_Phytologia vol, 14 1966 -67 MissouR! BoTANICAL GARDEN LIBRARY PHYTOLOGIA Designed to expedite botanical publication Vol. 14 November, 1966 No. _ CONTENTS HELLER, A. H., & HAWKES, A. D., Nicaraguan orchid studies —1. MOLDENKE, H. N., pogo materials toward a sogpahiiie =: is the genus Callicarpa. HE Ge ores feat, WA OPN A. L.. Bodk revews: 2a ce ee et Wossour! Botanicw#t NOV 14 1966 GARDEN LIBRARY Published by Harold N. Moldenke and Alma L. Moldenke ( i 303 Parkside Road - Q Plainfield, New Jersey 07060 Price of this number, $1; per volume, $5. 75, th wet : x $6 at close of volume NICARAGUAN ORCHID STUDIES — 1 A.H. Heller and Alex D. Hawkes * The present series of articles will deal with new or otherwise noteworthy orchids of Nicaragua. This is the largest of the Cen- tral American countries, and it has until very recent times been essentially unknown botanically. Even today the casual collector omes up with important additions to the indigenous flora with truly remarkable frequency, in vi rtually all groups of plants, The senior author has resided in Nicaragua for some ten years. During this period, on several occasions with the junior author, he has visited many parts of the republic, paying especial atten- tion to the Orchidaceae. The plants have been transported to the Jardin Heller, at Quinta Monte Fresco, near Managua, where they are cultivated under glass, in lath- houses, or on trees or Special tables outdoors. Studies of specimens bloomi ing, season after season, under these admirable eaittinn# offer materials infinitely superior to those to be encountered in the desiccated or pickled contents of the herbarium. Detailed drawings, color pho- tographs, and critical descriptions are made of each plant which flowers. this study of the Nicaraguan Orchidaceae was commenced about 1956, a total of 139 species and varieties were recorded from s e reader is referred to the series of articles by Hawkes, "An Orchid Collecting Trip to Nicaragua,"' published in The Or- chid Weekly, Volumes 1 and 2, 1959, and reprinted in a revised form in The Orchid Review, Volumes 72 and 73, 1964 - 1965, for pertinent locality data and general information on the floristics of the land. this series, the genera and infra-generic taxa are consider- ed in alphabetical sequence. EPIDENDRUM L., The total number of representatives of the huge aggregation gener- ally considered to comprise the genus Epidendrum L. which occur in Nicaragua is still uncertain. This since many of the collections * Monte Fresco, Box 1869, Managua, Nicaragua, and Editor, The Orchid Weekly, P. O. Box 435, Coconut Grove, Florida 33133, res- pectively. pe 2 FRISGLOGI2 Vol. 1h, no. 1 ‘ by the writers have obviously been sadly mis-identified by the orchidological "experts'' to whom they have been submitted are undergoing further critical study. The precise confines of Epi- dendrum L., sensu lato, are also being taken up by us, and by by several of our colleagues, at this time Until we have our decisions fully Sita ied, the following singularly spectacular ''true'’ Epidendrum has been found and requires establishment as a new species. EPIDENDRUM HAWKESI A.H.Heller, sp.nov. Planta ordine eo gee erecta, basin radicante (non supra). Foliis magnis, rigidis. acemis acute. longe-pedunculatis, us- que ad 60-floris. 2 pg in sectione speciosis. Labellum valde 3-lobis, margine dentatis oe lobum medium cuneato- flabellato, ‘rues retuso; c s 2, alatis, ad apicem conver- gentes, medio lamellum pease sigmoideum ornatum, Fe normally epiphytic, but taking root on sharp overcut gras- sy slopes as a stilt-rooted plant with other customary epiphytes. Stems crowded or borne from the rhizome at some slight distance from one another, simple, rigidly erect, with profuse basal (no ee Sse white, rarely branching roots; stems to 1m tall, up to 6m iam., below concealed by sheaths which are usually ee fle or flushed with dark dull purple, slightly fractiflex in leaf-bear-_ ing upper portion. Leaves 3.75 - 11 cm long, 4 - 4.5 cm broad, ob-, ovate to oblong-elliptic, rather concave, with rounded slightly retuse long, on a peduncle up to 50 cm long, 20 - 60-flowered, the eine vinous-purple, mostly covered oie scarious sheaths; rachis 4 - 6 cm long; a secondary branch customarily appears on each inflores- cence the second season, this with an enlarged base, branching off from base of the original rachis. Floral bracts triangular-ovate, 4 mm long. Pedicellate ovary erect, lightly arcuate, dull vermilion, - 4-5 cm long. Flowers very showy, 3 cm long, 2.5 cm broad, spre@ ding, long-lived, vivid carmine lake with dark yellow calli on lip, th® column~-apex also yellow, the anther-cap apple-green. Sepals and pe tals rather rigidly heavy-textured, acute, nervose with raised nerve® those of the sepals 5; dorsal oblanceolate, 1.5 cm long, 4. 5mm broad, reflexed above middle; laterals oblique, narrowed basally, unevenly spatulate, being expanded or lobulate on inside margin, }- 6 cm long X 4mm broad. Petals 7-nerved, obovate, acute, the mat” gins above middle lightly toothed, 1.5 cm long X 5.5 mm broad ab- | ove middle, ascending. Lip adnate to apex of column, deeply 3-lobed: basally truncate, 8mm long X 1.2 cm broad, the margins irregu- ) i i 1966 Heller & Hawkes, Nicaraguan Orchid Studies 3 tuse to form a bilobed segment with an apicule in the sinus, 4 mm long X 6mm broad. Disc with 2 prominent flap-like calli at base whose apices converge; a prominent erect sigmoid keel extends from base to apex of lip. Column almost straight, with conspicuous flaring angles especially towards apex, clavate, 1 cm long. Capsule long-stalked, to 4 cm long and 1.5 cm thick, with 3 prominent blunt keels, when mature heavily flushed with dark dull vinous-purple. NICARAGUA: Dept. Zelaya: Rama (Roosevelt) Highway, be- yond Rio Mico bridge, frequent stilt-rooted orchid on moist gras- Sy roadside bank, alt. about 400 - 600 feet, 30 March 1966, A. H. Heller & A. D. Hawkes 8344 (Type in Field Museum of Natural History). Epidendrum Hawkesii is an ally of the widespread E. ibaguen- se HBK, but differs from that species, which habitually occurs at far higher elevations, in a number of vital respects. Erect-grow- a rigidly so, it bears roots only basally, not scattered up the viney stems of the ally, which are in our area sprawling or even clambering into adjacent umbrage. The leaves of the novelty are of divergent shape, heavier texture, and larger dimensions. And floral dimensions, color, and shape are conspicuously different. Details of all parts of the flowers— which are extremely nume- rous per inflorescence (up to 60), and particularly the lip with its complex callosities, and the column, are im portan ild, this splendid orchid was first Gein as a solitary pecimen some years ago by the senior author; this occurred as atrue epiphyte, but regrettably no materials were preserved, except for a sketch. The habitat fram which the present type ma- terials were gathered is an exceptionally interest i ing one, that the highway cuts through hillsides at a somewhat higher elevation dant and spectacular when in flower (they would otherwise pass un- noticed), can be considered as almost epiphytic, since the found perched loosely on intricate stilt-root systems on leafy humus on Sheer hillsides along with such other normal epiphytes as a hand- some form of Gongora maculata Ldl., lonopsis utricularioides a ) sal. ,and Polystachya cerea cerea Ldl. Additional orchidaceous ne eig bors are large clumps of Sob Sobralia fragrans Ldl. and an as yet un- identified member of that genus. In this region the original forests in which these Epidendrums doubtless occurred as epiphytes were cut over about the year 1960. It would appear that the new species has admirably made a necessary adaptation to a radical change of environment One nddisieanl specimen of the novelty, growing —— on a bare twisted Bauhinia liana in a hot, humid, palm-aroid jun near La Esperanza (Rio Siquia) was collected in March, 1966, Bok 4 PHYTOLOGIA Vol. 14, now 1 did not survive. And several plants fram the Rfo San Juan, near San Carlos, were found by H. F. Cross, of Managua, who presently has them in his extensive greenhouses there. Epidendrum Hawkesii has been introduced into cultivation in- honor the junior author of these studies, and of the recent Ency- clopaedia of Cultivated echd (Faber, London, December 1965). HELLERIELLA A.D.Hawkes, gen. nov. Among the most exciting of our discoveries in the Nicaraguan o chid flora to date has been a large epiphytic plant which rat represents a genus new to science. HELLERIELLA A.D. Hawkes, gen.nov. Planta epiphytica magna conspicue ramosa. Inflorescentiis racemosis lateralibus terminalibusque. Floris in subtribo ma- jusculis, patentibus. Sepalis petalisquam valde amplioris. Pes columnae valde prominens. Pollinia 4, This new genus is a member of the subtribe Ponerinae. It is | closest in its affinity to Platyglottis L.O. Wms., i Pe endem- — ic to Panama. Helleriella, however, differs ini conspicuous branching habit, lateral as well as terminal ss ciaediiaaiain larger | spreading flowers of different shape with sepals much broader than © e petals, a prominent column-foot, and four pollinia instead of six. The species described below is the sole member of the genus known to date. HELLERIELLA NICARAGUENSIS A.D. Hawkes, sp.nov e epiphytic plant, to 3m long (averaging about 1 m long), the new growth leafy and erect, the old growths branched and pen- dent, leafy only on upper parts of main stem and branches. Roots _ flexuose, fleshy, to 15 cm long and 5mm in diam. Rhizome woody; ' stout, black, up to 8mm in diam., with stems arising at intervals of 2 Stems to 3m long, terete below, compressed above, to 1 cm in diam., woody and bamboo-like, with Ate center, grad- ually deciduous from the base after the first year's growth, when branches up to 30 cm long appear, with co 4 roots at the nodes; base of branches often very slender (1 - 2mm diam.), in- creasing in diam. upwards. Stems and branches concealed by stiff, Striated leaf-sheaths averaging 2 cm long; lower parts of stems whe? exposed often turning black or red-brown. Leaves ligulate, charta- ceous, articulated with leaf-sheaths, the margins recurved, the ba- sal margins somewhat crenulate, sharply carinate, the apex ob- liquely retuse, those of the new growth up to 13 cm long and 4 cm 1966 Heller & Hawkes, Nicaraguan Orchid Studies 5 broad, those of the branches up to 8 cm long and 1.6 cm broad. Inflorescences terminal on new growth and secondary branches, and lateral on old leafless stems and branches; peduncle stout, very abbreviated, occasionally up to 5mm long, with a number of dry scarious bracts at base; rachis flexuose, up to 1.5 cm in length, with up to 6 successive flowers borne about 2.5 mm ap- art. Flowers rather large for the subtribe, up to 3.5 cm in di- am., Spreading, lasting about one week, the sepals and petals greenish to yellowish-white, often with either red or green ner- vation, the lip with 5 red median nerves. Floral bracts ovate, closely appressed, purplish, up to 3mm long. Pedicellate ova- ry up to 1.3 cm long, arcuate, usually purple-striate. Sepals stiff, 5-nerved, with an acute fleshy apex, the basal part a bit fleshy, verruculose outside, the dorsal one 2.1 cm long and 6 mm broad near base, the apex slightly reflexed, the upper half shallowly concave, the lower portion convex; lateral sepals ob- ique, 2.2 cm long and 7mm broad at base, slightly concave, dorsally carinate, subacuminate, forming a prominent mentum with the column-foot. Petals concave, 5-nerved, the median nerve sulcate on rear, ovate-lanceolate, acuminate, 1.8 cm long and 4mm broad below middle, narrowed to an abbreviated 2mm broad basal claw. Lip freely mobile, subpandurate, the basal 1/3 fleshy and at right angles to rest of segments in natu- ral position, very shortly clawed, 1.8 cm long and 9 mm broad above middle, 5 mm broad at constriction, and 6mm broad be- that, the margins of upper half finely serrate or vaguely er- ose, the apex sharply apiculate; disc with a prominent rib exten- ding from base to apex, where it terminates in an apicule, with Column slender, arcuate, 9mm long, with a prominent foot 4 mm long, apically with short lateral teeth and a tridentate tooth at the rear; anther quadrilocular, with 2 pairs of equal, com- pressed, triangular, waxy pollinia, each pair with reflexed fari- naceous broad stipe as long as po NICARAGUA: Dept. Jinotega: cen Jinote ga Rock Quarry, epiphytic on scattered trees in and around pasture, alt. about 4700 feet, December 1959, A. H. satis & A. D. Hawkes 1270 (Type in Field Museum of Natural History). Saint locality, June 1960, A. H. Heller 2684. Helleriella nicaraguensis has been found to date in only one locality in Nicaragua, in the vicinity of the Jinotega Rock Quarry, between Matagalpa and Jinotega. Here it is reasonably frequent, and occurs with an extraordinary wealth of other orchidaceous plants, many of them notable rarities. Its seemingly restricted semination is perhaps explained by the remarkable scarcity of production of seed-capsules. Careful self-pollination of livin specimens at Monte Fresco has thus far failed to set these fruits. 6 PHYTOLOGIA Vol. 14, no. 1 The junior author accompanied Heller when this unusual and attractive orchid was first BOs and it is a distinct per- sonal pleasure to commemorate this active orchidologist's labors, once again, this time in Sy glorious adopted land. LEPANTHES Sw. Of 66 species of this highly technical genus recorded from Mexico and Central America, only one had been authenticated from Nica- ragua prior to the commencement of this study of the indigenous orchid flora. To date, we have found twelve species of Lepanthes our area, including the following charming epiphyte here estab- lished as new to science dditional botanical cloation of Nicaragua will doubtless in- crease the number of these pleurothallids here, since there are 35 species in Costa Rica and 4 in Honduras which have not as yet been detected within our territory of consideration. LEPANTHES HELLERI A.D.Hawkes, sp.no Planta sere ee parva, gracilis. Toliia cehiniacs, forti- ter concavis vel tubulari-cucullatis, margine saepe tangentibus. Floris usque ie 40 succedaneis, minutis. Petalis verrucosis, pa- pillosis. Labellum forcipatum, medio lineato, elongato, ad apicem tuberculatis ciliatis erectis ornato. Plant erect, caespitose, 3 - 7 cm tall, the secondary stems ve~ ry slender, concealed by 4 - 7 tubular, black, somewhat dilated, acuminate sheaths with reflexed margins and verrucose longitudinal ° Leaves orbicular, sometimes broader than long, the lar- gest about 2.1 cm long and 2.2 cm broad, always deeply concave and tubular-cucullate, with margins often touching, green to yel- low-green in color, conspicuously marginate, abruptly contracted to a petiole 3mm long, the apex rounded and minutely retuse, oc~- casionally with an indistinct apicule in the sinus. Inflorescences 1 - 3 per growth, usually shorter than the leaves, about 2 cm long, bearing up to 40 successive flowers | on each distichous raceme. Flo- ral bracts 1mm long, ovate, acuminate, verrucose. Flowers spreading, about 5mm aa the ceaale light greenish-yellow, the lateral lobes of the lip and posterior lobes of the petals ochraceous bordered with magenta, the anterior lobes of the petals ochraceous, remainder of lip and column magenta. Pedicellate ovary filiform, 1.5 mm long, the pedicel persistent. pals elliptic-lanceolate, acute, l-nerved, the dorsal 2 mm broad at middle, the laterals uni- ted to about the middle, 1.5mm broad. Petals transversely bilobed, with a minute apicule in the sinus, the surface and margins minutely but densely papillose, the anterior lobe triangular, subacute, small, 1,1 mm long and 0.5 - 0.6 mm broad, the posterior lobe broadly 1966 Heller & Hawkes, Nicaraguan Orchid Studies 7 oblong with obliquely obtuse apex, 1.5 - 1.8mm long and 0.7 - 0.9mm broad. Lip forcipate, bilobed, about 1mm long, with fleshy verrucose falcate-lanceolate lateral lobes, the apices of which converge around the column-apex, the midlobe a linear wards base, 1.5 - 2mm long; anther large, ovoid, with 2 lin- ear pollinia. NICARAGUA: Dept. Jinotega: Jinotega Rock Quarry, epi- phyte on dry branchlets of small trees, common where found,alt. 4700 feet, August 1964, A, H. Heller 5163 (Type in Field Mu- seum of Natural History). Cerro Volcan Grande, alt. 4000 feet, A. H. Heller 1370. Finca Santa Fé, alt. 3200 feet, A.H, Heller 4529. Dept. Matagalpa: Finca El Roblar, alt. 2650 feet, A. H. Heller 4023. Depts. Matagalpa & Jinotega: Between Cerro Matapalo (Tuma Grade) and Jinotega Rock Quarry, alt. 3200 - 4700 feet, A. H. Heller (& A. D. Hawkes, in part) 1080, 1088, 2539, 2893, 3023, 3078, 3166, 3241, 3695. eee This fascinating Lepanthes, with its jewel-like flowers borne inside the cup-like leaves, is somewhat allied to L. rotundifolia L. O. Wms. of Panama, differing in the foliage, the not malleoli- pillose petals, lip, and column. Lepanthes turialvae Rchb.f., f Guatemala, Honduras, Nicaragua (new record!), and Costa Rica, differs from L. Helleri in elliptic, flattened leaves, an al- most obsolete midlobe of the lip, ciliate margins of the lip's la- teral lobes, and the apex of the anterior lobes of the petals, m (especially Acacia spp.) with Masdevallia chontalensis Rchb.f. or on guavas (Psidium Guajava L. ) with Oncidium crista-galli Rchb.f. Both in the wild and in cultivation, this elfin new Lepanthes is es- sentially ever-blooming. PLEUROTHALLIS R.Br. As of this writing, fifty-three species of Pleurothallis R.Br. have been found in Nicaragua. Until Heller commenced his activtities in the republic, the total of recorded representatives of this fasci- nating and complex genus stood at seven. Five very interesting Pleurothallis fram the Nicaraguan collec- tions are here described as new to science. A forthcoming paper in this series will discuss the records new to the regional flora. 8 PHYTOLOGIA Vol. 14, no. 1 PLEUROTHALLIS ALEXIL A.H.Heller, sp.nov Planta epiphytica, erecta. Foliis linearo-ligulatis, coriaceis, ad apicem tridentatis. Inflorescentiis filiformibus, paulo elonga- tis, 2-floribus. Floris nutantibus, campanulatis. Petalis carno- sissimis, concavis. Labellum ligulatum, sulcatum, ad apicem convexum, verruculosum, ad margine parce glanduloso-pilosum, supra basin auriculae parvae rotundatae ornatae. caespitose, about 5 cm tall, the rhizome creeping, bearing secondary stems at intervals of 1 - 2mm. Secondary stems 1 - 1.5mm long, concealed by scarious sheaths up to mm long which also enclose the petiole. Leaves linear-ligulate, tridentate at apex, coriaceous, including the 1 cm long petiole to about 5 cm long, to 7mm broad at middle. Inflorescence ho- rizontal for half its length, then obliquely erect, the peduncle reddish-green, filiform, 6 - 8 cm long, with three nodes on basal third each with a 4mm appressed sheath, with a fourth some- what inflated sheath 5mm long at middle. Flowers two, borne successively, large for the plant, about 1.5 cm long, nodding, the apical half of sepals dark wine-purple, the lower half stri- ped with white; petals dark wine-purple; lip yellowish with red calli and an ochre apex; column ochraceous, with the anterior margin and foot reddish, the anther-cap white. Pedicellate ovary filiform, arcuate, verruculose above, maroonin color, 8mm long. Sepals connate at base, 3-nerved, the exterior sparsely verrucose, the dorsal one ovate-lanceolate, shortly acuminate, 1.3 cm long and 3.8 mm broad near base, concave there, otherwise convex, the midnerve strongly carinate, the lateral nerves slightly carinate; lateral sepals connate to 3mm from apex, 1.5 cm long and 7 mm broad when expanded, the midnerve ancipitously carinate. Petals very fleshy, concave, the surface sparsely glandular, 3.5mm long and 2mm _ broad above the middle, obliquely expanded upwards on anterior mar- gin, the apex subacute. Lip ligulate, sulcate, 4mm long and broadly rounded and broadly retuse, with an erose apicule on the underside; a rounded auricle is set on each side of the narrowed portion about one-third from base; entire surface finely verru- culose, the margins sparsely glandular-pilose; calli paired, small, basal, pilose. Column conspicuously clavate, broadly winged, 3mm long, with a 2mm long foot, elevated at base of foot and glandular on sides, the apex with three teeth, the pos- terior one broad and denticulate, the lateral ones acuminate. NICARAGUA: Dept. Matagalpa: Cerro Matapalo (Tuma Grade), epiphytic, alt. about 3200 feet, July 1965, A. H. Heller 8631 (Type in Field Museum of Natural History). A pickled specimen is on deposit at the Oakes Ames Orchid Herbarium of Harvard University. 1966 Heller & Hawkes, Nicaraguan Orchid Studies 9 Pleurothallis Alexii ea to have no particularly close allies in the Central American flora. It can be compared with the widespread “ Sei chant? S. Wats., but differs in its droop- ing campanulate flowers, the very fleshy concave petals, and the structure of the lip. It is named to honor the junior author of this series of studies. PLEUROTHALLIS CARNOSILABIA Heller & Hawkes, sp.nov. Planta caespitosa, variabilis, caulis secondariis gracilis, vaginae striatae, inflatae ornatis. Floribus fere duobus. La- bellum late ligulatum, carnosum, longitudine cavae duae papil- losae ornatae, ad basin lobis parvis tenuis porrectis dolabri- formis ornatis. Plant caespitose, variable in size, in bloom from 2.5 to 12 cm tall, the secondary stems slender, 7mm - 4 cm long, con- onaee by two scarious, striated, somewhat inflated sheaths. Leaves eee aca elliptic to oblong-elliptic, coriaceous, 2 - 7.5 cm long including the 5 mm long petiole, 1.2 cm broad at middle, the apex tridentate, with the middle tooth the longest. Inflorescence from base of petiole, arising from a sheath 2 - mm long, lax, up to 2 cm long, spreading, usually 2-flowered, the peduncle flexuose, filiform; old peduncles frequently persist. Floral bracts 3mm long, white, tubular, with an inflated apex, connate from above middle to base, with an apicule at the trian- gular apex. Pedicellate ovary arcuate, 5 - 6mm long, reddish. Flowers about 1.5 cm long, the sepals magenta, white at base; petals green with vivid maroon nerves; lip white, with basal part of callus brownish-red, with three short red carinate nerves on underside; column maroon. Sepals 8mm long, the dorsal join- ed to laterals above base, elliptic-lanceolate, iWacus: con- cave, 4mm broad at middle, with three carinate nerves on back, sparsely verrucose between these; laterals connate almost to the ved, the nerves carinate, broadly ovate-elliptic. Petals some- what oblique and narrowed toward base, 4.5mm long and 2.1 mm broad, spreading, sub-spatulate to obelliptic, with three rominent raised papillose veins on inner surface, the one on anterior margin raised as an erect lobule above the middle, the surface verrucose between the nerves, the apex rounded, cucul- late and muricate. Lip 6.5 mm long and 3.5mm broad above the middle, the lamina broadly liglulate, entire surface except apex scurfy, the apex smooth, rounded, sub-bulbose; unusual longi- tudinal papillose excavations occur on each side of median cal- lus in upper half of lip; the base of the lip bears two thin por- rect dolabriform lobes 2mm long; disc with a verruculose rai- sed callus above base of claw, 1mm broad near base and near 10 PHYTOLOGIA Vol. 14, no. 1 apex, extending down center of lip to within 1mm of tip about, narrowed in middle where it is strongly pilose-verrucose; thic margins of the lip curve inwards to mask some of the excava- tions. Column 2.5mm long, with a foot 1mm long, narrowing upward to a triangular, acute, denticulate apex, the stigmatic cavity large, orbicular, me ; anther erect, vaguely sigmoid and narrow in lateral view, as denticulate apex; pollinia 2, lin- ear, white, erect and parallel to column in natural position. NICARAGUA: Dept. Chontales: Cerro Pistacho, Babilonia district, epiphytic on large mossy tree, alt. 2150 feet, May 1962 (flowering June and July), A. H. Heller 6537 (Type in Oakes eae ‘cechie Herbarium of Harvard University). There are apparently no Central American species of Pleuro- thallis even remotely related to P. carnosilabia. The closest part ally would seem to be the Colombian P. triquetra Schltr., in that lip has somewhat similar longitudinal excavations, but the al taxon differs — in other respects. The basal part of the lip is much like that P. pteroglossa Schltr., also from Colombia. The specific ee is given in allusion to the extremely fle- shy labellum. PLEUROTHALLIS CHONTALENSIS Heller & Hawkes, sp.nov. a parva, caespitosa. Foliis erectis, obovatis, triden- tatis. Pedicellis glandulo-pubescentibus, alae ancipitae promi- nentibus ornatae. Labellum oblongo-ligulatum, adapicem ro- tum obtusumque, ad basin lobulis eerectis parvis denticula- tis ornatis. Plant caespitose, small, to 6 cm tall, the roots slender, flex- uose, white, to 2 cm long, the secondary stems 5 - 7 mm long, concealed by white scarious sheaths. Leaves subcoriaceous, er- ect, obovate, tridentate but not oblique at apex, 3 - 4.5 cm long including the 5 - 10 mm long petiole, 0.8 - 1.3 cm broad above middle. Inflorescence solitary from base of apace erect, the peduncle filiform, to 4 cm long, glandular, with s as sheath, usually bearing a single flower (this with ee sepals rmost in most instances) with an abortive attendant bud; on mm long. Flowers not opening fully, about 1.5 cm long, ochra- ceous greenish-yellow, the petals and lip-callus orange. Pedicel- late ovary glandular-pubescent, the ovary triangular with 3 high ancipitous wings and a lower one between each, the ovary 3mm long, borne at right angles to persistent 1 cm long pedicel. Se- pals ovate-lanceolate, acute, 1 cm long, 3-nerved, the surface somewhat r, the margins ciliate, the keeled nerves cil- , the median one of the dorsal sepal ancipitous, praninent, min segment 3mm broad below middle, its inner surface with 1966 Heller & Hawkes, Nicaraguan Orchid Studies a conspicuous bullate or ocellate spots; lateral sepals joined to within about 2mm from apex, 4mm broad above middle. Pet- als thin-textured, spatulate, with wide lobe on anterior margin, the apex falcately acute, narrowed basally, 2-nerved, 3mm long and 1.1mm broad above middle. Lip parallel to column, oblong- ligulate, with a rounded obtuse apex, broadest at base, 3.5 mm long and 1 mm broad near apex, fleshy, verruculose, sulcate, the apex more fleshy, margins ciliate, with an erect lobule on each side 1 mm from base; callus elevated, bifid, papillose, exten- ding from lip-base to lateral lobules. Column slender, clavate, winged, 2.5mm long, apically with 2 acuminate lateral teeth and a broad denticulate posterior one, the broadly elliptic, alate foot 1.2 mm long, at right angles to column; anther helmet- shaped, apiculate. CARAGUA: Dept. Chontales: Rio Mico, epiphytic, alt. 1500 feet, August 1960, A, H. Heller 3735 (Type in Oakes Ames Orchid Herbarium of Harvard University). Same locality, alt. 1400 feet, in bloom December, A. H. Heller 3575. Same lo- cality, alt. 1400 feet, in bloom May and June, A. H. Heller 7827. Cerro El Chamarro, La Libertad district, epiphytic, alt. 2175 feet, June, A. H. Heller 1036. Pleurothallis chontalensis belongs to the P. glandulosa Ames lian P. glandulosa Ames, P. pertenuis C.Schweinf., and P. vitarii- folia Schltr. iP closest ally appears to be the Costa Rican P. barbae Schltr., from which it differs in the following respects: broad, not linear acer, although the pedicel is glandular-pu bescent, the ovary is only sparsely so, and has very Siedialinsut ancipitous wings; the floral bracts are densely glandular and ac- ute, not apiculate; the inner surface of the dorsal sepal has bul- late or ocellate spots of some prominence, these not noted in P. barbae; the lip is of divergent shape, and the two lateral lobules are very small and denticulate; the column-foot lies at right an- gles to the column, is broadly suborbicular, separated from the column by a constriction or isthmus, and is eeply excavated with wing-like margins; and the lateral teeth at the apex of the column are acuminate, while the broad triangular posterior one is lace- rate. This pretty novelty is named for the spectacular Nicaraguan department in which it occurs with some frequency. PLEUROTHALLIS EXESILABIA Heller & Hawkes, sp.n Planta caespitosa, parva. Foliis crassis, shbaboak 5 tri- dentatis. Inflorescentia singula, filiformis, 3- 4-floribus succe- daneis producens. Labellum panduratum, exesum, inflatum, fo- ramen in medio discum occuparum. 12 PET TOLOGIS Vol. 14, no. 1 Plant caespitose, small, in bloom to 4 cm tall, the secondary stems almost obsolete, 3mm long, concealed by scarious white sheaths. Leaves thickly coriaceous, 1.2 cm long, including a petiole of about 2mm, 3.5 mm broad above the middle, oblance- olate, tridentate at the apex. Inflorescence pes to 4 cm in length, the peduncle filiform, reddish, bearing a solitary infun- dibuliform nract near middle and 3 or 4 foward. these opening successively. Floral bracts tubular, 1mm long. Flowers open- ing well, about 1 cm long, the sepals and petals pale greenish - py ibe with reddish longitudinal veins, the lip reddish-brown th pale greenish-yellow basal portion, the column greenish- tie. reddish between wings on anterior surface. Pedicels persistent, the pedicellate ovary reddish, 4mm long. Sepals lanceolate, acute to subacuminate, with strongly serrated mar- gins, 6mm long, the dorsal 1.5mm broad, with 3 carinate nerves, the laterals free almost to base, lanceolate, vaguely falcate, with 2 carinate nerves, 1.8 mm broad below the middle. Petals ovate, long-acuminate, concave, sparsely toothed above middle, l-nerved, 3mm long and 1 mm broad. ip pandurate, Fy 5mm long and 0.8 mm broad, hollow and inflated, the open- ing at center of disc, with 2 small teeth on claw and a large and broad falcate porrect lobule on each side at base of lamina, these lobules with incurved apices; callus raised, smooth, extending from claw to middle of lip, narrowly retuse at base; above the opening at the middle of the lip there is a slight swelling, but up from there to apex the lip is concave; apex of lip rounded, blunt, with long pilose hairs on margins and underside. Column 1.9 mm long, slender, somewhat arcuate, shallowly winged, with an entire apex; anther agi _ovoid, contracted to a truncate apical neck, the 2 pollinia NICARAGUA: Dept. acts cs: near Finca Santa Elena, Ma- ge - eg Highway, on fallen branch with Pleurothallis faegil alt. 4510 feet, in bloom July 1962, A. H, Heller 6497 ee in 7 Ames Orchid Herbarium of Harvard Univ- ersity). In habit, structure of the sepals and petals, as well as gen- eral shape of the lip, this remarkable novelty is reminiscent of Pleurothallis samacensis Ames, of Guatemala and Costa Rica. It differs, however, in the hollow lip, this with a small orbicu- lar opening at the center of the disc. The apical portion of the labellum, further, bears long hairs which are lacking in P. sa- macensis. . The specific name is derived from the Latin, exesus, hollow- ed out, and labium, lip or laebellum. J PLEUROTHALLIS HELLERI A.D.Hawkes, sp.nov. Planta pro genere multo procera, erecta. Foliis subcoria- ceis, nervosis, ellipticis, ad basin apicemque contractis, mag- 1966 Heller & Hawkes, Nicaraguan Orchid Studies 13 nis. Inflorescentiis pluribus, flexuosis, gracilis, multifloris. Floris patentibus. Labellum carnosum, verrucosum, dio 3-lobatum, lobum medium oblongo-ovatum, obtusum, ad ba- sin callis dette elevatis mammillatis ornatum. Plant large for the genus, to 50 cm tall, the rhizome abbre- viated, stout, woody, set with scarious fibers and flexuose white roots 1mm in diameter. Secondary stems erect, borne at inter- vals about 1 cm apart, terete, glabrous, to 35 cm long, 2 - mm in diameter, with a striated sheath 1.5 cm long at base and anathise one 4 cm long near middle. Leaves obliquely spreading, subcoriaceous, nervose, elliptic, aivuptly contracted toa sub- petiole 1.5 cm long, up to 20 cm long and to 6 cm broad at mid- dle. Inflorescences 2 to 6 straggly, flexuose racemes to cm long, arising from a 4mm keeled spathe, arching to sub- -pendent, the peduncle slender, 1 - 2 cm long, the rachis bearing 10 to 20 flowers about 1 cm apart. Floral bracts triangular-ovate, infla- ted, 1.5mm long. Flowers spreading, about 2 cm long when ex- lip and its callus. Pedicel persistent, filiform, 3.5 mm long, the ovary 2.5mm long. Sepals lanceolate, long-acuminate, setace- ous, dorsally carinate, 1 cm long, the dorsal 2.6 mm broad near ovate base, 3-nerved, the laterals connate to the narrowly trun- cate apex, 4-nerved. etals narrowly linear, attenuate, seta- ceous, l-nerved, 9mm long and 0.6 mm broad at base, spread- ing. Lip fleshy, 2mm long and 0.7 mm broad, slightly arcuate in natural position, verrucose, with a short membranaceous cu- cullate claw, 3-lobed below middle with thin, erect, rounded la- teral lobes which are difficult to spread out, the midlobe oblong- ovate, obtuse; disc with an elevated mammillate callus along the base of each lateral lobe. Column short, 1.5 mm long, stout, clavate, finely papillose especially on the prominent rounded col- umn-foot; anther subquadrate, with 2 verruculose pollinia. NICARAGUA: Dept. Chontales: Cordillera Amerrisque, ep- iphyte, alt. 2700 feet, December 1962, A. H. Heller 3784 (Type in Field Museum of Natural History; isotype in Oakes Ames Or- hid Herbarium of Harvard University). Rfo Mico, alt. 1150 feet, 2 ter 1963, A. H. Heller 7870. As yet, this singularly attractive Pleurothallis is known o from two localities in Nicaragua's Departamento de Chontales. It may, however, be more widespread here, since its habit is superficially much like that of the common and widespread P. gelida Ldl., and its distinctiveness would not have been noticed unless specimens were encountered in bloom. Pleuro othallis Hel- leri is distantly allied to P. ruscifolia R.Br., distributed from Cuba and Guatemala to northern South America. The inflores- cences of the latter, however, are. glomerate or fasciculate, with 1h raerTtoprec6c Tra Vol. 1h, no. 1 obsolete peduncles, while in the present novelty up to twenty flo- wers appear on each of the multiple rather elongate racemes. Floral structure, too, of P. Helleri, particularly details of the lip, differs from all other members of the genus with which we are familiar. The new species is named to honor the senior author of this serial, A. H. Heller, who has immeasurably increased our know- ledge of the orchids of his adopted land, Nicaragua. SOBRALIA R. & P. Of the 35 Sobralias recorded from Mexico and Central America, only three had been authenticated from Nicaragua prior to the present study; eight species had been reported in the past, north of Costa Ric Thirteen fapresciitatives of Sobralia R. & P. are now known from Nicaragua, and the immense colonies of these spectacular tical studies continue. Four new species are herewith described. se re rl CHATOENSIS Heller & Hawkes, sp.n epiphytica, brevis. Foliis rigidis, sian nervosis. rie cate ube: speciosis. Labellum ellipticum, ad basin tubulosum, parve trilobatum, crenulatum, disco fossa depressa ornato. Plant epiphytic, to 30 cm tall, profusely rooting. Stems lea- fy, with leaves spaced 3 - 4 cm apart, the leaf-sheaths spotted with red. Leaves rather stiff and rigid, to 16 cm long and 5 cm broad, with 9 major nerves, the median one of which has a dorsal keel, on each side of which are 2 minor accessory nerves. Flowers showy, short-lived, spreading, about 10 cm across in na- tural expansion, the sepals and petals magenta, lighter towards base; apical margins of lip richmagenta, the throat and disc ochreous-brown, the sulcate portion holding median nerves gold- ochre, the basal calli rich yellow; column yellow on anterior face, ~ with purplish apical teeth. Sepals 7 cm long and 2.2 cm _ broad, oblanceolate, acute, shortly clawed. Petals 7 cm long and 2 cm broad above middle, oblanceolate, with slender claw, obscurely crenulate on apical margins. Lip elliptic, forming a firm tube clasping the column, 7 cm long Ga 5 cm broad when spread out, obscurely trilobate, the apical margins crenulate; disc with a flat depressed channel 5 mm broad down the middle which encompas~- ses 5 raised median nerves which extend somewhat beyond middle of lip; there are no tubercles between the basal calli, which mea~ sure 1.5 cm in length. Column 4.5 cm long, clavate only near the apex. NICARAGUA: Dept. Boaco: Cerro Chato, epiphytic, alt. 2500 feet, January 1964, A. H. Heller 9553 (Type in Field Museum of Natural History). 1966 Heller & Hawkes, Nicaraguan Orchid Studies 15 Sobralias on Cerro Chato are very rare, and this sole collec- tion appears amply distinct from all others known to us from Cen- tral America. The median channel on the lip is especially unique. SOBRALIA HAWKESI A.H. Heller, sp.nov. Planta speciosa, caulis parce maculatis, gracilis. Foliis el- ee -lanceolatis, magnis. Floribus profusis, succedaneis, spe- osis. abellum ad basin tubulosum, ad sino apicem 5 nervis aes elevatis pustulatis ornato. Plant epiphytic, erect to spreading, up to 15 dm tall, pro- fusely rooting, the roots forming sizeable mounds. Stems slen- der, 6 - 8mm in diameter, slightly maculate, the leaf-sheaths maculate and verrucose. Leaves elliptic-lanceolate, acuminate, 1l-nerved, up to 20 cm long and 7 cm broad, light green, with prominetnt nea Flowers produced successively almost through- out the year, lasting rather well for the genus, to 15 cm in diam- eter, the sepals, petals, and apical margins of the lip rich lus- ous magenta-red, basal half of throat yellow, light lilac to- wards lateral margins, with a brown-ochre band between the yel- ow of throat and magenta-red lip-apex, the lateral teeth of the column purple. Sepals 7.5 cm long, apiculate, with 9 main nerves, the dorsal oblanceolate, 2.1 cm broad, the aes elliptic-lan- ceolate, 2.5 cm broad. Petals elliptic-spatulat with a short narrow claw, 7.5 cm long and 6 cm broad above sues the diverging nerves with an intricate network of short chateaus trans - verse veins, margins crenulate in upper third, the apex subtrun- cate with a small triangular apicule. Lip tubular basally, 7.6 cm long and 6 cm broad when spread out, cuneate-obovate, somewhat constricted on crisped-undulate margins 1.5 cm from apex to form a — deeply retuse apical lobule; disc with two yellow calli at base which are 1.5 cm long, and 3 median raised nerves which saa from base for three-quarters of entire length of the lip; a lateral nerve is present on each side, extending to the cen- ter of the lip, and a shorter nerve branches off from the base of each of these laterals; 5 mm from the apex of the three median nerves occur five somewhat blistered radiating nerves 5mm in length. Column 3.5 cm long, triangular in cross-section, witha thin spreading wing on each side of the anterior face. ARAGUA: Dept. Granada: Volc4n Mombacho, frequent i 3950 feet, January 1960, A. H. Heller & A. D. Hawkes 3025 (Type in Field Museum of Natural History). Same locality, A. H. Hel- ler 7728. This strikingly beautiful orchid is a frequent inhabitant of the marvelous forests of Mombacho Volcano, haunt of an extraordina- ry array of differentk kinds of Sobralias. In addition to the present 16 PHYTOLOGIA Vol. 1h, no. 1 S. Hawkesii, at least three others appear sufficiently distinct to warrant erection as new species; these will be discussed at a la- ter date in this serial, upon flowering of additional living mater- ials, and the complex forms which appear to be natural hybrids betwixt these populations will as well be considered. Sobralia Hawkesii has no immediate allies with which we are familiar. It is related, obviously, to the as yet unnamed collec- tion A. H. Heller 9132, also from Volc4n Mombacho, but the light green (not dark blackish-green) leaves are more lanceolate and less stiff, the maculations on the leaf-sheaths are m less pronounced, floral form and dimensions are radically different, and the five short, raised, blistered nerves near the sinus of the lip are as well distinctive. It is a species of considerable horti- cultural potential, one which flowers profusely almost throughout the year at Jardfn Heller, Quinta Monte Fresco, altitude about 1800 feet. SOBRALIA HELLERI A.D.Hawkes, sp.n Planta epiphytica, speciosa, erecta. Senin robustis, flexu- osis. Foliis plus minusve in plano singulo disponentibus, rigi- dis, fortiter plicatis, magnis. Floribus speciosis, non paten- tibus, singularis. Sepalis ad basin connatis. Labellum expan- sum ellipticum, prope saccatum, margine prope integrum Plant epiphytic, tall, tomore thanlm, erect-spreading. Stems stout and stiff, somewhat flexuose, up to 6 mm in diame- ter, smooth olive-green with distinct red-brown maculations on the leaf-sheaths. Leaves more or less in one plane, green, pli- cate, fairly stiff, borne 3 - 4 cm apart, the largest ones to 23 cm long and 8 cm broad, elliptic-lanceolate, acuminate, with 9 sharp~ ly carinate nerves, sulcate above; the laminas of the leaves over~- lap one another to a considerable extent. Flowers borne succes-~- sively and frequently over a space of several months, lasting lon- ger than usual for this genus, about 6.5 cm long, not widely expan~ ding due to basally connate sepals, which are pure white; petals pure white; lip on inner surface rich blood-red, outer surface white except for red median nerve; column-face red. Floral bracts several, 3 - 4cm long, forming an inflated, lanceolate, acuminate, rigidly erect sheath-like structure to 8 cm long. Pe- dicellate ovary slender, decurved, 3 cm long. Sepals soft-fleshy, 7-nerved, elliptic-lanceolate, apiculate, concave, 6.5 cm long, connate for 1 - 2 cm at base, the dorsal one 2 cm broad, the late- rals oblique, 2.3 cm broad. Petals 7-nerved, the three median ones close together and raised, obliquely elliptic-lanceolate, api- culate, 5.5 cm long and 2 cm broad, the median nerve sulcate un~ derneath, the — apical margin recurved, acts 7 cm long and 4.5 cm broad, cuneate in natural psosition, candle elliptic when spread out, deeply conical to almost saccate, apical- a El 1966 Heller & Hawkes, Nicaraguan Orchid Studies 17 ly retuse with a convex bulge in the sinus, the margins entire ex- hase for indistinct serrations on extreme apical part, nerves nu- rous, indistinct, radiating, none of them elevated; with two cha erose papillose calli 2 cm long at base, these terminating in two to three very short ancipitous teeth. Column disproportionately elongate, 4 cm in length, slender, clavellate, 8mm broad below apex, 2.5 mm broad at base. NICARAGUA: Dept. Rivas: Volcan Maderas, Isla de Ometepe, Lago de Nicaragua, epiphyte in mossy rain forest, alt. 4000 feet, May 1963, A. H. Heller 8029 (Type in Field Museum of Natural History). Sobralia Helleri appears totally distinct fram all other des- cribed members of the genus. In the wild plants, at least, the leaves lie distichously in very much the same plane, with one conspicuously overlapping the other; in cultivation, this condi- tion seems to vary to some degree. Though the flowers do not expand fully, because of the basal connation of the sepals, the pure white color of the outer segments beautifully sets off the in- credible hue of the lip, a startling rich blood-red which is un- known elsewhere in the Orchidaceae, at least to this writer. The labellum, with its entire, unfrilled or non-crenulate margins, pos- sesses a bulging, concave medial portion which is very fleshy and difficult to spread apart; some 5 mm inside the margins the lami- na thins in texture and furls under towards the apex. There are no raised nerves nor lamellae, except the basal elevated calli. The white underside of the lip sports a striking red median nerve. This is, in the opinion of the junior author, one of the most ex- citing of Heller's Nicaraguan discoveries to date. It is an oe, a beautiful Sobralia, one which Pea RAE 5 attracts the atten- n of the viewer, and its ease of culture (at pciicider ably: eae pepe than its cloud-wreathed ittine Siastiat) and profuseness of bloom at once recommend it to the connoisseur collector values these remarkable orchids. It is indeed to be hoped that the fascinating Sobralia Helleri will ere long be made available, to selected orchidists, outside of its isolated, volcanic, insular home. appears that this new species is a narrow endemic to Vol- can Maderas, on the island of Ometepe in Lake Nicaragua; its twin peak, Volcan Concepcion, the taller of the pair, is persistently ac- tive. As of this writing, it is the sole species of its genus which has been discovered on the island, an area which has already pro- duced some remarkable new genera and species for the Nicaraguan flora and for science (see Heller's article, ‘Notes fram Nicara- a: Houlletia tigrina,"' in Amer. Orch. Soc. Bull. 35: 213 - 214. March 1966. ). SOBRALIA TRIANDRA Heller & Hawkes, sp.n Planta epiphytica, rigida, erecta. Foliis iiglle lai eckaul patentibus. Floribus plerumque 2, patentibus, triandris. Label- 18 PHYTOLOGIA Vol. 14, no. 1 a lum 7-nervosis, callis obsoletis. Plant epiphytic, up to 1m tall. Stems stiff, erect, yellow-green, slightly compressed, to 5mm in diameter, the upper part with about 7 leaves, lower part concealed by dry leaf-sheaths, these like the stem with minute brown maculations. Leaves elliptic-lanceolate, yellow-green, spreading, to 13 cm long and 5 cm broad, borne 4a 5 cm apart, fairly stiff, subcoriaceous, the upper surface smooth, set with 9 nerves that are sulcate on the underside, abruptly Pedicellate ovary clavate, to 3cm long. Flowers usually 2 borne successively, widely expanding, tomore than 10.5 cm across, the sepals and petals white often tinged with lavender; apical part of lip blue-lavender to violet-blue, the raised ner- ves in the tubular throat rich yellow. Sepals linear-lanceolate, acute, the dorsal one 6.5 cm long and 1/2 cm broad, the later- als oblique, 6.2 cm long and 1.5 cm broad above the middle. Petals oblanceolate, acute, with serrated margins in upper half, 5.8 cm long and 1.8 cm broad above the middle. Lip tightly tu- bular, with flaring apex, 3-lobed, obovate when spread out, con~ volute at base, 6 cm long and 4 cm broad, the apical lobe deeply retuse and with a blunt rounded apicule in the sinus, the apical margins crenulate and erose to deeply lacerate; disc with a flat sulcate smooth callus commencing at 1 cm from the base, this giving rise to 7 slightly elevated nerves which become more el- evated and pronounced to beyond the middle of the lip, where they become sinuate, crenulate, and verrucose; there are often 1 or 2 lateral, eige ee elevated nerves as accessories; basal lateral calli and fovea obsolete. Column 3 cm long, clavate, slightly alate from top to bottom, the apex with a broad forward- projecting posterior tooth; clinandrium bears a shallow, serra- ted lobe on each side; anthers invariably 3; sometimes the la- teral anthers have four divisions and on other occasions, two. NICARAGUA: Dept. Jinotega: Macizo de Petas Blancas, phe PS alt. 2500 feet, April 1961, A. H. Heller 7260 (Type Field Museum of Natural History). Finca Santa Fé, alt. 3200 ar June 1960, A, H. Heller 4571. Dept. Chontales: Cerro Pistacho, alt. 2150 feet, February 1961, A. H. Heller 6879. Dept. Matagalpa: Finca La Isla, alt. 3800 feet, March, A. H. Heller 8964. In the seven raised nerves on the lip-disc, this species is a bit reminscent of Sobralia Bouchei A. & S., of Panama. But it differs from that taxon in being consistently triandrous (apparent- ly an unique condition in the genus), in that the raised nerves do not extend to the basal part of the lip, and in the naked disc there. Further, the flowers are of a different color, with the apical portion of the lip blue-lavender to violet-blue, a distinct- 1966 Heller & Hawkes, Nicaraguan Orchid Studies 19 ive hue for a Sobralia. When the first collections of this extraordinary plant were made, the senior author of course thought that the three anthers were merely an aberration. But Sobralia triandra has now been found in widely separated localities in Nicaragua, and the trio of fertile anthers is invariably present narrowness of the slender, tubular lip might not admit the lar- ger usual pollinators for similar appearing large flowered Sob- ralias. VANILLA Sw. As of this writing, five species of the genus Vanilla Sw. have been found in Nicaragua. Vanilla planifolia Andr. and V, Pompona Schiede have previously been recorded from the area, as noted by L. O. Williams in his Enumeration published in 1956 (Ceiba, Vol- ume 5). Critical exploration by A. H, Heller has brought to light Van- illa Pfaviana Rchb.f. , previously known from Mexico, British Hon- duras, Guatemala, and Costa Rica; V. odorata Presl, previous- ly known only from Ecuador and Bolivia; and an unusual plant, new to science, which is here described in his honor. VANILLA HELLERI A.D. Hawkes, sp.nov. Planta scandens, elongata, caulis flexuosis, subquadratis. Foliis carnosis, oblongo-ellipticis, ad apicem abrupte acumina- tis contractis. Floribus pro genere parvis, sub-patentibus. La- bellum fortiter 3-lobis, lobo medio appendicis elongatis retrorsis complanatis ornato. Columna gracilis, sub-clavellata, infra stig- mato pilis gracilis glandulosis ornatis. Plant scandent, elongate, the rooting stems flexuose, sub- quadrate, sulcate on two sides, 1 cm in diameter, dark olive- green. Leaves 1 - 14 cm apart, fleshy, 12 - 14 cm long, 3.5 - 4.5 cm broad, oblong-elliptic, abruptly contracted to an acumi- nate apex 1 cm long, below abruptly contracted to a sulcate pet- iole 1 cm long. Inflorescence 10 cm long, with an almost obso- lete peduncle, subumbellate, bearing a succession of up to 20 flo- wers. Flowers small for the genus, about 6 cm in diameter, semi- spreading, short-lived, the sepals whitish with underside of upper half greenish, the petals whitish, the lip bright yellow with orange appendages on the midlobe, the column white. Floral bracts trian- 20 PERT -TOLD 0G TA Vol. 1), no. 1 of the ovary green. Sepals 4 cm long, slightly concave, with 7 - 9 nerves, the dorsal oblanceolate, 1.4 cm broad above the middle, the laterals 1.4 cm broad at middle, oblong-elliptic, oblique, sub- obtuse. Petals 4 cm long and 1.3 cm broad, oblong-elliptic, ob- tuse, set with thick 3-nerved median carinae on both surfaces. Lip strongly 3-lobed, 4 cm long and 3 cm across the lateral lobes when spread out , adnate to base of column and without a claw; midlobe densely covered with retrorse complanate elongated appendages ; lateral lobes multi-nerved, with entire margins; a penicillate crest is located in the exact center of the lip, and from this point to the base there is a slightly raised verruciform callus which be- comes rather dense and more tuberculate at the base. Column slender, rather straight, sub-clavellate, with fine glandular hairs extending for a short distance just below the stigmatic orifice. NICARAGUA: Dept. Chontales: 2 miles south of La Liber- tad on the road to Santo Tomas, growing as a vine on a wild ayo- cado (Persea sp.) tree, alt. 1900 feet, April 1962, A. H. Heller 7946 (Type in Oakes Ames Orchid Herbarium of Harvard Uni- versity). his interesting, small-flowered Vanilla has no close Central American allies. The structure of the lip in particular is most distinctive. 1966 Heller & Hawkes, Nicaraguan Orchid Studies 21 All plates by A. H. Heller Plate 1. Epidendrum Hawkesii A.H.Heller, sp.nov. Left, habit of flowering plant and inflorescence, with semi-ma- ture seed-capsule. Upper right, flower, approx. natural size. Center, seed, considerably enlarged. Center below, sepals and petal. Lower center, column and lip in cross-section. Bottom, largest leaf. Center right, lip enlarged. Lower right, anther, two views, enlarged. Plate 2. Helleriella nicaraguensis A.D. reggie: gen. et sp.nov. Left, second year growth of flowering plant. Bottom left, anther e sg growth of plant. Top right, column and foot, in cross-section. Top center, oblique face-view of column. Center right, column and lip in lateral view. Plate 3. Lepanthes Helleri A.D. Hawkes, sp.nov. Upper left, habit of flowering plant. Lower left, leaf expanded, with stem-apex. Center left top, flower, enlarged considerably. Center left bottom, petal and lip, lateral view. Center top, pet- al, enlarged. Right top, lip expanded and lip with column. Cen- ter right, pollinia and anther. Bott right center, late view 1. of lip, enlarged. Bottom right, stem with dilated costate sheath. Plate 4. Pleurothallis Aléxii A.H.Heller, sp.nov. Top left, flowering plant. Top center, lateral and top view of flower. Top right, dorsal and lateral sepals. Center left, base of plant, showing attachment of inflorescence, Center, column and lip. Center right, anther-cap. Bottom left, column, front face. Bottom center left, lip, top view. Bottom center right, petal, frontal view. Bottom right, petal in cross~section. Plate 5. Pleurothallis carnosilabia Heller & Hawkes, sp.nov. Top left, flowering plant. Bottom left, anther. Top center left, floral bract. Below this, petal. Below this, flower. Below this, lip in natural position. Center top, flower, showing sepals and lip. Center right, column in lateral cross-section. Center om, two sections through lip. Top right, column and lip in natural position. Petal sectioned, Column seen in lateral sec- tion and in frontal view. Section of lip, column, and lateral se- pals. Underside of lip-base, showing its carinae. Center right, section through petal. Bottom right, basal lobes of lip expanded. 22 PHYTOLOGIA Vol. 1h, no. 1 Plate 6. Pleurothallis chontalensis Heller & Hawkes, sp.nov. Top left, flowering plant. Center left, column in face view. Cen- ter right, lip. Right, section through dorsal sepal, plus dissec- ion Ower, except lip. Left center, pedicellate ovary, column, and lip, in lateral view. Right center, section through lateral syn- sepal and this segment. Bottom left, section through pedicellate ovary. Bottom left center, pollinia and anther. Bottom right cen- ter, apex of leaf. Plate 7. Pleurothallis exesilabia Heller & Hawkes, sp.nov. Top left, flowering plant. Top center, longitudinal section of a leaf. Right, flower, dissected. Center left, lateral view of col- umn and lip, and lateral cross-section of lip, showing cavity. Bottom left, column with basal callus, next basal auricle, nex front of column, and to right, underside of lip. Bottom center, anther-cap and pollinia. Plate 8. Pleurothallis Helleri A.D. Hawkes, sp.nov. Left, flowering plant. Top right, flower, dissected, Lower cen- ter right, lip expanded, plus lateral view of lip and same in cross- section. Bottom right, lateral view of column, frontal view of the column, and anther-cap with pollinia below. Plate 9. Sobralia chatoensis Heller & Hawkes, sp.nov. Left, leaf, with portion of leaf-sheath. Flower, dissected. Bot- tom center, section across lip-disc. Center right, lateral and frontal views of column Plate 10. Sobralia Hawkesii A.H. Heller, sp.nov. Dissected flower on outline of average leaf. Bottom left, lip in natural lateral position. Center right, column with apex of ped- icellate ovary in lateral position, and column in frontal view. Bottom right, cross-section through column. Plate 11. Sobralia Helleri A. D. Hawkes, sp.nov. Flowering plant, showing apical sheathing bract-system. Bottom left, underside of lip, and anther. Center right, lip expanded, plus sepals and petal. Bottom right, lip in lateral view, and col- umn in lateral view. Plate 12. Sobralia triandra Heller & Hawkes, sp.nov. Left, apex of flowering plant. Bottom left, apex of column with anthers removed. Top right, apex of triandrous column, Upper right, median anther, and to its right, one of the lateral anthers. Plate 13. Vanilla Helleri A.D. Hawkes, sp.nov. Left, portion of leafy stem with flowering inflorescence. Center left, lateral view of column. Top center, frontal view of column. Dissection of floral parts. Plate 1. Epidendrum Hawkesii A.H.Heller, sp.nov. 2h FRY TOL0:6:1:2 Vol. 1h, no. 1 Plate 2. Helleriella nicaraguensis A. D. Hawkes, gen. et sp.nov. Heller & Hawkes, Nicaraguan Orchid Studies Lip Spread With column 2mm, sp.nov. Lepanthes Helleri A. D. Hawkes, Plate 3. PHYTOLOGTIA Front Vol. 14, no. 1 Section Plate 4. Pleurothallis Alexii A.H, Heller, sp.nov. 1966 Heller & Hawkes, Nicaraguan Orchid Studies 27 —— Floral Bract Sec. of Lip, Column and Lateral Sepals 4mm. Underside of Base of Lip and carinae LLP Section C-C' of Petal Section A-A' QD . _RB! Basal Lobes of Lip Section B-B Spread Plate 5. Pleurothallis carnosilabia Heller & Hawkes, sp.nov. 28 PHYTOLOGIA Vol. 14, no. 1 Plate 6. Pleurothallis chontalensis Heller & Hawkes, sp.nov. 1966 Heller & Hawkes, Nicaraguan Orchid Studies Plate 7. Pleurothallis exesilabia Heller & Hawkes, sp.nov. ———— lcm. Longitudinal Section of Leaf. ® Ny Underside of Lip. Callus LES Ye 30 PHYTOLOGIA Vol. 14, no. 1 | Plate 8. Pleurothallis Helleri A.D.Hawkes, sp.nov. 1966 Heller & Hawkes, Nicaraguan Orchid Studies 31 Section across Disc Sobralia chatoensis Heller & Hawkes, sp.nov. P RY.7-01..90.0.14 Vol. 1h, Section A-A' Plate 10. Sobralia Hawkesii A.H. Heller, sp.nov. ee 1966 Heller & Hawkes, Nicaraguan Orchid Studies 33 Plate 11. Sobralia Helleri A.D. Hawkes, sp.nov. 3h PHYTOLOGIA Vol. 14, no. 1 Plate 12. Sobralia triandra Heller & Hawkes, sp.nov. “ | iy Lateral Anther '——— 4 mm. Heller & Hawkes, Nicaraguan Orchid Studies 1966 — Vanilla Helleri A.D. Hawkes, sp.nov. Plate 13. ADDITIONAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS CALLICARPA. III Harold N. Moldenke & ARBOREA Roxb, Ridley (1923) records this species from a Island, giving its general distribution as "India, Burma, Sumatra". Craib gives its general distribution as “India, Burma, Assam, Yunnan, Malaya" Lam (1919) says that "In the bibliography about the subject we discover some indications, that some authors cannot find and dif- ference between C. arborea Roxb. and C, lanata a (e.g., Mia. Fl. Ind. Bat. Suppl. 1, 243 and 569; Koorders en Valeton, Bijdr. Booms, Java.VII, 175). We too, think so. So we have combined the two species "under the name C, lanata a, finding no essential difference of any importance, that could separate them distinctly. Even if it should be shown that there are a treelike and a shrubby form, it would not be exact to consider them as more than two Ks eties of one single species. -- It has a strong affinity with Geunsia farinosa, var. a, from which it is different, however, by its opposite leaves, l-merous flowers and fissury opening of the anther-cells,." King & Gamble (1908) describe their C. arborea var. villosa as follows: "Leaves 6 to 8 in. lone, 3 3.5 to 4.5 in. broad, ovate, caudate-acuminate at apex, rounded lightly decurrent "at base. Tomentum of branchlets, young Terres and inflorescences very thick and pale, chiefly consisting of branched hairs. This var- resembles C. lanata, Linn., of southern India and Ceylon in its leaves, but the Flowers are those of C. arborea. Perak: Scorte- chini.” For the typical form of the species he cites Ridley 5607 from Kedah, Deschamps s.n. [Batu Gaja jungles] and Curtis 699 from Penang, and Wray ray 182) & 3957, Ridley 7179, Scortechini § s.Ne, and King's Collector 972 (in part) from Perak, as well as Tei js- mann ),388 and Forbes 1530 & 2601 from Sumatra. He gives the species’ distribution as_ ae gress to Assam, Sonthal Pargan~ as, Chota Nagpore, eastern » and Sumatra Bakhuizen van den Brink (age) Syme C. arborea te Ce. tectoniolia Wall., and C, villosa Roxb, in the synonymy of C. to- mentosa (L.) Murr. Pei (1932) places C. vastifolia Diels in the synonymy of C. hina. The material above cited _—< very closely with the typical Indian form of Galligarpa axtores arborea Roxb. Rock 7197 has smaller leaves which are acute at both en ends, but the other charac- ters are same as in C, arborea Roxb. Rock 6977 seems to be identi- % 1966 Moldenke, Monograph of Callicarpa 37 cal with Diels' species, having the subcordate and rotundate leaf bases but acute bases are also found on the same specimen." Dop (1932) places C. farinosa Roxb. in the synonymy of C, ar~ borea, but it belongs, - rather, in that of C. tomentosa (L. Aa Murr. Craib (1912) cites Kerr 691 and Hosséus 6. 618 i aig i He gives +e species' distribution as "India, Burma, » Xunnan, " and says of Hosséus' C. lanata: "Hosseus! ase "in the shape of the leaves is very similar to Callicarpa villosa, Roxb. (Callicarpa arborea, var. villosa, King et Gamble) which ori came from Sylhet. tn no case can it be referred to the Indian Callicarpa lana Common and wonnaoetar names recorded for C. arborea somes st" tapoeéng" *wsuk Id kai", "stnga", "tepong", "tepong tepong", "espong-tépone”, "tind jaoce", and "turmong". The villosa Bald., referred to in the synonymy above, is a synonym seas C. americana L., while C, villosa Vahl is a valid spe- cies, which si see; C. arborea Merr. is a synonym of C. arborea var. psilocalyx (H. J. Lam) Moldenke, C. arborea Miq. is 5 actually Cc. tomentosa (L.) Murr., and C. arborea Wall. is C. vestita Wall.; 3 Ce ianata Gamble is actually Cc. vestita Wall., C. “Tanata L. and Cc. lanata Wall. are c. tomentosa (Le) M Murr., C. - lanata H. J. Lam is c. arborea var. psilocalyx yx (H. J J. Lam) Moldenke, C. lanata “C. lanata Lam. is Premna - tomentosa Willd., and C. lanata Schau., C. “lanata Vahl, and Cc. lan lanata Zipp. are C. pedunculata R. Br.; C. tomentosa Bakh. is in part C. arborea and in part C. integerrima Champ., Champ., C. tomentosa Hook, & Arn. and C C. tomentosa Willd. are C, loureiri Hook. & Arn., C, tomentosa Ktnig : is C, macrophylla Vahl, | C. tomentosa L. is C. erioclona Schau., C. tomentosa Lam. is Cc. candicans (Burm. f. Peg Hochr., C. tomentosa Murr. is C. tomentosa (L.) Murr., C, tomentosa Thunb. ae Ce. Sa Lan., and the identity of c. tomentosa ga Vahl is not yet e established. King & Gamble eee distinguish their C. maingayi as follows: "A Callicarpa arborea, Roxb., foliis obtusis subtus pilis minutiori- bus aureofulvis tectis, corollae tubo longiore et lobis brevioribus distincta." Ltrzing 5857 has the inflorescences tremendously galled, while Rock 7197 | 7197 has as both the leaves and the inflorescences galled. The Herb. Hor Hort. Bot. Bogor. 8.M. specimens, cited below, were grown from seeds eeds brought from Sumatra -- there are toothed leaves on some 38 PHYTOLOGIA Vol. 1h, no. 1 of the sheets, Yates 1078 in the University of California herbar= ium has a photograph | of the plant mounted on the sheet. Nair & Rehman (1962) describe and illustrate the pollen grains and cite Nat. Bot. Gard. 2209, slide 2653. ~~ Material of C, arborea has been misidentified and distributed in herbaria under the names C, farinosa Roxb., ee lanata var. typica H. J. Lam, C. pentandra Roxb., C C. pentandra var. typica f. farinosa (Blume) Bakh., C, reevesii Wall., C. wallichiana Walp., Geunsia | farinosa Blume, and G. pentandra (Roxb.) Merr. On the other h hand, the | the Carr 11317, distributed as Cc. arborea, is actual- ly Cc. macrophylla Va Vahl, and H. 0. Forbes 1530 and Rock 983 are Ce tomentosa (L.) Murr. ~~ Merrill (1903) cites his no, 687 from Mindanao, but it is probable that this is actually var. psilocalyx. He says "No. 5h represents a related form, but quite distinct!’ He gives the he dis- tribution of the species as "Himalayan region to Sumatra and New Guinea". Ridley (1920) cites his no. 6592 from Koh eee Malaya, and gives the distribution of the species as mere ly "Malay Penin~- sula". In his 1923 work he cites Nur 726k from rey nape Bukit Kramat Kuda Dop (1932) cites the following specimens: THAILAND: Kerr 691. INDOCHINA: Annam: Chevalier 32505; Poilane 1612. Cambodia: Har=- mand s.n. [Pnom Penh]; Pierre e 1220. Cochi: Cochinchina: Pierre 182. Laos: “Harmand 1199; Massie s.n. 1. [Attopeu]; Perrot s.n. sn. (Attopeu]; Thorel s.n. “gen. (Paklai]. Tonkin: dn: Balansa 3811, 3812 » 3613; chev 29497. He comments: "Pour H. J. Lam et Bakhuizen van fan den “Brink le binéme C. arborea est aa en synonyme pour des raisons de ar & la monographie de Schauer ig Fem le Prodrome de De Candolle étant en ohalersl insuffisantes. Comme le binéme C. arborea a été util- isé par tout les auteurs des travaux modernes sur sur les flore o-malaises: Kurz, 8 te In all, 207 he rbariun specimens have been examined by mi Additional citations: PAKISTAN: East Bengal: W. Griffith “6037 (Cp, Cp, S). NEPAL: Ram 300 Oe — sans e 7056 (N), 9 (ms Ee Thomson s.n. [Sikkim] (Bz: 18666). INDIA: AS sam: hand 152 (a), 5098 (Mi), 595 5 (8) “hatterjee son. (Shak hati, = ase (Po——-61,810); Collecto wn, [Assam] (W—9972) ; Jenkins s.n. [Assam] a aaer ee Bz-—18670, Bz—18671, Bz—18673, Bz—-1867h); Koelz 22718 (Mi), 2h502 peed ses (ui), 25043 (ui), 26u78 (Mi), 28877 (Mi), 30119 (M4); W—261235), 220 (W—2612 123k); Simons | s.n. [As san] fend a a Watt 11305 (Na—12912). Khasi States: Hooker & Thomson s.n- {Mont. Khasi] (M, S); G. Mann s.n. [Khasia] ] (Bz—~18661) . Madras: Koelz 1086 (Mi). Manipur: hissboia 7120 (S). Uttar Pradesh: Har- 1966 Moldenke, Monograph of Callicarpa 39 sukh 500 (S); Koelz 2053) (Bv, Bv, N). West Bengal: C. B. Clarke 11723 (W—-802338), 11613b (Bz—18667); Mukerjee 1%1 (S). State undetermined: Duthie 2,835 [Shimala] (Ca~—-269793). BURMA: Tenas- serim: Brandis s.n, [Pegu] (Bz--18659); Falconer 505 (Bz--18657); Helfer 6037 (T). Upper Burma: Forrest 2)68h (W—1377450); Khalil son. [Southern Shan States, 1893] (W—369326); Khan 139 (Bz— 18658); Kingdon-Ward 22196 (Bm). ANDAMAN ISLANDS: Barren: Prain sen. [April 1891] (W-—-261236). CHINA: Kwangsi: Ching 7291 (N, W 1248676); Tsang 23914 (N). Yttman: Forrest 21079 (W—1278917); A. Henry 9551A (W-57158), 12093 (N), 12093a (N); J. F. C. Rock 6977 (Ca—328045, W--1332122), 7197 (Ca--328)16, W—1332123). THAILAND: Bunkird s.n. (Herb. Roy. Forest Dept. 25225] (Bk); Chamruaengsri 39 (Herb. Roy. Forest Dept. 1945] (W—206)781); Mrs. D. J. Collins 1251 (W—~1701059); Khantchai 350 (Herb. Roy. Forest Dept. 15879] (Mi), 1080 (Herb. Roy. Forest Dept. 24283] (Mi); Khit 128 (S); Kostermans 55 (Bz—-73295); J. F. C. Rock 1589 (W-—-12145h7); Sind- en ett 72616); Collector undetermined s.n. (Bz-~18620); Cramer 91 (Bz— 18637); Diepenhorst 2563 B.B. (Bz—18617); Endert 65 (Bz—18633, Bz—1863)); Idenburg 32 (Bz—18608, Bz—-18609); Jacobson 20 (Bz— 1862); Junghuhn sen. (Ut—53392); Karta 54 (Bz-18694, Bz—18695, N), 72 (Bz—18696); Koorders 901* [29479b] (Bz-—186h9), 1OL77b (Bz—1868), 29179 (Bz—-18650, Bz—18651); Krukoff 248 (Bz—-18615, N), 3h9 (Bz-~1861, N); LOrzing 3135 (Bz——18627, Bz—-18628), 3522 (Bz—18630), 5857 (Bz—-186kL, Bz-~18645), 7933 (Bz--18635), 9984 (Bz—-18612), 11239 (Bz—1863); Nur 7264 (Bz—-18697); M. M. S- Tei jemann 1261 H.B. (Bz—18618, Bz—16619), 116b-2563-3525 (Bz--18621), 3525 HB. (Bz~-18616, Ut—53391), 4388 H.B. (Bz—-18640, Bz—-186h1, Ut-- 5339), sen. (Bz—-18699, Bz—-10700); Toroes 10h5 (N), 366 (N), 1,363 (N)j Van Steenis 3 (Bz—19613), 6262 (Bz--1870k); Voogd 1211 Tob 216316) (B2-—18698); H. S. Yates 629 (Ca—226102, Mi), 1078 (Be- 72799, Ca—-251168, Mi, N, 8), 2651 (Bz——18675, Ca—325716, Mi, N, ho PHT f0O-L 0G-bA Vol. 1h, no. 1 S). MELANESIA: NEW GUINEA: Papua: Brass 528 (Bz—-17462), 3675 (Bz—18676); Carr 11535 (N); M. S. Clemens 10870 P (Ca--180781); Hoogland 3905 (Ng—16872, Ng). CULTIVATED: Hawaiian Islands: Fe Brown 1276 (Bi, Bi). India: Herb. Hort. Bot. Calcutt. D.8 (Bz— 18660), sen. (Br, Bz—18663, Ed, Mu--951, T, X); Wallich 1826 (Ed, Mu—~1),27, S), s.n. (Cp, Cp). Java: Herb. Hort. Bot. Bogor. X1.G.83 (Bz—25791, Bz—25792, Bu—-26526, Bz--26593, Bz, Bz, N, N), X1.G.83a (Bz), Sen. (Bz—18688, Bz—-18689, Bz—~18690, Bz— 18691, Bz—-18692). Pakistan: Kurz s.n. [Sibpur, lower Bengal, 5/ 67] (Ba--18664). Sumatra: Herb. Hort. Sibolangit 246 (Bz—26507). eter ate B. [Sitapong] (Ut—53393). CALLICARPA ARBOREA var. PSILOCALYX (H. J. Lam) Moldenke, Phytolo~- gia 4: 125. 1952, Synonymy: Callicarpa magna Schau. in A. DC., Prodr. 11: 61. 1847. Callicarpa manga Schau. apud Elm., Leafl. Philip. Bot. 3: 865, sphalm. 1910. Callicarpa lanata var. psilocalyx H. J. Lam, Verbenac. Malay. Arch. 81. 1919. Callicarpa tomentosa var. magna (Schau.) Bakh., Bull. Jard. Bot, Buitenz., ser. 3, 3: 22. 1921. G ia straminea Elm. ex Bakh., Bull. Jard. Bot. Buitenz., ser. Je3? 22; in ayn. 1922. Callicarpa magna var, lilacina Eln., Leafl. Philip. Bot. 9: 3222. 193). Callicarpa arborea Merr. ex Moldenke, Prelim. Alph. List Invalid Names 9, in syn. 1940 [not C. arborea Miq., 1885, nor Roxb., 181), nor Wall., 1829]. Calli- carpa lanata H. J. Lam ex Moldenke, Prelim. Alph. List Invalid Nanes 11, in syn. 190 [not C. lanata Gamble, 1878, nor Hosséus, 1912, nor L., 1771, nor Lam., 1821, nor Schau., 1893, nor Vahl, 1856, nor Wall., 1940, nor Zipp. 181]. Bibliography: Schau. in A. DC., Prodr. 11: 61. 18h7; Miq., Fl- Ind. - 2: 889. 1856; Bocq., Adansonia 3: [Rev. Verbenac.] 192. ° a Bot. Buitenz., ser. 3, 3: 21 & 22. 1921; E. D. Merr., Enum. Phil- ip. Flow. Pl. 3: 386. 1923; H. J. Lam in Lauterb., Engl. Bot. Jahrb. 59: 89. 192i; Elm., Leafl. Philip. Bot. 9: 3222. 193h; Moldenke in Fedde, Repert. Spec. Nov. 0: 49. 1936; Moldenke, Alph. List Common Vern. Names 3 & 20, 1939; Moldenke, Prelim 1966 Moldenke, Monograph of Callicarpa ral O, 141, & 177. 19493 Moldenke, Alph. List Cit. h: 1259. 19h9; wotdente, Phytologia : 77, 122, & 125. 19523; Moldenke, Biol. Ab- tr. 27: 98h. 1953 3; Moldenke, Résum 6 182, 2h, 2hh, 245, 2h7, 295, 3. 19593; Moldenke, Résumé S ppl. 3: 30 (1962) and 1h: 7. logi 8 surface and the calyx glabrous or with only a few scattered hairs It is described as a tall tree, 22--30 feet in height; trunk h 1/2 ey 8 inches in geese] at breast height; bark smoothish, blades membranous, elliptic or oblong-ovate, 15--22.5 cm. long, short-acuminate at the apex, rounded at the = and shortly pro- longed into the petiole, glabrate above when mature, venos pas ie “ re e-canescent with stellate tomentum beneath; canmpanulate, twice as long as the calyx, its limb velutinous a ve The ee was based by Lam on Cuming 1266 from the province of Al Luzon, Philippine Islands [cited | from Manila by Bocquil- lon (1863)], on ye Ramos = Towne 1910] and 15038 from San An- tonio, province of Laguna, » and on Nyman 1053 1053 from Friedrich Wilhelmshafen, October 1, amy Rodatz & Klank 185 | and Schlechter 1645) from Northeastern New Guinea. He cites Lauterbach 1200 & 1241 from Constantinshafen with a question. In his 192) work he is in doubt only about the 1200 collection and spells the collec- tor's surname "Klink", The type of C. magna is also Hugh Cuming's no. 1266; that of Ce. var. lilacina was collected by Adolph . Daniel Edward El- mer (no. 22377), and that of Geunsia straminea is Elmer 8193, both also from the island of Luzon. Bakhuizen van den Brink (1921) fenced C. sorsogonensis Elm. the varietal soar Nahr [including C. sorsogonensis] as sean "Folia adulta subtus dense tomentosa vel su sa mollia v subrugosa, eat in sicco subpersistente + elena, at aS longa; Geel 0 0M. lata; cymi mediocres vel majusculi, peduncul petiolum sub-aequante vel saepe majore, raro breviore; calyx bra vel pilis sparsis stellatis; corolla extus sige vel dorso laciniarum velutina; ovarium glabrum raro sparse is vestitum, glanduli Fags crebris densissime tectimn.* Elmer 16939, but this I regard as Geunsia palotnsis h2 PHYTOLOGIA Vol. 1h, no. 1 Callicarpa arborea var. psilocalyx has been found growing in old secondary forests, at 700--800 meters altitude, flowering in May, July, and October, fruiting in ps a ae R. S. Willians 1003 epee includes a wood gig Saas ar names re- verve r the plant are "at{mla" ‘sageitbtrdy he "nagilig". Ma- pe tie been misidentified and * distributed in herbaria as Premna sp. In all, 22 herbarium specimens, including type material of all the names involved, and 3 mounted photographs have been examined Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Luzon: Cuming 1266 (N--cotype, Z--photo of cotype); Elmer 8193 (Bz— 18686, N), oe (Bz--1868), N, Ut—26960, Vt), 17575 (Bz--18687, ca—271612, Ny, S, Ut--67267), 22377 (Bz——18096, Ca-~7763, Mi, N)5 M,. Ramos 35 (photo. of ot pe, Ut-—-221 95--cotype, Z--photo of cotype); R. S. Wi N, W—707311). Mindanao: D. Fairchild rit crn wiskey 287 287 (Mi). — AREOLATA Urb. in Fedde, Repect. Spec. Nov. 20: 3h6. 92h. Bibliography: Urb. in Fedde, Repert. Spec. Nov. 20: 36. 192h} fs - Kew. Suppl. 7 36. 1929; Moldenke in Fedde, Rep- pec. Nov. 39: 301 (1936) and 0: 63-6), a ant : 129. 1933 Moldenke, Geogr. Distrib. Avicenn. . 1939; Popes: Geogr, Distrib. Ver a ed. 1, 2h & 86. 1942; mus ” alph List Cit. 1: 75 & 185 (19)6) 2s 187, 648, 649, "e 651 Psst 31 713 & 929 (1949), and 4: 103). ae Molde: — ween trib. Verbenac., [ed. 2], 2 ‘ 176. 1949; n in Leén & Al ain, FL. Guba bs 305" 309. 19575 Moldenke, hes oe ise 19593 J -« Clark, Gen. Sp. Pl. n Rec ent collectors aie found this plant growing on limestone and oh the borders of limestone cliffs, at 60 meters altitude, flowering July, and frui ruiting in May and July. Material has been misidentified and distributed in herbaria as C. grisebachii anc: ta eo. Urb. ee ae In all, 2) herbarium specimens, Rete age type (now de- stroyed), ” and 11 mounted photographs have examined by me. Additional citations: CUBA: Oriente: team "1m8) (N); Acufia & Barreto 17442 (Es); Acufla, Cereza, & Montenegro 1718 1718 (Es); Alain & Morton 5222 (2); G. C. | Bucher cher 102 (Herb. Roig 7604] (Es), 10558 (Es); Ekman 361) (E—photo of type, photo of type, W-- Photo of type), W487 (N); Leén 17158 (Ha, N), 17269 (Ha, N); Leén & Victorin 17331 (Ha); Matos s.n. [Leén 17738] (Ha, N), sen. [Le- én 18378] (Ha, N). 1966 Moldenke, Monograph of Callicarpa 43 ae Nohdacihe * Résumé ane & 43. 1959. the young po ns of the branchle ts and the buds ee eee eee the older branches becoming cinereous; glabres- ex, ser _ long the margins, subentire toward the base, acute or acutely at- tenuate at the base, loosely floccose on the venation on both sur- pea. otherwise glabrous, minutely sharilacctankitees beneath; ndaries 7--10 per side; cymes ample, greatly many-flowered soon glabrescent; peduncles equaling the petioles or rarely twice as long; bractlets linear, deciduous; calyx-tube cupuliform, minu- tely granular~glandulose on the outer surface, the rim obsoletely 4-dentate; corolla-tube turbinate, glabrous, somewhat longer than the semi-orbicular recurved lobes; anthers exserted, minutely granular-glandulose. No type is cost qneted for this species and it is said by the author to grow in "Sikoku, Kiusiu, Lutchu", with the dag rr cd name of "oh-murasaki". He classifies it in the pectioa Cyathi morphae and says of it "Haec species C. japonicae affinis, sed a r sensim valde multiflora." I have seen no authentic material of it and am inclined to think that it may prove to be synonymous with C. Japonica var. iuxurians CALLICARPA BARBATA Ridl., Kew Bull. Misc. Inf. 1929: 260—261. 1929. Bibliography: H. N. Ridl., Kew Bull. Misc. Inf. 1929: 260—261. id A. ve nee age Ind. Kew. re 8: 37. 1933; Moldenke, erbenac » 65 & 86 (1942) and ed. 2, 145, libs & 177. = 19195 Moldenks, gm er 193, & by3. 19593 Anon., Index 1 9-1956, 59. 1 “ek niece shrub; pee ves decussate-opposite; leaf-blades hartaceous, large, o t 1 cm. long, 1 wide, acute at the apex, undulate and denticulate along ns, uate at the base and decurrent into the petiole, the mar venation red-hirsute with plumose hairs; es 10 per side; veinlet reticulation elevated beneath, sparsely rufescent-pilose above; panicles robust, borne on the stems in the axils of fallen cm. long, 8-cm. wide, many-flowered, stellate-pubescent; s and inflorescence-branches l-angled; pedicels slender, ea ments very short; style e 3 8 idee pe ape patelliform, 7 mm. eye fruit ep ad 5 mm. wide, -celled cell 1-seeded. e of this species was collected by Geo Darby Haviland (no. 3043) at Rejang Kapit, Sarawak, in March, 1893, and is depos- hh PHYTOLOGIA Vol. 1h, no. 1 ited in the herbarium of the Royal Botanic Gardens at Kew. The eer says of the plant "C,. basilanensi Merrill approximat, sed is magnis barbatis, floribus in paniculis robustis longe pedi- suriice glabris corolla 5~loba staminibus 5 distincta....The leaves, the stem bearded. The tt Bos not Kim but possess a lateral branches, plumose." s known to me of this weasins except what is stated about it in the bibliography. CALLICARPA BASILANENSIS Merr., Philip. Journ, Sci. Bot. 3: 263. BA Bibliography: se! md ” Philip. Journ, Sci. Bot. as 263. 1908; Prain, Ind. kK - kh: 3h. 1913; H. J. Lam, Verbenac pena Arabs 5253 8 & pall. 19193 Bakh., Bull. Jard. its Bui- Aes benac bates! Phytologis Re 93. 19453; Mold denke, Known Geogr. Distrib. Verbenac., ed. 2, 140 & 177. 1949; Moldenke, Rhounb 1 182 ‘& M3. 19595 Moldenke, Pray Suppl. — 8. 1965. Small tree, 5-10 m. tall, cauliflorous, the young parts covered with dense rust-colored hairs; Leaves de ecussate-opposite; petioles 1—2 cm. long, j leaf-blades subcoriaceous, oblong ovate or elliptic-oblong, 25 em. long, 1 long caudate~acuminate at axi 2.5 cm. long, etek Be stellate-tonontoses 3 flowers glabrous; calyx 3 mm, long, 2.5 mm. wide at the apex, the rim a teens; corolla 7--3 mm. long, the lobes 4 or sometimes 5; filaments 5 mm. long; an o> mm stigma more or less ieclotiedls fruit glo- bose, 5 mm. wide, nieasoone of 4 pyrenes, pecies was based by Merrill o n two collections made b Wallace ee Falstae (Herb. Philip. Forest Bur. 397) & 62h in January and July, 1 ao 6 oN 1 an unnumbered one made Gottfried Hallier in Faas , all on the island of Basilan, Is » and rol Bete in the herbarium of the uu pe amen at Manila, but stroyed. The Hallier specim e The Pascua s.n. {Herb Philip. rv Bur. 30230] ee catraay ae "Basilan and Zam paiigh its label and Oo may actually consist of material collected on both these i siting Me (1908 says gos this is "A species well characterized by its large, acuminate, entire leaves » fascicled cymes from the axils of fallen leaves, jp ather large glabrous flowers » which 1966 Moldenke, Monograph of Callicarpa Th are rarely 5-merous, hence intermediate between Callicarpa and Geunsia." Lam (1919) ~ "This plant is nearly related to Geun- sia ‘sia Cumingiana var, . by its sometimes 5-merous flowers." Bakhuizen van den Brink *Peoot) places it in the synonymy of what he calls C. pentandra var. cumingiana f, typica Bakh. (=Geunsia cumingiana a (Schau. ) Rolfe]. ~~ The spe e species has been found growing in nO pain and "parang" at altitudes of 300 meters. Vernacular names recorded for it are "eagayuc" and "linagop". Material has been misidentified and distributed in herbaria under the names C. pentandra tandra f. pubescens Bakh., C. pentandra var. cumingiana f. typica Bakh., and C. pen- tandra var. cumingiana f. genuina Bakh. In all, 9 herbarium specimens, including cotype material, have been examined by me. Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Basi- lan: Hallier 25 (Le—937.287-53--cotype); W. I, Hutchinson s.n. [Herb. Philip. Bur. Sci. 612] (N—cotype); Pascua s.n. (Herb. Philip. Forest Bur. 30230] (B, Ca~-320987, N); C. B. Robinson s.n. {Herb. poten, Bur. Sci. 11502) (Bz--18202). Mindanao: Ramos & . [Herb. Philip. Bur. Sci. 36727] (Bz—18196). “Santa Cruz: 3 Br Gised 953 (Cp), 990 (Cp). ee BASITRUNCATA Merr. ex Mold oc. Venez. Cienc. Nat. 11: 9, nom. mud. (19h7), Prytofogia tae 06-107. Eek Bibliography: *Moldenke, at Soc. Venez. Cienc. a 5 1917; Moldenke, Known Geogr istrib. Ver rbenac., e ed. 2, 135 & i07. 1949; Moldenke, Phytologia 3: pr aaianey 1951; eases’ Biol. Abstr. 25: 3050. 1951; Moldenke, Résumé 17h & hh3. > im, tall; bran ches and br: ree hg tne light- colored, *medullose, * obtusely tetragonal, o rely furfuraceous or stellat e~pube erulent , becoming glabrate i aah twigs very slender, brown, rather eo achean ee entellous with aoe cinereous hair, densest toward th se cae | aia not venation beneath when young, glabrescent in age (sometimes a few stellate hairs on the upper surface, especially along the midrib, ure) ~axi much abbre usually only bractlets and prophylla minute; calyx about 1 mm. long and wide, very minutely and sparsely stella llate-puberulent or glabrate, its rim very shallowly and obscurely -toothed; corolla white ,» its h6 PHYTOLOGIA Vol. 1h, no. 1 tube about 2 mm. long, glabrous, its limb l-lobed, glabrous; sta- mens exserted. The type of this species was collected by J. Linsley Gressitt (no. 1168) at Liamai (Leng Mun), on a bank by a Hainan Island, at 20 meters altitude, in June or J 935, and is deposited in the Langlois Herbariun at the Catholic “University of America in Washington. The species is known to me rom the type specimen Citations: CHINESE COASTAL ISLANDS: Hainan: Gressitt 1168 (I-- type). neeacaare BAVIENSIS Moldenke, ome 13: 205. ibliography: Moldenke, Lloy dia 13: 205. ate ee Phyto- logia 3: 380. 1950; Moldenke, haves 175 & 43. 19 Shrub; branchlets and twigs slender, obtusely tetragonal, densely sordid-puberulent; nodes often annulate; principal inter~ nod ve, on beneath, elliptic-ovate or ovate, l--8.5 cm. long, 2. 23 CMe wide, short-acuminate at the apex, entire, varying from acute to obliquely truncate at the base, minutely puberulous on both sur- faces, especially along the larger venation, glabrescent on the lamina » pronin ndar’ filiform, 4 or 5 per side, arcuate-ascending, y plane above and slightly p nulous beneath; veinlet ftp eps rather a- neg su axillary, cymose, usually shorter than the iabbaasing petioles rather few-flowered; peduncles filiform, to 6 mm. long or rackets, densely puberulent; pedicels filiform, 1--2 m. long, densely puberulent; bractlets setaceous, about 1 mm. long, densely puberu- lent; ¢. te, about 1 Wi more or less puberulent and resinous—punctate, its rim subtruncate, minutely h- apiculate; corolla » White, its tube mm. long, densely r on the outside and microscop- inous-punc ically puberulous, its lobes about 1 mm. long, erect, resinous- punctate on the back. The type of this species was collected by Paul Alfred area (no. 2636) in an open ec at about 600 meters altitud Mount Bavi, in the province of Sont&y, Tonkin, fale. on June 4, 190, and is deposited es the Britton Herbarium at the New York Botanic. al arden. The species is known to me thus far only from the type specimen Citations: INDOCEINA: Tonkin: Pételot 2636 (N—type). CALLICARPA BICOLOR A. L. » Ann. Mus. Hist. Nat. are, [tn t4s 1806 [not C. renee aceee 1886, nor F. ‘Vall. eae Bibliography: A. L. Ju Juss., Ann, Mus. Hist. Nat. eis 73 "1. 1966 Moldenke, Monograph of Callicarpa 47 1806; sheet, Nom. Bot., ed. 1, 137. 1821; Spreng. in L., Syst. Veg., 16, 1: 420 (1825) and 5: 126. 18283 D. Dietr., syn. Pl. ls ido. “4833 Steud., Nom. Bot., ed. 2, 257. "1810; Walp., Repert. h: 128. 18)5; Schau. in A. DC., Prodr. i 62. 1847; Miq., Fl. Ind. Bat. 2: 889. 1856; Bee: ’ Adansonia : [Rev. Verbenac.] 192. 1863; Benth. & Muell., Fl. Austral. Lap rr "1870; 0; Vidal, Fl. For. Filip. Atlas pl. 74, fig. c. 1883; oe Journ. Linn. Soc. Lond. Bot. 21: 315. 188; Jacks. in Hoo Me rik nt ss Ind. Kew. 1: 386. 1893; Briq. in Engl. & Prantl, Nat. cia nfam. 18953 Koord., Meded. Lands Plant-tuin el tai, 19: 558. 1898} E. e 1 906; H . Je Bot. Buiteng., ser. 3, 3: 10 & 19—20. 1921; E. D. Merr., Enum. Philip. Flow. Pl. oh 388-389. 1923; Bakh. in Bakh. & Lam, Nov. Guinea 14, Bot. 1: 167-168. 192h; Stapf, teat Ind. Lond. ls: 525. 1929; Moldenke in Fedde, Repert. Spec. - YO: 99 & 109. 1936; Moldenke, Phytologia : 122--12h. 1952; ‘uotdenee, Résumé 182, 19h, 198, & seed 19593 ape cee. Résumé Suppl. 3: 21. 1962; Moldenke, Phytologia 13: 499. Illustrations: Vidal, Fl. oe Filip. Atlas pl. 7, fig. c. 1883. Shrub, about m. tall; stems to 2 cm. in diameter; wood hard; branchlets su pherecticde with furfuraceous tomentum, sub- tomentose, lightly whitish-tomentose; leaves decussate- opposite; "esti ned short, subferruginous with furfuraceous t mentum; leaf=blades dark-green, elliptic or elliptic-lanceolate to pt he Pinerrennce i almost 10 cn. eek es ee: e at the apex, mate or mucronate-cre ong th ate-angustate — the base and attenuate into the eentoas ealiseas and shiny above, eglandulose, penninerved, venose-reticulate be- subglabrate, sparsely punc paceous, about the size of a grain of mustard (sin sh oF ihe) violet, l-seeded. and is deposited in the Jussieu Herbarium at the Muséum Na d'Histoire Naturelle in Paris. Sprengel (1825), Dietrich (1839), and Steudel (180) make the remarkable claim t this ne se ak Para, b BS ees England, U. S. A.] — what they in- tended . Brit.” [=New coe Vabeoens #) Bri- quet (1895 ae ess Pe 5 diptviebion as "auf d askarenen un den Philipp n, hauer (1847) records it ng ote Sechelles (ex have never seen any material of it from the slands dict. and doubt very much if geogr. yn, Seychelles nor from the Mascarene I 48 PHYTOLOGIA Vol. 1h, no. 1 it occurs there in the wild state. The species has been found growing in secondary forests and on coral rock, at altitudes to 50 meters ae sea-level, flowering & Sei. 48132] and on Feuilleteau de Bruyn 280, and as "violet Ramos & Edaflo Son. {Herb,. Philip. Bur. Sci. 49172], but the latter statement may be an tags for the fruit-color instead of +t flower-color, since the collection is both in flower and fruit. The C. bicolor of iar. referred to in the synonymy above, is a synonym of C. candicans (Burm. f.) Hochr., while that of Fernandez Villar is C. formosana Rolfe. Jackson (1893) reduces "C. bicolor, Juss." to C. cana L. [=C. candicans (Burm. f.) Hochr. ] and "C. bicolor, | F. Vi Vill." to C, blancoi Rolfe (=C. for- mosana Rolfe]. ~~ Bakhuizen van den Brink (1921) includes C. paucinervia Merr. in the synonymy of C. bicolor, but I regard I errill's taxon as a variety of C. erioclona Schau Gahan, Bentham & Mueller (1870) regard C. bicolor as "precisely the common Archipelago form of C. Cana." Admittedly, C. bicolor is very closely related to both Cc. . candi- cans and to Cc. erioclona, ntteriel has been misidentified and distributed in herbaria r the name of C. cama L. On the other hand, the Kjellberg 96 pris distributed as C. bicolor, is stuns C. candicans, Bish 68 and Elmer 18086 are eC, erioclona Schau., and F, L. Stevens 1429 is Buddleia asiatica L. in the Loganiaceae. Schauer (1847) says of C. - bicolor “Proxima C. canae, sed di- versa pube alien pracci pus | in eigen anrto eager ut puncta glandulosa neuti areant, et cymis us pedunculatis bipartitis." He athe uming 1083 from Tog Pesiiceine:. Miquel (1856) says "C. canae affinis, tomento distincta" and also cites the same Cuming collection. Bakhuison van den Brink Mone cites Feuilleteau de Bruyn 280 from Schouten Island in Dutch New Guinea. In all, 24 herbarium specimens have been examined age me. Citations: WESTERN PACIFIC TSLANDS: PHILIPPINE ISLANDS: Alabat: Ramos & Edafio s.n. [Herb. Philip. Bur. Sci. 8132] (Ca—-321616) . Cebu: Je v. Barrow Sen. [Cebu, 190)] (N); Usteri sn. Sasha (N). Luzon: E. D. E. D. Merrill 2715 (N), 3bh2 Mindanao: s & Edaflo s.n. [Herb. Philip. Bur. Sci. 9172] (Be--17263, eaand N). Negros: E. D. Merrill 207 (Ut—22)75). Panay: Martelino Edafio s.n. [Herb. Philip. Bur. Sci. 35531] (Bz—~17461). INDONESIA: GREATER SUNDA ISLANDS: Buton: KJjellberg 96 (Bz—~17288, Bz--17289)- Celebes: Kjeliberg 14 (Bz—17287), 1275 (Bz—17276) ; Koorders ighg90b [3921] (Bz—17283), 19491b [58] (Bz—1728h), 19hghb [98] (Bz—17285, Bz~-17286) ; Lam 2471 (Bz--17277); Noerkas 422 (Bz— 1966 Moldenke, Monograph of Callicarpa ho 17278, Bz-~17279, Bz—-17280). MOLUCCA ISLANDS: Halmahera: Anang 426 (Bz--72770). Ternate: Beguin 1698 (Bz—17)59). CALLICARPA BODINIERI Lévl. in Fedde, Repert. Spec. Nov. 9: 56. 911. : Callicarpa seguini Lévl. in Fedde, Reyert. Spec. Nov. 9: 455—~56. 1911. Callicarpa feddei Lévl. in Fedde, Repert. Spec, Nov. 10: 39. ag Callicarpa giraldiana var. subcanescens Rehd. in Sarg., Pl. Wils. 3: 360. 1916. Callicarpa grisea Hand.- Mazz., Anz. Akad. tt Wien Math.-nat. 58: 230. 1921. Callicarpa giraldiana subcanescens Rehd. apud Worsdell, Ind. Lond, Suppl. 1: 160. 1941. Bibliography: Lévl. in Fedde, Repert. Spec. Nov. 9: 455--)56 (1911) met 10: 439. 1912; Fedde, Repert. Spec. Nov. Gesamtverz. 58. : & Kew. Suppl. 5: 3. 1921; Hand.—Mazz., P . Sin. Forts. 1h: H - ,» Anz. . Wiss ien Math.-nat. on 230. 2 Mottet, ha ashe 9: O-—391. 19233 H. H. Chung, Mem. Sci. Soc China (2): 26. 1725, Saathoff, Gartenwelt 29: Sl. 1925; A. W. ah Suppl. 7: 36. 1929; Ptei, Mem. Sci. Soc. China (3): I-37, ss) 2 es Svenson, Brooklyn Bot. Gard. Record 22: 7. 19333 Rehd., Journ. Arnold Arb. 15: 321--323. 1933; Hand.-Mazz., Symb. a. 7: 900. 19363; Moldenke in Fedde, ae ke Spec. Nov. he 297, 303, & 30) (1936) and 40: 93-96. 1936; R ourn. Arnold Arb. 18: 2)3. 1937, Moldenke, Annot. es t 108. st Rehd., Man. Cult. Trees, ed. 2, pr. 1, 803, 80h, & 932. 1905 gpd Prelin. Alph. List Invalid Names 9-—~13. 1903 Moldenke, Known Geogr. Dis- & 86. 1942; Moldenke, Alph. List So. i: : tologia 2: 32. "19473 Moldenke, Alph. List Invalid Names Suppl. 1 3. 1947; H. Nw & en Re Moldenke, Pl. Life 2: a 58, & 61. oct Moldenke, Alph. List Cit. 2: 398, 108, 43h 560, 561, 577, 601, 617, “ 618 (1918), 3: 666, 742, omy & 90h. (91), ana i: 1olo, 1018, 1136, 1202, 1243, & 1288. 1949; Rehd., Bibl. Cult. Trees ~ 199; Moldenke, Known Geogr. Distrib. Verbenac., ed. 2, 130, 19493 Dict. Gard. 1: 358 & 359. 1951; . Blackburn, Trees & Shrubs East. N. Am. 103. 1952; sae Bg “riytol ogia k: 75. = Molden=- ke, Journ. Calif. Hort. Soc. 15: 1954; Moldenke, Résumé 167, iv, 21 213, 2h1, 243-245, 247, & ree 1959; Krtissmann, ae. Laub- geh. 1: 25); & *o5S, 19595 T. H. Everett, New Illustr. Sones Gard. 3 Rehd. . Cult. Trees, ed. 2, pr. 9, 80 & 2: 295. » 80h, 932. 1960; woldenke, Sask tects 20 03, 2a on Li, Mor- ris Arb. Bull. 1h: 3, bh, & 7, fig. 1k. 1963; sy prey yene lo- a 9: 43k. are se 5g Résumé Suppl. 13: 6 (1966) and 1): 3, gmt Le A9GG: Grimm, N. Y. Herald Trib., sect. 2, Dp. 20, Feb. 6. Tiss bea raga Ptei, Mem. Sci. Soc. China 1 (3): pl. 2. 19323 Li, es Arb. Bull. Lys h, ks 1—}. 1963. ge woody bush or shrub, 0.9--3 m. tall, or tree to 6.9 m. 50 PHYTOLOGIA Vol. 14, no. 1 tall, the pubescence denser throughout than in var. giraldii; setinian jong By 0.5-—3.5 cm. long, sharply sulcate, densely pubescent or pube rulent~ tomentellous; leaf-blades char ences or he eous, comparatively narrow, varying from elliptic or broad= ly elliptic, elliptic-oblong, elliptic-lanceolate, or vuntediat ) to oblong-elliptic, oblong=lanceolate A ovate~lance Olate, ovate, closely denticulate, dentate, or densely and broadly mucronulate- denticulate along the margins, brownish-green or rubescent above neath; midrib impressed above; secondaries 6-9 per side, flexu- ous~arcuate or oblique, branched, free, pubescent; tertiaries of- ten loosely reticulate, prominent beneath; inflorescence corym- bose, densely pubescent or densely tomentose throughout; pedun- cles’ slender, 4-10 mm. long, as long as or shorter than the pet- linear, - ripe mm. eke: flowers fragrant; calyx cupuliform r tur- binate, about 1 mm. long, subsessile, pilosulous or minutely hir= tellous to densely pubescent on the outer surface, subglabro within, the rim mimtely h-toothed with broad and obtuse or sable er obtuse teeth; corolla infundibular, pink to pinkish, lilac- pink, rose, red, purple, reddish-blue, or white, 22,5 mm. long, near. ame densely pubescent or pilosulous to glabrate or gla- brous outside, glabrous within, the limb slightly longer than th tube, the Lobes 1/3 the length’ of the limb, ovate, obtuse at the i Ss x5 ensely ipa very small, globose, 3-1), mm. wide, blackish-blue or violet te ea purple, or bright-purple, yellow when immature, pubes- cen pecies was based by Léveillé on a collection made by Léon ca Martin (no. 2365) in the environs of Gan-Pin on June 19, 1898, one by Léon Fr. Frangois Martin and Emile Marie Bodinier (no. 1966 Moldenke, Monograph of Callicarpa 51 1278) from among rocks at Grandes Rocailles on October 28, sp one by Pierre Julien Cavalerie (no. 1905) from Pin-Fa on oe: 1903, all in Kweichow, China. The he type of C. seguini was Peek lected by Pierre Julien Cavalerie (no. 231) in the environs of Tou-chou, ae China, in May, 1898, an and is deposited in the Bodinier erbariun. The type of C. feddei dded was collected by Joseph Henri a (no. 468) at Kouy-Tehéou, Kweichow, China, in June of 1905. The type pe of C. aldiana var. subcanescens was gathered by Augustine Henry (no. | 586) without precise locality in western j Ar- boretum at Cambridge, Massachusetts. The type of C. sea was gathered by Wang-Te-Hui [distributed as Handel-Mazzetti 1 182] around the coal mines at Pinghsiang, at 600 meters altitude, in | western Kiangsi, China, 1920 Ptei (1932) ” includes Ce lyi Lévl. in the synonymy of C. bodin- ieri, but I regard this as a distinct variety, C. bodinieri ‘eri var lyi lyi (Lévl.) Rehd. Rehder (1949) cites the original 1 publication ‘of C. feddei erroneously to page "320" instead of 439, and in his 193] publ a eevee places C. grisea in the synonymy of C. bodinieri baht Spectes has been found growing on slopes and hillsides, a- long roadsides, and in meadows, thickets, open thickets, and swamps, sometimes in clay soil, at altitudes of 200 to 2300 meters, flowering from May to September, and fruiting in February — aad to October. Ching calls it ee in Kwangsi; Wilson says tha "fairly common" in Kwangsi Yin Pessoa ee ee & as Pes sd which, of course, is quite meous . described as "pink" on Bullock 197, Ching ning 5365, and S tsang 27635 27835 & 28380; "pinkish" on Keng 1g 5955, ceeworeare on A. Henry 12119 and and Yin 102; "reddish-blue" on Tsang 277h0; "white and rose" on Handel- Mazzetti 8691; and "white" on Cimung & Sun 298. Li (1963) says that "The species is native to central and wes- rn China. It grows in thickets of the hilly districts but does a seem to be common, It flowers from July to September fruits in a — October. pee. oe is readily distin- guished by t. e margins 0 e leaves Rehder (i95h) in “wali aes soabatert of the oldest name for that very variable species generally known as C. Giraldiana and agrees with its pubescent form described as var. subcanescens. The type of the latter is slightly more pubescent than than Bodinier's no. 2365, one of the syntypes of C. Bodinieri eri and that which should be considered the t ectot; of the ies because it is from Bodinier's pattie ctype) of the species,» flowering material, sao and has the older leaves glabrescent, approaching those of Giraldii. Callicarpa Feddei hardly differs from the type ex- 52 PHYTOLOGIA Vol. li, now 2 cept that the leaves are narrower and smaller. The type of Ce Giraldiana var. subcanescens is somewhat more densely pubescent and agrees in this respect with C. Seguini. It seems hardly pos- 2 cee _ densely pubescent var. Lyi is more distinct partic- ty ts pubescent corolla." Léveillé's original description of C. seguini is "Affinis C. rubellae a quo differt foliis conspicue a nlaties affinis etiam C. Chaffanjoni a quo differt foliis subtu poseces sed dense to- mepiiets et inflorescentia tota esticas riginal descrip- of C. bodinieri is "Rami glabri, apic Sayin eo o-villosi; : Seams magna; folia 12 x cm. sarerks denticulata ovata utrinque attemata petiolata supra rubescentia subtus villoso reticulata; flores in cymas axillares, dichotomas, quintam folii partem aequan- tes prepa, sepala et petala pilosula; stamina exserta; stylus breviter emarginatus staminibus aequalis; fructus minimi jucunde violacei. © His ga te of C. feddei is "Rand griseo-rosei; folia ovato-lanceolata a nata, in petiolum sensim vel abrupte contracta denticulata, er glabra, brunneo-viridia subtus ful nervis flexnoso-arcuatis, ranosis, pubescentia; petioli pedunculi flores rubri; calice minimo hirtello, cupuliformi; corolla gla- blata; oe, Longe exserta, stylo ¢amen filiformi et apice bi- fido bre Rehder (1936) distinguished his C. giraldiana var. subcanescens what be now call Cc. bodinie ri var. giraldii as Ba 8 "A typo recedit indum ento omni iore, ramulis pube farinoso Tisvindeate atone obtectis, foliis supra pba et te fas asciculato-pilosis subtus densius’ praecipue in nervis et i 8 acacia: 3 the lea r art ane ois rather long fascicled hairs. The leaves scoala ogee Ee ay rag to silisticciumecteen e larger ones not exc 36 width, while those of wise No. 1528 are ore elliptic or ob- ovate and up to 6 road." Handel-Mazzetti's "original description of C. grisea is "Frutex (%) ramis hornotinis elongatis medullosis, totus pilis fasciculatis brevissimis cinereo-tomentosus, foliis subtus praeterea glandulis scutellatis de luteis adspersis su supra tantum calvescentibus. Folia hg ae 1.5-~16 cm. 1g. at 2——3 angustiora acuta vel anguste data (vel obhe ; basi contracta et in pet- iolum 10—35 mm. lg. argute 4 6 — id te mucronulato-denticulata, herbac supra impre nervis 6——9 nis obliquis arcuatis Uibelte vg trabeculis reueia Pon saepe venis laxe reticulatis a ae axillares d compositae myrianthae, pedunculis tenuibus usque ry 1 cm. Agies 1966 Moldenke, Monograph of Callicarpa 53 1.5--2 cm. lg. 3-- cm. 1t. densissimae irregulares. Bracteae ad mm. usque lg. lineares. Calyx subsessilis turbinatus 1 m. lg. intus glaber dentibus brevissimis latis obtusiusculis. Corolla rosea 2—-2.5 mm. lg. infundibularis intus et tubo extus glabra, Limbo paulo ores ad 1/3 in lobos ovato ubtuso fisso. Stamina ad 5 mm. lg. antheris 2/3 mm. lgis. late niiiptdele dorso parce et minute glandulosis, rima longitudinali dehiscentibus.....Flor- ibus C. Americanae et longifoliae et ningpotnsis, sed indumento pec ris." Li (1963) distinguishes this taxon from its immediate hag in popular cultivation as follows: On vegetative characters —~ Le eaves ve enpeainet the petioles 1--3 cm. long or about 1/3 01/5 t Brine of the crenate-serrate blades -C. americana. ac rated Pectin late, the petioles 2—15 mm. long or about 1/5 to tee the “longth of the serrate ors errulate to dentate or den 26 Psi arson gesetaatirs ae ee the blades glabrous to pubescent beneath. 3. Leaf-blades glabrous or nearly so, the margins serrate to serrulate more or less toward the central part only. h. sassrbiades elliptic or obovate, smaller, about 1.5-—3 . (rarely to 5 cm. or more) long, acuminate at the apex C. dichotoma. ha. fea Miro Bsn re: or ovate-lanceolate, lar arger and 2.5 cm. (sometimes to -. cm.) long, petal Posts at the apex C. japonica. 3a. Leaf-blades pubescent on both surfac the ee den= tate to denti bende: or eechaleciacit milake ppaleon 0b the whole lengt bodinieri. 2a. Leaf-bases rounded or subrounded; leaf-blades thickly stellate-tomentose beneath mollis. On ee ee characters he distinguishes the same taxa as follows oe Mince and fruit-clusters sxe or subsessile..C. americana. la. Flower- and fruit-clusters p 2. Flower— and fruit-clusters pce peduncles longer than 3. Fruit smaller, t 3--l mm. wide; anthers opening by a slit; cere meet generally supra-axillary.C. dichotoma, 3a. rents larger, about )--5 mm. wide; anthers opening by a nal pore; inflorescence axillary........C. japonica. 2a. Peon and fruit-clusters pubescent; peduncles about as long as the petioles |. Fruit bright-purple, about 3--l, mm. wide; rowers lilac, the stamens exserted.. podinieri. ha. Fruit dull-purple, about 5 mm. wide; flowers Tite pur plish-rose, the stamens Lluded......e..ee+eeeC- mollis. Material of C. bodinieri has been misidentified and distributed in herbaria under the names C. bodinieri giraldii (Hesse) Rehd., 54 Pet 201 667-4 Vol. 14, no. 1 C. giraldiana Hesse, C. giraldii Hesse, C. japonica Thunb., and C. longifolia Lam. On the other hand, the E. H. Wilson 116) and Herb. Univ. Nanking 1095, distributed as C. bodinieri, are actu- ally var. giraldii (Hesse) Rehd., while Fang 2529 is C. japonica var. angustata Rehd, P'ei (1947) records C. bodinieri from Sikang. In his 1932 work he cites the following: CHINA: Honan: L. H, Bailey s.n. [Chi- kung-shan, June 1917]. Hupeh: A. Henry 586. Kiangsi: Wang-Te- (Ed); L. F. Martin 2365 (Ed). The Cavalerie 1026 which he also cites is C. bodinieri var. iyi (Lévl.) Rehd. “He regards E. H. Wil- son 1319 and A. Henry 12119 as var. giraldii (Hesse) Rehd. In all, 54 herbarium specimens and 10 mounted photographs, in- cluding type or phototype material of several of the names invol- ved, have been examined by me, Citations: CHINA: Anhwei: R. C. Ching 320), (Ca——3)8070). Che- kiang: Chiao 106 [Herb. Univ. Nanking 15345] (Ca-——325372); Keng 595 (Ca—-361626). Fukien: H. H. Chung 2115 (Ca—-420358). Hupeh: Chun & Chien 5090 [Herb. Univ. Nanking 8065] (Ca—-261516); A. Senry 1185 (800128), 3017 (N), 3999 (W—800732) ; E. H, Wilson 439a (#—7770uh), 1319 (N, W-5967L3), 2196 (W—777382), 2213 (), Sun 252 (N), 298 (N) 182 (N—photo); E. H. Wilson 1528 (N--photo, W—-777228). Kwangsi: Ry C. Ching 5365 (N); Steward & Cheo 451 (Bz—1786, N); Tsan, 27TWO (W—1757182), 27835 (W—1757260), 27963 (W—1757361), 28 380 (W—1757681). Kwangtung: Tak & Chow 2788 [Herb. Canton-Chr. Coll. 1649] (Ca--319),13). Kweichow: Steward, Chiao, & Cheo 565 (N, S, W--159888)) ; Tsiang 5453 (S). Szechuan: Fang 1235 (N), 4648 (N, W~-1626995), 5719 (N), 8h02 (N)5 Yin 102 (N). Yttnnan: Forrest 1207) (S); Handel-Mazzetti 8691 (Diar. 1664] (W—1607241); A. (W-158868), 12119¢ (N); J. F. Rock mmany: Herb. Hort. Berol. s.n. [Sept. 1842] (B), s.n. [July 1815] (B), Sen. (Oct. 1846] (B, E— photo, N-—photo, Z--photo), s.n, (B, B, B, N—photo, Z—photo); Herb. Link s.n. (B, N—photo, S--photo, Z~=photo) , CALLICARPA BODINIERI var. GIRALDII (Hesse) Rehd., Journ. Arnold Arb. 415: 322. 193). Additional & emended synonymy: Callicarpa graiciane Hort. ex C.K, Schneid., Il]. Handb. Laubholak. 3: 1048, nom. nud. 1912. Callicarpa giraldiana Hesse, Mitt. Deutsch. Dendrol. Gesell. 21: 366, hyponym. 1912. Callicarpa giraldii Hesse ex Rehd. in L. He Bailey, Stand. Cycl. Hort. 2: 628 & 629. 1914. Callic longi-~ folia Hemsl. apud Rehd, in Sarg., Pl. Wils. 3: 306, in syn.(in > 1966 Moldenke, Monograph of Callicarpa 55 part). 1916 [not C. longifolia Benth., 1966, nor Blume, 1936, nor Hance, mer nor Hook., 1932, nor L., 1820, nor Lam., 1783, nor ahl, 1 Callicarpa mairei Lévl., Sert. Yunnan. 2. 1916. Cal arnoldiana Kelsey, Lriatcuite for the Garden 37, nom. nud. 1932. aa bodinieri var. giraldii (Rehd.) Rehd. ex earns in Fedde, Repert. Spec. Nov. 0: 93, in syn. 19363; Rehd., Journ. Ar- a bes 28: 2h3. i937. Callicarpa giraldii aii Reha. ex Moldenke Repert. Spec. Nov. 0: 93, in syn. ~~ es Alph. ned a apr Names 10, in syn. 1940. Callicarpa giraldii (Hesse) Rehd . ex T. H. Everett, N. Y. ts aan Fal Exch. List 1951: 5. 1950. Callicarpa bodinieri giraldi Mattoon, Pl. Buyers Guide, ed. 6, 83. 1958. Callicarpa giraldiana Pamp. ex Moldenke, Résumé 2h3, » in syn. 1959. Callicarpa bodenieri giraldi Rehd. ex Kelsey Nursery Service, Short Guide Cat. 167: 25. 1961. Callicarpa bodinieri var. giraldiana (Hesse) Rehd. ex Li, Morris Arb. Bull. 1h: 7, in syn. 1963. Callicarpa bodinieri giraldi Grimm, N. Y. Herald Trib., sect. 2, p. 20, February February 6. 1 Bibliography: Hemsl., Journ. Linn. Soc. Lond. Bo : 253. , Bot. Jahrb. 29: 548. Ill. Handb. Launholzk. 2: 1048. 1912; Hesse, Mitt. Deutsch. Den- drol. Gesell. 21: 366, 368, & 369. 1912; pend. in L. H. Bailey, Stand. Cycl. Hort. 2: *628 & 629. py ite: Rehd. in Sarg., Pl. Wils. 3: 366——367 & 369. 1916; Stapf in Curtis, Bot. Mag. 142: pl. 8682. 1916; Lévl., Sert Yunnan. 2. 1916; Lévl., Cat. Pl. Yun-Nan. 277. 19 9173 Prain, Ind. Kew. Suppl. 5: 3. a Ae Mottet ete -_ « My: 390391. 1923; W. J. Bean, Shrub 26 & 2. » Mem. Sci. Soc. China 1 (1): 228. cae. Stipp, 291 Bi. 1925; Leray, Rev. Hort. 97: 416. 1925; Rehd., Man. Cult. Trees, ed. 1, 775 & 776. eT} Cloud, Cult. Shrubs 121. 1927; , 148. 19265 Stapf, Icon. Bot. Ind. mdi. 1: 525. 19293 L. He = E. 2. tot us 111. 1930; Hottes, Fre Laubgehtlze, ed. 3, L11. 1931; a Mem. Sci. Soc. China 1 (3): [Verbenac. China] 16, 23, 31—3h, 36, & 55. 19323 Sal oes in Kelsey- jursery Cat. 1932: 37. 19325 gp eas Br Bot. . Record 22: 7. 1933; Junell, Symb. Bot. Upsal. k: 81 & 83. 193k; Rehd., Journ, Arb. 15: 322—-323. 1934; L. H. Bailey Flor. mac., 935; Patermann, ol. Verben- ac. 30, [55], & [56], pl. 3, fig. 23--27, & pl. 5, fig. 6 & 7+ 19353 Moldenke pert. Spec. Nov. 39: nee 03, & 304, hO: 9 » 98, 120, 6, & 131. a Sl Moldenke, Decorat. Pl. Winter "Months (aj. a Rehd., Journ. Ar= nold Arb. 18: 243. 19373; Hottes, Book Shrubs - 3, 168. 19373 Moldenke, Cult. Pl. 35. +938; Uoldenke, Geogr. Distrib. Avicenn. 56 FEY? O4..002a Vol. 14, no. 1 as Set Moldenke, Annot. & Classif. List 108. 1939; Moldenke, lim. Alph. List Invalid Names 9--11. 1 90; amie » Man. Cult. ed. 2, pr. 1, 80) & 932. 190; sae, orf Bot. Gard. » ahs Sin. 1: 3. 1947; Moldenke, Alph. List Invalid Names Suppl. 1: ge 1947; He No & A. L. M Moldenke, Pl. Life 2: 61. 1918; Moldenke, Alph. List Cit. 2: 398, 08, 43 h, 514, 563, 601, 617, & 618. (1948), 3: 666, 7h2, air. &'90h (19495, and h: 1010,"1018, 1136, 20 fe . 1949 é 9 anta J. Bean, Trees & Shrubs Hardy Brit. Isles, ed. 7, 1: 333. 1950; T. H. Everett, N. Y. Bot. Gard. Seed Exch. List 1922: : 360. 39. W. 358 & 3 19a s Bs Oy arereis: Trees & Shrubs East. N. Am. 103. 1952; Mol- denke, Journ, Calif. Hort. Soc. 15: 80, 198i; Slettcon: Pl. Buyers Guide, ed. 6, 83. ; Krtlssmann, Handb, Laubgeh, 1: “25h. 19595 Moldenke, Résumé 167, 17h, 213, 21, 243, 2h, & 43. 1959; Rehd., Man. Cult. Trees, ed. 2, pr. 9, 80) & 932. 1960; Moldenke , Résumé 7 - 196 i rri Bull. 1h: ustrations: Hesse, Mitt. Deutsch, Dendrol. Gesell. 21: 368 & 369. 1912; Stapf in Curtis, Bot. Mag. 142: pl. 8682 [in color]. 1916; Mottet, Rev. Hort. 9h: opp. 390 [in color]: 19233 Stipp, Gar t 29: Si. rhe Leray, Rev. Hort. 97: _ = color]. 