PHYTOLOGIA Designed to expedite botanical publication Vol. 11 November, 1964 No. 1 CONTENTS MOLDENKE, H. N., Materials toward a monograph of the genus Te eee ane oe ve ee OSWALD, F. W., A new color form of Stachys from ante Island. : wees OD MOLDENKE, A. L., Book reviews........ oe ae Published bby Harold N. Moldenke ne Alma L. Moldenke re oe a5 Bleckewk een — : Sg BE gee _ Yonkers 5, New York, USAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXUII Harold N. Moldenke ENA PLATENSIS Spre Additional itt emean Rév. Hort. 1896: 188. 1896; Kuntze, Rev. Gen. Pl. 3 (2): 257. 1898; Haage & Schmidt, Cat. Général 93. 1898; Chod., Bull. Herb. Boiss., sér. 2, 2: 818» 19023; Chod., Plant. Hassler. 9: 197. 19023; Ed. Rodigas, Bull. Arboricult. Belg. 1902: 113. 1902; L. H. Bailey, Cycl. Amer. Hort., ed. 1, 6: 1912 & 1916. 19023 Briq., Arkiv Bot. Stockh. 2 (10): 9--10. 190k; Briq. hod, & Has sler, Bull. Herb. Boiss. *,s sér. a he: 1056--1 057. 190k; Briq. in Chod. & ners ere Plant. Hassler. 10: 477—-li78. 2505 Macloskie in W. B. Scott, Rep. Princeton Univ. Exped. Patag. L ley, Stand. Cycl. Hort. 6: 3445. 1917; Sanzin, Anal. Soc. Cientif. Argent. Buenos Aires 88: 98, 112, & 13h. 1919; Molfino, ne 5: 285. 1921; Olmsted, Coville, & Kelsey, Stand. Pl. Kaneo, ° 521. 1924; Hegi, Illustr. Fl. Mittel-Eur. 5 (3): 22)0. 1927; “aa ville & ge Rep. Bot. Exch. Club Brit. Isles 1927: 452. 1928; rule Urug. 106. 19305 Seckt, Rev. ‘Univ. Nac. Cérd oba Agron aT 202 193k; L. He & Ee Ze Bailey, tortiia: new rev. + Ry 631 & 632. 1935; Dermen, Cytologia 7: 161 & 162. ae. He Se rover Kew Bull. Misc. Inf. 1936: 9h. 1936; Troncos ——e ana 3: 53 & ah. «19375 pigs Revist. Sudam. Bot. h: 186— 19373 P. C. S » Field » Publ. Bot. 18: 1015. 1938; ee ke, Alph. Serer Common Names 32. 19393; L. V. Barton, Contrib. Boyce Thompson Inst. 10: 01, 410, 1], E08, & 25. 1939} Beale, Chron. Bot. 5: 517. 1939 Parodi , Darwiniana lh: 55. 1940; Beale, Journ. et. 0: 336-—-3)0, 32, "348, 3 350, 35h, Fe =i 1940; Moldenke, Suppl. List = pets Names 17 & 23. 1910; idenke, Suppl. List In- lina h: 18—21. 19423; Moldenke, Known Geogr. Distrib. Verbenac., {ed. 1], lh, 7h, & 102. 1942; Moldenke, Alph. List Inval Names 6 & Lownie 3 19423; Herter, Revist. Sudam. Rot. 7: 225. 19433 Schnack & reo Darwiniana 6: {71 473, & va 194; Rosengurtt, Estud. Prad. . Urug. hye 30 30. 19h Schnac Covas, Darwiniana Ts 7h: 72. ihe; faraey Bol. ° ee Bot. 1: ji ot 195; Ind. Taxonom. 1 (1h- 3 ate & Janaki Chromosome Atl. 270. 195; a Phytologia 2: 87 & 116. Scie arton & ae © Sans Inst. 1h: » 2h6, & 252. 196; p ahaa ge) - Prad. Nat. Urug. 5: 395. 1946; Augusto, Fl. Rio Grande Hg k 232. 9iby Moldenke, Alph. List Cit. 1: 18, i, 2 50 “Te, Pes 85, 87, , 212, 219, & 264. “19h6; Moldenke, Alph. st Invali pp £10," 2%, & i ats Moldenke, Phyto- 387, én, “u'629 (1918), 3° 873, 688, 536, , 166, 780, 781, 799, 874— 2 PHYTOLOGIA Vol. 11, no. 1 876, 909, 913, & ag (1949), and 4: 1010, 1084, 1100, ag & 130k. ig Moldenke, Known Geogr. Distrib. ’ Verben ac. , fede 2 Bol. Mus. Nac. Hist. Nat. Chile “253 70. 1921 Moldenke in Chit- tenden, Roy. Hort. Soc. Dict. gents 6: 2208, 2209, & 2212. 1951; Be J. Salisb., Ind. Kew. : 101. 1953; Lae oo - Cult. Montevid, 232. Sotky und Moldenke, Journ. Calif. Hort. Soc. 15 80. 195); ar in n Hunber, Fl. Madag. 17h: 8. “1986, Mat- a 3 122, 12 8, 125, 22h, 296; ° 357, ° 360, . 362, :371, s12, "3 4 30; & 736 1959; Moldenke, Résumé S Suppl. 1: 7 (1 959), 2: 12 &13. ag. Moldenke, Phytologia 8: 120, 121, & 123 (1961) and 8: 18h, 257, 280, & 316. 1962; Moldenke, Résumé Suppl. 3: 29 & ho" see) and’ 5: 8. 1962; A. Putz, N. Y. Herald a age h, p. 18, Feb- ruary l,. 1962; Moldenke, Résumé 6: 9 & 11 ae 63) and 7: - & oe 1963; Moldenke, Phytologia 8: 69 (1963), 9: 178 Rare 316, 329, 330, & 333—3% (1963), 9: 35h, 366-368, & 388 sens, and 10: 128, 129, 288, & 289. 19h. Illustrations: Paxt., Mag. Bot. "6s 23 [in color] - 2h. 1838; Floricult. Cab. 7: 22 [in color]. "1839; Hook. in Curtis, Bot. Mag. 65: pl. 369 [in color]. 1839; Maund, Bot. pia , pl. 197 {in oak "1839--18 Ls Je We Loud., Ladies Fl.-Gard. Ornam. Per- enn, 2: pl. 91 [inc lor}. 18h; Vilm., Alb. de Cliches, ed. 2, fig. 7c6Be. - 1888; sby, Hort. 1896: 188. 1896; L. H. Bailey, cel. Amer. Hort., ed. +a 6: 1912 & 1916. 1902; L. H. Bailey, Cycl. Hort. 6: BLS. 19173; Sanzin, Anal. Soc. Cientif. Argent. Buenos Aires 88: 112. 1919. or perennial, tufte » procumbent or creeping herb, usu- hed, stiff or sometimes almost suffrutescent at the base and then to 2 feet tall and branches ascending-erect or rooting at the nodes near the base, spreading-hirsute or firmly hispid, the stems subterete, the branches tetragonal; internodes abbreviated; pee decus sate- opposite, sessile or subsessile, rather rigia, reen, varying from ovate, lanceolate-oblong, or subovate-el laptic to rather - lo ab short~apiculate “at the apex, shortly cuneate and entire at the base and narrowed into the short petiole, sinuately serrate or ns mm. and 2——5 mm, apart, or irregularly and deeply inka phenie or even subpiamatitid dentate, more or less rugose-wrinkled and glandular-hirtellous or pilose-pubescent above, ert goer fot or hirsute-tomentose beneath and hispid on the venation, t venation usually more or less deeply impressed above, prominent 196), Moldenke, Monograph of Verbena 3 or is pammeete beneath, pinnate, anastomosing toward the margins; nara eet solitary or ternate, long-pedunculate, dense or very dense, sont considerably elongate even during anthesis with closely imbricate flowers, or at first in ri spikes, glandular-hirtellous; flowers erect, fragrant in and night, with the odor of jasmine (Jasminum) ; sss ye subulate- lanceolate to lanceolate-acute, half to one-third as long as the he commissural face pass i bisulcate, the 2n = of this handsome cma “was collected by Friedrich Sellow = Montevideo, MERE OTs one was eee, in the herb. ium of the Botanisches eres t Berlin, now destroyed. Macbride's photograph of it (no. 1752) “2 inscribed " sendy teucrioides var. platensis", Another Sellow collection, inscr inscribed "Brasilia" i type photo no, 34356), in the same herbarium was the type o scores Verbena mc oides was based on collec- Port saadtargeipd in the Andes Mountains. Briquet's V. platensis var. latiuscula was based on Regnell I:A.3649 and A.3905 from Paragu- ari, Paraguay, collected in eat and deposited in the Deles- sert Herbarium at the Conservatoire et Jardin Botaniques at Gen- eva, the former said to be "spe odbiden densius pilosa". Briquet says "Specimina dmani ad hanc cam varietatem ent. Planta 30--50 cm. alta, validor. Folia subsessilia, — vel latiuscule Sor tare ARE superficie ad 3x 1,3 cm., + iter crenata, cren culminibus 1--1,5 mm. altis et Si mm. distantibus, nunc pores ie 2 Spicae longius pedun ae I re this as typic latensis Spreng. hamaedryfolia f. Sodtaans tae ed on Hassler 428, 4 PHYTOLOGIA Vol. 1, no. 1 collected on a campo near the Rfo Capibary, Paraguay, in Septem- ber, and deposited in the Delessert Herbarium at Geneva. Of it Chodat says "forma foliosae i.e. foliis latioribus, tenuibus. Her- ba 0,3-—-0,5, petala violacea." Concerning V. platensis var. lat- iuscula Briquet says "Plants 30--50 cm. alta, validior. Folia subsessilia, ovata vel latiuscule Gvateunellipttca, nice ad 3 x 1,3 am., regulariter crenata, crenarum culminibus 1--1,5 mm altis’ et 2-=5 a distantibus. Spicae longius TES Herba 0,3——0,5 m. » petala violacea, in campo prope fl, Capibary, Sept., uasalenl” ne 428; suffrutex 0,5--1 m, altus, petala cyan- ea, in valle fluminis Y-aca, in nie ss Dec., n. 662h; suffrutex 0,3--0,6 m. altus, petala r » prope Tobaty in rupes- tribus collium, Sept., no. 6357; eutirates 0,3--0,6 m. altus, pet- ala coerulea, in valle fluminis Y-aca, in campo prope Valenzuela, Jan., ne 6936. Nous rattachons encore au V. platensis var. lati- uscula un échantillon unique: 'herba 0,25 cm. alta, SoroLia viola- cea, in culmine coll. Cerritom Jan. n. 102'. Cet échantilion Pp. ment cette forme de 1'étudier sur des matériaux =e abondants." Verbena platensis has been found in sandy rocky dry sunny soil aiong mg roadsides, roc! rocky fields, hillsides and rocky hills, rocky moun rocky slopes and rocky places in general, oot grassy Slopes, meadows, campos, sandy soil by streams, in good soils and Rosengurtt describes it as frequent in grassy clearings, and Hau- mann calls it "common" and "abundant"! in Buenos weenie On the other hand, Legrand calls it "rare" in Uruguay and Schulz = Sg 1% te Mrare® in Salta, while Osten avers that it < only found fern-covered ground. Common names recorded for the species include "flor de vovia", "germanderartige Verbene", "germander-leaved verbena", ngermander like vervain", frat ype a blanca", "margarita colorado", an a ita punza", "piedra", "verbena", verbena blanca", tyerve odorante", and weeresisie teucrioide", while STANDARDIZED PLANT _ “recommends the name "fragrant verbena". species seems to have been introduced into cultivation in 1837; a 181 it was in cultivation in Switzerland inasmuch as it is said to have been one of the parents of V. anais Hort. in that year. Troncoso (1937) reports it as cultivated as an orna- mental plant in gardens and parks in Buenos Aires. Bailey G9) al iy by Vilm “ 5922 by Haage & Schmidt. Mattoon (1958) says "V. eudineial past pga by Hirst & my London. In cultivation the plant is o flower in and autumn, It is recorded as adventive in in Pugiand by Douglas i. Kent in a letter to me dated November 28, 1949 -- cfr. also Rep. Bot. Exch. Club Brit. Isles 1927: 452 ( (1928) | where a Melville & Smith collection from Dagenhem, Essex, 196) Moldenke, Monograph of Verbena 5 Lemaire (182) gives ”~ following interesting description: "La Verbena teucrioides découverte pas le docteur Gillie es sur les parties les plus 61 oo des monts ers, au Chili, a plus de 3,250 matres au dessus du niveau de la m Elle fut trouvée plus tard croissant aussi sur les er asl el environs de Mon- tevideo, ainsi que sur le Pain-de-Sucre, dans la province de verte de toutes parts de poils alain souvent glandul ;’ munie de feuilles lancéolées-oblongues, profondément taahabea: sessiles; a nervures enfoncées profondément sur la face supbrioure, sail- lantes inférieurement; 4 fleurs feubrsinke Ss, os pis allongés, serrés, et § calyces allongés, se tordant aa lt an- thése. Ces fleurs, d'un blanc jaun&tre passant au blanc rosé, exhalent une odeur trés suave rappelant assez l'aréma de Jasmin, et dont l'tintensité est surtout remarquable de soir et pendant la nuit ° Rttapler (1873) describes it: "Brasilien. -—- Einj&hrig, im ewtic Me ch, die » all ungleich eingeschnitten-gez#hnt. Blumen sehr wo wohlriechend, weisz ichten und doldenartigen, dann verltngerten shar ga 2 jede Blume hat ein pfriemliches, behaartes Deckblatt, das zwei Mal kttrzer, als der Kelch, welcher seinerseits wieder viel bere ist, als die Rthre der Blumenkrone. Letztere ist grosz, auszen glatt, mit fast re- Abtnderungen erfahren betrachtet sie und die folgende als den Ausgangspunkt fast aller jetzt in den Garten kultivierten Varie " ita paquiriza ze tallos radicantes perennes ie florece en primavera, Avi daghsdose hasta el vesuting las rn son fragrantes Ps “sername Vive en campos virgenes, pobres y astosos, dregos Briquet (190) says vanes avait cité dubitativement le Vv. platensis 4 propos du V, teucrioides Gill. et Hook, M. 0. Kuntze a la premier établi la synonymie des V. platensis et V. erica des par l'examen des originaux de Sprengel. De notre Bag! possédons 4 1'Herbier copies la plante de Sellow s mere Sprengel @ basé son V. platensis et qui a plus tard até preor es a nouveau par daiane . sous le nom de V. scordioides, La synonymie ci-dessus nous paraft done maintenant définitivement paca Tor Le V. platensis ayant souvent &t4é confondu avec d'autres espéc du group, nous croyons utile d'en donner ici une eer com= pléte." Whereupon he gives a detailed description, in however, he makes the amazing statement that "Corolla eaunaian vel 6 PHYTOLOGIA Vol, 11, no. 1 olacea" — apparently indicating that the only specimen or spec- imens before him which gave any indication of corolla color on their labels were of f., violacea Moldenke. sais u nervures longuement poilues-ciliées e Il se sépare aussi sans peine V. chamaedrifo = Juss. (in oe Mus. VII, 73, ann. 1807); Briq. Verb. Balans. ar, p. 2. = V. peruviana 0. Kuntze Rev. III, 2. 2573 non etlice = Erinus peruvianus L. Sp. ed. 1, p. 630, ann. 1753). Ce dernier type s'en écarte par son ouiine notablenent e dépassant pa: r On a dtailleirs trés rant yen ee de V. chamaedrifolia avec des espéces voisines. C'est ainsi que le no: Sl de M. tan attribué au V. peruviana et provenant de l'Assomption, n'a tient nullement 4 cette espace, mais bien au V. eae eek var. truncatula, = Verbena peruvian est - ee une plante de l'Uruguay et de l'Argentine, figurée p 11ée (Journ. observ. ve etc. ot 25, fig. 5 ann, 1 728), at manque 4 la flore du ar of In my “opinion, the Morong 51 mentioned above is typical V. if at Montevideo in January, 1931, says: "Ein weiteren grober a thum Briquet's betrifft v. teucrioides Gill, fur den zweifelhaften Namen Sprengels 'platensis' nimmt, dem er ae lette Blttten zuschreibt. V. teucrioides rane fis Paraguay auscheinend nicht vor und die von Briquet's pla s Spr. ge- nannte Pfl. is V. marrubioides Cham. Hassler te mir sein Einversttndniss d: damit.....¥. teucrioides Gill. hat wohlriechende ig Uruguay, Sierra Pampeanas (Stiden v. Bras.) gesammelt worden, ebenso habe ich sie aus den (alten) centralen Gebirge argentiniens iste Cordoba) wahrend sie in die jtngeren Cordilleren noch nicht eingedrungen zu sein scheint. Es ieheins mir ein Fehler zu sein, alte, kannte sich Langem gebrauchte Namen wir teucrioides iia vielleicht witere aber nicht einmal sichere Namen zu vertauschen. Cf. Walpers Repert. IV p. 33 sub no. 90, ‘platens: msis: srasiee haec inter types of both names were preserved in Berlin and Geneva compared there by Kuntze and by Briquet, so there seams to be no doubt now about their conspec ity. Walpers (1845) kept V. scordioides Cham. separate from V. teu- crioides Gill. & Hook. and classified both in his Section Verben- 196, Moldenke, Monograph of Verbena 7 aca, Subsection Inermes, Group Foliosae, Subgroup Macranthae, and Secondary Subgroup Melindres along with 9 other species. Standley (1938) applied the name V. teucrioides to the plants now more properly termed xV. hybrida Voss, saying "Verbena. Cultivated in gardens as an ornamental plant. Native of southern ommon gard with flowers v: rom white pink, or purple, is a favorite ga of Central aque. especially in the rate regions, The pro- per specific of the garden plants, which may be in part of hybrid rere rf somewhat uncertain." Similarly, the work re- ported on by Barton (1939) as having been done with V. platensis was actually done with xV. hybrida. It should be noted here that the V. anafs Hort., V. teucrioides var. anais Lem., V. teucrioides var, al anais Lem., and V. teucri- oides anai anais Lem. regarded by me in previous publications as syno- nyms of Ve | V. platensis, are actually to be placed in the synonymy of xV. hybrida Voss since Lemaire definitely states that they are based on ee hybridization. The names V, teucrioides auri a Regel, V. teucrioides var. auriculiflora ra Regel, V. tancrlatdas ipieide Vilm., and V. teucrioides x venosa Hort. also refer to xV. , hybrida, while V. teucrioides x chamaedryfolia Osten is xV. osteni Moldenke. The only definitely known and named nat- ural hybrid involving V. platensis is with V. peruviana (L.) Brit and is xV. osteni . The Herter 750b specimen in the United States National Herbarium at Washington has flowers that appear to be rather small for V. platensis -- possibly xV. osteni is in- volved here, The Vilmorin illustration cited above does not look exactly right for V. platensis -- it may represent xV. hybrida stead, I suspect that most, if not all, of the records of angry or slightly rosy corollas represent flowers that are fading due to @. fs Specimens of V, platensis have been misidentified and distrib- uted in herbaria under the names V. chamaedryfolia Juss., V. coc- cinea Hort., V. odorata Meyen, and | V. peruviana (L.) Britton. On the other hand, the Calderén 646 and Herb. Hort. Bot. Basil. s.n. [Aug. 181] distributed as V. piatensis are actually - rida Voss; Archer 666 and Jtrgensen nm 3772 are V. incisa thek-4 and Hassler 1105 is V. kuntzeana Moldenke. Schauer er (1847) | cites a ites a Sellow s.n, from "in campis Brasiliae meridionalis" and an Arséne Isabelle s.n. from Montevideo; Augus- to (1946) also cites Isabelle s.n. from Montevideo, while Tronco- so (1937) cites her no. 306 from Buenos Aires. Previous records of this species from Paraguay by me in various of my publications were basen 2 on Hassler 11052, aden: is now regarded by be as being Ve kuntzeana Moldenke. In all, 116 herbarium specimens and 16 mounted illustrations, 8 PRITOLOGIA Vol. 11, now 1 including type or phototype material of most of the names invol- ved, have been examined ° Citatdons: CUBA: Havana: Van Hermann 876 (N). BRAZIL: State undetermined: Sellow s.n. (Brasilia; Macbride photos 3356] (Bm, Br, F-~photo, Kr-~photo, | N--photo, N--photo, Si--photo, Z-~photo). URUGUAY : Je Anderson 131 (Bn); Arechavaleta 25 (Ug); Berro 72 (N), 5521 (N), “679 (N); Cabrera 3852 (N); Castellanos s.n. [Nov. 2 ‘ 196; Herb, Inst. Miguel Billo 15776] (N, N)}; Collector undesig~ nated s.n. ee March 10, 1885] (Ug), sen. (Ug); = i cocatamanaennmemmmanl iene Campal, Prcaigetaseests < hgeeet tell Andis Herter 750 (Herb. Herter es (B), 750a (Herb. Herter seate Herb. Osten 22635] (Ca~l07186, N, S, Ug, W—2l4hh23), 750b (Herb. Herter 85476] (Ca—l53205, N, "Ww—1934633), 750¢ THerb. Herter 91079] (N), 750d [Herb. tapes 50367] (B), sen. [Estancia Santa Clara, Sept. ae 1926; Herb. Osten 19119} (S, Ug), son. {Santa Clara, Oct. 2, 1926; Herb. Osten 19188] (Ug); Legrand (Ug), 1567 (Ue), 3066 (Ug), 4508 (N, Ug); Montoro Guarch rch 579 ®), 2903 ms N); Née a (Q), 93 (Q); Osten 2876 (Ug, Ug), 3177, in in part (Ug), in part (F--photo, N--photo, Sg--photo, Ug, Z—~photo), soi om te), 20080 (Ug, Ug); Romachako 566 (Ug—981h); Rosen- gurtt B.810 (Ca--79856, N), B.2133 (N), B.2082 (i). B B.3010 (N), B.l577 (N)} Schrtder s.n. [Piedras de Afilar; Herb. Osten 19455] (Ug); Sellow s.n, ; SeN. (Montevideo; Macbride photos 17452] (Kr-—photo of type, N-— N--photo of type, N--photo of type); Teisseire s.n. [Es- tac. Moreno, 1913] (Ug--l76). ARGENTINA: Buenos Aires: Boffa 18 (N, N, S)3 Bruch 4091 (N); Cabrera 5166 (N), 6570 (N); Dusén 6261 (S)3 Eye erdan, Be Beetle, & Grondona ndona 23419 (Ca~—627L92) 5 Hauman - SNe [Sierra de “Tanai, rae -1905) feet sen, [Sierra de la . Ventana, a, I. cerning sateen (N); Reutzell 1101 (ca—-3312); Ruiz Huidobro 1175 (S), 1261 (S), 1575 (S). Catamarca: Jtrgensen 1297 THerb. Osten 11350) (Ug, # 1121006). Cérdoba: Hieronymus s.n. [10.X1.1881] (N); A. T. Hun- ziker 7033 (N), 7205 (N); Eossen 117 (Ba, Um—~h6); A. G. Schulz 5991 (Sz); Sparre 1376 (S); E. Wall s.n. (La Falda, 15/12/L6] (Ew); Wall & Sparre s.n. {La Falda, 15/12/16] (Ew). Corrientes: Bonpland 109 (N, N, W—2343h95, W—2343197), s.n. [Corrientes] (N). Salta: Parodi 13h67 (N); Schreiter 6620 [Herb. Inst. Migu- el Lillo 32820] (N, N); A. G. Schulz 2692 (N). San Luis: Varela 651 (Ca). Tucum4n: Monetti 1943 [Herb. Inst. Miguel Lillo 31338) (Vi). State undetermined: 1 Friedman sen. [1923] (Be)5 I Kurtz 6780 {Guasapampa] (N). CULTIVATED: Belgium: | M. Martens s.n, [h. b. 196), Moldenke, Monograph of Verbena 9 lov. 1839] (Br), sen. (Hort. Cels.] (Br). Ecuador: Fimnin (N, W--1420001), El Salvador: P. C. Standley 23652 Tis). Germany: Herb. Prager 18610 (Gg——3159). Massachusetts: L. He Bailey s.n. [Feb. 190] (Ba). New York: T. H. Everett s.n. [N. Y. Bot. Gard. Cult. Pl. 97/46] (N); H. Ne Moldenke 18652 (N). Uru- guay: Moldenke & Moldenke 19939 (N). MOUNTED ILLUSTRATIONS: Paxt., Mag. Bot. 5: 23. res i» 24h. 1838 (N); Vilm., Alb. de Cliches, ed, rh fig. 25685 ( VERBENA PLATENSIS var, STENOD ries io oe Chod, & Hassler, Bull. Herb. Boiss., sér. 2 sgn 1056. 1 Syno : Verbena chamae aedryfolia ra pie Chod., Bull. Herb, Boiss., sér. 2, 2: 610. 1902. Bibliography: Ch hod., Bull. Herb. Mar sér. 2, 2: ~ eee Chod., Plant. Hassler. ry I97 < + an Chod. & H Bull. Herb. Boiss., sér. 2, h: 10ss*-1056. 190h3 Briq. i poh & Hassler, Plant. Hassler. 10: L477—l,78. 190); Moldenke, Sup List Invalid Names 8, 191; Moldenke, Alph. List Invalid Names and 4: 1090, 1091, 1093, 2. 19493 Moldenke, Distrib. Verbenac., [ed. 2], 106 & 199. sity Moldenke, gia 3: 290 & 378. "19503 Moldenke, Résumé ork & 73. 19595, Moldenke, Résumé Suppl. 6: 7 (1963) and i os 1963. riety differs from the typical form pe "the species in having much narrower leaf=blades, oblanceolate-cuneate, subcoria- ceous, rugose on both surfaces, the venation more robust beneath, fragrant flowers, the corollas white or blue to yellow, the tube pale-pink on the outside. The type of this variety was collected by Emil Hassler (no. 5758) on the campo near Caraguatay, Paraguay, in recstaeinel de and is deposited in the Delessert Herbarium at the Conservatoire et Jar- din Botaniques in Geneva. Chodat says of it "foliis oblanceolato- cuneata subtus robustius nervosus, subtus et supra rugosis subcor- iaceis, caulibus subsimplicibus. Suffrutex 0,3—-0,), petala coe- t 1--1,5 mm. altis et 2—3 m. distantibus. Spicae (an constanter? ) brevins pedunculatae. Suffrutex 0,3--0,l m. alta, petala coerulea The variety i been found by collectors on hillsides, among praia on mountain slopes, in dry river valleys and among roc pen rs, Augus ary. The Herb. Osten 11350 specimen cited below bears a note to the effect that "the habit of the mountain plant is different from that of the plains, namely the leaves are narrower ~~ probably 2 10 PEYTOLOGIA Vol. LL, no. 1 vars. or subvars." Material of this variety has been misidentified ~ Pavol pote in herbaria under the names V, platensis Spreng., V. teucrioides Gill., and V, teucrioides Bog and as "V, Stet var. fol, ear agent a abe ee ae In » 4 he ia bie tls and 2 mounted photographs have been examine d by Citations: ener si Buenos Aires: Mansel s.n. [Bahia Blanca, 188} (Bm); Sparre 370 (S). Catamarca: JOrgensen 1297 (Ca--205333} s.n. [Andalgal4, Dec. 1916; Herb. Fac. Cienc. Med, Buenos Aires 1297] (N--photo, Sp--2578), Z-~photo), s.n. [El Suncho, Dec. 26, 1916; Herb. Osten 11350] (S); Rodriguez Vaq Vaquero 899 899 (N). Cérdoba: Castellanos s.n, [Valle de los Reartes, 15.X11.1920; Herb. Osten 15288] (Ug); Fe A. Roig 1046 [Herb. Ruiz Leal 17828] (Sm). Cor- rientes: Bonpland s.n. carters (W~-1705992). Jujuy: Burkart & Troncoso 11003 (N a La Rioja: Castellanos s.n,. {Herb. Mus. Ar~ gent. Cienc. Nat. 33891] (N). Salta: R. B. Fries 615 (S); Garo- & Romero sen. (La Silleta, 16-I-19h7] (Gg--353218, N); Ts Meyer 3451 [Herb. Inst, Miguel Lillo 35567] (En, N), 3456 [Herb. Inst. Miguel Lillo 35572] (En, N); Moldenke & Moldenke 19736 (N); O'Donell 4933 (N); Ruiz Leal 14186 6 (Rl); Schreiter 5079 [Her [Herb. Osten 20020] (Ug, W--1591L55), 6620 {Herb. Inst. Miguel Lillo 32820] (Ug—-h946), 9476 [Herb. Osten 22901] (Ug); A. G. Schulz 6581 (Z); Sparre 1201 (S); Venturi 5143 (S, W~1591h63), 7976 (Herb. Osten 2080h] (Du--372)03, Gg—166628, Ug, W—1hh331h, W— 1591397); Wall & Sparre s.n. (mw. of Salta, 1/12/46] (Ew, Ew, Em, Ew, Ew). Tucumdn: Bailetti titi 167 (Herd, Inst, Miguel Lillo 3137] (N); 3 Lillo 2899 [Herb. Inst. Ki Miguel Lillo 31340; Herb. Osten 8459] (N, Ug); Monetti 1943 [Herb. Inst. Miguel Lillo 31338] (N); Venturi 2,08 (W—159125)). VERBENA PLATENSIS f. VIOLACEA Moldenke, Phytologia 3: 177. 19h9. Bibliography: Moldenke, Phytologia 3: 177. 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 100 & 199. 199; Molden- ke, Résumé 120 & 473. 1959. This form differs from the typical form of the species in haw ing its corollas always violet or rose-purple in color. The type of the form was collected by Cornelio Osten (no. 3195, in part) at Arroyo Grande, Tala, in the department of Soriano, , U= » on October 3, 189%, and is deposited in the Osten Her ium at the Museo de Historia Natural in Montevideo. The specimen was originally annotated by Osten as "V, teucrioides Gill. forma flor. violascentibus". It is mounted on a sheet with a branch of the typical white-flowered form from the same locality. Osten re- ports that the form was growing along with "V. erinoides", but is not a hybrid with it. With this assertion I agree. er found the form on a stony campo at Santa Clara, at 200 meters altitude, and describes the flowers as rose-purple (xeno= 1964 Moldenke, Monograph of Verbena 11 chromia) and fragran all, 2 fatectas specimens, — the type, and 4 moun- ted photographs have been examined Ie) Citations: URUGUAY: Herter s.n. Herb. Osten 19188] (Ug); oe ten 3195, in part (F-~photo of type, N--photo of type, Sg--phot of. type, Ug--type, Z-~-photo of type). VERBENA PLICATA Greene, Pittonia 5: 135. 1903 Synonymy: Verbena plivata Greene ex Moldenke, Alph. List Cit. 2: 630, sphalm, 1948, Verbena cinerea H.B.K. ex Moldenke, Résumé 362, in syn. 1959. Bibliography: Greene, Pittonia 5: nak 1903; Prain, Ind. Kew. Suppl. 3: 187. 1908; Perry, Ann. Mo. Bot. Gard. 20: 248, 261, 07-1110, 1122, 1137--1139, 1150, 1151, 1163, 117k, 1183, 11877 1212, 1213, 12207 1228, 1230, 1236, 1239, & 129k. 1949; Moldenke, Known G eogr. Distrib. V Verbenac., fea. 2], 2h--26, 33, & 199. 19193 Moldenke, Phytologia 3: 132. 1919; Waterfall Rhodora 52: 20 & 38. 19503; Shinners, Spring Fl. Dallas 326. 1958; Moldenke, Résumé 26, 29, 31, 32, 39, 362, & 473. 19593; Moldenke, Résumé Suppl. 2: 3, hb, &12. ge Jones, Rowell, & Johnston Flow. Pl. & Ferns Tex. Coast. Bend 1h. 1961; Irwin, Roatinaiie Flow, Tex. 190. 1961; Lewis & pep Am. Journ. Bot. Ja 641, fig. 20. 19613 Moldenke, Phytologia 8: 12h (1961) and 8: 279. 19624 Moldenke, Résumé Suppl. 3: 7 & 10 (1962), 5: 4 (1962), 6: 3 4 (1963), and Al 36 ie Moldenke, Phytologia 8: 471, 473, & 7? (1963)" an : 3h, 36, 156, 165, & 199. 19635 Woldenke, Résumé mary 8: 1 aoc ) and 10: 2. 196); ldenke in Shreve & Wiggins, bi 9 tk Son. Des, 2: 12h]. 196) solace, Phytologia 10: 182 & 196), és ‘Tlustrat ions: Lewis & Oliver, Am. Journ. Bot, 8: 60, fig. 961 low, — or perennial, erect herb, branched at the b teLiows} branches Mes or ascending, to 2 dm. long, preading; leaves decussate-opposite; ee winged; lower Laat Nladee ob- long-spatulate or elliptic-ova -- cm. long, narrowed at the base into a margined petiole of ng he the same length, broadly obtuse at the apex, plicate, coarsely incised or incised-dentate, 12 PEYTOLOGIA Vol. LL, no. 1 often 3~lobed, more or less canescent, rugose and peer apetee sed-hirsute a above, hirsute beneath and prominently marked ( ticularly near the margins) with whitish veins, the ncn r ones similar but smaller, often appearing spatulates spikes terminal rdinari ompa ng 1.5-— ° e, attenuate from the base to the apex, acute at the apex, rsute, the midrib often noticeable; calyx 3.5--), mm, long, more or less “aig nao conan 3 its lobes very seg or subtruncate terminating ab rupt a@ vary bluish-lavender, light luis h-lavender, blui ar bluish~violet, or dark-blue to blue~purple, purplish-blue, purplish-violet, purplish, purple, lavender, pink-Lavender, or magenta, its tube scarcely longer than the c a "4-6 mm. de, the anterior lobe retuse; cocci ee eee » one « long, shallowl scrobiculate above, changing to indefinitely "striate toward the base, the commis sural faces finely muricate-scabrous to ee ly smooth, not reaching the tip of the coccus; chromosome number a Ts The type of this species was collected by Samuel Mills Tracy and Franklin Sumner Earle (no. 30) at Barstow, Ward County, Tex- as, on April 1), 1902, The specimen on which V. cinerea is based seems to be J. Reverchon s.n., collected on August 9, 1877, in Barnard Coll k an ems tain that this is an error in nee sow Pm for V, canescens H.B.K. In Texas V. plicata is said to occur chiefly in open sandy ground, flats, prairies, and ravines, and along roadsides, fre- quent on gravel hills, widespread in northern, central, and south- ern portions of the state from Childress and Y to Gonzalez igneous or * ary clay-loan veg re and or brickred-colored loose sand, coarse sand and sandy 1 andy red soil, red sand over gypsum, red Permian shale, Rass te alluviun, fine sandy loam brownish sandy~loam soil, orange-colored sandy loam or gh aid known b , pastures, oak areas, depressions, and wet ravines. It oc~ curs on cliffs and gypsum bluffs, ri shoulders, plains and brushy or high rocky plains, sand dunes, banks, rough, rocky, or dry rocky hillsides, yo ridges pia large yuccas, Prosopis- Condalia flats, open hillsides and flats, and on the sandy bottoms of floodplains. One is apt to encounter it among brush on sand- 196k Moldenke, Monograph of Verbena 13 stone ridges, along rivers and roadsides, and near irrigation ditches, at altitudes of 30 to 5400 fest, flowering and fruiting from February to October. Benson states that V. plicata ta is a component of the Desert Grassland vegetation type in Arizona. Waterfall reports it from "gravelly soil in Larrea-Prosopis—Gutierrezia association" in Reeves County, Texas, and in "silty soil in valley between gypsum ridges" in Oklahoma. Cory reports it as "frequent on roadsides" in Ward County. Brown calls it a "rare perennial in re in Brewster County, while Hanson avers that it is "frequent rocky Slopes" in that county. Scudday asserts that it is found aabeonie. out" Andrews County in low places, Pammel says that it is "common According to Runyon, it is "abundant in the Beeville region" of Bee seme Be Texas, and "frequent in Hidalgo and Starr Counties on the gravel hills"; he also states "flowers in terminal racemes, rare in the region near Brownsville, prefers sandy soil". A ® * 2 se | c alk iter — TT ° me ee U a Po 7 [ . uscrry Pan . Q — tm = oe a \ Pomotas PHLLips Se_ ° om valley ° Gms © 5 cues ms py ae "etm ’ sehom one Scam se => Crue | wae FONVILLE . porn a. 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WILKINSON n sn ROSA ORAL OOSAY « ee wants \ 0: iles SCHLEICHER meal ANGELINA TEWTON® e * LOE soe ‘oo wa ] SAN SABA ‘waDISON TTY // PO ee SPER RAPIOES Mid pana Shana WASHINGTON gp Tome ®, i 0 100 200 u : wine iy — : ream. on (st WALKER ‘Beaumecano | MLEN yarn es | — ne WILLIAMSON isa , OS be Ce am pe ae . sawn ay wan SI TANG vias oar went San . PAHOA! u TETTE PTY H —_ ter SE apps) HAROIN Tatas rrenson ) MAI — crt] “2 - rt) MAP NO. 125 UBERTY Lcatad st 1BERVILLE, i na a d = vee was > 80-6 Oo pont h. c A . ma WAYS st & wre rt OPV RIGHT sie ey vermiLion —Y BERA FY Sel [bane st co ~ ‘OMAL JEFFERSON } BERNARD, A G : I 6. Fa a | AMERICAN MAP COMPANY, INC. st. WARY ourcit} 4 Une aac cmon "sa ORIGINATOR } — ure Feamanes [ome ‘ait ba > Feacnont . eum ORS SOLE PUBLISHERS é eave an error, Argen says "Semina e Bonaria fuisse allata ferunt; ex aftinitate typi tamen regnum Mexicanum patriam autumaverim, Mire similis V. cil- 20 PHYTOLOGIA Vol. 11, no. 1 jiatae, vero tamen distincta pube, fol. laciniis magis elongatis, corolla et antheris super. omnis vestigio appendicis carentibus." In Chittenden's "Dictionary" this species is said to be "very similar to V. diffusa, but differing in having the calyx=-pubes- cence substrigose, not wide-spreading." Verbena diffusa, however, is a synonym of V. urticifolia L., with which our plant has no close resemblance whatever. Probably V, dissecta Willd. is in- Walpers (185) classifies V. pogostoma in his Section Verben- aca, Subsection Inermes, Group Foliosae, Subgroup Macranthae, and Secondary Subgroup Aubletia, along with ten other ie ae Ui ac was introduced into cultivation in or before 1839. A canm for it is "Argentine verbena", It has been collected in aathesis in August and from October to Agee ya In all, 4 herbarium specimen: d 6 mounted photographs, in- cluding photographs of the pai Se both names involved, have been examined by me wing ARGENTINA: La Rioja: Castellanos s.n. [Herb. Mus. Argent. Cienc. Nat. 33885] (N). San Juan: Cuezzo 2255 (N). Santa Fé: oop Balegno 82) (N). Tucum4n: Varela s.n. [22.1V.19b4] (Ca-- 165879). CULTIVATED: aarp! Herb. Hort. Bot. Berol. s.n. [Oct. 1839; Macbride photos 170, in part] iy ie ees of cotype, N— photo of cotype, N--photo of cotype), sen. [Aug. 183; Macbride photos 17,0, in part] (Kr--photo of cotype, Bpticto of cotype, N--photo of Cot: ype). VERBENA PORRIGENS R. A. Phil., Linnaea 333 195.1 Synonymy: Verbena porrigenes R. A. Phil. ex uolaene, Known Geogr. Distrib. Verbenac., [ed. 1], 42, sphalm. 192. Bibliography: R. A. Phil., Linnaea a 33: 195. 1864; F. Phil., Kew. 2: 1179. 1895; Moldenke, Known Geogr. Distrib. Verbena {ed. 1], 42 & 102. 19423 Moldenke, Lilloa 10: 345. 19hh; qcinscks, Alph. List Invalid Names Suppl. 1: 26. 1947; Moldenke, Known Ge- ogr, Distrib. Verbenac., [ed. 2], 102 & 199. 199; Moldenke, Alph. List Cit. 3: 688, 813, "e 940. 1949; Acevedo de Makes os ‘Mus. Nac. Hist. Nat. Chile 25: 59. 1951; Moldenke, Résumé 12 vo 473. 1959; Muffoz Pizarro, Espec. Plant. Deser. Philippi “he 19603 Moldenke, Résumé Suppl. lj: 5. vot Moldenke, Phytologia 8: 192 & 204 (1962) and 10: 298 & be Suffrutescent, foal ee As Ree with short white spread- ing hairs throughout ; stems several, rising from a woody base, e- rect or asce » 15--)5 cm. long; "leaves decussate-opposite, the larger ones about 1.6 cm. long and almost as wide, multi ite, strigose, with 2 or 3 lobes on each side, the lower ones 2- or 3-lobed, the rest entire, all the lobes linear, the appressed white hairs; peduncles 1.l\--5 cm. long; inflorescence capitate, later elongating; bractlets lanceolate, about 3.5 mm. 196 Moldenke, Monograph of Verbena 21 long or half as long as the calyx; calyx about 5 mm. long, hispid; iol rose, its tube 8 mm. long; anther-appendages clavate, ex- serted. The type of this little-known species was collected — Christ— vatis, exsertis munita, quae antea innotuit, est V. sulphurea Sweet, quae colore floris, colore viridi foliorum, eorumque lac- iniis’ longioribus, angustioribus primo adspectu ditfert. * The Skottsberg & Skottsberg 843, sometimes regarded as representing V. porrigens, is actually V V. sulphurea D. Don. Acevedo de Vargas (1951) cites the following specimens not as yet seen by me: CHILE: Aconcagua: Landbeck s.n. [near Quili- mari, XII .1861] (Sg--54729--type). Coquimbo: Reiche s.n. [Los a, 1.1899] (Sg—68380), sen. [IX.1905] (Sg-~54730). Valpar- afso: Collector undesignated s.n sen. [Quillota] (Sg—h2l78). In » one herbarium specimen n and mounted photographs, in- cluding be of the types of both names involved, have een examin agar CHILE: Aconcagua: Landbeck s.n, [near Quilimari + 1861; Herb. Mus. Nac. Hist. Nat. Chile 54729; Macbride of isotype, N--photo of isotype). Coquimbo: Herb. Mus. Santiago 1k (N). XVEFRBENA PROSTIBULA Moldenke, Phytologia sid : Rone 1955-6 Synonymy: Verbena hispida x simplex De » Cytologia 7: 170. 1936. Verbena hispi hispida la Ruiz & Pav. x av. x V. 8 pri Lehm, ex Moldenke, Résumé 366, in syn. 1959. Verbena simplex ae: x V. hispida Ruiz & Pav. ex Moldenke, Résumé 37 7u, in sy. in syn. 1959. Bibliography: Dermen, Cytologia 7: 170. 1936; Moldenke, Phyto- lo 19 ol. Abstr. 30: 1093. 1956; Moldenke, Am. Midl. Nat. 59: 359. 1958; uoldenke, Ré- sumé 22), 366, 37h, & 473. 1959; Moldenke, Phytologia 8: (1961) and = as 1963. his is the hybrid between V. pispids Ruts & Pave one 2 plex Lehn. pect oe artificially by Dermen in Massachusetts and described * him in 1936. Since these two species do not grow in the wild together anywhere, as far as I am aware, there is little oe of ever finding this hybrid in Nature. Both species, how- occur in cultivation -—- especially Europe — the hybrid might have horticultural possibilities. PULCHELLA Sweet, Brit. Flow. Gard., ser. 1, 3: pl. 295. 1829 [not V. pulchella Greene, 1903]. 22 PHYTOLOGIA Vol. 11, no. 1 Synonymy: Shuttleworthia pulchella Meisn., Pl. Vasc. Gen. 198. 1839. Shuttelworthia pulchella Meisn. ex Steud., Nom. Bot., ed. 2, 2: 575. 1841. Verbena pulchella Hort. ex Steud., Nom. Bot., ed. 2, 2: 750, in syn. 1841. Schuttleworthia pulchella Meisn. ex Morren, Belg. Hort. 20: 111, in syn. 1870. Verbena pulchella Spreng. ex Regel, Gartenfl. 28: 372. 1879. Verbena laciniata var. albida Herter ex Moldenke, Suppl. List Invalid Names 9, ins 1941. Schuttelworthia pulchella Meisn. ex Moldenke, Alph. List Invalid oe fi aa 1: 19, in syn. 1947. Verbena pulcherrima Harvey e nke, Alph. List Invalid Names Suppl. 1: 26, in syn. 19h7 ict v, ‘paharvin “Vilm., 1865]. Bibliography: Swe , Brit. Flow. Gard. 3: pl. 295. 1829; Lodd., Bot. a 16: pl. rece 1829; Matind Bot. Gard. 3: pl. 70. oe 1830; Swee » Hort. Brit., aay? ° 1830; Reider, Ann. Walp., Repert. 4: 13 & 3h. 185; Payer, 115. i857; Bocq., Adansonia 2: 106, 107, . ‘ “130. we sa61 1862; Bocq., Rev. Verbenac. 2G, 27, Si, & 52. 1862; Vilm., Fl. Pl. 111—-112. 1870; Ulrich, Internat. Worterb. Pfl., ed. 1, 250. ey & - in Macaig Illustr. Blumeng., ed. 1, 1261--1262. 1873; Ulri Internat, Worterb. Pfl., ed. 2, 250. 1875; Vilm., Alb. de Cliches, ig 2, fle. 25680. 1888; Jacks » in Hook, f. & Jacks. es . Kew 895 & 1179. 1895; L. H. Bailey in A. Gray, Field Forest & Gard. 3316 (Ug), 3523 3523, in part (Ug), . 3923 (Ug), le), auer6 (Ug), 22006 (Ug, Ug) > 22096 (Ug); Rosengurtt B.992 (N), Be2116 1/2 (N); Safford s.n. [Independencia, oct. 2 Oct. 2h, 1886] (H-992284) 5 Schrtder 3523a (oe), Son. (Herb. Osten 16546, in part] (Ug); Seijo 0 650 (Ug), § Son. Nov, 188] (Ug); Teisseire s.n. (Ug). GORRITI I ~ TSLAND: — (Ug). CHILE: Antofagasta: Barros Valenzuela 8030 (N)- “ARGENTINA: 26 PHYTOLOGIA Vol. 11, now 1 Buenos Aires: Boffa 330 (Ug); Cabrera 10352 (W—1988265); Ryerdam & Beetle 23039 (Ca—-652),28) A. Te .. T. Hunziker er 539 (N, N, S); Kra-_ kra~- mee 3086 (N); Martinez Crovetto 32 0 32 [A. G. Schulz 1899] (Sz); 2678 (Ug), 2707 (Ug); Parodi Parodi 8563 (Herb. Osten 20938] (Ug, a “220 ist Rodrigo 2339 (N)3 Troncoso 172 (N); Wall & Sparre sets [3/11/46] (Ee, Ew, Ew, Ew). Mendoza: “Smitt sen. [1890--91] (8, 8) a FH del Estero: Frenguelli 105 105 (N), 137 (N). Tucu- ee Venturi 4295 (W--15914)2). CULTIVATED: France; Herb. Hort, Bote Huber Huber 753 (M (M). Italy: Herb. Harvey s.n. [h. P., 1848] (Du— 166,93). New York: Je H. Cowen son. n. [Cornell U., Ithaca, April 1900] (N). LOCALITY OF COLLECTION UNDETERMINED: Collector undes- ignated s.n. (Ug, Ug). MOUNTED ILLUSTRATIONS: Lodd., Bot. Cab. 16: pl. 1550. 1829 (N); Sweet, Brit. Flow. Gard. 3: pl. 295. 1829 (N); Vilm., Alb. de Cliches, ed, 2, fig. 25680 (N). ENA PULCHELLA f. LATILOBATA Moldenke, Phytologia 3: 281. 1950. Bibliography: Moldenke, Phytologia 3: 281 & 289. 1950; Molden- ke, Btaset 120 & 473. 1959. thi mm differs from the typical form of the species in hav- ing the beat gataeer’ very broad, 3--7 mm, wide, and obtuse at The type of the form was collected by Mariano B. Berro (no. 4751) at ye Santa Lucia, in the department of Montevideo, U: U- ° October 30, 1907, and is deposited in the Britton pacha ay con the New York B a Garden, I; is known thus the Stee tang? URUGUAY: Berro 2 bre (N-type). VERBENA PULCHRA Moldenke, Phytologia 2: 476--,77. 198. Smee a ldenke, Phytologia 2: 476--,77. 1948; Moldenke, Castanea = 4 - 1948; Moldenke, Known Geogr. Distrib. Verbenac., fed. 2], 107, & 199. 19495 Moldenke, Phytologia 3: 75 & 136 (1919) bar 3: 289. 1950; $s naman Trib. Farmac. 19 (10): 167. a E. J. Salisb., Ind. « Suppl. 11: 263. 1953; Angely, Fl. aran. 73 é & 13 (1957) ‘ea “ 17. 1958; Moldenke, Résum ie re & 473. 1959; angely, Fl. Paran. 16: 79 (1960) and 17: 3 cumbent or ascending, several, not branched, sidects ieccugtual. sulcate between the angles, ’ scabrous wi many short reflexed hairs; principal internodes 3-8 cm. long; nodes more or less annulate; leaves numerous, decussate-opposite, usually with clusters of smaller ones in their axils; petioles slender, 5~-3 mm. long, rather sparsely strigose with short an- ‘acuminate % the base, rather irregularly eee from the apex lane to a base with rather broadly triangular a teeth, slightly scabridous-strigillose above, rather softly plane 1964 Moldenke, Monograph of Verbena 27 Li yesige beneath with bulbous-based hairs; midrib very slender, plane above, prominulent beneath; veinlet reticulation mostly in- discernible above, conspicuous (but not elevated) bemeath; inflor- escence terminal and in the uppermost axils, the floriferous por- tion very dense, at first flattened-subcapitate, later elongating to 3 cm, but t remaining v ery dense; peduncles rather stout, 6--3.5 cm. long, tetragonal and sulcate like the stems, reflexed—pilose and more or less rough to the touch; bractlets lanceolate, l-~9 mn. long, 1--1.l, mm. wide, attemate-acute or subacuminate at the apex, glabrous (or very sparsely pilosulous) except for the long- ciliate margins; calyx tubular, 10-~12 mm. long, about 1 mn. wide, 5-costate, purplish especially toward the apex, ciliate~ pubescent on the ribs, the teeth unequal, 1--2 mm. long, filifom- appendaged; corolla showy, blue or Binewiotet to rose-violet or violet, its tube 15—-17 mm. long, very sparsely and minutely pub- erulent or merely pulverulent on the ubaite above the calyx, its limb to 15 mm. wide, pilose in the throat. The type of “re species was collected by Per Karl Hjalmar Dusén (no. 9334) in wet thickets at Calmon, Paran4, Brazil, on March 13, - 1910, and is deposited in the herbarium of the rin historiska Riksauseum at Stockholm. The calyxes on the t species inhabits campos and, oe to Arechavaleta, "tierras one herbosas", blooming ’ from anuary to April and in October and November. Material has been tetaemifiod and distributed in herbaria under the name "Verbena aff. phlogiphlora Cham." In all, 19 herbarium specimens, ee the type, and mounted photographs have been examin Citations: BRAZIL: Parand4: Braga & Langs 255 (Bm); Dusén 9334 N--isotype ) S--type, Si-- Photo of type, Z--photo of type); Hatschbach 4369 (Z). Rio Rambo 30980 (N). S#o Paulo: Brade 16268 [Herb. Jard. Bot. Rio de Jan. 42222] (N, N). URUGUAY: Arechavaleta 35 (N), sen. [Tac- uarembé, Nov. 1903] (Ug, Ug); Legrand 2561 (Ug); D. Wright son. [Cufapira, — (Bm). ARGENTINA: TNA: Misiones: Ekman n 1980 (N (N); Ge 3 G. Je Schwarz 2523 N). VERBENA PULCHRA var. PALUDICOLA Moldenke, a 23 woh 19h8. Synonymy: Verbena menthaefolia Herter ex Moldenke, Résumé Bos rg As 9, in syn. 1963 [not ve aniedass Benth., 1839]. graphy: Moldenke, Phytologia 2: Le? ( (1948) and 3: 75. 19195. Molnenke’ Known Geogr. Distrib. Ve: , [eds 2], 199. gh Moldenke, Résuné 120 & 73. 19595 Yoldenke, Résumé Suppl. - 1963. thie variety differs from the typical form of the species in its leaves being shorter, the blades only 2.l4—l.3 cm. , ae 1 2 cm. wide, and the corollas white except for a red " 28 PHYTOLOGIA Vol. 11, no, 1 The type of the variety was collected by Wilhelm Gustav Herter (no. 99937) in a rather wet sandy marsh, me an ape of 150 meters, exposed to the sunlight in eanqueras, Riv- era, Uruguay, between March 2 and 27, “1907, oe pa cage “see bk the Britton Herbarium at the New York Botanical Garden. The ta is known thus far only from the type specimen. Citations: URUGUAY: Rertar 99937 (N--type). VERBENA PUMILA Rydb. in Small, Fl. Southeast. U. S., ed. 1, 1010 & 1337. 1903. mymy: Verbena inconspicua ogriiny Pittonia 5: 137. 1903. Verbena bracteoso-aubletia Hall ex M oldenke, Suppl. List Invalid Names 6, in syn. 19h1. Verbena permila Rydb. ex Moldenke, Suppl. List Invalid Names 9, in syn. syn. 1941. Verbena pmila Rydb. ex G. L. Fisher, Am. Bot. Exchange List 196, sphalm, 1946. Verbena pum- ila Greene ex Moldenke, Résumé Suppl. 3: 40, in syn. 1962. Bp scary Eggert, Torreya 2: 12). 1902; J. K. Small, Fl Southeast. U. S., ed. 1 1010 & 1337. 1903; ci Pittonia Ss 137. 1903; Prain, Ind, Kew. Suppl. 3: 187. 1908; J. K. Small, Fl. Southeast. U. SG ed. 2, 1010. 1913; E. D. Schulz cand Wild Fis. 178. 1922; E. D. Schulz, Texas Wild Fls. 337. 1928; Perry, Mo. Bot. Gard. 20: acta A 312, 314, 338—339, & 356. 1933; cory, Texas Agr. » Bull. 550: 89. 1937; E. L. Little, Am. Mid]. Nat. 19: jose i930; “Woldenke, Prelin. Abn. List Invalid Names 45 & 6. 190; Sperry, Sul Ross State Te ach. Coll. Bull. 22: h1. 1941; Moldenke, Suppl. List Invalid Names 8 & 9. 19); Geogr D 2356 s 2: 398, foo, fo3. Lo 7, LS0—-23, 470° Vlcl79) 488, h95— eth T SAD 520, 522, 526, 539 sl, 55h, 559, a oy ats 19 678, 22, 1129 38, 1139, 1143, 1h9—~1151, 1162, 1264, 1166, 1172, 1182118, 1193, 1203, 1207, 1212, 1216, 1220, 1225, 1226, 1238, 1239, 1241, 196) Moldenke, Monograph of Verbena 29 = aes Pre & ie orf Moldenke, Known Geogr. Distrib. 2),.4802hy 255.33, £199 19495 Moldenke ak peer 3: 132. ‘ais, cat Sp allas 325. 1958; Moldenke & 22, 26, 29, 31, 39, ripe 367, 372, & 473. 1959; Jones, Rowell, & Jo » Flow. Pl. & Ferns Tex. 1961; Irwin, Roadside Flow. Tex. 189— 190, gh 39. 1961; Lewis & i . - 48: 639-641. 1961; Moldenke, Phytologia 8: 123 (1961) and 8: rips rapa 31, 279, 401, 435, & 437. 1962; Moldenke, Résumé suppl. 3: 2 657 & bo (1962), heh (1962), 5: (1962), 6: 3 & 10 (1963), : 2 & 3. 1963; Moldenke, Phy- tologia 9: 17, 18, 2h, PP 143, 195, 197, 199, & 202. 1963; Shinners, Sida 1: "298. 196k. oo Lewis “A Oliv., Am. Journ. Bot. 48: 60, fig. 2. green; branches” decumbent, to 15 cm. ae leaves decussate- opposite, deltoid to ovate, 1.5--3 om. long, often obtusely tri- lar with a cuneate or truncate base contracted into a short narrowly margined petiole, trifid, occasionally lobed, the div- isions variously FooEpets incised, or es jag sogcaneni long as the calyx, Mehichiterte: flowers very small, eg corolla small, hypocrateriform, varying from blue or ggg oral e purple, purple, or prt preamp with a minute yellow Neye", glands minute or absent; fruiting-calyx about 6 mm. long, pubes- finely glandular, its lobes short and subulate; fruit cylindric; cocci 2,25—3 um. long, deeply wrinkled or reticulate~scrobicu- late except at the base, the commissural faces muriculate; chro- mosome number: n = 10, The type of this common species was collected by Benjamin Franklin Bush (no. 1275) at Catoosa, Rogers County, Oklahoma, on May 8, 1895, and is deposited in the Columbia University Herbari- um at the New York Botanical Garden. Cotypes of V. inconspicua met collected by Samuel Mills Tracy and Franklin Sumner Earle — » 106, on the plains west of Pecos, Reeves County, on April 20, ote and no. 178, in the foothills of the Davis Mountains, Jeff Davis County, Texas, on April 23, 1902. The type of V. bracteoso- aubletia was collected by Elihu Hall (no. 431, in part) indry at or near Austin, cies County, Texas, in May, 1872. On 30 PHYTOLOGIA Vol. 1, no. 1 his labels Hall seens to credit the hybrid name to Asa Gray, say- ing "Verbena bracteoso-aubletia? so says Gray, but a distinct spe- cies" and "V vg "V. bracteoso-aubletia fide Gray, but a good fertile species It peeala be noted here that the V. pumila f, albiflora Standl. is actually a synonym of Vv. quadrangulata Heller. The "V. pumila” of Irwin (1961) is Vv. tenuisecta Briq.,as is plainly indicated the plate. Tolstead & Cory (196) refer to V. pumila as "an inconspicuous- ly flowered weedy plant.” Runyon describes the fruit as "a cylin- dric capsule". C.M. Rowell 4776, cited below, bears a notation on its label: "with pale purple to white corollas", and so may be a mixture with f. albida Moldenke. Verbena pumila has been found by collectors at the edge of woods, os roadsides, gravel or rocky clay hina haga railroad tracks and rights- seh arroyos, and the green e por of dried- up streams, growing in moist or dry soil, sandy, d andy, or rich sandy gett. black or shaded soil, rocky or posicarense soil, red or s clay soil, limestone soil, much overgrazed andy clay loam, sandy red sandy clay, ms or low open ground, low damp or moist ground, soon fields, old pele? and otitelvated fields, nae the edge of oak woods, an sandy fields, wet or open rocky P ods, open or wet woods, open oak stoak woods, alluvial woods, e forests and sandy pastures, in draws and creek-bot llanos and catch-basins, glades and > or open pastures, lowlands, limestone Aan se apron’, desert scrub, granite or cultivated areas, in dry draws and dry pastures, moist banks and roadside ditches, in creek valleys and river-bottoms, yards, flat sandy areas, floodplains, dry open pastures, low scrub Woodford chert formation, and sand formation of the Wil- cox group. It is described as a oheasee in abandoned fie has been collected also on mesquite e floors of can- yons, gravel and sand bars, small streams, cliff est on 1 s and mesas, xeric rocky limestone outcrops, the sides and tops of buttes, dry, » or granite slopes, va aren road shoulders, plains and high rocky plains,flats sum, or dry granite foothills, as well as on hills and sand hills, rocky sandstone, or gypsum hills, I4 is said to grow on prairies, sandy or See ee prairies, and on calcareous and mesquite prairies. in general, it is described as an inhabitant of fields, gee tovay and cha aparral, meadows, pastures, river-terraces, woods, and sandy slopes and hilisides, blooming from January to August, in the western, central, and southern parts of the state, 196, Moldenke, Monograph of Verbena 31 from Reeves, Jeff Davis, and Brewster to Cameron, east to Wharton and tp oon and north to Grayson and Childress Counties. Actual- oS Pee + has’ been collected in anthesis in every month of the year, fruiting from ru , August, November, and December, tudes of 10 to 2830 meters. Jones, Rowell, & Johnston tone state that it p e ome road— sides, mostly on sandy loam, flowering in spring. Clemens says ae it inhabits the Metores i life zone"; Hanson states that it i p aie a "frequent weed in fields"; Warnock found it "scattered on aad silt flats", while Runyon says "frequent in sandy soil" and ommon throughout Cameron County; in the sandy i pe this plant a freely at the nodes; it is frequent but not common in Ken- County", Texas. Bray maintains that it is "very common in dry ground in Travis County, where Warnock found it "frequent in andy banks" and Ely "in black soil containing 38 percent silicon Sart ie derived from basalt". In Archer County, Texas, Whitehouse found V. pumila to be "very abundant and forming masses of color in scattered mesquite" and in woods. owell calls it "occasional" in See Patricio County. Shinners (1958) de- scribes it bia "sandy, silty, or gravelly ground, frequent and locally co , Prairie Border westward. Late March--May". McCart found it vin Light tan loose soil of stabilized sand dunes", while Cory says that it is "frequent in waste ground around shallow ponds" in Hamilton County and "infrequent on roadsides" in Johnson Hamby notes "also in Medina, Zavala, Real, Dimmit, & Bandera Coun- ties", Williges found it "occasional in undisturbed Cees pid In Garza County it is described by Rowell as "occasio loam in flats below rim of canyon", while the same oma: in San Patricio County, notes eae frequent but not e num- bers in deep loose sand along and adjacent to river." In the same county Gould & Hycka found it in "grassland-shrub area along river bank." La il P xeric Soe with granite outcrops and juniper, Yucca, and cac- tu In CidLahona it is said by Eskew to be "common along streams", by Engleman "common in clay banks", and by Stevens "common in open Woods"; Waterfall found it "in shallow soil on limestone conglom- erate", and Robbins "in moist grassy depressions on open roc slopes in open oak-hickory woods", while Hopkins reports it from dry rocky pastures originally shortgrass prairie", "in and along ins ure: um mon (Diospyros) and hackberry (Celtis)". In Sonora it is said tap Gentry to be found in "arroyos and milpas, Lower Sonoran; scattered in the eerorse but more colonial in the milpas” and in the "Short- =e re vernacular names recorded for this species are wictenachar peimeng "hairy verbena", "low verbena", "pink ver- 32 PHYTOLOGIA Vol. 1, no. 1 bena", "pink vervain", "small-flowered verbena", and "wild verbena" Material of V. pumila has been misidentified ” and distributed in renee under the names V. ambrosiafolia Rydb., V. aubletia Jacq., V. bipinnatifida Michx., V. bipinnatifida Nutt., V. bract— eata var. brevibracteata Gray, V. bracteosa Michx., V. bracteosa var. brevibracteata Gray, V. canadensis (Le) Britton, — ¥. canescens HeBK., Ve ciliata Benth., V V. ciliata var. longidentata | Perry, Ve delticola Small, V. lambertii Sims, V. quadrangulata Heller, V. racemosa Eggert, v. wrightii A. Gray, Helleranthus quadrangulatus (Heller) Small, and even Glecoma hederacea L. in the Lamiaceae. The Carleton 13 collection bears a note reading "V. bipinnatifida Nutt., a dwarf form. This species and V. aubletia evidently run together." Hayes 610 was first identified as V. bipinnatifida, then as V. ciliata, and finally as V. pumila. ota On the other hand, the R. Runyon . 2359, 59, 2366, & 2h99, distribu- ted as V. pumila, are f. albida Moldenke; Shreve 7169 | is actually the type collection of V. bajacalifornica Moldenke; Reynaldo Ra- mirez 36 & Waterfall 13097 are V. bipinnatifida Nutt,;H. C. Hanson 30h is V. ciliata var. longidentata Perry; B. H. Warnock 11693 is Ve V. quadrangulata _ Heller; B. H. Warnock 46135 & - 4701 are Ve ra race- mosa Eggert; and G. J, Goodman 2396 & 2hie are V. wrightii A. Gray. The A. A. Heller 1365, previously cited by me as V. pumila, is actually V ve. ciliata Benth. , While Clover 567 is V. teucriifolia Mart. &Gal. The E. Hall 431. and M. Hawkins 37 are mixtures with V. bipinnatifida Nutt.; He CG. CG. Hanson 322 and and Ruth 110 are mix- tures with V. ciliata var, longidentata Perry; Ro Rose, e, Standley, & Russell 12745 is a mixture with Ve. gooddingii var. huoeti toile Tidestr.; Parry, Bigelow, Wright, « & Schott 830 is a mixture with V. racemosa Eggert; and Wencurtin s.n, [5-2-1916] is a mixture ently Asa Gray combined three of Palmer's field numbers under the one number "2),6": his original no. 21, collected on April 6 in a County, appears to be the V. v. caaaienia portion, and his riginal no. 167, collected on May 22 in the same county, is Ve Seats leaving his original no. 438, collected on June 2h in Garvin or possibly in Johnston Count y, as V. pumila. Edw. Palmer 247 is also a mixture with V. canadensis —- — here Gray combined two of Palmer's original field numbers under "2:7": his original no. 260, collected on June 11 in Garvin County, seems to have been vec anadensis, while his original no. 390 , collected on July 1 either in Johnston or Atoka County, s' seems to be V. pumila. printed labels on all these are inscribed "On the False Washita between Fort Cobb and Fort Arbuckle" 196, Moldenke, Monograph of Verbena 33 Lundell & Lundell 10226 was determined by me and distributed as V. quadrangulata Heller, but the United States National Herbar- ium sheet does not exhibit beaked cocci. The E - J. Palmer 13562 specimen in the same herbarium has cocci with Tetintte bea beaks and is therefore cited by me under V. quadrangulata. E. L. Little says of his no. 472: "Ve bracteosa. Possibly a hybrid with ve canadensis as 3 the b bracts are short", but to me the specimens | ap= pears to be typical V. pumila, The Lindheimer 1075 collection may be from either Comal or Comanche County -- its label is in- scribed "Comanche Spring: New Braunfels, etc." The printed labels on C. Wright 1500 are inscribed "Coll. N. Mex.", but Perry (1933) Hore ‘the collection from "southwest Texas, 1851-2". J. Frank and's surname is incorrectly written orman" on some labels. (E). Johnston Co.: H. W. Houghton 2570 (G). Murray Co.: Emig 4Ol (E); G. W. Stevens 28 (E, G); Stratton 11 (E). Rogers Co.: B. . F. Bush 1 1275 (E--isotype) . TEXAS: Bastrop Co.: E. J. Palmer 33367 (E). set Co.: Be F. Bush 1185 (E); Clemens & & Clemens 976, in part (E); Eggert s.n. . [San Antonio] (E); G. G. Jeray "209 09 (G), s. n. [Bexar Co.] (E)} Trelease 99 (E); Wilkinson 8 (E), “108 (E). Blanco Co.: E. J. Palmer 13281 (E). Cameron Co. He C. i Hanson 322 (G), son. .. (Brownsville, 23 Jan. 1919] (E). Comal C ae Lind- heimer 43 (E), 1075 (E, F, G)- Comanche Co.: Eggert s {Round Top Mountain, 9 May 1900] (E), sen. [near fasta 10 May 1900] (E). Dallas Co.: J. Reverchon 738 (E), 1963 (E, F, G), 2117 (E). Gillespie Co.: 3 Ge Jerny 18h ( (E). Haskell Co.: Morton .n. {Haskell, 1898] (E). “Hood Co.: Eggert s.n. [near Granbury, 6 May 1 900] (E). Howard Co.: Eggert s.n. [Big Spring, 11 June 1900] (E). Jeff Davis Co.: Tracy & Earle 178 (E, F, G). Kerr Co.: E. J. Palmer 9278 (E), 10003 (E). Reeves Co.: Trac racy & Earle 106 (E, F, G). San Saba Co.: E. J. Palmer 11843 (E). Tarrant Coe: Ruth 110, in part (Polytechnic, 10 April 41 1913) (E), 110, in part Trerrent Co., June 1913] (G). Travis Co.: E. Hall 428 (G), M3, in part (E); E. J. Palmer 9339 (E); M.S. Young s.n. [Austin] (E, G). Uvalde Co.: B. J. Palmer 13562 (E). Victoria Co.: Eg- ert s.n. [near Victoria, 11 April 1900] (E, G) can coe: Be J. Palmer pm E Wharton Co.: E. J. Palmer O (E), (8) oo Court tae aces Lindheimer III.501 (E, F, G); C. Wright G). NEW MEXICO: County undetermined: C. Wright 1a aan (D). MEXICO: Sinaloa: M. E. Jones 23243 (E). Son- ? Edw. Palmer 326 (G). She says nyerbena pumila has often been saategant with V. ve ee oe ata and, superficially, sitet resem- bles it, but is readily separated on the beakless matlete.s on "Reverchon 1963" which ee is cited by me below as "J. Re 3h P3ITTOLOGIA Vol. 11, no. 1 erchon s.n. [Curtiss 1963]" ~—- her "NY" specimen is actually in the Barnard College Herbarium, Her "NY" citation for C. Wright 1500 is based on a Torrey Herbarium specimen. She cites Lindheim- er 34 from Comal County, but I cite it below from Bexar County. She cites "V. brevibracteata Eggert, Torreya 2: 12h. 1902, not V. bracteosa var. brevibracteata Gray, Syn. Fl. N. Am. at: 3%. 1878" as a synonym of V. pumila. However, Eggert plainly bases his binomial on the trinomial of Gray and the latter belongs in the synonymy of V. bracteata Lag. & Rodr. The names, V. brevi- bracteata Eggert and V. brevibracteata (A. Gray) Secor’, there- fore, which in my previous publications have been reduced to V. pumila belong, instead, in the synonymy of V. bracteata. In all, 608 herbarium specimens, including the type collections of all the names involved, ers 3 mounted illustrations and clip- pings have been examined by m Citations: ARKANSAS: Aa Bm Co.: Brinkley 47 (Au). OKLAHOMA: Beckham Co.: C. T. Eskew 1605 (Ok). “Blaine Co.: Cutter sen. [Salt Creek Canyon, May y 5, 1951] (Ca—-986212, Ok); J - Engleman . 1270 (0k), 127k (Ok), 1275 (Ok); G. J. Goodman 2377 (Gg-= 310869, 0869, 1o--13272, N, Ok, Se—L1665); J. B. Taylor 112 (St). Caddo Co.: D. H. Dunn a (st—-20059) ; Edw, Palmer r 2h5— (C, C). Canadian Co.: Ke Hopkins 1399 (Ok). Carter Co.: M. Hopkins 4693 (S); A. G. payne 21 37 (0k); Rice 36 (Ok); Rinehart 192 (Ok). Cherokee Co.: C. Wallis 5u6 (St). Cleveland Co.: Hambrack 51 (Ok—1028h); Jeffs “is "(= 1956) M. Nielsen s.n. nog 16, 1924] (Ok--10278). Comanche Co.: P. Re Beck 1yh (Ok); Bull 38 (Ok—-14911) ; Chilcott s.n. [Lawton, 1926] (Uk (im—-20); We M. Ke Clemens 11752 (Ba); C. T. Eskew 15143 fs ok), 1570 (Ok), 1676 (0k); 8. Fry 173 Fats anys S.n. [April 23, 1938] (Gg—~310870); M Me Gates s sn. n. {April 2, 1950] (Ok); | Hopkins, Nelson, & Nelson 199 (Au--122517, Ca—72198, Ok), 9 970 (Ok); E. L, Little L72 (Ok—16614) ; D. McLean 273 (Hi--152271, “St—17800); W. S. . y= ers 31, Ber Waterfall 11938 (Au--12251), st), 13073 (St, Pare 2260919 9). ter Co.: Mericle 495 (0k). Garvin Co.: Co.: Edw. 246, in part t (38) (T). Grady Co.: Dahms a. St~-9257) 3 ee » & Merkle 107 (Ok). Greer Co.: Bull sen. [April 25, 1927] (ok—1503); Lo Logan 50 an 50 (Ok). Haxmon Co.: Waterfall 9410 (St), 13126 (Au--123206, St, W--2260941). Harper Co.: Ge Je Goodman 2398 (Ok). Jackson Co.: B. E. Cole 45 (St--21763); Demaree 12218 (Ok=-20379) ; E. L. Little 532 (Ok—10317); Small & Wherry 1220 (N). Jefferson Co.: Rohrbaugh 493 (Ok). Johnston Co: He We Houghton s.n. [G. W. Stevens 3570] (N); Edw. Palmer 27, in | part [390] (T); G. T. Robbins 3256 oer sas Ok). Kingfisher Coe: 3 (St—1L4]452); J. Eng 196), Moldenke, Monograph of Verbena 35 C. Smith 510 (0k). Major Co.: Goodman & Waterfall 420, (Au— 122518, N, Ok, St). McClain Co.: M. H Hopkins 1278 (Ok). McCurt- ain Co.: Demaree 12630 (Ok—22232); Waterfall 15897 (Hi—195395, Hi--200233, St). Murray Co.: J. F. Brenckle 48066 (N, W-~20038 23); Coleman 18 (We); Couch 0-6 (Ok) 5 E. E. E. Dale . 19 9 (Ok), 297 (0k); paeree 12292 (Ok=-222)5) 3 G. J. Goodman 2079 (Ok—17715); M. Haw- , in p part (Ok), 39 (Ok); 1 M. Hopkins 2757 (Ok), 2766 (Or), 1802 ony 5091 (Ok), 5735 (Ok), 5961 (Ok); Hopkins, MacDowell Copeland 6390 ( (Ok); Hopkins, Nelson, & Nelson 139 (Au-~-122516, &). 150 (Ok), 159 (Au, Au--122519, Ca-- 7,233, Ny , Ok, S, St—26117, W—1862729), 905 (Au, Au--122515, Ca—733376, Mv, Ok); D. J. Hutch inson 75 (St); Rice 23 (Ok); G. T. Robbins 2hh9 (Ca--9039%6, Ok) ; Stratton 11 (N, St--26120, St—22121); G. W. Stevens 28 (Du--6569h, N, Ok, Ok, | St--9221, Ur, ¥--588952), 28b (Ok—20731), A»28 (St— 9220); Tripp 122 (St--9258) . Muskogee Co.: M. Hopkins 3978 (Ok); E. L. Little 1576 (Ok—~10316). Oklahoma Co.: M. A. Carleton 13k (Ka, W); Waterfall 389 (Ok), 1117 (0k, st), 2600 (Ok), 2622 (Ok, St). Payne Co.: F. Clark 254 (St--9256); Kenworthy 23 (Au). Ro- gers Co.: Be. F. Bush 1275 (C--type). Sequoyah Co.: Ce S. Wallis 6490 (Au—17137h, St). Stephens Co.: Mericle 61h (Ok); Waterfall 3652 (Ok, St). Washita Co,: Stratton 295 (St~-26133, St—2613h); Waterfall 11916 (st). County undetermined: G. W. Stevens A.15.5 (Ok). TEXAS: Archer Co.: Purcell (au), sen. [Archer City, April 4, 193k] (Au); Whitehouse 9555 (N, “No—1)970), 9614 (Mi, N). Atas- cosa Co,: Mebane s.n. [Pleasanton road, 3/2/1926] 6] (Wi); E. J. Pal- mer == Serato ks Bastrop Co.: Bogusch 748 (W-~1285713); He val 1) (W—~1569751); Marrs s.n. , (Smithville, 3-17-1930] (au); Sa 5705 (Au--12280); E. Je Palmer 33367 (N, W--1532535). Baylor Co.: Whitehouse 9630 (Mi ay) Bee Coe: Fe 2 Fe Be Jones 1732 (Wr) 5 M. E. Jones 29191 ~(Po—-199869) « Bell Co.: Normand s SNe ne (h/ 4/28] (Au); Wolff 607 (Tr--16210), 1373 (Hp, Tr--16212). “Bexar Co.: Clare 52h (Gg—267606); Clemens & Clemens 975 (Po--69915), 976, in part + (N, Po--69562); Havard s.n. [San . [San Antonio, March 1882] (—11,757h, W-~218872); G. Jermy 8h (W-—315683), 85 A Fee Sn. [190k] N), sen, [San Ant.] (N, N); Kidder sn. [San Antonio, 25 Mch, ony Gay he [een Lindheimer 134 (Ka); Metz ee (N), = (I, I, S), 2479 (N), 32h6 (P1—151578); ); Patoni s.n. (San Antonio mee "out (me (Me, Me); Rip Ripley & Barneby 413 (Ge—-30132h, N); E. + Schulz 767, in part (Wi), son. [So. S. S. A., h-1)-22] (Au), Sone i mi. 50 so. San Antonio] (Wi, Wi). Blanco Co.: E. J. Palmer 13281 (B); Tharp s.n. [3/13/38] (Ca--882799). Bowie Co.: Tharp son. (Salted ibe eae occ; (Au). Brazos Co.: Cory s.n. [College ege Sta- tion, 49-26] (Tr--1620h); H. B. Parks 14h (St), sen. [College Station, December 1h, 1916] - (Au=—122569) § | Reeves 198 (Cs); Rowell 36 Set TO O00 1h Vol. ll, no. 1 9051b (N); Trew 188 (Au--122577). Brewster Co.: Cory 28682 (N); Cutler 672 (Tr--211)1) ; C. Newnan s.n. (Sperry T.127] (Om); Parks & Cory 18716 (Tr), 18717 ( (Tr); Sperry 493 (W-—-1679313), T.127 (W- 1679250), T.565 (Om), T ), 1.566 (Fs, Om); W. P . Taylor sen. [3~31-hbh]) (Au); B. H. Warnock C.800 (N), W W.166 Gia ‘Sake Co.: Ecology Class Univ. Texas s.n, (Falfurias, 3.2.30] (Au); Lundell & Lundell 10071 (Id, N). Burnet Co.: Cory 51,029 (Rf); E. We Lathrop 2020 (Lw); C. L. Lundell 15050 (Rf); C. M. Rogers 6668 (Au—-122565) 5 Whitehouse s.n,. [Marble Falls, he1303 30] (Au). Caldwell Co.: Mc au sen. [3/6/31] (Au), sen. [Spr.-Sum. 1931] (Au). Cameron Nn COs Je Fe Brenckle )7-338 (N); Cory 28300 (N), 28308 (N); Fleetwood saz" (Au--192005) 5 H. C. Hanson 30) (N), 322, in part (Bl—l2329, Ka~60567, Mi, N, Ur, W—982768), s.n. [Brownsville, Jan. 30, 1919] (Au); F. L. Lewton 21) (Ar——271609, W); Tharp 1871 (Au, W-~ 1203149) ; Ca He , Townsend 8 (W~279251). Childress Co.: Biology Class Univ. Texas s.n. 3 8.n. (Childress, May 1930--31] (Au). Coke Co.: Cory 37143 (N, N). Comal Co.: Lindheimer 1075 (Au, Br, Ca~- 17516, de ag Me, N, Ok--10350, Up--18538, W iy Par ncaa Comanche Co.: Phi Ne (Camanche, April 1931) (Au). Cooke Co.: De Se Correll Tee (Rf). Crosby Co.: C. 0. Erlanson 1160a (ui) 3 3 Wolff 2031 (Tr--16213, W--1623983). Dallas Co.: Boll s.n. [1871] (Cb); I. E. Diehl 6 (Po—2l8006); Lundell & Lundell 8578 (Ld, Mi, Mi, N, N), 9178 (id, Mi), 12818 (Rf, Sm); J Reverchon 738 (Up--17102, ¥, W--1],7573) » Sen. (Curtiss 1963] (Be), SoM. [Curtiss 19634] (Go), sen. [April 1878] (Bc); M. R. Stephenson 199 (W--1580366) , sen. (vicinity of Dallas, Apr. 1, 1928] (W—-1580365), sn. [near » Apr. 6, 1929] (W—15801,88) . Denton Co.: B. Be "Harris SMe tspeiiie 126] (Nt, Nt); Lundell & Lundell 8431 (Ld, Mi, N); N)} McCart 1597 (N, Nt). Dickens Co.: Parks & Cory 13619 (Tr=~16205) « Dim- mit Co.: Hoglund s.n. [Carrizo Spr., 4.6.30] (Au). Duval Co.: Croft 6515 (W--147572). Eastland Co.: Cox s.n. [Rising Star, l- 22-31] (Au); H. E, Parks 14393 (Md); parks & Cory 13146 (t2--26201), 14392 (Tr—16201) ; Wolff 2057 (Tr-~16211). Ector Co.: Lundell Lundell 11)01 (N, Rf). Edwards Co.: Cory 37068 (Au), 37070 7070 (N, “"), 53678 (N); Cutler 789 (Ba); M. E. Jones 26220 (Du-~235291, Ge— 23786, Po--178867), | 26228 (I, Po~178363). Hrath Co.: Gough s.m- [3/25/22] (Au). Fannin Co.: irs. J. M. Milligan s.n. (W-—50,332) « a eee Fisher Co.: T. M. Shepherd s.n. [Ray's Pasture, Grady, Mch. 31, 1901] (W--13916). Frio Co.: Barkley, Painter, & Lucas 14201 (S)3 Painter, Lucas, & Barkley 14201 (Au, Ok), 14213 (Au, B, Ok, Ok, S)- Garza ie Cc. M. ~ Rowell 5636 ( Au--195257). Gonzales Co.: Bogusch son. [2-20-1926] (Au); Johnston & McCart 5226 (Au--122h73); E. De Schulz s.n. [Ottine, Spring 192k] (Wi); Tharp s.n. {ottine, 36] (Au). Grayson Co.: Harold G Gentry 87 87 (Au), 50-94 (Au--12257h) 5 196) Moldenke, Monograph of Verbena 37 McCart 1575 (Au, N, Nt, W--1825822); Van Meter 11 (N,N). Guada- lupe Co.: Johnston & MeCart 5037 (Au--122),7h). Iamilton Co.: Cory 54000 (Rf); Wolff 2091 (Tr). ee Cos: C. Re Ball 111) (N, W—501587) ; Herb, | Univ. Texas sen. [Quanah, 4-21-1931) (Au); arks & Cory 13373 (Tr--16209); S ae Dept. Univ. Texas s.n, Wa/3 31] (Au). Harris Co.: G. L. Fisher 330 (W--503329). Hays Cos: Friesner 101,01 (Bt-—-387)0); Stanfield s.n. [San Marcos, Aug. 1897] (N).» Hidalgo Co.: Cory 28214 (N). | (N). Hockley Co.: Rachaner 131, in part (Au), 135 (Au, N). Howard Co.: Parks & Cory 12538 (fr). Jack Cos: Whitehouse 15311 (sm). Jeff Davis Coe: Tracy & Earle 178 (Au, Cm, Es, It, N, N--photo, Tr, W--l4l1837, Z--photo) ; Warnock, | Hinckley, & McVaugh 3795 (N). Jim Hogg Co.: McCart 71,80 (Ok). jim Wells Co.: Freeborn & | & Freeborn 315 (N). Johnson Co,.: Cory 53968 (Rf); Whitehouse s.n. [Cleburne, May 3, 1931] (Au). Jones Cos: Ne C. Henderson 63-32 (Au--217678). Karnes Co.: Col- lector undesignated s.n. [Brackenridge Park, 3-13-19] (Wi); de | 2 C. Johnson 1128 (Z), 53-870 (Au--122),70). Kaufman Co.: Harold Gent- ry ry 2699 (Hi--116717); Fe Fe. J. Tyler sen, [vicinity of Terrell, April 23, 190k] (W—500805), sen. [vicinity of Terrell, May 13, 190h)] (W—-500806). Kenedy Co.: Harold a B66 (Au--122505) ; Lun- Gell & Lundell 8731* (Mi). Kerr Co.: « Palmer 9278 (Bi, “Du 205593, S). Kimble Cos: M. E. Jones Ls T (Po--190085). Kinney Co.: Mearns 1282 (Du~—2,193, “W—-229528). Kleberg Co.: M. C. John- ston 53253.19 (Au—~122h67) ; J. F. Sinclair s.n, [Kingsville, Spring 190] (Au); Tharp & York - 51-240 (Au—122583, St). Liveoak Co.: Tharp son. [March 2. 1931) (Au); Whitehouse s.n. [March 22, 1931] (Sm). Llano Co,:Barkiey, Webster, & Tharp 47268 (Au, N); Ge L. sisher s.n. [Bluffton, Apr. 20, 1931) (B, Bt--37796, Gg— 22255); ¢ E. Jones s.n, [Llano grande, March 27, 1932] (Po 199868); A Sassier 95 (Au--16965k) 5 en AS & Warnock tee) (Au); Whitehouse sn. ., [Enchanted Rock, Spring 131] (au). Lubbock Coe: Demarse 750 (Du—205682, W--1699018) ; Studhalter 1118 (W- 1285908). Maverick Co.: Bogusch s.n, (February 17, ary 17, 1927] (Ob== 50827). McCulloch Co.: Studhalter 1106 (W—-1285900). McLennan Co. Ricker 33% (W--501878). Medina Co.: C. L. Lundell 10961 (N, Rf, Rf, W—1926939). Milam Co.: G. L. Fisher s.n. [Rockdale, Apr. 18, 1931] (Ew). Mitchell Co.: Finley 9a Sa (Au); Re Re W. Pohl 48 48143 (sm). Montague Co.: Whitehouse 15038 (a, N). “Nolan Coe: V. Ee Stanfield 3 (Au). Nueces Co.: E. Be Jo « Palmer 11222 (Ca— h2séen). Palo Pinto Cos: M. Parks s.n. [Palo Pinto, April] (Au). Parker Co.: Shinners 10954 (11); Wadsworth s.n. [Milsap, 1930-~31] (Au). Randall Co.: Lundell & Lundell 11135 (N, Rf). Bh Co.: Wild Flower Contest 2 (Au). Red River Co.: Wilfong 16 (Sm). Reeves Cos? Cory 27906 (N); Tracy & Earle 106 (Cm, T (Cm, Tr, W-—-Lh1808) . 38 PHYTOLOGIA Vol, 11, 34 Refugio Co.: F. B. Jones 1712 (Ww). San Patricio Co.: Gould & Hycka 8061 (Au--19266); F. B. Jones 1808 (Ww); C. M. Rowell 4776 (Wr), 5255 (Au—195926); E. De Schulz 845 (Wi); Tharp sen. [Sin- ton prairie, 3/23/31] (Au); Williges 25 (Au—175629), 357 (Au-- 179741). San Saba Co.: Lewis & Oliv & Oliver r 5435 (Nb); B. Norton Sone [Cherokee, 4-1-1931] (Au); E. J. Palmer 116h3 (Au--122565, Ca——— spe Schleicher Co.: Cory - hy (N, i. Shackelford Co.: Ne . Henderson 63-38 (Au--217678)> Fok Rohrbaugh 70 (0k). Sutton Cost 7 E. Jones 28512 gee. Re We Pohl 4759 (Sm); Rohrbaugh 372 (Ok). Tarrant c oo og (Ku, W—1h65237); Ruth 110, in part (Cm, Io--1l 366 SE, Up, Ur, W--1188947). Taylor Cos: Tolstead 7071 (Au, Au, mn,’ Ca~2882601 , Mi): Tom Green Co.: Cory 40913 (N, N); Lewis & Oliver 51,69 (Nb); Parks & Cory 5620 (Tr), 5621 (Tr). Travis Co.: CG. C. Albers 31,006 (Au); Armer 5361 (au, W--1,38995), sno (Austin, ]-2-29] (P1—151557); Bodin 217a (W); Bogusch & Molby 3 3009 (Ur); Seay 2 98 (N); Ely sn. [Pilob Knob, 21 March 195] (Au—122192); E. Hall 431, in part (N, Po—2i8007, Pr, W), son. (Austin, 1872] (W--771876); Herb. Univ. Texas sen. [Deep Eddy, h-26-36] (Au, Au, Au, Au); Long & Ferguson sen, [Bee Creek, ay (Au); Mart artindale Son. [Austin] (Up—17103); 3. J. Palmer 9278 (W--1531907), 9339 (Bi, Du—205592, S, W—153191h), 10003 (W-=1532016); Tharp 1362 (Au, W--111):13), "1364 (Au, ee), (Au, Ca—882798, N), sen. (Country Club, ub, +728] rage Be Be H. Warnock 20623 (Au, N), , 46086 (Au, Ok, vi); York s.n. {Austin 3-18-08] (Au); M.S. Young 11h (N, Po—161333), 15 (Au, Au, Ur, Vi), Sen. [Deep E Eddy, 11/15/13) (Nt), Sete [2/28/14] (Au, N), SMe [Sandy Creek, 4/22/15] (Ca~882802), sen. [Austin, 3/9/18] (Ca— 882803), sen. (Austin, 3-12-18] (Mi, N), sen. [Austin, 8-25-18] (Au), sen. [west of IAGN Ry.] (Au). Uvalde Co.: i sen. [U- valde, 4/17/31] (Au); H. J. Hamby 573 (Ar--206662); Herb. Hapeman sen. (April 17, 193i] (S)3 M. E. Jones 28298 (Po—188288), 28518 (Po--190121); E. J. Palmer 13562, “in part rt (B); Parks & Cory - 8392 (Tr—16207). Val Verde Co.: Cory sen. [Del Rio, 3-17-1927] (Tr, Tr——-16206); H. C. Hanson 543 (W—-952686) ; Tharp & Havard 1.9360 (Au--122559). Walker Co.: E. J. Palmer 13175 (W--160 3455) . Ward Cos: Lundell & Lundell 11382 (Sm). Webb Co.: M. E. Jones 29189 (Po--199870, Po--199871) « Wharton Co.: E. J. Palmer 480 0 (Po-= 217663, W--1581011), 4977 (Po—-217662, W—1581010). Wichita Co.: Rohrbaugh 488 (0k); * jhitehouse 9513 (Ca—7h571, Mi, N, W--1947162), 952, (Au--122560, N, Tl), 966 (N), 9669 (N), 10903 (sm) « Wilbar- ger Co.: Whitehouse 9817a ee hice N). Willacy Co.: J. Fe Brenckle ),7-366 (N); M. C. Johnston 54170 (Au—122)68). Williamson Cos: Wolcott 121 (Au). Young Co.: Correll & Johmston 22126 (Id); Hayes 610 (N); McCart 862 (Nt); Whitehouse 15323 (N). Zavala Co.? Herb. Univ. Texas s.n. , (Crystal City, ]-6-1930] (Au); B, Stewart 40 196), Moldenke, Monograph of Verbena 39 (Ok, 0k). County undetermined: Collector undesignated s.n. [/ 1/26] (Sr); Lindheimer III.501 [Herb, Reichenb. f. 9221] (ca-- 21,8243, V, W); Metz 3095 [Texas Military Inst.] (St); Sarpin Sen. (Au, Au); C. Wright 1500 [N. Mex.] (Ca—~27678, T, 7). MEXI~ CO: Eddy Co: Cory sen. [Carlsbad, Apr. 192] (rr); De rope ies 109 (W--1203666), “County undetermined: Parry, Bigelow, Wright, & Schott 830, in part [Valley of the Rio Grande de below Dofiana Dofiana] Fy 56216); Wencurtin s.n. [5-2-1916] (Au). MEXICO: Coahuila: Johnson & Barkley 16205M (Au), 16208M (Au), 16212M (Au), 1627" (Au); War- 3 War- nock & Barkley 1798M (Au). Nuevo Leén: Alonis, Johnson, & Bark- ley 1611M (Au), 16175" (Au); Frye & Frye 2376 (Hp); J. N. Weaver 00867, En— En— 1033 (W—21 34239). Sinaloa: Me E. Jones 23243 (Ca—l00 6122, Gg~-172668, N). Sonora: is Sart S. . Gentry ry 1311 (ca-61s6302, Fs, Ge, Ge——2955h0, I, Ka--94678, Me, Me, Mi, N, N, S, S); Rose, Stand ley, & Russell 12745, in part (N, W#—6 35561). Tamaulipas: Painter & Barkley 11383 (Au). CULTIVATED: Texas: E, Hall s.n. [Austin] (W). LOCALITY OF COLLECTION UNDETERMINED: Edw. Palmer 168 (Sd— 6781). MOUNTED CLIPPINGS: Pittonia 5: 137.1903 (N). IA PUMILA f. ALBIDA Moldenke, Résumé Suppl. 7: 3, nom. nud. (1963), Lf, nov. Haec forma a forma typica speciei corollis albis recedit. This form differs from the typical form of the species in hav- ing white corollas. The type of the form was collected by Robert Runyon (no. 299) - Sie sandy ground, sandy loam soil, on Highway 96 at the edge Hag motes, 2 at the New York Botanical Garden. Cory reports that his no. 53500 had the corolla-limb 2-3 mn. wide, but was ¢ “along with no. 53501 whose corolla-limb is 5 mm, or more wide; frequent on graded roads" in Pecos oon Texas. McKelvey 1931, actually hav been collected t k pale-gr e toothed or dissected, and the & cylindric capsule, "frequent in s soil, the plants root at the lower nodes. s that his no, 2366 was collected on January 17, 191, in the same locality as the type. About his no. 2359 he says “in the sandy region ae plant roots freely at the nodes “8 it is frequent but not common. arium material has usually “a misidentified and distribu- ted a as a. adrangulata Heller. C. M. Rowe 11 4776, fram San Pat- Ticio County, cited herein under V. pumila, bears a notation on its label reading "with pale serie to white corollas". In all, a herbarium specimens, including the type, have been by m Chtattenss TEXAS: Bexar Co.: Kidder s.n. [San Antonio, 25 Mch. 0 PHYTOLOGIA Vol, 11, no. 1 1885] (Oa--10739). Cameron Co.: R. Runyon 2366 (N). Kenedy Co.: Be Runyon 2359 (N), 2499 (N--type, N--isotype, N--isotype). Kle- rg Co.: Correll & ieee 15616 (Rf). Pecos Co.: Cory e860 (Du-=362506); Parks & ory = (Tr). Val Verde Co.: McKel- vey 1931 (ac sbOKG yy W--1581390 VERBENA QUADRANGULATA Heller, Contrib. Herb, Franklin & Marshall Coll. 1: 84--85, pl. 6. 1895. Synonymy: Helleranthus quadrangulatus (Heller) Small, Fl. Southeast. U. S., ed. 1, 1011 & 1337. 1903. Helleranthus quadran- gulatus Small ex Prain, Ind. Kew. Suppl. 3: 87. 1908. Verbena pum~ tia fs “albiflora standl., Field Mus. Publ. Bot. 4: 256. 1929. Verbena quadrang. quadrangularis Heller ex Moldenke, Résumé Suppl. 2: 13, in syn. 1960 {not V. quadrangularis Vell., 1827]. Verbena guadran- gularis Heller ex Moldenke, Résumé Suppl. 6: 10, in syn. n syn. 1963. Verbena & pandrangulate Heller ex Moldenke, Sheek Suppl. ll: 7, so graphy: Heller, Contrib. Herb. Franklin & Marshall ant 1: BBS pl. 6. 1895; J. Ks Small, Fl. Southeast. U.S 1011 & 1337. 1903; Durand & Jacks., Ind. Kew. Suppl. 1: Wen. 19085 Prain, Ind. Kew. Suppl. 3: 87. 1908; J. K. Small, Fl. Southeast. U. S., ed, 2, 1011. 1913; Standl., Field Mus. Publ. Bot. : 256. 1929; Stapf, Ind. Lond. 6: 31. 1931; Perry, Ann. Mo. Bot. Gard. 20: 2h5, 2kb--250, 311, 313--3ih, 339, & 356. 1933; Cory, Texas Agr. - 550: 89. 1937; Moidenke, Prelim. Alph. List u Invalid Names 25 & 50. 1942; Moldenke, Known Geogr. Distrib. Verb- enac., ae 1}, 13, 19, & 102. 1942; Moldenke, Alph. List Cit. 1: 126, feat 162, 176, 269, & & 29h. "1946; Moldenke, Phy- tologia 2: “tbs (1946) and 2: 328°& 383. 1947; He Ne & A. L. Molden- ke, Pl. Life 2: ae 1948; Moldenke, Wrightia 1: 230. 1948; Molden- ke, Alph. List Cit. 2: 393, 398, 69, 492, 493, 496, st 527, 55h, » 915, 2'(1949), and kh: 990--992, 99h, "100 1099, 1113 ty “39, ish, iy 1212, 1213, 1217, laid, & 125 "1993 ion denke, Phytologia 3: 132. 19,95 He Ne & Ae Le. Moldenke, Anal. Inst. Biol. Pps 20: 1h. 19493 Moldenke, Knom Geogr. DMatrif. Verbenac ., [ed. » 33, & 199. 19493 Shinners, -giag Pie vi — _ Br Se iosby Jones, Rowell, & Johnston, Flow ast. Be bh. 1961; Moldenke, Résumé 30, 39, 298, 313, ie nr Pa on Molden- ke, Résumé Suppl. 2: 13, 1960; Lewis & Oliv v., Am. Journ. Bot. 8: “ 639-641. 1961; Moldenke, Phytologia os a (1961) and 8: 279 & Las. in Moldenke, Résumé Suppl. 3: 7 (1962) and 6: 2, & 10, 1963; Mol- enke, Phytologia 8: ire ti9 63) and 9: 1, 2h, & 62. 1963. Tllustrations: Heller, Contrib. Herb. Franklin : “Marshall Coll. 1: ag 6. 1895; Lewis & dliv., Am. Journ. Bot. 48: 60, fig. 3. ow annual herb, decumbent or prostrate to ascending, more or 1964, Moldenke, Monograph of Verbena hl less diffusely branched from the base; roots fibrous; stems pros-~ trate to ascending, rooting at the lower nodes, hirsute; bark pale-lavender, light-violet, or pale~blue, or even red, its tube slightly longer than the calyx, practically glabrous outside, the limb ing-calyx 5--6 mm. long, hirsute, particularly along the ribs, its lobes short, subulate-tipped; fruit cylindric, schizocarpous, con~ stricted along the lines of cleavage, the cocci about | mm. long, surmounted by a smooth rather obtuse beak, shallowly reticulate- Scrobiculate above, longitudinally striate and somewhat abruptly enlarged at the bas se; chromosome number: n = 10. The type of this species was collected by Amos Arthur Heller (no, 1388, in part) in open grounds at both the northern ae southern ends of the town of Corpus Christi, at an altitude 10 to 35 feet, Nueces County, Texas, on Marc ch 5, 189), where "it was described as being "plentiful", and is deposi ted in the Brit- ies Herbarium at the uae York Botanical Garden, inscribed "Verbe- quadrangularis", rial under this collection number on ait no. 46200 in ‘i ginny State College herbarium is Cressa a- phylla in the Convolvulaceae. Heller (1895) says: "This most peculiar plant has the ha d general appearance of the species of Verbena which bear ce large and showy flowers, as V. bipinnatifida and and V. Aubletia, but known, it probably an undescribed genus." This doubt on the part of its original author would probably cause some literal interpreters of the International Rules of Botanic Nomenclature to maint. hat the binomial has not been validly published by Heller and must be replaced! It should be noted here that V. quadrangularis Vell. is a syno~ nym of V. bonariensis L. The type of V. pumila f. albiflora was collected by Lydia E. Pagel (no. 2208) at San Antonio, Bexar ty, Texas, in the spring of 1926, and is no. 580190 in the herbar~ ium of t. the Chicago Natural History Museum Verbena quadrangulata has has been collected in sandy, loose sandy, 2 PHYTOLOGIA Vol. 11, no. 1 rocky, clay, limestone, gravel, and grayish calcareous soil, in sandy loam, brormish sandy loam, and dry clay-loam , in loose sand, O00 places, woods, swales, and fields, mesquite brush and brushland, in waste or open er eared mesquite scrub and open desert s and on road shoulders or sandy road shoulders and caliche slopes, as we railroad rights-of-way, flowering from cribed as it: waste and open grounds, fields, and desert scrub, clay slopes, sandy beaches, blo from February to in the southern and western portions of the rom flowe pe d hilltops in Hidalgo and Starr Counties." Warnock reports it "fre- ent" c in Webb qu in Crane County, and Hamby "more or less common" We refers to it as "a weed groves"; Crutch- fielec & Johnsto ed: sandy clay-loam along r. pasion way"} McCart found it "in soil grading pale orange light brownish ne loam"; and caiman (1958) writes bina in Rio Grande Plain, reported from Tarrant Co., far out of range, probably because se plop ee: of specimen, but possibly intro- duced "pasture sae! on well-drained sandy or clayey loams and cal- oe spr abundant ® in Kleberg County. The only common name reported is i "regard to flower color, most collectors fail to mention heir labels, but it "is given as "white" on ckel vey 1786 ane Passed 1780, 2183, 2496, & 2561, as "whitish" on Cory on ory 51369, Re R. Runyon on 628, an and Turner h96, as "nearly white, tinged hiner" on n Lundell & & Iundel1 9803, as as "nearly white, tinged laven- der" on Lundell & Lundell 1012) & 10226, as Mnearly white, tinged bluish-lavender" on Lundell & Lundell 9790, 9793, 10050, 10078, & 10141, as "pink with white throat” on Ecology Class Univ. Texa Texas Sen. [Pearsall, 2-27-30], as "pale lavender" on c. Le » Lundell 10692, as "light-violet, nearly white" on F, D. Jones 1773, and as ‘Wpale~blue" on Frye & Frye 2376 and Cc. L. Lundell Lundell 13621. Williams "flowers red or white", More field study study is eigen be- fore we can be sure if os color forms are involved here merely cases of fading due t e. » Davis, in a letter res me dated March 25, 1912, refers to only half as L. two forms of this species -~ one with the corolla 1964, Moldenke, Monograph of Verbena 43 wide as the calyx is long and the other form (from La Joya) with the corolla-limb 6-~8 mm. wide and exactly equal to the calyx length. The soil in the two localities, he says, is entirely different. Material of this species has been misidentified and Sepeoait ted in herbaria under the names V. ambrosifolie 2 Rydb., V. bipin- natifida Nutt., V. bracteosa Michx., V. canadensis (L.) "site: V. canescens H.BoK. » V. ciliata Benth., V. ciliata var. longiden- tata Perry, V. pumila Greene, and V. pumila Rydb. On the other hand, the Fleetwood a 32, G Gould & Hycka 8061, Mc Cart 7480, and C. M. Rowell 1776 & 5258, distributed as V. quadran = are actually V. pumila Rydb., ., while R. Runyon 2359 & 2366 are V, pumila f, albida Moldenke, A.A. Heller 1365 is a mixture witty v. ciliata var. longidentata Perry. Croft s.n. [San Diego, 1885] was S apparently first identified as V. ciliata Benth., then as V. pumila Rydb,, and finally as V. racemosa Eggert by Farwell. Edw. Palmer 105 was annotated by Perry as "Aff. V. delticola Small” and a pencil notation on the sheet says "seed has commissure long-warty." E, J. Palmer 13562 was cited by Perry as V. based on St. Louis | and Was Washington specimens, and I annotated the. — specimen as V, pumila also, The Washington specimen, how- » has been examined by me and is cited below — it has cocci with’ definite beaks! Similarly, Frye & Frye 2376 was regarded by me as V. pumila, based on Minden and Washington specimens. These collection should also be re-examined, since the Washington sheet of this number nap eceoiag sea not have beaked cocci and is there- fore cited by me as V. pumila. Without rota fruit it is almost impossible to mandierion te these two spec The Herb. Univ. Texas s.n. [ Pear oe "ited below as agian Wells County, Texas, may actually have been collected in San Pa ricio eases bie Sabai 4a inscribed merely "Alice--Mathis", but Dr. Tharp, in a letter to me, states “likely in Jim W County"; similarly, Tharp s.n. (h-19-35) and s.n. (Apr. 19, 1931], aoe from Jim Wells County, m: may have been collected in Du unty, since the labels are merely inscribed maison Ome Diego", send gh Dr. Tharp, in his letter to me, says probably Jim Wells Geuntye, Hoglund s.n. [h-6-30], cited from there County, may have come from from Zavala, since its label is inscribed merely "Carri- zo Spr.—Crystal City"; Hoglund s.n. [.6.30], cited from Medina County, may have originated in Frio, because its label is inscrib- ed merely "Devine—Dilley"; McKelvey 1786, cited from Webb County, bh PHYTOLOGIA Vol. 01, nowd label is inscribed "Kennedy~-Portlan Perry (1933) cites the following a additional ee not as yet seen by me: TEXAS: Atascosa Co.: E. J. Palmer 1747 (E). Bex- ar Co.: Pagel 2208 (F); E. J. Palmer 2039 9 (G). Cameron Coe: Neal- ley s.n. [Brazos Santiago, 1889] (F. W). Frio Cos: Eggert s.n. Trailroad north of Moore Station, 5 April 1901] (E). Kinney Co.t Trelease 101 (E). Kleberg Co.: High 7 75 (E). Nueces Cos: Ae Ae Heller 1388, in part (E--isotype, G—-isotype) ; Ee Je sung TOL6 (ee Uvalde Cos: E. J. Palmer 23592 (E, G). Val Verde Co.: “Or eutt s.n. [Devils River, 15 May 1913] (E). Webb Co.: Berlandier "1485 = 225" (G); aera sen. [Laredo, 6 April 1901] (E, G); oe cutt 55h2 (E), 5730 (E)3 J. Reverchon 3902 (E, G). MEXICO: aulipas: Berlandier "3018 = 1516" (E). ~The Berlandier "1485 = 205", cited by her as from Webb County, may have come from on, plate where, since its label is inscribed si se Laredo and Bejar Her "Canby 193 193" is cited below by me as Canby, Sargent, & ae age 193; her "Nealley Sen." may possibly = the the Nealley 11 11/36 9 cited below by me; her e3; her Croft 78 from "NY" is actually in lly in the Columbia University Herbarium. She says "Verbena quadrangulata is an anom= alous species often confused with V. pumila on account of the strong similarity in habit, but is p perhaps more closely related to V. delticola, the only other known North American species with a sarang toward developing a nutlet with a beak. The style is be enlarged at the base into a persistent subhemispherical body; hence, when the nutlets split apart they appear as if elongated at the apex into a beak~like appendage. Although the anthers are not glandular and the flowers not showy, the species seems to be- long to the section Glandularia." In all, 169 we aa cen including the types of most ed d of the names * involv: unted illustrations and clippings have been examined by me. Citations: TEXAS: Bee Co.: Lundell & Lundell 10050 (Id, N); Tharp 5538 (Au, W—1468817). Bexar Co.: Clare 755 (Hp); J. B. Daw 16 (Ca—25137); Metz 755 (I); E. D. Schulz 475 ; (W-—-1074921), 767, part (W-~12),3198, Wi), s Sm. [May 1921] (Wi), s sen. [s0. S. Ae, Le aoe, (Au), sen. [9 mi. s 0, San Antonio] (Wi). Cameron Co.: Me Cannon s.n, {La F Feria, February 25, 1926) (Mi), sen. (Santa Rosa, February 7, 1927] (Mi, Mi); Cory 51369 (Sm); A. M. M. Davis sen. [Ol- mito, Summer ig (Au); C. L. Lundell 10692 (N, Rf, W—1926931); Nealley 114/369 (W); L. H. Pammel S.n. (Brownsville, Feb. 12, 192] (Io), sen. [Point Isabel, 2-25-29] ~ (To—-11h036), Sen. [Rio Hondo, 3-2-29] (Io-~11)036); R. Runyon 628 (Rr, W—122),221), 1780 (Rr), 2183 (Rr), 2496 (N, N)j ‘Schott s.n. [Rio Bravo del Norte ‘te, March 1852] (N); Small & Wherry 11903 (N); G. H. Thomas s.n. (Fort Brown} (N); Wherry s.n. [Brownsville, April ih, 1925] (Up). Cherokee Co! Je Fe Brenckle 47-379 (N). Crane Co.: B. H. Warnock 1693 (Rf). 196) Moldenke, Monograph of Verbena Ls Dimmit Co.: Cory 28067 (N); Hoglund s.n. [Carrizo Spr.~-Crystal City, 4.6.30] (A (Au). Duval Co.: Croft 7 (C, Pa), 8 s.n. [San Diego, 1885] (Mi); M. C. Johnston 54106 (Au--122587) ; Lewis & & Oliver 5409 (Nb); Ramos, Sandoval, & “& MeCart 8007 (0k). Frio Co.: Ecolo- gy gy Class Univ. Texas sen. SNe (Dilley, 2227630] (Au), s.n. [{Pearsall, 2-27-30] (Au); Herb. Univ. Texas s.n. (Dilley, 2-27-1930] (Au), s. n. (Pearsall, 2-27-30] (Au); C. L. Lundell 13621 (Sm). Goliad Cou Ce Be Williams 9 (Au). Hidalgo © Co.: Clov Clover 596 (Mi, N); Fleet- wood _ 3200 (Au-~180075); Lundell & Lundell 9803 (Ld, N, W--1888830); Re Runyon 2561 (N)3 B. L. Turner LL96 (Au--168766) Mrs. E. J. Walker ql (au, (Au, N). Jim Hogg Co.: Salazar, Santos, & McC McCart 33 (Ok, Ok); Sanchez, Rodriguez, & McCart 8311 (0k); Tharp, “Johnson, & & Webster 18-71 (N, St). Jim Wells Co.: Herb. Univ. Texas s.n. [Al~ ice-Wathis, — 3-2-1930] (Mi); Lundell & Lundell 10078 (Ld, N); Tharp s.n, Siise-tan Diego, ily ve 1931] (Au), sen. [Alice-~ San Diego, -19-35] (Mi). Karnes Co.: Tharp 5610 (W--1468818) . Kinney Co.: Canby, Sargent, & at 193 (W—-369717); Cory 28994 (N). Kleberg Go.: M. C. Johnston 5441 (Au--122588, St), 53253615 (Au-~122589); Fe Bs , Jones 1773 (iv); R. Runyon 428 (Mg, N). La Salle Co,: Botello 9 (Ok); Cory 28548 (N); Lundell & Lundell 1011 (Ld, N). Maverick Co.: Havard san. [Eagle Pass, April 183) (W (W--218871, W--250657). McMullen Co.: Thompson & Gra- ham 61 ager Medina Co.: Hoglund s.n. [Devile—Dilley, 1.6.30 6.30] ( Nueces CO.? Ae Ae Heller 4 i oat part (P1--72008), 1388, in a (C-~isotype, Sn canto dione, Ca~-10)86--isotype, Es—i type, Io~~387),5--isotype, Mi-~isotype, N-~pho Tu--isotype, Ur-~isotype, W—~1,8059--isotype, Z—-photo of isotype); F. B. Jones 1718b (Wr); S. M. Tracy s.n. [Corpus Christi, 4/6/1905] (Tr, Tr). Pecos Coe: Parks & Cory 18351 (Tr). San Patricio Co.: F. B. Jones 2833 (Ww); E. D. Schulz sen. [4/22/1938] (Wi); Tharp s.n. (Mathias, 3-2-1930] (N). Starr Co.: Alvarez, Guajardo, Salazar, & McCart 8042 (Ok); Lundell & Lundell 9790 (id, N, W--1887592), 9793 (MN); NeCart 7234 (Ok); Ramos, Sandoval, Salazar, & McCart 7879 (0k); Tharp, Johnson, & Webster 18-73 (Au--122596, "N). Tarrant Co.: Parks & Cory 11278 (Tr). Upton Co.: Lundell & Lundell 10226 (Id, N, W Preeian Dvalde Co.: Dobie s.n, (Uvalde, Apr. 13, 1931] (Au u); M. EB. Jones 28298 (Ca--)79296, Du-—~239730), 28998 (Bm), 98298 (Gg—237850); E. J. Palmer 13562, in part (W--1603)56), 33592 (N); Tharp & Havard ward 9346 (Au--122591). Val Verde Co.: Small & Wherry 12010 (N). Webb Co.: Alwarez, Guajardo, Salazar, & Meco: McCart 7632 (0k); Blair & al. 48-198 B (Au--12258h) 5 Cantu, Chapa, & MeCart 14 (0k); C. K. Dodge @ 154 (W--8L1668); H. J. any 2 515 (A (ar- 206561); Mecart 7150 (Ok, Ok), 7361 (Ok); McKelvey 1786 46 PHYTOLOGIA Vol. 11, no. 1 1581389); Edw. Palmer 1046 (Pa); J. Reverchon 3902 (N, Po--6l699, W—5O1157); Shute 51-171) (Au--122506), Wharton Co.: J. Ke small sen. [near Wharton, April 12, 1925] (H—h373h, Hi—7905, 1738993, We); Small & Wherry 1182) (N, N). Willacy Co.: J. eS Brenckle 47-65 (N); | Cory 511,87 51487 (Sm); Lewis & Oliver 5426 . (Nb) . Zapata C Coe: He J. Hamby 55) ly (Ar—-206670) 3 Hi Herrera & era & McCart 7699 (Ok); Lundell | & - Lundell 1012), (Ld, N); MeCart 7206 (Ok) 3 Ramos, Sandoval, Salazar, & McCart ‘art 7848 (Ok). Zavala Co.: Cory 43814 (Au). County undetermined: Parks & Cory 178h1 a MEXICO: Nuevo Leén: e& 376 (Se--65763, W--1790996) ; » Palmer 1045 e-s61857; ie & . 11,508 faecal areola (Au, N). Tamaulipas: Crutchfield & Johnston 517la (au—178530) MOUN- TED ILLUSTRATIONS: Heller, Contrib, Herb. Franklin & acne Coll. 1: pl. ee 1895 (W, W). MOUNTED CLIPPINGS: Heller, cantetb. Frank- lin & Marshall Coll. 1: 8--85. 1895 (W, W). VERBENA RACEMOSA Eggert, high 2: 123. 1902. Synonymy: Verbena puichella Greene, Pittonia 5: 136. 1903 (not V. pulchella Sweet, 1829, nor Hort., 181] ° Bibliography: Eggert, Torreya 2: 123. 1902; eras Pittonia 5: 136. 1903; Prain, Ind. Kew. Suppl. 3: 187. 1908; Ann. Mo. Bot. Gard. 20: 312, 332--333, & 356. 1933; Slaues “Cytologia 7: 161--166 & 168, fig. 9 & 25. 1936; Cory, Texas 8 Agr. Exp. Sta. 550: 89. 1937; Moldenke, Prelim, Alph. wet Invalid Names Bull. 48. 190; 0. E. Sperry, Sul Ross State Teach, Coll. Bull. 22: U1 19413; Moldenke in Lundell, Fl. Texas 3 (1): 19 ‘ ums. 19423 oldenke, Known Geogr. Distrib. Verbenac. {ed. 1], 13, 1h, & 102. 1942; Moldenke, Alph. List Invalid Names ho. 19423 Se aca , Anal. Inst. rr Sta. Catalina ): ee chance Schnack & Covas, *Dar- « 19h; Dancy aki Ammal, Chromosome Atl. 270. 1953 —— ogia * 196; Moldenke, Alph. List Se rag tare = age pig 78 & 526. 1948; Moldenke, Wrightia ls = Pies: es ty tanea 13: 112. 1948; Moldenke, Am. Wild 450. af Pg denke, Alph, List Cit. 3: 682, 71, 769, 7B, a3, “387, 873, 7h, 3 95h, 965, & 966 (1949) and h: 989, 100k, 1007—1009, 1198, 1212, 1213, & 129). 1949; Moldenke, Kn Geogr. Distrib. Ve sre ig [ed. 2), 2h, 25, 33, & 199. 199; Me Dougall & Sperry, Pl. Big Bend Natl. Park 146. 1951; Moldenke, wiptoksete 5: 330. 1958; Moldenke, Am, Midl. Nat, 59: 349 & 35h. rare) Résumé 3: 7 (1962) ~ a - 1962; ‘Molden, plytologis 9: , Illustrations: Dermen, ane nae 163 tie. 9, 165, fig. 25. 1936; Lewis & Oliv., An. Journ. Bot. ib: 6 U 1, low, annual herb, einen: eh cetiee to spread- ing~erect er in erect clumps; stens several from a common Sale: ascending-erect, branched at the base, pubescent, sometimes bloom- 196) Moldenke, Monograph of Verbena 47 ing when only 2.5 cm. tall; branches procumbent or decumbent to peti re Sate usually about 25 cm. long; leaves decussate- posite, small, firm, the lower ones petiolate, the upper ses- sile; “earebladés: oval or ovate in outline, about 2 cm. long and linear, somewhat pubescent or hirtellous on both surfaces; spikes terminating the untine » short-pedunculate or subsessile, cylin- mm. long, usually surpassing the bractlets, sparsely glandu Sistel lone Gs or rough-hairy, its teeth short, shorter than the tube, acute~ or linear-subulate; corolla white or essentially so, de. spreading reticulate-scrobiculate almost to the — the commissural faces muricately scabrous; shesmoents number: x = 5; n= 10 (Lewis & Oliver) or 2n = 30 (Derm The type of this pu AT SA “was collected by Heinrich Karl Daniel Eggert in low sandy ground in valleys near Sierra Blanca, El Paso Verbena bipinnatifida with which it has been associated. It dif- fers from the latter species in the narrow leaf-segmmts and the er, pale and persistent corollas. It hybridizes with Ver- — eee? " This hybrid has not as yet been recognized by me. y hybrids known to me which involve v. recmass are wi ah A (L.) Britton inxie = ferax Moldenke] and with V. ten- uisecta Briq. [=xV. kondai Moldenke]._ The type of V. " pulchella of Greene was collected by S Mills Tracy and Franklin Sumner Earle (no. 1068) na the pamece| of hong Davis Mountains, Jeff Davis County, April. n Texas V. racemosa is said to inhabit een Py aor valleys, and foothills, blooming from January to June, the winter form of- ten blossaning when only an inch tall, in the western and central portions of the state, from El Paso to Brewster to La Salle, Bex- ar, and Houston Counties. Collectors ag oe it from reddish, limestone, and alluvial soil, eiconag and low foothills, - leys and mesa margins, rocky limestone valleys, deserts, borrow abe = ores floodplains. They eee found it on oO od flats, dry flats, plains, rocky hillsides, desert soectaine, sand and gravel bars, and pies es abene of rock an 48 PETTOLOG TA Vol. 11, nOs:2d that it often forms "large plants 2 feet broad in gravel and ae bed on rocky slopes" in Pecos County; McVaugh calls it "abundan quent" in Brewster County, "frequent along highways" in Pecos County, "frequent" in Val Verde County, and "infrequent but wide- spread along highways in gypsum soil" in Reeves County. Cory 53500 bears a label reading "corolla-limb 2--3 mm. wide; frequent on graded road with no. 53501 in which the corolla-limb is 5 mn. broad or more." 0. E. 8 perry 493 represents the winter form, the plants actually only a an inch or so tall and in full flower in Jan- uary. As to the color of the corollas, most collectors who mention it at all claim it to be white or essentially white; on D. S. Cor rell 20699 it is said to be "whitish", on — & McBryde 14750 "mostly white", on Whitehouse 19633 Mearly white , lavender tinted", on McVaugh 7940 "very enna nem, es off white", on Warnock )7011 "white to light-lavender", and on McVaugh 7951 "vary- ing to pink or blue", Warnock T.6 is Bee by the collector to have been eTreeey ee white-flowered", whatever that means Lewis & Olive ts are ts e cepting 2n = 30 for V. es ao Eggert (ors 1936). aor aa of n = 10 suggests hx and 6x races for the taxon, or, alternately third possibility: Dermen's material, which I have not seem, may pees been misidentified. Lewis & Oliver's material has been check- 2 me. Material of V. racemosa has been misidentified and distributed in herbaria under the names V. ambrosifolia Rydb., V. bipinnatifi- da Nutt., V. ciliata Benth., a pubera Greene, V. pumila Rydb., Ve f, albiflora | Standl., and ve iV. wrightii A. Gray. On the oth- er hand, the 0, E. Sperry 1.1072, “distributed as V. racemosa, is actually V. ambrosifolia Rydb., Hamby 500 and R. F. Smith i are V, ciliata Benth,, R. F, Smith ¥.10 is is V. seions Small, an : albiflora Woldenks. Parry, Bigelow, oe “& Schott 830 is a mixture with V. oe Ry’ The McKelvey 19 Sollestion, cited below as from Val Verde County, “Texas, 1 yas, may actually have been gathered in Brewster County, since its label merely reads "between Langtry & Marathon", H. Re Reed 34064 has no locality of collection indicated on its abet but V. L. Cory, in a letter to me dated November 15, mest soe that it was probably collected in Pecos County — nthe 196) Moldenke, Monograph of Verbena ug has "H. N. Moldenke* ac on its label as collector, but po’ se certainly erroneous, ery (1933) deen the following 7 additional specimens not as its sn by me: TEXAS: El Paso Co.: Eggert sn. (low valleys near Sierra oe ‘Us May 1901] (E--isotype, G--isotype); Orcutt 618) (E). Jeff Davis Co.: Tracy & Earle 106a (F, G). Upton Co.: Cory 664 (G). NEW MEXICO: County undetermined: C. Wright 1501 (G). Her "Parry, Bigelow, Wricht & Schott in part" is the 330, in part, cited by me below; the Wright 1498 which she cites fron "NY" is actually in the Torrey Herbarium. She says “Fine pubescence, short calyx-teeth, small leaves, and elongated profusely florif~ rp haa Copa are the distinctive characters of this species. It is bably a close relative of ve ciliata. " in all, 129 herbarium specimens, “including type material of all the names involved, have been examined by me. Citations: TEXAS: Brewster Co.: D. S. Correll 20699 (Id); Cory 30083 (N), 30620 (N), 31653 (N), 43930 ) (au), 53209 9 (Ca—73539, Du—~320340, Sm, “Sm, W—1946557) 3 Cottam am 10305 (Fs); Goodman & Water~ fall 457k (Ok, St); Rose-Innes & 3 & Warnock k mock 527 (Au, N), 21430 (Au); o. Es Sperry 93 (Fs), 614 (W—1699350), 625 (Om, W—1699359), 1380 (Om, W—1731035), 1734 (Om), T.1071 (Ca—88280h); Steiger 1275 (N); Tharp & Havard 9307 (Au—122606); B. H. Warnock 118 (an), 20105 (Au), 47011 (Rf), T ToS (N, N)5 Warnock & & Turner 7654 (Rf, RE); W 3 Waterfall ae (St); Wh Whitehouse 19633 (Mi, N). Cul- berson Co.: 0. Ee Si 0. E. Sperry erry T.128 (Qn, W--16792 251). El Paso Co.: Eggert son. {5 May 1901] (N-type). Glasscock Co.: Cory 42070 (au, N N). Houston Co.: Molby 722) (Au). Hudspeth Co.: Tharp & Havard 49294 (Au-~122607). Jeff Davis Co.: Cory 522h5 (N); Tracy & Earle 106a (Au, Cm, N, Tr, W—1809), LaSalle Co.: Cory 28553 (N). Montgomery Co.: Havard s.n. [Stockton, June 1881] 7 ee 225358). Pecos Co.: Cory 52155 (Sm), 53500 (N, W--1925)05), 53501 (N), 53502 (N), sen SoNe ~ [h-9-1936] (Md); Hinckley were (N)5 Lundell & Lundell 10190 0 (Ld, N, W—1887636), “10206 (La, N R. McVaugh 790 (Ar--23392h, Au—-1782h8, Du—~355362, mi "Nelson &] & Helson jelson 1,985 (W—197032); H. R. Reed 3,061 (N, N); Ripley & & rneby 1188 (Gg—~301325, N); Tharp 43-905 (Au, N), 43-806 (Au, , 43-007 (Au, N, St), 43-608 (Au, Ca—882806, Ny N, W— 1888995)2 Tharp & Havard 14,9359 (Au--122608); B. H. Warnock 46 (W— 1730930), 7692 (Rf, Rf), 13862 (Rf), 46135 (Ok, Rf, S), 6136 (Au, N), T.46 (Au, Au); Whitehouse 19536 (Ca—771833, ¥--1973687) - Pre~ Sidio Cos: Re. M McVaugh 1 7951 (Mi); Steiger 64 649 (N)» Reeves Co: Correll & Johnston 18576 (Rf); Cory 31309 (N), 52133 (Sm); = & Tracy 106 (N)3 Nelson ¢ & Nelson 1985 (Au, Du-~331360, Ka), (Au, Du-—331381) ; Roane Roane 28 (Ar—256371); E. Wall 5, in part (S ay B. H. Warnock 8593 (Rf) (Rf). Terrell Co.: Be ; He , Warnock 2100 age 50 PHYTOLOGIA Vol. 11, no, 1 & Turner 66] (Au--122575); Warnock & McBryde 14750 rien ° Upton Co.: Cory 5587 (Po-—213639). Val Verd Verde Co.: McKelv (W—1581343); B. H. Warnock 1400 [Parks & Turner 183] oie) County undetermined: Schott 8.n. {roadside between Indi San Antonio] (N). NEW MEXICO: San Miguel Co.: Me. L. piety 8. ne [Pecos Valley, Mar. 1, 1917] (Gg—31296). County undetermined: C. Wright 1498 (T), 1501. (T). MEXICO: Coammila: Lewis & Oliver 6 (Nb); Warnock & - Barkley 14853M (Au). Nuevo ee ae & ye& Frye 225a (Se—65655, W--1791003) ; Painter, Lucas, & Barkley 15281 (Au). LOCALITY OF COLLECTION UNDETERMINED: ok 34065 (Ur); Parry, Bigelow, Wright, & Schott 830, in part [Valley of the Rio Grande below Doflana] (W--56216). VERBENA RADICATA Moldenke, Phytologia 9: 18. 1963. Synonymy: Verbena radicans Gill. & Hook. in Hook., Bot. Misc. 1: 170. 1829 [not V, radicans Moench, 1794]. Verbena radicans Gill. ex R. A. Phil., “Anal. Ui Univ. C isa Bm 35: 190——191, 1870. Glandularia radicans (Gill. & Hook. ) Schnack & Covas, D Shh. Bibliography: Moench, Meth. cise 1794; Steud,, Nom. Bot,, ed. 1, 874. 1821; pee 7 Bot. Misc. 1: 170. 1829; Maund, Bot. Gard. . DC., Prodr vee Say, Hist. Fis. Chile Bot. 5: 12. 1849; R. A. Phil., re Univ. Chil. 35: 190--191. 1870; F. Co ks. i Moldenke, Alph, List ge ewe rate Suppl. ls “lo. 1947; Moldenke, own Geo ogr. Distrib, [ed. 2], 102, 107, & 199. 19h93 entaaske, Phytologia pas Be 0 (1920) and 3: 467. 1951; Acevedo de Vargas, Bol. Mus. Nac. Hist. Nat. So 25: 70. 19513 Moldenke in gprs Roy. Hort. Soc. Dict. Gard. 6: 2209 & 2212. 1951; E- J. Salisb., Ind, Suppl. 11: 101. 1. 39535 Moldenke, Résumé 122, ae 22h, 296, & L73. 19593 T. He Everett, New Illustr. Encycl. ard. 13: 21,06. 1960; Moldenke, Résumé Suppl. 2: 13. 1960; Mol- denis, a 8: 123 (ase) and 8: =. a ot oa ealdereey ésum 1. 3: hO (1962) and 7: 5—7 ldenke, teloei "oe 472 (1963), 9: eR 70, 18h, $ 130 ett Fy and 10: 18. 1964, Moldenke, Monograph of Verbena 51 Illustrations: Sanzin, Anal. Soc. Cientif. Argent. Buenos res 88: 127. 1919; Schnack & Covas, Darwiniana tt pl. LB. 19ls. Soca bier ; blades atid.” completely glabrous on Pst surfaces, the segnents ish colour when dry, and the spikes are te 5 oa ary. It is remarkably procumbent, rows out roo om the lower part of the woody stem." Schauer (187) says virtually he same: "Habitu prope accedit ad V. microphy: cs « Be sed folia majora magisque divisa glaberrima, exsiccata flavi tia," and "Vix diversa a sequenti [V. tenera] -" Gay (189) aK. The species has been collected i dy playas, ground, and on montane plains at altitudes of 6000. . “3000 ens flowering in November to January. As to flower color, Sparre 1329 is said to have had blue flowers, Sparre 1035 wpieiekiions Lillo 8644 violet, and Sanzin 666 white flowers. The species is eeds are offered to the trade by the Royal Horticultural Society. Rrecett (1960) describes the plant as "a vigorous grower that forms a perfect flat carpet of greenery studded with heads of li- is The name, V. radicans, of Moench, is a synonym of and actually only a new name for V. supina L., apparently proposed by Moench merely because he did not like Linnaeus' epithet. Yet i invali- dates the similar binomial proposed by Gillies and Hooker 35 years later. The Loud., Hort. Brit. Suppl. 2: 680 (1339) Rae 2 The Mf (1834-35). entire Diiveteniian identified and often cited as V. radicans, actually gbemien Vv. berterii S Spreng. Herbarium material of V. radicata has been misidentified and 52 PHTTOLOGIA Vol. 11, no. 1 distributed in herbaria as V. santiaguensis (Covas & Schnack) Mol- denke, On the other hand, the Collector undesignated s.n. [Hs H. Loc. 1837], distributed as V. radicans, is actually V. berterii Spreng., while Carette 339 is vy. V. sulphurea var, intermedia Kuntze. m ei3, 9 herbarium specimens, a. ae material of all the names involved, have been examined Ce Citations: ARGENTINA: Catamarea: Sparre 1035 (S). Mendoza: SEES 339, in part (N); Gillies s.n. [Mendoza] poet [Herb, Osten 12811] (N, Ug). Tucum4n: Lill oa Osten 8471] (N, Ug); C. C. Olrog s.n. [P. Olrog ees toe) } Sparre 1329 (S). vee eae var, GLABRA (Hicken) Moldenke, Phytologia 9: 18h. 1963. se! onyny : Verbena radicans var, glabra Hicken, Darwiniana 1: 1923. Bibliography: Hick Darwiniana 1 - 1923; Moldenke, Knom Geogr, Distrib, Be sie {ed. 1], ih & og (1942) and [ed. 21, 107 & 199. 199; Moldenke, résumé 128 & 473. 1959; Moldenke, Phy- tologia 9: 18). 1963; Moldenke, Résumé Suppl. 7: 6. 1963. s y differs from ihe typical ies of the species in having the outer surface of the c i or almost brous h very few short scattered he ice ~teeth less wane the bractlets glabrous except for He very minu gins, and the arrthergl ve and difficult to discern. n. The calyx is to 12 mm. — and the corolla 22——-25 mm. long, glabrous on the outer surfac The type of the werkate was collected by Cristébal Marfa Hicken (no. 32) at Arroyo Calabozo, Argentina, in December, 1911. The variety is known to me only from the original deseripticns VERBENA RAMBOI Moldenke, Phytologia 3: 27-28. 1951. Bibliography: Moldenke, Phytologia 3: 27-28 & Sh. 1951; Mol- denke, Biol. Abstr. 25: 3051. 1951; G. Taylor, Ind. Kew. Suppl. 12: 149. 1959; Moldenke, Résumé 111 & 1959. rb, apparently about 30 s. tall; stems apparently cong tp 1 etween the angles much more minut ap- pressed, a on the angles white, spreading or exed; nodes not annulate; prin ern a - long; leaves decus- pres rger rib tender; impressed above, Pa min beneath; secondaries slen- 196) Moldenke, Monograph of Verbena 53 pee. about 3 per side, impressed above, prominulous beneath, not xtending directly to th e tips of the teeth or lobes, but brane h- ia early, with the branches e in e teeth et reticulation indiscernible on both surfaces; e termin- al, composed of 3 abbreviated spikes; peduncles slende ° long or less, acute e ) shor pan, florifer~ ous portion of the spikes subcylindric, 1-~2 cm. lo long-ciliate margins; calyx cylindric, about 8 mm. long, conspic- uously 5 271272 1933; Moldenke, Prelim. Alph. List Invalid Names 45. 19k 3 Molden- S, gre Distrib, Verbenac 19 & 102. ae 3 Mol- eo, [ed. denke, Alph. List Invalid Nanes 6. 1912} Moldenke, Alph. Lis Cit. 1: 53, 10h, Us, & 203 (1946) and 2: 356, 393, 10, 573, & 607. 1948; H. aris Moldenke, Pl. Life 2: ihe 1948 3 Moldenke, spi pilosulous; leaves decussat e-opposite; es prided margined; leaf-blades thin-textured, lanceolate-oblong to ovate-lanceolate or ovate, 4-9 cm. long, 2--2.5 om. wide, pth 2 the apex, sharp- ly and somewhat irregularly serrate or serrate—dentate along the margins, cuneately narrowed into the petiole at the base, strigil- 196), Moldenke, Monograph of Verbena 55 lose on both surfaces, minutely pustulate above, reticulately veined; spikes RURETOUS — pat vane apices of the stems and usually strict, 4--7.5 om. long, nes much wiaceatiaig after anthesis; bractlets ovate-lanceolate, equaling or a little shorter than the comm bac -5~-2 mm. long, meres striate, the eerie sural surfaces meeting at right angles, oth. practically smoo The type of this species was collected by Friedrich Heinrich Alexander von Humboldt and Aimé Jacques Alexandre Bonpland (no. 4066) between Pachuca and Cerro Ventoso, Hidalgo, Mexico, and is deposited in the Humboldt Herbarium at the Muséum National d'His- toire Naturelle at Paris. The species has been collected in llanos, along mountain streams, in moist clay agen, in brown clayish volcanic or dark soil in’ pine forests, and along grassy roadsides in open pine forests, at altitudes of Voge to 10,000 feet, flowering in July Bee jonh iw. t Hinton states that the plant is medicinal. The only stlmoh name recorded for it is “verbena Waljers (1845) ciansttise it in his Section Verbenaca, nmnaastied Inermes, Group Foliosae, Subgroup Micranthae, and Seco ondary Su group oloivtian with with 22 other species. Sahener (1847) and Jack~ son (1895) reduce it in bin onymy under VY, carolina L. It was in- troduced into cultivation in or about 1819. It should be mah ron that the v. recta of Caesalpini is a Species of Lycopus in the Lamiaceae; "V. rect recta Dodon. and V. recta Trag. are V. officinalis L.; V. recta “canadensis etc. Moris., V. recta canadensis seu virginiana maxima, urticae foliis Moris., V. recta canadensis s. S. virginiana maxima articns foliis Moris., and V. recta canadensis urticae folio Moris. are all synonyms of Vv. LT NT ne Lee Material of ve recta has been misidentified and distributed in herbaria under the names V. carolina L., V. caroliniana L., V- hya H.B.K., or r (by Salazar) as "Vv. mexicana L., and V. polystachya H.B. stac var." and (by Nelson) as "Verbena aff. V. lystac H.BoK." Kenoyer s.n. [Pedrigal, 7-7~38] isa mixture with V. caro~ iina L., while Herb, Hort. Bot. Genev. s.n. (18 Aug. 1826] i isa mixture with V. urticifolia var. leiocarpa Perry & & Fernald. The 0. M. Clark THB, cited by me in Phytologia 8: 9) (1963) as V. caro- lina, seems actually to be V. recta. ies ollowing 8 additional specimens esi Sag oe anes as t+ as yet seen by me: MEXICO: Federal District: Pringle 13597 (G). Hidalgo: Humboldt & Bonpland 4066 56 PHY TOLOGIA Vol. 11, no» (E--photo of type). Morelos: G. L. Fisher 320 (E). Oaxaca: Pringle 4769 (D, E, F, G). Puebla: Orcutt 3950 (F). She cites the the Pringle 13596 eckieation (from Cima Station) as being from the state of México; the Pringle 769 which she cites from "NY" is actually in the Columbia University herbarium. She comments: "Verbena recta, generally determined in the material at hand as ‘aff. V. polystachya HBK.", is easily distinguished from this sloasiy related species by the rs nares ae pubescence, the h : f erally the Seatcaue calyx of V. carolina is more acutely ovoid and the subtending bract is much sharper at the apex." In all, 82 herbarium specimens and 2 mounted photographs, in- uk. phototypes of all the names involved, have been examined me. Citations: MEXICO: Federal District: Barkley, Rowell, & Web- ster 222 (Au-~1232)3, N); Bourgeau 117 (Cb); D Degener & Degener 26257 (Ny (N, W); G. L. Fisher s.n. Son. (San Angel, July 18, 192h] (Hp); Kenoy Kenoyer s.n. [Pedrigal, 7-7—38 } cee Matuda 18986 oy 3 Moldenke & Moldenke 19848 (B, Es, F, Fy, , Mg, Mr, N, ; Rs, S, Sm, me) Pefialosa 795 A Ses: * pringls 13597 Sider er Cp, Ge—h21275, Mi, RE, 5, Vt, W-H62777, W—1960738); J. N. Weaver 78h (N, ad BOR Sui Wdelaotis I. Bavis 229 (N); Humboldt & Bonpland 4066 [Macbride photos 39478] (Kr—photo of type, type, N-——photo of type); Moldenke & Moldenke 19845 (Es, N); F. Salazar sen. [Re- al del Monte, July 12, 1913] (W--1013230). México: Berlandier 1222 (W—1169),02) ; Hinton 4607 (N, W--184761); E. Lyomet 601 (N, W--16)2926) ; Matuda 21296 (N), 21301 (N), 21575 (N), 25302 (Cb), 28366 (Cb), 28407 (cb), 2 28610 (Cb); Rowell - 2050 (N); J. D. Taylor 8 (nN) (N). Morelos: G. L. Fisher 320 (W—1 20709), Sen. TAug. 192k] (Mi); Matuda 25597 (Z)3 Rose & Hay 5310 (W—395067); Wyatt 59 (Cb). Gerad Be We ’» We Nelson 1105 (W—25022h); Pringle 769 69 (Bm, Br, C, Ca—10877 eaater Io--3873, Me, Me, ae 15392, Ss, Vt, 1 2shaso, W-~1418489). Puebla: 0. M. Clark 7348 (N). Vera Cruz: Matuda 1277 ae CULTIVATED: Belgium: Herb. Martens s.n. [h. b. lov, , 1840] (Br, Br). Italy: Herb. Harvey rvey 5.D- th. R. P. 1819] (Du—166462 2). Switzerland: Herb. Hort. Bot. Genev- Son, [18 Aug. 1826] (W—26)565). LOCALITY OF COLLECTION UNDETER- MINED: Née 80, in part (Q). ‘A REGNELLIANA ent mse » Phytologia 2: h25--h26. 1948. ibatoerepliy; Moldenk , Phytologia 2: 125-126 . 1948, Moldenke, Castanea 13: 117. Tosti wordenke, Alph. List Cit. 3: Bb. 1995 mae Knowm Geogr. . Verbenac., [ed. 2], 9h & 199. 19493 E. J. Salisb., Ind. Kew. As: 11: 263. 1953; Moldenke, Résumé 111 & 473. 1959. erb; stems often decumbent and rooting at the nodes; branches 196) Moldenke, Monograph of Verbena 57 cending or erect, obtusely tetragonal, brunnescent in drying, athe slightly mar; gined, sparsely pilose or glabrescent; princi-~ pal internodes 1-708 on em, long; nodes annulate; leave dant, decussate-opposite, often with a cluster of small ones na their. axils, brunnescent in drying; petioles slender, 2--5 mm lainly winged, sparsely pilose; leaf-blades Pt ey da darker above than beneath, ovate in outline, 1.5--.5 cm. long, th terminal and also axi pa cb pe in the uppermost axils, the florifer- ous portion apparently short and dense or elongating to about cm. after anthesis; peduncles very slender, brunnescent, 3--7.5 cm. long, very sparsely pilosulous; bractlets ovate-lanceolate, about 3 mm, long and 1 m, wide, sharply attemmate or subac acumin~ ate at the apex, subglabrate pees for the ciliate margins and s calyx tubular, he type of this distinctive species was collected by Anders Fredrik Regnell (no. III.939} -- in whose honor it is named — on the swampy banks of the oes SO Rio Verde at Caldas, Minas Gerais, Brazil, on November 20, 1867, and is deposited in the herbarium of the Naturhistoriska * Rikemuseum at Stockholm. It is known thus far only fram the type collection. In barium specimens, including the type, and mounted photographs have been examined by me. Citations: BRAZIL: Minas Gerais: Regnell III.939 (F—~photo of type, N--isotype, N--photo of type, S—type, “$—isotype, Si—photo of type, Z—photo o of type). VERBENA REICHEI Acevedo de Vargas, Bol. Mus. Nac. Hist. Nat. Chile 25: 60—62, fig. 3. 1951. Bibliography: or de Vargas, Bol. Mus. Nac. His Chile 25: 60--6 ede 1951; Biol. Abstr. 26: 90h. 195h3 a ke, Phytologia 5: oe "195h; Moldenke z Résumé 122 & 473. 19593 G. 2 9 Ss Illustrations: Acevedo de Vargas, Bol, Mus. Nac. Hist. Nat. Chile 25: 62, 6 De! 1952s subshrub, branched from the base; branches ascending or decumbent, about a5 em. long, hispid, apparently sparsely foliose on the lower parts (at least when ary); leaves separate, sessile, at 4 E gat gee 58 PHYTOLOGIA Vol. 11, now 1 1—1.7 cm. long, 6—12 mm. wide, obtuse at the apex, revolute a- long the margins, attenuate at the base, 3--5-fid, pilose, the middle segment larger, broader, and 3-toothed, the pubescence longer and acclinate above, the venation hispid beneath; inflor- escence with 1—3 heads at the tips of the branches; bractlets linear—lanceolate, 3--.5 mm. long, ciliate along the margins; calyx 7--10 mm. long, slightly dilated at the upper part, hispid, the rim with subulate teeth, the larger ones to 2.5 mm. long; corolla glabrous or lightly pubescent (under a handlens), its tube 1—1.3 cm. long; upper stamens usually with exserted anther- en el herbario con V. laciniata (V. erinoices) especie que se reconoce f&cilmente por ser m&s o menos hirsuta, por los segmen- tos de sus hojas m4s largos y por los ns Bates del céliz cortos y por los apéndices d e las anteras inclusos en el tubo octet ian También presenta nrentiven semejanza con Verbena araucana Phil., pero el tipo de esta especie, conservado el Museo, es at fe) re she named the species], locality of collection, or date of col- lection. The species is know to me only from the original description. VERBENA REITZII Moldenke, Phytologia 3: 119--120, Bibliography: Moldenke, Phytologia 3: 119-~120 (sis) and 3: 289. 1950; E. J. Salisb., "Ind. Kew. Suppl, 11: 263. 19533 Angely, Fl. Paran. 7: 6. 1957; Reitz, Sellowia ll: 57 & 13h. 19535 Mol- denke, Résumé 111 & 473. 1959; Reitz, Sellowia 12: 152. 1960; Moldenke, Phytologia 10: 160. 196). Shrubby perennial, about 1 m. tall; stems and branches rather stout, acutely tetragonal, glabrate throughout; branchlets more slender, retoses 4 tetragonal, sulcate between the angles, glabrous and shiny throughout; nodes somewhat anmulate; principal inter- nodes 1—3.5 cm. long; leaves numerous, decussate-opposite, ses- sile; leaf-blades chartaceous, firm, rather uniformly bright- green on both surfaces, narrowly elliptic or oblong, 1.4-- cm. long, 2-6 mm, wide, acute at the apex, amplexicaul at the base, the upper ones entire, the lower ones rather irregularly sharp- serrate with 1—3}3 teeth per side, glabrous but very slightly scabrid above; midrib very slender, plane or impressed above, prominent beneath; secondaries 1--5, very slender, ascending, ob- scure or subimpressed above, subprominulous beneath; vein and veinlet reticulation not visible; inflorescence terminal, spicate, lly in » te bractlets lanceolate, 3—— mm. long, 1 mn. Wide at the base, reg- ularly tapering to the acute apex, glabrous; calyx tubular, ) mm. long, 5-apiculate, glabrous or microscopically strigilloses 196) Moldenke, Monograph of Verbena 59 corolla red or indigo, infundibular or 9 Aap ly its tube about l, mm. long, strigillose, the limb --5 m e type of this very distinct s species was were by Rau- lino Rotts (no. 2366) —- in whose honor it is named -- in a wet campo + Campo | dos Padres, at an altitude of 1900 meters, Santa Ca aban, Brazil, on December 16, 1948, and is deposited in the Britton Herbarium at the New York Botanical Garden. The regen c te", and "jurupeba", Reitz 3663 is marked. "typus", but is not the type col- lection -- perhaps it is a topotype. Herbarium material has been poten aes and distributed in herbaria as V. montevidensis In all, 7 herbarium specimens, including the type, have been examined me. Citations: BRAZIL: Santa Catarina: Reitz 2366 (N--type), 23h (N), 3663 (S), 372 (S); Smith & Klein 7787 (N, Ok, W--2251312). VERBENA RIBIFOLIA Walp., Repert. k: 29. 18h). Bibliography: Walp., Repert. : 29. 1844; Schau. in A. DIC., & Jac ic Ind. Kew. a tee 1 895; Briq. Bot, Gandy, be. 27, 1900; Reiche, Fl. Chile $1 285 & 286. “1310;" S anzin, Anal. Soc. Cientif. Argent. Buenos — 88: 98, 112, & Tag AAS Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 2, & 102. 1942; Moldenke, Alph. List sap i ko SUE) as 33 roe 1949; Moldenke, Phytologia 3: 75 & 76. 19493 Mold » Known Geogr. Distrib. Verbenac., [ed. 2 102, si & 199. 39193" Aceve- do de Vargas, fot Mus. Nac. in ee Chile 25: 62-~63. a Moldenke, Résumé 122, 128, & 473. 1959 4h Per ct9%, Résumé 3: 15 (1962), hi: 5 (1962 “a and 7: 5. Diffuse, b dul branchlets subtetragonal, pubescent; leaves decussate~opposite; petioles short; leaf-blades membranous, slightly grayish, oval, + 4 cm. long, 41h mm. wide, very obtuse at the apex, trifid or 3—-many-lobed, long-cuneate or cuneiform and attenuate into the petiole at the base, the middle segment pinnatifid, the lobes sub- cuneate, 2= or S-dentate or irregularly crenate-dentate, revolute along the margins, eate-rugose and rather scabrous or strigose- hirtous above, sesiely hispid or hirsute-subvillous beneath, sub- canescent, the venation prominent beneath; Spee solitary, ter- minal and’ axill short-pedunculate, many~ cr eulcapitate-ovate or subfastigiate during Pree ater becoming oblong, pubescent, reddish in drying; peduncles short; bractlets linear or linear- ar-subulate, somewhat shorter than or only one-third 60 tess OLS Ts Vol. 11, no. 1 as long as the calyx, hirtellous-pubescent; calyx about 6 mm. long, hirtellous-pubescent, the teeth lanceolate, plicate-subu- late, deciduous, the posterior one smaller; corolla rose or purple, one-third longer than to twice as long as the calyx, its tube cy- lindric, glabrous within to the slightly Led. po eg 8 ee Oe 2h, 27, 33 10, “h2, iS, 7, 53, “Se, "oh, 98-100, 102, 107, "188, 122, 135) 126, 15h, 16h, & 199. ~ 9l95 Delect. Sem. Hort. Pisa 1949: 13. 199; Moldenke, Am. Wild Fls. 293 & 450. 1949; He eae Pe Voldenke, Anal, Inst. Biol. Mex. 20: lh. 1995 oe Burpee, Walter, Seed Annual 1950: WS1, WSh, bS8, & 467. 1951; Moldenke, Inform. Mold. Set 46 Spec. fh. 19513; Moldenke, Sp. Subsp. Cont. Mold. Set.45 [3]. 1951; N. Y. Bot. Gard. Seed Exchange List sae - 1951; Stellfeld, Trib. Farmac. 19 (10): 167. vont Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 2208, 2209, & 1951; ps piece Stepp, & Walter, Everyt a for oasie 1952: 22. 1952; We Ae B ee Seeds 1952: 55, fig. 4030. 19523; Mol- enke, Phytologia “hi oe 1953; eles te Inform. Mold, Set 18 Spec. [h]. 1953 W. A. Burpee, Burpee Seeds 195i: 58, fig. 030. af 3; Rambo, Sellowia 6: 60 & 84. 1954; Moacyr do Amaral Lisboa, Aspect. Fl. Ruder. Ouro Preto 3. 1954; Thorne, Am. Midl. Nat. 52: 313. 1954; Moldenke, Journ. Calif, Hort. Soc. abs 82. 195h3 Fe Co Hoehne, Relat. Anual Inst, Bot. S. Paulo 1951: 100. 1955; Moacyr do Amaral Lisboa, Anaid V Reun. Anual Soc. Bot, Bras. 10. 1956; Moldenke, Inform. Mold. Set 51 Spec. 4. 1956; Moldenke in Hum- vert, Fl. Madag. 17h: 5 & 10. 19563 “Rambo, Sellowia 7: 260. 1956; W. Ae B e, Burpe 1956: 70, fig. 3923 Burpee § 1957: 6h & 128, fig. 3923 (1957) and 1958: 62. 19573; Ang « 7: 13. 1957; Alain in L , Fl. Gu 2 O—15, 23, 30, 47, 49, 53, 55, 61, 62, 111, 115, 118, 120, 122, 128--130 5 156, 16h, 182, 191, 214, 22h, 226, 2% 93, 310, 373, 37h, 378, 379, r; 473. 19593; Reitz, Sellowia 11: 87 & 134. 19593 Uoldenke, bs ogia 5, a rhe 1962; Park's Flower Book 1962 tz, No. Y. H +, Sect. Uy ourn 39641 120 & 12h (1961) eh es sists 267, 317, 36. A. Pu otf & Rehman, Bull. oi ‘et. Gard. en now 76: 5, fig. i. 19623 Moldenke, Résumé $ 15, 27, & 29 (1962), hs 1—3, 10, & 20 (1962), 3 Ay: "2 (i962, 6 2, zy 95 & 11 (1963), and 72 2, 3 3, 6 & 10 33 Flower Book 1963: 39. 19633 Mol denke, Phytologia "93 hi? 87, BT, 190, 296, = & 316 (1963), 9: 375 & 376 (196k), and 10: 132, 10, 964; Putz, N. Y» Herald Trib. Home & Gard. Sect. 5, pet. pos "soeht Moldenke, Résumé Suppl. 10: 7. 196. 196, Moldenke, Monograph of Verbena 67 Illustrations: Curtis, Bot. Mag. 59: pl. 3127 [in color]. 18323 Sweet, Brit. Flow. Gard. 6 [ser. 2, 3]: pl. 207 [in color]. rit. Flow. ng bed ’ ° OW. pl. 262. 1883; L. H. Bailey, Cycl. Amer. Hor ed. 1, 6: 1916. 19053; F. M. Bailey, Weeds & Polson: Pl. Queensl. fig. 245. 1906; ri & Pulney Crawford, Addisonia 8: pl. 258 [in color]. 1923; "C.", Gard. Chron., ser. 3, 76: fig. 99. 192h; Hegi, Illustr. Fl. Mittel-fur. 5 (3): 2240. 1927; Junell, Symb. Bot. Upsal. h: 19, fig. 17. 1935 soa Bienew Beitr. Pharmakog. Borag. 97, pl. 7, figs 1. 1935; Ce Argent. reek 1s 262, fig. le. i Be een : go ay oh Distr. Ibiti ° 75 [in’ color]. 19) 1950: 56, fig, “Ebs0 (1950), 1952: oe re: "i030 30 (3982), w agShe "3, fig. 4030 (195k), 1956: 70, fig. 3923 (1956) aie 64, fig. 3923. 1957; Alain in Leén & Alain, Fl. Cuba 281, wee Lewis & Oliv., Am. Journ. Bot. 118: 640, fig. Poi 1961; Nair . -« Nat. Bot. Gard. Lucknow 76: [F " Stiff, tufted, rhizomatous or stoloniferous, rabies sakees cent, perennial herb of erect-spreading habit, usually low, 15 mm see s sinple or little branched, usually 2--6.5 dm. tall, ascending or erect, decumbent or creeping at the base, a= cutely tetragonal, very scabrous-hirsutulous, scabrous-pubescent, abundantly foliose to the middle, about 3 m. in di- 10 CMe long, 1--2.5 cm cm. wide, ent on the rigs ones, more or less semi-amplexicaul or subcordate and adnate at the base on the upper ones, acute at the apex, une sub coarsely incised-serrate wi a eeth, with somewhat revolute margins on older leaves, conspicuously ve~- nose, , scabrous and more or less hispidulous on bot surfaces or callose-strigose above and rather shiny beneath and hispi on eins, lineate-rugose above, rough to the touch; mi ins ed above, prominently reticulate pecans face, the primary nes excurrent in the teeth; spikes lateral o erminal, in 3's or VIER EY: Seale, Pmt in pester short, sessile or the lateral ones pedunculate, fastigiate, usually dense-flowered org Gendasted at first, pte cylin- 68 PHYTOLOGIA Vol. 11, no. 1 ¢ and elongating to 5 cm.; bracts usually present at the base of the peduncles, foliaceous, lanceolate or lanceolate-subulate, actlet ceolate. acuminate-subulate, usually closely imbricate and longer than the calyx, often 2--3 times as long as the calyx, purple when pola hairy or glandular-pubescent, ciliate; flowers very showy d handsome, blooming from below upwards ge succession, imbricate, Pech ta crowded in heads or spikes; calyx red or green, cylindric, shorter than the subtending bractlet and nao by it, 4--6 m long, 5-angled, glandular-pubescent or hirtous, h 5, near- equal, oblique, red, acute, with short mucronulate-subulate tips; corolla rat large, mostly about 10 mn. " ft gpsabent to violet, rose-violet, aE pty al eta ghia lilac, purplish-blue, red-blue, intensely violet-blue, or blue, very lightly villosulous or subpilose-pubescent on the outside, inflated toward the middle, its tube slender, 2--l; times as long almost bifid, mostly purple segments, the mouth slightly hairy; stamens ||, inserted below the middle of the corolla-tube; fila- ments short; anthers nib is arte ee te oblate~- spheroidal, 3236 p x 35-=39 ually 34 36}, the endocolp- ium lalongate, 2.8¢x 10.5; pistil about ); mm. long; style about half as long as the corolla-tab i tate, with a spur at its base; sta oval, glabrous; fruiting- calyxes closely aieioety, dilated below the fruit; fruit separat~- ing into oblong cocci, on one of which the style sometimes re- ains, and enveloped by the persistent calyx, which is closed at its mouth; cocci trigonous, slightly broader by the base than at of dorsal nerves, margined, the apex subareolate, the commissural surfac hens esa muricately scabrous; chromosome number: x= 7, n= 21, 2n = The type of this a described Sprengelian species was col- lected by Friedrich Sellow somewhere in Rio Grande do Sul, Brazil. It was examined by Otto Kuntze, who states that it is without doubt conspecific with the common plant at that time usually know under the later name of V. a the type of which was collected by John Gillies on the pampas n r Buenos Aires, Argentina. Spren- gel's type (which may have ee: ti plant photographed by Macbride as his type photograph no. 174) is now destroyed, having been de- posited in the herbarium of the Botanisches Museum at Berlin. The type of Chamisso's V. scaberrima was collected by Sellow somewhere in "Brasiliae meridionali®, and was also deposited at Berlin. A NEW COLOR FORM OF STACHYS FROM LONG ISLAND Fred W. Oswald STACHYS HYSSOPIFOLIA f. OSWALDIAE Oswald, f ing pure white flowers instead of the normally purple- and white- marked pink flowers. The type of the form was discovered and collected by Eileen Wolde Oswald, in whose honor it is named, in a small tree sur- rounded, dried-up, and unnamed pond, 500" feet east of Middle Is- _-——— ee BOOK REVIEWS Alma L. Moldenke "Seaside Plants of the World", by Edwin A. Menninger 3 PPe, illustr. Hearthside Press, New York £70 New sol Mas 196). $9.95 This interesting work lists alphabetically according to plant types, describes, limits ecologically, and photographs (over I 4,00) more than 1500 species in 875 genera of those shoreline p victorious in varying degrees in the ge battle neta the Salt, sand, and wind associated with the sea Authentic text, well written by a highly qualified author, ex- plains the nature * of the growth problems in such habitats and the conquest of them, gives directions for planting and soil mainten- ance, and demonstrates effective conservation means for protect- ing beach areas and for reclaiming marshlands. The plants are grouped as sag ss » grass-like forms, other herbs, shrubs, trees generally, and palms specifically. Common and Scientific names are ver In a work of such scope it is difficult to avoid a few errors. On p. 20 "cause" appears in place of "course". The index lists a "Lantana" reference on p. 235, where, however, palms are pictured. The name name, "Albizzia", is spelled correctly in the index but incor rectly on pp. 171 and 172. aa" 23 should be labeled Duranta 70 PRITOLOUGIA Vol. 11, no. 1 repens var. variegata. On p. 158 Clerodendrum thomsonae Balf. f. is misspelled and accredited to the wrong author. On p. 213 the name, Vitex altissima, should be accredited to the younger Lin- naeus, not the father, and the name, Vitex parviflora A. L. Juss., should have been used instead of "V. altissima Naves ex F. Villar? There is a good index, a helpful pepe alee and an especi- ally valuable Nevgangens listing the plants according to types and grading them according to their salt ey cold tolerances. The photographs are indeed attractive. "Dynamic Aspects of Biochemistry", by Ernest Baldwin, 55h pp., illustr., ed. ). Cambridge University Press, New York 22, New York. 1963. $7.50 Because I had ence, studied the excellent previous edition of this eke finding it — in organization, explanations, diagrams, — -—- es than most texts on the market - I welcomed the appearance of nage fourth edition which is also excellent on all of pry oc In the short six year span between the printing of these two editions so much biochemical research has been accomplished the world over and such important new basic concepts aching that the text had to be considerably rewritten and augment Dr. Baldwin stil orients his materials about broad principles that x: n coenzym carbon, oxidative decarboxylation, citric acid and cognate Ses nucleic ~—— nucleoproteins, and biochemical genetics ae had t give e way for all this new material: the pili sees was lost. The continuing pace of biochemical work will probably require a fifth edition within a short time. May it be as valuable a ashe th as this onel "The Picture Book of Perennials", by Arne and Irene Nehrling, pp., illustr. Heathside Press, New York 16, New York. 1964. $5.95 The clearly written plant descriptions, the 250 good draw- ings, photographs, and color plates, the help ful techniques for al experience of the authors make this book both useful pleasurable, The evaluation of 300 questionnaires returned by 196) Moldenke, Book reviews 71 landscape architects, professional gardeners, Legon Saris: writers and teachers, nursery firms, and garden club leaders gives a practical viewpoint especially since the eet caea asked for the negative features about perennials which are naturally overlooked in most seed and stock catalogues. "Biochemistry and Physiology of Plant Immnity", by B. A. Rubin and Ye. V. Artsikhovskaya, translated from the Russian by Helen Wareing, ix +358 pp., illustr. Pergamon Press, New pe ll, New York. 1963. $1h.00 This work deals minal res rd with the intricate pez aet ships existing between plants and their pathogens, and the con- comitant physio~chanical changes in their protoplasm. It ts special features of peat as a whole, characteristic to it.” The translation sere easily. A few errors in spell our abstracting services, increases the value of this work con- siderably. "The Ecology of North America”, by Victor E. Shelford, xdi + 610 pp., illustr, University of Illinois Press, Urbana, Tlinois. 