ss GONTRIBUTIONS FROM THE UNIVERSITY OF MICHIGAN HERBARIUM, NO. 8 FLORA OF EASTERN TABASCO AND ADJACENT MEXICAN AREAS CYRUS LONGWORTH LUNDELL STUDIES OF AMERICAN SPERMATOPHYTES—III BY CYRUS LONGWORTH LUNDELL ANN ARBOR UNIVERSITY OF MICHIGAN PRESS SEpremBeErR, 1942 CONTRIBUTIONS FROM THE UNIVERSITY OF MICHIGAN HERBARIUM, NO. 8 FLORA OF EASTERN TABASCO AND ADJACENT MEXICAN AREAS BY CYRUS LONGWORTH LUNDELL STUDIES OF AMERICAN SPERMATOPHYTES—II1 BY CYRUS LONGWORTH LUNDELL ANN ARBOR UNIVERSITY OF MICHIGAN PRESS SepremBer, 1942 CONTENTS Fora orf EASTERN TABASCO AND ADJACENT MEXICAN AREAS...... SrupIES OF AMERICAN SPERMATOPHYTES— INDEX PLATES (Plates I to IV face page 88) PLATE I. The Usumacinta River with a view of Boca del Cerro. Il. The Usumacinta River with an inner view of Boca del Cerro. III. The expedition crossing a savanna east of Palenque. IV. Javalinero on the Chacamax River below Palenque. TEXT FIGURES FIGURE PAGE 1. Sebastiania confusa Lundell. Matuda 3172, half natural size.. 39 9. Erblichia Standleyi Steyermark. Matuda 3319, half natural size 46 3. Hugenta Papen Lundell. Matuda 3361 (type collection), Age oR MND CTH EIU As 0h eo ete ar eS ae Goce amo a6 48 4, Hugenia Aner Lundell. Matuda 3561 (type collection), Ahead HA bUeENL GIVA Sng 2 eee ee RM RE a 6 cums iia pene 0 5, Diospyros salar aad Lundell. Matuda 3843 (type collection), al Pana Pune esi AOr son tte rea de ha a. 2 ey ee eee me ee 56 6. Stachytarpheta ae Moldenke. Matuda 3218 (type col- lection), half natural size... 6.6... ee eee ee 62 ths SS Unie ae um Standl. Matuda 3670 (type collec- tion) halfanaturalesisee i. cog ead ot os one 66 8. Chione opens Standl. Matuda 3675 (type collection), half TANABE TWAclo ke 5 aa ee ee PER Etat 2.0 oly aod 3.6 68 MAP MAP 1. Map of the lowlands of southeastern Mexico (facing)....... 6 FLORA OF EASTERN TABASCO AND. ADJACENT MEXICAN AREAS* HROUGH 1938 six expeditions for botanical exploration in the Mayan area were sent out under the joint auspices of the University of Michigan and the Carnegie Institution of Washington. These undertook surveys in Yucatan, Campeche, Quintana Roo, British Honduras, the Department of Petén, Guatemala, and the Huastecdn country of San Luis Potosi. None of the expeditions visited the extreme western part of the Yucatan Peninsula and adjacent lowlands of southeastern Mexico. Few areas of equal size or importance in Mexico had been so com- pletely neglected by botanists. Prior to 1939, not a single plant collection worthy of note had ever been made in eastern Chiapas, eastern Tabasco, or extreme western Campeche. The writer in 1932 explored the San Pedro de Martir River in Petén almost to the Mexican border,! and this preliminary reconnaissance trip established that the forest of that area was similar in aspect to the forest of central Campeche and central Petén. To obtain addi- tional data regarding the vegetation, a representative plant col- lection from various sectors of the region was desirable. Mr. Eizi Matuda, who has collaborated with the University of Michigan Herbarium in his study of the Mexican flora, was in- duced by the writer in 1939 to botanize along the Usumacinta River and tributaries. The trip was financed in part by a grant from the Carnegie Institution of Washington and by the sale of duplicate sets of specimens. From May 9 through July 28, 1939, Matuda collected in the region. His field work began at Balancan, extended up the San Pedro de Martir River to the Petén border, across to Tenosique through San Isidro, along the Usumacinta River from Estapilla south again to the Petén border near Piedras Negras, across to the Chacamax River in Chiapas, up to Palenque, * This paper has been published with funds derived from the income of the endowment of the Horace H. Rackham School of Graduate Studies of the University of Michigan. 1C. L. Lundell, The Vegetation of Petén, Carnegie Inst. Wash. Publ., No. 478, 1937. 6 CYRUS LONGWORTH LUNDELL northeast to Aguacate and EK, Zapata, and down the Usumacinta River to Palizada, Campeche. From Palizada he went to Vapor where the final collections of the trip were made, The principal localities are designated on Map 1. Matuda obtained 911 numbers of plants, his series 3008 through 3914, a total of 5230 specimens representing approximately 700 species. The first set of the collection is in the University of Michigan Herbarium. His collection localities, some of which were only temporary camps (champas), are listed below approximately in the order in which he visited them. Matuda often abbreviated localities on the labels, and his abbreviations (or misspellings) are enclosed by parentheses. Further, in giving the locality in which a particular collection was made, he indicated first the last large town through which he had passed, or the principal town in the district. Hence for his collections at La Palma, which is quite removed from Balancan (Map 1), his labels read “Tabasco, Balanedén, La Palma.” Although at times confusing, his practice has much to commend it where obscure localities are involved. TABASCO I. Balancaén—on the Usumacinta River below the mouth of the San Pedro de MArtir River, 2. Mercedes—west of Balancdén on the road to Las Cruces. 3. Achotal—about eight miles east of Balancan. 4. Ojo de Agua (Ojo’ Agua)—about five miles southeast of Balancan. 5. Reforma—a finca east of Ralancdn and north of Limon on the San Pedro de M4rtir River. Most of the collections were made at a camp set up by Matuda and his assistants in the high forest about ten miles north of Reforma. 6. La Lucha—about six miles south of Balancdn. 7. Provincia—near Reforma on the San Pedro de Martir River. Near Reforma the river is practically impassable, and boats operating into Petén make headquarters at Provincia. 8. La Palma—on the San Pedro de Martir River east of Teno- sique and near the Petén border. a ty ; : 95" | MAP OF | LOWLANDS OF SOUTHEASTERN MEXICO SCALES Kilometers yp a 10 030 0 50 * Miles Comalcalco A Va Atasta \ Huimanguillo nite \ titted ' “n ‘“ / MAN ey Cot Wvato Obregon (Frontera Piedras Negras tea, ' oo. ah ‘ es) ‘ a sy) to § <2 ~ J illahermosa (6-Juan Bautista) ! Macuspana ’ SMacusoand San Diego N ay Nan xs i P V B aP s 2 3 Simojovel — © e cee « oe fee Pe, Bee ly % exer” te i 2 ‘ ESA Sar Ococingo =) / Gr... 2 ty ‘ Z loge ° om! M Ke. ae 94° 93° 92°. Eresls Map 1 FLORA OF EASTERN TABASCO 7 9. San Isidro (S. Isidro)—an overnight muleteer camp (champa) in high forest about midway between La Palma and Tenosique. 10. Juarez—about five miles from Tenosique. 11. Tenosique—on the Usumacinta River south of Balancan. 12. Retiro—south of Tenosique near the Petén boundary. Col- lections from this locality were made southward into Petén and northward to a point on the Usumacinta River called Santo Tomas (a champa). 13. Estapilla—on the Usumacinta River north of Tenosique. 14. Boca del Cerro (Boca Cerro)—a gorge on the Usumacinta River southwest of Tenosique (Pl. I). It is near the place desig- nated on some maps as Santa Margarita. CHIAPAS 15. Javalinero (also spelled Jabalinero)—a ranch on the bank of the Chacamax River about twelve miles below Palenque (PI. IV). 16. Palenque (Rvina)—ruins of the ancient Mayan city. 17. Sierra de Palenque (Cerro de Palenque)—the ridge south and southwest of Palenque. 18. Aguacate—a ranch in the savanna country midway be- tween Palenque and E. Zapata (formerly Montecristo). CAMPECHE 19. Palizada—a town about midway down the Palizada River. 20. Vapor—a ranch on the Marente River east of Palizada. From the collections, Matuda’s diary, which gives a brief ac- count of the country traversed, and his photographs, it is evident that the vegetation of the region differs little from that of Petén. In the vicinity of old towns and settlements, savannas alternate with second growth, and these give way to old advanced forest in outlying districts. The savanna country of the Palenque sector (Pl. III) is almost identical with that of the La Libertad country of Petén; differences in the flora are negligible. Old advanced forest is to be found in remote localities such as San Isidro, Retiro, Boca del Cerro (Pls. land II), and Javalinero (Pl. IV). Most of the 2 Thid., Pl. 35. 8 CYRUS LONGWORTH LUNDELL species have been collected elsewhere in the Yucatan Peninsula, and the others doubtless will be found in adjacent areas of Cam- peche and Petén. Although his collection probably represents less than a third of the total fern and phanerogamic flora of the areas traversed, we can anticipate that the whole flora is essentially the same as that of the southern two-thirds of the Yucatan Peninsula. Explorations of Mr. Llewelyn Williams in the isthmus of Tehuantepec, Mexico* and the writer’s collections from Tamazun- chale, San Luis Potosf,‘ indicate that many elements of this wet lowland flora range northward along the coast to the Panuco River. Since the collection is the first of importance from eastern Tabasco and adjacent Mexican areas, the following enumeration is the first major contribution to our knowledge of the flora of the region. Approximately 700 species are listed in the following cata- logue. Although no novelties are described in this report, thirty- eight published elsewhere have been based primarily on the collection. All notes on habit, habitats, vernacular names, ete., were supplied by Matuda on the labels of the first set of specimens now in the University of Michigan Herbarium. Six of the species described from the collection and two others of particular interest are illustrated. POLY PODIACEAE (Determined by W. R. Maxon, Washington) ACROSTICHUM DANEIFOLIUM L. & F,—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3547. ADIANTOPSIS RADIATA (L.) Fée.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3330. ADIANTUM DEcOoRATUM Maxon & Weatherby.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3328. ADIANTUM LATIFOLIUM Lam.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3678. * Llewelyn Williams, ‘‘Arboles y arbustos del istmo de Tehuantepec, Mexico,” Lilloa, 4: 137-71. 1939. 4 C. L. Lundell, “Studies of Mexican and Central American Plants— IV,” Field and Lab., 6: 9-16. 1937, FLORA OF EASTERN TABASCO 9 ADIANTUM OBLIQUUM Willd.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3325. ADIANTUM PETIOLATUM Desv.—Tabasco: San Isidro, June 7— 11, Matuda 3336. ApIANTUM VILLOsuM L.—Tabasco: Reforma, in advanced forest, May 22-26, Matuda 3205. Aprtantum Wiisont Hook.—Chiapas: Javalinero, in adv anced forest, July 6-9, Matuda 3616. ASPLENIUM CRISTATUM Lam. forest, June 1-6, Matuda 3307, AspLENIUM FoRMosUM Willd.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3331. ASPLENIUM sERRATUM L.—Tabasco: Retiro, on limestone in ad- vanced forest, June 19-25, Matuda 3463. BLECHNUM FRAXINEUM Willd. (?).—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3612. BLECHNUM SERRULATUM Rich.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3088. BOLBITIS CLADORHIZANS orci) Ching.—Tabasco: San Isi- dro, in advanced forest, June 7-11, Matuda 3379. CyYCLOPELTIS SEMICORDATA (Sw.) J. Sm.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3436. - DicryoxXIPHIUM PANAMENSE Hook.—Chiapas: Javalinero, in ~ advanced forest, July 6-9, Matuda 3606. Dryopreris pentata (Forsk.) C. Chr.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3363, 3366A. DryYopreris GONGYLODES (Schk.) Kuntze.—Tabasco: San Isi- dro, in swamp, June 7-11, Matuda 3338. Dryopreris LirsMannit M. & M.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3603. DRYOPTERIS SERRATA (Cav.) C. Chr.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3301. GyMNOPTERIS RUFA (L.) Bernh.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3416. NEPHROLEPIS BISERRATA (Sw.) Schott.—Tabasco: Mercedes, on palm tree at riverside, May 9-14, Matuda 3021. Tabasco: La Palma, in advanced 10 CYRUS LONGWORTH LUNDELL NEPHROLEPIS PENDULA (Raddi) J. Sm.—Tabasco: La Palma, on tree at edge of forest, June 1-6, Matuda 3332. POLYPODIUM ANGUSTIFOLIUM Sw.—Chiapas: ruins of Palenque, on tree, July 10-12, Matuda 3705. Po.typopium PatmMert Maxon.—Tabaseco: La Palma, on tree, June 1-6, Matuda 3804. Campeche: Vapor, July 25-28, Matuda 3884. PoLyPopiuMm PEcTINATUM L. (?).—Tabasco: San Isidro, June 7-11, Matuda 3372. PoLyPopiuM PHYLuITIDIs L. (?).—Tabasco: San Isidro, on tree at lakeside, June 7-11, Matuda 3368. PoLypoprum PoLypopioipEs (L.) Watt—Tabasco: Boca del Cerro, on tree, July 1-5, Matuda 3572. PrEeRIS GRANDIFOLIA L.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3442; Boca del Cerro, at edge of swamp, July 1-5, Matuda 3584, 8554. Preris LONGIFOLIA L.—Tabasco: Boca del Cerro, on limestone, July 1-5, Matuda 3535 STENOCHLAENA RECURVATA (Fée) Liebm.—Chiapas: Javalinero, on tree in advanced forest, July 6-9, Matuda 3652. TECTARIA HERACLEIFOLIA (Willd.) Underw.—Tabasco: Retiro, June 19-25, Matuda 3416. VITTARIA LINEATA (L.) J. E. Sm.—Tabasco: Reforma, May 22-— 26, Matuda 3212. SCHIZAKACKAE (Determined by W. R. Maxon, Washington) LOPHIDIUM ELEGANS (Vahl) Presl.— Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3662. LYGODIUM HETERODOXUM IXunze.—Chiapas: Sierra de Palen- que, in advanced forest, July 14, Matuda 3720, a vine. LyYGODIUM MEXICANUM Pres].—Chiapas: Sierra de Palenque, at edge of forest, July 14, Matuda 3716. SALVINIACEAR (Determined by W. R. M axon, Washington) SALVINIA AURICULATA Aubl.—Tabasco: La Palma, in lake, June 1-6, Matuda 3274. FLORA OF EASTERN TABASCO VA LYCOPODIACEAE (Determined by W. R. Maxon, Washington) Lycopopium cERNUUM L.—Chiapas: Aguacate, in savanna, July 16-18, Matuda 3781. SELAGINELLACEAE SmLAGINELLA sp.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3676. SELAGINELLA sp.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3691. SELAGINELLA sp.—Chiapas: ruins of Palenque, July 10-12, Matuda 3702. CYCADACEAE TAMIA FURFURACEA L. f.—Tabasco: Ojo de Agua, southeast of Balanedn, in savanna, May 9-14, Matuda 3117. POTAMOGETONACEAE PoraMOGETON sp.—Campeche: Vapor, in lake, July 25-28, Matuda 3876. ALISMACEAE SAGITTARIA LANCIFOLIA L.—Tabasco: La Palma, riverside, June 1-6, Matuda 8251. Campeche: Palizada, in swamp, July 25- 28, Matuda 3840; Vapor, in swamp, July 25-28, Matuda 3886, flowers white. The specific determination is somewhat uncertain, and more than one species may be represented. GRAMINEAE (Determined by Agnes Chase, Washington ; LeRoy H. Harvey, Ann Arbor) ANDROPOGON BICORNIS L.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3726. Anprorocon BourGcart Hack.—Chiapas: Aguacate, July 16—- 18, Matuda 8795. ANDROPOGON CoNDENSATUS H. B. K.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3741. ANpRopoGoN SEeLLoanus (Hack.) Hack.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3723. 12 CYRUS LONGWORTH LUNDELL ARTHROSTYLIDIUM SPINOSUM Swallen, Journ. Wash. Acad. Sez. 28: 6. 1938.—Tabasco: Retiro, in advanced forest, June 19-25. Matuda 3454, 7-9 m. high, 2-3 em. in diameter, clambering. Axonopus Purpusir (Mez) Chase.—Tabasco: Achotal, east of Balancan, May 9-14 Matuda 3089. Chiapas: Palenque, in savanna July 9-14, Matuda 3737. BoureLovua Hirsuta Lag.—Tabasco: Estapilla, in savanna, June 27, Matuda 3506. CENCHRUs Brownt R. & S.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3248. EcHInocuLoa Waurenrt (Pursh) Heller.—Campeche: Palizada, in swamp, July 25-28, Matuda 3839, 8842, Eracrostis Hiniorritr S. Wats.— Chiapas: Aguacate, July 16- 18, Matuda 3799. ERAGROSTIS HYPNOIDES (Lam.) B. 8. P.—Tabasco: Reforma, May 22-26, Matuda 3158. Campeche: Palizada, swampside, July 25-28, Matuda 3864, vernacular name, pan caliente. ERIocHLOA puNcrata (L.) Desv.—Tabasco: Retiro, riverside, June 19-25, Matuda 3481. Chiapas: Palenque, in savanna, July 9-14, Matuda 3759. IRIOCHRYSIS CAYENNENSIS Beauv.—Chiapas: Palenque, in savanna, July 9-14, Matuda 38750. UADUA ACULEATA Rupr.—Tabasco: Reforma, May 22-26, Matuda 3198, trunk 8 em. in diameter. HETEROPOGON CONTORTUS (L.) Beauv.—Tabasco: Istapilla, June 27, Matuda 3509. OMOLEPIS ATURENSIS (H. B. K.) Chase.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3736. HYMENACHNE AMPLEXICAULIS (Rudge) Nees.—Campeche: Vapor, swampside, July 25-28, Matuda 3892. Lastacis pivaricata (L.) Hitche.—Tabasco: La Palma, in second growth, June 1-6, Matuda 8242. LASIACIS PROCERRIMA (Hack.) Hitche.—Chiapas: Sierra de Palenque, southern slope, July 14, Matuda 8718. Lasracis RUSCIFOLIA (H. B. K.) Nees.—Tabasco: La Palma, June 1-6, Matuda 3292. FLORA OF EASTERN TABASCO 13 LeprocoryPHiuM LANATUM (H. B. K.) Nees.—Tabasco: Acho- tal, east of Balancin, May 9-14, Matuda 3106. Chiapas: Palenque, in savanna, July 9-14, Matuda 3760. MANISURIS GUIANENSIS Hitche.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3144A. According to Mrs. Chase, this species has been known previously only from British Guiana and Brazil. ManisuriIs RAMOSA (Fourn.) Hitche.—Chiapas: Aguacate, July 16-18, Matuda 3774. Mesosetum BuaKei Swallen.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3738, 3748 (det. Swallen). OLYRA YUCATANA Chase.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3244. OryzA LATIFOLIA Desv.—Campeche: Vapor, in swamp, July 25-28, Matuda 3901. Panicum CYANESCENS Nees.—Chiapas: Aguacate, July 16-18, Matuda 8769, 3792. PANicuM HirsUTUM Swartz.—Tabasco: Boca del Cerro, in second growth, July 1-5, Matuda 3577. Chiapas: Javalinero, in second growth, July 6-9, Matuda 3632. PANICUM HIRTICAULE Presl.—Chiapas: Javalinero, in second growth, July 6-9, Matuda 3640. PANICUM ICHNANTHOIDES Fourn.—Chiapas: Palenque, in savanna, July 9-14, Matuda 38722. Panicum LAxumM Swartz—Campeche: Vapor, July 25-28, Matuda 3899. Panicum RupcGeEr Roem. & Schult.—Tabasco: Achotal, east of Balanedin, May 9-14, Matuda 3087. PANICUM ZIZANIOIDES H. B. K.—Campeche: Vapor, July 25—- 28, Matuda 3896. PAspALUM CAESPITosUM Fligge.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3289. PASPALUM LINEARE Trin.—Chiapas: Aguacate, July 16-18, Matuda 3772, 3800. According to Mrs. Chase, this is the first record for the species on the continent north of Costa Rica. PASPALUM ORBICULATUM Poir.—Tabasco: Reforma, May 22- 26, Matuda 3206. 14 CYRUS LONGWORTH LUNDELL PASPALUM PLICATULUM Michx.