1925; Pate termann, Beitr Zytol. Verbenac MAC » pl. a § Ze 23~-—27, & pl. 5, fig. 6 : ve 1935; Silva Pe cry tomer: & Schneid., Insere Frei- land-Lau “ge Reap 931; Li, Morris Arb. Bull. 14: ) & 3 This variety differs from the typical form of the species i having its leaf-blades glabrous above and sparingly fascic sabes and the with resinous golden aa tes are hidden on more densely stellate~pubescent lea Recent collectors Salis this plant as a small woody or bushy slender shrub, 0. m. tall, a bush, a much-branched herb, or to co . bove, pale neath; peduncles purple; flowers small, e@ in bud; corolla me inkish, Si perm or SS oe Pink to purple, senapnicgibe ed, 1966 Moldenke, Monograph of Callicarpa 57 This plant has been found growing in sandy soil, in partial o dense shade, in woods and thickets, open woods and open tate. i d b. 0 to 3 meters altitude, blooming in February, May to August, = October, and fruiting from July to November. Wilson records it as "com- s ang: in dry silt" in Kwangtung; Chin calls it "common" in Anhwei, while Ching refers to it as "rare" in the same province. Ching likewise describes it as both "common" and "rare" in Chekiang and as a "fairly common shrub in thickets" in Kwangsi; Ching & Tso refer to it as both "common" and "rare" in Ki gsu ancock says "clusters fof fruit] white at “aug fi meats 2 considered of ornamental value". It these fruits mature, his rey = C,. americana L., C. dichotoma (Lour.) K. Koch, and C. japonica Thunb. Ching [on n no. 363] describes the flowers as is purple, but doubtless intended th this Limpen hi Syot to apply to the fruit, since also described a "purple" on Hu 1621 and on Rock 9195. They are described as une on Hu 28), "pinkish" on Ling ing 1213, "pinkish- purple" on Chun 3593, "reddish-purple" on Chung & Sun 20h, "rose- purple" on Wilson 1531, "purplish-pink" on Ching & Tso 1318, Ek Bie th on | Ling 2565 » "red" on Tsang 100 ig 100, “and | "yellow" on on 1319 Rehder (1916), in speaking of what he then called C. giraldia- nha, says "This species seems to be most closely related to C. for mosana Rolfe and C. ja ponica Thunberg; the first is easily dis- tinguished by the he denser pubescence, the longer peduncles which exceed the petioles, and by the rounded or obtuse base of the leaves; C. japonica differs in its glabrous narrower leaves, smalle rous inflorescence on a peduncle almost as long as the petiole and in its longer anthers opening at the apex with a pore. It is also near the Indian C. psilocalyx Clarke, which differs chiefly in its densely pubesc: scent branchlets, in its long-acuminate — Sep obtuse or rounded at the base, shorter petioles, inflorescence, and in the filaments scarcely exceeding the athe toe Wilson's Nos. 439, 139e, 2193 and 2196, Giraldi's Nos. 1360, 1361 and 5896 and Henry's No. 5992 agree very well with rtain. Von Rosthorn's No. ae is more pubescent and approaches var. Eecceeaener? edigaret reas the Arnold pe tum herbarium, the following: CHINA: 2992, 7312, 71973 Silvestri iol, 1950, 0, ase. Pgh. ABs E. H, a? a ine Gee sates eee 58 PHYTOLOGIA Vol. 1), no. 1 Wilson 439, 39a, 2193, 2196. Kiangsi: E. H. Wilson 1531. Shensi: Giraldi 13 1360, 13 1361, 58 5896, son. [Tu-kia-po, October 1897]. Szechu- an: an: Von Rosthorn rn 1896, E E. H. Wilson 219k, 5100. Yttnnan: A. Henry 12119, | 12119a, Tll9c. “CHINESE COA: COASTAL - ISLANDS: Hainan: A. Henry 6127. CULTIVATED: Germany: Hesse s.n, [Weener, August and Octo- ber, 1910]; Kache os [Weener, July and October, 1912]; Rehder S.n. dT ocseind Ma July 191 2) The A. Henry 12119, 121192, & 12119¢ - i. Wilson 19a & 2196 which he cites I regard as typical c. boinbnt Ptei (1932), also speaking of C. giraldiana, says: "This spe- cies is allied to Callicarpa formosana Rolfe, and C. longifolia e pee and from the latter by its me flowers, scarlet gl and relatively broader ovate leave Li (1963) says that this variety "is ste to central, western southern EE : =e ry) A. [or i 3, sparingly cent beneath. The inflorescence is also less pubescent. The teeth along the leaf margins are oe ger gee and not dis- tinctly dentate as in the typical forn The C. longifolia of Bentham and of “tao, referred to in the synonymy above, belong to the eet of C. longissima (Hemsl.) Merr.; that of Blume, Linnaeus xburgh, and Vahl is C. longifo- lia Lam.; that of Hooker is c. ‘warps (Benth, ) Hance; and that of Hemsley and of Diels is in part G e japonica var. angustata Rehd. The Leray (1925) reference igen in the MR a above is sometimes incorrectly written "96: 116," cultivation, according to Rehder (1940), C. bodinieri var. Siraldii blooms from July to September and erate in September and Octo October. He avers that it was introduced into cultivation in and), and ill Nursery oking, Surrey, England). The See Nursery Service (1961) of- fers 2 1/2 inch potted plants a She o xate of 2 for $3 or 10 for $8.50. Mattoon (1958) lists ten s 1966 Moldenke, Monograph of nice 59 Material of C. bodinieri var. giraldii has been misidentified and distributed in herbaria under so names C. dichotoma (Lour.) Cc. ieee: Cc. Tees Lam., ¢ rubella Lindl., and C. sinensis Hort. On the other hand, the R. C. Ching 3204 & 5365, Chung 2145 2145, Fang 6)8 & 802, A. Henry 12119a 12119a, Herd. U Univ. Nanking 1095 1095, Ip 1809, J. F. C. Rock 2523, Steward, Chiao, & Cheo 565, and E E. i Wilson i,30a, 1319, & 2196, 7 as C. bodinieri var. . gir aldii, are all typical C. C. bodinieri Lévl.; W W. XY. Chun 240 [Herb. Univ. Nanking 566] & s.n. [Herb. Univ. Nanking 5805] are C. can- os var. sumatrana (Miq.) Moldenke; Chang & Metcalf 197 & & 216, Chin ng 1638 1838, H. H. Chung 231k, 2677, & 3021, and | and Herb. gurton Sie “Goll. 12008 are e. ee nates Chiao = (Herb. | Univ. ilneied” sideman «Sanat See Fad Bi Pane aca ane Candee var. angustata Rehd.; J. F. C. Rock 9195 and E. H. Wilson 439 are Cc. Japonica var. rhombifolia H. J. Lam; Herb. C. Canton Chr. Chr. Coll. 12560 and Peng, Tak, & Kin 561 are C. longipes Dunn; and Tak 850 (Herb. Lingnan Univ. 16349] is isa cotype collection of C. rubella f. robusta Plei. Prei 1738) cites the following additional specimens: CHINA: Anhwei: C 2761, Ching 2290. Chekiang: Cheo & Wilson s.n. [Mo- kan-shan, Ju at 1926], Ch Chiao 106, R. C. Ching 363h, 4927, 4987, a Ra Sen. [Vicinity of Ningpoo, 1908], Tang & Haia aia 30. F Fu- kien: Chen . 3080. Honan: Hers 2574. Hupeh: Chun 3620, 3682, 4392, A. Henry 3107, 7497, 71117, Ke Ling s.n. [Ichang, 1926], , Silves- tri 198, 1950, 1952, 3158, . H. Wilson 2193. Kiangsi: Hu 20h, Ip s.n. s.n. n. (Lushan, ~ June 1921, greene 22h, E. H, Wilson ion. Kiangsu: R. C. Ching 1,863, L872, Ching & Tso 598, 650, 1310, ), 1328, Chun 2000, Nex N. 1095", Hancock SMe (Shanghai, Sept. 1925]. Kwangsi: | Re Cs Ching 639. Kwan Kwangtung: "ex C.1751, 3353", A. Henry 6127. Kweichow: Handel-Mazzetti 2054. Shensi: Giraldi s.n. [Kin- gua-shan, July 1897]. Szechuan: Bock & Von Rosthorn s.n. [1896], W. P, Fang 2061, 6423, E. H. Wilson 2194, 5100. THIBET: Forrest 1926, 1926. The R. C. Ching 320k, J. F.C. Rock 2523, and EB. H. Wilson 1319, which he cites, are regarded by me as typical C. - bo~ dinieri Lévl., and the Barchet 557, which he cites, I regard as C. japonica var. angustata R In all, 96 herbarium specimens, including type material of most of the names involved, and 13 mounted photographs have been exam- 60 PEI T OL:0074 Vol. 1h, no. 1 ined by me. Additional citations: CHINA: Anhwei: Cheng 1,008 (W--167h272); K. Ling 1127 (Herb. Univ. Nanking 7717] (Ca—258805), 123 [Herb. Univ. Nanking 779k] (Ca—-259031); Sun 1176 (N), 1260 (N). Cheki- ang: Cheo & Wilson 107 [Herb. Univ. Nanking 12753] (Ca--316)00), s.n. [Herb. Univ, Nanking 12753] (Ca-—35l,682); Chiao 1022 [Herb. Univ. Nanking 14321] (Ca—325396), s.n. [Herb. Univ. Nanking 14321] (W—126777); R. C. Ching 1652 (Ca—281671, W--126536), 3634 (Ca—343153); Hu 1621 (Ca—362323); E. D. Merrill 11298 (Ca— 992129, N). Fukien: c, H, Cheng 3080 (Ca—299362) ; L. XY. Tai 11793 (Ca-—325796). Hupeh: Y. Chen 736 [Herb. Univ. Nanking 15086] (Ca-—343481), 2009 (Herb. Univ. Nanking 15303] (Ca—3h3537)3 Chow 595 (N), 766 (N), 1570 (N), 1805 (N)5 We Yo Chun 3593 (W— 1506373), 3965 (W—1279530), bo2k (W—1279531), 4320 (Gg—2342hb); A. Henry 5992 (W--801)88), 7312 (W-598666); E. H. Wilson 439 (Ge 31995, Ge-~31997), 39a (Ge—36135), 1164 (N, W—596841), 2196 (Gg--31996). Kiangsi: Chung & Sun 20) (N); Tsiang 1005, (N). Ki- angsu: M. L. Hancock s.n, [Herb. Univ. Nanking 12075] (Ca—-29),8h9)3 Herb. Univ. Nanking 1095 (Ca--230385, Io—11,023, W--1279692); Keng 1728 (N); K. Ling 2565 [Herb. Univ. Nanking 12515] (Ca— 31600); Shan 1095 (Mv). Kwangsi: R. C. Ching 5365 (N), 5666 (N), S39h (N, W--121867h). Kwangtung: Peng, Tak, & Kin 678 (Herb. 3 Tsang 20717 (B, Bz—17603, Canton Chr. Coll. 12677] (Ca--275059) N), 20797 (N). Kweichow: Steward, Chiao, & Cheo 565 (N). Shensi: Giraldi s.n. [Monte Kin-qua-san, 10.V11.1897] (Ga—-38200—iso- type). Szechuan: W. P. Fang 2529 (N), 4648 (Du—252638, N), 802 (Du—252693, N). Yttnnan: J 1332125). Province undetermined: W. T. Tsang 850 [Herb. Lingnan Univ. 16349] (I). CHINESE COASTAL ISLANDS: Hainan: How 71071 (B, N)5 We T, Tsang 100 (Herb. Lingnan Univ. 15599] (N, 3, W—12498L5) - JAPAN: Enoshima: Sawada s.n. (?rovince Sagami] (Ca--382277). CUL- TIVATED: California: McClintock s.n. [Nov. 16, 1959] (Gg—-l188L9, Gg--,18850). Germany: Herb. Kummer s.n, (Hort. Bot. Monac., Aug. 1848] (Mu—1)0, N--photo, Z—-photo), s.n. [Hort. Bot. Monac., 20. X.1855] (Mu—1))1), Sen. (Hort. Bot. Monac., 15.VII.186l] (Mu-- 1439, N--photo, Z—photo) ; K. Koch s.n. (B, N--photo, Z—photo). Massachusetts: E. J. Palmer 6712-1 (N); Rehder s.n. (Wilson 39 seed] (Ur); Cc. K. Schneider s.n, [Wilson 633 seed] (Ar—-19786)- MOUNTED ILLUSTRATIONS: Hesse, Seite 98 (Cp); Stapf in Curtis, Bot. Mag. 12: pl. 8682. 1916 (Cp, Oa). CALLICARPA BODINIERI var. LYI (Lévl.) Rehd., Journ, Arnold Arb. 18: 23. 1937. h abroad Callicarpa lyi Lévl. in Fedde, Repert. Spec, Nov. 10: 39. 1912, me 1966 Moldenke, Monograph of Callicarpa 61 Bibliography: Lévl. in Fedde, Repert. Spec. Nov. 10: Big let Fedde, Repert. Spec. Nov. Gesamtverz. 58. 191); Lévl. Tchéou 39. 1915; Prain, Ind. Kew. Suppl. S: 3. 1921; sre on oh Sec. China d Ar ~~ Be @ @s . Q B. #2 lo Ut ao | ‘Oo w ns) we ww 2. a . 7 cq, z J . ourn. e im. Alph. List Invalid Names get Borer Moldenke, Known Geogr. Distrib. Verbenac., ed. 1, 56 & 86 ree See EE ‘Alph List gens Names 10. 192; Moldenke, erbena ed. 130 & 177. ie Meldenke, puytaiiers is 7. 19825 aiaes.” ke, suas 167, 245, & 1959. er (193k) sas of Se taxon: "This at first glance looks es with the grayish white tomentum of the caady nas “of at spore of the stem and the inflorescence in- cluding the calyx and corolla, but in all other characters it a- grees with typical C, Bodinieri. By P'ei it is referred “a Cc. Giraldiana var. subcanescens as a synonym, but is easily distin- guished by the much denser whitish pubescence, particularly the upper part of the corolla is fairly densely pubescent outside, while in the type only scattered stellate hairs are present or often ey lacking." ecent collectors describe the plant as a shrub, 5 féet tall, with aril e-red flowers, growing on brushy slopes, "at 720 meters altitude, fruiting in June and July. The type of the variety was collected by Pierre Julien Caval- erie (no. 1026) at Pin-Fa, Kweichow, China, on June 3, 1903, and is deposited in the herbarium of the Royal Botanic Garden at Edin- burgh. P'ei (1932) regards it as Sid ical C. bodinieri (which he calls C. giraldiana var. subcane Léveillé's original (1912) deseription is: "Ramuli pulverulen- ti; totia ampla usque ad 18 x 7 om. petiolata, petiolo pulveru- lento obovata acuminata ad basim mite attenuata supra pilosa a subtus cinereo-tomentosa, sub lente y eeesiaoel inconspic- ue denticulata; flores composite coryubosi, rubri; pedun- culi, pedicelli et calyces cine soci mins calycis dentes breves cee, corolla calyce 2=plo longior; st longe exse Species indumento cinereo-tomentosa et colore Pisa Rehder (oak) places C. grisea Hand.-Mazz. in the synonymy here, but I regard it as typical C. bodinieri Lévl. Besides citing an isotype of C. grisea, Wang-Te-Hui s.n. [Handel-Mazzetti a ‘ ae the Arnold Arboretum herbarium, he cites the following: nam-Hupeh border: L. H. Bailey s.n. [Chikungshan, June 18, ae. . Kweichow: Cavalerie e 1026 (Am (A—-merotype, sais 2 Szechuan: W. P. (A). Fang 1235 Only 2 herbarium specimens and 2 mounted photographs have been examined me. Citations: CHINA: Hunan: Fan & Li 262 (Bz—-1729, N, N—photo, Z=~photo) . 62 PHYTOLOGIA Vol. 14, no. 1 CALLICARPA BODINIERI var, ROSTHORNII (Diels) Rehd., Journ. Arnold Arbe 1 23. 193k. Syno: rs 15 Galiicacs longifolia var. rosthornii Diels in Engl., Bot. Jahrb. 29: = 1900. Callicarpa giraldiana var. rosthornii Rehd. in Sarg., Pl. Wils. 3: 367. 1916. Callic giraldiana var. rosthornii (Diels) Rehd, ex P'ei, Mem. Sci. Soc. China 1 (3): [Verbenac. China] 37. 1932. Bibliography: Diels in Engl., Bot. Jahrb. 29: 548. 1900; Rehd. in Sarg., Pl. Wils. a 367-~368. 1916; Mottet, Rev. Hort. 9: 391. 1923; H. H. Chung, Mem. Sci. Soc. China 1 (1): 225. 192h; Pied, Mem. Sci. Soc. ae rae [Verbenac. China] 16 & 37. 1932; Rehd., Journ. Arnold Arb, 15: 323. 19343 Moldenke in Fedde, Repert. Spec. Nov. 40: 95 & 98. 1936; Moldenke, Prelim. Alph. List Invalid Names 11. 1940; Moldenke, Known Geogr. Distrib. Verbenac. ° & 86. 19125 Melee, Alph. List Invalid Names 10. 192; Moldenke, gr. Distrib. Verbenac., ed. 2, 130 & 177. soho: Moldenke, Résumé 167, 243, 2h5, & 43. 1959 This variety differs from a typical form of the species and from var. giraldii (Hesse) Rehd. chiefly in its smaller and nar- rower leaf-blades, which are very gradually narrowed into the pet- iole, in the denser pubescence, and in the somewhat longer anthers. The type of the variety was collected by A. von Rosthorn (no. China, in August, 1891, and was deposited in the herbariis of the Botanischer Garten in Ber: lin, but is now destroy Diels' original description (1900) of this tase is "Foliis membranaceis utrinque crebre glandulosis subtus praeterea stellato- subcinereis Sieaciets vel obovato-oblongis basin versus lon- ime Blttter (mit Einschluss der stielartigen Basis) 8--10 x ae Cisse Dure a r bisher beschreibenen Formen des Kreises zu erkenn P'ei (1932) describes the plant as "A shrub; snus pubes- ent with stellate hairs. Sows oblanceblate: or Pig rice ties ate at the apex, tapering at the base, chartaceous, about 7 lateral nerves on each side of the midrib. Petiole 0 rs to 1 cm. in length. Calyx truncate, densely pubescent without. Corolla Rehder (1934) records the variety from Kwangsi and cae % has been collected in flower in August. The Chi 5666, tributed as Cc, ee Bag var. rosthornii and cited by Plei (as32), is actually Gs japonica var. angustata Rehd, I know no nothing about this taxon exc except what is stated in the literature quoted above, 1966 Moldenke, Monograph of Callicarpa 63 oar ype hoe emg Moldenke, Phytologia h: 285--286. 1953. Bibliography: Moldenke, Phytologi ah: 285--286. sep Woldenke, Biol. abet: 27s 3121. 1953; Moldenke, Résumé Suppl. 1): h. 1 stems coarse, very conspicuously tetragonal with the nodes; nodes not annulate; principal internodes 6--9 cm. long; leaves decussate-opposite, very large, sessile, somewhat amplexi- caul at the base, obovate, thin-chartaceous, gray-green on both midrib on both surfaces, often densely ous above, abundantly sinciporsegtaprear zai som the margins numerous, ° ies, obscure above, prominent beneath; veinlet eee abun- dant, obscure above, prominulous beneath; Antiors ence ary, tomentellous throughout; peduncle es, tationes-ans tetee Ss, and pedicels very slender, the pedicels much elongated, 7—-15 mm ong; calyx campanulate, canescent—puberulent throughout, with a few larger branched hairs near the >t about : = long and e, i minu S-to 3c and s not well —— developed for Seoort ston: pistil saabealn one; style inal; np ere me The type of t very distinct species was collected by F. _ Endert (no. eg at W. Koetai in central eastern Borneo, at altitude of 1100 meters, on September 29, 1925, and is ani in the Herbarium Bogoriense at Buitenzorg. Unfortunately, the infl nfe th fungus and the Mr. Joseph V. Monachino, however, has examined the plant careful- ly and agreed with me that it is probably verbenaceous, and, if so, then probably a species of this gemus. It was also sent to me by the curators of the Buitenzorg herbarium as verbenaceous. The species is known thus far only from the type specimen; only me. Citations: INDONESIA: GREATER SUNDA ISLANDS: Endert 3723 (Bz-- 72607—type, N—-photo of type, Z--photo of tore CALLICARPA BRACTEATA Dop, Bull. Soc. Hist. Nat. Toulouse 6): 507—~ 3 Bibliography: P. Dop, Bull. ons Hist. Nat. Toulouse 6): lb 508 & 512. 1932; P. ee in Lecomte, Fl. Gén. Indo-Chine psc 1935; Worsdell, Ind. Lond. Saris wis 160. 191; Moldenke, istrib. Verbenac. 58 & 86 (192) and ed. 2, eee 277. 1993 Moldenke, résumé ex & Pbrase 1959. ustrations: P. Dop in » Fl. Gén. Indo-Chine :799. 1935 BOOK REVIEWS Alma L. Moldenke "Flora — the Prairies and Plains of Central North America" Per. Rydberg, vii & 969 pp., illus., reprinted for the N vor Botanical Garden by Hafner Publishing Company, New York & London, 1965 [Facsimile of the original edition of 1932]. $12.00 For quite some time this valuable work has been out of print and yet still in demand. The offset process has made its reap- pearance possible as it has done and will continue to do for sev- sre gems in several fields. Dr. Rydberg planned this flora as a complete manual of the Spermatophyta and Pteridophyta of Kansas, Nebraska, cee aaa! ta, the Dakotas, southern Manitoba and sctindistern katc extension it also includes most of the species hy the pie gs regions of Illinois, southern Wisconsin, northern Missouri, and in the plains regions of eastern Colorado and eastern Montana. It = new in its field and nothing has surpassed it since. Of urse new plants have — recorded for this area since the book Bay a and there have been re inden ural changes in this nterval. sian 8 more t "Flora of Missouri" in- pete some of these changes for phe oneanite é areas. "Illustrations to the Flora of Delhi" by J, K. Maheshwari, xx & 282 pp., illus., Council of So he and Industrial Re- a New Delhi, India. 1965. Rs. 28.00 or Sh. 56 or 00 . ‘ spe =e a represent 206 genera and 66 families. The book contains a few pages of text addenda and nomenclatural changes "Guide to Pig Soot of the Everglades National Park" by Alex D. O pp., illus., Tropic Isle Publishers, Inc., Coral Gables’ Florida. 1965. $2.00 Choosing the more immediately apparent and the more spectacular indigenous plants from this unique 2300 square mile area, the auth- or has descri illustra x bAd ee Tae F 4 ail 7 [o7) a te photo- graphs. This is indeed a lovely souvenir for the visiting — as well as for the general tourist in the south Florida area 64 PHYTOLOGIA Designed to expedite botanical publication Vol. 14 December, 1966 No. 2 CONTENTS SWALLEN, JK... Notes on presses 2 0 gs i veo te ee MOLDENKE, H. N. eee, materials toward a sel Seda of the genus Callicarpa. IV go oie epee we : MOLDENKE. Ai. Book seve 6 2 NOTES ON GRASSES Jason R. Swallen The following notes have accumulated during recent years. They should be en as notes and not as re- visions of the groups concerned. Retirement has given me the time and opportunity to prepare them for publication, Parvifolia Group of Panicum First glume broad, 2/3 the length of the spikelet, Spikelets 2 mm. or more lon NZ. First aioe S-nerved. Blades glabrous or nearly so. Blades narrow...secesecee Ll. P. carannasense Blades broad...eece 2. P. caaguense Blades pilose on both surfaces. P. errabundum First glume 3-nerved. Blades sons about 3 mm. wide, linear, stiffly spreading, glabrous.. . P. wettsteinii Blades mostly f- 5 mn. wide, shorter, pilose on both surfaces. - P. pseudisachne Spikelets less than 2 mm. long. Spikelets 1.7-1.8 mm. long. P erect. Plants annual, 6. P. kaietukense — Plants perennial from rhizomes. arum Plants decumbent-spreading, wither rhizomes; blades cordate or subcordate, pubescent. Blades 1-2 cm. wide, pubescent. - P. semitectum Blades less than 1 cm. long, glabrous. 9. P. helobiun Spikelets 1.2-1.5 mm. long. C mostly erect, at least not decumbent- spreading. Culms stout; blades more than 5 mm, wide, Blades mostly not over 7 mm. wide, not clasp cyanescens Blades as much as 15 mm. “wide, the base of the blades conspicuously clasping. =.¢ ee Culms slender, the blades distan Blades narrow, at least some of them inv olute; spikelets 1.6 mm. long. 12. P. orinocanun 65 PHYTOLOGIA Vol. lu, no. 2 Blades broader, not at — involute; spikelets 1.) mm. long. 13. P. vimnulun Culms slender, decumbent or cre wane Blades 1-3 cm. long, usually spreading or reflexed, glabrous; culms nearly simple. 1h. PB. parvifolium Blades much longer, pubescent or pilose on both surfaces; culms with short dense branches from the upper nodes. P. egleri First glume sional narrow, usually acute or sub- acute, rarely more than half the length of the spike 3 el Spikelets pilose. maior eve oS; 8-1 mm. lon ng. Ligule arcuate; culms about 30 cm. tall; blades appressed, 5 mm. wide. 16. P. subinclusum Ligule straight; culms not more than 20 cm. 1, usually much less; blades samemaing: Culms Seelge ed branching, not more than 1 @ll..... 17. P. obovatum Culms svaniy branching gies all the nodes. 18. P. froesii sidgesier ae 1 mm. lon > y partly enclosed in the sheath; ales on short stiff pedicels. 19. P. siccaneun Spikelets 1, — mm. long. Ligule stra 8 aemsiic. long and tangle ed. P. fonticolum Culms erect or decumbent, not creep oe Fic seca 2 - P. tam amayonis Ligule arcua Annual; i tufted, erect or spreading at the b Blades tigsee. not more than 1.5 mm. wide. P. gracilissimum Blades _ onan ovate. or ovate- eolate, up mm. wide. vive ne cag narrow, gn ee remote. graniticun First glume broad, not remote. - P. pandun Perennial?; culms Senueh aes ocecadtne or creep Culms very slender, widely creeping, becoming tangled, 2 - E. yavitaense 1966 Swallen, Notes on grasses Culms relatively slender, decumbent at the base only. Panicles small, rather dense, 1-2 em. cE pele at aay. - 26. P. petrense Panicles very open, up to cm. long. CDR nash ae simple; spikelets 1.1 mm. ONGececesee 27. P. mauryi Culms freely branching; spikelets 1.3- mm. long. 3 20. P. goeldii Spikelets glabrous. i ata Spikelets 0.9-1 mm. long. Blades involute, soft and fine; culms decumbent- spreading, branching, 29. P. petilum Blades flat. RS ants annual, Culms with ahoge branches from the node; panicles to 2 ecm, ag about as wide as hae. Sar, ol Culms simple or with a or "two branches; panicles as much as 8 cm. long, much longer than wide. 31. PB. arctum Plants perennial, igule arcuate; blades elliptic-lanceolate. “Gulns erect or ascending; panicles relatively larger, ss prea Foliage glabrous or nearly s 32. FP. grantliferm Foliage conspicuously pilose or hispid, luish-glaucous, 33. P. spissifolium Culms widely creeping; panicles small, mpprrreg partly enclosed in the shea 3h. P. kuhimanni Ligule peta yw blades linear 5. P. kappleri Spikelets 1.2-1.); mm. long. Culms very slender, creeping, becoming tangled. P. blackii Culms erect or decumbent at the base, not creep ants e Lateral spikelets with pedicels as long as r shorter than the spikelets. yas ae aniticum Lateral spikelets with rine at much longer than the spikelet P. angulosum Plants perennial Ligul e straight. Blades about 1 cm. long crowd 8. P. densifolium OGsevewes ss 68 FRT'TOLOGIA Vol. 1h, no. 2 Ligule arcuate. Blades linear, 4-5 em. long, 2 mm. wide. 39. P. micranthun Blades een ic, )-5 mm. wide des thin, lax, spreading, sparsely pilose or nearly glabrous; spike- lets pointed, the second glume as long as the fruit. d r e Blades oct appressed, glaucous, gla- rous; second glume rere shorter than the oo yl. » politii : ae ee riage vig epee Mez, Notizbl. Bot. Gart. Bm ft Toe wees micot enue Minas Gerais, near Carannas, Sellow 20h. 2. PANICUM Segre Henr. Med. Rijks. Herb. Leiden h73 2. Type: poten Caaguayu, Balanse 22. 3. aie ERRABUNDUM Hitche. Contr. U. S. Nat. Herb. hs Fe BS SY 7922. eige “Briti sh Guiana: Parika, Hitchcock 16817. its coraaase WETTSTEINII Hack. Ergeb. Bot. Exped. Akad. iss, Sudbras. 12. 1906; Denkschr. Akad. “Wino. io Naturw. (Wien) 79: 73. 1908, P. rectissimum Mez, Notizbl. Bot. Gart. Berlin 7: 73. 1917. Brazil: Glaziou 17955. Type: Brazil: =o Paulo, near S. Bernardo, Wettstein & Schiffn 5 Ss eepne: enaneny al Notizbl. Bot. Gart. Lin 7: 71 eodas * Beaultt Sexton 13330. 6. PANICUM KAIETUKENSE Tutin, Journ. Bot. Brit. & Ter 30. fe. 11. 193. Type: British Guiana: “Kaietuk" savanna, Potaro River, Tutin 688, 7. PANICUM SAVANNARUM Soderstrom, Mem. N. Y. Bot. Gard. 12(3): 2. 1965. Type: British Guiana: Essequibo District: Kaieteur Plateau, Cowan & Soderstrom 2195. 8. PANICUM SEMITECTUM Swallen, nom. P. emergens Doell in Mart. Fl. oe os. 2(2): 269. 1877. Not P. emergens Hochst. Flora 38. Type: Brazil: S&o Paulo, Riedel 1961. 1966 Swallen, Notes on grasses 69 9. PANICUM mien aoe os ex oe Med. Rijks. Herb. den. 0: inde Argentina: Nie hocite Loreto, Ekman 650, 10. PANICUM CYANESCENS Nees in Trin. Gram. Pan. 202. 1826; Agrost. Braz. 220. 1829, Type: ny, spp, Brazil. (Link. Mertens)." 11. PANICUM NERVOSUM Lam. Encyel. lh: 717. 1798. e: French Guiana: Isle de Cayenne, Desfontaines, 12. PANICUM ORINOCANUM Luces, Journ, Washington Acad. Des. Ser .oe tO. 3913. Type: Venezuela: Alto Orinoco, Puerto Ayacucho, Ll. Williams 13090, £3 PANICUM VINNULUM Swallen, sp. nov graciles, erecti, hO-50 cm. alti; ligula arcuata 0,3 mm. longa; laminae anguste adscendentes vel appressae, 2.5-l1.3 cm. longae, 2.5-3 mm. latae, acuminatae; parrot L.Seiel mm. DEES longe- or subobtuse, 3-nerved; second glume equal, just covering the fruit, 5-nerved, the ra tp lemna highae a staminate flower; ’ Ppa the U. S. National Herbarium, No. 1961151, collected Ha banhado, Estag&o Experimental, Ponta Grossa, anf, Brazil, efi hepa 31, 1946, by Jason R. Swallen (No, 837). 14. PANICUM PARVIFOLIUM Lam. Tabl. Encycl. 1: 173. z Type: "Ex Amer. merid. communis. D, Richard." 15. PANICUM EGLERI Swallen, sp. nov. erecti vel decumbentes 30-40 cm. alti ramosi; ligula brevissima, arcuata; laminae Bi Pac 5 cm. longae, 4-5 mm. latae, adscendentes, pilosae ve Slabrae; spiculae 1.5 mm. longae, glabrae, real setdis brevis rigide divergentibus; gluma prima spicula 1/3 r. brevio Perennial; culms erect or decumbent, reddish, 30-40 em, long, branching; sheaths of the main culm 70 PHYTOLOGIA Vol. 1h, no. 2 mm. ly glabrous on both surfaces, the leaves of the branches reduced; panicles shortly exserted, 2-l, cm. lo with a few widely spreading solitary branches; spikelets 1.5 mm. long on relatively short, stiffly spreading pedicels, glabrous; first glume 2/3 the length of the spikelet, broad, triangular, subacute, 3-nerved; second glume and sterile lemma equal, just enclosing the fruit. e in the U. S. National Herbarium, No. 2307286, collected at Rio Cururu, Tapajés, Par&, Brazil, July 17, 1959, by W. A. Egler’ (No. 9213. 3 16, PANICUM SUBINCLUSUM Swallen, Mem. N. Y¥. Bot. Gard. 9: 260. 1957. Type: Venezuela: Amazonas: Rio Atabapo, Bassett Maguire 29271. § iy a aie a OupvAtiy Doell in Mart. Fl. Bras. 2(2): 7 = G, ies ; Type: Brazil: Amazonas: S. Gabriel, Spruce 231. 18, PANICUM FROESII Swallen, sp. nov. i graciles, erecti vel adscendentes, ramo- 1 - longas laminae 1-2 cm, longae, 1-2 mm, latae; spiculae 0.8 longae, obovatae, papilloso-hispidae; gluma prima spicula 1/2 brevior, late acuta; fructus albidus 0.6 mm. longus. Annual; culms very slender, erect or ascending, more or less geniculate, branching at most of the ond lemma equal, slightly pointed beyond the fruit, strongly 5-nerved, the lemma empty; fruit white, about 0.6 mm. long. Type in the U. S. National Herbarium, No. 191078L, collected on Sandy river b 8, Irauasst, Aiary, Igana, Amazonas, Brazil November 6, 1915, by Ricardo de Lemas Frées (No. 21383). 1966 Swallen, Notes on grasses yal Colombia: Vaupes: Rio Vaupes: piceness Schultes & Cabrera 19698; Cerro Circasia, Schultes 56 & Keith 11702; Pimichin-Yavita trail, Maguire, Wurdack & Kesh h1792; between Maroa and Sabanita, Maguire, Wur & yore g 36423. Brazil: Amazonas: Tunu Lutzelburg 2 296s hia aid Rio Branco: Campos Gerais, region of Caracarahy, Frées 23618, 19. PANICUM SICCANEUM Trin. Linnea 10: 298. 1836. Type: Brazil: Colares, Rio Amazonas, Poeppig. 20, PANICUM FONTICOLUM Swallen, Mem. N. Y¥. Bot. Gard. 9: O. 1957. Type: Venezuela: Amazonas: Cerro Duida, Maguire, Cowan & Wurdack 29636. 21. PANICUM TAMAYONIS pei Bol. Soc. Venez. Cienc. %, 1 : 1953. ‘ Type: Sccamaniet ot Fa Gran Sabana, Tamayo 3061, 22. PANICUM peor ere Swallen, Mem. N. Y. Bot. Gard. 9: 259. 1957. e: Venezuela: Amazonas: Puerto Ayacucho, Maguire, Wurdack & Bunting 36043. alti; ligula arcuata 0.3 mm. longa; laminae 1-1.5 cm. longae, 2-l. mm, latae, longipilosae; spiculae 1.2-1.3 mm. longae, glabrae vel sparse pilosae, pedicellis brevis rigidis; gluma angusta, acuta, spicula 1/3 brevior, paula remota. ual; culms erect or ascending, freely branch- ing, 9-15 cm. tall, the ietas geen pearded, the hairs long; sheaths much shorter than the internodes, arcuate, 0.3 mm. long; blades 1-1.5 cm g, 2-k . wide, somewhat cordate, pate Her more or less long- pilose on both surfaces; panic -5 em. long, 2-3 cm. wide with a few stiffly spe vies to reflexed branches; spikelets 1.2-1.3 mm. long, glabrous» to s glume narrow, acute, 1-3-nerved, about 2/3 the length of the spikelet, somewhat remote; — second glume and t fruit; fruit 1 mm. long, obovate, acute, obscurely Aiea H or scabrous, yellowish. n the 0. 3. National Herbarium, No. 243451}, akipang Summit, Type collpeted. on exposed granite, Irain Te PHYTOLOGIA Vol. lh, no. 2 Kanuku Mountains, British Guiana, altitude 3500 feet, uot’ 18, 1963, by R. Goodland & P. F. Maycock (No. 53) e: Venezuela: Mount Auyan-Tepui, Tate 1287. 25. PANICUM cach paabeces Swallen, sp. no es, ramosi, iebldonté: “30 em. longi, nodis barbatis; ae arcuata, brevissima; laminae 2h. PANICUM PANDUM Swallen, Brittonia 3: 150. 1939. Typ pcre culms oe branching, forming tangled masses, as much as 30 cm, long, the nodes bearded; sheaths much ae ea than the internodes, long-pilose; ligule arcuate, very short; blades 2.5-5 cm. long, 2-6 mm. wide, thin, long-pilose; panicles spikelet, peabonecio acutish, 3-nerved; second glume and sterile lemma equal or the lume a little shorter ae covering the fruit or slightly pointed beyond its fruit <*? mm. long, white, shining, in the U. S. National Herbarium, No. 2307253, sollecies along Yavita-Pimichin trail near Yavita, Territory Amazonas, Venezuela, June 10, 1959, by Jo. dack ot This species was also collected at Yurupari, Vaupes, Colombia, by J. Cuatrecasas No. 6968, 26. PANICUM Snipes td Swallen, Mem. N. Y. Bot. Gard. 9:2 Type: Venezuela: Amazonas: Rio Atabapo, Bassett Maguire 29259. 27. PANICUM MAURYI Swallen, Fieldiana 28: 26. nna P. micranthus var. hirtum Maury, Journ. de Bot 3: be. 2889. Type: "Cerro Carichaud, savane seche, J. Chaff., ns» 26)" 1.5-2.8 om. Soudan 1.5-l mm. la atae, appressae vel adscendentes, sparse pilosae; spiculae obovatae 1.3- -. mm. longae, pilosae; gluma prima spicula 1/2 brevior. Perennial; culms decumbent at the base, freely 1966 Swallen, Notes on grasses 73 base; panicles up to 6 em. long, with ascending oe about 3 cm. long; spikelets obovate, 3 3-1. long, pilose; first glume broadly triangular, eave half as long as the spikelet; second glume and sterile lemma equal or the glume a little shorter; fruit smooth, shining, pale yellowi e the U. S. National Herbarium, No. ms io collected on Estate Gavinho, Marajé Island, Par Brazil, June, 1918, by Andre Goeldi (No. 222). Additional material examined: peli One kilo- meter NW. of Viltzberg, Coppenane River, Schulz & Donselaar 10555, 10559. 29. PANICUM PETILUM Swallen, sp. nov. Culmi gracillimi basi decumbentes, Specter ramosissimi, 30-0 cm. longi; ligula 1 laminae LiS=2,5 em. longae, involutae, edacenhantes vel reflexae; spiculae 1 mm, longae, glabrae; gluma prima spicula 1/2 brevior, subobtusa. rennial; culms very slender, decumbent spread- ing at the base, ascending, 30-l.0 cm. long, freely ake hing; sheaths shorter than the internodes on the in culm, much longer than the internodes on the peasuliss. Cop ert ged: ligule less than 0.1 mm, long, glabrous; blades 1.5-2.5 em. long, involute, ascend ing to rorinaeat panicle 1.5-3 cm. long, “a ae i mm 30. PANICUM POLYCOMUM Trin. Mem. Acad. St. Petersb. Nat. l: este 183k. Type: ny, SDD. uian 31. PANICUM ARCTUM Swallen, Bull. Torrey Club 75: 87. 194.8. Type: Surinam: Tafelburg, Maguire 221). 7% PHYTOLOGIA Vol. 1h, no. 2 32. PANICUM GRANULIFERUM H.B.K. Nov. Gen. et Specks 105. 1815. e: Venezuela: "Crescit in ripa fluminum Apure, Atabapo et Tuamini (Prov. Novae Guayanae).' 33. PANICUM SPISSIFOLIUM Swallen, Bull. Torrey Club 753 86. Type: British Guiana: Kaieteur Savanna, Maguire & Fanshawe 23097. 3. PANICUM KUHLMANNI Swallen, sp. nov. Culmi 10-20 cm, longi, graciles, ramosi, basi decumbentes; ligula brevissima, recta; laminae 1-2.5 en. eee 1-3 mm. latae, appressae; spiculae obo- vatae, 0.9-1 mm. longae, glabrae, pedicellis brevi- rg rigidis, glabris; gluma prima enervis, spicula 1/2 brevior as culms 10-20 cm. long or longer, a freely branching, decumbent spreading, forming a angled mat, aeeiticds sheaths usually much shorter than the internodes, glabrous or with a few hairs on the margins; ligule lige 2 short, straight; blades l- 2.5 em. long, 1- e, appressed, glabrous; panicles — =auinaed in the sheath, 1-2.5 em about em. Wide with a few ascending branches, ong glabrous ema Gather ots obovate, 0.9-1 mm. long, on relatively short stiff Ai ake glabrous; than half the eolsuctes on Bees do Pute between Arado Melgaes and Pimento Bueno, Mato Grosso, Brazil, June 1918, by Kuhlmann (No. 1778). 35. PANICUM KAPPLERI Steud. Syn. Pl. Glum, 1: 90. 1854. Type: Surinam: Kappler 177. 36. PANICUM BLACKII Swallen, sp. no Culmi gracillimi, Saguehented. an 30 om, Longs laminae 1-1.5 cm. longae, 1.5-2 mm, latae, adscen dentes, — ligula brevissima, recta; spic oulac 1.3-1.) mm. longae, a et gluma prima spicula 1/2 brevior, acuta, 3-ner Perennial?; say "He Slender, decumbent-spread- ae, about 30 cm. long, glabrous; sheaths usually h shorter than the internodes, glabrous; ligule 3 ae shor traight; blades 1-1.5 em. long, 1.5-2 mm. wide, stiffly ascending, blunt, villous on both - fa P rather prominent whi gins; panicles 3- cm. long with few to several slen ender, ascending, Tee-Piskaced branches; spikelets 1.3-1.) mm. long, 1966 Swallen, Notes on grasses 7 glabrous; first glume 1/2 as long as the spikelet, acute, 3-nerved; second glume and sterile lemna equal, just covering the fruit, obtuse, the sterile lemma containing @ well-developed staminate flower; fruit white. Type in the U. S. National Herbarium, No. 2238102, selected. on campo, Fazenda Capela, Territory of Rio Branco, Brazil, November Tg APs by George Black (No, 51-1071). 37. PANICUM ANGULOSUM Swallen, sp. nov. i graciles, erecti vel divergentes, ramosi, ap een usque ad 22 cm. longi; ligula arcuata, mm. longa, glabra; laminae 1.5-3.)1 cm. longae usque ad 3 mm. latae, planae, sparse pilosae; spiculae 1.4. mm. longae; glabrae vel + scabrae, purpurascentes; giuma prima acuta, spicula 1/2 brevior. Annual ; culms slender, erect to spreading, genic- ulate and branching at the nodes, as much as 22 cm. long, glabrous; sheaths much shorter than the inter nodes, glabrous or sparsely nee ligule prenenes -6 mm, long, glabrous; blades 1.5-3.\ cm. long, much as 3 mm. wide, flat, glabrous to sparsely Ppp panicles up to 8 cm. long, 11 cm. wide, ninco few solitary widely spreading to reflexed bran rather few-flowered, glabrous throughout; oe tiiiaco 1.4 mm. long, glabrous or somewhat scabrous, purple; spikelet; second glume and sterile lemma equal or the glume slightly shorter than the fruit, rounded, 5- nerved; fruit as long as the spikelet, rather thin, ee the U, 8. National Herbarium, No. 2380390, soil in full sun near brink collected in san roc of Kaieteur Fall, British Guiana, March 7, "1962, by R. S. Cowan & T. R. Soderstrom (No. 206k). Additional specimens referred to this species: Venezuela: 105 km. from El Dorado to Santa pene Estado Bolivar, Foldats 2786, Brazil: Territory 0 Amap&, Rio Ara aguari, Pires, Rodrigues & Irvine 61139. 38. PANICUM DENSIFOLIUM Swallen, Fieldiana 28: 25. 1951. Type: Venezuela: Amazonas: Esmeraldo (Cerro Duida), Steyermark 57755. 39. PANICUM MICRANTHUM H.B.K. Nov. Gen. et Sp. 1: 105. a5: e: Venezuela: "Caracasana.....juxta Villa de Cura.....", Humboldt & Bonpland. 40, PANICUM RIVALE Swallen, Bull. Torrey Club 753. 87. 198. 76 PHYTO L000 Ta Vol. 14, no. 2 Type: Surinam: Tafelberg, Bassett Maguire 2732. 1. PANICUM POLITII Swallen, sp. nov. Culmi erecti vel decumbentes, ramosi, ca. 30 cm. alti, glabri; ligula brevissima, arcuata; laminae adscendentes vel appressae, 1.5-l cm. longae, 2-l, mm 1 cm. longis; gluma prima spicula 1/2 brevior, acuta, l-nervis long, the slender solitary branches widely ascending or spreading, few-flowered; spikelets 1.3-1.). mm. long, glabrous, tinged with purple, on pedicels approximately 1 cm. long; first glume half as long as € spikelet, acutish, l-nerved; second glume obtuse, about 1/3 shorter than the fruit; sterile lemma 4 slightly longer than the fruit, acute; fruit shining, yellowish brown, e in the U. S. National Herbarium, No. 200162, 19148, by Bassett Maguire, H. R. Kunhardt, Jr. & Louis Politi (No. 27306). Additional material examined: Venezuela: Territory Amazonas: Esmeraldo Ridge, Maguire & Wurdack; Santa Barbara Savanna, junction of Rios Ventuari and Orinoco, Maguire, Cowan & Wurdack 3204. Brazil: Par&: Rio Cumina, Sampaio 533h. New Species of Panicum FPasciculata PANICUM OBLITUM Swallen, sp. nov i erecti basi decumbentes, 20-35 em. alti, Tm. latae, mmm. spicula 1/2 brevior, acuta vel obtusa; fructus minute rugosus. Annual; culms erect, or decumbent at the base, 20- 35 cm. tall, densely pilose; sheaths a little shorter than the internodes, densely pilose like the culms; blades 3-8 cm. long, 5-8 mm. wide, thickly covered : “Bes wide, > branches narrowly ascending or appressed, densely flowered, the main axis, branches 1966 Swallen, Notes on grasses TT and pedicels long-pilose; spikelets 2.|-2.5 mn. long, inflated, glabrous; first glume almost half as long as the spikelet, acute or eben broad and clasping; second glume and sterile lemma equal, just covering the fruit, 5-nerved, inflated, abruptly acute, whit- » the cells visable, the lemma containing a very a ink in the U. S. National Herbarium, No. 3181160, icateahed along "Carretera Central," between oer cla, Province Huaro ochiri, Department Lim Peru, March 2, 1952, by Ramén Ferreyra & ° Hutchison Pyrularia pa cn ICHUNENSE Swallen, sp. t Paw ciedt culms slen mder, decumbent spreading, freely branching, the branches erect, as much as 20 cm. long, glabrous or with a few long hairs at the nodes; sheaths all shorter than the internodes, gs sparse- ly pilose with long hairs; ligule arcuate, 0,.8-1 mm long; blades 2-l, em. _ Long, 3-10 mm. wide, ‘v ery thin, ter the fruit, 5-nerved; fruit 1.7 m. RONG tea, acute, pale, finely papillose-roughen in the U. S. National He eo No. 2362348, h° h6', Long. 63° 18!, state of Bolivar, Venezuela, December 28, 1961, by Julian A. Steyermark (No. 90359). Also collected to the north of Salto Maria Espuma, Steyermark 90292. Laxa cm. longae, 6-10 mm, latae, attenuatae, ae paniculae 35 em. longae, pyramidales, ramis usque ad 78 PHYTOLOGIA Vol. 1h, no. 2 12 cm. longis, ramosissimis; spiculae 1.4-1.5 mm. longae, ovatae, glabrae; gluma prima lata, spicula 1/2 brevior,. Perennial; culms erect, somewhat geniculate and rooting at the lower nodes, firm, glabrous, 125 cm. tall; sheaths about as long as the internodes, loose, glabrous; li arcuate, about 0.2 mm. long, minute- ly ciliate; blades 25-0 cm. long, 6-10 e numerous branchlets along them; spikelets 1.4-1.5 mm. long, ovate, pointed, glabrous, appressed to the branches; first glume broad, clasping, 1-3-nerved, ela, June 19 1954, by John J. Wurdack & L. 8. Adderlay (No. 2986). This species resembles Panicum hylaeimm Mez, but the blades are not at all cordate and the spikelets are pointed rather than blunt. Stolonifera PANICUM PIRINEOSENSE Swallen, sp. nov. Culmi erecti, 1.3 m. alti, rhizomatosi; vaginae internodiis multo breviores, inferiores dense pubes- centes, superiores glabrae; laminae 12-18 cm. longae, -2 cm. latae, pubescentes; inflorescentia 21 cm. bb - i 3-5-nervis, spicula 1/2 brevior, in parte superiore pubescens; lemma sterile 2-glandulis 1/y infra extremitatem praeditum. ; Perennial; culms erect 1.3 m. high, apparently glabrous, the collar area densely pubescent; ligule ciliate, about 1 mm. long; blades 12-18 em. long, 2-2 em. wide, acuminate, finely pubescent on both surfaces to nearly glabrous; inflorescence 21 cm. ‘ : t lower ones 3 cm. long, floriferous to the base, the base densely pubescent; spikelets 3.2 mm. long, short- pedicellate in two rows on one Side of the rhachis, 1966 Swallen, Notes on grasses 79 the second of a pair of spikelets seis abortive; first glume triangular, acute, 3-5-nerved, about half as long as the spikelet, rote BE rene peabes or on the mar gins coarse aera hairs 1 mm, long, pubescent or pilose across the back, the lemma bearing 2 glands about 1/); below the summit ; fruit i. i Pap long, yellowish, smooth and shining. h . S. National ey rapa No. 2236507, oniaacs between rocks, at Pirineos, Goifs, Brazil, October 18, 1956, by Amaro Macedo (No. 4.80 05). Penicillata PANICUM LONGIPEDICELLATUM Swallen, Sp. nov. Culmi basi decumbentes 1.6 m. longi, glabri; ligula 1 mm. longa, firma, hispido-ciliata, 2 mm. longa; laminae planae 35- hl ¢ em. longae, 2.3-3.6 mm, latae, seat pal poe eN it scabris; panicula Culms erect from a lo decumbent base, l. long, glabrous; sheaths longer than the inte the uppermost elongate, os ap eee papillose-hispid le fi 1-l. em. long; first glume clasping, as s spikelet or nearly so, occasionally longer than the hairs near the tip, and on the margins; second glume and sterile lemma equal, strongly 5-nerved, papillose- roughened; fruit 3.5 mm. long, striate, the margins of the lemma not inrolled, the summit somewhat crested, e in the U. S. National Herbarium, No. 1255492, ‘ a de Itatiaia, Rio de Janeiro, Brazil, 1200 M., January 18, 1925, by Agnes Chase (No. 8342). Additional specimens examined: Brazil: Paran&: Tres Barras, Dusén 17587; Santa ie Alta ot narrower, usually shorter blades 80 P ey? 0 L:0°G-TA Vol. 14, no. 2 Lorea PANICUM FONTANALE Swallen, sp. nov. i erecti, 45-50 em, alti, glabri; ligula deficiens; laminae usque ad 30 em. longae, 3.5 em. latae, ad basin angustatae, glabrae; spiculae 3.2 mm. longae, pedicellis spiculis multo longioris; gluma prima spicula 1/3 brevior, erennial; culms erect, 5-50 em. tall, glabrous; sheaths glabrous elongate extending nearly to the base of the panicle with a much reduced blade; ligule of the sheath, glabrous, even on the mar ins; panicle 9 em. long with spreading to reflexed few-flowered sterile lemma equal, acute or subacute, 5-nerved, _ glabrous, the lemma with a very well developed stami- nate flower; fruit 2 mn. long, acute or subacuminate, white, glabrous, : Type in the U. S. National Herbarium, No. 23786l1, collected “on top of waterfall," Steyermark eee on i oO g long attenuate blades, and larger and laxer panicles of PANICUM LUTZII Swallen, sp. nov. Culmi caespitosi, rhizomatosi, 35-50 em. alti, foliis basi aggegatis; ligula brevissima, ciliata; laminae 15-30 cm, longae, basi conduplicatae, usque 5mm. latae; spiculae 1.8-2 mn, longae, pedicellis plerumque breviores; gluma prima spicula 1/3 brevior, ule very short, ciliate; blades 15-30 om. long, firm, wide; inflorescence 8-10 om, long, about 1 em. wide, the branches appressed, as much as 5 em. 1 ng, naked at the base; spikelets 1.8-2 mm. long, glabrous, the pedicels usually shorter; first glume 2/3 as long as 1966 Swallen, Notes on grasses 81 tong, aeeRg aN acute, brownish. e the U. S. N acaenail orn No. 22.0718, 66Llokbed on Tijuca, Rio de Janeiro, Brazil, February 1914, by Adolpho Lutz (No. 687). Dichotoma PANICUM THELMATUM Swallen, Sp. nov. decumbentes, ramosi, pilosi vel papilloso- pilosi; ligula dense ciliata ca. 1 mm. longa; laminae usque ad em. longae, & mm. latae, acutae appressae, marginibus scabrae et papilloso-ciliatae; spiculae 2.9 mm. longae, sparse pubescentes vel pilosae; gluma prima 2 mm, longa, poi 3-nervia; fructus 2.4 mm dense line of hairs about 1 mm. long; blades as much as 5 cm. long, 8 mm. wide, acute, appressed, sparsely hispid on the upper surface, the margins scabrous and papillose-ciliate; panicles partly Sree in the sheaths or shortly exserted, 3-1.5 cm. long, about 1 m. wide, with few aan? narrowly ascending few flowered branches, the main axis and branches pilose; spikelets 2.9 mm. long, Saeeads pubescent or pilose; first glume 2 mm. long, broadly triangular, acute or rounded, 3-nerved; second glume and sterile lemma equal just covering the fruit, many nerved; fruit 2.) mn. long, yellowish, minutely striate roughened in the U. S. National Herbarium, No. 2182190, collected in burnt part of boggy central area, Serra do Sol, Bolivar, Venezuela, 2100 m., December 28, 195i, by Bassett Maguire & Celia K. Maguire (N ho “ie Pe telmatum appears to be somewhat related to Sian e Doell, but it is caespitose rather than ree with larger and longer blades, and usually larger panicles Ungrouped PANICUM ITATIAIAE Swallen, sp. nov Culmi basi decumbentes, ramosi, ae 72, ‘Lone. : Vaginae internodiis longiores vel eriatnyetse glabrae, marginibus pee as mm. longa, minutissime ciliata; laminae 10-12 oe longae, 10-12 mn. latae, subcordatae, glabrae, marginibus scabrae et papilloso-hispidae; panicula 15 cm. longa, ramis patentibus, paucifloris, glabris; spiculae b=le5 Tm. longae, glabrae; gluma prima 3-5-nervis, angusta acuta; gluma secunda et lemma sterile 11-13-nervia, 82 PHYTOLOGIA Vol. 1h, no. 2 giluma fructu breior, lemma fructu longior; fructus ini numerous internodes, glabrous on the neoks papillose- hispid on the margins; ligule 0.3 mm. long, minutely ciliate; blades 10-12 cm. long, 10-12 mm. wide, sub- cordate, acuminate, glabrous on both surfaces, sca- brous and papillose~hispid on the margins; panicle long-exserted, 15 cm. long, about as wide, the branches spreading, few-flowered, glabrous, the axis more or less glandular-spotted; spikelets heh .S mm long, glabrous; first glume narrow, acute, 3- ce onereetl second glume and sterile lemma 11-13-nerved, the glume a little shorter, the lemma a little longer than the fruit; fruit 3.8 mm. long, 1.6 mm. wide brown, the margins not inrolled, a Fs a small pubes- cent crest at the summi Type in the U. S. National Herbarium, No. 1255836, collected under bamboo on mountain slope below Maecieiras, Serra de Itatiaia, Rio de Janeiro, Brazil, 1700-1800 = ges itnaes January 18, 1925, by es Chase (No, 832 PANICUM INFUSCUM Swallen, sp. nov i erecti vel decumbentes, 30- 60 em. Longi plerumque multinodosi; laminae 6-9 om. longae, rid 12 mm. Leora subacuminatae, dense pubescentes; spiculae ongae, nervosae, glabrae; gluma prima spicula 175. 3/\k brevior, enervis vel 9-nervis; fructus fuscus, lucidus, eee us, So Perenni 5 rect from a decumbent base, rooting at the govas neaek. 30-60 cm. long, usually with many short intern odes: sheaths about @as long as the internodes, rather densely pilose, the hairs spreading, with a dense rim of hairs on the collar; blades 6-9 cm. long, the uppermost reduced, 6-12 mm. wide, subacuminate, softly and densely pubescent on -8 cm spikelets 1.8 mm, long, strongly nerved, glabrous; first glume 1/4 to 2/3 as long as the spikelet, nerve- sterile lemma as long as the fruit, subacute, 9- nerved; fruit light brown, smooth and shining, elliptic, apiculate Type in the U. S. National Herbarium, No. Lhh8L.755 collected on atcer shaded slope on trail to Tijuca, Guanabara, Brazil, altitude 700 m., April 28, 1930, by Agnes Chase (No. 1215). 1966 Swallen, Notes on grasses 83 PANICUL hp tro aging Swallen, sp. no C recti vel decumbentes, 55 ig longi, = ca. 25; ligule 0.3-0.4 mm. longa; laminae ..5-7 longae, 7-11 mm, latae, wh gce rorbetcree dense ea centes; spiculae 1.8-2 mm. longae, hirsutae; gluma prima spicula ca. 1/2 brevior, coer 3-nervis; fructus fuscus tenuiter rugosu Perennial; culms erect to nisiakdines 55 em. long, with about 26 nodes; sheaths all longer than the -0.. mm long; blades .5-7 cm. long, 7-11 mn, wide, spreading, slightly asymmetrical, subcordate, finely and densely pubescent on both surfaces; panicle 12 cm. long with a few ascending to spreading few-flowered branches; spikelets 1,.8-2 mm, long, hirsute; first glume a little more than half the length of the perenne broad, 3-nerved; second glume and sterile lemm obtuse, Senerved, the second glume shorter sg the fruit; fruit Light brown, ovate, acute, finely cross- era mer Type in the U. S. National pega No. 1612918, collec Sed ie open forest, Bragan azil, June 27-29, 193, by Jason R. Swelien (tO. 5037). The Serrata Group of Ichnanthus Blades 4-5 mm. wide, about 30 cm. long; panicles lax, the pedicels slender, usually much longer than the spikelets...ccsces 1. I. serratus Blades 8-11 mm, wide, usually more than 50 cm. long; anicles stiff, the pedicels usually shorter Panicles large, open, 30 cm. ae blades ciliate for 15 cm. above the ba >. Se, neblinaensis Panicles relatively small, usually narrow, not more thi O em. long. Blades attenuate to a fine point, meee and shining. Spikelets 3.4-3.6 mm. 3+ To vongifoiue Blades acuminate, not shining. ———— a mm. long, sparsely pilose with long hai I. ephemeroblepharis ietciabe “h-5 mm. long, glabrous. Spikelets 5 mm. long; blades long-a attenuate to the base; panicle — spreading. 5. I. tectus Spikelets ma. long; blades ek but not long-attenuate to the base; panicle nar- row, the branches ascen . 6. I. angustus 8h PHYTOLOGIA Vol. 1h, no. 2 1. ICHNANTHUS SERRATUS Swallen, Fieldiana 28: 30. 1 . Venezuela: Steyermark 58298, 2. ICHNANTHUS NEBLINAENSIS Swallen, sp. nov Culmi erecti, 1.35 m. alti, glabri; ligula cili- ata 1 mm. longa; laminae usque ad 55 em. longae, 8-11 mm. latae, firmae, glabrae, marginibus irregulatim et fortiter scabris; panicula 30 cm. longa, 10 cm. lata, ramis adscendentibus usque ad 15 em. longis, dense seabris; spiculae 3.5 mm. longae, glabrae; gluma prima 2.5 mm. longa, acuta; fructus 3 mm, longus, acutus, — ca. 1.5 mm. songs: mnial; culms erec 1.35 m. tall, flattened, glabro vane lower sheaths Nib longer, the upper ones 4 little shorter than the pt oi age glabrous or cili- ate on one side; ligule a dense line of hairs about 1 mm. long; blades flat, Pbsecsg as much as 55 cm. long, 5-11 mm, wide, firm, flat, — on both sur- faces, the margins ciliate for as much as 15 cm., the em. long, about 10 cm. wide, pyramidal, with ascend- ing branches up to 15 cm. long, the branchlets spread- » the main axis, branches, and branchlets densely and finely scabrous; spikelets 3.5 mm. long, glabrous; first glume 2.5 mm. long, acute or rounded; second glume and sterile lemma equal, 5-nerved, the lemma ain a well developed palea a little longer than the lemma; it 3 mm. long, elliptic, eres straw- colored, yop wings about half as lon, n the herbarium of the New York cle Garden, eollested on west escarpment slopes of Cafi Grande east of Cumbre Camp, Cerro de la Neblina, Rio Yatua, Territory of aes 1100-1300 meters, Venezuela, November 25, 1957, by Bassett ROO John J. Wurdack & Celia K. Maguire (No. 2239). related species, but differs from all of them in the ge open panicle and the long ciliate base of the blades. 3. ICHNANTHUS LONGIFOLIUS Swallen, Fieldiana 28: 29. Venezuela: Steyermark 58020, hk. ICHNANTHUS EPHEMEROBLEPHARIS rei & Prées, Bol. Inst o. Norte 15: 5. 1948. RS: “Pr es 22898. 5. ICHNANTHUS TECTUS Swallen, Mem. N. Y. Bot. Gard. 9: 265.1957. Venezuela: Cowan & Wurdack 31139. 1966 Swallen, Notes on grasses 85 6, ICHNANTHUS ANGUSTUS Swallen, Mem. N. Y. Bot. Gard. 9: 266. 1957. Venezuela: Phelps & Hitchcock 1.76. A New Species of Sacciolepis SACCIOLEPIS PUNGENS Swallen, sp. nov Culmi caespitosi, erecti, 90 cm. alti; ligula 0.2 mm. longa, minute cili hee laminae firmae, nervosae, planae, usque ad 25 cm. longae, 3 mm. latae, supra minute pubescentes; spiculae 1.8-2 mm. longae pedi- cellis pubescentibus; gluma prima spicula 1/3-1/2 brevior, acuta, S-nervis; fructus l- mn. longus, albus, nitidus, summo minute hispidulus. Perennial; culms tufted, erect, 90 cm. tall, gla- brous, with numerous short internodes toward the base; sheaths, at least the lower, much overlapping, gla- brous; ligule 0.2 mm. long, minutely ciliate; blades firm, strongly nerved, flat below becoming involute toward the tip, as much as 25 cm. long, 3 mm. wide, sere pubescent on the upper cietae: toward the bas otherwise glabrous; spikelets 1,8- 2 mm . Song, lets, pubescent with spreading hairs; first glume 1/2 to 2/3 as long as the spikelet, acute, 5-nerved; sec- ond glume and sterile lemma equal, strongly 5-7- shining, minutely hispidulous at the tip. Type in the U. S. National Herbarium, No. 2208531, colieehan« in marichal at northwest base of Cerr Gavilan (Cerro Carichana), Bolivar, Venezuela, 100 meters altitude, December 21, 1955, by John z. Wurdack & J. V. een ta (No. 4.08 Sh). New Species of Olyra OLYRA WURDACKII Swallen, sp. nov Culmi simplices, 3.3 m. alti, “glabri, nodis bar- batis pilis appressis; laminae 15-26 cm. longae, 5 cm. latae, glabrae; panicula 27 mm. longa, rami - Osis, appressis, verticillatis; spiculae masculae 6- 10 mm. longae, acuminatae vel attenuatae, glabrae, appressae; spiculae femineae 7-0 mm. iy pilosae, glumis acuminatis, non aristatis; fructu longus, albus, lucidus Culms simple, 3.3 m. tall, glabrous, with dense bands of white appressed hairs at the nodes; sheaths on the margins near the base, glabrous on both sur- faces; panicle 27 cm. long, composed of numerous 86 PHYTOLOGIA Vol. 14, no. 2 appressed verticillate branches, making a dense mass of almost entirely staminate spikelets; branches 10- 18 em. long, ascending or appressed, densely villous in the axils; staminate spikelets 8-10 mm. long, acu- minate or attenuate, glabrous; appressed; pistillate spikelets 2 or 3 at the ends of the branches, 7-8 mm. ng. e in the U. S. National Herbarium, Nos. 2307269 and 2307270, collected at edge of open rock on crys- talline dome on right bank of Rfo Siapa just below d - tude 300-500 meters, July 21, 1959, by John J. Wurdack & L. S. Adderley (No. 1.35)0). OLYRA MARANONENSIS Swallen, sp. nov. erecti, 85-90 cm. alti, glabri; laminae 19- 22 cm. longae, 3-l, cm. latae, glabrae, nitidae ; ~3.5 mm. longae, acuminatae, pilosae; spiculae femineae 10-13 mm. longae; gluma arista flexuosa 8-12 mm. longa; fructus 8 mm. longus, marginibus dense villosus. Culms erect, 85-90 cm. tall, glabrous; lower sheaths distant, glabrous, much shorter than the internodes, the upper overlapping with well-developed blades; blades 19-22 em. long, 3-l cm. wide, as et- @mm. long; fruit 8 mm. long, densely villous on the gins at the base, the margins near the summit less villous, but distinctly so; palea acuminate, as long as the lemma, Type in the U. S. National Herbarium, No. 2382316, collected in r orest on ridge crest of Quebrada Chuivi, valley of Rio Marafién near Cascades de Mayasi, elevation 500-550 m., Amazonas, Peru, September 20, 1962, by John J. Wurdack (No. 1936). Ov. validi, erecti, ca. 2.5m. alti, nodis ramosi; laminae 25-35 cm. longae, 6-7 cm. latae, sub- cordatae, petiolatae, glabrae, marginibus scabris; aniculae 12-17 cm. longae, ramis adscendentibus vel divergentibus, subverticillatis; spiculae masculae 1966 Swallen, Notes on grasses 87 8-10 mm. longae, breviaristatae; spiculae femineae 2- -& cm, longae; fructus 8 mm. longus, cylindricus, glabrous. Perennial; culms stout, erect, about 2.5 m. tall, branching at the upper nodes; lower sheaths bladeless e an the appressed hairs above the nodes, crowded upward; blades 25-35 cm. long, 6-7 cm. wide, somewhat cordate the branches stiffly ascending or spreading, roughly verticillate, the upper ones solitary, scattered; staminate spikelets appressed to branches, 8-10 m. long including the short awn, glabrous, the stamens mm. long; pistillate spikelets 2 or 3 at the ends of the branches, 2-2.8 cm. long including the awns, e body of the glume 8-12 mm. long; fruit about 8 mm, long, slender, cylindric, glabrous. pe in the U. S. National Herbarium, No. 24325), collected in swamp forest at Riber&éo Grande, Muni- cipio Jatai, Goifs, Brazil, July 25, 1956, by A. Macedo (No. 4.386). Also collected in the same locality Macedo 4.380. The second specimen is undoubtedly the same species but the blades are all much smaller, not more than cm. long and 3 cm. wide. A New Species of Imperata IMPERATA FLEXUOSA Swallen, sp. nov. Culmi erecti, 120 cm. alti, rhizomatibus promin- entibus; laminae usque ad 80 cm. longae, 1 cm. latae, Culms erect from deep prominent rhizomes, 120 cm. tall; lower sheaths becoming fibrous with age, the upper ones much longer than the internodes, glabrous or somewhat pilose near the summit; blades as much as 80 cm. long, attenuate, long-narrowed to the base, as much as 1 cm. wide, glabrous, the margins ciliate for ding, the axis slender, the branches in distan e oups, few-flowered, slender, flexuous, of unequ hairs about 10-12 mm. long on the callus and glumes. e the U. §. National Herbarium, No. 1913732, collected "En conucos. Kavanoyen. Selva de Oparuma. Guayana Venezolana" Venezuela, May 39, 1946, by T. Lasser (No. 1886). 88 PEI TOLoeia Vol. 1h, no. 2 Eriochrysis Spikelets not crowded, not greatly obscured by the airs, Spikelets ovate-acute. Panicles 8-1 cm. long, spikelike, the racemes appressed; spikelets 5-6 mn. E. holcoides Panicles 17-22 cm. long, not spikelike, the racemes ascending; spikelets l-5 mm. long. ee warmingiana Spikelets sy ser eae aa rounded or ~plunt at the Sts ° jaxa gs gE crowded, “mostly obscured by the hairs. Hairs of the spikelets light brown or tan; spikelets not glomerate. clues or at least most of them irregularly lobed, obtuse, foliage glabrous.. l.. E. glabrifolia Glumes acute; foliage densely appressed villous. villosa Hairs of spikelets dark brown; spikelets usually dense, RLOMORAUEL : wc cs coc ces, Ge cee cayennensis 1. ERIOCHRYSIS HOLCOIDES (Nees) Kuhlm. Comm, Linhas Telegraph. Estrat. Matto Grosso 67: 89 Annexo 5S, Bot. “Ils 89> 1922. Saccharum holcoides Hack. in Mart. Fl. Bras. 2(3): 254. 1003. Anatherum holcoides Nees, Agrost. Bras, 32h. 1829 Specimens examined: Brazil: Paran&: Pinhoe Dusén 8927, 15820; spas Swallen 8526; Tabatinga, 0114; Klein 776; Reitz 2710; Mafra, Smith & Klein 10668. Rio Grande do Sul: D, Petrolini, Swallen 7307; cy ee Swallen 821); Sao Francis co de Paula, Rambo 5h, 2. ERIOCHRYSIS WARMINGIANA (Hack.) Kuhlm. Comm Li Telegraph. Estrat. Matto Grosso “67: 29. Saccharum warmingiana Hack. in Mart. Fl. Bras. 23) Specimens examined: Bolivia: Santa Cruz: Buena Vista, Steinback 7032. Brazil: Mato Grosso: Aquidauana, Chase 11063; Swallen 9533 1966 Swallen, Notes on grasses 89 3. ERIOCHRYSIS LAXA Swallen, sp. nov. i caespitosi, erecti, ma cm. pave nodis pilosis; vaginse glabrae 3; ligula mn. longa, minute ey Eee ig eae phan filiformes usque ad cm. longae et 2mm. latae, glabrae, eae culmorum usque ad 60 cm. longae et ) mm. latae, su- periores multo breviores usque ad 6 mn. latae, supra dense pilosa; inflorescentia 22 cm, longa, 2 om. lata; spiculae sessiles ca. | mm. longae, obovatae, glabrae, marginibus pilis 3 mm. longis; gluma secunda angusta marginibus supra ciliata; lemma sterile hyalinm, marginibus pauciciliatum Culms densely tufted, erect, 130 cm. tall, gla- brous, the nodes appressed-pilose; sheaths all longer than the internodes, glabrous; ligule about 1 mm. long, minutely erose or ciliate; blades of innovations as much as 15 cm. long, subfiliform below, sometimes broadened above to as much as 2 m., glabrous, caaaa of the culm as much as 60 cm. long and l mm. wide, e upper ones much shorter as much as 6 mm, wide, densely pilose on the upper surface; inflorescence 22 cm. 2cm. wide, interrupted below, the anne and branches glabrous; sessile spikelets about l: mm. long, usually obovate, the tip rounded, the margins bearing hairs as much as 3 mm, Lang the back glabrous, papil- late-roughened; second glume narrow, the margins cili- ate in the upper half; sterile lemma hyaline, icin ly ciliate on the ek ee with relatively strong hairs. Type in the U., S. National Herbarium, No. 1256173, collected in wet ground near streamlet, Lauras, Minas arent Brazil, March 5, 1925, by Agnes Chase (No. 729 It is possible pe ysipi this is — taxon described by Hackel as Erioc sis cayennens var. laxiuscula. The description is too insufficient to be aare or 3%; and no type was found in y Mrs. Chase or Prof. Hitchcock, a] me better ae to describe it as a distinct spe Additional ed BE examined: Colombia: teen eatets Bolivia: Santa Cruz, Steinback 6975 bis. Lacus Ypacaray, Hassler 12:92; Posta Ipihend, cp 13307. Argentina: Prov. Corrientes: Estancia Santa ar prc Londrina, Swallen 8728; SHo Paulo: Pindo een oe Viegas 3381; Pickel s.n.; Minas Gerais: Glaziou 17899 is, Chase 11365; Santo Rita do Araguaya, Chase 12038, 12039; Rio Verde, Chase 12089; Mato Grosso: Dour ados, Chase 10945, 10956; Campo Grande, Gehrt 11h. 90 PHYTOLOGIA Vol. 1h, no, 2 rescentia 16 cm. longa, 5-10 mm, lata, densa, spic- ulae sessiles 3 mn, longae, obovatae; gluma prima dorso glabra, marginibus pauciciliata, summo dense hispida; spiculae pedicellatae ca. 1.3 mm. longae; lemma fertile ca. 2 mm. 1 back, glabrous, or pilose near the summit; ligule 1 mm. long, short-pilose at the tip; blades of innova- more or less ciliate on the margins, with dense tuft of hairs at the tip; pedicellate spikelet about 1.8 mm. long, densely hairy at the tip and on the margins; fertile lemma about 2 mm. long. Type in the U. S, National Herbarium, No. 1960865, collected in banhado, Estacdo Experimental, Ponta osso, Parané, rater! January 28, 1916, by Jason R. 2 .* Additional specimens examined: Brazil: Paran&: Guarapuava, Swallen 8803. Santa Catarina: Curiti- banas, Reitz & Klein 11806; Snith 11103; Indios, Smith 11220, villosis; spiculae 3-l mm. longae; gluma prima margin- ibus villosa pilis 2-3 mm, longis; gluma secunda summo et marginibus pilosis; lemma fertile 2 mn. longum mar- ginibus pauciciliatum, S densely tufted, erect, about 1 m. tall, glabrous, the nodes distinctly appressed villous; sheaths densely ascending or appressed villous, usu- ally longer, occasionally a little shorter, than the internodes; ligule about 1 mn. long, obscurely cili- ate; blades flat, 1-30 em. 1 mn. wide, firm, densely appressed-villous, the uppermost reduced; é 1966 Swallen, Notes on grasses 91 inflorescence 22 cm. long, dense, somewhat interrupted below, the axis and branches appressed-villous; spike- at the summit; fertile lemma 2 mm. long, sparsely ciliate on the margins. in the U. S. National Herbarium, No. 1961568, collected in banhado, Rio Cacador, Santa Catarina, re January 21, 191.6, by’ Jason R. Swallen (No. Additional material examined: Brazil: Santa Catarina: Lages, Swallen SIDES Smith 1222; Reitz 6456. Rio Grande do Sul: Jaquirana, Rambo 5209; Cambara, Rambo 36176. 6. ERIOCHRYSIS CAYENNENSIS Beauv. Ess. Agrost. 8. pl. eS aren 812. This is the “most widespread of all the species of Eriochrysis. It ranges from Mexico and the West Indies to Bolivia and heared It is composed of several elements, none of whic an be separated satisfactorily with the aabortal> avntiubase Erianthus Nodes conspicuously bearded. Glumes entirely glabrous; spikelets 3.5-l mm. long; terile lemma and lemma Pe etcé E. asper Glumes villous across the back or “eiliate on the pice a only. Spikelets 6-8 mm. long; sterile and lemma ciliate Glumes villous across the back; culms usually 2-3 m. tall; panicle large, dense, pale, many- Ges 2. E. trinii ins; culms less than 2 all, usually not much more than tall; panicles small, Diener few- flowered, tinged with pur Sp ot lemma and lemma ciliate, ge blades row, not much widened a ur ureus Sterile lemma and lemma villous on the k, the sterile lemma awned; blades broadened above, as much as li, mm. — . biaristatus Nodes glabrous or ae optic Pact oe the hairs, very shor Panicles exserted. 92 PHYTOLOGIA Vol. 1h, no. 2 Culms glabrous below the panicle. 5. E. angustifolius Culms villous below the panicle, Panicles entirely or only partly included in the eath; callus hairs longer than the spikelet. Culms stout; inflorescence included rived we the base; sterile lemma and lemma cilia glabrinodis Culms eonionde slender; inflorescence usually irely enclosed in the ven er sheath; ster- iis lemma and lemma glabrou 1% "R. clandestinus 1. ERIANTHUS ASPER Nees, Agrost. Bras. 315. Hato Type: Brazil: "Habitat in ripa porte Sa cisci pone Salgado, et alibi in prov. Minarun, " a. = TRINII Hack. in DC, Monogr. Phan. 6: 135. 1889, EH. saccharoides spp. genuinusvar. trinii in Mart. Fl. Bras. 2(3): 258. 1583. 3% i PURPUREUS Swallen, sp. nov. erumque ca. 1m. alti, infra inflores- centia appresso-hispidi, nodis barbatis; laminae angustae; spiculae = mn. longae; gluma et lemmata oo — pe the U. S, National Herbarium, No. 1960827, collected in banhado, near Curitiba, Parand, Brazil, 573). Additional specimens examined: Brazil: Paran&: itibanos, Klein ane Mun. peco, Smith : Klein 11627; Mun. P. Unido, Smith & Reit 862) Sho Paulo: a Grande Minas Gerai H eaves, Chas 10257. Rio Branco: Serra Tepequem, Maguire & Hagutae hOosS he ERIANTHUS BIARISTATUS (Hack, ) Swallen, comb. nov. E. saccharoides ssp. angustifolius var biaristatus Hack. in Mart. Fl. Bras. 2(3): bts) i803. 1966 Swallen, Notes on grasses 93 Glaziou ron: one side; gett 8919. Santa Catarina: Santa Cecilia, Reitz & Klein voting ert Paulo: Fazenda Campinha, NW of Moji-Mir ay Bolivia: Nor Yungas, Hitchcock 327373 Agel 5. mest ANGUSTIFOLIUS Nees, Agrost. Bras. 316, 1 29. Type: ",...ad flumen Rio Grande do Sul dictum prope fines regni Paraguayani. (Sellow)..." 6. ERIANTHUS GLABRINODIS (Hack. ) en. comb. nov. &. saccharoides ssp. genuinus var. trinii subv. labrinodis Hack. in Mart. Fl. Bras. 2(3): E. trinii subv. glabrinodis Hack. in DC. Monogr. Phan. 6, 135. 1889. Specimens examined: Brazil: Rio Grande do Sul: S. Leopoldo, Dutra 560; Passo Fundo, Fazendo da Brigada, Swallen 7781. S&o Paulo: Ypiranga, Usteri. Minas Ger ais: Ouro Preto, Chase 9373; Juiz de Féra, Chase 8601; 7. ERIANTHUS CLANDESTINUS Swallen, sp. nov. Culmi graciles, glabri; inflorescentia in vagina inclusa; glumae villosae; lomnata sterile et Practite glabra, Culms slender, glabrous; inflorescence included in the upper sheath; glumes villous ghee ae the back; etgeert ee fertile lemmas glabrous, entir mens examined: Brazil: eyes “seh ah Grosso, Swalon “B67 s Curitiba, Dusén 13843; Campo Grande, Dusén 421). Santa Catarina: Bom Re tiro, ays Fp 287k; Poste Uniao, Reitz & Klein 12628; Smith & Bern J01Bes Klein 12192; Campos et Reitz & Klein 11658; Rio Cagador: Swallen 8255. Rio Grande do Sul: Porto Alegre, Rambo 11390; pant 13ee; Pelotas, Sacco 313; Mun. Rio Par o, Jurgens G63. Rio de Janeiro: Chase a easy Lutzelburg 9. Mato Beckee: Dourados, Chase 6 New Species of Trachypogon ligula 1 mm, Lanes; laminae rsnied eg erectae, atbaanaths 12-18 em. longae, dense hispidae; spicula fertilis 10 mm. Pp callo 3 mm. longo dense hispido; gluma prima brevihispida; arista nag cm. Perennial; culms erect from pr rominent cass. 75 cm. tall, glabrous, the nodes short appressed hispid; 9h PHYTOLOGIA Vol. 1h, no. 2 sheaths all longer than the internodes, appressed hispid; ligule inconspicuous, less than 1 mm. long; blade erect, curved, involute, attenuate, 12-18 cm. long, densely hispid on both surfaces, becoming flat with age as much as 3 mm. wide; fertile spikelet 10 mn. long, the callus 3 mm. long, densely hispid; first glume evenly hispid, the hairs “trig awn 9-10 cm. long, 2-geniculate, hispid, the hairs 1-1.5 mm. long; staminate spikelet about 8 mm. long, hispid like the fertile spikelet. Type in the U. S, National Herbarium, No. 1961900, site on campo azenda de Criac&o, Ponta Grossa, Brazil, Potetass 6, 1946, by Jason R. Swallen (io. anaes). TRACHYPOGON DENSUS Swallen, sp. nov, ense caespitosi, yah 60-90 cm, ete. nodis hispidis; ligula ion nga; lamin ea volutae (vel planae), attqinetas, 7 ae htyge 10- "25 ¢ longae, 2mm. latae; racemus solitarius, 5- longus; spicula fertilis 9 mm. longa, callo mm. sae gluma prima sparse pilosa vel a tone oe .8- cm. longa, hispida pilis 1-1.5 mn Perennial; culms densely tufted, erect, °60-90 cm. tall, glabrous, the nodes appressed hispid; sheaths mostly longer than the internodes, the upper ones much longer, glabrous or sparsely hispid near the summit; ligule inconspicuous, about 2 mm. long; blades mostly involute, or becoming flat with age, wide; racemes itary, erect, long-exserted, 5- 9 cm. long; fertile aeibelet 9 mm. long, dark brown, the ca. mm. long; first glume sparsely pil with hairs 1-1.5 mm. long; — spikelets 7 mm long, sparsely pilose or hispid. Type in the U. S, National Herbarium, No. i collected in campo grr de Criac&o, Lages, Santa sponge bg January i, 1 » by Jason R. Swallen (No. 9). TRACHYPOGON PARVIFLORUS ae sp. erecti, 60-70 cm. alti, nodis Mebiavias longa, allo 2 mm, longo dense barbato; thes prima hirsuta pilis usque ad 3 mm. longis; arista lh cm. longa, Rote 2-geniculata, hispida. Perennial; culms erect, 60-70 cm. tall, glabrous, the tak bearded to almost glabrous; sheaths gla- ous, sparsely hispid on the collar, the uppermost nearly bladeless; ligule 3-10 mm. long, decurrent; 1966 Swallen, Notes on grasses 95 blades narrow in the lower part, stiff and rigid, be- comi flat, as much as 3 mm. wide toward the ends; the callus 2 mm. long, densely barbate in the center; first glume conspicuously hirsute, the hairs longer near the summi as much as mn, long; awn cm. long, slender, weakly 2-geniculate, the first two segments hispid, the terminal obscurely so; staminate spikelet 6 mm. ion ng, the hairs not as long as on the fertile spikelet. pe in the U. S. National Herbarium, No. reid collected on dry pepo: Itapicuru, Bahia azi October 1919, by G. Pinto (No. 0193). New Species of Sorghastrumn SORGHASTRUM AMPLUM Swallen, sp. nov Culmi 1. ti, caper ligula 3-l. mm, longa, firma; lamin ae lanae usque ad 60 em. longae, 7 mn. latae, glabrae; taf lavcaswthin 35 em. longa, ramis adscendentibus usque a 2 cm. longis, glabris; spiculae 6-6.5 mm. longae; gluma prima villosa, 9- nervis; lemma fertile angustissimum, obscure ciliatun, bifidum, arista 2-geniculata, 2.6 cm. longa, parte secunda’ villosa, Perennial; culms 1.lk m. tall, glabrous, or obscure- ly pubescent at the nodes; sheaths glabrous, the lower shorter, the upper longer than the internodes; ligule -l. mm, long, firm; blades flat, as much as 60 long, 7 mm. wide, glabrous; inflorescence 35 cm. long, ing = > > ed Spikelets light t o-6, aad mm. ae the ent glume Lier villous pe Fee white hairs, the second glume od =) 5 ry ph Oo oO +2) Qe A 14) ja? eq ctr QO he | joe +89) ct i) “ a oO 8 b 5 densely villous rey white silky hairs. Type in the U. S. National Herbar =e: No. 1128238, collected on open hills, S&o Jo&o, S&o Paulo, Brazil, prin: 47 Pie 22, by E. W. D. Holway and Mary M. Holway No. 16 Thi ae is allied to Sgeehestns stipoides, but differs in having a much larger panicle, larg Sa an 96 PHYTOLOGIA Vol. 1h, no. 2 collected at Tremembe, S&o Paulo, by E. W. D. Holway and Mary M. Holway. ‘The fact that these two specimens were collected at different localities and at widely separated times, points to their being a distinct taxa, SORGHASTRUM CHASAE Swallen, sp. nov Culmi scandentes, 3 mm. ongi, ramosi; ligula ad 2 cm. latae, scabrae, ibus scabrissimae; inflorescentia 20-30 cm, longa, ramis adscender.tibus summo hispidis; spiculae 1..5-5 al?; culms clambering in small tree, 3 m. high, branching, rooting at the lower nodes, glabrous; sheaths all shorter than the internodes, glabrous; and branchlets hispid; spikelets 1.5-5 mm. long, — green, the first glume 9-nerved, conspiuously hispid, the second glume 5-nerved, glabrous; sterile lemma thin, hyaline, pilose on the margins; fertile Lemma very narrow, glabrous, bearing a relatively stout 2- geniculate awn 2.5 com. long. Type in the U. S. National Herbarium, No. 1501345, collected in sandy cerrado, Santa Rita do Araguaya, Mato Grosso, Brazil, April 5-6, 1930, by Agnes Chase (No, 11861). : is species is related to Sorghastrum scaberrima as indicated by the hispid ends of the pani branches, but differs in being much larger with broad flat blades and is apparently annual rather than perennial SORGHASTRUM FLEXUOSUM Swallen, sp. nov. i dense caespitosi, erecti, 80 cm. alti; ligule 2-3 mm. longa, decurrens; laminae usque ad 20 em. longae, infra angustae, 3-l, mm. latae, attenuatae, tenuiter scabrae; inflorescentia ca. 20 cm. longa, ramis adscendentibus summo prominente hispidis; spiculae 1.5 mm. longae; gluma prima hispida, 6- nervis; lemma fertile marginibus minute ciliatun; arista 2-geniculata ca. 15 mm . Perennial; culms densely caespitose, erect, 80 cm. tall; sheaths all longer than the internodes, glabrous; ligule 2-3 mn, long, decurrent, firm; blades up to 20 cm. long, narrowed below, widened above to 3-l. mm., attenuate, smooth to the touch, but 1966 Swallen, Notes on grasses 97 finely scabrous on both surfaces and on the margins; inflorescence about 20 cm. long, the flexuous branches ascending, prominently hispid at the ends, rather loosely flowered, glabrous or nearly so in the axils; spikelets .5 mm, long, medium brown, the first glume hispid, 6-nerved, the second glume glabrous, 5-nerved; sterile lemma very thin, hyaline, minutely ciliate on the margins in the upper half; fertile lemma sparsely and minutely ciliate on the margins, bearing a 2-genic- ulate awn about 15 mm. long. Type in the U. S. National Herbarium, No. 2252125, collected on campo, Fazenda Ernesto Schiede, Campo egre, Santa Catarina, Brazil, by R. Reitz « R. Klein (No. 5347). The following specimens, although they are some- what variable, apparently belong to this species: Brazil: Paran&: Guarapuava, Swallen 8862, 8900; Santa Catarina: Porto Unido, Smith & Klein 10771; Campo Alegre, Smith & Klein 71.90, 8531; Lages, Swallen 811; Smith & Klein 8131; Curitibanos, it Klein 8291; S&o Joaquin, Reitz & Klein 7416, 745, 7763, 8156; Bom Retiro: Reitz & Klein 54.85; Rio Grande do Sul: Vacaria, Herb. Bradeanum 21611. SORGHASTRUM RIGIDIFOLIUM Swallen, sp. nov. Culmi e ti ‘ . alti, glabri; ligula firma, 1-3 mm. longa; laminae 25-30 cm. longae, 3 mm, latae, 2 ramis flexuosis, glabris, paucifloris; spiculae 5.5-6 ulata 15 mm. longa. Perennial; culms erect, 1.5 m. tall, glabrous; sheaths elongate, longer than the internodes, gla- brous; ligule firm, indurate, 1-3 mm. long; blades long, the slerder branches flexuous, glabrous, or sparsely hispid, few-flowered, the axis pubescent or pilose at the nodes; spikelets light brown, 5.5-6 mm. from between the teeth of a bifid apex. e in the U. S. National Herbarium, No. 2302971, collected in brushy savannas, Herradura, Pinar del o, Cuba, by E. L. Mus. Botan. Stockholm Plantae Indiae Occidentalis, No. 11603). This species has be erroneously referred to en Sorghastrum stipoides (H.B.K.) Hitche. which grows in South erica from Colombia to Brazil and 98 PHYTOLOGIA Vol. 1h, no. 2 Argentina. It has denser, less flexuous panicles with smaller spikelets, and the awns are shorter, scarcely more than 1 cm. long. The tip of the pan- icle branches are also glabrous All specimens from Cuba assigned to S. stipoides Sea i referred to S. rigidifolium, SORGHASTRUM VIRIDE pec. sp. nov Culmi erecti, 125 cm. alti, glabri, nodis penne tha ranis brevibus ad basin floriferis; spicula 5. 5-6 mm. longa; gluma prima 9-nervis, sparse villosa; lemma fertile angustum, bifidum; arista 15 mm. longa, 2- geniculata, Perennial; culms erect, 125 cm. tall, glabrous, the nodes ecpeaeet ni talk: sheaths all. longer than hee ME ene odes, the uppermost elongate, glabrous; gule 3-l. mm. long, the uppermost reduced, firm, Led ey decurrent; blades as much as 10 cm. long, |, mm. s 5-6 mm. long, green, the first glume 9-nerve sparsely villous, the second glume 5-nerved, glabrous, i. lemma narrow, bifid, awned from between the teeth of a bifid apex, the awn 2-geniculate about 15 mm. long; gic cob -. mm, long. in the U. S. National Herbarium, No. 2378565, ecarostes in open sedge meadow, Campos das Sete Possibly eueres to the same taxa are one speci- men collected at Lapa, Paran& (Swallen ges and another at Lages, Santa Catarina (Smith 1223). ADDITIONAL MATERIALS TOWARD A gens OF THE GENUS CALLICARPA. Harold N. Moldenke CALLICARPA BRACTEATA Dop cept at the base, long-attenuate and decurrent into the petiole at the base, almost glabrous above except for the venation, sparse- ly stellate—pilose on the vein and veinlet reticulation beneath lend fl " cm. long, 3—-l cm. wide, puberulous; peduncles 2— 2.5 cm. long; bractlets te; pedicels very short mm. long, stellate-pubescent, the rim » With very mi: te teeth; corolla violet, 2.5 mm. foie: glabrous and glandulose, the tube dilated, the lobes 4, subequal, 0.5 mm. long, rounded; sta~ t e the mens exse: 3 erted a of corolla- tube; ers glandulose; style equaling stamens; stigma capitate; ovary villous; sw ° of this species was collected by Henri Francois he (no. 2659) at Doi Ngang, Tonkin, Indochina. Dop (1932) ee "Cette @ espéce est voisine du C. longifolia Lam, Elle s'en dis~ tingue aisément par les pédoncules des cymes pape plus longs, les bractées foliacées. La longueur du pédoncules la rapproch~ serait du C. longipes Dunn de Chine et de Hongkong; mais les milieu Nothing is knowm to me of this taxon besides what ‘+ stated in the literature. CALLICARPA BREVIPES (Benth.) Hance, Ann. Sci. Nat., ser. 5, 5: 233. 1866. Additional & emended syno allicarpa longifolia longifolia var. brevipes Benth., Fl. alae ta 270. ‘ie Callicarpa brev. breviceps Hance, Journ. Linn. Soc. Lond. Bot. 13: 116, sphalm. a a nile ° icarpa lo olia - spud Pret, Mem, Sci. Soc. Chi- 1 (3): 45, in syn. 1932 ees C. longifolia Auct., 1965, nor 9 abe 196, nor Blume, 1936, nor Hance, 1932, nor Hemsl., 1916, ~— L., 1820, nor ig » nor Roxb., 1827, nor Vahl, 1936] carpa » dichotoma Bakh. Prei, e Sci. Soc. a 1 (3): 45, in 8a 1932. Callic brecipes (Benth.) Hance apud P'ei, Mem. Sci. Soc. China 1 (3): iB, sphalm. org Calli- brenipes Hance ex Moldenke, R Suppl. 14: 6 1966. 100 PHYTOLOGIA Vol. 1h, no. 2 Bibliography: Lindl., Bot. Reg. 10: pl. 86). 182h; Hook., Exot. Fl. 2: pl. 133. 1825; Benth. Fl. Hongk, nts acd Een ee » Bull. J tenz., s = Sci. Soc. China 1 (1 De 225. 192h3 E. D. Merr., Lingnan Sci. ourn. = gs es Ptei, Mem. Sci. Soc. China 1 (3): "Trea chinal 36 L5—L6, ry 50, & 56. 19323 Pe Dop, Bul « Soc. Hist. Known Geogr. eae o 1, 56, 58, 59, 71, & 1942; Moldenke, Invalid Names 8 & 10. 1 191,25 "Voldenke, Alph, List Cit. Z "530 ( (1948) and 3: 658, hee & 932. 1949; Mol- denke, Known strib. Verbenac » 130, 135 a 1s £177. 19195 olde, etalegin Dertke Gdkireee fa, 7 76. 1952; Moldenke, Biol. Abstr. 26: 1171 + 1952; Moldenke, vi 2h1, 25, & hh3. 1959; Moldenke, "Résumé 2) and 1h: mip & ustrations: Lindl., Bot. Reg. 10: pl. 86h. 182h; Hook., Exot. Fl. at pl. 133. 182 25, Hookets original description (1825) was "Foliis lanceolato- 8 superne serratis adultis glabriusculis » pedunculo peti- ceedingly numerous, and such as to give it a m mealy appearance to the naked eye. Leaves always opposite, 5 or 6 inches in length, lanceolate, somewhat waved, serrated in upper part, the extrem ity nearly entire, dark green above r beneath, the younger ones covered with a stellated pubescence, the old ones pubescent only on the nerves beneath, all petiolated, wi petioles scarcely more than half an inch Bob: 3006 Cymes ax~ » small. peduncles or main stalks s the length of the petiole, the pedicels short, a te, linear-lanceolate bracteas at their base. Flowers small, dro when fully expanded. Calyx small, cup-shaped, with four *short and very obtuse teeth. Corolla subc subcampanulate » 4-lobed, the lobes erecto-patent, of a white colour, iano with pink. Stamens , inserted at the base of the corolla, and exceeding it in length. Filaments white, glabrous. Anthers oblong, yellow. hose ger- men superior, small, spherical; style about as long as the stamens, filiform, white; stigma obtuse, scarcely capitate. ot cots white (Roxb.).° Sent to me by Mr. S Shepherd as a species of Callicarpa which was received by him from China, and which at present has reached only to a height of two feet in the stove of the Botanic 1966 Moldenke, Monograph of Callicarpa 101 Garden at Liverpool. It appears to me to agree in with the figure and p Rc Aoattoade of the C. » ongtfoli of Lamarck, which that author states to be found in Malacca Sonnerat. *Roxburgh's character, above referred to, pena the ves downy undernea this latter = the der foliage the pubescence, if present, is confined wholly to the veins on the under side. The plant described in the Flora Indica inhabits Prince of Wales Island. Very nearly allied to ete sent species, as far as I can collect from the characters, the C. japonica of Thunberg, and the C. rae of Jussi robably belongs to an altogether different genus." ance 8. erii rope Cantonem Page varietate habat Benthamius, cymis multo majoribus, ampli- oribus fructibusqe minoribus ramisque glabris. Stirps shnenckienes est C. brevipes mihi." Maximowicz (1886) cites: CHINA: Kwangtung: Hance s.n. (prov. Canton]. HONGKONG: Hance s.n., Wright s.n. He comments ™ eo video: ramos flavido furfuraceo pubesce entes, folia haud viscida distanter obiter serrata, cymas fere sessiles, flores pauciores majores, antheras magnas partim tan fructus sphaericos piperiformes. In Callicarpa 1 ongifolia Lam. gra recentia valde viscida, cymae laxioris pedunculi longiusculi, flores mmerosiores triplo minores, antherae late ovales decuplo minores, filamenta capillaria longe exserta, fructus san planus P'ei (1932) describes this plamt as follows: "a shrub more or less pubescent or subglabrous. Leaves linear—lanceolate, 9.5 to chartaceous Petioles nearly none, if present, not exceeding 5 mm. in length. Cymes shortly peduncled; peduncles not exceeding 5 m. in length. rudim without truncate, with ) en > glabrous within, about 2 m. high. Corolla -lobed, gl Without, twice as long as the calyx. S ilaments just as long as the c tube; anthers exserted, oblong, 1.5 m. the s mens, glabrous. Ovary densely glandular. Fruit glandular, about 3 m. in diameter......Hooker's pncid pie was based on a living ented Callicarpa plant from China, and he thought that it repres 102 PHYTOLOGIA Vol. 1h, no. 2 lo olia Lamarck. Bentham recognized that the plant Hooker de- scribed was not Lamarck's species and treated it as a varie brevipes. Hance a few years later raised this to specific rank as Callic brevipes (Benth,) Hance." P'ei also says that C. longissima (Hemsl.) — differs from C. brevipes by having long narrow leaves with the leaf-base attenuate. eee Leveillé (1916), es describing his C. mairei, says "Affinis C. Brevi a qua distincta foliis frequenter dentatis et floribus scrngoetg I regard Léveillé's C, mairei as C. bodinieri var. giraldii (Hesse) Rehd. Bakhuizen van den Brink (1921) regards C. brevipes as a synonym of what he calls C. japonica var. dicho~ toma —. ) Bakh. [*C. dichotoma (Lour.) K. Koch]. ~~ ‘The C. longifolia of Bentham and of oer referred to in the synonymy above, belong to the synonymy of C, longi selne pore Merr.; that of Blume, Linnaeus, Roxburgh, and Vahl is C. longifol- ia Lam.; that of amtey and of Diels is, in a C. bodiniert var. giraldii (Hesse) Rehd. and, in part, C, Japonica var. angus~ tata Rehd. ~~ In my 1936 work I regarded C. brevipes f. serrulata P'ei and f. yingtakensis P'ei as synonyms of C. brevipes, but I now regard the former as a distinct form and the latter as a synonym of C. col- lina Diels. Collectors describe C. brevipes as a small shrub, 1 a. tall, the flowers pink, the young fruit iggy ypromine 4 n grasslands and open thickets, flowering in September, fruiting in feo and September. They record the vernacular name — poene"”, The boundaries be- tween the typical form of the species and its several are very obscure and ill-defined. I also have doubts whether this very narrow- almost sessile-leaved taxon is the original of Bentham, who compared his plant with C. longifolia of iil . Bentham's type must be seen in order to clarify this po terial has side: and Ma ntifi preheat in herbaria der the names C, brevipetiolata Merr., C. eriophylla Ridl., and "Ge lo: olia Lam. On the other & Clemens 99 & 5055, distributed as C. brevipes, are actually C. brevipes f. annam ensis Moldenke [the first~mentioned mmber being the type collec- tion], W. Y. Chun 6803, R. C. Ching 2105, F. A. McClure 2664 [Herb- Ganton Chr. Coll. 9222], and Peng, Tak, & Kin 296 [Herb. Canton Chr. Coll. 12295] are C. brevipes f. serrulata P'ei [the second and cotype collections; t-menti being a cotype collection of f, subglabra P'ei]; Herb, Canton Chr. Coll. 12295 12556 are C. longissima (Hemsl.) Merr otype collec- tions of C. brevipes Sa mergeces Pisehy pnd C. Ching 1760 is Ce rubella var. dielsii ( Li [the type collection of C, rubella var. hemsleyana f, sub Preij. Forbes & Hemsley (1890) cite Champion s.n., Hance s.n., Wilford 1966 Moldenke, Monograph of Callicarpa 103 8.n., and C. Wright s.n, from Hongkong, deposited in the Kew and British Museum herbaria. P'ei (1932) cites Chen 5005, 5307, & 5648 from Szechuan, China : * ae all, 17 Westies ipielinios and |, mounted photographs have een me. Additional citations: CHINA: coh oong 4589 nO tl HONGKONG: Ford s.n. [Hongkong] (N); C. Wri Weigh 2.2. 5.0. a. (Hong Ko (W—1,908). CHINESE COASTAL iaieae Hainan: 7 ¢ C. How ae (Bi, S); eMa a 62267 (S), = (La, N); Sa 33985 (N). INDONE~ SIA: GREATER S$ INDA ISLANDS: Sum umatra: Boeea 8017 7 (8). CULTIVATED: England: aeteirg 8.n. ck. Liverpool] (uJ. Hongkong: Herb. opeeye wot Bot. Gar Gard. 8. 8.n. (E--photo, N~-photo, N——photo, S, S, > 3, I CARPA BREVIPES f. ANNAMENSIS Moldenke, Phytologia h: 1. 1952. Bib Liogeenty Moldenke, Phytologia : 1 & 76. 1952; Moldenke, Biol. Abstr, 26: 1171. 1952; Moldenke, Résumé 175 & 43. 1959 Tite fas ere Bate typical f orm of the species from the several other described forms in having its leaf-blades decidedly elliptic, the Sento) ones about 15 cm. long and 6 cm. wide, minutely serrulate from below he middle to below the ter~ minal acumination, attermate-acute at the base. The type of this form was collected by Joseph and Mary Knapp Clemens (no. 4j,\99) at Tourane or its vicinity, Annam, Indochina, between May and July, 1927, and is deposited in the Britton Her~ barium at the New York Botanical oc The collectors describe it as a shrub with lavender—pink fl Mat en mis- eae ee and distributed in haieaetn & as C. brevipes (Benth. ) In “al, , herbarium specimens, including the type, have been examined ned by me Citations: INDOCHINA: Annam: Clemens & Clemens = 499 (Ca 340h16—~isotype, N—type, Ut—136a—isotype), 505! 5055 “(Ca~3L0126) . ALLICARPA BREVIPES f. SERRULATA ig Mem. Sci. Soc, China 1 (3): [Verbenac. cna) 47. 1932 Synonymy: Callic revipes f. subglabra ra P'ei ex Moldenke, — Biot 3: 30, in syn. 1962. : Prei, naog inet) Sci. Soc. China 1 (3)3 [Verbenac. lio China] MT. 19325 Moldenke, Known Geogr. Distrib. Verbenac., ed. 1, » 58, & 86. 1925 Woldenke, Alph. List Cit. 1: 28h (ighe$ and 3: 78. 19195 Molde Known Geogr. Distrib. Verbenac., ed. 2, 130, 135, & 177. 19895 "Woldenke, Fepere 167, 17h, & hh3. 19593 Moldenke, Résumé Suppl. 3: 19 & 30. 1962. This form differs from the ctoaa form of the species in having its leaf-blades ovate-lanceolate, dentate, acuminate at the apex, subcordate at the base, and the fruits 5 mm, in diam- Pred (1932) also says "This form differs from the type by its 10, PHYTOLOGIA Vol. 14, no. 2 leaves which are widened at the middle and distinctly toothed and rounded to subcordate at the base". It was based by him on four collections: (1) Ching 2198, collected at Taisuan, Chekiang, Chi- na, in July 192), (2) Ching 2105 ae) Digg ie at eee meters altitude between Pingyung and T. , Chekiang, July 192) (3) Chun 6803, collected at sarmenas in November 28 and (45 Fe 4. McClure 2 {[Herb. Canton Chr. Coll. 9222], collected at T'ahon, Hainan Island, in April 1922 Collectors describe the plant as a slender, small, woody shrub, 0.8—1.5 meters tall, sometimes scandent, with a few branches, the flowers white or white and and yellow, and the young fruit green or pale-green. It has been found growing in forests, forested ravines, or thickets, in the shade of dense woods, along roadsides, in light w ong the borders of ypersseetl or climbing on rocks, at altitudes of Foor Ace 1800 pak Setorontng § in April, May, and we August. flowers are described mated nd Ve on Peng, Tak, & Kin 296. The type of C. brevipes f. subglabra was collected by To Kang Peng, Ts'ang Wai Tak, and Ts'ang Un Kin (no. 296) in Kwangtung, hina, and is deposited in the herbarium of the University of California at Berkel Callica noe: brevipes “t - serrulata differs from C. longissima (Hemsl.) Merr. (which has ona or subentire leaf=b nlades) - its te ng Cc terial of both names involved, and 2 mounted photographs have n exani by me. Citations: CHINA: Chekiang: R. C. Ching 2105 (W—12h6179— peta k Kwangtung: Peng, Tak, & Kin 296 [Herb. Canton Chr. Neseens Chun & Tso 3,02 (N); | How ow 72007 (Bz—~17295), 7206 ps 17296) 5 8 64235 (N)j F. A. McClure 266) [Herb. gb. nton Chr. ee 9222] (Ca——366337—cotype, N--photo of cotype, Ph-—cotype, “ Rapes ace Wang 36818 36818 (i). HONGKONG: Le ae 14 6803 ed CALLIC. Pog =f alin ca Merr., Philip. Journ. Sci. Bot. 1h: 9 Synonymy: Callicarpa eriophylla Ridl., Journ. Malay. Br. Roy. Asiat. Soc. 1: 8. 1923. Bibliography: E. D. Merr., ina-egg Journ, Sci. Bot. 1s 2h9 1919; H. N. Ridl., Journ. u om Asiat. Soc 1923: A. W. Hill, Ind. Kew. aoe 34 (1926) and 7 %. 19295 Fedde & Schuster in Just, Bot. ae: 59 (2): vt 1939; Mol- , Known Geogr. Distrib. Verbenac, y eel, » & 86. 1966 Moldenke, Monograph of Callicarpa 105 19h24 Meldenke, Alph. List Cit. 1: 25h. 19h6; Moldenke, Known « Verbenac., ed. 2, 143, lah, & 177. 19493; Moldenke, vom a ve gents, lad Gree Moldenks, Résumé 158, 160, 187, 189, 3s 19595 Moldenke, Ri 3: 23 (1962) and ht 7. 1962. » open bush, or shrub, i feet tall, or small tree; RE tye terete, 1.5—-2 mm. in diameter, very densely and uni- h egl pepe eaf-blades c » brittle when letibetlahé or oblong-lanceolate, 7--18 cm. long, 1.5—-7 cm. wide, narrowe: the slenderly a te apex, denticulate or rat rounded or eayeins long; pedice els ; m. long during anthesis; calyx cupuli. form, tomentose; corolla lavender, deep er a old-rose, or rose-purple, infundibular, about 1.25 mm. long, the tube nar- rowed at the base, dilated upwards, the lobes —, stamens ex- frui serted; anthers oblong; oo egg any s dense, subglobose, 1--2 cm. in diam ruiting-pedicels sicat 6.25 mm. long; fruiting-c io eat 2 mm, wide, stellate- alyx pubescent, the rim -toothed; fruits very numerous, crowded, globose, pink or lovely mauve-purple, 2—-2.5 mm. in diameter, black and rugos The type of this species was collec ected by Harley Harris Bart~ lett and Carl Downey La Rue (no. 323) at Kampong Bintang Mariah, Karoland, on the east coast of Sumatra, Indonesia, on August 10, the ppine Bureau phyla and is deposited in the herbarium of the Royal Botanic Gardens at Kew Ridley (1923) scum "Abundant in the open country among la- y shrub with its white leaves and bright as well to 7000 seen flowering in April, June, and August, to and December. The leaves on 106 PHYTOLOGIA Vol. 1h, no. 2 Groenhart 163 are the largest I have seen in this taxon, being to 18 cm. long and 7 cm. wide. The flowers are said to have been "lavender" on Chand 5799, "old-rose" on Chand 1611, and "rose- purple” on Koelz 23422. Yates 1563 is described as a topotype. Material of C Cc. . brevipetiola iolata has been misidentified and dis- tributed in herbaria under the names Cc. cana mbes C. cuspidata Roxb., C. lanata Vahl, "C. pedunculata var.", C, rubella Lindl., and [by Koorders] NGallicarpa sp. nov," artiste 7747 in the U- nited States National Herbarium bears a label with the name Luffa labels. The vernacular name, "mata poena”, on this label may, therefore, not actually apply to our plant. This name, however, is also recorded for C. brevipes (Benth.) Hance. the other hand, the Boeea 8017, distributed as C. eriophyl- ia, is actually C. brevipes (Benth.) Hance. In all, 57 herbarium specimens, including type material, have been examined by me Citations: PAKISTAN: East Bengal: Griffith 6036 (Tf). INDIA: Assam: Chand 1611 (Mi), 3202 (Mi), 5799 99 (Mi), 6225 (Mi); Koelz 23422 (Mi), 26055 (Mi), 32138 (Mi). Khasi States: Hooker & Thom son s.n. (Mont. Khasia) (M). West Bengal: C. B. Clarke 11652 (Bz—-1 ). ‘INDONESIA: GREATER SUNDA ISLANDS: Java: lava: Backer 11383 (Bz—-17676, B2—17677, Bz—-17678, poate Brsnienaa 540 Gondth0)s Burek 107 (Bz~-1767h, B2—17675); Doc n-Rei jnvaan - 1070 (Bz—-17h90); Groenhart 163 2a Cie sath8a) 5 Koorders 29459b [339*] (Bz—-17680), 36198b [1756%) (Bz--17h91), 38202b [255%] (Bz—-17h93), 38262 [1999*] (Bz—-17192); Kuntze 5896 (N); Oosten 33 (Bz—17672); Steenis 20 (Bz--17)88)3 Stoutjesdi jk 148 (Bz--17673). Sumatra: | H. He Bartlett 7747 (W— ene mame tentetee—— eeme 493 (Ba-T oS 2), 5956 (Bz—17553), 2s 17551), 3558 (Bz—-17555), 9029 (Bz—-17556), 9813 (Bz—-17550); Meer Mohr 147— (Bz—-175)8) ; Ouwehand 56 (Ba-~17557) 5 Steenis 5826 (Bz: (Bz 3 Toroes 1220 (N); R. Wind 9 [Boschproefstation BB 9826] (Ba—-17516); He S. Yates 101 (W--1261151), 1563 (B2—17544, Ca— 259330, Mi, N), 2313 (Bz—1755, Ca—-308375, Mi). pear: ea Moldenke in Fedde, Repert. Spec. Nov. 40: 67— > 9 7 : Callicarpa bucherii Moldenke, Résumé Suppl. 1h: 6, in syn. 1966. Bibliography: Moldenke in Fedde, Repert. Spec. Nov. 39: 300 1966 Moldenke, Monograph of Callicarpa 107 (1936) and hO: 67—68, 119, 122, 125, & 127. 1936; Moldenke, Geo- gr. Distrib. om "k. er loldenke, Known Geogr. Di strib. 2 1 196; H. No & A. L. Moldenke, Pl. Life 2: 52. 19),8; Moldenke, Alph. List cit. 2: 487 (19485 and 3: 757 & 867. 19, Known Geogr. Distrib, Verbenac, e Salisb., Ind. Kew. Suppl. 10: 38. 1 1947; Alain in Leén Cuba h: 30) & 305. 19875 Moldenke, Résumé 50 & 3. 19595 Moi tenic, Résumé Suppl. 14: 6. 1966. in ali, 12 herbarium specimens, including the ¥ ore of all the names involved, and ) mounted photographs have been examined by me. Additional Citations: CUBA: Oriente: J. B. Acufia Galé 17877 (Es); Acufla Galé & Diaz Barreto 170 (Es, N); Alain & 1 & Lépez 212 (2); G. C. Bucher Oy (Herb. Roig 965] (Ba-Asotype) ; ro cimaess 6602 (MN), | 7269 (N (N). CALLICARPA CANDICANS (Burm. f.) Hochr., Candollea 5: 190. cea Additional & emended synonymy: Mamanira Rumph., Herb. Amboin kh: 123, pl. 58. 173. Urtica candicans Burm. f., Fl. Ind. 197 (sphalm. "297"). 1768, “Callicarpa cana cana L., Mant. Pl. Alt. 198. 1771 [not C. cana Dalz. & Gibs., 1919, nor Gamble, 1881, nor Wall., 1863]. Callicarpa tomentosa Lam., Encycl. Méth. 1: 562— 563. 1783 [not C. tomentosa a Bakh., - 1932, nor Hook. & Arn., 1918, nor sre 1893, nor L., 7198s nor (L.) Murr., 177k, nor Murr., 177k, nor Thunb., 1959, nor Vahl, 179k, nor Willd., 1808]. Cal- lace americana Lour,. ical Vahl, Symb. Bot. 3: 12. 179k (not | Cc. americana Blanco, 188), nor Hort., 1936, nor L., 1753, nor Lam., 1966, nor Roxb., 1945, nor Sessé & Moc., 1893, nor Willd., = ee Callic cana: foliis serratis subtus tonentosis L Symb. Bot. 3: 12, in syn. 179h. C Callicarpa toment: tomentosa, ptane ovato~lanceolatis acutis serratis, subtus tomentosis albis, baccis 8 nigris distinctis Lam. at Vahl, Symb. Bot. 3: 12, in syn. 1794. Callicarpa macrocarpa Raeusch., Nom. Bot., ed. 3, 37. 1797. Callicarpa foliis ovatis denticulatis per petiolum semidecurren- tibus, subtus Villoso—ca: o-canis, paniculis dichotomis dichotomis Le, apud Willd., Linn. Sp. Pl. 1: 620, in syn. 1797. Callicarpa (c iicarpa (cana ) foliis Serratis subtus tomentosis L. apud Willd., Linn. Sp. Pl. 1: 620, heynii Roth ex Roem. & Schult., Linn. Syst. Veg. 3: 96. 1818. Callicarpa cana; foliis ovatis denticulatis per petiolum Semidecurrentibus, subtus 3 villoso—canis, paniculis dichotomis Vahl apud Roem, & Schult., Lin Linn. Syst. Veg. 3: 94, in syn. 1610. | Cal- licarpa cana; foliis serratis subtus tomentosis L. apud Sims in 108 PHYTOLOGIA Vol. 1h, no. 2 Curtis, Bot. Mag. 47: pl. glk in syn. 1819. Callicarpa sinensis Nois. es Steud., Nom. Bot., ed. 1, 1: 137. 1821. Calli to- mentosa L. ex Spreng, in $a seis. Veg., ed. 16, 1: 419, in syn. 1825. Callicarpa dentata Roxb. ex Wall., Numer . List 87, no. 183k. 1831 [not C, dentata Roth, 1818, nor Sessé & Moc., 190, nor Wall., 1893]. Callicarpa rheedii og Allgem, Med.-pharm, Fl. 3: 829. 1834. Callicarpa sinensis Hort, ex Steud., Nom. Bot., ed. 2, 257. 180. Callicarpa latifolia Zipp. ex Span., Linnaea 15: Bot. Bogor. Alt. 136. 18h). Callicarpa cana Vahl ex Pritz., Icon. Bot. Ind. 2: 55. 1866. Callicarpa bicolor Schau. apud Vidal y Soler, Rev. Pl. Vasc. Filip. 208, in syn. 1886 [not C. bicolor A. L. Juss., 1806, nor F, Vill., 1880) . Callicarpa cana var. a. Saat (Roxb.) H. J. Lani Verbenac. Malay. Arch. 71.1919. Callic cana var, Sertoeri told He J. Lam, Verbenac, Malay. Arch. 7h. 159. Callicarpa cana var, integrifolia f. typica H. J. Lan, Verbenac Malay. Arch. 7h. 1919. Callicarpa cana var. latifolia H. J. he; Verbenac. Malay, Arch. 71. 1919. Callicarpa cana var. latifolia carpa cana benac. Malay. Arch. 72. 1919. Callicarpa cana var. typica H. J. Lam, Verbenac. Malay. Arch. 70. 1919. Callicarpa runcinata Zipp. ex H. J. Lam, Verbenac. Malay. Arch. 73, in syn. 1919. Callicar- pa candicans var. typica (Bakh.) Hochr., Candollea 5: 190. 193h- Callicarpa canescens (Burm. f.) Hochr. ex Moldenke, Alph. List Cit. 2: 358 & 359, sphalm, 1918. Callicarpa candidans (Burm. f.) Hochr, ex Van Steenis, Act. Hort. Berg. 15 (2): he, Senge 1949 Gallicarpe haynii Roth ex Moldenke, Résumé 2); Pi Callicarpa cana Spreng. ex Woldenke, —— sti it & in syn. Bibliography: Rheede & Munnicks, Hort. Ind. Malab. l: pl. 60. 1683; Rumph., Herb. Amboin. }: pl. “eg: ree sce Sp. Pl., ed. l, 1: 121. 17533 L., Mant. Pl. Alt. 198. 1771; J. A. Murr. in | » Bot. tag e Acad, mac. at pl. a1. 1819; Wall Pa Roxb., | gl Ind., ed. i 3 : tee & Wall, ise 06——] 107. 1820; Roth, Nov. Pl. Sp. 82. 1966 Moldenke, Monograph of Callicarpa 109 1821; Steud., Nom, Bot., ed. 1, 1: 137. 1821; Link, Emm, Pl. Berol. Alt. ii: 12h. 1821; Spreng. in L., Syst. Veg., ed. 16, 1: 419 & 420. 1825; Blume, Bijdr. Fl. Nederl. Ind. 817. 1826; 3. Ke . s H. me pera, Mant. a 53, & 55. 1827; Spreng. in L - 1828; an. 29) & 87 185. 1831; Roxb., Fl. Ind., ed. 2 [Carey], 1: 392. 18323; Kostel., Allgem, Mediz .~pharm. Fl. 3 828-829. 18 Decne.e, Nouv. Ann. acques & Jard. Europ.] 3 a c02 . 1845—1862; Schau. in A. DC., Prodr. li: 642—-6),3 & 645. 13h7; Royle, Fib. Pl. India 310—311. °18553 Miq., Fl. Ind. 8 Bat. Suppl. 2h3 & 569. 1860; Dalz. & Gibs. re Ae 200. 18623 Rosenthal, Syn. Pl. oma 30 & 1130. 1862; » Rev. Verbenac. 129 & 137, pl. 8. 1863; Bocq., Adansonia 2: 59 . 137 (1862) and 3: 192 & 263, pl. 8, fig. 12—-22. 1863; apne Rev. Verbenac. 129, 137, 192, & 263, pl. "8, fig. 12—22. 1863; Dec -, Nouv. Ann. Mus. Hist. Nat. Paris 3: 01. 1865; Hassk., Neu "Bokltis . Rumph,. Herb. guboin. 5 Ind., ed. 3 (C. B. Clarke] 131. 187k; Blanco, Fl. Filip. Atlas Ba 427 bis. 1878--1880; Gamble, Man. Ind. *Timb. 283. "1881; F Muell., First Census 103. no PHYTOLOGIA Vol. 1h, no, 2 India 3: 260. eg | F. N. Will., Bull. Herb. Boiss., ser. 2, 5: 430. 1905; Bailey & Mill., Cycl. Am. Hort. 1: 217. 1906; E. D. Merr., Philip. Jo 9 Wright in Cassell, Dict. Pract. Gard., ed. 2, 1: 156 & Gamble, Journ. Asiat. Soc. Beng. 7 (2) extra = 802 & 807. 1908; King & Gamble, Mat. Fl. Malay. Penins, 21: 1016-1018, 1909; C. K. Schneid., Illustr. Handb. Laubholzk. 2: 59h. 1911; Craib, Kew Bull. Misc. Inf. 9: 2. 1911; Gerth van Wijk, Dict. Plantnames 1: 217. 1912; Craib, Contrib. Fl. Siam Dicot. 163. 1912; Jacks., Proc. Linn. Soc. Lond, 12), Suppl. [Ind. Linn. Herbd 49. 1912; Elbert, Meded . Rijks Herb. Leid. 12: 15, 1912; Koord., jonsfl. Java 3: 13h. 1912; F. M. Bailey, Compreh. Cat. Queensl. Pl. 382. 1913; L. H. Bailey, Stand. Cycl. Hort. 2: 629. 1914; H. J. Lam in H. gue Meded. Rijks Herb. Leid. 37: 32-- 3h. 1914; Gerth van Wijk, - Plantnames 2: as 1916; Heyne, Nutt. Plant. Nederl. Ind. ol of, 1917; Ewart - North, : gee" Phili 362—~383. 1923; H. N. Ridl., Fl. Malay Penins. 2: 61h, 616, elk ates Chung, Mem. Sci. Soc. China 1 (2): 225. 192) He Je » Engl. Bot. Jahrb. 59: 88—89. 192); S. Moore, nin Bot. Lond. 2 suppl 80. 1925; fgg a Nutt. Plant. NederL Ind. ch ies 1927; E Lingnan Sci. d ourn. S: 157. 19275 Domin, Bibl, Bot. ry (oye 1108 & 1109, text fig. 179. 1928; Stapf, Ind. Lond. 1: 525. 1929; A. W. Hill, Ind, Kew. Suppl. 7: 36. 1929; L. H. & E. Z. Bailey, Hortus 112. 1930; C. A. Gardn., ] 15, 1932; Kanehira, Fl. Micron. 156, 1933; Crevost & Pételot, Bull. Econom. Indo-Chine 37: 1289—1290. 193h; Hochr., Candollea 5: & 33. 1942; uw " Muses 2: 7h, 78, 83, & ae a ie == 1945; H. St. John, Bot. Club 7a 288. 19 , Al ° Bull. Torrey a 1: k, 3h, 5h, 60, 108, 120, 161, 198° 2355. & 277. 196; Moldenke, Phytologia 2 . 19h73 Moldenke, Bol. Soc, Venez. Cienc. Nat. 11: 37. 19h7; H. St. John, Pacific ag 2: 273. 1918; wet pp - L. Moldenke, Pl. Life 2: 64 & 78. 1948; Moldenke, Alph. Cit. 2: 358, 359; ok, ko8, 121, L3k, 47, 490, 556, $61, 565, 565, 1966 Moldenke, Monograph of Callicarpa 11 581, 58h, 63h, 643, & 6a (19h8), 3: 1558+ 10, 765, 810, 852, 877, 878, 932, & 957 (1909), and bs 987, 63, 1118, 1206, 199; Moldenke’ a “Hy + aat—ue. 191,95 Mold a gneg — Distrib. Verbenac. » Od. 2, 12h, 125, 130, L. Mold Ma) | 3. 1950; Moldenke, Phyto- logia 3: 29h & 296 (1950), kh: 83 & 121—125 (1982), and 6: 215. 1958; F. R. Fosbe c Sci. 12: ; Fosberg, Pac : 20. 1958; Moldenke, Résumé 158, 160, 168, 17h, 175, 177, 179, 182, 185-~167, 189, 191—196, 200, 203, 208, 213, 2hl—2h7, 319, 355, & h3. 19593 Anon., Kew ; sumé Suppl + ( : 23: 87. 1963; Backer & rts Fl. Java 2: 601. 1965; Moldenke, Résumé Suppl. 13: 6 (1966) and 1h: . 3, 6, & 7. 19663; Moldenke, Phytologia 13: 26, 439, 475, 478, 99, & 502. 1966. Monac ° * Verbenac.] pl. 8, fig. 12—22. 1863; Vi in, Fam Lefi, Filip. pl. 3h, fig. C. 1883; Domin, Bibl. Bot. 22 (89): 2 sateetion referred to above is said by Stapf (3929) to be in color, but the copy in the library of the York Botanical Garden is uncolored. King & Gamble (1908) mend: that "Clarke in Fl. Br. Ind. does not quote Bot. Mag. 2107, which Bentham had said probably represented a much more woolly With this Plant, though it might be a@ var. of cana. th we ietnest It should be noted here tube the C. americana of Blanco, re- ferred to in the synonymy above, is @ synonym of C. formosana Rolfe, that accredited to mi ee "is Cc. longifolia : -_ am., that of Linnaeus i is a valid species, while that of Lamarck, of Roxburgh, and of Willdenow are C. americana L., and that of Ther & Mocifio is C. pringlei Brig. “The C. bicolor of A. L. Jussieu is a valid species » While that of Fernandez Villar is C. formosana Rolfe. The C. cana of Dalzell & Gibson is C. tomentosa he) Murr., that of pedunculata of Roth is C. pedunculata R. Br., that of Seasé & Mociflo is Corm- tia andifolia (Schlecht. & Cham.) Schau., and that of Wallich is C. longifolia Lam. The C. tomentosa of Bakhuizen van den Brink is in part C. arborea Roxb. and in part C, integerrima Champ., that of Hooker & Arnott and that of Willdenew are C. loureiri Hook. & Arn., that of Ktnig is C, macrophylla Vahl, that accredited to Lin- naeus is C. erioclona Schau., that of Murray is = tomentosa (L.) Murr., a valid species, that of Thunberg is C. longifolia La Lam., and that S Vahl is something whose identity has not yet been estab- shed, In my 1936 work I mistakenly included C. bicolor A. L. Juss., C. ne PAT#TOL06.14 Vol. lh, no. 2 ana var, eer aH. J. Lan, C. cana var. integrifolia hs Giabcinaat a H, “% sot C. cana var. sumatrana (Miq.) H. J. La, C. chinensis Hort., and C. suma sumatrana Mig. in the synonymy of this Hayek, and ¢. var. sumatrana (Miq.) Moldenke, Lam (191), 1919) also 0 included C. bicolor A. L. Juss. in the synonymy of C. cana, while King & Gamble ble (1908) and Domin (1928) included both C. bicolor and Ce. — ee acuminata Roxb., given as a synonym of C. cana by Steu- del "CSB: is actually C. mdiflora Hook. & Arn.; the he Ano: baccifera, verticillata, - folio molli, & incano Pluk..and pn? variant orthography, ionehae baccifera, verti verticillata, folio gt incano Pluk., given as synonyms by Loureiro (1790, 1793) | aides in the Synonymy of the true C. americana L.; the C, wallichiana Walp., — as a synonym by D Dalzell & Gibson (1860), is C. tomen- tosa (L.) M "Gatiens. Oreiced & Pételot (193) seem to maintain C, ameri~ cana Lour., C. cana L., and C. facet Kostel. all as valid spe- cles, with only c. eakoes 1 Lam. as a also dates the original publication of C. adenanthera errone~ ously as "1827", that of C. dentata as "1828", that of C. rheedii as "1836", and that of C. sinensis as "181", The original publi- teing of C. os. is Lides sarees given as "Roth, Nov. Pl. Spec. 82. ntly i The "Journ. Roy. katie Soc. Bengal mh ore 1012 & 1018" (1909) * reference sometime King & Gamble lliography above is eemaninet 22 in- priya EY) ee. —- actually, pages 1119 onwards in this work that were published in Yi36. "The Hassk., Cat. Pl. Hort. Bot. Bogor. nt. nae (1844) reference is sometimes raha as "Hassk., Cat. Lands P: itenz. 2: 136", al e title. "Roxb., Fl. a = 1 (Carey & Wall.], i: 392 (1820)" is sometimes given as a reference to C. candicans, but in error since this taxon is not mentioned on that page. The type of C. candicans was collected in Java and its leaves impo mentosis., Habitat in Java. h Frutex ramis compressiusculis, albido-tomentosis. Folia ceponi te, petiolata, ovato-lanceolate, serrata, spithamea, supra nervis subtus tota tomen: tosa, Panicu- 1966 Moldenke, Monograph of Callicarpa 113 lae axillares, dichotomae, vix foliorum longitu tomentosae: Bracteis mimutis j subulatis, ad singulan ok Gia Eeanal oppositis. Calyx cylindricus, vix quadridentatus, laevior. Coro b- uliformis, quadrifida. Filamenta capillaria, corolla duplo lon- giora, Germen psa Stylus statura filamentorum. Sti capitatum, obtusum Vahl (179)) says: "In editione Syst. Voxet. MURRAYI omittitur ha a Callic lanata tome coopertis, denique denticulis marginis majoribus in omnibus +e] is, nec ut in Callicarpa lanata, in aliis foliis mllis in obsoletis. Facie ad Callic. americaram propior accedit, cate acute denticulata, paniculae parvae minus ramosae, ramis non di- oblongae, conveniunt coeterum figura foliorum." reference to the fruit as a capsule is amazing. Roxburgh's account (1820) of C. cana is of great interest: glandular—dentate-s 3 -This shrub Garden from the Moluccas in 1798, and in three years had attain- 6d to the height of from four to eight feet, considerably parts downy and the ligneous ones covered with a smooth ash coloured bark. They are in blossom in March ar—dentate-serrates upper side soft _ of a deep green colour, but very » and pale underneath. Cymes axillary, dichotomous » scarcely longer rs the satholan, Villous. -~ Flowers numerous, small, pale red colour. — Bractes length of the filaments. Stigma two-cleft. -- Berry round, smooth, deep purple; seeds four......1t differs from Cc. anericana, Willd. in naving a woolly toothed calyx, two-lobed cer and in the form of the leaves." Link (1821) comments under C. cana L.: "Non distincta est C. tomentosa W. E. 158", Actually, however, C. tomentosa Willd. is regards C, adenanthera R. Br. and C. 114 PHYTOLOGIA Vol. 1h, no. 2 giving C. acuminata Roxb. as a synonym of the latter! However, I re eget C. acuminata peas as a synonym of C. nudiflora Hook. & | very different plant. fosteleteky (1831) notes "Auf Bergen in Ostindien -- Blttht dort fast stets......vorztiglich in MYrz und arbre ape Bl&tter al Absond adhe mighonls co Other records of economic uses are known. Rosenthal (1862) says "Die Blatter worden vorzugsweise bei Flech- wirksam gehalten.....Wurzel und deanery “en gegen Fieber, Leberleiden, Flechten, Krttze u.s.w." 1917) says "Struik met geneeskrachtige bladern (De Clereg No. oa roy Volgens Filet... appliceert men de bladern op wonden om gwelling te voorkomen; te Batavia gebruikt man ze tegen steenpuisten." Lam (192) reports anes "die Monghinae es: Bl#tter werden zum Bettuben von Fischen an- and the fi a specimen at the Linnean Society, nh so named by Dr. Wallich, It is quoted with a query by Dennstedt for the Callicarpa phylla of Vahl." He i ferring here to plate 60 in Rheede's Hort. Ind. Malab. (1683). The reference here to the Linnean Herbarium is of interest because ens spec C. cana. Actually, I have examined the material in the as “cana 7?" by Smith (according to B. D. Jackson), but it is actu- ally C. | nudiflora Hook. & Arn, However, specimen number 3, com- pletely unnamed, with the single notation "Ind. or." is the true C. cana L, Perhaps it is to this specimen that Dillwyn refers as having been identified as C. cana by Wallich; if so, Wallich made no notation of his identification on the sheet. Perhaps the a (also enforced when I examined the material terial) that seine eben ed on any of the sheets was in force & Mueller (1870) say "The species istered over the In- dian Archipelago to the Malayan peninsula and the Philippine Is- Schauer with doubt to C. longifolia Lam, appears to me to be the true C, cana." a mmte G8 e (1881) onde "Common in forests and along roadsides in probably aothlenrds. te. the Ganges. oy sie pectty tie Fie It is, however, probable that he is 1966 Moldenke, Monograph of Callicarpa 115 here referring to C. macrophylla Vahl, rather than to C. candi- cans. This applies also to his description of the bark and wood. Balfour (1885) curiously regards C. cana L., C. tomentosa L., and C. americana Lour. as synonyms of f what he calis Cc. lanata L. (now known as C. tomentosa (L.) Murr.]