THE FERN GAZETTE is a journal of the British Pteridological Society and contains peer-reviewed papers on all aspects of pteridology. Manuscripts may be submitted, and books etc. sent for review, to: Prof. M. Gibby, Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 SLR, UK Telephone: 0131-248-2973 E-mail: FernGazette@eBPS.org.uk fa _ Instructions for authors are on page 215 of this volume and also available at http://www.eB _ Copyright © 2008 British Pteridological Society. All rights reserved. No part of this _ publication may be reproduced in any material form (including photocopying or storing it in any medium by electronic means) without the permission of the British - Pteridological Society. _ THE FERN GAZETTE Volume 18 Part 3 was published on 27th August 2008 : _ Published by THE eas PTERIDOLOGICAL SOCIETY c/o Department of Bo ‘The Natural aoe Museum, Laoanrell Road, London SW7 5BD, UK ee "Printed by Geos Fic Cavin erbert pices siectelan, bine Portsmouth, PO6 IRU, UK ae oo co. beg : DEC 052008 = sp GARDEN LIBRARY AN ANNOTATED CHECKLIST OF THE MONILOPHYTES (FERNS) AND LYCOPHYTES OF EL SALVADOR FERN GAZ. 18(4):120-215. 2008 J. MONTERROSA! & A. K. MONRO? 'Herbario LAGU, Asociacion Jardin Botanico La Laguna, Urbanizacion Industrial Plan de La Laguna, Antiguo Cuscatlan, apartado postal 1197CG Email: jorg t jardinbotanico.org.sv *Botany Department, The Natural History Museum, London SW7 SBD, U.K. Email: a.monro@nhm.ac.u Key words: checklist, El Salvador, pteridophytes, monilophytes, lycophytes, ferns, Central America ABSTRACT We report a total of 397 species, 89 genera and 28 monilophyte and lycophyte familes for El Salvador. 29 of these are new records for the country and we have excluded 28 previous records. We summarise the extent of collecting within El Salvador and list the major collectors for the country. For each record, the local name, herbarium collections (verifiable biodiversity data), distribution within El Salvador and conservation status (where assessed) are given. This study is the result of a literature review and a review of 6397 herbarium specimens at BM, F, ITIC, LAGU, MHES and NY. RESUMEN Se reportan un total de 397 especies, 89 generos y 28 familias de monilophyte y lycophyte para El Salvador. 29 de las especies son nuevos registros para el pais y hemos excluido 28 registros anteriores. Se explican como el alcance de las colectas en El Salvador y se listan los principales colectores para el pais. Para cada registro, nombre comun, muestras de herabrio (datos de biodiversidad verificable), Distribucion dentro de El Salvador y estado de conservacion (donde se valora) son dados. Este estudio es el resultado de una revision de la literatura y 6397 muestras de herabrio de BM, F, ITIC, LAGU, MHES and NY. INTRODUCTION El Salvador is located on the Pacific coast of Central America and covers 21,000 km”. It has a population of 6.9 million (The World Bank Group, 2007), making it the most densely populated country in Latin America. It is also, by far, the most deforested country in continental Central and South America. Holdridge (1975) estimated that 99.5% of El Salvador would have been forested prior to human occupation. The vast majority of the country’s biodiversity including ferns, would have been associated with that forest cover. In 2001 a forest survey of the country based on sattelite data estimated undisturbed natural forest cover to be 7% (Ventura & Villacorta, 2001). The loss of so much natural forest suggests that a huge proportion of El Salvador’s fern diversity is at risk. Conservation tools such as checklists and identification guides are therefore of critical importance since they enable the accurate identification and monitoring of diversity. A fact recognised by El Salvador in the Estrategia Nacional de Biodiversidad 12] FERN GAZ. 18(4): 120-215. 2008 (Ministerio de Medio Ambiente y Recursos Naturales, 2000) and the Global Strategy for Plant Conservation (Secretariat of the Convention on Biological Diversity and Botanic Gardens Conservation International, 2002). Past studies of the Salvadoran Pteridophyte flora Ferns represent one of the most studied groups of organisms in El Salvador. The first study of Salvadoran ferns was undertaken in 1930 by Maxon & Standley, Ferns of the Republic of Salvador . This was based on the collections of Standley and listed 87 species for the country. In 1941 Calderén & Standley published the Lista Preliminar de Plantas de El Salvador (Preliminary list of the plants of El Salvador) which included 97 species. Thirteen years later Léetschert (1954), based at the ITIC herbarium of the University of El Salvador, published Nuevas Pteridofitas para El Salvador (New pteridophytes for El Salvador) in which 75 new records were included, bringing the total unmber of species to 172. In 1982, Ralph Seiler, a US Peace Corp worker based at the Museo de Historia Natural de El Salvador (MHES) from 1980 to 1982, published Contribuciones a la Pteridologia Centroamericana (Contributions to Central American Pteridology) in which he recognised 316 species for El Salvador. This was based on a combination of his own collections (denoted by a ‘*’ in Seiler, 1982) and those that he studied at ITIC. Towards the end of the 1980’s Pfeiffer-Berendsohn, published Listado Basico de la Flora Salvadorensis. Pteridophyta. Cuscatlania 1(1). This was based on the PLANTAS collections database of the research section of the Jardin Botanico La Laguna and listed 346 species for the country. This number was reduced with the publication of Flora Mesoamericana (Volume 1: Psilotaceae to Salviniaceae, Davidse et al.) in 1995. Flora Mesoamericana recorded 304 species for El Salvador although the collections at LAGU and ITIC were not consulted. MATERIALS AND METHODS The checklist is based on an inventory of the fern collections at BM, F, ITIC, LAGU, MHES and NY. All material was critically studied using the following reference works: Listado Basico de la Flora Salvadorensis. Pteridophyta. Cuscatlania 1(1) (Berendsohn, W. G., 1989): ‘The Pteridophytes of Mexico’ (Mickel & Smith, 2004); and Flora Mesoamericana Volume | (Davidse et al., 1995). In additions colleagues in Costa Rica and Mexico (Alexander Rojas Alvarado, Jardin Botanico de Lankaster, Costa Rica y Daniel Tejero-Diez, Universidad Nacional Autonoma de Mexico, Iztacala, Mexico) were contacted where there were doubts over the indentifion of a species. In total 6397 collections were examined: 1329 at BM, 931 at F, 523 at ITIC, 1523 at LAGU, 992 at MHES and 1099 at NY. Species were considered as being part of the Salvadoran flora where a herbarium collection of known Salvadoran origin was seen by the authors, or where a species was cited in one of the publications listed in Table 1 and referenced to a herbarium collection. Species records published elsewhere, or where cited in a publication in Table | but not referenced to a herbarium collection (or where the identification resulting in a citation was incorrect) were excluded. Of the species that met the above criteria for inclusion in the checklist some were excluded where a herbarium collection was based on cultivated material. Examples of other species that were excluded from the checklist are: Athvrium dombei Desv., a species described from Peru and whose citation for El Salvador is known only from Seiler (1982) but was not based on any herbarium material from E] Salvador; and Asplenium castaneum Schlecht. & Cham. (C.D. Adams, 1995) MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 122 whose citation was based on the misidentification of a Salvadoran collection (Seiler 160, F) of Asplenium polyphyllum Bertol. Where species were cited in one of the publications but no specimens were seen by these authors, this is indicated in the checklist by the symbol: ‘P’. Species were considered new records where that species, or its basionym / synonym had not been cited as occurring in El Salvador in a previous floristic publication encompassing the country (see Table 1). Such species are indicated in the checklist by the symbol ‘ ¢@’. The classification of ferns used in this study follows Smith et al. (2006). RESULTS A total of 397 species, 89 genera and 28 pteridophyte familes are reported for El Salvador (summarized in Table 3). This compares with previous estimates ranging from 87 to 346 (Maxon & Standley, 1930; Calderon & Standley, 1940; Léetschert, 1954; Seiler, 1982; Berendsohn, 1989; Davidse et al., 1995). The records are listed below. 25 of these are new records for the country and we have excluded 28 previous records. Only two species are endemic to El Salvador: Elaphoglossum microproductum A. Rojas (known only from the type collection) and Anemia salvadorensis Mickel. New records for El Salvador 29 new records were encountered during the course of this work. They are indicated by the symbol * #’ preceding the taxon name in Appendix 1. Asplenium cirrhatum Rich. ex Willd. Asplenium cristatum Lam Asplenium monodon Liebm. Asplenium theciferum (Kunth) Mett. Bolbitis bernoullii (Kuhn ex Christ) Ching Ctenitis equestris var. erosa Stolze Dennstaedtia globulifera (Poir.) Hieron. Dennstaedtia spinosa Mickel Diplazium drepanolobium A.R. Sm. Elaphoglossum lonchophyllum (Fée) T. Moore Elaphoglossum minutum (Pohl ex Fée) T. Moore Hymenophyllum crispum Kunt Lastreopsis effusa subsp. divergens ( Willd.) Tindale Megalastrum pulverulentum (Poir.) A. R. Sm. & R. C. Moran Olfersia cervina (L.) Kunze Pityrogramma ochracea (C. Pres|) Domin Polypodium diplotrichum Mickel & Beitel Polypodium thyssanolepis A. Braun ex Klotzsch Polystichum muenchii (Christ) C. Chr. Psilotum nudum (L.) P. Beauv. Pteris erosa Mickel & Beitel Terpsichore delicatula (M. Martens & Galeotti) A. R. Sm. Thelypteris angustifolia (Willd.) Proctor Thelypteris linkiana (C. Presl) R.M. Tryon Thelypteris obliterata (Sw.) Proctor Thelypteris serrata (Cav.) Alston 123 FERN GAZ. 18(4): 120-215. 2008 Thelypteris standleyi (Maxon & C. V. Morton) R. M. Tryon Trichomanes poids gga Bosch Vittaria bradeorum Rosenst Species excluded from the checklist Asplenium auritum Sw. var. auritum The only record of this species (Seiler, 1982) is an incorrectly identified collection of Asplenium monodon Liebm. (Seiler 466, MUHNES). Asplenium castaneum Schlecht. & Cham. The only record of this species (Adams, 1995), is an incorrectly identified collection of Asplenium polyphyllum Bertol. (Seiler 160, F). Asplenium hastatum Klotzsch ex Kunze Seiler (1982) referred to the presence of this species in El Salvador, but referred to its resence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Asplenium otites Link Seiler (1982) reported this species for El Salvador. No specimens or additional citations associating this species to El Salvador were encountered and reports of this species rom Mesoamerica as a whole are uncertain (Adams, 1995; Mickel & Smith, 2004). Asplenium radicans L. Seiler (1982) reported Asplenium radicans var. partitum for El Salvador. This variety has since been placed in synonymy with 4. flabellulatum. (Adams, 1995). Asplenium rigidum Sw. This species was reported from El Salvador (Adams, 1995) based on misidentified material (see Mickel & Smith 2004: 90, under discussion for Asplenium cuspidatum Lam.). No collections of this species from El Salvador have been located. Athyrium dombei Desv. This species was reported from El Salvador (Adams, 1995) based on misidentified material (see Mickel & Smith 2004: 137 under discussion for Athyrium bourgeaui E. Fourn.). No collections of this species from El Salvador have been located. Dennstaedtia dissecta (Sw.) T. Moore This species is restricted to the Greater Antilles, Costa Rica, Panama and the Andes (Navarrete & Ollgaard, 2000). It is likely that Seiler (1982) mistakenly identified material of Dennstaedtia cornuta (Kaulf.) Mett., which was previously treated as conspecific to D. dissecta (see discussion of D. cornuta in Mickel & Smith, 2004). Elaphoglossum firmum (Mett.) Urb Seiler (1982) referred to the presence of this species in El Salvador as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Elaphoglossum formosum (M. Martens & Galeotti) Urb. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Elaphoglossum longifolium (Jacq.) J. Sm. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Elaphoglossum pilosum (Humb. & Bonpl. ex Willd) T. Moore MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 124 Seiler (1982) referred to the presence of this species in El Salvador as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Elaphoglossum revolvens (Kunze) C. Chr. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Elaphoglossum rigidum (Aubl|.) Urban Although Stolze (1981) reported this species (as a synomym of E. flaccidum) from El Salvador, Guatemala and Mexico, Mickel & Smith 2004 do not recognize this species from Mexico and Mickel in Davidse ef al. (1995) treats E. flaccidum as native to South America. Grammitis heteromorpha (Hook. & Grev.) C. V. Morton Seiler (1982) referred to the presence of this species in El Salvador as dubious because he did not see a collection from the country. No collections of this species were located by Monterrosa & Monro. In addition, Mickel & Smith (2004) state that G heteromorpha is restricted to the Andes. Grammitis meridensis (Klotzsch) F. Seym. Seiler (1982) referred to the presence of this species in El Salvador, but referred to its presence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Huperzia mexicana (Herter) Holub Seiler (1982) referred to the presence of this species in El Salvador, but referred to its presence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Huperzia verticillata (L.f) Trevis. Seiler (1982) referred to the presence of this species in El Salvador, but referred to its resence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Lellingeria delitescens (Maxon) A. R. Sm. & R. C. Moran Seiler (1982) referred to the presence of this species in El Salvador, but referred to its presence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Notholaena affinis (Mett.) Hook. ex T. Moore The only record of this species (Seiler 88, MHES, NY; Seiler, 1982) is an incorrectly identified collection of Notholaena meridionalis Mickel. Polypodium arcanum Maxon Seiler (1982) referred to the presence of this species in El Salvador, but referred to its presence as dubious because he had not seen a collection from the country. No collections of this species were located by Monterrosa & Monro. Polypodium montigenum Maxon Seiler (1982) referred to the presence of this species in El Salvador but did not cite a collection. No collections of this species were located by Monterrosa & Monro and it is possible that Seiler incorrectly identified a collection of Polypodium alansmithii R. C. Moran as P. montigenum. Polypodium ptilorrhizon Christ This species was reported in El Salvador by Seiler (1982) and Berendsohn (1989) and is based on a misidentification by Seiler (1982). 125 FERN GAZ. 18(4): 120-215. 2008 Polystichum ordinatum (Kunze) Liebm. Maxon & Standley (1930) and later Calderon & Standley (1941) reported this species from El Salvador. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he did not see a collection from the country. No collections of this species were located by Monterrosa & Monro. Pteris pulchra Schitdl. & Cham. Maxon & Standley (1930) reported this species as present in El Salvador. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he did not see a collection from the country. No collections of this species were located by Monterrosa & Monro. Pteris polita Link. Seiler (1982) cites Preris polita as present in El Salvador based on the misidentification of material of Preris orizabae. Thelypteris opposita (Vahl) Ching Seiler (1982) cites Thelypteris opposita as present in El Salvador on the misidentification of material of material of Thelypteris balbisii (Spreng.) Ching and 7: resinifera (Desv.) Proctor. Vittaria lineata (L.) J. Sm. Calderon & Standley (1941) reported this species as present in El Salvador. Seiler (1982) referred to the presence of this species in El Salvador as dubious because he did not see a collection from the country. No collections of this species were located by Monterrosa & Monro. Uses Marattia and Equisetum have documented (Pers. Obs.) medicinal uses: Marattia excavate as a contraceptive; Equisetum giganteum and E. myriochaetum as diuretics and hypoglycaemics and Equisetum species “cola de caballo” are commonly sold in local markets for these properties (Pers. Obs.). Polypodium pleurosorum “calaguala” is also sold as a medicinal plant, however the purported properties are unknown. Cyathea divergens, Nephrolepis spp. “cola de ardilla” and Polypodium pleurosorum are used as ornamentals in gardens and landscaping and the stems of Cyathea species are sold as a growing medium for the cultivation of orchids. No other uses are known for Salvadoran ferns. Table 1. State-wise distribution of fern species records based on collections at LAGU. State # State records State # State records Ahuachapan 14] San Miguel 34 Cabanas 24 San Salvador 107 |Chalatenango 142 San Vicente 46 Cuscatlan 10 Santa Ana 236 La Libertad 71 Sonsonate 87 La Paz 12 Usulutan 47 Morazan 122 Total st MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 126 Also of economic significance is the invasive species Preridium aquilinum var. feei (bracken) that has been responsible for the poisoning of cattle throughout the country (Ministerio de Agricultura y Ganaderia, Pers. Comm., 2005). Locality distribution data within El Salvador Species records for a locality are the product of species distribution and sampling effort, with differences in the number of species records from one state to the next not necessarily reflecting differences in fern diversity or sampling effort. Combined with knowledge of the geography and vegetation of a country and collecting effort as indicated by herbarium records, however, they can be used to give an indication of both variation in sampling effort and diversity. Analysis of the 1523 collections held at LAGU indicates that sampling effort has been very uneven in El Salvador - Santa Ana, Ahuachapan, Chalatenango and Morazan accounting for ca 75% of species-records whilst Cuscatlan and La Paz account for just 1.7% of records. In part this correlates with the expected distribution of fern diversity based on altitudinal gradients in these states. The unevenness of the records is, however, much greater than would be expected from altitudinal gradients and it appears that there has been collecting bias. Most likely collection effort has been focused on protected areas: the cloud forest of reserve of Montecristo (Santa Ana), the lowland tropical moist forest reserve of El Impossible (Ahuachapan), El Pital (Chalatenango) and the Rio Sapo oak and pine forest reserve (Morazan). This also corresponds with the authors’ own knowledge of collecting effort. A future collection program focussing on the states with the fewest species records (Cuscatlan, La Paz, Cabafias, San Vicente and San Miguel) and agricultural (e.g. shade coffee farms) and private land would be expected to provide valuable baseline data essential for the conservation of ferns in El Salvador. Principal collectors and collections The principle collectors of ferns in El Salvador have been William Létscher (also known as Loetschert, first set of collections at ITIC), William Maxon (first set of collections at US), Paul Standley (main sets of collections at US and F), Calderon (main sets of collections at US), John Tucker (first set of collections at DAV), Antonio Molina R. (first set of collections at EAP), Norman Fasset (first set of collections at WIS), Louis Williams (first set of collections at F) and Margery Carlson (first set of collections at F) and Ralph Seiler (first set of collections at MHES and F) DISCUSSION El Salvador’s fern diversity in a national context Ferns are one the best studied groups of organisms in El Salvador, alongside birds and reptiles. The first checklist to the ferns of the country dates to the 1920s and several have followed (see introduction). This comparitively strong knowledge of the fern flora of El Salvador makes ferns a good candidate for developing national inventory protocols and conservation strategies that will contribute to El] Salvador’s commitment to the Global Strategy for Plant Conservation (2002). A Darwin Initiative project initiated in 2006 with the Ministerio de Medio Ambiente y Recursos Naturales (Ministry of the Environment and Natural Resources), ‘Tools and training for fern conservation and monitoring (El Salvador)’, seeks to do exactly this and use ferns to develop a country-wide monitoring strategy that can subsequently be applied to other groups of organisms. 127 FERN GAZ. 18(4): 120-215. 2008 Fern species-records for Salvadoran ferns are very unevenly distributed between states, indicating an apparent sampling bias towards the country’s national parks and against private land and ‘disturbed’ vegetation (secondary forest, agroforest, small forest fragments). The relatively poorly sampled disturbed vegetation represents the greater part of the El Salvador’s forest cover (19% versus 7%) and further sampling in this areas should generate many new species-records at the state and possibly national level. As an example of this potential, an inventory of the ferns growing within 27 shade coffee farms enumerated 38 fern species (Monro et al., 2002) and accounts for a significant proportion of the fern records for the state of Cabajias. It should therefore be seen as a priority to sample such areas. The economic importance of ferns to El Salvador although yet to be evaluated, is likely to be minimal and restricted to medicinal and ornamental uses and the toxicity of the invasive Pteridium aquilinum. El Salvador’s fern diversity in a regional context El Salvador has relatively high species diversity per km° compared with the neighbouring countries of Central America. This is possibly a sampling artifact, El Salvador having been more intensively collected for ferns than other countries in Central America. Although the most active collectors of ferns in El Salvador expended greater collecting effort in Guatemala, Guatemala has a greater area - over five times the size of El Salvador. El Salvador’s high fern diversity could also be ascribed to its large altitudinal variation (2418 m) and variation in rainfall across the country (1100- 1700 mm pa), for its size. The transition from the dry coastal lowlands to the wetter highlands inland, combined with the presence of a number of isolated volcanic peaks that reach altitudes greater than 1700 m (Santa Ana, Coatepeque, Apaneca, Izalco, San Salvador, Ilopango, San Miguel, San Vicente, Tecapa and Conchagua) has resulted in a great diversity of forest and non-forest habitat types (Ventura et al., 2001) at altitudes associated with high fern diversity elsewhere (Kluge et al., 2006), many of which are isolated from each-other. Kluge et al. (2006), studying fern diversity along an altitudinal gradient in Costa Rica found that fern diversity peaked at mid-elevations (ca 1700 m) and that the mid-domain position together with humidity and temperature were the most powerful explanatory variables species diversity. It may be that a combination of the relative frequency of mid-domain positions in voleanoe-dominated landscapes is high and that this, combined with a relatively steep rainfall gradient across the landscapes contributes to the relatively high diversity of ferns in El Slavador. As is the case for Neotropical ferns in general the level of endemism is very low at ca 0.5% of the species. Table 2. Current estimates of fern diversity for Central America. Country Fern diversity (source) esas 5 ee Po area km |range (m) |1000 km Guatemala 652 (Stolze, R. G., 1976, 1981, 1983)| 108,900 4220 6.0 Belize 234 (Balick et al. 2000) 22,960 1124 11.9 El] Salvador |397 (this article) 21,000 2418 18.9 Costa Rica /1112 (Hammel et al., 2004) 51,100 3810 Pa ie Panama 938 (Correa et al., 2004) 13,320 3475 12.4 MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 128 BIBLIOGRAPHY BALICK, M.J.,.NEE, M. H. & ATHA, D.E.. 2000. Checklist of the Vascular Plants of Belize. 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New taxa, new records and redefined concepts in the — sect. Elaphoglossum subsec. Pachyglossa (Lomariopsidaceae) from Mexico and Central America. Revista Biol. Trop. 51(1): 1-32. ROJAS- ALY ARADO, A. F., 2003. Notes on Elaphoglossum (Lomariopsidaceae) section Polytrichia subsection Hybrida in Mexico and Central America. Revista Biol. Trop. 51(1): 33- SECRETARIAT OF si CONVENTION ON BIOLOGICAL DIVERSITY AND BOTANIC GARDENS CONSERVATION INTERNATIONAL, 2002. Global strategy for plant conservation. Montreal: Secretariat of the Convention on Biological Diversity. Botanic Gardens Conservation International, Richmond . SEILER, R. L., 1982. Contribuciones a la Pteridologia Centroamericana. Enumeratio Filicum Sancti-Salvatoris. Brenesia 19/20: 381-391. SMITH, A. R., 1981. Pteridophytes. In Breedlove, D. E. (Ed.) Flora of Chiapas. 2: 1- 370. California Academy of Sciences, San Francisco SMITH, A. R., PRYER, K. M., SCHUETTPELZ, E., KORALL, P., SCHNEIDER, H. & WOLF, P. G., 2006. A Classification for Extant Ferns. Taxon 55(3): 705-731. SMITH, A. R., KREIER, H., HAUFLER, C. H., RANKER, T. A. & SCHNEIDER, H. 2006. Serpocaulon (Polypodiaceae), a new genus segregated from Polypodium. Taxon 55(4): 919-930. SIOLZE. Kk, G., oe Fern and Ferns Allies of Guatemala. Part I. Ophioglossaceae through Cyatheaceae. Fieldiana Bot. 39: i-v, 1-130. STOLZE, R. G, 1981. Fern and Ferns Allies of Guatemala. Part II. Polypodiaceae. Fieldiana Bot. n. s. 6: 1-522. STOLZE, R. G, 1983. Fern and Ferns Allies of Guatemala. Part III. Marsileaceae, Salviniaceae, and the ferns allies (including a comprehensive index to Parts I, II, and III). Fieldiana Bot. n.s. 12:1-91. TEJERO-DIEZ, J. D. & PACHECO, L., 2004. Notas Taxonémicas y de Distribucion en Polypodium plesiosorum y P. rhodopleuron (Polypodiaceae: Pteridophyta). Anales Inst. Biol. Univ. Nac. Auton. Mexico, Bot. 75(1): 11-37. TEJERO-DIEZ, J. D. & PACHECO, L., 2004. Taxa Nuevos, Nomenclatura, Redefinicion y Distribucion de las Especies Relacionadas con Polypodium colpodes Kunze (Polypodiaceae, Pteridophyta). Acta Bot. Mex. 67: 75-115. THE WORLD BANK GROUP, 2007. El Salvador Data Profile. http://devdata.worldbank.org/external/CPProfile. asp?SelectedCountry=SLV&CCO DE=SLV&CNAME=EI+Salvador&PTY PE=CP VENTURA, N. E., VILLACORTA, R. F., 2001. Mapeo de Vegetacion Natural de Ecosistemas Terrestres y Acuaticos de El Salvador (CD Rom). San Salvador: Ministerio de Medio Ambiente y Recursos Naturales. ACKNOWLEDGEMENTS We would like to thanks the curators at BM, F, ITIC, LAGU, MHES and NY for providing access to the collections. In particular we would like to thank Alison Paul (BM) for help with the identification of some of the material at BM, Robbin Moran and John Mickel (NY) for their hospitality and kind help in the identification of material whilst at NY, José Daniel Tejero-Diez of the Faculty of Iztacala (UNAM) for the revision of the checklist and help with the nomenclature of a number of Polypodium MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 130 names, Alan Smith (UC) for his kind help in the identification of material of Thelypteris, David Barrington (VT) for his kind help in the identification of material of Polystichum and Alexander Rojas Alvarado (Lankester Botanic Garden) for his help with the identifaction of Cyathea and Elaphoglossum collections. We would also like to thank the Association La Laguna Botanic Garden for permission to undertake this research and access to the PLANTAS database and Gerrit Davidse and Jim Solomon at MO for provding a dump of Salvadoran fern records in TROPICOS. Jorge Monterrosa benefited from a Darwin Scholar Programme Award (No. EIDPS7), was able to visit F as part of Darwin Initiative project 640/15/015 and was able to visit NY as part of the Norwegian Agency for Development Cooperation project “Desarrollando Capacidades y Compartiendo Tecnologia para la Gestion de la Biodiversidad en Centroamérica” through The Natural History Museum of El Salvador (MHES). We would also like to thank Harald Schneider and Bob Press (BM) for their comments on earlier drafts of this manuscript. APPENDIX 1. CHECKLIST OF THE MONILOPHYTES (FERNS) AND LYCOPHYTES OF EL SALVADOR Ferns or monilophytes are arranged by family in alphabetical order following Smith et al. (2006) asignment of genera to family. Lycophytes are arranged by family in alphabetical order following the asignment of genera to family of Flora Mesoamericana (Davidse et al., 1995). Conspectus of the monilophytes and lycophytes of El Salvador (see pages 131-132) New records for El Salvador are indicated by “#”. Endemic species are indicated by “@”. Species for which no collections have been seen but where we are confident of their inclusion by previous authors (see Table 1) are indicated by “>”. DIVISION LYCOPHYTA LYCOPODIACEAE Huperzia cuernavacensis (Underw. & F. E. Lloyd) Holub, Folia Geobot. Phytotax. 20: toe 1982. Lycopodium cuernavacensis Underw. & F. E. Lloyd, Urostachys cuernavacensis (Underw. & F. E. Lloyd) Hetter ex Nessel Global distribution: Mexico, Guatemala. Salvadoran distribution: Chalatenango: R. Seiler 157 (MHES). Huperzia dichotoma (Jacq.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874. Lycopodium dichotomum Jacq., Urostachys dichotomus (Jacq.) Herter . Local name: candelillo. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, French Guyana, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Ahuachapan: Arana s.n. (JBL04315) (LAGU); R. Villacorta et al. 908 (B, LAGU, MO); W. Berendsohn & M. Sandoval 1634 (LAGU). Chalatenango: H. Sipman ef al. s.n. (JBL02486) (LAGU). 131 FERN GAZ. 18(4): 120-215. 