “flower ~~ anches . ar three or four pileb ale ” The blue color gned to the ligules om = original description was ev idently not, obtained from field notes but was doubtfully applied to the dried ieee by Nuttall. Some specimens are to be found in co Hectiaae frees the Santa Barbara region in which the heads are definitely pedunculate in an chiens t panicle rather Lege clustered along the branches of a virgate-panicalate inflorescence. The achenes of some of these s apeci mens are pale brown, 5-groo okey po scmemtak ose, all are attached he base in ethan of 3 or 4, not plumose to the hase, and break one ather een, other tearaeiee, absence of beamed a es angles, is met with sporadically cheokghus the species. Also iy areuers® having the short cro athe achene have grooves and low tubercles rather than smooth faces on the achen Stephanomeria tomentosa, described ie Greene from Santa Cruz Island, Santa Barbara County, is described as a stout plant having the inflore escence and achene characters ef S. virgata be vse igs as “‘white hoeseniuer throughout when young the i aiarunceolee glabrate.” This type of pubesc pace of soft, er multi- cellular, unbranched ea ie is ‘fea in varying abundance in other spec oie tise islands and adja cent fe and. oe can even be demonstrated on specimens having the ‘ ‘elata’® type oF alien which are found in same tiloria mescens, described from Napa Valley, and P. pleurocarpa from Redding, nasa Counts, as_ well other roadhg of the complex, occur he hout northern and central Pwitocts and the S ierra Nev a to he rs quite as often on plants es many heads on virgately disposed branches. Ptiloria sletecuehe is distneuianea se from Stephanomertia virgata var. vi poate by pale brown achtn es. 158. RAFINESQUIA Nutt. Trans. Amer. Phil. Soc. IT. 7: 429. 1841. amis and slightly ae * peed annuals with fistulose stems and pinnatifid lea Heads rather large, with white or rose-tinged flowers. Involucre conic or cylindric, with h 7-15 linear acuminate pylaries, somewhat fleshy at biog with a few 8 shape outer ones. Achenes terete, somewhat fusiform, obscurely few-ribbed, uate into beak. Pappus white or cet of 8-15 slender bristles, these softly a from the base to near the tip. [Name in honor of Consiantine Rafinesque, an American naturalist x0. aia eler.] A genus of 2 species, natives of southwestern United States and adjacent Mexico. Type species, Rafinesquia californica Nutt. Rays about 5-8 mm. long; beak of the achene as long as the body: pappus capillary and 1 ical ae straight Rays about 15 mm. long: beak of the achene shorter than the body; pappus flattened at base, See se with ‘arach- noid hairs, these sometimes lacking at the attenuate tip. SUNFLOWER FAMILY 575 1. Rafinesquia califérnica Nutt. California Chicory. Fig. 5993. Rafnesquie cloacae seh Trans. Amer. Phil. are Il. 7: 429. 1841. Greene, Pittonia 2: 193. 189 ms stout, a3 dee, high, ool ‘hook or sometimes branched above. Basal leaves coarsely to othed, soon withering ; stem-leaves oblong ba worse eolate, auriculate- siaaping dentate to Scteapesestenell d, the divisions rather broad and n _— the —— the upper leaves much reduced; heads several in a paniculate- cep ome a S anericen m. hi a ‘i igules 5-8 m long, white ; outer achenes pubescent and somewhat uderenians! beak slender, equaling the bo o pappus dull white or sordid, plumose with steeight hairs to the top of the bri Foothills and ebay: ee Sonoran Zone; Humboldt and Mariposa sacthow ies ricer 0 coast: California and norther r California; occasionally eastward at higher elevations to the Sescua wee Colorade Deserts and es tah yee ona. Type locality: “‘Near the sea-coast in the vicinity of St. Diego, Upper Cali- fornia.”” April—July. o sinneei neo-mexicana A. Gray. Desert Chicory. Fig. 5994. R A. Gray, Smiths. Contr. 56: 103. 1853. Na emoseris neo-mexicana Greene, Pittonia 2: 193. T6931; Stems simple or branched from Ay base, 1.5-5 dm. high, often purplish, rather weak and usually jaaneobted by desert shrubs. Basal leaves 3-9 cm. long, mostly runcinate-pinnatifid, acute at the apex, u sally per — during anthesis, the stem-leaves sessile and auriculate-clasping, much Ase gist abov and becoming feached te in the inflor rescence, deeply runcinate-pinnatifid, the nonincised p OE the feat spony. ne 6 lanceolate to linear ; flowers solitary at the ends of the branchlets ; igh arid about 2 cm. long, linear-lanceolate, ak aa ct , the outer calyculate bractlets much sho rter, ligules 1s. -20 m m. long, laciniate-toothed at the apex, white within, tinge with rose or purple without; achen 3 12 15 mm. long, obscurely angled, tapering abruptly into a beak shorter than. ie body of the ppl a surface npoearin grayish-mottled with a papillate puberulence, the pappus- Fang ors bright white, the bas mewhat flattened, plumose with soft arachnoid ha airs, hens often lacking at the inn of the ‘bri mre Mesas and canyons, usua my in the shade of shrubs, — Zones; Inyo County, California, south through the Mojave: gad feeeete Deserts to central Lower California and eastward to souther ice ey and through Ari- zona we Texas, and in adjacent Sonora. Type losabeee El Paso, Texas. March—-May 159. MALACOTHRIX DC. Prod. 7: 192. 1838. Annual or perennial herbs. Leaves alternate or basal, poangped Bim Heads long- dumeae panicled, or solitary; flowers yellow: or w ite or mes purplish- tinged. uo en ng, en rr 8 so " ~~ hair, in ict to ‘ie soft woolly nabencaee: on the leaves of the original s A wae ae American genus of about 15-20 species. Type species, Malacothrix californica DC. 576 COMPOSITAE pera chog 5-5 dm. high. Out phollaies well Pasicatiarns in 3-4 series, orbicular to narrowly or broadly ovate with conspicuous s margin Pappus-bristes 4, persisting on the achene; outer phyllaries orbicular, 4-5 ae pate of desert arid a co feneiie blatlo ae persisting on the achene; outer phyllaries bilan Piet 2 mm. bro: vai plants of Channel Islands. . M. thaiiadie fatale Outer sohlasaea? in 1—2 series, the outer much shorter than the i a with narrow scarious margin Stems leafy to the inflorescence; leaves not much reduced abov: gpsinsinl erect; leaves laciniate-pinnatifid. 3. M. Plants depressed (or erect) ; musts n the oh obtuse. 4. M. harose Leaves mostly basal, reduced and sparse abov Basal leaves pinnatifid with a ona filiform lobes; flower-heads large, the ligules 12-16 mm. long. 5. M. californica. Basal ald al pinnatifid Brey short, broad, mostly toothed lobes; flower-heads medium to small, the i 2-10 mm pie. pod" 4-7 mm. high; achenes finely 10-15 striate with the 5 ribs not markedly more promi- fo, about 2-3 mm. long; achenes with > tues denticulate crown and 1-2 persistent bristles; leaf-margins without tufts of w 6. M. clevelandii. Ligules 3-5 mm. long; achenes with an entire crown and no persistent bristles; leaf-margins with tufts of wool. vs M. floce ifera. —- 8-13 mm. high; achenes 15-striate, ever ib es 2-3 mm. long; ee, a all alee jntieimleie essentially aiabsaws. . M. sonchoide. ays boa 4 sation long; 1-8 pappus-bristles persistent; ee eta a Pg gland- ped 9. M. torreyt. Perennials; shrubs, stout kaha aceous plants or the more or ee ene forms from a woody root-crown. igules yellow; dune plan pied is sebeng ag spina from a 10. M.4 Ligules pink, rb: or V ect oc tanesine plants Heads wt Mn A eats herbaceous or shrubby ony at base (except M. savratilis var. implicata); leaves not obovate or oblong-obovate. Shans ‘Sechenous, — sisal from a ate seated, branched root, the entire est on if present, acute or acuminate at the . M.s Stems several from a woody root, usually siitcboieiend at base; entire bicvet ue aan eat or broadly acute at : the apex (except in vars. implicata and ‘tenu ifo " vii . M. saxatilis. Heads 9—12-flowered; shrubs; leaves obovate or oblong-obovate. a M. blair 1. Malacothrix coulteri Harv. & Gray. Snakes-head or Coulter’s Malacothrix. Fig. 5995. Malacothrix coulteri Harv. & Gray in A. Gray, Mem. Amer. Acad. II. 4: 113. 1849. ne ee coulteri Heller, Mihlechefeee 2: io. 1906. ms branching above and sometimes also at base, erect or sometimes more or less decum- bent, US dm. high, glabrous and pale. Leaves sinuate-dentate or somewhat pinnatifid, the basal m. uline claspi i ; oblong or mea ny ong, the cauline clasping, ovate to lanceolate i - 1 : rminatin branches, short-peduncled; involucres hemispherical, 10-15 i phyllaries well cated, suborbicular to ovate, with broad scarious margins and a broad, purplish reen, central band; ligules pale, yellow to nearly , th ter ones so p (6) white €- times lined with reddish purple, 5-18 mm. long; achenes light greenish brown, 2-2.75 mm. long, 4-5-angled, with 2 striae between the sharply sive Ses these extended above the body of the achene in short ead: a ee ‘appus-briste eek Flats er Sonoran Zone; Saas a, San Joaquin, and Mariposa Counties, central California, south to northern: peed, Catifornia and east in the desert r regions to southwestern Utah and Arizona. Type ineaticg: California. Collected by Coulter. March-— May Malacothrix coulteri var. co 2 ta Jepson, Man. FI. Pl, Calif. 1001. 1925. Heads and phyllaries as in M. eoatert var. coulteri; stem-leaves pris in but waited nearly to the midrib into linear divisions; achenes and pappus-bristles as in M. coulteri var. coulteri. Ra rely collected; Santa ( Cruz and Santa Rosa Islands in Santa Barbara County, the San Pedro hills in phe ‘Raasies County (Abrams a), = Tia Juana, San Diego County (Jones), California. Type locality: Santa Cruz Island. Collected by Bran 2. Malacothrix insularis var. squalida (Greene) Ferris. Island Malacothrix. Fig. 5996. Malacothrix squalida gyi a Calif. Acad. 2: 152. 1886. aE. Williams, Amer. Midl. Nat. 58: 507. 1957. Malacothrix Santarls 9 var. Mais Ferris, Contr. Dudley Herb. 5: 102. 1958. Glabrous annuals, the stout ig decumbent or sometimes erect, 1.5-4 dm. high, quite densely leafy se counhiout, branched at base and above. Leaves sessile, 2-7 cm. long, broadly lanceolate to ovate in outline, with ent, Saaanied: acute lobes, the anaes upper leaves a oe, laciniate and clasping at the base; heads many, on short, at bracteate peduncles ; aot s 9- uter oO : oran es ete — Santa Rosa, Santa Bertone and Anacapa Islands off the coast of southern terraces, Upoe eraik. eres locality : Santa Cruz Island. Collected by Greene. April-July. SS Sout Baviars tcland appear to be quite close to M. insu lacie Wax, ino SUNFLOWER FAMILY 577 3. Malacothrix foliédsa A. Gray. Leafy Malacothrix. Fig. 5997. Malacothrix foliosa A. Gray, Syn. Fl. N. Amer. ed. 2. 17: 455. 1886. Glabrous annuals, the rather slender stems often purplish, erect, 1.5-3.5 dm. high, leafy throughout, usually simple below ip Dp rses branched above, rarely with r 3 branches ase. anceo : ious m mare , calyc late bracts, 2 or 3 of these sometimes about one-half at’ length . the inner phyllaries; ligules bright yellow, surpassing the inner ohyllaries ne 2. —-3 m achenes 1.2-1.3 mm. long, o obscure ely angular, 12-15-ribbed or striate, “ce ibs cele of ‘equal width, aie on the angles not noticeably larger; pappus-bristles all deciduous. Hillsides and t es, Upp ae Zone; Santa Barbara, San Clemente, and Coronados Islands off the coast = southern California oat "hort thern Lewer California. Type locality: San Clemente Island, Los Angeles County, California. Collected by Nevin and Lyon. April—July. 4. Malacothrix indécora Greene. Fleshy Malacothrix. Fig. 5998. Malacothrix indecora Greene, Bull. Calif. Acad. 2: 152. 1886 Malacothrix foliosa var. jadocore E. Williams, Amer. Midl. Nat. 58: 507. 1957. Low, leafy annuals 2-4 cm. high, at first single-stemmed, becoming much br anched from the base, pete depressed. Leaves gues us and succulent, sees or often oblanceolate, more or less pinnatifid with ‘oie obtuse lobes, the lower 2-3 cm. long, the upper about half as long; heads Sty. P57, 5995. Malacothrix coulteri 5996. Malacothrix insularis 5997. ote foliosa 5998. Malacothrix indecora 578 COMPOSITAE on short bracteate peduncles ; involucre 6-7 mm high; inner phyllaries linear- lanceolate, pe greenish, the outer “area nd p urple-tinged ; ligules shorty greenish yellow. achenes about 1 m long, 5-angled and 2-3-st hal hi n the obtus e angles; pappus dec hi na ring with no date er ent bristles, the peiiea tte ee cilate at base, barectine abavey Fadeplees arith minute palea An insular species, occurring on Santa Cruz and San Miguel Islands, Santa Barbara County, and San ingen Island, Ventura County, off the coast of southern California. Type loc ality : Santa Cruz Island. April-Sep 5. Malacothrix califérnica DC. California Malacothrix, Fig. 5999. Malacothrix californica DC. Prod. 7: 192. 1838. caulescent scapose annual. Leaves basal, espn Se dense rosette, Fpeanchegs 6-12 cm. long, the lobes na arrowly linear or ep atin filiform, more or less Bageort tbs oolly- ntose when young, partially glabrate in age; scapes usually several, 10-35 ¢ high, opt or ; vimewtin decumbent at labrous, simple and bractless, bearing a rt al A head with pale yellow ligules; involucres broadly campanulate, m igh, densel fe) se; outer phyllaries nar- 1 t uch er and linear-lanceolate, with broadly scarious ma ; receptacle-bristles usually present but delicate ; achenes about long, s e nar r and e base, 4-5-ribbed, with 2 slender striae between the ribs: ; pappus white, about 8 mm — ng, the 2 outer bristles persistent, the inner ones united at base pe falling away together. r gravelly soils, Sonoran Zones; inland and along the coast, Contra Costa County, and Fresno and Tulare Wane south to Lower California; also on the Wwebter edges of the Mojave and Colorado Deserts. Type locality : ira sg: Collected by Douglas. arch— May hrix ca atte var. glabrata A. Gray nD: C. Eat Bot. King Expl. 201. 1871. (Malacothrix reer Syn. Fl. N. Amer. 12: 422. 1884 2 ‘Growth habit, ‘and Lig of M. californica var. californica; a gahrons throughout, rarely a little woolly on young ba “ee ; flower ring stems bearing 2 or 3 "aces heads (ra 2): 1 or 2 canine leaves sometimes present; aches pli asin M. ay Sal eal var. californica, Sandy ae Deathereatecl Idaho and eastern Oregon a and Nevilis idle the deserts oF Lalit nia and Arizona to Lower California; occurring rarely with the species in the San Fossum Valley in ae pee nty, California, and eastern part of the Cuyama Valley. Santa oa bara County. Type locality: 7 eal City (Anderson!) Foothills of the Trinity Mountains, Nevada.” Mare h-July. Desert Dandelion. 6. Malacothrix clevelandii A. Gray. Cleveland’s Malacothrix. Fig. 6000. Malacothrix clevelandii A. Gray, Bot. Calif. 1: 433. 1876 Annual, the stems usually several from the base, paniculately branched above, the branches ascending, 1-4 dm . high, glabrous, often reddish. Leave s mostly basal, more or less. Lira 2-10 cm. u teeth 3. ste. wid “ye e many, e slender paniculate branches ; involucre narrowly campanulate, 67 mm. high, the in “i igllaries i n, O i d i y about 2 m long ; achenes linear, minutely striate-costate; pappus rat deciduous pir rgd oy 1 (erely 2) rersictent bristle (this 1 usually oe deciduous) and a circle of minute white teeth. Chaparral and open Ayre espec n burns or disturbed places, mainly Upper Sonoran Zone; occa- sional in the foothills of the Sie 5 ora California, from ipisrahan County southward and the Coast Ra anges from Glenn County to gathers. Califo orets, wher more and nor Haan x teat Catitecata: also on Santa Cruz Island "(Brandegee) and occasional in rie gs ns Ag pny Mase Deserts to southwestern Nevada and Arizona, and Sonora. Type locality: dar Sax Diego. April-June. a pica ne (DC.) Blake. Woolly Malacothrix. Fig. 6001. Senecio flocciferus DC. Prod. 6: 426. 1837. reaengio 1ae4 ottnat Ben x =e Hartw. Poi 1849, ora Benth Pout Soneihive — Pace U. S. Nat. Herb. 22: 656. 1924. Annual, stems simple at base and erect or often several with the outer somewhat decumbent, ba ie ee aad above, 1-4 dm. high. Leaves nearly all broad, oblong or oblong-spatulate, 1.5-10 ing t 1 long, pinna “5 or the smaller often merely dentate, often bearing tufts of wool the lower surface peciall the lobes; involucre narrowl camp ofe Oe : ad ; inner phyllaries 4-5 mm. high, linear, acute or short-acuminate, with broad s mar gins, often i h purplish at least at the tip; ligules white or sometimes pale yellow, Seen: tinged — pink, 5-10 mm. long; achenes oblong-obovoid, entire at summit ; pappus-bristles all decidu Dry slopes and rocky banks, Upper Son moran and Tranisien Zones; Coast Ranges from ad sdvott County to poor ad Pinos, Ventura County. trattear nia and in the Sierra Nevada gs ses Concle to Mariposa County; tern Nevada. Type locality: California. Collected by Doupite:: April-Oct. 8. Malacothrix sonchoides er. pan & Gray. Sow-thistle Malacothrix. Leptoseris sonchoides Nutt. Trans. Amer. Phil. Soc. I hs 7: 439. 1841. Malacothrix sonchoides Torr, gre & |. N. Amer. 2: 486. 1843. Be make runcinata A. Nels. Bull. Tocca Club 26: 485. 1899, ual witha single erect stem or with several stems and somewhat decumbent at base, eee ing ork 1-5 dm. high, glabrous or essentially so. Leaves in a basal tuft, rather thick, 3-10 c lon long i in outline, regularly pinnatifid, the lobes and teeth callous-tipped, lower — sir few, similar to the basal, the uppermost reduced to small subulate bract cts; heads few to numerous, short-pedunculate ; involucres s campanulate, 7-8 mm. high, the inner phyllaries tispise ielabitate ; SUNFLOWER FAMILY 579 _ bright yellow, about 1 cm. hg achenes with 5 prominent ribs and 2 finer ones in the rvals ; ibe fete all — andy o avelly slopes and w vdesanh Zones; desert a of Inyo and San Bernardino Counties, California, oe pt ramen oe Great a region to western Nebraska. Type locality: ‘“‘The plains of the Platte. a pri 9. Malacothrix torreyiA. eey. amet Malacothrix. Fig. 6003. Malacothrix torreyi A. Gray, Proc. Amer. Acad. 9 Malacothrix sonchoides var. torreyi E. Williams, Anas wea Nat. 58: 5030 1957: Annual, the stems simple below and erect or often branched at base and somewhat decumbent, 10-30 cm. high, herbage glabrous or usually spar quit puberulent with gland-t ipped hairs. Leaves 2-6 cm. long, pinnatifid, the lobes divergent or run hg lower wing-petioled, 3 ee several to many, sessile or nearly bis with the lobes ent oothed ; heads ee pan ye gh seink slender peduncles ; involcre 7- m. high, the phyllaries ahr e, 16-20; r ligu gai ges ong; achenes about 4 m tee nen + DET keeled on the angles, aris ecun: sak sistent pappus-bristles 2- 5, white, 6-7 mm. long. Sandy or gravelly places, ae ‘Arid esac Zone; southeastern Oregon in Harney and Malheur Coun- ties, east to Idaho and Utah, and south to Nevada. Type locality : Salt Lake City, Utah. May-July. 10. Malacothrix incana ( Nutt.) Torr. & Gray. Dune Malacothrix. Fig. 6004. Malacomeris incanus Nutt. Trans. Amer. Phil. Soc. II. 7: 435, 1841. Malacothrix incana Torr. & Gray, Fl. N. Amer. 2: 486. 1843 Perennial with a gi rather deep-seated root, ee whole plant more or less densely white- woolly capectills when young, the stems branching from the root-crown, short and stout, forming tufts or mats, or Sreiches elongate, 2-3 dm. long, iti decumbent. Leaves spatulate, entire or some ating \ 5999 ) 6000 6001. 6002. Malacothrix sonchoides 580 COMPOSITAE what pinnatifid with a few broad lobes or teeth, 3-6 cm. long ait riers the winged petiole, 5-20 mm. wide, those of the stems reduced; peduncles several t to many, bracteate; heads 15-30 mm. b road; faetueres sis mm. high, attenuate a base; phyllaries in 4 or 5 5 eries, the upper linear-oblong. acute, those on the = in base much reduced ; ligules lemon i achenes oblong, 15-striate ; pappus- Take all deciduo On o ogre dun he coast, Upper Sonoran Zone; Pismo, San Luis Obi ispo County, pee ag wo to Carpinteria a hig adleets dnlands off the acon of Santa Ba rbara County. Type locality : age oa n the bay at San Diego, California, bg not since collected in that region. April-Aug. Some collections offman tiem Santa Rosa Island app var. succulenta. alacothrix inc - a var. suce rinse (Elmer) E. Williams, Amer. 58: 506. 1957. (Malacothrix ucculenta Elmer, Gaz. 39: 44. 1905.) Growth habit of the pal ‘ut yee os glabrous or the young leaves sometimes AoA beneath. On dunes along the coast, southern San Lui s Obispo County (Oso Flaco Lake) to Surf, Santa Barbara Connie, California, the type locality. 11. Malacothrix saxatilis (Nutt.) Torr. & Gray. Cliff Malacothrix. F ig. 6005. Leucoseris saxatilis Nutt. Trans. sy Phil. = II. 7: 440. 1841. Malacothrix saxatilis Torr. & Gray, Fl. N. Amer. 2: 486. 1843 Stems stout and fi ie erect or as mbent Sa a woody base, 3-7 dm. high, simple and very leafy blow: branched and leaf above, glabrous or somewhat floccose at the leaf-axils. L thickish an Rus o > it) [nad wn co a QO a oO =) co 3 i) i f Loe 9 of a2 ee 98 ie) H. M. Hall, Univ. Calif. Pub. Bot 268. 1907. (Mala- ly ce eae Nicolas, San Miguel, Seite = pig an aay ee Islands off the coast of San’ erhar ra, Cali- ocality: San Nicolas Island, Ventura County. sects i a = tenuifdlia (Nutt.) A. Gray, Syn. FI. Amer. Pt ie 1884. rr lag tes temuifola Nut, Trans. . Phil. Soc. II. 7: 440. 1841; Malacothrix fenuafoha & Gra . Amer. 2: 487. ae.” G ee eae: ipenissine unz, Man a lif. 591, 601. 1935.) Stems seve mt gt woody root, isa 4 igh, often gion oa | glabrous or som t floccose ‘throughout; ss Eas agg pinnatifid, upper stem-leaves reduced ste rie linear-filiform or ne parted into linear div fd ost evra form. Hillsides and cany coastal gages from the Santa Maria Ri oer. SSant am Bar bare Con to Orange County, Coliteruia, ‘and achlgeonage o the eastern border ced Los yetes County; ales Santa Cathlina sland. Type locality: “St. Barbara, on the ies untains near the tow Malacothrix saxatilis var. altissima dees Ferris. com = udley Herb. 5: 102. 1958. (Malacothrix altissima Greene, Bull. Calif. ng 4: 1885.) Leafy-st aes plants is m. hi - typically giabrous throughout; leaves thin, 8-20 c¢ ong, ifncauee and fecpiy Secale: lobed and also enticulate, the _upper- most leaves ys to be catite, | See ate at apex, tapetiae at the hips oe Mountains , Kern er ok westward to the unt Pinos re on, Ventura Poin A and the higher lopes o f the mountains in Santa Barbara County Tiovic ca Mountains in Los Aserles Comnee: Type locality: near Tehachapi rd tion, Kern County. Collected by Curran. Intergrading forms are to be found that show relationship with var. gheein o while others speiich var. comm othrix saxatilis var. arachno wort ‘(McG regor) E. Williams, Amer. Midl. Nat. 58: 509. 1957. (Malecothete arachnoidea McGregor, Bull, Torrey Club 36: 605. fig. 3. 1909.) Densely leafy-stemmed plants 2 m. high; herbage persistently and densely arachnoid-tomentose throughout; leaves firm, em, long, narrowed to a petiolar fae usually entire or marge imes denticulate, acute at apex; crown of the pee minutely crenulate. Hill slopes; known only from Carmel Valle ey, Monterey County, ap ince the type localit _.Malacothrix saxatalis var. commutata sess & Gray) Ferris, in Munz, Aliso 4: 100. 1958. (Hievactaos californicum DC. Prod. 7: 235. 1838, not sta a califor ornicum DC, vl cit. 192; Sonchus californicus pod a Arn. Bot . Beechey ope “se ae a 's californicus Nutt. Trans. Amer. Phil. Soc. II. 7: 441. 1841; alac commutata Torr. & Gray, Fl. N. eer 2: 487. 1843.) Raeely aeesy themenen plants 0.5-1m high, glabrous “si spars ad hnoid-t out the leaf-bases, midveins, and lower stems; leaves firm 4-9 cm. ong, narrowed at the base, mostly ‘denticulate, so metimes schice. iF og “un lobed these much shorter than the width the leaf, pot at apex; t ph Hobe paiesitely. crenulate. Salinas Valley, eastern Mead 2 the Santa Lucia Mountains in Mowers. on See fine tise Counties, and in a “onal Santa Barbara Cow The collections upon w hick the names listed = x sed were made by Douglas ogee Soe his journey from Monterey to Santa Barbara, an area wher M. pte’ J var. commutata commonly “The specimen SUNFLOWER FAMILY 581 collected by Douglas deposited at Kew insecrate d on a sheet with one of Coulter’s collection) is of the inflorescence only, but corresponds to this form rather than M. saratilis var. tenuifolia, This entity, with the related M. sara- tis var. prucieiothes: is quite distinct 1 in typical — from M. savatilis var. altissima, but intergrading forms e to be found at the southern extension of its ra 12. Malacothrix blairii (Munz & Jtn.) Munz. Blair’s Malacothrix. Fig. 6006. Stephanomeria blairi Munz & Jtn. Bull. Torrey wie — 301. 1924, Malacothrix blairi Munz, Man. S. Calif. 591, 601. Coarse straggling shrub 1-2 high, acu aa fleshy, thinly brownish-tomentose, gla- brate, scarred with persistent sabdigary subtending mall, brownish-tomentose bu Leaves ap- a ere ri u : or oblong-obovate, 6-13 cm. long, 3-6 cm. wide, rounded or obtuse, cuneate at base, irregularly sinuate- Sone ith 3-4 pairs of very blunt, unequal teeth, thin, light green, at first thinly tomen- i 0. cm. c i an gon. n Be cahiray Seciticas. of about 25 white, minutely Lapis git bristles. Known only sea the type soeelity, rocky canyon wall near Lemon , San Clemente Island, Los Angeles County, California. Late summer 6003. Malacothrix tprreyi 6005. Mal. cothrix saxatilis 6004. Malacothrix icana : he 582 COMPOSITAE 160. GLYPTOPLEURA D. C. Eaton, Bot. King Expl. 207. pl. 20, figs. 11-18. 1871 Low, tufted or matted, glabrous and somewhat fleshy, winter annuals with a deep tap- root. Leaves with a broad midrib, pinnatifid, the margins white-crustaceous and prickl toothed, often equaling or surpassing the flowers. Involucres cylindric, the phyllaries 7-12, by cross ridges; apex of achenes constricted into a short cupped beak. Pappus bristles i i i fer- A genus of 2 known species of the arid d t regi f th th t United States. Type species, Glypto- pleura marginata D. C. Eat Ligules little exserted; margins of the calyculate bractlets lacerate-toothed at the apex, lacerate-fringed belo . Gm een inata. Ligules much surpassing the phyllaries; margins of calyculate bractlets lacerate-fringed at the dilated a entire or nearly so below. . G. setulos 1. Glyptopleura marginata D. C. Eaton. — seed. Fig. 6007. Glyptopl ginata D. C. Eaton, Bot. King Expl. 207. pi. 20, figs. 11-18. 1871 Glabrous, fleshy, densely flowered annual, much br rae ad and forming er ce ss 5-15 cm. in diameter. Leaves 14 cm. long, pinnatifid, oblanceolate to obovate, narrowed to broad flat- tened see the lobes pee shed white-margined and pectinate ts toothed ; favolers iekalhy =. dric, on short peduncles and usually not surpassing t pad he ves; inner phyilla aries linear and a nate, 9-12 ong, roa “te eekvaicnts rgined; calyculate bractlets loosely ste rb spatu ulate and pectina e at apex, the apex with broad petite bees - gee er — t deeply lacerate- fringed; ligules aay little exserted, white turning pin achen mm. ong ; pappus very soft and fine, longer than the achene, the inne F beistieg. Scgtly ‘artited z the bas Dry sandy washes rd east of the Sierra copa to oo Mojave feast California, and east through Nevada. Type tocality' Sandy —— plain in Truckee Pass of the Virginia Moun ntains, in a cafion of the Trinity Mountains, and in Uni le Valley, Nevada.” April-— & i 2. Glyptopleura setuldsa A. Gray. Large-flowered Carved-seed or Keyesia. hep ej Glyptopleura setulosa A. Gra r. Acad. 9: Glyptopleura marginata var. pelea Baa Man. FI. ej eas 1008. 1925. Low, depressed, densely flowered annual closely resembling the ape ing species in general habit. ty hae es "Tike thos e of ‘the prece edink speci, the white-crustaceous margin usually narrower ; ai Sopabeee =o al ate, scarious-margined; calyculate hsgaige wit the white margins at x pectinately discested into slender teeth as long as the widt e green, Stine, a tip; ligules ay at ted, 1 mm. long, twice as long as t aos invouere, cream-colored or yellowish, aging pink; achenes and pappus similar to prec senditie spec andy oa flats, Lower Sonoran poe western and northern 2s Desert in southern Kern County and in Inyo County, California, eastward to northwestern Arizona and southwestern Utah. Type locality: near St. George, Utah. April-J 161. CALYCOSERIS A. Gray, Smiths. Contr. 5°: 104. pl. 14. 1853. Annuals branching from the base, glabrous and glaucescent below, glandular with tack- shaped glands above. Leaves pinnately parted into linear divisions. Ligules showy, in rather ee heads terminating the branches. Phyllaries linear, scarious-margined, much narrowed above into a short beak tipped by a shallow denticulate crown. Pappus-bristles white and copious, all united at base and falling away in a ring. [Name from Greek mean- ing cup, alluding to the shallow cup at the summit of the achene, and serts, chicory. A genus of 2 species, inhabiting southwestern United States and adjacent Mexico. Type species, Calycoser is wright A. Gray. Lanes 5 “en, purplish-veined; ribs of achenes more or less tuberculate; tack-shaped glands pale bags ish pol or agi ceaa mene ; ribs of achenes smooth; tack-shaped glands blackish-purple-tipped. 2. C. parryt. 1. Calycoseris wrightii A. Gray. White Tack-stem. Fig. 6009. Calycoseris wrightit A. Gray, Smiths. Contr. oy a. pl. 14. 1853. Calycoseris wrightti var. californica Brandg. 155;..19 Stems several from the base or cay nears erect or ascending, simple or usually branching, SUNFLOWER FAMILY 583 1-5 dm. high, pat green and glaucous or sometimes rose-colored below, glandular above with light ethos hier -shaped glands. Leaves basal and cauline, the lower 6-15 cm. long, pinnately divided into remote, linear or almost filiform and divergent segments, the uppermost reduced and bract- like; te campanulate, 10-12 mm. hi a phyllaries 12-18, ascending, linear-lanceolate, i i wish, tack-shaped minate tips more or less recurved; ligules 2-3 cm. long, white with rose-colored veins on the back extending down the tube from the rather prominent, attenuate teeth; achene shoe rk brown, tuber- culate on the eg 6 mm. long oe the beak; pappus-bristles mm. “a Desert hillsides, Lower Sonoran Zone; Shoshone and Death Valley regio nd Inyo County south along A western dues of the Colorado Desert of prac at California east to southern Utah rong Texas, and in northern Sonora. Type locality: ‘‘Stony hills around El Paso,’’ Texas. March—-May 2. Calycoseris parryi A. Gray. Yellow Tack-stem. Fig. 6010. Cal is p i A. Gray in Torr. Bot. Mex. Bound. 106. 1859. Stems several from the base, erect or ascending, simple or branching above, 6-30 cm. high, glandular above with tack-shaped, blackish-purple glands. Basal leaves more or less sthiicone ivi i nar line and fro e nar ear glands; phyllaries 10-13 mm. long, linear, rather ae attenuate at apex, broadly s ait margined, greenish dorsally or more or less purplish at the tip; calyculate bractlets few, 2-4 m long, scarious ; ligules showy, 1.5-2 cm. long, yellow; rachis light brown or gray with smadoth ribs. Usually in sandy soils, Upper and (mainly) Lower Sonoran Zones; desert regions from vatgh County, Cali- fornia, and adjacent Nevada, south through the Mojave and Colorado Deserts to northern Lower California and east to southern Utah and Fein Seon locality: Reported as “Mountains east of Monterey, California’; but probably not from that locality. Collected by Parry. March—June 584 COMPOSITAE 162. ANISOCOMA Torr. & Gray, Bost. Journ. Nat. Hist. 5: 111. 1845. Annual herbs with basal leaves toothed or pinnately parted and several monocephalous scapes. Involucre cylindric, the phyllaries with broad scarious margins, the outer short, broad and obtuse, the inner linear and acutish. Receptacle flat, the chaffy bracts linear- filiform and scarious. Ligules pale yellow. Achenes terete, linear-turbinate, 10—-15-nerved, the truncate apex with a narrow cup-like rim within whi ch is inserted 10-12 long, white, plumose pappus-bristles. [Name Greek, meaning unequal and a tuft of hair, in reference to the pappus A monotypic genus of the desert regions of southwestern United States. ie ES: acaulis Torr. & Gray. Anisocoma or Scale-bud. Fig. 6011. Torr. & Gray, Bost. Journ. — Hist. 5< 114, #. on figs. 7-11. 1845. esis hae runcinatus Kell. Peer: Calif. Acad. 3: 21. fig. winter annual with a rather a seated taproo ee in a rather dense basal rosette, pinnatifid, 2-7 cm. long, 3-10 mm. wide, the later Shee poeacadie and woolly-tomentose beneath, plants otherwise Rianeoe us ; scapes usually several to many, 5-25 cm. high, glabrous and bract tless : Se € Lan3.c mag, the phyllaries often margined with reddish brown, the outer short and peas the EE aetinont elongated and oblong-linear ; ligules conspicuous aed ale yellow Sandy desert washes, Sonoran Zones; Tehachapi Mountains ene east of the Sierra ati in ide County and ara Nevada south in the desert regions to no n Lower California and northwestern Arizona. Type locality: not indicated. April—July. 163. ATRICHOSERIS A. Gray, Syn. Fl. N. Amer. 1°: 410. 1884. Annual herb with a branched stem, glabrous throughout. Leaves in a basal tuft, cuneate or obovate, spinulose-denticulate. Stem-leaves reduced to small foliaceous bracts. Stems solitary, slender-branched above. Heads in a cymose panicle on slender peduncles. Phyl- laries scarious, lanceolate, about 15, with a few small ones at base. Receptacle scrobiculate. Ligules white or pinkish. Achenes oblong, corky-ribbed, without pappus. [Name Greek, meaning without hair, and seris, a name applied to chicory. ] A monotypic genus. 1. Atrichoseris platyphylla A. ay. Tobacco-weed or Parachute Plant. Fig. 6012. tncotheks: We — Proc. Amer. Acad. - 214. 1874. ray, Syn. Fl. N. Amer. 17: 410. 1884. sail a pasteee: and somewhat glaucous, the stems solitary, erect, cymosely branched above , 3-7 dm. high. Leaves basal, wide-spreading and usually flat on the ground, ob- long-obovate or oblong- set be olate, 3-10 cm. long, 2-5 cm. wide, narrowed at base A a usually u a about 6 mm. long, ovate-lanceolate, with broad scarious margins; corollas white or pinkish, the — te 10-15 mm. long, prominently toothed at apex; achenes white or stramineous, 4 mm. sy my oe ashes, Lower Sonoran Zone; Mojave and Colorado Deserts of California. Inyo County to Im Sucre oer y, California. and crear eee evee, east to southwestern Utah and western Arizona. Type locality: near Fort Mohave, Arizona. Collected by Coope Feb. —May. : 164. TRAGOPOGON [Tourn.] L. Sp. Pl. 789. 1753. labrous biennial or “shaded perennial herbs with ions taproots. Leaves entire, grass- stems like, with conspicuous nerves, clasping at base. Peduncles terminating the long, stout, often thickened and fistulous ; ptoliacce | in 1 series. Heads yellow- or purple-flow- early morning, usually closed by n Receptacles without bracts Achenes fusiform, long-beaked or the outer beakless, 5—10-rib . Pappus a St row of long, plumo: se bristles cuneate at the base. { Name Greek, meaning goat’s- enus of about 35 species, all natives of the Old World. Type species, Tragopogon packet Er hie scarcely inflated even in trait; outer Higules equaling the phyllaries. 1. T. pratensis. phy 11 Ligules uae to deep Siclet: urple; atly 3 8-9. 2. T. por is age Ligules pale yellow; phyllaries wea 13; 3. T. dub i = eeeere Peaioeie te Meadow Salsify. Fig. 6013. Tragop pratensis L. Sp. Pl. 789. 17 Stems from a hpi usually ani rather slender, 4-10 dm. high. Leaves keeled from the more or less clasping base to a long-acuminate, usually recurved tip, the margins concave and SUNFLOWER FAMILY 585 — below the heads even in fruit; phyllaries 8 or 9, rarely 13, broadly va teh gat acuminate, shor than or rarely slightly exc ceeding the chrome- -yellow ligules ; marginal achenes striate, ends or r pacagenri usually dark brown, the inner paler, 20-25 mm. long, Seely tapering to the beak; Papp Fields and w aii » Sparingly adven in western Washington and Oregon to Shasta County, Cali- fornia, and more hy deel eo east of 4 Coctade seh Pll hag seberialis in Uni ion an og Wallowa y sense vey Oregon; also northern Arizona. Widely distributed in Canada and eastern United States. Native of Europe. June—Sept. 2. Tragopogon porrifolius L. Salsify or Oyster Plant. Fig. 6014. pape evince Sp, PI. 789. ‘1753. mple from a — aproot, 5-12 dm. high. Leaves tapering uniformly to the apex, dilated and ‘chaping at base; peduncles much inflated and hollow from 2.5~4 cm. below the heads S" phyllaries 8-9, rarely 12, dir nia te, pyar usually much longer than the outer ligules ; fenles Ail chee about 3-4 m m. long, the outer ones covered with scale-like tubercles, espe- cially on the ribs, the beak ene longer seat the body and abruptly narrowed; plumose branche Ss of the —— -bristles ee erage t- Naturalized as a weed in the Pacific Sas a eS less frequently, eastward across the continent. Native of the Old World. en Nov 6011. Anisocoma acaulis ae Tragopogon pr: 6012. Atrichoseris platyphylla . Tragopogon pera 586 COMPOSITAE a. Ee a dubius Scop. Yellow Salsify. Fig. 6015. Jregepogen dubius Scop. FI. Carn. ed..2. 2: 95.. 1772. Jacq. FI. hae Belect. Icones 1: 19. pl. 29. 1773. “Stet ee om a taproot, 4-8 dm. high, rather bushy, the branches from near the base. Leaves linear- scaliontate: 12-15 cm. long, tapering bition from base to apex, racner floccose when young, oming glabrous and glaucous; heads many- “fbw ered, the peduncle much inflated below the involucre ; phyllaries about 13, about 33 mm. gre ng, aes ceolate to linear- a 5 parents ligules pale yellow, shorter than the pha tlaties achenes slender, 25-36 mm. long incl udin the bea k, the dy gra eral narrowed to the beak, the outer achenes pale eae the omer i and straw- colored; pappus whitish. In fields snd waste places in southeastern Washington and adjacent Idaho and occurring sporadically — ward; also eastward to Kviiets and Texas. It has been collected in Siskiyou, Santa Clara, and San Bernardin Counties, California, Native of Europe. May—Sept. The recent work on natural hybridization and amphiploidy in Tragopogon (Amer. Journ. Bot. 37: 487-499. 1950.) Marion Ownbey demonstrates the eee Pf two new amphiploids which oy cya ber — hybridization in southeastern Washington and agg col Idaho, where all three previously kno The amphiploids are Tragopo “ : mirus Ownbey, Am ourn. Bot, mils 497. 