. King & Gamble (1908) give a detailed . description of what they call C. cana, but which may contain elements that apply ase to C. macrophylla » With which these authors apparently at leas sometimes pontuant Cc. candicans. "A shrub, branchlets, al beneath, and inflores- tomentose when young, afterwards glabrous, shining and dark- coloured, glandular-punctate; lower more or less whitish-grey stellate-tomentose, sometimes rugose, but more usually softly and conspicuously hairy, glandular-punctate beeen the tomentums; » exc at uneate here Tinga ep ent above; main nerves 10 to 12 pairs, impressed above, stiestiy ra raised be- neath, starting at an angle of ho to 45° with the midrib and curving to the margin, joined by re transverse near- ly parallel dole .25 to .75 in. long, upper part bord ered by the d ecurrent 6 blade s Ovary somewhat depressed; style sa Beg am often twisted, as long as filaments; stigma funnel-shaped. Drupe globose, under .1 in. long & in diam., 3 Pyrenes h. pra Binaries Teysmann. ested that it might be a species Cali I consider “that T Teysmann is correct in his suppo- ssttae, as the description is unmistakably that of a Calli while the yaetl is a fair representation of Callicarpa ae for the ve drawn inflorescences. I am of the opinion that the widely distributed Calli cana Linn, was the 4, es. nothing approaching this ‘Species is a Amboina a Sortectenaer” axtioast Rel. - hehe 2465, from Macas~ Sar, Celebes, is unquestionably referable to it. re thes field work in Ambo Amboina will doubtless clear up any doubt there may exist as to the exact identity of Mamanira." 16 PHYTOLOGIA Vol. 1h, no. 2 Lam (1919) says "In our opinion, C. cana is a very polymorphous species, the varieties of which show an often uninterrupted series of ry forms between each other. Though there are certain species, the affinity of which with C. cana is evident and even close, their general habit makes them distinctly different from the present, genase » the varieties of which are conform in nalts the length s and peduncles, and the size of the leaves." This = el Na SP org worker divides C. cana into numerous Fisrie- 9 ips categories and cites (192) numerous specimens of subspecific taxa I will discuss elsewhere, but "those that I renee as belonging to typical C. candicans should perhaps have his de- scriptions and citations ; reproduced here: var. typica H. J. Lam (C. heynii Roth, C, latifolia Zipp., in part) — "folia membrana~ cea vel subchartacea, late ovata vel ovato-rotundata usque ad sub- stellato-tomentosa, basi integra, ceterum serrata, nervis sec iis urtinque osscther 10; 71h cM. Anew, , 1/2--8 cM. lata, peti- = 0.6-—2.8 cM. longo." -—~ WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Luzon: Cuming 1283. INDONESIA: GREATER SUNDA ISLANDS: Celebes: Teijamann & De Vriese s.n. (Le—~908.267-1061, Le—908. 267-1062). Java: Elbert 20467, Forbes 1252, Junghuhn 1 Kor=- thals 580, Mousset 1055, Waitz s.n. (Le—908.266-1230), Z linger 157. Kambangan: D De Vriese s.n. (Le—908 265-1459), sung 3 SNe (Le—-908 +265-380). LESSER SUNDA ISLANDS: Lombok: Elbert 20 es cmenaee! Seen? Some! ne CLL, gl prope suffulta; nervis secundariis utrinqe 8; 7--15 1/2 CM. longa, 2 1/2—5 cM. lata; petiolo 1.2—-2.5 cM. longo; co- glabra." -- WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: pean “Reillo 19265. PALAU ISLANDS: Koror: Ledermann 11,126, Ray- mundus 189. INDONESIA: GREATER SUNDA ISLANDS: Celebes: Collector undetermined s.n. (Le—-908 .266-1231, Le—908.266~-1232). "As in var. @ [sumatrana], it is distinguishable from C, longifolia by i ee. tomentose never floccose texture of the lower side of Var. latifolia H. J, Lam — "folia valde membranacea, meee o- vata, mumquam mquam tetragona, basi attemata, . longiuscule ac inata, margine ut in var. @ [typica], crenata, vel tiaertete; adulta pig glabra, subtus densius ee stellato-tamentosa, nervis secundariis 8-10, 13-19 cM. longa, 6—~11 cM. lata, pet- iolo 2-3 a. longo," Var. latifolia f. typica H. J. Lam (C. latifolia Zipp., in part) — "flores tetrameri". — WESTERN PA PACIFIC ISLANDS: PINE ISLANDS: Negros: E. D. Merrill 207. PALAU ISLANDS: Yap: Vol- 1966 Moldenke, Monograph of Callicarpa 117 kens 439. ‘INDONESIA: GREATER SUNDA ISLANDS: Sumatra: Collector undetermined s s.n. (Bz——l381). LESSER SUNDA ISLANDS: Lombok: El- bert 736 & 1992. Timor: Decaisne s.n. (Le—908.265-299), ae 5m. (Le—~908 0265-937, Le--908. 908 .265-938, Le—908. 265-917) « ESIA: BISMARK ARCHIPELAGO: Hermit: Kramer 51. ATE de es f. SESS 3 H. J. Lam — "flores pentameri" .— LLECTION ETERMINED: Collector undetermined s.n. Cinen300,2os-1108, L908 MSeLhhS as haces. sos Vea oe ar. dentata H. J. Lam (C. runcinata Zipp.) — "folia membran- acea vel subchartacea, basi attermata, acumine longiusculo, vel — subtruncata, adulta supra glabra, subtus aaxineel le stell~ entosa, margine basi integra, hae grosse e6 iter serrato-dentata, nervis secundariis utrinque 78, 7 1 7 cM. longa, 3--7 cM. lata, petiolo 0.7=-2 cl + longo." LESSER SUNDA ISLANDS: Sumbawa: Elbert 3505 es Timor: Zippel ea (Le—908 .266-13). pies if Var. integrifolia H. J. Lam — "folia membranacea vel subchar- su na dulata vel subintegra, adulta supra glabra, opaca, subtus brunneo- luteo- vel flavido-tomentosa vel cate cede | corolla pilis in vittis 4 in lobis positis vestita vel glabra Var. integrifolia f. typica H. J. Lam — eaika subtus dense tomentosa; corolla pilosa." — WESTERN PACIFIC ISLANDS: ISLANDS: Saipan: Fritz s.n. (B), Volkens 1). Tinian: Schnee s.n. se PALAU ISLANDS: Yap: Volkens 210. MICRONESIA: CAROLINE IS- : Truk: Kraemer 36. MELANESIA: BISMARK ARCHIPELAGO: Hermit: ETS 10. Var, integrifolia f. glabriuscula H. J. Lam — "folia subtus glabriuscula; corolla glabra." ——- WESTERN PACIFIC ISLANDS: MARIA- NA ISLANDS: Saipan: Fritz s.n. (B). INDONESIA: GREATER SUNDA IS- LANDS: Salajar: Weber s.n. (Le—908.267-916). MICRONESIA: CAROLINE ISLANDS: Truk: Ledermann mn 14019. In discussing his C. cana var. latifolia f. pentandra, Dr. Lam says "In some cases, like this, Callicarpa may be pentamerous, and in this point, form a transaction [transition] into Geunsia. The always gare and alternate. As there are 5-merous Callicarpae, 50 peat also l-merous Geunsiae. Yet there are no doubtful ke, or C, erioclona f. glabrescens Moldenke. Unfortunately, I have not as yet had the opportunity to examine the specimens he cites, 118 PHITOLOGIA Vol. 1h, no, 2 and, ae ee the specimens which I cite have apparently not been seen by him, I would be not at all surprised if his C. cana var. integrifolia and var. integrifolia f,. typica should prove to be C. erioclona or C. acicetoun var. paucinervia, and that his C 4 aS f. a ’ ee a change in name for the latter. allicarpa cana cana var. repanda Warb. is certainly C. erio- clona. ~~ Grevost & Pételot (193) refer to a "C. americana L." from In- dochina, but certainly mean C. americana L Lour., which is a synonym stella » on the upper surface (often with the exception of the large nerves) glabrous, oval~elliptic-oblong-obovate, with a contracted, decurrent bas obtuse or acutely to obtus uminate, serrate—de 5-25 cm by 2 1/2—15 cm; petiole 3/l— 3 1/2 om. Cymes on 1/2—1 cm long pedunc ns _ cm across; c. 11/2 m high, densely stellate- 3 teeth mimte, obtuse; corolla violet, 2 1/2--3 m high, rather deeply lobed; tube glabrous; segments ro n unded, with a eat honen? glands; a c. mm; drupe globose, dark vi- t Erec Recent eeitestare daaasthe tne plant as a small, erect, woody os» or subshrub, 0.5--l, m, tall, scnstilis only H emi-woody, or tree; trunk 0,6—1 cm. in’ diameter; bark subaromatic and anmal » Whi brown veins beneath; flowers ill smelling; pedicels light green- ish-brown; calyx light coat cena corolla varying from white to rosy, red, purple, or "li ght-violet, green within"; stamens purple or purplish; filaments light-violet; anthers Light-yellow; style Light-riolet; stigma very light~violet; fruit green or pale- immature, later deep-violet, dark-purple, purple-black, or one "etnies, Koorders 27608b has two of its leaves deeply es Weis eee a the leaves coarsely serrate. The fl ers on Ts'ang 515 are described as "red", on Yates 25 as mpurple", on Agama 1050 ee as "rosy to purple", and on Tsiang 873 873 and Ying 873 as “white Watt “(a589) makes the Sol souing comments about this plant: "A shrub of Bengal; common in forests and along road-sides in the to plant as reporting of this fibre, says: 'It is much too weak for either sailcloth or cordage. It, however, possesses all the free and 1966 Moldenke, Monograph of Callicarpa 1g ndly nature of flax, and even omnne like flax. It is easily worked, with little or no waste, ; Recent collectors have found the te ant growing in grassy wilds, brushwood, light forests, scrub jungles, thickets, ravines, open places, steppe-like grass formations, sandy soil, and loan, near streams, at the edge of jungles, along roadsides, on dry hills, hillsides, open slopes, ont ground, and level land, in open places, and on the strand, at altitudes of 7 candicans grows on "Ebene und Gebirge". Prain (1905) reporte.” c "namanirang "méniran kasar", "meniran kébo", "mSniran oetan”, "wéniran utan", "palis# , "papalsis", "poeltak-poeltak", Mred-fruited tampang b¥si", "resepo", "Rheede's Wirbelbeere", "roidi", "sai fa min", s 5, *tampah b¥si", "tampang b&si", "tampang bési merah", "tampong bési puteh", "tapoeng-tapoeng", "tfigau", "t{gau-na-itim", "tiptipinagut", "toembar bési", and "tfibang-dalég". It is very probable, however, that some of these names apply to what we now call var. sumatrana, C. erioclona, or C. erioclona var. paucinervia, which see. The bSsi" is applied also to Vitex vestita Wall. The fiber of C. candicans, as has been noted above, e, is know as “aroosha fiber" in India. It is of great interest to note me various authors have given the distribution of C. candicans (or C. cana, as most of 1903 case WSouthern Asia, Malayan region, to Australia"; King & Gamble (1908) “Malay Archipelago, gg* i S Sumatra, Banka, Java, Timor, Saleyer-Ins., Minahassah, Luzon, Negros, Hermit-Ins., Palau-Ins,, Neu-Mecklenberg, Yap und Truk 120 PHYTOLOGIA Vol. 1h, no. 2 Ins., Karolinen, Saipan und Tinian (Marianen), N. Australien" {this obviously includes C, erioclona and C. erioclona var. pauci~ nervial]; Lam (1919) "Calcutta, Malacca!l, Malaya (Sumatral!, nr: Javal, Timor!, Saleyer-Isl.!, bthelasiea! ): Philippines (Luzon Negros!), Bismarck-arch, (New-I Ireland, Hermit~isl.!), palatal. ‘, Marianne- Li isl.!, Caroline-isl.!J, Bourbon (Hallier)*" [agai obvious- ly including c. erioclona and Cs erioclona var. pauc taaeviali Bak~ huizen van den Brink (1921) “Yalabar, Silhet, Chittagong, eget Malacca, Cochin-china, China, Philippines, Malay Archipelag Guinea, Polynesia, Australia, & Bourbon (? cult.)"; Ridley art 923) “open country, edges of woods and clearings", Pahang, Malacca, Negri Sembilan , Foret, Wellesley, ere & Perlis » “Malay islands Mi hilip- pinen ustralien jutmen sealant is oe GO " fhe pip two of cg unnumbered collections of Febraury 1910 from Queensland o notes oF toe "forma vel varietas foliis minoribus, circa - longis ——-3.5 cm. latis excellens"]; L. H. & E. 105 (Ba— 17391), 2783 (Bu—-1707), 3436 (Bu—17333), 3625 (Bz—17393) , 3667 (B2—17339), 3779 (Bz——17302), 3822 (Bz—17301), 4736 (Bz-- 17361), 4764 (Bz--17360), 14806 (Bz--17303), A.302 (Bz—-17307) 5 Birkhoff II (Bz—170h); Blokhuis s.n. (Mei 1919] (Bz--1733h, Be— 17335); Boer 3515 (Bz—~17369); Brinkman 10 (Bz—~17321); Broekhui-~ jsen 15 (Bz--17305) ; Burck & Monchey s.n. Sen. (Buitenzorg] (Bz—-17328, Ca-—265961) 5 er 5620 (Bz—-17327)3 Bttsgen 97 (B, Bz——-17381); Buwalda 7212 (ean 70857), 7555 (Bu--72895), 7502 (Bz—-72896) ; Clason C.87_ (Bz-~-17309), D.10 (Bz—~17310) ; Clason-Laarman 966 (Bz- 17314); Cor Cordes 16 (Bu—17l02); Den Hop 35 (Bz--17386); Docters F Leeuwen-Rei jnvaan 773 (Bz--17363, Bz—1700) , eee ee — 17368), s.n. [1 Juni 1909] (Bz—-17338); Edeling s Tjira] (Bz—17373, Bz-~1737h); Elbers s.n. [Mai art capac Af Hallier 269 (Bz—17330, Bz—17331, Ut—53170), sen. [16.VIII 1896] (Bz—-17329, Bz—-17332); Hardenberg s.n. (Bz—17)03); Herb. Hort. Bot. Bogor. s.n. [Japara] (Bz—17401), son. (Bz-—17323)35 Jensen sen. (Cp); K Karta 147 (Bz—-17428) 5 Koorders 9706b (Ba— 17128); Koorders 9706b (Bz--17h2h), 21928b [91*] (Bz—-1709) , 21998 [39%] (Bz—-17)08), 22505b [2348] (Bz—17h11, Bz--17h12), 26211b [15)*] (Bz--17h21, Bz—17422), 27608b [588*] (Bz——17L10), 28L71b [738] (Bz—17i19), 28h78> [648%] (Bz—17i20), 29595 (Be— 17118), 29935b (Bz—17h25, Bz—-17426), 32377 [2628*] (Bz—17h23), 33682b (Bz—17416, Bs—17L17), 35734b [2168*] (Bz—17b1h), 35735b (Bz—-17L15), 426147 [365*] (Bz—17h13); F. Kramer 32 (Bz——17377, Bz—17378); L8rzing 783 (Bz—-17376), 983 -(be—17375, Ba-—17379) 5 Mousset 1055 (Bz—-1705); Oosten 15 (Bz—-17306); Ploem sen. [Java oceid.] (Bz—1736h, Bz—-17365); Rant hh (Bz—17311, Hace Z-— photo); Scheffer s.n. (Bz—17366, Bz--17367); Thorenaar 319 (Bz— 17336, Bz—-17337); Thunberg s-n. (S); tée 9 dia toed ae (Bz- 17362); Van Steenis 5299 (Bz--17315); Vorderman s tavia] (Bz—17370, Bz—-17371, Bz—-17372); Wisse Wisse 19 Tp (isi 7385), 794 (Be— 17325), 827 (Bz—1730h). Kangean: r 26851 (Bz—-17130), os (Ba —171]0, B Bz--17hh1, Be—172, sims y, 277k (Be—17 Bz——1 7445), 28113 (Bz—17h31) ; Dommers 9h (Bz —17138) , 166 (Bz-- 17437). Madura: Backer 18943 (Bz—-17uh6), 19512 (Bz—17uL9) , 19800 (Bz—-17447), 20899 (Bz--174h8). Paliat: Backer 2 29521 (Ba-— 12h, PHYTOLOGIA Vol. 1s, no. 2 17436). Saboenting: Backer 29790 (Bz—-1739). Saoebi: Mahl- meister 21 (Bz--17),29, Bz—-17432). Sepandjang: Backer 28803 (Bz- 17435). Sepapan: Backer 28438 (Bz—-17433, B2-—-17h34). “Sumatra: De Vriese s.n. (K). LESSER SUNDA ISLANDS: Soemba: Iboet 292 (Bs- 17450, Ut--1698a). Sumbawa: Rensch 769 (Bz—17h51) . Timor: Cas- tro 2 (Bz—17457, Bz--17458); Grijp 2 (Bz--17452, Bz--17453)3 Teijsmann 8922 8922 (Bz-~17)56); Voogd 2255 (Bz—-17454). AUSTRALIAN REGION: AUSTRALIA: Queensland: Collector undetermined s.n. [Edge- water Bay] gence Macgillivray 76 (Go). AUSTRALIAN ISLANDS: - White 10166 (N, N). CULTIVATED: France: Bouton Sone [1835] ain Herb. | Hort. Malmaison s.n. (B); Herb. Hort. Paris 8. Se n. (Br); Herb. Kum Kummer s.n. \. (Mu—1)29). Germany: Herb. Kummer 8.Ne ‘(u—1,30) ; Herb. >. Schwégrichen son. (Mu—1)28). India: Wallich 328 (Cp), sen. ne (CP, S). Java: Herb. Hort. Bot. Bogor. s.n. (Bz 17299). Réunion: Richard s.n. [Jard. Bot. de Bourbon] (P, P, P)- Trinidad: Herb, Hooker 3.n. rete LOCALITY OF COLLECTION UNDETERMINED: ane undesignated 30 (Q); Herb. Alstroemer iv (S); Herb. Burman s.n. (Le); Herb, Lugd.-Bat. 908.265-1uuh (Le), 908 265-1h¢h (Le) (Le); A. | 7% "Richard 67% (Du-166515). MOUNTED ILLUSTRA- TIONS: Ferd, Bauer, Icon. Nov. Holl, 96h (V), 96a (V), sen. (V). CALLICARPA CANDICANS £. LACINIATA Moldenke, Phytologia kh: 125. 1952. Bibliography: Mol » Phytologia h: 12h & rate 1953; Molden- ke, ana Abstr. 27: em ets 1953; Moldenke, Résumé 1 & bh3. 1959. *t orm differs from the typical form of the ae in ee its leaf-blades ee along the margins. The type of the form was collected by Maria Ernestine Walsh- Held (no. 474) at Nipel, Timor, on April 18, 1929, and is deposi- ted in the Herbarium Bogoriense at ltintaeres Thus far the tax- on is known to me only from the type aang oe “ie vente 1 her- barium specimen and 2 mounted photographs have be Citations: INDONESIA: LESSER SUNDA ISLANDS: whats Waleh-Held L7h (Bz--17455--type, N—photo of type, Z~-photo of type). CALLICARPA CANDICANS var. PERRYANA (Dop) Moldenke, Phytologia 2: + Callicarpa cana var. perryana Dop, Bull. Soc. Hist. ography: P. tig 28 ages - Hist. Nat. Toulouse 6): — 1932; Moldenke, Geogr. istrib. Verbenac. 2 a 1, 59 & 86. 192; Moldenke, Pigtologia hg 310 & 34. 19h7; denke, Known pe ie Distri rbenac,, ed. 2, 138 & 177. iols; Moldenke, Ré- sumé 1 , a2, ‘ke “3. 1959. ty differs from the typical form of the species in navace. ite 1 leat-blade obovate or ovate-elliptic, rounded and shortly acuminate at the apex, and conspicuously and irregularly serrate along the mer portions of the margins 1966 Moldenke, Monograph of Callicarpa 125 The type of this variety was collected by G. E. Perry — in whose honor it is named -~ at Poulo Condor, Cochinchina, Indochina, i nothing of the taxon except what is said of it in the lit- erature. It is apparently known only from the original collection. nay lawn rye var. SUMATRANA (Miq.) Moldenke, Phytologia ks - 1952. Synonymy: Callicarpa sumatrana Miq., Fl. Ind. Bat. 2: 886. 1856. Callicarpa chinensis Hort. ex C. K. Schneid., Illustr. Handb. Laubholzk, 2: 59), nom. md, 1911. eed cana var. sumatrana (Miq.) H. J. Lam, Verbenac. BEAT oc eee 1919. meee nceenaes Sims in Curtis Be hv: ple pits 1819; Miq., Fl. Ind. Bat. 2: 886. 1856; be ot in Hook. f+ & acks., Ind. 3 22," aut, & 43. 1959; Moldenke, Résumé Suppl. 3: 19 & 20 (1962) and 6: + 1963; Moldenke, Dansk Bot. Arkiv 23: 87. 1963; Molden- ke, Résusé’ 3 Suppl. 1h: 3. 1966. Titastrations: Sims in Curtis, Bot. Mag. 7: pl. 2107 {in color]. 181 This vari yaa ae fers from the mer ar form of the species in having its corolla-lobes densely white-tomentose on the ou ee the leaf-blades oblong or near eee WEI: 6.518 5 cm. long, 3-10 cm. wide, acuminate or attemate at the apex, at- tenuate at the base, and * denticulate along the aaa except at the base, and the petioles 0.5—2.5 cm. long. The leaf-blades are membranous or subchartaceous, glabrous a- bove when mature, densely white-tomentose beneath, the secondaries s mM. Meng the trunk 1 cm. in diameter, be leaves "stinking, hot~ benea mauve, Seana: or red, and the fruit ert ¥ immature, violet or deep-purple to nn piper k or black when mature, small, inch in diameter, sweet e type of the variety was collected by Johannes Elias Teijs- mann (no. 1159 H.B.) = Sumatra and is deposited in the Herbarium Bogoriense at Buitenz I have examined the type and helen iso- s and find the mets oe labels inscribed m8", "32", an. Lam (1919) describes this taxon as follows: "var. sumatrana membranacea, vel subchartacea, lanceolate-ovata, basi attenuata, integra, apice acuminata, margine denticulata, adulta supra gle- 126 PHYTOLOGIA Vol. 1h, no. 2 bra, subtus dense albido-tomentosa, nervis utrinque 8--10; 6 1/2- 12 (—18 1/2) cM. longa, 3--l, 1/2 (—-10) cM. lata; petiolo 1—2 1/2 cM. longo; corolla pilis densis in vittis in lobis positis, tecta." He cites: MALAYA: Malacca: Griffith s.n. (Le--908.265- 1440). Penang: Delessert s.n. (Le--905.265-1]2;). INDONESIA: GREATER SUNDA ISLANDS: Banka: Collector undetermined s.n. (Ut-- 49888). Celebes: Elbert 2538. Java: Junghuhn s.n. (Tt—19887); Zollinger 35 & 157. Sumatra: Junghuhn s.n, (Le—-908 266-808) ; 1100, Le—908 265-1126, Le—908 .265-367, a pt Bi He coments that "Perhaps there is a more glabrous form, belonging to this variety aA oreo ite it seemed to us, however, not conspicuous enough, to base upon it a new forma. There are tran~- sition forms to var. y and 4 [longifolia & latifolia}. It is distinguished from C. longifolia, with w with which it is nearly conform in regard to the leaves by the always tomentose, never laxely floccose texture of the lower side of the leaves, and its hairy rolla" Bakhuizen van den Brink 9) describes the variety as "Cor- olla on the lobes densely white tomentose o' Aap leaves oblong or lanceolate-ovate, attenuate, at Pee ends, ns snall- denticulate; 10—15 cM, long, 3--10 cM. wai so peti ieiee 0.5--2 cM. Sumatra: Padang (TEYSM. 11595 3081) 5 Lampong (TEYSM 1381). Distribution: Sumatra! Malacca g" It will be noted that Lam (iil, pe includes C, albida Blume in the synonymy of this taxon, I place t his in the synonymy of C, longifolia Lam, Pei nea lists C. chinensis Hort. as a doubtful species. He apparently regards the _— as the typical C. candicans, since he cites for * cana the C. Ford s.n., A. Henry B-D2, Tsang 83 & 315 [as BSN), ‘and | Chun 1554, ci ited b by me below as var. ", Sumatrana. van den Brink (1921) regards plate 2107 in curtis ae Bot. tags and dates it "1820". Sims same plant oak ie cana of the Mantissa, Retzius affirms on the authority of a specimen received from the author himself. The specific name of tomentosa has been since applied to a dif- ferent species. Our plant differs from americana in having the stems and underside of the leaves much more S pecially in having the rac » the berries in the lat- together so as to look like m whence it has called the Bermudi "Native of Malabar, Cochinchina, Java, Sumatra, and the straits of Sunda, Requires to be kept in the stove. Introduced to the Kew Garden in 1790, by the Right Hon. Sir Joseph Banks, Bart. Ke B.3 but does not appear to have blossomed there at the "time of the publication of the Hortus Kewensis, in 1810. Our drawi taken plant communicated by ep Barr and Brookes in 1966 Moldenke, Monograph of Callicarpa 127 June 1818, from their very extensive collection at the Northamp- ton Nursery, Newington Green." His calling the inflorescences of this "racemes" is an amazing error, since they are typical S. Collectors have found this plant growing in sandy soil or loan, , ugh lime- on dry level land, open slopes, open hillsi and ro hills, along s, in forests, sandy river’ » and ground, at altitudes of sea-level to 1700 meters, flower. from March to June and Septembe: November, ti arch, June, July, November. Lau describes it as "fairly common" in sandy soil of thickets on steep slopes and as "fairly common scattered shrubs" on H sland; Lei t "scattered shrubs in village greens" but "fairly common" or "abun- dant", and Chun refers to it as "common" on the same . Hen- derson reports it "common amongst boulders" in Mal ind- aya, hipongse calls it "common in deciduous forests" in Thailand. Chun describes it ns an "herb", but this is surely an error in observa- tion. Henderson says "fruit white", but there are only flowers (no fruit!) on “the sheet bearing this label, so I question the of the statement — no other white-fruited examples lar names "khao tawk" and "tampa besi" are reco The flowers are described as "red" on Lau 1026 ae Tsang (Tak] 315, "pinkish" on Chun 1554, "pink" on Lau 139, Lei 155, and Liang 61557 & 66541, "purple" on Sindhipongse es mauve" on Mrs. D. J. Collins 230 | and M. R. Henderson 18202, and "pale- e-lilac" on Squires 188. Material has been misidentified and distributed in herbaria r the names C, cana L., C. candicans (Burm. f.) Hochr., Cc. giraldiana Hesse, C. longifolia Lam., and even Urticaceae! In all, 105 herbarium specimens, including the type of two of the names “involved, ~— 3 mounted photographs and illustrations have been Citations: vats West Bengal: Griffith s.n. [Bengal] (Bz— 17483) ; Kurz s.n. [Sibpore, 5/67] (Bz—-17482). CHINA: Kiangsi: as 10342 (N). CHINESE COASTAL ISLANDS: Hainan: W. Y. Chun O (Herb. Univ. Nanking 5646] (Ca—233793), 155 0 . Univ. ne 6751] (Ca--2h2623), sen. (Herb. Univ. “Wanking 5805] (Ca— 233793) 3 c. Ford s.n. (N); A. . Henry SMe {Hainan, 28 3.93] (N)5 SA: Lau 139 (B, Ca——525256, I, Mi, N, W--1629209), 1026 (N), 2801 (Bi, S), 3379 (Bi, S); Lei 155 (B, Ba, Bi, Ba--17h67, Ca— 612232, N), 731 (Ba, Bi, Bz—17)68, Ca--612572, N); Liang 61557 (B, N, W--1669691), 61931 (N, S, W—-16707h9), 64567 (N), 652208 (Go, %), So (N); Ts [Tak] 83 [Herb. Lingnan Univ. 15582] (ca—316 S771, N, W—2 ), 319 [Herb. Lingnan Univ. 1706h] (Ca 356897, N, Ls Pose rces Wang 33351 (N)- THAILAND: Mrs. D. Je 128 PT T.0 L.06-5-4 Vol. 1h, no. 2 Collins 1252 Pare 70 NO) 2340 (W--1701677); K. aces 9168 (2); Sindhipongse 76 [Herb. Roy. Forest Dept. 6020] (W--206]525). IN- DOCHINA: Annam: Clemens & Clemens 10 (Ca—3)0720, st gia ‘i Mi, N, Ut-—Sla, W—1)27862); C. B. Robinson 1273 (N, W—713355)3 Squires 188, = Lat re La), 791 (Bz—-1770, N, S). Coc 5225 (B, Bu—72839, Ca-—36108, Ca—Sh657, $34, seis res 188, in part (Bz—27h8h, pment Thorel T19 (3B, Sy Sate 56, W—1758020) . Laos: F. K. Ward 8911 (N). BOOK REVIEW Alma L. Moldenke "Fundamental of Phytomorphology" by A. D. J. Meeuse, xi & 231 PPpe, illus. Ronald Press Company, New York, New York 10010. 1966. $10.00 Herein the author champions the New Morphology promulgated by H. H. Thomas in 1931 and documented primarily by Hagerup, ee oic in ospermous-Bennettiales form. The loser, slain by sharp deductive logic, references to morphogenesis and reinter- pretations of fossil material, is the Old Morphology with its roach. The single main force evoking adaptation through positive selective tendencies was the advent of entomophily and it resul- ted in versal my 0: nd of angiospe the gynoecia into closed structures. The female cycadopsid gymnosperm prototypes had developed ovules that were fertilized by siphonogamy leading to an early stage of double fertilization, bitegmic with a third layer of cupular origin, borne in groups on a ad, gynoc The encasing of the ovules 8S required the formation of stigmatic areas bide ovular coats and/or carpel walls and eventual stylar The poe cycadopsid prototype became similarly arranged in compound androclads also subtended by bracts. These androclads and the gynoclads separately or together,with some sterile leafy organs, formed strobiloid anthocorms There were spear am re~ ductions and differentiations producing flowers, The very last paragraph in the book sol a plea to paleobotanists to hunt for There is much meat “a for discussion and many ideas for ting speculation. PHYTOLOGIA Designed to expedite botanical publication Vol. 14 December, 1966 No.3 “ | CONTENTS GUEVARA, A. E., & KING, R. M., Vernacular names for some species of Colombian Compositae. . . . J ges gee eee i MOLDENKE, A. L.. Book reutews. 2-2. 0 a ep ~ DWYER, J. D., & HAYDEN, Sister M. V., Three new species f of Neea (Nyctaginaceae) from Panama ... . a ee 137 MOLDENKE, H. N., Additional materials toward a monograph of the genus Callicarpa. V. ..--+-+-++-+ + + 140 e Published by Harold N. Moldeake and Alma L. Moldenke — 303 Parkside ons ae Jer VERNACULAR NAMES FOR SOME SPECIES OF COLOMBIAN COMPOSITAE A.E. Guevara and R.M. King During the summer of 1965 » While collecting material for jroo study of Com ompositae, it became very apparent that in some cases, th common or colloquial name was very useful in identifying the plants. name The obtained primarily from annotations on labels on herbarium specimens in the United States pone Herbarium and the herbarium of the Instituto de Ciencias Naturales » Universidad Nacional de Colombia, but are supplemented by the co own inquiries in the field. The gts erg aad of the species have not necessarily been verified by specialists SCIENTIFIC NAME VERNACULAR NAME fcanthospermum australe (Loefl.) Kuntze cancer de loma 4+ hispidum Dc. pec mortal, espuela de caballo, ericito playero, cac js eL trejo. Achyrocline satureoides (Lam.) DC. Juan Blanco. Acrocliniun zoseum Hook. Inmortales dobles. Agsratum conyzoides L. Chiba, Manrubio, yerba de chino, Marrubio blanco, yerba de chivo, Mastranto. Ambrosia cumanensis H.B.K. Altamisa, Artemisa. Anthemis nobilis L. Manzanilla de Castilla. Le Artemisia sodiroi Hieron. Alucema. Aspilia tenella (H.B.K.) Blake Papunga falsa, Guasquilla. aster laevis L. A. marginatus RSI: Hortensis Cerrajita cimarrona, 130 TayYToLogya A. sinensis L. Baccharis cinnamonifolia H.B.K. decussata (Klatt) Hier. censstelioldes — scen poe moe Sp - x. [od [od Id Ib [ee] latifolia (R. & P.) Pers. lehmannii Klatt macrantha H.B.K. prunifolia H.B.K. tridentata Vahl Bidens cynapiifolia H.B.K. B. pilosa L. [63 bd Ito Ibo B. pilosa var. radiata Sch.-Bip. Calea glomerata Klatt C. penn i Blake C. eee: H.B.K. c yuruparina Cua re Calendula officinalis L. eel Centaurea cyanus L, Chaptalia nutans L. Chrysanthemum frutescens L. C. leucanthemm L. Cc. pathenium (L.) Bernh. Chuquiraga jussieui G.F. Gmel. Vol. 1h, 0. 3 Montenegro. Chilea, chilca tefiidora Armanga. pte Chica " pegajosa, pe “rusia, Chilca, Chil Chtice-rusia. Cc * Ciro de sierritas, Chilco. Ciro ceniciento. Cadillo menudo. pung see Cadillo de perr eee Papunga de masta Carrasposa, Chicharrén, Calendula, Maravillose- Albarina. arita. itén. ounandbtineg matricaria, Manzanilla roman Chiquiragua. 1966 Guevara & King, Vernacular names 131 Clibadium asperum IC. Chiu-Chiu, Sicuri Catalina, Chisi-chisi, Chilca amarilla e+ Sur surtnanenss 1: Pintadilla. c. omccranagy AE ) Baill Barbasco. Conyza bonariensis (L.) Cronq. Venadillo, Yerba de aball O. C. bonariensis var. leiotheca (BL.) Cuatr. Chilquilla, hoji- sauce. Cosmos bipinnatus Cav. Guatemala, Correo. Dahlia Lehoanntt Hieron. Dalia de huertas. D. yar: yariabilis Delia amarilla. piplostephium bacchar ideum Blake Romero de Monte. D. cinerascens var. centrale Cuatr. Guasguin. D. flortbundun 8 ubep. putas “hetaaiiienss Cuatr. Chilca rusia. D. revolutum Romero. D. ro peace Benth. Romero. Elephantopus mollis H.B.K. Yerba de caballo, Suelda-con-suelda. Erechtites valerianifolia (Wolf) DC. Bolador, Tachushiash, Venadillo. sopeletia congestiflora Cuatr. Frailejon. E. glossophylla Mattfeld Tabaco de la sierra. E. glandulosa Cuatr. Frailejon. E. grandiflora H. & B. Frailején. E. hartwegiana Frailején, lana de ove ja. E. hartwegiana Cuatr. var. brachyphylla Cuatr. Frailején. E. incana Cuatr. Frailej6én. E. iimenez-quesadae Cuatr. Frailején negro. E. lopezii Cuatr. Frailején E. murilloi Cuatr. lején. E. Petiolata Cuatr. var. escobensis Cuatr. Frailején. E. phaneractis (Blake) A.C. Smit Frailején. E. pleiochasia Cuatr. Frailején. E. schultesiana Cuatr. Frailején fish. torium acuminatum H.B.K. Patinegra, trebol aromati a 132 PHYTOLOGIA amygdalinum Lam. angustifolium (H.B.K.) Spreng. brs bullatum Klat humile Sars ) Hier. inulaefoliun H BK. rE E. laevigatum Lam E. odoratum L. 3 prsnoces slum Lose. E. stoechadifolium L.f. Galinsoga parviflora Cav. Gazania speciosa Less. Gnaphalium elegans H.B.K. G. spicatum Lan. Gynoxys parvifolia Cuatr. Helichrysum bracteatum Andr. Helipterum manglesii Muller ex Benth. Hypochoeris radicata L. H. sonchoides H.B.K. chthyothere scanden 8 Blake = terminales ference -) Blake Jungia soororses | Cuatr. ferruginea L J. f a: moschata sets i Pisbum bouplandit Cass. me eee ) Less. Vol. 14, no. 3 Santuaria. Chilca blanco-ceniza, Amargoso. Carrasposo. radi Salvia anarga , Salvia some ee “Chilquita, Yerba de chivo. Chilquilla. See RSS carrasposa. Vira-v guasca, chipaca. Gazania remarcada. Vira-vira, Yerba gallinoza. Rama blanca. Oo, iInmor- tales dobles, Siempre- viva, Inmortales de racteas. Siempreviva, Rodantes- Serraja de p&ramo, Serraja. Cuicenia cimarroné. Barbasco. Bayra, Jarilla blancé- Manito de oso. Mano de Oso. Mano de Leén. Yerba de Santa Marfa. Santamarfa de abejas- Santa Mar 1966 Guevara & King, Vernacular names 133 hs Bap: om ane a L. vulcanicum Klatt Lycoseris crocata (Bertol.) Blake L. latifolia (Don) Benth. Matricaria chamomilla L. Mikania cordifolia (L.) Willd. M. guaco H. & B. M. micrantha H.B. M + Pachydictya B. L. Robinson M. ruiziana Poepp. Hontanos lehmanii parkas Be Blake “ M. ovalifolia Sete 00 eg . woe clematis L.f. M. glabrata Cuatr. Neurolaens lobata (L.) Britt. Qliganthes discolor (H.B.K.) Sch. Bip. poseris cuoveroldes (H.B.K.) Rob. Pectis elongata H.B.K. Fluchea purpurascens (Swz.) DC. Pollslesta colombiana Aristeguieta Folymmia pyramidalis Tr. Bejuco amarillo. Santamarfa. Patuda. Amapola, Mapola, Mapola de monte. Ma Manzanilla. Guaco. Guaco Bejuco Guaco. Falso Guaco. Guaco. Guaco lanoso. Arboloco. on. Arboloco. Clavellino e Clematide, Clavellina de bejuco, Be juco wnat Clavellina. Contra-gavilanea, Salvia gavilana. Cenizo, Susaca Come jenero, Sauce, Hoja de Santamarfa. Santa Marfa. Comino silvestre. Tabaco cimarron. Cenizo. Arboloco bogotano, Arboloco hueco. 13h Poe Ys. O:L 0-6 Ta P. riparia H.B.K. Porophyllum ellipticum (L.) Cass. P. macrocephalum DC. P. ruderale (Jacq.) Cass. Pseudelephantopus spicatus (Juss.) Rohr. Pseudogynoxys bogotensis (Spreng.) Cuatr. Pterocaulon alopecuroides (Lam.) DC. Senecio abietinus Wedd. Ss. quericanus ~~ 22) ic. formosus H.B.K Ica} @uicanensis Cuatr. E oe H ophyllus H.B.K. sisrochante lary Cuatr oo Boa it. rufescens IC. vaccinioides (H.B.K.) Sch. Bip. Siegesbeckia cordifolia H.B.K. Sonchus oleraceus L. ZEREPEE Ita |ta . Spilanthes americana (Mutis) Hieron. Vol. 14, no. 3 Pauche, cumana, Jiquimilla. Yerba de gallinazo. Yerba de chulo purranga, chipaca. Venadillo, Ruda de p oped Brora Galle Sueldo-con-suelda, Yerba de caballo. Amapola del camino. Venadillo grande. Panque Romero, Romero chiquito. Duarte Arnica, Arnica de Bogoté, Arnica de pa. Romero, Chilquilla nudosa. Frailején blanco. Guasguin. Trompetilla. Serraja, Serraja lechosa, Cerraja grande, Diente de leén lechos0- Santa Marfa, Chisaca de ecafetal, Yuyo quemado, Chisac Chisac salaeiane ' acest : 1966 Guevara & King, Vernacular names 135 Steiractinia aspera Cuatr. - oyedaeoides Blake Stevia lucida Lag. Sylibum marianum Gaertner Tagetes apetala Posada T. erecta L. T. patula L. T. pusilla H.B.K. T. terniflora H.B.K. 7 - zipaquirensis H. & B. Taraxacum officinale Weber Tessaria integrifolia R. & P. Tridax procumbens L. Trixis radialis (L.) Kuntze Verbesina crassiramea Blake Pterophora Blake - turbacensis H.B.K. I M. Victoria Hayden 29 (MO); 67 (MO); A. Robyns (MO). DARIEN: Piocgans’ to Yaviza, Duke 5163 (MO). 3. NEEA LAETEVIRENS Standley, Field Mus. Publ. Bot. : 20h. 1929+ eet tele mot Colorado Island, Sr. - ¥iatorss he 1 (MO); 12 Colon, Tyson 2317 (MO). er 5158 ); toatl, between Paya & FIs de ies letras, ern, Chambers, r & Ebinger 196 (MO); between Paya & payita, Stern e ter ee )3; between Paya & Boca de Paya, stern et al 436 (MO). . NEEA ELEGANS Dwyer & Hayden, sp. FRUTEX, ramulis primo Gel ateeid “altime compressis minute rufo-pubescentibus . FOLIA See pe: petiolis 5-15 cm longis, rufo-pubescentibus; lamina anguste elliptica, 7-17 cm longa, 3.5-5.0 cm lata, apice scoumivaccspatns a, acumine ad 1.2 cm ngo, basi acuta, tenui-papyracea, puberula, ciliolis minut rt c areolis intervenialibus patulis. INFLORESCENTIAE terminales, cymoso=paniculatae, ad 7 em longae, ad 5,5 cm latae, pedunculo ad 3 cm longo, ad 0.2 em lato, rufo-pubescente, ramis paucis proximis vel distantibus, supra medium cymulos patulos paucifloros ferentibus, floribus terminalibus solitariis. seantidees floribus d bracteolis ge su FLORES (hic o") perianthio sessili elliptico-urceolato, ca 5.5 ml PANAMA: Darien: rd fr El Real to Pinogana, Duke Sub (MO, Holotype). The new species is readily recognized by its salicoid leaf blades and short, relatively few-flowered inflorescence. 5. NEEA DARIENENSIS Dwyer & Hayden, sp. nov ARBORES ad 10 m altse, ramlis teretibus laevibus glabris- FOLIA alternata, petiolis 1-2” em longis inconspicue alatis; elliptica (aut foliis minoribus elliptico-rotundis ve obovato-rotundis, apice rsa 7-5-20 cm longa, 5-9 cm lata, ap acuminata, acumine ad 2 cm longo, 0.8 cm lato, crasso=- , omnino glabra, venis lateralibus ca 10, arcuatis evan~- escentibus. INFLORESCENTIAE (nic in fructu) terminales pyramidato- paniculatae in vivo rubrae, > Ppedunculo subplano-com~ 1966 Dwyer & Hayden, New species of Neea 139 presso lignoso, ad ).5 cm longo, ca 0.3 cm lato, ramis inferior- Se ad 4.5 cm longis, superioribus ca 5 bene distantibus, reflex- s 8 ) a cicatrice calycis nullo, mesocarpio carnoso, endocarpio fibroso, seminibus solitariis, exalbuminosis, testa papyracea, toca tual ibus carnosis, ca ) mm longis, ca 3 mm crassis, eis c pact pera an oo aoe tegentibus : vic Paya, Rio Paya, Stern, Chambers, Dwyer & Ebinger 129 ( (M0, Eo} otype). The thi papyraceous leaf blades ft perha better described as Salto subcoriaceous, and the lignose ag te inflorescence are strikingly different from ey other Central a rare ws of the genus except Neea amplexicaulis described new below. 5 davieueseies® on the other hand, has conspicuously poem and site rnate Leave 6. NEEA ATES ICAL ae canee & & Hayeens Spe nov neagvoae 0 EIRP FOLIA amp rotundo-truncata vix cordata, coriacea glabra in sicco nigro- b a, costa supra promimula, infra subprominente, venis lateral _ 15-20 (folia maiora), 1-2-5 om distantibus, late divergentiby ne stricte et integre 2/3 ad marginem extendentibus tunc late bifureatis, are olis intervenialibus patulo-reticulatis. INFLORES- CENTIAE terminales, ad 10 cm longis, in vivo rubrae, pecan ae ad 6 cm longo, 1.8 cm lato, rufo-puberulo, r primariis 2, cendentibus, ramulis secundariis alternatis vel oppositis alvere- entibus, 0.5-2 em lo is, c parte superiore r Persistentibus, floribus digitate dispositis. FLORES (hic o) ) Sessiles, bracteolis conspicue late triangularibus, ca 0.6 mm longis, obtusis sracknol4eo-gutaegemenemndine liberis sed infra Perianthum ternate dispositis, periantho oblongo, ca 2.3 mm longo, ca lm lato, obtuso, carnoso, praesertim basi arachnoideo-pub- escente; stamina ca 6, antheris ca 1 mm longis. FRUCTUS inflorescentiarum cranes Tigmoets . S 2022 mm crassis, cicatrici- bus anth , anthocarpis ovato- ocarporum conspicue annular elispticts, ca lcm Pyemcis. ca OE | cm Jatis, fibroso-lignosis, s Holotype); Sanka Yo; Laks 1,02 ()F Was apenas ato Buena Vista, Rio Chncunaque above cont luence wi » Stern, Chambers, Dwyer & Eb Bbinge z @ species i readily recognized by its opposite and amplexican] sons ADDITIONAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS CALLICARPA. V Harold N. Moldenke CARPA CANDICANS var. SUMATRANA (Miq.) Moldenke Additional citations: INDOCHINA: Tonkin: Bois 255 (Ca—5)658, W—1758019). State undetermined: lest 3327 [Thua-thien] (Ca—-38107). MALAYA: Perak: Burkill 12418 (Ca--346270). MALAYAN ISLANDS: Aor [Johore]: M. R. Henderson 18202 (Bz--17465, Ca 318663). WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Luzon: Lete 193 (Ca—365687). INDONESIA: GREATER SUNDA ISLANDS: Java: Backer r 9490 (Bz--17352); Sundalandschap. Urwaudenflora 3-6000 (Ut—l9887). Kangean: Dommers 10 (Bz--17472). Sumatra: Backer 3213 [113] (Bz--17474); Collector undetermined 534 (Ba—-1 7473) 3 Lorzing 11140 11140 esse? as 1159 | H.B. (Bz--1775——is0- type, Bz—17h77—- hoto of type, Ut—53389—isotype, Z— photo of type), 3008 (Bea 7hBl), 308; #.B. H x (Bz—-17,80, Ut— 53388), 4381 H.B. (Bz--17478, Bz--17L79, 17479, Ut--53390). CULTIVATED: Java: Herb. Hort. Bot. Bogor. X1.G.27 (Be—25722, Bz--26518, Bz, N), s.n. (Bz--17471). MOUNTED ILLUSTRATIONS: Sims in Curtis, Bot. Mag. 7: pl. 2107. 1819 (N). ol. Bibliography: Chang, Acta P Phshokae in, 1: 305. 1951; G. “Tay- ler, Ind. Kew. Suppl. 13: 21. 1966; seis, Résumé Suppl. 1s: Nothing is known to me of this species since the work in which Res Was originally described has not yet been available to me. The plant is said to inhabit the provinces of Kiangsi, Ki- angsu, Kwangtung, and Kwangsi, China. Bae Bonn oe Malay. Arch. 60.1919. Callicarpa caudata var. typica te J. Lam, Verbenac,. Malay. Arch. 60. 1919. Callicarpa caudatifolia Max. ex Moldenke, Résumé Suppl. 3: 30, in syn. 1962 [not G. ee a folia Koids., 1925, nor Merr., 1962]. Callicarpa mo ex Moldenke, Résumé Suppl. 1h: 7, in syn. 1966 [not “ea ae Matsumura, 1922, nor Req., 1839, nor Shirasawa, 1949, nor Sieb. & Zucc., 18h, nor Willd., 18)0} : CALLICARPA CAUDATA Maxim., Bull. Acad. St. pe tt se 76. 1886. we =o 5 Bibliography: M Maxim., Bull. A : - Sci. St. Pétersb. 31: le 1886; Maxim., M4. Biol. 12: 2 50650. 1886; Durand & Jacks, . rt & Merritt, Philip’ Journ. Sei. Bot. 5: 380—381.°19103 E. D. Merr., Philip. Journ. Sei. Bot. 10; Tl. 1915; H. J. Lam, Verbenac 140 1966 Moldenke, Monograph of Callicarpa 141 Malay. Arch. eli eee 65, & [361]. 1919; Bakh., Bull. Jard. Bot. Buitenz. “Se 4 3: 23. 19213; E. D. Merr., Enum. Philip. Flow, Pl. 3: 383. "1923; Moldenke, Alph. List Common Vern. Names » 17, 20, & 28. 1939; Moldenke, Prelim. Alph. List Invalid Geogr Names 10. 19405 Moldenke, » Distrib. Verbenac., [ed. ] Mec Speen hy Mee List Invalid NW ae 1942; Moldenke, P' ogia 945; Moldenke, Alph t Cit. hytol 1: 22h (1946), 2: Ok & M3 (aol), 3: 765 (19195, and *y 1289. re ee » Known Geogr. Distrib. ae a { oo ’ . 1949; Moldenke, Phytologia 3: 6h (1951) “and hz 122— : i982, “Woldenke,, Résumé 168, 182, 19h, Sor 198, 200, 242, & li, 195951 Moldenke, Résumé Suppl. 3: 21 & 30 (19625, 12: 8 (1965), and 1h: h, ‘. & 7. 1968, Moldenke, Phytolo gia 3: 431, re £7499 (1966 966) and 1h: 121. 1966. Slender large bush, shrub, “or small tree, 2--10 m. tall, the young whitish-yellow floccose or tomentose; trunk 2—h cm. in diameter; stems floccose-tomentose; leaves decussate-opposite; petioles short, 3-7 m. long, hago s fos floccose—tomentose; leaf~blades membranous narrow. eolate or linear-lanceolate, narrowed at bo 2-345 cm. cae 1.2--6 cm. wide, whitish-yellow floccose or to~ mentose; buds pink; calyx 0 -75—i m. long, hispid with long simple hairs, glandular—dotted, with many glanduliferous hairs Some large scales, the rim eaebitaly Serre corolla ape mm. long, about twice as long as the calyx, r lavender-pink to een, Ee es purple, or violet, aaneelin or with some and some glands, its tube broadly cylindric, the lobes y 3 the lesg reer: of the tube; stamens slightly exserted or twice as The type of this species was collected by Hugh Cuming (no. 1095) in the Philippine Islands and is po ne = the a ardin Botani Principal at L (1886) says of it: "Folie Calli pilosissimae a rat 5 a, lamina 11: 2 cm, Pedunculus 15 m., cyma diam. 20 m. remeen ae 4 ve a. 6 longa." Hayek (1906), in speaking of C. elegans Hayek, ey eA caudatae tie et serratura maiore nec non indumento ramorum diverso." Lam (1919) divides the species into four varieties, but his var. 1h2 PHYTOLOGIA Vol. 1h, mo. 3 lata, petiolo 0.5 cM. longo; utrinque dense aimee stellato-tomentosa; cymi 1.2 cM. diam., peduncul 0.5 a Jones calyx dense pilosus pilis simplicibus, *pilis planduli fer — singulis. Distribution: Philippines (Max.). "Var, — magna H. J. Lam, nov. var. — folia cum eis var. 4 conformia, sed dinensiociins differentia, 9 1/2 — 16 cil. —— Merritt, & Soshokive 18127, Elmer 8616, and Herb. philip. Forests Forest. Bur. 1643 & 16h. 1 ndanao: Elmer 11333. He notes that what he calls C. reevesii Wall. (now know as as C. mudiflora Hook. & Arn.) is related to C. caudata and to C. pilosissima. He also avers that C. pedunculata R. B R. Br. is related to it, but never has its leaves with the acute or cuneate base seen in C. caudata The original publication by Maximowicz is dated a 1887" by Du- rand & Jackson (1901) and by Lam (1919) « It cate wi be noted here that the C. caudatifolia of Koidzumi, rred to in the synonymy above, is actually a synonym of C. sae vare eet sana var. glabrescens Moldenkes; the C. mollis of Matsamura is » okinawensis Nakai, that of Shirasawa is x0. 3 xC. shirasawana Mak., et of Siebold & Zuccarini is a valid species, and that of "Req." and of Willdenow is C. acuminata H.B.K. Philippine Islands, in March, 1915, and is deposited in the her- barium of the University of California at Berkeley. zen van den Brink (1921) places C. caudata in the sym onymy of what he calls C. cuspidata Roxb., along with such di- verse taxa as C. rubella Lindl., C. acutidens Schau., C. sessili-~ 1966 Moldenke, Monograph of Callicarpa 143 ye and in rocky and mossy soil on ridges, on open stream- banks, pine and damp forested slopes, and small coral islands, ~ at stream margins, from = to 2000 m. altitude, flowering fruiting in March, April, June, and August to January. he Fines are described as fate on Brass 5520 & 31788 and on Herb. Philip. Bee Sei. 43310, "lavender-pink" on on Clemens & Clemens 326k, "lavender" o} m Brass 31563, "purple" on Womersl 1323 “ & “Wpale— purple" on Hoogiand & & Pullen 5328, cag "yiolet® on Herb. Philip. Bur, Sci. 4,895 and on Verst teegh B.W.3056 Vernacular n names recorded for the we. are “anayop", terrae Hy "arayop", "diapu", "gital", "harayhai", "kabatiti", Nicumukwireh" , "lumis via", "mama", "minari", "nimnam", "nimname", "sakoita", "saykohadza", and "suba". Clemens hb” has a note “with Urt. & Astile rusts". Material has been misidentified and distributed in herbaria under the names C. cana L., C. data Roxb., C. lancifolia Merr., C. longifolia Lam., C. macrophylla Vahl, C. merrillii Mol- denke, C. pedunculata R. Br., C. pentandra var. paloensis (Elnm.) Bakh., C. rubella Lindl., and C. stenophylla Merr. On the other hand, the De Vore Vore & Hoover hi, “distributed as C. caudata, is C. merrillii Moldenke . In ~ Inall, 9) herbarium specimens, including type — of most of the names involved, have been ned as Citations: WESTERN PACIFIC ISLANDS: PHILIPPI TSLANDS: Bohol: M. Ranos son. (Herb. Philip. Bur. Sci. 3310] (Ca—-2h2439)- Lu- tcosa 7 7 THerb. Philip. Forest. Bur. 30020] (Ca—2917h7) 5 Curran, Merritt, & zschokke s.n, (Herb. Philip. Forest. Bur. 18127] (Ba— 17507), s.n. (Herb. Philip. Forest. Bur. 18163] (Bi); Elmer 578h (B2—-17525, sh 5870 2, 8646 (B2—17505, N, Vt); F.C. Gates S455 (Ka—61,623); Loher s.n. [Caraballo Mountain, Mar. 1915] (ca— 229198) 5 McClure rise (Ca——307573); E. D. Merrill 1727 (Bz— 17508), 4613 (N) @), 238) 11689 (Bz—17509); M. Ramos 8-Ne THerb. Philip. Bur. Sei. 772)] (N); Ramos & Edafio sen. [Herb. Philip. Bur. Sci. 37481] (Bz—17510), sen. [Herb. Ph ave Bur. Sei. 38035) (Bz— ), Sen. [Herb. Philip. Bur 40505] (Bz—17508), fee Pp. Bur. Sei. 44895] cea aderene: N), sen. (Herb Philip. Bur. Sci. 4863h) (acs 2208, N); R. S. Williams 1060 (N, N), 1158 (N, N). : M. K, Clemens s.n. [Herb. Herb. Philip. Bur. Sei. 15676] (Ca—-268581, pote Elmer 11333 (Bi, Be—17517, N, Vt); mene & Butohinson sn. as (ue. Fe lip. Forest. Bur. 1755] (B2—1 751 3); Ramos & Edaflo s.n. (Herb. Philip. Bur. Sci. seo} ans --17512), 8 2.5, Tier, PRtip ns Sci. 49295] (B, Bs—17520, Ca—32h45h, N)3 Ra Pascasio s.n. (Herb. Philip. Bur. Sci. 34775] (HN); R. 5 "piles Wiilizes 2507 ti Santa Cruz: S- wisen 1062 (Cp). Ticao: W. ¥. W. Clark s.n. [Herb. Philip. Forest. bh PHYTOLOGIA Vol. 1h, no. 3 Bur. 2534) (Bz--17523, N). Island undetermined: Herb. Philip. Bur. a Sen. (Gg—-3199h); Née 2h (Q). INDONESIA: GREATER SUNDA : Celebes: T, Kaudern 12 (N); W. Kaudern 313 (N); Koorders 19485b (B2—-17535, B Bz—17536), 1 19k98b [448] (Bz— oak i Bz——17538). LESSER SUNDA ISLANDS: Timor: pachienses Bole 8942 (Bz—-17497); Walsh 467 (Ba—17h96). MOLUCCA ISLANDS: Amboina: C. B. Robinson 299 (Bz—1798, NW). Halmahera: Anang 569 (Bz—-7295h), 638 (B2—72699)- Morotai: Anang 179a (Bz—72982), 235 (Bz--72981) ; Main & Aden 947 (Bz—~72705). Sanana: Bloembergen 1,31 (Bs—17),99) . Ternate: Beg- vin 1214 (Bz—17502). MELANESIA: NEW GUINEA: Dutch New Guinea: D. Bergman 3 (S); Docters van Leeuwen 10106 (Bz—72659); Kane- hira & Hatusima 13539 (Be—1750h); Mayr 10h (Bz—72635); Versteegh B.W.3056 (Ng—20223). Northeastern New Guinea: Brass 31087 (W— 2391917), 31563 (N, W—2392210), 31788 (W-—-23923h1); M. K. |. K. Clemens Tuk (B), 7852b (B), 11028 bis (Mi); Clemens & Clemens 326) ; (Br)5 Draper s.n. (Ng—-16957); Hoogland & Pullen allen 5326 (Bi, Ng—16911, Ne, W—231L880); N. G. F. 3065 (Ne—20222); Womersley 1323 (Ng—6L91)- Papua: Brass 5520 (N); C. B. Carr 13163 (N); Giulianetti s.n. (Mb). BISMARK ARCHIPELAGO: Mussau: Kgie & Olsen 1363 (Gp, Z). New Britain: Waterhouse 373 [Herb. Yale School Forest. 29198] (N)- ss ye CAULIFLORA Merr., Philip. Journ. Sci. Bot. 7: 338—339- Synonymy: Callic. carpa pentandra f, hexandra subf. cauliflora (Merr.) Bakh. ex sere Résumé hg ieee in syn. 1959. Bibliography: E. D. Merr., Philip. Journ. Sci. Bot. 7: 330— 339. 1912; H. J. bas Verbenac. Malay. Arch. 50, 83, & [361]. 1919 93 Prain, Ind. Kew. Suppl. 5: 43. 1921; Bakh., Bull. Jard. Bot te. Buitenz., ser. 34 3: d3« 19213 E. De. Merr., Journ. Malay. Asiat. Soc >» Emm. mat Flow. Pl. 3: 383. 1923; Moldenke, Known G Distrib. errenly 2& 1942) and ed tetragonal and glandular or rather densely covered with entire, gradually narrowed or attenuate to the base, glabrous a- bove, somewhat paler, » and Pilose on the midrib es, and veinlet reticulation be- neath, denser on the midrib, the hairs often plumose; sec 5 1h pairs, prominent th, anastomosing; veinlet reticu- lation prominent, rather lax; ence borne on P infloresc fasciculate, the cymes sessile, cauliflorous, hemispheric, crowd- ed, rather dense, 2—3 cm, wide, slightly hirsute or brownish- 1966 Moldenke, Monograph of Callicarpa 145 floccose, glandular; pedicels 3—5 mm. long; bractlets linear- FO Op oa 1-~2 mm. long; calyx oblong-ovoid, about 3 mm. long, 1.8 mm, wide , its rim equally )-toothed, the teeth narrowly ovate, 0.5 mm. long, acute at the apex; corolla dark-red, about 7 mm. long, cylindric, slightly glandular and pubescent on the outside, the limb - lobed; anthers oblong, 3 mm. long, glandular on the back; fruit globose » the lower half enc pete by the fruiting- pyrenes. The type of this species was collec ted by Harry Nichols Whit- ford and Wallace Irving Hutchinson [ Philippine Forest Bureau 9321) in canyons in forests, at an altitude of 20 meters, at Por- to Banga, in the district of Zamboanga, Mindanao, Philippine Is- lands, on January ya ae 8 and was deposited in the herbarium of s now destroyed ac fascicled inflorescence, in this character —— from all the other species of the genus known to me, Allied to C. ramiflora Merr., but with a quite different indumentum." Lan "also affirms that its af affinity is with C. ramiflora. Bakhuizen van Brink (1921) reduces it to synonymy under what he calls C. pentandra var. typica f. hexandra (Teijsm. & Binn.) Bakh., which I regard as the very different Geunsia hexandra (Teijem. & Binn.) Koord. In all, 3 herbarium specimens have been me. Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Leyte: - Ramos s.n. [Herb. Philip. Bur. Sci. 150] (Bz—1822h, Bz— lies Z). CALLICARPA CHENAULTI Fairchild, Expl. Plants 51, nom. mud. 1930. Bibliography: D. Fairchild, Expl. Plants 50-51. 1930; Molden- ke, Known Geogr. Distrib. Verbenac. ee ed. 1, 71 & 86. 19h25 es N. 19, Moldenke, Ere Distrib. Verbenac., ed. 2, 156 &177. lis, Moldenke, dad 213 & 43. 1959. S known to me about this plant except what is stated a Feizehiia (1930). His reference, in full, is as follows: e hardly credible that in this climate, in so short a time, one could so 16 PRITOLOeIA Vol. 14, no. 3 transform a place and almost entirely with comparatively newly introduced things, but when one has been a nurseryman and dealt th a great variety of plants for fifty years one knows just what to plant. I venture to think that there were more rare spe- cies of plants in these few acres than are to be found in many a so-called 'Botanic Garden', At the risk of being pedantic and for the purpose of proving my point, let me enumerate those I saw in the brief period of my stay there. I only noted those which attracted my fancy. Rosa sericea petrolutea, Himalayan rose. orange-throated flowers. Rhus cotinus atropurpureus (Rehder's Cotinus cogeyeria, Scop.), a charming form of the smoke tree. Ri- like fuchsias which, in its simpler form, is wild in California. Rubus deliciosa, a handsome, large, white-flowered raspberry from Colorado. P us capensis, a handsome Scrophulariaceae from the Cape, known as the Cape Figwort. Ele. reflexa, 4 variety, according to Rehder, of E. ens from Japan. Calli- carpa chenaulti, a species which I find described nowhere in ny books ." apex, floccose- ntose on the younger parts, glabrescent in age, with scatter er wood; nod Spicuously annulate; principal internodes },.5-—-7 cm. long; leaves decussate-o and cm dens branchlets; leaf-blades subcoriaceous, grayish-green on both sur- faces, broadly elliptic, 14—-19.5 cm. long, 8—10 cm. wide, short- pex. Pp cymose; cymes 2 per node, about as long as the sub tioles, about twice bi- , ong tending pe ’ ssinted ranc floccose-tomentose with grayish hairs like the peduncles; bracts 1966 Moldenke, Monograph of Callicarpa 147 very conspicuous, a pair beneath each furcation of the cyme and numerous among the flowers, l--15 mm. long, linear-subulate, densely floccose-tomentos se; calyx campanulate, 2——-2.5 mm. long, very ue Socealy grayish-tomentose, its rim l-toothed; corolla British North Borneo, on July 22, 1933, and is deposited in the Herbarium Bogoriense at Buitenzorg, Java. The taxon is known to me only from the original collection, of which I meee examined | specimens, including the type. Spat INDONESIA: GREATER SUNDA ISLANDS: British North Borneo: Clemens & Clemens 3036 (Bz--18332—isotype, Be--18333— isotype, Bz—-1333—type, Z—isotype) . Baars it 5 COLLINA Diels, Notizbl. Bot. Gart. Berlin 9: 1030. : Callicarpa brevipes f.yingtakensis Ptei, Mem. Sci. Soc. China 1 (3): [Verbenac. China] 7. 1932. Bibliography: Diels, Notizbl. Bot. Gart. Berlin 93 ei eh 47—h9, pl. 5. 1932; A. W. Hill, Ind. Kew. Suppl. in Ey aphoel dell, Ind. Lond. Suppl. 1: 160. 191; Moldenke, Known Geo Distrib. Verbenac., ed. 1, 56 & 86 (1942) and ed. 2, 130% te. 1949; Moldenke, Alph. List Cit. h: ae Le: Moldenke, 168 & 43. 1959 3 Moldenke, Résumé Sup go 1966; Moldenke, ee 1h: 102. 1 tions: P'ei, Men. Sci. Soc. China 1 (3): [Verbenac. China) Pl. 5. 1932. st Small wo woody shrub, 11.6 m, tall, with a spread of 3 ag fulvo- tellou finally glabrescent; branchlets pubes- subs petiolate: tioles, if ent, not surpassing 7 mm. in length; leaf-blades' h P chatte lanceola ceclate, 8.525 cm. long, 3. *wide, the larger leaves al- ways at the w of the branchlets, the uppe ae ek at the apex, the lower ones obtuse or roun~ pe all crenately serrate, callose~serrulate, or entire The type of this species was collected by Hsen-Hsu Hu (ne: 2250) on shady hillsides at Tzuchi-hsien, at 600 meters altitude, 148 PHYTOLOGIA Vol. 1h, no. 3 ium of the Botanisch her Garten in Berlin, but is now Diels (1926) says: "A C. rubella landl. *affinis ramis robustiori- bus, foliis majoribus fere integris, basi ipsa vix cordatis, in- florescentia densius conferta, corolla glabra, filamentis brevi- oribus distinguitur". P'ei (1932) cites an isotype in the Arnold Arboretum herbarium and also Hu 1125 from Kiangsi. The plant has been found growing on ality hillsides, in shady alleys treans Kiangsi, China, on July 12, 1921, and — deposited in the herbar- destroyed. places in v; 7 Pp. adsides, near s = on dry level land, at altitudes of 600 to 9 rs, flows to July flowers are described as "white" on Hu ay nes & ae Lau 20221, and Tsang 20416, and as "pinkish-white” on on T Prei (1 oy in his description, — that the leaves are "5— mm. tin 1428" but this is surely an r for "lata"; ke notes Hu 112 carpa collina Diels from which it differs by its sessile and cor- date leaves which are nearly glabrous, while the leaves of C. col- lina Diels are pubescent beneath at least along the nerves, ~ cune= ate at base and narrow lanceolate." His C. brevipes f. = sis is said to differ from the typical form of C. brevipes and leaves, the blades ovate-lanceolate, acuminate at the apex, sv cordate to rounded at the base, crena rrate re 1 in woods at Yingtak, Kwangtung, China, in July, 1929, and is de- — in the Britton Herbarium at the New York i CEN, Garden. In all, 7 herbarium specimens and one mounted photograph, in- cluding a or papeerare material of all the names involved, have examined by me aoe CHINA: Kiangsi: Hu 1250 (N--photo of isotype). Kwang- s S. XY. Lau 20221 (N), 2027h (N)5 OF W. T. Tsang 20416 (Ba, N, bt ~ 20606 (N); x. K. Wang 290 CARPA CRASSINERVIS Urb., Symb. Antil. 7: na gl. Additional & emended synonymy: Callic va Griseb. ex Urb», Symb. Antil. 7: 357, in syn (in part). 1911 ce C. fulva A. Riche, 1850, nor A. Rich. apud Millsp., ee A a Yugaefolia C- Lect. 9: 2, sphalm. 1937, Bibliography: A. Rich in in Sagra, Hist. Cuba 2 ans 115. agi te Pl. Bright... 2s 2: 529. 1862; Prain a, Ind. + Spec . 393 #500 (1936) foe" riot sts, 65—67, “hs, 128, & 131. 19%; Moldenke, Brief Course Syst- 1966 Moldenke, Monograph of Callicarpa 149 Bot. Lect. 9: 2. 1937; Moldenke, pone Distrib. Avicemn. ). 1939; ca Brief Course, Syst. Bot., ed. 2, 8. 1939; Moldenke, relim. Alph, List Invalid =F ames 10 & 13. 190; Moldenke, Known - Distrib. Verbenac., e - 1, 2h & 86. 1925 Moldenke, Alph. 13. 19h9 Moldenke, Known Geogr. Dis trib. Verbenac., ed. ery ‘ye & 177. 19493; Alain in Leén & Alain Cuba : 305 & 309. 19573 Moldenke, Résumé 50, 243, 2h7, & M3. 3 1959; Moldenke, Phytologia ba — 13593 Moldenke, Résumé 'S Suppl. 13: 6. 19663 J. A. Clark, en. Sp. Pl. n.d. ver oe be noted here that C. fulva fulva Griseb. is in part C. crassinervis Urb. and in part Cc. grisebachii Urb.3 A. fulva he Rich. is a valid species, but Cc. fulva "A. Rich. apud Millsp." is C. hitchcockii Millsp. In all, 31 herbarium specimens, including type material of all the names "involved, and 22 mounted photographs and other illustra- tions have been e Additional & emended pa Beye CUBA: Oriente: Acufla, Alonso, & Pina eee oe Clemente 6617 (2)5 aoe ee NG ie type, Mi—photo of isotype, S—-photo of type, W——photo of type). CARPA CUBENSIS Urb., Symb. Antil. 5: pa mst Fendt Additional & emended synonymy: Aegiphila incana +, Bull. Soc. Imp. Nat. Mose. 3% (2): 218. 1863 [not Callicarpa estar Roxb., 1814]. Callicarpa reticulata A. Rich. apud Urb., Symb. cubensis var, cubensis Alain in Leén & Die Fl. Cuba : 305. Bibliography: Sw., Prodr. 31. 1788; Schau. eA li: 62. 18 1847; A. Rich. in Sagra, ist. Cuba 1): “th. 78505 pst ks Pi. Brit. West Ind. 199. 1861; tures, B et ‘pal. ea von 2): 218. 1863; Sauv., Fl. Spat Anil cote ook. f. papi resieg few. 2: Ub. 18995 tide, Antil. 86 (1 and 7: rage 1911; ° Kew : OE oe oS eee et — Bei (17): 108. 150 PHYTOLOGIA Vol. 14, no. 3 denke, Prelim. Alph. List Invalid Names 2 & ene 1940; Molden- Known Geogr. Dis compa Phew “ep bos ed. 1, 2h, 2 & ish Moldenke, Alph. Invalid Names 2'& 92-11. 1942; Molden- e, Phytologia 2: 9h. 19h dilatkoss8 ‘Alph. List Cit. 1: ki, 1S, 63, 65, 66, 112, 136, 18h, 186, 187, 208, 2h6, 273, 286, 307, nee 470, ust, 490, 199, es > his. me Moldenke, Inform. Mold. Set 6 Spec. 2.19 Recg Biol. Abstr. 26: 643. 1952; Roig y Mesa, Dicc. 3172 from sees de 5 Gamemanes riinethcer C. F. Baker 5176 and Van Hermann 5078 from Havana, and Rugel 352 from Cajimar, Havana, deposited in in the bevbartie of the nes moon Museum in Berlin, but now destroyed. Of these, Sagra 210 has been chosen as the lectotype. The various Sagra specimens without he: i rugosa is Sessé, Mocifio, Castillo, & Maldonado — 516, from some~ in Jescettadta the Madrid herbarium pe (1 1908) observes that "Planta jamaicensis, C. reticulata anc ana recentioribus non collecta et in museo Holmiensi - Lindman in lit. deficiens e descriptione foliis elliptico- taneaanite ; wats breviter ae stylo bifido, baccis extus sericeis sine dub of his C. selleana (1929) he says rehire Calli siigs sdbices cubensi Urb., quae foliis Tis basi Phases ha margine integris, inflore escentiis amplioribus distat". n speaking of his C. sordida (1911) Fs notes "Longius distant Gailtesepe ecsinatd s Urb." In my 1936 work | = I adopted the name, C. incana rons Lorde Moldenke, for this taxon, but this binomial prove ater homonym of c. ae of Roxburgh [=C macrophy! la Pont * wea of the tated by me at t that time were returned to various herbaria mith the incorrect anno ectors describe C. cubensis as a sin to 1.8 nm. a pink, light pink-violet, lilac, or purple the fruit pink. It has been found growing in thickets, thickets near b dges, ruit- 168h0, "light hae seistas’ on Ekman 17316, "pink" on Webster 3709, 1966 Moldenke, Monograph of Callicarpa 151 and "purple" on C. Wright 3172. It should be noted here that the C. reticulata Sw., referred to in the synonymy above, is a valid species, while C. reticulata Poepp. is a synonym of Petitia domingensis Jacq. Turezaninow's original description of Aegiphila incana (1863) is: "Ae. ramis, petiolis et panicula pulverulento-incanis; foliis ellipticis, ovatis obovatisve utrinque obtusis, rarius apice culis, margine revolutis integerrimis, crebre reticulatis, supra pulverulentis punctatisque, subtus ob rete elevatum rugos- issimis, flavido-albicantibus; cymis bus folia quantibus; calycibus turbinatis corollae tubo latioribus atque sublongioribus; genitalibus exsertis. Folia in genere parva, pollice breviora. , Ramon de la Sagra." Roig y Mesa (1953) records the vernacular name "cerato" and notes * abanas de serpentina.....Es una espec= ie de Filigrana, comfin en Guabasiabo, Holguin". The names a and "filigrana de mazorquilla" are also recorded for t. rial has been misidentified and distributed in herbaria of C. cubensis at my request and presented the following report: "I + This information is supplied herewith by means of di- agrams at approximately x10 diam. magnification, with the follow- ing observations. "The material fell roughly into two groups based on the depth of the sims between the lobes and the acuteness of the lobe it~ self. There was also some correlation between this character one other — namely the pubescence of the inflorescence axis and stem. No other correlations — e.g., leaf-shape, size, pubes~ — could definitely be c le stages. The pubescence of the eemed to fairly well correlated with inflorescence axi is a list of the spec examined: (1) With low and fairly blunt calyx- 4nflorescence wi teeth, the e with ence: Britton, Britton, & Shafer 126, Britton & Earle 7607, Britton, Earle, & Wilson Lob, Leén 971, Rugel 352, Sagra s-n. (2) With prominent and somewhat sharper calyx-teeth, the inflores- cence axis with an obviously subpilose pubescence: Britton & Wil~ son 5651, Curbelo 619i, Leén, Edmund, & Fortfin 8566. (3) Two in- termediates were also noted with the deeper simses of the soonae group, but with the blunt teeth and short pubescence of the firs 5078 thought that the calyx was culiar way of drying with the points 152 PHYTOLOGIA Vol. 14, no. 3 collection). Since the sinuses are heavily covered _— hairs and the points less so, the hairs tend to ‘level off' the calyx~ rim to the casual observer. In the drawings I have eae to lg- nore the pubescence of the calyx." Casual observation Optical section ee Britton, Britton, & Shafer 126 ay Britton, Earle, & Wilson 622 Just prior % young ruit ( ] { j Britton & Earle 7607 Leén 97 on 10k Britton & 1966 Moldenke, Monograph of Callicarpa 153 Me. Rugel 352 aa t 4 bud prior to Britton & Wilson 565) Curbelo Sigh oo flowers young t young fruit Leén, Edmund, & Forttin 8566 C. F. Baker 5176 15h PHYTOLOGIA Vol. 14, no. 3 In all, 13) herbarium specimens and 0 mounted photographs, in- cluding type or phototype material of all the names involved, have been examine tddena: & emended dod citations: CUBA: Camagtfey: Leén 16263 (Ha, N). Havana: C. F. Baker 5126 (Ca—10132, rT, ee Oa, Po—64812, W297); Br Britton, Earle, & Wilson 622 (F— 281179) ; Guio 13 (Q); Leén 250 (Se—28355), 971 (Ha, Vi), 5689 (Ha, N), 6238 (Ha, N, Vi), 7218 (Ha, Ha, N), 13757 (Ha, N, I— 1632h); Moldenke & Moldenke , 19866 (Es, le; N, Ot, Sm); Sagra 12 (De, De, De), 211 210 (E—photo of type, F—976786—isotype, Mi —photo of isotype, S—photo of type, V—isotype, W—photo of type); Van Hermann 5078 (Po--64814); G. L. Webster 3709 (Mi, Mi); C . Wright 3172 [1860--186),; Herb. Sauvalle 1775] (E—-119135, F—2hh618, Hv, I—photo, Pa), 3172 (1865; Herb. Sauvalle 1775) (Hv). Las Villas: Alain 4003 (2); Bri Britton & Wilson 565) (Mi--photo, W—65869l); Ek- man man 16840 (Mi, N); Leén — Edmund, & Forttin 8566 (Ha). Matanzas: Britton, Britton Britton, & Shafer 126 (cm). Pin Pinar del Rfo: Acafe & & lain 15 15837 (Bs), (Zs), 18545 (Es); Acufia & Roig 10872 (Es); Acufla 4.1209 (N); "Britton & Earle 7607 (F—205759, a an) Ekman 12875 (N), 17316 (Mi, N). Province undetermined: Auber (D—€12061, D—612062); Collector undesignated 15 (Q); Herb. Short s.n. (E—11913k); Sagra 514 (De); Serre s.n. (P); “Sessé, Mocifio, Ca Castillo, & Maldonado 516 [296] (F—850973, Q). CALLICARPA CUBENSIS var. PARVIFLORA Moldenke, Phytologia 3: 487— Synonymy: Callicarpa cubensis var. parvifolia Moldenke, Phyto- logia 3: 48h, nom. mud. 19 Bibliography: Moldenke , Phytol aoe 3: 48h & 487-88. 19515 Moldenke, Inform, Mold. Set 46 Spec. 2. 1951; Moldenke, Biol. Abstr. 26: 643 & 3h)6. 1952) Mo Moldenke, Phytologia : 450. 19533 Alain in Leén : uba 4: 305. 1957; Moldenke, Résumé 50, 2h2, 418, & 43. ig This varie having its leaf-blades much smaller and narrower, 1.2——-2.7 Cie when ma ture. The type of the variety was collected by Juli4n Baldomero A- cufla Galé and Juan Tom4s Roig y Mesa (no. 16765) at La Cajabana, La Palma, Pinar del Rfo, Cuba, on March 8, 1951, and is deposited rk cal G unfortunate error in transcription, the varietal name was of- Hotaiy and hee dma d published as "parviflora". There are, how- lier times! Material of this variety has been misidentified and distribu- ted in herbaria as C. shaferi Britton & P. Wils. In all, 3 her- 1966 Moldenke, Monograph of Callicarpa 155 barium specimens, including type material of both names involved, have been examined by me. Citations: CUBA: Pinar del Rfo: Acufla & Roig 16765 (Es-~iso- type, Lm—isotype, N-type). RES oan ageetpane Britton & P. Wils., Mem. Torrey Bot. Club : 976 Biblio acer; gree & P. Wils., Mem. Torrey Bot. C ub 16: 97. 1920; Urb. in Fedde, Repert. Spec. Nov. 18: 120. 1922 A.W. Hill, Ind. Kew. cig 6: 3h (1926) and 7: Sa 1929; Moldenke in Tt. 5 129, & 130. 1936, “Woldenke, Geogr. Distrib. > reo he 19393 Moldenke, Known Geogr. Distrib. Verbe: ed. 1, 2h & 86. get Moldenke, Alph. List Cit. 1: 18h—186 "(A9h6), 3 929 (19h9), and ks 1016 & 103k. 1949; Moldenke, Known mi. re Distrib. Verbenac., ed. 2, 42 & 177. 199; oe Rev. Soc. Cub. Bot. 13: 37. 1956; Alain in Leén & Alain, Fl. Cub. h: 305 & 307. 1957; Conde, Hist. Bot. Cub, 221. “1958; TAT “Résumé 50 & hh3. 1959; J. A. Clark, - Ind. Gen. Sp. *Pl. n Morton found this plant growing in Pinus cubensis woods on limo and says: “agrees with isotype of cuneifolia in U. S. Nat. Herb., but in my opinion the species is too close to C. fer- a Swartz. C. revoluta Moldenke seems to be the same also". Material has been misidentified and distributed in herbaria as C. nipensis Britton ne P. Wils. In all, 11 herbarium specimens, including the type, and 6 ted hothenbohis: hi have been examined by me. Additional & emended eee CUBA: Oriente: Alain & Lépez Figueiras 848 (Z); Ekman 3497a (N); Marie-Victorin & n & Clément 21730 (Uim--25259, ba os7e); Shafer TTL (W—696369—isotype) - rome eg DENTICULATA Merr., Philip. Journ. Sci. Bot. 3: 30-- Bibli ography: E. D. Merr., Philip. Journ. Sci. Bot. h: 430-- 431. 1909; Prain, Ind. Kew. Suppl. h: 3h. 1913; H. J. Lam, Ver- benac. Malay. Arch. h8, 65-66, & [361]. 1919; Bakh., Bull. : Py OTT e, Jard. Bot Z 27. 1921; E. D. Merr., Emm. Philip. Pl. 3: 383. 1923; Moldenke, 3 Common Verb. 21. 1939; Moldenke er ed. 1, 62& be 19 25 ldenke, Phytologia 2: 94. 1945; Moldenke, b. L-} ke, Résumé 1 2 & 443. 1959; Hatusima, Mem. Fac. Agr. oshima Univ. 5 (3)s- 17 & 7. 1966; Moldenke, Résumé Suppl. 1h: 4. 1966. Shrub or small tall; branc eht-ereys glab- rous b hl ry 2——3 Be aly ote ’ 3 branchlets densely or very dens s umose~pubescent; leaves sted pitisies 1—1.5 cm. long, densely ste or stellate-pl 156 PHYTOLOGIA«A Vol. 14, no. 3 ero minute, yellow glands beneath; secondaries 5 or 6 pairs, very prominent; veinlet reticulation distinct, brown; cymes solitary, hairy widely scattered hairs, the lobes elliptic, about 1 mm. long, ob- tuse at the apex; s stamens 6 mm, long, long-exserted; Solieene a- bout 6 mm. long; anthers glandular, 1.6 mm. long; style very slender, 1 cm. long; ovary somewhat daccegit dates fruit glo- bose, about 3 mm. wide. The species was based by Merrill on two collections made by Eugenio Fénix, one [Philip. Bur. Sci. 023] from Camiguin, and the other [Philip. Bur. Bur. Sci. 3622] from Santo Domingo de Basco, Batan, Philippine Islands, on May 30 30, 1907, both a rapeod in the herbar- ium of the Bureau of Science in ii » but now unfortunately stroyed. Merrill (1909) says: "A species oi eetee ae by its relatively broad leaves, few cymes, and these only in the up- per axils, the very long-exserted stamens and style. It has more the facies = — than of Callicarpa." The common names “anaif and "mayop recorded ~ this plant, and it has been found growing 4 the he ithe ral b Lam (1919) notes also that "Its habit reminds us that of a Premna", Hatusima (1966) places C. kotoensis Hayata, C. japonica belonging in the synonymy of C. japonica var. lumrians Rehd. His Botel Tobago i therefore, is to be excluded. He also records CG. denticulata from Babuyan and cites his nos. 28691 and 29198 - The "Datan® cay in my Résumé (1959) sess to be a 2 typographic — for "Batan", The species is known to only from the liter- CALLICARPA DICHOTOMA (Lour.) K. Kooh, Dendrol. 236. 1872. Additional & emended syno: dichotoma Lour., Fl. Cochinch., ed. 1, 1: 70. 1790. Mme dichotoma Raeusch., Nom. Bot., ed. 3, 37, nom. md. 1797. Callic purpurea A. L. JusSe, Ann. Mus. Hist. Nat. Paris 7: 67. TE0S. Gallicarpe Toliis subsee subses- silibus, lanceolatis, serratis; racemis dichotomis Poir. in Lalle, sla Méth. Suppl. 2: 34, in syn. 1011. Callicarpa fruticosa: ramulis parti Sr ea ee eat silibus, ianceolatis, S, serratis, basi cunea teg- errimis, supra glabris, sibtus ose punctatis; corymbis ax- bus, parvis, dichotomis; floribus glandulosis Roxb. apud J- 1966 Moldenke, Monograph of Callicarpa 157 A. & J. H. Schult., Mant. 3: 54, in syn. 1827. calliearps Eee ilis Rheede ex Sieb., Jaarb. Konink. Nederl. Maa anmoed. uw [Ann, Hort. Pays-Bas], ser. 3, 1845: 72. Teh . Cal Callicar- pa gracilis Sieb. & Zucc., Abh. Akad. Muench. 3 (4): 15-155. 1846. Callic jamamurasaki Sieb, ex Miq., Ann. Mus. Bot. Lugd.-Bat. 2: 98. 1865. Callicarpa japonica var. angustifolia Seraiier. Livres Kwa-wi 78. 1873. Callicarpa sieboldii Zipp. ex oJ. Lan, Verbenac. Malay. Arch. 83, in syn. 1919. Callicarpa mabihane (Lour.) Raeusch, apud Bakh., Bull. Jard. Bot. Buitenz., ser, ane 3: 25, in syn, 1921; Nakai, Fl. Sylv. Kor. 1): ee slept Calli aponica var. dichotoma (Lour our.) Bakh., t. Slteas ser. 3, 3: 125. 1925. Callicarpa dichotoma spam ty: Merr. ex P. Dop, Bull. Soc. Hist. Nat. igrcdng 6h: re 1932. Callic Carpa koreana Hort. ex Viln., Dendr. e 87: 82, nom. nud. 1933. Galphimia hirsuta Hort. ex Moldenke ¥ peal. Repert, Spec. Nov. 0: 84, in syn. 1936 [not G. hirsuta Cav., 1799]. Callicarpa muricata Hort. ex Moldenke, , Prelim. Alph. List Invalid Names 12, in syn. n syn. 1940. Callicarpa sinica Colebrooke ex Moldenke, Prelim. Alph. List Invalid Names 13, in syn. 190. Cal- iicarpa japonica var. dichotoma Bakh. apud Rehd., Bibl. Cult. Trees 583, in syn. 1949. Callic allicarpa dichotoma K. Koch ex Kasapli- gil, Pl. oct 102. ee Bibliogr: hinch., ed. 1, 1: 70 (1790) and ed. 2 Witla), ‘ls 1 B16 “17935 Rasusch., Nom. Bot., ed. 3, 37- 1797; Cav., Anal. Nat. 1: 37. 1799; Ae L. Juss., Ann. Mus. Hist, Nat. Paris 7: : er 1806; Poir. in Lam., Encycl. Méth. Suppl. a oo Prep Roem. & Schult., Linn. Syst. Veg. 3: 97. 1818; Wall, L)+ 15h “phys. cl. ; ie hago : 154—- 155. "18L6; ‘Sieb. & Zucc., Fl. Jap. Fam. Nat. 2: 30. 18465 Schau. 202. 1859; Lindl. d. Chron. 1859: 96. 18593 Hort., pers h, 8: 7306- & 107. 1859; eee oh & es Guide hs 192 & 193 3, 18615 Boe ep coral 2: 103-105, 129, & 137 (1862) 3s 192 & > ee fig. 8—-ll. 635 Bocq., 103—105, 123, bt 192, & 263, pl. 8, fig. 8—1L . 1863; 334 Aan inn. “ss. Bot. Lugd. “Bat. 2: 98. 1865; pb fat sy Bot. Ind 158 PHYTOLOGIA Vol. 1h, no. 3 1866; Miq., Prol. Fl. Jap. 30. 1866; M. T. Masters, Gard. Chron. ier ae "fg. 38. wr me Koch, De ndrol. 336. 1872; Savatier, Kwa~wi 78. 1873; Ro , Fi. Ind., ed. 3 [C. B. Clarke], cs, 187; Franch. & 08 Enum, Fl. dap. 1: 358. 1875; *& Arbris. 179. 1877; D. Thompson, The Gardener 1880: s18-—519, 1880; Franch., Mém. Soc. Sci. Nat. Cherbourg 2h: hort axim., 509. — Maxim., Bull. - Imp. Sci. St.-Pétersb. 31: 78. Garden. liste 9: 323. o7887; F. ~ Muller, Am. Gard. 10: rs “18895 Forbes & Hemsl., Journ. Linn. Soc. Lond. Bot. 26 — Fl. Sin. 2]: 25h. 1890; G. W. Johnson, sores Diet. 157. 1890; erat Gard. Chron., ser. 3 aah 101'& 103, fig. 13. 1891; Meehan, fyb 6: 490. 1891; F , Am. Flor. 7: 1069. 1892; tennis, ardening 1: 276. 18935 Je Jacks. in Hook. f. & Jacks., Ind. Kew. 1: "386. 18933; Bois, Dict. Hort. 232. Blam ie Jacks « in ok, f. & Jacks., Ind. Kew. 2: 607. 189; S Coll. Agr. Tokyo sete Univ. 2: ae Laubh. fanteraust. sustel ” ‘pl. 1h, fig. 9. 1895; Briq. in Engl. & Prantl, Nat. P (3a): 165 & 166, fig eee. orton 5: Meehan, Gardening sr 9T. “18973 els in Engl., Bot. Jahrb. 29: ShB. 1 1900; W. P. Wright in Cas- - 1902; "xX ey 1907; Robinson & Fern. in A. Gray, New Man. Bot., ed. 7, ”690 & s & 783. “oh; +8) Seatheene: Field Book Am. Trees 363. 19153 Reba. - S. Sarg., Pl. Wils. 3: 370. near hore » Flowers Wint. 1 & 200. 29173 W. Trelease, Wint. Bot., ed. 1, 331. 19183 E. D. Merr-, 1918; Yabe, Prelim. . 96 195 H. Jd. Lan, ver- . o Arch. Sl 83~86, & 362. 1919; Ge Ve Nash Journ. N.Y. Bot. Gard. 21: 58. 1920; , Bull. Jard. Bot. Buitens., Bui sere 3, 3: 25-26 & 125. 1921; Nakai, Fl. Sylv. Kor. 1k: 27--29, pl. 5. 19233 H. H. Chung, Mem. Sci. Soc. China 1 (1): 225. 192h5 Olmstead e, & Ww Nakai, Trees & Shrubs Indig. Jap., ed. 2, 1: 450—l51, fig. 213- 19273 Rehd., Man. Cult. Trees, ed. 1 2, 77 778 . "176. 1927; 8 yee 1928; Blossfeldt, Art Forms in Na seed Tone pl. 9a. 1929; W. J- Bean, 1966 Moldenke, Monograph of Callicarpa 159 Trees & Shrubs Hardy Brit. Pi ed. 5, 1: 280. 1929; Stapf, Ind. sen ae 526. 1929; L. H. & E. Z. Bailey, Hortus 111. 1930; T. W. » Encycl. Gard., ed. 19, 10. 1930; Hottes, te Shrubs, ed. 2, 2, 267." 1931; Macself, Sanders Encycl. Gard., ed. Tl. 1931; - Dop, Bull. Soc. Hist. Nat. Toulouse 6): 198° 5, fol, 507, *& owt, aNeees Pred, Mem. Sci. Soc. China 1 (3): (Verben- ac. China] 1 Fea tee & 5153. 1932; Grey & Hubbard, List Pi Ins Pl. Grow. sor - 38.19 n Sho~ kubutsu Zufn [Jap. a Illustr. Album] 159). 1933; Mak., Gensyoku Yagai-shokubutu {Nature-Col. Wild Pl.] h: 296 on, B ard. R 22: 7. 1933 » Bull. Soc. . France 87: 82. 1933; Macself, San . Gard., ed. 21, pr. 2, 71. 193k; Rehd., Journ ld Arb. 15: 32h. 193h; L. H. Bai- er Fl. Gén. Indo-chine : 798. 19 E. D. Merr., Trans. Am. Philos. Soc., new ser., 2) (2): is a Deis Reamer: Pagel en 2, éla. a Moldenke in = (19 & 3. Moldenke, Geogr. Distrib. sree oe 2& 6. 19395 Poa gris Sone. & Classif. List 108. 1939; Moldenke, Prelin. vat? list Invalid Names 10-13 & 37. wes, Gates, rl. —: 190. 190; Rehd., Man. Broo Bot. Gard. Record 30: “33. "19h; Toredehi, Ind ip Foy Suppl. 1: 160, 191; Hottes, Book Shrubs, ed. }, abr—166. 192; Macself, - Gard., ed. 21, = si - 1942; Van Melle, Journ. N.Y. Bot. Gard. 43: 36 & 2. 1942; Ne : Herald Trib. hi tae Known Geogr. Distrib. Verbenac., ed. 1, 3, ames 5 p. ate RP Ee e re. = R & “y oe ag » 55, 59, y 225, Sg 78, 280, 282, & 299. 196; Van Melle, dogia ~ £255 15472 Tatnall, Fl. Del. 218. 1917; N. R. Sm., N. ¥ , November fig Garden Gate 73 19174 Molienie, Phytologia 5, 387. 1 , Enum. Sperm s ibo—a85. ta its Aliph. tae ‘cit. ite Bs 355 355 30, 51h, & 640 (198), 3: 3 Got, 666, 698, 702 ss fat oe 783 * oor 728 732, ae, 7 , ’ 827, 837, 8 i eg a 997 998, 1011, 1013, ag! 3,051, | 899, a ‘on Bog "tos, 1036 71400, 71102— los; m7 38 1147, 1176, 1177, 1200, 1 1238, 12h2, 1251) & 3, oe ; Moldenke, Phytologia 3: 7138 & 139. 19155 160 PHYTOLOGIA Vol. 14, no. 3 Moldenke, - Distrib. Verbenac., ed. 2, 5, 6, 125, 130, 133, 135, ao, 7, & 177. 1949; Rehd., bibl. Cult. Trees 583. 199; . Times s Spring Gard. Sect. 19, March 19. 1950; N. Y. Bot. salad "Be ed Exch. List 1951: [1]. 3350, W. J. Bean, Trees & Ss 8 - Bi 33 380 (1950) ana 33 449. 19513; W. J. Bean in Chittenden, oe Hort. Soc. Dict. Gard. 1: 358 & 359. 1951; N. Y. Bot. Gard. Exch. List 1952: 2. 1951; Chaput, N. Y. Herald Trib., sect. k, hii September 16. 1951; B.C. Blackburn, = & ee t ct & ih fe 19525 Moldenke in G ew Britton & Br. intast 197; 13058 a5 “1982; eerie py tbiset & & 8, pes Moldenke, Journ. Calif. Hort. Soc. 15: 8 “ty 1955 5 Ease Herald Fl., pr. Krttssmann andb. Laubgeh. 1: 253. 1959; Nut Tree Nurseries Spring '59 Price List. 1959; ge Ta Résumé 8, 9, 160, 168, 171, 172, 175, 182, 213, 242—2h7, 336, & hh3. 1959; Rehd., Man. Cult. Trees, ed. 2 3 Kelsey Nurs Eve = yel. Gard ‘ 1960 Moldenke, Phytologia 7: 429. 1961; Moldenke, Biol. 36: Loss. eee ere — Su Sek: 39 33 27 ( (1 962) and a8 kn 434. 196; Santamour, Morri Psa = 2 16: 52. 1965; Moldenke, Résumé Suppl. 1): ea & 6. 19663 Pie age gia 13: 72 & pl. 29 [in color]. abe Ou Ae Hérinca, Hort. Frang., ser. 2, 3 pe ms baer Floral World & Gard. Guide lh: 192. Scan A Bocq., Adan- sonia 3: pl. 8, fig. 8-11. 1863; Lig Rev. Vi NAC » 8 fig. pe 1863; M. T. Masters, - Chron. 1871: 173 1871; W. Robinson, The Garden 23: nae. *392 fin color]. 1883; Gard. Chron., ser. 3, 10: 101, fig. 13. 1891; Shirasawa “Asad fig.’ 38-_ ie 1911; Varacek, Oesterr, Gart.-Zeit. 8: 277, fig. 115 115. Fl. Sylv. Kor. 14: pl. 5. 1923; Nakai, Trees & 5 radi Jape» - 2, 1: 451, fig. 213. 1927; Hottes, Book S Sowa a 1, 148. 1928; Blossfeldt, Art Forma in Nature’ pl. 9a. 1929; Hottes, Book 1966 Moldenke, Monograph of Callicarpa 161 Shrubs, ed. 2, 167. 1931; Terasaki, Nippon Shokubutsu Zufu [Jap. Bot. Illustr. * Album] 1591. 1933; Mak., Gensyoku Yagai-shokubutu Epeeeconsieas Wild Pl.) 4: 296 [in color]. 1933; Hottes, Book s ed. 3, 167 (2237) and ed. h, 167. 1923 N. x. Herald Trib. , Debiber 25. 19423 No Re Smo, Neo Yo Times s 22x, November 9. 19h73 haput, N. Y. os Trib., sect. 4, 17, September oy 1951; Vol- denke in Gleason, New Britton & Br - Illustr. Fl., pr. 1, 3: 138. Heral ° tus, 3- Ss: ramis auitte ; routes » debilibus. Folia lanceolata, ser- rata, punctata, subsessilia, opposita. Flos purpureus, racemis dichotomis, axillaribus. Bacca globosa, violacea, carnosa, plur- , minima, Habitat in collibus provinciae Cantoniensis Sinarum." The type of C. sinica was collected by Nathaniel Wallich (no. 523) from cultivated material in the Calcutta Botanical Garden, Bengal, India, and is deposited in the herbarium of the Universitets Bo- taniske Museum in Copenhagen. The type of C. muricata i s Herb. Hort. Bot. Berol. s s.n. deposited in the herbarium of the “mae historiska Riksmuseum at Stockholm. Recent smerny describe this arid as a shrub, small shrub, undershrub, or bush, woody, erect, bushy, or even as a climb- er or vine. [Chiao 2787), 2--10 feet ett with a spread of : meter, the stem to 1 inch in diameter, the branches purplis bark pinkish, buds lavender, flowers fragrant, corolla aie ‘pink pete purplish-pink to eifictnbered purplish, purple, white, e yellow, red, lish-red, or "somewhat n, lilac Purplish-lilac, lavender, pasets violet, Mpeautifully violet", or even "black", shiny, 1/12 inch in diameter. The Nut ci s Nurser- ies catalogue (19595 ieastahee hd fruit as "berries in pe indenies ropes". The flowers are said to have been "pink" on Charette 1896, Lau 2011, Tsang 23862, and Tso 20h07, "pinkish" on Chiao 27687, — *pinkish-red” on on Hu 1634, be aera on Tso 815, "purplish" on Keng 72, "purple" on Peng, Tak, & Kin 199, "white" on Chiao 2617 and Ts. Tsang 2250, and "white @ and red" on Tsui 395. ~~ The species has been found growing on hills and | brushy or rocky Slopes, under the Pers of bamboo, in good soil, dry sandy soil, or silt, in thickets, woods, and forests, loose summer-green woods, Pine woods, open sites and open fields, eys, swamps, upland Peat bogs, and rocky streambeds, along s streamsides and Siaaatte: at the foot of hills, the base of mountains, and the base of high frui une t . In India it pag from April to September and the fruits ripen at the close of the rainy season. Ching states that it is common in Anhwei, but rare in Kwangsi; Wilson reports it rare in Korea} Fosberg found it "rare in hedgerows between cultivated fields" on 162 PEYTOLOGIA Vol. 14, no. 3 Ikema island. Tsang describes it as fairly common in thickets in K t only as "scattered shrubs” in — 1. und low bank of the "Roanoke River in Martin coonky,53 North Carolina; Radford found it in an upland peat bog in Henderson County. Tat- nall (1947) avers that the historic Delaware station is now de- stroyed. Gates (190 0) ee that it is "seldom spreading" from cultivation in Riley C as. He refers to it as a nano- phanerophyte. Wisler (19122-1913) lists it as cultivated in Dela- ware County, Pennsylvania, I have personally seen it in cultiva- tion at Great Neck, Long Island, New York, in November, 1938, and at Plainfield, New. Jersey, in 1966, but have not made collections from these plants. It was recorded by me (1952) as naturalized in Loudoun County, Virginia, Herb. Arnold Arb. 10151 was growing in Massachusetts from seed collected in Korea ‘ea by E. H. Wilson. —— (1956) records it as cultivated in Jordan. Common names for the plant include ee Yo berry", "beauty- berry", "Bermuda mulberry", "callicarpe 4 fl s purpurines", "Chinese beautyberry", "Chinese baladearhens “WFrench-mulberry", muras "ko-murasaki" no formal description, so his binomial must be rejected under the present edition of the International Rules of Botanic Nomencla- ture, although it seems obvious that he prrpeass flaky binomial as a new ic ecanipsan for L = eereinrie 0 rphyra dichotoma, and many later 2 include C, ra H. J. Lam in the synonymy of C, dichotoma, 45 also did in my 1936 work, but Lam's plant is now regarded by me lamii Ho fesaiane, hotoma var. koreana Hort. is regarded ¢. japonica ca Thunb. Roemer & Asin yi (7818) describe the fruits of the present 1966 Moldenke, Monograph of Callicarpa 163 plant as "baccis trispermis"., Jussieu (1806), Roxburgh —_ 5 Hooker & Arnott (1828), and Walpers rs (1845) all refer to the in- florescences as "corymbs"; actually they are cymes. S aan (1965) reports the chromosome number as n = 18, and regards the species as tetraploid. Wallich (1820) says "I cannot but consider this as Loureiro's quoted above which Jussieu has justly placed among the gems Callicarpa. It seems to be nearly allied to C. japonica, but dif- fers chiefly in wanting the smoothness, the short stamina and pis- um and the acute stigma of the latter. The flowers are be- sidea of a retest Rag re SOLASS > especially within, while those of Thunbe white." Hooker : ary (1828) say "Of this we have two forms before us, That from Bonin has oblong-lanceolate gr ted axi leaves, and corymbs: the other, Loo » has round- ish-oval suddenly acuminated leaves, inflorescence from @ uppermost axils only. In both, the young foliage is covered, particularly on the under side, with escence, but in the adult state it is Sabie is spects both agree with Roxburgh's eeettsotice in the Flora Indica.” Their Ryukiu Islands collection was probably Cc. sapoiiee ca Thunb., and that from the Bo- nin Islands was phir: C. glabra Koidz. ly c Rehder (1916) says of Cc. dichotoma: "This species is closely related to C. japonica Thunberg, but can be distinguished from it by the young branchlets being slightly grooved or obscurely angled, by the smaller, less acuminate leaves with fewer teeth ra slender-stalk orescence, and by the smaller flowers with the anthers dehiscent to the base. It is of more southern distribu- - China, Korea, cult. apan; rarely escaped in E. Intr. 1857. Zone (V)." Bean (19 951) also ete 1857 as the date of introduction into cultivation, but Van — (1947) maintains that 18h5 is the correct date. According to Roxburgh, however, species was introduced = cultivation am the Botanic Garden at Calcutta pate oe in 1 (1919), a etn ard , elegans Hayek, says: "Its affin- ity is with C. purpurea. Points of sone are the acuminate apex of the leaves, which is never abrupt, and always serrulate, upper side without any hairs and with some Ag and corolla- lobes as long as the tube." In speaking 0 GC. Jepontcs Thunb., he notes that "Its affinity is also with C. Ptei (1932) says: "Callicarpa dichotoma dichotoma (Lour.) ‘Raeuch., Cc. Japonica Thunb., and C. japonica Thunb. var. angustata Rehd. are ng tC. japonica......--Vale ta.....differs trom C. dichotoma.......which has has short ovate leaves, and which are 16) PETTOLOGAIA Vol. 1h, no. 3 toothed only in the upper part. C. japonica.....differs from C. ee its variety by its broadly ovate leaves which are usually suddenly eta sce at the apex. The anthers of C. japon rae cane C. japonica + angustata.....usually open by later oe sab while those ec: Cc. “dichotoma.....open by longitudinal Li (1963) notes that "This species is native to Korea, Japan and central and southern China, It is perhaps the most widely planted species of Gal ttbarpat: It flowers in August and fruits in around October through November. In its native habitat, it prefers aoe places and grows especially abundantly at base of hills waters.....It was introduced into western gardens around 1857 | by Robert Fortune. Lindley (1859) recorded the first fruiting plant in exhibit at the autumn 1858 meeting of the Horti- cultural Society in St. James' Hall in London ‘excited more in- terest’ than any seme plant. This species is closely allied to ies axill ores dehiscing by a longitudinal slit instead of an apical pore. It is gp Figen cba as in its original home it is generally of 4 more etley “(2935) Uh ee as handlers of ppeian Sanford, the c Garden, L. Sp&th (Beran), 3 - T. Hood & Company Maryland Nur= jewport, e Island), Outpost Nurseries (Ridgefield, sp aurea Henry F. Witchel1 (Philadelphia, Pennsylvania), Hill Top (Hartford Michigan), Joseph Breck & Sons (Boston, Massachusetts), Rosedale Nurseries (Tar Eastern Nurser n, Massachu- setts), and Kallay Brothers (Painesville, ohio). Mattoon (1958) | s sources, In 1959 it was selling in the United States 4 ee for a 3}foot bush wane greenhouse plant, we. cultivation although rather in habit. Its shoots are clothed with site serrate leaves, and it °. e or ame ttings, and young —— 7 olaptoal ne mended for Ardisias are best; old plants 1 tom foliage, and become 'leggy' and unsightly.” In 1892 ¢ Sag pers yay in 1966 Moldenke, Monograph of Callicarpa 165 pots, at chrysanthemum time for $5 apiece. Bush-Brown (1963) al g drastic abs in early spring is needed to restore its vigor." Of particular interest are some amusing but on the whole jus- tifiable | Sad the horticultural standpoint) remarks made by Mas- ters (1871). He says, in part, concerning the genus Callicarpa, but with special reference to C. dichotoma: "See them when in ar their company'; but see them when in fruit ’ and every one cries out how beautifult......Two or three species have from time to time been introduced into our gardens, and some even were figur— that criminals portraiture of the photographic camera, in order that they may be recognized in the future, and their worthlessness made apparent. carpa with purple berries, so beautiful that every spectator was loud in its praise. Dr. Lindley led the way in lavishi li- ments on the plant, and as it would never do to let such a beauty g0 unnamed, he called it 'Callicarpa aod Saal Very sagaciously he out a im after the name, knowing by experience that it is no easy matter to determine the — of Callic Varacez (1913) comments that "Im Sp&tsommer aes die Kelche der zahlreichen Blumen und sind erst grttn, f¥rben sich je- doch etwa Ende September zu einem wunderbaren, auffallenden a? C. purpurea ihren H&hepunkt erreicht und verbleibt so dutch den ganzen Herbst und den halben Winter hindurch. Die Beeren anthal- dieser Menge von Samen sollte man annehme en, die schtne Pflanze kOnnte in erbentont sein. Dich welche 'geheimnisvolle Her- kunft' hat diese Spezies{" Van aehee (1942) notes that "Callicarpa dichotoma.....is 4 use- ful and decorative Japanese-Chinese shrub, grown mainly for the beauty of its small rounded clusters of lilac berri rne autumn in the axils of the graceful, arching branches of the cur- rent season's . A unique and valuable effect, not duplica- ° - To about l ft. high, fairly ve er Sark City. If occasionally killed to the ground, it will de- P 8 on base. In his 193 work, this same author elaborates as follows: "Calli- carpa dichotoma......and C. japonica.....are small shrubs, normal- ly about 5 to 7 feet high, ae perfectly -—- i.e., not reliably — hardy in our zone and likely to be killed back at least partly; @ 166 PUT-TO LOCA Vol. 1h, no. 3 matter which does not interfere greatly with their usefulness since they flower and fruit on growth of the current season and may be treated as die-backs. In that case they will grow little more than 3 to feet high. Their one, but considerable, contri- bution to cig border the ee of their attractive, clustered, —_ berries s the branches, ripening in and remai ng time h rect-growing, with darker, long-tapering leaves to 6 inches long. Ce dichotoma is of a more ee’ nna and has smaller, shorter, light-green leaves. ~~ tious as to soil and succeed They are worth poms in a small way for the =— of their wean det iae; which are among the most decorative autumn effects." Hottes (19h2) says "The Beautyberries produce attractive leaves and the shrubs are quite graceful though upright. In plan them, give a rather protected place. Although the fruits are The Tsui 601 collection, cited below, bears great resemblance to C. japonica Thunb. Pételot 3039 is athe not typical — its leaves are large and the cymes cymes long~pedunculate. F. C. Gates 16499 exhibits a fasciated branch. Hayek (1906), in speaking of C. elegans Hayek, says: "a Cc. Juss. foliis coriaceis nec chartaceis in acumen elongatum productis, rele dentatis nec cre punctis glandulosis aureis nitidis ec purpureis a" Schauer (187) — Colebrook s.n. and Wallich 1828 from China, a 8.n. from Cochinchina, and Zollinger 340 from Japan. Mi- quel (1865) cites hoetiiy 3h8 from Japan. Maximowicz cites Ford S- . Rehd oh, (1916) cites the following: CHINA: Chekiang: D. MacGregor 5-Ne [Ningpo, 1908]; F. ue Meyer 425. Fulcien: Dunn s s.n, [Hongkong ong Herb. 3383]. Kwangtung: Hance 335. JAPAN: Honshiu: Herb. K. Sakurai S- n. [Omiya, June 18, 1911]; M Maximowicz s.n. [Yokohama, i862]. Kiu- Shiu: Oldham 628, Ptei (i932) cites the following: CHINA: Anhwei: Ching 2766; IP Sn. Chekiang: Ching 3803; Hu eee & 1634; Keng 72 & 916; D. Mac Gregor s.n.; F. N. Meyer 25. : A. Ne Steward 171y. Hunan: 1966 Moldenke, Monograph of Callicarpa 167 Handel-Mazzetti 2691. Kiangsi: Chun 42793; Ip i A. N. Steward 8.n.; Wang-te-Hui a 235 « Kiangsu: Hancock sn; Hers 2301; Ling 2489; A. Ne Steward s Seno; Tso 815 & 1716. Steere = Chun 5538; Sotensllloeseeetliemmmmmmceeneeens aati Speer Semen nee , ISLANDS: Hon: A. Henry 435, but I regard this as C. formosana Rolfe. Material of Cc. dichotoma has been wn misidentified and distributed — a —_—_——— a cementite caer carrie SeNRCEReN SR, 8065, and Tak & Chow 2708, distributed as C. dichotoma, are actu- ally C. bodinieri Lévl.; C. H. Cheng 3080, R. C. Ching 1652 "1652 & 639k, M. L. Hancock s.n. [Herb. Univ. Nanking ng 12075], . Herb. Univ. Nen- TTT & 779k, 1127 & 1243, BE. D. Merrill 11298, and Peng, Bu Ta & Kin Gein eke 0 Chr. Coll. 12677) are C. bodinieri var. giraldii (Hesse) Rehd.; Swartz s.n. [Jamaica] is the type collection of C. ferruginea Sw.3 Ae A. Henry 435 is C. formosana Rolfe; Chiao sen. {Herb. Univ. Nanking 18896] is C. japonica Thunb.; Barchet 557, Cheng 578 & 390k, Chiao 1606, , 1612, & 2916, Re C.C Ching 5666, Hayakawa's c collector Ads, Herb. Canton Chr. r. Coll. 12677, Herd. Univ. Nanking 775k, 18595, & . Lau L109, C. 0. Levine s.n. (Herb. Canton Chr. Coll. ste & coe Ling ng 1189, E. D. Merrill 11212, A. N. Steward s.n. (Herb. Univ. Nanking 2426], ee Arnold Arboretun herbarium, is Ligustrum rset z; i Nakai in the Oleaceae, but the United States National tional Herbarium specimen of the same sane mmber, from Ullung Island, is plainly C. dichotoma (although its label is inscribed "Lilium tigrimm"). Apparently there has been some serious mix-up in labeling involved here. re aioe 286 herbarium and fruit specimens and 11 mounted photo- ve been emma gare tn & emended steite tions: NORTH CAROLINA: Henderson es Radford 189 (Hi—1,8259) . geet Co.: Fox, ee & Boyce 44392 as INDIA: Uttar Pradesh: W. Griffith 335 (S)- State undeter- mined: W. Griffith s.n. (oer (S); jee 1; (S). CHINA: } (c De conic . Nanking 2). Be Se sg oa Bi B53 (ca-_943265); Hu 163k (Ca—362316); BN ee oe eens (ca Meyer 425 (ar--19785, Ca—218962, Du— 168 Pat tO 20-@ 5:4 Vol. 1h, no. 3 A. N. Steward 171, (Herb. Univ. Nanking 98)2] (Ca--270559, W-- 1370062). Hunan: Fan & Li 353 (Bz--17568), 487 (Bz--17564). Ki- angsi: Ip 119 [Herb. Univ. Nanking 770] (Ca—258866); S.K. Lm 4521 (S, W--1753185); A. N. Steward sen, [Herb. Univ. Nanking 2714) (Ca—230391, W—1279529); Tsiang 9882 (N). Kiangsu: H. 7. Chang 34h (Du—-250177) ; M. L. Hancock s.n. [Herb. Univ. Nanking 1207h] (Ca—298)8); K. Ling 2489 [Herb. Univ. Nanking 1240] (Ca—316h01); A. N. Steward 2763 (Ph). Kwangsi: Ching 5201 (N); Steward & Cheo 930 (N, S); W. T. Tsang 225h0 (s), 23062 (N), 2uh17 (N). Kwangtung: Herb. Bot. Mus. Stockholm s.n, [Canton] (S)5 S. K. Lau 201) (Bz—-17567, | Ca—611459, N, N); Liou 1660 (N); Peng, Tak, & Kin 199 [Herb. Canton Chr. Coll. 12198] (Ca— 27692, S, W-12)7568), 92h [Herb. Canton Chr. Coll. 12923] (Ca— 27lb10, S, W-—-12h8134); Tso 20h07 (N, N), 21075 (N, N); Tsui 395 (Ba, Bu—17565, N, W—1754642), 150 (N), 601, in part (Ba, Bi, Bz-~17566, N), 666 (N, W—175)800). Shantung: Chiao 2787 (N, ¥— 1596247), 2916 (N, N); Zimmermann 210 (Bz—-17655), 465 (Bz— 17654). Province undetermined: Fortune A.96 (K); Hu 492 (Ca— 246815). CHINESE COASTAL ISLANDS: Honam: C. 0. Levine sen. [Herb- Canton Chr. Coll. 1197] (Ka--62899, W--1010299, W--1173183) - INDOCHINA: Tonkin: Du Pasquier 3039 (Ca-— 1896 (W—22)7891); Collector undetermined sen. [Musachi] (W— 1133078), s.n. [Musashi, Oct. 22, 1893] (W—20618); Furuse s.n- (20 Oct. 1955] (s), s.n. [23 Oct. 1956] (S), sen. [17 Aug. 1959] ( 380 (B, Ca--955796, Go, Mg, Mi, N, Vi, W—22)215h), s.n. [Hondo, Jul. 13, 1951] (S); EB. H. Walker 5712 (W—206)953). Kiushiu: Herb. + Bot. Stockholm s.n, []/9/1913] (S). Island undetermined: Bridges s.n. (K); Herb. Lugd. Batav. s.n. [Japonia] (S); H. L. 2)3 Mi s.n. [Bizen, Oct- Vasey s.n. [Agricultural Grounds] (W—5613h); Ward s.n. [Agricul- tural Grounds] (W—-2)5 1966 Moldenke, Monograph of Callicarpa 169 Florida: Fennell 61 [U. S. Pl. Introd. 102922] (Qa--9183), 70 [U. S. Pl. Introd. 102922] (Oa—-9179). Germany: Her 4 Hort. Ber- a son. (1854) (B, S). India: Herb. Griffith 6043 (B); Herb. _ ;. Bot. Calcutt. son. (B, Bz--17661, Ed, Ed, Mu--988, Mu--1161, 7, tmri Voigt s.n. [H. B. Seramp.] (Cp, Cp, "op); Wallich Wallich 523 (Cp), 1828 (B, B, B, Ed), s.n. (Cp). Java: Herb. Hort. Bot. Bogor. XV. KA.XIV.6 (Ba-—26)31, Bz——26)32, Bz, Bz, Bz, Bz, Bz). ae F. C. Gates 11533 (Ka—71i9h, Um—821), 14548 (Ka—71506), 11963 (Bt- 12678, » B--10798%6, Ka--71391, Ob—50626, Vi), 1699 , See ey: Maryland: Coville s.n, [Garden of Whitman Cross, ‘3, Chevy Chase, Oct. 29, 1926] (Ar—-19787). Massachusetts: Blazic s.n. [July 21, 1922] (Gg—31979); Herb. Arnold Arb. 1088-2 (B), s.n. (Sept. 30, 1913] (Gg—31985); Kidder s.n. [21 Aug. 1925] (Oa-—-107K6); Leavitt s.n. Cambridge, Oct. 13, 1698] (Oa); E. J. Palmer s.n. [Herb. Ar Arnold Arb. 10151; seed of E. H. Wilson 9348) (Ur); Rehder sen. [1088-1] (Ur), s.n. [Arnold Arb., 10.IxX.1898] (Ur), s.n. [Arnold Arb., 21. 1.1898] (Ur, Ur), sen. (Sept. 28, 1915] (Ur), sen. (Aug. 2) (5), Sin. [Oct. 31] (B), sen. [Arnold Arboretum] (E—119313, Mi—-photo) ; RE E. Torrey s.n. [Amherst, Sept. 12, 192] (lis); A. P. mymans $: Se n. [July 23; Arn. Arb. 1088] (o-31570), sen. (Oct. 75 Arn. Arb. 119263); Shaw School of Botany s.n. eteiee oo “New erin H. FS Ne Moldenke 2689 (N), 9315 (Ba, Br), 19138 (N, nat 20130 (Sm). New York: K Eee R. Boynton 8.: s.n. [Bronx Park, ark, July 8 2h] (Ca 513050); W. W. Denslow s.n. [July 186k] (us), sun. PoaBe: 1865] (Ms); Quinn S.n, [6-10-38] (N)3; Tidestrom 085 (Ar—-19791). ages Carolina: | Biltmore Herb. 4189 [August 2nd, ao lech ee — 26 702009, vt, W— se RS oy Boe ai Bn rear a Ape Se Herb. "tac 71506) (al, B, Ur, Vt, Vt, We); Le Tate so. [Chapel Hill, #47, 1938) (Hi—3167) 5 Lindley s S.n. n. (Greensboro, Oct. sets ” (ii——£91217) 5 ; = C. Mathews Mathews s.n. (Chapel Hil Hill, October 5, 1938] (Hi—10083) ; Packard & d & Harrison s.n. [Biltmore Estate, Sept. 13, 1899] (Ar— 19790), san. [Sept. 20, 1899] (1o—29668); P. 0. Schallert sn. (Winston-Salem, 7/25/26] (Vt), sn. [7/10/27] (Bw), 3- seca (Winston Salem, 6/20/3k] (V). Ohio: Rob b. Burton sn. [October 19h6] (Ws). Pennsy1 eee MacElwee s.n. [West Philadelphia] (D—514126); Mar—- tindale S.n. [Meehan's Nurseries, July 1876] (Pr); Porter s-n- 1 [La- fayette Coll Satteed, Easton] (F—317329); W. H. Rhoades s.n. ; sen. (North Wales, August 193h] (Bt--29653, Bt—Wi906, Hs); Westerfeld 2502 (We). Russia: Virchou sen. [Batoum, 10 Oct. 1811] (B)- Virginia: Towne s.n. (October 1695] (No--256Ws). LOCALITY OF COLLECTION UN- 170 PHYTOLOGIA Vol. 1h, no. 3 DETERMINED: Muenscher & Burkholder s.n. (Ca—-5130)9). CALLICARPA DICHOTOMA f. ALBIFRUCTA Moldenke, Phytologia 7: 429. 1 Bibli ography: Moldenke, Phytologia 7: 429. 1961; Moldenke, Bi- ol. Abstr. 36: 036. i rene Résumé Suppl. 3: 27. 19625 Hocking, Excerpt. Bot. A.5: differs from the sei form of the species in haw ing white fruits. The type of the form was collected by M. Togasi (no. 1667) from plants in cultivation at Settsu, Yamamoto, Honshiu, “Japan, on September 21, 1957, and is deposited in the herbarium of the Na- turhistoriska Riksmuseum at Stockholm. etops 2 3 herbarium speci- peg | been exami luding type, tations: CULTIVATED: Japan: Togasi 1667 Chanctnotchiits S—tyPe, 42335139 levies) CALLICARPA DICHOTOMA var. SINUATO-DENTATA Dop, Bull. Soc. Hist. Nat. Toulouse 6): 507. 1932. Bibliography: P. Dop, = Soc. Hist. Nat. Toulouse 6: 507. 19323 Moldenke, Geogr. Distrib. Verbenac., ed. 1, 59 & 1942) and ed. 2, 135 & 177. 1949; Moldenke, Résumé 175 & h3- s variety differs from the typical form of the species in having its oa gers mors ig cages and sinuate-dentate along of ae type of the argon was collected by Benedict Balansa (no- 926) at Ouonbi, Tonkin, Indochina. Dop (1932) says: = 1' absence de fleurs je fais de ostas plante une simple variété. ne connais- sance plus compléte de ses caractéres permettra eathtne 2 de 1'éle- ver au rang d'espéce". I know nothing of this taxon besides what is stated in the original description. CALLICARPA DOLICHOPHYLLA Merr., Philip. Journ. Sci. Bot. 7: 339— 30. A912. Synonymy: Callic caudata var. glabriuscula H. J. Lam, Ver~ benac. Malay. Arch. a Pde Bibliography: Schau. in A. 645. 18473 E- De Merr., Philip. Journ. Sci. ig i 330 (1912) and bos 108 & 301. 1917; H. J. Lam, Verbenac, Malay. Arch. )7, 50, 6 Sie eae ag 1919; Prain, Ind. Kew. Suppl. 5: )3. 1921; Bakh., _—_* Prelim. Alph Moldenke, Alph. List Invalid Names 8 & 10. 192; Moldenke Geogr. Di erbenac., ed. ts 62 & 82 (a9k28 and ed. 2, 177. 1995 Moldenke, Résumé 182,22, & lh3. 19595 i Moldenke, Phy- tologia 1h: 12. elas rous or the A ona leaves, especially 5) oath 1966 Moldenke, Monograph of Callicarpa 171 stems 2—}3 cm. in diameter; branches terete or somewhat compres- sed, brown, smooth, glabrous or somewhat pubescent when young; branchlets somewhat hairy with simple hairs or subglabrous; leaves decussate-opposite; petioles 0.5--2.5 cm. long; leaf—blades chartaceous, lanceolate, 12—30 cm. long, 2.5--7 om. wide, acum- inate or rather slenderly acuminate at the apex, subentire or crenate- to crenmulate-denticulate along the margins, acute or a- simple hairs on ture, lower surface more densely » mostly entir or sub- us on one or both surfaces when mature, the lower surface distinctly yellow the glands in minute pits; seconda . ~glandular, pa ries 13—16, distinct, curved-ascending, anastomosing, the retic- than the petioles; calyx cupuliform or som bular, 1— 1.5 mm. long, glabrous, its rim l-denticulate, the teeth short, obscure; corolla somewhat exserted, violet or reddish, 2.5--3 m. white when mature, containing l pyrenes. The type of this species was collected by Hugh Cuming (no. thors to regard Schauer's trinomial to belong, in part, at least, in th Howe uer E Ie : F H oe designation for the typical form of C. longifolia Lam., I feel that it should be placed only synonymy of that species. His citation of Cuming's plant was only a misidentification on his part. Merrill (1912) cites, in addition, a Ramos s.n. [Herb. Philip. Bur. Sci, 8268] from Luzon, in the Philippine Byreau of Science Herbarium, now destroyed. He comments that Schauer's description particularly well 172 PHYTOLOGIA Vol. 14, no. 3 the M. Ramos 2037 collection cited below, and the two species are certainly closely related. Lam (1919) says of C. dolichophylla: "It has the habit of C. longifolia, but is essentially different by its simple hairs" and "folia quam in var. > (=C. caudata Max- im.] angustiora". Bakhuizen van den Brink (1921) placed it in the synonymy of C. longifolia Lam. with a question. Merrill (1923) comments that "This is remote from Callicarpa longifo olia Lam., where Bakhuizen has erroneously placed it as a " SN a C. longifolia var. subglabra Schau., C. longifolia var. subglabra- ta Schau., and Cc. Puan ¥ var. typica f. genuina Bakh. In all, 9 herbarium specimens, including type material of both names involved, and one mounted photograph have been examined by me. fede WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Lasont Cuming 1330 (U—isotype, V--isotype, Z--photo of isotype); M. Ra- mos any ee 7 (Bz—18533, N), s.n. (Herb. Philip. Bur. Sci. 1063] (N)5 Ramos Ss & Edafio s.n. [Herb. Philip. Bur. Sci. 16779] (B, Bz-—17570, Ca—309301, N). CARPA ELEGANS Hayek in Fedde, Repert. Spec. Nov. 2: 88. rid Synonymy: Callicarpa serrulata Zipp. ex H. J. Lam, Verbenac 1 bliography: Hayek in Fedde, Repert. Spec. Nov. 2: 88. 19063 E. D. Me - Sci. Bot. 7: 338. 1912; Prain, Ind. 3 Kew. Suppl. 4: 3h. 1913; Fedde, Repert. Spec. Nov. Gesamtverz. 58- 1914; H. J. r - Arch. 50, 73, & 83-—8h. 19195 ‘ . Jard. Bot » ser. 3, 33 25 & 26. 1921; BE. De M Tle, En Flow, Pl. 33 38h. 19233 He.ds Bot. Jahrb. 59: 25. 192k; K » Fl. Micron. 56. Mol" » Alph. Lis ar ag Vern. Names 29, 62, oF. B 86. 779595 Mol~ denke, Phytologia 94. 19h5; Moldenke, Alph, Invalid Names Suppl. 1: 3. 19h75 Yoldenke, Alph. List Cit. 2: es (1948) meh Na 1259. 1949; oe , Known Geogr. Distrib. Verbenac., 142, & 177. 1949; Moldenke, Phytol ogia 3: 140 (1949) and a 2 19523 Moldenke, Résumé 182, 185, 186, 2h2, & h3. 1959; Mold Résumé «3: 21—23. 1962; Voldenke, Phytolo a 1h: 122, nz, 1yl, 162, 163, & 166. 1966. Slende ender bush or very lax shrub, 1—-3 m, tall, or a shrubby vine; stem to 1.6 cm. in diameter ; wood soft, SAPPY» white, odorless, mrtatnens bark smooth, grayish-white; branches arising from below the middle, slender, lax, re wipes teri floc~ cose-tomentose or with sparse stellate ptt when young, ous in age, eglandular, re~branched; ves decussate-opposi short-petiolate, nearly flat, clantiy horizontal; ee pies 1966 Moldenke, Monograph of Callicarpa 173 mm, ant leaf-blades chartaceous (or en aia weap h att: to Hayek and oe ee lanceolate, :.5—1) cm. long, h.5 cm. —— shiny bright-green above duller and paler et acuminate a serps serrate or serrulate along the margins except near the base, at- tenuate into the petiole at the base, glabrous on both surfaces, sparsely glandulose above with shiny golden glands, more densely so beneath, with some stellate hairs on the midrib and some Larger pairs glands it; ondaries )y 3 inflorescence s few-flowered, slender, loose, 1-~3 cm. long ’ 3— 1/2 as as the leaves, nearly glabrous; peduncl lish-blue, 0.5--1.5 cm. long, almost twice as long a 3 rs odorless; vie ok pee of a mish tinge in the blue, 1—1.5 mm. long, t truncate or hardly en seaitet sbeutka blue, pluish-yellow, or Violet-blue to rose, purple, or pinkish-white, its tube 1.5 long, the lobes 1.5 ong, somewhat glandulose; stam ‘ long; filaments blue; anthers yellow or deep-yellow, glandulose on bo 3 styl sh, mm. long; stigma greenish; ovary glandulose; fruit green when immature, blue to purple or violet when ripe, globose, about 3—6 m wide, hanging, very pretty, somewhat depressed above, glabrous; eglandular, subtended by the persistent green fruiting-—calyx. The type of this species was collected by Hugh Cuming (no. 1460) in the province of Camarines Sur, Luzon, Philippine Islands. Hayek, in his original description (1906) says: "Foliorum forma Callicarpae caudatae parse simillima, differt foliorum glabritie et serratura maiore nec non indumento ramorum diversa; a C. pur purea Juss. foliis Perry: nec chartaceis in acumen elongatum says "margi ariter serrata"), tis glandulosi is nitidis nec reis diversa e ae This is first reference that I can recall to any " on the leaves of C. di- 6g alt chotoma (Lour.) K. Koch! Kanehira (1933) misspells Hayek's sur name "Heyek", Mrs, Clemens says of her no. 16800 "probably 4 Shade form of C. formosana; note droop of f inflorescence". Bak- hmizen van den Brink (1921) saat C. Sees, S C. serrulata, C. iongifolia var, brevipes Benth., vipes (Benth.) Hance in the synonymy of what he calls rae: eatin var. dichotoma [=C. Gichotoma (Lour.) K. Koch]. In ny y 1936 work and my y other papers previous - to the eae one I also placed C. serrulata Zipp. rrect. The e reference sometimes appears to there, but this seems given to Just, Bot. scinabie-. 