BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY ’ ‘ ee. 2 ee | PRINTED AND PUBLISHED AT THE BOTANICAL MUSEUM CAMBRIDGE, MASSACHUSETTS BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY VOLUME 1 BOTANICAL MUSEUM CAMBRIDGE, MASSACHUSETTS 1932-1933 TABLE OF CONTENTS NumsBer I (June 7, 1982) Nomenclatorial Notes on Plants growing in the Botanical Garden of the Atkins Institution of the Arnold Arboretum at Soledad, Cienfuegos, Cuba. By F.’Tracy HupBarb AND ALFRED REHDER. NuMBER II (June 29, 1932) A New Epidendrum from Spanish Honduras. By OAKES AMES Two New Orchids from Yucatan and Trinidad. By Oakes AMES AND CHARLES SCHWEINFURTH On the Identity of Kirsanna. By F. Tracy HvuBBARD NumbBer ILL (July 28, 1982) An Extension of Range for Epidendrum rhyn- chophorum. sy OAKES AMES NuMBER IV (February 7, 1935) A New Octomeria from Spanish Honduras. By Oakes AMES The Honduran Species of Lepanthes. By OakrES AMES [v ] PAGE NuMBER V (February 24, 1933) A New Oncidium from Honduras. By Oakes AMES Epidendrum Edwardsii NuMBER VI (March 20, 1933) An Addition to the Genus Hexadesmia. By OakEs AMES A New Bletia from Spanish Honduras. By OAKES AMES NuMBER VII (April 24, 1933) An Addition to the Genus Epidendrum. By Oakrs AMES NumBer VIII (June 14, 1983) A New Epidendrum from Honduras. By OakrEs AMES Pleurothallis Lewisae NuMBER IX (August 12, 1933) A New Genus of the Pleurothallidinae. By Oakes Ames Saunders Refugium Botanicum Volume II. By F. Tracy Huspsarp NUMBER X (October 24, 1933) A. New Sobralia from the Republic of Honduras. sy Oakes AMES A New Bletia from Mexico. By Oakes AMES AND CHARLES SCHWEINFURTH [vi] PAGE 32 INDEX OF ILLUSTRATIONS Roman numerals refer to the numbers, Arabic numerals to the pages. Bletia papillifera 4dmes ........ 4. .VIi:7 Bietia tenutionaud. BG. 2 2 es ees BT Epidendrum comayaguense Ames. . . . . . VIIL:3 Epidendrum Edwardsii Ames ....... . Vi7 Epidendrum hondurense Ames... . . . .) VILE Kpidendrum rhynchophorum A. Rich. & Gal. . ILL: Hexadesmia hondurensis Ames... . . . . . Viti Lepanthes Edwardsii Ames... ...... .I1V:7 Lepanthes hondurensis Ames... 2 2 2. 2. TVS Lepanthopsis anthoctenium (Reichb,f.) Ames . UX :29 Lepanthopsis densiflora (J@odr.) Ames... . UX:18 Lepanthopsis floripecten ( Reichb.f.) Ames 1X :17,21,29 Lepanthopsis melanantha (Reichb,f.) Ames. . IX:29 Lepanthopsis microlepanthes (Griseb.) Ames . UX:25 Octomeria hondurensis Ames . . . . . 2. . PV: Oncidium hondurense Ames... . . . . . . Vib Pleurothallis Lewisae Ames ....... . VIILI:7 Pleurothallis ruscifolia (Sw.) R.Br. 2... 2). UN:5 Sobralia Edwardsii Ames .......... Mii Stelis Endresii Retchd ff... °0>ess s ss. VED Stelisnubis Ames... 7. eee ss ws ED Stelis Powellii Selif. 2. See re. ws . ERED Stelis vestita Amée . "vee et. fw SURO [ vii | INDEX TO GENERA AND SPECIES ACTINOPHLOEUS Becc. 1:1 Maearthurii (1. Wend.) Becc. apud Wigman . . Is im ANDROPOGON Auct., non L, Sorghum Brot. subsp. sativus Hack. [ var. ] Durra Hack, 1:10 ARECA L. Sechellarum Hort. apud Ciee . . oo ae ASTROCARYUM G.F.W. Mey. aureo-pictum Hort. apud Verschaff. ... . 4 aureo-punctatum Lem... 1:8 Borsigianum C.Koch . . 1:8 Borsigii Hort. apud C.foch. <0. .- > ee pictum Hort. apud Balf.f. 1:9 BLETIA Ruiz & Pav. papillifera dmes 2. VE: Pottsii S. Wats. 2... X36 tenuifolia Ad. & S. 2. . XG tuberosa (L.) Ames , X36 BRACHYCHITON Scholl & Endl. diversifolius R.Br... . Ts populneus (Scholt & Endl.) i ae a, CAESALPINIA L. pulcherrima Sw. var. flava Hort. apud L.H. Bail. & Rehd. . [:9 CALADIUM Vent. .. .1:8 bicolor (4it.) Vent. . . 1:8,4 esculentum Vent. . . . . Lit CALAMUS L. Zalaccea Gaertn. . . . . 1:9 CATHARANTHUS G. Don 1:4 pusillus G.Don . . . . [s4 roseus (L.) G. Don [34 var. albus (Sweet) cS} OY a een uae a: var. ocellatus (Sweet) GOO ine. ee COLOCASIA Scholt I antiquorum Schott . .. 1:5 esculenta (L.) Schott I var, antiquorum (Schott) Hubb. & Rehd. . 2 2 1:5 COUMAROUNA Aubl. . . 1:5 panamensis Pittier 2. . 1:5 DIPTERYX Schreb. . . . 1:5 panamensis ( Pittier) Hubb. & Rehd. . 2. 2 155 DRYMOPHLOEUS Zipp. 1:1 EPIDENDRUM L. arbuscula Lind/. . . . Uh: [ ix ] bisulcatum Ames . Vil Brenesii Schltr. VII Broadwayi 4. & S. . I comayaguense Ames VII Kdwardsii Ames. Aoribundum HBK. hondurense Ames mf : VE paniculatum Ruiz & Pav. [1:1 I:1 E35 [:1 I1:1; V:6 1:4 Ts1 VII:2,4 rhynchanthum Sechitr. . 1:2 rhynchophorum 4. Rich. & Gal. . II1:1,2 tipuloideum Lind/. 11:6,7 vieji Reichb f- VIII: GARDENIA Ellis. a et augusta Merr. me Gt. Hlorida L. . E56 jasminoides Ellis . ae es GLIRICIDIA HBK. Lambii Fernald 1:6 maculata Walp. 1:6 sepium (Jacq.) Walp... 1:6 HEXADESMIA Brongn. crurigera Lind!/, Vi: hondurensis Ames VI:1 HOLCUS L. 1:10 Caffrorum Thunb. 1:10 Durra Forsk. 1:10 halepensis 1. 1:10 lanatus L. 1:10 Sorghum Ll. . 1:10 var. caffrorum L.H. Bail. 1:10 var. Durra L.H.Bail. 1:10 KENTIA B/. Macarthurt Hort. ; 142 LEPANTHES Sve. appendiculata Ames IV :3 densiflora Rodr. IX:11 Edwardsii Ames LV :3,4,8 guatemalensis Sch/tr. . IV:8 hondurensis Ames IV :3 Johnsonii Ames IV:8 secunda Rodr. IX:11 Tuerckheimii Schltr. IV :8 turialvae Reichb,f. IV :3,4 LEPANTHOPSIS Ames 1X25 anthoctenium (Reichb.f- ) Ames densiflora (Rodr.) Ames 1X :1 floripecten ( Reichb f. ) Ames . 1X:7,16,19,23,28 1 IX :2,7,11,16,19,20,27,28,31 melanantha (Reichb f. ) Ames 1X :7,19,28,24,27,28 microlepanthes (Griseb. ) Ames IX :19,24 LOCHNERA Reichb. . 1:4 rosea Reichb. . 1:4 LOCHNERIA Heist. . . 1:4 LOCHNERIA Scop. . 1:4 LONCHOCARPUS HBK. maculatus DC. . 1:6 seprum DC, ae Fi MACHAERIUM Pers, Tipu Benth. T:11 MARANTA L, divaricata Rose. ie 3 gibba Sm. ae by OCTOMERIA R. Br. brevifolia Cogn. IV: hondurensis 4 mes IV: ONCIDIUM Sie. glossomystax Reichbf. = Vit genomus Krénzl. Vi: hondurense Ames ve pusillum Reichbf. . Ve Titania Sehltr. eat PHOENICOPHORIUM H.Wendil. ..... .1:7 Borsigianum (C. Koch) Stuntz 1:8 Sechellarum H.Wendl. — . 1:9 PLEUROTHALLIS R. Br. anthoctenium Reichb.f. EX :2,7 congestiflora Cogn, . . IX:11 Horipecten Reichb. f. 1X :1,2,3,7,11 ies 19 VIIL:5 Aoripicta Lindl. Lewisae Ames . melanantha Reichb.t. 1X :7,16,19,23 microlepanthes Griseb. IX :16,19,24 ruscifolia (Siw. ) R.Br. IX :2,3,4,19 sertularioides Lind/. . 11:4,5 unilateralis Cogn. .TX:11 yucatanensis 4. & 8. . [1:4 POECILODERMIS Schott & Endl. populnea Schott & Bnd. . bs nN POINCIANA L. puleherrima L. var. flava (Hort. apud L.H. Bail. & Rehd.) Hubb. & Rehd. 1:9 PTYCHOSPERMA Labill/, Macarthuriti H.Wendl. . 1:2 ROBINIA L. maculata HBK. 2... 36 Sepium Jacq. . . . . . 1:6 variegata Schlecht... . 1:6 SALACCA Reinw. . . . . 1:9 edulis Reinw,. . . . . . 1:9 SALAKKA Reinw. apud BI. 1:9 edulis Reinw. apud Bl. 1:10 SOBRALIA Ruiz & Pav, crocea Reichb,f. X:1 Edwardsii Ames . . . Xit SORGHUM Moench 1:10 Caffrorum Beauv. 1:10 Durra Stapf . 1:10 vulgare Pers. var. caffrorum (Thunb. ) Hubb. & Rehd. 1:10 var. Durra (Forsk. ) Hubb. & Rehd. 1:10 STELIS Sw. Endresii Reichb,f. IX:8 nubis Ames IX :8 Powellii Schltr. IX:8 vestita Ames IX :8 STEPHENSONITA Hort. apud we feemtte 6.0, 2. AST grandifolia Hort. Veitch . 1:9 STERCULIA L. eaudata Heward . . . 2 1:2 diversifolia G. Don By STEV ENSONIA Dunean . 1:7 Borsigiana 1..H.Bail. . . 1:9 grandifolia Duncan Sechellarum Hort. apud Van Houtte... . 1:9 TALINUM Adans. paniculatum (Jacg.) Gaertn. forma variegatum (Hort. apud W. Mill.) Hubb. & ST a re 47 e patens Willd. var. variegalum Hort. apud We ee TT © lh Ce Bell A TIPUANA Benth. Tipa Lillo . a tipu (Benth.) Lillo apud Stunt . VARNERIA Stickm. augusta Stickm. . VICIA L. Ervilia (L.) Willd. VINCA L. L:11 rosea La. var. alba Sweet Bed ey Be var. ocellatus Sweet . var. oculata W. Mill. P35 ZALACCA Rumph. Blumeana Mart. . 11:8 edulis Reinw. apud BI, [ xii | ERRATA 1:2, line 1 for Marcarthurii read Macarthurii 1:10, line 22 for Sorghum vulgare L. read Sorghum vulgare [’ers. T:11, line 15 for Tipuana tipu ( Benth.) Hubbard & Rehder comb. nov. read Tipuana tipu (Benth.) Lillo apud Stuntz in Invent. Seeds & Pls. Introd. no. $1 (1914) 88 (Bur. PI. Indust.) VIII:1, line 3 for Epidendrum viejii read Epidendrum viejt. VIII:5, line 4 for Izabel read Izabal VIIL:5, lines 11 and 12 for Guatemalen read Guatemalan IX :2, line 25 ; for P. ruscifolia (Sw.) R.Br. read P. rusejfolia (Jacq.) R.Br. IX :8, line 16 for Sretis NuBIA Ames read Stretis NuBIS Ames [ xiii ] *y wig +> vite ‘ i. aug? BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CampripGe, Massacuusetts, JUNE 7, 1932 No. NOMENCLATORIAL NOTES ON PLANTS GROWING IN THE BOTANICAL GARDEN OF THE ATKINS INSTITUTION OF THE ARNOLD ARBORETUM AT SOLEDAD, CIENFUEGOS, CUBA BY F. Tracy HuBBARD AND ALFRED REHDER In preparing a list of the plants growing in the botanical garden at Soledad, several instances of nomen- clatorial difficulty have been revealed. Ordinarily the new combinations made necessary by the transfer of a name from one genus or from one specific name to another are clear without further explanation, but in some cases the nomenclature has become so involved that it has seemed advisable to discuss it. The following notes are issued in order to publish those new combinations which are necessary and to clarify those points which seem ‘involved and uncertain. ACTINOPHLOEUS Beccari in Ann. Jard. Bot. Buitenz. 2 (1885) 126, an teatu. Beccari in Malesia 1 (1877) 42 originally charac- terized the group as a subgenus of Drymophloeus Zipp. The spelling of the subgeneric name is 4 ctynophloeus. The species cultivated in the garden is: 1 Actinophloeus Marcarthurii (H. Wend.) Beccari apud Wigman in Bull. Dépt. Agric. Indes Néerl. no. 81 (1909) 1, nomen — Beccari in Webbia 4 (1913) 154 — Radermacher in Ann. Jard. Bot. Buitenz. 85 (1925) 12. [The first complete description of the species |. Kentia Macarthuri Hort. apud Belg. Hort. 27 (1877)241, nomen (as Mac Arthuri)— H. Wendland apud ‘I’. Moore in Florist & Pomol. 1879 (Aug. 1879) 115, text cut — H. Wendland apud III. Gar- tenz. 28 (Dec. 1879) 265, t. 36. _Ptychosperma Macarthurti H. Wendland apud Kew Rept. 1882 (1884) 55. This species, nomenclaturally, has had a compli- eated history. Originally introduced about 1877, it was placed in the genus Aentia and was described (rather inadequately) in 1879. Later it was transferred to Ptychosperma. In horticulture it has been frequently cultivated under both names, but usually as Kentia Macarthuri. In 1918 Beceari removed it to Actino- phloeus where we believe it more correctly belongs. Brachychiton populneus (Schott & Endl.) R. Brown in Bennett Pl. Jav. Rar., pt. 3 (1844) 284. Sterculia diversifolia G. Don in Loudon, Hort. Brit. (1830) 392, nomen—G. Don, Gen. Syst. Gard. & Bot. 1 (1881) 516, non Brachychiton diver- sifolius &. Brown in Bennett Pl. Jav. Rar. pt. 3 (1844) 234 which is Sterculia caudata Heward in Herb. Cunn. apud Bentham, FI. Austral. 1 (1863) 230. Poecilodermis populnea Schott & Endlicher, Melet. Bot. (1882) 33. Following Engler and Prantl and recent authors it has seemed best to accept the genus Brachychiton Schott & Endl. It is not, however, possible to apply the earl- 2 iest specific name ‘‘diversifolia’’ on account of the earl- ier Brachychiton diversifolius R. Br. Robert Brown's binomial undoubtedly refers, as pointed out by Bentham (Fl. Austral. 