1950 (T. Pe 3th x P crriine and 7. miscellus Ownbey, op. . 498, escced from T. pS te xX pratenst. 165. SCORZONERA L. Sp. Pl. 790. 1753. Perennial, rarely annual, herbs. Leaves alternate, usually entire, often grass-like, some- times pinnately obed or dissected. Flowering heads on long peduncles, yellow, rose, or lilac. nvo. i i in tH 5 - o n 72 o <2 os ot oy ' Ww fo) ad rt) et & (aa = o ° c o Les 3 | <> me = ro) = oO wo 3 or a ss n rus ° bas | a o + co = ye ro) =| ee a oO rar o co pale a) branches slender. Achenes linear, subterete, many-nerved. Pappus-bristles in more than series, unequal, serrulate or more or less plumose. [From the old French, meaning serpent. A genus of about 120 species, natives of the Old World. Type species, Scorzonera humilis L. ? aoe hispanica L. Black Salsify. Fig. 6016. Scorzonera hispanica L. Sp. Pl. ed. 2.1112. 1763. Scorzonera glatifolia Willd. = Pl, 3: 1499. 1804. Branching perennial 6-10 dm. high from a fleshy, black- ya tanrtet, the herbage woolly- idee cae to nearly glabrous. Le eave s oblong to lanceolate or linear, the margin entire, undulate, tapering below to a Rosy loris ae the stem-leaves few, much re eds ced above; peers poser i the ends of the branches, the peduncles 15-30 cm. long; involucre urceolate in flow 2. long, the ligules yellow; phyllaries few, rs geod d-seriate, hie -acum “id somewhat olly ee em ag e the inner; slender, many rved, of the ou Y fener’. “ge tied eebertubste: aaaceal bristles | ae cuca length, 1 cm. long or Seng ihidiase oe soft hai A native of southern and central se which has become established on roadsides and waste places in Napa and Mendocino Counties, California. June-July. Viper’s Grass. 166. HYPOCHOERIS L. Sp. Pl. 810. 1753. aceo cts! to linear, ge constricted above or the outer truncate. koure? * of a peek i me 1 of "Theophr astus for mber of this t About 50 species, natives sigs ok the Ee tote R Region, Asia, and South America. Type species, Hypochoeris glabra L. Annual; heads 4-6 mm. broad; t , not beaked LH: glabra. Perennial; heads 20-40 mm. broad; achenes all beaked. 2. H. radicata. 1. Hypochoeris glabra L. Smooth Cat’s-ear. Fig. 6017. Hypochoeris glabra L. Sp. Pl. 811. 1753. l, mple or often bocyuntiiat sely Srasiched: 1-4 dm. high, giabed ous, bracteate especia ally at the | base oF the branches. Leaves spreading on the ground, denticulate to pinnatifid, “oblanceolate to oblong-oblanceolate, 2. s etimes cili ar panulat, 10-16 mm. long; ligules scarcely exceeding the phyllaries; achenes dark brown, the ermost beakless, the wars with slender beaks as lo ong as the body; pappus 1 cm. long. a Cale on roadsides, fields, and waste places, Son d Transition Zones; Washington, Oregon, and California, also eastern United States and Canada. Waties. = the Old World. March- Sept. 2. Hypochoeris radicata L. Hairy or Long-rooted Cat’s-ear. Fig. 6018. Hypochoerts radicata L. Sp. Pl. 811. 1753. Perennial ; stems several, slender, 3-8 dm. high, branched or rarely simple, bracteate. Leaves SUNFLOWER FAMILY 587 basal and spreading on the ground, oblanceolate to obovate, pinnatifid to dentate, 5-30 cm. long, hirsute; involucre oblong-cylindric, 2-2.5 cm. long; ligules usually well exceeding the avtiotas achenes bro own, all with slender beaks as long as "the get sein plumos Roadsides, waste places, and pasture land, Sono tion Zones; ‘ideal weed throughout the Pacific States and across the Continent. Native of the. Old. World. ke il—-Nov. 167. HEDYPNOIS [Tourn.] Mill. Gard. Dict. abr. ed. 4. 1754. ual herbs with branched stems and yellow flowers. Phyllaries in 1 series, narrow short scales or sometimes absent, the inner of bristles dilated toward the base. [ Name given by fits toa kind of endive. cies, natives of the Mediterranean Region and the Canary Islands. Type species, Hedypnois aoe; Mill. gh ihc hedypnois i, 1753). WIG, Vs 6015. Tragopogon dubius 6017. Hypochoeris esa 6016. Scorzonera hispanica 6018. Hypochoeris radi 588 COMPOSITAE 1. Hedypnois crética (L.) Willd. Crete Hedypnois. Fig. 6019. ad goes nies. L. Sp. Pl. ed. 2. 1139. 1763. eticus All. Fl. Ped. 1: 226. 1785. a see cretica Willd. Sp. Pl. 3: 1617. 1804. Stems simple or usually branched from the base, 1-3 dm. long, hispidulous, the hairs simple or ‘ciinaaby rp the tip. Basal leaves oblanceolate, tapering to a winged petiole, entire or sparsely m bed, 2-10 cm. | idulous ; denticulate or someti ie ong, mm. wide, Seerdal hispi id peduncles erate solitary, or on more branched stems; involucre 8-10 m igh, enlarged in fruit, his- ulous peat ne ete on the margins; achenes appressed- scabrous on the ribs; pappus- Locally natural ayo dry interior valleys of the Inner pe igo and the Sierra Nevada foothills west to = coast ied south to San Dieys, California. Native of Eurasia and originally described from Crete. March—May 168. PICRIS L. Sp. Pl. 792. 1753. large, a or paniculate heads of rail flowe volucre c ate or cupulate, its inner phyllaries in 1 series, nearly equal and erect, its exterior ones in 2-3 series, small r large and spreading. Receptacle flat, short-fimbrillate. Rays truncate and 5-toot t apex. Anthers sagittate at the base. Style-branches slender. Achenes linear or oblong, slightly curved, terete or angled, 5—-10-ribbed and transversely alpen narrowed at the ase and apex, or beaked in some species. [Name Greek, meaning About 35 species, natives of the Old World, or one species possibly sl i in Alaska. Type species, Picris pairs Re Ary i 1. Picris echioides L. Bristly Ox-tongue. Fig. 6020. Picris renee wp, PE 792: 1753; au ial with branched hispid stems 5-12 dm. high. Leaves beset with coarse, barbed, usually pustulate- based cass ae basal and lower leaves spatulate or oblong, obtuse, ro mage dentate, 5-15 cm. lon: i th late, the uppermost ese e and entire; heads many, usually rather crowded, sheet is tees d, about 1. oad ; outer obyilaries of ein in oo 4 or Ba by sowie subcordate, hispid-ciliate ; inner pistlevies tanckata thin and tipped swith, a prickly A native of Europe, naturalized in waste places, sieaahe eae roan hag distributed but most common in central and pethers: California; also Michigan east to the Atlantic seaboard. May—Nov 169. LEONTODON L. Sp. Pl. 798. 1753. Annual or eee herbs with heads borne on simple or branched, naked or scaly- bracted scapes. Leaves in a basal rosette, entire, toothed, or pinnatifid. Heads many- ime pappus of the marginal flowers often of scarious paleae only. [From the Greek words meaning lion and tooth, in allusion to the toothed leaves. ] enus of a 45 species, natives of Europe, western and I Asia, and northern Africa. Type species, teteee hispidus 1. Leontodon léysseri wet) G. = Hairy Hawkbit. Fig. 6021. Crepis nudicaulis L. Sp. Pl. 805. ee (N Leontodon hirtum of authors, not t. ed. 10.1194. 17 59. oseris taraxacoides Vill. Prosp PI aa ee Thrincia hispida Roth, C B 822. Leontodon nudicaulis erat. Ann. Sci. Nat. 22: 109. 1831. Loontetes eatbie Ball, Journ. — — 16: +. 1878. Leon Beck, Fl d-Oesterr. 22: 1312. 1893. yon nudicaulis subsp. nits Schinz & Theil, Bull. — =e II. 7: 389. 1907. Leontodon nudi, Thell. | Scapose perennial with fibrous roots and short ars Leaves many in a basal rosette, 5-15 cm long, narrowly oblanceolate, tapering to a etiolitcras base, midvein prominent, sometimes nearly entire, more often shallo oothed to runcinate-pinnatifid, with hirsute sprea t ence ; heads solitary, the s scapes several to many, 10-35 cm. jong, n ve, spreading-hirsute toward the base; involucre 6-10 mm. long, the inner phyllaries subequal, lanceolate, gla = ta ad —— usually medially darkened, the ites reduced to small calyculate eee achenes long, fusiform, more or less sho srtly beaked above, striate, the striae muricate, pappus of the SUNFLOWER FAMILY 589 marginal flowers usually reduced to short eae scales, that of the inner of plumose bristles, exceeding the achene in length and flattened below to a scarious base. A European soe of lawns, waste places, and roads ifs from pe Columbia west of the Cascade Moun- tains south to central California; eae intron I in eastern United Stat Because of the oe created by efforts to assign a precise por he of the basonym Crepis nudi- caulis, authors using the ne Prahor me available name here followed. Leontodon sueuaahine Le Spe, 798. “F753: Differs from L. leysseri in having the a ce of sev- eral heads, the peduncles sealy-bracted and_enlarged below the head, and the onppas ner and outer flowers a single row of plumose bristles. A European weed rather commonly distributed in the case but otha in pos a og ‘eregy be from a single collection: % in Bellingham, Washington (Blake, Leaflets West, Bot. 7: 285-86. 1955) so from 170. SONCHUS [Tourn.] L. Sp. Pl. 793. 1753. Annual, biennial or ere rather fleshy coarse herbs with a mostly auricu- late-clasping, enti natifid, prickly-margined leaves. Inflorescence corymbose, paniculate, or ser af ie heads yellow-flowered. Involucre in sea coming thick- ened and indurated at base in age, the phyllaries somewhat peri cis = thin tips reflexed in age, subtended by calyculate bractlets. Receptacle flattened, . Achenes obcom- pressed, ribbed or striate, often rugose, truncate at apex. Pappus much surpassing the achene in length, of copious, soft, capillary bristles ptecnac at rie. hase and falling away ina ring. |The Greek name for the sow-thistle. ] An Old World genus of about 45 species. Type species, Sonchus oleraceus L. Hedypnois cretica 6021. Leontodon leysseri a4 Picris echioides arvensis 590 COMPOSITAE Perennial with deep vertical root and horizontal creeping root-branches; flowering-heads 4-5 oe broad. arvensis. Ber Ae or rage al; ee ng-heads less than 4-5 cm. broad. Au s of the leaves acute; achenes striate, agantagai! wrinkled. ome blades serra ead r with lobes 0.5—3 cm. wide; achenes about as broad as thick. 2. S. oler Leaf- yer s pinnately parted with narrow me mostly less than 0.5 cm. wide; achenes ya Bees. slightly = k ag of the — rounded; achenes usually conspicuously 3-nerved, smooth or ness A, pt ae 4 Sper 1. Sonchus arvénsis L. Perennial Sow-thistle. Fig. 6022. Sonchus arvensis L. Sp. Pl. 793. 1753. Perennial with c reeping rootstocks ; stems nth dm. high, glabrous below, Seal tre mabe bristly above, pi ae the pedun cles. Lea s 3-10 cm. lon ng, runcinate — ee d or sometim merely dentic ulate, the margins a, -denticuate, the lower narrowed t Sasa! petioles the upper sensiie, — ulate- aos at base and rounded; inflor ce seeiitelah es mbose or even subumbelliferous ; involucres 14-20 m me nine the chelated shins ate at apex, cinahieslae-liintly as are also the Stduiihens ligules scieht: orange- -yellow, the heads up to 3-5 cm. broad at ant si achenes cs 8-10 mn “ong ned, 2 2.5-3.5 mm. long, usually 5-ribbed on each face, yale tie rugose papp ng. ralized fro rope but not common in the Fecite ote Puget Sound ar d Whitman County, Wash- ing Mths Ppa Sing and west = the Cascade Mou n Willa eats Valles: occurring sporadically in Cali- forma whoa it has been collected at a few scattered pager throughout the state. April—July. us arvensis var. poms Guenth. Graeb. & Wim , Enum. Stirp. Phan. Siles. 127. 24. (Son chus A nm nosus Bieb. Taur. Cauc. 2: 238.. 1808.) Resemblin ng ome i arvensis var. arvensis in habit and variability of ave: “son lacking the ‘glandula ar bristles on involucre and peduncles of that taxon, tle a pbmrioe! are often m ale. uropean weed more common in north — = northeastern United States, and occ ng sporadi cally in the Pacific States as a weed but less common n S. arvensis var. pieeasis 2. Sonchus oleraceus L. Common Sow-thistle. Fig. 6023. Sonchus oleraceus L. Sp. Pl. 794. 1753. Plants usually stoutish, 5-10 dm. high, sparingly leafy, glabrous or with a = glandular hairs on the peduncles and involuc cres, oiten glaucescent. Leaves variable in shape, ovoid or narrower, si rari or runc a -pinnatifid, toothed but not prickly-margined, amplexicaul, ine hacen straight and the lobes acute; —s yellow; achenes about 2 mm. long, striate -nerved, Be ae trans- banal rugose- Seabee s on nerves and intervals, not strongly flattened on the mmon —~ pes roadside weed throughout the Pacific States and across the continent. Matte of Europe SpringsAnteane 3. Sonchus tenérrimus L. Slender Sow-thistle. Fig. 6024. Sonchus tenerrimus L. Sp. Pl. 794. 1753. Sonchus tenuifolius Nutt. Trans. Amer. Phil. Soc. II. 7: 438. 1841. Plants rather slender, leafy-stemmed to the inflorescence, glabrous, 3-8 dm. high. Leaves ob- long in stad eg deeply divided into several pairs of aie Spreading lobes, hee entire or denticu- ate, the au ricles narrow, lanceolate to ovate, sometim uate ; Pate it cea — rt corymb, mostly with few heads ; " phyitairies “sergbinngee age 81: fica site thic a ietow age; ligules yellow; achenes narrowly ovoid, 2. mm. long, = little eae longitudiealty striate and finely and distinctly rugose on both s striae ary intervals. Introduced from Europe, sandy and rocky soil; Aapecas oso the Channel _—, Santa Barbara County, south e western San Diego County, California, s outh to Lower California. March— Jun 4. Sonchus asper (L.) Hill. Prickly Sow-thistle. Fig. 6025. Sonchus oleraceus var. asper L. Sp. Pl. 794. 1753. Sonchus asper Hill, Sebastian Brit. 1: 47. 1769 Similar to Sonchus oleraceus but stems usually movies. ferdat ed angled, and more leafy. Leaves undivided, lobed or sometimes pinnatifid, the margin nulose-dentate or spinulose-den- ticulate, the low wer and basal obovate to spatulate and petioled, t the. upper oblong or lanceolate and clasping by an auriculate base, the auricles helicoid and appressed to the stem, the lobes ype ligules jonas achenes 2.5-3 mm. long, st ronely usually 3-ribbed on each face, not at all t versely rugose, the this, sine like a corte rrulate. A fairly common field and roadside weed in the ech States and also across the continent. Naturalized from Europe. June—Nov 171. LACTUCA [Tourn.] L. Sp. Pl. 795. 1753. Leafy-stemmed annual or perennial herbs, glabrous or nearly so. Leaves linear and entire to broader and pinnatifid. Heads 5-56-flowered ; ligules yellow or blue, in small or medium-sized, paniculate heads. Involucre cylindrical, the phyllaries usually imbricated in several series. Receptacle flat, naked. Ligules truncate, 5-toothed at the apex enes oval, oblong, or linear, flattened, ribbed on each face, abruptly or gradually beaked. Pappu ious, of soft capillary bristles. [The ancient Latin name, from lac, milk, referring to the milky juice.] SUNFLOWER FAMILY A genus of about 50 or more species, natives of the northern hemisphere. Type species, — sativa L. baat common garden lettuce is Seeulonkity’ collected in waste places but apparently does not become estab- lished 591 Aemeal: or biennial; ligttles not long-exserted, shades of yellow (bluish in L. biennis and forms of L. /udoviciana). 1-nerved on each side. Frviting involucres nS as m. high; pappus 5—7 mm. long. 1. L. canadensis. Rubehnetaat Subtaapeats3 re 22 arp aigre baie 7-12 mm. long. 2. ha pareminted Achenes with a long filiform beak as long as or longer than the achene. es black, conspicuously eae ne 3. L. virosa. chenes gray or brownish, no Inflorescence broadly paniculate rig peste ap branches; stem-leaves bine to elliptic in outline Infl picif P lat 1 tly linear- EN Rg — 5. L. saligna. Achenes with a short —- sates ageen than the achene or beakless Pappus tawny; ligules 15-35, white or bluish (in ours) 6. L. biennis. Repose demic gules -e yellow. 7. L. muralis. Perennial wit ; ligules long-exserted and showy, blue or bluish purple. 8. L. tartarica pulchella. 6023. Sonchus oleraceus 6025. us asper 6024. Sonchus tenerrimus 6026. Lactuca canadensis 592 COMPOSITAE 1. Lactuca canadénsis L. Trumpet Fireweed. Fig. 6026. Lactuca canadensis L. Sp. Pl. 796. 1753. Lactuca elongata Muhl. ex. Willd. Sp. Pl. 3: 1525. 1804. Lactuca pase Rydb. Bull. Torrey Club 38: 23. 1911. Stout, weedy, simple-stemmed annual or _ 3-25 dm. high, glabrous or sometimes beri coarse ain hairs. Leaves entire, toothed, or deeply pinnately lobed, sagittate and often nar- rowed at base, 10-35 cm. long; panicle elongate , spreading, the peduncles —— heads 13- "2. flowered with yellow ligules ; fruiting Pe i usua — 10-15 mm. high, t phyllaries i in 4 or series ; achenes black, obovate, flat, transversely rugose with 1 prominent peeoe on eac = the beak about one-half as long ‘to equal the body of the sat di ndnc white, 7-10 mm. lon Fields and_woodland, introduced from the eastern United States, occurring ieee in adinder | in Wal- lowa County, Oregon, and in Siskiyou, Shasta, Plumas, and Amador Counties, California; also Idaho. Type locality: Canada. July y_Sept. A taxon quite variable as to leaf-shape and many named forms have been described. Both the entire-leaved and pinnatifid-leaved forms are found in the Pacific States. a ett ludoviciana (Nutt.) DC. Western Lettuce. Fig. 6027. Sonchus ludovici Nutt. Gen. 2: 125. 1818. Lactuca ludoviciana DC. oer 7: 141. 1838. Lactuca campestris anata Pittonia 4: 37. 1899. Biennial, t ms leafy the inflorescence, pahiculately bce attr above, 6-20 dm. high. Leaves a clasping yp uaaddienaeulats sinuate-lobed or pinnatifid with spinulose eee — setos sores id on the midrib, otherwise gla a direcahant: heads numerous re | pen pa tet hele pedirictos bes, er fruiting involucre 15-22 mm. high, cyclindric or pote evi ndric, a These: s, the phyllaries in 4 or 5 series and peg Mes shorter and broader ; ligules yellow ; achenes oval to obov te "i ms ttled with gray or i k, transversely rugose, 1-ribbe on each fac e, beak about equaling the “at dilated below ; white. SENS introduced sa ae visas arene Klickitat County, Washington, Wallowa County, Oregon; Shasta semety ig be ia, and vic of San ardino, southern California. Type locality: ‘‘Fort Mandan on the 3. Lactuca virdsa L. Wild Lettuce. Fig. 6028. Lactuca virosa L. Sp. Pl. 795. 1753. Biennial, the stems stout, often 6-8 dm. high. Leaves ample, broadly obovate, 6-15 cm. long, strongly rae sharply dentate or sometimes. divided, glabrous, the midvein often prickly on the dorsal side ; Lees Borns often 30 cm. long and 15-20 cm. broa d; heads 6-12-flowered, yellow Grohe s 10-12 m. high; ac a hee i long, n acronis obovate, black, 4-5-ribbed P gored the thickened w sommes margin, cnuicasy spinulose a apex ; beak slender, ‘about equaling the of the pe ee ; pappus whi Seams lly on satan or yerey: shaded slopes; a European weed rather sparingly introduced in the San Francisco Bay region, " Californ . June 4. Lactuca serridla L. Prickly Lettuce. Fig. 6029. Lactuca serriola L. Cent. Pl. 2: 29. 1756; Amoen. Acad. 4: 328. 1759. Lactuca scariola L. Sp. Pl. ed. 2.1119. 1763. Biennial, the stems leafy, ey soem branched above, 5-20 dm. high, glabrous throughout o prickly-hispid below, pale green or s colored. Leaves oblong or Ree cee Scions ae ing, lobed or pinnatifid, the tow 2 : i the upper much re : a5 ulose-hispid on t ag “ea midrib and margins, otherwise glabrous or more or less hispid; heads s in : usually large, seh panicle, 6-12- flow ate Scales ier sonal subcylindric or i Te conical, 12-16 mm. high, the phyllaries ee oiaste 4 i pigraegae in about 4 series, linear-lanceolate ; ieee obovate- oblong, about equaled by the slender beak; pappus white Be ids and waste places; a common weed in the Pacific States, ‘eo generously distributed in the United States. Native of Europe. Pe ns Ph ft. The plant with the leaves simple instead of lobed or divided has been recognized as a form (L. serriola var. integrata Gren. & Godr. Fl. France 2: 320. 1850). 5. Lactuca saligna L. Willow Lettuce. Fig. 6030. Lactuca saligna L. Sp. Pl. 796. 1753. Annual, glabrous throughout, the stems usually several from a rather stout taproot, dec . high. v e , 3-7 Leaves pale green, narrowly linear-lanceolate, attenuate at a sagit- tate-clasping at base, en sometimes thed or f sn to : 8-10-flowered, yellow, in rrow spicate panicles with short pe cedis branches volucres narrowly cylindric, 1 se long ; achenes light brown, 3 mm. long, oblo Sie Caned os rather abruptly attenuate at a ribs 5-7 on each, a ‘spiculate near the apex, the beak slender, 5-6 mm. long; pappus- eae et — as long s the beak. Ae Aue wee ig has become well established in many places throughout California and has also been found in Dour and Marion Counties, Oregon; also eastward to the Atlantic seaboard. Type locality: Europe. 6. Lactuca biénnis (Moench) Fernald. Tall Blue Lettuce. Fig. 6031. Sonchus biennis Moench, Meth. 545. 1794. Sonchus racemosus Lam. Encycl. 3: 400. 1789. Not Lact osa Willd. 1804. SUNFLOWER FAMILY 593 Sonchus leucophaeus Willd. Sp. ™ 3:3:1520. Lactuca — a Hit yet in Britt. & ile 7; Fl. 3: 276. 1898, as to plant described; not as to basic type Sonch Peeteanes Lactuca biennis “iakrg ‘Bocioce 42: 300. 1940. Biennial, glabrous throughout, the stems simple, stout, erect, 1 -3.5 m. high, leafy. Leaves pin- nati id o or ru poor irregularly i sharply dentate, the upper auric culate-clasp ping at base, 15-30 hea d 1 d cm. long; heads in an elongated, many-flowered panicle, ligules little-exserted, cream- colored to bhaish Eblgercs 9-14 mm. high, siege constricted above, the phyllaries pe pena at tip; achenes 4 mm. long, mm. broad, narrowly winged on the margins, 3-ribbed on each side, gpa ome with different ah of brown, narrowed at apex to a very short beak; pappus tawny, o times tinged with purple. Wet grassy aa in rich soils, Transition and Canadian Zones; nee British tapers and Olympic Mou tains, Wace a, to Humboldt Ff eatpeget California, and east a s Washing and Orego: on to the caiters United States and Can da. Type locality: not indicated. Fude-Ades 6027. Lactuca ludoviciana 6029. Lactuca serriola 6028. Lactuca virosa 6030. Lactuca saligna 594 COMPOSITAE 7. Lactuca muralis (L.) Fresen. Wall Lettuce. Fig. 6032. Prenanthes muralis L. Sp. Pl. 797. Cicerbita muralis Wallr. Sched. Crit. 436. 22, Mycelis muralis Reichb. Fl. Germ. Excurs. a 1830-32, Lactuca muralis Fresen. Taschenb. 484. 18 Slender yan annual or kere stems simple, 4-9 dm. high. Lower stem-leaves thin and often vagelich: lyrate- or runcinate-pinnatifid, narrowed below and auriculate-clasping, the ter- minal segment broad and angularly lobed, up to 18 cm. long, the upper stem-leaves few, much re- duced; sessile and as eee Byrn ited: inflorescence a divaricately branched panicle; involucres narrowly cylindric, 9-11 high, 5-flowered, the ligules yellow; phyllaries in 2 series, the inner long, the outer fis crop it Me about 4 mm. long, dark brownish or reddish, flattened, several-nerved on eac ary the — k pale, Pack shorter than the achene. AE n species woods, that has become established on Vancouver Island, British Columbia, and in laleat a Challe yer Wakianiea: yt New York and Quebec. June—Aug. 8. Lactuca tartarica subsp. pulchélla (Pursh) Stebbins. Blue Lettuce. Fig. 6033. Sonchus pulchellus vai: FI. ony pee 502. 1814. Lactuca pulchella DC. Prod. 7: Lactuca tartarica subsp. sides Shite, Madrojfio 5: 123. 1939. Plants with a deep perennial rootstock, glabrous throughout and somewhat glaucous, the stems leafy up to the inflorescence, 5-12 dm. high. Leaves variable, linear to lanceolate or oblong enti, dentate, lobed, or pin a the lower sometimes petioled d, the others sessile or som hat clasping, 5-20 cm. long; heads in an elongated eens often leafy below ; invo c; Sy gh; iy lat ries cael bashed cated” the outer succ apts shorter, ovate-lanceolate, often purplish ligules blue or violet, well exserted ; en elied ehince ladeesi sate flat, short- beaked ; pap- < c + oO Q = wn res a] Psee Moi und, Arid Transition Zone; British Columbia southward east of the Cascade Motateins through eastern Wathinetie and Oregon to boc Oe | hag the hg —_ as a weed in the Bw n Josie Valley; east to western Ontario and Kansas. Type loca roe t anks of the Miss ssouri.”’ June 172. PRENANTHES [Vaill.] L. Sp. Pl. 797. 1753. Leafy-stemmed, perennial herbs. Leaves alternate, petioled below and sessile or auricu- late-clasping above. Inflorescence thyrsoid or ee culate, the flowers drooping. Involucres cylindric and usually narrow, 7~-15-flowered, the phyllaries in 1 o seri Receptacle naked. Ligules white shades of pink or Pirie or yellowi h. Style-branches slender. Achenes beg = kag ni or terete, ia e; Bye us COpLONE, white or reddish brown, A genus of 27 species or less, natives Ea sea Europe and ga North America, and the Mediterranean Basin. Type species, Prenanthes purpure 1. Prenanthes alata (Hook.) D. Dietr. Western Rattlesnake-root. Fig. 6034. Sonchus hastatus Less. seaenee 65 99. 1831 Nabalus alatus Hook. FI. B nee . 294. os 102. 1833. Prenanthes alata D. Dietr. Pat Pi. evens mines Rosas aM. E, wee , Bu : fe all Biol. Ser. No. 15: 47. 1910. Not Thunb. 1784. git Hul én, Fi. Aleut. Isl. 335. 1947. Perennial herbs si ori I Sogpoe ero — 8 or somewhat branched above, 3-6 dm high, glabrous or nearly so below Toes Lower and middle leaves 5-12 cm. long, paler below than above, deltoid to rary apraton ietenanke ee ee ae arply aaa irregularly ent t tiolar base, the upper scarcely narrowed bel nd shorter than the lower ; heads 10-15-flowered, the ligules purplish, loosely and so hat paniculately cory ; peduncles tomentose ; lucre narrowly campanulate, about 12 mm. high, the phyllaries linear, becoming glabrate, the calyculate bractlets very small ; oepen es about 5 mm. long, finely striate po pts the ribs ; pappus- bristles capillary, ra gins brow ish. ich soil in ele Humid Transition — dian “Zones Alaska eure along the Pacific Pity oy ‘Daten: and Hood River Counties, Oreg ae ocality: Fort Vancouver, Washington. Col- lected by Scouler, June— 173. HIERACIUM [Tourn.] L. Sp. Pl. 799. 1753. Perennial herbs, the stems scapiform or leafy, arising from a branched caudex, stout rhizome, or from slender stolons. Leaves entire or toothed in most of ours, more abundant b ew- to ma oO yellow, white, or orange-red ligules. In calcite cyli indri or campanulate; phyllaries in 2 or 3 series, often obscurely imbricate. Receptacle flat, mete naked. Achenes truncate, striate; pappus a single row of white or tawny, capillary bristles. [From the Greek, mean- ing hawk. 4 SUNFLOWER FAMILY 595 A variable genus, in which nearly 1000 species have been described, native of both hemispheres Ly <, My A j . Type species, 1 native species, perhaps Ly a i degree in H. horridum, ape ve field and experimental studies should be made to at ive at an understa nding of true r etatic Sots nahi ips. Then so. * e names here listed as synonyms may be found to represent —- bislogical entities te least at the Subspecific level. Probably in the albertinum- tleri-cynog lo mplex, a situation exists compar. cin to that which has been demonstrated to occur in the polyploid apomictic pond 9 ne yi of the oe us 5 uro re not stoloniferous; native s ith long eaves Goltwad ng oety long bates tiated ‘iarite of the ae River Gorge. cies sal st sae al — rather small os pico sainraeene those . rs many and mostly larger. Ste etose lon “Lea longiberbe. T.eaves essentially glabrous; plants widely distributed in Canada and northern United Asap nadense. Stems without long setose hairs; leaves four to twelve times as long as wide. Basal po subbasal leaves the ri arkedly reduced upward Stellate cueastasne: lacki wers white, 19-30 p per head; plants 3-12 dm. high; widely seen in Fagen t Pomoatg States. is pe mbellatum. rger, persistent, those of midstem few or none, progressively and sometimes Flowers yellow, 5—15 per head; plants 1-3 dm. high; restricted to the ‘Siskigon region. . H. bolanderi. sets pubescence present (sometimes only on the inflorescence), sparse or dense nts canescent throughout with a a stellate osaaan pelo elie with long setose hairs on the lower leaves.) ae Ai. greene. Stell restricted to leaves or inflorescence, obsc MNearl¥ all leaves art but conspicuously sinuate-dentate; plants of aan fgg ect and southern Califor 3. Leaves all entire or a few obscurely denticulate; plants not of west-central and stick Cali- fornia. Basal i eo glabrous or if at all pubescent lacking setose hairs; err. Baggs stemmed plants, mostly of moist mountainous si tions, res = & cile. Basal and pent eaves when present dens or apeesely covered with ‘i setose hairs; plants principally of yp Bh open or set d area: Heads (5)10-12-flowered, the involucres narrow; plants crinate, Mendes, Te 1-3 — high. 0 Heads ae 50-flowered, the involucres broad; plants if crinate rather stout, %. $- 7 dm. Hathess sparsely to moderately long-setose above (occasional on involucres) and sometimes glaucous; plants of Cascade, Tiikiners, ry Sigre Bl Megs ranges, Herbage egg long-setose shove d_ below, — 3, glawcous ds tea ae of the Cascade, Siskiyou, sis Gos rthe a Sierra es. Involucres and stems pega sparsely long- a meee ney spl . H. cynoglossoides. Involucres and sometimes the stems crinate, sss dai ‘if at ai glandular. Be d 3. H. albertinum. tol ; introduced spec Inflorescence o él 5—10 heads, orange- es orange-red-flowered. Heads solitary, yellow-flowered. 13. H. aurantiacum. 14. H, pilosella. 6031. Lactuca biennis 6032. Lactuca muralis 596 COMPOSITAE 1. Hieracium scotleri Hook. Scouler’s Hawkweed. Fig. 6035. Hieracium scouleri Hook. Fl. Bor. Amer. 1: 298. 1833. Hieractum ~~ Gr aan es 3:10). 1895. Hieracium cinereum Howell, Fl. N.W. Amer. 396. 1901. a Tausch. 1819. ro tea cineritium Nel s. & ser Bot. Gaz. 61: 47. 1916. dog Bull. ~ Bot. Fr. 65: * 1918. — paddoense Pips a Op. cit. Hieracium washingtonense Gandoger oe Hier ais pepo Zahn, Piieneak nl 1127. 1922. ?Hieracium parryi Zahn, op. cit. 1128. Hieracium ae as Zahn, op. cit. 1129. Hieracium hemipoliodes Fabri, op. cit. 1130. Perennial from a short rootstock, the stems simple, eens slender, 2.5-7 dm. high, sub- glabrous and often ee above, long:setose | sg es and m or less stellate. Lower leaves n base m, 8-25 cm m. wi i T > owe Oo ase, oblanceolate to lanceolate, esse ially e ntire, - stem-leaves few, progressively reduced in size, sre ene -hairy and ine onspicuoushy stetinte ; inflorescence rather open, the branching tend- ing to be paniculate, the heads rt ew to many and about 15-20-flowered ; involucre 7-11 mm. high, the phyllaries as conspicuously imbricate; phyllaries linear-lanceolate, finely stellate-pubescent with some gland-tip 63 bristles and some setose hairs often present ; achenes 3-3.5 mm. long, exceeded by the sordid pap Open woods, Transition and Canadian Zones; commonly occurring west of and cheough, the Cascade Si cilntalers from baa h Columbia fe ati i and Psy san ocr a northern California; also occurring less ca ear pa! eastward to Mon and western Wyom Type locality: mouth of the Columbia River. Col- lected by Scouler. June—Aug. ini aah boy species merging to the south with var. nudicaule and with H. cynoglossoides and H. alberti- num in its more easterly range. Hieracium scouleri var. nudicatle (A. Gray) Cronquist, Vasc. Pl. Pacif. Northw. 5: 238. 1955. (Hieracium a Sn var. nudicaule A, Gray, Proc. Amer. aac 19: 68. 1883; H. nudicaule Heller, lictaeies 22149. 1906; Pecan fgets Zahn, Pflanzenreich 428°: 112 1922; H. babcoc. kit Zahn, loc. cit.; H. babcockti var. setosi- oh in, loc. cit.; H ianum Zahn og: to oot, es h cyn glossoides and also glandular. Dry oods or gravelly flats in the southern yf a poo Siativen’ Mountsins, Oregon, south through sasiheie California to Placer Cocnty in the Sierra ere and to Trinity County in the Coast Ranges. Type locality: Sierra County, California. Collected by Lem 2. Hieracium cynoglossoides Arv.-Touv. Houndstongue Hawkweed. Fig. 6036. Hieractum cynoglossoides Arv.-Touv. Spicil. Hier. 20. 1881 ogioie cium ro Rydb. Mem. suet Bot. any 1: 464. 1901. Not Form. 1896. deracin , Bull. Soc. Bot. F Hi | . Planaes: nreich 4750; 1130. ?Hieracium roi st. aa & Warren, Proc. Biol. rs ack: 41: 108. 1928. Habit much zegesobls ing H. scouleri var. scouleri but setose and not subglabrose or meaycone. Leaves and inflorescence as in the preceding taxon; involucres finely seule also with shor blackish, dag sped bristles at some long setae; achenes and pappus as in H. scouleri oe i; albertis nu Wa Sut: t low elevations, Sonoran and Arid Transition Zones; British Columbia and Alberta south to eastern re gion a rn Oregon, and Modoc County, California, pe - Utah and Wyoming. Type locality: no . June Aug. Intermediate between H. scoulert and H. albertinum. 3. Hieracium nae a Farr. Western Hawkweed. Fig. 6037. ig ieracium albertinum Farr, Ottawa Nat. 20 1906. acium absonum Macbride & Payson, Conk: nia Herb. No. 49: 71. 1917. re r coarse perennial 4-12 dm. high, the stems 1 or more from a stout rootstock, leafy espe- cially below, with the uppermost leaves few and somewhat reduced in size, abunda ntly pubescent throughout with long setose hairs and rather sparingly stellate. Leaves 8-25 cm. long, 1- fe) Se) e rrowing to a petiolifo ish, the upper leaves sessile; inflorescence usually with many heads, corymbiform to agro ye corymbiform, the peduncles 5-20 mm. long; involucres 9-12 mm. high, fate fewer as 5 20) ; phyllaries loosely imbricate, linear, with dark mudaes 06: stellate-pubescent and Sona y long- setose usually with black-based bess these sometimes dense enough to conceal the phyllaries ; achenes 3 mm. or more long, t the pappus whitis Dry open places rairies and pen © pine forest ‘Arid T ition Zone; southern British Columbia = Alberta soy to eastern | ashington and northeastern co a tek to Idaho aoe western Montana. Type loc ity: betwee and Moraine Lake, Rthetis. July—Aug. 4. So _— Howell. Long-bearded Hawkweed. Fig. 6038. giberbe Howell, Fl. N.W. Amer. 395. 1901. vere piperi St. John pe Sal Proc. Biol. Soc. Wash. 41: 109. 1928. Perenni i 3-6 dm. high, the stems erect, 1 o e from a short woody Caste rather slender, erect, reddish below and rather s sabe 4 ‘bes et wr tora setose hairs 5-10 mm. long, nearly gla- brous above. Laps pighhg tem- oon withering, the midstem-leaves 65 cm. long, ise uppe what smaller, all eave, ptscheertthy oblanceolate, or elliptic, narrowed to a petiolar t so base, fe hes sessile above, entire or rarely denticulate, thin, green but pubescent with setose SUNFLOWER FAMILY 597 hairs up to 10 mm. long, conspicuously so on midrib and margins, ee glabrous; et ie open, the heads mostly few on peduncles 3-5 cm. long, the flow yellow; involucre to p high, the phyllaries with dark geadrib, lon, often ack based, setose hairs and sparsely stellate ubescent, not glandular ; achenes 3. 5 mm. lon ng, exceeded by white or sordid-white pappus Cliffs and rocky bluffs, Humid niet an Columbia River poe Skamania County, Waikinwtca cad Malone County, Oregon. Type locality : Columbia —— near The gp ronotei> July tinct local form penognies ble by its unusually setose hairs, suggesting Pelatinaaiin with the scoulert- ynog ous rioleal complex on one hand and with the ppc pee penne atum ccanpiex = the other. The habit and the eek veduced and more numerous stem-leaves indicate relationship to the latter gro 5. Hieracium argutum Nutt. Southern California Hawkweed. Fig. 6039. Hieracium argutum Nutt. Trans. Amer. Phil. Soc. II. 7: 447. 1841. Perennial, 3-7 dm. high from a branched woody caudex, the stems beset with long setose hairs below, jae lacking or r nearly so on the upper stems. Lower leaves often densely clustered below, 6033. tartari cium scouleri 6034. os a ee —. cynoglossoides 598 COMPOSITAE 10-20 cm. long, 1+4 cm. wide, oblong or oblong-lanceolate, narrowed to a petiole or a petioliform base, the margins remotely but t deeply eg ae hey tate or -denticulate, setose-hairy; middle and a ae techn sessile, often much reduced, linear, essentially entire, stellate-pubescent to nearly a e€ linear her sparsely black- ‘stipitate-glandular with some cao “stpitate glandular halts, the ie saetbetack sparse; achenes 2.5-3 mm. long, dark brown, the papp whitish. Rocky and wooded big nag Lge Sonoran and Transition Zones; Santa Rosa and Santa Cruz Islands, Santa Barbara County, and on the inland in San hon gee County, where it merges with forms of the following variety. Type si na ~— Barbara. bgt by Nuttall. a Hieracium argu r. pari bg ay) Jep Fi. lif. 1009. 1925, (Hieracium parishii A. Gray, Proc. Amer. arts 19: 67. ness: HW teen ‘Greene Bull Cait. a 1: 195. 1885; _ oa Eastw. Bull. Torrey Club 32: 217. 1905.) Leaves -: . Hi. um var. ar - , Be northern for: ore often denticulate and not dentate; inflorescence usually re Sane a ranched; of the ohyttes fk merely stellate-pubescent (H. parishii) or more Conemeety a with pellow: stipitate ‘lander — the black-stipitate- glandular hairs pliers A hy Jacking or sparse. Dry wooded canyons, Santa Lucia Mountains, Monterey County, California, south to mountains of Los a — and Ped Bernardino Cosun Ss. Tis lo cain rock crevices, Water- man Canyon, San eruue ino Mountain: 6. Hieracium albifldrum iis White-flowered Hawkweed. Fig. 6040. Hieracium albiforum Hook. FI. Bor. Amer. ?Hi ium va vert Arv.-Touv. Sve Hier. 10. 1881. Hieracium helleri Gandoger, Bull. Soc. Bot. F 1918. Hieractum leptopoda m Gan , loc. ci Hieracium candelabrum Gandoger, op. cit. 52. Hieracium albifiorum subvar. rosendahitt Zahn, Pflanzenreich 42: 1123. 1922. Hieracium albiflorum f Ae Zahn, cit. Hieracium albiflorum subyv anh carinens Zahn, loc. cit. ?Hieracium pacificum Perennial, 3-12 dm. h ee = stems usually eat te arising from a woody root and commonly unbranched short caudex, desthes ss or with few stem-leaves, with long setose hairs below and mostly naked above. Leaves mostly basal, 8c ng, Shige eolate, narrowed to a petiole except in the stem-leaves, moderately pyres entire or repand-denticu sy i inflorescence corymbiform, the rather small heads several to many, the peduncles oie , 2-7 cm. long, the flowers white or pale yellow (?); involucres narrow, usua salty 18-30. flower ~ phyllavies: saree imbricate, the longer inner ones 7-9 mm. long, line ar-lanceolate greenish or male ckish, glabrous or often sparsely olathe lar te pale or black hairs, sometimes with a few setose hairs ; Schenes about 2.5-3 mm. long, the pappus white or sordid, longer an the achene Open ah in moist or dry situations, Upper diuaewk Zone to Hudsonian Zone; Alaska and northwestern Canada ne to southern California and Colorado. Type locality: ‘‘mouth of the Columbia.”