33: 88 (1905) to be erron~ Callicarpa elegans has been found growing in coral reef forests, and forests on thickets near the sea » open forests © mine the Summit of hills, on dry forested baph oely stony-gravelly soil of 17h PHYTOLOGIA Vol. 1h, no. 3 riverbeds, and near rivers, at altitudes of 10 to 665 meters, flowering in February, April, May, July, August, and December and fruiting in February, April, May, September, October, and De- cember, Vernacular ar names recorded for it are "g4rnei", igerruchu", "gerucdut™ » "gorrua "haruei", and tambalabési". The flowers are gulusated as "pinkish-white on Williams 3L0, “bluish-yellow" on Herb. Philip. Bur. Sci. 49011, "blue" on Elmer 1214), "violet-blue" on Herb. Philip. Bur. Sci. 43103, and ‘pur ple" on Takamatsu 1218. Lam (192) says "die Bl&tter werden zusammen mit Betel gekaut" in the Palau Islands. In his i919 work he comments that "Its af- finity is with C. purpurea. Points of difference are the acuminate apex of the leaves » Which is never abrupt, and always serrulate, upper side without any hairs and with some , and corolla- lobes as long as the tube", In this work he cites a Ramos S-N- {abra, Luzon, Feb. 1909} eee eat bel while in his 192 work cites Ledermann 11,358 from Babelthua Ledermann 11065, ihe & 14257a and ae 129 from Pad Under what he calls CG. ca cana var. far. latifolia H. J. Lam am [=C. candicans (Burm, f.) Hochr a) he cites a Volkens 439 from Yap with a question and a note “cf. C» elegans", ~~ Takamatsu 1218 does not appear to be typical; the collector de- scribes it as an abundant shrub in dry fields. Merrill (1912) says: "The species appears to be quite widely distributed in the northern Philippines". He cites, in addition, sen. (Herb. Philip. Forest. Bur. 7018], Hallier s.n., Mem ritt & Darling s.n. [Herb. Philip. Forest. Bur. 1399k], and M. Ra mos s.n. (Herb. Philip. Bur. Sci. 815] from Luzon, and Merritt ritt - — Philip. Forest. Bur. 6725] from Mindoro. aterial has been misidentified and distributed in herbaria un ase the names C, cana var. glabriuscula Lam., C. formosana Rolfe, In all, 43 herbarium specimens and 2 mounted perenne) cluding material of the type collection, have been examined by me- Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Bohol: M. Ramos s.n. (Herb. Philip. Bur. Sci. 43103] (Ca—-2h2blh), sD (Herb. Philip. Bur. Sci. Bas) Hiarsthgihe « Leyte: Edafio $Me (Herb. Philip. Bur. Sci. 167) 7658, Ca—2 3917h). Lugont M. S. Clemens 16800 (Ga-28551) "17073 (Ca--285316); Cuming 1460 (N—-isotype); Edafio s.n. (Herb. Philip. Bur. Sci. 48817] (ca— 321236); Haenke 73 (Ca—280929); Loher 11887 (Ca-—24:3059) , BBs (Montalban, May 1915] (Ca—229197); Otanes s.n. (Herb. Philip. Bur. Sci. jie (N), s.n. (Herb. Philip. Bur. Sci. 3316] (Ca—216187)5 Re Williams 340 (N (N). Mindanao: Ramos & Edafio s.n. (Herb. Philip. es Sei. 9011] (ca--32h035). Panay: " Edatio s.n. (Herb. Philip. Bur. a 42365) (Ut--6305a). Sibuyan: “Elmer 1 12th (Bi, Bz 1966 Moldenke, Monograph of Callicarpa 175 17659, eal N, Ut~-28763, V, Vi, Vt, Z--photo). MARIANA ISLANDS : : Kanehira 1758 (N). Saipan: Kanehira 1026 (N), a6 (N). "Toland Gaiet Sei ed: Haenke 798 (N). PALAU AU ISLANDS: thuap: Takamatsu 1218 (Bi, Ca--995254). Koror: Herre 53 (Bi, w), 6 (Bi, N). Makarakol: Hosokawa 9273 (Bi, Mi). Todaiyama: Kanehira 1886 (N). Yap: Kanehira 120) (Bi, N). Island undeter- mined: Hosokawa 7195 (Bi); Tuyama Sen. [29 Aug. 1937] (Bi). MIC- RONESIA: CAROLINE ISLANDS: Auluptagel: Hosokawa 7454 (Bi, Mi). Corol: Kanehira 248 (Bi, N), 249 (N), 1995 (N), 22 (N). ppepticrs ns ERIOCLONA chat in A. DC., Prodr. ll: 643. 187. ynonymy: Callicarpa cana var. repanda Warb. in Engl., Bot. pre & 13: 426. 1891. | Callicarpa repanda (Warb.) K. —_ Bs Warb., Nobizbl. Bot. Gart. Berlin 2: ly. 1898. panda K. Schum, & Warb. apud Thiselt.-Dyer, Ind. Kew. ss, oe 32. 190h. Callicarpa erioclona var. repanda (Warb.) H. J. Lam, Verbenac. Malay. Arch. 78. 1919. Callicarpa erioclona var. typ- ica H. J. Lam, Verbenac. Malay. Arch. 77. 1919. Callicarpa fu- mata Zipp. ex H. J. Lam, Verbenac. Malay. Arch. 77, in syn. 1919. Callicarpa erioclona var. typica f. typica Bakh., Bull. Jard. Bot. Buitenz., ser. 3, 3: 19. 19. 1921. Callic candida Elm. ex E. D. Merr., Emm. Philip. Flow. oe 3: 304, hyponym. 1923. Cal- licarpa erioclona var. repanda H. J. Lam in Lauterb., Engl. Bot. Jahrb. 59: 89. 192k. Calicarpa soaee Schau, ex Moldenke, Alph. List Invalid Names Suppl. 1: 3, in syn. 1947. Verbena eri~ Oclona Schau. ex Moldenke, Résumé Suppl. 3: 37, in syn. 1962. Callicarpa 1 lagunensis Merr. ex Moldenke, Résumé Suppl. 4: 11, in an re. 1908: Callicarpa erioclona var. typica f. gemuina Bakh., erb ete ae oe Bibliography: Schau. in A. DC., Prodr. 11: 643. 1847; Miq., Fl Ind, Bat. 2: 889. 1856; Benth. & Muell., Fl. Austral. 5: 57. 18703 Scheffer, Ann. Jard. Bot. Buitenz. 1: ll. — Vidal, Revis. Pl. Vasc. Filip, 208. 1886; Warb. in Engl., Bot. Jahrb. 13: 426. 1891; Jacks. in Hook. f. & Jacks., Ind. Kew. se 2 366. 1893; K. Schum. & Warb. in K. Schum., Notizbl. Bot. Gart. Berlin 2: - 1898; K. + Toviou 2 Moldenke in Fedde, Secces — 500, 502, & 512. 1932; ©) Nov. 0: 112. 1936; Moldenke, Alph. List Common Vern. Names V,. 23, 26, & 29-31. 19393 Moldenke, Prelim. Alph. List Invalid 176 PHYTOLOGIA Vol. 1h, no. 3 Names 10 & 12. 1910; Moldenke, pony Geogr. Distrib. faxpanne'ay ed. 1, 59, 62, 6h, 65, 67, 68, & 86. 19425 Moldenke, Alph. List m nvalid Names 8—io. ioles Mold a sorte ogia 2: 5h. 19453 rd , Alph. List : hk, 22h, 25 (196), 2: 03s Ui 3 : 566 (198), ot 5° (a9i9)° and "us ih 5, 1158, 1161, 1226, 1259. 19495 M wi e, Known Geogr. Distrib. Ve got ~ ae 135, 140, 150, & 177. ~ 15195 Bal steke, 5 Bot. 8: 172. tones *Moldenke, Phytologia 3: 293-- bogs & 380 3 (1980) and : 78, 87, & 121--123. 1952; ey pan 114 & 122. and straggling or shrub, 1-10 m. “tall, tree t t or semi-climbing; stems 1.5--6 cm, in di- ameter, qopesiy branched from below the middle; wood rather hard, , odorless, tasteless; bark yellowish, especially be- neath the einottiah epidermis; branchlets very hirsute leaf=blades submembranous or membranous to su subchartaceous, divers in size, deep- or dark-green above, glaucous or yellowis te ng, subabruptly acuminate at the apex, denticulate tebant at the base) or shortly and unequally dentate along the margins, some- times irre ly so, acuminate or subabruptly acuminate to cune- ate or obtuse at the base, the upper surface stellate-hairy when immature, smooth and glabrous when mature, swe wee usu subequ. r sho ding petiole, sometimes longer, lanate-tomentose with ferruginous hairs and also white-farinose; pedicels very short, stellate- tomentose; flowers odorless; calyx exiguous, 1—1.3 mm, long, shorter than corolla, white-stellate-tomentose and -plumose villous its red, vi lobes ok. mn, long, with lines of yellow glands; s 1966 Moldenke, Monograph of Callicarpa 177 long, long-exserted; anthers yellow-brown, densely glandulose; pistil long-exserted; style |, um, long; stigma somewhat incras- sate; ovary villous , glandulo ose; young infructescence erect, green; fruit small, globos se, green when immature, blue or dark-blue to blue-black, pink, red, blood-red, violet, * purple, or black when ripe, sometimes white (G jellerw rup 698 *698). The type of this interesting species was collected by Hugh Cu- ming (no. 911) in Albay province, Luzon, Philippine Islands, with specimens of the type collection cited fy Schauer Salt Bs in the Berlin, Kunth, and Lucae herbaria. The t type of C. da was collected by y Adolph Daniel Edward Elmer (no. 151k) at Tro at Irosin, ount Bulus in the province of Sorsogon, m, Luzon, in November, 1915. The ne of C. lagunensis was gathered by Maximo Ramos (Herb. Reap. Bur. Sci, 23798] at San Antonio, in the province of » Luzon, in October, 1s and is deposited in the United States National Herbarium a Washington. It exhibits large coarse- ly toothed leaves and s spre Sion aijines stems Callicarpa erioclona has been found by collectors growing along streams and forest streams, along the ocean strand, on woode edges of cliffs, in dry fertile soil, rainforests and dense rain- forests, new and sec ondary forests, forested valleys, banana Plantations, woods and forests, open forests, and open wet areas, at altitudes of 10 to 126 me ters, but chie efly at low altitudes, from March to December, and fruiting from April to De- cember, Herre describes it as "common everywhere” in the Palau Islands; Takamatsu calls it "common in dry places in open forests" on Yap. Pancho — it "in open waste places throughout the Philippines", and G calls it "common" on Lele islet. Smitinand describes Ve it are "alo-loy", ta tiptipinagut", nantiptipinagut®, "bal4- *, ong", "palis nN "salingérau", "tambalab4si", tigaot, stipe "tigoo", maitbang-dal gg", Of these Ido not know which is the more aut The fies are described as — on Janowsky 515 ae » "pink" on Kostermans 2681, "violet! on Herb, Philip. Forest. “rose-colored” on Zwickey 1h, ee enta" on H. St. St. John john 21502, "blue" on Herb, Philip. Bur. Sei. 46162, "yellowish- ™m" on Herb. Philip. Bur. as 1.9282, "whitish-brown" on Herb. ick. Bur. Sei. | u931h, “purple” on Glassman 2720 and Smitinand a 1451, "dusty- white” on | Castro & Welegrite 17 rito 171k, and "white" on Agama 10h9, GJellerup 698, Guarrero 7, Hallier L027a, Pancho 1385, Peekel 63, and Vanoverber h 17598. Miquel (1856) wes "Tomento dimorpho insignis". Bakhuizen van 178 PEYTO LOG 1a Vol. 1h, no. 3 den Brink (1921) elaborates this as follows: "Young parts, 1 side of leaves and inflorescence with dense hairs of two sorts, pounded with abuse’ “ bee then strewn on the water of streams. The leaves are employed as a fish-poison also on the Palau Islands according to Kanehira, while on Ponape, according to Burton, this is their finest available fish~poison, somewhat like Derris in its action. Lam (192) says "die zerissenen Blatter werden zum Betfuben von Fischen angewendet." This same author, in his 1919 work, cites Peekel 73), from New Ireland under what he calls C. cana var. typica H. J. Lam with a note "(cf. C, erioclona var. -y)", © this variety being var. repanda- He distinguishes his va: his ad tA of C. erioclona as follows: Var. typica H. J. Lam -- "folia membranacea vel subchartacea preeter basin, i ariter, denticulata; nervis secun- @ 712; 9-26 cM. longa, 3 1/ 9 1/2 cM. Lata; esas 1—2 1/2 cM. —— /2 — kh cM. longi, 21 2 cM. lati, pedunculo 0.7—-1.5 cM. longo, interdum quam petiole longiore." He cites Forsten s.n. (Le—908 —— from Celebesj re eg ee ae Pe a Ye Pe Datel Tee New Guinea, and Parkinson i912 fron jin Oe Var. latifolia H. J. Lam — "folia chartacea, ovata vel oblong? rote te ggg > apice acuminata, margine, praeter basin, grosse dentata dentibus produndis, nervis utrinque 8, 717 cM. longa, h 1/2 — —iu 1/2 cM. lata, petiolo 0.7—3.2 cM. longo." Here he cites DeVries? sen. (Le—908.267-1091, Le—908.267-1092), "Herb. var. botan.” 5: R. (Le—-908.265-110), Le—908.265-1110, Le—908 .265-1hl6), and Forbes 1355 from Java. I regard this taxon to be C. inaequalis Teijam. & Binn., which see. Var. repanda (Warb.) H. J. Lan — "folia subchartacea, vel BOP branacea, late ovata, basi apiceque subabrupte acuminata Spe oe ta, nervis utrinque 10, 22-2 cll. longa, 1213 ol. lat, petiolo 2—3 1/2 cM. longo, crasso (0. cM.); cymi majores 5 oll. longi, 9 cM. lati, pedunculo 1 1/2 — 2 em. long." Here he cites Dahl s.n. from New ew Britain and Peekel 63 from New Ireland. 1966 Moldenke, Monograph of Callicarpa 179 He comments that "The species is well characterized by its of- the f te tomentum, sig and the ‘sae denticulate or d omate margins. Its affinity is with C. cana and C. paucinervia." He gives its general dis tribu- tion as a Celebes, Philippine Islands, New Guinea, New Brit- » and New’ Ireland. Bakhuizen van den Brink Sn! proposes C. erioclona var. typ- ica f. rivularis (Werr.) Bakh, and C. erioclona var. subalbida (Eim.) Bakh., but I regard both these taxa as distinct —— C. rivularis Merr. and C. subalbida Elm., which see. The "N Britain" reference given m by this same catia under C. aie A. L. Juss. doubtless refers to C. erioclona instead. “It is interes- ting to note in this connection that Bentham & Mueller rif say "C. bicolor Juss.......and C. erioclona Schau.....appear t both to be precisely the common Archipelago form of C. Cana” fas candicans (Burm. f.) Hochr.]. It is also worth noting here that the original citations for C. re oa is often given as "K. Schum. et Warb., Fl. Neu~Pomm. Lh, r instance, by Lam (1919). Bekimisen van den Brink (1924) Pg the original publication of C. cana var. repanda as cite La . 59: 89 as "1925" instead of 192h. Schauer *CibuT) and Miaial gees cite a Lesson s.n. from New Guinea, not as yet seen by me; Vidal (1886) cites his nos. Lok from Bataan and 83 from Camarines Sur, Luzon; and Lan - (192h) notes that the branches on Peekel 63, from New Ire land, are "braun- oder schwarz-filzig". He (192) and Kanehire & Hatusima agli give the general distribution of the species as Java, Celebes Guinea, and the Bismark Archipelago. Bakhuizen (192k) gives te aa the Philippines, Celebes, New Guinea, New Britain, and New Ire The Stockholm specimen of Herb. Philip. Bur. Sci. 30258 has been cited as C. candicans, but the University of California spec~ Glassman 2710, Kanehira . 1006, 12h2, & 1301, Pe Nelson 531, and Takamatsu 182 182k, distributed as C. erioclona, are actually var. Paucinervia (Mi (Merr.) Moldenke, while M. S. Clemens 16799, Haenke 81, Loher 12347, Pierre 5227, and Ramos "& Edafio s.n. n. (Herb. Phil- In all, 155 herbariun specimens, including type material of most of the names involved, and one mounted photo examined by me Citations: THAILAND: Smitinand 1355 [Herb. Roy. Forest. Dept. 180 PHYTOLOGIA Vol. 1h, no. 3 7311] (Ss), 1451 (Herb. Roy. Forest. Dept. 7799] (Z}. INDOCHINA: Annam: Pierre s.n, [9/1865] (N). Cambodia: Bejaud 47 (N}. Co- chinchina: Pierre s.n. [9/1865] (B). WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Alabat: Ramos 8 & Edaflo s.n. (Herb. Philip. Bur. Sei. 4831h] (Ca-—~321758). Bohol: R. C. MacGregor s.n, (Herb. hilip. Bur. Sci. 12h] (N); M. Ramos s.n, (Herb. Philip. Bur. Sci. 42770] (Bz—17579, Ca—2h2hh2), son. (Herb. Philip. Bur. Sci. 42853] (Ca—2h2hh3). Catanduanes: RB Ramos & Edafio s.n. [Herb. Philip. Bur. Sei. 75120] (Ca—9190, N), s son. sn. (Herb. Philip. Bur. Sci. 75628] (Ca—hh9228). Coron: G. Lopez s.n s.n. (Herb. Philip. Bur. Sei. 413h2] (Bz--17271); M. Ramos sen. (Herb. mien Sci. 1148) (Bz—1727h). Culion: E. D E. D. Merrill hi (N). seid Q. Ruiz s.n. (Herb. Philip. Forest. Bur. 30005) (Ca——291732). Leyte: Edafio s.n s.n. (Herb, Philip. Bur. Sei. 1685] (Ca~-239201); Elmer 7367 (Bu—17586, N); Kondo & Edaffio 112 [Philip. Nat. Herb. 36778] (Bi); Wenzel 1328 (Oa), 1479 (N, Oa), s.n. [June 21, 1915] (0a), sen. [June 2h, 19: 1915] (Oa). Luzon: Ascuru aru 150 (Gg—-31993); Ahern's Ahern's collector s.n. [Herb, Philip. Forest. . 148) (Bz—~1726h, iS sen. (Herb. Ph Philip. Forest. Bur. 3300] bees 3 I. E. Borden s.n. (Herb. Philip. Forest, Bur. 1595] (N); Cuming 911 (li--isotype, N—isotype); Edafio s.n. (Herb. Philip. Bur. Sci. i. 36333) (Bz— 17578), s.n. [Herb. Philip. Bur. Sei. 8678] (Bz—17575, Ca— 322081, S); Elmer ct (N), 806 (Bz—-17589, N), 15082 (Bz— 17592, Ca—27233h, N, S, Ut--673h3), 1512 (Bz—17269, Ca— 272868, yu, 8, menenests 17598 (Bz—17590, Ca~-271432, N, Ut-- F.C. Gates 6599 (Ka—60447, Ka—66826), 7169 (Mi), 7787 (Ka- 62320) ; Haenke 579 (N); Holman s.n. (Sept. 2h, 24, 1910) ( (Du—668h5 5)5 B.C; Metregee sen. (Herb. Philip. Bur. Sci. 22910] (Bi, Bi); Be D. Merrill 2536 (N); Pancho 1385 (Ba): I. Ramirez s.n. [Herb. Philip. as Bur. 29915] (Ca—268045); M. Ramos 1901 (Bz— 17591), sen. (Herb. Philip. Bur. Sci. 1039] ieee, N), SMe (Herb. Philip. Bur. Sci. 23798] (W—1239180), s.n. [Herb. Philip. Bur. Sci. 32719] (Bz—17593), sen. (Herb. Philip. Bur. Sci. 33179] (NW); Reillo s.n, [Herb. Philip. Bur. Sci. 19265] (N); Robinson & Merritt s.n. [Herb. Philip. Bur. Sci. 612] (N); F. L. Stevens 23 (Ur), 54) (Ur); Tamesis s.n. (Herb. Philip. Forest. Bur. 15363] (Bi); Yenoverberg 17598 (Ca—32h65); H. N. Whitford 487 (N)- Mindanao: M. K. C ik Clemens 271 beri 278) Elmer er 11190 (Bi, Ba— 17267, ree H tlie 4027a (Ca—-918517); Kanehira ae (MN); Quadras 335 a Quezon 1 (Herb. Philip. Forest. Bur. 3025 (Ca—320973); Ramos & Edafio s.n. (Herb. Philip. Bur. Sci. 149013] 1966 Moldenke, Monograph of Callicarpa 181 Sesronyei sb sn. [Herb. Philip. Bur. Sci. 9282] (Bz—~17573, 321), s.n. (Herb. Philip. Bur. Sei. 49314] (Bz=-1757, Ca- pe) Ramos & Pascasio s.n. (Herb. Philip. Bur. Sci. 7) (Ca—215160) ; Wenzel 3389 (Ba—17275, Ca--351975, N)3 R. S. Wil- liams 296) (N, N)3 Zwickey 14 (N). Mindoro: Bermejos s.n. 1. (Herb. Philip. Bur. Sei. 153k) (N); 1 M. Ramos s.n. (Herb. Phi: Philip. Bur Sei. 0933] (Bz—17577). Negros: Usteri sn. [25/1/03] (N). ‘Pal- awan: Fénix s.n. (Herb. Philip. Bur. Sci. 15568] S cata Re. B. Fox - 106- [Philip. Nat. Herb. 1301] (Mi). Panay: [Herb. Philip. Bur. sei. 46162] (Bz—17268, Ca—309637, N i) Es ‘Eepid- osa s.n. (Herb. Philip. Forest. Bur. 301}6] (ca—29192h) Ramos & Edatio s.. Son. (Herb. Philip. Byr. Sci. 31537] (Bz—17265, N). Is- land undetermined: Herb. Philip. Forest. Bur. 9761 [Palis] (Bt, N); milks S.n. cr ~ MARIANA ISLANDS: Guam: & Glassman 81 (Ur). Tini- ? Schubel 28 (Ur). PALAU ee Angaur: H. St. a. sale ee 21502 (8, eae. Koror: Herre 17 (Bi, N). Ponape: R. E. '. Burton 129 (Bi); Hosokawa 58381 (Bi, Mi); Kariyone 31 (Ca—287231); Taka- a 796 (Bi, Ca—805723). Yap: Takamatsu 182h, in part (Ca— 805722). INDONESIA: GREATER SUNDA ISLANDS: Banguey: Castro & Melegrito 673 [field no. 68] (Ca—2h168h), 1714 [field no. 209] (Ca~-2,1306). Celebes: Bish 68 (Bz—17281); C. B. peeeat 2465 (Bz—17282). Java: Zollinger 35.7062 (S). Miangas: H. J. 1 3372, in part (Bz—-17290, Bz, N). British North Sirweey az ama 1049 (351) ms TROIS CARD CAROLINE ISLANDS: Corol: Kanehira 79 (N). 26604, Bz). Mansinam Island: Teijsmann 7788 (Bz—1758), Bz—-17585) . Northeastern New Guinea: Brass 29352 (W—23909)2). BISMARK ARCHI- PELAGO: New Britain: Commerson 8.0. s.n. [Port Praslin] (P); Water- house 290 [Herb. Yale School Forest. 27601] (N). 18% Biol. Abs Abstr, 5 an 1961 B Moldenke, Phytologia 1): 117, the mature leaf-blades varying from rather sparsely to very sparse- ly stellate-pubescent or even glabrescent and resinous-punctate on the ee eee surface, The type of the form was collected by Takahide Hosokawa (no. 8398) at Tadiu, Truk, Caroline Islands, on August 11, 1936, and is oergrm ngeshe rs herbarium of the Bernice P. Bishop Museum in Hono- 182 PHYTOLOGIA Vol. 1h, no. 3 Collectors describe this plant as a shrub, flowering in Janu- Nigatar® and ary and Spi known as gateri”, g ng mo at the seashore. It is used in the treatment of boils on the arms by ue pati vea of the Caroline aoe 24] ote the ground-up young lea are placed in the mouth to s emetic. Material has "been misidentified and distributed . ox hesbarta under the name C. cana L. tr all, 3 este eens specimens, including the type, have been examined ned by m Citations: " ICROMESIA; CAROLINE ISLANDS: Param: Takamatsu 323 (Bi). Truk: Hosokawa 8398 (Bi—type) . CULTIVATED: “Caroline I3- lands: Abbott & Bates 5 (Bi). CALLICARPA ERIOCLONA var. PAUCINERVIA (Merr.) Moldenke, Phytolo- gia 8: 57. 1961. Synonymy : cada paucinervia Merr., Philip. Journ. Sci. Bot. A Peedi as 2 es graphy: - Merr., Philip. Journ. Sci. Bot. 9: 134— 15. S. 19k —. yang r+ Sg Tokyo 32: 136. 1918; H. J. Lam, Ver- « Malay. Arch. 49, 76, 78, & 362. 1919; Prain, Ind. Kew. Suppl . 5: 43. 1921 ; Bakh., Bull. Jard. Bot. Buitens. op BOF> 3, 33 19. eo EB. D. Merr., Boum. Philip. Flow. Pl. 3: 389. aoe Kane er- Steenis Males. umé 12: 8. ae Moldenke, Phytologia 1h: 1,8, 117--120, & 122. 1966. s variety differs from the typical form of the species in its smaller leaves, which usually have only 5 pairs of secon- and z 0 col 9-11 cm. long, l--5.5 om. wide, ro al 1.3 glabrous, the lobes mm. long; stamens exserted; anthers glandulose; fruit dark-blue ws 1966 Moldenke, Monograph of Callicarpa 183 dark-purple to purple-black or black, 2 m. wide, glabrous. The type of the variety was collected in rocky places at Asan, Guam, in Jamary, 1912, and is Guam Exp. Sta. 292. Collectors have found the plant growing on or at the edg ates oF of cliffs, on pure coral outcrops, along the strand, under cocomt trees, ins or clay soil, and in thickets at low altitudes, usually f. rom sea- level Ky Pe mn. altitude, flowering in n January, April, May, and July ber, fruiting in Janu wary, April, and June to S ye Bs “te flowers are described as "lavender" on R. Moran 4525, "lilac" on R, C C. Cowan s.n., “purplish~lavender" on 3. H. Br, ommon dry ar names recorded for the plant "garawow", “hamlat", "hatar", and "memangmah". Kondo ne ret 1 1952) has Ipomoea congesta twining on it. In In regard to P. K. Nelson 531 Merrill says "prob. = cana L." , but with this suggestion I do not agree. This collection is labeled as actually being a topo- inervia. errill (19 says of the plant here under discussion "A spe- cies mch resembling the Philippine Callicarpa erioc 8 ee Points of difference are the number of nerve-pairs the stel- never plumose, texture." Bakhuizen van den aiees (1921) in- cluded C. inervia in the — Cc. 7 thoetae A. L. Juss., but with this disposition of it Ic agree. Material has been misidentified and ae been in herbaria un- der the names C. cana L., C. candicans (Burm. f.) Hochr., and C. | Schau. In | all, 43 herbarium specimens have been exam- Citations: WESTERN P PACIFIC ISLANDS: MARIANA ISLANDS: Aguijan: Kondo VI (Bi), s.n. [June 7, 1952] (Bi). Guam: E, H. Bryan 11,2 (BE), 1218 (Bi, Bi, Ca—995253); H. M. Mayo sen. [Oct. 2h, 1947] Bi); GC. Moore 339 (Ca—7hSiil); R« Koran 4525 (Bi, Ca—51979, M,N); P. Ke Nelson 531 (Bi, Bi, N); Rodin 599 (Ca—789L76); 0. H. Swezey s.n, [June 3, 1936] (Bi), s-n. [Machanao Rota: K Kondo s.n. [June 25, 1952] (Bi). Saipan: Kanehira 2006 (Bi, N). Tinian: R. C. Cowan s.n. [April 3, 1945] (Bi). Is- land unde undetermined: Haenke Sn. . [Anno 1792] (Bi); Hosokawa 7751 (Bi). e yaad ISLANDS: Babelthuap: Takamatsu 1229 (Bi), 1355 (Bi). Ulithi: Lessa 2) (Bi). Yap: Hosokawa 8857 (Bi, Mi); Kanehira 12h2 (N) 5 ceemaates 2 182) (Bi); C. . ¥. Wong 326 (Bi). MICRONESIA: INE ISLANDS: Kusaie: Hosokawa okawa 6190 (Bi); Kanehira 1301 (N). Salas & F. Glassman 2710 (Bi, W). Truk: Hosokawa 8348 ) (Bi); Ce 18h PHYTOLOGIA Vol. 1h, no. 3 C. XY. Wong 135 (Bi). Island undetermined: Maddren s.n. [1899] Du, Du). MELANESIA: BISMARK ARCHIPELAGO: Mussau: Kgie & Olsen 1283 (Cp). Island undetermined: Dissing, Kgie, & Olsen 2556 (Cp, Z): CALLICARP. yg 3 agente Merr. & Chun, Sunyatsenia 5: 178—~179, 0 a a 19h) . iblio nenket wat » Sunyatsenia 5: 178—179, pl. 27. 190 oct g Phytoingie iy 18 1952; Moldenke, Résumé 17) & 1959 ustrations: Merr. & Chun, Sunyatsenia 5: pl. 27. 190. Erect shrub, 0.5—2 m . tall, subglabrous; branches tere glabrous; ultimate branchle ts slender, scarcely 1 m. thick, shortly stellate-pubescent downwards with scattered hairs; inter- and calyx persistent, disciform, 2 m. wide, red-glandulose on the outa surface, its rim hapten: fruit globose, mm. wide, gl@- rous. The type of this species was collected by F. C. How (no. 72515) in a forest at C ha village, at an altitude of 100 feet Po-Ting district, Ma~Tiu Chung, Hainan Island, in anthesis on "May The authors c from Hainan and commen ‘A striking differential character of this species, which mani- festly belongs in the group with Callicarpa dichotoma thurs) Raeusch., are the cert rd shining, red glands of the lower , faces of the * leaves, and to a less degree on the midrib above, ° the petioles, and on the inflorescences." the In all, 3 herbarium specimens and 2 mounted photographs of type collection have been examined by me Citations: CHINESE COASTAL ISLANDS: Hainan: F. C. How 72515 (Bi—isotype, Bz—17596—isotype, N—isotype, N—photo of isotype, Z—photo of isotype). CALLICARPA FASCICULIFLORA Merr., Philip. Journ. Sci. Bot. 17 Liography: E. D. Merr., Philip. seg ci. . 3b. 19205 E. D. Merr., Emm, Philip. Flow. Pl. 3: 38h. ‘1505 he - Hill, Ind. Kew. Suppl. 7: 36. 1929; aati Known G 1966 Moldenke, Monograph of Callicarpa 185 trib. Verbenac., ed. 1, 62 = = sag and ed. 2, 140 & 177. 199; Moldenke, on 182 & “us. A shrub mall tr gros r parts villous and stellate- tomentose, sepeek hy ihe Rrensihepe and petioles; petioles 1--2 om. long; leaf-blades thin-chartaceous, oblong to elliptic, 2h cm. long, 6--12 om. wide, rather slenderly acuminate at the a- gins, acute at the base, olivaceous and sparingly stellate-pubes- cent on the midrib, secondaries, _ veinlet reticulation; secon- mostly borne bLo form, 3 mm. long; calyx apnpicise aids one or ek ahae omentose, its be cylindri at the apex; stamens exserted; anthers 3 mm. long; fruit elobose, about 2.5 mm, wide, glabrous, enclosed by mye fruiting- The type of this species was collected by Maximo Ramos and J. Pascasio [Herb. Philip. Bur. Sci. 35123] in open forests at low altitudes, Bucas Grande, Philippine Islands, on June 9, 1919, and was deposited in the herbarium of the Bureau of Science at Manila, but is destroyed. Merrill (1920) says "This species is read- ily mop elle phate from its congeners by its fascicled, sessile or subsessile flowers, this type of inflorescence being unknown to me in any other species of the knowm to me genus Not is me of this esse except what is stated in its bibliography. CALLICARPA FERRUGINEA Sw., Prodr. 31. 1788. Additional & emended synonymy: Callicarpa foliis lato~lanceols tis serratis subtus scabriusculis, cymis terminalibus axillaribus- we Sw. apud Willd., Linn. Sp. Pl. 1: 620, in syn. 1797. Callicar- - pa maestrensis Urb. in Fedde, Repert. Spec. Nov. 18: 119—120. 22. liography: Sw., Prodr. 31. 1788; J. F. Gmel-, Linn Nat., ed. 13, 2: 1; Willd., Linn. Sp. Pl. 1: 620. 17975 Ind. Occ. 1: 260.°17975 aeusch., Nom. Bot. 37- 17975 Ed oi: 133. 1805; We. Te Ait., Hort. Kew. ed. rr; 1: 2h7. 1810; Poir. in Lam., Méth. Suppl. 2: 32-33. 10115 Roem. & Schult., . Syst. Veg. 3: 95. 1818; Steud., Nom. Prnit) pa 3 rane tae Spreng. in L., pie eo 20 (1 3 . ‘oi Ze bs . 36 - . 26 : 1 aged eg Pl. 1: 343. 1842; Walp-, Repert. St. Cuba 2 (11): cI aTIC “B66 Tapes hound Bot. 5: 709. 1800; Griseb., Fl. Brit. W. Ind. 99. op se lacb., Cat. Pl. Cub. 26. 186 PHYTOLOGIA Vol. ly, no. 3 1866; Sauvalle, Fl. Cub. 113. 1868; 0. A. Reade, Pl. nates 62. 1883; Lefroy, Bull. U. S. Nat. Mus. 25: 97. 1684; W. B. Hemsl., Rep. Scient. Res. Voy. Challenger Bot. 1 (1): ‘aby, a5 G. We . Nat. e AC @ S76 & 656. 1902; Millsp., eiake Cola. a Publ. Bot. 2: 1906; H. B. Small, Bot. Bermad, 8—19. 19035 its St ’ athe Spec. Nov. 18: ‘9-120 (1922) rata 20: tent oi A.W. Hill, Ind. Kew. Suppl. 7: 37. 1929; Moldenke in F epert. Spec. Nov. 39: 298, 300, & 308 (1936) and 0: is, “Sic oe 73, 78, 117, 119--121, "123-126, & 128-131. 1936; Moldenke, List Common Vern. Names 12 & 26. 1939; Moldenke, Geogr. traci Avicenn. 4, 6, & 35. 1939; Moldenke, Suppl. List Common Vern. Names 23. iohd; merenres. Bijan Alph. List Invalid Names 12. 190; a Known — Verbenac., ed. 1, 2h, 25, & 86 Ss 5 « 1949; Moldenke, Known Geogr. Distrib. Verbenac., ed. 2, 4¢s “6, 157, & 177. 19493 Roig y Mesa, Dicc. Bot. 2: 389, 390, & 9%- 1953; Alain in Leén & Alain, Fl. Cuba lh: 30h & 306. 19575 "eek dane ke, R Fn 50, Sh, 213, 245, & hhh. 1959; Moldenke, Résumé Suppl. Ui: + > e A. Cc ik .Ind. Gen. Sp. ° ° Recent collectors describe this plant as a low shrub, 1. 3 - tall, with h-white corollas, large fruit-clusters, and the fruit round, red or magenta to purplish or purple, fl vulets, and o at altitudes of 1000 to 3950 feet. Webster suckin it as "com mon in rather disturbed places on rocky banks covered with moist in Oriente. Hemsley (1885) calls it "a common West Indian species os. el (2883), Le Lefroy Cw), and H. B. Small (1913) alt record C. f but this is based on an obvious misidentification, as regen i i ge ed atiy {ndicate. Their plant is C. americana L., the only species of the genus known from the Bermida Islands. The common name, vhf sarey®, which they record, therefore, belongs to C. americana and not to Ce fer- inea ruginea Y Mateialt nf Ge C. ferruginea has been misidentified and distribu- ted in herbaria under = Tames C. apiculata Urb., C. fulva A- Rich., and C. . L. Juss. On the other hand, the “the C. Wright 430 [1860] Sra y a Grisebach (1866) and distributed a5 S: ferruginea, is actually C. fulva. Acufla 12689 greatly resentles C. fulva A. Rich. Len, Clément, & Nestor 5300 is a mixture ™ 1966 Moldenke, Monograph of Callicarpa 187 Ce. TS Britton & P. Wils. In all, 98 herbarium specimens, including type material of all the names involved, and 29 mounted photographs have been examined Addi tional & emended citations: CUBA: Oriente: Acufia 9868 (Es), 10208 (Es), 12689 (Es, W--1881216), 13327 (Es), 13861 (Es); Alain, Saleen, & Chrysogone A.877 (N); Alain & in & Lépez see ne 2am. (Bm); Britton & Cowell 12755 (W—698)10); Mrs Bi, Clément ément 4377 (Ha, N); Ekman 1787 (N); Leén Len ES te i 172h5 (Ha, N), 21297 (Ha, N); Leén & Clément 20186 (Ha, Ha, N), 23128 {duly 19h7] (N); Leén, Clément, & | & Nestor 5300, in part (Ha), ~ 5b? (Ha); Le6n, Clément, & Roca 1016) (Ha); Leén & Victorin 2076 (Ha, N); Linden. 2088 (P); Lépez Figueiras 2191 (2); Shafer 8075 (W— 696470); G. L. Webster 1015 (Mi). JAMAICA: N. L. Britton 208 (F— Hort Sh 7h); W. Harris 5292 (Us, W--307712)5 Harris & ;& Brit- ton 1055 (F—243)07); W. Hooker s.n. (P); McNab s.n. [Jamaica] (Ed); Perkins 1182 (F-38990); | G. Re Proctor 1011h (N); Swartz 3.0, [Jamaica] ~(Cp--isotype, De--isotype, De, Mi--photo of type, pe, S— isotype); G. L. Webster 5608 (Mi, S). CALLICARPA FLOCCOSA Urb. in Fedde, Repert. Spec. Nov. 20: 345— 92 - 192 Bibliography: Urb. in Fedde, Repert. Spec. Nov. 20: 345-36. 192k; A. W. Hill, Ind. Kew. Suppl. 7: 36. 1929; Moldenke, in Fedde, Repert. Spec. Nov. 39: 303 (1936) and 0: 51—52, 119, & 123. 1936; Moldenke, Brief Course Syst. Bot. Lect. 9: 2+ 19373 Course int — ed. 2, 48. A ge We. I ow me 73 %. 1929; Pei, Ven. China 1 (3): [Verbenac. China) 15 3 » 3h, & 36. 19325 L. H- ailey t Florists Handl. enac. mss. 19 mins, Sci. Bot. 61: 80. 1936; Kanehira, | heir ga net ae ed. 2, 643, fig. 599. 1936; a in - pec. Nov. 39: , 1936) and 40: 93-100, 102, il, 120, 222, & 130. 15385 omdonne Alph. List Common Vern Names 2, 15, 23, | & & 20—2. 1939; Moldenke, Geogr. Distrib. ie t 1939 Prelim. Alph. List Invalid Names 9 LO; 1966 Moldenke, Monograph of Callicarpa 189 611. 1945; E. D. Merr., Chron, Bot. 10: 210. 196; Moldenke, Alph. List Cit. 1: 33, 103, 17h, 198, 22h, 235, & 28h. 1946; Mol- denke, Bol. Soc. Venez. Cienc. Nat. 11: 50. 1917; H. N. & Aw Lif Pp - 2: 35, Ok, 0, & (19b8), 3: 718, 721, 727, 728, 732, 765, 805, 837, B40, 895, & 969 (1949), and : 984, 1011, 1012, 1103, P » 1200- 1202, 122), 1234, 1238, 1259, 1260, 1289, & 1297. 1919; Moldenke, Known Geogr. Distrib. Verbenac., ed. 2, 130, 133-135, 140, 157, & 177. 1949; H. N. & A. L. Moldenke, Anal. Inst. Biol. Mex. 20: 3- 1950; Moldenke, Biol. Abstr. 26: 1471. 1952; Moldenke, Phyto- logia k: 1, 77, 121, 122, & 125 (1952) and : 191. eel Molden- eegeet > Zk e ke, Résumé 168, 172—17) 2, 213 : 959; Mol- nke, R Suppl. (1960), 3: 19°& 21° (1962 ht 11. 1962; Li, Woody Pl. T 818—8 fig. 330. 1963; Mol- 3 e Go ° S. Ecol. Res. Humid Trop. Veg. 3h. 1965 F Moldenke, Phytologia 13: et 475 (1966) and 1y: 46, 48, 53, 57-59, 107, 108, & 111. Illustrations: Blanco, Fl. Filip. pl. 427bis [in color]. 1878-1880; T. Ito, Taiwan Shobukutu Dzusetu [Illustr. Formos. Pl.) pl. 601. 1927; Kanehira, Formos. Trees, ed. 2, fig. 599. 1936; Li, Woody Pl. Taiwan fig. 330. 1963. Recent collectors describe this plant as a bush or shrub, a bushy or low tough shrub, half- shrub, tree or small tree, woody or semi-woody, slender, erect, deciduous, 1--5 m. tall, with yellowish scurfy pubescence; stems 1 or several from the same root—cluster or branched from near the base, to 8 cm. in eter at breast height; pith large, white, tasteless; bark smoo us, conduplicate on the upper green or pale-green sur face and yellow-green beneath, or dark-green above and paler be- neath, the apex urved, the upper sometimes narrow-elliptic, the uppermost sometimes lanceolate {cfr. f. angustata Moldenke] ; Sci. 48766, "Lilac on Elmer 13l41 and Glassman 476, “reddish” on Conklin 186, "pink or purplish" on Ching 1838, "purplish" on Kien- 190 PHYTOLOGIA Vol. 1h, no. 3 holz 327 and Tsiang 2158, "purple and red" on Herb, Canton Chr. Coll. 12008 and Peng, Tak, & Kin 9, "pinkish-purple"” on Chun L67h, "pale-purple" on E. D. Ke Merrill 1 is, "purplish~lavender" on on Mf, ee Clemens 17,00, "purpureo-violascentes" o n Owatari s.n., "purple" on on Ying 347 37, "pinkish" on A, Henry 71 and Keng K Keng K.1021, "pink" on Au 905, Gressitt 721, Keng , K.1052, Tsang 20496 & 20596, Tso 20315, and R. S. Williams — 2577, * » "violet" on Herb. Phil Philip. Bur. Sei. Wa7k5, by 15067, & 4638 146383, an and "blue" on Elmer mer 10985 and Tsui —~tmnis plant has been found growing in wet or dry sandy soil, sandy gravel soil, silt or sandy silt, clay, ordinary highland in forests and prime forests, shady woods, wooded ravines, thickets and Acacia thickets, betel=nut groves, rich woods, river land, on level land, river banks, parang slopes, hillslopes and rocky hillslopes, grassy or wooded hillsides, village commons, steep slopes, forested slopes, on small or grassy hills, rocky sides, or on hillsides ae ter scattered trees, as well as long roadsides and open roadsides, creeks and streamsides, and mountain roads, at ahs rodes of to 2333 meters, flowering in every month of the year, and fruiting i n February and May to De=- cember. Merrill (1926) points out that the seeds are dissemina- ted by birds. Sasakt s.n. (Herb. Govt. Formosa 21435] has its leaves rather and narrow, Mori s.n. [August 10, 1936] has the uppermost leaves lanceo olate, “while La Lau 96 has the upper leaves narrow~ elliptic. These collections may represent f. angustata Moldenke or var. longifolia Suzuki. Elmer 22211 is a small-leaved form. on Hainan Island, Tsang reports it as fairly common or abundant in Kwangtung, and Wilson describes it as common or abundant on Formosa. Bailey (1935) informs us that it is offered to the hor ural twigs are pounded and used as a fish poison; Castillo, Juliano, and Buen also say that it is used as a fish poison, while Merrill states that it is employed to "stupify fish". Chung, Peng, Tak, & Kin report that it is employed in Chinese native medicine. v names recorded for this plant are numerous y inclu- "anadhiu", "anoyop", "anoyot", Natélba", "buraun", "chho- khng-&", "chim mei ip lo hai ngan", "for chai tsai", "horai- mura ", “horai-murasaki", "hérat-murasaki", "kabahaboha", "koodi~itu", = kai t's2" francient mustard seed] , “lo had — "lo hai ngan muk", lo hai ngan shue", "mfig-chiong~kun", nan yang", "palis", hpardiatan, "pe th=c hho~khng-4", "sai hong 1966 Moldenke, Monograph of Callicarpa 191 hun", "sai ip un mat” (=small leaf origin close], eban-0ern Anas "sigbat-ta-caran", "tai-teng-h6ng", "tfagau", "tigau", "tfgau", iefeuueticad™, meigbabéat", , neinbabssts "ts'u-k’ang", "tubang dalag", "tubang-dalag", —dalég", "tubaybési". Of these, I do not know which is the more pies Pi which is probably the Uredo callicarpae Petch recorded by Arthur & Cummins (1936) as infesting this species It should be noted here that the Cc. attenuata of Wallich, re- Bees, to in the synonymy above, is actually a synonym of C. ion- gifolia Lam.; the C. bicolor of Jussieu is a valid species, |, While aes attributed to Schauer is C. candicans (Burn. f. ) Hochr.; and the C. americana of Linnaeus is a valid species, while that at- tributed = "Hort." is C. longifolia Lan., that attributed to Lamarck, to Roxburgh, and to Willdenow is C, americana L., that practi to Loureiro is C. candicans (Burm. f.) Hochr., and that of Sessé & Mocifio is C. pringlei Briq. Bakhuizen van den Brink (1921) reduces C. formosana to synonymy under C, pedunculata R. Br., a species to which it is admittedly formosana, but I feel that they may be worthy of nomenclatural recogniti Maacats. original (1837) description of his Callicarpa ameri cana is as follows: "Callicarpa de America. Tronco ee con 4 Campana, en cuatro par tes redondeadas: una 4 un lado y tres al otro. Estam. cuatro, pags en la base de la corola, casi iguales _ altura, pe pero la ineerdion de los dos, mas alta que la de los cion varia. Ant. larguitas, todas fertiles. Germen de figura de - Estilo mas largo que los estambres. Estigma con dos la- bios. Baya con cuatro huesecillos. = Es un arbolito conocido, de la altura de Dig e's & cinco varas. Las flores son pequefias y en- Carnadas, hojas tienen un page algo ait estas con ie pint outes chiquites, 6 solas y estregadas den a del agua co * emborrachan al pescado, y de ahi le viene aga ag Se ees ee a ee a Rolfe's o: siete description (1882) of C. oe as oe Branches becoming glabrate with age, but when young, well the under side of the leaves, the cymes, pedicels, and calyx, 192 PRETO LOG TS Vol. 14, no. 3 covered with a pale brown tomentum of stellate hairs. Leaves el- liptical, acute or shortly acuminate, serrulate or denticulate, base wounded; 2 — 31/2 in. long, 1 1/2 — 21/2 in. broad; dark brown, sparingly pubescent above, slightly scabrid with age; veins al es branch long Pedicels 1/2 lin. long. Calyx campanulate, scarcely 1/2 lin. long, segments minute. Corolla 1 lin. long, lobes rounded, short- er than tube. Stamens exserted 1 lin., anthers elliptical, notched at each end. Style equalling stamens. Stigma capitate. Fruit globose, scarcely 1 lin. diameter. — Tamsui » Oldham, Nos. 388, 389; Watters, Nos. 23, 82. At the foot of hills on S.W. side of Formosa, Wilford, No. 93. Also from Gregory and Swinhoe. A ve e made into po 8 Lam's description (1919) of C. ovata is: "A shrub, 2 M. high; branchlets, cymes and petioles densely ferruginous stellate- tomentose, glabrescent; leaves opposite » membranous, ovate- elliptic; base subrotundate, apex acutely or obtusely acuminate c cM.; calyx with somewhat unequal lobes; 0.09--0.12 cM. long corolla pink, 0.23—0.25 cM.; tube 0.12—-0.16 cM.; lobes h, 0.1 cM. broad; stamens , 0.27—0.3 cM. long; anthers 0.07 cM.; style 0.50.6 cH., with l-lobed stigma; ovary globose, 0.05 cM.; drupe glabrous, 0.25 cM. Distribution: Mindanao. We did not see any imen of this species, but the description gives rise to the supposition that it may be identical with C. Blancoi." The type is R. S. Williams 2577 from above Darong, in the province of Da- Mindanao, collected at an altitude of 600 feet m (1913) of C. ningpoensis is 45 fokiowss “shrub; branches, petioles and inflorescence thickly vao, southeastern on April 9, 1905. Ma fil e s serted, very slightly exceeding the stamens, dilated at the top; fruit a berry, glabrous, 2 m. in diam. (Chang. no. 103). ae species of the gems are difficult to distinguish from each ee a PHY TOLOGIA Designed to expedite botanical publication Vol. 14 January, 1967 No. 4 CONTENTS REED, C. F., & ROBINSON, H., Some seks jee the Bh cep of Mexico. . . : : ee - ELIAS; T. S., Notes on the genus fuga. 2 os 4 se OS DEGENER, O. & I., Partial review of Doty & Mueller-Dombois’ ‘‘Atlas’’, and new taxa in Hawaiian Rubiaceae. ..... + 213 MOLDENKE, A. L., Book review =. 20 2 es a ee SO MOLDENKE, H. N., Further novelties in the Verbenaceae....- - 216 MOLDENKE, H. N., Additional materials toward a ——— of the genes Cailicarpa. Vie Soe ve Fe ed "Published by Harold N. Moldenke and Alma L. Moldenke SOME BRYOPHYTES FROM THE HIGHLANDS OF MEXICO Clyde F, Reed * and Harold Robinson *+ During the past five years, Dr, Jerzy Rzedowski, egy He Poli- tecnico Nacional, Escuela Nacional de Cieneias Biolo ogica » Mexico, has sent several collections of mosses and liverworts from nities + senior author, st of the collections are from altitudes from 1 to Two new species have been described from these collections: Oreo= Weisia mexicana Robinson (Bryologist, 68: 331, f. 1-5. 1965) and Metz= Reria rzedowskii Robinson, in the present pores Complete sets a as ecimens of these collections are in the Reed at aah ee The localities in Mexico in which these collections were made are — below, along with the proper ecological data and a number, be referred to along with the collector and collector's number for ees species e Contreras, alrededores del 4° Dinamo, ladera 00 m. ? FEDERAL I: Canada andesitica con Sepstavthe de bosque de Abies religiosa, alt, Mar, 28, 1965, FEDERAL II: Desierto de los Leones, bosque de Coniferas, sobre los arboles, Sept. 23. 1964. FEDERAL III: bireconaens de la Estacion La Cinca, Serrania del Ajuseo, laderas basalti con vegetacion de pinar abierto con abun- dantes gramineas, alt, 3000 m, June 13, 1965. FEDERAL IV: Desierto de los Leones, cerca de Venta, bosque de Abies, alt, 2900 m Sept. 23, 1964. nn eetiniositibissetianionmegis ce et a * Reed Herbarium, 10105 Harford Road, Baltimore, Maryland, ** Division of Cryptogams, United States National Museum, Washington. 193 194 PHYTOLOGIA Vol. 14, no. 4 FEDERAL V: Desierto de los nes alrededor del Convento, sobre — las ramas caidas de los calgon bosque de Pinus y Abies, siendo ¢ primero el dominante, May 30, MEXICO I: Bosque de pino~encino, 1 km al W de Cahucan, alt. 2700 & | Mar, 28, 1965. | MEXICO II: Camino Amecameca—Tlamacas, Volean Popocatepetl, sobre las piedras, en lugares my humedos en el gor de Abies, alt. 3000 ™ June 20, 1964. CO III: Laguna de Santiago Tilapa, cerca de Victoria, munici- pio de Santiago Tanguistengo, taludes sombreados de un arroyo cercand, alt. 2600 m Feb. 7, 1965s CO:IV: 3 kmalE ne Se _ municipio Tlalmanalco, verses | W del Iztaccihuatl, sisoey n vegetacion de bosque mesofico 196 Prunus, Garrya, Ilex, pothegg ee y Abies, alt. 2700 m May 21; 5. V: 1 km, al SE de Cahucan, encinar sobre suelo rojo arcil- loso, alt. "2650 m, Sept. 6, 1964. MEXICO VI: Vertiente W del gig 2-6 km al E de San Rafaeh municipio de Tlalmanalco, alt. 2750-3000 m, Feb, 28, 1965. yr MEXICO:VII: Carretera a Sultepec, km, 18 approx., faldas del a de Toluca, bosque de Abies, sustrato arbustivo abundante, may humedo alt. 2500 Me. Aug. 9. 196k, MEXICO VIII: Bosque de encino y pino, 1 km al SE de Villa del Carbon, alt, 2700 m, Mar. 28, 1965, MEXICO IX: Nevado d e Toluca, bosque de Pinus, sobre la tierra sculta, alt, 3500 m. Aug. 14, 1964. MEXICO X: Tlamacos, Volcan Popocatepetl, alt. 4000 m July 1962+ , MEXICO XI: Volcan Popocatepetl, bosque de Abies, sobre los rocé®s alt. 3000 m, June 20, 196k, es aga CO XII: Km 75 de la carretera a Tlamacas, en las faldas del scbuneti, bosque de Abies, alt, 2800 m, Nov. 22, 196k. MEXICO XIII: 2 - al S de Atlapul Chala, pulco, sobre la cerretera 4 orilla de camino, 2800 m Feb. 7, 1965, homie XIV: Cumbre del Cerro Gordo, cerca de San Martin de las ? 3000 mm, july 4, = con vegetacion de bosque de encino, alt. | 1967 Reed & Robinson, Some Bryophytes 195 MEXICO XV: Tlamacas, Popocatepetl, tundra alpina, sobre ceniza voleaniea, alt, 4100 m, June 20, 1964, MEXICO XVI: Vertiente E del Cerro Gordo, cerca de las Piramides, ladera basaltica con vegetacion a metre de staentin: Zaluzania y Mimosa, alt, 2500 m, July 4, 1965. MEXICO XVII: Crater Nevado de Toluca, sobre las rocas, abundante, tundra alpina, alt, 4100 m, Aug, 20, 1962. MEXICO XVIII: Camino la Palma—Basura Veracruz, suelos lateriticos muy humedos, alt, 100 m Mar, 21, 1965 MEXICO XIX: Nevado de Toluca, bosque de Abies de estrato arbustivo my denso, alt, 3300 m. Aug. 20, 1962 MEXICO XX: Carretera Ixtapan de la Sal-Cacahua milpa, a la orilla de la cerretera, alt, 2000 m Oct. 4, 1964. MEXICO XXI: Carretera Mexico-Toluca a la altuca del Posque Nacio— nal de € Salazar, a la orilla de la cerretera, alt, 3000 m Oct. 3, 1964. MEXICO XXII: Nevado de Toluca, reg. de San Pedro Tlamixeo, alt. 3400 ms Oct. 1958, : MEXICO XXIII: Carretera Popo-parck-Ozumba, suelo my humedos, asundante, con esporangios inmaduros, Aug. 1964. bo DURANGO I: Puerto Buenos Aires, cerretera El Salto, Mazatlan, Sque de Coniferas, alt, 2600 m Sept. 21, 1961. O km al NNE de Cuernavaca, sobre la autopista a MORELOS I: ae 7” toe ie Quercus y Abies con Styrax y Ternstroemia, alt. 2150 m MORELOS II: Lagunas de Zempoala, bosque de Pinus y Abies, alt. 2900 m. Aug. 5, 1962, sé I: 10 km al ESE de Cuajomulco, km 59 supercarretera eve ladera basaltica con vegetacion de bosque mesofilo con Guaesrmaraca Abies, Meliosma, Styrax, alt. 2150 m, May 25, 1965. “st VERACRUZ I: 10 km antes de Huatsusco, carretera Huatusco—Cordoba, “que de Liquidambar, my humedo, alt. 1450 m. Oct. ll, 1 a dover ad Z II: 12 km al E de Cordoba, cerca de setede ee cafetal dhe mbra de bosque secundario en ladera caliza, alt. 650 m Oct. 12, ne ra 196 PHYTOLOGIA Vol. 14, no. & VERACRUZ III: Cerca del Punta sobre al Rio Jamapa entre Huatusco y Coscomatepec, sobre las liedras a la orilla del Rio, alt. 14,00 m Oct. 11, 1964. VERACRUZ IV: Alrededores de Orizaba, sobre la corteza de los ar= boles, clima templado frio, a las faldas del Volcan Pico de Oribaza. May 5, 1964. VERACRUZ V: En los lugares my humedos, sobre la cotteza de wee arboles, alrededores de Jalapa, Salto del Gato, alt. 1400 m, Sept. 19, 1964. OAXACA I: Cerce de Santa Rosa, municipio de Juquila, km 195 pont retera Oaxaca—Puerto Escondido, ladera granitica con vegetacion de que seco de pino y encino con Clethra y Saurauia, alt. 1300 m. Apr. 13, 1965. OAXACA II: 10 km al N de Lachao, municipio de Juguila, km 170 — carretera Oaxaca—Puerto Escondido, ladera con vegetacion de bosque sé de pino y encino, alt. 1800 m Apr. 15, 1965, GUERRERO I: Cerro Teotepec, municipio de Tlacotepec, rocas grani- tieas cubiertas de msgos, alt. 3200 m Aug. 10, 1964. GUERRERO II: Cerro Teotepec, municipio de Tlacotepec, bosque de Pinus y Abies con abundantes gramineas, alt. 3200 m, Aug. 10, 1964. GUERRERO III: Aserradera Agua Fria, municipio de Tlacotepec, bos que de Quercus, Abies y Clethra, alt, 2600 m, Aug. 11, 1964 HEPATICAE LOPHOCOLEACEAE 1. Lophocolea bidentata (Linn,) Dum, — FEDERAL I: Rzedowski 19467. JUNGERMANNIACEAE 2. Jungermannia Linguifolia Gottsche ~~ FEDERAL I: Rzedowski 19458. PORELLACEAE 3. Porella arborea (Taylor) Spruce — MORELOS I: Rzedowski 18550. 1967 Reed & Robinson, Some Bryophytes 197 LEJEUNEACEAE 4. Ceratolejeunce flagelliformis (Steph.) Fulford — VERACRUZ I: Ma. E. Sanchez R. 71. PELLIACEAE 5e Noteroclads contiuens Tayl. ex Hook, et Wils. -—- FEDERAL I: Rzedow~ 19460; MEXICO I: Roberto Cruz C, 189. METGERIACEAE 6, Metzgeria conjugata Lindenb, —~ MEXICO II: Ma, B. Sanchez R. Te Metzgeria rzedowskii H, Robinson, n. spe Longissima, pallide, virens, Frons ad 10 em, longa, pauci- ramosa, Costa valida, subteres, Cellulae costae anticae 3-8 plerum- que é-seriatac ; postdcae 4~11, plerumque 6~-7-seriatae denseque setu- 1 € valde decurvae, subinvolutae, in facie nudae, mar s dense dotuinats, setuli mar aia oppositus, Cellulae frondis 50-75 x 40-50 m, trigonis nullis, HOLOTYPUS: Mexico, Dist, Federal; : ada de Contreras, alrededores del 4° Dinamo, ladera andesitica con vegetacion de ——- de Abies religiosa, alt, 3100 m, Mar. 28, 1965. eegeuskd 19476, I SOTYPUS in REED HERB.; PARATYPUS: Mexico, 3 km al alma ilo Garrya, Ilex, Quercus, Pinus y Abi 2700 me May 27, 1965« ¥ ies, o Reedowsid 19% 19869- HOLOTYPUS in U.S. livia. Metzgeria reedowskil is very close to M. gigantea Steph. of Bo Both have large fronds with hairs on the margin and the undersurface of the costa both, t 6 * he fully develo costa has 6 or more Cells on each surface and S07 bf ells, he Mexican materi~ - gowever, the ma 1 cells are distinctly paired on the ’ © th tea vale more clustered and not complet a. t ginal hairs. The hairs of M. gigantea are gen aig Sinuous and thinner eng 15 m wide), The hairs of M. rzedowskii ae Mostly wide and are strongly curved, giving the plant the e on the ventral surface of the costa, short, deeply cup-shaped ser ttn i roW essentially entire margined slit on the upper side, and co sally 3 or 4 antheridia, 198 PHYTOLOGIA Vol. 1h, no. MARCHANTIACEAE 8, Dumortiera hirsuta (Sw.) Nees — MEXICO I: Roberto Cruz C. 191; OAXACA I: Rzedowski 19585, 9. Marchantia domingensis Lehm, et Lindenb, — FEDERAL I: Rzedowski 19484; VERACRUZ I: Ma, E. Sanchez R. 10, Marchantia paleacea Bertol, —- MEXICO I: Roberto Cruz C. 191; MEXICO III: Rzedowski 19422, 11. Marchantia polymorpha Linn, —= MEXICO I: Roberto Cruz C. 195; MEXICO III: Rzedowski 19882, REBOULIACEAE 12, Asterella elegans (Spreng.) Trevi MEXICO V: Rzedowski 18738. ~ 18737, 18750; MORELOS I: siesdatrele4 clea 13. Reboulia hemi schacrica (Linn,) Raddi — MEXICO III: Rzedowski ~ 19421, m Campylium hispidulum, TARGIONIACEAE 14. Targionia hypophylla Linn, — MEXICO VI: Roberto Cruz Co. 125. ANTHOCEROTACEAE 15. Anthoceros laevis Linn MEXICO III: Rzedowski 19421A, con asec toren (Hedw. ) Schimp. 16, Anthoceros punctatus Linn, — FEDERAL I: Rzedowski 19473. 17. Notothylas orbicularis (Schweinitz) Sull. — VERACRUZ I: Ma. 2. Sanchez R, MUSCI FISSIDENTACEAE 1. Fissidens asplenioides Hedw, — MEXICO III: Rzedowski 19420. 1967 Reed & Robinson, Some Bryophytes 199 DITRICHACEAE 2. Ceratodon purpureus (Hedw.) Brid, ——- MEXICO VII: Giovanini y Sau- geda, con Bryum argenteum, 3. Seratodon stenocarpus B.S S.G, -— MEXICO VIII: R, Cruz g. 172; MEXICO IX: Ma, E. Sanchez; MEXICO X: G, Guzman; GUERRERO I: Rzedowski 589. DICRANACEAE 4. Dicranum sumichrasti Duby -- GUERRERO II: Rzedowski 1858). 5. Oreoweisia mexicana H. Robinson — GUERRERO I: Rzedowski 18589 (HOLOTYPUS = US; dupl. in RE=D HERB. and Rz edowski HERB, 6. Smleoharis helicophylla Mont. — FEDERAL I: Rzedowski 19478; : Rzedowski isn, MEXICO IV: Rzedowski 19860; MEXI vis ou ion GC. 117, 125A, 96; MEXICO XI: Ma, E, Sanchez R.; MEXICO II: Ma, E, Sanchez R.; FEDERAL V: Salgado y Villegas; DURANGO I: G.Guzman; MORELOS II: Rzedowski; GUERRERO Il: Rzedow ski 1858,A, LEUCOBRYACEAE 7s Octoblepharum albidum Hedw, —~ VERACRUZ II: Rugdowski 19095. POTTIACEAE 8. Barbula bescherellei Sauerb. — MEXICO XIII: Rzedowski 19412; Ma, E, Sanchez R, 9» Bryoerythrophyllum recurvirostrum (Hedw.) Chen, — FEDERAL I: Rzedowski 19179~ 10, Didymodon campylocarpus (C.MUL1.) Broth. — MEXICO VI: Rzedowskd 18741 y 1874). Tie Stephila involuta ouket Jaege & pars (He yertala (Schwaegr. ) lampe), — VERACRUZ III: Ma, E, Sanch: a ae (C.MAl1,) Mitt. — OAXACA II: Rzedowski + Leptotontium wlocalyx (CMe ) Mitt, — MEXICO II: Ma, E, Sanchez Re; Giovanini y Saucedo. MEXICO VII: Gio 200 PHYTOLOGIA Vol. 1h, no. 4 14, Morinia ehrenbergiana (C.Mi11.) Ther. — FEDERAL I: Rzedowski 194,69. 15. Rhexophyllum subni (Mitt.) Ther, et Hilp. — MEXICO XIV: Pectocant gaye ooo 16, Tortula andicola Mont. —- MEXICO XV: Ma, E. Sanchez R. 17, Tortula caroliniana ae ~- MEXICO I: Roberto Cruz C., con Orthot re pycnophyll 18. Tortula fragilis Tayl., ~- MEXICO XIV: Rzedowski 20059. 19. Tortula ruralis (Hedw.) Smith — MEXICO XVI: Rzedowski 20029. 20, Trichostomum ¢ylindricum (Bruch) C.Mill, — MEXICO V: Rzedowski 18746, GRIMMIACEAE 21. Grimmia fusco—lutea Hook, — MEXICO XVII: R. Cruz C, 22, Grimmia trichophylla Grev, — MEXICO XIV: Rzedowski 20056. FUNARIACEAE 23. Funaria hygrometrica Hedw, — FEDERAL I: Rzedowski 16,80; oe ~ XVIII: Roberto Cruz C, 125; MEXICO XIX: R, Bruz C,; VERACK Ma, E, Sanchez R, 76. er BRYACEAE 2h. filiforme gg Husn, — MEXICO: Rio Frio. May 7; 1962, G.Guzman; MEXICO : Rzedowski 18739, Dera y 18740. 25» Brachymenium spirifolium (C.Mull.) Jaeg, — VERACRUZ IV: David Garcia S, 26, Brachymenium systylium (C.Mull,) Jaeg, —- DURANGO I: G,Guzman. 27. B == argenteum H 5 RE I: Rzedowski 18591; MEXICO XV: Me, E, Sanchez R; » Sanchez R.; MEXICO XX: MA. 3 Ma. E, Sanchez R.; MEXICO Elena Sanchez de Gruz 23 extn XXI: Roberto Crus C, 61. 28. Sryum ca =z capillare Hedw, — MEXICO IX: Ma, E, Sanchez R.; MEXICO VII: Giovanini y Saucedo, lms A eS Saucedo 1967 Reed & Robinson, Some Bryophytes 201 29. Bryum procerum Schimp, in Besch, — DURANGO I: Gaston Guzman; MEXICO XI E, Sanchez R : Ma, E, a 30, Bryum trun — MEXICO V: Rzedowski 18742; MORELOS I: doe apa aie III: Ma, 5, Sanchez R. 96, 31. Pohlia integridens (C.MU11.) Broth, — FEDERAL I: Rzedowski ng con Ano iliforme; M=XICO II: Ma, E, Sanchez R; MEXI Eaedowsid, 18735; GUERRERO I: Rzedowski 18587, 18586, 18588,y MNIACEAE 32, Mnium rostratum Schrad, — MORELOS III: Rzedowski 19830. BARTRAMIACEAE 33. Anacolia intertexta (Schimp.) Jaeg. — MEXICO VI: Ry Cruz CG, 122. 3he Bartramia microstoma Mitt, —~ FEDERAL I: Rzedowski 19483. 35. Bartramia schimperi C.MU11. — FEDERAL I: Rzedowski 19463; MEXICO VII: Giovanini y Saucedo, Breutelia deflexifolia Card, -- FEDERAL I: Rzedowski 19461. 37, Philonotis fontana (Hedw.) Brid, — MEXICO XXII: G,Guaman. TIMMIACEAE Timmielia gnomela (Bruch & Schimp.) Limpr. — MORELOS III: Rzedow- ski oe iets Ment ICO VI: Roberto Cruz C, Ds, | con Targonia hypo- PTYCHOMITRIACKAE 39. Pt Chomitrium lepidomitrium age ) Sohinpe — FEDERAL I: Rzedowski 19485; FEDERAL V: A, Ma, Salgado y M, Villegas. ORTHOTRICHACEAE M. Macromitrium fragilicuspis Card. — VERACRUZ V: Ag Rebolledo. 202 PHYTOLOGIA Vol. 14, no. 4 Al. Orthotrichum pycnophyllum Schimp, — MEXICO XVI: Rzedowski 20054; MEXICO I: Roberto Cruz C. 176, con Stereodon falcatus y Tortula pabatialans: MEXICO XV: Ma, E, Sanchez R. 42, Schlotheimia rugifolia (Hook,) Schwaegr. — VERACRUZ V: A, Rebolledo. RHACOPILACEAE 43. Rhacopilum tomentosum (Hedw,) Brid, — VERACRUZ I: Ma, E, Sanchez R. 71, con Mnjum rostratum y Mittenothamnium reptans; Ma. E, Sanchez YF, Sanchez 75, HEDWIGIACEAE 44, Braunia secunda (Hook,) B.S.G. — MEXICO XXIII: Rzedowski. 450 Hedwigia ciliata (Hedw.) Beauv, — VERACRUZ III: Ma, E, Sanchez Re 92. LEUCODONTACZAE 46. Leucodon cryptotheca Hampe — FEDZRAL II: Ivonne Rojano. PTEROBRYACEAE 47, Calyptothecium duplicatum ceieners) Broth, —— VERACKUZ V: A, Re- bolledo, con Met Meteorium teres, METEORIACEAE 48, Meteorium teres Mitt. — VERACRUZ V: As Rebolledo. 49. Papilleria deppei (Hornsch.) Jaeg. — VERACRUZ V: Aa Rebolledo. NECKERACEAE 90. Neckera ehrenbergii C.MM21. — MORELOS I: Rzedowski 18498. 1967 Reed & Robinson, Some Bryophytes 203 HYPOPTERYGIACEAE ole Hypopterygium tamariscinum (Hedw.) Brid, — VERACRUZ V: A, Rebolledo. LESKEACEAE d26 Herpet ineurum toccoae (Sull. et Lesq.) Card, — VERACRUZ I: Ma, E. Sanchez R., con con Rhacopilum tomentosum, 53. Leskea angustata Tayl. —- MEXICO XIV: Rzedowski 20055. 54. Lindbergia mexicana (Besch.) Card. —— MEXICO V: Rzedowski 18745. 55. Thuidium delicatulum (Hedw.) Mitt. —- VERACRUZ V: A, Reboliedo. 56, Thuidium philbertii Limpr. — MEXICO II: Ma, E, Sanchez R. AMBLYSTEGIACHAE 37. Campylium hispidulum (ria. -) Jatt — MEXICO III: Rzedowski 19421, con Reboulia hemispha BRACHYTHECIACEAE 58. Brachythecium corbierei Card. — FEDERAL I: Rzedowski 19462; MEXI- CO II: Espinoza Russi. 59 Brachythecium frigidum (C.MAll.) Besch. — FEDZRAL I: Rzedowski 19457 y 19471. 60. Brachythecium plumo ; Ki sum (Hedw.) B.S.G. —- FEDERAL I: Rzedows 194.68, 19472, 19486, 19478 y 19459. ol, B Brachythecium stereopoma (Spruce) Jaeg. —- MEXICO I: R R, Cruz 174. 62. B Bryhnia stokesii (Turn,) H. Robinson — FEDSRAL I: Rzedowski 19475, 1947h y y 19465. $3. Surhynehium riparioides (Hedw.) Rich, — VERACRUZ III: Ma. E. San- ; chez R. 93, THUIDIACEAE F E ; & PHYTOLOGIA Vol. 1h, no. ENTODONTACEAE Entodon aeeevistus (Schimp, ex Besch.) Jaeg. — MORELOS III: a 19836; MORELOS I: Rzedowski 18,99; MEXICO XI: Guzman, n Symblepharos helicophylla. Entodon erythropus Mitt. — FEDERAL III: Rzedowski 19926. Entodon jamesonii (Tayl,) Mitt, —- MORELOS I: Rzedowski 18501. Rozea bourgaeana Besch, : R, Cruz C3; MEXICO IV: Rze- dowski, con Symblepharis eatiecueriet MORELOS II: Rzedowski. PLAGIOTHECIACEAE Pagiothectun denticulatum (Hedw.) B.S.G, — FEDERAL I: Rzedowski 19482 y 19466, SEMATOPHYLLACEAE Sematophyllum caespitosum (Hedw.) Mitt, — FEDERAL IV: Rzedowski 18765, Sematophyllum cuspidatum Mitt, — VERACRUZ III: Ma Sanchez Re9h+ Sematophyllum lindigii (Hampe) Mitt. — MORELOS II: Rzedowski. HYPNACEAE Stereodon falcatus (Schimp, ) Fleisch, — MEXICO XIV: Rzedowski 20058 y 20053; MEXICO I: R, Cruz C,,con Orthotrichum pycnophyllus; MEXICO V: Reedowsk 187133 FEDERAL V: Salgado y Villegas; DURAN- GO I: Gast POLYTRICHACEAE Pogonatum cuspidatum Besch, — MEXICO I: C. 175; GUERRERO I: Rzedowski TE 585, aa : Pogonatum subflexuosum (Lor. ) Broth. — FEDERAL I: Rzedowski i MEXICO XIX: Cruz C; MEXICO V : Rzedowski 18734; GUERRERO i: Rzedowski as GUERRERO I: Rsedowsici 18583. Polytrichum juni: djuniperinum Hedw, — MEXICO II: Ma, E, Sanchez Re NOTES ON THE GENUS INGA Thomas S. Elias!?@ Department of poe phn i rere University ondale, Dllinoi While working towards a revision of the genus Inga (Legumi- hosae), I found seven species and one variety as yet ign Be an adi additional taxon has been acavkieet to sieckete while tenth taxon has been elevated from forma to variety. Inga is a large genus (ca. 400 meron) restricted to the New World tropes, George Bentham (1345) divided the genus into five sections, the sections being based one on the indument of the flowers and the shape of the pit ben ot All the new spy and combinations belong to eg Inga Series Inga which is char- INGA EGLANDULOSA Elias, sp. r Arbor parva, ramili subt soaker: "dense brevi-pilosi. Olicla 3-5 juga, 8-18 cm longa, 48 cm lata, elliptica, rotunda ad basim ilosa Pilosa, ale late ovatae; 1 naalad rhachiais absentiae vel inconspicuae. rminal4 picata i cm longa, singulat t vel in superiori ribus axillis disposita, SAG ot a acteae mee tig 3-5 mm longae, acutae, cee villosae; gemmae sllipsoideae, acutae, dense villosae; pedunculi 1,5-2.5 om longi “ise pilosi; calyx 10-12 mm longus, tubulosus ad sw beampanulatus, se st um dorsifixae, 0,25-0,50 mn 1 pa: 0. ongae; pistillum 3.5. ovariun ans A = longum, bisulcatum, glabrum, stylus p1itormis, Stigma Cap: 1) Present address: Department of Biology, Saint Louis University, Saint Louis, Missouri, 2)I ame H, Mohlenbrock for his specially grateful to Dr. Robert H. assistance and ea Dr, John D oor for reviewing the manuscript. 206 PHYTOLOGIA Vol. 14, no. gumen subterete, 60-80 cm longum, 1.5=-2.0 cm in diametro, sulcatum, dense brevi-pilosum. This species bears a superficial resemblance to Ie codonantte but can be distinguished from the latter species by the non- pedicellate flowers, partially winged rachis, and Pineda corollas The type specimen was collected in Colombia at 950 m where it flowers Bs: Marche This species is known only from the type collection Type: Uribe 94, from Colombia. Specimens cited: Colombia: Valle de Cauca: Ginebra, Uribe uh fi ty ste engi gminen Elias bor 5-10 m alta; rami Seeietek: glabri, parce lenticellatiy ne subteres vel angulati, parce ferrugineo-villosi, parce lenticellati. Foliola aa juga, 8-22 cm longa, h=-11 em lata; late elliptica vel obovata, rotunda ad ba: basim, brevi-acuminata ad acuta, glabrescentia ad parce hirtella supra et subter, sub- nitentia supra, opaca subter, subchartacea, laterales venae 16-36 jugae, parce conspicuae subter; petiolulus 3-) mm longus, any 88 ferrugineo-villosus, nigrescens maturitate; petiolus l-6 c longus, parce villosus, exalatus, parce tumidus ad ppeinns rhachis parce scr eg 11-16 mm lata, oblonga ad elliptic > paree eae aaaensiae supra et subter; stipulae subpersistentes, 8-12 mm longae, cordatae, profunde striatae; glandulae rhachidis 2 mm in diametro, Saepe contortae, sessiles, crassis caxteintnd Inflorescentia spicata, 3-5 cm longa, 2-1 per senskey oes in superioribus axillis disposita, 2-40 floribus; brac persistentes, 6-10 mm longae, lanceolatae vel anguste soa long 16 sessilis, striatus, lobis 1-2 mm longis, anguste ovatis, parce is; corolla 26-30 mm — tubulosa » parce dilata ad apicem, oct lobis 5-7 mm longis, lanceolatis ad e ovatis dense longo-sericeis; tubus aan inc lusus $ filamenta alba $6 oa nga; antherae non visae; pistillum 6-7 em longum; ovarium 3 mm longum, bisulcatum, glabrum, stylus filiformis, stigm capitatum. Legumen tetragonum, 15-70 cm longum, 1.2-1.3 cm latum, sessile, sulcatum marginibus dilatis, valvis conspicuis, 6-10 mm dense villosum. This oe is known only from the type location. The length of flowers and the cordate stipules distinguish this 1967 Elias, Notes on the Gemus Inga 207 new species from other members of Section Inga. ree 5. tocarpa = of the legumese This species is possibly an intermediate between Series Inga and Series Pettachanae The specific Sig was chosen because of the distinctly winged margins of + Type: Little 6278, from Ecuador. Specimens cited: Ecua raldas: San Lorenzo near Rio Nadadero, Little 6278 F nie: US isotype). INGA VALIENSIS Elias, sp. bor; ramuli subteres oat a angulati, dense ferrugine Villosi, Foliola h-6 juga, 6-16 cm longa, 2-7 cm lata, oblonge- ° Pilosa subter praeter dense pilosas venas, subnitentia supra, Space subter, subcoriacea, laterales vense 10-16 jugae, conspicuse subters Bh rasa 1.5-2.20 mm longus, dense villosus; petiolus cm longus, dense villosus, exalatus; rhachis 6-15 cm longa, a ta Caducae, 5 mm longae, ovatae; prpentetre rhachidis usque ad 1 m in diametro, orbiculatae, sessiles cum marginibus angustis. Inflorescentiae racemosae, 1.0-2.5 cm longae, productis tim vel geminatis, terminales vel in superioribus axillis dispositae, 1 10-18 floribus; bracteae caducae, 1-2 mm longas, Cvato-lanceolatae; gemmae i-8 mm longas, globoideae, rotundeae, dense villosaes pedunculi 344) cm longi, dense villosi; (exeludus pedicellus) S10. mm longus, late campanulatus, dense inparibus, 1.5-3.0 mm lo dense villosis la 536 mgis, acutis, dense 22-25 em longa, late tubulosa, dilata ad apicem, sericea, lobis filame 5 pte d rece, ” usque mta virdi~alba, 5-7 cm longa, antherae dors ’ to mee mm longae; pist ium pron fre longum, pte os 2.5~ is Capitatum legumen immaturum, terete, sulcatum, dense ferrugineo- villosun, forma The + cimen was originally determined I. spuria racemaria’ by WECP. Killip. i: ison with the type of I. spuria Torma racemaria showed the two to be distinct taxa. its a ee is easily distinguished from I. ingoides by er 208 P.B:Y..0:2.0-G6-1 i Vol. 14, no. Inga vallensis has been collected at altitudes of 1050- 1100 m. Type: Cuatrecasas 23072, from Colombia. Specimens cited: Colombia: Valle de Cauca: Ansermanuevo, Cuatrecasas 23072 (F holotype, US isotype). beet EXALATA Elias, spe bor 8-12 m alta; rami teretes, glabri; rami tere in basim, acuta vel brevi-acuminata, pilosa supra practi dense pilosa in venis, dense pilosa subter, subnitentia s » obs laterales ferrugineo-villos i He staminun ae antherae non visae; 3 pistillum .55.5 lo ovar 2-3 mm longum, bisulcatum, glabrum, stylus filiformis, "stigma capitatum. Legumen ignotum. The presence of the tiny, eres leaf-rachis glands be easily overlooked. glands along with the unwinged rachis distinguish this tee This species is known only from the type collection. Type: Brenes 363), from Costa Rica. imens cited: sidan trian’ 3634 (F BS sana Rica: Cartago: Jesus de San OREALIS Elias, sp. Arbor 10 m alta; ramli mubbaretas » dense ferrugineo juga, 6-1) cm longa, 265-15 n venas, parce puberulenta vel 1967 Elias, Notes on the Genus Inga 209 villosus; petiolus 2-3 cm longus, de osus, exalatus vel e alatus, ala l\-6 mm lata, unpar; rhachis <6 cm 1 nga, dense villosa, alata, ala 6-12 lata, obovata, ce puberula Supra, tomentosa subter; stipulae caducae 2-3 ae, tae; ndulae rhachidis 2.0-2.5 mm in diametro, orbiculatae, sub- Sessiles, marginibus conspicuise Singulatim vel 2-4 per fasciculmm, in superioribus axillis 1, latae vel elliptico-oblanceolatae, 12 » persistentes, €nse villosae; gemmae |\-12 mm longae, oblanceoideae, rotundea ad apicem, dense villosae; pedunculi 1-) i, dense villosi; lyx 8- lo bulosus, substriatus, dense villosus, Sessilis, lobis 2-3 mn longis, anguste ovatis, dense villosis; corolla 18-22 mm longa, anguste tubulosa, dilata ad apicem, sericea, lobis 2=3 mm longis, angus » sericeis; tubus longae, dorsifixae; pistillum 5-6 em longum, ovarium 2-3 mm longum, bisulcatum, glabrum, stylus filiformis, stigma capitatum. legumen ignotum. This species is known only from the type collection. Inga TES, Tesenbles I. xalapensis in that the bracts are persistent. € 0 ceolate bracts and longer corolla distinguish Ie borealis from I. xalapensis The specific epithet was chosen because this taxon is on the horthern-most range of the Zenus o Type: King 275, from Mexico. cimens cited: Mexico: San Iuis Potosi: 2 miles W of Spe Tilitia, King 4275 (NY holotype, US isotype). Arbor; ramuli valde angulati, ferrugineo-villosi, lenticellati. peiiola 3-1 juga, 8-18 cm longa, h-8 om lata, rotunda vel sub- 210 PHYTOLOGIA Vol. 1h, no. 4 Inflorescentiae spicatae, 2-4 cm longae, pera is s latim vel geminatis, in superioribus axillis dispositae, ees floribus; Snankane qaducee, -2 mm longae, ovatae, dense villosae; calyx 220-225 mm longus, campanulatus » parce villosus, sessilis, lobis 0.5 mm longis , ovatis, parce villosis; corolla 6-8 m longa, as Ake dilate ad apicem, parce sericea; tubus staminum includus; filamenta 2.0-2.5 cm longa, antherae dorsifixae, 002 mm longas 3 pistillum 2.5-3.0 cm longum, plurisulcatum, glabrum, sessile, stylus filiformis, stigma capitatum. Iegumen ignotum,e tiny, sparsely pubescent flowers zeadig distinguish this pe from other species of this serie Type: Buchtein' 3778, from Bolivia. ns cited: Bolivia: Nord Yungas: near Coroico, SO va 8 78 (US holotype, GH isotype). INGA CHOCOENSIS Killip ex ge tele obovata, parce pilosa supra et subter, venae conspicuas subter3 stipulae caducae, -6 mm longae, ovatae, ote vel brevi- acuminatae; glandulae rhachidis 1-2 mm in diametro » orbiculatae» subsessiles, marginibus conspicuis. Inflorescentiae ine eulonten » 12-16 mm longae, productis singulatim vel geminatis, terminales vel in superior ibus axillis dispositae, wee £ flor ibus; bracteae caducae, 3.5-6.0 mm longass lanceolatae vel anguste ovatae, sericeae; gemmae 3-7 mm 10 ONgae y campanu dila: lobis 2-4 mm longis, acutis, pilosis; corolla 1-17 mm longa, tubulosa vel dilata ad apicem, sericea, lobis 2-3 mm longis acutis, sericeis; tubus staminum includ us; filamenta alba, hs longae; pistillum 45 cm longum, ovarium 2.5-3.0 m longum, bisulcatum, glabrum, stylus filiformis, stigma capitatum. 1967 Elias, Notes on the Genus Inga 211 gumen subteres, 8-1), = longum, 1.5-2.0 cm in diametro, sessile, valvis conspicuis, 8-1 mm latis » mMarginibus sulcatis, dense pilosis. This species is easily distinguished from all other species of Series Inga by the branchlet and calyx pubescence. Inga chocoensis has been collected at altitudes of 150m where It occurs intr requently along river banks. Type: Cuatrecasas 16372 16372, from Colombia. Specimens cited: Colombia: Choco: Rio San Juan, Palestina, Curtecasas 16913 (US)e Valle del Cauca: Rio Calima, La Trojita, 5a — holotype, F isotype); Rio Calima, between a 7 casas 21169 ied nn seer or > Cuatrecasae 1763 (03); la de a, Cuatrecasas 16035 (US) 3 3 Rio os, oatredibis 14319 (i. 18)3, between El Aquacate and Quebra la Teeny Ceperrcnoee INGA COCIEENSIS Pittier var. MEGANTHA Elias, vare nov ehh ab var. cocleensis calyx 7-10 mm longus, corolla Nga a Aa! oe. 1520 This variety is different only in the larger flower size. The calyx in var. megantha is 7-10 mm long, while the corolla pastures 15-20 mm Tong. This variety can be distinguished rom I. multijuga by the sulcate legumes. variety extends the range of the species into Colombia. Previously Ie cocleensis was limited to Guatemala, Nicaragua S collections of the variety were made at altitudes of 1700-2100 m Type: Arbelaez & Cuatrecasas 658, from Colombia. es Specimens cited: Colombia: El Valle: between Sevilla and icedonia, Arbelaez & Cuatrecasas eS (US holotype); north of Alban, Dugand Jara ret Trek MINUTULA (Schery) Elias, stat. 8 edulis var gent Rs Se Mees Anne cant Bote Garde 3722096 ies 9 Species may be partially o Mays peter winged. Other variations can be found in the shape rk e calyxe bi Several Colombian and Ecuadorian specimens have had th Tame was il. Seve ee applied to them by Killip; however, = rachis = lished. The small — ’ “hig leaf rac lace this 4 glands, and ially to sig acta winge cis 212 PHYTOLOGIA Vol. 1h, no. b I. edulis. Inga cuatrecasasi, another herbarium name of Killip's but never va published, is synonomous with I. minutula. Type: Allen 968, from Panama. Specimens cited: Guatemala: Escuintla: near Barranco Hondo, tea 63873 (A)+ Car agua, Honduras: Atlantida: vicinity of Tela, Standley 56617 (08)« Costa Rica: Cartago: Tucurrique, Tonduz 12705 Ct S)3 Tres Rios, Leon 1235 Sagih US s Del Toro: Changuinola valley, Dunlap 582 - “a F). Darisn: ee iaity of EL. Pena: Allen 968 (MO h holotypes Fs 0 isotypes). Panama: around Go ne 2696 (NY, F; GH) it Canal Zone: Barro Colorado > Island,” se Bangham 395 (US, A), vicinity GH). Grenada: Annandale, Broadway 3353 (NY); Saint George, Broadway se ne (Gi). 1629 (M0) To ass go: without exact aurea tity B Broadw Colombia: Antioquia: 0S), 2276 (es — Uribe 725 (Sin) 5 seg ie == Pereira, Daniel 3 3860 (COL, F). Choco: Palestina, Cuatrecasas 16919 (08) bank sean ee Rio San Juan, near tae, as Caparrapi. Sebiacierciee 7 7 LoS, “Haught 1268 (0s). NOTRE x de Santander: EI Indio, Cua aa ecasas 13131 (US). e de Cauca: “ eins & around erate Pittier 10 US)3 shady Delgate 317, (US). Tovar Iston 70L8 ( : Ecuador: G t seeeRe itcheock 3Ce BN tus; ea osiaes iy Rio. Quininde, Tittle 6195 (F, US). Brazil: Ceara: : Guaramirangs, Catler B16" (MO, NY, US). INGA SPURIA var. RACEMARTA (Uribe) Elias, state nove SS Uribe, Caldasia 3: Jes ~ 19S Spuria forma racemaria Uribe, ae ott 5282. 19L8- variety differs from vare s in the long- jaliserrgte flowers and the Resa or = leaf -rachis a eene This variety is known own only from Cali, Colombia. Type: Uribe 964, from Colombia. Specimens cited: Colombia: Valle de Cauca: Cali, Uribe 264 (COL holotype, US isotype). Literature Cited Seg Ft ow epee 1845. Notes on Mimoseae. Hooker's Jour PARTIAL REVIEW OF DOTY & MUELLER-DOMBOIS' "ATLAS", AND NEW TAXA IN HAWAIIAN RUBIACEAE Otto & Isa Degener In reading wise good "Atlas for Bioecology. Studies in Hawaii Volcanoes National Park," Univ. of Haw. Bot. Sc. Pap. 2: I-507. (June) 1966, f page 153, as "VASCULAR PLANTS by F. R. Fosberg." Familiar with leading botanical institutions, the present checklist in our opin- ion leaves much to be desired. *Fagerlund and Mitchell in 19h performed the actual spadework; while the so-called "partial re- Vision," appearing 22 years later, hardly revises! In the present "revision", Dr. Fosberg's "sensu Hawaiian authors" or "of Hawaiian authors" placed under a binomial he has iar available during the time he studied at the University of Hawaii ° dec o and, conversely, ar modern works Neal, by In fact, of the binomials so glibly listed as new for the Hawaii- an flora, he might have easily gleaned them by making a few pur- chases in Honolulu bookshops or by a little study in the library On page 186 under "Grevillea robusta A. Cunn.", he cites as syno- uym "Stylurus robusta (A. Cunn.) Deg.", evidently not realizing that this latter binomial was published June 30, 1932 before the genus Grevillea ted a nom. cans. He appa alized that this sheet in the "Flora Hawaiiensis” had been re- ro by one entitled "Grevillea robusta A. Cunn.” on March 15, Just to mention a few his l four genera of Hymenophyllaceae dua: Remennte stan and nine combining Dianella sandwicensis with D. lavarum, uniting the Hawaii species 1. GOULDIA KAALA var. RUSSII (Fosb.) Deg. & Deg. (Syn. G. ter” Mus. Bull. 73 Rinalis var, kaala f. Russii Fosb. in B. P. Bishop 213 214 PHYTOLOGIA Vol. 14, no. 4 49. 1937.) 2.G. OVATA var. HETEROPHYLLA (Fosb.) Deg. & Deg. (Syn. gs term, var. ov. f. heterophylla Fosb, ibid. 52.) 3. G. OVATA - KALAUPAPA (Fosb.) Deg. & Deg. (Syn. G. t. var. o. f. Se You, ibid. 51.) Me J G. OVATA var. LYDGATEI (Fosb.) Deg Deg. (Syn. G. t. var. o. f. Lydga tel F Fosb, ibid. 53.) 5.G. OVATA var. MAKAWAOENS ENSIS (Fosb. ) se: & Deg. sto G. t. var. 0. : makawa0o~ ensis Fosb. ibid. 50.) 6.G. OVATA var. MAUNAHUI (Fosb.) Deg. & Deg. (Syn. G. t. var. o. f. maunahui pea sin ae ) is G. iter var. MEMBRANACEA (Fosb. e) Deg. & Deg. (Syn. G. t. branacea Fosb. foes 53-) 8.G. OVATA var. oS ar oc isee: 3 os Deg. (Syn. G. t. var. o. f. obovata Fosb. in Bull. Torr. Bot. Cl. peas 391. 193. Ee # ns , OVATA var. PETIOLATA (Fosb.) Deg. & Deg. Syn. G. t. ¢ petiolata Fosb. in B. P. Bishop Mus. Bull. 147: 53. 1937. #), B.. G. OVATA var . PUNAULA ULA (Fosb.) ef & Deg. (Syn. G. t. var. o. f. Fosb. ibid. 53.) 11. G. OVATA var. SANTALIFOLIA (Foeb:) Deg. & Deg. (Syn. G. t. var. 0. rt santa lia Fosb. ibid. 51.) 12.cG. ‘ais var. STOREY (Fosb.) Deg. & Deg. (Syn. G. t. var. 0. f. Storeyi Fosb. ibid. 52.) 13. G. OVATA var. SUEHIROAE (Fosb.) Deg. Deg. (Syn. G. +. var. o. f. Suehiroae Fosb. ibid. 52.) 1h. G. OVATA var. WAILAUENSIS (Fosb.) ae & Deg. (Syn. G. t. var. o. f. wailauensis Fosb. kid. = 82. ) 15. * cORDATA var. ACUMINATA (Fosb.) Deg. & Deg. (Syn. G. t. var. cordate £ acuminata Fosb. ibid. 5.) 16. G. CORDATA vax. MOLOKATENSIS (Fosb.) Deg. & Deg. (Syn. G. t. var. cord. f. molokaiensis Fosb. ibid. 5-) 17. G. CORDATA var. NEALAE (Fosb.) Deg. & Deg. (Syn. G. t. vare cord. f. Nealae Fosb. ibid. 45.) 18.G. AXILLARIS var. HAWAIIENSIS (Fosb.) Deg. & | Deg. (Syn, G. Hillebrandii var. hawaiiensis Fosb. ibid. 61.) 19. G. AXILLARTS var. NODOSA (Fosb.) Dee. & car (Syn G. Hillebr. var, nodose. f. eunodosa Fosb. ibid. 61.) 20. G. AUI- 2 MICROPHYLIA + (Syn. G. He var. t. f. mi wylla Fosb, ibid. 60.) a over 60 categories of Dr. Fosberg's putative hybrids may urk specimens % are true forms and even true varieties without He should continue to gather and s study representatives of this genus, Whose 2n varies from approximately 72 to 105, in ad questions of hybridization Ce home owners on the edge of Hawaii Volcanoes National Park, the reviewers intend to publish an additional "partial review" 1967 O. & I. Degener, Partial Review 215 of the "Atlas" after more extended field work and after checking the determinations oe critical voucher specimens cited. *Pagerlund, G. 0. & Mitchell, A. L., A checklist of the PLANTS, HAWAII I NATIONAL P PARK, Kilau ea-Mauna Loa Section, with a discus- sion of the vegetation. Nat. Hist. Bull. 9: 175. 19h. BOOK REVIEW Alma L. Moldenke "Botanical Latin —- his story, Grammar, Syntax, Terminology and Vocabulary" by William T. Stearn, = . "956 pp., illus., ne Nelson & Sons Ltd., Edinburgh, 105 sh., and Hafner lishing Co., New York City Ou, *$16.75. 1966. What a Wonderful service to the field of taxonomic bo tiz, ee ” nm some py the eighteenth century terms have gone changes for "the coining or introduction of new words provides material for the action of linguistic natural selection ort the — Gonciee, necessary and expressive ones pass into use and the Perish." How ps it to be learned by coming generations of botanists when our secondary a no longer teach Latin and aeons ele bs includes ™ms, abbreviations and symbols, Greek derivatio: explai = this t he plant . The price 4 ea : k will circumvent the plea for writing descriptions 5 other €s as is permitted by the paleobotanists and zoologists. 216 PUY TOL 0 o Fk Vol. 1h, no. 4 therefore expect to find this work a highly advantageous substitute for his stock of school day Latin grammars and dictionaries. If my husband had not kept earlier issue of this journal! This delay should be interpreted as further confirmation of the great value of "Botanical Latin". FURTHER NOVELTIES IN THE VERBENACEAE Harold N. Moldenke CITHAREXYLUM PACHYPHYLLUM var. CANESCENS Moldenke, var. nov. Haec varietas a forma typica speciei foliis dense minutissime- que subincano-puberulis recedi ° orm differs from the typical form of the species in hav- ing its leaf-blades and petioles when mature densely subincanous- hair, Se € variety was collected by Hugh H. and C. Iltis, no. 488) in thickets along the road to Yauriviri, at the last point where one can still see Puquio, on more or less cultivated slopes with huge basalt blocks, just above Puquio on the road to Abancay, at an altitude of 3700 meters, » Peru, on Decembe 962, is deposited in the H. N. Moldenke herbarium at Plainfield, New Jersey. The collectors describe the plant as a shrub, - tall, the f: sh-brown or dark in color. LANTANA CHIAPASENSIS Moldenke, sp. nov. Frutex; ramulis dense incano~pubescentibus; petiolis brevibus dense incano-pubescentibus; is foliorum ovatis 5-8 cm. lom ‘ » ad apicem acu- truncatis plerumque : ets subherbaceous, tetragonal, incanous-pubescent with e divergent tul rnodes elongate, |,.5—9 cm. long; leaves de- cussa’ Pposite; petioles sheets ck mm. long aneth hi pubescent; leaf-blades membranous, dark-green above, much lighter =. distinctly ovate, 5-8 cn. long, 2.5--l; cm. wide, acute or attenuate at the apex, ly serrate from apex to base, rounded or subtruncate and often asymmetric at the base , sometimes acuminately prolonged into the ely short- 1967 Moldenke, Further Novelties 217 pubescent above, extremely and uniformity Sy AY over the whole surface beneath; antiengge ® solitar capitate; peduncles rather slender, 3.5--6 cm. long, very densely white-pubescent, the hairs not oe in Urouocas 3 heads flowered, to about 1.5 cm. wins outer involucral teustloke eolate-elliptic, to 8 m. long and 2 m. wide, densely short- pubescent on both surfaces, attenuate at the apex. e of this species was collected by Keith Roe, Eunice Roe, and Scott Mori (no. 999) in cutover secondary growth wood- ra, Clusia rosea, and Psittacanthus schiedearms, in tropical de- edduous vegetation 4 km, southwest of Las Rosas on the road to | Venustiano Carranza, at an altitude By Bo meters (16920! Nj 92 30' W), Chiapas, Mexico, on August 8, 1965, and is deposited the H. N. Moldenke herbarium at ue New Jersey. LIPPIA HYPOLEIA var. OVATIFOLIA Moldenke, var Haec varietas a forma typica speciei laninis ngatidiels dis- tincte meri recedit. fers from the typical form of the species in having i 2 Leaf-blades distinctly and uniformly ovate. The type of the variety was collected by Keith Roe, Eunice ¥), in the highlands of Chiapas, Mexico, on August 10, 1965, and is deposited in the H. N. Moldenke herbarium at Plainfield, 8 New Jersey. The collectors describe the plant as a shrub, 1.5 m tall, with white and yellow flower a TA ROSMARINIFOLIA var . LATIFOLIA Moldenke, var aec varietas a froma typica speciei foliis anguste te ellipticis wme, - 7 m. latis recedit. S variety differs from the typical form of the species in having its leaf-blades narrow-elliptic and to 7 m. wide. The type of the variety was collected by Alban Stewart (no. 3309) on James Island, Galapagos Islands, on December 26, 1905, and is deposited in the: herbarium of the California Academy of Sciences in San Franc LANTANA FRUTILLA var. OBTUSIFOLIA Moldenke, var. nov Haec varietas a forma typica speciei laminis foliorum obtusis vel obtusiusculis recedit The type of this variety was collected by K Roe, and Scott Mori (no. 680) in open disturbed ee thickets in a disturbed semi-deciduous forest with sca Quereus and large ferns, along the cool moist Pacific escarpment 218 PHYTOLOGIA Vol. 1h, no. 4 8 km, south of Quezaltenango on highway 9s » near the hydroelectric plant 1/2 km. north of Zunil, at an altitude of about 2075 m., Que , Guatemala, on July 31, 1965 (14°15' N; 91°30! W), and is deposited in the H. N. Moldenke herbarium at Plainfield, New Jersey. The collectors describe the plant as a shrub, 1.5 m. tall, with a pungent odor and with white flowers. ADDITIONAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS CALLICARPA. VI emt Harold N. Moldenke CALLI' =: & emended bibliography: Sieb., Jaarb. Ko erl., Maatsch. : . Hort. Pays-Bas] 18h): 25. 1844; Lindl Paxt. in Paxt., Flow. Gard. 2: , fig. 221. 1853; Schef- ° tenz. 1: 1. 1876; P. Henderson, Hand. Pl., re 6 Gartenfl 1; Lefroy, Bull. U. S. Nat. Mus. 25: 97. 188); "B.", Gard, Illustr. 9: 323. 1887} J. Matsum., Bot. Mag. Tokyo 3: 115 & 318. 1889; P. Henderson, hne us & 385 b—1. 1911; Bakh. in Lam & Bakh., Bull. Jard. Bot. Buitenz., ser. 3, 3: [1], 3, 9-27, [106]--108, & Addit. [I]. 1921; E- D- Merr., Enum, P' + Flow. Pl. 382--389. 1923; Nakai, Fl. Sylv. Kor. ih: 27—33 & 133, pl. 5—9. 1923; Gamble, Pl. Presid. Madras 1085 & 1091—1092. 192h; Sci. 3 : 1932; Suzuki, Trans. Nat. Hist. Soc. Formos. 25: 130—131. 1935; Sympos Impact Man Humid Trop. Veg. 229- 3 Darlington & Wylie, Chromosome Atl., pr. 2, 323 & 503. 1961; 1967 Moldenke, Monograph of Callicarpa 219 Rep. G. W. nape’ ae 362. 1961; Hanelt, Kulturpfl. 11: oh 1363; Qnisumb., » Sarawak Sympos. Ecol. Res. Humid Trop. at a 1965; ; Harvill, ji Marmecn 67: i 1965; Moldenke, Phytolo- gia 13: 66—506 & 508 * (1966) and 1h: 36--63, 99-~128, & 1)0--192. 19665 Stearn Botan. Latin 26h. 1966. Bean (1951) says of the members of this gems "They love the Sunshine and thrive in good loamy soil. Easily increased by te fruit are borne when several plants are closely associated —- common occurrence however, with fruit-bearing shrubs." Yamamoto (1938) has described a mint, Taitonia callicarpoides Yamamoto, f rom rom Formosa, which he says has much the aspect of a species of Callic carpa, CALLICARPA ACULEOLATA Schau. Additional bibliography: alisb., Ind. Kew. Suppl. 10: 38. 1947; Moldenke, rnytetegts 13s “130-130. 1966. Sager ARPA ACUMINATA H.B.K. pair tional bibliography: Sieb. & Zucc. e~phys. Cl. Ktnig. Baier. Akad. Wiss, Mttnchen -% (3): 156. tenes | Moldenke, Phytologia 13: 66 & 475 (1966) and 142. 1966. Recent collectors describe this junk as a shrub and found it growing in aint ghly disturbed by cutting now mostly secondary growth thickets. The ers are uascetbed as "white" on Roe, Roe, & Mori 1353. Rae to Siebold & Zuccarini (18]6) Hasskarl rl claims that C. acuminata H.B.K. was introduced into Java from Japan! It seems Rost Probable, however, that he is referring here to C. acuminata Tombs» 8 synoRous designation for what we now call Ge mudiflora Additional citations: MEXICO: Hidalgo: H. E.Moore Jr. 3392 (Ws). Quintana Roo: Roe, Roe, & Mori 1339 (Ac), 1353 (RE). Vera —_a! Cruz: Cavender s.n. |Alazam, Dec. 19, 1963] (Ws). NS Sc : 45. dditional b 1938; Moldenke, binlioeraniy: 4 13: 166-16? (1966) goes ie. 1966 ANA L. al. Additional bibliography: Thunb., Nov. Act. Reg. ‘ k: 31, 1783; P. Rae Handb. Pl., ed. 1, si. “i381; 2 = Pl. Bermud. 62. 1883; ede Bull. ee" § 188h5"P. Hienderson, Handb, Pl-, ed 2 zo aio) and ed. Ae Oe ez peners are anid Se endo? fling in ane . 220 PHYTOLOGIA Vol. 1h, no. & Small (1913), but = "C, americana L." of Thunberg (1783) is really C. japonica e Additional etieisina : LOCALITY OF COLLECTION UNDETERMINED: Herb. F. J. YX son. [ex N. “Americal (Ws). CALLICARPA ANGUSTA Schau. Additional bibliography: Moldenke, Phytologia 13: 98--500. Merrill ae aaa in addition, an unnumbered Hallier col- lection from Luz The Ahern 8119, “distributed as C. angusta, is actually C. for mosana Rolfe, while Kollmann s.n. [Java, 1638] is Cc. longifolia CALLICARPA ANGUSTIFOLIA King & Gamble Addi tional bibliography: Fletcher, Kew 1938 412 & 413. 1938; Anon., Kew Bull, Gen. Index say *9e “585 Moldenke, Phytologia 13: 500—SO1. 1966. ALLICARPA ARBOREA Roxb. Additional bibliography: H. N. Ridl., rig Peta mae States 938: » Kew Bull. 7 "1959 Moldenke, a ios 5m1--506 (4966) and aby 15639; Fletcher (1938) places C. villosissima Ridl. in the synonymy of C. arborea. He places C. villosa Roxb. and C. lanata oe es Lam in the synonymy of what he calls C. arborea ¥. var. 7. villosa I regard Lam's name as applying to C. arborea var var. psilocalyx eregee ie cae Moldenke. e W. T. Tsang 2391h, distributed as C. arborea and so cited ad in Phytologia 1h: 39 (1966), is actually C. integerrima CALLICARPA BICOLOR A. L. Juss. Additional bibliography: Mol enke, Taytelocts 1h: Lb6—h9, 108, 11, 112, 1h, 121, 179, 183, & “9, The . Barnes s s.n. (Herb. moa Forest. Bur. 55], distri- buted as 3c. SS bicolor, is actually C. formosana Rolfe. CALLICARPA BODINIERI Léveillé age ne semen & SEE bibliography: Moldenk coer Dis- 167, 168 168, 1 im, 213, i, e —35, : oat ag 2} olden, xe, 8 néouné 2 ‘ss 2 Sod 3, & 19! 95 °. ke, Phytologia 1h:'49—62 & 167. 5 Additional synonymy: Gallicarpa arno arnoldiana Hort. ex Moldenke in Fedde, Repert. Spec. fon _ 88, in syn. 1936. iy Lingnan Sei. Journ. 11: 07. 1967 Moldenke, Monograph of Callicarpa 221 tag L. - & E. Z. Bailey, Hortus ga 670. ere A.W. Hill, id ow. uppl. 9: ie eRe faa Es J. Sali b., Ind. sr ysnrae 10: 3. 19h75 Mol desks, gia 1h: 50--60, é2, 99, “103, 127, & the Baileys (1935) and rom se | (1947) regard C. arnoldiana Hort. as a synonym of C. japonica Thunb., as I also did in my 193% work. It seems to me now, Saver that this name belongs more likely in the synonymy of C. bodinieri var. giraldii, a tax- on with whose introduction into American gardens the Arnold Arbor- etum was so closely associated. The Tsang 850 [Herb. Lingnan Univ. 16349], distributed as this variety, is actually C. formosana Rolfe. CALLICARPA BRACTEATA D onal bibliography: A. W. Hill, Ind. Kew. Suppl. 9: 5. 19385, Moldenke, Phytologia 1: és & 99. 1966. CALLICARPA BREVIPES (Benth.) Hance Addi tional bibliography: Moldenke, Phytologia 14: 55, 58, 99— 10h, 106, 147, & 148. 1966. CARPA BREVIPES f. ANNAMENSIS Moldenke ¢fidtitional bibliography: Moldenke, Phytologia 14: 102 & 103. CARPA BREVIPETIOLATA Merr. Additional bibliography: Moldenke, Phytologia 1h: 102, 10L— 106. & 121. 1966, CALLICARPA BUCHERI Moldenke ; Additional bibliography: E. J. Salisb., Ind. Kew. Suppl. 10: 38. 1947; Moldenke, Phytologia lj: 106—107. 1966. Additional reine hi cana var. typica Bakh. ex eal Kew Bull. Misc. inf. ~ 1938: 3: 413. 1938. neal onal bibliography: sin Kew Bull. - Inf. Baath 2 1938; A. W. Hill, ere Suppl. 9: 45S & 46. 19385 Phytologia 1h: 37, L6, ie "59, 107—126, 140, 142, 17h, iE 178. 179, 182, 1283, 91.1966. : The 1569, S thts s.n. [College campus, January 3, 193), and g Sime CE oe distributed as this species, are all actually C. formosana Rolfe. EAA Fg ery . SUMATRANA (Miq.) Moldenke Addi ti nea : er ae Phytologia 1h: 121, 122, "Tee 58, a Piece CALLICARPA CAUDATA Maxim 1h: 140—1h4 & Bea oe bibliography: Moldenke, Phytologia 59, 12, lg, 222 P80: %, 0.46.7 & Vol. 1h, no. 4 The R. S. Williams 1158, distributed as this species, is actu- ally Cc. ~ formosana f. angustata Moldenke. CALLICARPA CRASSINERVIS Urb. Additional synonymy: ca icarpe rugifolia C. Wright ex E. J. Salisb.» Ind. Kew. Suppl. 10: 30, in syn. 19h7. onal biblio, oa < J. Salisb. Ind. X Kew. Suppl. 10: 38. 197; Woldenke, Phytologia ll: 1)--149. CALLICARPA CUNEIFOLIA Britton & P. Additional bibLicgrspty: ekdenke’™ Phytologia 2 ai 155. ag oh The Morton & Alain 9122 cited by me - Rich. much of the aspect of C. cl aap rect as a oent a hybrid sa be tween these two taxa. Seen CALLICARPA Sect Gen?) K. Koch Emended syno: A. L. Juss., Ann. Mf Hist. Nat. Paris 7: ome: C. purpurea Hort. ex Lem., “1859, nor Hort, Bass Meltenke, ere nor sg a 1923, nor Van Houtte, be emended b ibliography: Miq., Cat. Mus. Bot. L 9 Journ ed. Aaeng e Nakai, ri. - Sylv. Kor. lk: 28--30 & 133, pl. 5. 19235 DearS Lingnan Sci. Journ, 11: ),07. 19323 A. W. Hill, Suppl “ MS & Eom Moldenke, Phytologia 1h: 156--170, 173, eit, 18h, Ys purpurea Nakai, refer erred to above, is a synonym of C. japonica oe that ascribed to "Hort. ex Moldenke" is C. longi-~ folia Lam., and that ascribed to "Hort. ex Lem." and to Van Houtte is C. rubella Lindl The Chiao 1612 [Herb. Univ. Nanking 18601], Sawada s.n. [17- VII .1926], and Ta a Takamatsu 22, distributed as C. dichotoma, are all actually C. Miquel (670) ext cites a a Siebold specimens, 3 Btirger Sectnene x2 Specimens, and 1 Maximowicz specimen fram Ja- CARPA ELEGANS Additional Sitdieerari: Ind. er, Suppl. 9: l6. 1938; Moldenke, Phytologia uh! liz. CARPA ERIOCLONA PAUCINERVIA (Merr.) Moldenke 2 a wihitograniis i Oldenke, Phytologia 1: 17h, 179, & Rarer pee FERRUGINEA ioe tional bibli. 1 185-187, eT po ted Moldenke, Phytologia 1): 155, ’ The Alain & Morton 5078 and c. G- Bright 130 koe [1860], distributed as this species, are actually Ce. fulva A 1967 Moldenke, Monograph of Callicarpa 223 CALLICARPA FORMOSANA Rolfe Additional bibliography: Metc., Lingnan Sci, Journ. 11: 07 & 408, 1932; Suzuki, Trans. Nat. Hist. Soc. Formos. 25: 130. 1935; Ee. J. Salisb., Ind. Kew. Suppl. 10: 38. 1947; Moldenke, Phytolo- gla ls: 142, 167, 173, 17h, & 187—192. 1966. Matsuda's original description (1913) of C. ningpoensis con- times as follows: "According to Hemsley this is the case with C. fommosana Rolfe, C. japonica Thunb., C. longifolia Lam., and C. Juss., all of which are closely allied. My plant is al- ed to the species of this group, but as I can not satisfactorily identify it with any of them, I have to propose a new name." Handel-Mazzetti's original description (1922) of C. aspera is: asper m. diam. viridiflava. Prov. Kwangtung. Sept.; Ad vias montis Lunghaotung montium Lungtou-schan, 900 m.?, 8 IX. 1917 Mei1 (Nr. 920). Calycis lobis praeter indumentum ceter- aque insignis." ‘ gi description (1919) of C. blancoi follows: "A shrub with omentose texture on innovations, es and petioles; leaves op- Posite, membranous, ovate or dhl stiguovabe: base acute or somewhat ae to truncate, margins more or less irregularly crena 1 j nerves, not very distinct, 7-9 pairs; 5 1/2 — 9 cM. Ong, 2.25 cM. wide, petioles 0.5—1 cM.; texture varying pper side, from stellate- ry in youth, and glabrous when rag c . Tt thed rous 6 » Ol cM.; corolla glabrous with ) short, glab- 0.26.5 Somewhat hairy, lobes, and 4 lines of little glands, : -3 cM.; stamens exserted, anthers on both sides, 1 are than the epar ene sorences, which, with the different form of the — Z te it from that species." He cites Teijsmann s.n- (Le— 22h PETZOLOGtT Ss Vol. 14, no. 908 .265-307) from Banda with a question, and Lilles 42 and Cc. B. Robinson 9695 from Luzon and Elmer 10985 & 1341 from Mindanao. He keeps C. eeps C. formosana Rolfe separate from C. blancoi in his key and text, but makes the comment "There were no , specimina of this spe- cies "within our reach, but from the Sasaki nrene we should say that it is identical with Cc. Blancoi." Prei (1932) sums it all up in the following statement: "Distri- ution: Formosa and the Ae g pe its other distribution in Se laysia uncertain. Callicarpa form formosana Rolfe differs from its ro usually 6 to 15 cm. long, 2.5 to 7 om. wide, the upper surface rough due to falling off of the stellate hairs, glandular densely pubescent beneath; glands s agate yellow 3 petioles about 1 e 9 ; out, glabrous within. Corolla-lobes wide spreading about 2 m. in diameter, sparsely pubescent and ular without.. Stamens }, e- . erted; anthers oblong, 6 mm. in tang Sa: exserted. Ovary - Fruit globose, 2 m he cites the ar - in following specimens: CHINA: Chekiang: "Chiao 326, Ching 1838, Keng ua. Pukien: Chang & Metcalf 197 & 216, Cheng 103-1 & 2:39, Chang » abs, as Kel, dy 2198, & 6677, En 2607, Fong 212 & 21 Sing 506 "$306 & 6166 $766, Po 12120, and Tai 11793. meccres ka 905. Kwang tung: Chun 5706, ar, 5715, 5828, & 7555, Herb. Canton Ghr. Coll. 32, Mel 920 (A), E. D. Mer Merrill 10838, Peng s.n. oe chow region, April 1919], Peng, » ae, & , Tak 3 & Chow 2221 & 23h, Tsiang 20315, and Ying 13)2 Tee CHINESE COASTAL ISLANDS: Lantau: Sean MoT HONGKONG: Chun 517 and Ying 3h7. enry (1896) reports the species from Lambay Island. Matsumura (1899) & cites Hiraoka s.n. [Shinchiku] Kelung, Pikoh, Shalyootoo, Taipe], Owatari s.n. [Panlyau & Too- tsumra & —— , pmo} Girancho; Tensonhi, X1.1899], Miyaki sn. - [Kus- rere et Shintengai, 1899], Yano s.n. [in monte Taiton, —Vart. 1897), and Yokoyama 48, Li (1963) cites, also from Formosa, Chen 8.0, to £31, Sasaki 211,35, Tanaka 89, T 82, Wilford 193, and E. E. Hy Wilson = toe a bs 20238, 10 , 11012, & - Henry 10L8 from sand ac Tana 1967 Moldenke, Monograph of Callicarpa 225 10397 from Lanyu Isl ~~ Material of C. Sa has been misidentified and distributed in herbaria under the names C. angusta Schua., C. bicolor A. L. Juss., C. cana L., C. dichotoma (Lour.) K. Koch, C. giraldiana mae C. giraldiana Pamp., C. japonica Thunb., C, macr On the other tent e Lizardo 30, McClure 3038 [Herb. Canton Chr. Coll. 9591], and T reac 21989, “distributed as C. formosana, are actually f i S.n, [Herb. P Philip. ak Sei. 145203 & 7223] are var. glabrescens ne We pis aa Chun 5828 is the type aerate of var. chinensis dii eee eee ee ————eee In all, 29 herbarium specimens and 5 mounted photographs, in- ae ee photo totype material of some of the names involved, ve Additional map age “CHINA: Chekiang: Chiao 828 (Herb. Univ. Nanking 1128) (Ca——325125, W--1426628), 1838 2 Ei erenetpa 14128 (Go, N); R. C. Ching 1838 (ca—28120h, S). Fukien: T. C 197 (Vi), h645 (Du—200031) ; Chang & Metcalf 197 7 (Ge cehg0e6), 2b 216 (Ca—2)9173); c. H. Cheng 1031 (Ca—286996), 1088 (ca-~286958), 2298 8 (co 565676), 118 (Ca-—563776), 1439 (Ca—286958), 300k Gr 7602); H. H. Chung 881 (Ph), 120) I, (Ca—-22h680), 21:5 (Bz— 18168). 231 (Ca—2329h5), 2677 (Ca--232857), 3021 (Ca—2h36 Ca—20297), 7727 (N), 7865 (N), 828) (N); En 2607 (Ca—299582) ee 212 (Ca—300065), 218 (Ca--300042); Ging 5326 Setar se be Kiang (i), 6766 (ca—322328); Po 12120 (Ca--325772, Um—8l). si: S.K. Lau 392 (S, W—1752677, W—19h1h18), boo (S, W— 1753137) . Kwangsi : W. T. Tsang 23820 (N), 2h6h2 (uN). Kw Kwang tung: eee Canton Chr. Coll. 12008 (S, W—12h7385); S. K- Lau 696 (MN); oat. 920 ( Levine sun. S.n. [Herb. Canton Chr. Coll. 32] (i— 776523); Mell 220 (N—photo); E. D. Merrill 10838 (Ca—300797, oe. Gg— 237836) ; Preng & & Groff 2802 ( (cg—31972, N), Sn. se {[Herb. Canton Chr. Coll. 2802] (Ca—301022) ; , Tak, & Kin 9 (Herb. Canton Chr. Coli. 12008] (ca 274877) 5 Sin 9004 Girt Te & Chow 23h (Herb. Canton Chr. Coll. 14295] (Ca—316962), 2621 [Herb. Canton Chr. Coll. 1682] (Ca—319357); To & Tsang 12008 (N); W. T- aie 20496 (Ba, Bz—17599, W--1753685), 20596 (Ca—611373, N), “a 226 Pur Toroe re Vol. 1h, no. 4 (Ca—11185, I, N, N, S), 21388 (Bz—-17601, Ca--112h6, I, N, N, S), 21586 (Ba—17600, Ca—11127, 1, N, N, S); Tsiang 2158 (N); Tso 20315 (N), 21315 (N), 21160 (N); Toui 118 (W--1751503), 119 (Ba, Bz—17598, N), 212 (x, T—I75)555), 6 613 (Ba, N, W—175)765) Ying 13h2 (Ca—360133, 1 W—1513175), 2158 (Bz —18166, Bz—18167). Prov Gressitt 721 (I); S. K. Lau 96 (B, Ca—525179, I, Mi, N, W—-1629076 3706 (Bi, S); te Tsang 650 g 850 (Herb. Lingnan Univ. 16349] (N, S)5 Wang 33688 (N). Lantau: W. Y. Chun 874 (Ca—357970, Ca—37)136) HONGKONG: N. K, Chun 077i (Gg—237830) ; W. ¥. Chun 5147 (Ca— 2 Ge) Ying 347 (Ca—358380). WESTERN PACIFIC ISLANDS: FORMOSA: & al. s.n. [April 13, 1961] (Lb—-18707); Faurie 305 (Du-—- ui, “v—825h); Gressitt 6 (N), 215 (N); ta, Kanehira, & Ta- naka 263 (Ca—31)5488); A. Henry 435 (N, W—L553N6), Td (N, #— 455423), 1048 (N), 1947 (N, N); Hosokawa 9905 (Bi); Kanehira 21154 (Ca—344599); Keng K.1021 rane (W—20357h1); Kuntz 028 (W—=2336852)} Le L. L. Liu s.n. [April 10, 1961] (Lb—9l2h, Mi); Mor: Mori s.n. [Aug 1 1936] (W—2063359) ; y W. Ream s.n, [May 1h, 1950) (Ws); Sas wt 21435 (Ca—3h4h30, N), s 8.n. (Herb. Govt. Formosa 20962] (W—photo), s.n. (Herb. Govt. Formosa 21135] (La, W—137298); T. Tanaka 89 (La, S, ate 20991 eae TS N), 10988 (N); Tanaka & Shimada 10988 (B, B, Go , S, W—1577h61); E. H. Wilson 10770 (Ph, W--1053010), "10880 ¢ #1053033); Yamamoto 2367 (N)- HILIPPINE ISLANDS: Babuyan: R. C. McGregor s.n. eco b. Philip. Bur. Sci. 10108] (Bi). Batan: M. Ramos s.n. (Herb. Philip. Bur. eg 80655] (Bz--17597). Bohol: R. C. McGregor s.n. [Herb. Phil- - Bur. Sci. 1239] (Bz--18136, N); i. M. Ramos s.n, [Herb. Philip. a Sei. 42808 (Ca—2h2hh0), s.n. (Herb. Philip. Bur. Sci. Se igo orci: Catanduanes: Ramos | & Edafio s.n. [Herb : Ga (Bes 166511). CHINESE COASTAL ISLANDS: Hainan: Chun & Tso 757 (B, 8 ; Bacani s S.n. [Herb. Philip. Forest. Bur- Forest. Bur. 29136] (Ca--2680)2, S); M.S. Clemens 5881 (Ca— sal ge (Ca—27L174), 17400 (Ca—304358, N), san. (I1iea", an) Bae safe sen. (Herb. Philip. Bur. Sci- ca--321810, N)5 29 (Bz--181)1, N), 8151 (Bz— 181bh, N), 14352 (ae, ete N, S, Ut-67399), 17621 (Be—18133, Ca—271LH1, ¥, , Ut—67247), 22211 (Bz—17293, C8 1967 Moldenke, Monograph of Callicarpa 227 7817, N); Fénix s.n. (Herb. Philip. Bur. Sci. 12642] (Cm); F.C. Gates 5270 (Wis), 6286 (Ka—l615), 6600 (Mi); Haenke 79 (Ca— 260936); Juliano 9 (Ca--308171); Lete 87 (Ca—30H611)3 Lizardo s. n. (Herb. Philip. Forest. Bur. 29935] (Ca—268043, N); Loher s.n. (Rizal province] (Ca—229195); Madrid s.n. [San Quintin, Sept. 1925] (Ca—2919))6) ; R. C. McGregor s.n. [Herb. Philip. Bur. Sci. 11507] (Gm); E. D. Merrill 145 (Bz—16153), 2522 (N), 2688 (N), Sp. Blanc. 637 (Bz-~1813), N)3 R. Meyer s.n. [Herb, Philip. For- est. Bur. 2520] (Bz—~18145, N, Po—6815); Paraiso sen, (Herb. Philip. Forest. Bur. 30917] (N); Quisumbing 2261 (Ok—17136), 1884 (Mi); M. Ramos 1663 (Bz—-18131, N), s.n. (Herb. Philip, Bur. Sel. 13281] (Bz--161])2), s.n. (Herb. Philip. Bur. Sci. 13950] (Cm), son. (Herb. Philip. Bur. Sci. 33019] (N); Ramos & Edaflo s. n. (Herb. Philip, Bur. Sci. 29051] (Bz~18130), sen. (Herb. Phil- 4p. Bur. Sci. 4573] (Ca--257639, N), sen. [Herb. Philip. Bur. Bur. Sci. 45067] (Bz—~-18139, Ca--2576h1, N), sen. [Herb. Philip. Bur. Sei. 5203] (N), s.n. (Herb. Philip. Bur. Sci. 75712] (Ca— 9195, N)5 C. B. Robinson s.n. [Herb. Philip. Bur. Sci. 6808] (Bz—-181),3); F. L. Stevens 225 (Ur), 537 (Ur); Vanoverbergh 177h3 (Ca--359365) Weiss 3630 (Bz—-18157); Whitford Ok (N); Re Se Williams 115 (N). Masbate: E. D. Merrill 3375 (N). Mindanao: ine 9020 (Bz—~18151); M. S. Clemens 190 (W—619268); Elmer 10985 (Bi, B2—18150, N, Ut—-276h3, Vt), 13441 (Bi, Bz—-181L8, Ca--27208h,, N, Ut--3352h); Fénix sen. [Herb. Philip. Bur. Sci. hee (Ca—238)90); C. M. Weber 1055 (Ca—239439, Cm) $ Be Se ams 2577 (N, N). Mindoro: Abaca 1 (Herb. Philip. Forest. Bur. 29531 (ca-2oseobl), Conklin 196 [Philip. Nat. Herb. 17535] (i); Kienholz 327 [Herb. Philip. Bur. Sci. 15310] (Ca--26301h), = (Herb. Philip. Bur. Sci. 1532] (Ca—263008); Mangubert s.n. erb. P M. Bene lip. Bur. Sci. 16383] (Ca—3091)0); Velasquez 10 (Bi). * R. G. Smith s.n, (Aug. 1945] (Ur). Island undetermined: Née 25 (Q), 28 (Q), 32 (Q). CULTIVATED: Formosa: Keng K.1052 ~~*035769).” Martinique: Duss Wk2 (B), sen. (B). Philippine Islands: A. Castillo s.n. {College campus, January 3, 1931] (Hp). ALB wan 821 & 9h, nom. md. 1963. a graphy: Li, Woody Pl. Taiwan 821 & 9h. 1963; Moldenke, Suppl. 8: 3. 196k. = As yet I have not been able to ascertain the original place _ trinomial, if, indeed, it was ever — a? Published, I assume that the taxon differs from the typi 228 PHYTOLOGIA Vol. 14, no. h of the species in ha white corollas. If so, then it is ae probable fe % Lizard Lizando 36 and the specimens cited below belong wee since they are described as having had white flowers. n all, only 2 —— specimens have been examined by m Citations: CHINA: Kwangtung: Wotlure 3038 [Herb. Canton sets Coll. 9591] (Ph); We . T. Tsang 21989 (S). CALLICARPA FORMOSANA f. ANGUSTATA Molderke, Phytologia ): 125. 1952. Synonymy: Callicarpa fesse fi ers ta (Rehd.) Moldenke ex Li, Wo Pls Pads <3 823 - 196 brie graphy: M Neda; Par tsing ia nck 121, 122, & 125. 19525 eiteae, Boke inte. 27: 98h. 1953; Moldenke, Résum$ 172, aha Lh. 19595 1 » Woody Pl. Taiwan 823. 1963; Voldenke, Phytolo ly: 189 & 190. 1966. z orm i aitrece from the typical form of the species in hav- ing its leaf-blades unifo eola a e type of the form au Aeggoem by "Har ley Harris Bartlett (no. 6082) at Taiheisan, Formosa, at an altitude of 3500-000 feet, in September, 1936, and is deposited in the Britton Herbarium at the New York Botanical Garden, It seems very probably to me that Susuki's C. formosana var. zonglfolia Suzuki is the same as the form here under discussion, i but not having seen any type material of Suzuki's plant, I hesité to make the reduction. form, rather than a varie ety, my epithet remains the valid one in any case. The Sasaki s.n, {Herb. Govt. Formosa 21135], Mori S-Me (August 10, 2s and S. K. Lau 96, cited by me under C. formosana tually represent f. angustata instead. Li (1963) 18 is in error sion he regards my f. ane as a new combination for ta th tirely different type, as is plainly stated The two taxa are abundantly distinct. . is described by collectors as a bush or shrud, inch in C. blancoi Rolfe, C. caudata Maxim., C. formo Rolfe, C. micrantha Vidal, Cc. pedunculata Re Be, and Cc. steno phyla Wi In all, 11 herbarium specimens, including the type, have been Citations: WESTERN PA 6082 (N—type) ACIFIC ISLANDS: FORMOSA: H. H. Bartlett . a 3 Keng & Kao K.1273 (W—2036017); E. H. Wilson (W==1052891, W—1052852),— TA (W—105311,, w-—¥092623). PHIL“ 1967 Moldenke, Monograph of Callicarpa 229 IPPINE ISLANDS: Luzon: E. D. Merrill 1718 (Bz—-181h7); Ramos & Edafio sen. [Herb. Philip. Bur. Sei. 6955] (Bz—18138, N); Ea Se Williams 1 1158 (N, W=-70732 a); CALLICARPA FORMOSANA va - CHINENSIS P'tei, Mem. Sci. Soc. China 1 (3): gimewatlyy China] 30. 1932. Synonymy: Callicarpa pedunculata var. chinensis (P'ei) Metc., apo pot Journ. 11: ts 405,06. 1932. graphy: Ptei, Mem. Sci. Soc. China 1 (3): [Verbenac. China) 1 "ls & 30. 19325 "Vete., ees ceed l: — 11: 405-06. Ca anees Known Geogr. Distri nac., ed. 1 Highs & “i d. 2; TE arr. 19495 | ‘esihenke, Résumé 168 & 1959; Moldenke, Résumé § ike Foe his variety differs from the typical form of the species having its leaf-blades oblong to elliptic and entire or subenieete and the inflorescences lax and diffuse. Prei (1932) describes the Plant as a shrub, about 2.5 m. tall branches branchlets de pubescent ung; leaves de- 8 meath; secondaries about 8 on each pore of the midrib; inflorescence axillary or sometimes sub terminal, and diffuse; peduncles 2—5 om, long; fruiting-calyx glandular; vate globose, 2.5 mm. wide, glandular, orange when type of this variety was collected by Woon-Young Chun (no. 828) in open brush between Changkiang Chengkou dge, — are two sheets of this si sg The taxon is known thus ar only from the original collection. all, 2 herbarium specimens —_ one photograph of the type d by m Citations: CHINA: Kwangtung: W. "r. Chun 5828 (Ca—34728)—iso- type, N—isotype, N—photo of type). een, RMOSANA var. GLABRESCENS Moldenke, Phytologia hi: —l2. 1952, Synonymy: Callicarpa caudatifolia Merr. ex Moldenke, Rés “ge ie ll, in syn. 1962 [not C. caudatifolia Koidz., 1925, nor 962]. “el ee » Phytologia h: 1-2 & 1. 19525; Molden- ke, Biol, Abstr, Oke mer - 1952; M Ncldenke, Résumé 1 gar 19593 Moldenke, Résumé S Suppl. he + i562; Moldenke, ea 1s 142. tits variety differs from the typical form of the species in having the lower leaf-surfaces entirely glabrous or else sparsely and only on The type of liected by Maximo Ramos and Gre- orto B. Edafio tacucveini een of Science 29137] on Mount 230 PHYTOLOGIA Vol. 1h, no. Tulaog, in the province of Tayabasi, Luzon, Philippine Ts}ands, in lec y Richard Crittenden McGregor (Herb. Philippine Bureau of Science Seti in the province of Laguna, Luzon, between June and August en and was deposited in the herbarium of the Philippine Science at Manila, but is now destroyed. The C. caudat~ ifolia 2 Koidzumi is a synonym of C. japonica var. angustata ta Rehds, while that of Maximowicz is C. caudata Maxim. Collectors describe this variety as a plant 2 m. tall, the 3 cm. in diameter, growing in thickets and forests, in open places, ae in March, May, and November. The flowers are described , ee Cea on Ramos & & Edafio s.n. [Herb. Philip. Bur. Sci. 5203 & i2e3hs Material has a misidentified and distributed in herbaria e C. formosana Rolfe, C. longifolia Lam., C. micrantha Vidal, and C- ta —_—_——— In all, 15 herbarium specimens, sednlhne type material of both names involved, have been examined me. Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Luzon: dafio s.n. [Herb. Philip. Bur. Sci. 8817] (M5 B R. C. McGregor S$: ne [Herb, Philip. Bur. Sci. 22919] (W—898252); Me Ramos 05 SoNe [Herb. Philip. Bur. Sei. 27662] (Bz—17660), sn. Po ecaiey ways e . Bure 28513] (Bz--18135), s.n. (Herb. Philip. Bur. Sci. 29137) age sn. (Herb. Philip. Bur. Sci. 45203] (Ca—308530), sn. [Herbd. Philip. Bur. Sci. 561i] (Bz--181,0, N), sen. [Herb. Philip. Bur- (Bs——-17656, Bz—17657, N). Mindoro: M. Ramos sn. [Herb. Philip. Bur. Sci. {612] (Ca—308873, N). CARPA FORMOSANA var. LONGIFOLIA Suzuki, Trans, Nat. Hist. 50c- Formos. 25: 130. 1935 Bibliography: Suzuki, . Soc. Formos. 25: 130- 19353 Li, Woody Pl. Taiwan Seer & ees 113634 Moldenke, Résumé Suppl 8: 3. 196k; Mo: Moldenke, Phytologia 1): This ha Ha differs from the ae Ponte of the species in Ae having the leaf-blades thi branous, oblong-lanceolate to ov tas lanceolate, caudate-acuminate at the apex, cuneate at the e, miu mucronate-dentate along the ieraine 1S ee long, 2--3 cm. wide, glabrous beneath except for he puberul midrib and venation, ype of the variety was collected by Sigetaka Seeks (no. ae Doba, Formosa, on July 28, 1929. In his original 8 on the collector notes that "The distinguishing Secs the shape ppecies and the present variety seen to be found in oad sien at = viz. ate latter has longer and more cune 1967 Moldenke, Monograph of Callicarpa 231 mode of oecological adaptation. The present variety has been rarely Picci in low altitudes of Formosa as a chief element of secondary orests ." It seems highly likely that C. formosana f. angustata Moldenke is the same plant as Suzuki's variety, but | ety, but not having as yet seen any type material of Suzuki's plant, I hesitate to make the re- duction. If the taxon is to be regarded as a variety, Suzuki's epithet must be adopted, but if its status is to be that of a mere form, then my epithe t remains valid. CALLICARPA FULVA A. Rich. in Sagra, Hist Cuba 11 (2): 15. 1850 (not C. fulva Griseb. p aoadle ed Callicarpa fulva in Leén & Alain, Fl. Cuba li: 307, 1987. Cal Callicarpa fili~ D. “ks 1788; : Rich. in Sagra, Hist. fal Sauv Cub. rey oa 1: 386. 1893; Millsp., Field Columb. Mus. 1 (1908) and 2: 313. 1909; Urb., Symb. Antil. i "rr 386. z “57, 1911; mae & Millsp., Bahama Fl. 373. 1920; ee - Kew. : 1. 5: 43. 1921; Urb. in Fedde, porerts Spec. Nov. ty Moldenke in Fedde, Repert. Spec. Nov. 33: 1k] (1933), 398 i (1936), and 40: 53, '56—s9, 61, 6h, Noe, 67, 68, 73, 78, 82, 9, oe 125, 126, 128, 129, & 131. 19%; agro ths . st Inval vicenn e, Pre V & 86, 1942; Moldenke, Alph. Invalid Names 8, 3 &l. 19125 ¥ Moldenke, ee List ‘cite | 1: Bs 7%, 18h, 185, 192. 222 & 343. 1946; E. J. Salis Kew. Suppl. 10: aur Moldenke, Alph. List Cit. 2: sie, 87, 647, & 649-651 19, 1; 109; Geo, » Résumé 50 Sp ee 148, 119,186, '& 187. 1966; J. A. Clark, ° Rod Richard! s original (1850) tundsdn ary is worth repeating in full here because of ge es in interpretation given his Species by various pa acdsee ag the igetes "Callicarpa tota ee “a~floccosa, fulva; foliis oppositis ovali-lanceolatis aut o 232 PHYTOLOGIA Vol. 1h, no. 4 ovali-lanceolatis acutis, basi sensim attenuatis margine subcremu- latis, superne glabriusculis, subtus shied Dred pigueary yo ea 7 panulato glabriusculo, ore dilatato et subtruncato; drupa eve glabra, magnitudinem seminis Sinapeos nigrae vix excedente. flores rosadas 6 color de lila, est& caracterizado por el ei vello que le recubre casi en totalidad. Se aproxima algo, po 4 fi de called "filigrana", growing on limestone rocks and cliffs, beatin hillsides, in charrascales and wet woods, at altitudes na one to 1000 meters, flowering also in April and December, 6063 in July. The flowers are described as "pink" on R. A. Howard 6053, "white" on Le6n 12015, and "white or slightly purplish" on Leén, Clément, & Roca 10019. ton & P. Wils. and may possibly represent a The sen al 13327 specimens cited below are almost as glabrescent as mater Which is typical of var. glabrescens Moldenke, while his 13326 is very pubescent. Clément 3396 has extremely large and very tose leaves. The C. fulva ascribed to Grisebach, referred to in the synonyay above, is in part C. crassinervis Urb. and in part Cc. grisebachit Urb., that of "A, Rich. apud Millsp." is C. hitchcockii Millsp., eC. fulva f. foliis lanceolatis Griseb. is Cc. lancifolia ——— illspaugh (1906) makes some extremely interesting —, ~ bout what he then called C. fulva but later re-named C. hitch cockii Millsp.: "Mr. Hitchcock's plant [from scrub land and — pice, Cat Island, Bahamas], while having (on account of its grea reduced Cuba, May 30, 1859) nevertheless has no other characters of ee entiation. It is fairly well connected in leaf size and form t Far. Sept. 29, 1860, which is in the same fruit stage as the Hitchcock plant. Of this Farallones plant Mr. Wright says, 'A slender bush 6 to 10 feet high: 4 4 J ~++-++-considers this species synonymous with the Jamaican C. fer Swartz, this however is not the case, the differences are broad and evident. Grisebach in his Catalogus Plantarum Cubens inc C. fulva Rich, Mr. Wright's Majanabajo, Cuba 3173 with the remark forma foliis lanceolatis'. The sheet of this nur 1967 Moldenke, Monograph of Callicarpa 233 ber in Herb, Gray, Cambridge (one of Mr. Wright's original ser- ies), proves to be an entirely different species" which he then Proceeds to name C. lancifolia Mills Sp. Material of C, fulva has often been misidentified and distrib- se in ‘aie as C, ferruginea neh On the other hand, the A- 9868, tate & Wetorin ferruginea Sw., age while. Clément, Alain, & Chrysogone 6698 6698 is > floccosa Urb. R. A. Howard 6053 is a mixture with Cc. C. fulva v var. glabrese. rescens Moldenke. In all, 7lh erbarium specimens, including type material of all hapa hanes involved, and 22 mounted photographs have been examined Addit & emended citations: CUBA: Oriente: Acufia 12690 mel 13326 ( (Es, N), 13327, in part (Es, N); Alain & 1 & Acufla 7686 ); Main Alain & Lépez Figueiras 7150 (Bm), 7229 (Bm); Alain & Mor- tn 5078 (s (Ss), 51a) (2); G.C. "Bucher 13 » (Rg--8153) ; Mrs. G G. Ce Bucher 128 (N, Rg—6152), 728° (Hia); Clément 3396 (ia, W)p clésent & Alain 3945 (Ha, Ha, N); Clément & Chrysogone 3396 (Mv); 8773 (); R.A. Howard 6053, in p in part (N); Leén 12015 (Ha), eas (Ha), 20186 (WN), 20276 (Nl); Leén & Clément 20276 (Ha, N), 23029 (N); Leén, | Cement, & & Roca 2009 (Ha); Leén & Victorin 17295 (Ha); Linden 19 1969 (P= otype); | sen. [Leén 18551] 51) (N)3 Morton & Main | 8993 (2281), sit lie eis CEBSAAD) | Van Hermann 11766 (Bs); Vict, Victorin & Alain 21730 (Ha, N); G. L. Webster 376h (Mi); Ce ¥right 1,30 [Jan.