—Tabasco: Achotal, cast of Balancan, May 9-14, Matuda 3069. PASPALUM VIRGATUM L.—Tabasco: Kstapilla, in savanna, June 27, Matuda 3519. PENTARRHAPHIS SCABRA H. B. K.—Tabasco: Kstapilla, in savanna, June 27, Matuda 3503. Chiapas: Palenque, in savanna, July 9-14, Matuda 3742. PHARUS LATIFOLIUS L.—Chiapas: J avalinero, in advanced forest, July 6-9, Matuda 3604. SETARIA GENICULATA (Lam.) Beauv.—Tabasco: Retiro, river- side, June 19-25, Matuda 3430. SETARIA TENAX (L.) Desv. Matuda 8806. SORGHASTRUM SETOSUM (Griseb.) Hitche.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3733; Aguacate, July 16-18, Matuda 8770, 3778. STREPTOGYNE CRINITA Beauy.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3637. TRACHYPOGON ANGUSTIFOLIUS (H. B. K.) Nees.—Tabasco: Achotal, east of Balancan, May 9-14, Matuda 3105; Wstapilla, in savanna, June 27, Matuda 3506. TRICHACHNE INSULARIS (L.) Nees.—Chiapas: Javalinero, in second growth, swampside, July 6-9, Matuda 3628. Chiapas: Aguacate, July 16-18, CYPERACEAE (Determined by Hugh O'Neill, Washington) CALYPTROCARYA GLOMERULATA (Brongn.) Urban.—Chiapas: Javalinero, at edge of river, July 6-9, Matuda 3610. CLADIUM JAMAICENSE Crantz.—Tabasco: La Palma, riverside, June 1-6, Matuda 3316 (det. Lundell), 2-3 m. high. CYPERUS ARTICULATUS L.—Tabasco: Estapilla, swampside, June 18, Matuda 3483. Campeche: Palizada, in swamp, July 25— 28, Matuda 8868. CYPERUS BREVIFOLIUS Rottb.—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3559. CYPERUS DIGITATUS Roxb.—Campeche: Palizada, in swamp, July 25-28, Matuda 3851. FLORA OF EASTERN TABASCO 15 Cyperus Eacrrsi Boeckl.—Tabasco: Retiro, riverside, June 19-25, Matuda 3441. Cyprrus GIGANTEUS Vahl.—Tabasco: San Isidro, in swamp, June 7-11, Ma/uda 3350, 2.5 m. high. Cyperus HUMILIS Kunth.—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3574. Cyperus LIGULARIS L.—Tabasco: Retiro, riverside, June 19- 25, Matuda 3452. Cyperus Luzuuas (L.) Retz.—Chiapas: Javalinero, swampside, July 6-9, Matuda 3629. Cyperus MACROCEPHALUS Liebm.—Campeche: Palizada, in swamp, July 25-28, Matuda 3869. Cyperus ocHraceus Vahl.—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3578. CypErRus UNIOLIOIDES R. Br.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3744. DicHROMENA coLorata (L.) Hitche—Tabasco: La Palma, riverside, June 1-6, Matuda 3260. ELEOCHARIS CELLULOSA Torr.—Tabasco: La Palma, riverside, June 1-6, Matuda 3259. ELEOCHARIS GENICULATA (L.) R. &. 8.—Tabasco: La Palma, riverside, June 1-6, Matuda 3262. FIMBRISTYLIS MONOsTACHYA (L.) Hassk.—Tabasco: Kstapilla, June 27, Matuda 3497. FurreNA simMpLEx Vahl.—Tabasco: La Palma, riverside, June 1-6, Matuda 3321, 3322; San Isidro, in swamp, May 9-14, Matuda 3360. Hypo.yTruM NICARAGUENSE Liebm.—Tabasco: San Isidro, June 7-11, Matuda 3337. RuyNCHOSPORA CEPHALOTES (L.) Vahl.—Tabasco: Ojo de Agua, southeast of Balancan, in savanna, May 9-14, Matuda 3120 RHYNCHOSPORA CYPHROIDES (Sw.) Mart.—Campeche: Vapor, July 25-28, Matuda 3911. Ruyncnospora rascicuLaris (Michx.) Vahl.—Tabasco: Acho- tal, cast of Balancan, in savanna, May 9-14, Matuda 3057 ; Ojo de Agua, southeast of Balancan, in savanna, May 9-14, Matuda 3128. 16 CYRUS LONGWORTH LUNDELL RHYNCHOSPORA sp.—Tabasco: Achotal, east of Balancdén, in savanna, May 9-14, Matuda 3052. RHYNCHOSPORA sp.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3090. RHYNCHOSPORA sp.—Tabasco: Ojo de Agua, southeast of Bal- ancan, lakeside, May 9-14, Matuda 3127. RHYNCHOSPORA sp.—Tabasco: Provincia, May 29, Matuda 3222. RHYNCHOSPORA sp.—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3539. SCLERIA BRACTEATA Cav.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3092, vernacular name, navajuela. SCLERIA MELALEUCA Reichenb.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3326. SCLERIA MICROCARPA Nees.—Tabasco: Reforma, May 22-26, Matuda 3228. PALMAE ACOELORRAPHE sp.—Tabasco: Achotal, cast of Balanedn, in savanna, May 9-14, Matuda 3070. Bacrris sp.—Tabasco: Reforma, May 22-26, Matuda 3192. Bacrris sp.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3478, 2.5 m. high, 1.5-2 em. in diameter. Bacrris sp.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 8479, 2-2.5 m. high. CHAMAEDOREA sp.—Tabasco: Reforma, in advanced forest, May 22-26, Matuda 3153, 1.5 m. high. CHAMAEDOREA sp.—Tabasco: Reforma, in advanced forest, May 22-26, Matuda 3232, 2 m. high. CHAMAEDOREA sp.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3275, 80-100 em. high. CHAMAEDOREA sp.—T'ahasco: Retiro, in advanced forest, June 19-25, Matuda 3466. JHAMAEDOREA sp.—Chiapas: Javalinero, July 6-9, Matuda 3650, 1-1.3 m. high. DEsMONCUS sp.—Tabasco: Reforma, May 22-26, Matuda 3196. GEONOMA sp.—Tabasco: Retiro, June 19-25, Matuda 8448, FLORA OF EASTERN TABASCO 17 GEONOMA sp.—Tabasco: Retiro, in advanced forest, June 19- 25, Matuda 3477, 3 m. high, 2-3 cm. in diameter. ARACEAE Axruurrum myosurorpes (H. B. K.) Endl—Chiapas: Java- linero, on tree in advanced forest, July 6-9, Matuda 3644. MoNSsTERA TUBERCULATA Lundell, Lloydia, 2: 78. 1939.— Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3308, fruits white, edible, vernacular name, chile montana; Estapilla, June 27, Matuda 3495. This is the first record for the species out- side of British Honduras. Pistia Srratiores L.—Tabasco, San Isidro, aquatic, June T- 11, Matuda 3387. SPATHIPHYLLUM LACUSTRE Lundell, Contrib. Univ. Mich. Herb., 6: 4. 1941.—Tabasco: Ojo de Agua, southeast of Balancan, lake- side, May 9-14, Matuda 3136 (type collection). SyNGONIUM PODOPHYLLUM Schott.—Chiapas: Aguacate, epi- phyte, July 16-18, Matuda 3825. XYRIDACEAE Xyris aMBIGuA Beyr.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3728. No authentic material of the species has been available for comparison, but the specimens from Chiapas agree closely with Malme’s description (North Amer. Fl., 19: 8. 1937). This plant, widespread in the southern United States, has been re- ported from Cuba and British Honduras (tbid.). Xyris Jupicat Rich—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3109, corolla yellow. BROMELIACEAE Arcumea Krenasrit BE. Morr.—Tabasco: San Isidro, on tree at riverside, June 7-11, Matuda 3365. BILLBERGIA vrrIpIFLORA Wendl.—Tabasco: San Isidro, on tree, June 11, Matuda 3400. Caropsis MorRENIANA Mez. Matuda 3394 (det. L. B. Smith). Tabasco: San Isidro, June 11, 18 CYRUS LONGWORTH LUNDELIL PITCAIRNIA PUNICEA Scheidw.—Chiapas: Javalinero, epiphyte, July 6-9, Matuda 3660, 3661. TILLANDSIA BRACHYCAULOS Schlecht.— Campeche: Vapor, epi- phyte, July 25-28, Matuda 3882. TILLANDSIA BULBOSA Hook.—Tabasco: Ref orma, epiphyte, May 22-26, Matuda 3180. TILLANDSIA FESTUCOIDES Brongn.—Tabasco: Reforma, on tree, May 22-26, Matuda 3144; San Isidro, June 7-11, Matuda 3389. TILLANDSIA JUNCEA LeC.—Tabasco: Reforma, epiphyte, May 22-26, Matuda 31898. TILLANDSIA POLYSTACHYA L.—Campeche: Vapor, on tree, July 25-28, Matuda 3880 (det. L. B. Smith). TILLANDSIA ScHIEDEANA Steud.—Tabasco: Reforma, on tree, May 22-26, Matuda 3143. TILLANDSIA STREPTOPHYLLA Scheidw.—Tabasco: La Palma, on riverside tree, June 1-6, Matuda 3305. TILLANDSIA USNEOIDES L.—Chiapas: Aguacate, on tree, July 16-18, Matuda 8812. Campeche: Palizada, on tree at lakeside, July 25-28, Matuda 3828. COMMELINACEAE COMMELINA ELEGANS H. B. K.—Chiapas: Aguacate, July 16- 18, Matuda 3802, flowers blue. PONTEDERIACEAE PONTEDERIA LANCEOLATA Nutt. (?).—Tabasco: La Palma, edge of lake, June 1-6, Matuda 3302. Chiapas: ruins of Palenque, July 10-12, Matuda 3688. ZOSTERELLA LONGITUBA Alexander, North Amer. Fl., 19: 52. 1937.—Campeche: Vapor, in lake, July 25-28, Matuda 8872, flowers yellow. LILIACEAE DRACAENA AMERICANA Donn, Sm.—Tabasco: Tenosique, in ad- vanced forest, June 14-16, Matuda 3404, tree 8-10 m. high, 40 em. in diameter. FLORA OF EASTERN TABASCO 19 SMILACACKAE (Verified by C. V. Morton, Washington) SMILAX ARISTOLOCHIAEFOLIA Mill.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3015. SMILAX LANCEOLATA L.—Chiapas: ruins of Palenque, at edge of forest, July 10-12, Matuda 3674. Sminax LucULENTA Killip & Morton, Carnegie Inst. Wash. Publ., 461: 289. 1936.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3709. Sminax MoLtLIs H. & B.—Tabasco: Achotal, east of Balancan, in advanced forest, May 9-14, Matuda 3072; Estapilla, June 18, Matuda 3487. Chiapas: Aguacate, July 9-14, Matuda 3762. Smriuax spinosa Mill.—Tabasco: Ojo de Agua, southeast of Balancan, in second growth, May 9-14, Matuda 3135; Reforma, May 22-26, Matuda 3159, 3177, 3199; Boca del Cerro, in advanced forest, July 1-5, Matuda 3556. HAEMODORACEAE XIPHIDIUM CAERULEUM Aubl.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3701, flowers white. AMARYLLIDACEAE CURCULIGO SCORZONERIFOLIA (Lam.) Baker.—Chiapas: Agua- cate, July 16-18, Matuda 3794, perennial herb. HyMENOCALLIS LITTORALIS (Jacq.) Salisb.—Chiapas: Aguacate, in wet savanna, July 16-18, Matuda 3797. Campeche: Vapor, in swamp, July 25-28, Matuda 3885. ZEPHYRANTHES CITRINA Baker (?).—Tabasco: Tenosique, in second growth, June 14-16, Matuda 3407, flowers white. DIOSCOREACEAE DioscorrA Bartrierri Morton, Carnegie Inst. Wash. Publ. 461: 242. 1936.—-Tabasco: La Palma, in second growth, June 1-6; Matuda 3318. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3669, a vine. DrogscorEA BERNOULLIANA Prain & Burk.—Chiapas: Aguacate, in savanna, July 16-18, Matuda 3767. 20 CYRUS LONGWORTH LUNDELL IRIDACEAE Crpura PALUDOosA Aubl.—Tabasco: Estapilla, in savanna, June 27, Matuda 3511, flowers white. Chiapas: Palenque, in savanna, July 9-14, Matuda 3729, flowers white. Marica Gracinis Herb.—Tabasco: La Palma, June 1-6, Matuda 8281. NeMASTYLIS Buquarrti Standl., Journ. Arnold AD Lis a7, 1930.—Chiapas: Aguacate, July 16-18, Matuda 3815. MUSACEAE (Determined by L. B. Smith, Cambridge) HELICONIA AURANTIACA Ghiesbr.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3885, HELIcONIA CouLinstana Griggs (?).—Chiapas: Aguacate, in advanced forest, July 16-18, Matuda 3828, flowers scarlet. HELICONIA crassa Griggs.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3385A. This species probably is not distinct from H. aurantiaca Ghiesbr. HELICONIA HIRSUTA var. CANNOIDEA (L. C. Rich.) Bak.— Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3639, flowers red. HELIcONIA LATISPATHA Benth.—Tabasco: Estapilla, June 27, Matuda 8494. Heiconia Liprara Griggs.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3465. Heviconia Scuteprana Kl. (?)—Tabasco: Boca del Cerro, riverside, July 1-5, Matuda 3541. Heiconra sp.—Chiapas: ruins of Palenque, in forest, July 10-12, Matuda 3683. ZINGIBERACEAE Costus VILLosisstmus Jacq.—Tabasco: Reforma, May 22-26, Matuda 3218. RENEALMIA AROMATICA (Aubl.) Griseb.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3386. FLORA OF EASTERN TABASCO 21 MARANTACHEAE MARANTA ARUNDINACEA L.—Chiapas: Aguacate, July 16-18, Matuda 3791. GALATHEA MACROCHLAMYs Woodson & Standl., Field Mus. Bot., 22: 134. 1940.—Tabasco: Tenosique, in advanced forest, June 14-16, Matuda 3399 (type collection), flowers yellowish. CALATHEA mIcANS (Math.) Koern.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3418, flowers white. PLEIOSTACHYA PRUINOSA (Regel) Schum.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 8680. BURMANNIACEAE BurMANNIA CAPITAtTA (Walt.) Mart.—Chiapas: Aguacate, in savanna, July 16-18, Matuda 3778. ORCHIDACEAE (Determined by Charles Schweinfurth, Cambridge) BRASSAVOLA Noposa (L.) Lindl.—Campeche: Vapor, on tree, July 25-28, Matuda 3879, flowers white. CATASETUM INTEGERRIMUM Hook.—Campeche: Palizada, on tree, July 25-28, Matuda 3830. Epipenprum aLatum Batem.—Tabasco: Ojo de Agua, south- east of Balancdn, May 9-14, Matuda 3111, flowers reddish, fra- erant. Campeche: Vapor, on tree, May 25-28, Matuda 3898. Epmenprum cocHugatum L.—Tabasco: La Palma, on tree in advanced forest, June 1-6, Matuda 33038. Epipenprum Dram Schlitr.—Tabasco: Reforma, on tree, May 22-26, Matuda 3163. Campeche: Vapor, on tree, July 25-28, Matuda 3900. EpmeNnpRUM DIFFORME Jacq.—Tabasco: Reforma, May 22-26, Matuda 3165, flowers yellow. Campeche: Vapor, on tree, July 25—- 28, Matuda 3878. EpripENDRUM IMATOPHYLLUM Lindl.—Tabasco: Balancén, May 20, Matuda 3179. EpipENDRUM NocTURNUM Jacq.—Campeche: Palizada, July 95-28, Matuda 3826. 22 CYRUS LONGWORTH LUNDELL HPIDENDRUM POLYANTHUM Lindl.—Tabasco: San Isidro, lake- side, June 7-11, Matuda 8849. IK}PIDENDRUM RADIATUM Lindl.—Chiapas: Aguacate, on tree in savanna forest, June 16-18, Matuda 3824, flowers fragrant. E}PIDENDRUM RIGIDUM Jacq.—Tabasco: Ojo de Agua, south- east of Balancdn, in advanced forest, May 9-14, Matuda 3114; La Palma, on riverside tree, June 1-6, Matuda 3317. IoNoPsIs UTRICULARIOIDES (Sw.) Lindl.—Tabasco: Retiro, on tree at riverside, June 19-25, Matuda 3464. Chiapas: Javalinero, on tree, July 6-9, Matuda 3646. Oncipium LinpENTI Brongn.—Tabasco: Reforma, on_ tree, May 22-26, Matuda 3230. ONCIDIUM PUSILLUM (L.) Rchb. f.—Tabasco: San Isidro, lake- side, June 7-11, Matuda 3352. PLEUROTHALLIS Grosyt Lindl. P. marginata Lindl.—Tabasco: Reforma, on tree, May 22-26, Matuda 3150, flowers white. Cam- peche: Vapor, July 25-28, Matuda 3895. CHOMBURGKIA TIBICINIS Batem.—Tabasco: Balancdn, May 18, Matuda 31454. PIPERACEAE PEPEROMIA CHICBULANA Trel., Field Mus. Bot., 17: 230. 1937.— Tabasco: San Isidro, June 7-11, Matuda 3370. Peperomia Lunpexui Trel., Field Mus. Bot., 12: 406. 1936.— Tabasco: Reforma, May 22-26, Matuda 3164. This species is very near if not conspecific with P. glutinosa Millsp. PrPEROMIA PETENENSIS Trel., Field Mus. Bot., 17: 230. 1937.— Tabasco: Reforma, epiphyte, May 22-26, Matuda 8214. PEPEROMIA TREMULAEFORMIS Trel., Field Mus. Bot., 9: 276. 1940.—Chiapas: Palenque, on trees in advanced forest along river, July 10-12, Matuda 3671. PIPER ANDRESENSE Trel., Field Mus. Bot., 17: 280. 1937.— Tabasco: Achotal, east of Balancan, in advanced forest, May 9-14, Matuda 3065. The collection agrees so closely with the isotype of P. andresense from Petén (Lundell 3203) that I do not hesitate to refer it to this species. PIPER auRITUM H. B. K.—Tabasco: La Palma, June 1-6, Matuda 3234. FLORA OF EASTERN TABASCO a Prrer HIspiIpuM Sw.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3411. Prrer Marupat Lundell, Lloydia, 4: 44. 1941.—Chiapas: along Chacamax River at Palenque, July 10-12, Matuda 8697 (type collection), spikes white. Piper psItoRHAcHE C. DC.—Tabasco: La Palma, in forest, June 1-6, Matuda 3246, a shrub. PIPER USUMACINTENSE Lundell, Lloydia, 4: 44. 1941.—Tabasco: Boca del Cerro, in advanced forest, July 1-5, Matuda 8589 (type collection), a shrub. PIPER YUCATANENSE C. DC.—Tabasco: San Isidro, in ad- vanced forest, June 7-11, Matuda 3382, a shrub. SALICACEAE Saprx cHtteNnsis Molina.—Campcche: Palizada, July 25-28, Matuda 3856. FAGACEAE QuERCUS OLEOIDES Cham. & Schlecht.—Tabasco: Ojo de Agua, southeast of Balancdn, in advanced forest, May 9-14, Matuda 3129, a tree, 13 m. high, 60 cm. in diameter. Chiapas: Palenque, in savanna, July 9-14, Matuda 3782, a tree, 15 m. high, 55-60 cm. in diameter, vernacular name, roble. ULMACEAE TREMA FLORIDANA Britton.—Tabasco: Boca del Cerro, in sec- ond growth, July 1-5, Matuda 3575. MORACEAE Crcropra sp.—Tabasco: La Palma, June 1-6, Matuda 3240. CHLOROPHORA TINCTORIA (L.) Gaud—Campeche: Palizada, in swampside forest, July 25-28, Matuda 3850, a tree, 5-6 m. high. 30 cm. in diameter. CovussAPOA OLIGOCEPHALA Donn. Sm.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3300, a tree, 20 m. high, 45 cm. in diameter. Dorstenta Contrragerva L.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3276. 24 CYRUS LONGWORTH LUNDELL Ficus invoutura (Liebm.) Miq.—Tabasco: Boca del Cerro, swampside, July 1-5, Matuda 3552, a tree, 12 m. high, 50 em. in diameter. Ficus Orrsrepiana Miq.—Chiapas: Palenque, in savanna forest, July 9-14, Matuda 3724, a tree, 10 m. high, 50 em. in di- ameter; Aguacate, in savanna forest, J uly 16-18, Matuda 3805, a tree, 15 m. high, 60 em. in diameter. Ficus papirotia H. B. K.—Tabasco: Reforma, May 22-26, Matuda 3176, a tree. Ficus rapuLa Willd.—Tabasco: Reforma, riverside, May 22- 26, Matuda 3162, a tree, 10 m. high, 45 em. in diameter; San Isidro, lakeside, June 7-11, Matuda 3346, a tree, 14 m. high, 45 cm. in diameter. Ficus Turerckyermi Standl.—Tabasco: La Palma, riverside, June 1-6, Matuda 3250, a tree, 20 m. high, 60 em. in diameter. PIRATINERA TERRABANA (Pittier) Lundell, Carnegie Inst. Wash. Publ., 478: 208. 1937.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3625, a tree, 10 m. high, 25 em. in diameter. POULSENIA ARMATA (Miq.) Standl.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3645, a tree, 15 m. high, 30 cm. in diameter. URTICACEAE (Determined by E. P. Killip, Washington) BOrHMERIA CYLINDRICA (L.) Sw.—Tabasco: San Isidro, in swamp, June 7-11, Matuda 3357. BOEHMERIA RAMIFLORA var. CUSPIDATA Wedd.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3441. PILEA puUBESCENS Liebm.—Tabasco: Retiro, in virgin forest, June 19-25, Matuda 3462. Povuzouzta OBLIQUA Wedd.—Tabasco: Retiro, in second growth, June 19-25, Matuda 8475 (det. Lundell). URERA ELATA (Sw.) Griseb.— Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3699, a small tree, 3 m. high. LORANTHACEAE PHORADENDRON CHEIROCARPUM Trel. in savanna forest, July 9-14, Matuda 3727. Chiapas: Palenque, FLORA OF EASTERN TABASCO 20 PHORADENDRON FRANCISCANUM Trel., Field Mus. Bol., 17: 236. 1937.—Campeche: Palizada, on tree, July 25-28, Matuda 3849. PHORADENDRON PIPEROIDEs (H. B. K.) Trel—Chiapas: Palen- que, on tree in savanna forest, J uly 9-14, Matuda 3721. PsirraACANTHUS AMERICANUS (Jacq.) Mart.—Campeche: Vapor, on tree, July 25-28, Matuda 3898, flowers red. SrPRUTHANTHUS cassyTHoIDES Millsp., Field Mus. Bot., Seis 1930.—Tabasco: Reforma, May 22-26, Matuda 3200. SPRUTHANTHUS MACRosTAcHYUS Lundell, Lloydia, 4: 45. 1941. —Chiapas: Palenque, in savanna forest, July 9-14, Matuda 3731 (type collection). QrRUTHANTHUS PHANERONEURUS Standl., Field Mus. Bot., 9: 285. 1940.