? Collected by Scouler. othe widely distributed species in the Pacific States both altitudinally and geographically. Quite variable as to type or pie of the lower leaves and in vesture y = A sa dpc es having in common proett habit, linear- lanceolate phyllaries, absence of stellate pubescence, and ream-colored flowering hea 7. Hieracium greénei A. Gray. Greene’s Hawkweed. Fig. 6041. a greenet A, Gray, Proc. Amer. Acad. 19: 69. 1883 m howelli A. patton Bot. Gaz. 13: 73. 1888. "Hirao barbigerum Greene, Pittonia 3: 228. 1897. yee rage 1896. teracium oregonicum Zahn, Pflanzenreich 42%: 1130 Peskaiies 1.5-3.5 dm. high, the stems few to several from a iy a de ig mont branched caudex, stoutish, erubnt aged densely sicliase tecavlake throughou lower leaves also long-setose. ie and low stem-leave s 4-10(15) cm. long, rather thick, oblanceolate to an late, narrowed to iolar tiene the base shorter and becoming sessile, the m one or irregularly sebelah desdicadete Aereecbnce panienes e with hve varicate ‘brane hlets, nthe sev BES Ow Leo = rs ou =b (=) Oo o8 Re oo - 5 Oo wn Bp & ao Ld ~ Ree bat ERE vy | = oO a Oo has | ae ae 3 3 > =] itis) bry w tory S, a es 5 25 eho o@ rrow, -NOW' m broadly linear, obtusis apes, pie is sri and more or less ste te-pubescent ; achenes about 5 mm. long, the pappus onger achenes, white or brownish. = rocky places, often in serpe ee iain and Canadian Zones; ‘Klamath County in the south Cascade Mountains and Fodeohine ‘County in — Siskiyou acer age ag| Oregon Humboldt County, Cc California, and Shasta County and also g as far as Sierra County. Tone inca pine woods, Scott untain, Si aves County, Cali forts. = sn In some of the Californian specimens, he stellate pubescence is quite sparse but the plants possess the char- acteristic stiffly divaricate inflorescence of H. greenei and are here considered to be aberrant forms of that species. 8. Hieracium bolanderi A. Gray. Bolander’s Hawkweed. Fig. 6042. Hieracium bolanderi A. Gray, Proc. Amer. Acad. 7: 365. 1868. Hieracium siskiyouense M. E. Peck, Proc. Biol. Soc. Wash. 47: 188. 1934. Rather slender perennial 1-3 dm. high, the glabrous and glaucous stems solitary or few from a | simple or branched, woody caudex, unbranched below the inflorescence, Seni pean abe a inflorese asal lea late, sessile (the longest leaves usually with pétiolifermn bases), thin, entire or obscurely denticu- late, conspicuously but not densely long-setose ; inflorescence open with sseendiee branches, the SUNFLOWER FAMILY 599 heads 3-10, borne on slender, glabrous or Med sparsely glandular peduncles (2)4-8 cm. long. about 5-10-flo wered, -- ei rarely white; involucres na epi the phyllaries few, scarcely imbri- ate, the 1 p to 1 cm. long, . 1.3 mm. wide, acute ap ex, often perme -tinged a au- cous, entirely abouk 4 or oe ight ery few s ikea gland- tipped hairs ec = occasional long oeta: achenes about 3 mm. long, the ae exceeding t the achen s, white or whitish. d or ~— _ apes Arid ee and Panett Aas’ mountains of Curry, For iy pie and Jackson Counties, Oregon, south to Siskiyou and Humboldt Counties, California. Type locality: Red Mountain, Per eetdt aN June-Ji bite. 9. Hieracium gracile Hook. Alpine Hawkweed. Fig. 6043. Hier ee acile Hook. FI, we Pomp 1: 298. 1833. Hieractum hookert Steud. Nom 422 763. 1840 Hieracium triste var. oie - ray, Bot. Calif. + 441, Hieracium gracile var. detonsum A. Gray, Proc. Amer. Aca Hieracium gracile subsp. gracile var. detonsum subvar. ie bea Pflanzenreich 4°; 1133. 1922. nnials, 1-3 dm. high, the subscapose slender stems clustered, few t ral from a hor sce. pcr and short caudex, simple or sometimes branched, Ba me a 2} plots WA, gus ! 1b } « Ds re ins 6037. Hieracium albertinum 6038. i dens 6039. Hieraciim argu’ 6040. Hieracium rr boca 600 COMPOSITAE above, to glabrous. Leaves 3-10 cm. long, many-clustered at the base, the occasional stem-leaves srantie bracteate ; spatulate to oblanceolate, narrowed below to a petiole 1-5 cm. long, essentially glabrous; heads yellow-flowered, solitary or 2-15 in racemes, pedunculate; involucre high, the phyllaries with black, usually gland-tipped setae or these sorts or lacking, also often stellate- Nee achenes 2 mm. long, surpassed by the white p Meadow open places, Boreal Zone; Alaska and Canada south in Hes Pacits States to the Siskiyou hs and the ee ning Nevada in California and the Rocky Mountains region south to northern New Mex Type locality: Rocky Mountains. Collected by Drummond. July—Sept. 10. Hieracium horridum Fries. Shaggy Hawkweed. Fig. 6044. Hieracium horridum Fries, Ep — — 154. 1862. Pilosella relicina Sz. Sz. Flora 4 Hieracium breweri A. Gray, Proc. ne ee por 62/553. 1865: Rather slender pernie! 1-3 dm. high, the tenagd Elackered, simple or with few slender lower ranches, arising from a woody ro Sta and stout, simple or branched caudex, oe to nearly glabrate and also with fohee oe te hates 6 these la Hee nodes on the upper stems. Basal an lower stem-leaves many, m. long, phicacce nate to sale aut pee narr red to a petioli- form base, shaggy with long, nie es g, setose, white or brownish hair ahs Thats Selene ae bracteate inflorescence sessile, similar, longer than the internodes and su greatly reduced; florescence Bae branched, corymbiform, with few e y heads, the heads a small, welin ow, miele red, the pedu uncles mostly 1-1.5 = lon sate e 7.5-10 m m. high, the phyllaries cely im bribed cate, the er series very shor atelinte ‘b aie giibeate. the usuall y darkened midsectio n with long, fate blacks based setae; maatern about 2 mm. long, the pappus tawny, longer than oe ks, mostly Boreal Zone; eastern Lane County, Oregon, in the Cascade Mountains, to the tice methert “California Bay southward in the Sierra Meveee to Slare County; also the higher mountains of ‘Sauen California. Type locality: California. Collected by Bridges. July—Sept. 11. Hieracium SRsoE gig Canada Hawkweed. Fig. 6045. Hieracium canadense Michx. F1. Bor. Amer. 1803 Hieracium columbianum Rydb. fant nate Pak, 28: 513, 1901. ial, 4-12 dm. high, the stems 1 o e from a short woody caudex, rather stout, pu- hegeene below with long spr caiiie ete) labo us to sparingly stellate-pubescent above. Basa and lowest stem-leaves rather small, s withering and deciduous, the midstem-leaves about 6-12 cm. 1-4 cm. wide, cage Slee: east the internodes, mostly lanceolate to oblanceolate, entire or sparingly denticulate, sessile and subclasping at base, rather thin, the margins and often veins and undersurface with spreading setose hairs, rarely sparingly ’ stellate- Pabeseseik: in- florescence corym rary o-akagene e ag spreading peduncles, the heads 6-20 or more, yel ow flowered ; involucres 6-10 m igh, the phyllaries imbricate, dark, glabrous oo or sO; © eetian about 3 mm. long, purplish bled, oo rites tawny, a little ‘longer kan the Woods pa moist open places, Transition and lower eee Zones; Canada south to Wha tseton: the Rocky a region, and eastward to New Jersey. Type locality: Canada. July—Sept. Much resembling H. auher 12. Hieracium umbellatum L. Umbellate Hawkweed. Fig. 6046. racium umbellatum L. Sp. Pl. 804. 1753. Hieracium caesarean! Schwein. Long. Exp. 2: 394. 1824. Not H. umbellatum var. scabriusculum Farw. 1927. Nutt. —— Amer. Phil. Soc. II. 7: 446. 1841. Hieracium suksdorfii G ; Bull. Soc. Bot. Fr. 65: 51, Perennial, 412 dm hi ae the stems 1 or more from a short woody rootstock, stout, glabrous or nearly so below, usually stellate- Sher ulent above. — and — stem-leaves rather small, soon rithering and deciduous, the midstem-leaves 6-10 cm. long, 1-: . wide, sessile, not ianping nd someti narrowe ‘t se, many, much longer goose the fesindis.: entire or sparingly denticulate, lanceolate to narrowly oblong, rather firm, stellate nat toes or less scabridulous « e pubescent, the heads 3-10 or more, yellow-flowered; involucres 6-13 mm. high, the phyllaries imbric aed gla — or staat a so; achenes 3-3.5 mm. long, reddish, the pappus tawny, about equal- ing t chen Moist Pe Se and woods, Transition and lower Boreal Zones; circumboreal southward in North — to Michigan and Wisconsin and i in the west to Colorado and northwestern Oregon. Type locality: Europe. July—Sept. 13. Hieracium aurantiacum L. Orange Hawkweed. Fig. 6047. Hieracitum aurantiacum L. Sp. Pl. 801. 1753. Resembling H. pilosella in habit, the stems 2-5 dm. high, —_—e- bearing 1 or 2 reduced leaves, long-setose, and also stellate and glandular above. Leaves 4-20 cm. long, oblanceolate to elliptic, narrowed to a petiolar base, usually ss above and below; inflorescence a form, the heads 5-25 on short peduncles, the flow red-orange; involucre 6-8 mm. high, t yllaries long-setose and also bearing black, Shand teal hairs, peti slightly tomenasane tienes pappus as in H. pilosella. A European weed sparingly established in Bremerton, Kitsap County, Washington, and in Multnomah and Deschutes Counties, Oregon; also introduced i a eastern United States. June-Sept. King-devil. SUNFLOWER FAMILY 601 . Hieracium greenei an ‘ ncn asoseyghas . Hieracium horridum 6045. Hieracium canadense 602 COMPOSITAE 14. Hieracium pilosélla L. Mouse-ear Hawkweed. Fig. 6048. Hieracium pilosella L. Sp. Pl. 800. 1753. Stolon oii perennial, the slender stems 5—20 cm. high arising from a rather slender t- stock, scapose, rarely with a single leaf, more or less spreading-pubesce h gland-tipped, often black hairs 12 ng (those f the stolons small), oblanceolate to spatulate, tapering to a petiolar base, entire, pale beneath with a dense stellate pubescence a ith som se hairs, green a glabrous exc long setose hairs; heads solitary, yellow- d; in- volucres 7-11 igh, the phyllaries stellate-pubescen at and also with spreading, often gland- mm, high, t tipped, black hairs, ese lal ith long setose hairs; achenes about 1.5-2 mm. long, the pappus exceeding the achene, so A European weed spari tor stieRiahad in lawns and waste places, Willamette Valley and Multnomah County, Oregon; also introduced in eastern United States. May—Sept. 174. CREPIS* L. Sp. Pl. 805. 1753. Perennial, biennial, or annual herbs; stems scapiform or branched. Leaves chiefly basal, the cauline alternate, many — or deciduous. [From the Greek, meaning sandal, the ancient name of som plant. ] A genus of 196 species, natives of Eurasia, Africa, and North America. Type species, Crepis biennis L. Leaves dentate or parted, not novos or obovate; not dwarf alpine plants. troduced annual or biennial species (perennial in ie ipragann above capillaris, Achenes beaked, the beak slender or sacs Perennial; beak filiform, about twice the length of the achen i. rsifolia. Annual or biennial; beak n ae fibtotan: about the length ‘of ae ‘bene or slightly Lig: Be oi: Involucres and stems strongly setose Aged ovale bristles Involucres and tulose with ne black ee re ON aoubveais Saviinastfulta: . high; native perennials. Stems and leaves ar aE glabrous and glaucous; cauline leaves absent or rem so; plants of moist situat 6. C. runcinata, Stems and 1 won r less para get = addition to A r058 types of sthacthiee: 1-3 cauline leaves ose Rory es C. : iheratur pa); of psig situ Herbage and ‘eveluetas wba? Sok ahh gland- duoat bite: outer heared i nia lh -lanceolate. Herbage and involucres variously pubescent, if glandular of short hairs only; outer phyllaries lanceolate or ovate- or r bot mspicuously setose but not sine Inflorescence a 1-9 ae sone phyllaries 10-15. . C. modocensis. Inflorescence of 6-70 heads; inner phyllaries 5-10. - C. barbigera. setae sparse or terra Heads 5—10-flowered; aed phyllaries 5-7. Cauline leaves 1-3; phyllaries labs or evenly and tad tomentose. . C. acuminata. pe leaves reduced or weone: phyllaries elabrovs - midportion, conspicuously omentose on the margin C. pleurocarpa. Heads a flowered; inner i s 8-10. Divisions of leaves linear or narrowly lanceolate, mostly entire; achenes greenish. 12. C. atribarba. Pipi Se of dares lanceolate or deltoid, some usually toothed; achenes yellowish or Plants ‘mostly 3 dm. or less high; involucres broadly cylindric; heads 12—30- owered Leaves grayish-tomentulose, not glandular; ee ee not aS parent above. 1 It. Leaves green and glandular; peduncles fistulose Fie ss he yl above. Plants mostly 3-6 dm. high; involucres soled” de og es heads 8—10-flowered. Se weit Leaves all entire or few-toothed, spatulate or obovate; dwarf alpine plants. C. na 1. Crepis bursifdlia L. Italian Hawksbeard. Fig. 6049. Crepis bursifolia L. Sp. Pl. 805. 1753. Perennial, 0.5-3.5 dm. high, stems several, decumbent or arcuate, arising from a woody caudex and vertical root, E plese: tomentulose, cymosely branched above, bearing 2-14 heads. . and text a ed from Babcock and Stebbins (Carnegie Ivst. Pub. No. 504, 1938) and Babcock (Univ. Calif. Pub. Bot. 22: 1947), where complete synonymy is also to be found. SUNFLOWER FAMILY 603 Leaves mostly basal, the cauline few, linear and bract-like, the basal leaves 5-25 cm. long, oblanceo- late in outline, lyrately pinnatifid, the obtuse terminal portion dentate or denticulate; heads 30-60- flowered, on peduncles 0. cm. long; aie cres 9-11 mm. long, cylindric, canescent-farinose ~ sometimes somewhat yellow-setulose ; outer phyllaries unequal, one-fourth to one-third as lo ong a the inner, these 8-10; achenes 6-7.5 mm "tae 10-ribbed, pale brown, fusiform and seh bite a filamentous beak about twice the length of the body of the achene; pappus mm. long. A native of Italy found growing in waste places in the San Francisco Bay region, California (Berkeley; Stanford). May—June. 2. Crepis capillaris (L.) Wallr. Smooth Hawksbeard. Fig. 6050. Poe misc Sp2Fk 812: 1753. Crepis v cit. ed. 2. 1134. 1763. Crepis apilaarts Wallr. Linnaea 14: 657. Malacothrix crepoides A. Gray in J. G. Cooper, Pacif, R. Rep, 122: 53. 1860. Annual or biennial, 0. oy 9 dm. hi gh, with several stems from a ig thse ae erect, rather Pic, ilspicialous at the base and sometimes above. Basal leaves 3-30 c g, la nceolate in outline, runcinate-pinnatifid or rately pinnate, petiolate, glabrous or tevidiious beneath on the 6046. Hieracium umbellatum 6048. Hieracium pilosella 6047. Hieracium aurantiacum 6049. —_ bursifolia 604 COMPOSITAE midrib with ser yellow | hairs, sometimes sparsely oe above, me cauline leaves ecesils, e oO bifo an 60-flowere , peduncles 0. cm. long; involucres 5-8 mm. high, ett bang in fruit ; peak phyllaries ay Bin to one-half the lengt th of the inner, glabrous or tomentose, the inner phyllaries 8-16, lanceolate, glabrous, tomentose or sparingly gla en sometimes spars sely setulose with black glandular hairs ; receptacle glabrous; achenes 1.5-2.5 mm. long, 10-ribbed, “heuuteids yellow to dark brown, fusiform and beakless ; pappus 3-4 mm. Habe A native of Europe, eivelig in meadows, lawns, and waste places, widely introduced in the Pacific byron from western Washington to central California; also occurring in Canada Seite eastern United States. May Crepis nicaeénsis Balbis ex Pers. Syn. Pl. 2: 376. 1807. Ann r bienn ial, 2.5-11 dm. high, po el to and posse C; capillaris; differing in having more eh a oy pate: involueres 8- /?— pris high and cg st. ger gener is fruit; receptacle aero achenes 2.5—4 mm. long, golden-brow of sonal n.Europe and s gly introduced in eastern United States, and known in the Pacific’ Pe pial one col- fenton at WMaryeviie.: ‘Cohutta: County, Washi ington repis pilchra L. Sp. Pl. 806. 1753. antints: 3-7 - . high from a slender taproot, pubescent below, glabrous above; basal leaves 3-15 cm. long, oblanceolat ate, acute or obtuse, runcinate-pinnatifid, eae toa winged petiole, ‘pubescent; stem-leaves somewhat reduced, etic “ahd amplexicaul and less divided than the basal; involucres cylindric, glabrous; achenes 4-4.5 mm. long, the outer compressed and often without pappus, the inner mica rin i A egon. rope rubra L. Sp. Pl. 806. 1753. A garden annual, native of Italy and the Balkans, mostly 1-2.5 dm. os wri — dentate or Dah: yee pinnatifid leaves an nd sc gy Feria stems, os branched below, termi- nated by pink- or white-flowered heads, nodding before anthesis; achenes 9-20 mm. long, with a coarse or slender beak as long or twice as tong as the body. Rarely escaping from cultivation; in the Pacific States it has been collected at petty Marin County, California Cre me: LocSp. Pi 807; 1783. Glatinun or puberulent annual, 1-10 dm. high, with. runcinate- pinnati if teal gsr Poze Scaatie, linear cauline leaves; icccenenre usually with few heads; inyolucres cylin- dric-campanulate, 9 m igh, the inner phyllaries pubescen on the inner faces; achenes usually 3-4 mm. long, 10-ribbed, dark pba: — fusiform and attenuate se i but scarcely beaked. 7% native of Europe, sparingly introduced in Canada and ihe aed Staten: on the Pacific Slope, introduced in British Columbia and to be expected in northern Washingt 3. Crepis setésa Hall. f. Rough Hawksbeard. Fig. 6051. Crepis setosa Hall. f. Roem. Arch. Bot. 12: 1. 1797. Annual, 1.8-8 dm. high, the erect stem from a taproot, simple or branched from near the base and also above, hispid. Basal leaves mostly few, 6-30 cm. long, oblan regs in ou = ne, narrowed i i ion oot at ape abov eads : ; : hispid and ochoae with yellow bristles; involucre mm. long, somewhat turbinate in fruit, the outer ap hoe not more that one-half the length of the inner, the inher linear-lanceolate, thick- ened at the base, setose with yellow nef pete the yellow ligules often reddish on outer face; re- ceptacle sk ge Pe 7 shoe 3 .5-5 mm. long, 10-ribbed, fusiform and tapering into a beak .5 mm. long; pappus 2.5-5 mm. long. A native of sae Europe, Sa sc pl" Sai not common in the Pacific States; in the Willamette Valley, Oregon, and Humboldt County, California. Jun ug. 4. Crepis vesicaria subsp. taraxacifolia (Thuill.) Thell. Weedy Hawksbeard. Fig. 6052. a Thuill. Fl. Par. 409. 179 Cres vestcaria a oitea taraxacifolia Thell. ex a & Keller, Krit. Fl. Schweiz ed. 3. 361. 1914. Annual or biennial Coe perennial), 0. the dm. high, stems 1 to several from a thickened —— alten branched near the base as well as above, more or less purplish, striate, more or less tom Leaves mostly paral. the cauline leaves sessile and amplexicaul, made! entire to pin- nately aad the tae leaves 10-20 cm. long o what longer, mostly narrowly oblanceolate in ease mostly rancinate-pinnatifid or lyra sea ere with a large rain lobe, I pubesc on both surface .. = gla ss hairs; inflorescence istiiskr set m, with s heads, pe icon cles 1-13 om vee wolne e 8-12 mm. high ; outer phyllaries short, Rencncetig or ovate-lance oat, calaiee the A As lel 9-13, lanceolate, obtuse = en glan- ular-pubesc and usually with ‘black glandless setae ; eeceiaele ciliate; act mm. se Ocribbed. pale brown, fusiform, attenuate into a slender beak as long as or oat than the ~ the achene ; pappus mm. ‘long. riable siibiagiedies native of Europe and widely introduced elsewhere, becoming more abundant in waste places an ot borders of woods in California; A ec a te Marin, Alameda, cy Mateo, and Santa Clara Counties, and also adventive in Los Angeles County. April-June oe 8 #8, Comal 5. Crepis nana Richards. Dwarf Hawksbeard. Fig. 6053. Prenanthes pygmaca Ledeb. Mém. Acad. St.-Pétersb. V. 5: 553. 1815. Crepis nana Richards. in Frasil. lst Journ. Bot. ps6: 7s Bee -A 833. Crepis humilis Fisch. ex Herder, Bull. Soc. Impér - Nat. oe 43: 190. 1870. Dwarfed tufted abo acer the vertical or creeping, underground stems arising from a taproot, the leafless uppe ag ees sued ae t ie m. high, about as long as or a li ie ee mags she sie s. ves ofte mewhat purpli Bs many-clustered basally, u cm. vate-acute to pebcrtacnlas: the blade tapering ‘abruptly into the petiole, eibein. entire or a tee sda ceed: involucres 24 on the est ranches; inner phyllaries about 0-13 mm. long, subtended by the paivcatan outer pei es a ge very short, Sele lee SUNFLOWER FAMILY 605 hana with purple; achenes sara striate, pias aac smooth or rugulose ribs, somewhat narrowed and attenuate above white, 4-6 m ue stly in loose gravel and sick "ee Alpine Zotie? ra te ne to the Atlantic Ocean and south to Utah and Nevada, and in the agg States from Washington to = higher peaks of southern California; also eastern Asia. Type locali Arctic ast “O per-mine River.” July—Aug ; sy pe, from C. su y the br ith heads borne well above the bas ves. British Cotuanhie ay Idaho south a Tulare County in the southern Sierra Nevada, California. mat Lecsite: above Lake Constance, Jefferson County, Washington. 6. Crepis monticola Coville. Mountain Hawksbeard. Fig. 6054. Crepts occidentalis var. crinata A. Gray, Bot. Calif. 1: 435. 1876, in part. Crepis monticola Coville, ee U.S. Nat. Herb. 3: 562. pi. 22. 1896, Perennial, stems 1.3-3. é ws high, from a simple or 1-forked caudex and a s Fee hong near the base, spar and den leaves 10-20 cm. long, pinna atifid eit rather tt Be angular, dentate lobes, atten a broadly winged betiols the sae -leaves every gradually reduced above and becomi ng os div ided, eiuiie, what i the pubescence of the leaves like that of the stems; i inflor res- of 4-20 he At the seh ads 16-20-flowered; involucre campanulate, densely hirsute with long ppc om hairs ; phyllar s 18-24 m Spee sak e inner lanceolate, attenuate at the apex, the i fe) eed lanceolate to linear; achenes 5.5-9 m he pappus t woody root Ope Arid T sectiens Zone; Swshkiet: oR and Lake Counties, Dremiais ig in the Coast eunges wee County, California, and Mount Hamilton, Santa Clara Count - nd fro A oc County south to rag County. Type locality: Yreka, Siskiyou County, California. May- 6050. Crepis capillaris 6051. Crepis setosa 6052. ees vesicaria 6053. Crepis nana 606 COMPOSITAE 7. Crepis occidentalis Nutt. Western Hawksbeard. Fig. 6055. Crepis occidentalis Nutt. Journ. Acad. Phila. 7: 29. 1834. Perennial, stems 1-3, fog stly 1-4 dm. high from a somewhat swollen Nae and long slender root, ‘es stems and leav » witha close dense tomentum. Basal leaves long, variable, laciniate-dentate to pinnatifd w ith toothed ety ‘the stem-leaves somewhat feduced and sessile; ads 12- ed NW wm 10 mm. long, af ribbed, slightly attenuate at the apex, mostly medium brown, saepaltat t shorter ore the pus. Dry rocky ens ie Transition Zone; southeastern Washington south pote: Nevada and northeastern California heme the east face of the an Nevada to the Mount Pinos region and Bear Valley in the San Ber- nar 7 va ns; also from Idaho and w n Wyoming south to New ee Type I ocality: “‘on the borders an the Wicinity of the river Columbia. , timid: Bd id hers May—Jul is occidentalis ra costata (A. Gray) Babe. & Stebbins, Ciriveiss Inst. Wash. Pub. No. 504: 124. 1938" “Ct repis occidentalis var. costata A. Gray, Bot. Calif, 1: 435. 1876; C. grandifolia Greene, Pittonia 3: 107. 1896.) Stems —4 reaps Bah involucres, peduncles, and usually the upper cauline leaves with conspicuous, dark Garey pate). gland. Seped setae; gare ory rather narrow, of 15-30 heads, the heads 10—14-flowered, usually = I 8 in phyllaries. Gravel y or ro open slopes and flats; common from British Columbia southward, the Ca Bite Mountains, to motthark: California and east to South Dakota. Type locality: Stansbury Island, Peng pe Lake, Utah. repis occidentalis f Sueae. poets (Rydb.) Babe. & Stebbins, Carnegie Inst. Wash. Pub. No. 504: 128. 3 (Crepis pumila Rydb m: N.Y. Bot. Gard. 1: Re a 0.) Stems 1-4 dm. high, Prag 8 above, glandular pene eoeeeerety See ie heads few to many, 1 0-flowered, the inner phyllar Rocky open slopes and flats, ring locally in the mountains of ti ble x iA and eastern Washingtin io more com- mon in Nevada, teas ae rn | pote California as far south as Kern and Ventura Counties. Type locality: Bridger Mountains, Montan repis occidentalis oe sp. conjuncta (Jepson) Babe. & Stebbins, Carnegie Inst. Wash, Pub. No. 504: 134. 1938. (Crepis ses Sen var. fouivacta Jepson ex mers * Sg trig, (oc. cit. : (Ge porienigec var. nevadensis Kell. Proc. Serer Acad. 5:50. 1873, in part.) Stems 0.5—2 high, branched from near the base, the cg ches Counties, Oregon, and the mountains of adjacent California south to Mendocino County, and souk A n the Sierr: Nevada _to Placer County; also southeastern Washington and wtheecatet? gr pioac Type aeality: Cisco, Placer County, California 8. Crepis bakeri Greene. Baker’s Hawksbeard. Fig. 6056. Crepis bakeri Greene, Erythea 3: 73. 1895. ennial, stems 0.8-3 dm. high Ries a Sety ine ex and slender fee pom reddis also glandular, the stem-leaves few, reduced above, sessile; inflorescences ds, the eads large, 11-40-flowered, on stout peduncles; involucres 11-20 mm. high, vhroadle cylindric to cyathiform, dark green; inner phyllaries 10-14, lanceolate, sparsely toatl entose and cons es Vv to irds i - a. mm. long, slightly contracted at the a pet 13-ribbed, yellowish to dark brown, ‘ie. pappus about equalling to a aa — than 1 ac hen Rocky slop rid Transition Zone; wakes Washington in Klickitat and Kittitas Counties; outhern Draka in Cake, ghee od “Josephine ig ooret in California south in the Coast Ranges to Mendo- cino ty a Pe oy in eastern California from Cou o Placer County. Type locality: Egg Lake, Modoc oun bakeri uae cusickii (Eastw.) Babe. & Stebbins, epeste Inst. Wash. Pub. No. 504: 140. 1938. oat se ‘sick Eastw. ae oye Club 30: ae 1903.) Di ig g from C. baher. rscrong” Sobers in having involucres 10-14 mm. long in flower. 13-17 m * ve in fruit, the p: s 6-9 mm. long, and achen atten mel at the Be rather than ‘ef seresty bisected: Lake and Jackson Uatiaicn, Gran, — to Siskiyou and Lassen Counties, California. Type locality: 15 miles east of Ashiand, Jackson County, Oregon 9. Crepis eee Greene. Low Hawksbeard. Fig. 6057. oo eeaggelst da Greene Jey 1895. ovi ille, Con r-U.S. tate Herb. $2565. pl. 24. 1896. erennials, rcs 1 to several, rather slender, 0.5-4 dm. high, from a simple or 2-4-forked caudex and woody root, sparsely leafy, branching above, glabrate to tomentulose and bear- ing stiff sella setae. Basal leaves narrowly elliptic in outline and deeply pinnatifid or bipin- natifid, the segments acute; stem-leaves sessile, less divided, glabrate or tomentulose but bearing some stiff yellowish setae on the is; inflorescence of 1-8 heads, the heads 10-60-flowered ; involucre cylindric-campanulate, the phyllaries 13-16 mm. long, i ther | ire beset throughout with stiff black setae; achenes 7-12 mm. long, som ewhat striate to nearly smooth, dark, greenish black to reddish brown, mostly oT the pappus in length. Sagebrush slopes a Arid Transition and Canadian Zones; Montana to British Columbia south- ward, east of the Camtede ¢ Mountains ios one — ad Washingt ging to northeastern California and east to Colorado. Type loc ssa lava beds, y, Califor = odocensis Ae i bia (Kell, Babe. ie meee. Carnegie Inst. Wash. Pub. No. ima ie. 1938. (Crepis occidentalis yvar.- Subacerts Kell. Proc. Calif. sap i 50. 38745 og . occidentalis var. nevaden Kell. loc. cit.) Stout stems 6-20 c high branching from e base; setae of stem and petiole straight an a a involucres sparsely heset with black setae, oe se Rapes 7 A 0.13.5 mm. long, than eq the achenes, which a rede rk, re or less brownish, and not on all beaked. atom places, greet r Mountains. Take sg ot Oregon, south t Bearaid County, California, and also in the pate Bernardin no Mou Type locality: Cisco, Plac r Cou unty, Californ s modocensis Sigel (Cov.) Pele, & Stebhine, Carnegie tisk Wash. Pub. No. 504: 152. 1938. {Ci Se dy i usivate Coville, cone U.S. Nat. Herb. 3: 564. pi. 25. 1896; C. occidentalis var. crinita A. Gray, Bot. SUNFLOWER FAMILY 607 if. 1: 435. 1876, in part.) Stems 1.5-3 dm, high, rather stout, densely covered with c or tomentulose; involucres 14-17 mm. high; i pappus- ee a 10 mm. long, shorter than i da lumbia sout to Klickitat york bil ery ioe Type locality: n Crab ashington. Crepis Oca ensis subsp glareésa (Piper) Babc. & Stebbins, Carnegie Inst. wie ae fa) 2 ee te: 1938. ( vars glareosa Bag ssi! yo Saal — 28: 42. 1901.) Stems 0.6—1.3 dm. high, tomentulose, the whitish crisped setae lacking o arly so; 5 anaes s 11-13 mm. high, the phyllaries hirsute with whiti pappus- printies shorter “than the “ore itish crisped setae; nish oe yellowish ream which are attenuate at the apex. Known only from the type locality at Riconane: “Kittitas County, Washington 10. Crepis acuminata Nutt. Long-leaved Hawksbeard. Fig. 6058. Crepis acuminata Nutt. Trans. Amer. Phil. Soc. II. 7: 437. 1841. Perennial, 2-7 dm. high, stems 1-3 from a swollen, 2-3-forked caudex and woody, deep-seated root, stout, tomentose at least at the base. Basal leaves 1 cm. long, pinnatifid with rather even lanceolate lobes, attenuate into a stout, narrow-winged petiole, densely to sparsely canescent- e€ ile, reduced above; inflorescences with mig fe hea x oe heads flowered, the peduncles short and slender; involucres 9-1 ong, cyli c-cam- anulat te; inner eS 5- lanceolate, glabrous or tomentulose, the outer cieviiaties 87 ing ceo- Eas, deltoid, mostly 1-3 mm. long; achenes 5.5-9 mm. long, aa 12-ribbed, pale yellow or brown- ish, equaling the iy Ae Dry ope ces, Arid Transition Zone; eastern Washington. southward to the mountains of southern Cali- fornia, cast to rnatews and south to northern Arizona and New Mexico. Type locality: ‘‘Plains of the Platte.” ay—Aug FSC, bee: a 608 COMPOSITAE oe wes lag nae 1% ice Naked-stemmed Hawksbeard. Fig. 6059. ay, Proc. Amer. Acad. 17: 221. 1882. pies witermeiie ar ae se rpa A. Gray, ee Fl. N. Amer, 1°: 432. 1884. Crepis acuminata var. pleurocarpa Jepson, Man. FI. Pl. Calif. 1011. 1925. al, stems mostly 2-4 dm. high, from a simple or 1- forked, thickened caudex Cr form base, the stem-leaves very few, sessile, much reduced abov e; inflorescences mostly f 15-3 heads, the wine sma au 12-flowered ; involucre land. -campanulate, 8-16 m ong, the phyl- laries densely fl tomentulose, the midportion at, Ral ote. ‘glabrate ; inner phytare mostly 5(6-8), certie a ith scarious nape fete r 5-6, 1.5-4 mm. long; achenes 5-8 long, 10-ribbed, dark brown, shorter than the p In forests or on rocky slopes, goals Arid Trans sh ion goede Curry County to Douglas County, Oregon, and the mountains in adjacent California south to Lake ‘County: oid Wosstebne Mountains in central Washington. Type locality: heateaters of the ' Sareaheauis River. Collected by Pringle. June—Aug. 12. Crepis atribarba Heller. Slender Hawksbeard. Fig. 6060. Crepis occidentalis var. gracilis D. C, Eaton, Bot. = Expl. 203. 1871, in part. Crepis atribarba Heller, Bull. Torrey Club eo 314. 1899, Crepis gracilis Rydb. Mem. N.Y. Bot. Gard. 1: “461. 1900. Not. Hook f. & Thoms. 1876. Crepis exilis Osterh. Muhlenbergia 1: 142. 19 n part. Crepis atribarba subsp. typica Babe. fat Erba Inst. Wash. Pub. No. 504; Suppl. 1939. eoeonis, stems 1.5-3.5 dm. high, rather slender, 1 or 2 from a simple or 1-forked caudex and slender woody root, usually branched at or above ‘the middle, ericishitemieheuition, becoming glabrate. Basal leave wis cm. long, pinnatifid, with lanceolate attenuate lobes; stem-leaves similar, pew aes ced i e, pubescence like the stems; inflorescences of 3-18 heads, the heads up to 65-flowered; cine Ke oun campanulate, canescent-tomentulose to glabrate ; phyllaries 14 mm. long, lanceo late, the inner and sometimes the outer with black glandless — : achene m. long, rather strongly pitied. mostly greenish, about equaling the pappus Grassy open places and in yellow pine forests, Arid Transition Zone; common from British Columbia to : Montana south : o Colorado, Utah, Nevada, and central Oregon. Type locality : Lake Waha, Nez Perces County, daho. May-— s atribarba subsp. originalis Babc. & Stebbins, Carnegie Inst. Wash. Pub. No. 504; Suppl. 1939. (Crepis a ocladuialis var. gracilis oa Eaton, ore Pray | Expl, 203. 1871, in part; C. ag eth Rydb. em. Bot. Gard. 1: 461. 1900, Not Hook. f. & Thoms. 6; C. exilis subsp. originalis Babe. & Stebbins , Carnegie Inst. Wash. Pub. No. 504: 162. 1938.) Stem ~ a 3.5-7 dm. high; leaves pinnatifid with linear falcate divisions and groaned glabrate; heads 10-40 n inflorescence, ie flowered; pie aries nearly or completely without setae. Grassy slopes, mostly at lauae pewwtions than the Keon bod and less common in the Rocky Mo ai ein Ty Semcebhe Columbia to Montana south to Utah, Nevada, and c vee Oregon. Type locality: near Hedle Brit 13. Crepis intermédia A. Gray. Intermediate Hawksbeard. Fig. 6061. Crepis intermedia A. Gray, Syn. Fl. N. Amer. 12: 432. 1884 epts acuminata var. duterieedin ral Man. FI. Pl. Calif. 101 1.51925, Perennial, 3-7 dm. high, stems 1-2 — a swollen, simple or 1-forked caudex and woody, deep- Denier root, rather stout, senda at or above the middle, more or less canescent-tomentose. Basal leaves 15-40 cm. ong, pinnatifid, the janeeeiate lobes entire or dentate, narrowed below into i i th -leav i e; ostl ; involucre cylindric-campanulate, 16 mm. long, canescent, glabrous, sometimes with a few g _— less setae; inner phyllaries 7-8, lanceolate, outer ba dae 6-8, lanceolate-deltoid, one-fift one-third as ode * as the inner ; achenes 5.5-9 mm. long, 10-12-ribbed, vellowinh to brown, toceer than ae e€ pappu In forests aoe open places, Canadian Zone to Arid Transiti Zone; eastern Washington south to Inyo cated hy B Caltornia also southern Alberta to Colorado, northern falsiia 5 ates) New Mexico, Type Seaiie: in aa cated ne (Univ. Calif. Pub. Bot. 22: 601. 1947) as Yosemite Valley, based upon a collection of Bolander. 14. Crepis barbigera Leiberg ex Coville. Bearded Hawksbeard. Fig. 6062. Crepis barbigera Leiberg ex Coville, ae U.S. Nat. Herb. 3: 565. 1896, Perennial, stems 1 to several, 2-8 dm. high, from a swollen, simple or 1-2-forked caudex and rather yomerg woody root, mostly branched above the middle, the herbage ae to a soately ee stout, often with yellow or greenish, nonglandular setae. Basal lea 1-4 a lon pinn bipinnately toot q or parted, the lobes acute or acuminate, tapering Tbe low 7 yas. narrow, pind petiole, the cauline leaves few, similar, encarta length, the estnoet pas rag : infloresc of about 7-30 heads, the heads 8-25-flowered ; jeavabier re cylindric, with spreading veclicuiel seta ue oapecqgar ge ge phyllaries say ee - lon ng, Lissette acute, the outer less than e length of the inner, similar outline; achenes 6.5-10 m ae a gly ri , oblon: a with a broad soa olive- -green : a i sHightly — hats ‘the Dry open places, Arid Transition Zone; eastern Washington east to northern Idaho and — to ee County, eastern Oregon. Type locality: Alkal i Lake, Douglas County, Washington. June—July. SUNFLOWER FAMILY bt?” tev GEC . Crepis pleurocarpa . Crepis stan: 6062. Crepis barbigero 6063. Crepis runcinata 609 610 COMPOSITAE 15. Crepis runcinata subsp. hispidulésa Fig, Coed Babe. & Stebbins. Meadow Hawksbeard. Crepis ial apt bree Pittonia 3: 27. 1896 tspidulosa avant x Rydb., — N.Y. Bot. Gard. 1: 461. 1900. leolat e, Leaflets Bot. Obs. 2: G venie pallens eeoeag on: cit. Crepis runcinata subsp. hispidulosa (Howell) Babe. & Stebbins, Carnegie Inst. Wash. Pub. No. 504: 96. 1938, Perennial, stems 1-3 from a swollen, simple or 1-forked, thickened caudex and a fleshy root, usually 2.5-5 dm. high, glabrous or glandular-hispid. Basal leaves obovate, rounded at the apex, 6-25 cm. long, cm. wide, the margin toothed toward the base, dentate above; cauline leaves redu ved o bracts; inflorescences of 10-30 heads, the heads 20-50-flowered; involucre 8-12 mm. high, campantlate phyllaries linear-lanceolate, acute, more or less glandular- hispid, the outer few and short, much shorter than the yee ee .5=5 mm. long, light to dark brown, mostly 10-tibbed, narrowed but not eernrs. ait us 4-8 mm. lon regpey moist alkaline mea Arid Trans sata tibem ses pee and Oregon in the Svat Stat also Montana, Idaho, Gis. so northern Utah. Type locality: base of Steen Mountains, Harn County Oregon. June—Jul is runcinata tehap . imbricata Babe. & Sethian, Kosuegie Hr Wash. Pub. No. 504: 102. 1938. rea Oregon soc to ilk and Rad Canuaea Nevada. Type locality: Alvor d Valle ey, Moree antes regon is runcinata subsp. andersonii (A. Gray) Babc. & Stebbins, Carnegie Inst. Wash. Pub. No. 504: 104. 1938. repis andersonti A. Gray, Proc. Amer. Acad. 6: Joa: reo — Dearie 6-20 heads; basal leaves ke imbricata; phyllaries attenuate at the apex, the inner 13-2 ; achenes 6—8 mm. long, more oa tebe strongly beake d. Alkaline ee Sierra County, California, pe iH aoe Ormsby, and Esmeralda gee Bhd ae Type locality: Carson City, Ormsby County, Nevada. nata subsp. hallii Babe. & Stebbins, pain, Inst. Mera ib. No. es 6.5 leaves pee ca eads of yellow Beers "alaaty at the ends ed holl In- volucre shone. or campanulate, its inner phyllaries in 1 series, slightly united at base and often corniculat h , the r in several series of shorter, somewhat erect, reflexed or spreading ones. Recep at, naked; ligules yellow (in ours), truncate, 5-tooth t the mit. Anthers sagittate at the base; style-branches slender, obtuse. Achenes oblong or linear-fusiform, -angled, 5-10-nerved, roughened or spinulose at least above, tapering into a slender beak. Pa appus of many unequal, filiform, persistent eae [Name from the Greek, meaning to stir up, alluding to medical virt enetically complex genus of f= erhaps 50 or 60 species, though many more have been descr ibed; occurring in << = =a New Worlds and in both the northern and southern hemispheres. Type species, Taraxacum offi- cinale Only Tinnited synonymy is shown for the taxa given below. For a more — oe of the taxa described see: a iorete der Gattung Taraxacum, Handel-Mazzetti, 190 7; North - n Species of Taraxacum, Sherff, Bot. Gaz. 70: 329-359, 1920; and various papers by Haglund and Dahlste Introduced, widespread, weedy species; outer phyllaries reflexed to spreading (often less so in T. laevigatum). Leaves usually not deeply grimconley the terminal lobe larger than the lateral ones; inner peak scarcely or not at all corniculate. 