2008 TABLE 3. Conspectus of the monilophytes and lycophytes of El Salvador. N Division Class Order Family Genus p p. Huperzia 11 DIVISION vous Lycopodiales Lycopodiaceae Lycopodiella 3 LYCOPHYTA copodium Z Selaginellales Selaginellaceae _|Selaginella 13 PSILOTOPSIDA Ophioglosales |Ophioglosaceae on ; Psilotales Psilotaceae Psilotum z EQUISETOPSIDA |Equisetales |Equisetaceae Equisetum 4 MARATTIOPSIDA |Marattiales Marattiaceae Marattia 2 Osmundales Osmundaceae Osmunda I - oy = Based |Hymenophyllum| 9 : si pili Trichomanes 9 |Dicranopteris Bl Gleicheniales Gleicheniaceae a a Seo Ea Gleichenella | Sticherus 4 ee Lygodiaceae Lygodium | Anemiaceae Anemia 6 Marsileaceae Marsilea * Salviniales ae Azolla 2 Salviniaceae : zz Salvinia 2 Culcitaceae Culcita 1 DIVISION Cibotiaceae Cibotium im! EUPHYLLOPHYTA Alsophila | 3 SUB-DIVISION Cyatheales Cyatheaceae Cyathea a MONILOPHYTA Sphaeropteris | | POLY PODIOPSIDA Dicksoniaceae a ©) ophosoria a ; Lindsaea l Lindsaeaceae pores i Dennstaedtia E Dennstaedtiaceae a Me x Hypolepis | 4 Pteridium 3 |Acrostichum 2 Adiantum 18) Polypodiales Ananthacorus l Anogramma 2 Bommeria l Pteridaceae Ceratopteris 2 Cheilanthes 11 Cheiloplecton l Doryopteris 1] Eriosorus l Hemionitis | 3 MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 132 TABLE 3. (continued from previous page) Division Class Order Family DIVISION EUPHYLLOPHYTA SUB-DIVISION MONILOPHYTA POLY PUDIVUT SIDA Polypodiales Pteridaceae Pityrogramma Polytaenium Pteris Scoliosorus Aspleniaceae ; a Thelypteridaceae Th Woodsiaceae — Blechnaceae — Dryopteridaceae nr] CO] Wi | NO] ii | SO] | Go i) Ks) Phanerophlebia Polystichum 1 j nds Cyclopeltis r Nephrolepis Tectariaceae Tectaria Polypodiaceae |Micropolypodium Terpsichore ampyloneurum ho Serpocaulon Nd] Bi —| rol | KL AL NJ ee EY Be] BO] MO] Gy BY] |] OOP NM |] ef Total (vs) ‘oO — 133 FERN GAZ. 18(4): 120-215. 2008 Huperzia hippuridea (Christ) Holub, Folia Geobot. Phytotax. 20: 73. 1985. Lycopodium hippurideum Christ in Pittier, Urostachys hippurideus (Christ) Herter ex essel Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: R. Villacorta et al. 1284 (B, LAGU, MO). Chalatenango: R. Villacorta et al. 2052 (B, LAGU, MO); R. Seiler 158 (F, MHES). Huperzia linifolia (L.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874. Lycopodium linifolium L., Urostachys linifolius (L.) Herter Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Suriname, Ecuador, Brazil, Bolivia. Salvadoran distribution: Santa Ana: H. Weber 277 (ITIC); R. Seiler 247 (MHES); S. Winkler 8b (ITIC). Huperzia myrsinites (Lam.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874. Lycopodium myrsinites Lam., L. skutchii Maxon, Plananthus myrsinites (Lam.) BP; Beauv., Urostachys myrsinites (Lam.) Herter Global distribution: Mexico, Guatemala, Honduras, ee Costa Rica, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Ecuador, Salvadoran distribution: Santa Ana: R. Seiler 235 (F, ITIC, ores 288 (MHES); R. Villacorta et al. 1278, 1280 (B, LAGU, MO). Chalatenango: R. Villacorta et al. 2057 (B, LAGU, MO); R. Seiler 401 (JBL00110) (F, LAGU, MHES); 747 (F); S. Winkler la (ITIC). Morazan: R. Seiler 1021 (F, MHES). Huperzia orizabae (Underw. & F. E. Lloyd) Holub, Folia Geobot. Phytotax. 20: 75. 1985. Lycopodium orizabae Underw. & F. E. Lloyd, Urostachys orizabae (Underw. & F. E. Lloyd) Herter ex Nessel Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Kovar 2594, US (Ollgaard in Davidse et al. 1995). Santa Ana: R. Seiler 764 (MHES) Huperzia pithyoides (Schidtl. & Cham.) Holub, Folia Geobot. Phytotax. 20: 76. 1985. Lycopodium pithyoides Schltdl. & Cham., L. gigas Herter, Urostachys pithyvoides (Schltdl. & Cham.) Herter, U. gigas (Herter) Herter Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela. Salvadoran distribution: Santa Ana: R.A. Carballo et al. 90 (B, LAGU); R. Seiler 310 (MHES); S. Winkler 6b (ITIC). Chalatenango: R. Seiler 420 (JBLO0111) (LAGU); R. Seiler 746 (MHES). Morazan: R. Seiler 982 (F, ITIC, MHES), Huperzia pringlei (Underw. et F. E. Lloyd) Holub, Folia Geobot. Phytotax. 20: 76. 1985. Lycopodium pringlei Underw. & F. E. Lloyd, L. tuerckheimii Maxon, Urostachvs pringlei (Underw. & F. E. Lloyd) Herter ex Nessel, U. tuerckheimii (Maxon) Herter, Huperzia tuerckheimii (Maxon) Holub MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 134 Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: H. Weber 294 (ITIC); J. Lagos 12 (ITIC): R. Seiler 180, 767 (MHES); R. Villacorta & H. Sipman 1275 (B, LAGU, MO); S. Winkler 3 (ITIC). Chalatenango: R. Seiler 159 (ITIC, MHES): 743 (MHES). Morazan: E.A. Montalvo s.n. (ITIC); R. Seiler 1001 (F, MHES): 1022 (MHES). Note: Seiler (1982) and Berendsohn (1989) treat L. pringlei and L. tuerckheimii as distinct species. Huperzia reflexa (Lam.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874. Lycopodium reflexum Lam., Urostachys reflexus (Lam.) Herter Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Sonsonate: R. Villacorta & M.L. Reyna de Aguilar 716 (B, ITIC, LAGU, MO). Santa Ana: A. Molina ef al. 21712 (F, ITIC); M.C. Carlson 979 (F); R. Seiler 267 (F, MHES); 883 (F, ITIC, MHES); S. Winkler 7a (ITIC). La Libertad: M.C. Carlson 249 (F): R. Villacorta & E.A. Montalvo 2314 (B, LAGU, MO). San Salvador: A. Molina et a/. 21862 (F). San Vicente: E.A. Montalvo s.n. (ITIC). Morazan: R. Seiler 1013, 1085 (F, MHES); 1052 (MHES); 1077 (F, ITIC, MHES). Huperzia taxifolia (Sw.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248. 1874. Lycopodium taxifolium Sw., Urostachys taxifolius (Sw.) Herter Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia. Salvadoran distribution: Ahuachapan: A. Molina et a/. 21760 (ITIC); E.A. Montalvo 5463 (ITIC); R. Seiler 959 (F, MHES). Santa Ana: G. Davidse er a/. 37244 (B, BM, ITIC, LAGU, MO, L). Gonzalez 1981 (ITIC); R. Seiler 307 (F, MHES); S. Winkler 4 (ITIC). San Vicente: J. Lagos 2381 (ITIC). Huperzia watsoniana (Maxon) B. Ollg., Ann. Missouri Bot. Gard. 79: 701. 1992. Lycopodium watsonianum Maxon Global distribution: Costa Rica, Panama. Salvadoran distribution: Molina et al. 16823, F (QOllgaard in Davidse ef a/. 1995). Santa Ana: R. Seiler 232 (MHES): 289 (F, MHES). Chalatenango: R. Seiler 744 (MHES) Lycopodiella alopecuroides (L.) Cranfill, Amer. Fern J. 71: 97. 1981. Lycopodium alopecuroides Global distribution. USA, Mexico, Guatemala, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Urugua Salvadoran distribution: Morazan: R. Seiler 1078 (F, ITIC, MHES). Lycopodiella caroliniana (L.) Pic. Serm., Webbia 23: 165. 1968. Lycopodium carolinianum L., L. meridionale Underw. & F. E. Lloyd, L. carolinianum L. var. mer —— (Underw. & F. E. Lloyd) Nessel, seme s meridionalis (Underw F. E. Lloyd) Holub, Lycopodiella caroliniana (L.) Pic. Serm. var. es (Underw. & F. E. Lloyd) B. Ollg. & P. G. Windisch. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Panama, 135 FERN GAZ. 18(4): 120-215. 2008 Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru. Salvadoran distribution. Morazan: R. Seiler 1014 (F, MHES). Lycopodiella cernua (L.) Pic. Serm., Webbia 23: 166. 1968. Lycopodium cernuum L., Lepidotis cernua (L.) P. Beauv., Palhinhaea cernua (L.) Vasc. & Franco. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Tropicos y Subtropicos del Viejo Mundo. Salvadoran distribution: Santa Ana: B. Bechyne 3908 (ITIC); E. Pefiate s.n. (ITIC); R. Seiler 39 (F, MHES); 1204 (F); Thomson 55 (ITIC). San Salvador: N.C. Fassett 28601 (ITIC); W. Eggler 682 (F). Chalatenango: A. Diaz s.n. (ITIC); M.C. Carlson 592 (F); R. Villacorta & A. Araniva 980 (LAGU, MO, B, ITIC). Usulutan: J. Lagos 2 (ITIC). Morazan: R.A. Carballo et al. 1032 (LAGU, MO); R. Seiler 978 (F, ITIC, MHES); 1012, 1043 (F, MHES). Lycopodium clavatum L., Sp. Pl. 2: 1101. 1753. Lycopodium aristatum Humb. & Bonpl. ex Willd., L. piliferum Raddi, L. trichophyllum Desv., L. aristatum Humb. & Bonpl. ex Willd. var. incurvum Grev. & Hook., L. aristatum Humb. & Bonpl. ex Willd. var. robustius Grev. & Hook., L. clavatum L. vat. raddianum Spring, L. clavatum L. var. aristatum (Humb. & Bonpl. ex Willd.) Spring, L. clavatum L. var. desvauxianum Spring, L. clavatum L. var. jamaicense Spring, L. aristatum Humb. & Bonpl. ex Willd. var. desvauxianum Spring, L. trichiatum Bory vat. desvauxianum Spring, L. eriostachys Fée, L. clavatum L. var. eriostachys (Fée) Nessel & Hoehne, L. clavatum L. var. piliferum (Raddi) Nessel & Hoehne, L. clavatum L. vat. equisetoides Schwacke ex Nessel & Hoehne, L. clavatum L. var. trichophyllum (Desv.) Nessel & Hoehne. Local name: guia de roble. Global distribution. USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Europe. Salvadoran distribution: Santa Ana: A. Molina et al. 12538, 16715 (F; B). Pfeiffer- Berendsohn 21 (B, LAGU, MO); E.A. Montalvo s.n., 3128, 4136 (ITIC); J. Lagos, 8, 1480 (ITIC); M.L. Reyna de Aguilar 1029 (MHES); N.C. Fassett 28310 (F, ITIC); R. Seiler 115 (F, ITIC, MHES); 769 (F, MHES); R. Villacorta er al. 1046 (B, LAGU, MO); W. Berendsohn et al. 1720 (B, BM, LAGU, MO). Chalatenango: J. Tucker 1051 (F); R. Villacorta et al. 2077 (B, LAGU, MO); W. Berendsohn et al. 1565 (B, LAGU, MO). Lycopodium thyoides Humb. & Bonpl. ex Willd., Sp. Pl., ed. 4, 5(1): 18. 1810. Diphasiastrum thyoides (Humb. & Bonpl. ex Willd.) Holub, Lycopodium complanatum L. var. validum Weath., L. complanatum L. var. adpressifolium Spring, L. complanatum L. var. thyoides (Humb. & Bonpl. ex Willd.) Christ. L. complanatum L. var. tropicum Spring . Local name: helecho guia de roble. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia. Venezuela, Ecuador, Peru, Brazil, Bolivia, Argentina. Salvadoran distribution: Padilla 170 (US) (Ollgaard in Davidse ef al. 1995). Chalatenango: W. Berendsohn et a/. 1567 (B, LAGU, MO). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 136 SELAGINELLACEAE Selaginella acutifolia (Stolze) Valdespino, Mem. New York Bot. Gard. 88: 558. 2004. Selaginella pallescens (C. Presl) Spring var. acutifolia Stolze Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 635 (B, LAGU): J.M. Rosales 1062, 1169 (AAU, B, BM, LAGU, MO). Sonsonate: D. Rodriguez & M. Trejo 77 (B, LAGU, MO). La Libertad: K. Sidwell et a/. 509 (B, BM, ITIC, LAGU, MO); M. Renderos & R. Villacorta 286 (B, EAP, LAGU, MO); R.A. Carballo ef al. 48 (B, LAGU, MO). Chalatenango: R. Seiler 365, 549 (MHES). La Paz: J. Lagos 4 (ITIC). San Miguel: R. Seiler 541 (MHES); R. Villacorta et al. 2753 (B, BM, EAP, LAGU, MO). Cabafias: R.A. Carballo et a/.289 (B, LAGU, MO). Morazan: R.A. Carballo et al. 1067 (LAGU). Selaginella bernoullii Hieron., Hedwigia 41: 192. 1902. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Chalatenango: R. Seiler 733 (F, MHES). Note: M.E. Fraile in Dvidse et a/. (1995) cites this name as a synonym of S. porphyrospora A. Braun. We have chosen to maintain the species distinct following the criteria cited by Mickel & Smith (2004). > Selaginella convoluta (Arn.) Spring in Martius, Fl. Bras. 1(2): 131. 1840. Lycopodium convolutum Arn., Selaginella longispicata Underw. & Millsp. Global distribution: Mexico, Guatemala, Honduras, Antilles, Brazil, Bolivia, Paraguay, Argentina. Source: Léetschert (1954): 23; Seiler (1982): 382; Berendsohn (1989) 1: 24. > Selaginella estrellensis Hieron., Hedwigia 41. 200. 1902. Global distribution: Costa Rica. Source: Fraile in Davidse et al. (1995): 32; Alston 1955 (no visto). Selaginella hoffmannii Hieron., Hedwigia 41. 184. 1902. Local name: yayuco colorado. Global distribution: Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 666 (B, LAGU); J. Lagos 5 (ITIC); J.M. Rosales 1231, 1232 (B, BM, LAGU, MO); 1257 (AAU, B, BM, LAGU, MO); 1300 (B, BM, EAP, LAGU, MO); 1619 (B, LAGU); R. Villacorta e7 al. 905 (B, LAGU, MO). Santa Ana: A.K. Monro & R. Douglas 3462 (B, BM, ITIC, LAGU, MO); R. Seiler 705 (F, MHES); R. Villacorta 2179 (B, LAGU, MO). La Libertad: A.K. Monro ef al. 3082 (B, BM, ITIC, LAGU, MO); R.A. Carballo ef a/. 40 (B, LAGU, MO). San Salvador: Grupo Ecologico Canton El Zapote s.n. (JBL03316) (LAGU, MO); R. Seiler 634 (F, MHES); 1509 (F). Chalatenango: R. Seiler 143 (F, ITIC, MHES). Selaginella lepidophylla (Hook. & Grev.) Spring in Martius, Fl. Bras. 1(2): 126. 1840. Lycopodium lepidophyllum Hook. & Grev. Global distribution: USA, Mexico. Salvadoran distribution: San Salvador: R. Seiler 48 (F, ITIC, MHES). 137 FERN GAZ. 18(4): 120-215. 2008 Selaginella martensii Spring, Acad. Roy. Sci. Belgique 24: 129. 1849. Selaginella solmsii Baker Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution. Santa Ana: A. Molina et al. 12633, 16866, 16883 (F); E.A. Montalvo 4137 (ITIC); G. Davidse et al. 37242, 37304 (B, BM, ITIC, LAGU, MO); J. Lagos 14 (ITIC); J. Tablas s.n. (ITIC); J. Toledo s.n. (ITIC); M.C. Carlson 890 (F); K. Seiler 108 (F, ITIC); 184 (F, MHES); 472 (F, ITIC, MHES); R. Villacorta ef al. 1279 (B, LAGU); 2679 (B, F, LAGU, MO). La Libertad: F. Lopez s.n. (ITIC). Selaginella pallescens (C. Presl) Spring in Martius, Fl. Bras. 1(2): 132. 1840. Lycopodium pallescens C. Presl, L. cuspidatum Link, Selaginella cuspidata (Link) Link, S. cuspidata (Link) Link var. elongata Spring Local names: helecho tuya, resurreccion Global distribution: Mexico, Guatenata, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Suriname, Brazil. Salvadoran distribution: Ahuachapan: E. Sandoval et a/. 602 (B, LAGU); 1338 (K, LAGU); J.M. Rosales 1299 (B, BM, EAP, LAGU, MO); 1403, 1586 (AAU, B, BM, LAGU, MO); M. Sandoval & E. Sandoval 20 (B, F, LAGU); T. Croat 42064 (ITIC). Sonsonate: R. Seiler 8 (F, MHES); T. Croat 42239 (ITIC). Santa Ana: A. Molina ef al. 21881 (F, ITIC); T. Croat 42300 (ITIC); V.M. Martinez s.n. (CMC00262) (B, LAGU). La Libertad: J.C. Gonzalez & M.A. Hernandez 361 (B, EAP, LAGU, MO). San Salvador: M.C. Carlson 331 (F); R. Seiler 53 (F, ITIC, MHES); 506 (F, MHES); Tobar s.n. (12-802) (MHES). Chalatenango: D. Sloot & A. Reina 262 (MAG00691) (B, LAGU, U). La Paz: R. Villacorta et al. 293 (B, LAGU, MO). San Miguel: W. Eggler 663, F (ITIC). Cabafias: R. Seiler 521 (F, MHES); R. Villacorta & R.M. Rivas 2119 (B, LAGU, MO). Cuseatlan: F. Wunderlin 3026 (ITIC). Morazan: A.K. Monro et al. 3833 (B, BM, ITIC, LAGU, MO); R. Villacorta et al. (B, BM, LAGU, MO). Selaginella porphyrospora A. Braun, Ann. Sci. Nat. Bot., sér 5, 3: 286. 1865. Selaginella bulbifera Baker, S. binervis Liebm. ex Baker Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador. Salvadoran distribution: Sonsonate: R. Seiler 609 (F, MHES). Santa Ana: J. Williams 30 (ITIC); R. Seiler 877 (F, ITIC, MHES). San Salvador: H. Weber 7 (ITIC). Selaginella sertata Spring, Nouv. Mém. Acad. Roy. Sci. Bruxelles 24(1): 104. 1850. Selaginella nicaraguensis Baker Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: J.M. Rosales 1603 (AAU, B, BM, LAGU, MO). Santa Ana: R. Villacorta et al. 1237 (B, LAGU, MO). La Libertad: R. Villacorta & D.Lentz 1109 (B, LAGU, MO). Chalatenango: R. Seiler 147 (MHES). La Paz: R. Villacorta & S. Villacorta 1154 (B, LAGU, MO). San Miguel: R. Seiler 543 (MHES). Cabafias: R. Seiler 526 (MHES). > Selaginella silvestris Asplund, Ark. Bot. 20A(7): 30 (F). 3-5. 1926. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Bolivia. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 138 Salvadoran distribution: Tucker 1174 (UC) (Fraile in Davidse et al. 1995), Source: Stolze (1983): 60; Berendsohn (1989) 1: 24; Fraile in Davidse et al. (1995): 28: Mickel & Smith (2004): 596. > Selaginella stellata Spring, Flora 21: 194. 1838. Selaginella galeottii Spring lobal distribution: Mexico, Guatemala, Belize, Honduras, Costa Rica, Panama. Source: Berendsohn (1989) 1: 24. Selaginella tenella (P. Beauv.) Spring, Bull. Acad. Roy. Sci. Bruxelles 10: 234. 1843. Diplostachium tenellum P. Beauv., Lycopodium tenellum (P. Beauv.) Desv. ex Poir., Selaginella cladorrhizans A. Braun, S. subcaulescens Baker Local name: helecho tuya. Global distribution: Mexico, Guatemala, Belize, Honduras, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Brazil. Salvadoran distribution: Ahuachapan: E. Sandoval & O.Rivera D. 1376 (B, LAGU): J.M. Rosales 1298 (B, BM, EAP, LAGU, MO); 1340 (B, LAGU). La Libertad: R. Villacorta 316 (B, EAP, LAGU, MO). DIVISION EUPHYLOPHYTA, SUB-DIVISION MONILOPHYTA ANEMIACEAE Anemia guatemalensis Maxon, N. Amer. Fl. 16: 46. 1909. Local name: petatillo. Global distribution: Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 15 (F, LAGU, MO); R. Villacorta et a/. 1264 (B, ITIC, LAGU, MO). Chalatenango: D. Sloot & A. Reina 109 (MAG00538) (B, LAGU, U); L. Lara 220 (MHES); R. Seiler 562 (F, MHES). Morazan: J. Monterrosa & C. Rivera 712 (B. BM, LAGU, MO); R. Seiler 1050 (F, MHES). Anemia hirsuta (L.) Sw., Syn. Fil. 156. 1806. Osmunda hirsuta L., Ornithopteris hirsuta (L.) Bernh., Anemia hirsuta (L.) Sw. var. achillaeifolia M. Martens & Galeotti, 4. opaca Fée Local name: helecho pluma de perico. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval et al. 484, 1317 (B, LAGU); F. Chinchilla s.n. (ISB00522) (AAU, B, LAGU); s.n. (ISBO0501) (AAU, B, F, LAGU); s.n. (ISB00763) (B, LAGU); J.M. Rosales 898, 972, 1404 (AAU, B, BM, LAGU, MO); M. Sandoval & E. Sandoval 10, 39 (B, LAGU); R. Villacorta ef al. 406 (B, LAGU, MO): 952 (B, LAGU): W. Berendsohn & E. Sandoval 1481 (B, LAGU). Sonsonate: R. Seiler 35 (F, MHES): R. Villacorta 2402 (AAU, B, LAGU). Santa Ana: B. Pfeiffer- Berendsohn 57 (B, LAGU, MO); M.L. Reyna de Aguilar 765 (MHES); R. Villacorta & H. Sipman 1270 (B, LAGU, MO). La Libertad: N. Ventura 5013 (ITIC); O. Rohweder s.n. (JBL01254) (LAGU). San Salvador: J. Lagos 1076 (ITIC); R. Seiler 69 (F, MHES); 445 (MHES); R. Villacorta & M.A. Sanabria 402 (B, LAGU, MO). Chalatenango: R. Seiler 366 (F, MHES); 553, 569 (MHES); R. Villacorta 2362 (B, i323 FERN GAZ. 18(4): 120-215. 2008 LAGU, MO). La Paz: E.A. Montalvo 6388 (AAU, B, ITIC, LAGU, MEXU, MO). San Vicente: N.C. Fassett 28348 (F). Usulutan: D. Williams 217 (LAGU); 354 (B, LAGU, MO). Cabaiias: J. Monterrosa & R. Carballo 295 (B, BM, LAGU, MO). Morazan: J. Monterrosa & C. Rivera 664 (B, BM, LAGU). La Unién: S. Calderon 2374 (F). Anemia oblongifolia (Cav.) Sw., Syn. Fil. 156. 1806. Osmunda oblongifolia Cav., O. humilis (Cav.) Sw., Anemia oblongifolia (Cav.) Sw. B humilis (Cav.) Hook., A. pilosa M. Martens & Galeotti @ brevistipes Liebm., A. cornea Prantl, A. seemannii Hook. Local name: helecho rosquilla pacha. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Argentina. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Pérez 1347 (B, LAGU, MO); W. Berendsohn & R. Villacorta 1383 (B, LAGU). Sonsonate: R. Seiler 36, 494 (MHES); 73 (MHES). Santa Ana: O. Rohweder s.n. (JBL01255), s.n. (JBLO1257), 2660 (JBL0O1256) (LAGU). San Salvador: E.A. Montalvo 3775 (ITIC). Chalatenango: L. Lara s.n. (JBL02202) (B, LAGU); 77 (MHES); R. Seiler 558 (F, MHES); S. Calderon 2474 (F). Usulutan: D. Williams 140 (B, LAGU, MO). Morazan: J. Monterrosa & M. Rivera 602 (BM, LAGU); K. Sidwell e¢ al. 773 (B, BM, ITIC, LAGU, MO); M. Renderos et al. 629 (AAU, B, LAGU, MO); R. Villacorta et al. (B, BM, LAGU, MO). Anemia pastinacaria Moritz ex Prantl, Unters. Morph. Gefasskrypt. 2: 110. 1881. Anemia pilosa M. Martens & Galeotti B /ongistipes Liebm., A. longistipes (Liebm.) C. Chr., A. helveola Fée Local names: petatillo. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 60 (LAGU, MO); R. Seiler 134 (MHES). San Salvador: R. Seiler 205b, 782 (F, MHES). Chalatenango: D. Sloot & A. Reina 183 (MAG00612) (B, LAGU, U); R. Seiler 145 (F, MHES); 565 (MHES). Morazan: J. Monterrosa & C. Rivera 665 (BM, LAGU). Anemia phyllitidis (L.) Sw., Syn. Fil. 155. 1806. Osmunda phyllitidis L., Anemidictyon phyllitidis (L.) J. Sm. Local names: helecho cola de raton, helecho de montafia. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. Salvadoran distribution: Ahuachapan: B. Smalla & N. Kilian 81 (B, LAGU); E. Sandoval & H. Rivera 1319 (B, LAGU); J.M. Rosales 1005 (B, LAGU); M. Sandoval & E. Sandoval 53 (B, LAGU); R. Villacorta et al. 876 (B, LAGU); 955 (B, ITIC, LAGU, MO). Santa Ana: J. Monterrosa er a/. 260 (B, BM, LAGU, MO); 674 (B, BM, LAGU); 921 (LAGU); 1027 (B, LAGU, NY); R. Villacorta 2172 (LAGU). San Salvador: M.C. Carlson 332 (F); R. Seiler 65, 781 (F, MHES). Chalatenango: R. Seiler 566 (MHES); 570 (F, MHES). Usulutan: A.K. Monro et al. 3004 (B, BM, ITIC, MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 140 LAGU, MO); D. Williams 127, 131 (LAGU); 302 (LAGU, MO). Morazan: J. Monterrosa & C. Rivera 674 (B, BM, LAGU); R. Seiler 1087 (F); 1090 (F, MHES). ® Anemia salvadorensis Mickel, Amer. Fern. J. 74: 113. 1984. Local names: helecho de rosa, helecho plumerillo. Global distribution: Endem Salvadoran distribution: Miuadivele: E. Sandoval & F. Chinchilla 556 (B, LAGU); F. Chinchilla s.n. (ISB00541) (B, LAGU); s.n. (ISB00542) (LAGU); W. Berendsohn er al. 1395 (B, LAGU, MO); R. Seiler 947 (MHES, NY). ASPLENIACEAE Asplenium abscissum Willd., Sp. Pl., ed. 4, 5(1): 321. 1810. Asplenium polymorphum M. Martens & Galeotti. Local name: helecho quezalio. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, French Guyana; Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Uruguay, Paraguay. Salvadoran distribution: Ahuachapan: E. elle & F. Chinchilla 41 (LAGU); G Davidse eft al. 37420 (B, BM, ITIC, LAGU, MO); R. Seiler 901 (F, ITIC, MHES). Sonsonate: R. Seiler 619, 623 (F, MHES). Santa Ana: R. Seiler 111, 182, 363 (F, MHES). Asplenium achilleifolium (M. Martens & rete — Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. Ser. 51: 249. Caenopteris achilleifolia M. Martens & ae A a conchatum Fée, Asplenium conchatum (Fée) T. Moore, A. grande Fée Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Peru. Salvadoran distribution: Sonsonate: A. Molina et a/. 21623 (F):; R. Seiler 624 (F, MHES). Santa Ana: J. Tucher 1248 (F); M.B. Siu s.n. (ITIC); M.L. Reyna de Aguilar 762 (MHES); R. Seiler 468 (F, MHES); R. Seiler 469 (F, ITIC, MHES): R. Seiler 269 (ITIC (F, LAGU (photo), NY)); R. Seiler 872 (ITIC); S. Winkler 37 (ITIC). San Salvador: M.C. Carlson 417 (F); R. Seiler 346 (F, ITIC, MHES). Asplenium alatum Humb. & Bonpl. ex Willd., Sp. Pl. ed. 4, 5(1): 319. 1810. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 309 (MHES). R. Seiler 465 (F, LAGU (photo), NY). Asplenium auriculatum Sw., Kongl. Vetensk. Acad. Handl. 38: 68. 1817. Asplenium salicifolium L. var. auriculatum (Sw.) Proctor, A. semicordatum Raddi, A. pimpinellifolium Fée & Schaffner ex Fée Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Ahuachapan: D. Witsberger 686 (MHES); E. Sandoval & M. Sandoval 1423 (B, LAGU); R. Seiler 821, 823, 961 (F, MHES); 910 (F, ITIC, MHES); S. Winkler 6a (ITIC). Sonsonate: R. Seiler 615 (F, MHES). Santa Ana: A. Molina er 14] FERN GAZ. 18(4): 120-215. 2008 al. 12650, 16760, 16802a, 16806 (F; B). Lopez s.n. (ITIC); B. Pfeiffer-Berendsohn 32, 36 (B, MO, LAGU); E.A. Montalvo 12 (ITIC); G Cerén & D. Rodriguez s.n. (JBL04659) (B, LAGU, MO); G. Davidse et al. 37297 (B, BM, ITIC, LAGU, MO); J. Monterrosa & R. Carballo 554 (B, LAGU, MO); J. Tucker 1250 (F); K. Sidwell ef al. 849 (B, BM, ITIC, LAGU, MO); M.C. Carlson 995, 999 (F); M.L. Reyna de Aguilar 685 (MHES); R. Seiler 92 (F, ITIC, MHES); 361, 646 (F, MHES): R. Villacorta 1121 (B, LAGU). San Salvador: M.C. Carlson 428 (F); R. Seiler 338 F (MHES). Morazan: R. Seiler 1037 (F, ITIC, MHES); 1061 (F, MHES). Asplenium blepharophorum Bertol., Novi Comment. Acad. Sci. Inst. Bononiensis 4: 443. 1840. Athyrium macrocarpon Fée, Asplenium macrocarpon (Fée) T. Moore, A. myapteron Fée, A. commutatum Mett. ex Kuhn, 4. onustum Christ, A. subvestitum Copel. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles. Salvadoran distribution: Sonsonate: R. Seiler 15 (F). Santa Ana: R. Seiler 11 (ITIC, MHES); 85 (F, MHES). @ Asplenium cirrhatum Rich. ex Willd., Sp. Pl., ed. 4, 5(1): 321.1810. Asplenium radicans L. var. cirrhatum (Rich.) Rosenst. Global distribution: Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, French Guyana, Colombia, Venezuela, Ecuador, Peru. Brazil, Bolivia. Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar 848 (LAGU, MHES). @ Asplenium cristatum Lam., Encycl. 2: 310.1786. Asplenium cicutarium Sw. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Guayanas y Galapagos, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Anuachapan: A.K. Monro et al. 1993 (B, BM, ITIC, LAGU, MO). Asplenium cuspidatum Lam., Encycl. 2: 310.1786 . Asplenium lacerum Schltdl. & Cham., A. mexicanum M. Martens & Galeotti, A. fragrans Sw. var. mexicanum (M. Martens & Galeotti) Christ, 4. auritum Sw. vat. hia asi Kunze, 4. pyramidatum Liebm., A. auritum Sw. var. bipinnatisectum Ct, Local name: helecho trensillo de altura. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Brazil and Paraguay. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2008 (B, BM, ITIC, LAGU, MO); D. Witsberger 687 (MHES); E.A. Montalvo 60 (ITIC); E. Sandoval & M. Sandoval 1292 (B, LAGU); R. Seiler 827, 911 (ITIC, MHES); 962 (F, MHES); 963 (MHES); W. Berendsohn & M. Sandoval 1633 (LAGU); W. Berendsohn et al. 1519 (B, LAGU). Sonsonate: R. Seiler 183 (F); 617 (MHES). Santa Ana: A. del Cid 1618 (ITIC); J. Monterrosa & R. Martinez 949 (B, LAGU, MO, NY); L. Gonzalez 1617 (ITIC); M.L. Reyna de Aguilar 816 (MHES); R. Seiler 83, 183 (MHES); S. Winkler 47 MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 142 (ITIC). San Salvador: R. Seiler 213 (ITIC, MHES). Chalatenango: R. Seiler 384 (JBL00083), 414 (JBL00084) (F, LAGU, MHES). San Vicente: R. Seiler 319 (MHES): 330 (F, MHES). Usulutan: A. del Cid s.n. (ITIC); A.K. Monro, ef al. 3010 (B, BM, ITIC, LAGU, MO); D. Williams 52, 137 (B, LAGU, MO); D. Williams & R.W. Herrera 299 (LAGU); E.A. Montalvo 5033, 5045 (ITIC). Morazan: R. Seiler 975 (F, ITIC. MHES); 1018, 1036 (MHES). Asplenium feei Kunze ex Fée, Mém. Foug. 5: 194. 1852. Asplenium sanguinolentum Kunze ex Mett., A. sarcodes Maxon Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Suriname, Brazil, Bolivia. Salvadoran distribution: San Salvador: M. Carlson 428a (F). Asplenium flabellulatum Kunze, Linnea 9: 71. 1834. Asplenium flabellulatum Kunze var. dentatum Klotzsch, A. radicans L. var dentatum (Klotzsch) Bonap., A. flabellulatum Kunze var. lee Klotzsch, A. partitum (Klotzsch) C. Chr., 4A. radicans L. var partitum (Klotzsch) H Global distribution: Mexico, Guatemala, Honduras, ce es Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru. Paraguay. Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar 824 (MHES). Chalatenango: R. Seiler 406 (JBL00086) (F, LAGU, MHES). Asplenium formosum Willd., Sp. Pl., ed. 4, 5(1): 329.1810. Asplenium formosum Willd. var. incultrum H. Christ, A. subalatum Hook. & Arn. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Africa, S India, Sri Lanka. Local names: helecho peine de nifio, helecho peinetero, helecho de rosquilla. Salvadoran distribution. Ahuachapan: E. Sandoval & F. Chinchilla 99, 541 (B, LAGU); E. Sandoval & F. Pérez 1264 (B, LAGU); R. Seiler 927 (F, MHES); R. Villacorta et al. 939 (B, LAGU); W. Berendsohn et al. 1401, 1432, 1611 (B, LAGU): W. Berendsohn er a/. 1453 (LAGU). Sonsonate: R. Seiler 76 (F, MHES). Santa Ana: B. Pfeiffer-Berendsohn 18 (B, LAGU, MO); M. Magaria 28 (MP-00028) (LAGU); S. Winkler 14 (ITIC). La Libertad: E.A. Montalvo 6002 (ITIC); R. Seiler 713 (F, MHES). San Salvador: R. Seiler 49 (MHES). Chalatenango: J. Lagos 1720 (ITIC): L. Lara 234 (MHES); R. Seiler 146 (F, MHES); 372 (F, ITIC, MHES). Usulutan: A. del Cid 1031 (ITIC); A.K. Monro et al. 3014 (B, BM, ITIC, LAGU, MO); D. Williams & R.W. Herrera 315 (LAGU, MO); D. Williams 503 (LAGU); L. Gonzalez 1085 (ITIC). Cabafias: A.K. Monro et al. 3748 (B, BM, ITIC, LAGU, MO). Morazan: J. Monterrosa & C. Rivera 676 (B, BM, LAGU, MO); R. Seiler 1089 (F, MHES); S. Winkler 5 (ITIC). Asplenium fragrans Sw., Prodr. 130. 1788. Asplenium auritum Sw. vat. tripinnatum E. Fourn., A. cuspidatum Lam. var. tripinnatum (E. Fourn.) C.V. Morton & Lellinger, 4. foeniculaceum Kunth, A. cuspidatum Lam. var. foeniculaceum (Kunth) C.V. Morton & Lellinger, 4. cortifolium Liebm. Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, 143 FERN GAZ. 18(4): 120-215. 2008 Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Chalatenango: R. Seiler 151 (F, MHES); 422 (JBL00087) (F, ITIC, LAGU, MHES) Asplenium harpeodes Kunze, Linnea 18: 329.1844. Asplenium erectum Bory var. harpeodes (Kunze) Mett., A. pendulum Fée, A. donnell- smithii Christ . Local names: helecho lloron, quetzal silvestre. Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: A. Molina et al. 16802 (F); E.A. Montalvo 15b, 19 (ITIC); J. Monterrosa et al. 221 (B, BM, LAGU, MO); M.L. Reyna de Aguilar 1458 (LAGU); R. Seiler 120 (F, ITIC, MHES); R. Villacorta e¢ al. 1039, 1281 (B, LAGU, MO); S. Winkler 2 (ITIC). Chalatenango: J. Tucker 1105 (F). Morazan: J.L. Linares & A. Pool 2285 (JBL01930 ) (LAGU); J.L. Linares & C. Pefia H. 2749 (JBL01928) (LAGU). Asplenium hoffmannii Hieron., Hedwigia 60: 258. 1918. Asplenium membraniifolium Maxon Local names: helecho de roca. Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad and Tobago. Salvadoran distribution: Ahuachapan: J.M. Rosales 1104 (B, LAGU, MO); J.M. Rosales 1233 (AAU, B, BM, LAGU, MO). Sonsonate: D. Rodriguez & M. Trejo 98 (B, BM, LAGU, MO), NY. La Libertad: R. Seiler 718 (F, MHES). Asplenium laetum Sw., Syn. Fil. 79, 271. 1806. Asplenium lugubre Liebm. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Ahuachapan: M. Sandoval & E. Sandoval 297 (B, F, LAGU). Asplenium miradorense Liebm., Mexic. Bregn. 243. 1849. Asplenium bradeorum Hieron., A. schlechtendahlianum Hieron., A. standleyi Maxon Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, y Brazil. Salvadoran distribution: Sonsonate: A. Molina & E.A. Montalvo 21777 (JBL01478) (LAGU). Morazan: R. Seiler 1038 (F, MHES). Asplenium monanthes L., Mant. P|. 130.1767. Asplenium monanthemum L., A. arcuatum Liebm., A. obtusissimum Fée, A. galeotti Fée, A. leptophyllum Fée, A. curvatum Fée Global distribution: S USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Uruguay, Chile, Argentina, Africa, Hawaii. Salvadoran distribution: Ahuachapan: F. Wunderling 1226 (ITIC); Santa Ana: B. Pfeiffer-Berendsohn 80 (LAGU, MO); G. Cerén & D. Rodriguez s.n. (JBL04646) MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 144 (LAGU); J. Monterrosa ef al. 209, 908 (LAGU); J. Monterrosa et al. 931 (B, LAGU, MO, NY); M. B. Siu s.n. (ITIC); M.C. Carlson 724 (F); M.L. Reyna de Aguilar 673 (MHES); R. Seiler 313 (F, MHES). San Salvador: R. Seiler 210 (F, MHES); 516 (MHES). Chalatenango: R. Seiler 387 (JBLOO088) (F, LAGU, MHES); R. Villacorta et al, 2073 (B, LAGU, MO); W. Berendsohn et a/. 1572 (B, LAGU, MO). San Vicente: R. Seiler 331 (JBL04807) (F, LAGU, MHES). @ Asplenium monodon Liebm., Mexic. Bregn. 247. 