—Jul. 1859] (E-—photo, F-24613, | 263, errata Seay Photo) , 430 [1860] (Ca--93679h), 430 [1860-—6h; H Sauvalle eH (iW), 5070 [ieet, Savalas TPR (e_iSE, Suite et CALLICARPA FULVA var. eae Moldenke in Fedde, Repert. ate: Aa uw. a Bibliogr Meloadte res Fedde, Repert. Spec. Nov. 33: 1h1 733), 39: 299, (1936), and 0: 59° 119, 126, & 131. i936 Mol- denke, sCeoer. Dis rade erage 5. tf, uoldenke, ait me & 86. rier ee it. 1: 273 2 310° (198), "9: 420 & 647 sis), 3: 3: Boh | (1919), and hs TS & & 1158. 1949; Moldenke e, Known Geog Di - Verbenac., 1949; Alain in e in L hs ir voice, Résumé 50 & hi. 1959; J. ‘% "clark, Card Ind. Gen. This plant has been collected in anthesis in July. R- A. How- = $053 appears to be a mixture of the typical form of © Cc. - fulva var. glabrescens, at least insofar as the Britton Herbarium metial 4s concerned. Acufla 13327, cited by me under typical C- suet. is almost as glabrescent and typical material of this var- and may be , 18 h satearee ai cociadeests including the type, and 7 moun~ 23h PHYTOLOGIA Vol. 14, no. & examined b ear ceed pitas: CUBA: Oriente: Acufia 14117 (Es); Mrs. G.C. Bucher 11456 (Es); R. A. Howard 6053, in part (N, S)5 Leén & Clément Sas rh (N), 23308 (N); Linden 2087 (Br); C. Wright Ae bell (Br--isotype, D—-612063—isotype, E—119139—type, —~—2),1,612——isotype) . CALLICARPA FULVOHIRSUTA Merr., Journ. Straits Br. Roy. Asiat. Soc. 76: Pe Qt de 1917. Synonymy: allicarpa fulvo-hirsuta Merr., in herb. Biblio cesoigs D. Merr., Journ. Straits Br. Roy. Asiat. exe’ 76: 113--114. 3 A. W. Hill, Ind. Kew. Suppl. 6: 3h. a! on oe Roots Se coer + Dietrib. Verbenso ey, #53; él & 86 (19h2 hairs; leaf-blades paler beneath, chartac in elliptic, 12—1) om. long, Soabycks cm. ane: 5 bebocially = the sharply acuminate apex, sharply dentate along the margins ex cept for the entire or subentire basal portions, subequally apt rowed to the acute base, the per surface densely hirsute on tne pale, shiny, small, waxy ace of the midrib, promi nent, curved, ? . ‘culation dark-brown in contrast to the. pes lamina surface; Satlorsececns densely fulvous stellate-tomentose or —pubescent intermixed axillary rsute; flowers }, calyx cupuliform oat long, the te, obscurely l-denticulate, lla with sige pang on the outer surface; coro white, - long, externally glandular, subequally })-lobed, the lébes oblong, about 1.5 m. 1 to obtuse at the oes stanens ij on the back; fruit depressed-globose, red when mature, about 3 mm. in di. sparingly The 't ype of this species was collected by Mary Knapp ge (no. 9846) between Kibaya and Keung, Mount Kinabalu, British No 1967 Moldenke, Monograph of Callicarpa 235 Spects to Geunsia havilandii King & Gamble, but the indumentum on its leaves is less dense, sien its flowers are smaller and with five, not four stamens » and is hence a true Callic ." The taxon is eae thus far only ane: the original collection. It is the bove n all, 2 herbarium specimens, including the type, and 2 moun- ted photographs have been examine ° Citations: INDONESIA: GREATER S ISLANDS: British North Borneo: M. K. Clemens 9846 (Candi hies—aaotepe, N—-photo of type, Ph--type, Z——photo of type). ‘A FURFURACEA Ridl. , Journ, Fed. Malay States Mus. 10: CARPA pose 1920 blio ography: H. N. Nes roan fiat, - State 150. 1920; H. N. Ridl. Mala eui-él6, 192 19234 > - Hill, ote . Suppl. “ee “34 (2926) aad 7 eae Lee 3 Moldenke, Tom G eogr. Distrib. Verbenac., ed. 1, 60 & 86 (1942) and ed. 2, 38 & 177. 1949; Moldenke, Résumé 179 & yh. 1959. Branches brown~sc curfy; “Leaves decussate-opposite; petioles d 12.5 mm, long; leaf-bla ranous, oblong, 12.7 cm. sities 7 : ris dentate along the of truncate at the base, smooth abo th n tion, pale beneath, with whitish veins, the veinlets and reticula~ bees: scurfy, the midrib scurfy above; cymes paniculate, ter- ? cm. long or longer, many-. red, brown-scurfy; pedi ree at the apex; stamens )); filaments ie inserted at the ise of the corolla-tube and shorter than it; anthers large, ob- ", Slightly exserted. The type of this ag oR species was collected by Evans at Gunong - Ridley (1920) anew that "This is of the section ise C. arborea, but dis ti. ed by its Pr ned fur and leaves white beneath tn with brown, cpa reticula- ons I know nothing of the plant except what is stated in the bibliography listed above. CALLICARPA ee —! Bot. Mag. Tokyo 32: 56. 1918 [not C. SynOREy ¢ subpubescens Maxim. ex Koidz., Bot. Mag. “, 32: 56, in syn. 1 sa = po [not C. ens Hook. & Arn., Bibliography: J. Matsum., Bot. Mag. Tokyo 3: 318. 1889; Hatto- ri, Journ. Coll es 23 (10): 3h. 1908; J. Matsum., oll. Sci. Univ. cage 3 (10) — wie i %. . Kew, Suppl Ponte Spec. er. 40: 84. 1936; Koidz. in Shirasawa, Icon. Essenc. orest. Jap. 2: fig. 2487. 1938; Moldenke, Prelim. Alph. 236 PHYTOLOGIA Vol. 14, no. k valid Names 13, Roe Moldenke, Known Geogr. Distrib. Verbenac., ed. 1, 58, 61, ie 9h23 Moldenke, Alph, List Invalid Names ll. 1925" Hara, nie: yi? rm. Jap. 1: 183. 1948; Moldenke, Known Got Distrib. Verbenac., ed. 2, 133, 140, & 177. 1949; Moldenke, Inform. Mold. Set 48 Spec. [2]. 2954; “ddr, Phytol ogta ss 28. Bg Moldenke, Résumé 172, 181 » 27, & bh. 1959; H Phytogeog. Japan 87. 1959 section’ nan Suppl. 4 ke 8: °19625, Moldenke, Phytologia 1): 163. 1966 Illustrations: Koidz. in Shirasawa, Icon, Essenc. Forest. Jap. almost completely glabrous shrub; branchlets terete, at fi sely s te-farinose but the pubescence soon evanes- cent, those Pa foe: k-purplish or black, those of last - » the older ones grayish; leaves decussate-opposite, deciduous; cog Ape —2.5 cm. long, at first tellate- the corolla; filaments glabrous; anthers oblong, resinous~ punctate on both sides; style long-e xserted, glabrous; fruit glo- pigeons € cymes, 3—~l mm, wi common name "shima-murasaki", rtenee: de a the plant as a shrub or bush, 5—12 Feet tall, growing on hillsides and in open villages, at 75 meters altitude, flowering in April, August, and September. The flowers are described as "pale-1 e-lavender" on Walk. awada, & Aman bree reports that the species is "common" on Mukojima. The Ce known as > lamii Nneehasie: seg tede we ed : Tn alt 6 are herbarium specimens have been examined by me. 1967 Moldenke, Monograph of Callicarpa 237 Citations: WESTERN PACIFIC ISLANDS: RYUKYU ISLAND HIPELAGO: kKinawan Islands: Okinawa: Field & Loew 6m (Mi, Wa-191,2288). Saki- shima Islands: Iriomote: Walker, Tawada, & Amano 6507 (Le, W— 2 Z). BONY ISLANDS: Mul iit tines Ee Be Wilson 8332 (ij Las porty GLANDULOSA Fletcher, Kew Bull. Misc. Inf. 1938: 199. 9 Bibliography: Fletcher, Kew Bull. Misc. Inf. 1938: 199, 12, & 4. 19385 1 Moldenke, Known Geogr. Distrib. Verbenac., ed. 9& 87. 192; 'E. J. Salisb., Ind. Ken. Suppl. 10: 38. 19473 foe Geogr. Distrib. Verbenac., » 137 & 177. 1949; Anon., ew Bull. Gen. Index 1929-1956, oe pi me eg Résumé 177 & ull. 1959, shrub, of diffuse habit, about 2 m. tall; branchlets obtuse- ly tetragonal, often terete, * prownis sh-gray, glabrous; leaves de- c c Lag c Ereen above pars ate—pubescent, with erous ro red sessile glands, gray-green beneath; midrib conspicuous above, Prominent beneath; secondaries pea pairs » conspicuous above, pro~ ous beneath, uately joined at the margins; ter- tlaries t rse, alandah: te more or less ; inflorescence rs ainda’ ; peduncles 3—5 cm. long, some- t pubescent; pedicels 2 m. long, glabrous; bractlets smal » to 2 mm. long; fl not seen; frui e, globose, I » With sessile yellow glands at the apex. wi The type of this little-known species was collected by Ar Francis George Kerr oe 11169) at Chumpawn, Ta Ngaw, Surat, Thailand, by streams in an evergreen forest at an altitude of a-~ dae lamk. affinis sed foliis glandulis rubris itis, nervis lateralibus paucioribus differt." paces GRACILIPES Rehd. in C. S. Sarg., Pl. Wils. 3: 371. eae e Rehd. in C, S, Sarg., Pl. Wils. 3: 371. 1916; H. Kew. Suppl. 6; iL. 1926; Plei, Mem. Seis Soc 1 (3): Lverbe- hac. China] 15 & 26. 19324 Moldenke, Known Pegi F Verbe nate, ed. 1, 56 & 87 (1942) and ed. 2, 130 & 177. 19493 Koldenke, Résumé 168 & ih. 1959. , her any ee above , elliptic-ovate a ovate-oblong, 238 PHYTOLOGIA Vol. 1h, no. k remotely denticulate along the margins except on the lower one- » sometimes almost entire, at first sparsely farinose-stel- rous bove, elevated and distinct beneath with a flavescent color; cymes few-flowered, small, 1—1.5 cm. in diameter, yellowish far- flowers not seen; f ng-c sparsely stellate-pilose, glabres= cent, the rim obsoletely -denticulate; fruit apparently purple, subglobose, scarcely 3 mm, ameter. The type of this distinctive species was collected by Augustine Henry (no. 7690) somewhere in western Hupeh, China, and is deposited in the Gray Herbarium of Harvard University. Rehder age —— ments that "This species does not seem to be closely related to any other Chinese species; it is characterized by its capensis: generally elliptic-ovate leaves, dark green above, whi es, because of its short internodes. It is allied to Cal- licarpa cana L. Dag which it differs by its smaller leaves which are shining | above Only one aye tica specimen of this species has been examined me, Citations: CHINA: Hupeh: H. C. Chow 769 (N). CALLICARPA GRISEBACHII Urb., mgr th Ansa, 7: 356—357. 1911. Bmended synonymy: Callicarpa fulva Griseb. (in part) apud Urb., Symb. antil. 7: 357, in syn. 1911 oe C. fulva A. Rich., 1650]. Known “ad - V *» 1942; Moldenke, , tS t Invalid Names 9. kes Woidenke, & x? oe Lint Sor 1: 75 187, & 306. 196; KR. %, - L, Moldenk 3, 1 62. - wid Moldenke, alph. List cit. 2: 120 & 187 Cus cs atu goths is. 1949; Moldenke, Known Geogr e.. ae ey & wih 1895 Alain in Leé be 305 & 09. "19873 M Résumé Résumé 50, 23, ei lids 2 19593 3 olden ke, Phytologia aie lave & & U9.” "The flowers are sagem rose-violet" on Ekman 8672. Fie Alain & Morton 520, opengl buted as this species, is is actually C.areolata | Urb. Tn all, 29 herbarium specimens, including type material of all 1967 Moldenke, Monograph of Callicarpa 239 the names involved, and 20 mounted photographs have been examined me Additional & emended citations: CUBA: Oriente: Clément 5930 (N)5 Elman 8672 (Go--photo, N), 8926 (Br--photo), 9217 (I—photo, N), 15633 (Mi, Mi--photo, N); C. Wright 1357, in part [Sept. 1859 an. 1860] (Br--isotype, E--1191h1—isotype, E—photo of type, ig pag ted ad Mi--photo of isotype, S--photo of type, W— 9. ° CALLICARPA HAVILANDII (King & Gamble) H. J. Lam, Verbenac. Malay. Arch, 52. 1919. Synonymy: Geunsia beccariana Briq. in Engl. & Prantl, Nat. Pflanzenfam. I, (3a): 165, nom. nud. 1895. Geunsia havilandii King & Gamble, Kew Bull. Misc. Inf. 1908: 105. 1908. Callicarpa havilandii var. pentamera H. J. Lam, Verbenac. Malay. Arch. 52. 1919, Callicarpa havilandii var. tetramera H. J. Lam, Verbenac. Malay. Arch. 52. 1919. Callicarpa havilandii H. J. Lam apud A. nag, Ind. Kew. Suppl. 6: 34. 1926. Callicarpa ferox Bakh. ex 0. e Résumé 2 in syn. 1 & Bibliography: ret & sean” bee Bull. Mise. Inf. 1908: 105. 1908; King & Gamble, Journ. Roy. Asiat. Soc. Bengal 7 (2), extra no, 801-502. 1908; King & Gamble, Mat. Fl. Malay. Penins. 21: oar 1909; H. J. Lam, Verbenac. Malay. Arch. 6, 52, & 362. 19195 : 3 Names 9, 2h, z 5 1: 266 & 267 ? & 25. 19423 Moldenke, Alph. List Cit. 1: ° 1946; He Ne & A. L. Moldenke, Pl. Life 2: 63. 1948; Moldenks, Alph. Mist Cit. 3: 751 & 763 (1949) and hk: 1173. 1949; Moldenke, T. Distrib. Verbenac., ed. 2, 145 & 177. 191,95 Moidenke, Phy- ones 4: 81 & 82. 1952; Moldenke, Résumé 192, 193, 23, & hhh. or tree, about 3,3 m. tall; branchlets densely and rugose- ly Stellate-pubescent with tawny or ferruginous hairs; leaves de- jah. gesely stellate- and glandular beneath with simple and ferru- ginous permanent hairs on the venation only, the re : Jang and peltate; midrib coarsely prominent; secondaries 7--15 ee veinlet reticulation conspicuous; cymes small, axillary, — Sile, few-flowered, 1.5~-3 cm. long, 3—l cm. wide, scarcely 2h0 PHYTOLOGIA Vol. 1h, no. 4 ing the subtending petioles, densely and rugosely stellate-hairy or ferruginous 3 peduncles 3--10 mm. long, rugose- ly stellate-pubescent; bractlets minute, subulate; pedicels short, slender; calyx campanulate, 1—2 m. ong, 1—-1.5 mm, wide, glab- rous on the inner surface, strigose-hispid or densely and rugosely stellate-hairy with ferruginous hair and gland th large resin-glands on the outer surface, its rim shortly 4- or 5-toothed back) with dense resin-glands; style filiform, j stigma capitate or subpeltate, more or less obscurely 5-lobed; 0- vary glabrous, densely glandulose with 1 bright~yellowish res- in-glands, |, lobed, the cells 1- or 2-ovulate; fruiting-calyx somewhat » persistent, flattened; f. ad when fresh, globose, rather large, 3—5 mm. wide, somewhat depres~ sed at the apex, us when mature, with sparse resin-glands, purple~blac —8 [—12] l-seeded pes ecti mn dry, with },. King & Gamble (1908) based this species on four collectio: rom Sara} and Haviland 889, 3549K, & 3549L, the been collected near Kuching and the others with no specific locality mentioned. The type of C. ferox is Tei jsmann 8500 from Borneo, deposited in the Herbarium Bogoriense Lam (1919) says "Without any doubt King & Gamble's species is 4 carpa (only opposite leaves, leaves denticulate, spt ®, fruit with 4-5 (not 7=-12) pyrenes). As in some other spe cies of this gems there ntamerous fom." Hi » pent wak, collected in flower and young fruit in November, 189). It is rth emphatic in stating that this plant has 11) t, but gen van den Brink (19 publi by Lam (1919) and by Bakhuizen van den Brink (1921), but actually eppeared in 1895. The latter author comments that it "is a doubt- Species.....Its relation with the C. Havilandid (King/Gamble) He J. L. is not yet certain.n cee Callicarpa havilandii has been found growing in forests. The flowers are reported as "white" on Clemens & Clemens 2178h, Moul- — 1967 Moldenke, Monograph of Callicarpa 2h ten 79, and Native collector 2827. Haviland & Hose mistakenly refer to the fruits as “berries”. » 11 herbarium specimens and mounted photographs have INDONESIA: GREATER SUNDA ISLANDS: Borneo: Jaheri 1734 ind Bz~~17608); Teijamann 8500 (Bz--1760h, Ba—17605, Be-- , N). Sarawak: Clemens & Clemens 2178) [field no. 6161] (eat Ko}, N); Haviland & Hose & Hose 3549 (V); Moulton 79 [Native col- lector 2827] (N—photo, N—-photo, Ph, Ph, Z--photo, }, Z==photo) . aire nant erry Millsp., Field Columb. Mus. Publ. Bot. 2: —=313 ed synonymy: Callicarpa fulva "A. Rich." apud Millsp., Pela Columb. Mus. Publ. Bot. 2: Pie sea! 1906 [not C. fulva Griseb., 1911, nor A, Rich., 1850]. Callic hitchcockiana 1% + ex Moldenke in Fedde, Repert. Spec. Nov. 40: 59, in syn. Bibliography: A. Rich. in ae, a Cuba 11 re 145. _ Milisp., Field Columb. Mus. Publ, Bot. 2: 180—181 (1906) and 312-313, 1909; Prain, Ind. Kew. maeis h: 3h. 1913; “Britton e og 2h 9. isha; ie at Phytologia 2: hs 1945; Moldenke, Alph. sas a l: £6, 6 65, 1799. are age , 308, ea 3 196; H. Me Moldenke, Pl. 6h. 10185 "\woldenk t Cit. 2s wise & Sis i9 (ashe), 3 928 & 929 (1919), ant bt "103 & 1039. 19k9 » Snom peo ct: m8 _ benac., 177+ 19) j Alain in *Leén he: 305 & 08s “39875 olden “ot Foeprrap ae A 3 50, 23, “ ie «1959 35 woidenke, Phytologia 1h: 1h9. cs Gen. ° e Ne In all, “mn ati rm se nN / fackeding type ——_ of all eg anes! involved, and 15 mounted photographs have been Additional & emended citations: BAHAMA ISLANDS: Andros: L. J. K. Brace 6965 (F-21369), 7100 (F—21i95). Cats Britton & ME- Spaugh 5819 (D—532333, F198620, F--198621, 01, W—645021), 5913 ©—198707—type), S9k6 (F—198726); A» S. Hitchcock s.n_ [Pore Howe] (E--1,73)2, F-17219). New Providence: Eggers LL69 (8, K, N, S). CUBA: Camagtley: Shafer 256 (F— — 250995, SLT.) 2791 (P~251258, W—B8931) . CALLIDARPA THABQUALTS Teijsm., & Bim. ex H. J. Lam, Verbenac Malay. Arch. 77—78, in ages 19195 * Bakh,, Bull. Jard. Bot. tenz. sur’; 3 3: 18—19. 1921. Synonyuy: Gallicarpa ericclona var. latifolia H. J. Lam, Ver 22 PHYTOLOGIA Vol. 14, no. benac. Mal Moldenke, Résumé 21, in syn. 1959. Callicarpa inae is "T. & + ex Bakh." apud Backer & Bakh., Fl. Java 2: 600. 1965. Bibliography: H. J. Lam, Verbenac. Malay. Arch. 77--78 & 362. Bakh., Bull. Jard. Bot. Buitenz., ser. 3, 3: 10 & 18-19. » ind. Kew. Suppl. 7: 36. 1929; Moldenke, Prelim. List Invalid Names 10. 190; Moldenke, Alph. List Invalid Names 9. 1942; Moldenke, Known Geogr. Distrib. Verbenac., ed. l, 6 & 87 (1942) and ed. 2, 1h & 177. 19493 Moldenke, Phytologia h: 79, 80, & 83. 1952; Moldenke, Résumé 187, 189, 19h, 21, 2h2, & + 1959; Backer & Bakh., Fl. Java 2: 600. 1965; Moldenke, Phyto- logia 1: 178. ‘ An erect shrub, 2—3 m, tall; branchlets obtusely tetragonal, rugose, very densely stellate-farinose with subferruginous, brow te » resinous-punctate beneath the tomentum, sparsely cussate-opposite, long-petiolate; petioles 3--5 cm. long, distinctly callose-connate at the base, te am ® ous—puncta 4 with scattered peltate glands on both surfaces, reticulate~vei ned; ec: es so » very densely incanous- or ferrug < or appressed-pubescent, the margin glabrescent, resinous~ punctate on bo: surfaces, the rim very shortly -dentate or ~denticulate to subtruncate: pi te both surfaces, its t Sed Wes Wee, 1 T/8 20 these is léce os the lobes hk, 1.5~-2 Lo: , 1--1,2 mm. wide, forn, oblong, obtuse ad & ddle, op 6=<- ; . tate; style terete, filiform, mm, long, short—exserted; stigma peltate, obscurely lobed; wt globose, densely s te~pilose with white hairs and resinous- punctate, l-celled, the cells 2-ovulate; fruiting-calyx scarcely 3.5 j Sruit drupaceous, rather large, depressed-globose, 2-9— -? Um. wide, dark-purple or black when mature, sparsely stell ape tes seed L--8, laterally compressed~ e type of this was not clearly desigmted by 8 zen van den Brink (1921) when he f and diy - 1967 Moldenke, Monograph of Callicarpa 2h3 ~ epee which Lam 529) had merely taken up in synonymy. How- , it seems clear that sheet no. 17613 in the gecdergtac ip We an un ted collector, is intended to be the type, and I have so desig- nated it [=lectotype]. Callicarpa backeriana is based on an un- numbered collection made by Cornelis Andries Backer —- in wh honor it was named -- at Bandjar, Java, on December 27, 1910, “and is = deposited in the Herbarium tm Bogoriens nse at Buitenzo Lam (1919) regarded this taxon as a mere re variety of C. erio- characters: "folia chartacea, ovata oblong: ta, i cune- ata, plerumque subrotundata, serupte a attenuata, apice acuminata, , “Bete basin, grosse dentata, den 8 utrin petiolo 0.73.2 cM. longo." He based his var. latifolia o Forbes 1355 and De Vriese Son. (Le—908.267-1091, Le—908 .267- 1092) from Java. He gives as the basis for C. inaequalis "3 spec- Herb. Var. botan. Sey Acad. Lugd.-Bat., in He L.-B. sub nos. 208-265-110), 1110 and 1446." @ species has been found growing in light forests and is said to bloom all all through the year at altitudes of 50 to 500 meters. The flowers are described as "pink" on Iboet 221. The leaf-blades are merely denticulate on Ltraing & Jochems 7427, and this specimen d here. = all, 37 herbarium specimens, including type ma material of all re involved, and 2 mounted photographs have been examined picmecte ie : GREATER SUNDA ISLANDS: Celebes: Zijll de soy ‘el [Bose bouwproefst. bb.20215] (Bz--18703). Java: Backer (Bz--17625, poe ), 33426 (Bz--17627, Bz—17628), sn. (Bandjar , 27.12.1910] (Bz—25), ah Bakhuizen van den Brink 1798 (B2—-17617), 1833 (Bu—17619, Bz—-17620, Bz--17621, 1, Ba-—17622, Ca— 235073, Ca—205055, pense Ut—25)68, Ut—-6700h) , 5933 (Ba-— 17623, B2—1762,, Ca——23661h, N, Ut—80253), 7209 (B, Ba—1761h, Be-—~17615); Collector tate s.n. [Tjiampea, Bogor] (Bz— 17612—isotype » Bz-—17613—type, N—photo of tyie Cee of tee); d!Arnaud Gerkens 361 aoe 217618, Ca—23199k) 5 7 Rei jnvaan 7691 (Bz—17616, Ca—301563); 3 Van pare rey, (B2—17610, Bz—17611, N). Sumatra: Iboet 2 2 (Bz—17629, Bo— 17830); j Loring & & Jochems 7427 (Bz—17631, ia. seein INTEGERRIMA Champ. ex Benth. in Hook., Journ. Bot. & Kew Gard. eats 5: 135. 1853. n emended synonymy: Callicarpa integrifolia epee *% ri & Hens J Journ. Linn. eps Lond. Bot. 26: ae mi So q ° Ty Prei, Mem, Sei. Soc. China 1 oo "Seber China] 22, in syn. Calticarpa tomentosa Bakh. (in part) apud P Pei, Mem. Sci. 2h) PHYTOLOGIA Vol. 14, no. Soc. China 1 (3): [Verbenac, China] 22, in syn. 1932 [not C. to- mentosa Auct., 1962, nor Hook. & Arn., 1918, nor Ktnig, 1893, nor ise 1959, nor "L. [ex Spreng.]", 1825, nor "L. [ex mild.]*, 1966, r (L.) Murr,, 1774, nor Lam., 1783, nor Murr., 177), nor Thunb, 1959 » nor Vahl, 1794, nor Willd. 1868, nor sensu uetave 1906}. Callic errima Lindl. ex on nl in Fedde, Repert. Spec. Nov. 40: 107, in syn. 1936; Prelim. Alph. List Invalid Names ll, in syn. 1940. Callicarpa integerrima var. diffusa P'ei ex ae ke, Résumé Suppl. 3: 30, in syn. 1962. Bibliography: J. A. Murr, in L., Syst. Mee a 13, 130. 1775 Jacq., Select. Stirp. Amer. Hist. Picta, ed. 2 the 2595 ve ge 0 » Bull. Sci. ; Maxim., “at Biol. 12: . 1886; Forbes & Hemsl., Journ. Linn. Soc. Lond. Bot. 26: 353. 1890; Jacks, in Hook. f. & Jacks., Ind. Kew, 1: 386. 1893; Dunn & Tutcher, K ew Bull. — heey ot Ser. 10: 202. 1912; Balin, Bull. Jard,. Bots Buitenz., 21. a H. H. Chung, e Sel. Soc. China 1 (1): 36. 1 g2hs Pies, . diffusa was collected by Rem t no. as) in the reeion of Kingyuan, at an alti a z kiang, China, in August or oat La pnt and is deposited i an United States National Herbarium at Washiné It should be noted here that the species is reduced to synom 1967 Moldenke, Monograph of Callicarpa 2h5 under C. tomentosa (L.) Murr. by Bakhuizen van den Brink (1921). The C, tomentosa of Bakhuizen van den Brink is in part C. integer- rima and in C. arborea Roxb.; that ascribed to "Auct.", to Hooker & Arnott, to Willdenow, and "sensu Matsum." is C. loureiri Hook. & Arn.; that ascribed to Ktnig is C. macrophylla Vahl; that ascribed to Linnaeus is C. erioclona Schau. » but the C. tomentosa L. [ex Spreng.] is C. candicans (Burm. f.) Hochr. as is also c. tomentosa Lam,; C. tomentosa (L.) Murr. is a valid species, with the homonym ascribed to Murray along and to "L. [ex Willd.]" as — ee es =ntegrifolia of Jacquin is Aegiphila integrifolia (Jacq.) Jacks, The flowers of C. integerrima have been described as "white" on TZ. W- Tsang 21107, “pink” on W. T. Tsang 2391h, and "purplish- red® on W. T. Tsang 21650. The vernacular names "lo hai ngan" "lo hai ngan muk" have been recorded for it. Champion's original (1853) description of the species is worth repeating here in view of differences of opinion on the validity m: mis folia subaequantibus, floribus parvis mumerosissimis, calyce lanato minute l-dentato. — Folia 3—h poll. longa, 2 — 2 1/2 eo etie nt t® (inferiora mihi desunt), supra viridia v. subflaves- the 32 Mollia, punctis glandulosis inconspicuis sub lana recondi- 138, petiolo semipollicari. e e, laxae, ak ; 1/2-pollicari, ramis primariis elongatis. Flores magnitudine Ss Mas iiae. Hongkong, The broad entire leaves and the dense golden-coloured tomentun readily distinguish this species Her the C. macrophylla, C. lanata, and others to which it is al- Pret (1932) amplifies the description as follows: "A shrub of about 10 feet tall; with branches, branchlets, petioles and in- florescences clothed with a dense floccose golden-colored tomen- tum or wool, Leaves broadly ovate-oblong, to nearly oblong or o- te to “te, shortly acumina obtuse ro o 14 cm. long, RE: O cm, wide, ckly chartaceous, entire or slightly te, glabrous or nearly so above, dense os pe ‘ ae Golden colored beneath, lateral nerves 7 or 8 on each side o: exceedj gla ng the stamens. Ovary densely pubescent. Frui via ing Subglabrous at maturity, 2 to 3 mm. in diameter, subglo- 26 PHYTOLOGIA Vol. 14, no. & Material has been misidentified and distributed in herbaria as C. arborea + and as C. siongsaiensis Metc. ~~ Forbes & Hemsley (1890) cite Millett s.n. from somewhere in China and Champion s.n., C. Ford s.n., § S.n., and C. Wright s.n. from ongkong. Maximowicz (1886) cites the same Ford and Wright col- lections and adds Bentham s.ne from southern China and Forbes Sn. from Hongkong. P'ei (1932) cites the same Ford and Wright col- lections and adds Ching 2125 from Chekiang and Sargent s.n. [Nov. 5, 1903] from Hongkong. The Ford and Wright collections are often ref d to as topotypes of the species. ie coer 28 herbarium specimens and 2 mounted photographs have by me. Addi tional & emended citations: CHINA: Che easgts Re © #3 ea 2h25 (Ca—261468, N, W—12i72s6). Kiangsi: S. K. Lau a al, 1752793). Kwangsi: Re ge Ching 6993 (N), 8034 (N); W. T. rang 2394 (N). Kwangtung: W. T. Tsang 21107 (Bz—-17633, C a—1118, » N, S), 21650 (can aaziee I, N, N, S). Province ear i, hairy S.n, sn, [Wongneuctung] (N) . HONGKONG: C. Ford s sen. [Hong- kong] (N); C. Wright s.n. [Hong Kong] (T, w—L907)- CULTIVATED: New York: N. Taylor ‘g.n. [New York Bot. Gard. Cult. Pl. 1s2hh] (Mi--photo) . CALLICARPA ae var, SERRULATA Li, Journ. Arnold Arb. 253 ene pag . Arnold Arb. ab, Fe) 4,25. 194; Moldenke, Piytologia a3 oldenke, Known Geogr. Distrib. Verbenace, ne voll ; ar. Eag Mcldenke. "Résumé 168 & ly. 19595 Moldenke, Fees Bg ewe oe the typical form of the species in having the seek Sinies serrulate along the margins. e type of the variety was collected by Wai-Tak Tsang (n0- 25228) at Ch'an Woh Tung village, T! district, Sam Kok by m ns Recract HONGKONG: C. Wright s.n. [Hong Kong] (W—91L)- oo pce Merr., Journ. Malay. Br. Roy. Asiat. Soc. 1+ Synonymy: Callicarpa involucrata f. clemensae Bakh. ex Moldenkés Résumé wt, aye. 1988S pees” 32. e54 ae will, Tad, few, Kew. ee Re sas Me a te 145 & 177. 1949; Moldenke, Alph. Lis * Gat. 3 k: al ¢ 82, af 525 Ul , 243, 1967 Moldenke, Monograph of Callicarpa 2h7 y glabrous shrub, about 3 m - tall; branchlets obscurely tetragon, the “ast aa ones sometines minutely puberulent, the ul ameter; leaves decussate-opposi te; fected 1.5=-2 om. tomes septs chartaceous, subolivaceous, poh Racer ahene a cm. long, 8-~1) cm. wide, narrowed to the s Be F ie F & to the bot base, shiny, glabrous and conspicuously pitted~g homme on th piece es, with a few widely scattered = and sessile the upper surface with numerous similar crowded glands at i very meees in- Po age aig cauline, sparingly pubescent, sessile during anthesis 23.5 cm, Wide, the flowers dense ely crowded, fascicled or de- Pauperate-cymose, subtended by several suborbic mm with a few distant discoid glands near the rim; corolla leben ort or cream-white, its tube 5 mm. long, glabrous, the teen rbicular-ovate, subequal, about 3 mm, wide, rounded or ob- 8 at the apex; stamens ; anthers oblong, 3 mm. long, slightly 1395) on forested slopes at low altitudes, Batu Lima, near Sanda- sh Nort to ascertain if or are it was formally described and published te from all ‘herto described forms, but somewhat resembling Callicarpa cauli- or , or as a 1—12 "recumbent peepee Hg S—-20 fick tall, the trunk inches : in diameter, buds cream-colored, flowers borne on the trunk, 248 PHYTOLOGIA Vol. 14, no. and fruit light-green or green to cream-green or cream-colored whe nally red 29965, "milk-white" on Endert 3651 and "white" on Boden-Kloss 19173, Clemens & Clemens 3 27651 & 2 & 28285, Pascual 1090, and M. Ramos 1395 & 1927. ~ Glemens & Clemens 27651 has a secondary fungus on the scale in- Kloss 19173 Merrill comments: "in type the infl. (anthesis) is is sessile. e axis elongates as the fls. are produced. See C. woodii Merr. which is probably involucrata." ~~ Material has been misidentified and distributed in herbaria In all, 35 herbarium specimens, including the types of all the names involved, and as pecethe Me ig have been examined by me- (A, Bu--72602), 3836 (Bz ae N); Pascual 1090 (N)5 Me Ramos 1395 [field no. 32h] (Bz—1763%—isotype, N--photo of type, Ph— Z—=photo of type), 1927 (Bz--1763h, N, N--photo, Ph, Z— IS Masam., Trans. Nat. Hist. Soc. Formos. 25: 19353 A. W 0, seg iy - Hist. Soc. Formos. est 25h. poh As isn: eg pl. 9: 5. 1938; Moldenke, a 13 a9 Molden, Sa a1 tuk i559; Moitonke, RSSt- : 2. branches spreading roeeally, ashy-gray, glabrate, scarcely lenticellate; branchlets dar’ Stata toseatenal leaves decussa Rta: peisaten coset 3 3 sn long, fuscous, steLlate- BE blades dots when dry; cymes axillary, their branches slender, stel- late-pubescent, and bracteate; a gs ovate, 1—1.5 m. long} fruit 1967 Moldenke, Monograph of Callicarpa 2h9 globose, 3--l, mm . The type oe this. species oe pstyperen by Genkei Masamune island o omote slands, on July 2, 1933. Recent collectors have found in the shade of large i the species growing trees, in dense low scrubby forests, in wet gulch bot "occasional" in ravines at the sides of wet wooded guiches, at 120 to 175 meters altitude, fruiting in August. In all, only 3 herbarium specimens have ses oe by me. ND ARCHIPELAGO: Sakishima Islands: Iriomote: Walker & Tawada Con oF Miyako: F. R. Fosberg 37191 (Z), 37289 (Sm 1a CALLICARPA JAPONICA Thunb., Fl. Jap. 60-61 [as acs ca"). 178h3 3 ja- J. A. Murr, in L., Syst. Veg., o 14, 153. mica Hort. ex rritzel , 4066, r "Thunb. auct." 3 Additional & emended srpiones Callicarpa fol. serratis gla- apud J. 153, i natis glabrie Willd., Linn. Sp. Pl. 1: 621, in syn. 1797. Callicarpa (Japonica) foliis serratis glabris Thunb. apud Willd., Linn. Sp. Pl. 1: 621, in syn. 1797. Callicarpus japonica Thunb. apud Hassk., Cat. Pl. Hort. Bot. Bogor. Alt. 136. 164k. Callicarpus Hort. apud Hassk., Cat. Pl. Hort. Bot. Bogor. Alt. 1%, Maatsch. Tuinb A meee Pays—Bas] 18h: 25, nom. md. 18h5 Sieb, & oes Math.-phys. Cl. Ktngl. Baier. Akad. Wiss. rary h (3) Gi 356.3 1846. Callicarpa mimurasaki Hassk. apud Sieb. ° oe Baier. Akad. Wiss. Mttnch- % “ h (33: 6 terece rage bee “ie pinaeeete japonica f. olia Miq., Cat. Mus. eng Poe ee 70, nom. md. 1870. Calli longifolia var, japonica (Thunb.) Kuntze, Rev. Gen. Pl. 2: 03. 1891. Callic carpa mimurazaki Hassk. apud Jacks. in Hook. f. & Jacks., Ind. Kew. 1: 306, in syn. 2893+ Callic aponica var. ca H. J, Lam, Verbenac. Malay. . 1919. Callicarpa Japonica var. typica Bakh., Bull. aakas ‘Bot. Buitenz., Sere yf 3 25. 1921. Callicarpa a Nakai, Fl. Sylv. Kor. 1h: 30, Syn. 1923 [not C. pres. Hort. ex Lem., 1859, nor Hort. ex Yel- » 1941, nor A. L. Juss., 1806, nor Van Houtte, 1932]. Calli- carpa japonica f. clabre P pai: Mem. Sci. Soc. China 1 (3): [Ver- carpa dichotoma var. koreana Hort. ex ee t Alph. List Invalid Names 10, in syn. 1940. Callicarpa shirasé 250 PHYTOLOGIA Vol. 1h, no. k Hort. ex Moldenke, Prelim. Alph. List Invalid Names 13, in syn. 1940 [not xC, shirasawana Mak., 1910]. Callic japonica & subglabra Schau. ex Moldenke, Résumé 2h, in syn. 1959. Callicarpa Japonica var. japoniica Kobayashi ex Moldenke, Résumé Suppl. 3: 30, in syn. 1962. Callicarpa japonica L. f. ex Moidenke, Résumé Suppl. 1h: 7, in syn. 1966. : sen Nov. te Soc. Sci. Upsal. : 31 1783; Thunb., Fl. ° . J Murr L., Syst. Veg-, ed. 1h, 153. 178k; Jacq., Ind « Syst., ed. 1h, 32. 17855 A 1. 1: 307. 1789; Lam., Tabl. Encycl. Méth. [Illustr. Gen.] 1: 293. 1791; J. F. Gmel., t. Nat., ed. 13, 2: 2h6. 1791 » Symb. Bot. 3: 14. 1794; Willd., ° Pl. 1: 621. 1797; Raeusch,, Nom. 37. 1797; Murr. & Pers., Linn. Veg 215 [ eo], 159. 17973 J. Ae Syst. Veg-, » Nom. Bot rs., Syn 343. 1842; Hassk., Cat. Pl. Hort. Bot. Bogor. Alt. 136. 18hh Sieb., Jaarb. Konink. Nederl. Maatsch. Tuinb. [Ann. 18h: 25 (18h) and 18h5: 71 & 72, pl. 5 & 6. 1845; Walp., Repert- h: 129. 845; Sieb. & Zuce., Abhand, Math.-phys. Cl. KUngl. Baier. Maxim., Mél. Biol. 12: 50h & 508—~S09. 1886, G. W. Johnson, Gard. Dict, 891. 1890; Forbes & Hemsl., ot tim, Soc. Lond. Bot. 26 (Ind. Fl. Sin. 2]: 252 & 253, 1890; P. Henderson, Handb. Pl-s ed. 2, 6h. 1890; Kuntze, Rev. Gen. Pl. 2: 503, 1891; Jacks. in Hook. £. & Jacks., Ind. Kew, 1: 386. 1893; Bois, Dict. Hort. 1+ 32. 18 . Agri d Imp i Engl., Bot. Jahrb. 3h, Beibl. 75: O-- & Mill... Cycl. Am. Hort. 1: 217. 19063 We Ps » Dict. Pract, Gard., ed. 2, 156. 1907; snirasaré, 1967 Moldenke, Monograph of Callicarpa 251 Nippon Shinrin Jumoku Dzufu [Icon. Ess. Forest. Jap.] 2: pl. 70, aes 1—10. 1908; Nakai, Fl. Kor. 2: ¥ 1909; P. Henderson, we ie; s U Handb, Laubholzk. 2: 591-59, $m ge aes & 385h—-1. 1911; Hay- ata, Veget. Mt. Fuji 67. 1911; J. Matsum., Ind. Pl. Jap. 2 (2): 529. 1912; Hickel, bull. Soc. Dendrol. France 28: 111. 19135 A. H. Graves, Winter Key Woody Pl. 8 & 30. 191.35 Rehd. in L. H. Bailey Stand. Cycl. Hort, 2 628. 191h; Nakai, Fl. Quelp. ealns "16. 1918; Nakai, Fl. Mt. Chirisan )3. 1915; Rehd. in C. S. Sarg., 3: 367, 368, & 370. 1916; Nakai, Rep. Veg. Diamond Mts. 183. Ts ni 3 x - Bur. Agr. Co. 12: (Prelim. Rep. Fl. Tsing-tao] 96. “i993 H. J. Lam, Malay. Arch. 51, 84--85, & 362. 1919; Bakh., Bull. Jard. "Bot. Buit er. 3, 3: DL & 2 . 1921; Nakai, Fl. Sylv. Kor. 1h: 30-32 & 133, pl. 6—8. 19235 Olmstead, Coville, & Kelsey, Stand. Pl. Names, ed. 1, 61. 192k; W. Trelease, Wint. Bot., ed. 2, 333. ) Be ° - a oe 1925 5Wa J. an, Trees & Shrubs rit. Isles, ed. lL, og 1925; Rehd., Man t. Trees, ed. 1 & 776. 192753 oud, Cult, 120 & 121. 192 akai, Trees & Shrub Pheme a Icon. Bot. Ind. Lond. 1: 526. 1929; W. J. Bean Hardy Brit. Isles, ed. 5, 1: 280. 1929; L. H. & Bs 2: in Zytol. Unte Self, Sanders Enc 1 yel. Gard., ed. 21, pr. 2, 71. 19 Pa - Bot. Upsal. : 81 & 82, fig. 127. 193h5 Grevost t Pételot, - Econom. Indo-chine 37: 1290. 193; Hand.-Mazz., Ann. Ho > Fedde ert. Spec. Nov. 39: 297 » & (1936) and 40: 8h--92. 9 36, 38s jo2" 1s, 120 2212s, & 130. Sanders Mag. T rele Ap 52. 1937; eg ee Book Sy ed. 3, 168. 1937; Meldenke, Cult. Pl. 35. 19383 A. W. Hill, Ind. ew. S . January nets 1939; M 6 og asamune, Trans. Nat. ° 19405 Rehd., Man. Cult. Trees, ed. 2, a re 80h. “15405 252 PEITOLOGILA Vol. 1h, no. Moldenke, Prelim. Alph. List Invalid Names 10-~13. 1910; Doney, Brooklyn Bot. Gard. Record 30: 23. 191; Worsdell, I nd. hrw ° Fe ‘s $ Q a) o ak Be 36 . Encycl. Gard., ed. 21, pr. h, nl. 192; Mold enke, Known Geogr. po SeePenens, ed. 1, 56—59, 61, 62, Ti, &'87. 1942; Molden- Alph. Li st Invalid Names ees Se 19425 Wisler, rthmore Pl. 7 , 1081, 1ioh, 1115, 1%, 3 69, 122h, 1253, 125), & 1289. 1949; Moldenke, P _— 3: 78 & 139. 1949; Moldenke, Known Geogr. Distrib. Verbenac. ed. 2, 7, uaeee 133-135, 140, 157, & 177. 1949; Matsum., List Seed Exch. 2.°1949; E. D. Merr., Ind. Raf. 20). 1949; Rehd., Bibl. Cult. r 123. —_ Darlington & Wylie, Chromosome Atl., pr. 1, 323 & 503. 2 53 Spector, Handb, Biolog. Data 12. "1956; lattcon = — » Od. 6, 83. 1958; J. & L. Bush-Brown, Am. Gard. Book, ati Res” kSr thes “ 92 935. 1958 at. + 40: 77. 1958; Moldenke, Am. t. 592 335. 19595 — Résuné 9, 168, 171—173, 17s, 8, 83, 193, 213, 2h2- » 427, & hhh. 19595 Ha Hara, Outline P Phytogeog. Japan 7. 13595 , : : 253-265. 1969; il sae, Suppl. 1s 12 & 13. 19595 ied. » Man, aa me ed. 2, pre 9 2 6.4 & 932. 1960; T. H. Everett, New Illuste. Bnoyel | may 503. 19 sas ae brn ig & Wylie, Chromosome Atl., pr. 2, 6 sumé Suppl e 4 od 2 is (562), 3 st rs (2962), and 7: 7, 1963) = 25, 38" 96 wg : . 30, & 376. 1963; Li, Morris Arb. Bull. 1+ 1967 Moldenke, Monograph of Callicarpa 253 ee 2: 601. Stig: Santamour, Morris Arb, Bull. 16: 52. 1965; Ne ts. 0 ie a sec, 2. ce a od es peek Mol- re > Rhowé Supp : 5 (1966) and 1966; Mol- ologia 2 $ 2 (1966) and 101, U2, 456 Ist 33: «ite ses ) ahs 13, $3, 4, 51, 39, tihetesr icy Ay Jaarb. Konink, Nederl. Maatsch. Tuinb [Am. Hort. Pays-Das] ibuss pl. 5 & 6 (in color] - 185; Lindl. & Paxt. in Paxt., Flow. Gard. 2: 165, fig. 221. 1853; Hérinca, Hort. Frang., ser. 2? : pl. h. 18615 M. T. hens, Gard. Chron. 1871: 173, fig. 39. 1871; Shirasawa, Bull. Goll. Are - Tokyo Imp. Univ. 2: pl. lk, fig. 10. 1895; Usef. Pl. Jap. - 656 [in color]. ndrol. Winte: Nippon Shinrin Jumoku Dzufu [Icon. Ess. Forest. Jap.) 2: a 70, fig. 1—1o (in color]. 1908; C. K. Schneid., Illustr. Lenthal 2: fig. 3le--e & 365h--1. 19125 Rehd in L. He Bailey, Stand. Hort. 21 628. 191s G. V. Nash, Addisonia 3: pl. ae [in (in ester]. }. 29285 a vbgee Kor. 1h: pl. 6. 19235; Nakai, ee — Jap., ed. 2, 1: 452, fig. 21h. 1 esa tee owa Acad. Sei. 35: 11% fig. 32. 1928; Schwencke, Zytol. Untersuch. Verbenac. fig. 12--14. 1931; Junell, Symb. Bot. 3 Li, Mor ; Akt h & 6. Ste lee 19645 . Recent collectors describe this as a 8 r low shrub, 0.6—- er 1--1.7 m. sp and moderate relative g buds lavender, the flowers fragrant or odorless, and the fruit ge green Sige ture, then lilac, deep-lavender, or purple en ripe. number is gi 2n = 16 or 18 [or are described pig as Mhaploia ein cecne 2 eer 188 ")]. The flowers as "lilac" on F, R. Fosberg 38215 & 38546, "deep-pink" on Dorsett & Morse 787, "pinkish-lavender" on F. R. Fosberg 38180, mreddish- Purple" on on Ch hung & Sun 20h, "purplish" on Hurusawa ra lll, "light- Purple" on Takenchi 1)-C, "purple" on Chun & Sun 176 & 106, Hiroe 13906, » Is Kimura 1. 1j-B, Suzuki UC.699, E. H. Wilson m 10422, | and Yin 102, and “white? on Chiao 2617. ‘ ~The Plant has been found by recent pniscng-tcnn growing in shade r half-shade, in clay soil or in hums on mountainsides, in woods, an at rock cliffs, on es ina "spike", I feel ae aD these statements are errors in observa t 7 rts the species as rvation or terminol . Wilson repo ais i. ogy + “common in thickets", while Aedes "Sate it to be 25h EETTOLTOG Tx Vol. 14, no. the undergrowth of scrub forests, in Pandanus scrub on limestone, in graciagh between cultivated seChak ead » and in rocky grassland with shrubs on the Ryukyu Islands. It should : be noted here that the C. j Japoni ca of ce ex Prit- zel", referred to in the synonymy above, is a synonym of C. rubel- ia Lindl., while that on "Thunb, auct. ex Raf." is C. ies Lam. The C. purpurea of A. L. Jussieu is C. dichotoma (Lour.) K. Koch, that of "Hort. ex Moldenke" is Cc. longifolia Lam., while that ascribed to Van Houtte and to "Hort. ex Lem." is C. rubella L The C + japonica angustifolia Kwa-wi, C. = depont ca f. angustata (Rehd.) Mf Foote C. japonica f. angustata (Rehd.) Ohwi, and C. japonica are telate, Mattoon are all synonyms of C, japonica var. angustata Rehd., which see; C. japonica leucocarpa Aul, C. japon~ dea f. leucocarpa (Sieb.) Rehd., C. japonica var. mica var. leucocarpa C. japonica var. | var. leucocarpa Sieb., and "CG, japonica ® C. leuco- = Sieb." are all synonyms of C. japonica £. albibacca Hara; C. japonica var. angustifolia Savatier, C. japonica var. dichotoma Bakh., and C. japonica var. dichotoma (Lour.) Bakh. are C. dichot~ oma (Lour.) K. Koch; C. Ssponies f. latifolia Miq., C. japonica f. rugosior Miq., C. j japoni a subsp. luxurians (Rehd.) Masamune, and C. japonica var. kotoe oat (Hayata) Masamune are all synonyms of C. japonica var. luxurians Rehd.; C. japonica var. luxurians es ieucocarpa Nakai is a synonym of Ce: japonica f. albifructa ifructa Hara; =f epee Li Fhombifolia Miq. is C. japonica var. rhombifolia He J. Lamy Ce Japonica var, taquetii Nakai is a synonym of C. japon- eola Pm rous shrub, with a few stellate hairs when : aves glabrous, glandular beneath, ovate-lanceolate, crenately a-tibe as as Style subglabrous. Ovary subglabrous- Miguel (1870). arated § eae er specimen collected by Birger and one collected by siebold in Japan. 1967 Moldenke, Monograph of Callicarpa 255 Hara (1948) and — (1949) maintain that Amictonis japon- ca Raf. is a synonym of C. japonica Thunb., but I place it under C. lo longifolia Lam. Danae yan den Brink (1921) includes C. brevipes (Ben (Benth.) Hance, C. dichotoma (Lour.) Raeusch., C. elegans Hayek, C. glabra H. J. Lam, C. gracilis Sieb. & Zucc,, C. jama~ at Sieb., C. longifolia. var. rar. brevipes Benth., C. oligantha Merr., C. ea A. L. Juss., C. serrulata Zipp., C. sieboldii Zipp., and i, dichotoma Lour. in the overall synonymy of C. Japonica, albeit not in the typical form of the species (which he calls 0. C. japonica var. typica Bakh.), where he places only C. mi- murazaki Hassk, and C. murasaki Sieb. _ ‘The Baileys (1935), Moldenie (1936), and Salisbury ( (19h7) on C. arnoldiana Hort. as one oF we sro C. japoni it seems to me now that thie is r homonymous Tot nation for the C. arnoldiana of Kalear > 13932), gt iey in turn, is a name that clearly applies to the C. bodinieri var. giraldii (Hesse) Rehd. Ee eeeretness ae our country through the Arnold Arboretum. authors, including Nakai (1923), Moldenke (1936), and Hara ( (1948) saben ¢. longifolia var. _subglabrata Schau. (in part) Lam belongs in the synonymy of that whens regardless of what speci- mens he cited as representing it. Sprengel, on the other hand, in his 1825 work reduces C. japonica to synonymy under C- longifolia, but in his 1828 work he reinstates it as a valid species Lindley & Paxton (1853) make the following comments: ugiebold & Zuccarini have pointed out (Florae Japonicae sy EAE ater oF ) the error committed by M. Schauer i erring this species to C. longifolia, a still less attractive ee figured in the Botanical Register, t. now @ spasenty lost in gar fles not branous, calyx. C. Cs a ata plant, sts more a mag than this, which we believe occurs exclusively in Japan, whence we have wild specimens from Zuccarini, SS only in a loos and long er a and larger leave: = Stee Joponics was actually first snes by rg cana L.", a clear case of IS tae EEE later iy him. Nakai (192 ives its rep on vCorea media et austr. —o Ko ee a et Saeetet cus mame is 0 mulberry", "guiou-sa6-si ama-murasaki", ‘"jama mura saki", "Japanese’besutyberry”, i Sesent sat Schtinbeere" , "ko-mourassaici", 256 PHYTOLOGIA Vol. 1h, no. h "méka-sogui", "mimurasaki", "mi-murasaki", "mi mura saki", "mi- murasaky", "mi-mura-saki", mourassaki-skibou", “murasaki*, “mura saki", "murasakishikibu", "murasaki-shikibu", tmurasakisikibu", "murasaki~sikibu", "o murasaki-shikibu", "passalnitenaa” "tama- mourassaki", "tama-murasaki", nyabu-murasaki", d "yanamurasaki". Of these, "French mulberry" is applied also ate the other or eeu species of the at "Japanese penittenae is the rec d common name in Rehder (1940) and Bean (1921) state that C. japonica was intro- oe into cultivation in 185 [when Siebold introduced it into at pc mae Ay aoe also gives the same date, but Van elle (19 tains that it was introduced as early as 181. ecu tne lists only 6 horticultural sources, but Bailey (1935) rts that at that time it was offered to the horticultural trade by H. P. Kelsey, Sanford, Brooklyn Botanic Garden, Katzen- stein, Kew, Knap Hill, Eastern. Nurseries (Holliston, Massachusetts), cultive- tion in Greak Neck, Nassau County, New York, on November 20, 193" but did not make any collection of it. The New York Times ie in spring." (1965) get "No explamation can be offered for ys r (x = 8, 9) [of the chromosomes } in the ploidy of C. 5 sponte red hx) However, the Verbenaceae ° OT contains many genera for which two basic numbers have been po even in the same species. The existence of diploid and ple forms in the same species is more common and be the case in Us ra) her chers (if any were tate the specimens on which this cyto- ; _ very widespread misidentification in port on or apringgante Studies of this plant and illustrate the 4 arance ngs. Gressitt 532 is anomalous, two Kinashi s.n- collections have the leaves gray me under C. di- eS hairy snp is cited by me ves ertainly is not that species! The E. D. Merrill Lue, ao cited by me under Cc, dichotoma, may likewise prove oa represent C, Jponice instead. “\sushina & Mort (1958) report : : + lumurians Rehd. is often found near the eashore" on the Shinoki ta Pestees Peninsula, Japan, Pry TOLOGIA Designed to expedite botanical publication ———————. o Vol. 14 February, 1967 No. 5 CONTENTS WURDACK, J. J., Certamen Melastomataceis XI. ....-.. + + 257 MOLDENKE, H. N., Additional notes on the genus Verbena. Ill . . 275 MOLDENKE, H. N., Additional notes on the genus Avicennia. |. . 301 MOLDENKE, A. L:, Book reviews 2 Sp es we | - Published by Harold N. Moldenke and Alma Lenk a = a 303 Parkside Beat ae Plainfield, New Jerer 9 07060 U, f CERTAMEN MELASTOMATACETIS XI. John J. Wurdack Dept. of Botany, U. S. National Museum In continuing the routine adjustments needed in the > eae for the Flora de Venezuela, some long-delayed delenit ies. of capsular-fruited tribes of the femily has been essentially completed (which a vot construe as evi ce that e work of th t ars has not been mere logom ys fortunately for my innate des leisure, the Miconieae are bar be The usual distractive inflow of collections from elsewhere in tropical America has not ceased, 50 her Colombian and Peruvian novelties have been included here; cytologic leverage has even extruded some Mexican nsi ost o Hutchison’s superb Peruvian collections have been given formal status; the material was ges dag, in 1963-1964 during the Seventh Botanical Expedition to the Andes of the University of California 8 an n 8 & ct 3 c) | ® 2 oO kK wa hs YO n 2) g B e BUCQUETIA VERNICOSA Glea Originally eter sos a rom Téchira (Venezuela) material, Acad Species has also been collected in Pet Santander, Colombi anga, St. John 20604; ea $ _ ole CUTERVOANUM Wurdack, sp. no Campanulari (Bonpl. ) Triana arrinis, sed fol pubesecnee subtus sparsiore, floribus in panicu a Sa Sesregatis, hypanthiis graciliter petalonie differt. ic hom ta laevia. Ramuli quadrangulati fet “petioli folia che afore centia hypanthiaque modice setulosi, P is SPice acuto ba e acuta si ea s(ot)-4 ne s Subtus manifestis et laxe reticulati 6, Bu Boanshn 1 modice et graciliter str FL h-meri nutantes in conte Plur rum s regati; pedice cere E 6 mm mox asatoudh thium zum) 3 2. 3; ecalycis tubus 0. 6-048 m altus, lob rea} x ae 3(H) a a xtus sparse strigulosis intus ieee 257 , 258 PHYTOLOGIA Vol. 14, no. 5 2.5-3 mm versus sparse strigulosis. Petala 15-15.5 X 8-9 mm subrhomboidea apice acuto ciliis caduce parang sae longis orn is 4 mm sum. Collection: A. Sag&stegui 5384 (holotype US 2469209; isotype TRP), collected ; at La Pucarilla along the road between Socota and San Andrés, Prov. Cutervo, Depto. Cajamarca, Peru, elev. jake - 2h May 1965. "Arbusto. Flores morada-oscuras, pedicels (ca. 1 mm diam.), and the hypanthia are densely loose- strigose; generally also, the older branchlet pubescenc appres and leaves are rigid rather than chartaceous. The quantitative pubescence distribution gives B.c te & P relative has narrowly lanceate = Soasenea’ glabrous sepals, s well as appressed OS a8 hai TIBOUCHINA HUTCHISONII I Wurdack, sp. Sect. Diotanthera. T. incarum Siac son et T. stenopetalae Cogn. affinis sed inflorescentia pauciflora pet talis maioribus differt. ge dichasiorum b -8-1.5 om is lanceatis persistent ibusi res 5-mer dicellis 2-3 mm longis sicut h hi glanduloso-setulosis, pilis ca. 1 m longis, bracteolis ca 1.5 is ellipticis persistentibus ca. 1 m infra nypanthit bases insertis. Hypanthium (ad torum) 4. 5-5 mm longum; calycis tubus 0.5 mm altus, lobis 2-2.5 mm longis pentormicngte ae rotundato) g glanduloso-ciliolatis intus glabris. Petala 15-21 X i — mm glandul rarum ecis Connectivis 0.6-0.8 vel 0.3-0.5 mm : basim interdum tuberculatis lobis ventralibus 0.3-0.5 mm longis hebetibus. St truncatum; stylus 13-14. oe - 0. 0:5 mt mm glaber; ovarium ice setulosum pilis P- p. glandul POS Collection: P. C. Hutchison z- i Straw 6034 (holotype US BOTs , teotypes U ae cohaectei on wet steep banks at lower Boqu rolongatis dorsaliter ad 1967 Wurdack, Certamen Melastomataceis 259 San a Peru, elev. 480 m, 25 July 1964. "Flowers lilac pink. t Para H. A. Allard 22102 (US), from ssa gn Pass between Rtg “Maria j and d Puca: allpa, San Martin, Peru, elev. 1.00 mM. "FL a rose-pink.” oe stenopetala has well-developed spe deir ip nh cater ah a on mm long) dichasial bracts 6 ee s 6-7 mm ong > ee with eglandular stout-based hairs, and pe ae aS mm ae the collections of T. hutchisonii, well-developed Heuecs of 6034 are meee! plinerved but all visible leaves e. 22102 are basally nerv TIBOUCHINA SANDIENSIS Wurdack, sp__ nov. 349) Sect ° hera. T. mollt ae ee ) Cogn. affinis, sed foliis flor busaue minoribus diff li obscure quadran ealatt Ap petioli folia subtus itl cccteciae hypanthiaque pilis laevibus erectis 0.2-0. (-0 >) mm longis basim versus paulo expansis non te = at ae -2-0.3 mm longis hebetibus. Stigma truncatum; stylus 9-5 X 0.35-0.2 mm glaber; ovarii apex sparsiuscule strigulosus, Pilis 0.15-0.4 mm longi Type Collection: os Ferreyra 1667] (holotype US 2h69214; isotype USM), collected in low forest below Sandia, Depto. Puno, Peru, elev. 1800-1900. m, 13 May 1966. "Sufruticosa; flores rado-purpureas mo: ratype: R. D. Metcalf 30606 (NY, wl ht near Oconeque, Sandia, Puno, Peru, elev. 1800-2100 m, 22-25 May 1942. “Perennial growing from cracks of moist shaded rock raat calyx red- en bouchina mollis has leaves 5-10 cm long, hypanthium 5-6 a long and about equalled by the lobes, and anthers 4-5 mm long; ea. ses 2. sandiensis. I have not studied any of the collections cited in the Flora of Peru as T. mollis, but feel sure that they are 260 PHYTOLOGIA Vol. 14, no. 5 all referable to other species. Tpouchine dimorph oph, yil Gleason eau pubescence similar to that much larg leaves and flowers as a as attenuate anthers. eee longifolia (Vahl) Baill. has longer and proportion- gayana (Naud. ) - and (ex char.) T. solmsii Cogn. have larger and Sree. at teniate anthers as well as vegetative difference TIBOUCHINA BREEDLOVEI Wurdac sp. nov - durangensi Standi. ah anter affinis, sed foliis minori- bus Dabs saetit aritie floribus 4-meris differt. Ramuli a cute quadrangulati sicut folia utrinque spar s strigulosi, pili laevibus 0.5-1 longis Petioli 3.0.5 em +3 -5-2.5 X 0.5-1 cm elliptica vel paulo ovato- elliptica utrinque acuta subcoriacea, tr a nervis secun- 6 em, ramulis pedicellis hypanthiisque modice glanduloso-setulosis, pedicellis ca. 3 mm longis, bracteolis ca. n 1 longis lanceatis caducis. Hypanthium (ad torum X 2.5 mm; calycis tubus 0.4 mm altus, lobis 2-2.2 X 1.7 mm tri ribus, extus glanduloso-setulosis, intus glabris. Petala 6.2-6.8 X 5-5.3 mm ovata, apice rotundato, versus s rse ciliol ta, pilis p. glanduliferis. Stamine dimorphica glabra, filamentis 6. 5.5 longis, thec oblongo-subulatis poro ventraliter inclinato 0.2 , s ecae 5 mm longae, connectivo 2.5 m pro- : longato, appendicibus Pe adapta 1.5 X 0.5 mm hebetibus. Stamina minora: thecae 3.8 mm longae, connectivo 0.3 mm prolongato, appendicibus verre! thus i 2 0.6 mm hebetibus. Stylus 7 X 0.5- 0.2 mm glaber; stigma punctiforme; ovarium 4-loculare, apice ipse e 6 mm te glanduliferis coronato. Semina 0,5 X.0,3 - ccochieata muriculata llection: D. E. Breedlove 10060 (holotype US eueseor; isotype DS), collected on a slope with Pinus and o de Montabello, 40 km ai: la Trinitaria, itari emg La Trin a, Chiapas, Mexico, elev. 1600 m, 27 May 965 - aang 8 purple; 3 feet tall. ratypes (both Se ee: Breedlove 9693 and Rzedowski ‘ibouchina d urangensis has leaf blades below the nee cence — X1.5-2.5 cm, these aah sparsely loose-strigulo above; the f rs are 5-merous ese ventral conane re. “lobes of the stamens are 1 X 0.7 m. T he e had qualms proper generic Sele ae of both rT. reeset Lovet d chtabensisi certainly b tive of Ovelty seems to belong, other than pode as ‘Bite flowers, in Tib ouchina Sect. Pseudo- s but rather with those 5-merous Mexican species with 1967 Wurdack, Certamen Melastomataceis 261 long ventral lobes at the connective base, such as Tf. galeottiana tik ) est < Pie an unpublished species of Gleason based on Orcutt 3939. the -merous flowers might also indicate that placing take n Pterolepis sh be considered, re no nched intercalycine hairs in either o se a br anomalous novelties and the stamens are quite different from those of Pterol lepis. TIBOUCHINA CHIAPENSIS Wurdack, sp. nov. T. breedlovei Wurdack distant er affinis, sed r hypanthiorumque pilis minute barbellatis tolls latioribus differt. pilis ubique sparse vel modice minuteque tae am lati sicut n ovata apice late acuto vel obtuso basi obt irme membranacea, Supra sparse vel modice strigulosa, subt ice strigu trinervata vis secundariis supra obscuris subtus elevatis nervulis subtus laxe reticulatis. epee Hs sub- multiflora ca. 4 em longa lataque sicut pedicelli hypant Sepalaque cppresso-setulosa pilis 0.5-1 mm longis pe crretlets flores 4-meri, pedicellis 3-5 mm Seen practeolis ca. 3 m longis ieeetins at open caduc Hypanthium 4-4.5 mm longum; Calycis lobi 3-3-2 mm longi, bine oblongo-triangulares per- Sistentes. Petala 8.5 X 7 mm suborbicularia densiuscule glandu- ina hi loso-ciliolata ut videtur rosea. m imorphica glabra; filamen vel 6.5 mm longa, antheris oblongo-subulati Stamina maiora: thecae 5-5-5 longae ut videtur eae, connectivo 3-3.5 mm prolongato, appendicibus ventralibus 1.8-2 X 0.3-0.4 nm hebetibus. Stamina minora: thecae 3-8 m longae ut videtur ia teal vs ona oe 0.6-1 mm prolongato, appendicibus ventralibus 1.2-1.6 X 0.3 mm hebetibus. Stylus 10 X 0.3 mm laber; stigma mig ere ovarium 4-loculare, apice modice ace “eae Parra: . (Matuda Herbarium Collection: Mac Do 26061) (nolotype Us HO cacy 1B -Monserrate, Chiapas, 0, 9 Marc ont 8: c. . Purpus 10200 (April, 1925) and 10627, segs G both from mountai eater, Fénix, Chiapas, Mexico. Tibouchina breedlovei has smooth hairs and le a il “yf 0.5-l cm. wide. The fruiting s collections had been £ the — Monochaetum but excluded from that genus by psec t i . — ose resting and _ foliage of T that species Suggestive of those of Monochaetum pulchrum Dene. ne ant be — nguished gre by the patent caul Pubes: TACHIRENSE Wurdack, SP- landii (Kunth) meet: afrind 8, sed caulis aa eubins pills appressis praeditis, sepalis ovato-lance fert. 262 PHYTOLOGIA Vol. 1h, no. 5 i paulo quadrangulati sicut petioli foliorum lamin ae complanata, apice hebeti-acuto Stamina minora: anthera thecae 2:53: 5 X 0.1-0.2 mm Lnéavee steriles, appendice pales 1.3-2 X 0.2-0.3 mm apice hebeti-acuto. Stylus 15 X 0.3 mm; Type C A VEN 10545; isotype US), fgeenciter in wet thickets between El Cobre Venezuela, elev. 1520-1820 m. 8 July © a Q 5, mo) R Ang i ° . E 194, 5 fee Sony, leaves deep green above, silvery below; petals deep rose-orchid as are fil ts and style. at dubious as to the exact relationship owe ioieiese, as well as in the sede mn sepals; M. meric has mu uc sepals and larger petal MONOCHAETUM VILLOSUM Gleason subsp. VENEZUELENSE Wurdack, subsP- nov. peo Hypanthium glabrum; calycis lobi ciliati; tori regio extus ra. Type Collection: F. J. Breteler 3370 cute ME; isotypes US, VEN), collected in a2 shade along road to Barinas 79 km eas Mér (apparently ca. Estad skoda aestnes red; leaves herbaceous, slightly glossy and medium green above, below. EB green. Corolla white. Filaments e typical subspecies of M. villosum, still known only from the Colombian Ginite) type collection, has evenly fine-setose thia with a zone of setae at the torus line. The other 1967 Wurdack, Certamen melastomataceis 263 Venezuelan species of Gleason's group Dicranantherae, M. jahnii Pittier, resembles M. villosum subsp. venezuelense in its glabrous hypanthia, but has a dense zone of toral hairs external- & glabrous ovary. Bretele a distinct cr ° gland-tipped ovarial hairs up to about 1 on, his origina description of M. villosum, Gleason noted that the hypanthial hairs in part were caducously gland-tipped; his ke treatment of the South American species is therefore misleading. Perhaps M. villosum will not be maintained as spec 11 distinct from M glanduliferum Tr when more collections are ’ iana available; for the present, however, the eglandular vegetative hairs and longer cauline pubescence can serve to separate th Huila species from the earlier-described one CARDONAE Wurdack, sp. Sect. Anisostemones. A scieakabiin “37419 Monographiae beethnees: differt foliis supra sparsissime gracili-setosis oe LS « rulata, dentibus ca. 8 per cm et 0.3-0.8 mm altis setuliferis. Inflor- rescentia anguste thyrsoideo-paniculata foliolosa usque ad 23 cm longa, ramis brevibus plerumque 3-4 cm longis bracteosis ca. d-floris; flores breviter (1-2 mn) pedicellati. ium (ad torum) 5.7-6 mm longum sicut sepala dense glanduloso-setulosum Pilis gracilibus 0.6-1 (in calyce ad 2) mm longis; saat tubus ltu 1 llection: F. 8 (ho ase VEN; isotype o Paragua, Type Co F. Card 0 NY), collected "en playa, Salto Rfo a en Edo. Bolivar, Vi enezuela, elev. 380 m, 1 March fe @ sf s ted relatives have the upper es 26h PY 0:8 602i Vol. 1h, no. 5 her species placed in this alliance by recent describers are i an forme with "subsetaceous"” calyx Amon group characterized by Cogniaux as subeglandulose, R. verbenoides Cham - differs (ex char.) in the stout petioles, rigid and re tively narrower leaves, and Fcuneininenat calyx lobes. deg £1 imensions given for R. cardonae are f a low $k just before erent thus the filaments and style are so somewhat imma RHYNCHANTHERA Wurdack, sp. Vv Sect. Se ke ag R. se Sere eee Naud., R. leucorrhizae S. Moore, et R. cacerensi Hoehne affinis, differt floribus ’ minoribus. quadrangulati sicut inflorescentiae hypanthiaque Os uli modice glanduloso-setulosi, pilis gracili ibus 5-1 mm longis. longis foliolosis secundifloris; flores sin pedicellis ca- 0.5 in fructifero ca. 1) mm longis. Hypanthium (ad torum mm 3 calycis tubus 0.3 mm altus, lobis 2-2.2 (in statu fructifero ca. 2. 7) densiuscule Hackl ese pete Petals T-T- obovata, apice late acuto et glan i i expansum O67 mm shane ovarium 3-loculare, apice inconspicue ani sbcpterp Pilis 0.1-0.2 mm longis. Type Collection: Leandro Aristeguieta & Ge tulio Agostini — (holotype US 232 L167; “isotype VEN), ool f the Se de San Camilo, between Guacas and Guasdualito, Ho. pease ener is, March 1960. "Planta fruticosa, alrededor 1 ™ alta. moradas All c relatives (ex char.) have petals 13-25 mm long an¢ anther sin Hd 3- Ong- Rhynchanthera secundiflora, from y, has Puree calyx lobes about as long (4-5 mm) as the hypanthia. o Brazilian relatives, with anther rostra about om th SBo Luiz 1967 Wurdack, Certamen Melastomataceis 265 described Venezuelan species with austro-Amazonian taxa, rather than with those from northern South America, is striking. “eaepora B65. Prodr. 37 °127T):2808. BC op Progr. 3: 092: 282 —- Raf., Sylva Telluriana 98. 1838. To Adelobotr 3, de Candolle or pe papa ascribed only A. Scandens (Aubl.) Dc. (Melastoma eonaiie ubl.). Under Dav. ) e listed fo e followi ta h species in moe D. panicula = ra D. guianensis ad D. peruviana DC.; of these, the first two were transferred in a fine Triana Meriania, while the latter two wer wh d by him und A. adscendens (Sw.) Triana. Gleason (1932) distinguished A. guianensis (Dc.) Gleason from A. adscendens by anther structure. In his discussion of Da Mem. Melast. 17. 1829), de Candolle indicated the generotype as D. guianensis by his rises J'ai primitivement &tabli ce genre sur une espéce recueille 4 e WO genera described in the Prodromus. In his treatment vya, he placed in Sect. Eudavya D. bra DC., D. claussenii Naud., and D. calyptrata Naud.; in Sect. Adelobotrys (Naud. DC. were placed D. ciliata Naud d D. scandens (Aubl.) Naud. Naudin apparently not seen the is type collection of D- guianensis DC., commenting at the end of the treatment ft under Species fortassis addendae): "10. ? guyanensis DC., 1.c.--Nonne affinis D. ciliatae?” Both Triana and nhlaux subsequently treated Davya as a section of Meriania and Adelob otr ys 2 nus. Naudin's failure to include D. guianensis in his Sect. ux! s part i alignmen Swartz's genus surely needs adjustm gender of the generic name ries s was not estab- lished ay de Candol Triana's usage was feminine with the trys should not be followed; however another respected botanical Soccmeagicn because the first part of the word, “Adelo-" Greek, has disagreed with this interpretation. ® ct fi 266 PHYTOLOGIA Vol. 1h, no. 5 In describing the genus Xeracina, Rafinesque placed there definitely three species of Aublet: Melastoms villosa 18 Placed by Naudin in his monotypic genera Desmoscelis and Nepsera. The thi cies had already (1828) been used b de Candolle as the generotype of Adelobotrys ave LS= plac i X. scandens (Aubl.) Raf., Sylva Telluriana 98. 1858, 4 is s herewith designated as the rong of Xeracina Raf. The gen description of Xera s sufficiently vague (and acon quae inaccurate in Senne gota that no taxonomic evaluation is needed as to Rafinesque's generotypic concept. Squetiwers IPAPOANA Wurdack, sp. no silifoliae Triana affinis, a foliis bene petiolatis septaplinervatic = ert. Arbor parva; ramuli hebeti-quadrangulati sicut folia subtus Hie esaanti on epeeeetdinics primum densiuscule furfuracei, pilis nigro-rufis crassis barbellatis 0.1-0.2(-0.3) mm longis gaint longi ad eae infra bigibbosi, gibbis ellipsoideis vel sphaericis 2- longis hebetibus; lamina (8-)12-18 x (5-)8-14 em, elliptico- ovata vel oblongo-elliptica, apice rotundato vel late hebeti- obtuso, basi ca. 3-5 m i obscure longi, 1-1. diam.; flores ob tori cicatrices evidenter S-meri. thium ( orum mn; torus rane fimbriata paulo ind arma calyx evidenter in lobis 2-3 evi ongis persistentibus fissus, tubo torum supra c4 1 mm longo. Capsula trilocularis, apice in tubum 2.5 mm longum lacera longato, dentibus 0.3-0.5 vam eS ares numerosa 1-1.2 X 0.2-0.3 mm, recta laev Collection: ire & & L. poles 25h Tee “(nototope, NY; isotypes US, VEN), collected in the Campo Grande, pty Sipapo, Terr. Amaz zonas, Venezuela, ed ero m, 20 ner Paratype: Maguire & Politi 28272, from Cafio Profundo, ne@r Cafio Negro, Cerro Sipapo, elev. 1460 m , 10 Jan. 1949. . Graffenrieda sessilifolia has pubescence, foliar texture 20 venulation, calyx lobes, and (in subsp. occidentalis Wurdack) te ? s ovary. Because of difference in the foliar venulation and pubescence, I do not believe that 1967 Wurdack, Certamen Melastomataceis 267 Sipapoana is closely related to G. rupestris Ducke; that ers es has a 5-celled ovary with similar aera te a Despite the distinctive leaves, the description of G. sipapoana has been see postponed because of the lack of flowering material (cf. - N. Y. Bot. Gard. 8: 136. 1953, sub G. versicolor); however the “hope for ‘such collections has become progressively IT a during the inimical late rainy season. The nodular at, eta at the petiole apices of the Sipapo tree are reminiscent of those found in the Ecuadorian Miconia gibba Markgraf. RAFFENRIEDA LATIFOLIA (Naud.) Triana subsp. MERIDENSIS Wurdack, ae nov. Folia 5- vel indiatincte Lae a Petala a Type Collection: A. L. Bernardi 1025 (nolotype ™), collected in clondoeesees at La cotta toe ,» Edo Méri Venezuela, elev. oe m, 2h Oct. 1953. "Arbol de as * de alto." Paratypes 3 elev. 2200 m; Bernardi 2258, Pueblos del Sur, elev. 1800-1900 m; pemantt seteemark 56018 oe San Isidro, above La Carbonera, elev. SEA 75 m; Steyerma. «x 56490, Canagua-El Molino, elev. 2530- 715 The typical subspecies, known from the West Indies, Trinidad, and the Venezuelan coastal cordillera ( Distrito Federal, Miranda), has 7-9-nerved (and generally Yasar} leaves and apically rounded petals. ‘The related G. emarginata (R. & P. ) from decellea ovaries, resembles G. latifolia. INGA GLANDULOSA (Gleason) Wurdack, comb. nov ‘ 1931 Macrocentrum glandulosum Gleason, Bull. Torrey Club 58: 422. SALPINGA PUSILLA (Gleason) hk 1931. Macrocentrum pusillum ee a — Club 58: 422. My doubts sencet oa the original generic disposition of these two Guayana species (Phytologia 9: 417. 1964) have been confirmed ce. a study of Venezuelan Bertolonieae. Both spec®s k, and S. &ck; the specie have OR Re et Stamens with eieiate "Nemenatne spurs MACROCENTRUM STEYERMARKII Wurdack, sp- oe M. rubescenti Gleason et M. minore gi pe affinis, cellis 1 ioribus differt apentrre 7-15 cm alta; ramili sicut petioli modice setulosi, pilis gracilibus caduce glanduliferis 0.7-1 m longis. pide Plerumque 3-8 mm longi; laminae membranaceae in quoque rs © Sequales vel paulo (1:1.5) inaequales raro valde dispariles, ith th generotype), S. nore (Gleason) Wurdac Wurd Macrocentrum sed 268 PHYTOLOGIA Vol. ly, no. 5 in folia matura ut videtur uninervatae sed in folia juvenilia Giseemerous trinervatae, nervis secundariis tertiariisque laxe ticulatis satis inconspicuis, plerumque 1.2-2.4 X 0.7-1.3 cm, sup os versus glabrae margines ver sparse glanduloso- setulosae, subtus sparsissime glanduloso-setulosae. ore = meri in ramulis terminales demum laterales plerumque solitarii; glanduloso-setulosum; ecalycis tubus 0.3 0.7-0.8 X 1.7 mm secus i meh ae sxperdori tis hebetibus paulo (0.1 mm) eminentibus. Petala 5-5-5 X 2.4 mm ga apice rotundato et setula glandulifera unic . 0.2 oO. Filamenta ca. 2 longa; antherarum thecae ca. €, connectivo infra loculos ca. 0.3 mm prolongato, appendice dorsali descendente acuta 0.6 mm longa. Stylus ca- : sti e ber apsula 3-locularis. cepa Th Bassett Maguire, Julian A. Steyermark, Celia K. Maguire 64 Th ee VEN; isotypes NY as), socked on top of moist ane facing escarpment, Cerro Uroi, Rio Uroi inage of Rfo Soper 6°15'N, 61°50'W, Edo. Bolivar, ‘ Venezuela, 13 Se 2. "Locally frequent. Flowers white. r ocentrum rube escens has pedicels 2-3 mm long, branchlet : es ek and -gthpbe 9-11 mm long. Macr ocentrum minu cauea as ie Wurdack, Fe i Trian ees Gleason (ex char-);, gies Mich ve: afens: sed foliis supra laxe strigu- sae differt. Ut videtur arbor exclusa) 15-35 -25 cm apice aay oT tas em caudato-acuminato Si paulo (0.7-l c¢ Je rdata, membranacea et integra vel obscure undulato-ser sup: ice longo-strigulosa, pilis a 1.3 mm longis laevibus gracilibus laxiusculis een subtus in venis venulisque primum modice sq glabrata in pr s e resinoso-granulosa demum glabra, T7-n ervis secundariis plerumque 0.5-1 cm inter se distantibus nervulis Us paulo elevatis et reticulati areolis ca. 0.3 mm latis. Panicula 22-26 cm longa submultiflora, ad apices bibracteolatis, bracteolis 2.5-3 mm longis anguste ovatis usque ad anthesim 1967 Wurdack, Certamen Melastomataceis 269 persistentibus; flores 5-meri. Hypanthium (ad torum) 4 X 3 m; calycis tubus 1.7-2 mm altus, lobis 0.5-0.7 X 4 mm oblatis apice rotundato, dentibus exterioribus vix elevatis nec liberis nec eminentibus. Petala 16-18 X 14-16 mm, extus dense amorpho- rum _ ro nato. Stamina isomorphica; Raoye eh ues mn longa modice ili Juxta thecas obsito, dorsaliter prope basim per 3 elevato ad basim dorsaliter dente hebeti alain 0.2-0.3 mm longo ornato. Stigma capitellatum 1.6 mm diam.; stylus 17.5 X 1.1-0.8 mm basim versus per 3-41 mm densiuscule glanduloso-puberulus in Ovarii apicem 0.5 mm immersus; ovarium 3(?)-loculare 1/2 inferum glabrum. Type Collection: Kjell von ee = (holotype S), collected in primaeval forest at La Costa, El Tambo, Depto. Cauca, Colombia, elev. 1000 m, 24 June 1 sg, type: von Sneidern 787, from the same locality, 7 Sept. 1936. The three suggested relatives have the general flora features of M. ingens, including puberulent hypanth ia and petals, large petals, expanded stigmas, and d anther csinanelewh: ait however, ha: leaves glabrous above at maturity (more- or-less ate-lepidote when young) th acute bases. Miconia centronioides (ex char.) has glabrous fi s and style and (as br 1 the connective glands only nea = thecae bases. More distantly relate M. axinaeoides, with a Pune tat stigma, and oblong-obovate leaves which are glabrous above. acing of M. centronioides, M. a. » and M. axinaeoides in Sect. Adenodesma does not seem satisfactory. The f cies above mentioned, along with M. notabilis Triana and a M. boecilantha Uribe, form a related species-group, but the most satisfactory sectional disposition is not obvious at prese ent. MICONIA RAVENII Wurdack, sp. nov. M. globuliferae Naud. affinis, sed laminis foliorum subtu Pilis brevissime s stipitato-ste ellatis instructis, ppm ap eme Pinas a glanduloso-setulosis, floribus plerumque ie pi air OT ea sicut petioli inflorescentia bypanthtaque pilis barbellatis vel brevissime (stipite 02-063 Ova im (aa 0.5 cm) cordata, membranacea et distanter (2 h ig Serrulata, supra modice strigulosa, pilis simplicibus 0.+-1. ee longis, subtus sparsiuscule (5-6 per mm quad.) stipitato (stipite 0.1-0.15 mm longo, ramis 0.15 Secundariis ca. 5 mm inter se distantibus, nervul 270 PHYTOLOGIA Vol. 14, no. 5 flora sparsissime glanduloso-setulosa, pedunculo 4-6 cm longo, mm -5-O.7 mm altis oblatis, agi tie bs Veece pest setula terminali 0.3 mm longa inclusa ca. 0.6 mm eminentibus. Petala 5-5.4 X 4h 2 om oblongo-obovata, apice rotundato, minutissime granulosa. Filamenta 3 mm longa glabra; antherarum thecae 3.8 X 0.7 X 0 mm, poro 0.2 mm diam. paulo ventraliter inclinato. Stigm punctiforme; stylus 10 X 0.5-0.3 mm basim versus sparsissime eifachs fone ti amans ovarium sine re 9/10 inferum apice in ollum 0.9 mm altum circum stylum protracto collo modice glandu- loso-setilose, pilis 0.1 mm longis. ollection: D. E. Breedlove & P. H. Raven 12928 (holotype Me 2469212; isotype DS), collected on a arr slope near ridge crest in the paraje of Banabil, Municipio Tenejape, tiaree, Mexico, elev. 2800 m, 10 Oct. 1965. "Plant 3 feet ea conia Sanaa has the lower leaf surface ial with @ =. about 0.25 mm long and the arms about 0.06 mm eglandular tart ieacénce pubescence, and ee ati gutiy 5 merous 1 ler 2938, det. Boqueron in Chiapas no great distance from Huehuetenango For the study of M. globulifera, Purpus from Ce sani Boguer én, More dis are M. anisotricha YS ) or Triana; the latter species “enon py me recent collections Nelson 2 Pringle 8101, Pringle 13495, Rzedowski 15533, Conzatti 3474, and Balls L710, from Vera Cruz, Hidalgo, and Oaxaca) is quite ieguan ag: of Leandra melanodesm® (Naud.) Cogn. in penecel: aspec a generic problem. Of the examinable flowers in wate Cees, 71 were 4-merous and we 5-merous. JITOTOLANA Wurdack, sp : M. ravenii Wurdack affin ora i foliis nies supra Strigulosis, inflorescentia modice glanduloso-setulosa, Ploribus Lerumque Ramuli teretes sicut petioli inflorescentia acagaaitinviy % pilis stipitato-stellatis (stipite 0.2-0.6 mm o, ramis O-1- 0.2 mm longis) erectis densissime armati. aes (1-)2-4 cm 3 lamina 5-11 X 2.5-5 cm ovato-elliptica, apice per = 1967 Wurdack, Certamen Melastomataceis 271 gradatim a basi late acuta, membranacea et integra, ra rse strigulosa, pilis ca. 1 mm longis simpli- i i scule uad., subt rr Puberula, pilis stipitato-stellatis stipite ca. 0.1 mm longo et ramis 0.2 mm longis, breviter 5 i ervorum pare interiore 0.5-1 cm supra basim scbakterans te” inserto, nervis secundariis 4-5 mm inter se distantibus, sane laxe reticu- a ob ubtus ob part Inflorescentia pauciflora sicut hypanthia sparsiuscule vel modice glan oie 4 gage legge. pili - 0.5 mm longis laevibus, pedunculo ca. 2.5 cm longo, ramis primariis plerumque tribus 1.5-2 cm ri 0.6-0.7 mm poro -15 mm diam. ventraliter inclinato. Stigma truncatum non expansum; stylus 8 X 0.5 mm glaber; ovarium S-loculare 9/10 inferum glabrum, apice in collo 0.7 mm alto circum stylum pro- rac Type Collection: D. E. Breedlove 11951 (holotype US 2469210; isotype nie ollected on slope along road to Pueblo Nuevo Solistahuacén, 8 km south of Jitotol, Chiapas, Mexico, wae Ea m, 19 August 1965. "Flowers white; plant 5 feet Of the flowers examined in Breedlove 11951, 43 showed 5-mery and 3 were 4-merous. As is obvious in the discussion un M. ravenii, the lan species M. echinoidea Stan Steyerm. and M. tixixensis Standl. . inly not ere in Miconia Sect. nS sears SNEIDERNII seine sp. nov. multuplinerviae Cogn., M. penicillatae Gleas., et M. ta Wurdack affinis sat foliis non plinervatis supra retes sicut petioli foliorum su vel inflorescent iaque pilis erectis stipitato-stelletis (atipite ah athe mm longo, ramis 0.5 mm longis dense uti. ee » 272 PHYTOLOGIA Vol. 1h, no. 5 glomerulis ad ramulorum apices aggregati, seething: eas Ee5 ox O.4 mm basalibus, pedicellis 0.7 mm longis. Hypanthium (ad lon ; i yeis tubu re oribus 0.4-0.6 mm longis oblongis, dentibus exterioribus lobos 0.3-0.4 mm glaber in ovarii apicem immersus; ovarium 3-loculare 1/3 inferum apice setulis ca. 8 ace steasic 0.4-0.5 mm longis coronato. Type Collection: Kjell von Sneidern 5525 (holotype ey collected at Pueblo Rico, La Selva, Depto. Caldas, Colombi elev. 1500 m, 28 December 1945. "Flowers white. The suggested relatives have plinerved leaves which, on the ia t nature of the anther form for sectional delimitation (paytenogta 11: 390-391. nas are still pertinent. Apart from the anthers, the general aspect of M. sneidernii is like that of M. barbi- nervis (Benth.) Triana and its allies (Inetuting M. stipitata Gleason and M. cionotricha Uribe ex char eit et ges ivcion SIS Wurdack, sp. Amblyarrhena. M. ate oan. et M. amabili Cogn. eeriien: bad petiolis linea paulo elevata in “nodis connectis differt. Fru 0.8-4 m. altus. Ramuli obtuse — “nates demum a. sicut folia turlatesdenbiaque squamulosi mox glaberrimi. Petioli 0.7-1.2c floes oes Tamin ies X 3-5 cm gages Lapa deope apice = ee seen Be inato, basi acuta (in petiolem ca. 0.5 cm decurrens), pare J obviis nervulis reticulatis areolis plerumque 0.7-0.9 mm diam. Inflorescentia submultiflora 7-10 x 4-6 cm, ramulis teretibus 1 paulo ‘sphere flores sleet: pedicellis 1-2 X 0.7 m, peuto intra 2.5 K 0 -6 mm mox caducis juxta hypanthii basim vel s orbicularia dense granulosa. Stamina isomorphica; eilamen 4 - entraliter apicem versus sparsissime puberula, ee non glandulosis 0.2 mm longis; antherarum thecae 2.8-2.9 X 1-3 ° ula ad to igma 1.5 mm diam. capitellatum; stylus 5 X mm pilis gracilibus non eiaamiiodds 0.2 mm longis densiuscule 1967 Wurdack, Certamen Melastomataceis 273 indutus in ovarii apicem 0.5 mm immersus; ovarium 3-loculare 1/2 infe ice conico sparse puberulo. Collection: P. C. Hutchison, J. K. Wright, & R poy La 5935 (holotype US 2469205; isotypes UC, ed biog: at Carpish, above Acomayo on the road to Tingo Maria, Depto. co 1964. "Leaves red-margined, Shiny green, lighter beneath. Flowers creamy white; ive ae cle Breen; anthers and filaments golden, the anthers later orange bro ge =] B c “ © d “ 0) a oO Ny ne) © 8 B “ | oo win Cy = ratypes (both Ned NR get . Ferreyra 8066, 1 Oct. 1950; =. Asplund 13090, 15 Sept. oth M. bangii and M. am a bee es similar puberulou enh but smaller petals; aos lack the elevated rnteetiolee line at the branchlet nodes. Miconia lucida Naud. (ex descr. hypanthium and truncate calyx limb as well non-e d istant relati M. saltuensis, all with much smaller flowers and glabrous styles, include M. tere M. cookii Gleason, M. e. The description of M. adrienii Macbride might be somewhat sugges of M. saltuen- an icularly, pleiostemony - a feature not noted in the original description. Recent collections of M. adrienii (which have been compared with the holotype) are: Ferreyra 990 (US), La Divisoria, San Martin; 2 lund 12567 (S), from Divisoria, Hudnuco; and Asplund 13294 from between Pumahuasi and Divisoria. ibe Srey sed ONII Wurdack, sp. t. Aublyerrhens- M. Reger Naud. affinis, we foliis eben in nervorum primariorum axillis setulosis, stigmatibus capitellatis differt uadrangulati demum teretes sicut folia (poculis lob dentibus exterioribus omnino adn % 2.4 ores aequantibus. Petal eigen quan iy marginato- Stamina glabra ° marg e connectivo crasso basaliter 0.3 mm prolongato- Stigma i sd tum 0.8 mm diam.; stylus glaber 3-3 X 0-3-0- 6 mm; ovari 27h PHYTOLOGIA Vol. 1h, no. 5 inferum 3-loculare, apice eee: lection: P. C. Hutchison & . Wright 6638 (holotype US 2469206; —— UC, can. a on the summit km above Huancabamba, on the road to Canchaque, Prov. Huanca- bamba, Depto. Piura, Peru, elev. 3200 m, 17 Sept. 1964. "Shrub -l1 m. oo alypifolia, known from southern Ecuador and northern a: (Cag rea), = Se ees paaned vein axils on the — 11 @ both of Sect. Creman rted from northern Peru have the general aspect (gla isbrout, “anal -Leaved) of M. hutchisonii; M. vace Huoldes tear Naud. has nodding T-merous flowers and i a. » vaccinioides “rea nd met been ae ee since Weberbaier’ s field days not seen any Peruvian colin? tion of Me buxifolia -(idberbauer 263, from Huambos, Cajama: was so cited a the Peru and am somewhat ceptinet of this record. Bly coun nie lowers in Hutchison & Wright 6638, 42 were 5-merous and 30 were 4-merous. MICONIA MEDIA (D. Don) Naud. subsp. CAJAMARCENSIS Wurdack, subsp. nov. p- media foliis phen subtus ubique laxe setulosis, pilis laxis luteis 1-1.6 m longis, in superficie tarde glabratis differt Type C ollection: P. Cc. ee & J. K. Wright 5028 (holotype US 246920); Stee UC, USM), co. AE an on a steep brushy hillside 1 km northeast of Sin Pablo, Depto. Cajamarca, to the p v 1s and primary vein bases ace (as does subsp. borealis Mis subsp. censis, th ir pr leaf veins are more obvious the typical subspecies and the yi lets an inflorescences are more densely (but caducously) stellulate- eous are no floral differences appa af Peruvian populations; an Ec rian subspecies (the description uad Ow in press for the Memoirs of The New York Botanical Garden has glandular-puberulous styles. ADDITIONAL NOTES ON THE GENUS VERBENA. III Harold N. Moldenke VERBENA [Dorst.] L Additional synonymy: Verbena [Tourn.] L. ex Migula, Exkursions- fl. Deutsch. 2: 13). 1906. Verbenia L. ex Moldenke, Alph. List cate serene Suppl. 1: 28, in syn. 197; Runner, Rep. G. W. Groff oll, soy tonal. & amended bibliography: Rumph., Herb. Amboin. 7: 60 d : se ivssler , Gemain. Handb. Gew. 2: 816 & Reg. 1815; W. D. J. & S ° : - 1845; W ch. & Schw, Fle, ed. 25 681—682 & 1206. ret asc. (18 . 4, xliii, 00, & 583. ; Ry sce & Mitteldeutsch., ed. 2 , 252-253, & h62. 18565 Guillard, . Soc. Bot. France : 122. 1857; W. D. J. Koch, Taschen Deutsch, Fl., ed. ’ xliii, & gy Hane nia 23 86, 106-107, 110-113, 11%, 118, 123, 12h, 26-128, "02133, 136, 139, 140, 142, kh, 16, 1h?, “thd, ied-ish, 157, & 158 (1862) and ak "178, "180, "181, "183, 185, & 201—204. 4 Bocq., Rev. ; o, 106-10 113, 115, 118, 123, 12h, 126— roe Le I35 136, 139, 140, 1h2, 1b, UT, als, 152—15h, 7, 1 a 10, 181, 183, 18 & 201 1863; W. D. Je ‘ Taschenb. 3, 5, & 583. eke eit Langethal, ° xliii Bot h., ed. 2, 48 & 8 fake ate Cat. Mus. e Lugd.—Bat. 70. 1870; E. Hall.in W. D. J oc Deutsch. Fl., ed. 7, 03 & 802 (1678) and ed. 8, 03 & $02, 1881; ‘ Twat 188; K. Pflanzenfr. erate rs 6 Berar Essent. Bot. ooh "bale tere "J. Matsun., 4 relyo 13: (iis}—alh aa 18595 395 Téhimura, Bot. Mag. Tokyo 13: 116. « Deutsch. 2: Kawakami, Lis ei Acad, Sci. sl it 1: pep 3 Dunn & Tutcher, Kew Bull. Misc. Inf, Addit. Ser. 10: 201 & 202. 2. do125 E. D. Merr., sade “a Rumph. Herb. haba, 5u7. ell Robertson, Psyche 29s 35 “ga ‘ 1922; Gamble, Fl. Madras 1085 & 106 192h; $. Sasaki, 276 PHYTOLOGIA Vol. 1h, no. 5 ill., Gard. x. 1935; Fletcher, Kew Bull. Misc . Inf. 1 938: 408 & 10-11. 1938; Hottes, Book Shrubs, ed. h, “yo gi 192; Plakelock, Kew Bull. Misc. Inf. 1949: 539. 1949; Brenan, Kew Bull. Misc . Inf. 1950: ‘ r, os . he Sig hile, 573c, 956a, 1137f, & 3018f. 1952; Ke He er, Arkiv Bot., a 2, 2: 08. 1952; H. C. Greene, Trans. Wisc. Acad. Sci. 2: 70. 1953; Bor & Raiz zada, Some Beaut. Ind. Climbers [136]. 1955 Berhaut, Fl. Sénegal 65. 195k; rg: jute & Wylie, Chromosome Atl., pr. 1, 322—323 & rent 1955; E. W. Lath- rop, Univ. Kans. Sci. Bull. 39: 12h, 125, 137, 172, & ins. 19583 Mostert, Bot. Surv. S. Afr. Mem. 31: hl & 191. 1958; T. A. Rao, - Bot. Surv. Ind. 1: 11). 1959; K. H. Reckinger, ag it ew Bull. Moss, Fl. Alberta 397. 1959; Nath, Bot. Surv. South. rop T Wi tice el ers 5. 456k} . Kans. Sel ee sc. Sit Pent 23! he 1 Ungar Bull. 46: 18, 19, & 28. 1965; Percival, Flor. Biol. "Ts—132. 19655 Backer & Bakh., Fl. Java 2: 695 & 596. 19655 J. E. Weaver, Native Hawke D 43 51). 1965; Liogier Rhodora 6 Po : 19683 Solbrig Castanea 30: 173-—17h. 2 1966; Rickett, Wild yore (2): 361-36, Bh oie 112 . 7 C. By Clarkson Castane 3h: e = nan, Biol. Abstr. 7: 615. 1966; T. Swain = an BA & re CG e bad & 170. 1966; Harborne in 7. C : 285. 1966; Moldenke, 2h3—276, 307, 366, Ol, 509, & 512 (1966) and Us: 175. 1966; G. Taylor, Ind, Suppl. 13 e 1966; ewe f Anone, Costa Ric. Phan. 10. 1966; Moldenke, RSsumé Suppl. 1}: 13, 5,9 & 10. 19663 Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 196855 67- 966 Results arakor 132. 1966; J. Gémez, Biota 6: 103, 107, 109, 11)—116, & 119. "i966 Rs C. Jacks., Reg. Veget. bas 59. 1966; Stearn, Botan. Latin hi? ah S461 & 8.162. 19663 Herbst B08 Sonn Se eee erymen 23, 1966; G B. Foster, Herbs for Every G ° ; gee s-Craig, Drawings Brit. Pl. 23: we 38. 1966. 1967 Moldenke, Notes on Verbena 277 Robertson (1922) and Percival (1965) report that ae verbenae is an oligolectis bee of the family Panurgidae which col- lects solely from three species of Verbena, viz., V. hastata L., V. stricta Vent., and V. urticifolia L. Hostert (1958) reports that an unidentified prostrate herbaceous Verbena species was re- corded from Bloemfontein, South Africa, represented by G. Potts 2079. er (1965) sided but does not a a "woods found in Nebraska only in woodlands, n the prairies. The Heddle 2816, distributed as a species sig (soe is actu- ally Lantana 1 macropoda Torr., while M. C. Johnston 266h, Waterfall & Wallis 13476, and Weintraub & Roller 173 are Priva grandiflora (ort.) Moldenke, and Bowman s.ne - (Sept. 29, Maro is Salvia occi- dentalis Sw. in’ the Lamiaceae VERBENA oe Additional bibliography: Boeq., Adansonia 3: [Rev. Verbenac.] 202. 1863, Moldenke, Phytologia 335 181. 1966. A AMBROSIFOLIA Rydb. Additional bibliography: Darlington & Wylie, Chromosome Atl., heed 1, regs (1955) and pr. 2, 323. "19613 i Phytologia 13: 19 Additional citations: TEXAS: Brewster Co.: D. S. Correll 33820 (La). VERBENA BERTERII (Meisn.) Schau. Additional bibliography: Bocq., Adansonia 2: 126 (1862) and 3: 202. 1863; Boe ocq., Rev. Verbenac. "j26 & 202. 1863; Moldenke, Phy- tologia 13: 183. 1966. VERBENA BIPINNATIFIDA Nutt Emended syno: : Verbena b ipinnatifida Schau. apud Bocq-, Adan- Sonia 3: 202, ert Additional bibliography: Bocq., Adansonia 3: [Rev. Verbenac.] 202. 1863; eaarheony S pig taidess U8 ahy--2h6» 1966; Rickett, Wild Fls. U.S. 1 (2): 36h. 1966. Jones found this species growing in post oak savannas. Additional citations: TEXAS: Bexar ee Martz & Martz s.n. (June 3, 1959] (Ws). Brown Co.: a2 & McCart 933k (id); Ewing 27 (Ld). Burnet Co.: J. Jones 1 ( Ld); F. Sylvester 2 er 2 (Ld). Cole- man Co.: J. Folimer 13 (Ld); H. Spoon 13 3 (Id). Coryell Co.: Se Jackson 11 (Id). Eastland Co.: Wheeless 18 (14). Turner 17 17 (Id). Garza Co.: B. Jensen 8 ( Ld). Kaufman Duke lo (¢ (Ld). San Saba Co.: “Calhoun 13 (Ld); E. Howell 1 (14). Williamson Co.: J. Sargent 6 oF eS xVERBENA BLANCHARDI Moldenke Additional bibliography: E. W. Lathrop, Univ. Kans. Sci. Bull. 278 PHYTOLOGIA Vol. 14, no. 5 39: 172. 1958; Moldenke, Phytologia 13: 18). 1966. Le Additional bibliography: Guill., - Pl. Déc. Voy. Iles So- ciété 209. 1837; Bocq., Adansonia 3s eer Verbenac.] 202. 1863; Dunn & Tutcher, Kew Bull. Misc. Inf. Addit. Ser. 10: 201. ee Chromosome Atl., pr. 2, 323. 1961; B & Fl. Java 2: 96°, 1965; a lieeiar, Rhodora 67: 349+ 19653 } Moldenke, Phytologia 13: 245.1 Recent collectors Resists this plant as having the calyx green below to reddish above, the corolla whitish Bs reddish in the central region, and lavender above, with white hairs, the anth agricultural fields, Cinchona plantations, grassfields, and along roadsides. Mostert: (1 1958) ess that ae be Africa it is found on streambanks, » flooded an ee areas sub- ject to wetness and Feoord “severe grazing, and trampling, while Sidey refers to it there as a "shrub 5--6 feet t tall", with purple fl rs. Sampaio (1937) iecdas the vernacular name "cambaré de Additional citations: JAMAICA: Crosby, Hespenheide, & Anderson 231 aged Yuncker 1867) (Mi). sou AFRICA: Cape of Good Hope: 2-(8). ca BRACTEATA Lag. & Rodr. Additional bibliography: Bocq., Adansonia 3: [Rev. Ver benac «] 202. 1863; E. W. Lathrop, univ. Kans. Soi. Bull. 39: 173. 1958; E. H. Moss, Fl. 7. 1959; Ungar, Univ. Kans. os Bull. 46: 28. 1965; Rickett, wad Fls. U. S. 1 (2): 362, pl. 1966} ais 15 . lg » Castanea 31: 100. 1966; Moldenke, Ra pik 13? dditional illustrations: Rickett, Wild Fls. U.S. 1 (2): Pl 112 fin bss th 1966. Holmgren & Reveal 1023 is said to have had "blue-violet™ co- arpet vervain' more or gs erent and blue to purple corollas, growing on "stream fla roadsid waste places in the t ~~ region® of Alberta, Eanrep (1958) deseribes the habité of the species as "fields, waste places, and pastures” e Ur rad (1965). tells te that it is "prominent" on plaegi on. Additional rata NEVADA: White Pine Co.: 1023 ~ a Buffalo Co.: Guthe s s.n. (Kearney, Bolngren & Boss] 0H). : Floyd Co.: Purvis 21 (id). 1967 Moldenke, Notes on Verbena 279 VERBENA BRASILIENSIS Vell, Additional synonymy: Verbena brasiiensts ae hes in herb. Additional bibliography: Sampaio, - Nac. Rio Jan. ed 20. 1937; Moldenke, _ Paptolagia 133 "he "39665 Rickett, Wild Fls U.S. 1 (2): 36h. 1966. Recent collectors describe this mate as 2—3 feet tall, with a minty odor, growing at low as 200 meters altitude in Peru. e of a large le , in poor soil — h-brown fine with a little clay — in t+ was originally cerrado eta- tion; Ugent found it on sunny lich overed piles anong In- weeds along 4 potato fields, and among weeds in fertile deep-soil cabbage (Brassica oleracea) fields along with abundant Solanum calcense and also Rumex, Er Eragrostis, Oxalis, Polygonum, Galinsoga, a, Tagetes, etc. The flowers on G. Eiten m 1595 are described as having been le, Additional citations: PERU: Cuzco: Iltis, Iltis, Ugent, & gent 891 (Ws); D. Ugent $35) (Rf). Junin: 1tis, tis, Ugent, & Ugent 347 (Ac). Lima: Tltis, Iltis, Ugent, t, & Ugent 390 (af). BRAZIL: S&o on G. Eiten 1595 (N). VERBENA CAMERONENSIS L. I. Davis ‘ To wo ge bibliography: mained, Phytologia 13: 2h5 & 257. VERBENA CANADENSIS (L.) Britton Additional witbece te Bocq., Adansonia 2: 126 (1862) and 3: 202. 1863; Bocq., Rev. Verbenac. 126 & 202. 1863; Darlington & Mog e, Chromosome Atl., pr. 1, 323, 1955; E. W. Lathrop, Univ. - Sci. n Atl., pr. 2, hele 1961 wor denke Phytologia 13: 2h5—2h6. 19665 Rickett, W se 1 (2): 362 & 3h, pl. 112. 1966. Additional’ iastednens micheks; ba Fls. U. S.1 (2): ple a5 Sipe fei his plant is frequent and prom 19 tells us that this s inent es in the spring, in rocky prairie gg and wanda: open woods, and along roadsides dditional citations: TEXAS: Henderson Co.: Correll & Correll 23960 (Ld). County undetermined: T. Drumond 263 (S)- oBoK. benac -] ography: Bocq., Adansonia 3: [Rev. Ver 202. 1863; Moldenke, Phytologia 13: 26. 1966. Recent collectors ve found this growing on dry grazed and cacti, in te grasslands now becoming desert. flowers are — "blue" on Roe, Roe, & Mori h. 280 PHYTOLOGIA Vol. 14, no. 5 Additional citations: MEXICO: Coahuila: Roe, Roe, & Mori 4 (Ws). Puebla: Ugent & Flores C. 2539 (Ws). INA CANESCENS var. ROEMERIANA (Scheele) Perry Additional babldography Molderke parscioa> ia 13 Bovis 1966. Runyon describes this branches above the base, Peace on open clay ry on with rs. dditional citations: TEXAS: Brown Co.: Benson & McCart 9335 (Ld). Cameron Co,: R. Runyon 6067 (Ld). Coleman Co.: Be E. Ho. Hol- land 21 (Ld). Travis Co.: f0.: Webster & Webster 115 (Mi). CAROLINA L. Additional peas a a Bocq., eae 38 Eee veheee 203. 1863; A. Stewart, . Calif . Sei., 8 1911; Moldenke, Pigtologia 13: 2hé, eel, "& 258. 1966; etre Résumé Suppl. ib: ah Breedlove found this species growing on grassy slopes, grassy » and in large moist pastures. He describes the flowers as "blue! on his nos. 10457 & 1152 and as "pale-blue" on no. 10519. Feddema 1659 ; is said to have had the "corolla white with ring in thr . Bowman 81, distributed as V. carolina, is actually ¥. betula Moldenke. Additional citations: MEXICO: Chiapas: Breedlove 1057 (9s 10519 (Rf). Guerrero: Feddema 1659 (Mi). México: Roe, 0e, Roe, & Mori Mori 1530 (Rf). Michoac&n: Ugent & & Flores C. 2217 (iis), 22h5 (Ws). GUATEMALA: Quezaltenango: Breedlove 11,52 (Rf). neem 6, 287, & mal bibli raphy: Moldenke ee 13: ’ , 273. “i966, Moldenke, résumé Suppl. 1: 1 1966. F describes this plant as an " enone weed of ¢ . fields" in tigen His no. 2335 is saidto have had nee der" flowe owers on Mick & Roe 15 are said to have been ovtelats tin, gelloé;‘ but “thee ox Roe, Roe, & Mori 10, 23, & 40 were "lavender", Recent collectors have found this plant growing along grazed atoms and drainage ditches with Yucca, Bouvardia, a eg cacti in mes grasslands now becoming desert, on heavily graz rosa sides with species of Yucca, Acacia, and cacti in highly distur mesquite grassland vegetation with cornfields, and on dry grazed 1967 Moldenke, Notes on Verbena 281 Slopes with Agave, Opuntia, Acacia, and Solanum polyadeni Mick & Roe call the seaken "common" in deserts with Sriruad desert pavement", along with Agave, Alo&, and cacti. Additional citations: MEXICO: Coahuila: Mick & Roe 15 (Rf); Roe, Roe, & Mori 10 (Ws), 23 (Ac). Jalisco: Feddema ma 2335 (Mi). Wueve Le Teeny Roe, Roe, & Mori O (Ws). Puebla: Ugent & Flores C. 2522 (Ws). VERBENA CILIATA var. LONGIDENTATA Perry Additional bibliography: Melsenke. » Pigtologia 13: 26 & 2h7. 1966; Moldenke, Résumé Suppl. 1): 1966 CILIATA var. PUBERA (Greene) Perry Additional bibliography: Moldenke, etclenta 13: 2h6, 2h7, & 257. 1966; Moldenke, Résumé Suppl. ih: 1. 1966. VERBENA CLAVATA Rufz & Pav Additional biblio graphy: Bocq., Adansonia 2: 126. 1862; Boca., Rev. Verbenac. 126. 1863; Woldenke, Phytologia 13: 190. 1966. VERBENA COCHABAMBENSIS Moldenke Additional Se reteerarta? Moldenke, peep tna 10: 210s i: Recent collectors have found this * plan Se bluffs above roadsides, flowering and fruiting in , Additional citations: BOLIVIA: Cochabamba: Ugent & + “Ugent 5223 Ac, Additi gg ree Adansonia 3: [Rev. Verbenac.] onal bibli hy: gh hoas le) a ad bd 202. 1863; Darlington & tylie, Chromosome Atl., pr. 1, 323 (1955) and pr. 2, 323. 1961; Moldenke, Phytologia 13: 27. 1966. VERBENA CRITHMIFOLIA Gill. & Hook. Additional bibliography: Darlington & Wylie, Chromosome Atl ek. toe i, 3 (1985) and pr. 2, 323. 1961; Moldenke, Phytologia 1: VERBENA CUNEIFOLIA Ruiz & ap ; Additional bibliography: dansonia 3: [Rev. Verbenac. 2 203. am 35 Noliceks, Pryectogia 33: ATL. 2 1966; J. Gémez, Biota 6: 107, > ates bénes "(1 966) ec for this species the vernacular name paraiso", a name also applied to V. fasciculata Benth. VERBENA DELTICOLA Small : Ey sper bibliography: Moldenke, Phytologia 13: 27 & 27h. , Adansonia 3: [Rev. V GA DISSECTA woe 203. 18 18635 Da eat ese & Vylie, peigge hacmeoea Atl., pr. i, 322 322, (1955) 282 PHYTOLOGIA Vol. 14, no. 5 and pr. 2, 322. 1961; yaldonke, Phytologia 13: 247—2h8, 251, 255, 256, 262, 263, 268, 270, & - 1966. Bocquillon nm (1 863) refers o% “this taxon as "y. dissecta Walp.", a binomial referred by me to V. sulphurea D. Dae; fasaks Recent collectors have found V. dissecta growing on weedy, pre- viously cultivated, but now fallow slopes along with Solanum boliv~ iense, S. radicans, Chenopodium, Tagetes, Setaria, and d Plantago. Additional i tathiaat BOLIVIA: Chuquisaca: Ugent & Cardenas C4rdenas 4988 (Ac, Rf). VERBENA DISSECTA £. ALBA Moldenk Additional bibliography: Moldenke, Phytologia 13: 248 & 255. 1966. ENA ELEGANS H.B.K. Additional bibliography: Moldenke, Phytologia 13: 28. 1966; Moldenke, Résumé Suppl. 1): 2. ai vo Rec collectors have encountered this plant on steep slopes with Quercus and on east~facing ciausbane cliffs with Acacia and Agave in Quercus scrub vegetation on rolling hills in the mesquite- grassland zone. The flowers are described as "purple" on Breed- love 12421 and as "maroon-red" on Be oe, 2 aes & Mori 123. Additional citations: MEXICO 3: Breedlove 1221 (Ac). San Luis Potos{: Roe, Roe, & Mori 3 tan)? ENGELMANNII Moldenke Additional bibliography: Moldenke, Phytologia 13: 2h8. 19665 gene! Résumé Suppl. 1h: 1. 1966. One of the A. R. Moldenke 821 specimens previously cited 38 de~ posited in my personal herba: ae bigicrall is now in that of the Texas Re- search Foundation at — Additional citations: NEW YORK: ‘County undetermined: J. Torrey 8.n. (Mi). ILLINOIS: ates Co.: A. R. Moldenke 1261 (Rf). MIS- eee Ralls ge A. R. Moldenke 1278 (Ac). Randolph Co.: A. Re Moldenke 1281 (Rf). MOUNTED ILLUSTRATIONS: A. R. Moldenke Kod. 9b ORES 22 (Z), 23 (Z). VERBENA FASCICULATA Benth. Emended synonymy: Verbena mathewsii Briq. ex Phytolo- 4 9: eee in syn. 1983; a. Gémez, Biota 6: 103, os 5 "a9. — onal bibliography: Moldenke, Phytolo : 57. 19655 = Bots 6: witha 107, & 119. 1966. Gémez (1966 the name "parafso" for this Species, a name ata applied to V. Sanaa fot Rufz & Pav. FECUNDA Moldenke Additional bibliography: u , Phytologia 13: 195. 19665 % Taylor, Ind. Kew. Suppl. 13: negeres 1966. 1967 Moldenke, Notes on Verbena 283 Moldenke Additional bibliography: Moldenke, Phytologia 13: 195. 1966; G. Taylor, Ind. Kew. Suppl. 13: 143. 1966. VERBENA FLAVA Gill. & Hook. Additional bibliography: Dar’ nm & Wylie, Chromosome Atl., tet i, ae (1955) aad pr. 2, 22. "E5614 Moldenke, Phytologia ll: GLABRATA H.B.K. Additional bibliography: Mol dariea, Fong ooapiage 11: 458. 1965. The A. Stewart 3317, previously cited by me as V. glabrata, is actually the type collection of V. glabrata var. temispicata Mol- denke. LABRATA var. TENUISPICATA Moldenke, var. nov. Haec varietas a forma typica speciei spicis temissimis recedit. its inflorescences much branched, the peduncles and individ- = gpg being extremely slender, the fruits sparse and not con- The type of this ew was collected by Alban Stewart (no. 3317) at Villamil, Albemarle Island, Galapagos Islands, on m Au- gust 23, 1905 ani _ # Aarons a in the Britton Herbarium at New Hew York Px n . The collector notes + the tis om 600° ter. at 2780 feet. This taxon was included by me in typical V- rata H.B.K. in previous installments of these notes. Citations: GALAPAGOS ISLANDS: Albemarle: A. Stewart 3317 (Ge— 31375--isotype, N--type). VERBENA GLUTINOSA Kuntze ae Additional bibliography: Moldenke, Phytologia 13: 196. 1 "5 aed citations: ARGENTINA: Mendoza: Ruiz Leal 7665/14 GOODMANI Moldenke Additional bibliography: Moldenke, Phytologia 13: 197. 19663 G. Taylor, Ind. Kew. Suppl. 13: 143. 1966. VERBENA GRA GRACILESC. e erter aa eat See Be , Chromosome Atl., e-igeaow ography: Darlington & Wyli ae § i, 23 *(1985) and pr. 2, 323. 1961; ssatihesiin Phytologia 13: GRISEA Robinson & Greem. gee mal bibliography: A. Stewart, Proc pat. ioe Sci., ale 23 23- 19125 Weldenice, Phytologia 92 tag se bangs tie rare species was collected by G. on Duncan I Islands, in August, SS ited in the Gray H sang, Galapaste I Harvard University at Cambridge, Mas- 28h PHYTOLOGIA Vol. 14h, no. 5 sachusetts. Additional citations: GALAPAGOS ISLANDS: Duncan: G. Bauer 180 (Du—photo of type). a . nal bibliography: Bocq., Adansonia 3: [Rev. Verbenac.] 203. “18635 Moldenke, Piptoncgia 13: 197. 1966. ENA HALET ee eb iocechy: Moldenke fon se 13: 2h7——-2h9 & air 1966; Moldenke, Résumé Suppl. ui: 1 9 cates citations: al a Bell a Me Latham 7 (Ld). Brow so Ewing 28 (Ld); J. Wheeler 1), (1d). Eastland Co.: Wheeless 17 (1d). Frio Co.: Drews 6 ms 6 (Ld). Kleberg Co.: J. R. Crutchfield 1088 (Ld). Medina Co.: C Cope 26 26 (Ld). San Saba Co.: L. J. Jones 18 ( (Ld). HASSLERANA Brig : Additional habiineceas: Moldenke, Phytologia 13: 199. 1966; Stearn, Botan. Latin 295. 1966. HASTATA L. Additional bibliography: Bocq., Adansonia 2: 112 (1862) and 3: aby 1863; Bocq., Rev. Verbenac. 112 & & 203. 1863; W. Stone, Pe N. J. State Mus. 1910: 660. 1911; C. Robertson, Psyche 29% 159-15. By fc E. Ms Lathrop, Univ. Kans. Sci. Bull. 39: 173- 19585 : Acad. Sci. 53: 195. 196k; Ungar, “9 ae oa cS - 16: 18, we & 28. 19655 Percival, Fl Biol. 131. 1965; R. B. Clarkson, - 1966; Moldenke, ‘s: Phytologia 13: 249 & 260. 1966; Rickett, Wild Fls. U.S. 1 (2): es we Additional illustrations: Rickett, Wild Fls. U. S. 1 (2): pl. 112 Se erat m (isan) and Percival (2965) report that the py tic eat Verbenajus verbenae amily Se) prairie fe yale in Bonxant! While Ungar (1965) describes it, from the same state, nt sum Scirpus HAMPSHIRE: Carroll Co.: A. R. Molden- ke 127 es ee ). Fair field Co.: A. R. Hcldenke 1238 (Ac). : A. Re Moldenke 1252 (Rf). ILLINOIS: Clay Co:: A. Re Yol dene 125k (ac). Fayette 1967 Moldenke, Notes on Verbena 285 Co.: A, R. Moldenke 1263 (Rf). Macoupin Co.: A. R. Moldenke 1268 (Rf). Marion Co.: A. R. Moldenke 1258 (Ac). Montgomery Co.: A. R. Moldenke 1265 (Ac). MINNESOTA: Ottertail Co.: P. Johnson 545 (i). KANSAS: Dickinson Co.: A. R. Moldenke 1297 (Rf). Osage Cow: A. R. Moldenke 1303 (Ac). - MISSOURI: Monroe Co.: A. R. oy denke 1279 (RF). Ralls Co.: A. Re Moldenie 1277 (Rf). Buffalo Co.: Guthe s.n. (Kearney, . Aug. 1 ug. 1893] ] (Mi). MOUNTED ILLUS- TRATIONS: A. R. Moldenke Kod. 7b (Z), 8b (Z), 9a (Z), 11b (2), lye (Z), 16 (Z), 17 (Z), 20b 0b (Z), 21b (Z). HASTATA f. ROSEA Cheney Additional bibliography: enaniey eee se - 1966. Additional citations: MISSOURI: : A. R. Ko ldcrke 1276 (Rf). MOUNTED ILLUSTRATIONS: A. Re apr Kod. “73 (Z), 19 (Z)> 20a (Z), 2la (2). VERBENA HIRTA Spreng. oe bibliography: Moldenke, Phytologia 13: 249—-250. Reitz & Dsin 17L8 represents the very conspicuously white- hirsute _ m or stage of this species; the flowers on their no. 1700 are descathas as "roxa™, ~~ Kdditional citations: BRAZIL: Paran4: Reitz & Klein 1700 (W-— 2465578) , 17484 (Ac). VERBENA HIRTA var. GRACILIS Dusén t Additional bibli ee powepon. Phytologia 13: aes ie . T owers on Reitz & Klein 17883 are described as it was a ceetectes at 950 m Sees = Ttitate, S iseuiees in De Mebtabed ditional citations: BRAZIL: P. Paran&: Reitz & Klein 17616 (Ac), 17883 (Ac). ENA HISPIDA Ruiz & Pav. : Additional bibliography: Bocq., Adansonia 2: 112 (1862) a 3: 203. 1863; Bocq., Rev. Verbenac. 112 & 203. 1863; Moldenke, P 0. 1966. ien sia Inca ruina alo ong with Solanum Se canasense, ag Bidens, Mutisia, Baccharis, rage “Talpoclaria aria, Casts sey Veronica, Euphorbia, Rumex, Capsella, Brassica, pew be a liaria, Hypoxis ——————— tiun, Ste F » Lupinus, and species of non-tu Solamm, as well as in rocky soil along open Spee with Sone Rum soukupii, Senecio, Zagetes, Capsella, Brassica, Eaas a Ce rastiun, and non-tuberous species of Solanum, frui bru- ary. Steinbach ee the flowers "flor violeta vivo base ox” terior r pirpurea ro iso ° Additional nfed> title PERU: Cuzco: vee F Seaba & vest 2 & 308 Se Puno: Ugent & Ugent 11543 (Rf). BOLIVIA: © Es 286 PHYTOLOGIA Vol. 14, no. 5 bach 8) (S). HOOKERIANA (Covas & Schnack) Moldenke Additional bibliography: Barlincten ‘ Wylie, Chromosome Atl ee tta 35 ae a and pr. 2, 323. 1961; Moldenke, Phytologia 13: HYBRIDA Voss Additional synonymy: ieee hybrida Grtml. & Rpl. ex Backer & Bakh.,, Fl. Java 2: 596. 1965. oxigen Uethisography: S. Sasaki, List Pl. Formos. ze: 1928; Bodger tros 61, 95, & 105, pl. Shg, bile, 573c, 956% 1137f, & tr 1952; ps Bhd & Wylie, Chromosome Atl., pr. 1, = (1955) and pr. 2, 323. 1961; Clapham, Tubing | & Warburg, Fl. t. Isles, ed. 2, Backer & «, Fl. Java 1965; Moldenke, Phytologia 13: 250 & 253. 1966; eal enke, Résumé Suppl. 1h: 10. 1966; Harborne in T. Swain, — Phytochem. 285. 15 Herbst Bros., Seeds oad Perham lh 2 | ~J e & Mh we & 53. 1966. dditional illustratio : Bodger Fls. pl. Shg, bile, ste alin color], 9568, » 1st, e sole [ AP cat ies aki (1928) rec the common zyo-zakura", "hana- cor and "siki- cat ide for a plant which he calls V. ogiflora Cham., cultivated in Formosa, but I suspect that his plant is actu- ally xV. hybrida. Bodger (1952) states that there are about 12,500 seeds of this per ounce. He lists V. hybrida grandiflora, V. flora ora Ye rightness, melee a affirms that in southern California the average dates for planting this subject February, nee July, and the harvesting of fruit in September, ing of the seeds in ber. The bst Brothers (1966) offer seeds of fo varieties: Ame + 1966, Crys te), t ( dor (purple and white), Starlight, and White. These are mostly de- scribed as dwarf forms » but also offered are Large-flowered Mixed Dwarf Upright Rainbow Mixed. ILLICITA Moldenke Additional bibliography: M Moldenke, Phytologia 13: as as (RE). KANSAS: Wabaunsee Co.: A, R. Moldenke iG, RE). MO TED ILLUSTRATIONS: A. R. Moldenke Kod. 1 (Z), 2 (Z), 13 (Z)- Addi opp — 966 tional bil Moldenke, 1 13: 250. 1960 Woolston describes this plant as ~ er Ae 3m. tall, erect a the flowers growing in wet wet places in low campos, and blooming in January. : Additional citations: PARAGUAY: Woolston 785 (S). 1967 Moldenke, Notes on Verbena 287 oe — ihe _ k. nal bibliography: Darlington & Wylie, Chromosome Atl., op. ices i; 2 (1955) and pr. 2, rs 1961; Moldenke, Phytologia 13: VERBENA LACINIATA (L.) Briq. A nal bibliography: Bocq., Adansonia 3: [Rev. Verbenac.] 203. 1863; Lefroy, Bull. U. Nat. Mus. 25: 9 88; Darlington & » Chromosom «, pr. 1, 323 (1955) and of 2, 323. 1961; acker & Fl. Java 2: 596. 1965; Solbrig : 17 B Bakh. 17h. 1966; Moldenke, Phytologia 13: 250—2 al,” 560, & 263. 1966. The Verbena laciniata described by Backer & Bakhuizen van den Brink (1965) from Javan gardens is V, tenuisecta Briq. as far as material seen by me is concerned. I doubt if the true V. laciniata occurs in Java, Bocquillon (1863) refers to V. laciniata as as Vv. erinoides Hook.", a binomial which I refer to V. berterii (Meisn.) pant datros (188) records V. laciniata from m Bermuda, probably Pee, ve Rufz Leal 7665/1h, distributed as V. laciniata, is ac- tually Vv. glutinosa Kuntze. VERBENA LASIOSTACHYS var. SCABRIDA Mol Additional bibliography: ee mre 11: 469. 19653 Hocking, Excerpt. Bot. A9: 36. 1965. VERBENA LASIOSTACHYS var ONALIS Moldenke J Additional bibliography: Moldenke, Phytologia 11: 169. 19655 » Excerpt. Bot. A9: 36h. 1965. VERBENA LILACINA Greene Additional bibliography: Hocking, Excerpt. Bot. A9: 36L. 19655 Moldenke, Phytologia 13: 20—205. 1966. VERBENA LINDMANII Additional pr Ppa : Hocking, Excerpt. Bot. A9: 36h. 19655 Moldenke, Phytologia 1 13: vip Pa 1966. LIPOZYGIOIDES Walp. tional synonymy: Verbena liposygoides Walp. ex Bocq., Adan- sonia otk 203. oe ore at ha 3: (Rev. Verbenac.] ogra: <5 ss: idee, Mae 203. 18 "18635 io Moldenke, Phytologia 13: 251. 19665 Mold . 1k: 10. 1966. LITORALIS H.B.K. . Verbenac.] Additional a Bocq., Adansonia 3: a h, 1: 13h. eect 1863; A. Stewart, Proc. Calif. Acad. Sci-, ‘Wylie, tl 1961; Moldenke, 13: 251—252. 19655 i + ly: 2. teen 3. see Biota 6: 107, 116, & 119- . 288 PHYTOLOGIA Vol. 1h, no. 5 Recent collectors have encountered this plant in pastures, a- long roadsides, in cloud~forests, and in somewhat dry roadsides with Caesalpinia, Salvia, Calceolaria, and Tagetes. Feddema re=- fers to it as "an oc canicind. @ eed", Wiggins reports the chromosome Seunt as n= 21 for his no. 18558, but makes the qualifying statement "one cell 90 percent zone, in partial shade among trees and shrubs, and Pood - the Scalesia Zone". The flowers are described as "lavender" i. L. Wiggins 18508, on Breedlove 1163) they were "blue", on Dressler & Jones 177 “blue-violet", an on ses 1667 "corolla pale-lavender, throat darker purple- Sampaio (1937) regards V. Sonera ae Vell, as synonymous with V. litoralis and ager. She xermestiar names "herva do pai Caetano" and therva A. R. Moldenke 1232, cited ice “under V. parvula Hayek, is re- garded by my son (the collector) as a mere ecotype of V. litoralis. He tells me that V. litoralis was growing commonly all through the area Lage this collection was made, with ‘ stature, and that the above-mentioned cbtection merely Peeper its waaliont alpine form. If this is an accurate description of situation, then it is possible that vais binomial should be ced to form status under V. litoralis. Additional citations: MEXICO: Michoacan: Feddema 1667 (Mi). Vera Cruz: Dressler & Jones 177 (Mi). GUATEMALA: HALA: Huehuetenango: Breedlove 1163) (Rf). COSTA RICA: Alajuela: A. R. Moldenke 1203 (Ac). Cartago: A. R. Moldenke 1320 (Rf). Guanacaste: A. R. Mol- . Limén: Ae Re R. Moldenke 1201 (Ac). COLOMBIA: Cundinamarca: aliases s.n. "Que- tame, Oct. 196L] (S). PERU: Cajamarca: Ugent & Ugent 342 12 (Ac)« Se AGOS te ome oe Itow 179 (Du--561,532) . Indefatigable: 5/57) (Ge—h619h8); I. L. Wiggins 18508 er iBes8 tbe SEkB39)" VERBENA LONGIFOLIA Mart. & Gal. Addit at bibliography: Moldenke, Ns ta 13: 251 & 252. 1966; Moldenke, Résumé Suppl. lj: 2, ib ke, REé- B Lsograpiy loldeaes, Pay Piptalce a cong 307. 1966; Moldenke, sumé Suppl. - 1966. cs Citations ; meriees Oaxaca: Breedlove 12292 (Z—type). Additional biblio om 365. 19655 ography: Hocking, Excerpt. Bot. A9: : , Moldenke, Phytologia 13: 207. 1966. = 1967 Moldenke, Notes on Verbena 289 DOUGALII Hell vo ga bibliography: err , Excerpt. Bot. A9: 365. 1965; Moldenke, Phytologia 13: 208. 1966. VERBENA MACROSPERMA Speg. Additional bibliography: Moldenke, eat 10: 119—121. 1964; Hocking, Excerpt. Bot. A9: 365. VERBENA MARITIMA Small Additional bibliography: Hocking, Excerpt. Bot. A9: 365. 19655 Moldenke, Phytologia 13: 208. 1966. [A MARRUBIOIDES Cham. Additional bibliography: Hocking, Excerpt. Bot. A9: 365- 196535 Moldenke, Phytologia. 13: 208. 1966. A MATRITENSIS Moldenke Additional bibliography: Moldenke, Phytologia 10: 129. 196h; Hocking, Excerpt. Bot. A9: 365. 1965. MEGAPOTAMICA Spre wets dditional bibliograohy: Darlington & Wylie, Chromosome Atl., pr. 1, 323 (1955) and pr. 2, "323. 1961; Hocking, Excerpt. Bot. Ag: 365. 19653 Moldenke, Phytologia 13: 208—209. 1966. MENDOCINA R. A. Phil. nal bib ton & Wylie, Chromosome Atl., ce) Bry 2 323 (1955) vas br 2, 323. 1961; Moldenke, Phytologia 13: VERBENA MENT LIA Benth . Additional bibliography: Moldenke, Phytologia 13: 252 & 276~ 1966; Moldenke, Résumé Suppl. li: 2. 1966. in association with Solanum michoacanum, S. tamm, S. Stolonifermm, Physalis, Gaura, Oenothera, cum. i Additional citations: KEXICO: Chiapas: Breedlove 11161 (Ac). Michoac&n: Ugent, Ugent, & Flores C. 5802 (Ws). Sinaloa: Fed- Gema 1808 (Mi). tho nal SG eae ganinat ee graphy: : [Rev. Verbenac ol] 203. ue Moldenke, Phytol se ‘Bs 252, $2, 25 7x62, & om. 11966 - Pe ay tors ate h Solanum -_ open village fields, on pare hte panks wit acaule, and on vast ——— grasslands with a few round cacti, Solanum acaule, Tetraglochin (strictun?), and a few other small shrubs, with ith calcareous outcrops. 