1. T. officin Leaves usually deeply fy cornicalate the terminal lobe scarcely or not larger than the ere pat inne phyl- y cornicu. aevigatu Nati e M4 Fe EK | a Svleenes brown, olive-brown, or stramin neous en ripe, obscurely quadrangular Inner phyllaries mostly peeelate; plants circumboreal in mountains of ‘Washington Oregon, and prob: ably northern Californ eo ceratophorum. Inner phyllaries seldom if at a corniculate; plants of southern California 5 aah : 4. T. californicum. Achenes red or reddish brown when ripe, mostly sharply quadrangular. 5. T. eriophorum. 1. Taraxacum officinale Weber. Common Dandelion. Fig. 6064. eontod. L. Sp. Pl. 798. 1753 ah. — Leontedo fee Lam. FI. Franc. 2: 113. 1778. } acum officinale Weber in Wiggers, Prim. Fl, Holst. 56, 1780. ey psd cadeee dens-leonis Desf. Fl. Atlantica 2: 228. 1799. Taraxacum palustre var. vulgare Fernald, Rhodora 35: 380. 1933. thick and deep. Leaves oblong to spatulate in outline, pinnatifid, sinuate-dentate ot arid nearly entire, usually with a large terminal , 1-3 dm. long, 2-5 cm. wide, glabrous or ent ; scapes usually several, 540 cm. high ; heads solitary, .2.5-5 cm. broad, with numerous Coton flowers ; i inner pis ttasiee es erect, linear to linear-lanceolate, maney way fasion ; outer phyl- in auihe but shorter and refle xed ; sates tee ed cee rown, - orm, spinulose SUNFLOWER FAMILY 611 Fields, roadsides, and lawns; naturalized from Europe; Alaska to southern Coitornts. well established in the Pacific States as well as in the eastern states. Native of the Old World. Ma: ay—Oct 2. Taraxacum peak tien (Willd.) DC. Red-seeded Dandelion. Fig. 6065. rae neste laevigatus Willd. Sp. Pl. 3: 1546. raxacum laevigatum DC. Cat. Hort. t. Monsp. 149. 18 axracum erythrospermum Andrz. ex Bess. Enum. Ag neat 75. 1822; and general habit similar to the prec ceding species. Leaves deeply Lap a Speers usually long- agabints , the ° 5 ¥4 | =] ro) oo o Se I O- o.. 3 = Bo o 328. ae ~ oo oO S = 4 et © +s f=} 9 a = > oO i *'3 > BR oO. s ro : many-flowered, pas dosh roa a involucral phyllaries glaucous, spe outer spreadin ng or somewhat reflexed, wy ay at attenuate, the inner linear, 7-10 mm. long, erect, each usually corniculate just below the tip; flowers sulphur-yellow or the outer usually mec dor He achenes red or red- dish brown, rather te ribbed msi Jokela omen spinulose above, the beak slender, not over twice the leng th of the body; pappus 4-7 mm. long, rather dull white Fields and dooryards; naturalized aa Europe ra = common in the Pacific States than T. officinale; common hiouphase ‘the United States. Ma ay—Oct 3. Taraxacum ceratOphorum (Ledeb.) DC. Horned Dandelion. Fig. 6066. pete a fee aoa Ledeb. Ic. Pl. Ross. 1: 9. pl. 34. 1829. Taraxacum ceratophorum DC, Prod. 7: 146. 1838. halos oe M. E. Peck, Madrojfio 5: 247. 1940. Roo k, blackish on the surface. Leaves procumbent, ascending, sf —_ ene or ate * dents te or often retrorsely psa or lobed, glabrous ; scapes s long as the leaves (short r in flower), w fs exceeding them uit; outer p hyllaries erett ceoeate rnaclate to ovate, glabrous or villous; inner ipliylkaties tinenY lanieolae te, 10-12 mm. long, dei distinctly weed ib cok at apex, dare f oh ec scarious-margined ; rays s well exserted, purple-veined ; achenes stra us or brown, pr rominenty muricate above the middle, the beak as long as or npsepars ing the Bode: aes hits, 5-6 mm. lon and meadows, Seceet stag circumboreal southward in the mountains from Alaska to northern Cali Banks fornia; also Rocky Mountains and eastern United States. Type locality: Kamchatka. June—Oct. Taraxacum lyratum ine aha DEC, er 7: 148. 1838. (Leontodon lyratus Ledeb. Fl. Alt. 4: 152, 1833.) e her nd “wt OL \ 6064, T. pie ae ee 6065. Taraxacum laevigatum 612 COMPOSITAE 4. Taraxacum californicum Munz & Jtn. California Dandelion. Fig. 6067. Taraxacum californicum Munz. & Jtn. Bull. etalon — $22/227.. 1925. Taraxacum ce mene nerne var. peices n. Fl. Pl. Calif. 1004, 1925. Root thick and dark, the crown sim Lil or sometimes divided into 2 short Crown, the herbage glabrous. pee asce ending o or widely spreading, bine mero , 5-12 cm. long, 1-3 cm. wide, obtuse to acutish, subentire to sinuate, dentate, rarely i aa incised, ls vhs ous and maid green; scape solitary to several, ascending, longer than the Pilea in fruit, labrou ; involucres broadly cylindric, high, nearly truncate at base; inner phyllaries ae “or corniculate ly obscurely so; outer phyllaries ovate-lanceolate, barely half as long as the inner, glab flow ellow, erect or nearly so, 2-3 mm. longer than the involucre; achenes light brown, abou mm. pene c rugose below, dial ie above, abruptly attenuate into the beak, this 7-9 mm. lon pappus white m. lon Moi ey Arid Transition it Canadian Zones; San Bernardino Mountains, i Bernardino County, ieitliets "California: Type locality: Bear Valley, San Bernardino Mountains Jun e—Aug 5. Taraxacum acai Rydb. Rocky Mountain Dandelion. Fig. 6068. Taraxacum ores Rydb. Mem Bot. Gard. 1: 454. 1900. Taraxacum olympicum G. N, Jones, 2. Wash. Pub. Bot. 5: 263. 1936. Perennial, saras esembling T. ceratophorum in growt a habit, and leaves; inner phyllaries usually not ociéenlon; outer phyllaries and summit of scape with some woolly hairs; achenes red bs apo brown at maturity, sharply quadrangular Boreal Zone; Alaska, the Olympic Mountains in Washington, Montana, and Wyoming. Type k Secalay: matey eg Madison County, Montana. May—Sep 176. LAPRANA I, Sp-Pi, Sl, 1753, Ann Ssacaise Involucre ra ie the bracts erect in 1 series ses few minute outer ones at base. Corolla yellow. Achenes oblong-obovoid, rounded at x, 20-30-nerved. Pappu a Receptacle naked. ncient Greek name of obscure aera on. |] n Old World genus of 9 species. Type species, Lapsana communis L 1. Lapsana comminis L. Nipplewort. Fig. 6069. Lapsana communis L, Sp. Pl. 811. 1753. Ste — below, ie sag — from the annual root, branching aceon above, 3-10 dm high, aiake s above, m r less villous-hirsute below and sometim s glandular. Lower leaves ovate or deltoid. ovate, 3-5 ¢ cm. lene ‘a. pubescent or glabrate ; ek cles often as long as or longer NYS a, hig ei =< bpd’. 6067. Taraxacum californicum SUNFLOWER FAMILY 613 ee the blade, often with 2 to several lateral lobes ; rap leaves often deltoid or agonal eres h shorter winged petioles or nearly sessile; hea ads slender peduncles, usually n ous ; in- be rang cyli oa ric, 4-6 oe . hi one with about 8 aripcival clotaiies bia a ae all outer ones at base, these erect in fru ostly in ane sean stations Lehane wed ee eae Ee — og dl bac grado sey — of the Cascade Mou to Oregon and south a the coast in California to San o County; also the eastern United States. Wate “rom Eurasia. i ud or . shoheaaites axim. Bull. Acad. St. Pétersb. 18: 288. 1873. A eeerens. poreregt stemmed annual Gay of the mature heads divaricately spreading. A native of Japan, grow a ground cover on se berg Mataua County, Orego: ee = Wages \h Ae 6068 6068. Taraxacum eriophorum 6069. Lap nis xy aa P ti? cy 4 z j* s Ks ? » ra bag 3 ‘ ‘ : 2s + ‘ + ¥jpit 4 SL3thav $ pe Be | i eee Sar DV : ’ 7 a i { z Ee “ iw £ 7 a eee ee & 5 axe Sup ee SF Gist ee Rees Sree ; - ; eeie act ‘ fare , : ni ‘ ’ Hal Ae. i yee : j u E "ad “ 4 ; % = VES ’ #4 z { E 2 7 ie ba ire aiahes = Prenat Sekt ornia; flowers — in globose c 120. Loganiaceae 3: 350. Herbs a if — Og fen bce Turricula), not as in the precedi ‘Sat oe. wen present opposite (except Ver- um Capsule 1-celled; stamens 4 or 5. 121. Gentianaceae 3: 350. Capsule 2-celled; stamens 2 or 4 (5 in Ver- bascum). 136. oe 3: 686. Stem-leaves al — aka — the base, the sepals distinct. APPENDIX 623 Flowers mostly solitary, sessile or pedi- — te IM tue tear-advlio, Margin Of rollas aoe or nearly so (ex- ae t Cre. 126. Pcietrulaceee 3: Inflorescence cymose, the pie usu- ally coiled; corolla s lobed. 131 ydr ge bo eae 3 Calyx popermees or shor iy ibe agente so in Fruit a many-seeded berry. 135. Solanaceae 3: 662 Stamens at least twice as many as the corolla- lobes (petals united only at base in some genera).* Anthers opening by ongituctinal slits; spiny shrubs of California deserts. 129. Fouquieriaceae 3: 474. Anthers opening by pores or tubes; gone not of the desert. erbs; bas corolla-lobes united only at the 112. Pyrolaceae 3: 287. Mostly shrubs or trees; ie caisraaulen or urceolate (lobes nearly free in Clado- thamnus, Ledum). 4. Ericaceae 3: 297 Corolla irregular, bilabiate or spurred, with lobes obviously unequal.t Fruit 10-25 cm. long. capsule ee membranous. 137. Bignoniaceae 4 Herbs, introduced, or ‘indi igenou the desert; capsule woody beneath the deciduous exocarp, the persistent style ailing pr 2 curved horns. 138. Mar astyainces F t than 2 cm. long. Plants th with dissected leaves bearing minute bladders (except ae one re Pres bogs, with tire ~~ leaves) 140. Lentibulariaceae 4: 1 Plants ad aqua’ Fruit a conse abruptly narrowed basally and be lgare glare Abe ir elastically dehiscing; t 13. Fruit if a t dehi i =) nor riarrowe cy Fruit of. 2-4 nutlets; wail: genera wit matic herbage. Ovary 2-celled with 2 nutlets; oi cone (Phyla, Aloysia). 133. Verbenaceae 3: 609 Ovary —— and -lobed with 4 nutlets; style eric 3 Pies depresnions between the ovary lobes 34. Men e (Labiatae) 3: 614. Fruit a several- vs many- seeded capsule, opening by a or pace 136. Scrophulariaceae 3: 686. Ovary = aig (partially so in Styracaceae). Stamens 10; errer: shrubs ae nut- Seay 1-seeded fruits. 118. Styracaceae 3: 345. Siasiens 2-5; if shrubs the fruit other Stamen res ‘ead S opposite Mistake in Campanuloideae). tipules always present (leaf-like in Galium, the saves Ye me segretins whorled). ubiaceae 4: 22. Stipules mig (except Sambucus in Caprifoliaceae). Fru ry, gh or often modified achene (dry, 3-celled, 1-seeded fruit in Linnaea). rubs or woody es joes a creeping ga Pee en her - baectiallaians 4: 42. Perennial or annual herb: Flowers irregular, Pit tube usually spurred 3 gibbous. 145. Valerianaceae 4: 56. 6. Di ipsacaceae 4: Fruit a capsule, dehiscing by valve-like openings “ _cpliting between a sibs (Campanu- loideae). . Campanulaceae 4: 7. Stamens with. either filaments dus i or both parts ieee or partially fused; eae alternate letely basal in eee itae). Individual towers small, a into a h volucre; the whole appearing flower- ike * salt oe 149. Campesitact 4:98. sect ae in heads not aggregated as in the sarees penete prese: uicy berry; stamens opening by pore Gaultheri ith ogee f calyx). Soa ciaiacies e326, Fruit capsular or a dry berry (pepo) either indehiscent or rupturing auksee stamens ope Ti 1 & . —_ aoe subtended by a calyx-like ning by slits. Flowers monoecious or dioecious, regular ape ae’ parted in Marah); vines with ten . Cucurbitaceae 4: 65. Flowers pittect: irregular; herbs ida. 148. Campanulaceae 4: 72. * See Choripetalae for genera with petals cohering at the base in Crassulaceae, Oxalidaceae, Mimosaceae, Malvaceae. + See Choripetalae for genera with petals partly united in Fumariaceae, Fabaceae, Polygalaceae, Krameria- ceae. t Marginal flowers of the heads often with colored rays, as in sunflower and daisy, or all flowers of the heads bearing such appendages, as in da ndelion, or all flowers without such appendages, as in thistle; corolla vestigial in e Ambrosiinae). ; Sebi : <= : 3 ‘ Cee at ¥ : ah $f aoe Nie Hl ETA | ’ tee © ae ere TS oe Per Tartan : i ie nu » | 7 pirat tes Ges ae i US ene SE OSs, Gin & | | ie | : | | Ss ee ada io eS ahead J : * Fens * | | * ’ iD rs ‘ : Lal ‘ ie | * Sb oe * ri : os AS Ochs a ess oa y angrier + Pe e Sgt ew ao ea {82 ‘ Seale $ eter rias my Abrojo California, ITI, 61 Gray, III, 61 Spiny, III, 61 Abronia Alpine, u Sir Beach, I Co ville’ s wal li, -1d a Sa a 8 Pe oney- ‘scented II, 108 Mojave, II, 11 Red, II, 1 10 Transmontane, II, 111 ee Dwarf, III, Acacia, oe a sb 475 Acaena, Califor I Acalypha, California, Tfi,.2 Pisce FAMILY, TVj44 eo s- aeage: Fam Adder’ s ah I,2 “California, ns Pia neg ae Adelphia, aacads: pe —— III, 6 Small-leaved, mt 626 1, 1V, 566 arge- flowered: IV, 565 Orange-flowered, Seaside, IV, 566 Short-beaked, IV 562 re chee TV, 565 Tall, lV, 564 Agrimony, Tall Hairy, Tf; Ajamete, III, 378 Alder Red, i Ree Sitk: de I, 514 Wavy-leaved, I, 513 peeiaits a 541 All-heal, IV, 56 Allon, “+d 104 locarya Adobe, ITI, 561 —<— IT, 565 Slender, III, 567 oth-stemmed, 62 Stipitate, a 563 All-se I, 86 Aloysia, “Wei S; III, 6 Alum er II, 381 AMARANTH FAMILY, as seh Amaranth Catarsta: II, 99 Fringed, II, 99 Green, II, 98 a II,9 AMARYLLIS eases I, 459 Amblyopappus, IV, 248 Ammannia, Long leaved, II, 165 Ammi, Toothpick, III, 230 le, = eee I, 414 Amsinckia Doula’, oe 604 — ini III, 606 “Shor. caerad: re 367 Woolly, III, 3 y, ‘hechuae kau 541 genera in Roman; sac California, III, 337 m, 311.337 II, 337 omens flowered, I, 410 Ane Desert, II, 19 mmond’s, “ee 191 ieee II, 191 Northern, II, 191 Small- flowered, II, 191 Western Wood, 191 III, 378 Pil AMILY, II, 469 Cside: i225 Divaricate, I, 125 Purple, I, Reverchon’s, E427 Schiede’s, I, 125 ede’ oe I, 427 epedtiedka: IV, 420 Heart-leaved, IV, 424 i IV, 422 INDEX OF COMMON NAMES [Families in SMALL CAPITALS; Roman numerals refer to volume numbers.] Arrow-grass Mar. rp I, oe Seaside, I, 9 ames ribbed I, 96 Arrow-hea Aru Suaved I, 101 Broad-leaved, I, 101 Arrow- leaf, Iv, 498 Arrow-weed, IV, 466 Arti ches ke Ga rden, IV, 513 Pe M FamIcty, I, 345 Apisonds 118 347 California Flowering, Dwarf, III, 347 Fringe-flowered, Foothill, III, 347 Oregon, III, 346 -IV,3 reese ccd iV, 3 — California, hac IV, 341 6 32 326 En nti leaved, EV 313 Geyer’s, IV, 316 Gorman’s, IV, 3 Great Nor ie: IV, 314 Greata’ . IV, 316 Hall’s, , 319 Heath- ee L¥vS22" , 1V, 334 Kicks IV, 326 IV 333 a 1 , 328 Leafy, IV, 318 Little Gray, IV, 322 Long-leaved, IV, 320 Marsh, IV, 316 Rough- oe iV San Se ty TV, 322 Shasta, IV, 334 Showy, IV, 313 Siskiyou Rayless, V3 Athysanus, Dwarf, II, 298 Avens Bell-shaped, II, 449 Drummond’s Moun- tain, II, 44 rge-leaved, II, 447 1 Awlwort, Water, Pancha 57 eed, II, 6 Avena, Tit; ve Azalea, Western, oy "309 Azolla, Fern-like, I, 35 errd Blue-eyes, 482 Baby s —— IV, 32 Bace ~_ i , 384 Desert, IV, 384 Douglas’, 1V, 386 mory, IV Plummer, IV, 384 Salt Mars ; IV, 215 Small-rayed, IV, 216 Southern, IV, 218 Woolly, IV, 216 Dissected, IV, 221 INDEX OF COMMON NAMES New Mexico, IV, 222 Balm nae EEE se emon, IIT contain i 4 Balsam-apple, Wild IV Balsa Arrow eae pels 106 California, IV, sn arey’s, IV, Deltoid, IV, “9? Hairy, I ooker’s, IV, Serrated, IV, 110 Silvery, IV, 108 Woolly, IV, 109 Baneberry, Western Red, II, 178 Saweene FAMILy, II, 216 Barley Little, I, 245 the as 3 Bar satan lil, 73 Bartonia ie Iden III, Batis Bou II,.114 oth ating I, 58 BayBErRRY FAMILY, Bean caper, Rectan Ill, 7 Bearberry, Red, III, 312 Bear Brush, III, 287 Beard-tongue a, IV, "35 Matthews’, IV, 40 ey s, 1V,4 st ‘tae ne 34 Rothrock’s, IV, mover fruited ea Ply anta — IV, 34 9 uoia, IV, 37 ge branched, v3 bie Rough, i ot TV; 3 Wall IV, oT, 28 llow, IV, 32 Y olla Bolly, IV, 34 BeEEcH Famicy, I, 514 oade: fern (se Fern) Beet, Common, II, 68 cian be IV, 130 nia, Wild, I IT, 50 5 cpa TEE, 80 2 7 Bequilla, Ii, 562 Bergia, Texas, III, 119 riot Western Pos Cuteness II, 474 Oso, II, 469 Salmon, I I], 457 Thimble, II, 455 Besseya, Red, te 800 Big Tree, I, 6 Bic GNONIA ed 1 , 383 os H FaMiny; I, 510 Glandular, ia 511 Low, Western, I, 51 Paper, it sh oR a $11 a prunes, Ill Pointy III, 85 Pallid, III, 856 almate-bracted, Pringle’s, III, 859 Purple, Stiffly- branched, lil, 853 Tight- ——* HEIL, 853 Viscid, I 855 Ya kima, a 859 Birds- on-Nest, ITT, 855 ge WORT I, 534 Biscuit Root, III, a hn hop’s Cap, IT, 3 Bis mi ae 4 on — III, 230 a, Moj jave, IL 1 159 Bistort, Western, II, 62 135 ieee alec 00 ackbe yeeros! II, 458 Black-eyed Susan, IV, 117 BLADDER-NUT FaMILy, 1],.55 ss Double, TL 286 Oregon Double, II, BLADDERWORT FAMILY, IV, Bladderwort Mo untain, ie 12 Blip ing Heart Few-flowered, II, 234 Pacific, II, 234 Blepharipappus, ee 192 Blo §, EVs silat pits — IV,-72 Bluebell Common Alaska, soba III, 67 greene IV,8 Large- lowered, 80 eigen Blue arin Hernandes, HB om ou mn, Lit wered, = vod Francia eS Greene’s, II Large-fowered, Ill, ees iruited, Lf, 779 Mrs. Bruce’s, III, 779 Efi, Small- flowered, 778 pone s, III, 780 ade s III, 780 -flag, Western, Pacific, z 206 INDEX OF COMMON NAMES Piper’s I, 2 Pringle’s, . Ae San Bernardino, I, 201 0 ss oO al OQ, gs bo ¢ Blu Blue Witch, III, 680 Boe — Com mI, 1 rare Valley, sy 103 03 W Bog- eerie aie — i “a Boisduv pe een III, 180 ea: III, Large lowered, IlRA Narrow. caved 100 & Pale, i a ao ooth, Sy - andra gered Ik, 350 ornia, 1, 255 Bouncing Bet, II, 171 phar ne III, 21 Box — California, H1,5 Bas de III, 45 Boxwood, Oregon, III, 54 in, II, 3 Round-leaved, II, 352 Bracken, Western, wee Bract-scale, II, 82 Bramble Five-leaved Dwarf, Hairy-fruited Dwarf, i, Snow Dwarf, II, 455 ates IV, 502 California Spear- leaved, IV, 502 Desert, IV, 504 Greene’s, IV, 506 Gum-leaved, IV, 500 Knapp’s, IV, 501 Large-flowered, IV, 502 Little-leaved, IV, 504 w-leaved, IV, 504 Nevin’s, IV, 504 Shrubby, IV, 501 Spear- mai , IV, 502 Sweet, I Willow- leaved, Bristleweed Annual, IV, Goodding’s, IV, 271 ike, IV, 272 Rush-like, IV Brittlebush, IV, 123 , 409 Many- Sorenel. I, 409 ban Bro Hungarian, = 229 Brooklime, American, 1h? Brookweed, III, 344 room California, II, 552 ant Chaparral-, wy 386 French, IT, 519 an 9 Silky Californix, L Turpentine, III, 18 White-flowered, II, 519 BROOMRAPE FAMILY, 3 Brown Turbans, IV, oe Penny, it Bucks FAMILY, III, 3 Bucidean III, 366 Bu Common, III, Socsere: III, 66 627 BUCKEYE FAMILY, tf ee Buckeye, California, Ill, BucKTHORN FamILy, 5 WHEAT ’ FAMILY, ti a Buckwheat Desert, II, 39 Neale Si II, 49 Bis — ie sh “California, se agi G asi bees sae Utah, acd 350 B Sour, III, 16 esr fgg one 681 i ads II, 179 False, IT, seg Hood, II, 179 aie 756 eels ‘weed, Northern, III, 658 Bugloss, Small, or 541 Bugseed, 11, 90 “Searle III, 756 Utah, III, sic can en 4273 River Ambrosia leaved, ati: eedeit ae 40 San Die Woolly- taht Bur-chervil, HI, 224 Bur-hea 3 BurR-REED Pee thks I, 80 Bur-reed 628 [Bur-reed] Reoad: fruited, I, 81 Coe 1 ot Gre s, I, 82 = a leave d, I, 82 Simple- awa I, 82 Small Bush-cranbe exty; High, IV; Butter- yr Eggs, 817 iit, 7 Butter atte reed © 214 Bloomer’s, II, 203 Cooley’s, thie a 1, 66. 207 Cursed, II, 210 oe os 210 Desert, II, 211 Eschechol's, ET, 207 Field, II, 205 oe i IT, 208 oy 0 Tai nt 2 eeae caved II, 206 Low. 11,2 Lobb’s Water, fo 214 mate Prickle- elk ge 205 Pubescent-fruited, II, 206 Parsh’ efile Sacramento athe T, 200 hot oe Se 207 Small-flow red, Strai sr beaked, ieee: line, II. 207 Water, II, 214 Waterfall, T2101 ater Plantain, II, 208 Western, IT, 2 ntact ous a 211 Butterweed Alkali-marsh, - 438 pin eee kek IV, 444 Afoceet — 436 asin, Black-tipped, on ong Bloch INDEX OF COMMON NAMES Clark’s, IV, 4 Cut- leaved, a 450 ae Atay Age 444 Dwarf Mou LV. "434 Elmer’s, IV, o Ni Mountain-marsh, IV, 440 Mountain Meadow, IV, 436 Narroe: leaved, IV, 434 Olympic, IV, a Porter's, IV, 4 Puget, IV, on pe sh aa ibd Rayless Mountain, IV, 4 Rocky Mountain, IV, 4 St Bernardino, ks ae Obispo, m Sweet “marsh, IV, 438 Tall, FV; Tehachapi. ca 442 Woolly, IV, 442 aes Common, ai Button-bush, IV, 24 oe bl 274 Squ 1,255 Wild. nfs Slender, Th, 255 Yellow Skunk, I, 346 Reker i ILY, Ill, cae s Beaver-tail, es 148 Foxtail, Ill, izzly Bear, III, 149 Pincushion, III, 162 Saints’, San Diego Barrel, Sp 156 meth IIT, 162 Seaside, os 122 Calla alltanda, “Hairy ved, I Calon es 135 Cal Common, I, 399 Sierra, I, 400 CALTROP als III, 14 Calton eM 111,46 Calyc But Oe si “19 . Dwarf, IV, 1 Freniout, iv. 190 alyptridium Common, II, 13 Four-petaled, II, 131 Party's, 11,13 Rosy, II, Camass ommon, I, 415 Cusick’s, I, 415 Death, I Rose, II, Scouler’s, II, 166 Seely’s, II, 1 Canaigre, II, 50 Canbya an White, II, 232 Ca rete ile, FET: 234 False, III, 233 III, 115 Carpet-weed, III, 115 Carrot FamIty, dia | abe Carrot, Wild, III, 228 Carved-seed, IV, 582 Large-flowered, 582 ae Sl sap: a Armed, II, 479 Castor Bean. U1, 30 Casto pp Plan IIT, 2 Cate fly Common, II, Many- dae 4 160 pes ye wering ees rooted, Sad 586 Sm hme Yellow oor Car TAIL Bice. 1,79 Cat-tail . leaved, I, 80 eee w-leaved, I, 80 eh “Calor, 113254 eo ping-leaved, 253 et ade II, 254 Glaucous, II, 257 hie aly podded, ees oe Cean 5c ‘ode, one 73 ,76 Greenchavied, LE 71 Hairy, TI, 67 et Poe LEE, :75 Island, III, 73 Wavy-le ved, III, 70 Woolly-leaved, III, 67 Ceda > TIT wane Weshilagtse: III, 352 Cereus, Golden-spined, I 153 Chae iiesee: leaved, IV, 242 Branching, eae 238 oad-flow Common Yellow, IV, 244 Cusick’s, IV, ~ V2 V, 240 Inner Coast Range, John ey wh 241 Large-flowered, , 242 Nevins’, IV, 242 Northern Sierra, , 238 Parish’ s1V;2 Sentelics. TV 2237 Serpe , 1V, 244 Shasta, Southern Sierra, V523 Thompson’s, IV, 238 Xantus’, IV, Chaetadelpha, IV, 568 Winonied IV, 268 INDEX OF COMMON NAMES Chaff-bush, IV, 262 Chaffweed, III, 334 S 2 ebatia, Southern, I, 45 Cae et f, III, 668 amise roesow on, II, 4 Thin- leaved. “ aa Chamomile Downy: sheathed, 236 suey. 232 Sm og flowered, Che iis Bloom, a Aig Cheese-weed, III, oke, IT, Holly leave dit, 467 Tudias Ill, 115 pc an II, 143 r Mouse-ear, hi ry, III; 20 Chinc Chinese Case: III, 34 ee Houses Downy-leaved, TIE: O75 Purple-and-white, Lit 775 Round-headed, I Southern, TL, 775 Chloris, Silky, I, 176 Chlorocrambe, II, 242 Choll e eee 1; Parr Bs Pert olitte: i 15 Pink, II, 1 Prosirate it 5 a 116 t IL-1} an Francis co, II, 6 pees II, 11 Cor tie eter IV, Chuckwalla’s aoa Chuperosa, IV, 14 II, 43 Viffuse, II, 429 tecnico leaved, II, 434 et II, 438 TI 439 > ionnanw omental... ™ 8 ap 5 8 es) & 3 boon onl > w o -lamath, II, 435 629 Many-leaved, II, 435 Marsh, II, 4 Mount Rainier, oe 434 Newberry’s, II, 430 Short- leaved, IT, 435 Shrubby, II, 438 Rage Mater Slender, i. ie k Coenen: ITI, 298 Clam eed, Western, 11, 327 Clarkia a III, 184 We ir driv 1, 123 Cleavers, IV, 2 Clematis Chaparral, II, 194 Columbia, IT, 194 Small- feaved, IT, 194 Cleome Br er eatk a 3° ee Short- pedicelled, Small-flowered: II, 326 Su rf’s, I, 30 Chiff Bush, IT, 387 Clintonia Red, I, 453 Single-flowered, I, 452 Clotbur, Spiny, IV, 153 to to net RaFe Es WwW Ww Pad ° g — a - 3owl, II, 630 [Clover yer] Branched Indian, TES Brewer’s, II, 524 hy tT Ga aero shi Clover, 8 BA Ff) Gray’s; II, 533 Shag: Il, 526 a — a ata Large- edhe IL $27 onterey, ” 534, 535 Mountain, II, 534 Mustang, i 42 —— leaved Sack, i; 5 Notch: leaved, II, 523 tancheria, IT, 531 ted, IT, 530 hasta, IT, 526 howy Indian, IT, 531 ymall-headed, II, 533 2 ta FAMILY, I, 43 Clu b-m Alaskan, 1,45 Bog Fir, oe 5 Pockic TI, 171 Corn, II, 159 Seared IV, 152 Soke tecas — 63 California, II Sierra, 111 III, 62 Co Nuthalls: IIT, 534 INDEX OF COMMON NAMES Palmer’s, IIT, 534 Plicate, III, 534 Shr Hert III, 533 Collom piewne ITI, 405 Bristle-flowered Diffuse, III, 403 gi flowered, III Pret ITI, 406 nat row-leav ed, I, 403 Rh Delt? 405 Talus, ITI, 4 Coltsfoo Arrowhead, ~ 456 eet, I Colubrina, behevviis! oe ae sie II, 180 Northwest Crimson, Hey Van aris s, II, 180 Yellow, II, 179 Co dra Pale, I, 528 omarostaphylis, Cali- fornia, III, 308 Comfrey, Rough, , 540 Conyza Coulter’s, IV, 382 South American, PV 4 Coral-r Bieelow's, 1, 484 1 Leafy-stemmed, TV. 432 Mount vgragatn iV Silman’ ae: 134 Coreth Ca “ak, IV, 338 Common, IV, 340 San — ardino, Vv; Coriander Hp2 Cornel, Dwa rf: 4 284 m orn Salad, IV, 61 Coronilla, I], 614 rydalis Costmary, Iv, 396 Cotton, ITI, tint Pian, IV, 469 Cotula, ‘AGettalian: Cous, ik2 Cowania, ctindinete II, 450 Cow-bane, Western, III, 245 Cow-herb, IT, 171 Ss A eat, American, III, 8 Coyote -brush, IV, 386 e Me lon, IV, 66 282 Crab- soot, Giaewe Cra Pela Lisropean: Ill, 329 —— Bush, III, 16 Cre Field, II per ee pied Hoary, ies 258 Pg IIL, 389 Crocke a IV, 22 Crossosoma FAMILY, Crossosom Bigelow’s, < fi. 406 g Catalina, II, 406 i ton, Califor rnia, me Bsc BERRY FAMILY, III, 4 gobi Black, III, 4 ccbeieils FAMILY, II, 174 Crowfoot Bristly, II, 2 Somes 11 wt 207 211 Crownbe: IV, incendie IV, 126 va wn-sca 1,79 cillo Calor, III, 61 a Spiny, II Crypis, ise leaved, T, 142 Cryptantha Bedked, IIT, 595 Bearded, III, 585 — ger ITI, 583 Bushy, III, 580 ree ter, ITI, 591 ee III, 594 Coas , 59 Co mat fo aie III, 585 Colusa, III, 595 : a a) Bs No bo np — to wm oe) 25 ceed like, I, 257 pity FamIty, 1, 70 as I, 74 ae ‘aie 117 Annual, IV, 3 he Alpine, IV, 3 Alpine nt a IV, 354 ieee ca hes Basin Rayless, IV, 360 Bloomer’s, IV, 3 Branching, IV, 373 Brewer's, IV, 368 ee — Rayless, pomtek of 352 V,3 Eastwood's, IV, 348 INDEX OF COMMON NAMES Eaton’s, IV, 366 Engelmann’s, ‘358 ig ry Fals Fle exudus, iV. 366 Foothill, [V, 364 G sty: erg: 2% Iv, 122 Paris Sane bg YY, it 67 Philadelphia, IV, 349 Piper’s, IV, 362 Supple, IV, 3 Tigi, IV, ab Three-vei ea Utah, 1V 3 Volcanic, ie 362 allace’s Woolly, IV, Wandering, IV, 346 ¥ er Ca bbage, eee 366 Deer eee As 362 Jeer Weed, I Jesert pede II, 466 , 460 Jesert Gold, ITI, 419 Jesert Holly, II, 84 Jesert-hollyh I I I I De Desert Candle, Des 4 I I I ock, III, 88 Desert-mallow, III, 88 Coulter’s, III, 87 Currant- caved, III, 90 Emory’s; ITI, 8 III, 380 Dita California, III, 26 Common, III, 2 Glandular, III, 2 25 ow-leaved, III, 25 heii III, 26 Yellow, ‘IL, 51 631 Dopper FaMILy, ITI, 390 Dodder ; I, 395 “it e, III, 39 Es say ue Soh Wester Westees' Fel ae 391 DocBANE FAMILY, III, 367 Clasping-leaved, Spreading, III, 369 Western, III, 369 Dog easel IV, 388 Docwoop FAMILY III, 283 Do ood os American, III, 283 Blackfruit, III, ee r III, 283 Ii, 2 Western Red, ne 284 417 Double-horned, IV, 90 S rats ft IV, 92 sreat Basin, IV, 94 oover’s, IV, Maroon- spotted, IV, 90 pares IV,9 outh American, 1,9 nuunsZ TOs ae ca ats II, 295 41,297 Douglas’, II, 297 ptr II, 295 Southern Californix, Volcste. IT, 295 632 [Draba] White Mountain, it Yellowstone, IT, 295 ps II, Dro Shahin I, 145 DuCKWEE e’s, II, 337 Lance-leaved, II, 336 pig? said TL, 336 Lax Panama, ah ey wder Setchell’, a ae an’ a tirecches Dyer’s-weed, II, 328 Dyer’s Woad, IT, 271 yssodia Cooper’s, IV, 252 San Felipe, IV, 251 Thurber’s, IV, 252 Sa ee ii, 233 twoodia, IV, 138 o Echinopsilon, Hyssop- hse: II, 90 EL-GRASS FAMILY, 1,93 Eel- -grass et ees Egbertia, IV, 568 rah sly Il, 459 | Chaparossa, IV, 204 Elderbe at IV, 45 IV, 44 pen Red, IV, 46 44 Desert, IV, Mountain Red, IV, 46 Southwestern, IV, 44 PROORE INDEX OF COMMON NAMES ee PAS oe 466 Ei is I, 523 Elm, ri a 378 Elephant’s Head, Little, III, 807 Ele phant Tree, Small- leaved, III ns Spiny, rf 49 Ence pass EV, California, iV, ie Rayles: Virgin iver, wv, 124 EpHepra FamIity, I, 77 Ephedra ee I, 78 Green, I, 77 Nev dk ‘I, 78 Ries ce forked, I, 78 —— L: 187. Creeping, I, 186 Low, I, 188 2 ib exican, 87 Orcutt’s, I, 187 and, I, Vis cid, ina! Eremocarya, - 578 “Abrans III, 438 randegee’s, ‘TI, 438 Desert ITI, 435 e, III, 434 asl Swerve — 437 Hoover’s, III, 4 Ericameria Boundary, IV, 288 Cuneate-leaved, ‘. Eastwood’s, IV, 286 Palmer’s, IV, 287 Parish’s, IV, 290 eer oe: Awned, jag a Eri Angle: “stemmed, es 29 t.41,4 es ’ Douglas’, II, 23 e, II, 47 Biscant I sae Es: meralda, 1h, 3} Flat-crown, II, 34 Flat-top, II, 42 a ee Hatiar ee flowered, II, 43 Onion- pein ed, Il. 24 Ord’s, 30 isi cae a 43 ish’s, 41, 3 Papi ty Fonte Pine, II, 38 -iper’s, IT, 2 roliferous fe 43 Pu "rola leaved, ns 24 Race § lender Woolly, II, 36 Snow, Spurry, II, 29 sulphur-flowered, 2 IV, 231 Jepson’s, IV, 226 Nevin’s, IV, wn Pringle’s, IV, 2 White Woolly, Vv, 230 W V, 230 ieee IV, 228 79 o, III gti iti ‘leaved, III, 281 cde TH, 28 1 Prickly, III, 27 eae saa en 2 79 y’s, III, 28 Escobilla, IV, a Escobita, III, 814 Eschscholzia Euploc III, 5 Bustoma, all, gia 355 Eva Trae. IV, 495 Erect, , 494 Involuerate, pal ASS EVENING-PR Cespitose, ITI, 195 Dandelion- je By 196 oe Sad, Kern County, Bh 205 Large Desert, III, 194 “el re wered, Woody. Pted vac Desert, ITI, vening Snow, III, 4 rlasting Isabella, eS ibe Lo M “adow a) 480 Narrow- leaved, IV, 478 e, IV, 46 White. margined, I Woolly, IV, 478 Fagonia, California, III, 16 Rough-fruited, I, 456 Fairy Bells Farewell-to-spring, II, 187 Fawn Lily Alpine, I, 428 California, I, 428 Giant, I, 42 Henderson’s, I, 429 Howell’s, I, 430 Lemon-colored, I, 429 Purple, I, 430 Sierra, I, 427 Small flowered, I, 427 Yellow, I, 4 oe Ill, ae Nn Famity, I, 5 aa Acpiateas Parsley, i, 22 Beech-, Brittle-, 4 a Cl oak- California, I, 33 ones’, I, 33 Gold- Licorice, 1,8 ip Cleveland’s, I, 29 tal, 12 Viscid, L, 26 INDEX OF COMMON NAMES Male-, I, 16 Oak-, I, 13 Shield- Cuaifotate k, 11 udley’s, I, 12 Lemmon’s, I, 10 Swor Tinbitcathde I, 10 Western, I, 10 Venus-hair, I, 24 Wood- Coastal, I, 16 Downy, I, 14 Fiddle-neck Large- flowered, III, 606 Rigid, III Se ouler’s, Bn 606 Small-flowered, Ii; : Tessellate, III, 604 White, III, 480 Fig hee pee a 119 Indian Bae Sea, III IGWORT Pete Hy, ” Figwo Black lowered, III, 7 oe tT 771 Hare, ITI, 771 Arizona, IV, 4 California, IV, sp arrow-leaved, IV, 490 Filaree Large-leaved, III, 5 Long-beaked, III, 7 Musk, III Red-stemmed, til 7 Texas, III, White stemmed, ae near tylis Grass- “Exe. I, 268 Big Cone, I, 64 Grand, I, Lovely, I, 65 Noble, I, 6 Red, I, 67 ew ‘Lucia, I, 68 2 Shas I, 67 Firecracker, Eaton’s, III, 758 Fire- cracker Flower, I, 41 Fireweed, III, 171 Cut-leaved Coset; IV, sr Eastern, IV, 460 Harves he Ayo earaniee 607 Toothed boas bh 460 Trumpe Fi Leica? ee Phe t, 111; 97 Flaming Trumpet, III, 4 Flat- bed II, 283 FLax FAMILy, tFF-9 i 1B & og wer’s Dwarf, IIT 14 Caliloruis Dwarf, Hit: yee trogde Dwarf, Fae Yellow, III, 12 Drymaria Dwarf, Dutch, IT, or False, II, 29 Gl cenbomare Oaee 14 Little Wi 13t, 43 Marin Dwarf, III, 14 arrow-leaved, III, 11 Northwestern Yellow, HE Slender Dwarf, III, 14 Small-flowered Dwarf, Ti, Small-fruit False, Western Blue, HE, 12 633 Fleabane, Salt-marsh, IV,4 leh 2 FLow salah: Quituwoer Fam Flowering ‘eater I, 95 Flower-of- -an-hour, ate iis d-leaiveld ITI, 786 Sharp. leaved, III, 786 Fluminea, I, 197 Fog- fruit, ITT, 612 w-leaved, 11, 613 Fogweed, II, 82 Footsteps-of-spring, I, 220 Forget- -me-not, IIT, 542 FouguteEriA FamIty, Ill, 474 Four-o’cLocK FAMILY, ed, IT, MacFarlane’, te 106 Foxglove, Purple, Ill, 792 18 RAN KENTA Poa. II, 119 — Palmer’s, ae , Fra Wii stemmed, iil, 362 Fr hatka, I, 4 Many flowered, z 424 deste <— Page Purp ts Poe ecaahia. I, 423 Scarlet, I, 422 Siskiyou, I, 424 ine-colored, I, 425 ellow, I, 426 7. View trae ITI, 612 634 — cae leaved, III, 169 hie ia, III, 1 Hoary California, ti, 1 Pied ad California, III, 170 FuMEwort FaMILy, T1233 Fumitory, ne VA f Furze, II, Gaillardia, Great- bees - IV, 206 Galins: Ciliate Tv, 136 Ss sae flowered, 137 Galium, Watson’s, y! Galleta, I, 109 James’, 1,1 109 Galvezia, Shaws. Gambelia, III, 783 Gaura California, III, 211 lagi Gayop ee yang III, ie ue 's; TZ 209 Much-branched, 31,2 Natal’ eee 208 TIAN FaMILy, , a0 Gentian Alpine, ITI, 356 ew Zealand, III, 3 INDEX OF COMMON NAMES Oregon, III, 4 KS TNTATA Ss — i =e) — ° — Se om PRA H o > —_ — B20 bo ia Abrams’, ITI, ite Argus ILL 4 Bell-flow ea mle 474 Blue Field, Ill, 463 Bridges’, ITI, 4 aah flowered, awe Br oad 9 Ea ITI, 462 California, III, 463 Ca away-leaved, III, 470 pert Jil, 472 Desert, “we Gache 455 Great Thing III, 471 Inyo, III, 474 Many-flowered, III, Many ea 465 Bd s, III, 462 Rock, III, 471 Spreading, II, 460 ar, III, 470 igelow' sII,91 rish’s, II, 91 san Diego, il, 94 a FL, a4 N oody, II, seen? ehnia, pildaioin III, 273 linus, III, 117 lobeflower, American, 176 Q a Sinna wee + oO ni Be ne és wn ° a) utheni Contidut, III, 45 Goat’s-beard, II, 413 Godetia Botta’s, ITI, Cylindrical, Hl, — Ts4 ea Sea oni tne III, 187 Witlea's het, III, 189 Golde Golden Line II, 2 Golden Death Valley, LY.,.t11 Nevada Showy, V,1 G Shockley, IV, in Alkali, IV, 218 Coast, IV, 214 Woolly, IV, 216 Gold-threa Idaho, II, 178 Oregon, II, 1 GOOSEBERRY FaMILy, 8 Gooseberry Alpine, II, 398 B ath, earaiaite. ah 401 Mount Adams, II, 399 Oak, II, Parish’ 5; II, 398 , II, 403 Seathern ‘California, II, 402 Stra ggly, II, 396 TE GoosEroot Famity, II, 6 Goosefoot California, II, 72 Coast, if, 71 Nettle-leaved, II, 70 Red, II, 71 Stinking, II, 70 Tasmanian, II, 72 Wall, II, 70 Gourp Famicy, IV, 65 Gou Vinee: leaved, IV, 6 eer leaved, IV, =e Gra ppetet! I, 178 Black, I, 179 Blue, I, 179 Needle, I, 177 Purple, I, 178 Rothrock’s; I, 178 Sand, 1,177 Side-oats, I, 177 GRAPE Sater III, 81 Gra California Wild, III Ze Desert III, 82 Aparejo, I, 139 sricn I, 17 Mar tite, I, 147 oe l; 107 eae rk vata howered, I, 149 e Mountain, 1, 22 Narrow. flowered, a leaved, I, 227 Orcutt’s, I, 231 e, 1,2 Suksdorts, . 229 kI2 Woodland I, 230 nch- e bo Colusa, I, 188 INDEX OF COMMON NAMES Cord- Alkali, I, California, Ea75 Cot a 10 Crested ‘nae s-tail, Satie 15.141 Deer-, I, 146 te) Lyme-, Sea, I, 248 anna- Davy’s, I, 212 *ew-flowered, I, 212 “owl, I, 213 Nit-, 1 156 Nut-, Yeliow, 259 Sores I, 168 _ 1172 erican, [ Geyer’s, I, 190 Showy, I, 191 Sierra, I, 192 Small, I 1 Orchard-, I, 195 Orcutt, I, 1 Awnless, I, 182 Pine-, I, 1 Tufted, I, 160 Prairie-, I all; 1,195 Rattlesnake, I, 233 eed- Bolander’s, Pig Brewer’s, I, oe & aa 159 alas go feprs I, 145 sine ll, I kite I, 249 Na rrow-spiked, I, 238 cific, I, 251 re Soaeigk I, 237 Yell eed godt III, 544 cribner, I, 239 — T, 186 Scouler ay = rrey’s, I, 9 635 Tickle-, I, 154 V Water Stare I, 349 V heat- Alpine, I, 243 ss Mcxidin, i; 1438 Smooth, 2 112 Western, I, 113 Wood, Siender, I, 148 Poa II, 89 Grease-bush Low, TIL 55 Nevada, ITI, 55 Greasewood; II, 94 Small- headed, IT, 445 Weste Grbehetilaieta, Li; 557 Grindelia Big, IV, 25 Great Vaiss, ae 258 San Francsico, IV, 257 sens IV, 257 well ga Pai 548 III Pahoa bk III, 329 636 Guelder-rose, IV, 47 Gum — t, Coastal, IV;.2 Gum- “weed, i 170 ar y at veueae III, 407 Small flowered, III, 407 Hackberry, Douglas’, Halimolobus, Diffuse, oe. Whitney, IV, 2 Hardtack, IT, 453 Harebel ees, te 78 Common Alaska , 74 Olympic, IV, 76 Parry’s Northern, H H il, I Harpagonclla, ifr, 609 Hasseanthus Blochmans, II, 342 Long-st — - 339 San Diece. if Hawkbit, Haig v. 588 Haw d Baker’s, IV, 606 Bearded, IV, 608 , range, IV, 600 Scouler’s, IV, 596 Shi 0 INDEX OF COMMON NAMES Common, IV, 285 Island, IV, 2 Hazelnut, California, fleal-all, ITI, 628 rt-nut, 11,628 Phils ns II, 8 par ios Hain a ne 297 _ a ge - 305 A Alkali, ITI, Heathberry, ey 46 Heather Brewer’s Mountain, E303 Canyon, II, 41 Mock, IV, 286 Pink Mountain, III, Purple Mountain, III, Wale Mountain, ITI, 305 Yellow Mountain, III, Hedgehog, Eight- spined, sa Hedge-hy Brabileas TIL, 731 Common American, TEE zat Hedge-nettle (see Nettle Hedge- parsley California, III, 222 Knotted, III, 224 , J apanese, III, 224 nois, Crete, ae Hedyp Bi Hed Mackense’s. Hy 614 Selec IV, 120 y Mountain, Hel Risdybed IIT, 534 7 Common, on 630 Herba IV, 489 Herb- or “Gilead, III, 626 Hermidium, Wing ed; Herniasia, Gray, II, 114 iron sR ra ITI, 529 Heu Alpi 3 asd 3 Cylindrical, II, 382 uran’s, II, 383 reen-flowered, II, 383 Island, I, = . Laguna, IT, Oval- ebay A 383 Pink, II, 38 Seaside, IT, 3 Urn- Aowered, 382 Hibiscus ees er 112 ock-, III, 1 a cba. del A ae IV, 38 Hoarhound, Common, IIT, 62 H me be An Small- leaved, II, 48 Hogweed, IV, 382 Hole-in-the- sand Plant, , 250 Hollisterin:. II, 2 Holozonia LIV, 157 HoNnEYSUCKLE FaMILy, IV, 42 Honeysuckle en) Bi 4 - bd wm V;55 Southern, IV, 54,55 ay, 5 Honey: sie Arizona, II, 100 Hookera ee E 406 arf, I, 405 NnNWOmY | all wn in rh 24 | £6 “> o & = > So a Hop- tree, Western ae Horke s, II, 418 presporlg II, 41 Carrot-leaved, II, 421 Cleveland’s, II, 418 Dense-flowered, II, 421 Howell’s, II, Leafy, II, 4 Parry’s, II, 41 Three- toothed, II, ee ea leave te Sai oy ns, IL 4 Wilder's, II, 418 Heekaepns FaMILy, Pe | Pralenece’ ET; Horse 00%, Ps pr 615 Hors ie Cat Little. oe fh aa 2 Horsehead, IV, 466 orse-mint Nettle- ange oe 624 Western, III, 6 Horseradish - en H AIL FAMILY, Horsetail California, I, 40 Con:mon, I, 39 Giant, I, 39 Marsh, I, 40 Sedge-like, I, 41 Swamp, I, 4 Horseweed, IV, 382 Hosacki Bentham’ a.t1, 551 opper- -flowered, I, 542 ecumbent, - 547 arge ne ee ere ed, II, 542 Narrow- leaved, II, 542 Soot I, 542 nete- —— - 541 fer e, II, ] ee TI, 543 ose-flowered, II, 541 Sulphur-flowered, ce un =] - 8 Toothed, II, 545 Torrey’s, II, 542 Gra d, 11, 5 Western, oy ae HUCKLEBERRY FAMILY, I, 326 Shot III, 3 Red- rayed, IV, 202 San Diego, IV, 203 Short- big bl 203 Hunger Wee 205 — hinsia, sss a 290 uk Lilac-flowered Wild, ie Water, I, 349 HypRANGEA FamILy, II, 384 Hydranthele, Western, FFL;-732 ‘ol iineneae Weiske 5; IV, 393 Hynienoxys, Arizona, IV, 207 Ice-pla rece III, 118 Slender-leaved, III, 118 Imperata, Hooker’s, 12 licheniae; FV.1 _—— Paint Gita ual pas lipped, III, 846 arge-flowered, II, 846 Small- flowered, I, 846 preenss ee oe Brewer Ewan’s, III, 837 INDEX OF COMMON NAMES Franciscan, III, 844 Oblong: leaved, oo 839 Obscure, III, 8 Ow nasi IIL, oe Pale, III, 828 Parrot’s Head, ITI, 825 oO QO o L al — — Oo Eee LT,3 Sho hela California. III, Pe hs a 832 Stiff Yellowish, Suk ane TIT, 835 Tiburon , 841 * Torch- like III, Wallow betes stalt III, 829 e, III, 828 833 hited’s White- ak SE 842 Wide-flowered, III 830 bare s, IIT, 836 lly, III, 842 Mettow-hiaived, ITI, 832 Indian Pink Cpewte oe vat INDIAN-PIPE FAMILY, III, 292 Le re Pipe, IIT, 2 dian Warrior a ane Me ndian’ s Dre Indigo-bus ig California II, 558 II, 557 Inkweed, II, 95 Iodine Bush, II, 91 Mii Weed, IT, 95 s Famity, I, 461 “Caifoena, I, 465 Siskiyou, I, 464 Slender- tubed, I, 465 Tough-leaved, I, 463 Western, I, 4 2 Ironwo Catalina, II, 409 Desert, II, 562 socoma Coastal, IV, 284 Pi IV, 284 tem — A, 422 Muir’ Pickering o Al 424 Plum Shockle “ is be tes Silver-haired, II, 423 Tahquitz, II, 423 Tweedy’s, II, 426 Webber’s, II, 426 Yosemite, IT, 424 Ivy German, IV, 453 Ground, III, 627 Kenilworth, III, 786 Jamesia, a pila IV, 198 Jep J Jewel Flo rae & ieee Mount “Diablo, ti 250 a IT, 250 JEWEL p FAMILY, Hy 59 Jewelaveed Spurless, III, 59 Western, III, 59 Jim-bru sh, bad vers! Jimson-weed, III, 683 pare 28 -up, IIT, Jo ius: "Tuck, III, 817 Nei We Weed, IV, 499 ua Tree, I, 446 Jase, Il, 478 "Caliorna, a 3 arf, oe Mountain, I, 76 Utah, Westeri. I, 76 637 Jupiter’s Beard, IV, 63 Kallstroemia, Cali- fornia, III, 17 Kalmia ee bs 302 rn Sw , 302 Kalmiopsis, Ill, — Kelloggia, IV, 2 iste Manic III, 112 a heirag Geran, ITI, Alpine, II, 62 Austin’s, IT, 58 Beach, II, 56 Bidwell’s, IT, 61 Bolander’s, Il, 61 iver sheathed, II, 56 spurry, II, 5 Swamp, II, ye Viviparous, HH, White- br phincety II, 59 Yard, II, 56 Yellow lowered ah 58 Knotwort Fam iT; at Kobresia, Bellard’s, [, 281 — pie 170 Koto, I, 377 KRAMERIA fe I, 627 Kruhsea, I, 458 Kuha, ITI, 139 ‘ae Tea, 298 Coastal, III, 2 Glandular, ae i Lac eflow ila aes Ladies’ Tre Hooded, I, "479 638 [La sneit Tresses] n, I, 479 Lady: se III, 68 Lady-fingers, Pink, 1a Lady’s- comb, III, 224 s Mantle, Lake-agoseris, ‘Alpine, IV, 553 TLamb’s Lettuce, IV, 60 Lamb’s Quarters, II, 70 Hanaupah, ye 200 nyo, IV, Narrow-le aad: Large-headed, IV, 200 Larch Alpine, I, 61 mooth, IV, 220 Yellow-rayed, IV, 220 Larkspur Anderson’s, IT, 189 ue Mountains, II, 183 Coast, IT, 186, 187 Cuyamaca, II, 189 Arabs wm, TT. 1 REL Fami Ly, IT, 222 California, i223 i IT, 66 1; Lav haere Cotton, IV, Laxflower, IV, 196 Beach, IV, Colusa, IV, 160 Comanche, IV, 159 IV, 200 INDEX OF COMMON NAMES Fre eg ne. 159 60 eat flower, ‘Play: 105 Le sig nde II, 554 me ood, Western, Tit. Legenere, IV, 8 Lemmonia, III, <3 ee peal THE; 51 LENNOA Fam i, Lesquer Lessingia Dwarf, IV, 378 Lemmon : V7 Sa n Pianniecs’ TV 376 Sierra, IV, 380 Slen der-stemmed, IV, 378 pice IV, 380 in Wild, I 2 Willow, IV, 592 Wire, IV, ger eae Western, HHT: ne Lewis Alpin ne, CTE 13 Columbia, IT, ne EL, 134 Kelloge’s, II, 135 Lee’s, II, 1 plus ag ay | Licori ote TI, 613 inook, IT, ilacopsis, Western, III, 218 oy Famity, I, 379 rge-flowered, III, 426 Lemmon’s, ITI, 418 Madera, III, 429 Many-flowered, ITI, 43 Lindernia Long-stalked, III, 688 Short-stalked, III, 688 nseed, III, 11 Daint “¢ wie ig TAIL FaMicy, Be Pe Famity, IV, 81 LoBELIA SUBFAMILY, V,81 Lo belia Brook, IV, 96 Kalm’s, IV, 96 Rothrock’s, IV, 97 Water, IV, 96 parent tree, II, 562 Locoweed Angle- stemmed, II, 607 09 Blue Mountains, II Mae hs eit IT, 606 597 ein II, 591 Eventi? II, 594 Fremont’s, II, 597 cron s Dwarf, *loatnig, III, “ Many-flowered, IEE; 217 Umbellate oe — = woe California, I TIT, 344 Small-headed, IV, 2 e aurandya Valley, Ill, eras Mayweed, IV, 388 Mazus, “capi iE 6 Pr erastei FOAM FAMILY, III, 46 White, III, 49 Woolly, III, 49 mall- flowered, II, 225 Medic Black, II, use Purple, II, 520 mooth- barred 41,520 eae ae Medusa H I, 248 Melampodin, IV, 139 ew ahaa ie i 2 Nodding, I, 192 Sm Menziesia, Pacific, 301 Mesquite, II, 478 False, II, 475 Mexican hy ce te 72 Mezencum FaMILy, , 163 Srl acat American, III, 351 S Cruz, IV, Sierra Foothills, IV, 560 MIGNONETTE FamILy, Mignonette, ea 328 I ore aster, Tejon V,520 i. ILKWEED FaMILy, ipa oe si Milkw California 111: 375 TE ie Milkwor a III, 23 Small-flowered, IIT, 519 Mimetanthe, Downy, III, 706 Mimosa Famity, II, 474 peed : ee III, 1 Be s fee ii,.127 Mint Famity, III, 614 Horse, III, 661 cheery ITT, 65 Round-leaved, III, 661 Sierra, 58 Westy, Ul, 661 Mirabili Frocbel's II, dd Greene’s, II, 1 Sa actiaae - 106 -vigoust < I, 423 Mi E FAMILY, }. 