1849. Asplenium auritum Sw. var. monodon (Liebm.) E. Fourn. Global distribution: USA, Mexico, Guatemala, Honduras, Costa Rica, Panama. Antilles, Venezuela, Colombia, Trinidad and Tobago, Ecuador. Salvadoran distribution: Santa Ana: R. Seiler 466 (MHES). Asplenium polyphyllum Bertol., Novi Comment. Acad. Sci. Inst. Bononiensis 4: 443. 840. Asplenium polymeris T. Moore, A. bertolonii Donn. Sm., A. trichomanes L. var. viridissimum Christ, A. viridissimum J. Bommer ex Christ Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 41 (LAGU, MO); J. Monterrosa & R. Carballo 252 (B, BM, LAGU, MO); J. Monterrosa & R. Carballo 837 (B, LAGU, MO); M.L. Reyna de Aguilar 764 (MHES); R. Seiler 464 (F, MHES); R. Villacorta & L. Lara 2663 (B, LAGU, MO); V.M. Martinez s.n. (CMCO01046) (B, LAGU, MO); L. Lara 238 (MHES). Chalatenango: R. Seiler 160 (F, ITIC, MHES). Asplenium potosinum Hieron., Hedwigia 60: 247.1918. Asplenium regulare Sw. var. latior E. Fourn., A. /unulatum Sw. var. latius E. Fourn., A. hesperium Mickel & Beitel Local names: helecho cola de pavo real. Global distribution: Mexico, Guatemala. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 40 (LAGU, MO): R. Seiler 824 (JBL04817) (F, ITIC, LAGU, MHES); R. Seiler 903 (F, ITIC, MHES); 955 (F, MHES); W. Berendsohn & E. Sandoval 1513 (B, LAGU). Usulutan: A.K. Monro et al. 3008, 3015 (B, BM, ITIC, LAGU, MO); D. Williams 53 (LAGU, MO); D. Williams 413 (LAGU). Asplenium praemorsum Sw., Prodr. 130. 1788. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: M. Magafia 33 (MP-00018) (LAGU); R. Seiler 819 (F, MHES). Morazan: R. Seiler 1035 (F); 1072 (F, MHES). > Asplenium pulchellum Raddi, Opusc. Sci. 3: 291.1819. Asplenium pulchellum Raddi var. subhorizontale Rosenst. Global distribution. Mexico, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay. Source: Seiler (1982): 387; Berendsohn (1989): 5; Mickel & Smith (2004): 114. 145 FERN GAZ. 18(4): 120-215. 2008 Asplenium pumilum Sw., Prodr. 129. 1788. Asplenium anthriscifolium Jacq., A. leucothrix Maxon, Athyrium verapax Christ Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Africa. Salvadoran distribution: Sonsonate: M.L. Reyna de Aguilar 1453 (LAGU); R. Seiler 74 (MHES); 495 (F, MHES). Santa Ana: R. Seiler 703 (MHES). San Salvador: RK: Seiler 582 (F, MHES). Usulutan: A. del Cid 1785 (ITIC); J. Lagos 1786 (ITIC). Asplenium seileri C.D. Adams, Novon 2: 291.1992. Global distribution: Mexico, Guatemala, Costa Rica. Salvadoran distribution: Ahuachapan: A. Molina et al. 21797 (ITIC); G. Davidse er al. 37372 (B, BM, ITIC, LAGU, MO); R. Seiler 957, (F, MHES). Sonsonate: R. Villacorta 751 (LAGU). Asplenium serra Langsd. & Fischer, Pl. Voy. Russes Monde I: 16.1810. Asplenium serra Langsd. & Fischer var. incurvatum H. Christ, A. insigne Liebm., A. progrediens Fée Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay, Uruguay. Salvadoran distribution: Santa Ana: A. Molina et al. 16755 (F); J. Monterrosa & R. Carballo 860 (B, LAGU, MEXU, MO); R. Seiler 106, 191b (F, MHES); 114 (MHES); 192 (F); V.M. Martinez s.n. (CMC01095) (B, EAP, LAGU, MO). Chalatenango: J. Tucker 1045 (F); R. Seiler 424 (F, MHES); W. Berendsohn et al. 1574 (B, LAGU, MO). Asplenium sessilifolium Desy. var. sessilifolium, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 322.1811. Asplenium sessilifolium Desv. var. guatemalense Hieron. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Sonsonate: R. Seiler 625 (F, MHES). Santa Ana: A. Molina & E.A. Montalvo 21512 (JBL01516) (F, LAGU); B. Pfeiffer-Berendsohn 55 (LAGU, MO); J. Monterrosa & R. Martinez 936 (B, LAGU, MO, NY): R. Seiler 121 (JBL04803) (F, LAGU, MHES); R. Seiler 13 (ITIC, MHES); 84 (ITIC); 354 (F, ITIC, MHES). Chalatenango: L. Lara 233 (MHES); R. Seiler 378 (JBLO0085) (F, LAGU, MHES). Asplenium tenerrimum Mett. ex Kuhn, Linnea 36: 97. 1869. Asplenium cuspidatum Lam. var. tenerrimum (Mett. ex Kuhn) C.V. Morton & Lellinger, A. fournieri Kuhn ex E. Fourn. Global distribution: Mexico, Honduras, Nicaragua, Colombia, Venezuela, Ecuador. Salvadoran distribution: Santa Ana: R. Seiler 545 (F). @Asplenium theciferum (Kunth) Mett., Ann. Sci. Nat. Bot. sér. 5, 2: 227. 1864. Davallia thecifera Kunth, D. lindenii Hook., Darea thecifera (Kunth) E. Fourn., Loxoscaphe lindenii (Hook.) T. Moore, L. thecifera (Kunth) T. Moore Global distribution: Mexico, Honduras, Nicaragua, Costa Rica, Panama, Antilles, MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 146 Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Africa. Salvadoran distribution: Santa Ana: J.L. Linares 391 (JBLO1410) (LAGU). Asplenium virillae Christ, Bull. Herb. Boissier, sér. 2, 4: 1090. 1904. Asplenium alfredii Rosenst. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Colombia. Venezuela. Salvadoran distribution: Sin loc: P. Standley 19774, US (fide Adams, 1995). Santa Ana: J. Monterrosa et al. 1457 (BM, LAGU, MO, NY): 130. BLECHNACEAE Blechnum appendiculatum Willd., Sp. Pl., ed. 4, 5(1): 410. 1810. Blechnum glandulosum Kaulf. ex Link, B. meridionale C. Presl, B. occidentale L. var. minor Hook., B. occidentale L. var. pubirhachis Rosenst. Local names: helecho esqueleto de pescado, petatillo. Global distribution. USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1994 (B, BM, ITIC, LAGU, MO); F. Pérez s.n. (ISB00003) (B, LAGU, MO); G. Davidse et al. 37417 (B, BM, ITIC, LAGU, MO); J.G. Sandoval ef a/. s.n. (ISB00027) (B, LAGU, MO), US; K. Sidwell er al. 814 (B, BM, ITIC, LAGU, MO); S. Castillo s.n. (ISF00594) (B, F, LAGU, MO). Sonsonate: R. Villacorta & A. Gonzalez 693 (B, LAGU, MO). Santa Ana: B. Pfeiffer- Berendsohn 65 (LAGU, MO); D. Rodriguez & G. Cerén s.n. (JBL04618) (LAGU); E.A. Montalvo 3654 (ITIC); J. Monterrosa et al. 907 (B, LAGU, MO, NY); M.B. Siu s.n. (ITIC); M.L. Reyna de Aguilar 684 (MHES); R. Seiler 349, 477 (F, MHES): V.M. Martinez s.n. (CMC01096) (B, EAP, LAGU, MO). La Libertad: A. Mangandi ef ai. s.n. (JBL04492) (LAGU); A.K. Monro ef al. 2276, 3086 (B, BM, ITIC, LAGU, MO); J.C. Gonzalez 415 (B, LAGU): M. Renderos et a/. 412 (LAGU). San Salvador: A.K. Monro ef al, 2217, 2223 (B, BM, ITIC, LAGU, MO, L).O. Williams ef a/. 15232 (F): M.C. Carlson 329 (F); R. Seiler 29 (F, MHES). Chalatenango: D. Sloot & A. Reina 216 (MAG00645), 433 (MAG00862) (LAGU, U); L. Lara 218, 225 (MHES): M. Renderos & R. Villacorta 141 (B, LAGU); R. Seiler 413 (JBL00091) (LAGU, MHES): W. Heed 26 (ITIC). San Vicente: E.A. Montalvo 4615, 4618 (ITIC). Usulutan: L. Gonzalez 1089, 1787 (ITIC). San Miguel: K. Sidwell ef a/. 885 (B, BM, ITIC, LAGU, MO): M. Renderos ef al. 687 (LAGU, MO). Cuscatlan: S. Dar s.n. (JBL03666) (LAGU, MO). Morazan: J. Monterrosa & M. Rivera 615 (B, BM, LAGU); J. Tucker 608 (F). Blechnum appendiculatum Willd. x Blechnum gracile Kaulf Global distribution. Mexico, Honduras, Nicaragua, Colombia, Venezuela. Salvadoran distribution: Santa Ana: R. Seiler 763 (MHES). Chalatenango: L. Lara 209 (MHES). Blechnum x caudatum Cav., Descr. Pl. 262. 1802. . . Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Suriname, Ecuador, Peru, Brazil. 147 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 59 (B, LAGU, MO); C. Salazar s.n. (ITIC). Morazan: A.K. Monro et al. 3804 (B, BM, ITIC, LAGU, MO); J. Monterrosa & R. Carballo 794 (LAGU). Note: Blechnum x caudatum is a hybrid between Blechnum gracile y B. occidentale (Mickel & Smith 2004: 146, Fig.: 55C). Blechnum divergens (Kunze) Mett., Ann. Sci. Nat. Bot., sér. 5, 2: 255.1864. Lomaria divergens Kunze, L. exaltata Fée, Struthiopteris exaltata (Fée) Broadh. Global distribution. Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 765, 892 (F, MHES); 1156, F (ITIC). Blechnum ensiforme (Liebm.) C.Chr., Index Filic. 153: 1905. Lomaria ensiformis Liebm., Struthiopteris ensiformis (Liebm.) Maxon Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: G. Cerén & D. Rodriguez s.n. (JBL04660) (LAGU); R. Seiler 299 (F, MHES). Chalatenango: R. Seiler 389 (JBL00092) (F, LAGU, MHES). Morazan: R. Seiler 1026 (F, ITIC, MHES). Blechnum falciforme (Liebm.) C. Chr., Index Filic. 154. 1905. Lomaria falciformis Liebm., L. deflexa Liebm., Struthiopteris falciformis (Liebm.) Broadh. Global distribution. Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Sonsonate: R. Seiler 632 (F, MHES). Santa Ana: A. Hernandez s.n. (ITIC); M.C. Carlson 725, 891, 952 (F); R. Seiler 348, 478 (MHES): 863 (F, ITIC, MHES). Chalatenango: N. Ventura 5026 (ITIC); R. Seiler 409 (MHES). Morazan: R. Seiler 990 (F, MHES). Blechnum fragile (Liebm.) C.V. Morton & Lellinger, Amer. Fern J. 57: 68. 1967. Lomaria fragilis Liebm., L. polypodioides (Sw.) Desv., L. onocleoides (Sw.) Spreng., L- mexicana Fée, Blechnum binervatum (Poir.) C.V. Morton & Lellinger subsp. fragile (Liebm.) R.M. Tryon & Stolze, B. onocleoides Sw., B. polypodioides (Sw.) Kuhn, Osmunda polypodioides Sw. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 654 (F, MHES). Morazan: R. Seiler 1041 (F, ITIC, MHES). Blechnum gracile Kaulf., Enum. Filic. 158. 1824. Blechnum intermedium Link., B. subdimorphum Copel., B. fraxineum auct., non Willd. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname. French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Chalatenango: M.B. Siu s.n. (ITIC); M.C. Carlson 594 (F); R. Seiler 572 (F, ITIC); (MHES). Cabafias: A.K. Monro er al. 3746 (B, BM, ITIC, LAGU, MO). Morazan: R. Seiler 1083 (F, ITIC, MHES). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 148 Blechnum lehmannii Hieron., Bot. Jahsb. Syst. 34: 473. 1904. Blechnum I’herminieri (Bory) C. Chr. subsp. /ehmannii (Hieron) Lellinger, Struthiopteris maxonii Broadh. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Sonsonate: J.L. Linares & C.A. Martinez 3033 (JBL01933) (LAGU). Santa Ana: F. Wunderling 1245 (ITIC); G. Cerén & D. Rodriguez s.n. (JBL04650) (LAGU); R. Seiler 894 (F, ITIC, MHES). Morazan: R. Seiler 979, 1039.F (ITIC, MHES). Blechnum occidentale L., Sp. P|. 2: 1077. 1753. Local names: helecho esqueleto de pescado. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1929, 2984 (B, BM, ITIC, LAGU, MO): A. Sermefio 98 (JBL00999) (B, LAGU, MEXU, MO); E. Sandoval & M. Sandoval 864 (B, LAGU); J.M. Rosales 1188 (B, LAGU, MO); 1715, 1723 (B, BM, LAGU, MO); M. Magafia 19 (MP-00032) (LAGU); M. Sandoval & E. Sandoval 136 (B, F, LAGU); R. Villacorta 622 (B, LAGU, MO); S. Castillo & B.A. Martinez s.n. (ISF00406) (AAU, F, LAGU); W. Berendsohn ef al. 1316, 1368, 1382, 1393 (B, LAGU, MO). Sonsonate: D. Rodriguez & G Trejo 203 (LAGU). Santa Ana: G. Davidse et al. 37226 (B, BM, ITIC, LAGU, MO): J. Monterrosa & R. Carballo 509 (LAGU); K. Prestegard 10 (MP-00031) (LAGU); M.L. Reyna de Aguilar e¢ al. 1439 (B, ITIC, LAGU, MO). La Libertad: F. Wunderling 461 (ITIC); N. Ventura 5001, 5004 (ITIC); O. Tejada et al. s.n. (JBL04504) (LAGU); R.Guerra et al. s.n. (JBL04502) (LAGU); R.Menjivar & J. Sermefio s.n. (JBL04483) (LAGU). San Salvador: M.C. Carlson 4a (ITIC); R. Seiler 457 (F, MHES); 783 (F, ITIC, MHES). Chalatenango: E.A. Montalvo 30, 3202, 3778 (ITIC); L. Lara 498 (MHES); M.B. Siu s.n. (ITIC). Usulutan: D. Williams 54 (LAGU, MO). San Miguel: A.K. Monro et a/. 2944 (B, BM, ITIC, LAGU, MO): A. Salgado 66 (MHES). Cuseatlan: E.A. Montalvo 3312 (ITIC). Cabafias: A.K. Monro & K. Sidwell 2778 (B, BM, ITIC, LAGU, MO); G. Davidse er al. 37083 (B, BM, ITIC, LAGU, MO). Morazan: J. Monterrosa ef al. 45 (B, LAGU, MO) Blechnum polypodioides Raddi, Opusc. Sci. 3: 294.1819, non (Sw.) Kuhn, 1868. Asplenium blechnoides Lag. ex Sw., Blechnum unilaterale Sw., B. blechnoides (Lag. ex Sw.) C. Chr., B. scabrum Liebm., B. angustifrons Fée Local names: helecho peineta amarilla. . Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: W. Berendsohn er al. 1469 (LAGU). Chalatenango: M.B. Siu s.n. (ITIC); L. Lara 230 (MHES); R. Seiler 564 (MHES): 571 (F, MHES). Usulutan: D. Williams 55 (LAGU, MO). Morazan: J. Monterrosa & C. Rivera 690 (B, BM, LAGU): J. Tucker 643 (F). 149 FERN GAZ. 18(4): 120-215. 2008 Blechnum schiedeanum (Schltdl. ex C. Presl) Hieron., Hedwigia 47: 239. 1908. Lomaria schiedeana Schitdl. ex C. Presl, L. spectabilis Liebm., L. longifolia Schltdl. ex M. Martens & Galeotti, L. danaeacea Kunze, L. acrodonta Fée, L. varians E. Fourn., L. costaricensis H. Christ, Struthiopteris costaricensis (H. Christ) Broadh., Blechnum danaeaceum (Kunze) Christ, B. varians (E. Fourn.) C. Chr., B. costaricensis (H. Christ Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution. Anuachapan: B. Pfeiffer-Berendsohn 51 (B, LAGU, MO); 53 (B, LAGU); J. Monterrosa & R. Carballo 262 (B, LAGU, MO); R. Seiler 196, 681 (F, MHES). Santa Ana: A. Hernandez s.n. (ITIC). Morazan: R. Seiler 1034 (F, TTC, MHES); 1051 (F, MHES). > Blechnum serrulatum Rich., Actes Soc. Hist. Nat. Paris I: 114. 1792. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Australia. Sources: Calderén & Standley (1941): 30; Seiler (1982): 389; Berendsohn (1989), 1: 6; Moran in Davidse ef al. (1995): 330; Mickel & Smith (2004): 151 Blechnum stoloniferum (Mett. ex E. Fourn.) C. Chr., Index Filic. 154. 1905. Lomaria stolonifera Mett. ex E. Fourn., L. ghiesbreghtii Baker, Struthiopteris stolonifera (Mett. ex E. Forun.) Broadh., Bleohiign — (Baker) C. Chr. Global distribution: Mexico, Guatemala, Honduras Ric Salvadoran distribution: Santa Ana: R. Seiler 678 (F); 780 a ee Chalatenango: R. Seiler 731 (F, MHES). Blechnum wardiae Mickel & Beitel, Mem. New York Bot. Gard. 46: 89. 1988. Global distribution: Mexico, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Santa Ana: A. Molina et al. 16837 (F); E.A. Montalvo 17 ITIC) >» Woodwardia semicordata Mickel & Beitel, Mem. New York Bot. Gard. 46: 403. 1988, pro hybr. Global distribution: Mexico, Costa Rica. Source: Mickel & Smith (2004): 694. Woodwardia spinulosa M. Martens & Galeotti, Mém. Foug. Mexique 64. 1842. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: G. Davidse et al. 37216 (B, BM, ITIC, LAGU, MO); V.M. Martinez s.n. (CMC00979) (B, LAGU, MO); s.n. (CMC01024) (B, BM, EAP, LAGU, MO); R. Seiler 471 (F, MHES); W. Berendsohn e¢ al. 1679 (LAGU). Chalatenango: R. Seiler 727 (MHES). Morazan: R. Seiler 1002 (F); 1027 (F, MHES). CIBOTIACEAE Cibotium regale Verschatf. & Lem., Ill. Hort. 15: under pl. 548 (F). 1868. Cibotium guatemalense Rchb. f. ex Kuhn, C. wendlandii Mett. ex Kuhn MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 150 Local names: cabeza de nifio. Global distribution: Mexico, Guatemala, Belize, Honduras. Salvadoran distribution: R. Seiler 151, MO (fide B. Pérez-Garcia in Davidse et al. 1995). Santa Ana: R. Seiler 186 (MHES); 304 (F, ITIC, MHES). La Libertad: K. Prestegard 36 (MP-00025) (LAGU). Usulutan: D. Williams & R.W. Herrera 471 (B, LAGU, MEXU, MO). Note: listed as ‘at risk’ by MARN, 2004. CULCITACEAE Culcita coniifolia (Hook.) Maxon, Annual Rep. Board Regents Smithsonian Inst. 1911: 488, t. 13C. 1912. Dicksonia coniifolia Hook. Global distribution: Mexico, Guatemala, Honduras, oa Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivi Salvadoran distribution: Molina R. 12682, GH (Pérez- peices in Davidse et al. 1995). Santa Ana: A. Molina et al. 12682 (F); R. Seiler 485 (F, ITIC, MHES); 1129 (F, ITIC); L. Lara 279 (MHES). Note: listed as ‘at risk’ by MARN, 2004. CYATHEACEAE Alsophila firma (Baker) D. S. Conant, J. Arnold Arbor. 64: 372. 1983. Cyathea firma (Baker) Domin, C. mexicana Schltdl. & Cham., C. trejoi Christ, C. articulata Fée, Nephelea mexicana (Schltdl. & Cham.) R. M. Tryon Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Ecuador. Local names: helecho de espina, helecho cuerpo espin Salvadoran distribution: Ahuachapan: M. Sandoval & E. Sandoval 9 (B, LAGU); W. Berendsohn et al. 1455 (B, LAGU). Alsophila salvinii Hook. in Hooker & Baker, Syn. Fil. 36. 1866. Cyathea salvinii (Hook.) Domin, Alsophila muenchii Christ Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Peru. Salvadoran distribution: R. Seiler 899, MO (Riba in Davidse ef a/. 1995). Santa Ana: A. Molina et al. 16749 (F); R. Seiler 296 (F, ITIC, MHES). Note: Listed as ‘threatened’ by MARN, 2004. Alsophila tryoniana (Gastony) D.S. Conant, J. Arnold Arbor. 64: 371. 1983. Nephelea tryoniana Gastony Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: R. Seiler 1154, MO (Riba in Davidse et a/. 1995). Santa Ana: R. Seiler 236, 300 (F, ITIC, MHES). Morazan: R. Seiler 976 (F, ITIC, MHES). Note: Listed as ‘threatened’ by MARN, 2004. Cyathea bicrenata Liebm., Mexic. Bregn. 289 (reprint 137). 1849. ee Alsophila bicrenata (Liebm.) E. Fourn., A. scabriuscula Maxon, Trichipteris bicrenata (Liebm.) R. M. Tryon, 7. scabriuscula (Maxon) R. M. Tryon ES FERN GAZ. 18(4): 120-215. 2008 Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Santa Ana: R. Seiler 301 (F, ITIC, MHES). Note: Listed as ‘at risk’ by MARN, 2004; listed as Apendix II by CITES. Cyathea costaricensis (Mett. ex Kuhn) Domin, Acta Bot. Bohem. 9: 107. 1930. Hemitelia costaricensis Mett. ex Kuhn, Trichipteris costaricensis (Mett. ex Kuhn) Barrington . Local name: helecho pescuezo de violin. Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: K. Prestegard 18 (MP-00026) (LAGU); R. Seiler 929 (F, MHES); W. Berendsohn et al. 1352, 1414 (B, LAGU, MO). Sonsonate: J. Tucker 1355 (F). Santa Ana: S. Winkler s.n. (ITIC). La Libertad: A.K. Monro et al. 2307 (B, BM, ITIC, LAGU, MO). San Salvador: R. Seiler 224, 368, 535, 787 (F, MHES). Note: Listed as ‘at risk” MARN, 2004; listed as Apendix II by CITES. Cyathea divergens Kunze var. divergens, Linnaea 9: 100. 1834 Alsophila subaspera H. Christ, C. divergens var. hirta |. Losch, C. divergens var. minor Rosenst., C. pelliculosa H. Christ, C. subaspera (H. Christ) Domin. Global distribution: Costa Rica, Panama, Colombia, Venezuela, Guayana, Suriname, Ecuador, Peru. Salvadoran distribution: Santa Ana: R. Seiler 778 (F, ITIC, MHES). Note: This species is cultivated at Los Planes, Montecristo national park (Pers. Obs.). Cyathea divergens vat. tuerckheimii (Maxon) R.M. Tryon, Contr. Gray Herb. 206: 56. Cyathea tuerckheimii Maxon, Cyathea juergensenii E. Fourn. Global distribution: Mexico, Belice, Guatemala, Nicaragua. Salvadoran distribution: Santa Ana: J.Monterrosa & D.Rodriguez 1539 (LAGU); R. Seiler 276 (MHES). Morazan: R. Seiler 980 (F, MHES). Note: Listed as ‘at risk” MARN, 2004; listed as Apendix II by CITES. » Cyathea fulva (M. Martens & Galeotti) Fée, Mém. Foug. 9: 34. 1857. Alsophila fulva M. Martens & Galeotti, Cyathea delicatula Maxon, C. compersa H. Christ, C. furfuracea H. Christ, C. mollis Rosenst., C. molliuscula Domin, C. onusta H. Christ, C. papyracea H. Christ, C. underwoodii H. Christ Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador. Note: Listed as Apendix II by CITES. Sources: Mickel & Smith (2004): 237. Cyathea godmanii (Hook.) Domin, Pterid. Dominica 262. 1929. Alsophila godmanii Hook., A. mexicana Mart., Cyathea mexicana Schidtl. & Cham., C. valdecrenata Domin, Trichipteris mexicana (Mart.) R.M. Tryon Global distribution: Mexico, Guatemala, Honduras. Panama. Salvadoran distribution: Santa Ana: R. Seiler 243 (F, ITIC, MHES). Note: Listed as ‘threatened’ by MARN, 2004; listed as Apendix II by CITES. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 152 Cyathea schiedeana (C. Pres!) Domin, Pterid. Dominica 263. 1929. Alsophila schiedeana C. Presl, A. crassifolia H. Christ, A. chnodédes H. Christ, Cyathea chnoddes (H. Christ) Domin, Trichipteris schiedeana (C. Presl) R.M. Tryon, T. crassifolia (H. Christ) Gastony Global distribution: Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Carballo 848 (LAGU); R. Seiler 172, 237 (F, MHES). Chalatenango: J. Tucker 1129 (F); R. Seiler 760 (F, MHES). Morazan: J. Tucker 788 (F); R. Seiler 1082 (F, ITIC, MHES). Note: Listed as ‘threatened’ by MARN, 2004; listed as Apendix II by CITES. Sphaeropteris horrida (Liebm.) R.M. Tryon, Contr. Gray Herb. 200: 20. 1970, non Cyathea horrida (L.) Sm. Cibotium horridum Liebm., C. princeps Linden ex J. Smith, Cyathea princeps E. Mayer, C. bourgeaui E. Fourn., C. glauca E. Fourn., C. muenchii Christ Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: R. Seiler 242 (MHES), F (ITIC). Note: Listed as ‘at risk’ by MARN, 2004. DENNSTAEDTIACEAE Dennstaedtia bipinnata (Cav.) Maxon, Proc. Biol. Soc. Wash. 51: 39. 1938. Dicksonia bipinnata Cav., D. adiantoides Humb. & Bonpl. ex Willd. Local name: helecho palma de paraiso. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia. Salvadoran distribution: P.Standley 19760 (GH) (Moran in Davidse et al. 1995). Ahuachapan: E. Sandoval & F. Chinchilla 123 (LAGU); S. de Moore s.n. (12-909) (MHES). San Salvador: R. Seiler 1214 (F). Dennstaedtia cornuta (Kaulf.) Mett., Ann. Sci. Nat. Bot,, sér. 5, 2: 260. 1864. Dicksonia cornuta Kaulf. Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles. Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Argentina. Salvadoran distribution: Santa Ana: R. Seiler 165, 271, 693, 1192 (F, MHES). Dennstaedtia distenta (Kunze) T. Moore, Index Fil. 306. 1861. Dicksonia distenta Kunze Global distribution: Mexico, Guatemala, Costa Rica, Panama, Antilles. Salvadoran distribution: Standley & Valerio 49449, GH (Moran in Davidse et a/. 1995). Morazan: Tucker 726 (F). @ Dennstaedtia globulifera (Poir.) Hieron., Bot. Jahrb. Syst. 34: 455. 1904. . Polypodium globuliferum Poir., Dicksonia globulifera (Poir.) Kuntze, Dennstaedtia gracilis A, Rojas & Tejero Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, cami ‘ Antilles, Colombia, Venezuela, Ecuador, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. 153 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: San Salvador: S. Winkler 74 (ITIC). @ Dennstaedtia spinosa Mickel, Amer. Fern J. 58: 90. 1968. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: W. Berendsohn et al. 1320 (B, LAGU, MO). Note: This collection was cited as Dennstaedtia cicutaria (Moran in Davidse ef al. 1995), however it posesses spines that are diagnostic for D. spinosa. Histiopteris incisa (Thunb.) J. Sm., Hist. Fil. 295. 1875. Pteris incisa Thunb., Litobrochia incisa (Thunb.) C. Presl Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Africa, Asia. peeps distribution: R. Seiler 1195 (UC) (Moran in Davidse ef al. 1995). Santa Ana: A. Molina et al. 12534 (F); R. Seiler 253 (F, MHES). Chalatenango: J. Tucker 1122 (F). Hypolepis blepharochlaena Mickel & Beitel, Mem. New York Bot. Gard. 46: 219. Global distribution: Mexico, Guatemala, Honduras, Costa Rica. Salvadoran distribution. Santa Ana: A. Molina et al. 12528, 16775, 16885 (F); R. Seiler 249, 294 (F, MHES). Chalatenango: R. Seiler 427 (JBL00108) (LAGU, MHES). Hypolepis bogotensis H. Karst., Fl. Columb. 2: 91. 1865. Dennstaedtia munchii H. Christ, Hypolepis flexuosa Sodiro Global distribution: Mexico, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivi Salvadoran distribution: ciataienstaas Tucker 1121 (F). Hypolepis nigrescens Hook., Sp. Fil. 2: 66, t. 90C. 1852. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: J.Monterrosa & D.Rodriguez 1532 (LAGU) Hypolepis repens (L.) C. Presl, Suppl. Tent. Pterid. 162. 1836. Lonchitis repens L., Cheilanthes repens (L.) Kaulf., Hypolepis mexicana Liebm. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: Tucker 1303 (F, LAGU (photo), NY). Pteridium aquilinum var. feei (W. Schaffn. ex Fée) Maxon ex Yunck., Publ. Field Mus. t. Hist., Bot. Ser. 17: 308. 1938. Ment Jeei W. Schaff. ex Fée, Pteridium feei (W. Schaffn. ex Fée) Faull Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R. Seiler 777 (F, MHES). Chalatenango: R. appeal (JBLO00109) (F, LAGU, MHES). Morazan: J. Tucker 696 (F); R. Seiler 1058 is i: MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 154 Note: An invasive species that has been responsible for the poisoning of cattle throughout the country (Ministerio de Ganaderia, Pers. Comm.. Pteridium arachnoideum (Kaulf.) Maxon, J. Wash. Acad. Sci. 14: 89, 1924. Pteris arachnoidea Kaulf., P. aquilina L. var. arachnoidea (Kaulf.) D. C. Eaton, Pteridium aquilinum (L.) Kuhn var. arachnoideum (Kaulf.) Brade, P. caudatum ( LJ Maxon subsp. arachnoideum (Kaulf.) Lellinger . Local name: helecho petatillo Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Urugua Salvadoran distribution: Ahuachapan: D. Witsberger 665 (MHES): B. Sandoval & F. Chinchilla 429 (B, LAGU); R. Seiler 665 (MHES). Santa Ana: R. Seiler 306 (F, MHES). Chalatenango: D. Sloot & A. Reina 310 (MAG00739) (LAGU, U);: R. Seiler 266 (F, MHES). San Vicente: R. Seiler 333 (MHES). Morazan: R. Seiler 1093 (F); 1094 (F, MHES). Pteridium caudatum (L.) Maxon, Proc. U.S. Natl. Mus. 23: 631. 1901. Pteris caudata L., P. aquilina L. var. caudata (L.) Link, P. aquilina L. var. mexicana Fée, P. caudata L. var. mexicana Fée, Pteridium aquilinum (L.) Kuhn var. caudatum (L.) Sadeb., P. aquilinum subsp. caudatum (L.) Bonap. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Carballo 251 (B, LAGU, MO); J. Tucker 1298 (F). La Libertad: J.C. Gonzalez 372 (B, LAGU, MO); J. Lagos s.n. (ITIC). San Salvador: M.C. Carlson 404, 418 (F); S. Calderon 461 (F). Chalatenango: M.C. Carlson 611 (F); R. Villacorta & A. Araniva 981 (B, LAGU, MO). Usulutan: D. Williams & R.W. Herrera 451 (LAGU). Morazan: J. Tucker 697 (F). DICKSONIACEAE Dicksonia sellowiana Hook., Sp. Fil. 1: 67. 1844. Dicksonia gigantea H. Karst., D. ghiesbreghtii Maxon Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Uruguay. Salvadoran distribution: Santa Ana: A. Molina et al. 12523, 16757, 16915, 16917 (F): M.C. Carlson 964 (F); R. Seiler 240 (F, ITIC, MHES); S. Winkler 100 (ITIC). Chalatenango: J. Tucker 1127 (F, MO) (Pérez-Garcia in Davidse er al. 1995): R. Seiler 416 (JBL00094), (JBL00095) (F, LAGU, MHES). Note: Note: listed as ‘at risk” by MARN, 2004; listed as Apendix II by CITES. Lophosoria quadripinnata (J. F. Gmel.) C. Chr. in Skottsberg, Nat. Hist. Juan Fernandez I: 16. 1920. . Polypodium glaucum Sw., P. quadripinnatum J. F. Gmel., P. pruinatum Sw., Alsophila quadripinnata (J. F. Gmel.) C. Chr., A. pruinata (Sw.) Kaulf. ex Kunze, Trichosorus glaucescens Liebm., T. densus Liebm., T. frigidus Liebm. Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Be FERN GAZ. 18(4): 120-215. 2008 Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia, Chile, Argentina. Salvadoran distribution. Santa Ana: A. Molina et al. 16773 (F); G Cerén & D. Rodriguez s.n. (JBL04655) (B, LAGU); s.n. (JBL04656), s.n. (JBL04657) (LAGU); R. Seiler 292 (F, ITIC, MHES). Chalatenango: J. Tucker 1150 (F). Morazan: R. Seiler 1030 (F, MHES). Note: Listed as ‘threatened’ by MARN. DRYOPTERIDACEAE Arachniodes denticulata (Sw.) Ching, Acta Bot. Sin. 10: 260. 1962. Polypodium denticulatum Sw., Aspidium denticulatum (Sw.) Sw., A. jucundum Fee, Polystichum denticulatum (Sw.) J. Sm., Dryopteris denticulata (Sw.) Kuntze, D. denticulata f. jucunda (Fée) C. Chr., Rumohra denticulata (Sw.) Copel., Byrsopteris denticulata (Sw.) C.V. Morton Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Sonsonate: R. Seiler 607 (F, MHES). Santa Ana: A. Molina et al. 12497 (F); F. Wunderlin 543 (ITIC); G. Cerén & D. Rodriguez s.n. (JBL04653) (B, LAGU, MO); J. Monterrosa et al. 204 (AAU, B, BM, INB, LAGU, MEXU, MO); M.C. Carlson 953 (F); R. Seiler 126 (F, ITIC, MHES); 179 (MHES); 239 (F, MHES); V.M. Martinez s.n. (CMC01027) (B, LAGU, MO). Chalatenango: J. Tucker 1060 (F); R. Seiler 154 (F); 410 (F, ITIC, MHES); 423 (JBL00082) (LAGU, MHES); W. Berendsohn et al. 1573 (B, LAGU, MO). Morazan: J.L. Linares & A. Pool 2291 (JBLO1780) (LAGU); R. Seiler 1065 (F, MHES). > Bolbitis aliena (Sw.) Alston, Kew Bull. 1932: 310. 1932. Acrostichum alienum Sw., Gymnopteris aliena (Sw.) C. Presl, Leptochilus alienus (Sw.) C. Chir Global distribution: Mexico, Honduras, — Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivi Source: Calderon & Standley (1941): 31 a uptmetisiis alienus); Seiler (1982): 389; Berendsohn (1989) 1: 9. ¢ Bolbitis bernoullii (Kuhn ex Christ) Ching in Christensen, Index Filic., Suppl. 3: 47. Acrostichum bernoullii Kuhn ex Christ, Gymnopteris bernoullii (Kuhn ex Christ) Diels in Engler & Prantl, G tuerckheimii Christ, G donnell-smithii Christ, Leptochilus bernoullii (Kuhn ex Christ) C. Chr., L. tuerckheimii (Christ) C. Chr., L. donnell-smithii (Christ) C. Chr., Bolbitis donnell-smithii (Christ) Ching in C. Chr. Local names: helecho cola de leon. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: M. Sandoval & E. Sandoval s.n. (ISBO0801) (AAU, B, F, LAGU); M. Sandoval & E. Sandoval 299 (B, F, LAGU). La Libertad: W. Berendsohn ef al. 1454 (LAGU). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 156 Bolbitis portoricensis (Spreng.) Hennipman, Amer. Fern J. 65: 30. 1975. Acrostichum portoricensis Spreng., A. cladorrhizans Spreng., A. irregulare Liebm., Gymnopteris portoricensis (Spreng.) Fée, Leptochilus cladorrhizans (Spreng.) Maxon, Bolbitis cladorrhizans (Spreng.) Ching in Christensen . Local names: helecho colchon de leon, helecho salamandra, pescadillo. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1954, 2987 (B, BM, ITIC, LAGU, MO); A. Sermeijio s.n. (JBLO1118), 97 (JBLO1000) (B, LAGU, MO); E. Sandoval 1149 (B, LAGU); F. Chinchilla s.n. (ISB00314) (LAGU); J.M. Rosales 6 (AAU, B, BM, EAP, LAGU, MEXU, MO); 2450 (B, BM, LAGU, MEXU, MO); M. Sandoval & E. Sandoval 269 (B, F, LAGU); P. Standley er al. 2604 (F); R. Seiler 923 (MHES); T. Croat 42096 (ITIC); W. Berendsohn 1317 (B, LAGU, MO); 1724 (B, LAGU). Santa Ana: J. Monterrosa et al. 84 (B, LAGU, MO); M. Magafia 45 (MP- 00023) (LAGU); R. Seiler 706 (MHES). La Libertad: A.K. Monro et a/. 3771 (B, BM, ITIC, LAGU, MO); B. Pfeiffer-Berendsohn ert al. 67 (B, LAGU, MO): F. Wunderlin 460 (ITIC); K. Sidwell et al. 479 (B, BM, ITIC, LAGU, MO); M.A. Hernandez s.n. (JCG00562) (B, LAGU, MO); W. Heed 27, 28, 29 (ITIC). San Salvador: K. Sidwell er al. 569 (B, BM, ITIC, LAGU, MO); R. Seiler 314, 454, 458 (F, MHES). Chalatenango: R. Seiler 144 (F, MHES). Cabafias: A.K. Monro ef al. 3750 (B, BM, ITIC, LAGU, MO); R.A. Carballo et a/. 259 (LAGU). La Union: A.K. Monro et al. 3537 (B, BM, ITIC, LAGU, MO). Ctenitis equestris (Kunze) Ching var. equestris, Sunyatsenia 5: 250. 1940. Lastrea ciliata Liebm., Polypodium alsophiloides Liebm., Aspidium bourgeaui E. Fourn., A. scabriusculum Davenp., Drvopteris davenportii C. Chr., D. equestris (Kunze) C. Chr. var. mentiens C. Chr. Global distribution: Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2013 (B, BM, ITIC, LAGU, MO): G. Davidse et al. 37379 (B, BM, ITIC, LAGU, MO); R. Seiler 814, 906 (F, MHES); 954 (MHES): W. Berendsohn & E. Sandoval 1510 (B, LAGU). Sonsonate: A. Molina et al. 21630, 21763, 21775 (F, ITIC); 21770, 21773 (F); R. Seiler 612 (F, MHES). Santa Ana: A. Molina er a/. 