1 (1886) 280) to Sterculia caudata Heward in Herb. Cunn. Consequently the oldest available specific name is ‘‘nopulnea’’ based on Poecilodermis populnea Schott & Endl. and the correct combination is Brachychiton pop- ulneus (Schott & Endl.) R. Br. CALADIUM Ventenat, Deser. Pl. Nouv. Jard. Cels, livr. 3 (1801) t. 80 and in Roem. Archiv. Bot. 2 (1801) 847. In discussing this genus, W. F. Wight (in Safford in Contrib. U.S. Nat. Herb. 9 (1905) 208) advanced the idea that Caladium Vent. applied to the genus com- monly known as Colocasia Schott. The basis of his argument was that Ventenat drew his generic name from Rumphius who used this name for certain species of Arum. ‘This is undoubtedly true. Starting with this premise, Wight argues that the only species common to both Rumphius and Ventenat is ‘‘esculentum’’, which is likewise the fact. Granted the truth of both statements, we cannot agree with Wight’s deduction that Caladium Vent. must apply to that part of the material included in the genus typified by ‘‘esculentum’’. In the first place, from the point of view of Ventenat’s genus the governing factor is what did he describe, not the source from which he drew the generic name. . The generic description seems broad enough to cover both types of plants in- cluded in his subsequent list of species composing the genus. However, the species to which he refers through- out the text and which he illustrates is C. bicolor and no mention of ‘‘esculentum’’ is made until at the end he sums up those species of Arum which he believes belong 3 to the genus. Ventenat’s treatment of Caladium in Roemer Archiv fiir die Botanik follows the same course, discuss- ing C. bicolor and at the end mentioning ‘‘esculentum’’. Consequently, we feel that the standard species of Caladium is C. bicolor and not C. esculentum as advo- cated by Wight and accepted by certain recent authors. CATHARANTHUS G. Don, Gen. Syst. Gard. & Bot. 4 (1838) 95. Lochnera Reichenbach, Consp. Reg. Veg. (1828) 134, non Lochneria Heist., non Lochneria Scop. On account of the earlier uses of the name Lochnera Reichb. must be replaced by Catharanthus G. Don. Dalla Torre and Harms (Gen. Siphon. fase. 6 (1904) 406) are obviously in error when they refer Catharanthus G. Don to Vinca L. as Don treated but two species — C. pusillus and C. roseus — both of which are included in Lochnera by Engler and Prantl. The transfer of the plant known both as Vinca rosea L. and Lochnera rosea Reichb. was made by G. Don (Gen. Syst. Gard. & Bot. 4 (1888) 95). The synonymy of the two varieties of this species growing in the garden is: Catharanthus roseus (L.) G. Don var. albus (Sweet) G. Don, Gen. Syst. Gard. & Bot. 4 (18388) 95. Vinca rosea 1... var. alba Sweet, Hort. Brit. (1827) 274. Catharanthus roseus (L.) G. Don var. ocel- latus (Sweet) G. Don, Gen. Syst. Gard. & Bot. 4 (1838) 95 (sphalmate occellatus ). Vinca rosea V.. var. ocellatus Sweet, Hort. Brit. (1827) 274. Vinca rosea 1. var. oculata W. Miller in L. H. Bailey, Cycl. Am. Hort. 4 (1902) 19385, t Colocasia esculenta (L.) Schott var. anti- quorum (Schott) Hubbard & Rehder, comb. nov. Colocasia antiquorum Schott in Schott & Endlicher, Melet. Bot. 1 (1882) 18. Authors in general, who separate this variety from the species, have made it the species and have reduced C. esculentum (L.) Schott to varietal rank. While there is hardly any doubt that var. antiquorwm is the phylo- genetic type of the species, it is nomenclaturally in- admissible to reduce an older specific name to varietal rank under a name of later date of publication. — In consequence the ranks must be reversed and C. esculenta become the species with a variety antiquorum. Dipteryx panamensis (Pittier) Hubbard & Rehder, comb. nov. Coumarouna panamensis Pittier in Contrib. U. S. Nat. Herb. 18 (1917) 236. The genus Dipteryx Schreb. is conserved over Coumarouna Aubl. ‘The transfer of this species makes a new combination. Gardenia jasminoides ///is in Phil. ‘Trans. 51, pt. 2 (1761) 985. Varneria augusta Stickman in Linnaeus, Amoen. Acad. 4 (1759) 186, nomen. Gardenia florida Linnaeus, Sp. Pl., ed. 2 (1762) 305. Gardenia augusta Merrill, Interp. Rumph. Herb. Amb. (1917) 50, 485, 547. Merrill undoubtedly went too far when he accepted Stickman’s Varneria augusta as valid publication. According to the International Rules, it can only be considered as a nomen nudum as the Rhumphius refer- ence given is to a plate lacking analytic details and furthermore at the time of publication the genus to o which the species belonged was undescribed. Gliricidia sepium (Jacg.) Walpers, Repert. Bot. Syst. 1 (1842) 679 — Standley in Contrib. U. S. Nat. Herb. 23 (1922) 482. Robinia Sepium Jacquin, Enum. PI. Carib. (1760) 28. Robinia maculata Humboldt Bonpland & Kunth, Nov. Gen. et Sp. 6 (1824) 398 (Quarto ed.), 309 (Folio ed.). Lonchocarpus sepium De Candolle, Prodr. Syst. Nat. 2 (1825) 260. Lonchocarpus maculatus De Candolle, Prodr. Syst. Nat. 2 (1825) 260. Robinia variegata Schlechtendal in Linnaea 12 (1886) 301. Gliricidia maculata Walpers, Repert. Bot. Syst. 1 (1842) 679. Gliricidia Lambi Fernald in Bot. Gaz. 20 (1895) 538. There has been considerable difference of opinion as to the authority to be given for the combinations Gliricidia sepium and Gliricidia maculata. According to Index Kewensis 1 (1895) 1038, ‘‘H.B. & K. Nov. Gen. et Sp. vi. 393 in nota’’ [Quarto ed.] is given as the place of publication of both combinations. Neither combination, however, was made in either the quarto or folio edition of Humboldt Bonpland and Kunth, Nova Genera et Species Plantarum... or in Kunth, Syn. PI. 4 (1825) 81. Urban, Symb. Antill. 2 (1900) 288, and other authors refer both combinations to Steudel, Nomencl. Bot. ed. 2, 1 (1840) 688. While both combinations exist there, they are distinctly synonyms and are cited as being made by Kunth [HBK.]. Consequently, as they are published in synonymy, they have no valid standing. The earliest actual publication of the combinations which we have been able to find is in Walpers, Repert. Bot. Syst. 1 (1842) 679. Maranta gibba Smith in Rees Cycl. 22 (1819)no. 2. Maranta divaricata Roscoe, Mondr. Pls. (1828) t. 7. K. Schumann in Engler, Pflanzenreich LV. 48 (Heft 11) (1902) 126 cites M. gibba Sm. in Rees in synonymy under M. divaricata Rose. It is evident that Schumann had no knowledge of the date of issue of volume 22 of Rees Cyclopedia as he fails to cite the year. It contains the earliest name of the species. PHOENICOPHORIUM JH. Wendland in Ul. Hort. 12 (Feb. 1865) Mise. 5 and (May 1865) t. 433. Stevensonia Dunean, Cat. Roy. Bot. Gard. Mauri- tius (1868), xomen — L. H. Bailey, Gentes Herb. 2 (19380) 192. Stephensonia Hort. apud Van Houtte in FI. des Serres 15 (May 20, 1865) 177, in synon. We are unable to agree with the deductions of Prof. L. H. Bailey in Gentes Herbarum 2 (1930) 192 and,— in agreement with H. C. Skeels (in Bull. Torr. Bot. Cl. 58 (1981) 49),— we cannot accept his reasoning that the generic name Phoenicophorium should be out- lawed. ‘There is no provision in the International Rules that we feel could be construed to cover the case. The genus Phoenicophorium was proposed and well described by H. Wendland in [llustration Horticole 12 (Feb. 1865) Mise. 5 and later illustrated by him in the same periodical (May 1865) t. 433. Apparently Prof. Bailey has overlooked the date of publication of the Miscellanées when he states, “‘A portrait of the palm appeared in L’ Illustration Horticole bearing date April 1865,...’’ The footnote date on page 5 of the Mis- cellanées is Février 1865 and we can find no reason why this date should not be accepted. ~ ! The generic name Stevensonia was first published by James Duncan in 1868 (Cat. Roy. Bot. Gard. Mauri- tius). It isanomen nudum. Furthermore we are not able to agree with Prof. Bailey that Van Houtte in Fl. des Serres 15 (May 20, 1865) 177, t. 1595-1596 pub- lished or even intended to publish a generic description under Stevensonia. The fact that Phoenicophorium was used as the page heading and as the title entry in the Table des matiéres on the back cover of the Livraison with Stevensonia grandifolia Dune. in italics and clearly in synonymy convince us that Van Houtte was con- sidering the species as a Phoenicophorium. Moreover the date of publication of the ‘‘Onziéme livraison du tome XV” is clearly given on the cover of the copy in the Gray Herbarium, which is bound with the covers in place, as **Paru le 20 Mai 1865”°. We are therefore unable to accept Stevensonia Dune. as the generic name. The species cultivated in the garden is: Phoenicophorium Borsigianum ((C. Koch) Stuntz in Invent. Seeds & Pls. Introd. no. 81 (1914) 88 (Bur. Pl. Indust.). A strocaryum aureo-pictum Hort. apud Verschaffelt, Cat. Ill. no. 12, mise. page 5 (fide Van Houtte in Kl. des Serres 15 (1865) 177, in synon.) — Ver- schaffelt, Cat. Hl. no. 74 (1864) 12, nomen. Astrocaryum Borsigianum C. Koch in Wochenschr. Giirt. u. Pflanzenk. 2 (Dee. 22, 1859) 401 —C. Koch apud Regel in Gartenfl. 10 (1861) 29. Areca Sechellarum Hort. apud C. Koch in Wochen- schr. Giirt. u. Pflanzenk. 2 (1859) 401, mention. A strocaryum Borsigu Hort. apud C. Koch in Woch- enschr. Giirt. u. Pflanzenk. 2 (1859) 402, mention. Astrocaryum aureo-punctatum Lemaire in Il. Hort. 6 (Dec. 1859) in nota sub t. 229, nomen. fa) Stevensonia grandifola Wunean, Cat. Roy. Bot. Gard. Mauritius (1868), 2omen. Stephensonia grandifola Hort. Veitch apud Proce. Roy. Hort. Soe. Lond. 4 (1864) 134, nomen. Phoenicophorium Sechellarum H. Wendland in II. Hort. 12 (Feb. 1865) Mise. 5 and (May 1865) t. 433. Stevensonia Sechellarum Hort. apud Van Houtte in Fl. des Serres 15 (May 20, 1865) t. 1595-1596, in synon. Astrocaryum pictum Hort. apud Balfour f. in Baker F]. Mauritius (1877) 388, a synon. Stevensonia Borsigiana L. H. Bailey, Gentes Herb. 2 (1980) 192. Poinciana pulcherrima L. var. flava (Hort. apud L. H. Bail. & Rehd.) Hubbard & Rehder, comb. nNOv. Caesalpinia pulcherrima Sw. var. flava Hort. apud L. H. Bailey & Rehder in L. H. Bailey, Cycl. Am. Hort. 1 (1900) 206. The combination for this horticultural variety has never been made under Poinciana. SALACCA Reinwardt in Syll. Ratisb. 2 (1825) 3. Zalacca’ Rumphius, Herb. Amb. 5 (1747) 118, t. 57, fig. 2— Reinwardt apud Blume in Roemer & Schultes, Syst. Nat. 7, pt. 2 (1880) 1334. Salakka Reinwardt apud Blume, Cat. Gew. Bui- tenz. (1828) 112, nomen. The oldest valid post-Linnean spelling of the generic name is Salacca of Reinwardt, and from the standpoint of Latin it is also more correct than Zalacca. The species involved is: Salacca edulis Reinwardt in Syll. Ratisb. 2 (1825) 3. Calamus Zalacca Gaertner, De Fruct. et Sem. PI. 9 2 (1791) 267, t. 139. Salakka edulis Reinwardt apud Blume, Cat. Gew. Buitenz. (1828) 112, nomen. Zalacca edulis Reinwardt apud Blume, Roemer & Schultes, Syst. Nat. 7, pt. 2 (1830) 1384. Zalacca Blumeana Martius, Hist. Nat. Palm. 8 (1838) 201, t. 123; t. 159, fig. 3. SORGHUM Moench, Meth. (1794) 207. The recent trend has been to conserve Holcus L. for Holeus lanatus 1. and to place H. halepensis L. and H. Sorghum 1. in Sorghum Moench. In doing this, two new combinations are made necessary, namely: Sorghum vulgare Pers. var. caffrorum ( Thunb.) Hubbard & Rehder, comb. nov. Holcus Caffrorum Thunberg, Prodr. Pl. Cap. 1 (1794) 20. Sorghum Caffrorum Beauvois, Agrost. (1812) 181, 164, 178. Holeus Sorghum W. var. caffrorum L. H. Bailey, Gentes Herb. 1 (1923) 1838. Sorghum vulgare 1. var. Durra ( Forsh. ) Hub- bard & Rehder, comb, nov. Holeus Durra Forskal, Fl. Aegypt.-Arab. (1775) 174. Andropogon Sorghum Brot. subsp. sativus Hack. [var.] Durra Haeckel in De Candolle, Monogr. Phan. 6 (1889) 516. Sorghum Durra Stapf in Prain, Fl. Trop. Afr. 9 (1917) 129. Holeus Sorghum WL. var. Durra 1. H. Bailey, Gentes Herb. 1 (1928) 132. The synonymy of the varieties of Sorghum vulgare is very involved and the identity of many varietal names 10 is, at present, not well known. Consequently, although evidence tends to lead one to believe that ‘‘caffrorum’’ and ‘‘Durra’’ are not the oldest varietal names for these forms, it has seemed advisable to retain them until a more complete study of the species has been made. Talinum paniculatum (Jacg.) Gaertn. forma variegatum (Hort. apud W. Mill.) Hubbard & Rehder, comb, nov. Talinum patens Willd. var. variegatwm Hort. apud W. Miller in L. H. Bailey, Cycl. Am. Hort. 4 (1902) 1767. The combination for this horticultural form does not appear to have been made previously under 7°. pani- culatum. Tipuana tipu ( Benth.) Hubbard & Rehder, comb. nov. Machaerium Tipu Bentham in Hook Journ. Bot. & Kew Gard. Mise. 5 (1853) 267. Tipuana Tipa Lillo, Contrib. Conoe. Arbol. Argent. (1910) 58. Lillo in transferring this species to T’puwana elected to use the other and possibly more correct spelling of the vernacular name. In consequence the adoption of Bentham’s earlier specific name necessitates a new com- bination. 