—Campeche: Palizada, on tree, July 25-28, Matuda 3856. The Campeche collection considerably extends the range of this parasite, which was recently described from Honduras. The original description, based evidently on imperfect material, may now be amplified. Leaves obovate-elliptic or elliptic, up to 6 cm. long, 3.0 cm. wide. Inflorescences axillary, solitary or fasciculate, narrowly paniculate, usually sessile, up to 5 em. long. Flowers sessile or subsessile, about 4.2 mm. long, in clusters of 3, the clusters borne on peduncles about 1.5 mm. long. Bractlets deciduous. Calyx truncate, obscurely denticulate. Petals 6, linear, 3.4 mm. long. Style straight, 3 mm. long. OLACACEAE Hersteria MEpIA Blake, Contrib. U. S. Nat. Herb., 24:3. 1922. H. Chippiana Standl., Field Mus. Bot., 11: 130. 1932.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3690, a tree, 20 m. high, 40 cm. in diameter. XIMENIA AMERICANA L.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3074, 3075, a shrub, 2 m. high. BALANOPHORACEAE Herosis MEXICANA Liebm.—Tabasco: Boca del Cerro, in ad- vanced forest, July 1-5, Matuda 3448. 26 CYRUS LONGWORTH LUNDELL POLYGONACEAE CoccoLoBA COZUMELENSIS Hemsl.—Tabasco: Reforma, May 22-26, Matuda 3173, a tree, 15 m. high, 35 em. in diameter. CoccoLoBa MAYANA Lundell, Bull. Torrey Bot. Club, 64: 547. 1937,—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3634, a tree, 12 m. high, 50 cm. in diameter. GYMNOPODIUM FLORIBUNDUM Rolfe (?).— Tabasco: Reforma, May 22-26, Matuda 3182, a small tree. POLYGONUM MEXICANUM Small.—Tabasco: Boca del Cerro, swampside, July 1-5, Matuda 3544. CHENOPODIACEAE CHENOPODIUM AMBROSIOIDES L.—Tabasco: La Palma, in sec- ond growth, June 1-6, Matuda 3256. AMARANTHACEAE ACHYRANTHES ASPERA L.—Tabasco: La Palma, in forest, June 1-6, Matuda 3294. TRESINE NIGRA Uline & Bray (?).—Tabasco: Reforma, May 26, Matuda 8208. NYCTAGINACEAE NEEA BELIZENSIS Lundell, Contrib. Univ. Mich. Herb., 7: 9. 1942.—Tabasco: Achotal, east of Balancdn, in advanced forest, May 9-14, Matuda 3076, a shrub, 50 em. high. PHYTOLACCACEAE ACHATOCARPUS MEXICANUS H. Walt.—Tabasco: Kstapilla, riverside, June 18, Matuda 38489, a shrub. Rivina numiuis L.—Tabasco: Retiro, riverside, June 19-25, Matuda 3446. Campeche: Palizada, July 25-28, Matuda 3852. NYMPHAEACEAE CABOMBA AQuatica Aubl.—Tabasco: Reforma, May 22-26, Matuda 3191. Campeche: Vapor, in lake, July 25-28, Matuda 3S74, NeLUMBO LuTEA (Willd.) Pers. (?).—Campeche: Palizada, in lake, July 25-28, Matuda 3833, flowers white. FLORA OF EASTERN TABASCO Path NYMPHAEA AMPLA (Salisb.) DC.—Tabasco: Reforma, May 22- 26, Matuda 3190, flowers white. Campeche: Vapor, in lake, July 25-28, Matuda 3877. MENISPERMACEAE CigsAMPELOS PAretrA L.—Tabasco: Achotal, east of Balancan, in second growth, May 9-14, Matuda 3058. Chiapas: Aguacate, July 16-18, Matuda 3809, a vine. CISSAMPELOS TROPAEOLIFOLIA DC.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3457. Chiapas: ruins of Pa- lenque, in second growth, July 10-12, Matuda 3686. HyrerBAENA NECTANDRIFOLIA Standl.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3380, a trec, 10 m. high, 25 cm. in diameter. ANONACEAE CYMBOPETALUM PENDULIFLORUM (Dunal) Baill—Tabasco: La Palma, June 1-6, Matuda 3237. GUATTERIA AMPLIFOLIA Triana & Planch.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3684, 3685, a small tree, flowers yellow. XyYLOPIA FRUTESCENS Aubl.—Tabasco: Achotal, east of Balan- cin, in advanced forest, May 9-14, Matuda 3094, 80965, a tree, 13 m. high, 25 em. in diameter. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 8607, a tree, 15 m. high, 30 cm. in diameter. MONIMIACKAE SIPARUNA NICARAGUENSIS Hemsl.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 8413; Boca del Cerro, July 1-5, Matuda 3558. LAURACEAI CAssyTHA FILIFORMIS L.—Chiapas: Aguacate, in savanna, July 16-18, Matuda 3780. NEcTANDRA PERDUBIA Lundell, Lloydia, 4:47. 1941. —Tabasco: Boca del Cerro, in second growth, July 1-5, Matuda 3576 (type collection), a tree, 10 m. high, 25 cm. in diameter. NECTANDRA SANGUINEA Rottb.—Tabasco: Mercedes, in second erowth along river, May 9-14, Matuda 3027, a tree, 4-5 m. high, 28 CYRUS LONGWORTH LUNDELL 10-15 em. in diameter; Tenosique, in advanced forest, June 14— 16, Matuda 3402, a tree, 7 m. high, 39 em. in diameter, flowers white. Although the flowers are smaller than those typical for this tree, the species is quite variable, and the collections appear to be referable here. NECTANDRA TABASCENSIS Lundell, Lleydia, 4: 48. 1941.— Tabasco: La Palma, June 1-6, Matuda 3299 (type collection), a tree. PHOEBE LONGICAUDATA Lundell, Bull. Torrey Bot. Club, 64: 548. 1937.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3648, a small tree, 3-4 m. high. CAPPARIDACEAE Capparis TupRCKHEIMII Donn. Sm.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3099, a tree, 2-3 m. high, flow- ers white; Reforma, May 22-26, Matuda 3185, a shrub. CRATAEVA GLAUCA Lundell, Bull. Torrey Bot. Club, 69: 389. 1942.—Tabasco: Boca del Cerro, at edge of swamp, July 1-5, Matuda 3588, a tree, 5 m. high, 25 em. in diameter. Campeche: Palizada, swampside, July 25-28, Matuda 3863 (type collec- tion), a small tree, 7m. high, 30 em. in diameter, flowers greenish yellow, vernacular name, crustta. DROSERACEAE DROSERA CAPILLARIS Poir.—Veracruz: Minatitlan, lakeside, May 1, Matuda 3023. Tabasco: La Lucha, south of Balancdn, in savanna, May 20, Matuda 3175. ROSACEAE CHRYSOBALANUS Icaco L.—Tabasco: Ojo de Agua, southeast of Balancén, in savanna, May 9-14, Matuda 3182, vernacular name, cacco. CouFPIA DODECANDRA (DC.) Hemsl.—Tabasco: Achotal, east of Balancan, in advanced forest, May 9-14, Matuda 3102, a tree, 15-18 m. high, 35 em. in diameter; Boca del Cerro, riverside, July 1-5, Matuda 3585, a tree, 15 m. high, 55 em. in diameter. Chiapas: Aguacate, in savanna, July 16-18, Matuda 3798. HIRTELLA AMERICANA L.—Tabasco: La Palma, in advanced FLORA OF EASTERN TABASCO 29 forest, June 1-6, Matuda 3270, a tree, 20 m. high, 55 cm. in diameter, vernacular name, icatillo ; San Isidro, in advanced forest, June 7-11, Matuda 3343, a tree, 20 m. high, 40 em. in diameter. HirreLta RAcEMosA Lam.—Tabasco: Achotal, east of Balan- cn, in second growth, May 9-14, Matuda 3097. LICANIA HYPOLEUCA Benth.—Tabasco: Achotal, east of Balan- cAn, in advanced forest, May 9-14, Matuda 3054, a tree, 3-4 m. high; Ojo de Agua, southeast of Balancdn, in advanced forest, May 9-14, Matuda 3110, a tree, 7-8 m. high, 25 cm. in diameter; Ojo de Agua, Matuda 3142, a tree, 10 m. high, 35-40 em. in diame- ter; Boca del Cerro, in advanced forest, July 1-5, Matuda 3565, a tree, 15 m. high, 40 cm. in diameter. CONNARACEAE Rovurea GLABRA H. B. K.—Tabasco: Mercedes, lakeside, May 9-14, Matuda 3004, a woody vine, flowers white. LEGU MINOSAI: J. MIMOSEAE Acacta FARNESIANA (L.) Willd—Tabasco: Boca del Cerro, in second growth, July 1-5, Matuda 3587, 3588, a shrub. ACACIA MAYANA Lundell, Carnegie Inst. Wash. Publ., 478: 210. 1937.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3410, a shrub, 3 m. high. This exceptionally well-marked species has been known only from the type collection made in southern Petén by Mercedes Aguilar H. ACACIA USUMACINTENSIS Lundell, Contrib. Univ. Mich. Herb., 4. 1940.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3323, 3324, a tree, 15 m. high, 45 cm. in diameter; Boca del Cerro, July 1-5, Matuda 3540 (type collection), a tree, 14 m. high, 40 em. in diameter. ALBIZZIA TOMENTOSA (Micheli) Standl.—Tabasco: La Palma, at edge of forest, June 1-6, Matuda 3267 (det. Morton), a tree, 13 m. high, 55 cm. in diameter. CaLLIANDRA Dramu (B. & R.) Standl—Tabasco: Reforma, May 22-26, Matuda 3202, a shrub, flowers red; San Isidro, in ad- vanced forest, June 7-11, Matuda 3340, a tree, 3m. high, 9 cm. in diameter. C. Deamii and C. yucatanensis are quite variable, 30 CYRUS LONGWORTH LUNDELI especially in pubescence. When additional collections become available, a further division of the complexes may be desirable, CALLIANDRA YUCATANENSIS (B. & R.) Standl.—Campeche: Palizada, July 25-28, Matuda 3871, a shrub, flowers red. INGA EDULIS Mart.—Tabasco: Achotal, east of Balancén, May 9-14, Matuda 3088, a tree, 3-4 m. high, 30 em. in diameter. INGA LEProLoBA Schlecht.—Tabasco: Boca del Cerro, swamp- side, July 1-5, Matuda 3548, a tree, 10 m. high, 40 cm. in diameter. INGA PINETORUM Pittier.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3098, a shrub, 1.5 m. high. InGa puncrata Willd.—Tabasco: Reforma, May 22-26, Ma- tuda 8220, a tree. LYsSILOMA DESMosTAcHys Benth.—Tabasco: Estapilla, edge of savanna, June 27, Matuda 3520, a tree, 10 m. high, 45 em. in diameter. MiImMosA SOMNIANS Humb. & Bonpl.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8734. NEPTUNIA PUBESCENS Benth.—Chiapas: Aguacate, Ju 18, Matuda 3810, flowers yellow. PITHECOLOBIUM ALBICANS (kunth) Benth. Albizzia rubiginosa Standl., Carnegie Inst. Wash. Publ., 461: 58. 1935, not Miquel, 1855. Albtzzia Lundellii Standl., Trop. Woods, 52: 26. 1937,.— Chiapas: Aguacate, in savanna, July 16-18, Matuda 3801, a tree, 10 m. high. Campeche: Vapor, July 25-28, Matuda 3912, a tree, 7m. high, 30 em. in diameter, flowers white. PITHECOLOBIUM ARBOREUM (L.) Urban.— Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3290, a tree, 10 m. high, 40 cm. in diameter. ->ITHECOLOBIUM BELIZENSE Standl., Field Mus. Bot., 4: 212. 1929.— Tabasco: Tenosique, riverside, June 14-16, Matuda 3409, a tree, 7m. high, 40 em. in diameter, PITHECOLOBIUM CAMPECHENSE Lundell, Phytologia, 1: 481. 1941.—Campeche: Palizada, in swampside forest, July 25-28, Matuda 3866 (type collection), a tree, 7-10 m. high, 30-60 em. in diameter, vernacular name, tinta. PITHECOLOBIUM DISCIFERUM Lundell, Contrib. Univ. Mich. Herb., 6: 27. 1941.—Tabasco: Reforma, riverside, May 22-26, Matuda 3211, a tree, 10 m. high, 380 cm. in diameter; Retiro, on ome y 16- FLORA OF EASTERN TABASCO 31 limestone rock at riverside, June 19-25, Matuda 3468, a shrub. Campeche: Vapor, July 25-28, Matuda 3908 (type collection), a tree, 10-15 m. high, 30-40 cm. in diameter. PrrHEcoLtoBium ENGLEsINGu (Standl.) Standl., Trop. Woods, 34: 40. 1933. Inga Englesingii Standl., Trop. Woods, 17: 27. 1929; Field Mus. Bot., 4: 211. 1929—Chiapas: ruins of Palenque, in ad- vanced forest, July 10-12, Matuda 3665, a tree, 15 m. high, 35-40 em. in diameter. PirHEcoLoBIuM GENTLEI Lundell, Contrib. Univ. Mich. Herb., 6: 28. 1941.—Chiapas: Javalinero, in the Chacamax River, July 6-9, Matuda 3620, a shrub, 2 m. high, flowers white; ruins of Palenque, riverside, in forest, July 10-12, Matuda 3693, a shrub. PITHECOLOBIUM LEUCOCALYX (B. & R.) Standl., Field Mus. Bot., 4: 308. 1929. Samanea leucocalyx B. & R., N. Amer. FL, 23: 34, 1928.—Tabasco: Estapilla, riverside, June 18, Matuda 3486, a tree, 15 m. high, 60 cm. in diameter. PirHecoLospium pacuypus Pittier—Campeche: Palizada, swampside, July 25-29, Matuda 3837, 3845, a shrub or tree, up to 10 m. high, 60 cm. in diameter, vernacular name, lokin. Il. CAESALPINIEAE BAUHINIA pivaricata L.—Tabasco: Tenosique, in second growth, June 14-16, Matuda 3397, flowers white. BAUHINIA PALENQUENSIS Lundell, Contrib. Univ. Mich. Herb., 6: 24. 1941.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3666 (type collection), a small tree, 3m. high, flowers red. CAssIA BACILLARIS L—Tabasco: Boca del Cerro, in second growth, July 1-5, Matuda 3593. Cassia DIPHYLLA L.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3051, flowers yellow; Ojo de Agua, southeast of Balancdn, in savanna, May 9-14, Matuda 3122. Chiapas: Aguacate, July 16-18, Matuda 3793. CassiA FLExUOSA L.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3068. Cassia GraNnpIs L—Tabasco: Boca del Cerro, swampside, July 1-5, Matuda 3536, a tree, 15 m. high, 50 cm. in diameter. CAssta HISPIDULA Vahl.—Tabasco: Achotal, east of Balancan, 32 CYRUS LONGWORTH LUNDELL in savanna, May 9-14, Matuda 3045; Estapilla, June 27, Matuda 8521, Cassia Kinuirn Rose.—Chiapas: Aguacate, July 16-18, Ma- tuda 8788. Cassia Tacera L.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8740, 3751. CAssIA UNDULATA Benth.—Tabasco: Achotal, east of Balancan, in second growth, May 9-14, Matuda 3035. CYNOMETRA COLIMENSIS Britt. & Rose.—Chiapas: Aguacate, swampside, July 16-18, Matuda 3796 (det. Morton), a tree, 15 m. high, 35 em. in diameter. DIALIUM GUIANENSE (Aubl.) Sandwith.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3248, a tree, 15 m. high, 45 cm. in diameter. Swarrzia LuNDELLU Standl., Carnegie Inst. Wash. Publ., 461: 61. 1935.—Tabasco: Boca del Cerro, on limestone, July 1-5, Ma- tuda 3586, a tree, 12 m. high, 35 em. in diameter. III. PApInionaTAE ANpDIRA GALHOTTIANA Standl., Contrib. U. S. Nat. Herb., 20: 217. 1919.—Tabasco: Ojo de Agua, southeast of Balancdn, in ad- vanced forest, May 9-14, Matuda 3125, a tree, 10 m. high, 55-60 cm. in diameter. ANDIRA INERMIS (Sw.) H. B. K.—Tabasco: Boca del Cerro, July 1-5, Matuda 3569, a tree, 15 m. high, 39 em. in diameter. CENTROSEMA UNIFOLIATUM (Rose) Lundell, Bull. Torrey Bol. Club, 69: 391. 1942.—Tabasco: Estapilla, in savanna, June 27, Matuda 3498, a vine, corolla blue. CLITORIA RUBIGINOSA Juss.— Chiapas: Aguacate, July 16-18, Matuda 8786, a perennial herb, corolla white. CROTALARIA RETUSA L.—Campeche: Palizada, July 25-28, Matuda 8834. CROTALARIA SAGITTALIs L. (?).—Tabasco: Estapilla, in savanna, June 27, Matuda 3527, a perennial herb, corolla yellow. DaLBEerGIA Browne (Jacq.) Urban.—Campeche: Palizada, swampside, July 25-28, Matuda 3870, a woody vine, vernacular name, balbasco. FLORA OF EASTERN TABASCO So DALBERGIA GLABRA (Mill.) Standl—Tabasco: Reforma, May 22-26, Matuda 3195, a woody vine, corolla white; Retiro, river- side on limestone, June 19-25, Matuda 3444. Campeche: Pali- zada, July 25-28, Matuda 3865, a shrub, corolla white. DALBERGIA PAUCIFOLIOLATA Lundell, Contrib. Univ. Mich. Herb., 4: 8. 1940.—Tabasco: Reforma, riverside, May 22-26, Matuda 3167 (type collection), a small tree, corolla white. DrsmMopium BARBATUM (L.) Benth. «& Oerst.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3091. Chiapas: Palenque, in savanna, July 9-14, Matuda 3746. DESMODIUM FRUTESCENS (Jacq.) Schindl. (?).—Tabasco: Re- forma, May 22-26, Matuda 3221, a perennial herb. Chiapas: Palenque, in sayanna, July 9-14, Matuda ays. Drepanocarpus Lunatus (L.f.) Meyer.—Tabasco: Boca del Cerro, swampside, July 1-5, Matuda 3549, 3546, a woody vine, 20 cm. in diameter. ERioseMa pirrusum (H. B. K.) Don.—Tabasco: Achotal, east of Balanedn, in savanna, May 9-14, M atuda 8048, Estapilla, in savanna, June 27, Matuda 3526, corolla yellow. Chiapas: Agua- cate, July 16-18, Matuda 3768, 3817, a perennial herb. GaLAcTIA sTRIATA (Jacq.) Urb.—Chiapas: Aguacate, July 16-18, Matuda 3783, a vine. INDIGOFERA LESPEDEZIOIDES H. B. K.—Tabasco: Estapilla, in savanna, June 27, Matuda 3499, corolla salmon. INDIGOFERA suFFRUTICOSA Mill.—Tabasco: Estapilla, river- side, June 18, Matuda 3488. Campeche: Palizada, July 25-28, Matuda 3862. Loncnocarpus Castiuuor Standl., Trop. Woods, 32: 15. 1932. Tabasco: Reforma, May 22-26, Matuda 3217, a tree, Lier high, 35 cm. in diameter. LONCHOCARPUS HONDURENSIS Benth.—Tabasco: La Palma, riverside, June 1-6, Matuda 3249, a tree, 15 m. high, 45 cm. in diameter. LoNcHOCARPUS LATIFOLIUS (Willd.) H. B. K.—Tabasco: Re- forma, May 22-26, Matuda 3186, a tree, 13 m. high, 40 cm. in diameter; Estapilla, riverside, June 18, Matuda 3482, a tree, 15 m. high, 40 em. in diameter. Campeche: Palizada, July 25-28, 34 CYRUS LONGWORTH LUNDELL Matuda 3886, a tree, 7-10 m. high, 30-50 em. in diameter. MACHAERIUM sEeTULOsUM Pittier (?).—Campeche: Palizada, in swamp, July 25-28, Matuda 3841, & woody vine. PHASEOLUS GRACILIS Poepp. (?).—Tabasco: Estapilla, in sa- vanna, June 25, Matuda 3516. Chiapas: Palenque, in savanna, July 9-14, Matuda 3761, a vine, corolla salmon; Aguacate, July 16-18, Matuda 3775, corolla salmon. PIscIpIA PIscIPULA (L.) Sarg.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3012, a tree, 8-10 m. high, 35 em. in diameter. RHYNCHOSIA MINIMA (L.) DC.—Campeche: Palizada, July 25-28, Matuda 3838, a vine. STYLOSANTHES GUYANENSIS (Aubl.) Swartz.—Tabasco: Acho- tal, east of Balancdn, in savanna, May 9-14, Matuda 3041, 8047, corolla yellow. Chiapas: Palenque, in savanna, July 9-14, Ma- tuda 8735. More than one species may be represented. TEPHROSIA LITTORALIS (L.) Pers. (?).—Tabasco: Estapilla, in savanna, June 27, Matuda 3517. TEPHROSIA NITENS Benth.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3758, VIGNA REPENS (L.) Kuntze.—Tabasco: La Palma, June 1-6, Matuda 3261, a vine. VIGNA VEXILLATA (L.) A. Rich. (?).—Tabasco: Boea del Cerro, swampside, July 1-5, Matuda 8594. ZORNIA DIPHYLLA (L.) Pers.—Chiapas: Aguacate, July 16-18, Matuda 3821. OXALIDACEAE OxaLis NEAEI DC.—Chiapas: Aguacate, July 16-18, Matuda 3819. OXALIS YUCATANENSIS (Rose) R. Knuth, Notézbl. Bot. Gart. Berlin, 7: 291. 1919. Lotoxalis yucaltanensis Rose, Contrib. U.S. Nat. Herb., 10: 116. 1906. Oxalis yucalanensis (Rose) Standl., Contrib. U.S. Nat. Herb. 23: 518, 1923.—Chiapas: Aguacate, July 16-18, Matuda 3820. ERYTHROXYLACBRAR ERYTHROXYLON spp.—Tabasco: San Isidro, swampside, June 7-11, Matuda 8383, a small tree, 3m. high; Boca del Cerro. FLORA OF EASTERN TABASCO 35 swampside, July 1-5, Matuda 3549, a small tree, 4 m. high. Chiapas: ruins of Palenque, in advanced forest along river, July 10-12, Matuda 3706, a small tree, 5 m. high. Campeche: Vapor, swampside, July 25-28, Matuda 3894, a shrub, 4 m. high. At least two species belonging to the complex of EL. areolatum L. are represented. RUTACEAE TANTHOXYLUM CARIBAEUM Lam. (?).—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3007, a small tree. ZANTHOXYLUM NiIGRIPUNCTATUM Lundell, Bull. Torrey Bot. Club, 64: 551. 1937.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3708, a tree, 8 m. high, 30 cm. in diameter. SIMARUBACEAE PICRAMNIA ANTIDESMA Sw.—Tabasco: Boca del Cerro, in ad- vanced forest, July 1-5, Matuda 35738, a small tree, 7-8 m. high. SIMARUBA GLAUCA DC.—Tabasco: Ojo de Agua, southeast of Balancdn, May 9-14, Matuda 3116. BURSERACEAE Bursera Srmarusa (L.) Sarg.—Tabasco: Reforma, May 22- 26, Matuda 3204, vernacular name, palo retinto. Prorium Copau (Schl. & Cham.) Engler.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3414, a tree, 20 m. high, 40 em. in diameter. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3698, a tree, 8 m. high, 35 cm. in diameter. MELIACEAE CEDRELA MEXICANA M. Roem.—Tabasco: Hstapilla, riverside, June 18, Matuda 3492, a tree, 20 m. high, 70 em. in diameter, ver- nacular name, cedro. GuAREA ExceLsA H. B. K.—Tabasco: Achotal, east of Ba- lancén, in advanced forest, May 9-14, Matuda 3085, a tree, 4 m. high, fruits red; La Palma, in advanced forest, June 1-6, Matuda 3969, a tree, 15 m. high, 45 cm. in diameter, fruits red. Guarea sp.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 8458, a tree, 15 m. high, 60 cm, in diameter. The speci- 36 CYRUS LONGWORTH LUNDELL mens, in fruit, are not barbate in the leaf axils beneath; the leaves are scarcely to be distinguished otherwise from those of G. excelsa Dek: TRICHILIA CUNEATA Radlk.—Tabasco: Reforma, May 22-26, Matuda 3155, a tree, 3 m. high; Provincia, May 29, Matuda 3225. TRICHILIA HAVANENSIS Jacq.—Tabasco: Reforma, in second growth, May 22-26, Matuda 3229, a tree, 3m. high. TRICHILIA HIRTA L.—Tabasco: Estapilla, riverside, June sEoy Matuda 3485, a tree, 12 m. high, 27-30 em. in diameter. TRICHILIA MONTANA H. B. K.—Tabasco: Boca del Cerro, in advanced forest, July 1-5, Matuda 3570, a tree, 3-4 m. high. TRICHILIA YUCATANENSIS Lundell, Bull. Torrey Bot. Club, 69: 392, 1942. Tabasco: Reforma, May 22-26, Matuda 3154, a tree, 3m. high; Provincia, May 27, Matuda 3219; La Palma, in ad- vanced forest, June 1-6, Matuda 3268, a tree. MALPIGHIACEAE BANISTERIA LAURIFOLIA L.—Tabasco: Achotal, east of Ba- lancan, in second growth, May 9-14, Matuda 3029, 3030, a woody vine, corolla yellow; Retiro, in second growth, June 19-25, Matuda 3460, a woody vine; Kstapilla, at edge of savanna, June 27, Matuda 3507, a woody vine. BUNCHOSIA LANCEOLATA Ture. (?).—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 361 7,asmall tree. BuNcHosIA SWARTZIANA Griseb.—Tabasco: Reforma, May 22— 26, Matuda 3157, a small tree, 3 m. high, 15 em. in diameter. MAvpiGcHia LUNDELLII Morton, Carnegie Inst. Wash. Publ, 461: 138. 1936.—Tahasco: Reforma, May 22-26, Matuda 38181, a small tree, 6-7 m. high, 15 em. in diameter, corolla reddish. MASCAGNIA DIPHOLIPHYLLA (Small) Bullock, Kew Bull, p. 300. 1937. Hiraea dipholiphylla Small, N. Amer. Fl., 25: 122. 1910. Mascagnia polycarpa Brandeg., Univ. Calif. Publ. Bot., 10: 409. 1924.—Chiapas: Javalinero, in advanced forest, July 6-9, Ma- luda 8635, a woody vine. STIGMAPHYLLON LINDENIANUM A. Juss.— Tabasco: Mercedes, in second growth, May 9-14, Matuda 3022, corolla yellow. TETRAPTERIS SELERIANA Ndzu.—Tabasco: Achotal, cast. of FLORA OF EASTERN TABASCO oi Balanedn, in advanced forest, May 9-14, Matuda 3099, a woody vine. VOCHYSIACEAE VoCHYSIA HONDURENSIS Sprague.—Tabasco: Achotal, east of Balancén, in advanced forest, May 9-14, Matuda 3077, a tree, 15 m. high, 40 cm. in diameter. POLYGALACEAE PoLYGALA CONSOBRINA Blake—Tabasco: Estapilla, in sa- vanna, June 27, Matuda 3513, corolla blue. PoLyGaLA HYGROpPHILA H. B. K.—Chiapas: Palenque, in sa- vanna, July 9-14, Matuda 3756. PoLyGALA LoNGrcauLis H. B. K.—Chiapas: Palenque, in sa- vanna, July 9-14, Matuda 3748. PoLyGALA PANICULATA L.—Tabasco: Retiro, riverside, June 19-25, Matuda 3427. SECURIDACA DIVERSIFOLIA (L.) Blake.—Tabasco: Achotal, east of Balancan, in second growth, May 9-14, Matuda 3060. EUPHORBIACEAE ACALYPHA sETosA A. Rich.—Tabasco: Retiro, in second erowth, June 19-25, Matuda 3459. ACALYPHA TABASCENSIS Lundell, Lloydia, 4: 51. 1941.—Ta- basco: Retiro, June 19-25, Matuda 3417 (type collection). ACALYPHA VILLOSA Jacq.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3424. CAPERONIA PALUSTRIS (L.) St. Hil—Tabasco: San Isidro, lake- side, June 7-11, Matuda 3348, flowers white; Boca del Cerro, swampside, July 1-5, Matuda 3542. CROTON FLAVOGLANDULOSUS Lundell, Phytologia, 1: 403. 1940. —Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3042 (type collection), flowers white. CROTON GLABELLUS L.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3420, a tree, 5 m. high, 25 cm. in diameter. Croton perostusus Lundell, Phytologia, 1: 405. 1940.—Ta- basco: Reforma, May 22-26, Matuda 3160, a small tree, 2.5 m. high, 15 em. in diameter; La Palma, in advanced forest, June 38 CYRUS LONGWORTH LUNDELL 1-6, Matuda 3327 (type collection), a tree, 18 m. high, 40 em, in diameter. CROTON REPENS Schlecht.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8747, a perennial herb, Croron TAaBascensts Lundell, Phytologia, 1: 408. 1940.— Tabasco: Reforma, May 22-26, Matuda 3145 (type collection), a tree, 7m. high, 25 cm. in diameter. DALECHAMPIA SCANDENS L.—Tabasco: Achotal, east of Ba- lancdn, in second growth, May 9-14, Matuda 3064, a vine. KUPHORBIA HETEROPHYLLA L.—Tabasco: Kstapilla, in sa- vanna, June 27, Matuda 3500. EUPHORBIA MARGINATA Pursh.—Campeche: Palizada, swamp- side, July 25-28, Matuda 3860 (dct. Wheeler). Garcia NuTANS Rohr.—Tabasco: Boca del Cerro, riverside, July 1-5, Matuda 3568, a tree, 10 m. high, 35 em. in diameter. Hvura PoLyanpra Baill.—Tabasco: Hstapilla, riverside, June 18, Matuda 3480, a tree, 3 m. high, 15 cm. in diameter, flowers white. JULOCROTON ARGENTEUS Didr.—Tabasco: Boca del Cerro, swampside, July 1-5, Matuda 3548. Campeche: Palizada, swamp- side, July 25-28, Matuda 3846. MABEA OCCIDENTALIS Benth.—Tabasco: Achotal, east of Balancin, May 9-14, Matuda 3108, a tree, 4-5 m. high, 15-20 cm. in diameter. PERA BARBELLATA Standl.—Tabasco: Ojo de Agua, southeast of Balancan, in advanced forest, May 9-14, Matuda 3184, a tree, 7m. high, 35 em. in diameter. PHYLLANTHUS ANTILLANUS Var. HYPOMALACUS (Standl.) Lun- dell, Phytologia, 1: 337. 1940. Phyllanthus nobilis var. hypomala- cus Standl., Carnegie Inst. Wash. Publ., 461: 68. 1935.—Tabasco: Achotal, east of Balancdn, in advanced forest, May 9-14, Ma- tuda 8096. Campeche: Vapor, swampside, July 25-28, Matuda 3887, a tree, 6 m. high, 35 em. in diameter. PHYLLANTHUS comprREssus H. B. K—Tabasco: Retiro, river- side, June 19-25, Matuda 3471 (det. Wheeler). PHYLLANTHUS DiIFFUsUS Klotzsch.—Tabasco: Achotal, east of Balancdn, in savanna, May 9- 14, Matuda 3086 (det. Wheeler). FLORA OF EASTERN TABASCO 39 Fig. 1. Sebastiania confusa Lundell. Matuda 3172, half natural size. 40 CYRUS LONGWORTH LUNDELL PHYLLANTHUS LATHYROIDEs H. B. K.—Tabasco: Retiro, river- side, June 19-25, Matuda 3471A (det. Wheeler). PLUKENETIA ANGUSTIFOLIA Standl.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3288, a woody vine, SEBASTIANIA CONFUSA Lundell, Lloydia, 2: 99. 1939 (Fig. 1).-— Tabasco: Reforma, in advanced forest, May 22-26, Matuda 3149, a tree, 7m. high, 15-20 em. in diameter, vernacular name, chitzen; Reforma, Matuda 3172, a small tree, 3 m. high. SEBASTIANIA STANDLEYANA Lundell, Lloydia, 2: 97. 1939. Sebastiania longicuspis Standl., Field Mus. Bot., 11: 184, 1932, nomen confusum.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3476, a tree, 15 m. high, 25 em. in diameter. AQUIFOLIACEAE ILEX PANAMENSIS Standl.—Tabasco: Ojo de Agua, southeast of Balancdn, in advanced forest, May 9-14, Matuda 3126, a tree, 5-6 m. high, 30 em. in diameter. CELASTRACEAE Mayrenus Purpusm Lundell, 4 mer. Midi. Nat., 20: 238. 1938. ~——Tabasco: Reforma, at edge of { orest, May 22-26, Matuda 3147, a shrub. The collection has staminate flowers, and it appears to be referable here. Mayrenus Scurper Lundell, Phytologia, 1: 305. 1939.—Ta- basco: Ojo de Agua, southeast of Balancdn, in advanced forest, May 9-14, Matuda 8140, a tree, 7-10 m. high, 35 em. in diameter; Mstapilla, in second growth, June 18, Matuda 3490, a small tree, 3m. high. Chiapas: Aguacate, in savanna forest, July 16-18, Matuda 3766, a small tree, 2.5 m. high. MyaInpba FILIPEs (Sprague) Loes., Notizbl. Bot. Gart., 13: 226. 1936. Microtropis filipes Sprague, Kew Bull., p. 363. 1909.— Tabasco: Mercedes, in second growth, May 9-14, Matuda 3018, a shrub, 30-50 cm. high; San Isidro, in advanced forest, June 7— 11, Matuda 3875, a small tree, 2 m. high. MYGINDA PUBERULA (Lundell) Lundell, Bull. Torrey Bot. Club, 64: 553. 1937. Rhacoma riparia var. puberula Lundell, Carnegie Inst. Wash. Publ., 478: 213. 1937.— Tabasco: Retiro, riverside on FLORA OF EASTERN TABASCO 41 limestone, June 19-25, Matuda 3429. The species is very close to M. filipes, and additional collections may show that it 1s conspe- cific. RHAcoMA MACROCARPA (Brandeg.) Standl., Contrib. U. S. Nat. Herb., 23: 681. 1923. Myginda macrocarpa Brandeg., Univ. Calif. Publ. Bot., 6:56. 1914:-— Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3689, a tree, 10 m. high, 30 cm. in diameter. Winmertra Barrierri Lundell, Bull. Torrey Bot. Club, 65: 467. 1938.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3287, a tree, 20 m. high, 50 cm. in diameter. HIPPOCRATEACEAE Currvea Mrrcue.iar (Johnston) A. C. Smith, Brittonza, 3: 401. 1940. Hippocratea Mitchellae Johnston, Contrib. Gray Herb., 81: 93. 1928.—Tabasco: Boca del Cerro, July 1-5, Matuda 3571. This fruiting collection was distributed by the writer as a new species, but flowering material is needed to determine definitely whether or not it is distinct. Pristimera ceLastromwes (H. B. K.) A. G. Smith, Brittonza, 3-371. 1940. Hippocratea tabascensis Lundell, Contrib. Univ. Mich. Herb., 4: 16. 1940.—Tabasco: Estapilla, June 18, Matuda 3484 (type collection of H. tabascensis), a woody vine. The recent treatment of the American species of Hippocrateaceae by A. C. Smith (Brittonia, 3: 341-555. 1940) is conservative, and various species which he reduces to synonymy doubtless will be revived when additional material becomes available. Hippocratea voutusiLis L.—Tabasco: Achotal, east of Ba- lancdn, in advanced forest, May 9-14, Matuda 3079, a woody vine. SAPINDACEAE CUPANIA AURICULATA Standl., Field Mus. Bot., 8: 20. 1980.— Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3656, a tree, 5 m. high. MATAYBA RETUSA Lundell, Lloydia, 4: 52. 1941.—Tabasco: Tenosique, in second growth, June 14-16, Matuda 3403 (type collection), a small tree. 42 CYRUS LONGWORTH LUNDELL SERJANIA PTERARTHRA Standl., Carnegie Inst. Wash. Publ., 461: 70. 1935.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3017, a woody vine. TALIsta FLorestt Standl., Trop. Woods, 26: 14. 1931.—Ta- basco: Reforma, May 22-26, Matuda 3193, a tree, 15 m. high, 30 cm. in diameter. THourntpiumM Marupar Lundell, Lloyard, Be 53.194]. Ta- basco: Boca del Cerro, on limestone, July 1-5, Matuda 3590 (type collection), a tree, 7 m. high, 20 em. in diameter. RHAMNACEAE SAGERETIA ELEGANS (H. B. K.) Brongn.—Tabasco: Hstapilla, in second growth, June 18, Matuda 3491, a woody vine. VITACEAE AMPELOCISSUS ERDWENDBERGIT Planch.—Tabasco: Reforma, May 22-26, Matuda 3201, a woody vine. , CISSUS GOSSYPIIFOLIA Standl., Field Mus. Bot., 8: 28. 1930,— Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3314, flowers red. CIsSUS RHOMBIFOLIA Vahl (?).—Chiapas: Javalinero, in ad- vanced forest, June 6-9, Matuda 361 4, flowers red. VITIS CARIBAEA DC.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3306. TILIACEAE CARPODIPTERA AMELIAE Lundell, Field and Lab., 6:18. 1987. —Tabasco: Boca del Cerro, on limestone, July 5, Matuda 3626, a tree, 15 m. high, 35 cm. in diameter, leaves purplish, flowers violet. Muntineta CaLasura L.—Tabasco: Provincia, May 27, Matuda 3228, a tree. MALVACEAE ABUTILON HIRTUM (Lam.) Sweet.—Tabasco: La Palma, river- side, June 1-6, Matuda 3280, flowers yellow, vernacular name, malva, Hisiscus cosrarus A. Rich.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3758. FLORA OF EASTERN TABASCO 43 Histscus Furcentatus Lam.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3048, corolla reddish. Hiztscus sororius L.f.—Tabasco: San Isidro, in swamp, June 7-11, Matuda 3367. Hisiscus sp.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8753A. Hipiscus sp.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8753B. MaLacura capitata L.—Tabasco: Mercedes, lakeside, May 9-14, Matuda 3010 (det. Wiggins), corolla yellow. SipA LINIFOLIA Juss.—Chiapas: Aguacate, July 16-18, Ma- tuda 3777. STERCULIACEAE HeLicreres GUAZUMAEFOLIA H. B. K.—Chiapas: Palenque, edge of savanna, July 9-14, Matuda 3752. Me.Locuta PYRAMIDATA L.—Tabasco: Reforma, May 22-26, Matuda 3189. DILLENIACEAE CURATELLA AMERICANA L.—Tabasco: Ojo de Agua, southeast of Balancdn, in savanna, May 9-14, Matuda 3123, a tree, 7 m. high, 30-35 cm. in diameter, vernacular name, caticon. Chiapas: Aguacate, July 16-18, Matuda 3808, a tree. Davita Marupat Lundell, Phytologia, 1: 371. 1940.—Chi- apas: Javalinero, in forest, July 6-9, Matuda 3647A (type col- lection), a woody vine. TETRACERA MOLLIS Standl., Field Mus. Bol., 8: Atay, UYU Chiapas: Javalinero, in forest, July 6-9, Matuda 3647, a woody vine. ACTINIDIACEAL Q\URAUIA LEUCOCARPA Schlecht.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3458, a tree, 17 m. high, 35 em. in diameter; Boca del Cerro, in advanced forest, July 1-5, Matuda 3555, a tree, 8 m. high, 30 cm. in diameter. Chiapas: ruins of Palenque, in advanced forest, riverside, July 10-12, Matuda 3700, a tree, 10 m. high. The species has been widely confused with S. pauciserrata Hemsl. 44 CYRUS LONGWORTH LUNDELL OCHNACEAE OURATEA NITIDA (Sw.) Engler.—Chiapas: Aguacate, in savanna forest, July 16-18, Matuda 3804, & small tree, corolla yellow. OuraTEA Puck Riley, Kew Bull., p. 109. 1924.—Tabasco: Reforma, in advanced forest, May 22-26, Matuda 31 61, a small tree, 3 m. high; San Isidro, lakeside, June 7-11, Matuda 3355, a shrub, 3 m. high, corolla yellow ; San Isidro, in swamp, Matuda 3371, a small tree. SAUVAGESIA ERECTA L.—Tabasco: Ojo de Agua, southeast of Balancan, in savanna, May 9-14, Matuda 3138, corolla white. MARCGRAVIACEAE SOUROUBEA TRIANDRA Lundell, Phytologia, 1: 244. 1937.— Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3679. THEACEAE TERNSTROEMIA TEPEZzAPOTE Cham. «& Schlecht.—Tabaseo: Achotal, east of Balancdn, in savanna forest, May 9-14, Matuda 3104, a tree, 7-9 m. high, 20-25 cm. in diameter. GUTTIFERAE CiusiA LunpELLu Standl., Carnegie Inst. Wash. Publ., 461: 72. 1935.—Tabasco: Ojo de Agua, southeast of Balancan, in ad- vanced forest, May 9-14, Matuda 3139, a tree, 10 m. high, 25-35 cm. in diameter. HYPERICACEAE VISMIA FERRUGINEA H. B., ix.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3599, a tree, 10 m. high, 30 em. in diameter. VIOLACEAR CORYNOSTYLIS ARBOREA (L.) Blake.—Tabasco: San Isidro, swampside, June 7-11, Matuda 3356, a woody vine, corolla white. Campeche: Vapor, swampside, July 25-28, Matuda 3890, 4 woody vine. HYBANTHUS CALCEOLARIA (LG. Seniilze=—Dabascd: Ojo de Agua, southeast of Balanedn, in savanna, May 9-14, Matuda FLORA OF EASTERN TABASCO 45 3119. Chiapas: Aguacate, July 16-18, Matuda 38787, corolla white. Rinorea Hume Sprague, Kew Bull., p. 307. 1921.—Ta- basco: Achotal, east of Balancén, in advanced forest, May 9-14, Matuda 8101, a tree, 3 m. high, 10 cm. in diameter; San Isidro, in advanced forest, June 7-11, Matuda 3353, 3354. FLACOURTIACEAE CASEARIA ACULEATA Jacq.—Tabasco: Estapilla, in savanna, June 27, Matuda 3518, a small tree. Campeche: Palizada, July 25-28, Matuda 3853; Vapor, in forest, July 25-28, Matuda 3914, a tree, 5 m. high, flowers white. CaseartA Barrietri Lundell, Lloydia, 2: 104. 1939.— Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3470. CASEARIA gAvITENSIS H. B. K.—Tabasco: Achotal, east of Balanedén, in advanced forest, May 9-14, Matuda 3100, a small tree. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3696, a tree, 3-4 m. high, fruits red. CASEARIA NITIDA (L.) Jacq.—Tabasco: Mercedes, riverside, May 9-14, Matuda 3011, a tree, 3 m. high, 10 em. in diameter; Achotal, east of Balancdn, in second erowth, May 9-14, Matuda 3038, a shrub, 3-4 m. high, 10-15 cm. in diameter; Reforma, in second growth, May 22-26, Matuda 31 78, a shrub; San Isidro, in advanced forest, June 7-11, Matuda 3341, a tree, 3 m. high, 15 em. in diameter; Retiro, in second growth, June 19-25, Ma- tuda 3433, 3461, a tree. Hasseuria prorca (Benth.) Sleumer (?).—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3003, a tree, 7 m. high; Reforma, May 22-26, Matuda 3156, a small tree, 3 m. high. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3597, a tree, 8m. high, 25 cm. in diameter. LartiA THamnra L.—Tabasco: San Isidro, June 7-11, Ma- tuda 3346. Chiapas: Sierra de Palenque, in advanced forest, July 14, Matuda 3715, a tree, 10 m. high, 40 cm. in diameter. ZUELANIA Guriponra (Sw.) Britt. & Millsp.—Tabasco: Ojo de Agua, southeast of Balancan, in second growth, May 9-14, Matuda 3118, a tree, 3-4 m. high, 15-20 cm. in diameter; Reforma, May 22-26, Matuda 3233, a tree, 8 m. high, 20 em. in diameter. 46 CYRUS LONGWORTH LUNDELL Pia. 2. Erblichia Standleyi Steyermark. Matuda 3319, half natural size. FLORA OF EASTERN TABASCO 47 TURNERACEAE ERBLICHIA StaNDLEYI Steyermark, Field Mus. Bol., 22: 306. 1940 (Fig. 2)—Tabasco: La Palma, in advanced forest, June 1— 6, Matuda 3319, a tree, 25 m. high, 70 cm. in diameter, vernacular name, San Pedro. PASSIFLORACEAE PASSIFLORA FOETIDA var. crbtaTa (Dryand.) Mast.—Tabasco: La Palma, June 1-6, Matuda 3247 (det. Killip). Passtrtora Rovrrosar Killip.—Tabasco: La Palma, in second erowth, June 1-6, Matuda 3283 (det. Killip). PASSIFLORA SERRATIFOLIA L.—Tabasco: Reforma, May 22- 26, Matuda 3207 ; Retiro, in second growth, riverside, June 19-25, Matuda 3434; Boca del Cerro, in second growth, July 1-5, Ma- tuda 3583. CACTACEAE Dram1a Testupo (Karw.) B. &. R,—Campeche: Vapor, July 25-28, Matuda 3897 (det. Clover), flowers white. EpreHyLLuM CRENATUM (Lindl.) G. Don.—Tabasco: La Palma, riverside, June 1-6, Matuda 3310, flowers white. Rurpsauis CassutHa Gaertn.—Tabasco: San Isidro, lakeside epiphyte, June 7-11, M atuda 3384. RuIPsaLis cORIACEA Polak.—Tabasco: San Isidro, on tree, June 7-11, Matuda 3369. LYTHRACEAE (Determined by Robert C. Foster, Cambridge) CUPHEA CARTHAGENENSIS (Jacq.) Macbr.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3749, corolla blue. CuUPHEA DECANDRA var. PURPUSII (Brandeg.) Bacig.—Tabasco: Retiro, riverside, June 19-25, M atuda 3426, corolla blue. LytTuruM GRACILE Benth.—Tabasco: Reforma, riverside, May 22-26, Matuda 3170, corolla purple. RHIZOPHORACEAE Ruizopuora Maneue L.—Tabasco: La Palma, riverside, June 1-6, Matuda 3329. 