12654, 21496 (F); 21513 (F, ITIC); D. Rodriguez & G. Cerén s.n. (JBL04604) (LAGU); J. Tucker 1216 (F); K. Sidwell er a/. 440 (B, BM, ITIC, LAGU, MO, L). Lara 227, 228 (MHES); M. Magafia 23 (MP-00010) (LAGU); R. Seiler 19 (MHES); 350 (F, MHES); S. Winkler 67 (ITIC); T. Croat 42232 (ITIC); V.M. Martinez s.n. (CMC00819) (B, LAGU, MO); s.n. (CMC01000) (B, EAP, LAGU, MO). San Salvador: M.C. Carlson 427, 477 (F); R. Seiler 208 (MHES); 209 (F, MHES). Chalatenango: R. Seiler 435b (JBL04943) (LAGU, MHES). San Vicente: R. Seiler 332 (F, MHES). Morazan: R. Seiler 1006 (F, MHES). @ Ctenitis equestris var. erosa Stolze, Amer. Fern. J. 67: 41. 1977. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R. Seiler 484, 690 (F, MHES). Ctenitis erinacea A.R. Sm.. Mem. New York Bot. Gard. 88: 223-224. 2004. Global distribution: Mexico, Guatemala, Honduras. 157 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: R. Seiler 890 (UC) (Mickel & Smith, 2004); Croat 42398 (CAS, MEXU) (Mickel & Smith, 2004). Santa Ana: B. Pfeiffer-Berendsohn 52 (B, LAGU, MO): J. Monterrosa & R. Carballo 287 (B, BM, LAGU); R. Seiler 136 (MHES); 312 (F); 479 (F, ITIC, MHES). Chalatenango: Tucker 1140 (LL, UC, US) (Mickel & Smith, 2004). Ctenitis excelsa (Desv.) Proctor, Rhodora 63: 34. 1961. Polypodium excelsum Desv., Dryopteris excelsa (Desv.) C. Chr. Local name: helecho colchon de leon. Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles. Salvadoran distribution. Anuachapan: A.K. Monro et al. 1945, 3078 (B, BM, ik. LAGU, MO); E. Sandoval 1148 (LAGU, MO); E. Sandoval & F. Chinchilla 488 (B, LAGU); R. Seiler 922 (F, MHES). Santa Ana: R. Seiler 879 (F). Ctenitis hemsleyana (Baker) Copel., Gen. Fil. 124. 1947. Polypodium hemsleyanum Baker ex Hemsl., Dryopteris hemsleyana (Baker ex Hemsl.) C. Chr., Ctenitis costaricensis R.C. Moran Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Morazan: R. Seiler 1024 (F, MHES). > Ctenitis interjecta (C. Chr.) Ching, Sunyatsenia 5: 250. 1940. Dryopteris interjecta C. Chr. Global distribution: Mexico, Guatemala, Belize, Honduras, Costa Rica. Source: Léetschert (1954): 27; Seiler (1982): 388; Berendsohn (1989) 1: 9. Ctenitis nigrovenia (Christ) Copel., Gen. Fil. 124. 1947. Nephrodium nigrovenium Christ in Donnell Smith, Dryopteris nigrovenia (Christ) C. Chr. Local name: helecho trecillo. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Peru, Bolivia. Salvadoran distribution. Ahuachapan: D. Witsberger 692, 655 (MHES); E. Sandoval 1135 (B, LAGU, MO); R. Seiler 817 (F, ITIC, MHES, NY). Ctenitis sloanei (Poepp. ex Spreng.) C.V. Morton, Amer. Fern J. 59: 66. 1969. Polypodium sloanei Poepp. ex Spreng. Global distribution: USA, Mexico, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia. Salvadoran distribution: Ahuachapan: N. Herrera s.n. (JBL03707) (LAGU, MO). San Salvador: A.K. Monro ef al. 3453 (B, BM, ITIC, LAGU, MO). Dryopteris karwinskyana (Mett.) Kuntze, Revis. Gen. Pl. 2: 813. 1891. Aspidium karwinskyanum Mett., Nephrodium karwinskyanum (Mett.) Baker in Hooker & Baker . Local name: helecho pata de paloma. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: J.M. Rosales 1063 (AAU, B, BM, LAGU, MO); M. Sandoval & E. Sandoval 69, 102 (B, F, LAGU); W. Berendsohn et al. 1642 (B, BM, LAGU, MO). La Libertad: N. Ventura 5010, 5024 (ITIC). San Salvador: K. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 158 Prestegard 34 (MP-00011) (LAGU); R. Seiler 70 (F, MHES): 438 (F, ITIC. MHES). Chalatenango: R. Seiler 522 (F); 554 (F, MHES). Cabaiias: R. Seiler 520 (F, MHES). Morazan: J. Monterrosa & C. Rivera 679 (BM, LAGU). > Dryopteris maxonii Underw. & C. Chr., Amer. Fern. J. 1: 96. 1911. Dryopteris patula (Sw.) Underw. var. moreliae C hrist in Lecomte Global distribution: Mexico, Guatemala, Costa Salvadoran distribution: R. Seiler 1485 (MO) ead in Davidse et al. 1995) Source: Moran in Davidse et al. (1995): 213. Dryopteris nubigena Maxon & C.V. Morton, Proc. Biol. Soc. Wash. 50: 179. 1937. Local names: helecho petatillo de montafia. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: Santa Ana: R. Seiler 691 (F, MHES). Chalatenango: W. Berendsohn et al. 1570 (B, LAGU, MO). Dryopteris patula (Sw.) Underw., Native Ferns, ed. 4, 117. 1893. Aspidium patulum Sw., A. paupertinum Kunze, A. mexicanum C. Presl var. obtusilobum Kunze ex Mett., A. apertum Fée, Dryopteris aperta (Fée) C. Chr., D. simplicior Mickel & Beitel Local names: helecho cola de — real, helecho pata de comadreja, helecho pata de paloma, helecho pluma de faisa Global distribution: Mexico, eieciie Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paragu Salvadoran distribution: Ahuachapan: E. Sandoval & M. Sandoval 1371 (B, LAGU); F. Chinchilla s.n. (ISB00538) (AAU, B, LAGU); s.n. (ISB00659) (AAU, B, F, LAGU); M. Sandoval & E. Sandoval 19 (B, LAGU); M. Sandoval 66 (B, F, LAGU). Sonsonate: R. Seiler 34, 611 (F, MHES); T. Croat 42245 (ITIC). Santa Ana: K. Prestegard | (MP- 00012) (LAGU); M.B. Siu s.n. (ITIC); R. Seiler 82 (MHES); 500 (F, ITIC, MHES); 660 (F, MHES). La Libertad: E.A. Montalvo & R. Villacorta 6268 (B, LAGU). San Salvador: R. Seiler 25, 52 (F, MHES); 437 (F, ITIC, MHES): 441 (MHES): S. Winkler 79 (ITIC). Chalatenango: R. Seiler 567 (JBL04812) (F, LAGU); R. Seiler 142, 568 (F. MHES). San Vicente: R. Seiler 318 (JBL04806) (F, LAGU, MHES). Morazan: J. Monterrosa & M. Rivera 643 (B, EAP, LAGU); J. Monterrosa & C. Rivera 648 (BM, LAGU): S. Winkler 84 (ITIC). Dryopteris wallichiana (Spreng.) Hyl., Bot. Not. 1953: 352. 1953. Aspidium paleaceum D. Don, A. wallichianum Spreng., A. donianum Spreng., A. paleaceum Sw., A. parallelogrammum Kunze, A. crinitum M. Martens & Galeotti, Dryopteris paleacea (Sw.) C. Chr., D. parallelogramma (Kunze) Alston Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Africa, Asia, Hawaii. Salvadoran distribution: Chalatenango: Hammer s.n. (12-274) (MHES); R. Seiler 163 (F, MHES). San Vicente: S. Winkler 77 (ITIC). > Elaphoglossum affine (M. Martens & Galeotti) T. Moore, Index Fil. 4. 1857. Acrostichum affine M. Martens & Galeotti . Global distribution. Mexico, Guatemala, Costa Rica, Panama, Colombia, Venezuela. 159 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: M.C. Carlson 730, F (Mickel, 1995: 256). Source: Mickel in Davidse et al. (1995): 256; Mickel & Smith (2004): 285. Elaphoglossum albomarginatum A.R. Sm., Proc. Calif. Acad. Sci. ser. 4, 40: 220 (F). Global distribution: Mexico, Guatemala. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2016 (B, BM, ITIC, LAGU, MO). Santa Ana: R. Seiler 876 (F, ITIC, MHES); S. Winkler 62 (ITIC). Elaphoglossum baquianorum A. Rojas, Rev. Biol. Trop. 51(1): 35. 2003. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. oo distribution: Santa Ana: R. Seiler 290 (F, MHES); R. Seiler 688 (F, MHES, Y). Elaphoglossum ellipticifolium A. Rojas, Rev. Biol. Trop. 51(1): 5. 2003. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 16855 (F); T. Croat 42423, holotipo MO, isotipo NY; R. Seiler 170 (F, MHES). Chalatenango: R. Seiler 397 (JBL00100) (F, LAGU, MHES); R. Seiler 429 (JBL00098), F (LAGU). Morazan: R. Seiler 992 (F, MHES). Elaphoglossum erinaceum (Fée) T. Moore var. erinaceum, Index Fil. 9. 1857. Acrostichum erinaceum Fée Global distribution: Mexico, Guatemala, Honduras, — Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Boliv Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar’ 839, 872 (MHES). Elaphoglossum eximium (Mett.) Christ, Monogr. Elaph. 107. 1899. Acrostichum eximium Mett. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: A. Molina et al. 16824; B. Pfeiffer-Berendsohn 35 (LAGU); J. Lagos s.n. (ITIC); R. Seiler 174 (F, MHES); S. Winkler 617 (ITIC). Usulutan: L. Gonzalez 1942 (ITIC). > Elaphoglossum glabellum J. Sm., London J. Bot. 1: 197. 1842. Acrostichum glabellum (J. Sm.) Klotzsch Global distribution: Mexico, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, Ecuador, Peru, Bolivia Salvadoran distribution: Ahuachapan: P. Standley 20172 (US). Source: Mickel & Smith (2004): 290. Elaphoglossum glaucum T. Moore, Index. Fil. 10. 1857, Acrostichum glaucum Fée, Elaphoglossum eucraspedum Christ Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R. Seiler 668 (MHES). Chalatenango: R. Seiler 399 (JBLO0101) (F, LAGU, MHES) MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 160 Elaphoglossum guatemalense (Klotzsch) T. Moore, Parker's Cat. 1858. Acrostichum guatemalense Klotzsch, A. salvinii Baker ex Hemsl. Global distribution: Mexico, Guatemala, Belize, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 16868 (F). Chalatenango: R. Seiler 396 (F, MHES). San Vicente: R. Seiler 321 (F, MHES). Morazan: R. Seiler 1066 (MHES). Elaphoglossum herrerae A. Rojas, Brenesia 45-46: 13. 1996. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: Morazan: R. Seiler 1080 (F, ITIC, MHES). Elaphoglossum lanceum Mickel, Amer. Fern J. 69: 101. 1979. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R.A. Carballo 649 (B, LAGU, MO); R. Seiler 104, 173 (F, MHES); 684 (MHES). Chalatenango: R. Seiler 161, 739 (F, MHES). Elaphoglossum latifolium (Sw.) J. Sm., J. Bot. 1: 197. 1842. Acrostichum latifolium Liebm., A. sartori Liebm., A. scapellum Kunze ex Fée, Elaphoglossum latifolium (Sw.) J. Sm. var. lutescens Rosenst., E. sartori (Liebm.) Mickel, E. scapellum (Kunze ex Fée) T. Moore Global distribution: Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia, Brazil. Salvadoran distribution: Santa Ana: G. Cerén & D. Rodriguez s.n. (JBL04645) (LAGU); G. Davidse et al. 37290 (B, BM, ITIC, LAGU, MO); J.L. Linares & C.A. Martinez 1520 (JBL01904) (LAGU); J. Monterrosa & R. Carballo 859 (LAGU); J. Monterrosa et al. 174, 179 (LAGU); R. Seiler 103, 175, 657 (MHES); R. Villacorta ef al. 1286, 1018 (B, LAGU, MO). Chalatenango: W. Berendsohn er al. 1596 (B, LAGU); R. Seiler 150, 749; 753 (F, MHES). Note: Rojas-Alvarado (2003) notes that Elaphoglossum latifolium is taxonomically difficult because of variation in rhizome habit and scales, frond scales, size, apex and base shape. Rojas-Alvarado (2003) and Mickel & Smith (2004) suggests that E. latifolium sensu stricto may be restricted to the Caribbean islands. It would seem probable that the collections cited here as E. /atifolium include more than one species. ¢ Elaphoglossum lonchophyllum (Fée) T. Moore, Index. Fil. 360. 1862. Acrostichum lonchophyllum Fée, Elaphoglossum acutissimum Christ Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 28 (LAGU, MO). > Elaphoglossum longistipitatum A. Rojas, Brenesia 47-48: 6. 1997. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: Santa Ana: W. Léetschert 375 (US). Source: Rojas-Alvarado (2002): 997. Elaphoglossum mevaughii Mickel, Brittonia 32: 336. 1980. Global distribution: Mexico, Guatemala. 161 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: Santa Ana: J. Monterrosa et al. 1011 (B, LAGU, MO, NY). San Vicente: R. Seiler 322 (F, MHES). Elaphoglossum mesoamericanum A. Rojas, Rev. Biol. Trop. 51(1): 8. 2003. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: M. Carlson 753 (F). San Salvador: M. Carlson 593 (F). Chalatenango: R. Seiler 258 (F, MHES); S. Calderén 2448 (F). Note: Mickel & Smith (2004) cite Seiler 578 as Elaphoglossum yourkeorum (see below), this specimen may be conspecific to E. mesoamericanum but the two species have been maintained distinct here pending further research. Elaphoglossum mexicanum (E. Fourn.) A. Rojas, Rev. Biol. Trop. 51(1): 42. 2003. Acrostichum mexicanum E., Fourn. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R. Seiler 130 (F, MHES); W. Léetschert 369, US (Rojas-Alvarado, 2003). ® Elaphoglossum microproductum A. Rojas, Brenesia 47-48: 9. 1997. Global distribution: Endemic. Salvadoran distribution: Chalatenango: R. Seiler 742 (MHES, NY). @ Elaphoglossum minutum (Pohl ex Fée) T. Moore, Index Fil. 12. 1857. Acrostichum minutum Pohl ex Fée, A. conforme Sw. var. alpinum J. Bommer ex H. Christ?, A. leptophyllum Fée, Elaphoglossum leptophyllum (Fée) T. Moore Global distribution: Costa Rica, Panama, Antillas, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Brazil. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2018 (B, BM, ITIC, LAGU, MO). Morazan: R. Seiler 1016 (F, MHES). Elaphoglossum muscosum (Sw.) T. Moore, Index. Fil. 12. 1857. Acrostichum muscosum Sw., Elaphoglossum hookerianum Underw. ex Maxon Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru. Salvadoran distribution: Sonsonate: R. Seiler 601 (MHES). Santa Ana: B. Pfeiffer- Berendsohn & W. Berendsohn 4, 24 (LAGU, MO): MLL. Reyna de Aguilar 665 (MHES); R. Seiler 90, 354b, 653 (MHES). Chalatenango: R. Seiler 428 (JBL00097) (LAGU, MHES); R. Seiler 740 (MHES). Morazan: R. Seiler 994 (F, MHES). Elaphoglossum paleaceum (Hook. & Grev.) Sledge, Bull. Brit. Mus. (Nat. Hist.) Bot. 4: 95. 1967. Acrostichum paleaceum Hook. & Grev., A. squamosum Sw., Elaphoglossum squamosum J. Sm., E. hirtum sensu auct., non Acrostichum hirtum Sw. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Hawaii. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Carballo 857 (B, LAGU); R. Seiler 291, 645 (F, MHES); S. Winkler 55 (ITIC). San Salvador: R. Seiler 337 (MHES). Chalatenango: R. Seiler 403 (JBL00099) (F, LAGU, MHES): 741 (MHES). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 162 Elaphoglossum peltatum (Sw.) Urb., Symb. Antill. 4: 60. 1903. Osmunda peltata Sw., Acrostichum peltatum (Sw.) Sw., Rhipidopteris peltata (Sw.) Schott ex Fée, Peltapteris peltata (Sw.) C.V. Morton . Local names: sombrillita. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Bolivia. Salvadoran distribution: Sonsonate: R. Seiler 597 (F, MHES). Santa Ana: A. Molina et al. 16754, 16850 (F; B). Pfeiffer-Berendsohn & W. Berendsohn 27 (B, F, LAGU, MO); G. Davidse et al. 37298 (B, BM, ITIC, LAGU, MO); K. Sidwell er al. 846 (B, BM, ITIC, LAGU, MO); M.C. Carlson 882 (F); O. Rohweder s.n. (JBL01258), 2649 (JBL01243) (LAGU); R. Seiler 97 (F, ITIC, MHES); 868 (MHES). Morazan: R. Seiler 993 (F, MHES). Elaphoglossum petiolatum (Sw.) Urb., Symb. Antill. 4: 61. 1903. Acrostichum petiolatum Sw., A. viscosum Sw., A. schiedei Kunze, A. schmitzii Mett. ex Kuhn, Elaphoglossum viscosum (Sw.) J. Sm. Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil. Salvadoran distribution. Santa Ana: G. Davidse et al. 37303 (B, BM, ITIC, LAGU, MO). Morazan: R. Seiler 1029 (F, ITIC, NY). Elaphoglossum piloselloides (C. Pres|) T. Moore, Index Fil. 13. 1857. Acrostichum piloselloides C. Presl Global distribution. Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil. Bolivia, Chile. Salvadoran distribution: Santa Ana: J.L. Linares & C.A. Martinez 1519 (JBL01916) (LAGU). Elaphoglossum sartorii (Liebm.) Mickel. Mem. New York Bot. Gard. 46: 181. 1988. Acrostichum sartorii Liebm., A. scapellum Kunze ex Fee, Elaphoglossum scapellum (Kunze ex Fée) T. Moore, E. subarborescens Rosenst. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: San Vicente: R. Seiler 327 (F, MHES). Elaphoglossum setosum (Liebm.) T. Moore, Index Fil. 366. 1862. Acrostichum setosum Liebm., A. recognitum Kunze ex E. Fourn., Elaphoglossum catharinae Underw. ex Maxon Global distribution. Mexico, Guatemala, Nicaragua, Costa Rica. Salvadoran distribution. Chalatenango: J. Monterrosa ef al. 1619 (BM, LAGU, MHES, NY). Elaphoglossum squamipes (Hook.) T. Moore, Index Fil. 15. 1857. Acrostichum squamipes Hook., A. ovatum Liebm., Elaphoglossum subaequale T. Moore Global distribution. Mexic na Venezuela, Ecuador, Peru, Brazil, Bolivia. o, Guatemala, Costa Rica, Panama, Antilles, Colombia, 163 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: Chalatenango: R. Seiler 425 (JBL00096) (LAGU); R. Seiler 149 (F, MHES); 745 (MHES). Elaphoglossum tenuiculum (Fée) T. Moore ex Baker, Ann. Bot. (Oxford) 5: 491. 1891. Acrostichum tenuiculum Fée, Elaphoglossum stellatum Mickel Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: San Salvador: R. Seiler 31, 54, 515 (MHES). Chalatenango: W. Heed 31 (ITIC). Elaphoglossum yourkeorum Mickel, Mem. New York Bot. Gard. 88: 314-315. 2004. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Chalatenango: R. Seiler 578 (F, MHES, NY). ¢ Lastreopsis effusa te divergens (Willd.) Tindale, Contr. New South Wales Natl. Herb. 3: 299, pl. 21. 1965. Polypodium ee Willd. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Santa Ana: J.Monterrosa, D.Rodriguez & E.Pérez 1553 (BM, LAGU). @ Megalastrum pulverulentum (Poir.) A. R. Sm. & R. C. Moran, Amer. Fern. J. 77: 129. 1987. Ctenitis pulverulenta (Poir.) Copel., C. karsteniana (Klotzsch) Vareschi, C. pulverulenta (Poir.) Copel. var. heydei (C. Chr.) Stolze, Polypodium pulverulentum Poir. in Lamarck, P. karstenianum Klotzsch, Dryopteris pulverulenta (Poir.) C. Chr. in Urban, D. karsteniana (Klotzsch) Hieron, D. karsteniana (Klotzsch) Hieron var. heydei C. Chr., Megalastrum pulverulentum (Poir.) A. R. Sm. & R. C. Moran var. heydei (C. Chr.) A.R. Sm. & R.C. Moran Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Martinez 944 (LAGU); J. Monterrosa et al. 961 (LAGU, NY). ¢ Olfersia cervina (L.) Kunze, Flora 7: 312. 1824. Global distribution: Mexico, Guatemala, Belice, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Cabafias: G. Cerén 1788 (LAGU, MEXU, MHES); J. Monterrosa & D. Rodriguez 1633 (BM, LAGU, MHES. MO). rhs a juglandifolia (Humb. & Bonpl. ex Willd.) J. Sm., J. Bot. (Hooker) 4: 187. 1841. _ Juglandifolium Humb. & Bonpl. ex Willd., Cyrtomium juglandifolium (Humb. & Bonpl. ex Willd.) T. Moore Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 164 Colombia, Venezuela. Salvadoran distribution: Tucker 1288, US (Yatskievych in Davidse ef al. 1995). Sonsonate: J.L. Linares & C.A. Martinez 3045 (JBLO1790) (LAGU). Phanerophlebia macrosora (Baker) Underw., Bull. Torrey Bot. Club 26: 213. 1899. Aspidium juglandifolium (Humb. & Bonpl. ex Willd.) Kunze ex Klotzsch var. macrosorum Baker, Cyrtomium macrosorum (Baker) C.V. Morton, C. guatemalensis (Underw.) C.V. Morton, Phanerophlebia sneelatnacal Underw. Global distribution: Mexico, Guatemala, Costa Rica, P Salvadoran distribution: Chalatenango: R. Seiler Te ELDOOe ee LAGU, MHES). Polystichum alfaroi (Christ) Barrington, Rhodora 94: 328. 1992. Polystichum aculeatum (L.) Roth var. alfaroi Christ Global distribution: Mexico, Guatemala, Costa Rica, Panama. Salvadoran distribution: Sin loc: P. Standley 21567, NY (Mickel & Smith, 2004: 514). Ahuachapan: J. Monterrosa & D. Rodriguez 1434 (B, BM, LAGU, MO, NY). Polystichum distans E. Fourn., Mexic. PI. I: 91. 1872. Polystichum pallidum E. Fourn. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: R. Seiler 12, 18, 504 (MHES). San Salvador: R. Seiler 207 (MHES). Chalatenango: R. Seiler 381 (MHES). San Vicente: R. Seiler 315 (MHES). Polystichum fournieri A. R. Sm., Amer. Fern J. 70: 27. 1980. Polystichum muelleri Mett. ex E. Fourn. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: Sonsonate: R. Seiler 613 (MHES). Santa Ana: M.L. Reyna de Aguilar 667 (MHES); R. Seiler 181, 461 (MHES). Chalatenango: W. Berendsohn et al. 1575 (B, LAGU). Polystichum hartwegii (Klotzsch) Hieron., Hedwigia 46: 355. 1907. Aspidium hartwegii Klotzsch, A. trejoi Christ, Polystichum grande Fee, P. drepanoides E. Fourn., P. trejoi (Christ) Christ Local name: helecho trenza de sanson. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia?, Venezuela, Ecuador, Bolivia. Salvadoran distribution: Ahuachapan: D. Witsberger 654 (MHES); F. Chinchilla & E. Lopez s.n. (ISB00311) (LAGU); N. Herrera s.n. (JBL03863) (B, EAP, LAGU, MO). Santa Ana: D. Rodriguez & G. Cerén s.n. (JBL04613) (LAGU); J. Monterrosa ef al. 208 (B, LAGU): J. Monterrosa & R. Martinez 940 (LAGU, NY); 948 (B, LAGU, NY); K. Sidwell et al. 833 (B, BM, ITIC, LAGU, MO); R. Seiler 293, 345 (MHES); V.M. Martinez s.n. (CMC00845) (B, LAGU); s.n. (CMC00880) (B, LAGU, MO); s.n. (CMC00879), s.n. (CMC00897), s.n. (CMC00972) (B, EAP, LAGU, MO); W. Berendsohn er al. 1541 (B, LAGU, MO). Usulutan: A.K. Monro er al, 3007 (B, BM, ITIC, LAGU, MO). Morazan: R. Seiler 1011 (MHES); R. Seiler 1033 (JBL04820) (LAGU, MHES). 165 FERN GAZ. 18(4): 120-215. 2008 > Polystichum mickelii A. R. Sm., Amer. Fern. J. 70: 25. 1980. Global distribution: Mexico, Guatemala, Honduras. le distribution: P.Standley 20135, GH (Barrington in Davidse ef al., 1995: 222): Sources: Barrington in Davidse er al. (1995): 222; Mickel & Smith (2004): 518. ¢ Polystichum muenchii (Christ) C. Chr., Index Fil. 585. 1906. Aspidium muenchii Christ Global distribution: Mexico, Guatemala. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2009 (B, BM, ITIC, LAGU, Polystichum muricatum (L.) Fée, Mém. Foug. 5: 278. 1852. Polvpodium muricatum L Local names: helecho pata de leén, helecho rosquilla de altura, helecho trenza de Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Venezuela Salvadoran distribution: Ahuachapan: E. Sandoval 1134 (B, LAGU); F. Chinchilla ef al. s.n. (ISB00091) (B, LAGU, MO); M. Sandoval & E. Sandoval 92 (B, F, LAGU); R. Villacorta et al. 906 (B, LAGU, MO).Usulutan: D. Williams 58 (LAGU, MO). Polystichum turrialbae Christ, Bull. Herb. Boissier, sér. 2, 6: 163. 1906. Polystichum smithii Mickel & Beitel Local name: helecho trenza de sanson Global distribution: Mexico, Costa Rica, Panama, Bolivia. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1970, 2010 (B, BM, ITIC, LAGU, MO); W. Berendsohn ef al. 1427 (B, LAGU, MO). EQUISETACEAE Equisetum giganteum L., Syst. Nat. ed. 10, 2: 1318. 1759. Local name: cola de caballo. Global distribution: Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Uruguay, Argentina, Chile. Salvadoran distribution: Santa Ana: M.B. Siu s.n. (ITIC); R. Villacorta & H. Sipman 1260 (B, LAGU, MO); V.M. Martinez s.n. (CMC00048) (B, BM, LAGU, MEXU, MO); V.M. Rosales 10 (ITIC). Chalatenango: B. Lopez s.n. (ITIC); E.A. Montalvo 5435 (ITIC); R. Seiler 255 (F, ITIC, MHES). Equisetum hyemale A. Braun ex Engelm. L. var. affine (Engelm.) A. A. Eaton, Fern Bull. I: III. 1903. Local name: cola de caballo. Equisetum robustum A. Braun ex Engelm., E. robustum A. Braun ex Engelm. y affine Engelm., E. hyemale L. subsp. affine (Engelm.) Calder & Roy Taylor Global distribution: Canada, USA, Mexico, Guatemala, Asia. Salvadoran distribution: La Libertad: M. Renderos & R. Villacorta 291 (B, LAGU, MO). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 166 Equisetum myriochaetum Schltdl. & Cham., Linnea 5: 623. 1830. Equisetum mexicanum Milde Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru. Salvadoran distribution: Santa Ana: R. Villacorta et al. 1049 (B, LAGU, MO). Chalatenango: E.A. Montalvo 5279 (ITIC). Morazan: M. Renderos ef a/. 628 (B, BM, LAGU, MO). Equisetum x schaffneri Milde, Verh. Zool.-Bot. Ges. Wien II: 345. 1861. Global distribution: Mexico, Guatemala a Peru. Salvadoran distribution: S. Calderon 1864 (F, ITIC, US). Note: Equisetum x schaffneri is believed to be a hybrid between E. giganteum and E. myriochaetum (Hauke, 1995: 326). GLEICHENIACEAE Dicranopteris flexuosa (Schrad.) Underw., Bull. Torrey Bot. Club 34: 254. 1907. Mertensia flexuosa Schrad., Gleichenia flexuosa (Schrad.) Mett. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution. La Libertad: M.C. Carlson 251 (F). Usulutan: D. Williams 141 (LAGU, MO); J. Lagos 1777 (ITIC). Morazan: K. Sidwell et al. 736 (B, BM, ITIC, LAGU, MO); R. Seiler 1049b (F, ITIC, MHES); R. Villacorta er al. (B, LAGU). Diplopterygium bancroftii (Hook.) A. R. Sm., Amer. Fern. J. 70: 26. 1980. Gleichenia bancroftii Hook., Mertensia bancroftii (Hook.) Kunze, M._ bancroftii (Hook.) Kunze var. vitellina Kunze, Dicranopteris bancroftii (Hook.) Underw., Hicriopteris bancroftii (Hook.) Ching Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution. Santa Ana: J. Tucker 1184, F, K; Puller s.n. (12-548) (MHES); R. Seiler 238 (F, ITIC, MHES); S. Winkler 68 (ITIC). Gleichenella pectinata (Willd.) Ching, Sunyatsenia 5: 276. 1940. . . Mertensia pectinata Willd., M. glaucescens Humb. & Bonpl. ex Willd. var. mexicana Fée, Gleichenia pectinata (Willd.) C. Presl, G. nitida C. Presl, Dicranopteris pectinata (Willd.) Underw. ; Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution. Santa Ana: A. Molina et al. 12577 (F); R. Seiler 460 (F, MHES): 1199 (F). Chalatenango: E.A. Montalvo s.n. (ITIC); M.C. Carlson 590 (F); R. Villacorta & A. Araniva 968 (B, LAGU, MO). Morazan: R. Seiler 983 (F, MHES). Sticherus bifidus (Willd.) Ching, Sunyatsenia 5: 282. 1940. 7 — Mertensia bifida Willd., M. fulva Desv., M- gleichenioides Liebm., Gleichenia bifida (Willd.) Spreng., G liebmannii T. Moore, Dicranopteris bifida (Willd.) Maxon, D. fulva 167 FERN GAZ. 18(4): 120-215. 2008 (Desv.) Underw. Local name: helecho cien pié. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval & M. Sandoval 1375 (B, LAGU); J.M. Rosales 1294 (B, LAGU, MO). Santa Ana: H. Weber 293 (ITIC); M.C. Carlson 997 (F). La Libertad: M.A. Hernandez s.n. (JCG00590) (B, LAGU, MO). Morazan: J. Tucker 792 (F). Sticherus brevipubis (Christ) A. R. Sm., Amer. Fern J. 70: 27. 1980. Gleichenia brevipubis Christ Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: J.L. Linares 482 (JBL01980) (LAGU). Santa Ana: A. Molina et al. 12520 (F); G. Cerén & D. Rodriguez s.n. (JBL04661) (LAGU); G. Davidse et al. 37309 (B, BM, ITIC, LAGU, MO); M.L. Reyna de Aguilar 682 (MHES); R. Villacorta et al. 1065 (B, ITIC, LAGU, MO). Chalatenango: J. Tucker 1183 (F); R. Seiler 725 (F, MHES). Morazan: R. Seiler 989 (F, MHES). Sticherus palmatus (W. Schaffn. ex E. Fourn.) Copel., Gen. Fil. 28. 1947. Mertensia palmata W. Schaffn. ex Fée, M. palmata W. Schaffn. ex E. Fourn., Gleichenia palmata T. Moore, G palmata (W. Schaffn. ex E. Fourn.) C. Chr., Dicranopteris palmata J. H. Schaffn. ex Underw. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Carballo 273 (B, BM, LAGU, MO); R. Seiler 233 (F, ITIC, MHES); 241 (ITIC). Sticherus underwoodianus (Maxon) Nakai, Bull. Natl. Sci. Mus. Tokyo 29: 31. 1950. Dicranopteris underwoodiana Maxon, Gleichenia underwoodiana (Maxon) C. Chr. Local names: helecho de mano, helecho pié de gallina. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 16747 (F; B). Pfeiffer- Berendsohn 38 (B, LAGU, MO); B. Lopez s.n. (ITIC); G. Davidse et al. 37269 (B, BM, ITIC, LAGU, MO); R. Seiler 195 (MHES); 230 (ITIC, MHES); 251 (F, ITIC, MHES); W. Berendsohn et al. 1547 (B, LAGU). Chalatenango: R. Seiler 726 (F, MHES). HYMENOPHYLLACEAE Hymenophyllum crassipetiolatum Stolze, Amer. Fern J. 66: 15. f. I. 1976. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 12638, 16767, 16811 F; 12488, 12504 (F, ITIC); M.C. Carlson 894 (F); R. Seiler 102 (F, MHES); 687, 876 (MHES):; R. Villacorta et al. 1023 (B, LAGU, MO); W. Berendsohn et al. 1698 (B, LAGU, MO). Chalatenango: J. Tucker 1062a. Morazan: J.L. Linares & A. Pool 2296 (JBL01929) (LAGU) MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 168 ¢ Hymenophyllum crispum Kunth, Nov. Gen. Sp. |: 26. 1815. Sphaerocionium crispum (Kunth) Klotzesh, S. schiedeanum C. Presl. Hymenophyllum schiedeanum (C. Presl) Bosch Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Antilles, Colombia. Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Chalatenango: J.Monterrosa ef al. 1618 (BM, LAGU, Hymenophyllum fucoides (Sw.) Sw., J. Bot. (Schrader) 1800(2): 99. 1801. Trichomanes fucoides Sw., Meringium fucoides (Sw.) Copel. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 22 (LAGU, MO); 31 (B, LAGU, MO); M.L. Reyna de Aguilar 877 (MHES); R. Seiler 284a (ITIC, MHES); 295 (F, MHES); 297 (F, ITIC, MHES). Chalatenango: R. Seiler 433 (F). Morazan: R. Seiler 996 (F, ITIC, MHES). > Hymenophyllum microcarpum Desv., Mém. Soc. Linn. Paris 6: 333. 1827. Sphaerocionium microcarpum (Desv.) Copel., Hymenophyllum angustifrons Christ, H. microcarpum Desv. var. lanceolatum C. V. Morton Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Dominica, Colombia, Venezuela, Guayana, Ecuador, Peru, Brazil, Bolivia. Source: Stolze (1976): 66; Berendsohn (1989) 1: 13; Pacheco in Davidse et al. (1995): 68; Mickel & Smith (2004): 344. Hymenophyllum myriocarpum Hook., Sp. Fil. 1: 106, t. 37D. 1844. Mecodium myriocarpum (Hook.) Copel., M. nigricans (C. Presl ex Klotzsch) Copel.. Hymenophyllum nigrescens Liebm., Sphaerocionium nigricans C. Presl ex Klotzsch Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru. Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 285 (F, ITIC, MHES); R. Villacorta 1119 (B, LAGU, MO); S. Winkler 92 (ITIC). Chalatenango: R. Seiler 434 (JBL00105) (F. ITIC, LAGU, MHES). San Vicente: R. Seiler 334 (F, ITIC, MHES). Morazan: R. Seiler 995 (F, ITIC, MHES). Hymenophyllum polyanthos (Sw.) Sw., J. Bot. (Schrader) 1800(2): 102. 1801. Trichomanes polyanthos Sw., Mecodium polvanthos (Sw.) Copel., Hymenophyllum Jalapense Schltdl. & Cham., H. millefolium Schltdl. & Cham., H. botrvoides Bosch Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Africa, Asia. ae Salvadoran distribution: Sonsonate: A. Molina et al. 21767 (F. ITIC); R. Seiler 599 (F, MHES). Santa Ana: A. Molina e¢ al. 12629, 12661 (F, ITIC); 12685 F; G. Davidse er al. 37215 (B, BM, ITIC, LAGU, MO); R. Seiler 282 (F, MHES). Morazan: R. Seiler 988 (F). 169 FERN GAZ. 18(4): 120-215. 2008 > Hymenophyllum pulchellum Schltdl. & Cham., Linnea 5: 618. 1830. Sphaerocionium pulchellum (Schltdl. & Cham.) C. Presl, Hymenophyllum pannosum Christ Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, razil. Source: Seiler (1982): 384; Berendsohn (1989) 1: 13; Mickel & Smith (2004): 345. Hymenophyllum tegularis (Desv.) Proctor & Lourteig, Bradea 5: 385. 1990. Davallia tegularis Desv., Hymenophyllum elegantulum Bosch, Sphaerocionium elegantulum (Bosch) Copel. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: A. Molina et al. 12505 (F, ITIC); 16813 (F); R. Seiler 231 (F, ITIC, MHES); 683 (MHES); S. Winkler 91 (ITIC). Chalatenango: R. Seiler 435a (JBL00106) (F, ITIC, LAGU, MHES). Hymenophyllum tunbrigense (L.) Sm., Mém. Acad. Roy. Sci. (Turin) 5: 418. 1793. richomanes tunbrigense L., Hymenophyllum fucoides (Sw.) Sw. var. frigidum Liebm. Global distribution. USA, Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Venezuela, Ecuador, Bolivia, Chile, Argentina; Europe, Africa, Australia. Salvadoran distribution: Santa Ana: R. Seiler 125 (F, ITIC, MHES); 284 (F, MHES). Chalatenango: R. Seiler 432 (F, MHES). Trichomanes capillaceum L., Sp. P|. 2: 1099. 1753. Vandenboschia capillacea (L.) Copel., Trichomanes tenellum Hedw., T. trichoideum Sw., T: trichoides Sw., T. schiedeanum Miill., T. hypnoides Christ Global distribution: Mexico, Guatemala, Belize, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru. Salvadoran distribution: Santa Ana: A. Molina et a/. 