11 BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY Campripce, Massacuuserts, JUNE 29, 1982 No. A NEW EPIDENDRUM FROM SPANISH HONDURAS BY OakKES AMES From Mr. J. B. Edwards, who has been collecting in the vicinity of San Juancito, Honduras, there has been received a species of Epidendrum which appears to be undescribed. It is closely related to EH. arbuscula Lindl. differing from it in both vegetative and floral characters. ‘The unbranched stems, linear-lanceolate leaves, shorter floral bracts and pedicels, and the some- what smaller flowers in a paniculate rather than racemose inflorescence are conspicuous differentiating characters. In H. arbuscula the naked portion of the peduncle is very short and stout, while in the new species the pe- duncle below the flowers is elongated and very slender. Epidendrum Edwardsii 4 mes, sp. nov. Herbaterrestris. Radicesalbidae, glabrae, carnosae. Caules plus minusve erecti, foliosi, foliis distichis lineari- lanceolatis, infra folia vaginis persistentibus arcte ad- pressis. Pedunculus elongatus, satis gracilis, basi bractea scariosa vaginanti instructus. Inflorescentia laxe paniculata, pauciflora. Sepala similia, oblonga, acuta, patentia. Petala spathulata, acuta. Labellum trilobatum, columnae adpressum et cum ea in tubum adnatum; lobi laterales subdolabriformes, margine irreg- ulariterdenticulato ; lobus medius subsimilis,emarginatus cum apiculo obtuso interjecto; discus prope apicem columnae conspicue bicallosus, per medium leviter cari- natus. Columna sectionis. Plant 4.5-6 dm. high. Roots whitish, smooth, coarsely fibrous. Stems slender, erect, about 4 mm. in diameter near the base, concealed by the tubular closely appressed sheaths with which the leaves are articulated. Leaves numerous, distichous, the lower ones 3—6 cm. long and 1 cm. wide, lanceolate, acute, their sheathing bases 3.5-5.5 em. long, tinged with violet-purple, the upper ones up to 16 cm. long, 1—1.7 em. wide, linear- lanceolate, acute, chartaceous when dry, 1.5—2 em. apart on the stem. Inflorescence erect, subpaniculate, termi- nal, 6.5-12 em. long. Peduncle with the rachis 5.5—11 em. long, provided with one or more sheathing acute scarious bracts. Flowers 5 to7, fleshy. Pedicels about 8 mm. long, smooth, slender. Bracts of the panicle 3.5-10mm. long, usually much shorter than the pedicels, linear to triangular, strongly coneave. Ovary about 5 mm. long, smooth. Lateral sepals 10-11 mm. long, 2.75 mm. wide, oblong, acute, spreading, margins strongly revolute. Dorsal sepal similar, 12 mm. long, erect. Petals 11-11.5 mm. long, 2-3 mm. wide, spatu- late, acute. Labellum distinctly 3-lobed, including the basal part that is adnate to the column 11-12 mm. long; free portion up to 8 mm. long, 10 mm. wide across the lateral lobes, with a more or less clearly defined central keel on the upper surface, and with 2 hemispherical complanate calli on the dise in front of the apex of the column; lateral lobes sub-dolabriform, about 5 mm. wide with an irregularly dentate or crenulate margin; middle lobe up to 11.5 mm. wide, about 3.5 mm. long, 9 ~ emarginate with a blunt apicule, margin entire, undulate or sometimes sparsely dentate. Calli 1.5 mm. long, 1 mm. high. Column 6 mm. long, very slender toward the base, dilated upward; margin of the clinandrium dentate, tinged with purple. RepuBiic OF Honpuras, Department of 'Teguci- galpa, Vicinity of San Juancito, old lumber road at Rosario. ‘Terrestrial on exposed rocky ledges at 5000 feet altitude. Sepals and petals white to pale pink; lip white with purple dot. February 14, 1932. J. B. Hd- wards 122. (T'ypE in Herb. Ames No. 87891.) TWO NEW ORCHIDS FROM YUCATAN AND TRINIDAD BY OakES AMES AND CHARLES SCHWEINFURTH Among orchids sent to us for identification there are two undescribed species, a Pleurothallis submitted by the Field Museum of Natural History and an Epi- dendrum collected by W. EK. Broadway. These may be characterized as follows: Pleurothallis yucatanensis Ames & Schwein- Surth, sp. nov. Herba epiphytica, pusilla. Rhizoma repens, cauli- bus numerosis brevissimisornatum. Folia oblanceolata, apice rotundato minute bilobulata. Racemi laxiflori, pauciflori. Flores pusilli. Sepala lateralia anguste lanceolata, uninervia Sepalum dorsale majus, lanceo- latum, concavum. Petala triangulari-linearia. Label- lum oblongum, apice late rotundatum, parte inferiore leviter dilatata. Columna apice alato-dilatata. Plant very small, up to 5 cm. tall, epiphytic, recall- ing P. sertularioides Lind]. Rhizome creeping, entirely invested by imbricating scarious nervose sheaths. Roots fibrous, glabrous, stout for the plant. Stems very short, approximate, ascending, unifoliate, about 2.5 mm. long, when young entirely invested by imbricating tubular searious sheaths. Leaf oblanceolate, 8-15 mm. long, about 2.8-3.5 mm. wide, broadly rounded at the apex with aminutely bilobed apiculate tip, gradually narrowed to a 1-jointed petiole, very rugose in the dried specimen. Inflorescence conspicuously surpassing the leaf, solitary, about 3 cm. tall; peduncle capillary, adorned below the middle with one close tubular sheath; racemes very lax, 2—~ to 5-flowered. Floral bracts loose, infundibuliform, I. twice or thrice surpassed by the slender ascending pedi- cels. Flowers small, segments of the perianth sub- parallel, slightly thickened toward the apex. Lateral sepals free, narrowly lanceolate, 2.5 mm. long, 1— or obscurely 3-nerved, acute, carinate on the outer side. Dorsal sepal slightly longer, lanceolate, about 2.8 mm. long, obtuse, concave, obscurely 3-nerved. Petals linear-triangular, 2.5 mm. long, 1—nerved, acute, slightly oblique. Labellum much shorter, simple, very shortly clawed; lamina oblong or lanceolate-oblong in outline with the erect sides of the lower half slightly dilated from a cuneate base, about 1.4 mm. long, broadly rounded at the apex, 3-nerved, minutely auriculate at base. Column short, almost 1.2 mm. long, with an abrupt quadrate wing on either side of the upper half, extended in a foot. Anther cupuliform. Pleurothallis yucatanensis suggests P. sertularioides Lindl., but differs in having racemose inflorescences and simple lip. Mexico, Yucatan, Campeche, Tuxpena, C. L. Lun- dell 912, November 8, 1981. Common on tree trunks and old logs. (‘TyrE in Herb. Ames No. 37882). Epidendrum Broadwayi Ames & Schweinfurth, sp. NOV. Herba humilis, epiphytica. Caules caespitosi, tenuiter subclavati, omnino velati, mono-vel diphylli. Folia anguste linearia, graminea, acuta. Inflorescentia singula, quam folia multo brevior, laxe racemosa. Sepala lateralia oblique elliptico-lanceolata, apice conspicue carinato-mucronata. Sepalum dorsale oblanceolato- oblongum. Petala oblanceolato-linearia. | Labellum apici columnae adnatum, trilobatum, lobo terminali majore subquadrato triangulari-acuto vel retuso et apiculato. or Plant small, epiphytic, up to 17 em. tall, closely related to EH. tipuloideum Lindl. Stems approximate, slenderly subclavate, about 4—-5.5 em. high, altogether invested by close tubular scarious sheaths which waste into fibres, one- or commonly two-leaved at the apex, rugose in the dried state. Leaves narrowly linear, up to 11.7 cm. long and 4.2 mm. wide (generally much shorter), acute, the mid-nerve more or less prominently suleate above and carinate beneath. Inflorescence much shorter than the leaves, the longest scape about 4.5 em. high, clasped at the base by « loose tubular sheath which is up to 1.6 em. long. Raceme loosely 4— to 7-flowered. Floral bracts minute, lanceolate, membranaceous, many times exceeded by the slender pedicels. Pedicellate ovary up to 1.8 cm. long, laxly spreading. Flowers large for the plant, yellow in the dried specimen, rather fleshy in texture. Lateral sepals obliquely elliptic- lanceolate, prominently adnate to the column at the base, 11 mm. long, 3.2-4 mm wide, dorsally carinate- mucronate at the tip, prominently 7-nerved near the base. Dorsalsepal oblong-elliptic or oblanceolate-oblong, about 10 mm. long, 8.1-8.7 mm. wide, acute, conspicu- ously 7-nerved below. — Petals oblanceolate-linear, 8.6-9.3 mm. long, more or less oblique, acute, 3-nerved. Labellum adnate to the apex of the column, 3-lobed; lamina about 5.1—5.5 mm. long from the center of the cordate base to the tip, about 6.4 mm. wide when ex- panded, bicallose at base; lateral lobes semiovate or rounded, obtuse, extending to about half the length of of the lip, erose on the margins; terminal lobe much larger, irregularly subquadrate, with a prominent tri- angular acute apex, or retuse and apiculate. Column stout, about 6.2 mm. long, dilated above. Anther broadly ovoid. Pollinia 4, complanate-ovoid. Epidendrum Broadwayi is very similar in habit to 6 Epidendrum tipuloideum Lindl., but it has a dissimilar anterior lobe of the lip and 8-nerved petals. The flowers also are apparently of a different color, being yellow in the dried specimen. TrinipAb, North coast, Saut d’Kau W. HE. Broad- way 7444, January 18, 1981. Grows in tufts on a tree. (Type in Herb. Ames No. 37893). ON THE IDENTITY OF KIRSANNA BY F. Tracy HuBBArbD In September 19380 Dr. C. S. Coon of the Peabody Museum submitted specimens of seed for identification under the name of Kirsanna. These had been collected for him in Morocco by Gordon Browne in the Valley of the Iherrushen where they are grown by the Tribe of Gzennaya, Rif. The following notes of economic interest in regard to this seed were furnished by Dr. Coon. In the Rif the seeds are used for making bread after they have been thoroughly soaked to extract the poisonous element which they contain. They are placed in flat wicker baskets and weighted down in streams and allowed to remain until the poisonous element is judged to be re- moved. In the Rif the Berber vernacular name of the seed is Shasanna and the Arabic name is Kirsanna. Dr. Coon further states that the name of the seed in Armenia is Kushna and that it is used there for feed- ing cattle after the poisonous element has been removed by soaking. After several unsuccessful attempts the seed was finally grown to maturity at the Botanical Garden and proves to be Vicia Hrvilia (1..) Willd. BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY Campripae, Massacuuserrs, Juty 28, 1932 AN EXTENSION OF RANGE FOR EPIDENDRUM RHYNCHOPHORUM BY OAKES AMES THE MEXICAN ORCHIDS published by A. Richard and H. Galeotti, in Annales des Sciences Naturelles in 1845, and described with exasperating brevity, (the average length of the descriptions hardly exceeding three lines of print), have been a source of perplexity for students of the Mexican flora. Several of the species are known only through the type specimens in the herbarium of the Muséum d’ Histoire Naturelle de Paris. Among the ex- tremely rare orchids is Mpidendrum rhynchophorum. This species was originally described as follows: ** Pseudobul- bis ovoideis 1—phyllis; folio oblongo-lanceolato, acuto; flor. luteis, racemo 4—5 floro: labello trilobo, adnato, lobis lateralibus trunecatis, intermedio angusto-lineari. ”’ Very recently, while collecting for the Arnold Arbo- retum in the Department of Comayagua, Honduras, Mr. J. B. Edwards found Hpidendrum rhynchophorum in full flower in May. It has seemed worth while to publish an account of Mr. Edwards’ specimens and by means of a drawing, to remove the ambiguity caused by Richard and Galeotti’s inadequate description. ‘The extension of range from Mexico to Comayagua, Honduras, is remarkable, but in this regard it has been found that several Mexican orchids extend to Comay- agua, indicating that intensive exploration may prove that many more may be characterized by a wider range than has been thought probable. In the original description, HMpidendrum rhyncho- phorum is referred to the section Encyclium, although the plants are described as being pseudobulbous with a divided labellum adnate to the column. John Lindley, when he prepared his monograph of Epidendrum for his Folia Orchidacea, placed the species in the section Aulizeum where it seems properly to belong. In con- formity with the characters laid down for Aulizeum the labellum is adnate to the lower half of the column, with the lateral lobes erect and concealing the anther. The following description has been prepared from dried specimens, and flowers preserved in alcohol. Epidendrum rhynchophorum 4. Rich. & Gal. in Comptes Rend. Acad. Sci. Par. 18 (1844) 805, 512, nomen; in Ann. Sei. Nat. ser. 8, 8 (1845) 20. EKpidendrum rhynchanthum Sch/tr. in Beihefte Bot. Centralbl. 36, abt. 2 (1918) 467 sphalm. Rhizome stout, prorepent, emitting coarse, fibrous roots. Pseudobulbs approximate on the rhizome, 3—5.5 em. long. lageniform, slender or sometimes much swol- len, monophyllous, when immature completely concealed by ample, tubular sheaths. Leaves 13-26 em. long, up to 1 em. wide, linear-oblong, unequally bilobed at the apex. Peduncle terminal, 7-9 cm. long below the ra- ceme, slender with an elongated, spathaceous sheath at base in the axil of the leaf; raceme 6-12 cm. long, loosely 3-7-flowered, with the flowers 1.3-2.5 em. apart. Bracts of the raceme squamiform, about 8 mm. long. Pedicels with the ovary 1.5-2 cm. long, glabrous. Flowers brownish near the base of the sepals, otherwise yellowish Yd E.PIDENDRUM Iifryncdophorum fj i | \ i \ | : | \ | v1 / ii / | \ Ny . ) f \ | y iy H } | | ; INS y Ua j; ? / ; | f) y | /} Y % 4 / | 4 mS / | Hf \ | NH V7 : | ? | y f o I Hy | H 4 Ne , | Nick § Val or yellowish green with the lateral lobes of the labellum whitish. Sepals about 2.5 em. long, scarcely 2 mm. wide, directed obliquely forward, linear-triangular, acute. Pet- als similar, about 1.5 cm. long, 3-nerved. Labellum up to 13.5 mm. long, conspicuously three lobed, adnate to the lower half of the column; mid-lobe up to 9 mm. long, about 1 mm. wide, linear, tapering gradually toward the acute tip; lateral lobes about 8 mm. long and 2.5 mm. wide, rotundate-dolabriform, erect, appressed to the apex of the column; disc conspicuously bicallose, the calli pa- piliform. Column stout, about 5 mm. long, adnate for about one half its length tothe labellum, with a conspicu- ous terminal rostriform appendage which is dorsal, por- rect and blunt. Mexico, Vera Cruz, on the parasitical Andromeda at 4,000 feet, Galeotti 5077. (Tver in Herb. Mus. d’ Hist. Nat. Par.). ReEPuBLic OF Honpuras, Department of Comayagua, Las Jaguas in the vicinity of Minas de Oro. Epiphyte in open mountain forest, at 4,200 feet altitude. Flowers yel- low and brown. May 8, 1982. J. B. Hdwards 143. EXPLANATION OF ILLUSTRATION Plant drawn natural size from Honduran specimens ( Edwards 143). 