1963. $10.00 Bs tstanding work is a printed culmination of the silo ery life work of the founder of the science of and the firs scientific society for ecology. On the basis of careful field studies and on the collation of the early published writings, it peti the biotic communities in North America as they existed around 1500 A. D., before the advent of European settlement (ex- cept for the Norse) » and so serves as a comparative basis for the study of the subsequent changes in the flora and fauna wrought by our civilization, The author lists the most characteristic plant and animal spe- cies originally inhabiting this continent, along with quantita- 72 PHTTOLOGIA Vol. 11, now. 1 _ data on populations, feeding habits and other plant and ani- in employed are not always the most recently accepted ones. In the Verbenaceae and allied families the following obsolete names were noted: Avicennia nitida for A. germinans, Lippia nodiflora for Phyla nodiflora, Lippia ligustrina for Aloysia gratissima, and Verbena carnea . tor Stylodon carneus. This excellent work should certainly achieve the author's goal to present "the structure, composition, and dynamics of the inal commnities of North America before they finally suc- cumb before the advance of civilization." “Principles of Cog Sa gd Taxonomy", by P. H. Wale and V. He Heywood, + 556 pp., iliustr. D. van Nostrand, Princeton, New jog "1963. $15.00 This writing is a very thorough theoretical treatise on the subject by qualified taxonomists of Liv wespee ol and Edinburgh who are familiar not only with classical taxonomy but also with the newer approaches through genetic polyploidy, mathematics, phyto- chemistry, and machine processing. The emphasis is naturally European. It invites favorable comparison with two as si cellent and somewhat similar American works - "Plant Tax Methods and Principles", by Lyman Benson, and iptaninins of ve imal Taxonomy", by George Gaylord Simpson The main topics are: roots of taxonomy, systematics after Dar- win, units of classification, concept of characters, all types of ation of species, hybridization, and lastly an excellent conclu- sion that recapitulates the major principles and issues This book will orient the thinking of many in the field of taxonomy for several years to come, ani yet the authors know that its and all taxonomists' "decisions are subject to revision in time. I, no case is an ideal or final classification “possible, nts to be reconsidered and probably modi fied. axonomy is a series of progressive stages of knowledge." i ount Figure 14. Distribution of Verbena pumila in the United States ¢ jes are asked to gend it to th d, so that future editions of e author for verification and this map may be more complete Mapping by counties done by Andrew R. Moldenke é ] 20 21 2 I 23 a. 4 u 7 , ees 5 8 ee 9 10 * ™s . ’ », Ri WwW ‘. ®. Hq LB E ~~ c | R a OL U TA SA QD rT = ©» ® MB I : S kK A M AN Oo N $ = { TCHEWawyN TO PF 4 | ’ Lr -~— c ras oso ae s ba) mae ld Ed SreRioAn came as * em . | Ld RENVKLE | portneay sauce Trowmen | caves mae ATTSON mal . ROosz¥¢ 7 WOLLas ° s MOUNTRAIL Leseenid Unt OF a® Se + wes PERCE Tasty | was was THe woos roocncnns | A gs, ° a =~ ale BENSON Tea % mie eo? ft Le ‘elisha eed McnERIE NELSON | GRAND FORKS POLK PeRmmCTOR ki a i \ a ie Seat ay Gang | ED UKE "3 oO caeacael SHERIDAN Wels on TTASCA 7 “> = N T , = LS E re H D TRAILL ap S ORM aa etal G Paine . BiLLINGS rose Gricgs uEeAnO| 6 N | oT} BURLEIGH | WODER | STUTSMAK STEELE eM oss (e ao Custer s ot . ia cuAY mec — = mau foace AR wonton ) rr eer soe YFIEL rmeASuRE FaLon 7 oor crt a - - Sore WETTINGER CAaNT OTTER TAIL GOGEBIC rm) J ° EnaneOnes LOGAN LA MOURE RANSOM DICKINSON | DEL _— RON oe of \ @ Bowman ‘ADAMS svoux or M I N N Ss A ws” i , STAumaren 816 Wom WeinTOSH DICKEY SARGENT \ tooo Ch TG pone Jonsraunn| SAWYER | asuano . WORRISON CARTER com. — Caan o PRICE DA FLORENCE ) ame POWOER Riven ie AROING PERS a los 2 td . — @ —_sunneTT 4 ae — L/ Campset | MePHERSOM BROWN MARSHALL 7] oF a — 1 sorts FOREST . . “\ rn a a; : wie ‘SrEvERS | POPE STEARNS TSANTI A <—f anon | sk 7 a), f ~ « nar srone re Se WALWORTH TOMUNDS 17 y SHERBURNE :% i Php’ gs Pax ) Paix - zeBACH J] Dewey ‘o sTome i TAYLOR A 4 Pm Pmt ey * i 6 Wome 2 5 mem 2 my wOBeRTS 7 — wom |E —) Clan ue SHERIDAN . ae 8uTTE chinT MEEKER -{ waicHT t ma oy G =f C owen 2 om s 1 POTTER FAULK PINK ENG fast cro Swan a le 7 Son MEADE CuK | CODINGTON CHIPPEWA ar I S ARMSTI uc WeLEOD W eae Cc Oo N cc = Pr | a j JOHNSON = SULLY WrDE | HAND _ DEVEL oui PARLE ment PIERCE oe E i a ay = —- Micison eT] 7 STANLEY YELLOW Me sini = 3 ara ‘i onan ' : LYON svBLEY 7] 2 JACKSON TANTO, Le FREMONT Lawrence an H A) K/O rae ae Hy a cee x & Cy , WESTON xz Unc 9 HOT SPrings : PENRINGTON BEADLE KINGSBURY I Mose WOMROE 3 H SOMES ura BUFFALO | JERAULO | SANBORN] MINER | LAKE | MOODY re mommy | = om = o : WACKSON ousted ‘ cusTER BLUE EARTH STEELE acre” | Fomo OU LAC ns Moony POmex “AniBoy bias es naTROMA . ~ SAE AROMA tons] Taccoox | Wanna ROOK [NOBLES YAGKSON waa — | FARIBAIAT | FREEBORN | MOWER Fico aman SUNN : - s Y CONVERSE : WASHABAUGH MELLETTE Toe e — RICHLAND F NOBRARA FALL RIVE L . 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S of. ‘i “" a THIRD JUDICIAL DIVISION MEK? ro oa - Ts ad ° utay re _— j= K : Scale of Milee Pre @ te F nm : ; = ~ aia ——2 snake as i Copytight, American M i ) Ean La PS Map Co., Inc., New York, No. 14324 . 23 “ \ z 1 i 20 I 2 I 2 it ce it u 15 16 | 17 | 18 i I z I I I i i 5 I a I Pe 3 | f 10 I ii 2 3 4 I PHYTOLOGIA is financed entirely by its sega each one paying in advance for the entire cost of supe, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be cee among the eS upon the completion of a Spopaet in ee to the space which they _ Each contributor fore a shareholder in the magazine, ae his ee the apie and ste in the profits, if any accrue. : Mache number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary tly. A volume will contain about 32 signatures, 512 pages, or a smaller ee pages with an equivalent number of plates. This plan i insures a of all accepted manuscript. n: ; will be published according to the desires of the authors. No made for line drawings, such as are ordinarily reproduced in ) so-called popular r type, and sant polemics will not be ‘al Advice of manuscripts will he solicited, if iemrent ge from. i PHYTOLOGIA Designed to expedite botanical publication = EEE Vol. 11 December, 1964 No. 2 ) — le . a ze = E f MOLDENKE, H. N., ae s te Verbena. XXIV ......... Do Ria si ats a as le MOLDENKE, A. L., Boo ST TET SPECIES OF ICHNANTHUS IN SOUTH AMERICA RELATED TO I, ICHNODES (GRISEB.) HITCHC. & CHASE Jason R, Swallen There have accumulated through the years at the U. S. National father ry a number of specimens of Ichnanthus which have seemed to ielené to I, ichnodes, and have been so nam with reservations that some of them might represent distinct havi past pert er every species the fruit has small inconspicuous ites about 1/4 or 1/5 the length of the spikelets. Other characters, such as the size of the plant, the form of the panicle and the size “9 the spikelets, size of the blades, and the pubescence, make very diverse group of species. None of them can be identified with Le _tchnodes of Trinidad which is a large plant with rela- tively small spikelets. Some may turn out to be extremes of poset aa when sufficient exploration has been done and enou material is available for examination. The current study will form the basis for any future work. Sheaths densely pilose with agg pang yellowish hairs 5-8 mn, long. Spikelets 2,.8-3 mm, long.......1. I. villosissimus Sheaths glabrous or peed but ae with long = foals h hairs. Blades 10-25 cm. long. Spikelets 4.5 mm. long 2 I. amplus Spikelets not more the 3.9 mm, long. tufted, Blades, or re epee the ieogl ones, conspicuously ase Blades densely ahelpit dhgunt adel I. vestitus Blades glabrous or nearly so. I. erectus gal ejcsgenpe at the base. Sheaths papillose- I. vimineus Culms inveasent actemaie Blades with co vatioabae white ——o densely velvety 6 ndutus Blades without white margins, not ee ee 2.9-3 mm. long t glume nearly as “long as the Me, mend 3 lanceol ee of ° pallidus Spikelets 3.5-3.9 mm, isne a5 atus ercg or at least some of them, more haar os” cm. long. Pa Aste yg or clambering. 73 7% PHYTOLOGIA Vol. ll, no. 2 een rte piped sales! a, hee oa ae cm, 10 Neer pineots or nearly so; Sgepen “sortly a cm, wide, at least on the low of the plant. Blades soon below into a petioleriike base; sheaths distinctly auricula wis Blades not anlar ekes ae not ion Panicle large, as broad as long; siteetats 27: 5 mm. long 12. I. silves Panicle 2-3 times as long as broad; spikele = h mm, long 13. I, chaseae 1. ICHNANTHUS VILLOSISSIMUS Swallen, sp. nov. basi decumbentes internodiis mitis 2- em, longis; -§ mm. lo e - dense pilosae; paniculae 30-5 em. longae; spiculae 2,8-3 mm, longae; gluma prima spicula 1/2-1/3 brevior; fructus 2,4 mm longus alis 0.) mm. longis. Perennial; culms de cumbent at the base with numerous short somewhat asymmetrical, softly and densely : oatiek “with hairs about 0.5 mm, long; panicles 30-5 cm. ae , with 5 or 6 whorls of ascending branches, e lower one mach a s 20 cm. long; spikelets 2,8-3 mm, long, the first ae poy 1/2-2/3 as ometimes with hairs; second glume and ote lemma equal, blunt, smooth or somewhat scabrous; fruit 2.) mm. long, yellowish, the wings «= .0ne. the U. S, National Herbarium, No, 1255849, col- erat on steep brushy banks about 2000 ft., La Merced, Peru, aes Resse: 1929, by J. Francis Macbride (No. 5535). » & specimen collected by A. S. Hitchcock ia 22128), at Colonia Perené, Junin, Peru, undoubtedly belongs 8 species 2. TCHRAMTHUS AMPLUS Swallen, sp. n decumbentes, 1.) m, at, ” vaginae internodiis longtoner » Glabrae vel marginibus ciliatae; laminae 1-22 cm a 15-17 mm, latae, acuminatae; panicula 0 cm. longa, 25 cm, lata, ramis patentibus 13-15" em. longis; spiculae ).5 mm. longae; gluma prima acuta spicula 1/5 brevior; fructus 3 mm. longus alis 0.6 mm. longis, Perennial; culms spreading, 1./, m. long, sparsely pubescent at the nodes or glabrous; sheaths longer than the te becoming longer upward, glabrous or ath ciliate margins; ligule membranaceous, about 1 mm, long, some lacerate or ciliate; blades 14-22 cm, long, 15-17 am . wide, pers cast scabrous on the margins, and with a ew long hairs at the base; sagpted about O cm. long, 25 oft yrs the branches spreading, 196k Swallen, Species of Ichnanthus 75 longest about 13-15 cm. long, mostly naked at the base; spikelets .5 mm, long, the first glume acute, 0.5-0.8 mm, shorter, scabrous on the keel; second glume and sterile lemma equal, acute, the glume somewhat scabrous toward the summit; - long, pale lead-colored, the wings 0.6 mm, long. Type in the U. S, National Herbarium, No. 1613273, col- lected in brushy borders, Campo Salles to Crato, Ceard4, Brazil, April 13-15, 193k, by Jason R, Swallen (No, 4332). 3. ee Pin yeh Swallen, sp. n espitosi, erecti ca. yp cm, alti; vaginae dense pibeacentes, sad iaene vel glabratae; laminae 13-15 cm. pay ag a2 3.2-3.5 mm, longae; gluma prima spicula 1/3 brevior pilis paucis longis praedita; fructus 2.6 mm, longus, alis ca. 0. mn, lo ial; culms tufted, stiffly erect, about 60 cm, tall; shesthas eather closely overlap owe: the s » softly and densely pubescent or villous to glabrate; blades 13-15 em. lon & a de, t Tr ones teral sho gt er y the base, velvety isahenis ree Pow 22 cm. long, with widely spreading or ascending branches as much as 13 cm, long, densely pilose in the axils; spikelets 3,2-3.5 mm. long; first on the keel and usually with a few oe hairs; second glume 3 lected around dry ledges of Chimant4-tepui, Bolivar, Venezuela, altitude 1700 meters, nay 21, 1953, by Julian A. Steyermark (No, ene This Ss was also collected on Cerro Bolivar by Bassett Batra (No, 32671). 4. ICHNANTHUS ERECTUS Swallen, sp. Culmi erecti 70-85 cm, alti Saket laminae 16-22 cm, latae acuninatae, pilis paucis base praedita Paniculae 17-21 em, longae, -7 cm. latae, ramis adscendentibus usque ad 8 om, longis; spiculae 3,8-.2 mm. longae, glabrae, appressae; gluma prima spiculam aequans; fructus 3 mm, longus alis 0.8 mm. longis. peunuens culms tufted, erect, 70-85 cm, tall, glabrous; with al hairs and a pubescent collar; ligule about 1 mm. long, membranaceous, ciliate, the hairs about 1 mm. long; “blades 16-22 cm. long, 1 cm, wide, subattenu- ate = acuminate, gradually narrowed to the base, the lower ores smaller, with white scabrous margins and a few long hairs Hoe the base, otherwise glabrous; panicle 17-21 cm, long, 7 em, wide, with ascending branches as much as 8 cm, long; 76 PHYTOLOGIA Vol. 1, no. 2 spikelets 3.8-,2 mm, long, glabrous, the pedicels appressed; um: and sterile lemma equal, acute, similar; fruit 3 mm. long, the win ng. in the U. S, National Herbarium, No, 1613193, col- lected in brushy borders Campo Salles to Crato, Ceara, Brazil, April 13-15, 193k, by Jason R. Swallen (No. 4322). 5. ICHNANTHUS VIMINEUS Swallen, sp. nov Culmi erectl 130 cm, alti; vaginae "papi llosae; laminae 11-15 em. longae, 1-1.5 cm, latae, papillosae = ink apg hispidae; panicula 25 cm. longa ramis adscendentibus usque ad 12 em. longis; spiculae 2,8-2.9 mm. longae; sane woaaetond page suba ; fructus 2.) mm, longus alis : : longis. al; culms erect, 130 cm, tall, the nodes appressed- culm, acuminate, hispid, especially on the collar; panies about 25 cm, long, 10 cm, wide th rather stiffly oe or ches as mu much as lected on "ilhas de mata e campo misturados, beira de estr até Viana," Porto Velho, Territory of Guaparé, Brazil, May 27, 1952, by G. A. Black & E. Cordeiro (No, 52-1;560). Also col- lected at Porto Velho by E. Cordeiro & J. F. da Silva (No. 2h7). 6, ICHNANTHUS INDUTUS Swallen sp. nov Culmi graciles basi decumbentes ea 60 em. longi, papilloso-pilosi; vaginae internodiis longiores, pilosae, ried giles dense papilloso-pilosae; laminae 8-13 cm, longae 8-14 mm, latae, dense pubescentes; Sank 6fiue 12-15 cm, longae, ramis adecententibus solitairiis usque ad 6 em, longis; pa 3.5-l mm, longae; pluma prima spiculam subaequans, 5-nervi pilis paucis praedita; fructus 2,5 mm, longus alis 0.) mm lo ° al; culms slender, hard and woody, decumbent at the cm long, 8-14 mm. wide, softly and densely pubescent on both surfaces, with rather conspicuous white finely scabrous margins; panicles vei em, long, with solitary ascending branches as much as 6 cm, long; spikelets 3.5- mm. long; first glume nearly as long as the spikelet, 5-nerved, very scabrous on the keel, 1964 Swallen, Species of Ichnanthus 77 sometimes with a few hairs along the margins; second glume and sterile lemma equal, mae toward the tip, with a few scattered hairs; fruit 2.5 mm, long with well developed wings 0. mm, long. e in the U, S. National saps No. 1255815, col- lected in brush on clay slope, Campo Santo to the sea, Bafa, Bafa, Brazil, by Agnes Chase (No. 806). 7. ICHNANTHUS LUTZELBURGII Mez, Bot, Jahrb. Engler 56: 9. 1921. The following specimens have been referred to this species: Brazil: Parahyba: Pedras de Fogo, Pickel 172k. Pernambuco: Garanhuns, Chase 7792, 7832. Tapera, Pickel 1363, — — Vao de Faria, Lutzelburg 5540. Ceara: po Gran 4585, MaranhSo: Barra do Corda to Grajaht, sieliees erie; 3636. Goias: Goiabeira, Chase 11500-I. 8. ICHNANTHUS PALLIDUS Swallen, sp. nov decumbentes 80-90 cm. lo nodis pubescentibus; vaginae internodiis longiores, pilosae vel hispidae, marginibus se i Pe pilis 2 mm, longis; laminae 16-26 em. longae, 15-23 mm, latae, basi pilis lo ongis praeditae; Pos ca. 30 cm, ‘here 5 om, lata, ramis adscendentibus usq longis; spiculae 3 mm, longae; gluma prima spicule 1/6-1/5 brevior; Prectas 2, = h mm, longus alis 0 Perennial; culms decumbent, 80-90 cm. tebe the aden 15 cm, covered with much overlapping sheaths, the nodes pubescent; sheaths all longer than the internodes, variously pilose or hispid, with a prominent line of papillose hairs on the margins about 2 mm, long; blades 16-25 cm, long, 15-23 mm, wide, acumi- nate, gradually narrowed toward the base to the width of the a branches as much as 1) cm, long; spikelets 3 mm. long Reon glume 3-nerved, },/5-5/6 as long as the spikelet, scabrous 0 the keel; second glume and sterile lemma equal, blunt, nadievs toward the tip; fruit 2,.3-2.) mm. long with wings 0.5 mm, long. Type in the U. S. National Herbarium, No. —— col- lected ts patch of forest in scrub savanna, at Yupukari, basin of Rupununi River, British Guiana, October 15, 1937, by A. C. Smith (No. 2263), Le agree ACUMINATUS Swallen, Fieldiana 28: 28, 1951. following specimens are ay able to this species, although they vary somewhat from the type. The main difference is in the size of the leaves, seo may be as little as 10 cm, long and 1 em, wide, or as much as 35 cm. long and 3.5 cm. wide. zag Santander: Mesa da los Santos, Killip & Smith zuela: Bolivar: 105 km. from El Dorado to St, Elena, Foldats 271m Gcine-Catathiy Aeabiebeds Miawiah of Sate Teresita de 7 Kavanayén to Rfo Karuai, Steyermark 59371; ee San eresita de Kavanayén and base of Ptari-tepui, severe’ 60335; 78 PRIFOLOCIA Vol. 11, no. 2 between and Uaduarapara in Valley of Rio Kuken4n, Steyer- 90 Su t Bejuma road, Box 3886, 3887. Aragua: Mountains north of Maracay, e Guiana: Kaieteur Savanna, Jenman 902; Sandwith 1289, Orinduik Falls, Ireng River, Harrison 1461; Imbaimadai, — & ge ng 3226). Brazil: ar&: Campo Grande, Swallen 523. Par&: Pires & Silva 259, 10. ICHNANTHUS HISPIDUS Swallen, sp. n Cc radicantes 1 m, longi nodis RELontae vaginae inter-~ nodiis multo longiores dense papilloso-hirsutae; laminae 20-30 em. longae, 1,5-2 cm. latae, acuminatae, age pooeh Segre 9S panicula 25 cm. longa ramis usque ad 12 em, longis; spicula 3.8 mm. longae; gluma prima —- ie brevior; fructus 2. 8 mm, longus alis 0,-0.5 mn, gis. Perennial?; culms stra a 1m, or more long, white- ee at the nodes and for a short distance below; sheaths port ascending branches arranged in 4 or more whorls, as much 0-12 em, long; spikelets 3,8 mm. long, the first glume }/5 28 long to nearly as long as the spikelet, eeakeves on dh mee tip; fruit es ion mm. long, the wings small, 0.4-0.5 mm, or the U. S, National Herbarium, Nos. 2182142 and 2182113, arctan on Cerro Altamira, 10 km, east of Ciudad Piar -650 m, altitude, State of Bolivar, Venezuela October 16, =i a Bassett Maguire, John J. Wurdack & George Bunting (No. 35877). Also collected on river banks of Rio ica » Caronf, at Venezuela, by F. Cardona (No. 291). ll. ICHNANTHUS AURICULATUS Swallen, sp. nov. Culmi erecti 2,3 m, alti, ee deg§ “nodis pubescentibus; r asi petiolatae, infra et supra pilosae; panicula 55 cm, longa ramis adscendentibus vel pat usque ad 20 cm. longis; spiculae 3. ior (4.0) mm, longae; gluma prima spicula 1/3 brevior; 2.6 mm, oe alis 0.5 mm. longis. M yesatniele culms erect, 9 3 m, tall, more or less pilose with pubescent nodes; sheaths shorter than the internodes, glabrous, with prominent au ricles me to 8 mm, long, and densely villous collar; blades 28-31 cm, 1 ie em, wide, acuminate, grad narrowed to a petiole Laine ha » pilose on both sur- faces, the petiole densely hispid with ati about mm. long, the margins finely scabrous; panicle 55 cm. long, ee owly ascending to spreading branches as much as 20 cm, , pubes- Ee eee 1964, Swallen, Species of Ichnanthus 79 cent to pilose at the base, and with a few long hairs inter- mixed; spikelets 3.6-3.8 mm. long, rarely to k mm.; first glume 2/3 as long as the spikelet, scabrous on the keel near the summit, otherwise glabrous; second glume sterile lemma “iting acute, glabrous; fruit 2.6 mm, long, the wings 0.5 mn. ng. e in the U. S, National Herbarium, No, 1,88 col. lected among brush, wooded slope, between Itabira oe olatairs, from Annapolis and. Goias, Goias, Brazil, altitude March 22, 1930, by Agnes Chase (No. 11);88), Additi tional material examined: Brazil: Goias: Neco of Goiabeira, Chase 11500-II, 1150. Minas Gerais: Betwe Uberlandia and Rio Paranahyba, Chase 12107. 12, ICHNANTHUS SILVESTRIS Swallen, sp. n basi decumbentes, 180 cm. leg glabri; heat 28-32 om, longae, 18-33 mm. latae, glabrae; panicul cm, oe ramis usque ad 18 cm. longis; spiculae 3.3-3. : a longae; uma prima spicula 1/3 brevior, pilis paucis praedita; fructus 3 ee. 8 mm. longus alis 0.5 mm. longis. Perennial; culms rather coarse, decumbent at the base, 180 cm. long, glabrous; blades mostly 28-32 cm. long, 18-33 mm wide, obo " the margins scabrous; sheaths a little longer than the internodes, glabrous on the back, the margins pilose- ciliate, the collar shortly emg sea panicles 32 cm, long, aa as ——— pyramidal, the branches in ) or more pag ney 3 as much as 18 cm, a “phtose in the axils; ae a = 5 ° long; first glume triangular, about 2/3 as long as orang with a few pei hairs along the seabrous nae on the margins; second ciese and sterile 1 equal, blunt, Sessa or usually with a few scattered faire on the keel and margins; fruit 2.6-2.8 mm, long, obscurely yellowish, the oie re bf oe long. e U. S, National reap ERG: Nos, 1614126 and 1614125, poetic: in sandy forest, Japanese concession, 35 km. north of Monte Alegre, Parad, B Brazil, January 28, 193h, by 3 n Although there is some "difference in the length of the Panicle and in the general aspect, the following specimens seem no belong to this species: Brazil: Par&: Aramanahy, apajos, Swallen 3225; Obidos, "eaten 50795 Jo&o Coelho-Vigia, Black S2-UA7S; B - Arapiuns, Pires & Silva 20. Territory of Amapa: Maca capdé, Black & Froes * 110283 Rio —— down river from Porto Platén, Pires, Rodriguez & Irvine 51006. 13, TCHNANTHUS CHASEAE Swallen, Contr. U. S. Nat. Herb. 29: 71. 19h9, 5 This species is very closely allied to 1. acuminatus ie ae The first glume is longer, the spikelets are larger Pina cde and do not have the long hairs that they Sede in 2 nat These characters which seem to be enou Separate the two species now, may turn out to be only variables 80 PHYTOLOGIA Vol. 11, no. 2 of the same species when more material is available or studies are made in the fi Venezuela: Carabobo: South side of Lake Valencia, Chase 123h0. Aragua: Parque Nacional, Chase 12,63, MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXIV Harold N. Moldenks VERBENA RIGIDA S Verbena venosa f, umbrosa is based on a collection made by Cornelius | args (no. 10637) : in shady aah on rocky hills at in the same herbarium. Verbena .unusta is based on a collection made by Joseph Hicks Pyron at the old reservoir in Clarke County, Georgia, on July ), and is Se ee in dl herbarium of Duke University. I know nothing of V » parviflora except what is said of it by Hegi Cette Raat Waut ea? eatace belt Friedhof von Mannheim [1913}." The species is apparently native from central Brazil south to northern Argentina, but has been introduced and became naturali- zed in Chile, the West Indies, Mexico, Costa Rica, the southern slands > Emgland, Sweden, South Africa, India, the Pacific islands, and Australia, ertie practically throughout the year. It is widely cultiva- ted in many parts of North and South America, dies, Europe, Java, . roduced into cultivation in 1830. La " en 1839 Franc g e de t 1 {in ° Plata". Mattoon (1958) states that it is offered to the horti- cultural trade by gowgee Delbard, keperuam & Simpson Liepsen) ‘ Hillier :" Sons (Winchester), La Unwin, (New York and Chicago), Sutton & & Sons aetn). Thompson “ sais gan, Hurst & Son (London), John Forbes (Hawick), and Pearc (Moorestown, New Jersey). It is a very showy plant, especially when f large patches of brilliant purple, as I personally observed it in Bra- zil and in the southern United States. The typical form has 196) Moldenke, Monograph of Verbena 81 purple flowers — plants with a paler gerd blossom have been called var. lilacina, which see, It is not certain how many of the color variants listed above in the tatesp description as reported by yooasn een really represent different color oe and how many are merely the result of different termino age Apelgcrtan on the part of the observers. Whether there inite colors for the flowers: "purple" —~ Barclay & Purdue 7h, Hassler 2l6a, Jtrgensen 26, Lundell & Lundell 11206, Mexia 53878, Meyer 289, Q Questel 3855, Rojas 3407, Sidey 1966; "purplish" — Walther 1975 "bri. "brilliant purple" — E. C, Smith s.n.3 “vivid pur- ple*® — Clemens SNe "rich brilliant pink-purple" — Cronquist Libhs "red-purple*® —— Lundell & Lundell 10502; *rose-purple" -= Burdick s.n.3 "deep r reddish-purple" — Correll & Rollins 210613 Grtmer 90, Ha: Harris ris 11599, Hatschbach 1617, Ibarrola 3393, Legrand 1189, Rodrigue ez 545, Rojas 13118, Schwarz 1319; Wdark-violet® —— Anglade 625, Osten ten 38615 * “bright-violet® — Osten 11636; "laven- eae ee an 10646; "rose" — Sampaio 1920 io 1920, Schwarz 12h95 Secacaaee — Jorgensen 2637; "dark-rose® — - Jorgensen 264; "lilac-rose” — Herb. Inst. Bot. SXo Paulo 870; “magenta” - Erlanson 5613; "light~lavender® — McLeod | 5 Ne5 "lilac* — Ber- Tbarrola 3503 & 3069, Montes 1050, Pickel 5491, Ruiz Huidobro 4607 & 5083, Schwarz 2081, Venturi 2731; “dark-blue" — = Venturi 2731 & 9953, The species has numerous common names, as is to be expected for a plant so widely cultivated and introduced, among which are "camaradinha", "camaradinhas", "Felsheide", formosa sem dote*, “hardy garden verbena", "hardy garden vervain", "hardy verbena", Philippi (1896) says "Es propiamente planta oer merid- Schauer, Su tallo tiene 80 centimetros bs altura i un poco mas de 3 milfmetros de grueso en la base. Las hojas son en nuestro 82 PHYTOLOGIA Vol. ll, no. 2 ejemplar bastante apartadas i no 'aproximadas', las mayores, que hallan en el medio del tallo, miden 10 c entimetro s de largo i 21/2 orgs de ancho, i unos 7 dientes de fade lado, Hl pedtinc entral tiene (incluso la espiga) 6 centimetros de lar go, los aldo 10 cent{metros. La lonjitud de las bracteas es de 2 1/2 milfmetros, la del c4liz 5 a 6 milimetros, miéntras se- gun Schauer las brdcteas han de ser mas largas que el célizs la corola mide 10 milfmetros. (En la obra de Gay se ha olvidado de mencionar la)." Reiche (1910) says "A esta especie brasilefia i arjentina se refiere una sola muestra (escapada de un jardin?) que se encontré cerca de Santiago." Walpers (185) classifies the species in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Macranthae, and Sec- ondary Subgroup Melindres, with 10 other species. It has been collected in both moist and dry soil, sandy waste ground, sandy or ous soil, weedy tar open soil, formerly cultivated soil, open dry sandy soil, and black earth, at altitudes of 3 to 2300 meters. Collectors have found it in fields and grassy pastures, open places along the edge of railroads, grassy hollows cite tains wet places and meadows, low "woods and secondary for- ests, moist fields an ures sandy vacant lots and waste ditches, chalky or sandy roadsides, riversides, and railroad rights-of-way, under Eucalyptus trees, and on bare hills, grazed hilltops, aye shellmounds, low or dry ridges, rocky plateaus, pinemoate on sandy Soh)" in Tyler ee: Barclay & Purdue say along fences and roadsides" in Harris County, while por maintains "likely introduced by State Highway Dept." in Wal- ler County. Hall found it "adventive from cultivation on prairies 196), Moldenke, Monograph of Verbena 83 at Houston” in 1872, but Bebb in 1903 says “very showy in patches -—- this must be rare -—- no specimens at F.C. M. and only noted at Houston by Small." Shinmers (1958) states “rarely cul- tivated; found once as an escape on vacant lot in Dallas (more frequent in southeastern Texas). May-~June." Coker calls it "an old garden favorite in the South", Langlois found it on the "bor- ders of the Mississippi" and "rare on roadsides" in 1885 in Pla- quemines Parish, Louisiana. Between 1878 and 1880 Mohr it as "extensively naturalized", "perfectly established", and "a- round dwellings, introduced, perfectly naturalized" in Mobile Alabama. Ahles found it "forming colonies on roadsides" County Ts in South Carolina. Kuhlmann & Ktthn lr call V. rigida a nomophyte; Rambo found it in a region of 2 meters anmal rainfall, rare snows, and a temperature of wn Cos I found it abundant in dry sandy Eucal- yptus woods in Rio Grande ae Sul, Brazil. Thorne (195) reports it from "roadsides and waste places, frequent"; Jorgensen found it "common on campo" at Villarica, tent oe Bartlett describes it from McDuffie County, ere as growing on "subsoil a sandy red loam formed in situ princeton composition of the Piedmont gneisses, overlain by white sand of ae Ata formation.” In ra ntina Parodi found it palling 1k he Andropogon lateralis on the climax steppe in dept. San ped, Corrientes, along with sotabseordih, Zephyranthes, Polygala, Cuphea, Aristolochia, a, etc." Reitz describes it as "ruderal”" in Santa Catarina. is ae in ae tae ak in Saget ohm while Rosengurtt reports it as "gregarion. Uruguay. Domin (1928) stim”! sg "In Argentinien einheimisch; in Sttd~- Queensland und N. S. Wales an einzelnen Stellen vollig einge- burgert, so auch in der Mulde zwischen den Tambourine und Beech Mts." In Bermuda it is described as "frequent bern sop et Ae in ts en epa" in Corrientes erved it planted with beautiful effect in Hyde -adhegg prchigg Troncoso Ale = shee it as cultivated in Buenos Aires. Gon- Galves da Cunha ves Sobrinho cite it from Santa Maria in B tol Naturalists Soc. 1937: 261, and is called an adventive in by Douglas H, Kent in a letter to me dated November 28, Ps For those interested in plotting the spread of this species pa regions other than its native ones, I submit herewith the na ~~ Allendale in 1958, Barnwell County in 1962; Georgia -- Bibb County in 1888, McDuffie County in 1907, Clarke County in 1925, 8h PHYTOLOGIA Vol. 11, no. 2 Grady, Hancock oie Davis, Jefterso on, Monroe, Putnam, Screven, Telfair, Twiggs, Washington counties in 19623 Florida -- Walton County in 1938; * lation -- Mobile County in 1868, Sumter County in 1934; Mississippi — Lincoln County in 1882, Claiborne Count in 1925, Warren County in 1927, Forrest County in 1931, Jefferson Pik ties M 1865; Azores — 1937; Sweden —- 1930; Switzerland — 1912; New 8; queensland -- 1928. Un- doubtedly there are ELE records, but these are tue earliest noted thus by m It is worth cates here that Jtrgensen 2637 shows leaves rath- er broad and hairy. Model 298 in the Glass Flowers exhibit at the ovary, stamens, pistil, and opened corolla of this species. In 199 a specimen of ve rigida was sent to Dr. R. P. Wodehouse by a physician who stated that he had a patient with an allergy like that of poison-ivy (Toxicodendron radicans) caused by this species, Patermann (1935) reports "Zwergpollen kamen nicht zur Beobachtung" in this species. It should also be noted here that the V. nervosa of Scheele is actually Buchnera elongata Sw. in the Scrophulariaceas, Vv. rugosa Michx, is a synonym of V. stricta Vent., V. rugosa Mill, “is a doubtfully valid species (which see), Ve Tugosa Muhl. and Vv. re & Willd. are V. simplex Lelm., V. scabra Vahl is a valid spe- cies (which see) “of which v. scabra Gray ray is a synonym, and Ve scabra Muhl. is Phyla lanceolata (Michx.) Greene. In Meded. Rijks Herb, Leiden 29: 43 (1916) our plant is misidentified as Y. hispida Rufz & Pav. by Herzog. Morphological notes on V. ri S oemeniaememaemnel da are given in Svensk Bot. Tidsk, 32: 231 (1938) and by Ju- nell (1934). Hooker (1829) states that the fruit of this spe- cies is very like that of V. intermedia Gill. & Hook. In his 1832 work he says that ve rigida is allied to ¥. bonariensis L., differing in its much shorter eee and yeti goes Te flowers. He might have added that the the 1 usually is very different. Riewk in Arkiv Bot, 2 (10): ‘10-1 (90h): reduces his V, bonariensis var. reineckii what we 1964, Moldenke, Monograph of Verbena 85 call Vv. rigida, but I am not at all sure that this variety is really invalid. In Ann. Conserv. - Jard, Bot. Genév. 7-8: 291— 292 (1904) he reduces V. ri to varietal rank under VY. bonari~ ensis, following Kuntze. He states that it is olnesterand a by its 1: Targe corollas whose tube surpasses the calyx-teeth by 5--8 mm. and whose limb measures j--7 mm. in diameter. He claims that it represents a transition between typical V. bonariensis and his so-called var. brevibracteata as var. longibracteata, which I regard as V. intermedia Gill. ook. Rttmpler (1573) says of ve. ean "Brasilien. - Einjahrig, im doldenformigen, dann verlingerten, gewhnlich zu drei bein einan- der stehenden Aehren, von denen die seitlichen gestielt und Boden kultiviert, mit Glasfenstern deckt und strengen Frostes schtftzt, so treibt sie im Frtthjahre mit noch grUszere Kraft, als im ersten Jahre und die Bltthe ist dann frtth er und reicher,” In Gardening 1: 563 (1879) "J. ti writes that "This is a most useful plant, as it grows and flowers abundantly in al- most any kind of soil or pr rel ren It looks aa the brighter for drenching rains, and lasts very late in the season. When the ordinary varieties of Verbena fail, this one is sure to give satisfaction. It is easily Kept kept through the winter, and if its fleshy roots are stored thickly in boxes any number of — may be propagated in spring from the young shoots that are thrown out. It should be planted rather thickly, and cakeua down until the ground is covered, when ae til the last of the summer flower are removed or destroyed by the frost." Potz (196) states hat . 2 days are required for seed germina Haage & Schmidt (1898) say: "reisend, fttr Rabatten und Einfas sungen; tres prope pour massifs ou bordures." poconetpti cra (1934) coments that "Verbena venosa is hate mentioned by growers look- ing for plants which have proved their value. I4 has been listed in the catalogues for a long time and has been "peti with success park superintendents and the gardeners on private estates, Smateurs in this section [New York] seem to have ovasnakied it. This verbena is distinctly different from Produces great quantities of bright purple flowers all summer. It be difficult to find a better subject for bedding purposes. garden rs have been disa in this Sucbaan. tes because Sod ag teramt it as a hardy perennial, certain catalogues 86 PHYTOLOGIA Vol, ll, no. 2 having listed it among trotted plants. It is a perennial, the verbenas in Is- land, and yet it must be considered as a tender nie sign and started fron. seeds each spring, the seeds being rig Henderson (1950) lists it as among the most popular hardy per- ennial plants. He calls the flowers ‘sate tatelioe. nl 1 feet tall, blooms summers, sunny locations, any soil, useful for bor- of from 60 to 70 deg., cov seeds to a depth of only four times their size, press dom with a board f. ’ water with a fine spray, and do allow e to dry out. T as they require, if plants in pots are desired, when the seedlings have formed 2 or 3 leaves, and plant out in the garden after dan- + 2064. Park sells the seed at the rate of 25 cents per packet of “loo, us cents for 200, 1/8 ounce for $1.25, 1/h i for $2.25. Jones says "Hardy border’ plant, plant a few feet apart; 2——3 he 2 ft. tall." Chittenden calls it "a good bedding plant, the tubers of which may be kept like dahlias over winter." Bai~ —_ sa reports that it is offered to the trade by Andorra Nur~ es (Philadelphia), <— ee eres c Garden, puist, Burnett, Burpee, Carters (London), Cheltenham, rChneter? Jay Hunt, Cronamere , Dreer Po Nursery (Askov , Minnes ota), Floraire, Gardenside, Great Valley Mill Garden (Paoli, Pennsylva- & ° Haage Se on, Hocker, Hunt, Hun John Waterer Sons (Twyford, Berks.), Kaye, Le N sery Co. rt > McDonald, Michell, Mulley-Seely, Schling, ar RR ot & Walter, Sutton, Thompson-Morgan, Trivetts, and Wayside. In Journ. Roy. Hort. Soc. Lond. “sae nor” (i938) it is reported that seeds were secured from Bodger Seeds Monte, California erial of V. rigida has been misidentified and distributed in herbaria under the names V. angustifolia Michx., V. aubletia Jacq., V. bonariensis L., V Ey eeeleats eee & Rodr., Ve beasts ensis Vell., V. canadensis . (Le) Bra ton, V. caroliniana Sieg Ve: chamaedrifolia Juss., ve Eiepiae ak & Pave, ve > Ve littoralis Tenth, V. officinalis L., V. phlogiflora Cham., ve scabra a Vahl, V. Vv. teu- carneus (Medic,) Moldenke, S. carneus Moldenke, and Lippia sp. On the other hand, the Kaspiew 1169 9 distributed as V. rigida is acm tually V. bonmeientti L., Archer 182. is V. intercedens Briq., 1964, Moldenke, Monograph of Verbena 87 Bertoni 1156 & 2508, Montes 1058 & 9347, Pedersen 5205, Ruiz Hui- dobro 1607, J. Ge G. Schwarz 12h9, 1319, & 2081, and Schwindt at 271 are are V. da var. obovata . (Hayek) Moldenke, and Walther 197 7 (at least insofar as the britton Herbarium specimen is concerned) is Cochranea anchusaefolia (Poir.) Gttrke in the Heliotropiaceae. The reference in Loud., Hort. Brit. Suppl. 2: 680 (1839) is come transposed on #4 illustration and the picture of ¥, rigida is fig. 3 — fig. 2 represents lythrum roseum Marnock, a name ap~ parently overlooked by the Index Kewensis, Bauman 5, L. H. Bailey son. (May 17, 1930}, and Herb, Univ, Calif, L. A. Sen. . [Los Angeles, April 31) do not have any indica~ tion on their lab labels that they originated from cultivated mater- ial, but I am assuming that they did. Pearson (1901) cites Wilms 1176 from the Transvaal and records the species from Saint Helena. Hayek (1908) cites Campos Novaes 8.n, [Campinas, XII.1900], Wacket s.n. [prope Rio Grande inter Santos et urbem Sfo Paulo], and Wettstein & Schiffner s.n. [prope S. Anna ad flumen Tieté haud procul ab urbe SHo Paulo, 500 m. S. m.) from S%o Paulo, Brazil. Troncoso (1937) cites her nos. 375, 376, & 383 from cultivated material in Buenos Aires, argentina; Augusto (196) cites Bormmttiler s.n. [ao norte de Pindorama] and Enrich S5.n., Augusto s,n., and 3.n., and Edésio sn. from Rio Grande do Sul, Brazil; and Ahles (1958) cites ites Correll 5348 (H) from — chester County, South Carolina, He also cites Ahles 12%8, 258, & 15505, but these collections are cited by me (see bel ee les & Bell San i ons, _ The pollen-grain description for the species is provided by Nair & Pinan (1962) based on a specimen apparently cultivated at know, India -~ "NBG [National Botanic Garden] 2582; sl[ide] yet seen by me. Perry (1933) cites the following 23 additional specim @s yet seen by me: NORTH CAROLINA: Martin Co,: Randolph 688 688 (6) ALABAMA: Mobile Co.: £. W. Groves 525 (E).- LOUISIANA: East Rouge Par.: Billings 49 2 49 (G). Saint Tammany Par.: Arséne s.n. [(Covington, Aug. 1919] (W). TEXAS: Harris Co.: E. Hall 433 (E, G); E. J. Palmer 12001 (E). Walker Co.: E, J. Palmer 12036 (E). WEXICO: Vera Crus: Pur 6413 (E, F, G). BERMUDA ISLANDS: Mains Brown & Britton 153 (D); Brown, Britton, & Wortley 16h5 (D)3 Fe S. Collins L5 (| 5 (Q), . 269 (@)3 Ba Harshberger s.n. (nr. Devil's Hole, June 13, 1905) (D, E, G); Moore 296 (G). Saint David's: S. Brown 693 (D). JAMATCA: W, Harris 11969 (E, G); Perkins 102h (¢). 88 Pewee OL UOI £ Vol. ll, no. 2 She says "This South American species is commonly cultivated. It i and me hereinafter as Langlois 9 and is deposited in the Barnard Col~ lege herbarium, In all, 715 herbarium specimens and 15 mounted clippings and illustrations, including the type collections or paca ka of most of the names involved, have been examined by m Citations: NORTH CAROLINA: Cumberland Co.: inves" & Hammond 24461 (Hi--104)10). Duplin Co.: Ahles & Haesloop 28470 (Hi-- » Edgecombe Co.: Radford 36843 (Hi--104408). Forsyth Co.: P. 0. Schallert 570 (Ok). Johnston Co.: Radford 218% (Hi~ 92950). Martin Co.: E. J. Alexander s.n. (7/20/23) (Hi--59486) 5 Randolph & Randolph 688 (Ba, Vi). Northampton Co.: W. H. Rhoades 8.n. [near Jackson, 5-1935] (Bt—61222, Dp). Onslow Co.: oe & Phe S00 CAROLINA: Allendale Co.: Ahles & Bell 12581 ( piers te Re Moldenke 421 (Fg). Bamberg Co.: Ahles & Haes~ loop 22154 (a aa 9h), 25955 (Hi--104412). Barnwell Co.: A. Re Moldenk e 426 (Fg). Beanfort Co.: Ahles & Bell 12368 (Hi—-92863, St). pete Co.: Radford 21215 (Hi--92918). Colleton Co.: Ahles & Bell 15505 (Hi—92864); C. R. Bell 2292 (Hi--92855). Fair- field Co.: C. Re Re Bell 7260 (Hi—~93)00). Horry Coe: Ce ee - Bell 6220 (Hi—-929)9). Newberry Co.: C. Re Bell 7056 (Hi=-92951)> | Sal~ uda | Moldenke 3h (Fg). Bibb Co.: Eddy s.n. (Macon, May '38] (Ob-— 92283) 3 A. R R. Moldenke 367 (Fg); Fe i “He Sargent hg (We), 59 (W- 2067070). Bleckley Co.: pe Re Moldenke 374 (Fe (Fg)» Burke Coss Ae R. Moldenke 09 (Fg). Calhoun Cos: Re Fe F. Thorne 3597 alien N, Vi, W—1929863). Clarke Co.: Cronquist (N, Tl, 1927651); Holder s.n. [July 7, Se Be baguire LEO [May 3, 4925) (Gu--E.1054); Miller & Maguire 126) (Ua—16577)5 L- M. Pe 020 (N, W-~16047L5); Pyron 8.n. s.n. [old reservoir, J (#169 bs Se Je M. Reade s.n. [Athens, May May 26, 1928] (Ba, N). Clinch Cost Godfrey | 55558 (N). cr. Crisp Cos? He He Hume SMe {h mi. south of Cordele, 13 May 1937] (Fl—26628). Dodge Co: A. R. Moldenke 373 (Fg). Dooly Co.: Fattig s.n. [Vienna, 5-328] (Sa); W. He Rhoades s.n. [near Vienna, July 1929] (Bt, N). Grady Co.: | Cron- quist 5470 (Gu—29321); A. R. Moldenke 301 (Fg, S). Greene Co.: Re Be Duncan 1570 (N). Hancock Co.: A. R. Moldenke 394 (Fe). Jack- et Boyd s.n. [May 30, 1931] (Gu--£.5505). Jeff Davis Co.: 196k Moldenke, Monograph of Verbena 89 A. Re Moldenke 355 (Fg). Jefferson Co.: A. R. Moldenke 02 (Fg); P. 0. Schallert 570 (B, Hi--5Sh9l6, Je—7002). Lamar Co.: Hamlin (Gu—£.8028). Lincoln Cos: W. H. Duncan 10646 (No—21LL6). McDuffie Co.: H. H. Bartlett 1122. (Mi, Po--64522). Monroe Co.: Ae Re Moldenke 381 (Fe), 386 (Fg)» Pike Co.: W. A. Jenkins s.n. near Z 5717/36] (Ga). R. Moldenke 411 (Fg). Telfair Cos: A. Re Moldenke 357 (Fg). oe oe Co.: Biltmore Herb, 9585a (S). Twiggs Co.: Davis & Davis 11735 (Hi—TII526); 2. R. Moldenke 383 (Fg). Washington Co.: A. Re Mo Moldenke 396 (Fg ). FLORIDA: Walton Co.: E. G. Hume s.n. [Northwest paiten Co., 5/11/38] (Fl-—-29766). ALABAMA: Crenshaw Co.: C. T. Reed 2087 (Au). Mobile Co.: Crawford, »& Segars 8 1033 } (Au—122635) ; E. W. Graves 525 25 (W—98LI0) 5 W. E. 56. (We=719L9 ) 5 Co ae . Mohr gl hr 91 (Du—90886), —_— 1323138), son. (Mobile, May 1868] (W—~147593), Sen. - {Mobile, 1875] (Ka), son. , Mobile, 1876] (W~771880), sen. (Mobile, May 19, 1878) (Vt), son. (Mobile, 1878] (Pr, Vt), s.n. [Mobile, May 1880] (Du— 91138), 8. 8.n. (Mobile, April 1893] (Hi--591:66) , sen. (Mobile) (Mi, Kae Sumter Co.: Crawford & Harvill 1071 (Au--1226 33); Re Me 3186 (Ba, N, Se—-15932). Dauphin Islands S. B. Jones jones 600 (abs 210618). MISSISSIPPI: Adams Co.: A. R. Moldenke 734 (S)-_ Claiborne Co.: F. A. Cook sen. [Grand Gulf, June 15, 1925] (W— 1325892). Clarke Co.: J. D. Ray 5070 (Hi—199560, N). Forrest CO. F. He Sargent Bole ~[Hattiesburg, Sept. 16, 1931) (uN). Han- cock Co.: Denaree 31888 (Sm). Hinds Co,: Mabel Parke 27583 (Tr— 1628), Jefferson Co.? McDougall 1081 (W—1925263), 1290 (W=- 1925705) 5 Ae Be Moldenke 739 (S)- Lincoln Co.: M. B. Flint s.n. (Brookhaven, April 15, '82] (Io—92226). Pike Co.: J. D. Ray 5459 (1in199685, N). Rankin Co.: Webster & Wilbur 3352 (W—- 208807). warren Coz Ae Re Moldenke 76 (83 W. He — $.n —" (near ney July 1927; He a. Woldenke 17081) (Er, H1K0, Ec, Hs, Ml, N, Nd, St, Va), wai. ete a eine, July 1931 va N, Ob—50856), son, [National Cemetery, Vicksburg] (Hs). ARKANSAS: Craighead Co.: Demaree 2920 (Sm). LOUISIANA: Ascension Par,: Penfound 8.n. {March 24, 1937] (T1). ——e Pars: W. H. Rhoades s.n, [Athens] (Hs). East Baton Rouge Par Correll & Correll 9118 (H--7663h, N, N), 1ObL9 (H—78936, N, 5 S.W. # Le = 7882 (En); Joor s.n. [Baton R Rouge, June ll, 187k} (a, 3 A. L. Richardson 36 (Ur). East Feliciana Par.: we He Rhoades $3. {ethel, July 1931) (N, Up). Orleans Par.: D. Humel sn. [17/1/1958] (S). Plaquemines Par.: Langlois 9 (Be), sat (21.VII.1879] (I), s.n. [June 1880] (Ca—192902), s.n. [July (Al), s.n. [Pointe a la Hatche, 21.VII.1885] (Mi, N, Pa)3 90 PAT.-T 0.1.0:6-5:4 Vol. lL, no. 2 H. R. Reed s.n. [Belle Chasse, 1 May 1933] (W~-1600285). Saint Mary Par.: Correll & Correll 9316 (H—-78935); G. L. Fisher 35011 bp s.n. (Morgan City, Apr. 21, 1935] (Gg—-267618, Hp); Lambert ert s.n. [September 1, 1938] (Up); Perkins & Hall 2634 263 (Po- ae Um--21); W. H. Rhoades s.n. [Paterson, Aug. 19%] (B (Bt, W)- Saint Tammany Par.: Arsene T1080 (#--103157h), act a (W—-1033055); Bomhard s.n. (Mar. 25, 1927] (Cm); Cocks s,n, (Slidell 1901] (TL), son. [May 10, 1901] (11); T. F. Hall s.n. [Apri 25, 1939] (Tl); A. Re Moldenke 22 (Fg); F. W. Pennell I 20, (Mi, N, Up—62326). Terrebonne Par.: Arceneaux 25 (It). Washington Par.: H. P. Riley s.n. (May 9, 1937] (Se--7523) « West Feliciana Par.: Een 3 3.0. (31.3.1951] (Ss); Ewan 187), (T1); Penfound s.n. [April hk, 1936) (11); F. W. Pennell Il 4332 (N (N, Up-62322). Parish unde- : Dormon | Sone . [South La., April 1930] (N). TEXAS: Brazor- ia Co.: H. B. Parks sn. [Cory 29582] (N); THATP 820s sen. [7/2/39] (Au, Av, A Au, ,» Ma, N, Sm). Cherokee Co.: H. Je H by 1764 251841). Fort Bend Co.: Tharp 253 ple Sen. [7/2/39] (Au). Gal- veston Co.: Mally s.n. [Hulen, Sept. 2 2, 1895) (W--227686)3 Mrs. Ae Ae Ee Nelson s.n. []-20-)2] (Au, WN, itt E. Ce saith 5M. (Dick on Bayou, 5-2-1942] (Fe). Harris Co.: G. C. Albe: (oe); Barclay & Perdue 7h (1d); R. Bebb 120 (Ok, Gt Ge Le Fisher W--503510), “625 (W—-503560) , 51001 (co, S), 5eR. 460 en REE [May 28, 1913] (vi), Son. 1. [Hosuton, Sept. 1s, 1913] (Hp), 52m. [Houston, Apr. 11, 1930) | eee. Ew, Qg-—22251h), son. (Sept. 10, 1932] (Du--2301,50, I, Ms), sen. {Hosuton, May 5, 197] (B, Go) 3 E. Hall 433 (N, Pa, Po--71137, W—227686), sen. [Hosuton, 1872] (N); E. J. Palmer 12001 (Ca—L2560k); Reid 8745 (Au); Small & Wherry 11613 11813 ( (N, |) W—1738007); Thurow s.n. [Houston, Sept. oe 1915] (W—865599), s 8.n, (Houston Heights, April 28th, 1923] (Hu); Warner s.n, [near Houston, May 20, 'hO] (Hu). Jefferson Co.: J Cory 11282 (Tr). Lee Co.: J. F. Brenckle 148077 (N). Montgomery Cos: McLeod s.n. [near Conroe, May 15, 1960] (Au--18048). gg ange Co.: M. S. ¥ 662 (Au, Au, Po—161331, W-~110)638, rae s.n. [Orange] (1o—-104883). San Angustine Cos: Co D. Smith sae 26 (Nb).» San Jacinto Co.: Gould & Reaves 8230 (Au~-199210). Shelby Co.: Cory 56512 (W—2007675)3 Lundell & Lundell 10502 (Ld, N, W—1887685). Travis Co.: Herb. Univ. Texas s.n, (Austin, 5/ 2/35) (Au); Tharp s.n, (Austin, 5/2/35) (Au, Au, Bt~-—25),23, St—9281) - Tyler cee Pa Turner, & Johnston 544841 (Au—122628, St). Wal- ~ Palmer 12038 (Ca—l25605); Parks & Cory 2035 (Tr), 7880 =, = at (Tr). Waller Co.: Correll & Rollins ae. Ga Cory iors, Rf, St). County undetermined: C. C. ar ay = aan bers 8 (nn, Au). CALIFORNIA: Sonoma Co.: M.S 196) Moldenke, Monograph of Verbena 91 (Gg--2182)2). MEXICO: Vera Cruz: Purpus 6113 (Bm, Ca~—169),91, W--566990), 15705 (Ca-l64353), sen. [Zacuapan, May 1919] eke 200693). COSTA RICA: Cartago: H. , Pittier 2929 (Br). BERMUDA Is- IANDS: Main: S. Brom 153 (N)3 Brow, Britton, & Wortley 1645 (N); Fe. S. Collins 2 269 (N, (N, W—717563)3 O« QO. Degener 1301 1301 (N), 8 Ben. n, (July 2h, 1921] (Ms); B Harshberger s.n. (nr. Devil's Hole, June 13, 1905] (Up—43130, W--8]7617); McCallan s.n. {Agric. Experiment Station, July 4, 1921] (Ba), sen. [July fake Ade Reet Saint David's: S. Brown 693 (N, W--13)105)). Saint George's: W. R. Taylor ,9=1189 (Mi). CUBA: Oriente: Clément 662h (W--1959956); Hxman _ (S), 8769 (8); Herb. Estac. Exp. . 2 15776 (Es); m & Maurel 2520 (N)5 Leén - ‘3910 (Ha, N); Lépez = 2615 (W—2 egg a CAs WG. L. Britton tton 3176 (N)3 Chrysler 1101 i (Ru); » Harris 11969 (Gg—31389, N, 8, W—791016), ot Jy Bane (27107922) C thi); We W. R. N). LEEWARD ISLANDS: Guadaloupe: Duss “¥ Philipson 689 ( : Duss N); Questel 3 3855 (N). WINDWARD scenes Martiniqe: Duss 1697 (N, N). BRAZIL: Minas Gerais: Campos Porto 1839 [Herb. Rio de Jan, og (B); Mello Mattos s.n. [1tabira do Campo, June 1902; Herb. Rio de Jan, 46762) (N); Mexia 5387a (N). Parané: Dusén 3638 (Ja—h6571, N, S), 7786 (N, S), 8772 (S), 8825 (N, 3, W— 181762), 13329 (s), 15793 (S, W—1h81767), s.n. [Herb. Rio de Jan. 3638] (N); Hatechbach 1617 (N); Mello Mattos 702 (N); E. A. Moreira 60 (W—-2369335); S Stelifeld 28 (Herb. Mus. Paran, 1397) (N), 1397 (S); Tessmann 2788 (S), | (S), 8m. [Herb. Mus. Paran. 2788 gece eiro: Herb. Be Bae Ge sane Dek 005 & ampaio 1920 5h), 2351 ‘ates. Rio Grande do Sul: Jurgens 223 (B), 45. (B) 7 5 ta), sen. (B)3 Leite 246 (N); Lindman 4.473 shy 1/2 (N,S, 8); Malme 50h (S), 50a (S), 10u6 0s tise Vol- Genke & waste Sy ( (B, pain _ — , 1g, Im, Mg, Mr, N, No, Ot, Rs, S, Sm, 990 (Sp--50992), 27143 (N), (N), 27286 (N), ones “Ot aes a 75), ar , S), 51646 (W--2102123), ak 21647 (s), 572u8 (S); Reineck & Czermak 68 (Herb. Osten 1158] (x, S, Ug); A. Re Sclmlte 09 (N)y Schwacke 2759 (Herb. Saldanha 5070] (Ja--L6580); Sehnem 3505 (B, Gg——-356u11, N); J. Vidal s.n, (Allemoa, March 1939] (Ja—W6558), son. [Santa Maria do Boca do Monte, March 1939] (Ja~h65u7, Ja—Lé519) . Santa Catarina: Dusén c.as7 (N, S); Schwacke 183 fuels. ie Mus. Nac. Rio ref. no. rr} (Ja); L. B. smith 5610 (W—2120177, 2); Smith & Klein 7488 (N, Ok, W—-225126],); Smith & Reits 10200 (W-22h9368). So Paulo: Araujo h (s Sp--2000) 5 Brad Brade 5742 (N, S, Sp—67Ab) » 5769 (S)s Can- Pos Novaes 918 (W—-389580); Camp Campos Porto 2983 (Herb. Rio de Jan. ] (B, )5 are Bit seen Peet Agron. S. » aul 8150} (Be—37287) ); Edwall s. - 18, 1893; Herb. Com. Geogr. & Geol. 653] Tis ts67335 ecb Jara. erage Rio de Jan. er a eee 92 PHYTOLOGIA Vol. ll, no. 2 1406 (N); F.C, Hoehne s.n, (Butantan, Nov. 7, 1917] (N, Sp-—-870), BMe [Itapira, May 14, 1927] (Sp--2030)), s.n, (Serra Negra, June I, 1927] (Sp—206hh), s son. (Serra Negra, June une 3, 1927] (N, Sp— 6)3 Hoehne & Gehrt s.n s.n. [Caraguatatuba, Dec. 8, 1939] (N, Sp- 41852); Kiehl & Castro s.n. (Herb. Inst. Agron. S. *Pmlo 3777) (4N, W—-1775580) 3 Kiehl & Fr Franco s.n, (Herb. Inst. Agron. S. Paulo 5161) (N, Sp--l44300) ; Me Kuhlmann 77 77 (Sp~h7879), 481 (N); Leite 3726 (El); B. Luts s.n. rea Bocaina, Jan. 1925] (Ja--15093)5 Tuederwaldt s.n. Ipiranga; - Mus, Pa 92) (N, Sp~ 5716); Pickel 1760 (N, sf), "1678 (Sf), 5165 (Sp, Sp—hh9h7), S. Paulo 392] (Ba, N, W-—1593220), s.n. [Herb. Inst. Bot. S. Pau- lo 735) (#1591593); Usteri s.n. (Capital, 1905; Herb. Polytech. S. Paulo 247] (N, Sp—I5711); Viegas & Zagatto s.n. [Jaguari, 10 Jan. 19393 Herd. Inst. Bot. S. Palo 3792] (#1775883) « State undetermined: Sellow 1,28 (Vt), s.n, (Bras. merid.; Macbride pho- tos 17h] (Br, F—photo, Kr—photo, N, N—photo, N—photo, N— photo, N--photo, Si-—photo, Vt, Z--photo, z-~photo), S.D. tBra- silia] (N). BOLIVIA: Chuquisaca: Troll 53 (B). Santa Cruz: Car denas 5192 (W--2250729); T. Herzog , 1696 (S)3 Steinbach 6052 (Herb. Inst. Miguel Lillo 38012] (N). PARAGUAY: Fiebrig 6305 rig 6305 (W--1134887); Grosse & Lindman 3899 (Ja—28233); Hassler 1560 (N), N), 6508 (N, S)3 aoe 3770, in part (N, Ss, pri a leta s.n, (Herb. Osten 3747) (Ug); H. He caer 21008 (Mi); Berro 2362 (N), 5535 (N), 7856 (N)} Cantera 34 (Ug, Ug, Ug); Cas- tellanos s.n. (Playa Atlantida, Dec. 29, 1945; Herb. Inst. Miguel Lillo 15164] (N, N), son. [arroyo ¢ Catalancito, Jan. 30, 1948; Herb. Inst. Miguel Lillo 15049] (N); Collector undesignated s.n. (Marzo 10 de 1885] (Ug), s.n. (Ug, Ug); Hauman s.n. (Montevideo, X1/1922] (Br); Herter 268e [Hi {Herb. Herter 80738) (He), son. [Herb. Osten 17051] (Ug); Kuntze s.n. [Sierra de Solis, Nov. 1892] (N, N, N); Legrand 3492 (Ug), 1189 (Ug); Lombardo bhe (N); Miler & Melchers s.n, [Harb. Inst. Miguel Lillo 11762] (N); Osorio B-De {Valle Eden, Feb. 19, 197] (N, yar ert Osten 386L (Ug), 5302 (W-—113490h), 11636 (Ug), 11637 (Ug), 160h9 (Ug); Rose: 1920 (N, N, Ug——5609), B.2705 (N), B.2069 (N), B.3035 (N), B.5301 (20 48512), sen. [He Ne Moldenke 11203] (N); Schroder BDe [Herb. Osten 19)5h] (Ug); Tappen s.n. (Herb. Osten 5302] (Ug)- ARGENTINA: Buenos Aires: Gillies s.n. [Macbride photos 3)3h9] 196k Moldenke, Monograph of Verbena 93 (Kr-photo, N--photo); Parodi 7382 [Herb. Osten 20939] (Ug). Cha~ co: Jtrgensen 2/64 (Herb. Inst. ;. Miguel Lillo 31553; Herb. Osten 11869] (N, Ug); T. T, Meyer 289 (Ug——10941); A. G. Schulz 1480 (N); Venturi 5 (W--10)3590). Cérdoba: A. T. Hunziker 7134 (N). Cor rientes: Ibarrola 3393 (N), 3505 (N), 388 (N); Parodi 12623 (N). Entre Rios: T. Meyer er 10097 (N). Formosa: . Jorgens en 2637 [He [Herbd. Osten ieee co) Oe) Misiones: Bertoni 2008 page e » 2961 (N, 1,067 (N), eee (W—1858339); Ekman 1981 (S), 1962 ee N, a3) 2028 (S); G 90 [Herb. Osten 23177] (Ug); Iilliesktld. co [vicinity =a Siceia Ba); Stearn 8. {ioldenke & Moldenke 9157] (N).- France: Laplace 5388 (La), S.n. n. [Duffour 3888] (Vi). Germany: Collector undesig- nated 2109 )3 “He. G." 26 (Bm); Herb, Martius s.n. gen. [h. Monac. pie 1), sen. (Br) Herd. Prager 16641 (Gg—3h25); C. J. Mey- » duli 1897) 0B); | wtthibach Bon. (Potsdam, “7 Aug ar Fil (B). Hawaiian Islands: C. He Hitchcock s.n. {Kauai} (Dt). Illinois: E. E. Green s.n. [Garfield Park, July 2h, 1935] iuiiee « aecmtsmencancier eniemcandam ucaerare lanponton Bes Ss oe Ss Be (Bz, Bz--26h30), li (Bz, Bz--26)38). Maryland: W. H. Cowgill 563 te. 6. Introd, 121505] (Oa-~9236); McCann s.n. (9-1 (Md) « Massachusetts: L. H. Bailey s.n. (Cambridge, Aug. 31, 1921] (Ba), Son. (Cambridge, | August 6—-12, 1929] (Ba); Leavitt oe [Cam- bridge, 11 Oct. 1898] (Rf, Rf). Michigan: Baer 1 New Jersey: H. N. Moldenke 9289 (N). New York: L. H [Sept. 25, 1922] (Ba), son. [C. U. greenhouses, “lay 1, Oy) 23) (Ba), sen. [C. U. gardens, Aug. 1h, 1925] (Ba), son. [Ithaca, Oct. 10, 193k] (I), sen. [Vilmorin-andrieux & Cie. seed 80080] (Ba); Bas- tedo s.n. [21.S.1897] (N); Burdick s.n. [Sept. 26, 1922] (Ba)3_ Kalergi sen. (7/2h/h2] (N); H. N. Moldenke 807) (N), 10611 (N)5 Moldenke & Moldenke 11891 (N); Nanton ea; ton 8. [9/3/40] (N)3 Ge Vo Nash s.n. (N. Y. Bot. Gard. Cult. Pl. 4553) (N), Sen. (22.5. 1898] (N)5 R. 5 Se Schneider s.n. (N.Y. Bot. Gard. Cult. Pl. e * Ne 7i9h8). Spain: Herb. Hort. Matrit. AGF Sweden: Alm s.n. [15- VIII.1947] (Bm); Bauman 5 (Ew); Blom s.n. [1931] (Go); Herb. Mus. Bot. Stockholm a fi8uL) (s)5 Sepa ack, sen. [7/1897] (3)3 REO terman s.n. [13/9/1886] (Go); Trolander s.n. [Juli ego (Bs Ee Ee Wall 5 [1878h)) (om), 5 [26/840] (Ew), (Ew), sn. (2/70) swit- zerland: Herb. Hort. Basil. s.n. [Jun. 1030] (Mi), sae Pa 1839] 196), Moldenke, Monograph of Verbena 95 (M). Texas: E. Hall son. (W—71978). LOCALITY OF COLLECTION UN- DETERMINED: Collector undesignsted 49 (Q); Dunlop s.n. (aug. (Pa); Miers s.n. Ben. (Zanjon, Desmochadez] ( Bm); Tweedie a oe MOUNTED ILLUSTRATIONS: Gard, Chron., ser. 3, 76: 279, 192 (Ba); G. V. Nash s.n. [N. Y. Bot. Gard. Cult. Pl. *Hi3533 “(N)5 ie pee (N). MOUNTED CLIPPINGS: Gard. Chron., ser. 3, 9. RIGIDA var. ALBA Mange Moldenke in Chittenden, Roy. Hort, Soc, Dict. Gard. 6: 2 Sl. : Verbena venosa ilies Diets: Cat. Suppl. 1933 Nov= elty List A 19334 Moldenke, “Aiph. List Invalid Names Suppl. 1: a tant, in syn. 1947; Moldenke, Résumé 378, in syn, Bibliography: Trivetts, Cat. Suppl. 1933 Novelty List 6. 1933; Le “i Bailey, Cat. Florists Handl. Verbenac., mss, 1935; Moldenke, Alph. List ike pera Names 7 oe lz: 27. 19473 ne in Chit ttenden, Roy. « Soc. Dict. Gard. 6: 9 ke, Journ, Calif. hyp Soc. 15: 80. 195h3 Moldenke, adr 22h, Journ 378, & 473. 1959. Haec varietas a forma typica speciei corollis albis recedit. This Reotauen differs from the typical form of the species in Bailey (1935) lists this as only offered to the horticultural trade - ree , but I am not certain that the reference given above i ually the original publication by Trivetts. In my 1947 eA Saee my I accidentally Seon the author name as "Triretts" and reduced the trinomial to synonymy under V. rigida As yet I have not seen any material which was definitely this variety, Rags eae var, GLANDULIFERA Moldenke, Phytologia 6: 331. Synonymy: Ver See See omens ene oe ae Be Clark, Card Index Gray Herb. issue 229. 1958. Bibliography: Moldenke, Phytologia 6: 3h ST ten - Brig. sum Card. Index Gray Herb, issue i 1958; Moldenke 473. 1959; angely, Fl. Paran. 16 79 (4960) and 17: 7. 196; Moldenke, Résumé Suppl. 7: 9. 196 Ai S variety differs from the typical ey of the species in having the peduncles, rachis, bractlets, and calyxes densely -soor at troche a d the bractlets only very shortly and Sparsely ciliolat The type of the "variety was collected by Gert Hatschbach (no. kata) Ae: the roadside at Laranjeiras do Sul, in the municipal- ay of Guarapuava, Paranf, Brazil, on November 15, 1957, and is ceposived in the Moldenke ning at eee New York. Only 2 erbarium specimens the type, have been examined by me. 96 PHYTOLOGIA Vol. 11, no. 2 sitettemes BRAZIL: Parand: Hatschbach },212 (Z--type), 4213 (Ok). VERBENA RI - LILACINA (Benary & Bodger) Moldenke, Phytolo- gia Ss 33. "1988. Synonymy: Verbena venosa lilacina Benary & Bodger ex Harrow, Journ, Roy, Hort. Soc. Lond. 61: G01. 1935. Verbena rigida var. lilacina Hort. ex Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1), me ees 1942. Verbena venosa lilacina Dreer ex Moldenke, % Invalid Names Suppl. 1: 27, in syn. 42875. Pearce, 196 Garden Aristocrats 20. 1964. Verbena rigids lilacina Moldenke Chittenden, Roy. Hort. Soc. Dict, Gard, 6: 2209. 1951. Verbena ida lilacina Hort. ex Moldenke, Résumé Hoag 3: 40, in syn. 26 Bibliography: Purdy, Perenn. Pl, Fall 1928 Spr. 1929, p. 53-6 5 343 Harrow & Hy ee Verb Moldenke, Known Geogr. Dis- trib. Verbenac., [ed. 2), are & 102. 1942; Moldenke, Alph. List Invalid Names Suppl. 1: 27. 1947; Moldenke, Known Geogr.-Distrib. Verbenac., [ed. 2], 161, : 199. 19195 Moldenke in Conntyndens Ps . ache: Sec. Dict . Gard, 6: 2209 & 2212. 19513 Moldenke, . Hort. Soc. 15: 80. 1954; Moldenke, Phytologia Ss 3. oo; “Woldenke, Biol, Abstr. 30: 1092. 1956; Moldenke, Résumé 22h, 373, an & 173. 1959; Te H. Everett, New Illustr. Encycl. Gard. 13: pl. 1i-1la. 1960; Moldenke, Résumé Suppl. 3: 29 & h0 (1962) po 7 9. 1963; Pearce, New Highlights in Fls. 20. 1963; Pearce, 196) Garden Aristocrats 20. 1 astra tions: T. H. Everett, New Tllustr. Encycl. Gard. 133 pl. li-lla yoga color]. 198. This variety is said to differ from the typical form of the been collected in tbat in June in California. It is not at all certain how much variation there actually is in the color fran’ the flower of this horticultural variety, and how the ida Spreng. actually belong in this variety and that some of the ‘apparent color variations listed there actually belong here The first description of the variety knom to me is the one by Purdy (1928) who has this to 5 of it: “hardy, — quite a ieee cold. It args by underground runners to make masses: he flowers are and quite pretty.” He pect a 2 cents per plas, 52.6066 per dozen plants. Farrington (1934) is the first m know as yet to me to have published the varietal epithet, 1964, Moldenke, Monograph of Verbena 97 but this was done in a newspaper column and therefore is of doubt- ful nomenclatural standing. He says of the csi "A variety of this verbena which is somewhat new is called Venosa lilacina and will be set ete for its color, a very delicate auie of lavender blue. Like the original venosa, it is good for beds and borders, and even for the rock g ° In my 1942 work I used the name "Verbena rigida var, lilacina Hort." before I had determined the basinym. Even now, it is not at all certain to me that the variety should be credited to Benary & Bodger in the 1935 reference given above. It may very well have been named and sp orci a er by these authors elsewhere, or by someone else Harrow (1935) jays on ite ‘characters of V. Venosa, but flowers very pale lavender—lilac", and t se ere obtained fran Benary & Bodger. Bailey (1935) ste. ve it is offered by Dreer be by Schling. Pearce (1963) calls it a verbena "of merit...... grown as snnala.... perennial in mild tage Seabee rah showy long~bloomer, near to sky-blue. pgpaioling 11 winter-store" and sells it for 26 eects per pooner 2 seeds. ti pecimens d cantons zo this variety have en her’ examined by me pag or pe CULTIVATED: California: Bracelin 2113 (Gg—-3h772h, “9 RIGIDA var, OBOVATA (Hayek) Moldenke, Rev. Sudam. Bot. 5: - 1937. Synonymy: Verbena rigida f. obovata Hayek in Engl., Bot. Jahrb. 42: 162. 1908, Bibliography: Hayek in Engl., _ yahib 42: 162. 1908; Mol- denke, Rev, Sudam. Bot. 5: 2. 1937; eogr. Dis- trib, (Werbenac., 1}, le& 102", shes Moldenke, Alph. {ed s Suppl. Re 26. 19h73 Moldenke, Alph. List cit. ls 26h, 19hisy a Moldenke, Phytologia 2: 337. 19li73 Moldenke 13 119, igh; Moldenke, Alps. List Git. 32 761 & 782 (1949) and 33 oo 19495 Moldenke, Known Geogr. Distrib, Verbenac., [ed. 2], 99, 100, 107, & 199. 19h9; los woldenke, ‘Résumé 112, 8, 12, 128, This variety hinrere’¢ raat the typical form of the " species in The type of this variety was collected by Emil Hassler (no. oa) in the vicinity of Caaguazu, Paraguay. The variety has as been found in m nedium~dry to moist generally gered grasslands, on cam- ong a Gill. & Hook. The variety is not we marked and oes sossible that some of the specimens cited by me 98 PHYTOLOGIA Vol. 11, no. 2 under typical 25 rigida, especially those from Paraguay and Mis- iones, actually belong here. In all, 30 herbarium specimens, including the type collections of all the — involved, and 2 mounted photographs have been examined by Citetionas “BRAZIL: Santa Catarina: Smith & Klein 11968 (W~ 2251838, Z). PARAGUAY: Hassler 6508 (Ca—9l)359), 8911 (N—iso- type, N--photo of isotype, e, V--isotype, Z—-photo of isotype); Ped- ersen 5205 (S). URUGUAY: Osten 6569 (Ug), son. (near Piriapolis, Warch 30, 1911] (Ug). ARGENTINA: Misiones: Bertoni 1156 (N), 1513 (Ca--164687), 2508 (N); Ekman 1973 (Mi, N, S)3 T. Meyer 6659 (It 11508b); Montes 1058 (N, N), 240k (N), 9347 (Au--122909, Iu, Lu); J. G. Schwarz 12h9 (N, S, S, $), 1319 (N), 2001 (N); Schwindt 271 (N)- ee hee San Mateo Island: Rufz Huidobro 4607 (Au--122636, N, Ok). Mair ie var. REINECKII (Briq.) Moldenke, Phytologia 2: 150. 9 : Verbena venosa var. reineckii Briq., Ann. Conserv. & Jara. Bot Bot. Gen Genév. 3: lok. 1899. Verbena bonariensis var, rein- eckii Briq., Arkiv Bot. 2 (10): 10, in syn. 190]. Verbena rigida var. latifolia Hassler ex = Holden, is oe in syn. 1959. Bibliography: Briq., ard. — Genév. 3: 16h. 1899; Briq., Arkiv Bot. 2 2 (ioye 2 10-1. 190k; i idenke, Phytolo- gia 2: 150. 1946; Moldenke, Alph. List Invalid ani Suppl. 1: 22 & 28. 19173 H. Ne & Ae Le ae re Et 2: — 19483 Molden~ olde b. Verbenac., [ed. 2], 99 & 199. 1995 1 usidenke logia 3: 45h. 1951; Moldenke, Résumé 111, 118, 359, 373, 378, 273. 195995 Moldenke, Résumé Suppl. 3: 15. 1962. variety differs from the typical form of the species in having re leaves much thinner in texture, much les — re broadly elliptic in shape, and less scabrous above, os eit ce much less prominent beneath. The plant is described by oer as suffrutescent, 0.8--1 m. tall, with blue-purple flows: The type of the variety was sored by Eduard Martin Reineck and Josef Czermak (no. 55) at Belém Velho, Rio Grande do Sul, Brazil, on October 31, 1897, and is deposited in the pomnecs Seertak tien at the Conservatoire et Jardin Botaniques at Geneva. I have not as yet seen the type, but the several semester cited below apparently represent the same taxon. The type of var. lati- folia was collected by Teodoro Rojas (no. 3365) a. the edge oft woods at the foot of Cerro Acahy, Carapegué, Paraguay, in April, 1919, and is deposited in the herbarium of the Museo de Historia Natural at Montevideo. Reineck & Czermak 68 [Herb. Osten 158] in the Montevideo herbarium is ; is inscribed "Verbena venosa vare reineckii var. nov.", but appears to represent the typical form of the species. The variety has been found in swamps, on hillsides, and at the 1964 Moldenke, Monograph of Verbena 99 edge of woods, flowering in April, Material has been misidenti- fied and distributed in herbaria under the name V. yenosa Gill. & Briquet (190k) cites the Ann. Conserv, & Jard. Bot. Rew: 3: 16h reference as "1898" and states that he naned this taxon as a variety of V eat edels L. there, which is not so. He also notes "La GiiaGen que nous avions faite en 1899 entre le V. yenosa Gill. et Hook, type et une vari6ét6 Reineckii basée sur Ta forme des environs de Porto Alegre, ne résiste pas a l'examen de matériaux abondants. Nous y voyons plut6t maintenant une simple forme locale." ¥ In all, 7 herbarium specimens and ), mounted photographs have een Citations: PARAGUAY: T. Rojas 3365 (Herb. Osten 18171] (N, N, Ug), Sen. [Hassler 216; Herb. Osten 1566] (Ag—-photo, peta N--photo, Ug, Z—-photo). ARGENTINA: Corrientes: Ibarrola (6g-35927555 Geo, gemue ORL (Gr-—262670), iclaaseT Wodinn 37 (Gg--35327h) < VERBENA RINCONENSIS Moldenke, Phytologia Fe 100. 1963. Bibliography: Moldenke, Phyto Ologia 9: 100. 1963; P sage Ré~ sumé Suppl. 6: h. 19633 Moldenke, B Biol. Abstr. 43: 643. 1963. Creeping herb; be. slender, rooting, mostly procumbent; branches asce piine, very elender, rather sparsely pilose with rather stiff and sr pledge ci te hairs; principal internodes e- longate, 2--9 cm. long; n ye annulate; leaves decussate- opposite, very variable aa pe and shape, petiolate; petioles Slender, Pa ager Ley rather sparsely spreading~pilose with erect whitis h hairs, more or less margined, especially toward the apex; leaf-blades a tndin-ohartapeone; rather uniformly von on both surfaces, ovate, 1.5--3.5 a Pools irregularly incised, the larger ones rather sparsely mie a both surfaces, the lobes and divisions oblong Sven recurved, attenuate-acute at the ag pilosulous on the back, long=ciliate; ¢ calyx cylindric, about 7 mm. long, rather densely subappressed=pubescent on the outer surface, plainly S- ribbed, the teeth 5, subulate, 2 long and 3 shorter: corolla hy- pocrateriforn, blue, its tube about 8 mm. long, venose and pil- °sulous on the outside toward the apex, the limb to 7 ma, wide. The type of this species was collected by George B. Hinton (no, 8011) on a hill at Rincén, in the district of Temascaltepec, co, a 8 on August 29, 1935, and is deposited in the her- barium of the Texas Research Foundation at Renner, The Collector describes the plant as procumbent, growing on a clay hill at 1960 meters altitude, flowering in February, haroh, and 100 PHYTOLOGIA Vol. 11, no. 2 August. It has in the past been confused with and distributed as V. pumila Rydb. In all, 11 herbarium specimens, including the type, have been examined by me Citations: MEXICO: México: Hinton 5589 (N, W--18222h)), 80121 Oe ee ee K—isotype, Ld—-isotype, N--isotype, Rf-- type), 8988 (K, 1d, N, W-182229h). VERBENA RIPARIA Raf., Herb. Raf. 69, nom, md. 1833; Small & Hel- ler, Mem. Torrey Bot. Club 3: 12. 1892 Synonymy: Verbena urticaefolia riparia Britton, Mem. Torrey pie Club Si 276. 109k. Verbena urticifolia var. " piperds Britton » Ann. Mo. Bot. Gard. Gard. 20: 267, in syn. 1933. Verbena ctlcharelie var. riparia (Raf.) Britton ex Moldenke, Suppl. List Invalid Names 10, in syn. 19). Verbena pb var. ripa- ria Britton ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 197. Bibliography: Nutt., Gen. 2: maf ates Raf., Herb. Raf. 69. 1833; Small & Heller, Mem. Torrey Bot. Club 3: "12. 18923 Ne. L. Britton, Mem. Torrey Bot. Club oy 26. 189k; Dur Durand & Jacks., Ind, Kew. Suppl. 1: 51. 1906; Robinson & Fern, in A. Gray, N Man. Bot., ed, a 688 & 92h. 1908; Mo Re Day, Check List —e 1908; N. Taylor, Men. ic Y. Bot. G (Fl. Vic. N. Ye] ‘ 1915; Perry, Ann. Mo. Bot, Gard. 20: 2h7, 259, 267, & tae 19333 7p @. > Man. Southeast. Fl. yan 1933; Moldenke, . Alph. List Invalid Names 6 & 49. 190; dare ae Suppl. Invalid Names 10. 191; Moldenke, Alph. id Names i7 & 51. 1942; Moldenke, ° . baa. st 9 denke, Alph. List Invalid Names Suppl. 1: 27. 1947; Moldenke, Alph, List Cit. 2: 170 & $08 (1948) and 3: 911 & 9h2. 19L9; BD» Merr., Ind. Raf. 205 & 295. 1949; Moldenke, Known Geogr ib. Verbenac., [ed. 2], 6, 7, & 199. 1949; Fern. in A. Gray, Man. Bot., ed. 8, 1209, 1210, & 1612. 1950; Moldenke in Gleason, New ae f Br. Illustr, Fl., print. l, oe 126-128 (1952) and . 2, 3: 126128, 19565 Moldenke, Am. Midl. Nat. 59: 347. 1958; Moldenke, Résumé 9, 10, 366, 377, & 73. tre “Yoldenke, résumés ra. 2 1 Neen us 20 (1962), and 7: 10. 1963; Gleason & C c. Pl. 579 & 580, 1963; bioldenke, Phytologia 8: “ibe «(1963), $3 ngs "3, 203, 204, & 220 (1963), and 10: 217 & 1961, oe a testes : Moldenke in Gleason, New Britton & Br. Illustr. Fl., print. 1, 3: 128 (1952) and print. 2, 3: 128. 1958. herb 1 he base, sparsely strigillose on both surfaces, the venation prominent beneath; spikes slender, elongate, paniculately dispo~ sed; bractlets lanceolate-ovate, as long as "the calyx during an- thesis, acuminate at the apex; calyx to 3 mm. long, minutely 1964 Moldenke, Monograph of Verbena 101 cent, subtruncate, the teeth minute; corolla one enarginate or notched at the apex; coces. oblong or ellipsoid, 2--2.5 mm. long. The species has been found in rich thickets, on riverbanks, and on the banks of streams, very local in distribution, at an altitude of 2100 feet, flowering and fruiting in June and July. Fernald, Small, and Perry all report that it grows in New Jersey, but cite no specimens from that state and I have not seen any from north of Virginia. The name, Verbena riparia, is usually apres to gies que "ex Small & Heller, 1 Mem. Torrey Bot. Club 3: 12. 1892", as, for instance, by Durand & Jackson (1906) and Perry 7 (1933), bat” ac- to E. D. Merrill (1949) it gu ag publi Rafin- esque himself (1833), although as a nomen nudum. gual & Heller (1892) give "1830" as the date for the pa publicstion, but Merril not seem to substantiate this. They also claim that ep ee type is preserved in the Columbia University herbarium I have not been able to locate it there. Merrill claims that Rafinesque based his name on material from "Long Is- land, New Jersey, & Chesapeake Bay", which is most amazing, since this. species is know to me only from Virginia and North Carolina. Nuttall's type of v. hastata var, oblongifolia does not enter the Picture question by Small & Heller when Rafinesque's name was validated by a description — this description being based on two North Caroli- na collections np bis ayes arently should be considered the real co- whee oa the & Hell Globe", is probably the actual collection here referred to and is arded by m Stanly Yadkin' referred to by Small were on the Yadkin River, in 102 PHYTOLOGIA Vol. 11, no. 2 tanly Co., North Carolina, and are also referred to as the ‘'Rap- ids of the Yadkin'. They were apparently in the vicinity of Ba- din, North Carolina, but are now under high water from a dam that was placed on the river just below there.....In Mem. Torr. Bot. Club 3: 20-2] (1892-1893) Small mentions the Falls of the Yadkin This would seem definitely to tie it into the area about Badin.” This specimen has not yet been seen by me. The original Rafinesque reference reads: "Autikon Maritimum. Collection of )) new or rare Plants from the Atlantic shores of ription. Nuttall's description reads as omaeey: "),, hastata. Flowers deep blue; rarely ever hasta 8 s there appears to be a very dist. variet; y near piiladslphia which I shall dis- tinguish by the name of f£.* Oblongifo lia, having oblong-lanceo~ late ves, merely acute, ot 0 not acuminated; this be V. paniculata of Lamarck? but the flowers are not imbri- cated, nor in the least corymbose, it appears to be equally re- mote from V. diffusa of the same, but assuredly intermediate, if not a weal betwixt V. hastata and =. urticifolia. It has only occurred to me twice on the banks of the Delaware." It seems to me much more Likely + that Nuttall's tsi is xV. engelmannii Mol- denke, as, Rafinesque's original plant may well also have been. I cannot agree with previous authorities who have placed Nuttall's name in ‘the synonymy of Vv. Dr. Small apparently also collected Vy. of se Le. on the Middle Fork of the Holston River, near Marion, Smyth County, Virginia, on July 6, 1892 (which see). The leaves of the Vv. ri- material are considerably larger than those on the V. . of- ficinalis specimens. Fernald (1950) distinguishes these two very closely related species as follows: V. officinalis — ma~- long. V. ri a — mature calyx 3 mm. long, about as long as the awa ractlet; sith bluish; mature schizocarp twice as as broad, Material of Vv. ri a has been misidentified and distributed 7 herbaria as pf ES Vahl. Taylor (1915) reduces the species V. urticifolia L Sees (935) « cites the Fellorine l, additional specimens not 48 seen : VIRGINIA: : J. K. Small s.n, (Marion, 6 July pain (, 0 6). NORTH CAR! cee ‘Caldwell Co. Co.: Small & Heller son. [near Globe, July 3, 1891] (F). Stanly Co.: small & “Heller s.n,. [near falls. of Yadkin, 18 Aug. 1891] (F). She sa says "The ma~ 196h Moldenke, Monograph of Verbena 103 terial at hand is too scanty and too immature to give many clues to the probable relationship of this species. It would seem as nearly related to V. officinalis as to either V. hastata or V. urticifolia. It is characterized by pinnatifid | or r tripartite mens minutely glenda ler-gobesoent inflorescence, and fruit a- t twice as long as thi In all, 34 herbarium te including the type, have been by me, Citations: VIRGINIA: Smyth Co.: J. K. Small s.n, [near Marion, dune 22, 1892] on : 1, Ok), sn gan (ise ddle Fork, Holston River, on, July 1, 1892) Fes. Ba, Ba, Cm, Cn, Fl—11173, es H— 3598), 5975, ob—50829, P, *po=269312, Up, Up, W 298398, We), Naatne Fork, Holston River, Marion, July -—" 1892] (al, c, cos "P6; 1020755, Ob——-50828, Ot, ¥-71939), 8.n. {July 20, 1892] " (Up), Bets ee Marion, July 22, 1892] (Ca— 25188). Ragged Island: Fernald & Long 12453 (N). NORTH CAROLINA: Caldwell ao Small & bette sen. [near Globe, July 3, 1891] (C— logotype). LOCALITY OF COLLECTION UNDETERMINED: Herb. Bot. Mus, iund, sen. (North America] (Lu). VERBENA ROBUSTA “ystmewe Pittonia 3: 309. 1898. : Verbena strato-stricta Palmer ex Moldenke, Résumé Suppl. 7 in syne 19 a5. Bib ograpiy: Greene, Pittonia 3: 309. 1898; Thiselt.-Dyer Suppl. 2: 191. 190k; Jepson, Man. Flow. Pl. Calif. iP. 859 & 12 36." 1925; Perry, Ann. Mo, Bot, Gard. 20: 248, 261, 292--293, & 356. 1933; Munz, Man. So, po Bot. 437. 37. 19355 Eastwood, . adhe Bot. 3: Bi 1941; Moldenke, Known Geogr. Distrib. [ed. 1), 15, 19, 80, & 102. 1942; Jepson, Fl. Calif. 3 "Oe 3 132 19,3; catalase’ Known Geogr. Distrib. Verbenac. Suppl. 1? m sf 1 h HTT, Wee “39, iss, is 7 a 75, 477 2, 483, 488, 492 2 “rt 16, S71, 58h, 5 é1 (sta), Ass a th , 859, 892, gs 3 3h » 976, & 7 (19h9), ot "1127, 1133 0, 982 1139, 1155) 1198, 1199, 12231227, 1229 1230, 1237, 1239, 12hh, 12h5, 1247, 1252, 1253, & 1290. Gentry, Hancock Pac 27, 1252, OnE 33 & 2hh. 799; Moldenke, Known Ge~ v + Distrib, Verbenac. oo. 3e She Moldenke, Phytologia 3: 141 Ql) 3s 264. 1 : * a Soc. Dict. Gard. 63 2209 & 2212, 19513 dl. Nat. 59: 34h. 1958; Moldenke, ane, ae least Sa 22h, & 73. 1959; Moldenke, Phytologia 8: 120 & 1961) and 8: 278 & 279. 1962 Hag ¥ asco “Hanns Suppl. 3: 9 (1962), 5: 5 1365. ho 6s od: (2963 » and 7: 9.1 Raven, Aliso 5: 302 & 3%. me » Phytologia 8: re thse3) and 9: 17, 3h, & 215. 104 PHYTOLOGIA Vol. ll, no. 2 r semi-erect perennial, 1 1/2 to 5 feet tall, sometimes with a Sent spread, often stiffly erect, harshly hairy through- out, sometimes very floriferous and with the athens quite cram usually e arenes or sparsely hirsute} branches ascending or erect, leafy; leaves decussate-opposite, ovate or oblong-ovate, 4--10 cm. long, rounded at the apex, cuneately narrowed toward the base and into the winged petiole, bright-green on both surfaces, usually 3-cleft, irregularly serrate-dentate with sharp apiculate teeth or incised, ose and scabrous-pubescent above and less harshly pubes- ry h to aces, the venation prominently reticulate; inflorescence showy; spikes P peasy » often crowded, subsessile, thick, usually densely many-flowered even after anthesis but occasionally elon- gated, densely glandular-hispidulous throughout; bractlets lanceo- late-subulate, equaling or surpassing the calyx, densely ¢ de lobes obtusish, terminating very unequal pas gaia teeth; corolla’ hypocrateriform, varying from purple o sh lilac, lavender, or violet, its tube 5--6 m. ert a little longer than the calyx, puberulent on the outer surface, hairy in- side, its limb 3--) ma. wide; fruit mostly contiguous; socci ob- long-trigonous » 2.5 mm. long, raised-reticulate above, the nia, "flowering in the middle of the dry season". The Greene collection cited below fran this same county may represent a co- type or isotype. The type of V. prostrato-stricta was collected by Edward P at San Luis Obispo, in the county of the same name, California, in 1876, and _ deposited in the United States National Herbarium at Was Verbena robusta differs ies the closely related V. lasio- stachys 1 Link in ha’ ae its corolla 5-~6 mm, long, the limb }+-h in dry streambeds, rocky loam, marshy ground, fallow fields, moist soil near the mouth of canyons, dry or rocky creek ek beds, creek bottoms, small canyons, small ravines, gravel, salt marshes, dry clay soil around rocks, and drainage ditches in fields, at sunny mo 1 canyon floors, and beach dunes, at altitudes of 5 to 3500 feet, flowering from April to December and M er. Abbott found it "common on 196), Moldenke, Monograph of Verbena 105 sea level”, Twisselmann mgt that it is "scarce in mud of small pool, broken off by cattle, in full sun, in a ies Sonoran sub- shrub association"; Fosberg also found it "in dry creekbed in Up- per Sonoran Zone." Ewan 9726 is described by the ait hess as follows: "herbage viscid, subslimy, plants 5 feet tall gravel alluvium, leaves roughish." Gowsiiden ( et yor says xr it is "common on the margins of springs, rarely in streambeds, 500 to a — nats Mt. Bisbal, June to nhac associated with Oenothera hookeri, allis arvensis, ae isin fa poe “i describes the tortscense as "small is of lilac poet common name recorded for the species on "Catalina vervain", Material of this species has been misidentified and distribu- ted in herbaria under the names V. bracteosa Michx., V. ciliata Benth., vy. hastata ta L., ve lasiostachys Link » Ve lasiostachys va var. scabrida M Moldenke, ve. Oly stac H.B Ke, v. prostrata Re Bre, V. scabra Vahl, ve stricta bart. and even Rumex sp. On the other hand, the Murbarger 218 8 and R. A. Plaskett 142, distribu- ted as V. robusta, are actually v. ~Jasiostachys Link, while V. Fe Hesse 2606 and R. A. Plaskett 98 @ are V. lasiostachys var. septen- trionalis } Moldenke. E. K. E. K. Abbott s.n. [Monterey, 1889] is a mix- ture with V. abramsi Moldenke, while M. W. Williams 15 is a mix- ture with a species of Salvia. Wiggins & Demaree L7ad is not typical —- the spikes are thin -- nor is ) Grant 5. sn. . [Avalon, Aug. 6, 1902], both originally determined as V. lasiostachys var. scabrida. Rattan s.n. [Shell Mound Park, , June 1880), Ferris 7656, = 2 Smith 13, and C a Globee oe S01 have the leaf~blades smooth (not eens) above. these coll ons ve and look Lag like a possible hybrid with V. lasiostachys, as, ote fact, the Clokey specimen in the Dudley Herbarium was actu- ly anno tated rig someone, Howell 00) is very floriferous and sh Owy, with rbage quite glandular. Gentry (asie)” states that the species "apparently originated on Catalina an ted e mainland". He claims that it is Cruz, Santa Catalina, and San Clemente. Peter H. Raven, however, t@, 80 this record is clearly based in turn on “that of Muns, Man- ual Southern California Botany, p. 437. 1935, which correctly lists 'y. prostrata! from San Clemente Island alone, of all the California islands. Therefore, the wa seems clearly to in- dicate that only V. lasiostachys is o San Clemente Island, only V. robusta on San ee. Rigi cisitane, ge Cruz, and Santa Rosa islands." The spec apparently introduced into culti- vation in 1843. Jepson (193) cites many specimens. 106 PHYTOLOGIA Vol. 11, no, 2 Perry (1933), under V. prostrata R. Br., cites the following 3 additional specimens not as ze seen by me: CALIFORNIA: Siski-~ you Co.: E. L. Greene 860 (E, F, G). Under V. robust she ott the following | 19 specimens not as yet seen by m 6: CALIFORNI Alameda Co.: Congdon s.n. [salt marshes, Oakiand, Sept. 1886] ae Prgelaas 8.M, bower back of Berkeley, 23 Oct. 1880) (E); E. 8.N. Thin Berkeley, May 1887] (W); Michener & Bio- Tet os “G ). dor Co.: Braunton 1263 (E). Marin Co.: East- wood 5.n. iraeree, July 26, 1900] (a). San Diego Co.: Edw. Pal- mer 310 (E, F), sen. [Jamel Valley, 26 June 1875] (G). San Luis Obispo Co.: Edw, Palmer 341 a (F, G). San Mateo Co.: Elmer 4950 (E). CHANNEL ISLANDS: Santa Catalina: J. I. Carlson 5.n. TAvalon, June 13, 1915] (E, ay Macbride & Payson 850 (G); Tra Trask s.n. (Avalon, Sept. 1896] (E), s.n. [Avalon, May 1897] (F). ~vEXI- CO: Baja California: C, R. Orcutt 1301 (E); Wiggins & Gillespie 3977 (E). The Palmer 310 which she cites Poke "NY" is actually in the Barnard College herbarium and the 341 1/2 is in the Colum- bia University herbarium. The E. L. Greens s.n. which she cites from "US" is probable the E. L. Greene 76 cited by me hereinafter. barium as "atypical". She says "Verbena robusta has been much confused with the nearly related V. prostrata, but is mite read~ ily distinguished from the latter by the brighter green color of the herbage, the scabrous upper surface of the leaves, and the dense spikes. Moreover, the mature calyx lacks the mark- ed tendency toward subconnivent i = a characteristic of V. prostrata, The schizocarps of t re very much alike. The collection Orcutt 1301 has a iseahag" siadeatee inflorescence and the rane are not particularly scabrous. The specimens Braunton 99 and 1263 show unusually long floral bracts. These are probab- ly atypical phases of the species or yoeetoly hybrids." Another possibility is that they may represent V. lasiostachys var. scab- Moldenke,. In all, 227 herbarium specimens, including the types or topo- types of all the names involved, been 6 y me. Citations: CANADA: Nootka Island: Née 90 (Q). CALIFORNIA: Al- ameda Co.: M. S, Baker 10504 (Ca—80l538); J. W. Blankinship s.n- (Berkeley, July 22, 2, 1991] (C. (Ca~-397h06); Brandt s.n. [Berkeley Hills, July 9, 1914] (Ca~-176060), s.n. (Wildcat Canyon, July 16, 191k) (Ca); Chesnut s.n. [Oakland] SiGe Collector undes- ignated s.n. [Oakland] (Sg—16088); Eas tman s.n, [near Livermore, Nov. 1 T8987 {ca—104873) ; E. L. Greene cmc (W-13231h2); J. T. How ell 19838 (Gg—20h)8h), 18173 (Gg—319113) ; Jepson 12935 5 (Ca); 3 Ce Ee H. Michener Bon. (Temescal, A i: 1891] (co_—7al); Hick Michener & Bioletti 1886 (Ca——35053), s Oakland, Aug. 1891] (Ur), 8-ne ——oot -f[ TLeona, Aug. 6, 1892] (c, da3s157, 12060, Pl--22603), 5M: enamel 196), Moldenke, Monograph of Verbena 107 {Berkeley, Aug. 9, 1892] (Ur); Rattan s.n. (Shell Mound Park, June 1880] ae a a 4 Walker oe pp Denese nia Co.: Braunton 3 (MN). tra Costa Co C11), 399 (Ga=S9155), 64 (comonti6), IBY (ox G157) kK. Brandegee s.n, [betw, Martinez & Port Costa, July h, 1907] (ca— 139763); Ewan 9726 (Ca--723797, En, En, Re—22 231k); J. T. Howell Ujh0 (Gg—172h421); H. S. Yates 1069 [U. S. Dept. Agr. Forest Serv. 5601] (Ca-12419). Kern Cos G. H. Horn s.n. [in Owens Valley and at Fort Tejon] (N); Twisselmann 1535 (Gg—391009). — Angel- es Co.: Blake 70k (Ur); Braunton 99 (W--l6510k); Ep & s-n. (Mint Canyon, May 1925] (La); G. B. Grant 5305 (Du--75581), Sen. [June 17] (Du~-75651)3 Knoche 879 (Du). Marin Co.: Eastwood 3.a. (Tiburon, July 26, 1900] (Ca--10)872); H. Edwards s.n. [Lag- unitas Reservoir, 4/77) (N), Son. [Saucilito, Apr. '77] (N) E. i = Greene B.D. (23 July 1891] (I); Jepson 21230 (Ca), s.n. (Tri- » July 20, 1891] (C). Mendocino Co.1 Eastwood 9320 (dg— ar; » 10647 (Gg—-31337) . Merced Co.: Abrams 5295 (Du--67811) . rey Co.: E. K. Abbott s.n, (Monterey, 1889] (Mn—-23135, Ma Me ere abe Esai ishaw 2731 (Ca--570390); Eastwood & Howe e—119973); Je T. Howell 23360 ye ay S), pay (B), Tale (2), 400k, (B); L. S. Rose 35605 (N). Napa Co: Je 3 rt. Howell 5599 (Gg—181600); Raven 3953 (Gg—366303). Orange Ewan 3612 (B (En), 7691 (En); ie A R. Fosberg S.5146 (S). San Pent Cos: J. T. Howell ell 11527 (Du—219218, Gg—21258); Raven 8855 (Gg—~h081k7); Rodda Ee Se gga May 16, 1926] (Ge— 10322), San Bernardino « L. Greene s.n. (Highland Sta- tion, Aug. 7, 1887] (Wa 56191) San Diego Co.: Abrams 3787 (N); Cleveland s.n. [Sweetwater Valley, June 15, 1885] (N, W—205393k); Epling, D Darsie, Knox, & Robison s.n. [Flin Springs, June 19, 1932] (Ca—520076, En, La); Gander ler 222 (Sd—11633), 251.3 (Sd— twooe.” st A.226 (Ba); Edw. Palmer 310 (Bc, Pa); S. G. Stokes s.n. Mesa, Aug. 189 Du--2)210). San Francisco Co.: Good 8.D. (188 7 pny ly San aD ae Co.: Suksdorf 3 foctepend $2: Luis Obispo Co.: K. Brandegee sen. [Lagoon] (Ca-~185316)3 "I. Js Co" aon. (Nov. 22, 1907] (Ca--l56151); R. J. Ferris 7656 (Du— 206261)5 Ba Edw, Palmer au (Pa), 32 ae (C, Pa, W—56223); M oe R. Es Gamers 5 SM. X, penis Pl—81730, 86, 5836) J. T. Howell AL7LS (Gg—269702); L. He Pammel s.n. (a Honda, Sept. 13, 192) (1o—103282) 3 L. W. Reinecke s.n. (Redwood City, June 13, 1937] &—~77001,6 ber 307 (Ca—615560); Suksdorf 30 (P1-- 138409) . oe Sotreiber a bt ecitaod 3 868 7 See), s.n. (Zaca 108 PHYTOLOGIA Vol. 11, no. 2 “i June 17--22, 1902] (N); Morton 8698 eee J. Torrey 416 (T); M. W. Williams 4 a m—35079). Santa Clara Co.: Abrams a (Du--1)3857)3 J. Be 28 peas 289 (Ca-~25162) Dudley s.n. Selle ieee ik , 1901) (Du-—2786 36 ), 8 8.M- {Oct. 10, eos (Du—2)180) ; F. We Johnson 52h (N)3 C. P. Smith 1155 (us); Re Je Smith 13 (Du——77583). Santa Cruz Co.: fs A. A. Reed s.n. [A- pril 10, 1919] (Gg-—31339). Siskiyou Co.: hak L. & Greene 860 (Be, Pa). Solano Co.: Jepson 20250 (Ca), s.n. {Alamo Creek, June 8, 1881] (Ca~-2515l). Sonoma Co.: J. Be Davy 865 (Ca--25159) 3 E. Samuels 161 (Ky). Tuolumne Co.: “ae ers ne 28 (Du—67519, N); Mrs. W. J. Williamson 189 Pee, Gg—313u5, AES fat ount nem ne en ree omen senna NEL ISLANDS Sabie. Catalina: K. Brandezes e sn. [May 1916 (Ca—185)15) ; J.°X. Carlson #.n. TAvaion, April 28, 1914] aa 31369, W—88059L), s.n. (Avalon, June 13, 1915] (Gg—31366, # 1070618); Dunkle 1955 (Hp, Po--1556 74) Eastwood 6500 "(Ge 31367, W~1100h81); ¥. R, Fosberg $.4606 (N), S.u7ul (Ca--882793, N), $.48)3 (Ca--625012, Ca—882792, I, N, S, W—1766310), S.5412 (Cam 625057, ca--88279h,, En, I, N, Up, W—1766351); G. B. Grant s.n. {Avalon, Aug. 6, 1902] (Du—91161); Poumey s.n. [Sept. 17, 1 159L) (Ca--104,871) ; Trask Sen. [Avalon, May 1897] (W-—3l0303), Sm. [Avalon, Aug. 1901] (N); C. B. Wolf 3605 (Du--30075h, Rs—3588), 212 ae et Du-300760, | Ra—3595, 1 W--18),5569) . nia? oe! 38 (S)3 T. S. Brandegee s.n. [1888] (Ca~-16922h)3 Clokey ae te ( T8800, a N), » SOI (Ca 3 i Ca—882796, = Du 261238 designated 38-291/38-292 (Mi); F. H. Elmore 291 )j Soldector we Me Ee Jones s.n. [Valdez Bay, 9-5~27] (Po—-172767, Ua—~1h8h3)5 Mrs. We W. Rand 26 (Gg—179175); Swain s.n. [1919] (Gg—313h8); Thatcher Son. (May 1921] (Ca); M. W. Williams 15, in part (Gg—310872), bo (Du-—289568); C. B. Wolf 1160 (Ca--882808, Gg——381235, Be 550)> MEXICO: Baja California: C. R. Oreutt 1301 (C, Ca-—2516 ealages Wiggins & Demaree 70 an Bish, wacesi2}} ‘ee & Gillespie 3977 (Du-—196037, Ge—179399, Me, yi i, N, O— 5082h, Po—-193008, Rs: Rs-—19050, Waer ee CULTIVATED: Sweden: (s sect natal ont net hte pthc VERBENA RUGOSA Mil. - Dict no. 18. 1768 [not V. vr oe sa D. Don, 1836, nor i “ash, nor Muhl., 1809, » 2845). paitiogenten P. Mill., Gard. Dict., ed. 8, no. 18. 1768; J. SB in Rees 2 Owes conte aes he "a8 Steud., Nam. Bot., ed. 2, 2: 750. 1845 Walp., Re 18h5; 3 Jacks. in Hook.f. & Jacks., Ind. ew, 2: 1179. 1 1 Bos Known Geogr. Distrib. Verbenac., [ed 196k, Moldenke, Monograph of Verbena 109 1}, 19 & 102 (1942) and [ed. 2], 33 & 199. eet: Moldenke, Résumé 4o"& 473. 19595 Moldenke, Résumé Suppl. 7: 8. 1963. Nothing 4 own to m on this plant oe Miller's orig- inal dees ean "Verbena (rugosa) diandra, spicis ovatis, fol- dis subrotundis serratis & rugosis, caule fruticoso ramoso. Ver- vain with two stamina to the flowers, oval spikes, roundish, san- ed, rough leaves, and a shrubby branching stalk. Sherardia ar- eiviatess nodiflora foliis serratis & rugosis flore purpure: purpureo, ae caer geen Senin tae nm gree eaten —siarnse@seesane _-manmememae these seldom appear in this country, and are not succeeded by Seeds heres but the plants are easily propagated by cuttings the summer months, and may be preserved many years in a moderate stove." Walpers (2845) places it in his group of f sepia at the end of the genus known by name only. Obviously, with only two stamens, it cannot ei a true Verbena. I suspect that it will prove to be a aaeputarpheta. Schauer apparently does not list it either a- mong or excluded species. Smith (1817) seems to regard it asa synonym of V. s simplex ex Lehm., which is not very probable. VERBENA "a A ining Phytologia 2: 25—26. 19h1. yonii Moldenke apud Plant. Exsicc. arey 27, in syn. 197; chit tenden, Roy. Hort. Soc. Dict. Gard. 6: 2305 & 2212, zine, Moldenke, sis Ae 2: 25—-26. 1941; Moldenke and 2: 328. 1947; Moldenke, Alph. List Invalid 26.197; H. N. & As Le Mo htia 1: 225-227. 19183; Mo denke, Alph. List Cit. 5 407, 1s3, 457, L9L~—-L96 (1948), 3: 678, 681, 683 806, , 873, 882, 883 & 939 (19h9), and kh: 98 060, 1284, 1213, 1218, & 1230 1949; Moldenke, Known bd. V ed. 2], 2h, 27, 16h, & 199. 19h93 Mol in ttenden, Boy mort 80 a Di 2209 & 2212, 1951 tg age ag ae 29535 xg » Rf 22h, 33 & um. Th 19585 ‘wel- Tall, erect, coarse, annual or perennial herb, usually 18—2) 110 PHYTOLOGIA Vol. 1, no. 2 inches tall; stems erect, green, rather stout, sharply tetragonal or square, hollow, larger below, sparsely hi rsutulous with short whitish divergent hairs especially on the angles and at the nodes, glabrescent in age, more or less scabrellous on the angles; bark green or dark-green; roots fibrous; internodes elongate; leaves decussate-opposite, sessile, clasping, dark-green, oblong in out- line, 2--6 cm. long, 0.8--3 cm. wide, more or less 3-parted or “lobed, each division pinnatifid-incised with broad acute teeth, ab ly hirsutulous on both surfaces with rather short whitish often bearing 1--3 pairs of much reduced leaves near the base, the flowers close together near the top of the spike, with a faint odor or odorless, very densely imbricate before and during anthesis, rather uniformly separated in fruit; peduncles (2-6 CMe long) and rachis slender, sharply tetragonal, rather densely or sparsely spreading-pilose or -pubescent, glandular, the pub- escence very short; bractlets linear~lanceolate, about 3 mm. » 8 attenuate, rather sparsely pu- berulent and glandular, the margins d irregularly ciliolate toward 6; calyx tubular, about 3 mm, lo t NE» glandular-pilose with short spreading hairs; corolla blue or light-blue to lavender, rages cogs violet, or purplish, a- bout 6 mm. ise, 1 its tube puberulent at the a apex on the outer surface, the limb about | mm The type of this wort was collected by Robert Runyon (no. 24,85) —- in whose honor it is named —- in clay soil in o moist ground and ditches in the El Jardin Tract, at an aoe m C ps deposited in ~ P sing ong Herbarium at the — ges Botani Garden, xutha Runyon says about V. runyoni: "rare in this region: it occurs in moist ground, never in dry fields", "occasional: it occurs only in moist ete eg “occasional in semi-dry fields", and "abundant springtime". Hanson reports that it is "frequent in openings in woods", while F. B. Jones calls it "locally abundant" in Nue- ces County, Other oe have found it in open or banks, prairies and coastal prairies, and along roadsides, at ring and It is called "Rio Grande vervain" and was a apparently in ced into cultivation in 1901. Suksdorf 2955 was collected at Portland, Oregon, presumably as a waif, on October 26, 1900 -~ 196) Moldenke, Monograph of Verbena lu seeds were taken from it and planted in a garden Bingen Washington, and specimens pap there on ieeuek 10, 1902. Runyon describes the fruit "a small capsule", but itisa schizocarp. L. I. Davis, ie a letter to me dated March 25, ae refers to several color forns of this species which he state that he is thinking of naming and desc ee _Thus far I nie of only one that has actually been published by Material of this species has been gprs Poe and distribu- ted in herbaria under the names V, halei Small, V. neomexicana (A. Gray) Small, V. neomexicana var. xylopoda Perry, V. officin- alis L., v. inthe Lehm., and "Verbena hybrid*. Ferris | & Duncan Duncan 3160 was annotated by Perry as "aff. V. mutha", R, Runyon 2560 is accompanied by a photograph of the . plant growing in situ. In all, 105 herbarium specimens, including type material of all names * involved » and 2 pines pho coureehe and clippings have been examined by me. Citations: TEXAS: Aransas Co.: Cory 903k (N). gr CO: Killip 42107 (W—2069798); Lundell & Lundell 110% (N, Rf, W—19269L0). Cameron Co.: J. J. F. Brenckle ckle 47-350 (N); re = Isis (Mi), 1695 (éi); Cory 28299 (N), 36467 (N, N), 51439 (Sm); Mrs P. Cottrell 8743 (Au); L. Davis 35 (Au--122639, Mg be Ie ois en, ised May 1942] (Au, Gg—-316103, N, Sm, W—1573783); Ferris & an 3160 (Du-~125)23, Gg—31h05, N); H. C. Hanson 496 Ce M. C. Johnston 253-7 (au—-12261); C. L. Lundell 20679 mr, Bile 10681 (Sm), 10709 (N, Rf, W-1926932); Lundell & 0012 teem Mi, N), 10753 [Plant. Exs Exsicc. Gray. wad a, ore , in, edie! Dp 31803, eect sabes, ou 23896, H--81531, 3014, I, Ka--95213, Ld, 14, it » Ms, N, , Or) t nadaatgh ‘be. nf, 8; Sn, §t—23303, m oe vi, ¥2-132578h, We); A. R. \Moldenke *190 (Fg, Z)3 Rose & Russel 24238 (1369527) Re , Runyon 2u85 (It--isotype, N--type, N--isotype, N-isotype), 2585 (N, N, N, Ny N, N, N, N, N, N, N— photo), 2691 (N, N), 2692 (N, N), 3178 (Herb. Texas Agr. Exp. Sta. 43663] (au), 4287 (au, N, S)3 Sixth Grade Brownsville 34 (Au), 5. n. [Nov, 1934] (ar (au); Tharp 1201 (Au, W—1116114). Nu Nueces Co.: Fe Be Jones 1925 (Ww), 6191 cae F. W. Pennell 10340 (N). OREGON: Wultnamah Co.: Suksdorf 2955 (Pl—-130h16). MEXICO: Nuevo Leén: Re F. Smith 76 Bee CULTIVATED: Washington: Suksdorf Sen. . (garden, B Bingen, Aug. 10, 1901; from Portland] (P1—-138362). MOUNTED CLIPPINGS: Moldenke, Phytologia 2: 25. 1941 (Wf). VERBENA RUNYONI f. ROSIFLORA L. I. Davis in A. M. T. Davis, Study Boscaje Palma 62. 192; L. I. Davis, Nature Leaflet [Lower Rio Grande Valley Nat. cua] 2s (h]. i9hS. : Verbena runyonii f, rosiflora L. I. Davis ex Moldenke, Alph. List Invalid Names 26, in syn. 19)7. Bibliography: A. M. T. Davis, Study Boscaje Palma 62. 192; nu2 PHITOLOGIA Vol. 11, no. 2 ager Known Geogr. Distrib. Verbenac. Suppl. 1: 2 & a 194433 vis, Nature ane Rio Grande Valley Nat. Club) tht. 19s Moldenke, Alph. List Invalid Names Suppl. 1: 26. 1947; Moldenke, Wrightia 1: 26. “T9483 Moldenke, Alph. List Cit. k: 1218. 199; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 2h . a. Renta’ Moldenke, Résumé 30, 373, & 47h. 1959. rom the typical form of the species in hav- ing ante eephae print tbee The type of the form was collected by L. Irby Davis at South- most, Cameron County, Texas, in May, 1942, and is deposited in . + is Davis May 1942] (Au—type, N--photo of type, Z—-photo of type). R Moldenke, Phytologia 2: 55--56. 191. Bibliography: Moldenke, Phytologia 2: 55--56. 19h13 ee Known Geogr. ° Ae ae Moldenke, Anal . Let Biol, Mex. 20: 1h. - Je Salisb», Kew. sul. ll: 263. 1953; Moldenke, sicee No & ih. 1959 erb, not at all nigrescent in drying; stems slender sharply istnigtiet. very minutely and obscurely evar hout; nodes annulate; principal internodes 2--6.5 cm. long} leaves decussate-opposite, the w st ones sessile, the lower and older ones petiolate; petioles to 1 cm. long, winged, hirsute- pubescent; lower leaf-blades elliptic or ovate-elliptic in out- line, 1-4 cm. long, 5--15 mm. wide, acute at the apex, irregu- larly incised-dentate along Damen margins » long-attenuate into the petiole at the base, aah 4 ensely hirsutulous and glandular on both surfaces, especially beneath, with stiff whitish hairs, the margins eden. ges. revolute in age; "midrib and secondaries flat or sub ssed above, p beneath; upper leaves ear-lanceo~ late, picts. Pee ts mm. long, irr ly incised-dentate or en- , & hirsutulous on both surfaces, with a very 3 ulately branched, the branches to 20 cm. long, dense 3 peduncles slender, sharply tetragonal, minutely ‘ly puberc- lent, often elongate; rachis densely glandular— puberulent; poked lets lanceo » 2--2.5 mm. long, attenuate to the sharply acu apex, densely glandular-pubes cent and ciliate~margined; c hie tubular, about 2 mm, long, densely glandular-puberulent, its rim Standley, and Paul George Russell (no. ) a moist field in the vicinity of Culiacan, Sinaloa |, Mexico, on April 21, 1910, and is deposited in the Britton Herbarium at the New York Botanical Garden. The species is named in honor of Paul 196) Moldenke, Monograph of Verbena 113 George Russell and the type was annotated by Perry as "aff. V. officinalis L." in 1922. Gentry found the species in heavy olay soil of the coastal plain in Thorn Forest association at an alti- tude of 100 feet, calls it a "common annual", and records the popular name "alusame", It has been collected in flower and fruit in March and april. In all, 6 a specimens, including the type, have been examined Gitatione: . MEXICO? Sinaloa: H, S. Gentry 7011 (Ak--21727, Du— 35600, Mi, N)3; Rose, Standley, “& Russell Russell 14850 (N-type) . ra~Chihuahua: H. S. Gentry 557m mtry 557m (Ak-~20007). XVERBENA RYDBERGII Moldenke, Revist. Sudam. Bot. k: 1920. 1937. Synonymy: Verbena paniculato-stricta Engelm., Am. 46: 100. 184). Verbena stricto-paniculata Engelm., Am. ee Sci. h6: 101. 184h. Verbena hastata x stricta Rydb., Bot. Surv. Nebr. 3: 18. 1894. Verbena paniculata x stricta Engelm. apud Rydb., Contrib. U. S. Nat. Herb. 3: 173. 1895. Verbena bracteosa x stricta Rydb., Fl. Rocky Mts. 740. 1917. Verbena has hastate x stricta P. Patermann, Beitr. Zytol. Verbenac. 43. 1935. Verbena bracteosa x stricta Deam ex Moldenke, Revist. Sudam. Bot. k: 20, in syn. 1937. Verbena bracteosa x stricta Eggert ex Moldenke, Revist. Sudam. Bot. h: 20, in syn. 1937. 1937. Verbena stricta x has- tata Eggert ex Moldenke, Revist. Sudam. Bot. %. hz 20, in syn. 1937. Verbena hastata x stricta Gates, Fl. Kans. 190. 19h0. Verbena stricta x hastata Hill ex Moldenke, Suppl. List Invalid Names 10, in syn. 191. Verbena stricta x hastata Schneck ex Moldenke, Suppl. List Invalid Names 10, in syn. syn. 191. Verbena stricta x urticifolia Stevens (in part) ex Moldenke, Alph. List Invalid Names sora 1: 27, in syn. 1947. Verbena stricto-hastata Pat- terson ex Moldenke, Alph. List Invalid Names Suppl. 1: 2 Syn, 1976 Verbena stricta x hastata Gates (in part) ex Molden- ke, Alph, List Invalid Names Suppl. 1: 27, te syn. 1947. Verbena ta x stricta Anderson ex Moldenke, Résumé 365, in syn. . 1959. Verbena stricta x hastata Patterson ex Moldenke, Résumé 375, in syn. 19 1959. Verbena stricta x urticifolia Patterson ex Moldenke, Résumé 375, in syn. 1959. Verbena stricta x urticaefolia Eggert ex Moldenke, Résumé 375, in syn. 1959. Verbena stricta x urtici-~ folia Glatfelter ex Moldenke, Résumé 375, in syn. 1959. Verbena wticeefolia x stricta Engelm. ex Moldenke, Résumé Suppl. Li, ry’ : 18, in Wa sgn ee Verbena rydbergi Moldenke, Résumé Suppl. fae Bibliography: Journ. Sci. 46: 100--101 Siivoy:betc iter Rees fa 3: 16. 1895 Ryd, Contrib. U. S- Wat. Herb,”3: 173, 189 95; Rydb., Fl. Rocky Mts. 740. 1917; Kanda Seen Gaz. 69: Sh—71, fig. 62, 73, & Th, ple are fig. b—6. 1920 4 PHYTOLOGIA Vol. 11, no. 2 sa » 2ytol. Untersuch. Verbenac. 3. 1931; Rydb., Fl. Cent. N. Am. 678. 19323 Patermann, Beitr. Zytol. Verbenac. 19 & \3. 1935; Moldenke, Revist. Sudam. Bot. ly: 1980; 19373 oe Fl. ans, 190. 190; Moldenke, Prelim, Alph. List Invalid s 45— 48. 1940; Moldenke, Suppl. List Invalid Names 10. isis “yoldenke, ph. List Invalid Names 5, 47, 9, & 50. 1942; Moldenke, Known Geogr. Distrib. secoctnoses lee 1}, 6--9, 11, & 102. 19:25 Deam, ebel, Yuncker, & Fries Proc. Ind. Acad. Sci. 5h: 95. 1953 Moldenke, Castanea 10: 38. “13Ue; Moldenke, Bot. Gaz. 106: 159. 195; Moldenke, Alph. List Cit. 1: 9, 109, 110; i322, ie ais 1 > & . List & 1261. ot aes cua: Geogr. Distrib. Verbenac., [ed. 12-15, » 21, & 199. oe Moldenke, Phytologia 3: a é 131 (359), 3 "Bh (1950), and " 67. 19523 Moldenke in nee New Britton & Br. Illustr. ~ St dees bets 132. 1952; Moldenke, Phytologia h: ies. 19534 *Voldenke in Aes New Britton & Br. ernie r. Fl., print. 2, 3: 126, 129, & 252 as 16—-19 3 Moldenke, Résumé Suppl. 2: 1 & 2. 1960; Noldeok , Phytologia me 3, 18, & 21 (1962), and $3 3. 1962; J. D. Poindexter, Trans. K Acad. Sci. 65: 410, 411, 123, 415, & “ar, mt woldenke Résumé prsinc 7: 2. 1963; Beto & Cron wits Man F580 wen and 10: 209 isu, & 155, ; Bate Moldenk See Suppl. 10: (2 96l) and 1 a. e, ‘PP eee ustr. Fl., print. a> 132 (1952) and print. 2, @” oe. This is the natural hybrid between V, hastata L. and V, stricta arb A aoe NN Vent. which occurs commonly where the r 8 of these two species over in central U. States —* south anada, It rem sembles V. hastata in habit, but its lea are more reticulate- 201-57, which are all typical = the hybrid and on which the polle® fertility was 21, 12, and 30 percent respectively. His 191-1 shor 1964, Moldenke, Monograph of Verbena ns ed 33 percent pollen fertility. Schwenke phar reports the hap- loid chromosome number for this hybrid 6. Poindexter (1962) compares this hybrid with its two parents as follows: Leaf index: V. stricta = 1.32.1 (average 1.8), xV. rydbergii = 1.7-—-3.5 (average 2.6), V. hastata = 3.2—5 (average 4.3); leaf-shape: Ve. stricta = elliptic to ova ovate, xV. rydbergii = broadly ovate to lanceolate, V. hastata = = lanceolate; leaf-base: V. stricta = broadly at Gallia to © sessile, xV. rydbergii = broad to narrow attenuate, some hastately lobed, V. . hastata = rounded & petiolate, hestetely lobed; leaf-apex: V. stricta = acute to tel xV. rydbergii = long-strigose to hirsute, V. hastata = | = strigose; nutlet length length: V. stricta = 2.2--2.9 (average . 2.6), xV Ve = 1.5=-3 (average 2.5), Ve hastata = 1.5—2 (average 1.7); mark- ings on back of nutlets: V. stricta = deeply ribbed, xV. rydbergii = deeply ribbed to faintly ribbed or striate, V. hastata = smooth to faintly striate; pollen fertility: V. stricta = 50—I100 percent (average 90 percent), xV. rydbergii = 3--92 percent (average 32.7 Percent), V. hastata = = 30--99 percent (average 79.8 percent); corolla-tube length: V. stricta = 4.5—5.5 (average 4.9), x¥. ryd- bergii = 2.55.2 (average i), V- hastata = 23.9 (average -365)5 calyx length: V. stricta = 3, Me Egy pone (average .3), xV. = 2.4—l.3 (average 3.2), V. hastata = 2.4-—3 (average 2.7); peti- Ole-lateral vein measurement: V. stricta = 3—6 (average 3.8), xV. rydbergii = 5—22 (average 9.7), V. hastata = 12--25 (average typical hybrid with hardly any fruit, on the same Lower leaves arehastate, His 509-19 had be “recast Sellen Sort iity and its inflorescence is almost typical V. stricta, but the leaves are hastate. G. E. Morley 805 and 105l-2 are the typical hybrid Pastures and overgrazed pastures, brushy overgrazed pastures, o- Pen woods, creek bottoms, and dry ponds, along roadsides or low erassy roadsides, creeks, and sloughs, at the junction of dry and noist bottoms, near low pasture draws, and on badly er or Prairies, f1 — cg = July to ee to 7hO feet altitude. ra on sandy kunk-cabbage ( 8 foetidun) oe Ahles & iipin cs it "freqent naith both parents" in pastures; Ahles found it "in marshy area around pond, with V. hastata, V. stricta, 116 PHYTOLOGIA Vol. 11, no. 2 and V, urticifolia"; Goodman collected it growing with V. stricta and ¥. halei Small; Isely found it "scattered in pastures", "while Lathrop saw it on "roadside bank agar e . prairie pasture with limestone surfacing in the pasture" and in a "waste place border- 5g a prairie park area, weedy ground," Tage enknecht found it on weedy rocky southfacing bluestem prairie ee Maceo Neeattered between parents in low moist areas in pasture” and "in ce in prairie", while sore pty a “common in local colonies in wet alluvium of floodplain" in Kan Miss Gaiser reports that “his fae tae NO. 1608re] is quite abundant about this region and stretches over r qiite an an area at Point Edward [Ontario] where the river front road sae he tye dump back of Metal Works, in sandy soil". For her a shi 29151 and L. L. H. Pammel s.n. (Hamilton, Sept. 8, 1918] & s.n- (Granite, Sept. 1, 1920] are mixtures with V. stricta — Gndica- Main pag the close proximity of at least one of .e of the parent specie The type of the hybrid, which is based on the Ve paniculato- stricta of Engelmann, was collected by George Engelmann on the banks of the Mississippi River at Saint Louis, Missouri, in July, 1843, and is deposited in the Torrey Herbarium at the New York Botanical Garden. Verbena bracteosa x stricta Deam is based on 4 plant collected by Charles Clemon Deam (no. 20603) in Washington County, Indiana, and deposited in the herbarium of Purdue Univer~ sity; v. hastata x stricta Anderson is based on a collection made by Jacob Peter Anderson in Decatur County, Iowa, on August 25, 1 ane is deposited in oe herbarium of Iowa State Collearh.n: Le s based on a collection made by H ich cal Daniel | Eggert Saint Louis —— ere’. on A = 10, 1877, and is pitted ey: in the herbarium of t cenege ton, Chicago, ge Prisciy Tilinois, on August 10, 1898, and is deposited in the herbarium of the University of Illinois stricta x hastata Patterson and V. stricta x urticifolia Patterson are both based on a collection made by Harry Norton se gen in the vicinity of Oquawka, Henderson County, Illinois, in and is deposited in the herbarium of Iowa State College; "Ve serie ta x hastata Schneck is based on a collection made by Jac Schneck in moist soil — the Utter farm, Mount Carmel Wabash County, Illino June, is raps in the herbar- ium of the University of tisenle; V. stricta x urticaefolia Ee- 196k Moldenke, Monograph of Verbena 117 gert is based on an Eggert collection from Saint Louis, Missouri, e in August, 1875, and deposited in the herbarium of Iowa state College; ‘7 stricta x urticifolia Glatfelter is based on a ae eae by Noah Miller Glatfelter near Saint Louis, Miss- £ 2308, klahoma, on May 26, 1913, and is deposited in the herbarium of the Univer- sity of Oklahoma; V. stricto~hastata Patterson is based on a col- lection made by Patterson near Oquawka, Henderson County, Tine Ois, in August, 1873, and “prea itas in the United States National Herbarium; V. stricto-paniculata Engelmann is based on a collec- tion made by Engelmann at a Louis, Missouri, in July, 1842, and is deposited in the Torrey Herbarium; and ‘ urticaefolia x stricta Engelm. is based on a collection made by Engelmann on the Ridge prairie, Vermilion County, Illinois, in August, 1860, and deposited in the Jesup Herbarium of Dartmouth College. It is worth noting here that the V. bracteosa x stricta of Britton, of Clothier, of Palmer, and of Schnack are xV. perriana Moldenke; V. hastata x stricta of Allen is V. stricta f f. albiflo- ra Wadmond, of Elmore is V. hastata L., and of Ravenel is 3 Stylo- don cameus (Medic.) Moldenke; V. stricta x hastata of Allen is Ve stricta Vent. and of Gates (in part) is ve . stricta Vent. and xv. “moechina Moldenke; Vv. stricta x urticaefolia of Gates, of Mackenzie, and of Pond are xV, illicita Moldenke, of Letterman and of Pammel are xV. siesta Moldenke; and V. stricta x urtici- = of Britton, of f Bush, ush, of | Eggert, of Gates, and of Schneck are xv ve aatcite Moldenke, and of Stevens (in part) is xV. illic- - perr XV. perrians, ita and "Thumesess collectors have noted the possible hybrid nature of Specimens and have commented on it. For instance, Hayden 3006 is labeled "Verbena hastata L. x V, stricta Vent.", while L. H. Pam- mel s.n. Lenaneeiens, fae Lee the notation "a possible _ rid between V, hastata and V. stricta." Material of this hybrid has been an misidentified and distributed in herbaria under the names V, hastata L., V. hastata Michx., V. hastata stricta Rydb., V, hastata x urticifolia Eggert, x¥. illic- ita Moldenke, V V. officinalis L., V. stricta Vent., V. stricto- urticaefolia Engelm., and V. ee en Engelm. On A. E. Allen s.n. en. (Talmage, July 11, 1091] and J. L, Sheldon s.n. ss July 21, 1900), identified as my. stricta x x hastata", and © E. W. Lathrop 963, distributed as xV. rydbergii, are also fall uho, Loomis typical v. stricta, a and the S. V. Fraser | omis s.n. (Alton, 118 PHYTOLOGIA Vol. 11, no. 2 Oct. 1938], and Muck 82, cited in Gates, Fl. Kans., as "V. stric- ta x hastata", are also typical V. stricta. A. E. Allen s.n. [Peru, Aug. 2, 1893], distributed as "V. hastata x stricta", a", is actually V. stricta f. albiflora Wadmond The cytology of this hybrid is iaeisase by Kanda (1920), who reports that Dr. Chamberlain is authority for the statement that initely that they were not mutants, but were hybrids. He further — that pears forms occur, but I have seen albinos only of ach parent, not as yet of the’ hybrid. Gates (19h0) records the hybrid from Cloud, Osborne, and Shaw- nee Counties, but I have not as yet seen material from those counties. He cites Clothier & Whitford s.n. [Elk Co., Aug. 20 & 21, 1897] and Kellerman Bon. [Galena, jalena, July 6, 1887] as V. stricta x hastata; Harman 0 as V. hastata x stricta; and J. B. Se Norton Son. n. (Webber, Sep Sept, | 23, 1895] as Vv. hastata. xVerbena “rydbergii has been recorded from Banner County, Nebraska, on the basis of 4 sheet of Rydberg 1564 in the Britton Herbarium which has a print- ed label reading "Banner er County", but the longhand inscription says "near Mullen" (as on the other sheets of this number), which is in Hooker County. Apparently an incorrect printed label was used for this particular en. In all, 193 herbarium specimens, including the Pag material of all the names involved, have been examined ; aoe ONTARIO: Lasbton Co.: Gaiser 1608PE ( (Gp, Gp, Mm), TOPE (Gp, Gp, Gp, Mm); Gaiser & Montgomery 2232PE (Gp, Gp, GP, Mm), 3233PE (Gp, oP, Dicgen lin). ILLINOIS: Bureau Co.: Ahles 4176 (Ur); Winterringer 8008 (Il~—37299). Champaign Co.: Ahles & & Gil- pin 8153 (Ur), ee (Hi—~103235, Ur); G. N. Jones 12525 (Ur). Son Bee Co.: Eo J . Hill 160/1898 (Ur, Ur, Ur)3 Lads Ne Johnson SM» (Evanston, 9 3-3-89) ( (Ur—17)62). DeKalb Co.: P. B. Whitford a 156 (I1--3392h). Fulton Co.: Winterringer 886) (I1—38680) « Hancock Cos: L. H. Pammel s.n. (Hamilton, Sept. 6, 1918] (Io--9l058). Henderson Co.: He H. N. Patterson s.n. [august 1872] (Al), ‘= [near Oquawka, August 1873) (W (W--1323127), sen. [Oquawka, (Io—92216, Io—92217), s.n. [vicinity of Oquawka] poet W--1323128, W--1323129, W--1h16723). LaSalle Co.: Thone 87 (Ur)- Macon Co.t A. R. Voldenke 819 (Lw, Ut, Z, Z, Z, Z, Z, Z)- Peoria Co. V. H. Chase 566 (Ur). Pike Co. ie Davis 32u1 (Ur), a [July 18, 1914] (Se—1h93k). Sangamon Co.: G. D. Fuller 6406 (Ii- peldogl gt F C. McGregor 1117h etipas). Vermilion Co. Boge? Ridge prairie, Aug. 1860] ( Dt); Storm s.n, [July 2, T5h9) Ci ie, Ur). Wabash Co.: Schneck s.n. [Mt. Carmel, Sep. 1, 1887] (Ur), son. (Mt. Carmel, June 1900] (Ur). Winnebago Co.? EB. W. Fell 5132h (1138586), 51328 (1138587), 51329 (m1-- 38609), 51u0l, (I1—38332). County undetermined: F, Brendel 8.0- {Illinois, 1573] (W—71993). INDIANA: Pulaski Co.: Friesner 22312 196), Moldenke, Monograph of Verbena pale) (N). Washington Co.: C. C. Deam 20603 (N, N, Pu). County unde- termined: Drew s.n. {northwestern Indi Indiana] (Ca--67759). IOWA: A- dams Co.: Isely 5706 (Gg--392102).- Cherokee Co.: L. H. Pamel s. ne (Cherokee, Oct. 17, 12h] (Io--11)907); Pammel GS ace Son. [TGoodell, July 12, 1922] (Io--1057h6). Clay Co . Hayden 3003 (To~1,900h) , 3005 (Ca=~999282, W--1835171), 3006 (Ces ag Io- 149007, N, W—1835172). Dallas Co.: L. He Pammel son. (Dawson, Aug. 16, 1918) (Io-~95629). Decatur Co.: 3 J. P. Anderson 8.n. {Aug, 25, 1904] (Io=-52107). Dickinson Co.: Shimek s.n. sen. [Aug. 8 1916] (N, Ur). Hancock Co.: L. H. Pammel s.n. (Garner, Jul. 27, 1918] (Io--96114). Hardin Co.: L, H. Pammel s.n, [Steamboat Rocks, 9-12-12] (Io--52669, Io--52670), Son. (xehoes, Oct. 5, 1924} (Io~~116221). Lyon Co.: L. He Pammel s - (Granite, Sept. 1, 1920] (Io--97830). Madison Co.: A, Re ey watieaks 942 (Lw, Ut, Zz). Mahaska Co.: A. R. Moldenke 960 (IL). Muscatine Co.: Shimek Sn. [Aug. 21, 1915] (N, Ur); Somes 370k (W--672)00). Palo Alto Cou: L. H. Panel sen. (Jul. 18, 1920] (Ilo—97793). Poweshiek Cow: Conard s S.ne [8-29-1927] (Ok—-10362) 5 Me Ee Jones S.ne (Grin- nell, Aug. “hug. 1877] (Du—-176622); A. Re Moldenke 966 (Im). Story Co.: L. H. Pammel s.n. [Ames, 1902] J (Io—-81968). Warren Cost Ae R. Moldenke 951 ( 951 (Lw). Winneshiek Co.: Tolstead s.n. [Decorah, July 23, 193h] (Io—=14h28h). WISCONSIN: Grant Co. A. R. Molden ke 997 (Lm); L. H. Pammel s.n. [IMuscoda, Aug. 1928] (1o—137607) « Rock Co.: Ee We Fell 57-73%, i in part (Ws), 57-740 (Ws); Skavlem Sen, (Janesville, July 23, '89] (Ws). Trempealeau Co.: Fassett & Wilson 9776 (Ws); L. H. Pammel s.n. [Trempealeau, 7-20-82] (Io 10815). x MINNESOTA: Hennepin | Co.: Sandberg 605 (Ka). KANSAS: Allen Co.: R. L. McGregor 13230 (Lw). Atchison Co.: A. S. Hitch- cock s.n, _ itektes Co., Oct. 1896) (Ka). Barber Co.: Horr & on. [July 6, 190] (Iw); R. L. McGregor 14598 14598 (L¥)3 Je D. Poind Gaiselactes 201-57 (Lw). Brown Co.: Clothier & mithtord aos. Touly 29, 1897] (Ka). Chautauqua Co,: We H. Horr s.n. [July 6 1930] (Le), Cherokee Co.: Kellerman s.n. [Galena, Jul, a 8, 1887] (Ka), coe ake W. H. Horr O91 (Lm). Comanche Co.: Bites McGregor uglas COs: Je D. Poindexter 191-1 as arate (Lw), Fon) (Lm), 229-8 (Lw), 229-19 (Im). Elk C Clothier & Waitford s.n. (Elk Go., Aug. 20 & 21, 1897] (xa). Fl- lis Co.; E. Runyon | Runyon 280 (Dm). Jackson Co.: By a Wagenknecht 3788 (Lm). Jefferson Co.: Harman 0 (Ka). Jewell Co.: J. B. S. Norton S:Re [6/14/30] (Iw). Marshall Co.s A. S. Hitchcock 8.0. (Marshall Co., Dec. 10, 1896] (Ka). Montgomery Co.: R. L. MoGregor 14395 (Lm). Nemaha Co.: A. S. Hitchcock e.n. [Nemaha Co., Dec. 11, 1896] (Ka). Pottawatomie Gost Clothier s.n. (St. George, "aug. 28, 1896) 120 PEITOLOGIA Vol. 11, no. 2 (Ka). Reno Co.: R. L. McGregor 12511 (Lw). Republic Co.: G. E. Morley 805 (Lm), B30 | aa 1054-2 (Lw). Scott Co.: V. Le Harms 1108 (Lw). Woodson Co.: E. W. We Lathrop 963 (W-~2235188), 1376 316 (H). County undetermined: Kleeberger s.n. (Gg--31419). MISSOURI: | Co.: Spellman $.311 (Lb--5152h). Saint Louis Co.: Eggert s.n. [19 Aug. 1877] (Cm, I); Glatfelter s.n. {near St. Louis, 8-11-9h4] (Mi). Saint Louis: Eggert 5320 (N), s.n. (St. Louis, Aug. 1875] (c, > Pettia Gatee s.n, (St. Louis, ess 1875] (an, B, het Ca-- 181598 , Ca—-25201, Cm, Du—202132, Gg—183217, Io--7986 6892, N, Po 192592, “H-75955), gon, sae Aug. 1877] (A) Engel- mann s.n. [July 1939] (Br), son. [St. Louis, July 1842] (Pr, T), sen. [banks of Myssissippi, St. Louis, July 183] (T—type), 5.n. TSt. Louis, Aug. 17, 1859] (Dt), s.n. [St. Louis] vib Glatfelter o.ks tet; Louis, 6-3-92] (W-~3096L3)) ne aes ARKAN- SAS: Benton 003 Heacox s.n. [July 20, 189] ( oon oy. Marion Co.: Demaree 29151, in| in part art (Au-~122726) « pede Cedar Coe: F. E. Clements 2663 1/2 (C, W—719l3). Hooker Co.: Rydberg 1564 (N, N, W—210369). Lancaster Co.: J. L. Sheldon s.n, [Lincoln, | July 20, 1898] (We). Thomas Co,;: Krautter s.n. {Halsey, July 23, 1907] (Up~—h57h0, Up--L57h1, Up—LE7hS, Up--ho7h8) « a Ge W. Stevens 1791 (Ur, W--589685). Marshall Co.: G. J. Go Sh85 (ok, Z). Ottawa Co.: Schendel 202 (St—17571)3 G. W. 7 ai vens 2308, in part (Du--66376, N, Ok—21028), A.2308 (St-—9278) « pat undetermined: G, W. Stevens ,032a (Ok). LOCA LOCALITY OF COL- : Engelmann s. 8.n. . (Dt); 3 Short s.n. [banks of LECTIO the Babéah} (Pr). VERBENA SAGITTALIS Cham., Linnaea 7: 259. sr Sat *: ad Cham., Linnaea 7: 259. re fo Ha Nom. Bote, ed. 2, 2: 750. 18h1; D. Dietr., Syn. Pl. 3: 603. 18h3; Walp., Repert. kh: 18. 1845; Schau. in A. DC., Prodr. ll: 53. 18h7; Schau. in Mart., Fl. Bras. 9: 190. 1891; Briq. in Engl. & Prantl, Nat. Pflanzenfam. (3a): 147. 1895 Jacks. in Hook. 2. ; Laas 4 ° Ace 2: 1179. 1895; Briq., Arkiv Bot, Stockh. 2 (10): oo. 190k idenke, Known Geogr. Distrib. Verbenac., [ed. 102 RORS and [ed. 2], hah & ae aes Moldenke, ae pare i aed 29M Moldenke, Résumé 111 & 7k. 1959; Moldenks, Résumé . “he i 962; reopen ap tenea 9: 14. 1 with creeping rootstocks; stems gusty babes rush-like, tetragonal, with an aphyllous aspect, con- tracted at the nodes, the angles subalate, the margined edges and surfaces scabrous, the surfaces canalic culate, scrobiculate in pre the depressions suborbicular, puberulent; internodes long- te, ed with small pits or depressions between the stria~ rg leaves small, sessile, less than 2.5 cm. long, ovate or triangular-oblong, acute at the apex, entire or subserrate along the margins, 3-nerved, strigose-scabrous; spikes cylin dense-flowered, subternately aggregate, becoming almost 2. 5 om. 196k Moldenke, Monograph of Verbena 121 long, thick; bractlets lanceolate, short, half as long as the ca~ wx © cuminate at the apex, the margins hirtulous; c long, with elevated ridges, the angles hirtulous, the teeth subu- late; corolla-tube scarcely longer than the calyx, t small; cocci about 2 mm. long, ferruginous on the back, slightly e edges. The type of this little~knowm species was collected by Fried- rich Sellow in "Brasilia meridionali" and was deposited in the herbarium of the Botanisches Museum in Berlin, now destroyed. Un- fortunately, it was never photographed, as far as I am aware, but chau ge Briquet (190k) ares it stb and nies tg =f aca pe t from V. ephedroid i Briq., and Junellia pseudo~juncea oF Gay) esenieg 5 aur (1852) points out that V. sagittalis is closely related to V. ala~ ta Sweet, but differs in its inflorescence, its bigger spikes and calyxes, in the character of the fruit, and most especi in the serobiculate stems. eitiars (18455 classifies it in his Sec- tion Verbenaca, Subsection Inermes, and Group Junceae, with 7 other species. It has been collected in anthesis in March. Only herbarium specimens have been examined by m Citations: BRAZIL: Rio Grande do Sul: Sobwacke IIT.299 (Ja, N). VERBENA SANTIAGUENSIS (Covas & Schnack) Moldenke, Phytologia 2: pst 196. onymy: Glan antiaguensis Covas & Schnack, Revist. Argent. Agron, 11: cor figs 2. hee Verbena santiagensis ip hnack, Revis ts Argent. Agron. : 92-—— 9h & 97, fig. 2. 19h pais & Darwiniana 7: 86 & 88. 19453 Schnack & Covas, Darwiniana 7: Tl, 73-1, & T1179, fig. 2 & 5. 195; schnack & Covas, Revist. ia Agron. 12: 222, 22h, & 228. 195; schnack & Gonzalez, 12: 285— Revist : 288, fie. 1 & 2, pl. 15. 1953 Moldenke, , Pigtologsa 2: 150 (1916) and 2: 348. 1947; Moldenke, Alph. List Names Suppl. 10. 19475 Moldenke, Alph. List Cit. 3: ore & & The 19195 Holdenke, ps $3 1962;” Moldenke, Résumé Suppl. chy (1962 ) and ‘a ga | Pohinaeeee =e} pen 9: 45, 6 10, & ci (1963), 9s hoo 135 + 133,186, 295, & 301 (196k), and 11: 52. 196k. 122 Pa rer Ol oer kk Vol. ll, no. 2 TNlustrations: Covas & Schnack, Revist. Argent. Agron. 11: 93, vas Perennial ing at the Pecyrss nodes, ascending toward the apex, cylindric- quadrang bescent, the hairs simple, rather rigid, 0.5 m. long, igs) retrorse; leaves decussate-opposite; petioles long, 0. outline, 2=-5.5 cm. long, 1.3--3.5 cm. wide, pubescent above with al, spicate, solitary, abbreviated during anthesis but sometimes elongating after anthesis; bractlets lanceolate, more than half the length of the calyx, ciliate along the margins, with subap- pressed hairs on the back; calyx tubular, 6—7 mm. long, pubescent with simple oblique hairs, rarely with subsessile * glands on the angles; corolla hypocrateriform, pale-lilac or purple, laxly pubes cent at the apex of the tube outside as well as on the lower sur~ face of the upper lobes and at the base of the lower lobes, the lobes all glabrous on the upper surface; stamens typical, the 2 upper ones with subcylindric glandular appendages, the thecae ° mm. long; cocci subcylindric, 2 mm. long, rounded at the pug truncate at the base, reiaslite at the middle on the upper face of the back; chromosome number: 2n = 10. The species is based on a specimen collected by Benno Julio Christian Schnack (no. 2111) from material cultivated ny Capital, emp as t ships are with V. dissecta Willd. and V. laciniata (L.) Briq- They distinguish - it from its immediate relatives as follows: 1. Stems erect or suberect, not rooting. 2. Entire plant covered with a dense pubescence of simple hairs mixed with glandular ones; leaves triparted-pinnatilobsd'. V. peraicit . 2a. Entire plant covered with a sparse pubescence of only sim- ple hairs; leaves pinnatisect.........++++0+sV. mendocima- la. Stems covering. rooting at the base, ascending at the tips. 3. Cocci 2 mm. long. lk. Bractlets one-third as long as the calyx; spikes not elon gating after anthesis V. dissecta- ha. Bractlets more than half the length of ee calyx; spikes elongating after anthesis V. santiaguensis- 3a. Cocci more than 3 mm, long. 1964 Moldenke, Monograph of Verbena 123 5. Corolla externally glabrous; glandular appendages of the connectives subcylindric, hardly visible ap aegiyn or included; pubescence formed of appressed ha ecu sew ee v. laciniata. Sa. oes externally pubescent; glandular appendages of the mnectives much compressed, exserted; esc formed at erect obli Crovetto & Rojo (1957) describe the plant as "Planta ; perenne s 8, radicantes cies. The only etal name recorded for it is "verbena", It has been collected in anthesis in April, July, August, and December. ¥. cheitmaniana Moldenke and in part V. tenera Spreng. In all, 5 herbarium imens have been examined by m Citations: URUGUAY: Lorents s.n. [Concepcién, eae "(8). : Corrientes: A. T. A. T. Hunziker 5528 (N). La Rioja: F, A. Tepe Sarak 19aré9), (N). CULTIVATED: New York: H. N. Moldenke N) VERBENA SCABRA V. Vahl, Eclog. Am. 2: 2. 1798 [not V. scabra Marnock, 180, nor aun, 1825]. Synonymy: Verbena polystachia H.B.K. ex Moldenke, Suppl. List Invalid Names 10, in syn. 191. Verbena urticifolia Jepson, Fl. Calif. 3 (2): 380, in syn. 1943 [not Ve V. urticifolia L., 1753, nor Gray & Small, 1962]. Verbena polystachya Jepson, Fl. Calif. 3 (2): 380, in syn. 19)3 (not V. polystachya H.B.K., 1818]. Verbena polystachys H.B.K. ex Moldenke, Alph. List Invalid Names Suppl. 1: » in syn. 1947. Verbena urticaefolia Griseb. ex Alain in Leén & Alain, Fl. Cuba 4; 281, in syn. 1957 [not V. urticaefolia L., 12h PHYTOLOGIA Vol. 11, no. 2 1811}. Ocymum nervosum Vahl ex Moldenke, Résumé 322, in syn. 1959. Verbena scabra Gray ex Moldenke, Résumé 37), in syn. 1959. Verbena urticifolia Griseb. ex Moldenke, Résumé 377, in syn. 1959. Berbena scabra Vahl ex Moldenke, Résumé Suppl. 7: 7, in a, 5 ag Bibliography: Vahl, Eclog. Am. 2: 2. 1798; Steud., Nom. Bot., 2: 79 Schau. in A. DC., Prodr. 11: 546. 1847; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 1179. 1895; H. H. Rusby, Mem. ee Bot. Club 6: 106. 1896; J. K. Small, Fl. Southeast. U. S., ed. 1, 10 (1903) and ed. 2, 1009. 1913; N. L. Britton, Ms Bermuda 310. 1918; Britton & P. Wils., Scient. Surv. Porto Rico 6: 138. wane Woldenke iat see Fai Southeast. Set 10. 1930; Stapf, rry, Ann, Mo. Bot. Gard. 20: 2hh, 25, aur, oe "210, aT 21S 278, & 356. 1933; J. K- Small Verb L Fe qd : ora 371 & sap 190; neyo Suppl. List Common Nanes 18 valid Names 160. 1945; Murrill, Guide Fla. Pl. 6. 1945; Moldenke, Phytolo- gia 2: 117. 1945; Moldenke, Am. Midl. Nat. 32: 576. 1945; Mol- denke, Castanea 10: 37. 19h55 W. H. Hodge, Journ. N. Y. Bot. qd 7; Fe oy tae Moldenke, Phytologia 2: 328. 191j7; Moldenke, Castanea 13: i, & 1h. 1948; Moldenke, oo ia l: a ia Molden- od ’ 692, ag 72 747, 756, 760, 773, 776, 778, 779, 787, 789, 73h, 796, 798, 825, , oo e = oo oo ~ . bad Le +] 3 . ‘Oo w = = 2) 2 1110, 1139, 63, 1166, 1170, 1176, 1178, 1179, 1181, 1182, 1192 1219, 1223, roest-3g90 1239, 120, 12h5, 1258 1256, 1255, 1259, 1288, 1289, & 1301. i9hi9; wokdenks, Known » (ed. 18, 493 2h, 26, 28, 33, 1, isto, 6 & 199. 1949; Moldenke, Jon Be ia 3: 196k Moldenke, Monograph of Verbena 125 3. 1993 powrengs ina sh eae Man, Bot., ed. 8, 1209, rer » fig. 1462. 1950; Castanea 16: hi3e 1951; usidoake, Prytologia. 38 50 (i983) efor ey eg 1952; Moldenke in Gleason, ie i & Br. Illustr. Fl., print. 1, 3: 126—~129. 1952; Mol- ne a hy he 166. 1953; Thorne ig Midl. Nat. 52: 313. sche * Moldenke Inform. Mold. Set 18 8 Spec. [4]. 195h; —— Rhodora Piece 288. 1955; Moldenke, Inform. Mold. Set 51 fa 3 1956; abate & Al ain, Fl. Cuba hi: 281, fig. tas "1957; ar pi in a New Britton & Br. Illustr. Fl., print. 2, 3: 126--129. 1958; Woldenke, Résumé 9—13, 15, 22, 23, 30, 32, 33, bo, ig, 53--55, 57, 59, 322, 372, 37h, ie tru. 19595: Meldiacien, sumé pel. ds 2's & i, (1959) and = ldenke, a 8: 143. 1961; Moldenke, Psa Suppl. 3: 3, 6, 7, &9 (362), ne 1-3 (1962), $: 1&2 (1962), wt a e 3 (1983 ), and 7:1 & 1 a oe Gleason & Cro « 579 & 580. 1963; ae Phytologia 8: 783, 48 487-489, “igh 0369), om Sout bi 93s 1655. £199 (1963), and 10: 140, bd, & Shreve & Wiggins, Veg. & Fl. son. ae = ee I ations: Marnock, Floricult. Mag. 5: pl. 54 [in color]. 1840; N. L, Britton, Fl, Bermuda 310. 1918; Fernal Gray, less scabrous and somewhat paler abet and also hispidwlous a- acuminate, a Roms as long as the calyx, hispidulous; calyx to 2mm. long; c a hypocrateriform, varying from blue, bluish, or pale-bive - le Coan purple, purplish-white, lavender, Violet, lilac, pale-rosy, rosy, pink, pinkish, or even white, its tube scarcely, if at all, longer than the calyx, usually vio with white hairs in the throat, the limb 2--3 mm. wide, the lobes usually light lavender-pink, obtuse at the apex; anthers yellow, Slandless; stigmatic surface of the pistil midway between 2 almost equal obtusish sterile lobes; fruiting-calyx 2.5--3 mm. long, o- void, hispidulous, with the eriginge unequal lobes acutely conni- vent, diverging f spike by an angle of 45 more; cocci Spare roel 1—1.5 m. 1 Striate, re te 2-striate on the back, the striae t ° iculations, the c s- Sural faces extending to the tip of the schizocarp, meeting sharply at right angles, muric t. Philipp Benjamin von Rohr (no, 35) "eelidabiaiie "in America meridi- 126 PHYTOLOGIA Vol. 11, no. 2 onali", and is deposited in the herbarium of the Universitetets Botaniske Museum at Copenhagen. This tt Benge nem and dis- damp or paige J land, low moist or marshy ground, open swamp ae open or wet ground, cultivated ate moist or ’ moist sandy ground, moist rich soil, and rich on muck, in moist or wet andy bogs, dry sandy 1 ds, low pinelands, satel: thickets, and moist ground at the And of marshes, on river- and streambanks, open moist streamb : dbanks and lake shores and river bot~ toms, sandy beaches and the margins of rivers, about salt lagoons, along roadsides, creeks, and rivers, at the edge of hammocks, the low woods, the edge of waterholes, and at highwater-mark on shores. Collectors report it from marshy areas, low carey sts a ea 5 eager, brown marshy swamps, low or sandy woods, , damp soil near springs, salt marshes, grassy hae nt eehincsan Pres: among dunes and low areas in open forests back of » among rocks, in hotel yards and low io? in true shale e woods sh near , and in low places or swanpy country in opie at altitudes of 5 to 7800 feet, flower- ing and fruiting from February to ie Texas it grows * mostly rich soil of low ground, marshes, t state, from Newton and Jefferson Counties west to Uvalde and sind Counties. Head describes it as "scarce" in Jeff Davis ° In es it is said to inhabit the i ag vse 9 Life zone according to C. B. Wolf, while Johnston regards it as "introduced! In Louisiana it is they cone sae defies cor PN Quercus and Stenotephran" on the banks of lagoons. On Sapelo Island, Gé~ orgia, an found it "in [the] open at side of roadway around eee lake fed by artesian springs". cae Palm Beach pecgorh £ in everywhere County it is "common in the everglades", dle ve "ep O'Neill. Rankin reports it "common in fields" in’ Bermuda, Ekman says "com~ mon in swamps" in the Dominican Republic, and Wright says "intro-~ duced on roadsides" in Las Villas, Cuba, Webster & Proctor found it to be "common in the drier areas" of J. quent pea area recently drained and burned". yurrill fishes) desortbie 2k as having small, pink or ate Sr, and er ruvian — a statement which is ridiculous, since neither V. carolina L ; nor V. scabra Vahl (both of which species have a nV. polystachy® 1964 Moldenke, Monograph of Verbena 127 in their synonymy) grows in Peru nor ever forms a cushion plantt H. M. Pollard says for his collection of V. scabra from north of Veronica Springs, Santa Barbara County, California, made on ? om Santa Barbara ~- this from a favored collecting spot probably uae off and on for the past 100 years. When did the plant ar "The Lindheimer 1077 collection cited below may be from either Comal or Comanche County, Texas, since its label merely states: ee Spring: New Braunfels, etc." It was cited in Lundell, Fl. Texas 3 (1): 23 (1942) from Bexar County in error. Dr. S. W. ep believes that the Bigelow s.n. cited by Perry from Texas was not collected on the present Leon River as indicated on the label, but oe the present Leona River in either Frio, Uvalde, Zones, from North Carolina to Florida and the Greater Antilles, West to oe Seer att: and south to Baja California, Coahu- and Nuev cttpers (1845) eee ee this species in his Section Verben- &ca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Secondary Subgroup Hol ophyllae, along with 22 other species. Schauer (847) places it doubtfully in synonymy under V. urtici- folia L. It should be noted here that the V. scabra of Marnock is ac actually V. rigida Spreng., while that of f Muhlenberg is Phyla lanceolata (Michx.) Greene. Bailey (1935) reports that the seeds of V. scabra are handled by Kew. Thorne (1951) reports the spe- ugherty and Lee Counties, Georgia, but I've not as yet seen substantiating specimens. Bogusch 1235 is a mixture mith V. xutha Lelm., while Fernald & Long 12453 is a mixture with xv. engelmannii Moldenke, The C. C. Deam 65942 collection shows fasciation, whi while Ekman 933 has one spike fasciated~flattened for tnehas: One of the GC Cory 38068 sheets in the University of Texas herbarium is inscribed "H. N. Moldenke 33068" in error. Rugel 639 639 in the United States National Herbarium has an origin- al label inscribed "ad fluv. St. Johns", to which someone has added "Ins. Cuba", obviously in error, since the Saint Johns Riv- er is in Florida. The Sapelo cams cited below is politically in MeIntosh County, Georgia. Braunton says "I found what is ev- idently polystachya x prostrata but ' Ts ia ik gene eenies has it asia ennai No. 520 and it 1 nlgheasee tit Small (1903 & ey uses the name V. po hya H.B.K. for V. scabra, but this name is a synonym “iC Yearotina L Comm names ep, Bs reported for ve 3 ae are "rough vervain", aa trerbena sons » and te vervain", White-flowered specimens (perhaps worthy, of a form name?) are Ahles & Leisner 32007, H. A. Allard 128 PHYTOLOGIA Vol. li, no. 2 8270, W. Harris 9937, and Radford 28495. ~~ Material of | Vv. scabra has been misidentified and distributed in herbaria under the names V. carolina L., V. carolinensis Michx., V. caroliniana L., 7. halei Small, He hastata L., V. lite toralis H.B.K., V. officinalis L., L., Ve BS pe cha H.B. HeBoKe,y Ve poly- stachya H.B.K., V. polystachys H.E.K., V. urticaefolia L., V. ur- ticifolia L., Vv. urticifolia x hastata Gates, and Priva echinata A. L. Juss On the other hand, the Kearney & Peebles 10506, distributed as V. scabra, is V.c carolina L.; Rav Raven, Lewis, & Thompson 12180 is ve menthaefolia | Benth.; and Ahles & Duke 18221 is V. urticifolia var. leiocarpa Perry & Fernald; W Wiggins 9157 is V. “orcuttiana Perry; T. S. Brandegee s.n. [Cafion Salado, June 1, 1897] and Edw. Palmer cae "sisted as V. polystachya, are V. peontinetolia’ Benth.; Ewan 1770) and I. L. Forbes s.n. [Colfax, VIII-20-27] are xutha Lehm.; and F. S. Blanton 6671 and O'Neill 6200 & s.n- [1 1/2 miles south of Leesburg, Sept. 1 are Hyptis mutabilis (A. Rich. ) Beige i Ghataniddene., aS) Se} ate in Rhodora 38: ]l2 3 (1936), says that in Vv. sca~ bra "the Bisicced surface lies between two almost equal sterile lobes....the fruiting calyx is strongly ae oat a the nutlets pci above". He cites Fernald & Long 6863 from Surry Coun= , Virginia, deposited in the Gray | Gray Herbarium "tise (1933) cites the following 65 additional meres not as yet seen by me: NORTH CAROLINA: a Hanover Co.: Randolph & Randolph 1012 (G). FLORIDA: Broward Co.: Small & Carter 1072 .