16870 (F); E.A. Montalvo 4152 (ITIC); G. Davidse er al. 37305 (B, BM, ITIC, LAGU, MO); R. Seiler 188, 286 (F, ITIC, MHES); R. Villacorta et a/. 1277 (B, ITIC, LAGU, MO); S. Winkler 90 (ITIC). Morazan: R. Seiler 1040 (F, MHES). Trichomanes diaphanum Kunth, Nov. Gen. Sp. 1: 25. 1816. Trichomanes bradei Christ Global distribution: Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Brazil. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 731 (B, LAGU). Santa Ana: A. Molina et al. 16804 (F); R. Seiler 250 (F, MHES): 303 (F, ITIC, MHES). Usulutan: D. Williams 247 (LAGU). Trichomanes hymenoides Hedw., Fil. Gen. Sp., t. 3. 1799. Trichomanes muscoides Sw., T. apodum Hook., Didymoglossum hymenoides (Hedwig) opel., Microgonium schaffneri Bosch ex Fée Global distribution: Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia. Argentina, Uruguay. Salvadoran distribution: Ahuachapan: R. Seiler 964 (F, MHES). Santa Ana: R. Seiler MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 170 302 (F, MHES); 357 (F, ITIC, MHES); S. Winkler 64 (ITIC). ¢ Trichomanes hymenophylloides Bosch, Ned. Kruidk. Arch. 5: 209. 1863. Trichomanes leptophyllum Bosch, Vandenboschia hymenophylloides (Bosch) Copel. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guyana, Suriname, French Guyana, Ecuador, Brazil. Salvadoran distribution: Santa Ana: J.Monterrosa & D.Rodriguez 1586 (BM, LAGU, MO) Trichomanes krausii Hook. & Grev., Icon. Filic. t. 149. 1831. Didymoglossum krausii (Hook. & Grev.) C. Presl, Hemiphlebium krausii (Hook. & Grev.) Prantl Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Local names: helecho musgo de pimiento Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 685 (B, LAGU). Santa Ana: S. Winkler 94 (ITIC). Morazan: R. Seiler 1020 (F, MHES). Trichomanes petersii A. Gray, Amer. J. Sci. Arts 15: 326. 1853. Microgonium petersii (A. Gray) Bosch, Hemiphlebium petersii (A. Gray) Prantl, Didymoglossum petersii (A. Gray) Copel., Trichomanes schaffneri Schltdl. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Chalatenango: R. Seiler 390 (JBL00107) (F, ITIC, LAGU, HES). Trichomanes radicans Sw., J. Bot. (Schrader) 1800(2): 97. 1801. Trichomanes scandens Hedw., T. kunzeanum Hook., T. Mexicanum Bosch, Vandenboschia radicans (Sw.) Copel. Local names: helecho plumajillo, helecho pluma de quetzal. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Europe, Asia, Africa. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1982, 1985 (B, BM, ITIC, LAGU, MO); E. Sandoval & F. Chinchilla 44, 518 (B, LAGU); J.G. Sandoval et al. s.n. (ISB00026) (B, LAGU, MO); M. Sandoval & E. Sandoval s.n. (ISB00799) (AAU, F, LAGU); R. Seiler 812 (F, ITIC, MHES); 958 (F, MHES): W. Berendsohn er al. 1363 (B, BG, LAGU, MO). Sonsonate: A. Molina ef al. 21771 (F): J.L. Linares & C.A. Martinez 3035 (JBLO1905) (LAGU); R. Seiler 618 (F, ITIC, MHES). Santa Ana: A. Molina et al. 12648, 16905 (F); J. Monterrosa et al. 172 (B, BM, LAGU, MO); N. Lopez et al. s.n. (JBL04627) (LAGU); R. Seiler 116, 364 (F, ITIC, MHES); R. Villacorta & M. Calderon 368 (B, LAGU, MO); S. Winkler 93 (ITIC):; W. Berendsohn et al. 1542 (B, LAGU, MO). San Salvador: R. Seiler 341 (F, MHES). Chalatenango: J. Tucker 1217 (F). Trichomanes reptans Sw., Prodr. 136. 1788. Didymoglossum reptans (Sw.) C. Presl, Hemiphlebium reptans (Sw.) Prantl 171 FERN GAZ. 18(4): 120-215. 2008 Global distribution. Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, ae Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argen Salvadoran distribution: Ahuachapan: E. Sandoval & . Chinchilla 576 (B, LAGU); G. Davidse et al. 37423 (B, BM, ITIC, LAGU, MO). Sonsonate: R. Seiler 598 (F, MHES). Santa Ana: S. Winkler 95 (ITIC). Chalatenango: R. Seiler 391 (F, MHES). Usulutan: A.K. Monro et a/. 3005 (B, BM, ITIC, LAGU, > Trichomanes robinsonii Baker, J. Linn. Soc. Bot. 9: 339. 1866. Global distribution: Honduras Source: Léetschert (1954): 25; Berendsohn (1989) |: LINDSAEACEAE Lindsaea stricta (Sw.) Dryand. var. stricta, Trans. Linn. Soc. London 3: 42. 1797. Adiantum strictum Sw., Lindsaea portoricensis Desv., L. microphylla C. Presl, L. multifrondosa Fée, L. moritziana Klotzsch Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Morazan: A.K. Monro et al. 3856 (B, BM, ITIC, LAGU, MO); K. Sidwell et al. 733, 734 (B, BM, ITIC, LAGU, MO); M. Renderos ef al. 605 (AAU, B, LAGU, MO); R. Seiler 1073 (F, MHES). Note: Seiler (1982) and Berendsohn (1989) treat L. stricta and L. portoricensis as distinct species. > Lonchitis hirsutus L., Sp. Pl. 2: 1078. 1753. Anisosorus hirsutus (L.) Maxon, Pteris lacinata Willd. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Suriname, Ecuador, Peru, SE Brazil, Bolivia. Source: Seiler (1982): 386; Berendsohn (1989) 1: 8. LOMARIOPSIDACEAE Cyclopeltis semicordata (Sw.) J. Sm., Bot. Mag. 72 (Compendium): 36. 1846. Polypodium semicordatum Sw., Aspidium semicordatum (Sw.) Sw., Dryopteris semicordata (Sw.) Kuntze, Polypodium semicordatum (Sw.) T. Moore Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: J.M. Rosales 488 (AAU, B, BM, LAGU, MO): R. Seiler 925 (F, MHES). Sonsonate: D. Rodriguez & M. Trejo 99 (LAGU). San Miguel: A. Salgado 70 (F, MHES); J. Tucker 925 (F). Nephrolepis biserrata (Sw.) Schott, Gen. Fil., t. 3. 1834. Aspidium biserratum Sw., Nephrolepis mollis Rosenst. Global distribution: USA, Mexico, Guatemala, Belize, Honduras. Nicaragua, Costa MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 172 Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Africa, Asia, Polynesia. Local names: helecho cola de pescado Salvadoran distribution: Ahuachapan: M. Magafia 17 (MP-00030) (LAGU). La Libertad: R. Seiler 1046 (F, MHES). Chalatenango: L. Lara 211 (MHES). Nephrolepis cordifolia (L.) C. Presl, Suppl. Tent. Pterid. 79. 1836. Polypodium cordifolium L Local names: centavito, helecho cola de ardilla. Global distribution: USA, Mexico, Guatemala, Honduras, Panama, Antilles, Peru, Brazil, Tropicos del Viejo Mundo. Salvadoran distribution: La Libertad: K. Prestegard 47 (MP-00028) (LAGU); M. Magafia 46 (MP-00027) (LAGU). San Salvador: R. Seiler 509 (F, MHES); Vaquerano s.n. (12-923) (MHES). Nephrolepis hirsutula (G. Forst.) C. Presl, Suppl. Tent. Pterid. 79. 1836. Polypodium hirsutulum G. Forst., Davallia multiflora Roxb., Nephrolepis multiflora (Roxb.) Jarret ex C.V. Morton Local name: helecho cola de quetzal. Global distribution: USA, Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Ecuador, Peru, Brazil, Bolivia, Asia, Polynesia. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 79 (LAGU, MO); E. Sandoval & E. Fuentes 1300 (B, LAGU); E. Sandoval & M. Sandoval 1016 (B, LAGU): R. Seiler 945 (F); S. Castillo s.n. (ISF00687) (B, F, LAGU, MO): T. Croat 42159 (ITIC). La Libertad: E.A. Montalvo 73a (ITIC); K. Prestegard 27 (MP-00034) (LAGU), San Salvador: R. Seiler 223, 373 (F, MHES). Chalatenango: L. Lara 141 (MHES). La Paz: R. Villacorta et al. 2855 (AAU, B, BM, LAGU, MO). Usulutan: D. Williams 51 (B, EAP, LAGU, MO). Morazan: R. Seiler 1044 (F, ITIC); 1088a (F, ITIC, MHES). Nephrolepis undulata (Afzel. ex Sw.) J. Sm., Bot. Mag. 72 (Compendium): 35 bis. 1846 Aspidium undulatum Afzel. ex Sw., Nephrolepis occidentalis Kunze, N. intermedia Fée Local names: helecho cola de iguana, helecho cola de quezallio, helecho quetzal. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Africa, Asia. Salvadoran distribution: Ahuachapan: E. Sandoval et al. 773, 1070 (B, LAGU); M. Sandoval & E. Sandoval 70 (B, F, LAGU). Santa Ana: B. Pfeiffer-Berendsohn 58 (B, LAGU, MO): R. Seiler 1215 (F, MHES). San Salvador: R. Seiler 44 (F, MHES); 637 (MHES). LYGODIACEAE Lygodium venustum Sw., J. Bot. (Schrader) 1801(2): 303. 1803. Lygodium mexicanum C. Pres Local names: alambrillo, helecho bejuco de san pablo, helecho crespillo. Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, 173 FERN GAZ. 18(4): 120-215. 2008 Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution. Ahuachapan: A. Sermefio 42 (JBL00946) (B, F, LAGU, MO); D. Witsberger s.n. (12-998) (MHES); E. Sandoval & M. Sandoval 1150 (B, LAGU); F. Chinchilla s.n. (ISB00307), s.n. ISB00377) (LAGU); J.L. Linares & On. Martinez 3085 (JBL01902) (LAGU); J.M. Rosales 241, 477 (AAU, B, BM, LAGU, MO); 1128 (B, BM, LAGU, MO); 1858, 2403 (B, BM, LAGU, MEXU, MO); M. Sandoval & E. Sandoval 106 (B, LAGU); O. Guerrero s.n. (ISF00448) (AAU, B, F, LAGU); O. Rivera D. & E. Sandoval s.n. (ISB00695) (AAU, B, F, LAGU); S. Castillo s.n. (ISFO0108) (B, LAGU); W. Berendsohn et a/. 1412 (B, LAGU, MO). Santa Ana: J.C. Gonzalez & R. Villacorta 121 (B, LAGU, MO); J.L. Linares & C.A. Martinez 1169 (JBL04295) (LAGU); K. Prestegard 22 (MP-00045) (LAGU). La Libertad: L.O. Williams et al. 15218 (F); N. Ventura 5008 (ITIC); O. Gonzalez er al. s.n. (JBL04501) (LAGU); R. Villacorta 1187 (B, LAGU, MO). San Salvador: M.C. Carlson s.n., 11 (F). Chalatenango: M.L. Reyna de Aguilar 269 (MHES). La Paz: R. Villacorta & S. Villacorta 1147 (B, ITIC, LAGU, MO); W. Berendsohn et al. 1194 (B, F, HBG, LAGU). San Vicente: P. Standley ef al. 3516a, 3669 (F). San Miguel: A. Salgado 68 (MHES): J. Tucker 816 (F). Cabanas: G. Davidse et al. 37095 (B, BM, ITIC, LAGU, MO); R.A. Carballo et al. 222 (B, LAGU, MO). Morazan: J. Monterrosa & C. Rivera 670 (B, BM, LAGU, MO). La Unién: A.K. Monro et al. 2762 (B, BM, ITIC, LAGU, MO); G.W. Barclay 2632 (F). MARATTIACEAE Marattia excavata Underw., N. Amer. Fl. 16: 22. 1909. Local names: helecho casco de mula, helecho casco de burro. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Sonsonate: J.L. Linares & C.A. Martinez 3046 (JBL01932) (LAGU). Santa Ana: A. Molina et al. 21493 (ITIC); G Cerén & D. Rodriguez s.n. (JBL04652) (LAGU); J. Monterrosa & R. Carballo 854 (LAGU); J. Monterrosa & R. Martinez 947 (LAGU); L. Gonzalez s.n. (ITIC); L. Lara 229 (MHES); R. Seiler 268 (MHES); R. Villacorta et a/. 1054 (B, LAGU, MO). Chalatenango: R. Seiler 436 (JBL00112) (LAGU, MHES). Morazan: R. Seiler 1031a (ITIC, MHES). Note: Used by local people in Montecristo National Park as a contraceptive. Marattia weinmanniifolia Liebm., Mexic. Bregn. 308 (reprint 156). 1849. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama. Salvadoran distribution: Santa Ana: R. Seiler 234 (F, MHES). MARSILEACEAE Marsilea ancylopoda A. Braun, Monatsber. Konigl. Preuss. Akad. Wiss. Berlin 1863: 434. 1864. Zaluzianskia ancylopoda (A. Braun) Kuntze, Z. mexicana (A. Braun) Kuntze, Marsilea mexicana A. Braun Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 174 Salvadoran distribution: La Unién: N.C. Fassett 28654 (ITIC); R. Villacorta et al. 2708 (LAGU); J. Monterrosa & R. Villacorta 1636 (BM, LAGU, MHES, MO OPHIOGLOSACEAE > Botrychium schaffneri Underw., Bull. Torrey Bot. Club 30: 51. 1903. Sceptridium schaffneri (Underw.) Lyon, S. pusillum (Underw.) Lyon, Botrychium pusillum Underw., B. schaffneri Underw. var. pusillum (Underw.) R. T. Clausen Global distribution: Mexico, — Honduras, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Bolivia, Argenti Source: Seiler (1982): 382; ae (1989) 1: 15; Mickel & Smith (2004): 163. Botrychium underwoodianum Maxon, Bull. Torrey Bot. Club 32: 222. 1905. Botrychium ternatum Hook. & Baker var. daedaleum Christ Global distribution: Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Brazil. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 46 (B, LAGU, MO); 49 (LAGU, MO): M.L. Reyna de Aguilar 778 (MHES); S. Winkler 8a (ITIC). Botrychium virginianum (L.) Sw., J. Bot. (Schrader) 1800(2): III. 1801. Osmunda virginiana L., O. Cicutaria Savigny in Lamark, Botrvchium cicutarium (Savigny) Sw., B. virginianum (L.) Sw. var. mexicanum Hook. & Grev., B. brachystachys Kunze, B. virginianum (L.) Sw. var. meridionale Butters, B. B virginianum subsp. meridionale (Butters) R. T. Clausen Global distribution: Canada, USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia; Europe, Asia, Salvadoran distribution: Santa Ana: F. Wunderlin 1230 (ITIC); J. Monterrosa et al. 197 (LAGU); 1023 (B, LAGU, NY); M.L. Reyna de Aguilar 679 (MHES),; R. Seiler 132 (F, MHES); 252 (F, ITIC); 1146 (F). Chalatenango: R. Seiler 382 (JBL00093) (LAGU, MHES). Ophioglossum reticulatum L., Sp. P|. 2: 1063. 1753. . Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Argentina, Africa, Asia. Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar 777 (MHES). OSMUNDACEAE Osmunda cinnamomea L.., Sp. pl. 2: 1066. 1753. Osmundastrum cinnamomeum (L.) C. Pres! Global distribution: Canada, USA, Mexico, Guatemala, Honduras, Costa Rica, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Asia. Salvadoran distribution. Sonsonate: R. Villacorta 768 (B, LAGU, MO). Morazan: R. Seiler 1009, 1075 (F, MHES); S. Winkler 7b (ITIC). ie FERN GAZ. 18(4): 120-215. 2008 Osmunda regalis L. var. spectabilis (Willd.) A. Gray, Manual, ed. 2, 600. 1856. Osmunda spectabilis Willd., O. mexicana Fée, O. palmeri A. E. Bobrov Local names: helecho real. Global distribution: Canada, USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. Salvadoran distribution: Ahuachapan: G. Davidse et al. 37401 (B, BM, ITIC, LAGU, O); N.C. Fassett 28718a (ITIC); N. Herrera s.n. (JBL03779) (B, LAGU, MO). Sonsonate: R. Villacorta 771 (B, LAGU, MO). Santa Ana: R. Seiler 779 (F). Morazan: R. Seiler 977 (F, ITIC, MHES). POLY PODIACEAE Campyloneurum amphostemon (Kunze ex Klotzsch) Fée, Mém. Foug. 5: 258. 1852. Polypodium amphostemon Kunze ex Klotzsch, P. angustifolium Sw. var. amphostemon (Kunze ex Klotzsch) Baker in Martius, Campyloneurum angustifolium (Sw.) Fee vat. amphostemon (Kunze ex Klotzsch) Farw. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia Salvadoran distribution: Santa Ana: A. Molina et al. 16764 (F, ITIC); J. Monterrosa ef al. 193 (B, BM, LAGU, MO); R. Seiler 89, 113 (F, MHES); 187 (MHES); 656, 669 (F, MHES); S. Winkler 49, 58 (ITIC). Chalatenango: R. Seiler 385 (JBL00203) (LAGU, MHES); 395 (JBL04945) (F, LAGU, MHES); 392, 754, 755 (F, MHES); W. Berendsohn et al. 1592 (B, LAGU, MO). Campyloneurum Bi oar (Sw.) Fée, Mém. Foug. 5: 257. 1852. Polypodium angustifolium Global distribution: USA, a Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: P. Standley ef a/. 19932 (F). Santa Ana: R. Seiler 707 (MHES). Chalatenango: R. Seiler 393 (JBL00196) (F, LAGU); 394 (JBLO0202) (F, LAGU, MHES). San Vicente: S. Winkler 69 (ITIC). > Campyloneurum costatum (Kunze) C. Presl, Suppl. Tent. Pterid. 190. 1836. Polypodium costatum Kunze Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Brazil. Source: Seiler (1982): 385 (as Polypodium costatum); Berendsohn (1989) 1: 17 (as Polyvpodium costatum),. Campyloneurum ensifolium (Willd.) J. Sm., Cat. Cult. Ferns 12. 1857. Secale ieii ensifolium Willd., P. angustifolium Sw. var. ensifolium (Willd.) Hicken, ampyloneurum angustifolium (Sw.) Fée var. ensifolium (Willd.) Farw. Lvaal names: sails helecho barba de cacique, helecho barbiquejo. Global distribution: Mexico, Guatemala. Salvadoran distribution: Jaurequi 87 (UC) (Mickel & Smith 2004). Ahuachapan: A.K. Monro et al. 2971, 3425 (B, BM, ITIC, LAGU, MO); E. Sandoval 1258 (B, LAGU): MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 176 J.L. Linares 516 (JBL01863) (LAGU); P. Standley er al. 2640 (F); R. Seiler 969 (F, MHES); R. Villacorta et al. 869 (B, ITIC, LAGU); S. de Moore s.n. (12-926) (MHES): W. Berendsohn ef a/. 1370, 1433 (B, LAGU, MO). Sonsonate: R. Seiler 3 (F, MHES). Santa Ana: A. del Cid 1614 (ITIC); A. Mangandi et al. s.n. (JBL04469) (LAGU); A.K. Monro et al. 2304 (B, BM, ITIC, LAGU, MO); E.A. Montalvo s.n. ge a Monterrosa & R. Carballo 512 (B, LAGU, MO); M. Magafia 32 (MP-00039) (LAGU); R. Villacorta 2173 (B, LAGU, MO); S. Winkler s.n., 50 (ITIC); V.M. Martinez s.n. (CMC00256) (LAGU). La Libertad: E.A. Montalvo 4298 (ITIC). San Salvador: P. Standley et al. 22672 (F). La Paz: S. Flores 9 (ITIC). San Vicente: E.A. Montalvo s.n. (ITIC). Usulutan: D. Williams & R.W. Herrera 455 (LAGU). San Miguel: A. Salgado 67 (F, MHES). Note: This species name has been placed in synonomy, or as a variety of C. angustifolium. Lellinger (1988) and Leon (1992) clarify the diagnostic characters (scale cell shape). See discussion by Mickel & Smith (2004) under C. ensifolium. Campyloneurum phyllitidis (L.) C. Presl, Suppl. Tent. Pterid. 190. 1836. Polypodium phyllitidis L. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: San Salvador: R. Seiler 971 (F, MHES). Campyloneurum xalapense Feé, Mém. Foug. 5: 258. 1852. Polypodium xalapense (Fée) Christ, P. weatherbyanum F. Seym., Campyloneurum caudatum Fée . Local name: cincho. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: M.L. Reyna de Aguilar 651 (MHES),; R. Seiler 907 (F, ITIC, MHES). Sonsonate: R. Seiler 595 (F, MHES); R. Villacorta er al. 329 (LAGU, MO). Santa Ana: A. Molina ef a/. 12658: B. Pfeiffer-Berendsohn 48 (B, LAGU, MO): 76 (LAGU, MO); J. Monterrosa et a/. 178, 968 (LAGU); J. Monterrosa & R. Carballo 278 (B, BM, LAGU, MO); 978 (B, LAGU, NY): M.B. Siu s.n. (ITIC); R. Seiler 122 (F, ITIC, MHES); 274, 670 (F, MHES); S. Winkler 57 (ITIC). San Salvador: M.C. Carlson 505 (F); S. Winkler 59 (ITIC). San Vicente: E.A. Montalvo 4655 (ITIC): R. Seiler 324 (F); 325 (F, MHES). San Miguel: M. Renderos ef al. 672 (LAGU), Ceradenia jungermannioides (Klotzsch) L. E. Bishop, Amer. Fern J. 78: 4. 1988. Polypodium jungermannioides Klotzsch, Grammitis jungermannioides (Klotzsch) Ching Global distribution. Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Brazil; Azores. Salvadoran distribution: R. Seiler 1194 (UC) (A.R. Smith in Davidse et al. 1995). Santa Ana: R. Seiler 124 (F, ITIC, MHES). > Ceradenia margaritata (A. R. Sm.) L. E. Bishop, Amer. Fern J. 78: 5. 1988. Grammitis margaritata A. R. Sm. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Tucker 1075 (UC) (A.R. Smith in Davidse ef al. 1995). hey FERN GAZ. 18(4): 120-215. 2008 Source: Smith in Davidse et al. (1995): 370; Mickel & Smith (2004): 173. Cochlidium rostratum (Hook.) Maxon ex C. Chr., Dansk Bot. Ark. 6(3): 23. 1929. Monogramma rostrata Hook., Grammitis rostrata (Hook.) R. M. Tryon & A. F. Tryon Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela. Salvadoran distribution: Sonsonate: R. Seiler 606 (F, MHES). Santa Ana: A. Molina et al. 12492, 16796 (F); J. Toledo s.n. (ITIC); R. Seiler 98 (F, MHES); 647 (MHES). Morazan: R. Seiler 997 (F, MHES); S. Winkler 46 (ITIC). Cochlidium serrulatum (Sw.) L. E. Bishop, Amer. Fern J. 68: 80. 1978. Acrostichum serrulatum Sw., Grammitis serrulata (Sw.) Sw., Xiphopteris serrulata (Sw.) Kaulf., Polypodium serrulatum (Sw.) Mett., P. duale Maxon Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Ecuador, Peru, Brazil, Bolivia, Africa. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1968 (B, BM, ITIC, LAGU, MO). Chalatenango: S. Winkler 45 (ITIC). Morazan: R. Seiler 1004 (F, MHES, NY). > Lellingeria apiculata (Kunze ex Klotzsch) A. R. Sm. & R. C. Moran, Amer. Fern J. 81: 83. 1991 Polypodium apiculatum Kunze ex Klotzsch, Ctenopteris apiculata (Kunze ex Klotzsch) Copel., Grammitis apiculata (Kunze ex Klotzsch) F. Seym Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia, Brazil. Source: Seiler (1982): 385 (as Grammitis appiculata); Berendsohn (1989) 1: 16 (as Grammitis appiculata). Lellingeria prionodes (Mickel & Beitel) A. R. Sm. & R. C. Moran, Amer. Fern J. 81: Grammitis prionodes Mickel & Beitel Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Sonsonate: R. Seiler 602 (F, MHES). Santa Ana: E.A. Montalvo 4155 (ITIC); J. Monterrosa et al. 226 (B, LAGU); L. Lara 237 (MHES); R. Villacorta & L. Lara 2664 (B, LAGU). Chalatenango: S. Winkler 81 (ITIC). Morazan: R. Seiler 999 (F, MHES). Loxogramme mexicana (Fée) C. Chr., Index Filic. Suppl. 3: 25. 1934. Selliguea mexicana Fée, Gymnogramma mexicana (Fée) Baker in Hooker & Baker, G. salvinii (Hook.) Hook., Grammitis salvinii Hook., Loxogramme salvinii (Hook.) Maxon Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: E.A. Montalvo 57 (ITIC); G. Davidse er al. 37411 (B, BM, ITIC, LAGU, MO); R. Seiler 822 (F, MHES). Santa Ana: R. Seiler 359, 503 (F, MHES). San Salvador: R. Seiler 342 (F, MHES). » Melpomene deltata (Mickel & Beitel) A. R. Sm. & R. C. Moran, Novon 2: 429. 1992. Grammitis deltata Mickel & Beitel MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 178 Global distribution: Mexico, Honduras. Source: Mickel & Beitel (1988): 199; Moran & Smith in Davidse et al. (1995): 381; Mickel & Smith (2004): 390. Melpomene firma (J. Sm.) A. R. Sm. & R. C. Moran, Novon 2: 430. 1992. Polypodium firmum Klotzsch, P. aromaticum Maxon, Ctenopteris firma J. Sm., C. aromatica (Maxon) Copel., Grammitis firma (J. Sm.) C. V. Morton, G aromatica (Maxon) Proctor Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Sonsonate: R. Seiler 603 (MHES). Santa Ana: J. Monterrosa et al. 182 (LAGU); R. Seiler 96 (F, ITIC, MHES). Chalatenango: R. Seiler 737 (MHES). Morazan: R. Seiler 998 (F, MHES). > Melpomene leptostoma (Fée) A. R. Sm. & R. C. Moran, Novon 2: 430. 1992. Polypodium leptostomum Fée, Polypodium productum Maxon, Ctenopteris leptostoma (Fée) Copel., Grammitis leptostoma (Fée) F. Seym. Global distribution: Mexico, Guatemala. Source: Stolze (1981): 258 (as Grammitis leptostoma); Berendsohn (1989) 1: 16 (as Grammitis leptostoma). Melpomene moniliformis (Lag. ex Sw.) A. R. Sm. & R. C. Moran, Novon 2: 430. 1992. Polypodium moniliforme Lag. ex Sw., P. moniliforme Lag. ex Sw. var. major Liebm. Ctenopteris moniliformis (Lag. ex Sw.) J. Sm., Grammitis moniliformis (Lag. ex Sw.) Proctor Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: L. Lara 98 (MHES); M.C. Carlson 900a (F); W. Berendsohn et a/. 1550 (B, LAGU). Chalatenango: R. Seiler 431 (JBL04944) (F, LAGU, MHES); R. Seiler 748 (F, MHES); 758 (MHES); R. Villacorta et al. 2074 (B, LAGU, MO): W. Berendsohn et al. 1568 (B, LAGU, MO). > Melpomene pilosissima (M. Martens & Galeotti) A. R. Sm. & R. C. Moran, Novon 2: 431. 1992. Polypodium pilosissimum M. Martens & Galeotti, Crenopteris pilosissima (M. Martens & Galeotti) Copel., Grammitis pilosissima (M. Martens & Galeotti) C. V. Morton Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Suriname, Ecuador, Peru, Bolivia, Brazil. Source: Seiler (1982): 386 (as Grammitis pilossisima); Berendsohn (1989) 1: 16 (as Grammitis pilossisima). Melpomene xiphopteroides (Liebm.) A. R. Sm. & R. C. Moran, Novon 2: 431. 1992. Polypodium xiphopteroides Liebm., P. rigens Maxon, Ctenopteris rigens (Maxon) Copel., C. megaloura Copel., Grammitis xiphopteroides (Liebm.) A. R. Sm., Grammitis rigens (Maxon) Proctor Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 93 (F, MHES); 95 (ITIC, MHES). 179 FERN GAZ. 18(4): 120-215. 2008 > Micropolypodium basiattenuatum (Jenman) A. R. Sm., Novon 2: 422. 1992. Polypodium basiattenuatum Jenman, Xiphopteris basiattenuata (Jenman) Copel., Grammitis basiattenuata (Jenman) Proctor, G. basiattenuata (Jenman) Proctor var. valens Mickel & Beitel Global distribution: Mexico, Guatemala, Honduras, Antilles, Venezuela. Salvadoran distribution: R. Seiler 1133, NY (Mickel & Smith 2004). Source: Seiler (1982): 385 (as Grammitis basiattenuata); Berendsohn (1989) 1: 16 (as Grammitis rigens); Smith in Davidse et al. (1995): 383; Mickel & Smith (2004): 400. Niphidium crassifolium (L.) Lellinger, Amer. Fern J. 62: 106. 1972. Polypodium crassifolium L., Pessopteris crassifolia (L.) Underw. & Maxon Local name: cincho. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Seiler 1529, MO (Pacheco in Davidse et a/. 1995). Santa Ana: R. Seiler 491 (F, MHES). La Libertad: O. Pank s.n. (MP-00041) (LAGU). Pecluma alfredii (Rosenst.) M. G. Price var. cupreolepis (A. M. Evans) A. R. Sm., Mem. New York Bot. Gard. 88: 437. 2004. Polypodium cupreolepis A. M. Evans, Pecluma cupreolepis (A. M. Evans) M. G. Price Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: J.M. Rosales 410 (AAU, B, BM, LAGU, MO); R. Seiler 914 (F, ITIC); 968 (F, MHES). Santa Ana: B. Pfeiffer-Berendsohn 81 (LAGU, MO); G. Cerén & D. Rodriguez s.n. (JBL04647) (LAGU); J. Tucker 1249 (F); R. Seiler 129 (F, MHES); 287 (MHES). Chalatenango: R. Seiler 385 (F, MHES). San Vicente: E.A. Montalvo 73b (ITIC). San Miguel: W. Eggler 665 (ITIC). Morazan: R. Seiler 1023 (F, MHES). Pecluma plumula (Humb. & Bonpl. ex Willd.) M.G. Price, Amer, Fern J. 73: 115. 1983. Polypodium plumula Humb. & Bonpl. ex Willd., P pulchrum M. Martens & Galeotti, P. pulchrum M. Martens & Galeotti var. minus E. Fourn.. Ctenopteris plumula (Humb. & Bonpl. ex Willd.) J. Sm. Global distribution: USA, Mexico, Guatemala, Belize. Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia. Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E.A. Montalvo 62 (ITIC); H. Weber 35 (ITIC): R. Villacorta et al. 872 (B, LAGU, MO); W. Berendsohn et al. 1373 (B, LAGU, MO). Sonsonate: R. Seiler 75 (F, MHES). Santa Ana: M.B. Sius.n. (ITIC); R.A. Carballo & E.O. Ramos 675 (B, LAGU, MO). La Libertad: M. Renderos 496, B, K (LAGU, MO); N. Ventura 5016 (ITIC); S. Guadalupe & F. Lessette s.n. (JBL04481) (LAGU). San Salvador: M.C. Carlson 356 (F). San Vicente: R. Seiler 323 (F, MHES). Usulutan: D. Williams 56 (LAGU, MO): D. Williams & R.W. Herrera 298 (LAGU). San Miguel: A. Salgado 64 (F, MHES). Usulutan: E.A. Montalvo 5035, 5037 (ITIC). Morazan: J. Monterrosa & C. Rivera 696 (B, LAGU, MO): R.A. Carballo et a/. 1044 (LAGU). Phlebodium pseudoaureum (Cav.) Lellinger, Amer. Fern J. 77: 101. 1987. Polypodium pseudoaureum Cav., Polypodium areolatum Humb. & Bonpl. ex Willd., P: MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 180 areolatum Humb. & Bonpl. ex Willd. var. /oreum J. Bommer, P. aureum auct., en parte L., Goniophlebium areolatum (Humb. & Bonpl. ex Willd.) C. Presl, Phlebodium aureum auct., en parte Local names: helecho calaguala, helecho cola de pavo real. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Brazil, Bolivia, Argentina, Paraguay. Salvadoran distribution: Ahuachapan: D. Witsberger 696 (MHES); E. Sandoval & M. Sandoval 1299 (B, LAGU): W. Berendsohn ef al. 1458 (LAGU). Santa Ana: G. Davidse et al. 37233 (B, BM, ITIC, LAGU, MO); J. Monterrosa & R. Carballo 284 (B, BM, EAP, F, LAGU, MO); R. Seiler 185 (MHES). San Salvador: R. Seiler 28, 56, 513 (MHES). San Vicente: R. Seiler 317 (MHES). Usulutan: D. Williams & R.W. Herrera 442 (LAGU). Morazan: R. Seiler 1000 (MHES). Pleopeltis angusta Humb. & Bonpl. ex Willd. var. angusta, Sp. Pl., ed. 4, 5(1): 211. Polypodium angustum (Humb. & Bonpl. ex Willd.) Liebm. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Local names: helecho mano de mapache, helecho pié de mapache. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2977, 3426 (B, BM, ITIC, LAGU, MO); D. Witsberger 688 (MHES); E. Sandoval & M. Sandoval 1298 (B, LAGU); F. Chinchilla s.n. (ISB00206) (B, LAGU, MO); M. Magana 16 (MP-00040) (LAGU); R. Seiler 913, 967 (F, MHES): S. de Moore s.n. (12-927) (MHES); W. Berendsohn ef al. 1456, 1459 (LAGU). Sonsonate: R. Seiler 627 (F, MHES). Santa Ana: B. Pfeiffer-Berendsohn ef al. 10 (B, F, LAGU, MO); G. Cerén & D. Rodriguez s.n. (JBL04668) (LAGU); J. Monterrosa et a/. 175 (B, BM, LAGU, MO); 176 (B, LAGU, MO); J. Monterrosa et al. 920 (B, LAGU, NY): 973 (LAGU); K. Sidwell er al. 436 (B, BM, ITIC, LAGU, MO); M.B. Siu s.n. (ITIC); R. Seiler 168 (F, MHES). La Libertad: W. Berendsohn et al. 1445 (LAGU); W. Eggler 681 (ITIC). San Salvador: A.K. Monro ef al. 2240 (B, BM, ITIC, LAGU, MO); M.C. Carlson 429 (F); @, Rohweder s.n. (ITIC); R. Seiler 26 (F, ITIC, MHES). Usulutan: A.K. Monro et al. 2999 (B, BM, ITIC, LAGU, MO); D. Williams 129 (B, LAGU, MO). San Miguel: K. Sidwell et al, 884 (B, BM, ITIC, LAGU, MO) Pleopeltis astrolepis (Liebm.) E. Fourn., Mexic. PI. I: 87. 1872. Polypodium astrolepis Liebm., P. elongatum (Sw.) Mett., P lanceolatum L. var elongatum (Sw.) Krug, Phlebodium astrolepis (Liebm.) Conzantti, Grammitis lanceolata Schkuhr, G. revoluta Spreng. ex Willd., Gymnogramma elongata (Sw.) Hook., Pleopeltis revoluta (Spreng. ex Willd.) A. R. Sm. Local names: helecho barba amarilla, helecho barba negra, helecho barbiquejo. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution. Ahuachapan: A.K. Monro & M.L. Fomtg 2973 (B, BM, ITIC, LAGU, MO); D. Witsberger 711 (MHES): E. Sandoval er al. 729, 1430 (B, LAGU); F. Chinchilla s.n. (AAU, B, LAGU); M. Sandoval & E. Sandoval 138 (F, LAGU); P. Standley e¢ al. 2516 (F); R. Seiler 928, 965 (F, MHES); W. Berendsohn er al. 1430 (B, LAGU, MO); 1466 (LAGU). Sonsonete: R. Seiler 77 (F). Santa Ana: B. 181] FERN GAZ. 18(4): 120-215. 2008 Pfeiffer-Berendsohn 20 (LAGU, MO); V.M. Martinez s.n. (CMCO0OO103) (LAGU). La Libertad: J. Monterrosa ef a/. s.n. (JBL04485) (LAGU); N. Ventura 5020 (ITIC); O. Rohweder 2634 (JBL0O1246), 2635 (JBL01245), 2636 (JBL0O1244) (LAGU); W. Zelaya et al. sn. (LAGU). San Salvador: R. Seiler 507 (F, MHES); 1515 (F). San Vicente: E.A. Montalvo 5098 (ITIC). Usulutan: E.A. Montalvo 5047 (ITIC); J. Lagos 1783 (ITIC). Cuseatlan: C. Salazar s.n. (ITIC). Pleopeltis mexicana (Fée) Mickel & Beitel, Amer. Fern J. 77: 21. 1987. Drymaria mexicana Fée, Polypodium lanceolatum L. var. trichophorum Weath., Pleopeltis macrocarpa (Bory ex Willd.) Kaulf. var. trichophora (Weath.) Pic. Serm. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: A. Molina et al. 12652 (F; B). Pfeiffer- Berendsohn 26, 78, 79 (B, LAGU, MO); 56 (LAGU, MO): G. Davidse et al. 37164, 37306 (B, BM, ITIC, LAGU, MO); J. Monterrosa et al. 171 (B, BM, LAGU, MO); R. Seiler 109 (F, MHES); S. Winkler 56 (ITIC). Chalatenango: R. Seiler 386 (JBL00201) (ITIC, LAGU); R. Seiler 152, 729 (F, MHES). Morazan: R. Seiler 1005 (F, ITIC, MHES); 1015 (F, MHES). Pleopeltis polylepis (Roemer ex Kunze) T. Moore var. interjecta (Weath) E. A. Hooper, Amer. Fern J. 85: 79. 1995. Polypodium peltatum Cay. var. interjectum Weath., Pleopeltis macrocarpa (Bory ex Willd.) Kaulf. var. interjecta (Weath.) A. R. Sm., P. interjecta (Weath.) Mickel & Beitel Global distribution: Mexico, Guatemala. Salvadoran distribution: Santa Ana: W. Berendsohn et al. 1714 (B, LAGU). Polypodium alansmithii R. C. Moran, Ann. Missouri Bot. Gard. 77: 845. 1990. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Sonsonate: R. Seiler 633 (MHES). Santa Ana: A. Molina ef al. 12562 (F; B). Pfeiffer-Berendsohn 77 (B, LAGU, MO): E.A. Montalvo s.n., 3641 (ITIC); G. Cerén & D. Rodriguez s.n. (JBL04667) (LAGU); G. Davidse et al. 37301 (B, BM, ITIC, LAGU, MO); J. Monterrosa er al. 165 (B, BM, LAGU, MO): 951, 964 (B. LAGU, NY); 942 (B, LAGU, MO, NY); 1007 (LAGU, NY); K. Sidwell et al. 847 (B, BM, ITIC, LAGU, MO); M.C. Carlson 726, 727, 970 (F); N.C. Fassett 28292 (ITIC); R. Seiler 87 (F); 91, 190 (F, MHES); R. Villacorta & Claus 2147 (B, LAGU, MO); W. Berendsohn 1303 (LAGU, MO). San Salvador: R. Seiler 214 (F, MHES). Chalatenango: R. Seiler 398 (JBL00198) (F, LAGU, MHES); W. Heed 36 (ITIC). San Vicente: R. Seiler 326 (MHES). Polypodium castaneum Maxon & Tejero, Act. Bot. Mex. 67: 87. 2004. Global distribution: Guatemala. Salvadoran distribution: Sonsonate: Croat 42213, MO. Santa Ana: J. Monterrosa e? al. 967 (B, BM, IZTA, LAGU, MO, NY). Polypodium colpodes Kunze var. colpodes, Linnea 23: 276, 316. 1850. Polypodium fuscopetiolatum A. R. Sm. Local names: helecho calaguala rojo, helecho de arbol, helecho pluma de ganzo. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 182 Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2988, 3427 (B, BM, ITIC, LAGU, MO); E. Sandoval ef al. 548, 1442 (B, LAGU); F. Chinchilla s.n. (ISB00658) (AAU, B, F, LAGU); J.M. Rosales 1061 (B, LAGU); 1301, 1491 (AAU, B, BM, LAGU, MO); 1420 (B, BM, LAGU, MO); M. Sandoval & E. Sandoval 58 (B, F, LAGU). Sonsonate: R. Seiler 38 (F, MHES); T. Croat 42237 (ITIC). Santa Ana: J. Lagos 1616 (ITIC); V.M. Martinez s.n. (CMC00306) (B, LAGU). La Libertad: N. Ventura 5017, 5021 (ITIC); R. Seiler 719 (F, MHES). San Salvador: R. Seiler 439, 584 (F, MHES); 514 (MHES). Chalatenango: L. Lara 210 (MHES); R. Seiler 473 (F, MHES). ¢Polypodium diplotrichum Mickel & Beitel, Mem. New York Bot. Gard. 46: 297. 1988. Goniophlebium lepidotrichum Fée Global distribution: Mexico. Salvadoran distribution: Ahuachapan: J.Monterrosa & D.Rodriguez 1446 (BM, IZTA, Polypodium fissidens Maxon, Contr. U.S. Natl. Herb. 8: 275. 1903. Global distribution: Mexico, Guatemala, Honduras. Local names: helecho calaguala Salvadoran distribution: Chalatenango: W. Berendsohn ef al. 1571 (B, LAGU); R. Seiler 156, 164, 756 (F, MHES). Polypodium furfuraceum Schltdl. & Cham., Linnea 5: 607. 1830. Polypodium nivosum Fée, P. furfuraceum Schltdl. & Cham. var. coronulatum E. Fourn., P. furfuraceum Schltdl. & Cham. var. rufum E. Fourn., P. margallii Rovirosa Local names: cola de ardilla, helecho peine de indigena, helecho peinetero rojo, palmita de café. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Peru, Bolivia. Salvadoran distribution: Ahuachapan: A.K. Monro & M.L. Fomtg 2972 (B, BM, ITIC, LAGU, MO); E. Sandoval et al. 1190, 1403 (B, LAGU); F. Chinchilla & E. Sandoval s.n. (ISB00428) (LAGU); R. Seiler 829 (ITIC); 970 (F, MHES); S. de Moore s-n. (12-929) (MHES):; W. Berendsohn et a/. 1429 (LAGU). Sonsonate: R. Seiler 5, 628 (F, MHES). Santa Ana: B. Pfeiffer-Berendsohn 19 (LAGU); E.A. Montalvo 4297 (ITIC); T. Croat 43037 (ITIC); V.M. Martinez s.n. (CMC00251) (LAGU). La Libertad: E.A. Montalvo 4302 (ITIC); N. Ventura 5005 (ITIC); V. Henriquez & C. Bermudez s.n. (JBL04482) (LAGU); W. Berendsohn 1452 (LAGU). San Salvador: E.A. Montalvo 5350 (ITIC); L.O. Williams et al. 16746 (F); S. Flores 8 (ITIC). San Vicente: E.A. Montalvo 5091 (ITIC). Cuscatlan: C. Salazar s.n. (ITIC). Usulutan: D. Williams & R.W. Herrera 300 (LAGU). San Miguel: A. Salgado 65b (F, MHES). La Uni6n: A.K. Monro et al. 2123 (B, BM, ITIC, LAGU, MO). Polypodium hartwegianum Hook. in Bentham, PI. Hartw. 54. 1840. Polypodium ellipsoideum Fée, P. pubescens Fée Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 12560 (F, ITIC); J. Monterrosa er al. 1031, B, IZTA (LAGU, NY); 1044, IZTA (LAGU); L. Lara s.n. (12-992) (MHES): 183 FERN GAZ. 18(4): 120-215. 2008 R. Seiler 80, 81, 189, 672 (F, MHES). Chalatenango: E.A. Montalvo 4811 (ITIC); R. Seiler 430 (JBL00199) (LAGU); R. Seiler 162, 426 (F, MHES). Morazan: R. Seiler 1092 (F, ITIC, MHES). > Polypodium hispidulum Bartlett, Proc. Amer. Acad. Arts 43: 48. 1907. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua. Source: Seiler (1982): 385; Berendsohn (1989) 1: 17; Moran in Davidse et al. (1995): 358; Mickel & Smith (2004): 494. > Polypodium lindenianum Kunze, Farrnkrauter 2: 83. 1851. Polypodium cancellatum Fée, P. verapax Christ Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica. Source: Stolze (1981): 408; Berendsohn (1989) 1: 17; Moran in Davidse et al. (1995): 362; Mickel & Smith (2004): 495 Polypodium muenchii Christ, Bull. Herb. Boissier, sér. 2, 3: 147. 1903. Pleopeltis muenchii (Christ) A. R. Sm., Microgramma muenchii (Christ) Copel., Microphlebodium muenchii (Christ) L. D. Gomez Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Chalatenango: R. Villacorta et al. 2075 (B, LAGU); R. Seiler 400 (JBL00200) (LAGU); R. Seiler 153 (MHES); Tucker 1070 (EAP, F). Polypodium plebeium Schltdl. & Cham., Linnea 5: 607. 1830. Polypodium cheilostictum Fée, P. karwinskyanum (C. Presl) A. Braun ex Kunze, Marginaria karwinskyvana C. Pres| Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Sonsonate: R. Seiler 608 (F, MHES). Santa Ana: R. Seiler 16 (MHES). San Salvador: R. Seiler 206 (MHES). Morazan: R. Seiler 1019 (F, MHES). Polypodium plesiosorum Kunze var. plesiosorum, Linnea 18: 313. 1844. Polypodium gonatodes Kunze, P. confluens Liebm., P. anisomeron (Fée) E. Fourn. vat. elongatum E. Fourn., P. anisomeron (Fée) E. Fourn. var. genuinum E. Fourn., P. anisomeron (Fée) E. Fourn. var. pectinatum E. Fourn., P. anisomeron (Fée) E. Fourn. var. abortivum E. Fourn., P. feei E. Fourn., P. van-heurckii E. Fourn., P. pachyrhizon Christ, P. rossii Christ ex H. Ross, Goniophlebium molestum Fée, G. anisomeron Fée, G. calaguala Fée, G. stramineum Underw. ex Conzatti Local name: calaguala. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: A.K. Monro et al.3428 (B, BM, ITIC, LAGU, MO). Sonsonate: A. Molina ef a/. 21758 (F, ITIC); R. Seiler 594 (F, MHES). Santa Ana: G. Davidse ef al. 37240, 37307 (B, BM, ITIC, LAGU, MO); J. Monterrosa ef al. 932 (B, LAGU, NY); L. Lara 223 (MHES); R. Seiler 167 (F, ITIC, MHES); 344 (F, MHES). Chalatenango: R. Seiler 388 (JBL00197) (F, LAGU, MHES); R. Seiler 757 (F, MHES). Polypodium plesiosorum Kunze var. plesiosori Tejero, Anales Inst. Biol. Univ. Nac. Auton. Mexico, Bot. 75(1): 24-28 (F). 1, 2a, 3g-h. 2004 Local name: calaguala. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 184 Global distribution: Mexico, Guatemala, Nicaragua. Salvadoran distribution: Santa Ana: Linares s.n. (JBLO1320) (B, LAGU, MO). Note: var. rubicundum is found in Mexico, Honduras, Costa Rica and Panama. Polypodium pleurosorum Kunze ex Mett., Abh. Senckenberg. Naturf. Ges. 2: 101, t. 2 CF): 6. 7, ESSt. Phlebodium inaequale T. Moore, Goniophlebium inaequale (T. Moore) J. Sm., Polypodium inaequale (T. Moore) Lowe, P. lowei C. Chr., P. guatemalense Hook. Local name: helecho calaguala. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: D. Witsberger 65b (MHES); E.A. Montalvo 3506 (ITIC); M. Sandoval & E. Sandoval 16 (LAGU); T. Croat 42145 (ITIC). Sonsonate: M.B. Siu s.n. (ITIC):; T. Croat 42207 (ITIC). Santa Ana: D. Rodriguez es al. 5 (B, LAGU, NY); J. Monterrosa & R. Carballo 224, 285 (B, LAGU); J. Monterrosa et al. 965 (B, NY, LAGU); M.B. Siu s.n. (ITIC); N. Lopez ef al. s.n. (JBL04626) (LAGU): R. Seiler 191, 280 (MHES):; R. Villacorta & M. Calderon 375 (LAGU, MO). San Salvador: R. Seiler 27 (F, MHES); 512 (MHES). Chalatenango: J. Tucker 1031, 1252 (F); R. Seiler 407 (JBL00195) (F, LAGU, MHES); 732 (MHES). Polypodium polypodioides (L.) Watt. var. polypodioides, Canad. Naturalist & Quart. J. ci. Sr 2. 3; 156, 1867 Acrostichum polypodioides L., Polypodium incanum Sw., Goniophlebium incanum (Sw.) J. Sm., Marginaria polypodioides (L.) Tidestr., Pleopeltis polypodioides (L.) E. G. Andrews & Windham Local name: helecho peinetero amarillo. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Uruguay, Argentina. Salvadoran distribution: Ahuachapan: F. Chinchilla s.n. (ISBO0568) (AAU, B, F, LAGU); J.M. Rosales 2537 (B, BM, LAGU, MO); 2673 (B, LAGU, MO). La Paz: R. Villacorta & S. Villacorta 1173 (B, LAGU, MO). Usulutan: A. del Cid 1782 (ITIC): D. Williams 50 (B, LAGU, MO); R. Seiler 588 (MHES). Morazan: J. Monterrosa & C. Rivera 705 (B, BM, LAGU). Polypodium polypodioides (L.) Watt. var. aciculare Weath., Contr. Gray Herb. 124: 33. 1939, Pleopeltis polypodioides (L.) Watt var. acicularis (Weath.) E. G. Andrews & Windham Local name: helecho peinetero rojo. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: E. Sandoval er al. 540, 1291 (B, LAGU); M. Sandoval & E. Sandoval 94 (LAGU); R. Seiler 828 (ITIC, MHES): 912 (F, ITIC, MHES): R. Villacorta et a/. 880 (B, LAGU, MO). Sonsonate: R. Seiler 78 (MHES). Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 11 (LAGU, MO); J. Monterrosa et al. 225 (B, BM, LAGU, MO); R. Seiler 169 (ITIC, MHES). La Libertad: R. Seiler 47 (MHES). San Salvador: R. Seiler 215 (MHES). Usulutan: D. Williams 243, 436 (LAGU); E.A. Montalvo 5044 (ITIC); J. Lagos 36 (ITIC); R.A. Carballo er al. 820 (LAGU). San Miguel: A. Salgado 65a (MHES). La Unién: R. Seiler 216 (MHES). 185 FERN GAZ. 18(4): 120-215. 2008 Polypodium puberulum Schltdl. & Cham., Linnea 5: 607. 1830. Polypodium ehrenbergianum Klotzsch, P. olivaceum Liebm. var. elatum Liebm., P. pubescens Fée, P. fournieri C. Chr., P. galeottii Mett. ex Kuhn, P. glaberulum Mickel & Beitel Global distribution: Mexico. Salvadoran distribution: Chalatenango: R. Seiler 752 (F, MHES). > Polypodium rhodopleuron Kunze, Linnea 18: 315. 1844. Polypodium isomeron E, Fourn., P. lesourdianum E. Fourn. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Source: Moran in Davidse et al. (1995): 360; Mickel & Smith (2004): 506. Polypodium sanctae-rosae (Maxon) C. Chr., Index Filic., Suppl. I: 62. 1913. Goniophlebium sanctae-rosae Maxon Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 9 (F, LAGU); 12 (F, LAGU, MO); R.A. Carballo et al. 115 (B, LAGU, MO); R. Seiler 166 (F, MHES). San Salvador: R. Seiler 32 (F, MHES); 55 (MHES); S. Winkler 27 (ITIC). Chalatenango: R. Seiler 257 (F, MHES). San Vicente: E.A. Montalvo 4610, 5087 (ITIC). Morazan: R. Seiler 1067 (F, MHES). Polypodium subpetiolatum Hook. in Bentham, PI. Hartw. 54. 1840. Polypodium biserratum M. Martens & Galeotti, P. schaffneri Fée, P. guilleminianum E. Fourn., P. hahnii E. Fourn., P. firmulum Maxon, P. maxonii C. Chr., PR. adelphum Maxon, P. teresae Maxon Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: San Vicente: R. Seiler 328 (F, MHES). ¢ Polypodium thyssanolepis A. Braun ex Klotzsch, Linnaea 20: 392. 1847. Goniophlebium thyssanolepis (A. Braun ex Klotzsch) T. Moore, G. rhagadiolepis (Fée) Fée, Marginaria thyssanolepis (A. Braun ex Klotzsch) Farw., Polypodium incanum Sw. var. fimbriatum M. Martens & Galeotti, P. rhagadiolepis Fée, P. purpusii Christ Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Chalatenango: J.Monterrosa et al. 1612 (BM, LAGU, IZTA, > Polypodium ursipes Moritz ex C. Chr., Index. Filic. 572. 1906. Polypodium ambiguum Mett. ex Kuhn Global distribution: Costa Rica, Panama. Source: Moran in Davidse et al. (1995): 354. Serpocaulon falcaria (Kunze) A.R. Sm., Taxon 55(4): 928. 2006. Goniophlebium calaguala Fée, G. invertens Fée, Polypodium falcaria Kunze. Global distribution: Mexico, Guatemala, Honduras. Salvadoran distribution: Santa Ana: A. Molina et al. 16774 (F; B). Pfeiffer- Berendsohn 23, 37 (LAGU, MO); J. Monterrosa & R. Carballo 852 (B, LAGU, MO): M.C. Carlson 728 (F); R. Seiler 123, 176 (F, MHES); 651 (MHES); V.M. Martinez s.n. MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 186 (CMC01094) (B, BM, EAP, LAGU, MO). San Salvador: R. Seiler 339 (MHES). Morazan: R. Seiler 1055 (F, MHES). Note: The name Polypodium loriceum has been applied by Léetschert (1954), Stolze (1981), Seiler (1982) and Moran in Davidse et al. (1995) to this species. See discution as P. falcaria in Mickel & Smith (2004), Serpocaulon triseriale (Sw.) A.R. Sm., Taxon 55(4): 929. 2006. Goniophlebium haenkei C. Presl, Polypodium haenkei (C. Presl) Liebm., P. triseriale Sw Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Local names: helecho calaguala, helecho yema de huevo. Salvadoran distribution: Ahuachapan: F. Chinchilla & M.J. Lopez s.n. (ISB00635) (AAU, B, F, LAGU); M. Sandoval & E. Sandoval s.n. (ISB00929) (B, F, LAGU, MO). San Salvador: E.A. Montalvo 20 (ITIC). Chalatenango: D. Sloot ef al. 41 (MAG00470) (LAGU, U). Morazan: J. Monterrosa & C. Rivera 695 (B, LAGU, MO). Terpsichore anfractuosa (Kunze ex Klotzsch) B. Leon & A. R. Sm., Amer. Fern J. 93: 86. 2003 Polypodium anfractuosum Kunze ex Klotzsch, P. saxicola Baker, P. induens Maxon, Ctenopteris anfractuosa (Kunze ex Klotzsch) Copel., Grammitis anfractuosa (Kunze ex Klotzsch) Proctor, Melpomene anfractuosa (Kunze ex Klotzsch) A. R. Sm. & R. C. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Guadalupe, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Sonsonate; R. Seiler 604 (MHES). Santa Ana: B. Pfeiffer- Berendsohn et al. 25 (B, LAGU); R. Seiler 648 (MHES). Chalatenango: E.A. Montalvo s.n. (ITIC). Morazan: R. Seiler 987 (ITIC, MHES); S. Winkler 80 (ITIC). > Terpsichore asplenifolia (L.) A. R. Sm., Novon 3: 485. 1993. Polypodium asplenifolium L., P. laxifrons Liebm., Ctenopteris asplenifolia (L.) Copel., Grammitis asplenifolia (L.) Proctor Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador. Source: Seiler (1982): 385 (as Grammitis asplenifolia), Berendsohn (1989) 1: 16 (as Grammitis asplenifolia); Moran & Smith in Davidse et al. (1995): 387. Mickel & Smith 2004: 617 Terpsichore cultrata (Willd.) A. R. Sm., Novon 3: 486. 1993. Polypodium cultratum Willd., Ctenopteris cultrata (Willd.) C opel., Grammitis cultrata (Willd.) Proctor Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: R. Seiler 1130 (UC) (Moran & Smith in Davidse ef al. 1995: 388). Santa Ana: A. Molina et a/. 16903 (F); R. Seiler 105 (F, MHES). San Salvador: S. Winkler 24 (ITIC). 187 FERN GAZ. 18(4): 120-215. 2008 ¢ Terpsichore delicatula (M. Martens & Galeotti) A. R. Sm., Novon 3: 486. 1993. Polypodium delicatulum M. Martens & Galeotti, Ctenopteris delicatula (M. Martens & Galeotti) J. Sm., Grammitis delicatula (M. Martens & Galeotti) Proctor Global distribution: Mexico, Guatemala, Honduras, Jamaica. Salvadoran distribution: Chalatenango: J.Monterrosa et al. 1624 (BM, LAGU, UC). Terpsichore lehmanniana (Hieron.) A.R. Sm., Novon 3: 487. 1993. Polypodium lehmannianum Hieron., P. sublongipes Christ, Ctenopteris lehmanniana (Hieron.) Copel., Grammitis lehmanniana (Hieron.) C. V. Morton Global distribution: Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Santa Ana: J. Monterrosa & R. Carballo 858 (B, LAGU); R. Seiler 171 (F, ITIC, MHES). Morazan: R. Seiler 1062 (F, MHES). Terpsichore semihirsuta (Klotzsch) A. R. Sm., Novon 3: 488. 1993. Polypodium semihirsutum Klotzsch, P. semihirsutum (Klotzsch var. fuscosetosum Hieron., eae semihirsuta (Klotzsch) Copel., Grammitis semihirsuta (Klotzsch) ba rton, G semihirsuta (Klotzsch) C. V. Morton var. fuscosetosa (Hieron.) Tellinger Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 34 (LAGU, MO); R. Seiler 94, 101, 652 (MHES). Chalatenango: R. Seiler 751 (MHES) » Terpsichore taxifolia (L.) A. R. Sm., Novon 3: 488. 1981. Polypodium taxifolium L., P. lherminieri Fée var. costaricense Rosenst., Ctenopteris taxifolia (L.) Copel., Grammitis taxifolia (L.) Proctor Global distribution: Costa Rica, Panama, Antilles, Colombia, Venezuela, Suriname, Ecuador, Peru, Brazil, Bolivia. Source: Seiler (1982): 386 (as Grammitis taxifolia); Berendsohn (1989) 1: 16 (as Grammitis taxifolia); Moran & Smith in Davidse et al. (1995): PSILOTACEAE Psilotum complanatum Sw., J. Bot. (Schrader) 1800(2): 110. 1801. Local name: helecho palma de esquipulas. Global distribution: Mexico, Guatemala, Honduras, Antilles, Colombia, Venezuela, Peru, Malasia, Oceania. Salvadoran distribution: Ahuachapan: E. Sandoval s.n. (MS-00325) (B, LAGU, MO); E. Sandoval & F. Chinchilla 54 (B, LAGU); M. Sandoval & E. Sandoval 247 (B, LAGU, MO). San Vicente: Léetschert 252 (EAP) (Palacios-Rio in Davidse ef al. 1995). @Psilotum nudum (L.) P. Beauv., Prodr, Aethéogam. 106, 112. 1805. Lycopodium nudum L., Psilotum triquetrum Sw. Global distribution: USA, Mexico, Guatemala, Belice, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guyana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Old World tropics, Australia, New MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 188 Zealand. Salvadoran distribution: La Union: R. Ibarra, R. Juarez, F. Franco, I. Vega & O. Jiménez s.n. (JBL05114) (BM, LAGU, MHES); J. Monterrosa & R. Ibarra 1608 (BM, LAGU, MHES, MO). PTERIDACEAE Acrostichum aureum L., Sp. Pl. 2: 1069. 1753. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa ica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Brazil, Paraguay, Tropicos del Viejo Mundo. Salvadoran distribution: Ahuachapan: R. Seiler 137 (MHES); R. Villacorta & E.A. Montalvo 826 (B, LAGU, MEXU). La Unién: K. Sidwell et al. 674 (B, BM, ITIC, LAGU, MO) Acrostichum danaeifolium Langsd. & Fisch., Pl. Voy. Russes Monde I: 5, t. I. 1810. Chrysodium lomarioides Jenman, C. lomarioides Jenman var. hastatum Christ, Acrostichum lomarioides (Jenman) Jenman., A. excelsum Maxon Local names: helecho de manglar, helecho de pantano. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Ahuachapan: J.M. Rosales 2191 (B, LAGU). La Libertad: M. Magafia 48 (MP-00008) (LAGU). San Miguel: J. Tucker 908 (F); R. Villacorta 2889 (B, BM, LAGU). Adiantum amplum C. Presl, Relig. Haenk. I(1): 63. 1825. Adiantum princeps T. Moore Local names: helecho de arriero, centavito. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela. Salvadoran distribution: Ahuachapan: A.K. Monro ef al. 1925, 3642, 3700 (B, BM, ITIC, LAGU, MO); A. Sermefio 106 (JBL01020) (B, LAGU, MEXU, MO); J.M. Rosales 5 (AAU, B, BM, LAGU, MEXU, MO); 2415 (B, BM, LAGU, MEXU, MO): M. Sandoval & E. Sandoval 304 (B, F, LAGU); R. Seiler 938 (F); T, Croat 42060, 42061, 42095, 42102 (ITIC); W. Berendsohn 1311 (LAGU, MO); 1390, 1391 (B, LAGU, MO). Santa Ana: L. Lara 221 (MHES). La Libertad: A.K. Monro ef al. 3770 (B, BM, ITIC, LAGU, MO); B. Pfeiffer-Berendsohn 68 (B, LAGU, MO); E.A. Montalvo s.n. (ITIC); J. Monterrosa et al. s.n. (JBL04488) (LAGU); K. Sidwell eg al. 465 (B, BM, ITIC, LAGU, MO); N. Ventura 5012, 5015 (ITIC). San Salvador: K. Sidwell et al. 582 (B, BM, ITIC, LAGU, MO); R. Seiler s.n. (12-326) (MHES); 72 (F, MHES);: 711 (F). Cuseatlan: S. Dar s.n. (JBL03659) (B, BM, LAGU, MO). Morazan: J. Monterrosa & C. Rivera 675 (BM, LAGU). La Union: A.K. Monro et a/. 2753 (B, BM, ITIC, LAGU, MO). Adiantum andicola Liebm., Mexic. Bregn. 266 (reprint 114). 1849. Adiantum cuneatum Langsd. & Fisch. var. angustifolium M. Martens & Galeotti, A. tenerum Sw. var. dissectum M. Martens & Galeotti, 4. amabile Liebm., A. 189 FERN GAZ. 18(4): 120-215. 2008 glaucophyllum Hook. Local name: helecho centavito. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colomb Ese eal distribution: Ahuachapan: A.K. Monro et a/. 2003 (B, BM, ITIC, LAGU, MO). Sonsonate: T. Croat 42212 (ITIC). Santa Ana: A. Molina ef a/. 16881 (F); D. Rodriguez & G. Cerén s.n. (JBL04597) (B, LAGU); G. Davidse ef al. 37231 (B, BM, ITIC, LAGU, MO); J. Monterrosa et al. 917 (LAGU, NY); 1024 (LAGU); L. Lara 226 (MHES); R. Seiler 135, 277, 463 (F, MHES). La Libertad: M. Renderos et al. 413 (LAGU, MO). San Salvador: E.A. Montalvo 3155 (ITIC); M.C. Carlson 383, 472 (F):; R. Seiler 211, 517 (F, MHES); S. Winkler 18 (ITIC). Chalatenango: R. Seiler 404 (JBLO0080) (F, LAGU, MHES); 730 (F, MHES); W. Berendsohn & H. Sipman 1585 (B, LAGU). San Vicente: R. Seiler 320, 329 (F, MHES). Usulutan: D. Williams 136 (B, LAGU, MO). Morazan: J. Tucker 607 (F). Adiantum braunii Mett. ex Kuhn, Linnea 36: 75. 1869. Adiantum braunii Mett. ex Kuhn var. cuneatum Mett. ex Kuhn, A. convolutum E. Fourn. Local names: helecho nido de arriero, helecho nido de pijuyo. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Colombia, Venezuela, Ecuador. Salvadoran distribution: Ahuachapan: E. Sandoval & M. Sandoval 1401 (B, LAGU). Santa Ana: W. Berendsohn ef a/. 1536 (B, LAGU, MO). La Libertad: A.K. Monro er al. 3083 (B, BM, ITIC, LAGU, MO); J.C. Gonzalez & M.A. Hernandez 364 (B, LAGU, MO). San Salvador: R. Seiler 1502 (F). Chalatenango: R. Seiler 550 (MHES). Adiantum concinnum Humb. & Bonpl. ex Willd., Sp. Pl., ed. 4, 5(1): 451. 1810. Adiantum lutescens Moungeot ex Fée Local names: helecho cola de arriero, helecho nido de arriero, helecho palmita. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1915 (B, BM, ITIC, LAGU, MO); E. Sandoval & F. Chinchilla 704 (B, LAGU); E. Sandoval & M. Sandoval 1402 (LAGU); 1521 (B, LAGU); F. Chinchilla s.n. (ISB00534), s.n. (ISB00736), s.n. (ISB00768) (AAU, B, F, LAGU); M. Magafia 7 (MP-00002) (LAGU); M. Sandoval & E. Sandoval 101 (B, F, LAGU); R. Villacorta et a/. 942 (B, LAGU, MO); T. Croat 42115 (ITIC). Sonsonate: D. Rodriguez & M. Trejo 83 (LAGU). Santa Ana: L. Lara 222 (MHES); M.L. Reyna de Aguilar & R. Villacorta 1438 (B, LAGU, MO). La Libertad: A.K. Monro et al. 2335, 3084 (B, BM, ITIC, LAGU, MO); J. Reyes ef al. (JBL04510) (LAGU); J. Williams s.n. (ITIC); K. Sidwell e¢ al. 483, 698 (B, BM, ITIC, LAGU, MO); M.B. Siu s.n. (ITIC); N. Ventura 5007, 5011, 5025 (ITIC); R. Amador et al. s.n. (JBL04461) (LAGU); R. Menjivar & J. Sermefio s.n. (JBL04484) (LAGU); V. Salzman s.n. (JBL02066) (LAGU). San Salvador: A.K. Monro ef al.3437 (B, BM, ITIC, LAGU, MO); J. Williams 94 (ITIC); K. Sidwell et a/. 577 (B, BM, ITIC, LAGU, MO); M.C. Carlson 2 (F); R. Seiler 511 (F, MHES); Tobar s.n. (12-10) (MHES); W. Heed 25 (ITIC). Chalatenango: L. Lara 207, 214, 217 (MHES); M.B. Siu s.n. (ITIC); R. Seiler 141 (MHES). Usulutan: A.K. Monro et al. 3000 (B, BM, ITIC, LAGU, MO). San Miguel: J. Tucker 976 (F). Cabafias: A.K. Monro et al. 2775 (B, BM, ITIC, LAGU, MO); G. Davidse et al. 37111 (B, BM, ITIC, LAGU, MO); R. Seiler 527 (F, MHES). Morazan: J. Tucker 640 (F). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 190 » Adiantum deflectens Mart., Icon. Pl. Crypt. 94. 1834. Adiantum deflectens Mart. var. tremulum Hieron., A. delicatulum Mart., A. dissimile Schrad., A. dolabriforme Hook., A. filiforme Gardner, A. flagellum Fée, A. lunatum Keyserl., A. subaristatum Fée Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay. Source: Stolze (1981): 22; Berendsohn (1989) 1: 19; Moran in Davidse et al. (1995): 111; Mickel & Smith (2004): 29. Adiantum latifolium Lam., Encycl. |: 43. 1783. Adiantum lucidum Sw. var. bipinnatum Mett. ex E. Fourn. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Africa, Asia. Local names: helecho rosquilla Salvadoran distribution: Ahuachapan: J.M. Rosales 1916 (AAU, B, BM, LAGU, MO). Usulutan: J. Lagos 1084 (ITIC); R. Seiler 591 (F, MHES). Adiantum lunulatum Burm. f., Fl. Indica 235. 1768. Local names: helecho cola de arriero, helecho pie de zanate, helecho salamanquera. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Africa, Asia. Salvadoran distribution: Ahuachapan: E. Sandoval et a/. 609 (B, LAGU); 1398 (B, K, LAGU); F. Chinchilla s.n. (ISB00605), s.n. (ISB00769) (AAU, B, F, LAGU); J.M. Rosales 1133, 1134, 1167, 1316, AAU (B, BM, LAGU). Sonsonate: D. Rodriguez & M. Trejo 64, B (LAGU, NY). Santa Ana: J.L. Linares 535 (JBL01985) (LAGU); K. Prestegard 43 (MP-00003) (LAGU): R. Seiler 709 (MHES); W. Berendsohn et al. 1539 (B, LAGU). La Libertad: A.K. Monro er a/. 3404 (B, BM, ITIC, LAGU, MO); N. Ventura 5006 (ITIC). San Salvador: R. Seiler 376, 446 (F, MHES). La Paz: E.A. Montalvo 6205 (B, BG, F, HBG, ITIC, LAGU, MO). San Miguel: R. Seiler 544 (F, MHES). Cabaiias: R. Seiler 524 (F, MHES). Morazan: J. Tucker 452 (F). Adiantum macrophyllum Sw., Prod. 135. 1788. Local name: helecho palma de esquipulas, helecho palma de roca. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval et a/. 106 (LAGU); s.n (JBL01386), 1293 (B, LAGU); F. Chinchilla s.n. (ISB00660) (AAU, B, F, LAGU); M. Sandoval ef al. s.n. (ISB00842) (AAU, B, F, LAGU); 273 (B, F, LAGU); P. Standley ef al. 2673 (F); R. Seiler 932 (MHES). La Libertad: D. Matamoro s.n. (JBL01317) (LAGU); M.C. Carlson 143 (F); R. Seiler 717 (MHES). San Salvador: E.A. Montalvo I5a (ITIC); R. Seiler 449, 508, 530 (F, MHES). Adiantum patens Willd., Sp. Pl., ed. 4, 5(1): 439. 1820. Adiantum lobatum C. Pres! Local names: helecho nido de arriero, helecho pata de paloma, petatillo rojo. Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Colombia, Venezuela, Ecuador, Peru, Bolivia, Africa. 19] FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: Ahuachapan: E. Sandoval et a/. 589, 1362 (B, LAGU). Santa Ana: R. Seiler 133 (MHES); 761 (F, MHES); T. Croat 42288 (ITIC). La Libertad: J. Monterrosa ef al. s.n. (JBL04500) (LAGU); N. Ventura 5018 (ITIC). San Salvador: R. Seiler 448, 636 (F, MHES). Chalatenango: D. Sloot & A. Reina 135 (MAG00564) (B, LAGU, U). Morazan: J. Tucker 641 (F); S. Winkler 19 (ITIC). Adiantum petiolatum Desy., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk 5: 326. 1811. Adiantum lucidum (Cav.) Sw. var. pinnatum E. Fourn. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Chalatenango: R. Seiler 138 (F, ITIC, MHES). San Vicente: R. Seiler 973 (F, ITIC, MHES). Adiantum poiretii Wikstr., Kongl. Vetensk. Akad. Handl. 1825: 443. 1826. Adiantum thalictroides Willd. ex Schltdl., A. pellucidum M. Martens & Galeotti, A. gratum Fée Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia, Chile, Argentina, Viejo Mundo. Salvadoran distribution: Santa Ana: R. Villacorta & L. Lara 2517 (B, LAGU). Chalatenango: W. Berendsohn & H. Sipman 1586 (B, LAGU). Adiantum pulverulentum L., Sp. P|. 2: 1096. 1753. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1973 (B, BM, ITIC, LAGU, MO); D. Witsberger 702 (MHES); R. Seiler 818 (F, ITIC, MHES). Sonsonate: D. Rodriguez & M. Trejo 205 (LAGU, NY). Adiantum raddianum C. Presl, Suppl. Tent. Pterid. 158. 1836. Adiantum cuneatum Langsd. & Fisch., A. mexicanum C. Presl, A. werckleanum Christ Local name: helecho colchon de guardabarranco. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, guay. Salvadoran distribution: Ahuachapan: M. Magafia 14 (MP-00001) (LAGU); W. Berendsohn ef al. 1392 (B, LAGU). Usulutan: A.K. Monro er al. 3024 (B, BM, ITIC, LAGU, MO). Note: A.K. Monro & D. Alexander 3084 is cited erroneously as A. raddianum in Monro et al. 2002, it is A. concinnum Humb. & Bonpl. ex Willd. > Adiantum tenerum Sw., Prodr. 135. 1788. Adiantum trapezoides Fée Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Antilles, Venezuela, Trinidad and Tobago. Source: Maxon & Standley (1930): 172 (as Adiantum trapezoides). Calderon & MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 192 Standley (1941): 29 (as Adiantum trapezoides). Seiler (1982): 384. Berendsohn (1989) 1: 20. Moran et al. in Davidse et al. (1995): 110. Adiantum tetraphyllum Humb. & Bonpl. ex Willd., Sp. Pl., ed. 4, 5(1): 441. 1810. Adiantum fruticosum Poepp. ex Spreng., A. tetraphyllum Humb. & Bonpl. ex Willd. var. costaricense Christ Local names: helecho ciempiés, helecho rosquilla pacha. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Bolivia, Paraguay. Salvadoran distribution: Ahuachapan: E. Sandoval et al. 1119, 1207 (B, LAGU, MO); F. Chinchilla s.n. (ISB0O0315) (LAGU); s.n. (ISB00661) (AAU, B, F, LAGU); M. Sandoval et al. s.n. (ISBO0815), AAU, B, F, AAU; 298 (B, F, LAGU); R. Morales et al. s.n. (JBL01274) (B, LAGU, MO); R. Seiler 924 (F, ITIC, MHES); R. Villacorta 623 (B, LAGU, MO); W. Berendsohn ef al. 1355, 1369, 1398 (B, LAGU, MO); 1428 (B, LAGU). Santa Ana: R. Villacorta & J. Maria 2203 (B, LAGU, MO). San Salvador: R. Seiler 531 (F, MHES). Chalatenango: R. Seiler 265 (F, MHES). Cabafias: A.K. Monro et al. 3747 (B, BM, ITIC, LAGU, MO); J. Monterrosa & R. Carballo 230 (AAU, B, BM, LAGU, MO). Morazan: J. Monterrosa & C. Rivera 673 (B, BM, LAGU, MO). Adiantum trapeziforme L., Sp. Pl. 2: 1097. 1753. Adiantum trapeziforme L. var. oblongatum T. Moore, A. trapeziforme L. var. plumieri T. Moore Local names: cabello de angel, helecho nido de arriero, lantrillo. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Antilles, Trinidad and Tobago, Bolivia, Paraguay; naturalised in Asia and the Pacific Islands. Salvadoran distribution: Anuachapan: E. Sandoval & F. Chinchilla 177, 266 (LAGU): F. Chinchilla & R.E. Sandoval s.n. (ISB00589) (AAU, B, F, LAGU): M. Sandoval & E. Sandoval s.n. (ISB00838) (AAU, B, LAGU); P. Standley et a/. 3024 (F); R. Seiler 937 (F, MHES); T. Croat 42103 (ITIC); W. Berendsohn er al. 1434 (B, LAGU, MO). Santa Ana: A.K. Monro & R. Douglas 3464 (B, BM, ITIC, LAGU, MO). La Libertad: O. Pank s.n. (MP-00004) (LAGU). San Salvador: K. Sidwell et a/. 580 (B, BM, ITIC. LAGU, MO); M.C. Carlson 20, 327 (F); R. Seiler 67 (F. ITIC, MHES); 785 (F, MHES). Chalatenango: R. Seiler 139 (F, MHES). Usulutan: E.A. Montalvo 5101 (ITIC); S. Winkler 16 (ITIC). Cuseatlan: S. Dar s.n. (JBL03658) (LAGU, MO). La Union: A.K. Monro et al. 3536 (B, BM, ITIC, LAGU, MO). Adiantum trichochlaenum Mickel & Beitel. Mem. New York Bot. Gard. 46: 29-30. 1988 Global distribution: Mexico, Guatemala. Nicaragua, Costa Rica. Salvadoran distribution: San Salvador: R. Seiler 455 (F, MHES). Adiantum villosum L.. Syst. Nat., ed. 10, 2: 1328. 1759. Local names: helecho ciempiés, helecho rosquilla pacha. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 486 (B, LAGU); ios FERN GAZ. 18(4): 120-215. 2008 J.M. Rosales 1419 (LAGU); M. Sandoval er al. 29 (B, LAGU); T. Croat 42087 (ITIC); W. Berendsohn & R. Villacorta 1350 (LAGU, MO). Santa Ana: M.L. Reyna de Aguilar 663 (MHES). La Libertad: R. Seiler 716 (F, MHES). Chalatenango: R. Seiler 140 (F, MHES). San Vicente: M.C. Carlson 645 (F). Usulutan: R. Seiler 592 (F, MHES). San Miguel: A. Salgado 71 (MHES); J. Tucker 848 (F). Cabafias: R. Seiler 225 (JBL04805) (F, LAGU, MHES). Ananthacorus angustifolius (Sw.) Underw. & Maxon in Maxon, Contr. U.S. Natl. Herb. 10: 487. 1908. Pteris angustifolia Sw., Taenitis angustifolia (Sw.) Spreng., Vittaria angustifolia (Sw.) Baker in Martius, Vittaria costata Kunze Local name: helecho barbiquejo. Global distribution: Mexico, Guatemala, Belize, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval et al. 439, 1303 (B, LAGU); J.M. Rosales 1466 (B, BM, LAGU, MO); 1513 (B, LAGU, MO); M.L. Reyna de Aguilar 652 (MHES): M. Sandoval & E. Sandoval s.n. (ISB00930) (LAGU). San Salvador: R. Seiler 459, 788 (F, MHES). Anogramma chaerophylla i Link, Fil. Spec. 138. 