290 PHYTOLOGIA Vol. 1h, no. 5 Additional citations: PERU: oon pes & Iltis sn, [10 km. SW. of Puno on road to Llave, Jan. 10, 1963] (Ac); Titis, Iltis, Ugent, & Ugent 1376 (Ws); Iitis & gent 1393 1393 (Rf). VERBENA MINUTIFLORA Briq Additional bibli apni: Moldenke, ee 13: 252. 1966; G. saviors Ind. Kew. Suppl. 13: 143. e flowers on Reitz & Klein 17805 are “described as "roxa" ee citations: BRAZIL: Parand: Reitz & Klein 17805 (Ac). mea Maldente Adas graphy: E. W. Lathrop, Univ. Kans. Sci. Bull. 39: 173. "i958; Rocking, GEORGIA: Baldwin Co.: He Ne ee Ts 23036 (Ac). JAMAICA: C , Hespenheide, & Anderson Webst Webster & Proctor s2h9 (il » BRAZIL: : Reite & Hein 17672 (1 isle. VERBENA RIPARIA 2 j Additional binri crea Moldenke, Phytologia 11: 179. 19655 296 PHYTOLOGIA Vol. 1h, no. 5 Rickett, Wild Fls. U. S. 1 (2): 362. 1966. VERBENA RYDBERGII RIMES Additional bibliography: E. W. Lathrop, Univ. Kans. Sci. Bull. 3921735 Sofa. Molderke, Phytologia 13: 20-261. 1 One eA. R. Woldenke enke 819 specimens, cited previously as 7 ane in my personal herbarium, is now in that of the Texas Research Foundation at Renner, Texas. Lather (1958) tells us that this plant may be expected in overgrazed prairie pastures, i , and this agrees completely with the observations of my son and mys ° Additional citations: KANSAS: Dickinson Co.: A. R. Moldenke 1298 (Rf). Jackson ay A. R. Moldenke 131), (Ac). Osage Co.? A- R. i akaldios 130 ae) Sans Nae eae Cost Aw Bs Moldenke 1260 MOUNTED I IONS: A. R. Moldenke Kod. h (Z), 5 (2) e( (2), Ta (Z), Sy ah fs (z), 12 Pia yb (Z). VERBENA SANTIAGUENSIS (Covas & Schnack) Moldenke Additional bibliography: Darlington & Wylie, Chromosome Atl ogr. CG Rees toe? (1955) and pr. 2, 323. 1961; Moldenke, Phytologia 13: SCABRA Vahl Additional bebliogranty: Moldenke, eytlewia 13: 261. 19665 Rickett, Wild Fls. U.S. 1 (2): 36h. In Jamaica this species is webitlnd to be "common in the drier areas of swampy country". The flowers on Webster & Proctor 533k are described as "lavender", while those on Webster & Wilson 5051 we were "pale-blue". The two Sevier collectors describe the plant an annual. ald tinea citations: LOUISIANA: Orleans Par.: T. Drummond S-Re (New Orleans, 1833] (S). JAMAICA: Webster & Proctor 533k (ii “is Webster & Wilson 5051 (Mi). SCABRA f. ANGUSTIFOLIA Moldenke, f. wal. Haec forma a forma typica speciei laminis meold orum anguste This form differs from the typical form of the species in hav" ing its tien narrowly or very narrowly elliptic or elliptic eolate, the i with the margins sharply and irregularly mucronulate- serra The type of the form was oat tected by John R. onsen (nO: 1837) in moist black loam at the edge of Beaver Cree 1967 Moldenke, Notes on Verbena 297 Maren Texas, on July 7 1966, and is deposited in oe Lundell Herbarium at Renner, Texa, The collector states t the flowers are light-pink. While eaves approximating these in size and Shape are sometimes seen in and near the young inflorescences in the typical form of the species, in the present form uniform to 76: 22 & 23, pl. 2, fig. 12, & text fig. 26. $e oe Nat. : . Taiwan PF mi nage 19625 feck 1967 Moldenke, Notes on Avicennia 329 Suppl. 803. 9625 McLuckie & McKee, Austral. & N. Z. Bot., pr. 5, 88, lbs, 459, 60, 658, 659, 662, & 700, fig. i 1962; Moldenke, Dansk Bot. Arkiv 23: 85. 1963; 1d, Woody Pl. Taiwan 15, 817-818, & 942, fig. 329. 1963; H. P. Riley, Fam. es Pl. S. Afr. 128. 33 "Legris, Trav. Sect. Scient. Inst. Frang. Pond. 6: 323, 547, Te & 556. 1963; Backer & Bakh , Fl. Java 2: 61h. 19655 Antes Biol. Abstr. 7 17): 18. 3 J. S. Beard, Descrip. Nee Pl. 91. 19 cnae, Austral. Journ. Bot. 1h: 67, & Th——76. 3 Ma 1966; Moldenke, Phytologia 14: 309. 1907s Addi tio nal & gir illustrations: W. Griff., Trans. Linn. Soc. Lond, 20: pl. 1, fig. 8-12. 1816; Talbot, Forest Fl. Bombay 2: fig. 457 Y (ai ¥ albal® 1911; Kanehira, Form. Trees, rev. ed., Sia pet 33936: Nair & Rehman, Bull. Bot. Gard. Lucknow 163 pl. 2, & text f : ss Introd ustral. & 1652 Bot., pr. 5, fig. 31. 19623 Li, Woody Pl. Taiwan fig. 329. Recent collectors describe this plant as a shrub, 1 m. or less in height, or a tree, 10—15 m. tall, the bark white or blackish Nair & Rehman Talbot (1911), who misidentified this pie as A. alba Blume, describes it as follows: "A shrub usually grvoved, angular, fi white-tonentose. Leaves ovate-lanceo- late, soe by [2 —1", acute at and base; evohted above, gray-tomentose beneath. Tnflorescence as in th species” Th offici erty ]. Stamens included. Ovary and style long, smooth, yel- villous. Capsule ovoid, Speier. ee xy Low when nee Closely allied to A. officinalis. India in ti- dal g Dna. cry Ae ommon throughout the Konkan; on the trap near the pparently throughout the Swanips, ee ie Bandra, Bombay. Fl. a bo & Bakhuizen vam den Brink (1965) report that in Java this plant grows in the "mangrove (also in the more saline parts), ring through the Pair som ther ones eles etimes dis from the 0 Jed 5 cm longs sige robust » C- 11/2 m long; stigmas an obtuse or 1 flowers were "dark-yellow" on Steward 3112, On Smitinand 5641. The latter Zollection is described as paged had "blackish bark", while on Smitinand 5642 the bark is said have been "white". This collector calls the : suas and reports the ar name "samae thale". 330 PHYTOLOGIA Vol. 1h, no. 6 Fron Ee wenewe East Africa we have reports of the names "mocho", ", "musso", and "staca", Riley (1963) reports that this species, which he misidentifies as A. officinalis L., is called ‘white mangrove" and grows in Mozambique, Natal, and the eastern such cig of Cape Province to the Kentani Division." n general, the distribution of Ae marina na may be said to ex~ tend from FESS to Arabia, along the shores of the Red Sea and the Oc Goo: western Indi ean to the Cape of H eastward along the shores of the Arabian Sea, the Bay of al, the northern and western Indian Ocean, the south China Sea north to Hongkong and Formosa, and the islands of the Philippine Sea the South Pacific to New Zealand. Beard (1966) reports that in seeded Australia it is called "white mangrove" and the flowers Doty & & Alvarez 14342 is described as having had a og algal coating on trunks and igor os no prop roots" -— metoeran of the trees in situ i unted on the sheet with the - Pierre s.n. (Siam, 1868] » was first a pea: as A. eres L. by ‘someone, then later as A. marina var. media (Griff.) Bakh. A note appended to F. A. A. Mendonca ie a- vers that it matches Burtt 155, R. M. Graham 251, J. W. Grego $.n., Hildebrandt 12), Mearns 215 & 2154, Schlieben 257, and Volkens 160 in the herbarium of the British Museum. In Biol. Anat. Abstr. (1966) it is aha that a new species of fruitfly has esting this ydney area the first colonizers of bare md fiat are Avicennia "of- ficinalis" and the shrub Aegiceras majus; above this is the saltmarsh of Salicornia australis; Save: the tide level is the grass zone of Sporobolus us virginic us, the rush zone of Juncus maritimus, and the sedge zone of Cladium junceum, then the sa- line she-oak forest of Casuarina jaarina glauca, 4 and then the freshwater swampy forest of Buc robusta. Hochreutiner comments as follows: "Dans quelle mesure l'A. serine différe-t-il spécifiquement de L'A. officinalis? L' examen nombreux spéc: nous laisse dans le doute a ce sujet, un eS resi de t décider. La forme des quée par Vierhapper est non seul suivant les indi- nom’ 1967 Moldenke, Notes on Avicennia 331 — ie aussi caractéres aléatoires. En revanche, le degré de de l'ovaire et du style, invo par Bakhuizen nous a paru “an difficile 4 observer et laisse souvent planer de doute au sujet de 1! soe age ie seco En réalité ltestrémité du style est toujours glabre. De sorte qu'il semblait plus judicieux de se contenter en Liceiabinaant d'une distinction variétale. t a la distinction variétale adoptée par Bakhuizen dans 1'A. marina, nous l'avons appliquée d'aprés ses descriptions mais, on se i gro compte combien elle est précaire, en lisant sa clef dont le premié e invoque un caractére ¢ = Sap et base morphologique. Nous serions tenté plut6t d'tadopter le point de vue de S r (Bot. Mitt. aus den Tropen Ill us les spécimens de 1'Ancien monde ala. officinalis, — distinctions ean seraient pees probablement enter ear ns express in Rumphiana & cause de leurs pétioles trés courts car c'est 1a, en earhiace la _ différence réelle que Bakhuizen reproduise sur s The F, pote (185) reference given in the bibliography above is often cited to Hochstetter, but the article in question is cre original! Nair & apertecs Hoes) ate « eke ee Bot. Gard. 4,828, slide 3122, from India, while Li (1963) cites Kori & & Morimoto 8. S.De from For- mosa. The Yates 1110 distributed as as A. or arina is actually A. al~ ba Blume, while Yates 2075 is A. officinalis ~~ Additional citations: a He Ge F paniaae 1574 @), 2 1591 (B); Tanner 26 (B, S), 342i (S)» PORTUGUESE EAST EAST AFRI Delgado: F. A. Mendonca soe tua), 1076 (U1); Torre & Paiva 9616 (U1). Inhambane: Gomes © Sousa 2096 (U1). lLourengo Marques: Bal- Sinhas 131 (U1), 232 (U1); Gomes @ -e@ Sousa 155 (U1), 3793 (m2); Je Su- nod 517 (U1). Mozambique: Torre e Paiva 1148) (U1). : » 50 nm Natal: J. M. Wood 12973 (Bi). INDIA: Madras: E. K, Janaki 311 (Mi). CHINA: Fukien: tes Steward se et sk INDOCHINA: Cochinchina: Pierre s.n. {Hataen, 1/187h] (Ca— ° Pierre s.n. [Siam, 1868] (Ca—5l863)5 Smitinand 5044 561 [Herb. Roy. Forest Dept. 22952] (Gg), 5642 [Herb. Roy- Forest Dept. 22985] (Gg). WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Cebu: Doty & Alvarez 1432 (Ac AVICENNIA MARINA var. RESINIFERA (Forst.) Bakh. : Additional & emended bibliography: W. Leh Linn. Soc Lond. 20: 1-7, pl. 1, fig. 1—7, 13, & uh. 18h6; F. M. Bailey, Wiss, Wien 89: , 191hz3 Ro T . Miss, Wien 89: 569, fig. 29. 290s Ss BL PALET*, iB, i5l6s Bal 332 PHYTOLOGIA Vol. 1h, no. 6 - & Lam, Nov. Guinea 1), Bot.1: se inet J. M. Black, Fl. South Austr. 3: 83—~8h. 1926; Moldenk Phy toLogial 7s 227— 232 & 259262. 1960; Moldenke, Biol. aisle, 35 ~ oe (1960) and 3%: 719. 1961; Hocking, Excerpt. Bot. A.5: 45. dastsonal evar W. Griff., Trans. i Soc. Lond. 1—7, 13, & 1h. 1846; Rechinger, Denkschr. Akad. = = wProos Roy. Soc. N. S. Wales 9: ae 35, lb, l6, & 19 Recent collectors describe this plant as a tree, 8m. tall, or prostrate, with yellow flowers and light yellow-green fruit. It b een "wallum" with Pimela, Leptospermum, Goodenia, Drosera, and Lobel ja, at 2 meters alti altitude, flowering in December and fruiting in February. The flowers are described as "yellow" on MacDaniels 250. E. H. Walker 5352 bears a notation "prostrate form". Bailey (1913) misidentifies this taxon as A. ene Le = records So gee oe Ver neralas names "egaie” seem Oo". imilarly misidentified by ecu Gsu), Baker “A9i6), Black oe and Laing & Blackwell (29h0) k tells us that it is n mud close to the sea all a sending DOWN roots; Avicennia sends UP aerial roots." Laing & Blackwell report that _Avicennis has an embryo with 2 fleshy coty- ledons unaccompanied by any trace of a rootlet. Therefore it has to germinate by rs erctas fit roots only. They say, inaccurately, that "A. officinalis occurs also in Australia, throughout Melan- coast north of Kawhia on the west and of Opotiki on the Rechinger (1914) notes "Mit auffallend schmalen Laubbluttern.. Solomons-inseln: Insel Buka. Auf dem Strandriff mit Aegiceras corniculatum Blanco und Rhizophora-Arten." Baker us a cusses the morphology and wood anatomy and records Fig bales His work is sometimes inaccurately ery ae an her of this variety has been p Aagretigy tse and distribu- ted tn herbaria as A. officinalis L. On the other hand, the Ka- jew ski 2344, in part, distributed as ‘. poner var. resinifera, is actually A. emcalyptito olia = EAP: Additional citatio CALEDONIAN I New Caledonia: MacDaniels 250k (Ca-51017) ; H. S. McKee 2114 iil (G0) « AUSTRALIAN REGION: AU: AUSTRALIA: Queensland: | Ma Ss. . Clemens s.n. [23 ere aed nn pt Rid South Australia: Kaspiew 36 (S), 539 (B)- orth Island: E. H. Walker 5352 (Ca-—923408) - | Rangi- 1967 Moldenke, Notes on Avicennia 333 toto Island: Meebold 5233 (Bi). AVICENNIA MARINA var. RUMPHIANA (H. Hallier) Bakh. Additional & emended bibliog raphy: Lam & Meeuse in Holthuis & Lam, Blum span ime 235. 19455 Moldenke, Phytalogis 7: 262—266. 1960. Lam & Meeuse (1945) record this plant from Karakalang Island in the G er ees Sunda Is Aes Additional citations: WESTERN PACIFIC ISLANDS: PHILIPPINE IS- LANDS: Mindanao: Elmer 10)5a (Bl), 12008 (Bi). MELANESIA: GREAT- ER SUNDA ISLANDS: British North Borneo: Cuadra s.n. [Herb. North Borneo Forest. Dept. A.3187] (W--2129128). AVICENNIA OFFICINALIS L. Additional synonymy: Avicennia officinalis Ue ) Kurtz ex oe Kew Bull. mee Inf. 1938: 443, in syn. 1938. A 1 W. A. Talbot, Forest Fl. Bombay 2: 361—36h, fig. 456. 1911} & Tutcher, Kew Bull, Misc. Inf. Addit. Ser. 10: 205. 1912; J. q - 1913; R. T. Baker, & Proc. Roy. Soc, N. S. Wales 257—281, pl. 35» 4S, ie, & & 48. 1916; Bose, Man. Ind. Bot. 253, fig. 220. 1920; H. N. Ridl., Journ. Fed. Malay States Mus. 10: 151 & 152. 1920; Despeissis, West & Trop. North, ed. 3. 1921; eared Re irs id. Madras 1105 & 1106. 192h3 H. J. Lam in ss 3: is 2 59: 29. 192h3 EAS Lever gen Austr. ° asaki, Li + Se “iB sl. 1965 oi Pflanz enanat. 10 (2): 233 & 380. 192) Schencke, Zytol. Untersuch. Verbenac. 7 & 31. 19715 Kudo, Bots, Mag. Tokyo 6: 15h. 1932; Patermann, Beitr. Zytol. Verbenac. h6, 48, & 49. 1935; Jovet, Bull. Soc. Bot. France 82: 219. ai Fletcher, Kew Bull. Misc. Inf. apn ay oe Bull. Misc. Inf. fee index 27. ‘0 3 oh ins tan moat to 7: 267 . 1960; Santapau, Fl. Seca zor 290 4 293. : ke 3, 1605 San *ourr. Sci. India 30: 424-25. 33h PHYTOLOGIA Vol. 14, no. 6 1961; E. L. little, Phytologia 8: 50--52 & ee ade ptern Bip: ’ Pakist. Journ. Forest, 11: 43—h5. 1961; ¢ - Jard. Bot. 203. LZ 23 Nair & Rehman, Bull. Bot. Gard. Lucknow 76: 22. 1962; Liu, Tllustr. Nat. & Introd. Lign. Pl. Taiwan 2: 1201. iA McLuckie & McKee, s: Acad. Sci., ser. B, 58: 45—-50 & 351, fig. 1~20. » Trav. Sect. Scient. epee emp paeee he OP. 323, L7, & 556. ri He & E.B ser. 2, an Steenis - Males. "poll. 193 1133. Dek. Feretee> Biol. Abstr. te 257 & B.13. Mae, B. we Bull. Bot. Gard. Lucknow 89: 1). 1964; Backer & Bakh., Fl. Java ny “oh. 1965; Moldenke, re Suppl. wer “is 1965; Mani, Bull. Bot. Surv. India 7: 1i,. Addi tional & emended illus ee W. A. Talbot, Forest Fl. mend 2: fig. 456. 1911; Bose, Man. Ind. Bot. fig. 220. 1920; ee a 3g eee Forest. 11: 43—h5. 1961; oh ea? Proc. - B, 58: layer; medullary rays white, unifom; weight 66 pounds to the cubic foots bark contains irewoo He ords it tannin; "used in India only for f a", as @ swamps of the Deccan Ceylon, common in salt marshes and along the banks of tidal rivers creeks, thro t the Konkan and K4nara, ering from March to May, the fruit ripe in July and August." In his 1911 work he erroneously places A. tomentosa Jacq. in the synonymy of ae oo ey icp that the thick-walled lignified pit- tem These are actually transformed parenc cell the sten, or mesophyll cell in the atch and lam- ina of the : leaf. They are formed by secondary sclerosis of these parenchyma cells. Baker (1916) also discusses the anatomy as well as the morphology, but his plant was probably A. seals . ae apetnitare (Forst.) aa ae & Rehman (1962) give e size o pollen grains as Patermann (1935) says: "Treub (8825 eo Avicennia officina- ida; dass die Verbenaceen in der Endospermwicklung wes wesentliche hiede zeigen die ohne Zweifel flr die verschiedenen Unter- familien charakteristisch sind....Immerhim dttrften wir in der 1967 Moldenke, Notes on Avicennia 335 Endospermwicklung von A. officinalis den spezialisiertesten Typus finden und dadurch auch h zytologische die Anordnung dieser Unter- familien (av neniitieA dine) als zu Recht bestehend best#tigen."” Dastur (1952) tells us that in India and Pakistan the astringent bark of this tree is used for tannin, the wood ashes for washing cuties » the green fruit as a poultice in the treatment of boils, and the ripe fruit as food; painters mix the ashes with their color pigments to make them adhere geet the wood is used to gr Hives beams and door-frames, but chiefly as . ani (1965) describes a subglobose, agglomerate, solid, fleshy, B Saat pouch-gall made by an Eriophyes Sp-, 2 mm. large, the outer surface densely clothed with hair-like parenchyma emergences Additional vernacular names recorded for this plant are "héi- -tang", "h4i-ka-tin", "hirugi-damashi", "hirugi-damasi", and mahi & synonym of A, marina (Forsk.) Vierh. Lam's note in Engler's Bot. Jahrb. (i92h) is 4s often erroneously cited as "1925", which is the title-page date for the completed volume. Boyko (196)) records this species as cultivated in Egypt, but I strongly sus~ pect that he is referring to A, marina. The ma, officinalis L." of F. M. Bailey (1913) and the vernacular names which he records from Queensland doubtless belong under A. marina var. resinifera lection actually is i asad G. F. Gaumer 635 nagit “abel IE Sass Schimpff 30, Snod- 11 A. Stewart 3266, 3267, 3268, 3269, 3270, 2a, 3272, 3273, & re distributed as A. A. officinalis, are actu- ally A, germinans (L.) L.; B+ K. Janaki 311, F. A. Menconee 1015, Pierre sn (Siam, 1868], and J. M. Wood 12973 are A. i. marina (Forsk.) Vierh.; H. S. McKee 211) and Meebold 5233 4r® A= var, resinifera (Forst.) Bakh.; and Elmer 10L5a & 20S ©°" ==* serine ver. rusphien (i oasrege "4 : H. St John 24 066 (Bi) “TT paaisohel citations: INDIA: o Ube 2066 ‘ MALAYA: Singapore: M. R. Henderee 34770 ( )(S). WESTERN PACIFIC n: Elgincolin s.n- s.n. (Herb. Philip. Kondo & Edafio Fo 92h2 Saat Fi d 2 ay af i ra: 2075 (Mi). ; NEW GUINEA: Papua: A. Hart ner te —- “Guinea Forest. Dept. 4530] (Bi) - Additional synonymy: go che tomentosa sensu Gooding, Loveless, & Proctor, Fl. Barbados 365, in pri ee 336 PHYTOLOGIA Vol. 1h, no. 6 dditional biblio : Moldenke, P ogia 7: 280—290. TOO Aneeny) Fl. Serer nbs 38 (1960) dias 15. 1961; Reitz, Seezenka LF 29, » & 109. 1961; Angely, Bibl. Veg. Paran. a5 - 196k; Moldenke, Résumé Suppl. 12: 3. 1965; Gooding, Loveless, , Fl. Barbados 65-366 & 65. 1965. Additional illustrations: Reitz, Sellowia "13: hS & h6. 1962. Reine (1961) calls the association in which this species grows Shearman a Hahn 654, cited below, is a mixture with A. germinans Additional = set a saree ISLANDS: Martinique: Hahn 65h, in part (Ca-~332)88). TRINIDAD: Cowan & Forster 1252 (N). BRA- ZIL: Cear&: Brouet 258 eee Guanabara: Ae kh. Castellanos 23460 [Herb. Cent. Pesq. Florest. 1720] (Ac); Scheiman 108 (Ja-— 12);208) ; J. Vidal sen. [30/5/193k] (Ja—36839), s.n. sen. [21 de Junho de 193k] (Ja—36838, Ja). Rio de Janeiro: Abreu s.n son. [VIII/1925] (Ja—31572); Dusén 1963 (Ja—l852); Glaziou 1362 . (Ja—11185) ; Schwacke 3111 [Herb. Saldanha 5962] (Ja--l)066, Ja, Ja), son. Daas, ae bial (Ja—l865); Ule 388) (Ja--37)3h) . Santa : 5088 (N); Reitz & Klein a (W--2281838), 1183 Cece F Fikes Island: Magalhfes Corr8a s.n. (Oct. 1936] (Ja—30128). Gobernador Island: Rente & grat sen. (18.11 21958) (Ac, ine Ja, Ja). Paquet&é Island: Descartes 7 (Ja—36350). Saravata Island: J. Vidal s.n. [Junho 193k] (Ja—31546, Ja, Ja)- AVICENNIA TONDUZII mM Additional 35 smn Moldenke, oN 7: 290—292. 1960; Moléenke, Biol. Abstr. %: 79. Additional citations: COSTA RICA e halectdedids Tonduz 6776 (Mi-~isotype) . ADDITIONAL NOTES ON THE GENUS PRIVA. IV Harold N. Moldenke PRIVA Adans. Additional synonymy: Tertula "Roxb, ex Willd." apud Brenan, Mem. N. Y. Bot. Gard. 9: 37, sphalm. 195), Additional & om biblio A. Le Juss., Ann. Mus. Nat. Wie Maks Vaehe\ ta agcee. Aprec ; ‘aris Linn, Ind Cat. Hort. + 2: 47. 1843; F. Krauss, Flora 28: 68. rg alba, Be beg pert 6: 687. 187; D. Clos, Ann. i soe 7109 . Bocq., Adansonia 2 lene Ph-—99 é a8, oo: 125, 190 , 1365 “139, 140, us6—iig, 152,"& 15h (1862) 1967 Moldenke, Notes on Priva 337 and 3: 180, 182—185, 211—213, & LA a — - 1863; Bocq., Rev. Fexbenas 8 ae J—99, 101, 109, 116, 125, 130, tig 3, ibe ake” 152, "sh, "180, *spotabs, Bocce 5 pi. my 1863; Griseb., Cat. Pl. Cuba 21h. 1866; Et ann, Enum. Hawai. Pl. ge 1867; Aschers. = .e Schweinf., Beitr. Fl. Aethiop. 278. 1867; R. A. Phil., Anal. . Chil. 35: 193. 1870; A. Gray, Syn. Ye wl tae 2 (1): 333 & "33, 18785 Franchet Sert. Somal. 50. 1882; A. S. Hitche., fee Mo. 117. 18935 Just, Bot. Jahresber. 23 (2): yr, 96 & “S05. “38975 ay %, 1900; Durand & uppl. a 207. 1921; dente: Fl. maces ioae & & ‘opomton. “192; Britton & P. Wils., Scient. Surv. Rico 6: 137 —1y5. 19253 A. W. Hill, Ind. Kew. Suppl. bs "166 (1926) and - ik 1929; Baeza, Nomb. Vulg. Pl. Silv., ed. 2, 63, 176, & 263. 1930; Schwe NAC « & 25, pl. cane & = eee - 5. 19353 Chiov., Racc. Bot. Miss. Consol. Kenya 98. 1935; H. B. e e orks Pringle 669. 1936; A. W. Hill, Ind. Kew. Suppl. 9: 223. 1938; Robledo, Lecce. Bot. 2: 99. 190; Schnack, Inst. Fitotéc. Sta. Cata~ lina - 18. 1942; Schnack & Co : k7l. ry: Dar- some 70. 70. 1915 Hill & r Salisb., 2956, 1955; Moldenke in Humbe rt, Madag. avn 3 & 06— bia: Moldenke, Inform. Mold. Set oe Spec. h. 1956; aay. ‘Cat. Anon., Biol. ihatr'. 29: 3h69. it . Bot. Fan. 17: 5. 1956; 1957; Alain in Leén & * 1. Cuba i280 & 302. 19575 Be He Johnson, Wasmann Journ. 16: 295. ag Pog Ind. ark sumé 1 ho—l 1959; Moldenke, Ré j ab 8 Bt, 16h, ih. Wess zt on 6). 68, 73, To— ac? dl Je , > 7 r) ry ’ ? ’ , ir tte ale ig a a 166, 167, 190, 222, 237, 238, 210, 2795 33lt, 20, 30%» i 67, & 468. : 4 3 ata, Bo 2? erp ag vi ha gely, Liv. Gen. Bot. . 35 & 52. 1960; Anon., Assoc. 5 Tax. Fl. : 60. Darlington & Wylie, Chromo~ ht Sig” ones &0 An. Journ. Bot. some i 48: 62, 1961; Runner, Rep. Ge We ‘ .« 1962 Bull. Jard. Bot. Brux. 32: Suppl. 794—795 re oe takckinaon & Bo Codd, & Letty, Wild Fls. Transv. 280 19625 Hepper 338 PHYTOLOGIA Vol. 1h, no. 6 Dalz., Fl. W. Trop. Afr., ed. 2, 2: are ee 1963; Rollins, Tax- on 12: ae. 1963; Huber,” Hepper, & Meikle in Hutchinson & Dalz. ‘ Fl. W. Trop. Afr., ed. 2, 2: 432. 1963; H. P. Riley, Fam. Flow. W. 34 Bol. Soc. Brot., ser. 2, 39: 131--132. 1965; Anon., Assoc. Etud. Tax. Fl. Afr, Trop. Index 1965: 67. 1966; Anon., Gen. Costa Ric. Phan. 10. 1966; G. Taylor, Ind. Kew. Suppl. 13:18, 109, & 19. 9 To be added to the list of excluded species: Priva mitchelii Endl., Cat. Hort. Acad. Vindob. 2: 47. 183 = ieptostachya L., eae om" (1963) gives the basic chromosome number for the genus Priva 2. The Hooker (1829) yaternes ori fin the bibliography above 4 ee mis-dated "1830", The "Bot. Zeit. 3: 587 Spr <9 and Flora 29: [207] (1846)" references sometimes given for gemus appear to be erroneous, since there is no gantiod of the genus on those pages of the works in question PRIVA ADHAERENS (Forsk.) Chiov. Additional synonymy: Verbena forskaelei Vahl ex Willd., Sp. Pl. 1: 118-119. 1797. Verbena forskaehlei Vahl ex Steud., Nom. Bot., ed. 1, 657, in syn. 1821. Verbena forskohlei Vahl ex Priva Steud., Moi Bot., ed. 1, 873, in syn. 1821. rens Chiov. apud A. W. Hill, Ind. Kew. Suppl. 7: 197. 1929. Priva septestachys Auct. ex Cuf., é 5 - 32: Suppl. + 794. 1962; Moldenke, Résumé Suppl. 12: 7. 1965. Krauss (1845) ‘cites Krauss 420. Smith (1817) records the common name "Arabian vervain", Additional & emended citations: SUDAN: Lord s.n. [Upper Camp, 1967 Moldenke, Notes on Priva 339 Hot Tamarib, en (Ca~—331299). ANGOLA: Mossamedes: Exell & Mendonca 2121 (U1). SOUTH AFRICA: Natal: Kuntze s.n. (Durban, TIT.9h] (W—633155). PRIVA AFRICANA Moldenke Additional bibliography: a Ind. Kew. Suppl. 10: 18h. 197; weldecke. Phytolo aF ae 108: 195): eter eae In~ 1953. Mold. Set 51 Spec. h. "F586; Moldenke, Résumé 15) & 167 my 1936 work, on page 36, there is an incorrect or of sights’ in the Latin pete! of this paisa, The p should read "dentibus remotis 8 Ss. weg sublovosdeie®. The Britton Herbarium or in my personal herbarium. aoe eencene Brees forming dens stemmed clumps 30- - tall, a 12-inch shewhleh. growing on Siaoeis veld on rch A. ppt in dry sandy Acacia thornveld in open Places and shallow pans, and in open spaces in dry sparse thorn- veld on alkaline and probably calcareous soils, at 1800 feet al- titude, blooming in March and December. The flowers are descri- bed as "pink" on Meeuse 10222, as "mauve" on L. E. Sede 86, and deep sandy $2, probably by preference on slightly brackish soil. Flowe rs large, white. Bs r Priva species I have collected » etc.” grow in light shade, have pinkish flow & emended a haended "PORTUGUESE EAST AFRICA: Lou- 1423 Trans-— ional renzo matooads Myne & C (Ul). vaal: L. E. Codd 3886 (Cb, Ss); Meeuse 9477 (Z), 10222 (s); van Van Dam son. (Dec. 192k] (Ss). PRIVA ANGOLENSIS Moldenke Bibliog graphy: Koldenke, Bol. S00» Brote, see = ee i Moldenke, Résumé coeds 17 12: 7. ry ping , Assoc. BL Afr. var L. Robinson apud A. W. Hill, Ind. Kew. Suppl. 8: 19). 1933. Priva grandiflora Moldenke ex J. Rzedowski, Anal. Esc. Nac. Cienc. Biol. 6: 105. 1954. Priva rhinanthifolia as & G.) ve, in herb. cir ges bibliography: Walp., Repert. 6: 687. 1847; Jacks. in Hook. f, & Jacks.,, Ind. Kew. er 628. Ar M, Kunz, Anatom. h. ns 55. 1912; Perry, Ann. Mo. Bot. Gard. 20: 342 & A. » ind. S : Anal. Inst. Fitotéc. Sta. Catalina : 18. 1942; Schnack & Covas, ana 6: 471. 194k; Darlington & Janaki ir Chromosome Atl. 270. 1915; E, J. Salisb., Ind. Kew. Suppl. 1: 202. 1953; J. Raedowski, Anal. Esc. Nac. Cienc. Biol. 8: x08. "1953 Moldenke, Phytologia 5: 72——73.(195k) and 6: 239. 1958; Moldenke, Résumé 38, 222, 3h0, 3hl, 355, 365, 373, 377, & 68. 1959; Moldenke, Phytol : 183 & ita ogia 8: 183, 27, (1962) and 9: 307 & 315. 1963. Recent collectors have found this p growing in grass by cornfields, in with scattered pines or stony outcrops, in roadside ditches through the rete in pedregal, and in dry rocky level grazed Acacia-Hilaria~Aristida-Opuntia a grasslands With many herbs like Calochortus, Tuellia, Oxalis, Artemisia, Salvia, Mimulus, etc.; also on stony ee a savannas, gentle slopes on reddish sandy loam + mountain tops, and "under oaken hills", at altitudes of 2000 om * 9665 neters. Hinton describes the plant as "procumbent", while Waterfall notes "roots fascicled", The corolla is described as "pale-lavender and white blotched” on Weintraub & Roller 173, "lavender" on Hinton 1320k, "Light- pink" on Waterfall all 13400, and "pale-bluish, nearly ly white” on R. McVaugh 18278. Schnack & Covas (19h) report the chromosome — er = eateetin id been misidentified ish bet ap in herbaria as Acalypha sp., Valerianodes sp., and Verbena sp. Additional y aeavial citations: spp Pi Aapseeeadenet, R. — Vaugh 18278 (Mi); J. Rzedowski 1,011 (I Chihuahua: Goldman (W—~33560 IF tasees iex.95 (Ca—712708, W-~1887830); Pringle 154) ringle 1549 (Ca—104892), 3057 (Gg—L21280, N, W—2296117). Durango: x. C. Johnston 2661, (Au--231742, Mi); E. W. Nelson 4635 Falne: ), 2h (c 118770, E—-118771, W-—-304369); Waterfall 13400 (Gg), 13476 (Ce), 15433 (Gg), 71488 (Ca); Waterfall & W & allis 13358a (St), ), 13400 (St), 13476 6 (St, W—2297161), 13539 (St)- aeedoal Dist District: Balls B & 1923 (Ca—68128), B.5665 5 (oe SOEs Oreutt s.n. [Oliver] ig "4 281902); Rose, Painter, & 9496 (W—LS . Hidalgo: exia 2 2743 A -BRT EE, “F--1023108); Salazar sn. [ 350 PHYTOLOGIA Vol. 1h, no. 6 Aug. 1, 1913] (W--1169845). Jalisco: J. N. Rose 3576 (W--302553); Setcicacks & Roller 173 (Mi). México : Bourgeau au 357 7 (W—-209558) ; Galvan sn. [23.V1.1963] chs); Paray 863 3 (Ip), 2957 (Ip); Rose, Painter, & Rose 822 (W--lS1916); Salazar s.n. [Teoloyucan, 1» Aug. , 1913] (W--1013206, W—11698,6). Michoacdn: Arséne 3499 (W-- 1003701), 8296 (E--841342, W—1003669); Hinton 1320) (Mi, Rf, Ur, W=-189108); Iltis, Koeppen, & Iltis 530 (S); Pringle 4147 (Ca— Park 1187 9, im—1 06, W—57689, Smet CULTIVATED: Herb. Pernhardi s s.n. [Hanover] (E—1180)8). MOUNTED IL- foeairsaie Kobuski dra anand oe ring 13 (E--925)06), oo (E--925),05) « PRIVA HUMBERTI Moldenke Additional bibliography: Moldenke, mye oe 5: 73. 195h5 Moldenke in Humbert, Fl. Madag. 17h: 29. 1956; Molcenke, Ré- sumé 156 & 68. 19595 Ge taylan; Inte cas Suppl. 12: 11). 1959. gts: a —— mal bibliography: & Salisb., Ind. Kew. Suppl. 10: 1h. 19 19h73 Mo Moldenice,, “Résumé él & & 468. 1959. PRIVA LAPPULACEA (L.) Pers. Additional & emended synonymy: Priva echinata A. L. Juss., Ann. Mus. Nat. Hist. Nat. Paris 7: 69. 1806. Verbena tetrandra, calyc. subrotundis erectiusc., sem. echinatis L. L. apud H. E. Rich~ ter, Cod. Bot. Linn. 35. 1935. Blairia Houst. apud H. E. Richter, Cod « Bot. Linn. 35, in syn. 1835. Scorodonia floribus spicatis purpurascentibus pentapetaloidibus Sloane apud H. E. Richter, er, Cod. Bot. Linn, 35, in syn. 1835. Priva lappulacea Pers. ex Benth., Bot. Voy. Sulphur 152. 1846, Priva lapulacea L. ex Patermann, Beitr, Zytol. Nat AE ss & fe te sphalm. 1935. Blairia "Houst. ex Adans." apud G - Kew. Suppl. 13: 1h9, in in syn. 1966. Privia la: lappul ieeea uy ) Pers., in her aa Additional & emended ner aa > Nats, oe: 10, 852. 1759; Adans . o, Fam. Pl. 2: , Cate Pl. Gaba 21h. 1866; A. 6 Syn. Fl. N. 2 (1): 3h. 1878; A. S. aaa, Rep. ay Lg G ard. \: 7. 18933 Hook. f, & Jacks Kew. 2: 638. 1894; Just, Bot. Jahresber. 23 (2 h7, 76, ‘%& 506. ga MU. Kunz, Anatom. Untersuch. ee Stewart, erb. Calif. Acad. Sci., ser. h, 1: 13h. 1911; Guillaumin, Ann. Mus. Colon. Marseille 9: 206. i911; 1967 Moldenke, Notes on Priva 351 Loes., Verh. Bot. Ver. Brand. 53: 80 [Abhandl. 245]. 1912; stad nd nd ‘be Suppl. Ws 33. 1947; Bravo Hollis & Ramirez Cantu, Anal. Inst. Biol. Mex. : 21. 1951; Asprey & Robbins, Ecol. Monog. 23: 411. 1953; Mol-~ denke, ad PY *OLOSAE 5: ¥ & lee 19543 Daring tds Wylie, t on, Wasmann Journ, Biol 1958; Moldenke, te oe a po 38, ho, 425, L7--50, 52, Su, 240, 3h0, al 153, 5, 36 bk, 366, 368-370, 378, 39h, : 1955 Darlingto & Wylie, Chromosome Atl., pr. 2, 323. 1961; Hep- ee R Dalz., Fl. W. Trop. Afr., ed. 2, 2: — 35. 19635 wotenite in Shreve & Wiggins, Veg. & Fl. Son, Des. 2: 1257. 196; Moldenke, es Suppl. 11: 8. 196h; G » Lo 2-363 & 481. 1965; Backer & Bakh., Fl. Suppl. 13: 18 & 149. roc é Java 2: 599, 1965; a: Taylor, Ind. Kew. 1966. Additional illustrations: ng eae Bot. 1: 37, pl. 2h. 176k; Jacq., Select. Stirp. Amer. Hist. Picta pl. 9. 1780; Patermann Beitr. Zytol. Verbenac. pl. 2, fig. 15-1, & pl. 5s As 5. 1935; Alain in Leén & Alain, Fl. cuba Recent collectors describe this P. nial herb, erect or decumbent, to 1m. “tall, with a ng =? small or mimite flowers, the calyx green inflated, partite, and on Herald & Clark 406 and nome i Ph 12162 and J. J. V. Santos 2221, se ourplish” on Abbott 429, *pale~purple" on Woytkowski 5618 Ki 5618 “Tpurple-lavender, white-streaked pet eh lobes" on B. TL. Turner 2100 Hinton 1967, "pale-violet" on Molina R. 263, *Light-violet, 8.5 p av? on Kunsell system per Nickerson color r fan" on Straw & Forman > "lavender" on Correll & Schweinfurth 15651 and Wiggins & wollte 396, "pale-lavender" on I. L. Wiggins 15626 a "pale-mauve sary Purseglove P.6221, "blue" on Breedlo , Breedlove & : bee 13347, C. L. 1 » and Yuncker Sek ee on Plund vein Breedlove 11793, J. Be Hall 2517, Irwin R-1l7, and Morton "pink" on Glassman 1715 and Koelz 34220, sone _ a1 on or ee eat 1/2 "light~blue with th white center" on Wilbur & w=” Wwhite" on Goll 28, lark 339, Hinton 1229, R- Tuck Ol, Wedel 2834, Worth, Morrison, & Horton = Zar) Se vabe’ white, white, petals oe atclet™ “on Cro rosby, 352 PrESTOLvye.s Vol. 1h, no. 6 Hespenheide, & Anderson Anderson 95, Thi southeas south to eastern Peru and Bolivia; introduced in Java and Ghana occasionally cultivated elsewhere. Recent collectors have found it growing in shaded areas, open or flat grazed areas, in loam soil, clay, or silty-clay soil, in alpine meadows, dry arroyos, pedregal » Prosopis brush, fields, barrancas, secondary torral, ie gy dense woods caeeael of woody Mimosaceae, Cereus-type cacti, araliads, etc., 6—10 m. tall, along or near roadsides ate roadsides, and “ bot- toms in s shady canyons . Sapdanine thickets, jacent to ditches, as well as on dry slopes and in coffee of peeps 9 ll from near sea-level to 1600 meters altitude. Schwenke (1931) reports the haploid chromosome number as 6. tional common names reported for the plant are "bur vervain" and "farolito", It has been collected on clay flats, open reek banks ’ ’ ial clay, calcareous clay-loam, and limey soil with rocks, in deep or partial woods, e hedgerows aro fields, swamps in low wet palm a » pastures, varied short brush » and in sand and limestone rocks in ’ Worth, Morrison, & Horton describe the style as "gynobasic", which is a characteristic of the - Herald & Clark re- fer to the @Mi-tt. " and that its hab- Fed loam, partial shade, of old fields and hedgerows" in Nay- ron County, Texas. On the Galapagos Islands it is said to grow in. the Scalesia zone ree in dense jungles of the Up- Per Transition zone with Pis sonia tree: a weed Sonoran to Tropical zones, from Florida and Texas ae the West Indies, Mexico, and Central 1967 Moldenke, Notes on Priva 353 naturalized in Java. Occurring in the Sonoran Desert near Matapé, n Hepper (1963) calls the plant "A perennial weed with spreading and decumbent or erect stems, markedly l-angled; flowers small, the corolla violet with paler stripes." He cites Hall "1891 from Asebu, near Cape coast, in Ghana, and notes "A native of the warmer parts of America and the West Indies; introduced into Java and possibly elsewhere, but not previously recorded from Africa." when the fruit is mature the calyx (with the enclosed off from the persistent pedicel and easily clings to passing ob- jects by the aid of upcurved bristles." The description of this plant by aearl as given by J. A. Murray (1774), is worth repeating here lappulacea. 7. V. tetrandra, calyc. rotundis erectiusculis, sem. echinatis. — Me. wie es be ° eria Loefl. it. 194. n. 69. Semina b In the Linnean Herbarium, under ro 35, rant specimen number "5" is labeled "Lappulacea" in the handwriting of Solander and bears the notation "Br". It is de ly as Priva lappulacea (L.) Pers. Specimen number ber "5", is labeled "lappulacea" in Linnaeus' own handwrit ing and also bears the notation "Br." [=Browme] aod" molander" [Ro- lander collect ed in Surinam after 1754]. It is Salvia occidenta~ iis Sw. in the Lamiaceae eae Keys st noted hes that the C. Junge 2993 a a cited below, bears a label inscribed Sami OF CHILE", but was actually collected in the Dominican b Ma akak rected 4 lappulacea has wor misidentified and dis- tributed in herbaria as P. aspera H.B.K. and as Fe mexicana (L.) Pers. On the other hand, the Tucker 786, distributed as P. lap- pulacea, is actually P. aspera H.P.K.; F. W. Gould 10810 is P. mexicana (L.) Pers.; W. Schumann 1081 is Aloysia gratissima Gilt & & Hook.) Troncoso; and J. Rzedowski 4 19977 and I. L. Wig- gins 18810 are probably mints. nroe Co.: Bl + Additional & emended citations: s.n. [Key West] (W--l3516); Duckett s-n- tia "I, 1930] (W— 1485601); Garber s.n. [Key West, Aug. —T877] (W—L3518)5; H- N- Mol- denke 61) (E—1 100289h, W—1581810) ; Edw. Edw. Palmer 395 (E— 118768, F—266379, W--H3517); Rugel son. (Florida, 5 ahe-I89) (E--11695h, W—512596). TEXAS: Cameron Co.: Correll & Johns n 17948 (Rf )3 c- L. Lundell 12466 (Rf); R. Runyon 26¢ 562 eS sy. Hidalgo Cons Correll & Schweinfurth 15651 (Rf). yextco: Baja California: T. S- Brandegee s.n. [corral Piedra] (Ca—10h890), sn. [San José del 35h PITT GOLOA TA Vol. 1h, no. 6 Cabo] (Ca—10891, W—l0072). Campeche: Goldman 458 eae 3 C. L. Lundell 883 (Ca--186969, E--1017041, F—700L92, 1494515); 3 Ce D. M Mell 2061 (W-~1588308). Chiapas: eee 7670 (Du—508668), 11793 (Rf); Breedlove & Raven 13347 (Ac). Colima: P. Golcsmith 89 (Ca—923500, Du—39897); Koelz 34220 (Mi); Edw. Palmer 1007 (W—209151); Worth, Morrison, & & Horton 8611 Sere 643505). Guerrero: R. Q. Abbott eg (Ip); B arkley, Webster, Paxson 17M739 (Au--123212, Au--209670); Herald & Clark 339 har: 406 (ii); Hinton 10545 (Rf), 10573 (Rf), 14391 (Rf); Lyonnet 183 (W--103)282); Edw. Palmer 551 (Ca~167893, E—118781, W——267026) 3 Paray 3039 (Ip); Rose, Painter, & Rose 9h19 (W--452907); Ryan & Floyed 79 (Mi); Willis & Crism Grisman | 239 (Mi). Hidalgo: Lundell & Lundell 12382 (Rf). Jalisco: Wilbur & Wilbur 1569 (Mi). wéxico: Hinton 4360 (Rf); Ortonburger, Paxson, & Barkley 16M816 (Ca— 763400, N). Michoac4n: Hinton 1222; (Mi), 13062 (Mi, Rf, Ur), 139k) (RE, Ur), 13970 (Rf), 13975 (Rf), 16120 (Ca—97622, Mi, N)5 B. L. Turner 2100 (1). Nayarit: Feddena a 529 (Mi); Ferris 5613 (#1269091). Oaxaca: R. M. King 7 (Ii), 1229 (W-~2301501), 1751 (Au--183)86, Mi, N, geet whee 1993 (Au, Mi, W—2301527 15 Martinez—Calderén 22 227 (Rf). tana R GF. dasaadie 1702 (B, E—952970, F—58500, 0, W—1267h1h), 1702 vas (D--6583L,7). San Luis Potosf: J. Rzedowski 10366 (Ip). Sinaloa: T. S. Brandegee s.n. (Culiacan] ~(Ca——LOL89 » W--873701); Gonzalez Ortega 1220 . 4220 (W--1083506), 6429 (D~651101), 6758 (F—709231)5. Mexia 3% 1, 1/2 (ca—367533, E— E-~-966856); Nervaez Montes & Salazar 389 (W—1035275) ; Edw. Palmer 1458 (Ca--770289, F—707669), X, in part (W—305280); J. N. Rose 1722 (W—300588), 32h0 (W--302205); Rose, Standley, & Russell 13371 TL (W-636195), 13711 711 (W--6 36510) 5 Je Rzedowsid 1 165L7 (Ip); Straw & Forman 1838 (Qi). Sonora: Wig- gins & & Rollins 396 396 (Mi); Edw. Palmer mer 75 (W- (W—57692). Tabasco: Rovirosa 531 (ca—251126, W— W—1323167). Tamaulipas: Graham & Johnston L681h (Au—17,3k5), 4683a (Au—171698, Mi); “Edw. Palmer 8 (E--777h75, W--L629))) , 502 (i (W—572731). Vera Cru Key, Rowell, & Webster 260 (Au=—12 tg Mohr s.n. (W—-771822 3 Or at a0 to : rong F—-279058); Purpus 2099 (Ca--13973)5 5 dete antos 2221 (Mi). Yucat4én: G. F. Gaumer 364, in part (E--118779, F—36167, W—268669) , 39h, in | n part G—Ki57), 24029 (F--552031, W—1268127), s.n. [Izamal, 1888] (F—181636), s.n. [Buena Vista] eter. Gaumer & sons 1702 (Ca—l,6012); C. F. Mill cee i248 (F-61518); Schott 22 (F—-l,0020), 23 (F—h0021); Seler & Se WSkh) . UATEMALA ier 3995 (F— Alta Ve: 2215787) ; Ee po 211 (W--1081323). El Petén: H. H. Bartlett 12162 (Du-—353971, F. 971, F—71005) ; Contreras 932 (Ld); C C. Le Lundell} 3693 (?--688127, F—689169), 15343 (Ld); J. A. Steyermark 16267 1967 Moldenke, Notes on Priva 355 (Mi, Rf). Escuintla: Rojas - (W--1166665). Izabal: Greenman & Greenman 5978 (E—-908315); P. C. Standley 23762 (W--1139L37), 2,635 (W--1150256), 2795 ee Santa Rosa: Heyde & Lux 3017 (W--57691, W--1323161). Solold&: Shannon 27 (W-13507, W— 1323166). BRITISH HONDURAS: Gentle 17h (F —713008); Karling 42 (F-59984, W—1h06569); C. L. Lundell 79 (F—597989), 363 (F— 712069); W. C. Meyer 139 (F—-713297)3 Schipp 845 (Ca—l81631, E— 1020212, F—71231). HONDURAS: Atl4ntida: Severen 25 (W--1168299); P.C. Standley 5369 (W--107865). Cortés: Thieme TS [J. De Smith 5613] (W—1323165). La Union: Morrison & Beetle 8770 (Ca— 643529). Moraz4n: Glassman 1715 (N, Ok, Ok, Ur); Molina R. 263 (Ca-- 792806). EL SALVADOR: Ahuachap4n: P. C. Standley 19719 (#— 1135644). La Libertad: P. C. Standley 23339 (W—-1139028). Mora~ z4n: Tucker 501 (Ca—1000902, Mi, N, Rf, Vi). San Miguel: P. C. Standley 2111], (Ww—1136927). San Salvador: Calderén 917 (Ww 1151880); Renson 16 (W—576119); Velasco 885 5h (W--829126, W— 1323156). Sonsonate: P. C. Standley 21905 (W ; (W-—-1137660), 22003 (W--1137758), 22097 (W—1137646). NICARAGUA: ene ‘Cake Baker 754 (W-862823)5 Maxon, Harvey, & Valentine 71 — 1160820). Grenada: C. F. Baker 166 (Ca—199lb, B—008110), 619 ered Maxon, eee & Valentine 7601 pers . Ma- René 66 (W—1209530). COSTA RICA: Alajuela: A. R. Moldenke 120), (Rf). Guanacaste: A. R. Moldenke amas (Ac). Limén: United Fruit Co. 71 (W—861477). ~~puntarenas: ike Echeverria 18) (Ca-— 7762uL); A. R. Moldenke 1219 ee wal (Ac). San José: A. Re Moldenke 1200 (Rf Rf). Pr Province undet ermined: Rowlee & Rowlee 505 (W=1036 1036486 86). PANAMA: Bocas a Toro: Carleton ie ape C. P. Cooper 129 (F—579557)3 Durl ae 293 (W--1205126 Wedel i 283 (Mi). Canal Zone: Duke . (Sept. 26, aseal Gs Be P. C. Sti Standley 2673 (W--1217023), 27157 oat Bi (rae. . Ep 20 12003 aang eh P. C. Standl 16 (W—12183 Ds Re a 1032 (W--678365). PEARL SoS. ame 278 (W— 165); H. Pittier 3560 *m678616) 3 Ps P.-C. Standley 27032 27031 “s Tames 27917 (W—-1127 RAHAMA ISLANDS: Andros: Small & Carter 8928 (W—758229) . Rleuthera: Britton & willspaugh 5534 (3 A A. S. Hitchcock s.n. [11-18-90] gig oie t Green Turtle: Brace 1482 (N). New Providence: Brace 2 3 (F—248455), 95 (4)> 483 (Kk); Britton & Brace 534 (N), 655 — ; Curtis 2h (E—- » W--L.2852); Earle 12 (N)j As Be = ant Ue ho (F—225221, N). Parro Millspaugh & Millspaugh 9196 ( Watling: Britton & 6166 (N). Island pant Oe as ve $3 Hitchcock $ +n gen. [12-3-90] a 356 PHYTOLOGIA Vol. 14, no. 6 (E--6910). CUBA: Havana: Leén s.n. (15-5-1909] (Vi), s.n. (Vi- 4027); Van Hermann 113 (E--969555, F--170529). Las Villas: Combs 220 (E--118796, F-35802); Pringle 9 (W--28)2h) . Meare Eggers 527h (W—1323162). Oriente: Alain 1992 (Z); Hioram Btiste 1323 (Um--100); C. F. Millspaugh 1043 (F--610h3), is) ioks (F— 61048); Morton & Alain 9013 (W--2285127); sh Shafer 1576 (Bi, W-- 659811). Pinar del Rfo: Shafer 111), (W--699300). CAYMAN ISLANDS: Grand Cayman: A. S. Hitchcock s.n. [Jan. 1890] (E--118795); C.F. Millspaugh 1286 (F-61286). JAMAICA: R. C. Alexander s.n. [Rio Cobre] (F-50988), Sen. (W--10,8225); $ S. , Brown 306 own 306 (D—53892h); Crosby, Hespenheide, & Anderson 95 (MMi); Fredholm 3081 (W--315932); W. Harris 11796 (E—-79259h, W—790858); A. S. . Ss Hitchcock s.n. [12- 9="90) (E—-11879); H. A. Lang 61 (D-=-SSh717), 62 (D—5' eee —_— ee _ Lloyd 100) (E—-118793, F—228293); Maxon 1657 (W--L27952), 2180 acteao ete z Killip 313 (W--1046009), 1605 (W—-10)6615) 5 xi 1 (F—60921), 1910 (F—87523); Orcutt 19b3 dadscn a Cet (Ca—l,30600) ; F. W. Pennell 11225 (D—645)h2) 5 Sauer 1865 (Ws); Yuncker 17028 (Mi, S). HISPANIOLA: Dominican Republic: Abbott 1198 (W—1078620), 646 (W--1078619), 648 (W— 1078621), 1026 (W——1078783), 1200 (W--1078865); B. Augusto 10L5 (N)5 Faris 303 (W--11)5872), 32h (W—11)5889); Fuertes 886 (W— 698029); C. Junge 2992 (W—22537h2); Raunkiaer 369 (W—1110063) 5 Valeur )))3 (W--1273793) ; 3 Wright, Parry, & oe eee el 354 (W—-h3512, W—l3513). Haftd: Ekman H.7958 (W—-1112888) 3 B.C. Leonard 293 (W—1075189), 3101 (W--1075358), 3302 cacao, W—1075596) 512h (W—1077716), $229 (W--1077836), 7506a (W--1149369); Leonard & Leonard 1389 (W—1152822), 15413 (W—153679). PUERTO RIGO: Goll 68 (W—l08661), 150 (W—l08737), 675 (W—l09218); A. A. Hel- ler 6181 (D--500557, E—-118286, W-1,26]17) ; Heller & Heller r 1L9 (W--I12569h) ; Johnston & Stevenson 1297 (W--1h759h3)3 C. F. Mill- spaugh 337 (F--60337); : Sintenis intenis 157 7 (W--1323151), 185 1857 7 (i 1323158); | Underwood & Griggs 20 201a (W—l0593)) , 529 Y (#05472) 5 Velez 2918 (N). VIEQUES: Shafer 21,25 (W--759977) « VIRGIN IS- LANDS: St. Croix: A. E. Ricksecker 167 (Ca--73250, E--118792, W--278058); L. A. Ricksecker 131 (E—118791, W--42529); Thompson 8h2 (D--619098), 866 (D—619082). St. John: Britton & Shafer 507 (W—756L417). St. Thomas: Eggers 26 (W--1110232), 365 a Ten ts s.n. [Novbr. 1882] (W--1323153), s.n. [Dec. 1886] (F— 141757) Ehrenberg 107 107 (E--114062); Krebs s.n, (F—131515). LEE WARD saceee came Antigua: Box 1021 (N). Guadeloupe: Duchassai n. (E—188790); Questel , L566 (W. (W—2),5366). Montserrat: rae 138 (W—695158). WINDWARD ISLANDS: Barbados: Bovell & Freeman S+ Ss Re (Bot. Stat. Herb. 139] (F—-200798, W—81553h); G. S. Miller 57 ————— —— 1967 Moldenke, Notes on Priva 357 (W—11)7613). Grenada: W. E. Broadway s.n. [Oct. 26, 190k] (E-- 118788, F--175765, W—-l29675), sen. [November 190k] (F—175766). Martinique: A. Duss 196la baong lie Hahn 962 (Ca—332h86); Ko~ haut s.n, [Sieber Fl. Mart. 316, in part] (E—118789). St. Vin- cent: Morton 826 (W—-1883987), sla (W--188),351). TOBAGO: W. E. Se L6h5 (W--759676) ; Eggers 5832 (W~1323159); Purseglove P6221 (W--2)29305). TRINIDAD: DAD: Britton & Hazen 162 (W—-101683h); I a a meead Archer 1])7 (W-—154,2191) ; Toro 328 (W--1342845)- Atl4ntico: Du~ gand 5428 (W—-2369330); Elias das 556 (W--1422489), 872 (W--15h,3262) , 1035 (F—700259); Paul 871 (F--694282), CoL2 (W—1281942); Fe We Pennell 12010 (D--61259h). Bolivar: Heriberto 16 ye sk Ts Killip & Smith 1043 (F-638258, W—13500L2); F. W. Pennell (W-~101,3036). Cundinamarca: Mutis 3689 (W--1562717); Me Sc! stere = 1006 (S). Magdalena: C. Allen 241 (101278), 461 (E— 101,158); H. H. Smith 545 Ca SO5 8, E—-118799, Hi--19731b, Mi, ¥--$33011). > 1b65 (D—-509362, E—~118798, W—53355L). Tolima: Hol- ton 496 (D--510664). Valle del Cauca: Holton 95 (D--61066)) 5 Walker 1265 (E--9280l7) . tar Bolivar: J. A. Steyermark 88186 (N). Carabobo: Asplund 15133 (S). Delta @ Amacuro: Rusby & Squires 306 (Ca--52343, E--119797, W--325638). Laré: Saer (F—6898),2). Sucre: W. E. Broadway 67 (W--1188701) . Trujillo: 2560 (Ve—28166) ; Moc- querys 909 (B), s.n. [Zulia] (N). BRITISH GUIANA: A. Se Hiteh- cock 1653 (W=—-1055889) , 16773 (W—-1056033); Irwin Roll? (Au— 165429, Au--178036); Persaud 402 (F—-532958). SURINAM: Lanjouw & rindenan ai (N); Rare Sas Sone [May 5, 1905] (D—599922). a GUIANA: W. E. Broadway 89 (W TW—-1068)63). ECUADOR: Guayas: As: plund snare (S); Gilmartin 690 (N, W--2428422). Los Rios: As- plund 5596 Naps ns 30558). Manabi: Anthony & Tate 118 (W— 1192206). GALAPAGOS I 3312 (BA 817128, W—921595). Staats Schimpff 151 (B—1072193, Ge— 2129). Indefatigable: Itow 139 (Du--56L525), 1398 (Du--5645214) 5 I. L. Wiggins 18385 (Ac), 16626 (Rf). PERU: Amazonas: 5617 (W—-23969kL), 5618 (W—23969K5) « co: Asplund 12096 (#— 222450h). Juntn: Macbride 5296 aha gggtn pane els “Loreto: Klug zs SC serale W——-1562h9); Ll. Ww 1 (F—-603732, a , 418 (F a, pS Saree 23s (7623688), 4158 (F—622962, W—-1L96401), Lh28 (F--624 741) - : Allard 21510 Seaseacayet . Tumbes: Weberbauer 7737 (F-S7186h) ‘ Department und : Rufz 4785 (W—2h9769) = BRAZIL: Amazonas: Chagas §-De ae Inst. Nac. Pesq. Amaz- 2033] (Bs); Herb. Inst. Nac. was Fes. aq. Ama. 939 [2] (Bs)- NEW SPECIES OF PASPALUM Jason R. Swallen Ceresia Ma a Beebe s fay Paaeee sp. nov. Culmi » 50-60 cm. alti; laminae erectae, appressae, 6-8 at longae, 2-3 mm. latae, prominente papilloso-hispidae; spiculae h.2 mm. longae; gluma secunda longipilosa; lemma sterile e marginibus ciliatun; fructus 3.l. mm. ‘longus, pallidus, sumo ciliatus. Perennial; culms stiffly erect, 50-60 cm. tall; cacebie ia. conspicuously Bpbi ee SSaneexm ena: sheaths all longer than the internodes, ut 15 completely ee the culm, Trebeoliat etkdes erect, appressed, 6-8 cm. long, 2-3 mm. wide, acuminate, prominently papillose-hispid; racemes 1 or 2, when 2, one a short distance below the other, 1 mm. wide with narrow margins; spikelets 1..2 mm. long, t cond pilose with long hairs on the lower part of the back, a few hairs exten n the margins to th i of se back, the margins ciliate on the upper half; fruit 3.) mn. Loris, pale, the lemma rounded, ciliate at the a. n the U. S. National Herbarium, No. 1612651, collected axtong rocks at top of rocky hill, Carolina to San on. lso c Leaaee approximately in the same locality (Swallen 077, and from Grajant to Porto Franco, Maranh&o {Sian 3796). Paspalum spissum is related to P. carinatum Humb. & ene but differs greatly in having flat appressed blades completely covering the culn, conspicuously villous cataphylls, and very narrow margins on the racemes, - carinatum the blades are involute, sheaths are scarcely pilose, and the margins of the racemes are broad. There is little difference in the Boece although those of P., carinatum average slightly larger. Notata 2. PASPALUM NITIDUM Swallen, bake nov. Culmi caespitosi, ae m. alti; Laminae l. witha cm. 1ongae, 2-3 mm, 1atvae, pliosae; racem1 ¢, sud conjugati, rnachi glabri; spiculae 1.5 mm. Longae, 358 Bh 967 Swallen, New species of Paspalum 359 zlab ghina ellipticae vel obovatae; fructus pallidus oes et lemnate sterile brevior : : ong, pilose on both surfaces; racemes 2, su conjugate, 1-2 spice cts rae rachis glabrous, even on the margins; aliehtis oF 5 mm. long, smooth, shining, elliptic, or mead ovate, the tip rounded or sometimes minutely + ; fruit pale, s Baan shorter than the second T poaeeated on campo at Monte Cri ree Garuv oer ee do Sul, Santa Catarina, Be ie ghee 3s leneei ks Be Reitz & R. Klein (No. 10900). Also col- Ginn aay eitz & Klein at Barra do Sul, Araquari = meee species is the small member of the Notata ie up, perhans more closely ag oP. ee kes, than anything else. It differs, however, ing smaller spikelets and densely pilose cet ea Livida Plants annua Racenes h- as 2-2.5 em. long, the rachis 0,5 mm. wide, gla abrous, the margins scabrous; spikelets l. 6- 1.7 mm. long, the tip rounded. Sure Racemes 5- Fe us a more than 10, =i ea 0.8 mm. wide, pubes cent or seabrous on the surface, with long hairs on the margins; spikelets 1.9-2 mm. long, very Pe apiculate. . eranis Pisnte pasa yk. Ee trims Lowermost sheaths densely pilose pene the base. Culms very slender; racemes 3-10, plage ire 1-2 m, long, on 4 precaoea axis; axils of the racemes glabrous; spikelets l. 8-2 sts long, apiculate; blades subfiliforn. 5. trichophyliwi Culms stouter; racemes mostly 10-20, ascending to spre agi: ene lower up to cm. long, long- pylas ils; epee 2-2,.2 ma, long, pions d at gk Civ. scexe P. goeldii lowermost sheaths glabrous or nearly so toward the -2.5 mm. long; culms dec umbent-spread- mite rooting at the anwar nodes; racemes relatively distant, spreadi the rachis 1.5- 2 mu. wide, usually wipes 2s ha. irs Jenticulatum Spikelets 263 ing, sides 360 PHYTOLOGIA Vol. 1h, no. 6 Spik sprter 1.4 mm. long; culms erect or geniculate lower nodes; racemes crowded, ab pRaKaeA, cls rachis 0.5 mm. wide, long- -ciliate 8 yee humigenum 3s ee ae PISINNUM Swallen, sp. ulmi erecti, 15-27 cm. alti, glabri; lenin ae ual; culms cers or cer 15-27 cm. tall, more or less geniculate at the node glabrous; sheaths shorter than the internodes, ah ctakenis , the uppermost glabrous; ligule 3 mm. long, whitish, somewhat decur- rent; blades flat, lax, 5-10 cm. long, 2-3 mm. wide, acuminate, scabrous-roughened on both surfaces, e midnerves and margins scabrous, with a few long hairs near the base; inflorescence 3-l, cm. long, composed of l-7 appressed or narrowly ascending racemes 2-2.5 cil. long, the rachis 0.5 mm. wide, glabrous, the margins scabrous; spikelets solitary, pale, 1.6-1.7 mn. long, broad and rounded at the summit, slightly obovate; second glume and steril green nerves contrasting with the pale body, the lateral nerves conspicuously scabrous; fruit pale, minutely peters : n the U. S. National eager tere — enige oats collected ra wet open ground near amp, Faz Nacional to Picos, Piaui, yrs April l-5, “Iga, by Jason R. Swallen (No. 217 lh. PASPALUM Peet Swallen, sp. Ts , Culmi 90 ¢ lti, erecti, ramosi, glabri; vaginae sparse Lonsisi ideas laminae planae, 17-28 c ngae, 3-4 mm. latae, acuminatae, sparse Longipilosae; pacentt 10-12, appressi, usque 3.5 cm. longi; spiculae 1. mm. longae, obovatae, apiculatae, marginibus scabrosae; fructus pallidus late acutus. : ual; culms about 90 cm. tall, erect, branching at the lower nodes, glabrous, or the nodes sparsely pilose; sheaths elongate, densely scabrous-roughened po sparsely pilose with long hairs; ligule 2.5-3 mn. Pa Rare ears decurrent; blades flat, 17-28 cm. 3- mm de, acuminate, densely scabrous— ct are Take t the sheaths and sparsely pilose with long hairs; inflorescences about 10 em. long, compos? 1967 Swallen, New species of Paspalum 361 sterile lemma scabrous, 3-nerved, the marginal nerves conspicuously scabrous; fruit pale, broadly acute. in Type the U. S. National Herbariun, abe 1613382, collected in Tyee pond, Crathefs, Cear zil, May aes 1934, by Jason R. Swallen (No. "1507) ° British Guiana: Rupununni Savanna, Melville (Her- onde No. 162). Brazil: Cear&a: Junco to Quixada, Swallen 4163; Cratheis, Swallen 4.510, Piaui: Picos, Swallen 1.225; Fazenda Nacional, Swallen 201. Maranhé&o: Caxias to Barra do Corda, Swallen 352, Bahia: Joazeiro, Dorsett & Popenoe l,06b. Se fe hae feta pads es Henr. Blumea ia fe 1941. il: lha Marajé, Kuhlman 2135; Black & ingelhard d 50- bo8h, ie -8985; Soure, Black “SO. S83 Black & Lobato 50-9301; Fazenda Tuiuid, Rio Arari, Black, Ledoux & Stegemann 52-1255; eeu Camburupy, Swallen vz Fazenda cayinhd, Goeldi 165 (type). Ilha Mexicana: ta Maria, Huber 6. PASPALUN GOHLDII Swallen, sp. no C erecti vel decumbentes, be om. alti, glabri; laminae ca. 25 cm. longae, 3-l. mm, latae, erectae vel patentes, glabrae; racemi 20, ede eaent ow usque 5 cm. lo oa epicutee 2-2.2 mm. longae, Par hacer dense ing racer rmost 5 em. long; 6 edo aked une abe 1 as wide, pubescent, the mar- Sins scabrous and with a few long, pousteven Spikelets 2-2.2 mm. long, Ng ge more © pai obovate, apiculate; second glum d sterile s strongly densely pubescent, 3-nerved, the marginal one Scabrous; fruit pale, striate e in che, 3. National Herbariun, eas hae collected on open savannas, Estate race eT ee Island, Perf, Brazil, June 1918, by André 97) ens examined: sone uchete Gavinho, Goeldl 7109, 180, nabs, Blac 192, 193, 200, 250; Fazenda Menino Deus, ‘ Rgelhard 50-9028, 50-9038, 50-8962 Soure: Ble ck & Camargo 8873, 8876; Monte Alegre, Swa Sobral, Avelino & Black k 118ce. 26. 7. PASPALU DENPICULATUM Trin. Gram. Pan. 111. 18 362 PHYTOLOGIA Vol. 14, no. 6 8. PASPALU Ra ER Swallen, sp. ulmi erecti, 20-2 cm, alti, nets lamin : pl i.i2 lonca >» 2-l. ma, latae, glabrae, eri ana. em, ibus scabrae; racemi 1-8, appressi vel adscendentes, usque 2 cm. longi; rhachis prominente ciliata pilis 7 mm. longis; spiculae 1.3-1.]: mn. longae, ellipticae vel Shegeitetning apiculatae, glabr ae. nmnial; culms erect or geniculate at the lower nodes, 20- aes em, tall, sometimes branching at the lower nodes, glabrous; sheaths all much shorter than the internodes, compressed, keeled, glabrous, the some 12 cm. long, 2-1 ma. wide, glabrous on both writ he margins Sie eeueS inflorescence 25 cr. cm. Monies? rachis 0.5 mn. wide e, pilose in the ae glabrous, the margins scabrous, jliote with proms iets white hairs about 7 mm. long; sail oie 1. long, elliptic-obovate, purple-tinged, sc ebelab: 3-nerved, the merginal nerves very scabrous; frui pale, tegeng? striate Type in the U. S. National Herbariun, No. oases. collected ap Sth Re inside zone of TIPE: moe sedges & Rio Salitre, 46 lm. west- fsa lak of Joa Bahia azil, Deccaeer 15, 1921, by Agnes ting (iio. "7931). Recta 9. we rece PETROSUL hp rncbed sp. no os Culmi caespitosi, erecti, 70- Bc em. alti, gilebris vaginae eek pieent oes laminae 35 cia - longae, 1n- volutae, basi papilloso-hispidae; raceri 5-8, adscen- dentes, inferiores l-5 em. longi; spiculae solitariae vel binae, 1.3- 1.9 mm. Merigaes glabree, pedicellis she Mote 0.5 mn. lon Perennial; culms tu tea, erect, 70-85 cm, tall, glabrous; sheaths ocuabaee at t i base, beconin, t 8 t e in the upper half, the inner surface pubescent; inflor- n, : em. long, composed of 5-8 ascending racemes, the lower li-5 em. long, the rachis O.l4-0.5 mm. pio: gletrouss adilen let solitary or sometimes paired, 1.0~ oes : long; second e and sterile lerma elliptic, »vroadly acute, 5-nerved, the midmerve much stronger t lateral, glabrous; fruit as long as the spikelet, tel : shining. the U. S. National Herbarium, No. 1910825, edge of mountainside, Cerro Camejen, 1967 Swallen, New species of Paspalum 363 about oy 72°u, Faun Gay Colombia, July 2h, 193, by Carl assl (No. o1s). aopaveatis aaee species belongs to the Recta aoe but the racemes are paniculate rather than solitary o in pairs. Quadrifaria 10, PASPALUM CHAPADENSE Swallen, sp. nov Culmi erecti, 90 em. alti, nodis pilosis; laminae erectae, 30-32 om. longae, 12 mma, latae, marginibus papilloso-ciliatis; racemi 28, adscendentes, 5-6 cm. longi; spiculae densae, binae, l. -O mm. longae, 0.8 mm. iatae, acutae, dense pibescentass gluma secunda sp ae pees or. nnial; culms erect, 90 cm. tall, the nodes apr teasediplikycnt onal sheaths all loncer than the glabrous, ciliate toward the summit, with long, the rachis 0.3-0.l. mm. wide, densely hairy in the axils; spikelets dense, paired, 1.6-1.8 mm. long, 0. mi. wide, acutish, elliptic- foie baie densely — the pedicels less than 0.5 mi. long; secon nd glum shorter than the spikelet exposing the light eas Striate fruit. Type in the U. S. National Herbarium, No. 2238394, collected in chapada between Caxias and ra do Corda, Maranhio, Brazil, February 16-26, Swallen (No. 3508). This species h northeastern Brazi as pense es Brejo wis to Séo José), Swallen 37h; ¢ Bae Se 1: Corda, parry 3519, 3562, 3503; 3565, * 3506, 360 B ae do Corda to Grajahi, Swallen 3708; Ger elanes a h an Anton te de Balsas, suallen agit Cear&: Sw icke bo52. __rernaubuco? cept Lise eee mr species, atl nur ane 4-28, a there are no other ete anes to 11. PASPALUM DASYWYRICHIUM Dusen, Sp. NOV. Culmi 11.0 em. alti, internodiis inferioribus i a, 1 cm. abbre a resso-villosi; ligula c Pcuthad aecae Lh ma As latae, ee . viculae 2 mm adscendentes, usque 9 cm. longi; spicu ewes 0.8 mm, latae, ellipticae; gluma secunda “frecta vi decumbent base, Perennial; culms hard, erect from @ ir 10 cm. “ball, the lower internodes ts ee accompanying leaves falling early, tees 36h, PHYTOLOGIA Vol. 1h, no. 6 the culm densely villous, the upper internodes glabrous, glaucous, the nodes appressed villous; sheaths densely ascending papillose- -villous toward the base becoming le d wide, firm, flat, appressed, papillose-ciliate, the hairs about 5 mm. long; inflorescence 25 cm. long, composed of 25 or more ascending racemes, the ah one as much as 9 cm. long, becoming shorter toward t summit, bearing short appressed branchlets, supeoteale toward the base; spikelets 2 mi. long 8 mm. wide, brown, elliptic; second glun in, nerveless, 1/2-3/4 s long as the fruit, often as lo ruit Fact nerved, the sterile lemma as long as the fruit, nerved; fruit light-colored, the same shape and size as the spike elet. in the U. S. National Herbarium, No. aan deliected among rocks, Villa Velha, Parana, Bra October 21, 191), by P, Dusen (io. 15700). This “qpoures is credite 4% to Dusen because the name is written on the label as a new lees mie it was never published. Also collected in Sao at Caietras by Kuhlman (No. 36620), and at Beitadetaes Tobias by Hoehne & Gehrt (No. 36726). Linearia nOVe : i jae i, gracillimi, 25-35 cm. alti; laminae 2.3-2.7 mm. longae, pallidae, ceateelits 1 ma. longis scabris; gluma secunda et lem: rile tenui nervia, glabra; gluma secunda fructu 1/5-1/l, brevior. mnial; culms erect, very slender, solitar Pe 2 together in small tough tufts, 25-35 em, tall, with one culm node, glabrous; sheaths glabrous, or occasion ally vein a few long hairs, the culm sheath bladeless; mm. long, decurrent; bla de s 5-15 om. pag pee scattered hairs; raceme 1, erect, “2=3 om. long; rasinde solitary or paired, somewhat inflated, 2.3- 2.7 mm. aoe pale, on round scabrous pedicels about 1 mm ms; second glume and sterile lemma thin, appar~ ently Jenerved, glabrous, the glume about 3/l-l;/5 as long a ma pale, pointed, minutely striate fruit re . ope Minas A190 co! Brazil, by es Chase (No. 11259). so collected in "Terreno brejoso beira correg° " Gampo Alegre, Sdo Paulo, Brazil, by Toledo & pp loes (Dept. Bot. S80 Paulo 3200). 1967 Swallen, New species of Paspalum 365 13. PASPALUM CRISPULUM Swallen, sp. nov. Culmi gracillimi,. dense caespitosi, 23-33 cm. alti; laminae usque 10 cm. longae, filiformes, sparse papilloso-pilosae; racemi 2, conjugati, adscendentes; spiculae 1,.9-2 mm. longae, 0.8 mm. latae, solitariae, elliptico-lanceolatae, glabrae; fructus albus, apicu- latus, minute striatus. t of the culm long-pilose; spikelets 1.9-2 mm. long, re Type in the U. S. National Herbarium, No. 2206290, collected on campo at Macido, Niquelandia, Goias Brazil, by A. Macedo (No. i i Lr, * us. ellipticae, glabrae; fructus albus, ute riggs mnial; culms erect, soft, compressec, : ere : 115 cm. tall, glabrous; sheaths overlapping om . lower part of the culm, shorter than th urface, in long, the rachis 0.5-0.6 mm. wide, el wide, narrowly margins; spikelets 3. tim. iO Lm lon : elliptic, on glabrous pedicels sg Se Senerved gila- Slume and sterile lemma equal, faint 7 punctate- brous, light green; fruit white, minutely r bane ee the U. S. National Herbarium, No. 1982916, collected in lake, at Santa teres teet Macedo (No. Gerais, Brazil, February 18, 3790s Toe the same time 2167). A second collection was mace end place (Macedo 216l)). 366 PHYTOLOGIA Vol. 14, no. 6 15. PASPALUM FESSUIi Swelien, sp. nov ; Culmi graciles, erecti, 50 cm. alti, glabri; aninae graciles, involutae, 1-19 em. longae, supra ae, 0.0 mn. latae, pallidae, pedicellis 0.5 m. long gis, erelee) glume secunda et lemma sterile pallida, tenuia, 5-nervia,— glabra; fructus albus, obtusus, maeeinibas parallelis. Perennial; culms slender, = from an ascending epee emt blades slender, inrolled, eng cm. long, glabrous on the lower surface, densely and minutely pubescent on the upper; racemes 2, conjugate, 1 greenish-white, round at the tip, the sides nearly parallel, ninutely striate Tyy the U, S. National Herbarium, No. anrGeh?, collected in sandy — soil on _— bank of the R Paranaiba, Praifio, Ituiutaba, Minas Gerais, meagitge September 3, 1950, by re Macedo (No. 253). 16, PASPALUM AMBUS?UM Swallen, sp. nov. Culmi dense caespitosi, erecti, 75-100 em. alti; laminae 30-50 cm. longae ad basim longiattenuatae, supra po a et molliter pilosae, marginibus scabrosis; racemi 8 v » 4-5 om. longi, appressi; spiculae binae, 3. 3 -6 mm. longae, 1.3 mm. latae, on gatas ial; as de vey:: tufted, erect, 75-100 cas tall; weeattd all longer than the internodes, the uoper- most part enclosing the bas the panicle; basal blades 30-50 em. long, long-attenuate to b densely and softly pilose above, glabrous beneath, the margins very scabrous, the r blades much wide, very scabrous on the margins; spikelets in pairs, -h=3.0 10m, long, 1.8 ma. wide, elliptic, bluntly acute, glab geet fruit pale, flattened, pani} Lose eugene. e the U, Ss. ae a Herbarium, No. 1961593; collected sec on pare ado, Hsta Sar a: eee Poni va spay wing Specimens probably can be referred to this species: Swallen 8129, 8791. faspelun ambustum appears to be between the 1967 Swallen, New species of Paspalum 367 Linearia and Vir gata groups. It has the habit of other species of aad Linearia group but the blades are v scab “A bbste§ bey on th : mar gins and the spikelets are in pairs, acters of Ke Virgata grou Eriantha Blades narrow, involute Spikelets ce mm. long; “racemes erect, appressed. ve P. album 5 hae he pe 3.6-3.3 mm. long; racemes ascending. Spikelets long-pilose or papillose-pilose, the heirs not obscuring the goer ah 1 Spil Ps gianent ines pikelets densely covered with long " si licy hair securing the spikelet..... 19. F. invo tian obsc Blades flat, broader pe le densely and softly pilose.. 20. PF. mollifolium ees glabrous or more or less hispid. Spil relets h-lh.2 mm, long. Blades glabrous, narrower at the base than the mouth of the sheath. 1. P. rigens Blades conspicuously spreading-oi ilose or hispid, narrowed but wider than the mouth of the 3 Cath. .secescesecees 22. P. intonsum Spikelets not more than 3.6-3. 7 om long. ades 8-10 mm. wide, evenly distributed. 23. P. naughtii ee eed: more than 5 mm. long, usually less, ostly crowded to the base of the plant. Spikelets 3mm. long; blades a ee P. sericatum Spikelets 3.60-3.7 mm. long; bla des hirsute. Culms loosely tufted, , Palnonatouss plades 3 -&8 ca. long. 2 . paucifolium Culms densely curted, not rhizomatous; blades 8-1) om. long....-++- 20+ EB formosulum Culmi sanesed, 90 cm. alti, eave nodis ogee laminae involutae, illa innovationun 12-17 cm. longae a fructus 4 . longus, acutus, pallidus, striatus. mni culms erect, 90 Ciil. tall, ous, the nodes bearded with ascending silky hairs; low sheaths appressed-pilose, the pi rie labrous, nearly bladeless; ligule >ro panaceous, ie long; blades loosely involute, those of mgr eros ogpiee 12-17 cm. long, pilose, those of the 45 em. long, eiabro ous or nearly so 368 PHYTOLOGIA Vol. 1h, no. 6 long, composed of 3-6 narrowly ascending racemes, the lower 7 cm. long; rachis 1 mm. wide, scabrous on ‘ ; mm. glume and sterile lemaa acuminate, densely villous, Senerved, the lateral abbas marginal; fruit m. long, vaca pale, striate in the-0. 8; +Hattoont Herbarium, No. 150020, setivctod in sand and gravel on campo, Diamantina, Serra de San Antonio, Minas Gerais, Brazil, altitude 1200-1300 meters, by Agnes Chase (No. 10397). 18, PASPALUM DIAMANTINUM Swallen, sp. nov. Culmi erecti, 65-75 cm. alti, compressi, glabri; laminae 17-26 cm. longae, plerumque plicatae, 1 m. O. mn. lata marginibus scabra; spiculae 3.6-3.5 mm. longae; gluma secunda et lemma sterile S-nervia, villosa pilis is Pere nnial; culms erect, 65-75 cm. ee lake Sait ; hs H + 3 pressed, keeled, mostly crowded toward ths base, the lowermost densely pubescent or pilose, the upper elon- gate, glabrous, nearly bladeless; ligule a membrana ceous rim 0.5 mm. long; blades 17-26 cm. long, usually folded, 1 mm. wide as folded, ascending, pubescent or pilose on the upper surface, or glabrous toward the tip, glabrous beneath; inflorescence composed of 3-5 approximate, ascending racemes 7-9 cm. long, the rachis 0.6 mi. wide, scabrous on ae margins; spikelets pha ares mm. long, the second glume and sterile lemma 5-nerved, the nerves almost Bote: are villous or papillose-villous with white, spreading hairs about 1 mm, long; fruit as none as the spikelet, greenish, oblong, Beery. acute in the U. 3. National Herbarium, collected on white at ween gravel, Diamantina, Serra de San Antonio, Gerais, Brazil, December 27-30, 1929, by Agnes Chase (No. 10,01). Also collected near the tyve local- ity by Agnes Chase (No. 10317). The racemes of the latter specimen are 1 or 2, and the blades are shorter and finer, Paspalum diamantinum is related to P. ammodes Trin. which has been placed by Agnes Chase in ene Ceresia group along with species having a winged rachis. How- ever, it belongs in a separate group closely allied to the "Erianthae," 14. PASPALUM INVOLUTUM Swallen, sp. caespitosi, erecti, 55- 270 ay alti, glabri; laminae 8-15 cm. longae, involutae, ca. 1 mm. latae, fere glabrae, racemi 3, 3.5-5.5 em. longi, adscend- entes; — 3.6 mm. longae, dense longi-pilosaes fructus spicula brevior, acutus, pallidus. 1967 Swallen, New species of Paspalum 369 Perennial; culms tu fted, erect, 55-70 cm. tall, glabrous; gpa sheath densely appressed-pilose, those of the culm elongate, glabrous, the uppermost - nearly Sictelsas: ligule about 1 mm. long, decurrent; axis about ‘ cm. Tone echia = al 2” TM. "wide, pur- ple, glabrous, the margins tS eat scabrous; spike- lets 3.6 mm. long, densely covered with long white hair obscuring the spikelet; geen) shorter than second glume ute, pa type the U. S. ational ph nro No. 1500757, collected in white sand and gravel at Diamantina, Serra de San Antonio, Minas Gerais, Brazil, 1200-1300 meters hhagh” December 27-30, 1929, by Agnes Chase 20. PASPALUM MOLLIFOLIUM Pi sar ae sp. no Culmi dense caespitosi, 55 cm. Seem “nodis appresso-barbatis; laminae 10-22 en. longae, an? mm. latae, acuminatae, dense pilosae; race mi 3 vel inferior 4.5 cm. longus; rhachis costa met marginibus ilosa; spiculae 3 mm. longae, solitariae vel binae, ae. mnial; culms in dense tufts, 55 em. tall, the hodes peep ote sppneased-beanded. with yellowish hairs; sheaths mostly basal, densely pilose with ascending or spreading hairs, the lower one densely tomentose at the 2 densely covered with long appressed white in eennna the enikeLes? fai t a little shorter than the second glume and sterile leaks. pale, acute. ae pe in the U. S. National reherg ‘Yo. Oe Oe eas collected on campo near Far ia, Serra da Boc gy Gerais, Brazil, altitude 1000-1200 ges fg Fanvary Oy 1930, by Agnes Goase.{io. 10sec) <.. Aap Go hee” the type locality by Agnes Chase (No. 1050 21. PASPALUM RIGENS Swallen, Sp. "0° C . se <: 85-95 ian alii, zlabri; vagin ? + laminae e 15-18 ¢ ain Tébaee, 3-5 mm. latae, acuminatae, slabrae; racemi Eee 6-9 em, longi, aa uaanneht oe vel patentes; spiculae . e. im. longae, {bones vanreee® ty 85-9 95 en. aged ee aa ; culn the lower nodes densely appressed-nispid; ri Caner brous, the lowermost densely tomentos 370 PHYTOLOGIA bmi toi blades 15-18 cm. long, 3-5 mn. wide, glabrous, nate, narrowed to the base, narrower than the outn ere the sheath; inflorescence composed of 3 or b. ogee ran or spreading racemes 6-9 om. long, on an axis 5-7 ¢ long; svikelets 4 mm. long, densely ised: siniey-vetiaus, completely obscuring the spikelets; fruit a little rter than the second glume and sterile lemma, acute, pale. Type in the U. S. National Herbarium, No. 1500399, collected of gravelly slope, 2-10 lon. west of Diaman- oni x 1500 meters, December 27-30, 1929, by Agnes Chase (No. 10luj2). Also collected at Faria, Serra da Bocaina, , Minas Gerais, by Agnes Chase (No. 10521); Serra de Cipo, George Black, 22. PASPALUL INTONSUM Swallen, sp. nov. ulmi erecti, 50-70 cm. alti, glabri; laminae 19- 2h. em. » 3-6 acuminatae, conspicue pilosae silis appressis vel patentibus lh mm, long1s; hi dscendentes vel appressi, inferiores usque 7mm. longi; spiculae 1.2 mm. longae, dense villosae. Perennial; culms erect, 50-70 cm. tall, glabrous; sheaths appressed-pilose to glabrous; ligule a membra- nous rin less than 0.5 mm. long; blades erect or ascending, rather stiff, 19-2) cm. long, 3-6 mm. wide, acuminate, narrowed slightly toward the base to the width of the sheaths, conspicuously covered on both surfaces with appressed or spreading hairs, mostly y mm. long, densely subescent just above the ligule 4s well; inflorescence poapcaee or he Sa sve or appressed racemes, the Lo wer as much as 7 cm. long; rachis 1 mm Wide, hispid on the ma aera spikelets 1.2 mm. long, covered with hits silky appressed hairs obseuring the spikelet; fruit a little shorter than the second glume and Peoraee lemma, pale e. in the U. S. National Herbarium, No. 1501433, doxiscted on hillside campo, Hargreaves, ilinas Gerais, Brazil, December 31-22, 1929, by Agnes Chase (Ho. 1026835) . 23. PASPALUM HAUGETII Swalle en, sp, nov. ; Culmi erecti, 130 em. alti basi dense mepwaige, nodis dense aparesse-villosis; yeeaphe marginipus papilloso-ciliatae; laminae 15-26 cm. longae, 0- G=1 cm. latae, marginibus base hausitcacce iliatis ceterun gla- brae; racemi 6, adscendentes, inferiores usque 5 ct. longi; spiculee 3.5 mm. longae, binae brevipedicellatae; glima secunda et Lerma sterile acuta, 5-nervia, dens¢ villosa; fructus ellipticus, acutus, pallidus, striatus, Spiculam aequans, ennial; culms erect, 130 em. tall, from a rhizone-like base, densely villous with white or reddish 1967 Swallen, New species of Paspalum 371 he Bab densely aopressed-villous; lowermost 2 elspa ager | villous toward the base, the upper ones blades@1d Be e back, papillose-ciliate on the margins; 5-26 em. long, 0.8-1 cm. wide, ascending, grad- eee: inflorescence 8- m. long, with 6 ascending - Scant the - r as much as 8 cm. long -9 mm. wide; pikelets 3-5 mn. long, paired, short et eaGed glun : and sterile lemma eoual, acute, 5-nerved, Sun ely villous with ubite nairs, the hairs longer on 3 a By age = Bt long as “the epticeets ptic, acute, pale, striate, as in the U. 8. National Herbarium, No. 1706822 collected in open bunches on llanos, about im. ° Col of Orocue, Comisaria El Vichada, Los Llanos, Olombia, by Oscar Haught (No. 2771). Pasr alum haughtii is closely related to P. : in aller culm and 1 distributed on the which onger and laxer racemes. Another specimen ee tes matches P. haughtii very closely was collected Ba aco, Rio Arinos, lato Grosso, Brazil, by J. Te ldwin (No. 3077). ah. te ue SERICATUM respon sp. nov. C aespitosi, erecti, cae om. alti; vaginae glabrae; pn 5 mm. ietage a acuninatae, glabrae; racemi vel 3, anguste adscendentes; spiteles. har cee 5 see eieake dense culm sheaths bladeless or nearly so, the base sligntly enlar d, tomentose; lower sheaths densely appressed-pilose, the upper glabrous or nearly 803 ligule a short membraneous rim about 0.5 mm. long; blades mostly basal, 5- long, 2-2.5 ma. wide, acuminate, glabrous of both sur- i toward the the axils; a colitary: 3 mm, long, densely villous with white or tawny appressed hairs completely obscuring the spikelet. 8 in the U. S. National Herbarium, No. 125528 ky e Sol, Serra do 7 collected on rocky open slope, Chapeo Ciné, Minas deen: Brazil, altitude 1000-1100 neters, March 28-April 1, 1925, by Agnes Chase HOw = So soecimens may rererrea © the Minas Gerais: Be e species, all from the State of Belo Horizonte, Chase 1 03153 Paria, serra da Bocaina, Chase 10512; Barbacena, Glazion 16558. 372 PHYTOLOGIA Vol. 1h, no. 6 25. PASPALUM cee Each sie Sp. Rey erecti 5 ce nov. 1, glabri, rhizomatibus curtis tomentosis; ‘Tamin em. longae 3-5 rin. latae, @ppressi, 2-2.5 en, longi; spiculee 3. Sa hl ram. longae, solitariae; gluma forge e rm ecunda hirsuta pilis longis urans; Spolings sterile pilosum; fructus pallidus, ate 5. i Perennial; culms erect in loose groups from snort tqiant pee rhizomes, 35 cm. tall, glabrous; sheaths rather densely hispid or papillose-hispid, those ees the culm nearly bladeless, almost glabrous; blades : long -5 mm. wide, sparsely to rather ar eels papal lose-hi sterile Pee ° acuminate, fruit, th scuring the glume, the sterile lemma pilos el ; ee aa evident; fruit shorter than the eeristet: pale, acute the U. S. National Herbarium, No. 1258166, ‘ umber and date not given). Oth er Specimens which may be referred to this species ar bs azil: Rio Grande do : S. Jeronymo, J. Dutra 5 Pedro Orth 19:3. Argentina: Provincia de Corrientes: Mercedes, L. R, Parodi 6295. 26. PASPALUM FPORMOSULUM S _Swallen, sp. no i ecti, oO cm. alti, etapa i; vaginae 3 inferiores dense pilosa ae, basi tomentosae; laminae 0- cm, longae, 3-5 mm. latae, acuminatae, hirsutae; i adscendentes, inferior ca. 6 Oe By ee mam» imped e€, pur i y white-h spikelets; iescng ak +c j pi Lon, cute Type in the U. S. National Herbariun, Wo. eis collected in small quite on hillside campo, Hargr pipet » 1350-1100 sr dad December 21-22, Bae (No. 10268). Also collected by es Chase near the type locality (No. 10226). 1967 Swallen, New species of Paspalum 373 Plicatula 27. PASPALUM INDUTUM Swallen, sp. nov. Culmi erecti, ca. 1m. alti, infra paniculas pilosi; laminae erectae vel patentes, 2-0 cm. longae, 7-10 mm. latae, dense pilosae; racemi 2-6, appressi vel adscendentes, 8-18 cm. longi; spiculae % mn. longae, -0-2.3 mn, latae, obovatae, dense etree fructus wom gee 22 at "Rio do moe pr n March 1, 1909, by P. Dusen (No. 7883) « ‘ Additional material examined: Brazil: Parana: Jaguariahyoa, Dusen 1611; Vilha Velha, Ponta Grossa, Pereira 5202, 52 BE. Mato Grosso: Campanario, Otero 299. Minas Gerais: Serra do Curral, Chase 8923. 28. PASPALUM LUTICOLUM Swallen, sp. nov. mo Culmi 1.5 m. longi, basi 7 mm. crassi, glabr spiculae 2.1 mm. longae, 1.1-1.3 mn. glabrae; fructus 2.1 mm. longus, ferruginosus, stria Mis acutus, : m, long, 7 mm 1; culms about 1.5 wre eLabrows; k alnos brous, the pedicels slender, anos pms pod rumpled, glabrous; second glume and steri light brown, acute, hin, glabrous; fruit 2.1 mm. long; eee re u. 8. ‘National ger i gl cur nage Faded age ee GN nad aah IOS Wilo Tt, Silva (No. June 16, 1952, by J. HM. 4298), 374 PHYTOLOGIA Vol. 14, no. 6 eo, PASPALUM LACUSTRE Chase, sp. nov. compressi, graciles, = m. longi; laminae Dlanae, 3.5-1l cm. longae, 2-l1 mm. latae, glabrae; racemi. l. vel 5, graciles, adscendentes eee cm. longi; spiculae solitariae, 2.5 mn. longae, glabrae; gluma secunda et lemma sterile sequalia. 3-nervia glabra, fructu longiora; fructus 2 mn. longus, O.9 mm. latus, pvallidu Aquatic perennial; culms submerged or floating, slender, compressed, el dbrous, 1m. or more long, the pe age h-15 em. long, shorter toward the summit; S pather loose, stiffly ciliate toward the summit, otherwise glabrous,. the upper longer than the int er- nodes; ligule membranaceous, 1.5 mm. long, erose blades flat, lax, 3.5-1l cm. long, 2-l: mm. wide, " gla- brous; panicle included a base, composed of or slender ascending to somewhat loosely spreading racemes, 3-5-6 cm. long; rachis thin, flat, 1-1.5 ms. wide, gle- brous, the margins scabrous; spikelets solitary, 2. 2 mm. long, on slender pedicels about 1 mua. long, in 2 rows, contiguous but not crowded; second glume and sterile lemma equal, thin, 3-nerved, pale, glabrous, pointed beyond the fruit; fruit 2 mm. long, 0.9 mm. Wide, pale, e in the at tage tee of the Instituto Agronomico do pre tng Belém, Paré, Brazil, No. 61326, collected in water of a deep. ol at Ambé, * AmapA, Brazil, July 9, 1951, by Richard L, Frées & peor ge A. Black (No. 27312). Photograph and fragment in the S. ifational Herbariun. 30. fate CORDAENSE Swallen, sp. nov Culmi 25-50 cm. alti, nodis inferioribus peawigeg ilaminae 8-25 em. longae, 1-6 mm. latae, papillo pilosae, marginibus en racemi 2, 3.5-8 ay “longi Spiculae binae, 3-3.1 mm. longae, 2.3 mn. Later obo~ vatae, dense pnilosae; fructus beneua nitidu al; culms erect or mp eacire, "25-50. em. tall, ing hairs, eae glabrous toward the summit; ligule in & - 8. National Baibars, Wo. 223839; 1967 Swallen, New species of Paspalum 375 to Grajaht, Maranh&o, Brazil, March 1-5, 193l, by Jason R. Swallen (No. 3618). Another specimen of the type collection has racemes, and blades as much as 10 mm. 31. PASPALUM CAPSRATUL Swallen, sp. nov. Culmi centes vel pilosae; racemi 2 vel 3, appressi; 6-8 mm. lata, marginibus scabra; spiculae 2.7-2 longae, 1.5 mm. latae, ellipticae, Fas gluna secunda et lemma sterile grosse ru Perennial; culms in small Garks: eee ho-50 cm. tall, glabrous: sheaths overlapping, completely cover- ing the culm, the lower ones densely pilose, the upper two on the culm glabrous below, becoming pilose toward ane summit, LSD Beanery beh bare ligule 0 1.5 mi. wide, elliptic, glabrous, on short, angular, scabrous pedicels about 0.5 mm. long; second glume and sterile lomia very coarsely cross-wrinkled, the glume ate ae the nerves evenly spaced. in the U. S,. National Ben bar es No. 1961377, hatiaat aa on hill above swamp ta Florestal, Pelotas, Rio Grande do Sul, Brazil, te °53, 194.6, by Jason R. Swallen (No. 920 0}. This species is characterized by the erect racemes and the strongly wrinkled BEACONS, glume and sterile ies with evenly Spaced lateral nerves of ie second glume. ‘ ; t a peeaur collected on the Faz zenda de Cria AL i Lages yan Catarina (Swallen 3138), is referred to this a natee oe. ea AWanNSEh SD. nov. ; 32. PASPALUM PARANAKNS# Swallen, 50 cm. alti, ulmi dense caespitosi, erccti, re Glabri, infra nodis glandulosi; laminas eracbecs = & cm. lo 2-), mm. latae, acuminatae, dense Pi ; pe eS , i; spiculae 2.3-2.5 longae, 1.5 mm, latae, binae, elli : ubglab rs gluma secunda lemma steril ogdeg er eeees brabus cia mm, longus, brumneus, luct = mnial; culms in small dense tufts, oh Pa about 50 em. tall, 3-noded, glabrous, with p a8 be lar depressions below the nodes; sheaths oeiBe ge trorsely pilose, ery_dense ane on rE: the upper two glabrous or pilose aan: ee Sone. 54. co a est mm. Long; blades erect, sae sic | cuminate, nar arrowed to the Beets og acante hem. a pilose, one naire Above the ligule; int 376 PHYTOLOGIA Vol. 1h, no. 6 long, composed of 3 erect racemes h-6 cm. long, the rachis 0.6-0.3 mm. wide; spikelets 2.3-2.5 mm. long, 05 af wide, elliptic or somewhat obovate, in pairs, the pedicels 0.5-1 mm. long, nearly glabrous; second glume and oe lemma shining, glabrous, ia giume 5-nerved, the nerves nearly evenly spaced, very sli aes orto than the ogre a at maturity; fruit dark brown, shining, 2.2 mm. e in the U. Ss. "Rational Bede eky No. 1961379, collected on campo nda Cap&o Redondo, Guarapuava, ke Men Brazil, March 20- 23, 194.6, by Jason R. Swallen No. 33; PASPALUI Se ere Swallen, sp. nov. ‘ C cillimi, erecti, 110 cm. alti; laminae elongatae, attenuatae, 15-30 em. longae, 0.5-1.5 mi. latae, glabrae; racemi 2 vel 3, adscendentes, usque 11 cm. longi; spiculae binae, 2.3-2.5 mm. longae, 1.3- 1.4. mm. latvae, glabrae; fructus brunneus, lucidus. or the upver ones shorter, glabrous; ligule firm, ¢ed- mm. long, protruding from the sheath, brown; plades elongate, attenuate to a fine point, man 30 em. long, pe5-1.5 mm. wide, gee ins curled or flexuous, glabrous; inflorescence 10'¢ - long, composed of 2 or 3 ascending e pedicels 0,5-1 mm. long; second glume 5-nerved, the lateral nerves not crowded, glabrous, at maturity 2 little shorter than the fruit: sterile lemma with a prominent midnerve, the margins broad, reaching almost to the omg ane glabrous; fruit dark brown, shining. . the U. S. National Herbarium, No. 15016, gollesked on campo between Jataby and Rio gerbe rs retien 3 Brazil, April 3, 1930, by Agnes Chase (No. 3l.. PASPALUM PONTANALIS Swallen, sp. nov. Culmi basi decumbentes-patentes, ramosi, ca. 1m. fa 1967 Swallen, New species of Paspele 377 sparsely pubescent; sterile lemma reddish brown with a broad light margin, the neat uate inconspicuous}; uit 2.2 “on long, dark brown, shining. Type in the U, S. National Herbarium, No. 1)866L, collected a sand and muck on Saino embankmen across pontanal, Porto Esparango, Mato Grosso, Brazil, February 28, 1930, by Agnes Chase (No. 11080). 35. PASPALUM MACEDOII Swallen, sp. nov. : Culmi erecti, 110-120 cm. alti, glabri; laminae m nu 2.3 mm. Ta atae; satel secunda { pilosa; lemma sterile glabrum — sparse pilosun; fructus brunneus, lucidus. mnial; culms 110-120 cm. tall, erect, simple, Sparsely pilose; fruit very dark with - wl aca hump in the middle of the lemma. Sons n the U. S. National pee tegretabe No. 2206201, eelkectea ne campo, Monte Alegre de =e motes. February 10, 1956, by A. rasuas “to. pine Also ererabes at Pirineus, Goias, by A. Macedo (No. 4359, 4.366) 36. PASPALUM LATIP#S Swallen, sp. as Culmi- erecti, ee em. alti, suebri vel no ae 15-30 cm. longae, 4-6 racemi 3 ser hy e 3.5-3-7 m Ae longae, 2mm, latae, ea lemma sterile margin Perenni culms erect, 13 oon moor A B the nodes appressed-pilose; sheaths ae inte fricdde;” keokeds glabrous to spars sel pilose . labrour or than the poe about 2 mm. long 378 PHYTOLOGIA Vol. 1h, no. 6 hairs spreading; inflorescence 10-13 cm. long, composed of 3 or 4 narrowly ascending racemes, the lower 9-10 cn. long, the rachis 1-1.2 mm. wide, scabrous on the mar- t lemma with a broad raised Slightly Welakiea margin; fruit dark _brown, striate, shining. e U. S&S, National Herbarium, No. oe collected on satan, Ituiutaba, Minas Gerais, Brazil March & se pale by A. Macedo (ito. 88). ecies is closely related to P. macedoii, but the spiccteta are differently shaped and glabrous. The specific name refers to the broad base of the spikelets. 37. PASPALUM GUARICENSE plgee Nn, sp. no : Culmi graciles, erecti, ramosi, oon 75 a g aude: laminae 15-20 en, longae, 4-7 mm, la atae, et condupli- catae, hirsutae; racemi 1-7, adscendentes, 3-6 cm. longi; spiculae binae, 1.8-1.9 mm. longae, 1.3-1.4. mm. latae, gna eaeaes gluma secunda et lemma sterile tenuia, 3-nervi ae 2 slender, erect, branching at the lower ce 55-75 em. tall; sheaths loose, compressed, keeled, ted cy pee than the internodes, the upper one elongate with a very much reduced blade: ligule about 3 mm. long; blades about 15-20 em. long, -7 mn. t ike wide, conduplicate at the base, hirsute like the eaths; inflorescence 9-12 ong, composed of li-7 ascending racemes 3-6 cm g, the rac 1 ma. wide, long; spikelet in Baee 1.8-1.9 mm. long, 1.3-1.-) mm. Type in the U, S, National Herbarium, No. 13.3986, collected in humid places, El Dividive, Guarico, Vene- zuela, September 11, 1927, by H. Pittier (No. 12520). 38. PASPALUM ATRATUM noe ae sp. nov. ulmi erecti, vala 130 cm. alti; laminae poo Sean marginibus scabris; spiculae 3.1 longae, 2mm. latae, obovatae, glabrae; fructus ferrugineus, lucidus, minute striatus. Perennial; culms erect, stout, 130 cm. tall; sheaths broad, loose, keeled, the lower ones crowded e toward the pase, the upper ones elongate, nearly Avie g the base of the panicle, glabrous or a few long hairs on the margins toward the summit; ligule 1-2 m1. long, hidden in the sheath; gpiaane elongate, 1.5-2 cm wide, very scabrous on the margins; inflorescence 2 cm. long, composed of sR aoahegs 20 rather narrowly ascendiné 1967 Swallen, New species of Paspalum 379 racemes, the lower ones as much as ll: cm. long, 1 mm. face? rather thickish, scabrous on the margins; spike- lets 3.1 cm. long, 2 mm. wide, elliptic, slightly obo- vate, glabrous; second glume and sterile lemma covering the fruit, reddish brown, the lateral nerves of the glume crowded at the margins; fruit reddish brown, shining, minutely striate, the lemma raised in a eee a ump n the U. S. National ager ey: No. 1501223, éolisotad ne moist spot in brushy c , Campo Grande, Mato Grosso, Brazil, by Agne es Chase “aids 108i1.2). Additional material examined: azil: Mato Grosso: Between Bonito and Rodonopolis, Chase 11899. Goias: Golas, Chase 118.6. Minas Gerais: S&o Miguel, Chase 39. PASPALUM FORMOSUM Swallen, sp. no i erecti, glabri, 155 cm. alta laminae 1U- 25 em, longae, 2-3 mm. latee, irmae, "attenuates, re lated, ‘ihicth. broader than the ae cn the uvpermost to 32 om. long with a much reduced blade; ligule ne long, decurrent; blades 10-25 cm. long, 2-3 mu. wide, firn, attenuate, pubescent on the upper surface, the margins scabrous; inflorescence 12 cm. long, composed of 6 ascending racemes about cm. long; pairs, 2,6-2, long, 1.5-1.6 ma. wide, elliptic, 1 mm. long; second glume and sterile lemma rather edie sae fa , th pat ba of the sterile lemma raised, ew obscure cross-wrinkles yp as Pe Geeta) Herbarium, No. 2014969, Collected at San nk Terezinho, tuiutaba, Minas Gerais, i 18 95 species is unlike pe else ae <— . 2 eve sp ulm apake ni ances bop fre oe relatively culm and completely covering it, small blades, are ya cuvegntatees hO. PASPALUM PALUDOSUM Swallen, Sp. HOV. : ue Culmi erecti, graciles, 95 cm. alti; Shaggy te 22 cm. longae, 6-9 mm, latae, erectae, Sh atentes, U5 papilloso-hispidae; racemi 1, erac cues ae, 1.0-1.8 em. longi; spiculae binae, 2-4-2. O Mi. ee edanene. = latae, ellipticae, glabrae; poeple ucidus. e erennial; culms erect from 4 ; i imot fe ge ects Slender, 95 cm. tall; sheaths crowae between sheath the internodes, the line of demarcation 380 PHYTOLOGIA Vol. 1h, no. 6 and blade not evident, the upper two shorter than the internodes, bladeless or nearly so; of tee- 5 mm rescence 6 cm. long, composed of l: slender spreading racemes about 1.5 cm. long; spikelets in pairs, 2.l-2.6 mm. long, 1.6-1.8 mm. wide, elliptic, glabrous, or the second glume pubescent, on pedicels 0.5 mn. long; sterile lemma reddish with pale raised margins, with few cross-wrinkles; fruit very dark brown, smooth and shini type in the U. S. National Herbarium, No. 1613888, = age sire in open wet art ces or in woods, Carolina to San Antonio de Balsas, Maranh&o, Brazil, March 20-25, 1934, by end R. Swallen (No. 3959). 1. PASPALUM RAMOSUM Swallen, sp. nov. : Culmi erecti, nodis inferioribus ramosi; laminae 8-28 cm. longae, 2-l mm. latae, giabrans racemi 5, erecti vel adscendentes, l-7 cm. longi; spiculae binae, -2 mm. longae, 2.1 mm. latae, leviter obovatae, glabrae, Perennial; culms erect, branching at the lower nodes, glabrous, shining; she aths loose, all longer than the internodes, paimonss ligule 1 mm. long, oe d 2 n, long, with a tuft of long hairs in the axils; rachis ma. wide, the margins scabrous, occasionally with 4 few long hairs; spikelets in pairs, 3.2 mm. long, 2.1 mn. wide, slightly obovate, on rather thick sonra pedicels, 1 and 2 mm. long; second glume and sterile lemma hag or the glume sometimes very ara pubescent 68, in the U. S. National Herbarium, No. 22052 collected if sandy moist groun pe at the ena bAsee. Agro~ ico do Sul, Pelotas, Rio Grande do Sul, Brazi November 10, 195k, by José a Costa Sacco (No. ooh). is is one of the very few of the perenni hed 4 of the Plicatula group of Paspalum which has bran enor t is also Dai aue in being almost eotircis labrou Aside from the type, it was also collected at raxtman by Costa Sacco “The. 208 he. PASPALUM CRUSTARIUM Swallen, sp. no 6 evecti, 25-35 cm. alti, sensed laminae O- longi; spiculae binae, 2.l:-2.5 mm. longae, obovatae, glabrae; fructus atrobrunneus, lucidus, striatus, valde convexum, 1967 Swallen, New species of Paspalum 381 Annual; culms erect, 25-35 em. tall, : glabrous; sheaths all longer than the internodes, glabrous, or Spreading racemes 2-l, em. long, the rachis 0.7 mm. wide glabrous; spikelets in pairs, 2 Renee mm ; glabrous, the sterile lemma flat, embossed, ee striate, a lemma strongly co ° : in th s. Netional Pennine: No. heey co lected in open we between Carolina and tonio de Balsas, Maranh&o, Brazil, March 20-25, 1931, by Jason R. Swallen (No. 3985) . z Additional material examined: erage Haranh&o: Grajahh to Porto Franco, Swallen 3826; Carolina to San Antonio de Balsas, Swallen 3947, 3963, 3995 i092, 4202. these specimens vary in size from the e, much as 60 cm. tall with blades to 5 mm. ciate and 5 racemes. The spikelets, however, are identical. 43. eons ise ae i Swallen, sp. no Cte occ ane alti, zlsbri; “laminae 20-35 pk longa “6 "ihe 7 mm. latae, ad basim attenuatae, piers: centes; racemi lh, erecti vel adscendentes, 13- longi; spiculae atrobrunneaze, 3.5 mm. longae; nedicelld breves, summo Eee gluma secunda et Sterile grosse rugosa ennial; culms erect, 2.3 m. tall, glabrous; sheaths elongate, all longer than the internodes, nar- rowed at the surmit, the lower ones pubescent or pilose; ligule 3 mm. long, brow, i er sheaths; blades 20-35 em. lon i attenuate toward the base to the eee of fue sh doti tly ok ra- and sterile lemma very coarsely transversely wrinkled covered by the glume and sterile lemma Type in the U. 8. Hiationsl 3 pombe cig No. 1961391, collected res small banhad da Palma, cee : reande do Sul ate Horie : 23, 1945, by Ja . wallen (lio. 918 is Lega a robust species of the Plicatula group, hy. PASPALUM Bedtenaruent Swallen, sp. nov. : compressi, LO cm. alti; laminae 3 marginibus scabrae; er mm. latae, conanpiababas, elabrae, 382 PHYTOLOGIA Vol. 14, no. 6 racemi yea; “mn ocr cake 3-5 cm. longi; spiculae bine 2.5-2.6 mm. longae, 2 mm. latae, ellipticae, glabra vel iiae secunda pubescens; fructus atrobrunneus, minute striatus. Perennial; culms erect, flattened, about 0 cm. tall; sheaths crowded toward the base, 3-0 cm. long, the culm sheaths elongate, conspicuously ye rei and keeled, glabrous, the basal ones as much a mn, wide as folded; ligule brown, membranaceous, 1 Sei long; blades conduplicate, BPepyeeee narrowed from the base, as much as 3 mm. wide as es ar nape pics: except for or the second glume more or less pubescent; pedicels short and thick, about 0.5 mn. wide at the summit; fruit dark brown, finely striate-roughened, Type in the U. S, National Herbarium, No. 1961358, collected in wet ground, margin of penhado, Fazenda da Brigada, Passo Fundo, Rio Grande do Sul, Brazil, December 19, 1915, Dy Jason Ee Swallen (Ho. 7790) « The following specimens seem to belong to species although the sheaths ane not so sanaceaaanaly equitant. Brazil: Rio Grande do Sul: Passo F ee Se 5200, 5252. Santa Catarina: Lajes, Reitz & in 13965, 1065; Cagador, Klein 3551; Ss. Joaquin, 6 Ra itz & Klein 7881: Santa Cecilia, Reitz & Klein 116. us. fest VALIDUM Swallen, sp. no grossi, erecti hin m. alti: laminae usque 50 sa longae -1.5 cm, jatae, prope margines pilosae vel hispidae; racemi 18, adscendentes vel patentes, imus ca. 9 em. longus; spiculae binae, 2.5-2.7 mm. longis; gluma secunda sparse eee fructu paulo brevior; Reascr - fuscus, lucidus C coarse, erect, 1.5 m. “tall; sheaths ew + well marked; blades as much as 50 cm. long, 1-1.5 cm. wide, usually pilose near tne margins, or sometines sparsely hispid; inflorescence 22 cm. long, pyramidal, composed of 15 ascending or spreading racemes, che pedicels, as much as 1.5 wm. long; second glume sparse- ly TeILO Res a little shorter than the fruit, the sterile emma glabrous, equaling the fruit; fruit dark reddish brown, shining, 1967 Swallen, New species of Paspalum 383 Type in the U. S. National pepeterctanrs! No. 1613226, collected at base of —- re in chapada, between o, Brazil, March 5-13, ty PL ~ 6 B a a po EP S BG an: Por 193k, by engi Rs Swalien rm 378). 6. PASPALUM VIALE Swallen, sp. nov. Culmi erecti, 55-80 cm. alti, glabri; laminae ca. 20 cm. longae, 2-h mm. latae, acuminatee, pilosae; racemi 3-5, adscendentes, inferiores 10 em. longi; spiculae 2,8-2.9 mn. loncae, 1.8 ma. latae, ellipticae; i eg hecoiaas et Lemma sterile atroferruginea, lucida, giabra, Perennial; culms tufted, erect, glabrous, 55-80 em. tall; sheaths all longer than the internodes, densely pilose sci spreading hairs as much as 5 ml. long; ligule 3-l mm. long; blades about 20 om. long, 2- mm, wide, gradually narrowed to the base, acuminate, pilose like the sheaths; inflorescence partly enclosed in the elongate upper sheath, composed of 3-5 ascending mm. long, 1.8 mm. wide, elliptic on scabrous pedicels 0.5-1 mm. long; second glume and sterile lemma dark reddi brown, both rather tly cross-wrinkled, shining, glabrous; fruit dark bernie smooth and shining, the back of th sed. t “ lemma evenly ra No, 1916368, Type in the U. 8. perm Herbarium, collected ane roadside, Campo Largo, near Curitiba, a “ge: cigs February 15, 19i15, py Jason R, Swallen °. 91 This species is definitely related to Paspal = Z pannuceum Swallen, which differs in being nearly gla brous end in having larger spikelets. th species = have the evenly spaced nerves on a second aii an the fruit evenly curved on the ba 47. PASPALUM OTHROII Swallen, sp. novs Aan Cu decunibentes, anpeecenrt Si —_ maciem i. glabri, usque 1.5 m. longi; laminae 6-15 cm. ccaet 3-5 mm. latae, acuminatae, narginis eS tlnes i. vel 5, adscendentes vel appressi; spic eg longus, ae a a dense pubescentes: fructu S 2. atrobrunr lucidus. an hati 1; culms " decumbent-spreadings ene ou rooting at the nodes, glabrous, 45 much 0 so cig, the internodes of nearly equal see ve ‘ é sheaths 1/2-1/3 as long; blades 9--~ wide, acuminate, nearly glabrous, oF ae = eee papillose-pilose toward the base, = or appressed, very scabrous; racemes h or 55" ase an LEO: Tie 3 mm, long, very —- dark brown, shin 38h PHYTOLOGIA Vol. 14, no. 6 Type in the U. S. National Herbarium, No. 1982726, collected at the Seccao Experimental de Agrostologia, ato Grosso, where it is considered to be a good forage grass. It is easily propagated by cuttings. Anachyris 48. PASPALUM PLANIUSCULUM Swallen, sp. nov Culmi erecti, 160 cm. alti, compressi, pilosi, nodis dense villosis; laminae usque 65 em. longae, l. 5 mm. latae, infra pilosae, supra glabrae, marginibus scabrae et hispidae, superiores 15 em. longae, 2 cm. atae; racemi ca. 0, adscendentes vel patentibus, penny usque 6-7 cm. longus; spiculae 1.6 mm, longae, rennial; culms erect, 160 cm. tall, conspicuous- ly flattened, apparently soft and spongy, appressed to 8 e airs; sheaths much longer than the internodes, com- pressed, keeled, as much as 38 cm. long, appressed- pilose or the uppermost nearly glabrous; blades as much as 65 em. long, 1.5 mm, wide, about half the length narrowed, petiole-like, pilose beneath, glabrous above, composed of about 10 ascending or spreading racenes, the lowest as much as 6-7 cm. long, the rachis 0.7-1 mi. wide, scabrous on the siete spikelets 1.6 mm. long, 0 : -8 mm, wide, Brabiows : the fruit white Type in the U. S, National Herbarium, No. one us d and Porto Franco, Maranhio, Brazil, March 8-13, 1934; by Jason R, Swallen (No. 381). ‘This species is char- acterized by the conspicuously flattened culms, and the elongate sheaths and basal portion of the blades. 9. gd igen TENUIFOLIUM Swallen, sp. nov. ecti nodis inferioribus ranosi, 1.5 m. aitt; soati: boda villosis; pe tenues, 9-i10 he gracilibus 0.5-1 mm. eg ogy culms erect, br anching at the lower nodes, 1.5 m. tall, the nodes densely villous; sheaths about as long as the internodes, papillose-pilose with spread- ing hair 3 blades thin, 9-0 cm. long, 1.5-3 cm. wes th s very scabrous, with a dense tuft of long hairs above the ligule; 1967 Swallen, New species of Paspalum 385 zpeceaeronnee about 22 cm. long, composed of nearly “ia ascending racemes, the lower 9 cm. long, the tee lie mm. wide, the margins scabrous; spikelets 1.6-1.7 mm. rong, 0.8-1 mm. wide, on slender pedicels 0.5-1 mm. ong. Type in the U. S, National Herbarium, No. {abana collected . wet banks along ditch, Caxias to Corda, Maranh&o, Brazil, February 16-26, 193}, or —. R. Swallen (No. 3602). This species also was collected Sacuane Barra do Corda and Grajahd, Maranha&o, Swallen (No, 3766), and between Carolina and San Antonio de Balsas, Maranh&io, Swallen (No. 4017). 50. PASPALUM COSTHLLATUM Swallen, sp. nov. : Culmi gracillimi, 12-28 cm. alti, erecti, glabri, nodis inferioribus ramosi; laminae usque em, longae, 4-8 mu. latae, hispido-ciliatae, infra et supra hir- sutae; racemi 3-6, patentes usque 2. Py em. longi; spiculae solitariae, interdum binae, 1.3 mm. longae, glabrae. ual; culms very slender, erect, glabrous, branching at the lower nodes, 12-28 cm. tall; sheaths mostly shorter than the internodes, keeled, glabrous or hearty so; blades as much as 6 em. long, -8 mm. wide hispid-ciliate, appressed-hirsute on both surfaces; racemes 3-6, spreading, up to 2.5 cm. long, the rachis 1-1.3 mm. wide, scabrous on the margins; spike ets nostly pol ehees sometimes in pairs, 1.3 mm. long, gla- brous; fruit whit Type rr Ae ve S. National Herbarium, No. 1613883, collected in chapada, Carolina to lags sees de Balsas, Maranh&o, Brazil, March 20-25, 193l, by Jason #. Swallen (No. 3955). his species was ARP ee in 1962 by Hiten & Eiten, in chapada near 'Grota ESO De " trail to Santa Barbara, Municipio de Loreto, Maranhao 51. PASPALUM HITENII Swallen, sp. no ulmi graciles, erecti vel yap 55-100 cm. alti, glabri, straminei; laminae elliptico-Lanceo- latae, l-16 cm. longae, 6-15 mm. latae, basi lon ner pk : tbe infimus usque 6 pilosae; racemi 7 chee Batentes, rasa - “Latee, ile oak. culms slender, erect or ascending, og 100 cm, tall, soft, glabrous, straw-colored, shinin the sheaths all shorter than the internodes, glabrous, lower up to 5 em. long, the rachis 0./ mm. wide, ga brous on the back, the margins scabrous; SP 386 PHYTOLOGIA Vol. 1h, no. 6 1.6 mm. long, 0.7-0.8 mm. wide » glabrous, the pvedicels about 0.5 mm, long; fruit white or light yellow, the lemma athe S-nerved. Type the U. S, National Herbarium, No. 23291, collected hn ravine of dry brook in extensive "caatinga" forest, between Rios Balsas and Parnaiba, “Ilha de Balsas" region, Rahat de Loreto, Maranh&o, Brazil, Avril 6, 1962, by George Hiten and Liene T. Hiten (No. 4091). This species was also collected in the same regio by Eiten ny Eiten (Nos. 3901, 398, 1078, 4089, and 273). Additional material exami ined: Brazil: Maranh&o: an Antonio de Balsas, Swall S68 Piaui: Swallen 2075 Cear&: Crato to Barbalha, SE 373; Baturite to ga rret rango, Swallen 1):28. ese So anal have been referred to aspalum malaco sues: frin., which is a perennial eeres: 3S, much different Fran PR. eitenii. Ungrouped 52. PASPALUM CLANDESTINUM Swallen, sp. no i erecti vel adscendentes, 73-28 cm. alti, cm. longus; rhachis 1 mm. lata, marginibus scabra; Spiculae solitariae 3 mm. longae, 1.5 mm. latae, ovatae, os a nervia, villosa; fructus 2.5 mn. ee? plano- -~convexus, ellipticus, p ae minute striatu culms erect or secon 13-28 cm. tall, ual; glabrous; sheaths all longer than the internodes, gla- brous on the b 2 oe ion tary, almost entirely enclosed in the sheath, about 10 cm. lone, slightly arcuate, the meee about 1 mm. wide, minutely iy oe on the margins; spikelets solitary, 3mm. long, 1.5 mm, wide, ovate, acute, the first glume aduate to it; second glume and sterile lemma spotted, equal, 3-nerved, the lateral nerves marginal, lightly covered with white Silky hairs; fruit 2 2 ong, ecEEieace broadly elliptic, pale, minutely striate Balsa anhio, Brazil, Mirchi =n oe, 193k, by Jason e Sista dey Gio, hoéh). cere ee be exserted later, but the spike- lets do ane mature 1967 Swallen, New species of Paspalum 387 53. PASPALUM SCHULTESII Swallen, sp. nov. Culmi dense caespitosi, erecti, 16-35 cm. alti, glabri; laminae involutae, usque 25 em. longae, supra dense scabro-pubescentes, infra scabrae; racemi 2, con- Jugati, adscendentes vel appressi, 1.5-3 om. longi; Spiculee obovatae, acutae, brunneae; pedicelli con- pressi, ciliati, 1 mm. longi; gluma secunda et lemma Sterile teuissima, nervis inconspicuis; fructus face, glabrous on the outer, sometimes curved or flexu- ous; racemes 2, conjugate, ascending or appressed, 1.5- 3 em. long, the rachis glabrous, even on the margins; Spikelets ovate, acute, brow, on flattened, ciliate pedicels 1 mm. long; second glume and sterile lemma very fragile, the midnerve suppressed, the marginal nerves faint; fruit obovate, dark brown, smooth and shining, Type in the U. S. National Herbarium, No. 2150293, collected near the Rio Paran& Pichuna (tributary of Rio Vaupés), Vaupés, Colombia, June 1953, by Richard Evans Schultes and Isidoro Cabrera (No. 1993l). This species does not belong to any group of Paspalum Imown. It is characterized by the involute blades and two conjugate racemes of brown spikelets. erennial; culms rather coarse, erect, pete ee 1m. tall; sheaths longer than the internodes, hispi¢ or papillose-hispid with long appressed Shorter on the uppermost nearly bladeless sheaths; ligule a brown membranaceous rim 0.1 rm. meee ss 2 amet flat, 30-40 cm. long, 8-12 mm. wid ees Stok ar ong, becoming shorter and more crowded towar , spikelets 2.6-2.7 mn. long, elliptic, on peels mi. long: first glume evident, ciliate; se 388 PHYTOLOGTI A&A Vol. 14, no. 6 little shorter and narrower than the fruit, appressed- pilose; sterile lemma as long as the fruit, appressed- pilose especially on the margins; fruit pale in the U. S, National Herbariun, No. 2205975, collected on the Macap4-Clevelandia roa ferritorio do Amap&, Brazil, July 22, 1951, by G. A. Black and R. L. Frées (No. Bre £2357) « 55. PASPALUM TELMATUS Swallen, sp. nov Culmi dense caespitosi, erecti, 50-60 cm. alti, glabri; laminae firmae, erectae, acuminatae, 20-30 cm. longae, 3-6 mm. latae, dense appresso-pilosae; racemi 5-7, rigide adscendentes, 3-5 em. longi; spiculae 3 mm. longae, 1.3-1.5 mm. latae, acutae; gluma secunda et lemma sterile tenuia, glabra, fructu 0.5 mm. lonsiora; oe ws 5 mm, longus, pallidus, obovatu nnial; culms in dense tufts, msi ey 50-60 cm. tall, ea bvckis: 1s @ a very short glabrous rin; sheaths mostly crowded toward the vase, of nearly equal 1 : cm. long, mm. Wide, densely oe ee aes 3 perro 6-8 cm. long, composed of 5-7 stiffly ascending racemes, 3-5 em. long, with a few hairs in c T tong, 1.3-1.5 mm, wide, acutish, on slender glabrous pedicels 0.5-1 mm. long; second glume and sterile lemma thin, glabrous, pointed beyond the fruit 0.5 m., glume with the lateral nerves very close together; frui 5 mm. long, pale, slightly obovate, rounded. Type in the U. S. National Herbarium, lio. 1500550, collected on wet sandy margin of lake, Logoinhes, be- tween Campo Grande and Dourados, Mato Grosso, Brazil, by Agnes Chase (No. 10926), 56. PASPALUM sn igh Swallen, sp. nov. 5-60 i erect om. alti, glabri, on ecoaigats Pee curtis; laminae riahas, planae, erectae, 6-10 cm. longae, 1.5-2.5 mn. latae, sparse papillosae ae papilloso-pilosae pilis 2-3 ma. longis; racemi l-6, Dg ee inferiores usque 7 cm. longi; spiculae mm. longae, 1.1 mm. latae; gluma secunda et lemma sterile tenuia, 3-5-nervia, glabra; fructus pallidus, lucidus, minute striatus. rennial; culms erect from a short rhizome-like t long, 1.5-2.5 mm. wide, with a few widely spaced large papillae, or papillose-ciliate, -” hairs spreading, 2-3 mm. long; inflorescence 5-7 em. long, composed of L-6 ascending racemes, the ene much as 7 cm. long, 1967 Swallen, New species of Paspalum 389 long-pilose in the axils; spikelets 2.8 mm. long, 1.) mm. wide, the second glume and sterile lemma rather thin, 3-5-nerved, glabrous; fruit pale, smooth and shining, minutely striate e in the U. S. National Herbarium, No. 1961595, collected in ahel isu a ct aoe pee ed, Fazenda Capo Redondo, Guarapu and, Brazil, March 20- 23, 1916, by Jason R. shalien: fist 8837). Swallen 8 collected at the same locality, belongs to this species. ST. PASPALUM MORULUM Swallen, sp. nov. Cu erecti, 35-60 c sei alti, glabri; laminae 15- 20 em, longae, 2-6 mm. latae, acuminatae, papilloso- Dilosae; racemi 2-l, adscendentes, 1-3 cm. longi; Spiculae binae, 2.l1 mm. longae, 1.8 mm, latae, plano- convexae, 1.5 mm. crassae, fructus pallidus, jucidus. culms erect, 35-60 em, tall, glabrous; t toward the acuminate tip; inflorescence 2-6 cm. long, composed of 2-l, spreading racemes 1-3 cm. long, with a few long hairs on the margins of the rachis and in the axils; Spikelets in pairs, 2.1, mm. long, 1.8 mn. wide, plano-convex, 1.5 mm. thick, slightly obovate, rounded, mottled, light-colored, finally turning to a deep brown or black; fruit the same size and shape as eff spikelet, pale, sm aye and shining. Ua in S ee sorb eral sate 24.3293, Brest 1, April 11, 1962, by oarad Hiten & Liene T. Hiten (No. 250). 58, PASPALUM SWALLENII Chase, sp. nov. o Swallen, racemis brev ioribus et spiculis brevioribus 1.5-1.6 mm. longis differt. imilar in all respects to P. morulum Swalien, except for the shorter ~ponepates and smaller spikelets, the latter _1.5-1.6 mm. long. Type in the U. S. National Herbarium, No. 223806, Collected in red sandy ao in AB pe between Carolina 8nd San Antonio de Balsas, Maranh&o, Brazil, March 20- Eos 193k, by Jason R. anal tes (No. 0). specimen was indicated by Mrs. Chase as a new siaeton many years ago, but no description was ev written BOOK REVIEWS Alma L. Moldenke "Wild Flowers of the United States", Volume I the Northeast States Parts 1 & 2 by Harold Willian Rickett, en vi, & 559 pp., illus., New York Botanical Garden and McG: aw-liEL1 Book Company, New York City, New York. $39.50. 1966. Irresistibly I have spent hours upon hours joyfully admiring this work of beauty, sotentiic accuracy and thoroughness. Many more will be so spent b elf and all f , regardless of their than half of the known total of wild flowering rage pra exclud 1, ly re graphs usually taken in situ for easier ee pee they are the glory of the book with artistic credit to each of the 51 con- tributing nature photographers, including the author. 350 helpful line drawings. The text is simple to read and has ac clear print. There is a glossary with clearcut definitions. is a amilies considered and then further keys species. on and scientific names are listed, and in the intro~ duction they are wisel lained for the ama e region covered is from Maine to Minnesota south to Missouri and Virginia. The price is modest considering the size (9 7/8 x 13 L/h s % the superior paper and type, the quality photographic reproduc and the research essential for so large an undertaking het toally, the price is partially subsidized by the David Rockefellers to whom the work is dedicated and some foundations "Die grossartige Weld der Sukkulenten" by Werner Rauh. 18h pps, 730 illustrations (62 in color). Verlag Paul Parey, Hamburg & Berlin, Germany. 