48 CYRUS LONGWORTH LUNDELL —— 2 $55: eee balancanensis Lundell. Matuda 3361 (type collection), h ah natur al size FLORA OF EASTERN TABASCO 49 COMBRETACEAE Buciwa Buceras L.—Tabasco: Ojo de Agua, southeast of Balancdn, in advanced forest, May 9-14, Matuda 3121, a tree, 12 m. high, 45 cm. in diameter; Reforma, May 22-26, Matuda 3166, a tree, 15 m. high, 60 cm. in diameter. CoMBRETUM MEXICANUM Humb. & Bonpl.—Tabasco: Mer- cedes, riverside, May 9-14, Matuda 3024, a tree, 7-10 m. high; Achotal, east of Balancan, in savanna, May 9-14, Matuda 3082, a woody vine; Boca del Cerro, edge of swamp, July 1-5, Matuda 3582, a woody vine, 25 cm. in diameter; Boca del Cerro, riverside, Matuda 3537, a woody vine. As currently interpreted, it is ob- vious that this species is a catch-all for various related forms of the complex. MYRTACEAE CanyprrantHes Karuinei Standl., Pield Mus. Bot., 8: 29. 1930.—Tabasco: Boca del Cerro, on limestone at riverside, July 1-5, Matuda 3564, a small tree. Chiapas: Sierra de Palenque, in advanced forest, riverside, July 14, Matuda 3717, a tree, 5 m. high, 20 cm. in diameter. EUGENIA BALANCANENSIS Lundell, Phytologia, 1: 481. 1941 (Fig. 3).—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3361 (type collection), a tree, 4 m. high, 15 em. in di- ameter. EvGEnIA BRACTEOLOSA Lundell, Contrib. Univ. Mich. Herb., 7: 31. 1942.—Tabasco: Reforma, May 22-26, Matuda 8194, a tree, 13 m. high, 20 cm. in diameter. EvGENIA BUMELIOIDES Standl., Carnegie Inst. Wash. Publ., 461: 75. 1935.—Tabasco: Boca del Cerro, on limestone at river- side, July 1-5, Matuda 3568, a small tree, 6-7 m. high. EuGENIA CAMPECHIANA Lundell, Contrib. Univ. Mich. Herb., 7: 39. 1942.—Campceche: Palizada, lakeside, July 25-28, Matuda 3832 (type collection), a small tree, 3-4 m. high, corolla white; Palizada, Matuda 3847, fruits edible, vernacular name, boroconte. Evucenia Caputt (Schl. & Cham.) Berg.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3018, a shrub. Eucenta LInpENIANA Berg.—Tabasco: Boca del Cerro, on limestone, July 1-5, Matuda 3592, a shrub, 2m. high. CYRUS LONGWORTH LUNDELL "la. 4. Hugenia tabascensis Lundell. half natural size, Matuda 3561 (type collection), FLORA OF EASTERN TABASCO 51 Eugenia TaBascensis Lundell, Phylologia, 1: 482. 1941. (Hig. 4).—Tabasco: Boca del Cerro, on limestone, July 1-5, Matuda 3561 (type collection), a small tree. SUGENIA sp.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3621, a tree, 10 m. high, 35 cm. in diameter. EvuGENIA sp.—Tabasco: San Isidro, in forest, June 7-11, Ma- tuda 3342, a tree, 4m. high, 13 cm. in diameter. Eucenra sp.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3241, a shrub. Myrcia RUFIDULA Schlecht.—Tabasco: Boca del Cerro, on limestone, July 1-5, Matuda 3557, a tree, 4 m. high, 15 cm. in diameter. Chiapas: Palenque, riverside, July 10-12, Matuda 3672, a tree, 4m. high; Aguacate, in savanna forest, July 16-18, Matuda 3811, a small tree. Pimenta OFFICINALIS Lindl.—Tabasco: Estapilla, June 27, Matuda 3496, a tree, 15 m. high, 35 em. in diameter, vernacular name, pimiento de tierra, MELASTOMACEAE (Determined by H. A. Gleason, New York) Acroris rosreLuata (Naud.) Triana.—Chiapas: Javalinero, edge of forest, July 6-9, Matuda 3609, corolla white. ACISANTHERA QUADRATA Pers.—Tabasco: Ojo de Agua, south- east of Balancdn, in savanna, May 9-14, Matuda 31388, 30 cm. high, corolla reddish. Bre.Lucta GRossuLARIOIDES (L.) Triana.—Chiapas: ruins of Palenque, July 10-12, Matuda 3772. LIDEMIA DENTATA D. Don.—Tabasco: Boca del Cerro, in ad- vanced forest, July 1-5, Matuda 3567, a shrub, corolla reddish. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3654, a shrub, 1-1.5 m. high, corolla reddish; ruins of Palenque, in ad- vanced forest, July 10-12, Matuda 3692, a shrub, corolla reddish. CLIDEMIA DEPENDENS D. Don.—Tabasco: Ojo de Agua, south- east of Balancdn, in advanced forest, May 9-14, Matuda 3130, a shrub, 2 m. high. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 36638, a shrub, corolla white. 52 CYRUS LONGWORTH LUNDELL CLIDEMIA Duppwana Steud.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3296, a small shrub, 50-90 em. high, corolla reddish; Boca del Cerro, July 1-5, Matuda 3581, a small tree, corolla reddish. CLIDEMIA RUBRA (Aubl.) Mart.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3049, a shrub, 2 m. high. Chiapas: Palenque, in savanna forest, July 9-14, Matuda 3725, a shrub, 1 m. high. CONOSTEGIA XALAPENSIS (Bonpl.) D. Don.—Tabasceo: Acho- tal, east of Balancdn, in savanna, May 9-14, Matuda 3062, a shrub, 1.5 m. high. CONOSTEGIA sp.—Tabasco: Achotal, east of Balanean, in ad- vanced forest, May 9-14, Matuda 3068, a small tree, 1.5 m. high, corolla white. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3601, a shrub, corolla white. HENRIETTEA SUCCOSA (Aubl.) DC.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3657, a tree, 10-12 m. high, 30 cm. in diameter, H&TEROTRICHUM OCTONUM (Bonpl.) DC.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3655, a shrub, 2 m. high, corolla white. LEANDRA MEXICANA (Naud.) Cogn.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3600, a shrub, corolla reddish. Miconia auBicans (Sw.) Triana.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3083, a shrub, 1.5 m, high. Chiapas: Aguacate, in savanna, July 16-18, Matuda 8782, a shrub. MICONIA ARGENTEA (Sw.) DC.—Tabasco: Achotal, east. of Balanedn, in advanced forest, May 9-14, Matuda 3080, a tree, 8m. high, 25 em. in diameter. MICONIA DODECANDRA (Desr.) Cogn.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 8613, a tree, 6 m. high, 18 cm. in’diameter. MICONIA IBAGUENSIS (Bonpl.) Triana.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3602, a shrub; ruins of Palenque, in advanced forest, July 10-12, Matuda 3687, a shrub, corolla white. FLORA OF EASTERN TABASCO 53 Miconra LAcERA (Bonpl.) Naud.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3624, a shrub, 2 m. high, corolla reddish. MiconIA PTEROPODA Benth.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3618, a tree, 8 m. high, 19 em. in diameter. Miconta puncrata (Desr.) D. Don.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3605, a small tree, 6-7 m. high, 15 em. in diameter. Miconra SCHLECHTENDALIL Cogn.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3376, a small tree, corolla white. Campeche: Vapor, July 25-28, Matuda 3907, a shrub. Miconta scorproipEs (Schl. & Cham.) Naud.—Tabasco: Ojo de Agua, southeast of Balancdn, lakeside, May 9-14, Matuda 3141, a shrub, 2 m. high. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3598, a small tree, 8 m. high, 15-25 cm. in diameter; Javalinero, Matuda 3638, a tree, 10 m. high, 15-20 em. in diameter. MovuriritA GLEASONIANA Standl., Field Mus. Bot., 22: 361. 1940.—Tabasco: Achotal, east of Balancdn, in advanced forest, May 9-14, Matuda 3098 (type collection), a tree, 4-5 m. high, 10-15 em. in diameter; San Isidro, June 7-11, Matuda 3339. The two collections were referred to M. Muelleri Cogn. by the writer, and distributed under that name. M. Muelleri is very closely re- lated if not conspecific. Mourtiria PARvVIFOLIA Benth.—Tabasco: Achotal, east of Balanedn, in advanced forest, May 9-14, Matuda 3046, a shrub, 3 m. high, 10-15 em. in diameter, corolla yellowish; Reforma, May 22-26, Matuda 8208. Campeche: Vapor, July 25-28, Matuda 3891, a shrub, corolla white. ONAGRACEAE Juss1apaA NatTANs H. B. K.—Tobasco: Boca del Cerro, in swamp, July 1-5, Matuda 3588, corolla white. JussIAEA SUFFRUTICOSA L.—Tabasco: Mercedes, lakeside, May 9-14, Matuda 3008. ot CYRUS LONGWORTH LUNDELI ARALIACEAE GILIBERTIA ARBOREA (L.) March.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3474, a tree, 15 m. high, 45 em. in diameter; La Palma, in advanced forest, June 1-6, Ma- fuda 3239, a shrub. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3636, a small tree; Sierra de Palenque, in ad- vanced forest, July 14, Matuda 3719, a small tree. GILipertTIA Marupar Lundell, Phytologia, 1: 372. 1940.— Tabasco: Reforma, in advanced forest, May 22-26, Matuda 3152 (type collection), a small tree, 3 m. high. OREOPANAX GUATEMALENSE (Lem.) Dec. & Pl.—Tabasco: Boca del Cerro, July 1-5, Matuda 3551. UMBELLIFERAE IRYNGIUM NASTURTIIFOLIUM Juss.—Tabasco: Mercedes, lake- side, May 9-14, Matuda 3014 (det. Constance). HYDROCOTYLE UMBELLATA L.—Campeche: Palizada, swamp- side, July 25-28, Matuda 8854. MYRSINACEAE ArpisiA Matupar Lundell, Lloydia, 4: 55. 1941.—Tabaseco: La Palma, in advanced forest, June 1-6, Matuda 3295 (type col- lection), a small tree, flowers white ; San Isidro, in advanced forest, June 7-11, Matuda 3364, a small tree, flowers white. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 8648, a small tree, 4 m. high, flowers white. PARATHESIS AERUGINOSA Standl.—Tabasco: J uarez, near Tenosique, June 14-16, Matuda 3391, a treelet, 3m. high, flowers red. PARATHESIS CUBANA (A. DC.) Molinet & Maza. Parathesis obovaia Standl., Field Mus. Bot., 4: 250. 1929.—Tabasco: La Palma, June 1-6, Matuda 3265, a shrub. STYLOGYNE LAEVIS (Oerst.) Mez.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3622, a tree, 3m. high, 30 em. in diameter, FLORA OF EASTERN TABASCO 55 THEOPHRASTACEAE JACQUINIA AURANTIACA Ait.—Tabasco: San Isidro, swampside, June 7-11, Matuda 3377, a tree, 15 m. high, 40 cm. in diameter. Campeche: Vapor, swampside, July 25-28, Matuda 3913, a tree, 7-10 m. high, 20-30 cm. in diameter. SAPOTACEAE Acuras Zapora L.—Tabasco: La Palma, June 1-6, Matuda 3254, a tree. BUMELIA MAYANA Standl., Trop. Woods, 31: 41. 1932.—Tabas- co: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3034, a shrub, 2 m. high. BuMELIA TABASCENSIS Lundell, Contrib. Uni. Mich. Herb., 4: 92. 1940.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3455 (type collection), a tree, 18 m. high, 35 em. in diameter. Lucuma CAMPECHIANA H. B. K.—Tabasco: Achotal, east of Balanecdn, in advanced forest, May 9-14, Matuda 3071, a tree, 10 m. high, 30 em. in diameter. Lucuma Durianpu Standl.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3297, a small tree. LucUMA LAETEVIRIDIS Pittier.—Tabasco: San Isidro, in ad- vanced forest, June 7-11, Matuda 3351, a tree, 7-10 m. high, 45 em. in diameter. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3713, a tree, 10 m. high, 45 em. in diameter. MAMMEA AMERICANA L.—Tabasco: Boca del Cerro, July 1-5, Matuda 3580, a tree, 15-20 m. high, 55-70 cm. in diameter, vernacular name, mamey. SIpEROXYLON LUNDELLU Standl., Carnegie Inst. Wash. Publ., 461: 79. 1935.—Tabasco: Achotal, east of Balancdn, in advanced forest, May 9-14, Matuda 3073, a tree, 8-10 m. high, 30-40 cm. in diameter. EBENACEAE Diospyros BUMELIOIDES Standl., Trop. Woods, 18: 31. 1929. —Campeche: Vapor, swampside, July 25-28, Matuda 3889, a shrub. 56 CYRUS LONGWORTH LUNDELL Diospyros campechiana Lundell. Matuda 3843 (type collection), Fia. 5. half natural size. FLORA OF EASTERN TABASCO 57 Diospyros CAMPECHIANA Lundell, Contrib. Univ. Mich. Herb., 4: 23. 1940 (Fig. 5)—Campeche: Palizada, swampside, July 25— 28, Matuda 3843 (type collection), a tree, 7-10 m. high, 40-45 cm. in diameter. Drosprros EBenaster Retz.—Tabasco: Boca del Cerro, river- side, July 1-5, Matuda 3540, a tree, 15 m. high, 45 em. in di- ameter, vernacular name, faucht. Drospyrros YUCATANENSIS Lundell, Carnegie Inst. Wash. Publ, 478: 217. 1937.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3378, a tree, 15 m. high, 35 cm. in diameter; Tenosique, in advanced forest, June 14-16, Matuda 3401, a tree, 5 m. high. OLEACEAE LINOCIERA OBLANCEOLATA Rob.—Tabasco: La Palma, in ad- vanced forest, June 1-6, Matuda 3284, a small tree, 4 m. high, 15 em. in diameter. LOGANIACEAE PoLYPREMUM PROCUMBENS L.—Tabasco: Achotal, east of Ba- lancdn, in savanna, May 9-14, Matuda 8059, 3108. SpriGeLIA HUMBOLDTIANA Ch: im. & Schlecht.—Tabasco: Re- tiro, in advanced forest, June 19 -25, Matuda 3438. QrRYCHNOS BRACHISTANTHA Standl., Field Mus. Bot., 12: 412. 1936.—Tabasco: San Isidro, in adv: anced forest, ane 7-11, Matuda 3374, a woody vine, 30 m. high. . SrRYCHNOS TABASCANA Sprague & Sandw., Kew Bull., p. 128. 1927. Strychnos panamensis var. hirtiflora Sande Field Mus. Bot., 11: 138. 1932. Strychnos hirtiflora (Standl.) ewdell Bull. norrey Bot. Club. 64: 556. 1937.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3285, a woody vine, corolla yellowish. GENTIANACEAE Covrousra spicata Aubl.—Chiapas: Palenque, in savanna, July 9-14, Matuda 37464. Eusroma ExALTAtum (L.) Griseb.—Tabasco: Reforma, river- side, May 22-26, Matuda 3177, an annual, corolla blue. 58 CYRUS LONGWORTH LUNDELL MENYANTHACEAE LIMNANTHEMUM HuMBOLDTIANUM (H. B. Kk.) Griseb.—Cam- peche: Palizada, in lake, July 25-28, Matuda 3827, APOCYNACEAE MANDEVILLA suBSAGITTATA (R. & P.) Woodson.—Tabasco: Achotal, east of Balancan, in second growth, May 9-14, Matuda 3055, a vine, corolla yellow. Chiapas: Javalinero, July 6-9, Ma- luda 8651, a vine. PLuMEnriopsis Anouat (L.) Rusby & Woodson, Ann. Missouri Bot. Gard., 24: 11. 1937. Thevetia Ahouat (Dy) As DC DG. Progr, 8: 344. 1844. Thevetia nitida (H. B. K.) A. DC., DC. Prodr., 8: 345. 1844.—Tabasco: Achotal, east of Balancan, in second growth, May 9-14, Matuda 3078, a shrub. Although the plant may be native to the Yucatan Peninsula, it has been observed and col- lected only in clearings and second growth. PRESTONIA GUATEMALENSIS Woods., Ann. Missouri Bot. Gard., 23: 339. 1936.—Tabasco: Tenosique, in second growth, June 14— 16, Matuda 3405. PRESTONIA MEXICANA A. DC.—Tabasco: Boca del Cerro, in second growth, July 1-5, Matuda 3582 (det. Woodson). Chiapas: Aguacate, July 16-18, Matuda 3808, a vine. TABERNAEMONTANA CHRYSOCARPA Blake.—Tabaseo: Ojo de Agua, southeast. of Balancdén, in second growth, May 9-14, Matuda 3124, a tree, 3 m. high; Estapilla, edge of savanna, June 27, Matuda 3522, a small tree, corolla yellow. Urecuires Anpripuxu Muell. Arg.—Chiapas: ruins of Pa- lenque, edge of advanced forest, July 10-12, Matuda 3677, a vine. ASCLEPIADACEAE ASCLEPIAS LONGICORNU Benth.—Tabasco: Mstapilla, in sa- vanna, June 27, Matuda 3510 (det. Woodson). ASCLEPIAS sETOSA Benth.—Tabasco: Estapilla, in savanna, June 27, Matuda 3510A (det. Woodson). FIscHERIA BRIQUETIANA Standl., Field Mus. Bot., 11: 139. 1932.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3482, FLORA OF EASTERN TABASCO 59 GonoLosus crenropHorts (Blake) Woodson, Ann. Miss. Bot. Gard., 28: 248. 1941. Vincetoxicum cleniophorum Blake, Contrib. Gray Herb., 52: 84. 1917. Vincetoxicum Lundellit Standl., Pzeld Mus. Bot., 8: 148. 1930.—Campeche: Vapor, July 25-28, Matuda 3908 (det. Woodson), a vine. METASTELMA BARBIGERUM Scheele.—Chiapas: Aguacate, July 16-18, Matuda 3765 (det. Woodson). CONVOLVULACEAE , ANISEIA MARTINICENSIS (Jacq.) Choisy. —Campeche: Palizada, July 25-28, Matuda 3857 (det. Martin), flowers white. Evo.vuuus Fitirers Mart.—Chiapas: Aguacate, July 16-18, Matuda 3790 (det. Martin). EVvoLVULUS SERICEUS Swartz.— Tabasco: Estapilla, in savanna, June 27, Matuda 3504, corolla white. IPOMOEA APHYLLA Standl., Field Mus. Bot., 11: 139. 1933.— Chiapas: Palenque, in savanna, July 9-14, Matuda 3739, corolla white. IpoMOEA CALLIDA House (?),—Campeche: Palizada, July 25—- 28, Matuda 3831 (det. Martin), a vine. Ipomoga cissorpEs (Lam.) Griseb.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3061 (det. Martin), corolla white. IPOMORA CRASSICAULIS (Benth.) Robinson.—Campeche: Pali- zada, edge of swamp, July 25-28, Matuda 3835. IpoMoEA RUBRA (Vahl) Millsp.—Tabasco: La Palma, riverside, June 1-6, Matuda 3252 (det. Martin), a vine, corolla red. IpoMOEA SAGITTaATA Cav.—Tabasco: La Palma, riverside, June 1-6, Matuda 3253 (det. Martin), a vine, corolla red. IpomoEA squamosa Choisy.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3733A (det. Martin). JACQUEMONTIA AZUREA (Desr.) Choisy.—Chiapas: Aguacate, July 16-18, Matuda 3763, a vine, corolla blue. JACQUEMONTIA TAMNIFOLIA (L.) Griseb.—Chiapas: Aguacate, swampside, July 16-18, Matuda 3813, corolla blue. OPERCULINA ORNITHOPODA (Rob.) House.—Tabasco: Estapilla, in savanna, June 27, Matuda 3508, a vine, corolla white. 60 CYRUS LONGWORTH LUNDELL BORAGINACEAE BOURRERIA OXYPHYLLA Standl., Trop. Woods, 16: 40. 1928.— Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3682, a tree, 8 m. high, corolla white. CorDIA ALBA (Jacq.) R. & S.—Tabasco: Mercedes, riverside, May 9-14, Matuda 3020, a tree, 8-10 m. high, 30 em. in diameter, corolla yellow. CORDIA DIVERSIFOLIA Pavon.—Tabasco: Boca del Cerro, July 1-5, Matuda 3584. CorpIA GLABRA L.—Tabasco: Reforma, May 22-26, Matuda 8216. CoRDIA STENOCLADA Johnston, Journ. Arn. Arb., 21: 353. 1940. Chiapas: Javalinero, in second growth, July 6-9, Matuda 3658, a tree, 7-8 m. high, 35 em. in diameter; ruins of Palenque, in advanced forest, July 10-12, Matuda 3695, a tree, 10 m. high, 40 cm. in diameter, corolla white. The two collections agree closely with the description of this recently proposed species. HELIOTROPIUM ANGIOSPERMUM Murr.—Tabasco: Reforma, May 22-26, Matuda 3226. HELIOTROPIUM FILIFORME Lehm.—Tabasco: Achotal, east of Balancén, in savanna, May 9-14, Matuda 3050, corolla white. TOURNEFORTIA GLABRA L.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3412, corolla white. TOURNEFORTIA HIRSUTISSIMA L.—Tabasco: Mercedes, in sec- ond growth, May 9-14, Matuda 3016. VERBENACEAE AEGIPHILA ELATA Sw.—Tabasco: Achotal, east of Balancan, in advanced forest, May 9-14, Matuda 3031, a woody vine; Achotal, in savanna, Matuda 3081, a woody vine; Tenosique, in second growth, June 14-16, Matuda 3406, a woody vine. CALLICARPA ACUMINATA H. B. KX.—Chiapas: Javalinero, in second growth, July 6-9, Matuda 3641, a shrub, 2.5 m. high. CITHAREXYLUM CAUDATUM L.—Tabaseo: Achotal, cast of Balancdn, in advanced forest, May 9-14, Matuda 3066, a tree, 4-5 m. high. Chiapas: Aguacate, in savanna forest, July 16-18, Matuda 3771, 8776, a treelet, 2.5 m. high. FLORA OF EASTERN TABASCO 61 CIPHAREXYLUM HEXANGULARE Greenm.—Tabasco: Boca del Cerro, on limestone, riverside, July 1-5, Matuda 3562, a shrub. Campeche: Palizada, July 25-28, Matuda 3861, a shrub, corolla white. CLERODENDRUM FRAGRANS var. PLENIFLORUM Schau.