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\PH OF THE PHEASANTS

FRONTISPIECE, PLATE

XXI.

WHITE-CRESTED KALEEGE.

BO eotsre

PLATE XXI

WHITE-CRESTED KALEEGE

Gennaeus albocristatus (Vigors)

Over the great spruce forests of Kashmir and Garhwa! chilly winds come rearing down from Tibetan
snow-peaks. But among the ferns and moss-muffled bases of the trees the air is still, and fragrant with the
odours of many forest flowers. Here the White-crested Kaleege live; here they scratch for grubs and tubers,
court their mates, make their lowly homes, and at twilight roost upon high, swaying branches. Before dawn
a few will succumb to the sudden attack of marten or weasel ; the others awaken in early morning, send forth

_ their challenge and begin anew their daily life.

MONOGRAPH OF THE
PHEASANTS

WILLIAM BEEBE

Curator of Birds of the New York Zoological Park; Fellow of the New York Zoological
Society and Director of the Tropical Research Station in British Guiana; Fellow
of the American Ornithologists’ Union and of the New York Academy
of Sciences; Member of the British Ornithologists’ Union ;
Corresponding Member of the Zoological
Society of London, etc.

IN FOUR VOLUMES

7 VOLUME II

PUBLISHED UNDER THE AUSPICES OF THE
NEW YORK ZOOLOGICAL SOCIETY BY

H. F. & G. WITHERBY,
326 HIGH HOLBORN, LONDON, ENGLAND
1921

FOL
6%
Cora ae
Pals X
14h
yd
RB

Pare pats:
NA

CONTENTS

PAGE

KALEEGE PHEASANTS ; : i 5 : : 5 : : 4 ; I
WHITE-CRESTED KALEEGE (Gennaeus albocristatus) P ‘ : 6 3 “ 10
NEPAL KALEEGE (Gennaeus leucomelanus) . é : : 4 . - 3 26
BLACK-BACKED KALEEGE (Gennaeus melanonotus) . 5 : 30
BLACK-BREASTED KALEEGE (Gennaeus horsfieldt) . ‘ : : : : : 43
LINEATED KALEEGE (Gennaeus lineatus) : ‘ A A ( : - 5 53
CHINESE SILVER KALEEGE (Gennaeus nycthemerus) : : ; . : : 63
HAINAN SILVER KALEEGE (Gennaeus whitehead?) . : : : ; : 5 72
EDWARD’S KALEEGE (Gennaeus edwardst) . Q : : : ; A : 76
SWINHOE’S KALEEGE (Gennaeus swinhoit) . ‘ : . é : : . 78
Witp KaLeEGE HysrRips . : : . : : 3 3 : - - 84.

CRESTLESS FIREBACK PHEASANTS . : : : : ‘ : Sh OO
MALAYAN CRESTLESS FIREBACK (Acomus erythrophthalmus) . . : : 102
BORNEAN CRESTLESS FIREBACK (Acomus pyronotus) . 6 : ; : : 109

CRESTED FIREBACK PHEASANTS . ‘ : : : ‘ : : ot BIE
SIAMESE CRESTED FIREBACK (Lophura diardi) . 5 : , : : spas 117
MALAYAN CRESTED FIREBACK (Lopfhura rufa). : : ; : ° . 122
BORNEAN CRESTED FIREBACK (Lophura ignita) . . : ; : “ 5 he SAS)

WHITE-TAILED WATTLED PHEASANT. : : - : : : =) 143
WHITE-TAILED WATTLED PHEASANT (Lodiophasis bulwert) . “ . 5 p> MKS

THE JUNGLEFOWL. ; : : : : : : : 3 ; ae LO,
RED JUNGLEFOWL (Gallus gallus) . ; é : : : : : ; ele
CEYLON JUNGLEFOWL (Gallus lafayettz) . : : 6 5 : : 5 ez
GREY JUNGLEFOWL (Gallus sonnerati) . ¢ c : 5 5 : 5a BY
JAVAN JUNGLEFOWL (Gallus varius). : : ; : A . : a 249

LIST OF COLOURED PLATES, PHOTOGRAVURES

AND MAPS

COLOURED PLATES

Prate XX]. WHITE-CRESTED KALEEGE Gennaeus albocristatus (Vigors)
Painted by G. E. Lodge. Frrontisptece
Over the great spruce forests of Kashmir and Garwhal chilly winds come roaring down
from Tibetan snow-peaks. But among the ferns and moss-muffled bases of the trees the
air is still, and fragrant with the odours of many forest flowers. Here the White-crested
Kaleege live; here they scratch for grubs and tubers, court their mates, make their lowly
homes, and at twilight roost upon high, swaying branches. Before dawn, a few will succumb
to the sudden attack of marten or weasel; the others awaken in early morning, send forth
their challenge and begin anew their daily life.