1, front view of column showing the rostriform dorsal appendage ex- ceeding the anther. 2, side view of labellum in natural position. 3, side view of labellum and column with the labellum depressed to show point of attachment. 4, labellum with the lateral lobes spread out. 5, the flower drawn to show the relative proportions of the sepals, petals and labellum. BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY Campriper, Massacuuserts, Frpruary 7, 1933 No. A NEW OCTOMERIA FROM SPANISH HONDURAS BY OAKES AMES AMONG ORCHIDS recently collected in Honduras by Mr. J. B. Edwards a new species of Octomeria has been recognized. There are no close allies of this species among the Octomerias recorded as natives of Central America. It isin habit similar to the Brazilian O. brevifolia Cogn. from which it differs in the size and form of the labellum. From dried specimens it appears to belong to the section of the genus characterized by semiterete leaves. Octomeria hondurensis Ames, sp. nov. Herba epiphytica, parva. caespitosa, caulibus gracilibus tereti- usculis. Folium subeylindraceum, leviter arcuatum, valde carnosum. Flores satis numerosi, densiusculi, fasciculati. Sepala oblongo-lanceolata, subacuta, trinervia. Petala similia. Labellum distincte trilobatum; lobis lateralibus erectis; lobo medio oblongo, apice tridentato ; disco prope basim bicarinato. Columna generis. I.piphytic herb about 10cm. high. Stems caespitose, terete, 8-9 cm. long,several-jointed, with fibrous remains of sheaths adhering near the base, monophyllous, about 1 mm. in diameter when dried. Leaf about 2.5 cm. long, about 1 mm. in diameter. semiterete, acute, lightly sul- [1] OCTOMERIA Onodur En SIS XN mes cate above, much exceeding the inflorescence. Peduncles 5 mm. long, faseiculate, in the axil of the leat, subtended by closely appressed sheaths. lowers opening in succes- sion, only one present ata time. Lateral sepals spreading, 4.5 mm. long, 2 mm. wide, oblong-lanceolate, acute or subacute, 8-nerved. Dorsal sepal similar. Petals 4 mm. long, 1.5 mm. wide, oblong-lanceolate, subacute, 3- nerved. Labellum 3 mm. long, 8-lobed; lateral lobes 1.5 mm. long, .5 mm. wide, erect and porrect, faleate, sub- acute or obtuse; middle lobe 2 mm. long, 2 mm. wide, quadrate-oblong, tridentate at the apex with the lateral teeth rounded and the middle one acute; dise bicarinate at base. Column produced at base forming a distinet foot, about 1.5 mm. long, Rereusiic or Honvuras, Department of Cortez, Kl] Jaral, Lake Yojoa. Epiphyte in dense tropical forest, 2,000 feet altitude. Sepals, petals and lip lemon yellow: column white. October 29, 1982. J. B. “dwards 304. (Tver in Herb. Ames No. 39043). ILtusrrRatTiOoN: Plant drawn natural size from dried specimens. Flower much enlarged (at right). Labellum much enlarged (at left). THE HONDURAN SPECIES OF LEPANTHES BY OAKES AMES THREE SPECIES of Lepanthes are now known to be natives of the Republic of Honduras; L. hondurensis Ames, published in 1931, L. turtalvae Reichb.t., recently collected by J. B. Edwards; and L. Mdwardsu, which is described below Lepanthes hondurensis A mes in Proc. Biol. Soc. Wash. 44 (1981) 43. This species is related to L.. appen- diculata Ames, a Guatemalan plant, but is larger with differences in the structure of the labellum. ‘The flowers are extremely attractive, suggesting tiny butterflies with orange wings bordered with mauve, spread out on the vellow background formed by the sepals. ‘The accom- panying plate was prepared from the type. Repusiic or Honpnuras, Department of Atlantida, Lancetilla Val- ley, near Tela. March 11, 1923. Ames IT. 171, IT. 176: Department of Comayagua, Pito Solo, Lake Yojoa. Kpiphyte in dense forest, at 2,000 ft. altitude. August 1982. J. B. Edwards 90, 93 and 230. ILLusrRATION: Plant about four fifths natural size, 1,inflorescence much enlarged. 2, labellum. 3, column (posterior view). 4, pollinia. Lepanthes turialvae Reichb.f. Beitr. Orch. Centr. Amer. (1866) 57. Reeusiic oF Honpuras, Department of Comayagua, Siguatepeque. Kpiphyte in open mountain forest, at 3,700 feet altitude. Sepals pink, petals and labellum red. September 26, 1982. J. B. Edwards 258: Fl Achote. Epiphyte in dense forest, at 3,900 feet altitude. Sepals light pink, petals and lip rose. September 27, 1932. J. B. Edwards 260. Lepanthes Edwardsii mes sp. nov. Caules se- cundaril vaginis laxis, hispidis, tubularibus omnino in- clusi, ascendentes, monophylli, caespitosi. Folia elliptica apice inconspicue tridenticulata, marginata, in petiolum contracta. Inflorescentiae folium excedentes, rachis frac- tiflexa. Sepala lateralia glabra, usque ad medium cohae- rentia, laminam ovatam bifidam formantia. Sepalum dor- sale lanceolatum, bene caudatum. Petala carnosa, lan- ceolata vel sub-semihastata, glabra, basi rotundata, apice subacuta. Labellum trilobatum; lobi laterales triquetri, anguste crescentiformes, acuti; lobus medius valde abbre- viatus, oblongus, acutus. Columna valde arcuata. Roots coarsely fibrous, whitish, smooth. Stems caes- pitose, monophyllous, 5-15 mm. long, slender, concealed by elongated tubular sheaths, erect or obliquely ascend- ing, rigid. Sheaths two to five, 2-6 mm. long, the basal one shortest, finely hispidulous on the prominent longitudi- nal nerves, dilated at the mouth, with the margins [+] XN 7 ee <7. ‘ Kond uUrensts Ames LEPANTHEW thickened and hispidulous, the uppermost one enclosing the petiole of the leaf and conspicuously dilated upward above the middle. Leaf elliptical, marginate, 5-15 mm. long, 4-7 mm. wide, contracted at base into ashort petiole, apex tridenticulate. Peduncles 1-4, equalling or exceed- the leaves, arising from the axils of the leaves, 7-12 mm. long to the base of the first flower, slender, smooth, usu- ally with a single bract near the middle, bearing six to seven flowers in succession, only one of which isexpanded ata time. Floral bracts infundibuliform, hyaline, hispid- ulous, equalling the short rigid pedicels. Flowers con- spicuous, the tips of the lateral sepals directed upwards, 13-14 mm. long from the tip of the dorsal sepal to the tips of the lateral sepals, whitish or yellowish with purple petals. Lateral sepals smooth, united to about the middle, forming a broadly ovate bifid 4-nerved lamina 6.5 mm. long, 4 mm. wide, free portion of each sepal triangular acute. Dorsal sepal smooth, 6.5 mm. long, 2 mm. wide, lanceolate, caudate. Petals fleshy, smooth, lanceolate or semi-hastate, obtuse orsubacute, 2-2.5mm. long, scarcely 1 mm. wide at the base, parallel, the inner margins ap- proximate above, closely appressed to the dorsal sepal. Labellum 8-lobed, with the lateral lobes fleshy. trique- trous, 8 mm. long, very narrowly crescentiform, closely appressed to the column at base, parallel or with the tips strongly incurved, at the tip closely appressed to the lateral sepals; mid-lobe minute, oblong, acute, about five times shorter than the lateral lobes, concealed by the column, ascending or abruptly inflexed. Column 1 mm. long, strongly curved downward over the middle lobe of the lip. Anther deeply retuse at the apex. This is a charming little orchid with flowers longer than its leaves, with membranaceous light yellow or whitish sepals and deep purple fleshy petals which hang [6] cH tES LEPANTHEW O Owed roost? downward, closely applied to the dorsal sepal and at base extend very little beyond the column, their basal ends close to the base of the labellum. It is closely allied to Lepanthes Johnsonu Ames from which species it differs in the relative proportions between the petals and the se- pals. In 1. Johnsoni the sepals are purple and twice long- er than in L.. HMdwardsiu while in both species the petals and labellum are nearly equal in length. From L. T'uerch- heimu Sehitr. and L. guatemalensis Schitr. it differs con- spicuously in the structure of the petals and in being vegetatively a much smaller species. Edwards no. 261 differs from all the other specimens referred to L. Hdwardsu in the color of the sepals which, except for a yellowish tinge at base, are purplish rather than yellow. It seems highly probable that the flowers of this species vary in color as they pass from youth to maturity. Repusiic or Honpuras, Department of Comayagua, Pito Solo, Lake Yojoa. Epiphyte in dense forest, at 2,000 feet altitude. August 26, 1982. J.B. Edwards 96. (Tyrer in Herb. Ames No. 39045) : Minas de Oro at Malcotal. Epiphyte in mountain forest, at 4,200 feet altitude. July 5, 1982. J. B. Edwards 196: San Luis at Coyocutena. In dense mountain forest, at 4,000 feet altitude. May 25, 1982. J. B. Edwards 171: Siguatepeque at El Achote. Epiphyte in dense forest, at 3,900 feet altitude. Flowers wine color, slightly lighter at base of petals. September 27, 1982. J. B. Edwards 261. ILtusrratrion: Plant natural size. 1, flower much enlarged. 2, labellum and column as seen from above. 8, labellum freed from the column to reveal the sharply upturned apex of the mid-lobe. 4, petal. 5, anther from the ventral side. 6, side view of the column and mid- lobe of lip, the lateral lobes forcibly spread apart. [8 | BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CAMBRIDGE, Massacuusetts, Fesruary 24, 193! No. A NEW ONCIDIUM FROM HONDURAS BY OAKES AMES From J. B. Edwards there has been received a spe- cies of Oncidium which appears to be new. It is related to O. glossomystax Reichb.f. and O. Titania Schltr. , but differs from them in having the dise of the labellum, in front of the callus, quite free from appendages. ‘The closest ally is undoubtedly O. gnomus Krinzl., but that species also differs in having a more complicated crest on the labellum. It is probable that the crest varies consid- erably and is not a reliable guide to specific distinction, but until more material has been examined it would be questionable procedure to merge the Honduran plant with O. gnomus. It is a charming species and with O. gnomus and O. Titania enjoys the distinction of being one of the smallest species of the genus vet known. Oncidium hondurense Ames, sp. nov. Planta minuta, ebulbosa. Folia plus minusve octo, ensiformia, equitantia, obtusa vel subacuta. Pedunculus foliis sub- duplo longior, uni-triflorus; bracteae minutissimae, acu- tae. Sepala lateralia libera, oblongo-lanceolata, apice bre- viter conduplicata, acuta, extus per medium carinata. Sepalum dorsale obovatum, apice conduplicatum. Petala oblongo-lanceolata, apice breviter conduplicata, acuta. Labellum unguiculatum; lobi laterales dolabriformes, [1] margine exteriore plus minusve crenulati; lobus medius multo major, quadrilobulus, lobulis lateralibus semi- orbicularibus, lobulis anticis minoribus, obtusis; totum labellum igitur in lobulos sex divisum. Crista usque ad apicem dilatata, apice quadridentata. Columna brevis. Alae gynostemii dolabriformes, vix crenulatae. Flores pallide lutei. In habit similar to Oncidium pusillum Reichb.f., but smaller in its vegetative and floral parts, mature plants hardly exceeding 2.5 cm. in height. Roots filamentous, whitish, smooth. Leaves ensiform, about eight, equitant, compressed, forming a fan-shaped plant, the lowermost ones often horizontal, the others obliquely ascending with the uppermost ones perpendicular, 7-18 mm. long, 2-4 mm. wide, obtuse or subacute, longitudinally nervose when dry, fleshy and smooth when fresh. Peduncle slen- der, one — to three — flowered, 2-8 em. long. Floral bracts about 2 mm. long, conduplicate, triangular, acute, much shorter than the pedicels, smooth. Pedicel together with the ovary about 5 mm. long, slender, smooth. Flowers vellow, 9-11 mm. long between the tip of the dorsal sepal and the tip of the six-lobed labellum. Lateral sepals free to the base, 2 mm. long, about .5 mm. wide, oblong- lanceolate, acute, somewhat conduplicate at the tip, cari- nate along the middle nerve on the outer surface. Dorsal sepal 3 mm. long, up to 2 mm. wide, obovate, broadest above the middle, tapering gradually toward the base, abruptly narrowed at the tip and shortly conduplicate, acute, lightly carinate externally along the middle nerve. Petals 