528 Mistlet California, I, 532 olorado Desert, I, 533 Constricted, I, 5 Dense, I, Dwarf American, I, 529 Douglas's, I, 531 Long-spiked, I, 534 itrewor fake II, 376 Ce TL, 376 ais leave: II, 152 Large-leaved, IT, 152 ohave Flower, III, 788 rda, Pla Large- + Saabs III, 6 eet i, 657 Mountain, IIT, 655 Pringle’s, III, 657 Robison’s, IIT, 654 k 651 rosace Candelabewn ITI. 701 oe, Islands Bush, oa: an weed, III, 701 Ciolate toothed, III, 704 Cleistogamous, III, 726 Cleveland's i 714 Com siae TIL 7 Rate S711, 730 INDEX OF COMMON NAMES Cusick’s, III, 719 Cut-leaved, III, 706 De ath ee It, 726 ior es) a g 3 ° 5 =f Zn Sms RS bo EE 7 Layne’s, III, 722 Least-flowered, jane Lewis’, III, 6 Linear-leave ot “err 698 pee Purple, III, 705 w Bush, ITT, 718 opie ae be 711 Ma II, 700. ariposa, III, 698 ean, IIT, arrow-leaved Bush, LU, 715 arrow-leaved Pansy, — — eee M M M M Mottled, I Mrs N N N , 727 otch- peal Bush, ITI, 715 Olympic Large, III, 707 One-sided, ITT, 723 Palmer’s, IT Pansy, aa Parish’s, Hi 7 Parti- colored, ne 699 i ; 726 III, 730 Santa ig Dwarf, Hi7 Se atl, nee rpentine, nr 723 shi ed Large, £8 =} d1- 70 hield-bracted, II, 704 aS aba Bale 2 Hit} Baie. Iii, ae = Ss Ss Sie S Slender-stalked, S lender stemmed II, 699 ae fea gecoatt Lae Smal Howered Smal ok font 710 III ree- i Ting F rple, ITI, 719 Tooth- leave ae 695 Torrey’s, IIT, 723 Varicolored, III, 721 Viscid, III, 722 Washington, III, 701 Weak-ste emmed Large, Hi, 712 Whipple’s, III, 701 Wide-calyxed, ITI, 712 W ide throated Purple. W throated Yellow, Hh Y diow-andaerne Tl, 721 beige gag thik III, Monkshood Columbia, i is Cla Diied. UY Ss warf, II, 75 Kosei Il, 74 Monolopia "Caen IV, het Crum’s, IV, 2 Cupped, IV, an Woodland, IV, 222 Montia Brandegee’s, II, 127 Coast dines I, 127 Com ia f Cordate- eaved II, 126 Dichot s, II, 124 Diffe rent | II, 129 Diffuse, IT, Henderson’s, II, 129 Howell’s, II, 126 Linear-leaved, * 124 Siberian, ITI, 1 Small- leavéed:; i 124 Water, II, 126 124 Ivy- leaved, es 381 Kern, IIT, 3 Modoc, ITI, ae Oregon, IIT, 384 an Sierra, I 641 Small-flowered, III, 389 Sonora, III Southern California, III, 384 Western. III awe oe Utah, ITI, 54 o Bills, Ne 342 nAs Little- leaved, oh 454 Smooth, IT, Trask’s, II, 4 Mountain Misery, II, 451 Mountain Pride, III, 768 Mourning Aart IV, 64 Mou (ue 367, : “TIL, 542 use-ears, Purple, 70 wort Novihees 4 III, 732 pestis III, 732 Mestuae Ae nb Coastal, IV, 4 Columbia ioe. V, 407 Douglas’, IV, 406 Michaux’s, IV, 408 Suksdorf, IV, 406 Lematen’ s, I, 139 Mountain, . 141 Porter’s, I, 141 Short- leave i 138 Slender Muilla Common, I, 397 Crowned, ‘ sid Rough, I, 3 Muk-ko, i a Mule- Sgt Af 102 Gray Me pe 104 642 [Mule-ears] Patselaicial IV, 104 Northern, IV, Woolly, II, 7 Musk Flower, III, 695 Lon bed, ITI, editerranean, II, 274 Rush, IT, 246 Tower; I 3 ia 1, alee 321 Tum 267 Whi sa bay 271 Myrtle, Ill, 368 in, 14, 223 ae tog Famity, I, 92 “Guadeloupe I, 93 e, 1,9 Sender, a. sg “Hispia ITI, 525 Marigold, III, 443. INDEX OF COMMON NAMES Mitre-fruited, III, 446 ountain, III, 44 ehama, ee vee ane med, White flowered, Peicous II, 2 Nemacladus Comb leaved, Af 84 ee nT, 8 ceca, io] 86 Long-flowered, 7 82 Mountain, IV, 8 Nuttall’ s,5V, vy Pe der ¥iV.§ asd lowered, IV, 83 Nem este eee III, 484 Great Basin, ve ff) Meadow, III, 4 Sierra, IIT, 482 Sm ee a4 ered, ake Canyon, III, 481 Variable. leave IL, $4 t-stra Desert, IV, 487 se eg pe rv 486 © FamI ty, I, 523 Carolina Horse, III, 682 men t:- 525 “Common, ne 630 Red, I Spoted mh, 630 Dw H 3 White, III, 634 Hoa “de 2 4 Lyal irs 525 rests ITI, 681 Small, I, 525 Western, I, 526 Nicolletia, IV, 250 Climbing, III, 676 Cutleaf, III, 676 Douglas’, III, 677 Forked, III, 676 Hairy, III, 676 Narrow- lcaved, II, 677 Pacific eee the IT? Pars III, 677 e, III, 680 Small “Enchanter’ s, Sma lowered Viecd bedi marina, aed are oe 11, 409 aca II, 67 Blsdows, 1; “at Parry’s, I, 4 Nun’s Veilin bh aL, 2 Nutmeg, California, tS1 Oak Alvord’s, I, 521 California Black, 517 foe ornia Live, I, 518 net re rub, Csifoena White, 518 ae 1,322 Huckleberry, I, ae Island, I, 521 Kellogg’s, I, zu Leather, I, 5 M aeDonals J 520 Tan-bark, I, 516 ee hivi Wild, I es owe Ey papa II, 414 paemet, III, 474 enanthe American, III, 238 Pacific, Mi poe Old Man, Old Manin on Sects. IV, 462 OLEASTER FAMILY, III, 163 Oligomeris, af Om | OLIVE FamILy, acd 346 Olive, Desert, III, 349 Many-flowered, I, 386 Narrow-leaved, I, 394 Roc 47 Spa vache han 4 weg PL I, 473 White-flowered, I, 474 Large Round-leaved, a ee 478 Ore Alpine, aT, 599 60 H Low, III, 602 Malheur, III, 6 ellow-e paces lig pe IV, 246 Oro California, III, 24 Great Aaetia bai fh a F a Orth froth baited: III, 817 ney Ill III, 8 Paint- ae IIT, 813 Rosy, ITI, 815 San] Bernardino, i j gi n Fraticisco, III, 819 Shasta III, 817 Smooth, ITI, 819 Thin. leaved, IIT, 815 Yellow, III, 815 Oryctes, sWosaste: I Osier, Red, III, 283 Ox-tongue, Bristly, IV, 588 Oxypolis, Western, Ill, Oxytenia, IV, 141 Oxytheca Chickweed, IT, 19 Nuttall’s, II, 18 ee-lobed, II, 19 Tree-like, IT, is Watson’s, II, White- marained ¥F49 Yellow, II, 1 Cusick’s Pendent Pod, i: why 2 Ye low lowered 11.6 peel Plant, IV, 585 Ozom Croak adie 1, 377 Pacific, II, 377 Varied: leaved, II, 377 Pale Face, III, 112 Pat Famicy, I, 344 Palm, California Fan, 345 INDEX OF COMMON NAMES Palo Maas re fia: Border Sree as: a 479 Pana mint Pains II, 240 a Weattin’ 4-175 Pansy Pinto, ITI, Me llow, ii na Paper- flower, IV, 194 Parachute Plant, EV; Parentucellia, Yellow, III, 800 Parishella, IV, 88 ghd IIT, 23 oh Mountain, TIL 2 Wooly Mountain, Tit; 2 pina fe i Cow, , III, 2 Cut-leaved ee 231 Hemlock Water, rae III, 278 Panamint Indian, Pestle, III, 2 Pasque- Andee Wessies, II, 193 Patata, IT, 74 PE Bones II, 480 Sarst. II, 6 622 Sierra Nevada, II, 622 Silky Beach, IT, 6 att II, 6 Swee II, 619 Teel er, II, 619 Torrey’s, “ 619 Tule, II, 6 Watson's ort 623 ri H FaMILy, IT, 465 ee eng rt, II, 466 she: pads: IL, 145 Beach, IT, 145 ae IT, 145 Sticky, IT, 145 Bristly oe Broad-fruited, III, 536 Chuckwalla, IIT, 536 Little, III, 537 Recurved, III, 536 Slender, ITI, 536 Winged, ITI, 536 Pellitory Florida sh, oe Pe cheseanta! I, 526 Pencil Leaf, II, 338 su, Pennyroyal nL 661 Dwarf, ITI, 645 Blue-etewinieil; 121; 770 Broad-leaved, III, 746 Ht: Cushion, III, 760 Cusick’s, III, 760 Cut- leive d, IIT, 764 Death Valley, oe 755 mnt! ngs Dese sig ate ny 763 Gairdner’s, III, 747 << te pag y, 111.7 Glandular, re 764 Glau ger Aaeb Globe. TT roan Sid LH 751 Mono, II, 750 Notable, III, 756 643 Owens ae? 758 PL. Redwood, III, 770 Rock, III, 76 Rose, 2755 Rothrock’s, III, 769 Sand-dune, ITI, 753 San Gabriel, III, 758 a ; wae Me Shas Share: gain gent 747 er III, 746 ephens’, III, 756 estern, ITT, oe be ase Vallow Washington, III, 739 Vhite a's, IF0, 751 a tt et x pe ee | IM > _ if y hite- margined, , 748 Wilcox’s, III, 746 Wo Bt eony, Western, 33-175 Pepi mt III, Pe epper-grass Dwarf, II, 262 English, IT, 259 Fremont’s, IT, 261 PEPPERWORT FAMILY, LE? Pepperwort, II, 282 airy, I, 34 Nelson's, I, 34 644 Pericome, Tailed, IV, rishi III, 368 sicaria “Amerian Water, II, 62 r* le, 66 Pennsyvania, II, 66 Spotted, II, 63 sas , Il, 63 Peruvian Pepper, Iz, 51 Petunia, Wild, IIT, 685 — n Nelsons a 497 Pediceliate, at td t-flow "50: Pringles: Piss Purple-bell, III, 516 Racemose Sho, bese ent 497 Sticky, III, a= Stinging, III kool} a 511 INDEX OF COMMON NAMES bonis Lu, ‘gra Tan y, III, 4 Thick ‘leaf, iI, 515 Pp irgate, TH, 500 Waterleat, Ill, 4 489 x Fam y, III, 396 Phio Bear Siew. ae “ Clustered, ITI, 4 Coville’s, ITI, jy Douglas’, ITI, 411 Gray, II, 413 Long- saves “ve 409 Moss, a peor III, 411 Prickly, III, 454 Showy, III, Slender, III, 413 bi II, 412 Stansbury’s, III, 409 Western Mountain, I, 41 Wo priests wr gee Woolly, ae Phoenic Broad- ded, "i 290 Comm Pholisma a ree Re v, I, eiient = 91 Pigmy-cedar, IV, 460 ig ees I, 52 Pin Big Cone, I, 59 Mountain White, 153 Nut, I, 5 One-leaved Pinon, Parry’s Pinion, I, 55 ns -cone, I, 60 Sugar, I, 53 Tamarack ye 59 Ea Sadie 1, §3 Western Yellow, I, 56 White-bark, I, 54 Pineapple Weed, IV, 400 V, 400 American, III, 295 eo III, 296 nk boniet IL. 174 Grass, IT, 171 Mullein, IT,,174 Pi Purp Pipe-vine, Califor 1.53 Pipsissiwa, EEE 292 Pitayita, III, 162 Pee PLANT FAMILY, , 329 — Pe Pitcher- plant, Cali- acl, 329 pe icons. IIT, 2 Plagiobothrys, loses EEL, sae FAMILY, II, 403 Plane-tree, California, II, PLANTAIN FAMILY, IV, Plantain Alaska, IV, 17 Bris st shih Large-bra cted, IV, 20 es’ Rattlesnake, 481 Pesican A Pac ific Soli iV,47 Ac aulescen, II, 487 Agardh eating WRAY II, 487 Eastwood’s, II, 489 Fresno, II Mojave, II, 489 Mount Hamilton, soeet 487 pi ve, II, 487 its 487 red, II, 487 Sunse t: 35, ong- spurred, IV, 62 Pink, I White a Pleuricospora, ITI, 296 Pleuropog California. I, 194 Nodding, I, 194 MBAGO FAMILY, , 344 Podistera, Sierra, III, 239 Th Soh ae in 647 Poison Hemlock, Poison Ivy, ITI, 51 Poison daa Pacific, Tt 5 Pole hetresl TIL, 397 Elegant, III, 400 California, II, 158 oa ur-leaved, II, 158 gala Caltornia Hi 21 Desert. III, 2 Polypody California, 1,8 ti. 7 We We n,I,8 meray. Indian, Tr 172 PoONDWEED FAMILY, , 83 Pondw eed Co oe Floating, 1,8 ees 1 3/ Eel-grass, I, 89 71,85 Various-leaved, cess stemmied, - “a Prt dog Bush, Penn III, 574 Poplar, Balsam, I, 486 Poporo 677 Poppy Berle L223 oppy Broad-horned Prickly, II, California, II, 227 Island Tree, }1, 226 Leafy Prickly, II, 230 Matilija, II, 226 Merriam’s Bear, FP 232 Mexican ee II, 228 Mojave, IT, Pale Rodeke tenia lla, Potentilla, eae II, 431 Potato hie gies Ill, sees fs , Poverty Bs oe 141 Prairie Clove Blue Mccinlatis: II, 559 Searls’, 55 Prairie- rocker, small- flowered, IT, 318 Prairie- Shdte. II, 449 Pretty Face, I, 399 INDEX OF COMMON NAMES Prickle- leaf, IV, 551 aise: _ ae 200 Beach, ITI, 2 Booth’s, ITT, om Clavate-frutied, I, 206 rt Plateau, III, 201 Refracted Desert, III, 197 Shredding Ill, 201 Sierra, III, 334 ender- dowered, IIL 197 Small, ITT, 2 Wallowa, ee ae Willow-herb, IIT, 200 Wing-seeded, III, 204 Primula Cusick’s, III, 334 Sierra, III, 334 s, IV, 232 2 Pterostegia, I Pteryxia Rock, III, oe Ill, 274 oak Ill, 548 Purple-bush, Ill, 24 Purple Mat, ITI, 525 Eee Ill purer: 1,135 American Marsh, III, 168 7 III, 117 Sea, II, 77 Southern Marsh, Paw W se a in, ie 18 the Single- headed, IV, 274 IV, 272 Say. IV, 274, 276 Quail Brush, IT, 86 QuassIA FAMILY, Queen Anne’s Lace, II, 22 228 QuiILtwort FaMILy, 3 Western, , a Rabbit-bru Cha ee IV, 300 Common, IV, 302 cae IV, 300 ng-flowered, IV, 300 Nectleeaved, IV, 298 Parry’s, IV, oe Pune ine iV52 Sticky-leaved, W, 298 Rabbit-foot, IV, 156 aa s Foot, I, 147 Radis rae a Tip2r5 Wild, II, 275 Ragwort, Tansy, 450 Raillardella Green-leaved, IV, 428 a V, 428 esti: 457 a siountain II Ww oe Sem IT, 457 645 “Glandular, II, 628 Pine, I, Msp s, IT, S75 Cade IT, 586 Cima, IT, 59 Clar a Hunt's, IT, 603 Clovelsadie IT, 603 Prov ‘idence Mountain, IT, 582 Rattan’ s, II, 603 ernardino, II, 602 Sand, IT, 587 San Diego, IT, 583 Redberry, Ill, 63 646 Red-bud, Western, II, 478 Red Maids, III, xal Red Molly, 3lu jolander's i, is 7 sre Slate leave i 314 3road-s t Bee ee I 306 Coas coteloe a tia ot ve 306 Dese I, 305 Dr caeuie II, 307 Hairy, II, 306 yori: Sea, II, 270 Dame’s, II, 321 Dyer’s, II, 328 Garden, II, 271 Sand, qt, 275 INDEX OF COMMON NAMES Sea, II, yi Wall, I Rock- ‘inge, LLY. rahe Rock ipa Des TIT; Rock- ae Parry’s, IV, S571 Rocknosz FaMILy, III, Rech’ they Elegant, Il,'3 Ro ck- ne nate FaMiny, ~s ge Ros Alderson's II, 462 Tehachapi, IL. 463 Wood, II, 465 Rose-heath, EV ;,332 Rosemary, III, 620 osettes oe Weed i 282 Rud California, IV, 118 ington, IV, 118 gee Pie TIE, 17 Rue pots 19 Western, IL, 1 6 5 Rusu Famity, e 349 Rush Baltic, I, 354 Basket, I, 355 Brownish, I, 280 White, ad 280 Black-grass, I, 356 Blunt- EN I, 360 On) Bolander's, I, 365 Brown-headed, 367 Cece I, 364 Clu Co ville's, i Dru aaa hg 352 raise L,:357 sa Je gh, I, 361 Sh Iris- ie ey I, 368 Jointed, I, 362 Knotted, I, 366 Long- anthered, J. 367 eae Mountain, 1,3 Salt, . 354 ri harg-funted. I, 366 pie 1,357 ickle- leafed, I, 359 San Jacinto, I, 264 Spiny, I, Spreading, I, 353 Straight-leaved, I, 358 Thread, I, 35 ree-stamened, I, 368 To Western, L 358 Woo Common, % - 1 Small- flowered, I, 369 Smooth, I, Spiked, I, mL. Phun aay I, 365 ‘Alderson’ ae one Common, III, eat TIL, re Sacaline, II, 66 Sacapellote, IV, 550 Saccaton, Alkali, I, 145 Pe ad IN; S42 age African, III, 636 Puen III, 639, 643 Red, II, 89 Rose, III, 6 Thick- oa oe 641 Thistle, III, 6 Cutleaf, IV, 414 Dwarf, IV, 412 Hoary, IV, 414 piven IV, Hey IV,4 are na,i1, Twining, I, 408 Salal, III, 306 Salsify, IV, 585 Coast, Coulter’s, II, 82 Crown, II, 7 Dav "$3 Nar Nuttall’s, 1a 87 i I Salt Cedar, I, 181 Saltweed, Gmelin’s, Tie 78 Salt. hae EET SASS Blac III, 333 SAL LVINIA Fan 1}-35 i Ill, SA Lwoop gts Death = soos ThE 133 Sprtiniet eaved, Ill, shining Iti}? re rsley, Wedel 1, 22: 7 ee ts Spurry Boccone’s, oat 455 Canadian, II, 155 La FP Medite erranean, 1,455 Middle. -sized, II, 155 Pu rple, II, 156 Sek 1; 155 oo II, 156 erbena goes i, Bs Beach, II, 110 ston Il, 108 INDEX OF COMMON NAMES Honey-scented, II, 108 Bear ne he ae 150 Beautiful, II, 150 ) leaved, LI; 152 eal, II, 147 II, 147 Thyme fies 455150 Sani petal III, 221 Bear’s- foot, ee 220 Coast heriag eh got Pacific, Ill, 221 sh, I Sh arp touted, = on Sierra oom Tubero a pon Sanseitatin: ’ Abert’s, IV; Sarekpatit: T, 458 bie. “i 215 Sat MakGene. 7 III, 646 San Miguel, I ase rics Searweet J Amer IV, 509 SAXIFRAGE FAMILY, erect 2 Pigmy, Il 358 Piper’s, II, 362 Purple, ii; Red-stemmed, II, 360 Rusty, II, 366 Sierra, II, 362 otte 97 Schoenoerambe II, 246 Schoenolir Tract d, 14 White ek LAt2 Scopulophila, II, 113 Scorzone Scouring- -rush (see Rus Screw-bean, IT, Screw~ Sores 520 Sea-blit Black, IL, 95 California II, 96 ET 95 Sea Sel -pink, III, Sea-scale, II, 84 edella Congdon’s, II, 333 Mount Hamilton, Sierra; 1, 33% SEDGE “ideas P2255 edge Abruptly Beaked, Alpine Sierra, I, 333 Ample-leaved, I, 327 Awl-fruited, I, 298 teen Bro : ade, I, 317 Clasping-bracted, 2 Cloud, I, 309 Clustered 647 Northern, we 315 Coiled, I,2 Flaccid- dea : Foot-hill, I, 2 Ox, a 296 Wes 295 Fr agile sheathed 1,'30: Geyer’s, I, 31 Golden. ‘fruited, 13322 4 ee net bary £2335 athed, I, 302 tet Orney’s 324 Mountain, T, 307 nd, I Inv ofute-leaved, I, 291 Jon desert Ss, 7 Je Large-headed, I, 293 Lemmon’s, I, 325 Long-awned Alaskan, Long-stoloned, I, 319 Luzula-leaved, I, 326 a-like, I, 325 Merten’s, I, 332 oe jE 8 Moana Nea 648 [Sedge] Nebraska, I, 334 L 97 Piper’s, I Porcupine, I, 343 Presl’s, I, 312 3 ocky Mountain, 3 Holm’s, I, 332 Ross’, I, 320 Rough, I, 337 Rough-sheathed, I, 333 Round- ag rf 319 Rus ty, ; Slender-beaked, Ss 302 oe ibs Smal ‘rated, 1313 oth Scag fruited, 1,330 Smooth-stemmed, 2 | Softcleaved 1 $14 “hick-fruited, I, wile “*hick-headed, I, : ‘hread-leaved, i, aa ) 7 + 4 4 Jn 3 ) , 30 riangular- fruited, I, 32 waar t2 At ERS 7330 Vernacular, I, 291 Water, I, 336 INDEX OF COMMON NAMES Well- fruited, . oe 1,339 Sedu aes vy aad petaled, "344 Seep- willow, IV, 386 ae GINELLA FAMILY, I, 46 Selaginella y Mountain, I, 48 ur oe III, 628 = pintaagihee MILY, IT, 478 na, Armed, II, 479 Sequoia, Giant, I, 69 , 471 Séabanis "Big-podded, II, 562 Shad-scale, II, 87 Shamrock, IT, 523 ais Fat, IT, 87 pod peta II, 579 Purple, II, 578 III, 224 Bee s Purse, Shield pod, California, gar nese aH 290 Shooting Sta Sidaltea-. Cre copies: TI, 103 Cut-le aved, III, 106 ged, III 98 Glancous, Ii 105 Hairy ake i a Hartweg’s, It, 01 H gress Powers III a 05 Spiked ae 102 SILK-TASSEL FaMILy, Silver-weed ecrae ee — Pacific, Siepasbates oid II, 267 Skeleton- plant, Thorny, V, 568 Skeleton Weed, II, 34 Skul Austin's Lit; 623 e, IIL 624 Beat leaved, 11,6 Sn a IIT, 623 Skunk Bush, ITT, 51 Smallage, , 228 Small-scale, IT, 83 Sm eed Smilax Californ “ anes fees Desert, EE Snake- a. III, 220 Snakes-head, IV, 576 | Ti6!: Ev; Western, “i 62 Snapdr. ewirs. TIL. 790 Dimorphic, ITI, 791 Garden, ITT, 789 Golden De sert, IIT, 788 Lax, HI, 783 Least, III, 791 Nuttall’s, Il 1,791 Oval-leaved, III, 791 ee ee III, 790 Talli 89 Violet eine ble ITI, 784 White, ITI, 790 Wir y, III, 790 Yellow Twining, ITI, 783 Sneezeweed Bigelow's, IV, 211 Ni 2t2 Western, IV, 210 Snowball, IV, 47 Snowberr Common, IV, 48 anoa, III, 374 oliva Common, IV, 400 South American, Solomon's Seal re I, 54 Spectacle Pod, II, 286 Speedwell American Alpine, ITI, 793 Broad-fruited Water, III, 79 Copeland’ 2 bi 793 nal Ill, 796 rine ara cle se 793 Persian, III, Sonar es me-le aved, on 793 abe destnas § HT 273 Spikenard, California, i aps woop : ne horned, ry 15 a, 25; < Spatula pleas iS : pil cae I, 411 soca “ 181 Ose- emo ;t81 Senn Beauty, Western £322 Spring Gold, IV, 454 Spruce , 1, 62 SPURGE any: 1128 Spu Abra, JEL, 39 Clift. III, 3 32 INDEX OF COMMON NAMES Cypress, III, = Desert, ITI, 3 Hall’s iets, TI, 24 Hoover’s, III, 4 Indian, III, 42 Large, III, 40 Leafy, III, 35 Louse, IIT, 40 III, 34 Reticulate- seeded, 2 Hl,.3 Ridge-seeded, III, 39 Shrubby, III, 24 Spurr 5 ehgele, II, 473 ye tail, I, 246 m1 254 Bottle-brush, I, 254 STAFF- TREE FaMILy, os 53 Pos Hill, IT, 369 Elegant, 1 Green banded, I, 441 Hairy, I, 434 Large- ees I, 437 Lobb’s, I, 436 pom veges I, 438 437 Long-stilked: Il, 139 Northern, II, 139 Rocky Mountains, t. Catharine’ s Lace, It Steers Head, II, rs Steironema, III, 3 a gy redores ITI, 731 Stensnthienk: Western, Stenophyllus, Hair-like, Stenotopsia, IV, 283 Chicory-leaved, Few-flowered, IV, 572 Gray, IV, 57 uarge- -flowered, Tv, 70 eee leaved, EY; 5 V, 574 Small-flowered, Veatch's s, M1, 139 Venta White brated, a J, 140 Tit 557 cere ds IIT, 550 O Pink, I Velvety, III, 555 Western, IIT, 549 Stick-tight, IV, 128, 129 1V:t368 res leaved, base 30 Mojav Seimpaceee bush, 1, 483 Stink Be see Stink-w California, II, 326 5,45 327 California, I; 130 t, I, 129 Com n, IIT, 116 pn Cansiian: TI, 417 Ry flowered, III, 117 Scouler’s, a 116 Stock, II, 3 ene ie ILY, oi ieee II, 346 Narrow-petaled, IT, 34¢ Pacific, II, 346 Star-fruited, II, 346 Poeiedierto air Il, 253 Strept Alpin ae 7s 248 Bearded, 248, 249 Brewer’s, II, 249 a Dien: II, 250 Samoe EE. 247 650 [Streptanthus : Tamalpais Varied- leaved II, 251 Styloclin Poebodus IV, 486 Hooked, IV, 4 Styrax, rayhena Hit. 3 Sugar Bowls, II, 193 Sugar Bush, III, 51 7 u only Sullivantia, Oregon, at, 35 Sumac Famity, III, 50° “Chapera LH;54 51 undew Obteng: leaved, II, 330 Round-leaved, II, 330 SUNFLOWER FAMILY IV, 98 IV, Woolly Dwarf, IV, 104 Seer sie eee es Sweetie, II, 458 gp goatee s, II, 614 614 8 lunt-fruited, I, “eas California, IIT, 22 Mountain, ITI oe INDEX OF COMMON NAMES Purple, III, 226 Western, III, 226 weet-clover White, a 521 Yellow 21 Sweet res III, 241 Sweet Gale, I, 508 pasting scented Shrub, Parry's, a, a Pine Shintae. 3 Oe 63 gl sab stemmed, Syeamore IT, 406 Syn bias IV, 234 Syrin California, me, 385 t-1E.3 ack he it} er Lewis’, II, 384 Tack-stem White, IV, 582 ellow, IV, 583 aaa Til, 373 Talin Oka cals. — 121 Spiny, III, 12 Tamarack, eis 60 TAMARISK Famity, 0 Five-stamened, III, 120 French, III, 12 Small- ry Ene II, 120 Tamarisk Salt Tree, III, tae aS 376 Cinquefoil, IV, 398 Dune, IV, 397 Sierra, II, 2 estern, ig tk TAPE-GRASS FamI_y, I, 103 Sle Tine, II, 616 on IV, 410 — eed IV, 192 Catalin, IV, 174 Chile 170 Cleveland . IV, 184 Hall’s, 1V; 17, Hayfield, IV, 182 Headland, IV, 170 Heermann’s, IV, 186 EV: 168, et 176 Small, IV, 1 So onoma nd a Tecate, IV, 178 Three- rayed, Ae 176 Tracy’s, IV, 1 Pid airins IV, ne schia be stew III, 243 artweg’s, III, 243 Hoover’s, III, 242 Howell’s, III, 243 Kellogg’s, III, 243 Parish’s, III, 243 Southern, III, 242 Strickland’s, III, 242 TAXODIUM FAMILY I, 68 Tea, California, II, 553 TEASEL Famity, IV, 64 Tea Fuller’ “f of 64 Wild, I lee Vien ITI, 66 Ec Weed, 1V;2 Tetra Holly. fae III, 24 Le 246 enersty iy "244 Red-purple, II, 242 Sasittate, IT, 243 Short-podded, II, 243 Siena ek wind. Yellow- flowered, 1. aps Therm Cabtoseaa, II, 482 Rocky Mountain II, 482 Silvery, II, 482 Slender, II, 482 Thin Tail, I, 238 Thistl Anderson’s, IV, 536 Ashland, IV, 518 Distaff, IV, 5 Douglas’, IV, 520 warf, IV, 532 Edible, IV, 528 Fountain, IV, 538 Franciscan, IV Peregrine, AV, $22 Red, I pas (ates IV, Russian, II, 96 Scotch, IV, 510 oor, flowered, IV.5 RAE ss 530 Smooth Distaff, iV, 512 Snowy, IV, 526 Sicen | Mountain, IV 3 PM ie 520 Surf, IV, 528 n San ee wee 635 San Diego, III, 635 Santa Cs. III, 635 Thousand Mothers, Th €; 1,27 Thrift, California, III, 3 Thyme Creeping ’ III, 6 Tiarlla, Laciniate, 371 ace II, 46 Tidy Tips, ee 160 Tillae ea, II, 3 Wild, I, Ti nidaeky iat toe ~ Tipton-weed, III, 1 ad-flax Bastard, I, arger Blue, 8 Indian, ITT, 685 — flowered; I, 684 Mexican, IIT, 684 Ban 4 Faas a III, 66 Tofieldia, Western, 372 Telguacha ITI, 682 II, 474 Large-flowered, III, 782 Small-flowered, Tooth- “cup, III, 164 Toothw Calton, II, 283 Tornillo, 1 478 Tor +; Small leaved, tL, a Touch- -me-not, Pale- w, III, 59 Townu Climbing, III, 373 ——— Ill, ah Tracyina, IV, 3 INDEX OF COMMON NAMES Tragia, Desert, III, 29 eciieee _ og Tre Tre af Hbdved) III, 19 Trefoil, Bird’s-foot Ti 353 Tricardia, III, Tri choc coronis, ey, 496 riodia Awnless, I, 184 Low, I, 184 Trisetum Beardless, I, 167 403 Trixis, ata IV, Tropid Caper- oak II, 285 Slender, IT, 2 Trumpet Desert, IT, 31 Little, II, 30 Yellow, IT, 30 Tule Potato, I, 101 Tu meniew ert II, 99 Tun Coville's omg 152 ae piesa TE 152 Turkey- lah fet 13738 Turkish Rugging, II, 6 Turnip, II, 274 Porvantinetedls LV; Turpentine Weed, I, 619 Turtleback, IV, 456 Twaybla Broad-lipped, I, 480 Heart-leaved, I, 480 Loese l’s, I, 481 Northwestern I, 480 Twin-berry, Black, Twin-flower, Western, Ea ted- stalk Clasping-leaved, I, 457 Simple-stemmed, I, 458 Umbrella > acevan Ji355 e, III, 20 Yellow, IV, 2 Uropappus, IV, 558 VALERIAN FAMILY, IV, 56 Valerian, IV, 56 Califottia, LV p57 Northern, IV, 57 e eria n, II, 2 pore lowered, a a | 1¢224 Vanilla aes 11219 V b,t1ij2 Venus’ ia, on IV, 78 Small, IV, 78 Vera Dulce, III, 613 erbena Cluster-flowered, III, 610 Mintleaved, bet 611 Rough, III, Wascen: a oi VERVAIN FAMILY, ae -— Ver Blue, e, Br aan aL "612 Desert, III, 6 Mu len- laved, “ie 611 ran tg II, 6 jee i, 617 Smaller ‘Common, i Spiny Milk, IT, 568 Spring, II, 615 Winter, II, 617 Woolly, II, 617 Vetchling Low, II, 619 Marsh, II, 622 651 Ag a II, 6 Viburnum, oe leaved, 7 Viguiera Laciniate, IV, 1 Leathe leav “298 IV; 21 Nev fing IV, 112 Parish’s, IV; 3i2 Vinegar Wee d, III, 619 Viotet Famicy, III, 123 Violet Alas ka, TEE pr B Butterfly, III, 129 California Golden, Canada, III, 129 i ae — 129 Douglas’, III, E te ; III, 132 Evergreen, IIT, 126 Golden, ITI, 125 Great Basin, ITI, 125 He 1 325 ooked- -spur, III, 129 ee Ill, 125, a Langsdorf’s, ITI, 1 prea ag s oe “ a III, 1 e, III, 12 Redwood, aoe 126 129 olymni, III, 129 5 s leaved, III, 126 piney Sweet White, III, 1 Waciee: Water, III, 132 Yellow Wood, III, 125 Small- leaved, ? 194 Western, hasenae robin Repand, IT, 315 Sierra, II, 318 Western, IT, 318 Watnut Famicy, I, 509 652 Walnut, California, Wart- -cress, Lesser, Ee? ht ete ae ae Wat r-cres s, Ti; 276 Water rate IV, 96 Water Hem Bola nder’ : a 238 Bulb-bearing, III, 237 \W cee rFackey ae 667 WaTE FaMIty, III, 476 Waterlea California, ap 478 Fe ndler Water- ecg Autumna IL. 43 Seon Ill, = se ornia, III, 4 batter r-wally, IV, 386 peieny Mountain, L 103 rend IV, 384 , 168 Wax-myrtle, California, ii, 33 Whisk-broom, IT, 35 Whisker-brush, III, 427 Whispering Bells, IE, 521 White-heads, Swamp, nolan rns tt, 294 INDEX OF COMMON NAMES Carolina, II, 292 ie eee II, 294 292 Whit itueya a EVs: W Sacer est Red, III, 329 WILLOW FAMILY, I, 486 illow — I, 500 ses be I, 496 arclay’s, I, 498 ea I, 503 Bebb, I, 506 Bluish, > an Bog, I, 4 Brewer's 4 504 c 1,4 False Mounain, 7. 497 Farr’s, I, 495 Fe athe: ovein: : ae Firm-leaf, I, 4 Rocky Mountain; I, 507 Canater: I, 505 Sierra, I, 499 Silver-leaved, I, 492 lavate-f ; Tit 177 elicate, III, 176 Elevated, III, 17 ai wf cadutsens IIl, 179 a. III, 176 Slender-fruited, III, 176 now Mountain, HI, 2 Stiff, 113,173 White-flowered, 9 RGREEN FAMILy, Winte Kou Spicy, III, 306 Bog, III Common, an 291 English, III, 291 Gree — fiawees d, Saber-shaped, I, 3 bee shaped, c er Woodsi Oregon, ky Mountain, a . Wane: -SORREL Fam Trillium-leaved, III, 8 BAL en breeches, 479 Woolt-heas, i 2 Seaside, IV, 228 seed, II, 315 Wormwood, IV, 4 Woundwort, Field, III, 631 Wyethia Bolander’s, IV, 101 Coville’s, IV, 102 Humb oldt, IV, 104 Southern, IV, ‘100 White-rayed, IV, 104 Xerasid Fremont’s, IV, 234 Lemmon’s, IV, 234 Yampah Bolandas’s III, 234 Parish’ 234 Prinele's, Ill, 234 — ommon, IV, 390 Yellow, IV, 227 Yellow Bell, I, 426 Yellow-c Blunt- Bae - med Colu mbia tIs2 ke: oake Rattle, i, _ Yellow Turban Yerba ‘Amavilia. ul, 13 Yerba Buena, III, 6 Yerba de Chivato, 1,19 Yerba de la Rabia, Yerba de soak Lt, $88 Yerba-de-tag beg del Panes 447 Bore nde II, 388 Yerba Mansa, I, 485 Yerba ses .A19 Palmer’s, = 12 Yerba Sant California, III, 526 ma LTE. 522 c,IIL, ny 6 San eg chat 527 Thick- = ne 527 woe Tit, 527 patric T5850 w, Western, I, 51 You -on- gene II, 374 Yu Fleshy- sheers I, 447 Mojavy 47 Desert, I, Fremont's, I, 375 Giant, I, Glaucous a 375 Sema 6 ocky ree OES gd Mn Poa 1,376 Abama, I, 372 californica pore décidentalis, I, 373 bi 65 concolor, I, 67 pale tag I, 65 I, 66 heterophstlo, Ag = magn ITI, 86 hastensis, I, 67 menziesti, I, 62 mucronata, I, 64 nobilis, I, magnifica, I, 67 pattoniana, I, 6 picea, I, 65 shastensis, I, 67 venusta, I, 68 williamsonit, I, 63 bronia, II, 1 acutalata, oz 110 alba, Il, alpina 3 oe 1 angulata, BH; Se arenaria, II, turbinata, II, umbellata, Il, aa 110 onyms and references in italic.] variabilis, r eT, villosa, II Mistiahind inten IV, 410 Abutilon, ne a 110 abutilon i eh pits newberryi, III, 84 m, IIT, 83 2 Acamptopappus, IV, 2 microcephalus, IV, a shockleyi, IV, 262 sphaerocephalus, 262 hirtellus, IV, 262 ACANTHACEAE, IV, 13 Aca meee enae corru- ealike II, 14 po olygonoides, {7,12 rigidum Acsdtioniateg he 635 carphacd artemisiae - oi V, 242 Acer, HI, 56 bern ardin num, III, 58 californicum, Ill, 58 modocense, Hil, 38 ps sie iiekce III, 56 torre ae a ACERA ACEA E, pet en ne pent III, 375 tom rentosa, III, 375 borealis, IV, 390, mae arenicola, IV, 39 vey Sate, a 391 A 1V;3 californica, vi 391 IV, 390 A ichyranthes lanugi- pecan dchyrodes, L 196 aur. Porites tL 111 Aconitu 190 laccaefolium, a ate rtia im icrocephala, V, 550 eee affinis, alvicaulis, III, 139 nitens, III, pectinata, IIT, 139 veatchiana, III, 139 Acroptilon picris, 653 INDEX OF SCIENTIFIC NAMES [Families and tribes in SMALL CapiItaLs; genera and species in Roman: sy IV, 541 | Acrostichum marantae, e es i ae A I, 20 Pes 78 arguta; ab 17 elata, II, 1 spicate, u, 17, 179 arguta, A rte: eri, le mmonii; odorata LIV, 0 Actinella biennis, V, 208 eee rYV;, mi pak rickardsonil canescens, IV, 208 Actinolepis lanosa, 1¥i2 lemmonii, IV, 234 ‘acet, 1V, 230 I, 347 neo-me-xicana, III, 349 Aden lon, IV, 416 bicolor, IV, 416 iuteprifo olium, IV, 416 Adenostegia bolanderi, canescens, IIT, 849 hansenii, Ill, 855 nelleri, III, 850 He ei involucrata, intra, III, 853 pringlet, III, 8 evibracteata, ITT, 853 tenuts, III, 857 viscida, III, 855 Adenostoma, II, 446 fasciculatum, II, 4 fas. ea Se densi 654 <5 Sete rum, II, 446 ra i = 446 630 gone gees nardos- a, IV, 457 ‘Adieu tum, I, 23 ca ilisemet: J; 23,24 emarginatum, I, 24 jordani, I, 24 ee i »1,24 pedatu m aleuticum, , 24 angiferinum, I, 24 hitolpicis alld; 79 ve entina ciliate, IV,1 Aegilot hystrix, I, 254 incurva, I, incurvata, | I, 239. des stalactites III, 445 Aes chynomene sesban, II, AESCULACEAE, III, 58 Aesculus, III, 58 californica, m1, 58 hippocastan III, 38 Agastache, ITI, 624 INDEX OF SCIENTIFIC NAMES on IV, 564 IV, villosa, IV, 563 gracilens, IV, 564 564 tsomeris, LV, 563 leonsinton aspera, IV, longissima, os 563 plebeia sae etl WV, 564 retrorsa, I nou ata IV, 563 ryposepala, = 446 Snes. II, 446 Agropyron, I, 239 arenicola, I, 249 bi a gt Arona 1, 240 occidentale my gmelin i pringle, . 244 in rm 242 leusneiniael I, 240 laev as we prin faeudorcne hi 242 eeaay T, PF riparium, I, 240 axicola, I, 244 smithii, I, 241 lle 1 spicatum, I, 244 ie eng ia eg mo e,1,2 puberulentum, y 244 pubescens, I, 2 eubeilfosin : pee m, I, 242 ongifoinm, 1242 239 californica, I, 153 canina aequivalvis, I, 149 ee I, 145 cruciata, 1, 176 crtanis I, 146 152 geminata, grandis, I, longiligula, I, 154 lut eo com Ger ma a4 morophl ss oe #1, 153 miliace, £135 a, 1,153 occidentalis, 1,152 or 6. 1155 yacemesa, I, 139 scabra, I, 154 retrofracta, I, 149 rossae, 7 Pi scabra seth ced Ts ‘ 154 oo uleri, 1, 15 sen. I, 55 spica-ve Pha i mi L an 150 pie aso 155 thu rberi variabilis, I, 153 varians, I, 153 Ranting ig IIE, 19 , 16 atro tee I, 165 aespitosa, I, 16 ve illaris, 1, 166 car reephyitea 1 166 cristata, I 169, 170 conia, I pea fraesitinge IV, 104 helenioides, 1V, 102 Albonia peregrina, III, 19 lcina saith y oe IV, a arcti- s, III nese rnd mE “ $4 pacificus, III Alguelagum oon pe maurorum, II, 614 Ebdon triodon, III pestitins III, 471 Ali seered niger ay, 63 geyeri, I, 98 slantaaen -aquatica, I,9 rostratum, I, 99 re te ‘ordatun, ‘, ” Auber a nae IV, 390 ae ae procumbens. I, 669 Allenrol ea, II, 90 ziy coccised; malacoides II, 104 cuspidatum, I, 390 gracile, I, 39 ambiguum, I ampeloprasum, I, 380 amplectens, anceps, I, 38 aberrens, ry 387 ewe , 385 fibrillum, I, 393 INDEX OF SCIENTIFIC NAMES fesbainiee I, 389 acre i 383 m,1,3 cere. ts 383 3 ace la, c ment a 390 phe cals III, 570 ey 1563, 567 anne pie, III, 561 arcolata, III, 571 III, 562 bracteata, III, 570 californica, III, 571 conjuncta, ll, 570 cooperi, ppd a At 565 Pict III, 562 ‘yocarpa dispar, III, 571 di stantiflora. III, 562 diva ricata, III, 566 PN Hoe Tit, 561 figurata, , 565 filicaulis, III, 567 hen nil, III, 587 hick sae 565 hirta, III, co eR IIT, 565 hispidula, III, 570 unda, III, 566 pea tree III, 566 laxa, III, 5 leibergii, i 570 leptoclada, III, 562 limicola, III, 559 m a, III, mollis, III, 559 stita, III, 559 nigra, III, 570 oricola, III, pratensis, III, 567 ramosa, III, reti culata, Ill, 571 t tenera, III, pe trachycarpa II, ursi num fuluescens, LE St Allasonas mucronatus, I, 31 Allotropa, III, 292, 293 virgata, III, 293 Ne nus, incana virescens, Ly occidentalis, I, 514 es 143 3 paliack 2, b pratense 143 , 1, 143 Alvan TI, 613 wrightii, III, 613 Alsinanthomin euro- jlnttielcs. III, 333 Bender! baicalensis, I, 138 bbcaeel iL i585 viridula, II, 139 was: shingtoniana, II, 139 Alsinella ciliata, He 145 aulis, II, 1 jectteutalin Il, oy AT oc, Diner, pe 3 4 é P t/ ¥ a, II, 146 douglasii, II, 146 656 [Alsinopsis] howe ili, TL, 146 pits ba, Tl, 150 calyctnum, II, 319 AMARANTHACEAE, II, 98 Aviavuntlids , II, 98 baictestes, TI; 99 albus, IT, 99 blitoides, II, 9 rassior, I californicus, II, 99 pe teo“gy ; 98 Amareta pe ay iis copelandis, 135359 lembertit , III, 359 Us procumbens, I, 323 Aaaavicioatcad: I, 459 Amauria dissecta, FY; 221 A oe gn ad dissecta, IV, 2 A ees moschata, muvicata ,1V, 542 Amblo cyna fimbriata, II, 99 ior its Staal IV, 248 neo IV.2 seiactie. IV, 2 Ambri artemisiifolia, integrifolia,] a Laas pilostachya, Vv. as cali — fie 148 pumila, 148 trifida, IV, eet IV, 146 ROSIINAE, IV, 140 INDEX OF SCIENTIFIC NAMES Amelanchier, II, 470 alnifolia covillei, II, 470 pallide, II, 473 hensis, II, 471 ice pl oe covillei, IT uneata, Il, ts cusiclei 21,471 1 ovalis semiintegri- folia, II, 471 pallida, II, 473 II, 471 Amaltax itais: IV, 263 Ametron pedatum, a Amide gracilis, 1V, 170 hirsuta, 1V, 170 Stat modenia, TL, 152 pests II, 154 peploides major, Amodi oregona, T¥;2 Am heat scorzone- raefolius, 1V, 563 woodtae, III, 607 prelate III, 607 prs: IIT, 606 , III, 606 elomerata, “ee Tried ei orio ran ndifiira’ in oe 4, IIT, 607 fe ller howell, III, 607 end! Ill, 607 media, III, 607 poe edia eastwoodiae, III, 607 ITI, 606 n heen: ut, 04 lycopsoides ITI, 603, 606, 607 i munzit, I III, 3 tomentosa, ITI, 368 AMYGDALACEAE, it 465 , este cnet, II, 466 fremontit, II, 466 ANA pies age IIT, 50 Anacharis canadensis, a 103” qe IV, 4 a Anatallis, TE 333 pitt, 333, 004 pees < coer, III scasae decumbens, HES Anaphals, nae 484 IV, 485 “occidentalis oy. 485 , 485 Cae IV, 485 wading: IV, 485 Ancistrocarphus fila- ineus, 1 8 Andro ocera lobata, ITT, 681 rostrata, III, 681 gers tag axillaris, III bracteesa, TAT 325 FE; 337 filiformis, III, 337 maxima, III, 337 occidentalis acuta, 3s Peter iar I, 410 nelsonia eurycarpa, = Anemone, IT, adamstana, cy is 3 quinquefolia grayi, 1y,19 lyallii, II, 193 canbyi, III, 272 genuflexa, III, 272 hendersonii, III, 269 RMAE, I, 79 Anisocarpus Sibeniiers IV, 165 madioides, 1V, 166 fan is, EV: ~ ran PIV.1 vosemitanis Iv. be Sas socoma, IV, 5 pence "TV, on a senda argensis, II, 543 argyraeus, IL, 543 brach cristata digi IT, 110 triangularis digitata, III, 110 shoe nanan geen IIT, doi ne asiana, es 194 longiflora, III, 195 pallida ee 195 Anoplantis piece IV, o poner mensiesil, IE. oi Ante a. IV, 474 alpina media, 1V,478 478 oe Bidar oer IV, 478 475 anaphaloides, IV, angustifolia, IV, 480 carpatica lanata, IV, 478 —_ IV,4 oncolor “ 4 IV,4 ra sponte IV, pis dioica corymbosa, kernensis, IV, 480 Moose IV. 480 INDEX OF SCIENTIFIC NAMES geyeri, IV, 478 gormannit, IV, 478 latisquama, IV, 484 eucophae IV, 478 ape daronty Tv, i argentea a, 1V oblanceolata, ie 476 pt atin sub- alpina, sti a =00 wn I marginata, IV, 482 l Vv sols ~— - IV, 480 nardin asglerm pe IV, 482 oblanccolata, IV, 476 parvifolia, IV, 482 pedicellata, IV, 482 ‘gee > eal rhodantha, IV, 482 aclV.4 pei IV, 480 Fiigoetigreis el 480 cabra, IV, 4 solstitialis aa oph tomentella, IV, 478 mbrinella, IV, 480 WFHEMIDESE IV, 387 As Anthemis, IV, 388 slsiitea IV, 389 arvensis, IV, 388 tinctoria, IV, 389 Anthericum ossifra- gum, I, 372 sation m, I, 430 Antheropeas, 1V, 230 Antho opogon simplex, Tit Ant ia. 1423 scandic pain IIT, 22 Anticlea Po fre meee fe 375 Antipas opposit folia sp pie b eae va 788 rrhinifior pes , 784 sppendiculatum, asarina, Pak 783 breweri, III, 790 nite lium, III, 790 canadense, Ill, 786 ee ertiflorum, poh Se III, 783 venosum coulterianum, IIT, 790 appendiculatum, I nevinianwm ; III, 790 reuttian em arginatum, 111, 790 glandulos ae 789, ad beten um, IIT, 783 ie t, III, 783 kingit II, 791 leptaleum, IIT, 790 linaria, III, 784 majus, ‘III, ret 789 mau spear paces orum, III, 4 m, ITI, subsessile, III, 791 orcuttianum, oo 790 ovatum, III, 79 putianad III, ong is subse ua Bene III, 778 pies Ill, 790 bolanderi, III, 790 breweri, III, 790 rimorum, Ill, 790 -rillo- tolyenlatene, “TIL. 790 breweri, III, 790 virga, Anulocaulis annulatus, +. A ey borealis, 556 Apargiinm, oh ae ée,IV,5 ‘ pein ts SL. I, ~ ra spica-venti, Apeton opis alni- folia A phanes sBogORae II, occidentalis, IT, 445 Aphantochacta erilis, pate: serrata, IIT, 26 Aphyllon arenosum, 657 IV,8 californicum, IV, 6 comosum, IV,6 cooper IV, 8 fasctoule tum, IV, 5 faicdeaiere ‘luteum, IV;5 nia Sib 5 widatase pir Soca. hh 6 sedi, IV, * idiavasen. iV; uniflorum occidentale. violoceum, IV,6 Apiastrum, III, 222 1, III, 230 IT, 228 Bie ~ Apleappus vent be ranted IV, ee haenkei, lV, 3 sthacrocephalus, IV lari tes IV, 336 androsaemifolium, III, 369 i oe Ill, Ng II, 3 Hinder, III, ie arcuatum, IIIT, 370 oe ae Al bicolor bola ner fe Se brewe calophsiluin, TIT, 370 uksdortii, 11 369 cardiophyllum, Br: 370 rei ne ium, III, 370 III, 370 oribundum, III, 369 ype ree Sorc salig- num, 369 = num, OTT. 369 — a 369 luri ERI 3 feb Sone: ae lg iun " oribundur Vv coil to m, ITI, 369 polycardivut, III, 370 658 [Apocynum] pubescens, III, 369 nneeserg Ill, 370 milum, III, 370 rebicenambeise: III, 370 rhomboideum, IIT 370 penryg tan ium III rubi ara III, 369 ealignim /I, scopu ulorum, IIT, 369 sibiricum salignum, III, 369 Sane? III, 370 ee an! oO 80 ea flavescens, II, 179 californica, II, 180 — ¥ =" osa ig 180 cens, II, 179 formosa Il 180 ’ ambigua intermedia, 1i;3 angustifolius, ~ Aad INDEX OF SCIENTIFIC NAMES deserti, II, 305 drum umondii, II, 307 6 pu iene pen II, 307 glabra, II, 302 xisocoratvule II,-311 hirsu s, II, 629 subpinnatifida, Th=3t1 beckwitht i, II, 311 E, z, I, 345 Aragallus acutirostris, II, 6 io sid II, 613 ARALIACEAE, III, 213 uva- ursi, Ill, 310, 312 Arceuthobium, I, 529 abietinum, I, ae americanum, I, 5 campylopodum, > — tsuge nse, 1,531 —— ngelica gmelinit, 269 percents, III, 269 rad TE, 244 Arctodracon camtscha- v acuta, arguta diversifoli ia, tit, 3 otiehouba: TIT, 322 campbelliae, Ill, 323 candidissima, III, 319 canescens, III, 3 onomensis, ine ao a, III, 319 cleveland, III, 308 columbian “TIL 312 320 cra III, por a, III, 3 coneeniiaaee od 325 crustacea, IT; $23,:325 rosei, 1,3 tomentosiformis, III, 325 cushingiana, III, 323 domahiere. III, 31 oo elegans, ca pera TH5323 Siena TE 323 hingiana, — 323 howellii, III, 3 mollis, III, ei zacaensis, Ill, o23 —_— , III, 326 micola, III, 326 a, ILI, 322 imstitarias , 316 -P be ns, = 317 323 ns —— pacea, rns III, Stes Fer Preraes ae ia 314 regismontana, iil, 322 rosei, III, 325 minus, IV, nemorosum, IV, 508 Arenaria, II, 145 aculeata vy a biflora, II, 1 brevi ‘folia eaeine , 146 burkei, ¥.15] californica, II, 146 art ty II, 150 aris ursina, # cephaloides, IE, 152 ompacta, IT, 151 gregaria, II, 147 oblongifolia, II, 154 INDEX OF SCIENTIFIC NAMES obtusa, II, 150 obt palustris, II, peploides major, Th; propinqua, II, 147 II, 145 150 ursina, propin pubescen au. 147 Pht ber ihe I, 482 Argemone, II, 230 c 3 intermedia corymbosa, II, 230 mexicana, II, 230 unita ARISTOLOCHIACEAE, Armeria, ITI, 344, 345 andina californica, III, 345 arctica, III, 345 II, 409 racia ‘armoracia, IL.2 Arnica, IV, 418 piperi, IV, 420 amplexifolia, IV, 420 angustifolia eradiata, IV, 422 aphanactis, oe 423 8 ‘ > — wo SBSs te 18, 419 chamissonia interior, Pgs folio IV, “419, 420, 424 foliosa bernardina, incana, IV, 420 jepsoniana, IV, 420 chandleri, IV, 424 — IV, 420 ordifolia “TV. 421, 423, aa 426 eradiata, IV, 4 humilis, IV, hh 42 macrophylla, IV, 424 pumila, IV, de 425 cusicki, IV, 4 discoidea, IV, Oe alata, IV, 426 eradiata, IV, 426 hirticaulis, ye humilis, IV, 4 eee ,4 latifol IV, 421, 423, 424 gracilis, IV, 424 viscidula, IV, 421 longifolia myriadenta, macrophylla, IV, 424 eacecdeng™ ea, oe 423 pS IV, on mollis 659 V, 420, 421, 424 0 sororia, IV, 422 iae, 14, 426 22 viscosa, IV, 426 Aromia tenuifolia, IV, 24 Arthenatherum, : fade avenaceum, | elatius Artemis ia, ee 6, 8, 403 wormskioldit, V, 410, 411 californica, IV, 408 s, [V, 408 caste: IV, 410, 411 pponacehhela: IV, 41 rv, camporum, IV, 410 IV, 414 bolanderi, IV, 414 candicans 405 c tilis, lV, 404 Fda IV, 414 desertorum, IV, 412 dou apes IV, 410 scouleriana, IV, 410 660 [Artemisia] i iccie IV, 408 ee ,1V,4 nap halodes, IV, 405 heterdbhs 404, 4 tacomste, IV, 405 kennedyi, lV, 406 latiloba, IV, 405 letbergii, IV, 407 wetiniae IV, ney erg 5, 406, 407 dougie IV, 406 gnaphalodes, IV, 405 incompta, IV, 405, 408 latiloba, IV, 405 typica 05 matricarioides, = IV, michauxiana, IV, 408 ht Sie IV, 405 nor Vv. Recah ne! 404 a, 1V erat sys pac Paige wv, 412 ,1V, 4 seams - 412 pycnostachya, IV, 412 pu agile: rv. 406 INDEX OF SCIENTIFIC NAMES unalaschcensis, IV, 406, 407 ss coe I amgustifaliy ‘ i arbuscula, ig 8G WV pi eh ay ae 12 rothrocki, ae 414 ence acumina otus. Il, 413 sylvester, II, 413 vulgaris, IT, 413 Arundo, I, 182 arenaria, I, 162 pers tenets 2 = canadensis, I, do nies sy 182, fet fes a, 12197 phragmites, I, 183 vulgaris, 1, 183 4sagraea ee if;s ~~ III, 783 ntir chiniflora: SCLEPIADACEAE, Tit; 372 Asclepias, III, tabe 378 albicans, III brevicornu, HE ae Bese Pil aed re. 378 cordifolia, IIT, 374, 375 cryptoceras, on 375 davisit, IIT, douglasii, fr ir a, LIE S75 pel sini Ill, Sif erosa ees 377 obtus: fasciewlaris | ASF ag fren 1, WE STF Loreto 7 latifolia, IIT, 375 leucophylla, Ill, 377 yale se a es vay mexicana, III, 377 upotactnttues. LH, 378 vestita, III, 375 parishii, III, 377 viridis, III, 378 Asclepiodora, III, 378 dec III, 378 Aseyrun ‘crux-andrece, Tih, 1 Peni oi6) 1, 452 officinalis, I, 452 Aspera spica-venti, 1 Asperugo, IIT, 609 procumbens, IIT, 609 a, IV, 25 Aspre gist aa 13255 Aspris ke pat llaris, I, 166 secnsronct I, 166 x, I, 166 Amveyihitise III, 372 acrddenta’, IV, 284 case ndens, IV, 320 fr it 320 P arryi, [V, 318 yosemitanus, IV, 320 adsurgens, 1V, 322 alpigenus, IV, 328 ee IV, 328 aydenii, IV, 328 mae IV, 328 oboe us, IV, 311 iple rifolius, ry a amplissimus, IV so pile ipen ged TV, 315 2, 1V,29F habevencte IV, 314 bellus , IV, 332 bre vibracteatus, FV33 benleenacunea: TV, 320 brickellioides glabra- tus, IV, 324 chilensis, ravi os os visheum ans: IV, 288 ciliomarginatus, TV; 318 Piles aes ne 316 commutatus crassne lus, 1V, 322 pence EViS1S c V, 28 u aoEV OF cusickii lyallii, IV, 318 cu abaies: IV, 290 curtus, TV, 310 def us, IV, 322 delectabilis, 1V, 320 I al a =. a & — fiveeoud: IV, 320 eatonii, IV, 319 lau Ren hae ledophelins, IV, fe paucicapitatus IV, ensatus, IV, 316 ericacfolius, TV;.332 tenuis, IV, oe ericina, IV, 286 ericoides, IV pes fF eriocaulis, IV, 318 exilis, 1V, 330 gustralis, IV, BM capus, I faleatus ea dhlae IV, 322 filaginifolius, foliateus, IV, 318 apricus, IV, 318 he. esperius, IV, 316 ban IV, 318 IV, 318 floss, IV, 319 ? ines IV, 318 glaucescen gla neopeaiee Ww. sé glossophyllus, 1V, 334 gormanii, IV, 326 latahensis, IV, 316 INDEX OF SCIENTIFIC NAMES latissimifolius sero- tinus, IV, 294 ledophyllus, IV, 324, ‘' lentus, 1V, 3 lepidus, IV, oo leucanthemifolius, Ves leucelene, IV, 332 linearifolis, IV, 283 gali, IV, 313 m I 4 mohavensis, IV, 336 molli. re m ponies ey columb sy? va Liv. 3 multradions, ahs 296 7 oO NS I $:5 delectabilis IV, 320 rme ediu BTV, 320 a 20 paladiedea. ‘ly, 318 ansus, IV, 322 parishii, IV, 290 paucicapitatus, IV, pe IV, vd ii resin osus, I" latifolins, IV, = sibiricus, 1V, 3 Seces: ad Pan IV, 314 siskiyouensis, IV, 324 piers ong EV. S10 S, TV, 322 sparlatus IV, 320 s, IV, 32 rmalis, 1V, 330 roimieants, IV, 294 us, IV, 323, a eg torreyi, IV, 3 tortifolius, Vv fae funercus, IV, 100, 336 tweedyi, TV, 318 umbraticus, IV, 319 unalaschkensis major, Vv. Py tee ‘ve tee re be a v scion, W, 298 IV. 3 ende ersonii, ri, = seston s. 11,6 adsur, ae TIL. toy agninus, IT, 597 albatus, II, 586 sreeud hte. Il. 578 —_— ai re arre conn eA 595 leibe , 594 arthur ar asymmetricus, II, 579 atratus meéensarus, TES ouyheensis, IT, 594 oral y uae 594 sustinie IJ, 609 austrinu , 606 be chewithé weiseren- ruceae. IT, 603 californicus, IT, 573 661 calycosus, II, 600 rise serotinus, II, canadensis, II, 607 ande larius, II, ae claris ineus I ¥ 579 ox ytropidoides, TE, consectus, II, 578 cu spidocarpus, a. 598 cyrtoides deflexu te a de nsifolns, II, 585 deserticolus, I, 582 di bei Siphnt ht en 610 pr a ermu arias STL. 5 dorvehnivies, iT 600 — jae i, II, 583 pa II, us, ‘it 597 ofenicas II, 594 i forwoodii wallowensis, franciscanus, IT, 585 fremontii eremicus, griseopubescens, harwoodii, IT, 586 hasseanus, II, 571 hendersonii, I1, 606 vest has II, 585 hoodianus, IT, 5 hooke crianus Liwesot II, 583 hornii, II, 589 tularensis, I1, 589 662 { Astragalus] howellii, II, 591 sus incurvus, II, 577 ineptus, II, 599 inflexus, 577 , 578 nsularis harwoodii, te 586 inversus, If, 571 ee kentr mee elatus, II, 568 kernensis, II, 599 laurentii, II, 574 layneae, II, 600 lectulus, Il, 578 lemmonii, IT, 609 idrie: nsec . ue subvestitus, II, 578 leucophyllus, 11, 579 leucopsis, II, 579 symmetricus, "ig 579 ipes, II, matus, II, 58 montanus impensus, 2 tegetarius, II, 568 tris, 11, 606 INDEX OF SCIENTIFIC NAMES canescens, TT, 6 oophorus, I, ee. xyphysus, II, Pik tinctu 578 pycnostachyus, II, 587 nosissimus, II, 587 60 ty daturiotts tejonensis, tener tr 603 e, II, 603 ra nHtanoides, if 603 yee 607 IL 603.” toreyi, it; nt iae, II, 5 pecans 1, 602 trichopodus, II, 582 antisellii, II, 571 capillipes, II, 582 gaviotus, II, 571 triforus morans, I, 586 meta ventosus, I vestitus iain 58 , var. miguelensis, iH, viarius, II, 578 violaceus, II, 583 virgineus, II, 587 viridus impensus, 6 +p be watsonii, II, 606 sinlayucsaines. IT, 583 Astrophia littoralis, II, 6 Asyneuma, IV, 78 canesce V,78 pre enanthoides. IV, 78 Ataenia gairdneri, HiT, 233 Gite alpini- forme, II, 590 alpinum, IT, 590 cottonit, II, 589 Sania Il, 590 wallowense, II, 589 Athamanta chinensis, Ih: 273 —_— a. oligo- stachyu 179 radicosus, 2 Pig Athyrium, I, 1 olaontes yeaa ame ene wD es cyclo. ry 19 filix- plese I,19 re poy ae 19 e451 unilateral, Il, 298 Atragene columbiana, “(ve rosseserrata, II, 194 Prtownts ris, IV, 584 pleleeey IV, 584 Atriplex 6 pepaiyi ts obtusa, confertifolia, II, 87 cordulata, II, 83 tularensis, II, 83 ssa, II, 83 elega ns coronata, II, 79 fasciculata, I1, 83 expansa mohavensis, IT, 82 f gmelinii, hasta leuc ophylla, fi 84 lindleyi, 11,79 linearis, II, 87 es IT, 82 SCUu nummular II, 8 py ropa gland ea anid III, 672 Atropis californica grandiflora, III, 639 ptstonge ee “pachystachyo, I, 641 ieia he ne III, 6 Prise Sieh 5 pet tata, III, 641 pes og ae 642 I polysta achya, III, 642 stachyoide. es, br 641 vaseyi, III Aulosperm um a tense acutifolium, Ill, 277 watsonii, ved 277 mpacta, III, 115 sidaeformis, III, 115 Azalea albifiora, californica, III, ee nao ncaina wiagie | Azale “um ali III 39 Hs 29 Azolla, I, 35 filiculoides, I, 35 Baccharis, IV, 6, 3 Bacchar rob baal IV, 384 brachyphylla, IV, "0 coerulescens, IV, 3 consanguinea, LY, = douglasi ii, IV, 386 runcata, IV, 219 anthemotdes, IV, 219 INDEX OF SCIENTIFIC NAMES mut ica, IV,2 V,218 caliente of “te ase chess IV, re 214, ahh oa gracilis, IV, 2 cle ementina, 4 Z = crassa, IV, 215 curta, IV, 215 nuda, IV, 215 paleacea, IV, 215 tener errima, IV, 215 peduaage! ats 215, 216 14.215 W,2 bakeri, IV, 2 peso Sa iv uy pauciaristata, IV, 214 thalassophila, IV, 214 maritima, IV, 218 ae IV, 216 min IV. 214, 216, on artemisiaefolia, IV, 228 artemisiaefolium Pst con yi 228 ‘perl them posh } = ere gracilis, 1V, 226 integrifolia, a 226 lanata, I brachypods, IV, 226 grandiflora, IV, 226 latifolia, 1V. "226 IV, 225, 226 neomexicana, he 222 palmeri, IV, 2 parv venir IV, oor rubella, 1V, 231 staechadifolium cali- ornica, lV, 228 nudicautis, IV, 196 Beets adiata,. IV, 196 ne IV; oecraliate IV, nh pleniradiata, IV, 19 Balardia platensis, Ir, 155 BALSAMINACEAE, III, Balsamita balsamita, IV, 396 m V, 39 alsamorhiza, IV, 106 balsamorhiza, IV, 108 0 intermedia, IV, 108 acne ver raf a 107 flocco elabrescens wv, 107 hirsuta, IV, 1 Lcsrcahals: oY. 108 neg leche IV, 108 hoo V, 106, 108, 109 Pek x ferehinthaces, Banalia occidentalis, Barbares, II, 275 mericana, IT, an pa Si eowtEy a: dolichacerbe, oy os sce, eee EAs vulgaris, I, 276 IIT, 139 aurea, III, 137 porbtatees a JIT, 335 Boris sete HE Dee III, UVISCOS. 4, i Basilima cihefeKun, II, 414 Bassia hyssopifolia, II, 90 Batanthes aggregata, III bridgesii, III, 460 663 pulchella, III, 460 pungens, III, 455 BatipaceEag, III, 11 Batidaea page dite II, 45 mesh II, 457 r, II, 457 Bates hium bak ’ , Beckwithia andersonii, It2 austiniae, II, 214 Beckmannia, I, 179 rucaeformis, I, 179 asatt a, 1,179 neulgars IT, 160 0 , 14 plu umbaginielia wlll Ben Says pens Ty 290 Bensonia, II, oregona, IL. 374 Benthamia micro- carpa, III, 607 BERBERIDACEAE, i, 216 aguifolon, II, 218 , II, 218 Berkheya, IV, 548 tases ier pA 548 Bernardia corpnifai, it, 28 Bermuilona Ca pan taee: I, 468 Berthelotia sericea, IV, 466 Berula, III, 231 posh oo I, S11 664 [Betula] alba, I, 511 papyraceca ghd el caee Si? papyrifer. occidentalis Se 512 subco fos 1, 512 uaa +, oe w iandililiels: ort it E512 BeTuLacEAgE, I, 510 scars chilentix: IV, 4 Ricweulla, Tt: 233 pT 233 Bic uspidaria involu- crata, III, 140 tricuspis, ‘TIL, 140 Biden s, IV, 1 128, 136 ellipt ptic a, 1V, 128 ch aa alg oides, I cus ibe IV, 128 ansa, IV, mandate IV, 129, 130 puberula, IV, 130 pes be rhe ke log. Ae lae lon oh. Be nV. 128 melanocarpa, Ae 129 aes 1 aes Ne pilosa, IV, 1 We Prionophyil, He 128 1V,4 IV, albi V, 296 arborescens, IV, 288 bolanderi, IV, 30 eolens aiticed lis, hololeuca, IV, 303 howardii nevadensis, INDEX OF SCIENTIFIC NAMES IV, 301 intricata, IV, 330 IV, 2 vscidifora, IV, 278 Bic i ' Big s, IV, I Bikweulla Ears sanity: 233 Formed II, 234 ettia ‘riparia; IV, Biscutella californica, Bistorta bistortoides, Tt: thi coe, TI, 61 Blechmum radicans, mice: he 248 chilenss IV,2 eee hid, Sore robus Ve Blepharipappus Vv. 192 carnosus, IV, 163 chespasitheneoulie: TVt douglasti, IV, 1 — ig hd iv 159 16 ties IV, 159 poe Aone af Sag glandu ete a Ly. heterot he IV, graveolens, IV, heterotr she a's ig hier acioides tv 1 oregan epee Vv 162 platyglossu ea thee 160 tv, "subphvmoss, Vv" 192 ida, IV, 192 Bane pM 2 m, Ii 72 cere Ti #i atum, II, 72 hcacbedindee II, 74 glaucum, II, 71 eo tegen II, 74 rubrum, II, 71 Blondia Sifeibite, Bloomeria, I, 398 aure cleveland, I, 399 croceo, I, 398 at Sg I, 397 serotina, |, 397 Boebera chrysanthe- motides, IV, 252 hirsut intermedi 1, 103 ia ewhe IL 103 3 ramulosa, II, 1 repens, II, 10 rel a pee t iq tra californica, 50 Hieahasedes iy 350 oregana, IT, 350 Bolbo re devant, TII. 8 Bolelia bicornuta, IV, 90 wpe eaters : 94 r,1 IV, 3 siaanitolia IV, 304 latisquama occiden- V, 304 occide ntalis, 1V, 304 Bonacixaceae TEE "S32 Borago, III, 540 aiheinaite: TIl, 540 Borkhausia lessingii, IV, 566 BorrAGININEAE, THT; Boschniakia, IV, 9 0 occidentale, I, 4 Botrys ambrosioides, Ii 72 aromatica, II, 72 fremontu, II, 71 esa aes II, 70 ie Pichia i 176 arenosa, I, 177 aristifpides: E177 b 8 palmata, septentrion ali is, Ett. Boykinia, TE351 aconitifolia, II, 352 elata, II, 352 humilis, 111, 164 cidentatis; UT, 352 lat a, ET; 352 cotumditolia SIE S52 ai vo eo ctis angustus, V; 332 side sa, B isugstanig fies: Brachyphragma braun- phos TI, 597 serenot, II, 595 Brachyris californica, IV, 260 euthamiae, IV, 260 microcephala, IV, 260 Brandegea, IV, 72 pe ipestlies TI, 172 schreberi, II, 172 Brassica, II, 274 a II, 274 271 arve msis SAE, 274 campestris, IT, 274 71 gra, II, 274 oleracea, II, 274 ety iy vi 321 ra SSinapistram, II, 271 ae ah 237, 6 , 207. pi IT, 302 reweria snininia, III, 389 Bremerina suffrutes- cens, II, 152 Bekele IV, 500 oe Paige pe wee We bee bicacilokine IV, 502 Zu mu ra, I nevi IV,5 blongifolia, IV, 504 gracilis, I, 404 INDEX OF SCIENTIFIC NAMES pen arre-i> po 405 majo Biss ino ete he wor I, 404 ee I, 402 acinthina, I, 400 Ait i e4. 409 paar ie! lugens, I, 399 0 breviaristatus, I. ; brizaefo ‘ 123 carin atus, I, 2 ealifocnicun, 7 228 — SE 22 hookerianus, % 228 linearis, I, 2 lane aot E23 paucthorus: 1; 230 Ss sus, 1 23 commutatus, I, 235 run; 1, 23 labrescens, 1, 2 leptostachys, I, 232 inermis, I, 229 japonicus, I, 236 — “: aristulatus, i Sy scented 231 1,231 hallii, I, 231 pacificus, I, 231 patulus, I, 236 S; pratensis, I, ren a bs esdirde I, 230 p unioloides, I, h I, 627 Bryanthus breweri, IIT, 303 pri hee ITI, 303 iflorus, III, 303 Buillardia aquatica, ie Bulbostylis annua, I cali ifornica, IV, 5 adithanelte. IV, A Bunias Cakile, 11, 270 Bapsihaln ses sagitta- IV,1 ielia chrysostoma 4 : song lept IV, ae 216 LVe21 a u ie bursa-pastoris, i;2 pastors, II, 291 ra, 111,20 pace 4:11), 2 microphylla, III, .. Cacaliop IV,4 Cal albicen 665 BurRSERACEAE, III, 2 ee occidentalis, i IIT, 45 Buxus, III, 45 chinensis, III, 45 Cacalia nardosmia, IV, 457 po orophyllum, = sis glab nordesmia IV, 457 Calais aphantocarpha, IN; bigelovii, IV, 561 densa, I, 159 fasciculata, I, 160 666 es ne tn folio: a, 1,15 elongata, I stenodes, I, 162 inexpansa, I, 162 barbulata, I, 162 cuprea 157 aco ouniana, I, 161 mis, I, 16 anes I, 159 oregonensis, I, 161 tte I, 159, 161 purpurascens, I, 158 retrofrac eta,’ T; tg rubesc Se stricta robusta, 1 Bee subflexuosa, I, eubedorf, 3 - s, I, 160 Persehdhy Seaiotiié (€ purpurascens, I, 158 ai , 158 vase vurietonnia: 1; 159 Calamintha chandleri, III, 646 Hil rot C allichroa. douglasii, 1V;,1i jones, IV, ae Matjaivisa: v. 160 INDEX OF SCIENTIFIC NAMES Calliglossa douglasit, IV, 158 Calligonum canescens, II, 87 Calliopsis atkinsoniana, bicolor, tV, 135 er invotucrata; one ey en 102 Callistachys: pyrenaica, I, 289 C allisteris aggregata, IIT, 460 bridgesii, IIT, 460 seleaie8 lia, ITI, 460 Cal litrichaceae, III, 42 Callitriche, 43 heterophylla, IIT, 43 longipedunculata, Ill, marginata, III, 43 lon gipedunculata, II palustris, III, 43 bolanderi, III, 43 na, , ipa a IIT, 43 ver a, III, 4 Caloboirya sanguinea, Ce al IV, 554 exuosus, I, 443 i 38 kennedyi, I, 440 pete I, 443 lilaci ns Pt 37 lobbii, 14 lo vai asdk I, 438 luteus, I, 444 9 carpus, I, 441 ma aes #4 & 441 438 eurycarbus, I, 439 sprue panini 437 wasttailit purdyi, I, Siaateusts: I, 437 speciosus, I, 440 roseus, I, vesta, I, 5 wiedit 445 obispoensis, I, urpurascens, 1,445 p C alerherant californica, 158 ro i EF, 475 asarifolia, rt Body sri! 11,3 howellii, Me ae piace tit +f ti tye de “howe Th 1 undifolia, Hy 176 , 1V, 189 elegans, IV, 190 fremontii, IV, 190 rosea, I 88 gi ar tae 1 as bicolor, TV; 4 i ose We 189 18 IV, 189 opposititolia, IV, 190 truncata, IV, 186, 187 ephala, mtv. scabrella, we ete villosa, IV, 1 Cavvenvrnacear, | ae Calyeanthus ¥E 221 §, 10, 224 Ot aiaie 4,221 oer montanum, 14 Calyoseris, IV, 582 V, 583 wi ins i, IV, 582 californica, IV, 582 I, 482 andru Ftd monospermum, IT, 130 nudum, II, 130 parryi, IT, 130 pygmaeum, II, 130 piety yor If, 131 ote TT, ase sale fis lum Calyeteta sa itt ail III, 382 sepium pubescens, TE: repens, III, 383 soldanella, III, 382 subacaults, cae 384 villosa, III, 3: Camassia, 1, ae sativ. a, IV, 76 56 tabiforaien, IV, 74 exigua, IV, 76 filiflora, IV, 78 las iocarpa, IV, 74 parryi idahoensis, IV, 76 perfoliata, lV, 78 —— ,1V, 74 piperi, IV, 76 pr ronanthoides, = 78 é,IVi7 CAMPANULOID Canbya, a, 232 pein nai Canotia holacantha, III, Cantua aggregata, I, 460 pungens, III, 455 hookeri, III, 455 Pease IIT, 455 4 237 caseanum, II, 235 ouleri, 11, 235 nse 11,235 Capnorchis chrysantha, , Capnorea campanulata, fulcrata, 111, 529 panberaen: III, 529 a, ITI, 529 IIT, 529 It; 3223717, Capraria erdtioibéitee, itt; CAPRIFOLIACEAE . IV, 42 Caprifolium ciliosum, ives Peiedte aren 1TV82 hispidulum, TV, 54 indorPiabiie, IV, 54 involucratum, IV, 52 arara en a, 263 INDEX OF SCIENTIFIC NAMES Cardamine, IT, ~i angulata, II, 2 pen arene: — 280 acu minata, pr dcuminate, IL. sgeatatieices a 280 ts, II, 279 palustris occidentalis, IT, 280 peer ii 2 nsy Ivanic a; 4 om prone hemtithatico, I, 280 peek EI 282 vallicola leBergit; 1i:2 pingauicis At , 305 Cardaria draba, II, 258 pubescens elongata, Cardiobatius nivalis, nema, TI, 114 , 1V, 540 mart lanes, 8, TV. co. 510 maritim S, s, IV, 528 avensis, 1V, 521 ne. cei IV, 540 nevadensis, IV, 524, 525 nutans, IV, 538, 539 occidentalis, IV, 525 pycnocephalus, IV, oe remotifolius, 1V, 5 pe ms congesta, I, minor, I, 3 aequa, I, Aa abide; 1325 albonigra, I, 331 llacea, 1, 343 angustior, I, 300 angustata verticillata, I, 335 anthericoides, 1, 293 aperta, I, 335 poda, I, 331 aquatilis, I, 336, 339 areta,-1,;315 aristata, I, 341 at airatiforuiés lL; —_ squama, I, 331 rea, I, 322 pert oe auricu lat ais ne 1337 i bisha iii bla bru eace ae boxieuant! T, 329 667 alifornica, I, 323 Tubia, I, 314 Peeing ar: 1,315 pitata, I, 290 ce a 1.322 cephalantha, I, 301 chrysoleu cinnamomea, va 325. circinata, I, 2 columbian i a como peace 12321 congdoni, I, Fest cooley LL 3 crawford ry 303 I, 303 erspiocorba 1,339 de a bootii, I, farwelli, I, 320 rossii, I, sa; I, deweyand, I, 301 olanderi, I, 301 sparsiflora, I, 299 diandra, I, 2 dives, I, 336 disperma, I, exsic 3 farwellii, 1, 320 festiva, I, de cum haydenian a1; 309 packystachsa, 4.344 stricta, ist flaccifolia, I, 323 flava recterostrata, I, 341 668 foetida, I, 291 vernacula, I, 291 fracta, T, 303 frigida, I, 326 fulva eee ona pie T 325 a I, 344 307 hartii, I, 343 hassei, I, 322 oe > 309 helleri, I, jamesii, I, 334. onesii Poser as laxiflor plontaginea, - 323 lem oe lenticular Ihfru ctus, 1 as I baton e 31 luzulaefolia, I, 326 INDEX OF SCIENTIFIC NAMES strobilan tha, I, 326 5 ony endocinensis, E325 pee si, 5,332 michauxii, 4 318 ” vic rochaeta, I, 328 mult icostata, 1 i multiflora, I, 2 multimoda, I, oe mutic nardina hepburni, I, 288 nebraskensis I, 334 nigricans, I, 290 nubicola, I, 309 nudata, I, firmior bnupt oederi, oregonensis, I, 340 oO ha, I, 300, 557 1 poddvenss, I, 289 pansa, pasicicontats I, 334 aucifiora, I, 342 pancifieccds, 7,313 eocephala, I, 307 ainda I, 300 Deere aoa I, 324 a fe 328 polygama, I, — polymorpha, Eg californica, I, prs hie trichoides, . ni S52 4 pse eudo-jaronica, I, 326 weer rpoidea, pico I, 282 aica, 1,2 89 pobre [322 quadrifida, I, 330 caeca, I, 330 nis, I, 330 qualicumensis, I, 339 —— : 329 343 richardson 13325 320 gerne I, 320 rostrata, I, 343 salina — . 321 salin rmis, I, 321 _— vlna, 1, 339 ximo a, 13S orn 1,317 pre ae I, 303 min or I, sco evi-fructus, I, 303 Pina ], 328 sphacrostacha, I, 314 327 sten isan, % 318 a, 5,291 , 304 straminifarads; 305 stylosa virens, I, 332 sub- practhahe 1348 ka isca, I, 310 erecta, I, 304 teneraeformis, I, 309 tenutrostris, I, 302 pprinienboe I, 297 la, I 297 pitino I, 33 umbellata globosa, I, 319 unilateralis, I, 302 urban nit, I, 341 a1; 292 mri er I, 343 vallicola, I, 294 variabilis, 1336 ve see I, 319 91 vertcilata 1 a bier sic eee osa, ni ee nchale I, 342 major, I, 342 obt rusian I, 342 a 1;319 vitrea, I, 29 viridula, I, 341 vulgaris alpina, I, 3 bracteosa, *: 333 ‘ee rpa, whitn 324 ee ee I, 318 yosemitana, I, 339 Carnegiea, III, 153 gigantea, 1E1, 353 Carpenteria, II, 386 californica, II, 386 Se by trae junceus, 5 arpobros chilensis, 19 lana 3 EY; si, 512 ieee EV,5 tinctorius, <. ar 512 Cartiera barbata THe? howellii, II, 248 lepto petala, II, 247 multiceps, II, 247 Carum, III, 231 carvi, III, 231 erythrorhizum, III, 2 gpetor latifolium, III, ‘dares 112233 lemmonii, III, 234 kelloggti, III, 233 oreganum, III, 234 YE E, Caryopitys mono- phylla fe ee Cascadia nuttallii, 358 ata, I, pio ae gery I, 515 15 ervirens, oe "i Castile III, oe sy IL, fee ome 836 iariefaia, 844 andrewsii, III, 8 pi aoe Lipo abbrevt- on sohhon ra, aa, 837 Cc err shee III, 824 brevilobata, III, 830 breweri, , 832 dixonii, III, 840 dolichostylis, I11, a8 douglasii, ITI, 84 contentiosa, Ill, tie pee 5,31; 841 w& © o meri, III, 8 pb ee ITT, 836 i, III, 837 gene: III, 837 INDEX OF SCIENTIFIC NAMES exilis, III, 846 filifolia » HT, 825 eed grisea, III, 8 gyroloba, “i 836 hispida, III, 8 abbreviate i, 841 III, 8 bg mannii, a. 837 hololeuca Shik = 42 howelli hyetophila,t in, ayer inconspicua I, 825 a, III, 835 abit Ill, roa insu s, III, 84 Pape III, pinnatifida, III, 836 wightii, III, 836 lemmonii, & eS 82 pallida camporum, lutescens, IIT, 832 muiniata, III, 839 septentrionalis, 833 foe nosa, ITI, 829 pa peed a 824 peckian: I, 840 erie ae 832 pilosa, III, 825 pinetorum, III, 837 II, 837 psitta a, III, 825 pumicicol, re 824 r , eis! Ill, ee 37 rubi da, III, 828 schizotricha, III, 824 entrionalis, 23 spiralis TIT, 8 , III, 836 xanthotricha, III, 832 Cathartolinun digy- m, III, 12 erulum, IIT, 12 22 austromontanus, califorwicts: 1H, 73 confusus, III, 78 669 cordulatus, 8 gii perplexans, TL 7 oe III, 75 gri s, III, 6 68 ho micantbiee: III, 68 utus, III, 67 prostratus. pinetorum, III, 79 profugus, III, 76 670 [Ceanothus] Cenc E carolinianus, I, 119 echinatus, I, 11 centau uriu 540 ine squarrosa, IV, 544 Centaurion centaurium, 1H,:352 exaltatum, III, 354 muh, bergii, FIT, 352 Centaurium, III, 351 INDEX OF SCIENTIFIC NAMES curvistamineum, Ill, 352 davyi, ITI, 352 erythraea, III, 352 ee Ill, 354 wei 35 a on — minimum, IIT, 3 muhlenbergi Conteuriiies Pontius: dum, III, — yonseretnas gti, TIT, 352 ichanthun, III, a ae Cen woubalie Seed tents IV, 180 . C. entrostegia leptoceras Pes tharbe o. II, 1 4,11; oa ae ao 334 minimus, III, 334 Ceph clnithavs austinac, I, 478 poset I, 478 SM oy —s TI, a @, 7153 maximum, - 14 brachypodum AI; e 3 longepedunculation, , 14 nutans, at 43 ech Patan, II, 143 142 CERATOPHYLLACEAE, II, 174 Ceratophyilo, II, 174 174 Cercidium, II, 4 floridum, II, 4 5 betulasfolins IT, 453 blan II, 453 betufaies, Tl, 453 douglasii, II, 453 forherslvies oe 453 II, 4 ninutifloru rus, I ae Cer _ pap a THT, Sicoteots TEL 485 ganteu is mojavensis, III, 155 munen, III, 155 Cer ophyllum douglasii, I, 391 spathianum, IT, 392 Ceropteris triangularis. viscosa, — Cha perce a 240 pi ge IV, 238 alpi dngestilelid, IV, 240 artemisiaefolia, IV, 242 Pine pions 246 a,I romhastie Sg tv, 246 bracteata, IV, 2 carphoclinia, oo 246 attenuata, IV. 246 cheilanthoides, achilleaefolia; V, 240, 24 fremontii, IV, 245 glabriuscu IV, 236, 244 aurea, IV, 244 curta, IV, 244, 245 uttiana, IV, 245 patielef dren IV, 246 panamientensis, IV, 240 santolinoides, IV, 237 indurata, IV, 327 stevioides, IV, 245, 246 brachypappa, IV, 246 Bona ne 238 incana, I epee is wate ta 7 a orden nn 244 tenuifolia, IV, 244, 245 thompsoni th vtec xantiana, integrifolia, Chaeroph Sauer neo V, G hectanthird aster- oides, IV, 266 gor Rare I, 118 californica, I, exilis, IV, 268 ene IV, 268 nii, IV, 266 a hamachatia, II, 451 alis, II, 451 amaenerion angusti- —— IIL,3 latifolium, UL ie Chamaeper sshecnoan unalaschkens III, 284 canadense, III, 284 Chamaesaracha, III, 668 nana, IIT, 668 —— III, 670 Cha —— mar- gi ey Ff Ch esheets chamo- V, 399 “IT, 318, 319 " thotdes 3S perennis, II, 318 INDEX OF SCIENTIFIC NAMES suffrutescens, II, 319 — hegre II, 318 ra hoid 15 yg ag: ‘glaucium, Brat nemorosa, III, 769 CHENOPODIACEAE, II, 66 Chenopodium, II, 68, 74 album leptophyllum, II, 70 rida, II, 70 sebrmidan Hi; 72 anthelminticum, hastatum, II, 72 berlandieri II, 70 botr »@2 lifor rnicum, TE 72 hybridum, IT, leptophyllum, IT, 70 subglabrum, II, macrospermum m halo- sani farinosum, Li maitidum, II, 74 oe 70 prac um, II, macr crospermum, a 71 I, 95 phila, ‘III, on Pete se Hi2 occidentalis, sa = umbellata occidentalis, II, 2 Chionanthus Fnpuies este Ill cage cali- > rnicum, III, 114 G — scaliate, 355 : 9 Gc ee austinae, Ci hioris, I, 176 curtipendula, I, 177 elegans,1,176 ° angu leichtlinti, I, 416 parviflorum, I, 414 pomeridianum, I, 413 purpureum, Chloropyron canescens, III, 849 maritimum, a 848 ae III, 8 palustre, iin, ‘4s —_ i, III, 8 Chon rophyla Sti can Fremont III, 359 NaUUSE osa IV, 302 “ hondrosium gracile, 179 folystachyo, I,178 CHORIPETALAE, I, 485 C quar nevadensis, V, 142 Chorispora tenella, Chorizanthe, II,4 spinosa, II, 1 staticoides, IT, 6 671 latiloba, II, 6 brie er II, 629 villosa, II, 6 simiintes Il, 4 eat II, 14 t 4 ntii, II, 7 leucotheca, II, 7 CG hrossostephium cali- fornicum, IV, 408 foliosum, IV, 408 Chrysam morpha, II, 329 californica, II, 329 Chrysanthemum, LV::3 ancthifolium, x 394 balsamita, IV, 3 _tanacetoides, mie 396 ina whe 394 maximu pith mV panthanies av, 396 IV, 3 segetum G hryshibis Me Ii 32 aarbore escens, IV,2 icmp IV, 288 cooperi, IV, 288 egg sbracteata. 265 boner IV, 278 californica, IV, 264 gracilis, 1V, 265 672 [Chr ye nig is] hir , 264 hirtella ae 361 II, 367 pean 13 onpostfoli m, econ 11-367 een douglasit, IV,2 gnaphalodes, iV. eo gramineus, IV, 300 greenei filifolius, 123 humilis, 1V, 299 lanceolatus, IV, st per IV, 2 ermus, a ie sega is, 4V 303 monacePhalus IV, 301 nanu oe Jeers 302 " pibiceutis, TV, 302 V, macrophslns, IV, 303 ospermus, IV, 303 mohavensis, IV, 303 occid wn hide: IV, 302 oreophilus, TV, 303 speciosus, IV, 302 INDEX OF SCIENTIFIC NAMES viridulus, IV, 303 viscosus, IV, 303 nevadensis, IV, 301 gar sa — 30 ccidentais, IV, 302 2. TV,303 or. rthophyilus, gs 302 paniculat us, IV, 297 cus, FV;3 301 ok 01 Chyli —- brevipes, IIT, poche a III, 205 clavaeformis, III, 206 oe cruciformis, ee II, 206 C ‘cede exoltate, TII, 354 Cicerbita muralis, IV, ac pie IV 3551 Ci chori EV, 552 ire MIL, sien Passed = ee 79 elata, II, jfoetida II, 179 iata, II, 179 palmata, II, 215 Cinerartia lewisii, V, 354 ula, I, 148 Circaea, III, 211 Ipina, III, 212 I ecaulescens: IV, 532 amblylepis, TV, 514, 535 americanum, IV, 532 callilepis, 1V, 534 andersonii, IV, 5 536 eiasene Ev, 314, 530, 532, 536 brewer avs califor V, 514, parishii, 1V, 514 eu candidissimum, IV, 526 canovirens IV; 514, 521, 525 ciliolatum V,514 518, 520 coulteri, TV, 525, 526 crassicaule, IV, cymosum, IV, 520, 522, Sao, oa douglasii, IV, 514, nee §32 cane edul 6:3R 528, 530 flaidilen: IV, 514, 532, 534 fontinale, IV, 514, 532,538 obispoense, IV, 538 hallii, 29, 530 hu spre IV, 25 hyaionham, — 520 marttimum, he 528 mendoci IV, ite we mohav TV, $21 secu tone EV;521 neomexicanum, IV, 524 nevadense; IV, 524 nidulum, IV, 525 id can ro aa IV, coulteri, IV, nf venustum, IV, 5 ochro contrat WV, oa oreganum, IV, 535 —— se, 1V, 518 salV¥e Sp 526 oh IV, s ceaeitial an 534 proteanum, IV, 514, _ 527 quercetoru V;514, 520, §30; 531, 532,.535,538 = 2 ee = =. 3 < : 532 EV;'5 psi ophstbit, IV, aa walkerianum, IV, 532 xerolepis, IV, ae IV, 528 re foliu odontolepis, IV, 534 oregonen. 3535 pick ie IV, 535 —— ey ite scabru soon ree ee 536 — IV, 514 fa a TV. 522 un ee ~ I cilotatim, IV, 518 utahense <3 514, 518, 524, 525 — hydrophilu m,1V, 520 florus, III, 298 Cl lodtithrt psa i II, 100 oblongifolia, I1, 100 Clarkia, III, 181 biloba, III, 189 breweri, III, 184 concinna, III, 184 delicata, III, 183 ciseniana, III, 183 e parviflora bellidifo lia gee 123 arayiony: 13 1}127 i, 11,126 ae es a 127 peer ophylla 1, 129 virginica, III, es grt ees ok 446 Ae TE 174; 193 biflora, II, 194 h ones > a = pauciflora, IL, 194 INDEX OF SCIENTIFIC NAMES suksdorfii, II, 194 ve rticillaris colum- gynandra, II, 322 cs EI 322 isom venge! ; io pinnata, stricta, IT, poner = 323 taurocranos, II, 323 eo ee incanum, 658 sminulodes, 646 pulche lla, IV, 92 pusilla, ek 92 uniflora, I, 452 Clo iadadels cooperi, IV, 252 laciniata, IV, 2 Porophsiodes, bas re ro LAciée 1,3 maritima, II, 319 ophacos know- oronopus, II, 263 oblongifolia, I1, 266 officinalis, II, 266 Coelopleurum actaci- folium, III, 269 — III, 269 gipes —— m, III, 269 Cogswellia altensis, III, 266 pasar sy et 266 carnitfolia patens, Tike255 cha willevi, IIT, 255 cottamii, III, 264 260 utricu ioe easireiet ITE 255 Coinogyne carnosa, IV 8 Co m TF brevicalyx, III, 534 c ens, III, 533 ma, IT, 454 Coleosanthus albicaulis, IV, 50 argutus, atrac tylo ides; IV, 502 californicus, IV, 503 desertorum, IV, 50 microphyllus, 1V, 504 multifiorus, TV, 500 nevinit, IV, 504 geet rete IV, 504 IV, 04 watson, IV, eee pre insia, phe 774 779 po III, ae barbat hartsaefola, III, 776 dav idsonii, III, 776 austromontana, con color 175 bracsphon, ‘TH, 780 bre 079 br ucea «Tit, 779 Jahowtl, III, 778 corymbosa, III, 776 673 davidsonii, III, 776 diehiti, III, 778 divaricata, III, 778 hirsuta, II, 776 SP pusilla, oe 778 septemnervia, — it sparsiflora, ITI, arvensis, III, sig bruceae, III, 779 collina, III, 778 franciscana, III, 776 ee Ill, 778 ricta, III, 776 colitara III, 778 te a, torreyi, III, 780 brachysiphon, ’ brevicarinata, III, 780 latifolia, III, 780 linearis, III, 780 vere) Collo mia, Il}, 400 = i] We mazama, III, eiekant Ki; I, 413 674 [Collomia] tracyi, III, 402 Coloptera parryi, TEE 277: Colubrina, III, 79 californica, III, 79 for. : laf. - 2 III, 79 Comandra, I, 527 cakforsica, I, 528 pallida, I, 5 528 ee omarobatia lasiococca, II, 455 Gi omaropsis pedata, Lis Comarostaphylis, III arguta, III, 308 diversifolia, III, 308 CG —— lustre, II, 4 Gi stokes: dubia, Composirar, IV, 8, 98 Conanthus multifiorus, III, 523 parviflorus, III, 523 Condalia, III, 61 divaricata, III, 61 globosa pubescens, , 1 tnid. TIT, 61 microphylla, ae 61 par reac aa Conioselinum, TH;:272 chinense, III, 273 docitines ‘TIT, 273 tataricum, III, 272 PS oes Ie. . f IV, 248 Conium ETE 230 ac III, 380, 474 it. Convolvulus, III, 382 althaeoides, 389 aridus, III, ntermedias, III, 384 “85 lobus, III, 384 teaeifotins UT, 384 arpa, ot 384 Resin stir TL 383 californicus, III, 384 INDEX OF SCIENTIFIC NAMES chartaceus, ee Sei tp Stirs me caaine 10 & 629 proliferum, im, il. 43 petnaatich I, 46 pusillum, IT, 34 ertoriinl: Il, 629 m1, 27 thurberi, II, 31 parishii, TE; 31 oO :& = membranaceum, subscaposum, II, 41 trachygonum, IT, 41 Eriogynia caespitosa, 7 endersonii, If, 411 ‘pectinato, it, 41 13» achillaeoides aphanac- tis 14.9 fein IV, 230 paleaceum, IV, 230 artemisiaefolium, V, 228 integrifolinm, IV, 226 INDEX OF SCIENTIFIC NAMES latifolium, IV, 226 gris cm tyra ceca chrysanthum, IV, 226 cineraria, IV, 22 6 confertiflorum, IV, 226, 227, 228 discoideum, IV, 227 latum, IV, 228 laxiflorum, IV, 226, 227 tanacetifolium, donii, IV, 228 IV, 225 jepsonii, IV, et 227 lanatum, IV, 2 achillaeoides, av. 225 IV. ae croceum, IV, 226 integrifolium, IV, 226 lanceolatum, IV, nay bov , eves obovatum arishii, IV, 230 , 228 artemisiaefolium, IV, depressum, IV, 228 tanacetiflorum, IV, 227 wallacei, he 224, 230, rie ustrale, IV per et Sg me rubellum, IV, 231 watsonii, IV, 226 Eritrichium, ITI, 557 barbigerum, IIT, 585 fe ‘UL 557 glomeratum humile, 2 hispidum leiocarpum, = z ma erum, IIT, ty submolle, III, 58 how : intermedium, TI, 585 hingi i; FH, 572 aac rpum, ITI, 594 le weophacwn IIT, 599 micranthu , 578 molle, I n, IIL, 5 pterocaryum, III, 582 pectinatum, IIT, 582 racemosum 80 torreyi, III, Erodiu m, III, 5 ae iideibin. lil;7 rys, iid 7 monta 115; 7 obt hisipabctiids EET::7 californicum, III, 5 cicutarium, assifolium, Ill, 5 obtusiplicatam, 11 7 texanum, ITI, 5 Erophila vulgaris, sativa, II, 27 Erucastrum Ces. Brews hirsutum, 615 tear se IT, 616 Radi ali smaefolium, ITI, 281 armsianum, III, 279 683 eae III, 28 axibracteum, rT, 281 packs istylum, IIT, 279 oblanceolatum, ITT, 281 par arishit, III, 282 io zmmephilum, ~ 319 asperu in eins ea “ 318 capitatum, Il, 318, 319 perenne, II, perfoliatum, IT, 321 repandum Erythraea centaurium, III, 352 curvistamineum, III, 352 dot uglasi, IIT, 354 exaltata, III, 354 flori kaa: Til, 352 minima W sont, I, 429 montanum, I, 428 multisca eum, I, 427 parviflorum, I, 427 684 [Erythronium] 9 Eschenbachia coulteri, IV, 382 Eschscholzia, Ii 2265227 caespitosa, 115227 ypecoide urceolata, II, 228 wrigleyana TI, 237. —_— - auiplexicas: cog | helontoides, at 106 Esula cypar Eclisia amplexicaulis, Hy 2 hispida, II, 250 50 = = Q: we emifolia, INDEX OF SCIENTIFIC NAMES III, 4 bipinnatfida, aa 485 foliosa, III, micrantha, py oe paniculata, III, 485 —— ovalifolia, 43 meevettaenh m, II, 39 Eulobus californicus, III, 196 Eulophus ambiguus, HE-25 americanus, ITI, 233 rt benignus, Iik, californicus sanctorus, {11,2 cuspidatus, III, 234 —— rusbyi, ITI, 234 prin nglei, ITI, 234 Bes:;.tasial, 11], 234 Ill, 727 austinae, III, 719 torreyi, III, 723 1 abramsiana, III, 39 albomarginata, III, 38 antiquorum, III, 30 arenicola, III, 36 arizonica, III, 40 arkansana, III, 3 cinerascens appendi- culata, III, 4 crenulata, ITI, 3 ranciscana, — 34 conju ta; III, flage elliformis, III, 36 floccosiuscula, III, 4 glyptosperma, III, 39 [ 9 misera, III, 32 — III, 34 sie rs aa III, 39 ton sil 4 ctentalle i, 39 a, 31,3 sli pe gece FRA iL, purisimana, III, 40 spathulata, in, 32 supina, I, st ra a III, 42 virgata, III, 35 EUPHORBIACEAE, 4,23 Euplecs, ITI, 534 — californica, lit, § ptiselaee ie LH. — pa ie a, III, 5 Eurotia, IL. ceratoides, II, 89 la eR 8 a — lucida, I, 249 Pie ri III, 249 Eustoma, III, 354 exd altatum Tit, 355 silenifoli ms III, 3 Eu eset: ‘californica, IV,2 Pada IV, 292 occidentalis, as 292 Eutoca albiflora III, 514 aretioide’s, ETI, 523 bra eliana, III, 508 a, 1V, 100 eae Mexus, II, 99 mula, IV, 496 sparsiflor EV: newt 495, 496 brevifolia, TV, 495 Bie a! So IV, 494 Exacum quadrangularis, TE¥351 Exochordia davidiana, II, 469 Fasaceag, I, 485; I, 480, 630 a, Tit, 16 haeane aI utinosa, III, 16 Fendlera utahensis, TI, 388 Fendlerella, II, 387 Penal concinna, III, 4 peered III, 424 III, 424 johnsonii a 15 rostii, III, 157 Mii: retin nt 156 pi la uifolia III, 387 dissoluta, III, 261 brevifo lia, I, 225 Auitans, I, 2 grayi, I, 219 hallii, 1, 223 kingit, oreg reflex INDEX OF SCIENTIFIC NAMES rubra, I, 223 pubescens, I, 223 am Filipendula, IT, 415 occidentalis, II, 415 rubra, II, 415 tlix fragilis, I, 7 Fimbristylis, I; 267 capillaris, 1, 267 congesta, I, cyperoides, I, 268 junciformis, I, 268 Floerkea, or 49:f alba, tube dou ae hetae occidentalis, fo 50 proserpinacoides I ro. acuminata parvifolia, neo-mexicana, III, 349 Forsellesia arida, pes evadensis, IIT, 55 aden FH, bs - tent ed glab sities, THI, 55 a, III, 474 cunei $folia: it: “442 ip eae II, irs sibbaldifolia, _ ey suksdorfi, froneatas he vesca, II, 442 chiloensis, II, 442 virginiana illinoensis, II, 4 platypetala, IT, 443 Frangula californica, III, 62 B ticinow I, ankenia, oT, 119, 394 pear) ITI, 119 ampestris, ITI, 120 evis, III, 119 latifolia, III, — palmeri, III, pulverulenta, ‘il, 120 Heirkanias CEAE, 152 sdeerile IV, 150 ubia, IV, 150 ins sularis, IV,1 chamissonis, ey i bipinnatisecta, I chenopodiifolia, I cuneifolia, IV, 150 confer iiflora: IV, 150 ten villosa, 5 Frasera albicaulis; III, 362 capes TEE ‘ei cusickit, III, 363 fasti — ate 362 neglec II, 363 spoil it, 363 umpquaensis, III, 362 gece us, IIT, 346 nericana oregona, TIL 346 anomala, III, 347 triphylla, III, 347 coriacea, III, 347 685 dipetala, III, 347 brachyptera, III, 347 6 I, 347 coriacea, III, 347 Fremontia, III, 114 ca lifornica, II x 114 crassifolia, III, 114 mex a, ITI, A he 115 nabeunis: TI obispoensis, i, 114 vermicularis, II, 94 F 425 amts ee AL I, 423 422 purdyi, nyrenaica, I, 420 va, I, 422 co a, 1/422 sectbleute. je 421 vir ee Sees Fum cnculri, i, 235 for a, Il, J MARIACEAE, £ 485; Hi; 233 piri nace m Atti. 372 ‘aowelitaen m, }, 373 gee idicort ape: II, hirtellum, ITI, 373 Gaertnera dumosa, IV, eriocentra, 1V, 151 ilicifolia, IV, 152 686 Gaertneria acanthi- DS $3 & lii, 1V, 206 pulchella, IV, 206 alardia amara, V,210 G alarhoeus cyparissias, I helioscopia, — 32 3 parviflora, IV, 136, 137 preneni IV, 136 Galliaria blitoides, ei zia, ITI, frticore, “ri ae sa, III, 783 Galina iu “Tv, 25 agreste echindsper- pincesedeshng TV..3 siski iyouense, iv 34 ardin m, IV, 38 h yssopifo Jeni, IV, = intermedium, IV,3 randegei, oft, IV, 33 Pel grande, IV, a INDEX OF SCIENTIFIC NAMES grayanum, IV, 4 arate Ae 42 hallii, IV, 3 h Sieienuee I 0, 41 subalpinum, IV, 41 bebnmne ape S1V;41 — IV, 3 muschaticun, 32 mi. 3 Pine S 30 litigiosum, IV, 28 margaricoc EV;36;.3 matthewsii, cum, mexicanum asperu- lum, IV, 25 Pes ahi EV, 35 mollugo, IV, 26, 32 multiflorum, IV, 40, 41, 42 hirsuta IV, 40 angen IV, 42 muricatum, IV, 36 nuttallii, 1V, 35, 36 ma reer IV, 25 pataerrars pt TV, 25 sparsifloru 3 spurium echinosper- mum, 1V, 28 chet dcae er, , dete um, 1V, 37 su pore Ne 36 m, IV, 32 tricornutum, IV, 30 trifidum columbianum, V, 30 cuspidulatun, IV, 36 pacificum, IV, 30, 31 iatena: IV, 30, 31 subbiflorum, IV , 30,3 tinctorium, IV, 30 triflorum, IV, 3 cite IV, 28 V5 2 Raa diffusum, IV, 28 watsonii, IV, 42 wrightii rothrockii, TV,38 Gambelia speciosa, III, 783 Gastridium i I. 156 eo rater III, 306; Gant ‘it, 209, 211 pee heterandra, III, 211 odorata, III, 21] parvillocs: TE 2t} sinuata, III, -. villosa, III Gau gw ieieles III, Gayophytum, III, 206 n, III, 209 hirtellum, III, 209 eorsesornenen III, 208 las m, II I, 208 criospermum III, 209 hoffmannii, III, 209 8 08 oe ee pumilum, III, 209 racemosum, III, caesium oe ramosiss TEL, 2 rile deflexum, III, 208 obtusum 5 va ista ceisdicom Ss 519 a sens a acuta, ITI, 3 affinis VIII, re forwoodit, ne 356 acTils 3 parvidentta, ch, 356 amare Til 3 pepsin III, 359 bisetaea, III, 358 californica — ~ ca alycosa oregana, ITI, 356 orfordit, III, 356 prostrata americana, I, 359 quadrangularis, LEE, 354 sceptrum, III, 356 humilis, III, 358 serrata holopetala, III, 360 oe ifl; 358 sim um, ’ bicknellii longipes, californicum, 1:5 canum, III, SE Aeded. iti.2 longipes, III, 3 cientariwn, 1TE.7 co incisu incisum, III, 4 laxum —, Ill, 4 strigosum, III, 4 syloatien m, III, 1 viscosissimum, IIT, 4 dia fruticosa, TIL 76 ers wae ak ude II, 50 ciliatun, II, 449 phyilum, II, 447 7 wrbanum, II, 447 regonense, II, 447 Gifola germanica, IV, Gigantabies taxifolia, Gilia, It, 396, 456 I, 466 abretan tyre ITI, 463 s, III, 463 heen III, 403 INDEX OF SCIENTIFIC NAMES atractyloides, III, 451 he aurea, III, 419 decora, III, 419 ae. III, 423 bicolor, III, 430 bee art — jones bola ae ur, ‘is, 422 ambigua, 419 brauntonii, i = aie, conges bevitifoliar III, 460 montana, palmifrons, III, 460 Pic terete III, 460 tylophyllum, , 424 davyi, ITI, 467 d demissa, III, 424 densifolia, III, 432 aber i ‘omontana, eal III, 434 mohavensis, vee dei ed sanctora, III, 4 dianthiflora, III reek dianthoides, III, 424 on bie ,4 farinosa, III, 424 dichotoma, III, 422 422 III, 448, 4 volcaniea, TH, 462 elmeri, III, 4 elongata, ie as eremica, ITI, 435 yagert, III, 434 zionis, III, 435 exilis, III, 467 fenslia, ITT, te filicaulis, III, 4 filifolia, III, aoe 437 ifus I, 434 ambigua /Ii, 435 a, III, 437 IE , 426 ax xillavis, Ill, on diffusa, III, 405 rin harknessii, III, 4 heterodoxa, III, 4 a heter ophylia, III, 405 ho mannii, III, 466 hookeri, III, 455 hivwicr dis, III, 405 huege nt Til, 432 7“ Ree IIT, 460, Ill, 471 Ccnretes III, 467 eu-inconspicua ochro- intertexta, AI, 43 inyoe brevasci, ut oe bicolor, III, 472 pve wig III, 472 687 eu- -leptalea, III, 471 meria, III, 471 fili. ipes I, 418 longitu, HHL a crocea, III, 4 eu- -longituba, on 430 ro a, III, Itea, TIL 430, "a eu- ine in 430 longistylus, III, 430 30 tiflora, III, 472 modest, ts 424 hs 467 atgtipe fi 427, 460 688 [Gilia] epnibeimeiaes, Pits ae em 426 pallida, III, 463 pusilla californica, III, 4 pygmaea, III, 417 rattanii, III, 419 eee III, 407 ripleyi, ITI, 462 sherman-hoytiae, II, 434 sinister, III, 472 sinistra, III, 472 ao Sajarae, III, 4 430 ella, TH, 403 418, 430 7 eu-icnuiflora III, 466 INDEX OF SCIENTIFIC NAMES genuina, III, 4 altissima, III, oo triceps, III, 467 acs a III, 467 evilis , III, 467 Z : 6 sa, II, 465 ambigua, III, 435 dasyantha, III, 435 filifolia, III, 437 flor ibu; nda, III, 434 heterodoxa, UL, 451 — IIT, 4 Gilmania, IT, ys Gilaasiaby Tateola, II, 2 iloba, I, 50 latifolius, IV, dl ere IV,8 culatio; des, we 80 peer IV, 80 labra, IV, 80 hirsuta, IV, 80 Glaucium, II, 228 fl II, 228 ’ ei ocarpa, I aridum ; stipulifera, III, 55 Glyceria, I, 211 bore alis, 1, 24 pauciflora, I, 212 (lurnema ninhe * III, 226 a occidentalis, III, 226 3 V, 470, 472 chilense, IV, 469, 472 confertifolinm, sien IV, ae decurrens, IV,4 alifornicum, v, :" dimorphum V,4 dio IV,4 Ng can IV, 489 SOE IV, 469 ies iT 472 IV,4 johnston, bid 469 leucocepha IV "470 tutes ee IV, 468 4 margaritaceum, microcephalum, IV, 469 palustre, IV, He V,4 nanum, IV, Spas IV. 469 ure V, 474 circles ha cee IV, 472 shel huletrs: wr - sprengelti, 1V, 4 eulphurescens, Wy, 469 therm e, IV, 469, 470 Gnaphalodes no es Si aia IV, 494 Gnetales, I, 50 Godetia, III, 184 deflexa micropetala, ce 187 pacifica parvo, ur ae lacunarum, III, 187 pa hain ; TF = ngs pro eAKisa uate alenes ne 185 davyi, III, 18 a eee Ou" sain: 17185 sehen IIE, 188 —_— menziesii, ain ania, II, 342 enieinias II, 346 burnhamit, II, 343 debilis, II, 3 Se thy we 343 glandulifera, II, 343 oa Gossypium barbade onan, HT, 112 — festu- m, I, Graticla: III, 731 anagallidea, III, 688 Grayia, polygaloides, II, 89 spinosa Gree eeuuie. 