1841. Gymnogramma chaerophylla Global distribution: Mexico, Antilles, Brazil, Paraguay, Argentina, Urugua Salvadoran distribution: San Salvador: H. Weber 24 (ITIC); R. Seiler 20 (MHES). Anogramma leptophylla (L.) Link, Fil. Spec. 137. 1841. Polypodium leptophyllum L., Gymnogramma leptophylla (L.) Desv. var. mexicana Kunze, Pityrogramma guatemalensis Domin, Anogramma guatemalensis (Domin) C. Chr. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Venezuela, Ecuador, Peru, Brazil, Eurasia, Africa. Salvadoran distribution: Sonsonate: R. Seiler 610, 616, 629 (F, MHES). Santa Ana: .A. Montalvo 4895 (ITIC); J. Williams 31 (ITIC); R. Seiler 110 (F, ITIC, MHES); 658 (F, MHES). San Salvador: Castafieda s.n. (12-897) (MHES). Usulutan: D. Williams & R.W. Herrera 424 (LAGU). Bommeria pedata (Sw.) E. Fourn., Bull. Soc. Bot. France 27: 327. 1880. Hemionitis pedata Sw., Gymnogramma pedata (Sw.) Kaulf., Gymnopteris pedata (Sw.) ‘le Local name: helecho mano de mapache. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: M. Sandoval & E. Sandoval 40 (B, F, LAGU). Santa Ana: R. Seiler 498 (F, MHES). San Salvador: R. Seiler 46 (F, MHES):; 640 (MHES); O. Pank s.n. (12-987) (MHES). Ceratopteris pteridoides (Hook.) Hieron., Bot. Jahrb. Syst. 34: 561. 1905. Parkeria pteridoides Hook. Local name: helecho acuatico. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 194 Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: J.M. Rosales 2674 (B, LAGU, MO). La Paz: N.C. Fassett 28557 (EAP, F); 29180 (F). San Miguel: N.C. Fassett 29391 (F). Ceratopteris thalictroides (L.) Brongn., Bull. Sci. Soc. Philom. Paris, sér. 3, 8: 186. Acrostichum thalictroides L. Local name: helecho de agua. Global distribution: USA, Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Venezuela, Suriname, Ecuador, Brazil, Africa, Asia, Australia, Pacific Islands. Salvadoran distribution: Ahuachapan: J.M. Rosales 1921 (B, LAGU, MO); 2070 (LAGU). San Vicente: N.C. Fassett 29157 (F); 29162 (EAP, F). Usulutan: R.A. Carballo & S. Santamaria 899 (B, LAGU, MO). Cheilanthes angustifolia Kunth, Nov. Gen. Sp. I: 21. 1815. Allosorus angustifolius (Kunth) C. Presl, A. angustifolius (Kunth) C. Presl var. minor Liebm., Onvchium angustifolium (Kunth) Kunze, Pellaea angustifolia (Kunth) Baker in Hooker & Baker, Cheilanthes venusta Fée var. aurata Fée Local name: helecho tuya morada. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 664 (B, LAGU); J.M. Rosales 1230 (B, LAGU, MO); R. Villacorta & Martinez 626 (B, LAGU, MO); W. Berendsohn & R. Villacorta 1384 (B, LAGU, MO). Sonsonate: R. Seiler 37 (F, MHES). Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 13 (F, LAGU, MO); S. Winkler 28 (ITIC). La Libertad: E.A. Montalvo s.n. (ITIC). San Salvador: H. Castafieda s.n. (12-899) (MHES); E.A. Montalvo s.n. (ITIC). Chalatenango: L. Lara 213, 231 (MHES); R. Seiler 559 (F, MHES); S. Calderén 2476 (F). San Vicente: E.A. Montalvo 5095 (ITIC). Usulutan: D. Williams & R.W. Herrera 301 (LAGU, MO). Morazan: J. Monterrosa & C. Rivera 689 (B, BM, LAGU, MO); M. Renderos ef al. 611 (LAGU, MO). Cheilanthes aurea Baker in Hooker & Baker var. aurea, Syn. Fil. 453. 1868. Aleuritopteris aurea (Baker) Ching Global distribution: Mexico, Guatemala. Salvadoran distribution: Ahuachapan: J.M. Rosales 1229 (B, LAGU); W. Berendsohn et al. 1644 (BM, LAGU). San Salvador: R. Seiler 23, 586 (F, MHES). Cheilanthes bolborrhiza Mickel & Beitel, Mem. New York Bot. Gard. 46: 107. 1988. Global distribution: Mexico. Salvadoran distribution: San Salvador: R. Seiler 45, 579 (F, LAGU, MHES, NY). Cheilanthes brachypus Kunze, Linnea 23: 307. 1850. Cheilanthes squamosa Hook. & Grev. var. brachypus Kunze, Notholaena brachypus (Kunze) J. Sm. Local names: helecho de bulto, helecho dorado, helecho pluma de buho. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 3644 (B, BM, ITIC, LAGU, MO); E. Sandoval & F. Chinchilla 468 (B, LAGU, MO); E. Sandoval & M. Sandoval 195 FERN GAZ. 18(4): 120-215. 2008 1404 (B, LAGU); J.M. Rosales 1289, 1587 (AAU, B, BM, LAGU, MO); M. Sandoval et al. 61 (B, F, LAGU); R. Seiler 926 (F, MHES); R. Villacorta & Martinez 404 (F, LAGU, MO); T. Croat 42155 (ITIC); W. Berendsohn et al. 1408 (B, LAGU, MO). Sonsonate: R. Seiler 7 (F, MHES). Santa Ana: K. Prestegard 30 (LAGU); T. Croat 42238 (ITIC). La Libertad: A.K. Monro et a/. 2919 (B, BM, ITIC, LAGU, MO); E.A. Montalvo & R. Villacorta 6266 (B, LAGU, MO); J. Flores s.n. (JF-00106) (LAGU); R. Villacorta s.n. (JBL00543) (B, LAGU, MO). San Salvador: R. Seiler 444 (F, MHES). Chalatenango: L. Lara 212, 216 (MHES); R. Seiler 367, 551 (F, MHES). San Vicente: O. Rohweder 2643 (JBL01248), 2644 (JBL01247) (LAGU). San Miguel: A. Salgado 63 (MHES); R. Seiler 540 (F, MHES); R. Villacorta 2391 (LAGU, MO). Morazan: J. Monterrosa & C. Rivera 709 (B, BM, LAGU, MO). La Union: A.K. Monro et al. 3540 (B, BM, ITIC, LAGU, MO). > Cheilanthes chaerophylla (M. Martens & galeotti) Kunze, Linnea 23: 243, 307. 1850. Allosorus chaerophyllus M. Martens & Galeotti Global distribution: Mexico, Guatemala, Honduras, Costa Rica. Salvadoran distribution: Chalatenango: J. Monterrosa et a/. 1611 (BM, LAGU, NY). Cheilanthes decurrens Mickel, Mem. New York Bot. Gard. 88: 191-192. 2004. Global distribution: Mexico, Guatemala. pope distribution: Santa Ana: R. Seiler 100 (F, MHES). San Salvador: R. Seiler 1209 (F, N Cheilanthes kaulfussii Kunze, Linnea 13: 145. 1839. Cheilanthes viscosa Link, C. glandulifera Liebm. Global distribution: USA, Mexico, Guatemala, Honduras, Costa Rica, Colombia, Venezuela Salvadoran distribution: Sonsonate: R. Seiler 10 (F, MHES): 1466 (F). Santa Ana: R. Seiler 680 (F, MHES). Usulutan: D. Williams & R.W. Herrera 348 (LAGU). Morazan: R. Seiler 1074 (F, MHES). Cheilanthes marginata Kunth, Nov. Gen. Sp. I: 22. 1815; 7: t. 669. 1825. Allosorus ciliata C. Presl var. minor C. Presl, C. hirsuta Link, C. rufescens Link, C. crenulata Link Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, Argentina. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn 47 (LAGU, MO); 54 (B, O); R. Seiler 298, 473 (F, MHES); 1208 (F); R. Villacorta & J.C. Gonzalez 1160 (LAGU). Chalatenango: R. Seiler 383 (F, MHES); W. Berendsohn & H. Sipman 1582 (B, LAGU). Morazan: R. Seiler 986 (F, ITIC, MHES); 1069 (F, MHES); 1071 UF). Note: Seiler (1982), Berendsohn (1989) and R.C. Moran in Davidse et al. 1995 recognise Cheilanthes hirsuta as a distinct species. > Cheilanthes myriophylla Desy., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 328. 1811. Cheilanthes paleacea M. Martens & Galeotti, C. intermedia Baker, C. fournieri C. Chr., MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 196 Myriopteris intermedia E. Fourn. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Chile, Argentina. Salvadoran distribution: R. Seiler 1471 (MO) (Yatskievych & Moran in Davidse et al. 1995). Cheilanthes pyramidalis Fée, Mém. Foug. 7: 38, t. 25 (F). 3. 1857. Pellaea angustifolia (Kunth) Baker var. elongata Rovirosa Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Santa Ana: B. Pfeiffer-Berendsohn & S.A. de Reyna 39 (K, LAGU); M.L. Reyna de Aguilar 766 (MHES); R. Seiler 42 (JBL04800) (F, ITIC, LAGU, MHES); 351 (F, MHES). San Salvador: R. Seiler 24 (F, MHES). Cheilanthes skinneri (Hook.) R. M. Tryon & A. F. Tryon, Rhodora 83: 133. 1981. Pellaea skinneri Hook., P. flavescens E. Fourn., Dorvopteris fournieri (Baker) C. Chr. Local names: helecho cola de pavo real, helecho morado. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia. Salvadoran distribution: Ahuachapan: E. Sandoval ef al. 521, 1400 (B, LAGU): J.M. Rosales 934, 951 (LAGU). Santa Ana: R. Seiler 1201 (F, ITIC, MHES). La Libertad: R. Seiler 1116 (F, ITIC, MHES). San Miguel: R. Seiler 538 (JBL04808) (F, LAGU, MHES). Cheiloplecton rigidum (Sw.) Fée var. rigidum, Mém. Foug. 7: 34, t. 20 (F). 3. 1857. Pteris rigida Sw., P. cartilaginea C. Presl, Pellaea rigida (Sw.) Hook. Global distribution: Mexico, Guatemala. Salvadoran distribution: Santa Ana: R. Seiler 1190 (F, MHES). Doryopteris concolor (Langsd. & Fisch.) Kuhn in v.d. Decken, Reisen Ost-Afrika 3(3): 19. 1879. Pteris concolor Langsd. & Fisch., Cheilanthes concolor (Langsd. & Fisch.) R. M. Tryon & A. F. Tryon . Local name: helecho de roca. Global distribution: Mexico, Guatemala, Honduras, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil Bolivia, Paraguay, Argentina, Uruguay, Africa, Asia, Australia. Salvadoran distribution: Sonsonate: M.L. Reyna de Aguilar 1452 (LAGU, MO). San Salvador: R. Seiler 585 (F, MHES). San Miguel: R. Seiler 539 (F. MHES): R. Villacorta 2481 (B, LAGU, MO). Eriosorus flexuosus (Desv.) Copel. var. flexuosus, Gen. Fil. 58. 1947. Gymnogramma flexuosa Desv., Grammitis flexuosa Kunth Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: J. Lagos 13 (ITIC); R. Seiler 177 (JBLO4804) (F, LAGU, MHES); R. Villacorta et a/. 1048 (B, LAGU, MO); W. Berendsohn 1069 (B, BM, LAGU, MO). Chalatenango: J. Tucker 1120 (F). Hemionitis levyi E. Fourn., Bull. Soc. Bot. France 17: 237. 1870. Hemionitis otonis Maxon Global distribution: Mexico, Nicaragua, Costa Rica. 197 FERN GAZ. 18(4): 120-215. 2008 Salvadoran distribution: S. Calderon 1710, MO (Ranker in Davidse ef al. 1995). Ahuachapan: J.M. Rosales 1187 (B, LAGU). San Miguel: R. Villacorta 2392 (B, LAGU, MO). > Hemionitis palmata L., Sp. P|. 2: 1077. 1753. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: P.Standley 19894 (GH) (Ranker in Davidse ef al. 1995). Source: Maxon & Standley 1930: 173; Calderon & Standley (1941): 31; Seiler (1982): 383: Berendsohn (1989) 1: 21; Ranker in Davidse ef al. (1995): 132; Mickel & Smith (2004): 326. Hemionitis pinnatifida Baker in Hooker & Baker, Syn. Fil. 399. 1868. Local names: helecho ostra. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 469 (B, LAGU). Sonsonate: R. Seiler 493 (MHES). Santa Ana: O. Rohweder 2641 (JBL01249), 2689 (JBL0O1251), 2690 (JBL01250) (LAGU). La Libertad: W. Berendsohn 1024 (LAGU, MO). San Salvador: M. Renderos 387 (B, LAGU, MO); O. Pank s.n. (12-959) (MHES); R. Seiler 442 (F, MHES); 505 (MHES). Chalatenango: R. Villacorta 2394 (AAU, B, LAGU, MO). San Miguel: R. Seiler 59 (F, MHES). Morazan: J. Monterrosa & R. Carballo 535 (B, LAGU, MO). Mildella intramarginalis (Kaulf. ex Link) Trevis., Rendiconti Reale Ist. Lombardo Sci., Cl. Sci. Mat., ser. 2, 9: 810. 1876. Pteris intramarginalis Kaulf. ex Link, Cheilanthes intramarginalis (Kaulf. ex Link) Hook., Pellaea intramarginalis (Kaulf. ex Link) J. Sm. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Galapagos. Salvadoran distribution: Sonsonate: R. Seiler 9 (F, MHES). Santa Ana: B. Pfeiffer- Berendsohn & W. Berendsohn 14, 16 (LAGU); G. Davidse et al. 37247 (B, BM, ITIC, LAGU, MO); J. Lagos 1422 (ITIC); J. Monterrosa et al. 223 (B, BM, LAGU, MO); 994 (LAGU); M.B. Siu s.n. (ITIC); N. Lopez er al. s.n. (JBL04635) (LAGU); R. Seiler 352 (F, MHES); 898 (F); R. Villacorta et al. 1159 (LAGU); 1052, 1257 (B, LAGU, MO). San Salvador: A.K. Monro et al. 2234 (B, BM, ITIC, LAGU, MO); M.C. Carlson 424 (F); R. Seiler 30 (JBL04799) (F, LAGU, MHES); 58 (F). Chalatenango: L. Gonzalez 1711 (ITIC); L. Lara 215 (MHES); R. Seiler 577 (F, MHES). San Vicente: J. Lagos 66 (ITIC). Usulutan: D. Williams 57, 60 (B, LAGU, MO). Morazan: R. Seiler 1088b (F, ITIC, MHES). Notholaena candida (M. Martens & Galeotti) Hook., Sp. Fil. 5: 110. 1864. Cheilanthes candida M. Martens & Galeotti, Aleuritopteris candida (M. Martens & Galeotti) Fée, Chrvsochosma candida (M. Martens & Galeotti) Kiimmerle Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Sonsonate: R. Seiler 497 (F, MHES). San Miguel: A. Salgado 62 (MHES) MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 198 Notholaena meridionalis Mickel, Mem. New York Bot. Gard. 88: 421-422. 2004. Global distribution: Mexico, Guatemala, Nicaragua, Costa Ri Salvadoran distribution: Ahuachapan: A.K. Monro et al. 3645 (B, BM, ITIC, LAGU, MO). Santa Ana: R. Seiler 88 (F, MHES, NY). Pellaea ternifolia (Cav.) Link. subsp. ternifolia, Fil. Spec. 59. 1841. Preris ternifolia Cav. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Chile, Paraguay, Argentina, Hawaii. Salvadoran distribution: Santa Ana: R. Seiler 679 (MHES); 467 (F). Pityrogramma calomelanos (L.) Link var. calomelanos, Handbuch 3: 20. 1833. Acrostichum calomelanos L., Gymnogramma calomelanos (L.) Kaulf., Ceropteris serrata Fée Local name: helecho pinta mano. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: E. Sandoval ef a/. 279 (B, LAGU, MO); 1103, 1301 (B, LAGU): F. Chinchilla s.n. (ISB00093) (B, LAGU, MO); J.M. Rosales 117, 1228 (B, LAGU); 788 (B, LAGU, MO): M. Magafia 8 (MP-00006) (LAGU); M. Sandoval et a/. 103, 312 (B, F, LAGU); W. Berendsohn ef a/. 1335 (B, LAGU, MO). Sonsonate: M.B. Siu s.n. (ITIC); M.L. Reyna de Aguilar 586 (MHES); R. Seiler 631 (F, MHES). Santa Ana: M.C. Carlson 841 (F); M.L. Reyna de Aguilar & R. Villacorta 1440 (B, LAGU, MO); R. Seiler 228 (F, MHES). La Libertad: K. Sidwell et al. 468 (B, BM, ITIC, LAGU, MO); N. Ventura 5014 (ITIC); R. Guerra ef al. s.n. (JBL04503) (LAGU); R. Menjivar & J. Sermefio s.n. (JBL04494) (LAGU). San Salvador: K. Sidwell et a/. 563 (B, BM, ITIC, LAGU, MO); M.C. Carlson 3 (F); N.C. Fassett 28599 (ITIC); R. Seiler 57 (F, MHES); S. Winkler 30 (ITIC); W. Eggler 680 (ITIC); W. Heed 33 (ITIC). La Paz: J.C. Gonzalez 102 (B, LAGU, MO); R. Villacorta ef al. 2854 (B, LAGU). San Vicente: P. Standley et a/. 3689 (F). Usulutan: A. del Cid 1941 (ITIC): D. Williams 14, 297 (LAGU). San Miguel: A. Salgado 75 (MHES); J. Tucker 977 (F). Morazan: R. Seiler 1047 (F). Pityrogramma dealbata (C. Pres!) Domin, Rozpr. Ceské Akad. Ved., Tr. 2, Vedy Mat. Prir, 51(15): 7. 1942 Gynnagnemns dealbata C. Presl, Ceropteris schaffneri Fée, Pitvrogramma schafjneri (Fée) Weath. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Ahuachapan: N.C. Fassett 28718b (F); S. de Moore s.n. (12- 916) (MHES). Sonsonate: J. Tucker 1340 (F). Santa Ana: A. Molina ef a/. 16844 (F); M.B. Siu s.n. (ITIC). La Libertad: N. Ventura 5019 (ITIC); V. Hellebyuck s.n. (12-92) (F, MHES). San Salvador: A. Molina ef al. 21829 (F, ITIC); M.C. Carlson 464 (F); R. Seiler 66 (F, MHES). Chalatenango: L. Lara 208, 219 (MHES). San Miguel: A. Salgado s.n. (JBL04801) (LAGU). Morazan: A. Salgado 62 (MHES); J. Monterrosa et al. 43 (B, BM, LAGU, MO); 671 (BM, LAGU). ea, FERN GAZ. 18(4): 120-215. 2008 Pityrogramma ebenea (L.) Proctor, Brit. Fern Gaz. 9: 219. 1965. Acrostichum ebeneum L., A. tartareum Cav., Pityrogramma tartarea (Cav.) Maxon, P. tripinnata Domin Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Sonsonate: R. Seiler 630 (F, MHES). Santa Ana: J. Monterrosa et a/.1029 (LAGU, NY); M.L. Reyna de Aguilar 669 (MHES); R. Seiler 40 (F, MHES); 462 (F, ITIC, MHES). San Vicente: S. Winkler s.n. (ITIC). Pityrogramma ferruginea (Kunze) Maxon, Contr. U.S. Natl. Herb. 17: 173. 1913. Gymnogramma ferruginea Kunze, G. bommeri Christ Global distribution: Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Peru. Salvadoran distribution: Sonsonate: W. Eggler 606 (ITIC). Santa Ana: R. Seiler 499 (F, ITIC, MHES). San Salvador: A.K. Monro et al. 2222 (B, BM, ITIC, LAGU, MO); M.C. Carlson 448 (F). Morazan: J. Monterrosa ef al. 11 (LAGU, MO). @ Pityrogramma ochracea (C. Presl) Domin, Spisy Prir. Fak. Karlovy Univ. 88: 8. 1928 Gymnogramma_ ochraceae C. Presl, Acrostichum caudatum Cay., Pityrogramma calomelanos (L.) Link var. ochracea (C. Presl) R. M. Tryon Global distribution: Honduras, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Usulutan: D. Williams 139 (B, LAGU, MO); 427 (LAGU). Pityrogramma trifoliata (L.) R. M. Tryon, Contr. Gray Herb. 189: 68. 1962. Acrostichum trifoliatum L., Trismeria trifoliata (L.) Diels in Engl. & Prantl Local names: helecho vertical. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Chile, Paraguay, Argentina, Uruguay. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 431 (B, LAGU): K. Prestegard 49 (MP-00007) (LAGU). Santa Ana: M.C. Carlson 840 (F); P. Standley et al. 3254 (F); R. Seiler 1207 (F, MHES). La Libertad: L. Williams & A. Molina 15192 (JBLO15499), LAGU (F). San Salvador: F. Lopez s.n. (ITIC); R. Seiler 518 (F, MHES). Chalatenango: F. Wunderlin 717 (ITIC); R. Seiler 264 (MHES); R. Villacorta et al. 2086 (B, LAGU, MO). > Polytaenium cajenense (Desv.) Benedict, Bull. Torrey Bot. Club 38: 169. 1911. Hemionitis cajenense Desv., Antrophyum cajenense (Desv.) Spreng., A. lacantunense Rovirosa Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles; Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Source: Moran in Davidse ef al. (1995): 146 (as Antrophyum cajenense), Mickel & Smith (2004): 526. Pteris altissima Poiret in Lamarck, Encycl. 5: 722. 1804. Pteris kunzeana J. Agardh, P. protea Liebm., Litobrochia kunzeana (J. Agardh) Fée, L. grandis Fée MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 200 Local names: helecho cola de pavo real, helecho palma de cristo. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: A.K. Monro e¢ al. 1978 (B, BM, ITIC, LAGU, MO); E. Sandoval & R.Chinchilla 1167 (B, LAGU): F. Chinchilla s.n. ([SB00330) (LAGU); R. Seiler 944 (MHES): W. Berendsohn et a/. 1425 (B, LAGU, MO). Santa Ana: S. Winkler 31 (ITIC). San Salvador: R. Seiler 536 (MHES). Pteris biaurita L., Sp. Pl. 2: 1076. 1753. Local names: helecho cola de ganzo, helecho palma real. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Old World Tropics. Salvadoran distribution: Ahuachapan: J.M. Rosales 490 (AAU, B, BM, LAGU, MO); M. Sandoval & E. Sandoval s.n. (ISB00873) (AAU, B, F, LAGU); R. Seiler 921 (ITIC, MHES); W. Berendsohn ef a/. 1399 (B, LAGU, MO); 1423 (LAGU); 1426 (B, LAGU). Sonsonate: D. Rodriguez 157 (LAGU). La Libertad: A.K. Monro ef a/. 2314 (B, BM, ITIC, LAGU, MO); J. Monterrosa et a/. s.n. (JBL04489) (LAGU): K. Sidwell et al. 699 (B, BM, ITIC, LAGU, MO); N. Ventura 5009 (ITIC); O. Tejada et a/. s.n. (JBL04511) (LAGU). San Salvador: B. Lopez s.n. (ITIC). Cabafias: A.K. Monro ef a/. 3751 (B, BM, ITIC, LAGU, MO); G. Davidse et a/. 37119 (B, BM, ITIC, LAGU, MO). Pteris cretica L., Mant. Pl. 130. 1767. Preris triphylla M. Martens & Galeotti, P. trifoliata Fée Local name: helecho pié de zenzontle. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Peru, Brazil, Argentina, Old World Tropics and Subtropics. Salvadoran distribution: La Libertad: E.A. Montalvo 6227 (B, LAGU). San Salvador: R. Seiler 222 (MHES). Chalatenango: R. Seiler 377 (JBL00206) (LAGU, HES). @ Pteris erosa Mickel & Beitel, Mem. New York Bot. Gard. 46: 322-323. 1988. Global distribution: Mexico, Guatemala. Salvadoran distribution: Sonsonate: R. Seiler 622 (MHES). Santa Ana: D. Rodriguez & G Cerén s.n. (JBL04599) (LAGU); J. Monterrosa ef al. 206 (B, LAGU, MO): R. Seiler 246 (MHES); V.M. Martinez s.n. (CMC00844) (B, LAGU). San Salvador: R. Seiler 212 (F, MHES). San Vicente: R. Seiler 316 (MHES). Note: Moran in Davidse et al. (1995) considered this name a synonym of Preris orizabae M. Martens et Galeotti. Mickel & Smith (2004) consider P. orizabae and Pteris erosa distinct based on differences in indusium margin and other characters. Pteris grandifolia L., Sp. P|. 2: 1073. 1753. Litobrochia grandifolia (L.) J. Sm., Heterophlebium grandifolium (L.) Fee Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil. Salvadoran distribution: P. Standley 21242, MO (Moran in Davidse er al. 1995). San Vicente: R. Seiler 974 (ITIC, MHES). 201 FERN GAZ. 18(4): 120-215. 2008 Pteris muricella Fée, Mém. Foug. 8: 73. 1857. Pteris mollis Christ Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Venezuela. Salvadoran distribution: Ahuachapan: R. Seiler 904 (F, ITIC, MHES). Pteris orizabae M. Martens et Galeotti, Foug. Mexique 53, pl. 13. 1842. Pteris apicaulis Liebm., P. hemipteris (Fée) Hook., Litobrochia orizabae (M. Martens & Galeotti) Fée, L. setifera Fée, L. hemipteris Fée Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: T. Croat 42417, MO (Moran in Davidse et al. 1995). Santa Ana: J. Monterrosa & R. Martinez 945 (B, LAGU, MO); R. Seiler 272, 692 (MHES). Morazan: R. Seiler 1032 (ITIC, MHES). Pteris paucinervata Fée, Mém. Foug. 8: 73. 1857. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Chalatenango: R. Seiler 408 (JBL00205) (F, LAGU, MHES). Pteris podophylla Sw., J. Bot. (Schrader) 1800(2): 67. 1801. Lonchitis pedata L. Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Suriname, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: R. Seiler 675 (F, MHES). Pteris quadriaurita Retz., Observ. Bot. 6: 38. 1791. Pteris plumula Desv., P. nemoralis Willd. var. major M. Martens & Galeotti, P. galeotti (Fée) Hook., Litobrochia galeotti Fée Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, anama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Bolivia, Argentina, Africa, Asia. Salvadoran distribution: Ahuachapan: G. Davidse et a/. 37435 (B, BM, ITIC, LAGU, MO): R. Seiler 813, 908, 951 (MHES): W. Berendsohn et al. 1347 (B, LAGU, MO). Sonsonate: R. Seiler 621 (MHES). Santa Ana: K. Prestegard 2 (MP-00035) (LAGU); M.B. Siu s.n. (ITIC): R. Seiler 1205 (MHES): S. Winkler 36 (ITIC); T. Croat 42306 (ITIC). La Libertad: W. Berendsohn er al. 1048 (B, LAGU). San Salvador: R. Seiler 217 (MHES). Chalatenango: J.C. Gonzalez & M.A. Hernandez 268 (B, LAGU, MO). Cabaiias: A.K. Monro et al. 3743 (B, BM, ITIC, LAGU, MO); R.A. Carballo et al. 253 (LAGU, MO). Preris vittata L., Sp. Pl. 2: 1074. 1753. Local name: helecho de banda. Global distribution: Mexico, Honduras, Panama, Antilles, Venezuela, Peru, Brazil, Argentina; Viejo Mundo. Salvadoran distribution: Ahuachapan: K. Prestegard 6 (MP-00036) (LAGU); R. Seiler 811 (F, ITIC, MHES). La Libertad: E.A. Montalvo 6340 (B, LAGU, MO). Scoliosorus ensiformis (Hook.) T. Moore, Index Fil. xxix, 1857. Antrophyvum ensiforme Hook. in Bentham, 4. falcatum M. Martens & Galeotti, A. carnosum Liebm., A. galeotti Fée, Polvtaenium ensiforme (Hook.) Benedict — MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 202 Global ‘hatin Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panam iia distribution: Ahuachapan: M. Renderos et a/. 430 (LAGU); R. Seiler 915 (ITIC, MHES); 966 (MHES). Sonsonate: E.A. Montalvo s.n. (ITIC); R. Seiler 614 (MHES); T. Croat 42216 (ITIC). Santa Ana: B. Pfeiffer-Berendsohn & W. Berendsohn 30 (B, LAGU, MO); D. Rodriguez et a/. 37 (B, LAGU, NY); J. Lagos 1619 (ITIC); J. Monterrosa et al. 528 (B, LAGU, MO); 938 (B, LAGU, NY): K. Prestegard 25 (MP- 00009) (LAGU); R. Seiler 15, 131 (ITIC, MHES): 360 (MHES); S. Winkler 54 (ITIC): W. Berendsohn ef al. 1710 (B, BM, LAGU, MO). Chalatenango: R. Seiler 412 (JBL04946) (LAGU, MHES). Morazan: R. Seiler 1017 (ITIC, MHES). ¢ Vittaria bradeorum Rosenst., Repert. Spec. Nov. Regni Veg. 22: 18. 1925. Global distribution: Mexico, Guatemala, Costa Rica. ee Panama. Local names: helecho barba de cacique, helecho cola de gorrio Salvadoran distribution: Ahuachapan: E. Sandoval & M. siidisdd 1439 (B, LAGU); W. Berendsohn et a/. 1461 (LAGU). Vittaria graminifolia Kaulf., Enum. Filic. 192. 1824. Vittaria filifolia Fée, V. dimorpha Miill. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Suriname, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: R. Seiler 820, 960 (F, MHES). Sonsonate: A. Molina ef a/. 21722 (F, ITIC); R. Seiler 605 (F, MHES). Santa Ana: A. del Cid 1621 (ITIC); E.A. Montalvo 3281 (ITIC); F. Wunderlin 559 (ITIC); G. Davidse er al. 37281 (B, BM, ITIC, LAGU, MO); J.L. Linares s.n. (JBL01322) (LAGU); J. Monterrosa et al. 950, 991 (LAGU, NY); M.B. Siu s.n. (ITIC); R. Seiler 17 (ITIC, MHES); 128 (F, MHES); 99, 283, 356 (F, ITIC, MHES). San Salvador: M.C. Carlson 426 (F). Chalatenango: R. Seiler 402 (JBL00208) (F, LAGU, MHES): W. Heed 39 (ITIC). San Vicente: R. Seiler 335 (F, ITIC, MHES). Morazan: R. Seiler 991 (F, MHES). SALVINIACEAE > Azolla filiculoides Lam., Encycl. |: 343. 1783. Azolla caroliniana Willd. Global distribution: USA; Mexico, Guatemala, Honduras, Nicaragua, Colombia, Venezuela, Bolivia, Chile, Argentina, Uruguay, E Asia, Australia, Hawaii. References: Maxon & Standley (1930): 177 (as Azolla caroliniana), Calderon & Standley (1941): 35 (as A. caroliniana); Seiler (1982): 390; Berendsohn (1989): 23. Azolla microphylla Kaulf., Enum. Filic. 273. 1824. Azolla mexicana C. Pres Global distribution: Canada; Mexico, Honduras, Nicaragua, Antilles, Colombia, Venezuela, French Guyana, Ecuador, Peru, Brazil, Bolivia, Argentina, Uruguay. Salvadoran distribution: Ahuachapan: R. Seiler 722 (MHES). La Libertad: E.A.Montalvo 3472 (ITIC). San Vicente: N.C. Fassett 28344 (JBL01484) (ITIC, LAGU). San Miguel: J. Monterrosa & R. Villacorta 1638 (BM, LAGU, MO). 203 FERN GAZ. 18(4): 120-215. 2008 Salvinia auriculata Aubl., Hist. Pl. Guiane 2: 969, t. 367. 1775. Salvinia rotundifolia Willd. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Chile, Paraguay, Argentina, Uruguay. Salvadoran distribution: La Uni6n: R. Villacorta et a/. 2706 (B, CR, LAGU, MEXU); N.C. Fassett 28631 (ITIC, MO) (Moran in Davidse et al. 1995); 28662 (ITIC); J. Monterrosa & R. Villacorta 1637 (BM, LAGU, MO). Salvinia minima Baker, J. Bot. 24: 98. 1886. Salvinia rotundifolia auct. non Willd. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. Salvadoran distribution: Ahuachapan: G. Davidse et al. 37384 (B, BM, ITIC, LAGU, MO); J.M. Rosales 2272 (B, LAGU, MO); N. Herrera s.n. (JBLO3862) (LAGU, MO); R. Seiler 900 (MHES). Santa Ana: C. Pefiate 4 (ITIC). San Salvador: K. Sidwell ef al. 534 (B, BM, ITIC, LAGU, MO). La Paz: N.C. Fassett 28515 (ITIC). Usulutan: N.C. Fassett 28683 (ITIC). San Miguel: N.C. Fassett 28556 (ITIC); R. Seiler 723 (MHES). La Union: K. Sidwell ef a/. 689 (B, BM, ITIC, LAGU, MO). TECTARIACEAE Tectaria heracleifolia (Willd.) Underw., Bull. Torrey Bot. Club 33: 200. 1906. Aspidium heracleifolium Willd., Polypodium cordifolium M. Martens & Galeotti, Tectaria heracleifolia (Willd.) Underw. var. maxima C.V. Morton Local names: helecho cisne, helecho cola de perro, helecho corazon, helecho palma de costa. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 84 (B, LAGU); 732 (B, LAGU, MO); J.M. Rosales 231, 745 (AAU, B, BM, LAGU, MO); 946 (LAGU); R. Seiler 941 (JBL04819) (F, LAGU, MHES); R. Villacorta & Martinez 409 (LAGU, MO); W. Berendsohn & R. Villacorta 1110 (B, LAGU); W. Berendsohn ef al. 1319, 1324, 1361, 1605 (B, LAGU, MO). Sonsonate: D. Rodriguez & G. Trejo 200 (B, LAGU, MO). Santa Ana: K. Prestegard 35 (MP-00024) (LAGU). La Libertad: L.O. Williams et al. 16779 (F); R.A. Carballo et a/. 38 (B, BM, LAGU, MO). San Vicente: P. Standley ef al. 3532 (F). San Miguel: A. Salgado s.n. (JBL04802) (LAGU); A. Salgado 77 (F, MHES). Cabaiias: A.K. Monro er al. 3745 (B, BM, ITIC, LAGU, MO). Tectaria incisa Cav. f. incisa, Descr. Pl. 249. 1801. Aspidium macrophyllum Sw. var. pittieri Christ, A. martinicense Spreng., Tectaria martinicensis (Spreng.) Copel. Local name: helecho samaritana. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilts 212 (LAGU); MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 204 1352 (B, LAGU); R. Seiler 930 (F, MHES). Sonsonate: D. Rodriguez 161 (LAGU). La Libertad: S. Winkler 70 (ITIC). San Salvador: S. Winkler 89 (ITIC). San Vicente: P. Standley et al. 3771 (F). Tectaria mexicana (Fée) Morton, Amer. Fern. J. 56. 133. 1966. Sagenia mexicana Fée, Aspidium latifolium C. Presl, Tectaria latifolia (G. Forst.) Copel., TZ. mexicana (Fée) C.V. Morton var. pilosula C.V. Morton., 7. dilacerata auct. non Kunze Local names: helecho cola de pavo, helecho falsa salamandra, helecho pié de leon. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 2002 (B, BM, ITIC, LAGU, MO); E. Sandoval & H. Rivera 1313 (B, LAGU): F. Chinchilla & E. Sandoval s.n. (ISB00276) (B, LAGU, MO); s.n. (ISB00670) (AAU, B, LAGU); J.M. Rosales 1314 (LAGU); M. Sandoval & E. Sandoval s.n. (ISB00870) (AAU, B, F, LAGU); M. Sandoval & E. Sandoval 93 (B, F, LAGU); P. Standley et a/. 2607 (F); R. Seiler 816 (F, MHES); 934 (MHES). Sonsonate: D. Rodriguez & M. Trejo 82 (LAGU); J. Tucker 1351 (F). La Libertad: A.K. Monro ef al. 2306, 3090 (B, BM, ITIC, LAGU, MO); J. Monterrosa et al. s.n. (JBL04486) (B, BM, ITIC, LAGU, MO); M.C. Carlson 141 (F); M. Magafia 38 (MP-00038), 39 (MP-00014) (LAGU); N. Ventura 5003 (ITIC); O.Gonzalez et al. s.n. (JBL04495) (B, BM, ITIC, LAGU, MO). San Salvador: A.K. Monro ef al. 3438 (B, BM, ITIC, LAGU, MO); A. Molina ef a/. 21848 (F, ITIC); K. Sidwell et al. 573 (B, BM, ITIC, LAGU, MO); R. Seiler 197 (F, MHES). Chalatenango: L. Gonzalez 1721 (ITIC). San Vicente: P. Standley ef a/. 3772 (F). Usulutan: D. Williams 132 (LAGU, MO). San Miguel: A.K. Monro er al. 2947 (B, BM, ITIC, LAGU, MO); A. Salgado 74 (MHES); M. Renderos ef al. 685 (AAU, B, LAGU, MO); R. Villacorta et al. 2824 (B, BM, LAGU, MEXU, MO). Cuscatlan: S. Dar s.n. (JBL03664) (B, LAGU, MO). Tectaria rivalis (Mett. ex Kuhn) C. Chr., Index Filic., Suppl. 3, 184. 1934. Tectaria rheosora auct. non. (Baker) C. Chr. Global distribution: Belize, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador Saktadoxan distribution: Ahuachapan: R. Seiler 943 (F, MHES). > Tectaria transiens (C. V. Morton) A. R. Sm., Amer. Fern. J. 70: 27. 1980. Tectaria incisa Cav. subsp. transiens C.V. Morton, T. ramkissoonii Jermy & T. G Walker Global distribution: Mexico, Guatemala, Honduras, i Costa Rica, Panama, Antilles, Colombia, Trinidad and Tobago, Ecuador, P Source: R.C. Moran in Davidse et al. 1995: 209. eee & Smith 2004: 614. THELYPTERIDACEAE Macrothelypteris torresiana (Gaudich.) Ching, Acta Phytotax. Sin. 8: 310. 1963. Polystichum torresianum Gaudich., Thelypteris torresiana (Gaudich.) Alston, T. uliginosa (Kunze) Ching, Aspidium uliginosum Kunze, Drvopteris uliginosa (Kunze) 205 FERN GAZ. 18(4): 120-215. 2008 Local names: helecho amarillo, helecho palma de pozo, helecho pata de paloma. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Africa, Asia, Pacific Islands. Salvadoran distribution: Anuachapan: A.K. Monro et a/. 1930 (B, BM, ITIC, LAGU, MO); E. Sandoval & H. Rivera 1276 (B, LAGU); E. Sandoval & O. Rivera s.n. (ISB00814) (AAU, LAGU); F. Chinchilla & E. Sandoval s.n. (ISB00270) (LAGU): J.M. Rosales 1052 (B, LAGU); M. Sandoval & E. Sandoval s.n. (ISB00843) (AAU, B, LAGU); s.n. (JBL01367), 305 (B, LAGU); 208 (B, F, LAGU); S. Castillo s.n. (ISF00688) (LAGU); S. de Moore s.n. (12-933) (MHES). Santa Ana: G. Davidse et al. 37227 (B, BM, ITIC, LAGU, MO); K. Prestegard 20 (MP-00049) (LAGU); R. Seiler 470 (F, ITIC, MHES); 839 (MHES). La Libertad: A.K. Monro e¢ al. 2309, 3733, 3775 (B, BM, ITIC, LAGU, MO); W. Berendsohn et al. 1047 (B, LAGU); 1444 (LAGU). San Salvador: M. Renderos 707 (B, LAGU, MO); R. Seiler 369 (F, MHES); 1504 (F). Cabaiias: A.K. Monro et a/. 