(F)s Collier Co.: O'Neill s.n. (Everglades, "2 hug. 1925] (B)+ Colum- bia Co.: Quaintance s.n. [Lake City, 21 July 1893] (BE). Dade Co+ Edw. Palmer 397 (E, G). Duval Co.: Curtiss 511) (£). Frank Cos: Ae ig Chapman sen. [Streets of Apa. Apa.) (2 (BE). Hernando Co.: A+ S. Hitchcock s.n. [June-July 1898] (BE). Lake Co.2 G. Ve Nash 12h8 (E, G). Lee Co.: A. S. Hitchcock 269 (E). Manatee Co.? Ss; 1s Tracy 6652 (E, G). Okeechobee Co.: E. Bc de Palmer 27162 (E)- Orange Co,: Fredholm suns (E, G). Saint Johns Co.: Rugel 156 (E, F). MISSISSIPPI: Jackson Co.: C. L. Pollard 1191 (E, G); Skehan s.n. (Ocean Springs, rv Aug. 1895] (£). LOUISIANA: Orleans 3 Par! Joor s.n. [New Orleans, 21 Oct. 1885] (E); Riddell s.n. [New Or leans] (G). Plaquemines Par.: Tracy & Lloyd 22 a 22 (E, a Se Comal Co.; Lindheimer 618 (E), 1077 (E, G), sn. [New 1847] (E). Harris Go.: Lindheimer s.n. {Houston, July 8h2) (E)- Jefferson Co.: E. J. Palmer 10692 (E). County eS Bigelow s.n. (Leon R: River, Oct. 1850] (G). ARIZONA: Pima Co Pringle 8.n. [Santa Cruz River, Tucson, 11 May 1881] (G), oats [banks of the Santa Cruz Cruz River near Tucson, 18 July 1884] (D, HM G). CALIFORNIA: Los Angeles Co.: A. Davidson s.n. [Los Angeles 196 Moldenke, Monograph of Verbena 129 1892) Re 6 (G). San Bernardino Co.: S. B. Parish s.n. [San Bernardino, Oct. 1891] (E); Parish & Parish h 10L3 (E, F), 21143 (G). County undetermined: Wallace s.n. (G). MEXICO: Baja California: C. R. Orcutt 1302 (E). Coahuila: Edw, Palmer 1040 (D, G). BERMUDA ISLANDS: Main: Brown & Britton 373 (D, G), 2631 (Ors FS Collins 268 (G); A. H. Moo Moore 2874 (G). Smith: hook H. Moore 29]7 (G). CUBA: Las Villas: Cambs 309 (E, G). Province un- determined: C. Wright 3659 (G). JAMATCA: “W. Harris 11808 (£, G); A. S. Hitchcock s.n. [Port Antonio, Dec. 1890] (E); Lang Lang 102 (D). HISPANIOLA: Hatti: Eyerdam 201 (G), 432 (G). PUERTO RICO: : Read Sen. (D); Sintenis 1074 (E) ; > 2010 (D). The collection which she Cites as "Randolph 1012" is cited by me below as Randolph & Ran- dolph 1012, while her "Leon & Edmunds 8719" is Leén & 8719. “The Curtiss s.n., Keeler s.n., and Pollard rd 1191, whic which she cites from "NY", are all actually in the Columbia University her- barium, while the Pr River near Tucson, July 18, J which she also cites from "NY" is in the Barnard ege ise sp She comments: "Verbena scabra, often confused with V. urticifolia, is easily separable by the very scabrous upper surface of the leaf and the rather conspicu- ruit face apparently between two almost equal sterile rb gag orate This condition is somewhat approached in Vv. carolina, other species examined the second sterile lobe has never 7 head so definitely developed." She cites Fuertes 391 & 1758 from Hatti, but they were actually gathered in the Dominican Republic. The "Krig 3187" which she cites is an error for Roig 3187. _iIn all, 464 herbarium specimens and 2 mounted photographs, in- cluding type material or phototypes of all the names involved, have been examined Citations: VIRGINIA: Isle of Wight Co.: Fernald & Long 12785 (Au, St, Vi). Princess Anne Co.: Fernald & Long ; Long 10799 799 ( = z W—1810833), 12453, in part (Gu--27186). Shenandoah Co.: H. : Allard 8270 (W—1829979 Surry Co.: Fernald & Long 6863 (N, 3 W-~-168 ze) NORTH bere Bladen Co.: ites & Duis 50975 Hi~110657) ; Ahles & Leisner 33335 (Hi--l0hl1). Brunswick Co.: Fox, & Beaman 50046 - 50046 (No—26200). Carteret Co.: R. K Soatrey 49809 (N)5 Re oo Ee 8.n. ee bah (H—50217); Lewis 229 (i). Chowan Co.: Ahies 147840 (Hi—10439); A. R. Moldenke 468 (Fg). <4 Coe: Ae é ee sn. {summer 1931] (Hi--13321). New Hanover Co.: Randol ih & Randol seers 22 h 1012 (Ba). Onslow Co.: Ah- ies & Leisner 32582 (Hi—10l)36) « er Co.: Ahles & Lelener 32380 (HI—I0LD35). Per quimans peo Ae "Re Moldenke SOUTH CAROLINA: Beaufort Co.: Ahles & Bell 15630 Teeay THe, i 92866), 18011 (Hi—92867). Charleston Co.: Ahles & Leisner 3209h 130 Pay tT 0:10.06 I-A Vol. 11, no. 2 (Him~104440). Colleton Co.: C. R- Bell 4675 (Hi--92868). Dor- prea poh Ahles & Leisner 32007 ‘(Hi—104]37). Georgetown Co.: on 277 (Ca—956876, Gg—290714, N, W--1837183); Rad- ont Ses (Hi=-50725), 31332 (Hi-—-92727) « Horry “ie Duke 19 ian Soe) Jasper Co.: E. 0. Mellinger s.n. [May 2 9 1959] (Hi--1,6718); Mrs. E. 0. ellinger 338 (Hi--13L657) . ‘Baer Hel- ena Island: Cuthbert s.n. [Oct. 1903] (Fl—-21119). Sea Island: Cuthbert s.n. (Aug. 1884] (Fl—21136). Sullivan's Island: Bilt- more Herb, 14892 (S). GEORGIA: Dapelo Island: W. H . Duncan 20258 (Lb—-]2772), 20268 (Au--167515, Gg—~L13460, Hi--106316, 6, W--22625 75) FLORIDA: Alachua Co.: Arnold & West s.n. serene ts 17 Sept. 1937] (Fl—27112); Cuthbert s.n. (Fl—2113h, 21135)3 Hume, Arnold, & West s.n. (Gainesville, 3 July 1937) Sl ae vard Co.: H. N. Moldenke 21503 (Hk, Ok, Sm, Ss, Z); W. H. Rhoades Son. (Merrit, “L/2/28) (Fl—11930). Broward Co.: He Ne Moldenke 591 (E, Go, H--5477, N, S, Up, Ur), 599a (E, Go, N, S, Up, Ur) Small & Carter 1072 On Clay Co.s J. N. McFarlin 7898 (N)- Col- r Co.: Buswell s.n. [April 10, 1936] (Bu); C. C. . Deam 65573 (Dm), 65856 (Es, (Es, N), 5; bone (ik, N), 6601 (Es, N); Je 7 McFarlin a ome O'Neill s.n. [Everglades, 27 Aug. 1925] (Io—1279Lh, 17260). Columbia Co.: Quaintance 209 (Fl--21131, Ga). Dade oad = C. Deam 6086) (N); A. A. Eaton 196 (Rf); Elder 6h (H—- 69375); O'Neill son, (March 1h, 1933] (1); Edw. Palmer s.n. [Bis- cayne Bay, 1874] (W—71951); J. K. Small 4020 (N, Be 8090 (S)5 Small & Small 520 (N). DeSoto Co.: J. K. Small 8186 (N). Duval Co.: Curtiss 5111 (C, W—22h92); Keeler 8 Sen. ae of May~ port and Jacksonville] (C); Lighthipe 406 (N); Rugel 639 (W— 264614). Escambia Co.: Cc. T. Mohr s.n. [: {Pensacola 5 vuly 5, 1872] (W—771853). Flagler Co.: West & Arnold s.n. (Andalusia, 10/10/ 40] (F1—32843). Franklin Co.: R. M. Harper 242 (N, W--1039060) « Highlands Co.: Brass 15556 (W—2065618) . Lake Co.: G. Ve Nash 1248 (C, Ca—10)867, Es, Mi, Mm~15)01, s, W--228276), rik i oes Lee Co.: Buswell s.n. ty 2h, 1933] (Bu); C. C. Deam 65658 (Da Es, N); A. S. Hi S. Hitchcock 269 (N, W—387303), 3.N. . (Myers, Jul Aug. 1900] (Ka); H. N. Moldenke 972 (B, E, Go, H--5478, N, Ob, 5» Up, Ur, W—118h)13); J J. P. Stan P. Standley 427 (W--910802) « Leon Coe! s Watson & Murrill s.n. [Gulf Hammock, 6/18/39] (Fl--31h55). Mana- tee Co.: S. M. Tracy 6652 (N, N). Monroe Co.: Meebold 27567 (Bi)- Nassau Co,: Biltmore Herb. 1112c (Po—267670); P. 0. Schalle: 14103 (Hi—1],6050). Okeechobee Co.: Harshberger s.n. (Okeechobee, June 22, 1912] (Up—-62867); small & Small 43hl (S). Orange a 11621). Osceola Co.: Fredholm 59h (W--717355). Palm Beach CO«t 196), Moldenke, Monograph of Verbena 131 Baker s.n. [Belle Glade, Sept. 1925] (Fl—21118); W. B. Fox son. [Delray Beach, May 25, 195] (We); O'Neill 852 (I); : Small & Small 4337 (N, S, W--1737793). Pinellas Co.: M. H. Williams son. [Max~ imo, March 12, 1926] (H--26130). Polk Cows Ce C. Deam 64204 (No= 21221) 3 J. B. McFarlin 5930 (Gg—237854). Saint Johns Co.: | Rugel 156 (Bm, W-—512025). saint Lucie Co.: Small & Small 341 (N). Taylor Co.: R. K. Godfrey 53548 (N).- Wakulla Co.: Redfearn & Kral 290 (N). Observation Island: Small & Small 1],03 (N). $ Saint Vincent Island: McAtee 176la (W—514792) . inte Island: Orrok Sen. (Sanibel, March 20, 1915] (W—-910360); S. M. Tracy s.n. [San- bel Isl., 18 May 1901] “(H). ALABAMA: Mobile Co.: C. T. Mohr s. n. (W--771852). MISSISSIPPI: Hancock Co.: Demaree 29588 (N). Har- ison Co.: Demaree 32278 (Le). Jackson Co.: Demaree 28367 (Le); os La roland a (C, W-~271773). ARKANSAS: Drew Co.: Demaree 6 (Sm), 26748 (Bm). Washington Co.: Herb. Univ. Texas s.n. = stevie 7 2/29/27] (au), sen. [Fayetteville, 7/29/27] roe is. LOUISIANA: Orleans Par.: Brenning s.n. [17.6.190h] (B); T. Drummond s.n, [New Orleans, 1832] (Lu, ae, » 5), Son. [New Or-— leans, 1833] (Lu). Plaquemines Par.: oyd & Tracy 22 (MN); Tracy & Loyd 22 (Cm, W—38353k). Saint sexes Par.: R. J. Lemaire 866 (Tl). Tangipahoa Par.: Correll & Correll 9239 9 (H--78635, N). Terrebonne Par: Arceneaux 383 (It). TEXAS: Bell Co.: Wolff 3243 (Tr--22252). Bexar Co.: Clare 782 (Ar—257309), sen. [San An io, pale (Ew); Metz 782 (I). Brewster Co.: Be) H. Warnock 8020 (Re ne Coe: * Lindhetnes 618 (Ka), 1077 See , Co AS o e, Me, N, Ok— Up—h8536, W— 2450) . Gonza les me Siuanh 969 tig tage — in part Pera ar Tharp son. [6.27.27] (Au); Tharp & Barkley 13850 (Au, N). Hardin Co.: GC. Le L. Lundell 1,079 (1d). Hays zap Ecology Class s.n. [San Marcos, 10.12.30] (Au); | Stanfield s.n. [San Marcos, sumer er 1898] (N). Jeff Davis Co.: T. Head 16 (Au--122665, St). Jefferson Co.: Kirn 2139 (Po—158770, Wi); E. J. Palmer 10692 (Du—205609, S, W- 1569398). Kerr Co.: G. L. Fisher 32217 (W--1624194), son. [Kerr- ville, Aug. 27, 1932] ( (Bt—}33826, St); Normand s.n. [Frio R w. of Hunt, 9.30. 1929] (Au); Parks & Cory 23921 (Tr), 23983 (Tr), "23984 (Tr); Whitehouse sen. [n. of Hunt, 9.1.30] (Au, Au). Kimble Co.: trandtmann s.n. [Aug. 19, 19)1] (an). Liberty Co.: E. J. Palmer 8557 T (og-—182i68, W—15318h7). Newton Co.: T sen. [7-20-39] Ge). residio Co.: Hinckley 213 (N, N), 3982 (N), 4122 (N). Co.: Cory 39708 (Au), 39709 (N, N), 4277h (au); Parks & Cory 27u15 (Tr-16228); 7 Tharp, Follan Follansbee, & Thompson 51-1663 (Au-- 122662). Reeves O08 sr Ea & Gong Lol (es, WE (Or), 120h6 ee Travis Co.: C. C. Albers 40003 (Au, Au, Au, Au, Au, Au); Ag 4. Barkley 13566 hae N); Herb. Univ. Texas s.n. [Austin, 10-1- pe a Se oR Rs ee I Vol. li, no. 2 34] (Au, Au); Strandtmann s.n. (Aug. 1, 1940] (Au); Tharp son. (Capt. Aldrich, 11/12/31] (hu, Au), sn. [Austin, 7-18-1] (Ca-- 882811, Sm). Uvalde Co.: Parks & Cory ry 23877 (Tr). Val Verde Cos ry 38068 (Au, Au), 38069 (N, | (N, N)- Walker Cost C. C. Alvers oI (Au). County undetermined: Collector undesignated L.25-538 [Skillman] (Sr); Cottle s.n. [Skillman, Aug. 23, 1931] (Sr), Sone (Skillman, Aug. 29, 1931] (Sr); Havard s.n. [Pefla etal (W=- 155628), sen. [Cottonwood Spg.] (W--221166); Lindheimer S.No {southern Texas, 18)9--51] (N), s.n. [1850] (Ka). ARIZONA: NA: Co- chise Co.: Blumer U.213 (W—-563590). Gila Co.: G. J. Harrison ni on o--158506, Up—1 7109, Vt, W). Pinal Co.: G. J. Harrison n 1897" (213677). Santa C Co.: Pringle s.n. [1 July 1862] (C). CALIFORNIA: Los Angeles Con Braunton 532 (Ca—-56839, Du-—-7559h, We-lj65514), s.n. (Sherman, Nov. 1902] (Du--91157); A. Davidson s.n. [Los Angeles, 1892] (Du— 91131); G. B. Grant 5057 ’(Du=-7565h) , Sen. (Sherman, 2 Nov. 102] (Po--26 7667) ; 3 Grinnell 5 s.n. [Los Angeles river bottoms] (Du— 86983); J. T. Howell Howell 3174 (Gg--176993); McClatchie s.n. [near Pasadena, 12/17/1892] (N); C. B. Wolf 1093 (Ca--729)98, Du-- 282943, Rs—~587%6, W—-18)5834). Orange Co.: L. M. Booth 1334 (Ca, Ca--537269, Po—-201879, Ua—1h737); Hwan 7731 (En, I); Je rs Te How ell 772 (Rs--655). Riverside Co.: Jaeger sen. (July 15, 1921] (Du--1)0388); McClatchie s.n. [near Elsinore, /23/1892] (N)- San Bernardino Co.: F. Re Fosberg S.44188 (N); I. M. Johnston 8.n- (Upland, July 1917] (Po--h013), sn. gen, [3 km. | south of San pated Berar dino, 2 Sept. 192k] (Po-~7883); Mun Munz & Johnston 11289 153388) ; Se Be Parish 5113 (Du--9539, W—-Wlh871), 5338 (N), n), 5939 (Ca~-1112h9), 6id (Au), 7149 (Po—126473), 9463 (Vi), Sone {san Bernardino, Oct. 1891] (Ca, Ca—-25152); Parish & Parish 103 (Br, ey ae Io--92220); Roos 964 (Se-—7735 53). Santa Barbara ara COs! llard s.n. [north of Veronica Springs, Aug. 8, 1957] ae 76857, Gg—412067). MEXICO: Baja California: C. R. Orcut 1302 (Ca~104823, Dt, Io), son. [7-15-85] (Ur), s.n. oe eas] (ii); Wiggins & Denaree 4734 (Ca—-511510, Du--215362, Fs, Mi, N, Ob—68,01, Po--213)38, W--1587985), 766 (Ca—~51151,6, pu—219987, Fs, Mi, N, Ob—65512, Po—-21355, W-—1587979). Chihuahua: Edw. Palmer 36ly (Pa). Coahuila: Edw. Palmer 100 (Io, Pa, ergs W—1323117). BERMUDA ISLANDS: Main: Brow & Britton 373 (N, 45684, W—524936), 1631 (N); F. S. Collins 268 (N, oe); "Oe Degener 130) (N), sen. [July 21, 1921] (lls), sn. [July 2h, 1922] (Ba); A. H. Moore 2874 (ce—-155393, N); Rankin s.n. Songs a 1, 1897] (Pr). Smith: A. H. Moore 2947 (Gg—-1553 CUBA: Havana: Boldo 60 (q), 112 (4); Ekman 78 (S), 933 (3), ago! aot 196), Moldenke, Monograph of Verbena 133 (S); Le6n 685 (N), 1343 (Ha, N), 13313 (Ha, N), 13U0 (Ha, N), 8. Re (Vi—L191) p Soe ~ [Almendares ] (Vi--l,026) ; Leén & Edmund 8719 (Ha, N); J. 7. Roig 3187, in part (Ha). Las Villas: Je Be Galé 1743 (Es, Es); Britton & Wilson 5771 (N); Cambs ; 389 (Io-— 15353, Ka--61357, N, W185 35) 3 3 C. Wright s.n. [near Manacal, Trinidad, 1865; Herb. sauvalle 743] (Hv). Matanzas; Britton, Britton, & Shafen Shafer 292 (Es, N). pia del Rio: Acufla & Zayas 17101), 8493 (Es). Piciiace ae ermined: C. Wright 3659 [1865] (Pa, W—71966). ISLA DE PINOS: Britton, Wilson, & Leén 25256 (N, W—-793637). JAMAICA: Re Ce Alexander s.n. [22 Jan. 50) (N); N. L. Britton 1527 (N)3 Britton & Hollick 270k (N)s Fredholm 3175 (W--316019); W. Harris 9937 (N, W--056765), 11808 (Gg~-3108, N, W—-790870); H artweg tweg 1558 (Lu); C. R. Orcutt , 2297 (Ca—130597) 3 Webster & Proctor 5334 (S , W~2227703). HISPANIO- : Dominican Republic: 1.13985 (s, W--1712018); Fuertes 391 (N, W--658353), 1758 (N, W—759366). Hafti: Ekman H.5 (N, S), #47229 (S), H8642 (S, W-1413151); Eyerdam 201 (W—130329k), 432 (Se-=20915, W--1303198). PUERTO RICO: Britton, Britton, & Brom 6019 (N), 7037 (N); Eggers 996 (Br), 1162 (W--1323133), s 3.n. [Si- erra de Luquillo, April 1883] (W—1159327), sen. [1883] (Cp); Je R. Johnston 742 (N); Moldenke & Moldenke 19546 (Es, lg, N); "F. Le S." 8359 (N); F. H. Sargent 695 (W—--1558570) 5 Setting 767 (8, Sg—16119, 1 W—1323135), 107k (B, N, N, Pa, W—~71961, W--132313h), 2010 (Es, Po--63865); Stevens & Hess 1260. (N); J. A. Stevenson 5335 (W#—1753)0) . WEST INDIES: Island undetermined: Swartz s.n. [ind. oce.] (s, s), sen. ioderaat (S). LOCALITY OF COLIEC-— TION UNDETERMINED: Swartz s.n. (S, S); Von Rohr 35 [America mer- idionali; Macbride photos T8] (Kr--photo of type, N--photo of type), sin, (s). VERBENA SCABRELLA Sessé & Moc., Pl. Nou, Hispan., ed. 1, 1: 6 {as womens’) 1887. Suppl. 7: 249. 1929; Perry, Ann. Mo. Bot. Gard. 20: 313 & 356. 1933; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 19 & 102 (1942) and [ed. 2], 33 & 199. 1949; Moldenke, Résumé 0 & pas Red | Moldenke, Résumé S Suppl. 7: 3 & 8. 1963. Mex. Ic. 16. Herba sesquidodrantalis caule simplici scabro e~ recto. Folia alterna oblonga — integerrima scabra. Spicae te-minales solitariae laxae o blongae. Flores purpurei. Bracteae singulis floribus ternae subalatae ciliatae, Semina in- tra caosulam oblongam. Habitat in saxolis Cuyuacami circuitibus. 134 PHYTOLOGIA Vol. 11, no. 2 ped: Augusto. ©, # Nothing is known to me about this plant except what is stated in the original description. Perry (1933) says: "Apparently this species does not belong to the genus, but anything fu urther re- t Maximo tinez tells me that the type locality is probably Coyoacan, south of Mexico City, Federal District, Mexico. XVERBENA SCHNACKII Moldenke, Phytologia 2: 150. 196. Synonymy: Glandularia peruviana x megapotamica Schnack & Covas, Revist. See: ent. Agron. 12: 222 & 22--229, fig. 1B, 2, & 3 He-L, & pl. 12 A, B, D, & G. 1945. Verbena megapotamica Spreng. x V. per- uviana 3) Britton ex Moldenke, Résumé 369, in syn. 1959. “Verbe- na peruviana (L.) Britton x V. megapotamica Spreng. ex Moldenke, Résumé 372, in syn. 1959. Bibliography: Schnack & sees Revist. Argent. Agron. 12: 222 & 22ln229, fg. 1B, 2,%3 , & pl. 12 A, B, D, & Ge 1955 Schnack & ez, Revist. Ar ot: Agron. 12: 290. 1945; Molden- ke, Meetsionit 2 2 ie. 196; H. N. & Ay L. Moldenke, Pl. Life 23 82. 198; MoIdenke, Known Geogr. Distrib. Varbemno., fase 2], 164 & 199. 1949; Moldenke in Chittenden, Roy. Hort. Soc - Gard 6: 2212. 1951; Moldenke, Am. Midl. Nat. 59: 361. 1958; uoidenk, Résumé 22h, 296, 369, 372, & h7h. 1959; Moldenke, Phytologia 8: 121 (1961) and 10: 133. 196h. liustrations: Schnack & Covas, Revist. Argent. Agron. 12: sce of Fe: & 1B, 2,&3 HL, & pl. 12 A, B, D, & Ge 195~ e artificially produced hybrid between V. megapotam- ica parce Task V. peruviana (L.) Britton, first produced in Arm gentina in 1945. The two parental species grow together in at least two states of Brazil, in Paraguay and Uruguay, and in at least one province of Argentina, so it is possible that this hy- brid may yet be found in the wild. In fact, some of the anomal- ous specimens distributed as one or the other of these genre represent this hybrid. It should certainly have considerable horticultural merit, as both parents are handsome~ red. The hybrid is named in honor of Benno Julio Christian Schnack (1910—), a specialist on the genetics of cul- tivated plants and on the taxonomy of the Verbenaceae. chnack & Covas (19hs) describe the plant as follows: "El porte vegetativo del hfbrido es intermedio respecto =< ambos Ce rg simple : doblemente lobulado-aserrado Bricte as florales de for- intermedia respecto a ambos padres (Fig. 2 ay. Pobepo ints del ealiz semejante a las de G. peruviana, con los pelos m4s ralos oblicuos. Flores con la oo swpesice: 1 limbo de color lila rosado [Este pe at parece eberse a ezcla de pigmento derive- do de pelargonidina con pigmento selgeae _ delfinidina. P0 otra parte ya es conocido en Glandularia (Lawrence, W. J. C+, and J. R. Price, Genetics and chemistry of flower colour variation, 196), Moldenke, Monograph of Verbena 135 Biol. Rev., Vol. 15, a4 1, 1940) que un gene que condiciona la produccién de pigmento derivado de pelargonidina es incomplete- Ib Pigmento derivado de delfinidina, produciéndose en consecuencia mezcla de los dos tipos oe pigmentos.]; la intensidad del color es del mismo grado que n G. megapotamica, es decir el car- 4cter dilufdo se ha auesereeie’ como dominante. Las papilas epi- dérmicas del centro de los 1ébulos en la cara superior del limbo, son de forma intermedia respecto a ambos padres (L4mina XII, D). ° to a ambos padres. Garganta de la corola con pelos moniliformes de color muy pélido, Limbo con los lébulos de posicién intermed- ia respecto a los padres (Fig. 1, B). Estambres superiores tg lébulos glandulosos apenas esbozados Urs. 2, B). Estigma forma y posicién intermedia respecto ambos" padres (Fig. 2, 2 Fertilidad del polen relativamente ctevedh (aproximadamente 0/o). En la meiosis del hfbrido se observan lhe Became srl adenas como ¢ e cromos y cromosomas no apareados (Fig. 3: I, J, Ky L), que indican que G. peru x G, megapotami un rido patehe hee heterocigota aka ag name Roe eg rido G. santiaguensis x G. laciniata [=x V. covasii ist dactcs' (Schnack, B., y G. Covas, Hibridacién interespecifica en Glandv- iaria (loc. oe. )], cuyo polen presenta una esterilidad proxima al 50 50 o/o, y cuyas especies paternas son relativamente poco dif- erentes desde el punto de vista morfolégico. El est del hibrido G. santiaguensis x G. megapotamica [=xV. vaga Moldenke] ico, tradue ae nae otre e completamente estéril; correlativamente las diferencias norfol6gteas don grandes entre G. santiaguensis y G. me lidad del polen y el hfbrido g G. peruviana x G, megapotamica, indican una estrecha entre G. peruviana y G. megapotamica. ra erencias m morteligicas entre ambas especies paternas son ente pequefias ." relativam ke, Known Geogr. Distrib. Verbenac. -» [@de 2], 107 & 199 19L953 E. J. Salisb., Ind. Kew. Suppl. ll: 263. 1953; Moldenke, Résum 128 & 474. 1959. Herb. 30—l0 cm, tall; stems and branches acutely tetragonal, hollow, pilose-pubescent with short spreading hairs, somewhat less so in age, the hairs whitish or sordid; nodes annulate; Principal internodes 2 -5—-7 cm. long; leaves decussate-opposite 136 PHYTOLOGIA Vol. 11, no. 2 and sessile; leaf-blades chartaceo 2 vee Ged a sagt okie on h surfaces, lanceolate, 2 ae co cm. - wide, nel (or the upper only somewhat sttomato-acwts) seg fia. apex, som what clasping as the Bled sharply and rather irregularly ppc | along the margins, scabrous above, scabridous and rather sparsely pilose Deoeath: especially along the larger veins; mid- rib slender, som impressed above, promin beneath; secon~ daries very slender, 6 or more per side, somewhat impressed above, 8 rominent beneath, ascending, slightly arcuate; v' reticulation rather obscure on both surfaces; inflorescence ter- ciliate on the margins; tubular, about 1 cm. long, costate, abundantly w write-pilose; © eats vivid-rose, the tube b t-rose, about 12 mm, long, densely pubescent on the outside above the calyx, the limb about 6 mm, in diameter. The type of this rare species was collected by Augusto Gustavo — (no. 284) — in whose honor it is named — in swampy gro la = December 7, 1933, and is deposited in the Osten Herbarium at the Museo de Historia Natural at Montevideo. Thus far the species is known only from the original collection. Two herbarium speci- mens, including the type, and |, mounted photographs have been @x- y me, Citations: ARGENTINA: Chaco: A. G. Schulz 28) [Herb. Osten 23148] tga of type, N—isotype, N--photo of type, Sg--photo of type, Ug--type, Z--photo of type). XVERBENA SCORTA Moldenke, Phytologia 5: sat 1955. : Verbena halei x prostrata D » Cytologia 7: 170. 1936. Verbena halei Small x V. Tek cteasin’ Link ex Moldenke, Ré- sumé 365, in syn. 1959. Verbena as ashostac Link x ¥. halei 959 Bibliography: Dermen, Rent Srp 7: "170. 1936; Moldenke, Phyto- logia 3: 167 (1951) and’ SPs 133. 19555 Moldenke, Biol. Abstr. 30: 1093. 1956; Moldenke, Am. Midl. Nat. 59: 361. 1958; Moldenke, Ré- , 5, 68, & ith. 1959; Amiens, Phytologia 8: 121 (1961) and oe "1hb 1963 s is the hybrid between V. halei Small and Vv. lasiostachys Link, produced artificially in cultivation by Dermen in Massach' setts in 1936. The two parental species do not actually grow gether anywhere in the wild, as far as I am aware, so it is not very probable that this hybrid will ever be found in the wild, ur less one of the parental species eventually invades the territory now occupied the other in the southwestern part of the United States. It does not seem very likely that the hybrid would ever be regarded as possessing much horticultural merit. 196, Moldenke, Monograph of Verbena 137 VERBENA SCROBICULATA Griseb. ex Lorentz, Veg. Nordeste Prov. Entre Rios, ed. 1, 122 & — hyponym. 1878; Abhand. Kaiser Gesell. Wiss. Gotting. : [Symb, Fl. ane. ] 275—276. Synonymy: Verbena hunzikeri Moldenke, Phytologia 2: 321—322. 1947. Verbena scribiculata Griseb. ex Moldenke, Alph. List Cit. 33 913, sphalm em 199. Bibliography: Lorentz, Veg. Nordeste Prov. Entre Rios, ed. l, 122 & 150" "18783 Griseb., Abhand. Kaiser. Gesell. Wiss. Getting. Prov. Entre Rios, ed. 2, 122 & 150. 1947; Moldenke, Phytologia 2: 321--322, 338, & 339° (1947) and 2: 4,82. hae Pars Cas- tanea 13: 117 & 119. 1948; eee Alph. : 364, 367, 368, 3h 376, 378, 389, 458, 53h, oo. TS, : 9, (age), ” 2 Ne 70h, 732, 745, 746 3: a 666, 672—67h4 “tea 71, bo, 80h, 858, 863, 869, 907, 913, 916, 1 922 sis, and k; 1088, 1091, ue 1166, 1207, i880, Pt 1293, & 1302. 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 98, 99, 107, & 199. 19193 Moldenke, Phytologia 3: 7 B . 7 (2318), 3: 289; 290, & 305 (1950), and 3: 468. 1955 | stelif Farmac. 19 (10): 167. 1951; E. J. Salisb., - Kew cee 263. =o arr se Inform, Mold. Set us "spe. [4] (195k) oat es Résumé 120, 22h, 360, 362, 366, oldenke, @ lth, kins uoldenk, Phytologia Bes parse (1963) and 10, "17, 175, a by7. modes ok 0.25-—-1 m. tall, somewhat woody at cm. long, 9-- 2h mm. wide, regularly narrowed from the broad base to - atten- uate-acuminate apex, truncate or subtruncate and entire at the incised belo ow, sparsely pilose or hirtellous-scabrous above, densely pilose-pubescent beneath, less densely so in age and the poy mostly concentrated on the larger venation; midrib very very slender, h--6 per side, ascending, only slightly arcuate, obscure or very slightly jitieorensed above, obscure or slightly 138 PHYTOLOGIA Vol. 11, no. 2 wlous beneath; inflorescence terminal, 2-3 cm. long, de~ - Wide, usu long as t even shorter, glabrate except for the long-ciliate margins, sharply attenuate or acuminate at the apex; c narrow-— 3 short-pilose on the ; parallel ribs, otherwise subglabrate, the 5 sharply acute teeth 1—-1.5 mm. long, irre or iter= i P from blue, deep-blue, or blue-violet to bluish- lilac, rose-lilac, d-rose, or vi white, rose-pink deciduous, its tube l. ong, less puberulent or pilose on the outside, hirsute insid he cm, wi the lobes deep ginate or cordate at the apex; stamens r ones subsessile, the upper ones abo ing the mouth of the corolla-tub jlaments short; anthers 2=celled; style surpassing the lower stamens; igi with one lobe capitate and the other linear; cocci about 3 mm. long, half as long as the fruiting-calyx, linear, smooth below, densely scrobiculate above except for the commis ssure. ype of this much misunderstood species was collected by Paul Gtinther Lorentz ee Georg Hans Emmo Wolfgang Hieronymus (n0- 244) near San Lorenzo, Jujuy, Argentina, ad was deposited in the Herbarium of the Botanisches Museum at Berlin, where it was photo- formation along the highway between Alto del Clavillo and Alpachi- _ entina, on September 18, 1916, and is deposited in the Britt ton Herbarium at the New York Botanical Garden. phlogiflora Cham. and V. incisa Hook., whose densely pubescent calyxes at once serve +t pro crchage eg them. Grisebach (1879) says "Proxima V. phlogiformi Cham. V. incisae Hook., distinc calyce brevior, corolla extus ahem et foliis etiam poets longiuscule et abruptim petiolatis." de has been collected in sandy mud near riverbanks or among treams, in moist sand of riverbanks, dense thidaber see soil, and erassy ground, in shade, on campos, al smal reams, and among tall herbs in ee wet soil, at Salta; J 3 " voorst says that it is a "characteristic element of Alnus woods" in Tucum4n., Material has been misidentified and distributed in 196), Moldenke, Monograph of Verbena 139 herbaria as ¥. pulchra Moldenke. The plant f. figured i in color as V. scrobiculata by Descole (194k) on my determination is actually V. peruviana (L.) Britton. Some 20 years ago I regarded V. scrobiculata as the correct name for what are now known as V. incisa Hook. and V. peruviana (L.) Britton, since at that time I had not seen a photograph of the collection of Grisebach's plant. This is the reason for Seach eee of the plant on which Descole's excellent plate based, The V. briquetiana f. campestris Osten, V. briquetia- na f, Silvatica Sates. V. chamaedryfolia hybrida Osten, V. chamae- dryf: ecanea Osten, and V. m ica var. truncatula Briq., reduced by me to V. scrobiculata in my Résumé 20, eee & 369 (1959), actually belong in the synonymy of ¥. in- ne cabinets collections distributed as V. scrobiculata or so cited by me in the past are actually V. incisa Hook.: Archer 1,666; Bailetti 58; Bertoni 1942, 2251, 23h2, .s Bolte {Herb. Inst. Miguel iillo 28112); Ekman 1978; Fries 110); Garolera & Romero 10; Hens 33438; F.C, Hoehne s.n. {Herb. Inst. ae S. Paulo 0712 & 23061]; Hunziker 7135 Leite 250; Lillo 3220; Lourteig 487; A. Lutz 25085 T. Meyer 9799 & 1587]; | Moldenke & | & Moldenke 19721, 1972h, 19739, & 1975h5 Mi Monetti 1623 & s.n. (Herb. Inst. Miguel Lillo 3135h]; Mon- tes 1091 & 213 24353 Pa: Pastore 2007; Peirano s.n. (Herb. Inst. Lillo 32193]; R Reitz 882; Rodrigues 576 & 907; Sampaio 7675 & 7705; sommester 315 515 & & 810k; A A. G. Schulz 277 & 3988; Schwarz 2052, 312k, & 3162; and Venturi ari 378. “The following are re Ve moricolor = Moldeaket Herb. Osten 8169 & 2253h; Lillo 18111; and Parodi 9179. The fol- lowing are V. nana Moldenke: Brade 7002; Edwall s.n. eos Inst. Bot. S. Paulo 15732]; T. Meyer 2943; and A. G. Schuls following are L. peruviana (L.) Britton: Archer 1625; Castellanos S.n. A sic ees : Rea 33875]; and T. Rojas 3395. adage in interpretati ion of names for this taxon, the pata a tes by Osten are of f interest: Zu Herb Ce Osten No. est Verbena 'Briquetiana’. Cf. Briquet Verbenac Balansanae in Annulaire du Conserv. de Genave mtg war) P. 268. 190). Cf. Briquet in Arkiv f. Botanik II no. be Die Arbeiten von Chodat (Plantae ea. oti ese my weit sie die Sectio 'Nobiles' Schauer betreffen, sind voll von Irrthumen. Ich habe Briquet von 2 Jahren eine grossere Sammlung Chamaedryfolia in Paraguay nicht vorkamme. Dis ist richtig so- weit es sich um V. Melindres Gill. handelt. V. melindroides Cham. habe ich von Misiones und ditrfte sie am Alto Parand jedenfalls 140 PHYTOL OGI A Vol. 11, no. 2 auch in Paraguay vorkommen, Wemn man aber die rotbltthenden For- men der Nobiles zu einer sp. coll. (turma, grex) V. chamaedr. zuU- sammenfasst, so gehtrt diese Form 8046 jedenfalls ; dazu. Ich habe hier in Uruguay, in Paraguay, in Argentinien gefunden di Bltttenfarbe der einzelnen Arten sehr constant ist, dass dieselbe rot, violett, weiss bltht, ist absolut susgeschlossen ( Ss . Chodat in Plant. Hassl.J). Soweit ich aus dem Beschr Briquet's ersehen kann, hat er diese Form zu shaegitioah *(eagape tamica var. Tweediana) als ac truncatula gezogen. Sie ha nichts damit zu tun. V. phlogiflora ist hockwichsiger, mit oe seren Blttten deren Farbe immer lila violett ist, deren Blttten- stande sich nach dem Verbitthen nicht verlm, verltngern sondern kopf- fUrmig bleiben. Ich habe diese Form 806 in meinem Herbar als "Briquetiana' bezeichnet und halte sie fttr den Uebergang von V. incisa Hook. zu der V. scrobiculata Griseb. (Symbolae no. 1735) « V. incisa Hook. aus der Araucariazone, s/Br. Uruguay [nur fluss littoral!]; V. scrobiculata = Tucumanzone, ‘Alles fittssti'" I regard Osten S06 as V. incisa Hook. In all, 28 herbarium specimens of the true V. scrobiculata and 6 mounted. photographs, including the Sapek ee phototypes of all the names involved, have been examin Citations: BOLIVIA: Tarija: J. West 8275 (ca--565128). PARA- GUAY: Ponder 7708 (Je--7708) . ARGENTINA: Catamarca: J&rgensen 1298 (Herb. Osten 11351] (N, N, Ug, W--921951). Jujuy: Eyerdam & Beetle 22302 (Ca—-652223), 22598 (Ca—655675), 22673 (Ca- 2 % Levent & ee 244 [Macbride photos 3 17445] (Kr- phot hoto of type); C. Skottsberg s.n. [near Urun= del, "seo ein (G0). Salta: Ryerdam & Beetle 22626 (Ca— 655895) 3 Killip 39612 ie ee Pierotti 1223 (S); Ruiz Hui- dobro s.n, [Salazuti, 8/VIII/14] (Ca); Schreiter 1167 (Ca 164770, Ut--115h10b); A. G. Schulz 3433 (Sz), 5185 (Cb); Schulz & Varela 5267 (S, 2); Venturi 5056 (W=-151,8943), 8673 (We et a Willink s.n. [Herb. Inst. Miguel Lillo 106667] (Ca)- - T. Hunziker 6812 (Ba, F—-photo, N, N--photo, on ee photo, pee xvas Mexia 1332 (Ca--560598); Sparre 922 (S)3 ¥ Ah yoorst 4260 (Le). 5. = Biol. Abstr. 30: 3551. 1956; Moldenke, Résumé 81 & ‘7h. 19593 G. Taylor, Ind, Kew. Suppl. 12: 19. i 59. Herb, about 1 m. tall; stems and branches tetragonal, rather conspicuously sulcate and ribbed, browmish, glabrous; nodes anmun late, often with white setulose hairs; principal internodes 3.9c cm. long on matibe leaves, glabrous or with a very few setulose hairs; leaf~blades thin-chartaceous, brunnescent in drying, ellip- 1964 Moldenke, Monograph of Verbena 141 tic, 4--8 cm. long, 1—2.7 om. wide, acute at the age, ee ir- regularly serrate from below the middle to the a: x, glabrou both surfaces or with a very few scattered edtiiods hairs; midrib slender, subimpressed above, prominulous beneath; secondaries very slender, about 6 per side, subimpressed above, prom ous be- arc veinlet reticulation rather abundant but often obscure sonetines the larger part Saou tage beneath; inflorescence ’ter- forming an open taitcle ¥ and 15 cm. wide, slang tatatsdanis the small flowers bass in spicate fashion on the. some ay the individual spikes to about 1) cm. long and 5 mm. wide, densely many-flowered, the flowers contiguous and ly pid, fie peduncles and inflorescence-branches slender, tetrag- nal, g tubular, about 3 mm. long, sparsely p sSuiliods, its rim bat oothed; corolla white, pstirt ar Seek; about mm. long, its limb about 3 mm. wide, The type of this rare species was collected by Robert I. Bowman (no. 81) at an elevation of 2500 feet on the north slope of the to me only from the original collection. Only 2 herbarium specimens, d me. tations: GALAPAGOS ISLANDS: Indefatigable: R. I. Bowman 81 (Ca-~13)99—type, 2~isotype) . = SESSILIS (Cham.) Kuntze, Rev. Gen. Pl. 3 (2): 257. i Limaea 7 ynonymy: Verbena stellarioides — sessilis Cham., 265. 1832, Verbena sessilis Kuntze apud Thiselt.-Dyer, Ind. iil, Suppl. 2: 191. 190k. Verbena stellarioides var, sessilis Cham. apud Moldenke, Prelim. Alph. | aie List Invalid Names 8, in <— ig Schnack & Covas, Bol. Soc. Argent. Bot. 1: 26h. 196. syn. 1 pasthgeaptiie tian; Cham. ; Tinaes 9 265 18325 Schau. in A. DC., - li: 52. 18475 Se Fl. Bras. 9: 187. 1851; 11. 190k; Thiselt .- . Kew. Su i a Briq. in : ee pe qe in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, k: 1059. ie Hassler, Flor. Pile. 101. 1909; Moldenke, Prelim. Alph. List Invalid Names 7 & 48. 19105 Moldenke, Alph. List Invalid Names 49 & 50. 1942; Mol- 2 28h. 19L6; Mo idenke, Alph. ast cit. as ing & & 263 3 Cigks) an and 2: 457, 458, 533, 600, & 142 PHYTOLOGIA Vol. 11, no. 2 628 (1948), 3: 665, 688, 70k, 7L9, 913, & 921 (19h9), and hs Be 127 & i Faia woldenke, Known Geogr. Distrib. Verben: ed. 2], 107, & 199. 19193 oe Te Pagtologsa 3s 76 (1913) = 3 37, 1950; Moldenke, R 118 # 37h, & U7. 1959; Moldenke, Résumé Suppl. o's 13 (1960$ and’7: 6 & BS “A oe weak herb, 60 cm. to 1 m, tall, with the aspect of Stellaria holostea but omen more fim and with the leaves somewhat thicker; stems almost 1 m, long, about the thickness of the quill of a feather, mostly simple, sharply angled, the angles subala a shortly attermate eg the apex, often serrulate, strigulose on the upper surface ("strigarum autem et dentium rudimenta et in var. & observabis identilatis speciei pignora"); bractlets shorter than the calyx; calyx praia about 6 mm. long, 5-veined, io the veins excurren ned eee the poe te; style setaceous, Hd Pome the stamens, the apex horn; cocci oblong, mm. long, slightly thickened at the base, 8 t venose, the transverse veins elegantly areolate, e areolae quadrangular, "color illi griseus pallide nigrescens carpello dorso octonervia, faciebus albidis angulo faciale parvo" so.” The type of this eign ee species was collected by Fried~ rich Sellow (no. 1563) in Brazil, and was deposited in the herbar- ium of the Botanisches epi at Berlin, now destroyed, but is conspecific with V. sessilis seems, rather, to belong in the sy? onymy of V. « stellarioides Cham. Kuntze (1898) says: "Chamisso hat in Linnaea VII: 26) unter V. stellarioides 2 ganz verschiedene Arten, die in der Inflores~ cenz Bracteen, Blttthe wenig Unterschied zeigen, aber in der Blut- tern so sehr verschieden sind, dass ich sie als 2 A rten behandelet @ decurrens Cham. = Verbena decurrens 0. Ktze. Folia ia decurrentis integerrima margine revoluta aspera ceterum glaberrima. p sessilts Cham, = Verbena sessilis 0. Ktze. Folia sessilia denticulata vel pie Paden: supra strigulosa. Hierzu ex descr. V. Morongit Ne BOOK REVIEWS Alma L. Moldenke “BOTANY - an Introduction to Plant Science", by mae W. Rob- bins, T. Elliot Weier & C. Ralph Stocking, 3rd > O14 pp., illustr. John Wiley & Sons, New York 16, New pang 1964. This new edition should continue the book's favored position graphs -~ are sensibly taken from other texts. Original nearly duplicate, illustrations are a waste of time, energy and money, =e since the functional life of any science text s y very short because the rate of scientific pane is so rapid. The introduction is well worth being assigned read A few desiderata are missed. There is no bibliography and there is very little reference throughout the text to the world's outstanding botanical writings and workers. The commendable showing of different systems of classification warrants an ex- planation. Indentations, letters and/or numbers would peo the key on page 17. There is no mention of sex in bacter page 373 despite so much modern experimental work in this tisid. The meaning of "stoma't is over~reached on page 86. This work must have slipped by the psesscenien's desk un- noticed. Some of the typographic errors Nand" for "as" on P. 3, and the misspelling of "vinifera" on “ee 8, "pond" on p. 53, "xylem" on p. 180, and "Sterculia" on p. 220. Labels should have been included on en drawing on p. 217; the legends on pp. 176 and “e need correction; the plant name should be given in the legend on p, 245; "anabolic" and "catabolic" should be in changed o on | pe 59. Split infinitives slipped through on pp. 65 and 67, and so did "referring back" on p. 55. ——_— = "Poisonous Plants of the United States and Canada", by John A. Kingsbury, xiii +626 pp., illustr. — Engle- wood cliffs, New Jersey. 1964. $13. This is a welcome reference book of value to botanists, chen- ists, pigstotogiote, cieterloglate. veterinarians and physicians because of abe 11 developed subject matter, its helpful Cornell Bulletin plan retool and the Shh pndied clear photographs, and 3 Uh PHITOLOGIA Vol. ll, no. 2 its ¢omprehensive bibliography of 1715 items. There are a tan- talizingly few fine color plates. The author first surveys the accumulated knowledge mt 700 in phylogenetic order, giving obvious botanical descriptions, poison symptoms, treatment and The text is easily readable. Just a few minor items for cor rection are noted: the legends at the top of p. 293 are in re- verse order. The following names in the Verbenaceae are to be preferred over those used in the text: Aloysia gratissint (Gill. & Hook.) Troncoso for "A. lycioides", Lantana camara var. acule- ata (L.) Moldenke for nL. aculeata", and Lant. Lantana Se nteviaensis (Spreng.) Briq. for "L. _ Sellowiana" "Bacterial pe piconet edited by ae — Anthony San Pietro & Leo P, » xvi + 523 ustr. Antioch Press, Yellow dina ’ Ohio. reapeert oe $6.00 This is Contribution No. 112 from the Charles F. Kettering Research Laboratory and consists of O excellent papers pre- sented at an invitational symposium, the first such symposium devoted exclusively to this topic. The authors are outstanding workers from all over the world; the dedication is to their physical and inspirational mentor, Dr. C. B. van Niel. The volume has the papers, each with its own bibliography and illustrative material, arranged according to the following topics: I. Components, structure » and function of the bacterial the fi directions for future Psd an additional bibliography, 4” a detailed index Since these bacterial systems have a more flexible nacanntet cal apparatus that can readily be manipulated, much ha and can yet be learned that is applicable to the Secunia of higher plants Herbarium curators who have material of this species fron additional counties are asked to send it to the author for verification and Mapping by counties done by Andrew R. Moldenke Figure 15. Distribution of Verb rigida in the United Stat ena rig e Unite ates record, so that future editions of this ma b t _—_— ’ p may be more complete 7 29 » 31 3 | 3 ei a: «ae - - = ‘i a 9 10 i 2 i 1d [ 15 I 16 I 17 [ 18 19 I 20 [ 21 2 hae [ 24 25 l 26 l ci | = L 1 - ] J L 8 E R H 1 | Tee Et Wan M A N TOBA Oo N j Lr J i | — ee . oa) en Four es ae a Uae Ta e PY TY gs Sa Hey . WaLey ee Es ee re SHERIDAN T= s 2 a rr a RENWILE | 0) HOUTEN Tinea ROLETTE [Towner | cavacien PEMBINA soos WIL Lawes ) WOUNTRAIL Mecneey ir y s en ine rawsey | WALSH . ay %. scAng - — | st Lous 9 ® ~ ues Meco cored stan “st 9 ° oe ween an « (eo mewnesy? = . NELSON | GRAND FORKS "= POLK cl] ‘ae OOK Co LD OTH Rasim | \ UF se aloe Uae Ss R 1 N a oamsom a N URN tT SHERIDAN WELLS EDOY 4 TASCA ‘ E ° & : T MERCER H D A TRAIL » Ko) we A N Prag . BLLINGs Foster macs a = ¢ 7, Nano BURLEIGH cua’ 7 ne oa Rideo | Over woo] stuTswan - sven cunt amma uss (ge a SOLDEN , = WALLEY Gute STARK Clay | BECKER CALLaty Ts, 7 ety = Les TOMAGOM wa -aaK TRE LEY on SWEET crass iol ‘SLOPE WETTINGER s cr CARLTON 1) mares " soni “ TELLOWSTone GRANT Tamons | Loca ser renee OTTER TAIL GoerBic “aon met poet spcKL ADAMS —l—| RON STILLWATER 816 Hoey © BOWhaAN SHOUK A M I N N i) ee — = oy 7 rr) . MciNTOSH DICKEY SARGENT l 1 ‘CROW WING ilLe = PINE waASNBUnn] SAWYER ASHLAND orn eo - Fe mi worrison | LACS 7 Cc 6 , Ane POWDER riven . = a Se 2 2 mee sheep 5 RICHLAND ONEIDA FLORENCE oo eee icc camps. | McPHERsom tadamngelanls proven: wae 4 anon 7 burner WARINETTE presque st =a a o ane s —_ an Cae Preto J a 2 ey H RAVENS. mt vee sng POLK [BARRON] RUSK TINCOUN a ‘ 4 Bes “~ nun OSSTED) eer es ri fea Apes TeeACH ) Dewey egestas EDMUNDS Day m Es : “a 4 axe Meme . °° $s TAYLOR Y ene SHERIDAN sts Aine aes Lge ROBERTS aT i OU ; — —atescoon [NCO FE Mcey hast CAMP REL | Mey POTTER FAULK SPINK GRANT gO sr crow DUNA MARATHON = \e 7 00K Moor ie ‘SHAWANO ¥ \ TASS CLARK] CODINGTON Car I No ste | ra] ae woe |e raancom | OSEMAM a at HNO suuLy woe ] van Deve Bayt rae = newt c\O N hc < om ied all a) VILLE wAuPi sis YELLOW MEDICINE ws ne out - s cv) ay wASianig : T pet weno] 3 snow 2! a ot H HES A K Oo ited svBLEy ie scan ° (ay osc Fetuaony ea ppt! BROOKINGS NICOLLET / Sueur | — RICE Fs qupnama Promemanpaonti| anon is mason HOT Springs E ry PimancTon BEADLE rungsBuRY ro 5 WonROt DAMMS xz acon | SARE H . TONES Tyan BUFFALO | ieRauLD [SANBORN | Mince | LaxE | MOODY gy PIPE-| MURRAY : — — eas (7 exanst " c= ad si AND) — TOM corronmoo0 | avo 0 Uncouy] SUBLETTE = CUSTER me BLUE EARTH STEELE OLMSTED aoe Resse |i en itt ar] GRArioT TRONA * ts en me Maccoox | Pook | ROBLES | JACKSON | WAR FamipauLy | Freteown | wower | FRU = By a Y COMVERSE SHANNON T WASHABAUGH WELLETTE eer Bi ca taammansaall base HT) _ sea) O M MOBeARA FALL RIVER TRIPP i; ‘ sapeaume aT) wai 2 re Tie Col . ll i N . BENNETT 7000 Chances OOUGLAS | HUTCHINSON | TURNER hd OSCE Poumon | EMMET | ROSSUTH] mumeneen] woRTH | MITCHELL] WOWARO = sux J cane = m= 5 sm Tirviason]| want me —T ‘ L. * four yosren [GAY [PAO AT HANCOCK Flova fowomaw ot inst H wn [om Sioux | oawes ee os GREGORY LNCOLs Faverie i » rr . = . . 80N | YANKTOW CERRO GRANT J sherioan | Cherny —e . . . HOMME cay [omy soRo0 . Tate | ROOK laa paca < 7] SMEETWaTER sami bon'e Saka ocmmerTn moma] wmnenT | FRavenkin | BUTLER | REMER user 7) pe rare | cosnen—e en ou ° —_ vista ur; . . nm ‘syst NaaW ocx nox — . iY j CEDAR conte ® WeaSTER ry Snecesoe ‘ cnt | ARE canon 1 a . segs [ANTELOPE] PIERCE ga Oe yd ‘ ee eae pac wal wooosuRy CALHOUN HAM T Tawa | BenTOR | Le somes Tanna | OGLE OY KAM ) COOK M : * +m 7 ro THURSTON EE —UGuowa | CRAWFORD [CARROLL ToRcEne [BOONE | STORY =e nae oi ene berg oa FC 2 Wooner | Thomas be cuDee e Bune [L0uF coarse] wnetien wADSon |sTTo8] CLAN A A CEDAR waareet ess ou) cant ome K Lean BOONE bai TSR [romemets| Towa | JOHNSON ssi proof > = oe "ome 2 aatnun | cPwenson | cbc | cusTER sae Mae cess nae J a ae hae vans came 2 mee» = B Wally | aReeiey mare [oom] poner carat) <7) a a am a] OnGGETT comity ina, 4 eases ont * 2 » om — 7 incon A ‘ POTTAWATTAMIE | CASS | ADAIR | MADISON] WARREN | MARION ogee Head ; ssa ar” i Pal Uinraa ror 8 . me . by Seman | Howanp NANCE sAuMOeRS Coser’ LOUIS wee H ts ra ors COR ‘mom 5 smells Sa Maen, / POX erent Cd (al OK Tigran |_ansracr] [LISTON | oat, Js & wo os Ps erry z TDMION [CLARKE LUCAS | MONROE as, eon > Sooo on StoGwice PERS DANSON BUFFALO | wa Your | Stwanp JuscaneP ~ cass { maa i CT FREWONT| PAGE | TAYLOR |RINGGOLO] DECATUR | WAYNE fa @ "0 BuNco CHASE MAYES aaa Om oan = a m f PMELPS. T KEARNEY Cay |Fuumone | sume ie AY oy Anson ] BOULDER wastincton J Yuwa TCMSOM, WODAWAT | WORTH ercen| PUTNAM L@) =, Fy SunoY HITCHCOCK cEnteY ‘SULLIVAN db = Gar . ee oad) mocnous | THAYER | ervcaso a 4 ome = ADAMS Loni 7 mit somes) (7) $P__ ds J “ pra! fetus con] SOM CHO ee) Sa? Sn 2 femme dope dee 2 2 i © _—_, oe wat ~~ fad < = ‘SuMMrT 7 HOE s honTom | Pears | Smith EWE REPUBLIC [WASHINGTON] MARSHAd | NEMAMA | BROWN ‘ M . Cc 7 xe ee UNCOLN Ta Carson guzel — a | a Seman Thomas ICHSOM, or | Oo Doucuas I PE rranaronne| — pst . nT GUNMISOm saa ne — ‘ ay & Pas wallace | cw SEE H 28 — + - a ; za = CHEYENNE sat sally X iil Chappe ) FREMONT ‘ MEE © acxsom m ae > | wu % r worms OSAGt | OOUGLAS | JOHNSON — | |cooren_) ne - p' _ lame UAC nae ee creeity | wicma | Scar — Tawn| a 8 = | 4 / N * N WMSDALE CUSTER " Case = "MI 3 | — ¢ ang BENT COFFEY [per Tj ‘ | oRcan |_| Fran | ie a, PROWERS nasa. Ton | KEaRwy | Finney F = avr. WonTEzona Juan RA st Cuan NTOM seer | fam, 0 Rane Roaosa ran | Gaay Prom ounun GatENWO0D FySooSOR] ALLEN | BOURBON VERNON MCKORT | coment oases] S ; a Ps i FORD ok AS = ARCHULETA LAS BACA STANTON | GRANT | HASKELL oma) Nc ‘i ae ; " c =e oem contvos J COsTis wae = ‘- ~ ann onc carton veut [Tous ne aw =—ss 2 TER] WRIGHT -_ me © ce —s WORTON | Stevens | sewano) MAZE Cam 3 BARBER SUMNER COWLEY CREENE wait 9 | Taos . — sa © = rs spi crautaugua) © canonerg (awmnct . Courax UNION : ° ow 2 s Taryn Cittaneon — som “awion iy DOUGLAS owt cover [UTUER | ; Ceaver HARPER 0005 ALFALFA] GRANT may we (mowatal CRAG | OTTAMA! arty cael wry * 7 NNEY OZARK = a ! j 4" 5 " o& ( swoovag OAL ® . s ELUS | wooowaRD GARFIELD WORE = Trocees ia ae > s bd © PR nanos Gout pews e s SHERMAN | HaNSFORO | OCHILTREE ipsceee — as waves =o coax) SOOKE HOM) BAXTER | FULTON : MAROING PAWMEE . saa ee ‘GREENE or nn | —)- r Pp ] ot amon | SANTA FE — LAWRENCE . = wane « Stent Moone [HUTCHINSON] ROBERTS | wexmpwaLL ees pn ee one sal TULSA WAGONER asaoncrony MADISON | NEWION SOT ot rane Ser, wl cont o/s | en s KnOEPENDENCE ea ae Jacscewnon) (PERRY Ye ayant . ) I at a LINCOLN POINSETT ey curs = i ne Y = ROGER MILLS es ee Bo Pe Imcon TPOPE van BUREN [CLEBURNE TPT SY onc SAME Temcnce] CLES Cad ven Pion 2 = porter | cansow feed =e wom U EMAIL | — O L JA |H Ol as cman Toss vateg seeay [FACE | fen . oon iis aman: ANOMA : . ‘ . GUADALUPE bd ‘aa Becanam | WASHITA = =1 = Mei TOSH 7 en i wie : | ana i" heer faa worse J) i TORRANCE DEAF sit) ‘i = eeTsT FRANCIS . unit = N E WwW . aasstionG | Dom Ey Pama gag acl TE Fone oa ye — es com | qe See em Q Q TRO E x I 4 GREER) Kiowa vers 7] - _ 5 i oat : Oo Comer pana WeCLAiN = oT bat rem 2 fax Le 3 Foam ae lec wan! % I OF ANA wwe | casteo Tsmisnen | BmSCOe Teal —Jomans j ans raw loa a ae Pairs a aa] atl eg CCT) x me wae aoe CER, . ° STEPNENS PUSHMATAKA 7 Lonone - = fi a ame [WTO Towar i? on pa) ni ER nant a a Taam _ an vena raowe i Games| jamais oa ° = 4 =) a -_ KEYED TO COLUMN A wom? ha ‘ - wuReay - CL ee be mE punt ot) 4 engi dex Book Feat “a 1a Puan Teoma] hana aTORA er od COAMOMA). en ata wom cr Ey awe of Cleartype Index JEFFERSON SONSTOM CUA « = farertt FF ERSOM aaa ¢ wane \ fort CHAYES SEVIER DALLAS ncOuM ” % Pie: worn *G en CARTER fuasasuc—F pevan | CHOCTAW CLEVELAND pL — 5 paosr suet cab tones Ne, esto R LovF Gat cuaY : — TER cur Rg Sera COCHRAN | HOCKLEY [LUBBOCK | GRoSaY | proxENS — he e° os” ome 1 = i bag PICKERS hae y Lod KNOK BAYLOR MOMER MONTAGUE | og GP | crarson My © Mr NEVADA QUACHITA peanut BOLIVAR uervone 2 TuSeAL00sA sano. . on are ss OTERO 7) : FANNON F. CALNOUN he wa em u oi nat Ue 7) wo =a ~ eamot P< Ly TAA ord ~~ cue as . Youu [Teeny | Lym GRA | newt Tg oa Co TACK oa = eta rr) ee lone Bea een . Sous Tmt po p oA 9) ‘ = OUNG Co TS ore = Re WOLMES wenstn i “4 ont ® ~ =] aa woud DONA AMA EDoy ‘ani 3 = ig edoror _)* ox mi wort > a P\t » ~ = oN MOREHOU' z) Ca i ae ; eas — Fowes a = a ws uN f ZY Iu een vor ae) es rari fae TRADE MARK REG. U.S. PAT. OFF. e jean F si ae (cd LAUDERDAL! eased pas wonreowert wusseLt _ ware arse) — 8 | BOssiER} e- citi MADISON SO BAL i 7 ore E ano / WADI LOWS eC nwa en I @ MoREWS warrin | HOwaRD | i SMITH me bi on reamed s = ® NTCMELL Wo n . e is co : j = | a a | en ee SAMA van ZANT “a - 000 A peraut Ce (ed Hy bag wre ra} wean aa a, Ca Sea @ Fi paso =e © ome « = _ 7 — TENSAS' ae oun . a ary id a es ra, MUOSPETH * ms = . moan] Tarton cS favaees O€ SOTO O ive wor aK ‘e wonnot a . CULBERSOM . EcTOR wiDUND Tau HENDERSON corre | paLt ne cu 7 come s 2 Loving we SASSCOCK | STERLING Cone RUNMELS | COLEMAN | amown\ COMANCHE a tel ee os weRsOn ‘ rotireent A enamee hry Bi WE a Laon a 2 Sao ~~ al ree . ue * Saran [OOO | nt ® WARD FREESTOME i FRANKLIN ee asnimcton = ore ws 3 t , CRANE UPTON TOM GREEN Wis \ MAMILTON ‘Wick Eman CIMESTONE Perey | GREENE cenevaA vous sem, | en to Seale of Miles i * iow omen cee r ese * 7) ® w /™ ‘ HOUSTON M4 WALTON | PECOS ‘5 E x MeCULLOCH a Ss A vane Seay vans for onst eee rarer 4 wat 0 100 * FALLS ns ane waeail AVOYELLES? WILKINSON cot * anne ADIN / carne 4 " | ‘e : LAMPASAS BELL ANGELINA WEWTON® ots st 9 ‘4 CAMOUR ta Ses 'e ROCKET | SCHLEICHER Wenn POReE ROBERTSON oman Tanty POLK TASPER uml per mason al 125 * Surnet mn | cnet iver . ¢ wy gener MAP NO. MASON caro | Aut eaeSON owe "=F owns TuaNo mee iy or gy Bunt UNE FH. erty we Gar ext UAYETTE Tom yy acum eo Ci ose TANG y pret) u I * pRewSTER ent _ mE = f Laon —" ae nt) aot PRESION HARDIN mvreesom ) ACADNA GULESPIE BURLESON CALCASIE . fl FOURTH | ) VAL VERDE EDWARDS CT ee — ai Ate on ee an ad Cal i wal AM 7 a f ’ JUDICIAL DIVISION H 4 ASHTON Pee mA Sal Coun fi a F | + aa RENOALL HAYS BASTROP cn CAMERON heawaiin d } oat ae ORIGINATORS a y 2 TRUS { al BANE Comet CHAMBERS 3 soem yacht | j * ‘oa oe iS a. CLEARTYPE MAPS “sae KMNEY WALDE wom Guapauer /eomzues [aca con TT YORK=" Ss K H + eto praguust wap. Tu lew v ° (rcs) ae MONICE—This us # oi Se of me, - ae grotto eson Doerol, by any proceme 100 PF / = De wiTT 7 RO ere sonal use or resale, without permamion 1 s CR DAMA Fro ; Ecol a ° — RARNES VICTORIA MATAGORDA ' rend sent Dit TASALLE ewe bee Se ; oasis v sue KAUAI / . e are w e HIRD / H A W A I I ° —— m sO ore acs aRaseTe 7 10N LIVE OAK Rif 4 JUDICIAL DIVISIO! / HONOLULU cr) — amore KAUAI / PART J yi DA Yaw wells sg os 4 UMA / oe re’ MUECES = oo g Ae) / / ; »~ ate ? ; { / OAHU / MAUI I é * ae n ; / f Ye x HLEBERG wt x ' / ; / KALAWAO a ’ poate Ts woos Tgaroons NENEDY ° : pe - a PY, a ° ARs 7 MOLOKAl ] ) * : - a > we ee a JUDICIAL Scale of Miles gp / 5 ? 0 Anas a STARR WDA 2 i roy ‘ / i fe * wu ay ead ‘ i 4 o? KAHOOLAWE \% ° if 1% 0»,; HONOLULU awe HAWAII A =’ % — a Y PART : 7 f = >) “Nee, _ a 9 ee LISIANSKI, : i . LAYSAN ‘ i | Pid - , 7 4 ‘ — = 4 _" . me tS 4, >. ALEUTIAN ISLANDS fy 1 & : 5 > és *. * THIRD JUDICIAL DIVISION NECKER 2 ‘ainend 5 ai Ma a fi org" ite j . York, No. 14324 | 2 mofay a 2,7 f >) ight, American Map Co., Inc., New Seale of Miles Ae 6 ».. ae eee HAWN ms Fy g Copytigh ee en ee 4 * 94d" Miles 300 KAUAI 7 ae Re eRe RE ~ , | caiaanael HONOLULU | a aie BE te >. . ? aid 17 18 19 20 21 : : : O 7 3 10 it 12 a i 15 I 16 ] L if it 8 PHYTOLOGIA is financed entirely by its contributors, each one paying : advance for entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon — the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his = * ae expenses and sharing in the putes if any accrue. Each canbe. consists of not less than 32 pages. All manuscript accepted fe Ate ding with rch nal nsf tan, ny seal PHY TOLOGI Designed to expedite botanical publication Vol. 11 December, 1964 i ee D b America. NEW SOUTH AMERICAN IcHNANTRUS /°°75 Jason R, Swallen ICHNANTHUS FIRMUS Swallen sp. Culmi 1.7 m, longi; ee teiekasa multo ge OPP Papilloso-pilosae; laminae 15 cm. longae, 1,-1.8 cm Pepilloso-pilosae, marginibus scabris; pani cul a 37 cm. Moe s uma p acuminata; gluma secunda lemmate sterili longior vel Seethie Snervis, scabra, ge greys longe pilosa; fructus 3,2 mn. longus, alis latis 0.7 mn. longis. Perennial; culms straggling, 1.7 m. long, papillose-pilose below the nodes; internodes mostly short, the sheaths much long, 1.4-1.8 om. wide, pia Llosa oe tepeteed on both sur- faces, Amcgels 80 just above the ligule, with white very 7 mar, h narrowly as nding branches as much as 13 cm » the lower gee ratios distant, the axis and ‘ennai papi llose-piose wit Spreading hairs; spikelets ,5-6.5 mm. long, nant doth callie, ac from shorter than the second glume and sterile lemma to exceed~ ing them as mich as 1.5 mm,, 5-nerved, scabrous, sparingly Covered, especially on the margins , with long papillose-based hairs; second glume and sterile Lonma equal or subequal, S-nerved, the glume acu te, scabrous, hairy toward the tip, the emma scarcely scabrous with only a few hairs at the summit; ne 3.2 mm. long, yellowish, with broad obtuse wings 0.7 mm. os in the Herbarium Bradeanum, No. apes collected at Manaus, Amazonas, Brazil, January 26, 1963, . Santos 1499 St al. A piece of the type is in the U. S. ibcnay Webern. ICHNANTHUS RECLIVIS Swallen, sp. no Culmi ranosi glabri 1.2 m alti; vaginae internodiis breviores, eae ramorm pilosae vel cenit inac~ol iteen 3 laminae yodue ad 12 cm, “enene 2 cm, latae, glabrae; ee ot cm, longa, ca. 5 om, lata, ramis adscendentibus Ongis; spiculae h-l.2 mn. longae, appressae, bieMigedibel tates; 3 gluma prima spicula 1/5 brevior, 3-nervis, — plow paucis ca. 4 mm, longis; fructus 3 mm, longus alis Perennial; culms erect at the woody ba i. % aed branch- sng, glabrous, up to 1.2 m, tall; sheaths, at least t hoe low = ®S, shorter than the internodes, glabrous, those of t Fanches pilose or papillose-pilose with long hairs, aa very i Povrenend pubescent greg blades as much as 12 cm, long, ®, glabrous, or younger ones of th Pubescent toward ee bears panicle 17 cm. long, gi 5 cm, 145 146 PHYTOLOGIA Vol, 11, 80.3 wide, with narrowly ascending branches, the Wi ones 7 cm long; spikelets short-pedicellate, appressed, -l|,2 mm . long; first glume },/5 as long to as long as the i elet, 3-nerv d, e about ) mm, ss secom glume ae es e lemma equ peices ong. n the U. S. National Herbarium, No. Stat col- lected on Sende rocks among brush, between Bonito Rondonopolis, Matto Grosso, Brazil, * April 8, 1930, oy Agnes Chase (No. 1189). ICHNANTHUS SUBINCLUSUS Swallen, sp. nov, Culm i duri, lignei, ramosi, infra nodis papillosi vel papilloso-pilosi; vaginae internodiis longiores, papilloso- aRge FN ee ca, 5 mm, longis; laminae 7-10 cm. longae, 1-1.7 cm, latae, anguste ovato-lanceolatae, infra dense spige ing supra scahian pilis longis dispersis; panicula 20 cm, longa 3~ on Joe densa, ramis adscendentibus vel appres sis; Spiculae 5-5,5 mm. longae, brevipedicellatae, Pas iano gluma prima pares 1/5 brevior vel spiculam aequans, 3-nervis, scabra; shh Neg 3.6-3.7 mm, longus alis 0.6 fs longis. Perennial; culms hard and woody near the base, branching at the middle node, with several (6-10) internodes of roughly equal length, papillose or papillose-pilose at least below the 1 with spreading hairs abo mm, long; bla 10 cm, long, 1-1.7 cm. wide, narrow ate-lanceolate, d ly and softly pubescent on th r surfa finely scabrous on the upper, often with long scattered hairs, especial the base, the margins finely scabrous; panicle rather ely flowered abou O cm, long, 3- cm, » partly enclosed in the up cent at the tip; fruit 3.6-3.7 mm, long, with small wings 0.6 mm, long. Type in the U, S. National eo No, 2382897, col- lected in cerrado, Campo Mourao, Paran4, Brazil, 625 meters, February 3, 1962,” by G. Metecthech (No. 8851). ICHNANTHUS pain sai Swallen, sp. nov ecti vel decumbentes ca. “65 om alti, nodis breve barhatis sy Licernoatis pilosis; vaginae dense pilosae; laminae 6-7 cm ae, 8-12 mm, latae, infra dense pu oP aac supra glabrae basi pilis longis praeditae; panicula 17 cm, longa, 5-6 cm, iat ramis solitariis usque ad 6 cm. longis; spiculae 4.8 mm, longae, appressae; gluma prima spicula 1/5 brevior vel paulum rr oe —— vel acuminata; fructus 3,3 mm, longus alis 0,7-0.8 mm, longis. =) ° 1964, Swallen, South American Ichnanthus 147 Perennial; culms erect or decumbent at the base, about 65 cm, tall, with shortly bearded nodes, the internodes “per opeeer pilose, especially below the nodes; sheaths nearly as long the internodes, densely pilose, the hairs becoming longer toward the summit of the sheath; blades 6-7 cm. long, 8-12 m., wide, acuminate, spreading, densely pubescent bepasth, pearls ous above, with a few long hairs at the base; r cm hes, the lowermo oer lets 4.8 mm, long, app pressed; first glume h/5 as slightly longer than the spikelet, acute or eerie e, ggg scabrous on the keel; second glume le lemma sube goperved, the glume acute, the yi blunt or rounded; fruit - long, e in the U. S, National Sieg No. 2304289, col- lected re campo cerrado, Fda. Morungava, Rio do Funil, Parané, ies "+ pai! 12, 1958, by G. Hatschbach and R. B, Lange No, 5276) This species is richng related to I, subincl hy aay. which has a sg r and denser ponies STN a an oe she th, erect o r narrowdy ascending panicle branches, d larger spikelets to 5.5 mm, long. ICHNANTHUS cose bes Swallen, sp Culmi erecti, 165 cm. nti, Mae vaginae internodiis mailto et habs laminae 12-22 cm, longae, 2,2-3.3 cm. latae, sm 2 cm, longae ramis patentibus ad basin ipingoe ahaa remoto usque ad 16 cm. longo; spiculae 4.5 mm appressae, nervosae; gluma prima spicula eae. ea pila 3-nervis; lemma sterile ened ations fructus 3.6 mn, " alis cieatricibus reductis, Per al; culms erect, 165 cm. high, glabrous or somewhat iabenccnt at the nodes; sheaths much shorter than the inter- odes » @labrous, densely pubescent in the throat; blades 15-22 mm long, 2.2-3.3 cm. wide, somewhat narrowed toward the base, very densely scabrous; Spikelets .5 mm g, appressed to the branches, strongly erved; first oo cy, 2-2/3 as — as the spikelet, 3-nerved, Scabrous near the tip; second glume and sterile lemma equal, S-nerved, the glume acute, the pee io cucullate Scabrous near the t tip, otherwise amare 3 fruit 3.6 ma. long, yello ona » the wings reduced to Type in the ZH ee National * aro pag No. 1645512, col- lected in a wood a pera, Pernambuco, Brazil, April 2h, 1935, by D. Rento id Piskel “(No. 3855). rg GLABERRIMUS Swallen, sp. n Culmi erecti, glabri, 60 cm. nti; ; “ligula obsoleta; laminae ca, 16-22 om, longae, 1, 8-2 em, latae, glabrae; panic *recta, 25 cm. longa, 2 cm, lata, spiculis appressis; spiculae 148 PHYTOLOGIA Vol. 11, no. 3 h mm, longae, purpureae, glabrae; gluma prima spicula 1/3 brevior, vel pre aequans; fructus 3.) mm, longus, albus, alis latis a; 5 mm. longis. Per Bos culms erect, 60 cm. tall, glabrous; leaves with no differentiation a sheath and blade, the blade about 16-22 cm. long, 1.8-2 cm, wide, acuminate, narrowed toward the base, glabrous, even the white margins; ligule wanting; ores erect, 25 cm. long, about 2 cm. wide (possibly wider in matur panicles), with appressed branchlets; spikelets mm. long, oar tinged with violet, glabrous or slightly scabrous; first glume 2/3 as long to nearly as oat as the cae ei second rant and sterile lemma equal, the glume a emma rounded; fruit 3.) mm. long, Pn the eis broad, 1.5 mm, g Type in the U. S. National Herbarium, hs 2382261, col- lected in Jalca Zone, 3 km, south of Pornacocha, east of Shipasbamba trail, Peru, 200 m. altitude, ip tame 20, 1962, by J. J, Wurdack (No, 1 a Ichnanthus glaberrimus is related to I. eens Pe eaiert but in that species the cece: are 5 mm. long, long scattered hairs, and the leaves have a aehtbate ree hg between sheath and tind; ICHNANTHUS te oy hea Swallen, sp. piu em. longi; vaginae intaeubdlse Tae glabrae, in colle dense villosae; leminae 11-18 cm. longae 1,6-2,.6 cm, latae, ad basin longe-villosae; panic om. ah spiculae 6-6,5 mm, longae, glabrae; gluma prima acuminata, — , mm. longa; fructus 3.5 mm. longus Slis fir firmis 1 mm. nni culms more than 60 cm,, perhaps more than 1 m. long; sheaths yiecter than the internodes, glabrous with a densely villous collar; blades flat, somewhat asymmetrica 11-18 em, long, 1.6-2.6 cm. wide, narrowed toward the base, but ~~ branches, the lower as much as 15 cm, long with short ng few-flowered branchlets; spikelets mostly in pairs, enpreasisd; 6-6.5 mm. long, glabrous; first glume acuminate, 3.5-4.5 mm. long, 3-nerved; second glume and sterile lemma : nate, or the sterile lemma somewhat blunt, 5-nerved; t Tapera, Pernambu » Fe 16, 1930, by D. Bento J. Pickel sassy 2267). roarhatse at Tapera, also by Pickel, Nos. 1361, ICHNANTHUS te Swallen, a ei Culmi graciles, lignei, eines "i-2 m, longi, 1-2 mm. assi; vaginae pecateia saabtorsl marginibus conspicue br pes e ligula usque ad 5 mm, longa; laminae 8-11 cm, longae, 196k, Swallen, South American Ichnanthus 149 0.8-1 cm, latae, glabrae vel plus minusve pubescentes; ; paniculae 5-6 cm, longae, 1-2 ~ Miah densae, axis et r dense villosis; spiculae 3,.5-3 . longae, erevibediballitels 3 gluma prima spicula 1/ ac erovige’ pilis longis preedita; fructus 2.7-2,8 mm. longus alis 0.) mn. longis. slender, woody, 1-2 mm, thick, branching, 1-2 m peed sheaths longer than the ebasrtred: conspicuously villous e margins and across the back of the co » the hair ak falling off; ligule a prominent line of rather coarse airs, as much as 5 mm. long; blades mostly 8-11 cm. long, 0.8-1 em. wide (or the uppermost smaller), glabrous or somewhat Spikelets 3.5- mm, long -pedicellate; first glume acuminate, 2/3-3/l as long as the spikelet, 3-5-n irs ne he tip occasionally the base, the hairs shorter downward; and sterile lemma equal, con 5-nerved, the glume with a few hairs near, the tip, the sterile lemma glabrous or nearly nef ade 2.7-2.8 mm, long, the wings - long. Type in the U. S, og Pee goer ae No, 196381, col- lected between shrubs of thicket, Campo de C ongonhas, S&o Paulo, Brazil, Obteber 23, 1948, by D. Bento J. Pickel (No, 5893). ICHNANTHUS CAMPORUM Swallen, sp. nov Culmi erecti 110 cm. alti, dense pilosi vel subglabri; t; ramis 6-8 adse Denes spitulae 5-5.2 mm. longae, inflatae, Glabrae; gluma prima a, 3-nervis, 3-3.5 mm, rege en fructus 3.2 mm, longus, neaties ahs: cicatricibus parvis inc ager Perennial; culms erect ag hort te 110 cm ’ 2 mm. long, elliptic, acute, white, eiosth Gd abtadog? Scars small and inconspicuous Type in the U. S. National Herbarium, No. 14487hh, eae lected in low brush, sort clay ott between Viannapo bv 3 " — ee (No. 1127h). M itional material examined: Brazil: Goias: Pirineus acedo “13783 between Viannapolis and Ponta Funda, Chase 113363 Anna ith saree Chase 11343. Some specimens show a relationship achnanthus procurrens (Nees) Swallen, comb, no cum tines Nees), in that the spikelets are hispid and/or the ades are shorter and relatively broader, but the spikelets 150 PHYTOLOGIA Vol. 11, no. 3 e 5-5.2 mm. long, and mines or i hispid, In I, procurrens the spikelets are < -3.5 mm These specimens, which seem to be hybrids, are pers TE ity a 11386, both collected near Annapolis, ICHNANTHUS RECLINATUS Swallen, sp. Culmi decumbentes ramosi, re a A 3-5 cm. longis sparse pilosis; vaginae internodios subaequans, dense papilloso- pilosae; laminae )-8 cm, longae, 5-10 mm, latae, infra et supra . e villosis; spiculae 5 mm. longae, appressae; gluma acute vel breviaristata spicula 1/h-2/5 cht pags fh ny 3. 5 - longus alis 0,6 m, gis. see more tha ae one S SETAERL RSs brane has D ard and woody, about 1 mm, thick, with numer internodes 3-5 cm. stir a sa densely Seeey Tis races ait erect or ascending, 4-8 cm, long, 5-10 mm, wide, slightly ovate, softly peveaceat, on both surfaces; inflorescence 3.5-5 cm. long with a few s r pedicels densely villous; spikelets 5 mm. long, appressed; first glume acute, apiculate or short-awned, 3/5-3/h as ong. as — Pskasuced p 5-(3-) nerved, sometime s bearing a few ong t margins; second glume and peta lemma acute or rather pine, 5-(7-) nerved, usually with a few hairs; fruit 3.5 m, long, pale yellowish, with wings about 0.6 mm, long. = nthe U. S, National Herbarium, No. 2380658, col- lected in A AR Alto Matador, Rio do Sul, Santa Catarina, ay R. Klein (No. 7257). One other specimen was examined which can be referred to this species: Brazil: "Rio de Janeiro: Nova Friburgo, Curran 635. ICHNANTHUS RIPARIUS Sw. wallen, sp. ni graciles, ramosi, ic ae Ag plus 50 cm. Np subglabri; vaginae eaieae: laainae 5-8 cm. longae, 5-8 mm latae, glabrae, marginibus scabrae; panicula 3-6,5 cm, lon ramis dense raiders ta spiculae 3 mm, longae brovipedicel lataes gluma prima acuta spi subaequans "pilis paucis praedita; gluma secunda lemma sterile aequans, 5-nervia, apeci gee ple joggaa fructus 2.5 mm, longus alis cicatricibus 0.) mm longis reductis. slender, branching, straggling, more than 50 cm. long, merprateie. or with a few long hairs; sheaths about as long = base; spikelets 3 mm. long, short-pedicellate; first glume cute, 3-nerved, beat get long as the spikelet, with a few ts at the tip; second glume and sterile lemma equal, 196k, Swallen, South American Ichnanthus 151 strongly 5-nerved, with a tuft of hairs at the tip of the pos de c 1894, by eee and Edwall (No ot This species collected in the same general locality by A. C. Brade, “April 2, 1951 (No, 208)1). roa. BACULARIUS Swallen, sp. Culmi graciles, radicati, 1m oh ngi ve vel longiores, internodiis pilosis; ae internodiis multo breviores, pr rae, marginibus ciliat os © 4 ° a o> ap EE Be ct ~ ' mM . qd 3 panicula 15 cm, longa, r ramis piaseclactttaa vel patentibus usque ad 9 cm. longis; spiculae 5-5.); mm, longae, glabrae, appressae; 3; fructus 2.7 mm, longus alis cicatricibus 0.5 mm, s reductis emia}? ; culms slender, tangled, leaning on other vegetation, over 1 m, long, with long stilt-roots from the lower nodes, the internodes pilose; sheaths much shorter than the internodes, glabrous across the back, with ciliate margins and a Nacesiy pubescent collar; ligule an erose or cilia membrane 1-2 mm, long; blades 7-11 cm. long, 10-15 mm, wide, t little longer, acute or acuminate, 5-nerved, scabrous the tip; fruit 2.7 mm. long, the wings reduced to a gan Type in the U. S, National Herbarium, No. 1500985, col- i te in eens colony, rain forest, southeast slope of Serra do Caparo, Esperito Santo, Brazil, November 23, 1929, by Agnes Chase (No, 1007 79). Also collected on Serra do Caparao by Inez Mexia (No, 4011). i TWO NEW GENERA OF OLYREAE FROM SOUTH AMERICA Jason R, Swallen Two ag toon fas related oeoere. of the Olyreae have been known androl from South Ame a Stapf was described in 1906, and Reitzia pl ep in > vhe latter from Santa Catarina, pommel hon The other species of Piresia is based on oye ica Doell, described from French Guiana in 1877. ad is : e most widespread species of these four genera, ranging Trinidad to Surinam, and Pernambuco, Brazil. All “a species are rare in forested regions, judging from the small number of specimens which have pes collected. The may have been overlooked, however, becaus inflorescences are usually hidden under’ the foliage and a yd apparent. Key to the Genera Glumes of fertile Bde er oa 3-nerved. Nerves of a of fertile spikelet ahi brs pubescent; " alee i form Nerves of climes, oe fertile spikelet Faint. Se feat ae es ous culms all a Glumes of fertile spikeist hbrcabos Stamens 2; fruit glab cone sess nees oe 16 Ouran Stamens 3; fruit cahasoante’ Culms bulbose at base. hh. Bulbulus pills albidis villos Culms of two cints, the fertile coe short, bladeless, or with very much reduced blades, mostly hidd en underneath the 2 t Nobo spikelets about 3 mm, lon sile, 1 (sometimes 2) below each pistillate spikelet; gi rerna spikelets 1 or 2, terminat- ing the main culm and branches; fruit elliptic, sessile, about 6 mm, eee the lemma and palea white-villous. ype species: Piresia goeldii Swallen. 1964 Swallen, Two new genera 153 Key to the Species Blades pacer ina L-S cm. long, not more Be, 7 on each culm, P. sympodica Blates” ie AR lt mostly less than A ae . long, as many as 20 on each culm, evenly spaced...2,. P, goeldii 1. PIRESIA SYMPODICA (Doell) Swallen, comb, Olyra dica Doell in Mart, Fl. me # 322. 1877. Radda a ae Hitche. & Chase, Contr. U. S. Nat. Herb. Material examined: Trinidad: Caparo, Broadway 2375 (typ nies biformis Hitche, & Chase), 8122; Tabaquite, ai iekosk er urinam: ps pevifoaia arts French Guiana: Karouany, fie ae 2. PIRESIA GOELDII Swallen i foliosi erecti, 7 7k (25) em, alti, retrorse pubescentes, nodis den BeOS F ase 2-1 ee pg’ Superiore congestis, L. rc em, longis, 2.5-6 mn, tc fertiles 5 cm, alti, geniculati, spiculis fo aye 2 appressis, 6 mn, ieneis. glumis 3-nervibus; fructus plus minusve obovatus, geo, ee palea pilis albidis villosis. mnial; leafy culms in small tufts, 7-1) cm. erbot wis spreading toward the top, densely Dei ey eaeeeonl in lines, the nodes spreading-pubescent, with 2-18 dis ener shorter than the internodes, glabrous, bladeles upper ones closely overlapping, Bi lose i the margins, the oe pilose to Blabrous; blades as much as 1,7 cm. long, 2.5-6 mm. wide, firn, ovate, sparsely bate pel par fertile culms about tall, bladeless, with 2 appressed pistillate spikelets and a or 2 stami nate spikelets beneath each pistillate one; st yr hge pe B mm, Jone, with 3 stamens; pistillate spikelet shout ? - long, firm, acute, glabrous, the glumes Ea h rust slightly obovate, the lemma and palea villou e in . National Herbarium, No. 1039650, col- lected in sandy ‘is umous places below second growth forest, Macuary-ass mde Railroad, Paré, Brazil, June 191h, by André sane’ (No. Additional ames examined: Brazil: Para: Breves, Pires, Froes & Silva 908, Surinam: Soderstrom 5li291. @. REITZIA SMITHII Swallen, Sellowia 7: 8. 1956. i.e erial ned: Brazil: Santa Catarina: Azam buja, gaits 6123 (type bag pm Reitz & Klein 122 Reits Soa Sdo Paulo: Morro as d3 Pedras, Brade 8107 (I.A Ne). 15h PHYTOLOGIA Vol. 11, no. 3 3. DEAMDGOL TRA BICOLOR Stapf, Kew Bull. Misc. Inf. 1906: 20h. 1 "Native country unknown, Raised at Kew from seeds communicated by Messrs, Sander and Son." Known only from type material, It is undoubtedly tropical American re: other genera, closely related to it, have been found t 4. BULBULUS Swallen, gen. no liosi, inflorescentsbus terminalibus; spiculae in escence terminating leafy pairs of spikelets, one of each pair pis sti llate, er s e, spikelets 7-8 mm, long, the glumes acute acuminate, 7-nerv the fruit 5.5 mm, long, villous, the marine 4 nrolled erering- tl the palea; staminate spikelets 5 mm, long, cigntodh: containing 3 stamens BULBULUS NERVATUS Swallen, sp. caespitosi, erecti ver tanbeaauitee: retrors truncate-r Ginked or slightly cordate at the base, glabrous 0 ae Ps romi- nent, acute or acuminate, ecumahak relaveds fruit 5.5 . feos Light-yellowish, pilose or hl Yous the margins of the nrolled covering the palea, in the U. S, National Herbarium, No. 1612746, col- ese in high sandy forest from Caxias to Barra do Corda " for 3538) February 18-26, 193i, by Jason R. Swallen o. ditional material examined: Brazil: Pard: Santarém, duct ten 3284; Castanhal, Estrada de Ferra de Braganga, Goel 303, Territory of Amapd: Rio Olapoque, Froes 26759 CE, A.N.)- MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXV Harold N. Moldenke ENA SESSILIS (Cham.) Kuntze Kuntze (1898) continues: "Schauer in DC. prod. und Flora bras- iliensis stellt noch ex descr. snp. dritte Art: ve montevicensis Spr. dazu, die aber nach dem Original-Exemplar gar keine Aehn= lichkeit- hat. Bei V. decurrens und sessilis sind die kopfigen Flat ran sehr d. dicht, oben zur Blttthezeit fast flach, 2 cm breit verjtingen sie keilig nach unten: die Bracteen sind flach, a rent he 1-1 1/2 cm lang in eine lange Spitze auslaufend ie haarig gewimpert; die schlanke Corollenrthre steht 1 cm — us dem Kelch hervor; die Blatter sind lineallanzettig (1: iss a und bis 8 cm lang. Bei V. montevidensis Spr. sind die kopfigen Inflorescenzen locker und mehr oblong Yhrenartig, nur }- a mi breit, die Bracteen sind kahnftrmig oval acut Fed nepr: ae m lang, nicht gewimpert; die breite Coro ret d Kelch kaum; die Bl&tter sind oval bis sderiianestate (1: 1 1/2~2 en nur — 1/2 cm lang." Verbena sessilis bits d swampy meadows, wet soil and wet oe grassy fields, shrubby campos, and wet taligrass pig ot and is found along rivers and highways. ci ance calls Common", It has been collected in anthesis in Jan it be ember. Rosengurtt (2913) de describes : as "Hemicriptéfita, florece a fines Pp vera rano abita os, ara” and cites his B.2351, OH. baffad s 1849), and OH.18L95. Schnack & Covas (1916) also cite Rosengurtt a (Si), while Briquet (190) cites Regnell A.2039 from Para- s “foliis sessilibus an- conspersis, bracteis ne no 1 brevioribus". original de escription is ";) sessilis foliis sessilibus lanceola- hoa obsolete serrulatis supra pagina strigulosis. E Brasilia . ridionali utramque varietatem misit Sellowius." Although these Uthors agree that V. sessilis differs from V. stellarioides in having it its leaves merely sessile, not decurr ent, joe peeureniay ar thar ty serrate, ponent rag pete Plo and the pes tle e cal is the only y dependable ahanaatar for separating t On the other hand, ~ Efiowing plated ta were distributed or cited by m 5 rreiioukly (H ~ a Ve Stes Sorgen 3773 Hy . Osten 222)1)], Kuntze s.n., Morong ‘a P (Herb. assler 1680; Herb. Oste salleeas > A. Ge “Sms 285 (Herb. ct co) = | o ct iB ct oa 4 i ° an 928 i) o 4 p Q oS KO o ct a 156 PHYTOLOGIA Vol. 11, no. 3 so 23149] & 1468. n all, 30 ‘erbarium specimens and 5 mounted photographs, in- sonata the types or phototypes of all the names involved, have been examined by me, Citations: BRAZIL: Rio Grande do Sul: Rambo 25786 (N). State undetermined: Sellow 1563 [Macbride photos 1747] (Kr—photo of type, N--photo of type, N—photo of type). PARAGUAY: Fiebrig 1,031 (Bm, V); Hassler 278 (Bm, V), 2478a (Bm, Cb, N--photo, V, a ee photo); Kuntze s.n. [Nord Paraguay, IX.92] (W—701065) 5 Lindman 4.2039 (S, S); Malme 928 (N, S, S); T. Rojas 174 (S). URUGUAY: Arechavaleta 1 19 (Ug, Ug), 3136, in part (Ug), sen. [Herbd. Osten 18494) (Ug); Berro 6721 (N), 7477 (N); Gallinal 742 [Herb. Osten 23356] (Ug); Gallinal, Aragone, Bergalli, Campal, & Rosengurtt A.1110 (N); Herter s.n. wp pee Osten 1649] (Ug), Son. {Herb. Osten 18195] (N, Ug, Ug). INA: Formosa: Jorgensen 2477 [Herb. Osten 1367] (Ug, pitrrerrerinon I. Morel 1707 (N)e ice SETACEA Perry, Ann. Mo. Bot. Gard. 20: 339--3h0, pl. 15. 33. Bibliography: Perry Bot. Gard. 20: 2h9, 250, cont 339--340, 356, & %2, pl. 2B. "1933; A.W. ores Ind. Kew. 93 at 1938; Worsdell, Ind ort cg 2: 486. 191 aaaune: eogr. Distrib. Verbenac. 1), 19 & 102. 19125 te hy Alph, List Cit. 2: 167 cass) ot 3: 833. 1949; Moldenk Known Geogr. Distrib. Verbenac., [ed. 2], 33 & 199. 199; Yoiden- ke, Résumé ho & bth. 1959; Moldenke, Phytologia 9: 176 & 177. 19635 Moldenke in Shreve & Wiggins, Veg. & Fl. Son. Des. 2: 12h6. Tlustrations: Perry, Ann. Mo. Bot. Gard. 20: pl. 15. 1933. Herbaceous or suffruticose at the base; stems decumbent or de- th short~pedunculate; bractlets linear-lanceolate, about 3/l as long as the calyx, soft-pubescent, ciliate, subulate-setaceous at the apex; calyx about 6 mm. long, villous-pubescent, its teeth 2 mm. long, about half as long as the tube, subulate~setaceous; corolla- tube about 7 mm. long, pubescent outside, the limb 6—~7 mm. wide; anthers not glandular; mature cocci not know type of this rare species was pe ollected by Carl Albert Purpus (no. 195) among rocks, at an altitude of 1600--1700 feet, at Calmallf, Baja California, Mexico, between January and March, 1898, and is deposited in the herbarium of Pomona College ed Claremont, California. The species is know thus far 0 the type collection and was originally misidentified and asatest- ed as V. ciliata Benth. Perry (1933) says "The gross habit © 1961, Moldenke, Monograph of Verbena 157 this plant is very like that of V. Gooddingii var. nepetifolia. Unf, are in anthesis, the inflorescence is immature and only a few flowers ant he so different in habit it would seam as if this were perhaps only a Superficial resemblance. The species is readily Batings shes by its “papa habit and Song setaceous calyx~teeth." e herbarium specimen and 3 mounted illustrations and elippings, ineiudite the type, have been examin me. Citations: MEXICO: Baja California: Purpus 195 (N--photo of » Po-~63919-~type, Z--photo of type). MOUNTED CLIPPINGS: Per- ry, Ann. Mo. Bot. Gard. 20: 362, pl. 15. 1933 (W). VERBENA SHREVEI Mo oldenke, Phytologia 2: 26-27. 1 Bibliography: Moldenke, Phytologia 2: 26-=27. ey Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1), 19 & 102. 192; H. Ne. & A. L. Moldenke, sh hye 23 83. "19185 ldenke, Known Geogr. Dis- trib. Verbenac 2], 33 & 199. i9l93 Moldenke, Alph. List cit. 3: 933 “asksh, ae i dels. 329195 Shreve & Wiggins, Camegie Wash. Publ. 591: 1 » ind i: 363. 1953; Moldenke, i Pans: rg ap & ih. 1959 *Woldenke, Phyto- logia 8: 196,'1962; Moldenke in Shreve & Wiggins, me & Fl. Son. eh 12? 1246. 19645 Moldenke, _Phytolo gia 10: 306+ 196). ual herb; stems decumbent at the base, slender, obtusely Netcasts, more or less dacedig.a preading-pubesc ent with coer dular which are ? Pubescent; leaves decus sate-opposite, petiolate or the uppermost aaa petioles very slender, 1--10 m, long, more or less ensely hi calyx tubular 1 rather densely glandular-pubescent ond also sot ar aimee centered widte-uiveutilous; corolla small, (8 m long, slightly projecting from the “ae. its tube minu- ely puberulent at the apex outside, the limb about l mm. wide. The type of this species was collected by Forrest Shreve (no. Tuy) —~ in whose honor it is named — at an elevation of 1900 f feet, 158 PHYTOLOGIA Vol. 11, no. 3 19 miles northeast of Comondén [=Comondi], Baja California, Mexi- ic mesas. Herbarium material has been misidentified and distrib- uted under the names V. ciliata Benth., V. gooddingii Briq., and V. pumila Rydb. It is 8 known thus far only from the type locality. In all, 5 herbarium specimens, including the type, and 2 mounted euithgnasia= have been y me. Citations: MEXICO: Baja California: H. C. Orcutt s.n. [Pinery, 10-6~1882] (Mi); Shreve 7119 (Du--265832—~isotype, Fs-——type, Mi— isotype, N-—-isotype, N——photo of type, Z—photo of type). VERBENA SIMPLEX Lehm., Nov. Act. K. Deut. Akad. Naturf. Halle 14 (2): 52h. 1825; Ind. Sem. Hort. Hamb. 17. 1825. te) Verbena nymy : ‘ . 1803 [not V. angustifolia Mill., 1768, nor Lam., 1947]. Verbena rugosa Muhl. ex Willd., Enum. Hort. Berol. 2: 633. 1809 [not V. sa D. Don, 1836, nor Michx., 197, nor Mill., 1768, nor Sweet, ]. Verbena integrifolia Michx. ex Hook., Comp. Bot. Mag. 1: a 1836 [not [not V. integrifolia sessé & Moc., 1889]. Verbena ru~ gosa Willd. ex Hook., Comp. Bot. Mag. 1: 176, in syn. 183%. Yer- bena angustata Michx, ex Moldenke, Suppl. List Invalid Names 5 in syn. 1941. Verbena angustifolia L. ex Moldenke, Suppl. List Invalid Names 7, in syn. 1941. Verbena amsifolia Michx. ex G. L. Fisher, Am. Bot. Exchange List, sphalm. 1946. Verbena langusti- folia Warns: ex Moldenke, Alph. List Invalid Names Suppl. 11 25, in syn. 1947. Verbena cuneifolia W Wallberg ex Moldenke, Résumé 363, in syn. 1959 [not V. cuneifolia Hort., 185, nor Raf., 1808, nor Rufz & Pav., 1798]. Verbena angustifolium Michx. ex Moldenke, Résumé Suppl, 4: 14, in syn = 1962 Verbena angustifola Mx. ex Moldenke, Résumé Suppl. li: & in syn. 196k. Bibliography: me -, Gard. ’pict., ed. 8, no. 15. 1768; Michx., Fl. Bor.-Am., ed. 1, 2: 1s, 1303; Poir. in Lam., Encycl. Méth. Bot. 8: 549. 1808; "yuna Emm, Hort. Berol. 2: 633. 1809; Pursh, : £17. 181h; J. Sm, in Rees Cycl. 36: no. 1. 18175 =2 Repert. l: Sus. 1847; Ae We Chapm., Fis Sout «Ue Se, ede pre i, 307+ 1860; Bocq., Adansonia 2: 113. 136i 186 Bocq, Rev. Verbenac. 33. 1862; A. W. Chapm., Fl. South. U. S., ed. 1, pr. 2, 307 (1865) 6 196) Moldenke, Monograph of Verbena 159 and pr. 3, 307. 1872; A. Gray, - Fl. N. An. & (1): 3%. aris Pickering, Chron, ght Pl. mos & "1220 1879; Pe, South. U. S., ed. 2, pr. 1, 307 (188), pr. 2, 301 (2887), Pr 3, 307 (089) and a . 307. 1892; Brig Engl Nat. Pflanzenfam, 1 Forest & Gard. Bot. a. "1895; Jacks. in Hook. f. & Jacks., Ind. Kew, 2: 1178 & 1179. 1895; cules Bull. Ohio Agric. Sta. 83: ak als Britton & Pel ae Tbes ¥Rlo pred. hy 30°71, Pig. Ss. ys “Uh. 1899; Selby, Bull Sin. = IB 360" 19065 ‘laea » Bot. Gaz. 43: 57. 1907; Robinson - in A. Gray, New + Botep ed. 7, 689 & 92h, fig. 879. 1908} Ue A. Day, Cheek List "128, 1 908; L. H. Bailey, ° 7d; Britton & Br., Illustr. Fl., ed. 2, 3: a fig. 3555. 1913; & 70, & 21: 133. 1921; K. V. 0. Dahlgren, Bot. Notiser Lund 1923: 10, 19233 A. B. Seymour, Host Ind. Fungi N. Am. 597. 19295 G. T. Sier, Trans, Kans. Acad. Sci.» 3h: 261. 1931; Schwencke, “_ Untersuch. Verbenac. 35 hy & 17--20. 1931; Stapf, Ind. Lond. 6: 8. 1921; Benner, Fl. Bucks Co. 255. 1932; Perry, Ann. Mo. Bot. Gard. 20: 2i6~~2h,6, 260, 2 i Journ. N. Y. Bot, Gard. 35: 133. 19343 Junell, Symb. yo ae kt 16h & 171. 193k; E. J. Palmer, Ann. Mo. Bot. Sane 1935; Patermann, ho Zytol. Verbenac. 6, 7, 1 oh. : - 19353 Demmen, Cytologia 7: 161163, fig. 1h. nce Mo oldenke, Revist. t ° Common Names Z. 190; Worsdell, Ind. Lond. Suppl. 2: 486-187. 19h; Moldenke suppl. List Invalid Names 7& 10, 19); Gates, Weeds Kans, ,2'2 220, 192; C. N. Jones, Ohio Journ. rw hl: 3h0. 19ha; Schnack K, Anal. Inst, Fitotée. Sta. cat alina y: 18 & 19. 1 ’ ’ own ], ill, 7k, 76, & 102. 1942; Moldenke, Alph. List Invalid Names h5 & 50. 192; Moldenke, Known Geogr. Distrib. Verbenac. Suppl. 1: 1— C. 1945; Me\denice , Bot. Gaz. 106: 158-160. 1915; Moldenke, Castanea ioe 36 ma 19453 Ge. N. Jones, Fl. Il]. [Am. Midl. Nat. Monog. 22) 216, apkes Darlington & Janaki Ammal, Chromosome Atl. 270. 160 PRs.) os OC Ts Vol. 1, no. 3 19k 5; al Kriebel, Yuncker, & Friesner, Proc. Ind. Acad. Sci. ae 95. 195; Tryon. & Moore, Rhodora 8: me 116, & xiv. 19h6; L. Fisher, Am. Bot. Exchange List. 1916; *kKriebel, Yunck- oe & Friesner, Proc. Ind. Acad. Sci. 55: ie. “19h6; Moldenke, Alph, List Cit. 1: 3—5, 7, 11, 13—16, 18, 25, 27, 28, 31, 33- ee some se 45, 46, 55, 57, s9--61, 63, Tl, 72, 77, 80, 61, 88, 100 0 lis, gang, ake 15, uA -a83, is1, is8, 162, 163, "Te-i77, i81—i83, 193-195, 197, 20, 2 5, 23h, 236, 2k5, 2h7, 2uB, 250, 257, 262? 567, ae 280, 282, 263, 291, 292, sg. & 297. 196; Molden- ke, *alph. List Invalid Names suppl. as 22; 25, & 26. 19475 Ray~ » 357, sae 393--397, 3 4o2, 4Oh-- 10s, 438, 50, ngs, ibs, "be, 467, U7, 473, 480, 461, 491 19h, 505, 507, 508, 510, Sil, peasy 517, 518, 520-522, 527— 576, 579, 581, 583, 587, 688, 98, G01, 60, 605, 613, 615, 617 618, 621, 633, 63h, 6ll; & 6h (1918), é7, 66 tte ee 111, ; » 1129, 1135--1137, 1145, 1149, 1158" 1162, 1167, 1168, 1176, 1177, 1179--1181, 1187, 1188, 1192, 1197, 1209, 1210, 1213, 1217, 1218, 1220--12227 12257 1227; 1228, 1233, 1236, 1252, » & 1289. ish9; Moldenke, Known Geogr. Distri rib. Verbenac., [ed. 2-8 453 17; °18; 20; 21, ‘16h, &199- 1949; He N. & A. L. Moldenke, Anal. I fol. Mex. 20: Ik. 19493 R. T. Clausen, Co - 19h95 Univ. Agr. Exp. Sta. Mem. 291: Moldenke, Phytologia 3: 72, 131, & 1g (1949) and 3: 283 & 375. 1950; Moldenke, Revist. Sudan. Bot. 8: 165 & 166. 1950; gn in Deam, Yuncker, & Friesner, Proc. Ind. Acad. Sci. 59: 51 (1950 res hae 87. 1951; Moldenke in Chittenden , Roy. Hort. Soc. Dict. : 2209 & 2212. 1951; Moldenke, Phytologia 3: 450, h67, & N68 (1952) and h: ve .— rei 1952; Moldenke in Gleason, New Bri itton & Br. Illustr. Fl. te 1, 32 126, 129, 131, & 132. 19523 Mol- denke, Phytologia fs 135 & 186. 1953; Moldenke, Inform. Mold. Set 48 Spec. (4). “195h3 Moldenke, Phytologia 5: 133. 1955; bert? ss 182. 1955; Moldenke, Biol. Abs 1093. 1956; Lehr, oe — Rot. ree Bh: 312. 1957; Straus~ augh & Core, Fl. W 71h & 777+ 1958; Moldenke, Am. Mi b Nat. 59: 343, 3hh, ju, 9) & net 119585 Moldenke in Gleason, New Britton & Br. Illustr. Fl., print. 2, 3: "126, 129, 131, & 132. 1964, Moldenke, Monograph of Verbena 161 1958; Moldenke, Résumé 4—19, 21 26 a Bead nce, sc 368, 373, 37h, "46, & Li7he 19594 Jes Beh gor 1&2 :1& lL, 19 60; enke and 5: 1, 1962; J. er, Trans. Kans. Acad. Sci. 6 418. 1962; Moldenke, Résumé Suppl. 6: 1&2 (1963) and 7: 1& 2. 1963; Gleason & Cronquist, Man. Vasc. Pl. 579 & 580. 19633 G. N. A. ie Fl. Ill., ed. 3 [Am. Midl. Monog. 7:] 213. 19634 Steyerm., 963; Moldenke, logia » 51 » 151, 165, 206, 21h, 215, ‘sig—-201, 293, t 296 (1963), ¢ ”372°& 373 (196k), 10: 151-155, Vil, 285, & 30h 196 Ae ee 8h, 86, & 109. 196k; Ssumé’ inedt i 1 #2 {a96l) ane : 1-3 & 8. 196k; A. a “aataeki, Phytologia 3060 (1 ; Gaz. 69: 58, fig. 17, 5 fig. fig. 60, 67, fig. 69 & 70, 70, fig. 28—In, 3) & mn! & pl. 6, fig. 1. 1920; G. T. Stevens, Illustr. Flow, “ba, Middl. Atl. & New Eng. States pl. 129. 1930; Merriman, Fl. Richmond & Vic. pl. 36, figs 9. = pe Dermen, en ‘i "161~263, fig. Se le 19365 Moldenke in Gleason, New Brit- - lustr, Fl., print. i; 3: 3 (1982) ae print. 2, 3: 132. 1958; Strausbaugh & Core, Fl. W. Va. 3: 777- 1958; Steyern., Fl. Mo. pl. 202, fig. 2. 1963. paadal herb; roots fibrous; stems chiefly erect, 1—6.5 dm. tall, very s slender, tetragonal, simple or sparingly fastigiate- branched above, occasionally branched from the base, sparse Strigose-hispidulous; branches ascending, usually spars ely strig- illose; leaves decussate~op e~opposite or sometimes apparently whorled, Sessile or subsessile, linear or linear-lanceolate to narrow- eolate, lanceolate, narrowly oblong, or spatulate, sometimes oblanceolate, 2,5--10 cm, long, 6—~8 mm. wide, bright~green, ob- tuse or subacute at the apex, long-attenuate or tapering into the Subsessile base, varying fron entire to minutely sharp-serrate, Ss e ly scabrous, somewhat prominently veined aiaeath: glabrate or Sparsely strigillose on both surfaces, especially along the rais- 3 litary at the apex of the stems a anches or subternat strict, el rather densely many-flowered, strigose- Scabrid; bractlets lanceolate-subula » commonly shorter than the calyx or subequaling its tube, glabrous or nearly 80; ca mm. long, the teeth acute; corolla hypocrateriform, varying fran pale nepal sci nder, or lilac to purple, pale Purplish~blue, pale-blue, light-blue, bagi or light-violet Snoothish, its tube scarcely longer than the calyx, with scatter- ®d hairs at the mouth, the limb 5-6 mn, wide; fruiting-calyx angulate, --5 mm, long, sparsely pubescent, the lobes acuminate TOOREVEG in urpassing the fruit; fruits crowded, the cocci linear, 162 PREE0L 0624 Vol. ll, no. 3 2.5-—-3 mm. long, seivederqticylase above, striate toward the base; chromosome numbe # It is worth noting i that according to Kanda the haploid number of chromosomes is . Patermann (1935), however, says "An~ derseits ist es ny auch nicht ganz von der Hand zu weisen, dass die untersuchte Pflanze ein Bastard war, weil, wie auch schon Kanda angibt, htufig Zwischen-formen zwischen den einzelnen Ver- bena-Arten vorkommen, was um so mehr verst&ndlich ist, als alle der Gattung Verbena die Haploidzahl 6 besitzen." Walpers (185) classifies this species in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Sec~ ondary Subgroup Ho Holophyllae with 22 other species. The type of V, rugosa Muhl. appears to be Muhlenberg s.n. [Herb. Willdenow lig] i in the Willdenow Herbarium at Berlin. The name V. angustifolia L., is based on Herb. Vigener s.n., from material cultivated in the Botanical Garden at Heidelberg, wong: 4 while V. cuneifolia Wallberg is based on Wallberg s.n. from material cultivated in "Hort. Proprio", Sweden, in 1602, from seeds secured from Pennsylvania. It should be noted here that the ¥. angusti- folia of Lamarck is V. gracilescens (Cham.) Herter, while that of Miller is Stachytarpheta angustifolia (Mill.) Vahl; the V. cunei- folia of Hort. is V. hispida Ruiz & Pav., that of Rafinesque eis Ve st: Stricta Vent., while that of Rufz & Pavon is a valid species, which see; the v. stiegriielss of Sess6 & Mocino is also a valid species; and the V. rugosa of D. Don and of Sweet are V. rigida Spreng., while that of =o is V. stricta Vent. Willdenow's description (1809) of Verbena rugosa Muhl. is worth repeating here: "*VERBENA rugosa. V. “spicis filiformibus solitar- iis terminalibus alaribusve, foliis lanceolatis, basi attenuatis integerrimis, lineato-venosis. Verbena rugosa. Mtthlenberg. V- (angustifolia) humilis erecta, foliis lineari-lanceolatis rariter subserratis, awiaia filiformibus. Mich. amer. 2. p. 1s. Habitat in Pensyl Tennassée. \y. F.* Verbena simplex has been found by collectors in dry or open dry soil, sandy or open sandy soil, clay or dry loam soil, —_ or hard dry or or Dek open roadside 8s, c limestone glades, old fields, prairie fields, woods and 8 ods , ar , shallow soil on limestone ou limestone quarri terops, es and rocky limestone slopes, the talus of limestone ledges, open and roadside waste places, moist meadows, weedy roadsides, ditches, cinder beds of railroads, dry rocky places around stonepiles in hayfields, and bare: rocky regions in general. It has been collected on sandy and low sandy prairies, aneeat or sterile prairies, bluffs and river 1964 Moldenke, Monograph of Verbena 163 bluffs, steep dry or sandy banks, exposed cliffs, sand mounds, ballast, river flats and rocky bottoms, railroad tracks and em- a limestone slopes along creeks, limestone barrens, and gravel bars along streams, around old Limestone quarries , in’ open woods, at shes edges of Besse and the foot of lime stone rocks, and along railroads, , clay, or open roadsides, rivers, and waysides, at altitudes: of 7 he to 2000 feet, flowering and fruiting from May to tobe: Pammel & Fiske describe it as "common in sandy soil of creek bottoms associated with V. stricta, Potentilla c s, and Scutellaria parvula" and "common zamens sandy soil associated oe Lithospermum canescens, Padus nana, and Fraxinus lanceolata”. Pennsylvania it is said by Stone (1945) to be "rare in —_ roadsides", flowering in early June, while Benne r (1932) "dass vibie it as "rare and local" in "ary fields and waste ns " it "local in dry sandy fields" in amentinn Camp fo Carolina, while Cronquist calls it a eoudatde weed" in Georgia. ueky by Says of it in dry situations or in low places in fallow fields", while Braun (2943) reports it from "dry limestone soil of open slopes and roadsides". C.K. Dodge avers that it is "plentiful" in Saint Clair County, Michigan. In Tennessee McGre~- gor calls it "abundant in shallow soil over limestone". In Ark- ansas it was found on basic soil on limestone by Robinson, while Dunn found it in "cutover oak woodland with regrowth 15--20 feet In Illinois it inhabits "sandy humus underlaid by limestone", Tlelds. apie Anes bclaneie hybridize with V. ss hastata, and V. bracteata", but F. E. McDonald calls it "rare" in Peoria County, aise 90) mi that it grows in dry soil most- in — southern 2/3 of eastern t of rl. kophtte’. oviey ectianviee th alah wpase Coast TE Sn Seonences, along the edge of bluegrass pastures, roadsides, open Sandstone areas, rocky s in the prairie, sandstone outcrops, Places in swales in bluestem 29, 7228 ravines in tall hluesten airies, open places among grasses on rocky prairie hillslopes and in ravines, in loam of moist thickets, on rocky prairies Slopes in the dry bluestem association, mstelands, sandy soil of 164 PHYTOLOGIA Vol. 11, no. 3 pastures and rocky prairie pastures, and in open places in ra- vines in rocky bluestem prairies. Hulbert calls it "scarce" and "“yery scarce", but ray 6 describes it as "common" or "scatter ed"; Horr found it "scat In Iowa, according to ensS it inhabits "sandy, alluvial d ary rocky ground, infrequent". Bush reports re as "com mon in barrens" in Missouri and McGregor found it "scattered to common on rocky hillsides" there, while Palmer ares says "Prai- ries, fields, and glades. Calciphile to circumn al. eo SOEee ents of a Marion and Boone counties to Jackson Co." The Seis “1958 sate tion cited below bears a notation to the effect that Prof. Tuck- erman first seever ee bags species at that locality. Harvard vary names recorded for V. simplex are a wiles vervain", "bur~ vine", "narrow-leaf vervain", "nar row-Leaved verbena", "narrow leaved vervain", "narrow-leaved ae ara "pigmy vervain", "rugose ein vervain", "sand verbena", "verbena m, and "verv feuilles étroites". The plant is ete by the following fungi: Aecidium verbenicola Ell. & Kellerm., Dicaeoma verbenicola Arth., Erysiphe cichoracearum DC., Puccinia bite "villas Arth, & Holw., and Septoria verbenae Rob. a Pe ae Specimens of V. simplex have been misidentified and distributed in herbaria under the names V. snes tifolia x hastata Hill, V. caroliniana L., V. halei Small, V. hastata L., V. officinalis Bs; V. stricta Vent., | Stachytarpheta a: castifolis ’ II Vahl, and e- ven a eee Raf. in the A cieisthcenh On the other os Re Bell 420, — lia o rv. simplex phen? Bs ve. "nalet i small; W. H. Chase s.n. Feet aaTsTT) and W. He Horr 6723 are ve. hastata Le; Clevenger s.n. (Fletcher, Aug- 3, 1897] is in part V¥ V. hastata and in part V. urticifolia 1.5 Daniels sen. [July 1903], Egger Eggert son. [Pacific, 4 July 1896], We H. Horr san. (Aug. 12, 1929] & sen. [July 12, 1930], G. Merrill 571, an and Rickett son. [Yancy Mills, Aug. 13, 1927] am xV. m moechina Moldenke; A, Hayden 2 and eat Gleason 8845 (in part) are V- Sime plex var - eggerti i Moldenke; | Re L. McGregor 13217 is the type col= lection of V. simplex f. albiflora Moldenke; B. E. Smith 57 is Stylodon carneus carneus (Medic.) Moldenke; Roy Morrison s.n. n. is Buchnera americana L, in the Sebobinl asl aseay Paull son. [oswego, 5/29/ 1886] is Lobelia pubernla Michx. in the Lobeliaceae; and Jacobs © 26 is Salvia azurea subsp. pitcheri (Torr.) Epling in the Lamiaceae- 3 2. D. E. Brom s.n. [June 5, 1899], A. S. Hitchcock 5-M+ [Neo- ho Co., July 1896], and A.A. he s.n. (Monmouth, June ne 12, 1932], cited below, were all cited by Gates in his Kansas Flora 8° 196k Moldenke, Monograph of Verbena 165 , simplex x hastata", but I do not believe that they represent Ss hybrid. M. P. anit 3301 is a mixture with xV. moechina, The S. D. McKelvey 2523 2523, ci! cited below, is labeled "between Springfield and St. Louis"; M.W. Twaddell sen. [July 22, 1869] may be from Augusta or Rockbridge County, Virginia, since its la- bel is inscribed "between Staunton and Lexington", The specimen of Usteri 531 is inscribed "Brasilia", ar was actually collected at Tottenville, Staten Island, New Yor The W. He Rhoades s.n. [Sevie ele Pike near Knoxville], cited of leaves." Bohn 436, Eggert sn. [12 Aug. 1875], H. A. Gleason 88h5 (in part), Guttenberg son. [Sept. 10, '79], A. Hayden 2, Koe- nig sen. {July 1878], Ruth s.n, [Knoxville, June 1900], Shacklette 2 “and Sinclair, Dunn, & & Spellman 913, all cited below, are rather too large, nanyobyenched, mx ae plants and probably should be regarded as var. eggerti Moidenk The M. S. Bebb Son. specimen from aids, cited below, is marked "sp. ven." Schauer (187) cites Michaux s.n. from some- where in North America, Peter s.n. from Kentucky, Ba Beyrich s.n. from Tennessee, and Durand s.n. from "Carolina", in the Berlin 1 ahi Pennsylvania; C. on Jones (1941) cites Bartley & Pontius S.n. from Jackson County, Ohioz EB. L. Braun (1 va recards the species from Barren, Bell, Grayson, Hardin, Lo Meade, Spencer Todd, and Wayne Counties, erga nee a9k7} records it from Sussex County, Delaware rchester County, Maryland; Dean, Yuncker, & Friesner “Tighé, mm 9h8, 1951) ree ord it fram Han- > m Co’ . ge o Journ. Sci. F nana “County, Ohio, on the y J. H. Schaffner Be jaa ei (1963) cecoraa the species ‘tron Atchison, Bates, Ben~ Fae ee Cal allaway, Camden, Carroll, Cedar, Christian, cls, : 4 erford, Dade, Dallas, Daviess, Douglas, Gasconade, ry, W he basis’ of Rpachanee in (te herbarium of the Missouri Botanical Gar den at St * Louis, The species mae per ently first introduced into cultivation in 1302 in Sweden, 181 in s MacDougal (1907) suggests that V. lex may hybridize in the wild with V. bracteata. However, he cites no examples, gives no description, and does not actually say that he ever saw Bia Ke such hybrid plants. I doubt very much whether the hybrid oc al- though G. N. Jones (1963) also claims that such a gi occurs in 166 PHYTOLOGIA Vol. 11, no. 3 Illinois. Van Vleet's surname is misspelled "Van Fleet*® on some labels in the herbarium of the a ea Pe of Oklahoma Perry ( 1933) 6 cites the fe sags | 129 additional specimens and 1 photograph not as yet seen by me: ONTARIO: County undetermined: J. Macoun 1305 [Belleville] (a). VERMONT: County undetermined: ¥. H, Blanchard 26 (G), 60 (G). MASSACHUSETTS: Berkshire Co.: Hoffman s.n. (Sheffield, 27 Aug. 1902] (G). CONNECTICUT: New Ha-~ ven Co.: D. C. Eaton s.n, [New Haven] (G); Harger s.n. [Fair Ha- ven, 18 June 1886] (G). NEW YORK: Dutchess Co.: Hoysradt s.n. [Pine Plains, 1875] (G). County undetermined: Burnham s.n. [Trip- oli, 25 July 1920) (G). NEW — Bergen Co.: K. Ke Mackenzie 4213 (E). Camden Co.: Brinton s.n. (Sicklerville, 21 June 159h] (G); Parker sen. [22 June = a) one Redfield 6430 (E). Union Co.: Trelease 8. s.n. (Scotch Plains, 13 Aug. ; Aug. 1877) (BE). PENNSYLVAN- TA: Chester Co.: Bartram 1021 (a) « Franklin Co.: Green 3.n. [Mercersburg] (G). Philadelphia Co.: Nuttall s.n. Philadelphia] (G)- York Coe: MacElwee 760 (E). County nty undetermined: Muhlen- berg s.n. (E—photo). MARYLAND: Dorchester Co.: C. P. Smith mith 3186 (G). Montgomery Co.s H. De House 1363 (E). DISTRICT OF COLUMBIA: Blanchard s.n. [Joy City, “Ig June 1891] (E); Sucre Bone Laas ington, 26 May 1889] (G). VIRGINIA: Bedford Co.: Curtiss (ie 1873] (E). Botetourt Co.: A. B. Seymour 8 (G), (6), 9 ( (OF ee Cost J. K. Small s.n. [about Cumberland Gap, 27 July 1892] (e F, G). Loudoun Cos: H. D, House 875 (W). Smyth Co.: Je Ks Small s.n, (Chilhowie, Aug. 1892] (F)- WEST VIRGINIA: Green- brier Co.: Grimes 3716 (G). Lewis Co.: Small & Heller 32 (G)- Monroe Co,.: Steele & Steele 3k (EB, G). “NORTH CAROLINA: Buncombe Co.: Biltmore Herb. 4759b (G). SOUTH CAROLINA: Abbeville Co-: Hexamer & Maier s.n. [Abbeville District, July ony (G). GEORG IA: Catoosa Co.: J. | J. K. Small s.n. [near Ringgol » O--12 Aug. 1895] (F). FLORIDA: County undetermined: Rugel 127 rrken 49} (E, F)- MISSISSIPPI: Oktibbeha Co.: C. L. Pollard 1312 (E, F, G). OHIO: Butler Co.: Overholts s.n. (Oxford, 16 June 1910] (E). Erie Co? Moseley s.n. [Castalia, 27 July 189k] (G), s.n. (Sandusky, 26 July 1894] (F). ILLINOIS: Cook Co.: Greenman 1981 (E); H. H. Smith 5670 (G). Gallatin co.: E. J. Palmer 15262 (E). Pulaski Co.: Ge R. Vasey s.n. [Mound City, June 1959] (G). Randolph Co.: L. He Pammel s.n. [Red Bud, 3 June 1888] (£). — Island Co.: Bngel- mam $.n. [Rock Island, 17 Aug. 1866] (E). Saint Clair Co.: Eg~ gert sn, sn. [bluffs near Prairie du Pont, 20 i 1876] (EB); Norton s.n. [East St. Louis, 9 July 1898] (E). Stomy Island: Greeman 30 (8); He H. Smith 5923 (E, G). INDIANA: Harrison Co.! CG» C+ Deam 20357 (G, W). Saint Joseph Co.: Nieuwland 2685 (E)- Tipton Cows Se Me peas Sen. (C. C. Deam 13619] (E). IOWA: Black Hawk Coe! 196k, Moldenke, Monograph of Verbena 167 L. H. Pammel 272 (G). Johnson Co,: A. S. Hitchcock s.n, [Iowa City] (E). Louisa Co.: L. H. Pammel 1699 (E). County undeter~ mined: Arthur a [17 July 1875] (E). KENTUCKY: Henry Co.: Eggles- Farland 109 “WE Lyon Co.: Eggleston 5237 (E). Simpson Co.: An~ derson | & Woodson 47 (£). Warren ren Co.: Price sen, [Bowling Green, 21 May 1899) (E). TENNESSEE: Blount Co.: Curtiss 1955 (E, F, G). Carroll Co.: Biltmore Herb. 759 (E, G). Cheatham Co.: Eggert s. n. [Kingston Spring, 5 Aug. 1897] (E). Davidson Co.: Eggleston _ 1,30 (E). Hamilton Co,.: Engelmann sn. (Chattanooga, 25 Aug. 1876] (E). Knox Co,: Ruth 765 (E), sen. [Knoxville] (F). WISCON- SIN: Sauk ~ Hale s.n. n. [Baraboo, 1861] (E, G). KANSAS: Ander- Son Cos: A. S. Hitchcock 791 (E, G). Linn Co.: Kellerman Son. (Mound oie S July 18, 1887] (); Rydberg & Imler 120 (BE). MISSOU- * Barry Co.: B. F. Bush 206 (E). Franklin Co.: Greenman 4125 (E); Kellogg 1130 1130 (B)} L. L. He H. Pammel s.n. (Washington, 25 June ne 1888] (E); Trelease 713 a3 8); “Greene Co.: Dewart 42 (B). Jackson Co.: B, F. Bush U7 ( (E, G)3 Tindall s.n. [Independence, 1 June 1895] (E). Jasper Cou: E. J. Palmer 200 (E), 2981 (B), 3319 (E). Jef- ferson Co.: Eggert 5 Sen. [20 June 1876] (E); | A. S. Hitchcock s.n. (Victoria, 8 July 1390] (E); Riehl 456 (E); Steyermark 292 (E);_ Trelease s.n. (Crystal City, 1 May 16987] (E); Wislizenus 280 (E). Lincoln Co.: J. Davis 552 (E). Macon Co.: B. F. Bush F. Bush 7586a (er; Marion Co.: J. Davis 3588 (E). McDonald Co.: E. J. Palmer 232 (E). Ozark Co.: E. J. Palmer 34766 (E). Phelps Co.: Kellogg 496 (E). Sainte Genevieve Co.: Greenman 4578 (E). Saint Louis Co.: Ti 3% (E), sen. [banks of Mississippi above St. Louis, Oct, j King s.n. [mineral region southwest of St. Louis, June Sun. Sana Station, Eureka Springs] (n). OKLAHOMA: McCur Co. He W. Houghton 3622 (E, G), 36 (E, G). Osage Co.: Ge ¥. Stevens 1980 (G). Ottawa Co.: G. W. Stevens 2299 (G). Rogers Cos: B. Fe Bush 433 (BE). Washington Co.: Ge G. W. S Stevens 2080 (G). The "Steele 314" which she cites is actually Steele & Steele 31h; her " Sabeogh is a misspelling of Hoysradt; the Curtiss 1955, P Pollard 1312, and Small s.n. (Chilhowie, Aug. 1892], which she cites frou "NY", are actually in the Columbia University arena while Short s.n. is in the Meisner Herbarium. She says "This spe- cles appears to | to be very closely related to V. ha pines In its typical form, it is readily distinguished by its elongate spikes and narrow leaves, Often, however, intermediate forms occur be- 168 PHYTOLOGIA Vol. 11, no. 3 tween the two, making it somewhat difficult to find clear lines of demarcation between then," In all, 973 herbarium specimens, including the types of at least some of the names involved, and one mounted clipping have been examin y me, Citations: QUEBEC: Jacques Cartier Co.: Holmes s.n. [Montreal] (T); Racine s.n, [Montreal, 17 juin 193)] (Um--255); Rouleau 1% (Um—-253). County undetermined: Dumas s.n, (Carré Chaptal] (Vi). Sainte Helene Island: E. Roy 2789 (N, Um—252, Vi, Vi), 3681 (Vi, W—1948911). Saint Paul Island: Rouleau 1125 (Vi). ONTARIO: Carleton Co.: Soper, Gillett, Calder, & Vick 3262 (Ca--882812, S). Hastings Co.: J. Max Macoun s.n. [July 10, ~1871] (Mm--950). Victoria Co,: Bassett & - Bragg 147) (Vi). Counts undetermined: J. Macoun 54634 (N), s.n. [near Belleville] (Bm). NEW HAMPSHIRE: Grafton Cost Josue sn. (25 June 1887] (Dt), sen. [June 189k] (Dt). VER- MONT: Bennington Co.: W. H. Blanchard 1 [1 (vt), 161 (N), son. [Ar- lington, July 1903] (vt). 1 Windsor Co.: Kittredge s.n. TWood- stock, July 2h, 1918] (Vt). MASSACHUSETTS: Berkshire Co.: E. H. Eames 10365 (Dm). Hampshire Co.: Jesup 1955 (Dt). Middlesex Co.: corals sen. (June 1894] (Dt). CONNECTICUT: Hartford Co.: T. 8.n. nN. [July 8, 1895] (We). Litchfield Co.: Eames Godfrey Frey 32h (ii (i), s 8.n. [Kent, June 24, 1905] (F1~—21059). New aid Coe: n Brandegee s.n. [New Haven] (Ca~-l68126); W. We Denslow s.n. [New Haven, Aug. 1, 1866] (Ms); E. H. Eames rp ry [VIII 14.1893] (vt); D. C. Eaton 8.n. [New Haven, 1053] (S), 8-Ne New Haven, 1859] (C); Harger 5032 (Vi); Kleeberger s.n. [New Ha~ ven, 6-22-187))] (Gg--31267); Safford 216 (W--515216); W. A. Set- chell Sn. (Westville, Feb. 18 885) (Ca--25177); W. W. Smith ho (We); J. W We Wood s.n. [New Haven, June 186] (Mis) « ~~ County unde- termined: W Wright s.n. [Connecticut] (Go). NEW YORK: Bronx Co. E. P. Bicknell 7337 (N), 7338 (N); Leggett son. [July 234 1871] (Te). Cattara taraugus Co.: Fenno s.n. [July 30, 1903] (Al). Colum- & Co.: McVaugh 2239 (Al, Up). Delaware Co.: P. Wilson s.n. {arkville, July 9, 1915] (N). Dutchess Co.:; Disbrow $.n. [Pine es Plains, Oct. 1875] (Tc). eavbe Co.: Boughton 5762 (Al). Nassau Co.s Ee P. Bicknell s.n. (Aug. 15, '03] (N). New York Co.: W. We Denslow a [July 1, 1867] (Ms). Queens Co.: Hexamer & Maier sn. . [duly 3, 31, 153) (cn); S. 0. Rusby s.n,. [Woodhaven, July 1878] (Qi). Rockland Co.: He | Ne] Moldenke 7545 (N); Whitney 5374 (Al). Sara toga Co.: Cotton s.n. [June 28, 1905] (Al); H. D. D. House rok [June 30] (a1), 10M, (Sept. 18] (Al). Ulster Co.: H. D. House 25779 (Al). Washington Co.: S. H. Burnham s.n. [June e 28, 191k] @)- Westchester Co.: P. Wilson s.n. [Sept. ie 1915} (N). — Is- land [Kings Co.]: Usteri 531 (B). County undetermined: W. W. 196) Moldenke, Monograph of Verbena 169 Denslow s.n. (New York] (Go); J. Torrey Tu9 (Al), sen. (New York] (a, c, C, Mi, Mi, Pa); Torrey & Gray | B.n. (N. Y.] (C). MEW JERSEY: Bergen Co.: Brinton s.n, [closter] (1 (Up-—~17073); Churchill s.n. (Englewood, July 1883) (1 (Ba); L. Johnson s.n, (Hackensack Swamp, Sept. 6, Pgh (Lh); K. Ke Mackenzie 1213 (N); Niederer s.n. (Carlstadt, July 26, 1883] (N); J. W. Wood s.n, (Lodi, July 1850] (Ms). ade Co.: Ge C. Stonart s.n. n. [Edmondson 2,28] (N). Glou- cester Co.: E. B. Bartram s.n. [. sn. [June 23, 1907] (Um--60); I. Burk s.n. [Red Loa Cup 707%). Hunterdon Co.: K. K. Mackenzie 5928 (N). Mercer co.: Slack s.n. [Trenton] (Up--51869). Middlesex Co.: Monachino s.n., (Cheesequake State Park, July kh, 1950] (N). Monmouth Co.: N. L. Britton s.n. [Seabright, June 1877] (Tc); 0. R, Willis s.n. [Freehold] (Mi). Morris Co.: F. C. Baldwin 81-1- 2 (Nm); K. K. Mackenzie 7850 (N); E. Wall s.n. eae, 19/6/ 31] (Bw, Ew). Passaic Co.: Hills Sone . (Paterson, June 2h, 189)) (Up--17073) ; Knieskern s.n, (Br); Schneck s.n. [In a field be- tween New York and Phi. Philadelphia, June ane 6, - 1902) (Ur); Seaman s.n. ig PENNSYLVANIA: Bedford Co.: Berkheimer 2675 (Ca~-805800) ; Jennings s.n. [6/26/41] (Cm, Cm). Berks Co.: Wilkens 5191 eee Up). Centre Co.: H. A. Wahl 1316 (Ca—882813, No— 8618, P1~-129972, Ur), 177k (Hi—168733). Dauphin Co.: J. K. Small. 106 106 (0b—-50753), sen. [August 13, 1888] (Vt). Delaware Co.: Smith & Smith s.n. [Rhodes Swamp, June 10, 1865] pp Othe Erie Co.: | Guttenberg s.n. [Sept. 10, '79] (Cm). Franklin C True 27 (Up). Fulton Co.: Bright s.n. (Sept. 2nd, 1918] toa). san. [6-1-19] (Cm). Lancaster Co.: Disbrow s.n, (Nm); L. J. Mil~ ler s.n. (N. Lanc, Co.) (Vt); T. C. Porter sn. [Aug. 26, 1862] (ca)5 J. x. Small s.n, [vicinity of Conewago, May ze 1889) (Ww 298656). “Lebanon Co-: Arndt s.n. [Adamsdale, Aug. 22, 1916] (Up). Mifflin co.: G. B. Grant 1434 [3060] tro 7632). Mont- gomery Co.: Mumbauer s.n. [July 1, 1907] (St--22309). Northampton Co.: Tyler s.n, [Easton, 1892] (Dt). Perry Co.: Adams & Adams 26 Tom, Up); Gress, Jennings, & Jennings s.n. [July 25, 1920] (Cm). Philadelphia Co.: Martindale s.n. (Philad., July 1876] (Du-=21,173, Je—~3149); C. F. Parker sen. [June 1876] (C), sn. (July 6, 178] (Pr). Schuylkill Co.: P. “Pe Re Wagner 7049 (Up). Snyder Goi Wade & Wade 1726 (Up). York Co.: MacElwee 760 (N, Une=13) 5 We Stone Bag (Up). — undetermined: Galen s.n. (Ur). DETAWARE: New Castle Co : Brinton sen. [Holly Oak, Aug. 19, 170 PHYTOLOGIA Vol. 11, no. 3 1888] (Up--17072). MARYIAND: Frederick Co.: W. (W-- 1927873). Kent Co.: Brinton s.n. [6/16/87] a A "eaten ery Cos: F. J. Hermann 9533 (Mi); C. S. King 2220 (La); Leonard & cre 388 688 (Ur); Pieters s.n. [Cabin John, May y 31, 1896] “(it), | 8. » [Chevy | Chase, July 1, 1897] (Mi). Prince Georges Co.: T. Holm Son. (Junii 5, 1900] (I). Talbot Co.: E. C. Earle 150k (Up), 1640 (Up), 2773 (Up); Jump s.n. [June 16, 1937] (Up). Baltimore: We R Re Jones s.n sn. (Baltimore, June 22, 190h)] (Mn--6887). DISTRICT OF COLUMBIA: F. “F, Blanchard sen. (19 June 1891] (N, Or--14720); E- S. Burgess s.n. (Ob—5075i); Camby s.n. [D. C., 1881] apt T. Holm s.n, [Junii 17, 1893) (1, S$); Pieters sen. [32nd Streets, July 30, 1899] (Mi); Rugel 196 (Br); c. 5 Se pie ae 5M. (June 1, 1881] (Al, 0b—~9 7280); E. ; E. S. > Steele ele s.n, (Washington, 1874] (Po--70657), son. [June 27, 7, 1896] (Ob—-50761); L. F. Ward 8.n. (Washington, 1888] (Ka), sen. (N). VIRGINIA: Albemarle Co.: F. Blanchard s.n. [Ivy City, June jane 19, '91] (Dt). Augusta Co.: Murrill s.n. (Ataunton, 23 May 1895] (Cm); M. W. Twaddell s.me . [between Staunton and Lexington, July 22, 1869] (Up). Bedford Cos: Curtiss 6715 (Mi), sen. [June 6, 1871] (Al, Mi), sen. (June 19, 1871] (S), 3. Sone [June 20, 1872] (Pu), Sone (Mi) « Botetourt Co: A. Be Seymour lo (Gg—~—358661, Lb=--16172 138 Campbell Co.: Britton, “Britton, | & Vail s.n. [July 1, 1892] (C); Murrill 5.ne [TLynchburg, 31 lay - 1927) (Fl--25520). Fairfax Co.: Se ee Blake 8473 (I); J. J. Carter s.n. (Falls Church, June 1876] (T1)5 Te Holm s.n. (Septbr. 3, 1898] (1); Kinsley s.n. [June 26, 1888] (Ob--50757); W. R. Taylor 2610 (Up—-75577) « gua! Co. Al- Up—17071, W). Loudoun Co.: J. H. Holmes s.n. (Aug. 1888] (Lb— 27617, Ua—11396), sen. (Ur—2290h); H. D. House s.n. [Bluemont, May 28, 1905] (W--92807). Mecklenburg Co.: F. R. Fosberg 1500 (Up). ‘Middlesex Co.: Leonard & Killip 53h (W——1101588) « vont- gomery Co.: C. H. Hitchcock s.n. [Blacksburg] (Dt). Page C+! \ Britton & Britton s.n. (Luray, Aug. 31, '85] (C); W. H. Camp 12 (MN). Roanoke Co.: C. E. Wood 5850 (Hi—51180). Rockbridge Co-% Brom, Hogg, Vail, Ti , Britton, & Britton s.n. [Na ms Bridge, June 6, 1890] (V)3 F. H. Sargent 6908 (Ge—393050). Roe ingham Co.: He H. Bailey s.n. Lecorierha rien: 1, 1918] seat id wi ieuas Smyth Co.: Britton, Britton, & Vail s.n- (gune 23, 1892] (C, Cm); J. K. Small s.n. [Chilhowie, Aug. i, 1692] =a ee curators who have material of this species from additional counties are asked to send it to the author f ificati Figure 16. Distribution of Verbena simplex in the United States record, so that future editions of this map i bigennnsypteriaty: nnn NY come See ty Anteet a inne SUR 7 ——— tm, ) vg Fe cy Shiga at. > sueear Bien fa coos ryt i | 4 ‘ 4 rl a Bis 14 i , = - mill i i a “ie oa ‘es in Pag | a wf oon on 7 fe 3 ' a 2 ' f rE \ Y? ‘ = ee ins, — = mENTISS| SM coset PRAIRIE E |ramaney = 9 SA (ve : KEYED TO COLUMN A of Cleartype index Book CLEARTYPE Ww Lt i Sten , “jee. eo Be TRADE MARK REG. U.S. PAT. OFF. HER Ss vl ae ies ) COUNTY OUTLINE PANOLA Tew Me oe Cosa a ela pat grec a cepa ase UNITED STATES 4 ? ane ¥ v ae . M xan wae b= ann Youn" Tomas [wroons rd gog a eal boy ay VERNOR NEWTONS ® = Scale of Miles 5 100 200 a SCHLEICHER 0 MAP NO. 125 AMERICAN ORIGINATORS ~Yoe CLEARTYPE MAPS Rue meenremee Copyright, American Map Co.,, Inc., New York, No. 14324 I a ST ee rT T z a 3 I 7] | a ae: See I Hi I FI I 2 i | EE = 3 u T u T ry 196, Moldenke, Monograph of Verbena 171 Washington Co.: W. H. Camp 1633 (N). County undetermined: R. C. Alexander s.n, [Valley of Virginia, 1850] (Ca~=-379993) ; Cc. Si Sheldon s.n. [Chain-bridge, 1 June 1881] (0b-~50752). Roanoke: Brown, Hoge, V. Vail, Timmerman, Britton, & Britton s.n. [May 29] (Bm); Small & Talker 132 (C). Wildl susie: GF Grimes 3738 (N). WEST VIRGINIA: Berkeley Co.: E. L. Core 5883 (We); 3 da he L. Sheldon 266 (We, We). Grant Co.: E. L. Core s.n. [Petersburg, July 13, 1935] (We); Davis & Davis 768 (We); West Va. Univ. Bot. Exped. 271 (We). Hanpshire Co.: Frye 15a (We), 1219 (We), s. sen. [June 20, 1933] (N); Koenig s.n, . [July 1878] (Cm). Jefferson Co.: Da~ vis & Davis 2908 (We); C.. C. F. Millspaugh 865 (C, We); H. N. Mol- denke 21764 (Bn); Puissant 5 Sens [Harper's Ferry] (Br); Straus— baugh 271 (We, We). Kanawha Co.: E. T. Harper 5687 (Cm). Miner- al Co.: Strausbaugh s.n. [June 16, 1933) (We). Monroe Co.: Steele & Steele 31, (N, W--1,90291). Morgan Co.: Burton s.n. (Shady Grove Park, | July 6, 1937] (Gg—267620, sd—20296); Mf Maysil- les s.n. [August 30, 192} (We). Pendleton Co.: E. L. Core 2 3676 (0b—50755), S.n. [H. N. Moldenke 6822] (N). County undetermin- ed: H. M. Gamble | s.n. (We). NORTH CAROLINA: Buncombe Co.: Beadle S.n. [Biltmore, - VII.1911] (La); Biltmore Herb. 4759b (N, W— 332101) . Burke Co.: Rugel s.n. (Morganton, Ju June 17th, 1872] (Ww 310736). Chatham Co.: ? Kessler ler 573 (Hi--137)59). Durham Co.: Blomquist 4710 (H—-20h)0), s.n S.n. ~ (h/30/38] (Fl--12344); Fox, God- frey, & Anderson 2h (No—~23139) . Forsyth Co.: P. 0. Schallert 1539 (H-—5L53). Gaston Co.: Columba 325 (I); P. I . Schmitt s.n. TJune 1881] (I). Granville Co.: Ahles & & Leisner Thuoe i 92929); Buell 1383 (No--8593); W. B. "Be Fox 910 (N). Jackson Co.: Cc. W. Galloway 2206 (Hi—-5060). Madison Co.: J. De Smith s.n. {Wam Springs, July 27, 1880] (W-13230)2). Orange Co.: Ashe s- n. (near Chapel Hill] (W-~-32781:8) 3 W. C. Coker son. [May i, 1910) (Hi--59456); Collector undesignated 8.n. sen. [May 20, '91] (Hi—59U52)5 A. Holland s.n. [May 21, 1929] 7 (Hi--S9L53) 5 He Re Totten s.n. [June 15, 1915] (i $9455)3 Warnock sen. [8/7/23] Metis Polk Co.: E. C. Townsend 91 (P1--07178), son. [Col~ , June 28, 1897] (WoTSL76E) » Bockdighen Cos: ot Son. (spray] (W— 367383) . Rowan Co.: Coit 1254 (No--3599)3 W. M. Davis S-n. [May 1882] (Up--17073). SOUTH CAROLINA: York Co.: 2 Ab les & Haesloop 27167 (Hi—~10399). GEORGIA: Bartow Co.: W. H. Dun can 8,27 (Gu—29950) . Catoosa Co.: J. K. Small s.n. (near | Ring- gold, Aug. 6—~12, 1895] (C, Io—20749). Cherokee Co.: W. H. Duncan 8349 (Gu--29977). Clayton Co.: B. B. Higgins s.n. [Lovejoy, 7-1- 29] (Ga). Dade Co.: W. H. Duncan in 2600 (N)- Floyd Co.: H. C. Jones meee 155 (Gu--19913). Fulton Co.: J. H. Barnhart 2253 [Herb. Barnhart 172 PHYTOLOGIA Vol. 11, no. 3 2697] (N). Richmond Co.: Cuthbert s.n. (Augusta, Aug. 7, 1900] (Fl—-21057). Meriwether Co.: Wetherill s.n. [Warm Springs, Aug. 1882] (Up). Walker Co.: Cronquist 5280 (Gu—-30233, N, Tl, W#— 1928668), 5299 (Ca-~777L67); P. Wilson 161 (N, W——384816). Whit- field Co.: R. M. Harper 385 (N, W—36)817). FLORIDA: County un- determined: F. W. Chapman sen. (C); Rugel 127 [18h2--9] (W-- 511956). ALABAMA: Blount Co.: C. F. Baker s.n, [Blount Springs, 5/5/98] (Po—63885), s.n. [Blountsville, 5/6/98] (cm, Ur, Vt, W-- 344791). Choctaw Co.: Schuchert s sen. [Cocoa, ene 13--15, 1896] (W-~288831). Cullman Co.: C. T. Mohr s.n. (Cullman, May 29, bet (W--771847); M. Morgan 69 GE pekaiy Bot ra Moldenke & oods 507 (S). Etowah Co.: Moldenke & Woods 508 (S). Jackson ae Earle sn. [Scottsboro , 6/30/99) (N)3 A Ae Re Moldenke 773 (S)- Mobile Co.: Dukes s.n. [Mobile, June 1905] (Rf). Montgomery TY COs? Farr s.n. (Montgomery, May 1h, 1911] (Up—52678). Wilcox Co.: Buckley sen, [July 180] (T, W—771847). County undetermined: M. Martens s s.n. (Br); G. R. Vasey s.n. (Alabama, 1878] (W); Winchell 138 (W). MISSISSIPPI: Oktibbeha co.: C. L. Pollard 1312 (C, Po— 70659, W—27168h). Panola Co.: Herb. Univ. Til. sen. [Long Creek] (Ur). OHIO: Adams Co.: E. L. Braun s.n. [June 23, '26] (Cn); D. M. Brown s.n. [July 27, 1910] (N). Butler Co.: J. Fer gusen sen. (July 9, 1932] (Ob—80588); L. Roth sen. [6/9/3k] | (or— soto, Obes Or—l3760, Or—-l5393); Wehmeyer & Waters 2 (Mi). Champai : E. C. Leonard 1610 [Herb. Leonard Ra) EXP I 2 19069 cra) Erie Cos: 0. E. Jennings sen. [July 8, 1911] (Cm); Mosel, . (Sandusky, July 28, 189)] @e-kn257); ep H, Panmel 67 (fo=-51037); T. Schoepfle s.n. [Sand Hill, May 31, 1896] ) Ot): York s.n. (Cedar Point, 10 July 1902] (Up). Hamilton Co.: Aiken 10673 (Ll), son. [aug. 20, 190] (Po—~26763k)3 E. L. Braun S-n- sen. [8/ 15/05] (Cn, Cn), s.n. [6/18/09] (Cn), sen. {aug. 9, '17] 17] (¢n)> Hig Co.s Bartley & Pontius 287 (N). Lorain Co.: Rudolph 5-2 5 -De (6-8-96] (Ob—-75327). Ottawa Co.: al Ss Britton s.n. [Lakeside, May 26, 1896] (F1-~21060); M. E. Day son. [July 30, 1893] (0b— 99795) Redick s.n. [July 15, 1890] OT (380589) 5 B. H. Thorne 8-N- (June 23, 1897] (db—-80590); York s.n. [Lake Side, 8-1-1902] (Dp, Up). ILLINOIS: Boone Co.: E. W. Fell 52652 (1138878); Fell & Fell F.46390 (I1—21217). Champaign Co.: Winterringer 569 9 (ur). Christian Go Co.: R. G. Mills s.n. [June 25, a (Ur). Cook Co.! 70) 1683 (Ok), 1684 (Ok), 1685 at oe SoD. 1926] (Bt—2]67, Bt—2h68); K. A. Chase sci Tir), 1620 (tr, Ur, W—Lh3181), son. (Aug. 5, 1899] (Ur); G. D. Fuller 1220 (Il— ast 1601s); Fuller & Link 348 (I1—16012); E E. Hall son. (W); Ee de Bee 196), Moldenke, Monograph of Verbena 173 152.1902 (Ur); F. W. Johnson 1753 (Go, N)s Lansing 2k2 (Ur), 1393 (Ur, Ur); Munroe $.n. [Riverside] (Mi); L. T. Nelson s.n. (Chica- go, June 1898] (N); Warne sen. [July 1870) (Al). DuPage Co.: Mun= roe s.n. [Hinsdale, Aug. 1876) (Po--70662); P. B. Whitford tee (1133828). Hardin Co.: W interringer s.n. [May 29, 19L9) (Ur). Jackson Co.: Fuller & Welch 918 (I1--1587) ; McCree 918 (Ur); A. B. Seymour s.n. {Aug. 19, 1800] (Ur—80010). Jo Daviess C aH anes 4399 (Ur); Vv. H. Chase 671 (Ur); Pepoon s.n. [Rush Tp., ~ 8h) (Ur); Winterringer 8039 (Il~—3793). Johnson Co.: Schnec 7 Sen. ee May 16, 1899] (Ur); Winterringer 317) (I1--28)07). - Kane > Boyce s.n. (Aurora, 1878] (vt). Kan (Vt). Kankakee Co.: C. C. Cramp~ ‘ad 363 (W—750626) ; E. H. Hill 72.1871 (Ur). Kendall Co.: Arth- ur s.n. [Oswego o, Aug. 2h, 187h] (Ah). LaSalle Co.: V. H. Chase 06s (ur) 3 Ge D. Fuller 2300 (I1—16011); McDougal I; (Ur); Re G. Mills sen. [duly i 19LL) ] (Ur); Raymound s.n. (Ur—-17h)57). Lee Co.: Keithley 460 (1123257) . Macoupin Co.: W. E. Andrews s.n. [June 21, 1889] (Ur). Marshall Co.: V. H. Chase 10660 (Ur, Ur). McHenry $353 Benke 5686 (N); G. Vasey s.n. [Ringwood] (C). McLean Cos: L. M. Underwood s.n. [Normal 1, July 17, 1880] (Cm). Ogle Co.: Ahles 1537 (Ur); W. S. Moffatt 480 (Ur). Peoria Co.: F. E. Mc- Donald s.n. (Peoria, Aug. 1096] (Ur); W. S. Moffatt s.n. (Romeo, Aug. Aug. 2h, 16 1895] (Ur). Pope Co.: Ahles 2782 (Aa). Pulaski Co.: Herb. Univ. Ill. 30011 (Ur); Pearsall 19 19 (I1~-22257). Putnam Co.: ve He H. Chase 10664 (Ur) (Ur). Randolph Co.: Winterringer 67. sci 34281). Saint Clair Co.: Brendel sen. [1850] (Ur); Eggert s.n. [across from St. Louis, 13.6.753 Herb. Geete 5697] (Go); J. “Fei 319 (I1—~23398); Welsch s.n. (Ur). Sangamon Co.: G. D. Fuller 2275 (I1—16008) ; “A. Be Seymour sen. (Springfield Junction, on, 6 Ju- ly 1879] (H—106l63). Will Co.: Boyce s.n. [Joliet, June 1h, 1881] (Io-~92229); Umbach 1702 (8), ), 5120 (Ca-bh1522, Ka), sane (Romeo, Aug. 25, 1897] (Ur), sn. [Rameo, June 18, 1898] (Hp, W—- 339476) . Winnebago Co.: M. Se 2 s.n. (Fount taindale} (Dt, N); R. Bebb s.n. (Rockford, Spring 1579] (Ok); E. W. Fell 51308 (I1— 38605); Fell e Fell F.i8-340 (Ur); G. D. Fuller 13746 (T1—~31276); Fuller & Haine 2h7f (11--16013), 220sh (I1--16009); L. H. Pammel 291 (Se--70052); Pammel & Fisk 291 (To--126997); We H. Rhoades s- af [Rockford] (Hs), s.n. [near Rockford] (Hs). County undetermin- ed: M.S. Bebb s.n. [Illinois] (Bm); R. Burgess s.n. [June 1883] (1092208); Regart s.n. [Bluffs in Illinois, 13 June '75] (Cm, Sg—-31271, Po--125373), sen. [Bluffs in Illinois, 20 June '76] (Cm, N, er tae Mead s.n. (C); G. R. Vasey 1 (I1--1600k). Sto- ny Is » H. Smith 5923 23 (Ca——982565)~ IND: INDIANA: Carroll Co.: Friesner 3056 (Bt—28517). Cass Co.: C. GC. Deam 17878 (Dm), 48838 17h PHYTOLOGIA Vol. 11, no. 3 (Bt—-28887), sen. (Logansport, 9/12/30] (Bt--5209, Crawford Co.: C. C. Deam 1648) (Pu), 20383 (Dm); S eis SNe [July 4, 1893] (Ur). Daviess Co.: H. J. Clements s.n. [7716/95] (Pu); C. C. Deam 25585 (Dm). DeKalb Co.: C. C. Deam 323) (Dm We). Harrison Co.: C. C. Deam 20357a (Dm), 50323 (N)} Fr Friesner 5669 (Bt--1h1h9, Ca-—Sh0h31, Ew, Lb—1354h), 14366 (Bt--50968, Gg--310873, Gu--21559, H--68383, Ky N). Jackson Co.: C. C. Deam 13519 (Dm). Jasper Co.: Welch ios. (Dp) « Jefferson Co.: Ce R. Barnes s.n. [Hanover, 6-23-76] (Pu); C. C. Deam 16195 (Dm) 5 A. He Young s.n. [Hanover, 1875] (Pr), s.n. (Hanover, May 12, 1880] (Pu), son. [Hanover] (Gg--31269). Kosciusko Co.: Friesner 15429 (Bt--51972). Lake Co.: F.C, Gates 1127 (ui); F. W. Johnson mn 389 (N, N); E. D. McDonald s.n. [Hammond, June 12, 1942] (W—1888L23) 5 Umbach 18h1 (ca—hl152 523). Lawrence Co.: Kriebel sen. [near Geor- gia, 6-l-33] (Bt--19474). Marion Co.: C. C. Deam 6953 (Dm). Ma shall Co.: C, C. Deam 31862 (Dm). Monroe Co.: V. Davis 5.n- [Bloomington] (In—2286); Gl Gheen s.n. [Bloomington] (In—-2325)} Gullion s.n. (Ellettsville] (In—-l225); Wynn 63 (Au, Gu--27366) Newton Co.: M. McKee 866 (Dp). Porter Co.: He | R. Bennett s.ne [August 18, 1957] (Go). Posey Co.: C. C. Deam 16808 (Dm); Fe Je Hermann 6610 (Mi, N). Putnam Co.: Grimes 163 (Dp). Saint Jo- seph Co.: Ni Nieuwland 2685 (N, W--6),386)) « aoton Co.: S. M. Deam sen. (C. C. Deam 119] (Dm, Pu). Vermillion Co.: Ce. Ce Deam 1194) (Dm). Warren Co.: Friesner 2285) (Ok, S, St). White te Cost Pipal sen. (Monon, 7/11/12] (Ah). IOWA: Benton Co.: J. J. Davis s.n. [Vinton, 1877] (S), s.n. [Vinton] (Du--2172). Black Hawk Cost A. S. Hitchcock s.n. fuetertnss bellied, setters Le He Pammel | $.n. [Cedar Falls, Aug. 13, '08] (Io—782%, Io—- Be) S.n. [Cedar Falls, VIT-18-1927] kee ren Pamel, a: Gilbert 272, in part (Io—-127005, N); Shimek s.n. ne [July ce Fei) (Ur). Cedar Co.: Fay 09 (Vi), 1017 (Ca—96 ). Cerro Gordo ee A. Hayden 1, a part (N), 3, in part (N, N, N, N). aera 6.2.6; R. Bal Ball s.n. [June 30, 1898] (Io—15308) ; L. He Pammel 7 (Io-53991). F Fiayd Co.: Collector undesignated oot (Io—15 306) Johnson Co.: A. S. Hitchco Hitchcock s.n, Ag City, June 1887] (Io-- 15305), sen. [Iowa City, June 1, 1889] (Ka), s.n. [Iowa City, 1889] (Yo—=15 5307), sen. [Iowa Mest (Io—15303); R. F. Thorne 17397 (W--2328928); Treat san. [July 1865] (Ms). Lyon Co.? Le He Pammel s.n. = Canale, Aug. 28, 1920) (Io—98606). Marion Co.: (Dm), 59003 (N); Ek sen. [6-12-1936] (Bt~-l2533); Erieensr $789 10 Brendel s.n. - (Ur). Wiscatine Co.: W. A. Anderson s.n, [east of Nichols, "eae: 9, 1936] (Ca—-88256)); Este & Brown s.n. [south of Yuscatine, summer summer 1935] (Io--1)6364); Shimek s.n. [June 28, 1913] (NW), sen. (aug. 21, 1915] (ur), s.n. (Moscow, June 28, 1917] 196), Moldenke, Monograph of Verbena 175 (N). O'Brien Co.: Winge son. (Calumet, Maj 1911] (Cp). Scott Co.: E. A. Ross s.n. [Davenport, Aug. 1889] (Se-—-95717). County pee a 3301, in part [Blackstrap] (W--672170). KEN- : Allen Co.: H. A. Gleason 8907 (N). Anderson Co.: Wilburger 1955 (Ka) « Bell Co.: H. A. Gleason 8845, in part (N). Bullitt Dum, & Spellman 913 3 (Lb—-51376). Christian Co.: Ahles 7540 (Ur). (Ur). Ednonson Cos? Li. Cos: Lix 21 217 (W—-1970472); Edw, Palmer s.n. n. [vicinity of Mammoth Cave, May 1699] (N, W—36016). Fayette Co.: —— Son. oe ton, June 14, 1882] (Ka); F. T. McFarland s.n. [Lex ington, May 27, 1915] (Ky——-1776); Re Peter | Sone wn. (Lexington, july 1833) (ui) 5 Short s.n. (Lexington, 1637] (Mi), sen. (1). Frank- Co.: F. W. Pennell 10719 (N). Hart Co.: W. H. Rhoades s.n. (near Macon, July 1927] (N). Jessamine Co.: ts T. McFarland 109 (ky, S, Vi); J. We McFarland 1092 (Ky); Wharton 5906 (Hi~-18d36k). Knox Co.: W. HK. Rhoades s.n. Sn. [Wilton, July ly 1927] ~(N)e LaRue Co.: F. C, Gates 17153 (Ka—80976) ; W. H, Rhoades s.n. [Attilla, July 1927] (N). Livingston Coe: Eggleston 158 (N). Lyon Co.: Eggles~ ton 5237 (N). Madison Co.: A. A. A. A. Wright s Sen, [June 10, '76] (Ob-—- 50758). Marion Co.: Wharton L67h (N). McCreary Co.: H. J. Rogers 630 (Ky). Mercer Co.: F. T. McFarland 2988 (Ky). Nelson Co.: Sen. [June 10, 1933) (I). Perry Cot R. Peter son, [July 6, (ky--1777). Rockcastle Co.: Potzger 3196 . (Bt--13397) 5 a seit & Hodgdon 3705 (N, W--1725668); Wharton on 2595 (Hi--188699) » Scott Go.: Singer r 178 ee Taylor Co.: E. S. lyon 12 (Ky). Union Co.: Shacklet 609 (Ky). County undetermined: Peter s.n. [Kent River, vais 1833] (C); S$ 5 i (1840] a). TENNESSEE: Blount Co.: Curtiss 1955 (C, 900). Carroll Co.: Biltmore Herb. 759 (N, W--332100); Denaree ser 30780 (Au--122691, Au—122693, Hi--51367, Le, Mi, Ok). Carter Co.: J. D. Smith s.n. (July 8, 1880) (W~1323041). Claiborn Co.: Fs E. Lloyd Sen. [Aug. ] (Ka— eG. sen. (Nashville, June ‘SET Ui); P. “ Ve Soak 326 (0k); : Bohn 136 (Lb—23710); Eggleston l30 (Gm, Dt, N, W—506150) 5 oo S.n. (Nashville, May 26, 193h] (F1--1203) 5 ee Palmer s.n (Nashville, Nov. 7, 1897] (W316 318) ; Quarterman 821 82. (H— ~T-108162, Va), 1183 (Au); Shanks 153h (Ca~-882563); Re H. Ward sen. [Nash- ville, March 1862] (Ur). Decatur Co.: Nason Sen. (May 1555) (Ur). Grundy Co.s Calef S65 (Ur); W. H. Rhoades s.n. (Mt. Eagle] (Hs); Richards & & Maguire s.n. (Mount Eagle, le, June 6, 1930) (Ua—8843). Hamilton Co.: J. H. Barnhart 2283 [Herb. ae 2727] (N)- picig Co.: A. R. Moldenk Moldenke 112 Humphreys Co.: Galloway s.n. [June 8, 1889) ( (Ob—50759). i (re). Co.: Biltmore aan: \759¢ (S); Rigel 127 (Bm). Knox Co.: Lamson-Scribner s.n, “Tvie. of Knoxville, 176 PHYTOLOGIA Vol. 11, no. 3 June 1890] (W-—-26002); W. He Rhoades s.n. [Sevierville Pike near Knoxville] (Hs); Ruth 731 1 (345916) ‘ 740 (N), son. (Knoxville, June 1895] (Gg, N, Ob——50750, Po--63886, S, Ur), 3.n sen. [Knoxville June 1900) (Dt). Lawrence Co.: G. Een sen. [26.7.1950] (S)- Me on Cos: Moldenke & Woods 99 (S). Meigs Coe: A. R. Moldenke 7 (S). Montganery Co.: | Ae eRe Moldenke 109 (Fg). Roane Co.: Ne- 7 517 (W—-2175281). Rutherford Co.: Demaree 45755 (S)3 Re Le’ McGregor 16899 (Lm), 17160 (Lw). Stewart Co.: A. R. Moldenke 10 140 (Fg). Wayne Co.: McDougall 1059 (W--1925242). White Co.: Weatherby & Weatherby 6276 (N). “Wilson Co.: C. C. Deam 61340 (Ns Si); Demaree 45737 (S); W. He Rhoades Sone a Lebanon, Aug. 1931; H. N. Moldenke 17076] (Bt, Bt--30065, N, St). County unde- termined: G. B. Grant 2808 [3060] greener AAP Whelden s.n. {May '91] (F-21088). a MICHIGAN: Lenawee Co.: D, Houghton s.Ne _— 14, 1838] (Mi). Saint Clair Co.: C. K. Dodge s.n. [Port » July 12, 1897) (Mi), sen. [July 8, 1903) (Mi, Ob—87LLh) 5 i sen. [Port Huron] ] (Cm). WISCONSIN: Lafayette Co.: He H. Smith 7783 (B, Ws). Milwaukee Co.: J. S. Douglas sen. (Wis.] (Wer 317); Shinners 531 (Ws). Sauk Coe: +. J. Hale son. [Bara~ boo, 1861] (Ws). County undetermined: Skavlem s.n. in. (Harmony, July 3» 1890] (Ws). KANSAS: Allen Co.: A. S. ;s Hitchcock s.n. [alien Co., July eg. (Ka). Anderson Co.: A. S. petchepok k 791 (Ka, N, 7 (Im). ee Fae 2 felleman = ie Ribs onakt ie — 1887} ae Bs me Thompson 541 (Lw). Butler Co.: R. L. eGregor 7257 (Lw). e Co.: Hulbert 339 (lw). Chautauqua Co.: A. S- Hiteh- cock s.n. IGhavteuana ee Co., Aug. 1896] (Ka); We He Horr sen. [July 5, 1930] (Lw); R. L. McGregor 10826 (Lw). Cherokee Co.: A. A. Ja~ cobs 70 (Ka--74127), sen. (Monmouth, June 12, 1932] (Ka—77596)$ a Pe” McGregor 1746 (Lw). Coffey Co.: W. He Horr 3476 (Lw)5 Wine sen. (Burlington, July 1896] (Ka). Cowley Co.: Re Le McGregor T0387 (im); M. White s.n. [June '98] (Ka). Crawford Co.: Ae Se Hitchcock s.n. [Crawford Coe, Aug. 1896] (Ka); We We Holland 298 (Iw); Hulbert 3750 (Lw). Douglas Co.: Le He Horr E.76 6 (Av=—12159%, coer Gg--396798, Lw, sg Sine k, 8, St, Tl, Tl, Ur, Vi, W 20 Tie) ; Re L. MeGr regor 117 (Lw, S); = a 675° (LW), SMe cast Ven ase 3 W. C. Stevens s.n. (Lawrence, June] (Ca-= 1048 32, beanie “Elk Co.: Clothier & eee wy sen, (Auge 20 & Sm. [June | "1898] (Ka); Roof Roofe s.n. [Spring Hill, May 1925] ine 196) Moldenke, Monograph of Verbena Ltt 71597). Labette Co.: Clothier & Whitford s.n, [Aug. 18, 1897] (Ka); Etter s.n. [June 1, 1950] (N); Imler 322 (Ms); Imler & Ryd- berg 322 (Bl—-42279, Lw). Leavenworth Co.: W. He Horr s.n. [6/ 23/30] (Lw). Linn Co.: Kellerman s.n. [Mound City, July 17, 1887] (Ka), sen. (Mound City, July 18, 1857] (c, Ka); H. N. Moldenke 2192 (Lw); Rydberg & Imler 120 (Im). Lyon Cos: CG. J. Elmore s.n. (Emporia, June 20, 1922] (Je—31k6); H. A. Elmore s.n, [Bmporia, June 20, 1922] (Je); W. H. Horr sen, [6/26/30] (Iw). Miami Cos: Clothier & Whitford s.n. (Aug. 8, 1897] (Ka); Re L. McGregor [Neosho Co., July 1896] (Ka) ; W. We Holland 108 (iw); Re L. Me Gregor 10275 (Iw). Osage Co.: Ze D. BE. Brown sn, [June 5, 1899] » July 23, 1927) (Ka--72217). Wabaunsee Co.: Carleton & Nor- ton s.n. [Mill Creek, 7-1)-1893] (Ka); Maus 7h (Ka--72513); Re L. McGregor 9160 (lw). Wilson Co.: Darnell s.n. (June 1890] (Ka); W. H. Horr son. [July 13, 1930] (Lw); R. L. McGregor 10260 (Lr); Vaughn sen. [Aug. 20, 1936] (Iw). Woodson Co.: Clothier & Whit- ford sen. (aug. 29, 1897] (Ka); W. He Horr san. [7/11/30] (Lm); E. We Lathrop 20) (Lw, W—-223)96]), 282 (W—2235008), 386 (Lm, W—22350L9), 611 (Lw, W—2235112), 828b (W--2235153). County un- determined: Ripley s.n. [E. Kansas, June 19, 1899] (Je—31L5). MISSOURI: Barry Co.: B. F. Bush 206 (Ka, S, W~318217), 15051 (Lb—15327). Barton Co.: Z. Williams sen. (Milford, Aug. 1928] (Ib—L172). Boone Go.: Daniels sen. [June 17, 1902] (Lb--918k), Sn. (Columbia, July 3, 1903] (Lb—l168, Lb--l169); Drouet 613 (Ib—-12122), 3122 (Lb—25690); D. B. Dunn 12553 (Lb—38025). Carter Co.: R. L. McGregor 17182 (Iw). Cass Co.: Keetch s.n. (Lisle, July 1, 1934] (Je--3148). Clay Co.: Gier sen, [VI-h-50] (Je—6799). Dent Co.: F. H. Sargent sen. [May 29, 1932] (No 10919). Franklin Co.: J. M. Mason 6.n. [Meramec State Park, 7/ 5/35] (Lb—34372); muenscher & Winne 16728 (Au--12269), Ca— 916312). Greene Co.: Blankinship s.n. (Willard, July 2h, 1919] (Po—638)5); McKelvey 2523 (W—-1581i04); P. C. Standley 8311 (W— 687272), 9336 (W—688251). Grundy Co.: Crookshanks 85 (Lb-- paaccghh Henry Co.: Drouet s.n. [May 22, 1932] (Lb—103h4). Iron ¢? Churchill s.n, [Ironton, May 25, 1918] (Ba). Jackson Co.: = = Bush 285 (C), 3 (Ok), &77 (C), 91h (W-~282373), 7647 (N, 38033), 8647 (Ur), son. [ 7-3-1892] (Lb—L17h); K. K. Macken- 178 PHYTOLOGIA Vol. 11, no. 3 zie s.n. (Lee's Summit, June 18, 1899] (N). Jasper Co.: Demaree 39338 (Z). Jefferson Co.: Hasse sen. [May 1), 1887] (N), Sone [June 1887] (Du--90901). Johnson Co.: E. J. Palmer 5590) (Lb— 50951). Lawrence Co.: Spillman 298 (P1—22609). Lincoln Co.: Je Davis 607) (po-~87289), 6432 (Vi), sen. (Silex, 29-5-15] (Au), 3. ne (Whiteside, 16-15] (Mi), s.n. (Whiteside, 18-9-15] apitre Sen. [7-6-16] (Vt). Macon Co.: B. F. Bush 7586a (N, Ur). Mi Co.: Damon s.n. [Kaiser, May 26, 1939] (Lb—3762) ; A. Re Saris ke 855 (Lw); Steyermark 13056 (Ky). Montgomery Co.: Drouet se [Sune 6, 1931] (Ib—918L); C. O. Erlanson 170la (Mi); Steyermark Truss (Lb—~1,3267) « Phelps Go,: Fleenor s.n. [May 21, 1933] (1b— T1716); Meek s.n. (Rolla, 7-19-89] (Io—15340, Io--22905). Pike sed J. Davis 370 (W—503460). Polk Co.: aes 2409 (N)- Reynolds Co.: Rickett s.n. [Aug. 18, 1927] ( Lb—L173 ae ose Louis Co.: Eggert s.n. [12 Aug. 1875] om Engelmann Pr), sen. [July 1839] (Br, Br, Br); C. Ep 681], (La La); G Gated Se ne n. [near St. Louis, 6-30-92] (Mi); M. Ma Me a ome sen. (Br). Cos: E. J. Palmer 2769 aman “W—-1581046). Taney eae ‘De Spencer 35 (Lb—35371), son. [8.1.40] (Lb--l,8913). Washington Co.: Brinker 708 (I). Wright Co.: Lansing 2976 (Ur), 22 (Ur). County undetermined: Z. Baldwin s.n. (Au); Engelmann Sone rich 7768] (Sm). Saint Louis: J. A. Drushel 1,888 ae i6LB) 5 Engelmann s.n, [St. Louis, Sept. 1041] (W), Sen. (St. Louis] (S)5 F, E. McDonald s.n, [St. Louis, June 1866] (Ca—-192066, Ur). AR- KANSAS: Baxter Co.: Demaree 28939 (Au--122692, PEW 3 N, Ok, We). Benton Co.: Heacox s.n. Y. [duly 20, 189] (Ob—50756). Car- roll Co.: E. J. Palmer Luo (W~1828079) « Clark Co.: Demaree 17178 (N), 17808 (N). Craighead Co.: Demaree 27792 (N)» Fulton Coe? | Demaree 30805 (Au-~122688, Le, St). Independence Co.: Dem~ aree 26752 (Au—122689, Lb—l7028, N, Ok, St). Izard Co.: F- = We Pennell 1 10698 (N); A. Robinson 1792 (Bed « Lawrence Co.: Demaree 26179 (N), 30LL7 (N)3 Rolfs s.n. [8/91] (Io—489h). Marion Co? Demaree 292h9 cahmnageo 8 Hi—51438, Mi, N, Ok, We), 47947 (Lw)- Randolph Co.: Demaree 30955 (Le). Searcy Co.: R. Bebb b leet Sal Turner s.n. [Summer 1939] (Au). County undetermined: F. Le , Har- vey sen. (N. W. Ark.] (Ka). NEBRASKA: Banner Co.: Rydberg 15) (N). Lancaster Co,: J. L. Sheldon s.n, [Lincoln, August — 1902] (We). OKLAHOMA: Adair Co.: C. S. Wallis 2386 (St). = Ce 5. sueidon We 49 (W). Cherokee Co.: Re Bebb 437 (0k, “Oks « S. Wallis (st), 238 (St), 720 (St). Choctaw Co.: Ripley & Barneby 7435 (N); Smith & Randel 69 (Ok); Waterfall 8901 (0k)- 196k Moldenke, Monograph of Verbena 179 Craig Co.: Waterfall 6852 (Ok). Delaware Co.: C. S. Wallis 2296 (St), 2bho (St), 3258 (St), L199 (st). Harmon Co.: | Demaree 12216 (Ok--22237). Leflore Co.: R, E E. Je" s.n. (Winding Stair Mts., 7/3/19] (Ok—10262). Mayes Co.: = Coryell 96 11 946 (St). McCurtain Co.: H. W. Houghton 3622 (N), 366 =~ batter i & Olmsted 10h (Ok— 10263, Ok--1026h). Murray Co. C. Harper 52 (St—23058). Mus- kogee Co.: E. L. Little 158) ‘oman 7) 5 1785 (Ok—10265), 2346 (Ok--10266). Osage Co. : G. W. Stevens A 1980 (St—-9205); Van Vleet 434 (Ok—-20509); Waterfall 1 8179 (Ok). Ottawa Co.: G. W. Ste- yens vens 2299 (N, Ur), A.2299 (St--9206). Payne Co.: Van Vleet 399, _ in part (Ok—-20508), s.n. [yale, 6-25-05] (Ok—-10315).P Pontotoc Co.: De McCoy 605 (ach tae (st). Rogers Co.: Waterfall 1453 (Ok, St), 1469 (S (St). Wagoner Co.: Re Bebb 32h (Ok). Washington Cow: G. W. Ste Stevens A.2080 (St-—920k) . County ty undetermined: Bur- rill s.n. (Indian Territory] (H~10663). UNITED STATES: State unde : Rafinesque s.n. {13.1.41] (N); Torrey & Gray sen. cet omi taste States] (Br). CULTIVATED: Belgium: Herb. Hort. Brux, 3 Herb. Jard, Bot. Brux. sen, (Br); Lejeune s.n. (1822) (Br es sen. [1836] (Br), s.n. (Br); M. Martens s.n. [183] etl son. (Br). Czechoslovakia: M Me Martens s.n. (Br). France: Herb, Decaisne s.n. on. (h. P. 1832] (Br, Br); Herd. W. H. Harvey Re [he Re P., Aug. 1846] (Du--176520). Germany: Bouché s.n. (Herb. Prager 18638] (Gg—31456); Herb. Mus. Bot. Stockholm s.n, . (herb. Hamb.] (S); Herb. Vigener s.n. [Botan. Garten Heidelberg] (La). Spain: Collector undesignated | sen. [1814] (Q); Herd. Hort. Reg. Matrit. 