111,557 circumscissa, III, 557 hispida, III, 557 Gregoria laevigata, nivalis, III, 336 Greniera tenella II, 147 arenicola, pachyphylta, "Ye 256 blakei, I basics, ss 260 “Ty en "258 australis, IV, 259 davyi, IV, 258 intertoris, lV, 258 parvifiora, ~s 257 collina, IV,2 col jan a discoidea, I integrifolia, IV, 258 turbinella, lV, 2 oregana, IV, 256 INDEX OF SCIENTIFIC NAMES wilkesiana, es, 256 256 Patyphl, IV, 258 258 ribrcanis "IV, 256 procumbe ens, I A virgata, 1V, 256 Grossularia, “ 395 a, Ti, 4 neues ata, ‘a 399 californica, II, 402 t 8 hystrix, II, tosma, II, 40 , II, 401 marshall, Lee cond i, II, 4 senilis, IT, 401 sericea, II, 399 osa, I, 403 echaina, Il, 399 II, 388, ruvelia reel tit 3 “aiuie im, boede iene II, 245 rostrata, IT, 253 pers abyssinica, Gutierreria, tv, on bracteata, IV, 2 Galltatnica: TV; = bracteata, IV, 260 ivergens, IV, 26 60 euthamiae, IV, 260 microcephala, IV, 260 glomerella, IV, 261 ionensis, Jee a0 laricina, IV, 2 pore ie! x 260 fe) preedives hala, IV, 2 V, Goemtean longis- pica, I, 47 Cornea rubra, III, 800 Gymnogonium spines- cens, II, 1 Gy mnogramma triangu- laris, viscosa, I, 21 G ymnolente enceli- Gymn lerigt es GYMNO M 50 ymnosteris, III, 407 leibergii, III, 40 minuscula, III, 40 nudicaule parvula, III, 4 nudicaistis; III, 407 Gymnostichum, cali- _fornic wm,t, 255 IV, 4 peers IV,4 Aaa i. I, 47 romanzofianum, I, 479 ees: I, 479 Habenaria borealis, viridiflora, I, 472 operi, 1,4 cuntekds, Ill, 551 deflexa, III, 550 americana, “i 550 IT, 554 Halim nium aldersonit, liga scoparium, III, 122 Ha sri Rae lg occiden- talis, Halioln, os 267 diffus lasio lob ny Halliopkstum ha, Hi pee SS III, 43, 212 Halostachys occiden- talis, II, 91 amosa acutirostris, 0 I, dre (ee sselehs. II, ae em a, Il, 606 rh II, 6 606 kelloggiana, Ul, 603 layneae, II, 600 brachylepis, igh 288 brickel V; se canus, IV, 284, 285 pegs oides. 1V,2 case, Ve? eryth reece IV, : maximus, TV, 272 690 (H. carthamoides] cooperi, IV, 2 cuneatus, IV, spathulatus, 1V, 290 poset IV, 286 — int tenuicaulis, ‘Iv, 275 lan oe IV, 278 lari us, is IV, 288 linearifolivs, IV, 283 nevaden nsis, IV ophit tidis v, 281 stenocephalu zV; virgatus, IV, 276 parishii, IV, 290 racemosu halophiloides, IV, 276 hilus, 1V, 276 liatriformis, IV vada longifolius, 1V, 276 lucida, 1V Hs pinetorum, IV, 276 practicola, 1V, 276 prionophyllus, INDEX OF SCIENTIFIC NAMES IV, 276 sessiliflorus, IV, 276 stenocephalus IV, 276 virgatus, lV, 276 radiatus, IV, 272 resinosus, IV, 282 spinulosus gooddingit, IV; 271 squarrosus, IV, 2 ib cae ie WV, me obtusus, IV, 2 steno clap ive stenophyllus, IV, a poem IV, 280 re s, 1V, 280 Pico IV, 275 tra iskae , 1V, 285 us, Vee H. arpaecarpus califor- nicus, 1V,171 exiguus, IV, 1 ma Pipes. Harpagonella, in, 609 palmeri, IIT Harrimanella, OE: 305 paynbided: III, 645 serphylloides, III, 647 ziztphoroides, III, 648 He i 7 annua, IV, 587 cretica, IV, 588 Hedys. arum, sg 614 nia decurrens era | Heleniastrum bigelovii, IV, 193, 456 Helenium, IV, 208 amarum, IV, ve autumndle,¥ grandi soci og 210 , 210 echhird. TV, 2u1, 212 rosilla, Iv, 2 tenuifolinm, Vv; 210 Heleochloa alopecur- oides, I, 142 ariz: onica Ai “4 ta; 1V “douglas, Ass on ianthemum, lik: 3122 greenei aegeri, IV, 113 lenticularis, IV, 113 macrocarpus, IV, 113 aridus, IV, 113, 114 cusick us, 1V, giganteus insulus, I uiahensis: IV, 116 Be get aasten IV, 114 hoo IV, 104 satearsfoNed IV, 114 niveus, 1V, 114 nuttallii, IV, 116 arishii petiolaris, TV, 113, 114 s, IV, 1 canus Ww. 114 utahensis, TV, 1 Heliamphora, II, 329 Heliopsis balsamorhiza, - — IV, 106 H californicum, m1, 534 Helogyne fasciculata, IV, 498 Helonias paniculatus, I, 376 tenax, I, 373 Helosciadiu m a a nicum, III, 238 leptophyllum lati- folinm, III, 222 Hemiev I, 350 Fanuc olia, II, 350 vio — a, II, 351 misphaeroca rya II, 598 Hemizonella durandit, FV, 172 minima, IV, 172 Hem a, IV, 172 angus, IV, 174 ephala, IV, 175 somalia WV. 1 arida, IV, 1 australis, I balsamifera, IV, 175 calyculata, IV, con se V, 180 merce IV, 182 conju a HY. 178 mbosa, IV, 1 “angustifolia, iv. as 7 decumbens, TV, ai douglasii, 1V, 188 durandi IV, 172 fragarioides, IV, 182 INDEX OF SCIENTIFIC NAMES lutescens, IV, 182 is = a1, 1 ma pstmt iv is vernalis, IV, 18 multiglandulosa, oor nos 189 we = 85 pons! ov 190 pallida, IV 174 pungens, IV, 1 180 % & 80 septentrionalis, ramosissima, rudi. s, IV sca abrella, iv. 188 sericea, IV, 182 ci truncata, IV, 187 vernalis, % se ed TV,A srmannit, Tv, 186 Heracleum, ITI, douglasii, III, 268 ana oH pert IIT, 106 Hesperastragalus com- pact H. esperevax aculis, IV, 495 brevifolia, IV, oe lescens, IV, 49 helunslis. IV, 495 sparsiflora, at 494 I pastas ~ 290, 319 Hes s, I, 411 ee ae we “41 esperochiron, III, 527 californicus, III, 529 benthamianus, III, 529 ciliatus, III, 529 latifolius, Ill, aoe pumilus, III, e estitus, I . io Hesperochloa kingii, I, 221 Hesperocnide, I, 526 tenella, I, 526 esperodoria hallit, IV, 282 Hesperogenia, II1, 242 s tricklandii, III, 242 micranthum, a gulinum, Ill, 14 taser omecon lineare, » 225 Hesperonia bigelovii, 3, californica micro- la, II, 107 cedrosensis, II, 107 glutinosa gracilis, Il. 107 tenuiloba, II, 107 Hesperonix accidens, bolanderi, II, 606 es II, 607 Hespe te fog merten- siana, I, 63 H csperoacordon ewisti, I, 400 Hesperoscordum, I, 400 hyacinthinum, I, 400 lacteum, I, , 298 1,211 Ill, 211 1,211 74 > e - g 8 3 F} . — o g + ° ¢ o he lia, II, 474 fremontiana, II, 474 salicifolia, 11, 474 691 Heterostylus grami- neus, 1,95 gee IV, 265 breweri, IV, 265 echieidal, IV, 264 floribunda, IV, 265 Z va, IV, 2 lamarckti, 1V, 265, 266 oregona, lV, 264 scabra, IV, 266 sessilifiora, 1V, 264 subaxillaris, oregonensis, II, 382 suksdorfi, 11, 383 tenuifolia, I], 383 Heyderia decurrens, L 71 Hibiscus, IIT, 1 californicus, ut, _ 2 denudatus, III, 1 pares tame lasiocarpus califor- albertinum, IV, 595, 596 692 [ Hieraci albifl pris had ne IV, 5 empl, a — m, I 97 hooker 9 leptopodanthu m, TV, 5 longi a. IV, 596 macranthum, IV. 600 pilosella, IV, 00, 602 piper a le ha TVs seabriuseulum, TV. 600 ouler: iV. 305, 5 596, 0 triste detonsum, IV, 599 umbellatum, IV, 597, 600 scabriusculum, TV, 600 vancouverianum, V, 598 washingtonense, IV, 596 Hierochloe, I, 124 INDEX OF SCIENTIFIC NAMES borealis, I, 124 macrophylla, . 124 odor gid a te Idia adpressa, EE Hobe caus II, 480 densiflora, II, 4 480 III, 403 squarrosa, III, 452 Hehe IT) ,.20 ryi, III, 20 Holes of 163 halepensis, . Fea ee Holochloa cylindrica, II, 382 clata, II, 382 glabella, IT, 383 Holodiscus, II, 414; discolor, e 414 fra us, II, 414 ir rp II, 414 dumosus saxicola, II, 4 glabr rescens, HE yo saxicola, II, 4 Holosteum, TH. ui bellatum, II, ve Renn tean fale 156 Rosie Wy ine os FEEHSTS, II, 574 entiieliit, Is bicristatus, - ome oe IT,.573 collin Ta 571 omopappus argutus, IV, 276 glomeratus, IV, 276 paniculatus, IV, 276 276 bicolor, I, 402 californica, I, 406 congesta, I, 408 coronaria, I, 405 douglasii, 1, 401 filifolia, 1, 407 printers I, 402 des Ingens, I, 399 leptanara, 1 go mi us at tH jae a es Pests P maritima pussone: anum, I, 246 muri num, I, 245 pusillum, I, 245 vulgare, I, 245 Horkelia, II, 416 Se ct as 1, 423 os uley 4; [fs aa ee _ 419 brownii, II, 418 ruleomontana, 8 OL californica, ye: 417 ee II, 417 fsa II, 417 campestris, II, 423 a enuduanite II, 418 tenella, II, 418 tenuiloba, TI, 419 glandulosa, IL, 417 gordonii, II, 427 alpicola, II, 427 grandis, II, 417 he nderaondk II, 419 ka ellog git, Il, 417 caer es ae I, 418 purpurascens, II, 421 congdonis, II, 421 tilingii, II, 421 tweedyt, Tl, "42 6 se arin II, 424 — aes oe Horkelila prpuras: I,4 Novia hl ae III, 643 Horsfordia, III, 83 alata, Hi, 83 brachycarpa, IT, 54 crassifolia, II, 541, 552 r gracilis, grandiflora pear eer = we ee JH, 552 lanthanides ee 542 ee — ies vs oe maritima, IT, 545 angus stifolia II, pies ochroleuca, 2 ornithorus,. 50 a iH, 5 sulphurea, II, 547 tomentella, II, 543 INDEX OF SCIENTIFIC NAMES tomentosa, IT, 5 glabriuscula, 49 ee torreyi, II, 54 nevadensis, II, 542 venusta, II, 550 raat II, 545 wrighti i multicaulis, IL 543 Houttuynia californica, I, 4 Howellia, IV, 88 aquatilis, 1V, 88 ihe: IV, 88 gelia abramsit, brauntonii, III, 434 densifotia, ITI, 432 austromontana, III, 434 mohavenstis, III, 434 conga III, 434 ere pars sitlore, U1, 437 hooveri, III, 4 lutea, III, be Sresanae “0 434 437 ppb: : i 435 dasyantha, III, 435 sapphirina, III, 435 Hulsea, IV, 202 algida, IV, 206 brevifolia, IV, 203 nyo ensis eni, 1[V, 204 a = — Ss es BS a $ _ & 04 eect Fusiasi- a IL 2 B ghee ic 290 calycin na, a, I, 302 II, 290 procumbens, II, 290 DRANGEACEAE, pares onic! , 384 Hydrastis carolinensis, I, Hydastylus, I, 468 borealis, I, 469 Hydroc otyle, III, 217 chinensis, III, 218 cuneata, III, 218 prolifera, III trira vulgaris, ydropeltis purhuree, II, 172 pig HYLLACEAE, III, 4 Hyd iat III, 478 albifrons, III, 478 5 macrophyllum occi- dentale, III, 478 binbesrinar prs) III, 487 oceiduniiia ITI, 478 e 1 478 8 Hymenantherum tenui- folium, IV, ger sis IV;2 ee ive as fsccnlat ~*~ 145 a, 1V,1 pentalep ix ry 145 salsola, V1 " veatetaade + 145 LT LAh. e PY hs tus, T, 290 pie Pas IV, 556 —— IV, 556 Hymenopappus s, IV, 392 eas halus, , 392 anus, IV, 3: 1V.2 07 biennis, IV, 208 californica, IV, 218, 219 calva, IV, 219 —— orata, IV, 208 Hyoscyanrus, ITT, 668 niger, ITT, 668 I Hyoseris cretica, V, 588 hedypnots, 1V, 587 taraxacoides, IV, 588 canes major, ve 117 ncinnum, IIT, 116 majus 17 mutilum, III, 117 rforatum, III, 116 116 tapetoides, III, 116 Hy oeris, 1V, 586 glabra, IV, 586 _radicata, IV, 586 1,25 Hypopitys, III, 295 fbr III, 295 latisquame, 9 295 Hyptis, IIT emoryi, te ree verticillata, III, 662 Hyssopus Reread tie sty III, x, i152 sts tee 15255 grits Ag 2 Bis oc jae he i, romansofianum, columnaris, III, 474 Ilex myrsinites, III, 54 Iliamna, III, 84 ae I, 84 a ges tea da ak 86 longis eae III, 86 remo I, 84 rivalaris, LECEBRACEAE, II, 111 cates pckiiiie. es 113 Tl Ill , pesca III, 688 694 a sora es] Infantea chilensis, IV, Inula, IV, 4 cricides 332 helenium, WY, Pe 466 scat IV;:2 subaxillaris, aE 266 INULEAE, IV, 465 Inyonia dysodioides, I Ionactis alpina, IV, 328 stenomeres, IV, 327 Ipomeria ageregata, III, 460 caurina, douglasiana, x 464 dia palustris americana, III, 168 ocoma acraden 84 leucanthemifolia, v. 28 microdonta, IV, 284 INDEX OF SCIENTIFIC NAMES Ser goa £35 oet 35 fees J 35 aupercula, I, 36 cutti, I, 38 occidentalis, ms 36 orcuttii, I, 3 steelers “ 36 underwoodit, I, 37 Isolepis carinata, 1, 270 koilepis, I, 2 Aapars 272 p can asi . i pti Lie} stipitatum, II, 176 , II, 426 gordonii, TI, 422, 427 lyc As odioides puenace, IT, 426 Pg ireapadicides IT, 427 427 egalopetala, II, scandularis, II, 427 megalopetala, II, 427 pea Poe ii, rd 424 purpurascens, II, 421 A=] i scandularis, II, 427 sericoleuca, II, 425 shoc a webberi, II, 426 Jac ager trachy- sperma, II, 327 Jac obtnia californica, IV, Jamesia, II, 387 americana californica, II, 387 eonuttalliana, IT, 354 ube i, II, 354 Selon. II, 354 te ag inaequata, ITT, 581 racemosa, III, 580 Juncoides, 1, 369 campestre, I, 371 divaricatum, I, 370 glab m, I, 369 Aidt I, 369 pipes pte cet! tgs a eubcongestom, T5370 Juncus, I, 350 scuminats I, 366 acutus, I, 350, 355 sphaerocarpus, T7355 alpinus rach ystyl us I 361 dudleyi, I, 357 dubius, I, 365 latifolius, I, 358 exicanus, I, 354 nevadensis, I, 364 eee 8,4; phae ialomeratus, on 367 gracilis pan aoe = 367 9 ‘a ncialis, I, xiphioides, I, 368 tus, I, 368 ibi E97 nana, occidentalis, I, 76 scopulorum, I, 76 sabina nie 75 sibirica, Like utahensis, I, 75 californica, III, 168 peploides, III, 168 Ka ee ITE-137 crophylia, ITI, 302 afta, III, 302 crophylla, III, 302 occidentalis, _ “sing polifolia, oc naan vr 302 Kalmiopsis, III, 301 achiana, III, 30 ruber, , Kentrophyllum baeti- cum, IV, lanatum baeticum, IV, 5i2 Kentrophyta impensus, montana, II, 568 KorneeLintacear: TTT, 55 Koeleria, I, 169 maritima, IT, 3 Kopsiopsis ic. ne onhle IV, 10 Krameria, II, 627 I, 627 pareja glandulosa, imparata, II, 628 KRAMERIA ee II, 627 Kruhse Sk yale I, 458 INDEX OF SCIENTIFIC NAMES tilingiana, I, 458 Krynitzkia Gfinis, III, ond ambigu i a es barbigera, III, 585 barbigera inaps, III, 585 chorisiana, III, 563 circumscissa, III, 557 operi, II ns, III, 583 dichotom Fe | 02 elomerate virginen- ois, 600 a, III, 580 micrantha, III, 578 micromeres, III, 583 sta micro chys, III, 595 mohaven ITI, 593 mollis, III, 559 yr obs abortiva, III, 5 mariewat, ae te oxy tahen “watsonii, 1H, 593 a $59 searlsiac, II, 559 mlienia cooleyae, II, 211 hystricula, II, 211 Kunzia glandulosa, II tridentata, II, 451 Kymapleura hetero- Dphylia, IV, 566 ental: 2 IV, 498 Labiatae, III, 614 Lachnostoma hastula- 5373 pulchella, IV, 594 racemosa, IV, 593 saligna, IV, 592 spica ata, Lamiaceee ye) bape mium, Il am amplexiauie TL — sine tum, ITI, 6 “TH, “628, "630 bot a 402 Langloisia a, III, 4 campyloclados, I, 453 Laothoe, I, 413 pe he I, 414 divaricata, I, 413 po intricata, megalocephala, IV, 200 villo shee sulias, VV. 508 minor, IV, 508 La , III, 548 I america a, III, 550 1,553 cusickii, III, 551 bella, III, 553 californica, III, 554 mundula, I re 555 ee deflexa americana, 695 III, 550 diffusa, III, 553 hispide, Ill, 554 texana columbiana, III, 549 velitina; Tit? 4 dolar: Hiss Lap a ensis, I, 60 laricina, I, 60 larix, = 6 wideateih ‘ght tinosa, III, 16 Lastarriaea, 1E3 3 graminifolius, II, 622 696 [Lathyrus] mas II, as jepsonii, II, 6 jacuaeeaa: Tt: oe — 11,622 lan 4, 215622 pay sraber) cr rigidus, IT, chaffneri, II, 626 plendens, II, 627 strictus, 11, 626, 627 sulphureus, II, 6 argillaceus, II, 623 svilvest II, torreyi, IT, tencllus, IT, het , EE clifornicus, II, 623 II, 6 e 11222 losemite ceuniiel I debilis serrata, 1V,97 Lau coy ilicifolia, 11.4 bau II, 467 fremontii, IV, 159 INDEX OF SCIENTIFIC NAMES emo a; IV, steadaieia. IV, 162 heterotricha, IV, 164 hispida, TV, 162 utea, IV rian IV, 162 162 — Iv, 162 160 fecovanga, IV, 159 munzii, IV, 1 nutans, 1V, 166 niculata, IV, 163 pentachaeta, IV, 162 albida, IV, 162 hansen septentrionalis, IV, 160 Lebetina cooperi, porophylloides, IV, 251 Leersia oryzoides, I, 120 Ledum ees ae eat, 298 columbianum, III, 299 seers: vias 298 = ost aati IIT, 298 palustre, III, te — enere, IV, 8 lim a, IV, 88 Le OK usta a biflora, IV, 78 V.78 Tl, 474 N Sicis ingles, TIT, 2350 Leighia lanceolata, IV, Leiostemon thurberi, III, 750 peng oy congdoni, IV, 1 Lemmonia, III, 522 ealitopaies, TIE S22 LENTIBULARIACEAE, IV, 10 Leonotis leonurus, Tif, Lesbkes triphylla, Ledtitedon: IV, 588 autumnalis, IV, 589 boreale, IV, 556 hirtum, I hivwuturs, IV, 566 hispidus aves 588 lacvigatus IV, 611 leysseri, IV, a 589 lyr ae IV, 61 nudicaulis, EM; mae ceratophorus, IV, 611 , 588 Lepargyraca argentea, 64 enna III, 164 iv peat hinia, TI, 643 Pe i jonesit, 259 a etalu win, IT, 261 bipinnatifidam; LE; 262 campestre, ee ss nes L264 sires Il, 261 es a Rt didymum, IT, 263 draba, II, 258, 259 flavum, II, 25 fremon idahoense, II, 261 intermedium, II, 261 jaredii, II, 259 jonesii, IT, 259 , 262 ect U1, , 258 me oe menziesii, os oi 262 montanum, II, 259 canescens, II, 259 j. FE, 259 repens, ruderale lasiocarpum, texanum, II, 261 monoides, III, 769 ae poco peed madi- oides, 1V,1 Lepigonum gracile, macrothecun m, IT, 156 Lepta us gramineus, Leptarrhena, 1B 355 I asea vespertina, 361 Lepraxi mensziesit, TE 374 Leptiton canadense, 382 Bebidas hloa, I, 180 uninervia, I, 181 Leptorchis ‘loe selit, Lepiedaétploni IIT, 454 5 _ _ aos 2 mn on on ft unr Pelican! III, 431 hallii, 455 peter = 455 pee ri, III, 455 aegeri, ‘IIT, 4 Haein: Ill, 4 pulchriflorum, ITI, 45 shastense, IIT, 455 pungen re hrifors, III, 455 pence TIs455 squarrosum, III, 455 subflavidum, III, 455 455 Leptoplectron, III, 784 Leptoseris sonchoides, IV, 578 Leptosiphon acicularis, LE, ee i 429 s ms osyne bigelovii, 132 valfomelea:} IV, 134 hamiltonit, IV, 132 pts AV, 13 ewberryi, IV, 2 iillmane: IV, 134 aenia anomala, II, 25 radshawii, Lit, 252 a, III, 267 denticulata, ie 257 leibergi oe min aTIL, 2 m ultifida, I sal) e ardina, II, 287 sone: 286, 287 peirsont ramulosissima, 377 TV.3 eee IN; nf) ighreeet lemm nii, IV, a7, 378 son cocoa The a leptoclada, IV, 380 arachn a IV, 380 halote euc 380 microcePhale, IV, 398 tenuis, 1V, 380 Pies IV, 378 micradenia, IV, 378, 379 INDEX OF SCIENTIFIC NAMES arachnoidea, IV, 380 glabrata, Se 379 nana, | eases, ae 378 lada, icr a, IV, 378 m erocephola, IV, 378 tenuis, I ce. VV. 377 subs tag ata, 1V, 378 tenuis, IV, 376 jared, IV, 376 mentosa, IV, 377 virgata, IV, 378 um cha- ‘macmelin, il 399 leu 1V, “a oe IV, 394 celene, IV, 332 ar pine IV, 332 ericoides, IV, 332 ste co ocraspedum albo- ntanum 1 Le ucOSseris Ladleacmibin: , 580 Rls. IV, 580 2 onuifolia, 1V, 580 Leucosyris carnosa, Vv davisiae, Ill, er Lewisia, gf 131 alba, curantice, " a ber 4 elle ae - poi br achycar rpa, IT, 135 sierrae, IT, 134 triphylla, II, 131 tweedyi, IT _ w w cu E71 onta) I, 70 Ligusticum, III, 2 ehtas aka: i. A abisileletie helleri, 2 apiifolium, III, 235 minor, III, 237 cacruleoMontanium; a 23 anbyi, III cUusi ee III, oo filici tenifolium, Ill; 235 grayi, III, 2 bor ns ‘ae ata occidentalis, TIL va LILIACEA I, 379 Liliorhica lanceolata, I,4 ium I, 416 bakeri, I, 420 bloomerianvan I, 420 atum, I, 420 boa 1, ree chee um, 1,4 ns, 1,417 washingtonianuit, I, 417 purpureum, I, 417 Limnalsine diffusa, I, 124 LIMNANTHACEAE, IE Limnanthes, III, 46 alba, III, 49 pia ae III, 49 baker Piatra ONG III, 49 697 ee be - ta, sulphurea, III, 46 versicolor, III, 47 parishii ah 47 Limi ores cola, IE f29 ee Fe . ed difolia, perfoliata, U 127 Jmmorchis, C es Limonium, ITI, 3 aiitonahied "TE, 3 commune neitfotuica III, 344 mexicanum, III, 344 mex m, III, 344 tica, III, 732 Linanthastrum nut- tallii, II flo bund, _ 431 413 rae ‘III, III, gu dacislopksim, Pe m, IIT, 424 d canores III, 426 698 [Linanthus] dianthiflorus, III, 424 farinosa, pe 42 4 ait septentrionalis, III, jonesii, III, 422 pharnaceoides, , 418 vall icola, III, 418 43 ori nudicaul, I, 407 oblanceolatus, III, 427 orcuttit , 426 luteolus, III, 430 Ill, 430 peirsoni, itt, 423 * naceoides, ‘M8. Be Ill pusillu pach shear ertsonii, 427 bE See ae 786 INDEX OF SCIENTIFIC NAMES longiflora, IV, 48 longifiora, IV, 48 : et Ee: IV, 302 rip llr IV, 288 4, 1V, 300 , 330 Ceilaeain: ot 303 depressa, 1V,3 howardit sense IV, 301 pari g = — 299 par adenophyltum, L312 angustifolium, III, 11 puberulum, ee puberulum, III, sperg be a III, 14 usi tatissimum Tl, 11 8 a, £ canescens, 613 rior, oe 613 lanceolate, it, - reptans, III, 6 wr ightii, Ill, dey Liquivitia lepidota, II, 613 Lisianthus exaltatus, i sty a caurina, I, 48' convallarioides, 1, 480 rdata, 1, 480 co nephroph _— I, 480 eri, II, 369 ig, Ss campanulat: a, > a oo a II, 370 mbalaria, Il, 370 laciniata, II, 369 parvi iflora, II, 368, 370 rupicola, TL, 368 seabrella, II, 369 nella TL 368, 369, 370 i he! Lithospermum, tL 546 arvense, III, 548 californieum, hhee 548 circumscissu , 548 eolatum, III, 548 rinctorium: on 574 a, IV,95 cardinalis graminea, IV, 96 multifiora, IV, 96 pseudosplendens, oo IV, er dortmanna, IV, 9 door. cael, Rag 97 falcata, 1V,9 graninea IV, 96 7 IV, 96 nea, 1V, kalmii, IV, 96 strictifora, 1V, 96 pu V,9 renathl lia, III, 454 pee aga a bg i Bai 3S re a, IV, 490 seth oe r ‘asd neers I, 236, 237 cristatum, I, 237 italicu 1, 237 multiforme 1-237, rictum, I, 23 arvense Lomaria spicant, 12 Lomatium, IIT, 246 alatum purpureum, III, 25 ambiguum, III, 253 angustatum, III, 261 avum, III, 261 :; caliters nicum, III, 267 psi Ii, 252 caruiielbat, 257 erythropodu: TIL, 257 solanense, III, 257 congdonii, 2 cous, III, 25 cusickii, III, 266 cuspidatum, Il, um decorum, ET medium, III, 258 I, 263 gelm anni ii, III, 261 humile, III, 257 id nse, III, 267 insulare, III, 249 jonesii, I l semivittatum, III, 260 marginatum, III, 255 rtindalei, III, 264 dicktindins. II, "253 nelsonianum, III, 264 nevadense, III, 257 parishii, III, 258 vevisdeicatall: I nudicaule; TIT, 268 pa ts m, III, 250 rollins sii, III, sg salmonifioru TH, °363 semisepultum, III, 258 serpentinum, ae 263 simplex, ITI, 2 aca iea- III, 266 sonnei, III, 2 suksdorfii, bo 267 tra 125 triternatum, ITI, 266 m, III, 266 pete III, 266 “TIT. 266. tuberosum, III, 263 utriculatum, III, 255 themifolt Tiie2s glabrum, III, rrupta subspicata, IV, 54 INDEX OF SCIENTIFIC NAMES Ehren IV, 52 e ae IV,4 camuni s, I Laphonths cusichii; a dea III, 626 urtcifolins, III, 624 Lophochlaena califor- THACEAE, I, 528 Lotodes californicum, II, castoreu FE, elipticum latifolium, IT, 554 macrostach yum, orbiculare, II, 554 nevadensis, II, 54 nuttallianus, II, 550 oeomes is > 542 pie river IT, 545 Loxanthes fasciculatus, wigia, 168 wires rade mr, 168 apetala, III, 1 ans Sek eg IIT, 168 palustris americana, IIT, 168 if nar recto! IV, — pipert, 458 rpentina, IV, 457 stricta, IV, 458 albo: albus, II, 483 699 aliceae, IT, 630 alcis-montis, IT, 630 alilatissimes, Il, 502 amphibius, IT, 505 i, 11, 51 sicki ; lobbii, 11, 495 torreyi, II, hoensis, II, 496 arvensi- -plasketti, II, 630 austromontanus, IT, 507 barbatus, II, 514 benthamii, borealis, Il, 1 brachypodus, 11, 496 b eei, IT, 630 bitoni, bru cu ro amblyophyllus, it. carnosulus, II, 494 caudatus, II, 497 rubeicaulis, II, 497 700 Bes ones oo al submanens, II, 630 Pir sition ag I, 497 Rete dona IT, 630 christinae, Tl, 630 i us 94 aiupee et ES 505. flexu Pe ormosus, II, bridgesii, IT, 514 ler , 11, 51 Ss. fragrans, II, 63 glareosus, Il, 487 INDEX OF SCIENTIFIC NAMES gormanii, TEAS LL II, 498, 505 amblyophyllus, II, 498 horizontalis, II, 486 johan s-howellii, 1; 631 jucundus, ay “hee ns 1, i; febabicna II, 631 ps eus, II, 487 hendersonianus, II, mollis, TI, 498 rmanensis, II, 498 II, 509 littoralis, II, 509 lobbii, I1, 495 longifolius, “1 = longipes, II eideericl microphyllus, II, 495 pachylo “ ee IT, 494 trifidus, I1, 495 microcarpus II, 487 dai 487 Orin. II, 487 indie. soba; II, 631 minimus, IT, 515 Be onthe’ It, ots pe eee: II, 631 moll m ollisifotius, II, 507 , II, 494 apricus, II, 494 1,502 ihe Tr, 511 ein II, 631 a I,.631 piperitus sparaipile- Pea II, 495 grandifolius, II, 515 pallidipes, II, 5 pondii, 491 intermontanus, II, 490 pocbaags% II, 503 631 , 506 gle datretaaess. II, priate II, 506 mpsonianus, A wallowensis, II, 631 sis, II, 510 subalpinus, ubhirsutus, 91 sublanatus, IT, 631 riceus, II, 518 transmontanus, II, 487 ucculentus, IT, 490 succ suksdorfii, II, 506 sulphureus, II, 498 Fe naa II, 6 Pct II, 515 ber w ayethit, yakimens is, II, 631 po eRe II, 518 Luzul glabrata, I, 369 lutea, 1, 369 andersonii ITI, 667 rticola, III, 667 gracilipes, III, 666 Aare . Mestre III, 667 has. aa ae ae mum, ITI, 666 patineri, FEL. oc parishii, ITI, 66 pringlei, III, pe richit, III, 666 INDEX OF SCIENTIFIC NAMES shockleyi, III, 667 pei Me 466 rreyi, III, 667 “fljorme, III, 667 osum, Ill, 666 slated. III, 667 Lycopersicon esculen- salica sanfordii, III, 167 tribracteatum, ITI, 167 Machaeranthera, attenuata, IV, 334, 335 s, IV, 334, 335 Mac rocarphus achilleae- folia, Os WV3 Macronema naades. IV, 300 discoideum, IV, 280 molle, TV, 280 pulvisculiferum, IV, ~ scopari TY. 2 suffraticosum, iV. ‘300 alpolee IV, 281 Marnapodtuin lacinia- tum, II, 244 orhynchus angus- == 8 Be | wn sep ide IV, 562 harfordii, I hetero i ee iv, 566 humilis, lessingt, We gor orsum, IV, 565 froximoides, IV, 564 PL ePHIL III. 732 . adaria corymbosa, fiber SN hispida, racemosa, IV, 168 M ht ate angusti- folia, IV, 1 carnosa, Vv G 701 douglasti, IV, 162 e s, IV, 160 heterotricha, IV, 164 hicracioides, nA 162 hirsuta, IV, 1 Madia, I ¥.4 densifolia. ag 168 hispida, IV, We vernalis, IV,1 whe re IV, vr exigua, macrocephala TV 17 filipes, Vv 171 macrocephala, IV, 171 cease ag 170 pilosa, IV, hallii, IV, 166 hispida, 1V subspicata IV. ia la, IV, 168 154 Madronella A le III, 657 candicans, III, 657 diss gers IV, 170 douglas oy Ie Ff lance Slat amplectens, 2 aquifolium, IT, ar 218 dictyota, II, 218 702 [Mahonia] fascicularis, II, 217 fremontii, II, Maianthemum, I, 454 bifolium dilatatum, I, 455 convallaria, I, 454 dilatatum, I, 455 Mataceag, II, 4 M. alacolepis coulteri, V, 576 Malacomeris incanus, IV, 579 Malacothamnus arcua- he - insularis squalida, IV, 576 obtusa, IV, 578 arviflora, 1V, 578 platyphyli, IV, 584 runcinata, IV, 576 meee “TW. 580 tg INDEX OF SCIENTIFIC NAMES tenuis, IV, 496 Malus, IT, 470 diorre ote, Be 470 coccine fsciulaa, aie 2 hederacea, III, mabachtoides. in "ie ro otundifolia, IL, 109 stellat sylv ve ris, See 109 i fragrans, ITI, 95 fremontii, Ill, 95 ce rcophorum, IIT, 95 um, ITI, 95 Te, III, grossu lariaefolia, jo hallii, IIT, 91 ri, III, 95 howellii, III, 95 cordatu 9 III, 94 fasciculata, III, 92 betas ITI, 95 m, III, 90 Mammillaria. ae oe alversonii, III, = as) ™ ~ > 8 ais 8 i) 25 Be mH + —_~* _ ot a i] se ow Tit: 159. i 60 simplex, III, 162 simpsonti, III, 159 Icolanata, III, 159 tetrancistra, III, 160 Marah, IV, 6 67 fabaceus, TV 76271 mis, IV, 70 macrocarpus, ve 70 major, 1V,7 micranthus, TV: 70 major, lV, micranthus, IV, 70 muricatus, IV, 68 vestita, I, 9 iN ea je | Martyn z, IV, 1 M en auernitolis. I altheaefolia, ne 3 mS, BS Ng % = = “ a vulgaris, lV, 388 M aus parvifolia, Ist 3 Matri ‘elath IV, 398 ae hang IV. 4 400 inodora, IV, 399 maritima, IV, 399 matricaricides. obliqua, IV, 39 occidentalis, IV, 400 odorata, IV, 3 parthenium, Iv, 396 tens, 1V, 3 pube estens: Iv, iy segetum, IV svovecieds IV, 399, 400 vulgaris, [V, 396 Matthiola, II, 321 incana, TI 32k ure wa flora, III, 784 pbs) val 784 stricta, Mazus, x bot japonicus, IIT, 688 rugosus, III, 688 Meadia clevelandii, III, 342 nee ee ah 341 5 Meconopsis crassifolia, i; heterophylla, II, 230 Medica iupulina, TI, 520 media Me ees ‘" sis 0 upaniana, II, 520 maculata, II, 520 n i kk eu Meg alodonta beckii, TV (13 hendersonii, IV, 130 Sd Pager lee IV, 130 pe IV, Megarrhiza secs, T¥;71 guadalupensis, 1V, 70 ve ah, 1 muricata, IV, 68 oregana, IV, 68 M egastachya uninervia, Lae Melampodium, IV, 138 americanum, IV, 139 il, IV, 172 ar lineare, III,8 M felandrinm oy Hekei- m, II, 1 Fier ie Pa Mel anobatus Smee nus, II, 457 sabi II, 457 ares m Te MEL NTHACE EAE, I, 371 Melarhisa Soiclaided; TV,.1 Melia sr OES I, 20 u canna is; Ti; MELIACEAE, III, 20 parishii, 1, 193 poaeoides, I, 193 ta 92 Melocactus wirideacent, II, 156 Mengea californica, Menodora, III, 349 helianthemoides, III, 349 glabrata, Ill, 662 pe IIT, 661 INDEX OF SCIENTIFIC NAMES piperita, III, 66 pulegium, III, 6 rotunaisaiin 4 8 spicata, III, 65 rotundifolia, III, 661 gilda Loe eremophila, III, 137 nitens, III, 137 lon globo, Ii], 135 micrantha, III, 140 nultiora, wee 135 S,L11, phila, I an 4a pechihte III, 139 peirsonii, III, 135 pinetorum, III, 139 puberula, 3 12137 MENYANTHACEAE, Si si nth es, IIT, 365 z 303 ferru giabelia III, 301 inter media, I. 172 unispicata, s ae tga artied, Fs jeines texana, bakeri amoena, 546 bella, III, 544 br achycalyz, IIT, 545 ciliata, III, 545 stomatechoides, IIT, 545 cu usickit, III, 546 a, III, 546 Presi III, 546 — iit; 54 ensis, ITI, 546 penile III, 545 pase borealis, ate pe 6 onicnaite:: ar. 546 pulmonarioides, III, 544 oe III, 544 stomatechoides lif, 54 45 subcordata, TIT, 545 seguilaterale, arr 119 chilense, ITI cordifaium, in, 119 tallin .118 pnb rari GC 119 edule, III, nodiflorum, III, 118 Mesorcanth us barbiger, 24 II, 249 fallax, II, 249 vimineus, II, 249 esosphaerum emoryi, Ait, M espilus amelanchicr, I , Metagonia ovata, 27 III, Micrampelis fabaceus agrestis, 1V,71 jranthes acquiden- a, II, 366 aitesii i; Au tip 3 ee howell i; 363 na, ruhdale Hoey Microbahia ot Hef rh a, ITI, 351 “aor IIT, ses gulari _ 351 | M icrolon chus salman- ticus , 544 703 Mi icromeria chamis- procera, leptoscpala, IV, 557 lindleyi, TV, 558, 559, 560 , * aphantocarpha, troximoides, 1V. $53 Microsisymbrium BP sien rr II, 246 III elves mage a, III, 462 elbelle ITI, 413 racilis, III, 413 To nike, ITT, 413 Abin IIT, 413 aski III, 462 Miium lendigerum, Miltitzia. III, 518 glandulifera, III, 519 704 [M. glandulifera] mimetanthe pilosa, III, 706 Mimosa scorpioides, IT; — MIM Minaiees: 7 me arvensis, III, 711 atropurpureus, I aurantiacus, III, 715 Tit; biolettii, III, 699 bolanderi, III, 724 III, 724 brachiatus, TiT,.723 clementinus, III, 711 evelandii, III, 714 clivicola, III, 718 pti re TET 722 e Ss, tH 7 decurtatus, III, 724 is, 21 subulatus, III, 700 INDEX OF SCIENTIFIC NAMES fremontii, III, 722 viscidus, III, 722 geniculatus, IIT, 700 eo 70 4 lyratus, III, 711 np siacter ss te 710 puber cts, in yee niet III, 71 12 hallit alvordensis, FILES? hirsutus, ¥1j, 712 moniliformis, Il, pea sai III, 704 n 69 se ssilifolins, Ill, —- eae ag mh s, 1] "bllolok. i = 1 subuniflorus, ie 730 nasutu s, III, 7 insignis, III, = mnicranthus, III, a procerus, III, 712 erulus, 111,707; 712 pulchellus, ITI, 727 veronicifolius, III, 707 viscidus, III, 722 washingtonensis, III, 70 whipplei, III, 701 whitneyi, ee Nes wolf, IIT, Minuartia So Sa II, 14 douglasii, II, 146 hoz ae jig sect es major, ee pusilla, Tt; Hy II, 47 Mirabilis, 106 bigelov ae ee ae a, aed trifida, Mitelstra, =a 375 II, 375 Miellopsis hookeri, 377 Gece Il; 376 pentandra, II, 376 ifida, 11, 377 Mit tophyllum diversi- folium, IT, ae eu ola, TIT, 8 ateliniant: ee 84 uthiiedds III, 84 Moehringia, IT, 152 pees ra, IT,152 ee 11, 152 MUSCO. 