2.5-3:mm. long, about 1.6-2mm. wide, oblong-lan- ceolate, shortly conduplicate at the acute tip, 1-nerved. Labellum about 7 mm. long, 3-lobed, the terminal lobe conspicuously bilobulate at the tip with a larger basal lo- buleon each side: lateral lobes 2mm. long, 2mm. wide, do- [2] O N y ; ab wi uy yp y, labritorm withthe margin entireor irregularly and coarse- ly crenate, 5-6 mm. wide from the tip of one to the tip of the other; isthmus .75 mm. long, about 2 mm. wide: terminal lobe 6 mm. wide at the base, about 4 mm. long. the terminal lobules 2 mm. long, 1 mm. wide, oblong, obtuse, entire or obscurely and irregularly crenate, sep- arated by a narrow sinus, inner margins partly contiguous or overlapping. Crest about 3 mm. long, shallowly sul- cate toward the base, dilated toward the anterior end where it becomes quadridentate (sometimes with suppli- mentary teeth or lobules), the anterior pair of teeth pro- jected forward on a quadrate base with the posterior teeth lateral one on each side and widely divergent. Column fleshy, 1.5 mm. long, with a pair of dolibriform wings at the summit; wings 1 mm. long, with the margin entire or obscurely and sparsely crenulate. Repusuic or Honpuras, Department of Comayagua, Pito Solo, Lake Yojoa. Epiphyte in dense forest, 2,000 feet altitude. Flowers yellow. August 28, 1982. J. B. Edwards 101 (Tyee in Herb. Ames No. 39050.) Intusrration: Above, plant drawn natural size. Below, plant much enlarged. [5] EPIDENDRUM EDWARDSII SINCE the publication of Mpidendrum Kdwardsii mes in Botanical Museum Leaflets, No. 2 (1982) 1, a drawing has been made from the type material to illus- trate the general habit of the plant and the details of floral structure. This drawing is reproduced on the fol- lowing page. ILtusrration: Plant drawn natural size. A single flower much enlarged. = SE cAm eC. EPIDENDRUM Codwardsti S NY pig N Yh = =~. ey LD ~ BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CamsripGr, Massacnuserrs, Marcu 20, 1933 No. AN ADDITION TO THE GENUS HEXADESMIA BY OAKES AMES THE NEW SPECIES of Hexadesmia described below might readily be mistaken for a tall, slender form of 17. crurigera Lindl., but in the texture and color of the flowers, and in the outline of the very different labellum it is quite distinct. Hexadesmia hondurensis Ames sp. nov. In habitu Hi. crurigerae Lindl. similitudinem gerit, sed labelli lobis et magnitudine differt. Caules caespitosi, valde elongati, diphylli. Folia anguste linearia. Pedun- culus quam folia brevior. Racemus plus minusve septem- florus. Sepala lateralia mentum rotundatum formantia, ovato-lanceolata, acuta, extus per medium leviter cari- nata. Sepalum dorsale oblongo-lanceolatum, concavius- culum. Petala oblonga, apice abrupte rotundata. Label- lum subpanduratum, apice valde bilobatum, prope basim bicallosum. Columna in pedem producta. Stems densely caespitose, erect, 12-15 cm. tall, min- utely pseudobulbous at the base, very slender and many- sheathed for about 6 em. then abruptly thickened to form an elongated secondary pseudobulb which is 8-9 cm. long, about 5 mm. in diameter when dry, subtended by several imbricating truncate sheaths and diphyllous at the apex. [1] Leaves linear, grass-like, 15-28 em. long, about 3 mm. wide, obtuse, apiculate, obliquely erect. Inflorescence racemose, arising 1n the axil of the leaves, 6-7 em. long, erect, with several imbricating scarious sheaths at the base, about 7-flowered. Bracts of the raceme about 5 mm. long, narrowly lanceolate, conduplicate. Pedicels with the ovary 1—1.8 em. long, slender, obliquely ascending, distichous. Flowers pale yellow with lavender striations on the labellum. Lateral sepals 5 mm. long, about 3mm. wide, ovate-lanceolate, acute, strongly concave, connate at the base forming a conspicuous rounded mentum, lightly carinate along the middle nerve on the outer sur- face. Dorsal sepal 5 mm. long, oblong-lanceolate, acute, closely appressed to the petals, 38-nerved to the tip. Petals 5mm. long, about 2 mm. wide, oblong, abruptly rounded at the tip, subacute. Labellum about 9 mm. long, 6 mm. wide across the bilobed tip, obliquely ascending then sharply deflexed, subpandurate from a rounded base; the apical lobes 3 mm. long, 8 mm. wide, rounded, slightly divaricate, obscurely erose on the margin. Dise with two obliquely placed basal calli from each of which a lightly raised keel extends nearly to the sinus formed by the ter- minal lobes, with a central keel between them and a short supplementary keel on each side. Column about half as long as the dorsal sepal, produced into a conspicuous foot. Pollinia six. Repustic or Honnuras, Department of Comayagua, Minas de Oro. Epiphyte in open mountain forest, at 4,000 feet altitude. Sepals and petals pale yellow, labellum pale yellow with lavender stripes. De- cember 26, 1982. J. B. Edwards 334. (Vyrer in Herb. Ames No. 39065); Epiphyte in open mountain forest, at 4,000 feet altitude. Se- pals and petals green with lavender ticking, labellum bronze. Decem- ber 28, 1982. J. B. Edwards 337. InLusrration: Plant natural size. 1, lateral sepal enlarged. 2, dorsal sepal enlarged. 3, petal enlarged. 4, flower enlarged. 5, lip and column viewed from the side, enlarged. 6, pollinia enlarged. [2] HEXADESMI taQ é Z, cy si Aono uUrensts CAmes A NEW BLETIA FROM SPANISH HONDURAS BY OAKES AMES AMONG recent collections made by J. B. Edwards in Spanish Honduras, a new species of Bletia has been re- cognized. It is without close allies and differs from all other Central American components of the genus in hav- ing the disc of the labellum closely beset with papilliform emergences below the base of the middle lobe. The fol- lowing description was prepared with the aid of flowers preserved in alcohol. Bletia papillifera 4 mes sp. nov. Cormus depresso- subglobosus vel pyriformis, vaginis pluribus scariosis ves- titus. Folia lineari-lanceolata, acuta, nervosa. Scapus erectus, elongatus, gracilis, pauciflorus. Sepala lateralia oblonga, acuta. Sepalum dorsale simile. Petala anguste oblongo-elliptica, obtusa, nervosa. Labellum supra me- dium trilobatum; lobi laterales erecti, triangulares, ob- tusi; lobus medius obcordatus, leviter carinatus, carinis crassiusculis; discus papillifer. Columna vix in pedem producta, elongata, prope apicem anguste bialata, prope basim utrinque lobata. Roots coarsely fibrous, whitish, smooth, suleate lon- gitudinally when dry, up to 7 em. long, about 2 mm. in diameter. Corm globular or pyriform, 2—2.5 em. long, covered with the fibrous remains of closely appressed sheaths. Leaves 12-20 em. long, 5-17 mm. wide, one or two in number, much shorter than the flower-scape, lin- ear-lanceolate, acute, tapering to a narrow sulcate base. Scape erect, about 40 cm. long, slender, about 2 mm. in diameter, with several closely appressed scarious sheaths. Flowers two to four, secund, whitish tinged with crim- [5] son, opening in succession, 2~4 em. distant from one an- other, the lowermost one with maturing capsule while the terminal one is in the bud. Bracts of the inflorescence about 1 cm. long, lanceolate, tapering gradually to an acute tip. Pedicels about 1 cm. long, slender. Ovary about 1 em. long. Lateral sepals 12-14 mm. long, 3.5 mm. wide, oblong, acute, lightly carinate beneath the tip or simply thickened, nervose. Dorsal sepal about 14 mm. long, similar to the laterals. Petals 14 mm. long, 3.5mm. wide, narrowly oblong-elliptical, tapering rather gradually to a rounded tip, many-nerved. Labellum 14 mm. long, 8-lobed; lateral lobes triangular, obtuse, erect, arising just above the middle, 9 mm. long to the base of the column, the nerves nearest the sinus thickened or verrucose; middle lobe 5—6 mm. long, 7.5 mm. wide near the middle, 3.5 mm. wide at the base, obcordate or cu- neate with a rounded retuse tip, with about eleven irregu- larly carinate or verrucose nerves, apical margin turned upward. Dise papilliferous along the nerves up to the base of the middle lobe, the papillae purplish. Column about 7 mm. long, dilated upward, with a rounded wing or lobe on each side at the base; anther 2.5 mm. long, conical, long-stipitate. Pollinia eight. Rereusirc oF Honpuras, Department of Comayagua, Kilometer 154, Tegucigalpa Highway. Terrestrial in marshy ground in open pine forest, 3,400 feet altitude. Sepals and petals cream with a scarlet tint; lip cream with many scarlet stripes. October 6, 1982. J. B. Edwards 277. (Tyrer in Herb. Ames No. 39066). ILLustrRATION: Plant natural size. 1, flower, front view, enlarged. 2, flower, side view, enlarged. 3, column enlarged. BisEITVIA A era Ames 2p \\ / f BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CamsBripGe, Massacuusetrts, Aprit 24, 1933 No. AN ADDITION TO THE GENUS EPIDENDRUM BY OAKES AMES IN 1923, while collecting in Spanish Honduras, | found near Tela, a species of Epidendrum which appears to be undescribed. It was growing in a swampy forest that re- ceives the flood waters of the Ulua River. The stems were pendulous from the trunk of a tree. Near Progresso, an inland town, I found another specimen of the same species growing in full sunlight on a rock beside a moun- tain stream. Owing to the pendulous habit of the stems, the flowers were inverted. This characteristic is clearly shown in the accompanying illustration. In August 1982, Mr. J. B. Edwards found this species near Lake Y ojoa, in the Department of Comayagua. Apparently it is widely distributed in Honduras although it has not yet been reported from neighboring countries. Epidendrum hondurense 4 mes sp. nov. Herba epiphytica, robusta. Caules foliosi, foliorum vaginis om- nino obtecti. Foliadisticha, oblongo-elliptica, acuminata. Inflorescentia terminalis, longe pedunculata, pedunculo vaginis pluribus ancipitibus ornato. Racemus multiflorus. Flores mediocres. Sepala lateralia oblanceolata, obliqua, concava. Sepalum dorsale simile. Petala angustissime spathulato-linearia. Labellum columnae valde adnatum, trilobatum, in basi cordatum; lobi laterales suborbiculares [1] vel dolabriformes; lobus medius bifidus; discus in basi bicallosus, per medium lineis tribus ornatus. Columna apice dilatata. Plant epiphytic. Stems tufted, 61-97 em. long, con- cealed by tubular closely appressed sheaths, the upper ones bearing leaves. Leaves jointed, seven or more, ob- long-elliptic ,acuminate, acute, distichous, obliquely as- cending, 15-21.5 em. long, 1.8-8.5 em. wide, gradually diminishing in size toward the summit of the stem, more or less coriaceous, becoming chartaceous when dry, the internodes 2—4 em. long. Inflorescence an elongated sim- ple raceme. Peduncle below the raceme 12.2—20.6 em. long, mostly concealed by distichous sheathing acute bracts which are 3.5 em. long and which gradually pass into the several widely separated bracts that arise alter- nately below the raceme and become brownish with age. Raceme about 10 em. long, hardly 4 cm. in diameter, many-flowered. Floral bracts triangular-lanceolate.acute, about 5 mm. distant from one another,the lowermost ones sometimes 1.9 em. long and somewhat surpassing the shortly pedicellate ovary, the upper ones becoming grad- ually smaller. Flowers about equal in size to the flowers of 1. paniculatum Ruiz & Pavon. Segments of the peri- anth spreading. Lateral sepals obovate-oblanceolate, 1.2 em. long, 8.5-4.4 mm. wide above the middle, acute, rather fleshy, greenish, somewhat verrucose on the outer surface. Dorsal sepal similar, oblanceolate, about 4 mm. wide above the middle, acute or subacute. Petals very narrowly spatulate, about 1 em. long, broadly obtuse, 1-nerved. Labellum adnate to the column for about half its length, lamina or free portion sharply deflexed, con- vex, trilobed, cordate at the base, about 5.2 mm. long; lateral lobes flabellate-suborbicular or dolabriform with the irregular margin often lobulate in front; middle lobe [2] EY PIDENDRUM | a Spee subquadrate-cuneate in outline, retuse or divided nearly to the middle into triangular-oblong lobules. Dise with two fleshy semiglobose calli at the base and with three raised longitudinal fleshy ridges through the middle. Column 7.1-7.9 mm. long. Anther semiglobose, 4- celled. Pollinia four. Epidendrum hondurense is allied to Id. paniculatum Ruiz & Pavon (12. floribundum HBK.), but is distinet in the outline of the labellum. In the type the sepals and petals exhibited a distinct tendency to become blackish with age. The lateral lobes of the labellum are sometimes almost free from lobules and exhibit only a slight tenden- cy toward a crenulate margin. This is true of Edwards no. 228. Reeusiic or Honpuras, Department of Atlantida, near ‘Tela. Epiphyte in swampy forest near the banks of the Ulua River. March 