3 (Q), h2 (Q). Sweden: Herb. Mus. Bot. Stockholm s.n. (S); Wallberg s.n, [ex Hort. Proprio 1602] (Lu). Switzerland: Herb. Hort. Bot. Basil. s.n. ca 1838] (C). LOCALITY OF COL- N UNDET 1 Lec: )3 H (bea tat ‘Herb. Ws. —— : Stools 8.0. ise Herb. Fe — Pitcher pias (T); J. Torrey Sen. qe A. Wood s.n. (Pa). CLIPPINGS: Lehm., Nov. Act. K. Deut. Akad. Naturf. Halle ul ( (2): 52h. 1825" (Ww). VERBENA SIMPLEX f. ALBIFLORA maaan: Résumé Suppl. 10: 1, nom. 96h; Phytologia 10: 172. 1 iu apenas tae, Résumé Suppl. 10: 1. 1964; Moldenke, Phytologia 10: 172. 196k. This form differs from the typical form of the species in hav- white corollas. he type of the form was collected by Ronald Leighton McGregor (no. 13217) scattered in low places in shallow soil over lime- stone in native bluestem prairie hay poke 2 miles south of Els- 180 PHYTOLOGIA Vol. ll, no. 3 more, Allen County, Kansas, on July 22, orig and is deposited in the herbarium of the University of Kansas at Ba nee. The form is known thus far only from the type spec Citations: KANSAS# Allen Co.: R. Le WoGregor 13217 (Lw—type). VERBENA SIMPLEX var. EGGERTI Moldenke, Am. Midl. Nat. 2h: 753. 1940. Synonymy: Verbena simplex x hastata Gates, Fl. Kans. 190, in part. 190. Yerbens simplex eggerti Moldenke, Résumé Suppl. 7: Bibliogra Pe te "F.C , Fl. Kans. 190. 190; Moldenke, Am. Midl. Nat. 2h: 7536 190; GAR aeAs Known Geogr. Distrib. Verben- ac., [ed. 1], 9 & 102. 192; Moldenke, Castanea 10: 38 & 39. 1953 Moldenke, Alph. List Cit. 1: 181 (1916) and 2: 05 & 587. 1948; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 17 & 153. lous Nolgeeke in Gleason, New Britton & Br. Illustr. Fl., pre 1, 3: 1 (1952) and pr. 2, 3: 131. 1958; Moldenke, R Résumé 2 2a Sine ‘& L7h. 19595 Moldenke, ésum wv p 1o =] nN a UL * = ® 0 * ON od nm ‘ee 5 Z @ 2 a sasuke abundantly branched from the base, with up to 35 branches per plant, each terminating in an elongated densely many-flowered spike, The type of the variety was collected by Heinrich Karl Daniel Eggert at Pacific, Franklin County, Missouri, on July h, 1896, and is thee diag. gorse in the ican Herbarium at "the New York Botani- cal Garden. The D. J. Johnson s.n., cited d below, was regarded by Gates (1940) as Vv. alex 2 x hastata stata, but does not seem to me to represent that hybrid. Ont = he the rather large, many~ actually represent var. eggerti. Among them are Shacklette (Ky) from Union County, Kentucky; Eggert s.n. [12 Aug. a tom) from Saint Louis County, Missouri; Koenig s.n- [July 1878] (Cm) from Hampshire County, West Viswinies Ruth s.n. [Knoxville, June 1900] (Dt) from Knox County, Tennessee; and Guttenberg s.n. [Sept- 10, '79] (Cm) from Erie County, Pennsylvania. It is very ry probable that ¥. multicaulis Raf. also belongs here. under the names V. angustifolia Michx. and V. simplex Lehm. 1 has been collected along r. tracks and "in grassy open woods, level area 20 feet from creek", flowering September, and fruiting in September. den found it in proximity growing to xV. moechina Moldenke and says that compared to the latter "this material seems to be typical Verbena angustifolia.” The Beason $85 8845 collection appears to be a mixture with typical V. Simpiex In all, besides the specimens mentioned above, 6 herbar specimens, including the types of “ae the names involved, it one mounted clipping have been examined ° 196), Moldenke, Monograph of Verbena 181 Citations: IOWA: Cerro Gordo Co.: A. Hayden 2 (N, N). KENTUCKY: Bell Co.: H. A. Gleason 885, in part , (N). KANSAS: Butler Cos: D.'J. Johnson s.n. [August 1926] (Ka--71717). MISSOURI: Franklin Co.: Begert sen. [Pacific, July 1896] (Ca—183208—isotype, N— Mi eee ¥, OUNTED CLIPPINGS: Moldenke, Am. Midl. Nat. 2h: 753. XVERBENA SOLBRIGII Moldenke, Phytologia 7: 85. 1959. ala Moldenke sehat Suppl. 1: 15, 18, 23, & 25. 1959; Moldenke, Phy- eines ty a, 1961. Illustrations: Schnack & Solbrig, Revist. Fac. Agron. La Plata 29: 255~-266, fig. 1~-k. 1953. This is the hybrid between Verbena tenuisecta Briq. and V. peruviana (L.) Britton, produced artificially in Argentina in hai The original statenent about its origin is "El hfbrido Glandularia laciniata x G. peruviana fué obtenido por ono de nos-~ otros en el Instituto Fitotécnico de Santa Catalina [Zérata, The two parental species occur toget ieee in the wild in at least one state of Braz - araguay, and nine prov- inces of Argentina, so the hybrid is iiveiy to ching ag encountered there. It should have considerable horticultural merit, since both parental species are extremely showy. VERBENA SPECTABILIS Moldenke, Phytologia 2: 26--l,27. Bibliography: Moldenke, Phytologia 2: l26--27. 15.85 ; “Yetdene, Castanea 13: 119. 1918; Moldenke, Known Geogr. Distrib. — 2], 107 & 199. 199; — ee List cit. ks 1256. 7919) oldenke, Phytologia 3: 306. 1950 alisb., Ind. Kew. Suppl. 1s 263.19 1993) ee, Résumé ny & vi "99h Moldenke, Résuné 959. Herb, to “50 cm. tall; stems rather short, obtusely tetragonal, mostly branched at or near the base, rather densely short~pubes- cent with reflexed brownish Mingo branches slender, erect or as- ending, rather obtusely tetragonal, usually sulcate between the ae densely short-pubescent with reflexed sordid-gray hairs; nod S annulate } principal internodes 2—-l; cm. long; leaves de- cussate-oppoai tes petioles very short or usually obsolete; leaf- blades chartaceous, rather uniformly green on both surfaces, o- on larger \eaves 8, ra nsely 54 soph aah above, _ ther Pressed short-pubescent beemaths midrib and the 5--8 pairs 182 PHYTOLOGIA Vol. 11, no. 3 secondaries very slender, usually impressed above, prominulous beneath, the secondaries. rather straight, ascending, branching at ps “apex and a branch extending to the tip of each tooth; inflor- e terminal and in the uppermost leaf-axils, congested- onisate: the pieces er oes portion 2,5--3.5 cm. long, densely many- ter slender, soenbs ye « long, densely short-pubescent with reflexed whitish es osten joined at about the midpoint and there bear~ ing a pair of lanceolate bracts 7--8 mm. long; floral bractlets species @LB gt +a mm. long, about 1 mm. wide at the base (or connivent before and after anthesis; corolla hypocrateriform, vi~- olet or purple, showy, its tube 10--15 mn. long, rather densely puberulent above the calyx on the outside, the limb about 9 mi. The type of this handsome sues was collected by Erik Leonard Ekman (no. a) along a small stream called Magdalena at Loreto, Posadas, Misiones, Arg entina, on a 6, 1908, and is depos= ited in the pactanaed of the Naturhistoriska Riksmuseum te Stock- holm. The species has also been found on low ground and on cam~ pos along the edge of 9 flowering in February and July. In all, 8 herbari specimens, ———- the type, and , moun- ted photographs have roses examined Citations: ARGENTINA: Chaco: T. Ronan "11785 (N, S$). Corrientes: Pedersen 4,8) (Hi--193799, N, S, “W--2283)22). Misiones: 1980 (F--photo of type, 5 lectype N--photo of type, S--typé, Si--photo of type, Z--photo of type). errs SPHAEROCARPA Perry, Ann. Mo. Bot. Gard. 20: 256--257- oe Bibliography: I. M. Johnston, Proc. Calif. Acad. Sci., ser- by 20: 88. 1931; P Perry, Ann. Mo, Bot. Gard. 20: 2h3, 219, 250, ie els & 356. 1933; A. W. Hill, Ind. Kew. Suppl. 9: 295. 193 Moldenke, Known Geogr. Distrib. Verbenac., {ed. 1], 19 & 102. own Geogr. Distrib. verbenac eds 2], 33, 3h, - 199. orcad | Moldenke, Résumé hO % 174. 1959} tsidbaics; Phytologia 8: 257 & 406 neers 6s. yea Sgt ae 9: 19h. 196h. Ann erb, ol 3 stems erect, tetr tragonal, branched Ae wre ams or nen Ss scabrous-pubescent; petioles short; feat clans ne. és tapering into the petiole at the base, sharply serrate along the ~ oO . i « 8 ct o fo cr ct ® f tad ~~ with the hairs minutely bulbous-based, or less short-strig- illose on both surfaces, Sreatanette arose beneath; peduncles to 196k Moldenke, Monograph of Verbena 183 9 cm. long; — sometimes short and dense, subsessile or short- pedunculate, or to cm. long, cymosely arranged; bractlets ont pa thar FSi than the calyx, acute at the apex, pub- i hi corolla hypocrateriform, light~pink or pluish-white, its tube protruding slightly beyond the calyx, the limb pe ae mm. wide; schizocarp about 1 mm. long and 1 mn. wide; cocci very faintly pad as a ate or essentially smooth, the commissural faces muricately scal The ¢ type of this strange endemic species was collected by Al- fred Webster Anthony (no. 380) on Socorro Is » co , Mexico, between March and June, , 1897, and is deposited pee the hertenton tani c been found on rocky soil, flowering and fruiting from May to July. Material has been misidentified and distributed in herbaria as vy. bonariensis L. an ve litoralis H.B.K. Reid oe, ina leben to me dated September 13, 1959, says “My specimens differ from Perry's er in having longer Spikes, whiting to 4 cm. long, borne on peduncles as long as 9 cm. According to my notes the Saar were light pink, though pereen, as quoted by Johnston.....said bluish-white." rry (1933) cites the following additional specimens not as yet s eoen by me: REVILLAGIGEDO ISLANDS: Socorro: A. W. Anth (E-type, Giedaotype)s Barkelew 231 (E, G). She says "This en- demic from Socorro Island is most nearly related to y. litoralis and has a similar habit, but is easily separated on on the distinct- ive characters of ee inflorescence. The spikes are shorter and denser, the flowers are smaller and so crowded that the lower ones appear to ae i acried at right angles ¥a. ity rachis of the spike; moreover, the is as or even » ully as broader than long, an unusual trait not found pos in the North American species of Verbena In ali, 16 ee Spree ie has the type collection, have been’ examined ee Citations: remind fee ISLANDS: Socorro: A. W. snttow ZC 380 (Ca-~10),821--isotype sot; —~ 3139 34—isotype) ; Barkelew 231 (Ca wr be28, seats TY N, Po—6h6L5, 39902); H. L. Mason 1612 (Gg—-186593, W--1569662); Patifio ee (Me)s 0, Solfs 20 (Me, W-—-126539h), 70 (W-~1268885), s.n- “san, [9 Maro 1925] (Me). VERBENA —— SEH Bol. Mus. Hist. Nat. Jav. Prado 7: 2h2, nym. Bibliogr. rae Raimondi, Bol. Mus. Hist. Nat. Jav. age - 22. 1943; Moldenke, Castanea lo: kl. 19h5; 3 Moldenke, Known Distrib. yforeenac., [ed. 2], 7h & 199. 1949; Moldenke, Résuné 85 Sietcccagear ss own to me of this plant except that the type is an unnumbered pois aekton made by Antonio Raimondi at Chanchamayo, 18), PHYTOLOGIA Vol. 11, no. 3 in the province of Tarma, Junin, Peru, in 1855. VERBENA STELLARIOIDES Cham., Linnaea 7: 26-265. 1832. nymy: Verbena stellarioides & decurrens Cham., Linnaea 7: 265. 1832. Verbena morongii Britton, Ann, 7. Y. Acad. Sci. 7: 198. 1893. Verbena decurrens (Cham.) Kuntze, Rev. Gen. Pl. 3 (2): 257. 1898 [not V. decurrens Moench, 197]. Varbina. stellarioides var. decurrens Cham. ex Moldenke, Suppl. List Invalid Names 10, in syn. 1941. Glandularia stellarioides (Cham.) Schnack & Covas, Bol. — dont on lz 28h. 1946. Bib aes es Linnaea 7: 26-265. 1832; an Nom. - 33 j vis ee: 1937; Moldenke, Prelim. Alph. List Invalid Names 7. 1940; Molden- ke, Suppl. List Invalid Names 10. 191; Mo eeaanie; Alph. List In- valid Names 9 i 50. 192; Moldenke, Known Geogr. Distrib. Verben- ac., [ed. 1], 39, ll, & 102. 192; Schnack & Covas, Bol. Soc. Ar- ah. 1 19465 ‘Troncoso & Burkart, Darwiniana 7: 211, , Known . Distrib. Verbenac., [ed. 2], 9h, 99 & 09. 195 * Moldenk Lgl: List Cit. 3: 688 & 923. 19493 Moldenk » Phytologia 3: 289 ‘ "378. 1950; Schnack & et Revist. gin. vagron. a eae lara he 8 1 eric denke, Résumé Suppl. 2: 13 (1960) and 7: 6. 1963; Moldenke, Phy- vologis 9: 130 (1963), 10: 159 & 160 (196k), and 1]: 1h] & 12. Illustrations: Troncoso & Burkart, Darwiniana 7: fig. 2a- 196; ee & Covas, Revist. Argent. Agron. 18: ee 2 fig. 9 Suffrutescent perennial herb, with the habit of Stellaria holo~ stea; rhizomes A nbn et erect, O cm. to 1 a. tall, slender, tetrag the pita subulate-acuminate at the apex, villous- Bagh Magid large, es cis be membranous, color ed, tub- ular, les out 6 mm. long, 5-rib villous on the angle 196), Moldenke, Monograph of Verbena 185 teeth 5, subulate-aristate, ascending, the 2 lower ones larger, the 2 lateral ones somewhat shorter, the uppermost one short; corolla large, hypocrateriform, varying from lilac, purple or vi- ing ¢ the stamens, elongat e and exserted from the calyx after the falling of the corolla, etavsts Hiisvadbits at the apex; stigma subapical, with an adjacent, minute, oblique, acute horn; fruit~ lo cylin h t oblong, about mm. long, slightly thickened at the base; cocci linear, obtuse, convex and elegantly sens on the back, the areoles quadrangular, the commissural surface angulate, smooth; x = 5. The type of this species was collected by Friedrich Sellow in grassy campos at Rio Jacuy near Banhado, Rio Grande do Sul, Bra~ zil, and was deposited in the herbarium of the Botanisches "wuseun at Berlin before World War II, now destroyed, but fortunately represented by Mac nrdda! a type photograph no. 17446. I regard this photographed specimen also as the type of var. decurrens, the ae al variety. I follow Schauer in regarding var. decurrens &S a synonym of the typical form of the species. The so-called var. sessilis Cham, I regard as V. sessilis (Cham.) Kuntze. The ee two species are not very clearly defined, but, in general, V. Stellarioides has the leaves decurrent, linear, entire, and Y glab- Tous (or strigose on the ma and Sih ‘veka ots longer rgins only), than the calyx, while in V. seaatité the leaves are merely ses- Sile, narrow-lanceolate obsoletely serrate, and sparsely strig- illose, and the bractlets are shorter than the calyx. The bract- however, to be the only reliable character. The type of V. morongii was collected by Thomas Morong (no. 600) -- in mare = honor $¢ As named —- along railroad tracks at allero, Paraguay, on January 20, 1889, and is deposited in the Columbia eb ae at the New York Botanical Garden. Verbena m morongii was regarded as conspecific with V. ae by me in my 190 re nt n. Verbena stellarioides has been found in swamps and — places, fields and moist grassy rena in foggy places, on low palms ong r oad tracks, and on sree on zr Campos, among er to Jan abundant ae Cohonis Bentten in Chaco, while JUrgensen says that it is also common m in open places in Paraguay. Walpers (185) classifies this species in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Macranthae, and Sec-_ ondary Subgroup Me Welindres, along with ten other species. It should 186 PHYTOLOGIA Vol. 11, no. 3 be noted here that the ve decurrens of Moench is a synonym of Stachytarpheta jamaicensis (L.) Vahl. While this binomial is ac- credited to Moench, actually Verbena is mentioned by him in his aig (1794) only on pages %8, 369, & 423 and no such binomial 8 on pages there, either; however, on page 150 he pang ae a eee as a new name for Stachytarpheta amaicensis. erbena decnrzens” is given in Kuntze, Rev. Gen. Te TG 2 257 ( 8 98) with ham.)" implied. The Morong, Britton, & Vail reference inthe tibataieas above is often credited to Britton alone and erroneous~ ly dated "1892", Troncoso & Burkart (1916) ons, mus stellarioides posee espigas terminales solitarias, rara vez 26 3 espigas por heeaina ees inflorescencia capituliforme re senen . 1 antesis; intermodio ter- minal (debajo de la espiga) visiblemente velloso; flores mucho mayores [as compared to V. tristachya], con limbo amplio y exten- dido, tubo corolar exteriormente velloso-lanuginoso; brfcteas muy su das, m4s largas que el c4liz, velloso-ciliadas; c4liz vel- loso en los 4ngulos......Ve seas cone también een Material of V. stellarioides has been misidentified and dis- tributed in herbaria under the names V. littoralis Kunth, V. ses~ silis Kuntze, and V. stellarioides sessilis Cham., while Rojas 1680 Was called "VY. stellarioides Cham. sensu lato” by Hassler. ~~ Schauer (1847, 1851) gives ve peredecet Spreng. as a syno- nym of V. stellarioides with a “question and the qualifying remark "ex diagnosi", but Sprengel's name actually applies to a very different species, which see. Schauer says of V. st stellarioides "Planta conspicua, facie Stellariae Holosteae haud omnino dis- similis, sed firmior." Chamisso (1832) s wy8. "cyma hac lege ter unispi curri omnino tranc — bifurci; bifurci etiam rarius ex inflorescentia, ramis ¢ summorum foliorum axillis evolutis foliosis spicreris spicam primariam alarem comitantibus" and of the cocci he notes "color illi griseus pallide nigrescens. Carpello dorso gp cence fac- iebus albidis nudiusculis, angulo faciale nervoso. Schnack & Covas (1951) studied the chromosome caine of this species and of V. tristachya Troncoso & Burkart and report "En G. arate | sobre la unica planta obtenida de semilias coleccionadas po el Ing. L. B. Mazoti en Ituzaingo (provincia de Corrientes), teal costatado en microsporocitos y en celula somaticas 5 pares de cromosomas y un fragmento centrico libre.-+- En todas las metafases mitoticas observadas esta presente el fragmento, lo que indica que se divide en cada mitosis. En 1964, Moldenke, Monograph of Verbena 187 proceso meiotico se observo que el fragmento pasa un cc) ® 3 E po) g as & & 8 Fy ] & b 5 Bs he BE: 8 un vaso se verifico a§ prometafase I la pres~ 28 herbarium speci- salt and 6 sina {lineation te cluding’ type or omits Lee material of all the names invol ved, ai be been examin Citations: BRAZIL: Rio Grande do $ ul: Jurgens 22 (B); Rando 10156 (Rb); Schreiner son. (Ja—6575, ms Sellow s sn. [Macbride photos 17kh6] (Kr--photo of type, N—-photo of type, N—photo of ype). PARAGUAY: actin 33 is rb. sen 2221} (Cp, Du-- m Les Fe N, Fe op N--photo, S$ 3812, Z~=photo) ; Kuntze’ s.n, [Nord Paraguay, aie 1692] (N)j Morong (C)5 Pedersen 4262 (W--22837)2); Rojas 1680 (Herb. Hassler r 1680; Herb. Osten 86)1] (N, Ug), 2527 [Herb. Osten 1818] (N, Ug). URUGUAY: Arechavaleta 18 (N, Ug, Ug, Ug), 36 (Ug), 5.n. Sdegeaie , Nov. ENTINA: mo es A. G. Schulz 285 [Herb. Osten 2319] (N, Ug), met (It, o genero,. Tag oma age). aioe cites a gait “sprays from Cerro yet seen by m In all VERBENA S I Moldenke, Le Se ay 2: 56--57. 19k1. Bibliography: Moldenke, : 56-57. x91; ge own Geogr. Distrib. pte ay oe i, 3h & 102. ag Fr ke, Castanea 13; 116. 198; H. N. & As Le Moldenke, te 2 8h. 1948; Moldenke, Alph. List Cit. 3: rte 2943p own Geogr. Distrib. Verbenac., [ed. 2], 71 & 1919; E. Je Salisb., Ind. Kew. Suppl. 11: 263. 1953; Moldenke, Pesaeb OL & his 1959. rb, more or less nigrescent in drying; st ~ branches very oan S the latter almost filiform, sharply te onal, terminal, srtmeies rather few-flow during anthesis the lower flowers often scattered after — “i nf s filiform, glabrous and shiny or very obscur soa ttered-malvesnt rulent; *practlets lanceolate, 1.5--2 Uminate at the apex, glabrous except for the ciliolate margins; Calyx tubular, about 2 mm. long, minutely appressed-puberulent; 188 PHYTOLOGIA Vol. 11, no. 3 corolla hypocrateriform, barely exceeding the calyx, its tube Stewart (no. 3320) -- in whose honor it is named —~ at Tagus Cove, Albemarle Is sland, Galapagos Islands, on March 27, 1906, and is deposited in the. Britton Herbarium at the New York Botanical Gar- den. The collector states that the species is common in lava beds at 300 feet altitude, It has been misidentified and distributed in herbaria as V. litoralis H.B.K., while the Bishop Museum iso- type is actually mounted on the same herbarium sheet as Stewart 3319, which is ¥. galapagosensis Moldenke. ~~ Verbena stewartii has been found in flower and fruit in March. In all, 3 herbarium specimens, snoeing the type, and ), mounted photographs have been examined Citations: GALAPAGOS ISLANDS: rans A. Stewart 3320 cael isotype, F—-photo of isotype, Gg-- --31381—-isotype, N—t photo of isotype, Si--photo of isotype, Z--photo of iia ee refs VERBENA STOREOC REOCLADA ae in cae & Hassler, Bull. Herb. Boiss., Synonymy: Vi Verbena rinse Chod. (in part) ex Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, h: oa in syn. 190k 1963, jog derlein, 1881, nor L. ex Wal Rea oepp., 1847, nor Spreng., 1830, nor Willd., 1947] : Perbent ss ord hc Chod. (in part) ex Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, he 1059 & 1060, in syn. 190). Verbena stereoclada Briq. ex angely, Fl. Paran,. 16: 79, sphalm. 1960. V Verbena humilis Cham. ex Mol- sér. 2, 4: 1059--1061. 1904; Briq. in Chod. & jeholee oF Plant. Hassler. 10: 82. A toa ghar Ind. Kew. Suppl. 3: 187. 1908; Moldenke, Known Geogr, Distrib. Verbenac., [ed. 1], 41 & 102. 1942; Moldenke ne List set 1: 263. 1946; Moldenke o- gia S 2 or & 386. 1947; Moldenke, Castanea 13: 117. 1918; Mol- denk - List Cit. 3: 693, 70h, & 766 (19k9) and h: corn 1172. ‘5i9 “Wold enke, Known Geogr, Distrib. Verbenac., [ed 9h, 99, 100, 107, & 109. 1949; Moldenke, Phytologia 3: 289. 19503 Stellfeld, Trib. Farmac. 19 (10): 167. 1951; angely, abel winged 7: 13. 1957; Moldenke, Résumé 111, 118, 120, 128, 363, & 47h. 19595 Angely, ri. Paran. 16: 79 (1960) and 17: 7. 19825 aie Résumé Suppl. 3: 11. 1962; Moldenke, Phytologia 8: 202, 117 420 (1962) nite 9: 12, 67, & 70. 1963; Moldenke, Résumé suppl. t 9+ 1963; Moldenke, Phytologia 9: 388 & 393. Herb LO cm, tall; stems many from a single root, rather weak, procumbent, somewhat angled, eipcet branches divergent- ascending, green, sparsely retrorsely pubescent; middle inter- nodes 2-6 cm, Long; petioles to 1 cm. long, winged, often a4p~ 1964 Moldenke, Monograph of Verbena 189 aring falsely se Pipi a because of smaller oO broadly cuneately decurrent into the ge axes, nate- e ur’ at the base, pinnately ese with the lobes obtusely c serrate, often rather wide, sparsely antrorsely adhe teicodet pubescent above, glabrescent beneath except for the prominulous veins; scence spicate, more or less pedunculate; peduncles Slender; spike inal, finally elongate to l cm., congested; s te actlets lanceolate, about 5 mm. long, shorter than the calyx, sparsely ri opel on the back, ciliate-pubescent along the mar= gins, spreading after anthesis; calyx tubular, about 7 mm. long, dexcls- pilose te short spreading or slightly ascendi s mi. Long oblique, the teeth setaceous, unequal, the 2 anterior ones lon- ger, about 2 mm. long, the r res st 1--1.5 mm. long, all finally con- vent; corolla hypocrateriform, rose or violet, surpassing the calyx-teeth by 1~-2 mn., pube erulent on ne outside, its tube regular, cylindric, somewhat exserted, the limb spreading, 5--6 mm. wide, the lobes obcordate, the pag ones about 2 mm. long 6, Fads téeniy 01 mm. at the apex; stamens = The name, V. humilis, seems to be based on Malme 1322 aha Sandy rather wet fields at fasta , Rio Grande de do Sul, Brazil, Collected on February 2, 1902, and ph in the ihicten of the Naturhistoriska Rikemuseum at Sto i It should be noted here that V. avinciaas and V. crinoides of Chodat are also in part V. calliantha Briq.; the V. erinoides of "Auth," is V. dissecta Willd., that of Humboldt, Bonpland, Kunth is V. ciliata Benth. , that of Linnaeus [ex Walpers] and of Lamarck is V, laciniata (L.) Brigq., that of Willdenow is V. lac- iniata var. contracta (Lindl.) Moldenke, and that of argh of Hooker & Arnott, of Poeppig, of Sprengel, and of Linnaeus Lorentz & Hledeflein] ie Ve berteril (Meisn.) Schau. Macbride's Photograph no. 24698, sited below, is of a specimen depoisted in the Delessert erbart: Verbena storeoclada Fe been found in fields and in sandy 190 P=: ¥°F°05L O.G:1-k Vol. 11, no. 3 rather wet fields, at sagem - 1100 to 1200 meters, flowering from January to Mar ch and in and September. In all, her- barium specimens, including “i “Aetertal of all the names invol- ed, 2 Ce Citations: BRAZIL: Paran4: Tessmann 3701 (Herb. Mus. Paran. 3701] (N). Rio weed do Sul: = Be (S); J. Vidal s.n. {near city of Torres, Feb. 1939; Herb. . Rio de Jan. Boul) (N), s.n. [Bocca do sate: March 1939) (Je--heshé) . Santa Smith & Reitz 10193 (W-2251660, Z). PARAGUAY: festa “M89 (ac bride photos ay } Fel a ae otype, Kr—-photo of oto of type, N—- isotype, N--phot type)» URUGUAY: Née 8) (N, Q). ARGENTINA: Santiago del ee H. H. Bartlett 20152 Th W--1907605) « VERBENA STRICTA Vent., Descr. Pl. Jard. Cels. 53, pl. 53- 1800- Synonymy: Verbena alopecurus Cav., Descr. 68, 1802. Verbena rigens Michx., Fl. Bor.-Am., ed. 1, 2: 1h. 1803. Verbena foliis ovato-oblongis, crenato-serratis ramisque pilosis Vahl apud Poir. in Lam., Encycl. Méth. Bot. 6: 549, in syn. 1808. Verbena (rigens), caulibus rigidé erectis; foliis obovalibus, subtomento- so-hirsutissimis; spicis Se Michx. apud Poir. in Lam., Encycl. Méth. Bot. 8: 549, in syn. 1808. Verbena ( stricta), hirsuta, albicans, spicis cylindraceis; foliis subsessilibus, vatis, | serratis; caulibus strictis, teretibus Vent. apud Poir. an rs Se ahs pt Bot. 8: 59, ta syn. 1808. Verbena cuneifolia .N. Y., ser. 2, 5: 360. 1808; Journ. Bot. Eperse 1: i. Shee oy Syne Verbena ringens Raf., First Cat. Gard sylv. Univ. 16, nom. “nom. nud. d. 182); Herb. Raf. 61, a nud. 1833. Verbena stricta @ mollis Torr., Ann. Lyc. N. ¥. 2: 234. 1827+ Ver- bena mollis Haf., Atl. Journ, 1: 146. 1832. Verbena scoparia ausch, Flora 19: 390. 1836 [not V. scoparia Gill. & = aes nor Hook., 1947]. Verbena stricta var. mollis Torr. ex Eaton right, N. Am. Bot., ed. 6, 470. 18h0, Verbena alopecuroides Hort. ex Steud., Nom. Bot., ed. 2, 2: 79, in syn. 1841. Verbena alopecurioides Hort. ex Steud., Nom. Bot., ed. 2, 2: 750, : i8hl. Verbena striata Vent. ex K. V. 0. Dahlgren, Bot. Notiser Lund 1923: 10, sphalm. 1923. Verbena stricta L. ex Moldenke, Suppl. List Invaiid Names 10, in syn. 19 iki. Verbena stricta Michx. ex Moldenke, Alph. List Tovalid Names 50, in syn. syn. 1942. ver Verbena strica Vent. ex G. L. Fisher, Am. Bot. Exchange List, sphalm. 19))6. Verbena rugosa Michx. ex Moldenke, Alph. List Invalid Names Suppl- 1: 26, in syn. 1947 [not V. rugosa D. Don, 1836, nor Mill., 1768, nor wuhl. , 1809, nor Sweet, 165]. Verbena stricata Vent. ex Mol- denke, ish: List Invalid Names Suppl. 1: 26, in syn. 1947. Ver~ bena stricta x hastata Gates (in part) ex Moldenke, Alph. List % In- valid Names Suppl. 1: ; 27, in syn. 1947. Verbena strieta Vent. ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 1947- 1964 Moldenke, Monograph of Verbena 191 Verbenia stricta thee ex Moldenke, Alph. List Invalid Names Sup- pl. 1: 28, in sy in - 1947. Verbena Sai ites Barkley, Nat. Mag. 50: [309], in jest. heer, Veebena stricta Vent. ex Moldenke, Résumé Suppl. 2: 10, in syn. 1960. Verbena stricta Benth. ex Moldenke, Résumé Suppl. 3: 41, in syn. 1962. Verbena stricta Willd. ex Moldenke, Résumé Suppl. 3: 1, in syn. 1962. Verbena stricta f. stricta Wallis ex Moldenke, Résumé Suppl. 3: 41, in syn. Nn. 19625 Steyerm., Fl. Mo. 1260. 1963. Verbena hastata stricta Rydb. ex Moldenke, Résumé Suppl. 5: 10, in syn. 1962. Verbena structa Vent., in e Bibliography: Vente, Descr. Pl. Jard. Cels. 53, pl. 53. 1800; Cav., Descr. 68. 1802; Michx., Fl. Bor.-Am., ed. 1, 2: lk. 1803} * Tran sylv. Univ - 16. 182k; Spreng. 4 Le, te Veg., & Hort. Brit., ed. 2, 418. 1830; Raf., Atl. pe Raf., Herb. Raf. st. 1833; Hook., Comp. Bot. mks 1: 176. re 750 D. Dietr. Walp., Repert. hs 23 & 33. 1845; Schau. in A. DC., Prodr. 11: 547 & 555. 1847; A. Gray, Syn. Fl. N. Am. 2 (1): 33. 1878; Pickering, Chron. Hist. Pl. 10h, & 1220. 1879; A. W. Chapm., Fl. South. U. S., ed. 2, 636. 1889; Wats. & Coult. in A. Gray, Man., ed. 6, 402. 1890; A. 3. Hitche., Trans. St. Louis Acad. Sci. 5: Sie. 1891; Coult. ag CORSE U. S. Nat. Herb. 2: 327. 1892; F. D- Bergen, Journ. Am. Folk-lore 5: 102. 1892; Briq. in Engl. & & Prantl, Nat. Pflanzenfam. (3a): 148. 189); Jacks. in Hook. f. & Jacks», Ind, Kew. 2: 1178 & 1179. 1895; L. H. eal in A. gee Field Forest & Gard. Bot. 341. 1895; Hitchc. & Bull. . Agric. Sta. 50: 54, pl. 6, fig. 91 (1895), 57: me “3, fig. 138. (1896), an : pl. 25 fig. 138. 1897; A. W. Chapm., Fl. South. U. S., Sa 3, 368. 1897; Hitche. & Norton, Bull. Kans. Agric. Sta. 76: pl. 9, fig. 7. 1698; Britton & Br+, TI Illustr. Fl., ed. 1, 3: 71, fig. 3061. 1898; . Hitche., Fl. Kans. pl. ite 1899; Ne Le Britton, Man. 778, tos a . Small, Fl. Southeast. U. S 100 1903; Michx,, travels West Alleg., ed. sa ict. "ots Robin- Son & Fern, in A. Gray, New Man. Bot., scale igs Page Ua + Day, Check List 128. 1908; L. H. Pamm se 218, 1911; Beal, Mich. aeons? {Bull. iia; Aeric. Exp. Sta. 267: ] 396. 1911; Gerth van Wijk, Dict. Plantnames 1: 1398. 1911; Blew itt, Rhodora 13: 89. 1911; L. H. Bailey, Botany 372. 1911; ang Ann. Rep. Mich, Agric. Exp. Sta. 25: 386. 1912; Iowa Geol. Surv. Bull. 4: 265 & ee 1913; Britton & Br., Illustr. Mm, ts, 3 192 PbS: FOL O.6:74 Vol. 11, no. 3 96, fig. 3556. 1913; Je Ks Small, Fl. Southeast. Ue Se, ed. 1009. 1913; N. Taylor, Mem. N. Y. Bot. Gard. 5 [Fl. Vic. Ne 3 ais 526. 1915; E. J. Palmer, Ann. Mo. Bot. Gard. 3: aay 1916; Goris Notiser Lund 1923: 10. 1923; A. B. Seymour, Host Ind. Fungi N. Am. 587--588. 1929; Strausbaugh & Core, Proc. W. Va. Acad. 4: h6. 1930; Gier, Trans. Kans. Acad. Sci. 34: 261. 1931; Stapf, Ind. Lond. 6: bi. 1931; Schwencke, Zytol. Untersuch. Verhenac. 3, 19, & 20. 1931; Wolden, Proc. Iowa Acad. Sci. 39: 122. 1932; BS 1932 Wadmond, Rhodora 34: 19. 1932; J. K. 1, Man. Southeast. Fl. 1136. 1933; Perry, Ann. Mo. Bot. Gard. 27, 260, 285—289, & an 19333 hainnsce | Symb. Bot. Upsal. h: “vel & 171. 193k; W. C Muenscher, Hi pre 1] Univ. Agr. Exp. Sta. Bull. 635: 10. 1935; Patermann, Beitr. Zytol. gf Awaad: 6:- 19, . 43. 1935; L. He Bai- ley, Cat. ’ Florists Handl. V. benac., "uss. 19355 s ape aos Ann. Mo. Bot. Gard. 22: éo9. » 19383 Le i. & EL Hortus, new rev. ed., 632. 19 935; Re Oo Eaton, Rhodora She ore “1336; Der=- » Cytologia 7: 161—16l, fig. 16. 1936; Cory, Texas Agr. Exp. Sta, Bull. 550: 89. 1937; Moldenke, Revist. Sudam. Bot. h: = - 1937; Truman, Rhodora 0: 2hh——2h5. 1938; Little, Am. Midl. Nat. 19: 385. 1938; Moldenke, Alph. List Common Names 7, 12, & to O: 83. 190; Gates, Fl. Kans. 191, map 1071. 190; C. C- » Fl. Indiana 797—-798 & 1232, map 1723. 190; W. A. Weber, prea di Fl. Northrop » mss. 1940; Moldenke, Suppl. List Com- honey mon Names 9, 15, & 23. 190; Moldenke, Prelim. Alph. List Invalid Names 1,8, 1510; seiderke. Suppl. List Invalid Lap 10. 191; Gates, Weeds Kans. 2, 90, & 91, fig. 19 pl. 6, fig. 91- 19h Moldenke in Lundell, Fl. Texas 3 (1): 16& eaotae 192; Co Ne Ed a Anal, Ee Fitotéc. Sta. Catalina h: 18 & 19. exe iebel, Yuncker, & Friesner, Proc. Ind. Acad. 127, 1912) and bos 10h. 1943; H. D. House, Torreya 2: 188. “19h34 E. L. Braun, Annot. Cat. Spermat. Ky. 116. 193; Moldenke, Known Geogr, Distrib, Verbenac. Suppl. 1: 1 & 2. 1943; Schnack & Covas, Darwiniana 6: Annal, (Am. Midi. Nat. Monog. 2:] 216. 195; Deam Kriebel, Yuncker, & Friesner, Proc. Ind. Acad. Sed. 5h: 95. 19h5; Seed Trade Buyers 609 & 610. 19 Guide 86, 1945; Moldenke, Am. - Bo : - 1953 ldenke, Bot. Gaz. 106: 159--161. 19,5; Moldenke, Phytologia 2+ 68, 4, 1, shat & 117 (1945) and 2: . 4, Ge a fee : ae pet Moldenke, Alph. Li Cit. la a 29-—31, 33--35, 37, lO, pus, Mt oe 2, 7 77, 80, aa 38, io 93, 94, 98, 100, 103, 10h, 107, 196), Moldenke, Monograph of Verbena 193 O9—-111, 115, 129, 130, 136, 137, lb2--145, 1y8—151, 153, 155, 157--160, 162-~16);, 169, 17h—-176, 182—~18h, 191--195, 197, 201, 207, 209-217, 22h--226, 236, 239, 2hh, me 218, Fis pe 262, 267° 268, 271, ate 278-2, 287, 291, » 2h, & 297. 196; Pell ett, Am. Honey Pl. the 420. But, gona Fl. Del. 218. 9173 sakdenies, ara sc 2: 326, 327; 3h8 Moldenke, Alph. List Invalid Nan s Suppl. 1: 26—~28. 1947; nig ‘Oo Ww . a) ve) , co we [-) "> ms . a4 iu e e wm) . qa co = cy B @ bd * A i : 198. ; astanea 13: “ie oa po ae Moldenke, Wrightia 1: 225. 1948 Moldenke, Alph 354, 356, 361, 391~L01 wecbsr lak, 4292 138, ye, “Ws, “168, 468, ud, LTT, ‘ie 5, 536, ehh? 6) 356.-c62, 568, 569, 577, eh 58h, 585, 587, 588, 592, 59h, 596, 597, 601, 60h--606, 608, 612, 61h, 615, 620, 621, 630, 63h, 639, & (1948), 3: 655, 657, 658, » 667, 668, 671, 673, 700, 707, ; , 72h, 726, 728, 737, 740, 741, 745, 750, 752, 253, 755, 761, 67" 768, 773, 790-793, 800, 802, 805, 806, 811, 816, 828, 831, 839, 843, 851, 852, 85), 856, 358 09, 869, 881, 886, 092, 895, 90h, 915, 917, 926, 927, 930~-932 , 946, 950, 953— 957, 963, 969-~971, & 978 (2919), and is oe 981, 983, 985, 7588, 988, 990, 992, 99h, 996, 998, 1002, 1005, 1010, 1013, 1015, 1083, >t 098, 1100 2 8 > 1187, 1189, 1191-1193, 1199, 1208, 1210--1214, 1216--1218, 1221— 122k, 1226, 1228, 1231, 1232, 1236--1238, 23, 1252, 1255, 1260, 1261, 1289, 1294; 1297, & 1299. 19495 H. Ne & As Le Moldenke, . 20; 99 ie 295. 1949; Moldenke, é 2-~5, 7, 12-21, 2h--26, 33, 16h, & 199. 1949; Re usen, Cornell Un: r - Sta. 291: 3. 1919; Moldenke, Am. e Exp 3 Wild Fls. 292, 293, & h50. 1949; Moldenke, Phytologia 3: 71, 72, 131, 132, & ui. 1 1919; M. L. Grant, Proc. Iowa Acad. Sci. 57: 950; W Revist. Sudam. Bot. 8: 166. 1950; Moldenke, Phytolog (1550) and 3: “50 & 468. 19515 Moldenke in Chit on, real Hort. » Dict - 6: 2 2i2. eit a eae in Gleason, New te & oy Timers ve, mie ae i 126, 129, & 132. 1952; H. C. Greene, Am, Midl. Nat. 48: 52. "195825 Voldenke, Phytologia 4: 66—~68 (19525 and hs 185, 186, & 266. 1953; Boivin, Rhodora 55: 225. 1953; Deam, Yuncker r, &F Friesn , Proc. Ind. be cad. Sei. 62s qi2. 1963, W. He can ao . Sci. 12: 100. 19533 H. Ce and 28; 3507. 1954; Hylander, ‘VacMil. d Flow. Book [331] & 340, pl. 166r. 1954; Moldenke, Inform. a, A 48 Spec. [lL]. 195k; Re F. Thorne, Proc. Iowa Acad. Sci. 62: 182. 19553 W. D. Barkley, Nat. Mag. 50: [309]. 1957; Strausbaugh & Core, Fl. W. Va. 3: 77k, 19h PEYTOLOGIA Vol. 11, no. 3 776, & 777. 1958; Moldenke in Gleason, New Britton & Br. reso Fl., print. 2, 3: 126, 129, & 132. 1958; Moldenke, Phytologia 6; 300. 1958; Moldenke, Am. Midl. Nat. 59: 345, 347, 352, 3, git 7 : : de - Wildfls. pl. 155. 1961; Moldenke Phytologia 8: 120, 121, & 12h os and 8: 2h6, 257, 268, 277, 280, 316, 317, 407, "& 435. 1962; ° Torrey Bot. Club 89: 56. 1962; Moldenke, Résumé Suppl. 3 2 2, 1 ae fs 283 296, 307, . 315 (1963), 556-359 Goalye bs “iiss, 2935 & 307-310 (196k), and i 8h, 103--105, & 113-118. 196); Moldenke, Résum 5. Suppl. 10s 1& Seen and 11: 13 & 8. 196k; Henry & Buker, Trillia 12: 39. v : & Norton s. p pl. | (fe e138. (1896). "66: ml. 12, fig. 138 (1897), and 76: pl. 9, fig. 47. 1898; Britton & Br., Illustr. Fl., ed. 1, 3: fig- Bull * fig. 3556. 1913; Kanda, Bot. Gaz. 69: 66, fig. 61, 67, fig. 72; 70, ny 42 & 45, & pl. 6, fig. 3. 1920; *Dermen, Gytologia 7: 163, fig. 16. 1936; Gates, Weeds Kans. 91, fig. 19, pl. 6, fig- 9 i9h1; Seed Trade Buyers Guide 86. 1915; Fern. in A. Gray, Man Bot., ed. 8, fig. 1166. ata Moldenke in Gleason, New Britton & Br. Illustr. rie, print. ’ 132, 1952; Hylander, Mscmil. ol Flow. Book p ey Fs {in color]. 195); W. D. Barkley _ da 19574 Strausbaugh & = — Fl. W. Va. 3: tt. i958), Moldenke in Gleason, New Britto r. Illustr. Fl., print. 2, 132. “1958; H. V. Sm., Mich. midis. pl. 155. 1961; soayetas) Fle Mo. pl. 302, fig. 4. 1963. Downy o: 196) Moldenke, Monograph of Verbena 195 throughout; —o pstmt elongate, often extending 1 m. or ovm int e ground; leaves decussate-opposite or approxi- fe) 1 fr ones often hastate-auriculate, sharply serrate or biserrate to incised or coarsely and acutely incised-serrate along the margins, hirsute and rugose above, strigose-hirsute or densely hirsute~- villous beneath, prominently reticulate-venose beneath, the re- ticulation hirsute; inflorescence spicate, terminal; spikes long and thick, solitary or frequently clustered or corymbose-fascicu- late, 30 cm. 2 or papel Be neulate, eylindris, strict, stout, compact, usually densely-flowe atrtdpabchireiee t thrvetent: "the flowers imbricate; peatee lan- ceolate-subulate or linear-setaceo ae approxima ately a s long as the mn violet, about twice as long as the calyx, its tube protruding slightly beyond the calyx, villous in the throat, pubescent or strigose-hirsute on the outside, the limb 8--9 mm t ee Gad and fruit c rowded, igiteysars the heeds “prowish, elon- in that city. The fact that the species is based on a cultivated plant indicates that it has been in cultivation at least as long ago as 1800. The type of V. alopecurus is a specimen cultivated in the Horto Botanico at Madrid, Spain, and deposited in the her- barium of the Jardin Botanico in that city, a phototype being de- posited in the herbarium of the Missouri Botanical Garden at St. Louis. The type of V. mollis is said by Merrill (1949) to be the "Arkansas Rivert", that of V. ringens from "Kentucky", and that of V. cuneifolia from "Pa, Md., & Va." The name, V. hastata stricta also xV. moechina Moldenke and xV. rydbergii rgii Moldenke; the V. sco- Paris of Hooker and of Gillies & Hooker are Diostea scoparia (Gi1I. & Hook. ) Miers. The name V. quintus was proposed in jest - D. Barkley, illustrated by dorsal and ventral views of coced of V. stricta with insect parts added and the legend "A rare prize is the verbena hardback, Verbena Py the male of which is the only known creature having an number of legs." 196 PHYTOLOGIA Vol. 11, no. 3 The "Verbena foliis ovato-oblongis, crenato-serratis ramisque pilosis" which Poiret accredits to Vahl, Enum. Pl. 1: 207, does not occur as a Verbena name in Vahl's book, but with "Stachytar- St a" as the first word is Vahl's description of his perfectly Stachytarpheta strigosa Vahl. Why Poiret shifted it to Ver- T, is not clear to me since he does not give Stachytarpheta strigosa as another synonym of Verbena stricta. Poindexter (1962), in his comparison of Vv. of V. stricta with V. has- tata, xV. rydbergii, and xV. illicita, describes Vv. stricta a as ae a leaf-index of 1.3--2.1 ci lederaca 1.8); leaf-shape ellip- pe to ovate; leaf-base broadly attenuate to sessile; leaf-apex rounded; stem pubescence hirsute; nutlet length 2.2--2.9 pp ge 2.6); markings on back of mtlet deeply ribbed; pollen fertility 50--100 percent (average 90 percent); corolla-tube length .5--5.5 (average .9); calyx length 2.8--5.1 (average 4.3); and petiole lateral vein peasurenen’ 3--6 (average 3.8). His data are apparently based, in part, at least, on the follow- ing of his collections, for all of which ao ept the second) he records a chromosome number of n= 7: P. pelinicter 3) 166-17 (pol- len f es 90 soit P “p01-hé whe peso) a as 7-27 (98 per- st -6 (98 percent), and 229-20 (73 percent). It shoul be noted me cate Schwencke (1931) eee oe haploid chromo- some number as 6, Walpers (185) places V. stricta in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Micrant thae, and Secondary Subgroup Holophyllae, with 22 other species. The os has bean collected in meadows and fields, cod fields and dry fields, sandy or grassy fields and clover fields, Psa anal axa” eroded fields, open sandy fields, pastures and or sandy tures, old or weedy pastures, dry o o. pastures, rocky prairie pastures, overgrazed or very T ocky hilltop pastures, dry lowland or dry sho ass prairie rangelan pastures, dry meadows, low or grassy meadows, woods open woods, low oak-hickory woods, ooded pastures, timber, open places and vacant sade road yards and old Amerind graveyards, rail ditches and roadside ditches, — bottoms and piney creek bottoms, draws and dry ponds, cke -elm woods and o pty ge openings of calcareous Taped ands, dry streambeds os andy valleys, moist places, high bottoms, and bottomlands, 1 wooded pond-margins, low sandy waste areas and sandy vall ne aii Collectors have encountered it in aa marshy areas along creeks, prairie draws, br pastures mowed each summer, breaks of prairie with Asclepias, ravines and ty ravines, waste areas in old feed lots, pine forests as well as fields bo paetne’ 6 woods, upland shortgrass and moderately grazed tallgrass ratios, 1964 Moldenke, Monograph of Verbena 197 eroded prairie ravines and low ravines in prairie pastures, grazed pastures on prairies, and the wet alluvium of floodplains. und in ) ground around shipyards, in low open ground, in soil, dry soil along edge of cultivated fields, dry sandy and gravelly soil, sandy or sandy dry soil, dry sterile or waste soil, thin loam soil, clay soil with many rocks, disturbed gravelly or alluvial soil, chalky or red gypsum soil, clay and black clay soil, moist clay or sandy clay soil, hard clayish soil, and Plain- field Fine Sandy Loam soil, in very rolling heavy clay, black ’ clayey and gravelly calcareous loam, moist to dry loam, mesophytic sand loam, beaches and sand mounds, railroad emb , and dry ballast a- jong railroads. They have found it on roadside and rocky bluestem prairies, dry and alluvial prairies, and sandhill prai ; S an livestock in bluegrass prairie pastures, and along streets and sandy roads. Gates (190) says that V. stricta grows in "Dry soils, valleys, ckets, and waste places" throughout Kansas "except southwest corner" and calls it a hemicryptophyte. Also in Kansas, McGregor refers to it as "common in low places in shortgrass pastures", Ne es", "common in low places", "com- sandy prairie ravines"; "abundant in distur meadows} "abundant in low ravines", and "abundant prairie pastures; tmostly in ravines"; "scattered", "scattered in disturbed parts" » "scattered in prairie ravines", and "scattered on prairie hillslopes"; "not common in dry overgrazed prairie pas- 198 PHYTOL OG TEE Vol. 11, no. 3 tures", and "not common in sandy pastures"; "large colonies in ravines in sandy prairie pastures", "large colonies in ro prairie pasture ravines", and tyery large extensive colony in over- grazed bluestem pas ture". Lathrop found it on "roadside bank bordering a prairie pasture with limestone surfacing". Vo e~ scribes it as "prevalent in dry loam, grasslands and roadsides" and "common in and undergrowth", Marsh describes it as "com~ mon r and "common Flint Hills limestone on roc hi prairies Harms avers that it is a ungrazed an- ally mowed prairies and "numerous oe. scattered in s loam". Unge tes itis " ocks". Hul andy refers to it as "sparse in sandy loam ote en uiree while Wagenknecht found it in "upland bluestem pasture near large limestone outcrops} in "disturbed area in slightly overgrazed bluestem prairie", "com— mon in large colonies in weedy overgrazed pasture along riverbanks} d a "common weed along edge of roadside ditches." Horr found it "scattered over [Trego] county in rich loam with fair soil mois- ture", "scattered all over [Meade] county in dry soil", "scattered in dry grassland, dry loam [Norton County]", "common in dry loam of overgrazed pastures [in Russell County]", and "common along roadside and waste land" in Coffey County. In Illinois it is reported by Cranwill as "very common" in Jackson County and by McDonald as "abundant't in Peoria County, while Jones (1945) says of it "roadsides and fields, common. J ept." In Iowa it is described by Pammel as "a common weed the Missouri loess, associated with Gaura coccinea, Chrysopsis villosa, and Heliopsis scabra" and "common in clearings, sandy clay soil associated with Asclepias verticillata, A. tuberosa, Coreopsis palmata, Heliopsis scabra, and Leptilon canadense", and common n weed, s sandy soil and 2 Sea li associated with V. bracte- ata, Plantago preslii, Delphinium penardi, and Viola pedata", W, and oomeson in sandy clay soil associated with Polygala sanguinea, Coreopsis palmata, Lespedeza capitata, Lepachys pinnata, and Par th and groves with Solidago missouriensis and ve bracteata", Hayden describes it as "common olis® Fawcett says that it is found "on barrens and prairies, Ohio “ Dakota, south to Texas and New Mexico", while Muenscher says "Na~ tive from Ontario westward to the Rocky Mountains; sparingly in- troduced in the northeastern states" exas it is found in pastures, "prairies, and waste vagy "dn ay so ing from u and on railro ankanen Om. June to September, mostly in the Blackland Prairies and Plai Country in southeas , from Lipscomb to Red River, Liber~ plains" in that state, while Hennen reports it as scarce sun" and Lynch says "infrequent to frequent in abandoned sterile fields". Clevender describes it as "common" in Platte County, 196), Moldenke, Monograph of Verbena 199 Nebraska, while in Biol. ahetes 9) at is said to be an "inva- der of degenerate prairi as » and Harms reports a "large colony in prairie " Groh (1944) says of V. strictat "Perennial. A weed in dry or gravelly soil" in Canada; Dodge s says beet abundant in dry open ground" at Point Edward, Ontario, in 1911. In South Dakota it was found by Hayward along "streamsides with Salix, Quercus, Populus" and Harms describes it as "frequent in black loam of lightly grazed prairie" and "common in gravelly sandy loam on er » While Robbins found it in "desiccated depressions along val ditatt” rights~of-way". Ramaley calls it "ruderal" in Colorado, while Weber & Anderson refer to it there as a "weed along road- side® In aosania it encountered by Ugent "in dry sandy soil with Opuntia", Arnot : eae it "in open dry roadside in former Me dahaee prairie", Iltis & Neess call it "a weed at edge of i an no c discovered it on a "very steep dry south- and southwest-facing Bouteloua-Andropogon goat prairie", while Iltis, Bell, Melchert, surfacing in oak-hickory woods", and by McGregor as "common in rocky open areas", Ewan found it in "rather dee uvium or river benchlands, full sun, among bushes back from the shore" nachino (1960) records the species from Bronx County, New Yorks; Tatnall says that it is "well established in waste ground ong rivers" in Delaware, while Earle found it in "waste ground around shipyard" in that state. Small s.n. [Sept. 25, 1691] ties ara £ County, Pennsylvania, bears a notation "in Oy E. J. Palmer (1916) records his nos. 199, 675, and 2980 from Jasper County, Missouri. In his 1935 publication he states that the species inhabits "Prairies, thickets, glades, and was ground, icone utral., General." Thorne (1955) calls it an "A- bundant weed Bad overgrazed pastures, railroads, and other open Places" in Iowa. Duncan (1950, 1953) "peoceds it from Douglas 200 PHYTOLOGIA Vol. 11, no. 3 County, Georgia. Lehr (1962) ue it from Rockland County, New York, where he notes that its predominant associates are Lo- belia spicata and Rudbeckia serot Weber records his no. 172k from Columbia County in the same state. Deam, Yuncker, & & Friesner their 1947 work they record it from Huntingto: unty; in 1950 publication, from Orange County; and, with. Kricbel Cu5hs), c T County, Iowa, record g for this species by me in my Résumé, page 18 (1959) is an error for Porter County, Indiana. Jones (1942) cites R. E. Shanks 5.ne from Wood County, Ohio, while Braun ied records the species from Jessamine and Tri unties, Kentucky. Steyermark (1963) records Verbena stricta from Andrew, Atchi- son, Bollinger, Buchanan, Butler ler, Caldwell, | ll, Cape Girardeau, Car- roll, Carter, Cedar, Christian, Clark, Cole, Cooper, Dade, Dallas, DeK alb, Daviess, Dent, Douglas, Dunklin, Gasconade, Gentry, Harri- son, Holt, Howard, owell, Jefferson, Johnson, Laclede, Lafayette, Lawrence, "Lewis, , Livingst , Madison, McDonald, Mercer, Mississippi, Moniteau, Monroe, 5 Paras New Madrid, Newton, Oregon, Osage, Perry, Polk, Pulaski, Ray, Ripley, Saint Clair, Saint Francis, Sainte Genevieve, Saline, Schuyler, Scott, Shannon, Shelby, Stoddard, Sullivan, Texas, Vernon, Washington, & Webster. Counties, Missouri, on the basis of specimens preserved in the herbarium of the Missouri Botanical Garden at St. Louis, and in Washington and comb The specific name is sometimes up~ pernenes for no valid reason. The plant has been collected in January and in May to October, and in fruit from M to “October and in B December, at altitudes of 200 to 7300 feet, usually in full sunl . Material of oe. stricta has been misidentified and slr uted in herbaria under the names V. bractiosa Michx., V. © canaden- sis Britton, V. diffusa Desf., V. hastata L., V. hastata x stricts Gates, v. lasiostachys Link, V V. macdouglalii Heller, V. officina~ lis L., a prostrata R. Br., and xV. rydbergii Moldenke, as well as Phyllanthus carolinensis hanes (Euphorbiaceae), Stac Stachys tenui- folia Willd. (Lamiaceae), Trian ema portulacastrum L. /L. (Tetragoni- aceae), Veronica arvensis L. = V. prostrata R. Br. ( Scrophulari- aceae). On the other ae the Over 11306 and Shear 149, distributed 58 V. stricta, are actually f. albiflora Wadmond; T. S. Brandegee S- ne (Providence nies May 26, 1902] is V. gooddingii Briq.; S. Wil wil- liams 96 is V. halei small; B. L. Wagenknecht 1927 is V. hastata Lez Rickett, $.n¢ [Rockbridge, July 9, 1927] 4s xV xV. illicita Mol- 196h, Moldenke, Monograph of Verbena 201 ee nyo Sorell oneal cenemmmneenemeetiianineneemnent macdougalii Heller; Demaree 29179 & 30963 and farce Son. {June 16, 1901] are xV. moechina Moldenke; Toumey sn. [Sept. 17, 189k] is V. robusta Greene; Glatfelter sen. [St. Louis, 6-3-92], Somes 370k, and B. L. Wagenknecht 3788 are xV. rydbergii Moldenke; F. C. Gates 3519 and H. W. Houghton 032 are V. urticifolia L.; C. C. Albers 33013, Condit s.n. (Houston, June 19, 1909], ae iv, and H. Ne H. Ness s.n. [June 26, 1925] are V. xutha Lelm.; A. W. C 8.n. (185) and | R. K. Godfrey 4239 are Stylodon carneus Wieden sae Moldenke; rag Porter r 5150 is Salvia sylvestris L. » (Lamiac eae). Cing~Mars s.n. [31 juillet 1951], A. S. Hitchcock s.n. [Ames, July '82), Pe Rydberg 122 are mixtures with f. albiflora Wadmond; Kraeger 1,75 is a mixture with V. hastata var. scabra Moldenke; iemetes 29151 and L. H. Pammel s.n. (Hamilton, Sept. 8, 1918] & S.n. (Granite, Sept. 1, 1920] are mixtures with xV. rydbergii Mol- denke; Z, Baldwin Bebe is a mixture with V. simplex Lelm.; and Schneck sen. [Sept. 1, 1887] is a mixture with V. urticifolia L. es G. E. Morley ry & 396, distributed as V. stricta with the omment ‘a very light color" of flower, are cited by me as f. rosei flora Benke, but may actually represent another soinontbes color form corresponding to the f. caerulea Moldenke of V. hasta- L. Miss Gaiser says of her 1382aPE "scattered plants", of 1759PE "this population seemed to have generally less deeply lob leaves", and of 2306PE "no hint of hybridization, just typical stricta in not sand hills along the beginning of lake toward Light Hous Shhesk sen. [July 25, 1897] has a spike fasciated at the a- pex; R. E. Shanks 1790 has rather slender spikes; C. R- Ball 2439 has two binary leaves; E. W. Fell 51309 has its leaves § unusually narrow; E. Hall s.n. [Athens, Aug. 1063) bears the EEE Pe no- tation. "unadulterated". Demaree 30203 and F. J. Hermann 9159 are abno normal, somewhat resembling xV xV. rydbergii, but a the spikes heavily fruited so I am regarding then as V. stricta. Shimek s- ne. (Oct. 14, 1919] was originally identified as "V. stricta x angustifolia"; A. E. Allen s.n. ae July 11) 1591], A. S. Hitchcock s.n. [Medicine L hag y 1892], and J. L. Sheldon 3.n. (Peru, July 21, 1900] were tia "Ve stricta x x hastata"; W. fo He Horr E.33 was distributed as "V. hastata x stricta"; L. H. Pan mel SN. [Comanche, Sept. 13, 1918] bear: bears the ambiguous identifi- cation "V. stricta x spicata"; and Loomis s.n. [Alton, Oct. 1938], Muck 82, and S. V. Fraser tho were cited as "V. stricts x hastata" ing Gates, Kans. Fl. — 1: regard all of these collections as s typical Verbena stricta, as also R. Bebb 14376, eaptiatiy thought by the 202 PHYTOLOGIA Vol. 11, no. 3 collector to represent a cross with V. simplex Lehm.; H. N. Pat- terson s.n. [vicinity of Oquawka), labeled led "Verbena stricta x annotated by Pe erry as "possibly a rica oe meee On the other hand, Evers 198 may b mg ton "since the tips of some of its spikes have the i emis austen, Deane s.n. [Med- ford, Nov. 13, 1887] consists of seeds only; S. F ", Norton s.n. [June 27, 19h] bears a notation averring that Pit was collected "at the exact center of continental U.S.A." The Herb. Calif. A~ cad. Sci. 31415 does not bear any indication on its label el that it came from cultivated material, but I am assuming that it did. L. H. Pammel s.n. [Waseka, Aug. 1920] is marked "Wisc.* on its label, but this — locality is actually in Minnesota. In my 1940 work I followed Perry in regarding V. stricta f. ae Ace Wadmond and f. roseiflora Benke as synonymous ious with Vv. stricta, but I have since decided that these represent distinct and nomenclaturally worthy color forms. The C. M. Rowell 4lls, cited below, s Me ag by the collector -“ ferieg had the ¢ cor=- ollas "lavend o pale purple" and so may possibly rep erg ts f. roseiflora. NeHitbeae Dixit, & Romberg 11689. is pei rf having wers white and lavender" and is cited by me sande f. albiflora. "ha corolla is described as "deep-blue" on Chamney i, "blue" on Horr 4548, 4662, & 4767, "dark-blue" on McGregor 5%7, "violet" on Marsh 1735, and ad “"bright-violet" on Marsh 1613. Bergen ith2) states that V. stricta is thought to be be specific ag. ever and ague — hence the common name of "fever-weed". H. V. Smith (1961) asus that yt jase we rmarkable resis- tance to drought and "The foliage is so bitter that cattle will not eat it even when forage is scarce, The root system extends than a yard in dry soil, open sandy ground, in barren seeds are a to the horticultural trade from Rex D. Pearce The following fungi are known to attack V. stricta: Aecidium abe Auct. Amer. (non Speg.), A. verbenicola Ellis & Kellem., Cercospora verbenae-strictae Peck, es verbenicola Ellis & Eve, Dicaeoma verbenicola Arth., Erysiphe horridula Lév., E. lampro~ carpa Auct. Amer. (in part), Phyllosticta verbenicola Martin, Puccinia sydowiana Diet., P. verbenicola Arth., P. vilfae Arth. & Holw., Septoria verbenae ROb., and Stagonospora verbenae Greene- uitcheli (1360) and my son, Andrew Ralph Moldenke, report that the flowers of V. stricta are "regularly visited by the bee, Calli- opsis sitindisaactha Cant ents the female taking pollen and nectar, the male taking only n and wecieeniac nanes recorded for V. stricta are ‘blue ee neem mec common verbena", "blue vervain", "blue vervian", "bur-vine™, "common ver~ 196) Moldenke, Monograph of Verbena 203 vain", a eBig “hairy v beeing "hoary-leaved vervain", erbena it; "bur-vine" is applied also to V. simplex and V. urticifolia; "blue verbena", "blue vervain", and "wild hyssop" are applied al- so to V. has hastata; "common vervain" is applied also to V. officin- ais and V. urti urticifolia; and "thimble-weed" is applied Y also to Anemone virginiana L. (Ranunculaceae), es purpureum (Vent .) we (Fabaceae), and Rudbeckia laciniata + (oannensee) Verben stricta is knom to hybridize freely -- the hybr hybrid with V. mactesie® Lag. & Rodr, is xV. deamii mT that with y. halei Small is xV. goodmani Moldenke, @, that with V. hastata L. is xv. rydbergii Moldenke, that with V. simplex Lehm. is xV. moe- china M Moldenke, and that with v. urticifolia L. is xV- illicita Netiorice separ Ade orge J. Go in a letter to me dated October 24, 1960, ristel that Bw. P Fileer > 361 Sacer "2,3" by Torrey } was collected in Johnson County, Oklahoma, on June 25, 1868, and not nA "the False Washita" or "Fort Arbuckle" as stated on the la- Hooker (1836) cites T. Drummond s.n. [St. Louis] in the eal rbariu. Schauer (187) cites hae ehi 196 and Engelmann from "In Am the DeCandolle Herbarium at Geneva, not as yet seen by me. Groh (1944) cites R. Cameron s.n. [Stamford, Welland Co., 1898], Dore & Groh 1805, Dore s.n. [N. of North Rideau, Carleton Co., 1943], Fisher s.n. [Sarnia, Lambton Co., 1902], Dent s.n. (Sarnia, Lamb- ton Co Co., 1909], Montgomery 709 {Waterloo Co.], Owens s.n. [Toron- to, York Co., 19h], Armstrong s.n. [Jermyn, Peterborough Co., 1903), and Scott s.n. [St. David's, Lincoln Co., 1898] from Ontar- » and Dore s.n, [sandy pasture field, ee Lake, Pontiac Co., 950} from Quebec, not as yet seen by m Perry (1933) rae te f. roseiflora Pediké and f. albiflora Wad- mond as synonymas with V. stricta, but I regard them as distinct. She gives the distribution of the species as "eastern bee —— United Stites s, from Pennsylvania westward through the Rocky Moun tains. ably oe into other localities." she cites the following 120'8 pecimens and 1 photograph not as yet seen by me: MASSACHUSETTS: - Bristol Co.: Sanford s.n. (Fall River, "2 Sept. 1903] (G). CONNECTICUT: Fairfield Co.: E. H. Eames s.n. [Bridgeport, 27 Aug. 1892] (G). New Haven Co.: Blewitt 1] (G). NEW YORK: Tomp 20h, PHYTOLOGIA Vol. 11, no. 3 Co.: A» J. Eames 12797 (G). PENNSYLVANIA: Lancaster Co.: Urban sen. [along Lincoln Highway at Gap] (G). OHIO: Montgomery Co.: Short s.n. [Dayton] (E). ILLINOIS: Hancock Co.: F,. C. Gates 9994 (&). Henderson Co.: H. N. Patterson s.n. (Oquawka, 1872] (Babe Jackson Co.: H. A. Gleason sen. [Grand Tower, 22 Aug. 1 1900] (G). LaSalle Co.: Thone 88 (E). “Macon Co.: H. A. Gleason 377 (G). Mad- ison Co.: Sherff s sen. {Mississippi River, bluffs north of Alton, Aug. 1910] (G). McLean Co.: Robinson s.n. (Bloomington, July 1886] (G). Menard Co.: E. Hall s.n. [Athens, Aug. 1863] (BE). Pike Cos: J. Davis 16 (E), 6370 (E). Richland Co.: Ridgway 2h31 (£)- Saint Clair Co.: Eggert s.n. [E. St. Louis, 28 July 1900] 0] (B). Will Co.: Umbach s.n. [Romeo, 26 July 1897] (E). Stony Island: H. H. Smith 5645 (E, G), 5952 (G), 6028 (E, G). INDIANA: Harrison Co.: C. C. Deam 20357a (G). Kosciusko Co.: C. C. Deam 5.n. (E); L. H. Pammel 85 (E, G). me Co.: Ball 1585 (E). Van Buren Co.: Graves 197 (E), 199k (E). KENTUCKY: Ballard Co.: McFarland & Anderson 253 (E). MICHIGAN: Saint Joseph Co.: G. L. Fisher 19 (E). WISCONSIN: Dane Co.: Trelease s.n. (Oregon road, Madison, ae, July 1839] (E). Trempeleau Co.: Hale son. [Trempeleau, 1861] (G)- Waupaca Co.: Garesche s.n. [Waupaca, 1907] (E). County undeter- mined: Eggert s.n. [Mirror Lake, 15 July 1903] (E). MIN NNESOTA Hennepin Co.: Schuette s.n. [St. Anthony, July 7, 1888] (G)- Hous- ton Co.: Freiberg s.n. [July 1912] (E). Kandiyohi Co.: We De Frost s.n. (Willmar, July 1892] (G). Ottertail Co.: Chandonnet sn. (Perham, August 8, 1912] (£). Wabasha Co.: Manning S$ -De [Lake City, 28 July 1883] (G). SOUTH DAKOTA: Brookings Co.: John- son s.n. (Windsor Township, 27 July 1903] (B). Jackson Co.: E. Eo Je Palmer 37627 (E, G). Lawrence Co.: W. P. Carr 118 (B, G)- Roberts Co.: Over 14386 (W). Sanborn Co.: Visher 450 (E). KANSAS: Hamil- ton Co.: C. H. Thompson 154 (E, G). ~~ Vontgomery Co.: Rydberg & In- ler 433 (E). Osborne Co.: | Shear 191 (G). Pottawatomie ie CO. Ae Ss. Hitchcock 972 (W). Riley Co.: J. B. S. Norton 391 (E, G)- MIss= OURI: Benton Co.: Trelease 717 (E). Camden Co.: Bmig 107 (5). Clay Co.: K. K. Mackenzie 261 (BE). Jackson Co.: B. F. Bush 475 (E). Jasper Co.: E. J. Palmer 199 (E), 2980 (E); Trelease ase 718 (E)- Lincoln Co.: Beckwith 8 (E). Marion Co.: J. Davis 1202 (E), yy, 2523 (E), 2791 (E), 2960 (E), 2962 @), 3227 (£), 3589 (E), 462i (Bhp 4h87 (E). Phelps Co.: Kellogg 198 (E). Saint Louis Co.: . Letter- man s.n. [Allenton](E); "stiaid 32 (E). Taney Co.: Be Fe ". Bush 570 (E). Wayne Co.: E. J. Palm r 6107 (E). Wright Co.: Lansing 3164 (G). Saint Louis: Rctieaha! ae on. (St. Louis, Aug. 18¥1] (G)5 Riehl 196 (E). ARKANSAS: Craighead Co.: Demaree 3553 (E). Wash- 196) Moldenke, Monograph of Verbena 205 ington Co.: F. L. Harvey 61 (E). County undetermined: F. L. Har- vey s.n. [Curtiss 1958] (G). MONTANA: Big Horn Co.: Blankinship S.n. [Crow Agency, July 1k, 1901] (G). WYOMING: Laramie Co.: A. Nelson 538 (G). Platte Co.: A. Nelson 505 (E, G). COLORADO: Denver Co.: Eastwood 90 (E, G). Yuma Co.: Eggleston 1522) (E). NEBRASKA: Custer Co.: Bates san. (Callaway, 27 June 1901] (G); L. H. Pammel s.n. (Broken Bow, 7 July 1897] (E). Harlan Co.: L. Ber Pammel s.n. {Alma, June 21, 1897] (E). Lancaster Co.: Webber as n. [Lincoln, 27 June 1885). (E). Lincoln Co.: H. Enge elmann 5.1 eg {South Fork of the Platte, July 1856] (E, G). Merrick Co.: A. A. Heller 14290 (E). Thomas Co.: Rydberg 122, in part [Dismal Riv- er south of Thedford, 27 June 1893] B93] (@).. 0 OKLAHOMA: Choctaw Co.: H. W. Houghton s.n, iste vens 4000] (E, G). Cleveland Co.: Emig 365 (E). Comanche Co.: Me Ke aan "11748 (E); G. W. Stevens 1354 1/2 (E, G). Ellis Co.: G. W. Stevens 2930 (G). Greer Co.: G. W W. Stevens 1034 (G). Harper r Co.: Stratton on 403 (EZ). Kay Co.: Ge We Stevens 1865 (G). Murray Co.: Eaig 787 (E). Pawnee Co.: Stratton 249 (E). Rogers Co.: Be F. Bush 135 (E). TEXAS: Dallas Co.: J. Reverchon 735 (E), Sone ene (G). Hemphill Co.: Eg- gert S.n. {near Canadian, 11 Aug. 1900] (£). NEW MEXICO: Santa Fe Cow: F Fendler 597, in part Lea ee a). WASHINGTON: Stevens Co.: Kreager 75, in part (G). CULTIVATED: Spain: Herb. Bot. Gard. Madrid s.n. ihaidt The Bain 328, Bush 475, Rydberg 122, and Sm Small s,n. which she cites from nek are E stagse fies in the Columbia University ry: what s as "Harvey ae from "NY" is herein cited as Harvey Hee godt 1958] in th peapeN a University herba ree her Reverchon s.n. from Dal ates t "NY" is probably the J. Reverchon s.n. (Dallas, May 1877] in the Barnard College herbarium; her Jones s.n. from Grinn "Aug. = taal ie probably the "Aug. 8, 1877" collection cited be- low; and what she cites as "Houghto n’ 4,000" is listed by me as Rouaiities s.n. [Stevens )\000]. Of Ley eee she says "A distinct species somewhat incapable | sharp delimitation 2 sg ae- In all, 1916 herbarium specimens and 2 mounted illustrations have been examined by sce Citations: QUEBEC: Deux-Montagnes Co.: Cinq-Mars s.n- Huron Gout Marie-Victorin, Rolland-ernain, & Dominique 45993 (Um—207). “Lambton Mal J. Dearness s-n. [Pt. Edward, 13th Aug- ust 1902} (Mg); C « Dodge Sone Sen. [Po Point nt Edward, July 27, 1902] (Mi, Mi), s.n. core (Ob-—-87659), son. [Point Edward, July ll, eee 206 PHYTOLOGIA Vol. 11, no. 3 1911] (Mi), s.n. [Point Edward, 8/27/40] (Au, Au); G. L. Fisher sen. [Sarnia, Aug. 5, 1902] (B); Gaiser 1382aPE (Gp, Gp, Gp, Mm), 1618PE (Gp, Gp, Gp, Mm), 1759PE (Gp, Gp, Gp, Mm), 2178PE (Gp, Gp, Gp, Mm), 2306PE (Gp, Gp, Gp, Mm); Gaiser & Montgomery 2234PE (Gp); Gaiser & Slobasky 1382 (Gp, Gp, Gp, Gp, ,» Gp, Mm). Middlesex Co.: Col- grove s.n. 1. [Strathroy, 31 Aug. 193] (Mg). MASSACHUSETTS: Middle- sex Co.: Deane sn. [Medford, Nov. 13, 1887] (Ba). CONNECTICUT: Fairfield Co.: E. H H. Eames s.n. [Bridgeport, VIII .27.1892] (N, ¥— 283608) . Litchfield Cow: Ee He Eames 10221 (In). NEW YORK: Al- bany Co.: H. D. House 18369 (Al). Cattaraugus Co.: M. Zimmerman son. [Red House, Aug. 8, 1935] (Ba). Columbia Co.: W. A. Weber —_—_——— Suffolk Co.: Disbrow s.n. [E. Hamp., 1894] (Nm). Tompkins Co.: A. J. Eames 12797 (S); Muenscher & Bechtel 163 (Ba, Gg—31h21, To—112276, Pl--63267). Ulster Co.: Elting ; 2073 (Al). Warren Co.: H. D. House 28901 (Al). Washington Co.: + Carpenter s.n. [Gra ville, Aug. 1911] (Vt); Pember sn, (Granville, Aug. hl, 3913) (Vt), s.n. [23 Aug. 1911] (Al). NEW JERSEY: Hudson Co.: Van Sick- le s.n. [New Durham, July 15, 1893] (Tc, W--3096)8), s.n. [Gran ton, Sept. 21, 1895] (W—21,3275) « Morris Co.: Neneste ne vino 462 (Ec, Jr, M1, N), s.n. [Budd Lake, Sept. 17, 1939] (N)- PENNSYLVANIA: Allegheny Co.: Bright s.n. [Aug. 27, 1922] (Cm, eas B. H. Pat terson s.n. [Sept. 2, 1916] (Cm, Cm, Cm). Lancaster Co.: Ae ee Heller 638 (W), s.n. "{coaewago, Sept. 27, 1901] (-L07055)4 ge K. Small 106 (0b--508)0), s.n. [vicinity of Dillerville, July 2, 1890] (Up—17126, W--29865]), s.n. [Conewago, August 30, 1890] pec tL sen. (Sept. 25, 1891] (C), s.n. [vicinity of Cone- ,» September 1892] (C). Philadelphia Co.: Eckfeldt s.n [June 20, tor) (Up). Westmoreland Co.: Shafer s.n. [July 19, y, 1900) (Cm), son. [July 20, 1900] (Cm). Presque Isle: Ammons, Straus- baugh, & Bartholonew s.n. [Presque Isle, July 29, 5» 1940) (We). DELAWARE: New Castle Co.: E. Tatnall s.n. [July 1881] (Pa). Sus- sex Co.: E. C. Earle 1781 (Ca--6869, Up); J. H. Holmes 2k2 1/2 (Ka); Re R. Tatnall 3122 (Up). WEST VIRGINIA: Pocahontas Co.! Herb, West Va. Univ. Bot. Exped. 1081 (We, We, We). NORTH CARO- LINA: Onslow Cos: Ahles 599 (Ur). ALABAMA: Cullman Cost Cc. Mohr s.n. (Cullman, May 29, 1892] (W--771847). OHIO: Butler Cost “w1- ier & & Roth sone [July 15, 1932] (Ob--80600). Champaign Co.: ee raper A.157 (Herb. Leonard 6100] (W--2161893); E. C. Leonard T7267 (W--1601267). Clark Co.: aoe k 36-709 a be —-1,7803)- Hamil~ ton Co.: Aiken 10691 (11), s.n. [Terrace Park, VII-25-1905] (Po- 267631); E. L. Braun s.n. [7/25/05] (Cn, Cn), s.n. (Terrace Park, Aug. 9, '17] (cn), s.n. [Terrace Park, Aug. 19] (Gn); J. Clark 8 TT 196), Moldenke, Monograph of Verbena 207 n. [Cincinnati] (Bm); Collector undesignated s.n. (Cn); C. Herrick sen. (Aug. 2h , 1889] (Ob—80601); M. Mohr 167 lg “ia Cow: He H. C. Beardslee s.n. [June 2nd, 1870] (ob--80599) . Montgom- ery Co.: “G. S. Grave: S. Graves s.n. [12 Segiat 1880] (Al); E. C. Leonard 2440 {Herb. Leonard 4382] (W-~2161775); Leonard & Mannakee 5552 [Herb. Leonard 393] (W--2161861); H. P. Smith 1958 (Dt). Sandus- ky Co.: Moseley sen. [Gibsonburg, Aug. 9 9, ~1920) (Mi). Union Cos: A. R. Moldenke Tb (. (Lw). Wood Co.: Moseley sn. [Bowling Green, June 12, 1921] (Mi, Mi); R. E. Shanks 1790 (N). County undete mined: rs Le ones son. (Du--95h); Jewett s.n. [July 12, 182] Mi); '» Lloyd yen. (Ll). ILLINOIS: Adams Co.: Brinker 1222 ae es 2691 ee 2729 (I1—-15907); E. W. Erlanson 18 (Mi); Evers 197 (T1—25583, N), 198 (ur), 1Uuk6 (Ur); Evers, Jones, & Jones 646 (Ur); A. B. Seymour . (Camp Point, 1 Aug. iets (H--106L6h) , sen. (Adams Co., 387 2] (H-~106465) « Bureau Co.: Congdon s.n. [Princeton, Oct. 1856] (Du--9515); Winterringer 1395 (I1--37248). Carroll Co.: ahles ),312 (Ur); Collector undes- ignated s.n, (I1—-15871); G. D. Fuller 1747 (I1--15802); Winter- niger ringer 81L5 (11-3737). Cass Co.: Ahles 299 (Ur); G. D. Fuller 11855 (I1--2)789); F. C. Gates 551 (iti); vinterringer 307 7 (1— 35517). Champaign Co.: M. H. Bell 161 (Ur); L. J. Blake s.n. (Urbana, 7-15-51] (Ur); R. Crane ue 73 (N); Evers 185 85 (Ur), ), 1487 (Ur); H. A. Gleason s.n. (Champaign, July 31, 1900] (In); F. We Johnson s.n. [Longwood] (N); G. Jones 12436 (Au, Ur), 12523, (I1—-15808, N, Ur), 12525 (N), 12667 (I1--15807, N, Ur), 1 15265 (Ur); Piaibercen sen. [duly 2, 1942] (Ur); Seymour & Waite s.n. (Ur--17],65); Stubbs 7 (We); Waite s.n. (Ur—-29662) ; "Winterringer LOL, (I1l—29329). Christian Co.: De Mott s.n. (Taylorville, 8/ 157"96] (Ur); Shouse s.n. [July 21, 1950] (Ur). Coles Co.: V. A. Anderson 1259, (I1--20713); G. D. Fuller 10643 (I1--21731, » il 21732). Co Cook Co. : He H. Babcock k 9283 (Bz--23790), sen. (Hyde Park, July 3, 1811] (N), 5 sn. (Chicago, July 3, 1871] (W--310735), son. (July 2, 187k] (Sg—-16102, W--71941), son. [Chicago] (Al); L. He Bailey s.n. (Chicago, July 27, 1880] (Ba); S. A. Cain sen. [aug 1926] (Bt—-2h6h), sen. [southern Chicago, 1926] ) (Bt--2h:73)$ Me Ae Chase s.n. (Cicero, July 20, 1896} (Ur, Ur); M. H. Clark s.n- [Lincoln Park, July 169] (uti); Davis & Davis 1635 (We); . Dixon & Gage 752 (W-—609),3h) 5 H. S. Fawcett san. [Berwyn, 8-25-03] (Io-- 7368L); G. De Fuller 1312 (11--15793); Gilman s.n- (Chicago, July '69] (Pr); Roman s.n. [13/9/1893] (5), Sens Sen. (Aug. 1693] (S); Ship- man 1358 (Mi); Umbach 591) (Ca—W1527 62531); Wessén 737 (Go, "S), Sen. [Jackson Par! Park, VII.1899] pe DeKalb Co. P. Be Whitford s.n. [July 19, 196] (Ur). DeWitt Co.: Winterringer 6766 T1~34),02). Douglas Co.: Winterringer 612 (Ur). Fulton Co.! 208 PHYTOLOGIA Vol. 11, no. 3 Winterringer 8832 (I1--38672). Hancock Co.: F.C. Gates 999 (Ur); Kirchherr 66 (I1—15917); Kirchherr & Fuller 46 (11--15803) 3 Mead sen. [August 18,5] (N); L. H. Pammel $.n. [Hamilton, Sept. 8, 1918) (Io—9)058). Henderson Co.: H. N. Patterson s.n. [July 1872] (Al), sen. (near Oquawka, August 1972] (N), sen. [August 14, 1873] (Al), sen. [Oquawka] (Cm), sen. [vicinity of Oquawka] (W--132312h, W--1323125). Henry Co.: Stark son. (Orion, July 16/ 23] (Dp). Jackson Co.: Cranwill s.n. [July 17, 1917] (Ur); a & Welch 915 (I1—15893); J. McCree e 915 (Ur). Jasper Coe: Shearer sn. n. [9/1/1928] (Ur). Jersey sy COs? P. P. Jacobs 68 (Ur). Jo Se Co.: Ahles 363 (Ur); Winterringer 8148 (T1—37434). Kane Co.? Pearsall 7279 (I1--15806). Kankakee Co.: Collector undesignated sen. [July 8, 1870] (Ur, Ur). Lake Co.: Earll s.n, [Waukegan, June 1878] (W--L91s2) ; F. C. Gates 3160 (Ur). LaSalle Co.: G D. Fuller 2299 (I1—15798); H. K. Henry 122 (Up); McDougall 5 (Ur); Thone 88 (Ur). Lawrence Co.: Sivert s.n. aes 23, 1949] “(Il 31977), s.n. [June 26, 1952] (Ur). Lee Co.: G, D. Fuller 11928 (I1-~-18896) ; Keithley. 129 (I1--15790), sn. eae 29, 1943] T (Ur); an s.n. [Amboy, 1696] (Dt); Mertz 251 (Cm Cm), son. [Dixon, Aug. '77] (Be), sen. (Dixon, Aug. 15, 1877] (N). Livingston Co. Ge We Fuller 919 (11-1591); H. N. Moldenke 7607 (N). Logan Co.: Ahles LOL (Ur); G. Randolph s-n. [July 17, 1959] preg et con Cos: H. A. Gleason s.n. [Je. 19, 1896] (Ur); H. Ne 21909 (Lw, Ut, Z); Winterringer 6502 (11—33798), 6683 ( awe 267) « Macoupin Co.: W. E. Andrews s.n. [June 20, 1889] (Ur); G. D. Ful- ler 22h6 (I1--15799). Madison Co.: Norwood son. [July 4 T9uB) (Ur). Marion Co.: M. S. Bebb s.n. [Salem] (T). Mason 60.8 05 De Fuller 223 (I1--15895); H. A. Gleason 9223 (N); Ge Ne Jones 22383 (Ur); Winterringer 61 (Ur), 51h (Ur). Massac [& Pop pe] Cost Whit- ing 1725 (T1--15800). McDonough Co.: E. Re Boardman 5.n- (prair- i (Go); C. H. Kauffman 458 (Mi); G. N. Jones 22401 (Ur); Re Me Myers 0.15 (I1—2394L). McHenry Co.: Ee Re R. Bennett s.n. [Augus 10, 1957] ab: Mason s.n. [Algonquin, July 27, 1878] pes — - [Ringwood] (Pa). McLean Co.: Calef 545 (Ur); G. D 7 7 (1115796). Menard Co.: E. Hall s.n. [1861] (Oe) “sane xr Tit Aug. 1863] (Pa). praca Co.: Winterringer 38 9 (I1 29265) Morgan Co.: E. Epling 6123 (La); A» Re Sct as 826 (Lm)5 E. S. Steele s.n. [about 1580 rae bea oultrie Co.: Winter- ringer 4870 (I1—3)908). Ogle Co.: Ahles 4547 (Ur); A. Le Hills 3392-0 (I1--15892, Ur). Peoria Co.: V. H. Chase 4585 (Ur); ee Fisher s.n. [9-3-1830] (Ka); Heading san. [July 1975] (Ur), SeMs Leet soe et (No--2522h); F. E. McDonald s.n, (Peoria, uen 1887] —~59473), sn. (Peoria, J 1901] (Ur), sen. [Peo 1903] (Io—76181, N); Panlbeendbebie 26, aL Piatt Co.! 1964 Moldenke, Monograph of Verbena 209 Ahles 3356 (Ur); F. C. Gates 2125 (W-~69182); G. N. Jones 2081k (tr); Re Pi Pearson s.n. (July 2h, 1949] (Ur); Ae Be Seymour s.n. (Deland, September 1889] (H--10772). Pike Co.: J. Davis 16 (Ur), 3239 (Vi), son. (Shepherd, 25-6-15] (Ur); A. R. Moldenke 835 (LW). Rock Island Co.: E. T. Harper s.n. [Port Byron, 1893] 31893) (Io--133475). Saint Clair Co.: E. W. sae geet (Ur); J, Neill 291 (11—22978). Sangamon Co,: Ahles 4590 (Ur); G. D. Fuller 1366 (1115791), 4194 (I1--15797), 176 (x1—16056) uh99 (T1=-15902), 4590 (I1— 15906), 4613 (I1—15910), 4660 (I1--15911), 4700 (I1—~15885), 7h2 (111590), 4799 (1118886), 4821 (1115782), 4880 (I1— 15912), Los (1115888) , 5024 (11—-15897), 5066 (I1—~15908), 5147 (1115783), 51h4 (15889), 5195 (1115887), 52hh (I1— 15898), 5348 (11--15787), 5470 (I1--15786), 9571 (1115788), 5591 (I1--15918), 5654 (1115909), 5691 (1115780), 5845 (I1—- 15890), 5882 (11~-1578k), 5974 (I1--15785), 6006 (I1—~15899), 6098 (I1-=15891), 6319 (I1--15781), 6385 (I1~15789), 8792 (I1— 15913); L. B. Mead s.n. (Ca--216738); A. R. Moldenke 823 23 (Lw); A. B. Seymour s.n. [Springfield Junction, § July 1879] (F-10646). Scott Co.: A. A. Flynn 14239 (I1—25153); A» Re Moldenke 831 (Lw, Ut). Shelby Co.: G. D. F Fuller 13090 (I1~-23753). Stark Co.1 V. He Chase s.n. [July 10 0, 1896] (Ur (Ur, Ur). Stevenson Co.: Serf £ 9833 (11=-15915), 9869 (T1-=15916). Tasewell Co.: V. H. Chase hi7h (Ur). Union Gow Collector undesignated s.n. fAug. 1, 1879] 9] (Cm); G. D. Fuller 667 (Ur); Fuller & Fisher 667 7 (1115795) « Vermilion Co.: | Storm s.n, [July 28, 1949] (Ur, Ur); )3 Winterringer 6520 (Il 33773). Wabash Co.: Schneck s.n. [Sept. 1, 1887} (Ur), sen oe {July 25, 1897} pees sen, [July Tsuly 1900) (Ur); Shearer S.n. eae armel, July 6, 1900] (Vi). Whiteside Co.: Ahles 4211 21l (Ur); G. D. Fuller 13662 (1131212); Winterringer 8167 (11--3738). Will Co.: Um- bach s.n, (Romeo, Aug. 25, 1897] (MM, W--339480). Winnebago Co.? a S. Bebb 9285 (Bz--23789), s.n, [Fountaindale, 1867] (Cm); E. » Fell 51309 (I1--38606), 51327 (I1—-38588), 51340 (12—-38616) , Sem: (11--38883); Fuller & Haime 15f (I1—-15792), 27h (1— 15 1589h) 5 W. H. Rhoades sn. [Rockford] (Hs); Swezey 5.n. gen. [Pecaton- fica] (Gg--31h16). Woodford Co.: Ve He Chase 9693 (Ur), 9963 (Ur), 9990 (Au--122715, St, Ur, Ur). Stony Island: “H. He Smith h 56k5 (Ca--882826), 6028 (Ca--882825) « County undetermined: H. H. Bab- cock 8.n, [Auge *6 '69] (Br); M. S. Bebb S.ne {Central Illinois, 1867) (Bm (Bm); J. Blake s.n. [Illinois] (S)} Buckley s.n. [1837] (1); Collector undesignated sen. (Gg--31h14); Fink s.n. (Blackberry, July 28, 1892] (1028699); Short s.n. (Dry prairies] (T); J. Tor rey Sen. [Illinois] (Pr); G. R. Vasey 4 (I1—-16006), son. (Ms). INDIANA: Carroll co.: Ce Co! Deam 1535 (In). Clay Co.: W. Me 210 Pet tO OO SS Vol. 11, no. 3 Rhoades s.n. [Spencer, July 1931] (Up). Daviess Co.: Tewell s.n. [Washington, 19 June 1936] (Fl--25879). Elkhart Co.: C. C. Deam 21001 (In); Friesner acer (Bt—5h7h7). Floyd Co.: C. C. Dean 157 7 (In). Fulton c iond Gs C. Deam 42167 (In); Friesner r 13655 (Bt—,6926). Gibson fen Pts eS) “Deam 25506 (In); Schneck s.n. (July 1888] (In). Greene Co.: | C. CG. Deam 25657 (In); Friesner 6513 (Bt—~14217); Kramer & Johnson s.n. [July 16, 1941] (Ok); W. Rhoades s.n. (near Marco, “July 1927] (Bt--2929h,, Ny). Hamilton Co.: Friesner 6043 (Bt--13L55, Ca--540466, Lb--13646, Po--201h15, Sd—23983). Harrison Co.: C. C. Deam 20357a (In), 4198 (In). Hendricks Co.: S. M. Deam 13675 (In). Howard Co.: EK S- sen, [7-9- 1938] (Bt--5673h). Jasper Co.: C. C. Deam 42088 (In); "Friesner 1611 (Bt—-50862); Welch s.n. (In--5538, In--5839). Jefferson Co.t Je M. Coulter 1h (V (Vt). Jennings Co.: C. C» Deam a (In); Hendricks | sen. [August 7, 192) (Hi--24737)~ Knox Gos: ree Deam 29283 (In); Friesner 7923 (Bt--20358, Gg--237861, Soni Se McCoy 630 (Dp--h67b) , 4233 (Lb--3727). Kosciusko Coes Ce Ce Deam S.n. (northeast of Winona Lake, August 2, 1897] (In, W—- 193309); Friesner 15436 (Bt—54877). Lagrange Co.: C. C. Deam 14980 (In), 31309 (In); Friesner 15621 (Bt--51608). Lake Cost Re Bebb 1682 (Ok); F. W. Johnson 348 (N); Lansing 2810 (W--75309h) « LaPorte Co.: C. C. Deam 31426 (in). Lawrence Co.: Kriebel 5.Me [7-3-33] (Bt--19)75); W Weatherwax s.n. (In-~2)0)); Wynn 89 (Au). Co.: H. H. Bartlett s.n. [July 13, 1903] (Mi); C. C. Deam 6957 (In); H. F. Dietz s.n. [Fall, 1910] (Du--9542); Friesner 16956 (Bt-—-622h9, st—-23607, We), s s.n. [Pleasant Run & & Brookville Rd., 71-25] (Bt--2h71). Marshall Co.: C. C. Deam 45070 (In); Ev ermann 970 (W—35780k); W. H. tthekdaa” s.n. [Plymouth] (Hs); Sco- ena Vell & Clark 970 [35] (Du--9538). Martin Co.: C. C. Deam 11383 (In). Miami Cos: Ek s.n. (near Bunker Hill, 8-16-1936] (Bt— 42529). Monroe Co.: V. Davis s.n. [Bloomington] (In~-2287); Lewis & Brown s.n. (Bloomington, Aug. 106) (Au). Montgomery Co.t Ce 2 Co Deam 17621 (In). Newton Co.: C. C. Deam 18108 (In); Friesner 1h703 (Bt--50598, Ky, N, Pl~132318, We); H. Hahn s.n. [Roselamn, 1905] (W-—-609230); M. McKee 10h (Dp). Owen Co.: 3. M. Deam 8952 (In); W. H. Rhoades s son. (Spencer, July 1931) (Hs, YN), 8 EBS yn. [near Spencer, July 1931) 1931} (3 Weatherwax s.n, [Spencer] (1In—10h2) « Parke Co.: Daubermire s.n. [7/7/30] (Bt—-6068); Disbrow s.n. [Sil- verwood, Sep. '03] (Nm); W. H. Duncan lgh (H—-2494). Pike ke Cows Ce C. Deam "16965 (In). Porter Co.: C. C. Deam 31560 (In); F. We Johnson 38 ( (N); Tryon 1519 (H=-1030)5) . Posey Co.: S. A. 48h (No--31721) ; C. C. Deam 16813 (In); W. H. Welch 6929 (Bl 2346, Ca--892820, Dp—1L5h7, We). Pulaski Co.: Friesner 22373 196k Moldenke, Monograph of Verbena 211 (Au--122723, N, Ok, S). Putnam Co.: Banker 1361 (N); S. M. Deam 13490 (In); Fri esner ToS (Bt—1) 85h); 3 Grimes 16) (Dp); McDougal 5.n. [7-30-89] (Dp, | Dp); XYuncker 3677 (Dp). Saint Joseph Co.: Ce C.D Deam 49230 (In), Starke Co.: C. C. Deam 17954 (In), 31835 (In); Friesner 1631.5 (Bt--56415); W. H. Rhoades s.n, [near near Knox] (Hs). Steuben Co.: Ce. C. Ce Deam 514226 26 (Dm). Sullivan Co.: Craw se ne [July k, 1930] (Bt—5283); C. C. Deam 7307 (In). Tippecanoe : Cummins s.n. (Battleground, July 2, 1936] (Ah); C. C. Deam 17689 (In); Dorner 102 (Pu), 248 (Pu), s.n. [near Lafayette, 1900] (In); Ek s.n. [8-1-19)0] (Bt-—56621); Erisman s sn, [LaFayette, Aug. 25, '9h) (Ah); Foster s.n. [Wea Bottoms, 10/17/01) (Pu). Tipton Co.: Ek s.n. [8-5-19)2] (B (Bt--6498). Vermillion Co.: C. C. Deam 1196 (In). Vigo Co.: C. C, Deam 25762 (In); Friesner 6666 (Bt ULé1, Bt—-1)525); Mohr & Mohr s.n. [Terrehaut, 168h] (W--771851). Wabash Co.: C. C. Deam jean 53985 (Dm). Warren Co.: C. C. Deam 97h0 (In), 11903 (N)j Friesner 1101) (Bt—-38616). Wayne Co.: Creek 160 (Ra); C. C. Deam 13585 (In); L. J. King 1663 (Ea), 1664 (Ea), 2620 — "MU. Be" 156 (Ba). White Co.: C. C. Deam 39359 (In). County undetermined: Dorner sen, [9/28/99] (Ur); A. Wood s.n. [Ind.] (Pay. IOWA: Adair Co.: A. R. Moldenke 937 (Lm); H. N. Mol- denke 21939 (lw). Adams Co.: Dunn & & Isely 5707 7 (W-2099962). Al- lamakee Cos: L. H. Pammel s.n, - [New Albin, Aug. 6, 1918] (Io— 94219); Shimek s.n. [Aug. 13, 1917) (Ur); Tolstead son. [Hanover July 15, 1933] (Ca—882818), sen. [Hanover Twp., pe, Auge 9, 1933] (To—1s),282) « Appanoose Co.: L. H. Pammel s.n. (Moravia, Oct. 1914] (Io). Benton Co.: A. R. Moldenke 977 (Iw). Black = Co.: Burk 527 221 (Ur); L. H, Pammel 673 (Io—1192h3), sen. [Ce- Falls, Aug. 13 , 1908] (Io--78234). Boone Co.: Collector un- Sesignated Son. fea: a (La); J. V. Ellis 155 (Io— o—51,363)« Carroll Co.: L. L. H. Pazmel s.n. [Coon Rapids, . June 26, 1921] (Io— 9992h). Cass Co. hk R. Moldenke 93) (Lw). Cedar Co.: Fay 689 688 (Hi—123798), 852 (Ca—965065), 970 (Vi). Cerro Gordo Co.: A. Hayden 1, in part (N), sen. [October 15, 193] (N); L. He Pammel Sn. 3.n. [Willow Cr., Sept. 20-23, 1902] (Io—28203). Cherokee Co.: L. B H. Pammel s.n. (Cherokee, IX.5.1920] (Io--98223). Chickasaw Cow: L. H. Pammel 8.n. [Lawler, July 1919] (Io—94013). Clarke Co.: Pammel & Pammel o Sen. [Osceola, Sept. 26, '2h] (Io—-114346). Co.t As Hayden 3005 (Io—~1h9006), 3007 (To—-1,8885); We We A Weber 1272 ( (B1--]8176). Clayton Co.: A. R. Moldenke 991 (Iw); Pease Pammel Sennen Co.: 7 He Pammel s.n. [Comanche, Sept. 13, 1918] Sanaa Se iaossts Oe s.n. faeay a3; 1903] (Io—~52108) . Delaware Co. Ms 212 PHYTOLOGIA Vol. 11, no. 3 Moldenke 987 (Lw). Des Moines Co.: L. H. Pammel 735 (Io-~118882), 961 (Io--119553). Dickinson Co.: W. B B. F Fox 7 (We); Gier s.n. {June 21, 1952] (Je~7173); Mrs. L. Jones s.n. {June 2h, 1928] (Ob—-83155) ; Pammel & Pammel s.n. ~ [Arnold's Park, 7-23-13] (Io— 73719); .M. Roberts s.n. [Spirit Lake, Aug. 1920] (Io--97107), s.n. [5/25/23] (Cm). Emmet Co.: Cratty s.n. [13 July 1886] (0b— 50835), sen. (Armstrong, August] (Io—92218); W. H. Welch 9700 (Au--122739). Fayette Co.: Fink 251 [July 12, 189L] (w--2ih893), 251 (Sept. h, 189k] Se s.n. (Fayette, 7-18-189),] (Io-- 26816, Io--31575); J J. R. Gardner 583 (N). Floyd Co.: Arthur s.n. {Charles City, July 13, 1872] (Ah), s.n. [Charles City, July 16, 1872] (Ah), s.n. [Charles ree 7/15/74] (Ah). Fremont Co.: Deer s.n. [southwest corner of the state, Aug. 193] (Cm); L. H. Pam- mel 704 (Io—119300), sen. ‘eaiear urg, Jul. 1h, 1923] (Io—111220); W. H. Rhoades s.n. [Hamburg, Anes 1928] (Hs Hs) « Greene Co.: L. He Pammel son. (Jefferson, June 26, 1921] (Io—95859). Hamilton Co.: L. H. Pammel s.n. (Stratford, ae 13, '2h] (Io—115186); Pammel & & Zimmerman 306 (Io—119090). Harrison Co.: Collector undesignated S.n. [Dunlap, July mei (NJ); Fay 3714 (Vi); L. Kello L. Kellogg s.n- [Halsted's Am. Weeds 17h] (Ah, C, Fc, 1o—15352, Mi, Ua-—11397, Ur, Vt, We); Rigg s.n. [June23, 1898] (Se--195,). Humboldt Co: Hask sen. [Humboldt, May 21, 1902] (Io--22917). Iowa Co.: Easter ley 729 (We); A. Re Moldenke 972 (lw). Jasper Co.: Elrod 3.ne [Prairie City, July 7, '85) (Mn—-18071). Jefferson Co.: L. He Panmel s.n. (Fairfield, Sept. 1919] (Io—-95191). Johnson Co.? Loufek s.n. (June 1938] (Io—1h7639, N); McConaha s.n. {August 1935] (Bt--390h1); Somes 3489 (W--672272). Kossuth Co.: E. Be Watson s.n. [Algona, 9=5-02] (Io—7h13h). Lee Co.: Re Ae “David- son 2269 (W—2239821); Fults 1089 (Io-—138822); Shimek s.n, [July 6, sees (Ur). Linn Co.: A. R. Moldenke 980 (Iw). Louisa Co.: ¢ Davidson 858 (W--2261035); L. He Pammel sen. [Fredonia, June Bari (To=129476). yon Cos: C. R. Ball son. [Little Rock, June 30, 1897] (Io--15325); L. H. Pammel s.n. (Granite, Sept.1, 1920] (Io—97830); Shimek s.n. 1. [Aug. 26, 1910] (N). Madison Co.: A. Re Moldenke 940 (Iw). Marion Co.: hs R. Moldenke 956 (Iw). Mehaska Co.: Augustine 261 (Ok); A. R. Moldenke 957 (Lw). Mills Co.: Fay 3593 (Hi--126891); A. R. Moldenke 92) (Lw, Ut, Z). Mitchell Co.: Tuttle 3la (Io—7)177). Monona Co.: ; L. He Pammel sen. [Onawa, J 13, 1923] (Io—109650). Muscatine Co.: Estle & B & Brown s.n. [south of Muscatine, summer 1935] (Io—16363); L. H. Pammel s.n. [Jul. 20, 1919] (1095283) ; Shimek sen. [Oct. 1), 1919] (Ur). O'Brien Coes Yuncker & Welch 4110 (Dp). Palo Alto Co.: Cratty sen. [West Bend, Jul. 10, 1919] (Io—96078); We A. Weber 1261 (Bl—18175, To—150767) « Polk Co.: Bakke s.n. [Des Moines, 9-21-12] (Io—- 1964 Moldenke, Monograph of Verbena 213 53554, Io--7l062); Mosier s.n. [Des Moines, June 11, 189k] (W—- 859086); L. H. Pammel s.n. (Des Moines, Aug. 3, '27] (Io--129878, Ok--10367, St--9269). Pottawattamie Co.: Demaree 23662 (Sm); Fay 3128 (Ca--l5357); A. R. Moldenke 930 (Lw); L. H. Pammel s.n. [Oak- and, Aug. 27, 1919] (Io--96584). Poweshiek Co.: M. E. Jones s. ne (Grinnell, Aug. 1877] (Al, Ca—380556, Cm, Du--176622, Gg— Tél » Son, (Grinnell, Aug. 8, 1877] (Du--151777, Po--70872); A. R. Moldenke 963 (Lw); L. H. Pammell s.n. (Grinnell, Aug. 2h, 1930] (Io-~136710). Story Co.: Arthur s.n. [Ames, July 25, 1877] (Ah, Ah); C. R. Ball s.n. [Ames, June 30, 1898] (Io—15331); C. E. Bessey s.n. [Ames, 1880] (Io--95122), son. [Ames] (Io—15327); Beyer s.n. (Ames, August 1887] (Io—l5076); J. R. Campbell 36 (Io- 84556); Carver sen. [Ames] (Io—15328); P. Clayton 11 (Io—8)559, To—-8)560); Cratty son. [Ames, Jul. 29, 1925] (Io—117338); He Se Fawcett 39 (Fl—-21123, Io~-52791, Io—76859, N, W--)93414); Feath- Srey 8-n. (June 24, 192] (St--9260); Haven s.n. (Sept. 1st, 1838] (Ob-~50839); A. Hayden s.n. [August 10, 1933) (N); A. S. Hitchcock S-n. (Ames, July 182] (Io—15322), sen. [Ames, 7/31/89] (Io 15323), san. [Ames, 1889] (Io—15329); Jeffs sen. [7/10/1907] (Ok- 10364), sen. [Ames, 1907] (Io--75062); Kolpin 38 (Bm); L. H. Pam- mel 85 (Fc, Fl—21126, N, W—2802h2), sen. [Ames, August 2, 1096] (Io—15330), s.n. {Ames, Sept. 1913] (Io—57775); Pammel & Cratty Sen. (Colo, Jul. , 1922] (Io--105536); Pammel & Pammel 61 (Io— 502k6, Io—84558); stafford son. [Ames, 1906] (Io--82002); Ee Be Watson s.n. (Ames, 71-02] (Io--82978); W. A» Weber 67 (Bl 48177). Wapello Co.: L. H. Pammel s.n. (Ottumwa, Jul. 12, 1919] all 53 L. H Pammel s.n. [Sioux City, July 7, 1919] (10--97279). Wright Co.: Whited 2359 (Or—12378) . County undetermined: Suksdorf s.n. [July 20, 1873) (P1--138359) . Alma L. Moldenke "The Life of Plants", by E. J. He Corner, xii+ 315 pp., illustr. World Publishing Company, New York, Ne Y- 196k. $12.50 = wonderfully thought~ fascinatingly readable and provoking, fasc botanically valuable this book is! 1t should become required Povege for all who have any interest in botany. Despite the act that the author has "lopped off most physics, chemistry, 214 PHYTOLOGIA Vol. ll, no. 3 physiology, genetics and anatomy" there is still an excellen text de eyntopes with fine illustrations, chapter references, ibe liography and index. | Because my feeble wording may not do the book justice, I will include short quotes from a few of the chapters PREFACE: "land plants are understood as seaweeds ap ha from the Fig multiplicity of thalassiophytes, dried off on the face, waxed against evaporation, rooted, gy a for water rope built up with the transparent bricks made from the excess of sugars, yet still reproduced by seaways. THE BREAKING WAVE: "In snore with the plant plankton of free cells, the seaweeds the plant benthos of many-celled, attached, and mortal tnaiyionale « The oat the concept of vegetation." ING VEGETATION: "Thus the plant body (seaweed) comes to en- he supply of mineral salts are j e tendency to apical gr r he rock, so or a single cell in ompetition for foothold and for light, fits the mechanical needs 9 ed. plants un= S grasses and Sapo, convert it as best they can into ete of the leaves." LIVING SEAWEEDS: "In clothing the rocks the seaweeds have alter- the shore. 2 ig floated by the rising aie or prostrated in its falling, they shade and shelter. Under, around, and u ee them spring up new pinta to be exploited by other seaweeds and by animals, and they in their turn create their situations for aeallet organisms....Every growth makes as a fixjure fo r some ne tine, ,* micro~ habitat with a micro-climate where micro2organi REPRODUCTION AND WASTAGE: "Parasitic seaweeds are rare. Many aweeds grow epiphytically on others but they do not pene~ nourishment therefrom in the manner of mistletoes. Seaweeds seem remarkably antibiotic, or they are not prone to relinquish their photosynthetic independence." THE LAND PLANT: With a description of a tropical forest as the botanical culmination in land vegetation: "There is a giant tree, ent in a forest that stretches to t ag skyline. On its canopy birds and butterflies sip nectar. its branches orchids, aroids, and parasitic mistletoes offer pia to other birds and ects. Among them ferns creep, lichens encrust, and centipedes and scorpions lurk. In the rubble that falls anong ¢ the geht roots and stems, ants build nests and even earthw d snails find homes. There is a minute munching of patapeiitass and the 196), Moldenke, Book review 215 silent sucking of plant bugs. On any of these things, plant o animal, fungus may = growing. Through the branches spread ite s keys, a halting squirrel, or a bear in search of honey; the ie dow of an eagle startles them. ig os done snags fungus an beetle have attacked the wood. There e fungus brackets we round the edge and bored a other beetles. A woodpecker Eanes In 11 Where tress-roots a tortoise is eating toadstools. An elephant has k and, in i tmarks rests, tory, and thence sprung the actin of the creatures that in- telligentiy ae: them," [ARD STRUGGLE: "The of palms, screw pines, banana~ des, and other pachycaul CM oS is the swampy flood zone of tropical rivers, where on newly forming soil they seem to show how the flo owering forest was pioneered.” THE SEED: "of oes four “oe inventions of land plants, namely internal air spaces between the cells, cuticle, lignin, and seeds, they are geologically the latest and by far the most complicated. HE FRUITING TREE: "From such arillate fruits (durian) all ating Kinds have been derived by loss of the aril, by drying up into te rattling pod or capsule with its dry seeds, by indehis- vence leading to the Tyee succulence of berries, by increasing senfication leading to stone fruits and nuts, and by decreasing Size leading to small sation of all kinds." THE FUNGUS: "They have explored by means of their hyphae the — world more effectively than any other grag of organism, In ‘rs they are the counterparts of insects. 216 PHYTOLOGIA Vol. 11, no. 3 FUNGUS PROGRESS: "Top soil is the gold in the basement of the erial; in their insides they can digest lignin. They generally t eir munching can be he may duce to edibility such material as bark, which is not at first palatable. The animals excrete, defaecate, and leave slimy tracks. The fungus excretes and Lékved tracks of threads. Bacteria mul- tiply on these. Protozoa multiply on the bacteria. ers, nematodes, and other minute animals multiply in these soil colo- nies and add their excretions. Beetles, spiders, centipedes, and 8 the scorpions prey on the soil ir quota of waste oducts, all of different chem ores abound, ger- ung ted, on these microscopic eggs m Ship rotifers, and mites feed, A mouldy sm compounded many odours, marks the solid, liquid, an gaseous deccieet eine in this microcosm of chanical engineering." BEYOND THE FOREST: "From all over the earth -- tundra, desert, mountain, ocean island, and lake -~- we gather the threads ife the « ct ° ct z o a trees flowering, cell grew into the plant form. But the forests, which show how trees were made, are going. They ar 2 vanishing nowhere faster than from the alluvial plains where the sactepee of the last creative acs of plant life, that prepared the way for the modern world, odern 7 weriive, The mo mouth is the people's, and theirs the new aed a ore machines the forest is defenceless. progress is clearing it with gathering sp o plant crops of quick returns. The botanist ould take the oppor- that a few brief centuries of his science have revealed; for soon there will be rice-field to every river-brink. The unmindful tree begot, indeed, the seed of its own destruction." I hope that many botanists will second effectively Dr. Corner's claim eat "Ye need a new generation of alert young biologists eager to enquire into the lives of plants. Laboratories are the a fy not the prisons, of research," Herbarium curators who have material of this species from additional counties are asked to send it to the author for verification and Figure 17. Distribution of Verbena stricta in the United States Mapping by counties done |! record, so that future editions of this map may be more complete FOSTER a ae Consow ~ lead BE ae eid. wee aa-eem.. . po aiininde 5 dedi ee ee a : | : ye es Ona midge. TEPMENS THLLMag ee i ar ee Eb, oe S * Ms v | 4 om 2% ¢ tS GREENE ee KEYED TO COLUMN A of Cleartype Index Book an co TT Fille Heel tem eI CLEARTYPE he AN Seosst (na } TRADE MARK REG. U.S. PAT. OFF. alk i ele oe rage COUNTY OUTLINE : fe UNITED STATES & Scale of Miles 4 ae 0 100 | soll - Bae: Se AMERICAN aps ‘ eal —_ FOS CLEARTYPE MAPS Suge) TOL ESS CT ey ! > WY a » walls. & yf ‘NG Wage sae el ~ 2 == a ks ie ~~, “~: ; . \ ; * > i ro ae 5 Pn Copyright, American Map Co., Inc., New York, No. 14324 ee Va ae E T 7) T 2 T 7) T ee ees oo a I [SP es ae SO OR ee I 2 It u if 3 EE) \ PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and chess ecuiees his con- tribution. All money received from subscribers, after the expense col- lections have been deducted, will be distributed among te aes upon the completion of a volume, in proportion to the space which they have used. contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. Each number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary . A volume will contain about 32 signatures, 512 pages, or a smaller number of pages with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra — is made for line drawings, such as are ordinarily msn - _ zine, or for diagrams, tables, or charts, provided they con limitations of size and proportion. An extra charge will be made be half "tones, depending on their size, as fixed by the engraver, with a minimum of © 7 shout — ible nine with research in all lines of hotany, in any smn . ie gth, biographical sketches, and critical reviews and sum of Ii = ue willbe considered for publication. Floristic lists, casual not fa | amateur or so-called popular t , will not be cpu Advice gly vk i- Unde te pron nf ining he he hasic rate for a page or fraction ximate See es ae This prices subject ntirely on the prices prevailing ues Ae tas structions concern : further information about the ‘magazine. In- : to either editor. : PHY TOLOGIA Designed to expedite botanical publication ‘eee January, 1965 No. 4 CONTENTS . KING, R. M., Chromosome numbers of Thailand Compositae . oe, 217 MOLDENKE, H. N., Materials toward a monograph of the 3 genes Verbena. XXVI cc. sven vec tewtcsssieneecansee sete 219 , : MOLDENKE, A. L., Book reviews bicker a he 287 Published by Harold N. Moldenke and Alma iL Moldenke — 15 Glenbrook Avenue yes é a Yonkers 5, New York, U. S. A. os Price of this number, $1; per voles: $5. 75, in advance, = or $6 at close of volume z: BOTH: “1, no. 2 was. issued December 2B, 1964; : = “il, no. 3, was, iapued Dee i 18 : : CHROMOSOME NUMBERS OF THAILAND COMPOSITAE by . M, King nc Amen 8.50, SSA, ould like to report chromosome counts for 15 collections represent ing 8 species of Compositae collected in Thailand Ea © summer of 1963, Buds were collected in field in 4 part chloroform, 3 parts absolute alcohol and 1 part glacial icdese LITERATURE CITED Cave, M. S. ed, 1956 ~62, Index to plant chromosome numbers. Un iv. of North Carolina Press, Chapel Hil Chuang, T., C, Y. Chao, Wilma W. L, and S, C. Kwan, Chromo- some numbers of the eels ais of Taiwan I Taiwania 8: 51- Darlington, C, D, and A. P, Wylie, 1956. Chromosome Atlas of Flowering Plants, MacMillan Co,, New York, Turner, B, L. and R, M. King. 1964, Chromosome Numbers in = Compositae VIII, Mexican and mere al American Specie The Southwestern Naturalist 9 (1): 27-39. 217 TABLE 1,--Chromosome numbers Species Ageratum conyzoides L, Ageratum conyzoides L, sumatre _— Retz, ca, Eupatorium Gnaphalium eee -album ssp, affine (D. Don.) Koster Gynura pseudochina (L,) Synedrella nodiflora (L.) Gaertn, ca. Vernonia patula eel ) rr. var. patul Wedelia biflora (L. DC. Wedelia biflora (L.) DC. Wedelia biflora (L.) DC, 24°) of Thailand Compositae, Is i] Voucher Collection Previou (by Province) Count Reference To n Previous Count Nakhon Rat Sima, King 5528 20 Turner & King, 1964 Nakhon Nayok, King 5537 Ranong, King 5584 Prachuab, King 5567 Pranakorn, King 5597 Chieng Mai, King 5469 7 Darlington & Wylie, 1956; Cave, 1960 Kamphaeng Phet, King 5438 Nakhon Rat Sima, King 5524 19 Cave, 1962 Pranakorn, King 5563 Pranakorn, King 5555 15 Chuang et al., 1962 Pranakorn, King 5556 Pranakorn, King 5596 VIDOTOLAH q tou “TT *TOA MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXVI Harold N, Moldenke bic mer eae ent. ibliography: Parks & King, El Crisol [Puerto Rico] ks pans 1960; then. Abstr. 45: 3037. $969; M. L. Buch, U. S. _ Agr. Handb. 6h: 13, 19 arks & King (1950) and Buch ee! report the occurrence of none acid throughout this plan Additional citations: uc “Ballara Co.: Anderson & Mc Farland 253 (Ky); McFarland & Anderson 253 (Vi), s.n. “gn, (Wickliffe, Aug. 16, 1923] (S). Campbell Co.: van Van Cleve s.n. ~ [Dayton] (Ky). Chicalsins Co.: Lix usb (W--191,8092) . Fulton Cos: Biltmore Herb. 2653 (N). Hickman Co.: H. A. Gleason 8978 (N). Kenton Co.: E. L. Br Braun s.n. [Devon Park] (Cn). Marshall Co.: Eggleston n 4837 __ (N). TENNESSEE: Chester Cos: S. M. Bain 328 (C). Obion Cow: vennings & Jennings s.n. [Reelfoot Lake, 192h] (Cm). Smith Cos: Gattinger Son. [Hickman] (W--1)7589). MICHIGAN: Alpena Co.: Cain, Raymond, & ie Kucyniak 583 (Go, Mg, S, Vi). Charlevoix Co.: ie He Ehlers 642 (Mi, Po--63839). Cheboygan Co: J. H. Ehlers 5165 (Mi), 5613 ( reas 82816, Mi); E.G. Voss 1038 (Mi), 1315 (Mi). aye Cows Scholes & Rafferty RL.97 (Mi). HBumnet Co.: J. H. Ehlers 2h53 (ii), 3489 (Hi--137h5, Mi), 4856 (Ca--l:98895, I), sen. (We of Pellston, July 1923] (Ka--68122); F.C. Gates 14450 (Ka--7001h). Grand Traverse Co.: Dieterle 1850 (a, ghar so Gratiot Co.: C. A. Da- wis $.n. (Elwell, Sept. 1900] (Mi, Mi). Kalkaska Cos: He Ne Mol- denke 21 (Lm) . Kent Co.: Bazuin 2232 (Mi); Mulliken s.n. (July 9, i 1897] ( (Mi); Parmelee 826 (Au--1915)8) . Livingston fingston COs: Out Fe J. Hermann 9159 (Mi, N). Menominee Co.: Grassl 2750 (Mi, N). Missau- Kee Cou: Be E. Ge Voss 3065 (Mi). Montcalm Go.: A. R. Moldenke 1033 (Lw). Oakland Co.: Farwell 6359 (Mi). Saint Joseph Cow: Ge Le ssenar S.n. [July 2, 1923] (S). Washtenaw Co.: Dodder s.n. . {July 1931] (Ok—=10 365); Frickelton sen. (July 13, 1921) 1] (Ok--10366) « ~~ fares F.C. Schmid s.n, (Romlus, August 1, 1943) (Mi). WIS- Adams Co.: Re T R. T. Brown 117 (Ws); J. We thomson son. (Aug. 1s, 9571 (Ws). Brown Co.: Shinners 3798 (Ws). Buifalo Co.t He ae Saith 712 7113 (B, Ws). Columbia Co.: Alberts s.n. [Sept. 29, 115) (We); Rob. Bu Burton 12h (Ws); H. P. Chandler 39 (Ca--25189); Fern- pols Biss. (Aug. 1 18, 1930] (Ws); He K: Herz 1 (Up); eke "s)} Orpunt s.n. [7/5/49] (Ur); H. L. Russell san. (Sept. hi, '85] (Wa) ; F. J. W. Schmidt s.n. [Oct. 1, 193h] (WS); Fe C- hepeomt 20280 (is); i H. Smith 6034 (B, Ws); J. Wright s.n. [September 219 220 PHYTOLOGIA Vol. 11, no. 29, 1945] (Ws, Ws). Crawford Co.: Fassett 9788 (Um--205, Kn. He W. Graham s.n. [July 9, 1928] (Cm); D. Marsh s.n. {Sept 1959) (Ws); A. R. Moldenke 993 (Lw); “Pammel | & Pammel s.n. pal e du Chien, Aug. 8, 1929] (Io—13383k); H. H. Smith 75 (B, Ws); = Sprague s.n. (7/ls/26) (cn, Cn); Ugent s.n. [July 25, 5, 1960] (Ws). Dane Co.: C. C, Albers 3301) (Au), 34007 (Au); Bakken = (Ws); Botany 162 Class s.n. (Sept. 28, 1935] (Ws); L. S.C s.n. [July 23, 1888] (Ws); H. M. Clarke s.n. [sept. er Tis, Ws); Greeley s.n. [Madison, July 3, 1949] T (Ga--90356); Grether Grether 8189 (Ws); H 3; Heddle 2421 (Ws), 2678 (Ws), 2728 (Io--109880); Koep: oep= pen 117 (Ws), 120 (Ws); Lapham m s.n. [1858] (Ws); McCabe son. [27 July 1945] (Ws), s sen. [13 August 195] (Ws); A. R. Moldenke 1012 (Lw); noe S.n, [October 10, 1959] (Ws); Reinert Son. [10/8/59] (Ws); Schlising & Musolf 60-111 (Ws); A. B. "Seymour - 1063 (H-- 107722, Ur, Ws); Shinners 2h39 (Ca——882819) ; H. L. Smith mith Sone {July 27, 1885] (Ws), s.n. [July 27, 1887] (Ws); “Ugent s sen, (July 31, 1960] (Ws); Van Arsdel s.n. [17 September 1952] 2} (Ws)3 P- P. Be Whitford 1821 (Ws). Dunn Co.: Weinzirl s.n. (Jy. 23, 189k) ] (Ws). Grant Co.: Iltis 9h8 (Ws); Iltis & Salaman 10103 (Ws); A- Re Moldenke 996 (Lw); L. H. Pammel s.n. (Muscoda, Aug. 15, 1: 1930) (Io—136579); Shimek s.n. (Aug. 11, 1921] (Ws); He H. Smith 7806 (B, Ws); Sylvester 1) 101 (N); Ugent s.n. [July 25, 1960) (Ws). Green Co.: E. W. F E. W. Fell 11 58-167 (Ws), ~ 56-611 (Ws). — Lake Coe? A. Re Moldenke 1017 17 (Lw). Lowa Co.: lltis sn. [end of June 1957] (Ws); A. R. Moldenke 1001 (Iw); Wills s.n. [July 2 » 1957] os Jackson Go.: Grether 6613 (Ws). Jefferson Co.: Shinners 2400 (Ws); Wadmond 3769 (Dm ree s.n. (Aug. 6, 1927] (Ws). Juneau Co.: Mearns 525 (W--670686) ; W. R. Taylor 2768 (Up--75581). La Crosse Co.: oe G. Hartley 1003 3 (W--2261909); King & Pammel 5-n- [La Crosse, Aug. 26, 1901] (To—-22920); D. Pammel s.n. [La Cros~ se, 1902] (Io~-22918) ; L. H, Pammel s.n. [La Crosse, Aug. 1882] (1015350) ; A. M. Peterson 187 (Ws); Ugent 433 (Ws). Lafayette Cos: Arnot 3.n. [duly 26, 1956) ] (Ws); Haggene, e, Melville, & & & Shaugh- nessy s.n. [Sept. 2, 1956] (Ws); P. K. Nelson son. (June ie 19h6] aval sen. [7 July 196] (Ws); Re 0 O. Wagner s.n. [Sept. 1956] (Ws). Lincoln co,: F. C. Seymour Tau (is, Ws), 12263 (is). Marinette Co.: J. J. Davis s.n. [July 3 areas) (Ws). Milwaukee Cow: Je Ss Douglas s.n. [Wis.] (Je~-3159). Monroe Co.: Iitis & Neess 8965 (Ws) ; bake | M. Peterson 15 (Ws). Oconto Co.: SC chlising & Schlising | 1018 (Ws). Pepin Co.: “0. Anderson 10 (Ws)- Polk C0+? rh ce (Ws); M. A. Fosberg 337 (Ws). Rock Co.: R. Be Anthony S-N- { 22, 1933] (Ws), s.n. [Sept. 2%, 1936] (Ws); E. W. Fell 57-059 1965 Moldenke, Monograph of Verbena 221 boygan Co.: Goessl son. (Sept. 1924] (Ws). Trempealeau Co.: Fassett & Wilson 9762 (In, Ws). Vernon Co.: H. H. Smith 7361 (B, Ws). Walworth Co.: Almon s.n. (Aug. 8, 1926] (Ws); Kleeberger s. n. (Whitewater, 8-25-1877] (Gg—-31h18). Washburn Co.: Goessl 8501 (B, Ws). Waukesha Co.: Fernholz s.n. [September 21, 1929] (Wis). Waupaca Co.: Goessl s.n. [July 28, 1910] (Ws); Rill s.n. (July 25, 1959] (Ws). Waushara Co.: Iltis, Bell, Melchert, Pat- n. [June 30, 1959] (Ws). County undetermined: W. H. Chase 8.n. o: N. Moldenke 7785 (N). Big Stone Co.: Eifrig s.n. (Ortonville, July 2, 1929] (Up); Gier s.n. (Ortonville, VI.30.52] (Je—-7183). Clay Co.: 0. A. Stevens 1271 (Ca--952815, W--2034558, Wp). Clear- water Co.: Terrell 1789 (W—2388691). Fillmore Co.: J. L. Leedy 8-n. (north of Rushford, June 29, 1937] (Bt—-5536h). Goodhue Co.: J+ H. Sandberg 28) (B). Hennepin Co.: G. B. Aiton s.n, (Minneap- clis, July 190] (N), s.n. [July] (Dt); F. H. Burglehaus 8166 (Se- aed Chandonnet s.n. [Minneapolis, Aug. 22nd, tat (In); Eaton S-n. (Minneapolis, July 1890] (N); D. Houghton 29 (T); J. H. Sand berg 7566 (Fe), sen. [Aug. 1879] (ca--28172), son. [Aug. 1869] (Ca—25171, W--71935), sen. [July 1891] (Dt); Schustte s.n. (St. Anthony, July 7, 1888] (N). Kandiyohi Co.: W. D. Frost s.n. (Willmar, July 1892] (Pl1—22612, Ss, W-~71936)- Lyon Co? L. H. Co.: F. Blanchard s.n. (St. Paul, July 31, 1890] (Dt); Detwiler 8598 (La); Mearns 525 (W—649505, W-~-6l,9506), sn. [Fort Snelling, 16667 (Ur), 16668 (We). Winona Co.: Hasse s.n. (Winona, July 1882] (n); Holzinger s.n. (Winona, Aug. '88] (Ca--199069), SMe (Aug. 1901] (N, Vt); Hotchkiss & Jones ll6h (W--1622897). County undetermined; H. S. Young Jr. sen. [6-10-1925] (Cm). NORTH DAKO- TAs Cass Co. 0. A. Stevens Sine [Fargo, July 25, 1934] (Ca-- 588086, cn). Ransom Co.: 0. As Stevens s.n. [Lisbon, July 25, 19b4] (W-~2008)69). Richland Got O. A. Stevens 318 (Ca—607217, 222 PHYTOLOGIA Vol. 11, no. Hi—1920h7, N, Ok, Um--212). SOUTH DAKOTA: Brookings Co.: L. H. Pammel s.n. (Brookings, Jul. 30, 1922] (Io--106919); J. J. Thorn- = 8.N. 1. (Brookings, August 1893] (Tu). Brown Co.: Griffiths s. 3. n. (duly 31, 1872] (P1—22613). Cherry Co.: Tolstead s.n. [July iz, 1935) (Io—-145164). Custer Co.: Barr sen. [July 25, 19)0] (Ba, Se--95483); Degener & Peiler 16269 (N, N, N)3 Hayward 1697 (Se--77057); Tollefson efson 16 (Sd--2529). Day Coos J. F. “¥. Brenckle 39-92 (N), sen. [July 23. 1938] (B). Deuel Cos: fu Brenckle 602° (N). Fall River Co.: J. W. Barnhart s.n. [Hot Springs, J 13, 189k; Herb. J. H. Barnhart 642] (N);Rydberg 932 (W--211121). Hanson Co.t G. F. Weber s.n. (Farmer, 8-18-27] (Fl—21120). Hutch- inson Co.: V. oe Harms 159 ( (Lw). Hyde Co.: Pammel & Fink s.n. (Highmore, June & July 1912] (Io—-79613). Kingsbury | Co. Oot Je Je Thornber s.n, [Iroquois, 8/11/9\] (Ca--12h662). Lawrence Cost F. L. Bennett 2 (Ua—l3209); W. P. Carr 118 (N, W-751127); He Ee Hayward 202 (N); P. Johnson 25 (Bt--6230h, Ky, Lb—-36500, N, Ur); G. N. Jones 19746 (Ur). Meade Co.: Forwood 299 ae: Kemp | s.n, [Bear Butte] (C). Mellette Co.: Moses s.n. [White er) (Pr); Tolstead sen. [July 9, 1935] es ewaey inden Co.: He N. Moldenk Moldenke . 7676 (N). Pennington : O. M. Clark 6044 (B); H. N. Moldenke 770k (N); Over 1507 (Mee; Visher 1129 (Se—11952), 1507 (N). Spink Co.: J. F. Brenckle 47-533 (3 Petrak sen. [31.VII.1950] (S, 8); A. E. Ricksecker 48 (Ca~- L75kh1, Ob—50841, Po--215279, Um—21l, Ur). Stanley Co.: Over 6168 (B1l—l2337). Turner Co.: V. Le Harms 11 (Lw). Washabaugh Coe: Kienholz 47 (Ur). Yankton Co.: L. He Pammel s.n. (Yankton, si: 6, 1919] (Io-~96322). KANSAS: Allen Co.: A. S. Hitchcock Se n. [Allen Cos, July 1896] (Ka). Anderson Co.: A. S. Hitchcock 8 Se R, [Anderson Co,, July 1896) (Ka); W. H. Horr s.n. [July 22, 1929] (i); A. R. Moldenke 880 (Iw). Atchison Co.: A. S. Hitchcock 8+Me [Atchison Go., Oct. 1896] (Ka); Horr & McGregor sen. [6/26/46] (Lw); Jeanette 72 (I); A. R. Moldenke 908 (Iw); Nolte 10 (I)- Bar- ber Co.t A. S. Hitchcock s.n. [Medicine L., July 1892) (Ka); Kel- lerman s.n, [Medicine Lodge, July 1h, 1888 (Ka); Re L. McGregor 1068 (Le), 14435 (Lw), 1sh63 (Lw), 14849 (Lw); J- ta De , Poindexter 201-6 (Iw); A. R. Moldenke 695 (Lw); B, L. Wagenknecht 1369 (L™)- Bourbon Co.: Kellerman sone sen. [Ft. Scott, | duly 15, 1887] (Ka); Zs a D. Thompson 587 (Lw). Brown Co.: Clothier & Whitford s.n- [guly 2 1897] (Ka); Garner sen. [Aug. 1925] (Ka--72013); We He iH, Horr ius (Iw); Re Le Hedreger r 17058 (Lw); A. R. Moldenke 915 (Im). ye Butler Cos: J. Da S.n. (Douglass, July 22, 1887] (Ka); W. H. Horr 3-Re (8/6/29) (iw) ee J. D. Johnson s.n. [July 1926] (Ka--71719) « Chase Co.: Kellerman s.n. [Strong City, July 27, 1887] (Ka); B. L. Wager ay 1965 Moldenke, Monograph of Verbena 223 knecht 432) (Lw). Chautauqua Co,: W. H. Horr sen. (July 5, 1930] (L¥)5 A. “= Moldenke 887 (Lw); B. L . Wagenknecht 1831 (Lm), 1853 (Lr). rokee Co.: N. B. Jacobs 16 (N); Kellerman son, [Colum- bus, sae 13, 1887) (Ka); | E. W. Lathrop 2619 2619 (Lw); ; Re be sicanegen 1708 (Lw); J. D. Poindexter. 207-27 (Lw). Cheyenne Co.: Bofman jofman 22 (ky), 311 (Ka--92665); Ve L V. L. Harms 1219 (Lw); A. S. Hitchcock s. n. [Chey enne Co., June 28, 1897] (Ka); W. H. Horr 689 (Lw); R. Le McGregor 9)2 (Lw); Roests "ya (Au--122717). Clark Co.: A. S. heock s.n. [Clark Co., Aug. 1896] (Ka). Clay Co.: Kellerman Son. [Clay Centre, July 2,1 1888] 53g L. McGregor 5012 nd L). Cloud Co.: Carleton s.n. (July 9, 1888] (Ka, Ur); Se | S. V. Fraser UO (Ka~-78937). Coffey Co.: W. He Horr 3475 (Lw, “}-d030701)5 Wimpey s.n. (Burlington, July 1396] (Ka). Comanche Co.: A. Se Hitchcock sen, {Comanche Co., Aug. 1898] (Ka); R. kL. McGregor i677 (Im). Cowley Co.: Clothier & Whitford s.n. [Aug. 22, 1897] (Ka); R. L. McGregor 10397 (Iw); H. N. Moldenke 21926 (Lw). Craw- ford Co.: Clothier & & Whitford s.n, (Aug. 12, 1897] (Ka). Decatur Co.: Kellern an s.n. . (Oberlin, July ti; 1888] (Ka); Re L. McGregor 9478 (Lm), 13530 (Iw); E. L. Richards "10148 (Lw), 2860 (Iw). Dick- inson Co.: A. S. Hitchcock s.n. [Dickinson Co., June 1896] (Ka); Lo H. Horr Son. . [July 19, 1930] (Lw); H. N. Moldenke 21931 (Lm, Ut); J.B. Ss N Norton s.n. [September 1895] (Ka). Doniphan Co.: Clothier & Whitford s.n. {July 31, 1897] (Ka). Douglas Co.: Channa” ’ eee 1 (iw); Garrett 8466 (Se--14953); Kellerman s.n. TE eating 20, 1887] (Ka); R. L. McGregor 115 (Lw, 8); J. D e 31 (Im), 229-6 (tm)° Be i6 Taye Bie ban (Sui) (F-21122); W ¥ Ge. Stevens 672 6728 (Lw), s.n. Se eT We (W--216103). Edwards A, S. Hitchcock s.n. [Edwards Co., Sept. 1, ree (Ka); Re Le “Ye Gregor 10538 (Lw ). Elk Co.: Clothier & Whitford Sen. [aug. 2 "20 & 2, 1897] (x (Ka); A. R. Moldenke 877 (Lw). Ellis Co.: A. S. Hitch Sock san. [Ellis Co., August 1895) (Ka); Imler s.n. (June 20, 1928] (m);"E. Runyon 175 (51—2348, Bt—Ul919, Ga, Go, Lb—33196, se 8, St—9263, Vt). Ellsworth Co.: A. S. Hitchcock s.n- [Ellswort Co., July 1895] (Ka); R. L. McGregor 9260 (Lw), 12388 (Lw); Ae g ae 2 899 (Lw); B. L. Wagenknecht 1907 (Lw), 1971 1971 (Lm), 3232 Finne ey Co.: Wilson & Miller 6728 (Lm), sen. [7-22-12] (L (L¥)- Ford Co.: Kellerman Son. Bucklin, June 26, 1888) (Ka). oC jeteaee 37h (Iw); A. S. Hitchcock s.n. [Franklin Co., July ] (Ka). Geary Co.: F yc C. Gates 1750 (Ka—80314), 19957 (Au), 250 (N); Gayle 510 (c); Hulbert 3298 (Lw); H. N. Moldenke 21932 (x) 5 52s F. Nort Norton s s.n. (June 27, af 19] (Ka). “Gove Cost Ae Gl icae Aiehcock san. S.n. n. (Gove Co., July 1 95] (Ka); Ge Je Inkenberry 59 18206), 3 8.n. (Sept. 10, ys (Ca—9l, 3061). Graham Coe: we 22h PHYTOLOGIA Vol. 11, no. 4 S. Hitchcock s.n. (Graham Co., June 20, 1897] (Ka). Gypeley Cost Me Reed s.n. [Tribune, Aug. 5, 1892] (Ka), sen, (Tribune, Sept. 11, 1892] (Ga—-10h870); Rose & Fitch 1707h (N, Saree Green- wood Cos: A. S. Hitchcock s.n. [Eureka, July '92] (Ka); W. H. Horr s.n. {guly 28, 1930] (Lw); Ripley & Barneby 7251 (N). “Hamil- ton Co.: C. He Thompson 15h (Ca—-1918h9, Ka, N, W--2658)0). Harper Coe: Kellerman Bon. 3.n. (Harper, July 14, 1888] (Ka); A. R. Moldenke 889 (Iw). Harvey Co.: V. L. Harms 669 (Lw); Herrick Sone . [Auge 1898) (Ka); Muck 82 (Ka--92760). Hodgeman Co.: ass Ss. ~ Hitchcock sen. (Hodgeman Co., August 1895] (Ka); R. L. MeGregor 10595 (LW). Jackson Co.: Norton & Clothier s.n. [July 5th, 1895] (Ka); M. Reed sn. [Avoca, July 25, 1891] (Ca--25170); E. L. Richards 919 (Lm) « Jefferson Co.: G. Latham 259 (Iw), 350 (Lw); D. L. Marsh . 1613 (Hi- 16208, Lw). Jewell Co.: R R. L. McGregor 7377 ” (Lw) « Johnson Co.! Kellernan s.n. [Olathe, July 7, 1887] (Ka). Kingman Co.: Branaman Sen. [July 29th 1895] (Ka); R. L. McGregor 1072) (Lw). Kiowa Co-: A. S, Hitchcock s.n. [Kiowa Co., Aug. 1096] (Ka). Labette Co.: Ae oi Hitchcock S.n. . [Labette Co., Aug. 1896] (Ka); = Le Wagenknecht 1890 (Iw). Lane Co.: A, S. Hitchcock s.n, [Lane Co., August 1895] (Ka). Leavenworth Co,: : Clothier & Whitford s.n. bing! 3, 1897] cael Herb, Western Fem, Sem. 5.n. [Stranger] (Pu); Sloan 4 (Ka-- 3196). Lincoln Co.: A. S. Hitchcock s.n. Fasise Co., July aan (Ka); R. L. McGregor 7hh6 (Lw). Linn Co.: Brooks & Brooks sen. [1926] (Cm, Cm); Kellerman 1 (Ka); A. R- nnaake denke 871 (L), 881 (Iw). Logan Cos: A. S. Hitchcock sn, [Logan Co., July 1895] (Ka); R. L. McGregor 908 (Lw), 17236 (Le) (Lw). Lyon Co.: He Ae more s.n. (Baporia, July 1, 1922] (Je); R. L. McGregor 1885 5 (ims B. L. Wagenknecht 1,180 (Lw) . Marion Co.: Griffiths 5050 (W oa 196552); A. S. Hitchcock sn. {Marion Co., August 1895] (Ka); Re be McGregor 10764 (Lw). Marshall Co.: A. S. Hitchcock S-ne [Mar- Co., Dec. 10, 1896) (Ka); W. He Horr LSuB (Lw). McPherson fo G. R. Babcock s.n. [Roxbury, June 23, 1887] (Ka). Meade Co-? W. H. Horr 3235 (Lw), 3275 (Iw), 3278 (Lw), 3450 (Lm, W--20 30692)» 3238 (Wj Kellerman s.n, [\feade Centre, June ‘ie A 1888] aes he De xter 166-17 (Lw). Miami Co.: A. S. Hitchcock s.n. [Pao Aug. 1892] (Ka). Mitchell Co.: H. ‘.; Dwyer 5.n. {July ay 1896] (xa). Wontgomery Co.: A. S. Hitchcock s.n. [Montgomery Co., August 1596 (Ka); W. H. Horr s.n. [July 5, 1930] (Lm); A. R. Moldenke 882 "aa Rydberg | & Tt Imler 33 (N). Morris Co.: Bs He “Horr sn. [July 1930] (Lw); Re L. i McGregor 1413) (Lw); Trembly s-n. $.n. [Summer 1854) (Ka). Morton Co.: E. L. R B. Le Richards 2959 ae 3533 33 (Lw). Nema Co.: A. S. Hitchcock s.n. [Nemaha Co., Dec. 11, 1896] (Ka); Pettit Px(ER)s Mencia Gos W. W. Holland 130 (Im); W. H. Horr 6728 (Lm) 1965 Moldenke, Monograph of Verbena 225 Norton Co.: F. C. Gates 16900 (Ka--76273); W. H. Horr 5022 (Lw); Kellerman s.n. {Norton, July 11, 1888] (Ka); | E. a ,. Richards 2781 oe Osage Co.: W. H. Horr £233 (B, Ca--k5170, Lw, No—12h02, “hg? TEL PBR 1 ee “W--2031022, We); Kellerman s.n. (Burlingame, at aig 1887] (Ka); B. L. Wagenknecht ht 4O4O (Lw). Osborne Co.: Dawley s.n. [1893] (Ka); Loomis sen. [Alton, Oct. 1938] (Ke— 90127);"R. R. L. McGregor 7418 (Lw), 9626 (Lw); Neher 115 Ae 80028); Shear s.n. [Summer 1894] (Ua-—2hhh7). Ottawa Co.: A. Hitchcock s.n. [Ottawa Co., July 1895] (Ka); H. N. Moldenke ahi (F). Pawnee Co.: A. S. Hitchcock Sn. iremee: Co., August 1895] (Ka). Philipps Co.: A. S. Hitchcock sen. [Philipps Co., June 29, 1897] (Ka); W. H. Horr L767 (Iw); R. L. if i: as 9542 (Lw); EB. Le Richards 2732 (Lr). Pottawatomie Co.: | F. C. Gates 3s 12613 (Ka— 72797, Lb—27620) ; 3 Kellerman s.n. [St. George, June 23, 1888] (Ka); L. D. Marsh 1735 (Hi--162532, Lw, W—2327297); B. L. Wagen- Knecht 3256a (Iw). Pratt Co.: A. S. Hitchcock s.n. [Pratt Co., Aug. Aug. 1896) ( (Ka); Hulbert 3189 (Lb—50539, Lw). Rawlins Co.: Ve L- Harms 12h) (lw); A. S A. Se ~ Hitchcock son, (Rawlins Co., June a7) (Ka); Re L. M McGregor 960 (Lw). Reno Coe: A. S. Hitchcock s (Tudor, July 1892] (Ka); R. L. McGregor 12510 (Lm). ys Republic C ses G. E. Morley 232 (Lw); D. K kK. “Thomas s.n. (Scandia, 1890] (Ka). Rice Co.: A. Se Hitchcock sane {Rice Co., August 1895] eg B.L. Wagenknecht 543 (Iw). Riley Co.: I. M, Atkins s.n. (Oct. 1 1935] (St—21L11); F. Cc. Gates 12660 (Wi); V. C. Hubbard s.n. (Sept. 1927] (Lb--27615); Maus s.n. [Rocky Ford, June 16, “527) (Mi); J. B. S. Norton 31 (Ka, N, W—353118); Payne s.n. [Manhat- — << SF a ST ¥. t Swingle s.n. funana tea June 20, 1888] (Ka). Rooks Co.: R. Ly McGregor 959 9590 (Lw); Wilkin s.n. fjone 15, 1887] (Ka). Rush Co. * A. Se Hit S. Hitchcock son, [Rush Co., August 1895] (Ka). Russell Co.: A. S. Hitchcock s.n. (Russell Gos, July 1895] (Ka); We He Horr 1937 (Lw); R. L. McGregor 5683 (Lw), 9283 (Ly). Saline Co.: Hancin 39) (Ka~-76791); A. S. Hitchcock s.n. a. eae ei June 1896) (Ka); H. N. Moldenke 21930 (Lw). sour Coe: Re Gregor 2367 (lw). Sedgwick Co.: T. L. Andrews s.n. [Whic war Get 2 To—28276) ; Herb. Wichita Univ. 12 (LW); 7 “ye Horr 8.n. (8/8/29) (Lw) ; A.S. Miller s.n. [i 3M. (Wichita, | Mar. 1892] (Ka). Shawnee Co.t We a. Dick s.n. sen, [6-12-87] (Ob--50837); Maus 847 (Ka-~72355, Um— 206); Volle 202 (Lw), 232 (Lw). Sheridan Co.: A. S. Hitchcock s- n. (Sheridan Co Co., June 21, 1897] (Ka). Sherman Co.: F. Ce Gates 20774 (Ka—76399)5 Ve Le Harus 1207 (Lm, N)3 A. S. Hitehcock son. (oodiana, Jan. 1892] (Ka). smith Co.: Vv. L. ” Harms 1312 (L¥); 3 Kel-~ ierman S.n. [Smith Centre, July 10, 1888] _ (Ka)3 E. | E. Le 7» Richards ED 226 PHYTOLOGIA Vol. 11, now § (Iw). Stafford Co.: R. L. McGregor 13711 (Lw); A. R. Moldenke 897 (Lw); Ungar 788 (Iw). Sumner Co.: Kellerman s.n. fArgonia, July 26, 1887] (Ka). Thomas Co.: A. S. Hitchcock s.n, [Thomas Co., mae 23, 1897] (Ka). Trego Co.: D. A. Horr 4173 (Lw); Re Le McGregor 9331 (Lw); M. Reed s.n. (July “8, 1892] (Ka). Wabaunsee Co.: Clothier s.n. [June 22, 1896] (Ka); Maus 30 (Ka-~7269, Ob— 50832). Wallace Co.: R. L. McGregor 17286 (Lw). Washington Co.: Clothier & Whitford s.n. [July 2h, 1997] (Ka); W. H. Bore HSS (Lw). Wichita Co.: M. Reed s.n. (Selkirk, Aug. 13, 1892) (Ka). Wilson Co.: Haller s.n. , (Roper, June 26, 1896] (Ka); W. H. Horr son. [July 12, 1930] (Lw); A. R. Moldenke 878 (Lw). Woodson Co.s Clothier & Whitford s.n, {aug. 29, 1097] (Ka); W. H. Horr s.n. {July 10, 1930) (Iw), sen. [7/11/30] (Lm); E. W. Lathrop 558 (Lm, W--2235099), 698 (N), 963 (lw); A. Re Moldenke . 879 9 (Im). Wyan- dotte Co.: M. Reed s.n. . (Argentine, J June 26, ne 26, 1890] (Ka); E. Le Richards 3220 (lw). County undetermined: Kleeberger s.n. [1876 a Gqe-aila y S. Norton s.n. (Aug. 11, 1902] (Mi); C. A- A. Ripl n. [E. K = sas, , July 20, 1890] (ob—-50797), son. [E. | Kansas, on 19, 1895] (Ob—-50833). MISSOURI: Adair Co.: 2 C. S. Sheldon s.n (Sept. 1, 1883] (Al). Barry Co.: B. F. Bush 210 (wo a8201). Bar- ton Cos: E. J. Palmer 51118 (Lb--50952). Bates @8 COs H. N. Mol- denke 2923 (iw). Benton Co.: H. N. Moldenke 21920 (LW). Boone Co.: Daniels s.n. (Columbia, Jun.-Oct. 1097] (Lb--9608), sen. [Co lumbia, July 3, 1903] (Lb—L182, Lb--l190), s.n, (Columbia, July 1903] ( Lb--187); Favor s.n. (Columbia, Sept. 3, 'Ol] (Lb--189), s.n, (Columbia, June 22, 1902] (Lb--188); Henson s.n. [June 30, 19h0] (N); Rickett s.n. (Sept. 1926] (H—106h66), s.n. [Rocheport, June 19, 1927] (Lb--7096); 0. E. Thomas s.n. [C Columbia] (Je— 5345). Callaway Co.: Schwab _ s.n. {hcCredie, July 192k] (Up). Cam~ den Co.: A. R. Moldenke 858 (Iw). Chariton Co.: Mrs. H. S. Young s.n. [June 3 1925] (Gm). Clay Co.: B. F. Bush 12696b (Bt— 15196); ¢ C. J. Elmore s.n, (Liberty, Aug. 25, ~3923) Lit srageard Gier 5132 wor K. K, Mackenzie 261 (S). Clinton Co.: C. R. Barnes San. (Converse, ~6-l-77] (Pu). Crawford Co.: A. Re Moldenke 848 (im). Franklin Co.; D. L. Marsh 1663 (Lw); J. M. Mason 8.ne , [Mer- amec State Park, 7/15/35] (Lb--34371); A. R. Moldenke 847 (L™)- Greene Co,: Blankinship s.n. sen. (Willard, July 9, 1919] (Po—638)8) 5 J- I. Cantrell s.n. (Springfield, June 17, 1929] (Lb--1bL83)3 Pe c. Standley 8325 (W—687286). Grundy Co.: Crookshanks 135 (Lb-— pope. a Cos: H. N. Moldenke 21921 (lw). Hickory y COot > « Moldenke 861 (Iw). Tron Co.: L. H. iH. Pammel SMe [Pilot Kno June 17, "B8] (Io—15351), Jackson Co.1 B. F. Bush h26 (C), u75 (C), son, [7-2-1892] (Lb--l,183); Erisman s.n. '. [Kans as City, June 26, 9k] (ah); K. K. Mackenzie s.n. [July 17, 1900] (N)+ Jasper 1965 Moldenke, Monograph of Verbena 227 Co.: Demaree 39337 (Bm). Lincoln Co.: A. R. Moldenke 842 (Lw). Maries Co.: | H. N fs N. Moldenke 21917 (Li). “Marion Go.: J. Davis 2791 sl 2680}; 3227 "(Se—11951), Lhe. (Du—-202359), 6246 (Um—209), FOM (Ur), s.n. [near Hannibal, 9-613] (In), SoD. . [Hannibal, 17- 7-13) (au, Au j Sede (Hannibal, 26-14] (Mi), son. (Hannibal, Uj-7-14)] (Au), 8.n. (Hannibal, 20-7-15] (Gg—216419). Miller Co.: A. R. Moldenke 856 (Lw). Nodaway Co.: J. Re Singleton 106 (Ib— 34596). Ozark Cos: Leopold s 8.n. [10.9.39] (Lb—L8919)3 Ri Rickett s.n, (Rock Bridge, June 26, 1927] (Lb—709h). Pemiscot Co.: A. J.D Davis s.n. (Hamlin, dais 7, 1917] (Se--15988). Phelps Co.s Droust s.n. [July 12, 1931] (Lb—92h7); A. R. Moldenke 849 (Lw). Pike Go.: Frite tchey 559 569 (Ib--18h). Platte Co.: > Radloff 8. gen. [July 2h, 1956] (Ur), son. [July 25, 1956] (Ur). Ralls Co.: J. Davis 40 (Ur), 3779 (Ur). Reynolds Co.: R. L. McGregor ings (Im). “Saint Louis Co.: Collector undesignated one Bon. [Between St. Louis & Iron Mts., Aug. 2h, 1878) (W—221165); M. Craig s.n. (West Richmond, aug. 1912] Passepeauel Prt sen. [Herb. Geete 5709] (Go); Glatfelter s.n. (near St. Louis, 7-20-92] (Mi); Let- terman 14 (Po), s.n. [Allenton, te 28, 1911] (Io--76363), son. [Allenton, July 6, 1911) (or-1h360); He N, Moldenke 21798 (Mi); L. H. Pammel s.n, [Jefferson Barracks, “6-1888) 8) (Io—2291h); Le L. Fe Ward sen. en, [AUg. 22, 1878] (W--117587) . Stone Co.: B. F. Bush Re a. (Oct. 9, 1936] (Ma) 5 Z. Williams s.n, [Hurley, August p27) (Do—12 4191). Taney Co.: De. L. Spence Spencer Tr SoM. (7-25-h0] (Lb—35420, Lb—1,8920), Warren Co.: A. R R. Moldenke 8 bh (Lw). Wayne Co.: E. J. Palmer 6107 (Po—21751h, W—15810L9). Wright Co.: Lansing 3164 316k ). Saint Louis: Prummond sen. [1832] (C); Eggert sen. [12 July 1877] (S); Engelmann 339 (Pr), sen. [St. Louis, July 1039] (S), 8- Re [St. Louis, Aug. 18)1] (W--7193b); Fendler s.n. (Herb. Krause 16832} (B); F. D. Kelsey s.n. (Sept. 189k] (Ob—50838); M- 3.n. (Br, Br); Be H. Patterson s.n. [St. Louis, Oct. 1, okt (on (ca) Fi Rieh1 196 (Br, M). County undetermined: Z. Baldwin sn. (AU); Buckley s.n. (T); Engelmann s.n. [July 1839] (Br, Br)s Geyer Sos (Upper Missouri, July] (C); Pringle s.n. ([Southeaste rm Missouri, 30 July 1885] (Vt); Spillman 298 (Pl—22615). ARKANSAS: Baxter Ot Demaree 20675 (N, Sm), 23606 (Sm), 2931h (N)- Benton Co.s Heacox 5.n. [July 20, 189] (Ob=-5083)). Boone Co.: Demaree 30203 (as—122703, Au--122725, N, Ok, St). Clay Co.: Demaree 27030 (N)- Conway Co.: Demaree 37218 (sm). Craighead Co.: Demaree 5 5085 (0k), 27113 (Au—-12270h, N, Ok, St). 30772 (N). Faullmer Co.t F. A Haas 1673 (W~1651392), Franklin Gos + Ruth 31 (W—1522032). Garland Cows Scully 33h (Ky). Green Co.: Demaree 26698 (Au--122712, N, Ok, St). Hot 5 Hot Spring Co.: Demaree 2760 Tw), LOSE 4 Be R. E. Earle 3. Re (Malverne, July 19, 1577] (N). Independence Co.: Denaree 23800 228 PHYTOLOGIA Vol. 11, no. (Sm), 26792 (N). Izard Co.: Demaree 2275 (Sm), 23376 (Sm), 23378 (Sm). Johnson Co.: Demaree 19995 (N, Sm). Lawrence Co.: Demaree 25748 (N), 26152 (Au~122738, N). Logan Co.: Demaree 17723 17723 (N, reg 2519 (Ba (Bm); Pyle 928 (De); Welch 1121 (Dp); Mrs. g. Si Young s.n. [June 16, 1925] (Cm). Marion Cos: Demaree 29151, in ait Tau-=i25707, N, st), 30032 (N), 30909 (Au-~122706, Le), 47937 (Lw). Mississippi Co.: McAtee 2053 (N, W--587275) - Phillips Co.: Demaree 1522), (Du—256558, Gg--267615, H--97679, Ok), 152bh (Ky, N, Pl--131530, Ur, Ur), 19251 (N, Sm). Poinsette Co.: Demar- ee 17568 (N, Ok). Pope Co.: Demaree 19871 (Mm, N, Sm). Pulaski Cos: Coville 3 (W--210602); Demaree 8365 (Bt—-11509, W--158239k) 5 F. W. Pennell 1065), (N). Randolph Co.: 0.: Demaree 29179 (N), 30963, in part part (Au--122705, Le, Ok); R. D. Hamilton s.n. . [Current River, 8/2/38] (Cm). Searcy Co.: Demaree 22251 (Hi--170690, Lb--53h1h, N, Sm). Sebastian Co,: Bigelow sen. [Fort Smith, 1853--h] (#— 71933) ; Rolfs s.n. [Ft. Smith, Aug. 1891] (Io—32271). Sharp Co.? Demaree 26270 (Au--122737, N). Stone Co.: Demaree 23495 (Sm), 27631 (N). Van Buren Co.: Demaree 2227) (N, Sm). Washington Co.: Braly sen, [Fayetteville, July 19, 1893) (St); Demaree 2235 (N, Sa), 22h53 (Lb--5315); F. L. Harvey 61 (Cm, Ca, Mi, , Mi, Ur), 262 (Pa), Son. oe Kay} “(Ur); Nix s.n. [June 7, 1893] (Ok). Coun- ty undetermined: . Harvey sen. [Curtiss 1958] (Be, C, Cm, Pa, Vt). MONTANA: at * ae Co.: Blankinship s.n. [Crow Agency, July 1h, 1901] (Fe). Missoula Co.: Kirkwo ae 1822 (Ca——352291, Mn-- 17746, N). IDAHO: Kootenai Co.: J. H. Christ 11832 (N); C. Epling Son. [Post Falls, July 1929] (La); Milburge 969 (Se-—15930)- WY0- MING: Albany Cos: C. L. Porter 7151 (Au—12: (Au——122709, Ca~-39483, Ge— 407318, Mg, N). Laramie Co.: Cantrell s.n. [8/15/47] (Ok) Fe Ve Hayden s.n. (Camp Carlin, July 1570) (Pa); A. Nelson 538 (Ur)- Platte Co.: A. Nelson 505 (c, Cm, Ur, W—-28)50h). Weston Co.: Degener & Peiler 16190 (N (N, N, N). County undetermined: F. V. Hay- den s.n, (Aug. 1 — Sept. 13, 1870] (W—2359L0). UTAH: Utah Co.s Q. Garrett 3172 (S). COLORADO: Baca Co.: Weber & Anderson 520h a--95429, No--23718). Boulder Co.: W. A. Weber 5270 (Wo—236h6)- Cheyenne Co.: Heustis sn, [Cheyenne Wells, , July 25, , 1906) (Bl 42311). Denver Go. Eastwood 90 (B1—233k, Ca-~148062, Ge St—-229)8, W--582335). Elbert Co.: Demaree 29416 (Le). El Paso Coss betgaupt 931 931 (Du--195879); Biltmore Herb. 147588 (s)5 Ferril sen. (Sept. 1, 1902] (Bl—l\2313). Jefferson Co.: Beth “Bethel s [Wheat tridge, Aug. 16, 1916] Eee? 3 Heustis s.n. Faestradge, coete (a3). Kit oo Co.: pa Saar, 3 (Ca--590153, Ae (Ca-~9 1965 Moldenke, Monograph of Verbena 229 1932] (Po--191051). Prowers Co.: Collector undesignated s.n. [Ark valley in eastern Colo.] (Fc). Washington Co.:; Piemei- sel 1576 (W--584372). Yuma Co.: Eggleston 1522) (Dt), 15528 (W-- 1); Ewan 12917 (En); Harrington & gton & Smith 417 (Fe); Pleneisel 1533 (W—58)326); Ramal ey 12768 12768 (Bl--2362). County undetermined ik L. Sheldon s.n, (Engelmann's Canon, Aug. 5, 1903] (We). rn KA: Adams Cos: “a. . Barnhart 468 (N). Buffalo Co.: "W. E. B."s.n. {Aug. 1922] (Ok==10361); J. He Holmes s.n. [Kearney, Aug. , 1889) (Fe~-2505, W—71938). Antelope Co.: Rohrbaugh 41 (Au--12272h). Cass Coot M. E. Day 221 (Ob—99771)5 He Ne Moldenke 21936 (Lm); Je Le Morrison m 1218 (Go (Go). Cedar Co.: Pe . Clements 2 2663 (Ca— 124661, W719). Cherry Co.: Tolstead [15 (Bt--35085). Chey- enne Co.: Osterhout s.n. [north of Sidney, July 7, 1932] (Po— 216659). Custer Co.: C. L. Dietz son. [2h July 1927] (Se— 21475); He J. Webber s.n. - fAnselmo, July 8, 1889] (W--216329). Douglas Co.: | Eastman son, [Omaha, Aug. 16, 1915] (Ur); Kleeberger S.n. [Omaha] (Gg--3120). Greeley Co.: Nolte 99 (I)- Harlan Co.: L. H. Pammel s.n. [Alma, June 21, 1897] (Cm). Hooker Co.: Ryd- berg” i422, in part [near the forks of the Dismal ~Nena Hooker County, on prairie, July 11, 1893] (Cc). Kearney Co.: H. Hapeman Sen. (Minden, July 1, 1833] (Gg--237860), sen. [June 3B, '91] (Hp), Sen. (July 1h, 1930] (En—2509), s.n. (Minden, 1930] eB war Sun. (Minden, June 29, 1932] (Bt—12318, Ua--11390), s.n. [Minden June 26, 1933] (Ca—-51229, Ew, Ew), sen. [July 1, 1933] (I, 14), Sn, [Minden » July 6, 1937] (Sd—-23838). Keith Coss C. 0. a son 2260 (Mi) 5 M. E. Jones s.n. [Ogalalla, June 16, 1925] (Po lt Knox Co.; Krautter s.n. [Niobrara Reserve, July 1903] (Up). r Co.: Herb. Okla. Coll. Women 311 (Ok); Herb. Univ. = 259 29 (ha) Pipal s.n. [Lincoln, 6-9-11] (Ah); Seal s.n. [Lincoln June 30, 1897] (W—2311053); J. L. Sheldon s.n. n. (Lincoln, september 1897] (We); Whited 2686 (Or—12379). Lincoln Co.: F. G. Miller 8. Bs [7-17-01] (Io—2290h, Io-—-22915); Plank s.n. [North Platte, July 1896} (N). Merrick Co.: A. A. Heller sieve Ws (Du--156905, 3e--88013). Nance Co.: L. Be Curtis s.n. [Genoa, June 22, 1903] (Ok--10363) « Nemaha Co.; H He N. _ Moldenke 21934 (Lw); J. L. " Sheldon s.n. (Peru, July 21, 1900] (We), s.n. (Peru, August 1900] (We); Shubert s.n- Peru, July 1933] (Ob=-50798); Vance s.n. [Peru] (Bt—10923)} Je ae M. Winter 20 (W—-1679069). Oteo Co.: A. E. Allen s.n. [Talmage, Ui, 1891] (Je); H. N. Moldenke 21935 (Iw); Wibbe s.n. (Salina, sae 1882) (Du--91129). Phelps Co.: McMillan s.n. [Holdrege] (Io--53117). ee Cos: V. L. Harms 581 (Lw). Platte Co.: Clevenger 5-n- [Co- Se e 21, 190h] (La). Richardson Co.: Grinstead 85 Med. He oi.denke 21933 Lw). Scotts Bluff Co.: F. G. G. Miller s.n. [Aug. 8, 901] ie amie) Sheridan Co.: R. E. Buchanan sn. [Spade 230 PHYTOLOGTIA Vol. 11, no. & Ranch, August 2-20, 1905] (Io—-6519); J. B. Hatcher s.n. [on the Running Water, July 1--10, 1886] (Ca—25173), sn. [July 1886] (Pr). Thomas Co.: Krautter s.n. [Halsey, July 23, 1907) (Up— 45720, Up—5S7h2, Up—L5743, Up—-hS7hk, Up—l577) ; 1422 (C, W--210363). Washington Co.: Conard $.279 (Se--19993)3 Lall- man s.n. (July 5, 1931] (Bt--32235). County undetermined: EB. K. Abbott s.n. (Gg—31)25); F. E. Clements s.n. (Nebraska, 1893] (W— Ti9h2); F. V. Hayden s.n, [Bad Lands, 1856] (Dt); LeRoy s.n, [Ne- braska] (C); R. de Lima 101 (Oregon Trail] (I); McClatchie s.n. [Endell, July 1889] (N); B. H. Patterson s.n. [July 6, 1895] (Cm). OKIAHOMA: Adair Co.: C. S. Wallis 2367 (St). Atoka Co.: V. Le Cory 56792 (St). Beaver Co.: Hindman 234 (St), 254 (St). Beckham Co.: B. Osborn 1368r (Ok). Bryan Co.: W. S. Blain s.n. [Durant, June 23, 1935] (Bt—29681). Caddo Co.: C. T. Eskew 1089 (Ok); Edw. Palmer 243 (C, Du--177391), 24h (T, W); Van Vleet 34 (Ok— Houghton s.n, (Stevens 000] (N). Cleveland Co.: Babb 108 (Ok— 7~3-19] (Wi); Demaree 13077 (Ok— Flowers 67 (Ok)3 George 29 (Ok); 22, 1926] (Ok); Personett 52 (Ok—1033h); Stemen 136 (Ok); C. Wha- ley 22 (0k-~10 che Co.: M. K. 11748 (Ba), 17h8a W. Stevens 2930 (Ok). Garfield Co.: Gephardt 769 (Ok). Garvin Co? St—9266, Ur). Harper Co.: Stratton 03 (St-~26129). Kay Coe? Me W. Barkley s.n. (July 1929] (Ok—13759); Davy son. (July 11, 1939] (St—1755); G. W. Stevens 1865 (Ok, St-—9262); J. J. Taylor 18 erate (St~17552). Latimer Co.: 0. M. Clark 200 (Ok—10318). Leflore Poot ces Sen. [G. W. Stevens 157] (Du—6630, Ok, St-—9269, Ur, W-589571); "R. E. J." s.n. [Winding Stair Mts., 6/30/19] (Ok— 1965 Moldenke, Monograph of Verbena 231 10335). Lincoln Co.: 0. M. Clark 2245 (B). Logan Co.: C. C. Smith G.689 (Ok—16362). Marshall Co.: G. J. Goodman 6475 (Bs, Ok, Ok, 2); Kelting 181 (0k); P. B. Riggs 1981 (S)} Stark Starks 3.n sen. [8 July 1950] (Ok). Mayes Co.: Flowers mers 427 (| (Ok) Ce Se Wallis 3348 (St), 767k (Au—171439, St). McClain Co.: Harding 553 (St- 20329); Me M. Hopkins 1336) (Ba, Ok—20523, St—12770); ¢ C. C. Smith 550 (Ok), 809 (Ok); Stratton 4770 (St--26143), 4771 (10, N, St— 261b4, St—261,5). McCurtain Cost Puckette 38 (au--12271h) « Mur- wd Co.: Emig 42 (Cm), 787 (Cm); C. T. Eskew 1019 (Ok); M. Hop- kins 1168 (Ok), 5207 (0k), 5u10 (Ok, Ok), 6023 (Ok); Hopkins & Cross ; 6450 (0k); Hopkins, Nelson, & Nelson m 659 (Ok); McLennan 122 (St—2h885); Merrill & Hagen 599 (Ok--19976)5; G. Mitchell ell 505 pos. Rutledge & Gage 561 (Ok--10333); Stratton 4730 (N, 730 (N, St--26139, t-—-26110). Muskogee Co. R. Bebb 5017 (Ok, Ok)3 EB. L. Little 161 fais le 1583 (Ok—10325), 1935 (Ok—1032h) ; W. Purcell cell 63 (Ok); C. S. Wallis 458h (St); Waterfall 10269 (St). Oklahoma Co.: A. R. Moldenke 113 (Fg); Waterfall 1403 (Ok, St). Okmulgee Co.: Autry 81 BI (St). Osage Co.: es 2 29 (st--17563) 3 eisute 2 272 Miller 62 (St—-2)972); G. W. W. Stevens 2308, in part (Du-86376), h.2312 (St—~9261). Pamee Co.: a — 2h9 (St--26125). Payne Co.: S. E. Myers s.n. (Stillwater, Aug. echt (Ca—882821); Waterfall 11) (St). Pittsburg tt =—_ Sen, [near McAlester, July 19, 1942] (Rf); McClary 58 (Ok— Tease Pontotoc Co.: Duffer 123 (St); D. McCoy 720 (st), 906 (St), 1717 (st); G. 7. Robbins 27h2 (Ca—966176, N, Ok), 2 (ca- 90434, Ok) ; Stratton 6169 (Hi--152269, M1, N, S St——26161). Potta- watanie Cos: E. D. Barkley 2h9 (Ok—-16731), 1485 (Mn—28630)5; Fe A. Barkley 1,85 (Ca—86201)); Deumer 1g (Ok); Hopkins & Demaree e 8 (0K); Jacobs 19 (0k). Rogers Co.: Waterfall 145k (N, St). Sequo- yah Co.: G. S. Wallis Uk89 (St). Tulsa Cow: M. Be Clark 366 (St)s Force s set 1s (6/25/28) (Ok—-10323); Gatti 689 (St--9259)5 Markley 71 (Se—69272 )3 Prewett 5 (Au); Stoneman s Sen. (Sept. 2h, 1938] (st— 14167); Tenney san. [June 22, 1931] (1)- Washington Co. D. Mc Lean 27), (St--17802). Woods Co.: P. J. White s.n. (July h, 1900) (In). Woodward Co.: J. P. Kimball s.n. [Fort Supply, June 22, 1891] (Ka); L. F. Locke 15 (W--1582636)- County undetermined: R- Bebb 4376 (Ok); Clifton 6 (Ok—10328); G. M. Merrill 625 (N); Saw- yer | & Schultz s.n. s.n. (Ok--10330); G. W. Stevens 575 1/2 (Ok), 2312 (ce), 4.1457 (Ok). ‘TEXAS: Cook Go.: Straudtmann s.n. [July 12) (Au). Dallas Co.: Boll s.n. [1871] (cb, Cb); Jj de Reverchon ia Regen May 1877] (Bc); Mrs. J. P. Stephenson s.n- (Ennis, ref 1928] (Id). Gonzales Co.: O63 Tharp | s.n. [Palmetto to State Park, / 26/3 39] (au, Au). Grayson Co.: H. Gentry 316 (Au-~122652); Hen- 232 PHYTOLOGIA Vol. 11, no. nen 221 (St). Hemphill Co.: Cory 50269 (Au, Ca——75),836, N, Sa, W--1885937) ; C. M. Rowell 4115 (Au—192086); Tharp 4502 (Au). Lib- erty Co.: Warner s.n. [Liberty] (Hu). Lipscamb Go Cot we H. Howell 6h (W-500189). Polk Co.: D. Lynch sen. [August 8, 19h7] (Au— 122699). Red River Co.: E. Adams s.n. [6-19-37] (Nt). Tarrant Co.: Ruth 162 (11), s.n. [Fort Worth, Aug. 27, 1909] (Po—267629). Wheeler Co.: et We P. Taylor s.n. sen. (Aug. 1, 1941] (Au). County unde- termined: M. Re Stephenson | 201 (W--1580539) - NEW MEXICO: County termined: Mrs. 0. Saint John s.n. (Uplands, Catskill, Aug. undet 1895] (Lb~-9608). ARIZONA: Coconino Co.: B. L. Wagenknecht 1,809 (Lw). WASHINGTON: Busser Co.: Kreager 75, in part (Ga-—002627, N, Pl—22578, Se—68370, Um--208, W—Lh1291) . MEXICO: Guerrero: F. A. Barkley 14079 (Au). + ere Belgium: M. Martens s.n. Thort. insul. 