—Ta- basco: La Palma, in second growth, June 1-6, Matuda 3282. CLERODENDRUM LIGUSTRINUM (Jacq.) R. Br.—Tabasco: Re- forma, riverside, May 22-26, Matuda 3168, 3169, a woody vine, corolla white. CoRNUTIA LATIFOLIA (H. B. K.) Moldenke.—Tabasco: Teno- sique, in second growth, June 14—16, Matuda 3398, a shrub, co- rolla purplish. Chiapas: Aguacate, in second growth, July 16-18, Matuda 3807, a shrub, corolla purplish. DuRANTA REPENS L.—Tabasco: Tenosique, in second growth, June 14-16, Matuda 3395. Lrppra HYPOLEIA Briq.—Chiapas: Javalinero, in second growth, July 6-9, Matuda 3630 (det. Moldenke). LIPPIA MYRIOCEPHALA Schl. & Cham.—Chiapas: Javalinero, in second growth, July 6-9, Matuda 3642 (det. Moldenke). PHYLA SCABERRIMA (A. L. Juss.) Moldenke.—Tabasco: Pro- vincia, May 29, Matuda 3224 (det. Moldenke). PHYLA STOECHADIFOLIA (L.) Small.—Tabasco: La Palma, June 1-6, Matuda 3273 (det. Moldenke). QrACHYTARPHETA TABASCANA Moldenke, Phytologia, 1: 4387. 1940 (Fig. 6).—Tabasco: Reforma, May 22-26, Matuda 3218 (type collection). Virex Gaumerit Greenm.—Tabasco: Ojo de Agua, southeast of Balancdn, in forest, May 9-14, Matuda 3114, a tree, 7-10 m. high, 45 em. in diameter; Reforma, in advanced forest, May 22- 26, Matuda 3231, a tree, 14 m. high, 55 cm. in diameter, corolla blue. Campeche: Vapor, in forest, July 25-28, Matuda 3905, a tree, 10-15 m. high, 40 cm. in diameter, corolla blue. LABIATAE Hyptis verticinuata Jacq.—Tabasco: Reforma, May 22-26, Matuda 3227. Sarvra coccinea L.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3272, corolla red. 62 CYRUS LONGWORTH LUNDELL y Pia. 6. Slachytarpheta tabascana Moldenke. Matuda 3218 (type collec- tion), half natural size. FLORA OF EASTERN TABASCO 63 SCUTELLARIA ORICHALCEA Donn. Sm.—Chiapas: ruins of Palenque, in advanced forest, riverside, July 10-12, Matuda 368!. Trucrium vesicarium Mill.—Tabasco: Mercedes, lakeside, May 9-14, Matuda 3006, corolla blue; La Palma, in second growth, June 1-6, Matuda 3257; Mstapilla, in savanna, June 27, Matuda 3512, corolla blue. SOLANACEAE ArHENAEA Nesonr Fernald.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3447 (det. Morton), a shrub. JUANULLOA MEXICANA (Schlecht.) Miers.—Chiapas: Aguacate, in savanna forest, July 16-18, Matuda 3822, on tree. PHYSALIS ANGULATA L.—Campeche: Palizada, swampside, July 25-28, Matuda 3867 (det. Morton). Puysauis sp.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3312. SoLANUM GUANICENSE Urban.—Campeche: Palizada, July 25- 28, Matuda 3848 (det. Morton). _SOLANUM HETEROCLITUM Sendtn.—Tabasco: Tenosique, in ad- vanced forest, June 14-16, Matuda 3449 (det. Morton). Sotanum Hovusronn Dunal.—Campeche: Vapor, swampside, July 25-28, Matuda 3883 (det. Morton), corolla yellow. SOLANUM JAMAICENSE Mill.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3633 (det. Morton), corolla white. SoLANUM NupuM H. B. K.—Tabasco: Mercedes, in second erowth, May 9-14, Matuda 3005; Tenosique, in second growth, June 14-16, Matuda 3396 ; Retiro, in advanced forest, June 19-25; Matuda 8435; Boca del Cerro, in second growth, July 1-5, Matuda 3579. Sonanum Purpusm Brandeg.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3649, (det. Morton), a vine, corolla yellow. Sotanum RovirosaNnum Donn. Sm.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3359 (det. Morton), a shrub, corolla white. SCROPHULARIACEAE ANGELONIA CILIARIS Robinson.—Tabasco: Achotal, east of Balanedn, in savanna, May 9-14, Matuda 3039, corolla blue. 64 CYRUS LONGWORTH LUNDELL Bacopa SaALzMaNnnul (Benth.) Chod. & Hassl.—Chiapas: Agua- cate, in savanna, July 16-18, Matuda 3764 (det. Morton), flowers yellow. BUCHNERA LITHOSPERMIFOLIA H. B. K.—Tabasco: Kstapil in savanna, June 27, Matuda 3514, corolla violet. BUCHNERA PUSILLA H. B K.—Tabasco: Achotal, east of Ba- lancdn, in savanna, May 9-14, Matuda 3143A, corolla white. CAPRARIA BIFLORA L.—Tabasco: La Palma, June 1-6, Matuda ye: KscOBEDIA LABVIS Cham. & Schlecht.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3044, corolla white. GERARDIA Harperi (Penn.) Penn.—Tabasco: Achotal, east of Balancdn, in savanna, May 9-14, Matuda 3040. USSELIA TABASCENSIS Lundell, Contrib. Univ. Mich. Herb., 6: 59. 1941.—Tabasco: Estapilla, in savanna, June 27, Matuda 3524 (type collection), corolla red. COPARIA DULCIS L.—Tabasco; Achotal, cast of Balanedn, in savanna, May 9-14, Matuda 3053. STEMODIA DURANTIFOLIA (L.) Sw.—Tabaseco: E stapilla, in savanna, June 27, Matuda 3528 (det. Morton). UROSKINNERA FLAVIDA Lundell, Harvard Bot. Mus. Leaflets, 9: 68. 1941.—Tabasco: Retiro, on limestone, June 19-25, Matuda 3425 (type collection), corolla yellowish. — a, BIGNONIACEAE ARRABIDAEA INAEQUALIS (DC. ex Splitg.) K. Schum.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3668 (det. Seibert), a woody vine, corolla reddish. CLYTOSTOMA OCOSITENSE (Donn. Sm.) Seibert, Car negre Inst. Wash. Publ., 522: 413. 1940. Adenocalymna (?) ocosttense Donn. Sm., Bot. Ga 18: 209. 1893. Clytostoma elegans Standl., Carnegie ae Wash. Pan 461: 86. 1935.—Tabasco: Retiro, in second growth, riverside, June 19-25, Matuda 3421 (det. Seibert). Campeche: Vapor, July 25-28, Matuda 3881 (det. Seibert), a woody vine. Cypista potosina (K. Schum. & Loes.) Loes. Clytostoma mayanum Standl., Carnegie Inst. Wash. Publ., 461: 86. 1935.— FLORA OF EASTERN TABASCO 65 Tabasco: La Palma, June 1-6, Matuda 3278 (det. Seibert), a woody vine; Estapilla, edge of savanna, June 27, Matuda 3528 (det. Seibert), a woody vine, corolla red. ENaLLaGMa LATIFoLIA (Mill.) Small—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3388 (det. Seibert), a small tree, 4 m. high, 20 cm. in diameter. PARMENTIERA PARVIFLORA Lundell, Lloydza, 3: 211. 1940.— Tabasco: Reforma, riverside, May 22-26, Matuda 3209, 3210 (type collection), a shrub. TYNANTHUS GUATEMALENSIS Donn. Sm.—Tabasco: La Palma, June 1-6, Matuda 3277 (det. Seibert), a woody vine, 20 m. high, corolla yellowish. GESNERIACEAE (Determined by C. V. Morton, Washington) CopONANTHE CRASSIFOLIA (Focke) Morton.—Tabasco: San Isidro, on tree, July 7-11, Matuda 3381, corolla reddish. CoLUMNEA SULPHUREA Donn. Sm.—Chiapas: Sierra de Pa- lenque, in advanced forest, on tree, July 14, Matuda 3714, corolla yellow. DRYMONIA CHIAPENSIS Brandeg. (?).—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3440, a vine, corolla reddish. LENTIBULARIACEAE UrrRICULARIA FOLIOSA L.—Campeche: Vapor, in lake, July 25- 28, Matuda 3878, flowers yellow. UrricuLaRIA oBtTusaA Sw.—Tabasco: La Palma, in lagoon, June 1-6, Matuda 3235, flowers yellow. Campeche: Vapor, in lake, July 25-28, Matuda 3875, flowers yellow. UrrIicuLARIA PUSILLA Vahl.—Chiapas: Agacuate, in savanna, July 16-18, Matuda 3779, flowers yellow. ACANTHACEAE DIANTHERA ANAGALLIS Hemsl.—Campeche: Vapor, swamp- side, July 25-28, Matuda 3888 (det. Leonard), corolla reddish. Justicia PECTORALIS Jacq.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3334. CYRUS LONGWORTH LUNDELL & Standl. Matuda 3670 (type collection), half natural size, Fia. 7. Blepharidium mexicanum FLORA OF EASTERN TABASCO 67 ODONTONEMA CALLISTACHYUM (Schl. & Cham.) Kuntze.—Ta- basco: Boca del Cerro, in advanced forest, July 1-5, Matuda 3566. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3694. RUELLIA NUDIFLORA var. yucATANA Leonard.—Tabasco: Re- tiro, riverside, June 19-25, Matuda 3469. | RUELLIA STEMONACANTHOIDES (Oerst.) Hemsl.—Tabasco: Re- tiro, in advanced forest, June 19-25, Matuda 3472. PLANTAGINACEAE PLANTAGO MAgor L.—Tabasco: La Palma, riverside, June 1-6, Matuda 3320. RUBIACEAE ALIBERTIA EDULIS (L. Rich.) A. Rich.—Tabasco: La Palma, June 1-6, Matuda 3298, a treelet; San Isidro, in advanced forest, June 7-11, Matuda 3362, a small tree, 3 m. high, corolla white. Campeche: Vapor, July 25-28, Matuda 3904, a shrub. ALSEIS YUCATANENSIS Standl., Field Mus. Bot., 8: 50. 1930.— Tabasco: La Palma, in advanced forest, June 1-6, Matuda 3293, 3333, a tree, 20 m. high, 40 cm. in diameter, vernacular name, papelillo. AmAtoua coryMBosa H. B. K.—Tabasco: Ojo de Agua, south- east. of Balancdn, May 9-14, Matuda 3131, a tree, 3 m. high, 7-10 em. in diameter, vernacular name, tarro de venado. Chiapas: Javalinero, in advanced forest, July 6-9, M atuda 3596, a tree, 10 m. high, 30 cm. in diameter. ANISOMERIS PROTRACTA (Bartl.) Standl.—Chiapas: Palenque, in savanna, July 9-14, Matuda 8730, a shrub; Aguacate, in savanna forest, July 16-18, Matuda 3789, a tree, 5 m. high, 20 em. in diameter. APPUNIA GUATEMALENSIS Donn. Sm.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3611, a shrub. BERTIBRA GUIANENSIS Aubl.—Chiapas: Javalinero, in ad- vanced forest, July 6-9, Matuda 3658, a shrub, corolla white. BLEPHARIDIUM MEXICANUM Standl., Field Mus. Bot., 22: 108. 1940 (Fig. 7).—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 8670 (type collection), a tree, 5-6 m. high, corolla white. 68 CYRUS LONGWORTH LUNDELL I'ie. 8. Chione chiapensis Standl. Matuda 3675 (type collection), half natural size. FLORA OF EASTERN TABASCO 69 BorRERIA SUAVEOLENS Mey.—Tabasco: Achotal, east of Balan- cAn, in savanna, May 9-14, Matuda 3056 (det. Standl.); Retiro, on limestone, riverside, June 19-25, Matuda 3467 (det. Standl.). CEPHAELIS TOMENTOSA (Aubl.) Vahl.—Tabasco: Achotal, east of Balancan, edge of forest, May 9-14, Matuda 3084, vernacular name, cresta de gallo. CEPHALANTHUS OCCIDENTALIS L.—Tabasco: Mercedes, lake- side, May 9-14, Matuda 3028, a shrub. Crrococca ALBA (L.) Hitche.—Tabasco: Retiro, riverside, on limestone, June 19-25, Matuda 3428, corolla yellow. Campeche: Vapor, July 25-28, Matuda 3906, corolla white. CHIONE CHIAPENSIS Standl., Field Mus. Bot., 22: 110. 1940 (Fig. 8).—Chiapas: ruins of Palenque, in advanced forest, river- side, July 10-12, Matuda 3675 (type collection), a tree, 8 m. high, 35 cm. in diameter. CoccocyesELUM HIrsuTUM Bartl.—Tabasco: Ojo de Agua, southeast of Balancdn, May 9-14, Matuda 3113, corolla purplish, fruits blue. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3667. Dropra riapa (Willd.) Cham. & Schlecht.—Tabasco: Achotal, east of Balancan, in savanna, May 9-14, Matuda 3107. FARAMEA occIpENTALIS (L.) Rich.—Tabasco: Reforma, May 22-26, Matuda 3184, a shrub; San Isidro, in advanced forest, June 7-11, Matuda 3344, a tree, 4 m. high, 25 cm. in diameter, corolla white. GoNZALAGUNIA PANAMENSIS (Cav.) Schum.—Tabasco: Retiro, in second growth, June 19-25, M atuda 3456, a shrub. Gurrrarpa Compsu Urban.—Tabasco: Reforma, May 22-26, Matuda 3187, a small tree, corolla white; Boca del Cerro, on lime- stone, July 1-5, Matuda 3591, a tree, 4—5 m. high. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3703, a tree, 8 m. high. GUETTARDA ELLIPTICA Swartz.—Tabasco: Reforma, May 22- 26, Matuda 3148, a small tree, 4 m. high, 12 em. in diameter; Provincia, May 27, Matuda 3216, a small tree; Boca del Cerro, on limestone, July 1-5, Matuda 3530, a small tree, 4 m. high; Boca del Cerro, in advanced forest, July 1-9, Matuda 3560, a 70 CYRUS LONGWORTH LUNDELL tree, 10 m. high, 25 em. in diameter. The continental collections, now referred to this species, may represent a distinct form, HAMELIA AXILLARIS Swartz.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3439, a shrub, corolla yellow. HAMELIA PATENS Jacq.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3245, a shrub. HOFFMANNIA REFULGENS (Hook.) Hemsl.—Tabasco: Teno- sique, in advanced forest, June 14-16, Matuda 8448 (det. Standl.). LINDENIA RIVALIS Benth.—Tabasco: Tenosique, riverside, June 27, Matuda 3498. Chiapas: Javalinero, in the Chacamax River, July 6-9, Matuda 3615. MACHAONIA AcumINATA H. B. K.—Tabasco: Kstapilla, edge of savanna, June 27, Matuda 3525, a tree, 3 m. high, 20 em. in diameter. Chiapas: Aguacate, swampside, July 16-18, Matuda 3814 (?), a tree, 7 m. high, 15 em. in diameter, flowers fragrant, corolla white. Campeche: Vapor, J uly 25-28, Matuda 3909, a tree, 5m. high, 25 em. in diameter, corolla white. MOoRINDA PANAMENSIS Seem.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3673, a tree, 8 m. high, 35 cm. in diameter. ORINDA YUCATANENSIS Greenm.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3026; Reforma, May 22-26, Matuda 3174, a shrub, corolla white. Two species may be repre- sented. PALICOUREA CROCEA (Sw.) R. & 8.—Tabasco: Achotal, east of Balancan, lakeside, May 9-14, Matuda 8036, a shrub. Campeche: Vapor, July 25-28, Matuda 3902, a shrub, corolla yellow. POSOQUERIA LATIFOLIA (Rudge) R. & S.—Chiapas: J avalinero, in advanced forest, July 6-9, Matuda 3619, a tree, 10 m. high, 25 cm. in diameter; ruins of Palenque, in advanced forest, July 10— 12, Matuda 3707, a tree, 10 m. high, 30 cm. in diameter, corolla white. PSYCHOTRIA BRACHIATA Sw.—Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3664 (det. Standl.), a shrub. PSYCHOTRIA CARTHAGINENSIS J acq.—Tabasco: Mercedes, in second growth, May 9-14, Matuda 3025 (det. Standl.), a shrub. FLORA OF EASTERN TABASCO call PsyCHOTRIA CHIAPENSIS Standl.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3423 (det. Standl.), a tree, 5 m. high, corolla white. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 8711 (det. Standl.), a small tree, 5 m. high, corolla white. Psycnorria cuspipata Bredem.—Tabasco: Achotal, east of Balancan, lakeside, May 9-14, Matuda 3037, a shrub, 2 m. high. Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3608, a shrub. Two species may be represented. PsycHoTrRIA FLAVA Oerst.—Tabasco: La Palma, June 1-6, Ma- tuda 3264 (det. Standl.), a shrub; La Palma, in advanced forest, Matuda 3286, a shrub, 2.5 m. high, corolla white; 5an Isidro, in advanced forest, June 7-11, Matuda 3373, a small tree. Chiapas: ruins of Palenque, in advanced forest, July 10-12, Matuda 3710 (det. Standl.), a small tree, 5 m. high. PsycHOTRIA FRUTICETORUM Standl—Tabasco: La Palma, June 1-6, Matuda 3266, a shrub. The species is either very variable or else several distinct forms are being confused with it. PsycHoTRIA INVOLUCRATA Swartz.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3623, a shrub. PsyCHOTRIA LIMONENSIS Krause.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3437, a shrub. Psycuorria micropon (DC.) Urban.—Tabasco: Estapilla, riverside, June 18, Matuda 3481, a shrub, corolla white. PsyCHOTRIA MOMBACHENSIS Standl.—Tabasco: La Palma, in advanced forest, June 1-6, Matuda 32386, a shrub, corolla white. PsycHorrRiA OERSTEDIANA Stand].—Tabasco: Reforma, in forest, May 22-26, Matuda 3146, a shrub, 2.5 m. high, 4-6 cm. in diameter. PsYCHOTRIA PUBESCENS Swartz.—Tabasco: Tenosique, in ad- vanced forest, June 14-16, Matuda 3393, a shrub, corolla yellow- ish. | PsyCHOTRIA SESSILIFOLIA Mart. & Gal.—Tabasco: San Isidro, June 7-11, Matuda 3890, a shrub. PsycHorrRIA UNDATA Jacq.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3366, corolla white. Ranpra AcULEATA L.—Tabasco: Mercedes, lakeside, May 9- 2 CYRUS LONGWORTH LUNDELL 14, Matuda 38019, a tree, 6-7 m. high, 30-40 em. in diameter; San Isidro, lakeside, June 7-11, Matuda 3347, a small tree; San Isidro, swampside, Matuda 3358, a small tree, 2.5 m. high. RicHARDIA scABRA L.— Chiapas: Javalinero, in second growth, July 6-9, Matuda 3659. RONDELETIA STACHYOIDEA Donn. Sm.—T'abasco: Juarez, near Tenosique, in advanced forest, June 14-16, Matuda 3392, a shrub, 3m. high. RuDGEA CERATOPETALA Donn. Sm.—Chiapas: Javalinero, in advanced forest, July 6-9, Matuda 3595 (det. Standl.), a shrub. SICKINGIA sp. Sickingia mollis Lundell, Contrib. Univ. Mich. Herb., 4:32. 1940, not Sickingta mollis Standl., Feld Mus. Bol., 11: 271. 1936.—Tabasco: Reforma, May 22-26, Matuda 3161 (type collection of S. mollis Lundell), a tree. In a recent treatment of the Mexican and northern Central American species of Sickingia (Standley, Field Mus. Bot., 22: 213. 1940), S. mollis Lundell was not taken into consideration. It is probable that the species is referable to one of the four now recognized by Standley, SPERMACOCE GLABRA Michx.—Tabasco: La Palma, June 1-6, Matuda 3238 (det. Standl.); La Palma, in second growth, Matuda 3265 (det. Standl.). CUCURBITACEAE ANGURIA DIVERSIFOLIA Cogn.—Tabasco: La Palma, edge of forest, June 1-6, Matuda 3318. MELOTHRIA GUADALUPENSIS (Spreng.) Cogn.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3309. SICYDIUM TAMNIFOLIUM (H. B. K.) Cogn.—Campeche: Pali- zada, lakeside, July 25-28, Matuda 3829. LOBELIACEAR (Determined by Rogers McVaugh, W ashington) LopetiA BrerLANpier1 A. DC.—Tabasco: Retiro, riverside, June 19-25, Matuda 3422, flowers blue. LOBELIA CARDINALIS var. GRAMINBA MecVaugh.—Tabasco: Tenosique, in savanna, June 14-16, Matuda 3408, corolla red. FLORA OF EASTERN TABASCO Tiss COMPOSITAE (Determined or verified by S. F. Blake, Washington) Asrur pxiuis Ell.—Chiapas: Javalinero, in second growth, July 6-9, Matuda 3631. CaLEA TRIcHoToMA Donn. Sm.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3756; Aguacate, July 16-18, Matuda 3784, a vine. CHAPTALIA NUTANS (L.) Polak.—Chiapas: ruins of Palenque, edge of forest, July 10-12, Matuda 3818. Cirsium mexicanum DC.—Chiapas: Javalinero, in second erowth, swampside, July 6-9, M atuda 3627, flowers white. EPALTES MEXICANA Less—Tabasco: La Palma, in second growth, June 1-6, Matuda 3315. ERECHTITES HIERACIFOLIA (L.) Raf.—Tabasco: Boca del Cerro, in swamp, July 1-5, Matuda 3581. Campeche: Palizada, swamp- side, July 25-28, Matuda 3858, flowers yellow. ERIGERON BONARIENSIS L.—Chiapas: Aguacate, in second growth, July 16-18, Matuda 3755. EUPATORIUM ALBICAULE Sch. Bip.—Tabasco: Reforma, May 22-26, Matuda 3197, a small tree, 3 m. high, 8 cm. in diameter; Achotal, east of Balancan, in advanced forest, May 9-14, Matuda 3067, a small tree, 3 m. high, 30 cm. in diameter, Tenosique, June 27, Matuda 3529. EupATORIUM MACROCEPHALUM Less.—Chiapas: Aguacate, in savanna, July 16-18, Matuda 3816. JeUPATORIUM MACROPHYLLUM L.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3268. CGoLDMANELLA SARMENTOSA Greenm.—Tabasco: La Palma, June 1-6, Matuda 3279. GyMNOCORONIS LATIFOLIA Hook. & Arn.—Campeche: Palizada, in swamp, July 25-28, Matuda 3844, flowers yellow. Har.eya oxyLepis (Benth.) Blake.—Tabasco: Retiro, in ad- vanced forest, June 19-25, Matuda 3445, flowers reddish. LIABUM SUBGLANDULARE Blake.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3311. MELANTHERA ANGUSTIFOLIA A. Rich.