20, 1923. Ames s.n. (Tyrer in Herb. Ames No. $3710): Department of Yoro, near Progresso. On rock overhanging the Rio Pelo. Sepals yellowish; petals and lip cream color, the lip somewhat yellowish on the disc, the expanded part sharply deflexed from the column. March 8, 1923. Ames s.n.: Department of Comayagua, Pito Solo, Lake Yojoa .piphyte in dense forest, at 2,000 feet altitude. Flower stalk 3 feet high. Sepals and petals light yellow, lip cream color. August 13, 1932. J. B. Edwards 228. InLusrrRATION: Plant drawn three-fourths natural size, from the type. Flower enlarged. [+] BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CamsripGk, Massacuusetrs, JuNE 14, 1933 No. A NEW EPIDENDRUM FROM HONDURAS BY OAKES AMES THE SPECIESof Epidendrum described below appears to be without close allies in the Central American flora. Superficially it suggests the rare Mpidendrum viejiu Reichb.f. and in its vegetative structure resembles /. bisulcatum Ames and EI. Brenesu Schltr. From EH. Bre- nesu it differs conspicuously in having a very dissimilar labellum. Epidendrum comayaguense Ames, sp. nov. Caules perlongi, laxe ramosi. Folia valde coriacea, ovato- elliptica vel oblongo-lanceolata, acuta, plus minusve con- gesta. Racemi terminales, perbreves, plus minusve quin- queflori; pedunculus infra flores valde complanatus. Se- pala lateralia concaviuscula,ovato-lanceolata, acuminata, acuta. Sepalum dorsale lanceolatum. Petala lineari-lan- ceolata. Labellum simplex, suborbiculare, margine bre- viter et irregulariter fimbriato. Columna brevissima, carnosissima, antice valde excavata. Roots coarse, spreading, white, smooth. Stems e- longated, 6 dm. or more long, straggling, with numerous short branches which are entirely covered by rigid tubu- lar sheaths and bearing near the ends two to four leaves and a terminal dense abbreviated raceme of dull purple flowers. Branches from 3-15 em. long, the older ones [1] devoid of leaves and sometimes terminated by the rigid remains of the rachis of a raceme. Leaves crowded, coria- ceous, 4-10 cm. long, up to 2.5 em. wide, elliptic-ovate or oblong-lanceolate, tapering gradually to a subacute apex, the terminal one exceeding the raceme. Leat-sheaths somewhat complanate, 1.5-8 em. long. Inflorescence 3—5 cm. long, from the axil of a terminal leaf. Peduncle below the flowers conspicuously complanate and __ slightly winged. Bracts of the inflorescence 1—1.5 em. long, linear- lanceolate, acute. Pedicel and ovary 1.5 em. long, rigid. I‘lowers three to nine, very fleshy. Lateral sepals about 2 cm. long, 9 mm. wide, strongly concave, porrect, asym metrically ovate-lanceolate, acuminate, acute, with the median nerve lightly carinate. Dorsal sepal 1.8 em. long, about 7 mm. wide, slightly concave, lanceolate, acumi- nate, acute. Petals 1.7 cm. long, 8 mm. wide, linear- lanceolate, acute, margin entire or minutely scabrid. Labellum up to 2.5 em. long, 2 em. or more wide, sim- ple, suborbicular, deflexed, very finely and irregularly fimbriate on the margin. Dise smooth, ecallose. Column extraordinarily fleshy, deeply excavated in front with the lateral walls strongly thickened and rounded at the en- trance to the excavation ; stigma concealed ; androclinium entire, with the margin forming an almost perfect circle: pollinia four, completely filling the androclinium. Repusric or Honpuras, Department of Comayagua, Minas de Oro. Epiphyte in open mountain forest at 4,000 feet altitude. Sepals and petals dirty lavender with bright lavender veining, lip and column very dark, dull lavender. December 25, 1982. J. B. Edwards 333. (Typr in Herb. Ames No. 39042.) ILLusTRATION: Plant drawn natural size. 1, labellum much en- larged. 2, column much enlarged. [2] We ‘ W aff MY ba wAAZ Yc Albetes iit AS woe J wS% sry EPIDENDRU XY CO Shh Say Se 7 Coad = ae BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CampripGr, Massacuusetts, Auausr 12, 1933 No. 9 A NEW GENUS OF THE PLEUROTHALLIDINAE BY OAKES AMES THE ORIGINAL COLLECTION of Pleurothallis floripecten Reichb.f., which I believe represents a new genus, was made by Hermann Wagener near Jaji in Venezuela and was described by H. G. Reichenbach in Bonplandia 2 (1854) 25. So far as I know, only two specimens were studied by Reichenbach when he described the species. One of these is preserved in the Reichenbachian herba- rium in Vienna; the other was presented to John Lindley by Reichenbach and is preserved in the Lindley Herba- rium at Kew. Later, what Reichenbach thought to be the same species was represented by a living:specimen in the collection of W. Wilson Saunders at Reigate, England. Saunders obtained this specimen from M. Linden of Brussels, but whether or not it came originally from Venezuela is not known. The relationship between Pleurothallis floripecten and its closest allies has heretofore never been thoroughly in- vestigated. There are two substantial reasons why this should be so. First, P. flortpecten has proved to be ex- ceedingly rare; secondly, those species which resemble it closely have also proved to be rare. ‘Two of these species which were originally assigned to Lepanthes by Barbosa Rodriguez, were transferred by Cogniaux in Martius’ [1] Flora Brasiliensis to Pleurothallis on the evidence of sketches, there being no specimens available for critical study. That these Brazilian species were aberrant is e- vinced by the fact that Cogniaux proposed for their re- ception the new section Lepanthopsis. Later, in Urban’s Symbolae Antillanae, Cogniaux recognized Pleurothallis anthoctenium Reichb.f., a West Indian species, as being referable to the alliance formed by the Brazilian species and he introduced the section Lepanthopsis in Symbolae Antillanae to receive P. anthoctenium. To establish generic boundaries in the Pleurothalli- dinae vegetative characters have proved to be without weight. The perianth has also proved inadequate as va- rious degrees of cohesion between the sepals, while relied on for generic distinction, have been found useless because of misleading exceptions. The only structure that is funda- mentally serviceable is the gynostemium, an organ that is desperately difficult to reconstruct and interpret when it has been crushed by the pressure used in preserving specimens forthe herbarium, although in living specimens it exhibits clearly marked characters which are service- able in differentiating major groups or genera. The gynostemium of Plewrothallis floripecten and P. anthoctenium is very unlike what obtains in P. ruscifolia (Sw.) R.Br., the type of the genus Pleurothallis. The receptive stigmas are widely separated (cf. plate of Le- panthopsis floripecten, fig. 4, the heavily stippled area) and at anthesis are not conspicuously confluent along the frontal margin of the clinandrium beneath the rostellum as in P. ruscifolia; at the base the gynostemium is apo- dal, and at the summit conspicuously dilated with the posterior margin of the clinandrium cucullate. In P. rus- cifola the gynostemium is more or less elongated, cylin- drical with a pulvinate foot, and an obliquely truncate apex. The stigmatic lobes of P. floripecten are suborbi- [2] cular and spreading. The anther is transversely elongated and contains two large, waxy pollinia. ‘The widely sepa- rated functional areas of the stigmas at the dilated sum- mit of the footless gynostemium are sufficient evidence to indicate that P. floripecten is generically distinet from P. ruscifolia. (ef. plate of P. ruscifolia). The mere mention of widely separated receptive stig- mas, to one familiar with the genera of the Pleurothal- lidinae, should suggest the highly technical genus Stelis, and were it not for the very unusual perianth and the le- panthiform vegetative structures, I think that one would be inclined to refer P. floripecten to Stelis. But this would be a debatable procedure. ‘To make this clear several flow- ers of characteristic species of Stelis are here figured. I do not believe that it would be conformable to sound practice to remove P. floripecten and its allies from Pleu- rothallis and transfer them to Stelis. They would consti- tute as aberrant a group in Stelis as they surely do in Pleurothallis. (cf. plate of Stelis species). Barbosa Rodriguez in his studies of the Brazilian spe- cies concluded that they were referable to Lepanthes. Vegetatively they are lepanthiform and the widely spread- ing connate sepals resemble very closely the sepals of cer- tain species of Lepanthes, but the petals, labellum and gynostemium are very different from what obtains in that genus. In my opinion the species of the section Lepanthop- sis represent a distinct genus. Lepanthopsis Ames, gen. nov. Sepala plus minusve subaequalis, patentia, breviter vel conspicue connata; lateralia altius connata. Petala multo breviora, membranacea, orbicularia vel elliptica. Labellum ad basin columnae sessile, simplex, valde mem- branaceum. Columna brevissima, apoda, antice utrinque lobo plus minusve carnoso instructa; lobis columnae stig- [3] EXPLANATION OF ILLUSTRATION PLeuROTHALLIs RUscIFOLIA(Jacqg.) R.Br. 1, leaf and inflorescence natural size, drawn from a Jamaican plant (W. R. Maxon 9486). 2, flower much en- larged. 3, labellum and column much enlarged (an- ther removed). 4, labellum much enlarged. 5, 6 & 7, column much enlarged, to show pulvinate foot, rostellum and stigmatic orifice under the rostel- lum (anther removed). 8, pollinia much enlarged. [4] PLEUROTHALLIS rusectholia matiferis. Anthera terminalis, opercularis, incumbens. Pollinia duo, cerea, subpyriformia. Caules secundarii caespitosi, monophylli, infra folium vaginis anguste in- fundibuliformibus instructis. Pedunculi terminales, ad basin folii solitarii vel fasciculati. Racemi elongati, flori- bus distichis saepe transversis, alternantibus. Lepanthopsis anthoctenium (Peichb,f.) Ames, comb, nov. Pleurothallis anthoctenitum Reichenbach filius in Lin- naea 41 (1876) 94; in Saunders Refug. Bot. 2 (Aug. 1878) sub. t. 118 °° —Cogniaux in Urban Symb. An- till. 6 (1909) 433. This species differs markedly from L. floripecten in the strigose hairs on the tubular sheaths that conceal the secondary stems and in the labellum being nearly as wide as the lateral sepals. In Urban’s Symbolae Antillanae, Cogniaux cites Wright's 1509 under Pleurothallis melanantha Reichb. f. Indeed some of Wright’s collections are mixtures of his numbers 3342 (P. melanantha) and 1509, otherwise it would be difficult to explain Cogniaux’s procedure in confusing two very dissimilar species. In Lindley’s her- barium at Kew, Wright no.1509 is mounted on the same sheet with Pleurothallis floripecten. In the Gray Herba- rium Wright’s nos. 1509 and 3342 are mounted on the same sheet and in this case it is apparent that both no. 1509 and no. 3842 are a mixture of Lepanthopsis mela- nantha and L. anthoctenium. Haiti, Poteau fide Reichb.f. loci cit. Cusa, Prope villam Monte Verde dictum, Cuba Orientale. Jan.- July 1859. C. Wright 1509: Valley of the Rio Bayamita, south slope of the Sierra Maestra, ten meters up on tree trunk, at 900~-1,050 meters altitude. April 5-7, 1907. W. R. Maron 3949. (U.S. Nat. Herb. 522574). qa) ’ . . , For discussion of date see page 82. [7] EXPLANATION OF ILLUSTRATION Srecis Enpresit Reichbf. 1, flower from the type specimen (8). 2, labellum side view. 3, label- lum as seen from above. 4, petal. 5, gynostemium (anther removed) showing lateral stigmas and the triangular rostellum. Srecis Poweiiiy Schltr. 1, flower from the type speci- men ( about 9). 2, labellum as seen from above. 3, labellum oblique side view. 4, gynostemium (anther removed) showing lateral stigmas and lin- guiform rostellum. 5, petal. STeELIs vEsTITA Ames 1, flower from the type speci- men ( about 10). 2, labellum as seen from a- bove. 3, gynostemium (anther removed) showing lateral stigmas and linguiform rostellum. Sreiis NUBIA Ames 1, flower from the type specimen ( about 7). 2, labellum as seen from above. 3, petal. 4,g¢ynostemium (anther removed) showing lateral pulvinate stigmas and linguiform rostellum. [8] STELIS Powellit Schltr. Lepanthopsis densiflora (Rodr.) Ames, comb.nov. Lepanthes densiflora Rodriguez in Vellosia 1, ed. 2 (1891) 119. Pleurothallis congestiflora Cogniaux in Martius FI. Bras. 3, pt. 4 (1896) 591, t. 118, fig. 4. Lepanthopsis densiflora differs from the other species of the genus in having the labellum nearly equal in length with the lateral sepals. The lateral sepals are about 1.5 mm. long, the labellum is 1.25 mm. long. Brazit, As avores dos lugares sombrios e humidos da matta que circunda 0 cume da serra de S. José d’El-Rei, provincia de Minas Geraes. Floresce em Junho e Agosto. Rodriguez. ILtusrration: Reproduced from Martius FI]. Bras. 3, pt. 4 (1896) t. 113, fig. 4 (as Pleurothallis congestiflora Cogn.). Plant, natural size. 3, lateral sepal. 