1827] (Br), sen. [h. b. lov. 1839] (Br). Denmark: Herb. Mus, Bot. Stockholm s.n, [Ex horto Havn.] (S). England: Stead sn. [Liverpool Garden] (T). France: Herb. W. H. Harvey S« n. [he R. P. 1820] (Du--166k)5). Germany: Collector undesignated 3.n. (Gg—3115); Herb. Martius s.n. (Br); Rodrian s.n. [botan. Garten Heidelberg] (La). New Jersey: G. H. M. “i. Lawrence P.528 (Ba); Lawrence & Lawrence P.2h0 (Ba). Pennsylvania: Martindale Sen. (Germantom, Sep. 1076] (W—-771850); T. Meehan s.n. [1885] (Up—17125). Spain: Herb. Hort. Bot. Matrit. "Sn. (Cp). Sweden: Herb. Beurling sen. [182] (S), s.n. [18)4] (S). Switzerland: Herb. Hort. Bot. Basil. sen. [1850] (M). LOCALITY OF COLLECTION UNDET : Collector undesignated 2109 (S), son. (S); Bngel- mann 8.n. Hah bor.] (Br); Friesner 8.n. (Bt~—2h,0h) Geyer 0b (Br); Herb. Torrey s.n. (North Fork of the Platte] (T); D. Hough ton s.n. [Falls of St, Anthony, July 1832] (Mi); James s.m- [Long's Ist Exped. ] (C); Werner s.n. [Aug. 30, 1885] (Ob——9729h) 3 As A. Wood Sn. (Pa). MOUNTED ILLUSTRATIONS: C. A. Ripley sen. [in color] (0b); Curtis, Bot. Mag. 5: pl. 1976 [in color]. 1618 (N). VERBENA STRICTA f. ALBIFLORA Wadmond, Rhodora 3k: 19. 1932- : Verbena striata var. alba Popenoe ex Moldenke, Suppl. bena stricta alba Gates ex Moldenke, Résumé Suppl. 7: 10, in sym- 1963. Verbena suricte f, alba Wadmond ex Moldenke, Résumé Suppl- 10: et) Bote syn. pecan N. F. Petersen, Amer. Bot. 5: 166--167. 1939; Bebb eh , Prel Alph. List Invalid thes 48. 1940; Moldenke, Suppl. List Invalid 1965 Moldenke, Monograph of Verbena 233 Names 10, 1941; Moldenke, Alph. List Invalid Names 58. 192; Mol- denke, Known Geogr. Distrib. Verbenac., [ed. 1), tg ee Pe ER : we 19h2; oo Castanea 10: 39. 195; M . a2: 7h & 117. 1945; Moldenke, pipe List C bite a: 109, 217, 22h, Ohl, & 267. ib, Fs wicinemges , Phytolog ia 2: 383. 1947; Moldenke, Alph. Steg oe oy Suppl. 1: 26. ll Mol ete , Alph. List cit. 2: 391, 00, » 406, 51k, 517, 520, 2h, 605, & 620 goo and 3: "800, "826, "90h, "927, & 949. 1949; Moldenke, di gia 3: 131. 1949; ‘Moldenke, Known Geogr. Distrib. V cna [ed. 2], 12, , ay 20, 21, & 199. 1949; M. L. Grant, Proc. Iowa Acad. sei. St: ¢- 19505 be Revist. Sudam. Bot. 8: ee 1950; + Gray oPetss ad. be 1211. 1950; Moldenke, lagi ie 7. 1935 | Moldenke’ in Gleason, New Britton & Br. Illustr. Fl., print. 1, 3: 129 (1952) and print. te 3: 129. 1958; Moldenke, 1 umé . : , 6 2 (1963), and er bed 1963; Moldenke, Phytologia 9: 506 & 220. 1963; Steyerm vee pat hi: 3. 1964; Moldenke, Phytologia 11: 117 & This form differs from” the typical fom of the rp te i "in hav- ing its corollas purplish-white, nearly white, or pure- The type of the form was collected by Samee] Christiensen ‘Wai- mond in southeastern Wisconsin. In the original description of He says te oaitreie en oe seen only once, along roadside where was interspersed with the normal urple-flowered form." Other sandy = Faint pastures, sandy waste places, ditches and open Toadside ditches. low ow marshy areas along small creeks, and sand of hog-lots > along rivers, roadsides, open roadsides, and rail- Toad tracks, and on dry bottoms, hilly prairies, bare bluffs, and Sand pene to 800 feet altitude, flowering fran June to Sep- tember, and fruit ting from June to August. Materi rial has been identified and distributed in herbaria as V. Stricta Vent. and as "Verbena stricta x --". The Cing-Mars S.ne (32 juillet 1951] and A. S. Hitchcock s.n. (Ames, July '82] are mixtures eins typical Ve stricta Vent. Common names recorded for the plant are "hoary vervain" and "white-flowered verbena". my 190 aohik oan I followed Perry (1933) in regarding it as Synonymous with V. stricta, but I now regard color pssisio such as this as worthy of distinct nomenclatural recognition Verbena hastata x stricta Allen is apparently based on Ae Ee Allen s.n., collected at Per Peru, Nemaha County, Nebraska, on August save S208 S.n. collected at Topeka, Shawnee County, Kansas, on July 23, 79, and deposited in the herbarium of Iowa State College at 23k PHYTOLOGIA Vol. 11, no. & Ames, Iowa; V. stricta var. alba is based on G, D. Fuller 1123, collected in an open [ooen alan © forest in Earl Township, Section on 21, La Salle County, Illinois, on September 10, 1918, and deposited in the herbarium of the University of Illinois at Urbana; and V. Tllinois Central Railroad tracks southeast of Havana, Mason Coun- ty, Illinois, on July 6, 1910, and is deposited in the herbarium of the University of Illinois. Hayden says "a colony of pure white flowered plants was grow same state, Chase says "locally not rare in sand prairie”. ansas Horr reports "about 50 plants growing with V. stricta in loam with fair moisture" and mot common in dry loam, ¢ grass- form in overgrazed sandy pasture" and a "colony of white form in prairie pasture ravine”. Poindexter 201-68 was found in a broad ravine below a pond and peoret to have 95 percent pollen fertility and a chromosome count of n e notes that in South Dakota "about 1 in every 100 was white". Terrell says "no color transition; with and outnumbering the he form in sandy waste places by roadside” (in Wisconsin), S reports “large colony, 25 percent white, 75 percent deep purple” (in Oklahoma), and Demaree notes "all these were pe many others were blue" (in Arkansas). The Davidson, Dixit, & Rom beng 4689, cited below, is described by the collectors as 3 as "flow ers white and lavender" and so is probably a mixture with f. ros~ eiflora Benke. Truman (1938) says: "In the summer of 193) the writer observed a very few plants of Verbena stricta Vent. forma albiflora Wadmond = several miles to the west of the station reported by Wadmond. + in 1932. Contimed observation of the colony during the summer of 1935 and 1936 revealed the fact that the white-flowered form was creasing in numbers and spreading to the adjacent roadside. In 1937 it was noted that not only had this form increased in its im portance in the original station, but also that several specimens had appeared in a farmyard a half mile family living upon the farm where the white-flowered form was first noted remember that in 1933 the station consisted entirely of purple-flowered plants. Their observation is not reliable, how- ever, for in 193, the relative number of white-flowered plants was so small as to be evident only to a practised observer. a "This form differs fran the typical V. stricta not only in tha’ its flowers are pure white, but also that its stems completely lack 1965 Moldenke, Monograph of Verbena 235 the . color that is invariably characteristic of the purple- flowered form. These differences seem to indicate that some change has taken e in the genetic make-up of the plant to in- hibit the vaiaeart of the pigment, presumably anthocyanin, which gives the typical form its color. In all other respects the white-flowered form resembles the typical, although in a doz- ed seeds, the white-flowered plants were all taller, sturdier than the purpl red. s difference may ha purely accidental. At no t r years ch this appeared any intermediate form. All individuals have been either typical V. stricta, or pure white. It is interesting to note here that this lack of intermediates has been observed by Benke.....-+ where V. stricta Vent, forma roseiflora Benke grows with typical Tm The corollas are reported as twhite® on Clinton s.n., 276888, Herb. Univ. Nebr. 364, Kellerman s.n., Over 11386 » Seymour Sone, | and Say Smyth h 3061; as "pure emer ai on Ve H. Chase 7852, & : ar Sheldon s.n.; and as on OW St on Grace +s Se ~ scone (1963) records this form from Daviess, New Madrid, ee Louis, and Stone Counties, Missouri, on the basis of speciinens p smeek in the herbarium of the Missouri Botanic examined Citations: QUEBEC: * Deux-Montagnes Cos: pe fio sen. [31 juil- let 1951] (Mg, Vi). ILLINOIS: Cass Co.: Seymour sen. [Chandler- e, July 6, 1886] (Ur--30015). Henderson Cost He “He Ne Patterson S-n, [Oquawka, August 1872] (N), s.n. [August 1873] ] (Al). Lasalle na, June 21, 1894) (Ur); F. C. Gates 3608 (Ur). Sangamon CO.: Ailes 11592 (Ur); Winterringer 7333 (11--36L65). saree Ve Ee Chas se 7852 (Ur). Wabash Co.: Schneck s.n. [Mt. Carmel, July 1885] Ch (Ur). Winnebago Co.: Wadmond s.n. [Roscoe, Aug. a 1931] (Ur). IOWA: Butler Co.: Isely ly 6508 icersaeetenes ea kinson Co.: A. Hayden 3001 (Io—-149002, N). Harrison Co.: V. L. Harms (Iw). “Story Co.: A. S. Hitchcock s.n. [ames, July a3 2] (1015322). WISCONSIN: Crawford Cos: Terrell 2415 (Ws). LaCrosse Co.? Hartley 264). KANSAS: Barber Co.: Re L. MeGregor “1600 (Lw); J. = Poindexter 201-68 (Iw). Comanche Co.: R. L. McGregor or 1,578 (Lm 236 PHYTOLOGIA Vol. 11, no. 4 1,679 (Lw). Decatur Co.: V. L. Harms 1276 (Iw). Greenwood Co. J. En Engionan 1079 (Ok). Marshall Co.: W. H. Horr 4556 (Lw). Mont gomery Co.: he L L. McGregor 15052 (Lw). ~~ Osborne Co.: Shear 1h9 (ca-~882822, “W--227296) Republic Co.: G, E. Morley 1302 1302 (Iw). Riley Co.: Kellerman s.n. Be ne 6=30-1887]} (Ka). SI Shawnee Cos: E. A. Popenoe s.n. (Topeka, July 23, 1879] (Io—10912), 8. n. (Topeka, July 2h, '79] ar er B. B. Smyth 3061 (Ka— 79497) . Washington Co.: W. H. Horr 1622 > (Lw). MISSOURI: Saint Clair Co.: Steyermark 82192 (Lb--L931). ARKANSAS: Stone Co.: Demaree 27888 (N), 2 278888 ( (Ok, St). NEBRASKA: Custer Co.: Herb. Univ. Nebr. br. 36h (Au). Grant Co.: Rydberg 1422, in part [in the Lake Region of Grant County, 2 miles NE of Whitman, Central Neb- raska, Aug. 1, 1893] (W~21036h). Lancaster Co.: Js L. Sheldon 8.n. (Lincoln, 8.11.98] (We). Nemaha Co.: A. Ee Allen s.n. [Pe- ru, Aug. 2, 1893] (Je). Thomas Co.: Krautter son. (Halsey, July 23, 1907] (Up—iiS722) « Valley Co.: Davidson, Dixit, & Romberg 4689, in part (Au--122711). OKLAHOMA: Cherokee Co.: Ce oy = 79h (St), 3395 (Au—122708, Au—-122710, St). Logan Co. 363 (Ok—19027). Marshall Co.: Alder s.n. [July 10, 1950] 7 (OE) « ee Pisin f. ROSEIFLORA Benke, Rhodora 34: 10. fie graphy: Schaffner, orm — ade Pa. 06 5 Rhodo fina me 10, 19323 Perry Os Gard. 20: 56, 287, & 356. 1933; Truman, Rhodora |0: : 28. i938; "F. C. Gates, Fl. Kans. 191. 190; Moldenke, Known Boe Distrib, Verbenac., [ed. 1], 6, » 9, & 102. 1923 Moldenke, Alph. List Cit. 1: 200 (19h6), 23 361 & 92 (1948), and 3: 827, 927, & 954. 1949; Moldenke, Known rib. Verbenac., [eds 21, = 13s 37; onion & reigned Proc Ac ‘Oo print. Phytologia s a Tost) a and i 185. - 19535 Cece in Gleas New Britton ky Illustr, Fl., prin » 3: 129. 1958; ecidecke: Résumé 6. 29; 2h. 20, aby & L7k. 19595 “foldenke, Résumé Suppl. 2: 2 & 3 (1960), a 7 (1962), and h: 3. 1962; Steyerm., Fi. Ho. 1260. 1963; Moldenke saiewh Suppl. 10: 1. 19 by » form differs fran the typical form of the species in having its corollas light~pink, pink, light-rose, rose-colored, rose~red, or roseate to lavender The plant is described as Qn} feet tall. form was collected by Hermann Conrad Benke Oe: Ne as one Concor~ dia, Cloud County, Kansas, on July 2h, 1929, “and is is deposited flora -- very luxuriant in the valley. About Concordia, among : most common plant life is Verbena stricta Vent., acres in exten- » however, is it to be classed as a weed, for it thrives on= 1965 Moldenke, Monograph of Verbena 237 ly in limited areas and at limited times, certain soil conditions ience lies to the northwest of the city. Here, in a field of creek bottom some twenty acres in extent, was found an almo solid mass of the species in the usual purplish-blue, = lesser of th pa s d rose-colored form. This variation in color made a very ¢ haved impression in t = Landacapets features. With no intermediate The form has bets Fail along roadsides and open roadsides, loam soil, dry or sandy tom, black loam and loam with fair soil moisture, on prairies, and along railroads, in grasslands, grass- land pastures, severely overgrazed creek valley pastures, and ra- ring and fruiting from June to Benke says that "in a field of sane 20 acres about a pron of the plants were of this color form" in the same state. Also in ve creek" (Bee - 464), "one or two plants found with very light cor- n loam over sandstone soil in grazed prai oe 6), and "one large plant of this very light color found" no. 396). ee Benke. Grant (1950) records it from Dic eee County, L. Kucera, in a letter to me dated February 2, 1955, sbatye "that there are no specimens of this form in the University of Missouri reo agi and that he cannot recall ever having seen it in that The flowers are described as "pink" on Horr 1,660 and Spillman S:Bej as “"light-pink" on Seymour s.n.; as as "light-rose" on McGregor I5T70; as "light-rose or pink" on Harms 1313; as as "roseate" on Benke 5164 and Fernald & Pease 3496; as as "lavender" on Porter 34135 and as "white and lave eee aan on Davidson, Dixit, & Romberg 609 erg 4689 (probably a mixture with f, albiflora Wadmond) . Tks § 2 sai " addition Benke 5164 (G--isotype), not as ma mi all, 2) herbarium LEP including the type collection, ve been examin by m Citations: ILLINOIS: "TaSelle Cos: G. D. Fuller 22998 (I1— 238 PHYTOLOGIA Vol. 11, no. 15801). Marshall Co.: V. H. Chase 11151 (Ur). McHenry Co.: Ben- ke 6150 (N). Sangamon Co.: Seymour s.n. [Springfield Junct., July 8, 1879] (Ur—176h). Vermilion Co.: Ahles 8092 (Ur). MICH= IGAN: Dickinson Co.: Fernald & Pease 3496 (Mi). WISCONSIN: Rock Coo: E. W. Fell 57-82) (Ws). We KANSAS: Cloud Co.: Benke 516), (N— isotype, W—1515910—isotype). Jewell Co.: W. He Horr L77h (1m). Kiowa Co.: Re L. McGregor 15170 (Iw). Morris Co.: R. Le ee 4356 (lw). Republic Co.: G. E. Morley 376 (IL), 396 (Iw), 464 (Lr), 79h (Lw). Russell Co.: V. L. Harms 721 (Iw). Smith Co.: V. L. Harms 1313 (Lw, N). Washington Co.: W. H. Horr 4660 (Lm). MISSOURI: County undetermined: Spillman s.n. [s.w. Mo., Aug. '89] (Pl—22614). NEBRASKA: Valley Co.: Davidson, Dixit, & Romber 1689, in part (Ur). WYOMING: Weston Co.: C. L. Porter 3413 (Au). OKLAHOMA: Cherokee Co.: C. S. Wallis 65 (St). A STRIGOSA Cham., Linnaea 7: 256—~257. 1832 [not V. strigo- sa Hook., 1836], Bibliography: Cham,, Linnaea 7: 256--257. 1832; Steud., Nom. Bot., ed. 2, 2: 750. 1813 D. Dietr., Syn. Pl. 3: 602. 18133 Walp., Repert. ks 236 1845; Schau, in Ae We, : 5. ° 1847; Sc Fl. Bras. 9: 186, 1851; Jacks. in Hook. f. & ks e « 2: 1179. 1895; Dusén, Arch. . an. 13: 36 & 9h. 1903; Hayek, Denkschr, Kaiser. Akad. Wissensch. Math.~nat. 79 (1): 29h. 1908; Moldenke, Kn . Dis 1), 39 & 102. 192; enke, Alph. List Cit. 1: mac., [ed. 3 1942; Mold 289 (1965 and 2: 598 & 62h. 1948; Moldenke, Castanea 13: 117. 1948; Moldenke, Alph. List Cit. 3: 688, 840, & 921—923 (19h9) and 4: 1249. 1919; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2), te. ib @, Phytologia 4: 188. 1953; Rambo, Sellowia 7: 260 & 268. 5 292. Small erect shrub, much branched, softly strigose-hirsute and incanous throughout, varying to subsericeous or very rarely almost bt acutely serrate with a few teeth toward the apex, entire and sub- cuneate at the base, 3—nerved at the base, the venation impressed above, prominulous beneath, the midrib usually pinnate above; florescence borne at the apex of the branches, single or ternate to quinate; spikes usually terminal, 1—20-flowered, mostly fer 1965 Moldenke, Monograph of Verbena 239 flowered, rather lax, solitary or densely cymose to racemose- culate on many axillary branchlets; bractlets broadly ovate, Berlin, where it was photo- graphed by Macbri s type photograph no, 17448, but is now destroyed. Walpers (185) places the species in his Section Ver- benaca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Secondary Subgroup Holophyllae with 22 other species. Schauer (1847) says "Praecedentis [V. hirta] formis parvifoliis quodammodo similis, sed jam calycis dentibus corollaeque barba magis conspic- ua discernenda. Variat foliorum figura et magnitudine neque P barba magis conspicua discernitur. Variat foliis basi magis min- usve attematis, obtuse et acute serratis, neque minus indumento 9) campos, and rocky crests, in grassy places, shrubby fields, bogs, and pinewoods, fran 750 to 2100 meters altitude, blooming in April, May, August, fruiting in i and October to February, t December. Rambo collected it in a region of 2 meters rainfall snows degre temperatures, Dusén (1903) says that it grows "In silvula ad mar- xutha Lem. The Dusén 313, distributed as V. strigosa, is actual- ly Ve lobata Vell. Rambo 1.9319 is said to have had blue flowers, and Hatechbach 216 violet ones. in Schauer (1817) cites Riedel s.n. and Sellow s.n. from S&o Paulo th trent erwin we cin _—_—_— a and Berlin herbaria, while Hayek (1908) cites —acket S.n. and Wettstein & Schiffner s.n. from the same state. Pm all, 30 herbarium specimens, including the type collection, 9 mounted photographs have been me. Citations: BRAZIL: Parand: Dusén 7969 (S); Hatschbach 4216 (0k), 24,0 Poot 0:1 0-025" Vol. LL, no. W——181970 ei 7ic kobinemn s.n, (Villa Velha, Aug. 28, 1939] (N, SP 41551); Rambo ‘.ea0R (N). Rio Grande do Sul: Rambo 30981 (N), 32450 (N), 34728 (S), 49319 (S); Sehnem 5155 (B). Santa Catarina: Reitz 3280 (N), 340k (S), $), 3u17 (N); smith & h & Klein 8101 (0k, ¥-—- 225134), 11107 (W—2251751, Z); Smith & Reitz 9994 (W—-22h936), 1002 (Ok, W—-2251638), 10163 (W--229306). S&o Paulo: Herter 1,360 (N); Sellow 4939 (Vt), s.n. wn. (Brasilia; Macbride photos 174)8] (Br— isotype, F--photo of Sectype, Rescplate of isotype, pcg of isotype, N--isotype, N—photo of type, N--photo of isotype, photo of type, "hai of porn Vt—-isotype, Z--photo of fied Z—-photo of isotype). besa” SUBPALUDOSA Malme ex Angely, Fl. Paran. 17: 7, nom. nud. 1 Bibliography: Angely, Fl. Paran. 17: 7. 1961; Moldenke, Résu- mé eee 7: 5. 1963. othing whatever is knowmm to me about this species except that it ri recorded by Angely — ‘biex, Brazil. I cannot find that Malme ever published the VERBENA SUBULIGERA Greene, Pittonia 1: 156. 1888. Synonymy: Verbena subli gera Greene ex oldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 19 & 102, sphalm. 192. Bibliography: Greene, Pittonia 1: 156, 1888; Durand & Jacks., Ind, Kew. Suppl. 1: 51,1906; Perry, Ann. Mo. Bot. Gard. 20: 2h9, [ea Moldenke, Alph, List Cit. h: 1292. 1949; Moldenke W7he 1959; ae Résumé Suppl. 2: ly 2 13, & 15. 1960; Moldenke, 36h» maha Tease: procumbent, sute- pibsanien x lence dette bik Scaioothe, the blades ovate, 1.5-—- sare long, gradually narrowed into a subpetiolar base, pinnately cleft, appressed-hirsute on both surfaces, especially beneath on the somewhat prominently reticulate venation, the larger segnents in- cised or sharply dentate; spikes short~pedunculate, elongate, com spicuously bracteate, interrupted in fruit; bractlets lanceolate~ subulate, twice as long as the calyx, ascending-spreading, reflex~ ed in age, hirsute; corolla-tube scarcely longer than the calyx; t about 3 mm. long, pubescent, hirsute on the vena~ carp; Anger subcylindric-trigonous, about 2 mm. long, only faintly striate, he commissural surface muriculate or snoothish, not reaching egy top of the nutlet. The type of this little-known endemic species was collected by A. Forrer in the Sierra Madre west of Durango, Durango, Mexico, at tan ktettuas oe 8100 feet, in in September or October, 1881. The specter has been found in overed mountains and on pine-c ountaintops, at altitudes of 2/00 to 2700 meters, flowering 1965 Moldenke, Monograph of Verbena 21 and fruiting from August to October. Material has been misiden- tified and distributed in herbaria as V. bracteata Lag. & Rodr. Perry (1933) cites the following 3 additional specimens not as yet seen by me: MEXICO: Durango: Forrer s.n, [Sierra Madre, Sep- tember and October 1881] (D--isotype, F—-isotype, G--isot; . The isotype which she cites from "NY" is actually in the ermine University herbarium, She notes that "Although the general of this species is much like that of V. bracteata, the latter differs in having obtusish rel eee and riage! reticulate- eeaets nutlets with commissural faces fully as long as the nut- t." In all, 9 herbarium specimens, a type material of all the 6 names ’ involved, have been examined by m Citations: MEXICO : Durango: Rorrer Sone feted Madre, Septem- ber and October 1881] (C--isotype, —-170277—-isotype, I-~iso- type, I--isotype, W—1)659. BO6GLeotoee, W~-11:65961--isotype) ; Waterfall 12703 (Mi, St), 15504 (st). XVERBENA SUKSDORFI Moldenke, Phytologia 2: es 1946. Synonymy: Verbena officinalis x prostrata Dermen, Cytolog 7: 170, 1936, Verbena suksdorfii Moldenke in Chittenden, a Hort. Soc. Dict. Gard, 6: 2209 & 2212, sphalm. 1951. Verbena iasiostachys var, septentrionalis seclidaiiee x V. officinalis L. ex Moldenke, Résumé 368, in syn. 1959. Verbena officinalis L. x V. dastostaciys var, septentrionalis Moldenke, Résumé 371, in syn. 59. sii aaa 5 Bibliography: Dermen, Cytologia 7: 170. 19365 ath @, Phyto~ logia 2: 150-152. 196; H. N. & As Le Moldenke, Pl. Life 2: 8h. ous Moldenke, Alph. List Cit. 2: 195 (1948) 8 and hs sophe 19495 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], & 199. al. N. e, ? 368, 371, 422 » & k7h. ae Moldenke, Phytologia 8: 121. 1961 Moldenke, Résumé Suppl. 3: 1. 1962; *Moldenke, Phytologia 10: 210 & 211, 1964. en sae a natural or artificial hybrid betreen oS icinalls | L. “ais V. lasiostac var. septentrio lo ke, with inter eee oe with intermediate characters; stems rather ape paherong ly puberulent wi th longer whitish forward-pointing hairs; princi~ Pal ro ape ae mostly elongate, 3--8 cm. long; leaves decussate- 2h2 PHYTOLOGIA Vol. 11, no. te, usually with clusters of smaller ones on greatly a viated branches in their axils; nodes plainly annulate; a teoens , hable inting bulbous-based hairs, more densely and lastingly s the venation beneath; inflorescence copiously branched at the a- pex of the stem; spikes greatly elongate, usually 15—l0 cm. long, densely many-flowere d, but apparently not sunpyiaed fruit; peduncles pking abbreviated, acutely tetragonal, margined; bractlets ovate, 2 mm. long, 6 -5—~-1 mm. wide at the base, ciliate-margined “8 the apex, puberulent ~ the back, acuminate, keeled; calyx slightly longer than the subtending bractlet, ght Z mm. long, : whitish-strigose; tempting sta equaling the calyx, s limb about mn, wide. The type of this remarkable plant was collected by Wilhelm Nikolaus Suksdorf — in whose honor it is named -- in a garden at scribed natural hybrids in this genus are also noteworthy for a paucity in fruiting, but none has the lack of fruit so complete in this one. The same hybrid has apparently been produced artific- dally by Dermen and is referred to by him as V. officinalis x prostrata. The source of Suksdorf's plant is not known, and neith er parent is known from the wild in Washington. I+ may be dist guished from V. officinalis in having its spikes much elongate, 6--16 inches long and d densely flowered, the flowers subimbricate, varieties it may be distinguished Ba the fact that the spikes are a. strigillose or puberulent and not set fruit well, if at all, 3 herbarium specimens, including type material of most be the names involved, a 2 mounted photographs have been examin ined ae tations: CULTIVATED: Washington: Suksdorf s.n. [Sept. 25, 190h] (P1—138360), s.n. [Nov. 21, ik (N—-isotype, N--photo of type, Pl—138361--type, Z—photo of type) VERBENA SULPHUREA D. Don in Sweet, Brit. Flow. Gard., ser. 2, 3? pl. 221. 183). Synonymy: Shuttleworthia diceras Meisn., Pl. Vasc. Gene 2: 198. 1965 Moldenke, Monograph of Verbena 243 1839. Shuttleworthia sulpimrea Meisn., Pl. Vasc. Gen. 2: 198. 839. Uwarowia chrysanthemifolia Bunge, Bull. Sci. Acad. Pétersb. 7: 278. 1840. Shuttelworthia diceras Meisn. ex Steud., Nom. Bot., Bert. ex Steud., Nom. Bot., ed. 2, 2: 750, in syn. 181. Schuttleworthia dissecta Walp. apud Schau. in A. aa Prodr, ll: 551, in syn. [as "Schuttlew."]. 187. Uwarowia sulphurea Bunge apud Schau, in A. DC., Prodr. 11: 551, in syn. 1547. Verbena dissecta Poepp. ex Schau. in A. DC., Prodr. 11: 551, in syn. 18h7 {not V. dissecta Morong, 1904, nor Schau., 1959, nor Willd., 1825]. Verbena ena sulfurea Sweet apud Schau. in A. DC., Prodr. li: 551. 1847. Verbena sulphurea ane apud C. Gay, Hist. Fis. Chile Bot. 5: 8. 189. Schuttleworthia sulfurea Meisn. apud C. Gay, Hist. Fis. Chile Bot. 5: 8, in = 1849. Uwarowia suypinres Schau. a~ a Jacks, in Hook. f. & Jac acks., Ind. Kew. 2: 1161, Verbena sulfurea a normalis Kuntze, Rev. Gen. Pl. 3 (2): 258. 1898. Verbena sulfurea D. Don apud Solered., Bull. Herb. Boiss., sér. 1, 6: 627. 1898. Shuttleworthia dissecta Walp. apud Reiche, Fl. Chile 5: 289, in syn. 1910. Shuttleworthia sulfurea Meisn. — Reiche, Fl. Chile 5: 289, in syn. 1910. Glandularia sulph- urea (D. Don) Schnack & elem Darwiniana 6: 475. 19uh. Glandu- sulfurea Schnack & Covas, Darwiniana 7: 72. 19h5. Verbena sulfurea Sw. ex Moldenke, nhowns 376, in syn. 1959. Ve erbena dissecta Walp. ex Moldenke, Résumé Suppl. 3: 37, in syn. 1962. Verbena lobata Meisn. ex Voldenke, Résumé Suppl. 6: 11, in syn. 1963 [nct V. lobata Arrab., 181, nor Vell., ay pabltography Sweet, Brit. Flow, Gard. 6 [ser 3): pl. 221. 1834; Lindl. Edwards, Bot. Reg. 21: pl. 1748 & 466. 13353 Meisn,, Pl, rag Gen. 2: 198. 1839; G. Don in Loud., Hort. Brit. ° e Ped. ores 2 (bot. ae 7881; F. Phil., Cat. Pl. Vasc. “chil. 2a, 1881, Briq. (» ete & Prantl, Nat. Pflansenfam. - (3a): 147 - Herb. e in W. B. Scott, Rep. Princeton Univ. . Patag. 8 (2): 688. 19055 Briq., Ann. Soabeev. & Jard. Bot. Gen . 10: 10h. 19073 ahh PHYTOLOGIA Vol. 11, no. Reiche, Fl. Chile 5: 273 & She ito 1910; Se aes k, Anal. Mus. Argent. Hist. Nat. Buenos Aires 24: {1h. 1 anzin, Anal. Soc. Cientif. Argent. Buenos Aires 88: 97, 98, th 121, 2 128, & 3): . ight; Moldenke, Known Geogr. Distrib. svcabaiine at fed ? 3 3 Mol graph. Distrib. Verbenac., [ed. 2], 102, 16h, & 199. 19493 C. Skottsberg, Medd. G&t. Bot. Tradg. 18: 151. 1950; Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 2209 & 2212. 1951; A- o de Vargas, Bol. Mus. Nac. Hist. Nat. Chile 25: ° 1951; Moldenke, Inform. Mold. Set 6 Spec. . 19513; Moldenke, logia 3: 68. aaah E. J. Peace td Ind. Kew. Suppl. 11: 101. 1953; Moldenke, Phytologia 5: 195h3 Moldenke, Inform. Mold. Set £8 spec. [h] (195h). lg see 3 (195k), and 52 spec. bh 19565 Mufioz Pizarro, Sin. Fl. Chil. 199, pl. 96 c--e. 1959; denk sumé Ee 2:10 & (1960 , 35, & 37. 1962; Mol- denke, Phytologia 8: 192 & 20h 2362), 3: "461 (1969), and 2} 67, 70, 72, 117, 130, & 192. 1963; Moldenke, Résumé S (1963) and 7: 8 & 10. 1963; iouiente Phytologia 9: er (96h), 10: 299 (196), and 11: 21 & 62. 1964. Illustrations: Sweet, “rg caer Gard. 6 [ser. 2, 3]: pl 221 [in color]. 1834; Lindl. in Edwards, Bot. Reg. a4: pl. 1718 {sn pac 1835; Fisch. & Mey., M Acad. St. P&tersb., ser. 6, {in color]. 180; Bocq., Rév. Verbénac. pl. ll. 62; ipl Engl. & Prantl, Nat. Pf am. (3a): 147, fig. STE. 1894; Sanzin, Anal. Soc. Cientif ~e Buenos > 1919; Mufioz Pizarro, Sin. Fl. Chil. pl. 96 c—e. 1959. Robust hairy perennial herb or Ser eratiaces, green or cinereous, 2060 on cM. pee, the ‘ieee often villous or hispid and ashy~- the lower ones remote, the segments all narrow or linear, rather uniform, simple and entire or trifid, obtuse at ne apex, wide- ding, stbeerslite or revolute along the margins; inflorescence capitate-globose, usually not elongating after anthesis, peduncul® te; heads many-flowered; peduncles short or elongated, leafless; 1965 Moldenke, Monograph of Verbena 245 bractlets subulate or lanceolate, 3--6.5 mm. long, acuminate * the apex, varying fron one-third as long to almost equaling th calyx, canescent—hirsute; calyx clavate-tubular or narrow o See Gric, 6--8 (—12) mm. long, plicately 5-angled, canescent-hirsut- ow and awl-shaped retrorse more dense hairs toward the apex, the limb 5- ~parted, about 8 mm. wide, barbate in the throat and at the mouth, the lobes cuneate, 5 mm. long, 2.5-—3.2 mm. wide, deeply retuse at the apex; anther-appendages varying from violet ion ticulate-rugose on the back, about half as long as the fraiti ng- calyx; chromosome number: 2n = 10. This + apparently a very variable species or else several taxa are herein included. It has been collected in sandy soil on ‘ 0 a Sandy soil just back from the ocean". A sulphur verbena". The plant has been in Gultt vation ayhe 1832 the coast and in the mountains, and that it also occurs in Peru. Schauer (1847) sotoniie cites a Meyen s.n. from "in Peruvia ad Chuquito", while Raimondi (1943) cites his no. 932) fran Junin, Peru, and no. 10562 from Hudnuco, Peru, but “of the latter he says "Flores de color morado claro". ‘Wiest Se, also cites ae I Hones that the Peruvian records may be based on misidentifica~ : I. M. Johnston (1929) asserts that the "Verbena sulphurea" and 'V. erinoides" of Philippi, Fl. Atac. 0 and Viaje Des. Atac. 2, sea (1860) are actually V. atacamensis Reiche. Skottsberg ~ Skottsberg 843 was first identified as V. porrigens R. A. Phil., then as V. berterii Schau. and so cited by me in Lilloa 10: 3k5 (194k), but™seems to be V. sulphurea eiendat _Smitioworthia diceras was based on nos. 1392 and 1809, collec- 1819. vio Giuseppe Bertero at Val in October, deposited paraiso, Chile, lobata Meisn. is based on Miers 335, also "from Valparaiso, ed in the herbarium of the British Museum in London. The 2h6 PHYTOLOGIA Vol. 11, no. Verbena dissecta of Morong is actually V. tenuisecta Briq., that of Schauer is V. berterii (Meisn,) Schau., and that of Willdenow is a valid species. The | Piste Lorentz & Niederlein reference given in the bibliography is sametimes erroneously dated "1889". The Lou- don reference (1839) is sometimes accredited to W. Baxter, appar= ently in error Material of ‘ve sulphurea has been misidentified and sistehiee in herbaria under the names V. berterii Schau., V. dissecta Spreng., V. erinoides Hook., V. erinoides Lam., V. atiiida Rutz & Pave, Ve re porrigens | R.A. Phil., and Shuttleworthia dissecta Walp. On the other hand, the Skottsberg & & Skottsberg 1395 distributed as V. sulphurea, is the type collection of f. alba Moldenke, and 885 Sy the type | type collection of var. fuscorubra skottsberg. Claude- oseph 3547 is a mixture with var. taltalensis Moldenke, and e, and Miers s.n. {Concan] is a mixture with V. berterii. ~~ Verbena sulphurea apparently is very variable, is a mixture of several tuck ty ate taxa, or else the published * descriptions are ed on misidentifications. Skottsberg (1950) sums up como é ly glabrous, both within and without'; de Candolle (Prodr. I Pp» 551) dice bees glabra', Gay (V p. 9) ‘tubo glabro en el ester- ior", Reiche (V 290) tubo corolar pelado'. La figura 3 de ut bubs con dientes largame visiones mayores del lim- bo corolar x 3 y los apéndices de los estambres ores 1.5 mm. (Bot. Register XM tab. 1145) las ti res son algo mayores que iedad fuscoree oe indi describes the "stamens" as b bly is ef the anther-appendages only. pe pt: er (1847) pa "Variat indumento magis minusve hirsu dove, canesc in his var. canescens, which see, Reiche speaks of the inflores~ cences becoming —— in age; possibly he refers here to what is now know as var, taltalensis Moldenke, Schauer (187) cites Bartero 1392 & 1809, Poeppig 157, Cuming 519, Gaudich s.n., Gay s.n., and Bridges s.n, from "In n Chili are~ nosis sterilibus n maritimis, ad Quintero et Valparaso, Concon, son as well as the Meyen san. from Peru. mentioned above, all deposi 1965 Moldenke, Monograph of Verbena 247 in the Berlin and De Candolle herbaria, the former now destroyed. Hauman-Merck (1913) cites a Garbers aun. preicmmnser n 348) from the plateau near Choele-Choel and Hier-Berg 12) from near de nin ag and San Blas, Rfo Negro, Argentina. , 96 herbarium specimens, including type material of some ee the names involved, and 3 mounted photographs have been examined me, Citations: CHILE: Aconcagua: s Valenzuela 803) (N); Glaude~Joseph 2516 orbs Eo E. he Kausel 2590 (N); Looser 5499 (N), 5503 (N); Rose & Rose 19395 (N, W——-761925). Atacama: Worong 16 (Cc), 1126 ©), Bar (Cc). Coquimbo: Barros Valenzuela 8037 (N)j We He Ja n, (Coquimbo, July—August 1856] (5); Looser 217 (N), OTe “ittehers & Skottsberg 789 (N), 843 (Go, N, S); Sparre 2680 (S). Malleco: Kunkel 166 (Cb). Santiago: BUcher, Hjerting, & Rahn 602 (S); Claude-Joseph 1725, in part (W- 1121280), Valparafso: K. Behn s.n. [29-VIII-1937] (N)5 Bertero 1392 (Du—166497, M, N--photo, S, Si—photo, Z—photo), 1809 (M), 3.n. [Valparaiso] (Br); Bridges s.n. [Valparaiso] (Bm, Br, Br); Buchtien s.n, [Vifla, 21/9/95] (S, #1177976); Clande-Joserh 3547, in part (W—12615)0), 3770 (W—12816)7), 4991 (W—-1422114); Col Gol- iector undesignated 275 [314] (ih); Garaventa 2438 (Herb. Looser 202k) (i); Goodspeed 4586 (Ca-—561316); Looser 2084 (N, N), 3997 (N,N), hO2, (N)5 Miers 72x (W—1343927), 335 (Bm), 8.m- [Concan] ip Moldenke & Moldenke 19762 age Lg, Mg, N, Ot, bei 19770 (B, Es, F, Fy, , Mr, N, No, Ok, Rs, “S, Sa Sm, Ss A a 2}; grea ieee aan J. hee Morrison 5 2 168s (ca-~629959, 8); R. A. Philippi 1299 (W—1323131)3 Rose & Rose 191k6 (N, W— 761710); Wall & Sparre 5 (Ew); Werdermann 17 (ca—238179, Ge— 34515, S). Province undetermined: Bridges 5.n- eg Claude-Joseph 2177 [Costa] apap: Cuming 450 (Br), SMe (Chile, 1835] (Br); Herb. Sullivant s.n. [Chili] (Pa); Miers 511 [south of Chile] (by, Siz Tenth of Chile] (Bm); Petré s.n. (S). CULTIVATED: Belgium: M. Martens s.n. [h. b. Brux. 1848] (Br). Italy: Herb. Harvey s.n. [hort. Corea: 1343] too-366196).. Switzerland: Herb. Hort. Basil. s.n. [Aug. 1841] (uM). Bibliography: Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 4,102 & 199. 1995 Moldenke, Alph. List Cit. 3: 940. 1949; C. — gee - G8. Bot. Tridg. 18: ag 1950; Acevedo de ca sts, Bo ac. Hist. Nat. Chile 25: 66. 1951; Moldenke, R 122° & “kth. 1959. form differs from the typical form of the species in hav- ing its corollas white or greenish-white. It is described as 4 2148 PHYTOLOGIA Vol. 11, no. perennial herb, 10—15 cm. tall, with leaves pubescent, bractlets L-—5 mm. long, ’ flowers very fragrant, calyx 8--8.5 mm. long, cor- olla-tube 10-11 mn. long, the lobes of its limb mm. long and 3 mm, wide, anther-appendages black, about 2 mm. long, and style 10 m . long. The type of this form was collected by Carl Johan Freder. Skottsberg and Inga Skottsberg (no. 739) at Estancia Frai pa at an altitude of 215 meters, Coquimbo, Chile, on August 13 or 1h, 1917, and is deposited in the herbarium of the Botaniska Trudgard- en at GUteborg. Worth & Morrison describe the "stamens" appendages only. The form has been found groring on —— page rocky oes at altitudes of 215 to 1800 meters, flowering in ust and November, Material has been distributed in Naehehe Bea the detention "Verbena sulphurea Sweet". In all, 4 herbarium 5; specimens, Yo aang the type, and 2 moun- ted photographs have been examined ions: CHILE: jae rie Shot isber & Skottsberg 739 (Go— type, N—isotype, N—photo , S--isotype, Z--photo of type)s Worth & Morrison’ 16690 radia Rs fond VERBENA SULPHUREA var. CANESCENS R. A. Phil., Anal. Univ. Chil. 35: "190, in nota: 1870. Bibliography: R. A. Phil. eet Univ. Chil. 35: 190. 18703 groeeeet Résumé mete Leet i to me about toi variety except that Philippi (1870) says "Se recojié una variedad 'canescens' de esta especie (Verbena sulphurea Sweet], que es una de las pocas que habitan en ambos lados de la cordillera." I assume that the name applies to the ashy-gray form of V. sulphurea ip to by me (above). More study is needed to determine if the gray and cane forms of the species are worthy of ane kim: separation SULPHUREA var. FUSCORUBRA Skottsberg ex Moldenke, Alph.- eres ah 3: 940, hyponym. 1949; Medd. G&t. Bot. Trudg. 18: Biblicgrepty’ Moldenke, Alph. List Cit. 3: 90. 1s 10. 19495 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 102 & 199+ Sept. 28. 1949; C. Skottsberg, Medd. Gtt. Bot. Tritdg. ery 151. 1950; Moldenke, Phytologia 3: 290. 1950; Acevedo de Vargas, | we Mus. Nac. Hist. - Chile 25: 66. 1951; Moldenke, Résumé 12 This variety differs from the typical form of the species having dark-red corollas. Skottsberg states that it also cers 2-—2.5 mm. long style 0 ae The type of tai vineey was senrestea a by Carl Johan Frederik 1965 Moldenke, Monograph of Verbena 2h9 Skottsberg and Inga Skottsberg (no. 885) at Loma de Frai Jorge Coquimbo, C , on August 19 ["August 19" acco label, but September 19, according to the original description], 1917 and is deposited in arium of the "Verbena sulphurea Sweet", It has been found in anthesis from August to October. In all, 3 herbarium specimens, including the type, and 2 moun- ted photographs have been exam me ons: CHILE: Coquimbo: Skottsberg & Skottsberg 585 885 (Go-- type, Bets of type, S--isotype, Z--photo of type). Ovalles Barros Valenzuela 8031 (N). VERBENA S ULPHUREA var INTERMEDIA Kuntze, Rev. Gen. Pl. 3 (2): 258 [as Hsulfurea®}. 1898. Synonymy: Verbena sulfurea f. intermedia Kuntze ex Moldenke, Suppl. List Invalid Names 10, in syn. 191. Verbena sulphurea f. intermedia Kuntze ex Moldenke, Suppl. List Invalid Names 10. 191. Verbena sulphurea intermedia Kuntze ex Moldenke, Résumé 376, in syn. 1959, Bibliography: Kuntze, Rev. Gen. Pl. 3 (2): 258. org WE Maclos~ kie in W. B. Scott, Rep. Princeton Univ. Exped. (2): 689. 1905; Moldenke, Suppl. List desl Naaes 10. ioidy j dendenke, Known Geogr. Distrib. Verbenac., [ed. 1925 Moldenke, Alph. List cit. Ir 93 "a9i6), 3: cat (ious), fie a "18 1949; Moldenke Known Geogr. Di ns rbenac., ite 8 a)..20 107, & 199. 19h9; Gonbaas de Var wea 1. Mus. Nac - iN Chile 25: ae 1951; Moldenke, résumé 122, 128, 316, & a hth. 1959; Moldenke, Résumé Suppl. 7: 10. 1963; Moldenke, Phytologia 9: 397 (196) a +} 52. 3 19 8 variety differs from the typical form of the species in ae the i ta to 15 mm. long, twice the length of the eas type of the variety “ collected by Francisco Josué Pas- casio Moreno and Tonini (no. 330) in Patagonia and is deposited in the Britton Herbarium at - cx New York Botanical Garden. The P. been collected along roadsides and in quebradas, at al~ ‘The p Sea te daacttvea bb ak hers Wits welts oes [Cuezzo & Barkley 20Mz538; Rufz Leal 1284] or white, purple int throat [Cuezzo esto & Barkley 20M 20Mz 378]. Kuntse says "Tubus calyce duplo longior — , 172 om longus.....Bei@ normalis ist der Cor ollen~tubus om. lang und therragt der Kelch nur wenig." In all, 16 pn Sher specimens, including the types of all the names involved, have been e me. Citations: CHILE: Coquimbo: Herb, Mus, Nac. Santiago 12 (N)- 250 PHYTOLOGIA Vol. 11, no. ARGENTINA: Mendoza: Carette 339 (N); Cuezzo & Barkley 20Mz378 (N), 20Mz538 (N); Ruiz Leal eal 126 (N) (N), 3627 (N), (N), 5130 (N), - (N), 7200 (RL), 10697 ( (Z); Semper s.n. [Herb. Rufz Leal Leal 319] (N ), Sn. n. (Herb. Ruiz Leal 4258] ( (N), sen. (Herb, Rufz Leal 10563] (R1), son, {Herb. Rufz Leal 13078] 8) (R1). San Juan: Castellanos 15923 (1 (W—— n, (Herb. Rufz Leal 13966] (Rl). Province undetermined: Moreno & Tonini 330 [Patagonia] (N—type). VERBENA S - LONOTTUBA Kuntze, Rev. Gen. Pl. 3 (2): 258 itt. wrtped od Logtasipe Runtae, Rev. Gen. Pl. 3 (2): 258. 18983 gone sioner seedy Dis ab Verbenac., [ed. 1], Pri = aes heart Castanea 13: - 19448; Moldenke , Alph. 628 (1948) and 3: “ere. 1949; Moldenke, Known i iegrionn “Distrib. Ver benac., [ed. 2], 102 & 199. 19l9; pote de Vargas, Bol. Mus. Nac. Hist, Nat. tite git 70. 1951; er ager Résumé 122 & 7h. 1959; Moldenke, Résumé Suppl. 7: 5 & 1963. This variety differs rae “the ‘gncek form of the species in its elongate corollas, which are to 20 mm. long, almost 3 times the length of t . The type of this variety was collected by Carl Ernst Otto Kuntze at Paso Cruz, at an altitude of 21,00 meters, O'Higgins, Chile, and is deposited in the wha ga eg Herbarium at the New York Botanical Garden. The s also been found at 1000 meters = pan flowering in are nr The Stockholm se cited be- curious pubescence on the calyx. Material has been identified and distributed in herbaria as V. sulphurea D. Don. In all, r ——— specimens, including the type, have been da iitetdons, "CHILE: Aconcagua: Herb, Mus. Bot. Stockholm s.n, (S). Coquimbo: Looser 216 (N). O'Higgins: Kuntze s.n. (Paso Cruz, 2400 m.] (N—type). SULPHUREA var. PEDUNCUIATA C. Gay, Hist. Fis. Chile Bot. be 8. 18h9. phy: C. Gay, Hist. Fis. Chile Bot. 5: 5. 18h93; Reich, 290. pet tegeroae Known Geogr. Distrib. Verbense 19 variety differs from the typical form of the having its leaves larger, with more remote segments, and the pedune~ les rae almost leafless, and : nothe ° was designated for this variet by Gay wees ing about iF except tat to sald by hin in hs original Tat eer’ nosis. His Spanish description is apparently meant to ale bo the variety and the typical form of the species, of which he says "Planta pte ae comun en las provincias centrales y del norte Valpa™~ 1965 Moldenke, Monograph of Verbena 251 ENA SULPHUREA var cevedo de Vargas, Bol. Mus. Nac. ne Nat. Chile "253 i odie ta 1951. b : Acevedo de Vargas, Bol. Mus. Nac. —_ Nat. Chile 253 67—~68. 1951; Biol. Abstr. 28: 90h. 195h3; Moldenke, Phytologia 5: 96. 195k; Team Résumé 122 & th 19595 Wolden- ke, Résumé Suppl. 7: 5. 4. This variety is said Py differ from the typical form of the species in being scabrous and more robust, with short rigid hairs, the The original descripti r as follows: fert scabre robustioreque » pilis brevibus, regidisque, denstoribus, longis, retrorsisque et nervio medio foliorum crassioribus simplisibusque, inferio t ores uate own at Huasco, Atacama, Chile, in September 1885, and is sheet no aa in the herbarium of the Museo Nacional de Historia Natural t Santiago, Chile. The plant is known to me only from the orig- inal description, in ae var. TALTALENSIS Moldenke, Phytologia 3: 281. bliography: Moldenke, Phytologia 3: 281 & 290. 19505 Molden- ke, “tad tee & h7h. 1959; Moldenke, Récumé’Suppl. 6: 6. 1963. This variety differs from the typical form of the spisiee in etary its pet Soreness elongated and spicate, .5--10 cm. long, even during an The type of this variety was collected by Ernesto Barros Valen- zuela (no. 8032) at Taltal, Antofazasta, Chile, on September Be 190, and is de deposited in the Britton Herbarium at the New York Botanical Garden, His no. 8033 was apparently collected at the Same locality on the same me day. Claude-Joseph 3547 seems to be @ mixture of this variety and the typical form of the species. In rye 3 mares specimens, including the type, have been ed by m Citations: | CHILE: Antofagasta: Barros Valenmela 8032 (N-- ae — (N). Valparafso: Claude-Joseph 3547, in part (WH VERBENA SUPINA L., Sp. Pl., ed. 1, 1: 21. 1753 [not V. supina 1 ° Syn nyay : Peristereon hyppios Diosc., Med. Mat. S: 496. 1529. Yerbenaca supina Diosc., Med. Mat. 5: 496. 1529. Peristereona — Diose., ware Mat. 5: ciate 1529. Verbenaca supina Clus., spina Gesn., 1683, nor Ruel, 1861). Sacra verhena hispanica — — or L'0bel, Icon. Stirp. 535. 1591. “Verbena supina clusit Tobe, Pg - Stirp. index n.p. 1591. Verbena s Clus., Rar. eaaat. - Stirp. : xlv & xlvi. 1601. Verbena tenuifolia C. 252 PHYTOLOGIA Vol. 11, no. & Bauhin, Pinax Theatr. Bot., ed. 1, 269. 1623. Verbenaca foemina Caesalp, ex C. Bauhin, Pinax Theatr. Bot., ed. 1, 269, in syn. 1623. Verbenaca supina (& foemina) Fuchs ex C. Bauhin, Theatr. Bot., ed. 1, 269, in syn. n syn. 1623. Verbenaca supina Fuchs apud Matth., Opera Omnis Comment. 71. 162. Verbena foemina Caesalp, apud Matth., Opera Omnia Comment. 71. 162),. Verbenaca supina J. Bauhin, Hist. Plant. Univ. 3: kl. 1651. Verbenaca supina Matth., Apolog. 17h. 1674. Verbenaca, supina, sive sacra herba Matth. apud J. Bauhin, Hist. Pl. Eur. 2: Ugh. 1689. Verbena temuifolia supina flo. coerul, R. Morison, Pl. Hist. Univ. Oxon. 3: "08" [=)18] & h19, sect. sect. ll, pl. 25, fig. 7. 1699. Supina sive foemina Park. secigg R. Morison, Pl. Hist. Univ. Oxon. 3: . 3: 9, in syn. 1699. Verbena; temifolia C. Bauhin apud Boerh., Ind. Alt. Plant. Hort. Acad acadi Lugd.-Bat., ed. 2, 1: 187. 1720. Verbena foliis decomposito-pinnatifidis, caule procumbente Toex Hort. cliff. 11. 1737. Verbena supina J. B J. Bauhin apud L., Hort. Cliff. 11, in syn. 1737. "Verbena foliis decomposito-pinnatifidis, caule diffuso L. apud A. van van Royen, m, Fl. Leyd. Prod. 327. 1740. Verbenaca supina | Cord. apud A. von Haller, Enum. Meth. Stirp. Helvet. Lo gee 661. 1742. Verbenaca supina s. femina Fuchs apud A. von Haller, Enum. Meth. Stirp. Helvet. 1: Sle 1742. Verbena foliis de composito-pinnatifidis caule diffuso L. apud Sauvages, ces, Meth. Fol. Pl. Fl. Monspel. 28.1751. Verbena procumbens Forsk., Fl. Aegypt+ arab. 10. 1775 [not V. procumbens Hayek, 1908]. Verbena radicans Moench, Meth. 369. 179 [not V. radicans Gill. & Hook., 1829]. Verbena tetrandra; spicis filiformibus, Ss foliis bipinnatifidis L. apud Desf., Fl. Atlant. 1: 17, in hs ‘syn. 1800. Verbenaca supina sive foemina Tabern. ath Desf., Fl. Atlant. 1: 17, in syn, 1800. Verbena sp spicis filiformibus, heared foliis bipinnatifidis; caulibus ri ramosissimis, decumbentibus Lam. 4 apod Poir. in Lam., Encycl. meyel. Méth. Bot. 8: Che, in syn. 1808. Verbena tetrandra, spicis filiformibus, solitariis; foliis pipinnatifidis L. apad Poir. in Lam., Encyel. Méth. Bot. 8: 545, in syn. 1600. Sacra verbena, hispanica, minor L'Obel apud d Poir. in Lam., Encycl- uéth. Bot. 8: 545, in syn. Tes. Verbena supina, sive = Tabern. apud Poir, in Lam., Eacycl. Méth. Bot. 6: 545, in syn. 1808. Verbena tetrandra; spicis filifomibus soitteriie, foliis foliis ee Kniph, apud Bt de Lapeyr. Me Hist. Abrég. 5 Fa Pyrén. 13, in syn, 1813. Verbena supina f hirsuta Ehrenb. ex Sweet, Hort. Brit., ed. 2, 418. 1830. Verbera esis; spicis filir. sotiteriis, fol. bipinnatifidis L. apud H. B. Richter, er, code Bot. L: Linn. %. 3. Bibliography: Diosec., Med. Mat. 5: 96. 1529; eco! Herb. Viv. Icon. 1: 120, 15323 Ruel, Pal Dioscorid. Anazarb., eds 2, 1965 Moldenke, Monograph of Verbena 253 1, 75. 1710; Boerh., Ind. Alt. Plant. Hort. Acad. Lugd., ed. 2, i: an (ar) and ed. 2 1: 187. met: L., Hort. tad ie 17315 A. » Fl. Leyd. Prod. 327. 1740; A. von H Meth. stir. “Helvet. 1: 551, 63h, & ser 1725 | sauages, ¥ Meth. Fol. Pl. Fl. - 248. 1751; L., Sp. Pl., ed. 1, 1: 21. 17535 L., Syst Kate, Aes 10, 852. 17593 maiphat, Orig. ae 12: no. : 3 Ver ena no. 3. 17685 J. A. Murray in » Syst. Veg-, ede 13, 62. att Forays Fl. Aegypt.-arab. 10. 1745; Sabbat in Ma- telli, : ll. 17 7753 C hristm. & pre totes flanz nteneyet Ray Fi 5: 130. 1779; RES 508 Stirp » 4t 226 2 . 1811; Picot de Lapeyr., Hist. Abrég. Pl. Pyrén. 12-13. 1813; J» Sm. in Rees, Cycl, 36: no. 2 1817; Steud., Nom. Bot., ed. 1, 87k. 1821; Spreng. in Le, Syst. Vege, ed. 16, 2: 750. 18255 Sib- horp & Sm ’ ci. Nat. Catania, ser. 2, 16: 125. 1860; Neilr., Aufz. 170. 1866, Ascherson in G. Schweinf., Beitr. Fl. Aethiop. i: 120. 75 Benth. in Benth. & Hook. f., Gen. Pl. 2 (3): 1146, 1876; cb Pe “apraarggey # Fl. Nontpel.. 53 533. 1876; J. Ball, Journ. Linn. 2229, 1 1879, Tristram, Surv. West. te Palest, ed. 1, 378 (1884) and ed. 2, 378. 1888; Batt. & Trab., Fl. Alg. 1: 717. 1888; Briq. in 25h PHYTOLOGIA Vol. 11, now & Engl. & Prantl, Nat. Pflanzenfam. (3a): 148. 1894; Jacks. in Hook, f. & Jac ks. » Ind. Kew. 2: 1179. 1895; Baker & Stapf in Thiselt.-Dyer, Fl. Trop, Afr. 5: 286. 1900; Fiori & Paoletti, Icon. tral. 127. 1909; Hayek, Pflanzend. Oesterr.-Ung. 1: 520. 1916; C. L. Pollard in Webster, New Internat. Dict. Eng. Lang. 2279 « ae A. Chev., Explor. Bot. Afr. Occid. Frang. 1: 50h. 1920; Vv Cun Arss oss r 7 1921; J. M. Black South, Austr. 3: 478. gt Hegi, I,lustr. Fl. Mittel-mur. 5° G): 2239. ape Stapf, Ind. Lond. 6: 31. 1931; Big in Repert, Beiheft 63: 153. 1931; Eig, Zohary, & Fein- ke, Prelim. Alph. List Invalid Names a — Worsdell, . Lond, Suppl, 2: 86. 191; Moldenke, Known Geogr. Distrib. Verben- ace, fed. 1], 5, Lh, 5, Ben & 102. "19h; eee Alph. List valid Names 51. 1912; Rozeira, Fl. Prov. - 164. 19h; Mol- denke, Phytologia 2: 339, “32, & 348. 19h7;3 "enaeaxe, Alph. List Invalid Names Suppl. 1: 27. 1347) Moldenke, Castanea "13: 12l. 19h8; H. Ne & A. Le Moldenke, Pl. Life 2: 19 & 3. 1948; Moldenke, Alph, List cit. 2: 356, 357, bik, 43, 482, 513, 535s 580, 5 s 1167, 1173, 1179, 1180, 1201, 1208, 1213, 1218, 1222, 1233, 12h1, & 1291. 19h9; Moldenke, Phytolo gia! a 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2 I, ‘1, 107110, 12h, 125, 165, & 199. 1949; Moldenke, Phytologia 3: 291. 1950; Moldenke in Chit~ den, Roy. Hort. Soc. Dict. Gard. 6: 2209 & 2212. 1951; Monod, Bull. Inst. Frang. Afr. Noir 1: 433 (1952) and 16: 28 & li. 195k; » rnat. Bot. 28. 195h; Sistine tie Nat. 59: 3h] & 342. 1958; Moldenke ls, 12 as 158, "i, 22h, 372, 378, 126, 427, & 47h. 1959; Moldenke, sien suné 8 & 11 (1959) and 2 13. oldenke, Phytologia 8: 20, U6. 1961; Moldenke, Suppl. 3: 27 (1962), kb: 10, 12, 15, & 19—21 (1962), and 5: 8. 19625 Cuf., . Bot. Bruxe 32: Suppl. 788, 19625 Z + Life Pa alest 16h. 1962; Molden- ke, Résumé Suppl. 7: 6 & 8. 1063) , Phytologia 8: 72 (1963), 9: 38, 198, 285, 286, 28 : cat (1963), 10: 210, 212, 217, & 282 (1964), and il: Si. Illustrations: Clus. » Rar. Rn a Hisp. pgs pot 2: 373- 15763 Dodon., stirp. Hist. Pemptad. 150. 1583; , Ico md Plant. 132. 1590; L'Obel, Icon. stom — 15915 1y Gerardo, B 580, fig. 2. 1597; Ger , Hist. , figs aot Clus., Rar. Plant. Hist, Stirp. ): pr nits 1601; Je Theat. Bot. 675. 1640; J, Bauhin, Hist. Plant. Univ., ed. 1 33 - 16 F) Je st. Pl. Bur, 2: Loh. 1689; Re Morison, Fi; Wake Wart. Ghent ta- debee ll, pl. 25, fig. 7. 1699; Kniphof, 1965 Moldenke, Monograph of Verbena 255 Orig. Cent. 12: no. 99 fin color}. ao Kniphof, ar i Viv. 2: pl. ier {in eames 176; Sibth. & Sm., Fl. Grae pl. 55h {in color]. 1827; ais Iconogr, 2: pl. 137 rr a i 1856; Reichenb., Pee Fl. Gem. 18: pl. 1292 [in color]. 1857; Fi $ Black, 1: 520. 1916; Murdeck, Acta Univ. Lund. [Lunds Univ. Arsskr. 2], new ser., 17 (1): 19. 1921; Javorka & Gsapody, Ic. 418. 1932; Monod, Phanerogames in Contrib. Etude Sahara rae 2: pl. 22. 1939. subterete or less pubescent or Prevetovsctd rtp branches mostly procumbent, stri- gose-hispid and more or less canescent, the tips often ascending from hypocraterifo rom blue or bluis purplish-blue or lilae, its ks “Sk mm. long, u s11y twice as long as the toad —-2 mm, wide; stamens lh; schizocarp the. original specimen of Linnaeus, preserved as sheet “0.83, 8 S. se, in the Linnean Herbarium at London, was collected iy bia George Clifford. garden in England an s and — $ not agree well with his arene ti of setae “1737 or 1753. In the Stockholm herbarium, however, there is a specimen of Loefling 16 which, in my opinion, should be regarded as the type- own ting there is written in pencil "Linné herb" and in ink " randra, spicis filiformibus solitariis, mp agree Linn. plant ing ve 1h in in Hispania 16 Loefl." The specimen is speuseithy of the supine form and thus agrees with the description of Linnaeus in 1753: b » VERBENA tetrandra, spicis filiformibus solitariis, foliis ipinnatifidie. Verbena tenuifolia. Bauh. pin, 269. Verbena supi- na. Clus. hist. 2. p. 6. Habitat in Hispania." | le (1962) 256 PHYTOLOGIA Vol. ll, no. cites the type of the species as "ex Hispania". Walpers (1845) classifies V. supina in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Sec~ ondary Subgroup Sc! Sehizophyllae, with 9 other species. The specif- ic name is sometimes uppercased, for no valid reason. Verbena supina has been collected in dry as well as mee places S lowlands and arid deserts, parklands and plains seaport ballast, at altitudes from sealevel to 90 meters, See ing in March and from om May to September, fruiting in te August, and September. It is native to the Mediterranean n and the Near East, introduced elsewhere. Baker & Stapf “1908 oe of it "spread through the Mediterranean rectonty Bentham (1876) Bere; “orbi veteri ph regionis Mediterraneae ab,ins. Canariensibus usque ad Orientem incola." Eig, riage apecginy (1931) re- port that in a it inhabits moist Bt part of the Crypsid min dis ecologi ation in that country. Mohr says that in 1893 it a "common st weed" at Mobile, Alabama, while Black (1926) states that it is common on parklands lai ar Adelaide, Sout ia. Douglas pa H. Kent, in a letter to me dated November 28, 1949, reports that it is adventive in England, and this statenent 4 ie borne out by the reference in the bibliography given above specimens cited below. It has apparently existed in mast Neg s in England sinc 799. It should be noted here that the Verbena supina of Tragus is actually Bidens tripartita L. in the Carduaceae, the V. femina of Brunfels is is Erysimum officinale L. in the Brassicaceae, the the V. procumbens of Hayek is V, hayekii Moldenke, and the Verbenaca su a pina of Ruel and of Gesner is a mint. C. Bauhin (1623) places Verbenaca foemina Caesalp. and Verbenaca supina (& foemina) Fuc in the sy synonymy of his Verbena communis caeruleo » flore (which is what we now call Verbena officinalis), , but I place @ them in the Be i! of Verbena supina. should perhaps also | be mentioned here that the L., Sp. Pl- (1755) ratehense is often erroneously cited to page "29"; the Clusius [=L'Ecluse] reference (1601) is Sen given as "2: 6" oF "2: 45-6, ig. et and the J. Bauhin (1651) reference in the bibliography ab e is sometimes misdated "1650" or "1750". The publication by Fein Ball (1878) is often cited as "Ball, Spic." Material of this species has been misidentified and saa in herbaria under the names V. canescens H.B.K., V- forskaelei Vahl, V. forskalei Vahl, v. a Desf., V. spuria L-, and Pri- va dentata Pers, On the other the Field & Lazar 190 (in part), Murbeck s.n. [Dar Farik, wT /E/B96T. and Ruhmer Sone [Ben- 1965 Moldenke, Monograph of Verbena 257 at igen a distributed as V. supina, seem to be f. erec- ta enke; C. B. Clarke 2018 is V. , officinalis L.j Pappi pi 2981 is v. sbrissistees var. prostrata Gren. & Godr.; Eyles 4349 is Vv. wrightdi A. Gray; G. Jermy s.n. [Ttara, 1853] is Heliotropium procumbens Mill. in the Heliotropiaceae; and Pitard 68 (at least insofar as the University of California at ra —— es specimen is concerned) is Teucrium campanulatum L. in the Lamiaceae. The Kulikowski 858, cited below, may TT aa f, erecta; Kneucker 266 4: is extremely white-hairy and may be taken as repre- senting Ehrenberg's variety hirsuta. The Lund specimen of Sin- tenis 1045 approaches f,. en cta, but the branches are too diffuse and the stems too hairy. Grossheim sen. [26,IX.25] is a mixture with Phyla nodiflora (L.) G ieee Schauer (1847) cites Tenors 2 s.n. from Italy, Duby s.n. from Sicily, Sibthorpe s.n. and Fra IS S.n. from Greece, Fenzl s.n. from along the river Danube in ——- Steven sen. from the Cau- casus, Kotschy 17 from Mesopotamia, motech 38 from Nubia, Acerbi S.n., Coquebert de Montbret s.n., Aucher-Eloy s.n., Mar- tins s S.n-, and Wiest 90 from Egypt, a, Setmes SoNe “fram Alge- via, Boissier sen. from Spain, and Broussonet & Chr. hr, Smith SNe fron the Canary Is. Islands, all deposited in the Berlin and D Decan- dolle herbari Irvine (18 59) records the species from Wandsworth, Surrey, England. Ascherson (1867) cites Cienkowsky sn. (seru, Fesoghlu, ? Marz 1848] and Ehrenberg s.n. [Chandek & Dar-Mahas, Nubia]. Baker & Stapf (1900) cite Ehrenberg s.n. [Nubia], Schweinfurth 578, Pfund 180, Fazokl s.n., Cienkowsky s.n., and Kotec hy 265 from the Nile aren; Chevalier (1920) records the species from Mauritania, citing Lt. G. Schmitt 28548. C. 1. Sandwith (1933) “cited vacant numbered @ollection Fre Baptist Mills, eon, England; Rozeira (194k) cites Ferreira s.n. [Braganga] and s.n. (Murga], Sampaio SeRe stem Sen. [Mirando-do-Douro], s.n. heehee and 8 -ne [Foz~Tua}, M. Lopes s. 8.n., Maria s.n., and Rozeira s.n. fro tugal; and Monod (1952, 195 5) sive his nos. 200 (227), 901h, “nd 908516 from Mauritania, Boucher cites ForsKk&l 2475. mY all, 14) herbarium eee, luding the type, and 7 mounted photographs have been examined by me. ey: Citations: ALABAMA: Mobile Co.: C. T. Mohr s.n. [Hun Sept. 5, 1892] (W--771870, W--771872), a: Tistiie, July 1893] (C). GREAT BRITAIN: England: Lousley s.n. [Sharpness Docks, 1 Juli 1957] (z). PORTUGAL: Ferreira s.n. [Julho 1880] (S); Mendes Pinheiro 1493a (Br); Rainha 219 (W--2109948). SPAIN: Alstroemer Sn. (S); Bourgeau 2163 (Du, N); Collector undestgnated Ste s.n, [in Hispania] (S); Herb. Mus. Bot. Stockholm sen. (S)j J z kenge ote Sots )3 (17 Jul. 1851-52] (5), sens (July 1052] (Du); Loefling 16 [ 258 PHYTOLOGIA Vol. 11, now h Linnaeus G.35, $.16] (E--photo of type, F--photo of type, Ls-- isotype, N--photo of isotype, N--photo of type, S--type, Sg-—-photo of type, esahete of type), sen. (S); E. Reverchon 81 (Pa); Sen- nen & Mauricio 7657 (Ba); Vahl s.n. (S). CZECHOSLOVAKIA: Leijeme sen. (Br). HUNGARY: Borb4s is 93 (B, (B, B, Go, S, W--811335); miler 1502 (B); Boros 5 (S), Sen. (30 Jul. 1933] (Ew, ag = 367621); Collector undesignated s.n. (Sg—-16111); Herb. Bormmtfller s.n. (B); eave Dorfler s.n. [1 stead 188] (S); Janka 1853 (Ca bee 91015, Du, Du), s.n. [26 Septembr. 1860] (Gg—- 2)3 Ge. G. Jermy Ss. ne ad: W--315852); Korenl s.n. [26 Julio 1877] (Go); Kovées 1,60, in part (Ba, Br, Gg—163265, Go, S, Ur, W-—13h6993)5 Lengyel s.n. [23.VIII.1914] (B); Ré&cz Son. (7.vrII -1911] (Go, S, Um—61); Tauscher s.n. [2 Julio 1878] (Go (Go). GREECE: Heldreich 99 (S), Sn. [31 Jul. 1852] (S); Marie & Petitmengin 375 (La)- ITALY: Herb. Martius s.n. [Palermo] (Br); Heuffel s.n. 1. (Br). SIC- ILY: Todaro 396 (Du, (Du, La, S), sen. [1850] (S), s-n- (S)- GRETE: Duffour 1507 (B, Vi). JUGOSLAVIA: Serbia: Sintenis 1669 (Iu). UNION OF SOCIALIST SOVIET REPUBLICS: Azerbaijan: Grossheim S.De (26.IX.25} (B). Donetz: Gorschkowa 18) (S). Stalingrad: Becker s.n. [Sarepta] (Du). Ukrainia: Kulikowski 858 (Go, S). MOROCCO: Herb. sen. [Tétuan, 1845] (Du—166)68); Jahandiez 240 (cp), 976 (Herb. Lacaitu 27915) (B, Bm); ie y, Rungs, & Sau- vage 2503 (S); Sennen & Mauricio 8866 (Um—l). ALGERIA: Bové s.n. [Septembre 1839] (Du--166467); L L. eee 81 (Du); Cosson Sen. [29 Juill, 1854] (S), sen. [20 Mai 1056] (Go); O- 0. Debeaux gon. (Pran, 28 avril 1683] (Br, Du), s-n. [1883] (Br); Jamin 3503 (Um—~63); Le Cesve s.n. [7/6/1934] (Ge—223437)5 funby $-Be [eirea lacum Halsula] la] (W—147591); Reboud 2209 (Du). TUNISIA: Doumet-Adanson & Bonnet s.n. [29 a vril 1884] (Br); Kralik 3a (Du); Pitard-Briau 468 (Go, Vt), sen. [Gabés] (Vt). ~CYRENAICA: Ruhmer 262 (Br). EGYPT: Aucher-Eloy 2700 (Du—166)50); Boucher 363 (Du—166500); C. G. Ehrenberg s.n. [IV. Cahiro] (S); A+ “he Grey 8.n. [Egypt & Nubia, 186 9} Ore, F. E. ~E. Hubbard s.n. [Feb. Bh; 1868) (W—71956); Kneucker 266 (B); Kotschy s.n. [Blue Nile] (T)5 13479 (Vi—126), sen. (22/3/47) (Bm), son. (20 fév. 18h8) (S), s.n. [Matarich, Cairo, Mart.] (W--1)7590); Letourneux $-Ne [april 1876) (1); Samaritani s.n. (S); $ Schweinfurth sn. [ Sone , 1880] (W—806208), s.n. [10 Juni 1887] (Br); Sieber 2 (Iu), SMe [Beniseuf] (Bm, Vt); Teickholm s.n. [Luxor, 23/1/1927] (Ka)5 As Wiest 90 (Herb, Prager 18642] (Du, Gg—-31454). SUDAN: Nubia: Kotschy 323 (Du—166L51, Lu, S). ARABIA: Yemen: Herb. Mus. Bote Lund. s.n. [Arabia felix] (lu). TURKEY: Ascherson 10h5 (Br); 5i® tenis I0L5 (Iu, S). ISRAEL: Jouannet-Marie 671 (Du). LEBANON: : Gombault L008 (s). JORDAN: Field & Lazar 190, in part (Du-—-2396%7)" 1965 Moldenke, Monograph of Verbena 259 805057). PAKISTAN: Sind: Kotschy $1 515 (S). MALAYA: Malacca: Col- lector undesignated sen. [circa Malaccam] (Lu). CULTIVATED: Bel- gium: 3 Lejeune s.n. (Br). England: Linnaeus s.n. [Hort. Cliff G. 83k, S.7; Bailey Hort. neg. 608] (N—-photo). France: Herb. De~ ee sen. (h. P. 1822] (Br). Germany: Herb. Calif. Acad. Sci. 3 (Gg); Herb. Prager 18637 (Gg—31)55). Italy: Herb. H san. [h. R. P P. 1819 ] (Du—16666). LOCALITY OF COLLECTION UNDE- TERMINED: Boisduval s.n. (W--71989); Collector undesignated 77 (Q); Hazslinszky sn. [Kisupszallas] (W—71958); Herb, Alm. 5.n. (S); Herb. Harvey s.n. (Du--166501); Herb. Mus. Bot. Stockholm s. n, [21/8] (S); Herb. Mus. Paris. s.n. (Ca—963026)} | Herb. Osbeck 2 (s); Mauricio s.n, [Melilla, VIIL.1939] (Vi); Osbeck $. sen, (S, 2 (5,8)- SUP: . ERECTA ease, Phytologia 2: 339, 30, & 32, nom. nud. (19h 7), £. nov Synonymy: Verbena supina var. major Foumerque ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 197. Verbena su- pina var. glabra Lojac. ex Moldenke, Résumé 376, in syn 1959. - Verbena supina var. erecta Bég. ex Moldenke, Résumé 37 376, in syn. 1959. Verbena supina f. erecta Sennen & Mauricio ex Moldenke, Ré- sumé Suppl. 1: 23, in syn. 1959. Verbena supina var. subglabrius- cula ee ex Moldenke, Résumé Suppl. 7: 10, in syn. 1963. Bibliography: Moldenke, cteie petec a 33% 340, & 342. 19475 Moldenke, Alph. List Invalid N :'27. 1947; Moldenke, abe’ List Cit. 2: 619 Gols), ae "sho, "33, "& 9h0 (19h9), and : » 1167, 1168, 1180, 122h, & 1919; Moldenke, Phyto- logia 3 Us & 138. 19 4 Moldenke, Known Geogr. Distrib. Verbenac,, [ed 12h, & 199. 1949; Moldenke, Résumé 130, Lite ial, 138, S06, 426, \2t, & 7h. 19595 Moldenke, Résumé Suppl. 1: 1, & 3 3 (1959), 2: & & 6 (1960), and 7: 10. 1963. Haec a forma typica caulis erectis recedit. 4 This a osen differs from the typical gsi of the species in having ts stems usually erect or suberect. The type of this form was collected by Elisée Reverchon fs 81) laces at Algeciras in southwestern Spain on June 2h hy SPT adopted, but I publication for any of them. It seems that the f. erecta of Sen- nen & Mauricio is based on their no. 8500 from Morocco, eco, while var. Subglabriuscula is based on H. Lindberg 89 from Spain. Material of this form has been collected in flower and fruit in June, Herbarium material has been widely distributed as V. supina L. More study is needed to determine the actual taxonomic status of this form, which seans at and intergrade with “the typical form 260 PHYTOLOGIA Vol. 11, no. or perhaps to hybridize with it. Material of Rainha 785, Sennen & Mauricio 7657, Kovécs 60, Balansa 193, Kralik 321, Letiurneux 8. n., n., Tackholm S.n., A. Wiest 90, Kotschy 323, Ascherson 1045, and Field & Lazar 1045, “cited by me under V. supina, may, upon re~- examinat tion, - both surfaces; spikes pedunculate, 2--10 an. long, capitate young, fascicle-like in anthesis, becoming cylindrie-elonget® in ves and often interrupted below in fruit; bractlets subulate or linear-subulate, strigillose, ciliate, abo ut half as long 45 the calyx; calyx 12--15 mm. long, strigillose, the longer teeth setaceous-subulate, 2=-l, mm, long; corolla hypocraterifom, vary~ ing from blue to purple or Fose-purple, its tube ee 1.5 cme js long surpass he calyx 2-- mm., scent on the oute surface and in the throat, the limb 10—12 m. ithe: the lobes b texas emarginate at the apex; anthers without a gland or ge very one on the connective of each of the upper pails fruiting-calyx 12--13 m. long, strigillose, the lobes 1965 Moldenke, Monograph of Verbena 263 very slender, unequal, subulate; mature schizocarp slightly con- stricted along the lines of cleavage; cocci cylindric or subcylin- at » about 7 mm. long, with a broadened base, furrowed or stri- ate toward the base and areolate or reticulate-scrobicul ate to- ward the seiag or coarsely eben ie the middle to the apex, the commissural ae devs minutely scabrou type of this species was SelLeseed by George Valentine Nash (no. 270) just back of the beach line at Ballast Point, near ences, The entire absence of the gland on the anther connective, a character of the section in which V. Aubletia is placed, at ased. De Ss swamp", Nebb comments “anther tips small or wanting" Curtiss 1963 is a mixture -- the part [19634] pial "sandy Sitges bordering th the ocean, eastern pest is V. V. maritima Small; the part [1963#*] from "roadsides near Charleston" is V. bonariensis L.3 and the | — from "low open woods, Indian River, June" is V. tampensis Nas Material = ‘Ve tampensis has been misidentified and distribu- ted in herbaria under the names V. aubletia L., Ve ¢ canadensis (L-) Britton, V. lambertii Sims, V. maritima Small, and Glandularia maritima (Small) Small. On the other hand, the Curtiss 5706, A- A. Eaton 5i6, W. H. Rhoades 6, and E. West s.n. [Chattahooc hee, 8 Sept. 19 1931], distributed as V. tampensis, are all V. maritima - The Collector undesignated s.n. specimen in n the University of Vermont herbarium is a mixture with Stylodon carneus (Medic .) Moldenke, a oe as yet se bes Curtiss 1963, in part (low open atid "Setar Bivée pa (E, G, W); Fredholm Im 580), (G). Br Co.: J. P. Young 737 (w) Dade Co.: Edw. Palme 6433 (E). pay ee G. V. Nash 2470 (G--isotype). Lee Co.: W. Kellogg s.n. [ ak 1907] (G); P. Ce . Standley Sen. [thicket along river, cad 8 March 1927] (W)- Manatee Coe: Rugel 305 (E, F); S. M. Tracy ‘6650 (E, G). Volusia Co.: Mell s.n. (Daytona, 2) Jan. 1907] (E). The Nash 2470 which she cites from "Ny" is actually in the Columbia University herbarium; the "Mattern" 3.n. which she cites is doubtless the Mattern & Mat- t is —<™ s.n. cited by me below; the "Small 8469" which she cites a 26h, PHYTOLOGIA Vol. 11, no. § typographic error for Small 8619; and the "Standley s.n." which she cites may be the P. C. Standley 52577 cited by me. She says "Verbena tampensis, apparently endemic in peninsular Florida, is very ery similar in habit to V. canadensis. It differs yas havi more shallowly incised or dentate leaves, a strigillose calyx, slightly longer nutlets, and essentially glandless anthers. The nutlets are enlarged at the base somewhat more than those of closely related species." In 7 herbarium specimens, ig the types of all the names inv olved have been exam xamined by m Citations: FLORIDA: Brevard Co.: Brows & & West sen, [Rockledge, 6-28] (F1—20981); W. M. Canby s.n. [Febr. 1889] (Pa); Curtiss 1963, in part [low open woods, Indian River, June] (Bc, C, Cm, 2 Os Ts Pa, Pa, Vt), 5706 (Ka); Mattern & Mattern s.n. [Titu srilte, March 3. 191h] (W-~910330); W. H. Rhoades s.n. [Cocoa, Sept. 7, 1936] ese 12861, H-~51418), sen. (Indianola, Feb. 3, 1937] (F1—25896)5 Je K. Small 8732 (N), s.n. [Cocoa, May 9, 1918] (Hi--7209, Up)- Dade Co.: Edw. Palmer 396, in part (Bc). Hillsborough Co. F. S- Blanton 6936 (M1, N), sen. [Dec. 1h, 1930] (I); A. P. Garber $e (Tampa, May May 1876] (C, W--265076); G. V. Nash 2470 (G=-type, | ype, isotype); C. L. Pollard s.n. [Tampa, March 7, 1698] (W--328227) - Lake Coe: J. K. Small 8649 (N). Lee Co.: Buswell s.n. [June iy (Bu); | Cy C. Deam 58711 (Dm, N), 58795 (Da, 5 (Dm, N), 60759 (N)5 P. C. Standley 52577 (W--1308777). Manatee Co.: C. H. M. Barrett 13 (W--893560); Cuthbert s.n. [Palma Sola, March 26, 1926) (F1l—- 20983) P. Dowell 7101 (N, Um--l7); Nebb s.n. (March 1887] (Pa); Rugel 305 (Bn, N, W—512258); S. M. Tracy 6650 (N, N, W--383903), 7808 (Cm); Weber & Kelbert s.n. . [Palmetto, 3-15-28] (F1--20980) - Orange Co.: He Ne , Moldenke 212 (E, N, Up, Ur). Pinellas COes Beckwith 772 (W—1200200); R. F. Thorne 10281 (Ca--965339). Polk Co.: McFarlin 4477 (Gg—237856). Saint Lucie Co.: Small & Matt haus 9635 (H—-l3153, S, W-—-1738h78). Sarasota Co.: Fattig 5.Me Noe 2-28] (Sa); H. N. Moldenke 1039 (B, E, G0, H—5080, ™ » Ur, W—-1567275); He B. Rusby s.n. (Sarasota, Febr. 12, 1335)" (x); B. H. Smith s.n. (Osprey, March 17, 190k] (Cm, ae 23135). Volusia Co.: F. Beckwith 773 (W—1200201); S. M. Dea 1831 (Du—8027)); Eastwood s.n. (Holly Hill, May '90] Gi ot278); Friesner 2037 (Bt-—5987, Bt--599h, Bt-—-6023); B. H. Patterson S-Be [Daytona, Feb. 23, 1918] (Cm), s.n. [Daytona, Feb. 1925] (Cm); de K. Small 8686 (F1—27705, N, N, S, W); Small & DeWinkeler B50 sy (S). Merritt's Island: A. a Baldwin 70 (N). County undetermino®® H. C. Beardslee s.n. [Cheney Highway, March 15, 1933] (Ob—97290) 5 Collector undesignated sen. (Vt, Vt). XVERBENA TEASII Moldenke, Phytologia 2: 28—29. 19. 1965 Moldenke, Monograph of Verbena 265 Synonymy: Verbena erinoides x hybrida Dermen, Cytologia 7: 163. 1936. Verbena hybrida x erinoides Dermen, Cytologia 7: 163, 165, tenuisecta Briq. ex Moldenke, Alph. List Invalid Names 7, in syn. 1942. Verbena teash Moldenke, Am. Midl. Nat. 59: 362, sphalm. 1958. Verbena tenuisecta Briq. x V. hybrida Voss ex Moldenke, Résumé 376, in syn. 1959. ‘in Bibliography: Dermen, Cytologia 7: 163, 165, & 169, fig. 3-5, 3 id Names 47, 1942; Moldenke, Known Geogr. Distrib. Te beet ze 1949; Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: uppl. 11: 263. 1953; Moldenke, Journ. Calif. Hort. Soc. 15: 82 & 83. 195l4; Moldenke, + Midl. Nat. 59: 362. 1958; Moldenke, Résumé 22h, 36h, 366, 376, & L7k. 1959; Moldenke, Phytologia 8: 121. 1961; A. Putz, N. Y. Herald Trib., ny 4, p. 18, February . 1962; Mdidenke, Phytolo- 3 Illustrations: Dermen, Cytologia 7: 163, fig. 3-5, 165, fig. 23 & 2h, & 169, fig. 43. 1936; A. Putz, N. Y. Herald Trib., sec. ’ Bag, February [as "V. erinoides"]. 1962. e anthesis; peduncles (1.5—7 em. long) and rachis slender, acutely vetragonal, densely hirsutulous or spreading-pilose, not glandular; practlets relatively very short, lanceolate, about | mm. long, a~ Sut 1/3 as long as the calyx, attenuate at the apex, rather dense- ly white-strigillose with appressed hairs, densely white-ciliate 266 PHYTOLOGIA Vol. 11, no. 4 toward the base; calyx eres f rbirpaagady 8-13 mm. long, densely short—pubescent with spreading hairs or *white-strigose with close- ly appressed ments corolla hypocrateriform, showy, varying from blue or purple to pink or white, if tube 15-~20 mm. long, about 1 1/3 times as “a as the calyx, brous throughout or slightly puberulent at the apex on the outer probly the limb 5—9 mm. de. This hybrid appears to have been made first in the so i of Edward Teas at Houston, Texas, in 1926, and was called "V. hybrida x erinoides" by him. It is typified, however, by a collection made by Glendon A. Stevens (no. 4) at Harrisburg, Dauphin County, vania, on June 20, 1933, and deposited in the herbarium of the Bailey Hortorium at Cornell University, Ithaca, New York. In 1936 it was produced artificially by Dermen in Massachusetts. Under the collective name, xV. teasii, I include the garden varieties called arpa lh Bellaire, Ceres Ceres, Madge Roberts, Rowena Ruth, and Teas Hybrid V or Tea s Hybrid Vervain, The corolla is white in Albion, apy in Rowena aa Ruth, purplish in Bellaire, and dark-red in Ceres. oe plant has been Collected in anthesis in April, June, July, and October, so apparently has a long bloam- period. Material has been distributed in herbaria under the names V. incisa Hook., "V. erinoides hybrid", and "Common verbena % ay wate anerte is said by Teas to be "probably the best of the rac Bailey says for his unnumbered spec ci- ted below, sh pane on July 22 and on October 3, 1927 ares seo . Teas, Houston, Tex., in spring of 1927"; his no. 7394 r sents Bellaire, 7395 is Ceres, and 7396 is Madge Robe oberts. A letter written | by G. A. Stevens ni abr Oe Horace McFarland Company, Harrisburg, Pennsylvania, ad d to Dr. Liberty Hyde Bailey and dated June 20, 1933, is of sie nepmett interest in the information it gives about the color forms of xV. teasii: "Dr. McFarland wrote you today é out the Texas verbenas. I am sending you by accompanying mail some samples which I hope will reach you in fair condition at least. "I am afraid that Dr. McFarland got the history of them a lit- tle mixed. Teas sent us originally four varieties. They were "1. A pu the botanical description of Verbena venosa. "2. A good light a variety, a little variable in a but is very similar to the purple sort, except that the flowe betwen are better ened and the plant seems to be a trifle nore goro "3. A A rather impure white variety which seems to have peewee able pink at times in the flowers. It differs from the former "h. A very ugly magenta-pink which was so bad that we threw it away at the end of the first year. I do not remember anything >” bout it beyond that it resembled the other three in a general Way 1965 Moldenke, Monograph of Verbena 267 "All four of these verbenas were reported to be hardy. They f cold frame without trouble. But six miles north of Harrisburg on the southern slope of the first mountain a friend of mine has a rock garden, to which I contributed some of each of the three varieties which we kept. They flourish exceedingly well and under a stone and take root there. You will note from the speci- mens that they are very flat trailing plants and root as they run. "It was in that garden that the red variety originated, evi- dently as a sport from either the white or pink variety. You will shorter growth. It is an exceedingly neat edging or bedding Plant "I am almost positive that no verbenas of the usual garden type were growing in the garden at the mountain last year, and I inoides", but certainly does not represent that species; it seems clear to me that the plant depicted is xV, teasii. In all, 21 herbarium specimens, including type material of all the names involved, and 2 mounted photographs have been examined me, Citations: CULTIVATED: California: Bailey & Bailey 9830 (Ba)- Haiti: Ekman H.9357 (S). New York: L. H. Bailey s.n. [July 27, 1927; Bellaire] (Ba), s.n. (July 27, 1927; ceres] (Ba), sen. [July a 1927; Rowena] (Ba), s.n. [July 27, 1927; Ruth] (Ba), S-De Scan Rag 5 2 (Ba), 3 (Ba), (Ba—type, N--photo of type, Z—-photo of type). t L. H. Bailey 739 (Ba), 7395 (Ba), 7396 (Ba); Teas 5.Ms (June 2,°1926; Albion] (Ba), s.n. [June 1926; Albion] (Ba), 5.R+ (June 1926; Bellaire] (Ba), sen. (June 1926; hybrids] (Ba), 5M: (June 1926; Madge Roberts] (Ba), sn. [June 192%; Rowena] (Ba), S:n- (June 1926; Ruth] (Ba). Baca VERBENA TENERA . in L., Syst. Veg., ed. 16, 2: 750. 1825. Synonymy : Starks tense x re Walp. Repert - ks 13. 1845. Schuttleworthia tenera Walp. apud Morren, Hort. Belg. 20: ’ Syn. 1870. Verbena geraniifolia Hort. ex Rttmpler in Vilm., Ill- 268 PHYTOLOGIA Vol. 11, no. ustr. Blumeng., ed. 1, 1261. 1873. Verbena geraniaefolia Hort. apud Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 1178, in syn. 1895. Verbena tenera Griseb. ex Alvarez, Fl. Sant. del Estero 106. 1919. Verbena apulchellas Sw. ex Augusto, Fl. Rio Grande do Sul 233, sphalm. 1946. Glandularia Lenora 3 ccarener metering Man. Fl. Al- reded. Buenos Aires 398. 1953. ta f, arenaria Herter ex Moldenke, Résumé 376, s Soaks we Verbena montana Lillo ex Moldenke, Résumé Suppl. 6: 11, in syn. ee “1963 {not V. mon- tana Hort., 1895]. Verbena tenella Michx. ex Yoldenke, Résumé _ Suppl. 6: 11, in syn. 1. 1963. Verbena tenera Auct. ex Moldenke, Résumé Suppl. 8: 6, in syn, 1964. Verbena tenera Briq. ex Mol- Bibliography: Spreng. in L., Syst. Veg: -, ed. 16, 2: 750. 18%; Reider, Ann. Blumisterei 8: [pl. 12]. pe Steud., Nom. Bot., d ed. 2, 2: 750. 1841; D. Dietr., Syn. Pl. 3: 60h. 183; a Repert. h: 13 & 3h. 18155 Schau. in fe R. Moldenke 458 (Fg). Cumberland Co.: Ahles & cs 104103). Duplin Co.: Ahles & Ranseur 21,010 + GTO) Hoke Co.: Ahles & Neuber 25065 (Hi—JOlh01). Robeson Co.: ables & Ramseur 23 23705 (it --T0Le32); Be Britt 2261 (Hi—180051). Scotland Co.: 294 PHYTOLOGIA Vol. 11, no. 5 Ahles & Hammond 21,790 (Hi—104400). SOUTH CAROLINA: Allendale Co.: C. R. Bell 261) (Hi-~101646); A. R. Moldenke hig (Fg). Bam berg Co.: Ahles & Haesloop 22108 (Hi--1h6192); A. R. Moldenke 26 (Fg). Barnwell Co.: A. R. Moldenke 423 (Fg). Berkeley Co.: Ahles & Haesloop 2237 (Hi—101645). Calhoun Co.: Ahles & Bell 1010 (Hi—92669). Charleston Co.: Vanderbilt s.n. [April 10, ISLi] (N). Chesterfield Co.: A. E, Radford 12h58 (Hi-~92871). Clarendon Co.: A. E. Radford 21219 (Hi—-1016h3). Darlington Co.: Be E. Smith 1459 (Hi—26h79), 1460 (Hi—-26478), 1h61 (Hi--26180) . Co.: Ahles & Ramseur 23307 (Hi—101629); Go SC 49027 (N). Dorchester Co.: Ahles & Haesloop 21836 (Hi—1)6193). Edge- field Co.: A, E. Radford 20371 (Hi~-10164)). Florence Co.: Ce. Re Bell 7541 (Hi--101642); A. R. Moldenke 7 (Fg). Georgetown Co.? A. E. Radford 21587 (Hi--101641). Hampton Co.: Ahles & Bell 10592 (Hi—-92870). Horry Co.: C. Re Bell 6269 (Hi—1016,0). Jasper Co.: Ales & Bell 1043 (Hi--92826). Kershaw Co.: A. R. Moldenke Lb (Fg); A. E. Redford 2370h (Hi--101639). Laurens Co.: C, R. Bell 5561 (Hi—101636). Lee Co.: A. R. Moldenke 1437 (Fe); A- E E > ag 5 & E d & bes 8 $ Io ir 1010, (N); W. H. Rhoades s.n. (Fitegerald] (Hs). Bibb Coo: A» Re Moldenke 370 (Fg); We H. Rhoades s.n. [near Macon, July 19293 H- N- Moldenke 17085] (Bt, N, St)- Brooks Co.: D. S. Correll 5549 (Dm, Dp, H~4023k); A. Re Moldenke 309 (Fg, Fg)- Bleckley Co+: Ae R. Moldenke 377 (Fg). Bulloch Co.: A. Baker 20 (Hi—186571)5 H» Y 33 (Hi—~198332); C. Owens 26 (Hi--166302); Pyron & ve Vaugh 136 (Gu--106); S. Taylor 27 (Hi—198297). Burke Co.t A. Re Mod denke 407 (Fg). Clarke Co.: Eggleston 510) (N). Clinch Coes As Re Moldenke 330 (Fg) P. 0. Schallert san. (h/29/u1} (Du--369013)- iquitt Co.: Kirkpatrick s.n, [Doe Run, 6-1h-30) (Ga). Crisp C0! He Hume s.n, (Cordele, 13 May 1937] (Fl—26626). Decatur Co! A. Re Moldenke 296 (Fg). Dodge Co.: A» R. Moldenke 365 (Fe)- Doughert; BeOll): | ¥ Co.s Eggleston 5104 (Cm, Vt, W-586011); Costing 80 (BY~ IF a Fs F 1965 Moldenke, Monograph of Verbena 295 355281, H--1255, Up). Echols Co,: P. O. Schallert Seb sapped (Ca—882635, Gg——-310859). Effingham Co.: Cronquist 1289 (Gu— 28157, N). Emanuel Co.: Craig & Craig 3392 Recast: Evans Co.: Miller & Maguire 322 (Ua—16571). Grady Co.: A. R. Molden- ke 299 (Fg). Hancock Co.: A. Re Moldenke 392 (Fg). Irwin Co.: Pyron & McVaugh 2118 (Gu~16 16575). Jeff Davis Co.: A. Re Moldenke 353 ( (Fg). Jefferson Co.: A. R, Moldenke 00 (Fg). “Lanier Cos: De'l Ee Eyles 5702 (Sa); A. R. Moldenke denke 32h (F; (Fg). Laurens Co.: Gronguist L875 L875 (Ca—788971, Gu—2761h, N, Tl, W--1927215). Lowndes Coes A. R. Moldenke 315 (Fg). McIntosh Co.: Chrysler & Johnson 350 (Hp, Ru). Mitchell Co.: Bai ley & Bailey sen. (Camilla, Mar. 29, 1933] (Ba). Muscogee Co.: S. F, Evans E.4270 (Up); Je Me Reade sen. [April 3, 1930] (Gu--E.4270). Richmond Co.: Duke 601 (112977). Sereven Coss Ahles 54307 (Hi—-200786); ¥. H. Duncan 12h1 (Gu—18203); A. R. Moldenke enke L12 (F (Fg). Tatnall Co.: Cronquist 953 3 (Gu~30565, N, W--1928907). Taylor Co.: Dress, Moore, & Law rence 633 (Bm, Go). Telfair Co.: A. R. Moldenke 358 (Fg). Thomas Cou: We H H, Duncan 1313 (Gu—-18199); A. R. Moldenke 305 (Fg); Oos- ting 9h (H--L0h26). Tift co: Mcatee 3509 (Hi—-52015); W. Be Rhoades s.n, (near Tifton, September 1935; H. N. Moldenke 17087) (au—122769, Br » Bt, Dp, N, St); Van fefor sen, (Tifton, nepal 26] (Ga), sen. (Tifton, April 1926] (Ga). Toombs Co.t J. S. H per 470 (Ns Ae M. Huger s.n, (Vidalia, na 1914] (romteBy. Treutlen Co.: McKellar Ly (Gu--1551h). Turner Co.: W. H. Rhoades Sen. (Ashburn, July 1929; H. N. Moldenke 1708k) (Br, Bt, M1, N, Nd, + Twiggs Coes A. Re poidedis 2 (Fg). Ware Co.: A. R. Mo ai 335 (Fg); P. 0. Schallert 25 part (B, Go, Hi-5l952, Je— 976, We), s.n. [ T8A/L0} at e7p6h). we Washington Co.: A. R. Mol- Gerke 397 (Fg). Wayne Co.: P. 0. Schallert 251, in part (Hi— “i 9, 5). FLORIDA: Alaclua Coz: Janish & danish 26) eT Murrill 111 (W—-18)6590); Small, DeWinkeler, & Mosier 0%) 5 Watson & Murrill Sen. rmtgh Springs, 4/30/39) ae: Weber & West; 8.n, (Paradise, 6-8-27] (Fl—21105). Baker Co.: L. Ky Ba Bailey s.n. (Glen Saint Mary, July 30, 31, 1918] (Ba); H. H. Hume S.n. [Glen St. Mary, Aug. 13, 1928] (Ba). Bay Co.: Davis ¢ & Davis lils2 (Hi—-115553). Calhoun Co.: A. R. Moldenke 272 (Fg)- Dads Cow: S Small, Small, & DeWinkeler s.n. [Deering Hammock, Dec. = (NW, N). Escambia Cost Ahles 7120 (Ur); Brinker 313 (I). ee Son. Gdaden Co.: Buswell s.n. (May 1928] (Bu); Exploration Party 1937 Sen. [1h Mar, 1937] » e928) 4 Kinkaid & West s.n. (ut. Pl Pleas- ant, Pei oa (Fl—21107); Mcatee 3349 (W--1368568); A. R. Mol- 1 (Fg); O'Neill sah EEE (7%); B.D. wn. [River Junction, | Apr. “19, sas} as 595); Alexander s.n. [Chattahoochee, 296 PHYTOLOGIA Vol. 1, no. 5 1931] (N, N); Small, Mosier, & Matthaus 12778 (H--82867, We); E. West s.n. [Chattahoochee, 8 Sept. 1931] (F1l--20997). Gulf Co.: Small, DeWinkeler, & Mosier 11225 (N, W—1738770). Hamilton Co.: West & Arnold s.n. (Jasper, 11 Apr. 1940] (F1--32028). Holmes Coe: z. S. Ford d 3551 (W—2230900). Jackson Co.: H. J. Banker 3685, in part (N); A. D. Brubaker 22 (St); Exploration Party 1937 sen. (15 Mar. 1937) (Fl—2703h); Grossman 7 (Fl--21108); H. N. Moldenke 8354 (N). Jefferson Co.: Be Le B. L. Wagenknecht 3615 (LW, Iw). Lake | Lake Co.: | L. H. Bailey 6812 (Ba). Leon Co.: Je Pe ~Gilles- pie 365 (Hi—112936); R. K. Godfrey 53039 (N); Hume “& Wast sone Tallahassee, 29 March 1936] (Fl—-20999); C. Jackson 16h (Ld); Ae R. Moldenke 288 (Fg). Liberty Co.: A, R. Moldenke 277 (Fg)- war ion Co.: H. N. Moldenke 1083 (Go, H~-560, N, Ob--68710, S, Up, Ur), 108a (N, S, Up, Ur). Polk Co.: McFarlin 5775 (Mi). Wakul- la Co.: Trott 100 (Ca—95388). Walton Co.: E.G. G. Hume 3 32-11 (Fl— 2127). Washington Co.: Hodges Son. (Chipley, April T 23, 1937) (Fl- 28083); F. H. Sargent s.n, [Careyville, June 20, 1950] (No--23913, Ok, We). County undetermined: McAtee 3590 [State tins (Hi--52016). : Autauga Co.: R. M. Harper 25 (N, W--1365901). Butler Co.? W. He posses s.n, (Georgiana, July 1931] (Bt, Bt--29647, N)- Es- cambia Co.: F. S. Blanton s.n. (Canoe, Oct. 25, 1932] (I); Gaines & Gaines 268 (N). Etowah Co.: W. H. Rhoades s.n. [near Ataltala, July 1918] (ky) » 3en. [near Attala] (Hs). MISSISSIPPI: George Co-? Demaree 32831 (Le); Diener 835 (T1, Ur). Hancock Co.: Diener 865 (Ur). Harrison Co.: Demaree 2 28400 (N), 29006 (Au—122772, N » Ole St), 30362 ptgriree N); Goodspeed s.n, (Biloxi, Apr. 1925) (Io—-1171,98); R. J. Lemaire 802 (Tl); A. R- Moldenke 25 (Fe)+ Jones Co.: J. D. Ray 5017 (Hi—199516, "N). Lincoln Lincoln Cos! Webster & Wilbur 3282 (W--2068016). Pearl River Co.: Hummel gone (Pearl River, 17.1V.1958] (co) 3 A. R. Moldenke 250 (Fe); We H. Rhoades 8- n. (Poplarville, Sept. 190; H. Ne Moldenke 17086] (er, Bt, N, St), Son. [near Poplarville, Sept. 1940] (N). Scott Coe: We He Rhoades s.n. [near Morton, July 1927] (N). Wayne Co.: Cooley r& Pease jul (Hi--193305) . TENNESSEE SEE: Henderson Co.: Sharp, Fairchild, a, cisbecty & Clebsch 9298 (0). ARKANSAS: Calhoun Co.: Demaree 22509 (N)-+ LOUISIANA: Beauregard Par.: A, R. Moldenke 708 (S). Bienville Par.: Moldenke & Woods 542 (S). Bossier Par-: Moldenke & Woods 5u6 (S). Claiborne Par.: Moldenke & Woods Skil, (3). Evangeline Par.: Ewan 19365 (Tl). Jackson Par. Moldenke & Woods 531 (S)- n Par.: Moldenke & Woods 539 (S); B. Williams s.n- (Ruston, March 28, 1950] (Ca—86352, T1). Natchitoches Par.: L. 8. Fox 402 (Fx), 407 (Ne), hO8 (Fx); A. R. Moldenke 699 (S); Ware e son. [June 2h, 1956] (S). Orleans Par.: Aydell s.n. (Feb. 1932] (Ti); Humet son. [New Orleans, 17/4/1958] (S). Vernon Par.: A. Re Mo. enke Herbarium curators who have material of this species from additional counties are asked to send it to the author for verification and record, so that future editions of this map may be more complete Mapping by counties done by Andrew R. Moldenke Figure 18. Distribution of Verbena tenuisecta in the Jnited States. ee te tc 1 7 = 7 ae | 13 l 7 15 I : 16 if 17 I 18 19 20 21 I 2 23 I ry) 25 26 27 28 29 30 31 it 32 I 3 I u I 35 I 36 e H 1 A Stas — er now. | sornnea = POLETTE =| TOWNER | CAVALIER PEMGINMA gy TTSON OSL X rocsivar Willams } * <7 . OUNTRAN — rent an = ae &a % maser | WASH er. THe woos aon | omer o fo ge Bexsom — en Tea 'o at cone pore — . ‘a a ot 7 Siewtnaw) . MELSON | GRAND FORKS " ry ng coOK bd ao | JPE pa wlan a U R : N SHERIDAN was 0 * TTASCA E ‘ _* ® JH | D 5* o PaAIRiE 4 Baines et FOSTER youn aomom v ~ *, suneen | K0D0ER ee HUBBARD ‘S *» Nemes aoe = ty Ouver ‘STUTSMAN STEELE CLEAR . waTeR ums (e s: ga . s was | cass + é oy CusTEe ST weaur DEN _ onTOR a ATOM X\ ” *, WaLLeY wee . Oo s a g ORTOMAGON om od * ae Sk SLOPE MeTTINGER 2 oo CARLTON J tasiiaas eee ‘ © 7 cour BARAGA rf om Tawons | ocan Tia en Oren TAR DOUGLAS 7. coutsie an man croc bor 2 ome @ SOWA ‘ADaats ‘OUR M I N- Ss :. som — » 7 DELTA iy a wm 7000 Crow WING] nit *e cate os \p nes paaee Pe TwasnbuRe| SAWYER | ASHLAND a POWDER eiven a ARO Ld aati . — . b = 7 PERKS a Ss =: - = PRICE 7 nenet Cc wee x < pee - | ; BENTON o wen : WARINETTE | peesout st < i z . EARRS, iSanTl Fores! * —_, — Tare Po BARRON | RUSK ~ ar we ’ mepacn J cewey ae SHERBURNE i“ mean noe oy 4 mat se) laps t Ana oan - 4 2 — a bur per waa |8 beso we” 2% Come . wees e O FRE sco owe comes porier wctxER HY weicht z CHIPPEWA =) coll ® je 7 ROOK fost cox | ou WARATHON ct —_ t ad < MEADE ooh SHAMAN loconTO oe ine] Fees Es b ARMSTRONG ta I bona od c ° | ee aa am as Wo OoONSIN a el - pak! ae REMVILLE CARVER PIERCE “ aceon |S luo oN aaah Maoisom [Tron STANLEY cay CLAIRE WOOD PORTAGE — | WAUPACA - ° LAKE ma | Pree Ss A oe ae. orn x - as = on = L e z an : se BUFF 3 © ste aaron pout anwn i" a fi r ‘ er WESTON nnd WCOULET/ SUEUR | RICE = cake TOM, ss aid WOT spmncs ry 2 errert) oi aa ~ ene = PENNINGTON BEADLE wuMGSBURY = ee 2 220 ADAMS, P aed —_ BUFFALO senauo [sansonn Tuimcr | axe | MOODY ig PIPE-| MURRAY waste Dons watt = seal wcosta | saa veo Uincouy) SUbLETTe CUSTER JACKSON ‘STONE coriowmane | waTowman a canst ng — Suan | NEw e Cameo var z awa Taunowa ue EARTH STERLE J toaee- | gon DY UN is cant Ld SHOR | washaaaucy mie aan] Meno | MimnETA ROK TRORLES PACKSON [mann | FARGAULT |Frecponn | woweR | FRIMORE a] SUNEAL eee i eorcas a mae from ty Tere < wousTOn mote) mm) Fy 2 ar ‘s s . . FJ or) BENNETT ) ‘CARLES [DOUGLAS | HUTCHINSON] TURNER a Se sn Lemuel PossuTayemmrnsen| wowts | MITOMELL| HOWARD | (Soe ran oun wa A _ sat SAU Dan or) es | ‘5 . - i iowa ed ——— Shur Vemen | cur ALO ALT WANCOCK CCAS Wnts Seow | oe ee ae Bon [esacton wan om an Faverre 7 ‘ 2 toa aoe yor car Gan mi KEYA PAHA tone = GORDO a “anwonts] RACI r WoT TuouTNToneROREE] BUENA Jrocmonrasfmomeaco] WRIGHT | FRANKLIN] BUTLER ] BREMER “ cane | rm) BROW VISTA CLAYTON = rT) ocx wwox cepan 5 7 ry a “ s * = eee eae mt beni Me DAVIESS STEPHENSON Hy woven | ad oun BLACK aoinant ANTELOPE] PIERCE cmumor| HAW . = wa weoosuRY CALHOUN Ton Tan [oenton [Lim | JONES cannon. [osu Te [wae (COO 4 eal cant T nooner | tomas] pune "™ORSTO" (Cowon | crawrord [Carman [oreene [S0one | STORY SO was poral BERRIEN . be ume Loup | GARFIELD] weenien Watson |stanrom] Commins A oun T CEDAR WHITESIOE Sas 7 Tate | ache ORAM rasa vel seat or som) Lacan | custen HARRISON | SHELBY “cuiame TOALAS [Pox [uASPER | roweswex| 1OWA Po ee | wk sareanef ST anaes [oat am WALLEY | GREELEY Pure ous] cooce , Ty SME w B A S A Tar or aed oa 0 err Cee OR | N ae marr peal eset WS BGTTAWATTAMIE | CASS | ADAIR | WADISOM] WARREN | MARION BAHASA | REORUN | WASr 1SLAMO| RARAKLL = ox Ps « 7 SHERMAA | HOWARD Burien |SUMOERSN occas LOUISA D> warkceR ea 5 ad Wl oe Y EP anne ernicx // POU TiwsstOn | usr s nouns ws bon ay pL er Za Taaaus Tomson ccanad [LUCAS [monn T warenio | i wasnincTom J yom ur SCOT . CHAMPN . of iP a & om v mooamay | worth urna . rarewELL s ff a fe ‘HITCHCOCK mn poate GAN TOE wit oa to OS - cg ia aed | Fumas CLARK wason} E rn Tao ae . wiLow Hanae | Fan waco] Tran Pare _— SIR Thon Ton Fins pe Le i = LAWARE . wor (GRUNDY DAMS one) aw! * alone A= pe be s wensTER ‘ . soy F CREYENING RICHARDSON AROREW DAVIESS Hl -~s Eat een) 4 pecaTuR ronan rues [sw Tew | rePuauc TO MARSHALL | REMARA i aes mao seeusy [manos oo : - ‘ waa SANGAMON cas as serum [Thoms Senco CHARITON was scorr a} % H Gun [Roos | osaome Puro] Fea Taney ron |e cus 'e GREENE PUCOUPIMY | custian) SHELBY mars 0 I — oe PK —— el OTTAMA ‘oe FAYETTE. | cevimcnais] JASPER WALLACE | LoGaN BOONE UNcOUn CHEYENNE beaded TREGO — Gans SHAWHEE nano, Soar SRST ans on ‘ - GEARY cay | om" worson 2° cay iain i WABAURSEE r Tiswoan > manner |cuam cs TuNTON — } Gretvey | wicnira | scott K ax _ ae ae - cooere ~ 7 ae . A na a TEFrERSON i a ae = N RICE Ss A CHASE Ss S I wasHacTOM com ‘ ae woauon nator | Fewer ee | ner ae ae ai = ote _ = , 3 STAFFORD "HE RSON mw ANDOU! na — 4 RENO HARVEY wiLeR Gaay BUTLER GREENWOOD CAMDEN LPS Ne, — EDWARDS S WOOOSOM| ALLEN = aN stanton | cant Paani oo u cor ~ CMA Som | ME - iat UACLEDE TEMS s 4 KonTow ] stevens | sewano | MAPE — } cLaan rao SUMMER COWLEY a ey mee Lg cHyTaQua| «© CHEROKEE @ JASPER Tan So = = an sm BEAVER hei ChaisTon DOUGLAS —T Oil a Hy HARPER wooos ALFALFA | GRANT uy OSAGE wfmowaral CRAIG | OTTAWA a" J ~ My i ; Taney — [OZARK a i i — DALLA a . . ELLIS’ | WOODWARD TT mide Mona s soemuan Tuansrono Tocm reer Toes . CARFIELO WOBLE ROGERS ‘ = ce 5 hee = meh . NARDIN WPSCOMk wAJOR pel MAYES -d BENTON CARROLL BOONE banTe » ves rad qmos| SANTA FE SHARP vg wicuet we HARTUY woort Tau DEWEY _ ] CHINSOM BLAINE | KINGFISHER | LOGAN CREEK 1S C “2 $6365) roserTs | MeMPHLL TULSA] WAGNER i wASHINGTON — so ITS Sanam s Ges ‘Quay meas TINCOUN CHEROKEE \ AMO PENDENCE, Banvecvont a aege e308 pee rT 2 , —— PONSETT ; ie. "1717 | nae porrer | caRSOW oT on ogrh MUSKOGEE sane Wcrnarongy™ | scrrcon] port [van BUREN CLEBURNE an ON : ee A H A SEQUOYAN TERN coattt ARON CURA PE . ee canapunn — | onLanoMa Site —T tess oreo gaint : | 3 ' OEM SMITH Gray . Pome ST) a ail wos r wert Lae \ ‘anuSTaomG | bom FY Poumssacay cagoo WuHES : es fecooaurt [ST FRANCIS Coe _—_ hed E xX I re) 4 5 creer Chow ms PITTSBURG — UF FLORE YELL pane MANNER ve TON] [ACORN je PARMER : MeCLAIN ° scort Hea wera ess DE BAA Castano | smsmen T beiscoe Tua — COMANCHE PomToToc SALINE Pennie = fun ie. 2 a . TACKSON L Com — = CARLANO PULASK! mame = ue) r TL mS | conve f LONOKE OA | LapaverTe i. sosevect Dairy * MURRAY McCURTAIN = So JEFFERSON | ARKANSAS PonTor . — shoes FLOW money [comme | “AROEMAW * o — TOMAR] PIKE var som [MN = ChavES = % JEFFERSON JOHNSTON cue pu Om enema | won KEYED TO COLUMN A SEVIER Foard wonta °Poony sq yee omen ie CALA CeLEvELaed p UMCOLN ape of Cleartype Index Book HOCKL WHLBARGER rl a DESMA = By COCHRAN fy [tussock crosay Tomas Tune r= a ‘a >» Pa . . me = = St aaestt 7 aaa =z 7 MRCHER MONTAGUE [Sone GO | cxarson hy ® amas cs wevaon TouacniTa DREW ce - R ram RED maven PO al . — BRADLEY LEFLORE! a : ’ somes ® — YOuNUM | TERRY Ther s MOCTA ARIA KENT aK DELTA nonveet STONEWALL | HASKELL | s0cemoevosT YOUNG Wise veron eae byes a ny TMON Tooy x forms | 2 5 CASS iat woNstN = aes Gawson T Booen | scumsy “ 1% “g Fswer — | sones ~~ [SHAcntirono | stepwens [Pao pwr | PARNER | TaRRANT [DALLAS [oct TAKS iC r, 2 ) os WOOD: tglal 08 . wesnowe | _ ME 7 DEROALL 5 woo mewre [UU TRADE MARK REG. U.S. PAT. OFF. 8 Maan HOWARD ns MITCHELL CALLAHAN | EASTLAND ERATH ‘MOOD JOMRSOM aus ea SaiTH 3 . Ba f+ — = YAN ZANDT GREGG CADOO aT Ne "2 ty Kory NO Sa soe s ss ate 2 oy wu | Pama at ULBERSON ona ECTOR noun] KS o ) Heo ° " Loving WML ER Wiovano [casscocn [tema | cont eee COMANCHE a agg Romeo 3 10 O ite gg es — BOSQUE ‘i . | nerves ANDERSON walp Tanne FreesTome san ce UNI l ED S I ATES % says = Yon GatH MILLS HAMILTON ‘te Erman \LIMESTOME _ " T TRON CONCNO ear TPERRY | GREENE : PECOS CORYELL ‘WOUSTON e F E X | aioe i S\ Aco Scale of Miles ‘ ‘nensas FALLS . uel | ¢ CROCKETT si \ ‘ 100 SCHLEICHER WERARD aia noeearsoe Gane aariots : . at Si \ Cried ki coell " 0 EF cans oe BURNET — - 3 oe = a Tiana err) S\- i ALLEN rr) ) 7 | ig Game's 7 roi ron sT pawns 4 fee a | i Paes BREWSTER TERRELL SUTTON iia wrLLlaMsone st ae i rancid AMMAN 2) Tt pl oN MAP NO. 125 010 Noe *. aston \PAROA Lea . la ~ GHLESPIE > Tee \ BURLESON came Tirtt som ) ACADIA abe (oon VAL VEROE TOWARO: — tans, BERTY Lead SNA wtRVULEL crevon pant Orr! U . = WASHINGTON gen 2 WOLUSA 2 KENDALL , bey st t Ht i oe ot ) BR CS AMERICAN ZOMPANY, INC. %» ‘BANDERA om. s : cui | \ camel Sartre Bia \ eons} rat ORIGINATORS Yom RS) SOLE PUBLISHERS / : eins ite ss a Tinmy WADE roy rene Jeans: (oe i= > TERREBONNE * aga [lenses om. u BEND amet ae j *, PRI feu CLEARTYPE MAPS Sage) COLORPRINT MAPS wi sow S wAnoo , ar Tenet 5 eo i fe. WHARTON BRAZORIA Passe \ NEW YORK ¥ MAVERICK | ZAVALA FRIO JACKSON wonce— 4 J TCE—This ss a copyrighted map. The lew A ATASCOSA ‘eTacoRA TT a s20ROUm" Petia prohibas che reproductmn or copying of same, WARNES, 3 or any pornon thereof, by any proces for per i sonal ust or resale, without permiamon KAUAI / Doar TA SALE WeMULLN ok cacaad . : BB A ctor / H A W A I I . REFUGIO wut st wok ms Coan frog = “ =| re ; Sain / S DA Tu WEL ges norway wae ork / wr wutces RET ad “' — / omu yy / s \ft / f J oe ial = / 7 Pd \ mLEBERC ame ‘ 7 Pa 5 L. . warn Ym woes Torooas Tw] wwe a nen ae — oo 7 MOLOKAI ) ry > - 0 Scale of Miles 20 f > Z . x — | LANAI a ? Tae rT) ial 1 . nue 7 rail a ) ° _—— ~ : a” [Moh KAHOOLAWE \ * 3 ; i, ~ avy HONOLULU om eC sation Y PART J ae @ So LISIANSE!, >) ———) * ~~ as . UAYSAN” 1 ] e” Y * . \ 5 ~ 8 ALEUTIAN ISLANDS a S “ " THIRD JUDICIAL DIVISION ; r NECKER teed 8 4 ww Ps @ 2 s.. - 5 . ° ” Miles 4 s i i ' ~ % of , 00 \ i Sf Copyright, American Map Co., Inc., New York, No. 14324 . * Zz . * om . I 1 | | % | I : [ ‘ 5 $ es i0 it 2 13 I is 5 16 i7 is i T = it I a T : 21 i 2 I 2 I u I 3 I 26 I Fa I Fy T Fz T ) T mM I 2 if 3 T xu if % Tr ry \ 1965 Moldenke, Monograph of Verbena 297 70k (S$). Webster Par.: Moldenke & Woods 5h5 (S). TEXAS: Angel- Se amma pent ina Co.: De S. Correll 15325, in part (Rf); Lewis & Oliver 522 (Nb); Lundell & Lundell 11065 (N, Rf). Chambers Co.: Miss Math- Le H. Pammel s SNe ate K |) 3-16-29) (1 (Io--132999) 5 Traverse erse 981 (au--17901), 1383 (Au--197025). Jasper Co. Lundell & Lundell 10511 (Ca—-767266, Du--364576, La, W--1887687), 10550 (Ld, W— Fe Rose~Innes & Warnock 21818 (Au, hs _ sen. [h/ ] (au, §),. Jefferson Co.: C. Ce Alb (au, Au), Be 2 (8/29/3h} (an); Je Fe te 2 48068 an “Yorteser Cons te a 4 og (Au). pee Co.: Correll, iene, we Edwin Ficio Co.: Fe Be pri fom (Fj). Shelby bo tin York, & == 11239 ) (Au). ‘Trinity Co.: ¢ Cory 54830 (Rf). Tyler Cos: ae & Cory 2210 (Tr); Tharp, Turmer, & Johnston 54849 (Au— 122754, St). Upshur Co.: Shinners 1012 (Gg—392103). Victoria oe Bindwall 28 (au); Parks & Cory 11105 (Tr). Waller Co.? Gory, S253 (Wo—-14789, Rf); Trew 202 (AU—122763). ARIZONA: Cow: & Ehlers & Ehlers — 1 (Mi); Di Fe He Gols S23 =S1007, N). CALIFORNIA: Los Angeles Cost L. S$. Rose 46296 (Gg); ures Ve Me S 3.0. ri Angeles, 192] (Ca—279772)5 Ms Winter s.n. (Los Angeles] (V). Santa Barbara Co.: H, M. Pollard Son. [Dec 1, 1950] (ag—~385736), sen. [June 7, 1957] (Ge— )$ Ge Fe Smith 2403 (ag—-355016). Ventura Co: He M. Pol~ lard sen. [apr. 29, ; 15th] (Og—330308), san. [Oct. 1, 19bh] (Ge— 330972), sn. [apr.. 28 » 1945) (Bg—-330972) » MEXICO: Yucatdén: E. C. Stewart 683 ( (Mi). HISPANTOLA: Dominican Republic: Ekman E15 (N (N, S, W~17115k6); Faris 178 (W—11L5997). PUERTO RICO: Stevenson & & Johnston 1845 (W—1)75867). WINDWARD ISLANDS: Barba~ dos: Bailey & Bailey 257 (Ba). VENEZUELA: Angostura: Bailey & Sat. . (Ciudad Bolivar] (Ba). Federal District: Ll. ans 10138 Seago te, PERU: Cuzco: Herrera 3450 (N). BRAZIL: : Dusén 9341 (N, S, W—-1481763); Hatechbach 4218 (Ok), 7hbb (Ca); atecibech & & Pereira 7689 (Herb. Hatschbach 10306] (Lv); 298 PHYTOLOGIA Vol. 11, no. 5 Hooker s.n. [Mai 1858] ai Swallen 8872 (W-~19331)1). Rio Grande do Sul: Arc her 233 (N, N, Sp--36826, Sp, W—1705702)3 Henz E.9591 (N) sz hering 373 73 (N, Sp-—20071); Jurgens 47 (B)s Os0- Tio | 8.n. [Cerro Tobatinga, Feb. 2h, 1947] (Ug-—-14136, ~“Ug——1s151) 5 Rambo 3939 mak. Inst. Bot. S. Paulo 50988] (N), 43778 (Rb), 51652 (W--2102128), 51703 (S)s A. Re Schmitz 165 (N). State un- Cruz: Kuntze son. [00 m., Sta. Cruz, 5.1892] (N); Paredo s.n. [Bl Pari, a-1eT (ag—-353278, N). Sucre: M. Cardenas 628 (N). PARAGUAY: C. Alvarez s.n. [Asuncion, aa (Je—883h); Balansa 1025 (N--isotype); Hassler 2660 (N), 6144 [Macbride photos 17451) ‘(Ca--94h365, Kr-~photo, N, N, N--photo, S), 7607 Sey ee 9 12411 (Ca-~929695, N, Coenen 12518 (Ca—929705, N, W—- 1057h06); Herter 84886 (N); JUrgensen en 377 [Herb. i * 22243) (Du—-197838, N, S, Ug, We-1483011), 3777 (N); Lindman a.%h3 (WN, , W—-1417046); 0 W—227580); Ponder 8.n. [Asuncion] (Je--7615); Ponder & Alvarez 8on. [10.18.1952] (Je—7617); Reales 253 (N); Ae = Robert O8 8h9 849 (N)3 T. Rojas 25% (Herb. Osten 1986) (0 (Ue), Dy 2998 | up2k (kK). URUGUAY: C. Alvarez s.n. [Trinidad cm 0 ae ee @ (ve, Ug), 22 cap 23 (Ug), 3 "a (Ug), 30 (Ue)s H. H. Bartlett t 19862 (Ca~-772525, ;, W~1904710); Berro 133 (N), Sr (N), 6022 (N), 7225 (MN); Cabrera 32h) {Herb. Osten 23345] (N,N, S, Ug)s Cabrera & Corte 9590 (W-—-2197981)5 sien Se ne [Belle Unién, Jan. wey 1948; Herb. Inst. Miguel Lillo ray (, N); Chebdatarorr son, [San Javier, 1939] (Ug--762)5 Herb: Mus. Hist. Nat. Montev. s.n. [dept. Minas, Feb.] (Ug), S-R« {banks of Rio Negro near Mercedes] (Ug); Herter 181h [Herb. Her ter 85536} (N), 1814 [Herb. Herter 96600] (N), bill (N), 50887 (N), Son. [Nueva Palmira; Herb. Osten 19032] (Ug), sen. [Paysan~ df, Sept.-Oct. 29075 _— Osten 20415] (Ug); Schrider 8-n- [dept - Paysandé, 192k; Herb. Osten 17303} (Ug, U 6) 3 rets- seire s.n, ‘oubeadiey "1913) (Ue) 5 D.¥ ight s.n. [Cunapiru, 1928] (Bm). “ARGENTINA: Buenos Aires: Boffa (Hy Gabreve 1860 1860 (N), 5179 (¥), 5628 enn oA: comersin 8 s.n, [Buenos- Montevideo] (B); Ryerdam, Beetle, & Grondona zu. ()) Hast 56 (N), = (N); Kuntze Ang FR , Nov. 1892) (N); Lefebre S- n, (Buenos Aires, (Br); Parodi 8962 (N), 12233 or Leal (im, Bm); Wall & 1 & Sparre s.n, [Sierra Bachiche, 2/11/46] (&, Bx), s.n. (Sloe El Volean, SFiL/é) (Ew, Ew); F. A. Wetmore 830 (Fe 1965 Moldenke, Monograph of Verbena 299 1065981). Catamarca: Wall & Sparre s.n. [La Negrillo, 28/11/16] (Ew). Chaco: R. M. Aguilar 109 (N), 643 (S, S)s JUrgensen 2465 (Herb. Osten 11867] (Ug, Ug, W--1055175); Malvérez 1330 (Es, N)3 T. Meyer 139 (Herb. Osten 22660] (Ug)3 A. io Ge Schulz 1h71 (N), i7ia (N), Ik7h (N); Venturi 9780 (N). Cérdoba: Castellanos s [Valle de los Reartes, Dec. 15, 1919; pat Osten 15250 ] (Ug), 8. on ne [Herd, Mus. Argen t. Cienc. Nat. 31195) (MN); Cozzo 8.n. [Barrio Ja X.1946] (N)s J. Gutiérrez 89 (Gg--352680, N), 8m. do ~II-191,7] (N)3 Hie 8 Sn, [1h-X1.1875] (N), sen, [near Cér- doba, 1.3718 ] (Br, Ca--082829), s.n. [in vicinity of Cérdoba, Nov, "18, 1876] (Ug); Kuntze s.n. {Cérdoba, 15/X11/1891] (W— 701723), sn. (Cérdoba, XII.1891] (N, N); Maldonado Bustos 95 (N), 103 (N, Ws Saas & & Moldenke 19712 (Es, Es, N); 0'Donell & (N); Ruiz Leal 12098 eae cala 10002 (N)j C. Skottsberg s.n. (Cérdoba, 1171071948) (Go); Serbs bile 633 (N), 671 (N); Troncoso 291 (N, N). Corrientes: Alboff s.n. BEE acne [9-11-1096] (HDs Bes & Berti & & dobro se te = N), 3769 (Es, N). ay St Selaart 316 Herb. on del Uruguay] (B); 7. Meyer 9993 (N); Rufz Hnidobro 3588 (N, We), 3637 (N)5 Schwacke si [Concepeion del Uruguay, 1¥/1960] (Ja~ 3)» Formosa: . + JOrgensen 21,70 (Herb. Osten 13675] (Ug) 5 ¥ Morel 530 (Bm); Pierotti 10 [Herb. Inst. Miguel Lillo 72639] (Ca 165859, 0 Ut—-26856b); Venturi 9099 (W—1591L17)5 J. West 84,70 (Can ). La Pampa: Cabrera 6735 (N)s J. Fortuna 1k (Ca), 17 (Ca— 1550) Paso s.n, atone ee April 190; Parodi 13647) (N)- Misi- Been ae 71); Bertoni 1970 peng, 2 ™ 1983 (wi anh Ree » Cuba: Ma Marcus 1962 (N). Dominican ene Eggleston 510) (Dt). aunnge peak bn (Potsdam, 9 August 192] (B). Hawaiian Islands: san. (Apr. 11, 1929] ”(B), | Ford (June 1, 191) (Bi), sn, (rob 28, “ii5) (Bi). Illinois: Herb. Div. Florie. univ. Tl. 177 (Ur). India: Dewan 253 eee Bogor. XV.K-A-XLV-10 (Bz, Bz—26]37, N); Visser Smits s.n. [1916] (Ba—23761). Kenya: Herb. Hort. Reg. Kew. 2615 (Br). “Louisiane: Je He iin cae aon, (Shaw's Garden, Aug. 25, aP (MN, 1, ¥—1325558)- New York: L Le He (San Sk Selle {Aug. h, 1922] (Ba), Ss a. [July 19, 192k) (Ba), son. (Aug. 10, 192h] (Ba), sone [Jy 1965 Moldenke, Monograph of Verbena 301 23, 1925] (Ba), son. [Aug. 7, 1925] (Ba); Burdick 2h (Ba); Hart- 5.n. Oe a Bot. Gard. Cult. Pl. 42722] (N); UcSweeney s.n 8.M. (N. Y. Bo’ Bot. G « Calt. Pls R.305/39} (N); He Ne Moldenke i262 62 OH, 18237 ae Yoldenke & Moldenke 11886 (N); R. C. Schneider s. . (N. Y. Bot. Gard. Cult. Pl. 2228] (N); Seringhaus s.n. [Ne Ys Bot. Gard. Cult, Pl. R.305/39] (N). Nicaragua: Garnier 615 (N); 1180 : ust 1937] (Or—36065). Pennsylvania: H. N, Moldenke 17095 (Au— 122770, St). ee Islands: M. S. Clemens 18433 (Ca—368059). Puerto Rico: E. E. er s.n. [April 21, 192] (Ba); Britton, Britton, & Boynton Bae (N), 8 8712 (N); Moldenke & Moldenke 19536 (N). Russia: fae Hort. Bot. Petrop. s.n. (W—-71 991). Sweden: Herb. Regnell s.n. (Hort. Bergiano, 8/1897] (S). Ah es Herb, Meisner s.n. [20 Jul. 1837] (M). Texas: C. L. Lundell — (NR Rf); Teas sn. ——— June 1926] (Ba). Trinidad: Bai Bailey s.n. [Feb. 16, 1921] (Ba); W. E. Broadway L997 (Ww—L2L7 Tey. VERBENA TENUISECTA var. ALBA Moldenke, Annot. & Classif. List 109 & 113. 1939. Synonymy: Verbena erinoides alba Benary ex Wittmack, Gartenfl. 49: 585. 1900. Verbena erinoides var. alba Moldenke, List Spec. Mold. Southeast. Set 12, nom. nud. 1930. V Verbena rbena erinoides alba Hort, ex Stapf, Ind, Lond. 63 429. 1931. Verbena tenuisecta lus. atbiflora Briq. ex Moldenke, Alph. List Invalid Names 51, in syn. 192. Verbena erinoides wars alba Hort. ex Moldenke, Alph. List Invalid Names Suppl. 1: 2h, in syn. 1947. Verbena tenmisecta alba Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 1 Bibli : 00; Bull. Soc ography: Wittmack, Gartenfl. 49: 585. 1900; Tosc. Ortic. 26: 36, 1901; Moldenke, List Spec. Mold. Southeast Set 10 & 12. 1930; Stapf, Ind. Lond. 6: 429. 1931; G tkins Ins logia 7: 161, 162, & 173. 1936; Moldenke, Annot. & oven List 109 & 11%, 19395 *Moldenke, oe List Invalid Names 8, 1911; Moldenke, Known Geogr. Distrib, 1], . Fito ta ar j Se "32: 576. “9k 55 talina 8. me - Mic Nat. : Moldenke, Alph. t Cite ls 1 re taph st Invalid anes St aa 197; ot Alph. Xho wetas lsiy (19h8), 32. 7h2 & ‘ns (2949) a2 he 97 ks 979 eae: » Phytol 72. 19493 Mo e oes Verbenac , [ed a2 21, 9, 10, ‘101, 5, 16e Be & fon 019} olden bytologia 3: 32 289 & "290 (1950 ool chit ttenden en, Roy. Hort. Soc. Dict. aon “ 2209 = a Hg 302 PHYTOLOGIA Vol. 11, no. 5 Journ. - Hort. Soc. 15: 83. — mena se 12, a 148, 128, oa, “ih, & 376. i Moldenke, Phytologia 8: 123. 1961; Moldenke, Résumé Suppl. 3: 41 (1962), i 3 "962, st ‘1 - 2 (1960), 6: 3 (196 3); ser 7: he 1963; Pearce, New Fls. 20. 1963; Pearce, 196 Garden Aristocrats 20. ies rie. llustrations: witimack, Gartenfl. 49: 585. 1900; Bull. Soc. Tosc. Ortic. Bist 96. 2 This var “espe coro. as a prostrate or trailing wear or perennial, sometimes suffre- tall, , the corolla-tube to 16 ma. wide, and the chromosome aatars 2n = 10. The type of this variety was collected by myself (no. 1163) in a dry sandy ditch at Sneads, Jackson County, Florida, on May 1930, and is deposited in the Britton Herbarium at the New York Botanical Garden. The type of V. tenuisecta lus. albiflora was collected by Teodoro Meyer (no. 142) at Fontana, Chaco, Argentina, where it was not frequent, preferring grassy places rather t loose sand, in October, 1931, and is deposited in the Osten Her us 2 re @ horticultural trade by Chester Jay Hunt (Little Falls, New Jersey), Muller-Seely, Trivetts, and Waterer. Earlier than 1935 seed were w a 25 , 6 inches tall, and sells the seeds 4 at cents per package. rE “a Bailey san. (May 17, 1923], cited below, bears a notation by the collector ‘with appendage or el anther" and E. P, Bradbury s.n. (Coolidge Nursery, 12/12/15] has @ note by him [Dr. Bailey] "with a wing below anther". His noe 12227 has a notation to the effect that a duplicate was sent mt to Dr. Briquet in Geneva. The variety has been collected in dry sandy ditches and Bowe V. canadensis (L.) Britton, V. tenera Spreng., and t, uisecta Briq. D. S. Correll 15325 is a mixture with iypasal We Lv. tenuisecta, For those interested in dating the spread 0 of species and var eties of — enus in new areas, foll e the earliest pe ar e — 1962. ey, Brook, Clinch, Decatur, Grady, Hancock, Jefferson, Lanier, 1965 Moldenke, Monograph of Verbena 303 Lowndes, Screven, Telfair, Thomas, Ware, & Washington Counties — rs Florida: Jackson County -- "1930; Calhoun, Gadsden, & Leon 1 4 Texas: Angelina County -- 1956. Wagenknecht refers to it as "a common weed in sandy soil along ges n all, 51 herbarium specimens, a pres ee ern of An R. Moldenke 310 (Fg, Fg). Clinch Co.: A. R. Moldenke 331 (Fg). Decatur Co.: As Re R. Moldenke 297 (Fg). Grady Co.: A. Re : A. R. Moldenke 300 (Fg); Pyron & MeVaugh 2195 (Gu--16187). Hancock Co.: A. Re Moldenke 393 93 (Fg). Jefferson Cot A. R. Moldenke 4o1 (Fg). Lan- fer Co.: A. | R. Moldenke 325 (Fg). Lowndes Co.: A. Re Moldenke 322 (Fg). Screven Co.: Ahles 54307a (Hi--200787); A. R. Moldenke his (Fg). Telfair Co.: A. R. Moldenke 395 (Fg). Thomas Co.: A. A. Ry Moldenke 307 (Fg, S). Ware Co.: A. R. Moldenke 32 (Z). Wash~ ton Co.: A. R. Moldenke 398 (Fg). ~~ FIORIDA: Calhoun Co.: Davis ¢ Davis 1162, (Hi—115552); A. R. Moldenke 27h (Fg); Be Le wae knecht 3620 (Lw, Lw). Gadsden Co.: Ae Re * A. R. Moldenke enke 292 2 (Fe Task Co.t H. N. Moldenke 1163 (Go--isotype, H— Yoo, HST saci, ¥—~ 3, S--is otype isotype, Ur--isotype, et Ra aA Oo6f ks Re Moldenks 289 (Fg)- OUTSTAMA® Jackson Par.;: Moldenke & Woods , 536 6 (S). Vernon | nm Par.: A. R. Mol- denke 705 (S). TEXAS: Angelina Co.: D. S. Correll 15325, in part (RE). "Pal ee Te Rojas 2528 [Herb. Osten erie (N, Ug). —e- Meyer © 1h2 (Herb. Osten 22659] (Ug)s A. Ge Sc 1473 (N), 2885" (N). Misiones: Bertoni 2265 (N); Ekman 2035 50). CULTIVATED: California: Bailey & Bail 7953 (Ba); E. P. Brad Sen. [Coolidge Nursery, I/inhey (ee . Cuba: L. H. Bailey 12227 (Ba). Java: Herb. Hort. Bot. Bogor. 5 REE (Bz=-23756) « New York: L. H. Bailey son. [May 17, 1923) (Ba), Sen. [Oct. 3, 1926] (Ba). Oklahoma: Stratton 6125, in part (N, St--26160). UISECTA var. GLABRATA Moldenke, Phytologia 4: 451, nom. ENA TEN mabart 1 ry 1960. a 259. ennis iste var. glabra Moldenke, Résumé 7h, 2: 13. ork typical “form of the species in having its aeetrniateg i Phytologia 3: aie 1949; Moldenke, Known Brae of Distrib. Verbenac., [ed. 2], 100 & 200. ag Molden- Teerme 120, LZ, & LT. 1959; sanute, Pytologia 8: 198. on form differs from the typical form of the specie: having white corollas, sins flowers are described as being chads rant or even ve t; -* deposited Uruguay, in November or December, 1892, and is = the Osten Herbarium at the Museo de Historie Natural in Monte- San Fos’ ton base caaaetas pbeky PbO rocky hills and » Stony cliffs, and campos in granitic soil with full ex~ posure £0 relate he ie Posgusd erect: Bt flowe: September to November, Material has been misidentified and dis- aran. to herbaria as V. balansae Brig. and as typical ¥. thymoi~ In ali, 15 herbarium specimens, ae the type, and moun- ted photographs have been examined by m Citations: URUGUAY: Arechavaleta sane [carro de Nico Peres & Si- 31h PHYTOLOGIA Vol. 11, no. 5 erra de Tapambai, Nov.—Dec. 1892; Herb, Osten 3752] ig rs of » F—photo of ae N--isotype, N--photo of type ’ Z—photo of type), sn. [Cerro de Nico Perez, Dec. S92 Herb. Osten 3751] (Ug); Ros 1055 (N), s-N. [Franqueras, Oct. 1900; valeta 29, a pests (De, Ug); Herter 11),5a [Herb. Herter 86120] (N); a Ce 2905 (N); Osten 11636 (Ug, Ug), 22375 (N, 8, Ug), 22375a (Ug), 2 a eles MOPHYLLA Briq. in Chod. & Hassler, Bull. Herb. Boiss., or 2, kz 1061-1062. 190k. synonymy : Verbena incisa Chod. ex Briq. in Chod. & Hassler Bull. Herb. Boiss., sér. 2, k: 1061, in syn. 1904 (not V. incisa Hook., 1839, nor Hort., 1 191.7}. Bibliography: Briq. in NEE & Hassler, Bull. Herb. Boiss., sér. 2, k: 1061--1062. 1903 Briq. in Chod. & Hassler, Plant. Hassler. 10: 48h. 190h; Prain, Ind. Kew. sae 3s 187. shag Mo. , Known Geogr. Distrib. Verbenac i, bb, 19h2; Moldenke, Alph. List Cit, 1: 263 cise), op 375 & 599 (98, and 3: 662 & 693. 1949; Moldenke, Known Geogr. Distrib. Verbenac., ef 2], 99, nad Se & ~, Red Moldenke, Phytologia 3: 290. 19505 e, Ré » 367, & L7h. 19595 aga Phytologia 3 361, 365, 361, ae 597 (A9@h) and 102 97 & n stems rt ea ltebong subacute or obtuse at the apex, ” pinnately lobed, cuneate at the base and prolonged into the petio ole, green on bot surfaces, shabcoeean above, sparsely pilose beneath especi along the veins, the lobes few, cg incis soa be aap at 8 visible aare, spikes short, terminal, to k Gk te and 1. - f the corollas more or ng 5 em. wide exclusive o: ’ “ser fl mm. long, somewhat shorter than the calyx, pose pa at the apory ent on the ely pubescent, ged urceolate at maturity, the tube 6 mm, long, the mouth note ably oblique, the teeth subulate, the anterior ones longer, 1+ le crassate toward the apex, somewhat bilobed, surpassing the aye. by 3 m. after anthesis; cocci sublinear-oblong, ees oy yellowish-green, 2.5 mm. long and 0.8 mm. wide, elevated-reti on the outside. The type of this species was collected by Bail Hassler (n0- 1965 Moldenke, Monograph of Verbena x5 295) in thickets at Caraguatay, Paraguay, in October, and is de~ os ted in the Delessert Herbarium at the Conservatoire et Jardin grassy places, on campos, in thickets, and "common" among trees, at 50 meters altitude, flowering from September to November and in March. The flowers on Hassler 3295 are described as Soh g on JOrgensen 26% as "vivid-rose", on Grier 115 as "purple*, on J0rgens en 3769, Lillo 8577, and Rojas 1362 as as "violet". Material has been misidentified and distributed in herbaria under the names ¥. calliantha Briq., V. erinoides Lam., V, ae ana Briq., and V. f. incisa Hook. JUrgensen 3769 is actually a ture with V. hasslerana | erana Briq. ani eg ; (1908) says: "M. Chodat a avec raison fait soivre 7 Subulées (courtes et simplement acuminées dans le v. incisa), en- fin par la corolle beaucoup plus petite, simplement pubescente extérieurement (deux a trois fois plus grande . étalé sur eus la m@me section (Verbenaca) SS V. incisa: il se place dans la 2. ty Ro Rai: Hassler 3295 (Macbride photos 24700) (Kr-—- Nh to of type, N--photo of type, Qu--photo of y deg uaen 2 3769, in part [Herb, Osten 22219) (My 8, Ue)3 T. nojea 2385 ae [es Herb, Osten 7900] (N, Ug)» ARGENTINA: Formosa: 2636 [Herb, Osten 14552; Herb. aa’ Miguel Lillo 31373) (N,N, Te, “W--1065616) . pp pasa Grtmer 115 [Herb. Osten 23176) (Ug). Tucunén: Lillo 8577 (Herb. Inst. Miguel Lillo 31568) (N)5 Venturi 2651 (W—1591262). xVERBENA : vedo an a areorgit £9 wen Ss cytologs Dernen, 7: 170. 19%. Verbena necmexicana (A. =e Small x V. urticifolia 316 PHYTOLOGIA Vol. ll, no. 5 L. ex Moldenke, Résumé 370, in syn. 1959. Verbena urticifolia L. x = neomexicana (A. Gray) Small ex Moldenke, Résumé 377, in . Cytologia 7: 170. 1936; ie age Phyto- Lecie ie FER (1981) and and a5 ha oi i Moldenke, Biol, Abstr. 30: « 19563; Moldenke, Am. Midl. Nat. 59: 362. 1958; Moldenke, Ré- 370, 377, & L7h. 19593 Moldenke, Phytologia 8: 121 (961) and 10: 178. 196k. the hybrid between V. urticifolia L. and V. neomexicana aa ae uty produced artificially by Dermen in Massachusetts in 1936. The two parental species do not grow together naturally ere, as far as I am aware, although both occur in Texas a variety of V. neomexicana has been found as a waif in Baltimore, Maryland, where V. urticifolia is common. It is not very probable that this hybrid will ever be found in the wild unless the two parental species invade each other's territory in the future. 3, fig. Syten a—d. “19355 A. W. Hill, Ind. Kew. Suppl. 9: 295. Worsdell, Ind. Lond. Suppl. 2: 486. 191; Moldenke, Known aa Distrib. Verbenac., [ed. 1], 34 & 102. 13425 ae Castanea 13: 116. 1948; H. N. & A. L. Moldenke, Pl. Life yee Moldenke, Alph. List Cit. h: 982. 1949; Moldenke, pei a ae eae [ed. 2], 71 & 200. 19h9; Moldenke, Bh" ‘a1 - 1959. Illustrations: Svenson, Am. Journ. Bot. 22: 271, pl. 3, fig- 1 a-—d. 1935. Glabrous erect herb, to 1m. tall; s tetragonal, about 3 mm. in diameter; rt Hi internodes Tt em. long; branches rigid, suberect, paniculate; leaves decussate-opposite, deeply Pa evolute along the margins; spikes elongate, to CH. long, ntti oe the apex of the branchlets, erect, filiform, the lower flowers becoming remote, often separated at maturity of the fruit by a vat of 15 mn.; bractlets ovate, about 2.6 mm. long, ciliate along th gins, otherwise glabrous; pedi out -5 mm. 1 flowers very sm calyx about 2.7 mm. long; eg : ac o os bearded in the throat, the limb G-parted, the lobes 1.5 mm. 1oné, the 3 lower ones slightly emarginate at the apex; stamens 4, ous; anthers broadly elliptic, 0.3 mm. long, equaling oF longer than the filaments; filaments ee Oo1-—003- ons egy ee bilobed; fruiting-calyx tetragonal, gla onset ose, of the teeth with incurved prot 0 $505 shorter and not curved; cocci h, brown, 1.5 mm. long, i inently venose on the outer surface, white-roughened on the ¢ 1965 Moldenke, Monograph of Verbena 317 missural maniee The type o s species was collected by Henry Knute Svenson (no. a in pravitiy s0il below the escarpment, at an altitude 1 @ herbarium the Brooklyn Botanic airkee? me ene e siothe that "It dit- fers from V. littoralis in its eae — narrow leaf segments, broader bracts, and m " It in honor of Charles Hasidns aecaaad a (2859-19) Ds a zool- ogist, director of the New York Aquarim, ed plants in Alaska and on the Galapagos Islands. This pedro uniel is knom only from the type collection and has been collected in an- thesis in April. Svenson cites also Stewart 3317, 3318, and 3319 from Albemarle Island, but the first of these is regarded by me as Vv. elabeata “RR "and the last two as V. galapagosensis Mol~ am all, 2 herbarium specimens and 3 — photographs of the es collection have been examined by m Citations: GALAPAGOS ISLANDS: Tedafatignblos Svenson 249 (F-- Photo of isotype, Gg-~237859~-isotype, N--isotype, e, N--photo of isotype, Si--photo of isotype, Z-—-photo of isotype). VERBENA TRACHEA R. A. Phil., Anal, Univ. Chile 90: 609. 1896. Synonymy: Verbena traquea R. A. Phil. ex Moldenke, Résumé 377, in syn. 1959 Bibliography: R. A. Phil., Anal. Univ. C os 90: 609. 1896; Durand & Jacks., Ind. Kew. Suppl. 1: 51. 1906; Moldenke, Known Geogr, Distrib. Verbenac., asi as Wwe & — ‘si2) 9% tea. 2h, an & 200. 19h9; Acevedo d 3 ; Rapes « lant. a Fay 10: 137. 196k. Suffruticose sass low, much branched, hirtous, with short Spreading hairs most dense on the branches; stems about 20 cm. tinate, the rim with very short triangular teeth as the calyx, glabrous on the outer surface; pa to 3.5 mm. The type of this greet species was collected by Chrie- tien Ludwig Landbeck in the Cordillera de C olchegus, at an alti- tude of 2000 meters, Colchagua, Chile, in December,'1861, and is oe in the herbarium of the Museo Nacional ay weateea teat ural at Santiago. Philippi says (1996) “La V. radicans difiere alt Nacsa Setoasateees lla Kth. por sus hojas mui Pequefias, del largo de solo 7 =e gus espigas en cabezu- 318 PHYTOLOGIA Vol. 11, no. 5 ela, etc." He places V. trachea in the section Glandularia of the genus. Only a single photograph of the type collection has been seen by me thus far, Citations: CHILE: Colchagua: Landbeck s.n. [Herb. Mus, Nac» Hist. Nat. Chile 2515 & 54732] (N--photo of type & isotype). XVERBENA TRANSITORIA Moldenke, J kewl 5: Sit 1955.6 Synonymy: Glandularia lacini ca Schnack & Covas, Revist. Argent. Agron. rr "ere z oa. oe: sg er laciniata x megapotamica Covas & Schnack ex Moldenke, Phytologia 5: 133, in syn. 1955. Verbena megapotamica Spreng. x V. tenuisec- ta Briq. ex Moldenke, Résumé 369, in syn. 1959. Verbena tenuisec- ta Briq. x V. megapotamica Spreng. ex Moldenke, Résum 6 376, in ~ Be ti graphy: —— & “pst oath di Pent ave Agron. i 222 2h. 1 19h5; Schnack & Gonz . Agron Toss os, Phytologia “ eT "G954) = a3: "133. 15555, olden ke, Biol, Abstr. 30: 1093. 1956; Moldenke, Am. Midl. Nat. 59: 362. 1958; ioiaae, Résumé 22), 296, 0, 376, & t lithe 39593 inicene, résumé Suppl. 2: 12. 1960; Moldenke, Phytologia 8: 12 (1961) and 10: 133. 96h, is the hybrid between V. temuisecta Brigq. and V ines. Spreng. produced artificially by Covas & Schnack i in 5 Anger tina in 1945. The two parental species grow together in neers? e 8 areas. It should have considerable horticultural merit and de~ serves further study and reckon rare Schnack & Covas ap~ parently misidentified the one parent as V. laciniata (L.) Briq- instead of VY. tenuisecta, and I mistakenly accepted their misiden- tification in my earlier references to this plant. TRIFIDA H.B.K., Nov. Gen. & Sp. Pl. 2: 273, Beg Synonymy: Verbena trifida Humb. ex Spreng. in L te "7 ed. 16, 2: 749. 1825. Verbena trifida Humb, & Bonpl. ex Steud, Nom, Bot., ed. 2, 2: Tol, Til. Yona trifida Humb. & Kunth ex D. Dietr., Syn. Pl. 3: 60h. 1843. ena trifida Kunth ex ep in A. DC., Prodr. 11: 539. 187. a ee aeitae yifidi Kunth ex Mol ke, Alph, List Invalid Names 51, in syn. 19 wt t HB.K., Nov. Gen. & Sp. Pl. ar 273, Pie 13h. j Steud., Nom. «, od. 1, 874. 18213; Spreng. *3 Veg., ed, 16, $ 7h9. 1525; Steud., Non. ; be, 2 23 751. 18lit} D. , es 3: ° the, Pl. a 169 Schau, in A. w., Prodr. 11: 5 * aXto, Fl. 1: ‘ 63 1850-1851; Lem., Jard. Fl. 1: Misc. 85. 1851; , olden . 192; Am. Mo. Bot : 3h2 & 356. 19335 ke, a ogee tes ———_ Verbenac., et 1). = & — ae if nk & 158 AKG) Be SST CSUR), code 4008 & 107k 9495 Be 1965 Moldenke, Monograph of Verbena 319 & As L. Moldenke, Anal. Inst, Biol, Mex. 20: 1h. 199; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 62 & 200. 1949; Molden- ke, Phytologia 3: 451. 1951; Moldenke in Chittenden, Roy. Hort. Soc, Dict. Gard, 6: 2213. 1951; Moldenke, Résumé 0, 69, 377, & Supp logia 9: 33 & 39 (1963) and 10: 140. 196; Moldenke, Résumé Suppl. 8: 3& 6. 196k. : ' aa Tllustrations: H.BK., Nov. Gen. & Sp. Pl. 2: pl. 13h. ag Paxt., Fl. Gard. 1: 169. 1850—-1851; Lem., Jard. Fl. 1: Misc. 85. 1851. Shrubby, to 1m. tall, canescent throughout; branches tetrag- onal, clustered in the leaf-axils, hirsutulous; leaves decussate- opposite, clustered, sessile, 1.5--2.5 cm. long, trifid or nearly 80, cuneate at the base, strigose-hispidulous on both surfaces, the segments lanceolate, acuminate at the apex, revolute-margined, tions conspicuous, subulate, subequal or slightly unequal; corol- la purple or pale-violet, medium in size, its tube slightly long- re: than the calyx, the lobes of the limb rounded, enarginate at apex, The type of this perplexing species is said to have been col- lected by Friedrich Heinrich Alexander von 1dt and Jac- @es Alexandre Bonpland between the village of Atotonilco and the Then draw a line to the west to Actopan. On this line, no a ri: a all tom named Sanctorum, and there is ver named Amajac. On this river is the Puente de Dantoe or Puente de Dios, a natural bridge 80 meters high. Santa Virginia is just a small village in the region.® It seems hardly probable that this distinctive plant should in ng areas. not again been found in Mexico, it seems probable that the Mexi- can locality is the result of an error in labeling on the part of Humboldt and Bonpland. I suspect that the species is endemic to Colombia and that the ype came from there, too. The species is Said to have been introduced into cultivation 1818 88 @ greenhouse plant and is described as "a curious plant". It has been found in Colombia at altitudes of 1800 to 3750 meters, in March, July, and September. A vernacular nme re- Corded for it is "sinamomo™, re acon (1847) cites Humboldt & 320 PHYTOLOGIA Vol. 11, no. 5 Bonpland s.n. and Hartweg 1353 1353 from the Kunth and Berlin herbaria. In ~~ In all, 10 10 herbarium specimens and 1 mounted illustration have been exam: xamined by me. Citations: COLOMBIA: Boyac4: Cuatrecasas 127 (W--1772927), i720 applets Linden 129) (F—876775)+ Cundinamarca: Apol- mr & (W—-603087) ; Hartweg 1353 (Lu, N)s Holton se Cartel TES Huertas & Camargo 733 (F.-1291995)1 Triana 2057 (Br, N). MOUNTED ILLUSTRATIONS: Fig. 103 TRIFIDA var. DESERTICOLA Moldenke, Phytologia 3: 281. 1950. liography: Moldenke, Phytologia 3: 281 & 290. 19503 Molden- sigh neers 122 & h7h. 1959. s variety differs fron the typical form of the species in ‘atin & more branched and diffuse plant, with more slender and much weaker branches, and with the divisions of the leaves uniform ly oblong or linear, 1--3 mm. wide, and more obtuse at the apex. The type ser the variety was collected by Ernesto Barros Valen- zuela (no. 8010) at Quebrada La Chimba, Antofagasta, Chile, on September 20, 19h0, and is deposited in the Britton copkt: at the New York’ Botanical Garden. The variety is knom t far only from the type specimen a Fabamaae: caren Antofagasta: Barros Valenzuela 8010 (N— ype XVERBENA TRINITENSIS Moldenke, Résumé Suppl. 3: 1h, 37, 39, & Wy rus BE side (1962), nom, erbena var, subbipinnatisecta Kuntze, PR saya Pl. 5 (2): 257. 59 98. Verbena tenera x incisa x or cides Hegi, Illustr. Fl. Mittel-Eur. 5 (3): 2240, in textu. 1927. » RE in syn. 1959. Verbena chamaedryfolia x subbipinnatisecta Kuntze ex Moldenke, Résumé 362, in syn. 1959. Verbena incisa Hook. x = 1 temuisecta Beta. ex Moldenke, Résumé Suppl. 3: 39, 39, in syn. 1962+ Verbena tenvisecta Briq. x V. incisa Hook. ex Moldenke, Résumé Suppl. 3: 1, in syn. 1962. Bibli phy: Kuntze, Rev. Gen. 3 (2): be 18985 ng th - Mittel-Bur, 5 (3): met Sy “W32h; Mold Alph. Cit. 2: 628. 19h8; Moldenke, Ré 2. 19595 hstesbes pl. 3: 14 39, & hl (1962), bz 19 (1962 : 6B. 19623 Moldenke, Phytologia 91 ih = yy & 279» 1965+ The type of Car. 8 Kuntze at T in sout and is deposited in rinidad, hern Paraguay, — Britton Herbarium at the New York Botanical Garden. I rounre. © plant as a natural hybrid between VY. incisa Hook. and V. teo uisecta Briq. Osten, in a memorandum written at moore’ in January, 193s says "Wie von OK angefthhrte V. chamaedryf. . subbipinnatisec 1965 Moldenke, Monograph of Verbena 321 OK, wegen der sich et den Kopf zerbricht ist der spontan Bastard: Verbena [Nobiles, chamaedryf.] Briquetiana (rot) x [Glan- dularia, erinoides] tenui Saati (violett) Briq. mit rosa BlUten, den auc! auch ich loco classico bei Trinidad zwischen den El Eltern traf (auch sonst in Paraguay nicht selten)." Hegi Seftl says that this hybrid has given rise in gardens to bunte Blumisten Verbenen” and "italienische Blumisten Verbenen" forms. rasdn, geen are what we now usually regard as V. ten- sera var, maonetti Regel, and are quite different morphologically fron Kuntze's and Osten's plants. specimen has been examined th ‘ar by me. Only the type : > pares PARAGUAY: Kuntze s.n, (Trinidad, aa paraguay] (N= ype VERBENA TRISTACHYA Troncoso & Burkart, Darwiniana 7: 208—215, bliography: Troncoso & farkart, Darwiniana 7: 208—-215, fig. 20 & Burkart, Darwiniana 2 211, fig. 2 c—p, & 213, fig- 3. 196; Schnack & coves, Bin Pa 10 Agron. Glabrous branched herb, probably biennial, erect, to 70 cm. philous; root perpendic cular, medium in size, with a- bundant fibrous superficial branches which extend from the crown and from the decumbent portion of the stem, with abundant adven- titious hairs; tens short-decumbent at the base (the decumbent rtion to 8 am the remainder erect, conspicuously fistu- t Ones linear and smaller, 2—l; om. long and 2-—l mm. wide, all a- bes at the apex, attenuate to the sessile base, not auriculate ee geen prc coarsely serrate-dentate along = margins with in 322 PHYTOLOGIA Vol. 11, no. 5 beneath; inflorescence dichasial; spikes terminal, biparous, mostly 3, rarely 1 (on less vigorous shoots) or up to 7 (ee very vigorous branches), pedunculate, subcylindric, dense-flowered 1,8——2 em, long during anthesis. and 7-—-8 mm, wide, elongate to 7 cm, after " anthesis, forming secondary lateral groups at the ends of lateral branches on very vigorous shoots, separated by leaves from the terminal group} flowers 7-—8 mm. long; bractlets lanceo- 6.5 mm. long » 5-cO Sag 9 on the pes otherwise subglabrous, the r , 4 of the teeth bequal, the 5th one very small; corolla pated ’ its ¢ tube 6.57.5 mm. long, sli sing the calyx, puber- gh uw on the exserted portion outside, sparsely pilose throughout ee otlay densely so at the throat, the limb exiguous, 2--2.5 mm. in diameter, the 5 lobes radiate, subequal, glabrous, obscurely emarginate at the apex; stamens |}, didynamous, inserted in the upper i ne the corolla-tube; filaments short; thecae ovate, separate; style filifom, 6--7.5 times as long as oa ay bi- lobed at the apex; stigma short, obtuse; ovary g » be celled; mature fruit separating into into l-seeded pox ant at maturity; ned achene-like, subcylindric, dark-nigrescent, 3— 325 mm. long, 0.6--0.7 mm, wide, striate below, reticulate on the back above 0. SCHAUER.....A esta serie emparentadas con la que criber, a saber: V. stellarioides CHAM., aaah dos variedades, Vs littoralis H.B.K., V. intermedia GILL. & HOOK., V. montevidensis SPRENG......., V. rigida SPRENG etc. De todas ellas 0s e diferencia af (sin entrar a considerar los caracteres , y peros o escabrosos (excepto en V. stellarioides y V. montevider” sis), mientras que en V. tristachya es ¢ co u obscuramente tetrégono, completamente liso y glabro. in jaanto* ala penne cencia, ve littoralis, V. intermedia y V. montevidensis florescencia multiespigada, mientras que V. F, stellarioides posee espigas terminales solitarias, rara vez hasta 2 6 5 espigas por 1965 Moldenke, Monograph of Verbena 323 v4stago; en la nueva especie es t{pica una trfade de espigas. Puede formarse una serie de especies con progresivo aumento en el nfmero de espigas por vastago: V. stellarioides (1—3 espigas), V. rigida (3), V. tristachya (1~-7), V. intermedia, V. littoralis, lari nodio terminal glabro, flores menores con limbo reducido, corola apenas pubescente en la parte superior del tubo, brdcteas menos We sas edliz ligeramente pubescente. La difundida V, littoralie se dif- erencia por las espigas delgadas, de 3-—1, mm, de diém, (en V. tristachya miden 6--8 mm. de didm.), flores de —-5 mm., con el tubo corolar de 3,5—- mm. de long., c&liz de 2,3--2,8 mm. de long., cubierto de pelitos breves uniformemente repartidos sobre gada, V. intermedia, otre de las especies afines...se dist : aparte de los rasgos ya camentados, por flores m&s grandes y Vistosas, de color viold&ceo intenso, limbo corolar bien desarrol~ lado, c&liz y br4cteas con pelos glandulosos, estilo breve y es- tambres fijos en la mitad inferior del tubo corolar....Creemos interesante hacer notar que ningfin autor cita el carécter fistu- loso del tallo en el género Verbena. Tan es asi que en un prin- cipio crefmos que éste era un cardcter especifico de la nueva especie, pero al efectuar cortes en los tallos de las especies afines, pudimos observar que V. stellarioides posee también tallo completamente fistuloso, aunque de cavidad menor. En V. littor alis las ramas superiores y los tallos jévenes son macizos, ob- Servaéndose que los tallos m4s gruesos y los entrenudos inferiores Son algo fistulosos, pero con una gruesa capa de - V. in termedia y V. montevidensis tienen los tallos macizos, £1 tallo El estudio anatémico del tallo de V. tristachya presenta c caracter{stica interesante, que le capa quima clorofilia~ nO 6s continua, es decir, que no interrumpida en lo 32k PHYTOLOGIA Vol, 11, no. 5 mismos autores consideran también diferencial y que en esta es- pecie no lo es, es la relacién longitud del estilo a altura del ovario. SCHNACK y COVAS.......consideran que en Verbena el esti-~ lo es tan largo o hasta tres veces m4s largo que el ovario, en Gl a, por el contrario, es varias veces m4s largo. En V. tristachya es de 6 a 7,5 veces m4s largo que el ovario, y en V. stellarioides es 10 veces y media mayor, Por los demds caracter- es, en cambio, V. tristachya es una auténtica Verbena. Seria in- teresante para resolver este problema, efectuar el estudio cito- légico de la nueva especies, el cual no pudimos re las semillas que g aron." The species has Pte collected only in "pajonal" and on river banks, flowering in December. In all, 6 herbarium specimens , in- cluding the type, pir 2 mounted photographs have been examine itabiedhe ARGENTINA: Lechiguanas Island [Entre Rfos]: Burkart 156hh (F—photo of type, It--isotype, N--isotype, N--photo of type, Si--type). Misiones: Ekman 202) (Mi, N, S). — TRTPEROUATA Re A. Phil., anal, Univ. Chile 90: 615, pl. 1. Synonymy : ae biternata R. A. Phil. ex Moldenke, Résumé 358, in syn. 1959. histececdsty: R. A. Phil., Anal. Univ. Chile 90: 615, 1895; Durand & Jacks., Ind. Kew, Suppl. 