—Tabasco: Ojo de Agua, 74 CYRUS LONGWORTH LUNDELL southeast of Balancdn, in savanna, May 9-14, Matuda 3137; Estapilla, in savanna, June 27, Matuda 3502, flowers white. MELANTHERA HAsTaTA Michx.—Tabasco: Retiro, in second growth, June 19-25, Matuda 3450. NEUROLAENA LOBATA (L.) R. Br.—Tabasco: San Isidro, in advanced forest, June 7-11, Matuda 3335, flowers yellow. NOTOPTERA SCABRIDULA Blake.—Tabasco: Achotal, east of Balancan, May 9-14, Matuda 3032, a woody vine. ORTHOPAPPUS ANGUSTIFOLIUS (Sw.) Gleason.—Tabasco: Esta- pilla, in savanna, June 27, Matuda 3501, flowers white. PaRTHENIUM Hysreropuorus L.—Tabasco: La Palma, June 1-6, Matuda 3291. PECTIS SATUREJOIDES ( Mill.) Sch. Bip.—Tabasco: Kstapilla, in savanna, June 27, Matuda 3515, flowers yellow. PLUCHEA CAMPHORATA (L.) DC.—Campeche: Palizada, swamp- side, July 25-28, Matuda 3859, flowers reddish. STRUCHIUM SPARGANOPHORUM (L.) Kuntze.—Tabasco: Retiro, in advanced forest, June 19-25, Matuda 3419. VERNONIA CTENOPHORA Gleason (?).—Tabasco: Ojo de Agua, southeast of Balanedn, in advanced forest, May 9-14, Matuda 3112, flowers purplish. WEDELIA PARVICEPS Blake.—Chiapas: Palenque, in savanna, July 9-14, Matuda 3757. WEDELIA TRILOBATA (L.) Hitche.—Tabasco: La Palma, in second growth, June 1-6, Matuda 3258; Retiro, riverside, June 19-25, Matuda 3478, Campeche: Vapor, July 25-28, Matuda 8910. STUDIES OF AMERICAN SPERMA- TOPHYTES—II1* PLANTS OF TEXAS, CHIAPAS, AND BRITISH HONDURAS nN a co-operative project between the University of Michigan Herbarium and Southern Methodist University, a long term botanical survey of Texas was inaugurated in 1941. Field studies undertaken in Texas in 1940 were continued in 1941 and 1942. From Texas three new species in the genera Xylosma, Bumelia, and Citharezylum, two subspecies and two varieties of Phlox, and an albino form of Leucophyllum frutescens (Berl.) Johnston are described. Included also are a new genus, Belicea of the Rubiaceae, discovered by Percy H. Gentle in southern British Honduras, and six new species from Chiapas and British Honduras in the genera Colubrina, Aegiphila, Hyptis, Solanum, and Belicea. The writer is indebted to the Faculty Research Fund of the University of Michigan for support in part of the Texas survey in 1940 and 1942. Colubrina mollis Lundell, sp. nov.—Arbor parva; ramulis pubescentibus. Folia petiolata, petiolo 0.8-1.5 cm. longo, lamina membranacea, ovato-oblonga vel ovato-elliptica, 8.5-16 cm. longa, 4.3-8 cm. lata, apice acuminata, basi rotundata vel sub- cordata et abrupte acuta, remote dentato-serrata, supra glabrata, subtus pilosa, 3-nervia. Flores axillares, pedicellis 1.5-3.5 mm. longis. Calyx 5-lobatus, lobis 1.7-2 mm. longis. Petala 5, glabra. Stamina 5. An unarmed tree, 12 to 15 m. high, 40 cm. in diameter; branch- lets slender, straight, densely pubescent with pale reddish brown hairs. Petioles pubescent, slender, 0.8 to 1.5 em. long. Leaf blades membranaceous, ovate-oblong or ovate-elliptic, 8.5 to 16 cm. long, 4.3 to 8 cm. wide, apex acuminate, base rounded or subcor- date and abruptly acute, the acute base triangular, paler beneath, margin coarsely and remotely dentate-serrate, the teeth rounded, * This paper has been published with funds derived from the income of the endowment of the Horace H. Rackham School of Graduate Studies of the University of Michigan. 75 76 CYRUS LONGWORTH LUNDELL at first densely pubescent on both surf aces, upper surface glabrous with age except along the veins, lower surface persistently and densely soft pilose or subtomentose, 3-nerved at base, the veins impressed above, conspicuous beneath. Flowers numerous, fasci- culate in the leaf axils or borne in dense subsessile cymes, pu- bescent. Pedicels slender, 1.5 to 3.5 mm. long. Calyx 5-lobed, the lobes triangular-ovate, 1.7 to 2mm. long, glabrous within. Petals glabrous, cucullate, subequaling calyx lobes. Stamens 5. Disk flat. Mexico: Chiapas, between Mazapa and Motozintla, altitude 1200 m., in advanced forest, July 19, 1941, Hizi Matuda 4873 (type in the University of Michigan Herbarium), The large thin leaves and persistent soft pubescence of the undersurface of the blade are characteristics by which C. mollis may be readily separated from C. glomerata (Benth.) Hemsl., the species to which it apparently has affinity. Xylosma blepharodes Lundell, sp. nov.—Frutex; ramulis novellis puberulis. Folia petiolata, fasciculati, chartacea vel sub- coriacea, oblanceolata, elliptica, vel ovato-elliptica, 1-5.5 em. longa, 0.5-2.2 em. lata, apice obtusa, basi cuneata, crenulato- serrulata. Flores fasciculati. Pedicelli fructiferi ad 6 mm. longi. Sepala 5, ciliata. Bacca subglobosa, ad 6 mm. diam. Slender spiny shrub, about 2 m. high; branchlets slender, at first densely puberulent; spines slender, straight, up to 3.5 em. long. Leaves alternate, usually clustered on short. spurlike branchlets of older wood; petioles 0.5 to 2.5 mm. long, puberulent above. Leaf blades chartaceous to subcoriaceous, green, only slightly paler beneath, oblanceolate, elliptic, or ovate-elliptic, 1 to 5.5 cm. long, 0.5 to 2.2 em. wide, apex obtuse or rounded, base attenuate-cuneate to broadly cuneate, margin crenulate-serrulate, the teeth rather remote, irregular and obtuse, puberulent above along the costa, glabrous otherwise, primary veins 5 or 6 on each side, slender, evident on both surfaces, reticulation inconspicuous. Flowers fasciculate. Pedicels slender, glabrous, jointed at base, up to 4 mm. long in fruit, usually short, the fruits sometimes sub- sessile. Sepals 5, unequal, ovate-oblong, 0.6 to 1.4 mm. long, acute, pubescent, conspicuously ciliate with stiff hairs. Fruits STUDIES OF AMERICAN SPERMATOPHYTES HI 77 subglobose, up to 6 mm. in diameter, glabrous; styles persistent, almost 1 mm. long. Texas: Cameron County, in Palm Grove, south of Browns- ville, April 6, 1941, C. L. Lundell and Amelia A. Lundell 10000 (type in the University of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University). Bumelia riograndis Lundell, sp. nov.—Frutex; ramulis glabra- tis. Folia petiolata, petiolo 1.5-6 mm. longo, lamina chartacea, obovato-elliptica vel elliptica, 1.6-3.5 em. longa, 1-1.8 em. lata, apice rotundata, emarginata, basi cuneata. Flores fasciculati. Pedicelli ad 3 mm. longi. Sepala 5, 2-2.5 mm. longa. Corolla 3 5-4 mm. longa, glabra. Ovarium hirsutum. Spiny shrub; branchlets reddish, short, nodose, glabrate, spine tipped. Leaves clustered on short. spurlike branchlets of older wood; petioles slender, 1.5 to 6 mm. long, covered at first with loose hairs, glabrous with age. Leaf blades chartaceous, glabrous except for loose hairs along the midvein, obovate-elliptic or ellip- tic, 1.6 to 3.5 cm. long, 1 to 1.8 cm. wide, apex rounded, shallowly emarginate, cuneately narrowed at the base or acute, conspicu- ously but finely reticulate veined on both surfaces, the veins whitish, primary veins 7 or 8 on each side, costa plane above, ele- vated beneath. Flowers numerous, fasciculate. Pedicels slender, up to 3 mm. long, with a few loose hairs at anthesis. Calyx glabrescent; sepals 5, unequal, ovate-orbicular or depressed orbi- cular, 2 to 2.5 mm. long. Corolla glabrous, 3.5 to 4 mm. long, the lobes suborbicular, about 2.2 mm. long, minutely erose; append- ages lanceolate, about 2 mm. long, acutish, erose. Staminodia petaloid, ovate, 2mm. long, acutish. Ovary hairy. Style glabrous, about 1.5 mm. long. Texas: Jim Hogg County, between Hebbronville and Rio Grande City, July, 1928, B.C. Tharp s.n. (type in the University of Michigan Herbarium). Closely allied to B. texana Buckl. PHLoXx NIVALIS Lodd. subsp. texensis Lundell, subsp. nov.— Suffruticosa. Caules pubescentes, ad 45 cm. longi, dense foliati. Folia rigidula, lineari-subulata vel lineari-lanceolata, 2.6-30 mm. 78 CYRUS LONGWORTH LUNDELL longa, ad 3 mm. lata. Cymae 1-5-florae. Pedicelli 3-25 mm. longi. Calyx glandulosus, 6.5-9 mm. longus, lobis tubo aequilongis vel subaequilongis. Corolla glabra, lobis bifido-emarginatis. Stylus -0—2.5 mm. longus; ovarii loculi uniovulati vel biovulati, Semina ellipsoidea, 2.2 mm. longa, rugulosa. Perennial, stems woody at base, trailing or decumbent, up to 45 cm. long, with numerous axillary latent shoots, often matted; flowering shoots lax, erect, up to 20 em. long, pubescent, the hairs inconspicuously glandular. Leaves persistent, sessile, very dense on the latent shoots, linear-subulate and canaliculate or linear-lanceolate and essentially plane, 2.5 to 30 mm. long, usually acicular, rarely up to 3 mm. wide, apex spinescent, usually re- flexed, the margin ciliate with stiff hairs. Cymes usually 3- to 5- flowered, sometimes reduced to 1 flower. Pedicels slender, glandu- lar-pubescent, 3 to 25 mm. long. Calyx glandular-pubescent, 6.5 to 9mm. long, the lobes equaling or subequaling the tube, spines- cent. Corolla glabrous outside, variable in size and color, fragrant, pink, rose-pink, bluish, lavender, or purplish, the eye reddish or purplish; tube up to 15 mm. long, villous within at base; lobes usually 5, rarely 4, obovate or cuneately obovate, up to 12 mm. long, typically with a conspicuous sinus up to 2.5 mm. deep, rarely only erose and shallowly emarginate, Stamens usually 5, rarely 4, usually included, rarely with tips of anthers exserted from corolla tube. Ovary 3-celled, usually with 1 ovule in each cell, sometimes with 2 ovules in a cell; style short, 1.5 to 2.5 mm. long; stigma 3-lobed, 1 to 1.3 mm. long. Capsule glabrous; seed ellipsoid, 2.2 mm. long, rugulose. Texas: Tyler County, south of Woodville, in rolling pine land, April 18, 1942, C. L. Lundell and Amelia A. Lundell 11282 (type in the University of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University); about 4 miles south of Woodville, in pine land, April 13, 1942, Lundell and Lundell 11122, 111265, 1 1126; about 5 miles north of Warren, in rolling pine land, April 18, 1942, Lundell and Lundell 11278; about 6 miles north of Warren, in rolling pine land, April 18, 1942, Lundell and Lundell 1 1280; south of Woodville, in rolling STUDIES OF AMERICAN SPERMATOPHYTES— I 79 pine land, April 18, 1942, Lundell and Lundell 11284, vernacular name, moss phlox. The Texas plant, first called to the writer’s attention by Edgar T. Wherry, is abundant in the pine lands off U. S. highway 69, between Woodville and Warren in Tyler County. Mr. and Mrs. G. M. Soxman of Pallas directed the author to this locality. The subspecies may be distinguished immediately by the pro- nounced sinus of the corolla lobes. In the typical form of the spe- cies, the lobes are merely erose or inconspicuously emarginate. The glandular hairs of P. nivalis texensis are scarcely evident in dried material. Further, the leaves of the latent shoots apparently are more acicular and denser than in the usual form of the species. Puiox PiLosA L. subsp. pulcherrima Lundell, subsp. nov.— Perennis. Caulis erectus, ad 60 cm. altus, pubescentibus. Folia sessilia, linearia vel anguste lineari-lanceolata, ad 12.5 cm. longa, 2-6 mm. lata, ciliata. Flores in corymbis terminalibus, pedicellis 3-18 mm. longis. Calyx eglandulose villosus, lobis linearibus tubo longioribus. Corolla rosea, glabra. Stylus 1-1.3 mm. longus. Perennial, erect, up to 60 cm. high, plants usually solitary; stems terete, pubescent with short eglandular recurved hairs, glabrescent, the latent shoots erect, shorter than the flowering shoots; internodes elongate above; nodes 7 to 10. Leaves opposite, sessile, chartaceous, linear or narrowly linear-lanceolate, up to 12.5 em. long, 6 mm. wide, usually less than 3 mm. wide, at- tenuate to the spinescent apex, sparsely pubescent, glabrescent, persistently ciliate at base with coarse hairs, upper surface ob- scurely papillate, the midvein impressed. Cymes in terminal corymb; the inflorescence variable, open to compact, usually few- flowered, up to 15 em. wide, 12 em. high; foliaceous bracts spread- ing or reflexed, those at base lanceolate, slightly wider than the leaves. Pedicels often less than 5 mm. long, sometimes up to 18 mm. long, pubescent with short recurved eglandular hairs. Calyx up to 13 mm. long, the lobes longer than the tube, subulate or linear, conspicuously awn-tipped. Corolla variable in size, rose- pink with darker reddish eye; tube elabrous outside, up to 21 80 CYRUS LONGWORTH LUNDELL mim. long, short villous within at base; lobes glabrous, obovate, up to 16 mm. long, 15 mm. wide, apex apiculate or rounded. Upper stamens barely included; anthers oblong, up to 3 mm. long. Ovary with 1 ovule in each cell; style very short, 1 to 1.3 mm. long; stigma up to 2mm. long. Texas: Angelina County, east of Zavalla, off highway 68, in pine land, May 8, 1941, C. L. Lundell and Amelia A. Lundell 10557. Cherokee County, northwest of Rusk, off U. 8. highway 69, in woodland, May 8, 1941, Lundell and Lundell 1 0568, Hardin County, about 2 miles southwest of Kountze, in pine land, April 14, 1942, Lundell and Lundell 11158, 11154 (type in the Univer- sity of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University). Jasper County, northwest of Jasper, off highway 63, in pine land, May 8, 1941, Lundell and Lundell 10558. Polk County, east. of Livingston, in pine land, April 18, 1942, Lundell and Lundell 11287. Sabine County, near Bronson, off U. S. highway 96, in pine land, May 7, 1941, Lundell and Lundell 10505; Brookeland, in pine land, May 7, 1941, Lundell and Lundell 10508. Tyler County, about 4 miles south of Woodville, in pine land, April 13, 1942. Lundell and Lundell 11124; northwest of Woodville, in pine land, April 13, 1942, Lundell and Lundell 11 164; about 5 miles north of Warren, in rolling pine land, April 18, 1942, Lundell and Lundell 11276 ; south of Woodville, in rolling pine land, April 18, 1942, Lundell and Lundell 11281, flowers faintly fragrant. Walker County, east of Huntsville, in forest, April 18, 1942, Lundell and Lundell 11295. The plant is currently passing as Phlox pilosa var. amplexi- caulis (Raf.) Wherry. The true identity of P. amplexicaulis Raf. is very doubtful, for the description is so broad and inadequate that it could be interpreted to cover several species, as well as various southern forms of P. pilosa. To assign the name arbitrarily can serve no useful purpose. Brand referred it with doubt to P. divaricata LL. (Pflanzenreich IV, 250: 72. 1907). P. pilosa pulcherrima is one of the finest of the Texas phloxes. The beauty of its large rose-pink corollas is scarcely equaled by any other form. In Tyler County, evidently the center of distribu- tion, it occurs throughout the pine lands. STUDIES OF AMERICAN SPERMATOPHYTES—HU1 81 The long straggly eglandular hairs of the calyx, the slender calyx lobes, the glabrous rose-pink corolla, the tall slender habit, the long linear leaves sparingly pubescent and persistently ciliate at base are the primary differentiating characteristics of the sub- species. The wide field observations indicate that there is a close correlation between the type of pubescence and corolla color. PHLOX PILOSA PULCHERRIMA var. typica Lundell, var. nov.— Calyx eglandulose villosus. Corolla glabra. The usual widespread form of the subspecies, typified by Lun- dell and Lundell 11154, has large glabrous corollas. Another form, apparently uncommon, has pubescent corolla tubes; it may be known as follows: PHLOX PILOSA PULCHERRIMA var. pubiflora Lundell, var. nov. Calyx eglandulose villosus. Corollac tubus parce pubescens. Tpxas: Cherokee County, northwest of Wells, off U.S. highway 69, in woodland, May 8, 1941, C. L. Lundell and Amelia A. Lundell 10561. Walker County, east of Huntsville, in forest, April, 18, 1942, Lundell and Lundell 11472 (type in the Univer- sity of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University), perennial, corolla rose-pink, the tube short villous, eglandular. Aegiphila wigandioides Lundell, sp. nov.—Arbor parva, ramu- lis cinereo-tomentosis. Folia petiolata, petiolo 1-1.4 cm. longo, lamina membranacea vel subchartacea, ovata vel lanceolata, 7-14 em. longa, 4-7 cm. lata, apice attenuata, acuminata, basi rotundata, supra villosa, subtus cinereo-tomentosa. Cymae axillares et terminales, pedunculatae, cinereo-tomentosae. Calyx ca. 1.2 em. longus, lobis 4. Corolla glabra. Stamina 4. A tree, 7 meters high; branchlets terete, canescent-tomentose, becoming glabrate in age and reddish brown. Leaves decussate ; petioles 1 to 1.4 cm. long, tomentose. Leaf blades membranaceous or subchartaceous, grayish green, paler beneath, ovate or lanceo- late, 7 to 14 cm. long, 4 to 7 ecm. wide, apex attenuate, acuminate, base rounded, margin entire, densely and persistently short villous above, the hairs multicellular and enlarged at base, canescent- 82 CYRUS LONGWORTH LUNDELL tomentose beneath, costa prominent on undersurface, primary veins conspicuous, 5 to 7 on each side. Cymes axillary and ter- minal, solitary in the axils, compactly flowered, canescent- tomentose; peduncles stout, up to 2.5 em. long; bractlets filiform, villous. Pedicels slender, up to 2 mm. long, villous. Calyx ob- conic-campanulate, about 1.2 em. long, 5 mm. wide, villous, 4-lobed, the lobes ovate-triangular, 2 to 3 mm. long, obtuse. Corolla glabrous, infundibular, the tube slender, 1.1 to 1.3 em. long, ampliate above; lobes 4, elliptic or oblong-elliptic, up to 5.5 mm. long, 4 mm. wide, apex obtuse or rounded. Stamens 4, equal, exserted. Ovary glabrous; style filiform ; stigma bifid. Mexico: Chiapas, Santa Rosa, near Escuintla, altitude 1600 m., in advanced forest, June 20,1941, Hizi Matuda 4253 (type in the University of Michigan Herbarium). Apparently, the species is related to A. Deppeana Steud., but the denser persistent indument and large calyx amply distinguish it. The superficial resemblance to Wigandia of the Hydrophylla- ceae is remarkable. Citharexylum spathulatum Moldenke & Lundell, sp. novy.