4, petal. 5, labellum. 7a, labellum and gynostemium as seen from the front. 71, labellum and gynostemium side view. Lepanthopsis floripecten (Reichb,f:) Ames, comb. nov. Pleurothallis floripecten Reichenbach filius in Bonpl. 2 (1854) 25; in Walpers Ann. Bot. Syst. 6 (1861) 175;in Saunders Refug. Bot. 2( Aug. 1878) sub. t.118. Lepanthes secunda Rodriguez Gen. & Spec. Orch. Nov. 2 (1882) 70. Pleurothallis unilateralis Cogniaux in Martius F 1. Bras. 8, pt. 4 (1896) 592, t. 122, fig. 2—Ames & Schweinfurth in Gleason in Bull. ‘Torr. Bot. Club 58 (1981) 346. The sketch of the Brazilian plant in Martius’ Flora Brasiliensis represents the leaves as being broader than in the specimens from Venezuela and Honduras, but the structure of the flowers presents nothing tangible on which to establish distinctions. The specimens collected on Mt. Duida have slightly larger flowers than the Bra- zilian and Honduran specimens, but there seems to be nothing more significant on which to rely in attempting [11] separation. In the flowers I have examined the labellum is by no means so cordate as in Reichenbach’s analytical sketches, but it is probable that in the plate prepared for Saunders Refugium Botanicum the tendency toward a cordiform base was overemphasized. Indeed, in the original sketch of the labellum of the Wagener plant, Reichenbach’s out- line of the basal portion indicates uncertainty, because he modified his drawing with supplementary lines. The plate in Saunders Refugium Botanicum shows yellow sepals strongly suffused with mauve along the nerves, petals which are yellowish on the lower half and mauve on the upper half and the labellum yellowish-green with a mauve border. In the dried specimens I have ex- amined, the flowers appear to have been yellowish throughout and in his description of Wagener’s plant Reichenbach described the flowers as being yellowish when dry. However, Saunders’ plant is structurally so similar to the ones from Mt. Duida and from Honduras that it would be unwise to regard them as distinct species until more material has been seen. Reichenbach’s draw- ing and his description of the gynostemium are mislead- ing and should be disregarded. The following description and notes are based on the specimens collected in Honduras by Edwards. Plants 8-12.5 em. tall. Roots glabrous, whitish, fi- brous. Secondary stems ascending, densely caespitose, variable in length, slender, 5-70 mm. long, completely concealed by two to six closely appressed elongated tubu- lar sheaths, monophyllous. Sheaths up to 1.5 cm. long, gradually dilated upward, terminating in an infundibuli- form marginate ostium; margin of the ostium muriculate or very finely hispidulous. Leaf coriaceous, oblong-ellip- tic, marginate, very shortly petiolate, 1.5—-8.7 em. long, up to 9 mm. wide, bidentate at the apex with the mid- [12] LEPANTHOPSIS densiflora (Rodr.) Ames nerve projecting. Peduncles one to four in the axil of the leaf, up to 6 cm. long, filiform, rigid, with several remote abbreviated tubular bracts. Raceme about 1.5 em. long, bearing as many as twenty distichously placed yellow flowers. Bracts of the raceme infundibuliform, scarious, equaling the pedicels of the flowers. Flowers .75 mm. apart, transversely attached, contiguous. Lateral sepals about 8 mm. long, connate to within 1 mm. of the tip, forming an oblong bifid lamina 1.5 mm. wide, conspicu- ously carinate on the exterior surface, acute or subacute, united at base with the dorsal sepal, 1-nerved. Dorsal sepal 2mm. long, 1 mm. wide, ovate, obtuse or subacute. Petals spreading, about .5 mm. long, oblong-elliptic, rounded at the apex, 1—nerved. Labellum 1 mm. long, scarcely 1 mm. wide, elliptical, conspicuously 3-nerved. Column very short, dilated upward; stigmas orbicular, widely separated. Anther papillose. Pollinia two. In the enlarged drawing of the raceme, two flowers near the summit are shown with the dorsal sepal inflexed. This condition seems to accompany the development that follows pollination. In flowers that exhibited this pe- culiarity I found pollen masses adhering to the stigma. The petals had become inflexed in such a way that they concealed the column; the dorsal sepal was closely applied to them, so as to form a protective covering over the pollinated stigmas. In Edwards’ specimens there is a wide range in the size of the plants. The production of the first raceme oc- curs when vegetative development is characterized by very short secondary stems. As the second, third and fourth peduncles appear, vegetative development has pro- gressed rapidly. This phenomenon is clarified by two drawings, one showing a young plant that is flowering for the first time, the other showing an elongated stem which bears four peduncles. [15 ] Repusiic or Honpuras, Department of Comayagua, Minas de Oro. Epiphyte in mountain forest, 4,200 feet altitude. Flowers sulphur color. July 5, 1932. J. B. Edwards 194; Siguatepeque. Epiphyte in open pine forest, 4,000 feet altitude. Flowers pale yellow. September 22, 19382. J. B. Edwards 252. Cotomsia, Reported by Schlechter as probably having been found in North Santanda. Venezupia, near Jaji. Hermann Wagener; Summit of Mount Duida, Epiphyte in woods, 4,400 feet altitude. G. H. H. Tate 825. Brazit, Croissant sur les arbres des foréts qui couvrent les mon- tagnes prés Rodeio et Ala Province du Ceara. Fleurit en Mars. Fr. Allemao 1490 fide Rodriguez loc. cit. ILLUSTRATION: 1, secondary stem (natural size) with four peduncles showing that the stem elongates as the successive peduncles are pro- duced. 2, general habit (natural size) showing that the stems are com- paratively short when the first peduncle is produced. 3, flower much enlarged. 4, gynostemium much enlarged. The heavily stippled areas represent the lateral stigmas. 5, labellum much enlarged. 6, petal much enlarged. 7, anther much enlarged showing the under side. 8, pollinia much enlarged. 9, peduncle and raceme much enlarged. ‘Iwo other species that appear to belong in Lepan- thopsis, namely Pleurothallis melanantha Reichb.f. and P. microlepanthes Griseb., are represented in herbaria by material that is difficult to analyze. P. melanantha was described from fruiting specimens with the perianth or- gans persisting. The gynostemium had suffered changes which obscure the structure that characterized the flow- ers before pollination was effected. P. microlepanthes in its post-pollination phases closely resembles P. melanan- thawhile at anthesis the gynostemium suggests the struc- ture that characterizes Lepanthopsis floripecten. It seems likely that both P. melanantha and P. microlepanthes are in the same alliance formed by L. floripecten and L. an- thoctentum, yet they appear to differ conspicuously from these species in the aspect of the inflorescence. Vegeta- tively and in the plan of the perianth they suggest LL. an- thoctenium very closely, with the dorsal sepal more ex- [16 | LEPANTHOPSIS | Merspecten mes tensively connate with the laterals. In the illustration of Lepanthopsis microlepanthes, fig. 5 clearly shows the post-pollination aspect of the gy- nostemium. The semi-globular protuberances, one with a pollinium attached to its summit, represent the ex- traordinary development that takes place after pollina- tion is effected. This development of the stigmas is also found in L. melanantha and in Pleurothallis ruscifolia, but if my conclusions are justified, the end-result simply masks, in P. ruscifola, the structural peculiarities ante- cedent to pollination. In adding P. melanantha and P. microlepanthes to Lepanthopsis it should be remarked that they differ con- spicuously from JL. floripecten in the nature of the in- florescence. The flowers are not transversely inserted, but are distichous and in two ranks. Lepanthopsis melanantha (Reich).f.) Ames, comb. nov. Pleurothallis floripicta’ Lindley in Mem. Amer. Acad. 8 (1861) 219, nomen tantum. Pleurothallis melanantha Reichenbach filius in Flora 48 (1865) 275—Cogniaux in Urban Symb. Antill. 6 (1909) 430; 8 (1920) 126. It was noted above under Lepanthopsis anthoctenium, that in Urban’s Symbolae Antillanae, Alfred Cogniaux had referred Wright’s Cuban plants distributed under numbers 1509 and 3342 to Pleurothallis melanantha. The only satisfactory explanation of this procedure seems to be that the plants representing these numbers were a mixture or at least that no. 1509 is represented by both species. In the Gray Herbarium where the plants num- bered 1509 and 8842 are mounted on the same herbarium sheet, Lepanthopsis anthoctenium and L. melanantha are represented under both numbers. Although these species [19 ] EXPLANATION OF ILLUSTRATION Lepantuopsts FLORIPECTEN (Reichb.f.) Ames Repro- duced from Martius Fl. Bras. 3, pt. 4(1896) t. 122, fig. 2 (as Pleurothallis unilateralis Cogn.). Plant, natural size. 1, flower. 2, dorsal sepal. 3, lateral sepal. 4, petal. 5, labellum. 6a, gynostemium as seen from the front. 6d, gynostemium as seen from above. 61, gynostemium side view. [ 20 | floripecten (Reichb.f.) Ames LEPANTHOPSIS 5+ 24,) are markedly distinct, they are very similar in the vege- tative structures and difficult to distinguish apart when sterile, unless, as I think is true, the margin of the leaf of L. anthoctenium always has the appearance of being serrate, while the leaf of LL. melanantha has asmooth or unserrated margin. In the specimens I have examined this distinction has been constant. Reichenbach’s description of Pleurothallis melanan- tha is, I think, misleading because in it he characterizes the petals as being ligulate. In his herbarium the sketch that accompanies the Wright plant shows the petals much longer than wide and conspicuously 1-nerved. In the specimens of Wright 3342 (the type number of Plewro- thallis melanantha), that I have examined, I have found the petals to be nearly orbicular and without a conspicu- ous median nerve. Reichenbach described the labellum as being cordiform, acute. In the flowers of the type number that I have examined the labellum is cordiform, obtuse. Perhaps there is much variation in the perianth organs of Lepanthopsis melanantha and my observations may differ from Reichenbach’s because my material rep- resents one of the variants. Thegynostemium of Lepanthopsis melanantha is diffi- cult to interpret because the flowers available for study are old with the subtending capsules about ready to de- hise. Unfortunately this species does not appear to be available in perfect condition. In the flowers of Wright 3342 the gynostemium has terminal stigmas. These stig- mas are very small and form two erect, conical or sub- globose masses which are contiguous through the centre of the clinandrium. It would seem that the generic af- finity is rather with Lepanthopsis than with Pleurothallis. Cuna, 1860-1864. C. Wright 3342. Santo Dominco, Prope Constanza in sylva frondosa 1250 m., Tuerckheim 3089 fide Cogniaux in Urb. Symb. Antill. 8 (1909) 126. [ 23 ] Lepanthopsis microlepanthes (Grisch.) Ames, comb, nov. Pleurothallis microlepanthes Grisebach F 1. Brit. West Ind. (1864) 610—Cogniaux in Urban Symb. Antill. 6 (1909) 480—Fawcett & Rendle Fl. Jam. 1 (1910) 65. My conception of this species is based on the speci- mens in the Herbarium of the New York Botanical Gar- den and on a drawing in my herbarium of the Wilson plant collected in 1857 and preserved at Kew. The gy- nostemium of the Cuban specimens suggests very closely the gynostemium of 1. melanantha. The flowers of Léon, Clement & Roca no. 10871 are described as having been yellow. Lateral sepals 1 mm. long, connate for the greater part of their length, forming an ovate, apically bifid lamina which is only slightly longer than the label- lum. Dorsal sepal very broadly ovate, obtuse, about 1 mm. long. Petals less than 1 mm. long, about half as long as the dorsal sepal is wide, narrowly elliptical, obtuse. Labellum obscurely or very shortly clawed, apparently sessile, orbicular, smooth, ecallose, slightly lessthan 1mm. long. All the perianth organs are tenuous and without conspicuous nerves. Gynostemium minute, slightly di- lated above the middle, the stigmas after pollination be- come much swollen and form erect, terminal, semi-glo- bular protuberances which occupy almost the entire area of the clinandrium. The diminutive gynostemium is very difficult to in- terpret from dried specimens, consequently information regarding the exact position and nature of the stigmas antecedent to pollination will have to await studies of fresh material. That the stigmas are terminal and in con- formity to the generic characters herein emphasized is in- dicated by post-pollination developments. (cf. illustration, fig. 5). That they are lateral and, in the early stages of [ 24 ] LEPANTHOPSIS microlepanthes (Griseb.) Ames a et a of anthesis, confined to the anterior margin of the androcli- nium, as in L. floripecten, seems probable. Cusa, Loma del Gato and vicinity, Cobre Range of Sierra Maestra. In woods at 1,000 meters altitude. July 11-August 14, 1921 Leon, Clement & Roca 10371. (Herb. N.Y. Bot. Gard. ) ; same locality. July 11—August 14, 1921. Leon, Clement & Roca 10548 (Herb. N.Y. Bot. Gard. ). Jamaica, Macfayden: Wilson: March: Mabess River, Harris. ILtusrraTION: 1, from a drawing of the Wilson plant in Herb. Kew., natural size. 2, from a drawing of a leaf and stem of the Wilson plant (enlarged). The ciliations of the sheaths not shown. 