1: 51. 3906) Reichs, ‘i. chile 3: 277 & 279. 1910; Stapf, Ind. Lond. 6: 431. 19 Lden= Known Geogr, Distrib. Verbenac., fed. 1), = tar: “loz ae a fed. 2], 102 & 200. 1949; Acevedo de Var Nat. Chile 25: 69. 1951; Moldenke, “Shedae 4i0 9 368, & & ‘The "19595 Mufioz Pizarr r Espec. Plant. De Deser e Philipp. lil. 9 ° Illustrations: R. A. Phil., Anal. Univ. Chile 90: pl. 1. 1895- he orymbs; petioles very short, about 3 mm. long, ampliate; leaf- blades semicircular in outline, Lg polgnd! or triternately dis sected, the larger ones about 9 m. long and 13 m. wide, the : é 5 Li ib He : E Be i B Arauc a, Arauca, Chile, in January, 1887, and is deposited in the oun of the! Museo Nacional de Historia Natural at Santiago, Chile. 7 i) error. Reiche (1910) says: "en las cordilleras de la araucantas | parece planta escasa. Observacion. La V, triternata iV. digi tats 1965 Moldenke, Monograph of Verbena 325 son mi poco diferentes; el cardcter distintivo featede.. « ia base del estilo debe comprobarse en muestras mas en vista de oa localidades nativas, distanciadas en "L3 idan no me atrevo reunir las dos a 28 Only a photograph of the type ha Be ha seen thus far by me. Citations: CHILE: Arauca: R. A. Philippi s.n. [La Cueva, Jan. 1887; Herb. Mus. Nac. Hist. Nat. Chile 83} (N—photo of type). ee Amn, Mo. Bot. Gard. 20: 322-~323. 1933. bliography: , Ann, Mo. Bot. Gard. 20, 2hh, 25, 2h8, 311, 322--323, & 6. 1933; Cory, Texas Agr. Exp. Sta. Bull. a9." PiPH: A. We Hill, Ind. Kew. Suppl. 9: ees 29385 tome own Geogr. Distrib. versevits {ed. 1], 13, 1h, 1942; Moldenke in Lundell, Fl. Texas 3 (1): "tg & 8. At vei Yol- » Known Geogr, » [ede 2], 255,235 19495 Moldenke, Résumé 30, 31, 0, & 7k. 1959; Moldenke, Phytologia fos 92, 98, 102, & 10h. 196k. Herbs stems 15-20 cm. long, decumbent-ascending, branching, ; ovate, 2—-l, cm. long, te a into the petiole, trifid, with the segments again — 8d or coarsely crenate-dentate, appressed-pubescent or s lose above, hirtellous beneath; spikes short-pedunculate, protrud- o- the base, hirsute, f. dular, teeth about 1.5 mm. long; corolla sain tonktin: the tube about 11 mm. long, puberulent on the outside, the limb 6—10 mm, wide; and is deposited in the herbarium of the Missouri Botanical Garden at St. Louis. The type collection is inscribed "Real County" on the labels, but the locality is in Edwards County. » species is own, from 1 from May Edwards oe eee Bexar Counties. It has been collected in fruit J ugust, and Nagra? at altitudes of 1225 to 2900 fest Marsh says acim wering in spring". Cory 509 consists of splendi Specimens ——— Ss Wright 455 cited below from El Paso County, Texas, is may ac have been made in one of several western counties of the state. The Colsctor undesignated 8.ne s.n. cited below is fran "Leon Spring" and 326 PHYTOLOGIA Vol, 11, no. 5 me B. C. Tharp, in a letter to me, states that this is probably eant to "be Leon Springs, in Bexar County -- it is annotated by reney as V. delticola Small with a question, Ce Wright 1503 is a mixture with V. bipinnatifida var. latilo- bata | Perry -- it was first identified as V. aubletia L., then as V. anbrosifolia Rydb. It may have been collected in western Tex- as instead of "N, Mexico" as stated on the label. E. F. Castetter, in a letter to me dated January 31, 1955, says that in twenty- seven years of botanical collecting in New Mexico he has never seen this species in that state. Material of V. tumidula has been misidentified and distributed in herbaria onder the names V. ambrosifolia ier ve ae Le, V. ciliata Benth., V. delticola Small, V. drummondii T Baxt., lambertii | Sims, and V V. pumila Rydb. Perry foe rated gels cites the following 3 additional specimens not yet. seen by me: TEXAS: Edwards Co.: B. J. Palmer 13512 (5—trre)> El Paso Co.: mt Wright 455 (G). NEW MEXICO: County undetermi C. Wright 1503, in part (G). The C. Wright a3 which she se from NY" is actually deposited in the Torrey - She ¢ ments that "This pia a pe undoubtedly belongs to she section Glandularia, although apparently the anthers are glandless. Super ficially, it resembles V. Gooddingii var. nepetifolia, but the spikes are not so large nor s0 6H Its distinctive characters are the ventricose nutlets and the inflated fruiting calyx. The brac are broader and peers than those of nearly related species of Ropes roup." In all, 21 hathaeiab specimens, haeiating material of the type collection, have been examined tations: TEXAS: Bexar Co.: Galtectar undesignated {Plains near Leon Spring, Sept. 7, 1952] (N). Edwards a oon 38940 (N)s E. J. Palmer 13512 (B—daotype, Co pa aah j Parks & Cory 2h83 (Tr), 26651 (Tr), 26826 (Tr). El Paso 2 Wright 155 (W--56208). Real Co.: Cory s.n. [7-19-19%] (ua) - L. Fisher 50103 (Ew, Gg-~3812h6); Parks & Cory 19093 (Tr), 1909h (tr); Tharp s.n. [Vance, 9/11/29] (Au). avai Ide Co.t Cory TY509 (au). “Val Verde Co.: XY. Anderson s.n. [Del Rio, Spr 1921] (Wi). NEW MEXICO: County undetermined: C. Wright 1503, resets (T, W--71927). MEXICO: Coahuila: Marsh 150 (Au, A Tamaulipas Eo H. Bartlett 10030 (Mi). VERBENA URTICIFOLIA L., Sp. Pl., ed. 1, 20. 1753. acad. Synonymy: Verbena p , foliis urticae Dodart, Mém. in syn. 1687. Verbena peregrina foliis urticae Dodart apud F- 1965 Moldenke, Monograph of Verbena 327 Herm., Fl. Lugd.=Bat. 5°, in sy>. 1690. Verbena recta canadensis siue vir; “giniana maxima urticae foliis J. Morison, Plant. Histor, Univ. Oxon. 3: : 3: sec. ec. ll, pl. 25, fig. 3. 1699. ites americana urticae folio Dodart apud J. Morison, Plant. Histor. Univ. Oxon. 32 "Bn '[=h18}. 1699. Verbena cretica urticae folio Zan. apud ; Morison, Plant. Histor. Univ. Oxon. 3: "Oo" [=18], in syn. 1699. Ve Verbena urticaefolia canadensis Tourn., Inst. Rei Herb. 200. 1700. Verbena lamii folio Barrel., Plant. Gall. Hisp. 30, pl. 1146. 171;. Verbena urticaefolia, canadensis Tourn. apud Bar- 20. Spicis filiformibus L., Hort. Cliff. ll. 1737. Verbena canaden- sis, f folio urticae Boerh. apud L., Hort. Cliff. 11, in syn. 1737. Verbena re recta canadensis seu virginians maxima, urticae foliis Moris. | apud L., Hort. Cliff. 11, in syn. 1737. Verbena foliis ovatis serratis, ae £114 formibus paniculatis A. van A. van Royen, Fl. Le. Leyd. Prod. 327. 17h0.- Verbena canadensis, folio urticae A. van an Fl. Leyd. ei. og in - 4 0. Srecto, spicis pyissesins a Dieta Le, Spe Pl., ed. 1, W, in syn syn, 1753. Verbena recta canadensis, urticae foliis Moris. apud L., Sp. Pl., ed. 1, 20, in syn. 1753. Verbena tetrandra, spicis f414formibus paniculatis, foliis indivisis serratis pet- iolatis E., Sp. Pl., ed. 1, 20, in syn. 1753. Verbena tetrandra, Spicis filiformibus paniculatis, foliis indivisis is serratis pet- jolatis L. apud J. F. Gronov., Fl. Virg Virg., ed. 2, h. . 1762. Ver- bena recta canadensis » 8. Virginiana maxima urticae foliis Moris. Sabbat in Martelli, Hort. Roman. 3: n, pl. 55. 1775. ratenn tetrandra, spicis filiformibus, paniculatis; foliis indivisis, Ovatis, serratis, acutis, petiolatis L. apud Poir. in Lam., n Lam., En- Rane uéth. Bot, 8: LB, in syn. 1008. Verbena foliis oretis caule erectos spicis filiformibus, paniculatis L. apud Poir. nn ceteneneetine > Lan mayel. Bek , in syn. 1808. Verbena (urtici~ culis div divaricatis; spicis . spicis filiformibus, i apud Poir. in Lam., Encycl. Méth. Bot. 8: 548 Verb — Moris, apud 328 PHYTOLOGIA Vol, 11, no. $ in 1808. Verbena diffusa Hort. ex Poir, in Lam., Encycl. cat Bot. 8: 550. 1808. Verbena urticaefolia L. ex Dum. Cours., « Calt., ed. 2, 22 623. “811. Verbena diffusa Willd. ex J. a in Rees, Cycl. 36: no. 18. 1817. Verbena diffusa Desf. ex Spreng. in L., Syst. Veg., ed. 16, 2: 748. 1825. Verbena tetran- dra, spicis filif, paniculatis, fol. indivisis ovat. acut. serratis petiolatis L. apud H. E. Richter, Cod. Bot. Linn, 35. 1835. Verbena fol. ovatis, caule erecto, spic. filif. panic. lL. apud H. EB. Richter, Cod, Bot. Linn. 35. 1835. Verbena lamii Tamil fol. Barr. apud H. E. Richter, Cod. Bot. Linn. 35, in syn. eevee OS. Ver- Linn. 35, in eg 1835. Verbena peregrina, fol. urtic. D Dodart apud Peterm., Cod. Bot. Linn. Ind. Alph. 196, in syn. 1640. Verbe- na recta canadensis etc. Moris. apud Peterm., Cod. Bot. Linn. . Inde Riph. 196, in syn. 100. Verbena diffusa Lam. ex Eaton & Wright, N. Am. Bot., ed. 8, 470, in syn. 1640. Verbena recta canadensis urticae folio Mrie. apud Walp., Repert. Syst. Bot. jot. ks 21, in syne 1845. Verbena diffusa Poir. ex Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 1178, in syn. 1895. Verbena urticifolia var. simplex Farwell, Papers Mich. Acad. Sei. 3: 103. 1924. Verbena urticifo syn. 19h1. Verbena urticafolia L. ex Moldenke, Suppl. List valid Names 10, in syn. 19 Verbena urtricifolia L. ex wide ke, Alph. List Invalid Names 51, in syn. 1942. Verbena urticae~ folium L. ex Moldenke, Alph. st, eal sa 8 Bi, in syn. 1925 E. J. Palmer, Handb, Nat. Hist. 297 & 663. 1949. Verbena urticS- folia x stricta Gates (in part) ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 1947 (not ¥. urticifolia x stricta Eggert, 1937, nor Norton, 1958]. Verbena urticifolia var. urts- cifolia (Moldenke] in Gleason, New Britton & Br. Dlustr. Wis; print. 1, 3: 129. 1952. Verbena Srtieetel* var. hirsutior Engelm, ex Moldenke, Résumé Suppl. ih Li, in syn. 1962. Verbena ee Llongissimis, laxis, culat , vald’ diffusis; foliis folie vato-lanceolatis, subpubescentibus, ate caule frutescente, anpaae Poir. ex Moldenke, Résumé Suppl. 4: 19, in syn. 1962. Verbena urticae folia canadensis Tourn. ex Moldenke, Résumé Suppl. k: 20, in syn. 1962. Verbena urticifolia Gray & Moldenke, Résumé Suppl. : 20, in syne 1962 Verbena scprielte- lia McCann ex Moldenke, Résumé Suppl. 8: 5, in syn. “syns 196k. Wer bena urricaefolia L. ex Moldenke, Résumé Suppl. 8: 6, in sy? 1965 Moldenke, Monograph of Verbena 329 1964. verbena urticifolia var. typica Haw. ex Moldenke, Résumé Suppl. syn. Bibliography: Dodart, liém. Acad, Sci, Paris kz 327, Lf 317. 9; Zanoni, Hist. Bot. 203-—20, fig. 77. 1675; Ray, Plant. 5%. 16865 P. Herm., Hort. Acad. Lugd.-Bat. Cat. ay. il P. Herm., Fl. Lugd.-Bat. 55. 1690; R. Morison, Pl. Hist. Oxon. 3: %,08" vee), sect. ll: pl. 25, via a 3- 20773 ae Inst. Rei Herb. 200. 1700; Boerh., Ind. "git. - Hort. Acad. Lugd., ed. a 75 1710; Barrel., Plant. oe o 5 ant rt. Acad. Lugd., ed. ed. 3, 1: 187. alt ee poe Reva pa i J. F. Gronov., & Virg., ed. 739; Fl. L 327, 17h0; pee Hort. row He 9. "TTBS Yizhel, int "Plant. Hort. Caes. Florent. 98. 1748; L., Sp. Ber = i, em. Serv, l'Hist. Pl., ed. 3, 627, pl. 35. 1758; Ls, Syst. Nat., ed. mia eras ted J. F. Gronov., Fl. are, ed. 2, h & 7. 1762; iy Sabbat. in Martegii, Hort. Roman. 3: 11, pl. 55. 1775; Christm. & Panzer, Volist. Pflanzensyst. Houttuyn 5: 127--128. 1779 he ka Stirp. Nov. 1: 22. 1786; Lam., Tabl. Encycl. Méth. Bot. [Illustr Gen.] 1: 57. 1791; ehh Sp. 2 : Ao 1797; C. Murr. Brit. Gard. 2: 79. 1799; Michx., Fl. Bor.-am., ed. 1, 21 15. 1803; Poir, in Lam., Encycl. grey Bot. 8: 548 & 550. 1808; Dum. Cours., Bot. meagre ed. 2, 2: 623. 1811; Pursh, Fl. am. Sept. 2: )16~—)17. 181); J. Sm. in Rees, Cycl. 36: no. 16 & 18. 1817; Michx., Fl. ed. 2 Wrig ed ot. Linn. gk idph. 156. 181; Steud., Nom. Bot., ed. 2, 2: 750 & 751. 16hl; Pl. 3: 601. 18,3; vis Torre, Fl. fs caiye Ni ft. Gs 52 t & 570. 183) Male, Repert. Syst. oom 20 & 21. 1845; W ° — ton, t. - 111 & 269. 18175 gtk in A. ‘De., Prodr. 5u6 & 555, 187; . W. Chapm., Fl. South. U. S., ed. 1, print. “i 18565 Ae and print. 2, 307. 1865; Griseb., Cat. Pl. Cub. 21h. 3 » Syn. , Chron Pi. 973 & 1220, 1879; S. Wats., Proc. Am, Acad. 18: 135. 1883; W. Mill., Dict. mg. Names Pl, 1h] & 261. 1884; A. W. Chapm., Fl South. U. S., e : Fls. ay 90, Rel & ety. “18314 cou Coult., Contrib. U. S. Nat. Herb. 2s Ui, Engl. & Prantl, Nat. Pflanzenf (3a): 148 : Dana, eat to Know Wild Fls., rev. ed., 122, 5, & 372. ep 330 PRY HL00.5.4 Vol. 11, no. 5 L. He Bailey in A. Gray Field d.Borogt & Gard. Bot. 31. 1895; Mill. & Whiting, Wild Fis. Northeast. States 488. 1895; Jacks. in Hook. f. e, Ind, Sih eh 78 & 11 1895; Hitche. & Nor- ton, Bull. Agric. Sta. 57: pl. 12, fig. 137 (1896) and 66: Ple 12 fig. 137 1897; Selby, Bul - 0 c. Sta. 8 rd ben: 330. 1897; A. W. Chapm., Fl. South. U. S 897; Britton & Br., Illustr. Fl., ed. 1, 3: 70, fig. 3058. cp Hitehe. & No - Kans. Agric. Sta. 76: pl. 9, fig. 48. 1898; A. S. Hitche., Fl. Kans. pl. 1h. 1899; J. K. » Southeast. U. S., 1 33 Selby & Hic Bull. Ohio . Ss ed. > 1009. 1 , Agric. Sta. 1\2: 118, 121, & 123, pl. 4. 1903; Michx., Travels West Alleg., ed, rete » 67. 190); Selby, Bull. Ohio Agric. Sta. 175: 350, pl. h. 1906; We Blankinehip in Engelm, & Gray, Pl. im. 3: 186. 1907; “icbingon & Fern. in A. Gray, New mp Bote, ed. 7, —. & 92h. 1908; M. ae tie Check List 128. 1908; B Mich, Weeds (Bull. Mich. Agric. Exp. Sta. 267:] 397. 1911; “gerth van Wijk, Dict. Retailing | - “1398, 1911; L. H. Bailey, Botany | 372. 1914; ony san. Rope. Mis me Agric. Exp. Sta. 25: 386. 19125 owa Geol. S - ks & 469. 1913; Britton & Br., Illustr. ri ed. Rs 3: ar fig. 3553. 1913; J. K. Small, Fl. Sout U. Se, ed. 2, 1009 1913; Shull, Plant World 17: 329--337. 191h5 Fl. V. N. + Mo. Gee d. 3: at 1516; Gerth van Wijk, Dict. Plantname 2: 228, 317, “& 161k. ; Farwell, Papers Mich. Acad. Sci. 3: 103. 192k; Goss, Journ, eg gi 29: 3h9--%2. 1924; Hegi, Illustr. s Rehmann, Am. Pl. for Am. Gard. 23, 96, & 130. 1929; Moldenke, List Spec. Mold. Southeast. Set 10.1930; E. J. Palmer, Journ. nold Arb, 11: 110. 1930; Merriman, Fl. Richmond 219. 1930; Trans. Kans. Acad. Sci, 3h: 261. 1931; Schwencke, Zytol. Unter- such. Verbenac, 10-12, 19, 23--2h, 33, 3k, & fig. h—-10 & 27-2 4321 Acad 1931; Stapf, Ind. Lond. 6: - 1931 Proc. Iowa ° Sci. 39: 122, 19325 Benner, Fl. Sacks a. 255. 19325 lg Fl. ‘lains *. & » Man pe ae. slg 39335 Moldenke, Obs. Fl. Watchung, fads Es e 203 1933 rry, ii 287, 20, "aha-278, & 356. 1933; Junell, symb. Bot. Upsal. h: Ul, 19, 171,'& 210, fig. 19. 193k; J. A. Harris, Physico-chem. Prope ; bd mss, 1935; E. Js Palmer * Mo. Bot. G 1935; W. C. Muenscher, Weeds 387.1935; H. T. Darlington, ih Mich. Acad. Sci. 22: 62. 1936; Fernald, Rhodora 36: pl. 45, fig. 1--h 936; gree uysele : —16h 7 1936; Cory, Texas Agr. Exp. S 550: 89. 19373 neo - Bull. 550: evist. Sudam. Bot. h: 18 & os "1937; Little, Am. Midl 385. 1938; Crocker, Bot. Rev. h: 256 & 26h. 1938; Fernald, 1965 Moldenke, Monograph of Verbena 331 Oz: 455, 1938; Moldenke, Alph. List Common Names 7, 9, & 33. 1939; Moldenke, Annot. & Classif. List 109. 1939; F. é. Holmes, 3 Phytopathogen, Virus. 180. 1939; F.C. Hoehne, Plant. & Subst. rs Med. 219-~250. 1939; Gates, Fl. Kans. 190 & 191, map 1072. 190; W. A. Weber, Summer age rare sy mSS. gto Mol- ana id Names 8 & 10. 191; Worsdell, Ind. Lond. Suppl. 2: 86. 191; Gates on Kans, 2 & 220, i9ha; C. N. Jones, Ohio Journ. Sci. kl: e 19413 Moldenke, Alph. List Invalid Names L5--4)7, 50, & “: ighss Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1); 1-- » 35, 7h, 101, & 102. 1942; F. A. Loew, Ann. Rep. Huntington pl We Bot. Gard. 73 STs 19h2; Oosting, An Midl. Nat. 28: 62, 6h, 7h, 82, & 109. 192; Schnack, anal. Inst. Fitotéc. Sta. Catalina exas 3 (1): 16 Be ep ol Se ~ 5 FS zs Bs Ae Bee ap me Kao = e Re -é 192; “- bor (a3) mar a bo "Tobe "hols, | Moldenke, Fl, N. Appal. Exp. rshed 28 & 56. 1943; Moldenke, Knom Geogr. Distrib. "Verbenac. Suppl. 1: 1. 19h3; Allard & Leonard, Castanea 8: 7. 193; Fs E. Egler, Bull. N. Y. State Coll. Forest . Techn. prod él: Ape 1943; rayton, perm. Ky. 116. 1943; C. N. Jones, Ohio Journ. Sci. eae 190. 19h Gier, Toc, Ind. Acad. Sci. ae ge 1944; Schnack & Covas, Darwiniana’ * denke, Phytol gs a 2: 70, "80, 88, & uu7. 1985 Moldenke, es fas 106: is8--161, 19453 G. Ne Jones, Fl. Il.. [Am. Midl. Nat. Monog. 2:] 215--216. 191,53 Moldenke, Torreya sis: "50 & 90. 19155 Moidenke, + Journ. Bot. 32: 609 & 610. 1915; , Kriebel, Yuncker, & Friesner, Proc, Ind. Acad. Sci. Bhs 95 5 (915) and 551 56. 1916; List. < * BE Moldenke, Alph. List Invalid Names Suppl. 1: 22, 2h, & 27. 1947 Woldenke, Phytologia 2: 326, 1 2T329, "aia & 362 (197) and 2: 478, 2948 Hausman, Begin. Gu & 375. 1948; Mol~ : List Fl, Obs. Beansbville 3. 1918; Leadeck: Castenen 13: 112 & 112, ine —— mr adaite l: 224. “r9l8; Ogden, Steinmetz, & Hyland) Bull. Joaselyn Bot. Gard. 8: 55. 1918; Gier, Trans. Kans, Acad. Sci. 50: T98. 198; Moldenke, Alph. List Cit. 2: 35367, 360, 361, 390-392, 394-02, LOh——LO7, 409, Lio, art 332 PHYTOLOGIA Vol. 11, no. 5 41k, 429, 438, 9, 451--456, 466, 468-70, 473, 475, 480, 181, 188; 489; Lo1--Lok, 97, 506, 507, S11, Sib—-S17, S19—-523, 525, 527=-530, 536, 538, Slb-— 519, 554, 556, 559, 560, 568, 569, 572, 576, 577, 519--581, 583, 58h, 587--589, 5 593, 59h, 596, eg 601, eit , eth 789-793, 800, 803, 811--813, 822, 824, 826, 827, 831, 839, 8 842, 8h, 89-852, 854, 856, 857, 866, 869, 870, 88h, 836, 887, 893, 899, 90h, 912, 916, 917, gst 927, 930--932, 939, a 9b, 946, 950, 953, 95h, 7705 & 978 (1949), and h: 979, 981, eS ah errr cob-=100l, 1010, 1013--1015, 1082, io, & 1299. 1949; Moldenke, Known own Geogr. Distrib. yet [ed. 2 2-21, 2h, 73, 165, 197, & 200. 1995 E. L. Palmer, Handb. Nat. Hist. 297 & 663. 191493 F. A. Loew, Ann. Rep. age dae Be Coll. Bot. Gard. 13: 37. 1949; R. T. Clausen, Cornell Uni Sta. Mem. 291: 42, 1949; Moldenke, Phytologia 3: 72, By ‘& ce (1949) and 3: 283 & 375. 1950; Moldenke, aorta Sudam. : 166. 19505 Fern. in A. Gray, Man. Bot., e ey 1209, 1210, & ‘1612, fig. 1463. 1950; Zim & Martin, Flowers Pr 19305 Me L. Grant, Proc, Iowa Acad, Sci. 57: 118. 1950 3 Deam, Yuncker, & Friesner, Proce Ind. Acad. Sci. 59: 51 (1950) ae 605 87. 19515 Moldenke in Chit~- enden, Roy. Ho - Soc. Dict. » 6: 2209 & 2213. 1951; isee oe Sp. Subsp. Cont. aa Set ro [3]. 19525 uoldenke, Phytologia 3: Lh9, 450, ‘isi, & 468 (1951) and h: 67 & 1952; iia List wild cul » Nat. Conf. 20. 1952; siete in Gleason, New 1955; R. Sy Pohl, Proc. Iowa Acad. Sei. 62: 182. 1955; Moldenke, Phytologia 5: 133. 1955; Moldenke, Inform. Mold. Set 51 Spec. b- 1956; Moldenke, Biol. Abstr. 30: 1093. 1956; Moldenke, Notes Pl. re hl. hi, 19575 Lehr, Bull. Torrey Bot. Club 84: 312. 19575 Alain & Alain, Fl. Cuba lj: 281. 1957; Strausbaugh & Core, Fl. r. 8 3: 77h & 775. 1958; Ahles, Bell, & Radford, Rhodora 60: 22. 1958; Moldenke, Am. Midl. Nat. 59: 3h6--348, 352, 353, 35% 357-360, & 362. 1958; R. J. Eaton, Rhodora 60: 68. 19583 Van Vechten & Buell, Bull. Torrey Bot. Club 86: 225 & Lbs « * wee Moldenke On, New Britton & Br. Illust oe - 126, 129, & 130. 1958; Moldenke, lyn12, W123, 25» 30, 85, 2 ‘ore 19593; Moldenke, Résumé Suppl. 1: 1, 2, & 23 (2959) and 2: 1-3, ll, & 13. 1960; T. B. Mitchell, ma, East. U. S. 1 [N- Ce» Agrs Exp. Sta. Tech. Bull. 1h1]: 292. 1960; H. Aas Kemp, Bull. T 1965 Moldenke, Monograph of Verbena 333 Bot. Club 88: 60. 1961; K. L. Brooks, Bull. teem 5 Bot. Club sn 3 420. 1961; Lewis & Oliver, Am. Journ, Bot. 8: 639-6 ig. 2. 1961; Moldenke, Phytolo 3 8: 120, 1A, onus & 143 (ag6i) and 8: 125, 281, 296, 356-- 3, "32, & 397 (19635, 30 100, 10-106, 109, 140, 18, 28; Con 178, 182, 210, 212, 217, 218, & 307—311 (196), ‘ Sei. a Hist. Bot. fig. 77. 1675; R. Morison, Pl. Hist. ann Ox- Sp. pl. 1146, 1714; Dodart, Mem. Serv. l'Hist. Pl., ed 5. eet eee ee Herb. ee 2 pl. 1158 [in color], Tray ities in Martelli, - Roman. 3: 55 [in color]. 1775; Maund, Bot. - 5: pl. nn fin color). 1831-855; oop & Waiting, y wha Fls. Northeast. States 88. 1895; Hitche. Norton, Agric. Sta. 57: pl. 12, fig. 137 (1896), 6: gat 13, fig. 37 (2897), and 76: ni, 9, fi rig. : . 1898; Britton & Br , Tiinste n-, Ld ’ benac. fig. 5) i —30. 1931; Junell, Symb. Bot. Upsal. \: fig. 19. 193k; v ri ‘aunitear: Seo 387. 19353 tard . Fernald, Rhod : 163, fig. 17. 1936; Hausman, Begin. Guide Wild. Fls. = Pra eee fy Nat. Hist. 297. 1949; Fern. in . ne wd oe et %, 7 af ° . 0 artin Flowers » fig. 1463. 1950; Zim & M a eeteeer, Fis; weit. 2, 32 334 PHYTOLOGIA Vol. 11, no. 5 most seen » nearly all oe upper nodes producing perfect or sos 7 cae decussate-opposite, up- igi = s ender; leaves gaa gig pS perce jan peathtee Saale short, Seay cae ee. narrowly nargined, rough-hirsute; leaf- — very t thin-membranous, dark-green both surf. rom ovate or broadly ovate to Scelk ee long-lanceolate, cps lanceolate, oblong-ovate, or lance-ovate, long, acute or short-acuminate at the apex, mostly rounded at the base and decurrent into the petiole, coarsely ser rate-dentate or serrate to somewhat doubly crenate-serrate with Ss, sim slender or filiform, green, interrupted, rather flaccid anthesis, to 15 cm. "Long or longer, aan sparsely flowered, us stif frui very short, ciliate-margined; flowers oo or minute, about 4 mm. long, seP~ arate or subopposite on 2s rachis, sessile; calyx tubular, 2— 2.3 mm. long when mature, green, 5-ribbed, strigose or _strlgose- le to minutely pubaonent. onereny along the veins), its lobes a or reieaee ae con= meeting at right angles; chromosome number: n * "t, én * herbarium from Linnaeus' own herbarium “a not of the type collection, since they seem to be fram a materi zn cultivated in Uppsala and are probably the basis for his des¢ a tion in Hort. Ups. (1748) —- the Nietzel specimen is inscribed 1965 Moldenke, Monograph of Verbena 335 the obverse ss ae in pencil, “cfr. Verbena 11 bh gb 2 — sp. pl. 20", and on the reverse side, in eat " and, in ink, "in Montin's handwriting, "Verbena Cavétoifolas) tetrandra, spicis filiformibus paniculatis, foliis indivisis ovatis ser: rratis. acutis petiolatis. Sp. pl. ed. 2. ne 13. Habitat in Virginiae, dis. Specimen ex horto Upsal. communicavit hortul. Nietzel." The first Linnaeus specimen is inscribed on the ob- verse side, in pencil, "Verb. 11. urticifolia L. sp. pl. 20" and, in ink, in'L us! ing, "11 urticifolia", and on the reverse side, in pencil, "Linné herb." and, us! omm handwri » "ex horto ups." hand "von Linné sen, tanh and then, in ink, "Herb. Alstroe- merii" and in Dahl's hand " f£." The second Linnaeus se Se is inscribed on the obverse side, in ink, "VERBENA" and on the bottam of the sheet in Linnaeus’ hand "urticifolia", and on the reverse side, in pencil, "Linn herb." and, in ink, "Verbena tetrandria, spicis filiformibus pan- iculatis, foliis age serratis. Lin. Spec. plant. 20, 11." Linnaeus? © 1.9 oh." Walpers (1815) pl places the seiatas in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, hae, and Sec- ondary Subgroup Holophyllae, with 22 other species, keeping V. diffusa Lam. as a distinct species. Hooker (1836), under V. hastata, says "From this the V. panic- ulata is probably not distinct, oad the V. urticaefolia seems too allied.” The first half of this statement, of course, is quite true, but the second half has no basis in fact at all. The description of V. diffusa in Rees, weg (1817) is as fol- lows: "Spreading Vervain, Poiret in Lam. no. lk, Pursh a. 7. _ sp ranc Leaves ovato-lanceolate, serrated, saewhat domy. — Native of North America; cultivated at Paris, according to Poiret, on whose authority Mr. Pursh gave this species a place in his Flora. He speaks of it as having a near resemblance to V. urtiei folie, at ag small, somewhat purple, flowers. We have seen no tt en Poiret (1808) describes V. diffusa as follows: "Spicis longis- Simis, laxis, paniculatis, valdé diffusis; foliis ovato~lanceola- tis, subpubescentibus, serratis; caule Frakescente, subglabro" and then he follows this with a variety: "Pf. i: ca to; = ead din Sept, ar pipiens ye ore variété 336 PHYTOLOGIA Vol. 11, no. 5 carolina L. or V. scabra Vahl. The former of these, however, was known to Poiret [as "10. V. caroliniana L. — Lam. — Willd. — Michx. — Dill. "Je so perhaps it was the See to which he is here referring. The phrase "somewhat purple" flowers leads me to suspect that V. brent itself does not belong in the synonymy of ve urticifolia, but bel: belongs with xV. engelmannii Moldenke. The actual specimen in ya Desfontaine H Herbarium would have to be consulted in order to determine this matter definitely. W. S. Dana (1895), in referring to V. urticifolia, says: "It almost excites one's incredulity to be told that this baancabig es esting-looking plant, which grows rankly along the highways, i portation from the tropics, yet for this statement the are eous s 8. e spelling oP ise specific part of the name of this plant has varied between "urticifolia" and eae in both ean and post-linnean work. How Linnaeus Repo: it as "urticifolia" in his eten it "01753), which is t pig nck aR ae Be ppuerclatcrs, and I see no justssteation for chang so many other writers have done Strausbaugh Pe sate (1958) Naetiaay spell it "ae" and "i" on page 774 of their work and "i" on page 775. The type of V. hastata var. oblongifolia was collected by Timothy Langdon Andrews (no. 146) in Montgomery County, Tennessee, and is deposited in the herbarium of Ipwa State College at Ames, while that of ¥. urticaefolia var. hirsutior was gathere George Engelmann at Saint Louis, Missouri, in September, 1836, and is deposited in the Jesup Herbarium of Dartmouth College at Han- over, New Hampshire. The V. urticifolia var. riparia of Britton is regar arded by me as a synonym e V. riparia Raf. This latter species, however, is reduced to synonymy under V. urticifolia by Taylor (sis), a disposition aa which it is impossible eds me to V. ree. Schauer (1847) reduces V. scabra Vahl to synonymy ze urticifolia, “a with a ee He cites the original L., SP» Pl. reference to e@ "29 : Barrelier (1714) cites saudi 6 chian Hernan. 399" in the syn~- onymy of our plant, but the "QVAVH CHIAN. Caulis coloris castane+ Sherer. fructus vt in Absinthio coloris castanei" of Hernande?, urticae folto estaba. foliis oe flore majore Tourn. is given by me in my Résumé Suppl. hz: 20 (1962) as a synonym of V sige urticifolia, but it actually belongs in the synonymy of V- officin sl. ~~Several hybrids involving V. urticifolia are known: that oe V. bracteata Lag. & Rodr. is xv. perriana Moldenke, with V. haste a L. is xV¥. engelmannii Moldenke, 1 with V. hispida Ruiz & Pav. is x¥s 1965 Moldenke, Monograph of Verbena 337 fabricata Moldenke, with ¥. lasiostachys Link is xV, inhonesta Moldenke, with V. neomexicana (A. Gray) Small is xv. torpa torpa Molden- ke, and ath vs stricta Vent. is xV. illicita Moldenke. The v. urticifolia x stricta of Eggert, of Norton, and of Gates (in part) are all xV. illicita, V. urticifolia var. paniculata Engelm. is ee . Verbena urticifolia has been cultivated in European gardens for several centuries. Linnaeus found it in the Clifford garden in in or before 1737. Hegi (1927) says of it "die bereits zur Zeit der Renaissance nach Schube in den schlesischen G&rten gepflegt wurde", He says that it is used medicinally "gegen Aus~ schlag", Hoehne (1939) + aa its use as a lotion against poison -- (Toxicodendron radicans) poisoning, while C. G. as AP eye "reg — Poindexter (1962), in his comparison . = urticifolia with V. Stricta, V. hastata, xV. engelmannii, and xV. illicita, describes Ve urti urticifolia as having a leaf-index of ese (average 2.2), leaf-shape broadly lanceolate to ovate, leaf-base rounded and de~ current into the petiole, leaf-apex gradually acuminate, stem pub- scence hirtellous, nutlet length 1.6--1.9 (average 1.7), markings S the back of the nutlets moderately ribbed, pollen fer rtility 63 99 percent (average 9.2 percent), corolla-tube 1 1.5--2.5 ee 2.1), calyx length 1.7—~2.3 (average 2), and eae - in measurement 1),—25 (average 19.6). Poindexter 9 had imen of VY. urticifolia that I have ever seen growing is cited below as A. R. Moldenke 65, from Botetourt County, Virginia — it was 8 feet 3 inches tall by actual measureme Demaree 20121 & 211,80 represent a very hairy form of the spe- Cles; F.C. Gates 20889 is also very hairy and was identified by the collector as "V, urticifolia x stricta", but I do not believe that it is a hybrid. H. N. Moldenke 1330 has the pubescence re~ markably short. E. M. Kittredge son. (Aug. 23, 1921] is very ab- normal, probably due to virus infection — it was examined in ton, Aug. 1h, 1593) has the nisieeter rennet pore needing infection and calle to mind Stachytarpheta jamaicensis f. monstirosa 338 PHYTOLOGIA Vol. 11, no. 5 (Moldenke) Moldenke. Van Dersal 303a is also anomalous and re- sembles xV. engelmannii, J. M. Fogg 11475 has its fruits very densely crowded on the spikes and may possibly be xV. engelmannii. Whitehouse 16426 is very close to V. carolina L. in appearance. E. E. Chambers 2 27 has inflorescences exactly like those of typical var. lelocarpa Perry & Fernald; C. J. Elmore son, [Talmage, July 6, 1891] may possibly be that variety. Je Torrey 77-0 in the nevberdien of the New York State Museum at Albany is inscribed "hy- brid between V. urticifolia & V. hastata", but certainly is not 4 hybrid of ~~ — Similarly, | R. Bebb 2107 is inscribed "probab- ly a hybrid f » but I do not think so. I regard both he specimens as eu esenting typical V. urticifolia. Wills {July 27, 1959] was identified as xV. illicita by the Beare but seems to me to be typical V. urticifolia. S. Watson (1883) states that Palmer 2037 is "perhaps a hybrid between V. poly- stachya or V. urticaefolia and V. tha", but I regard ad it as typ- ical V. ehrenbergiana Schau. __ Shinners 179 is said to have had "fls. pale bluish-white" and Harris s.n. [June 26, 1933] is inscribed "fls. pink". I+ is pos- sible that one or both of these represent var. incarnata (Raf.) Moldenke, which see. Hausman (198) describes the flowers of v- urticifolia as white or pale-blue. I eae that the pale-blue and pale bluish-white flowers were seen on specimens of xV. en- gelmannii, perhaps growing in close be ocosatins with V. u urtici- folia, as it often does, Shull (191)) Sapo br seeds of V. urticifolia after they were buried in mud covered with several inches of water for hl years. Goss (92) reports that he buried seeds of this species for twenty years -- 8 inches deep, gave 82.5 percent germina nation; 22 inches pe gave Res percent germination; 2 inches deep, gave 78 percent germinati The species has ssi found growing in shade as well as in di- ands, or 1 et bogey swamps, “ low land in general. Collectors report it from or wee eadows, low meadows and meadows in general, f PY 1965 Moldenke, Monograph of Verbena 339 dwellings, in woods, low rich or alluvial woods, open o damp woods, low or low moist woo ods, low woodlands bo ordering marshes, lowland woods along small streams, "Low brushy or bottom woods, up- woodlands and wooded spring creek valleys, bre or Bien ravines, shaded ravines and shaded ground under ecan tre swamp an floodplain forests, pine forests, cleared low vine rass areas, sandy low areas sax ricberctebars marshes, swampy places and open bogs a- long streams, in sandy or rich moist thickets, roadside thickets and thickets at ah edge of prairies, swamps, ’ swales, valleys, and arnyards. It grows on sandbars and sand flats, stream and creek banks, river and railroad banks » ary or steep banks, moist or al- as oms, on hillsides and shady steep hillsides , ary wooded hillslopes, high bottoms and pastured slopes, on low or loess ridges, mud and — flats, and prairies. It thrives along roadsides, dry or ry : lakes, pine~oak and rocky woodland borders, at the sides of rivers, at lake margins, and s open cedar Swamp margins. It January and from May to October, fruiting from May to November Carpenter describes the species as "common, roadsides at lower altitudes" ay Vermont. J. A. Harris (1934) says that it grows "in shady w and bogs and along edge of lakes" in wit! York, while Egler (a9) reports it from "forest edges, in silt loam with limestone rock in full to partial light; nda: single to several where present" in the same state. a. ennsylvania a Ben- ner (1932) found it in "rather dry soil of waste places, roadsides and fields; common", while Stone (195) ro “nroadsides, gardens, Waste places; common; fl, late June—-ear July". Merriman (1930) "found in open fields and pastures usually in poor sandy soil" in West. Virginia. Oosting (1942) avers that it occurs in oo eco= Logical communities -- birch dominance, mixed hardwoo ary and postclimax forest in the Piedmont Pf North Carolina, In entucky it is described as "a common coarse weed" by iain a and to grow aoe agen a , waste places, and laces in woods" by E un (1943). G. N. jones says (19h5) that it grows along Syoicuaats and open woods, common" » 3m. (Middletown, 18,0] (S). New Haven Co.: Kleeberger s-n. sen. [New Haven, Aug. 187k] (Gg—~31h39). New London Co.: Jansson $.M. (July 26, 1927] (S), sen. (Groton, Aug. 16, 1928] T (or--22020), 8+ n, (Groton, Sept, 2, 1929] (Go), s.n. [North Stonington, Aug. 12, 1952] (Go); W. A» Setchell s.n. (Norwich, July 28, 1883] (Ca-- 25190). Tolland Co.: M.T M. T. Travis 1918 (Up). Windham Co.: J» Le Sheldon s.n. {Central Village, July 1897] (We). County undeter- mined: A. Or Gray s.n. [Conn.] (S); Krebs s.n. (Cp); C. Wright 8-Re (Cm). NEW YORK: Al Albany Co.: H. D. House 26892 (Al); Whitney “7— igh (Al). Allegany Co.: A. R. Moldenke 1053 (Iw), 1055 (Lw)- Bronk Co.: Ahles 581 (Aa), 653 (Aa); Clute s.n Son. [Pelham am Park, July 12, 1899] (N)j H. M. Denslow s.n, [High Bridge, July 1864] (Ms); 2+ ¥ Edmondson Big ( (N); Gill: Gilly 110 (N). Cattaraugus Co.: Alexander & House 13023 (Al); A. R. olsenke 1052 (Lw). Chautauqua Co-t Fe We Johnson 1329 (N). Chenango Go.: EB. E. Davis 83 (al). columbia Cow: R. McVaugh 1408 (al, a 14767 (ML). Deleware Co.: P. Wil- 1965 Moldenke, Monograph of Verbena 349 son s.n. [Arkville, July 26, 1915] (N). Dutchess Co.: J. M. Fogg 7L07 (Up); Moldenke & Woldenke 19528 (Sm); Sherwood s.n. (Pawling, Aug. 5] (Sh, Sh, Sh, Sh, Sh, Sh, Sh), 5.n. [Pawling, Aug. 10] (sh, Sh, Sh), s.n. (Sh). Genesee Co.: E. Je re Hild 134.1855 (Ur); Muen- scher & Brown 210) (It); W. H. Rhoades s.n. [Buttermilk Falls, Aug. Aug. 8, 1915] (Hs). Greene Co.: F. We Johnson s.n. (Windom, June 25, 1921} (N); N. Taylor 129) (N). Jefferson Co.: Hotchkiss 873 (Al). Madison Co.: H. D. House 13648 (al ); Maxon s.n. wn. [vicinity of Oneida, July 22, 1895) (W-—674568). Monroe Co.: Barss s.n,. (Rochester, 7 September 1909] (Or--8873); Baxter s.n. [near Roch- ester, August 1925] (Ca--88283k);5 Holger s.n. [July 1863] (Al); We A. Matthews 3272 (Hi--29811, La); A. Re , Moldenke 9 {H. N. Mol- denke 20437] (N, No). Nassau Co.: He Ne ~Moldenke 19050 (N, Or); W. A. Weber 933 (Bl—)8170, I Io~-150189) . Niagara Coe: | E. C. Town- Son. [Aug. 1890] (P1—86886). Oneida Co.: Haberer s.n. sae 21-1886] (a1). Onondaga Co.: G. T. Hastings s.n. [Tully Lak July 31, 195] (N). Ontario Co.: A. R. Moldenke 7 [H. N. paidane 20435] (Mr, Mr). Orange Co.: Bohus-Jensen & Moldenke 1,1 (Jn); zs M. Denslow s.n. [Aug. 10, 1922] (N); W. He Lewis Jr. s.n. [Tuxe Park, Jy. 27, '99] (N); Lucian 406 (W1); Raup 7603 (vi). ae Co.: Lennon sen. [July 1893) ( (Al). Oswego Co.: : Hotchkiss 3314 (Al); ¢ C.S. Sheldon s.n,. (July 13, 1878] (Al). Putnam Co.: : Mol- denke & Moldenke 11852 (N). Queens Co.: Hexamer & Maier s.n. e [Lit- tle B. Bayside, July 9, '53) (Cm); L. Johnson s.n. [Fresh Pond, July 18, 1876] (Lh); Poggenburg s.n. [Flushing, July 11, '86] (Tc Danenetner Co.: Banker 138 (N) (N); Puissant s.n. (Br). Rockland Co.: Lehr 435 (N). Saint Lawrence Co.: lirs. 0. P. Phelps 800 (Gg— ). Saratoga Co.: J.M. Fogg 7132 (Up). Schenectady Co.: He D. House 23747 (Al). Steuben Co.: A. R. Moldenke 12 [H. N. Molden- ke 20163] (N). Suffolk Co.: Banker 3020 (N)j Je Ae . Harris C.15997, in part (Ca—-6101),8); L. K. ene ‘S.n, {Wading River, July-Aug. 1928] (cm). Tompkins Co.: L. H. Bailey s.n. [near Ithaca, Aug. 25, 1925] (Ba, Ba); W. C. Baker s.n. [Ithaca, 1905] (Ba); Clovis s.n. [September 1952] (We); Greenfield s.n. (July 1931] (N); Hoisington 74 (Ok—-15639); Laird s.n. [July 17, 1917] (Ms); B. Maguire 6661 (Wa—26813); A. R. Moldenke 4 [H. N. Moldenke 2027] (N), 1069 (Lm); Muenscher & Bechtel 526 (Ca--882835, Io--112239, P1--63399, , Ur); 2 616 (Ca--882642, Po-6682); Naylor & Smith s.n, (Ithaca, 3-5- Waly (5 (Bt—3767h); E. L. Palmer 1013 (S); 0. E. Pearce sen. [Six @ Creek, July 26, 1885] (W--210075); C. H. Peck s.n. . [Freeville] an), "J. M. R." gn. [July 26, 1926] (Gu--B-1055); Rendle s.n. [Mc- Clean, Aug. 16, 1926] (Bm); Cc. " skottsberg sen. [17/8/1926] (Go)- Uster Co.s Elting 2074 (Al); Je M. Fogg 7123 123 (Up); H. D. House 350 PHYTOLOGIA Vol. 11, no. 5 2492 (Al). Warren Co.: H. D. House 30092 (Au). Washington Co.? S. H. Burnham s.n. [August 1912] (Al); Dobbin 23) (Al); A. Fitch 3.n. 1. {Bast Greenwich, 1865] (Mi); H. D. House 2387h (Al), , 28329 Gl, Ca--8828)0), Westchester Co.: | Je He (. Barnhart hart 333 (N), > 18h i eienellllliemeneel aecummnmneneenmeerenaniiiceremmme ae ye Se a ee 2 (H. N. Moldenke 21120] (B, Fy, N, Rs); Moldenke & Acufia 19503 (Es); Waxenbaum 1281 (Wx). oes Co.: Clovis s.n. [September 28, 1952] (We); S. H. Wright s.n. (Mm--15),02). Staten Island [Rich- mond Co.]: Billberg s.n. [Staten Island, 1826] (5S); H. N- Molden- ke 19122 (N, Or, Si); W. A. Weber 207 (s1—-18172, To=—150808) « County ns ears Collector undesignated 1525 (N); Herb. Jard. Bot. Brux. s.n. (Long Island, July 24, '67] (Br)3 ovtley 16595 (Kr); J.T Je Se 747-0 (Al), sen. (C, Pa). NEW JERSEY: Bergen Co+! Dautun s sen. (Cresskill, Aug. 13, 1903] (La), sn. aase Hill, 1 aout 1908] (La); Oehler s.n. [1877] (N). Burlington Co.t Je Me Fogg 9240 (Up). Cape May Co.: W. Stone 16316 (Up). Cumberland Coes iW W. Adams 883 (Up, We). “Essex Co.: Lighthipe s.n. [East Orange, Aug. h, 1914] (Au), s.n. [East Orange, Aug. 7, , 1917) (N)- Hudson Co.: L. H. Bailey s.n. [Hoboken, Aug. 18, 1920] (Ba)5 ae 8.n. 1. [Hoboken, - 1826] (S). Hunterdon iio. A. R; Moldenke Lé (H. N N. Moldenke 21769] (Bm). Mercer Co.: Collector undesigna- ted son. (Aug. 2, 1876] (Pr), sen. (Pr). Middlesex Co.: B. D- Halsted 175 (Ah, C, Fe, Io--1535), Mi, Ua--11393, U Va--27136, Ur, Vt, We); L. H. Pammel s.n. [Perth Amboy, Sept. 3, '24] (Io— 116602) Vail s.n. (New Brunswick, July 18th 1890] (N)- Monmouth Co.: Re C. Alexander s.n. (Redbank, 1 July '6h] (Ca--379999)5 He N. Moldenke 21718 (Mm, Sm). Morris Co.: N. L. Britton s.n. [Green Pond, Sept. 21, 1886] (C). Passaic Co.: B. W. Berry s.n. (Passaic, July "189h) (Te); G. V. Nash 476 (Br, Lh). ~ Salem Cow: Je “J. Me Foss 10011 (Up). Somerset Co.: Lavoie 403 (Vi); Lighthipe s.n- ‘san. [Rocky Hill, Aug. 1, 1916] (Au, N)s A. R. , Moldenke 18 [H. N. Moldenke 20387) (N, Wh); H. N. Moldenke 1330 (E, Go, H--5483, N, N, Ny Ss Up, Ur), 2627 (N), 7u0l (Pl. Vic. N. Y. City Ll] (Go, N, tm—153, Ur), 8092 (Go, N, N, S, Ur); Moldenke & Moldenke 11687 (Go, N)- Sussex Co.: H. N. Moldenke 21539 (Ss); H. H. Rusby sn. (Franklin, Aug. 1879] (Mm--16103). Union Co.: H. N. Moldenke 21885 (B)- PENNSYLVANIA Adams Co.: J. M. Fogg 13598 (Up), 15216 (Up)- Alle gheny Co.: A. Dubois 337 (N), 351 (N)3 0. E. Jennings son. [AUS 1, 1918] (Cm); Millward s.n. (Wildwood, July 29, 1919] (Cm); Van Dersal 303a (Cm, Up). Armstrong Co.: J. M. Fogg 15338 (Up)+ Beaver Co.: Andriessen s.n. [July 1881] (Cm); Graham & Graham S.n- (7/29/ 23] (Cm). Bedford Co.: 0. E. Jennings s.n. (8/10/19h1] (Cn) . Berks 1965 Moldenke, Monograph of Verbena 351 Cows J. M. Fogg 5826 (Up); H. N. Moldenke 19371 (N). Bradford Co.: H. A. Wahl 16802 (Hi—168732); Westerfeld 919 (Cm). Bucks COs? He Re Bassler s.n. [July 27, '79] (Ka Butler Cot Bright s.n. {Plains Church, Aug. 2, 192k) (Cm); H. ¥. Graham s.n, ~ {Buhl, 8/28/22) (Cm); L. K. Henry 552 (Ca--882839, Hi--192854, W— 0936); 0. E. Ji Jennings s.n. [Valencia, Sept. 27, 190k] Sg XN. Bos H.3322_ (NR-1986] (We). Cambria Co.: 0. E. Jennings (Cn), s 8.n. [Flinton, ape els Cameron oe x M, Fogg Fen (Up); O.£ E. Jennings s.n. (Emporium, July 2, (Cm). Centre Co.: 0. £ E. Jennings | S.n. a 1/16/09] (om); A. R. aks 787 (Lw). Chester Co.: J. M. Fogg ‘oge 5772 (Up). Clarion Co.: 0. E. ? 0. B. Jennings 8.n. [Red Bank, July 21, 190] (Cm); H. N. Moldenke 21899 (Lw). Clearfield Co.: Westerfeld 23 (Cm). Columbia Co.: H. N. Moldenke 21892 (Lw, Ut). Crawford Co.: C. D. Curtis s.n. (Meadville, Aug. 193) (Po--65182); Shafer s.n. (July 23, 1901] (Cm, Cm). Cumber- land Co.: A. R. Moldenke [5 [H. N. Moldenke 21768) (Bm), 98 (Fg). Delaware Co,: “Brinton S.n. . (Wawa, Aug. 29, 1888] (Up-~17127); E. (Up). Erie Co.: N. H. Phillips 66 (Up). Franklin Co.: A. R. Mol- denke 42 [H. N. Moldenke 21765] (Bm), 43 (H. N. Moldenke 21766 (Ba, Gg, Mi); 3 Tidestrom 733) (W--7505]). Greene Co.: 0. E. Jen- nings s.n. [9/9/21] (Cm); Lohr 236 (Up). Huntingdon Co.: B. He Graham s.n son. [7/12/32] (Cm, Up, Up). Indiana Co.: 0. E. Jennings 8.n, 8.n, [July 11, 1908} precy Jefferson Co.: H. N. Moldenke e 21898 Gis). suntate Co.: J: M. Fogg 15537 (Up). Lancaster Co.: Bru- baker 59 (Up), 88 (Up), 956 (ip), 92 923 (Up), 986 (Up), 1496 (Up); A. A. A. Heller s.n. (Lancaster , August 29, 1900) (W--l07056); J. Ke Small 106 (0b~-50853). Lawrence Co.: Jennings & Kennedy s.n. {Sept. 18 18, 1919] (Cm). Lebanon Co.: C. He ~ H. Kauffman son. (Aug. 1889] (Mi). Lehigh Co.: Arndt s.n, [Allentom, July 1911] (Up); Q'Neill s.n. [Cedar Creek, Aug ins st 19, 1927) (Fl--21130, I). Lu- zerne Co.: J J. K. Small s.n. [Lily Lake] (Cp). lye Coes Gress, Jennings, & Jennings s.n. [Cedar Run, Sept. 2, 1920] (Cm); W. H. H. Rhoades s.n. [near Williamsport, Aug. 1937) (N); W. R. Tay~ lors 8.n. [Essick Heights, 1912] (Up~68767) « Monroe Co.: Bern~ hardt s.n. (Paradise Falls, Sept. 10, 1928] (Up); G. B. Grant Pe (Po—267626)5, Langnan 94 (Up). Montgomery Co.: J. M. Fogg 2140 (Up); A. R. Moldenke 103 (Fg); V. Wismer LOS (Up), L2h (N, ty. Northampton Co.? Re ze : L. Schaeffer 9450 (Au--122815); Tyler S.n. (Easton, Aug. 7, 1 ) (N). Northumberland Co.: J. M. Fogg 17687 (up), 18013 (N); “d se Moldenke 13 [H. N. Moldenke 2047) (N); M. T. Travis 2296 (Up). Perry “Perry Co. J. M. Fogg 17922 (Up); as Be lene 16 TH. N. Moldenke 20437] (N)- Philadelphia Co.: J. W. Adams U6 (up); Brinton s.n. [8.91878] (Pl—-115520); J. M. 352 PHYTOLOGIA Vol. 11, no. 5 Fogg 16194 a (Up), 162hh (Up); J. M. Greenman 137) (Or--14318)5 He A. Lang 51 1h (Up); Mack MacElwee 882 (Ka--71128, | 18; S, Um-~197), Some [West Philadelphia, Sept. h, 1895] (N). Pike Co.: Graham & = sn. (July 1930] (Cm). Schuylkill Co.: J. M. Fogg 1 11475 (Up); Ps R. Wagner 686 (Up). Snyder Co.: A. Re Moldenke 15 TH. N. Moldenke 20485] (S); Wade & Wade 143h (Up). Somerset Co.: = Je M. Fogg 15266 (Up), 15280 (Up); 0. E. Jennings s.n. {Keystone, oct. 9, 190] + ivan Co.: F. R. Fosberg 15778 (Up); W. Stone 15168 (Up)- Warren Co.: H. a Moldenke 15426 (We), 17983 (Br). “Washington Co.: st M. Fogg 18354 (Up); L. K. Henry s.n. [9726/40] (Cm); D. Wilson 52 (We). Westmoreland Co.: Denoise s.n. [9/1/35] (Cm), Sn. [7/ = (Cm); L. K. Henry s.n. (7/28/41) (cm); K. R. Holmes 8.Me Bet, July 15, 1902] (Gm); A. R. Moldenke 102 (Fe) H. Ne Mol- 19366 (N). Wyoming Co.: Osterhout s.n. “TTunkhannock, - July = 139i (Po--17326) « York Co.: J. M. Fogg 12643 (Up); Ke Be Hoover 4034 (Hi--196036). County undetermined: Schweinitz s.Me [1829] (Br). DELAWARE: New Castle Co.: Canby sen. (Sept. 28, 3, 1896] (Pl—22611). Sussex Co.: Lafferty s.n. [July 1865] (Pr). MARY- : Allegany Co.: Ahles & Gilpin 8182 (Ur). Baltimore Co.s Boldo 62 (Q)3 Jones & Ri Sen. [AUg. 15, 1903] (Mn—6891). Calvert Co.s Tidestrom 7797 (1). Carroll Co.: M. J. Fisher 121 (Up)j A+ R. Moldenke 495 (Fg). Dorchester Co.: c; P. Smith Smith 2936 3% (I). How- ard Co.: Arsene 72h (Or--14678); A. Re Moldenke ke Lok (Fg)- Kent Co.s He Ne Moldenke 13869 (z). Mewieoamey. 0 +8 Eee Ewan 17144 (Tl, Tl). Prince Georges Co.: Hyacinth 16h (Or—-1h730)3 A- Ah. R. Molden- ke 193 (Fg); Petrak s.n, [6.VIII.1950] (S); Tidestrom 1462 (1); True 564 (Up). Talbot Co.: E. C. Earle 1734 (Up). Washington Co.: A. Re Moldenke 39 [H. N. Moldenke 21760] im Worcester Co.: G. Ee eeldeake S.n. [H. N. Moldenke 6638] (N). DISTRICT OF COL *, Blanchard s.n. [Sept. 20, 1890] (Dt), sm. {near Long Bridge, 1 ig he 1891] (N); Boettcher 226 (Ca—-162h75)3 Je He = Hol- mes s-n. [1888] (Ka); leranier s-n. (Was [Washington, Juli 27, 9331 (8)3 We | H. Seaman s.n. (Mm--15)0]); BE. S. Steele s.n. [guly ear (Ob--50850) . = foes Albemarle Co.: H. N. Moldenke 19225 , Si). Augusta Co.: Murrill s.n. [12 July 1698] (N). Bedford te Curtiss s.n. [August 3, 1872] (S). Botetourt Co.: A. Re Moldenke 65 [H. N. Moldenke 21790] (Bm, Gg). Craig Co.: Je Me Fogg 13149 131k9 (Up). Fairfax Co.: J J. J. Carter sn, [Falls Church, July 1876] (11); A. R. Moldenke 497 (Fg). Fauquier Cos A+ Re Moldenke 66 [H. N. Moldenke 21791] (Bm). Giles oe Jeu [. Fogg 12772 (Up), 14787 (Up). Gloucester Co.: A. Re Mol ene 188 (Fe)- James City Co.: Arts 1161 (We, We). King Willian ee sage -. Re Mol- denke 487 (Fg). Loudoun Co.: A. R. Moldenke 67 [H. N- yoldenke 21792) (Bm, Gg). Montgomery Co.: R. Kral 11160 ta n15372) Nor- 1965 Moldenke, Monograph of Verbena 353 folk Co.: Ae R. Moldenke 47h (Fg, S). Northumberland Co.: A. R. aopconke 486 (Fg). Orange Co.: Nason s.n. [Gordo onsville, July 1867) (U (Ur). Prince William Co.: A. R. Moldenke 496 (Fg). Pulas~ s Co.: A. Re Moldenke 61 [H. N. Moldenke ke 21786] ~(Bm). Roanoke Cos: A. R. Moldenke denke 63 [H. N. Moldenke 21788] (Bm); C. E. Wood 328 (Ca--77281L), 6248 (Hi--51276). Rockbridge Co.: A. R. Mol- denke 6, (H. N. Moldenke 21789] (Bm). Rockingham Co.: H. Ne Mol- denke 19226 (N). Smyth Co.: A. R. Moldenke 59 [H. N. Moldenke _ 21784) (Bm); J. K. Small s.n. (Marion, July 20, 1892] (Ba, Ob-- 50846, We). Washington Co.: A. Re Moldenke 57. {H. N. Moldenke 21782] (Bm, Gg). Alexandria: A Ae Re Moldenke 498 (Fg). County un- determined: Herb. Mus. Bot. Stockholm s.n. (S);_ Herb. Retzius s.n. Lactealenaiae eapeaien oceania eaemme ieee ammee (We). Brooke Co.: E. A. Bartholomew B.1937—11 (We). Calhoun Co.: Re Harris 106 (We), s sen. (June 26, 1933] (We). Gilmer Co.: G. B. Rossbach s.n, s.n. [10/12/52] (We). Greenbrier 0931 E. E. Smith s.n. (August 3, 19h7] (We). Harrison Co.: Judy s.n. [summer 193k] (We). Jackson Co.: Neat VasBet Exped. s.n. [June 25, 1930] (Bt- (Bm). Kanawha Co.: Greenlee s.n. [Aug. 12, 1934] (We); ¥: o: Randolph 227 (We). Lewis Co.: Mrs. V. Joana s.n. [July 18, 195h] We). Marion Co.: Herb. West Va. Univ. s.n. (We); A. mace sun. [Valley Falls, June 27, 1957] (We). Marshall Co.: E. A. Bartholo- mew M.285 (We). Mineral Co.: M. Brown s.n. [August 23, '50] (We). Monongalia Co.: Anderson & Smith 360 (We); Sheldon s.n. {State Farn, 7/28/14] (We), son. [State Farm] (We). Ohio Co.: B. Drum mond 8.n. [October 1937] (Cm); Strausbaugh 3h) (We). Pendleton Cou: Re B B. Clarkson 1395 (We); E. L. Core 3188 sae aaree 6131 (We) 5 Ni Netting 117 (Cm). Preston Co.: J. Myers s.n. (July 2 pie (We). Randolph Co.: E. L. Core 62h (We). Roane Co. ve A. Bartholomew Ro-20 (We). Taylor Co.: Constable s.n. (Dry Run, v ee (We). Tucker Co.: A. H. Moore 2110 (Ca—~159478, Ge-- 1436). Tyler Co.: Bartholomew & Wilson 1.275 (We). Upshur Co.: = & Grose s.n. [Sago, ete ; 20, 1946) 7 (we) 5 We 3 Pollock apow Wirt Co.: West Va. Biol. Exped. s.n. [July 9, 1929] (#26357). NORTH AOE eg Rs Co.: Ahles & Bell 16843 (Hi--92729, Hi— 92730). Anson Co.: Ahles & Leisner 1995 (Hi--92731, Hi~—92732). Avery Co.: Britton & Britton s.n. [Cranberry Station, Sept. 13, 2885] (Cc). “Bertie Co. Ahles & Duke 16167 (Hi--121523, Hi— 12152h) 3 D. S. Correll 1987 987 (H—-hO2uS, N). N). Burke Co.: D. S. Cor- 354 PHYTOLOGIA Vol. 11, no. 5 rell 3088 (H—028)). Cabarrus Co.: Ahles & Leisner 19711 (Hi-- 92735). Camden Co.: D. S. Correll 2212 (H--9109); A. R. Moldenke 473 (Fg, S)- Caswell Co.: C. Re Bell s.n. (July 29, 1958] (Hi-- 121522); Oosting 33399 (H~15205). Catawba Co.: C. R. Bell s-n. (Sept. 8, 1958] (Hi—-1h6191). Chatham Co.: Houck 309 (Hi--9051h, Hi—92888) . Cherokee Co.: D. S. Correll 3559 (H—-0251). Chowan Co.: Ahles & Duke 7960 (Hi—-1029). Cleveland Co.: Ahles & Leisner 19143 (Hi--92734). Davie Co.: D. S. Correll 2739 739 (He 402)8); A. BE. Radford 118)8 (Hi--92889) ._ Edgecombe Co.: D. S- Correll 2Lh0 (H--l02)0). Forsyth Co.: Ahles & Duke 49010 (Hi 111296); D. S. Correll 2592 (H—-1987); P. 0. Schallert son. [7/ 1/33] (Du--236986). Gates Co.: Ahles & Duke 455 (Hi--11065h) « Granville Co.: Buell 1438 (No--8602, Faas WSO Greene Co.: De 8. aes 1368 (H—-L0252). Guilford Co.: Melvin s.n. [Aug- 1955) —-92891). Halifax Co.: Ahles & Leisner 17005 (Hi--92892). Hay- Co.: Ahles & Duke 50251 (Hi--112965); R. A. McLean 98 es A. R, Moldenke 51 [H. N. Moldenke 21775] (Bm); Oostang 31101 70422); Smathe: Smathers 3 57 (N). H son Co : Memminger s.n S.Ne ~[guly oe 1885] (Hi—-l9537); Oosting 351,87 (H-~31533) « Iredell Co.: C. Fe Hyams s.n. [State venaiie] ie %). Jackson Co.: C. W. Galloway 3101 (Hi—92738). Lee Co.: Houck 387 (Hi--92393) - “Yadison Co.: Ahles & to 146352 (Hi—121525). Martin Co.: A. E. Radford 39bh3 (H— )~ McDowell Co.: C. R. Bell 4505 (Hi—92 2894) « Mecklenburg ‘a Ahles & Duke 18501 (Hi--112968, Hi—~112969, Hi--112970)5 De Ss. Correll 3030 (H--9062). Mitchell Co.: Ahles & Duke 47078 (H—- 1OWL7s). Montgomery Co.: Ahles & Leisner 16305 (Hi—-9271)- Oram ge Cos: Blomquist 713 (H--20439) ; W. C. Coker s.n- [Sept. heb sol (Hi—59477); Radford & Stewart l99a (Hi--12595). Pasquotank C Ahles & Duke [8075 (Hi--10hl72). Person Co.: Duke 351 (ai-92782)- Polk Co.: D. S. Correll 3243 (H--l0239); Peattie 611 1/2 (Hi-- 594,80) ; E.C oy Townsend | 8.n. in. (7-20, 1899] (Se—-1495) « Randolph Co-! Millsaps s Son, {aug. 13, 1932] (Hi—3256). Richmond Co.? A- BE. Rad- ford 14h75 (Hi--92897). Rockingham Co.: A. E. Radford 1360, (Hiv 92899). Rowan Co.: D. 3. Correll 2801 (H--l02h7)5 A. B- Radford 14288 (Hi--92898). Stanly Co.: Ahles & Leisner 19855 (Hi Hi--92 1h, : Hi—-927k6). Surry Co.: A. E. Radford 19377 (Hi--92900). Swain Co? A. R. Moldenke 52 (H. N. Moldenke 21776) (Bm). Transylvania 00+? D. S. Correll 3398 (H—-N02h9). ‘Tyrrell Co.: A. E. Radford 39167 ic: Hi--10hL67). Union co.: Ahles 3050 (St); 4 & Haesl p juub2 : SIE) Bee Co.: Brimley 8 ( No—8601)’, a )- Wilson Co.: A. E. mains poe (Hi—104462) « fia Co.: A. E, Radford 15368 (Hi—-92003). County undetermined: Ash® 5: 1965 Moldenke, Monograph of Verbena 355 n. [July 19] (Hi--5948h). SOUTH CAROLINA: Abbeville Co.: A. E. Radford 25919 (Hi—92728). Anderson Co.: J. Davis s.n. {anderson, 8-7=19] (au); A. E. Radford 1027 (Hi~92886). Cherokee Co.: Ah- les & Haesloop 3100 (Hi--10L]27). Greenwood Co: A. E, Radford 26617 (Hi—-92737). Lexington Co.: A. E. Radford 27036 (Hi--92739). Marlboro Coe: A. E. Radford 15516 (Hi—92895). Union Co.: Ahles & Haesloop 31462 (Hi--104h06). York Co.: Ahles & Haesloop 3 (Hi—-10LL58, Hi--10h)59). Witherspoon Island [Darlington Co.]: Be E. Smith 1007 (Hi--2675). GEORGIA: Bartow Co.: W. H. Duncan 8535 (Gu—30152). Se: Co.: Drury s.n. [June 3, 1929] (Gu—U.6650);_ Holder £.8338 (Gu), s.n. [Athens, July 7, 1929] (Up); Miller & Maguire 1261 | 1261 (an-1696}, 1263 (Ba); Pyron s.n. (Athens, July - 12, 1929) (: (H—5l82). Richmond Co.: Cuthbert s.n. [Augusta, July 25, 1900] (Fl--21129). Thomas Co.: Mrs. A. P. Taylor sen. [Thomas- ville, July 28, 103] (Rf), sen. [Thomasville, Sept. 1903] (Rf). FLORIDA: Manatee Co.: S. M. _ Tracy 6652 6652 (W--38390,). Pasco Co.: C. Jackson 626 (Rf). ALABAMA: Baldwin Co.: S. M. Tracy 8037 (Au, N, N, tr, tr, * Tr, Up—50918, W--513695). DeKalb Co.: Moldenke & Woods 502 (S). MISSISSIPPI: Claiborne Co.: Voldenke & Woods 526 (8). Leflore Co.: Moldenke & Woods 511 (S). Smith Co.: S. Me Sen. (Taylorville, 3/21/1903] (W—510727), s.n. [Taylor- ville, 8/21/1903] (N). Sunflower Co.: Moldenke & Woods 510 (S). Tallaha chie Co.: Moldenke « Woods 506 (B, S). Warren Co.: Dem aree 25493 (Sm). Yazoo Co.: Moldenke & Woods 516 (S). OHIO: Bel- mont Co.: Guttenberg 251 (Cm). Butler Coe: Me Miller son. [July 7, 1932] (Ob-~80602) ; Wehmeyer & Waters 111 (Mi). Champaign Coe: Demaree 11704 (Bt—3039h, Ca--591979, “Du--2h070h, N, W--1632196); Jewett S.n. =n. (Urbana, July 28, 1838] ,(Mi); E. C.Leonard 1608 [Herb. Leonard 1,35: 4352] (W--2161287); A. R. Moldenke 797 (Iw). Clark Co.: F. P. Davidson s.n. (Springfield, July 160 July 1685) (Or—8872). Coshoc- ton Co.: Madison gn. (Co—-109); H. N. Moldenke 12095 (Co--39l) « Darke Co,; A. R. Moldenke Moldenke 800 (Lw). Delaware Co.: ot Re Crane 244k (N), 2506 (N). Erie Co.: C. L. Booth s.n. [August 2, T901} (Ob-- pee Hamilton Co.: Aiken 10693 (Ll), sen. [vIT~17-1905] (Po— 5 E. L. Braun s.n, [July 2h, 1917] (én); R. Buchanan §.Qe (11); J. Clark s.n. Gimtinnatt) (Bm, Bm); Frank s.n. eet 7] (Br) 5 C. G. Lloyd sn, [Sedamsville, July 1675] (Cn), 8.n. 1882; Herb. Prager 18635] (Gg—31h3h), sen. (July 31, gent (Ms), 5.n. (Cincinnati, July 25, 1883] (Ca--192071, Go); ue Mohr 455). naveison 66.1 A. Re Moldenke 788 (im, Ut). Holmes Co.: He N. Moldenke 1210) (co—-I01). Knox Co.: A. R. Moldenke 791 (LW). lake 00.0 i Co.: H. GC. Beardslee s.n. (Aug. 3, 1925) (Ob=97296), 2s 8 Ne adison] (0 295). c Co.: H. Le Jones sen. [Aug 1889} on] (oars En eens M. E. Day el (0b—=99797); Dick “ii 356 PHYTOLOGIA Vol. 11, no. 5 sn, (Oberlin, July 27, 1894] (Mn--18)92); F. E. Leonard 86/8 (Ob=-7531), sen. [July 20, 1896] (tim—18)) 3 Ee ae ee oo ee es 87871892) (cm) ; A. E. Ricksecker s.n 1, '9h] (W--217h41); A. A. Wright s.n. [July 30, sn a9} (Ob=ToHe Ob--75343), s.n. [Aug. 1, 1889] Passer a Lucas Co.: Sanford 3221 (S). Meigs Co.: C. H. Jones sen. {July 10, 1935] (N)- Miami : Clevenger s.n. (Fletcher, ay , 13, 1897] (W—152h139) 5 A. Re isiaaeke 799 (Lw). Montgomery Co.: H. P. Smith 1957 (Dt). Ottawa Co.: Re H. Moore sen. [Oak derbav. -galy 1927] (Ok—-10355), Sono (Oak Harbor, 1927] (Ok--10354). Richland Co.: E. Wilkinson 8467 (cm, Ob—8060h, Ob--80605), sen. [Aug. 1888] (Ob--00606). Ross” Co. Crowl 556 (N). Scioto Cos: Demaree 11518 (Ca--5920)8). Tus- carawas Co.: T T. S. Hopkins s.n, (Dundee, Aug. 1911] (Cm, Cm). Union Co.: A. R. elders 793 93 (Lw, Ut). Warren Co.: F. A. Bennett son. [July 16, “1884) (cm (Cm Wayne Co.: L. S. Hopkins Sen. . (6/25/ — li] (Br), sen. [7/3/11] as Thorne s.n, [Wooster, Aug. ug. 17/97] (Fe). ILLINOIS: Adams Co.: Brinker 2717 (I1--15819, Ur), 3489 (I1—15814), 3512 (I1--15813, Ur); Evers, Jones, & Jones 692 92 (N, Ur); A. Be Seymour s.n. {Camp Point, 19 Aug. 1876] (H--106L61) Brown Co.: V. H. chase 12760 (Ur, Ur, Ur, Ur). sige Co.: Ahles 2940 (Ur). “Champaign Co.: Evers 1484 (Ur); Fink s. . (Champaign, Aug. 16, 1892] (Io--28700); G. Ne Jones 12h39 9 (Ur), T9647 (Ur), 12829 (Ur), 1059 (Ur), 16528 (Ur), 1755 (Ur); McDougall 95 (Ur)5 Snyder s.n. (Urbana, July 1882 1] (Sd--h250). Christian Co.: DeMott sen. (Taylorville, 8/4/96] (Ur). Clark Co.: G. De ae er 10979 (T1—-19301). Coles Co.: V. A. Anderson 12595 (I1--2071h); “G. De Fuller 10700 (1121849). Cook Co.: H. H. Babcock 9282 Ae (ba=23793) Son. (vul. 1 13, 1814] (Sg--16116); Bakker s.n. [Gardner's Park, Sept. 1912] (Sd—1280)); R. Bebb 2107 (Ok); H. Re Bennett $Me {august 17, 1957] (Go), son. [August 31, 1997] (Go); Blatchford 2515, in part (11--15820), M. A. Chase s.n. [July 26, pie (Ur), sn. [Bowmanville, July 7, 1896] (Ur, Ur); Fuller & Link 1 2 (Il—- 15833); Lansing 1ild (Ur), 2806 (Ur); W. S. Moffatt 575 = 95719); Umbach 1303 (Ca--ki515). Crawford Co.: Ahles 4939 (Ur)- Cumberland Co.: Ahles 2574 (Ur); Winterringer 6071 (113326) - DeKalb Co.: P. B. Whitford s.n. [July 19, 19h6) (Ur). Douglas Co Winterringer 608 (Ur). Du Page Co.: G. D. Fuller 1355 Semone We S. Moffatt 293 (Ur), 575 (Ur); Umbach s.n. (Naperville, July 2 1898] (W--339L81). Edgar Co. A. R. Moldenke 809 (Lw)- getinghan rr Ahles 253 (UF) - Fayette Co.: O'Dell "Dell 337 (Ur). Hancock aacin - C. Gates 9948 (Ur). Iroquois Co.: Ahles 3211 (Ur); Winterring ee (I1—-34122). Jackson Co.: Fuller & Welch ee ais) efferson Co.: Ahles 568 (Ur); Winterringer 7803 (I1-—-3775 ° Daviess Co.: Ahles Bran) a it Kenko 1965 Moldenke, Monograph of Verbena 357 kee Co.: E. J. Hill 22.1873 (Ur, Ur); G. N. Jones ee (Ur). Lake Co.: R. Bebb 1681 (Ok); Umbach 5857 (Ca-—ll516, Ka). La Salle Co.: G. D. Fuller 1796 (T1—1583h); D. T. Ries Pee (I1— 15818) ; Thone 50 O (Ur); P iT Be Whitford 310 (11--32223), Lawrence Co.: Sivert s.n. {July 2S, 1952] (Ur). Lee Cos: —— (I1—31)90), s.n. [July 30, 1946] (Ur); Lyman s.n. (Amboy, 6-16- 94] (Dt). Livingston Co.: G. D. Fuller 9138 Ginaseis). Logan Co.: Winterringer 511 11 (Ur). Macon Co.: Clokey 253 (Ca—882652); A. R. Moldenke 817 (Lw); Winterringer 6599 (113110). Madison Co.: Demaree 27288 (N). Mason Co.: Ahles 298) (Ur), L110 (Ur); Ehringer 1210 (ITl1~-18978); G. D. Fuller 22h (I1—15811); F.C. Gates 3519 (Ur); G. N. Jones 22387 (Ur). Massac Co.: Winterringer 3554 (Il1—28979). McDonough Go.: Co. Re M. Myers M.83 (I1—23857). UcHenry Co.: Nason s.n. [Algonquin, Aug. 4, 1878] (Ur), s.n. (Il-- 20860). Mercer Co.: Winterringer & Fuller 2868 (I1--2769)). Mor- gan Co.: L. Carter 15255 (I1—~32313); A. R A. R. , Moldenke 827 (Lw). Ogle Co.: A A. E. Hills Is 3420-0 (I1--15821). Peoria Co.: Heading s n. (July 1875] (Ur); F. E. McDonald s.n. [Peoria, = 1887) (ur), Son. (Peoria, July 1889] (Ur), sen. (Peoria, July 1894] (Ca-- 10866), s.n. [Peoria, July 1903] 3) (N). Piatt Co.: anise 3310 (Ur); G. N. Jones 20771 (Ur). Pike Co.: A. R. Moldenke ienke 837 37 (Ly); H. N. Moldenke ES 13 (Lw), 21914 (Lw). Pope Co.: Whiting 1757 gl (T1-158h1). Pulaski Co.: Winterringer 5998 (I1--33174). Putnam Con: Rw G ate sen. (July h, 1941] (Ur). Randolph Co.: Winter- ringer ero (I1—3h00) . Richland Co.: R. Ridgway 1086 (Ur), 1155 (Ur), 1157 (Ur), 1158 (Ur), 3061 (Se~-29293), 3167 (La); Scherer 536 (T1—26358, Ur). Rock Island Co.: E. T. Harper s.n. (Port By- ron, August 1887] (Io—133293). Saint Clair Co.: Eggert s.n. [a- cross river from St. Louis] (Go). Sangamon Co.: ahles L591 ( (Ur); : oo be Puller Fuller 231 (I1—15831), 4870 (I1--15836), ieee oa (T1—15860), 5087 (I1~25840), 5237 (I1—15835), 5167 (I1— cy 5227 (1115037), 5257 (1115838), 5352 ‘ele Fes), 5565 (1115837), 6119 (1115826), sen. [8/5/k1] (I1--15825)5 Ae Re Moldenke 822 (Im). Scott Co.t A. R. Moldenke 830 (Lw). Stark Co.: V. He Ch La Chase | son. [July 10, 1895] (Ur). Tazewell Co: V. He Chase 3228 (Ca—8828),2, Ca--988091, [1—~-15817, N, Ur, Ur, Ur). Union Cow: G. De Fuller 676 (Ur); Fuller & Fuller 228 (1115810), 676 (I1—15830). Vermilion Co.: Ge Ne Jones nes 12359 (I1--15822, N, Ur), 12957 (Ur), 14376 (Ur); Storm s.n. {duly 28, 1949] (Ur, Ur). Wa- bash Co.: Schneck s.n. [July 30, 1879] (ur), sen. (Sept. 1, 1887] (Ur), s.n. [July 2088) (Ur); Shearer s.n. (Mt. Carmel, July 20, 1900} (Vi), sun. [7/20] (Ur). Wayne Co.: M. Walker s.n. [Sept. 5, 1949] (Ur, Ur). Whiteside Co.: Ahles 196 ligé (ur . BOOK REVIEWS Alma L. Moldenke "Chemical Plant Taxonomy", edited by T. Swain, viii + 513 pp-, illustr. Academic Press, London, England & New York, N- Y- 1963. 110 sh. or $16. 00° This is a valuable adventuresome book that bridges the gap be- tween yeste and tomorrow in plant taxonomy, as well as the gap between organic chemistry and this phase of botany. It con- sists of sixteen pertinent papers that were presented at a sym- posiun or ee cid ge hee PHYTOLOGIA Designed to expedite botanical publication = Vol. 11 March, 1965 No. 6 CONTENTS SWALLEN, J. R., & TOVAR, O., The grass genus Dissanthelium .... 361 WURDACK, J. J., Certamen Melastomataceis IX ....-++++000000 000+ 377 MOLDENKE, H. N., Materials toward a monograph of the genus Verkena, XXUI 2 Sag ee ee ge eae 400 REED, C. F., Cleome ornithopodioides L. on vanadium-slag at ts Canton, Baltimore, Maryland with notes on the biogeochemistry OF PABANMM cys oon ee eee Re ee seeeess 423 ba : ! ° -MOLDENKE, Ao is, Rook vevieine 0 te Ce eas eke ee ike Published by Harold N. Moldenke and Alma i ene - a Chauid Aves: - Yonkers 5, New York, U SAL ee . Price of this. number, $1; per solace. $5. 1, in a advance, = 162 at close of volume oS ce . eeorer "Sos 1f no. 4, was is sued. Jar ee MAR 8- 1965 THE GRASS GENUS DISSANTHELIUM Jason R, Swallen and Oscar Tovar genus Dissanthelium was founded by C. 8. Laban in pate triisece 10: 305. 1536) based on a single species, num, free Mire "in frigidissimis ad Cerro de Pasco Tinetibe ore, Poeppig)." It is characterized fae two awnless flatets: ihe exceeded by the equal glum The name Dissanthelium was derived from the Greek word eaSc08, dois and anthelion, a small flower, alluding to the mall florets (Hitchcock, "Manual of the grasses of the United States," p, 173. 1935), The type species of the — was first described by Presl as Brizo cal after was Soubiracaid to Poa by Tati aver eT at thoat hav having seen the plant. Hitchcock examined the type of es 8 spec treaioseens cinum) at the herbarium of the German “University in Prague ound that it was the same as the species (Dissanthelium supinum) which he examined in the ‘a ium of the Academy of uatetcet at Leningrad, Fragments from the types are in the U. S, National Herbarium. On grounds of prior- ity the Sane name must be adopted, The second Peruvian species s E Pilger described the e species ce Dissantheliv minimum, basing it on Weber aiiak No. 5451 fro Dissanthelium is a genus exclusively American occurring principally in the secre! Andes of South America at altitudes between 11000 and )800 meters, The greatest concentration of Species is in the Peru ante Andes. One species, D. californicum (Nutt.) Benth, , is confined t o islands off the southern os S.A. 3 s Mexico and the central Andes; D, patagonicum Parodi is pele nown from the Patagonian region of Argentina; and the other 1h cee aenetenaeetesinememerenenees Dr. Oscar Tovar is from the University of San Marcos, Lima, Peru, He studied grasses at the U. S. National Museum from September 28, 1959, until December 1960 under a Fellowship from the Jo imon Guggenheim Memorial Foundation. He prepared the uscript of "Dissanthelium" at that time. I had been inter- ested in the genus for some time previously and had designated Several new species which are described herein. J.R.S, 361 362 PHYTOLOGIA Vol. 11, no. 6 species are from the central Andes of Peru and Bolivi Dd. calycinum (Presl) Hitche, and D. minimum Pilger are “aa distributed dwarf species of the high Andes. eculiar Sigg tana a Gaede ares of the species pa se sitesi bisa ae central Andes is of interest. They oc only in the "and are never sagt in the "Paramo" of Ecuador and Koken, Neither are they found in the "Jalca" of northern Peru which has a high humidity in comparison with the A. S. Hitchcock, in his "Dissanthelium, an American genus of Grasses" recognized only 3 species. The apie paper includes 17 species, 10 of which are described as new. All the species considered have been examined in the U. S. National scien ops sau Smithsonian Institution. The types and isotypes e deposited there and without these it would have been impos- sible to solve certain problems satisfactorily. Most species of Dissanthelium are dwarf; few of them are “ more than 10 cm, tall. They are mostly perennial with only three annual species, The culms are mostly tufted, erect, glabrous, puatties decumbent at the base, and in a few cases rhizomatous, The sheaths are mostly glabrous, only in D. p atagonic™ my Dd, 1 ligulatum, and D. densum are they minutely sca pear a rauhii, e blades are usually narrow, involute, ina fe cases folded or flat, soft or subcoriaceous, mostly scabrous oF 4 scaberulous above, glabrous beneath, 5 eae or spreading, the apex acute » Subacute, or obtuse The panicles are dense or somewhat open. The branches may be appressed, ascending or spreading, floriferous to the base or sometimes naked below for a short distance. The pedicels are scabrous or scaberulous, except in D. semitectum where they are minutely villous, The small arg may be 2-3-flowered, but they are usually 2-flowered, the lower floret —— a little longer than the upper. The igen are > nearly so, 3-nerved, 4 tse ery a _ the . (Lechler 1836). Oraninastrus ti esr yon Krause, Beih, Bot. Centralbl. 32: 348, 191. Based o n Vilfa macusaniensis Steud. 36 PHYTOLOGIA Vol. 11, no. 6 Annual; culms . tall, slender, erect, more or less filiform, glabrous; witlcered glabrous, membranaceous at the base, 7 dense, the branches appressed, floriferous to the base; s 3-4.2 m, g, 0.5 mm, longer than the florets, ovate- lanceolate, glabrous or minutely scaberulous on the keel toward the summit, somewhat scarious toward the margins, 3-nerved; s lower lemma 2,5-2.8 mm. long, ovate-oblong, trifid, ng middle tooth longer than the others, membranaceous, hyaline oward the t erved, the lateral nerves see to the margin; palea a little shorter than the lemma, the narrow p somewhat bifid, minutely begin on oo keels; rachilla joint 0,3-0.) mn, Lone; anthers mm. long. Type: oot of Peru, ian Pisco and Ayacucho, Weberbauer 5) s aisle Open ground in the Andes of Peru and ecg between 3900 and F600 meters altitude. Peru: Ancash: Prov, Bolognesi: between Llamac and Jahuaco- cha, Dist. Pacllén, Cerrate 2323; Conococha, Cerrate has Mar- caré, Velarde 3202, Pasco: Cerro de Pasco, Hitchcock 222h0. Junin: between Tarma and Jauja, Killip & Smith 21951. L Prov, Huarochiri: ayo Drspe Lima and Oroya, Asplund 11775 Viso, between Lima and Casapalca, Macbride & Featherstone 628; around Tuctucocha Lipoid Cerrate 1856; Prov. Canta: near Antacocha, carro Colorado, east of Canta, Pennell 14654, Huan- re) n P~608, La vo Rauh & Hirsch P~668, a Paz: Copacabana, near Lake Titicaca, Asplund 1348h; La Fax; Puchtien 6420. Oruro: Challapata, on railroad between Oruro Uyuni, Asplund 9545, Larecaja: vicinity of Sorata, jr stetacnes between Pongo and Anilaya, Mandon 13h6. 3. DISSANTHELIUM PER oe M (Hees & Mey.) Pilger, Bot, Jahrb. Engler 37: 378, Phalaridium the teers Nees & Mey., Nov. Act. Acad. Caes. Yeon. Carol 19; Suppl. 1: 29. 181; 161. 18h3. ual; culms somewhat —. at the base, 2,5-13 cm tall, viteoas: sheaths glabrous, membranaceous at the base, the uppermost elongate reaching the Fes of the seeped: ligule 1 =k = es 1 1 ° flat, smooth, soft, glabrous on both surfaces, ascending, the uppermost rhe By the mn of the panicle; panicles 1-3.5 cm. 1965 Swallen & Tovar, Dissanthelim 367 2.8-3.5 mm, long, 0.8-1.3 mm. longer than the florets, ovate- lanceolate, glabrous or sca ~ ge on the keel toward the ed; 1 margins; palea a little shorter than the lemma, the narrow ano ee glebrous; rachilla joint 0,3-0.) mm, long; anther ong. ie "Lake Titicaca, Peru, Me istribution: meget ed to the ae Andes of Peru and Bolivia, Pepe 3700 and 200 meters altitude, Peru cash: an Bolognesi: vicinity of Lake Condorco- cha, pouaes "2087 ima: Prov. Huarochiri: sine between and dé ca? ro yacaja: Millpu, Puna Tocas, between Colcabanba and “site seas Tovar 1950; Misesesda Alalay, between Mariscal Caceres Pampas, Tover 1350; Prov. Castrovirreina: vicinity of Laguna eosin Tovar 2826; Prov. Huancavelica: Sachahuaccta, 6 Km. N.W. si m Conaica, Tovar 9 Bolivia: La Paz: Incachaca, Asplund 9549. Potosi: without locality, D'Orbigny 136, 186, 191; General Campero, on railroad between Viacha and Arica, near Corocoto, Asplund 1390 4. DISSANTHELIUM TROLLII Pilger, Notizbl, Bot. Gart. Berlin, 11: 778. 1933. Perennial, hig tall; sheaths iiaicuaiaaatas pte eee scaberulous, the lower shorter than the upper; li embranaceous, the tip toothed, 0.5-1 mm. long; blades 1.5-3 notes ng, 1-2 mm, wide, Spreading, flat or sometimes nearly involute, acute, soft gag erulous on both surfaces, scabrous on the margins; panicles 3-2 cm. long, 0.5 cm. wide, few-flowered, the branches appressed, floriferous to the base, branchlets 2 desing glumes equal, 4.5-5 mm. long, 1.5-1.8 mm, wide from keel to margin, as long as the florets - senations 0.3 mm, sani du Scaberulous toward the ih enindi on the margins, 3-nerved; lower lemma h-),3 mm. long, 1.2 mm. wide from keel to margin, obtuse, nes a rote 3-nerved; palea a little carhes than the tip sul prance glabrous, the nerves eye ssa seaciane porn 0.3 mm, long, stout; anthers 2-2,2 mm ng. Type: "Bolivia, San Edwardo bei La Union, C, Troll Te " Distribution: sete only at high ernst (Puna) o Bolivia, between 650 and 5000 meters altit Bolivia: La Paz: neous 30 Km, from - Paz, — eg : La Cumbre, railroad to Yungas, Parodi 100h2; e 10, 5. Dagsasenemttia PYGMAEUM Swallen & Tovar, sp. n culmi caespitosi, erecti, 2-3 cm. alts, glabri; vaginae Dans basin membranaceae; ligula 0.7-1 mm, longa 368 PHYTOLOGIA Vol. 11, no. 6 membranacea; laminae 1-2 cm. longae, conduplicatae vel planae, obtusse, supra sparse pubescentes, subtus glabrae; paniculae 1-1.5 cm, longae, 0.5-0.6 cm, latae, aoe pauciflorae, ramis appressis vel adscendentibus, glabris; glumae aequales, 45-5 mm, longae, quam flores 0.) mm. longiores, ion gs ornate es; acuminat nodium 6.3 mm, longum; antherae 2 mm. oblong-lanceolate, 3-nerved, the nerves prominent; lower lemm 4-k.5 mm, long, subacuminate, 3-nerved, glabrous; auias a little shorter than the lemma, the narrow tip more or less truncate, membranaceous, glabrous; rachilla joint 0.3 m, piel anthers 2 mm, long. Type in the U. S. National Herbarium, No, 2207161, lected in Huaytanayoce-Tansiri, near Manta, District of ear Deva and ude 1,500 45). ion: Known only from the type locality. 6, DISSANTHELIUM PATAGONICUM Parodi, Physis 8: 80, fig. 7. 1925. we rennial; culms tufted, erect, 12 cm. tall; sheaths sca- the lower shorter than the upper, membranaceous at the base, 6 uppermost elongate, somewhat coriaceous; ligule — * long, membranaceous; blades 1-2 cm. long, ascending, r es scaberulous; panicles 5,5 cm. long, 0.6 cm, wide, narrow, few-flowered, the branches appressed or sometime i floriferous to the base, the branchlets appressed oF SO! ascending, stiff, scabrous; s long, nearly as long as the florets or somewhat longer, linear- lanceolate or subacuminate, glabrous or scab s, coriaceous, 3-ne 3 lowe -4-2.5 mm, long, ovate-lanceolate, obtuse or subacute, densely villous, 3-nerved; palea a little shorter than the lemma, the — od bifid, villous on the keels; ae joint. es 8-1 mm e: "San a Crus: Bio Gallegos, leg. L. Dauber No. 180, verano v1922" pedantic Distribution: Known only from the type locality. Te east gy _— Swallen & Tovar, aespitosi, erecti, 43-16 a “atts minute oni. vg eteen vel ecaberulae basin age oa ligula 1-1.5 mm. longa, membranacea; laminae 6-10 cm, longae involutae, supra pubescentes, subtus glabrae vel minute scabrae, 1965 Swallen & Tovar, Dissanthelium 369 adscendentes vel appressae; paniculse 3.5-.5 cm. longae, 0.7 cm, 1 te pauciflorae, ramis appressis vel vix adscendentibus, rum, 3-nerve; palea lemmate paulo oe in carinis minute precedes rhachillae internodium 1 mm Perennial, somewhat ehdaemabeul fas culms tufted, erect, aE Svarsely scabrous, 13-16 cm, tal : ae glabrous or ely scabrous, membranaceous at the base; ligule membranaceous, 1-2.5 mm. long; blades 6-10 cm. long auenbeateds acute or subacute, some of them naked below; glumes equal, -3 mm. long, nearly as long as the cies linear-lanceolate, st on the keels toward tip, mee: the lateral nerves conspicuous only at the ana lower lemma 5.5-5.8 mm. long, linear-lanceo- te, scabrous, 3-nerved; palea a little shorter than the lemma, the tip bifid, “miméitely ‘ciliate -on-thé keels; rachilla joint - long. (No. 1292A), This specimen was mixed wit th ae 1292, Distribution: Known only from the type locality. 8. DISSANTHELIUM LONGILIGULATUM Swallen & Tovar, sp. nov Perenne; culmi erecti, basi decumbentes, 12-15 om. alt - glabri; vaginae elongatae, ” scabridae, compressae, suprema basin paniculae aequans; ligula 2-6 mm, longa, membranacea, vix truncata; laminae =a sony longae, ae rigidae, supra sca Scabrum, 3-nerve; palea 2.6-2, 8” mm, longa, membranacea, plabra, truncata; rhachillae internodium 0,-0.5 mm, longum; anther 9 moe 7 mm. longae. Perennial, rhisomatous; culms erect from a decumbent base, 12-15 om, tall, glabrous; sheaths peyton scaberulous, com- Pressed, the lower shorter than the upper, the uppermost reach- embre ar Spikelike, the branches somewhat appressed, some of them na at the base for a short distance, the branchlets ascending, stiff, scabrous; glumes equal, 6-6.5 mm. long, 2-2.5 mm, longer than the florets, oblong-lanceolate or subacuminate, glabrous, Sometimes scaberulous on the keels, 3-nerved; lower lemma 3,8-4 370 PHYTOLOGIA Vol. 11, no. 6 mm, long, 0,8-1 mm. wide from keel to margin, subacute, scaberu- lous, 3-nerved, the lateral nerves close to the margin; pa — 2.6-2.8 mm. long, somewhat membranaceous, glabrous, the nar tip truncate; rachilla joint 0.4-0.5 mm. long; anthers 0. wer’ 7 mm. long, Type in the U. S. National Herbarium, No. tae col- lected in La Paz, Bolivia, no at 18,900-20,000 feet January 20, 1926, by A. Guerrero s, Distribution: Known only ioe the type locality. 9. DISSANTHELIUM SEMITECTUM Swallen & Tovar, s erenne; culmi caespitosi, basi decunbentes, "9-13 em, alti, gies) vena glabrae, suprema basin paniculae aequans; ligula 0.5-1 mm, longa, membranacea, subacuta; laminae -7 cm. longae, involutae, acutae, supra scabrae, subtus glabrae, a esceneaennl paniculae 3.5-.5 cm. longae, 0,8-1 em - latae, ramis vix esdamensitel: basin floriferis, pene ee vix divergentibus; spiculae 2=3-florae (plerumque of hates) ? pediceliie v villosis; — aequales, ).46-5,3 mm. longae, ino 1.82 a palea lemmate vaulo brevior, truncata, in carinis minute scabra; rhachillae internodium 9,3-0.) mm. longum; antherae 0,/\-0.5 mm. longae, Perennial; culms tufted, decumbent at the oe BL seme scabrous eile) glabrous beneath, ascending; panic . em. long, 0,8-1 em, wide, rather loose, the branches spre Selow for 0 floriferous Fe the base or some of + a eked e dist ate, scarious toward the margins, 3-nerved, glabrous or minutely Scabrous on the keels; lower 2.5-2.7 mm. long, som oblong-lanceolate, acute or subacute, scabrous, 3-nerved; re e a little shorter than the lemma, th ow tip truncate, sca berulous on the keels, aesdiranscecuss rachilla joint 0.3-0.) m. long; anthers 0,-0.5 mm, e in the U. S. meena Herbarium, No. 1161 ip col- lected bn clumps on northeastern rock ledge to Huaron, Depart- ment of Junin, Peru, altitude about 1,000 feet, roe "12, 1922, by Macbride & = tan (No, 12 Distribution: Known only rte the type locality. 10, DISSANTHELIUM LAXIFOLIUM Swallen & Tovar, s Perenne; eculmi caespitosi, basi deci’, cau: 8-13 cm, — glabri; vaginse plabrae, — pete ranace i supr a 1965 Swallen & Tovar, Dissanthelium adscendentes, supra minute scabrae, subtus glabrae; paniculae 3-4.5 cm. lo ongae, 1-1.7 cm. latae, — adscendentibus vel glumae aequales, 3.5-l; mm. longae, PH 1,.2-1.6 mn, es, vix suba natae, glabrae v cari caberula i be nerves; lemma inferius 1 e- ira btusum mm, lon obtu acutum, minute scabrun, 3; palea Lenmate paulo brevier, in carinis minute ciliata; untested internodium 0.3 mm, longum antherae ce $s 5 mm. longae. erenni erect, elation, 8-13 cm. tall; heathe glabrous, membranaceous a panicle; ligule membranaceous, the uppermost 2-2,5 mm. long; blades 2.5-8 em, long — innovations filiform), soft, aaa or rarely somewhat flat, pointed, ascending, minutely s scabrous “go glabrous beneath, subacute or rether flexuous; panicles 4.5 cm. long, 1-1.7 cm, wide, loose, the branches ascending or ore ading, some of them naked at the base for a short distance, the branchlets ascending; glumes equal, 3.5-) mm mm. longer than the florets, line ar-lanceolate, or sometime subacuminate, arcuate, gon - eer aie on the kee “ate toward the summit, scarious argins, 3-nerved; lower lemma 1,9-2,1 mm, * long, wep pees or subacute, minutely scabrous in the upper half, 3-nerved, the nerves close to the margin; palea a little shorter than the lemma narrow tip truncate, more or less hyaline, glabrous or nd nut ely scabrous- Pa on the keels; rachilla joint 0.3 mm, long; anthers 0.4- mm : lected at cixipas along souharees valley, Province Bolognesi e 000 meters, Mav 29, 1956, by atcha Peru at altitudes from 4000-1600 — Peru: Junin: vicinity of Aguilar inicavalies: Prov. Castrovirreina, near Lake Siccicoule, Tovar 2833, li. fon nee BREVE Swallen ’ supre ema ’ basin paniculae aequans; 1-2.5 mm. longa, membranacea, vix truncata; laminae 1,5- 345 em, longae, adscendentes vel divergentes, conduplicatae vel planae, obtusae vel subacutae, supra scabrae, — glabrae; and. @ 2-3 cm. longae, 1-1.3 cm, latae, densae sdacenientibue, in et inferiore " nudis, ramlis recs eae a bus; glumae aequales, 5-5 mm, longae, floribu ngi- t, minute scabrae doh nerves; lemma inferius 2.5-2.7 mm. obtusum, minu te strane “Zen nerve; palea lemmate paulo brevior, truncata, in éortake minute ray FoThe ange internodium +3 mm. longum; antherae 0,5-0,.6 mn, al; culms tufted, erect, ag ares 5-8 cm, tall; Perenni sheaths glabrous, membrana ceous at the base, the uppermost 372 PHYTOLOGIA Vol. 11, no. 6 reaching the base of the panicle; ligule ip pe ree, 1-2.5 alo some t i runcate; blades 1.5-3.5 em. long, gubcoria- ceous, spreading or ascending, flat or folded, “obtuse or sub- acute, scabrous above gong beneath, dull, the nerves conspicuous; panicles 2-3 cm. long, 1-1.3 cm. wide, dense, the branches ascending, aad below for a short distance, the branchlets ascending; glumes equal, )|.5-5 mm, long, 2 mm, longer than the florets, oblong-lanceolate or subacuminate, sometimes arcuate toward the tip, glabrous or minutely scabrous on the keel, scarious on the margins, 3-nerved; lower lemma 2.5-2.7 mm. long, oblong, obtuse, scarious toward the tip an aaa minutely scabrous, 3-nerved; palea a little shorter than the emma, the tip truncate or somewhat bifid, minutely ciliate on keels; rachilla joint 0.3 mm. long; anthers 0,5-0.6 mm ong in the U. S. National Herbarium, No. 2181263, lected in Huaytanayocc-Tansiri, near Manta, District of Gonatce, Department and Province Huancavelica, Peru, — 00-500 meters, March 31, 1953, by Oscar Tovar is ane re Ande Peru a eru: Ancash: Prov. Bolognesi: hive a Dist. of Ticllos, Cerrate 2661; Hane nti Blanca, Rauh & Hirsch P-20h2. Hudénuco: Prov, Dos de Mayo: Cordillera a Raura, near glacier, Cardich s,n, Junin: Prov. Yauli: Anticona Pass, between Lima 282, 2878, 2887; Prov, Huancavelica: Huaytanayoce-Tansi Dist. of Conaica, Tovar 2525, Cuzco: Ausangate, Rauh & Hirsch Bolivia: La _— Chacaltaya, 30 Km, from La Paz, Buchtien 119}, La Cumbre, on railroad to Yungas, Parodi 10037. Prov. Larecaja, vicinity: be Sorata, near Anilaya, Mandon 135A. 12. DISSANTHELIUM CALYCINUM ne onagd Hitehe, Journ, Washington ha Acad, Sci, re 22h. - Pin calycinum nee "Rel. Haenk. 1: 281. 1 Pos ealyetia mth, Rev. Gram, i: Suppl. XXVIII. 1630; Pie ls 326. 1833. ar eerene Pai supinum Trin, Linnaea 10: 305. 1836. (Cerro a Saar Peru, Poeppig). Perennial; culms tufted, 2.5-8 cm, tall, glabrous; sheaths glabrous, mesbranaceous at the base, the lower shorter than the upper, the uppermost elongate; ligule membranaceous, 0,5-1 eee invol ing, scaberulous above, glabrous beneath, the uppermost a reaching the apex of the panicle; panicles 1-2 cm. long, obloné florife pirat to ume equal, 2.4-3 mm, long, 0.5 mm, longer than the florets, ovate- lanceolate or sometimes subacuminate, glabrous, 3-nerved, the 1965 Swallen & Tovar, Dissanthelium 373 margins scarious; lower lemma 1,8-2,1 mm, long, ovate-oblong, obtuse or ate te, scaberulous on the upper half, subcoria- ceous, 3-nerved, the lateral nerves close to the margin; palea a little shorter than the lemma, the narrow tip truncate, minutely ciliate on the neni membranaceous} ert joint slender, 0.2-0,.3 mm, long; anthers 0,5-0.7 mm. long. ype: Original locality not known, but probably Peru where it is widely distributed, Haenk Distribution: High altitudes (Puna) of Peru and Bolivia between 4200 and 900 meters altitude, Peru: Lima: Prov. Huarochiri: Chic la eaeraa Lima and i Prov, Junin: Hacienda Atocsaico, near Junin, Hitchcock 22207, 2221), Pistpee 22204; Prov. Yauli: Ticlio aed Rauh & Hirsch P-90, P-306; Prov, Jauja: Eda. Cachi-cachi, Km. 35 between — and Tarma, Velarde 2868, Huancavelica: Prov, ig i pe ina near Laguna Choclococha, Tovar 281; Prov, Huancavelica: Huaytanayocc-Tansiri, Dist, of Conaica, Tovar 1209, 2 hycadhioy Hebeses Huanta and Hacienda Pargora, fait Smith 23305, Puno: Prov. Carabaya: Hacienda Lacka, Macusani, Vargas 7891, Cuzco: pare pane - Hirsch P-1139. Bolivia: La P da, railroad between La Paz and Pongo, Hitchcock 20596 ‘fon "wBolivian Plateau," Bang 1873. 13. DISSANTHELIUM SCLEROCHLOIDES Fourn, Mex, Pl, 2: 112, _. Deschampsia mathewsii Ball, Journ, Linn, Soc, Bot. 60. 1885. (Casapalta, Peruvian Andes, Crocs al; culms tufted, erect or somewhat geniculate at renni the bases glabrous, 5-9 cm, tall; sheaths glebrous, membrana- ceous at the base, the lower shorter than the upper, the upper- memb: -1,3 1,5- cm. long, involute or sometimes somewhat folded, smooth, ascending, ged glabrous, the uppermost almost = the apex of the panicle; panicles 1.5-3 cm. long, 0.7-1 cm e long, ovate-lanceolate, scabrous, ed, i Bh the margin; palea a little shorter than the trun ate, a scabrous-ciliate on the a beater the tip; rachilla Bae nt 0,3 mm. long; pe inch 0.6-0. long. Fournier mentions wo specinens, Mevado de Toluca, Hahn, and San Luis de Potosi, Virlet 1s3, Mexico Distribution: High altitudes of Mexico; sail Andes of Peru — Bolivia, exi.co: Nevado de Toluca, Pringle 222; Beaman 1692; Maloe, Beaman 2329; Ixtaccihuatl, Purpus 1633; Matuda 374 PHYTOLOGIA Vol. 11, no. 6 3800. Tlaxcala: Malinche, Bosman 22ho. Peru: Huancavelica: P . Huancavelica: Huaytanayocc- Tansiri, Dist. of Conaica, sep Bolivia: Without locality, Mandon 135. 1k. DISSANTHELIUM repre Swallen & Tovar, sp. caespitosi, erecti, é-8 we nati, glabri; vaginae Eo scabrae, basin membranaceae, suprema basin paniculae aequans; ligula 1-2 mm, longa, membranacea; laminae 2-5 cm, longae, adscendentes, acutae, conduplicatae vel planae, scabrae; vaniculae 2-3 cm, longae, 0,7-1 cm. latae, densae, ramis rigidis, appressis, ad basin floriferis, ramulis adscendentibus vel appressis; glumae aequales, 3.5-3.7 mm longae, floribus 0,6-0,7 mm, longiores vix lanceolatae, glebrae, 3-nerves; lemma inferius 2,5-2,6 mm, longum, obtus minute hispidum, 3-nerve; palea lemmate paulo brevior, in carin is mimte ciliata; rhachillae internodium 0.3 mm, longum; antherse 0, siti 5s Sa longae, nnial; culms tufted, ee glabrous, 6-8 cm sheaths Pa minutely scabrous, membranaceous at “the base, the rmost reaching the base of lie panicle; ligule membrana~ fuhicae 1-2 mm, long; blades 2-5 cm, long, ascending, folded or sometimes flat, acute, minutely scabrous on both surfaces, the les 2 » 3-nerved, membranaceous toward the tip; p little shorter than the lemma, truncate, minutely pe Ge Re a the keels toward ate summit ; rachilla joint 0,3 mm, long; anthers 0,5-0,6 mm. long. nthe t U. S, National Herbarium, No. 2207162, col- ay Department and Province Huancavelica, Peru, altitude 500 meters, May 11, 1956, by Oscar Tovar (No. 257), Distribution: Known only from the type locality. 15, DISSANTHELIUM EXPANSUM Swallen & Tovar, sp. renne; culmi caespitosi, basi ctl ak "6210 cm, alti, glabri; vaginae sagtuts noabean basin membranaceae; ligula 1- um membranacea; laminae 1-l, cm, longae, conduplicatae vel planae, adacginieres. eee obtusae vel subacutae; paniculae 2-3,3 cm, longae, 1.5-2 cm, latae, laxae, ramis Ser eine basi nudis, ramulis adscendentibus 2 aoe scabris; glumae aequales, 3.5-l mm. longae, floribus 0.6 mm, longiores, sblongo-lanceclatac, . Rabees vel an carina minute scabrae, 3~ nerves; lemma infer 2.4-3 mm, longum, oblongum, obtusum, minute scabrum, 3-nerve; palea a lemmate paulo brevior, in carinis minute nity rhachillae internodium 0.3 mm, longum; antherae 09-1 mm. longae, Perennial; culms tufted, stout, somewhat decumbent at the 1965 Swallen & Tovar, Dissantheliun 375 base, glabrous, 6-10 cm. tall; leaves mostly basal; sheaths short, minutely seatirohes menbranaceous at the base, the upper- me ! cm. long, flat or folded, ascending, soft, glabrous on both margins; panicles 2-3,3 em, long, 1.5-2 cm. wide, lax, branches spreading, naked below for a short distance the branchlets ascending, sparsely scabrous; glumes equal, 3.5- mm, long, about 0.6 mm, longer than the florets, oblong-lanceolate, glabrous or pene i scabrous on the keels, 3-nerved, purple toward the s summit; lower lemma 2.):-3 mm. long, more or less oblong, obtuse, minutely scabrous, 3-nerved, the lateral nerves close to the margin; palea a little shorter than the lemma nearly bifid at the tip, ‘iuiuhals ciliate on the Senis toward the tip; meee joint 0.3 mm. long; anthers 0.9-1 mm. long. i on e eu. S. National Herbarium, No. 0, col- lected in eae Panticalla — Peru, altitude about ters, June 18, 1915, by 0. F. Cook & G, B, Gilbert (No. 1305). Distribution: High Andes of ies Peru: Pinasniocc, Panticalla Pass, Cook & Gilbert 1297, Fudmcos 1s miles N.F. of Hudnuco, Macbride & Featherstone 2183, ie, : Ticlio, pe of Rio Rimac, Prov. Huarochiri, Hutchison 16, ieee Mp en Swallen & Tovar, sp. te truncata, glabra, vel in carinis minute scabra peace internodium 0,-0.5 mm, longum; antherae 0, bb, 8 mm, Sone sia al; culms stout, tufted, glabrous, 2-l om ip tall the base of the panicle; ligule membranaceous, trun rather OY i 0.6-2 mm. long; blades os em, long, coria- nding naked 3 glume ®qual, subacute, 3-3.l: mm. long, more or less ovate, as pen as the florets or nearly so, glabrous, rather scarious toward the mar long; anthers 0,6-0.8 mm. long. e in the U, S, National Herbarium, No, 1161062, col- 376 PRETO L007 & Vol. 11, no. 6 lected on loose "doby" slope, at Yauli, Department Junin, Peru, altitude about 13,500 feet, May 25, 1922, by Macbride & Featherstone (No, 933). stribution: High Andes of Peru ru: Junin: Huarén, northern part, Cerro de Pasco region, Macbride & Featherstone 1153. Pasco: Cerro de Pasco, Hitchcock 22228, 22233, 222h2. 17. DISSANTHELIUM RAUHII Swallen & Tovar, sp. Perenne; culmi caespitosi, erecti, 6-9 ea opti, glabri; vaginae glabrae, membranaceae, suprema basin culae aequans; ligula h-5 mm. longa; laminae 1.5-3.5 cm. longae, adscendentes, coatuptinatne vel planae, obtusae, supra minute scabrae, subtus glabrae; paniculae 3-l; cm, longae, 0.8-1.5 cm, latae, densae, ramis adscendent ntibus ver appressis, in parte inferiore nudis, ramulis adscendentibus; glumae aequales, 2.5-2,6 mm, mg flores aequantes, ovate-lanceolatae, glabrae, 3-nerves; lemma inferius 2,1-2,2 m, lo re vix oblongo-lanceolatun, cenbacetams Sparse pubescens, 3-nerve; palea lemmate paulo brevior, carinis minute cil viata; pachitian internodium 0,3 mm, longum; antherae 0,5-0.6 mm. longae. Perennial; culms tufted, erect, glabrous, 6-9 cm, tall; sheaths glabrous, menbranaceous at the base, the lower ae rter than the upper, the uppermost sometimes readhing the vr of the panicle; 1 membranaceous, the uppermost -5 blades 1.5-3.5 cm. long, folded or somewhat flat, ponies ne Ss 2 ascending, obtuse, tely scabrous above, glabrous, beneath; panicle 3-l cm, long, 0.8-1.5 cm. wide, rather dense, the branches ascending or nearly appressed, some of them naked mm » 3J-nerved; lower lemma 2.1-2,2 mm. long, mo ees Sed subacute, sparsely pubescent, 3-nerved, the ~hay close to the margin; palea a little shorter than the emma, more or less elliptic, truncate at the tip, minutely ciliate on the keels; rachilla joint 0.3 mm, long; anthers 0,6-0.8 mm, long, Type in ake U. S. National Herbarium, No. 2180752, col- a +e pee Peru, May 5, 1957, by Rauh & Hirsc 0. P= Distribution: Known only from the type locality. CERTAMEN MELASTOMATACEIS IX. 964 27 Jo . Dept. of Botany, U.S. National Museum Except for several problems in species groups for which types or type-n at are currently not available, PS eierer en of the Depto. Amazonas Melastomataceae collected in 1962 been completed; the “field work was assisted by a grant toy the ma Miranda are c cnaman to furnish material a sica-sone te ric from somewhat further south in Peru, mostly in Still lacking for dufinitivn of the southern od ey er jalca adequate recent materials from the eastern cordillera in wi (but not the usual North American tyros) who can both exploration and return with aiken specime Also included herewith are initial comments on “the extensive dairies: yer ctions made in recent years by various Regnel- lidian expeditions; eg eet tkoal materials have been made available for is by D r and Naturhistoriska Blick. sda ,» Stockholm, through the intermediacy rr! Dr. eee Where appropriate, notes on Bg collections of Garden, Additional remarks on the extensive extraneous oe men collections will continue in the future. And hi Amazonas and indeed the 2 meee had oriented long-range plans for field work toward this region since 1950. Peruvians living n Amazonas use the vernacular name "Sarcilleja" ("ear-pendants" in Quechua, referring to the pendulous flowers) for all species 377 378 PHYTOLOGIA Vol. 11, no. 6 of Brachyotum found there; the same vernacular name is also used for species of Fuchsia. A local remedy in Molinopampa for pneumonia is a pe infusion from any of several species, but especially from B. weberbaveri Cogn. Seeds of several species Wi Beltsville B. multinervium, B. radula, B. cogniauxii, and (from the Colombian Paramo de Chisaca) B. {gre ram; of these, B. radula flowered in March, — in about one year from seed. Til of the numerous seedlings from each seed lot have been quite uniform aevened patent e effective pollinators of Brachyotum species in Amazona are aeinktarde and the genus must be the most peony ea food source for these birds, of more significance there by s abundance than Fuchsia, Tacciniacese, Labiatae, or iiealaanens it] oO a @ o o fom te 3 =) 4 be g a @ ‘3 be | ec ° < o & Dr. Alexander Wetmore (personal seomppg iescon say t hummingbirds may prefer pos flowe but are not © ekrestiwnly color-specific in feeding. Grant -hvetuelon 3: 82-97. 199) also noted that hummingbirds are not particularly ryan . field work, Segundo Sanchez Barrantes, had a discriminating eye for subtle morphologic variants; thus there rapidly accumlated species taxonomist could easily voaereke binomials; fo practicality has induced discursive treatment without * nomencla- tural formality. BRACHYOTUM QUINQUENERVE (R. & P.) Triana var. PUSILLUM ctor ny The northern variety is quite common in the jalca zone fe) white pe margined pink was once seen near Molinopampa (Wurdack 101), but Showed no vegetative evidence of gene infiltration from other species wae PARVIFOLIUM Cogn. One of the commonest unders shrubs in the Molinopampa scrub forest (Wurdack 135i P-Pey 1619), B. parvifolium reaches its flowering peak in the latter part of May or early June. The robustness criterion originally cited (Mem. N.Y. Bot. Gard. 8: 367. 1953) for gob onued rs B. barbeyanum is not valid, but the petal color and ovarial h r differences Still seem applicable. 4 One hybrid with the si ito var. illum collected in the Molinopampa loa ck 1354 p.p.)- ee bagi ae this collection is somewhat closer " enerve llum; reproductively, except for the l-merous flowers erately short-st thia, it resembles # folium (greenish-white corollas, large and adaxially strigose Sepals). One of the abundant understory shrubs on the lower 1965 Wurdack, Certamen Melastomataceis 379 eastern Calla-Calla slopes resembles a robust B. parvifoliun, wi k population merits specific recognition (no possible fe gat for hybrid origin being noted in the Calla-Calla ra i observations are needed. The ines sn shrubs nigh ceably 2 Poa abundantly in June, being mostly in young fruit on July 7 BRACHYOTUM RADULA Triana B, radula is common on Puma-urcu and Calla-Calla and quite constant there in 5-mery and pubescent petals; the closely related ck 143 Somewhat pubescent petals; can a slight infiltrat‘on of 3B. radula genes is represented in this collection. ene CRONE (sahara, tics comb. nov Pler coronatum nn, Soc. Bot. 28: hi. -~ a mak na See gic agrley Mart. Fl. Bras. lh, 3: sis es prechyotun racemosum Cogn., Jahrb. 42: 132. 1908. of the holotype hauaen 3211, as in con with recent pti pe pol ipevicne. ( ac and nee 799), leaves no doubt as to the generic adjustment. fhe corolla” e =a ope a 370. 1953) show somewhat longer upper leaf current collections, but the variability does not oot: ep B. coronatum ‘e an abundant shrub or small tree on = Tr ers ha een seen on the s. From the description and type photograph (Macbride 16807), Tibouchina virescens Cogn. ts sho n perhape as conspecific with B. coronatum; the gene deltentne be settled, however, without actual tibet me Trent colle BRACHYOTUM MULTINERVIUM Wurdack, sp. n De speciebus 20-28 revisionis es “Gittert foliis 7-ll- nervatis, Trichomata laevia. Ramuli novelli obtuse bat gre. oa demm teretes sicut petioli foliorum venae pri e Pedunculique sparse strigulosi pilis 0.4-0.8 mm longis healed 30 PHITOUOGIA Vol. 11, no. 6 arcte appressis, Petioli 2-) mm; lamina (2-)3- X 1.5-3 cm, temiter coriacea ovata, apice late hebeti-acuto basi paulo (3-6 mm) cordata margine integro et appresso-ciliolato pilis ca. 1 mm aa et 1 mm inter se distantibus, 7-9(-11)-nervata nervis cundarii supra invisis subtus paulo obscuris et 3-l, mm inter se distantibus tertiariis non vel (subtus) obscure evolutis, supra et subtu nervis primariis marginibusque exceptis glabra. Inflorescentia terminalis vel in ramis lateralibus terminalis; flores h-meri penduli terni vel in dichasiis tribus terni; pedunculus gracilis 2.5=-3.5 cm longus ad apicem bracteis duabus 9-10 X 1-2 mm acutis caducis armatus; pedicelli penduli 10-15(-25) mm longi 2-5(-10) mm supra basim articulati et bibracteolati bracteolis ay 5 X 0.3-0.4 mm mox (in alabastris) sande se Hypanthium (ad rum) 7=7.5 mm Longun sicut pedicelli ree: extus sparse Sinu acuto. Pet oe sparsis 0,1-0.25 mm longis eglandulosis vel valde caduce ie glandulosi robustis eflandulosis 0.2-0.6 mm longis, lobis apicalibus 0.6~0.9 mm longis. Type Collection: J. J. Wurdack 130k comet US 2h0h282 ; isotype USM; 13 additional isotypes to be distributed), collected ina ree Foss po below Km 15 of the slices Lvtaasuial og middle eastern Calla-Calla slopes, Prov. Chachapoyas, Dept Amazonas, 5 Paes — 3100~3250 m, 11 July 1962. "Shrub 0. whee 5 m, naar frequen a yellow-red; corolla black-purrls. app eons character. in B. mltinervium wi Pia, ge n the - genus; hina leaves ‘ee often red-tinged be young inflorescence features, perhaps B. s Lomantie gon ) Tr. imum t relatives. Wwordack 1263, re B. multinervium as one parent (multinerved leaves; pendul er flowers in dichasia, but with short peduncles). The pone r pare B. aff. parvifolium rien does not seem as likely as B. naudini>» q earn wrath Triana e northe pulation of this ee ea N. Y. Bot. Gard. 8: 383-38. 1953) is quite common both on the astern slopes of Calla-Calla (Wurdack isoestns ra & Acleto 15268, 15275) and on Puma-ureu (Wardack ay the Salient 1965 Wurdack, Certamen Melastomataceis 381 features are }-merous Sa nea purple petals, and pubescence min. Becsacedh dt being larger me in the typical population, the petals pink, hybrid, but also with B. naudinii genes slightly inaanatanteties was sampled in Wurdack 1350; the roughened hairs, tendency to terminal flower triads, slight predominance of 5-mery, an sparsely strigulose petals suggest B. radula as the other parent, Still another hybrid (Wurdack 1349) is almost exactly intermediate toward B, cogniauxii Wur > With slightly roughened hairs, arcuate=strigese branchlets, arb ey eeiirenr of floral bracts, and adaxially strigulose calyx 1 A fourt: fe hebsy (Wurdack 1133) intermediate in foliage, meeeidie roughening, and corolla Color toward B. an a Wurdack (with the perdiere anthers of B. naudinii and the pendulous dichasia of B. stifolium occurs on Puma-urcu ace with populations of its parent memate of the examinable flowers, 6 were 5-merous and 12, h-mer ie till abundant. Macbr., found as near @-Calla as Celendin (Cajamarca), is Seneiy. Suggestive in wipartictel aspect of some of the B. naudini ds aga ca BRACHYOTUM osc ms Wurdack, sp. nov. B. strigoso (L. f.) Tr. affinis, sed foliis ee fober: culato-stri gulosis, calycis lobis lanceolato-oblongis oesonaseg marginibus valde recurvatis, trinervata nervis later, non visis, supra in lineis 6-B(-10) dense ae a ulosa, subtus dense strigulosa. Flores 5-meri nutantes ramulis labettald bai singuli vel in dichasiis 3-floris; pedicelli 5-7 mm mm longi, bracteolis duabus plus minusve persistentibus 7-13 X 1-2 mm intus glabris extus modice strigulosis, 2-5 infra hypanthii basim insertis. d torum) 7-9 X mm; calycis tubus 0,.8-1 mm altus, lobis 11-13 X 3.5-h.5 mm extus sicut dense strigulosis vel centralite brevistrigosis intus glabris vel ad apicem ipsem sparse Strigulosis. Peta vide gro- rea 17.5-1 la ut tur Uy-15 mm elliptica apice rotundato et vix retuso ci glanduliferis glandulis mox caducis. Stamina (vix Saatrs) 382 PHYTOLOGIA Vol. 11, no. 6 glabra, antheris 3.5-3.7 mm apicaliter minute porosis, connectivo ventraliter ad basim Bonpentasntete, appendicibus ab thecis 0.9- fa liberis. Stylus 27 X 0.5-0.6 mm glaber; stigma punctiforme; ovarium ad apicem dense strigulosum ilis p. p. glanduliferis. Type Collection: A. Lépez Miranda & A. Sagastegui 3338 (holotype US 2338770; isotype TRP), collected in the jalca on the Travesia Laplap between Longotea and Bolivar, Prov. Bolivar, Depto. La saver VAs Peru, elev. 3600 m, 2 June 1960. "“Arbusto. Flores vinosas ve a Paratype: Z bs A. Sag4stegui 3511 (NY, TRP), from the de ayno-htne 0, Hu ajafltas, Hv. Pataz, Depto. La Libertad, Pom lev. 3350 - 23 May 1 wee has moch finer upper i surface pubescence, Dp r larger 5-nerved leaves) is B. benthamianum Tr., although the floral bracteoles in t Tuvian species never assume the bracteal proportions found in the Srcadosian congene BRACHYOTUM MULTITUBERCULATUM Wurdack The species is abundant along the Molinopampa-Diosan trail on the south side of the height-of-land (Wurdack 1606); it also oc pa. seerent hybrid with B. barbeyanum (Wurdack 1667) was found o oe ae AD sen sess it is rather eToser ¢ " B. ae eyanum and Si not pored anthers. The th h 7 preclude omuikesiin of any of the Seandant eolnge sm epesits near Molinopampa. The other wide-pored species of Amazonas, B. lyco oides Triana, was abundant in the cumbre of Cerro Melee al, ere no other congener occurred; the petals of this vegetatively bizarre shrub are lavender (Wurdack 223), a color not duplicated in other Amazonas species BRACHYOTUM ROSTRATUM (Naud.) Trian B. rostratum is a frequently seeds species oe the Calla- erest (Wurdack 1190). On the approaches to Cerro Campanario was seen a hybrid (Wurdack 1590) having the general vegeta tative features of B. rostratum (and pink petals), but sepals longer thm usual and adaxially strigulose, as well as calcarate anthers; perhaps the other parent was B. cogniauxii Wurdack, On ppe Slopes of » there is a able population characterized by leaf blades more-or-less Saternedsi te between B. rosteete and 3. stifolium (vide infra), nearly smooth trichomes ota n g. ar-setose hypanthia ough the pte Sa parent Species were not collected on -ureu, I still believe in hybrid origin of Wurdack 1697 and 786, the former vegetatively more like 3 3 and the latter like B. rostratum. The s e B. seor os ase . the combination of steheuleas peduncles and et e pedicels, but do r-s not show the marked leaf veinlet data. (lower qustece } of the Ecuadorian population. Dr. Ferreyra has collected ne 1965 Wurdack, Certamen Melastomataceis 383 Celendin (15042, 15126) ae | aspect of this complex, with moderately roughened trichomes, leaves as in typical B. rostratum, and inflorescences as in B. seorsum; accordingly, B. se seorsum cannot be maintained as specifically separable from B. rostratum. BRACHYOTUM ANGUSTIFOLIUM Wurdack locality is the Chach rene region; the dee is “ t on the nena Fok crest (Wurdack 1697, Ferreyra & Acleto 15330). On Puma-ureu above Chac = = yas, there "Lar eity as been hybridization with both B. maximowiczii mince Ly a the leaf blades and inflorescences in mediate, the petals pink (Wurdack 67h). sng MEXTAE Wurdack, sp. nov. M. hexamerae Sprague affinis, sed foliis ovato-lanceolatis fere Se peed ge si cordata floribus minoribus. i teretes graciles sicut folia ne ee tee floresque a gla el plerumqu Trees ad 1:2.5); petioli 0.3-0.7 em; ridin (5-)9- Pre: 0)x (2.5-)h-7(-8) em tenuiter nae integra o emer apice hebeti-acuto, basi O.-1.1 cm cordata, oe secundariis ws mm inter se dista ntibus rulis supra obscure vel paulo elevatis subtus paulo clevato-reticulatis, 2 subtus Sparse brunneo- ve gro-punctata, Panicula 22-27 cm longa laxe mltiflora; flores 6(-7)-me meri, pedicellis 1-2.5 cm longis plerum- que paulo supra medium articulatis, bracteolis delapsis non visis. 11-16 mm glabra non ciliata oblongo-obovata apice asymmetrice rotundata. Stamina isomorphica glabra; filamenta 8-10 mm longa ; antherarum thecae 7,5-1 subulatae ad apicem dorsaliter minute uni ae, connectivo in calcar acutum mm infra filamenti insertionem protracto dorsaliter per 3.h-).3 mm asi elevato, a ce dorsali asc nte non evoluta. Stylus 14-16 x 1-0.6 ber; ovarium 6(-7)-loculare glabrum Ose Type C ction: Ynes Mexia 6826 eu pired 8 US 1663013; isotype NY), “cal lected along river bank i e forest between Bafios and a, Prov. Napo-Pastaza, ays tale 00-500 m, 23 Feb, 1935. "Shrub 2.5 m, infrequent. Flowers dark cerise." elev. 1300 m, 9 Jan 1962, "Vine; sepals very dark purple; petals purplish red; filaments purple; anthers aceite Pearce }21(K), from "east of the Andes, 5000 ft."; Penland & Summers 155 (NY), from Ro Margar ites, ua, 12 m, 19 es ? o Margarjitas, Tungurah ind Fagerlind & Wibom hl from Rio Negr rahua; — 937 m Rio Zufiag, Prov. cpipesraetass, elev. m, 1h Feb, 1956. 17 observable flowers, 15 were 6-merous and ?-merous. M. hexamera has wexicpatiaied basally =e ty leaves, Ay 38h PHI710.140 OLA Vol. 11, no. 6 as well as hypanthium plus calyx 10-12 mm long. In Cogniaux' oo M. mexiae would key to M. rigida (Benth.) Tr., with smaller leaf blades that are distinctly petioled and apically rounded, ae -merous flowers, and evident (albeit very short) thick blunt external calyx teeth. Mexia 6826 was distributed as aay neee eooe eee Tr., which however has quadri-alate Lrtnctine, Tine ades, and an obvious dorsal bulb on the stame Asin enthg “unfort rtunately Rimbach 211 (NY, US), the aba i 9 been . hexamera is scmeuncda “+ ee Mokee with M. mexiae: Asplund 15378 (), Cashurco-Rfo Zufiag, elev. 1200 m; Lugo 55 (S), Mera. An apparent hybrid between the two species, with c leaf bases, well-defined upper leaf surface veinlet reticulation, hypanthium plus calyx 8-10 mm long, is represented by Penland © Topo, Tungurah 1939; the M. hexamera genes seem slightly predominant. ’\LEANDRA PELTATA Wurdack, sp. n De affinitate mihi fad fs sed ob folia peltata bene distincta, i teretes primum sicut petioli foliaque subtus modice setosi, pilis 1-2 mm longis plerumque non glanduliferis demum glabri. Folia per paria in dimensionibus plerumque satis (1: 2) isparil j m i floris; flores 5-meri, pedicellis 1-5 mm longis glabris ca. 0.5 mm infra hypanthii basim. bibracteolatis, bracteolis 0.)-0.7 mm long1s Subulatis. Hypanthium (ad torum) 1.4 mm longum modice ie ye 0.2 mm altus, lobis interioribus 0.5-0.6 mm altis ovatoninlenee® apice retnnaate et pilis 2-3 glanduliferis 0.3-0.5 mm lo erminato, dentibus pagel yu ca. O longis sacra ie non ne bus apice setulife Petala glabra 3.1 X 1. asymmetr ovata apice hebeti-acuto, Stamina isomorphica glabra; Mike a l- nome antherarum thecae 1.6 mm longae oblongae apice orae uniporoso, ef jemi ad — dorsa~ aber dente hebeti bilobulate 0.15 mm longo ornato, Stylus |1.1 7 0.4-0.25 mm glaber; stigma paulo expansum 0.45 mm diam; ovarium Seloculare 1/k-inferum glabrum apice in collum 0.4 mm altum et 0.1 mm 5-lobulatum protracto; bacca piopens non costata; semina paulo immatura cuneata laevia 0.6 X Type Collection: J. J. Wurdack 2u7h (holotype us 2404885 isotype USM; 2 additional isotypes to be distributed), collected 1965 Wurdack, Certamen Melastomataceis 385 on a forested ridge on right bank of Rio Santiago 3- km ahem mouth, Prov. Bagua, Depto. Amazonas, Peru, elev. 300-350 29 Oct. 1962. "Shrub 2 m. a ile L. peltata has somewhat the phon: of the 4-5-merou species of Clidemia cane diffuse inklomseoentes and soi Pe pitt fus (Cc. Saw woe Macbr. lies - Sm. semi Gleas, set hee and indeed a some = these species show near er uence to peltate leaves, the floral details camnot be reconciled sufficiently for placement in Clidemia. ni can this Santiago novelty be accomodated near any Species of Ossaea, Admittedly the petals are bluntly acute, but are less rounded than those of the Mexican L. cornoides (S. & C.) Cogn. In per pean peg nographic arrangement, L. peltata should peabehly be ed in Sect. Chaetodon where at least L. cornoides, L aligne sactigsts (Tr.) Cogn., and L. refracta Cogn. also have S=celled MICONIA CRASSIPES Triana Obviously the Peruvian type locality, not known to Cogniaux or Macbride, is the agg region, where this species is common in sheltered ar Recent collections in Depto. Amazonas, all from 22 00-2500 m ee ty are m Quebrada ne seit Chachapoyas (Wurdack 617), near Set (Fe 15548 15593), and the Laguna Pomacocha basin (Fe rreyra & iclete 1517h, k 827). MICONIA ALBORUFESCENS Naud. M. arirambae Huber, Bull. Soc. Bot. Gen. ser. 2, 6: 192. 1915. M. ceeee is Brade, Arq. Jard. Bot. Rio 16: Uh. 1959. en owering material from the Rio Doce, Minas Gerais, Brazil (Paiet 7113, US) has enabled the completion of the synonymy previously suggested (Mem. N. Y. Bot. Gard 5:1 =] Oo ce . Brazil to Venezuela. While there is a superficial resemblance to M. albicans (Sw.) Tr. (and nate ciesiint 3309 ca cited by Cogniaux as a mixture of the two spec » M. M. alborufescens can be distinguished by the agglonerated remy owers the discrete (stellulate) tomentum the young stems and inflores- cences, Through Dr. L. B. Satth ‘and Dr. Pabst, it has been Eepettie to borrow for examination the holotype (Pereira 1779) Paratype (Pereira a2) “ Tye “cachinbensis Onshtiene viants conrorm eX: material from other parts of the range of M. albo ir aieriipd MICONIA ERNSTII Wurdack, sp. nov prs Glossocentrum, M. tetra onae Cogn. et r’ rturbatae Wurdack in antherarum forma s, sed foliis ad basim paulo cordatis. acheiuesetine floribus seg alts in ni mart Sa ramis secundi Ramuli pri mam paulo sulcati mox teretes sicut petioli foliorum venae oh ade subtus inflorescentiae hypanthiaque 386 PHYTOLOGIA Vol. 11, no. 6 primm Sad ape ferrugineis minutissimis sparse obsiti mo. aber rati.: Petioli 0. rae 6 em; lamina tenuiter coriacea ‘shai supra nitidula 6-1) X 3-5 em oblongo-elliptica apice subabrupte -5-1.5 cm acuminato basi 0.2-0.5 em cordata, 3- vel obscure S-nervata nervi 0.5-1 em inter se dist. non vel obscure evolutis, supra et subtus in superficie venulisque glabr “0 ariis mox glabrata. Pa T- X 3-5 em oblonga, ulo adscendentibus secundifloris; flores 5-meri sasdctae bracteolis 0. 2-0. 3 X 0.2 mm triangularibus in alabastris valde caducis. Hypanthium (ad rece 2.2 X 1.5 mm bscure 10-costatum demum glabratum; calycis tubus 0.3 mm altus, lobis interioribus 0.2 mm altis ad apicem obtusis ad basim remotis, dentibus exteriori minutissimis obscure elevatis eminentibus. Petala glabra 2-2.2 X 1.6 mm grpchiesoutangs apice lo emarginato, Stamina paulo dimorphica glabra, filamentis 1.8-1.9 mm longis, antheris ventraliter minute (0.2 mm) uniporosis. s maiora: antherarum thecae 1 X 0.55 X 0.6 mm, connectivo usque ad filamenti insertionem 0.8 mm prolongato dorsaliter _ insertionem dente hebeti 0.5 mm longo armato. Stamina minora antherarum thecae 0.7 X 0.5 X 0.6 mm, connectivo usque ad file i apice sede conico 0. 3 mm alto sparse Sous dacsiel pube ebay ex: fructus immaturus paulo 10-costatus. Type Collection: W. R. Ernst 1723 (holotype US 236501; 9 isotypes to be distributed), collected on the south slope of Morne Macaque (Micotrin) on the road to Fresh Water Lake, Dominica, British West Indies, elev. ca. 760 m, 26 June 19h. “Shrub a tall. Flowers white to pale pink. single plant observed." aratype: a H. pad 513 (US), from the upper rainforest- clad slopes, minica, elev. 763- Both the vigutea eecateead have well-petioled non-cordate eter as elt e as a ar oe and smaller cgocyeton Jap monograph, M. ernstii would key t pthoidos a \% Nend.5 > that Brazilian speed, » eopeesally as ne var. cordifolia Cogn., vegetatively is reminiscent of bee Dominica congener, but shows pedicellate flowers ok line anthers, None of the West Indian species of Sect. Glossocentrum more recently described Urban (M. bisulcata Urb. Urb., and M. sdacettitia Ge & a “seen to be at aTl closely related to M. ernstil. M. trichotoma (De pc known from D differs in the larger well-petioled leaves and -secund flowers with shorter obliquely ge-pored anthers; from the anther fo 3 es would perhaps be better placed in Sect. Chaenanthera. Ss ia, there is a distr ted in herbaria as M. trichotoma, which suggests M. rnstii in reproductive features (secund flowers, nell-prolonged connective); however the St. Iucia plants show much larger 1965 Wurdack, Certamen Melastomataceis 387 well-petioled leaves with developed laxly reticulate tertiary veins; probably an undescribed species overlooked by earlier students of the Melastomataceae is represented, M. martipicens's Cogn, Speers has somewhat the aspect of M. ernstii, differs at least in the apically rounded leaf blades; only eae and fruiting collects ions nes 672, Hahn 1174) of this —97 placed by Cogniaux in Sect. Cremanium > have been seen by me MICONIA LOREYOIDES Triana 1li ipii Gleason, Bull. Torrey Club 52: 387. 1925, te Triana's statements, direct and implied, abou glabrous filaments and style in Triana 1,029, Macbride 's excellent Photograph (25982) agrees in all vi sible details with the various collections of M. killipii; pew significant in the photo- ~~ are the tendency to incrassate stem nodes, the lobed calyx, and anther and stigma niet Also there is a Triana Chocé Sees ine at Kew by N. L. Britton under M. jors oides’ 3 the Triana specimen, which i is perhaps a misnumbere syntype, also va BM, New York photograph N.S.5610) annotated by Trian ent Ecuadorian collection of M. loreyoides a As eae 167h¢ ae hei between El Volante and the a a, elev 1700 m t is especially unfortunate that aasamta commemorative epithet should disappear into synonymy. MICONIA GONIOSTIGMA Triana M. megasti Gleason, Bull. Torrey Club 68: 250. ity he @ pubescent and somewhat glabrescent forms are present on — sides of the —_ in Colombia and Ecuador. The following specimens with a reasonably compact spectrum of pubes- ae ave be Co . El Valle, Killip & Garcia 33936, Killip & in cnt 38770; ARE eg 2 Re sas ohh. = Ecuador, Esmeraldas, Mexia Ob1h, O53; Mexia 6687; po-Pastaza, Skateh 1558, Seeaett et. & Wibom URS Asplund 9529(8), 425(5). The population around Mera, Napo-Pas vestase (Hari Asplund 18431, Asplund 18696) is a quite glabrescent nods in M. Satoet Igma, with setulose petioles, eeresnedeos 14, ate leaf ee parsely setulose benea Sparsely setulose gb branches, and eset tulose. rezoned leaf blades in M. ostigma are 7-nerved — the racts and bracteoles rather persistent; in M. lorey » af blades are d e bracts and bracteoles early- le th Caducous in bud. The closest approach to M. loreyoides from 388 PHYTOLOGIA Vol. 11, no. 6 M. gontostl pes is represented by Camp E-609 (NY), from dense Tainforest in the Moro-Moro region km west of Portovelo, El Oro, Ecuador; this collection shows 7-nerved caducously ciliolate leaf blades which are very caducously strigulose, rather persistent map cence, or hypanthia. M. pichinchensis Benth. has the general aspe = f the men goniostigma; however, the anthers are proportionately narrow and exappendiculate, the styles are ay ae and the stigmas ale only minutely expanded (less than 1 mm diam MICONIA FLORIBUNDA (Bonpl.) DC. 1057, from montane rainforest along the Yambrasbamba- tween Yambrasbamba and Yanayacu 00 i mp M : Macbr. and M. Lasiostyla Gleas., both lack internal hypanthial ribs, as well as showing minor specific vegetative and floral differences. MICONIA CLIVORUM Wurdack, sp. nov. Sect. seer mena M. hirtae Cogn. et (ex descr. et photic.) M. pe aud.) Tr, affinis, sed floribus breviter pedicellatis stigmatibus expansis. Ramili p f e ei btus iincowmeentia recited dina dense pilis stellulatis vel pinoideis 0.1-0.2 mm longis obsiti et modice (vel in inflorescentia hypanthiisque modice vel sparse) ovata apice acuto vel paulo (1-2 cm) ee hebeti-acumina basi truncata vel obscure (ad 7 mm) cordata, 7- vel sunapenerraa nervis se is 3-5 mm ae se aheott teis supra pa obscuris nervulis obscuris sed densiuscule reticulatis, wap modice bullulato-setulosa bullis 0.3-0.) mm altis setis 0.5- 0.8 mm longis, subtus in nervulis dense setulosa. Panicula 8-12 cm longa oblongo-pyramidata multiflora; flores 5-meri 1-1.3 mm pedicellati, bracteolis 0.5-0.7 X 0.1-0.2 mm ad pedicellorum bases insertis ante anthesim valde caducis. Hypanthium (ad torum) 2.2 X 2,3 mm; calycis tubus 0.9 mm altus, lobis interioribus 0.5- 0.6 mm altis paulo pellucidis apice rotundato, dentibus exterior- ibus tuberculiformibus non eminentibus. Petala 2-2.3 X1.7- 2 ™ poro 0,25 mm re d ntraliter ine Linato, "iadiaaaw non prolongato e exappendi¢ ulato. Stylus ).5-h.8 X 0.4-0.5 mm basim versus pilis debilibus 1965 Wurdack, Certamen Melastomataceis 389 eglandulosis 0.1 mm longis sparse obsitus; stigma paulo expansum 0.7 mm diam.; ovarium (3- sve ine 1/3-1/2 inferum a conico et in collum 0.6-0.7 mm altum ca. 0.) mm h-lobulatum stylum aes sparse cemcttaiivean stellulato pe ceo. ollection: J. J. Wurdack 179) (holotype US 20430; isotype USM; 9 ie eye isotypes to be distributed), collected on the upper slopes of Puma-urcu just north of Levanto, Prov. Chachapoyas, Depto. part hi Peru, elev. 2700-2800 m, 29 Aug. 1962. "§ m, occasional; cauline hairs red-tinged. Corolla white." Paratype: Wurdack eee sees the middle slopes of Cerro Yama-uma (Cerro Car&n) above get — eich of Molinopampa, Prov. “Chachapovas, sat Peru 2700-3000 m, 11 A 1962, "Shrub 3-7 ™ sep oe but aes cng ae at this Stes tals whi ch. sparser upper Leaf surface pubescence, gland-tipped Ih i ot oe airs. The other species of Sect. woo warrhena in this alliance (spp. 392-398 of Cogniaux' costal ch also llulate differ i most have s pubescence, all obvious vegetative and reproductive features. In ' aes M. clivorum would probably run to spp. 384-385 or 389-391; most of these species lack stellulate pubescence and/or r pedicels, Sessile flowers and completely inferior ova th depressed apices; also insofar as sonescn 835 (US) is sieoaarn the simple Cauline hairs are much rter than in M. c ments and les are bee glandular-pu . Incidentally, scab And Strigulose style and prominently setulose ovary apex. Cogn., a species not represented recent collections, vitor in been 8 ani ned, osely related to M. clivorun, ‘ange the Ps leaf wartars es: tipped with short thick hairs and inflorescence with only appressed — densely yaaa late. ites, and the flowers considerably large MICONIA BRACHYANTHERA Triana Hiurdack k 1058 (montane rainforest along the Yambrasbamba- Pomacocha tra rov. Bongar&, Depto, Amazonas, Peru, elev. 2200-2300 » m, 26 Jun. 1962) has been compared minutely with the 390 PHY7T01L0 GLIA Vol. 11, no. 6 holotype (Mathews 1726, K) and agrees well in all details. The leaf blades are scarcely plinerved, the inner pair of primaries ication. M. brittonii Cogn. a extremely close to M. brachyanthera having the same minute glands on the inflores- cence wer leaf surface hairs, but differs in the moderately roughened sodeher than sparsely and obscurely barbellate) cauline hairs and strigulose style; the expanded stigma is perhaps of no specific import if heterostyly exists in this complex. er FOSBERGII Wurdack, sp. nov. - multuplinerviae Cogn. e et M. penicillatae Gleas. in antheFara forma affinis, sed follis supra planis. Rami i 4-8 cm; lamina 15-25 X 9-1 cm vel ultra chartacea elliptica apice acuto basi obtusa margine serrulato dentibus ca. 2 mm inter se distantibus 0.5 mm altis et ciliclato pilis 0.5-0.7 mm longis incurvo-erectis, 7-9-plinervata nervis subalternatim 0.3-3 cm supra basim insertis nervis secundariis ];-8 mm inter se distant- ibus tertiariis laxe reticulatis, supra primum densiuscule stellato-puberula et sparse pilis robustis 0.5-2 mm longis ee luteis simplicibus obsita demum glabrata, subtus in nervulisque modice et in superficie sparse stellato- acted, Panicula 10-15 cm longa pyramidata psabreteinsee de ramis ramilisque sparse pilis erectis 0.5-1 mm longis ad apice penicillato-stellatis obsitis sicut hypanthium modice stexietse puberula; flores 5-meri ad ramuli extremitates vel in ramlis capitato-aggregati; pedicelli 0.5-0.8 mm longi ad basim bi- bracteolati, bracteolis 0.6 X 0.1-0.2 mm acutis ad anthesim plusminusve ca cis. Hypanthium 1.8 X 1.8 mm; torus ciliolatus pilis 0.1 mm mn longis glanduliferis; calycis tubus 0.2 mm altus, lobis interioribus 0.3 mm altis ad basim remotiusculis 3 apice 1.7 versus curvata srcitnite (0.2-0.3 mm) emarginatae introrse biporosae. Style 3 x 0.2-0.3 mm glaber; stigma capitellatum 0.5 mm diam.; um 3-loculare 1/3-inferum apice conico 0.5 mm alto sparse elantalcso-setuloes Type Collection: F. R. Fosberg & M. A. Giler ae pagent type US 21h3072, iso ay pe USTp selin 5 collected in wet forest on a steep slope near Monos, + km W.n.w. of Zaruma, Zaruma Basin, ” prov. Ore, Ecuador, elev. 1230-1280 m, 11 Feb. 19h5. "shrub 2 m tall; Leaves glaucous beneath. Flowers white." Both suggested relatives have bullate-setose upper leaf surfaces and non-scutate nodes; all three species have stipitate- 1965 Wurdack, Certamen Melastomataceis 391 stellate cauline pubescence (although the hairs in M. fosbergii are more robust) and apically curved anthers with deeply e i= wit e possible exception ap “ rivetii -& in floral details, especially the anthers ; that eames ian Species, known to me by the cae photograph and several recent incomplete collections, ounnoee at least in the very fine (almost rm, formless) lower leaf surface pubescence and non-scutate nodes. In the search for ee — M. sosberes Sections Chaenopleura Cremanium, and _Susenanther ra “alSo canvassed because of the distin Ctive but secti onally Snconelueive anthers. M. polyneura Triana, M. i Macbr., M. = Comnes M. YV. a Cogn., and (ex descr . rhonhofiae | ave a few vege- tative characters in common with M. beara aie but differ in floral feature MICONIA PARADISICA Wurdack, sp. n Sect. Cremanium. M. verulentae R. & P. affinis, sed foliis 3- vel sub-5-nervatis N. polveruientas p, & P. affinis armatis alabastrorum hypanthiis glabris. Ramali primum obtusi-quadrangulati demum teretes s petioli inflorescentiae ramique primum modice pilis stellulatis 0.1-0.2 mm altis obsiti tarde glabrati; petioli 1-2 cm; lamina 7-13 X 3-6 cm chartacea elliptica vel paulo obovato-elliptica apice anguste hebeti-acuto basi late obtusa vel paulo rotundata 0.h-0.5 mm latis, supra p sparse hepa oe (costa excepta) Sree subtus in nervis Stellulato-pube t+ pilis barbellatis vel stipitatowetellatis rn. lo nervulis pilis stella 0.L-0.5 mm stipitatis modice obsita in superficie vera glabra. Panicula 7-10 em longa pyramidata; flor -meri in ramulis Ph do persistentibus. Hypanthium (ad torum) ~ 3 X 1.4 mm st ie 6 Collection: J. J. Wurdack 1055 (holotype US 20429; isotype USM; 3 additional isotypes to be distributed), collected 392 PHYTOLOGIA Vol. 11, no. 6 at the edge of the montane rainforest along the ae Pomacocha trail (between Yambrasbamba and Yanayacu u), Bongar&é, Depto. Amazonas, Peru, elev. 2200-2300 m; be pa 1962. "Shrub 3-6 m, occasional, Flowers white." lverulenta has distinctly aitaees leaf blades, the lower | sor ace veinlet hairs being sessile or ee bly (to 0.1 mm) stations the densely roughened cauline hai often 0.7 mm long and the young hypanthia are sparsely lebelnunites puberulous. Probably M. pulverulenta is dioecious, the type collection (fide Macoride photo T7IhL and Cogniaux! description) and capitellate stigmas. Wurdack 1055 is male, the aborti ovaries being barely raised above the hypanthial floor; thet compact ora ower leaf surface pubescence, and early-furfuraceous — » and M, galactantha Naud. (with, ex descr. and Mac bride —_ 17166, sharply f-angled stems and much less congested flowers), Both M. tiri Triana and M. coelestis Naud. have much larger flowers and different foliage; Naudin remarked (in his description of the misidentified M. verulenta) that the collection had oo anthers and a recent Hu&nuco sel mbetient (W 7998), agreeing pebre ectly with Macbride 's photograph ees M. tiri, shows anthers of Sect. Amblyarrhena rather than Cremanium. ~~ MICONIA PENNINGTONII Wurdack, sp. ct. Cremanium. M setnaphens: Naud. affinis, sed ra ad nodos non atato Sonminessts ramilorum petiolo orumque pilis s erectis ad apices non penicillatis, floribus plerumque )-meris. Rarmli obtuse quadrisulcati demum teretes sicut petioli pedunculusque modice pilis erectis 1. 5-2(-3) mm longis pote sete cm @ multiflora; flores )(-5)-meri in ramulis conferti pl Lerumque sessiles (interdum ad 1 mm pedicellati) bracteolis 1-1.5 X 0.15- 0.2 mm linearibus furfuraceis mox (ante anthesim) caducis. Hypanthium (ad torum) 2 X 2 mm une sparse stellulato-furfura~ 1965 Wurdack, Certamen Melastomataceis 393 ceum mox glabratum; calycis tubus 0,3 mm altus, lobis ea a ag 0.6 mm altis apice rotundato, dentibus exterioribus callosis eminentibus. Petala (sub lente) minutissime granulosa 1.5-1.6 X 1.2-1.3 mm obovata apice as ce truncato vel rot O. tamina isomorphica glabra; filamenta 2.8-3 mn; rum thecae 1.9-2.1 X 0.45 X 0.6-0.7 mm oblongae vel paulo obovato-oblongae late biporosae, connectivo ad basim ventraliter minute bilobulato dorsaliter imperspicue (0.1 mm) — Stylus ) X Onl mm laber exsertus; eon paulo expansum mm diam.; ovarium 2-loculare 1/3 inferum glabrum apice conico 0.5 mm alto et in collum 0.5 mm — circum stylum protracto. Type Collection: P. C. D. Cazalet & T. D. Penni = eee (holotype US iehosaiee isotype NY), collected in dwarf “fore the Oriente trail E.N.E. of preci Mountain, tn seme, uf Dec. ing. Leaves with _gelden-brown meal below, Corolla greeni on ee ices (s00 Ciattedseas & Jaramillo 12012, US) has notably Errpresser ai ted branch nodes, stem ji irs fray “: at the Pex, margin erst merely c callose. er close relatives ra droree M. ree Naud, (with much shorter rie apically te § llula cellate flowers), M. “ eura Triana * (with repre branchlets and pedicellate flowers), and M. cladonia Gleason (with stem and lower leaf surface hairs branched and with spicules 0.1-0.3 mm long, as well as relatively broader anthers). More remote chee in Sect. Cremanium include M. rophila Naud., M. sD. Sm., M. tuckeri ee: M. ere ficre nag and M, Trerele nta R. & Pl Of the rs counted in M tonii, 1] were eshte si a enact inns S-meror Sus 3 €spite this predominance of l-mery, the relationships do not seem at all to be with patie 1,82-1:85 of Cogniaux! arrangement. MICONIA JENTACULORUM Wurdack, sp. nov In systema Cogniauxii de _speciebus 434-36 differt foliis obtuse cuidsteigutest demmm teretes sicut petioli venilium costae supra inflorescentiaq ilis barbel- latis gracilibus cinereis 0.5-0.7(-1) ° dense obsiti Pilorum parte ap i demum caduca. +t ~ ongi; lamina 39h PHYTOLOGIA Vol. 11, no. 6 centia 2-3 cm longa confertiflora, bracteis bracteolisque 2-3 X 0.3-0.5 mm paulo puberulis conspicuis ad anthesim caducis; flores connectivo ad basim dorsaliter inconspicue (0.1 mm) hebeti- dentato ventraliter 0.25-0.3 mm prolongato et paulo bilobulato. phe X 0.4 mm glaber; stigma paulo clavatum 0.5-0.6 mm diam. ; um 3-loculare paulo inferum apice in collum conicum 0,8-] mm oo minutissime setulosum circum stylum protracto. Collection: J. Cuatrecasas 13530 thaletepe US ange isotype NY), collected In the Paramo del Almorzadero "extrema en Peralonso", Cordillera Oriental, Depto. Santander, ceksentic elev. 3100 my, 28 Nov. 191. "Frutex. Pétalos y estambres blanco~ amarillentos Par ss same locality data, fruiting, 19 Jul. 19h0, Cuatrecasas & Garcia Barriga 9925 (US). While M. jentaculorum has somewhat the vegetative “ee of M. rigens Naud., it does not seem florally closely relate sessile flowers would key the Peralonso endemic to M. segeen (Bonpl.) Naud i Naud., and M. caelata (Bonp ya all of which haves s ( th torus 1-1.3 mm long) and quite different large pubescen per’ ° rophila and also M, elaeoides Naud. are the closest relatives; a latter species ch thi glabres~ chosen for M. jentaculorum refers to the type MICONIA DIELSII Markgraf ex Eh This Ecuadorian Scastas: seems to show baheneanray Ce A female flowering collection (Hitchcock 2182), fro Bafios and Casnurco, Tungurahua) has abortive na an a capitate stigma on a well-exserted style; male material (Skutch 451k, from Puyo, Napo-Pastaza, originally distributed as brevitheca Gleason) shows a sat iedaveloped anthers, but a scarcely exse style with minute expanded stigma. A fruiting collection is As lund rohgo: re from Vera Cruz, Napo-Pastaza. Very minute own Sessile glands are frequent on the hypanthia and inflorescence mage as OB as Rs ah tipping the Spicules of the rather scurfy tiny dendritic inflorescence hairs. nne cent leaves and glabrous smaller hypanthia. The specific epithet p&ramo MICONIA iy ens is sp. a um, M. salicifoliae soaps ) Naud, affinis, sé foliis it or pee anmennas distincte trine is subtus in superficie glabris, calyci pana aa me eminentibus. Ramuli te s rn petioli inflorescentiaque primum dense pilis dense barbellatis 0.1-0.3 mm longis ad basim crassis primum - 1965 Wurdack, Certamen Melastomataceis 395 ravis demm fuscis armati, Petioli 1.5-2 mm; lamina 10-15(-20) x h integra apice acuto basi late acuta, supra glabra, subtus in venis primariis sparse vel modice furfurac n secundariis m sparse furfuracea mox glabrata, trinerva nervis primariis supra conspicue impressis subtus elevatis nervis secundariis supra obscuris subtus evolutis et ca. 0.3-0. inter se distantibus tertiariis non evolutis. Paniculae numerosae racemiformes paulo declinatae 3-7-florae furfuraceae; flores h-meri ca. 2 (demum usque ad 3) mm pedicellati, bracteolis sub- persistentibus 2 X 1 mm ovato-lanceolatis ca. 0. mm infra calycis tubus 0.2 mm altus, lobis interioribus 0.3 m ee tri- angularibus ad basim remotis, dentibus exterioribus 0.1-0 eminentibus ad apicem setiformibus. Petala granulosa 1 x “4 mm oblata apice rotundato. Filamenta 1.3-1.5 mm longa basim versus i aliter 1/2 inferum idan conico QO, 0.5-0. 7 mm ae modice setuloso setulis 0.2-0.3 mm Arey pi Dp. —_ feri Type C ion: . Wurdack "i218 Rete US 204271; isotype USM; 4 saa ont “isotypes © o be distributed), collected 1d + (2 walk south), Prov. Chachapoyas, Depto. Amazonas, Peru, pe Sig er m, 8 Jul. 1962. "Shrub 1m, occasional. Fruit purpli M. saltcifolia has longer leaves which are faintly ci basally and densely tawny-furfuraceous beneath, densely fu aceous thi und infra-marginal external calyx vont id Perhaps a closer relative, at least vegetatively, is an undes- cribed Ecuadorian species (to be aoe on se P-312), which mer pecies S suks Cremanium in Cogniaux' monograph, a vaceinoides (Boop Naud., at least as sanen the type cot and We “4 collections di axer xte pecie Collected on Puma-urcu above Chachapoyas (Wurdack 787) and resembles vegetatively M. ledifolia P(pc.) Waud., but with )-merous fruits similar to those of M, salicifolia. Unfortunately, and not as in M. paludi ena, flower remnants are completely lacking © the Puma-hroa sirab for MICONIA BULLATA (Turez.) Triana M. trichocaula Macbr., Field Mus. Publ. Bot. 4: 189. 1923. 396 PHYTOLOGIA Vol. 11, no. 6 After comparison of Mathews 873 sb with. Weberbauer - Soha there seems no valid separation of the two taxa eae Kew shows abundant simple cauline hairs as rie ones. M. bullata is now also known from rere "(Prieto P-287) and Amazonas, Peru (Wurdack 1705). MICONTA SECUNDIFOLIA Cogn. subsp. MALCABALENSIS Wurdack, subsp. nov. 8 nervatis venis tertiariis subtus planis vel paulo impressis rt. Type C sg geen J. J. Wurdack 1419 (holotype US 2h0299; omrre USM; 5 additional isotypes to be distributed), collected the summit of to Malcabal 1 (Genre Tunbe) 3 3-6 km southwest of itccotenn: Prov. Chachapoyas, Depto. , Peru, elev. 2850- 2900 m, 20 Jul. 1962. “Shrub 2 nm. coretaee, ested anthers yellow." n the typical subspecies, collected by Weberbauer on the snetabe slopes of the eastern cordillera, the petioles are 1.2- 2 cm long and the ovate leaf blades rae X h.5-5.5 cm; the tertiary veins on the lower leaf surface are Jab narne! reticu- late-raised and bees hypanthium plus peeve tube (dry) is 3.2- 3.4 mm long. the Malcabal shrubs, the eestcies are 0,-0.7 cm long, the itis blades 3-.2 X 1.8-2.3 cm, and the hypanthium plus calyx tube (dry) 2.6-2.8 mm long. Despite wre disparity in super- ficial aspect, bsp. malcabalensis seems ° her- elevation exposed-habitat variant of the typical subspecies. Subsp. secundifolia is still kno ° ° from the t col- lection; Macbride's excellent photograph (17159) and Gleason's notes from the Berlin speci c for the s natal level disposition of the M bal population, e aux Slope of Sect. Chaenopleura in the very narrow dehiscence of the anther thecae (extending for about 3/l of the thecal length) and are Somewhat suggestive of Sect. Chaenanthera. TOCOCA SYMPHYANDRA (Triana) Cogn. nee zariogate Mef., Notizbl. Bot. Gart. Berlin 15: 379- ni Poe barbicaulis Gleason, Bull. Torrey Club 68: 2h8. 