98 DM. Succulents have always been favorites among garden enthusiasts because of their bizarre appearance and often extremely handsome flowers. Among these the Cactaceae are the best known and most popular, = Petre are not included in this book. There are ucculents in other plant families and it is with these that aie present work is concerned. Many of these non=cac td 390 1967 Moldenke, Book reviews 391 are very often mistaken for and confused with true cacti by the uninitiated. The author divides his subject into stem-succulents and leaf- succulents. In the former group he discusses 50 —_ ra in the Apocynaceae, Asclepiadaceae, Compositae, Crassulaceae, Cucurbita- ceae, Didieriaceae, Euphorbiaceae, Foiadatandal Se ao Zeacinaoese, Uorscene, Passifloraceae, Pedaliaceae, and Vitaceae. In the second group he discusses 66 genera in the Compositae, Crassulaceae et nla Liliaceae, Mesembryanthemaceae, and Portulacaceae. One wonders why Agave, the xerophytic bromeliads, and some of the alpine saxifrages are ig plants, like Cakile, Mallotonia, Ipomo pes-caprae, Salicornia, and of Achyranthes are also certainly succulent, but are not included in the book, The entire begonia te is ignor An interesting map is provided showing the desert, semi-—desert, and semi-arid regions of the earth. The illustrations are magni f- val of the plants in cultivation is recommended and the months when pesned fron from and enj pong in this beautifully printed oe eee work, "The rape of Cultivated Plants" by Franz Schwanitz, transla e German by Gerd von Wahlert, vi & 175 illus. oe University Press Cambridge, Massachusetts, and Qx- ford Thiverstty Press, London. 1966. 275 Harvard has rendered another fine service to the field of the Biological sciences in bringing this valuable study to the general and one English reading public. € author shows clearly that the transition from the wild Species to the corresponding but distinct cultivated ones is due orti breeding ary lupines in Germany, the author illustrates how long cultivated plants probably evolved from wild speci culti vated forms as the cultivated traits appeared one after another 392 PHYTOLOGIA Vol. 1h, no. 6 through gene mutations and correspondingly how the wild traits gradually disappeared. He gives a striking example of how a somatic mutation inducing an increase in cell size caused the formation of doubled flowers simultaneously in a mixed head of ae chinensis. He stresses the importance of Vavilov's eight gene ee in reas the world in subtropical and tropical ou aus e of their greater tempe . i sure to stronger ultra-violet light, and their tremendous range in microclimates ease the mutations d establish more rapidly the successful ones with wild plant characteristics inherited as d ts in the centers of such areas, and with the omied eae ones inherited mainly as recessive traits on the peripheries, his book is crammed with valuable ideas and supporting facts presented in easy, direct language, and is well illustrated. "Gymnosperms — peek and Evolution" by Charles Joseph Chamberlain, & 18 lh pp., illus., paperback republication of o original. aileenerey of Chicago Press 1935. Dover Publi- cations, New York City, New York 1001). 1966. $2.75 This unabridged reprinting puts this classic back in circula- tion so that today's botany students can learn from its exce ia nt 397 illustrations and detailed text the anatomy, the life tories with their expanding sporophytes alternating with their RS pense the fossil records and the evolution i gymnosperms. The t+ part of the ia cats with the ancien Cyca dofilicales Sinibacot vat on the back er) and etal and their related modern cycads. The sagt part deals with the ancient Cordaitales and their related modern Ginkgoales, conifers, Gnetales, and the fantastic Welwitschia. The bibliography is the most detailed one in print up to the date 2° the oeisuat publication. -Patapconaaael gives assurance that the binding is sewn and "Drawings of ages Plants — Being Illustrations of the Species of Flowering Plants Growing Naturally in the British Isles G. Bell and Sons, Ltd., London. 1966. 12 sh. 6d. The same high quality of accurate and attractive black-and- white wae sueetaan is shown in papas work as in the previous parts. ere is an index. Time lapses between the appearance of aig Pei reasonably short, PHY TOLOGIA Designed to expedite botanical publication Vol. 14 May, 1967 No. 7 CONTENTS REED, C. F., Grasses new to eastern United States... .. ,° 3. 393 MOLDENKE, H. N., Additional notes on the genus Priva. V. ... . 394 MOLDENKE, H. N., Notes on new and noteworthy plants. XLVIL . .398 MOLDENKE, H. N., Additional notes on the genus Lippia. Il. . . .400 MOLDENKE, H. N., Additional notes on the genus Cornutia. Il . . .420 MOLDENKE, H. N., Additional notes on the genus Citharexylum. IV . 429 MOLDENKE, A. L: BOGk TEVMWS 4g Rr es eee <4 ee | DWYER, J. D., Three new species of Ouratea (Ochnaceae) from Posaes 8 eee ee a 439 JABLONSKI, E., Notes on neotropical E papa ue: 1. Synopsis of SOuth American S€MEM ee ee ee Ee JABLONSKI, E., Notes on — Siew Bc as 2. New species | tid heeslers = ee Se ee eee : - 450 Published by Harold N. Moldenke and Alma L. Moldenke 303 Parkside Road, p 0 Plainfield, New ens? 07060 Price of this number, $1; per volume, 85.751 in advance, — : fe t close: et volume < - Missoun Sorc se wns ice GRASSES NEW TO EASTERN UNITED STATES BY CLYDE F. REED* The following species of oi: yes seem to be new to the flora of Eastern United States, The specimens have been de- termined by Dr. Jason Swallen, perio of Grasses, United States National Herbarium, All the specimens are in the Reed Herbarium, unless noted otherwise. Andropogon traceyi Nash South Carolina: Roadsides, Sharon, York Co, Dec, 26, 1955. Reed 37164. Eragrostis oxylepis (Torr.) Torr. Virginia: at Fort Monroe, Hampton, Common in the sandy lawns. Elizabeth Co, Nov. 15, 1959. Reed 45919. Also in US. Festuca dertonensis (All.) Aschers. & Graebn. Delaware: Edge of woods just east of Noxentown, New Castle Co, June 13, 1960, Reed 47196. Also in US. Heleochloa schoeniodes (L.) Host. Delaware: Wharves, New Castle, New Castle Co. Oct. 21, 1863. Albert Commons. (Reed Herb, js Many records for species of grasses new to Eastern United os were pibitohed by the author in 1964 (Phytologia, 10(5)3 Beatty and 1965 (Phytologia, 11(5): 289). pong Herbarium, 10105 Harford Road, Baltimore, Maryland, 34. 393 ADDITIONAL NOTES ON THE GENUS PRIVA. V =.» Harold N. Moldenke PRIVA Adans, oe mG ca « Additional bibliography: Moldenke, Phytolo 73, 75=-79, 95, 98, 105, 107—111, & 511 (aes), 2: 232, 23h, 1 2, 239, & 22° (2958), = é: 511. 19595 Srinivasan & Agarmal, Bull . Bot. Surv, India é: 1963; S. V. Ramaswamy, Bull. - Surv. India 6: 17. 196k; acer t rtolbgta Lh: 336—-357. "1967. ADHAERENS (Forsk.) Chiov Additional bibliography: Moldenke, Phytologia 14: 338--339 & e 1967. PRIVA ASPERA H.B.K. Ton found this species growing on a slope with Quercus, Pinus, and Liquidambar, Additional citations: MEXICO: Chiapas: A. S. Ton 372 (Ws). PRIVA CORDIFOLIA (L. f.) Druc A garwal, Bull. Bot. Surv. India 5: 68. 1963; S. V. Ramaswamy, Bot. spol India 6: 17. 1964; Moldenke, Phytologia 1): 338 & PHS. 3 Ramaswamy (196),) anes that this plant grows oe “nedges along in India. —e CURTISIAE Kobuski og cr tional bibliography: Moldenke, Phytologia 1): 343, 3h7, & PRIVA LAPPULACEA (L.) P Addi tional Stitt coraees Moldenke, Phytologia 1): 3h5 & 350— 7. Additional citations: MEXICO: Oaxaca: Martinez-Calderén 7h) (Mi). oo — H. H. Smith 1165 (Mi). BRAZIL: “Mat- togrosso: F. hne Com. Rondon 5695 megilie Paré: Dahl, & dotle TH G05, Fah 3 D. A. Lima 53-1294 (ie—50855) Bo LIVIA: El Beni: H. H. ae eS ae (W--1,3521, Ws). La Paz: M. 1375 (E—118800, W—l3520); H. H. Rusby 698 (D— 645861, W--126L155). GHANA: J. B. Hall 2517 (S). INDONESIA: Den Berger 792 SO). Vea OF COLLECTION UNDESIGNATED: Herb. A. Gray 157 (E—- 118767) « MOUNTED ILLUSTRATIONS: Kobuski drawing 10 (£92506), 19 (E—925l05) « PRIVA MEXICANA (L.) Pers. Additional & emended synonymy: fructu aparines Dill., Hort. sian. SEGe a: Se Hee fig. x. 1967 Moldenke, Notes on Priva 395 1732. Verbena mexican. J. A. Murr. in L., Syst. Veg., ed. 13, 62, 177k. Verbena dian diandra, spic. laxis, calyc. fructus reflexis rotundato~didymis hispidis J. A. Murr. in L., Syst. Veg., ed. 13, 62. 17h. Verbena diandra, spicis laxis, calyethe fructus re- flexis rotundato—didymis h: hispidis L. ;. apud H. EB. Richter, Cod. ris Linn. 35. 1835. Verbena mexicana, trachelii fol., fructa « aparine: Dill. a H. E. Richter, Cod. Bot. Linn. 35, in syn. 1835. = bena mexicana, trachelii fol. etc. Dill. apud Peterm., Cod. Bot. Linn. Ind. Al; Tn ips 196, in syn. 1540. Verbena mexicana Pers. ex Steud., - Bot., ed. 2, 2: 750, in syn. 18h1. Blairia eee Gaertn. er Bocq., idaceechd a 3: 239. 1863. Priva fa mexicana oa Benth. in Benth. & Hook. f., Gen. Pl. 2 (2): @)s als: ae pies P. mexicana Sieber, 181]. Drepania mexicana Pers. ex M e, Phytologia 5: 108, in text. 195). Verbena uncinata baie ex Moldenke, Résumé Suppl. 3: lil, in syn. 1962. Priva spicata A. L. Additional & emended bibliography: Dill., Hort. Eltham, 07— 408, Be 302, fig. 389. 1732; e Syst. Nat., ed. 10, 852. 1759; J. A. Murr, in 1 yet: Veg-, ed. 13, 62. 177h; Dill., Hort. El- 1 Pflanzensyst. Houttuyn Ss 122. 1779; A. L. Juss., Ann. Mus Hist. Paris 7: 69. 1806; M. Mirbel, Ann. Mus. Nat. Hist. Nat. 3: 239. 1863; Bocq., Rev. Verbenac. 9--96 & 239. 18633 Hook. f. & Jacks., Ind. Kew. 2: 628. 189k; M. pom Anatom. Unter- I such. Verb. 55. 1911; Loes., Verh. Bot. Ver. Brand. 53: 80 [Ab- « 245). 1912; Britton & P. Wils., Sci Surv. Porto Rico 6: 1h5. 1925; Stapf, Ind. Lond. 6: 430. 1931; J. Rz Esc. Nac, Cienc. Biol. 8: 105. 1954; J. Rzedowski, Ci : 1k6, 147, & 150. 1955; , Dict. Terms og. 128. 1955; 9 , Act. C “ 8. 1959 Select. Guide Lit. Flow. Pl. Mex. 02 & 1010. ; feck; Résumé Suppl. 12: 12. 1965; Moldenke, Phytologia 1h: 3h Saeiremcee Dill., Hort. Eltham. pl. 302, fig. hee fa 8 Dill., Hort. Eltham. Add, Denom. Linn. pl. 302. 177k; M. Mi Ann, Mus, mit. Hist. Nat. Paris 15: pl. 1) (2), fig. 2 Porites Recent collectors describe this plant as an upright bo shrub tall, "s with several vir- gate stems from the base and apaticl ee of long spikes", the fruit easily detached, adhesive. They ha os it growing in open matorral, open shrubbery, and wet s in Myrtillocactus associ- 396 PHYTOLOGIA Vol. 14, no. 7 ation, and in steep south-facing Quercus woodlands with occasional Pinus, Arbutus, Ce » Mimosa, , Erythrina, etc., Saou the wn Pos poadebr of oak forests, on oak flats, tufaceous mo small trees of Acacia, Mimosa, and Opuntia, at altitudes of 50 to 2460 meters, flowering from flay to October, and fruiting from June to Dec - Waterfall encountered it near a mountaintop with a few scattered longleaf pines. It has been leer from the is- land of S A common name recorded for this plant is "Mexican vervain". corollas are described as "lilac" on J. A. Steyermark 16287; "pink" on Lundell & Lundell 12351; "purple" on J Je Rzedowski 4gh25 — on Breedlove 11)10 & 12272 and S. S. White 1566; "white and pink" on Stanford, Lauber, & Taylor 2063; "pinkish-white, most- ly white" on Stanford, Lauber, & Taylor 267h; and “purplish-red, the upper lip lined, the tube pale" on | R. Mc McVaugh 16665. R. M. King 1967 exhibits es and fruiting-calyxes that greatly re- semble those of P. lappulacea (L.) Pers., but the cocci are smooth. Possibly it represents a ° dditional & emended citations: MEXICO: nar manages Re Mc Vaugh 16665 (Mi). Chihuahua: Correll & Gentry 22770 (Ld); Gentry, Correll, & Arguelles 17962 (Ld); Pringle 1354 Gentoo oe ¥—1,81216). Coahuila: “Edw, Palmer 251 281 (Ca--139745, E—118805, W—336277); Pur- pus 1096 (Ca—1397h2). D Durango: E. W. Nelson 4709 (W—— 332763) 5 Bee = 278 (Ca—520602, E—-118777, E—118778, 7 pene (F—600913); Shreve 9138 (ca—731837) ; Waterfall 12 te ay 15376 (Gg). Federal District: Barkley & Rowell & Rowell 769 (Au--170035, Ip, Mi); G. L. Fisher sen, [July 18, 192k] (Vi); Matuda 18875 (12577279) Miranda & Bark 16958 a - Guana- duato: Purpus s.n. [Silao] (Ga--220601)- Hidalgo: V. H. Chase Ta0e Zoe (Ur); 7329 (U 7, ene 9287 (E—L8776, ei ee Rose & r 662 CF (W—li50198). Jalisco: R. MeVaugh 20652 (Mi). Méx- oe sar asiaaTosee (E—839806, F—1,8)993), sor (W—1003437); G. Le Fisher 316 (E—91793, F—555008, W—1207h07); Hernandez Corzo & Guamin sen. [9.VIII.1963] (Ip); j Lundell & Lundell 12351 (Rf, Rf); Matuda 26721 (z); Orcutt 3628 (E—118910, F—281921, W--567395)3 seray Jar ae pooemed : Arséne s.n. [Loma Sta. Maria aris E—8L,8)5, W—1003568), s.n. {Jaripeo, July 13, 1911 aie Bg i ant Koeppen, & Dtis bh3 (S). Nuevo : nm Ars Ss Mat iar | (685A), 3 178] Moir a Mueller see 2 Mueller 308 1967 Moldenke, Notes on Priva 397 12272 (Rf); R. M. King 1967 (N); E. W. Nelson 1822 (W—566114). a Arséne 96 (W—-1003559), ‘1195 Cr 1003556), 1340 (W— 567), Tol (H—1003560), 10207 (W--1003311); Nicolas & Arséne $280 (E—-SL)853, W--1003565); Purpus 2569 (Co-—13972), 352 (Ca= 1397h0); Ugent & Flores C. 2541 (Ws). Querétaro: Waterfall 1129 (Gg). San Luis Potosi: G. L. — 3313 Rah ON F. W. Gould 10810 (Mi); Parry & & Palmer 713 “in part (E—118783, E— 11876), W—L3503); Purpus 5518 Dg ete E—11880), W—61,002), 5519 egal J. Rzedowski 3800 (Ip), 49hi2 ap = (Ip). Sonora: H. S. Ge ay (F=-7 766070) ; Ss. S. Whi 35 (Mi). Ta=- maulipas: | Stanford aig Lauber, & Taylor 2063 2063 eats Hi—200919, W-—-2216858), 267) 267 (pa he3692, W—2216905); Stanford tet Retherford, & Northcraft 772 772 (Ca—713798). Vera Cruz: Bourgeau _ (F—26560). State undetermined: Foreman 2h Cate Herb. "Bernhardt 8.n, (E—11)056). GUATEMALA: Baja Verapaz: Tttrckheim im 11.2331 (W— 1323154). El Petén: J. A. Steyermark 16287 (N). (N). Sacatepéquez: Breedlove 11410 (Ac). Department undetermined: Tondus 787 [Brefias] (W—-108],758). CULTIVATED: Germany: Hort. Bot. Gvttingen s 3eNe [1795] (E—118773, E--11877h); — J Fe de “Young s.n. (Ws). MOUN- TED ILLUSTRATIONS: Kobuski drawing 1s o2ehee), ech wien . PRIVA MEYERI Jaub. & Spach Additional bibliography: Bocq., Adansonia 3: [Rev. Verbenac mt 212. 1863; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: - 189k; : 98 & 110—— Fl. Madag. 17h: 29. “aaron Moldenke, Résumé 143, oat "us, 151, 15h, 310, & Bb 1959; Moldenke, Phytologia 1: 3h7. 1 odin descri this Lan G t on a river bank. Peter found it growing at altitudes of 520 to 1500 meters. The corollas are described as "light-blue" as "white" on Peter 15509. Wart-like projections are found on the cocci of Barbosa 751, Jun Junod 277, and Torre 2425. The last-mentioned collection matches well the C. E. Moss 18268 specimen in the herbarium of the British Museum. The species has been collected in anthesis from January to April, as well as in August and November, and in fruit in March, April, and August. The L. E, Codd 52h9 and "Ee Sousa 5, di distributed as this species, are actually P. cordifolia var. abyssinica (Jaub. & Spach) M Bo Additional & emended citati TANGANYIKA: i: peice 12501 [0. III .156] (B), 19509 [0.1V.1h] (3), 51780 Pagel (B). PORTUGUESE EAST Peete. eee argues: gunod 277 oa) odin 12s (— 398 PHYTOLOGIA Vol. 1h, no. 7 206 3092) 5 Torre 2h (Ul). Manica e Safala: Barbosa 1151 (U1); ae 154, in p ie et Ul). SOUTH AFRICA: Cape of Good Hope: Bolus 306 (F— 9662); Drége a (E—~118803—cotype); MacOwan s.n. [Bosch- berg] (F—L6279) . ~jatal: E. E. Galpin 1782 (N). PRIVA MEYERI var. MADAGASCARIENSIS Moldenke Additional bibliography: Moldenke, Phytologia 5: 11]. ge Moldenke in Humbert, Fl. Madag, 17): *29—30. 1986; Moldenke, sumé 156 & 468. 1959. PRIVA Stipe # eae tio graphy: set Salisb., Ind. Kew. Suppl. 10: 28h. 19 19,7; Moldenke Spy telog da 5: 111. 1954; Moldenke, pésumé & 468. 1959, PRIVA PERUVIANA Moldenk ional biblio ogierha Hill & Salisb., Ind. - 10: Additi Kew 184. 197; Moldenke, Phytologia St 11. 195k; Slee co 8h & 468. 1959, Additional citations: PERU: Amazonas: A. Mathews 3158 (G— isotype) a3 PRIVA PORTORICENSIS Urb. Bibliography: Urb., Symb. Ant. h: 534. 1903; Prain, Ind. Kew. Suppl, 5: 207. 1921; Britton & P. Wils., Seient. Surv. Porto Rico 6: lsh & 145. 1925; Moldenke in Fedde, Repert. Spec. Nov. 1: 19-21. 1936; Moldenke, Résumé 58, 19, & 468. 1959. PRIVA SOCOTRANA 26 nke tional fais graphy: Hill & Salisb Suppl. 10: coed ant pees, Phytologia 5: lll. 19eh3 fasiaaies Résumé 959 NOTES ON NEW AND NOTEWORTHY PLANTS. XLVII Harold N. Moldenke CALLICARPA BICOLOR var. BERMEJOSI Moldcenke, var. Haec Ve varietas a forma typica speciei recedit laminis foliorum oblongo-lanceolatis vel oblongis 13-18 om, longis 3—5 cm. la- pre nes: te margine subintegris tis, ad apicem vel obscure sima: a This variety differs from the typical form of the species in having its leaf-blades oblong-lanceolate or oblong, _— cm. long, 3—5 om, wide, conspicuously long-acuminate at both ends, and subentire or obs simate-crenate . te along the margins The type of the variety was collected by J. Bermejos —- in 1967 Moldenke, New and noteworthy plants 399 whose honor it is named — at Bulalac _t Mindoro, Philippine August or deptinber 1906, and is number 1534 in a herbarium of the Philippine Bureau of Science, deposited in the United States National Herbarium at Washington. ee BICOLOR var. SUBINTEGRIFOLIA Moldenke, var. nov. Haec varietas a forma typica speciei laminis foliorum margine integris vel subintegris recedit. This variety differs from the typical form of the species in having the margins of its leaf~blades entire or subentire. ee type of the variety was collected by Jacin sniper at Pan- L Province, Luzon, Philippine Islands, o 27, 1958, isd is mmber 39680 4 the Philippine Nations): Rerbarius deposited in the United States National Herbarium at Washington. CALLICARPA FORMOSANA f. PARVIFOLIA Moldenke, f. nov asec forma a forma typica speciei laminis foliorun uniforme 1— 3 em. longis 7—15 m, Jatis recedit, This form differs from the typical form of the species in having the mature leaf-blades (on fruiting plants) uniformly o 1—-3 om. long and 7-15 mm. wide. The a of the form was collected by L. Escritor at a Bay, e province of Isabela, Luzon, Philipp June, i913, and is mmber 21155" in the eechae ane the red Seal pine Bureau of pe are es in the United states National Herbarium at Washingto LANTANA RUGULOSA var. PARVIPEDUNCULATA Moldenke, var. nov. Haec varietas a forma typica speciei pedunculis brevissimis recedit. This variety differs from the pence form of the species in The type o variety eas noihactat aty aoete Velarde Nuflez (no. 205) at 22150 5 aaa lagi he province of Caja- bamba, Cajamarc on November 16 1350, and is deposited in the United Static abs ee Herbarium at ¥ hington. SPHENODESME PIERREI var. THAILANDICA Moldenke, var. nov. edi variety differs from the typical form of the species in having the lower leaf-surface more or less densely pubescent or Pilosulous The type of the variety was collected by Kai Larsen (o- 8328) in scrub at Tha Ki Len, Thailand, ace wagngd pees Bape vy deposited in the H. N. ‘Moldenke Herbariun ™ sey. The collector notes that the plant is "very common in s all over the district." SYNGONANTHUS FROESII Moldenke, Herba anma caulescens mgr A, ea eiaciietion erectis 7—9 cm. 4,00 PHYTOLOGIA Vol. lk, no. 7 longis eee peor primaria terminali; vaginis dense hirsu- tulis; acillimis 3—7 cm. longis sparse villosulis; capitulis hemi, mead a albis )—5 mm, Anmal ender, dark-brown, 7-~9 am. long, hirsutulous or eventually slahesoet: toward rosulate, numerous, linear, erect or ascending, about 1 cm. pe . d— Pilosulous, often twisted * partly curled in drying; stem leaves very numerous, imbricate, ascending-erect, linear—oblong, 7--9 m. long, sharply acute at the snets sessile and obtuse or rounded at the base, densely spreading-hirsutulous on both surfaces especial- ly along’ the margins; inflorescence umbellate terminal e Wi pex rous . The type of this distinctive species was collected by Ricardo de Lemos Frées (no. 2990) — 2. whose honor it is named — in sandy soil in a wet area of the "campos gerais" at Conceogtio do » Brazil, on June 25, 1953, and is deposited in the H, N. Moldenke H Herbariun at Plainfield, New Jersey TEIJSMANNIODENDRON BOGORIENSE var. PENTAPHYLLUM Moldenke, var. nov. Haec varietas a forma typica speciei foliolis 5 recedit. This variety differs from the typical form of the species in The type of the variety was collected by A. Cuadra [North Bor- neo Fores De ent A.877) on rolling land, Compt 13, at Bili-Senit — Reserve, Elopura Forest District, S. ’ c andakan on August 5, 1948, and is deposited in the United States Settenas Herbarium at apne agi The collector describes the as a tree, 30 feet tall, 3 feet 6 inches = trunk diameter, with pale-violet flowers, called medang partes" in Malay. The wood is used for sawed timbers ADDITIONAL NOTES ON THE GENUS LIPPIA. II Harold N. Moldenke Houst. ; Additional & emended bibliography: N. J. Anderss., Vet, Akad. 1967 Moldenke, Notes on Lippia 401 Handl. “sacs 1853: 198-199. 185k; = cine bere te oe 80. 1859; » Adansonia 2: 89, 107, a, 12h, 12 5, i, in, 143, 147, 119, Neb -ase, 158, . "885 (2862 2), 296 (1863), and 3: 178, 180, oe 2h3-— & ee = A. S. Hitche., Ann. Rep. Mo. Bot. Gard. : 117. 1893; Robinson & Greenm., Am. Journ. Sci. 150 [ser. 3, 50]: Te & 162. 1895; B. L. Robinson, Proc. Am. Acad. 38: 196. 1902; K , List Pl. For- & 820. 191); 593, fie: 385a. 1911; A. Stewart, Proc. Calif. Acad. Sci., ser. l, Fae 3—134. 1911; P er haart Pl. Méd. Guin. 45. 1912; J. H. Hol- ao oe +. Ser. 9 [Useful Pl. ge 3): 517—518. cH Gamble, Fl. Madras 1085 & 1088--1089. 192); L. H. Bailey, Man. Cult. Pl., ed. 1, pr. 1, 628, 630, 826, & & 8h2 (192k) and pr. rvine 1930; Tu, elon amie Dict., abridg. ed., 460. 1933; nemeelns Bol. mes Nac . 13: 239 & 280. 19375 Dalz., Useful Pl. W. Trop. A Pass hs6. 1 1937 Fletcher, Kew bul. Misc. Inf. 1938: 408 & i0. 1938; Rabate, Journ. Pharm. & Chim. 28: 137—hh2. 1938; 226, & 510. “1957; E. W. Lathrop, Univ. Kans. Sci. - 39: 1% & 172. ca J. Hut pies a vig Flow Pl., ed. 2, 2: 395. 19593 i sm 1929=1 ie. Pers hath, Bot. Surv n te. ne “ 2 085) ; roff Coll. "361. 1961; F. R. Nat. 12. 19635 Quezel & Santa, Nouv. Thieret, Castanea 28: 170. 19635, San Santapau & India 5: 107. 1963; 7 . Bot. Surv. India 5: 26h. 963; Rao, Aggarwal, & Mukherjee, Bei Bot. Surv. India 5: 315. 1963; Huber, He & Meikle in Hutchinson & Dalz., Fl. W. Trop. Afr., ores 2; 2: b32” 1963; Meikle in Hutchinson & Dalz., aL. 8 "ed. 2, 2: 437. 19633 E. E. Lord, Shrubs & Trees Aus- 402 PHYTOLOGIA Vol. 1h, no. 7 Gard., rev. — 245. 1964; T. A. Rao, Bull. Bot. Surv. In- dia aia t 5h. 196k; & Sastry, Bull. Bot. Surv. India 6: 281. 1964; Singh gn Rog Bull. Bot. Surv. India 6: 1,0. 196; Marg. ala Introd. Flow. Pl. W. Afr 163 & 164, fig. h3e. 1965; Un- Univ. Kans. Sci. Bull. 26, h6, 48, 63. i968; Liogier, fnodore 67: 39. 1 108s, 3 5. E. Moo 26. 1965; Moldenke, Biol. A pala 46: 8h6h a 965), Tt "3296 “365), ne osta Ri Phan. Moore, Surv. Vasc. Pl. Lambton Co. - 1966; Fournier, sr ye Tree Fam, Costa Ric. 13. 1966; Moldenke, seat Suppl. 1): 1, & Do pdm, Assoc. Stud, Tax. Fl. Afr. Trop. Index 1 1965: “or. 1966; J. Ofmes, Biota 6: 101, 102, & 119. 1966; Rickett, Fis. U. S. 1 (2): 36h, pl. 113. 1966; Anon. Biol. Abstr’. "iT (20): $.89 & $.162. 1966; Moldenke, Phytologia 13: 343—368, 01, l5— h6, & 510 (1966) and Wj: 217, 325-326, & 3hb. 1967; Shinn, Univ. Kans. Ly Bull. \ : 790 80 8. 1967. Y Ol blOLOgical aApstracts s tne genus Lippia in the Phytolaccaceae in Biol. Abstr. ie: 8 Bh6ls aa for ained reason! Chittenden (1951) states that the genu: contains " —_ 90 species, mostly shrubs", Shinn ( soil records of this. genus, as well as from "Phyla strigosa" and ater geet (1965) points out that in Lippia and in Lantana the o- composed of only one carpel and the ; cross-wall pparently 2 bed the the common wall of two joined ek 92.04 such a wall would be at To be added to the list of dantales's Lippia baillonia Darlington & Wylie = Dipeted juncea (Gill. & Hook.) Miers Lippia canescens L. = Mentha alopecuroides Hull, Lamiac Lippia dubia Trev. = Phyla scaberrima (A. L. Juss.) Molde ie LIPPIA ABYSSINICA (Otto & Dietr.) Cuf. Additional & emended bibliography: Bocq., Adansonia 3: [Rev. 3 ax. Fl. Afr. Trop. Index 1965: 67. 1966. Dalziel (1937) ei "The ei leaves are odorous and are in- fused sa rr » Which diated by as well as the » and c tea. This is widely used as a sudorific se and laxative for colic, etc., in much same way 45 1967 Moldenke, Notes on Lippia 403 Ocimum; a common children's remedy for fever and constipation, accompanied by a light purge, useful also for common colds and chest complaints. The leaves are sometimes mixed with those of Sesamum or cooked along with other food to produce the same ef- fects. They are also a common ingredient in prescriptions for venereal disease. In Gold Coast the leaves boiled with palm-nu are given to women in partuition to assist delivery of the after- birth (Irvine). In French Guinea the decoction of the leaves is also used as a fumigation and bath, and as a hot application locally applied for ear troubles (Pobéguin.....). In Gambia hives are fumed with smoke from this herb to attract bees (Dawe)! He records many vernacular names: "afurati", "bahé~bahé", "bahé", diohuli", "efinrin-gogara", "fasau", "fetfetti", "borom-borom", "Gambian tea bush", "gane ba", "guilel guéri", "kani ba", "Kingkilli ba", Mimbo", ‘mbalhat", "mbormbor", "ngasu", "ngasuru", "nydna" . "sgaa=numum" [=the Ocimum viride of the dry open plains], and "sisiling hyamo". LIPPIA ABYSSINICA var. PUBESCENS (Moldenke) Moldenke Additional bibliography: Moldenke, Phytologia 13: 3h. 1966; Anon., Assoc, Etud. Tax. Fl. Afr. Trop. Index L965: 67. 1966. LIPPIA ALBA (Mill.) N. E. Br. dditional synonymy: Lippia gemuina H.B.K. apud Sampaio, Bol. Mus. Nac, Rio Jan. 13: 239, sphalm. alm. 1937. Fl. Madras 1088-1089. ingens rma AS wet as Nac. Jan. 13: 239 & 280. 1937; Le Cointe Arv. & Pl, Uteis, ed. 2, [Brasiliana, ser. 5, 251: 738 180 & twas. Aoki: Sebastine, Bull. Bot. ot. Surv. India 1: 5. 19595 Sentapan u & Wagh, Bull. Bote Surv. ndia 5: 107. 1963; Singh & Bull. Bot. Surv. India 6: 140. 196 bre Phytologia 3 Sie 36. 1966; Moldenke, Résumé The flowers on Saghategut 6035 are described as "white", but those on Sagdstegui & ve 5755 were "violet" and those on Woolston 818 were oe ee L. Se Smith describes the plant as a "shrub with low spreading branches at times almost eeu the flowers opening pale-violet with a large white c a, but owish in the throat, later becaning more or less alee violet mien His Arg.2 originally from Entre Rios, Argent. The H. C. elnacicr ha hoettiaa, | August 3, 1919), distributed as this species, is actually L. graveolens H.B.K., while Gaumer 830 is Lantana microcephala A. Rich. Sebentine ae cites | his nos. 817 & 1206 from India. Additional citations: COSTA RICA: Guanacaste: A. R. ered 132) (Rf). CUBA: Las Villas: Elman 16988 = WINDWARD ISLANDS: Miller 9485 (S). & que: Kimber wster, = = apices 6035 Loreto: Sag&s- Grenada: We A —< 587 (Ws). PERU: Amazonas: (Ac). LO PHYTOLOGIA Vol. 14, no. 7 tegui & Aldave 5755 (Ac). BRAZIL: Pernambuco: L. S. Smith Bras. Wi, (Ae). BOLIVIA: El Beni: H. H. Rusby 916 (Ws). “PARAGUAY: L. S. Smith Parag.1 th ARGENTINA: Entre Rios: L. S. Smith Arg.32 (Ac). Santa Fé: L. S. Smith ee (Ac). CULTIVATED: Argentina: L. S. Smith Arg.2 Pak Arg.13 (Rf). Paraguay: Woolston 818 (S). LIPPIA AMERT CANA L. Additional & emended bibliography: Bocq., Adansonia 3: [Rev. es 2h. 1863; Moldenke, Phytologia 13: 3i6—-347~ ore esters on Lépez & Sagdstegui 5225 are described a: suas att iaicuan citations: COLOMBIA: Magdalena: H. H. Smith 450 (Ws). PERU: Cajamarca: Lépez & Sag4stegui 5225 (Ac). LIPPIA AMERICANA f. PILOSA Moldenke Additional bibliography: Moldenke, Phytologia 12: 79 & 81-82. 1965. LIPPIA ANGUSTIFOLIA C Additional teerety: Moldenke, Phytologia 13: 347 & 362. BALANSAE Briq. Additional bibliography: Moldenke, Oa ae a 13: 348. 1966. Additional citations: PARAGUAY: u. S. Smith Parag.13 (Rf), Parag.17 (Ac). LIPPIA BOLIVIANA Rus Additional bibliography: Moldenke, Phytologia 13: 348. 1966. Additional citations: BOLIVIA: Cochabamba: M. Bang 979 (Ws— isotype) i LIPPIA BOLIVIANA var. ANGUSTA Moldenke Bibliography: Moldenke, Phytologia 1): Gag end Steinbach describes this plant as a shrub, ant leaves and white fragrant flowers Siseass Uometaiecnine ary Jen fl = at 2575 meters alti itude. Citations: BOLIVIA: Cochabamba: Re F. Steinbach 191 (S—type, me BRACTEOSA (Mart. & Gal.) Moldenke Additional bibliography: Moldenke, Phytologia 13: 349. 1966 Additional citations: MEXICO: Chiapas: Laughlin 659 (Ac). LIPPIA CALLICARPAEFOLIA eet Additional & emended bibliography: » Adansonia 3: [Rev. piper 2hh. ES Koldenke, Pigutlngis “Is: 3h9. 1966. s this plant as a coarse shrub 3m. tall, the Sint oom iy the corollas pinkish-white, then turning yellow finally near. nearly orange, found "occasional" on steep mountain- ke in pine-oak forest on thin soil over limestone. The Lang- 1967 Moldenke, Notes on Lippia 05 lassé 58h, cited below, is from "Cerro Verde" and may therefore have been ieee either in regents: or Michoac4n. The collec- ords rnacular tor rec the name "salvia" — mobs *"Aromati ae Fleurs rouge vif, plates de See Oe rere: rig A onal citations: MEXICO: Mi ni 9 hauela ssé 58h, (Mi); i R. McVaugh 22697 (Mi). LIPPIA CARDIOSTEGIA Bent Additional epituermeins Mohieckns pcb eyeerg 13: ates 1966. The flowers on Breedlove 1496 ar ribed as e~yellow"; itwas collected on a steep rocky nite ag Quercus Additional citations: MEXICO: Chiapas: Breedlove sdlove ilgh6 (Ac). ae CARVIODORA Meikle dditional bibliography: Anon., Kew Bull. Gen. Index 1929-1956, 12. 1959; Moldenke, Phytologia 1%: "550. 1966. LIPPIA CHEVALIERII Moldenke Additional peg oi J. Ge spa in Thiselt.-Dyer, Fl. Trop. Afr. 5: 280. 1900; achiabon me Fl. W. agen hg ed. De 2 (1): 2. 1931; Gait on & Dalz., Trop. Afr. og MOo. tg 2 Bate 19633 ‘ease, me eomnaee ee “350. _istide (1963) says that the L. adoensis of Fl. Trop. 900) and of Fl. West Trop. Afr., ed. 1 (1931) is in part Sota iin ae 8 He describes the plant as an erect aromatic su hrub to about 3 feet tall, the floral nets covered with whitish or in February and October to "December, fruiting in December. He cites the fol- lowing specimens: SENEGAL: Heudelot 103, Monod 8681, Thierry 227. : Brown-Lester , & 28, Dawe 30, , Fox 48, Ingram SDs, Saunders 108 . MATT: Chevalier 67. | PORTUGUES & GUINEA: Santo 2 23b1 & 3552. GUINEA: Caille s.n. [Herb. Saale 147 72], “Herb. “Mission Pharm, A. 0. F. 3/193h, Schnell 711. SIERRA LEONE: “Scott Elliot 1285. NORTHERN NIGERIA: Keay FHI.28012, Meikle 269, Onwudinjoh FHI. 24037. LIPPIA CHIAPASENSIS Loe mets png bibliograpiy: are ey Lg Saag aa ropa 1966. as renga pied) spo feat i atittide, ere Lage ona ai rocky slope at ? ee Addit: ap Poe gp Pape th MEXICO: Chiapas: Breedlove oes (Oe (Mi, r.. LIPPIA CHRYSANTHA Gre aiet tad saa Gerantii Moldenke, ee a ae oh & oe, gga Additional citations: MEXICO: Morelos: L. LIPPIA CONTROVERSA M 5 Additional rey nse 8 Moldenke, Phytologia 13: 351. 1966; Mol 1,06 PHYTOLOGIA Vol. 14, no. 7 denke, Résumé Suppl. 1): 1. 1966. Recent collectors describe this plant as a shrub, a feet tall, with yellow flowers (Keele 34190), flowering and fruiting in December, and growing on dry forested slopes. Additional citations: MEXICO: Colima: Koelz 34190 (Mi). Jal- isco: Koelz 3170 (Mi). ae CULMENICOLA Moldenke Bibliography: Moldenke, Phytologia 13: 5. 1966; Moldenke, Résumé Suppl. 1h: 1. 1966. Citations: MEXICO: Jalisco: R. McVaugh 23036 (Mi—type, Z— isotype, Z—photo of type). LIPPIA DOMINGENSIS Moldenke Additional bibliography: Moldenke, Phytologia 12: 146. 1965; Jiménez, Supl. Cat. Fl. Doming. 1: 217. 1966. LIPPIA FERRUGINEA H.B.K. Additional & emended bibliography: Bocq., Adansonia % [Rev. Verbenac.] 2h. 1863; Moldenke, “Phytologia 13: 353. 1966. LIPPIA FORMOSA T. S. Brandeg. Additional bibliography: Moldenke, Mrtelog.s Da ges — 8 found this plant growing on banks royo and describes the bracts as papery, asoes to tae meee and [on his no. 14536] the Sealine aee yellow 4 n anthesis, lavender in a age, lobes lave Additional citations: MEXICO: Baja ak oelitonetin I. L. Wiggins 14536 (Du—h38h)j)) . LIPPIA GARDNERIANA Schau, : es bibliography: Moldenke, Phytologia 13: 353 & 359+ PIA GEHRTII 3 the flowers adie "pale-violet on the lobes, with a white- ringed yellowish area in the center, later changing to more or less Sale Howes. with a pation are a in ber center, growing in sandy s "s Sabeeetie 4 ruary. ayttitional citations: mAZION S80 Paulo: L. S. Smith Bras.7h onal bibliography: Moldenke Logia 13: 35h. 19665 Moldenke, Résuné Suppl lis 2 1006? ee : LIPPIA GRANDIFOLIA Hochst Additional bibliography: Marg. Mielsen, Introd. Flow. Pl. W. 1967 Moldenke, Notes on Lippia 1,07 164. 1965; Maen Phytologia 13: 35h. 1 « iilsen (1965) s that this species is wk common else- ' than in we tens bgt LIPPIA GRAVEOLENS H.B.K ear eu & emended “bibliography: Bocq. : [Rev Verbenac.] 2h);. 1863; Moldenke, igo a is: eel 385. 19665. Moldenke, Résumé Suppl. 1): 1 & 9. 19 he flowers on Kruckeberg 4807 807" are described as "pale lemon- yellow", while on S. S. White 1938 they were "white". Kruckeberg reports finding the species in : scrub with Cordia and mesquite on limestone slopes Additional citations: TEXAS: Brewster Co.: H. C. Hanson 5.n. {Boguillas, oe 3, 1919] (Ws). Cameron Co.: J. R. Crutchfield 1108 (Ld). MEXICO: Coahuila: S. S. White 1938 (Mi). Nuevo Leén: G. L. Fisher sen. [July Me 192] (Ws); Kruckeberg 1807 (Du— 506122). Vera Cruz: L. S. Smith Mex.36 (Ac). Yucatan: G. F. Gaumer 832 [Field Mus. etn (Ws). LIPPIA HEDERAEFOLIA Mart. & Schau. Additional & emended bibliography: Bocq., nia 3: [Rev. Verbenac.] 2));. 1863; Moldenke, Phytologia 13: 385-356. 1966. LIPPIA HIRSUTA L. f. Additional saree net ae ae poh aepen rd ve 356. 1966. The Herb. F. J. Young s.n., distr buted as s species, is actually Lan Lantana achyranthifoli a it LIPPIA HIRTA (Cham.) Meis Additional perepysbensies Moldenke = L. S. Smith describes s plant meter tall, the corollas beso or less ringed yellowis Additional mepeeaae BRAZIL: Paran&: L. S. Smith Bras.61 (Ac). ie Bes ce 13: 356. eee + subshrub, 0 as with a ret LIPPIA HYPOLEIA Briq. Additional bibliography: Résumé S ~lys1& 1966; Noldenke, a ert 13: 66-387 & 362 (968) and 14: “aN. a shrub, 2 m. tall, this plant as Recent collectors describe seg with a yellow with a pung corollas white or W ; an apples Pepe it has been understory, in the tropic coastal plain in an are tude of only 5 Pua The corollas are descri 4,08 PHYTOLOGIA Vol. 14, no. 7 Roe & Roe 2218 & 230i, and as "white with yellow center" on Roe & Roe 2330. ~~ Additional citations: MEXICO: Hidalgo: Weber & Charette 11939 (Du--98875). Oaxaca: Janzen s.n. (13 January 1964] (Mi). San Luis Potosi: Roe & Roe 2218 (Z), 230k (Rf), 2330 (Ac). Vera Cruz: C. L. Smith 1076 (Ws). - OVATIFOLIA Molde The type of the variety was collected by Keith Roe, Eunice Roe, and Scott Mori (no, 1121) in scrubby Quercus woodlands on a plant as a shrub, 1.5 m. tall, with white and yellow flowers Citations: MEXICO: Chiapas: Roe, Roe, & Wort 121 (appt ps INOPINATA Mol ddi tional bibLiogranty: Jeiekt. Phytologia 13: 349 & 357. 1966, Moldenke, Résumé Suppl. 14: 1. 1966. [Herb. Inst, Miguel Lillo 32855] pahgmmeg LIPPIA soo ht sag (Burm. f.) S Spre pe onal bibliography: Comi - Surv. S. Afr. Mem. 33: td 1962; Moldenke, Be bag ons 13: 8. "1966. Comins (1962) re that this spec st » “He young ts pale-green, tinged reddish-purple, very obscurely |~sided, the leaves dull-creen above, gray-green be- neath, the ve reticulation of the veinlets sunken above ing in the shade of trees, flowering i February, notes that his material was "collected for chelabonth analysis os C. So I. Re "The lao Milne~Redhead & Taylor 1109, distributed as L. javanica, 1967 Moldenke, Notes on Lippia 409 appears to me to be more likely L. ukambensis Vatke. Additional citations: TANGANYIKA: Milne-Redhead & Taylor 10147 (S). SOUTH AFRICA: Transvaal: De Winter 7555 (S). LIPPIA JUNELLIANA (Moldenke) Troncoso 3 " SSonseaati bibliography: Moldenke, Phytologia 13: 358--359. 9 L. S. Smith Sponge Oe this plant as a subshrub, about 60 am. tall, the corollas opening more or less light-lilac with a white e sone ae tp ow in the throat, later becoming light- lilac to light-violet with a slightly darker center, and found it a on the top of a stony rise near a road cu cutting. dditional citations: ARGENTINA: Cérdoba: Stuckert 3909 (N). La ce, Stuckert 17145 (N). Santiago del Estero: ro: Burkart 20347 7 (N). LIPPIA LASIOCALYCINA Additional she iY Fay tol Moldenke, Biol. Abstr. 46: 846). 1965; Moldenke, Phytologia 13: 359. 1966. LIPPIA LASIOCALYCINA var. SAINTHILAIREI Moldenke Addi tional bibliography: Moldenke, Biol. Abstr. 46: 846). 1965; Moldenke, Phytologia 13: 359. 1966. LIPPIA repre ge BK. sav Bods; inawnla Si Fics Additional & emended bi bliogra: a 3: (Rev. Views o] 2h. 1863; Moldenke, Phytologia “Jo: H . ee eT. 1965. LIPPIA LORENTZII Moldenke sige oe ee bibliography: Moldenke, Phytologia 12: 260—262. Woolston describes this plant as a sprawling or erect shrub, 30--80 om. tall, with purple flowers in June, growing on lake shores Addi tional citations: PARAGUAY: Woolston 1263 (S)- LIPPIA LUPULINA Cham. dditional & emended bibliography: Bocq., Adansonia 3: [Rev. Verbenac.] 2. 1863; Moldenke, Phytologia Ts 359 & 360. sgh Additional citations: BRAZIL: Parand: L. S. Smith -60 (Ac). LIPPIA MICHOACANA Moldenke a ogia 13: 5-6. 1966; Moldenke, isotype, Z—photo of type). LIPPIA MICROC nded bi bliography: Bocq., Additional & emend ogra! Verbenac a 2h. Ke ele Phytologia is: Sr. eres .. 1 ke, Résumé Suppl. 1h: 410 PHYTOLOGIA Vol. 14, no. 7 LIPPIA MIC Schau. dditional bibliography: - U. S. Nat. Mus. 25: 96. 188; Liogier, Rhodora 67: 39. 1 1985; Mo Moldenke, Phytologia 13: 361. 1966. Additional citations: CULTIVATED: Trinidad: L. S. Smith Tr.8 (Z). LIPPIA MICROMERA var. HELLERI (Britton) Moldenke Additional bibliography: Liogier, Rhodora 67: 349. 1965; Jiménez, Supl. Cat. Fl. Doming. 1: 217. 1966; Moldenke, Phytolo~ gia 13: "361. Jiménez (1966) “eitan his no. 1025 from the Dominican Republic. LIPPIA MICRO Song Additional & emended bibliography: Bocq., — oan [Rev. Verbenac.] 2h). 1863; Modeniie ; Phytologia 13: 361.1 LIPPIA MODESTA Br Additional iM ioesashey: Moldenke, Phytologia 13: a 1966. Recent collectors describe this plant as a decumbent herb, 5-- O cm. tall, on low, dry, sandy, siial campos, “at alti-~ tudes of 60-80 m , flowering in December. The flowers are de- scribed as "yellow on R. E. Fries 616 and as "yellow and orange" on Woolston 413. The Fries collection cited below was previously incorrectly cited by me as L. trachyphylla Briq., while the Her- ter collection was cited as L. asperrima var. longipedunculata Molde Additional citations: PARAGUAY: Woolston 13 (S, Z). URUGUAY: Herter 2033a [Herb. Herter 50765] (S). ARGENTINA: Salta: R. E. Fries 616 (S). LIPPIA MORONGII Kuntze Additional bibliography: Moldenke, Phytologia 13: 362. 1966. Aoshsinoes citations: URUGUAY: Herter 989 (Herb, Herter 8259h] Ws PPIA MULTIFLORA Moldenke Additional bibliography: J, G. Baker in Thiselt.-Dyer, Fl. bi > Afr. 5: 280. 1900; F. R. Irvine, Pl. Gold Coast lxix, lx, & 263-26. 1930; Hutchinson & Dalz., ri. W. Trop. Afr., ed. 1, 2 (1): 270. 1931; Rabate, Journ. Pharm. & Chim. 28: hago 1938; Rabate, Rev. Bot. Appl. 18 (201) : 260-28. 1938; Palfray, Sebetey, & état, Chimie & Indust. 3: 367--370. 190; ‘hattitte ’ & Dale , Fl > tie. 306. 1963; are « Nielsen, ntrod. Flow. Pl. W.’ Afr. 163 & Ee: fig. h3e. 1965; "Melderks, 362. 1966, ustrations: Meikle in Hutchinson & Dalz., Fl. W. Trop. Afr., ed. 2. 23 ee. 306. 1963; Marg. Nielsen, Introd. Flow. Pl. W. Afr. 1967 Moldenke, Notes on Lippia 412 ve (1963) says that the L. adoensis of the Fl. Trop. Afr. 900) and of the Fl. W. Trop. Afr., ed. 1 (1932) is L. milti- branched inflorescences, growing in savannas. He cites: GUINEA: Herb. Mission Pharm, A. 0. F. 2, Martine 188. SIERRA LEONE: Smythe 121 121. IVORY COAST: Sty Lesieetbere "2066. GHANA: Irvine 1713, Johnson 542, Morton GC. 9551 & 9911, Vigne FH.1813 & 3881. DAHO- MEY: Poisson s.n. NORTHERN NIGERIA: Barter 768, K Keay FHI ae ES 722 & 1231, Noble 40. SOUTHERN N NIGERIA: Foster 133, K FHI.23433, Keay & Jones FHI.1)010, Latilo FHI.27331, Millen i. "Also in Rio Muni, Gabon, Congo, , Congo, Sudan and | Ubangi-Shari". | It has pig el Son nbd in’ anthesis in November and December and in fruit n and August. Irvine ne f3960)" gives L. adoensis Hochst. as a complete synonym n yellow centres, fragrant, id, stalk terminal panicle, nearly "1/2 in. long; fruit (Jan.). Local hives are smoked with this fragrant plant before being placed in trees, to attract settling of bees (M. T. Dawe, Gambia and Fr. Equat. Afr.). It is a scent plant. Root-ashes are occasi used as kitchen salt in Upper Ubangi (Chevalier). The oe . used in tea-like infusions for feverishness, a decoction of t ks and the treatment troubles, as a substitute for Combretum bath fo: Abbé Walker). The like rere colds and as a sudorific, being strong tea, is used for coughs tt . It is also used for piles. A slightly the tend le. fe ed me light purge an ever and constipation, accompani Sie tatne scmetions 42 PHYTOLOGIA Vol. 14, no. 7 mixed with those of Sesamum or cooked with other food to produce oe same effects (Dalziel). A leaf decoction is used in sae a fumigation and bath and as a hot application locally applied pal o- troubles (Pobéguin, Fl. Méd. Guin., p. 5). The leat a common ingredient in remedies for venereal rat leaves, boiled with palm-nmuts, are used by women to assist in the delivery of the after-birth. Like other oil-yielding plants it is used for rhino-pharyngal (i.e. at the back of the throat), mouth, and eye-trouble, especially Stage tient. the leaf-juice being applied in eye-drops or nasal drops as re quired. It forms part of various complex treatments for oer sickness, especi- ally for severe jaundice perp fever') (Kerharo and Vouquet). The medicinal properties of t S plant are mainly due to the pres- "tea bush", "ksuru", "Jaasuruu", and "Pasu", He cites pee ia: S-Re, Irvine 3, Johnson Sli2a & 58h, Kitson 8.n., Moor 208 & 217k, and Vigne ne 1813 & 3889. He gives the distribution of the species as “Waste ground; common on old fams (Savannah Forest); 'the bush of the Savannah! , Senegal to Cameroons and S, Africa." Nielsen (1965) states Shak “it is "fairly common throughout West Africa" among the savanna undershrubs and is called "Gambia bush tea". LIPPIA NIGERIENSIS Moldenke Additional bibliography: e in Hutchinson & Dalz., Fl. W. Trop. noi ed. 2, 2: 437. 19635 eaitecke: Phytologia 13: 362 3636 Meikle (1963) reduces this species to synonymy under L. rugosa A. Chev, LIPPIA OAXACANA Robinson & Greem Additional piace plea Moldenke, | Pigeelogia 13: 363.1 1966. Additional citations: MEXICO: 0 : L. S. Smith Mex. (Z)-. LIPPIA ORIGANOIDES H.B.K eK. Additional synonymy: Lippia opanenelcs H.B.K. apud Boca., Adansonia 3: [Rev. Serbehic «} 2hh. 186 Additional & emended bibliography: » Adansonia 3: [Rev. Verbenac.] 2hls. 1863; Moldenke, Biol. tet 62 86h. 1965; Kol- denke, Phytologia 13: 363. 19 Recent cr barat have found ‘this pec growing in sandy soil and report the vernacular name "“oulan cimmarén" for it. aottitiona citations: COLOMBIA: mp Saravia & Madrinan 9 (Z 2 LIPPIA PALMERI S. Wats. Additional bibliography: Moldenke, Phytologia 13: 36h. 1966» Recent ymggporane have found this’ spec oo ocky » small arroyos, medivne keer face wand in dense brushy porwr They describe the inflorescence as ct bert nt and the 1967 Moldenke, Notes on Lippia 413 plant as a straggly shrub 2 m. tall or less. The flowers are de- scribed as "white to lavender" on Wiggins, Carter, & Ernst 319 and "pale-lavender" on I. L. Wiggins ns 12399. Hastings & Turner found it in a small wash with Olneya tesota, Prosopis juliflora, and Franseria ambrosioides. Wiggins, “Garter, & Ernst found it as- sociated with as non =speors , Bursera microphylla, Colubrina glabra, Cyrtoc dulis, Ferocactus tus rectispinus, Forchemneris watsonii, Poaqente peninsularis » Jatropha cinerea, Lysil candida, Machaerocereus gummosus, Pachycereus thurberi, ae Sa Sebastiania sp., and Wilcoxia striata. Additional citations: MEXICO: Baja ja California: A. Carter 2667, in part (Du—-3),052); D. M. Porter 122 (Du—l51968), 300 (Du 451965); I. L. Wiggins 15399 Rirtaip et 15461 (Du-—l)53233) , 15588 (Du—l532h9); Wiggins, Carter, & Ernst 319 (Du--50L803); Wiggins & & Ernst 576 eas Ae ~ Sonora ra: Hastings & & Turner 6h- 5. (Du—507020). CORONADO ISLAND: 1. L. Wiggins 17489 (ui). LIPPIA PALMERI var. SPICATA Rose Additional bibliography: Moldenees: Phytologia 13: 36h. 1966. Porter describes this plant as oody-based shrub, 3 feet soa growing at the edge of a edit arroyo, flowering in Febru- Additional citations: MEXICO: Baja California: A. © 2667, in part (Du—3L052); D. M. Porter 380 (Du—Li51972) ay. LIPPIA PEARSONI Moldenk Additional sir or Lippia pearsonii Moldenke apud Anon., As=- soc. Etud. Tax. Fl. afr. Trop. Index 1 Index 1965: 67. 1966. Additional bibliography: Moldenke, Phytologia 13: 364. 1966; Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 1965: 67. 1966. LIPPIA PEARSONI var. SESSILIS Moldenke Additional synonymy: Lippia onii var. sessilis Moldenke apud Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 1965: 67. 1966. Additional bibliography: Moldenke, Phytologia 13: 36h. 1966; Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 1965: 67. 1966. eget Siser J. G. Bak ional bibli cerephy: Moldenke, Phytologia 13: 364. 1966; Mone, oes Etud. Tax. Fl. Afr. Trop. Index 1965: 67. 1966. LIPPIA PLICATA var. ACUMINATA (Moldenke) Moldenke Additional bibliography: Moldenke, Phytologia at es a Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 1965: 67. LIPPIA PLICATA var. PARVIFOLIA (Moldenke) Moldenke Additional bibliography: Moldenke, Phytologia aia Pas Anon., Assoc. Etud. Tax. Fl. Afr. Trop. Index 1965: 67. yy PEZTTOLOGIA Vol. 1h, no. 7 LIPPIA POHLIANA Schau, Additional & emended bibliography: Bocq » Adansonia 3: [Rev. Verbenac.] 2). 1863; Moldenke, chptelorte "13: 365. 1966. LIPPIA PRI} Briq. a el babliogrepty: Moldenke, Phytologia 13: 39, 351, & ve. eae this species growing at an altitude of only 83 ddi tional citations: MEXICO: Sinaloa: Breedlove 1572 (Du— 500726) 5 Edw. Palmer 1199 orseses). fananteh Soa ean ete PPIA PSEUDO-THEA (A. St.-Hil. ) Scha Additional & emended biblio eteaert Adansonia 3: [Rev. Verbenac.] 2h. 1863; Molcenke, Seine "12: 365—367~ 1965. LIPPIA P ham, Additional bibliography: Moldenke, Phytologia 13: 366. 1966; Moldenke, Biol, Abstr. 7: 3296. 1966. LIPPIA RECOLLETAE Additional “a spe Moldenke, Phytologia 13: 366. 1966. Woolston describes this plant as an erect aromatic herb, 30— 50 cm. tall, ins soil. g Additional citations: PARAGUAY: L. S. Smith Parag.5 (Rf), Parag.15 (Ac); "odivton.- 1637 (Ss). LIPPIA RECOLLETAE var. Sac ag (Moldenke) Moldenke Additional bibliography: Marfa, Pl. Valle Cochabamb. 2: h2. 1966; Moldenke, Prytol eda 13: 366. ee REHMANNI rv i. - Pearson dditional b: cere: a Phytologia 13: 366—367. 19665 Moldenke, gcore o Ly: 1966. LIPPIA ROSMARINIF LIA Anderss. Additional & ae bi ibliography: N. J. Anderss., Vet. Akad. Stockh. 1853: 198. 1854; N. J. Anderss., Galap. Veg. *80. 1859; Robinson & G . Acad. 190 Calif. Acad. sei., ser h, 1: 133~13h. 1911; Moldenke, Phytolo- gia 12: (Bas aaa ase 5) and lh: 217, 1967. , distributed “folie. ang tributed in herbaria under the designations Lipachaeta sp. and The species has been collected in fruit in Jamary. re is considerable confusion about the polae? pee ee of this species by ersson. It appears tha first proposed by him 4s in Vet. Akad. Handl. sok. "1863: 198, 1967 Moldenke, Notes on Lippia 41s which, however, was not actually issued until 185). The work with = ; 9 a paper not issued until 1859 is cited therein on page 30, c 59 as the earliest that it could have been ai ting th sued} hg fe a?) oats the following specimens not as yet seen by me: Abingdon: Snodgrass & Heller 827 (G). Albemarle: Baur 179 (G), “31 a "12 (G)3 Sno eapiates s & Heller 147 (G), 168 (G), a (G), 251 (G), 280 (G). The Snodgrass & Heller 888, 890, & 896 ee es Pp also cites are var. stewarti M Moldenke. ditional ended citations: GALAPAGOS ISLANDS: Albemarle: Ono 1307 (Ge--159677). Narborough: A. Stewart 3310 (Gg—31213). LIPPIA ROSMARINIFOLIA var. LATIFOLIA M Bibliography: Moldenke, Phytologia ie ar. 1967. At first glance this plan t looks much like L, salicifolia Anderss., but atnede inspection reveals that its leaf-blades are entire and revolute-margined as in typical L. ro folia, but It gh ages cee min Citations: GALAPAGOS ISLANDS: eas A. Stewart 3309 (Ge— 31211—type) . LIPPIA ROSMARINIFOLIA var. pare soe : binson os Am. Acad. 38: raphy 196. 1902; Moldenke, , Pavtalog! a oP ue Robinson (1902) cites Snodgrass & Heller 398 (G), 890 (G), & 896 (G) under L. oe lia iar 35., but comments that the trongly toothed. ec r tga must be se I have not as yet to the resent variety rather than to the spec LIPPIA RUBIGINOSA Schau Addi tional bibliography: Moldenke, Phytologia 12: k45—L7 (1965) and 13: 352. 1966. copier proces ional eeidine raphy: Hutchinso: pss 2 ia — Afr., ed. 1, 2 (1): 270. 93 Meikle in Hu on “pls Trop. 2c ein e; 437. 19633 Moldenke, Phytolo gia 12: Lipokls « f Meikl 1 refers to the original place of publication o this ean a Soc. alrgeiee 8: 191". He places L. ni- re and says that the L. adoensis nymy h rug’ He of the Fl. W. Trop. Afr., wy 1 (1931) 4a in part Le . ar to L, mitiflora Moldenke , notes that the spec is very simil ee a ie cies but has scabrid leaves with prominent 1 on. geriensis Moldenke in syno 416 PEETOLOG-TA Vol. 14, no. 7 he separates them as follows: Upper ga of “cio criti to the touch, the venation obscure; stems subglabrous or sparsely appressed-pubescent; apex 0. floral mesa tete. very uae, shortly mucronate.L. multiflora. ~ surface of leaves rough to the touch, oo venation promin- rugose-reticulate; stems usually dist tinctly appressed- roe cent; apex of floral bractlets acute or cuspidate....seee L. osa. = ee GUINEA: Chevalier 2010. NORTHERN NIGERIA: Meikle 82h, be 9, Hepper 1092, Latilo FHI.3797h, Lely 115 & 582, MacGregor iS Monod 9361 & "9872. SOUTHERN NIGERIA: Hambler 585. BRITISH CAMEROONS: Maitland 16 1697, Hepper 1269, Keay FHI. ” FHI .26522, Latilo & Daramola FHI .28795. ‘The 5; species has been collected ” anthesis from September to anuary . January; in fruit in December and LIPPIA SALT COPOLTA Anderss., Vet. Akad. Handl. Stockh. 1853: 198— 185), ua ee & emended bibliography: N. J. Anderss., Vet. Akad. gras “198-199. 1854; N. J. Anderss., Galap. peste G08. "185955 » L. Robinson, Proc . Am. Acad. 38: 9025 f. Acad. Sci., ser. h, 1: 13h. 1912; oldenke, Prstanaawas 12: hh 4L9—L50. 1965. inger found this plant growing in forests. Robinson (1902) red N There is considerstie Sanbiacor about the place and date of the original publication of this species by cacoentgeg oui It ap- ion and paper not issued until 1859 is cited therein on page 80, indicating — 1859 as the earliest possible date of its actual publica- tion o. ional citations: GALAPAGOS ISLANDS: Charles: Usinger s.ne Lay vob a 1964] (Z). LIPPIA SALVIAEFOLIA Chan Additional synonymy: Lipia saivifolis Cham. ex Bokerm., Stud. 6: bby & bh7, — ung leaf] —— numbers on a leaves of this species, Satay on new Saiicbad, as well as on those of Lantana hypoleuca Briq., in November, December, and January. ~~ citations: BRAZIL: So atlas L. Ss. Smith Bras Bras .70 1967 Moldenke, Notes on Lippia 417 LIPPIA SAVORYI Meikle Additional bibliography: Meikle in Hutchinson & Dalz., Fl. W. Trop. Afr., ed. 2, 2: 437. 1963; Moldenke, Phytologia 13: 367. Meikle (1963) describes this species as an erect woody peren- nial He cites: GHANA: Harris s.n. NORTHERN NIGERIA: Coombe 112, Dal- ziel 177, Gregory 201, Lely 69, 21, 320, & P.4O, McClintock 2hh, Meikle 1059 & 13h & 13kh, p: presumably all in the Kew herbarim. LIPPIA SCAPOSA Briq. Additional bibliography: Moldenke, Phytologia 12: 80. 1965; Moldenke, Biol. Abstr. 7: 3296. 1966. LIPPIA SCHLIEBENI Moldenke = Additional bibliography: Moldenke, Phytologia 13: 34h, 358, & Ted LIPPIA SCHOMBURGKTANA Emended synonymy: Li a “microphylla Benth. ex Bocq., Adanso- nia 3: [Rev. Verbenac.] ps » sy “in syn. 1863 [not L. microphylla Cham., 1832, nor R. A. Phil., Adistonal & eme aa tore Bocq., Adansonia 3: [Rev. benac.] 2). 1863; Moldenke, Pivaaegis “Je: 486—1)88 (1965) and 13: 352 & 354. 1966. LIPPIA ham. SIDOIDES C mole. Additional synonymy: Lippia sicotses sidoides Cham. nia 3: (Rev. Verbenac.] 2. 1863; Moldenke, Phytologia 13: ak sks 1966. LIPPIA STACHYOIDES Cham Additional bibliography: Moldenke, Phytologia 12: 498--500. Additional citations: BRAZIL: S¥o Paulo: G. Eiten 1575 [plant no. 1) (N). i sa ti LIPPIA SUBSTRIGOSA Turc ern biblionaniy; pesca ooh J hg 08 13: 351 & 3%7— 8. oldenke, R : Messe found this plant Sreoing on io “rocky slopes and on auce with Quercus Pinus. Additional citations: MEXICO: Chiapas: Breedlove 9368 (Ws), —_ (Ws). HONDURAS: Moras4n: Williams & Molina R. 23231 (Du— ). LIPPIA SUFFRUTICOSA (Griseb.) Kuntze : Additional bibliography: Moldenke, Phytologia 12: 505—506 418 P WY-T-0 L:O-G:T A Vol. 14, no. 7 (1966) and 13: 345. 1966. LIPPIA TEGULIFERA var Briaq. Additional oe peat bm Moldenke, "seb pmaee 132 368. i Additional citations: BRAZIL: Paran&: S. Smith Bras .62 (Z) « LIPPIA TRISTIS Brigq. — bibliography: Moldenke, Phytologia 13, 12--16, 29, 347, & 362. 1966, TURBINATA Griseb. Additional bibliography: Moldenke, Phytologia 13: 17—26 & 3h5. The Monetti Sete pi Inst. Miguel Lillo 32085], distributed as L, turbinata, is actually f. mento Moldenke, while the H. H. Bartlett ise, — by me in a previous seekalinent of these notes, is a is ac ctually « angustifolia Osten, LIPPIA TURBINATA f. ANGUSTIFOLIA Osten Additional bibliography: Moldenke, Phytologia 13: 21 & 23—2h. The Bartlett specimen cited below was e erroneously cited by me in a previous installment of these notes as typical L. turbinata eb. Gris i mre ge citations: ARGENTINA: Cérdoba: H. H. Bartlett 19237 aE edger f. tape S LIPPIA TURBINATA f. MAGNIFOLIA Moldenke Additional onal biblicgrapty: Hocking, Resenyt A.9: 367. 1965; Moldenke, pate ar da 13: 21 & 2-26, Additional citations: ARGENTINA: Sinn Monetti s.n. [Herb. Inst. ee Lillo 32085] (Du—317600). LIPPIA TURNERIFOLIA Chan, FF Seaman bibliography: Moldenke, Phytologia 13: 343 & 368. L. S. Smith describes this plant as 11/2 feet tall, the flowers opening yellow and later bec reddish-brown in the center, g along roadsides, blooming in November, The Herb. Herter eaeek, “distributed as L. "turnerifolia, is actually L. mo: morongii Kun Tatikional Giteinidas BRAZIL: Paran&: L. S. Smith Bras.66 (Rf). PARAGUAY: L. S. Smith Parag.12 (Rf). Sc ee LIPPIA UKAMBENSIS Vatke Additional bibliography: Moldenke Milne-Redhead & Taylor describe tis tall, ve tems Phytologia ane seo yee this plant as a shrub green, soon turning brown the “ped icels square, the li, deep dull-green above, paler beneath, the veins and veinlet reticulation sunken above ani raised beneath, the bracts and calyx yellow-green, turning green with age, the corolla 1967 Moldenke, Notes on Lippia ig creamy=white with a —— throat, turning black and persisting, and the anthers dull-yell abundant at the edge of clumps of upland rain forests, at 1890 m, altitude, blooming in ant Mater- ial has been misidentified and distributed in es ee as L. ja- vanica (Burm. f. NE» oe citations: TANGANYIKA: Milne-Redhead & Taylor 11019 S)e LIPPIA UMBELLATA Cav iy ae sae Be bibliography: Moldenke, Phytologia 13: 350, 352, The Janzen g.n. [13 Jamary 196] and the Weber & Charette = distributed as L. umbellata, are actually L Tee! ypoleia , While Edw. Palmer 1199 is L. pringlei Briq. eae citations: MEXICO: Federal District: J. Rzedowski 21961 (Mi). Michoacdn: King & Soderstrom 5161 (N). LIPPIA VELUTINA Schau. Additional & emended bibliography: Bocq., Adansonia 3: [Rev 966. 245. 1863; Moldenke, Phytologia 13: 163—~16) & 363. 1 PPIA VERNONIOIDES Additional bdbatnereneys Moldenke, Phytologia 13: 165—~167. Additional citations: BOLIVIA: El Beni: H. H. Rusby 927 (Ws). LIPPIA WHYTEI Moldenke Additional bibliography: Moldenke, Phytologia 13: 170~—171 & 358. 1966; Moldenke, Résumé Suppl. 13: he 1966. PIA WILMSII H. H. W. P Additional bibliography: 7s iabsanke » Phytologia 13: 171-177, 358, & 36h. 1966. LIPPIA WOODII Moldenke Additional bibliography: Moldenke, Phytologia 13: 17 & 176— LIPPIA YUCATANA L stoner: sabhicerentert Moldenke, Phytologia 13: 177--179 & 362. 1966 ADDITIONAL NOTES ON THE GENUS CORNUTIA. IT Harold N. Moldenke Plun,. Additional & emended bibliography: Westm. in Be wpe Tellur,. Habit. Incr. 59. 174; A. L. Juss., Gen. +s ed. 1, 107--108 (1789) and ed, rt 120--121. 1791; “Guill. Enun. MS "Déc. Voy. = Société 210, 1837; a ats Bot. Voy. Sulphur gh ha ad 2 88, y 20, 21 126, ‘dn, 133, 137-18, & cal CUnaey end bx aah 180, 16h, “il, 233— 3—25h, pl. 12, 1863; Rev. Verbenac iB, ng, ia, 123, 125, 436, Sa. 333, 137-139, i, i, *t80, 181, 18h, & 239-234, pl. 12. 1863; Tures. » Bull. Soc. Imp. ate Cali ioe. Verh. Bot. Ver. Brand. 53: 81 [Abhandl. 26). " 12; Cc. Standl., Contrib. U.S. Nat. Herb. 23: 1235, 1251, & 1252. 192h; K. V. 0. Dahlgren, Svensk, Bot. Tidsk. 32: 231. 1938; Ae We ; Ind. Kew. Sopp. 9: 5. 1938; Robledo, Lecc. Bot. 2: 199. 1940; C. i Iundell, Contrib. Univ. Mich. Herb. 8: 61 & 90. 1942; ele hop. td hogsiy, Cat. Estat. Gen. Bot. _ 17: 3. 1956; Angely, Fl. + 1957; Moldenke, Phytologia 5: 76, 478, & 509 (1987), 6 6 58 Cissh. "6: si 2325 & 22° aecee and 6: 509. 19595 Anon., Kew Bull. Gen. Index 1956, 8h. 1 1959; Angely, Liv. Gen. f. fa Abstr. 36: 3141. 1961; Saukup, joe h: 180. 1962; Paneth 20) 30 :106P, Menninger, Flow. trees ucts 83, 313, 316, & sac ag 41h, 1962; Shelford, "Ecol. N. Am. U2 & 559. 1963; Hoc Excerpt. Bot. A6: 533. 1963; Se mipinrst Guide Lit. Flow. Pl, ex. 515, 516, & 1010. sels Soloapey. Bt a 123 & 130. 196k; Ne n Gard, Hawaii 721. "T3655 *yol- a Llogia 12: ra é {3968 and 13: 32h & Sake 1966; Anon. can. fost es - Phan, 1966; Fournier, Imp. Tree Fam, Costa Ce e Westmann (1744) places this gems in his group "Plantae bacca- tae". Dahlgren (1938) places it in the Labiatae. The generic names Midi H » Sambucus Burmm., and Cornutioides L., listed by Angely (1956) as synonyms of Cormtia, actually b in the of Pramna. 5 (1 records the vernac names "toe " and "ulape" from Peru -~ the former being applied also to ~~ Same notes by Turezaninow we are of interest: "Cormtia e prov. Carabobo Venezuelae, Linden No. 1457, eadem videtur cum specimine ejusden collectionis su sub No No 1398 et cum mexicana Gale- ott No 757. Omnes ad Cc. ieniicclien Sch Schauer pertinent. Variat 1967 Moldenke, Notes on Cornutia 421 haec planta panicula descriptione lineari-lanceolata et late ova- ta ata. Habitu proxima sed staminibus omnibus ferti servatur in collectione mexicana Jurgensenii sub No 533. Generis propinqui sed verosimiliter distincti sunt specimina collectionis Sellowianae, No 510, filamentis duobus anantheris, foliis glabris, inflorescentia racemosa pauciflora, floribus multe majoribus cori- aceis et pees sertim calycis lobo inferiore cucullato-gibboso di- vergent Galeotti 757 is C. grandifolia var. purpusi oe ae Jurgensen 533 and Sellow 510 have not as yet been seen by m CORNUTIA COERULEA (Jacq.) Moldenke Additional bibliography: Moldenke, Phytologia 7: 381—382. se One of the original cotypes of this species from the Jacquin herbarium, preserved in the Vienna herbarium, was photographed sc ate Macbride as his type photograph number ional & emended citations: TIVATED: tria: Herb. eae. f. s.n. [Macbride photos 31,306] (W--photo of cotype). Germany: Herb. | Bernhardi s.n. (E—118976). Seat Ponsa ta ae Scheer: & Cham.) S Additional synonymy: Cornutia gr seactelia nye ae Imp. Nat. Mosc. 3% (2): og in ge 1863. eremiae grandifolia (Cham. & Schlecht.) Schau. . Stewart, “Proc. Calif. Acad. Sci., ser. h, 1: 395.1 Additional bibliography: Benth., Bot. Voy. rreg ee 18,6; meee Verh. Bot. Ver. Brand. 53: 8) [ . 26). 19123 A. ’ c. Calif. Acad. Sci., ser lz: 395. 1912; Pa Gs Stand] b. U. Se Nat. Herb. 23: 1251. 192h; E. J. b., 1961; Menninger, Flow. Trees World 283, pl. yh. a N. Am. bh? & $59. 1963; Neal, In Gard. Hawaii, ed. 2, 721. 9 eee Menninger, Flow. Trees World pl. 41h ional i color] 196 Rec reper peers goons® this plant as a shrub, aoe feet t tall, the foliage and in March and September The flowers are dondiiben as “blue” on euuhes 10094, and on Gilly & Hernandez Xolocotzi 172, e" on ae ie Ndark-purple" on Stork 3009, and “bright reddish-violet” on Rod- riguez C C .h27 422 P2TPtTG6:L0 OTA Vol. lk, no. 7 Stewart eee describes the plant as "bushes about 8 feet on of cliffs and on the banks of streams near Wa- fer Bay (No. 318). Further distrib. S. Mex.", but his specimen is the type ale of C. microcalycina var. anomala Moldenke. Menninger (1962) affirms that C. grandifolia can withstand temper- atures as low as 25°F, — "although rather tender and sometimes killed to the ground by frost, Cornutia recovers quickly and comes back from the roots." Shelford (1963) records the vernacular name "cuatro caras". Turczaninow (1863) cites Linden 1398 & 1457 from Venezuela and Galeotti 757 and Jurgensen 533 from Mexico. The first of these, owever, is ( Cc. odorata var var. colombiana Moldenke, the ta is C. mierocalyoton Pavon & | Moldenke, the third is C. grandifolia var. = bis, S, Stork 508, and Fonda oe [Tucurrique, Las Vueltas], ted by m me in my 1936 work as from "Province undetermined", sia now been localized and ele re-cited below. Material of this taxon has been misidentified and distributed in herbaria as C. grandifolia var. quadrangularis grst. & Molden- ke. On the other hand, the Paray "£0, espe as C. grandi- folia, is actually Cc. Teeer oa var. velutina M idenke. aaitlonal & emended citations: MEXICO: Bier Chtaoaa Breedlove 6469 (Z), 10094 (Rf), 10366 (Ac); Gilly & Hernandez Xolocotzi 172 (4); Paray 561 (Ip). Oaxaca: Lietmann 11306 (W—1315007); Mer- tinez & & Calderén 15 (Ca—938170). Vera Cruz: Plunkett 131 (F— Sri); $8 Schiede 81 (E—118976—-cotype). GUATEMALA: EMALA: Alta Verapaz: H. V. Johnson 895 (W—-108)015); Ttrckheim 21,8 (W--13233h0), 7935 Oe ace W—1323335), II 11.400 (W—-860706). El Petén: Contreras 7 (La, 3); C. L. Lundell 16450 (Ld). Izabal: P. C. Standley _ a8 (W--1150L7). “BRITISH HONDURAS: C. S. Brown 13 (F—58539h, Y—13032); Gentle 7968 (Ld, S), 9150 (Ld); Schipp 265 (Ca—39692, E—988623, F—713239, W—1)969],). HONDURAS: Atl4ntdda: A. M. Chicke 23 (F—689825); E. R. Mitchell 117 (F-58055, W— alvoza 809 (F—710562); P. C. Standley 53342 (F— 582699, eae Colon: Johansen 2 2 (F—572610). Cortés: Re- cord & Kuylen H.6 (W—1315362, Y—995: Y—9952). Province undetermined: Carleton 189 [Comagalpa] (E—921010, W—1208),72). EL SALVADOR: San Salvador: Calderén 47 (W—-1151L55); P. C. Standley 22990 (W— 1138699). NICARAGUA: Managua: Maxon Harvey, & Valentine 740k (W—1181115), Rivas: Lévy 1177 (cb). pas "‘Englesing Be 93 (F— 585038, Y—13275). COSTA RICA: Cartago: R. L. Rodriguez C. Li27 eet Mi); Stork 508 (W--1167057), Tee (Mi); Tonduz s.n- [ ,» Las Vueltas) (W—~1323330). Puntarenas: “Pittier & Tonduz on (W--1323327); Tonduz s.n. [Herb, Inst. Physico-geogr - 1967 Moldenke, Notes on Cormutia 423 Nat. Costaric. 9711] (W—1323326). San José: J. A. Echeverri 364 (Ca—-776219) ; C. Hoffmann 117 (Cp), 517 (B, B, B, B); He Pit tier 237 bis (W—1323329); Stork 3009 (Mi). Province undetermin- ed: Kuntze 2201 (F—297678, F—297691). PANAMA: Panam4: P. C. Standley 26662 (W--1217135). CORNUTIA GRANDIFOLIA var. INTERMEDIA Mol Additional tibliceranner Moldenke, ogo 7: 384—385. The flowers are described as "blue" on Breedlove 11399. This th des Additional & emended citations: GUATEMALA: rapaz: heim 935 (W—-1323338) « Escuintla: Seler & Seler 2560 et OOFELT). : Garcia S. 1472 (F--663850). Izabal: H. [. Pittier 8538 (W—1013483); P. Cc. Standley 24398 (W--115000) . ~ Quezaltenango: Holway 822 (W—863011, 1, W—863012). Sacatepéquez: Breedlove 11399 (2); Tejada 243 (W—862378). Santa Rosa: Heyde & Lux 2957 mel W—57683, W--G0107, W--1323333); Lehmann 1638 Pesan. 1ol4: Shannon 135 (W--5768h, W—1323332). Department undeter- re Heyde 79 [337] (W—-28336). HONDURAS: Santa Barbara: Thieme Sul (F—-57h65—isotype, W--2h664L—isotype, W-—1323339— type). NICARAGUA: Managua: Maxon, Harvey, & Valentine 7531 (W-— 118126). COSTA RICA: Alajuela: A. F. Smith 13 [Stork 1,113) (14), 1k [Stork 113] (i). Guanacaste: A. R. Moldenke 1323 (Rf). PA- NAMA: Bocas del Toro: G. P. Cooper r 529 y (F—579205, “Y—12149) . Coclé: R. S. Williams 963 (W=—678336) « Panam: P. C. Standley 25853 (W—-1216677). CORNUTIA GRANDIFOLIA var. NORMALIS (Kuntze) Moldenke Additional bibliography: Moldenke, Phytologia 7: 385—-386- Additional & emended pias COSTA RICA: Cartago: Tonduz n. (Herb, Inst. Physico-geogr. Nat, Costaric. 11507] (W—333918 a 365936, W—57753, eae q--1323337)- Puntarena —— 58) (W--132333h); Ae R. Moldenke 1322 (Rf); H. Pittier s-n- = : Inst. Physinneeear Me fat. Costaric. 11979] (w— 323325) 5 5 ora 5603 4512 (W— : Bocas del Toro: ’ 4512 (W—1323328). PANAMA pee Ee ree & Lasser 16528 (Mi); Gervais 122 (W#—716503) 5 Kuntze 1932 (E— 118973—isotype); Kaxon & sere 6610 (w—1180 803535 ie i. Patties 3702 (W—678770), 6711 (W—716765)5 P. C. Standley 2571 era Chiriquf: Cooper & per & Slater 225 (F—573158, 8 a, Y— ° Panam4: Bartlett & Lasser 16329 (Ki); Maxon & Harvey 6661 (i— —————— 2h PHYTOLOGIA Vol. 1h, no. 7 118002); H. Pittier 617 (W--679772); Riley 120 (W--1319063). oe ISLANDS: San José: C, 0. Erlanson 311 (Mi). CULTIVATED: an Islands: Hutchison 27uh [Ww ahiawa Bot. Gard. 5662] (Ac, CORNUTIA GRANDIFOLIA var. PURPUSI Moldenke Additional synonymy: Hosta longifolia Humb. & Kunth ex Benth., Additional bibliography: Benth., Bot. Voy. Sulphur 15h. 18h6; P. C. Standl., Contrib. U. S. Nat. Herb, 23: 1252. 192h; Woldenke, The flowers are described | as "purple" on Breedlove 6287 and as ee. on J. Ve == 305. Recent collectors refer to the plant as a shrub, | m tall, growing at altitudes of 2200--3600 meters, Flowering in J e and July. The Tessmann ),0)6, distributed as - this pasta G actually Sanango durum B durum Bunting & Duke, in the Addit i nat & emended citations: MEXICO: Chiapas: Breedlove 6287 (2); E. W. —e 3011 (W--233319, W—233320). Vera Cruz: Botteri 86h, (W—-57685), 1091 (E—118988, F--689828, W-—-57686, W-- 242022, na Mohr 323 (W—771906) ; E. W. Nelson 81 (W— 569125) ; Purpus 2647 (Ca—101607—type, E—118975—isotype, F— 343993—isotype, W—8h1773—isotype), U5 (Ca—143290), sn. (Veracruz, 1922] (Ca—02975); J. V. Santos 305k (ui). RNUTIA GRANDIFOLIA var. QUADRANGULARIS rst. & Moldenke Additional bibliography: Moldenke, Phytologia 7: 387. 1961. The Rodrfguez C. 27, distributed as this variety, is act typical C. grandifolia (Schlecht. & Cham.) Schau. Raaiaed citations: COSTA RICA: Cartago: grsted 11200 (W—~— 1315040—type) . CORNUTIA GRANDIFOLIA var. STORKII Moldenk Additional bibliography: Moldenke, hs totucke 7: 387--388. citations: COSTA RICA: San José: . Standley 313k (w-1206860), Stork 1743 (F--598926—type), 6 OF (FSS CORNUTIA JAMAICENSIS Moldenke Additional biblio erapty: Mold aad Prvtologia 7: 388. 1961. The Proctor 19689, distributed as this species and 30 cited by LATIFOLIA (H.B.K.) Moldenke Additional & emended bibliography: C. L. Lundell, Contrib. 1967 Moldenke, Notes on Cornmutia 425 ip Mich. Herb. 8: 61. 1942; Moldenke, Phytologia 7: 388-—389. gegen collectors describe this plant as a shrub or small tree, , the stem 12 cm. in diameter, the flowers with a r e flowers are des cribed as Hievaicies tina? on Gilly & Hernandez Xolocotzi 109, "grayish-blue" on Gilly & & Hernandez Xolocotzi 263 263, "purplish" on Matuda 3398, and "pale purple-blue" on R. M. King 992. The Set ieraneteer collector notes that it is a . Neommon tree growing in sandy loam in open ee in Vera Cruz. A vernacu- bik inne teported bar it-aé Mex e", Additional & emended citations: MEXICO: Campeche: Sauer & Gade 3184 (Ws); Steere 171 (F—668594). Chiapas: Matuda 3007 (Mi). Tabasco: G Gilly & lly & Hernandez Xolocotzi 109 (Mi), 263 (iti); | Matuda 3398 (Mi, Rf). Vera Cruz: R. M. King 992 (Mi); Orcutt 6330 30 (Ww 1266592) . Yucatén: Degener r & Degener 794 (N, W, Z)3 G. Fe Gau- mer 822 (E—~118983, 7 G08, W—268379)5 L apemeis & Lundell 7888 dalgo 54 (E--1067851, F—713088). BRITISH Pacis Gentle 16 (F-96392), 186 (F—662519, W—1585899), sen. [Lundell 11 476] (F— 683520); W. Ce M Meyer 38 (F—713307) 5 Winzerling III.2 III.2 (W-1266652). CORNUTIA LATIFOLIA f. ALBA Moldenke Additional bibliography: Moldenke, Phytologia 7: 389. 1961; Langman, Select. Guide Lit. Flow. Pl. Mex. 516. 196h,. CORNUTIA LILACINA Moldenke Additional synonymy: Cornutia liliacea Moldenke, Résumé Suppl. 6: 9, in syn. 1963. Adai tional bibliography: Moldenke, Phytologia 5: 95 (1954) and 7: 389—-390. 1961; Moldenke, Résumé Suppl. 6: 1963. Recent a io describe this plant as a shrub, I n, tall, g in forests and on creek banks, blooming also in July. The flowers are described as as “purple” on Williams & Molina R. 10100, "blue" on Williams & Molina Re 1027h, and "violet" on i Molina R - 83. The Molina R. 12981, ~“Gistributed as C. var. velutina Moldenke. Additional & emended citations: GUATEMALA: Zacapa: Co Ce 6383 (E11897h—type, W—579655—isotype) » HONDURAS: El cetao: Wild: & Molina R. 10274 paaggedl: Mi, Rf). Morazdn: mobi R. 83 (Gen-T52 70, “Ry; Williams & Molina R. 10100, in part 729291, Vi). Yoro: J» Be Edwards P.6u8 (Ca——522683, F ais, W—1588609). EL SALVADOR: La Libertad: Calderén 823 (W—1151780) . CORNUTIA LILACINA var. VELUTINA M Additional bibliography: rice eaediige 53 95 (195k), 7: lilacina, is actually 4,26 PRY 0:10 67% Vol. 14, no. 7 390 (1961), and 1): 422. fruiting in S ber. Molina R. reports it "frequent in wet matorrales", P, H. Allen calls it aprile eee and Allen & Severen refer to it as Prope i = t. Salva- El dor. The flowers are described as "blue" on Allen Allen & ‘Severe 6923 and as Ndeop-vialet* on P, aa a hr Throug vn error collection of this vari- ety is cited in Phytologia te 390 9 (ase) as "0. 38s" - Material has been misidentified and dis herbaria as C. grandi- folia (Schlecht. & Cham.) a — as cps C. lilacina na Mol- denke. Additional & emended citations: HONDURAS: Comayagua: wards P.385 (Ca—522721~-isotype, F--713810—isotype, + sta isotype). Lempira: Molina R. 12981 (W—2l,5627). Moraz4n: Wil- liams & Molina R. 10100, in part (Rf). Santa Rosa: Paray 560 (Ip). EL SALVADOR: Sonsonate: P. H. Allen 7073 (Rf); Allen & Severen 6923 (Rf). CORNUTIA MICROCALYCINA Pavon & Moldenke Additional & emended bibliography: Ba Je LOE ne Ind. oi Suppl. 10: 61 & 11). 1947; Moldenke, .G ie te ot Moldenke, Phytologia 7: $90-301 * (361) een 1h: fe & The oie on I. L. Wiggins 1092) are described as "purple", and the plant is said to grow " grow "over 6 m. tall", fruiting in Octo- — The Gilmartin 597, distribu buted under this name, is actually a ecvlacte & Tacs: nke. nded citations: COLOMBIA: amen, Beat 5094 ig ae ae eapnc Chimborazo: Rimbach sea Y=~22810). El Oro: L. Wiggins 1092) emer ba - Guayas: G 7 21a) PERU: Junin: Killip & Smith 26133 ial 1460129) . CORNUTIA MICROCALYCINA var. ete Additional bibliography: A arp aa A. » Proc. Calif. ere h, us 395. 1912; Moldenke, ghnsired ep 7: 391 (ise) and a1 Sond (1912) cites this taxon as "C. grandifolia (Cham. & Schlecht.) Schau." oy mended citations: PANAMA: Panam4: Goldman 1896 (W--71615h) » CORNUTIA MICROCALYCINA var. “hep catotngge Moldenke Bg wwary bibliography: » Phytologia 7: 391-392. Recent collectors describe this Plant as an aromatic bush, half~shrub, or shrub, 2—5 nm. tall, growing in virgin forests and seasonal orests » in March, nd fruiting in August. The corolla is described as nyiolet—biue" on 1967 Moldenke, Notes on Cormutia 427 Harling 4566, "violet" on Jativa & Epling 470, and "royal blue with white hairs" on Cazalet & Pennington 5259. Material has been misidentified and distributed in herbaria as C. microcaly- cina Pavon & Moldenke and as a member of the Labiatae. ti & emended citations: PANAMA: Canal Zone: Johansen 41 (W--1167493); H. Pittier 5690 (W--715977). COLOMBIA: Antioquia: Metcalf & Cuatrecasas 3 30194 (W—1833442). Boyacd&: Lawrance 764 aes 2428431) ; ag 4677 (S); sative & -& Epling U2 (La), 470 ec ee ee), ear Guayast As A. S. Hitchcock 20492 ca . Los Rfos: Asplund 5504 eae Seam hl). iaiatas coubaiat & Pennington 5259 (N, W—-2405728). CORNUTIA OBOVATA Urb. Additional Goahaganten Moldenke, Phytologia 5: 317 (1955) and 7: 392-393. 1961. mended citations: PUERTO RICO: Sintenis 2022 (E--11898 2—— isotype, F—-79572--isotype, We-1,03508—isotype) « CORMUEZA ODORATA Bee seit & Endl. a: dditional bibliography: Mol lie. "Phyto logia 5: 152 sia a 7: 393. ged prree, Biota ]: 180. 2 1962; Moldenke, Phytolo 32h (1966) and 1h: 421 & 422. Soukup (1962) reports the additional vernacular names "huitoto’, "toe", and "ulpe! for this species. The type specimen, Poeppig 2448, in the Delessert Herbarium at the Conservatoire et Jardin Botaniques in Geneva, was photographed there by Macbride as his type photograph number 7885. The Asplund 250 and Cazalet & Pennington 5259 5203 distributed as this species, The are ac tually C. 1 microcalycina var. pulverulenta Moldenke. thdiex Bente States National Herbarium deack of Asplund 8935 is actually Aegiphila elegans Moldenke. Additional & emended citations: PERU: Loreto: Klug 39 (W#— 1155396), 662 (F—630012, W—1h55661), 2159 (e-10L0L8, F—668862, pipmtigebat = $= 2448 [Macbride photos 7885] (F—645726—photo f type, W—photo of type); Ll. Williams 8223 (F-62337). San Martin: Klug 00h (W—1,58127).— CORNUTIA ODORATA var. CALVESCENS Moldenke Additional bibliography: Moldenke, Phytologia 7: 39h. er ‘ blades fimiy send con slat this plant feed he m. seg 1B? —e ae firmly memb , dull-green th raised venation, is: ie fruiting-rachis (Braet mgs the fruit dul blue-lavender Additional citations: COLOMBIA: Caldas: Sneidern 5232 (ii), 5432 (W—-2056616). VENEZUELA: Miranda: J. A- S Steyermark 9005) (Z). 428 PHYTOLOGIA Vol. 14, no. 7 CORNUTIA ODORATA var. COLOMBIANA Moldenke Additional bi biscereniee motdenles Phytologia 7: 394—395 98h) and 1): 421 & 22 The type imen of cae — Triana 2068, in the herbarium of the Naturhistorisches Museum in Vienna, was was photographed there by Macbride as his type photograph — eee The plant has been Feehan in anthesis also in tional & emended citations: COLOMBIA; Bolivar: Curran 160 pres ere Cundinamarca: Mutis 873 (F—712616, W—1562702);__ Triana 2068 [Macbride photos 34305] (W--photo of type). Tolima Juzepezuk 6867 (W196); F. C. Lehmann B.T,1108 (#_-$59867)5 F. W. Pennell 3371 (W--10)2802). ssp oF PUBESCENS Gaertn. f. ional & pares bibliography: Bocq., Adansonia 2: 126 cage) and 3: 137 & 23h, pl. 12. 1863; neargp Rev. Verbenac. 137, & 23k, pl. 12. 1863; Moldenke, Phytologia 7: 395. 1961. CORNUTIA TA Ls Additional bibliography: Bocq., Adansonia 3: [Rev. Verbenac.] 23k. 1863; Griseb., Cat. Pl. Cub. 216. 1866; P.C. a ik Con- Hill, Ind. Suppl. 9: 5. 1938; K. V. 0. Dahlgren, Svensk. Bot. Ti es 32 232. 1938; Jiménez, List Nom. Vernac. 8. 1961; Moldenke, prytologia 7 95—398. 1961. Recent collectors describe this plant as a shrub or tree, 2-- 10 m. tall, the trunk 20 cm. in ro eet the leaves aromatic, and the fruit black when sega Hd ng in dry woodlands, low % woods on headlands, Coperni cohen savannas on gray soil, and the edges of thickets in alluvial soil, at 420 meters alti- tude. The flowers are described as "blue" on J. S. Beard 502, "lavender" on Webster, Ellis, & Miller 9079, and Wbluish-purple" on G. L. Webster er 3877. Dahlgren (1938) places this taxon in the Labiatae. Jiménez (1961) records the vernacular name "palo vidrio" for it. The & Hernandez Xolocotzi 109, distributed under this name, is actually Cc. aetifolia (H.E.K.) Moldenke, while Lawrance 76 is C. microc rocalycina var. pulve rulenta Moldenke. Additional & manidaet citations: CUBA: Oriente: Palmer, & Palmer 193 (W—l,03120); Shafer 3490 (F—286225, — STIL. 3988 (F—286328, W—-697615), 8303 ) (W-696505); G. L. Webster 2877 (Qi); Cc. Wright 3178 (E—11898h). HISPANIOLA: Dominiean Republic Abbott 152 (W—-1076623), 2558 (W--115721), 2756 (W——-1147222), 2692 (W—114733h); Eggers 1538 (W~13233L1); Faris 85 (W—-101,91,0l)5 Fuertes 102 (E—706687, F—385116, W--658193); Raunkiaer 1119 (W— 1110135); Sauer 20L6 (Ws); Wright, Parry, & Brumel 222 mel 222 (W— 57687), 357 (F—575086). PUERTO RICO: E Eggers 26 (W——115932h), 626 (W—1323345). WINDWARD ISLANDS: Grenada: Eggers 6296 (ii— 1967 Moldenke, Notes on Cormtia 429 1323344). Martinique: A. Duss 1965 (W—8L97h1); R. F. Finch 28 (Mi); Hahn 899 (Ca—332)87, W--57688), s.n. [1870] (F—367191); Kohaut s.n en. . [Sieber FI. Mart. 158] (E——118980) ; Stehlé & Stehlé Lu7h (W—2h5 3688) , 5937 (W—253685), 6141 (W—2)536h1), 5875 (W- 2153687) . St. Lucia: J. S. Beard 502 (Ca--797552). LEEWARD IS- : Dominica: Stehlé ¢ & . Stehlé | 643 (W--2)53686). Guadeloupe: A. Duss 238), (W—8L9815); H. Stehlé 2759 (W--2453706); Webster, Ellis, | & Miller 9079 (Mi). CULTIVATED: Trinidad: W. E. Broa s.n, s.n, (Trin. Bot. Gard. Herb. 1376] (W—132332); Trin. Bot. Gard. Herb. 199 (W—1323343); Warming 185 (W—12348h6). CORNUTIA PYRAMIDATA var. ISTHMICA Moldenke 1 cer bibliography: Moldenke, Phytologia 7: 398-399. Emended citations: MEXICO: Campeche: C. L. Lundell 852 (Ca— 4869),2—isotype, Stee uate F—700445—isotype, W: 149712—~isotype) . Vera Cruz: C. D. Mell s.n. [Minatitlan] (W— 1406477). GUATEMALA: El Petear Cook Finds 128 (W—108),608) ; C. L. Lundell 3450 (F—685306) . BRITISH HONDURAS: C. L. Lundell a (F—598950, “W==19099) , 1O4 (F—598051), 433 (E—980201, F— T1205, W—1:72698) ; Winzerling III.20 [tree no. 20] (F—573771). CORNUTIA THYRSOIDEA Banks & Moldenke Additional synonymy: Cormutia thyrsoidea Moldenke, in herb. i. Sheen bibliography: Moldenke, Phytologia 7: 399 (1961) and 196 Proctor describes this plant as a small , om, tall, with violet flowers in May, growing along roadsides gt an altitude of ee feet. It was oneviously misidentified and distribu- in herbaria as C. jamaicensis sciirnior Additional & emended citations: : N. L. Britton 3766 (W—656325); W. Harris 5731 postage): "eae 9 (F-18395), 11066 (W—699858); Proctor 19689 (N, W--2286912). ADDITIONAL NOTES ON THE GENUS CITHAREXYLUM. IV Harold N, Moldenke CITHAREXYLUM B. Additional ae Citharexyhum Bocq., Rev. Verbenac. 128, pen 2: 85, 88, nded bibliography: Bocq., a 17, 38, 125, 126-128, “ip, 3a aa. ee 5-117, , 15h, nert7, & 160 (186 a tet 8, 185; 19? 197, & L Reo pl. 13. 1863; Bocq., Rev. Ve : 430 PHYTOLOGIA ae 88, 96 109, 117, 118, 123, 126--128, 130 i! 2 7 152, bl, ofa 160, 178, 180, 182--18s, he, 197, alsa 863; Aw Be Hitehe., Ann. Rep. Mo. Bot. G Gard be B76 18933 jaa Quinz. Cent. Pl, ie ios 15 Cc. ag ss Tilust. Handb. Laubholzk. 590. “Re 1, pr. 1, 628, 631, & 807 (3928) nie i 2. b2 631, "& B07. 1925) Sandw., Kew Bull. Misc. Inf, 1930: 210—21 a 2ih. 30; Sampaio, Bol. Mus, Nac. Rio Jan. 13: 267. 1937; Sandw., ew ecm "Wise. i 1938: 3 373. 1938; es - Bailey Man. Cult. Pile, c . 3 e L99.° 1947; cha etheien Roy. Hort. Soc. Dict. Gard. 1: 493. 1951; Asprey & Robbins, Ecol. Monog. 23: 37h, 382, 383, 400, & Ill. ‘ide 19535 Moldenke, Phyto ologia 53 te & 95 (1954) and 5: tos. el - denke, Say + Mo 1d. Set 51 . 2. 1956; Moldenke, Phytolog 509 1957; me ig & ‘a Re + oldenko, 1957 Christ. Bless. (3). 1957; uolderke, » 59: 33b—336. 1958; Moldenke, Phytologia 6: bhB~-$05 & on ire, Je BEOnL PCy Fam. Flow Pl., og 2, 2: 395. 19593 Anon., Kew Bull. Gen. index 1 1929-1956, ie 1979 93 Runner, Rep. G. W. Groff Coll, “362. 1961; Gupta & Mar- » Trav. Sect. Scient. Inst. Franc. (a): 78-79 Cr $9615 Srinivasan & Agarwal, Bull. Seb: Surv —_ — 80. 1963; E. E. Lord, Shrubs & Trees Austral. Gard., 1964; A. De Hawkes, Guide Pl, Everglades Nat. Park “hd nr ets 7. Fournier a. tg vem Costa Ric. 13. 1966; Moldenke, items from the French, fidele, referring to the "reliability" of the timber and therefore with NO reference to violins! one THAREXYLUM AFFINE D. Don Additional bibli ography: Moldenke, Inform, Mold. Set 51 Spec. Ya ner: Moldenke, Phytologia 13: 2 279.6 1 onal citat tions: MEXICO: México: G.L. Fisher son. fee July 26, 192k) (Ws). CITHAREXYLUM AMAZONICUM ee Additional bibliography: Cointe, Amaz. Bras. Pl. Uteis, ed. 2, 376. 1907} i Moldenke, Phytologia 13: 7 919-260. Le Cointe OokD records the vernacular names "pau-de-guitarra” and "pau-di 1a", and says of the wood "para marcenaria, caixoteira e fabrisantio tet instrumentos de cordas." CITHAREXYLUM ARGUTEDENTATUK Moldenke Additional bibliography: Moldenke,Phytologia 13: 280 & 303- 1967 Moldenke, Notes on Citharexylum 43 1966. ITHAREXYLUM HYANTHUM (A. Gray) A. Additional “Bablsogrep : er Pea Da 13: 310. 1966; Moldenke, Résum : Additio mal — oe ngs Luis Potosi: J. Rzedowski 7636 (Du-—-510645). Zacatecas: J. Rzedowski 9323 3 (Mi). CITHAREXYLUM CAUDATUM L. Additional bibliography: Asprey & Robbins, Ecol. 23: 400 & 11. 1953; Moldenke, Am. Midl. Nat. 59: 33k 1958} Moldenke, Phytologia 13: 262—28h, 292, 293, 312, & 31h. Recent collectors in’ Jamaica describe this oa there as a argins rus ed limestone hills, at altitudes of 2100—1l,000 feet. Baas on Yuncker 1767 are described as "white" and on Crosby, Hespenheide, & Anderson 692 as "white tinged with orangish in bud? Ekman 13902 represents a very small-leaved form of the species and may be wort worthy of varie rank, as gested by 0. E. Schulz. My son, — R. eee found the species growing in a mangrove Swamp in Thorne 33366 was collected in a patch of mixed woods with Aca- cia koa, | koa, Metrosideros collina, Hibiscus arnottianus, Cordyline terminalis, Antidesmia . platyphylla, ete., and is described as having been taken from "small introduced naturalized trees with white flowers", Its leaves are not at all typical of C. candatum, being too sharply acute, and have much the aspect of Gs ‘donnell-— smithii Greerm. These Hawaiian imens should be re-studied. & Proctor ae and Webster, ou Vs perkinsi ela —_ ee have very Li somewhat rough-reticulate above. The very fruit on the fruiting racemes, however, seems + doubt on the hybrid na- of this taxon. Perhaps it represents a nat iety, al- though its foliar characters seem to be intermediat The S. (1965) tell us that it exists in pe tion there, but I have seen no ma terial of it from India, or cultivated. They record the vernacular name ritiseisieen* for it there UM FRUTICOSUM var. SUBVILLOSUM Mo idenke Additional synonymy: Cit lum fruticosum var. pentadrum payers ex Jiménez, Supl. Cat. Fl. Doming. 1: 212, onkaba tis © Aeittdock, bibliography: Liogier, Rhodora 67: 350. 1965; Mol- 1967 Moldenke, Notes on Citharexylum 435 =. oe 13: 290. 1966; Jiménez, Supl. Cat. Fl. Doming. Sei 1 eine citations: CUBA: Oriente: Ekman 8638 (Mi). CITHAREXYLUM FRUTICOSUM var. VILLOSUM ee 0. B. Schulz Additional bibliography: Bocq., Adansonia 2: 96 & a a and 3: 223, pl. 13. 1863; Boog, se Verbenac. Later pl. 13. 1863; A. S. Hitche., . Bot. Gard. ie le ae Liogier, Rhodora 67: sou iss ete eae ety 290. 1 Additional citations: CUBA: Oriente: Ekman 7732 (Mi). CITHAREXYLUM HEXANGULARE Greemm,. Additional bibliography: Moldenke, Phytologia 13: 291--292 & MeVaugh found this as an occasional tree along lumber roads on steep slopes in humid forests of Abies, Quercus and other decidu- ous trees, with Pinus dominant on the ridges and summits, at alti- tudes of 2000 to 2250 meters. The leaves on his no. 23162 are exceptionally large and broad, being 16 cm. long and 7.5 om. wide. Additional eatatteed MEXICO: Jalisco: R. McVaugh 23162 (Mi). CITHAREXYLUK HIDALGENSE Moldenke . > iaprrinr ay bibliography: Moldenke, Phytologia 13: 292 & 301. i os tee ee a a 2631 are described as "white or lavender’ ons: MEXICO: Hidalgo: H. E, Moore Jr. 2631 (We); P ; Pringle 8969 a68 tyeckssuaeys CITHAREXYLUM HINTONI Moldenke dditional bibliography: Mold Inform. Mold. Set 51 Spec. 2. 1956; Moldenke, Phytologia 13: O98: 1966. CITHAREXYLUK HIRTELLUM Additional ninrtcacacairs re a Aitiek: Phytologia 13: 310. 1966; Moldenke, Résumé Suppl. 1h: 1. 1966. ee HYBRIDUM Moldenke dditional bibliography: Moldenke, Am. Midl. Nat. 59: 335. 1958; Moldenke, Phytologia 13: 292-293. 1966. ILICIFOLIUM H.B.K. jonal & emended bibliography: Bocq., Adansonia 3: [Rev. ed. a pre 1, 31 pono pee et lh 5 the type collection of C. sate tarta Greem. BOOK REVIEWS Alma L. Moldenke "Essentials of Biological Chemistry", second edition, by James L- 1 and Gordon L. our, xvii & 31) pp , illus. Reinhold yey nei Corporation, New York City, New York 10022. 1966. This new and much-improved edition with major additions of ma- terial on protein synthesis, nucleic acid structure, epirmctnet r try training, the language presentation is clearer than that in many other texts, “Principles of Agronomy" by Lorentz C. Pearson, xv & 3h pp illus. Reinhold Publishing Corporation, New York city, “New York 10022. 1967. $9.50 "Today the agronomist interested in weed control is dreaming of a tah which can be applied at the same time the crop is seeded, one which will destroy all weeds and weed seeds without inj ronomy". This text is Til Ledetne vel by stressing principles and reasons rather than minutia, therefore it is superior to most of the books in the field. t most universal 10th year subject this author have developed in depth more of the genetics and efor Siesta xt reads well 4 despite a ohm ot split pre pak a are jus on p. 12, The questions at the ends of the cha apters routine content ones. A he glossa x are given. Among the earliest plants cultivated are included raspberrie: 3 e kely just gathered crops from plants growing naturally near men's habitations. eee Tatra ate arranged as follows I. Introducti II. Basic Principles of Crop Production: rotation - seed - water - soils - weed other pest control - harvesting - agro- nomic research - pacar - plant breeding 1967 Moldenke, Book reviews 437 III. Field Crops: grains - forage - fibers - sugars - roots - timbers - drugs - tobacco - hops - etc. "Fantastic Trees" by Edwin ihe Menninger, xii & 30) pp., illus. Viking Press, New York City, New York 10022 and Macmillan Company of Canada, Ltd. 1967. $8.9 The author's years of training and serge in eal a pro- sah Donped interesting text and fantastically attractive illus- trations aa these fantastic marvels and monstrosities of the arboreal wi The baker “ts related with peapder ih accuracy, with correct ami ding leaders in the field. The name Avicennia nitida was changed to A. germinans at about the time that this book went to press. How can one put down unfinished a book with headings as: p- teasers, flowers and fruits underground, fruits on the wrong tree, s a reader "sniffs" gp itr nigh e~ it is only becaus Pl is too shallow; the book is about the extremes of adap- tation for survival that confront all organisms whether they are hi canno : the "mustard" of ane Bible was Salvadora persica. a mustard-flavored plant of the area, but its its small drupes could not be the minute "grain of mustard seed" which we believe be- Lipped The printing and photography are fine. Only "usable" s through misspelled on p. 136. The pibliogranty and notes are in- valuable, "Plant Physiology" by Robert M. Devlin, i & 564 pps, cago Reinhold et Corporation, New gat City, New 10022. 1966. $11.00 Here is a fine, Seong new text with clear explana and illustrations all so arranged that an instructor ORE, a profit by using it for either : semester or a year course, and ad aan and nie eee: nneretsg car- bohydrate metabolism involving enzymes, classification, fermenta~ 1,38 PHYTOLOUI«A Vol. 1h, no. 7 tion, pk! Sega and sugar translocation, () i gpa tig it us, Pigments, light/dark reactions and rat 5) mi eral nutrition, function, bre ation, nitrogen netabélin with protein synthesis, (6) plan t growth horm mones both and syn- thetic, and (7) growth and oe as effected ey photoperio- dism, vernaliz ation and do Often there are succinct s sumarie S at the end of topics. Good bibliographic material is provided at the end of each chapter. The "Biological Control of Insect Pests and Weeds", edited od sig 2B De Bach, xxiv & 84) pp., illus. Reinhold Publishing Corpora tion, New York City,” New York 10022; published 196), repriie ted i965. $22.50 This book is a crammed compendium f important, well illustra- ted information written by leaders "tg each field.’ Bedtied 1 dis- cusses the scope, importance and development of biological control principles and —— Section 2 stresses fundamental popula- tion ecology and natural control; Section 3 deals with systematics; Section surveys the importation and establishment of new natural enemies; Section 5 explains the conservation and augmentation of natural enemies; Section 6 describes all types of insect pathology; Section 7 deals with the biological control of we weeds; and esr 8 summarizes this work with "successes, tre possibilities". There are over 2500 literature references "datsng pared partly through 1961 in the bibliography; all are fully in- this thorough book is dedicated to the memory of one who initi- ated much of the work and was the mentor for most of the section —— Royalties from this book go to the Har arry S. Snith hiemor~ The binding on this review copy is already brok “Phylogeny and ae in vie Plant Kingdom" by Howard J. Dittmer, mii & us. D, Van Nostrand Company, Princeton, New Jersey Dies | 196k. $11.25 The author's preface describes his book as "a survey of the plant kingdom with descriptions of the structure reproductive This is a thorough survey ee plant morphology, finely illustra- ted, documented with a few references at the end of chapters, clarified by an excellent glossary, helped with a good index, and uninspi "home~work" questio sex in bacteria notwithstanding. There is much valuable material in this book, but its approach is "dated", BV wos by THREE NEW SPECIES OF OURATEA (OCHNACEAE) FROM PANAMA John D. Dwyer Department of Biology, St. Louis University; Missouri Botanical Garden In preparing a treatment of the family Ochnaceae for a foxthoceane issue of the Flora of Panama I encountered three new species of Ouratea i. OURATEA COCLEENSIS Dwyer, sp. Arbores ad 10 m altae, ramulis pas laevibus glabris. Folia | Polls orn ad 0.8 cm longis, oblongo-rotunda, ad 11 cm longa, ad h.5 cm lata (lamina foliorum angustorum 3 subtus prominente, venis lateralibus ca 15, prominulis sed non evanescentibus late arcuatis, venis intermediis paucis irregularibus, marginibus vel regularibus vel vix repandis, dentibus minutis, ca 3-5 mm distantibus; stipulae saepe per- Sistentes amplexicaules vel divergentes, rectangularss, ad 1 cm ongae, ad apicem angusto-cuneatae. Inf loressentias hic amido=paniculatae axillares, ad 1 longae, @ fag cm latae, longis, gradatim ad perianthum amplifi ovatis ad ovato-rotundis, ad apicem acutis; longa. Fructus non visi. petala ca 8 mm ANAMA: Cocle: Cerro Pajita, N of El Valle de antén, 3500 tt alt, Allen 49 (MO, holotype). The new Bate resembles 0. insulae Riley from which it differs inh a much more reduced panicle ae leaves with the erktnn of the blades more or less 2. OURATEA FLEXIPEDICELLATA Dwyer, Sp. no Suffrutices (?), ramulis teretibus Siageeea Sry Folia a be 0.8 cm longis, crassis ascendentibus bene a angus beer i ad 25 cm longa, ad 6.5 cm 1.2 cm lo ongo, basi obtusa, cams Supra p rominula, subtus — venis lateralibus ca 40 PUTTO0L0G Tu Vol. 14, no. 7 supra tenuibus vix prominulis, subtus conspicuis forte arcuatis, venis intermediis supra evanescentibus, subtus vee aliquibus prime leviter r arcuatis, tune ad marginem evanescent- ibus, marginibus vix crassis repandis, dentibus 7? distantibus, eis in parte tertia superiore divergentibus, ad mm longis,. Tar nocanee tan axillares vel terminales, anguste racemiformi~paniculatae, rhachidibus 8-30 cm longis, prime cea deinde ad apicem flexuosis, ramis arcuato-ascendent- vibus, vix ad 1 cm longis, floribus oS per bostrycem, yd ate 3 egies 0.5=2 cm distantibus. Flores pedicellis breviter articulatis ad 2 mm, filiformibus, in medio ca 0.75 mm tis, minute puberulis forte reflexis. Fructus (hic immaturi) previo Hon es ad 8 mm longo, drupis elliptico-rotundis, ca 4 mm lo PANAMA: Darien: forest near Pidiaque, 800-1600 ft alt, Duke 8050 (MO, holotype) The new species is readily distinguished by its slender bviously reflexed persoels- It appears to be closely do related to QO. patelliformi S Dwyer, described below as a new species, but differs in having a more rounded receptacle in the fruit. 36 es ceties PATELLIFORMIS Dwyer, sp. Arbores ad 6 m, ramulis teretibus avivel glabris. Folia bene distantia, petiolis ad 0.8 cm longis, fae crassis, angusto- pekictio., 15-20 em longa, 5-7 cm ad apicem cuneata, acuminata deinde ultime acuta, ee obtusa vel subcuneata, tenui-coriacea nitente laeve glabra, costa Supra prominula, ad petiolum saepe plana, subtus leviter rominen i us i s plano-compressis, ramis paucis, O.l-.5 cm aonte ts. pores solitariis vel geminatis, eis infra medium ori us 3-5 pee ferentibus, locis articulationis ad 0.3 cm inetes "pedicels ca 7 mm longis. Flores non visi. Fructus recep egale patelli- forme, ca mm longo, 6-10 mm lato, ob ibus druparum circularibus, ca 2 mm latis, forte depress PANAMA: Panama: Pacora, Allen 35 (MO, holotype). The saucer-shaped receptacle of the new estes is unique for the Central American species of Quratea NOTES ON NEOTROPICAL EUPHORBIACEAE 1. Synopsis of South American Sapium E. Jablonski New York Botanical Garden This is the first attempt to present a dichotomous key for this difficult genus. The great variability of characters, very few with diagnostic value has for a long time resisted all attempts to put this cerns in order. filler of Argau in 1863-1875 fighting these difficulties nth. iar = with a confused tri-nominal labyri We Botting pene in es oe vires a series of 35 drawings in Hooker's Icones. The analytical details are accur- ate and the dra seins beautiful, b but the nervature and margin of leaves are often misrepresented and are ieee the product of the draftsman's imagination. acques Huber's work & in 1906-1912 was no space ~*~ rane ie, ever a on this genus. Adhering to a clear nal that sui was unable to finish the Hylaea, the most important of all ar oor mann in 1912-192) with their usual thoroughness t did ee a ait the knowledge available up to that time bu le to treat the gies critically. Lanjouw in 1931 made very significant progress in ogee ing the taxonomy of S. klotzschianum and related spec Croizat in 193-5 described a number of new species from ia and giving has care- fale avoided to clarify their systematic positions. The key presented in the following no doubt requires in- provements. The binominals so far published have been reduc— ed to 58. Further improvements and reduction will be possible when more and better material will become available. A. Subgenus Sapium. Inflorescentia spicata cymulifera 1. Styli persistentes. Petioli longi (2-4 cm)- Secte Emmenosty lum 2. Lamina basi ariculata PHYTOLOGIA Vol. 1h, no. 7 hh2 3. Folia mire integerrima, 7-8 cm longa (Venez: DeFe, érida 1. stylare 3« Folia sat pepberiner serrata, 15-18 cm longa (Colombia)... 2. putumajense 26 i auri ealetey venae 15-25-jugae. Fotis 32 verun na basi non 10-1) cm longa (Colom 1. Styli non persistentes 4. Glandulae petiolares obsole vix pc btege Petioli i (Hy oli insertae. we Folia vel subcordata (West Hylsen) marmieri 5. Glandulae petiolares apice pe tiold insertae. a basi cuneata (West Hylaea, Colombia)... 5e e eglandulomin le Glandulae petiolares conspicuae (excepto Se haematospermo, foliis linearibus facile distincto Re petiolares, sub laminam inflexae saepe ul Folia apice plana, elliptico lanceolata vel oblanceo- ata; venae pauci-jugae (6-1 cm longa, 1.5-2 cm lata), obo- Te dentata, saepe revolu subtus ph geraoee sed non prominentes (oreeiaiia owianum orientalis)... se hides dyads ees cm longa, 3-6 cm rate), membran= 8. 1 chartacea, concoloria, viridia, longe acme aus margine obsolete dentata tig venae tk bite dupciaaae 6-12 jugae eeape Para rodentatan 7e Folia tsice babahlato Gortess. vel seAiuacteeanes 9- Petioli elongati, plus quam 1/ laminae longi (Rio de Janeiro). S. donghpes Ie Prout ae usculi, non 1/) laminae plisaee e Janeiro) .... ublanceolatus 6. ee pehsnaaiee; variae sed minquam " iiflex ub lami occultae branacea wes ga teaag viridia; venae subtus cai apeigen EG a apice plana 625-9 cm cae, 3 cm ata); venae bkainsl feeue ($-19) arcuato- sascendente 12. Petioli pp igareee (0.5-0.9 cm). Folia basi cuneata Ftos Ayacucho) ereeesen 10. contortum 12. Petioli longi (1.5-2.3 cm) oe Oe Fe 3- Venae a~T~ Jugs ae, vue prominentes Glandulae petiolares graciles, eylindricae (Br. ae na) 11. paucinervium 13. Venae S10 gugee, haud prominentes. Gland- ulae petiolares minores, hemisphaericae. Folia 8-10 cm longa 1967 Jablonski, Neotropical Euphorbiaceae Lh3 ly. Ovarium 2-loculare (Suriname)....ececsecs 12 1). Ovarium 3-loculare. Folia ovato lanceo- lata ay °9 cm longa, 2-)) cm lata) (Br. ian Gu 13. jenmanii ll. igen mpage cucullato-inflex a 15. Folia minora, obovata rs oblanceolata, minus quam eat elo longiora quam lata (7-10 cm longa, 305 cm lata) (Suriname, Delta, ae es Ales nifolium 15. Folia majora, lanceolata (10-18 em ome 265-3 cm la a Guiana, Bolivar, R. phage de Para, Ceara) eo. 16. yn oo petiolares patentes, longae Par + patens 10. Folia coriacea, pea vel bicoloria in siceo brunnea, nec viridia 17. Folia lata non linearia (minus quam 8-plo longiora quam lata 18. Folia margine integra vel tantum subserrulata 7 Folia Be plana, rariter subcucullata “21. Folia aoe vel subrotundata (minus quam bis longiora 7 lata) 22, Petiols ptaeeetl (Brazili orientalis) ecscccees 17. obovatum rg Petioll lone (1-2 em); venae 236 vais 1 Bua jugias prominulae; glandulae petiolares conicae Peru, Colombia) .18. bogotense 23. Venae se bapa glandulae sieabecaaoag subcylindricae 9. pavonianum 21% em icceata Seg quam bis long- ra quam la 2he “Petioll Bae (2-4 cm) “ ndu = petiolares ovoideae e (cf. S. eglandu- ae Al Negro, a 1 Guianas)...e. 20. aubletianun 25. Glandulae petiolares cylindricas distinctae; Vv 20-22 -jugae ecreeerese eee + 22 « myrmecophyllum PBs 20° 9:G;304 Vol. ly, no. 7 20. Costae secundariae laxae, graciles, patentes, saepius haud anastomas- mers dictyodromae, ultinae vi perspicuae 26. Petioli 2.5-) cm apace venae 20- jugae. a lh nea, = cm lata ahora re 2200 m tS eer e CaGeleoe chetresasasii 26. Pebiolt a 5 ¢ i dclengst oa S=jugae. Folia lhc longa, 6 cm lata, lanceo- lathe anaae — (Eucador) «+s. steyermart 19. Folia pice se age fe e S. ixia- masense) 27. arias subtus prominula, rigs venae rectae vel arcuato-ascenden 28. Petioli pengnnenntt (1.5-4 a) 29. Capsula sessi 30. Gland si tee anki lares graciles divaricatae, rectae vel cur- vatae 31. Folia lata (latitudo plus quam dimidi helguatense orientalis inter Cat te Sees 256 klotzschianum 31. Folia angusta (latitudo minus am 2 ce 336 i i Peeees fa hemsleyanum Venae Sasol teas TEES Folia ae poets ike W205 c ata) (Venez: resale Oey a. ) @eene 296 obtusilobum Snel Jablonski, Neotropical Euphorbiaceae Ls 30. Glandulae er te minores, hemisphaericae. Folia basi suorotundata, vel late cun 35. rarer supra opaca, metallice nitentia (Pe yla i oon 30. aereum 35. Folia supra nitida (Peruv. ndes eeeeee S16 poeppigii 296 Eh lpi gate i i a (9 cm longa, 5 cm lata). Venae Berl Sines (Venez: D.F., e)/overeee 32 ° nai abunes 28. Petioli 3 eT - Lami taceae Foli scure cucullata (2-8 cm longa, 2-3.5 cm Venae 12-15-jugae (Peru: Tara pot re) ) » peruvian 27. jNervatura, subtus obscura, a Folia minora a ata. 12-15-jugae. Petioli longiusculi (1.3 em) (Peru, Bolivia, Matto Grosso) ecceese ixiamasense 18. Folia margine distincte —— bat crenulata 376 reaie majora (plus quam 10 cm longa, ad 18 cm ga) lanceolata vel cae as elliptica (2-5 er segty ves a quam lata) ‘Petioli — (0.8-1.0 cm longi r 38. S)ecceccese 360 hasslerianum 38. Petioli tanetnseull (1 .0-3'« 5S tm longi). Nervatura laxa; — 12-15-jugae subtus ae s subimme ellitica, Slabeeinis long- iora quam lata hO. Folia margine 390 m e cremulata basi subrotundata, apice modice cucull- ata (Parana, Minas Geraes)eereee case pis ense ada dentata apice leviter cucullata vel plana cuneata (Ins. F. Noranha,Maturi Cristobal ?)eeccee 38. pg hO. Folia mel ne PHYTOLOGIA Vol. lk, no. 7 396 Folia anguste lanceolata, acumin - margine modice crenulata (Cc BO) Fhe enc 9. aucuparium hl. Glandulae petiolares graciles cylin- ’ aricata ara: - arem, Belem) LO. ciliatum 37- Folia minora (2. 5-10.0 cm) apice plana (cf. iste 2. Folia calloso-marginata, obtuse crenulata 43. Folia lanceolata, quadruplo longiora quar la S Paulo, Paraguay)e.... il. marginatum 43. Folia minora elliptico-Lanceolata, bis longiora quam hue Folia oratorelliptica eS om longa, 2 cm lata (Par ay) e- ee ed wh2e aoe 4h. Folia anguste ellipti 4S. Folia 5-7 cm longa “(Bans A) ccceece icatricosun Se Folia 2.5-3 cm Sian “eucuTlat (Goyaz, Piauhy) ese. le sie 42. Folia non calloso-marginata 46. Folia Sy Parpenpeniggen basi cun- eata, 2-34 oc Ne apice acumin- buco) 6. Folia elliptico-lanceolata h Petioli breviusculi 48. Folia cremlata sed non argute serrata h9. Folia haud distic eeeeeeenee h da 50. Folia non reticulata, con- coloria 51. Columna stylaris bre- vis (Piauhy, Rio Grande del S$ Urugu longa, l-2 c (Uruguay) eoe af . “pibertid 1967 Jablonski, Neotropical Euphorbiaceae bh,7 48, Folia regulariter argute cren- © h7e Petioli eee ein te Folia crenu- lata sed non argute crenata (Stae on Gi Re Gredel Sul) eerccecese 17. Folia linearia, §-20-plc “ongiora _aven lata. ioli brevissimi nee S24 Ovar rium 2-lo 53. Folia eas cm Sates 6 O.l:-04.7 cm lata (Bolivia) esses soscesce O2e bolivianum 53. Folia 10-20 cm longa, 1.5-2.5 cm latas distinc- tae (Argentina, Paraguay, ———— Pinney) (The Piauhy bine is 3-carpell oo C36 longifolium 52. Ovarium ete pce fa: Sue Folia mims pie 6 cm tenets? 0.5-0.7 cm lata P TU)ececeassvseessere « She ——— She Folia magis quam 10 cm longs 55. Folia magis quam 1.8 cm lata Lge tent S. saltense 55 Folia mimus quam 1.7 cm lat 56.6 _ pea: cm lata. Slandulse pet- lare: bs (krgentirs, Uruguay, P circ eres . haematospermum 566 Folia 0. “220.6 cm tate (Paragu ay)ee | Linearifoliun A. Subgenus Pseudo-senefeldera. es ali i we )Alto Orinoco) bd * oe eeesevee 58. yut aje ense 48 PHYTOLOGIA Vol. 1h, Plate 1 Distribution of Sapium in South America Figures in brackets indicate synonyms aereum Kl. 30 microdentatum Lanje albomarginatum Pax & H. 50 montanum Lanj. aubletianum Huber 20 montevidense Huber aucuparium Jacq. 39 muvelleri Hemsl. Huber 5 = myrmecophilum Croiz. bogotense Huber 18 naiguatense Pittier olivianum Pax & He 52 obovatum Kl. cicatricosum Pax & H. 43. obtusilobum Nevin Arg. ciliatum Hemsl. O occidentale Hube cladogyne Hutch. (13) pa um Hu claussenianum Hube (25) paranaense Pax & H contortum Croiz. 10 patens cremostachyum Johnst. (25) paucinervium ° cutrecasassii Croiz. 2 paucistamineum eee cu erum Herzo (52) pavonianum decipiens Preuss (21) peloto Pax & i. diandrum Pax (25) peruvia state nove eglandulosum Ule 5 petiolare Huber fragile Pax & H. (55) poeppigii Hemsl. gibertii Hemsl. 9 oh ° glandulatum P 25) prunifolium Kle glandulosum Morong (53) punctatum aricense Pittier (50) putumayense Croiz. haematospermum Mull. Arg. 56 rojasii Le hamatum Pax & H. (31) saltense stat. nov. hasslerianum Huber sceleratum y emsleyanum Huber 27 ~—s selllowia Hub ippomane E.P.Meyer serratum Mull. Arg. intercedens Huber (25) stenophyllum Huber asense Spe Ne steye kii spe ne jenm ‘“ 13 = stylare Mall fe klotzschianum Hube 25 sublanceolatum Huber lanceolatum Huber 15 bmarginatum Huber leitera Gleason ( h) leptadenium Huber (25) tenell & H linearifolium Hemsl. thomsonii Jumelle longifolium Huber 53 tijucense Huber longipe I 8 tolimense Jumelle marginatum Mull. Arg. 41 triste Huber m ri utile Preuss martii Huber (17) verum Hemsl. yutajense sp. ne 58 no. 7 NANO fal WL we ww Loe =a Were wywW 1967 Jablonski, Neotropical Euphorbiaceae Lg Distribution of Sapium in South America. Figures in circles indicate species with restricted distribution. Figures in hexagons indicate species with broad distribution. Outline of areas approximate. 300 600 MILES ° '- a ts NOTES ON NEOTROPICAL EUPHORBIACEAE } \\ 2. New species and transfers E. Jablonski New York Botanical Garden sem Beets Jable spe io lanceol ato iene a a quo at pl Pade pager pe grac- iba cylindricis divaricatis ne conici foliis basi subro- tundatis apice angus te acyminatis minusque “ened liatis differt. us. Brasilia: Para, Rio Delia, San Miguel, Froes 20372 — specimina visa: Para, Rio Guama, Irituia, Froes 20373, io Rio Tapajos Usheuld tthe, Pind fg 832, 81436. SAPTUM STEYERMARKII Jable Spe nov Ex affinitate S. ixiamasen: a A qua venis 5-6 jugis subtus prominulis arcuato-adscendentibus optime recedit. Folia late lanceolata margine pepe -e repando-dentata; venae ce ban ood glandulae peticlares subcylindricae. cuador: Prov. Santiago-Zamora, Setieaiay Soest moist on soos ve and bordering Rio Tintas a campamento, southeast of El Pan, alte 28602985 Julian A. Steyer- mark 53508. SAPIUM IXTAMASENSE Jabl. spe nove elene a) Ex affinitate Se lanceolati a qua foliis coriaceis bicolor- ibus superne nitidulis subtus Tap Re i inites differte A bor 30 Ogee) is, habe 1-2 pedes diametiente. Pet- ioli ype pag (1.3-1.5 c pe 5 Glandulae petiolares divar~- icatae conicae mims quam a mm altae. Folia 2 1/2- 3 1/2-plo longiora, quam alae plerumque 7-12 cm longa rariter ad 18 cm a 3- cm lata lanceolata a apicem basinque versus symmetrice Shane stata eetae oblanceolata basi cuneata apice caudata imo i U apice incrassata distincte cucullata vel rari Vix llate rarissime plana margine integerrima superne nitidula subtus opaca wat atst hers -viridula; costae seuatirtas 12-15-jugae superne fere prominentes subtus immnersae vix visibiles. Spicae ligerae 7-10 gae in axillis foliorum superi orientes folia ae vet ee apice flores paucos masculos ceterum femineos gerentes vel u xuales masculae; rhachis eee cymuli masculi SBtones. featn ei 1-2-flores; bracteae late triangulares subintegrae utrinque basi glandula tiviptios auctaés Ovarium sessile; — stylaris decidua. Capsula depresse~ globulosa matura 7 mm diametiens sessilis; styli decidui. Semin ee 301-3.) mm senate adie atr trobadia o-arillatae 891 (NY gel Bolivia: Ixiamas, alt. 1150 pedes, Re 5. Williams 450 1967 Jablonski, Neotropical Euphorbiaceae 451 pene cetera visa Mouth of Rio Santiago Tessmann 649 (NY) % Preeti: Solimoes, Froes 5662 (NY Falls of Madeira, Rusby 1239, 1238 (NY) % Nota. Rus by 1238 forma foliis apice non cucullatis. ee aoe Jable Spe wh oe 8 Os Ha § affinitate St. dichotomae Mull. Arge a qua hue ce eat weeietap ein et distinctius “serrato~dentatis, et basi glandulis minutis duabus ornatis optime ferte ex 2 metra lis nosulis in siccitate quas lignosis. F pposita vel ad apicem r o oR Sti- lae cito caducae ae _— lus 3- mm BUS na Sus angustata, imo apice wncaintaul, paulo succulenta, poset e argutissime serrulato-dentata, crenaturis angulo recto divarica- tis vel irregulariter curvatis, basi superne glandulis geminis i i liform i sicea 1-2 mm lata. _Capsulae 5-6 mm longae pericarpii basi coc- radiat ecopharun' el "gynobasin" dict nec usa centralem formanti. Semina 2.5-3.0 mm Sue subglobosa atro-badia, albo- ulata. Soeciad rs mucedinosa, ideo in pein valde nas ee Typus: Minas Gerais: = n da Rod srarge grin -Teofil Otoni.e Serra rochosa. Arbusto de 2 m relas, trates verdes. 1) Aug 1965, Romeu P. gnome ths, "helouine (UB); 1616 paratype (UB). SAPIUM YUTAJENSE Jabl. spe ni In Memoirs NY Botanical ic, in press. SEN PERUVIAN Gaskets) Jabl. stat. Bh oS Tu Sapium tii var. peruvianum Ma es in ora e Field Mus. Wet Histe “tal. Scare 13, 3a, 1: 195. 