— Frutex, ramulis glabris. Folia chartacea, glabra, spathulata, 0.8— 2.5 cm. longa, 2.5-5 mm. lata, apice rotundata, basi attenuata : petiolis nullis. Inflorescentiae 1- raro 2-florae, terminales. Calyx campanulatus, ca. 3.5 mm. longus. Corolla alba, 6 mm. longa. Stamina 4, Ovarium glabrum. A shrub, 1 m. high, openly branched; branches and branchlets slender, glabrous; twigs usually sharp pointed, quadrangular or subterete, the youngest shoots short hirtellous at the nodes, es- sentially glabrous otherwise. Leaf sears small, short stalked; leaves opposite on young shoots, clustered on short spurlike branchlets of older wood; petioles none. Leaf blades green, char- taceous, entirely glabrous or with a few short stiff hairs at apex, spathulate, usually 0.8 to 2.5 em. long, sometimes shorter, 2.5 to 5mm. wide, apex usually rounded, sometimes shallowly emargin- ate, acutish, or apiculate, base attenuate and very slender, entire, midrib discernible but inconspicuous, veins not evident. Flowers fragrant, usually 1, rarely 2, terminating the very short spurlike STUDIES OF AMERICAN SPERMATOPHYTES— III 83 branchlets. Pedicels very short, densely white-hirtellous. Calyx campanulate, about 3.5 mm. long, contracted at base into stipe about 0.5 mm. long, usually 5- rarely 4-ribbed, the rim bearing 5 thick reflexed teeth, truncate or toothed between the primary teeth, sparsely short appressed-hirtellous, ciliolate. Corolla white, sparsely hairy, 6 mm. long; the tube about 3.5 mm. long; the lobes ovate, rounded at apex, ciliate; throat and upper half of tube densely barbate. Stamens 4, included; filaments hairy; staminode rudimentary, hairy. Ovary glabrous; style 1 mm. long, basal half stipitate-glandular. Fruits unknown. Texas: Hidalgo County, 11 miles north of Mission, on sand in open scrub, April 4, 1941, C. L. Lundell and Amelia A. Lundell 9958 (type in the University of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University). This is evidently the plant described as Cztharexylum brachy- anthum var. glabrum C. L. Hitche. & Moldenke (Fedde, fepert., 37: 218. 1934). The glabrous longer leaves, glabrous branchlets, and differences in the flowers indicate that a distinct species is represented. In C. spathulatum the calyx rim bears 5 thick reflexed teeth terminating the ribs of the tube; between these primary teeth, the margin is truncate or shallowly toothed. The calyx rim is sugges- tive of the genus Lycianthes in the Solanaceae. In C. brachyanthum the calyx has 5 conspicuous teeth with well-developed sinuses be- tween each. Further, the flowering calyx of C. brachyanthum is somewhat smaller, the ribs are less evident, and the pubescence is dense. The corolla of C. brachyanthum is a third smaller. The interpretation of the flowers of C. brachyanthum is based upon Forrest Shreve 8421 from Coahuila, Mexico. Hyptis belizensis Lundell, sp. nov.—Herba erecta, ramulis novellis adpresse hirtellis. Folia petiolata, petiolo marginato, lamina subcoriacea, hirtella, anguste oblongo-lanceolata vel ob- lanceolata, 6-15 cm. longa, 0.7-1.7 cm. lata, apice acuta vel acuminata, basi attenuata, supra bullata. Inflorescentiae capi- tatae, axillares, pedunculatae. Pedunculi ad 3 cm. longi. Capitula ad 2.5 cm. diam. Calyx ad 1 cm. longus. Corolla alba, tubo ad 5 mm. longo. 84 CYRUS LONGWORTH LUNDELL An erect perennial herb, up to 90 cm. high, with woody root 5 cm. in diameter; stems quadrangular and densely appressed-hir- tellous above, subterete and glabrous below, reddish brown. Petioles narrowly winged to base by the narrowly attenuate de- current leaf blades, appressed-hirtellous. Leaf blades rigid, sub- coriaceous, minutely hirtellous on both surfaces, narrowly oblong- lanceolate or oblanceolate, 6 to 15 em. long (including petioles), 0.7 to 1.7 em. wide, apex attenuate, acute or acuminate, base at- tenuate, decurrent to the stem, margin irregularly serrate, costa and veins impressed above, prominent beneath , the upper surface bullate. Peduncles solitary, axillary, appressed hirtellous, up to 3 cm. long. Flowers in globose echinate capitula up to 2.5 em. in diameter, subtended by lanceolate acuminate spreading or re- flexed bracts up to 11 mm. long. Calyx at maturity up to 1 cm. long, the erect stiff subulate teeth subequaling the tube, the in- dument persistent. Corolla white, pubescent, its tube up to 5 mm. long. Nutlets smooth. British Honpuras: Toledo District, Swasey Branch, Monkey River, edge of hammock in pine ridge, Jan. 28, 1942, Percy H. Gentle 8896 (type in the University of Michigan Herbarium), flowers white, becoming reddish after four days. H. belizensis, which is nearest H. lanceolata Poir., differs from that species in having very narrow leaf blades, persistent indu- ment, and long subulate calyx teeth. The peculiarities of the British Honduran plant were first pointed out by Epling (Car- negie Inst. Wash. Publ., 522: 242. 1940). Solanum Gentlei Lundell, sp. nov.—Arbor parva; ramulis al- bido-stellato-tomentosis. Folia petiolata, petiolo ad 5 mm. longo, lamina membranacea, supra viridis, subtus albido-stellato-tomen- tosa, oblongo-lanceolata, anguste elliptica, vel oblanceolata, 8-22 em. longa, 3.5-7.7 em. lata, apice subabrupte acuminata, basi acuta. Inflorescentiae terminales, albido-tomentosae, cymoso- corymbosae, ad 6 cm. longac. Pedicelli ad 1 em. longi. Calyx ca. 2.8 mm. longus. Corolla ca. 6 mm. longa, profunde lobata. Bacca globosa, glabra, ca. 6.5 mm. diam. A small unarmed tree; branchlets slender, white tomentose STUDIES OF AMERICAN SPERMATOPHYTES—III 85 with fine stellate hairs. Leaves alternate, sometimes with a re- duced opposite leaf; petioles short, canaliculate, tomentose, usu- ally less than 5 mm. long, rarely up to 8 mm. long. Leaf blades membranaceous, dark green above and sparsely stellate pu- bescent, persistently white tomentose on undersurface with fine stellate hairs, the hairs sessile and stalked, oblong-lanceolate, nar- rowly elliptic, or oblanceolate, 8 to 22 cm. long, 3.5 to 7.7 cm. wide, apex subabruptly acuminate, base acute, often with sub- equal sides, margin entire, veins impressed above, conspicuous beneath, primary veins 5 or 6 on each side. Inflorescence terminal, white tomentose, cymose-corymbose, up to 6 em. long, with peduncles up to 2.5 cm. long. Pedicels usually recurved, slender below, thickened above, up to 1 cm. long in fruit. Calyx white tomentose with fine stellate hairs, campanulate, about 2.8 mm. long, 5-lobed almost to the middle, the lobes apiculate, scarcely accrescent in fruit. Corolla about 6 mm. long, the lobes ovate- oblong, about 3.5 mm. long, acutish, finely stellate-tomentose outside, glabrous within. Anthers blunt. Ovary white tomentose above. Fruits globose, glabrescent, about 6.5 mm. in diameter. BrrtisH Honpuras: Toledo District, Swasey Branch, Monkey River, in high ridge between river and pine land, March 7, 1942, Percy H. Gentle 3944 (type in the University of Michigan Herb- arium), small tree, leaves white on underside, flowers white, berries green. The species is related to S. salviifolium Lam. Solanum mayanum Lundell, sp. nov. Frutex parce aculeatus; ramulis stellato-tomentosis. Folia longe petiolata, petiolo ad 5.5 em. longo, lamina sinuato-lobata, membranacea, elliptica vel ovato-elliptica, 8-15 cm. longa, 3 5-10 cm. lata, apice acuta vel acuminata, basi rotundata et abrupte acuta, supra stellato-hir- tella, subtus stellato-pilosa. Inflorescentiae extra-axillares, cy- mosae, stellato-tomentosae, pedunculatae. Pedicelli fructiferi ad 1.8 cm. longi. Calyx stellato-tomentosus, 5 mm. longus, ad medium 5-lobus. Corolla alba, fere 1.3 cm. longa, fere ad basin 5-loba. Antherae 6-7 mm. longae. Bacca globosa, glabra. A shrub; branches armed with prickles up to 5 mm. long; S6 CYRUS LONGWORTH LUNDELL branchlets slender, sparsely aculeate with prickles about 2 mm. long, gray tomentose with appressed stellate hairs. Leaves alter- nate; petioles slender, sometimes armed, gray tomentose, canalic- ulate, up to 5.5 em. long. Leaf blades membranaceous, elliptic or ovate-elliptic, 8 to 15 cm. long, 3.5 to 10 cm. wide, apex acute or short acuminate, base rounded and acute, the sides often un- equal, margin sinuate-lobate or sinuate-dentate, usually with two or three shallow lobes on each side, sometimes entire or subentire, green and sparsely stellate-hirtellous above, paler and stellate- pilose beneath, the hairs with a short central ray. Inflorescence cymose, extra-axillary, up to 4.5 em. long, gray tomentose with stellate hairs. Pedicels of flowers slender, up to 1.3 em. long, thickened above and up to 1.8 em. long in fruit. Calyx thinly stellate-tomentose, 5 mm. long, 5-lobed to the middle, the lobes ovate-triangular, apiculate, not accrescent. Corolla white, about 1.3 em. long, thinly stellate-tomentose outside, 5-lobed almost to the base, the lobes lanceolate, attenuate. Anthers attenuate, 6 to 7 mm. long. Ovary very inconspicuously stipitate-glandular, essentially glabrous. Fruits globose, glabrous. British Honpuras: Toledo District, near Cowpen, Swasey Branch, Monkey River, in broken ridge, March 16, 1942, Percy H., Gentle 3976 (type in the University of Michigan Herbarium), vernacular name, broken ridge sosumbra. S. mayanum has affinity to 8S. H offmannit Bitter of Costa Rica, LEUCOPHYLLUM FRUTESCENS (Berl.) Johnston f. albineum Lun- dell, f. nov.—A forma, typiea corolla albo-recedit. EXAS: Cameron County, about 8 miles west of Boca Chica, in scrub on low ridge, March 12, 1942, C. L. Lundell and Amelia A. Lundell 10699 (type in the University of Michigan Herbarium, duplicate in the herbarium of Southern Methodist University), a shrub, 4 feet high, flower buds yellowish, corolla white. This rare albino form is well worthy of attention by horticul- turists. In typical L. frutescens the corollas are rose-pink and maculate in throat. Belicea Lundell, gen. nov. Rubiacearum.—Frutex. Stipulae persistentes, abrupte cuspidatac. Folia longe petiolata, magna, STUDIES OF AMERICAN SPERMATOPHYTES—IHIII 87 membranacea. Inflorescentiae capitatae, axillares, fasciculatae, brevissime pedunculatae, floribus sessilibus. Calyx cum hypanthio ca. 1.5 mm. longus, subtruncatus, minute 5-denticulatus. Corolla subtubulata, parva, lobis valvatis. Stamina, 5. Antherae semiex- sertae. Ovarium 4-loculare, loculis uniovulatis. Capitula fructifera paucicarpellata, mammosa. Type species, Belicea hoffmannioides Lundell. The genus, referable to the tribe Morindeae, is nearest Morinda. It differs from that genus primarily in having fruits which cohere at base only to form a mammose head. The fruits of Belicea superficially resemble those of Castilla elastica Cerv. In Morinda the fruit is a fleshy globose syncarp. In Mexico and Central America, British Honduras is known as Belice, the name which the genus commemorates. Belicea hoffmannioides Lundell, sp. nov.—Frutex. Stipulae abrupte cuspidatae, ad 4 mm. longae. Folia longe petiolata, mem- branacea, elliptica vel obovato-elliptica, 11-26 em. longa, 3-11 em. lata, apice subabrupte acuminata, basi attenuata. Inflores- centiae pauciflorae. Flores albi. Corolla puberula, 7 mm. longa. Filamenta parce pilosa. Capitula fructifera ad 1.5 cm. diam. A shrub; branchlets rather stout, densely puberulent, drying black. Stipules puberulent, persistent, ovate-triangular or sub- orbicular, abruptly cuspidate, up to 4 mm. long. Petioles slender, canaliculate, minutely puberulent, 2 to 3.5 cm. long, slightly winged above by the decurrent blade. Leaf blades membranace- ous, elliptic or obovate-elliptic, 11 to 26 cm. long, 3 to 11 cm. wide, apex subabruptly acuminate, base attenuate, decurrent, the margin undulate, costa and veins minutely puberulent on under- surface, glabrous otherwise, costa nearly plane above, prominent beneath, primary veins 9 to 12 on each side, anastomosing near the margin. Flowers capitate, sessile, white; heads borne on short peduncles fasciculate in the leaf axils; subtending bracts, pedun- cles, calyx, and corolla puberulent. Calyx, including hypanthium, about 1.5 mm. long, subtruncate, the 5 teeth minute. Corolla 7 mm. long, including lobes about 2 mm. long, the tube pubescent within throat, the lobes valvate, triangular-ovate, acutish. 88 CYRUS LONGWORTH LUNDELL Stamens 5; filaments inserted at middle of tube, about 1.8 mm. long, sparsely pilose; anthers versatile, attached below middle, linear-oblong, about 1.8 mm. long, apical part exserted from tube. Ovary 4-celled, with a solitary ovule in each cell attached near base of septum; style included, puberulent; stigma bilobed. Fruits coherent at base only, depressed apically, containing 2 to 4 flat pyrenes 4.5 to 6 mm. long; the heads mammose, up to 1.5 em. in diameter, borne on peduncles less than 5 mm. long. British Honpuras: Toledo District, Swasey Branch, Monkey River, near the falls, in high ridge, March 7, 1942, Percy H. Gentle 8945 (type in the University of Michigan Herbarium), In habit and leaf form the species suggests Hoffmannia. LUNDELL The Usumacinta River with a view of Boca del Cerro. LUNDELL PLATE II TEE vs fe: va Pie ra ‘S. The Usumacinta River with an inner view of Boca del Cerro. AB AAA AAS AAS LUNDELL The expedition crossing a savanna east of Palenque. LUNDELL PLATE IV Javalinero on the Chacamax River below Palenque. Abutilon, 42 Achras, 55 pasar eanea, 19 Andira 32 Anguria, 72 Aniseia, Anisomeris, 67 dae Arthr sttdium, 12 Asclepiadaceae, Asclepias, 58 eae 9 Aster, 73 INDEX (Synonyms in italics) Athenaea, 63 Axonopus, 12 Bacopa, 64 Bactris, 16 Balanophoraceae, 25 Bromelaccne a, 21 Burmanniaceae, 21 Bursera, 35 Burseraceae, 35 Soeur 26 28 alea, 7 Calliandra, 29, 30 Callicarpa, 60 Calyptranthes, 49 Calyptrocarya, 14 90 Caperonia, 37 apparidaceae, 28 Cae 12 Chamaedorea, 16 Champa, Chaptalia, 73 ee a Chenopodiun Chile ea Zr C ee ophor a, 23 Chr ee ae 28 Oitharacs ies: 60, 61 Cladiur 4 Gee 61 nese 51, 52 32 Combretum, 49 CYRUS LONGWORTH LUNDELL neve Commelina, 18 onnaraceae, 29 Eien 52 ee Goqicche. 57 Crataeva, 28 Cresta de gallo, 69 Crotalaria, 32 roton, 37, 38 Crusita, 28 Cucurbitacene, 72 , 64 Cymbopetalum, 27 Cynometra, 32 Cyperaceae, 14 Cyperus, 14, 15 Palbercia, 32, 33 Dracaena, 18 sas fea arpus, 33 era, 28 rosera, ees 28 Drymonia, 65 Dryopteris, 9 Duranta, 61 I po vaceae, 5D Eryngium, 54 Teena incene 34 Erythroxylon, 34 Escobedia, Bugenia, 48-51 Eupatorium, 73 Euphorbia, 8 Huphorbiaceae, 37 Eustoma, 57 eae 59 Fagaceae, 23 Faramea, 69 Ficus, 24 Fimbristylis, 15 Fischeria, 58 Flacourtiaceae, 45 Fuirena, Galactia, 33 Garcia, 38 Gentianaceae, 57 Geonoma, 16, 17 Gerardia, 64 Gesneriaceae, 65 Gonsloue 5) INDEX Gonzalagunia, 69 e Gymnocoronis, 73 Gymnopodium, 26 Gymnopteris, 9 Haemodor aceae, 19 0 Hippocratea, 41 Hippocrateaceae, 41 Hiraea, 36 Hirtella, 28, 29 Hoffmannia, Ms 88 Homolepis, Hypolytrum, 15 Hyptis, 61, 75, 83 Icatillo, 29 Iridaceac, 20 92 Jacquemontia, 59 Justicia, 65 Pabiaiac, 61 Leguminosae, 29 Lentibulariaceae, 65 Leptocoryphium, 13 Leucophy llum, 75, 86 73 Isrianthes, 83 Lopodi, i rye odiu ,l nee 30 Lythraceae, 47 Lythrum, 47 Mabea, 38 Macherium, 34 na 70 Macha a Malva, 42 CYRUS LONGWORTH LUNDELL Seen Va 58 Manisuris, 13 vere 21 Marantaceae, PA | Marcgraviaceae, 44 Maytenus, 40 Melanthera, 73, 74 Melastomaceae, 51 Meliaceae, 35 Melochia. 43 Melothria, 72 Menispermaceac, 27 Mimoseae, 26 Monimiaceae, 27 Monstera, Myrsinace ae, 54 Myrtaceae, 49 Nectandra, 27, 28 Nyctaginaceae, 26 Nymphaea, 27 Se ee 26 Ochnaceae, 44 Odontonema, 67 Olacaceae, 25 mene 57 Olyra Ope, 53 One Ot $ ee atea, 44 Oxalidaceae, 34 Oxalis, 34 Palicourea, 70 Palmae, 16 Palo retinto, 35 eran 54 Passifloraceae, 47 Pectis, 74 Pentarthaphis, 14 Pepero , 22 Phoradendron, 24, 25 Phyla, 61 Phyllanthus, 38, 40 Physalis, 63 Di 4 1 26 rhytoia ) See 35 eee Pim ,5 See de tierra, 51 3 Pistia, 17 INDEX Pitcairnia, 18 Pithecolobium, 30, 31 Plantaginaceae, 67 Plantago, 67 Pleiostachya, 21 Pleurothallis, 22 Pluchea, 7 Pristimera, 41 Protium, 35 Psittacanthus, 25 Psychotria, 70, 71 Pteris, 10 Quercus, 23 Randia, 71 Rhizophoraceae, 47 Rhynchosia, 34 Rhy oe 15, 16 Richardia 72 Rubiaceae, 67 94. R udgea, TZ Rutaceae, 35 Sager etia, 42 . 64 Serophulariaceae, 63 Scutellaria, 63 Sebastiania, 39, 40 Sicydium, 72 Sida, 4 Sider oxylon, 55 Smilacaceae, 19 Smilax, 1 Solanaceae, 63 Solanum, 63, iy, 84, 85 Sorgh st un Souroube Spathiphytum, irs Spermacoce, 72 Stachyt arpheta, 61, 62 4 Stemodia, Stenochl: aend, 10 CYRUS LONGWORTH LUNDELL Sterculiaceae, 43 Stigmaphyllon, 36 Stylosanthes, 34 Swartzia, 32 Syngonium, 17 Taber be aa cea 58 Talisia, 42 Tarro mae venado, 67 Tauchi, 57 ria, 10 Tephrosia, 34 Ternstroemia, 44 Tetracera, 43 Theophrastaceae, 55 Thevetia, 58 Thouinidium, 42 Tiliaceae Tillandsia, 18 Tinta, 30 Tournefortia, 60 7 rachypogon, 14 Trema, Trichachne, 14 Trichilia, 36 Turneraceae, 47 Tynanthus, 65 Ulmaceae, 23 Urosinine ra, 64 Urticace 24 Diricnlatie, 65 Verbenaceae, 60 74 Vinceto vicum, 59 Violaceac, 44 Vismia, 44 Vitaceae, 42 3 Vochysiaceae, 37 Wedelia, 74 Wigandia, 82 Wimmeria, 41 Ximenia, 25 INDEX Xiphidium, 19 Xylopia, 27 Xylosma, 75, 76 Xyridaceae, 17 Xyris, 17 Zamia, 11 Zanthoxylum, 35 Zephyranthes, 19 Zingiberaceae, 20 Zornia, Zosterella, 18 Zuelania, 45 No. CONTRIBUTIONS FROM THE UNIVERSITY OF bo eo = Cr io ~J wn MICHIGAN HERBARIUM . Rusts from British Honduras, by Edwin B. Mains. Notes on Agaries from British Honduras, by Alexander H. Smith. 28 pp., 2 pls. Price, $.25. (Issued ‘May 23, 1939.) Studies in the Genus Corhinartus I, by Alexander H. Smith. 42 pp., 12 pls. Price, $.35. (Issued June 30, 1939.) Revision of the Ametican Celastraceae I. Wi immeria, Micro- tropis, and Zinowiewia, by Cyrus Longworth landell: 46 pp., 10 pls. Price, $.40. (Issued Aug. 10, 1939, Studies of Tropical American Plants—I, by Cyrus Longworth Lundell. 32 pp. Price, $.25. (Issued April 1, 1940.) 73 pp., 32 pls. Price, $.50. (Issued Feb. 12, 1941. . Studies of North American Agaries—I, by Mexauten H. Smith. ) . Studies of American Sper matophytes—I, by C yrus Longworth 1¢ Lundell. 66 pp., 5 figs. Price, $.50. (Issued Sept. 1, Studies of American Spermatophytes—II, by Cyrus Longworth Lundell. 56 pp. Price, $.50. (Issued May 25, 1942. Flora of Eastern Tabasco and Adjacent Mexican Areas, by Cyrus Longw ah Lundell. Studies of American Spermatophytes—III, by Cyrus Long- worth Lundell. 95 pp. 4 pls., 8 figs., | map. Price, $1.00. mis eae : eee eee o pe reiete es see : se sit ne ie