1 Apr. ISbT- 3 A Gleason's notes on the Kew material (Triana 102, Kalbreyer 1836) _ Triana's floral sketches (K) have confirmed the ddentrt of Haught 522 and 5225 (both from west cf Tambo, maser anne elev. 1100 m, somewhat north of the type locality but 0° 1965 Wurdack, Certamen Melastomataceis 397 western flank of the western cordillera). The following Ecua- dorian collections (all Napo-Pastaza) are the same species as is ee rerrens by Hau mele ve a (US) and agerlind & Wibom 1167 (S), Puyo; Har 9 (S), Canelos; and Harling 3388 (S), Mera, Haught's collections show forms with green (5225) and purple-tinged (522), deep red in vivo fide Haught ) ooh leaf surfaces. Harling 3388 and Fagerlind & Wibom 1167 have green leaves, while the other two Ecuadorian collec- tions are like Haught 522). The upper leaf surface hairs (i. s.) vary from yellow to castaneous. Probably these color forms are analogous to those in Clidemia allardii Wurdack (Phytologia 9: 425. 1964). Markgraf'S excellent description, plus later Ecua- dorian collections, leaves no doubt as to the synonymization of T. variegata; Gleason's notes indicate that Berlin did not have f. hyandra represented in its collections, at least in 1930. ahaa a legalistic purist would care to designate a neotype or T. variegata, but this scarcely seems necessary. As suggested anthers, Adnittediy the anther shape may see impe Placement in Miconi Miconia, with reversion then to Triana's basionym. The white foliar macules noted by Markgraf are especially Prominent in Skutch 4h20, less so and less regular in the other Collections; these pale regions are not shacer enero but eptate f rather, fide Dr. Harold Robinson, colonies of a s ungus. CLIDEMIA DENSIFLORA (Standl.) Gleason Two Ecuadorian collections (Asplund 18620, 19602) from Mera, Napo-Pastaza, agree in all scoontiey details with the six Central erican specimens available for study. The Mera sheets show Somewhat firmer a ot retin ge ot all, of the more northern specime ted in the discussion of C. chocoensis (Phytologia 7: 238. 1960), the ots seems well accom Within Clid demia, but we engeon8 neither in Henriettella nor (even further astray) Henriettea per Bn abby po ae (Gr np i nig "affinis sed ramlis teretibus foliis’ = cisti ie é lps, me supra pustulatis. compres teretes sicut petioli to €~3(-) mm inter se a 7-nervata nervis s 3- mm inter se distantibus tertiariis nervulisque reticulatis, hig distincte pustulata et strigulosa pilis laxis 0,3-1 m longis, subtus in nervulis sparse setulosa pilis gracilibus laxis, 398 PHYTOLOGIA Vol. 11, no. 6 Flores in foliorum axillis congesti numerosi plerumque ]-meri eviter (ca. 0.5 mm) pedicellati, bracteolis ad her enon basim insertis 0.3-0. 5 X 0.2-0.3 mm lanceolatis eas 9 ’ setuloso. Petala glabra 1. X 0.9-1 mm oblonga apice emarginato- truncato, Stamina isomorphica glabra; filamenta 2.2-2.9 mm; antherarum thecae 2-2,3 mm longae ad apicem ventraliter minute (0.1 mm) uniporosae, connectivo 2s anes non Aetea po oe Stylus 6.8 X 0.25-0.3 mm glaber; stigma t xpansum ; pe 2(~-3)-loculare omnino inferum apice oerendet yt mires Pepe Collection: Lorenzo Uribe Uribe 3910 (holotype US 2371598; isotype COL), collected in Cano Garrapato, Tauramena, Casanare, Colombia, elev. 600 m, 2 Dec. 1961. “Arbustillo de 1.5 m de altura, muy ramoso, Flores blancas y eet violaceas." mena, Ca * 1-2 metros de altura. Follaje de un col r verde satinado muy elegante, Flores blancas."; Cuatrecasas 11805 (F, US), from Villavicencio, Meta, elev. 500m; S.C. Smith & J. M. Idrobo 156k (US), from a sandstone outcrop 2 km east of Rio Zanza above junction with Rio Gifejar, north end of Cordillera Macarena tee elev. 500 m. "Herb, woody at base, 0.5 m tall; flowers C. obliqua has + apaehae slightly wing-angled stems, with t+ th leaf blades merely rounded a base and plane e 3 s C. acutifolia Cogn, -bullulate leaf blades and a slightly evolved cymose inflorescence. The other species to me in this complex, C. fissinervia Gleason, i ermediate in so features between the above-mentioned pair that it may well represent a hyb in the species populations arou on the fruit oy: wank ungatiion aster potent pudtes placed in this complex by Cogniaux, C. rariflors (Bonpl.) ¢ — (the name a later homo is known to me the description, but —_ differs from C. uribei (plane leaves; inflorescences. 3-6 en long). Perha cause of the fascic tiolata » C. i would key in Cogniaux' monograph to C. ; c = a Sagot; the Pacis somewhat longer wae" leaf blade Which are pet beneath and have the inner pair of ase, as well as glandular-setulose hypanthia. ¢. micrantha 6s gst 1143, BR) differs in the finer 1965 Wurdack, Certamen Melastomataceis 399 (0.5-0.8 mm deep) leaf margin serrations, plane upper leaf surfaces, and rather densely short-setose (hairs 1.2-1.7 mm long) hypanthia; only one bud is present on the isotype, so plane upper leaf eg age At the moment, type material is not at hand to evaluate R Williams' treatment (Flora of Trinidad & Tobago) of C. debilis yr ) Cogn. to include C. aphanantha and C. bonpl ndii gn. In en S05, Gees 5-merous flowers were seen, but the other six observable flowers in this collection were )-merous re cime en reappraisal for me and o only ethiopia easy of determination. LOREYA UMBELLATA (Gleason) Wurdack, comb. nov Bellucia umbellata Gleason, Bull. Torrey “Club 58: 257. 1931. L. arborescens (Aubl.) Naud. and L. acutifolia Berg ex t re- s Sides of the petals about “Vb 1/3 below the apex, as well as some oreioaeent of sha shak flanges on the petals at this callused rescens is the generotype ; certainly for the FS shoveonentionsl Species, there is no justification for on with Bellucia, Rather, the closest generic affinities of (Rhodora. 65: “ie. 1963), Ecuador” (20 = st of Santa Domingo de los pains Pichincha, Cazalet & Pe be ones Quininde, BLAKEA BRACTEATA Gleas. subsp. nes ye Papert chen subsp. nov. subsp. bracteata foliis subtus in costa basim versus Sparsissime setulosis in venis secundariis se glabris, bracteis exterioribus extus glabris differt. Type Collection: P. J. Grubb, J. R. Lloyd, T. D. Pennington, & T. C. Whitmore 93 (holotype US 2369301), ¢ ee ted at the forest —- at Rigny > 15 km SSW of Tena, Prov. Napo-Pastaza, Ecuador, 9 Jul. 1560. "Liana, flowers ” fl Grubb et al 59, from the same locality, elev. 750 m. 1,00 PHYTOLOGIA Vol. 11, no. 6 ecuadorensis, the young branchlets, petioles, and pedicels are s caducously fine-setose ; the external bracts are very sparsely fine~setose marginally inside. In floral characters, the two subspecies are the same, ech calycine and ein an caducously gland-tipped. The typical subspecies has + nome in Ecuador at Canelos, nr ler (Harting "2208 Mef. must be rather si ea a, tae, ee 2; in ee hispid hypanthia and oo — ee -———ec ef = we ee Se eS Fe MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXVIII Harold N. Moldenke VENEERA URTIG IFOLIA L oe ti bibliogra Moldenke, Phytologia ll: 315, 316, 326-357. 195k. erepky rg pte citations: ILLINOIS: Winnebago Co.: E. W. Fell 5132 (I1~-38585); Fell & Fell F.46531 (1120632); Fuller & Haime 27lfa (I1—~15812), ~253hh (T1--15809). Woodford Co.: Ve He Chase 9991 (Ur, Ur, Ur, Ur); G G. N. Jones 14299 (Ur). county un- determined: Collector undesignated Ds $n. [Boewe}] (I1-—-1580); Va~- sey 3 (I1—16007). INDIANA: Allen Co.: C. C. Deam 1362 (Im Boone Co.: Friesner 16989 (penné2sei)} S. McCoy 5293 3 (Bt--51739)5 A. Re Moldenke 805 (LW), 806 (Lw). Cass iS Co.: Cc. C. “C. Deam 51078 (Dm). “Clark Go.: C. C. Deam 23792 (Dm). Clay Co.: S. Me ~Deam S. n, [C. C. Deam 13868] (Dm). Clinton Co.: Ek sen. [8-27-1911] (Bt—57380). Crawford Co.: Friesner pile ; (Bt—36886). Decatur Co.: C. C, Deam 58272 (N). Elkhart Co.: C. C. Deam 50h29 (Dm). Franklin Co.: Friesner 12418 (Bt—l,21,0); S. McCoy Coy 4172 172 (Dp— 4663). Fulton Co.: Friesner 23098 (Ca-—803863, Ok, S, — me son Co.: Schneck s.n. [July 1888] (Dm). Greene Co.: Friesn 22932 (B, N). Hamilton Co.: Friesner 60h), (Bt—13h56)3 A+ 7 Mol- denke 21912 (Lw). Hancock Co.: Friesner 18618 (N). Harrison Co. Co Co Deam Deam 41512 (Dm); G. Een sen. [16.7.1950] (S)- Hendricks Co. | Pesesoe a 17081 (Bt—-61356). Henry Co.: H. N. Moldenke 2191 (Lw). Howard Co.: C. C. Deam 5823 (N)3 Ek sone [1-2h-1935] (BY 12627), Sen. [7-29-1936] (Bt—L2528); Friesner 1167h (Bt—37H59)« Jasper Co. C. C. Deam 12039 (Dm); Walch 202 (Dp, Ur, Ur). Jeffe son Co.: M.S. Coulter s.n, [Hanover, 1873] (Pu); C. C. Deam wis (Dm). Jennings Co.: C. C. Deam 58280 (N, Ok); S. McCoy L17 ia? (Dp— 4675). Knox Co.: Cc. Ce Deam 51479 (D (Im). Kosciusko Co.: " Friesner 12664 (Bt——l172h). aera: Co.: C. C. Deam 51033 (Dm); Friesne® 15639 (Bt—55058). Lake Co.: Lansing 2806 (W—753090) « Laporte Cos? C. C. Deam 3723 (Du); Frdssner W991 (Bt—2373)3 We He 1965 Moldenke, Monograph of Verbena 401 Rhoades s.n. [near Westville, July 193); H. N. Moldenke 17075) (Br, Bt, N, St). Lawrence Co.: Wible 83 (In—h605); Wynn 90 (Au, Gu——27690).. Madison Co.: Friesner 17107 (Bt—62513); S. McCoy 2236 (Dp-——13911); A. R. Moldenke idenke 803 (Lw (Lw, Ut). Marion Co.: H. H. Bartlett s.n, [July 22, 1903) (i (Mi); Friesner s.n, ete s (Be— 2477); W Week. 1 H. Rhoades s.n. [Indianapolis] (Hs). Marshall ¢c Evermann 686 (W—357535); Scovell & Clark 686 [54] (en9534), Miami Co.: Ek s.n. [8-16-19 36] (Bt—-h2526). | Monroe Co.: Gullion s.n. (Ellettsville] (In—3316); Lewis & Brom s.n. [Bloomington] (xu); Lipps s.n. [July 3, 1941) (0k); Welch & & Price 145 (In). Montgomery | Co.: Friesner 3133 (Bt—-696l). Morgan Co.: Weatherwax Sen. (In—-922). Newton Co.: C. C. Deam 50597 (Dm); M. McKee 1345 (Dp). Noble Co.: C. C. Deam 34615 (Dm); Friesner 1263) (Bt— 41405). Ohio Co.: Friesner 12153 (Bt--l391h). Owen Co.: Fries- ner 4565 (Bt—-12051). Parke Co.: W. H. Duncan 180 (H--7807). Pike Co.: C. C. Deam 52667 (Dm) . Posey oe Ce. Ge Deam oo by Cov: 5. Me Dean sen. [C. C. Deam 13708] (Dm); Friesner 171k (Bt-~-61983) .- aes Go.: C. C. Deam 20885 (Dm), 56416 (H—L3335, ). Sullivan Co.: c.c. Dean 1 18226 (Dm). Tippecanoe Co.: Ek s.n. [10-1-19)2] (Bt 60675. Hip Tipton Co.: C. C. Deam 13621 (Dm Union Co.: C. C. Deam 47283 (Dm), 58171 (N). Wabash Co.: Plies ner 16015 (Bt--53162). Warren Co.: C. C. Deam 49625 (Dm). Wayne Co.: ™M. | "M. B." 155 (Ea); Creek 158 (Ea); Frenning 3 (Ea); Hudson 165 (Ea); Le Je King 1587 (Ea), 1762 (Ea), 2571 (Ea); Knipe s.n. gen. (In 2514). Wells Co.: C. C. Deam sen. [Sept. 13, 1896] (Dm). White Co,: C. C. Deam 58210 (N). County undetermined: Friesner 15826 (Kr). IOWA: Adair Cos: A. R. Moldenke 936 (Lw). “Allamakee Co.: Tolstead s.n. [July 20, 1933] (Ca—-802838), sn. [July 25, 1933] (Io—-1))283). Boone Co.: Macklin & Pammel s.n. wn. [Aug. 1912] (Io-- 52727); Pammel & Henning s.n. (Aug. 31, 1930) (St—9272). Cedar Co.: Fay 1368 (Vi). Clayton Co.: Melhus & Dietz s.n. (McGregor, Aug. 1933] (Io--1)1093); A. R. Moldenke 990 (Iw); L. H. Panmel s. n. (St. Olaf, aug. 10, 192h] (1o—11500h). Decatur Co.: J. P. An derson S.n. [July 43; 1903] (jonsz130). Delaware Co.: Bode s.n. fAug. ae A egelaee Des Moines Co.: L. H. Pammel 13 (Io— 118880) , in part (Io—11955h), sen. [Crapo Park, Sept. 8, 1930] Lon, Emmet Co.: Cratty s.n. [1883] (0050813), Sen. (Armstrong, June 2h, 1897] (1o--15335)- Fayette Co.: Fink 26h (aug. 1, 1894] (W--214890), 26 [Aug. 29, 189L] (W--214890), oshs [August 1, 1894] (Io—31576). Fremont Co.: L. H. Pammel 732 (Io—-119321). Hamilton Co.: L. H. Pammel s.n. (stratford, 6-1- 1913] (Io—77202). Hardin Co.: Pammel & Hagge s.n. [Eldora, Oct. 402 PHYTOLOGIA Vol. 11, no. 6 13, 1928] (Io—-134bh9). Henry Co.: Boley 688 (Se--31,87). Iowa Co.: A. R. Moldenke 976 (Iw). Johnson Co.: es Barker s.n. [Iowa City, 7-27-29] (Au); Huang 21,66 (Lb--54992) . Jones Co: L. He Pammel s.n. (Oxford Junction, mn, July 25, 1919] (Io~-97272). . Lee Co.: Fults 1500 (Io-~138821). Linn Co.: A. R. Moldenke 982 (Lw). Louisa Co.: L. H. Pammel s.n. (Fredonia, | June 1927] “(To--129L77) , s.n, (Fredonia, Aug. 1927] (St--9280). Madison Co.: A. R. Molden- ke 939 (Lw). Mahaska Co.: Augustine 4l2 (Ok, Ok); A. ike ~Moldenke 959 (Ti (Lw). Marion Co.: A. R. Moldenke 955 (Lw). Mills Co.: A. Re Moldenke 920 (Lw). Mitchell Co.: Tittle 31 (Io—-7h179). Musca- tine Co.: Pammel, Kelso, & Harlan s.n. [Jul. 20, 1919] (Io—950h3). Palo Alto Go.: L. H. Pammel s.n. [Lost Island Lake, Jul. 15, 1920] (Io--98810). Polk Co.: L. H. Pammel 412 (Io—32536); Pammel & Forbes s.n. [Bloomfield, Jul. 25, '27] (Io--129942); Pammel Pammel, Fran- kel, & Rieman 986 (Io—119680). Pottawattomie Co.: 0. Me . M. Clark — 6080 (B)3 Ae R. i Moldenke 931 (lw). Poweshiek Co.: M. = ;. Jones Se n. (Grinnell, August 1875] (Du--151781); A. R. Moldenke ce 96h, 6, (L¥) « Scott Co.t Suksdorf s.n. [June 29, 187h] (P1—138358), sme s.n. [July 28, 1874] (Pl--130358). Story Co.: Botanical Seminar 7 (Io-- 81132); W. P. Caldwell 18 (Au--12279h); J. Re Campbell 67 (to- 50252) 5 Pe Clayton yton 1h7 (To-~8562); Cratty s.n. (Ames, Jul. 1925] (Io—117327); A. S. Hitchcock s.n. (Ames, 1889] (Io—-15334); Jeffs sen. (ames, July 4, 1907] (Ok--10369); Pammel & Ball s.n. [ames, Aug. 21, 1897] (Io—15333). Tama Co.: Fisk son. [Jul. 1926] (Io 126759) . Union Co.: T. L. Andrews s.n. [Creston] (Io=-2292h) « Wapello Co.: Cratty s.n. (Ottumwa, Jul. 15, 1927] (Io--129751)- Warren Co.: A. R. Moldenke 945 (Lw); L. H. Pammel s.n. [Norwa Sept. 9, '27] (Io—130062). Webster Co.: Blumer hhil (1o--107739) 5 Paige s.n. [Fort Dodge] (Io--1310)5). Woodbury Co.: L. H. Pammel Sn. (Sioux City, July 7, 1919] (Io—-97278). County undetermined: Somes 3488 [Butler's Landing] (W--672271). KENTUCKY: Breckinridge Co.: R. Runyon 1260 (W—-138382). Edmonson Co.: Lix 478 (N)- Fay- Ft. cei 1915) (Ky-- 1780). Harlan Co.: F. E. Lloyd a [July 25, 1888] (Cm). Hendery son Co.: A. R. Moldenke 105 (Fg). (Fg). Jessamine Co.: F oS Ts Senes 92 (Ky, 8, Vi). Madison Co-t Wharton 835 (N). Nelson Co.: Agnes s.n. [July h, 1933] (1). scott Cow: Si Singer 308 (W--1465820). Union Co.: Shacklette 504 (By) » Webster Co.: A. A. R. Moldenke 106 (Fg). TENNESSEE: Davidson Co. Ainslie s.n, [Nashville, July 2, 1914] (Ka--66476); Gattinger $-M- {about Nashville] (Du--9132) . Hamilton Co.: Moldenke bode 2 (S). Henry Co.: A. R. Moldenke 111 (Fg). Jackson Co.: A. Re Mole denke 56 [H. N. Moldenke 21781] (Bm). Knox Co.: Ruth 780 (N)- Lewis Cos: C. B. King 176 (Au). Marion Co.: A. Re .. Moldenke Lea 1965 Moldenke, Monograph of Verbena 4,03 McMinn Co.: A. R. Moldenke 778 (B, S); Moldenke & Woods 500 (S). — Co.: A. R. Moldenke 779 (B, S). Montgomery Co.: Te L. An ews 116 (Io--83617). Obion Co.: Jennings & Jennings s.n. ~TReel- fone Lake, 1924] (Cm, Cm, Om). Sevier Go: ee S Moldenke 5) [H. N. Moldenke 21778] (Bm). Shelby Co.: Demaree 3 33199 (Sm). Tipton Co.: Byars s.n. (Covington, Aug. lh, 1898] (Ww. aaa), Wayne Co.: McDougall 1060 (W—19252h3). County undetermined: W. H. Camp 1806 [Smoky Mts. Nat. Park] (N), 1807 [Smoky Mts. Nat. Park] (N). MICHIGAN: Allegan Co.: C. H. Kauffman s.n. [New Richmond, Aug. ll, 1910] (Mi). Berrien Co.: Billington s.n. [July 29, 1919] (Mi). Gratiot Co.: C. A. Davis san. [Alma, Aug. 8, '39] (Mi, Mi), son. (July 29, '92] (Pl--2260h). Ingham Co.: L. H. Bailey s.n. (Lansing Aug. 4, 1885] (Ba); Collector undesignated s.n. (Lansing, 8/15/ 1867] (Lb--27623); E. B. Lake sen. [(8~7-88] (Or--887h); Yuncker 529 (Or--13548, Ur). Ionia Cos: Ce. fs Wheeler s.n. [Hubbardston] (MMi). Isabella Co.: A. R. Moldenke 1032 (Lw, Ut). Kent Co.: H. M. Bailey S.n. (Grand Rapids, July 29, 1892] (Mi, Mi); Daniels Sen. [Alto, June 1899] (Lb—~27622). Livingston Co.: Hamerstrom 197 97 (Mi). Me- nominee Co.: Grassl 2702 (N). Montcalm Co.: A. R. Woldenke 1034 (Iw). Ontonagon Co.: Es G. Voss 7169 (Mi). Saint Clair Co.: C. Ce. K. Dodge sche [8~28-92] (ui), Sn. 1. fAug. 15, 1910] (Mi). Washtenaw Co.: E. C. Almendinger s.n. [Ann A: Arbor, July 18, 1860] (Mi); K. Cc. Davis 108 (P ey, Harrington s.n. [Ann arbor, July 1870] (Mi); Ne A. Harvey s.n. [Ypsi, Aug. 16, 1916] (Mi); F. J. Hermann 9104 m5. “Ge De La Rue s.n. [Aug. 6, 1915] (Mi), s.n. tm. g. 7, 1915) (Mi); C. L. Lundell 8315 (1d); Whited 2108 (Or—-12381). Wayne Co.: Farwell - 1289 (C); Foote s.n. [Detroit, July 10, 1870] H. He Rusby s.n. (Detroit, Aug. 108] (Bz—23791, Bz—-23792, , se 149h6). Ww + WISCONSIN: Bayfield Co.: Shaughnessy & a SN. [Oct. 3, 1959] (Ws, Ws); Weber, Ugent, & Zimmerman s.n. [September 3, 1959] (Ws). Buffalo Co.: H. H. Smith 7180 (B). Columbia Co.: Botany 162 class s.n. [Sept. 28, 1935) (Ws); . Fernholz s.n. [Aug. 15, 1930) | (Ws); Koeppen 131 (Ws); H. L. Russell s.n. 11 s.n. [Poynette Aug. 20, '86] (Ws). or: Crawford Co.: “Fassett 9791 ‘(Um--186, Ws); fe R. Moldenke 99) (Lw); H. H. —— 7520 (B) (B); C. Wilson s.n. [Sep 3B, “1960] (Ws). Dane Co. C. C. Albers 33011 (Au), 33012 (Au); t: H, Chase s.n. [8/10/1877] (Ws); T. J. Hale sun. [Madison] (Ws); Hale & & McCabe s.n. [27 July 195] (Ws); Heddle 27266 (Ws); A. Re Moldenke 1011 (Lw, Ut); R. C. Monroe s.n. . [October 10, 1959] (Ws); Rickett s.n. (Madison, August 1, 1926] (Lb—27621); A. B. Seymour 1061 (H--107711, Ws); Stroesenreuther s.n. (Oct. 5, 1959] (Ws). Co.: A. R. Moldenke 1013 (Lw). Dunn Co.: Goessel 8956 (B); Weinzirl s.n. [Aug. 7, 92) (Ws). Grant Co.: Blazkovec s.n. [June Lok PHYTOLOGIA Vol. 11, no. 6 29, 1959] (Ws); Fassett 12528 (Ws); Iltis, Long, & Rudolph 13793 (Ws); Iltis & Salamun lanun 10058 (1 (Ws, Ws); A. Re Moldenke 998 2 998 (Im); L L. H. Pammel 1053 (Io—119695); Pammel & Johnson s.n. [Muscoda, Aug. 17, 1928] (Io—134359, St--9273); He ike Smith 7681 (B, Ws); Wun- derlin s.n. [Aug. 1h, 1960] (Ws). “Green Co.: E.5 « Chambers 27 — (Ws). Green Lake Co.: Kunz s.n. [Aug. Te, 1929] (Ws). owa COs: A. R. Moldenke 1002 (Lw, w, Ut). Juneau Co.: Bramschreiber gen. [July 15, 1959] (Ws). LaCrosse Co.: Hartley 1623 (Ws); A. Me Peterson 562 (Ws). Lafayette Co.: Amot sen. [July 26, 1956] (Ws); Iitis 9561 (1 (Ws); Shinners 2357 (N, Ws). Milwaukee Co.: J. S- Douglas s.n. (Je--3165); | Ramaley 3h); (Bl--423)3) 5 Shinners 1h79 (Ws); Strong s.n. (Wauwatosa, July yx 28, 1891) (Mi). Pierce Pierce Co.t Fassett 5631 (Ws). Richland Co.: Iltis & Salamun 9942 (Ws); i S.n. (Sept. 25, 1957] (Ws), son. [July 27, 1959] (Ws). Rock Co E, W. Fell 57-802 (Ws); Skavlem s.n, [Jonesville, July 23, 89] ge EE J. Wickham 15 (Ws). Vernon Co.: Fassett 20235 (Ws); Pammel & King s.n. (Portland, 8-26~'01] (Io--32602); H. H. Smith 7258 258 (B, Ws). Walworth Co.: Almon s.n. [July 23, 1926] ~(um—~20h) 5; Ne N. Hol- lister 137 (Um—196); Kleeberger s.n. [Whitewater, 8-1-1877] (Ge-- 31437). Waukesha Co.: Gull 236 (Ws); Schulte s.n. (Sept. 1959] (Ws). Waupaca Co,: L. Hansen s.n. [Sept. pt. 26, 160) (Ws); Rill se adr [July 16, 1959] (Ws). MINNESOTA: Hennepin Co.: F. H.B 8467 (Se—149k7); E. T. Harper III (P1—665h8) ; Je He Sore ¥3 n. n. [July 1890] (is). Rice Co.: Goldsmith 119 (N). NORTH DAKOTA: Stutsman Co.: Lunell 869 (Ur). SOUTH DAKOTA: Day Co.: Je Fe Brenckle s.n. [July 22, 1938] (B). Minnehaha Co.: P. Johnson 33 (Se--77271). KANSAS: bidire on Cos: As Se Hitchcock s.n. [Anderson Co., July 1896] (Ka); R. L. McGregor r 13169 (lw). Atchison Co.: A- S. Hitchcock s.n. [Atchison Co., Oct. 1896) (Ka); Horr & McGregor E.531 (AU--122795, B, Ca—ll13, No--15591, Ok, Rf, S, St, Tl, oi, Wi W--2031362), s sen. [10/L/)7] (Lw); Jeanette 165 (N)s As Re Moldenke 909 (Lw). Barber Co.: Kellerman s.n. sn. [Medicine Lodge, July =r nee 1888] (Ka); R. L. McGregor gor 14580 Gy), 14717 (Lw). Barton Co. A. S. Hitchcock s.n. [Barton | nm Co., August 1895] (Ka). Bourbon Co. Kel- lerman s.n. [Fort Scott, July 15, 1887] (Ka); Z. D. Thompson 50 (Lw). Brown Co.: Clothier & Whitford s.n. [July 29, 1897] (Ka); F. Garner s.n. [Aug. 1925] (Ka--7201k, Lb--27626); R. L. McG 170L8 (iw); A. R. Moldenke 914 (Iw). Butler Co.: A. S. Hitchcock s.n. [White Water, July 1892] (Ka). Chase Co.: A. Be “Pitcheock S- Hitchcock n. n. [Chase Co., Angust 1895] (Ka). Chautauqua Coe: A. Se Hitchoock .n. [Chautauqua Co., Aug. 8, 1896] (Ka); W. He Horr | Sone gee Ds 1930] (Lw). Cherokee Co.: A. S. Hitchcock sn. [Cherokee Co., May 1896] (Ka); A. A. Jacobs 21 (Ka--7h03h), 50 (N), 106 (Ka-~7)090)5 E. W. Lathrop 2758 (Lw); R. L. McGregor 15780 (Lw). Cloud Co.? 1965 Moldenke, Monograph of Verbena L05 Carleton s.n. [July 12, 1888] (Ka); S. V. Fraser 438 (Ka—789)1); A. S. Hi S. Hitchcock 8.n. loonctrdia, June e 17, 1897) (Ka). Coffey = Wimpey SoM. (Burlington, July 1896] (Ka). Cowley Co.: Clothier Whitford s.n. (Aug. 22, 1897] (Ka); A. Re Moldenke 885 (Lw); se berg & Imler L482 (Ly, N). Crawford Co.: Clothier | & Whitford S.n. (Aug. 12, 1897] (Ka). Dickinson Co.: A. S. Hitchcock s.n. [Dick- inson Co., June 1896] (Ka). Doniphan Co.: Clothier & Whitford s. Be [July 31, 1897] (Ka); F. C. Gates 20889 (Ka--89686); R. L. | R. L. Mc regor 15990 (Hi--1954)3, Lw)3 F. Schultz s.n. [Wathena, Oct. De] (Ka--7370h). Douglas Co.: Horr & McGregor E.531 (Ur); Kel~ ierman sen. [Lawrence, July 20, 1887] ¢ ] (Ka); Re L R. L. McGregor ape, (Iw); J. D. Poindexter 9 (Lw). Elk Co.: Clothier & Whitford s.n, {aug. 20 & 21, 1897] (Ka). Franklin Co.: Hetzer l19 (Lw); A. S.- Hitchcock s.n. [Franklin Co., July 1896] (Ka). Geary x pe Os Gates 18768 (Ka--87795); A. S. Hitchcock s.n. (Geary C 1896] (Ka). Graham Co,: re S. Hitchcock s.n. [Graham ies pi 20, 1897] (Ka). Greenwood Go.: rt Ss. Hitchcock s.n. (Eureka, 1892] (Ka); W. H. Horr s.n. [June 28, 1930] (Lw); R. L. McGregor 14699 (Lm). Jackson Co,: Clothier & Whitford s.n. [July 28, 1897] (Ka). Jefferson Co.: Clothier s.n. . (Perry, 0 Oct. one, 1896] (Ka); G. Latham 651 (Lw). Johnson Co.: A. S. Hitchcock s.n, (Olathe, Aug. 1892] (Ka). Kiowa Cos: A. S. Hitchcock s.n. [Kiowa Co., Aug. 1896] (Ka). Labette Co.: A. S. Hitchcock s.n. [Labette Co., Aug. 1896] (Ka); R. L. McGregor 16996 (Lm); B. L. Wagenknecht 1889 (Lm, lw). Leavenworth Co.: A. Se *, Hitchcock son. [Leavenworth Co., May 1896] (Ka); R. L. McGregor 840 (Iw). Linn Co.: Kellerman s.n. {Mound City, J July 18, 1887 1887] (Ka); A. R. Moldenke 869 (lw). Ly Lyon Co.: Re L. McGregor 189k (Lw). Marshall C0.: A. Se Hitchcock s S.n. (Marshall ¢o., Dec. 10, 1896] (Ka); W. H. Horr 4502 (Iw). McPher- Son Co.: A. S. ibaheneks s.n. [McPherson, July 1892] (Ka). Miami Co, A. ee Hitchcock s.n. [Paola, Aug. 1892] (Ka). Mitchell Co.¢ A. S. Hitchcock sen. (Beloit, June 18, 1897] (Ka). Montgomery Co.: Ae 3. Hitchcock s.ne pages COs ne 1896] (Ka); Rydberg & Inler 132 (Ka——7h56h, lw). Morris Co.: A. S. Hitchcock s.n. [Mor- es (Ka). Neosho Co.: A. S. Hitchcock s.n. [Neosho Pl ies 1896] (Ka); W. H. Horr 6729 (Lm); E. Le Richards 3176 (Lw). Osage Co.: 2.D. z. Brown 5.n. [July 28, 1896] (Ka); B. L. Wagenknecht 3969 (in), L086 (iw (Iw). Osborne Co.: Neher 348 (Ka——7993k) « Ottawa Co.: A. S. Hitchcock s.n. [Ottawa Co., July 1895] (Ka); A. R. Moldenke 901 ( (Lv). Philipps Co.: A. S. Hitchcock s.n. [Philipps | Co., o., June 29, 1897] (Ka). Pottawatomie Cost D. L. Marsh 1692 (Im); Pond s. 406 PHYTOLOGIA Vol. 11, no. 6 n. [July 2h, 1896] (Ka); B. L. Wagenknecht 3256 (Lw). Republic Coe: G. Ez. i 466 (Lw ), 810 (Lw); — Ae , Sait Selle apes si aes. mean ideo * Wiican nengh sen. [Aug. 16, iciages (Ka). Saline Co.: + Hanein 429 sate boned TL L. Andrews | sn. (Whichita] (Io--28226); pacts ee Univ. 168 (: (Lw). Shawnee Co.: Maus 842 (Ka-~72351); Norton & Clothier s.n. . [July 4, 1895] (Ka); Volle 327 (Lw). Smith Co.t A. S. S~» Hitchcock s.n. (smith Co., June 28, 28, 1897] (Ka). Sumner Co.: he Bi Hitchcock s.n. [Wellington, July 1892] (Ka). Wabaunsee Co.: J.B. S. Norton s.n. (Maple Hill, July 4, 1894] (Ka). Washington eet ee 80570); We He Horr rr 4655 (lw). Wilson Cost Haller s.ne n. (Roper, June 16, 1896] (Ka). Woodson Co.: Clothier & Whitford s.n. (Aug. 29, 1897] (Ka); W. H. Horr s.n. [July 10, 1930] (Lw); Horr & McGregor sen. (July 17, 197] (lw); E. We Lathrop 130; (N), 13u6a (Lw); Mrse 0.5 E. Rogers s.n. [Yates Center, July 11, 1928) (Ka--76503) « Wyan- dotte Co.: K. K. Mackenzie s.n, (Rosedale, July 12, 1896] (Ka, N)i E.L. Richards 3289 (Lw). MISSOURI: Bates Co.: A. R. Moldenke 868 (Iwy. “Boone Go.: Daniels s.n. [July 1903] (Lb—~],180, Tb—L181 15 : Dunn Aly (Bo—Is68) 3 Favor s.n. [Aug. 29, 1901] (Lb--)179); Jeffrey 105 30365, Lb—30372); Rickett s.n. (Sept. 1926] (P53), 3 S.ne sehiade wk July 16, 1927) “3927] (Lb—-7092). Camden . Moldenke 857 ( Clay bass Brouillette s.n. [Liberty, 7/ 1/33/38 oT = . "Bush 1269hd (Lb—1165) sist BS 8129 (S). sb ae Co.t Ae Re Moldenke 86 ary Greene C ship s.n, (Willard, July 9, 1919) Sd hl P. C. standley 62788 iy Both (W—687210). Grundy Co.: Crookshanks 11 (Lb—29612). Henry Co-? H. Nz Moldenke 21922 Ey A. R. Moldenke 859 (Lm)+ Jackson sen, [Kansas City, Jwy 27, '9L] (Ah); Ke ae Mackenzie SIT Ge), (We), sen. yen. [duly 19, 1896] (N). jefferson Co.: Favor 588 (Lb--178). Lincoln Co.: A. R. Moldenke 81 (lw). Maries ; Coet He N. Moldenke 21918 (Iw). Marion Co.: J. Davis 4592 (Ur). Me Donald Co.: BE. E. J. Palmer 4229 (W~1569310). Miller er Co.t Ae Re Mol- denke 854 (Lw). Nodaway Co.: J. R. Singleton 154 (Lb—-34597) « Pemiscot Co.: Dinmith 477 (Lb—L177). Phelps Co.: A. Re Moldenke 850 (Lw). Pike Go.: J. Davis 950 (N); A. R. Moldenke 339 (Lw, Ut)- Platte Co.: Radloff s.n. [July 5, 5, 1956] “(ur). Saint Saint Louis Co.t Collector undesignated s Sn. (Aug. 1893] (S); Eggert 19h9 (Lb— 1176); C. Epling 6816 (La); A. R. Moldenke 69 [H. N. ~Moldenke 21799] (MMi); H. N. Moldenke 21800 (Mi). Shelby Co.: Ripley SMe (Sep. 10, 1902] (Je—3162, Je—3163), Warren Co.: A. Re woldenke 1965 Moldenke, Monograph of Verbena 407 843 (Iw). Saint Louis: C. F. Baker sen. [St. Louis, Sept. '99] Cassie a Eggert s.n. (St. Louis, Aug. 1875] (au, Ca~~183210, o—~79866, Mn-—6897, N , W=751956); Engelmann s.n. (St. Louis, eons 18 38] (Dt), Sen. [st. Louis, Sept. 181] (W (W-—-71969), sen. [St. Louis] (Br); Ke i Martens s.n. [St. Louis] (Br); Riehl 135 — (Br, M). County undetermined: Engelmann s.n, (July 1839] (Br); Spillman 298 (Pl-~22608). ARKANSAS: Arkansas Co.: Denaree 14218 (Sm). Ashley Co.: Demaree 23898 (Sm). Baxter Co.: Demaree 23602 (Sm). Benton Co.: Heacox s.n. (July 20, '89] (Ob—508L). Brad- ley Co.: Demaree 20689 (Sm), 23865 (sm), 24845 (Sm). Clay Co.: Demaree 26997 (Au--122803, N). Craighead Co.: Demaree 25777 (N), 292k] (Le). Cross Co.: Demaree as (Lb—L67L4, N, Sm). Drew Co.: Demaree 13601 (Sm, Tl), 17620 (Du--256560, Ge-26761b, N, Ok), 17897 [ [Herb. Drushel 11619] (N, Ok, Ur), 17907 (N, Ok), 2hh62a, in in part (Bm, Sm), 24517 (Sm), 24748 (Sm). Faulkner Co.: Welch | 1292 (Dp). Franklin Co.: J. M. Mason s.n. [Meramec State Park, 8/16/37] (Lb—34373). Fulton Co.: Demaree 26391 (N), 26826 (Au—122816, N); Zirkle & Drouet s.n. [Aug. 19, 192 28] (Lb—-2762h, Lb—~27625). Garland Co.: Demaree 20470 (Lb-~53K16, N, Sm); Re Runyon 1487 (W--1568007). Green Co.: D Demaree 27097 (Au-—112817, N). Hempstead Co.: Demaree 21180 (Hi--170691, N, Sm), 46061 (S). Hot Spring Co.: Denare aree 15563 (N), 15609 (Io—152223, Ok), 16520 (Io-~152211, Ok, Or—-l)3132, Ur), 17h60 (N), 17887 (Bt—52h79, Io- 152221, Ky, Ok, ae Pl—-131515, Ur), ir), 18887 (N), 1 19321 (Sm). John- son Co,: ale 19993 (N, Sm), 20282 (N, | (N, Sm). Lawrence Co.: Demaree 30987 (Sm); Rolfs s.n. [8/91] (Io--h489). Lee Co.: = aree 37358 (Si (Sm). Logan Co.: P Pyle 831 (De). Lonoke Co.: L8818 (iw). Marion Co.: Demaree 20629 (N, Sm), 45947 (S). ~ Milter Co.: Demaree 1,898 (Ca). “Montgomery Co.: Demaree e 46623 (S). Neva- da Co.: M. P M. P. Hollister 110 (W--587798). Ouachita Co.: 46033 (S). “Phillips Co.: Demaree 37358 (Sm), 37402 (Sm, “Z)« Po Pope Coss | Demaree 19886 (Lb--53417, N, N, Sm). Prairie Co.: Demaree 15527 (N). (N). Pulaski Co.s Demaree 8193 (Bt-~10318, Ca——-717k1, Dm, Du—233%62h, N, W--158161), 8632 (W--1580200). Randolph Co: Demaree 23906 (Bm, Lb—1,7030, Sm). Sebastian Co.: Bigelow s.n. [Fort Si Smith] (W--71976). Sharp Co.: Demaree 26273 (N). Stone Cos: Demaree 27872 (N). Yell Co.: Demaree 20121 121 (N), 25121 (Sm). LOUISIANA: Bossier Par.: Correll & Correll 10057 (H--76636, N); Penn s.n. (Oct. 29, 191] (Tl). Bast Baton Rouge Par.: L. He Bailey s.: S.n. [Baton Rouge, July 16, 1918] (Ba). Lincoln Pars: Cocks son, [July 7, 1901] (Tl). Livingston Par.: Correll & Cor- Fell 9129 (H+~7859]1) « Natchitoches Par.: Biology Class 302 no. bettie 2169 [L. E. Fox 2169] (Nc); BE. J. Palmer 8000 000 (Gg~-182150). Or- 1,08 PHYTOLOGIA Vol. 11, no. 6 leans Par.: Cocks s.n. [June 1900] (Tl); T. Drummond s.n. [in vicin. New Orleans, 1832] (Lu). he ei Par.: Langlois 611 (N), Sen. [June 1882] (I). Rapides Par.: C. R. Ball 556 (N, W- a Saint Landry Par.: Ewan 19328 (Tl). Saint Martin Par.: Correll & Correll 9406 (H— 78593). Saint Tammany Par.: Arséne Trees t (W--1031192), 11391 (W—1031h96), 12069 (W—-1031916). Par- ish undetermined: Jos. Hale 2h] (T); Herb. Hapeman s.n. > Rid- dell 1261 (N). NEBRASKA: Adams Co.: H. Hapeman sn, [Aug. 5, '92] (Hp); R. de Lima 824 (I). Cass Co.: T. A. Williams 13 (W--71975) « Cherry Co.: Tolstead 706 (Io—-1i5167). Custer Co.: J. M. Bates 1962 (Io—35178). Douglas Co.: Kleeberger s.n. [Omaha] } (Gg——31438). Jefferson Co.: Rohrbaugh 167 (Ok). Kearney r COs: Hapeman s.n. [Min- den, Aug. 29, 1931] (Au). N Nemaha Co.: A. R. Moidenke enke 923 . (LW, Ut); Shubert sen. [July 10, 1933] (Ob--5087). Otoe Co.: A. oie Allen son, [Talmage, July 16, 1891] (Je-—-316h); C. J. Elmore | SoM. [tal-. mage, July 6, 1891] (Je—3166). Platte Co.: Clevenger Sen. [Colum- bus, July 11, 1904] (La). Seward Co.: Kiener 17088 (Ca~-36917) - county Aiedetesatneds Herb. Univ. Nebraska 436 (Au). OKLAHOMA: A- toka Co.: Cory 56795 (St). Bryan Co.: We W. S. S$. Blain son. [Durant, June 28, 1935] (Bt—29691). Caddo Co.: Edw. Palmer 2h) (T, W— 71955). Carter Co.: M. Hopkins 6366 (Ok). Cherokee Co.: C. S- Wallis 657 (st), 3396 (: (St). Choctaw Co.: H. W. Houghton 4032 4032 _ Gleveland Co.: R. E. B Berry 58 (Ok--10347); Bruner s.n. [8/2 /20/2h) (Io--117043, Ok-=10343); Collector undesignated 227 (Ok--10322);5 C. T. Eskew 2001 (Ok); M. Hopkins 109k (Ok), 4610 (Ok); Jeffs s- ne [6/22/1922] (Ok--10339); Personett 96 (Ok--103bh). Comanc jonanche Co.: M. K. Clemens 11719 (Ba), 117l9a (Gg--371L3). Custer Co! Waterfall 1 297% (Ok). Delaware Co.: Blevins 63 (St--18581); C. S» Wallis Wallis 2436 (St). Garfield Co.: Gephardt 655 5 (W--221182) . Johns- ton Co.: G. J. Goodman 6325 (Ok); G. T. R Ge Te Robbins 263 (Ca--9660L1, N, Ok). Kay Co.: Byler 190 (St); G. W. Stevens 1821 (Ok, St— 927k) « Latimer Co.: 0. M. Clark san. (Red Oak, 6/13/30] (Ok-~ 10341). Major Co.: G. W. Stevens 1742 (Ok). Marshall Co.s G- Je Goodman 5903 (Ca~~81888, | Ok, St). Mayes Co.: C. S. Wallis 2 (st), 3 5233 (St). McCurtain Co,: Hixson 100 (St—-13227)5 Iw y 25h 20371); Waterfall 12412 (ca--72ll2, St, W--2231160). ~ vurray pes Hopkins & Demaree 50 (Ba, Ok, St—~12768); G. M. Murrill 716 (N, RE); Stratton 738 (N, St--26137, St--26138). Muskogee Co.: E. L. Little 189 (Ok--10340), 1931 (Ok—-10342); C. S. Wallis 7719 (Au--L70516, St). Oklahoma Go.: Waterfall 2hh (N), aoe (OK, St), s.n. [June 2h, 1939] (Ok). Payne Co.: R. 0. Bain hh (PL =124970) Bogue s.n. Lye ei 1896] cco 97a Brensing 35 (Ca-—882833)5 R. M. House 95 29718) ; arn son. [duly 14, 1916] (St 9271); Pote 6 (Beh 282)" Stratton 168 (st ~-26123), 3608 (15 1965 Moldenke, Monograph of Verbena 09 Walton 143 (Au); Waterfall 123 (St); Waugh s.n. (Stillwater, 1893] (Ka). Pontotoe Co.: D. McCoy joy 807 (St), 1b 1421 (St), 1725 (Au-- 122805); G. T. Robbins - 3110 een 3 C. Thomas s.n. [Ada, l- 2-35] (Ok). Sequoyah Co.: C. S. Wallis 770h (Au--170517, St). Tulsa Co.: M. B. Clark 52h ao J. Re . R. Willis 3 (0k). Woods Co,: G. W. Stevens 1673 (Du--66082, N, Ok, Ok, Ok--2071h, st--9275, St--9277, Ur, W--589621). Woodward Co.: Engleman 513 (St). County undetermined: Clifton 91 (Ok--103)6); Herb. Univ. Okla. 10345 (0k). TEXAS: Angelina Co.: H. J. Hamby 1472 (Ar—-206312). Baylor Co.: Tharp 4501 (Au). Bell Co.: S. E. Wolff 1329 (W—1622h00), 32h7 (Tr, Tr--16239). Bowie Co.: “Tharp 277 (Au). Brazoria Co.: Tharp 2777 (Au), s.n. [8.21.29] (Au). Camp Co.: Moldenke & Woods 561 (8). Cherokee Co.: Ness s.n. [near Rusk, Aug. 15, 1926] (Tr). Dallas Co.: Boll s.n. [1871] (Cb); C. L. Lundell 11581 (N, Rf), 1022 (N, Rf, Rf); M. M. B. stephenson s SN. {vicinity of Dallas, June 20, 1929] (W--1432660) Denton Co.: B. B. Harris sen, [7-20-27] (au, Nt, Nt); Whitehouse 16426 (Mi, N)- Ellis Co.: 0, Sanders 5 Mi). Gonzales Co.: McCart 158 (Au--122793); Tharp s.n. . [Ottine, T-h-1929] (Au, N), Sen. (Ottine, 7-14-1929] (Mi), sen. [Ottine marsh, 6-26-39] (Au). Grayson Co.: H. Gentry 51-1786 (Au--122791). Hem enphill Cow: C. M. Rowell 5563 (Au--195L10). Hunt Co.: Thurow 8.n. (Clinton, “Oetobet 10, 10, 1924] (Hu). Jefferson Co.: G. Le jean ers S.n. [Nome, Sept. 2, 1937] (Ew, Ew, Ew). Lamar Co,: “Turner 8. ne [Deport, Sept. 188h] (Mi). Milam Co.: S. E. Wolff 430 (Tr he Montgomery Co.: Warner s.n. [R.R. yard, Conroe] (Hu). Nacogdoches Cow: W. H. Lewis 4999 (Nd). Newton Co.: Tharp sen. [7-23-39] (Au, Au, Ca--882832, Mi, N). Robertson Co.: Reeves 187 (N). Tarrant Coe: Killian 6933 (an, W—~165329); Ruth 503 (( (cm), 53 (N, Sm, Ur). Trinity Co.: Lewis & Oliver 530k (Nb); Tharp 2778 (Au, Au). Wise Co.: Whitehouse 1637h (N). Wood Co.: McMullen s.n. [Golden, 6/17/ 27] (Au), sen. [9-10-27] (Nt). CULTIVATED: Austria: Haldcsy s.n. (hab; Theresiani, Aug. 1871] (La). Belgium: M. Martens s.n. (Br). England: Linnaeus s.n. (Hort. Cliff. G.83h, S.5; Bailey Hort. neg. 6405] (N--photo of type). France: Herb. Harvey s.n. [h.R.P. 1819] (Du-166h4h), san. [h.R.P. 18h] (Du--166441), son. [h.R-P.] (Du-- 15543h); Herb. Molliano sen. [PJ] (Br); Herb. Schumacher s.n. [h. Paris] (Cp). Germany: Herb. Harvey s.n. [Pottstadt hort. bot. — Gosia R. Schulz sn. (Berlin, 11.8.1897] (B). Italy: sen. [h.P. 1520] (Du--1664)3). Maryland: McCann s.n. (Sas SeT te (Md). New York: Eggleston & Schneider s.n. [July 1 July 18, 1905] (N); R. C. Schneider s.n. (N. Y. Bot. Gard. Cult. Pl. 25117] (N). Pennsylvania: Je Me Greenman 1374 (Du--123485). Spain: Herb. Hort. Hispal. 67 (Q); 3 Herb. Hort. Reg. Matrit. 59 (Q); Pascal 66 410 PHYTOLOGIA Vol. 11, no. 6 (Q). Sweden: Linnaeus s.n. [hort. Ups.] (F--photo, F--photo, N-- photo, N--photo, S, S, » 5, 8; Sg--photo, Sg--photo, Z--photo, Z—photo); Nietzel s.n. fhort . Ups. ] (F--photo, N--photo, S, Sg--photo, Z—— photo); Reuterman s.n. [19/9/1888] (Go); vicars sen. [1925.56 VIII] (s ). Switz Settserlends Herb. Hort. Bot. Bern. s.n. [Aug. ape fot. Genev. 8.ne. (dul. 1625. Herd. HOt. LOU. GENSV. SNe Collector undesignated 63 (Q), 64 (Q); Dascomb s.n. [13.1] (0b-— 50852); W. Dearness 83 Toaklands]} (Mg); Disbro Disbrow s.n sen, (Nm); Engel- mann sen. [Amer. bor.) (Br, S); Freed s.n. ne (Amer, . bor.) (S)5 Gib Gib- bes s.n. (N); Herb. Buckley s.n. (T); Herb. Chapman s5.n- [New Eng- land] (C (C); Herb. Coll. Pharmacy s.n, (Pa); Herb. jerb. Liebmann Sone {North America] (Cp); Herb. Lin Linnaeus G.35, S13 (Ls, N--photo, Z-- photo); Herb. Martius s.n. sen. (Br); Her! 3 Herb. Mus. Bot. Stockholm 14 (North Am.] (S), sen. (S)} Herb. Saldanha 2767 [U.S.A. Herb. Mus. Nac. Rio ref. VII] (Ja); L. A. Koehler s.n. [U.S] (Pa); Linderall s.n. [N. Am.] (S); M. Martens sen. (Br); N. XY. Bot. Gard. Economic + 5750 (N); Osbeck s.n. sen. (S); Palisot de le Beauvois s.n. [Etates re 1819] (Du--1660); Schumacher sa sn. . (North Am.] (S)3 J. Tor- rey s.n. [ex Amer. septentr.] (Bm, S), Se sen. [Fl. Bor. An.] @s.- Weigand s.n. ek Berea pao a As huctens aes [N. Eng.] (Pa)- MOUNTED ILLUS Rhodora 38: pl. 45, fig. 5 a peg 1936 wye Ho. wi8 pb he. CTS (Nt) . ENA URTICIFOLIA var. INCARNATA (Raf.) Moldenke, Phytologia 7: 259. 1960 Synonymy : Verbena incarnata Raf., Atl. ea a ace Bibliography: Raf., Atl. Journ. 15h. 1832; ‘ 69. 1833; Jacks. a Hook «:ifo% Jacke. , ind. Kew. ne 79. 1058) Perry, Ann t. Gard Raf. 205 & 295. 1949; Moldenke, Phytologia 7: beg. 1960; “yoldenke, Résumé Suppl. 2: 1 & 15 (1960) ee 7: 9. 1963; Moldenke, Phytolo- gia 9: 93 (1963) and 11: 338. 1 This variety is said to ante from the typical form of the Species in having narrower ower leaves, the spikes not panic culate, and the flowers flesh-color instead of white. The type of the variety was collected by Samuel Constantine Raf- inesque along the Juniata River (this could be in Huntingdon, i lin, Juniata, or Perry Counties), Pennsylvania. The original de- scription is "Stem branched; leaves ovate lanceolate serrate rough, owers in — slender short spikes....on the Juniata R....one foot high, flowers flesh-colored. Differs from V. urticifolia nar rs not. Without an examination of the type specimen it is — —— to be certain of the status of this plant. metic mere 1965 Moldenke, Monograph of Verbena 412 color variant of V. urticifolia L., and possibly the Shinners 1479, whose fiowers are described by the collector as having been "pale bluish-white", and Harris s.n. [June 26, 1933], whose flow- ers are said to have been tpink" — both cited by me under typi- cal V. urticifolia — and F. R. Fosberg 15695, whose flowers are said to have been "pinkish-white" and Eames & Wiegand 12796, with "pink" flowers, cited by me as var. leiocarpa Perry & Fernald, be- long'to var. incarnata instead. It is also not beyond the realm of possibility that Rafinesque's plant was nothing more nor less han xV. engelmannii Moldenke, but I am inclined to doubt this. VERBENA ei var. LEIOCARPA Pe Fernald in Fernald, Rhod ex Kriebel, Yuncker, : Friesner, Proc. Ind. Acad. Sci. 51: 127. ike. . Verbena ultricifolia var. "le ae Perry & Fernald ex Moldenke pee a ae 609, e 1945. Verbena ur- ticaefolia var. leiophylla Fern. & eines ex qx byatl, Alph. List nvalid Names Suppl. 1: 27, in syn. 1947. Verbena urticaefolia f. ieiocarpa Deam, Yuncker, & Friesner, Proc. “Ind. Acad. Sci. 60: 87. 1951. Verbena urticifolia leiocarpa Perry & Fernald ex Moldenke in Gui teenie, Roy. Hort. Soc. Dict. Gard. 6: 2209. 1951. Bi or ag ae Perry & Fernald in Fernald, Rhodora i en pl. 450 - 5-8. 1936; C. C. Deam, Fl. Indiana 795, 7 bees map i 1940; Moldenke, Suppl. List Common gee et 1940; denke, Obs. Fl. Watchung Suppl. 1: 1. 1941; eorees & Friesner, Proc. Ind. Acad. Sci. 51: 127. "1925 Yoldenke, Geogr. Distrib. Verbenac., [ed. 1], 2--l, 6-8, ll, 12, 76, & 102. 1942; Moldenke, Known i Distrib. Verbenac. Suppl. is 1 Noldenke, “alph. List cit. 1: 5, 10, 1b, 15, 18, 25 , 125, 130, 139, 142, 1hh, 148-252, 153, “tbo, 175, 177, 182, ish, 197--199, 204—-206, 208, 213-215, 23h, 8, 253, , 267, 270, 271, 278-282, 2h, 267, 288, & 291— enk . Friesner, rote: rare Acad » 110 (a5u ye 5 i: 3. 19183 Moldenke, Wright 22h. iol; “Yoldenke, Castanea 13: ; & 121, 19485 ‘eideske, Alph. List Cit. 2: *296, peneden fi 396-00, Lo, 455, 469, L91—93, 196, “506, 512, 513, $20, Sah, 538, Shik 555, 558, pay 383, 58h, 589, 594, 606, 617, 620, & 630 (1948), 3 653, 98, 721, 7h3-—7h5, 747, 750, 759, 760, 763, 778, 785° Sie 823, 828, 82, 848, 887, 892, 898, 912, 927, 412 PHYTOLOGIA Vol. 11, no. 6 93h, foie 92, 963, & 977 gee ge and h: 979, 985, lgenrenag : 1095, 1098, 1109, 1112, 1234, 1117--1119, 1122, 1123, 1137, 17h, 1176, sy 1179, 1181) 1185; ap, 1211; 222, & 1287. isle) Wol- denke, Known Geogr. Distrib. Verbenac., fed. 12--18, 20, 2h, 165, & 200. 1949; Moldenke, Phytologia 3: 2, 10, «131 (1949) QO; i zaBs 129 & 130. 1952; Deam, Yuncker, & Friesner, Proc. Ind. Acad. Sci. 61: 75. 1952; Voldenke, Phytologia h: 185. "1953; Moldenke, Notes Pl. 2: 6. 1957; Moldenke in Gleaso: on, New Britton & ge tin xt i pe 2; 3: 129 & 130. 1958; Strausba’ augh & Cor 3: 77h. 1958; Moldenke, Résumé 6—11, 15—21, 23, 25, ioe “30; 2 205 f7h. 1959; ae, Résumé Suppl. te laws ae 139595, 2 2 & 13 (1960), 3: at, 36, & Ll (1962), h: 1-3 (1962), 5: 1 (1962), and 6: 1, 2, hoes Steyerm., Fl. Mo. 1258. 19633 loldenke, a Ang ogia 8: are (1963), Aa re (1963), 10: 101, 106, & 09 (196k), and ll: 55 & 128. 196; Moldenke, Résum§ Suppl. 8: 1& é 6 (él) ‘aa il. 1965 Woldenke, Saar 11: 338, 3h1, 342, Illustrations: Perry & Fernald in Fernald, Rhodora 38: pl. 50, fig. 5-~8 [by E. C. Ogden]. 1936; Moldenke in Gleason, New Britton ry Iliustr. Fl., print. 1, 3: 130 (1952) and print. 2, 3: 130. This variety differs from the typical form of the species having its leaves densely and uniformly puberulent, Bares oe or velutinous beneath, the ae to 0.3. me long, spread over the re scenc anches usually filiform, loosely ascending or diver sy eaenigpatva the mature calyx to 2 @ a yd 8 @ J g o J cE be | @ the — of the variety was collected by Merritt Lyndon Fernald, Bayard Henry Long, and John Milton Fogg, Jr. (no. 5013) in rich woods at Virginia Beach, Virginia, on September 10, 1935, and is deposited in the Gray Herbarium of ponies University. In the original publication the at CROWN RD are also cited: CON~ NECTICUT: Hartford Co.: C, Wright te yereee ery (G). NEW JERSEY: Union Co.: Long s.n. (Port Elizabeth] (G). VIRGINIA: _ of Wight Co.: Fernald, Griscom, & Long 6674 (G). Southampton Co? Fernald & Long 686) (a). . Sussex Co.: Fernald, Griscom, & Long OOl/ 6672 (G). NORTH CAROLINA: Durham Co.: Blom omquist 149 quist 119 (G). SOuTH Hf CARO LINA: Williamsburg Co.: Wiegand & Manning 271) (G). with stiff hairs up to 1--1.3 mm. long, or glabrate; the mature in florescence......with usually stiffly ascending strigose branches} mature calyx.....strigose, 2—-2.3 mm. long, the subtending bract 1965 Moldenke, Monograph of Verbena 413 1.5 mm. long; mature nutlets......about 2 mm. long and definitely corrugated or ribbed on the back. Var. leiocarpa, on the other » has the thin leaves.....velutinous or a beneath ry rarely 0.3 n lent, with very short (0.5--1 mm. long) bract; and the ti lets... --only 1.5 mm. long and quite smooth on the back. Although =) Sebievt teh of the New England Botanical Club from north of New Jersey. "Verbena urticifolia var. leiocarpa, although having the same general lax habit as #V. scabra Vahl, collected by Fernald & Long Creek, near Cros ek Landi south o. Point, no, 686 e first collection nie nort Wi , North Carolina), may be readily dist din flower by pose "pilobed character lies between two almost a sterile Lobes. The f calyx of V. urticifolia, var. lei is only slightly ppc At from the rachis and the nutlets are smooth; whereas, in V. scabra the fruiting calyx is s trongly divergent and the wart tintes is reticulate Thole Furthermore, after drying, the plants in question are easily separable on * foliar character, the upper surface of t leaves of V. urticifolia, var. leiocarpa being much less harsh to the touch than those of V. scabra." The variety is native to the eastern and central United ea with Common names recorded for it are wiiet tLe-Leaved verv "short-hair white vervain", "white vervain", and "white vervian". It has been collected in open or waste places » ary waste Places, ditches and wet ditches, bogs and boggy depressions, marshes, thickets and fencerows, fields, old or dry fields, wet sloughs, , low pastures and pasture land, wooded and small wooded ravines, woods, rich or alluvial woods, dry or moist woods, dense ponds, in mixed or oak-hickory woods, moist woodlands, low marshy land, “in flatwoods, semi-shaded areas, moist places near roads, ~hickory or pine-oak-hickory peers or mixed deciduous er ound of pastures, in dry or clay soil, swampy or moist peste soil, in low brusty soil, and sand or clay as well as in wallace gravelly sandy loam, on open or wooded limestone ly PHYTOLOGIA Vol. ll, no. 6 hillsides, limestone cliffs, wooded bottomlands, riverbanks and wooded riverbanks, stre am banks rocky creek bottoms and banks, creek bottoms and. flat open gulch bottoms, on dry wooded banks, wooded and loamy wooded slopes, — anid slopes, and grazed be ound at th oak-sugar maple — and at woodland and low woodland borders, in growing in the shade, at altitudes of 200 to 1200 feet, - owering from June to September, and fruiting from July to dcto- er. In New Jersey it is said by Mackenzie to be "common Ms ——- in Iowa it is described by Pammel & Spiker as growing weed in low ground in pastures, widely distributed, ass mocked oh with Scirpus atrovirens, Echinochloa crus-galli, and Pers Sous ad folia. Schallert says that in Wisconsin it is "weedy bu troublesome", while in Indiana it is called a "weed in a acne by Ek. In North Carolina it grows in the "maple-birch-magnolia association" according to Radford. Johnson says "not ¢ Material of V. urticifolia var. leioc has been misidentified and distributed in herbaria under the names V. caroliniana L., ve hastata L., V. hastata x urticifolia Eggert, V. polystachya H.B.K-, V. scabra Vahl, V. urticaefolia L., V. urticaefolium L., V. urtici- aefolia Le, Ve urticifolia L., and V. urticifolia Gray & Small. On the other hand, the Friesner 6044 and H. N. Moldenke 15426 distribu- ted as V. urticifolia yar. lelocarpa are trplenl 3. mati icifolia L. instead. The Herb. Hort. Bot. Genev. sen. [18 Aug. 1020], cited , is a mixture with V. recta H.B.K. H. Ne areas 13982 is infested with Erysiphe cichoracearum P. DC., determined by y Dr Fe De Kern. Plymale 52 has very short hairs. F. R. Frosberg 15695 is carnata (Raf.) Moldenke. P. Johnson 33 may actually be the typical form of the species. E. E. Chambers 27 has inflorescences exactly like those of var. leiocarpa, but all its other characters those of typical V. urticifolia, so I am citing it as that. The Bouché s.n. cited below does not have any indication on its ; assuming that it did. The "Stearns" County, West Virginia, recorded for this variety by me in my Résumé 9 (1959) appears to be an error in transcription sags Stearns County, esota. Eggleston 481, Grimes 3794, Visher 2263, Rydberg 1716, Aiton er ], an K. Small s.n. [Reed Creek] are all cited by Perry (1933) as ‘tyateat V. urticifolia, but this was 1965 Moldenke, Monograph of Verbena 415 — before she recognized the variety, which these collections ally represent. G.N. Jones (19455, ee variety to synonyt under V. urticifolia. J. A. Harris C. Lo A ee eee it from mtiny Delawaite Poa Hun tingt on, Orange, Owen, and In all, 379 herbarium iedtueen and 1 mounted illustration have en examin by m Citations: NEW YORK: Bronx Co.: T. W. Edmondson 1,92 (N); G. V. Nash s.n. [Bronx Park, July 1h, 1898} Ti) becca ne Co.: Standl ey & Bo: Bollman 11996 (W-~895562) . Orange Co.: Raup 7603 (N). Queens Co.: Ferguson s.n. [Kew Gardens, 8-1-19] (N). Steuben Co.: He Ne Moldenke 2055 (N). Suffolk Co.: Ferguson 4382 (N); J. A. Harris C.15997, in part. (B)s; L. K. Henry s.n. Sen. [July 23, 1926) (Cm); Shull fmll 51,3 3 (N). Tompkins Go.: Eames & Wiegand 12796 (S). Ulster Co.: Macfarlane s.n. (Phoenicia Valley] (Up--l6l1h); eee 4803 (Al). Westchester Co.: Herb. N. Y. Bot. Gard. s.n. (July 13 1896] (N), sen. [August 5, 1696] (N); Monachino 67 (Ec, +3 ¥. Ae Weber 1043 (Bl--\8171). County undetermined: Herb. Torrey Sen. [New York] (T); J. Torrey 747 (Al). NEW JERSEY: Burlington Co.: H. N. Moldenke 21875 (Fg). Camden Co.: J. M. Fogg 7633 (Up). @ May Co.: Fender 1159 (Up); Macfarlane s.n. [Holly Beach, Sept. 1, 1907] (Up-lj5857). Essex Co.: K. K. Mackenzie 349 (Dm, N). dindee ster Co.: J. M. Fogg 862 (Up); B. H. Long 26200 (Ca~-8828)3). Mercer Co.: J. a. Fogg 5376 (Up). Morris Co.: Reis s.n. [Lake Hopatcong, Aug. 189] (N). Somerset Co.: J. M. Fogg 1232s (Up), 12395 (Up); H. N. Moldenke 11788 (N), 18362 (Ec). Sussex on Krausser & Cohen s wn. (Aug. Oh, 1938] (Up). Union Co.: P. Wilson S.n. (Plainfield, Sep mae 1901] (N). Warren Co.: Ahles 929/48 (Ur). PENNSYLVANIA: Bedford Co.: Berkheimer 1188 (Up). Chester Cos: F. R. Fosberg 15695 (Up). Clearfield Co.: 0. E. Jennings s.n. [aug. 25, 1925] (Cm). Clinton Co.: M. T. Travis 2801 (Up). Columbia Cos: F, R. Fosberg 15987 (Up). Delaware Co.: ok Je M. Fogg 10069 (Up); W We S Stone one 5572 (Up) (Up). Franklin Co.: A. R. "Moldenke 100 00 (Fg). Greene Co.: 0. E. Jennings son. [9/3/40] ak ere 2 Glowenke 581 a eS Lawrence Co.: Henlen 61 (Cm). Nor pton Co.: Jennings, percha & Gress s.n. {July 28, 1920] (Cm); R. L. Schaeffer 3327 (Ok). Northumberland Co.: P. R. Wagner 9063 (Up). Pike Co.: DePue 356 (Up). Schuylkill Co.: “P. Re Wagner “5852 (Up), 6036 (Up). Snyder Co.: J. M. Fogg 17907 (Up). Somerset Co.: Shetler 290/3 (N). Sullivan Co.: F. R. Fosberg 15940 (Up). Wash- ington Co.: L. K. Henry son. [9/13/40] (Cm). Wayne Co.: Jennings, Jennings, & Gress s.n. (Aug. 21, 1920] (Cm). Westmoreland Co.: E. 416 PHEeT OLOG.24 Vol. 11, no. 6 H. Graham s.n. [July 15--22, 1923] (Cm); L. K. Henry sn. [9/11/ WO] (Cm). MARYLAND: Allegeny Co.: R. C. Alexander s.n. (Allegheny, July '9] (Ca--379998). Garrett Co.: J. D. Smith s.n. [Aug. 15, '78] (W--1323136). Talbot Co.: E. C. Earle 3112 (Up). Worcester Co.: Raymond 197 (I). DISTRICT OF COLUMBIA: F. Blanchard s.n. [12 July 1891] (or—14309); H. T. Holm s.n. [19/7/1912] (I, N, Ur), sen. [1912] (S); P. B. Kennedy s.n. [July 27th, 1899] (Gg--18649). VIRGINIA: Bedford Co.: Curtiss s.n. [July 0, 1872] (Pu). Carroll Co.: H. N. Moldenke 19271 TL (N, | Or). Charles City Co.: A. R. Mol- denke 8h (Fg). Greensville Co.: Fernald & Long 10801 (cn, H— 68436, It It, Ok). Isle of Wight Co.: Fernald, Griscom, & Long 6674 (N). James City Co.: A. R. Moldenke 482 (Fg). Loudoun Co.: Leo- nard & Killip 891 (Ur, “W--1220059) . Nansemond Co.: Kearney 1739 (W=356L0h); A. R. Moldenke ),78 (Fg). New Kent Co.: A. R. Molden- ke 483 (Fg, S). Norfolk Co.: A. R. Moldenke 1,80 (Fg). Page Cost E. H. Walker 2654 (W-~183991), 2668 (W--1039505). Southampton Go.: Fernald & & Long 686, (Up). Sussex Co.: Fernald & Long 7588 (N, W—1699826). Wythe Co.: J. K. Small sen. Lease Greek, at at base of Lower Rock, July 23, 1892] rte). Lynchburg: L. Be Henderson 143) (Bt—-L,857) . Williamsburg: Grimes 379) (N)j A. Re R. Moldenke 485 (Fg). WEST VIRGINIA: Barbour Co.: E. L. Core 5957 (We). Clay Coes E. L. Core 6337 (We). Greenbrier Co.: R. B- Clarkson 2679 (We). Hancock Co.: Sumpstein Sen. [7/38] (We) ape ai Guthrie s.n, [July 18, 1953) (We). Mercer Co.: E. Dickins (We). Monongalia Co.: J. L. Sheldon s.n. tier =a/i3] (We); Shunk 3) (We); Strausbaugh 3h (We). Ohio Co.: R. West 686 (We). Preston Cos: Davis & Davis 766 (We); J.C. Myers 5u5 (Ur). Ran- dolph Co.: BE. E. Hutton Sn. ee Aug. 26, 1938] (We); A. H. Moore 3541 (Up). Ritchie Co.: E. L. Core 5672 (We). Summe Go.: M. CG. Beals sn. [Camp Brookside, August ist 13-15, 1958] (We); Boone 251 (We). Tucker Co.: A. H. Moore 2110 (Io-~131756)- Wayne Co.: Plymale 542 (Ky). Webster Co.: Hinkle s.n, (August 17, 1957) (We). Wetzel Co,: Haught 626 (We). Wirt Co.: E. A. BartholaneW 1935-77, in part (We). Wyoming Co.: E. L. Core "1760 (Bt--8559) - NORTH CAROLINA: Alleghany Co.: A. E. Radford 30401 (Hi--10b)43u) - Ashe Co.: Ashe s.n. [July 29] (Hi—59h79); D. 5S. Correll 3981 (H~ 40253). Avery Co.: Ahles & Duke 47326 (Hi—10L33). Bladen den Co. Ahles & Haesloop 29219 (Hi--10l430). Buncombe Co.: D. S. Correll 79 (B (H—27637) . Burke Co.: A. E. Radford 2556 pice a well Co,: A. E. Radford 1,988 (Hi--92887). Catawba Co. Correll 2927 Mame tal Correll & Oosting 28) aang “Sher ‘okee Co Cow: AJ E rd 17363 (Hi—110652). Craven Cos: A. Es Radford seit tesa 28). Currituck Co.: Ables & Duke 16221 1965 Moldenke, Monograph of Verbena 417 (Hi--104),38). Duplin Co.: Ahles & Leisner 33221 (Hi--104hh7). Durham Co.: Oosting 33128 (H--17867) . Edgecombe Co.: A. E. Rad- ford 36752 (Hi—104Lh8, S). Franklin Co.: Ahles & Leisner 20668 (Hi-~92736). Gates Co.: Ahles & Duke 52483 (Hi--110653). Graham Co.: A. E, Radford 14217 (Hi--92890). Greene Co.: A. E. Radford 36592 ( (Hi--104Lh9). Haywood Co.: Beaven 415 (Gu--10085); Blom- quist L7Lk (H~5456), sen. [Lake Junaluska, July 1, 1930] (Bt— 17342); Ne N. Hall 87 (F1—29037) ; A. L. Price 425 (N). Hertford Co.: Ahles & & Haesloop 52358 (Hi—-112963, Hi--11296)). Iredell Co.: Ahles & Duke 4.8761 (Hi--112966, Hi—112967). Jackson Co.: A. R. Moldenke 2 53 [H. N. Moldenke 21777] (Bm). Jones Co.: A. Es Radford 37277 (Hi--104450). Lenoir Co.: A. E. Radford 28878 (Hi—-927)0). Lincoln Co.: C. R. Bell 15303 (Hi—1h6190). Macon Co.t A. E. Radford s.n. (Aug. 2h, 1946] (Hi--352h9), s.n. (Sept. 1, 1948] (Hi--3512)). Mitchell Co.: Ashe sen. [Roan Mtn., July 1893] (Hi--59467). Northampton Co.: Ahles & Haesloop 52500 (Hi-- 112971). Pitt Co.: A. E. Radford 39635 (Hi—-lOMK71). Polk Co.: E. C. Tomsend s.n. [ouly 11, 1899] (P1--86891). Robeson Co.: Britt 2339 2339 (Hi~-177878) « Sampson Co.: Ahles & Haesloop 29971 (Hi—~-104h68) . Scotland Co.: Ahles & Haesloo 28791 (Hi-~10L469) . Stokes Co.: A. E. Radford 37762 (Ht (Hi~~10L470) “Swain Co.: D. S. Correll 3709 (H--40250) 3 J. Mooney s.n. [Qualla] (W--21778L9) 3 A. E. Radford 17012 (Hi—-927h3). Wake Co.: R. K. Godfrey 990 (Hi-- 10166, No— No—8600). Warren Co.: C. R. Bell 199 (Hi--92802). Wash- ington Co.: A. E. Radford 38761 ~ (Hi—-10446h). Wilkes Co.: Rad- ford & Stewart 1 1755 (Hi—16062). Yancey Co.: Ahles & Duke L6724 (Hi--I21520, fase). SOUTH CAROLINA: Bamberg Co.: Ahles & Haesloop 305 35 (Hi--104)31). Berkeley Co.: Ahles & liaestoep 30751 (Hi—10L432). Charleston Co.: Gibbes s.n. (Aug. 19, 1859] (i). Clarendon Co.: A. E. Radford 2821) (Hi--92733). ata Co.: J. H. Rice s.n. {aug. L, 1927] (Hi—5947h). i Co.: W. 6.6 Coker s.n. [July 8, 1909] (Hi--59465); B. E. Smi 1462 (Hi--26)77). Dorchester Co.: Ahles & Haesloop 37876 (Hi sr); Ahles & Leisner 31948 (Hi—10)26). “Florence Co.: C. R. Bell 13353 eee Georgetown Co.: Tarbox s.n. (N). Lancaster Cos: Ahles 3,786 (Hi—10h)75). Lexington Co.: E. A. McGregor 258 ats W—984754) . Oconee Co.: A. E. Radford 17859 (Hi— 92896). Orangeburg Co.: Ahles & Leisner 31725 (Hi—104473). Pick- ens Co.: A. E. Radford 16874 (Hi--90515). Saluda Co.: A. E. Rad- ford 26869 (Hi--927h7). Sumter Co.: A. E. Radford 27478 (Hi-- 927hL). 0 Union Co.: 0. M. Freeman 56527 (Hi—92801). Williamsburg Cos: Godfrey & Tryon 66 (W—10373uh). York Co.: Ahles 34508 (Hi— 104460, Hi--104)61). Witherspoon Island: B. E. Smith ith 163 (Hi-- 418 PHYTOLOGIA Vol. 11, no. 6 26:76). FLORIDA: Leon Co.: Rugel son, [Tallahassee, Mai 18)3] (Go). OHIO: Cochocton Co.: H. N. Moldenke 12529 (Co--535), 12777 (Co—-1012), 12989 (Co--1179), 13142 (Co—1239), 13981 (Co-2170, Da). Hamilton Co.: E. L. Braun s.n. [Anderson's Ferry, July 2h, 1917] (Cn); C. G. —- $.n. [July 24, 1882] (Pl--72007). ILI- INOIS: Alexander Co.: G. N. Jones 12031 (I1—15823, N , UE)s Champaign Co.: N. He Bell , 160 (Ur); Herb. Univ. Ill. 17466 (Ur); moe sen. (Ur—1767) ; Waite s.n. (Ur—-29663) . Coles Co.: G. D. Fuller 10700 (I1--21850). McLean Co.: G. D. Fuller 211 (I1=~ eazy 74hl (Ur). Moultrie Co.: Winterringer 9129 129 (11—39)06). Piatt Co.: G Ge N. Jones 1921) (Ur). County undetermined: Stark Se n. [8/9/23].(Dp). INDIANA: Bartholomew Co.: C. & Deam 36909 (Dn); Friesner 12936 (Bt--l2833). Blackford Co.: C. C. Deam 173 (Bt—-l2189). Decatur Co.: Friesner 7198 (Bt--17515). Fayette Co.: C. C. Deam 52508 (Dm). Franklin Co.: C. C. Deam 3697) (Dm). Grant Co.: cc C. Deam Deam 247) (Dm). Greene Co.: Friesner 5243 (Bt-- ia Hamilton Co.: Friesner 17250 (Bt—63h0); He Morrison Ss. [Noblesville, July 17, 1910] (Du--95)3). Hancock Co.? EK | se Téer-asiz} (Bt (Bt—~6152h). Jay Co.: C. C. Deam 50722 (Dm). Jeffer- Co.: Friesner 13148 (Bt—13663)._ Jennings Co.: Co.: C. C. Deam 37007 (Dm). Madison Co.: C. C. Deam 2462 (Dm). Miami Co.: Ek § n. Sars 20, 191) (Dp—2708), s.n. [July 1, 192] (pp——33116)- Monroe ¢ : Prisener 6003 (Bt-- t—-12739) . Montgomery Cow: Co Ce Deam 32801, (Dm). Morgan Co.: C. C. Deam 2261 (Dm). Orange Co.: Friesner 22803 (B, N, S). Parke ern sen. [July 20, '29] (Bt—3509). Porter Co.: C. C. Deam 39523 (Dm). Randolph Co.: C. C. Deam 63096 (N). Ripley Co.: C. C. Deam 58103 (N)- Scott Co.: C. C. Deam Deam 2802 (Dm). Vermillion Go.: C. C. Deam 53143 (Dm). Washington Co.: C. C. Deam 37137 (Dm). Wayne C0.: Ce Ce Dean 507k (Dm). white Co.: C. C. Deam 39321 (Dm). Whit- ley Co.: C. C. Deam 57147 (Im). IOWA: Allamakee Co.: L. He Pam- mel att sas Ce. 18, 1929] (Io~13399h). Boone Co.: Pammel, Buchan- 3920 (Io—-33058). Butler Co.: Whaley s.n. [July 2h, i518] (To-138h72). Cass Co: A. R. Moldenke nke 935 (LW (Lw, Ut). Chick- asaw Co.: Pammel & Spiker 683 (Io—1191jh). Dallas Co.: L. He Pammel s.n. [Dawson, Aug. 16, 1918] (Io—95630). Delaware e COs? Ae R. Mol Moldenke 981, (Lw). Floyd. Co.: Arthur s.n. [Charles City, July 15, 1872] (ah), s.n. (Charles City, 7/16/1872] (Ah), sn. [Charles re 7/15/81) (ah). Greene Co.: L. H. Pammel s.n. (Churdan, Jul- 29, 1928] (Io--132870). Hardin Co.: L- H. Pammel s.n. [Steamboat Rocks, 9-12-12] (Io--52671). Howard Co.: L. H. Paumel s.n. [Sara~ 1965 Moldenke, Monograph of Verbena hig toga, Sept. 2, 1919] (Io~-96236). Jasper Co.: Sipe s.n. (Colfax, Jul. 1922] (Io—-105826). Marshall Co.: Pammel & Fogel 075 (Io— 42398). Poweshiek Co.: Mrs. E. J. Hill s.n. [Montgomery] (Io-- 107762). Story Co.: Fawcett, Tener, & Reinbott s.n. [Slater Sept. 13, '02] (Io—h2578); A. Hayden s.n. [August 10, 1932] "(N); A. S. Hitchcock s.n. [Ames, July 82] (Io--15332). Union Co.: T. a An s.n. [Creston] (Io--22903). Winneshiek Co Goddard — S.n. [7-6-95) (Io—72018); Holway s.n. [July 16, 1881] ‘ (To=38762). KENTUCKY: Edmonson Co.: Lix 433 (W--1948073) . Estill Co.: Whar— ton 5186a (N). Marshall Co.: : Eggleston 48h1 (N). TENNESSEE: Car ter Cow: 1] D. M. Brown 137 (H--h0221). G: Greene Co.: A. = fae 55°(Hs N. “Moldenke 21779] (Bm). Obion Co.: Jennings S.n. [Reelfoot Lake, 192] (Cm). Roane Co.: Nease Ae aero) Sevier Co.: Wehmeyer 361 (Mi, N). MICHIGAN: Ingham Co.: L. He Bailey s.n. [Lansing, 1879] (Ba); M. Craig 12088 (Ca—367851) Menominee Co.: Grassl 2702 (Mi). Monroe Co.: B. Robertson 318 (Mi). WISCONSIN: Beloit Co.: Herb. Scient. Assoc. Beloit Coll. s. ne (Ws). Brown Co.: P. 0. Schallert 336 (H—5))81); Schuette s.n. [July 7, 1878] (N). Buffalo Co-: H. H. Smith 7180 (Wis). Fond du Lac Co.: A. R. Molcenke 1015 (Lw, Ut). Green Co.: E. W. Fell 58- 620 (Ws); Struik gen. [August 19, 1956] (Ws); stuntz s.n. (July 1s, 1891] (Ws). Juneau Co.: L. S. Cheney 3711 (Ws). “Lacrosse Cos: T. G. Hartley 1962 (W--2262066). Lafayette Co.: P. K. Nelson Sen. [July 23, 1946] (Ws). Milwaukee Co.: Hasse s.n. [iilwaukee] (N)j Lapham sen. (Ws). Outagamie Co.: F. C. Seymour 10153 (Ws). Polk Co.: Mrs. C C. F. Baker sen. [C. F. Baker 10693] (Po—6)652); Benner & Benner 105 (Ws); M. an Sn. sae (ine. 15, 1960] (Ws); R. W. Pohl 516 (Ws). Rock Co.: J» W 16 (Ws). Saint Croix Goer De Byle s.n. [12 Aug. 1952] aS Sauk Co.: J. H. Zimmerman 1188 (Ws). Sheboygan Co.: Goessl s.n. [Aug. 1919] (Ws). Wal- worth Co.: Almon s.n. [July 23, 1926] (Ws). MINNESOTA: Beltrami Co.: Westley s.n. [Black Duck, Jul. 22, 1918] (Io—9363). Free- born Co.: L. H. P Pammel s.n, (vrIr-28-" 27] (Io--130033). Goodhue Con: N. L. Te N Nelson aoe [Goodhue, Aug. 1899] (La, Ur); + Be Sandberg s.n. [July 1886] (B). Hennepin Co.: G. B. Aiton 8467 o--6,6 650); Cratty S.n. [Minneapolis, Aug. 1, 192h) (Io—115),10) ; i. H. Sandberg 597 (Fo—2281), s.n. [July 1890] (Ca—25175, Ca-— 25176 ~~ Mower Co.: J. F. Brenckle Ke 47-548 (N). Ramsey Co.: Mearns S.n. (Fort Snelling, Aug. 21, 1891] (W—670318), s.n. [Fort Snel- Aug. 25, 1891] (W—670319). Rice Co.: G. K. = Knight 3.N. 243). NORTH DAKOTA: Ramsey Co.: Geyer 105 (W--71962). Ransom 420 PHYTOLOGIA Vol. 11, no. 6 Co.: Bergman 1077 (Cm). Richland Co.: W. B. Bell 242 (Ca~-58776h), Lé5 (Gu-—21808); 0. O. A. Stevens s.n. [Leonar rd, “July 25, 1949] (W- 200420h) . Stutsman Co.: Lunell 869 (Ur), son. [Jamestown, Aug. 22, 1913] (N), s.n. (Jamestown, Aug. 2h, 1913] (W—-893166, W— 893167) . SOUTH DAKOTA: Bennett Co.: Visher 2263 (N). Clay Co.: Over 037 (W—78972), sen. [Aug. 5, 1915] (Se--1h494). Minnehaha Cos: P. Johnson 33 (Bt--62601, Spd Lb--36393, N, Ur). Roberts Co.: Over 14305 (W—-12)2263); ; H. Pammel s.n. [Sisseton, Aug. 10, '17] (Io—-9977) . pe Eg Over 16175 (Gg--130641). KAN- SAS: Riley Co.: F. C. Gates 12775 (Ob--508)9). MISSOURI: Greene Co.: P. C. Standley 9435 (W—688351). ARKANSAS: Newton Co.: Dem~ aree 222)0 (N, Z). Phillips Co.: Demaree 19745 (N, Sm)- NEBRASKA: Cherry Co.: J. M. Bates s.n. (Valentine, Aug. 16, 1893] (Ur). Knox Co.: F. E. Clements 2743 (W--202568). Thomas Co.: Rydbe: 1716 (N, W--210372). Washington Co.: Lallman s.n. [Aug. 26, 1926] (Bt--32236). OKLAHOMA: Canadian Co.: Me M. Hopkins ns 2099 (Ok). Chero- kee Co.: E. L. Little 528 (Ok); Stratton 6592 (N); C. S. Wailis 928 (St). Cleveland Co.: W. S. Myers 5 (0k). McCurtain Co.: Hop- kins & Cross 2296 (Ok). Muskogee Co.: Waterfall 10150 (St). Ok- lahoma Co.: Stemen 334 (Ok); Waterfall 1773 (Ok, St). County un- determined: Flowers 16 (Ok); Waugh s.n. (Clarkson, 6/22/1893] (Fe 26693). TEXAS: Cass Co.: Whitehouse 17697 (Sm). CULTIVATED: Germany: Bouché s.n. (Gg--31435). Switzerland: Herb. Hort. Bote Genev. sen. [18 Aug. 1826] (W--264565). LOCALITY OF COLLECTION UNDETERMINED: Mackaness s.n. [eastern N. Y. and e. Pa., 1936--37] (Tl). MOUNTED ILLUST OSTRATIONS: Perry & Fernald in Fernald, Rhodora 38: pl. 450, fig. 5-8. 1936 (N). ae URUGUAYEASIS Moldenke, Castanea 13: 118, nom. md. 1918; Synonymy poor melindres x tenera Osten ex Moldenke, Résumé 369, in syn. 1959. Verbena peruviana (L.) Britton x Vv. tenera Spreng. ex Moldenke, Résumé 372, in syn. 1959. Verbena KO 24 pd i Qo Vv. peruviana (L.) Britton ex Moldenke, Résumé 376, Bibliography: Moldenke, oem 13: 118. 1948; Moldenke, Known pon he Hacer Verbenac 2], 100 & 200. 19h9; Mo Moldenke, Ré- sumé 120, 369, 372, 376, & ith, 19595 Noldenke, Phytologia 8: 121 (1961), ur 289 (1964), and 1: 271 & Planta inte V. peruviana (L «) bie et V. “tenera Spreng. hy- brida naturalis, suspect foliorum florumque parentis intermedio. This is the na ural hybrid between V. peruviana (L.) Britton and V. tenera Sp eee. which occurs where the ranges of these two rina, Brazil, in Uruguay, and in the Aires provinces of Buenos ’ <- Catamarca, Cérdoba, Corrientes, La Rioja, Santa Fé, & Tucum4n, Ar 1965 Moldenke, Monograph of Verbena 4a gentina, Osten, in speaking of xV. trinitensis Moldenke, says: "Er ent- spricht ganz genau dem spontanen Bastard der Uruguay flora: Ver bena [Nobiles, chamaedryf.] Melindres Gill. (rot) x [Glandularia, erinoides] 'tenera' Auct. ae (Briq. -- an Sprengel?) der ebenf. rei bd rosa Blttten ha » dadurch wie obriger leicht unter den Eltern zufinden ist." 7 plant should have considerable ra yan gh 4 merit. xVERBENA VAGA Moldenke, Phytologia 5: 133. 1955. Syno: : Glandularia santiaguensis x megapotamica Schnack & Covas, Rev. Argent. Agronom. 12: 222, 22h, & 228. 1945. Glandu- potamica Spreng. ex Moldenke, Résumé me in syn. 1959. ke, bol. Abstr. 30: 1093. 1956; Moldenke, Am. ‘yaa . Nat. ‘93 362- 363. 1958; Moldenke, Résumé 225, 296, 369, 373, ba 420, 421, & L7h. Gt: Moldenke, Résumé Suppl. 2: ad 1960; Moldenk Phytologia : 121. 1961; Moldenke, Résumé S$ ppl, 8: 3 te 3. 196s wotdente, Pitotogta 20s 133 Sec we u: Pn3e, sis natura between v. Pe ae (Covas & asda eka iaeg Y: cisctagets Sheet: found originally a- Wild state there. They also grow in parts of ee yo I have not as yet seen any wild specimens of both province of that country « The hybrid should offer r considerable nanetoat tect promise and deserves more experimen Under xV. schnackii Moldenke, in these notes, it us stated that the two parental | species have a relatively small affinity based on the evidence of the cytologic pattern and the completely sterile pollen. This shows a good correlation with the large morphological differences between the two parents. Schnack & Covas 2 say “Asi mismo hemos podido identificar Sn el jardin experimental de la vas scuela de Agronomfa de Mendoza el hfbrido natural Glandularia santiaguensis x G. megapotamica; Ps ssi también otros hfbridos naturales que aun no hemos estud- do." NA VALERIANOIDES H.B.K., Nov. Gen. & Sp. Pl. 2: 277. 1818 {not V. valerianoides St.-Hil., 1947]. h22 PHYTOLOGIA Vol. 11, no. 6 Syno : Verbena valerianoides Humb. ex Spreng. in L., Syst. Veg., ed. 16, “2: 77. 1825. Verbena valerianoides Humb. & Bonpl. es Steud., roles Bot., ed. 2, 2: 751. 1841. Verbena valerianoides Humb. & Kunth ex D. Dietr., Syn. Pl. 3: 600. 18)3. verbo valer- ianoides Kunth ex Schau. in A, DC., Prodr. 11: 5h2. Ts Bibliography: H.B.K., Nov. Gen. & Sp. Pl. 2: 277. canis Steud., om. Bot., ed. 1, 874. 1821; Spreng. in L., mig tng ed. 16, 2: e, Known Se 10 Moldenke, Alph. List Invalid aint 1947; Moldenke, Alph. last cit. es ss7. & 573 (98) ype Ay: rd & regis et Mol- nke ve Distrib. Verbenac., [ed. 2], Be ts Ane, Résum 8 69 & 7h. 1959; ee résumé Pets: “he 20 (1962) and 6: 5, Bd amrust phe ogia 9: 97 & 11k. 1963; Moldenke, Résumé S Ass os Glabrous herb, ort 3 weak; stems erect, tetragonal; branches erect; leaves Aras By: posite, sessile, iinear, about ther at the alyx; flowers small; calyx short-tubular, about 3 mm, long, strigillose-scabrous the rim mucronate~5-dentate; corolla small, ng from light-blue, blue, or blue-violet to lavender-blue or ben slightly surpassing the calyx-tube. e "type of this rather distinctive species was collected by bb viakek, Heinrich Alexander von Humboldt and Aimé Jacques Alex- th the Botanisches Museum at Berlin, photographed there by Macbride as his type photograph no. 391,76, but now dest Schauer (187) cites also Hartw Hartweg 1350 from "In aAndibus Columbiae" in the same herbarium. He says "Proxima et persimilis praecedenti (V. litoralis H.B.K.] tamen uti videtur foliis integerrimis caly- ceque longiore satis distincta.” sole (1845) places the species in his Section Verbvenaca, Subsecti on Inermes, Group Foliosae, Sub- group Micranthae, and Secondary ibd oup Holophyllae, with 22 other species. The original authors cament "Habitu Valerianis nonnullis valde accedens." It has been found in wet fields, fields at the base of hills, and along streams, at altitudes of 2600 to 3250 meters, flowering in May, August, October, and November. Schneider found it "very 4 bundant in moist meadows and brook margins", Langenheim calls it "common in marshes", while Barclay & juajibioy 4 describe it as "growing w up through shrubs at edge of small st am in great stret- ches of open paramo with dried slopes and wet flats." CLEOME ORNITHOPODIOIDES L. ON VANADIUM-SLAG CANTON, BALTIMORE, MARYLAND WITH NOTES ON THE BIOGEOCHEMISTRY OF VANADIUM Clyde F. Reed (1) The author (2-6) has pea several lists of the plants found on the various ore piles in Canton, Port of Baltimore, wport ipa Pp having been found on chrome ore, iron ore or manganese o In 1962-63 a pie pile of ieee um-slag was dumped along Clinton Street in Canton, and on this pile in 1963 about 25 specimens of Cleome ater Senay Be 4 Linn. came up. Specimens Wisc.) were sent to Dr. H. H. Iltis, Department of Botany at the University of Wisconsin, where identification was verified. This species is new to the flora of North America species came up again in 1964, spreading to cover the pile rt 10 ft. in diameter and 5 ft. high). Seedlings a- bout 6 inches tall were porlge for study. Some of these plants were in flower and even in fruit with ripe seeds at this time. (June 23, 196), Reed 66675). The seedlings were potted in normal garden soil. However, the plants showed little growth for three weeks. Then, I added some of the fine vanadium-slag to the soil and watered it in. Within two weeks the plants were lected on the slag pile on August 14, 1964 (Reed 58183). Seeds from No. 58183 were sent to Dr. Iltis for further study. Cl Sone Se Linn. Native in the southern Balkans and the Near (Persia bor. et austr.; Tauria; Cau- casus; Poe Te pie wk "Smyrna; Russia). annual, up to 60 cm. bate stems glandular-pubescent; An an leaves trifoliate, hein 2.5 em. long, including the 1 cm. peti- ole; leaflets oblong- linear; finden white with a tinge of pink, a 1. Botanist, gic Research Division, Agricultural Research _ vice, U. S. Department of Agriculture, Beltsville, Marylan 2. Reed, ae at 56: 178-181. 1954. 3. Reed, Castanea 26: 123-127. 1961. 4. Reed, Castanea 26: 128. 1. 2+ Reed, Castanea 27: 59-61. 1961. 6. Reed, Phytologia 10(5): 321-406. 1964. 423 42h, PHYTOLOGIA Vol. 11, no. 6 about 2-4 mm. in diameter; sepals sometimes with a blackish margin; stamens 6, free at the base; silique glandular-pubes- cent, 2-2.5 cm. long, with 9-10 black, smooth seeds. The seed- lings start to flower about the fourth node above the ground. From specimens studied, collected on vanadium-slag pile in Canton, Baltimore, Maryland, in 1963 and 1964. Exact data re- corded above. Vanadium is not usually considered a micronutrient Shey ment for most plants. However, for Cleome ornithoposio Linn ems to accelerate growth. Cannon(7) reported t that Cleome integrifolia had up to 70 ppm. in the ash. e most complete review of the biogeochemistry of vana- dium until 1950 is that by Bertrand(8), in which the occurrence of vanadium soil, animals and plants is thoroughly surveyed and critically evaluated. A fine bibliography for the subject is en. veral more recent works evaluate e tolerance, re- Sistance, toxicity, pa ia oa ee and nutrient value of vanadium in various types of plants Biebl(9) studied the resistance of plant plasm to vanadium, using solutions of VOSO, in percent solutions from 20 to 0.0001. eee 7 tm punctatun = 3.0; Plagiochila asp- Tenioides - 3.0; Bazzania trilobata - 10.0; and Trichocolea tomentella - 5.0. ng our cultivated erate of economic value Solanum ould tolerate solutions 0.0001 les pt 0.001; Rheum rhabarbarum, and Brassica oleracea var. re up to 0.01 Among some weedy plants tested, Polygonum lapatnifoLium, Atriplex patulum, Mercurialis annua, Avreps pels aoenee Solanum nis “= annuus 7. Cannon, Helen L. Effect of Uranium-Vanadium Deposits on the Vegetation of the Colorado Plateau. Amer. Jour. Sci. (10): 735-770. Oct. 1952. 8. Bertrand, Didier. Survey of Contemporary Knowledge of Bio- geochemistry 2. The Biogeochemistry of Vanadium. Bull. er. Mus. Mat. Hist. 94: 403-456, tables 1-5. 1950- 9. rier Richard. Ueber die Resistenz Pflanzlicher Plasmen g°6°? Vanadium. Protoplasma 39(2): 251-259, 2 tables. 19 1965 Reed, Cleome ornithopodioides 425 and — ope the could withstand concentrations up to 0.001 ntal pur- percent. Among the plants usually cultivated for ornamen ses, Begonia scharffiana (0.000 clamen persicum (0.001), Iris lida (0.01), Iris florentina (0.001), — — Me oer Iris trojana (0.01), Rhoeo =a (de Trad a 0.001) Tradescantia zebrina (0.0001) and aeoinlentiece aie a (0.001) were resistant to soli solutions up to the given percentages. Suzuki(10) found that the ig of vanadium in a culture (e]@) di the fruits became more sour. Chiu(12) found that growth was in- creased in rice seedlings by application sery beds of solutions of de, with the best results at 150 p to 0.001 ppm. of vanadium was added to the culture solution. Among the lower plants vanadium plays an important role in the nitrogen fixation of Azotobacter species (14-17). The data 10. Suzuki, S. On the action of vanadiun compounds on plants. Tokyo Bulls; Agr. 525L5e 1903s An ae Kumar, K. and S. B. Lall. Effect of vanadium on chemical composition of tomato fruits. Indian Sci. Cong. Proc. 40(3): 118. 1953 (1954). 12. Chiu, T. F. The effect of vanadium application on the paddy rice. Agr. Res. 4(2): 48-59. Sept. 1953. 13. Arnon, D. I. Microelements in culture solution experiments with iiaber plants. Amer. Jour. Bot. 25: 322-325. 1938. fo i - Konishi, Kametaro and Tsuge Toshihisa. Effect of inorganic constituents of soil solution on the growth of Azotobacter. Jour. Agr. Chem. Soc. Japan 9: 510-520. 1933. 15. Burk, D. and C. K. Horner. The specific catalytic role of molybdenum and vanadium in nitrogen-fixation and amide utilization by Azotobacter. Internat. Soc. Soil Sci., 3rd Congress, Trans. 1: 152-155. 1935. 16. Bore, J., Bore, C. and D. I. Arnon. Molybdenum and v ium requireme nts of Azotobacter oe growth and nitrogen pibattnes. (Abs.) Plant Physiol, 32(Sup.): xxiii. 1957. 26 Pot -T05 0-658 Vol. 11, no. 6 pertinent to the effect of vanadium on nitrogen-fixing organisms is reviewed by Nason(18). Jensen seek Spencer(19) studied the in- fluence of vanadium on the nitrogen fixation by Clostridium buty- ric and related organisms. Sampath(20) = abe that aces See in yeast cells was inhibited by 260 ppm. of vanadium, but wa quite favorable with 140 ppm. Bertrand(2l, 22) showed as vane dium acted as a growth factor in concentration of 2 x or Aspergillus niger. Allen(23) found that vanadium could not “a substituted for the molybdenum requirement of the blue-green alga, Anabaena. are green algae, Arnon and Wessel(24) found that there was opie increase in the growth rates in si green alga, Scenedesmus obliquus, upon the place addition of vanadium the purified nutrient solution. Later, Arnon(25) lees that 17 is H. Species differences in molybdenum i vana-~ dium r ie and combined nitrogen utilizatio [eatouesteetactan: Plant and Soil 16(2): 171-20L. oe 1962. 18. Nason, Alvin. The metabolic role of vanadium and pears in plants and animals. Chapter 19 in Trace Elements 64~ 296. 1956. 19. Jensen, H. L. and D. Spencer. The influence of molybdenum d vanadium on aaa fixation fod oe aye and related organisms. Linn. Soc. N. S. Wales Proc. 73-86. 1947. 20. Sampath, S. Effect of vanadium on yeast cells. Current Sctence: (India) 13: 47. lob. el. Bertrand, D. Vanadium in plants. Bull. Soc. Chim. Biol. 23: “6T 471. 191. ie ere iinc vanadium comme element are pour Soper rill niger. Inst. Pasteur (Paris) Ann. 68: 23. Allen, M. B. Sci. Monthly 83: 100-106. 1956. eh. Arnon, D. I. and G. Wessel. Vanadium as an essential ele- ment for green Sa Nature (London) 172: 1039-1040. 1953- 25. Arnon, Daniel I. The role of micronutrients in plant nutri- tion with special reference to photosynthesis and oar 2e assimilation. Trace Elements, Proc. Conf. Ohio Agr- Stat., Chap. 1. 1958. 1965 Reed, Cleome ornithopodioides 427 the maximal level of the Hill reaction in photosynthesis by isolated chloroplasts was raised by the addition of vanadium. Bechi(26) was the first (1879) to mention the existen of vanadium in the ashes of plants. ere is considerable aabiie ature dealing with the methods and techniques for testing the amounts of vanadium in soils, and in plant and animal tissues. I shall only indicate some of the critical species which seem to have high percentages of vanadium in their tissues. Among the fungi in 17 species tested, Anan itope* muscari showed from 60.8 to 181 ppm. in seven lots tested. Other “species of some note are: Lactarius tomninosus (12), Amanitina loides (2.4) yes) eae ee Amplariella spissa (1.0 ppm. ) and Bere solitaria (1.8 ppm.), as Soe by 5 Be asE TT Czapek(27) found vanadium in the ash of red beets, quant beets, grape vines, F Ss, Quercus, Picea, Abies and inu d he ntai £ oxide). Among the plants studied, Astragalus oiterininnt Gea Rica ), A. reussii arctus (3000 ppm.), A. see Se | opsis hymenoides (70 to 690 ppm.) a and Stanleya arcuata ss 13 al also store up large quanties —a se ane Other plants that stored vanadium were: At riplex confertifolia (10-90 ppm), A. canescens (25-30 ppm), Bahia nudicaulis (70 ppm.), Chryso- thamnus viscidiflorus (50-200 ppm.), Artemisia spinescens (70 1 bat mant period; 400 ppm. during the active period). In some trees Shrubs studied, the following ppm. of vanadium were observed; Ephedra viridis (70 ppm), Fraxinus anomala (5 ppm.), Juniperus monosperma (10-60 ppm.; 1¥0-4000 ppm. in roots), Pinus edulis —— 10 ppm). and Quercus —- (5- 90 ppm.; 1700 ppm in oots). A Spirogyra sp. showed 500 p a 26. rete E. Nuove richerche del st e del vanadio. Atti Roy. ccad. Lincei, III, 3: 186. 1879 27. Czapek, Fr. Biochemie der Pflanze, 2 Aufl. II. Bd. 1920. Jena. 428 PHYTOLOGIA Vol. 11, no. 6 Bertrand(28-30) reported vanadium in every sample of 62 species of plants studied. Most of the fungi had less than 0.5 nad most legumes had 3-4 ppm.; Vicia faba root nodules had 12 ppm. vanadium. Bertrand(8) presented a a correlation study of the vanadium intake for six lots of legumes Soil Roots Nodules Aerial y-vanadium pm. m Kidney bean 19.4 38.0 27.2 5.6 Black Tokyo 23.7 27.6 32.6 5.2 Soya Yellow lupine 54.3 24.2 5 12.8 White lupine 5h 3 56.0 20.4 5.3 Vendee White lupine 23.7 25.8 27.8 2.97 Versailles Broad bean 32.0 75.0 48.5 ie Demarcay(31) reported vanadium in the ashes of Pinus sy - vestris, Picea, Quercus, Vitis, Populus and Carpinus. Robins studied, and reported most notable: kidney beans, sugar pine needles and clover. However, “‘Dimetriev (33) found ina yaa 28. Bertrand, D. Recherches sur le vanadium chez les végétaux. Inst. Pasteur (Paris), Ann. 68: 58-68. 1942. 29. Bertrand, D. Distribution of vanadium in plants. Bull. Soc- Chim. Paris, V, 9: 121-133. 19h2. 30. Bertrand, D. Vanadium in fungi, especially Amanita. Bull. Soc. Chim. Biol. 25: 194-197. 19h3. 31. Demarcay, E. Sur la présence dans les végétaux du vanadium, molybdénem et we chrome. Compt. Rend. Acad. Sci. Paris, 130: 91-92. 1900. 32. Robinson, W. 0., L. A. Steinkoening and C. F. Miller. The ar pie vd rng of the rarer elements in plants a ar B U. - Agr., No. 600: 1-25. 1917. 33. Dimetriev, K. S. The effect of microelements in limed nen soils on the development and the crop of red clover. Pedole st peat No. 4: 114-133. 1939: Klim. Referat. Zhur., No. 1939. ce aelan 1965 Reed, Cleome ornithopodioides 429 had no effect on the development and growth of red clover in limed podzol soils of Russia. The presence of vanadium and other i not this and other oo are found only in relation to vanadium- carrying soils in Russia ium has been studied in the soils of Argentina by sites 2 om Amato(36), and its action on plant growth in Argentina nad < he also found tha plants that grew in the lava of this volcano contained eet quantities of the element. 34. Vinogradov, A. P. and Bergman, G.G. Chromium and vanadium in soils of the Soviet Union. Pochvovedenie, ioe, 569-573. Oct. 1949. 35. viagra E. V. and Levin, F. I. Content of vanadium, chro- mium a 8 n N. S. Avdonin, Povyshenie plodorodiya pochv nechernozemmoi polosy 1: 69-79. 1961 36. Trelles, R. A. and Amato, D. enico, vanadio y molibdeno Ars €n suelos y en algunos se ota de la Republica Argentina. Soc. Cient. Argentina An. 149: 93-107. 1950. 37. Ramirez, E. C. Vanadium and its action on plant growth. Am. Sei. Quim. Argentina 2(6): 145-146. 1914. 38. Nakamura, M. T. and Sherman The vanadium content of Hawaiian. Island soils. en — Expt. Sta. Tech. Bull. 45: 1-20. 1961. 39. sige L. Sulla diffusione del vanadio nel regno minerale vegetale. Atti Accad. Gioenia di Sci. Nat. Catania 17: 161-166, 1883. ko, Shibuya, K. and H. Saeki. Effect of vanadium on growth of plants. Jour. Soc. Trop. Agr. Japan 6: 721. 1934. 430 PHYTOLOGIA Vol. 11, no. 6 Shibuya and Saeki(40) studied the effect of gone on the growth of plants in Japan. While studying the effe molybdenum on plants in natire; Ter Meulen(41) found ik inant muscaria had 3.3 ppm. vanadium, garlic had 0.8 ppm., but tha onions had none. Although there is still no proof that vanadium is essential micronutrient element in the plant kingdom, thebe is t x some evidence that some lower plants and in a few higher plants, vanad ps in the physiological activities of those plants. re traces seem to accelerate growth activities, as Scenedesmus, Aspergillus, Azoto er, and various higher plants. Waringto - s ied the interrelationships between um and other microelements in the nutrition of soybean, flax, oats, eas, and has shown that there is an increase in the toxicity of molybdenum with an increase of va es pecially in soybean and fl at least when usin olutions of ie) up 1 Mazé and Mazé(45) indicated that vanadi was favorable to corn, but did not prove that it was a necessary nutrient element for good growth in that plant. Our studies of the effect of vanadium salts on the g of microorganisms and the higher plants started with the eo a. Pas b Ter Meulen, H. Sur la repartition du molybdene dans 1 nature. Rec. Trav. Chim. Pays-Bas 50: 491-504. 1931. he, Warington, Katherine. Some interrelationships between man- ganese, molybdenum and vanadium in the nutrition of soybean, flax oats. Ann. Appl. Biol. 38: 624-641. 1951. 43. Warington, K. The influence iat iron supply on toxic effects of manganese, molybdenum and vanadium on es peas and fl Ann. Appl. Biol. 41: Nes 9 4k, Warington, K. Inve ipa iveraen regarding the nature of t teraction. between iron and molybdenum or vanadium in nu Solutions with and Scnect a growing plant. Ann. A Biol. 44: 535-546. 1956. e in- aarp 45. Mazé, P. and Ma zé, J. Recherches sur la nutrition mineralé des végétaux esi fal Compt. Rend. Soc. Biol. 132: 375-377. 1939. 1965 Reed, Cleome ornithopodioides 431 Witz and Osmond in 1886(46). The concensus of opinion from the investigations up to date is that vanadium in concentrations up to 10 to 20 ppm. seems to be stimulatory, whereas higher concent- rations prove toxic to most higher plants. Cleome =e Linn. nl to note what other plants move in on the vanadium-slag pile in Canton. -—__ 46, Witz, G. and Osmond, F. Essai sur l'application des pro- priétés de 1' pxyoelluiocse au dosage du vanadium. Bull. Soc. Chim. Paris 45: 309-314. 1886. BOOK REVIEWS Alma L. Moldenke "Orchids of the Western Great Lakes Region", by Frederick W. Case Jr., xii+ 148 pp., illustr. Cranbrook msi of ackunte, Bloomfield Hills, Michigan. 196. $7.00 The Cranbrook Institute has rendered another valuable ig to science in publish hing this attractive, useful and accura survey by an author who has carried on years of aoe = nar work, systematics, and culture studies to these fascinating p The book begins with a brief description of the Orchidaceae, an re) our ds have migrated from — regions since econ nolan Sion about 10,000 years ago he book continues with excellent keys to genera, with descrip- tive text for each. Then there are clear cut keys to each species each Siesta with critical line drawings and each Necriet a0~ 432 PHY SOLOGIA Vol. 11, no. 6 cording to habit, season, habitat and distribution. These keys would certainly offer no difficulty to any amateur. Also for each species the author provides either an exquisite color photograph or very fine black and white ones, all in natural habitat, and geographic distribution maps. What a wonderful aid this book would be for a field trip in this area, or even just an armchair tourt "The Gardener's Fern Book", by F. Gordon Foster, xiv + 226 ppe, illustr. D. van Nostrand Company, Princeton, New Jersey. 1964. $7.95 Here is another fine book for the amateur naturalist, and outdoor garden hobbyist and horticulturist (May their a increase so that our sojourn on this earth may be more happily satis and the need for the conservation of our diminishing natural resources be better understoodt). It is written and il- ed a @ 2 5S cc) a & bh 6. 4 o pen ww 0g ® ctr fi 4 ing of hardy ferns for the purpose of growing them in home gardens, woodland plantings and indoors, ~— methods of fern propagation with a set of excellent diagrams (7) occupying the bulk of the sag forty hardy and twenty-seven gegen bel pata described critical- sori, culture ere are page dra one, The book ends with four useful features — comparative c » Glossary, list of commercial growers, and index. The specific name of Matteuccia pensylvanica is capitalized 0 anh 63, but, fortunately, is lowercased in the remainder of the Xt. PHYTOLOGIA Designed to expedite botanical publication Vol. 11 March, 1965 No. 7 CONTENTS REED, C. F., Lobaria quercizans on the Coastal Plain and Piedmont Plateau in Maryland and Virginiad.......++++.+++5- 433 MOLDENKE, H. N., Materials toward a monograph of the genus Verbena, XX IX PO Oe i eee Pee Perr Eee a eer eee 435 MOLDENKE. A. Li. Bock reviteg. 625 5 o.6s bi cee Ss bs vee es bed ae 507 Index to Authors in Volume Eleven ......-.+0eee0ee05 dese tuew se 508 Index to Supra-specific Scientific Names in Volume Eleven .......- 508 Dates of Publication of Volume Eleven ....0.eeerereceneercceere 512 Published by Harold N. Moldenke and Alma L. Moldenke — 15 Glenbrook Avenue Yonkers 5, New York, U.S.A. Hv Price of this number, $1; per volume, $5.75, in advance, or $6 at close of volume Vol. 11, no. 5, was issued February 6, 1965; Vol. 11, no. 6, was issued February 17, 1965 Misen: try Poranremg MAR «¥ 1965 LOBARIA QUERCIZANS MICHX. ON THE COASTAL PLAIN AND PIEDMONT PLATEAU IN MARYLAND AND VIRGINIA Clyde F. Reed LOST Halel/ reviewed our knowledge regarding ips distri- bution, of the Lobaria amplissima-L. quercizans complex in Europe rth America, stating that "L. quercizans in North America, er a tpetank Appalachian-Great Lakes distribution pattern, where 1G As Fogo abundant in mature conifer-hardwood forests with Lobar ulmonaria." His accompanying map indicates localities from apaeaied and Nova Scotia, through New England and Pennsylvania into the Blue Ridge region of Maryland, Virginia, North Carolina, South Carolina into Georgia and Alabama; then westward. There are no localities indicated for regions east or south of the Fall Line (geological divide between the Piedmont Plateau and the Coastal Plain). following specimens of Lobaria gquercizans Michx. repre- sent collections of this species from the Coastal Plain areas of ry. » the Delmarva Peninsula and Tidewater Virginia, and fr the outer areas of the Piedmont Plateau in Maryland and Virginia. Maryland: Anne Arundel Co.: Base of tree trunk, Bent near Benfield, near Head of Severn River. Dec. 16, 1915. ae, fi Sw.) an lobaria pulmonaria ios Hoffm. nize collected near here: On tree trunk between Benfield aw Calabi oe ahh c.c. Plitt 37 (USNFC and Reed Herb.). Baltim Co.: On base of tree, ddle River Neck. Dec. 30, 1905. C.C.Plitt See (USNFC); on tree in ravine near Camp Run. Oct. 3, oe eae C.0.Flitt 2 (USNFC). ‘This area is northeast of Baltimore, very near the Chesapeake Bay. Middle River is a tidal river on the Coastal Plain of Baltimore Patapsco River. Dorchester Co.: Low wo along Rt. US 3/50, oak, just N of Salem. April 12, LAN c. a Reed eyes. This ste lies just west of the Nanticoke River Virginia: ee a At base of oaks in low woods, Rt. 632, off Rt. us Ji60, Myrtle. April a6; 1963. C.F.Reed 6192h., This area is pe yin: of the Dismal Swamp. 1/ Hale, Mason E., Jr. Bryologist 60(1): 35-39, map. 1957. 433 43h PHYTOLOGIA Vol. 11, no. 7 the edge of the Piedmont Plateau, the following pig of Labaria ae = izans Michx. and Lobaria pulmonaria (L.) Hoffm. y be re faryland: Baltimore Co.: Lobaria Se ie Tao Pein tree, along Gunpowder Falls. 7210. oe a eae ua. inia: Charlotte Co.: eae ee on hickory, 2 mi. W ey Rayertiiss Virginia Rt. 720. Oct. 27, 1962. C.F.Reed a9722 The specimens cited above have been identified or verified by Dr. Mason Hale. They extend our known range for these re) species will be looked for more deligently. n my studies itis the Flora of Delmarva and Tidewater a ge these lichens add an other group of species usually found on the Piedmont Plateau or ain. Over 450 species of plents may now be recorded from iso- lated 'islands' on the Coastal Plain mre are otherwise known to be Piedmont and {ppalachian speci This the not too far. from ae oie River and the Soldiers Delight area. MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XXIX Harold N. Moldenke VERBENA L. Additional synonymy: Canadea Patermann, Beitr. Zytol. Verbenac. 13; Bet [Su], pl. 1, fig. 1& 2. 1935. ional & emended bibliography: Rtfling, Ord. Nat. Pl. rade wl; L * C. Rich., Act. Soc. Hist. Nat. Paris 1: 105. 1792; Ann > cy Hist. Nat. Paris 15: pl. 14, fig. 1~5. 1810; Kunth, tnd. - Hort. Berol. 1845: 7. rt = ai Ann. Sci. Nat, see 10: 319-381. 188; Sc hlecht., Ann . Nat., ser. 3, lh “3h. echt. 23: 7b 732 1850; R. A. peli, i Chil. 35: 190-192 18705 pa ace Bot. Notiser 6: ‘9-387. 1881; Hieron., Bol. Acad. daph. Fl. Argent 213-~215. 1882; Cockerell, Canad. Ent. 28: 158. 1896; Cockerell & Porter, Ann. Mag. Nat. Hist. (7 ) ks ie Rts, Cockerell & Atkins, Ann, Mag. Nat. Hist. (7): 10: hh. 1902; Crawford, Canad. Ent. 3h: 240. 1902; Charany, Journ. de Chim., ser. eye 2: 292--298. 1910; Lond, Chem. Soc. Journ. 98: 991. 1910; Chem, Abstr. 5: 2525. 1911; Robertson, Ent. News 25: 72. 1914; Rau & Rau, Journ. Anim. Behav~ ior 6: 368. 1916; Alvarez, Fl. Santiago del Estero 106. 1919; Kan- da, Bot. Gaz. 69: Sh--71, pl. 6--9. 1920; Robertson, Psyche 29: 171. 1922; Rau, Trans. Acad. Sci. are Louis 2h (7): 22. Pg 8 eee dst. se sa ae 7h: LOm-h5. 1925; Schnar i ‘ O (2): 6h, 2a 233, ane, IOs 3 688. 1929} Saeko Zytol. tuscan’ Verbenac. rb, 17-27, & 31. 19313 Patermann, Beitr. Zytol. Verbenac. 6-8, 1 ap CE 43, 7, & harmak (Sh], pl. 1, fig. 1 & 2. 1935; Bider, ce co . Borage 95——1 : ae) ae oe | 2 oskcemige & Evenari, Bull. Soc. Bot. Genév. 31: ° ~~ cg a oo orul. Cisiord. 363. 1948; Parks & King, El Crisol Tearto Rico) h: 7h-~ 80, 1950; Chen. Abstr. 5: 3037. "195; acaba k, Townes, U. S. barks Agr. Agric. Monog. 2 [Hymenop. N. Rie 1951; He Ce. Greene, Midl. Nat. 8s 37—5h. 19523 H. C. Gre ene, Biol. Abstr.” zr, "1885. 1953; Acevedo de Vargas, Rev. Chil. Hist. Nat. 54: h5. 1954; Moldenke, Biol. Abstr. 27: 98h, 2026, & 3121. 1 Santapau, Fl. Purandhar 103, 10, 158. 19573 Algonquin Prov. Mus. Check-list 19. 1957; Mufloz Piz » Sins F i —~200, pl. 96 c—e & 97. 1959; Reitz, Sellowia ne a: 51, 79, & 13h--135 (1959) and 12: 152 & is7. 1960; G. Rzedowski, Cie Potos. § - 1960; T. Be Mitchell, Bees “a U. S. [N. C. Agr. » Sta. Tech. B . 141): 292. 19603 M. Le Buch, U. S. D Agr. ey Handb. 6: 13. 1960; Mufioz Pizarro P List 8. 1961; Jones Tice & Johnston, Flow. Pl. & Ferns Tex. Coast. Bend 1h3-lhh. 1961; Twin, Roadside Flow. Tex. 189-190, 43 436 PHYTOLOGIA Vol. 11, no. 7 ta. 10: 50 & 65. ; Pinkava, Ohio bis Sei. rne in T. Swain, Chem. Pl. Tax. 372. 1963; Steyerm., Fl. Mo. 1257=-1261, pl. 302 & 303. 1963; Shelford, Ecol. N. Am. 77 & 607. 19633 Ge Ne Jones, Am. Midl. Nat. Monog. 7: [Fl. Ill., ed. 3) & Mat Gar tac ye Bo ash, 196 ete 3) ol 358—360, 372, 376--380, & 38h, pl. 3 & aes denke, Phytologia 8: 460-196 i565); sf "Si ior (1963), “to: ‘rok 91--115, 117~125, 127--1h43, 15-155, 157—161, & 490--50h ( ’ and 11:'1—68 & 72, fig. 13 & lh. 19643 A. & Ie Ne 131 & 15— 1964; Langman, ect. Guide Lit. Glow. Pl. Mex. 208, 329, 5 517, "580, 680, 73 & ape $ ea Menninger, Seaside Fa Pl. 35, 51, 54 265, 269, - 1964; Soukup, B cA 196h1; Seismic, résumé suppl. 10: 1, 2, yan & 7 (296k) and ll: 1—3, 1965 Moldenke in Shreve & Wigg ins, Veg. & Fl. Des. 2: 1235-1246, ae & 1738--1739. 1964; A. *1L. Moldenke, Phytologia T o Mel? Yonkers +o Ds Hoc 4Sh—)56 » 196k 5 Anon., Biol. A beta. be (2): 37130. “19055 : hisiaen, Phytologia 11: 219-286, 290--357, & 400--l\22. 1965; Anon., T a. 5 (5): 2. 19653 Anon., N. Y. erdia Trib., sect. 2, Pp. 19, Feb- » 1965. ” ‘addenda to list of excluded species: 6 Verbena suffruticosa reclinata foliis angustis serrato—dentato, pedunculis longis solitariis, floribus ribus conglobatis P. Bre Phyla stoechadifolia (L.) aa. Verbena undulata Reitz = Lantana undulata Schrank Addenda to list of abbreviations employed for the names of herbaria whose material is cited: Cd = Museo de Cérdoba Syaticnay: Argentina Fj = Fred B. Jones Herbar , Corpus Christi, Texas Ij = Science Museum, riptieate of Jamaica, Kingston, Jamaica Lb = University of meretes | Columbia, Missouri Sj = University of Puerto Rico » San Juan, Puerto Rico 4 Ww = Rob and Bessie cat a Wildlife Foundation, Corpus Christi, mining (1774) brackets Verbena and Phryma together and says tion Pediculares. In Alvarez (1919) se — name is in some places uppercased and in some places lo ased. Ver- Muesebeck, Krombein, & Townes (1951). pa that the genus "=" 1965 Moldenke, Monograph of Verbena 437 bena is regularly and apparently exclusively visited by the bees Calliopsis hirsutifrons Cockerell (C. verbenae Cockerell & Porter] and Ce. nebraskensis Crawford {C. verbenae var. var. nebraskensis Craw- ford]. Soukup (196) records the common names "azulina", "huallkjapaya” huayfu- ; hat, © a ys tratamiento de la terciana; la cataplasma de las hojas en la su- puracién del higado, los polvos bebidos en vino aflejo en = to e." ea from Verbena flowers which they called "pleasant-dream and apparently used it to quiet nerves and induce sleep. This may have been the first tranquilizer used in North America’ Spector (1956) says that this is one of fifty plants that are host in North America and Europe to the chrysanthemum nematode (Aphelenchoides ritzema-bosi), which is a vagrant endoparasite of e low concentrations of 2 ae pea slightly depending on cies involved NA ABRAMSI Moldenke Additional bibliography: Moldenke, Phytologia 10: 91 & 10 (196) and 11: 105. 196). XVERBENA ADULTERINA Haus Additional Hee fanentan “Yoldenke, Phytologia 10: 91, 210, 212, & 219. 196k. VERBENA ALATA Sweet Additional bibliography: Reitz, Sellowia 13: 67 & 110. 1961; Moldenke, Phytologia 10: 91, 122, *& 160 (1964) and 12: 121. 196h. VERBENA AMBROSIFO Rydb. or ing tu bibliography: Moldenke, Résumé Suppl. 10: 2 (196) 196; Moldenke, Phytologia 10: ee (1964), 11: 32, 3, & zie. (196h) , and 11: 273, 290, & 326. 1965. Jackson reports this species as very abundant along roadsides adalupe County, New Mexico, The Tharp & Janszen 49-110 dis~ tributed as this species is actually V. wr wrightdi A. G » Gray. D. Be Dunn 3050 is a mixture with V. bipinnatifida var. latilobata Perry. ~~ Additional citations: KANSAS: Douglas Co.: Wilson & Miller s.n. [6-15-12] (Iw). Linn Co.: B. F. Bush 15503 (Lb—-23431). OKLAHOMA: Se NR Logan Co.: hardt 940 (W-~2421207) . “TEXAS: Culberson Co. Tharp 1,38 PRTTOLOG@?TS Vol. 11, no. 7 & Janszen 49-1169 (Lw). NEW MEXICO: Guadalupe Co.: S. W. Jackson 12-1 =1 (Lw). San Miguel Co.: D. B. Dunn 6595 (Lb—-l1601). Socorro Co.: D. B. Dunn 3050, in part (Ib--)8])31). UCANA R. A. Phil Additional bibliogr. raphy: » Espec. Plant. Descr. ons aah 110. 1960; Moldenke, gee oeming io: 93 (196) and 11: 8 The Macbride photograph 17400, cited below, is of a specimen that was ee ed in the herbarium of the Botanisches Museum at sasiewd now destroyed. tional citations: CHILE: Linares: R. A. Philippi s.n. [Lin- on aapreis photos 1700] (N--photo). ENA ARISTIGERA S. Moor Additional bibliography Moldenke, Phytologia 10: 93 (196k) and 11: 28) & 290. 1965. VERBENA ATACAMENSIS Reic Additional Rene. ste. Moldenke, Phytologia 10: 93 (1964) and ll: 2h5. 1965. XVERBENA BAILEYANA Moldenke Additional bibliography: Moldenke, Phytologia 10: 93 & 210. 196 VERBENA BAJACALIFORNICA Moldenke Additional bibliography: Moldenke in Shreve & Wiggins, eke & Fl. Son. Des. 2: 1245. 1964; Moldenke, Phytologia ll: 32. 19 Langman, Select. Guide Lit. Flow. Pl. Mex. 516. 196h. VERBENA BALANSAE Briq. Additional bibliography: Reitz, Sellowia ll: 57 & 134 (1959) and 13: 67 & 110. 1961; Moldenke, Phytologia 10: 93 (1964) and 11: 312 & 313. 1965. RBENA BARBATA Grah. Additional bibliography: Moldenke, Phytologia 10: 93. 196h5 Langman, Select. Guide Lit. Flow. Pl. Mex. 329 & 1010. 196h. VERBENA BERTERII (Meisn.) Schau Additional synonymy: Glandularia berteroi Mufioz Pizarro, Sin. ohttticoal bibliography: Mufloz Pizarro, Sin. Fl. Chil. 199 & - 19593 Moldenke, Phytologia 10: ye (196k), “hh ol, 52, abs, 4 "89 (1964), and ll: 245 & 2h6. 1965. ” addi tional illustrations: Mufioz Pizarro, Sin. Fl. Chil. pl. 97- It should be noted here that the Macbride photograph 7853, ci~ ted ee is a photograph of two specimens in the rae gert Her- barium a t the Conservatoire et Jardin Botaniques in Additional citations: CHILE: Colchagua: Bertero Th “itacbride 1965 Moldenke, Monograph of Verbena 439 photos 7853, in part] (N~-photo of isotype). Valparafso: Bertero 1389 [Macbride photos 7853, in part] (N—photo). ae BIPINNATIFIDA Nutt dditional synonymy: Verbena a Nutt. ex Moldenke, sas Suppl. 10: 7, in syn. 19 Additi onal esis Sereciey$ “Irwin, Roadside Flow. Tex. 189, pl. 1; Jones, Rowell, & Johnston, Flow. Tex. Coast. Bend 143. ; 5 Was Wis T25r, 1 263, map 1846, pl. 303, fig. 3. 1963; Moldenke, Résumé we 10: 1& 7 (1964) and ll: h. 1964; Moldenke, ge he oa 10: 94-95, 217, 282--28h, 306, & 307 (196k), 11: 13, 25, 32, In, 3, ca & 48 (1964), and il: 290, 308, & 32%. 1965. Additional Pnstratidiie: Irwin, Roadside Flow, Tex. pl. 39 (in color]. aon Steyerm., Fl. Mo. pl. 303, fig. 3. 1963. The species has been found by recent collectors in pasture openings var dry rocky pr Le patois exposed areas of dry prai- rie hillsides, in calcareous clay loam soil, sandy soil of S, sandy loam over cutiche, chalky soil of readedite waste in chalky soil, rolling buffalograss prairies, prairie bluffs and limestone ep ee of steep prairie bluffs, overgrazed grasslands, rocky and dry prairie hillsides, rocky ridges with Buchlo¥, cal- iche outcrops, and prairies anohe, the edges of chalky bank banks. An itional common name is "prairie verbena". The corolla is described as "blue" on Horr 4766 & 4775, "bluish- purple" on Harms 1758, and "reddish to purple" on McGregor 5678. In Oklahoma the plant was found by McGregor on rocky prairies in Murray County. In Texas Harms found it abundant along fencerows and sparse on eroded gravelly banks in grassland in Prag a — It is described by Ireland as common on the prairie in W County. McGregor found it "in scattered pea dl on mixed a og prairie hillslopes” in Osborne, scattered in fencerows, in red gyp- Sum soil of rough dissected prairie, and in short sea ’ association Hodgeman, scattered on prairie hillsides in Tre ego, scattered on en banks inr ough rai prairies, common in red gypsum soil of ¢ rolling prairies, and scat- tered colonies in red gypsum soil of prairies in Barber County. The Same collector reports the species as common in barren places in Overgrazed prairies in Morton, scattered in overgrazed places in bottam of prairie ravines in Scott, scattered in open places of 4,0 PHYTOLOGIA Vol. 11, mo07 rocky prairies in Russell, common on rocky prairie hillsides in Ford, and scattered patches on rocky prairie hillsides in Jewell ains in on igen Singer hillsides" in ike pateigy mee "in scattered colo- nie falograss prairies" in Russ ell, and "plants in asantesen Stamee [sic] on rocky prairie hillsides" in Hodgeman County . Horr says that the epecked is ere all over Meade County", in dry loam wasteland and "common" in dry loam grassland in Rus- sell, "common throughout Clark County", in dry sandy loam of grassland and "scattered in dry loam of grassland" in Clark and Jewell Counties, "scattered" in dry calcareous loam of chalk cliffs in grassland in Phillips, and "scattered in dry loam of overgrazed grassland" an eee County. Horr & Horr report it common but not plentiful in dry loam of grassland and wasteland in Trego County, while i & ay piney describe it as common in dry sand of roadside ditches in Kiowa County. The D. B. Dunn 3050 (in part) & 6595, S. W. Jackson 12-1, and Wilson & Miller s.n. [6-15-12], distributed as V. V. bipinnatifida, are actually V. ambrosifolia Rydb.; D. B. Dunn 3050 (in part), 3267, & 6375 are V. bipinnatifida var. latilobata Perry; Hulbert 3572 2& 3581, E. Le | Richards | 23, and Wagenknecht 309k & 13L5 are aed Dunn & Lint 136] are V. ciliata Benth.; C. M. Rowell i702 4 is V. ciliata var. longidentata Perry; and D. B. Dunn 3081 and Wagenknecht 175) are V, wrightii A. Gray, while the F. Be Jones 1867, wegen as 3 "prob. albino of V. bipinnatifida" is Vv. ef ciliata Ben Additional coro KANSAS: Barber Co.: E. W. Lathrop 22h1 (Lw); R. L. McGregor 10665 (Iw), 12211 (Im), 1402 (Le), 1237 at 14876 (Lm); E. L. Richards 3715 (Lw); B. ~L. Wagenknecht 38 (Iw). Clark Go.: W. He Hor Horr co (Lr), LuS9 (iw). Ellis oe R. L. McGregor 9308 (Iw); H. Shepherd 83 ae aT). Elis- worth Co.: R. L. McGregor 9261 (Lw); B. L. Wagenknecht 2501 (Lw) - Ford Co.: R. L. McGregor 10917 (Lm). Gove Co.: Hulbert 3535 (Lb— 50383, Lw). Hodgman Co.: “Re L L. McGregor 601 (LW), w), 5101 (Lm), 10598 (Lw, Lw). Jewell Co.: Ww. H. Horr H. Horr 4368 (Lw), 7775 (Lw); Re L. McGregor 7369 (Lw). Kiowa Co.: Horr & McGregor 3809 (Lw)- Lo~ gan Co.: R. L. McGregor 17251 (Lw). Meade Co.: We H. H. Horr 3448 (Iw). Morton Co.: Hulbert 3582 (Ib—-SOh9h, Iw); R. L. McGregor 2953 (Lm). Norton Co.: E. L. Richards 2761 (Lw). Osborne Co.! Re L. McGregor 7,25 (Iw). Phillips Go.: W. H. Horr 4766 (Lw)3 Be ae mecieegey 9533 (Lw); E. L. Richards 27a (Lw Lw). Pot tawatomie Coe? McGregor 12288 (Lw) — Cow: Ge Ee oe 278 (Lw)- Rake Cow: V. L. Harms 1307 (Lw); W. H. Horr 1983 3 Re Le Me 1965 Moldenke, Monograph of Verbena 42 Gregor 9575 (Lw), 12452 (Lw). Russell Co.: V. Le Harms 923 (Lw); . He Horr 4922 (Lw); H Hulbert 3470 (Lw); R. Lé McGregor 5678 (Ly), Tee CY 12l10 (Lw); E. L. Richards 365 (Lw). Scott Co.: Fear- ing & Latham s.n. [August 1h, 1950] (Lw); R. L. McGregor 13007 (Ly). “Smith Coe: We He. Horr E.108 (Lw, Lw)3 ss L. Richards 2889 (Iw). Trego Co.: cA i ams 973 (Lm); Horr & & Horr 4208 (Lw); R. L. McGregor 935 (Lw), rns (Iw); E. L. Richards ds 411 (L (lw). Wa-_ baunsee Co.: Ireland 7 ~ (Iw). OKLAHOMA: Murray Murray Co.: R. Le Me oregor 15015 (tw). TEXAS: Baylor Co.: V. L. Harms ee (i¥). Kent Cos: Ve Vv. L. Harms 1758 (Lw). Nueces S COs! F. B. Jones 1718a rer San Patricio Co.: F. B. Jones 2838 (Fj), = Tas Cie) (Ww), “4523 (Wr), ey (Ww). Travis Co.: | Me Pea Pearce s.n. [Austin, April 1932] (Lb . NEW MEXICO: Catron Co.: Norris & Zweifel s.n. (9-h- 48} “Cie 36L3) VERBENA BIPINNATIFIDA var. LATILOBATA Perry Additional bibliography: Moldenke, Résumé Suppl. 11: h. 196k; Moldenke, Phytologia 10: 92, 955 ; 96 (196) and is 326— 1965, Additional citations: NEW : Otero Co.: D. 26 (Lb--8)58). Socorro Co.: D. ra 3050, in part =F 5—Tahoo) Torrance Co.: D. B. Dunn 6375 (i (Lb-- folic je XVERBENA BLANCHARDI carat ee ot bibliography: . Jones, Am. Midl. Nat. Monog. 7: ed. 3) 213. 1963; "imac were: 10: 95, 155, 16, & 18). 1964; Moldenke, Résumé Suppl. 10: 1. 196. he E. We Lathrop 369 collection is a mixture with V. canaden— Sis (L. ) Britton. W. H. Horr 4436, 4648, & L697, distributed as php conctcpciaan de reger Sc The hybrid has been found in overgrazed prairie pastures and in prairie pastures with limestone rock surfacing, flowering in Ma ay. Additional citations: KANSAS: Woodson Co.: E. W. Lathrop 369, in part (Lw), 488 (Lw). VERBENA BONARIENSIS L. Additional synonymy: Verbena bonaeiriensis L. ex Alvarez, Fl. Santiago del Estero 106, sphalm. 1919. Verbena banariensis L. ex Moldenke, Résumé Suppl. 10: 7, in syn. 196). Additional literature: gin Mus. Hist. Nat. Paris 15: pl. 1h, fig. 4. 1810; R. A. Phil., Anal. Univ. Chil. 35: 192. 1870; Alva- re 1 Sul’ 235. 1 196; ney Sellowia 11: 57 & 13h. 1959; arog — mé Suppl. 10: 7 (1964) and ll: h. 1964; A. & I. Nehrling Book Perenn. 131. 1964; Moldenke, Phytologia 10: 95, 128 160, "291 ‘ & 498 (196) and 11: hl, 62, 63, 8h--86, 88, 98, 99, & 183 (196k), and ‘a Ui Le & 3hl. 1965. ditional common name recorded for this species is "sangre Lh2 PHYTOLOGIA Vol. ll, no. 7 de Cristo", but this name = a applied to V. litoralis H.B.K. and to V. sat ana (L.) B The Bracelin 1512 & 2827, prercnes as V. bonariensis, are actually var. conglomerata Brig. » While Beamer s s.n. [April 3, 1953], Demaree 15103, Glaziou 6048, and S. M. Tracy 8706 are V. brasiliensis Vell. Additional citations: ARKANSAS: Drew Co.: Demaree 23251 (Lb-—- es PUERTO RICO: Alain 10365 (G, Ij, N, Sj, 2 Sj, 2, Z)» BRAZIL: anabara: Boechat 22 (Ja); Palacios, Balegno, & cuezzo L017a (a5), , 4020 (Ja—5405). Minas Gerais: Duarte & rte & Graziola 7864 (N). Rio de Janeiro: Lanna Soti 12 (Ja, yas ENA BONARIENSIS var. CONGLOMERATA Additional bibliography: Moldenke, Piytologie 10: 95. 196k. This plant has been collected in fruit in December. Additional citations: CULTIVATED: Calif rnia: Bracelin 1512 (N), 2827 (N). ENA BRACTEATA Lag. & Additional sleainenake; Spec pig Handb. Biolog. Data 39h. 1956; G. N. Jones, Am. Midl. Nat. Monog. 7: [Fl. Ill., oe . 213. rg Steyerm., Fi. Mo. 1257--1281, — 18hh, os 302, 1963; Moldenke in Shreve & Wiggins, . & Fl. Son. Des. 3: 12h2. 1964; Moldenke, Résumé Suppl. “to: i (96h) and i: ihe haa) A. L. Moldenke, Piytologis 10 10: 166. 39085 av Reg tag Moldenke » Phytolo: 10: 96, 152, 16h, 155, ata & 307-—311 (196) an aed 1%, 103-- 210 (196k a mie pc pr a 3h, 105, rent ae 117, 163, 165, 1987 200, 202, & 203 eae ue ition illustrations: Steyerm., Fl. Mo. pl. 302, fig. 3- 3 Spector (1956) reports that this species can be killed by rel- tively low concentrations of at least one formation of 2,l\-D. Recent collectors have found the plant s sandy areas, limestone ground, pastures and op ren an » ove areas, waste places along railroad tracks, sandy soil of was or te places, sandy c loam soil, loam or rich loam with fair soil moisture, clay soil in schoo » talus at base of cliffs, roadside waste land, adsides, bare breaks along rivers, and rocky soil on creek bottoms with Euphor~ bia, on roadside banks and clay ravine banks, prairies, the bot~ tons of dried ponds, banks of stationery sand dunes, railroad a and fla some count was n = 7, Sacer found V. bracteata "rare and scatter 1965 Moldenke, Monograph of Verbena 43 ed in grazed sandy areas" in Stafford County; Lathrop encountered it in overgrazed prairie pastures with limestone rock surfacing in Woodson ae while Wagenknecht found it in dry clay soil in at the base of river bluffs in Doniphan, scat- tered throughout rolling sandy prairie pastures in Stafford, and upland in native grassland" in Rawlins, and "sparse on gravelly banks in grasslands" in Russell County. Horr informs us that it is "scattered in wet loam of floodland" in Meade, "common in Wi by my number 7330" — and for his no. 7330 he says "Several of these plants were found which grew erect ge aon all the habit of the species. Typical plants were also s The corolla is described as "blue" on Wore 163 & 4772. An ad- common name is "low vervain". The Norris & Zweifel s.n. [9-L-h8], distributed as V. bracteata, is actually V. bipinnatifida Nutt., while Dalke s.n. [9.10.39] is ¥. canadensis (L.) Britton. Steyermark (1963 ) re records this species from And: from Andrew, Barry, Car- roll, Cass, Clark, Clinton, Cole, Crawford, DeKalb, Douglas, Gentry, Harrison, Holt, Howard, Johnson, Laclede, Lafayette, Lawrence, , Livingston, Macon, Madison, Moniteau, New Madrid, Newton, Osage, Ozark, Pettis, Pulaski, Ray, Reynolds, Saint Clair, Saint , Scott, Stone, Sullivan, Vernon, Washington, and Worth Counties, Missouri, on the basis of specimens in the herbar- ium of the Missouri Botanical Garden at St. Louis, not as gh seen by me. The Crookshanks 142 which he says "appears to be a hybrid between ¥. bracteata and V. V. hastata” is actually xV. perriana Mol- denk e. Additional citations: NEW YORK: Rockland Co.: Lehr 1018 (N). INDIANA: Madison Co.: A. R. Moldenke 801 (Lw). IOWA: Cass Co.: H. N. Moldenke 21938 (Iw). Clayton Co.: A. R. Moldenke 992 (Lm). Del~- aware Co.: J A. R. Moldenke 985 (Lw). Iowa Co.: A. R. 7: A. Re Moldenke 973 (Lw). Keokuk Co.: A. Re Moldenke 969 (Lm), 970 (Iw). Linn Co.: A. R. Moldenke 978 (Lw). Madison Co.: 2 A. Re Moidenke 9h (Li). Mahas~ hhh PHYTOLOGIA Vol. 11, no. 7 ka Co.: A. R. Moldenke 958 (lw). Marion Co.: A. Re Moldenke 952 (Lw). Mills Co.: A. R A. Re ~ Moldenke 919 (LW). Pottawatomie Co.: “he R. Moldenke 921 (Ly) s 933 (Iv). Poweshiek Co.: ALR. Moldenke 968 (Iw). Warren Co.: A. R. Moldenke 91,8 (Iw). Washington Cos: B. L. Wagenknecht 330 (Iw). MICHIGAN: Menominee Co.: A. Re Mol- denke 1025 (Iw). Wexford Co.: A. R. Moldenke 1028 (Lw). WISCON- SIN: Adams Co.: Rickett s.n. (Friendship, June 24, 1926] (Lb-- 27608). Dane Co.: A. R. Moldenke 1007 (Lw). Iowa Co.: A. R. Mol- denke 1006 (Lw). MINNESOTA: Polk Co.:Terrell 1833 (W--2300720). KANSAS: Allen Co.: A. R. Moldenke 874 (Iw). Anderson Co.: A. Re Moldenke 8 872 (Lw). Barber Co.: We Be: Horr 3118 (Lw); R. L. McGregor 12865 (Lw), ), 1766 (Lw), 14913 (Iw). Barton Co.: A. Re -, Moldenke 896 (Li). Brown Co.: A. Re R, Moldenke 911 (LW). Chautauqua Co.: A. Re R. Moldenke 884, (Lw). “Cheyenne Co.: We We He Horr 4725 (Lw), 470 (im); R. L. UcGregor 13566 (Lv). Cloud Go.: R. L. McGregor $33 (Lw). Coffey Co.: y CO: We H. Horr 3481 (Lw). Comanche Coe: R. L. Mc Gregor 14669 (Lw). “Cowley Co.: A. A. R. Moldenke 886 (Lw). Ce Coe: Coe: Re L. McGregor 13517 (Lw); E. Se Richards 284) (Lw). Dickin- son Cos: e« R. Moldenke 902 (Lw). Doniphan Co.: 2 We He Horr 4301 (Lw); B. L. Wagenknecht 3290 (Lw). Douglas Co.: ¥, Cc. Stevens 6719 (Lw). Elk Co.: A. R. Moldenke 876 (Lw). Blisworth Co+: A. R. Moldenke 898 (Lw); B Be A Wagenknecht 2506 (Iw). Finney Coes Wilson & Miller sen. [7-18-12] (Lw). Ford Co.: W. H. Horr 3613 (Iw). Franklin Co.: Hetzer 318 (Lw). Geary Co.: 3 Fe Ce Gabes 19783 (Lb—-7720); A. Re Moldenke 903 (Iw). Grant C Co.: Hulbert 3572 (Lw). Harper Co.: | Re Le McGregor 14697 (lw). Hodgeman Co. R. L. McGregor 10616 (Lw). Jefferson Co.: J. Latham 368 (Lw); A- Re Moldenke 90, (Iw). Jewell Co.: W. H. Horr 39k (Iw). Kingman Co.: Re L. McGregor 7322 (Lm); A. R. Moldenke 893 893 (Lw). Kiowa Co.? Horr &) McGregor 3790 (Lar) 5 Re het McGregor egor 15168 8 (Lw). Leaven- worth Co.: E. L. "Richards 2670 ( (Lw). Linn Co.: A. R. Moldenke 870 (Iw). Logan Go.: R. L. McGregor 9409 (Lw). ri Horr 3472 (Iw). Marshall Co.: W. H. Horr 569 (Lm). Meade Co.? W. H. Horr 3301 (Iw), 3345 (Lw), 3458 (Lw). Montgomery Co.: R. be UcGregor 159k (Lm) » Morris Co.: E. L. Richards 1786 (Lm). Mor= ton Co.: Hulbert 3581 (Lw); E. L. Richards 2455 (Iw), 2533 (Lw)- Nemaha Co.: R. L R. L. McGregor 12270 (Iw). Norton Co.: R. L. “L. McGregor 9497 (Lw). Pawnee Co.: B. Le L. Wagenknecht 450) (Lw). pratt Co.: H. N. Moldenke 21928 (Lw). Rawlins Co-? Hulbert 3498 (Lb—-50L61, Iw); R. L. McG L. McGregor 9469 (Lw). Reno Co.: R. L. McGregor 7330 (Lw), 7331 (Lw); B. L. Wagenimecht 309) (Lw). Republic Co.: G. Es EB. Mor SS ley 114 (Lw), 52h (lw). Rice Co.: BE. W. Lathrop 385k (Lw). Riley Co.: F.C. Gates 12748 (Lb—-27596). Russell Co.: Hulbert 3471 1965 Moldenke, Monograph of Verbena Lhs (Iw). Saline Co,: A. Re Moldenke 900 (lw). Sedgwick Co.: W. He Horr 6719 (Lw). Seward Co.: We H. Ho: Horr 3096 (Lw). Shawmee Co.: Volle 790a (Lw). Smith Co.: W. H. Horr 4772 (Lw). Stafford Co.: Ungar 819 (Lw); B. L. Wagenlmecht +t 43L5 (Le). Stanton Co.: R. L. McGregor or 14529 (Lw), “16134 (Lw). Sumner Co.: A. R. Moldenke . 888 (Lw). Trego Co.: Horr & | & Horr 4175 (Lw); J. D. . Poindexter dexter 18 (I (Lw). Wallace Co.: V. L. Han Harms 1189 (Lw). Wichita Co.: Hulbert 3262 (Lb=~50502, Lw). Wilson Co.: R. L. McGregor 15053 (Lw); A. wa he Moldenke 875 (Lw). Woodson Co.: E. W. Lathrop p 1276 (Lw). Wyan- dotte Co.: E E. L. Richards 3210 (Lw).» MISSOURI: Barton Co.: Es Je alae 5094, (Lb--509k4), 55331 (Lb--509)8); Z. Williams s (Lamar, Aug. 1928] (Lb—-h166) . nee Cost Ae ie , Moldenke “ie (Ly), 892 (Lw). Boone Co.: Daniels s.n. [Columbia, Jul. '97] (Lb— 116s); Drouet 791 (Lb-=12069), san sen. [Columbia, 8/31/30] (Lb-~ 8502, Lb~-8503); Favor sen. [July 16, 1901] (Lb~-h160, Lb--161); Rickett s.n. [June 29, 1929] (Lb—710)) 5 Van Horn sen. (Columbia, i ah 5) (Lb--1094k). Dunklin Co.: Steyermark 7934 (Lb--5325). 0.3 J. M. Mason s.n. (Meramec State Park, 7/17/35] (Lb— pines A. R. Moldenke BL5 (Lw (Lw). Grundy Co.: Giiblalianies’ 2 2h0 (Lb~-297h5). Henry Co.: Te R. Moldenke 86), (Iw); E. J. Palmer 55829 (Lb--50945). Jackson Co.: B. F. Bush §.n. [6-2-1892] (Lb— 62). Mississippi Co.: R. R. Kincaid sen. [Sept. h, 1927] (Lb-- 4163, Lb--l16h). Nodaway Co.: J. R. Singleton 169 (Lb—-31593) « ARKANSAS: Bradley Co.: Demaree 25503 (Lb—5 3498). Marion Co.: Demaree 7929 (Lw). Saint Francis Co.: Demaree 78,0 (Lw). NE- VADA: Clark Co.: Clokey 8096 (Lb--35773, N). COLORADO: Poulder Coe: Daniels 2 (Lb--27601). El Paso Co.: Naylor s.n. (Aug. 19, 1933] (Lb--2 (Lb--27600) . Huerfano Co.: B. L. Wagenknecht 2903 (Ly). Jefferson Co.: Perdue 034 (2122918) « NEBRASKA: Cass Coe: Ae R. Moldenke 917 (Lw). Otoe Co.: A. R. Moldenke 961 (Lw). OKIA~ HOMA: Pawnee Co: C. F. Coffman s.n. [Pawnee, Aug. 31, 1933] (Lb— 14291). NEW acai ore Fé Cow: Dunn & Lowe 6485 op aieetae Socorro Co.: De Be 71 (Lb--501). Torrance Cos: D. B Dunn 6125 (Lb—3897), Pas aes), 620h (Lb--385U5) CALI- A: San Bernardino Co.: Parish & Parish 2 2171 (Lb--27692). VERBENA BRASILIENSIS Vell. Additional bibliography: Reitz, Sellowia 11: 57 & 13h G97) and 13: 67 & ee 1961; Jones Rowell, & Johnston, Flow. P Ferns Tex. Coast. Bend 143. 1961; Moldenke, Phytologia 10: pad 97, 160, 217, 296, & 297. loshs Yoldenke, Résumé Suppl. 10: 1 (1964) and 1i: 3 & 86. 196k. The corolla is Fe as deep-lavender on Perdue 4153 and as purple on Beamer s.n. (April 30, 1953]. Recent collectors have found the plant in clay of roadside ditches, in clearings LL6 PeTITOLOGLA« Vol. 11, no. 7 near roadsides, and in clay banks and ditches. F. B. Jones de- scribes it as locally abundant in clay soil along ditches near railroads in San Patricio County, Texas. Demaree calls it "common sas. "Material has been misidentified as V. xutha Lehn. tional citations: FLORIDA: Escambia Co.: S. M (Lb—-27599). ALABAMA: Dallas Co.: Perdue 152 Ge Dhoaiss P a. KANSAS: Ouachita Co.s Demaree 18182 (Lw). Saint eager oa = aree 15103 (Lb--l,7033). LOUISIANA: Vernon Par.: Re L 15570 (Lw) (Lw). TEXAS: Harris Co.: Beamer s.n. [April 0, Sy i) Nueces Co.: F. B. Jones 6192 (Ww). San Patricio Co.: F. B. Jones 192) (Ww). BRAZIL: Rio de Janeiro: Glaziou 6018 (Ja—11150). VERBENA CALLIANTHA Briq. Additional bibliography: Moldenke, Phytologia 10: 97 (196k), li: A189 (1964), and 11: 290 & 315. 1965. VERBENA CAMERONENSIS L. I. Da Additional bibliography: Moldenke, Résumé Suppl. 10: 2. 196h; Moldenke, Phytologia 10: 97 (196) and 11: 307. 1965. The flowers are described as "rose-blue" on McGregor 16l2. The species has been found on the floodplains of small rivers, fruiting in February. Additional citations: MEXICO: Puebla: R. L. McGregor 16442 VERBENA CAMPESTRIS Molde Additional ididopwentes Rests, Sellowia 11: 57 & 134. 19595 Moldenke, Phytologia 10: 97. 196k. VERBENA CANADENSIS (L.) Britton Additional & emended synonymy: Anonymos caroliniensis Walt., Fl. Carol. 16h. 1788 [not Anonymos caroliniensis Walt., op. cit. Schnarf in Linsb., Handb. Pflanzenanat. 10 (2): 233 & 688, sphalm. 1929. Lobelia aubrietiae J0nsson ex ciate pete an. peso Verbenac. 6, in syn. 1931, caress aubletia canadensis f. canadniets ote Fl. Mo. 1260 & 1261. 1963. Additional bibliography: Jonsson, Bot. Notiser 6: 169~-187. sae Schnarf, Ost. Bot. Zeitschr. 7h: 1. 1925; Linsb., Handb. Zyt Verbenas. 6, 2h—-25. 1931; Patermann, Beitr. Zytol. Verben~ ac. 1315, ‘ex. ‘oe & (5h), pi. " fig. 1 & 2. 1935; D. B. Ward, Rhodora 6): 91. 1962; Steyerm Fi. . 1257 & 1260-1263, map 1845, pl. 303, fig. 1. 1963; $ & tukhopadhyay, Journ. Genet. 58: 359, 372, 38h, & pl. 12, fig. se fle 1963; Taylor & Hamblin, Handb. Wild Fl. Cult. 168-1 169 & 267. 1963; Moldenke, Résumé 1965 Moldenke, Monograph of Verbena bh7 Suppl. 10: 1& 7 —_ and 11: mae & 5--8. 196; Moldenke, Phy- tologia 10: 97--98, 11h, 122, 125, 15, 282, 283, 311, & 199 9 (1964), aa: 6-36, 32, 33, "1a. reo 47, 85, & 86 (196k), 7 1 nachdabencees illustrations: Patermann, Beitr. Zytol. Verbenac. pied, Pigs 1'& 2. mite Steyerm., Fl. Mo. pl. 303, Phd 1. 19633 shares & Mukhopadhyay, Journ. Genet. 58: 384, pl. i2, fig. 5h. Ward (1962) agrees that the Anonymos caroliniensis of Walter, + Carol., page 16 (1788) is sepa Verbena canadensis, but he is of the opinion that the similar binomial on page 263 pos- sibly refers to erinaceus Ball.., while the A. caroliniens on page 188 is Amphicarpa bracteata (L.) Fern. The corolla of Verbena canadensis is described as "reddish- violet" on Harms 1379. The haploid chromosome number is given as 6 by Schwencke (1931) and by Patermann (1935). Recent collectors have found the species in sandy grasslands, fencerows and roadside fencerows, bluestem prairie hay meadows, roadside ditches, thickets in Limestone glades, prairie hay fields and dry roadside prairie hayfields, basic soil of cedar glades loose soil of old limestone quarries, clay or dry rocky soil, or dry clay soil in cutover woodlands, on roadside and rocky prairie grazed and weedy rocky prairie hillsides, rocky wooded or shady oak-wooded hillsides, and prairies around the edge of lakes, and irie wood in grasslands in dry sandy clay soil", by Horr as "common on roadsides and wasteland in dry loam and clay", by Wagenknecht as occurring in " sandy soil along edges of dis turbed areas on arce in rocky area on southwest- ds Sandstone surfacing", "roadside bank of bottomland f g area, limestone surfacing", "overgrazed prairie pasture with Maclura along crest of gully", and "common on oak-wooded hill- pomifera side, ener outcrops". r calls the species "common on banks in rocky limestone ae ee colonies on prairie banks", "on ro urfacing, in native bluestem prairie meadow", "common on slopes, rocky overgrazed pastures", "common near lime stone escarpment on prairie hillsides", "common in area of limestone outcrops", Lh8 PHYTOLOGIA Vol. ll, nos? "common on prairie banks", "scattered on bank of little ravine in prairie", "scattered colonies in rocky wooded a “common on rocky prairie hillsides, limestone surfacin Ty, starting new plants = _ s elf-rooting stems. This species ee Bush 15503 is V. ambrosifolia Rydb.; W. H. Horr 4983, H ene 3582, Ireland 71 a, Re L L. McGregor 9295 & 9575, and B. L. - Wagenknech are V. are V. bipinnatifida Nutt. ; “Rickett sn. (Friendship, June 2h, 1926] is V. bracteata Lag. & Rodr.; R. Runyon 297 is V. ciliata var. longidentata Perry; W. H. Horr 3086 is x is xV. deamii “Moldenke; R. Runyon 2625 is V. delticola Small; B. L. Wagenknecht 3615 is V- alba Moldenke. As noted above, the R. Runyon 2197 & 2625, cited in Phytologia 8: 153. (1962) and 465 (1963) as V. canadensis, are pegs i V. ciliata var. longidentata and V. delticola, respect- Vv LS ely Steyermark (1963) records V. canadensis also from Benton, Bol= tantéal Garden at saint peer) not as yet seen by Additional citations: IOWA: Mahaska Co.: H. N. ifoldenke 21941 (Lw). TENNESSEE: Rutherford Co.: Demaree 47565 (Liv) « KANSAS: Allen Co.: R. L. McGregor 10012 (Lw), 12745 (Iw). Anderson Co.? Re be ucoreeer Lenk (Lw); E. L. Richards 328 (Lw); B. L. Wagen- Knecht 3651 (Iw). Bourbon Go.: EB. W. Lathrop 3789 (Lw); Re Rubs wceregor r 992k sa Brown Co.: Re L. McGregor 17020 (Lm). Chav- tauqua Co.: We a Horr s.n. [July 5, 19 30] (Lw)3 E. W. spore ud (19) 5 R. L. McGregor 4210 (Lw). cherokee Co.: 5. W. Lathrop ioe (Le); Re L. McGregor 1526h (Lm), 15332 (Lw). Clay Co.: Re as s Becesee ey 5530 (Iw). Coffey Co.: W. He Horr 3496 (Lw). Cowley De L. Marsh 426 (Lw). Crawford Co.: R. L. MeGregor 11176 (im). ~ Douglas Co.: C. A, Benson s.n. [B Baldwin, May 29, 1932] 932] (Lb- 27607); D. L. Marsh 1522, (Im); B R. L. McGregor 1,228 (Lw), 14289 Sa (Lw); Ungar la (Iw). Elk Co.: Fearing & Latham s.n. [August 10, 1965 Moldenke, Monograph of Verbena hho 1950] (Lw); R. L. McGregor 60 (Lw), 9059 (Lw), 11197 (Lw). Franklin Co.: Hetzer 91 (Iw); E Hulbert 3397 (Lw); ® Li McGregor 10048 (Lw). Greenwood Co.: R. L. McGregor 10322 (Lw, Lw), 11208 (iw). Jefferson Co.: G. Latham 7: 9 (Lw). Johnson Co.: B. io Wagenknecht 258h (Lw), 2605 (Lm), _ 99 (Lw). Labette Co.: Re Le McGregor 174)2 (Iw). Linn Co.: H. N. Moldenke a (Lw). Miami Coe: Re | R. L. McGregor 4242 (Iw). Montgomery Co.: WH. Horr s.n. [July 5, 1930] (Lw); Re: R. L. McGregor 4,331 (Lw) - 10189 (Ie). Neosho Co.: W. W. Holland 153 (Lm); 5. L. Richards 3160 (Iw). Pottawato- mie Co.: Re Le. McG: L. McGregor 5473 (Lw). Shawnee Co.: V. L. Harms 1379 (Lw); Volle 653 (Iw). Wabaunsee Co.: V. L. Harms 131 (Iw); Maus 36 (Lb—15169); R. L. McGregor 5548 (Lw). Wilson Co.: R. L. Mc Gregor 4300 (Lw);_ ‘Vaugn mM S.M. [May 1, 1937] (Lw). Woodson Co.: | Re L. McGregor 47h (Iw); E. W. Latnrop 34 (Lw), 169 (Iw), 369, in part (Lw). MISSOURI: Barton Co.: Ze Williams sen. [Lamar, August 1928] (Lb~-158). Boone Co.: Bell s.n. [Columbia, )/16/1801] (Lb- ); H. E. Bradley s.n. (Columbia, Apr. 29, '02] (Lb--l12); Daniels s.n. (Columbia, Apr. 1897] (Lb—-)143), s sen. (Columbia, May 11, 1902] (Lb—-h147, Lb—-1148, Lb—b15h), s.n. (Columbia, May 1903] (Lb--k1S1, Lb--)152, Lb—1,158) , Sen. (Columbia, June 1903] (Lb—-11,6); Drouet 2763 (Lb--25810); D. B. Dunn 12437 (Lb--37875); Favor sen. (Aug. 29, 1901] (Lb--l139, Lb—-4149), sen, (Columbia, May 2, 1903] (Lb--)153); S. Gray sen. (Columbia, May 17, 1898] (Lb—1,157); G. E. Moore 4 (Lb b—11),38); Oungst s.n. [April 15th, 1933] (Lb—11)6); Rick Rickett sen. [April 1926] (Lb--7099, Lb—7100), Sen. [Rocheport, July ‘ly 4, 1927] (Lb--7101), sn. [Bear Spring, A- pril 20, 1929] (Lb--7102), sn. [Grindstone Park, June 1, 1929] (Lb—7103) ; J. Re Singleton s.n sen. (April 18, 1936] (Lb—31:628) ; O. Le Steele s.n. (Columbia, Apr. 17, 1902] (Lb—-4145, Lb—-150); ze J. Sylvester 3 (Lb--15148); Van Horn s.n. (Columbia, 7/27/25] Ib=-10953). Cooper Co.: B. F. Bush 14425 (Lb—15596). Crawford on H. N. Moldenke 21916 (Lw). Franklin Co.: J. M. Mason s.n. [Meramec State Park » 4/19/37) (Lb—3)369). Greene Co.: M. Carter S.n. (Galloway, April 10, 1929] (Lb--14184). Harrison Co.: R. J. Lemaire 1188 (Lb--l9412). Iron Co.: Broadhead sen. (Lb—l1159). Jackson Co.: B. F. Bush s.n. [5-19-1891] (Ib—l156) . ae Cost Miles s.n. [5/1/1898] (Lb--47639). Lawrence Co.: E. 53390 (Lb=-509)6). Madison Co.: Swallow s.n. [July Se eT be (Lb— 2072) . Maries Co.: A. R. Moldenke 852 (lw). Marion Co.: Broad- head s.n, [May 1856] ~(Lb--9609) « Willer Co.: Damon s.n. (April 12, 1938] (Lb—34761). Morgan Co.: B. F. Bush UkS (Lb—~15681) Sedhen Co.: E. J. Palmer 53471 (Lb—-5091:7) « Phelps Co.: A. Mtil- ier s.n. (Meramec, March 30, 1935] (Lb--24970). Scott Cost i? R. 450 PHYTOLOGIA Vol. 11, no. 7 Kincaid s.n. [Commerce, Aug. 24, 1927] (Lb--1)0, Lb--11). ney Co.: ade Spencer 123 (Lb-~35300), sen. [9.16.40] Tipe ce Sone [4.12.41] (Lb--35530, Lb—L3916). co County undetermined: R. Baker sen. [Lovers Leap, May 18, 192] (Lb--9000); Dalke s.ne . [In- dian Trail State Forest, 9.10.39] (Lb--48915); Ryckman s.n. [Tir- isport, May 1st] (Lb—3326))) « ARKANSAS: Franklin Coe: Demaree 20890 (Lb--53396). Izard Co.: A. Robinson 1790 (Lw). Lawrence Co.: Demaree 25957 (Lb--7027). “Marion Co.: Dem Demaree 47941 (Ly) D. B. Dunn 12837 (Lb—-39826). Saline Co.: Demaree emaree 3477L (Lb—~ 53413). Stone Co.: D. L. Marsh 637 (LW). LOUISIANA: Cameron Par.: S. M. Tracy 8707 (Lb—27606). OKLAHOMA: Latimer Co.: Ve Le Harms 173 (Lw). Pittsburg Co.: McClary s.n. [Marsh 30, 1935] (Lb-=32371). TEXAS: Galveston Co.: Joor sen. [Galveston] (Lb— 27609). Harris Co.: Joor sen. [March 3, '77] (Lb—27598), SMe (March 9, '77] (Lb--27608). Marion Co.: B. L. Wagenkmecht 3529 (Lw). CULTIVATED: Missouri: Kucera s.n. (liar. 1957] (Lb——37518) « Wisconsin: H. N. Moldenke 21912 tia). VERBENA CARALEONSIS f. CANDIDISSIMA (Haage & Schmidt) Palmer & Steyerm. ddit tonal bibliography: Steyerm., Fl. Mo. 1260 & 1261, map 1845. 1963; Moldenke, Résumé Suppl. 10: 1. 1963 Moldenke, Phyto- logia 10: 98. 196). form has been found by MeGregor on rocky prairie banks at the edge of woods and rocky prairie roadside banks, flowering and fruiting in May. He gorge the flowers on his no. 7214 drying". dditional citations: KANSAS; Bourbon Co.: R. L. McGregor /e’e 7212 (Lw). Crawford Co.: R. L. McGregor 7214 (Lw). VERBENA CANESCENS H.B.K. tional bi se seby a oadside Flow. eee ae Poe Moldenke, ae: Supp 36:5 Moldenke, Phytol 100, 140, 11 16179, 1 182, “sal 186, 303, 30h, 306, ee S02 * 56h), Tl: 12-1), 35, oe & 43 (196k), and i: 256. 1 speci been found by Jones in brashy a a in sandy pt soil over coal dhe and in caliche outcrops, locally 4 , in sandy loam on caliche hillsides, flowering an fruiting in March and April. In Mexico it has been found by McGre~ gor and other recent collectors "prostrate on Socky. - wooded n rocky shrubby 2 in open pine forest, gypsum soil", and — Pe cite, at altitudes of 5370 to 7000 feet, flower in July. The flowers are described as "blue" on ion Baran 16612, “T66lds, & 16725 and Mc Gregor, Harms, Robinson, R. del Rosario, & Same. 98. Jon Jones 1822 rete eee saemanemencmmntemtams et ec The species is recorded from Amazonas, vot caabia (san Cristobal), by Troncoso, but is seems most unlikely that this is correct. 1965 Moldenke, Monograph of Verbena 451 Additional citations: TEXAS: Jim Wells Co.: F. B. Jones 1822 (Ww), 2935 (Ww). MEXICO: Nuevo Leon: R. Le McGregor 16725 (LW); McGregor, Harms, Robinson, R. del Rosario, | & Segal 98 (Le) (lw). Oa- 1664 GE R. L. McGregor 16512 (iy). San San Luis Potos’: R. L. McGregor 1 (Lw) ANESCENS f£. ALBIFLORA Moldenke ENA Peas zareg bibliography: Moldenke, Phytologia 10: 99 & 100. 196). NESCENS var. ROFMERIANA (Scheele) Perry Additional bibliography: G. Rzedowski, Act. Cientif. Potos. lj: 65 & 91. 1960; Jones, Rowell, & Johnston, Flow, Pl. & Ferns Tex. Coast. Bend 143. 1961; sitet abe Résumé Suppl. 10: 2. 196k; Mol- aa Phytologia 10: 99--100 & 182 (196) and 11: 13—~15 & 18. Lathrop found this variety growing in sandy soil of rocky gullies, with granite outcrops on the slopes with juniper, Yucca, flowering and fruiting in April. Lathrop 2075 is a mixture with ve. piicats Greene. The F. B. Jones 1822 & 2935, distributed as V. canescens var. roemeriana, seem to be typical V. canescens H.B.K.— eaeean ‘eatin TEXAS: Bee Co.: F. Be J Jones 1732a (Ww). Burnet Co.: E. W. Lathrop 2075, in part (Iw). VERBENA CANIUENSIS Moldenke dditional bibliography: Moldenke, Biol. Abstr. 27: 98). 1953; Moldenke, Phytologia 8: 82--83. 1963. VERBENA CAROLINA L. Additional bibliography: Schlecht., Ann. Sci. Nat., ser. 3, lh: 342—3hh. 1850; Moldenke, Phytologia 10: 100--101, 104, 129, 130, » lhl, 280, & 314 (1964) and 11: 13, 54-56, 86, 105, 123, 126- 128, & 16h. 1964; Moldenke in Shreve & Wiggins, Veg. & Fl. Son Des. 2: 1239. 1964; Hocking, Excerpt. Bot. A.7: 455. 1964; Langman, Select. Guide Lit. Flow. Pl. Mex. 7 748 & 1010. 1964; Moldenke, Phy- Fenn Cee ee Fearin & Thompson describe the corolla of this plant as "very Light-bluet. oe singed the plant in dense pine forests in black volcanic gs encountered it on rocky slopes. Shreve & mr erigeh) gt distribution is said to be "In diverse soils and habitats, Lower Sonoran to Hudsonian Zones, from Pima County, Arizona, through most of Ar pes gg ly soenere Baja California to El Salvador; ascending to 10,500 feet Additional citations: MEXICO: Chiapas: Breedlove “598 (Z). Mor elos: Fearing & Thompson 131 (Lw). VERBENA CATHARINAE Moldenk: Additional bibliography Reitz, Sellowia a 57 & 134 (1959) and 12: 157. 1960; Moldenke, Phytologia 10: 101. 196k. 452 PHYTOLOGIA Vol. 11, no. 7 Common names recorded for the species are "camaradinha", "for- mosa sem dote", and "jurupeba", but these are applied also. to most members of this genus in southern Brazil. VERBENA CHEITMANTANA Additional cahe pp tsi i TN Phytologia 10: 101 (196k) and 11: 123. 196k. ENA CHILENSIS Moldenke Additional bibliography: : A. Phil., Anal. Univ. Chil. 35: 190—-191. 1870; Mufioz Pizarro, Espec. ara Descr. “philipp i side 1960; Moldenke, Phytologia 102 101. 19 he isotype photographed ad wenger as his type photograph number 1717 was deposited in the herbarium of the Botanisches Museum at Berlin, now unfo rade destroyed a ae CHILE: Mendoza: R. A. Philippi s.n. [Men- doza; ac. Hist. Nat. Chile 2h)6 & 54790; Macbride photos nih gf Reda of isotype). VERBENA CILIATA Benth. Additional synonymy: Verbena ciliata var. ciliata [Moldenke] in rer & Fl. Son. Des. 2: 12hh. 1964; Moldenke, Phytologia 10: 101— ° 8, & 283 (1964) and lis {3, 3B, 49; 9, 105, ise, "158, 188, & 10 (196), and 11: 308, Rte ccna the veers of this plant as "reddish", while F. B. Jones says for 1867: “corollas white in con- trast to the usual abe var. longidentata] which was present at same locality; several specimens of white form noted; prob. albino of V. bipinnatifida." He found the plant in c soil along Rasetcons arbeagal found it prostrate in dry level rocky grassland; and Maguire reports it "on rocky granitic slopes flowering and fruiting in April and May. It has been misidenti- fied and distributed in herbaria as V. bipinnatifida Nutt. and as V. wrightii A. Gray. ues eee Shreve & Wiggins (196) describe its natural distribution as "On flats, sandy plains, dry limestone hills, and rocky slopes lated records from Tennessee and South Dakota. repent es aaa gin of the Sonoran Desert near Tucson and Aguila, Ariz The F. B. Jones 4478 and C. M. Rowell 4.702, race 2% as V. ciliata, are ras var. longidentata Perry. “Fadi tional citations: TEXAS: San Patricio Co.: F. B. Jones 1867 (Ww). NEW MEXICO: Otero Co.: D. B. Dumn 3276 (Lb—-LBLL7) « 1965 Moldenke, Monograph of Verbena 453 Socorro Co.: D. B. Dunn 3039 (Lb--,8h69); Dunn & Levine ue (Lb- 48192); Dunn & Lint 138 (Ib--53810). Torrance Co.: D. B. Dunn 6445 (Lb—],0930). ARIZONA: Graham Co.: B. Maguire ioe (xy. MEXICO: Jalisco: R. L. McGregor 16626 (Li). VERBENA CILIATA var. LONGIDENTATA Perry Additional Reigns, 2 Moldenke, Phy oloped 10: 102 & 283 sb? te lt : 32 & 3. 196k. colle ctors “ad found this plant in clay of open pas- rrminrstg pri in sandy pastures, while Rowell notes it as mre to localiy frequent in sandy loam and tight soils". His no. 702 is said to have had "pale-purple" corollas. The plant has been found in flower in February and March and in fruit in March. Her- barium material has been misidentified and distributed as V. bi- pinnatifida Nutt., V. canadensis (L.) Britton, and V. ciliata Benth. Runyon Zot was previously cited by me as V. canadensis, but this is err Additional tata TEXAS: Bee Co.: F. B. Jones 2842 (Fj). Cameron Co.: R. Runyon 24,97 (N, N, N)- San Patricio Co.: F. B. Jones hhh5 (Fj), LL76 (Ww); C. M. Rowell 4702 (Ww). VERBENA CILIATA var. PUBERA (Greene) Perry Additional bibliography: Moldenke, Prytologia = 102 (196) and 11: 48. 