3753 (B, BM, ITIC, LAGU, MO). Thelypteris subg. Amauropelta Thelypteris balbisii (Spreng.) Ching, Bull. Fan Mem. Inst. Biol. 10: 250. 1941. Polypodium balbisii Spreng., Aspidium sprengelii Kaulf., Dryopteris sprengelii (Kaulf.) Kuntze, D. balbisii (Spreng.) Urb., Thelypteris sprengelii (Kaulf.) Proctor Local name: helecho palma de rio. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1931 (B, BM, ITIC, LAGU, MO); E. Sandoval & F. Chinchilla 918 (B, LAGU); M. Sandoval & E. Sandoval 303 (B, F, LAGU); P. Standley et a/. 2499 (F): R. Seiler 918, 935 (F, MHES); S. de Moore s.n. (12-932) (MHES); W. Berendsohn et a/. 1346 (B, LAGU, MO); 1722 (B, LAGU). Sonsonate: J. Tucker 1331 (F). Santa Ana: G. Cerén & D. Rodriguez s.n. (JBL04658) (B, LAGU, MO): R. Seiler 1203 (F, MHES). La Libertad: E.A. Montalvo & R. Villacorta 6402 (B, ITIC, LAGU, MO); L. Williams & A. Molina 15203 (JBL01550) (LAGU). San Salvador: A. Molina et al. 21841 (F, ITIC); R. Seiler 221, 712 (F, MHES). Chalatenango: L. Lara 235, 236 (MHES). Cuscatlan: E.A. Montalvo 3313 (ITIC). Morazan: R. Seiler 1084 (F, MHES). Thelypteris cheilanthoides (Kunze) Proctor var. cheilanthoides, Bull. Inst. Jamaica, Sci. Sef. 5; SE. 1953 Phegopteris amplificata Fée, Drvopteris amplificata (Fée) C. Chr., Aspidium kunzei Fee Global distribution: Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Santa Ana: V.M. Martinez s.n. (CMC00980) (B, EAP, LAGU, MO) Thelypteris concinna (Willd.) Ching, Bull. Fan Mem. Inst. Biol. 10: 251. 1941. Polypodium concinnum Willd., Dryopteris concinna (Willd.) Kuntze, Amauropelta concinna (Willd.) Pic. Serm., Phegopteris elongata E. Fourn., Thelypteris concinna (Willd.) Ching var. e/ongata (E. Fourn.) C. F. Reed MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 206 Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: S. Calderén 1994, LAGU (photo), NY. Chalatenango: R. Seiler 256 (F, MHES). Morazan: J. Tucker 701 (F). Thelypteris deflexa (C. Presl) R. M. Tryon, Rhodora 69: 5. 1967. Nephrodium defelxum C. Presl, Drvopteris lindigii C. Chr., Thelypteris lindigii (C. Chr.) Alston, Amauropelta deflexa (C. Presl) A. Live & D. Léve Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Morazan: R. Seiler 1010 (F, LAGU (photo), MHES, NY, ¢ Thelypteris linkiana (C. Presl) R.M. Tryon, Rhodora 69: 6. 1967. Gymnogramma diplazioides Desv., G. polvpodioides Link, G. cheilosora Fée, Grammitis linkiana C. Presl, Dryopteris linkiana (C. Presl) Maxon, D. diplazioides (Desv.) Proctor Global distribution: Mexico, Guatemala, Belice, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: J.Monterrosa, D.Rodriguez & R.Gonzalez 1528 (BM, LAGU, MHES, UC). Thelypteris melanochlaena (C. Chr.) C. F. Reed, Phytologia 17: 292. 1968. Drvopteris melanochlaena C. Chr Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Ahuachapan: R. Seiler 826 (F, MHES). Santa Ana: G Davidse et al. 37234 (B, BM, ITIC, LAGU, MO). Thelypteris oaxacana A. R. Sm., Amer. Fern J. 63: end Fy es PP. Global distribution: Mexico, Costa Rica, Nicaragua, Pana Salvadoran distribution: Chalatenango: Tucker 1043 F MICH) (A.R. Smith in Davidse ef al. 1995). Thelypteris oligocarpa (Humb. & Bonpl. ex Willd.) Ching, Bull. Fan Mem. Inst. Biol. 10: 253. 1941 Polypodium oligocarpum Humb. & Bonpl. ex Willd., Drvopteris oligocarpa (Humb. & Bonpl. ex Willd.) Kuntze, D. columbiana C. Chr., Amauropelta oligocarpa (Humb. & Bonpl. ex Willd.) Pic. Serm., Thelypteris columbiana (C. Chr.) C. V. Morton, Phegopteris pilosula Fée Local names: helecho trensillo. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Pérez 1242 (B, LAGU). Sonsonate: R. Seiler 600 (MHES). Santa Ana: J. Monterrosa ef al. 935, 969 (B, LAGU, NY); R. Seiler 117, 279, 281, 347 (F, MHES); 671 (MHES); 873 (F, ITIC, MHES). San Salvador: A.K. Monro et a/. 3455 (B, BM, ITIC, LAGU, MO); M.C. Carlson 4b (F); R. Seiler 33, 362 (F, MHES). Chalatenango: R. Seiler 421 (F, MHES). Morazan: R. Seiler 1059 (F, MHES). 207 FERN GAZ. 18(4): 120-215. 2008 Thelypteris pilosohispida (Hook.) Alston, J. Wash. Acad. Sci. 48: 233. 1958. Nephrodium pilosohispidum Hook., Dryopteris pilosohispida (Hook.) C. Chr., Alsophila pilosa M. Martens & Galeotti Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Antilles, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: A. Molina et al. 16745 (F); J. Monterrosa ef al. 207 (B, LAGU); R. Seiler 248 (F, MHES); S. Winkler 66 (ITIC). Morazan: R. Seiler 1008 (F, MHES). Thelypteris pilosula (Klotzsch & H. Karst. ex Mett.) R. M. Tryon, Rhodora 69: 7. 1967. Aspidium pilosulum Mett., Dryopteris pilosula (Klotzsch & H. Karst. ex Mett.) Hieron., Amauropelta pilosula (Mett.) A. Love & D. Léve Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: G. Davidse ef al. 37310 (B, BM, ITIC, LAGU, MO): R. Seiler 1193 (F, ITIC, MHES). Chalatenango: R. Seiler 421 (JBL00207) (LAGU); R. Seiler 728 (MHES). Thelypteris resinifera (Desv.) Proctor var. resinifera, Bull. Inst. Jamaica, Sci. Ser. 5: 63. 1953 Nephrodium panamense C. Presl, Dryopteris panamensis (C. Presl) C. Chr., D. panamensis (C. Presl) C. Chr. var. gonzalezii Christ ex C. Chr., Thelypteris panamensis (C. Presl) E. P. St. John, Polypodium litiginosum Liebm., Aspidium exsudans E. Fourn., A. exsudans E. Fourn. var. majus E. Fourn., A. exsudans E. Fourn. var. feei E. Fourn., A. ghiesbreghtii E. Fourn. Local name: helecho palma de rio. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: A.K. Monro & M.L. Fomtg 2976 (B, BM, ITIC, LAGU, MO); E. Sandoval & E. Fuentes 1302 (B, LAGU); F. Chinchilla & E. Sandoval s.n. (ISB00152) (B, LAGU); J.M. Rosales 431 (B, BM, LAGU, MO); 943 (B, LAGU, MO); P. Standley er al. 2969, 2982 (F); S. Castillo s.n. (ISF00593) (B, LAGU, MO). Santa Ana: B. Pfeiffer-Berendsohn 64 (LAGU, MO); E.A. Montalvo 3367 (ITIC). La Libertad: A.K. Monro er al. 2291 (B, BM, ITIC, LAGU, MO); E.A. Montalvo 3417 (ITIC). San Salvador: A.K. Monro et ail. 2239 (B, BM, ITIC, LAGU, MO); E.A. Montalvo 3585 (ITIC); R. Seiler 784 (F, ITIC, MHES). Chalatenango: A. Molina et a/. 21556 (F, ITIC); R. Seiler 261 (F, MHES); R. Villacorta et al. 982 (B, LAGU, MO). San Vicente: P. Standley et al. 3409, 3697 (F). Morazan: A.K. Monro ef al. 3803 (B, BM, ITIC, LAGU, MO); J. Tucker 645, 700 (F); K. Sidwell et al. 750 (B, BM, ITIC, LAGU, MO); R. Seiler 1049a (F, ITIC, MHES). Thelypteris rudis (Kunze) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 64. 1953. Polypodium rude Kunze, Dryopteris rudis (Kunze) C. Chr.. Amauropelta rudis (Kunze) Pic. Serm., Phegopteris impressa Fée Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Ecuador, Peru, Bolivia. Salvadoran distribution: Santa Ana: A. Molina et al. 16745 (LAGU (photo), NY): R. Seiler 194 (F). MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 208 Thelypteris scalaris (Christ) Alston, J. Wash. Acad. Sci. 48: 234. 1958. Aspidium scalare Christ, Drvopteris scalaris (Christ) C. Chr., Amauropelta scalaris (Christ) A. Léve & D. Léve Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Ecuador, ' Salvadoran distribution: Ahuachapan: A.K. Monro et a/. 2015 (B, BM, ITIC, LAGU, MO). Santa Ana: R. Seiler 193 (F, ITIC, LAGU (photo), MHES, NY). La Libertad: K. Sidwell et a/. 708 (B, BM, ITIC, LAGU, MO). Usulutan: A.K. Monro er a/. 3021 (B, BM, ITIC, LAGU, MO); D. Williams 59 (LAGU, MO). Morazan: R. Seiler 985 (MHES, UC). > Thelypteris thomsonii (Jenman) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 65. 1953. Polypodium thomsonii Jenman, Dryopteris thomsonii (Jenman) C. Chr., D. stuebelii ieron., Amauropelta thomsonii (Jenman) Pic. Serm Global distribution: Mexico, Guatemala, Costa Rica, Panama, Antilles, Ecuador, Peru. Source: Stolze (1981): 487; Berendsohn (1989) 1: 26; Mickel & Smith (2004): 637. Thelypteris subg. Cyclosorus Thelypteris cretacea A. R. Sm., Univ. Calif. Publ. Bot. 59: 92. 1971. Christella cretacea (A. R. Sm.) A. Love & D. Léve Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica. Salvadoran distribution: Ahuachapan: A.K. Monro & M.L. Fomtg 2983 (B, BM, ITIC, LAGU, MO); J.M. Rosales 945 (B, LAGU, MO); R. Seiler 919 (F, MHES). Santa Ana: M.B. Siu s.n. (ITIC). La Libertad: R. Seiler 715 (F, MHES). San Salvador: R. Seiler 254 (F, MHES). Cabaiias: R. Seiler 523 (F, MHES). Thelypteris dentata (Forssk.) E. P. St. John, Amer. Fern J. 26: 44. 1936. Polypodium dentatum Forssk., Drvopteris dentata (Forssk.) C. Chr., D. mollis (Sw.) Hieron., Cyclosorus dentatus (Forssk.) Ching, Christella dentata (Forssk.) Brownsey & Jermy, Aspidium molle Sw. Local name: palmarin. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Paraguay, Argentina, Africa, Asia, Pacific Islands. Salvadoran distribution: La Libertad: K. Prestegard 44 (MP-00046) (LAGU). San Salvador: R. Seiler 219 (MHES); 710 (F, MHES). Chalatenango: R. Seiler 259 (F, MHES). La Union: R. Seiler 1048 (F, MHES). Thelypteris hispidula (Decne.) C. F. Reed, Phytologia 17: 283. 1968. Aspidium hispidulum Decne., Christella hispidula (Decne.) Holttum, Nephrodium quadrangulare Fée, Dryopteris quadrangularis (Fée) Alston, Thelypteris quadrangularis (Fée) Schelpe Local names: helecho de rio, helecho palma de rio, helecho trensillo, palma de barranco. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama. Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, 209 FERN GAZ. 18(4): 120-215. 2008 Uruguay, Africa, Asia. Salvadoran distribution: Ahuachapan: E.A. Montalvo 5061 (ITIC); E. Sandoval 206 (LAGU); 1147 (B, LAGU, MO); F. Chinchilla & E. Sandoval s.n. (ISB00426) (LAGU); J.M. Rosales 43 (AAU, B, BM, LAGU, MO, MEXU); 987 (B, BM, LAGU, MO); 340, 2667 (B, LAGU, MO); 688, 2177 (AAU, B, BM, LAGU, MO); M. Sandoval & E. Sandoval 251 (B, F, LAGU); R. Seiler 942 (F, MHES); S. Castillo s.n. (ISF00582) (B, F, LAGU, MO). Santa Ana: K. Prestegard 4 (MP-00047) (LAGU); M. Magajfia 3 (LAGU); O. Pank s.n. (12-1000) (MHES). La Libertad: A.K. Monro ef al. 3092 (B, BM, ITIC, LAGU, MO); N. Ventura 5022, 5023 (ITIC); W. Berendsohn ef al. 1443 (LAGU). San Salvador: R. Seiler 50, 220, 453 (F, MHES). Usulutan: D. Williams & R.W. Herrera 237 (LAGU). San Miguel: A.K. Monro et al. 2945 (B, BM, ITIC, LAGU, MO). Cabaiias: G. Davidse et al. 37115 (B, BM, ITIC, LAGU, MO); R. Seiler 227, 229 (MHES). Thelypteris interrupta (Willd.) K. lwats., Jap. J. Bot. 38: 314. 1963. Pteris interrupta Willd., Cyclosorus interruptus (Willd.) H. It6, C. tottum (Thunb.) Pic. Ser., C. gongylodes (Schkuhr) Link, Dryopteris gongylodes (Schkuhr) Kuntze, Polypodium tottum Thunb., Thelypteris totta (Thunb.) Schelpe, T. gongylodes (Schkuhr) Small, Aspidium gongylodes Schkuhr, Nephrodium paludosum Liebm. Global distribution: USA, Mexico, Guatemala, Honduras, Costa Rica, Panama, Antilles, Colombia, Venezuela, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay, Africa, Asia. Salvadoran distribution: Ahuachapan: G. Davidse et al. 37454 (B, BM, ITIC, LAGU, MO); J.M. Rosales 2447 (B, LAGU, MO); N.C. Fassett 28523 (ITIC); N. Herrera s.n. (JBL03767) (B, LAGU, MO); R. Seiler 950 (F, MHES). Thelypteris kunthii (Desv.) C.V. Morton, Contr. U.S. Natl. Herb. 38: 53. 1967. Nephrodium kunthii Desv., Dryopteris normalis C. Chr., Thelypteris normalis (C. Chr.) Moxley, Christella normalis (C. Chr.) Holttum. Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Antilles, Colombia, Venezuela, Brazil. Salvadoran distribution: San Miguel: R. Villacorta ef al. 2703 (B, BM, LAGU, MO); 2890 (B, BM, EAP, LAGU, MO). Thelypteris patens (Sw.) Small var. patens, Ferns S.E. States 243. 1938. Polypodium patens Sw., Dryopteris patens (Sw.) Kuntze, Christella patens (Sw.) Holttum Local name: helecho palma de pozo. Global distribution: USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Peru, Brazil. Bolivia, Paraguay. Salvadoran distribution: Anuachapan: E. Sandoval & F. Chinchilla 81 (B, LAGU); R. Seiler 936 (F, ITIC, MHES). Sonsonate: D. Rodriguez 160 (LAGU); J. Tucker [352 (F); M.B. Siu s.n. (ITIC). Santa Ana: E.A. Montalvo 3246 (ITIC); S. Winkler 87 (ITIC). La Libertad: A.K. Monro et al. 3731, 3734 (B, BM, ITIC, LAGU, MO); R. Villacorta 1068 (B, LAGU, MO). San Salvador: A. Molina et al. 21846 (F); R. Seiler 340, 519 (F, MHES). San Vicente: P. Standley ef a/. 3675 (F). Usulutan: D. Williams 261 (LAGU). Cabaiiias: A.K. Monro et al. 3744, 3756 (B, BM, ITIC, LAGU, MO); R. lied tna statin timate peli RA taal iccia citer atapagena = oly cham emai, aginst MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 210 Seiler 226 (F, MHES). Thelypteris patens (Sw.) Small var. smithiana Ponce, Darwiniana 28: 373. 1987. Global distribution: Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela. Ecuador, Peru, Bolivia, Brazil. Salvadoran distribution: Santa Ana: R. Seiler 762 (F, MHES). San Miguel: A. Salgado 73 (F, MHES). Cabafias: G. Davidse er al. 37118 (B, BM, ITIC, LAGU, MO). Morazan: Tucker 690 (F, UC) (A.R. Smith in Davidse er a/. 1995). Thelypteris puberula (Baker) C.V. Morton var. puberula, Amer. Fern J. 48: 138. 1958. Aspidium puberulum Fée, Nephrodium puberulum Baker in Hooker & Baker, Dryopteris puberula (Baker) Kuntze, D. feei C. Chr., D. augescens (Link) C. Chr. var. puberula (Baker) C. Chr., Thelypteris feei (C. Chr.) Moxley, Christella puberula (Baker) A. Live & D. Live Local name: helecho peloso. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Antilles. Salvadoran distribution: Santa Ana: J. Lagos 1622 (ITIC); R. Seiler 676 (F); V.M. Martinez s.n. (CMC00854) (B, LAGU); R. Seiler 876 (MHES). San Salvador: A. Molina & E.A. Montalvo 21653 (EAP, LAGU (photo), NY); R. Seiler 51 (F, MHES). La Libertad: J. Williams s.n. (ITIC); L.O. Williams er a/. 15092 (F); R. Seiler 1520 (F). Thelypteris subg. Goniopteris Thelypteris hatchii A. R. Sm., Amer. Fern J. 63: 118 (F). 1-3. 1973. Goniopteris hatchii (A. R. Sm.) A. Love & D. Love Global distribution: Mexico, Guatemala, Honduras, Costa Rica. Salvadoran distribution: Morazan: R. Seiler 1063 (F, ITIC, MHES). Thelypteris minor (C. Chr.) A. R. Sm., Phytologia 34: 232. 1976. Dryopteris nicaraguensis (E. Fourn.) C. Chr. var. minor C. Chr. Local name: helecho nido de guachota. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Anuachapan: A.K. Monro et al. 1977 (B, BM, ITIC, LAGU, MO); D. Witsberger 653, 690, 691 (MHES); E. Sandoval 42 (LAGU, MO); 1136 (B, LAGU); G. Davidse er al. 37373, 37376 (B, BM, ITIC, LAGU, MO); N. Herrera s.n. (JBL03861) (LAGU, MO); R. Seiler 815 (F, ITIC, MHES). Sonsonate: A. Molina et al. 21624 (ITIC). Santa Ana: M.B. Siu s.n. (ITIC): S. Winkler 88 (ITIC). Morazan: R. Seiler 905 (F, ITIC, MHES). ¢ Thelypteris obliterata (Sw.) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 62. 1953. Polypodium obliteratum Sw.. P. faucium Liebm., Goniopteris obliterata (Sw.) C. Presl., Dryopteris obliterata (Sw.) C. Chr. Global distribution: Mexico, Guatemala, Belice, Honduras, Nicaragua, Costa Rica, Antilles. Salvadoran distribution. Morazan: J.Monterrosa & C.Bermudez 1383 (BM, LAGU, MO, NY, UC) , . 211 FERN GAZ. 18(4): 120-215. 2008 Thelypteris tetragona (Sw.) Small, Ferns S. E. States 256. 1938. Polypodium tetragonum Sw., Goniopteris tetragona (Sw.) C. Presl, Dryopteris tetragona (Sw.) Urb., D. subtetragona (Link) Maxon Global distribution. USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: P. Standley 21647, GH (A.R. Smith in Davidse et al. 1995). Ahuachapan: R. Seiler 953 (MHES). Santa Ana: K. Prestegard 11 (MP-00050) (LAGU). Thelypteris subg. Meniscium ¢ Thelypteris angustifolia (Willd.) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 57. 1953. Meniscium angustifolium Willd., Dryopteris angustifolia (Willd.) Urb., Polypodium salicifolium Vahl obal distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay. Salvadoran distribution: Morazan: J.Monterrosa & C.Bermudez 1381 (BM, LAGU, UC). @ Thelypteris serrata (Cav.) Alston, Kew Bull. 1932: 309. 1932. Meniscium serratum Cav., Drvopteris serrata (Cav.) C. Chr. Local name: helecho gallardete. Global distribution: USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: F. Chinchilla & E. Sandoval s.n. (ISB00277) (LAGU, MO). Sonsonate: S. Winkler 34 (ITIC). @ Thelypteris standleyi (Maxon & C. V. Morton) R. M. Tryon, Rhodora 69: 8. 1967. Dryopteris standleyi Maxon & C. V. Morton Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia. Salvadoran distribution: Morazan: A.K. Monro et al. 3828 (B, BM, ITIC, LAGU, MO); J. Monterrosa & R. Carballo 795, 871 (LAGU). WOODSIACEAE Athyrium bourgeaui E. Fourn., Mexic. Pl. 1: 102.1872. Athyrium paucifrons C. Chr. Global distribution: Mexico, Guatemala. Salvadoran distribution: Morazan: R. Seiler 1056 (ITIC, MHES). Athyrium palmense (Christ) Lellinger, Proc. Biol. Soc. Wash. 89: 708. 1977. Cystopteris fragilis (L.) Bernh. var. palmensis Christ Global distribution: Mexico, Costa Rica. Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar 760 (MHES); R. Seiler 620 (MHES); 1202 (ITIC). La Libertad: W. Berendsohn et al. 1442 (LAGU). San MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 212 Salvador: R. Seiler 450, 580 (MHES). Athyrium skinneri (Baker) Diels, Nat. Pflanzenfam. I(4): 224.1899. Asplenium skinneri Baker Global distribution: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 527 (B, LAGU); J.M. Rosales 944, 1053 (AAU, B, BM, LAGU, MO). Sonsonate: D. Rodriguez & G. Trejo 107 (LAGU, NY). Santa Ana: K. Prestegard 9 (MP-00013) (LAGU); R. Seiler 701 (MHES). La Libertad: N. Ventura 5002 (ITIC); O. Rohweder 2625 (JBL01253) (LAGU); 2626 (JBL01252) (LAGU). San Salvador: R. Seiler 583 (JBL04814) (F, LAGU, MHES),; R. Seiler 68, 375, 510, 635 (F, MHES Cystopteris fragilis (L.) Bernh., Neues J. Bot. I(2): 26, t. 2 (F). 9. 1806. Polypodium fragile L., P. diaphanum Bory, Cystopteris fragilis (L.) Bernh. subsp. diaphana (Bory) Litard., C. diaphana (Bory) Blasdell, C. arcuata Fée, Aspidium fragile (L.) Sw. var. fumarioides M. Martens & Galeotti Global distribution. USA, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Chile, Argentina, Europe, Africa, Asia, Hawaii. Salvadoran distribution: Santa Ana: G. Davidse et al. 37250 (B, BM, ITIC, LAGU, MO); J. Monterrosa et al. 982 (B, LAGU, MO, NY): R. Seiler 353 (JBL04809) (F, LAGU, MHES); R. Seiler 278, 343, 502 (F, MHES). Chalatenango: R. Seiler 155 (F, MHES); R. Seiler 405 (JBL00090) (F, LAGU, MHES). Diplazium cristatum (Desr.) Alston, J. Bot. 74: 173.1936. Meniscium cristatum Desr., Asplenium arboreum Willd., A. denticulosum Desv., A. shepherdii Spreng., Diplazium arboreum (Willd.) C. Presl, D. denticulosum (Desv.) C. Chr. Global distribution. Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Guayana, Suriname, French Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina. Salvadoran distribution: Ahuachapan: J.M. Rosales 922, 947 (AAU, B, BM, LAGU, MO); M. Magajia 40 (MP-00015) (LAGU); R. Seiler 916 (F, MHES). Santa Ana: M. B. Siu s.n. (ITIC). San Salvador: R. Seiler 371 (F, ITIC, MHES); 532 (F, MHES). Diplazium donnell-smithii Christ, Bull. Herb. Boissier, sér. 2, 7: 270. 1907. Global distribution: Mexico, Guatemala, Honduras, iene, Costa Rica. Salvadoran distribution: Ahuachapan: A.K. Monro et al. 1980 (B, BM, ITIC, LAGU, MO); R. Seiler 825 (F, ITIC, MHES); S. de Moore s.n. (12-912) (MHES). Santa Ana: R. Seiler 358 (JBL04810) (LAGU, MHES); 689 (JBL04816) (F, LAGU, MHES); 273 (MHES); 474 (F, MHES); V.M. Martinez s.n. (CMC00896) (B, EAP, LAGU, MO). San Salvador: A. Molina & E.A. Montalvo 21852 (JBL01464), EAP (F, LAGU); R. Seiler 336 (F, MHES); S. Winkler 32 (ITIC). @ Diplazium drepanolobium A.R. Sm., Amer. Fern. J. 70: 21. 1980. Global distribution: Mexico, Guatemala. Salvadoran distribution: Sonsonate: R. Villacorta et al. 323 (B, F, LAGU, MO). Diplazium franconis Liebm., Mexic. Bregn. 256 (reprint 104). 1849. 213 FERN GAZ. 18(4): 120-215. 2008 Asplenium franconis (Liebm.) Mett., A. franconis (Liebm.) Mett. var. gracile E. Fourn., A. aspidiiforme Fée, A. distans Fée, A. shepherdii Spreng. var. bipinnatum Christ, Diplazium camptocarpon Fée . Local names: helecho pact, helecho crisalia. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Panama. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 334 (LAGU, MO); E. Sandoval & F. Chinchilla 43 (LAGU). Santa Ana: A. Molina ef al. 16896 (F); G. Cerén & D. Rodriguez s.n. (JBL04662) (LAGU); G. Davidse er al. 37225 (B, BM, ITIC, LAGU, MO); R. Seiler 178, F (LAGU). Chalatenango: R. Seiler 380 (F, MHES). Diplazium lindbergii (Mett.) Christ, Prim. Fl. Costaric. 3(1): 27. 1901. Asplenium lindbergii Mett., A. induratum Christ, Diplazium induratum Diels Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Ecuador, Peru, Brazil, Bolivia, Uruguay. Salvadoran distribution: Létschert 414, US (Mickel & Smith, 2004: 260). Morazan: R. Seiler 981 (MHES); 1028 (ITIC). Diplazium lonchophyllum Kunze, Linnea 13: 141. 1839. Diplazium inaequilaterum Liebm., D. actuale Fée . Local names: helecho de coqueta, helecho palma de angel, helecho pluma de tucan, helecho trencilla de bajio. Global distribution: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 130 (LAGU); E. Sandoval & F. Chinchilla 487, 1165 (B, LAGU); E. Sandoval & M. Sandoval 1372 (B, LAGU); F. Chinchilla & E. Sandoval s.n. (ISB00151) (B, LAGU, MO); J.M. Rosales 451 (AAU, B, BM, LAGU, MO): M. Sandoval & E. Sandoval s.n. (ISB00869), 289, 294 (B, F, LAGU); P. Standley et al. 2567 (F); R. Seiler 917, 931 (F, MHES): W. Berendsohn 1312 (B, LAGU, MO); W. Berendsohn ef a/. 1400 (B, LAGU, MO). Sonsonate: J. Tucker 1350 (F); R. Seiler 596 (F, MHES). Santa Ana: K. Prestegard 5 (MP-00019) (LAGU); R. Seiler 244 (F, ITIC, MHES): V.M. Martinez s.n. (CMC00853) (B, LAGU, MO); V.M. Martinez s.n. (CMC00873), s.n. (CMC00967), s.n. (CMC01010), s.n. (CMC01032) (B, EAP, LAGU, MO). La Libertad: J. Mauricio et al. s.n. (JBL04480), s.n. (JBL04490) (B, BM, ITIC, LAGU, MO); R. Seiler 714 (F, MHES). San Salvador: R. Seiler 218, 370, 451, 452, 533, 786 (F, MHES). Chalatenango: L. Lara 232, 236a (MHES). Cabafias: A.K. Monro et a/. 3749 (B, BM, ITIC, LAGU, MO). Cuseatlan: Grupo Ecolégico Canton La Bermuda s.n. (JBL03211) (AAU, B, BM, LAGU, MO). Morazan: R. Seiler 1064, 1086 (F, MHES). Diplazium striatum (L.) C. Presl, Suppl. Tent. Pterid. 114. 1836. Asplenium striatum L., Diplazium crenulatum Liebm., D. elongatum Fée, D. matudae C. D. Adams. Local names: helecho macho de altura, helecho negro de altura, helecho palma de calvario. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Antilles, Colombia, Venezuela, Trinidad and Tobago, Ecuador, Peru, Brazil, Bolivia. Salvadoran distribution: Ahuachapan: E. Sandoval & F. Chinchilla 211 (LAGU); E. Sandoval & M. Sandoval s.n. (JBL0O1385) (B, LAGU); F. Chinchilla s.n. (ISB00572) (AAU, F, LAGU), s.n. (ISB00662) (AAU, F, LAGU); R. Seiler 933 (F, MHES). ——e ee eee ee ee ee MONTERROSA & MONRO: CHECKLIST OF THE FERNS OF EL SALVADOR 214 Sonsonate: J. Tucker 1353 (F). San Salvador: S. Winkler 44 (ITIC). Morazan: R. Seiler 981 (F). Diplazium ternatum Liebm., Mexic. Bregn. 252 (reprint 100). 1849. Asplenium ternatum (Liebm.) Hook. Global distribution: Mexico, Guatemala, Honduras, Nicaragua. Salvadoran distribution: Santa Ana: R. Seiler 772 (ITIC); Morazan: J.L. Linares & A. Pool 2292 (JBL01931) (LAGU). Diplazium vera-pax (Donn. Sm.) Hieron., Hedwigia 59: 322. 1917. Asplenium vera-pax Donn. Sm., A. cordovense Baker Global distribution: Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama. Salvadoran distribution: Santa Ana: M.L. Reyna de Aguilar 659 (MHES). Diplazium werckleanum Christ, Bull. Herb. Boissier, sér. 2, 4: 969. 1904. Global distribution: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia. Salvadoran distribution. Ahuachapan: R. Seiler 902 (F, ITIC, MHES); 952 (F, MHES). Sonsonate: A. Molina & E.A. Montalvo 21625 (JBL01491) (F, EAP, LAGU), 21774 (JBL01555) (F, EAP, LAGU). Santa Ana: B. Pfeiffer-Berendsohn 50 (B, LAGU, MO); D. Rodriguez & G, Cerén s.n. (JBL04596) (LAGU); J. Tucker 1214 (F); J. Monterrosa & R. Martinez 939, 946 (LAGU, NY); K. Prestegard 31 (MP-00020) (LAGU); K. Sidwell et al. 844 (B, BM, ITIC, LAGU, MO); O.Pank s.n. (12-979) (MHES); R. Seiler 275, 483 (F, MHES): S. Winkler 1b (ITIC). San Salvador: R. Seiler 355 (F, MHES). Chalatenango: R. Seiler 379 (JBL00089) (F, LAGU, MHES). Morazan: R. Seiler 1025 (F). Woodsia mollis (Kaulf.) J. Sm., J. Bot. (Hooker) 4: 191. 1841. Physematium molle Kaulf., Woodsia guatemalensis Hook., W. fragilis Liebm. Global distribution: Mexico, Guatemala, Nicaragua. Salvadoran distribution: Sonsonate: R. Seiler 496 (F, MHES). Santa Ana: M.B. Siu s.n. (ITIC); R. Seiler 43, 661 (F, MHES); 501 (ITIC, MHES); S. Winkler 83 (ITC). 215 FERN GAZ. 18(4). 2008 INSTRUCTIONS FOR AUTHORS PAPERS should not usually exceed 20 printed pages and are generally expected to be considerably shorter. Review articles, as well as reports of original research, are encouraged. Short notes are acceptable e.g. new records. The senior author should supply a fax and email address to facilitate correspondence. MANUSCRIPTS should be submitted in English (British) in electronic format (preferably) or hard copy (two copies), in 10-point Times New Roman font and double spaced. Electronic versions of text and tables should be compatible with WORD, with figures as pdf or jpg files, and sent as email attachments or CDroms. All manuscripts will be refereed THE TITLE should reflect the content of the paper and be in BOLD CAPITALS (11- point) and centrally aligned. Generic and specific names should be in italics and any title containing a generic or specific name must be followed by the family and Pteridophyta in brackets e.g. TRICHOMANES SPECIOSUM (HYMENOPHYLLACEAE: PTERIDOPHYTA) IN SOUTHERN SPAIN AUTHOR ABBREVIATIONS should follow Pichi Sermolli's (1996) Authors of scientific names in Pteridophyta, Royal Botanic Gardens, Ke MAIN HEADINGS: should be in BOLD CAPITALS (10- point) and centrally aligne SUBSIDIARY HEADINGS: should be in bold, the first letter of each word in capitals, the rest in lower case and left-aligne AUTHORS' NAMES AND FULL ADDRESSES: follow the title and are centrally aligned. KEY WORDS: up to ten. ABSTRACT: should reflect the content of the paper. FIGURES: there is no distinction between photographs and line drawings in numbering. All should be presented in a form ready for reproduction, ideally in JPG format (please contact editor with queries), with a scale bar where appropriate. Lettering or numbers (Arabic) should be in the bottom left using uppercase Times Roman and be sufficiently large to be legible if reduction is necessary during printing. The number of photographs allowed in any one issue is limited by cost. Figure captions should be on a separate sheet TABLES: can be printed in either portrait or landscape format. Authors should consider this when preparing tables. Authors should ensure that tables fit the printed page size in a legible form. MEASUREMENTS: should follow the metric system. CHECHEISES: should follow the format of Baksh-Comeau, Fern Gaz. 16(1, 2): 11- REFERENCES: should follow the style of a recent issue of The Fern Gazette, e.g.:- HOOKER, W.J. 1864. Species Filicum, 5. Dulau & Co., London. MORTON, C.V. 1947. The American species of Hymenophyllum, section Sphaeroconium. Contr. U.S. Natl. Herb. 29(3): 139-201. STEVENSON, D.W. & LOCONTE, H. 1996. Ordinal and familial relationships of pteridophyte genera. In: CAMUS, J.M., GIBBY, M. & JOHNS, R.J. (Eds) Pteridology in perspective, pp. 435-467. Royal Botanic Gardens, Kew. JOURNAL ABBREVIATIONS: should follow Botanico Periodicum Huntianum & Supplements. Alterations from the original text at proof stage will be charged for unless they are minor points of detail. Twenty-five offprints will be supplied free to the senior author. Missouri Botanical Garden Libra re saris prenmotoaicaL soc ||| ||'||| || Registered Charity N Patron: HRH The Aa a athe Officers and Committee from April 2008 i) President: R.W. Sykes, Ormandy House, — amg Cumbria LA8 8BP il: President@eBPS.org.uk Vice-Presidents: A.R. Busby, R.J. Aare A.F. Dyer, M.S. Porter General Secretary: Dr Y.C. Golding, 7 Grange Road, Buxton, Derbyshire SK17 6NH. E-mail: Secretary@eBPS. org.uk Committee Secretary: ogy: Ackers, Deersbrook, Horsham Road, Walliswood, Dorking RH5 SRL Treasurer: rs G.J. Smith, Rookwood, 1 Prospect Road, Oulton Broad, Lowestoft, Suffolk NR32 3PT; E-mail: Treasurer@eBPS.org.uk Membership Secretary: M.G. Taylor, Westlea, ae Isleof Skye IV418PH mail: Membership@eBPS.orguk Meetings Secretary: R.J. Cooke, 2 Baden Road, Ketton, Rutland PE93RJ- E-mail: Meetings@eBPS.orguk Conservation Officer: Dr H.S. McHaffie, 180 Granton Road, Edinburgh EHS 1AH E-mail: Conservation@eBPS. org.uk Conservation OF icer Dr EJ. Rumsey, Dept. of B TI Cromwell Road, & Recorder. London SW7 SBD; E-mail: Consenaon 2eBPS.owk RevrereBPS.oi ak Project Officer: AC. rer Kersey’s Farm, Mendlesham ket, Suffolk IPI45RB E-mail: Projects@eBPS. org.uk Publications Secretary | MLH. Rickard, Pear Tree Cottage, Kyre, Tenbury Wells, Worcs. WR1S8RN & Editor of Pteridologist: E-mail: Publications@eBPS. org.uk, Pteridologist@eBPS. org.uk Editor of the Bulletin: Miss A.M. Paul, Dept. of Botany, The The Natural History Museum, — Cromwell apes London Swi SBD; E-mail: Bulletin@eBPS.org.uk Editor of The Fern Gazette: Prof. M prea EH3 5LR; E-mail: FernGazette@eBPS. org. uk Editor of BPS Website: A.C, Pigott (address above); E-mail: Webmaster@eBPS.org.uk Elected Committee Members: _ Prof. J.A. .A. Edgington, R. Golding, A.E. Greening, Dr M. Hayward, ee Dr B.E. Laue, Dr ae S.D. Martinelli, H.W. Matthews, P-H. Ripley, B.D. Smith, M.J. Stribley Booksales Organiser: Dr F, Katzer, 13 Hawdene, — Bigear eae Horticultural Information Adviser & Archivist: E-mail: H g.uk, Archivist@eBPS.org.uk Merchandise Organisers: Mr B.D. & Mrs GJ. Smith, Rookw ganda gran er ge 2 ce 23 Lowestoft, Suffolk NR32 3PT; E-mail: Merchandise@eBPS.org.uk Plant Exchange Organiser: Mr J.P. Coome, Kellys Cramer, Cottage, Tredilion, Abergavenny, Gwent NP7 SBB | E-mail: PlantExchange@eBPS.org.uk Spore Reclanse Organiser: ALR. Busby, phate above), E-mail: Spores@eBPS.org.uk ae _ Trustees of Greenfield & Centenary Funds: President, General Secretary & Treasurer The BRITISH PTERIDOLOGICAL SOCIETY was founded in 1891 and is still a focus for us THE FERN GAZETTE VOLUME 18 PART 4 2008 CONTENTS MAIN ARTICLES An annotated checklist of the Monilophytes (ferns) and Lycophytes of El Salvador J. Monterrosa & A. K. Monro 120-214 INSTRUCTIONS FOR AUTHORS 215