Pirate XXII. NEPAL KALEEGE Gexznaeus leucomelanus (Latham) . LHacung page

Painted by G. E. Lodge.

No white man has ever seen this bird in its native haunts, for it is confined to the
southern part of Nepal, where no Caucasian is admitted. It lives in the moss-hung oak
and spruce forests which cover the maze of tumbled mountain ranges of this little kingdom,
and is trapped wholesale by the Nepalese shepherds,

Prate XXIII. BLACK-BACKED KALEEGE Gennaeus melanonotus (Blyth)

Painted by G. E. Lodge. Facing page

Deep in the mossy, humid forests of Sikhim I have watched a pair of these pheasants

picking among the fallen leaves, and murmuring to each other in low musical tones. Later

the hen made her way to her nest among the ferns at the base of a great tree, and the cock

mounted slowly, branch by branch, to a lofty perch, and night settled quietly down over the
Himalayan wilderness.

Prats XXIV. PLUMAGES. OF THE BLACK-BACKED KALEEGE Gennaeus
melanonoius (Blyth) : : : ; . Lacing page

Drawn by H. Gronvold.

Even when young Black-backed Kaleege have moulted late, the shape, pattern and colour

of the adult plumage are not fully attained until after the first year.

Fic, 1. Chick in down four days old, collected in Sikhim, May 2oth.

Fic. 2, The sexes are clearly distinguishable in the juvenile plumage, the female showing
much warmer, more buffy tones, especially on the head, back and tail. In
this individual, the down is still present on the face, concealing the reddish
skin beneath.

Fic. 3. The juvenile male is darker throughout, with narrower white tips to the feathers.
The incoming dark, central tail-feathers are not clear black, but coarsely
vermiculated with grey.

Pirate XXV. BLACK-BREASTED KALEEGE Gennaeus horsteldi (Gray)

Painted by G. E. Lodge. Facing page
Among the moss-hung forests and the bamboo thickets of north-eastern Burma, I first
heard the bubbling murmur and cackle of this pheasant. In pairs or in small flocks they
work slowly through the ferns and over fallen logs, their scarlet facial skin glowing like the
ginger blossoms beneath their feet. A grouse-like whirr of their wings, or the sound of
scratching among dry leaves would often indicate their presence, but they were always on

the- alert, watching and listening for danger with keen eyes and ears.

vii

26

30

40

42

vill LIST OF COLOURED PEATEs

Prate XXVI, LINEATED KALEEGE Gennaeus Lineatus (Vigors) . acing page 52
Painted by G. E. Lodge.

A night spent in slumber on the branch of a tree; an early morning drink and search
for food; a midday siesta; again a feeding period and in the cool of early evening a leisurely
stroll to water and thence to roost; such is the epitome of the daily life of the Lineated,
as well as of most other pheasants.

They can be located often by the sound of their scratching among leaves, or the low,
undertone clucks and chuckles as they search for grubs or succulent tubers.

Prate XXVIII. CHINESE SILVER KALEEGE Gennaeus nycthemerus (Linnaeus)
Painted by C. R. Knight. Facing page ©2

This, the beautiful bird of “day and night,” ranges across the whole of south China,
from the Burmese border almost to the sea-coast of Fokien. It is found in open forest,
on clear, rolling hills and along swift rivers. While the Silver Kaleege is common in captivity,
yet no white man has ever seen the nest and eggs of a wild bird. Enormous numbers were
formerly killed for their plumage, but now the Chinese have developed a taste for their
flesh and trap and shoot the bird extensively.

PuatE XXVIII. HAINAN SILVER KALEEGE Geunaeus whitehead: Grant
Painted by G. E. Lodge. Facing page 72

This bird is very close to the Silver Kaleege, but as it is found only on an island .off the
south China coast, it must be recognised as a distinct species. There is less white on
the outer tail-feathers than in the silver bird, while the female is actually more like the
corresponding sex in the Lineated than in the Silver Kaleege. Mr. Whitehead, who obtained
the first specimens, died before he left the country, and since that time only Japanese
collectors have taken the bird.

Prate XXIX. EDWARD’S KALEEGE Gennaeus edwardst Oustalet . Facing page 76
Painted by G. E. Lodge.
Although known for a quarter of a century, absolutely nothing has been recorded of the

habits of this very distinct and beautiful pheasant. Four specimens have been obtained
from the province of Kaung-tri in Annam.