3, leaf and stem of the Cuban plant (no. 10371) twice larger than natural size. 4, flower of the Cuban plant (no. 10871) much enlarged. 5, column of the Cuban plant (no. 10548) showing the post-pollination condition of the stigmas, much enlarged, with a pollinium attached to one of the stigmas. 6, pollinia of the Cuban plant (no. 10371) much enlarged, imbedded in the glutinous material of the rostellum. NOTES ON THE FOLLOWING ILLUSTRATION: The petals of L. melanantha as interpreted by Reichenbach are nar- rower and longer than those of Wright’s 3342 (type num- ber) in the Gray Herbarium, and the labellum is acumi- nate rather than rounded at the apex as in the specimens of Wright 3342 in the Gray Herbarium. Furthermore the petals of the Gray Herbarium specimen lack a dis- tinct median nerve. The enlarged drawings of the flowers and the ac- companying analyses are from Reichenbach’s drawings in the Reichenbachian Herbarium. With the exception of g, which is much reduced, all of the analytical draw- ings are practically equal in size to the originals. It may be noted that the sepals in Reichenbach’s drawing of Lepanthopsis floripecten are longer in relation to the labellum than is true of Edwards’ plant. In this regard it is worthy of remark that Lindley’s sketch of a flower of the Wagener plant resembles the flowers of the Edwards plant very closely. Allowance must of course [27] EXPLANATION OF ILLUSTRATION Lepanruopsis FLORIPECTEN (Reichb,f.) Ames 1,plant drawn natural size from type sheet of Pleurothallis Aloripecten Reichb.f. in the Reichenbachian Herba- rium. a, flower from a tracing of the original draw- ing. b, front view of the column. c, posterior view of the column. d, labellum. Lepantuopsis aNruocrentuM (Reichb.f.) Ames 2, plant drawn natural size from a specimen in the Reichenbachian Herbarium. e, column. f, petal. g, flower (drawn somewhat smaller than the origi- nal). LepANTHOopPsIs MELANANTHA (Reichb,f.) Ames 3,plant drawn natural size from the Wright plant, the type of Pleurothallis melanantha Reichb.f. in the Reich- enbachian Herbarium. h, capsule with the persis- tent sepals. i, petals, column and labellum. j, dorsal sepal. k, flower and capsule slightly enlarged. Figures a~k copied from original drawings made by H. G. Reichenbach. [28 ] be made in the consideration of proportions in drawings that were made free-hand and unaided by the use of a camera-lucida. In the drawing of the plant of .. floripecten the mar- gin of the open end of the sheaths does not show the ciliation that is characteristic of the species. Reichenbach described his material as densely fimbriate at the dilated mouths of the sheaths. [31 | SAUNDERS REFUGIUM BOTANICUM VOLUME II BY F. Tracy Hupparp Votume two of Saunders Refugium Botanicum was issued in three parts. The dates of publication are open to doubt and have been a source of confusion. Fortunately Professor Ames’ copy of the second volume contains the original covers and these materially help to establish the dates, but nevertheless fail to remove entirely uncertainty regarding the actual dates of issue. The covers bear the following dates: Part 1, June 1869 contain- ing plates 73 to 96 inclusive; Part 2, October 1872 (corrected in ink to August 1878) containing plates 97 to 120 inclusive; and Part 8, June 1882 containing plates 121 to 144 inclusive. A discussion of the accuracy of these dates seems to be necessary. Part 1. Although the cover and plates are dated June 1869 the first notice regarding Part 1 seems to be in the Journal of Botany, volume 8 (March 1, 1870) 60 which reads, “‘The first part of the second volume of Mr. Wilson Saunders’ “Refugium Botanicum’ has just appeared.’’ From the evidence at hand it would seem that Part 1 did not appear until early in 1870. Part 2. While the plates all bear the date of October 1872 the evidence seems to indicate that Part 2 was not issued until a much later date. In the copy of volume two at the Gray Herbarium there isa paster sent out by the publishers apologizing for the late appearance of this part. Furthermore, in the American Journal of Science and Arts, series 3, volume 8 (1879) 155 a review referring to volume two is in part as follows: ““There is now a second part bearing the date 1878...’’ Consequently, it would seem advisable to accept August 1878 as corrected on the part cover in Professor Ames’ copy as the actual date of publication. Part 3. In spite of the fact that the plates of this part are dated November and December 1872, it seems certain that Part 3 did not appear until a much later date. In Just Jahresbericht for 1882 (volume 10, part 2 (1885) 76) there is a review and abstract of this part which definitely states the date of publication as 1882, which is in accordance with the date (June 1882) given on the cover of Professor Ames” copy. Therefore, on the evidence at hand, it would seem that Part 1 appeared early in 1870; Part 2 in August 1878 and Part 3 in June 1882. [ 32 | BOTANICAL MUSEUM LEAFLETS HARVARD UNIVERSITY CamsBripGe, Massacuuserrs, Ocroper 24, 1933 No. A NEW SOBRALIA FROM THE REPUBLIC OF HONDURAS BY OAKES AMES AmonG the described species of Sobralia,S. /’dwardsiu is most closely allied to 8. crocea Reichb.f., a native of Peru, differing from it in the color of the flowers and in the structure of the labellum. After pollination the ovary elongates excessively, bearing at its summit the swollen column and the withered vestiges of the tenuous peri- anth. This peculiarity, shared with S. crocea, gives the plant a remarkable and characteristic aspect. As the sec- ond flower expands, the capsule underneath the first flow- er, if pollination has been effected, may exceed 6 cm. in length. This condition is represented in the accompany- ing plate. Sobralia Edwardsii Ames, sp. nov. Herba epip- hytica, trifoliata, foliis elliptico-lanceolatis, tridentatis. Flores duo, succedanei. Sepala oblongo-lanceolata, valde apiculata, similia. Petala plus minusve membranacea, oblonga, subacuta, apiculata. Labellum elliptico-oblong- um, margine inaequaliter plicatum et undulatum, basi breviter bicallosum. Discus conspicue carinatus, carinis septem valde sinuatis ornatus. Columna gracilis, leviter alata, apice lobis falcatis ornata. Plant 26-31 em. tall, slender, trifoliate about 2 dm. above the base, with a conduplicate finely asperate sheath [1] 10 subtending each flower; sheaths tridentate at the tip, a- bout 2 em. long. Stems slender, about 3 mm. in diameter, partly concealed by elongated tubular sheaths. Sheaths 2—6.2 em. long, nervose when dry, finely asperate, closely appressed, obtuse or subacute, brown-annulate at the base. Leaves 6-8 em. apart, 7-nerved; lamina 4—14.5 em. long, 1-8.2 em. wide, elliptic-lanceolate, tridentate at the tip, continued below into elongated finely asperate closely appressed tubular sheaths. Inflorescence, with its subtending conduplicate bracts, arising in the axil of the uppermost leaf, 2-flowered, with the flowers opening in succession. As the second flower expands the first one is entirely withered,the remains of the perianth and column surmounting the elongate capsule. Lateral sepals 3.7 cm. long, about 8 mm. wide, pale green, narrowly lanceolate, conspicuously apiculate, lightly carinate on the outer sur- face along the median nerve. Dorsal sepal pale green, equal to the laterals and similar. Petals pure white, about 83.5 cm. long, 6 mm. wide, oblong, subacute, apiculate. Labellum 8 em. long, 1.5 em. wide, elliptic-oblong, ob- tuse, margin above the middle conspicuously folded or plaited in small regular undulations, unequally and in- conspicuously crenulate, but not at all fringed or deeply dentate, shortly bicallose at the base, each callus bifurcate or trifurcate, with one of the divisions extending along the dise almost to the apex of the labellum and becoming conspicuously raised into a sinuate keel above the middle, in addition five of the nerves become similarly carinate, and several others become shortly carinate or bear con- spicuous flattened protuberances near the margin on the distal portion of the disc. Column about 1.6 em. long, lightly winged on the sides, with two falcate erect retrorse terminal lobes or teeth. Capsule slender, elongated, up to 9 em. long when mature. Nm [2] = : ee Oe re oy rer Rerustic or Honvuras, Department of Cortez, El Jaral, Lake Yojoa. Epiphyte in dense forest at 2,000 feet altitude. Sepals pale green, almost white. Petals pure white. Lip white with a light yellow field marked with brown stripes, crinkled like crépe. October 27, 1932. J. B. Edwards 299. (Tyre in Herb. Ames No. 39404.) ILtustRATION: Plant drawn natural size. 1, labellum. 2, petal. 3, anther. 4, column. [5] A NEW BLETIA FROM MEXICO BY OAKES AMES AND CHARLES SCHWEINFURTH AMONG several orchids submitted by the New York Botanical Garden for determination, the following species from the Mexican state of Chiapas appears to be unde- scribed. Its very slender habit serves to distinguish it from all the other members of the genus. Moreover, the small flowers with shallow sinuses between the lobes of the lip differentiate it from the nearest allies, Bletia Pott- siti S.Wats. and B. tuberosa (.) Ames. Bletia tenuifolia Ames & Schweinfurth, sp. nov. Herba gracilis. Cormus subglobosus. Folia quattuor vel plura, angustissime linearia. Scapus folia paulo superans, saepissime subpaniculatus. Flores laxi. Sepalum dorsale lanceolato-ellipticum, acutum. Sepala lateralia oblique elliptico-lanceolata, acuta. Petala oblique elliptica vel ovalia. Labellum in circuitu ovatum, prope apicem tri- lobatum; lobi laterales semiovati; lobus medius suborbi- cularis; discus quinquecarinatus. Columna generis. Roots fibrous, coarse, flexuous, glabrous, longitudi- nally suleate when dry. Rhizome apparently creeping. Corm subglobose, about 1.2 em. long, rugose, shining, covered with the fibrous remains of sheaths. Leaves four to five in a cluster, very narrowly linear, articulated to convolute sheaths which in turn are loosely enveloped by several imbricating nervose scarious leaf-bearing sheaths; blades 15-83.5 cm. long, up to 8 mm. wide, complicate- acuminate, 3-nerved, erect-spreading. Scape up to 46 em. high, somewhat surpassing the leaves, slender, concealed at base by three short imbricating nervose sheaths and above by about three remote appressed tubular sheaths. Rachis commonly subpaniculate with a single basal branch, up to 15 em. long (the apical flowers in our [6] specimens consisting of immature crowded buds). Floral bracts lanceolate, long-acuminate, scarious, the lower- most about 5.5 mm. long. Raceme loose, flowers 1 cm. or more distant. Dorsal sepal lanceolate-elliptic, 1 em. long, 4mm. wide, acute, 5-nerved. Lateral sepals asym- metrically elliptic-lanceolate, about 9 mm. long, 3.8 mm. wide, acute, 5-nerved. Petals asymmetrically | elliptic, about 9 mm. long, 4.5 mm. wide, narrowed to an obtuse or subacute apex, the margins of the upper half irregular, 3-nerved with the side nerves branching. Lip ovate in outline, 3-lobed near the apex, articulated to the short column-foot, 8 mm. long, 6.1 mm. wide, broadly cuneate at base; lateral lobes semiovate, 6.2 mm. long, lobulate at their rounded apex; middle lobe much smaller, ee cular-quadrate, 2.1 mm. long and wide, shallowly retuse and apiculate, the margins strongly undulate and lobu- late; disc traversed by five parallel keels which are highest in the middle of the lip and become undulate near their apex; the three middle keels extend about to the middle of the mid-lobe, the lateral keels stop below the base of the mid-lobe. Column arcuate, 7.5 mm. long (measuring the posterior portion) ; clinandrium 3-lobed. Anther semi- globose, 2-celled, each cell 4-chambered. Mexico, Chiapas, Rio Leche. March 22, 1933. C. D, Mell 2084. (Tyrer in Herb. N.Y. Bot. Gard.) [7]