152 ar ea ou 788 brev III, 788 aia. III, 788 aide III, 788 Moluccella laevis, III, 630 Momordica echinata, I Monanthochlog, I, 181 Monardelta Ill, 648 onoides, III, 654 ausieets. IIT; ie breweri, ITI, 6 can ndicans, III, ee ae 48 = a . Loan) = inl Nn cinerea, IIT, 650 crispa, ITI, 655 c oriacea, III, 650 sia: ‘it a forsee pen nite 650 glauca, III, 6 plébos a, II hypoleuca, III, 651 saxicola, IT 1 lanceolata, III, 6 glandulifera, =~ as midveegliahs IIT, 655 sanguinea, III, 655 ledifolia, IIT, 651 le ucocephala, IIT, 657 4 odoratissima, “TIT, 654 glauca, ITI, hig ta, ie. a — IIT, 654 654 pallida 1.6 III, 650 suhaserath, IIT, 650 INDEX OF SCIENTIFIC NAMES z ‘iminea, III, 655 II, Monerma cylindvica, Motiéses:; TE, 291 grandifiora, III, = reticulata, III, 29 uniflora, IIT, 291 reticulata, III, 292 — YLEDONES Pine epis, II, 74 riuttallinne; II, 74 » e245, a4 EV 223 Mono ptilon, IV, 304 heliiforie, 4,305 gn oie IV, 305 mip tg x intricata, “IV,2 megoceala IV, 200 Mon ac 1il, 293. caifornee ae 296 a, III, 293 Paes III, 295 hypopitys, III, 295 californica, III, 296 laithepsdooas IIT 5 flora, III, 293 ROPACEAE IT, 1 gyps soils, il. 127 hallii, 11, 1 heterophylla, ae 129 howellii, IT, 1 7 129 pean se 129 spathulata, IT, exigua, IT, 127 stenophylla, a an sweetseri, IT, waskinete das HH; pikes ne ce oo mum, - line igh howell, “Nb ae Mortonia, IIT, 54 scudectla utahensis, III, 54 pede bai IIT, 54 utahensis, ITI, 54 Mudieas californica, pe rfoliata, II, depauperata, I, 138 filiformis, I, 140 folio sa ambigwa, T, 140 glomerat, A bre vifolt ie oe gracili age gracilina, i 40 leaiiodt “ei 3 sehreberi ‘ id uarr 38 ssleatico gr 140 cant, I, 141 Ss tenuis, uscaria ‘adscendens, IT, 359 penchgisg TI, 359 Mu E, IV, 549 Myaeeee panicilatum, , sativum, II, 291 Mycelis muralis, Myginda myrtifolia, ick a, 111,599 i IIT, 542 705 micrantha, ITI, 543 Ve niceahihle M1, are HI, redowskit, storpioides: III, 541, 542 peiatics , LIT, 542 palustris, 111, 542 pee n IL, 565, 566 sic LLA95 Myriopteris gracilis, 1 2F Myrrhis bolanderi, I claytonit, III, 226 Mysorrhiza californica, cooper cemhie: Iv, re grayana, pinorum, IV, ‘ I ese — iu 594 Nal E, I, 9 marina, I, 9 palsies: a 92 sie Naioc a ptt li-t Nama, 2 S25 ar aretioides: pen 523 biflora spathulatum, 1:52 pwn ti ITI, 525 706 [Nama] hispidum, III, 525 revolutum, III, 525 7 5 sagittata, WV, 456 specio , 455 Na oP ria ip 1,372 californicum p;\373 ssifragum occiden- tale, I, 37 Nassella chilensis, 113 Nasturtium armoracia, II — m, Il, 278 9, ~ 4 zs < mei pone fe bo fee iS On INDEX OF SCIENTIFIC NAMES diversifolia, III, 405 dubia, III, 4 pa AINE A III, 452 erecta, III, 443 eriocephala, III, 446 sail fallax 451 filicaulis, III, 449 filifolia, III, 437 diffusa pr opin qua, III, 443 involucrata, III, 440 lutescens, III, 435 antha, III, 451 matthewsii, III, 453 a, LH, 452 jaredii, III, 448 nigellaeformis, TIT, 445 radians, III, 445 nudicaulis, IIT, ae nuttallit, oc pe 449 pilosifaucis, III, 443 plieantha, ITI, pu ea ee II , 419 a, III, 451 III, 443 seneaiil IH, 452, 453 setiloba, ITI, 446 setosissima, III, 453 lig Bain fii III, 442 3 ~ = a — = te gs ms w wm rac III, 434 gymnocephala, ITI, 435 oligantha, III, 435 5 I purpurea, III, 452 ang cadiforwinuns, no vere capitata, II, 409 — IT, 409 monogyna alternans, II, 410 opulifolia mollis, Nemacaulis, 11.2 denudata, II, 2 foliosa, II, nuttallii, II, 2 Nemacladus, IV, 82, 87 denophorus, IV, 84 capillaris, IV, 87 landuliferus, IV, 86 ramosissimus, montanus,1V,8 pinnatifidus, IV, es rigidus, IV, 86 odiifolia, III, 480 erosa, IIT, 481 evermannii, a 482 exiqua, III, 4 exilis, III, reg rok ie ITI, 484 fallax, III, 484 glauca, III, 48 gracilis, III, 4 heterophylla, III, 484 integrifolis, mae 482 micran ok a, aoe aaa minutiflora spe = modes mcronat, es nana, III, 481 nemorenshe: III, 484 parviflora, III, 484 i 4 a bacelo(dee. IIT, 480 i 84 sepulte, Ill, 4 spatulata, III, nthe tenera, III, 484 Nemosceris californica, 357. neo-mext icana, 1V, 575 B pushoaedn pd ra 8 HE microcarpa, III, 162 seebney glu- tinosa, III, 16 eaciawhe colusana, I, 188 N. es ees integri- folia 51 ovata, os 51 Neowashingtonia fila- entosa, I, 345 N noche puberu- lum, Nephroohyllidium, EE; crista-galli, III, 366 Neslia, II, 291 paniculata, II, 291 Newberrya congesta, III, 297 longtloba, III, 297 spicata, III, 297 subterranea, III, 297 Nicolletia, IV, 250 s, IV, 250 683 nata multiflora, II, 684 ee III, 684 glauc ipo cinopitions no 684 multifiora, III, 6 méltivalvis = eg 7 ba mbaginifolia bige- qu vadrivalvis, in, 685 rustic Nigella damascena, Nitrophila, II, 67; 111;.3 occidentalis, II, 67 Nolina, I, 4 bigelow, “ a georgiana, I, 448 parryi, I, orta altissima, IT, 267 , LL, 267 Nothocalas sire TV-553 mergers): IV, 554 T. 163 mollis, I, 163 Nothoscordum aureum, T, 398 maritimum N be srs polysepalum, m: babes brandegei, I polysepala, IT, 17: NYMPHAEACEAE, II, 172 Obione bracteosa, II, 82 INDEX OF SCIENTIFIC NAMES confertifolia, II, 87 a, II, 82 Ochrocodon pudicus, iva tostomum, I, 412 oe oar californica, III, 1 alys Laatals LTT, 2 villosa i, nitida, III, 204 suffruticosa, IIT, 204 shelddoitihe Hi,'193 clav raeformis III, 206 co gnata, hive, ¥ heterantha, III, 197 ih T heterochroma, Abe a3 s, IIT, 2 hilgordit mt, ber” hir' 1H,:2 indy, is 187 clutei, III, 194. marginata, ITI, 195 purpurea, III, 195 osaiae age TIT, 202, abramsit, ny 204 707 exfoliata, III, 204 ignota, III, 204 jonesii, III, 204 minor cusickii, III, 20 scapoidea aurantiaca, Ill, clavaeformis, ITT, 206 purpurascens, III, 206 rsa, ITI, 206 strigulosa pubens, III, 202 tanacetifolia, III, 197 trichocalyx, III, 194 triloba ecristata, 96 venusta, III, 193 Onlifa ce californica, oraite IIT, 346 Omalanthus caniphora- tus, Omalotes camphorata, IV IIT strigosa, ITI, Ownacraceae, IIT, 167 708 Onopordum, IV, 510 acanthium, IV, 510 Onychium densum, L;23 rag Ar 1 au ssum, I, ae orrh 2 ope alternans, I, 410 oo II, 409 alvaceus, II, 4 409 opulifolius capitatus, Opuntia, III, 143 nthocarpa, my 145 pee hae III, 145 hotlines. III, 148 bra achyclada, III, 148 48 parkeri, ITI, 145 robustior, Ill, 145 ngelmannii littoralis, humistrata, III, 148 hystricina, III, 149 INDEX OF SCIENTIFIC NAMES incarnadilla, III, Seen intricat a, III, 15 indheimeri sol TIL, pn tola III, 152 mesacantha vaseyi, III, 1 missouriensis, III, 149 mohanennt ory 153 munzii, III, 1 occidentalis, in, 152 villet Ill, 1 145, 146 phaeacantha covillei, , 152 majavensis, ori 153 rigs EG Bea} ea canta, i, 149 rreee 146 raiesauel = III, ramosi sR III, 145 rhodantha, ITI, 149 ta megacantha, IT,.150 rubifiora, III, 152 aipite a 148 Orch porrfolnm, I, 479 um, 1, 479 man. Secueee I, 469 Orchi. dilatata, I, 473 elatum mil, 4 glondulosum, i, ao heckneri, II, kelloggi IL, 135 leanum, II, 1 longneialu, a. 134 sitifolium, II, 133 vee: 1,143) ecdyi, Oreocallis davisii, _ “WwW cd pss 598 suffruticosa abortiva, 1:5 mpsonii, III, 600 oo palitaretee, aa a eo ana: III, 2 Oreophila myrtifolia, I 15 Oreostemma alpigenum, IV, 32 andersonii, IV, 328 vulgar Onavan bicolor, IV, 389 mixta, IV, 389 Ornithogalum divarica- tum, > a: 1x10 Presi — III, 347 laremontensis, corymbosa, IV,7 ri, IV, 10 lu doviciana cooperi, V,8 latiloba, 1V,8& valida ,1V,8 mai ae IV, 3 praesent IV,8 multiflora arenosa, IV,8 pringlei, 1V,8 ino CEN porphyrantha, IV, 5 ramosa, IV, tuberosa, IV, 5, 10 uniflora inundata, minuta, 1V,4 occidentalis, IV, 4 purpurea, IV, sedi, 1V, 4 valida, ¥, Orobus littoralis, HH, Or cchaenacts, IV, 246 carpha, IV, 246 linearifolia, =‘ 241 lat 1-2 Paid nti a 810 attenuatus, III, 815 oldtiensis, 4 insalutatus, Bias Ey linearilobus, III, 814 lithospermoides, III, 813 olympicu pdchystachyus, III, 817 cuspidatus, III, 817 pallescens, eo rt parishii, III, 8 pilosus, III, oN pir dace Paes III, 824 monensis, III, 825 psittacinus, Ill, 825 nee rascens, III, 814 latifolius, III, 814 alailie ITI, 814 13 III, 813 schizotr aris III, 824 sonomensis, a 813 str ict us , I, 815 cuspidata, I, 136 A, 1135 Loeeeeee i, 1;135 hym eee I, 136 pape rembranace, I, 136 aitien a, I, ere eri, smadenia oe IV, 186 Osmaronia, II, 4 ce rasiformis, Pt Ne purpurea, _Osmunda incnelee lunovid: 1. 3,4 spic cant, I, 21 virginiana, I, 4 Ourisia californica, II, 529 OxXALIDACEAE, ITI, 8 Oxalis, ITI, 8 a Ill, 8 trilliifolia, III, 8 INDEX OF SCIENTIFIC NAMES Oxybaphus, II, 104 brandegei, Il, 106 u 104 1iz,2 pot inert ig 245 Oxyria, IT, 54 ine idea, II, emarginata, I : foli m cusickii, IT, 611 deflexa, IT, 613 luteola, II, 611 luteolus, IT, 611 mazama, II, 61 montana, II, kanoganea, II, 611 olympica, II, 6 oreophila, II, 611 parryi, II, 613 visci abn II, 613 O ryura chrysanthe- pacifica, 7 stauropetala, I, 377 rifida, II, 377, 378 pies fe Ae Pachistima, ITI, 54 canbyi, III, 54 myrsinites, III, 5 Pachylophis wept Paeonia, II, brow BT 175 gigan Palenlerte cornee, V, Pebener® debilis ser- ata, IV, Pauue Mr ileeaint III, 2 . Pie eh americana 1, 213 praise i211 osta chye, 1,212 mato, I = nerv. elat ree seidentalis, i ae iflor arizonicum, I, 112 barbipulvinatum, [3 Bowl re, 1,413 Be nihil tig dichotomum, ‘435 scopariu m, wa 116 ian longiglume, I, 114 ve ertiilatn, i138 ride, 1,1 psieso Be 9 230 pel si one seb 230 califor m, II, 230 heterophyllum rg ssifolium oe Prise 1,2 rho 23 II, 223, 629 Parentucellia, III, 800 floribunda, III, 800 viscosa, III, 800 Parkinsonia aculeata, II,4 sae? florida, microbll <7 479 torr 480 Poon ae pike II, Parnassia, II, 347 113 ‘- —— abeeiccas, pt nica, II, 558 simplifolia, Il, 558 mollissima, ved mollissim 1eglecta, II, parryi, II, 557 polyadenia, et 558 schottii, II, 558 SPIRO i. nag: epurvcarp 12 ert og II, — menziesil, er glabra 1,2 - uginosa, IL, 290 Parthenopsis maritimu 97 a vaseyanum, I, 111 virgatum pubiforum, Pastinaca, ee i sativa, III, 2 Patrini ia cera pa IV, 58 Paxistima, III, 54 5, 710 Pectocarya, III, 535 chilensis californica, 121; 53 gracilis platycarpa, iH; heterocarpa, III, 536 lin sao erocula, rs Bovaesinny ITI, 536 Ill, = penicilata, 15°536 heter pica iI, — plat ycarpa, Ill paddoensis, IIL. 8 805 m ii, IIT, 159 Pediomelum californi- cu ae 554 m, 11,555 parrintdahiy 11,5 Pellaea, I, 29 ndromedaefolia, I, 30 brachyptera, I, 31 na, wrightiana eae Peltiphyllum, TE, 355 peltatum, II, 355 Pennisetum villos 1,119 INDEX OF SCIENTIFIC NAMES Penstemon, Tit; 733 jg e Fe alsinoides, III, 742 amabilis, III, 746 ambiguus tharberi; Ti, 550 anguineus, III, 747 antirrhinoides, Hit; 769 microphyllus, III, sien hy III, 742 palustris, III, 743 austinit, III, 755 azureus, im, — 763 angust TTT, 761, jeffreyanus, III, 761 parvulus, III, 763 barbatus labrosus, FH; barrettae, III, 765 berryt, III, 768 bra cchyantins —? le breviflorus, ITI, 7 stabdlinpilod. III, 769 bridgesii, III, 764 cacuminis, III, 738 cardwelli, ry 768 carinatu conwrauentonia ITI, 756 i nae ait ITT: chionophilus, III, 738 761 caeruleo-purpureus, Lil, 738 seers eur ‘to globos br coreitofiae He 770 E770 vberalenis TIT, 770 IIT, 765 savagei, III, 74 sudans, III, 748 suffrutescens, III, 748 763 posse PW § dive ergens, ITI, 750 formos us, ITT, 738 f glaber occidentalis, 1H, 751 speciosus, Ets 754 ja ndulosa chelanen- s, III, 764 a sodailostis, III, 764 res eterodoxus, III, 742 stphelehnecs III, 742 hateiogitia III, 763 australis 763 palus, III, 760 ttatus, laevigatus, III, 734 seater Ill, 739 oo i, III, 769 leptophylins, III, 746 leuca 3 lepine ai; 765 — californi- s, III, 750 seoreus; Til, 747 lobbit TE 769 macbridei, se 760 I, 768 thom mpsonii, i, 768 geccneor 1, 738 sree sp: rn 770 minor, III, 747 rl, 751 monoensis, TH; 750 , 758 berryi, rupicola, IH, sonom oreganus, ITI, 747 ovatus, III, 746 pine storm; III, 746 pruinosus, ITT, pseudospectabilis, III, puberulus, III, 743 pumilus thompsoniae I pu rpus usii, III, 763 rattanii, ITI, 747 kleei, III, 747 minor, III, 747 recurvatus, III, 739 rex, 51 richardsonit, III, 764 roezlii, III, othrockii, ITI, 769 seorsus, III, 747 sonomensis 68 cee ti tf; gl pécio pet in II, 756 padre septentrionalis, III, 770 tho let ie td 750 ut heneta venustus, III, 764 washingtonensis, z, washoensis, III, 739 sima, ITI, 114 Dp $ es . ~ IV, 269 aphantochaeta, IV, 268 aurea, IV, 26 bellidiflora, IV, 268 exilis, lV, 268 aphantochaeta, IV, 268 discoidea, ae 269 grayi i, IV, 2 Peplis eanivicaié: ITI, 118 portula, III, 165 par veal I, 481 ens, I, 481 sone I, 481 repens, I, 48 pion Loti II, 473 m, II, 473 Prdciein satire cum, IV, 550 = TV, eg IV, 550 oa IV, 550 californica, III, 234 INDEX OF SCIENTIFIC NAMES gairdneri, III, 233 howellii, III, 233 psc a8 a III, 2 parishii, III, 2 a pir ba III, 234 Peri os a serrulatum, i 32 Peter: IV,1 californica, 1, 199 — Ae 7 dh orcuttii, IV, 199 panies palmer TV, 199 rothrockii, IV, 199 Persicaria alismaefolia, Hp 6d: chelanica, II, 62 covillei, IT, 63 sickii, II, 6 frenciscona i: 63 nee 63 ins a apts, 11 66 nnsylvanic, - 66 a la purpurata, II, 62 33 osteira laciniata, II, 3 pdr tree, II, 371 corym 5, IV, 454 bet certs dipetala, III, Plaid IV, 296 Fences abrotani- folium, III, 253 argense, III, 258 rine III, 258 ITI, 24 canbyi, III, 252 circumdatum, a7, 253 confusum, III, 250 cous 53 cusickii, III, 2 266 da P III, 258 donnellu, III, 263 euryptera, III, 249 [ TIE; 252 farinosum, III, 250 geyeri, III, 252 gormanii, III, 250 261 aberrans, III, 261 a Tl, 261, 264 has if leviouli, III, 252 2 I m, III, 260 arpum, III mileetiom IH, 261 ~ nevadense, Mt, latum, ‘i, 589 mudicon rs ellipticum, Hit, 2 calla III, 266 iriternatum alatum, ee 2 evifolium, se 266 pe arpum, III, 252 le ptophyllum, III, 266 macrocarpum, III, 266 platycarpum, III, 266 Peucephyllum, cens, 1 5 11, 571 conjurcta, IT, 595 711 crotalariae, II, 589 cusickit 582 cere leucephytie I, 579 miguelensis acanthominthoides, III adenophora, III, 519 argentea, — it, Se 493 austro Ill burkei, III, 500 AiZ [Pha californica, III, 498 bernardina, III, 498 otdensii TEE, $01 jacintensis, Ill, 498 circinata calycosa I; circinatiformis, IH, coerulea, III, 497 eremica e geraniifolia, III, 511 andulifera, IIT, 516 ETN ii, III, 511 grandiflora, IH, 514, 515 greenei, TIT, 505 grisea, III, 509 evmnoclada ese 518 oe heterophylla, eee 500 pygm ohiailens “8 eo heterosepala, IIT, 494 INDEX OF SCIENTIFIC NAMES eterosperma, III, 512 hispida, ITI, 494 brachyantha, III, 494 heliophila, HI, 494 lobata, ITT, 50 hydr ae iT. imbricata III, 498 Ill POS an oe en = 516 keckii, ITI, 511 anions? ITI, 516 lem i, TIT, 512 lenta, TIT 490 ] 6,414; — pias sepala, III, 501 u lineari 9 purpurascens, ik, hiteopurpurca, III, 516 lyon 492 magelanice angusti- ia, III, 500 ferruginea, III, griseophylla, Ill, ne 8 nemoralis, III, 500 ot do-hispida, Stirs IIT, 516 orogenes, pachyphylla, IIT, 515 palmeri, III, 508 rt III, 512 p i} s Le aay ul, 509 acger jis aceli ee Ss, a 507 pretviagia. inter- proce nuifolia, Tikes “i ormeet III, 5 = ITI, rt mbrosa, III, 493 valicole, He 507 vallis-mortae, IIT, 494 ampliata, Ill, 500 bernardina, III, 498. eta III, 514 whitlavia, ITT, 514 jonesti, ITI, 514 aco me cimae, is ied _weiserensis, TI, 5 — I, 183 rant dered IIT, 183 ea, I, brachystachys, 122 californica, I, 121 canariensis, 1 120.122 caroliniana, I, 123 erucaeformis, I, 179 oryz zoides, 1, on 120 me — acto morsa, * ‘ia ana re ahs pet spr taepal 381 imine III, 381 fie inti dryopteris, G feces 1:34 polypodioides, I, 14 neh pare californica, lutea Pinetorun, IV,6 ah a, IV, 5 ipterus multi- s, 111; 278 re) ae = Lona) ° a} = 7 — confus ordonianus, II, 384 helleri, IT, 384 lewisii, IT, 3 californicus, II, 385 platyphy ete II, 384 pumi 385 se lees eens TI, 385 stramineus, II, Philib aati hartwegit heterophylla, ty 4 SLs hirtella, III, 3 fe ibe pride Raven III, 373 linearis heterophylla, III, 373 hirtella, II, 37 hs aA a“ reas ee Phteum, I, 142 pratense, I, 14 Phlomis pare Ill, leonurus, III, 628 Phlox, III, 408 aculeata, IIT, sie vase urgens, III, 4 ustromontana, aed 411 12 longistylis, III,412 genes: III, 4 sees oe ITI, 412 go hrig occidentalis, III, 411 dolichantha. III, 409 bernardina, IIT, 409 douglasii, III, 411 austromontana, III, 411 Oo ae Ne: 413 humilis, ITI, 413 grayi, III, 409 hirsuta, IT I, 409 odii canescens, se = hoo III, 455 hus nis III, 409 lanata, III, 413 prasnae III, 411 linearifolia longipes, longifolia, III, 409 brevifolia, III, 409 calva, III, 409 da, 412 Seeronthifaia, ITI, 412 speciosa, ITI, nitic a; IIT, stan sbury i, 11L4 mosum, III, 480 paaiaces F238 incu wrcains et. 239 incurvus, I, 239 INDEX OF SCIENTIFIC NAMES Ph shale. 1, 531 califor . 531, 32 distans, I, 532 coloradense, I, 533 densum, I, 533 cues rum, I, incisa, ITI, lanceolata, III, 6 nodifl : canescens, "TIT. 6 reians, III, 613 a, III, 613 Phyllodoce Ill, Dregs bre ii, 3 paioh ns ni glandulifiors, IIT, 303 4 ogonum Iuteolum, Phyllospadix, 94 leri, I, serrulatus, C 95 torreyi, I Phymosia acerifolia, III, 84 longisepala, ee 86 II, 8 rivularis, I Physalis, ITI, es — pe 69 alkek III, 669 cardiashils IIT, 671 ct erp iss ia, III, 671 phylla, ITI, 671 digitalifoti, III, 671 rdifolia I greene I, 670 hederaefotia, III, 671 a, III, 669 pennsylvanica lanceo- III, 671 lata, , pruinosa, III, 670 pubescens, IH, alpestris, II, 286 mp As re II, 286 whberry i, II, 286 namintensis; IL 410 amurensis, II, 409 capitatus, II, 409 Ivaceus, IT, 409 basi ag sf 409 I, 409 Physo stegia ae Fit, ben Piar crassipes, I, 349 nto sa, TI, 483 amare biennis, IV, 208 canescens, IV, 208 c asplenioides. IV, 588 echioides, IV, 588 Picrothamnus deser- torum, IV, 412 Pieces engelmannii, 3 pizantcws, ITE TS3 Pilosella relicina, IV, Pilostyles thurberi, IV, 617 Pilularia, I, 34 , 163 Pimpinella apiodora, I, 235 gto nudicaulis, 11, 235 leptophylla, III, 228 parishit, III, 234 PinacEAgE, I, 52 californica, I, 57 713 commutata, - 61 ntorta eflexa, 1, 57 edgariana, I, 60 edul monophyla, 1,55 flexili albic nies i 54 fre at gg Phe ins pes 59 jeff itr 1, S7 3111, 365 prone rr = aS Pape Thi. lasioc Sante : lo phosperma, . 56 minima, nutta us, III, 296 viscosa, I, 2 714 gia sawsihs ai S72 arpus “TIL = llocaryoides, Ill, 571 arizonicus, III, 573 Samrat III, 573 fulvescens, ill, bi gracilis, III, 57 ursinus, III, te campestris, III, 574 nensis, III, 573 catolinensis, {I 573 chor a hickmanit, It, 365 cognatus, III, 567 DOL harknessii, Ill, 573 ypdieue sas. 565 wis cs i 565 3 micranthus, III, 563 strictus, TIL 561 e574, 575 parvulus humi ifusus, III, 575 INDEX OF SCIENTIFIC NAMES tener, III, 567 seer lg III, 574 to Hee etn S75 if 575 trac fini ae ae 570 uncinatus, III, 574 undulatus, IIT, 563 Planodes virginica, le 05 PLANTAGINACEAE, IV, 14 br , 20 califor ica, Iv, rs 22 caroliniana eriopoda, IV, 18 fastiee IV, 20 a, tV, 1 pinata te av Ef abra 2 hirtella galeottiana, TV As hookeriana californica, IV /21 oblenastohe he 18 pees: ae ston a pristela: IV, 20 cali ifornica, IV, 2 gnaphalivides, Ng 20 spinulosa oblonga, subnuda, IV, 17 Platanthera elegans, 1, leucostachys, 1, 474 stricta, I, 4 wrightit, II, 406 Pletyloma brachyp- Pi hee Peden scapi- gerum, IT, 2 pists temon, Ii} 223 ee sardie breweri, 249 lnaechl II, 248 pele ag Il, 249 a, hii 253 phe aa , II, 248 Pleuraphis jamesii, “igh I, 109 Pleuricospora, III, 296 fimbriolata, III, 296 longipetala, ITI, 296 leuropogon, I, 193 californicus, I, 194 refractus, I, 194 sabinii, I, 193 Pluchea, IV, 466 camphorata, Per 466 sericea, PLuM BAcINACEAE, lii3 Perit II, 344 Pneumonanthe affinis, IT S56 calycosa, DE & Bas new soa ~ zen sceptrum 356 setigera, ay ‘ie Poa, I, 197 absasiniae, 1 I, 186 a bul I, 211 cio I, 206 capillarifolia, I, 210 megastachya, I, 186 187 mu sehen c 206 nervata, I, 213 nervosa, I, 202 nevadensis, I, 207 olneyae, I, orcuttiana, I, 206 oxylepis, I, 185 pauciflora, I, 207 paucispicula, I, 203 ° > ie] — bolan- deri, III, 234 ostemma nyctagini- 7 Poinsettia eriantha I Polanisia, IT, 327 rave OLEMONIACEAE, GLE erryi, IIT, vid bicolor, IIT, 4! caeruleum, Te 398 INDEX OF SCIENTIFIC NAMES californicum, III, 398 calycinum, III, 398 tevisi, III, tricolor, 1346 98 scosum, III, 4 "Blister iT1L ss Polyanthrix hystrix, I, 254 pi tesa II, 158 depressum, II, 158 tetraphyllum, II, 158 Polyctenium bisulca par aa Li, 2 poniears: III, 21 fishiae, III, 21 Pha ty nT, 21 fishiae, IIT, lees III, 2 Po OLYGALACEAE, ‘aa 21 POLYGONACEAE, i, Polygonum, IT, 54; ,61 oe Tl, 62 um, IT, 59 emersum, II, erectum, II, 58 esotericum, II, 59 pare lapathifolium, Poy 66 incanum, salicifotium, 1 66 maj jus, Tz 2 pe ennsylvanicum, II, 66 persicaria, II, 6 rr riciasioieiees polygaloides, II, 59 tatum leptostachyum, II, 63 sawat 9 hast RE: spergulariaeforme, torreyi, ba oe Vivi afsal ope Ag ‘a wats: onit, Se 59, 61 succarinii, IT, 66 Polypa eee sericeus, V, Sew viae a 1,5 ypodium ‘anum, 1, 15 occidentale, I, 8 715 oreopteris, I, 15 pa achyphyllum, Ly J lutos sus, I, m scopulin Ponista casseiantia, PONTROERIACEAR, I, 349 Populus, I, 48 peesa tears IV, 95 sula, IV, 95 eximia, 1V,95 ulaca, ca Ie bs 8 aes 1135 wieder IIT, 119 mineus, heterophyllus, I, 86 hybridus, I, 84 latifo lius, T, 89 poo tides T, 88 716 [ Potamogeton] , 89 bejobates richard- ee wii s, 1, 88 proteus uc: I, 87 spirillus, I, 85 zizit, I, zosteraefolius, I, 89 POTAMOGETONACEAE, Potentilla, II, 427 a, II, 434 arguta glandulosa, II,4 convallaria, II, 439 =. oo a S rig Lom EN ce oO breweri, IT, 435 ose tata, aoe II, 4 ascadensis, I I ; one 417 fastigiata Poa er hy = “ INDEX OF SCIENTIFIC NAMES flabelliformis, II, 431 fragiformis villosa, hee igiat apidd. I grayi, IT, 435 Gs IT, 430 hallii ,U,4 hol opetala fastigicia, II, 4 howellii, II, 419 kelloggti marinensis, II, 419 rula, II, 417 longipedwnc ulata, II, 4 Fe Se II, 434 macrantha uniflora, II, 4 macropetala, IT, 430 9 a, IL, 4 monspeliensis, “id an muiri, II, 4 multijuga, a oe 438 senor 1; rape sob - 435 procu II, 429, 443 pseudosericea, IT, 430 a, II, 417 39 purpurascens, LE ving ia SET; 42 recta, 1 431 Ka Net II, 431 bea. -_ = fp ae a) — a Oo millegrana, II, 429 rosulata, II, 434 santolinaides, II, 422 saxosa, IT, 434 cesiaaea: Il, 4 419 shockleyi callida, II, 423 sibbaldii, i. gn noloba, IT paren es i, 431 te nu ilo »ba, uniflora, 423 Rede IT, 440 versicolor, IT, 438 villosa, IT, 434 viscidul wilierak it eee I, 440 Poteridium occi identole, IT. 445 Poterium sitchense, iG Prenanthes, IV, 594 alata, IV, 594 exigua, IV, 568 ha alpha: IV, 594 juncea, VW, a 569 594 Pri id Pept AE, 2s as Pritchardia filan istana, 1V louisianica, ie J lutea, IV, 2 Prosartes hookeri, I, 456 pagel trachyan- a, 1,457 menziesii, : 455 regana I, 475 chi Tena glandulosa, ui. psi II, 478 emor nity oe sa, I, 478 juliflora SF eave II, 478 odorata, II, 478 gana, II, 466 Psacalium strictum, IV, 458 Psathyrotes, IV, 456 annua, IV, 456 n inc 36 ramosissima, IV, 456 schottii ,4 IV, 232 Pseudotsuga, I, 63, 529 douglasii, I, 64 macrocarpa, 1 , 64 lindleya ebihhes: IV, 492 oreganus brevissimus, IV, 492 oreganus elatior, IV, 492 oregonus, IV, 492 tenellus, IV, 492, 493 tenuis, IV, 493 tenuis, IV, 493 Psilostrophe, IV, 194 castorea, LI, douglasti, I1, 55 oe purshii; II Ss ’ Psychrobatia pedata, II, 455 Ptelea;-J 11, 17 baldwinii crenulata, Powe: 18 nulata, III, 1 Jrifoliate, ce 18 pir se as lr PTERIDOPHY vTa, 1,1 Pteris andromedaefolia, bs < ropurpurea, I, 29 caw ” * 23 fee sca 1. 22 stelleri, Pterochiton occidentale, 11,8 Pterostephanus run- cinatus, IV, 58 . Pteryxia, III, 2 oeniculacea, IT Ptilocalais pin 9 IV,5 maior, IV, 556 nutans, IV, 556 Pvt sie TV, 556 gira: anihemoides, ike 219 Ptilonella laevis, TV.t INDEX OF SCIENTIFIC NAMES scabra, IV, 192 subcalva, IV, 192 Ptilophora major, laciniata, IV, 556 nutans, IV, 5 Ptiloria canescens, exigua, IV, 57 deanei, IV, 574 filifolia, IV, 571 lemmoni, I, Pugiopappus bioclowi, IVA Pie ons ciliata, Ti, oblongifolia, III, 546 a Fi 451: TEL pretends ml 451 Pycnanthemu III, californicum, III, 658 Pyrethrum halsamite; 396 acetoides, IV, 396 Peer iat IV, 395 thenium, IV, 396 asarifolia bracteata, III, 288 incarnata, IIT, 290 uliginosa, III, 290 m minor, III, 291 pallida, I IT, 290 Pyrrocoma apargioides, 274 ligt arguta, IV, 2 balsamitae, IV, 2 c ersthropappa, W. 272 folios ial pe on gossypina, IV, 2 grindelioidesi, = 286 roi 0 rt gifolia, 276 tenuicaulis, 1V, 275 ——— JV, 274 inihora 274 Pirie pal oa II, 470 fusca, II, 470 malus, I1, 470 pits 8 II, 470 rivularis, II, 470 slichensis, Il, 470 californica, Tl, 470 Quamasia, 1, 415 azu I, 416 Ae Quamoctidion froebelit, pile net, II, 106 multiforum obtusum, a, 1, 52 prapioniou L co tinctoria californica, I, 517 tecemieae: 4,521 turbinella, I, 520 wislizenii, I, 517 frutescens, 1,517 Radiana petiolata, EH, —_ TI, 276 ta, II, 279 patie Tl, 279 columbiae, II, 278 paniculata, IV, 426 pringlei, [V, 428 718 [Raillardella] pringlei, IV, 428 Raillardia. Petite IV,4 piles geek Raillardi loRets muirit, IV,4 sca abri at Pa pacha annio i: niera sivicta, eons 458 pred secunda, ITI, 291 Rémana capitata, II, 64 clevelandii, III, 642 vaseyt, ergs 23 sisenii; HE 7. bstiie sti II, 174, 629 Ranunculds, II, 197 acris, IT, 198, 199 aliinis cardiophyllus, }; alatus; EV; a alceus, II, paket: II, 208 alismellus, 11, 208 egii, II, — vg bakeri, IT, 2 areas II, 214 osus, canus, II, 200, blankinshipii, II, 200 laetus, ludovi ovicianus, II, 200 capillaceus, II, 214 ciliosus, II, 202 INDEX OF SCIENTIFIC NAMES cooleyae, IT, 211 cusickii, II, 210 cymbalaria major, E iy saximontanus, II, 211 detphinitotns, eee ersus, a nee te rrestris = 1t,.2 falcatus, II, 215 filiformis ovalis, II, 207 flabellaris, II, 211 0 , 207 hispidus, IT hydrockaris ee fa koe IT, 208 hystriculus, II, 211 inamoenus alpeophi- lus, II, 2 lobbii II,2 tadowiiiets IT, 200 macounii, II, 20 m armorarius, II, 202 s, II, 203 eg ss 8 EE, Vit petite Il, 205 entalis oreganus, oresterus, orthorh hallii, II, 203 peda phyllus, 11, 206 pennsylvanicus, IT, 204 bolitus, II, 203 ramulosus, II, 207 repens, II, 200 Bee hens) F207. rudis somolifaline veh 208 sa P tridentatus major, liv? triternatus, 11}207 sativus, II, 2 Rapuntium dortmawnis; I kalmii, IV,9 Razoumofskya abietina, (ih; 327 Rhagadiolus creticus, IV, 568 RHAMNACEAE, crassifolia, ITI, 63 cuspidata, III, 63 occidentalis, III, 6 rubra, III, 62 tomentella, III, 63 cathartica, ITI, 62 colubrinus, III, 79 crocea, IIT, 6 3 ilicifolia, it [, 63, 64 pilosa 63 laurifotia, Ill, 62 ensis, III, na a, Ill, 6 ee III, om oleifolia, III, 62 S-oes é Rhodode bdceh. Ill, 299 albi , III, 299 californicum, III, 299 leachianum, macrophyflurt, III, 299 occidentale, IIT, _paludos sum, III, 50 nomense, III, 299 RI todothannus leachi- s, III, 302 clophsla I, ss I, 50 avers, m1, 50 labra, ITI, 5 gi ctiden fale: Ill, . integrifolia, IIT, 5 Tit 5% trilobata, III, 51 Rhyso pterus, IIT, 239 peatige III, 239 Ribes, IT, aceriflium II, 391 391 co pa sare ‘IL 403 deductum, II, 394 wielaowce Fut II, gracillimum, II, 392 migratorium, , A mo m, II, 3 enum, it at tum, Li, oligacanthum, IT, 402 oxycanthoides lacus- t 390 II, rpusii, II, 397 parishii, II, 398 lum, IT, 390 ped tok ak # 396 roeshi. Tl. 403 INDEX OF SCIENTIFIC NAMES cruentum, II, 403 tiv bate m, 94 glutinosum, II, 394 malvatewn, IT, 395 Merrie II, 399 Re gt $,1 : Q2s8s = s 2: rx) wn Riddellia cooperi, Rigiopappus, IV, 234 leptocladus, IV, 234, 308 longiaristatus, I longiaristatus, IV, 234 princi 15 seudo-acacia, IT, 562 Robsonia speciosa, Rochelia patens, leib pees mendocina, III, 531 sitch austriaca, II, 2 columbia -: II, 278 curvisiliqua, = 278 iirc iH; Seu ON TE 276 obtusa, II, 278 Lerigvind e — palustri 278 earns II, 278 pacifica, II, 278 sinuata, II, 276 subumbellata, II, 630 pubescens, II, 462 ltramontana, II, 462 centifolia, II, 458 columbiana, I1, 459 copelandii, II, 462 covillei, IT, 46. crenulata, II, 465 ultram yainacensis, II, 4 RosAceas, if, 407.6 Ra IIT, 620 Rotala, ITI, 164 osior, III, 164 verticillaris, IIT, 164 Rottboellia cylindrica, pannonica, 1,238 1:7 ubacer parviflorum, IT, 455 eee ae 22 Rubus, IT, _ ardinus, eaeA 457 astwoodianus, II, 458 Hdaticies: II, 457 719 fruticosus, II, 454 helleri, 11, 457 hesperius, II, 457 lacer, II, 45 laciniatus, IIT, 458 lasicoccus, Il, 455 enziesii, Il, 457 ursinus, II, 458 Il, 4 Rud beck IV, 116 8 priemcnng IT 50 patie eel hc “s 54 gracilescens, II, 54 persicarioides, II, 53 II, 50 salicifotius, II, 51 nticulatus, IT, 51 _nontigenis, 12-5! eus, II, 51 sbinosas, 5 IL, 49 r venosus, IT, 720 —— viola 1251 Ramalioe oa Ruta chalapensis, III, 19 RuTACEAE, ITI, 17, 19 panes 1,279 ee a, 1; 28 aurea, 1,2 a3 capitis I, 280 Sabina, I, 70, 75, 531 I californica, I, 75 occidentalis, I, 76 scopulorum, I, 7 tahensis, 545 Sagina apetala barbata, II, 145 ciliata, II, 145 crassicaulis, If, 145 linnaet, II, 145 oc ccidentalis, II, 145 cumbens, I, ea LH 81 I, 394 par a, 119 bigelovii, IL, 91 — essa, II, 94 aea, II, 91, 94 iherbacea II, 94 herbac vehi 94 a, II, 91 senile Il, 91 pacifica, II, 91 subterminal, II, 91 sr 487 — 507 reso poh I, 490 arctica petrea, I, 500 stoma iy 1, 492 + ores F 7 hg Be I, 489 austinae, I, 502 bakeri, 1, 497 barclayi, 1, 498 bebbiana, I, 506 perrostrata, I, 506 INDEX OF SCIENTIFIC NAMES bella, I, 503 bigelo it, 1, 49 bolanderiana, I, 493 brachystachys, 1, 505 folia, I, 489 dru onitiuna,; I, 503 castwaddin e, I, ‘49 9 — uy 507 ys eaten. ss 505 fluviatilis, I, 492 goodding i, 1,4 490 age 491 leuco ndmudeaites: I, 491 hookeriana, _ 500 T, 500 lyallii, I, ro i fi ie L; 92 melanopsis, I, 4 bolanderiana, 4 gee monica, I, 501 ontanus, I, 507 mutta 1.565 chnophorg, - _ planifolia - pu ne ae e petal osha ho : ‘ei pseudomonticola, I, 497 ee es, oe I, 496 eg idahomaie 497 Saicheoa ae 672 oi III, 672 cone atriplicifoti, II, 74 hyssopifolia, II, 90 kali tenuifolia, II, 96 pestifer, II, 96 platyphlla, II, 74 II, 9 Salvia, III, — 639 mbar stan: hg ok dorrii, III, 639 carnosa, ITI, 639 gilmanii, III, 639 eremostachya, III, 641 funere a, III, 638 glau a, IV, 44 Pes anndensis mexicana, IV, 44 coriacea, IV, 44 veo A 44 ferax,.1V,4 fimbriat, ig 44 a, 1V, 44 zonica, IV, 4 neomexicana, oe intermedia neomext- cana, IV, 44 neomexicana, IV, 44 nigra, IV, orbiculata, IV, 44, 45 glabra, IV, 44, 4 puberula, lV, 44, 45 ae arborescens, IV, 4 ches arborescens, callicarpa; IV, 46 maritima, IV, 46 mcelanocarpa, IV, 45 ee IV, 46 seminata, IV, 45 eae IV, 44 trifida, IV, velutina, IV, 4 Samolus, ITI, a og americanus, III, 3 floribundas, III, at valerandit, III, 342 americanus IIT, 344 _ dbergia whitedii, I, 306 Suameeis II er nadencis latifolia, 445 ratbfatia: IT, 445 arguta, III, bipinnata, III, bipinna tifida, iL ae flava, III, 2 Roeinae iY ee rassicaulis, Hi 221 Il, nemoralis, IIT, 220 serpentina, oe 221 tripartia, III, tuberosa, ITI, oe SANTALACEAE, I, ead Santolina, IV, chidecrahie IV, 291 vace pe 8 As vs Saracha nana, III, 668 Sarcobatus, IT, 94 bai yi, TL, s, II, 94 Sarc oslemma hete ero- phyllum, IIT, 373 hirtellum, III, 373 Pande nia, II, 32' SAR ACENIACEAE, tie 329 ¢ aus riata, fey 99 Satureja, IIT, 646 chandleri, III, 646 INDEX OF SCIENTIFIC NAMES douglasii, III, 646 poet oh I, 485 Saururus cernu ‘eed I, 485 Saussurea, IV, alpina cordata, IV, 509 Savastana macrophylle, li? odorata, I, 124 Saviniona rer iad ia flora, III, 1 cletaieea, ane 109 rot angustifolia, IT, 363 breached s, II, 361 austromon ntana, vespertina, IT, 361 tiers nen pssgtty , 36 rugine: macounit, IT, 361 nivea, II, 361 fragarioides, II, 359 fragosa, IT, 3 daytomaclota: 3 5) gracillima, II, 366 granu ulata, II, 357 cant, themifolia é in- tegriflio ao 362 lyall lax wnalecetea a 354 , 360, 366 odontophyila, II, 360 nisciticagg be 359 ore rhea err ae, if as oregonensis, IT, 359 paddoens sts, II, 360 porolifoli, II, 3 355 ranu Sat! 350 virginiensis, ba et californica, IT, 365 SAXIFRAGACEAE, IT, 349; Saxifragopsis fragari- at ropurpurea, IV, 64 columbaria, IV, 64 Scandix, III, 224 peti gine tai 224 Sc heuchrera, 197 s, I, 97 B ecipatshcidcn Oe 1,9 Schinus molle, IIT, 51 Schizonotus discolor, II, 414 Schkurhia neomexi- cana, Schmalizia trilobata, 11.5 Sch inidtia subtilis, TI, 143, 144 S¢ ae II, 246 linifolia pinnata, es Espa ag so i 412 album, I, 412 721 bracteosum, I, 413 Schoenus, I, 281 albus, I, or aculeatu capitellatns, I, ca ‘1.2 Gevicie oe Schollera graminea, I, 349 Schroeterella glutinosa, III, 16 p. chubertia semper- virens, I, 6 Scilla pomeridiana, I, 413 Scirpus, I, 269 acicularis, I, 264 cuminatus, I, 266 acutus, I, 274 oc lenticularis, I, dct aly Tr 271 noes a: fF. cane fluviatilis, iy micranthus, I, 278 mi robustus cohipactus. peladoces, 1.25 rostellatus, I, 266 z Oo sylvaticus di igynus, I, 276 tatora, I, 274 EL EPS is 722 apa acon us, ITI, a Sclerocactus, II L157 polyancistrus, III, 159 Sclerocarpus exigua, I =n vochioa oem Sitooia hispanicus, iv Scopulophila, +H es pee et 55 7 Sco czonera, IV, 586 glatifolia, IV, 586 his t 71 californica, vt era cat it. da, i, m floribunda, ne Ae iepeonet: ITI, 771 ra, III, 771 IIT, 770 SCROPHULARIACEAE, TIL. 620 pada Poe sq IIT, 623 austiniae, III, 623 canescens, III, 623 antirrhi leraieain aaa TIT. 623 sanhedrensis, le 623 Patsy ITT, 6 , INDEX OF SCIENTIFIC NAMES bolanderi, III, 621 austromontana, III, 621 californica, III, 623 epilobifolia, III, 621 om ulata, , 621 riflora, III, 624 inar foi, ITI, 623 III, 6 tuberosa veronicifolia, U, 623 Se yondasatey: sere il, II fasciculata hallii, niv 46 oblo ngorhixeni, re 339 obtusatum, II, 3 boreale, II, an oreg II, 343 pumilum, IT, 331 pep aig II, 346 Fite ella, 1. 46 oregana, I, parishti, 1, 47 scopu , 1,48 struthioloides, I, 48 baronet I, 49 i, I, 49 INELLACEAE, I, 46 Selinocarpus, II 102 diffusus, IT, nevadensis, II, 102 elinum acaule, 1D il Bey 6 Re WA benthianis. PPL J278 canadense, III, 273 det ae ETT 243 hook ri 273 Senebiera didyma, Ib, 263 Senecio, IV, 430 IV, 44 subnudus, IV, 444 werneriaefolius, austiniae ,1V, 450 balsamitee narent: ensis, 1V, 4 bernardinus a sparsilobatus, oe 7e pursh ianus, ee 442 caulanthtfoliu. I chapa acensis, IV, 448 , 43 scorzonella, IV, 440 crassulus, IV, 436 cusickii, 1V, 436 cym ee aati sels. IV, 4 endinus, IV.4 eurycephalus, W. 450 austinae, IV, 450 focciferus, IV, 578 fodin m, IV, i Alec eb TV. 4 BY rophilodes, fates IV, 448 lithophilus, IV, 442 re es, IV,4 by Haskihik IV, are pacificus, IV, 4 etiforus, 1V, 448 lonceotars. are 436 e,I poeneets. gy ow leucocrinus, IV, 442 ochroleucus, IV, 438 lyallii, IV, 444 me ndocinensis, I mesadenia, IV, 438 serge IV, 453 4%. multilobatus, IV, 450 multnomensis, IV, 446 nelsonit uintahensis, — bsteri, IV, 440 occi ideudalis, IV, 434 saxosus, 1V, 444 pe yereremnry om 440 a, IV,4 “admirabilis, “ay, 436 altior, IV, 4 ‘duties. ee iss integriusculus, IV, 4 sdiede Bees IV, 436 spatuli speculicola, 1V, 444 sphaerocephalus, 40 sngustflis, oy wa neces ri Sent, subv estitus 7 IV,4 tr ichophyllus EAv; ae trigonophyllus, : gonophyllus, IV, 436 wintahenss, se 450 vaseyt, IV, 4 vilgattl IV, 430, 452 websteri, IV, 4 panei es I whippleanus, IV, 438 SENECIONEAE IV, 416, 456, 464 te Le greggii, 475 catia yi 6 pitas 7 70 sempervirens, I, 69 washingtoniana, I, 69 wellingtoniana, I, 69 INDEX OF SCIENTIFIC NAMES Serapias gigantea, helleborine, I, 478 oregonensis, IV, 3 califor rnicus, IV, me a be be sipet, paces 9 rs 323 S pile sige califor- nica Sevicotha pare II, 4 S pe te glabres- cens, II, obovate, nck Si pitt gec me aes. 1°25 ey yeoman alpina, arvensis, IV, 516 “ ii ee I, 562 sonorae, Il, 5 asco II, i rpa, IT, 562 A pselk Leiocarpum, TTt;2 pan III, 266 tritern natum:; III, 266 Sesuvium, ITI, 11 portulacastrum, 11 Sieyos angulatus, IV, 68 lobat peices LW. 68 Sida, III, 98, 110 abutilon, III, 83 pr IIT, 83 ristata, IIT, 110 delphinifolin, 105 soliseophe a, III, 98 grossulariaefolia, III, 9 hederacea, III, 110 alvaeflora, calycosa campestris, OL, 106 confinis, III, 105 diploscypha, TII, 98 minor, III, 98 hartw weit III, 101 ‘Gan ella mite 103 II, 105 multifida, III, 106 neo mexicana, III, 105 cusickii, ITT, 1 parvifiora, III, oe thurberi, a pan 05 pedata, III, 1 ramuneulacea, ne 102 — ITI, 103 na, III, ainda IIT, 102 rhi — III, 101 r I, 105 virgata, III, 105 vitifolia, III, 106 ideranthus gooddingti, IV, 271 gracilis, I IV, os junceus, IV, 2 Sieversis, II, 449 campanulata, oe ss canescens, II, 4 aperta, II, 16 723 behen, II, 160 bk, 167 mmondii II, 166 etiam 167 engeimanni behrii, oe | iisect, II, 166 pec platyota Siliquastru -” occiden- tale, II, 478 Silphium laeve, 1V, 105 724 Silybum, IV, 509 mar m, I SIMAROUBACEAE, III, 19 i ia, III, 45 guinkets, ATT, 45 ca IV, 125 a Hr ee IV, 124 Sinapi 271 a, II, 274 Sophos is floribunda , III, 4 hallii, 111, 431 altissi IT, 266, amphibium, II, 276 palust , 278 re, 1 canescens californi- cum, II, 269 plese Hae al II, 269 curvisiliqua, II, 278 diffusum, II, 267 jaegeri, II, 267 hartwegianum, II, 269 incisum, II, 269 eastwoodiae, 1, 467 elmeri, I, 469 flavidum, I, 468 funereum, I, 467 pabstog? - “gpl I, 467 green. 7 Ralophyiions I, 466 hesperium, 1, 467 ida 6 entosum, I, 466 Segetum, I, 4 septentrionale, I, 466 Sitanion, I, 253 anomalum, I, eviaristatum, I, 254 brodiei, I, 252 INDEX OF SCIENTIFIC NAMES pia depnae 254 cinereum, I, 254 elymoides, I, es 254 m, 1,2 ngu sti ifoli wm, III, 231 cicutaefolium, III, 231 pale ITI, 238 erectu 23 nie erophstum, Baer eee III, 231 pusillum, III, 231 rigidius, TI, 24 is uniflora, I, 452 a 1 454 race bra achystyla, A; a, 1, 459 rotundifolia califor- nica, I, 459 Se myrnium aurea, Ill, 241 cordatum, III, 241 cupuliferum, III, 680 douglasii, III, 676, 677 dulcamara, III, 676 een TE 681 flavidum, III, 681 di wee ie — 680 REL: tenuilobatum, — 677 texense, III, 6 triflorum, aoe umbelliferum, 4 680 califor rnicum, III, 681 algida, IV, 296 bellidifolia, 1V, 296 perce Me 292 aper Vo eect os 292 canadensis IV, 292, 294 elongata, IV, 292 salebrosa, IV, 292 subserrata, LV .292 a, IV, 292 sale alutinos sa, IV. 2 a,1V,2 oD ty 292 major, IV, 29 V, 296 hu saline, so ha ery Wn? ana, 1V, 296 lanceolata major, IV, 2 one Wy. 292 caurina, IV, 292 elongata, IV, 29 subserrata, IV, 2 missouriensis, IV, 294 tolm a, IV, 294 mutiradiata, IV, 296 copu ulorum, E¥; — educa ilis, IV,2 arvensis, IV, 590 uh 1ginos Sonnea hevbacthe TE; $73 hi. spida, TE 523 rs iscosa, II, Sorbarle pa ps hl II, 414 Sorbus, IT, 46 californica, I1, 470 ascadensis, II, 470 domestica, II, 4 cidentalis, II, 470 sitchensis, II, 470 talifornica,.11, 470 densa, II, 470 Sorghum, I, 108 eh seco e, I, 108 SPARG ANIACEAE, I, 80 2 multipedunculatum, I,8 Plisetia I, 174 “gfe m scoparium, I,5 br WANS IT, 519 S| — bryophora, S ahr biflora, perfoliata, TV, ~ salsuginea bracteata, Tt villosa, TL 356 as perian thes unalaschen- sis, I, SPERMATOPHYT a, I, 50 S; Se goi Siochmaniad, I, 643 cling, III, 6 monticola, III, 88 INDEX OF SCIENTIFIC NAMES rosacea, III, 88 rugosa, III, 88 angustifolia cuspi- eh a; ev edlensis. III, 87 variabilis, III, 87 er sea III, 88 ex vilis fa patos IIT, 92 pest III, 91 i, III, 92 III, 92 nesiotica, IIT, 94 nuttallii, III, 94 fendler californica, III, 87 fremontii, III, 95 ce ereophora, ITI, 95 exfi a; 115,95 Phi et ty III, 90 subrhomboidea, ne enntica. III, 94 orbiculata, III, 91 clem entina, Il, 95 1H, stellata, III, 88 subrhomboidea, ; Sphaeromeria cana, IV, potentilloides, 1V, 398 Ss di sa salsula, T, GY S ohare alys- soides, III, 200 alyssoides ‘macrophyl- 0 parishii, III, 202 chamaenerioides, III, 200 contortum, III, 202 flexuosum, III, 202 micranthum, III, 202 exfolia a III, 204 nevaden , III, 200 orthocarpun III, 202 stl: ermum 204 irae 11,197 strigulosum, I 02 utahense, III, 2 veitchianum, III, 204 Sepesmhel I, 169 _ lo = I, 169 poe Brin ade bade AIT, 273 pe aesatiig III, 273 m, III, 2 RY Piesia oasenne 11 een hl, 613 Sp in acia: oleracea, II, 75 aruncus, II, 413 corel jee 411 ro. ca spies sy 411, 412 capitata, 409 i Bb IT, 412 corymbosa lucida, II, 411 cuneifolia, II, 411 dens oe; 410, 411 411 glabrescens, II, 414 douglasii, II, 411 2a ye tt iipendua, t “ati helleri henderson 1, 411 lucida, I I, menzi milo, TI, 414 oc I, 415 pany am Neuiethore: pectinata, II, 413 pyramidata, II, 411 I, 47 Spirodela, I, 346 polyrhiza, I, 346 725 Ss Lintiliede ng occiden- s, II, 91 Sporsbolus, . 144 airoides, I, 145 asper r,1,1 is asperifolius, 1,145 bolanderi, I, 20 6 composite iy 145 44 jones} pulchella, II, 1 pulcherrima, IT, 629 umbellata, II, 129, 130 caudicifera, 130 montana, II, 130 Sprengeria flava, it:2 minuscula, II, 259 watsoniana, II, 259 Stachys, ITI, 630 acuminata, III, 631 ajugoi II, 634 pinch III, sa lei el rgit, III, 632 littoralis, III, 6 malacophylla, it 634 sestnapes leptostachya, ITI, 6 Rear pilosa, rigida, III, 633 726 + hy - Tanata, III, 634 quercetorum, ITI, 633 rivalaris III, 634 riv s, IIT, 634 opianine ITI, 632 63 I,6 ve ronicefolia, aa 634 vestita viarum. vi poe et maareniza: 254 is oe? II, 240 confertifiora II, 240 elata, heterophaila, II, 240 pinnata 240 Pinar oe 240 rara, II, cagieks: 1 confertifiora, ss 240 retica californica, 1 z; armeria, IIT, soi pies hal PT 3 limonium californica, II, 344 Steironeme ciliatum, ny cidentale, III, 332 lacviigaiiten, UIT, 332 Stenactis glauca, IV, 350 ramosa, lV, 373 Speciosa a: 348 strigosa, IV. Stenanthella occiden- INDEX OF SCIENTIFIC NAMES alis, I, 374 Stenochloa californica: & Sten Me I, 267 capillaris, I, 267 Steuonepetl interior, S linearifolia, IV, 283 i sie lor, IV, 283 ee 278 kennedyt, IV, 278 lanug: IV, 278 li Ss an IV, 283 interior, IV, 283 lyallit, TV, 277 stenophyliue: IV, hl St SS tips per IV, 5 blairi 81 cchovincea, IV, 570 a, IV, 572 coronaria, IV, 573,574 pine hd 574 WV 372, 573, 574 runcinata parishii, 7 nid +, 1V, S72 Stevia Seer tee iy ae 248 aris, IV, 2 finenettoli ba: III, 3 paucidentata, i, og spinulosa, ITI, 2 ante ge 1, 5 Stipa, I, 1 loo mor, : 137 californica, I, 1 ch eek vine. I, 129 hymenoides, I, 136. kingti, I, 136 fie ie lemmoni, 133 iy hra, I, 131 viridula lettermani, Fist sabdscou: me 129 robusta, I, 133 williamsii ; L; 132 Stobaea heterophylla, Streptanthella, TE; 253 sid Bice 1, 246 poe idus nplasiceutts! 1;-253 rg hs hesperidis, II, 249 callistus, IT, he campesteia: I I, 247 bernardinus, aT. 247 , 247 i 1, 2 254 lem 4,11; 255 crassicauli ia TE, 255 pulchellus, IT, 249 glaucus, II, 257 gracilis, II, Ate hallii, 11, 2 hesperidis, eA 249 resect Ag | IT, 250 hispidus, II, 249, 250 howell, II, 243, 248 1 55 1 ps egies scale I}, 25 pa II, 250 parryi, II, os prittacs II, 2 polygaloides, ‘T 251 245 II, 478 Si ieee californi- cum, I, 4 Struthiopteris, 1,21 spicant, I, 2 Stylocline, i 486 acaule, IV, 495 Stylomecon, II, 228 heterophylla, TI, 230 eee elatus, ede aT IV, 565 Stylophyllum, II, 337 idum, IT, 338 viscidum, IT, 3 hs