PrateE XXX. SWINHOE’S KALEEGE Geunaeus swinhott (Gould) . Facing page 78
Painted by G. E. Lodge.

In the deep inland forests of the mountains ot Formosa this splendid bird makes its
home. None but the natives have observed or captured it, and~there is no record of the
nest and eggs being found, but the bird lays readily in captivity and is not rare in zoological
gardens.

PuaTtE XXXI. RIPPON’S HYBRID SILVER KALEEGE . 2 . Facing page 92
Painted by G. E. Lodge.

This bird has been ‘given a name (Gennaeus rippond), but no two birds are alike, and there
is no doubt but that it represents individuals which have a very large percentage of the blood
of the Silver Kaleege, with a relatively small amount of Black-breasted ancestry. I found a
typical bird in a flock of eleven in northern Burma. They showed great variation, and could
not be classified with any specific exactness, and only one approached the description of this
hybrid form.

PLatE XXXII. OATES’S HYBRID KALEEGE . . . Facing page 94
Painted by G. E. Lodge.

This bird, which some ornithologists have considered to be a species (Geunuaeus oatesz),
shows great variation in the dozen skins I have examined. We must either give a half-dozen
new names to them or consider them as hybrids. Oates’s Kaleege represents birds with the
blood both of the Black-breasted and Lineated Kaleege, but the latter greatly dominant.

LIST OF COLOURED PLATES 1X

Puate XXXIII. MALAYAN CRESTLESS FIREBACK <Acomus erythrophthalmus
(Raffles) . : : : . ; : : Facing page 102

Painted by G. E. Lodge.

These birds are exceedingly pugnacious, and I have even seen hens fighting madly with
each other. Both sexes are armed with spurs. They live along the small creeks which wind
through the low jungle, and haunt the water-holes of the buffalo. The sound of warning or
challenge, uttered when two cocks threaten one another, is a deep, hoarse drawl, almost a snarl,
sounding as much like a small cat as a pheasant.

Pirate XXXIV. BORNEAN CRESTLESS FIREBACK Acomus pyronotus (Gray)
} Painted by G. E. Lodge. facing page 108

Low, wet jungle along the coast of Borneo is the home of this bird. I have seen a pair
feeding beneath a berry tree, upon the fallen fruit, together with the insects which have collected.
A small bird called out in alarm at the sight of some danger, and both birds squatted flat
among the begonias.. A scale-bird sang, and confidence was at once restored. A few minutes
later a great branch crashed to the ground, but the birds hardly glanced up, so perfectly did
they judge between actual danger and a harmless jungle happening.

Prats XXXV. SIAMESE CRESTED FIREBACK Lophura diardt (Temminck)
Painted by G. E. Lodge. Facing page 116

Sportsmen in Siam who sit up on the look-out for tigers, sometimes see this magnificent
bird step out from the jungle and walk slowly past, its fiery golden back flashing even in
moonlight. It lives in dense bamboo thickets, and comes into more open jungle, often near the
. banks of a river, to feed and drink. It is as easy to trap as it is difficult to observe, and the
Siamese bring many to the Bangkok market.

PratE XXXVI. PLUMAGES OF THE BORNEAN AND SIAMESE CRESTED
FIREBACKS Lofphura ignita and Lophura diardi . . Facing page 120

Drawn by H, Grinvold

Fig. 1. Chick of Bornean Crested Fireback, Lophura ignita.
Figs, 2. and 3. Bornean Crested Fireback, Lophura tgnita.
Juvenile plumage of the female (2), and male (3).
Fig. 4. Siamese Fireback, Lophura diardz.
Transition plumage between juvenile and first year, showing sequence and nodes of moults
and the unusual pattern complexity of the immature wing-feathers.

PratE XXXVII. MALAYAN CRESTED FIREBACK Lophura rufa (Raffles)
Painted by G. E. Lodge. Facing page 122

A single glimpse of these birds after many hours of nerve-racking search is the usual
reward. They are very shy and for ever on the alert for the slight crack or rustle of twig or
leaf, but their beauty is worth the longest stalk and the most wearisome wait. They are found
in pairs or in small families, and when alarmed go off with a headlong rush through the under-
brush in preference to attempting flight.

PratE XXXVIII. BORNEAN CRESTED FIREBACK Lofhura ignita (Shaw)
Painted by CRE night, facing page 128

This splendid bird chooses to live in low, swampy jungle, coming out to feed in the small,
shaded glades. In one such place, embowered with green and decorated with great elkhorn
ferns, a family of Firebacks spent many weeks. Although not far from a native Dyak house,
yet it was also close to a burial-ground, and the fear of evil spirits gave safety to the birds.