1964; Moldenke in Shreve & Wiggins, & Fl. Son Des. 2: 125. 196. Maguire, Richards, & Moeller assert that this plant is "common i grass of ee mutica type, with erescets Yucca elata, on ats", flower ring and fruiting in May. In Shreve & Wiagine ( 196) its OB hel bi is given as "In cindery soil, on sandy clay flats, sandy and grassy plains, and washes, mainly Upper S noran Zone, southern California to western Texas. Possibly enter- ing the Sonoran Desert along the eastern margin in Arizona." Additional citations: NEW MEXICO: Grant Co.: Maguire, Richards, & Moeller 11503 (N). VERBENA CLAVATA Ruiz & Pav. Additional bibliography: Soukup, Biota 5: 42 & 13h. 1964; Mol- denke, Phytologia 10: 102. 196). Soukup (196) records the vernacular name "tacpa". VERBENA CLAVATA f. ALBIFLORA Moldenke Additional age gt dy Moldenke, Phytologia 9: 32. 1963; Soukup, Biota 5: 2. 196). VERBENA CLAVATA var. CASMENSIS Moldenke Additional bibliography: Soukup, Biota 5: 2. 1964; Moldenke, Phytologia 10: 102. 196). xVERBENA CLEMENSORUM M Additional aaa: a Phytologia 9: 33—3h (1963) Lou PHYTOLOGIA Vol. 11, no. 7 and 10: 102 & 210. 196). CLOVERAE Moldenk eee vee “sisograpty: Jones, Rowell, & Johnston, anh - Bend 14). 1961; Moldenke, Résumé Suppl. 196k Sci cunaee pyc 10: 102 & 178 (1964) and 11: "03. 96hp - Le Moldenke Scag init 10: 238. 196k. ecent c ecto s have found this species on sandy slopes near creeks, on sandy sil shoulders, in open pastures, and scattered around depressions in dry rocky mesquite range. Williges calls it rare" in San ed it "usually on slopes" in Kleberg County. The flowers on R. Le McGregor 16764 are described as "blue". ~~ Additional citations: TEXAS: Bee Co.: F. B. Jones 57 (Ww). Kleberg Co.: F. B. Jones 2977 (Ww), 3801 (iw). San Patricio Co.: F. B. Jones 6177 ( (Wer) 5 Williges Se (Ww). Webb Co.: R- Le McGre= gor - 1676 1676) ( Cap a XVERBENA CONATA Moldenk a Gia ecaetes Moldenke, Phytologia 10: 103 & 210. 196 CORRUPTA Moldenk ; Yai ihanitherente': Moldenke, Phytologia 10: 103 & L98. 9 — CORYMBOSA Rufz & za Re dditional bibliography: z Pizarro, Sin. Fl. Chil. 199. 1959; Moldenke, Phytologia int 3 103. 1964; Soukup, Biota 5: 2. 964; Aw & I. Nehrling, Pict. Book Perenn, 131. 196k. XVERBENA COVASII Moldenke Additional bibliography: Moldenke, Phytologia 10: 103 (196h), li: 135 (1964), and 11: 290. 1965. VERBENA CRITHMIFOLIA Gill. & Hook. Additional Mibiogrepiy: Hieron, Bol. Acad. Nac. Cérdoba li: 409. 1881; Hieron., Pl. Diaph. Fl. Argent. 215. 1882; Moldenke, Phytologia 10: “103. 110, 120, & 288. 196k. been CUNEIFOLIA Ruiz & ah 96h Mo 1- Addit ional Sieaiaiepenes 4 Soule Biota 5: 42 & 134. 1 iP =: denke, Phytologia 10: 103, 287, & 296 (196) and 11: 158 & Soukup (1964) records the common name "china verbena". Peco Moldenke 126 mal bibliography: 60 & 1261, m aati 3 1963 Moldenke, pa or 20h @ 34 “1396h) and’11: 202 Sareea hs vo Moldenke » Résumé Suppl. 10: oop terial of this hybr id has been mis identified and distributed 1965 Moldenke, Monograph of Verbena 455 in herbaria as V. canadensis (L.) Britton. On the other hand, the ~ H. Horr 1679, distributed as xV. deamii, is actually V. Ls Satna citations: KANSAS: Gray Co.: W. H. Horr 3086 (Ly). VERBENA DELTICOLA Small Additional bibliography: Moldenke in Shreve & Wiggins, Veg. & Fl. Son, Des. 23 123. 196h; Moldenke, Phytologia 10: 104 (19¢h), li: ae ane 4h (196), and 11: 326. 19 Re Saneydie 2625, cited below, was erroneously cited in Phy- Pista es “G65 (1963) as V. canadensis (L.) Britton. In Shreve & Wiggins (1964) Vv. delticola is described as being found "In sandy, di adobe, clay, u soils, even estone or ky in diverse situations, Sonoran Zone, central Sonora to Tex- as and sou Puebla and Vera Cruz, Mexico. Often in cutlivated aster oo in fie and ore ids hards,." dditional citations: TEXAS: Hidalgo Co.: R. Rumyon 2625 (N, N). VERBENA DEMISSA Moldenke Additional bibliography: Moldenke, Biol. Abstr. 27: 2026. 1953; Moldenke, Phytologia 9: 202. 1963. Mer Pe sens Wilid onal & enended bibliography: Reitz aa te 13% 10. el, Voldenke, Phytologia 10: 10h, 145, 189, 295, 339. é 311 (196) and 11: 20, 2h, 25, 61, 122, & 189 ciseky” "and il: 23, 2h6, 210, 273, 276, 280, 290, & 291. The type specimen of this species, photographed by Macbride as his type photograph no. 1760, was originally deposited in the her- barium of the Botanisches Museum at Berlin, get os now destroyed. C4érdenas found this species growing on rocky s Additional citations: BOLIVIA: Cochabamba: . cdrdenas 5578 (W— 2429778). CHILE: Province undetermined: Née s.n. (Herb. Willdenow 11139; Macbride photos 17160] (N--photo of type). xVERBFNA DISSOLUTA Moldenke Additional bibliography: Moldenke, Phytologia 10: 10) & 498 (196) and 11: 290. 1965. VERBENA DOMINGENSIS Urb Additional bibliography: Moldenke, Phytologia 9: 203 (1963) and 10: (22 217, 219, & 230. 196k. iona nal citations: HISPANIOLA: Dominican Republic: B. Augusto 1047 (a), 1087 (N), 1109 (N). VERBENA EHRENBERGIANA Schau. Additional Eahidogsariert nagar te ft Linnaea 23: 7il--722. 1850; Schlecht. eet gee Nat., ser. : 342——344. 1850; Select. Gui ow. Pl. Ae ‘so & a at Woldenke, Phy- tologia 10: “Toh ‘ “sho (1964) and 11: 338. 1 McGregor and his associates found this espe "scattered in open 456 FHTTOLGCOI«A Vol. 11, me 7 places in dry rocky open wooded hillside" in Nuevo Leén and "scat- tered" on rocky very moist open hillsides in San Luis Potosf. Macbride's type photograph no. 171) is of the type specimen in the herbarium of the Botanisches Museum at Berlin, now destroyed. Additional citations: MEXICO: México: Ehrenberg 713 [Macbride photos 1714] (N--photo of type). Nuevo Leén: ueGregor, Hams, Robinson, R. del Rosario, & Segal 156 (Iw). San Luis Potos!: | Mc Gregor, Gregor, Harms, , Robinson, Re del Rosario, & Segal 906 (Iw). NA ELEGANS H.B.K. Additional bibliography: Moldenke, Phytologia 10: 105 (196k), ll: 25 (196h), and 11: 308. 1965. NcGregor and his associates describe the flowers of this species as "purple" (176), "dark-purple" (51), or "reddish-blue" (254) on open mountainsides, and, in Nuevo Leén, "scattered in dry rocky o- pen wooded hillside" or in "moist open forested area, well scat- tered", Additional citations: MEXICO: Nuevo Leén: McGregor, Harms, Rob inson, R. del Rosario, & Segal 51 (Lw), 176 (Iw), 25h (Iw). ‘san Luis Potosf: | R. L. Me L. McGregor or 16662 (Lw). LEGANS var. ASPERATA Perry Additional bibliography: Moldenke, Phytologia 10: 105 (196k) and 11: 25. 1964; Moldenke in Shreve & Wiggins, Veg. & Fl. Son. Des. 2: 12h). The Gizantoro s.n. [9.66] collection cited in Phytologia 9: 203 (1963) as from an undetermined locality, appears, after a a ering of the longhand inscription by my colleagues at Lund, have been collected in Germany from Sachi which had Beaty from cultivation there. In Shreve & Wiggins (196h) the poser huey of this variety is given as "On limestone ven bare sandstone outcroppings in moun= tains and canyons, in Wurisce da wots va situations but characteris- tic of arroyo margins, ascending to 8,000 feet, chiefly Upper 50~ noran and Transition Zones, Sonora to Texas idalgo, and Zacatecas i Lae the Sonoran Des ert near Hermosillo and Magdalena, — VERBENA ENGELMANNII Moldenke Additional & emended synonymy: Verbena engelm anni Moldenke, Ré- sumé Suppl. 2: 11, in syn. 1960; Steyerm., Fl. Mf Biss, map 18L0+ 1963. Verbena hybrida Pech. ex Moldenke, Résumé pions 11: 8, = syn. 196; [not xv. hybrida Voss, 1873]. Verbena xengelmanii denke ex Henry & -Buker, Tr. Trillia 12: 125, sphalm. 196k. 6 Additional bibliography: Steyerm., Fl. Mo. 1258, map 1840. 1963 Moldenke, Résumé Suppl. 10: 1 (1964) and 11: 1, 2, & 8. 196h; Hem & Buker, Trillia 12: 125. 1964; Moldenke, Phytologia 10: Los es 109, 130, 15h, & 160 (1964), 11: 102, 127, & 201 (196k), and 3-338 & 340-342. 1965, 1965 Moldenke, Monograph of Verbena 457 Recent collectors have found this natural hybrid in flower and fruit in July and August. McGregor found it "scattered among pa~ a ° as Coun and reports its pollen fertility as 1 percent and the a number as n = 7. Morley found it feseseated in creek bottom good loam soil" in Republic County and describes the corolla Pe "light lavender with lighter center" or "light-lavender with lighter throat when fresh". My son and I observed it in kira localities in six states this bees summer and in all case it to be very distinct from both parents not only in the ee of the corollas, but also in its greater height than either parent and its wide-spreading inflorescences. Additional citations: NEW JERSEY: Passaic Co.: A. R. Moldenke 785 (Lw). ILLINOIS: Edgar Co.: A. Re Moldenke 811 (Lw). Macon Coe: A. R. Moldenke 821 (Lw, Ut, "Z, Ze Pike Cos: Ae R. Moldenke 836 (Iw). Scott Co.: A. R. Moldenke 832 (Lw). IOWA: Clayton Co.: rey R. Moldenke 989 (Lw). Iowa Co.: A. R. Moldenke 974 (Lw, Ut). arion Co.: A. R. Moldenke 953 (Lw). Poweshiek Co.: A. R. Molden- ke 965 (Lw). Warren Co.: A. R. Moldenke 947 (Iw, Ut). MICHIGAN: Montcalm Co.: A. R. Moldenke 1036 (Iw). WISCONSIN: Grant Co.: A. R. Moldenke 999 9 (LW, Ut). KANSAS: Barber Co.: R. L. McGregor (Lm). Brown Co.: A. R. Moldenke 912 (Lw). Douglas Co.: J. D. Poindexter 191~23 (Iw). Greenwood Co.: R. L. McGregor 14700 (Lw). Republic Co.: G. E. Morley 80h (Lm), 813 3 (Iw). MISSOURT: Pike Co.: Fritchey 563 (Ib--l175). LOCALITY OF Z COLLECTION UNDE- TERMINED: Collector undesignated s.n. (Lb--27615). EPHEDRO Additional bibLiograpiy: orl Acad. Nac. Cérdoba : 408. 1881; Hieron., Pl. - Fl. Argent. 21). 1882; wig eer Phytologia 10: 106, 122, ce aah ( 196k) a “la: 12.. 196k; Hoc erpt. Bot. A.7: Shi 96h. XVERBENA FABRICATA Moldenke Additional bibliography: Moldenke, Phytologia 10: 106 (196) and ll: 337. 1965. VERBENA FASCICULATA Benth. Additional bibliography: Soukup, Biota 5: 2. 196h; mg Phytologia 10: 106. 196; Hocking, Excerpt. Bot. A.7: 456. NA FERAX Moldenke kt wg bibliography: and 11: 7. 196k. Moldenke, Phytologia 10: 106 (196k) VERBENA FERREYRAE Moldenke Additional bibliography: Moldenke, Phytologia 9: 118. 1963; 458 PHYTOLOGIA Vol. 11, no. 7 Soukup, Biota 5: 2. 196k. VERBENA FILICAULIS Schau. Additional i Moldenke, ph este 10: 106 & 120. 1964; Hocking, rpt. Bot. Ae7: 456 VERBENA FLAVA Gill. & Hoo Additional oe Re A , Anal. Univ. Chil. 35: 191. 1870; Moldenke, Phytologia the Lae, 196. GALAPAGOSENSIS Moldenke Additional bibliography: — Phytologia 9: 12h (1963), ll: 188 (1964), and 11: 317. 1965. GENTRYI Mol Additional litinrenty Moldenke, Phytologia 10: 107. 196h; Langman, Select. Guide Lit. Flow. Pl. Mex. 516 & 1010. 196k. |A GLABRATA H.B.K. Additional bibliography: Soukup, Biota 5: 42. 1964; Hocking, Excerpt. Bot. A.7: 456. 196k; Voldenke, Phytologia 10: 107 (196k) and 11: 317. 1965. VERBENA GLANDULIFERA no Additional bibliography: « Bot. Ae7: 456. 196h5 Moldenke, Phytologia 10: 107 & ne aie (age) gor 11: 273. 1965. The JOrgensen 1026 collection cited in Phytologia 9: 129 (1963) is a mixture with Vv. tenera Spreng. GLUTINOSA Kuntze Additional bibliography: Hocking, Excerpt. Bot. A.7: 456. 196h5 Moldenke, Phytologia 10: 107. 196. h isotype of this species photographed by Macbride as his type photograph no. 1718 was deposited in the herbarium of the Botanisches Museum at Berlin, but is now destroye Additional citations: ARGENTINA: Mendoza: Kuntze 2 [Macbride photos 17,18] (N--photo of isotype). VERBENA GONZALEZI Moldenke Additional bibliography: Moldenke, Phytologia 10: 107 & 301. VERBENA GOODDINGII Briq. Emended s ynonymy: Verbena gooddingii var. gooddingii i Briq-. 6x Moldenke, Résumé Suppl. 3: 39, in syn. 1962; Shreve & Wiggins, Veg « & Fl. Son. Des. 2: 123. Additional bib] dograpty woLdenike in coach = Wige ins, Veg. & fi. Son, Des. 2: 123. 196l5 Hocking, ‘Ss A.7: 456. 196k5 Yoldenke, Phytologia 10: 107, mS, beer leh) is 32, 15% 10 & 200 196), and i: 326. : In Shreve & Wiggins (1961) the distribution of this species is 1965 Moldenke, Monograph of Verbena hs9 given as "In sandy, limestone, and granitic soil ondry rocky slopes a gravelly hillsides, dry sandy washes, canyon floors, ravines, roadbanks, gravel bars, and grasslands So onoran to Arid Transition Zones, southern Utah and N evada. to southern Cali-~ fornia, Sonora, and no orthern Baja California." VERBENA GOODDINGII var. NEPETIFOLIA Tidestr. apes bibliography: eigen 2 Shreve & bd ago bsg Fl. Son. Des. 2: 123-12. 196h; H » Excerp 456. "T96h; Moldenke, Phytologia 10: pea i (196), is "32 # "157 (196), and 11: 326. 1965. In Shreve & Wiggins (196) the distribution of this variety is given as "In granitic soil, dry and rocky slopes hill- Sides, the sides of ravines, on bar of small streams, and along arid roadsides, often associated with Prosopis phe Simo mdsia, and Upper Sonoran Zones, from Arizona and southern Cali for- nia to central Baja California and Nuevo Leén." XVERBENA oldenk Aaeitbouns pabitacanaion Moldenke, Phytologia 10: 108 (196) and 11: 203. 196). sso epee (Cham.) hes _— ional bibliography: Phytologia 10: 108, 210, 211, 216, "Zl, fae Ge (96h) and uy a6 ‘ "162. 1964; Hocking, "excerpt. Bot. A.7: VERBENA GRACILIS Desf. Additional bibliography: Hocking, Excerpt. Bot. Ae7: 456. 196k; Moldenke, Phytologia 10: 108, 112, *379, 182, & 186 (196), tas 13 & 14 (196k), and 11: 256 & 308. 1965. VERBENA GYNOBASIS Wedd. Additional bibliography: Mufioz Pizarro, Philippi 110, ie Moldenke, Phytologia 9: "tac ato. er meded kup, Biota 5: ae The no s ecimen photographed by Macbride as his type photograph no. 1719 was deposited in the herbarium of the Botan- isches ot at Berlin, but is now unfortunately destroyed. citations: CHILE: Tarapac4: R. A. Philippi s.n. (si- baya; marae photos 17419] (N-—photo). VERBENA GYNOBASIS var. STRIGOSA Wedd. Additional bibliography: Moldenke, Phytologia 10: 108. 196k; Soukup, Biota 5: 42. 196k. VERBENA HALEI Small Additional Oe eee ee Jones, Rowell, & Johnston, Flow. Pl. & Ferns Tex. Coast. 1h. 1961; pe Roadside Flow. Tex. 190. paca, Moldenke in eg Wiggins - & Fl. Son. Des. 2: 12338— 1239. 1964; Moldenke, Phytologia 10: "3, 108, 140, 141, 178, 179, 1,60 PRTTOLDOGIA Vol. 11, no. 7 182, 186, Se 193, 210, 211, 216--218, 30h, & 502 (1964) and ll: 13 » een, 1% 16h, 200, & 203. 196; Hocking, Ex- cerpt. rege "7: 56. orollas are eee on Duncan 2-214 as "lavender-purple} on Rowell 779 as "pale-purple", and on McGregor 16762 as “blue”. Recent collectors have found this species in and around aban- doned trick gardens, on low ridges, along ditches, indry rocky mesquite range, in sandy soil of rocky xeric hillsides with Yucca, gia 6 in tight clay loam", and "frequent in black loam". e F. B. Jones 1925 distributed as V. halei is actually V. run- yoni Moldenke. In Shreve & Wiggins (1964) the distribution of Ve halei is given as "In sandy or calcareous meadows, prairies, fields, and w woodlands, on rocky hillsides and along roadsides, Sonoran Zones, from iene to Oklahoma and from south-central Arizona to Qu uerétaro.! Goodman pore that the pale purplish-blue flowers on his no. 7470 are much larger than the flowers of the V. halei of southern Vv. halei is variable. I've known it best in its small flowe form in Marshall Co., Oklahoma." Additional citations: GEORGIA: Sapelo Island: W. H. Duncan 2021, (Lb~-)2773). FLORIDA: Alachua Co.: B. L. Wagenknecht 2599 echt 2599 (Iw). MISSISSIPPI: Jackson Co.: J. Skehan s.n. [Ocean Springs, June 1892; ens pe 109] (Ib—-16h1h). Lb LOUISIANA? Camer racy (lw). TEXAS: Angelina Co.: = L. McGregor 15558 ‘an. w) -_ soaneeioe Ee We Lathrop 2016 2046 (Lw). Montgomery Co.: | Co.: Beamer Sn. {April 30, 1953] (Iw). San Patricio Co.: G. J. Goodman 7h10 (Z)3 F. Be Jones 4558 (Wr); CM. Rowell L779 (Ww), 869 (Ww), We52 (Ww). Webb Cov: Re L L. McGregor 1 16762 (iL ( 7 an VERBENA HASSL Brig. Additional bibliography: Hocking, Excerpt. Bot. A.7: 56. 196h5 Moldenke, Phytologia 10: 108 & 498 (196) and li: 315. ec Peder rsen found this species growing in wet ground and ports VERBENA HASTATA L. Additional synonymy: Verbena hastata f. hastata Steyerm., Fl. Mo. 1258. 1963. Verbena lastata Taylor Taylor & Hamblin, Handb. Wild Fl. 1965 Moldenke, Monograph of Verbena 461 Cult. 294, sphalm. Addi tdnad Sag notihe Schwencke, Zytol. Untersuch. Verbenac,. 3, 12—1h, 19, 26, 33, & 43, fig. 11. 1931; Patermann, Beitr. Zy- 1 Bees East. U. S. 1 (N.C. sag Exp. Sta. Tech. Bull. Js ie gonquin Pro n. Billie Bear Plant List 8. 1961; Steyerm., Fl. Mo. 19671280, m map 1841, pl. 302, fig. 5. 1963; Pinkava, Ohio Journ. Sci. 63: 125. 1963; Taylor & Hamblin, Handb. Wild. Fl. Cait. 168, 257, 279, & a 1963; Selle, Prog. & Abstr. Stud. Research Rep. 12th Annual ann High School Biol. Cong. 10. 196k; Moldenke, Résumé Suppl. 10: : ke 964) and 11: 1-3 & 8. 1964; A. & I. Nehrling, Pict. Book dai nal illustrations: Schwencke, Zytol. Untersuch. Verbenac. fig. 11. 1931 Steyerm., Fl. Mo. pl. 302, fig. 5. 1963; Taylor & Hamblin, Handb. Wild Fl. Cult. 168. 1963. It is of interest to note that b le Schwencke (1931) and oo. mann (1935) report the haploid number of chromosomes for this cles as 6. Poindexter gives n = 7 for his nos. 35, 195-13, and 229-13, with pollen fertility of 93 percent and 98 percent for the Recent collectors in Kansas have found this plant in boggy ground along the edges of springs, in sedge meadows, moist grassy areas, overgrazed pastures, moist sandy prairie ravines, sandy soil in low marshy areas along creeks, low places near small streams, along creek banks in mixed forest association, wet marshy places, Sandy waste and stabilized sanddune pastures, in rocky prairie pas- tures in association with Lactuca scariola, Poinsettia dentata, and Tragopogon, and with Rudbeckia on creekbanks. McGregor reports it from "sandy bluestem pasture, moist ravine, scattered in the area" in Fr river’ Cheyenne ium" of thighete on floodplains in a County, "scattered" in wet sand of marshy areas and "common" in dry loam of grassland yee tures in Washington County, "scattered in moist loam of floodland" in Meade County, "scattered in wet loam of roadside ditches" in Jefferson County, and "common" in artesian marsh in Meade County. In Republic County it was found by Morley "scattered near center of marshy area" and "common along head of lower seepage area". Its corollas are described as "purple" on W. H. Horr }648 and as "blue" 62 PHYTOLOGIA Vol. 11, no. 7 on his no. 697. This same collector regarded his nos. 1436, 4648, and nd L697 | as xV. blanchardi Moldenke and his no. L679 a: as xV. deamii Moldenke . si son and I noted this past summer that when this species in dry pasturelands, as it frequently does in the midwestern portions of its range, it develops a "field form" with the leaf- blades more canescent—puberulous beneath, slightly rough above, and the stem many-branched with many groups of een inflores- cences more or less + nid 1. . the main Examples of this field form may be Moldenk ~ 505, 922, 927, 967, 971, 981, 983, 988, 1014, 1018, 1026, 1035, 1040, Moldenke 21937. Mitchell (1960) reports that the bee, Calliopsis nebraskensis e~ male taking pollen and nectar, but the male taking only nectar Taylor & Hamblin (1963) say that this is a plant for "moist woods" and that "When growing in large colonies, blue vervain casts a violet mist over the ape Single specimens are not so ef- fective since only a few flowers are out at one time. his, tallest of the verbenas, is of pice in a planting of wild flowers and ironweed......This species is found in moist places in moderate- ly acid soil from cia Scotia to British talents south to Florida, and west to Nebraska and Ar of” Pinkava (1963) reports V. hastata from Sandusky County, Ohio. a se ene records it from Ad Adair, Andrew, ig a Audrain, Buchanan, Caldwell, Carroll, Cedar, Chariton, Clay, Dade, DeKalb, ‘ erg Harrison, Holt, Howell, Jefferson, Knox, Laclede, gomery, » Oregon, Ozark, Perry, Pettis, Phelps, Pike, Putnam, fandolgh, eis "Saint Charles, Saint Clair, Saline, Schuyler, Shelby, Sullivan, uri aney, Vern . bster Counties, Missouri, ° the basis of specimens preserved in the herbarium of the Missouri Bot. Garden at St. Louis, not yet seen by me n p. 126 f wor s "A specimen from ounty (pastures, July 28, 1937, D. R. Crookshanks in U. Mo. Herb.) a i. brid between V. bracteata and V. hastata." I have examined this specimen and regard it as xv. perriana Mf Moldenke. The V. L. Harms 903 & 1107, R. L. McGregor 1855, J. D. Poindexter 201-3, and B. ye Wagenknecht 31L9, distributed as ’V. hastata, all appear to be var. scabra Voldenke; D. B. Dunn 3179, 3217, 17, 6138, 6215, & 6703 are Ve macdougalii Heller; W. H. Horr 4691 and Re |e lieGregor 11385 ah xV. rydbergii Moldenke; and Drouet s.n. [July 12, 1931] and “and Be Le Wagenknecht 1890 are Ve stricta Vent. Additional citations: QUEBEC: Chambly Co.: A. Dubois 220 (N)- Or TARIO: Elgin Co.: A. R. Moldenke 1015 (Lw). Essex CO? Ae Be Be as denke 101 (Iw). Haldimand Co.: A. Re Moldenke 1018 (Ly) « “Kent Cov! hi Re R. Moldenke 1044 (Lr). Lambton Co.: Gaiser 38806 (Z); 13876 (2), 1965 Moldenke, Monograph of Verbena 463 4296Bos (Z). Norfolk Co.: A. R. Moldenke 1046 (lw). Welland Co.: A. R. Moldenke 1049 (Lw). Walpole Island: Gaiser 091W (2), Los6w (Z), Liisi (Z), 4136w (2). MASSACHUSETTS: Suffolk Co.: Fe: Allegany Co.: A. R. Moldenke 105), (Lm). Broome Co.: A. R. Molden- ke 1062 (Lw). “Cattaraugus Co.: A A. R. Moldenke 1051 (Iw). Chaun- tauqua Co.: Hayhurst s.n. (Fredonia, | July 30, uly 30, 1902] (Lb-~27613). Chemung Co.: A. R. Moidenke 1057 (lw). Chenango Co.: A. R. Mol- denke 1063 (Iw). Delaware Co.: A. Re Moldenke 106) ‘(um ; Wilkens S.N. s.n, (Stamford, August 5, 1919] (N). Erie Co.: ay Re Moldenke 1050 (Lw). Orange Co.: A. R. Moldenke 1066 (Lw). Rockland Co.: Lehr 4l2a (N), 412b (N). ~ Schuyler Co.: A. R. Moidenke 1058 (Lm). Steuben Co.: A. R. Moldenke 1056 (Iw). Sullivan Co.: A. R. Mol- denke 1065 (Lw). Tioga Co.: A. Re Moldenke 1061 (Lw). Tompkins Cos: |, A. Re Moldenke 1060 (Lr). PENNSYLVANIA: ? Armstrong Cos: He. N. Moldenke 2 21902 (Lw). Centre Co.: A. R. Moldenke 786 (Lw). Clar- ion Co.: H. N. Moldenke 21900 (Lw). Clearfield Cow: H. N. Molden- ke 21895 (Ly). Columbia Co.: H. N. Moldenke 21891 (Lm). ~ Jeffer— son Co.: H. N. Moldenke 21897 (iw). Luzerne Co.: H. N. Moldenke 21890 (Lw). Northumberland Co.: H. N. Moldenke 21896 (Iw). OHIO: Darke Co.: H. N. Moldenke 21906 (Lw). Delaware Co.: A. R. Moldenke 796 (Lw). Harrison Co.: A. R. Moldenke 789 (Lw). Knox Cos: H. N. Moldenke 21903 (lw). Miami Co.: A. R. Moldenke 798 (Lw). Morrow Co.: H. N. Moldenke 2190 (Lw). Tuscarawas Co.: A. R. Noldenke 790 (Lw). Union Co.: A. R. Moldenke 792 (lw). ILLINOIS: Cass Co.: Geyer sn. (Beardstom, July 15)2] (Lb--27610). Douglas Co.: A. R. Ho Moldenke 813 (Iw). Edgar Co.: A. R. Moldenke 810 (Lw). Macon Co.: A. R. Moldenke 818 (lw). Morgan Co.: A. R. Moldenke 828 (Li). Moultrue Co.: A. R. Moldenke 81h (Lw). Pike Co.: A. R. Moldenke 83 (Lw), 838 (Lw, Ut). Sangamon Co.: A. R. Moldenke 82h (Lm). Scott Co.: A. R. Moldenke 829 (Lv). INDIANA: Boone Co.: A. R. Kol- denke 807 (Iw). Hamilton Co-: A. R. Moldenke 80 (Lw). Henry Cos: | H. N N. Moldenke 21910 (Lw). Madison Co.: A. ke ae Moldenke 802 (Lw). Pulaski co.: Friesner 9768 (Lb-~3018h) . “Randolph Co.: rie N. Moldenke 21907 (Lw). IOWA: +: Adair Co.: A. R. Moidenke 938 (Lw). Buc Coe: A. | A. Re Moldenke 986 (Li). Cass Co.: H. Ne Moldenke 21937 (Lw). Clayton Co.: A. A. Re R. Moldenke 988 (Ly). ~ Delaware Co.: A. Re R. Moldenke 983 (Ly). Towa a COet A. Ost A. Re “Moldenke 971 (Lw). Johnson on Co.: Huang 2580 (Lb--5,991) . Tinn Coe: A. Re R. Moldenke 279 (lw), 981 (Ly, Ut). Madison Co.: A. R. Moldenke e 9hl (Lw). Mahaska Co: | Ae Re Moldenke 962 (Ly, Ut). Marion Co Cos: A. Re Mol- denke 95], (Lw). Mills Cos: A. Re Moldenke 925 (Im), 929 9 (Le). Pottawatomie Cos: A. Re Moldenke e 922 22 (LW). Poweshiek Cos: A. Re L6), PHELTTOLOGIA Vol. 11, no. 7 Moldenke 967 (Lw, Ut). Warren Co.: A. R. Moldenke 946 (Im), 949 (Lw). MICHIGAN: Charlevoix Co.: A. R. Moldenke 1027 (lw). Dick- inson Co.: A. R. Moldenke 1026 (Ly) . Grand Traverse Co.: Dieterle 1374 (S). Gratiot Co.: H. N, Moldenke 21945 (Lw). Ingham Co.: Collector undesignated s.n. [Lansing, 8-15-1887] (Lb—~27612); A. R. Moldenke 1037 (Lw). Ionia Co.: H. N- Moldenke 21946 (Iw). Isabella Co.: A. Re Moldenke 1031 (Lv). Jackson Co.: A. R. Mol~ denke 1038 (Lw). Kent Co.: Daniels s.n. [Alto, Jul. 1898] (Ib— 27 27611). Mecosta Co.: A. R. Moldenke 1030 (Lw). Menominee Co.: A. R. Moldenke 102) (lw). Montcalm Co.: A. Re Moldenke 1035 (LW). Osceola Co.: A. Re , R. Moldenke 1029 (Lw). Washtenaw Co.: A. Re R. Mol~ denke 1040 (Lw). Wayne Co.: A. R. Moldenke 1039 (Lw). “WISCONSIN: Crawford Co.: A. R. Moldenke 1005 (Lw). Dane Co.: A. R. Moldenke 1008 (Iw); Rickett s.n. [Madison, July 1h, 1926] (Lb--2760h). Fond du Lac Go.: A A.R. Rs Moldenke 101) (Lw). Grant Co.: A. Re Mol- denke 1000 (Lw). “Green Lake Co.: A. R. Moldenke 1016 (Iw). Towa Co.: A. R. Moldenke 100) (Lw). Marinette Co.: A. R. Moldenke 1022 (Lw). Oconto Co.: A. R. Moldenke 1023 (Lw). Outagamie Co.: A. ae Moldenke 1020 (Lw). Shawano Co.: A. he Ba Moldenke 1021 (Lw). Wau- paca Co.: A. R. Moldenke 1019 (Lw). Waushara Co.: A. R. Moldenke 1018 (Lw). MINNESOTA: Becker Co.: Terrell 1777 (W--2300680). Ot- tertail Co.: P. Johnson 545 (Lb—-36h11). KANSAS: Brown Co.: Ae Re Moldenke 913 (Lw). Cherokee Co.: E.L. Richards 1888 (Lw). Ghey- enne Co.: W. H. Horr 679 (Lw), 1,697 (Lw); R. L. McGregor 13587 — 15221 (Lw). Comanche Co.: R. | Re Le McGregor rv 14676 (Lw). ~~ Dick- n Co.: A. R. Moldenke 906 (Lw). Douglas Co.: R. re oe i ce 93668 (Lw), 14570 (Im); J. D. Poindexter 33 (Lw), 229-13 Sak lis Co.: E. Runyon 280 (Lb--33195). Ellsworth Co.: 6 Ae knecht 1927 (Iw). Franklin Co.: Hetzer s.n. [2) Aug. ae atoT te “im 5 R. L. McGregor 12699 (Lm). Greenwood Co: ot Re L. McGregor 14698 1 Harper Co.: Re L. McGregor 14577 (Lw); A. R. Moldenke ike 890 Ly ponte — etenl Jefferson Co.: We H. Horr 4228 (Lw); A. Re ; Moldenke 905 (Lw). Kingman Co.: A. R Re , Moldenke e 894 (Lw). Kiowa Co.: fa Cot Re he Me meer 15155 (Lw). Meade Co.: Fea Fearing & Latham s.n. [August ; hy. pie ] (Lm); W. He Horr 3302 (Iw), 35h0 (Lw). Morris Co.: Re be cGregor 14105 (Iw). Neosho Co.: W. He Horr 6723 (Lw).- Reno Cot J J.D. Poindexter 195-13 (Iw). Republic Co.: G. E. Morley 197 (I¥), 1062 (Lw). Rice Go.: R Re L. McGregor 12545 (Lw) . ~~ stafford Coot Ae R. Moldenke 907 (Lm); Ungar 349 (Lw), 599 (Lw, Lw), 673 (L¥)- Washington Co.: W. He Horr | (Lw), L648 (Iw); R. Le L. McGregor patric: Co.: E. W. Lathrop 1211 (Lw). MISSOURI: ton Co.: E. J. Palmer 510 (Lb--509h9). Bates Co.: A- R. Molden- = ea ke 891 (Lm). iia Co.: A. R. Moldenke 862 (Iw). Boone Co-! 1965 Moldenke, Monograph of Verbena 465 Daniels s.n, (Columbia, July 1903] (Lb—-193, Lb--h19h, —— Drouet 797 (Lb—~—12568), 846 (Ib--12183); Rickett s.n. [Centra Rd., July 17, 1927] (Lb--7097, Lb--7098). Callaway Co.: ee ler 93 (Lb--51500). Grundy Co.: Crookshanks 147 (Lb—29603). Henry Co.: A. Re Moldenke 865 (Lw). Jasper Coe: Z. Williams s.n. [Jasper, Aug. 15, 1928] (Lb--196). Lafayette Co.: Steyermark 83114 (Lb—-5016). Lawrence Co.: E. J. Palmer 56661 (Lb--50950) . Lincoln Co.: A. R. Moldenke 840 40 (Lw). Maries Co.: A. R. Moldenke 853 (Lw). Nodaway Co.: J. 1 J. R. Singleton 179 (Lb--3459). Saint Louis Co.: Eggert s.n. [Aug. ES 1877] (Lb—-h192). Scotland Co.: Drouet s.n. [Sept. 7, 1931] (Lb—-9465). NEBRASKA: Hall Co.: R. J. Lemaire 228) (Lb—-52671). Nemaha Co.: A. Re. Moldenke 927 (Ly), 928 (Iw). Otoe Co.: A. R. Moldenke 926 (Lw). Richardson Co.: A. R, Moldenke 916 (Im). VERBENA HASTATA f. ALBIFLORA M osaoreing ses Ny ge yl Mold » Phytologia 9: 281--282. neg Ss «, Fl. Mo. 1258, map 181. "19835 Moldenke, Résumé Suppl bs PS 2 136); Moldenke, *Phytolo gia - 1965. teyemark (1963) records this form from Barton and Saint Louis Counties, Missouri, on the basis of specimens preserved in the her- bariun of Fig Missouri Botanical Garden at St. Louis, not as yet seen ENA HASTATA f. CAERULEA Moldenke, Phytologia 10: 490. 196k. Bibliography: Moldenke, Phytologia 10: 490 (196) and ll: 201. 1964; Moldenke, Résumé Suppl. 11: 1. 19 This fom differs from the typical form of the species in hav- Sagan corollas The type of the form n was collected by Andrew Ralph Moldenke (no. 10:3). in a moist swale at Amherstburg, Essex County, Ontario, Canada, on August 28, 196, and is a peaong in the H. N. Moldenke Herbarium at Yonkers, » New Yo There out six husky plants of this distinct color form growing enue “100 of the normal form and 30 of ns rosea Cheney. Only the type specimen has been exam- ined by m Siteticess ONTARIO: Essex Co.: A. R. Moldenke 1043 (Z—-type)- VERBENA HASTATA f. ROSEA Cheney Additional bibliography: Steyerm., Fl. Mo. 1258, map 181. 1963; Moldenke, Phytologia 10: 110. 196h; ahve Sy Résumé Suppl. li: 1 96h; Henry & Buk 12: 125. 196k. in bluestem surg pasture; corolla roseate! Steyermark (1963) records it from Livingston County, Missouri, based on a specimen preserved in ert —— of the Missouri Botanic al Garden at St. Louis, not as yet s y me. Additional lire eit ONTARIO: Essex Co.: A. R. Moldenke 102 (Z). IOWA: Warren Co : A. Re Moldenke 950 (Li). _ KANSAS: Franklin 1,66 PEAY TOLOGIA Vol. 11, no. 7 Co.: Re L. McGregor 12700 (Lw, Iw). HASTATA var. SCABRA Moldenke Additional bibliography: Moldenke, Phytologia 10: 110 (196k) and 11: 201. 1964; Moldenke, Résumé Suppl. 10: 1 (196) and 11: 2. 1964. Recent collectors in ages have found this plant in ne ra- vines, the margins of ponds in prairies, and in sandy loam creek banks. Harms calls it "gcattered™ in sandy loam of vie low area, Wagenknecht calls it "common" in soil of low ground around edges of fields, and Poindexter reports the pollen fertil- ity as 97 percent and the chromosome number as n= 7. Although the above, the variety appears to inhabit the same wet habitats of the typical form rather than the dry pastureland habitat of the "field om", Additional citations: WISCONSIN: Dodge Co.: A. R. Moldenke 1010 (Z). Jefferson Co.: A. R. Moldenke 1009 (Iw). “KANSAS: Barber Co.: Re L. McGregor 14858 (Lw); J.D. Poindexter 201-3 (Iw). Rice Co.: V. L. Harms 903 (Lm). Scott Co.: V. L. Harms 1107 (Lm). Stafford Co.: B. L. Wagenknecht 3149 (Lw). HAYEKII Moldenke Additional bibliography: Soukup, Biota 5: a3 135. ees Mol~ denke, Phytologia 10: 110 (196) and 11: 252-- mer p (196) records the common name Netiioes ccora® “for this HIRTA Spreng. Additional bibliography: Reitz, Sellowia ll: 57 & 13h. 19593 Moldenke, Phytologia 103 110 & us6 (1964) and 11: 239. 1965. Additional citations: BRAZIL: Paran4: Hatschbach & Pereira T7941 {Herb. Hatschbach 1085),] rier ; VERBENA HISPIDA Rufz & P. Additional bibLiogranty: Ken th, Ind. Sem. Hort. Berol. 7. oes Hieron., Bol. Acad. Nac a hy: 408. 1881; Hieron. Pl. Dia Fl. Argent. 214. 1882; are ee Phytologia 10: 110, 15, 192, 210 ’ & 296 (1964), 11: 21, 8h, 86, 148, & 162 (196), and 11: 336. 1965 NA (Covas & Schnack) Mol VERBENA HOOKERIANA or bibliography: Moldenke, rotenceia 10: 103 & HO PoP pe open 127 Additional bibliography: Moldenke, Phytologia 10: lll ie 129, 289, & 3k. 196k. : ; xVERBENA HYBRIDA Voss 1965 Moldenke, Monograph of Verbena 67 Additional synonymy: Verbena eee x coerulea Schwencke, Zytol. Untersuch. Verbenac. 15 & 2 ae Additional bibliography: Schwencke, Zytol, Untersuch. Verbenac. ; Roadside ppl. 8. mm. re rie Moldenke, Phytologia 10: 111 “311, Ess. & oy (seh), the aes and 11: 265, 266, 28h, & 290. 1965; Anon., Time as (5): 42. mn iddltsonar naa, A. & I. Nehrling, Pict. Book Perenn. a9 Sebmencke (1931) says "Da es sich bei V. phlogifiora x V. coe- rulea einen als Gartenhybride geztichteten Bastard handelt, “ist die U Bude des a der in grosser Anzahl au iirebaien Zwergpollen in diesem Falle zum mindesten zweife teee-Die Tar- petenzellen ltsen sich rae wthrend die Pollen gefaltet sind." It should be noted here that the name, V. hybrida Pech., is a Synonym of xV. engelmannii Moldenke. The Agrelius > 3, Hall, Lovejoy, & Maroney sen. [8-18-18], distributed as xV. V. hybrida, is is actually xv. perriana Moldenke. Burpee has developed recently the "Miss Susie" variety of this hybrid with "two rows of petals". o 17 (i). citations: HISPANIOLA: Dominican Republic: B. Augus~ o XVERBENA ILLICITA Moldenke Additional bibliography: Fl. Mo. 1258, map 182. 1963; Moldenke, Résumé Su ty pose I “t196h) and ll: 1 & 2. 196ls5 Moldenke Phytol ogia 10: 111 & 310 (a 11: 117, 196, 200, & 203 (196f), and 11: 328, 337, 338, | & 1965. Poindexter reports t e chromos Sakae of this plant as n= 7 and the pollen bebop ia as 399 andl on his no. 207-19 and 2 percent on his no. 207-22. He collected the plant in waste areas Kansas. Horr found it "common" oan of grassl ington County, while Morley describes it as "scattered" in creek b te bottoms in Republic County of the same state. Additional citations: ILLINOIS: ion Co.: A. R. Moldenke 820 (Lv). Pike Co.: A. R. Moldenke 833 (Lw, Ut, Z)» IOWA: Iowa Co.: A. R. Moldenke 918 (Im (Iw, Ut, Z). Madison Co.: A. R. Moldenke (Ly). WISCONSIN: Crawford Co.: A. R. Moldenke 995 (Lw, Ut). Iowa Co.: A. R. Moldenke 1003 (Iw). KANSAS: Atchison Co.: A. R. Moldenke 910 ( (Lw, Ut). Cherokee Co.: J. D. Poindexter 207-19 (Lw), 207-22 (im). Republic Co.: G. E. Morley y 806 (Lw). Washington Co.: Ss H. Horr 4435 (Lw). MISSOURI: Boone Co.: Rickett s.n. [Rock Bridge, July 9, 1927] (Ib—7095). Henry Co.: A. R. Moldenke 863 (Lm, Ut). Morgan Co.: B. Fe Bush 12927 (Ib—137lh). VERBENA INCISA Hook. Additional synonymy: Verbena incisa Schau. ex R. A. Phil., Anal. 468 PHYTOLOGIA Vol. 11, no. 7 Univ. Chil. 35: 191, 1870. Additional bibliogra phy: yA: eae Ri nce Ee 191. 1870; Moldenke, ue, a a logia 10: 111, 128, 133, 136, 17h, oe , iserlde, e “503 "ase, li: 7 & 138—1h0 (96h)? and 11: 266, wn elts 290, 315, & 1965. XVERBENA INHONESTA Moldenke Additional bibliography: Moldenke, Phytologia 10: le (196k) and 11: 337. 1965. VERBENA INTEGRIFOLIA Sessé & Moc Additional “+ pebangin Voldenke, Phytologia 10: 112 (196) and 11: 158 & 162. 196k. INTERCEDENS dditional bali tertniy: Moldenke, Phytologia 10: 112 & 291 (196) and 11: 86. 196k. RBENA INTERMEDIA Gill. & Hook. Additional bibliography: peg peers ae 112 & 160 (196), 11: 8) & 85 (1964), and 11: “300 & 323. Additional citations: A {ERs Corrientes: i Alan 5435 (S, W—21;32865) , sen xVERBENA KONDAI eelenks Additional bibliography: Moldenke, Phytologia 10: 112 (196k), 11: 47 (196k), nrg 11: 290. 1965. KUNTZEANA Moldenke Additional bibliography: Moldenke, Phytologia 3: 387-388 (1963), 10: 132, 133, 175, & 499 (1964), and 11: 7. 196k. VERBENA LACINIATA (L.) Briq. Additional bilisoprapiy: Kunth, Ind. Sem. Hort. - Te 185; Hieron., Bol. Acad. Nac. peso be le. "18535 “peronsy 08 Diaph. Fl. Argent. 215. 1882; Mufioz Pizarro - Fl. © 200. aoe Sharma & & Mukhopadhyay, Journ. ealiet 58: 359, 3 312, et & 38h, pl. 12 , fie» 53 19635 Moidenke, Résum é opt 10: he Ay Moldenke, Phytologia 10: M2113, 128, 136, 137, 1h6—-1L8, 186, 190, eum 21, 294, 295, 99--301, 307 310° ons 58: sg pl. 12, fig. 53. 1963. he corollas a re described as "mauve" by Sharma & Mukho opadhy ay (1963). Alvarez (a9is ) records the common name "sacha-alfa" for ates plant, but indicates that it is also applied to V. temra preng. The Brttcher & Brttcher s.n. [Rio Pisavil, 30/1/49], cited below, was originally distributed as v. s antiaguensis (Covas & Schnac ck) 1965 Moldenke, Monograph of Verbena 69 Moldenke and was then cited by me as VY. tenera Spreng. It is not conspecific with the other material gathered by these collectors d cited by me under V. cheitmaniana Moldenke. Additional citations: ARGENTINA: Catamarca: Brttcher & Brttcher son, (Rio Pisavil, 30/1/49] (Ss). soi 6 cake i eae VERBENA LACINIATA var. CONTRACTA (Lindl.) Moldenke Additional bibliography: Moldenke, Phytologia 10: 113 (196) and 11: 188 & 189. 196. VERBENA LACINIATA var. SABINI (Sweet) Moldenke Additional bibliography: Moldenke, Phytologia 10: 113 (196)) and 11: 271. 1965. VERBENA BECKI R. A~ Phil. Additional bibliography: Muffoz Pizarro, Es Plant. Descr. Philippi 111. 1960; Moldenke, Phytologia 10: 113. 196he VERBENA LASIOSTACHYS Li Additional Hye pesca Moldenke in Shreve & Wiggins, Veg. & Fl. Son. Des. 2: 122. 1964; Moldenke, Phytologia 10: 113—1 am 186, 210, & a11 (196), 11: 104-106, 127, 136, 119, 140, 141, 200 (asely, and 11: 2h1, 22, & 337. 1965. ve & Mingo (196)5 the distribution of this species is ann 5 sandy or granitic soil of waste places, often in moist places along aie eae and canyons, Sonoran and Arid Transition Zones, northern Baja California to Oregon but barely entering the Sonoran Desert in eastern San Diego County, California. Introduced in New York." LASIOSTACHYS var. SCAERIDA M Additional bibliography: prac aoe pn 9: 462, 466—L67, & 169 (1964) and 11: 105 & 106. 196k. baer LASIOSTACHYS var. SEPTENTRIONALIS M tional biblio ography: Moldenke, Athiet to} 210 & 211 (1961), 11: Bus (1964), and 11: 2h1. th Additional citations: CA RNIA: Ventura Co.: Holzwart & Det- mers s.n. (Ojai Valley, foe 5, 1929] (ib—27616) « — LILACINA — graphy: Moldenke, Phytologia 9: 175--177 (1963), 9: 9: Ltn (196h), “lor Ih ee and 11: 157. 1964; Hocking, Excerpt. Bot. Ae7: 456 VERBENA LINDMANII B Additional ia ie Moldenke, Phytologia 9: 477—L80 (196k) and 11: 121. 196k. VERBENA LITORALIS H.B.K. Additional & emended bibliography: Hieron, Bol. Acad. Nac. Cér- 470 PHYTOLOGIA Vol. 11, no. 7 doba k: 08. 1881; Hieron, Pl. Diaph. Fl. Argent. 21). 1882; gusto, Fl. Rio Grande do Sul 211, 21h, 233, & 235, fig. 102. 196 Mufioz Pizarro, Sin. Fl. Chil. 199. 1959; Reitz, Sellowia 13: 67 & 110. 1961; T. Mey., Op. Lillo. 10: 65 & 125. 1963; Soukup, Biota 5: 135. 196k; cre Biol. Abstr. 5: 8025. 1964; Moldenke, hytologia 10: 114, 151, 160, 182, 210, 217--219, 285, 291, 296, & 297 (196) uu: "86, 128, 183, 186, & 188 (1964), and 1: 317, gee, te 3235 1 Alvarez (1919) records the common name "sange de Cristo" for this species, as he does also for V, peruviana (L.) Britton and V. bonariensis L. He ie the generic name with a rey oo in- itial letter. Soukup (196) records the names "verbena" ‘verbena del campo". The corolla is described as eiebe on MeGre- gor, Harms, Robinson, R. del Rosario, & Segal 721 and on Alston _ 5430, and as Saende™ on M. M,C. . Carlson 3772. ia dipestcte: the species was collected by Alston in waste ground at 900 meters al- it was titude; on found by — at 1100 feet altitude. McGregor his associates found it in an open moist mead 8000 feet altitude, but the yeep a does not appear to be typical. with Quercus at ),800--5100 tent altitude. Additional citations: MEXICO: Chiapas: Breedlove 6089 (Z), 6196 (Z). San Luis Potos{: McGregor, Harms, Robinson, R.del Rosario, & Segal 721 (Lw). VENEZUELA: Federal District: Alston 51,30 (8, 24,0232). HAWAIIAN ISLANDS: Hawaii: M. C. Carlson 3772 (S)- AUS- TRALIA: New South Wales: Gray s.n. [Herb. Austral. 5377] (LW). VERBENA LITORALIS var. CARACASANA (H.B.K.) Briq us pee bibliography: Moldenke, Phytologia 10: 76—78 & 160. ENA LOBATA Vell. Additional & emended bibliography: Resta, Sellowia 11: 57 nee 13h (1959) and 13: 67 & 110. 1961; Mold Phytologia 10: 79-82 (196k) and 11: 239, 243, & ake. 1968. VERBENA LOBATA var. HIRSUTA Moldenk a Joon bibliography: ueiaeike, Phytologia lu: 82--83 & 132- VERBENA LONGIFOLIA Mart oe htttttonal biblicgaaiers Me idhiis Phytologia 10: 83-85 & 10. VERBENA MA CDOUGALII He ue Additional ubllograpty: mane Phytologia 10: eat a & 307 (196) and 21: 200 & 20 1. 1964; Moldenke in S ‘ Lg S, Veg. & . Des. 2: 120 - 1964; Molcenke, hr ae ’ 10: 7 (1964) and ett ns "1964j Moldenke, Biol. abstr. 5: 8025. 1965 Moldenke, Monograph of Verbena 471 ecent collectors have found this plant in the ponderosa pine ieee ae Arizona, while in New Mexico it has bee encountered in sm. ows in stands of yellow pine, with deciduous oaks, two ecletared edits of junipers, pinyon pine, and Pseudotsuga on ns ridges, along running streams in canyons with meadows and cl arings, in wet soil along streams with Abies, yellow pine, and aspen, in pinyon-juniper formation and 5 uniper-pinyon belt, in meadows and along streambanks, some in deeply shaded oaks in assland but on reddish-brown soil. Material has been arms describes the plant as "numerous" in roadside cl pei, cog Fs ponder- osa pine forests. In Shreve & Wiggins (196) the distribution of the species is given as "In open ser a mostly above the Lower Sonoran Zone, from southern Wyoming to western cores vied argo and Arizona; ’ enter ing the Sonoran Desert at Mountain of Tucson, Pima County, Arizona", The B. L. Wagenknecht 711809" d atitetb- uted ved this spe pe is actually V. stricta besos XICO: US (Lb=-526h) « neta Co.: V. L. Harms 1802 (Lw). Otero Co,: De B. Dunn 3157 (Lb--l6329), 3179 - (Lb==52468), 3217 (Lb——l8536) . San Miguel Co.: De. B. Dunn 6703 03 (Lb--39457). Torrance Co.: D. Be Dunn 6138 (Lb--381L6), “6215 (Lb=-1536). ARIZONA: Coconino Co.: Perdue L119 (W—2422173). VERBENA IMA Small Additional bibliography: are yeh Seaside zon : Su. 196k; Moldenke, Phytologia 10: 122—126 (196) and lie 23 a 965. w Menninger (1964) records the common name eset =o ce deed Chan Additional & emended bibliography: Reitz, Sellowia 13: 67 & — 2985 Woldenke, Phytologia 10: 127--129, 289, & 498 (196) : 6 & 60. 196k. haga sa taster el cierantey “You ldenke, Phytologia 10: inperes 17h, 92, 193, “136-199, & 503 (196k), 11: 6, 13k, 135, & 139 (196k), and’ 11: 290 & 318. 1965. VERBENA MENDOCINA Re A. ‘tutor cece Additional ieitocnuaye zarro, Espec Philippi 111, 1960; Moldenk p payealogia io: 36-138, 186, 6 5h, 295, 301, & 313 (196k), li: i 2 (1964), and 11: 28h & 290. "1965. VERBENA MENTHAEFOLIA Benth. Additional bibliography: Moldenke in esises & Wiggins, Lie & jay hy, 1238. 1964; Moldenke, a 10: 99 2, 182, 186 210, 212, as, 286, rh, 7 ie (igek3, 22 2 27 196 « Son 138——1),' & 128 (beh) and 11: 308. 72 PHYTOLOGIA Vol. 11, no. 7 The corolla is said to have been "blue on McGregor, Harms, Robinson, R. del Rosario, & Segal 4,82 and "purple" on Fearing & Thompson 60. McGregor and his associates found the plant growing in Saya depressions of dry rocky open range in San Luis Potosf al "loc olo near a small din Nuevo Leén. Fearing & Phuc found it in red clay soil of dense oak forests. In Shreve & Wiggins (196) its range is given as sd ra- g es, streamsides, and along roadsides, Sonoran Zones, W edge of Colorado Desert to Texas and south into northern ei read te fo d on" axac Addi tional citations: MEXICO : Hidalgo: Fearing & Thompson 60 (lw). Nuevo Le6n: McGregor, Harns Harms, Robinson, R. del Rosario, “& Segal 492 (Lw). San Luis Potosf: McGregor, Harms, Robinson, Re piaiiadisyi pendant Seale aad XVERBENA MERETRIX Moldenk — tbiioererte: Moldenke, Phytologia 10: 145 & 210. 19 VERBENA MICROPHYLLA H.B Additional synonymy: “Lantana microphylla Mart. in Mart. & Spix, Reise Bras. 2: 792. 182 Additional efbtiogreniy x & Spix, Reise Bras. 2: 792. 28; Soukup, Biota 96h; Moldenke, Phytologia 10: Us6— ae & 90-196 seus Te 7.) & 53 (196k), ’and 11: 273, 290, & Soukup (196) records the common names "salvia de la Virgen" and "verbenita del campo" for this species in Peru IFLORA Briq Additional eb imaiene Moldenke, Phytologia 10: 150-151, 159, & 160. 196k. XVERBENA MOECHINA Moldenke Additional bibliography: steyerm - 1260, map 183. 1963; Moldenke, Phytologia 10: e157. (196h) and 11: 117, 16h, 165, 180, i290, 198; 201, & 203. 1964; Moldenke, Résumé Suppl. 10: ° (136 ) and oan oi sel. aree 8 rid growing on i in Arkans Additional t baci, Gace, Elk ie nig Are 883 “(im Ut). Lyon Co.: W. H. Horr s.n. (Aug. 12, 1929) (Lw). Wilson Co+t W. H. Horr s.n. [July 12, 1930) (Lw). MISSOURI: Boone Co.: Dan- iels s.n. [July 1903] (Lb—L167) . Phelps Co.: Rickett s.n. [Yancy Mills, Aug. 13, 1927] (ILb—H170). Saint Louis Co.: Favor 8 Soe [June 16, 1901] (tb—-L185), sen. [June 17, 1901] (ed —Ii86)- stone Co.: B. F. Bush 15697 (Lb—2307). ARKANSAS: Lawrence Co.: ee 5249 (N). Randolph Co.: Demaree 29179 (Lb--h7029), 30963 b—-]6715) . 1965 Moldenke, Monograph of Verbena 473 eePiepenacig a iat pea SMe nal bibliography: Moldenke, Phytologia 10: 122, 151, 1s8--161, “173, & 199° (196k), 1: 59, "155, & 186 (196k), and 11: 322 & 32 3. 1965. dditional sae: BRAZIL: Parand Hatschbach & Pereira 7827 {Herb. Hatschbach 1042] (Iw). ARGENTINA: Corrientes: Pedersen 5074 (W--2132703, ¥--2132708) , oo MORICOLOR Moldenk Additional bibliography Moldenke, Phytologia 10: 133 & 173-- 17h (1964) and 11: 139. 196k. VERBENA MULTICAULIS Raf. Additional bibliography: Moldenke, Phytologia 10: 17 (196k) and 11: 180, 196). VERBENA NANA Moldenke Additional yas Ph Moldenke, Phytologia 10: 175—-176 (1964) and 11: 139. 196k. VERBENA NEOMEXICANA (A. Gra v) Small Additional biblio graphy: raged ae ee 10: a 140, ye 192, 193, 210, 218, 58S, "396, 30h, 306, & 502 Se aa 13--15, 18, & in (196k), and. is "ng, 316, & 337. 1965. VERBENA NEOMEXICANA var. HIRTELLA Perry pcg ry biblicgrapty: Moldenke in tl & Wig » Vege & S. 2: 12h1. 196k; Moldenke, Phytologia 10: “1b, 180-— ih, ro “as, 05, 3 30h, & 306 (196h) li: 13 & 14 (1964), and ; 1965. In Shreve & Wiggins (1964) the distribution of this paar is given as "In dry arroyos, plains and rocky hillsides, oft@m shaded situations, Lower Sonoran to Arid Transition Zones, us ee Arizona, to Texas, Nuevo Leén, and at least to vicinity of El Mérmol, Baja california." OMEXICANA var. ernie Additional bibliography: Mol ee Phytol ogia 10: 140, peng re 285, 306, & 502 (19 yr ae hi. 196k; Mold denke in Shrev ins, Veg. & Fl. Son. J e.-2t i n Shreve & ‘Wieese (196) the distribution of this variety is givens as i fost soil on hillsides, mountain valleys, dry plains, open, s aie to Arid Paneition Zones, southern ornia to Texas, San Luis Potos{, and northern Baja California. Introduced in Florida." XVERBENA NEQUAM Moldenk Additional sinbiegeacters Moldenke, Phytologia 10: 186 (1964) and 11: 290. 1965. XVERBENA NISA Moldenke Additional bibliography: Moldenke, Phytologia 10: 189-190 & 301 474 P.EYTOLOGIA Vol. 11, no. 7 (1964) and 11: 290. 1965. VERBENA NIVEA Moldenke Additional Tit topeapky: Moldenke, Phytologia 10: 147, 148, & 190—-192. 196. ENA NIVEA f. ROSEA Moldenke Additional bibliography: Moldenke, Phytologia 10: 147 & 191— 192. 196). NOTHA Moldenke Additional bibliography: Moldenke, Phytologia 10: 178 & 192-- 193. 196. VERBENA OCCULTA Moldenke Additional bibliography: 0 aa Biota 5: 135. 1964; Moldenke, Soukup (1964) records ae common name "verbenita del campo” or this species. Saunders describes the flowers as ic ae mauve....outer surface of petals mauvish-white" on his no. 961 and "as for 961 but with consi taxable lighter flowers" on on his | no. 962. He sag the leaves "dark green, Baa on upper sur face, lighter green with fine iis hairs beneath," Additional pi bolted PERU: La Libertad: ry & Sagastegui 2718 (W--2l,30832); G. E. Saunders 961 (2), 962 (Z Je NA OFFICINALIS L. Additional synonymy: Verbena aubrietii Kern., in herb. » Bull. Soc e Florul. Cisiord. 33.1 1948; M. L. Buch, U. S. Dept. Agr. Agric. Handb. 64: 13. 1960; Irwin, Roadside Flow. Tex. 190. 1961; Hatus~ , Mem. South. Indust. Sci. Inst. aap Univ. 3 ay 31. 1962; enters "Résumé Suppl. 11: $ & 8. 196; Moldenke, Phytolo- gia i76--178, Si, 186, 194--236, z1in=282, 285, 307, "sh, & 502 (1964), 11: 55, 86 j02 ,, 103 113, 128; 16h, 200, & 203 (196k), and 117 2hi, 2h2, 256, 257, 273 290, ee 336, 30, & 31. 496 ris Pogson sip & e nal illustrations: Schnarf, Ost. Bot. rene 7h: yl 1925; Schwencke, Zytol. Untersuch. Verbenac. fig. 1--3 & 16—-2%- 1961; ae) Beitr. Pharmakog. Borag. 97, pl. 7, fig. 1- 19356 Ss spec and Patermann (1935) report the haploid chromosome number of this 1965 Moldenke, Monograph of Verbena 75 species as n= 6. Reitz (1959) se go the evagy as cultivated B t from t Is —— es eee Seantneeemmmeesaamnenaliliamattttiiemeeeees een tiene ve halei Small; Gordon & Dunn 878 is V. perennis Wooton and Crookshanks 2 is xV. perriana if Moldenke. —— VERBENA OFFICINALIS var. ALBIFLORA Stro pres ere bibliography: Moldenke, SP otha 10: 212 & 277— 2 19 VERBENA OFFICINALIS var. GAUDICHAUDII Briq Ss ag Co hea bibliography: Moldenke, ae ae 10: 217 & 279— 0. VERBENA OFFICINALIS var. GRANDIFLORA Hausskn, : spnshionel bibliography: Moldenke, Phytologia 10: 217 & 280. 96h. VERBENA OFFICINALIS var. MACROSTACHYA (F. Muell.) Ben i Ay aon bibliography: Moldenke, Phytologia 10: a & 280—- 281. 19 _— Ppl arc var. PROSTRATA Gren. bliography: sige sae DG Sats Zina Sees & SS Rae tw (96h) and 11: 257. 196 VERBENA ORCUTTIANA Perry oy hisitdonal Dts cones Moldenke in Shreve & Wiggins, Son. Des. 2: 1240. 196k; rep Phytologia 10: 182, "OB, : 26-986 (1964) and 11: 128. n Shreve & Wiggins Soda Be “the ‘distribution of this Fe er is given as "On tablelands and m » ary ts, and t. f dry pools, Sonoran Zones, sider | Berd Baja ia neat , ae ORIGENES Re A. Phil. ra Mabe Cn aire Verbena origens R. A. Phil., Anal. Mus. Nac. ies 2: 58. lL Additional “eititogrepty: Re A , Anal. Mus. Nac. Bot, 2: 58. 1891; Mufioz Pizarro, Sin. Fl. chi. 1 *% 200. 1959; Mufloz Pizarro, Espec. Plant. Descr. Philippi 110 & 111. 1960; even? Phytologia 10: 286-~288. aps: icidacia, Résumé Suppl. li: 8. 19 According tuflioz Pizarro (1960) the type of V. origenes was "encontré en 108 mds altos ed de Coquimbo, en la cerro del Toro. Herb. Chil. no. agp ag Gay, while that of Ve od origens was "Colec- tada en la quebrada de Paipote, Atacama." It s that the names are just accidental orthographic variants aot sda: other, XVERBENA OSTENI Moldenke Additional bibliography: Phytologia 10: 283-289 (196) and 11: 7e 196). 4,76 PHYTOLOGIA Vol. 11, no. 7 VERBENA PARAGUARIENSIS Moldenke Additional bibliography: Moldenke, Phytologia 10: 292 & 50h. 196). VERBENA PARODII (Covas & Schnack) Moldenk Additional bibliography: Moldenke, Phytologia 10: 293--295 & 313 (1964) and 11: 123. 196k. VERBENA PAULSENI R. A. Phil. Additional bibliography: Muffoz Pizarro, Espec. Plant. De Philippi 111. 1960; Moldenke, Phytologia 10: 298--299. 1 is6h. boo PERAKII Ses & Schnack) Moldenk dditional bibliography: Moldenke, Phytol ogia 10: 189—190, 295, & 295-301 (196), zt 122 (196k), and 11: 280 & 290. 1965. VERBENA PERENNIS Additional babtie ography: Moldenke, Phytologia 10: 183 & 301-- 306 Scag “ 1l: role 6h. Norris this species in "occasional oak, pinyon tivated” in land in open places" in Lincoln County. Material has been mis- identified and distributed in herbaria as V. officinalis L. Additional citations: NEW MEXICO: imei Co.: Gordon & Dunn 878 (Lb—l0886). Otero Co.: Gordon & Norris 5h9 (Tb--5)095) XVERBENA PERRIANA Moldenke Additional bibliography: Steyerm., Fl. Mo. 1261. tg Sarr Ul Résumé Suppl. 11: 1 & 2. 196; Moldenke, Phytologia 10: 210, 218, & 307-313 (1964), 11: 117 & 202 (196k), and 11: 3%. 19 z Steyermark (1963) says "A specimen from Grundy County Nawkagh img July 28, 1937, D. Re Crookshanks 142, in U. Mo. Herb.) appears to be a hybrid between V. bracteata and V. hastata." I have eosin specimen, originally identified as Vv. V. officinalis L., and find rri this it to be « perriana. Additional cltatione: IOWA: Mills Co.: As Re Moldenke 918 (LW). Pottawatomie Co.: A. R. Moldenke 932 (Lm). —yISSOURI: Grundy Co.? Crookshanks 142 (1b—-29613) . XVERBENA PERTURBATA Mol jeisiitional eikitoemnags "waldisiie, Phytologia 10: 295 & 313. ae ana te cy wee ography: Soukup, Biota 5: 135. 196; Mol mn Phytologia 10: 175, 288, 289, Ol, "313, 319, 490, lot, Ca soe, & 50h (1964), 1: 3,'6, 72 9, 13h; 135, 139 (196), and 11: 272, 273, 280, fn 320, & 32le 1965. 1965 Moldenke, Monograph of Verbena 77 Soukup (196)) goes the common name "melindres colorado" and "pampas" te this VERBENA PHLOGIFLORA Cham. Additional bibliography: Moldenke, Résumé Suppl. 11: 8. 196k; Moldenke, Phytologia oe 492-501 (196h), 11: 27, 86, 138, & 140 (196k), and 11: 290. 1965. VERBENA PHLOGIFLORA f. ALBA Moldenke Additional bibliography: ree ae Biol. Abstr. 27: 2026. 1953; Moldenke, Phytologia 10: 501. VERBENA PINETORUM Moldenke Additional bibliography: Moldenke, Phytologia 10: 501—503. 196; Langman, Sd ect. Guide Lit. Flow. Pl. Mex. 516 & 1010. 196). VERBENA PINNATILOBA (Kuntze) Moldenke 2 sacalaalia bibliography: Moldenke, Phytologia 10: 98 & 503. i VERBENA PLATENSIS Spreng. tional bibliography: tor, Handb. Biolog. Data 186. ft Addi eileen rertaieesa. 10: 203 ch (196) and 11: 1--l1, 2h, 25, 1 Spector (1956) reports on the temperature effect on the life span of seeds of this species stored in sealed containers. With a moisture content of 6 percent, the median life span is 3 years and the maximum life span is 6 years at 2° C. At 5° C. the median mi span is 9 years and the maximum life span is 13 gor 4 while s h° ¢. the median and maximum life span is 1 VERBENA PLATENSIS f. VIOLACEA Moldenke ap oar aes. bibliography: Moldenke, Phytologia 11: 6 & 10--ll. 196h. VERBENA PLICATA G dditional Ech terrastyy Moldenke, Résumé Suppl. 11: 3. 196); Moldenke, Phytologia 11: 11—18, fic. 13. 196k. VERBENA PLICATA var. DEGENERI Moldenke Additional bibliography: Moldenke, Phytologia 11: 13, lh, & 18—19. 196). VERBENA PORRIGENS Re Ae Addi tional nt ek a Se ag ON, Phytologia 11: 20-~21 (196k) and 11: 245 & 2h6. 1965. VERBENA PULCHELLA Swi Additional biblio sits Moldenke, Phytologia ll: 21--26 & h6 (196) and 11: 269—271, 273, 277, 278, & 290. 1965. 478 PHYTOLOGIA Vol. 11, no. 7 VERBENA PULCHRA Moldenke Additional bibliography: Moldenke, Phytologia 11: 26--28 & 139. 96h. VERBENA PUMILA Rydb. — rn tg Moldenke, Résumé Suppl. 11: 3. rfc » Phytolo : 26-1, 43, "ub, 48, 100, & 158, fig. (96h) ma ll: 285, 250, 308, & "326, 1965. PUMILA f. ALBIDA Mo ee dditional bibliography: . Bot. Ao7: 55. 96h Moldenke, Phytologia 9% 188 88 (1963) ae “la: 32, 39-—-li2, & 13. VERBENA QUADRANGULATA Additional erp Moldenke, Résumé eee li: 8. 196k; Moldenke, Phytologia 11: 30, 32, 33, 39 a os 48. e "Verbena quandrangulata" cited by m etre het been pub- lished in synonymy in my Résumé Suppl. i, page 7, was actually published on page 8 of that work b gat RACEMOSA Eggert dditional bibliography: Moldenke, Phytologia 11: 21, 23, 32, 13, & 6-50 (196) and ll: 290. 1965. VERBENA RADICATA Moldenke Additional bibliography: Hocking, Excerpt. Bot. A.7: lish. fod rte re Phytologia 11: 50--52 (196) and 11: 249, 252, 273, & sel & VERBENA RADICATA var. GLABRA (Eiken) Moldenke Additional bibliography: Moldenke, Phytologia 11: 52. 196k; Hocking, Excerpt. Bot. A.7: lok. 196k. VERBENA RECTA H.B.K. Additional bibliography: Moldenke, Phytologia 11: 54-56 (196k) and 11: 327. 1965. A REICHII Acevedo de Vargas Additional bibliography: Acevedo de Vargas, Rev. me Hist. Nat. 5k: 45. 1954; Moldenke, Phytologia ll: 57--58. I am omy friend, Benkt Sparre, for ae at- tention to the fact that Acevedo de Vargas (1953) has emended her original discussion of this species by stating that Reiche collec~ ted the type in the Cordillera de Santiago at 2500 meters altitude in January, 1892. RIBIFOLIA var. FOETIDA (R. A. Phil.) Acevedo de Vargas 196l Additional bibliography: Moldenke, Phytologia 11: 60 & 61. 1964+ VERBENA RIGIDA Spreng. Additional bibliography: Moldenke, Phytologia 10: 99 (196h), 14 1965 Moldenke, Monograph of Verbena 79 62—-68, 80--99, 127, 158, 162, & 190, fig. 15 (196), and 11: 266, 322, & 323. 1965. VERBENA RIGIDA var. LILACINA (Benary & Bodger) Moldenke gistttone bibliography: Moldenke, Phytologia ll: 81 & 96—97. 1964. VERBENA RIGIDA var. OBOVATA (Hayek) Moldenke geen bibliography: Moldenke, Phytologia 11: 87 & 97—98. 1964. VERBENA RIGIDA var. REINECKII (Briq.) Mold etme bibliography: Moldenke, erneets li: 84-86 & 98— 99 VERBENA RINCONENSIS Molde Additional wpapieeninas oo king, Excerpt. Bot. A.7: Loh. 196k; Moldenke, Phytologia 11: 99~-100. 196k. ARIA R Peitcewnin sre eee Moldenke, Phytologia 11: 100--103 (1964) and 11: 336. 1965. VERBENA ROBUSTA Greene Adg: Scns bibliography: Moldenke, Phytologia 11: 103-108 & 201. 196). VERBENA RUGOSA Mill. Additional bibliography: Moldenke, Phytologia 11: 108-109, 158, & 190. 196k. a ee a tional bibliography: Langman, Select. Guide Lit. Flow. Pl. es oie & 1010. 196k; Moidenke, Phytologia 11: 112-113. 196k. RYDBERGII Moldenke Additional bibliography: Moldenke, Phytologia ll: 113-—-120, 190, 195, 196, & 200--203. 196). SANTIAGUENSIS (Covas & Schnack) Moldenke Additional bibliography: Moldenke, Phytologia ll: 121--123 & 135" (196) and 11: 273 & 290. 1965. VERBENA S! ahl Additional bibliography: Moldenke, Phytologia li: 84, 86, 102, & 123-133 (1964) and 11: 33%, 3h2, 3ub, & 346. 1965. Naser SESSILIS (Cham.) Kuntze ditional bibliography: Moldenke, Phytologia 11: lil--l\2, Iss 166, 185, & 186. 196). VERBENA SHREVEI Moldenke 480 PHYTTOLOGIA Vol. 11, no. 7 Additional bibliography: Langman, Select. Guide Lit. Flow. Pl. Mex. 516. 196; Moldenke, Phytologia 11: 157--158. 196). VERBENA SIMPLEX Lehm. Additional bibliography: Moldenke, Phytologia 11: 158-181, 190, & 201—203, fig. 16 (196) and ll: 328, 3h0, & 341. 1965. ALBIFLORA Moldenke ona ietboeracty: Moldenke, Phytologia 11: 164 & 179-- 180. 1 VERBENA SIMPLEX var, EGGERTI Moldenke Additional bibliography: Moldenke, Phytologia 11: 16h, 165, & 180—181. 196). XVERBENA SOIBRIGII Moldenk Additional Setiadaentv Moldenke, Phytologia 11: 181 (196k) and 11: 290. 1965. ENA STELLARIOIDES C Additional bibliogr: ea Moldenke, Phytologia 11: 155 & 18h— 187 (1964) and 11: 302 32h,. 1965. FNA STRICTA Vent. Additional eae on Moldenke, Phytologia 11: 158, 162-- 164, & er 3, fig. 17 (196k) and 11: 219-238, 328, 337, 3395 31, & 342. 965. A STRICTA f. ALBIFLORA Wadmnon Additional bibliography: Se estion » Phytologia 11: 200-203 (196h) and 11: 232-237. 1965. STRICTA f. ROSEIFLORA Benke Additional bibliography: Moldenke, Phytologia 11: 201—203 (1964) and 11: 235—238. 1965. VERBENA HUREA D. Don For ape bibliography: Moldenke, Phytologia ll: 242-251 & VERBENA SULPHUREA f. ALBA Moldenke oo bibliography: Moldenke, Phytologia 11: 2h6—2h5. VERBENA SULPHUREA v: ESCENS R. A. P stains tht Geresters Moldenke, Sivan ee li: 2h6 & 2h8. VERBENA SULPHUREA f£. FUSCORUBRA Skottsberg Pe pec bibliography: Moldenke, Phytologia 11: 246 & 2h8— 1965 Moldenke, Monograph of Verbena 481 VERBENA SULPHUREA v TALTALENSIS Moldenke ont rc ere DEliogterky: Moldenke, Phytologia 11: 2h6 & 251. VERBENA SUPINA f. ERECTA Moldenke ME er bibliography: Moldenke, Phytologia ll: 257 & 259-- - 19 VERBENA TAMPENSIS Nash Additional bibliography: Moldenke, Phytologia 11: 262—~26 & 290. XVERBENA TEASII M Additional tipadogtante Moldenke, Phytologia 11: 26l—267, 28h, & 290. 1965. VERBENA TENERA Spreng. Additional bibliography: Moldenke, Phytologia 11: 267—280, 28h, 290, 291, 302, 312, & 320. 1965. ee Tomah eee Regel Soa Wenisoetanies hey prey Phytologia 11: 272, 273, 276-280, 290, & 321. Se VERBENA TENUISECTA Briq. dditional bibliography: Moldenke, Phytologia 11: 290-30), 318, be A 320, & 52k; fig. 18. VERBENA TENUISECTA var. ALBA Moldenke Additional bibliography: Sy wiles » Phytologia ll: 291 & 301— 303. 1965. VERBENA TEUCRIIFOLIA var. COROLLULATA Additional bibliography: Moldenke, Sept gl li: 308 & 310— 311. 1965. XVERBENA TORPA Moldenke Additional bibliography: Moldenke, Phytologia 11: 315--316 & 337. 1965. XVERBENA TRANSITORIA Moldenk ae bibliography: moldsike: Phytologia 11: 290 & 318. XVERBENA TRINITENSIS Mol Additional bibliography: oe Mai, Phytologia 11: 290 & 320-- 321. 1965. VERBENA VALERIANOIDES H.3.K. It should be noted here that the Verbena valerianoides of Saint- Hilaire is a synonym of V. ephedroides Cham. 4,82 PHYTOLOGIA Vol. 11, no. 7 In all, 1) herbarium specimens er 2 mounted phototypes of this species have been examined b Citations: COLOMBIA: Boyac4: parelay & Juajibioy 7632 (N); Lan- nas 3367 (Ca~-78027). Cundinam : Apolinaire-HMarie 706 Mg--50); Hartweg 1350 (Br, Lu, N); "Bolton 507 (T)3 Humboldt & pat pai san. (Macbride photos 3976] (Kr—photo of type, N—p of of type); F. W. Pennell 2329 (N, W--1042255), 2475 (N, thaes Ahan 3 M. Schneider 137 (S); Triana 2052 (Bm). Narifio: Triana s.n. [Pas- to, 1853] (Bm). VARIABILI ey ldenke, Bot. ag eae 16h. 194k. Bibliography: Moldenke, Bot. G : 162 & ah. 1944; Molden~ ke, Alph. List ‘ee 3: 728 (19L9) wt! 7% "1078. 19195 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 7h & 200. 19h93 Ee Je mage 8 Ind. Kew. Suppl. 11: 263. 1953; Tel io “Résumé 85 & 47h. panel herb, 0. i. m. tall; stems tough, subterete slightly _tetragonal, htly puberulent-strigillose, stabreacdat toward the base eget na branches short, very slender, more $ 3 8 Bg 5 g e margins; pe - long, more or less strigillose-puberule ent, especiall y toward the apex, usually with 1 or 2 pairs of much reduced weinertite densely many~flowered; rachis dens g-puberulent; bract~ lets lanceolate, about 5 mm. long, long-acumina + the apex, densely glandular-puberulent with erect spreadin j_calyx a an altitude of 3050 to 3300 meters, in the province of Castrovir- reina, Huancavelica, Peru, on March 27 or 28, 1942, and is depos~ ited in the United § tates National Herbarium "at lope eo The tion. In all, 2 sects Sieg eta, igi the type, and on mounted photograph have been examined by m Citations: PERU: Huancavelica: R. D. Metcalf 30285 (ca-691:968- isotype, N--photo of type, W—-183),978—-type). VERBENA =: re can Moldenke, Known Geogr. Distrib. Verbenac., [ed- 1), 1965 Moldenke, Monograph of Verbena 4,83 Biblio ony Moldenke, Knowm 1 « Ee Distrib, Verbenac., [ed 1) lh, 79, & 102. 192; Moldenke, Cas 13: 119. 1948; *Wolden- ke, Alph. List Cit. 2: 375 (1948) and is 1088, 1091, 1092, & 1293. 19195 4 Aone Boe eogr. Distrib. Verbenae , [ed. 2], 107 & 200. 199; E. » Ind. Kew. Suppl. 11: 263. 1953; Molden- ke, Résuné 138 & ithe 19895 Moldenke, Résumé Suppl. 6: 7. 1963; Mo idenk asa ap 9: 397.6 196). Herb; stems asc » herbaceous, tetragonal, rather densely hirsute ths preading whi te hairs of various lengths; nodes not annulate; principal internodes 1.5--3 cm. long; leaves decussate- ten peduncles 4.5--6 cm. long, spreading-hirsute with stiff ee hairs like the stems; spikes densely many-flowered, 3--3. long and wide in anthesis (including the corollas); Srudiiots lanceolate, about 3 mm. long and 1 m. wide, sharply acute at the apex, the margins long-ciliate with stiff white antrorse hairs; calyx n rtly scurely toothed; corolla hypocrateriform, violet or rose~lilac, ce tube 10——13 mm. long, glabrous on the outside, the limb 6--9 mm. wide, The t of this species was vie oa we! Santiago Venturi (no. 949) —- in whose honor it i n islands in and oil sandy campos, at altitudes from 450 to 900 meters, blooming in January, March, July, Septem- Material has been misidentified and distribu- ted in herbaria under the name V. erinoides Lam In all, 18 herbarium specimens, coon the type, and ) moun- ted photographs have been examined Citations: pa NA: — E. Fielding sen. [Cordova] (Bm). es a. Poe.) “Ta Rioja: Venturi 780 [Herb. Osten 20563] (oes, " ag—-166779, Ug, W—1h3307, W--1591393). Salta: Schreiter 9472 [Herb. Osten 22898] (Ug); Venturi 3677 (W— 1591278), 10301 (Bm, N, 2S Tucumén: Venturi 949 [Herb tant si 172h3; RES Inst. Miguel Lillo 31731] opote. f type, N-type, N--photo of type, Si--photo of type, Ug--isotype, T— eoie35— isotype, rales Lets of type), 2ih1 (Herb. . 17256] (Ug). IFOLIA Hayek in Engl., Bot. Jahrb. 2: 164--165. 1908. Bibliography: Hayek in Engl., Bot. Jahrb. 42: 164—165. 1908 ; 48h PHYTOLOGIA Vol. 11, no. 7 ain, Ind. Kew. Suppl. 4: 25. 1913; Moldenke, Known Geogr. Dis- trib. Verbenac., [ed. 1], 35 & 102 (192) = fed. 2], 7h & 200. 199; Moldenke, *alph. List Cit. 3: 688 (1949) and h: 1112 . 1167. anes Moldenke, Inform. Mold. Set 51 Spec. k. 19563; Mold 6 85 & 47h. 19593 Moldenke, Résumé Suppl. 6: 6. 19635 sich etka, Prytologia 9: 296 (1963) and 10: 103 & 110. 196k. shrublet; branches prostrate, the youngest parts densely Oe core 3 leaves decussate-opposite, cuneately trifid, gradually attenuate into a short petiole at the base, appressed-villous on both surfaces, the middle lobe cuneate and }lobulate at the — the lateral Lobes 2-lobulate; midrib and secondaries prominent beneath; spikes terminal, subsessile, dense, few-flowered, at first fastigiate, later decoming shortly cylindric, the rachis e t he ute at the apex, villous i rege ise 4 mm. long, appressed-pilose, 5- dentate, accrescent rolla rose or lilac oo violet or even veer sh, only atentay. Sodas than the calyx; w spl sise ini not ap ed; cocci testaceous, about 3 mm, long, Perobinttlat’ type of this diminutive species was collected by August inten (no, 258 2582) along a pera at about 3900 meters alti- tude at La Oroya, Junin, gets and was deposited in the herbarium of the aaay’ of Fant t Berlin, now destroyed, but where it was fort graphed acbride as his type photograph M no. 17458. pivek rsney says of it: "Ein kleines Spalierstr#tuch- lein aus der nichsten Nectiarncentat der V. inflata Kth., im Habi- tus aber der V, microphylla sehr *hnlich, “aber durch die dichte Be- haarung und den Mangel ree Konnektivanhtngsels leicht zu unter- schieden. Von V, inflata, der sie sich in der Ausbildung des Kelches an meisten nzhert, = sie durch den Wachs, die viel weni- Se tief geteilten Blatter und die dichte Behaarung verschieden. iit Tt sie auch der mir unbekannten Vv. pogonostoma se, stehen, doch hat letztere nach der Bes chreibung scmmalers, flederepaltige Blattabschnitte und sich nach dem Verbltthen stark verlungernde Inflorescenzen.*" Verbena ene inhabits grassy steppes at altitudes of 10,000 to 13, es feet, and has been collected in anthesis in March, April, Septemb and November, in fruit in September. Materi has been misidentified and distributed in herbaria as Vv. cuneifolia Rufz & Pav. and as V. hispida Ruiz & Par. ve Asher, Kates, & Cia. 18003, cited below, is referred here only doubtfully. all, 9h erbasiun specimens and 2 mounted photographs of the pan eetlortien have been examined me. Citations: PERU: Ancash: Cerrate 2567 (Ok). Cuzco: Rose & Rose 19072 (W--761654). Huancavelica: Tovar var 7143 (Z), 119k (Ss) - ~ dunia: A. S. Kalenborn 22a (W--104)270); Kunkel 37 372 (Gg); Fe L. Stevens 7 (W~1199960); Weberbauer 2582 [Macbride photos 17)58] (Kr-—photo of type, N--photo of type). Department undetermined: Asher, Kates, & Cia. 18003 (Kr); Collector undesignated sen. (N). 1965 Moldenke, Monograph of Verbena 485 VERBENA WEBERBAUERI Berek in Engl., Bot. Jahrb. 42: 163. 1908. Biblio erephy: Hayek in Engl., Bot. Jahrb. 2: 163. 3308, Prain, Ind, Kew. Suppl. 4: 245. 1913; Moldenke, Known Geogr, Distrib. Verbenac., P23 1j, 35 & 102. hog eer Castanea 13: 116 & 117. 1948; H. N. & A. L. Moldenke, Pl. Life 2: 43 & Lh. 1948; Moldenke, Alph. List Cit. ree 688 (2949) ‘or 4: 12 ie 1Aghe 19495 Moldenke Geogr. rib. Verbena & 200. 1995 Soukup, Biota 1: 181. "1956; Moldenke, Résumé ge nig, “ atl 1959; Moldenke, Résumé Suppl. 8: 2. 1964; Moldenke, Phytolo 10: 148. 196). Very small shrublet; roots woody; stems appressed to the ground, 5--10 om, long, simple or branched; leaves decussate~opposite; pressed setae on both surfaces; pertiar cylindric » 2-~3 cm. long, dense; bractlets past Pgs cpa gee than the calyx, ve at the apex, setose-hispid; calyx o 2--3 mm. long, shortly toothed; corolla small, lilac or Sltishakate: 3--l, mm. long, The type of this tiny species ° was collected by August ace aaa (no. 40) -- in whose honor it is named -- in open mats at 3700 meters altitude at Pucard, a station on the Puno--Cuzco ora and was deposited in the herbarium of the Botanisches Museum a he species has been found on dry cliffs, at 3700 meters alti- tude, flowering in March. Material has been pee and distributed in herbaria as vy. microphylla H.B In all, 2 eee specimens and 2 ssanted choles have been examined by m Citations: PERU: “Puno: Weberbauer 1,0 [Macbride aay 17459] (Kr—~photo of type, N--photo of type). BOLIVIA: La Paz: Buchtien i102 (N, S). nay XVERBENA WINGEI Sangre Bot. Gaz. 106: 164. 194. Synonymy: Verbena tenera x aubletia Winge, Proc. Linn. Soc. Lond. 150: 236. oa. Yerbene aubletia Jacq. x V. tenera Spreng. ex Moldenke, Résumé 358, in sy! syn. 1959. Verbena canadensis (Le) Britton x V, temuisecta tofioge ex Moldenke, Résumé 360, in syn. 1959. Verbena tenuisecta Briq. x V. canadensis (L.) Britton ex Moldenke, Résu Résumé 376, in Bring 1959. Bibliography: inge, . Linn. Soc. Lond. 150: 235. 1938; s. A. ie "Fo und. Pl. oe "335. 19h; Moldenke, Bot. Gaz. 106: 16h. red Yoldenke, Alph. List Invalid Names Suppl. 2s 27 nie Mol- own Geogr. Distrib. Verbenmac., [ed. 2], 165 & 200. 19,9; . J. *daliab:. Ind. Kew. Suppl. 11: 263. 1953; Noldenke, Am, Mid). 486 PHYTOLOGIA Vol. 11, no. 7 t. 59: 363. 1958; Moldenke, Résumé a 358, 360, 316, & h7h. 1959; Moldenke in Chittenden, Roy. Hort. Soc. Dict. o 6¥ 2213. 19635 Moldenke, Phytologia 8: 121 (1961), 8: 435, ion 8 & hhO (1962), and ll: 271 & 65. his is the hybrid between v. canadensis (L.) Britton and V. tenuisecta Briq., produced artificially by Winge in Engian din 1933. Its characters are intermediate between those of the two the hybrid is to be expected throughout the area. VERBENA WRIGHTII A. Gray, Syn. Fl. N. Am. 2 (1): 337-338. Shy Synonymy: Verbena pinnatifida M. E. Jones ex Moldenke, 1 List Invalid Names 10, in syn. 191 [not V. pinnatifida Daly 1791, nor Nutt., 1941, nor Sohen., 1895]. Verbena aubletia var. bipin- natifida Fendler ex Moldenke, Résumé Suppl. 3: 36, in syn. 1962. anes bipinnatifida var. lata ifolia Perry ex Moldenke, Résumé 1. 3: 36, in syn. 1962 [not V. bipinnatifida var. latifolia Schell, 1960}. ae ore eg Wats. sen ey py Sei. 18: 1%. "1883; ide Cont: rad iy Ue Se Nat. Herb. 2: 38. 1892; Briq. in Engl. & Prantl, *Nat. Pflanzen- fam. (3a): 148. 189); Jacks. in Hook. f. & Ind. 2 2: acks., 1180. 1895; Eggert, Torreya 2: 123. 1902; Shreve, Carnegie Inst. Wash, Publ. 217: lif. 19 915; Perry, Ann. Mo. Bot. Gard. 20: 28, 312, 327-329, 331, & 356, piv. 319335 dane, » Syab- Bot. Upsal. bt & lk. 1935 Cory, . Sta. Pull. $90: 89. 19373 203, 226, 242, 246, 22, 270, 279, 280, & 295--207. 1916; Moldenke, Phytologia 2: 327, 329, & 338 9h?) be ‘Ss 478. eg yoldenke, Wrightia 1: 233. 19h; H. N. & Ae Le Moldenke, Pl 7 1948; Moldenke, Alph. List Cit. 2: 302, 400, 107, oe, 50, 467, 1965 Moldenke, Monograph of Verbena 87 550% 598, 617, & 639—6h:1 4 Te, 722s 7 3 (iste), and 4: 990—-992, 996, 997, 100k, 1086, 1093, 1107—1109, 111, i115, 1138, 1139, 1241; 1142, 48, 1156, 1151, sh, us%, 116, 1167, 117k, 1203, 1213, 1223, 1225, 1226; 1229-~1231, 12h0; 1243, 12kh, 1252, 1255, & 1269-1291. 19/95 Moldenke, Known Geogr. Distrib. Verbenac., i Se 17, eS, 1949; Moldenke, Am. + 292 & 450. 19493 Moldenke, Phytolo- gia 3: 73 & 132 (1sh9) ete 3; 458. 1951; McDougall & Sperry, Pl. Big Bend Nat. Park 146 & 148, fig. 146. 1951; oe in Chitten- den, Roy. Hort. Soc. Dict. Gard. vr 2209 & & 2213. 19513; Moldenke, Inform. Mold. Set 8 Spec. []. 195); Klots, Desert Life 7, 8, & 61, fig. 8. 195); Moldenke, Résumé 21, 25, 26, 30--32, hO, 119, ris He bey & 474. 1959; Irwin, Roadside Flow. Tex. 189. 1961; Lewis ver 1 Moldenke, eas aves 8: 1. — Moldenke in Shreve & seins, Pics Perry, Ann. - Gard, 20: pl. 13. 1933; McDougall & Sperry, Pl. Big Sint oe "park 148, fig. 1y6. 1951; Klots, Desert Life eae 8. 1954; Lewis & Oliver, Am. Journ. Bot. 8: 6h0, fig. 5. 1961. rect annual ie o 60 cm. tall, often globose or in clumps, branched from the 2 Be iehad spreading, with a short taproot; erect, branched, sparsely hispid-hirsute; leaves subsessile or contracted into a short margined petiole, decussate-opposite; leaf-blades 2—l cm. long, bipinnatifid or trifid, with the divis- narrow hirtellous or hirsute on both surfaces; spikes short~ pedunculate, more or less compact in fruit; bractlets lanceolate, shorter than the calyx, acute or acuminate at the apex, hirsute- ciliate; flowers fragrant, yess when in bloom; corolla hypocrat~ erifom, varying from pink, rose-pink, lilac-pink, ails hg at- tractive bright-pink, bright Sasa ahowtnie pinkish-lavend deep pink-lavender, pinkish, or reddish to lavender, abkcietatbir, rose, rose-purple, ’ purplish, purple, light-purple, Cerise, mauve, magen or even blue, its tube 11--12 mm. long, pubescent around po ee the limb 6--8 mm. wide, the lobes retuse at the apex; rui ekent aces muricately scabrous; chromosome number: n 488 PHYTOLOGIA Vol. ll, no. 7 The Pf of this species was collected by Charles Wright (no. 150), in part) -- in whose honor it is named -- near Frontera, on the borders of Texas and adjacent New Mexico and Chihuahua, and is (1895) ascribes the species, from its original publication, to "Mexico", Parry, Peeeln, Wright, & Schott s.n. in the Britton Herbarium is probably a topotype, collected on March 22, 1852. The specific epithet is often uppercased. ype of V. pinnatifida M. E. Jones was collected by Marcus Eugene Jones in . Bernalillo County, New Mexico, and is deposited in the herbarium of Pomona College; that of V. aubletia var. bipin- natifida was gathered by August Fendler (no. 586) in waste fields and low valleys in the mountains of Santa Fe County, New Mexico, and is deposited in the herbarium of Dartmouth College at Hanover, New Hampshire; and that of V. bipinnatifida var. latifolia Perry was collected by R. M. Scholl (no. 11) in Culberson County, Texas, and is deposited in the herbarium of the University of Texas at pack (but the same trinomial accredited to Scholl himse on his no. 8, is V. ambrosifolia Rydb.). The V. pinnatifida of tountsk is a synonym of V. hastata L., while those e of Nuttall and of Schauer are V. bipinnatifida flutt. E. D. Schulz 057 has a notation on its label to the effect that the "flowers [are] mich deeper in color than those of V. bi- pinnatifida." For the benefit of those interested in the color variations of this species [or, what may be the more like ly situa- tion, the color-concepts and color—definitions of the collectors!], I give herewith the corolla color as recorded on various of the cited collections: "lavender" - Lundell & Lundell 10222, Nelson & Nelson 983; "rose-lavendert - Archer 3427; Mpinkish-lavender" - - Lundell & Lundell 10177 & 102h4; "deep pink-lavender™ - E. Re Schulz ; Sen. (near Alpine, Aug. 4, 1928]; "pink" - Lundell & Lundell 10255, MeVaugh 12300; "pinkish" - Nelson & Nelson 5017; "deep-pink" - Correll & Ro & Rollins 23676, Lundell & Lundell 17 16941; "rose-pink" - Lundell & Lundell 13127, 13185, & 14339; "bright purplish-pink" - McVaugh 7868; "attractive bright-pink (not purplish)" - McVaugh 7508; "lilac-pink" - H. N. Moldenke 10080; "rose" - Ferris & is & Duncan 2647, C. He. Mueller 8215; "rose-purple" — - E. D. Schulz s.n- “(8731/- 1931], Pe Ge Standley 0625; "reddish" - R. L. MeGregor 167125 "purplish" — [undell | & Lundell 10247; "purple" -- Whitehouse jouse 195375 "light=purple" - D. W. Lee 3; "cerise" - Correll & Johnston 1BLO25 and "blue" - Stanford, Retherford, & Northcraft 2 Little & Campbell (19]3) Fesard ve ambrosifolia Tx ngs as a syno~ nym of V. wrightii, but, although they are admittedly very closely related, I feel that both are worthy of being upheld. Specimens th intermediate characters may prove to be hybrids. Verbena wrightii has been found growing on plains and outwash 1965 Moldenke, Monograph of Verbena 89 es dryish gravelly banks and dry flats, on rocks and ridges, gravel bars, on sandy plains and rolling gras slands, on rocky c walls >» and the Saciite of mountain streams. Collectors have en- 7 dry valleys, dry gra creekbeds and dry draws, arid aeolian deposits, deserts and desert washes, rolling or desert grasslands, fo othills and valleys, sandy draws in the hills, burrow pits and desert areas, in the juniper belt, in washes at the base of buttes, and in eravel and sand beds on rocky slopes, at altitudes of 2100 to 7800 feet, flowering and fruiting from February to November. In Colorado it was found by aso ridge "in dry light adobe clay soil on gentle southwesterly slope, ena er meee area, associated with Juniperus monosperma, Pinus tion with Artemisia spp., Eurotia, Boatalous, ot a ite. re percent", of mesa in black volcanic rock and soil derived from it in dry xeric plateau." In Texas it is said to be found generally in fields, flats, — dry gravelly creekbeds and bars and on dry or semi-desert hi sides and slopes, blooming from February to November, in the wes- tern and central sections of the state from El Paso and Presidio to Bexar and Guadalupe Counties. In Jeff Davis County it is de- scribed by Hinkley as "frequent", by McVaugh as "common....{in] und", Hangon calls it "frequent on rocky hillsides" in sidio Comty, while York found it there — creosote-blackbrush in buff silt uvi- Prosopis association" in Reeves, "in Yucca-Agave association" in eee, Paso, and "in gypseous soil on level plain in Prosopis- Gutierrezia association" in Culberson County. WM. a Young calls it "common in open valley"; Cory says "infrequent ghway ps ye a os monn and Warn ock calls it jabebeared! or "infre- soil" in oe County. The Jast-named collec- " f ent quent along highway in low ASiankcie hills" in Hudspeth, "frequent on limestone flats" and "frequent on rocky igneous soil" in Brew- ster, "infrequent in rocky igeneous soil" in Jeff Davis, “infre- 490 PHYTOLOGIA Vol. 11, no. 7 quent on limestone Blopee® in El Paso, "frequent along highway in red sandy soil" and "abundant" in Presidio, "frequent" in Reeves, "desert grasslands with some Nolina" and "on stony Sistas: are Nolina and Acacia"; P. C. Standley says that it is "common" in Slaughter Canyon in the Guadalupe Mountains. Benson reports it gor rep it as "scattered in dry ", while Stan- , Retherford, & Northcraft found it "in playa valleys with con- siderab surrounding th Larrea, Acacia, zona near Same to Colorado, Oklahoma, and T to San Luis Potos {." G. J. Harrison 1960 has the pubescence of the calyx and bract~ surpa, distributed as "Ve Wiptndatd Pada x wrightii", seems to be V. am- brosifolia Rydb. Eggert (1902) states that V. wrightii hybridizes with V. zeosnons Eggert, but I have not as yet seen any specimens of such a hybrid, nor does he cite any. Verbena wrightii is called "desert verbena" and "Wright's ver~ na". Wy Wyman & Harris (191) report that the Navajo Amerinds make m this plant, called e.?ncj folia Rydb., V. ambrosifolia Rydb., V. ambrosiifolia Rydb., Ve jag bletia L., me aubletia (intermediate)*, V. bipinnatifida wittc. V. bipinnatifida x wrightii Wooton, V. canadensis Britton, V. cili~ ata Benth., V. ciliata var. longidentata Perry, V. ciliata var. pubera (Greene) Perry, V. gooddingii Briq., V. goodingii Bria-, Ve neomexicana (A. Gray) Small, V. pinnatifida Lan., V. pubera Greene, Ve. pumila Rydb., V. supina ‘tae V. teucriifolia Mart. & Gale, and even Labiatae. cites oe oe On the other hand, the Archer 739, Arsane 21865, Hitchcock & Stanford 6782, 0. B. Metcalfe 126, C. C. H. Muller s.n. [Cat- tail Cm 1965 Moldenke, Monograph of Verbena 491 June 20, 1901], B. H. Warnock W.283, Wooton & Standley 3635, & Mueller 572, and Me S. Young s.n. [Davis Mts., 9/18/18], dis- tributed as s Vv. wrightii, are actual ly ¥. ambrosifolia iegetises the Edw. Palmer 1050, distributed as "V. wrightii Gray var.?", is al- so V. ambrosifolia; Holzinger s.n. [Santa Rita, Aug. l, 1912] and Peebles 1423h, MacDougal 317, Shreve 4367, Townsend & Barber 139, Urs. C. C. D. Walcott s.n. [Grand Canyon, Aug. 1938), | and Warnock & McBryde 14866 are _ are V. en np Griffiths 4833 is V. is v. bi~ me ee eenaennenietanmminemeingy tween men et em te een ea v. ciliata var, pubera (Greene) Perry; Barneby 2806, Eggleston 10823, and E.G. Marsh 1225 & 1258 are V. elegans H HB Kj E. W Nelson 3896 a: and d Edw. Palmer 1052 are ue "elegans v var. asperata Per- ry; ry; Borel] s.n. [Bonita Canyon, Sept. 8 » 1935), Cutler 2002, D. R. Goddard ie 5 & 825, Goodding 76-52, Griffiths 439, Ca¥. Harbison 41178, 1 r, W. Jones sone s.n. (Dragoon, Aug. 18, 1920], Kearney & Peebles 9720, Tae & 12669, P Peebles 6732, Hooke s.n. [Kingman, ngman, Aug. 13, Ill}, and ES sen. [Tucson, May lay 16, 1896] are V. gooddingii Beh-i 8.n, | Poaceae: Aug. 1882], and Mearns arns 190 & 191) ar are V. gooddingii var. nepetifolia Tidestr.; while B. Ba He Warnock TL &T. 45 are V. wrightii f. albiflora Moldenke. C. C. Ellis 17 and Wooton 364 are mixtures with V. ambrosifolia; s 3801 B. H. Warnock 1.66 is 4 mixture with V. ciliata; M. E. Jone and T: Tracy & E & Earle 162 are mixtures with V. ciliata var. puters Weneurtin Sen. [5-2-1916] is a mixture with ve .V. pumila Rydb Ay Ne Nelson son 11338 has cr identified by me e in some Saat a as V. anbrosifolia; Eastwood 8468 in the herbarium of the California Academy of Sciences and J. Skehan J also gg resenble V. ambrosi- folia and may be that taxon or else a hybri ~ Tharp 46090 in the Stockholm herbarium “Anseribed "Dona Ana Co., Texas" in error. The P. C. Standley 0625, cited below from Otero County, New Mexico, may ay have been collected in Chaves or in Eddy Counties -- its label is inscribed "Slaughter Canyon in Guada- 92 PHYTOLOGIA Vol. 11, no. 7 or Pinal Counties, Arizona -- the label in inscribed "Reddington— Mammoth"; Eastwood 15697 may be from Bernalillo or McKinley Coun- ties -- the label is inscribed "between Albuquerque and Gallup"; Worth & Priest 5h0b may have come from Lincoln or Socorro County, New Mexico — the label reads "between Carrizoso and Socorro"; Tharp 3691, collected in Paisano Pass, could be ges either Brew- ster or Presidio County since this pass leads from one to the ot- her. Cottam 102), is labeled "Old Fort Butterfield, venat, and was pr obably collected somewhere along the route of the Butter- field Overland Mail (18581861) which followed a route from Gray- son through Cooke, Jack, Nolan, Upton, and Pecos Counties to El Paso. Dr. Geiser thinks that the locality refers to the oe collected Verbena specimens re akg Contrib. U. S. Nat. Herb. 2: 328 reference in the bibliography of this species is often cited as "189", but appar- ently should be 1892. Perry (1933) cites the ap lst 3 additional specimens not as yet seen by me: COLORADO: La a Co.: Baker, Earle, & Tracy 531 (E, G). TEXAS: Brewster Co.: pen & Steyermark k 3301 (E)- Cul- berson Co.: Eggert s.n. [flats near Van Horn, 13 May 1901] (E, G); Young s.n, [Guadalupe Mountains, 9 Sept. 1916] (E). Hudspeth Co.: Havard 97 (G); Thurber 143 (G). Jeff Davis Co.: Ferris & Duncan aie (E); E. J. Palmer 30672 (E); Tracy & Earle 162, in part rt (E, F, G G, W). Presidio Co.: H. H. C. Hanson 551 (E). Ferrell Coe: Eo Je Palmer 33583 (E). Ward Co.: Tracy & Earle 61 (E, F, G)- county — undetermined: C. Wright 150, in part [Fronteras] (G--type), S-0e S-n- [mountains near © Fronteras, March 1852] (E). NEW MEXICO: Berna- Lillo Co.: C. C. Ellis 17, in part (E); Kammerer 33 (E). Dona Ana Co.: P. C. Standley s.n. [Organ Mountains, 11 June 1906] (E)- Grant Co.: Eastwood 8L68 (c); Eggleston 17231 (E); E. L. Greene S- n. (near Silver City, , 1880} (E); 0. B. Metcalfe 126, in n part art (EB, G); Pilsbry s.n. [Gallinats Canyon, Black Range Range, at Aug. 1915] (D)- Lincoln Co.: Wooton 364, in part (E, G). Otero Co.: 0. B. Metcalfe 1090, in part (E, G); Rehn & Viereck s.n. (Alamogordo, 7 April--2 May 1902] (D, E); Viereck san. [High Rolls and vicinity, 21--28 May 1902] (D). San Miguel Co.: Mulford 37 (E). Santa Fe Co. Hel- ier & Heller 35% (£, G). County undetermined: Wislizems 1,83 {Rabbit Ear Creek] (£); C. Wright 1503, in part [1851-52] ()- ARIZONA: Apache on ae 1,22) (E). The Bigelow s.n. which s cites from "NY" i ually in the Columbia University soctastim} her "Heller 3536" in nit same collection cited below as Heller Heller 3536, and the Wright s.n. which she cites as from “Fronteras and deposited at "Ny" is the Parry, Bigelow, Wright, & Schott s-M- cited below. The Clawson 13085 Shick abe eater is” 3 regarded by me as representing V. ambrosifolia, Her Ellis 17, Wooton xh, and Met- 1965 Moldenke, Monograph of Verbena 493 calfe 126 & 1090 appear to be mixtures with V. ambrosifolia. The H. S. s. Barber 106, Griffiths 4757, W. Hough 109, Peebles 5353, Tou- mey 304 1/2, S58 Stokes as [Snowflake, July 30, 1897], cited both by | Perry and by me as V. ciliata Benth., were all annotated by T. He woh ae the United States National Herbarium as "Verbena Wrightii ? Perry 9a) says of V. wrightii "On account of its strong var- liability, this species has been exceedingly difficult to define. It was first set apart as erect annual, with anther-glands as The characte appear to be the glandular somewhat viscid pubescence hil nog Pini the very short acute-subulate calyx-teeth, and t pact spikes." In all, 5) herbarium specimens, including type material of most of the names involved, have been examined me. Citations: KANSAS: Finney Co.: W. R. Dudley s.n. [near Garden City, June 10, 1896] (Du-~2171). COLORADO: Boulder Co.: A. G Vestal 373 (Du-—10526) « El Paso Co.: R. B. Livingston 3131 co 63112). Fremont Co.: Degener & Degener 27016 (N, Z)- - (N, Z)- La Plata Co.: Baker, Earle le, & Tracy 531 (Cm Cm, Du--1517 797, Mi, N, Po--63889, W~~337 397) ; Eastwood 5368 (Ge--32289); Ferril s.n. (May 11, 1906] (Bl--,2301); Loughridge 461 (Bt-~2h702); Re W R. We ~ Pohl 2002 (Up); Remaley 16163 16163 (B1—-3765); 1 Way s.n. [Oxford, 1916] (Bl—-2358). Las Animas Co Co.: Herb. State Agric. Coll. 418 (N). County unde- penchant OEY A, Sconce Ta Cited ntem ra eee ae enna a Sem = Stevens 18h (5065886) . ~~ Harper Co.: G. J. Goodman 239 i os a 350). TEXAS: Bexar Co.: Havard s.n. [San Antonio] erick) - rewster Co.: Collector undesignated d L.1~3 (Sr); Ferris & Duncan 2876 (Du—12,214), 3000 (Du--12L09); G. L. Fisher s.n. (Alpine, Aug. 23, 1932] (Wi); a Univ. Texas s.n. (Marathon, 6/4/31] (au, Au, Au); Lewis & Oliver 5451 (Nb), 545h (Nb); C. L. Lundell 13185 (N, Rf); Lundell & Tundell lo2hh (1d, N, Dorgan EP Be Mc Dougall s.n. (10/5/35) (iv); R. | McVaugh 7 Ar--233632, Ar— 178200 200, Mi); A. as Moldenke 165 (Fg); M j Moore & Steyermark 332) (Ca— 471228, pe 22li78 , Gg--19h 780, Mi, N, W--15667L9); C. H. Mueller 8140 (Au, Mi, ,N); L. T. Murray s.n. (Chisos Mts., July 2, 1938] (Ar—-219761); ; chen & & Nelson 5025 (Ka); Parks & oe 6471 (tr), 7337 (Tr), 9B (Tr); Rose-Innes es & Warno ck 586 (A )3 uu); E. D « Schulz 5.n. [near ear Alpine, Aug. 4, 1928] (Wi), sen. Basie (Tj 92 8 QO. E. Sperry 75 (W-- 1681178), 1737 (Om), T 7.126 (Om), T.130 (Om 1679253), 1.573 (Fs, Om), 1.1073 (Ca—882852), T eine fe ee, 49k PHYTOLOGIA Vol. 11, née Steiger 1053 (N), 1133 (N); Tharp 880 (Au, Au), sen. begs Mts., 5-10-37] (Mi); B. H. Warnock 66 (W--1730933), > (N), 4000/ 7051 (Au—122863), 1700/5562 (Au--122862), 9953 (Rf), 20022 a Ok, S), €.802 (N), T.66, in part (Au), W.167 7 (N), W.283 (Au, N); M. S. Young s.n. (8/12/15) (Au), son. [9/12/15] (Au). Crane Co.: lundell & Lundell 10222 (Ld, N). ~Gulberson Co.: Hitchcock & Stan- ford ford 6782 (P1—116714, SAE Sie Se—69)17); Janszen 20 (Au-— 121659); Lehman s.n. [June 26, 1939] (Tr); Scholl 11 (Au--121658); 0. E. Sperry 1.129 eg aap re Waterfall 3775 (Tu—116233). El Paso Co.: B. Bar Barlow s s.n. [Canutillo, 12 July y 1911) (ca--882860, La, W—~—1430157); ; Benedict 16 (W—-1568212); M. K. Clemens s.ne gat Paso, Oct. 2h, 1916] 16] (Po—1737h9); Knobloch 1692 (i); L Lundell & Lundell 16941 (Rf); R. L. Oliver 95 (Nb); Shinners 899) . (Sm); Be Be H. Warnock 1650/7358 (Au——122853), | 7358 (Rf), f), 10422 (RE); Water- fall 3922 (St, Tu—128h43). Guadalupe Co.: Earle & pers 1,30 (N)5 K.S S. Young s.n. [9/9/16] (Au). Hudspeth Co.: Havard 97 (W— 1L759h); C. He H. Mueller 8215 (Sm); Thurber 143 (C); Be. He H. Warnock 1100/5912 ~ (Au-~122652), ~ 13813 (Au--171956, Tey Jeff Davis Co.: Bequaert s.n. (May 1, 1961] 61] (Au--208171) ; Correll & Johnston 1802 (Rf); Cory 17569 (Md), 27903 (N), 53075 (Sm), sen. [Sunny Glen Canyon, 6-29-27] (Tr); Karle & Tracy 162, in part (N); Fer- ris & Duncan 2647 (Ba, Du--126171, Gg--31h29, N, Po=-127357)3 Es Hamby 1112 (Ar--206808); Herb. Univ. Texas 71 71 (Po~-161328)3 Le Le Ce Hinckley ey 11h (N), son. [Mt. Livermore, 10-13-3h] (Au, Au), 5M {H. 0. Canyon, July 27, 1937] (N), sen. [8/2/37] (Fs); Lundell & Lundell 10255 (Ld, N), 13127 (Rf, Sm); R. McVaugh 7508 (Mi ee Sm); A. R. Moldenke 647 (S); Moore & Moore 2 (Ca—755008, N, EB. J. Palmer 32195 (Au); Parks & Cory 9599 (Tr), 9635 (tr), "17568 (Tr), 17653 (Tr), 18893 (Tr); H. R. Reed 101 (ar--307333, N)j Ee D. Schulz L057 (Wi); Tharp 51-22 Lngaaaee man & Janszen 9-10 (au—121573, Lw); Tracy & Earle 162, t (Au), 1628 (N); B. H. ee 5900/8681, Cu=I22581,), 13517 *(au--198386, 6, Bf), 21676 (N); S. E. Wolff 1666 (W—162395h); M. S. Young sen. [Davis Mts., 9/18/18] (Ur). 1 Montgomery Co.: Brenckle & Brenckle Ke 51103 Le). Pecos Co.: G. L. Fisher 36105 (W--1679361), sen. (July uly 20, 1936] (Se--69271); Goodman & Waterfall 1,565 (Ok); Lundell & Lun~ dell 10177 (Ld, N, W—1807631); B. H. Warnock 13956 (RE), C- 802 (au, N)j Whitehouse 19537 (wi, Nj; Ss E. Wolff 4390 (Tr). Presid- io Co.: Correll & Rollins 23676 (Ld); Cory 31173 3 (N); Drushel 11102 (N)j H. C. Hanson 551 (N, W--982093); L. C. Hinckley 1934 (N), Sen. an 9, 19h1] (Au); D.C. Ingram 2388 (Herb. Forest ae Ce es 96393] (s), ser t tHerb. F Forest Serv. Us Se De A- 394] (Au--186858, W--23k7L5), 2533 [Herb. Forest Serv. U. Se D- ri 96395] (Au--186861s) ; 3. Knight s.n. (Marfa, 9-29-32] (Av); 1965 Moldenke, Monograph of Verbena 95 Lundell & Lundell 1027 (N), 14339 (N, Rf); Tharp 3691 (Au); B. HE. Warnock 4300/5601 (Au--122851), 14594 (Rf); C. L. York 806) (Au—122008). Reeves Co.: Cory 52059 (N 9 (N); Nelson & Nelson 4963 (Au, Ka), 5014 (Au, Ca--710303), 5017 (W--197h035); Parks & Cory 18358 (Tr); B. H. Warnock 9945 (Rf); Waterfall 386 (N, Tu--31203). Val Verde Co.: * D. Schulz s.n. Seip (Wi). Ward Coe: Tracy & Earle 61 (Au, Cu, Es, N, N, Vt, W~h1790). mia unde- termined: Cottam 102h) [01a Fort a ee (Fs); Parry, Bige- low, Wright, & Schott s.n. [Frontera, Mar. 22, 1852] (N--topotype). NEW MEXICO: Bernalillo Co.: C.C. Ellis 17, i part (W--890528); M. E. Jones 457 (Po—-83728), son. [Albuquerque, Sept. 5, 188)] (Gg--154)21, “N, Tu), s Sen. (Po--65185); Kammerer 33 (N); D. F. Morgan s.n. [summer, 1939] (0k); Mulford s.n. d s.n. [near Albuquerque, Sept. '95] (Ur); H. H. Rusby s.n. - [Albuquerque, Sept. 5, 1909] (N). Catron Co.: 0. B. Metcalfe s.n. [Mangas Spgs., Aug. 1901] (W--562286); A. Re Moldenke 155 (Fg). Chaves Co.: G. J. Iken- berry 389 (St); Steiger 290 (N); Waterfall 10566 (St). Dona Ana Co.: Archer 327 (H--68552); Ellison 783 (La), s.n. [Organ Mts aie , 1929] (La); M. E. Jones s.n. (Rincon, 5-16-1890] {Poo 1163); Munz 13263 (Ca--537030, Po--20)037, Ua—-25323, Va--1)830) ; Ks C. Standley s.n. [Mesilla Valley, May 21, 1906] (W—560978), Sen. .. [Van Patt Pattens, June 11, 1906] (W-—560979) ; Tharp 46090 (Au, N, My, 8)3 Wooton 17h. (Du—-91155), 1900 (Au), s.n. [Organ an Mts., May 15, 92] (Ca--112317), sen. [Organ Mts., Apr. 30, 1893] (Hp, Ww alee. sen. (Sept. 15, 1893] (We), sn. [Organ Mts., Aug. 29, 1894] (C, N), sen. [Van Pattens, Aug. 29, 1894] (W—736872), s.n. [Organ Mts., May 8, 1895] (Po—70880) , sen. [Apr. 15, 1899] (Bl 42342), sen. [June 7, 1900] (Du--91154), s.n. [Organ Mts., 1900] (N, W~-560635), son. [May 26, 190] (Fs), sn. [Filmore Canyon, May 25, 1905] (W--736871), sen. [June 10, 1906] (Ob--50878). Ed- Co.: V. sen. {Carlsbad Cave, Apr. 15, 192k] (W-~122014)); D. W. Lee 3 3 (W--1203610) ; A. Nelson 11401 (Ca~-500719, S); Water- fall 3720 (Tu--115260), 1061) (St); Wilkins 1736 (En). Grant Co.: Bigelow sen. [Copper Mines, July 28th 1651] (C); Blumer Ly (= 563467); Eas Eastwood 81,68 (Gg--3127, W—1102096); Eggleston 17231 m 17232 (N); Emery s.n. [Oct. 17th 18h6] (C); G. L. Fisher 36132 — 1679375); E. L. Greene s.n, [near Silver City, June 1880] (Po— 71129), s.n. [26 July 1550) (Vt), son. [Aug. 3, 1880] (Pa); Hol- zinger s.n. (Fierro, July 29, 1911] (W-—-660556); Layton s.n. {Santa Rita, Sept. 1931] (Io--138119); Mearns 108 (Du--2h198); Oo. B. Wetcalfe 126, in part (Po-—70888); Mulford rd 681 (N); W. Stew- art s S.n. [Mimbre i a. June 26, '36] (La); C. B. Wolf 2598 (1 (Ba, Rs—-9997). Guadalupe Co.: F. C. Gates 20979 9 (Ka--922651); Har wood s.n. [Vaugn, 9~10-1921] (Po--18602); A. A. R. Moldenke 126 ( (Fg); 496 PHYTOLMOGTIA Vol. 11, no. 7 A. Nelson 11338 (Du—218830, N, ns 11388 (Ca--500720, S)j Be Le Wagenknecht 475 (Lw). Lincoln Co.: “De Bus Busk s.n. (U.S. Dept Agr. Soil Conserv. Serv. 6005] reason gon. [U. Ss Dept? Agr. Soil Conserv. Serv. 6006] (Tu-~10)396); Earle & Earle 236 (N, W--382398); L. C. Hinckley 750 (N, N), 776 (N)3 Hitchcock, Rethke, & Raadshooven 26ka (CG: (Ca--603732, , Gg=-267613); A. Lee 9 (Au, N); | F. G. Plummer s.n. [Lincoln National Forest, 1903] (We- 563048); J. Fiestian 5 han 5 (W--35012h); Wooton 364, in part (Ca—138816, Du--75577, Ka, Po—7112h, Ur); Wooton & Standley 3635 (Du--9536). Luna Co.: Abrams 7259 (Du--98h13), 3 SMe TMay 1919] J (Du--98125) ; M. E. Jones 3 3001, in. in part (Gg--13052h) ; Mearns 109 (Du--2)191, W— 56226), s.n. [April 17, 1892] (Du--24195). McKinley Co.: East- wood 15697 (Gg—161738). Otero Co.: F. S. Earle 539 (N); Goodman & & Waterfall 1995 (Ok); G. J. Ikenberry y 256 (Ok, St); Layton s.n. {Alamogordo, Sept. 1931] } (Io—1387h3); Orcutt sen. [Cloudcroft, July 23—26, 1926] ager Rehn & Viereck s.n. [Alamogordo, Apr. 16, 1902] (TI, , Mg--35, Po—18610); E. D. Schulz 549 (Wi); P. Co Standley ee a temo Waterfall 10603a (St); Wooton SN. . [Divide a above Mescalero Agency, June 23, 1895] (W-~736223) » s.n. [Along Tularosa Creek, Aug. 18, 1899] (W--736870), s-ne (White Sands, Aug. 25, 1899] (W—-736883) « Quay Co.: G. L. Fisher sen. (Nara Visa, Apr. 21, 1911] (Gg); G. J. Ikenberry s sen. [May 1, 1937] (St). sandoval Co.: Arsane 20890 (Du-=253899) ; Hedgecock 218 (Se--95967). San Miguel Co.: G. G. Je Goodman 2308 (in——26305, N); G. Be Grant 5758 (Du--75652); Mulford 37 (Io—3u0h7, Ny ye A. Nelson 11566 (Ca--500723, Du—210532, S); Sturgis sen. [May 1902] (Ca--216687). Santa Fé Co.: Arséne & Benedict 16555 (13 Fendler 586 (Dt); Heller & Heller 3536 ee Ka, N, Ob-- 50861, Pl—72006, Po—-63892, Ur, W--306h23); D. F. Morgan $.Ne (summer, 1939] (0k); A. Neison 11757 (ca--500722, pu-—210831) 5 W. W. Robbins 8242 (Bl--[2298); Snow s.n. [Santa Fé, Aug. '80] (H--107719), s sen. [Santa Fé, Aug.] (Ca—-67417). Sierra Co.: Cut- ler 2065 (0k); 0. 0. B. Metcalfe 1090, in part (Gg--31h28, N, Po $4188). Socorro Co-: "S. B. Benson 323 (Ca—526578); D. B. Dunn —_——— = (Lb--,8h9); C. L. Herrick 70h (W-~-73723); A. R. Moldenke x (5); H. H. Rusby 337, in part (Pr); Wooton SMe [Devil's Park, is. 9, 1900} (W--736934) ; Worth & Priest 5,0b (Ba, N). Torrance Co.: F. Clark 460 (St—-9203). Union Co.: Wooton s.n, [20 miles west of Patterson, June 21, 1892] (We=-562252). Valencia Co.: 0. Degener 1,853 (us) ; Wooton s.n. [west of Grant's Station, Aug. 2, 1892] (W-~562253), Some [south of San Rafael, July 29, 1906] (W— oo County undetermined: De Busk sn. [Bottomless Lake a, - S. Dept. Agr. Soil Conserv. Serv. 6007] (Tu—-104395); ae Ehlers 6304 [Wilma] (uh, Mi) ; Mexican eee) Surv. sn. (N N)3 vasey 1965 Moldenke, Monograph of Verbena 97 Sen. (Ka); Wencurtin s.n. [5-2-1916] (Au). ARIZONA: Apache Co.: LD D. Benson 9555 (Du-~326793, Po--267659, Tu--7643); Deaver 178) (Fg); Griffiths 5190 (W—-L9658h); A. R. Moldenke 153 (B, Fe, Fe, S)3_ R. L. Liver 91 (Nb). Cochise Co.: Cottam 10201 (Fs); J. J Thornber (Tu). Coconino Co.: H. H. Rusby 337, in part “(ons 67972). — Cos: Eggleston 19912 (W—152497h); Maguire & Ma- guire 10117 (Ca--676071, N, Pl--130552, Ua--l710), 10179 (Gg-- 295127, N, N, Ua—l7h08); Spangehl s.n. (Thatcher, May fay 1936) (Tu); Thornber & shinies 7899 (ok). Gree Greenlee Co.: Maguire, Richards, & Moeller 10095 (N, N, Ua--709), ic1e (N, N, |; Ua--h7hil)» Mol iichave Cow: Ke: Kearney & Peebles 12669 (To). vajo Co.: A. Nelson 10358 (Bt—196l6); Peebles 961) (3621359), Peebles & Fulton 961) (To); L. F. Ward s.n. [Woodruff, ea 0, 1901] (N). Pima Co.: Spaulding 3 37 (Tu). Pinal Co.: Ge J. Harrison 1960 (To); J. J. Thornber s.n. pela Suess ia 21--23/05] (Tu). MEXICO: Coahuila: G. | G. L. Fisher s.n. (Aug. 17, 1930] (I); Gregg sen. [Sal- tillo, 6/67) (C); E. G. Marsh 707 (St), 152) (St); Re: R. L. McGregor 16712" (Lw); R. McVaugh 12300 (Mi); Stanford, Retherford, & North- craft 2h1 (Ca--71361, Se--70282); Waterfall 15348 (St), 15782 (St); Waterfall & Wallis 13273 (St); Wynd & Mueller 572 ‘sp Nue- vo Leén: Painte: Painter, Lucas, & Barkley 1281 (Au). San Luis Potosf: C. L. Lundell 5590 (Du--266212). SOUTHERN RHODESIA: Byles 439 (Rh). CULTIVATED: New York: T. H. Everett s.n. [H. N. |, Moldenke 10080] (N); Quinn s.n. [N. Y. “Bot. Gard. Cult. Pl. SD.228-37] (N). VERBENA easy f. ALBIFLORA Moldenke, Résumé Suppl. 6: 3, nom. nud. (1963), f. nov Haec oS a forma typica speciei corollis albis recedit. Biblio ene Moldenke, Résumé Suppl. 6: 3. 1963; Moldenke, Phytologia 1i : 48. 196. This form differs from the typical form of the species in hav- ing white corollas. The type of the form was collected by Barton Holland Warnock (no. Pld) on the campus of Sul Ross College at Alpine, Brewster County y, Texan, on April 2, 1936, and is deposited in the Britton rataed Rages t+ the New York * Botanical Garden. The stems are branched at the base, the branches erect or x ae ding, to 20 cm. or more in length. The plant is described = as "infrequent on graded road 12 1/2 miles northeast of Fort Stockton" in Pecos County. Warnock calls it "infrequent on Jim Nichol's Ranch, Glass Mountains" in Brewster County. The Moores found it on sandy flats at 2200 feet altitude in Reeves County. Material has been misidentified and distributed in herbaria V. racemosa Eggert and V. wrightii A. Gray. It has been aatated in flower and fruit in March and April. In all, 8 herbarium specimens, including the type, have been ex- dq mé. 498 PETTOLOGIA Vol. 11, no. 7 Citations: TEXAS: Brewster Co.: B. H. Warnock Tl (N-type, W--1730928--isotype), T.l5 (N, N, W—-1730929). Pecos Co.: Cory 53502 (N). Reeves Co.: Moore & Moore 4 (Ca-~755057, N). VERBENA XUTHA Lehm., Del. Sem. Hort. Hamb. 1834: 7—8. 183k. Synonymy: Verbena strigosa Hook., Comp. Bot. Mag. 1: 176. 1836 [not V. ae osa Cham., 1832]. Verbena lucaeana Walp., Repert. Bot. Syst. 4: 23. 1845. Verbena matthesii Turcz., Bull. Soc. Nat. Mosc. 36 (2): sat 1863. Verbena xanthi Gray ex loldenke, Suppl. List Invalid Names 10, in syn. syn. 191. Verbena xutha Schm. ex Molden- ke, Suppl. List Invalid Names 10, in syn. Syn. 191. Verbena zutha Lehm. ex Moldenke, Suppl. List Invalid Names 10, in syn. 19 Verbena xantha Lefm, ex Moldenke, Résumé 378, in syn. 1959. ver- bena xanthii Lehm. ex Moldenke, Résumé 378, in syn. 1959. Verbena xertha Lehm. ex Moldenke, Résumé Suppl. 6: 11, in syn. 1963. Ver- bena x xuthia Lehm. ex Moldenke, Résumé Suppl. bi: 11, in syn. 1963. Verbina mutha Lehm. ex Moldenke, Résumé Suppl. 6: n, in syn. 1963. Bibliography: Lehm., Del. Sem. Hort. Hamb. 183k: 7--8. 183k; Linnaea 10: Litt.-Ber. 115. 1836; Hook., Comp. Bot. Mag. 1: 176. 1836; Walp., Repert. Bot. syst. he 23. 1815 Steud., Hom. Bote, ed. Mosc. 3% (2): 196. 1863; A. Gray, Syn. Fl. N. Am. 2 Wats., Proc. Am. Acad. Sci. 18: 135 & 136+. 1883; Cowlt., Con- trib. U.S. Nat. Herb. 2: 327. 1892; Briq. in Engl» & Prantl, Nat. Pflanzenfam. (3a): 148. 189); Jacks. in Hook. f. & Jacks., = : Small, Fl. Southeast. U. S., ed. 2, 1009. 1913; Pallett, Am. oney ed. 2, att 1923; Seymour, Host Ind. Fungi N. Am. 587— 88. 1 ee , Man. Southeast Fl. 1137. 19333 Perry, Ann Mo. Bot. Gard. 20: = ald, Rig along os 355, & 356. 1933; Cory, Texas Agr. ° 50: ~ ana ; Moldenke, Pre Names 9 & 10. 191; Moldenke in Lundell, Fl. Texas 3 (1): 26--27 & h6—l7. 19h2; uoldenke, Alph. List Invalid Names 48, 50, & 51. 1942; Moldenke, eogr. Distrib. Verbenac.., [ed hig as a0, » Cas : 40. 1945s Moldenke, Phytologia 2: 87 & 117 (ais). earner 1916: G. L. Fisher, Am, Bot. Exchange List. 19h63 Pl., ed. 420. 1947; Moldenke , Phytologia be 328, 329, & 18. 197} abe Wrightia Ls 225 & 226. 1948; Moldenke, Alph. Liat: Cit< 28:39 ’ 399, 405, 416, 439, “ise, u71, 473, W7h, 78, 7189s 98 506, 520, 522, 558, 598, 618, 627, & 632 (1948), 3: 657, 665, 671, 678, 679, 683-685, 696, 708; 713, 743, 750, 752, We. "163°? 440, 781, 788, 1965 Moldenke, Monograph of Verbena 99 795, 796, 798, 806, 84), 851, 855, 882, 91h, et mats 2b of (1919), and li: 986, 989, 990-993, 1003, 1069, 1 1113, 1122, ae 122, 11 eer ce 1162, 18s *a66, “BB, Hort. Soc. Dict. Gard. 6: 2209 & 243. 1951; Moldenke, In form. Mold. Set 8 Spec. [h]. 1954; Shinners, Sp - Dallas 326, ig 8; Moldenke, Rés é 1h, 15, 205 4O, 225, 368, 0 0. "1960; Lewis H Oliver, An. soteise Boe 48: 639-641, fig. 2h. 1961; Jones, Rowell, & Johnston, seg Pl. & Ferns Tex. Coast. Bend 1b. 1961; Moldenke, Phytologia 8: 12 & 435. 1961; Moldenke, Résumé er ee ar 6 & 8 (1962), h: rh & 20 (1962), 5: 3& h (1962), 6 11 (1963), and 7: 2. 1963; Voldenke, Phytologia 8: h7l S. » 478, & i930 (1963), 9: %, 81, 160 & 215 (1963), 10s ‘Bodh, 104, 108, 110, 140, 178, 182, 186, “2, 217, 218, 30h, & 502 (196k), and 11: 13, 110, at 127, 128, & 201 196k; ictheke, Résumé Suppl. 8: 1 & 3 (196i) and 11: 3 96h: Moldenke, Phytologia ll: _* 6:2: Illustrations: Lewis & Oliver, Am. Journ. Bot. 48: 640, fig. 1961. Usually a tall, erect, annual or perennial herb, to 2m. in height, coarse and spindly, sometimes bushy or even fil in outline, 5--12 cm. long, the upper ones Lnhgedepiemah Tia. “the a 3 violet, purplish-violet, bluish-violet, blue~lavender, bluis lavender, lavender, or pinkish-white, hirt rtellous, its ’ tube approx-~ imately as long as the calyx, the throat white, pubescent outside, villous within, the limb medium in size, 5 mm . wide; cocci a~ bout 2 mm. long, raised-reticulate on the upper half, faintly Striate below, the commissural faces reaching the apex of the coccus, muricate or muricately scabrous; chromosome number: n = 21. 500 PHYTOLOGIA Vol. 11, no. 7 The type of V. matthesii was collected by Benno Matthes (no. 13) ~~ in whose honor it was named -- in Fayette County, Texas. Hooker's V. strigosa was based on two collections of Thomas Drum- mond -- no. 253 bis from New Orleans, Orleans Parish, Louisiana, in 1832, “and s.n. from New Orleans in 1833 -- deposited in the herbarium of the Royal Botanic Gardens at Kew. Verbena lucaeana is merely a new name for V. strigosa, although Tainturier s.n., from New Orleans, is the only specime en cited. Hooker's description of his V. strigosa is as follows: "Ver- mens, both hg Mr. turier, so that it must a very common plant; yet it does not appear have been taken up by any author, nor have cimens been transmitted to affinity is, perhaps, with V. stricta, but the leaves an strong- ly pinnatifid, harsh, with the princi: principal veins more prominent on the under-side, yet not exhibiting such a strongly-retic culated Fa pe the spikes are longer, more lax; the flowers much er, more distant, and not pressed to the rachis. It is 4 strong- owing plant, two to three feet high, quite hispid with appressed hairs or bristles." Walpers (185) places this species in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Sec= ary Subgroup Schizophyliae With 9 other species. The specific name is often uppercased for no valid reason. Verbena xutha inhabits sand, red or brown gravelly silty sand, open dry soil, low sandy or sandy loam soil, most streamsides, low roadsides, and the edges oi ne fields, and roadsides. It has been found in waste ground, open or ped rocky ground, and sandy n Te tly and roadsides and in blackland prairies, in the eastern and so southern west to Jeff Davis and Brewster Counties, flowering from March t October. Bush reports it as "common on prairie" in Brazoria Sat 1965 Moldenke, Monograph of Verbena 501 The Lundells found it along "roadside through pineland" - Hardin Chambe County, Traverse in "Panicum amarulum complex" in and "in ditch in mectrond egies right-of-way, on sand with some ne assoc- iated with gras and various herbs" in Harris County; L. D. Smith "between field and wood border of stream" in McLennan, F. W. Gould in "partial sami along road through po forest" in Liber- ty, Garberson "in sand-loam soil alo in Brazos, M. C. Johnston "in ai othe “en of swale in prairie" yn Aransas County. a is, "occasional in dense herbaceous vegetation in sand along the road" in Jefferson, "occasional in sandy fields" in Galveston, and ‘in- frequent in open woods along creek" in Anderson County, while C. sition black soil to sand, in red clay and sandy loam, prefers presence of aed in Montgomery Coun » Turn & Johnston found it ‘ithal abiting a "gumbo poo (a local prairie ie ‘ei cht marly clay wre surrounded by pines on sand)" in Polk County, McDougall in "old field weed stage" the side" in Hardin County. Jones, Rowell, & Johnston (1961) state that it inhabits tgwales, ra avines, ditches, etc.", flowering in summer and fall. In Louisiana the species was found by Shinners "in yellow-gray Clay on highway rights-of-way". It has been in cultivation in Europe since 18:0. Common names reported for it are "Gulf ver- ", "hoary vervain", and "vervain". C. L. Lundell 50)7 is a low many-lobed form. Demaree 32009 in the Leiden herbarium shows a fasciated a The flower color o fv. xutha is described as follows by vari- ous collectors: "lavender" — Ferris & Duncan 3268, C. L. Lundell 11921 & 14135, and Lundell & Lundell 11522 & 13107; "blue- lavender" -- C. L. Lundell 1062; "lavender-blue" -- Shinners 23982; "pale lavender-blue" -- Traverse 770; “bluish-lavender" —- C. L. Lundell 15082; "bluish~violet" -- F. B. Jones 2695; "gray- Lilac" —— Kuntze 23611; "violet" — Brenning s.n. (17.6-190k] & C. M. Rogers 65%; "purple" ~- Tharp, Turner, & Johnston 51,706; "pinkish-white with white throat” -- Traverse 2510. This plant is often attacked by the fungi Cercospora papillosa Atk. and C. verbenicola Ellis & Ev. according to Seymour (1929). Material has been misidentified and distributed in herbaria under the names Verbena carolinense (Walt.) Gmel., V. carolinensis (Walt.) J. F. Gmel,, V.c V. caroliniana L., V. halei L., ve halei Small, V. hastata L., vel littoralis H.B.K., V. neomexicana (A. Gray) Small, i officinalis L., V. polystachya H.B.Ke, V. scabra Vahl, V. stric- ta L., V. stricta Vent., Phyla incisa Small, Stachytarpheta sp., and even Lythrum californicum T. & G. 502 PEYTOLOGIA Vol. 11, no. 7 On the other hand, the F. B. Jones 192, distributed as Verbena xutha, is actually V. brasiliensis Vell.; Edw, Palmer 10,7 is V. canescens H.B.K.; Lindheimer III.500, McKelvey 1710, and E. De Schulz 526, as well as E. J. Palmer Palmer 9183, are V. canescens 8 var. roemeriana (Scheele) Perry; 3 OM. E. Jones 28296 is either V. canes- Jones 29188, Molby 7221, “and Williges Se are V. cloverae Moldenke; A. M. Hildebrandt s.n., E. J. Palmer 911] & 9405, and E. D. Schulz See ee a enangemamnanenee iene ee eee — — — folia Santis: E. L. "ales Sen. [Prinos Ties Mts., Aug. 2. 1880], S. S. Marsh 3099, C. H. Mueller 8138, and Tharp 493 are V. neo- reed Sceieneallieeeadeememeeeeeee te cen tte a erm ; ry, Bigelow, Wright, & sence s s.n. [Valley of the Rio Grande below Dofiana], Shreve 9089, “Tharp 3687, 87h, s.n. [Devine-Dilley, March i. 1930], Sn. [Wilson Hane Ranch, ~ 6/19 31], & sen. [6-19-31], and M. S- Young 101 are V. neomexicana var. hivtelta Perry; Goodding 22h6, Herb. Le LeRoy | sen. Ariz. '8h], LeSueur 875, Pringle s.n. [Santa Rita Mts., May 11, 188k], Bs Runyon 55, Schery es and Wynd & Mueller 525 are V. neomexi War Syled erry; C. T. Mohr s- n. [ballast ground , Hobie, 5/30/188 7] . — officinalis Le3 E-Ce Marsh 1378 is V. perenni cre McKelvey 1756 is V. plicata Greene; and H. C. Hanson 96, F. B. Jones "oe, Killip 2107, Re Runyon 522, and Tharp 1201 oe oni Moldenke, while the E. W. Nelson on 6096, distributed as "Aff. V. halei or V. xutha" is V. pine ee Moldenke. - Ce Albers 33017, R. Runyon 1777, and L. F. Ward sen. [Hearne, ae n, 1877] are mixtures with ve “VY. halei Small; “Bogusch 1235 1235 is a mischaee with V. scabra Vahl; and T T. Drummond sn. {New Orleans 1832] is a mixture with Stylodon carneus (Medic.) Moldenke. Watson (1883) maintains that Edw, Palmer 2037 is "probably a hybrid between V. polystachya or V. urticaefolia and V. xutha", but I regar d it as aries typical Vv. ehrenbergiana Schau. H. Ness, in a letter to Dr. J. K. Small dated pages’ 2 2h, 1899, says "T aeal include in this letter a specimen of Verbina, to which not be cies before. It he app cee ide and is often found in com~ pany with V. zutha a is much lar fe 2 5 feet high, leaves broader, and the lower much less ced tha V. zutha. It is also more hairy, and bears larger flowers. If this species is a strang- er to you I will send you better specimens. I would like to have ie opinion on this plant.....[It] is a perennial, with pestis parcely branched stem." Dr. Small identified the plant as V. x~ 1965 Moldenke, Monograph of Verbena 503 tha, and with this identification I fully agree. Pellett (1947) reports that V. xutha yields nectar sparingly for honey in Texas. The Coulter reference given in the biblio- graphy above is ph ap cited with the date "189", which is incorrect for the page in question. The Appleman s.n. {Hot Springs] cited soe oa not actually have "Texas" on its label, but I am assuming that it was from that state, rather than from any of the localities with similar names in Arkan ansas, Montana, North Carolina, South Dakota, or Virginia. Schauer (1817) cites Tainturier s.n. fron Louisiana and Lind- heimer sn. from near Houston, Texas, i in the DeCandolle Herbarium at Geneva and cultivated material in the Botanical Garden at Ber- jin. Ras comments "Hab. in America tropica. Conf. V. Ehrenbergia- nam. feel (1933) cites the following 35 additional specimens not as yet seen by me: ARKANSAS: Miller Co.: Eggert s.n. [Homan, 10 June 1898] (E); Letterman s.n. [Texarkana, Aug. 1881) (E). " LouIs- IANA: Cameron Par.: S. M. Tracy 8708 (E, F, G). Natchitoches Par.: E. J. Palmer 7556 (1 (E). Orleans Par.: T. Drummond 253 bis (K). Rapides Par.: “Sal R. Ball 605 (E). Saint Martin Par.: Langlois s.n. [St. Martinsville, x 30.V.1893] (E). Parish undeter- mined: T, Drummond s.n. (G); J. Hale s.n. (E, G). TEXAS: Austin Co.: Lindheimer 15 (E). Brazoria Co.: B. F. Bush 899 (E), 1275 (E). Brazos Co.: H. Ness s.n. [sandy land, “Brazos Co., 1899] (G); J. Reverchon son. (College Station, oh July 1899] (E). Gregg Co.: Eggert s.n. (near Longview, 7 June 1899] (E); J. Reverchon 2532 (E). Guadalupe Co.: Groth 187 (F, G). Harris a J. Ball Sen. [Cypress City, May 1877] (E); | Ferris & Duncan 3 8 (E); Lindheimer pon (E, G). Jackson Co.: J. A. , Drushel = (E). Jef- ferson Co.: Tharp 3166 (E). Montgomery Co.: Dixon 473 (F, G). avarro Co.: J. Reverchon 2118 (E). Palo Pinto Co.: /: Lindheimer S.n. [Brazos, “July 1899] (E). Travis Co.: E. D. Schulz 701 (W). Washington Co.: E. Hall 43h (E, F, G). The "Munz 1607" cited by her is undoubtedly the same collection cited by me as Munz 1601. b nc sete trifid alee, : It has often been confused with V. canescens var. roemeriana, but the latter is a smaller and more compact plant," In all, 345 herbarium specimens and 2 mounted photographs, in- a vy type or Lan ghcay aty material of most of the names involved, have been examined by Citations: ALABAMA: "Mobile Co.: C. T. Mohr s.n, [Navy Cove, Aug. 1889] (W-—771855). Montgomery Co.: Koepper, Justice, & Isely Sen. [Oct. 3, 1943] (Dp--38319). MISSISSIPPI: Harrison Co.: Dem- aree 32009 (Le); Small & Alexander s.n. (near Pass Christian, May 50h PHYTOLOGIA Vol. 11, no. 7 1931] (N). Hinds Co.: W. H. Rhoades s.n. [Jackson, July 1927] (Bt, N), sen. [Jackson, 1927] (Bt--5921). Jefferson Co.: McDou- gall 1033 (W--1925220). Warren Co.: Moldenke & Woods 523 (B, Fe, S)3 1 ": 5. H. Rhoades s.n. (Vicksburg, July 1927; H. N. Moldenke 17088] (Br, Bt, Bt--301h], Hs, N, St), sen. (Vicksburg, July 1931] (N), sn. rhakelbevenndl Bluff, 7-1931] (Bt-—-61221). ARKANSAS Ashley Co.: Demaree 2518 (Lb--6760, Sm). Chicot Co.: Demaree 23265 (Sm). Cleveland Co.: Demaree 23313 (Z). Hempstead Co.: Demaree 4667 (Ca). Miller Co.: Eggert s s.n. [along railroads, 10 June 1898] (N); Letterman s.n,. [Texarkana s.n., 1881] (W--719 77). LOUISIANA: Ascension Par.: T. L. Andrews s.n. fAscensian, 13-1] (Io—83618). Avoyelles Par.: e R. Moldenke 72h (S); Penfound s. n. (May 16, 1946] (Tl). Beauregard Par.: A. R. Moldenke 707 (S). Bossier Par.: A. R. Moldenke 678 (S). Cameron Par.: S. HM. [. Tracy 8708 (Es, Lb—27628, N, Tr, Up--50343, W—510600). East Baton Rouge Par.: Joor sen. guly 6, 1874] (Mi). East Feliciana Par.: M. Carpenter s.n. [Feliciana, May 180] (W--771857). Grant Par.: I. L. Forbes s.n. [Colfax, Aug. 20, 1927] (Ca--35157h, Io—129293). Jefferson Par.: L. H. Bailey 7339 (Ba); L. E. Fox 2107 (Fx). La- fourche Par.: Arceneaux 93 (It). La Salle Par.: Shinners = (Hi--1976)5, Tl). Natchitoches Par.: E. J. Palmer 7 7556 (Gg 182147, W--1531646). Orleans Par.: Brenning s.n. ie (hE 5 agob] (B)5 Cocks s.n. [New Orleans] (Tl); T. Drummond s s.n. [New Orleans, 1832] (Lu, S), son. [New Orleans, 1833] (Lu), s.n. [env. N. 0-] (Vi); Holzer s.n. [New Orleans] (Mi) 3 J. M. Mearthur s.n. [7/21/ 28] (Tl). Plaquemines Par.: Langlois 123 ) (W--1323140), 612 (Cm), sen. [July 1878] (Du--91128), s.n. [25 June 1884] (Al). “Rapides Par.: C. R. Ball 605 (W--3828kh). Red River Par.: A. R. Moldenke 689 (S). Saint Charles Par.: Ewan 1770, (Tl). Saint James Par.: Small & Alexander s.n. [near Convent, Apr. 1931] (N). Saint Mar- tin Par.: Correll & Correll 9447 (H--78927, N, No—18)08); Lang~ lois s.n. [St. Martinville, 30.V.1893] (Ba, I, I, N, W——1,65963) « Terrebonne Par.: Bynum , Ingram, & Jaynes s.n. [3 mi. w. of Cocod= rie] (N); Munz 1601 (Po--12h052). Parish undetermined: Dormon S- n. [1930] (N); Herb. Chapman s.n. [Louisiana] (C). TEXAS: S$: Ander- son Co.: Cory 54630 (No--15143, Rf). Aransas Co.: Cory 45878 (N), 45879 (Au, Sm), 51177 (Sm); M. C. Johnston 53253.1 (Au--122882), 53253.2 (Au--122883), 532533 4 (St); F. Be. F. B. Jones 3195 (Ww); U Uzzell 17 (Ar--123390). Austin Co.: F. We Pennell 10279 (N); He wurzlow son, — 1891] (Ur). Bastrop Co.: B. H. Warnock T6439 (AY, N)- Bell Co.: S. E. Wolff 829 (W~1622068), 1016 (W--1622581), 436 (Tr). ‘suas Co.: Let Letterman 394 (Du--91133), 5477 (N), SMe [Texarkana] (W--98668),). Brazor-: Brazoria C0.: B. F. Bush 899 (We); Kil- 1965 Moldenke, Monograph of Verbena 505 lip 40591 (W--2005590). Brazos Co.: Garberson 293 (St); W. S. Jennings | 155 (0k); H. Ness s.n. [Aug. 1; Reeves - 202] (Cs), : S.n. [vuly 16 16; “Reeves 203] (Cs), 8 s.n. [Aug. 16; Reeves 205] (Cs), SM. [July 1899] (N), s.n. [June 2%, 1925] (tr); H. B. Parks s.n. [6-_ 7-47] (Au), s.n. [White Creek, "9-7-7 ] (S); Reeves 191 (N), ant’ (Cs). Brewster Co.: Applenan 8 sen. [Hot Springs] (La). Burleso Co.: Trew 10) (Au—122827). Calhoun Co.: Harold Gentry 57 ms N, N). Cameron Co.: Clover 1695 (N); Mrs. Cottrell 8743 (Au) 3 Ecology Class Univ. Texas s S.n. [Palm Grove, 3.1.30] (Ay); R. Run- yon 1777, in part (Rr). Chambers Co.: Cory 50987 (Sm); Tharp s. n. [Anahuac, July 22, 1929] (Au, Au); Traverse 2510 (Au—208),79) « eae Co.: C. Me Siaers 6536 (Au--122826). Dewitt Co.: M. Riedel sen. [7-h-l1] (Au); E. D. Schulz s.n. [5/11/1938] (Wi); Webster & Rowell 7065 (Au, N). Ellis Co.: Cory 53386 (N). Fayette Co.: Matthes 13 [Macbride photos 34343] (Kr—photo, N--photo); B. H. Warnock rnock 146297 (Au, N, S). Fort Bend Co.: Munz 1470 (Po— 121658). Freestone Co.: Harding 399 (St--17058). Galveston Co.: Cory 51020 (N, Sm, Sm, W—1886098); Dapprich 7764, in part (Sm); G. L. Fisher 212 (W--107h6L0), s Sen. [San Leon, July 7, 1929] biog 33687, Gg——290860); Mrs. A. F. Nelson Sone [11~2-11) (Au); P & Sony 20201 (Tr). Goliad Co.: C. B. Williams 18 (au). b ie Co.: C. C. Albers 16092 (Au , N); B Bogusch 970 (Au), 1235, in part (Po—161325), Tet [Her [Herb. Univ. Texas 970] (W—~13285)0); Flemming s.n. [Rutledge Swamp, Oct. 9, 193k] (Au, Au); Parks & Cory y 10068 (Tr); Tharp 253 (Au). Gregg Co.: J. Reverchon 2532 (N). Grimes Co.: Eee 2 s.n. [Plantersville, "10728 31] (Au); J. N. Weaver 1039 (N); T. Vv. Weaver 1039 (M1). Guadalupe Co.: Groth 187 (Dt, | (Dt, Gg— 7432, Io—75837, N » W—-717776). Hardin Co.: C. L. Lundell 1,082 (N, RE); Lundell & Lundell 11522 (N, Rf). Harris Co.: Armer s.n. [Houston, “July 1927] (Au); L. %. fe Bailey 310 (Ba); Boll s.n. [Cy- press City, May 1877] (E--117233); Boon él (Au), 291 (Au, N), 20001 (Au, N); Condit s.n. [Houston, June e 19, 1909] (Ca——l561L9); O. Degener 5171 (Ms, N); Eifrig s.n. [Houston, June 1926] (Ca— 882857, Mn——35073); Ferris & Duncan can 3268 (Du--12519, Ge—31433, N)3 G. L. Fisher 82 (W--503153), 51048 (Ew, Go), smn. [Houston, June 9, [912] (Vi), sen. [Houston, May 18, 191] (Hp), sen. [Hous- ton, May 16, 193k] (Gg--237866); Fitzgerald 296 (B1l--42367); Joor sen. [July 16, 1875] (Mi); Killip 15699 (W—2234120); Kuntze 23811 (N); W. H. Lewis 5300 (Nb); Lindheimer 1.154 (Ca--2h8)31, W- 1870923), s.n. [Houston, 1643] (Pr, W—18700h6); Lundell & Lundell 13107 (N, Rf) C. T. Mohr s.n. [Harrisburg] (W--771956); 0. Muel~ ler 10-14 (Wi); E E. 2% Schulz 37-93 (Wi); Tharp 498 (au); Traverse 770 (Au--168129, Rf). Hays Co.: S. W. Stanfield s.n. {San Marcos and vicinity, summer 1898] (N). Jackson Co.: J. A. Drushel 2842 506 PHYTOLOGIA Vol. LL, nos 7 (Ur); A. R. Moldenke 207 (Fg); Tharp sen. [8/30/41] (Au), sen. [Sept. at, , 1941] (Au). Jeff Davis Co.: 5 E. J. Palmer 34370 . (N) Jefferson Co.: C. C. Albers 34005 (Au, Au); Cory 50962 (Sm), 50968 (N, Sm, Sn, W~-1886083) ; 3 Golub s.n. [Beaumont, July 8, 1938] (is); Kolthoff s.n. [lh Maj 1927] (S), sen. [10/5/1927] (S); We He Long $.n. (Beaumont, May 28, '01] (Au); C C. L. Lundell 1135 (N, Rf, W— 1927002); Parks & Cory 19878 (Tr); Tharp 3166 (Au), 3167 (Au), S- n. (McFadden Beach, 9/10/37] (Ca-~882855), sen. (McFadden Beach, 9/11/37] (Au, Au, N, P1--151555, w--187378h), sen. [9/12/37] (Au, Au); M. S. Young s.n. [Port Arthur, )/21/18] (Au). Kaufman Co.: Ee Hamby 1285 (Ar--206803). Lavaca Co.: G. L. Fisher 122 (W— 503192). Lee Co.: J. F. Brenckle 18076 (N), 49075 atic (N)- Leon Co.: E. C. Smith sen. [Oakwood, 5-12-1942] (Fe). Liberty Cos F. W. Gould 6922 (Au—122868); Harding 172 (St--17565); C. L. Lundell 15082 (Rf, W—2297897). McLennan Co.: Le D. Smith 649 (Au--122873, St). Milam Co.: S. E. Wolff 4023 (Tr). “Montgomery Co.: Dixon 73 (N); Warner s.n. [Willis] (Hu). Navarro Co.: J. Reverchon 2118 (W--387015); Lloyd Williams 7 (Au). Polk Co.: , Turner, & Johnston 54706 (Au--12290h, , St). Refugio Co.: i Re ‘Woidenke 199 (Fg, S). Robertson Co.: L. Morris 2 (Au); Fe J. Tyler s.n. [Calvert, May 13, 1904] (W--50080)) ; Le ¥. Ward SMe [Hearne, Sept. 11, 1877] (W-~1;7568). San Patricio Co.: F. F. Be Jones 056 (Ww). Travis Co.: F. A. Barkley 13365 (Au, N)} Cohn & Barkley 13253 (N); L. C. Hinckley 1,03 (N); Ce L. Lundell 11922 (N, Rf, Rf, RE); R. H. Painter 16 (Ka—717h:2) 5 he D. Schulz z 619 (W-~110L9L5, Wi, WH), 700 CH); Strandtmann s.n, [July 2%, 1910] (Au); Tharp 667 (Au, Au, W—110\0)1, Wi), 668 (Au, W--110L6L2, Wi), sn. (Walnut Cr Cra, 7/12/20] (Io—104882), s Sen. { Walnut Creek, spring 1921] (Io—104881), s.n. Tle] (Au); York, Copeland, & Johnson 6048 (Au). Tyler Co.: Tharp, Turner, & Johnston on 54838 (Au--122903, St). Victoria Co.: F. B. Jones ee (tw); F. Le Lewton 765 (Ar— 271725); E. D. Schulz 2195 (Gg--1622hu, Wi). Walker Co.: Albers & Warnock 15136 (Au, N); Parl Parks & Cory 10437 (Tr); Tharp s-M- (67i1/ 35] (Ca—882856, Vi); Warner sen. (Au, W—111)303). 3). Waller Coe: We H. Lewis 5372 (Nb); H. B. Parks 2432 (Au). Washington Co.: C. Oe Albers 33013 (Au), 33017, in part (Au); Brackett 253 (Au), SMe (Apr. 21, 1939] (Au); Gander 7596 (s4—25183)5 E. Hall L3h (W, - 709 , Pr, W—~71996). Wharton Co.: J. A. Drushel 1 TOL35 (N)3 As Re Moldenke 210 (Fe); E. J. Palmer 6622 (Po--2l "EO, W——-1581013). Will- iamson Co.; Whitehouse s.n. n. (Cedar Park, July 8, 1929] (Au, Au) + High Island [Galveston Co.]: sis M. Clark 39hh (B) . County ye mined: Belfrage s.n. [28.h.1 S); Herb. Univ. Texas sen. (AU, Au); Kuntze “eet 190) wy ee Sane [Fasc. IV, 16h9] (Ka); H. Ness 14h (Tr, Tr, Tr, Tr, Tr, Ir, Tr, Tr, Tr, Tr, Tr, 7%» 1965 Moldenke, Monograph of Verbena 507 Tr, Tr, Tr), sen. (Tr). ARIZONA: Cochise Co.: R. D. Camp 2611 (Wi). CULTIVATED: Belgium: M. Martens s.n. (h. b . LOV. v. 1840) (Br). New York: G. V. Nash s.n. (1903; N. Y. Bot. Gard. ‘cult. Pl. 18690] (N); N. Taylor sen. [1903 N. Y. Bot. Gard. Cult. Pl. 18690] (N). Texas: Ee Hall SMe SiS fad LOCALITY OF COLLECTION UNDETER- MINED: McKay ay 2109 09 (S) - — — eae BOOK REVIEW Alma L. Moldenke "Algal Cultures and Phytoplankton Ecology", by G. E. Fogg, xiii + 126 p illust. The University of Wisconsin Press, Madison, Wieconsih. [1964] 1965. $5.50 Now readers . —- the same privilege of a about this information as did the listeners to Dr. Fogg's lecture 6 [from which this PA er book is compiled] at the Saismeetis of essaereegs se "introduction" the author explains how ele control- a ted pata and biochemical studies of culturing t phyto— planktonic algae can offer much to interpreting the detailed obser- vations made in natural bodies of fresh and salt water er is known to survive in the dark even with supplemented substrates, and the conclusion that there is no single criterion for measuring eae xt he explains "the growth of algae in continuous and synchron- ous pti using the Myers and Clark turbostat and the Monod chemostat for studying population density in controlled media. lative amounts of the rganic nutrients when ¢g similar conditions, even t is known that algal groups are charac terized by r pigments, carbohydrates, stero g "the general features of the phytop in es and the sea" is mo goiaayinian their patchy horizontal and verti- cal distribution with the growth rate of the biomass varying great- ly with cell size, Ce crease and intermingling of holoplanktonic types en in the wa- ter and of meroplanktonic types emerging from resting stages in bottom deposits. "Some other aspects of phytoplankton periodicity" are discussed. The "distribution and seasonal succession" of the various organ- 508 PHYTOLOGIA Vol. 11, no. 7 isms in the biomass includes its pioneers and its climax form whose positions are controlled by the rates of growth and the bio- chemistry of the internal and external environments Although the author stresses the need for study of truly plank- es index, a fine asia and several helpful illustrations -- some even in color _— Index to Authors in Volume Eleven King, R. M., 217 Oswald, F. Moldenke, 1 L., 69, 143, 213, Reed, C. a “ts, By 433 287, 358, 431, 507 Swalien, J. R » 145, 152, 361 Noldenke, He ‘.. oe "50, 155, Tovar, O., x , 290, 400, 435 Wurdack, J. J., 377 Index to Supra-specific Scientific Names in Volume Eleven Abies, 307, 29, 71 os Lh7 Acacia, 90 a peed, Acer, hls iahelensholdes + An Acrospermum, 312 raliaceae, Aecidium, i6h, 202, 3h2 Araucaria, 2)5 Aedes, hedes, 360 Arenaria, ae Agastache, 341 ai stoloots Agave, 9 Ageratum, 218 Albizzia, 69 ene 428, 433 scone i a Aspidella, 29 jn ’ Aspilia, ri 322 Alnus, 138 Aster, Alopecurus, 289 Astra sous 430 Aloysia, 72, ly Taiaareetboeen, 1b Alsinaceae, 308 Atriplex, 426, 430 Amanita, 431, 433 Atropa, 426 Amanitina, 29 Aubletia, 20, 60 Amanitopsis, 29 Avicennia, 72 Amblyarrhena, 388, 389, 391 Axinaea, 38h: Ambrasia, 339 Azotobacter, 427, 43k Amphicarpa, 7 Amplariella, 29 Bazzania, 25 Anabaena, 428 Begonia, 426 Anagallis, 105 Bellucia, 399 Andropogon, Abe. 199, 3h1 Berbena, 12h Anemone, Beta, 1965 Index 509 Bidens, 256 Cyclamen, 126 Boraginaceae, 3h1 Datura, 426 Bouteloua, 199, 3h1, 489 Daucus, 426 Brachyotum, 377-383 Delphiniun, a Brassica, 26 Deschampsia, 373 Brassicaceae, 256 Diandrolyra, 152, sh Brizo » 361, 372 Dicaeoma, 16h, 202, 3h2 Bromus, Digitaria, 3) Buchnera, 62, 81, 164 Diospyros, 31 Bulbulus, 152, 1h Diostea, 195 Calliopsis, 202, 343, 437, 462 Dissanthelium, 361-376 Callirho'¥, 285 Duranta, Canadea, 135, h6é Echinochloa, 340, Lik Gentes a 203, 2 edr carer Eragrostis, 289 bas 431 Erigeron, 218 TaetTTE , 307 Erinus, 6 Celtis, Celtis, 32 Eryngium, 307 Centrospermae, 359 simum, 256 Cephalanthera, 26 Erysiphe, 16h, 202, 342, 343, Cephalan Cercospora, 202, 501 Eucalyptus, 82, 83 Chaenanthera, 366, 391, 396 Eupatorium, 218, 31 Ghaenopleura, 391, 396 Euphorbiaceae, 200 Chaetodon, 385 Euphydras, Chlorella, 508 Eurotia, 489 Chiorobacteriaceae, 1h} Fabaceae, 203 Chrysopsis, 195 Fagus, Chrysothamnus, )30 Festuca, ig 307 Cleome, he3s-L25, 432, 435 Flour: Clerodendrum, 70 Foliosae, 7 e. 55, 60, Tlidemia, 305, 397-399 —I37, 162, 185, 196, nh “he Clio 3%0 335, 22° 500 Clostridium, 28 axinus, 163, 430 hranea, 87 Fuchsia, Conposttse, 217, 218, 359 Galium, 311 Condalia, . Gaura, gr Gerardia, ranvolvilacee, kl Gaeiaris, ks 50, 2 2, 13h Coreopsis, 198, 310 Ze, 263, 268; 280, 315, 38, eet 321, 323, 32h, ha, 438 Cremanium, 387, 391-395 Glecona, 32 ssa, Glossocentrum, 385, 386 Cruciferae, 359 Gnaphalium, 218, 286 ’ Graminastrum , 365 Cupressus, 307 Grandularia, "262, 280 510 Gutierrezia, 13, 489 G po al Haplopappus, 30 deoma, 199 Henriettella, 397, 399 Hilaria, Holophyllae, 55, 127, 162, 196, 22 Ichnantmus, 73-79, 145~151 3, 00g Oa. kek, hel s aOey 185, 196, 239, 256, jodina, 271 Iridaceae, 286 b Jasminum, 3 Junceae, 121 Juncus, 105 Junellia, 121, 312 Juniperus, 13, 430, 489 Labiatae, 378, 90 Lactarius, 429 Lactuca, FHI TOLOGIA Vol. lL, no. 7 Lobelia, 16h, 200, Tobeliacese, 16h, Loreya, 399 Lycopersicum, 26 iythrum acs acranthae, 7, 20, 60, B2y.aam Gattoniols » 432 Melastomataceae, 377 Melindres, 82, 185 Miconia, 385-396 Micranthae, 55, 127, 162, 196, 239, 2 oes 335, 422, 500 Mnium, ) Monodus % onstropels, 433 a, 307 Woblise, ra 1,0, 321, 422 Nolina, 9 Nothoscordun, 83 Ocymum, ta 105 » 343 Lea ‘ite, 153 Opuntia, 13, 1 199, 451 61 Tamiaceae, 325 554 225, 16k, 200, ee + aoe 63, ls, 436, 472 Lespedeza, 198 Liatris, 196 Linaria, 290 Lippia, 72, 86 Liriodendron, 359 Lithospermum, 163 Lobaria, 133, L3k ¥2 Cae, 30 Peristereona, 251 Persicaria, 339, 3h0, 1) Petalostemon, 203, 285 1965 Index $11 Phalaridium, 363, 366 Sanolus, 32 Phlox, 290 Santalaceae, 271 Phryma, 36 Sarcobatus, 1,30 Phyia, 72; 8b, 127, °257, 4%, Satureja, 311 Ol Scenedesmus, 428, 433 Phyllanthus, 200 Schizachyrium, 285 Phyllosticta, 202 Schizophyllae, 256, 500 Picea, L29, 431 Schuttelworthia, 22, 273 Pinus, 285, 286, 307, 430, 431, Schuttleworthia, 22, 243, 267, Scirpus, 1} Piresia, i = Scrophulariaceae, 8, 164, 200, —S 738 331, Se, 2 antago, 19 Scutellaria, 163 Platanus, 30 Senecio, ]26 Pleroma, 379 Septoria, ee 202, 343 oa O, 361, 372 Sherardia, somnsettne, ee 22, 2h3 ’ 2u2, aa, i a Smuttleworthia, 22, 242, 243, Populus, 199, 341, 431 Silene, 199 ulacaceae, 286 Simmondsia, li5 Potentilla, 163 Solanum, 340, 360, 26 Primulaceae, 3)2 Solidago, 19 56 Priva, 120, 2 Prosopis, 12, 13, 453, 459, L Spermatophyta, 358 eee k7l Sphaerotheca, 3)3 inia, 16h, 202, 343 Spirogyra, Pycnanthemum Quercus, 126, 199, 286, 30, stacivs, le poacibadie tices Bal nada ane ao ue 162, 16h, + “er od 203 ss = Stanl 3 aphanus, 426 eya, Reitzia, 152, 153 Stellaria, 12, 18h eum, 126 pene a pal Rhoeo, 26 Stenochloa, #3, 365 sites 340 Stenotaphrum, 1 Ringentes 4,36 ah hn 143 Rudbeckia, 200, 203, h61 Stylodon, 72, 86, 16h, 21, ’ a oe fe 252 Rutaceae, 359 ae waeem, Salix, 199, 3h1 Synedre Salvia, 16h, 201, 336 Tetragoniaceae, 200 512 PHITOLOGIA Vol. 11, no. 7 Teucrium, 257, 31 293, 295, 297, 299, 301-305, Thiorhodaceae, 1) 307-329, 331, 333, 335-347, tia 2 349, 351, 353, 355, 357, 400, iboushina, 379 4O1, 403, 405, 40 415, rer Pn ye 417, ae 435-13, LL5- Tilia, li) Wh8, 450- * Wee-hg3, l95, Tithymalopsis, 285 497-503, $08" Tococa, 396, 397 Verbenaca, 6, 7, 20, 55, 60, Toxicodendron, 8h, 337 62, 121, 127, 162, 185, 196, Tradescantia, 426 259, ae iar 256, 315, 322, Tragopogon, 161 cent eee poe Teseesceess 72, 13h, 139, 1bh, Eepee olen; ee Verbenia, 191 Triodia, 13 Verbenna, 290 Lenreee, 457 Verbina, 63, 498, 502 am Vermicularia, Umbelliferae, 369 woe ae ? panna a 3 Veronica, 200, 327 Vacciniaceae, 378 Varcdeastte. he Bi hcicditaeintnielstudhak ind 2 Valeriana, Verticillatae, 36 Valerianaceae, 3),2 Vicia Nospent, 191 Vilfa, 361, 365 Verbascum, 30 Viola. 198 Verbena, 1, 3-7, 9-15, 17-35, Vitex, 70 ping aa eee 3-89, 91, Wedelia, 218 155-159, 161-165, 167 Xanthium, 2 69 AAD 30 171, 173, 175, 177, 179-191, Yucca, 13, 31, 451, 453, 460, 207, 209 B 211, 213, 219, 221, 223, 228, zephyranthes, 83 227, 229, 231-243, 2h5-253, Se ee 255-257, "259-287, 290, 291, -—- we we wee ei eee ee ee ee Dates of Publication of Volume Eleven Number 1 -—- October 17, 196) Number 2 — December 23, 196) Number 3 -- December 24, 196) Number 7 -- March 19 , 1965 Figure 20. Distribution of Verbena xutha in the United States Herbarium curators who have material of this species from additional counties are asked to send it to the author for verification and record, so that future editions of this map may be more complete Mapping by counties done by Andrew R. Moldenke = ana RRS. a bec ‘Z Se a ee ee, | is RMON mi Rance Sy, st ee" ieee _ ae CT ONcKEY os ms MePHE RS: o bel Simmons C ROORS Eo ea i LoGaN con | | . | © wircncocx | Rep naa "Low s 2 SE sem (AAMRINS DecaTUR | NORTON . i tee Tans Tee AURORA or ANTELOPE] s @. ‘SOM Sn 6 para puymouTH Son MERCER nor S rweena wy vee wa wort # ; r 7 a Tone CEDAR MUSCATINE 0 QUISA CuuNTOR corr y 2 (SUA OER ‘ye INCKSON D caRRO WHITESIOE Lee | ena | BUREAU | or i} rey : = ae t ae K/ a we = © Y re © - — YS 0 ot ca OE ae A tae ete ee na aly I ae | DMA [ao = 5 ws} A cae ae Q ae ngs KEYED TO COLUMN A r] eee, of Cleartype index Book * s , AG . eames 24 eae CLEARTYPE ney : ‘ TRADE MARK REG. U.S. PAT. OFF. tie Scale of Miles : fag 0 100 200 eee MAP NO. 125 FOURTH ee JUDICIAL DIVISION = COPYRIGHT 4 AMERICAN ZOMPANY, INC. ORIGINATORS . SOLE PUBLISHERS rc CLEARTYPE MAPS OLORPRINT MAPS fe pe THIRD JUDICIAL DIVISION | WwW Copyright ic< , ~ | Pytight, American Map Co., Inc., New York, No. 14324 9° ; First aX yupicia ONION s : . fs \. 9 Scale of Miles gp ro) NX ee. KYRE J on g ee o” \ 1,7 HONOLULU . % ° PART ¢e LISIANSKI, 3 «* AYSAN : 2. ALEUTIAN ISLANDS e ™. THIRD JUDICIAL DIVISION - e = cs re Et aoa ° OS OE il iles ~ - 92d g__Miles 390 z i I 5 I é I 1 a I 3 [ 10 [ ti aa a DRY “ES PO I 2i I 2 i es es z = L at a ES a eee SE = ! 2 , x , es . *