1951. SAPIUM SALTENSE (O'Donell & Lourteig) J state n Sapium haematospermum var. as as “ gr cae in Lilloa i: C86. 19e. 452 PHYTOLOGIA Vol. 1h, no. 7 SEBASTIANIA RHOMBIFOLIA (Rusby) Jable comb. n apium rhomb st usby in Bull. Torey” Bot. Club 28: 307. 1901. (Plate I Corrected SR and additions to original description: Petioli 0.5-1.0 cm longi (nec 1-5 cm), eglandulosi. Lamina ctae Fruct Bolivia: Falls of Madeira, Rusby 182). 1967 Jablonski, Neotropical Euphorbiaceae Plate II Sapium ixiamasense Jabl. Spe Ne Fige Awe Habit lle B. leaf, nervature exaggerated l,/5 Cy Cudcliats tip of leaf /1 D. Ma ower 15/1 E. Male part of inflorescence 6/1 F. Female flower 12/1 Plate III Stillingia acutedentata Jabl. spe ne Fige A. Habit ),/5 Be Male cymile 12/1 De Capsule h/1 Ee Gynobase }/1 Plate IV Sebastiania rhombifolia (Rusby) Jabl. comb. n. Fige Aw Habit h/S B. Leaf, nervature exaggerated l/5 2h/1 ym 4/1 Female flower 8/1 453 45h PHYTOLOGIA Vol. 1h, no. 7 Plate II 1967 Jablonski, Neotropical Euphorbiaceae Plate III Y 55 456 PHYTOLOGIA Vol. 1h, no. 7 ——<— PHY TOLOGIA Designed to expedite botanical publication Vol. 14 May, 1967 No. 8 CONTENTS SMITH, L. B., Notes on Bromeliaceae, XXV . dd SMITH, L. B., & SMITH, R. C., Itinerary a William = Burchell in Brazil, 1825- 1830 492 MOLDENKE, H. N., Additional notes on the genus pre eee, v.50 MOLDENKE, A. L., Book reviews... -+- eee oal Index to Authors in Volume Fourteen. .-+++++* +t 0 3° 513 Index to Supra-s pecific Scientific Names in Volume Fourteen 513 520 Publication Dates of Volume Fourteen - - + +++ +2 ** 5 "°° Published by Harord N. Moldenke ‘ek Alma L. Moldenke "303 Parkside Road, a, Nee = Jersey 07060. U.S.A- Price of Bes: number, $1; per volume, $5 1, in advance, or $6 at close of v NOTES ON BROMELIACEAE, XXV Lyman B. Smith t is over 30 years since Mez made a comprehensive survey o ati Mez's pollen distinctions, there are many changes ahead in the Bromelioideae. Any synonymy since the Pflanzenreich is noted under the appro- priate genus BROMELIACEAE 1. Mature seed appendaged or if not (Pitcairmia in part and Navia) then the fruit at least partly capsular and dehis- cent; ovary superior or largely so in most genera, to in fe rior; fruit c epauier or if not (Pitcairnia in part) then the Seed appendag : Seed~appendages witire or slightly divided (Broechinia pani- culata) or lacking (Pitcairnia aphelandriflora and Navia); eA: usually dehiscent; leaves mostly spinose-serrate; in- dument of finely to scarcely divided auras ee usually terrestrig loi s capitiform or rarely spiciform with fascicles of flowers in the axils of ample bracts (Nidularium peat) often involuc Se ess 1,60 FHET.OLOGT & Vol. 14, no. 8 8. Flowers pedicellate. 9. Stamens equaling or exceeding the rounded erect petal- blades; plants of Chile...... -....24. OCHAGAVIA 9. Stamens shorter than the acute aprendthne petal blades; plants of eastern Brazil to Amazonian South America. 25. NEOREGELIA 8. Flowers sessile. 10. Epigynous tube very short or lack ing. ll. Outer bracts of the central inflorescence foliaceous; petals spreading, obtuse; usually some of the flowers unisexual; plants of eastern Brazil....26. C s 11. Outer bracts of the inflorescence bracteiform; petals erect ‘or if spreading (Nidularium in part) then acute. ts amp ie rg dean plants from Mexico and Venezuela to Chile 28. 10. Epigynous tube elongate; scape elongate, slender; inflo- rescence not involucrate; plant of eastern Brazil. 29. ANDREA 2s genehs forming a tube to which the fleshy petals are along their centers but with their margins free; pianes of Mexico and the West Indies to Argentina ruguay Gs . = 30% TA 1. Petals appendaged wisn ovals naked (ascnnes in uy then nh 14 Ovary is art superior; scape naked; inflorescence simple, strobilate, aap oS eas of Pages ania razil and adjacent BLEAS+ sees. ee vie tlcs erior 1 Bee aps: -bracts foliaceous or the scape lacking; sepals free. 16. sepals neesly or quite symmetric; petal -appendages well Lope Efe Planents of second series adnate to the ogame scape sually present; plants of eastern Bra nee RTHOPHYTUM 17. Filaments all free; scape lacking; plants of eve. FASCI CULAR 16. Sepals definitely asymmetric; petal-appendages vestigial; Plants of North and South Amer "38. AECHMEA (in part) 15. — based ied from the leaves, or if there is no eviden (Aechmea then the sepals much re ¢ . 18. Inflorescence involucrate; sepals unarmed, nearly or q free in most species; plants of pe ema aaik: 1967 Smith, Notes on Bromeliaceae L61 19. Petals completely fre ..34. CANIS 19. Pet artially nin as eee ce ek often free et the base and exposing the bases of the filaments of RE TBCE LER 6.4.6 scales wsea € - > 35. WITTROCK. 18. teticcameeccs not ivr Vaseate or if lcneane so then the se mucr ented mt Bae con 20. Inflorescence nate. d (simple in depauperate re aac of Rabanbinita Sie as flowers in ststelaat pikes, much compresse va Epigynou us tube very salt or lacking; pollen-grains ollen-grains with more 4 pores; plants of Costa Rica and the West o northeastern Brazil.... -e-+-37- GRAVISIA imple if compound thet Min flowers not in strobilate spikes. ee. Flowers sessile or if rarely pedicellate (Aechmea in rt ee. 23. Sepals mucronate or ent or if blunt then small and the ° aaah es PSR plants of North and South Amer ....-39. AECHMEA (in part) 23. Sepals “unamned oF pipe uuilcutates ovules unappendaged. 2h. Ovules num 25. Petals best Bi erect or suberect; pollen-grains baad pores; flowers sessile; plants of eastern ER pees: vekene cease tceee« .- 39. QUESNELIA 25. Beaty. Giines aiviroith eta or A ecard in a spiral; dry pollen-grains usually with a single longitudi- nal fold; flowers sessile or pedicellate; plants of Central America to Argentina and Uruguay. 40. BILLBERGIA 2h. Ovules few; ere ee regular; plants of northeastern Bra rrre ey ree etre OGLAZTO peaaae Viesces pedicellate; Pemeats connate; eolian-rcaine with more than 4 pores; plants of eastern Brazil. 4O, PORTEA 13. Ovaries Serre with each other and with the fleshy bract orm syncarp; inflorescence usually with an apical coma, as cet sepals never mucronate 26. Inflorescence with a small inconspicuous coma, never pro- ducing basal shoots; plant propagating by elongate rhi- zomes; petals bearing vertical folds; plants of Faraguey nd adjacent areas...... PS. 26. Inflorescence ieumaie ‘bie: a : lees conspicuous coma, baal with basal shoots; rhizomes lacking; petals usually bear ing well developed scales; plants probably native from Paraguay to Amazonian Brazil, now pantropical..10. eee ee ee eee I. PITCAIRNIOIDEAE 1. PUYA Molina, Sagg. Chile 160, 351. 1782. Type: P. 462 PHYTOLOGIA Vol. 1h, no. 8 ontsene’s Molina. ENCHOLIRIUM Mart. ex Schult. f. in Roem. & Schult. Syst. T, pt. aBy Bs Mabe 3: Seg Type: E. spectabile Mart. Smith, Phytologia 7: 171. ecda Type: F gishesthe (Lindl. y ee 'B. Smith (Pitcairnia micrantha Lindl.). indmania Mez in DC. Mon. Phan. 9: 535. 1896, in ative part but not as to type. ; oa N. E. Brown, Trans. Lin er. II 66. 1901. Type: C. augustae (Rich. ‘Sehonburek) 3 N. E. Brown ee rium Suen etas “aieh. Schombur, Schult. f. in Roem. & Schult. Syst. 7, pt. 2: ixiv, ee Meee ihe. Type: C. florida Schult. f. Lindmania Mez in DC. Mon. Phan. 9: 535. "1896. Type: L. guian- ensis (Beer) Mez (Anoplpphytum guianense eth 6. DEUTEROCOHNIA Mez in Mart. Fl. pt. 3: 430, 506. 1894. Type: D. longipetala (Dake) Ri ‘paki actiolate er). 7. ABROMEITIELLA Mez, Bot. Archiv. (Berlin) 19: 460. 1927. SA pulvinata Mez. 8. PITCATRNIA L'Hérit. Sert. Angl. 7. Jan. 1789 (nom. cons.)-. - bromeliaefolia L'Hérit. 9. BROCC Schult. — in Roem. & Schult. Syst. 7, pt. 2: ee) tees 1830. ‘Type: paniculata Schult. f. - NAVIA Schult. f. Fi Roem. & Schult. Syst. 7, pt. 2: son 1195. 1830 (nom. cons. propos.). ‘Type: N. caulescens Mart. eros 11. DYCKIA Schult. f. in Roem. & Schult. Syst. 7, pt. 2: lxv, 1194. 1830. Type: D. densiflora Schult. Prionophyllum K. Koch, Ind. Sem. Hort. Berol. 1873, App.: T- 1874. Type: P. selloum K. Koch. The distinction of Flowers partially unise xual doe does not seem strong enough by itself for oo from Dyckia. - HECHTIA Kl. Allg. Gartenzeit. 3: 401. 1835. ‘Type: H. Bakeria André, Rev. Hort. 61: 84, pl. 1889, non Seem. 1864. Type: B. tillandsioides igre dré. Bakerantha L. B. Smith, Contr. Gray Herb. 104: 72. 1934. Type: B. tillandsioides save} L. B. Smith (Bakeria tillandsioi- des An André). - TILLANDSIOIDEAR 13. TILLANDS ae as 286. TS; Gen. Pl. ed. 5. 138. 1754. wees t a, pace Ge oe SEA Lindl. eo Bee 29: 10. 7 Feb. 1843 thine ) fata: romel. T (orth. et nom. cons. Type: pe ‘tieoina (Hook. ) vind a et tots psittacina look. ). jum emend. sensu Mez, Bull. Herb. Boiss. ser. II, 3: 9. Thec 1 pes & 1903, non André i8a Smith, Notes on Bromeliaceae 1,63 1967 Cipuropsis Re corneas Bot. Ver. Brandenb. 48: 148. 1907 : C. subandin le. £5 ~ GUZMANIA R. i P. Fl. Peruv. 3: 37, pl. 261. 1802. Type: G. tricolor R. & P. oa André, Bull. Soc. Bot. France 24: 167. 1877. ‘Type: S. ta Sodiroa aramini folie André. cophyllum André, Bromel. Andr. 107. 1889. Type: T. poo + ik Cf. Smith & Pittendrigh, Journ. Washington ae nek ais: 401. 1953. Chirripoa Suesseng. Bot. Jahrb. 72: 293, pl. 4, fig. ll. 1942. : C. solitaria Suesseng. Monotypic genus, a taxonomic syno- nym of eumen ta polycephala Mez & Werckleé. 16. OBROMELIA L. B. Smith, Proc. Am. Acad. 70: 151, pl. l, fig. el #6 1935. ‘Type: M. bicolor L. B. Smith 17. CATOPSIS Griseb. Fl. Brit. W. Ind. Pak 186). Type: C. nutans (Sw.) Griseb.. (Tillandsia wee 18. GLOMEROPITCAIRNIA Mez Bull. Herb Be Ss. ser. qe 1905. Type: G. penduliflora ’ (Griseb. J. i ere ee flora Griseb.). I. BROMELIOIDEAE ee ante 19. 1889. Baker, Type: F. itati- a8 mig gt Baker (Bromelia itatiaiae Wawra - RONNBERG TA E. Morr. & André, Ill. Hort. Be 120, pl. 177- 1874, Type: R. morreniana Linden & André. 21. ARAEOCOCCUS Brongn. Ann. Sci. Nat. ser. II, 15: 370. 1841. : A. micranthus Brongn. STREPTOCALYX Beer, Flora 37: 348. 1854. Type: S- Poeppig- ii Beer. 23. ANDROLEPIS Brongn. ex Houllet, Rev. Hort. 42: 12. 1870. Type: A. skinneri Brongn. ex Houllet : 168. May 1856; 2h OCHAGAVIA Philippi, Anal. Univ. Chile 1 Bot. Zeitung 14: 647. Sept. 1856. Type: QO. elegans Philippi. Eiacseptelis Espinosa, Bol. Mus. Nac. Hist. Nat. Chile 23: 5- O47. : P. rebecae Espinosa 25. NEOREGELIA L. B. Smith, Conte. Gray Herb. 104: 78. 1934. Type: Billbergia meyendorffii Regel. Lindm. Gfvers. Akad. Holm. 542. 1890, non Schau : R. meyendorffii Ble: Lindm. (Billbergia ese dorffii Regel Aregelia sensu Mez in IC. vee 7. 9: 61. 1896, non Kuntze 1891. Kuntze's Aregelia was aming of Nidularium and there- fore can not be applied to a searegate ree ae c rtenzeit. 4: 297. 1836, S Otto & Dietr. Allg. Ga Cc. A pase? Otto & Dietr 2 non Osbeck 1757 goat acne dg pore i Rgpseinn og Lem . Fleur. 4: pl. 411, misc. 60. 1854. ns Lem pe ge ono fey: Gen. 2: 698. 1891. Type: Nidularium ens Lem. An unnecessary renaming ou of the suppos duplication of Nidularia Bull. in the fungi. 28. GREIGIA Regel, Gartenflora 14: 137, pl. At Roe: Type: plivep ys (R. & P.) Regel (Bromelia sphacele te 46h PHYTOLOGIA Vol. 14, no. 8 Hesperogreigia Skotts sberg, Acta Horti Gotoburgensis 11: 220. 1936. Type: H. berteroi (eins Skottsberg (Greigia ber- teroi Skottsberg 9. 29. ANDREA Mez := - Mon. Phan. 9: 114. 1896. ‘Type: A. sel- lowiana pt Rekeee: ae @ esnelia selloana Baker). 30. Pl. 285. 1753; Gen. Pl. ed. 5. 138. 1754. Type: = inac eseran — i - Phan. 9: 12. 1896. : D. ur- SP ring (Mez) Mez (mm thotostachvs ebeciann hey The character or Piles th inflorescence by itself is not enough to distinguish anthon , ch ec ally as sovetaa species of Bromelia are very littic branch = CHYS Kl. in Lk., Kl. & Otto 2. Rar: 1840. Type: A. he ob Sfaniane f: - e pl ; Gobeibercia strobilacea Schul PHYTUM Beer, Flore 37: “3h7, 1854. Type: QO. es ae (Prantleia glabra ez). Beer indicated no species in his description of Orth Sincoraea Ule, Bot. — sig 191. 1908. Type: S. amoena le Cryptanthopsis Ule, Bot. Jahrb. 42: 193. 1908. ‘ype: C. saxi- cola Ule. 33. FASCICULARIA Mez in Mart. Fl. Bras. 3, 33 Ne 189}. Type: F. bicolor (R. & P.) Mez (Bromelia wideten nee . ). 34. CANISTRUM E. Morr. Belg. Hort. 23: 257. 1873. Piste Cc. aurantiacum E. Morr. 35. WITTROCKIA Lindm. Svenska Vet. Akad. Handl. 24, no. 8: 20. 1891. ‘Type: W. superba Lindm 36. HOHENBERGIA Schult. f. ah Roem. & Schult. Syst. 7, pt. 2: ides: genie 1830. Type: H. stellata Schult. r. 37 RAVISIA Mez in Mart. Fl. Bras. 3, pt. 3: 179. 1891; 299. 8 G-. exsudans (Lodd.) Mez (Bromelia exsudans Lodd. It is veconing increasingly difficult to separate this genus from Aechmea ES ‘AECHMEA R. & P. Prodr. 47. 1793. ‘type: Ae. paniculata R. ~.. Stax Raf. Fl. Tellur. 4: 25, 1838. Type: E. glauca Raf. nom. illeg. (Bromelia melanantha Ker-Gawl.). Chevalieria Gaud. Atl. Voy. Bonite pl. 61. 1843. Type: Cc. sphaerocephala Gaud. Mes m maintained this genus on the basis of naked ae yet the original plate clearly shows them to be appen i ad enthus =: Fl. des Serres 3: pl. 227. 1847. Type: D. i kia Sion” ri Mart. Fl. pt. 3: 179, 274. 1 Nov. 891. Type: W. ingulata (L.) “ie ieee lingulata L.). Contrary to Mez's description, the petals are Be roo 39. QUESNELIA Gaud. Atl. Meee Bonite pl. abs 1842. Type: Q. rufa Gaud. ‘ BILLBERGIA Thunb. Pl. Bras. Dec. 30. 1821. aype: B. spe- ciosa ores. qi. OVIA Mez in Mart. Fl. Bras. 3, pei 35°179— 1691. 1967 Smith, Notes on Bromeliaceae 65 hae N. variegata (Arr. Cam.) Mez (Bromelia variegata Arr. am 4O,. PORTEA Brongn. ex K. Koch, Ind. Sem. Hort. Berol. ae App.: 7. 1857; Ann. Sci. Nat. ser. IV, 6: 368. 1857. Type: kermesiana Brongn. ex K. Koch. 43. PSEUDANANAS Hassler ex Harms in Engler & Prantl, Pflanzen- fam. ed. 2, 15a: 153. 1930. Type? P. macrodontes (E. Morr.) Harms Ananas macrodontes E. Mor 44. ANANAS Mill. Gard. Dict. ae uae 4. 1754. Type: Bromelia ananas L. DOUBTFUL OR EXCLUDED ANACYCLIA Hoffmannsegg, Preiss-Verzeichn. Pflanzen 1833: 1833, in adnot. Type: A. farinosa Hoffmannsegg. Identity uncer- pe) P Neck. Elem. 3: 150. 1790. No type indicated. Cited by some as equivalent to Bromelia, but impossible because of the unilocular ovary. Possibly a member of the Zingiberaceae or [AETHAKYA Herm. Mus. Zeyl. 55. 1717- Based on Bromelia, — a etc. L. Fl. Ceyl. 54. 1747, where cited. Accord- ing to Mez in Fngler, Pflanzenreich IV, 32: 636. 1935, this be- longs i the Pandanaceae. DYCKIA The four largest genera in the subfamily Pitecairnioideae are Pitcairnia ckia and Navia, in that order. All but all directions and it is necessar interminably. The present key is a working key and much les stable than the others, but I am ey eae it here to give other people a chance to use and improve Previously, I tried to make a pica division on whether the with the filaments were connate or not above the common tube e others, so I have decided to make what it is no wo than use I can of it. Thus, the present key is a completely artificial one to serve the purpose of identification while I grope for something more Significant. eeneweceeereseeeee te Cee lb Wens Sue Rhee ae eee e eek a eee SS ALE Sr ee ke 466 PHYTOLOGIA Vol. 14, no. 1. Pedicels 20-35 mm long, much exceeding the iets bracts; af eaves over 1 m long, a me cs ia with t a: mn lo Brazil: Minas Gerais : “i - pedicellata 1. Pedicels not over 15 mm long. 2. Inflorescence tripinnate or more divided; med bracts not more th ge long; sepals not more than 5 mm; some xual (Prionophyllum). 3. Bik irae omy densely Fe pad eben Brazil: Sant Catarina, Rio Grande do Sul..ccsccccecscccecce De fet cas 3- Inflorescence wholly pre do Sul or einen “Rio Grande D. selloa 2. Inflorescence not more than ‘bipinnte or the eal bracts and sepals larger 4, stanene long-ex gateas (by more bes the length of the — Shares lax. y: 5 Pedice ls [-11 sig Sela sepals ears mm long; inflorescence lep idote D. igenis or Pedicels Lae yaad noe: “sepals 6 mm hans tiPiorescence ac shit 6a ieiw vie ~-5. D. exserta h, sliets ite exserted by m iy eS of the anthers. 6. la an ape lacking; Aoepiadepiaiae enete, 4 om long. Brazi Goids ieesevwe D. odorata 6. Scape e Present ¢ or © where “not known, the inPlorescence much mo an long. {. Upper scape-bracts nearly equaling to exceeding the inter- nodes (scape unknown in D. vestita and D. vir abkey I I [- Upper scape-bracts much shorter than the coe 8. Inflorescence compound; pedicels not over 7 mm pag Subkey IL 8. Inflorescence simple (or few-branched and probably simple at times 9. Filaments Pee above the common tube with the petals. Subkey Iilt 9. Filaments connate above the common tube with the petals. Subkey IV Subkey I compound. 2. Axes ae f fosat} of the inflorescence densely lepidote or tomentose -lepidote 3. Pedicels to 15 m Longs floral bracts exceeding the sepals. Beaetes MOS GOPNIG os te 7. D. princeps 3. Pedicels not over Fy mm long; floral bracts serrulate. 4, Lowest floral bract e fl ured by o. dense tiaete indu- 1s ae —. the 15 mm lo af sepals. Gera D. ursina 5- Margins of the pomeicee Bein ad visible 6. Flowers subsessile; sepals mm long, aan abate obtuse; indument of the Fete Pa ence fulvous 1967 Smith, Notes on Bromeliaceae 467 PEAR VAY S25 a's! Mulia ie. ale, Haw ein ewe whe alee oe9s D. ae 6. Flowers on distinct aeiinels ee -5 mm Lanes ‘inten to inflorescence ferruginous. 7. Sepals strongly incurved at apex, mena to 10 mm long. Brazil: Minas Gerais, Santa Catar -10. D. monticola 7. Sepals straight or nearly so, 5-8 mm ep ing a. — at least wee 6 as long as broad, acute, cari- Brazil: Par waned ie or Be cee ait than twice as long as broad with a broad apex, ecarinate. Zil: Sao ve to Rio Grande do « Ds ee 4, es floral bracts equaling or pe cepa the flower tals carinate 9. past 8 mm Longs leaf-spines 3-5 mm long; edcaaieranines usually much branched. Brazil: SAo Paulo to Rio Grande do SUL ds5 tees owes 12. D. act iotias 9. Sepals 11-12 mm Loki; r the leaf-spines less than 2 m long; inflorescence fev-brancheds petals saacaly if at Tia al nat is 10. Tnflorescence siiernios eg leaf spines 2.5-3 mm et oes 13. D. cinerea LO6 inflorescence texruninoas-lepelote: “Jeafaspines 1.5-2 mm a Be Filaments free above the common tube; style very short. 14. D. trichostachya 11. Filaments connate = oe A mm ea the eae tube; style 1/3 the length o Pore Boe 2. Axes of the inflorescence depeche nti ae "appressed - lepidote to glabrous le. issn acute, atiecng ls incurved, lies mm long. Brazil: — tearing sss ess -»-16. D. cabrerae le. eee broadly ro oniled; eee or nearly so, 6-7 mm long. 13. Sepals ropa ay peas than wide. Brazil: Minas Gerais. Lis D- la 13. Sepals deltoid-ovate, as wide as long. Brazil: Gola 18. D. etki 1. Inflorescence simp 14, Axes (at eee of the inflorescence densely lepidote or tomentose -lep 15. Leaf-blades pach a “lepidote on both sides. 16. sag bracts ample, covering most of the sepals; pedicels ort and obscure; filaments free above the common tube. LT. a aceon villous; floral bracts entire razil: Rio Grande do Sul. ‘ ~-19. D. choristaminea 17. erenapey: lepidotes “floral ‘beens serrulate. Brazil: s Gerais. . eeee020- D. simulans 16. Floral eon eatecus ceklania "distinct; " Pilaments ; connate above the common tu 18. oo nearly 50 cm see, the got 30 mm wide, laxly errate; sepals 11 mm long. Brazil: Minas Gerais. 13. D. cinerea 18. Leaves 9-17 cm long, the blades 8-10 mm wide, re repand- 1,68 PHYTOLOGIA Vol. 14, no. 8 serrate; sepals 6 mm long. Brazil: Parand. D. fosteriana 21. 15. Leaf-blades soon eke above or on both sides. me Vi ah ag eee floral bracts ee filaments free above the ommon tube. Uruguay. Southern Brazil. ee. D. aa 21. eeibinns floral bracts ee filaments connate abov common tube. razil: Minas Gerais to Santa poaiine t cc eeenee 23. D. minarum 20. Inflorescence Diididiea Aacaeati ite e2. Floral bracts entire; sepals 15 0 mn long, > te margins cbecured by the dense lanate indument. azil: Minas b wales . : ath il D. ursina 22. Forel pacts Sata. se la “8-1 mm long. 23. U. cape -bracts end: lower yes bracts with a blade several times longer than the base; torso very lax at base; sepals fimbriate. Brazil: Minas Gerais eeeee 04. D. duarteana 23, see bie agora and floral nance with narrowly ak. Floral bracts Jsree, exceeding most of the flowers; cels 5 mm long, stout. 25. tesa sepals alate-carinate; filaments free above the common tube. Brazil: Rio Grande do 25. D. irmgardiae 25. Posterior sepals carinate but not alate; filaments onnate above the common tube. Brazil: as Gerais, Pa a, Santa Catarina 26 D. reitzi 24. Floral bracts shorter than the flowers; pedicels to 10 long, slender. Brazil: Parand. 27. D. hatschbachii 19. Sepals broadly r oo or subacute, obtuse or apiculate 26. Lower floral bracts ma ally 20-25 mm long, equaling o or m tube; se -blades sparsely steiteisrd a Ame eihdkts. steal Goids De eniriens Ag oo connate above the m tube; Leei-biades densely einereous-lepidote, nar ti a : Mina EBit oakk sy uals es br s tanta 26. nae — tenebel ahegitie: ae than 20 mm pat or exceeded he sepals or both. 28. deve. and lower floral bracts serrulate. 29. Leaves 5 cm long. Brazil: Minas Gera 29. D. densiflora 29. Leaves See long. 30. Floral bracts pes ee flowers erect or suberect; near very short. Brazil: Minas Gerais De lagoensis 30. 30. Floral bracts ovate or lanceolate; flowers | spreading; 1967 Smith, Notes on Bromeliaceae 469 style 1/3 as long as the ovary. pot on Sao Paul to Rio Grande do Sul....-eeseeee D. pa 28. pages and a floral een wine: or very D. 31. Petals glabrous; plant 3-5 dm high; Lear< -spines to 3.5 : lower floral bracts equaling or exceeding the sepals. 326 eibaeties free above the common tub 33. Sepals 8-12 mm long. Uruguay. athe Brazil De renotiflora Gro 33. Sepals 6-7 mm long. Brazil: Goids, Mato aes D. Neate 32. Filaments connate above the common tube ; sepals ae a ! mn lo Brazil: Minas Gerais....33. D. schwackeana 14. Axes of an ase oor gag glabrous or sparsely and fugaciously lepido 34. Leaf-blades very fin ou and regularly white-striate a o the inclusion of the scales in the deep d and thick. Brazil: Minas Gerais ne revolts iethadines broad Sant. bea bes oo 3te 34. Leaf-blades not requir semin te henentlss the ce mor or less covering the nerves. 35. Scape-bracts all with linear blades; entire; flowers 9 mm long, half as long as the inter- nodes. Brazil: Goids, Mato Grosso (?). leaves entire or sub- + De burchellii 3 35. Scape-bracts diverse, the upper ones pladeless. 6. Sepals triangular-acute or acum uminate, carinate. 37. Floral bracts ample, exceeding the lower flowers; inflo- rescence dense toward apex; pedicels stout, to 5 mm long; filaments connate. to 16 mm long; indument of in- lorescence whitish. — Parana...36. D. susan 38. Po ner alate, 8-9 mm long; indument of i cence peas de Brazil: Minas Gerais, cand, ta na. és ey ROH Ds reitzii 37. Floral bracts naerosts infloreecence ex} sepals incurved bracts span: fila- e common tube. Brazil Minas Ge Catarin 39. Scape-bracts snd floral sunaes en = laments scarcely connate above azil: Santa Catarina....-cserceesees 6 36. beast reas rounded or obec: obtuse or apiculate. 4O. Scape-bracts and lower flora 1 eecks serrulate. 41. Filaments connate prone the c be he, Horst bracts ample, the lowest simacdind the flowers, razil: Distrito Fede mrad Gk tek Shea brasiliana 470 PHYTOLOGIA Vol. 14, no. 8 hO, Floral bracts narrow, the lowest about ee the sepals. Brazil: Rio de Janeir 38. ee 41. Filaments free above the common tube. 43. Leaves 7-8 cm long. Brazil: Minas Gerais. cmmhcaden tee 20. D. simulans 43. Leaves 25-50 c¢ hh. Pe long. tal-blades ecarinate, suborbicular; lowest visio rs equa or exceeding the flower BS GRrAte ia Sua chs -l4. D. ate yy, Petal-blades PLE a carinate; Lanne is apt bracts about equaling the sepals. razil 39. D. silvae ho, ——-- and floral bracts entire or nearly | so. 45. Lower floral br: e. Paraguay. “ho. De vizgste 45. Lower floral bracts equaling or nematinn the er 46. Filaments connate above the common tube. Braz Minas Gerais euele atten D> See 46. Filaments free above the common 47. pose oe tube. racts michuned ovate and sesh U Brazi wee jangular lade Brazil: Goids, aha iw eae eee < D. tenuis about as long as the base. Grosso ee ee ee ee a Subkey II 1. Axes (at least) of ae inflorescence densely lepidote or tomentose-lepidote. 2. Margins of the se paibae obscured by the dense lanate indument; petals barely exceeding the 15 mm long sepals. Brazil: auasee sceeevceesss D. ursina 2. Margins of the sepals clearky: visible e. D 5. recat. steataet or pate so, 5-8 mm long. ‘breil Parana D. frigida kh. Sepals broadly subacute or rounded, entire or cot aioe - Scape-bracts entire. Brazil: Mines Gera as D. weddelliana 6. Scape-bracts serrulate. T- Floral bracts broadly ovate and npaeureernt barely exceed- ing the lower pedicels. Brazil: Bahia 2. D. maracasensis 7. Floral bracts narrower, ac much snansing 4 the pedicels. uminate, Brazil: S&o Paulo to ia ae o Sul. 2. D. hd a 3- Indument of the inflorescence SS or fulvous 1967 Smith, Notes on Bromeliaceae 471 Argent Paraguay. Southern Brazil 8. riladents | free above the common tube with the petals. 9. Style ati equaling the ovary. Paraguay. Argentina: Mision Brazil: Santa Catarina.......43. D. distachya 9. Style Oey & sho 4 10. Floral bracts apiculate or ssa flowers lax; anthers included. Bolivia. Brazil: Mato Grosso. Paraguay. gentina: Formosa, Chaco PEs. ecee ett. D. ferox 10. Floral bracts sigankivate: flowers dense; sortase somewhat exserted. Argentina: La Rioja, Jujuy, Tucum hs. D. Pidiaelibn 8. Filaments connate above the common tube with the petals. 11. Sepals 11 mm long; style none; oe bracts acuminate, equaling the sepals. Paraguay....-- +29. D. vestita 11. Sepals 7 mm long; style trifid, "2: 5 mm “long; floral bracts apiculate, shorter than the sepals. Salta el ae ppeeae ve NG. ch 1. oes of the iielerdaciridis ‘pigtvdus or Aagiy and nee ote. 12, Sepals triangular-acute to acuminate; 7 free above e tube with the baat or nearly Brazil: n shes ines oss oe Ss. « oe ee eae = . D. cabrerae 12. Sepals trey: eros ie “eetuas: entire or e apiculate ° 13. Filaments free abo he common tube with the foe 14. Floral bracts exceeding the flowers, acumina Paraguay. D. velloziifolia 14. Floral bracts shorter than the sepals. 15. Floral bracts much ——— the lower pedice 16. Style well developed, di LT. stinct, ca 2 mm pee igh resem parplines sepals 3- Pa ong. Para- rana to piece: D. microcalyx 17. branches nisin gone sepals 6-12 mm long. 18. Sepa ehissigins more or less carinate, only about half as tong = e petals. Southern Brazil. raguay. -49. D. leptostach 18. Peary on ee pereei; wot “enpinslte; “distinctly more than half as long as the petals. 19. Leaves very laxly serrate with nearly straight vals. Brazil: Minas Gerais...50. D. mello-barretoi 19. Leaves sinuate-serrate. Brazil: Santa Catarina. 51d. 16. Style short (not over 1 mm) o 20. Floral bracts acuminate; ones eps ibiramensis 20. Floral bracts apiculate Paraguay - D. affinis 21. Rhachis ered very slender; flowers more or les secund. Boli i D. pulquinensis 21. Rhachis not more tesa lexuous; flowers not oboe 22, Anthers exserted. prong " Cérdoba WP ES A ers hee ce es ey 22. Anthers incl San Iuis h, ae ee uded . 23. Style lacking. Bolivia. Paraguay: .- Brazil: Mato 472 P.HYT0.4.0G-1-4 Vol. 1h, no. 8 Grosso. Argentina: Formosa, Chaco, Santa Fé. . 44. D. ferox Paraguay. - D. commixta 15. Floral . about equaling to shorter than the lower 23. Style short, 1 mm, but distinct. pedic ok, Petal-plades itn gent ~~ almost none. Brazil: Ce « Cer ek 8 - D. _otobanchotdes Hig 6 etal Mequiasers« «057. De tobatiensis 13 Pa aco Sonate above the common tail wih the petals 25. Anthers included. 26. Pedicels to 5 mm Jonge mostly exceeding the floral bnac ts Brazil <(Mato: Grose0is wieese cess S0e5es De inii 26. Pedicels very ets pata esoeiod by t ‘his floral Mr in Brazil: Minas Gerais ieee aiele& ole ace --59. D. rariflora 25. Anthers exserted. 27. Style 2.5-5 mm long. Argentina: Misiones. O. D. niederleinii 27. Style 1 mm long. Argentina: Santa Fé, Santiago de BEORO sy pass tae wre wee Wa asia exe --61. D. ragonesei Subkey III 1. Axis (at least) ie Ae inflorescence densely lepidote or tomentose -lepid 2. Indument of the inflorescence ferrugin 3. Margins of the sepals obscured by the oe lanate indument; petals . saat At, the 15 mm long s 1s. Brazil: Minas G 3. Margins of a sepals clearly widibie 4. Upper scape-bracts ower floral prac ts with a linear blade several times longer than the base; i fimbri- five po nesuaee Decureina ate. Brazil: Minas Gerais 4. Upper scape-bracts with triangular blades. 5- Lower floral bracts and upper scape-bracts entire; petal- blades pacer pag ecarinate. Brazil: Minas Gerais. 62 . D. sordida 5- Lower floral bracts and upper scape-bracts serrulate}; petal-blades elliptic to gee carinate Sao Paulo to Rio Grande 2. Indument of duarteana Brazil: Liven etes. De encholirioides n e Sisiabadeeins | a. or white 6. Anthers exserted; style very sho Paraguay. 63. D. tomentella 6. Anthers es _— distinct 7. Flora ~—. mont of the Fue aa one thern Brazi D. renotiflors {- Floral bracts pe bE icin exposing mB or all of t sepals. Brazil: D. oligantha 8. Upper sp cart = lower floral bracts bi Para~ - Upper sca er se lower floral bracts a Minas 64 1967 Smith, Notes on Bromeliaceae 473 guay. Argentina: Misiones. Brazil: Santa Catarina. 43. D. distachya 1. Axis of the inflorescence glabrous or sparsely and ~fugaciously vestite 9. Lower robes bracts rounded, subacute, obtuse or apiculate. 10. Floral bracts from shorter than ie iy eg) exceeding the 3-8 mm ge ie sepals 7-8 mm 11. Pedicels 8 mm long; petal-blades goon ecarinate Brazil: Anas Gatnle: Seles A ASD “biflora 11. Pedicels 3 mm long. 12. Flowers arian petal-blades suborbicular. Brazil: inas iea5o. De a 12. Flowers vo lax: petal-bledes houbies Brazil: 66. D. aaaked 10. Floral bracts much ies sin the pedicels. 13. Style Sgernigies much mor an 1 mm long. 14. ssiek ae racts ample, peitioe most of the sepals. South- zil. Paraguay. Argentina. U Jv 22. D. remotiflora 14. taba bracts inconspicuous, exposing nearly all of the is sepals 15. Flowers dened sepals 3-6 mm long. Paraguay. Brazil: Parana, Mato GroSsO....++++eseeeeeee- 48. D. microcalyx 15. Flowers lax; sepals 6-12 mm long. 16. Sepals emarginate, eendake ceeaminte Vote: Northwestern Argentina.......+- Ci ca cide vetese OTs D. tweediel 16. Sepals obtuse, not ciabetnate, soon sen aee uthern Brazil. pete ee Boliv spay Nat A leptostachya 13. Style short, not over 1 mm, or non 17. Rhachis geniculate, ey slender; Piovars more or less secund. Bolivia ee te De pulguinensis 17. Rhachis not more that Wlenioa: Powers not secun 18. Anthers exserted. 19. Petal ebiadse carinate. Lage Cérdoba, San Lui Ta Rioja. a epitalate. 476 PHYTOLOGIA Vol. 1h, no. 8 23; ap an included; indument of the inflorescence erruginous; style 1.5 mm long. Brazil: Bahia ye. D. maracasensis 13. Anthers exserted. 14. Style 6 mm long. Argentina: Misiones ..66. D. mitis 14, Style 1 m long. Argentina: Santa Fé, es del Estero 61. D. ragonesei 1. Axis of the inflorescence glabrous or sparsely and ac Fe lepidote 15. Petals eosin a distinct claw and blade, narrowly elliptic; leaf-blades erect, 13-32 cm long, the spines nearl igmas subsessile. Brazil: Goids, Mato Grosso...87. D. horridula ile oo with a sharp dietinction between thie: narrow claw and bl broad blade. 16. Lower floral bracts acuminate. 17. Anthers exserted. Argentina: Santa Fé....61. D. ragonesei 17. Anthers equaling the petals or shorter. 18. Scape-bracts ‘ig ping serrulate 19. Pedicels 3-10mm 20. Sepals rounded, site or obscurely apiculate. 21; oo pectinate-serrate with flat spines 6 mm Brazil: Minas Gerais.........88. D. a pad Be eco with spines laxer, uncinate and not ove +5 tm 22. Pedicels not over 5 mm —— 23. Floral bracts ample, vering w of the sepals; leaf-blades ooo tnt and resins above. Brazil: Distrito Federal... .ccecescceeees37- De. brasiliana vering much if any of the sepals; leaf-blades densely —— above at least initially. Brazil: Minas ok. Style. nearly as aii as the ovary; leaf-spines 3.5 -69. D. argentea 2h. Style son ices leaf-spines 1 mm Lon ng- D. spinulosa oes — to 10 mm long; filaments almost — 1 connate. Brazil: Minas Gerais - macropoda 20. Se Ps acute; style very short 25. Posterior sepals a straight, convex. Brazil: Rio de Janeiro to Para 38. D. pseudococcinea 25. Posterior a henge inflexed and carinate. azil: URE ING 2 ois oes -+-16. D. cabrerae 19. Pedicels not over 2 mm mm long, very. short “end stou 26. slopes bracts exceeding - lowest —- and all the sepals. Brazil: Minas Gerais D. lagoensis 26. —— oe all aus by the sepals. Brazil: Rio e do Sul (7)..s.0.4. Be seitowlons 18, Pein entire or very obscurely ser ton 27. Flowers to 15 mm long; petals about twice as ee as the sepals. Brazil: Minas Gera rais.........91. D. warmingii 1967 Smith, Notes on Bromeliaceae 477 27. Flowers 8-13 mm long; petals much less than twice as long as the sepals. 28. Leaves about 5 cm long; nee een’ short; flowers 8 mm long. Brazil: Minas Ger 3 ‘consiuitts 28. Leaves much longer; inflorescence lax, akan 29. Flowers secund, very lax; lower floral irate bapily exceeding the pedicels; petals brown. Brazil: Bahia 93. D. secunda 29. Flowers not se 30. Leaf-blades 5 mm penn 10-15 cm long. Brazil: Goids. 94. D. stenophylla 30. Leaf-blades much wider and lon 31. Style 1/3 as long as the seery? “Sees broadly acute and apiculate. Brazil: Piauf, Bahia, Minas Gerais 83. D. dissitiflora 31. Style very short o 32. Petal-blade elliptic, ecesp than wide. Braz inas Gerais 186s sas saxatiiie 32. Petal-blade very bie sd obovate, tere: Bae long. Brazil: Goids..... --3e.95. D. aurea 16. Lower floral bracts rounded or “wronaly. aaauta, obtuse or apiculate 338 Se 7 mm wide as mostly subopposite spines 5-7 Brazi hag dds ioc ee D. dawsonii 33 ORL es wider and with much smaller spine 34. Floral bracts barely if at es oo the meter 4-6 long pedicels; style d 35. Leaf-blades densely lepidote Eee ee twice as long as the floral bracts. Brazil: Goi - D. racemosa 35D. beet views densely lepidote on both sides; apaicele slightly shorter than the floral bracts Brazil. - D. lutziana 34. Floral bracts much ge the pedicels. ar ey than i ice the floral hese style 45 ae ies “ si as vary. Bolivi itaeeesGss aveee De greciiia BT Serais ‘tcl suke or ‘pubreni iform, 5-7 mm = fone htly if at all exceeding the floral bracts; style a s long as the ovary. Argentina: Misiones. 38. awe "ok mm long, stout, angled. 60..Ds Se 38. Pedicels lacking s ecsaks? et 86. D. mitis 36. Style short, not over 1 mm, or none. 39. Co Cicvad tabi Eas mm long, from nearly peeps ra oe Ce ee So eee eee ee 6 4O. Floral bracts narrow, not covering the sepals. Brazil: Minas Gerais....+---++--0022-99+ De rariflora 478 a ims 6 Me a Vol. 1h, no. 8 39. athe floral bracts 4-5 mm see much shorter than the sepals; leaves 50-120 cm long 41. Flowers pedicellate for 3 mm. ‘Brageys gg atk D. paraensis 41. Flowers subsessile. Paneinyiw is 6 ew ees hoe D. virgata 1. °D; ICELLATA Mez in DC. = Phan. “9: a3 1896. Type: Schwacke shee rosin with a mre Dyckia is uncertain ecause of the lack of petals and stamens. The ape might be an Encholirium. re Ds Bereta Baker, Handb. Brom. 136. 1889. Type: Tweedie mophyllum maritimum (Baker) Mez in DC. Mon. Phan. 9: Soy ee Dyckia tonentoss Mez in DC. Mon. Phan. 9: 515. 1896. Type: Sellow Brom. Pa 3. D. SELLOA RS Koch) Baker, Handb. Brom. 136. 1889. Priono- phyllum selloum K. Koch, Ind. Sem. Hort. Berol. 1873, App. 4: 7. 74. ; llow c+ 137. Type: Sellow Brom. Paris 75. - myriostachya Baker, 1. c. Type: Sellow Brom. Pari Paris 76. 4. D. INSIGNIS Hassler, Ann. Cons. & Jara. Bot. Genve 20: 316 Feb. 1919. Type: Fiebrig “N65. D. insignis lusus flaviflora i ig 4867 insignis va Hassler, op. c Type: Fiebrig 4867. D. i r. macran Hassler, op. c. ; ae Fiebrig 5310-a. D. insig- nis var. obtusiflora Hassler, 1. Type: Fiebr a0. D. randiflora Mez, Fedde Rep. Bice 5 Nov. 16: 68. Nov. 1919. ‘Type: Fiebrig L615. ‘ - D. EXSERTA L. B. Smith, Rev. Argent. Agron. 8: 192, fig. 6 Rojas : 6. D. ODORATA L. B. Smith, Phytologia 10: NS5} phe, rie. 8, 5. on - PRINCEPS Lem. Jard. Fleur. 3: Pl. 224, 225. 1853. Type: T- Desssipitn & plates. - URSINA L. B. Smith, Arquiv. Bot. Estado S. Paulo n. ser. 72 109, ‘= 112. 1943, Type? Foster 636. hers peed Ann. Cons. & Jard. Bot. Genéve 20: 315. 1919. : Fiebrig 10. bg MONTICOLA Korg & Reitz in Reitz, Sellowia no. 14: 104, fig. 3. 1962. Type: Reitz & Klein mre er, Handb. Brom. 134, 1889, nomen in synon., quite possibly based on a descendant of D. frigida. nee D. ENCHOLIRIOIDES (Geud. Mez in DC. Mon. Phan. 9: 507. Ba Var. a. ENCHOLIRIOIDES. Garrelia encholirioides Gaud. A Voy. Bonite pl. 115. 1851. Type: Gaudichaud 130. mesioetiinn garreli Beer, Bromel. 27. 1857, nom. nov. kia catharinensis x; a Ind. Sem. Bee Berol. 1873, App. 4: 4. 187k. ‘Type: Geudichaud s.n. D. altissima sensu su Baker, Handb. Brom. 134. 1967 Smith, Notes on Bromeliaceae 479 1889, in part, as to Garrelia encholirioides Gaud. ? D. cathari- nensis Beet ~ dentete Wittm. Bot. Jahrb. 13, Beibl. 29: 17. 1891. k 456. Encholirion catharinense (K. ma Benth. & Hook. = ex a ae in > Engl - Pflanzenreich IV, 32: 320. 1935, nomen. Axes of the inflorescence and t the sepals yellow. Var. b. RUB 1g 13, Beibl. 29: 16. 1891. Type: Schenck 619. Axes of the inflorescence and the sepals red. 13. D. CINEREA Mez in Mart. Fl. Bras. 3, pt. 3: 469. 1894. Type: Glaziou 18570. 14.°D. TRICHOSTACHYA Baker, Handb. Brom. 133. 1889. Type: Sellow Brom. Paris 59. D. micracantha Baker, op. c- 135. Type: Sellow 2021 "15. D. BRACTEATA (Wittm.) Mez in Mart. Fl. Bras. 33 Ps 3: 470 1894. D. aiseteieeae var. bracteata Wittm. Bot. Jah Beibl. nih 16. 1891. Type: Schenck 16. D. CABRERAE Smith & Reitz in Reitz, Sellowia no. 14: 101, fig. 2. ie Type: Smith, Reitz & Sufridini 9262. 17. D. ELATA Mez in DC. Mon. Phan. 9: 508. 1896. Type: Schwacke 8739. 18. D. GOIANA L. B. Smith, sp. nov. A D. elata Mez, cui veri- Similiter affinis, Zioatia andes deltoideo-ovatis differt. PLANT flowering to 2.5 m high. LEAVES lacking (but judging from the lower scape-bract ay narrowly triangular, pungent, very d bot: serrulate. INFLORESCENCE laxly compound, 1 m long, finely cinereous-lepidote at first; primary bracts like the upper scape- bracts, to 7 cm long; main axis stout; lateral branches suberect, slender, sitantiy flexuous, to 5 dm long; floral bracts broadly t e Ww tinct, 1 mm long. Pl. I, fig. 1: Margin of nepes scape-bract x 1 fig. d: Flower x 13 ae 3: Sepal x 2; fig. 4+: Young stamens x cae fig. 5: Young pistil x 2. ee Goids Galeensy Cérrego Estrema, ca 42 km northeast of Form Serra do Morcégo, alt. 800 m, 20 April 1966, H. S. Irwin et al. “ines “tos, type: NY). ISTAMINEA Mez, Fedde Rep. Spec. Nov. 16: 71. 1919. : wee 20. D. SIMULANS L. B. Smith, Sets Bot. Estado S. Paulo n. sy gti 108, pl. 110. a Type: Foster 570- D. FOSTERIANA . Smith, ee Bot. Estado S. Paulo n. wee se 107, pl. 106. ae Type: Foster 1154. 480 FETT ODO rs Vol. 14, no. 8 2. D. REMOTIFLORA Otto & Dietr. Allg. Gartenz. 1: 129. 1833. a. (Sellow) Hort. Berol Var. a. REMOTIFLORA. D. rariflora sensu Graham, Bot. Mag. 62 pl. 3449. 1835; sensu Lindl. Bot. Reg. 21: pl. 1782. etd non Schult. f. 1830. D. rariflora var. "Ue remotiflora" Baker, Handb Type: Cunningham. D. vaginosa Mez in Mart. Fl. Bras. a oi ars 490. 1894, ie? Glaziou 154907. Upper scape-bracts and fitral bracts with broad Ba: summits; sepals cucullate, 8-10 mm long; aga 17-23 mm long. Var. b. MONTEVIDEN. irs (kK. ae L. B. Smith, Arquiv. Bot. Estado - Peis n. ser. 1: 108 D. ursitevidandia K. Koch ex Hauman & Vanderveken, An. Mus..Nac. Hist. Nat. Buenos Aires 29: 239. 1917. Upper scape-bracts and floral bracts with broad apiculate summits; sepals n nearly or quite straight, 6-8 mm long; Var. c. ANGUSTIOR L. B. Smith, Arquiv. Bot. Estado S. Paulo n. ser. 1: 108. oe idk Bornmueller B5t 5 E in Mart. Fl. Bras. 3, pt. 3: 483, pl. 91. 1894, Type: ‘Beanell 1 1-283. ah. D. L. B. Smith, sp. nov. AD. urs B. Smith, cui pears bracteis florigeris TERT ge aE distinctis, sepalis minoribus differt. linear blades, serrulate, all but the highest exceeding the i s INFLORESCENCE ieee ed with one-flowered branches, very lax, over 3 dm ong, densely ferruginous-lanate; lower floral bracts like the Bion Tess pedicels suberect to spreading, to 5 ae mm long. SEPALS broadly fig. 6: Margin of inner leaf x 1; fig. 7: Flower x 1; fig 2 RAZIL: Minas Gerais: Wet campo, km 137 on Estrada da Concei- &o, Serra do Cipd, alt. Sees m, 21 April 1955, A. P. Duarte 2749 RB va type; phot. US 25. D. IRMGARDIAF L. B. "Smith, Phytologia 13: sone pl. 7, Ties S. 3. cee Type: L. B. & R. Cc. Smith, I. Schultz & Oya n. 26. D. REITZII L. B. Smith » Anais Bo Bot. Herb. Barbosa Rodrigues 2: 14, pl. 1-3. 1950. rath Reitz 2690. 27. D. - B. Smith, sp. nov. AD. ememaen (Gaud . ) Mez, cui verisimiliter affinis » bracteis fPlorigeri conspicuis, pedicellis elongatis, sepalis acutis differt. 1967 Smith, Notes on Bromeliaceae 481 PLANT known only from fragments, flowering 1.5 m (! si a LEAVES to 85 cm long; sheaths cae i ous; bla (upper) slender, ferruginous-tomentose; scape-bracts imbricate, linear-triangular from an ovate base, serrulate. INFLORESCENCE simple, sublax dm long, many-flowered, ferruginous-tomentulose except the pétals; floral bracts narrowly triangular, the lowest a straight, slender, to 10 mm long. SEPALS ovate, acute, © mm long, carinate; petals 14 mm long, the blades subrhombic, curved- spreading; stamens included; filaments free above the common tube; styles 3, to 2 mm long. Pl. I, fig. ea Leaf-margin x 1; fig. 10: Flower and bract x 1; fig. 8 DS Es a Bg BRAZIL: Parand: Mun. Paranagud: Low woods os ee er 1 Nov. 1951, Hatschbach 2725 (US, type). 28, x ane ; Bot. Jahrb. 30, Beibl. 67: 5. 1901. Type: Glaziou 201 29. D. TEHTELORA Schult. f. in Roem. & Schult. Syst. 7, pt. 2: sco 1830. Type: Martius s. n. Ose Ds Aster Mez in Mart. Fl. Bras. 3, pt. 3: 483. 1894. we: Warming 2171. D. FERRUGINEA Mez in DC. Mon. Phan. 9: 533. 1896. Type: ein Ss. n. 32. D. TENUIS Mez in Mart. Fl. Bras. 3, pt. os waite 1894. Type: Gardner 3479. D. morreniana Mez in Mart. Fl. Bras. 35 ih 3: ho6. 18901. Type: Hort. Liege. D. kuntzeana Saf in DC. Mon. et: — o25; 1896. Type: Kuntze s. n. CKRANA Mez in Mart. Fl. Bras. 3, pt. 3: 478. 1894. SCHWA! a ‘Glaziou 1 8572. D. FOLIA Baker in Saund. Ref. Bot. 4: pl. 236. 1871. Hi mie Bert Kew. D. sulphurea K. Koch, Ind. —~ ort. Berol. 1873, App. ee 3 3. 1874. Type: Hort. Berol. D- riflora sensu Wittm. Bot. Jahrb. 13, Beibl. 29: 15. aes non acnuta. f. 1830. in Mart. Fl Rs Oa re oats 493. 1894, in part., non Lem. 1853. 2, oe "E. “Morr. ex Mez, Op. c- Loh, Type: Hort. Liége. . D. BURCHELLII Baker, Handb. Brom. 131. 1889. ‘Type: am ere L. B. Smith, sp. no A D. pseudococcinea - B oH ui affinis, bracteis ages amplis flores in- be ring to 1.5 m high. LEAVES numerous in a dense ro- » sheaths suborbicular, castaneous; blades wide, laxly retrorse-serrate with slen- labrou inely waiteslepidote, soon Ector one secape-bracts lance-ovate, larly longer and shorter than uminate, serrulate 482 PHYTOLOGIA Vol. 14, no. 8 ded; filaments connate abov e the omc oe style very short PL. 1, igs 3: mee ented x 1; fig. 14: Flower and bract x 1; pal RAZIL: piotrtid Federal: Campo cerrado, Brasilia, 14 N 1958, E. Pereira 4685 & Pabst 5011 (RB, phot. US); Kontad: § in teeeees mite nest, cerrado, Brasilia, alt. 975 m, 13 Nov. 1965, H. § Irwin et al. 10268 (us, type: > NY)? cerrado and gallery marrin, 3 km north of Sobradinho, alt. 1225 m, 9 Nov. 1965, Irwin et al. 10113 (NY). Very old material > Spa og referred here: Cerrado os: 5 km north of Planaltina, on road to S&o Gabriel de Goids, Goids, =e 36. D. PSEUDOCOCCINEA 1. B. Smith » Arquiv. Bot. Estado 8. Paulo n. ser. 1; 108, pl. 109, fig. q; 1943. Type: Foster 1144. 39. D. SILVAE L. B. Smith, Phytologia 13: 15%, pl cy, Fig. 4; 15. 1966. ‘Type: N. 7. Silva’ O. D. VIRGATA Mez, Fedde Rep. Spec. Nov. 16: 68. 1919. ‘Type: WEDDELLIANA Baker, Handb. Brom. 132. 1889. ‘Type: easel 258. l2. D. MARACASENSTS Ule, Bot. Jahrb. 42: 197. 1908. ‘type: Ule ce DISTACHYA Hassler, Ann. Cons. & Jard. Bot. Genéve 20: 308. it 1919. Type: Fiebrig 5648. D. distachya forma induta Me 4. D. FEROX Mez in DC. Mon. Phan. or S11. 38 Type: Kuntze S-n. D. meziana Kuntze, Rev. Gen - 3, Pt. 2: 302. 1898. : Kuntze s.n. D. hamoes Mez, Fedde Rep. Spec. nov. 3: 33. 1906. iebr kg Lecto ferox subsp. h a (Mez) Hassler, op. c. 30h, D. x f. hamo- 3 iS t5n Cas ice An. Mus. Nac. Hist. Nat. iacse Ties 37: . VELASCANA Mez in Mart. Fl. Bras. 3, pt. 3: 476. 1894. oe: Hisronmus. & Niederlein 66. a 7 oe 5 “An. Mu - Nac. ea Nat. Buenos ice te 373 es 1931. Type: ‘icine 8s. AT, r Mez, Fedde Rep. Spec. Bove "16: 70. 1919. Type: iaasler Eee 48.°D. MIC ome CROCALYX Baker, Handb. Brom. 133. 1889. ‘Type: Var. a. MICROCALYx. D- microcalyx var. inermis Hassler, Ann. 1967 Smith, Notes on Bromeliaceae 483 Cons. & Jard. Bot. Gentve 20: 307. 1919. Type: Hassler 8787. D. microcalyx var. micrantha Hassler, op. c- 30 Type: Fiebrig 5941. D. minutiflora Mez, Fedde Rep. Spec. Nov. a 67. 1919. Type: Fiebrig 5941. Sepals 3-4 mm long. Var. b. OSTENII L. B. Smith, Contr. Gray Herb. 104: 73, pl. 3, fig. 16. 1934. ‘Type: Osten & Rojas 9097. Sepals 6 mm long. ho. D. STACHYA or Chr + Ho: Kew. OD. Type: Balansa 4742. D. “egg ERTS Mez, ‘op; ec. 524. Type: D*or- bigny 858. D. hassleri Mez, Bull. Herb. Boiss. ser. II, 3: 134. 1903. Type: Hassler 3261. D. rojasii Mez, — Rep. Spec. Nov. 16: 67. 1919. Type: Rojas in Hassler 10884. - apensis rage op. it 3 ep Houas in Hassler 10500. D. remotiflora var. a ne sensu ith, th, Anais Bot. Herb. Barbosa Bariiiee es 45. 1950, not as to Seb? 50. D. MELLO- Ol. i. B. Smith, hei og 1 109,- pr. 1, fig. tig 1960. ‘Type: Mello Barreto 5 I SIS Reitz, Sellowia no. rset 104, fig. 4. 1962. Type: “Reitz & Klein 2635. 52. D. AFFINIS Baker, Handb. Brom. 133. 1889. Type: Balansa 53. D. PULQUINENSIS Wittm. Mededeel. Rijks Herb. Leiden no. 29: 88. ei gite Type: Herzog 1849. 4. D. FLORIBUNDA Gri Arg . Abh. 2h: 331. 1879. ‘Type: Hieronymus 566. D. gilliesii Baker, Handb. Brom. 136. 1889. Type: Gillies s. n- 55. D.C XTA Hassler, Ann. “Cons. & Bot. Gentve 20: 305. 1919. Type: Chodat dat & Vischer 97 € ae: D. OROBANCHOLDES Mez in Mart. FI. fea. 3, pt. 3: 4T5- Ss. ne - D. TOBATIENSIS Hassler, Ann. Cons. & Jard. Bot. Genéve 20: 58. D. IRWINII L. B. Smith, sp. nov. A D. racemose Baker, cujus bracteas en cpeait valde imitans, inflorescentia ramosa, stylo brevissimo differt flowering to 2. 25 m high. LEAVES 6-7 dm long; sheaths eet suborbicular, 4 em long, castaneous with pale margins; laxly serrate eageet scape -brac rowly tr iar from a broadly Sa wohtolincecuk, ase, much shorter than ih es rnodes, serrulate SCENCE laxly few-branched, 1 m long, finely sinavecus-leptiote, a ees glabrous with age; primary bracts +7 upper scape- : ALS broadly ovate 1 mm oie S tas least), the blade suberect, broadly obovate; stamens 48h PHYTOLOGIA Vol. 14, no. 8 included; filaments connate above the common tube; stigmas sub- sessile. CAPSULES ovoid, acute, 15 mm long; seeds with a broad ag al. 16923 (US, type; NY); eereans® aid abate margin, 75 ich south of Xavantina, alt. 525 a 17 June 1966, Irwin et al. 17340 (NY). RARIFLORA Schult. f. in Roem. & Schult. “Syst. fs pt. 22 u s. . D. NIEDERLEINII Mez in Mart. Fl. Bras. 3, pt. 3: 474. 1894 : Niederlein 2 ee p- D. missionum Mez,.op. c. 477. ‘Type: Niederlein 229-8. ps Ds messtonvm var. breviflora Hassler, Ann. Cons. & F aa Bot. Genéve 20: 316. 1919. Type: Hassler 142. 6l. - RAGONESET Castellanos, Lilloa 10: 454, fig. 2. 1944. Type: cuttatincs s. 62. D. SORDIDA ee Handb. Brom. 132. 1889. Type: Saint Hilaire 402. 3. D. TOMENTELLA Mez, Fedde Rep. Spec. Nov. 16: 69. 1919. Type: Hiebrig hohe. 64. LIGANTHA L. B. Smith, Arquiv. Jard. Soak Rio de Janeiro os 329, fig. h-l. 1958. Type: Duarte 2748. 65. D. BIFLORA Mez i n Mart. Fl. Bras. 3, oe e 86. 1894. 19919. - DUCKEI L. B. Smith, Bol. muss Paraense Emilio Goeldi n. ser. ~ Lt. 4, Sige Reo 1958. Ducke s. n. 67. D. TWEEDIEI Mez in Mart. Fl. Bras. 3 3, pt.3: 485. 1894. Type: neidtee S. n. . D. HEL HELOISAE L. B. Smith, a Misc. Coll. 126: 26, 65, fig. 16. 1955. ype: L. B. Smit Beta al. 66 69. D. ARGENTEA Mez in Mart. Fl. . 3, pt. 3: 492. 1894. aia ——. ee FOLIA Baker, Handb. Brom. 131. 1889. Type: ‘tien Bien ced i010. Mez in DC. Mon. Phan. 9: 530. 1896. Type: Saint Hilaire gek. ~~ 72. D. MARNIER-LAPOSTOLLEI L. B. Smith, fg oe Soc. Bull. 16: Type: Marnier-Lapostolle S&S. ne CEDOL ith, Arquiv. Bot. Estado S. Paul ger. 2: 195. 1952; “Suitheonia Misc. Coll. 126: 67, 68, HG: "8. 1955. Type: Maced 4 4. D. SICKIT Le = Smith, Arquiv. Jard. Bot. Rio de Janeiro 15: 330, fig. m-q. 1958. Type: Sick B-613. (5. D. PUMILA L. B. Smith, Phytologia 13: 151, pl. 7, fig. 13- ae Type: Irwin & Soderstron 7365. 76. D. in Mart. Fl. Bras. 3, pt. 3: 487. 1894. LO2, Bel 1966. INERMIS me Niederlein eo - D. GLANDULOSA L. B. Smith & Reitz, sp. nov. Ab omnibus wit ee adhue cognitis inflorescentiae indumento glanduliforme facile distinguenda. PLANT flowering 7 dm high. LEAVES to 23 em Long; sheaths sub- orbicular, 2 cm long; blades narrowly triangular, acuminate to an 1967 Smith, Notes on Bromeliaceae 1,85 abruptly acute pungent apex, 23 mm wide, laxly serrate with unci- nate antrorse spines 2 ong, covered beneath with cinereous APE bic, obtuse, carinate; stamens included; filaments free above the common tube; style 1 mm long. Pl. I, fig. 20: Leaf-margin x 1; fig. 21: Trichome x 100; fig. 22: Flower and bract x 1: fig. 23: 2 RAZIL: Minas Gerais: In smallsparse groups among rocks at the base of the Serra do Itambé, 19 March 1963, M. Magalh&es 18959 (Hb. Bradeanum 25726, type: phot. U 78. D. DELTOID FA (L. B. Smith) L. B. Smith, stat. nov coccinea var. amneae L. B. Smith, Arquiv. Bot. Estado S; “Paulo n. ser. 1: 107. 1943. Type: Dusén Last D. tuberosa var. = toidea na B. Smith) L. B. Smith, op. c. 2: 119. "79. D. TUBEROSA (Vell.) Beer, Bromel. 157. 1857. essa quberose Vell. Fl. Fluminensis 135. 1825; Icon. 3: pl- 1835 Type? Desort yao late. ag gocednee Mez in an Bras. 33 ele 1. 1894. low e. n. 23". TIANA L. B. ras Los gaat County Mus. Contr. h bes no. a ee masa a 1957. Type : Dawson 14153-a. 81. D. ELONGATA Mez in DC. Mon. Phan. 9: 529. 1896. Type: Sellow Brom. Paris 58. 82. D. MACROPODA L. B. Smith, sp. nov. D. duarteanam L. B. Smith (cf. supra) simulans sed bractets florigeris er haud lineari-laminatis, sepalis rotundatis, filament tis alte con- natis See rt. owering about 4 dm high. LEAVES 25 cm long; sheaths enti lowest exceeding the sepals; pene spreading, slenderly eylin~ dric, to 10 mm long. SEPALS bri ovate, rounded or apiculate, 7 mm long, strongly convex, the posterior more or teks carinate; petals 14 mm long, orange (! E. Pere ira), the blades broadly rhombic, carinate; stamens included; filaments almost wholly 486 PHYTOLOGIA Vol. 1h, no. 8 connate; styles 3, very short. Pl. I, fig. 25: Leaf-margin x 1; fig. 26: Flower and bract x 1; fig. OT: Sepal x 2; fig. 28: inas Gerais: Diamantina, Rio das Pedras, 29 May 1955, E. Pereira — (RB 90642, type; phot. US 83, D. DISSITIFLORA Sch wd Raat in Roem. "e Schult. Syst. 7, pt. 2: 1194, a Type: Mart - D. SAXATILIS Mez in ais fos Phan. 9: 518. 1896. Type: Schwacke 8948, e D. ULEANA Mez in DC. Mon. Phan. 9: 517. 1896. ‘Type: Ule Al Aires 36: 51. Pls 2.19290. : Spegazzini s. 87. D. HORRIDULA Mez, Bot. Salix, 30, Beibl. fr: 5 2, - D. PECTINATA L L. B. Smith & Reitz, sp. nov. A D. dawsonii th, cujus folias valde ag bracteis florigeris angustis, a inclusis differ PLANT flowering 75 cm high. ane 14 cm long; sheaths broad- ly ovate, 2 cm long, stramineous basally; blades very narrowly tr Ge; r, subulate-acumina mm wide, covered Boe pos sides with appr ressed cinereous scales, becoming more or rous above, subdensely pectinate-serrate with flat spreading spines 6 mm long. mm in diameter, glabrous; lower scape-bracts cies if at all carinate; petals 9 mm long, red-orange (! Pabst), the te, carinate; stamens included; filaments short- onnate above the short common tube; stigmas sessile. Pl. iy Pa. e9: leaf-margin x 1; fig. 30: Flower and bract x 1; fig. 31: Sepal x 2; ;fig. 32: Stamens x BRAZIL: Minas Gerais: In xerophytic habitat, outcrop of Trias- sic sandstone, between eng rege do Rio Verde and Cambuquira, 16 = 1957, G. F. J. Pabst 4129 (Hb. Bradeanum 3766, type; phot. US). D. SPINULOSA L. B. Smith & Reitz, sp. nov. A D. —, Mez, cujus inflorescentiam simulans, foliorum spinis parvis stylo a a rt. PLANT ong, covered with appressed cinereous scales on both — ber coming more or less glabrous above. SCAPE lateral, 4 mm wi sparsely and finely a Scape-bracts ovate, sve gi the lower dense and wi inear te shorter the Rebar ng simple, laxly * aes flowered, to 27 cm long, finely tie te at first; Y iatel 1967 Smith, Notes on Bromeliaceae 1,87 bracts like t e upper scape~bracts, serrulate, the lowest equal- ing the midpoint of the sepal; pedicels curved-ascending, slen- ALS ovate, rounded he common tub i ibsessile. Pl. I, fig. 33: Leaf-margin x1; fig. 34: Flower and bract x 1; fig. 35: Sepal x 2; fig. Stamens x BRAZIL: vee Gerais: Varzea da Palma, Fazenda da Mae d'Agua, bn Nov. 1962, A. P. Duarte 7409 (Hb. Bradeantit 27225, type: phot. 90. D. SELLOWIANA Mez in DC. Mon. Phan. 9: 520. 1896. Type: Sellow Brom. Paris 52. 91. D. WARMINGII Mez in Mart. Fl. Bras. 3, pt. 3: 481. 1894. Type: Warming s. n. 92. D. CONSIMILIS Mez in Mart. Fl. Bras. 3, pt. 3: 479, pl. 90. 1894. : Weddell 1407. 93. D. SECUNDA L. B. Smith, sp. n Ab omnibus speciebus adhuc mo ron inflorescentia sivieatis secundiflora, petalis brunneis differt. PLANT inverts 9 dm high. LEAVES o ver — em Long ; sheaths brous ab SC. subfoliaceous, the upper small, remote, tri ar, acuminate, Subentire. INFLORESCENCE simple, very laxly secund-flowered, to 23 cm long, sparsely and aciously pale-lepidote; floral bracts entire, the lowest distinctly exceeding the pedicels; a SEPALS ovate, rounded n mm cities . 3r ag ae f-margin x 1 fig. 30: bract x 1; fig. Bee Sepa) x 2; fig. 40: Stasens x 2; fig. 41: Pistil = 2, BRAZIL: Bahia: Drainage of the Rio agnor western Bahia, Campinas, ca 10 km south of Rio Piau, ca 150 oath sae Berrei- ras, 850 m, 13 April 1966, Irwin et al. 14723 grin type; phot. US oh. STENOPHYLLA Smith, sp. nov. AD. saxatile Mez ep lh Inflorescentiam simulans, foliorum laninis eaustianials diff T flowering - 3 dm high. LEAVES numerous in a dense glo- bose rosette, to 16 cm long; sheaths suborbicular, 2 cm long, brown; blades pbars fiaely 5 mm wide, laxly microscopically serrulate = acuminate, remote. ENCE simple, laxly few-flowered to 10 cm long, fugaciously white-lepidote; floral bracts like the 488 PRI70:20G6-I1.4 Vol. 14, no. 8 upper scape-bracts, the lowest equaling the sepals or shorter, microscopically serrulate; pedicels ca 1 mm long, stout but dis- tinct. SEPALS broadly ovate, rounded, 6 mm long; petals 8-10 m long, orange, the blades broadly elliptic; stamens included; filaments connate above the common tube; stigmas sessile. Pl. I, fig. 42: Section of leaf x 1; fig. 43: Flower and bract x 1; rig. 4k: Sepal x 2; fig. 5: Young petal x 2. BRAZIL: Goids: Mun. Cristalina: Rooted in soil-filled rocky crevices, local it~ common, 5 ~ south of Cristalina, alt. 1175 mn, 1 Nov. 1965, ir pein et al. 9728 (US, type; NY). 95. D. Al B. Smith, sp. nov. AD. saxatile Mez, cui affinis, Rest t laminis latissime sueiac differt flowering ca 1 m high (! Irwin). LEAVES to 23 cm long; eee suborbicular, 2-3 cm lo ng, stramineous; Liters strongly urved, narrowly triangula ar, acuminate to an abruptly acute ue apex, 3 cm wide, laxly aes with recurved pai eat ta mm long, finely lepidote in the narrow grooves beneath, glabrous above. SC lateral, 4 mm oan sparsely white-lepidote; scape- bracts broadly ovate, acuminate, the upper less than half as long INFLORE as the internodes, entire. RESCENCE simple, laxly many- flowered, 3 dm 1 long,- sparsely and fugaciously white-lepidote; floral bracts broadly ovate, acuminate, entire, the lowest ex- ceeding the sepals; pedicels subcylindric, stout, 4 mm long. 1mm long. Pl. I, fig. 46: Leaf-margin x 1; fig. 47: Flower and j E ri BRAZIL: Goids: Campo, 15 km west of Cristalina, Serra dos EAR alt. 1200 m, 6 March h 1966, Irwin et al. 13613 (US, ype; NY 96. D. DAWSONII L. B. Smith, Los Angeles County Mus. Contr. Sci. no. 17: 2, fig. 2-4. 1957. Type: Dawson 15236. 97+ D. RACEMOSA Baker, Handb. Brom. 132. 1889: Type: Gardner 98. LUTZIANA L. B. Smith, Arquiv. Bot. Estado S. Paulo n. ser. 1: ‘ict, pl. ge ag tg Type: Foster 1144-b. 99. D. GRACILIS in DC. Mon. Phan. 9: 516. 1896. ‘Type: D'Orbigny 1006 400, 0, ence L. B. Smith, Phytologia 13: 150, pl. 7, fig. 10-12. 1966. Type: Frdes 30 030. INDEX (by species numbers of Dyckia) ee maT affinis 52; "altissima" Baker 12; apensis 49; argentea * bifior ora 653 boliviensis 49; bracteata 15; brasiliana 373° bre- vifolia 34; burchellii 35. cabrerae 16; catharinensis 12; chaguar 46; choristaminea 19; cinerea 13; coccinea 79; var. deltoidea 78; commixta 55; consimi- lis 92; conspicua 49, 1967 Smith, Notes on Bromeliaceae 489 dawsonii 363 deltoidea 78; densiflora 29; sapere 83; var. bracteata 15; distachya 43; duarteana 24; duckei 66; dusenii 36. elata 17; elongata 81; eminens 28; encholirioides 12; exserta ferox 44; ferruginea 31; floribunda 54; fosteriana 21; frigida dar gemellaria 34; gilliesii 54; glandulosa 77; goiana 18; gracilis 99; grandiflora 4; grandifolia hamosa 44; “basslers 49; hatschbachii 273 heloisae 68; hilaire- ana "LS horrid eto apt» a) inblenké 4; interrupta 43; irmgardiae 25; irwinii 58. kuntzean lagoensis Ea leptostachya 49; linearifolia 70; longifolia 49; pire be macedoi 73; machrisiana 80; macracantha 3; macropoda 82; mara- aa "ho; maritima 2; marnier-lapostollei 72; mello~barretoi 50; meziana 44; micracantha ihe microcalyx 48; minarum 23; minu- tiflora 48; missionum 60; mitis 86; sietertleuis 22; monticola 10; morreniana 32; myriostachya 3. odorata 6; ods ha 64; orobanchoides 56. paraensis 100; pectinata 88; pedicellata 1; princeps Hort. 34; il princeps Lem. 7; pseudococcinea 38 se oat 533 a 75. racemosa 97; ragonesei 61;"rariflora” 22; "rariflora” dl rariflo Tet 5oe "reriflora" 3h rega- lis 11; reitzii 26; remotiflora 22; "var. ebitavidonsiet B. Smith 195 ee 49; rubra 12. saxat 4; schwackeana 33; secunda 93; selloa 3; sellowiana 90; seep: oc silvae 39; simulans 20; sordida 62; spinulosa 89; stenophylla 94; subinermis 76; sulphurea ah tenuis 32; he aang en tomentella 63; tomentosa 2; tricho- stachya 1}; tubero a 793 . deltoidea 78; tweediei 6 uleana 85; vaginosa 22; velnacics 4s; velloziifolia 47; vestita 9; virga- ta ho. warmingii 91; — rls se ENCHOLIRION cathar se - garreli le. GARRELIA encholirioies PRIONOPHYLLUM maritimum 2; “ee 3% TILLANDSIA abebcee 196 APPENDIX (Doubtful and excluded taxa, named hybrids) acaulis Baker, Handb. Brom. 137- 1889 = NAVIA ACAULIS Mart. ex Schult. f, in Roem. & Schult. Syst. 7, pt. 2: 1196. 1830. altissima Lindl. Bot. Reg. 27: 84. 1841. ‘he description: gri cord with " scapi sq not al that rhs be used as a lectotype in default of original mate- rial, so it seemsbest to keep this name in the dubious category. 490 PHYTOLOGIA Vol. 14, no. 8 ee hereon Dict. Gard. 1: 497. 1885 = HECHTIA ARGEN- TEA Ba n Hemsl. Biol. Centr. Am. Bot. 3: 317. 1884. ies Wena: Brom. 135. 1889. = CONNELLIA AUGUSTAE (Rich. 2 ae Ha E. Brown, Trans. Linn. Soc. ser. II, 6: 66, pl. 13. 1901 a — ee: Brom. 137. 1889 = NAVIA CAULESCEN. Mart Schult. f. in Roem. & Schult. Syst. 1, pt. 2: 1195. 1830 decom aie Baker, Handb. Brom. 136. 1889 = DEUTEROCOHNIA LON- PETALA (Baker) Mez in Mart. Fl. Bras. 35 Pt. 32-506, pl. 95» vdeo ane Baker, Bot. Mag. 120: pl. 7340. ae HECHTIA DES- TANA (Bak marae Mez in DC. Mon. Phan. 9: 551. 1896. uaa antea K. Koch, Ind. Sem. Hort. Berol. 1873, APP pe 4: 4. wal 2 nomen iites., wrongly attributed to Lindle De As used by D i unnecessary substitute name for D. prin- re got Mez equates it with D. altissima Lindl. as rhaps Koch a whip odo. I have found no material so namedand there laziovii pont Handb. Brom. 133. 1889 = NEOGLAZIOVIA VARIE- GATA (Ar ies: — ) Mez in Mart. Fl. Bras. 35° Wher 32 427, phar Bd, Tipe t ris —] Baker, Handb. Brom. 130. 1889 = ABROMEITIELLA BRE- VIFOLIA chit 2 ) Castellanos, An. Mus. Nac. Hist. Nat. Buenos Aires 36: 371, pl. ey 34,6. 1931. x Lad Cutak M. B. Foster, Bromel. Soc. Bull. 5: 35. 1955; 9: 61. 1959 = x LEPTOSTACHYA. laxiflora Mart r, Handb. Brom. 134. 1889, nomen in sy- but material of Ackermann from Minas Gerais, Brazil in Brussels, has serrulate floral bracts and can scarcely be the same. It may well be an er action. ~ at oh oma = no. 97: 5. 1874; Mez in Pflan- —— IV, 32 abe; "193 1 tala Baker, Hand gS 135. 1889 = DEUTEROCOHNIA LON- on A (Bake Baker) Mez in Mart. Fl. Bras. 3, pt. 3: 506, pl. 95. remosa Hort. ex K. Koch, Ind. Sem. Hort. Berol. 1873, App. 4 4. 1874, nomen fey synon. Equated with D. altissima vo and D. which are not the same, its identity therefore is in. wholly un pp obectabile Baker, Handb. Brom. agi 1889 = ENCHOLIRIUM SPECTA- . 2 ILE Mart. ex Schult. f. in Roem. & Schult. mat. T, ge. 2: 253° 3 ese real Handb. Brom. 135. 1889 = ENCHOLIRIUM SUBSE- oe Mez in DC. Mon. Phan. 9: 540. 1896. = Morr. Belg. Hort. 27:354. 1878; in Pflan- PR imag ye 7 Es 340. 1935. Judging from the scanty description and area of origin, this is probably a species of Dycki 1967 Smith, Notes on Bromeliaceae 491 Plate I D. macropoda; 29-32: D. pectinata: 33-36: D. spin- ulosa; 37-41: D. secunda; 42-45: D. stenophylla; 46-49: D. aurea. ITINERARY OF WILLIAM JOHN BURCHELL IN BRAZIL, 1825-1830 Lyman B. Smith and Ruth C. Snith Thanks to the cage ont of the Bric and staff of the Royal Botanic Garden » England, it was possible to consult However, the problem is quite different, because Burchell did not publish ny account of his collecting nor indicate localities or even states on his labels. Urban, in the introductory volume of the Flora Brasiliensis, s dication of gen ae: areas, but Burchell's collections deserve better treatme Fortunately, his numbers are chronological and thus can used as a guid d . e have made a list of the numbers and the dates and ee? at which these numbers were collected. Armed with this list, we have consulted the Millionth Maps of Hispanic America compiled and drawn by the American Geographical Society of New York, 1940, SS at The Smithsonian and Map Division of the Library o Congress. We also found old ae of modern locations and other information in the Enciclopédia $ Munierpice 2 fe published by the Instituto ed cp ag e Estatistica. In this way we have located a large tae. +g a places a ich Mr. and cl together to plot out his itinerary with a reasonable amount of sting the numbers, their dates and the localities by lstitude ih longitude as fox as we we m the old details that may clarify ae Seanad) Our s are in parentheses and everything else after each number is directly quoted from Mr. Burchell's is books - Burchell started his collections within the city of Rio de Sa the then national capital, now part of the State o Guanabara, with the numbers 701 - 1103 between 16 August and 29 September. of 1825. TOL. = » 16 Aug. 1825, along the aqueduct a pile and half above — convent of Santa Teresa (in the old part of Rio, S 22° 5 W 43° 10-12 é Bog - 818, Near Métopdércos, on stems of orange trees (in old city on northern rn slopes of Morro de Santos Rodr peda Aas of Gloria). 492 1967 L. B. & R. C. Smith, Itinerary of Burchell 493 819 - 897, 8 Sept. 1825, along the aqueduct, as before 898 - 991, 15 Sept. 1825, along the aquedue+ from the Convent of Santa Teresa to the spot where it crosses the ridge in sight of the Valley of Laranjeiros (as) (S 22° he w 43° 11-12"). . 992 - 1007, Near Lagéa das Freitas on the road between Botafo- go Bay and the Botanic Garden (S 22° 57’, ) Nov. 1825, on the Ilha do Governador (in ae nabara Bay, S 22° 47-50', W 43° 10-15". ‘This completes the 1105 series). The numbers 1106 - 3040 conclude the Rio de Janeiro and 49h PHYTOLOGIA Vol. 14, no. 8 vicinity collections. 1106 - 1152, 6 Dec. 1825, on the Corcovado Mountain, on the summit and in the 5 es, which clothes the highest part of the mountain (S 22° 57', 3754 9%) 1153 - 1180, mee the more open part of the road, from the forests to the cae of the Aqueduct. 1181 1192, Along the upper end of the Aqueduct as far as the spot wee the an chads x ve the northern side, and where the collection of 15.9.25 en ea 1250, 19 Dec. jan sine the Aqueduct as on 8 Sept., no. 19. 1251 - 1273, 24 Dec. 1825, along the Aqueduct mostly on the left or SE side and the thick woods 1274 - 1283, (Same day, on the Saiie hill) quitting = aque - duct by the first opening or va ley just above ise stone quarry. oes 1291, (Same day) above the stone qua 1eg92 - 1371, 29 Dec. 1825, Barros Vermelhos ahs Rio Comprido (S 22° 29 eg W 43° 10% 1372 1hé1 dite Dec. 1825, on the Corcovado Mountain by the way of hirartel 1462 - 1487, 1 : Fei. 1826, about Catete, near the bridge and Praia do Flamengo (both sider S 22°55-56', w 43° 10°). 1488 - 1564, 10 Jan. 1826, in a walk from eatin Grande (Nic- —— or Niteroi, S 22° 53', w 43° 08") to the village of Sao Joa Tearai, part of Niteroi, § 22° 55', W 3? 08") ae ss miles fi 1565 - 1655, 18 =a 1826, in a walk to the village of Sao Christovao — of eres = gh Nacional is soeatal, S 22° 5h", W 43° 13") and Engenho 1656 - 166) 22 Jan. 1826, at poss "(8 20°55-56", W 43° 10") near the bridge. 1665 - 1668, 26 Jan. 1826, Barros Vermelhos. 1669 - 1718, 26 Jan. 1826, Mo Morro da 1719 - 1731, Catombi (Catumbi, east side of Morro Santos Rodrt 32 = 1862, Along the Mons of Catumbi 1732 - 1760 Aqueduct 1761 - 1812 ana rage of a aah untain on the NW side and along the aqueduct 185 » Frexal, or Frechaes (S. Aleixo? S 22° 33", W 43° O4F) between Majé (Mage S 22° 397, w 43° 02") and 18). s Org&ios about the head of River Paquequé uequ Sallie aes dwelling house (near Teresopobis, S 22° 26", W h2° sor), 99T - 2004, 8 Feb. 1826, northward to the Mule C Curra. e Horse Cur Curral and Milho ey (probably on Mr. March's eneraeey 55 - 2140, 11 Feb. 1826, mountain in front of Mr. March's e1la - 2158, 9 Feb. a River Pacaque by Mr. March's house. 2159-2213, 13 Feb. 1 (Same location). tion 2214 - 2366, 15 Mai esha 1 1/2 miles south of Mr. March's 1967 L. B. & R. C. Smith, Itinerary of Burchell 495 ouse. 2367 - 2hh1, 20 Feb. 1826, River Pacaqué near Mr. March's alo - 2488, in Feb. 1826, 1 1/2 miles north of Fazenda de Pacaqué - March 2489 - 2506, ss date and place) Bec 2507 - 2575, Forest, descent from Serra dos peer to Frechal (S. Aleixo? S 22° 33', w 43° ol 2576 - 2584, Foot of mountains Frec és 2585 - 2704, 25 Feb. 1826, Frechal to eat (S 22° 39", W 43° 2705, - 3 E133 28 Feb. 1826, two miles down Rio Magé from Mage, om 2136, - - 2743, dl Feb. 1826, Town of Mage. 2Thh - 2777, 1 Mar. 1826, up Rio em, to Fazenda da Niteroi, S 22°" 53°, W 2833 - 2917, Me Sy $6 the Island of Boa Viagem (part of i 2918 - 2935, Ch eat 11 June 1826, Catete (S 22° 55-56", an 2936 - 2976, 10 June 1826, Valley "bagi spice 297TT - 3006, Catete and suburbs 3007 - 3028 17 Aug. 1826, grates er gety Mountains (S$ 22° shows 3029 - fet "20 Aug. 1826, between Catete and Botefogo Bay. ell goes to the area of Santos where finishes his 3 3 - 3082, 16 Sept. 1826, in the fore at close above the feiate ce Monastery at Santos (s oe 57', W 46° 19"). 3083 - 3108, 20 Sept. 1826, same . - 3152, 2h Sept. 182 re level ground westward of os towards Cubat@o (S 23° 53t, | - Ge > 25' ry. bet - 3182, 26 Sept. 1826, to Chapel of Nossa Senhora de Montserrat. 3183 - 3204, 29 Sept. 1826, at Sentos in the 3205 - 3226, 1 Oct. 1826, at the Barra along hee ae of the Bay, thence t oods back to the Town o Z T - 3261, 18 Oct. 1826, Outeirinhos (a "small elevation of land on the left bank of the River pe front of Morro Jupu- 262 - 23312 22, es “Oct. 1826, Santos to S#o Vicente fe 2?” = 58", w h6° 3313 - 3361, "95 Oct. 1826, in the forest under the Mountain (336 5) see - 3386, 3 Nov. 1826, on Montserrat (in Santos on north- 96 PHYTOLOGIA Vol. 14, no. 8 east part of Island of S&o Vicente). 3387 - 3411, 9-11 Nov. 1826, near be Benedictine Monastery in the chacara where I resided (in Sa. 3412 - 3439, 12 Nov. 1826, forest adore Benedictine Monastery. 3440 - 3450, 16 Nov. 1826, (same locality). 3451 - 3461, 1 Dec. 1826, (same ar ha 3462 - 3464, 2 Dec. 1826, (same locality). Eni of Bound Volume I und Volume II, with numbers Pei: - 5336, Mr. Burch finishes in Santos and ‘eagandna in rear seas its cephere and starts north in S&o Paulo toward obs Gera 462 - 3483, 8 Dec. 1826, near Cubat&o (s. 23° 53", W46° 25') at Rio das ie ay 23° 52, W 46° 28') in Mr. Smith's Citio and immediate vicini 3484 - 3514, 9 i: 1826, oad Ze sylvis. 3515 - 35 Uh, 10 Dec. 1826, 3545 3594, 13 Dec. 1826, in a thé lower part, re aye of the Bae up the Serra de Cubat%o (s 23° id A hee 3595 - 3628, 15 Dec. 1826, at Rio das Pedr 3629 - 3679, 16 Dec. 1826, in the tices about 2/3, of the Ascent up the Serra de Cubattio: 3680 - 3725, 22 Dec. 1826, in the upper mei of the Ascent up the Serra de Cubat@o - summ it or Pico da 3726 - aS, 22 Dec. 1826, from the Pico. i Seria 1 1/2 miles toward S&o 3824 - Pe Sy Dec. 1826, 15 Jan. 1827, about Rio das Pedras and Cubata 3865 - 3685, 17 Jan. 1827, | summit of Serra de Cubat&o to the village of So Bernardo (S 23° 42", w 46° 33! 886 = 3 18 Jan. 1827, about the wiliaes a f S&o Bernardo. 3985 - 2080, 20 Jan. 1827, Sfo Bernardo to oho. Paulo (S 23° 3 33", W4 ho80-2 - 4080-10, Jan. 1827, sé 0. 4081 = 4153, 3 Feb. 1827, Sdo eee about 1 m. NW by N from the oe de oe as far as the Caveira Santa, in the —— dry shru ee - ges 10. Feb. 1827, at St. Paul's for about 2 miles along the Rio de Janeiro 4210 - he78, 15 Feb. 1827, along the road from the Tanque de Zunica to the Convento da Luz (in the northern part of the city, abit east of the center) and thence along the Sta. Anna road goes due north from So Paulo) over the River Tieté Viekiad: across the northern part of Sao Paulo, the crossing at Sta. Anna 7 oe oy pate W bye ror) cde a few huts which are about 3 miles “tere - “1218-6, lere - eae = Paget 1827, vicinity of St. along the S. Amaro road. "ie: - forte (agag). 22 Feb. 18er, "Otc Pauls. uoethward 2 miles towards Agoa-Branca ca). 4320 - 4384, 4 Mar. 1827, at eoreus (Marumb{ is the district on the outskirts of modern S@o Paulo where a great new football 1967 L. B. & R. C. Smith, Itinerary of Burchell 497 stadium and buildings for other sports bi located), over the hilly pastures east of the House. (Northwest me of city how): 4385 - 403, 6 Mar. 1827, Morumbe Road NNW. 4uok - 4437, 7 Mar. 1827, Morumbe, to the Porto SSE. 4438 - 4452, 7 Mar. 1827, Morumbe, walk to the Porto. Hous 4550 - 4588, 14 Mar. 1827, Morumbe, to the Flint Hill “and thence to the Iron Stone Hill. 4589 - 4626, 16 Mar. 1827, Morumbe to Sao Paulo - to Rio dos Pinheiros 4589 - 4616 (a river starting in the southwest of the city, flowing north S the Tieté hée7 - 4681, 4681-1 - 4681-63, "4682 - 4683, 24 Mar. 1827, Praga da Megria, at sroucheis, west towards Agua Branca about 2 m from Tanque d 4684 - 4705, “bT5=1 - 4705-17, 4706 - 4712, 1 Apr. 1827, S. Paulo - Santos s far as church de Nossa Senhora da Gloria. 4713 - i721, gti . 1827, Varge do Carmo by Rio Road. 4722 - 4728, me, ‘ 4729 - 4736, 25 Apr. 1827, Same. 4737 - 47hh, 3 May 102%, Same. 745, Tanque de Zun LTh6 = ye 9 — 1827, Tanque de Zunica to the ae eee (Northern S&o a bit east of the center of the c ghee keh - Week ¢ ear 1827, Tieté to Sta. Anna (S 23° 4766 - 1766-2, 11 May 1827, "Boa Morte" in ripa rivuli Tamandaty (‘Tamandatet, a river which goes north northwest through the city into the Tieté - it later became a ca nal). 4767, 29 May 1827, Ponte do Marchél 4768, 6 June ar "Caveira Santa 4769" - ee . Paulo. 7 4TTT, 23 Juné 1827, Ad Jaragué (a town and a easier h! ‘pias west of So Paulo, S 23° 27', W 46° 45"). 4778 - 4779, 10 July 1827, Morumi 4780 - 4781, 19 July 1827, Tanque a arvalho 4782 - 4785, 25 July 1827, Rio Tieté at Ponte "do Coronel. Anastacio. say is og ert e ape travels pa oe S&o Paulo Belém unis. mee, 6 Saly grey “6 ie! Site do Coronél age - 4836-3, 27 July 1827, between the Rio Tieté and the Rio Juquirf. 4837 - Pia at July sal eed carrie of Francisco da Paula da S Whee 4843. - 1853, “99 duly 1827, Se to me Nes do Felis. 48sh - 4868, 29 July 1827, Rancho do Felis 498 P-H.Y 7.0.L0.:6 LA Vol. 14, no. 8 1869 - yio8s 30 July 1827, Rancho do Felis to Jundiahy (S 23° 1) 46° 53! ‘habe - 4916, "31 Pica 1827, Jundiahy campest : 917 - 4923, 1 Aug. 1827, Jundiahy to Rio Geet teief (9 23703", 46-57"). hoek - 1933, 1 Aug. 1827, at Rio Capivary ho3h - 257s 2 Aug. 1827, Rio Capivary to town of Campinas or S. Carlos re 22° 53", W 47° 05"), the country entirely thick forest and hilly 4958 - ho7l, 3-6 Aug. 1827, Campinas, woods east of rancho. ho72 - hoo7, 8 Aug. 1827, same, dry stunted shrubby hill = 5019, ~9 ee 1827, Campinas to Fazenda peal Coun- try entirely forest; and soil a red-brown stron, 020 - 0, 10 Aug. 1827, Bucaitiva to Reet ings 5041 - 5067, ahd to Restinga, forest Restin 5073 - 5104, 11 cay "1827, Restinga to Mogi-Merim (S 22° 26', T 9105 - 5183, 15 Aug. 1827, Mégi-Merim, campestris, sylva 5184 - 5196, 19 Aug. 1827, Mégi-Merfm to brisdnga (Rio Origén- ga, S 22° 12°, wW 46° 57°). 5197 - 008, 20 ey 1827, brisénga. : 5209 - 52h2, 20 Aug. 1827, brisénga to Itupéva (Rio Itupéva, 9 - 5280, 21 Aug. 1827, Itupeva to Cocaes (S 21° 50%, w 47° : a ~ 5325, 22 Aug. 1827, Cocaes to Tambavi (S 21° 42", w 47° 2 : * 5326 - 5336, 23 Aug. 1827, Tambaii to Cereado (Cerrado? 8 21° 33", W 47? 2785, End of Bound Volume IT Bound — III continues with numbers 5337 to 6817 and takes Mr. Burchell as far as the city of Goyéz (Go oat 53hs, 2h Aug. 1827, Ollarfa theclee, S 21° 3h", wh7? ihe sae 25 Aug. 1827, Ollarfa to Rio Pardo (Ss 21° 28', WAT 5356 - 5382, 26 Aug. 1827, Rio brppes to Cubat&o mae Rio og") Pardo crosses the Rio Cubat%o (s a5", whe 5383 - 5408, 27 mre 1827, cece to Retiro. 5409 - 5419, Re 5420 - 5435, ba pie 1827, ico to Servo 5436 - S40, 29 Aug. 1827, Se S441 - 5iss, Be Aug. 1827, dig to Ribeir@o de Paciencia (S 20° 50°, W 47° 25"), 5is6. - 5461, ae de Paciencia. fee. - 5487, 30 Aug. 1827, Rio de Paciencia to Ribeirao de - 5497, 31 Aug. 1827, R. a = ~ 5524, 1 Sept. 1827, R. Maaco to Villa Franca (S 20 1967 L. B. & R. C. Smith, Itinerary of Burchell 499 32', W47 akt). 5525 - 5551, 5 Sept. 1827, Villa Franca to Cachoeira. 5552 - 5562, 6 Sept. 1827, Cachoeira 5563 - 5582, 6 Sept. 1827, Cachoeira “to Fazenda Buritf (Buri,- tys, $20 13', W47 4h). 5583 - 5617, 7 Sept. 1827, Burit{ to Retiro. 5618 - 5632, 8 Sept. 1827, Retiro to Rio Grande at Pérto Alto prints Alta, 8 19° 59', W 47° 46"). MINAS GERAIS 5633 - 5646, 9 Sept. 1827, Rio Grande to dil de Tenente. 5647 - 5665, 9 Sept. 1827, Retiro do Tenent 5666 - 5691, 10 ao 1827, Retiro do eats to Farinha Podre (Uberaba, S 19° Ast, 55"). 5692 - 5702, 11 a 1827, Farinha Podre. 5703 - 5719, 11 Sept. 1827, Farinha Podre to Lanho bee - 5752, - Sept. 1827, Lanhoso to Tejuco (Rio. a juiles . Agua Limpa, S 19° 35', Wca. 48°). ote - 5789, a Sept. 1827, Tejuco to Verdba-legit 5790 - 5836, 14 Sept. Jeet, Verdva-legitima to Rio si Velhas (S 19° 05", Wca. 47° 5837 - 5859, 15 Sept. 1827, Rio das Velhas to Furnas (Indi- Hess ae, S 19° 03°, 2 5 - 5880, 17 Sept. 1827, Furnas to Pisar%o (Cascalho Rico, Ss 18° 3h, W 470 se 5881 - 5907, a Sept. 1827, Pisar&o to S&o Domingos 5908 - 5913, 20 Sept. 1827, S&o Domingos to Rio furanad be and — do Matto (Borda da Mata on the Rio Paranafba at S$ 16°: 26%, Safes 5913 - 5931, Same in sylva densa antiquissima. ‘i GOIAS 5932 - 5934, 22 Sept. 1827, Borda do Matto to Ridcho (AGS: S 18° 10', w 47° 58" 93 35. - 5954, Ridcho to Catalfo (8 18° 10°, W:47° 5T").- 5955 - 5974, 24 Sept. 1827, Arraial de Catalfo to the Pé-do- é6rro ° 5975 - 6007, ni Sept. 1827, Pé do Morro to Rio Verissimo (s 27° SBF Whe? 2 008 - 6015, a Sept. 1827, Rio Verissimo to Brago do Rio Ve- rissimo (§ 17° 52', W 48° 06! "5, 6016 = 6019, 27 ae 1827, Rio Booey do Verissimo to Lages (Corrego da Lajes, S$ 17° 35°, W 48° O 036, 26 Sept. 187, Léges . Palmitél (Ribeiro Palmi- tél, s 17° ape 48? 60 037 - 604k, a hey es Palmitél to Pé da Roc 6045 - 6061, 30 Sept. 1827, Pé-da-Rocha to Rio iain ee od 26, W 48° 1 6062 - ae 2 Oct. 1827, Rio Corumba to Brejo. 6073 - 6077, 3 Oct. 1827, Brejo to Bat, the Engenho of Miguel 500 BEE BO +L-0.0 Is Vol. ly, no. 8 Ribeiro (‘The Enciclopédia dos Municipios lists a small stream named Bau near Pires do Rio, 8 17° 26° Ww 46%46'). 6078 - 6090, 4 Oct. 1827, Bail to cela Aberto (Alegre? On the 1940 American Geographical Society map there is a Campo Alegre Just northeast of Pires do Rio at § 17° 13, W 48° 17", which is not the larger Campo Alegre de Goids further east). 6091 - — 2 Oct. 1827, Campo Aberto or Campo Alegre? to S. Basil or S. Ba 6111 - ay ee Oct. 1827, Vasio (Basio?). 6122 - 6158, Vasio to Bomfim, partly woody campo, partly. 9 open. er ~ 6162, 7 Oct. 1827, Bomfim (Silvania, S 16° ho! » Ww 48? 36" BS aa ~ 6170, 8 Oct. 1827, Bomfim to Rio Piracanjuba (S 16° 33' W 48° sot 6171 - 61 196, 9 ee 1827, arena to As Antas (Angpolis, S 16° 20", W LB? 5 8"), P. Borge's. 61 dl - 6209, 1b Oct. 1827, As abbas to Forquilho 6210 = 6232, 11 Oct. 1827, ee ag to Forna (aubuas de Fur- nas? io sl 15° 56' WW 46* ea = - 6241, 12 eee 1827, ~ 6252, ar). Oct. 1827, mae to Méia Ponte (Pirendpolis, § ra ( Corrego de Cahyssara west and a little south of Pirendpolis). 6327 - 6334, 24 oct. 1827, Caisdéra to S8o Joaquim. 6335 - 6353, 25, 6s 27 Oct. 1827, Sdo Joaquim. 6354 - 6358, 28 Oc 1827, Sko Joaquim to Sapezél. 6359 = 6362, 29 aie 1827, Sapezél in the forest called Matto Grosso (S 15° 53", w 49° 18: 6363 ~ 6370, 29 Oct. 1821, Sepezal to Retiro (Corrego do Reti- s T? 371 - 6386, 30 Oct. 1827, Retiro to Conce eigko. 6387 - 6h; 3lfets mgd Pig long to Campo Alegre (Ribeir&o s 5 37, 1 Nov. "1827, Campo Alegre to Urii (Rio Uri, S 15° 38 - “60 Bie. 1827, Uri to Cidade de Goyéz (Goias, § 15° 56", W 50° 08! 6451 - 6541, "9 Jan. 1828, 6542 - 6556, 18 Jan. 1828, at high ground at S. Eastern entrance. 6557 - 6572, Goyéz, in woods along the Rio Manuel Nunes. 6573 - 6609, 28 Rives 1828, Goydz, i: along road to Arraial okt do Ferreiro (acs: 15* 52° Ww 50° 6610 - 6631, 5 tg 1828, Goyéz, left bank of pan Vermelho = the Rio de Manoel Gomes to the Carioca fountain (The Rio lho comes from the biethedine into Goyéz and flow out t the northwest 6632 - 6700, 11 Feb. ere road to Cuyaba as far as the Rio agem (Ri iro Bagagem 8 due south from Goydz). 1967 L. B. & R. C. Smith, Itinerary of Burchell 501 6700-2 - 6700-18, 17 Feb.. 1828, Gerke. along the first 3 same of the road to Aldeia de Ss. JOZ 6701 - 6747, 21 Feb. 1828, Goydz, tare e Forca WSW about 2 n-, Cantane de Nene; country uneven and sie wooded. T 56 - 6817, 23 Mar. 1828, Caminho de Nene, Sitio of Zacharia Ferreira to City This completes Volume III of Mr. Burchell" s Brazilian Cata- Para logu We continue the trip through Goids to and Belém with numbers and places listed in his Large Field Book. There are no dates liste in a "Memorand tanica” manuscri - chell states that "The Brazilian Herbarium (including those o Portuga d » and riffe) collected between 25 March 1825 (arrival at Lisbon) and the 1 of February 1830 (sailing from Pard) wa to be unpacked am on e ended on 15 Feb 1850." This indicates that these later numbers were collected between the last date listed in the Catalogue, 23 March 1828, and 10 February 1830. e hiibeats in the Field Book start with 6818 which immediate- 6818 - , (Go 15° 56° 208" aminho de Bacopary. 6927 - , (Goyadz), Caminho de Carreira. 6995 - , (Goy4z), Morro de Canta Gallo. 7063 - , (Goyéz), no entry T7070 - , (Goydz), peak near > Canto Gallo. 7159 - , (Goydz), suburbanae. 7164 - , (Goydz), Rio Bacalh&o 7213 - , (Goydz), Rio de Hanoel Gomes. 7289 - , (Goydz), suburbanae. 7310 - , (Goyéz), Chapel of S. Amaro. 7341 - , (Goydz), suburbanae. Resumption of northward travel from Goids toward Belém 1358 5 “Uri - Rio dras (Rio Uri goes west from Goids, s 16°. wo" ° and then northwest to Ribeiro das Pedras (AGS: S 15° 4s', W 49° 55° (ser. = s Road to Retiro (Corrego do Retiro, AGS: S 15° 55°, m 7389 - , Retiro. 7397 - , Retiro to Goiabeira. 7411 - , Goiabeira. 7436 -, Goiabeira to Corrego de Jeragua (Jaragué - Serra de Ji and a city of Jaragud nearby at S 15° enti W ao Thhg - , Corrego de Jeragua to Rio Doiateste. 7463 - , Rio Bonafacio to Estiva (there is a “Ribeirfo da Esti- va just east of Matto Grosso de Goids at AGS: S 15° 50', w 49° 20' but we could not find a stream or corrego ) 502 PHYTOLOGIA Vol. 1h, no. 8 26', but he would be retracing his steps). 7 -, Estiva to Capao de S. Joka (Morro de S. Jo&o? AGS: Ss 15° aa? w 49? T Siete e S, rag to Ribeirfio dos Macdcos (Rio dos Ma- edcos, S “ye 45), ‘ “488 7503 - , Ribeir&o dos ar to Olhos de Agoa (Olhos de fares ca. 16 - just above on AGS map and Flt below on Inst. B w 48° 45" on AGS map and 48° 36' on Inst. Bras. map. “s thie move he a abe his steps also). 75 1h - , Olhos de Bom Je 7542 - , Bom Jestis to Fazenda Seca pci af S eH Seca is due north of Olhos de Agua running from S 15° 10', W 48° 45* to S 15°, W 48° 50'). - , Fazenda Seca to Rio Maranh&o (Ss 14° 59°, w 48° 45*). ranhéo. r. 7580 - , Guardamér Fs Ribeiro d de S. Jo& 7601 - , Ribeiro de S. tote fg Ribeiréo. da Vendinha (Corrego Vendinha, just south o 613 - , Ribeirfo da Yendinha to Trahiras (Trairas, Tupiracaba s 14° 30 W eu pe + seo S. Joze (S. José de Tocantins, Niquelan- dia, S 1407 28", "4B? 2 7631 - , no Ae, Listed. 7647 - , S. Joze to R. Bagagem (S 14° 20°, w 4g? 14"), 7681 - , Rio Bagage to Rio da Serra (s 14° 21!, w 48° 10*). TT19 - , Rio da Serra to Rio Tucantins (Rio Tocantinzinho, S$ Ian 48°), TTT - , Rio Tucantfns to Ribeirfo Padre R The Pret of 48° W longitude northwest to join the Rio MaranhZo at about S 13° 37", W 48° O7*. We think Mr. Burchell met it at about S 14° - bre ts 7 ho to to As Lages (There is a Lajes on the Inst- Bras. map at S he 03', W 48° 08° which would mean oat he re- es his steps to the westward TS 21 - , As Lajes to Rio Moqué (me ey Rio Moquém is a Ri- beirfo back south at AGS: S 14° 30! 48° 08" near the town of i ce (Muquém) of the same latitude ae w 48° 10' longitude. This would mean that he further retraced his steps. i There is some pabilnnc 6 as to whether he did this, whether similar place names further northeast are lost, or whether he got his numbers 7860 -, Aréra Areias at AGS: S 14° 17", w 48° sec which is between sacekiei and Cavalcante and might be this ref 7876 - , As Ardras to es be re 13° 47", W 47? 30°). 7897 - , Cavaleénte. 7923 - , Cavalcdnte to Ribeirfo Gambd gu - , Ribeiro Gamb& to Rio dos Bois (S 13° 34", w 47° 16°) oO. TI5T - , Rio dos Bois to Rio Parandn (Rio Parand, S 13° 27’, 1967 L. B. & R. C. Smith, Itinerary of Burchell 503 W47 16"). 6 - , Rio Paranén to Taldya (Riaucho Atalaia, AGS: S 13° 30'-25', wh6? 53"). 991 - , Talaya to Rio Bezérra (S 13° 03', W 46° 57°). 7996 - , Rio Bezérra to Rio Chupet éiro. 8017 - , Rio Chupetéiro to Arrdyas (Arraias, S 12° 56", W 46° 8036 - , Arrdyas to Corrego Rico ge ct - , Corrego Rico to Sape ( Eapetetrot AGS: 8 12° 53°, W 93: Pe 8104 - , Sape to Sta. Brizida (Serra Sta. Brigida? S 12° 42%, 8121 - , Sta. Brizida to Rio Palma (S 12° 28', W 47° 08"). 8133 - , Rio Pélma to Conceig&o (Conceig&o a0" Norte, S 12° 14! W 47° 16"). 8146 - , Conceig&o. 8156 - , Conceig&o to Borac&o. 8169 - , Borac&o to S. sts © (acs: S 12° 05", w 47° o7'). 8177 - , S. Bento to Rio Cangdélho (Ribeir&o Cangalhe, S71 a oe 12°,” w 47° yi bs 199 - , Rio Gangeine to Rio Manoel Alves (Ss 11° 40-54", 47°-48° , he Bre oe crossed it at S 11° 53", W 47° 37'). 820 Rio. Manoel Alves to Rio Saldbre (8 oe khe, w h7? 8226 - , Rio Saldbre to Natividade (s cM ely W NT uty. 8251 - , Natividade to Chapada (S 11° 37', W 47° 52" 8267 - , Chapada to Rio das Pedras (AGS: 8 11° 25%, Ww 47? 45") Cénga. 280 - , Rio das Pedras to 8289 - | Cénga. 8311 - , hee a, do Boi (Morro Cabega do Boi or Mau- zoleo, AGS: S 10° ie, o hs 3- oabese do "pot ps Passatrés (Ribeiro Passa Tres (S e - Paseatede to Corrego ey , Corrego Raiz to Cdrmo spo do Carmo, S 10° 45', ? C4rmo to Corrego Fin 8393 - , Corrego Findo to Rote hed (Porto Nacional, S 10° BY 25*)i. - , Porto Real to Iga rapé (Igarape is a ee word for a channel between two islands or an ages d the mainland). in Porto Real, Ferry to mouth of Igara 3 8482 - , Porto Real, about village. | 8520 - , Porto Real, ford of Igarape- 8555 - , Porto Real, west side of river. 8600 - , Porto 1, Ba 8637 - , aioe pest te entrance to villeg gl-, ab the Manga, sg ” (Rio dos Mangues? 8 10 22°, 8 at 36" yee “ meets the Tocantins). 8753 - , Porto Real, various. 8765 - , Porto do hi eire, at Porto Real. 50h PHYTOLOGIA Vol. 14, no. 8 From this point on, Mr. Burchell evidently traveled on ec — Tocantins and did much of his collecting at portages around t falls 8788 - , Rio S. Jo&o (Ss 10° 25! w 48° = - it enters the To- cantins from the east opposite the Mangu 8801 - , mye aciosa. 8828 - o place lis “ oo hg Soo eloaas (Piltes, cachoeira or falls, S 9° 50, W 48 20! 8847 - , Mares (Cachoeira dos Marcos? § 9° 48", w 48° opt), : i 1 borhan (Cachoeira Lageado and townof Tageado, s 9° es 8 » Funfl scaebon iets s p* het, w 48° 20°, town of Funfl and Seda. lower falls, 8 9° hot, w 48° 0 Stl 8888 - , no place 1 teted. 8897 - , Red clay aA slate. 8912 - , no place listed. 8915 - , peas gs 1din Deg 2 oar (So Pedro de AlcAntara, AGS: S&o Pedro, S OF Ww 8962 - - , and ee - , no place listed. 8972 - , Sobradinh 9015 - - , Sta. Ma 9023 - , Tres Barras (Cachoeira de I. de Tres Barras, S 6° 13°, WNT oe see falls would be a likely place for him to ee! °o collect but they are way down river from Carolina in Maranh®o, so that he either went ahead and then back or got his numbers out of order). 6 - , Carolina (S 7 20', W47 28°) in MARANHKO. red tae - » no place listed 9080 - , Ss. iboats (Cachoetra, BS ow Were SB). 9082 - - , Sandy I PARK OP cin “key "s » Araguay (S&o Jofo do Araguay or Araguaia, S 5° 2k', W 912k - , no place 1 9126 - , Former Reafetro (Border between Goidés and Pard). 9132 - , Large Sand Is . 9135 - , Above Itahury (Tauiri? AGS: S 5° O7', W 49° 17" on the river). ~ » Anaraquarra (Cachoeira Aranaquara, AGS: S 4° 56', W 9165 - , Itaboca (Cachoeiras, S 4° 20-23!, w lig® 23"). aye? - , Guariba (Cachoeira just above Arroio, s 2 mi ee 18 - , Arroyos (AGS: Arroio, S 3° 57", W en ae 9238 - , no place li sted, 9241 - , no place listed. 9255 - ia de Espirito Santa (Ilha de A crs Santo, AGS: 5 2 Ob, Ww hg® 27), 1967 L. B. & Re C. Smith, Itinerary of Burchell 505 9259 - , Baifo (Ss 2° hot, w 49° hor), 92 933 gn feed Gaon da Sta. Anna in Rio Tocantins at S ee 3a. W i hig? 2 9358 - , Brits U (at the city of Moju, S 1° 53', W 48° 46"). 9378 - , no oe listed. 9379 -, Porto Real to Pard (Arrival in Belém do Pard? S$ 1° 26', W 48° 29! 9382 - , Pond vadgery near the house. Pard, Za 9152 - , Pard, a a st O47 - » Pard, my house ee pone anes must have been on outskirts oe Belém). a - , Pard 9546 - , Penh S. Joze to the arsenal (in an old map of the city of Belém, there was an arsenal on the northwest corner of the city on the be. ae i 9614 - , Para, Nazareth, Rosinha near to. ° ne) \o I . uuburban. 9908 - , Paré, Rivulet de Cerveja above the Arsenal. 9950 - , Pard, north of Campinha and Largo de Pélvora. 10072 - , Pard, woods SSE of S. Joze. PHYTOLOGIA Vol. 1h, NO. 8 | | | | a | Rio/de Janeiro | 16 August 1825 | 506 50 40 | \ i | Aiaieat re sp | ITINERARY OF BURCHELL | | (Goode base map no. | 103, copyright 1937 | by University of | Chicago) | \ | | ee | a i dA Rn sp Awan 2 — Pe EES re AL en SEE “ae caer C | \ _, Belém do Pard | ‘ iH" 4 210 February | 1830 % gers | Y Baigo } i / Pa : o——, Araguay / Carolina | ‘ Bo eiesien a ee AFunil Redan) Senate iim Be eestor staat 2 5 ANE eile St re Porto Real # ye 4& Natividade 4 Conceig&o Trahiras~@ * aveicente s a ee | ans ih he 7 Goyaz & WW Meia Ponte a Be ; i ¥ Cataldo a “XL Farinha P, aes . ce : vs rote | MPambaby 2s . S. Pa: oP ties ‘ 0 “a 5 ee a ee eae : : | he tos BS, ££ a a on aes . ADDITIONAL NOTES ON THE GENUS CITHAREXYLUM. V Harold N. Moldenke CITHAREXYLUM CHARTACEUM Moldenke Additional bibliography: Moldenke, Phytologia 13: 284 & 30k. CITHAREXYLUM FRUTICOS Additional Gin: coven Moldenke, Phytologia 14: 433—l35. (Sen & Naskar (1965) record this species as cultivated in India, t I have not as yet seen any specimens of it from that country. ag Trp KUNTHIANUM Moldenke mal bibliography: Sen & Naskar 7: 10. 1965; 3 ee Phytolog Sen & 9655 record this plant as cultiva ted in India, but I have as at i no actual specimens to substantiate this claim Bull. Bot. Surv. India ak CITHAREXYLUM LUCIDUM Schlecht. & Cham, Additional bibliography: Moldenke, —— 13: 296. searhgrs Rzedowski refers to this plant as a te - tall, found t growing on limestone hills with Quercus and sgt denies, at 1200 BOA oe alti tional citations: MEXICO: San Luis Potosi: J. Rzedowski Addi 10502 (Mi). CITHAREXYLUM LYCIOIDES D. Do: ude bibliography: Moldenke, Phytologia 13: 296, 311, & x CITHAREXYLUM MACROPHYLLUM Poir. Additional bibliography: Moldenke, Phytologia 13: 296—297, 302, & 31). 1966. n addition to the comments given previously, Berti also says liares s cia racimos espiciformes, axilares, verticilados. Flores zigo- gen roan rola 5-mera gamofila. Estambres pfs C4liz gamofilo." tional citations: VENEZUELA: Delta Amacuro: 166 (i— 2458308, W—21,58385) . ae CITHAREXYLUM MOCINNI D. Don Additional bibliography: Moldenke, Phytologia 13: 298. 1966; - Uy: 1. 1966. Recent collectors describe this plant as a tree, 10—12 m. tall, 507 508 PHYTOLOGIA Vol. 14, no. 8 the trunk 12—20 cm. in diameter, growing as "an occasional tree" in deciduous forests with Podo » Distylium, Ostrya, and Quercus or in "bosque meséfilo de enciino y pino con Abies y sniranthe dendron", at —— of 1500—-2700 meters, flo flowering January and Warch, f ruiting in March. The flowers on R. Me es 23221 and Rzedowski & MeVaugh 312 are described as White"; the racemes on the former collection are to 42 cm. in length! Smith & Tejeda describe the fruit as orange in color; their col- lection, cited below, constitutes a oo og voucher for plant : pages used in the United States of ee eee Departmen eee National Service Center, aeneeeais scree’ rogram Additional citations: MEXICO: Guerrero: Rzedowski & McVaugh 312 (Mi). Jalisco: R. McVaugh 23221 (Mi). Vera Cruz: Smith & & Tejeda 483 (W—2)71186), HONDURAS: Moraz&n: Williams & Molina R. 13703 (Ws). UM MONTANUM Molde: ional bibliography: Moldenke, Mem. N. Y. Bot. Gard. 9: 176. “19555 Moldenke, Phytologia 13: 310. 1966. CITHAREXYLUM MONTEVIDENSE (Spreng.) irene Additional bibliography: Chittenden, Roy. Hort. Soc. Dict. Gard. 1: 493. 1951; Molderice, Phytologia 13: 298-299. 1966 ITHAREXYLUM MYRIANTHUM Cham, Additional bibliography: Moldenke, Phytologia 13: 299—300. Woolston describes this plant as a tree, 6~-10 m. tall, the bark gray-browm, smoothish, the res ss growing at * the edge of woods, and called "sira-noro Additional” citations: PARAGUAY: ieciahoe: 610 (S). CITHAREXYLUM OLEINUM (Benth.) Moldenke Additional synonymy: Citharexylun oleinum Moldenke, in herb. Additional bibliography: Bocq., Rev. Verbenac. 197. 1863; Moldenke, Phytologia 13: 300--301. 1966. Rz wski describes the habitat of this plant as "Hab. ladera caliza con vegetacién de encinar", at 1850 meters altitude; Quin- tero found it g rowing on a slope with pine-oak woods. Additional stebbisnue MEXICO: Hidalgo: Quintero 2856 (Mi). San Luis Potosf: J. Rzedowski 6030 (Du—513552). CITHAREXYLUM PACHYPHYLLUM Moldenke Additional ee Moldenke, Phytologia 13: 301 (1966) 967. and 1h: 216, CITHAREXYLUM PACHYPHYLLUM var. idenke BibLi ography Moldenke, Phytologia ety 316. a Citations: PERU: Ayacucho: Iltis, Iitis, Ugent, & Ugent 18 © 1967 Moldenke, Notes on Citharexylum 509 (Z--type) CITHAREXYLUM PENTANDRUM Ve hs (1862) and 3: 223. 1863; A Rev. c. 88," 123, 130, & 2236 Dowiag 2 Liogier, Rhodora 67: 350. 19659 , Supl. Cat. Fl. ie gone 1966; Moldenke, Phytologia 1H — & 301 (1966) 34. xCITHAREXYLUM PERKINSI Moldenke Emended de jab Citharexylum perkinsii Moldenke, ee Suppl. 10: 5, syn. 196k; ooh ora potores ee 350. 1965. Additional babtaeranien Li , Rhodo 350. 1968; Mol- denke, Phytologia 13: 310 (A966) and 1): 31 oer The collectors of the three specimens cited below describe this plant as a tall shrub, | m. tall, with orange or bright- orange fruit [called "berries" in error by Webster & Proctor], ° ne hills, and top of Dolphin Head in jooeg at altitudes of 1000—2 2500 feet, fruiting in July and August. Material has been misidenti- fied and distributed in herbaria as C. caudatum 1. The abundant fruits on the racemes cast some doubt on the hybrid nature of these plants, but the foliar characters are certainly intermedi- ate between those of C. caudatum and C. spinosum L. Additional citations: JAMAICA: Webster & Proctor Sul, (Mi); Webster, Proctor, & Powell 5358 (Mi); Web: Webster & Wilson 5069 (Mi). es rane nie POEPPIGII cage nal bibliography: Adansonia 3: [Rev. Verbenac.] 223, “18635 idence re talocia "3! 95 (1954) and 13: 301—303. Additional citations: VENEZUELA: TAchira: Steyermark & Rabe 96629 (Z). rage CITHAREXYLUM POEPPIGII var. MARGARITACEUM Poepp. & Moldenke Additional bibliography: Moldenke, Phytologia 5: 95 (195k) and 13: 302—303. 1966. CITHAREXYLUM PUNCTATUM Greemn. punctatum Greemm. ex Moldenke, Enended synonymy: Aegiphila Résumé Suppl. vee 9, in syn. 1965; A. Marfa, Pl. Valle Cocha- bamb, 2: ‘ Additional se eas . eee Pl. Valle Cochabamb. 2: 1. 13: 30 (1966) and 1k: __ 1967. arfa (19665 cites his nos. 6 sien «85 & ‘oie trom Cochabamba, Bolivia. C4rdenas found the plan on rocky slopes at 32 ters alti Additional citations: BOLIVIA: Cochabamba: 6176 (W—272185). Department undetermined: M. pang 1517 sor 7 isotype). 510 PHYTOLOGIA Vol. 1h, no. 8 CITHAREXYLUM QUERCIFOLIUM Hayek Additional bibliography: Moldenke, Phytologia 13: 303 & 317. 966. ITHAREXYLUM SESSAEI D. Don Additional bibliography: Moldenke, Phytologia 13: 312--313. The Pri ringle 8969, distributed as C. sessaei, is actually the type collection of C. hidalgense Moldenke. CITHAREXYLUM SPINOSUM L Additional bibkinenaie: Bocq., Adansonia 3: [Rev. Verbenac.] 223. 1863; Lefroy, Bull. U. S. enh ag 253 an ser Achart, "apes Cent. Pl. Ind. 145. 1905; - Bailey Cult. Pl., ed. > pied on & 807 (192k), pr. rs “631 & 807. (1925), and pr. en's & 807. 1938; Sandw., Kew Bull. ae Inf. 1938: 373. 1938 H. Bailey, Man. Cult. Bees ed. 1, pr. 4, 631 & 807. 19lh; Chitten- den, Roy. Hort. Soc. Dict. Gard. ie “ip. 3 955 ——— Fieldi- ana Bot. 28: 1083. 1957; Moldenke, Am. t. 593 365. 1958; Anon., Kew Bull. Gen. Index 1929-1956, ye 33591 Gupta & Marlange, Trav. Sect. Scient. Inst. Frang. Pond. 3 (1): 78 Srinivasan & Agarwal, Bull. Bot. Surv. India 5: 80. 1963; E.E rR wer lw y 3 * ial is very - often misidentified and distributed in herbaria as C. fruticosum L, tan in hedges clay slopes aqeed to ere lord (196) adopts the name C, 4lld. for this plant, but says that the C. of Jacquin and the C. cinereum of Jacquin are C C. spinos um. He points out that the leaves in this species are longer than those of C. fruticosum and have a "drawn out end". It requires protection from frost when ustralia, Achart (1905) describes the plant as "Arbre de 20 2 60 pieds de haut, acclimaté dans 1'Inde. Le bois sert & faire des instruments 1967 Moldenke, Notes on Citharexylum 511 aye d'oi son nom de bois 4 guitare." Srinivasan & Agar- wat "(19 ) Sate it from Lahore and from Ferozepur, Punjab, and West ere eenetbar's as naturalized wild plants or cultivated is no Additional citations: Abciaoge pe Re J. Wagner 578 (S). LEE WARD ISLANDS: Dominica: Kimber 9 (Ws). “WINDWARD ISLANDS: Mar- tinique: Kimber 52), (Ws), 1895 rey 2035 (Ws). TOBAGO: Webster & Miller 9829 ( (Ss). CITHAREXYLUM SUBFLAVESCENS Blake : Pgs bibliography: Molcenke, Phytologia 13: 315--316. 966. Additional citations: VENEZUELA: Mérida: Bernardi 2069 (S). CITHAREXYLUM sg Fr lg Additional bibliography: Molde mke, Inform. Mold. Set 51, 2. - Liogier, Shodera 67: 350. 1965; a ets Phytologia 13: 317. 1966; Moldenke, rte Suppl. 1h: 8 ece’ llectors ribe sage bie. 9 feet tall, or a tree, 15 feet are “ak ‘black fru fruit in graeecty growing in Additional citations: CUBA: Las Villas: R. A. Howard 6629 (N). Oriente: Ekman ),87) (Mi). nb Yuncker 18324 (Mi). "An Evolutionary Survey of the Plant Kingdom" by R. F. Scagel, R. J. Bandoni, G. E. Rouse, W. B. Schofield, J. R. Stein, and T. Me. Ce Taylor, xi & 658 PPe, illus. Wadsworth Publi Company, Inc., Belmont, California, second printing, 1966. $11.50. A series of successful classroom experiences in group or mas- ter teaching of a on botany course for second-year students e year at the University of British Columbia, Canada, has been trans- formed from the ephemeral oral form to this excellent text writ- dali pating is holistic, yet modern and intellectually stimulating. The con- tent is rich, full, accurate, modern, and definitely not diffi- cult nor stuffy. an educational treasure to put before a student s Terminol is te consistent, drawing illustrations are ny = 4 , and the print is clear, well clean glossary helpful, and the index tho. 512 PB2.2 0:40.06 TA Vol. 14, no. 8 "several inches" into different orders! "Flora of Texas" by Cyrus Longworth Lundell and collaborators. Volume I, xi & 07 pp., illus. Texas Research Foundation, Renner, Texas. 966. With this presentation volumes 1 and 3 are now completed. This volume includes monographic treatments of the Pterido D. S. Correll, Potamogeton by E. C. Ogden and Scleria by E. L. Core. The work is very carefully done. The many any full plate drawings are very clear and well detailed, "A Handbook of Systematic Botany" by Subhash cate Datta, xv & 435 pp., illus. Eastend Printers in Calcutta 1, Asia Publish- ing House in New York City 10019 and Taplinger Publishing Co. distributors in New York City, N. Y. 10003. 1965. $9.50. Here is a needed, Smprebsenive, readable text on a topic not ivpiouie nc for the whole country of India. Following the cut but old-fashioned le. Parts II and III comprise the - of this work analyz the plant familie al orders". Obvious line f pertinent floral and other characteristics drawings o clarify the text. Part IV deals with such special topics as evo- lution, biochemistry, etc., but not with sufficient exposure to modern studies in serology, chromosome patterns and numerical tax- onomy. Part V consists of such aids to taxonomic studies as fam- ily keys 2 la Gray, identification of picpenr a.m genera and a glos- sary of native plant names from 17 of the most common languages. e is a useful appe ng scientific 9 valuable reference list of "musts" for the s nt which hopefully the college lib can p de. From the author should have a &@ presentation technique that is more interesting and challenging. The index is useful. Beca of poo mediocre photographic printing the central plates could better have been replaced by more S. rint is better than that texts; nevertheless, theory on p. xv, do they have to choose between "thalams long" and "thalamus shorter" no concrete . Index to Authors in Volume Fourteen Chung, I.-C., 321 215, 320, 390, 436, 511 Degener, 0. & I., 213 Moldenke, H. N. 0, Dwyer, J. D., 439 216, 216, 275, 3 OL, 3 25, 326, Dwyer, J. D., & Hayden, M. V., ae 39h, 398, 1,00; 420, 29, 07 Elias, T. S., 205 Reed, C. Fe, 393 Guevara, A. E., & King, R.M., Reed, C. F., & Robinson, H., 193 th, L. B., Jablonski, E., Wurdack, J. Jo, 257 Moldenke, A. Le, wee ut? 1%, Index to Supra-specific Scientific Names in Volume Fourteen Abies, 193-195, 197, 435 Amblystegiaceae, 203 Abromeitiella, 458, 462, 490 Ambrosia, 129 Acacia, 7, 150, 217, 279, 281, Amictonis, 255 ete 339, 3h9, 396, 408, Anacardiun, at Anachyris, 3 eee eee ioecclia, 201 Acanthaceae, Anacyclia, 6 Acanthospermum, 129 en eS Acanthostachys, 60, 6k Andrea, 1460, Acanthus, 30) Androlepis, "459, 463 Acer, 21 Andropogon, 393 Achyranthes, 391 ici oetmaned, 263, 26h Achyrocline, 129 Anomobryum, 200 Acrocliniun, 129 Anonymus, 112 Acrostichum, 312, 316 Anopheles, 306, 332 Adelobotrys, 265 Lop » 402 Adenodesma, 269 Anthemis, 129 Aechmea, 460, 461, 46k Anthoc 198 Aegiceras, 304, 330 erotaceae, 198 A Ta, 151, 245, 427, 509 Antidesmia, 431 Agave, 217, 279, 261, 282, 391, es 413 396, 408 wn agama’ eratum, 129 jpn 459, 463 Alcantarea, 159 Arbutus, 3 Allenrolfea, 315 Aregelia, 163 o8, Aristida, 319 Aloysia, 353 Artemisia, 129, 319 b na, 272, 273 oe 301, 31 3 514 Asclepiadaceae, 391 Asclepias, 28 Avicennia, = » 305-31 325, 315—3 31 ri sab, 437 Bennettiales, 128 Bennettitales, 392 SS 203 ae age 75, 286, 292, 437 Braunia, Breutelia en Brocchinia, 457, 458, 462 Bromelia, 1,60, +65 Bromeliaceas, 57, 459, k61, 463 165, L67, 69, WT, 473, 75, P8323 D:L.0-6-3:4 Vol. 15, no. 1 nt 479, 481, 483, 485, 487, 89 Bronelioideae, 457, 459, 463 Calceolaria, 285, 288, 292 Calea, 1 Calendula todon, 199 Ceratolejeunea, 197 Ceresia, 3 1967 Index 515 Cereus, 352 Cyrtocarpa, 412 Ceriops, 30 Dahlia, 131 Chaptalia, 130 Dasyphyllum, 321, 322 Chenopodium, 282 Datura, 20 Chevalieria, 6) Davya, 265 Chiranthodendron, 508 Deinacanthon, 16) Chirripoa, 1,63 Derris, 178 Chrysanthellun, 321 Desmoscelis Chrysanthemum, 130 Seatarceohtia, at 462, 490 Chuquiraga, 130 Dianella, 213 Cincho: Dicho , 81 tharexyhum, 1,29 Citharexy. lun, 216, 429-35 4 507- 511 Citharoxylon, 32 Cladium, 330 Combretum, )11 Conanthes Cornutioides, 420 Cosmos, 131 Cotinus, Cottendorfia, 458, 62 Dicranaceae, Dic ranantherae, 263 Dicramm, 199 Didieriaceae, 391 Didymodon, 199 Diostea, 02 Diotanthera, 258, 259 Dipelta, Diplostephium, 131 Disteganthus, 6) Distichlis, 23) Dis » 308 Ditrichaceae, 199 Drepania, 395 Drosera, eee Dumortiera, Dyckia, 157, “Ts, 462, 465-188, 516 PHYTOLOGIA Vol. 15, no. 1 Eriostax, 6) Gravisia, 61, 463, 46) » Greigia, "460, 463, 464 Espeletia, 131 Grevill 213 Eucalyptus, 330 Grimmia Eudavya, Grimmiaceae, 200 rium, 131, 132 Guzmania, Euphorbia, 285, 292, 391 Gymnospermae, 512 Euphorbiaceae, 391, 41, 50 G 132 um, 203 Hechtia, 58, 62, 490 Euschlegelia, 33 Hedwigia, 202 Eustoma, 28 Hedwigiaceae, 202 Excoecaria, 30) Heleochloa, 393 Exema, 3 Helichrysum, 132 Fascicularia, 460, 6h Helipterum, 132 Fasciculata, | 6 Helleriella, 4, 5, 21, 2h Fernseea, 59, 1163 Hernandea, 2 Fe . He Herpetineurum Festuca, 393 Hesperogreigia, “yal Fissidens, 198 Beets ot Fis sidentaceae, 198 idlaria, , 3k9 Forchammeria, 1113 aseceress. 461, 464 Fosterella, 18, ms foaal sake Fouquieria, }1 oulle pane 391 enoniylacoe, 213 Franseria, h13 Hymenophyllum ia, “20 eat aS 25 Punariaceas 200 Hypericum, 289 a Salinasaay tar: 279 Hypnaceae, 20) ers 157° Hypochoeris, 132 Garrelia, rae “i » 489 Hypopterygiaceae, 203 ‘ Garrya, 19h, Hypopterygium, 203 Gaura, 289 8, 205, 292 azania, 13 Icacinaceae, 391 Geraniaceae, 391 Ichnanthus, 83-85 Geunsia, 36, 38, Wi, 45, 117, Tchthyothere, 132 1L5, 225, 239, 20 a Ginkgoales, 392 Imperata, 87 29h, shh Inga, 205-212 Gloneropiteaimia, 59, 163 Lonopeia, 3 Gnaphalium, 132 Ipomoea, 183, 391 Gne Ss, 392 Jatropha, 13 Gongora, 3 Juncus, 330 Goodenia, 332 ungermannia, 196 Gouldia, 213, 21) Jungia, 132 Graffenrieda, 266, 267 Tabiatas’ of h27, 512 1967 ularia, 306-308, 31 Lamiaceae, AAS we, 3 Lan 402, 8 107, : eee P Index 312-316, 3, ho2 517 Marantaceae, 65 Marchantia, 19 Marchantiaceae, 198 217, 277, 325, 399%,Masdevallia, , 16 7 Matricaria, 133 Maytemus, 327 Melastoma, 265 Melastomataceae, 257, 259, 261, OT, 208,720 218 195 Mesembryanthemaceae, 391 nearia, 36h 9 Lipachaeta, blk Wittenothamnium, 202 Crp riggs | Mniaceae, 201 Lippia, 217, snr 344, 100-419 ’ Mnium, 201, 202 bi a bes ? ’ qos ambar, 195, 39h, 421, 3h, aertie. seh Livi 389 Monochaetum, 261--263 Lobelia, 332 Montanoa, 133 Loganiaceae, see 122, heh Moraceae, 391 Lophocolea, orinia, Lorea, 50 Musci, 198 eT Nutisia, 133, 285, Tumnitzera, 30h Navia, L57, Ls8, 62, “es, 1,89, Lupinus, 285, 292 ~ 190 Neckera, eae - Neckeraceae, 202 . Neea, 137-139 Machaerocereus, bh ois Sian. Why 6h, 196 3 8 Mamanira, 107, 115, 142 Mangle, Mangle, 317 Widularia, 63 meee, ©. Ye, 463 518 PHYTOLOGIA Vol. 15, no. 1 Noteroclada, 197 Phrymaceae, 338 Notothylas, "198 Phygelius, 146 ctaginaceae, 137 Phyla, ) Obione, 30! oe a 225 Ochagavia, 60, 63 Physalis, Ochnaceae, 39 rtataccacoae, 02 Ocimum, 403, 11 Pimela, 3 Octoblepharum, 199 Pinus, 1s, 194, 280, 289, 299, Octomeris, 271 is te Tad 417, 421, 435 Oenothera, 285, 289, 292 Pipturus, Oepata, 317 Piqueria, 2, 325 Oleaceae, 167 Pisonia, 3 * on ae a Oliganthes, 133 Pitcairnia, 457, ’ ’ mR eya, L13 Pitcaimnioidess, 157, bl Olyra, 85, 86 Placseptalia, 163 Oncidium, 7 Plaglotheciacene, 204 Guosaria, 133 Plagiotheciun, 20) Opisthocomus, 315 Plantago, 282 285, 292 Opuntia, 195, om, 281, 31, Platyglottis, ls 9, 39 Pleurothallis, 7-1), 21, 22, 26- Orchidaceae, 1 . 2 Oreoweisia, 193, 199 Plicatula, 373 Orobanchaceae, 392 couches, es Orthophy tum h6h, oc : Orthotrichaceae, 201 Pogonatum, 20) Orthotrichum, 200, 202, 20) Pohlia, 201 Ostrya, Polemoniaceae, 512 Ouratea, 39, 0 Pollalesta, 133 Oxalis, 279, 3h9 Polygomum, 279 Pachycereus, 1,13 Po a, 133, 134 andanaceae, 165 Poly: Pandamus, 25) Po. chaceae, 20) Panicun Po chum, 20) Panurgidae, a 28h, 298, Ponerinae, Papillaria, Nk cot gsc serehis; 298 Parvifolia 6s orophyllum, 1 Paspalum, 358-389 Porphyra, 156, 162, 255 Passifloraceae, 391 Portea, Pectis, 133 Portulaca, 213 Pedaliaceae, 391 Portulacaceae, 391 Penicillata, 79 Potamo: Persea, 20 Pottiaceae, 199 Petitia, 151 Pourretia, Phalacraea, on 325 Prantleia, 16) Philonotis Premna, h2, 156, 248, 420 , Prionophyllum, 462, 466, 478 1967 Priva, 27». 336-361, 353, 355, 357, 39s rd Pseudopterole Pap Se errelaea, ee Pterolepis, 261 Ptychomitriaceae, 201 Ptychomitrium, 201 Puya, » él, L65, 490 77 Quadrifaria, 363 Quercus, 191-197, 217, 280, 262, 289, i hora, 30h, 307-309, 312, , 36, 327, 332 Rhus » 407 Index 519 Rumex, 279, 285, 292 S acciolepis, 85 Salicornia, 305, 315, 327, 333, Seblegelies 433 chlotheimia, 202 Sematophyllum, 2 Sincoraea, 46), Sobralia, 3, lb-19, 22, 31-3h Sodiroa, 3 Solamm, 279-282, 285, 289, 292, 3 ’ Steiractinia, 135 é ’ 292 Stereodon, 202, 20h 520 PHETTOLOGIA Vol. 15, mo. 1 Stevia, 135 Stillingia, 451, 453 Trichomanes, 213 Stolonifera, 78 Trichostomum, 200 Streptocalyx, 159, 463 Tridax, 13 Stylurus, 213 Trixis, 135 Styrax, 195 Uapaca, 08 Sueda, 305, 315 Uredo, 191 Sy. Mae Urt. 7 Symblepharis, 199, 20) Urticaceae, 127 Symblepharos, 20, Each saa Vanilla, 1 tabebuia, 10 % Snes cues pogete®, 1» nek Bh le yc ne ie stand 216, 256, 392, 433, Bdiacinsnati hamerear ak 12 Verbenajus, 277, 28h, 298, 300 Ve 275 Taraxacum ae Verb 2, 13 Targionia, 19 Vernonia, 135, 28h Targioniaceae, 198 Veronica, 285, 292 Teijsmanniodendron, }00 Ternstroemia, 195 Vitaceae, 391 2S Tetraglochin, 289 Vriesea, 459, 62 Tertula, 336 Wae Tessaria, say Wedelia, 135 Tetragonae, 20 Welwitschia, 392 Thecophyllum, 162, 463 Wilcoxia, 413 Petcare | ee Willrussellia, 162 , a ’ ack sed Wittmackia, 16), . china, Wittrockia, 461, 46h Tillandsia, 459, 62, 463, 485, ipsa 736 “Tis Tillandsioideae, 157, 458, 62 Xeracina, 265, 266 Timmiaceae, 201 Yucea. 279. 28 Timmiella, 201 Fe ae aD core Tortula, ag 202, 348 195 Tingiberac eae, 65 Zinnia, 136 Publication Dates of Volume Fourteen No. 1 —- November 1), 1966 No. 5 —- February 3, 1967 No. 2 -—- November 23, 1966 No. 6 — March 21, 1967 No. 3 — December 1h, 1966 No. 7 — April 28, 1967 No. —- Jamary 10, "1967 No. 8 —May 18, 1967 7e &) ee