When standing quietly, the colours of the Fireback cocks merge with the hues of the
jungle, but at the slightest movement the glittering back detaches itself from the spots of sun-
light, and the blue-black body from the dark shadows, and the bird stands out in its full glory.

oe LIST OF COLOURED PLATES

Purate XXXIX. WHITE-TAILED WATTLED PHEASANT CPEs bulwere
Sharpe : : 2 : . Hacing page 146
Painted by G. E. Lodge.

Perched in a tiny half-cave in the rocky side of a deep gorge in central Borneo, I watched
the flying lizards, gorgeous as butterflies, pass swiftly from tree to tree, while long-tailed paradise
flycatchers swooped after flying insects, in and out of the shadows overhead. Without warning
there stepped into view three chevrotain, the tiniest of jungle deer, and with them a quartet
of White-tailed Pheasants, one a fully adult male with great, spreading tail sweeping the

ground. They walked slowly beneath me, and still unalarmed, passed from view in the direction
of the river.

PratE XL. RED JUNGLEFOWL Gadlus gallus (Linnaeus) - . . Facing page 172
Painted by G. E. Lodge.

To the human race this is the most important wild bird living on the earth, for it
represents the ancestor of all varieties of domestic fowl. It ranges from the border of Kashmir
to Singapore, and is found in the wildest regions, as well as close to native villages, with the
fowls of which the wild birds frequently cross. Its crow is bantam-like, and sounds strangely
out of place when heard in deep jungle. These Junglefowl are usually monogamous, the hens
lay from five to eight eggs.

Pirate XLI. CEYLON JUNGLEFOWL Gallus lafayette Lesson . . Facing page 212
Painted by G. E. Lodge.

In the dusk of early dawn the Junglefowl begin to leave their roosts and make their way
through the thorn thickets of south Ceylon to open glades. Here they send forth their loud
crow, chuck-George-Joyce! Were they battle with each other, armed with sharp spurs, and here
they scratch vigorously for worms and grubs. Sometimes a dozen birds can be heard crowing
in various directions, but after sunrise they cease, and with the coming of heat the birds seek
shelter under the dense foliage.

Prate XLII. GREY JUNGLEFOWL Gadlus sonneratc Temminck . . Facing page 234
Painted by G. E. Lodge.

The Junglefowl of southern India range from the sea-shore to an altitude of five thousand
feet. Mated pairs appear to remain together throughout the year. At the breeding season the
birds retreat to distant parts of the jungle, but at other times they often feed openly in trails
leading out of villages, especially where cattle are pastured. The spots on the hackles of the
cock are like drops of sealing-wax, and unlike any character found in other Junglefowl.

Puate XLII]. PLUMAGES OF THE GREY ea S Ald Gallus sonnerati
Temminck . . , . Facing page 246

Drawn by H. Gronvold.

The chick in down has very distinct patterns and coloration, and the wing feathers sprout
rapidly, so that it can fly a short time after hatching.

The juvenile plumage of both sexes resembles that of the adult female, with usually a hint
of the sealing-wax spots on the median wing-coverts of the cock.

A full-grown cock in the eclipse plumage has the specialised neck hackles replaced with
black ones. This partial moult lasts only for three months after the breeding season.

Prate XLIV. JAVAN JUNGLEFOWL Gallus varius (Shaw and Nodder) Facing page 248
Painted by G. E. Lodge.

Early in the morning, before the sun appeared, picking my way quietly through a tangle
of cactus on the low-lying Javan coast, with a shrike sitting in every tree, with bulbuls singing
from every thicket, there would come across the valley a sharp, crisp, virile chaw-au-auk! the
challenge of the Javan Junglefowl. With erect iridescent comb and plumage glittering with
dew, the splendid wild fowl was leading his family from their roosting-place to the nearest
pool of water for their sunrise drink,

LIST OF PHOTOGRAVURES ~ xi

PHOTOGRAVURES |

PHOTOGRAVURE 16. GARHWAL—THE HAUNTS OF THE WHITE-CRESTED
KALEEGE . ; : . Facing page 14

Photograph by William Beebe.

Deodars, spruce and silver firs cover many of the slopes of lower Kashmir and Garhwal, and
offer sanctuary to White-crested Kaleege Pheasants. In the pleasant twilight of these forests, each
glade seems to have its own dominant perfume. One is starred with myriad,.long-stemmed straw-
berry blooms; in another the pale blue faces of a multitude of violets show faintly. Shrubs are
hung with bright panicles of bell blossoms, and the background of ferns is here and there picked out
with the white lacework of saxifrage. Here we find the nest and eggs of this Kaleege, brooded by the
patient, brown mother, and here the little chicks, at her alarm note, dive among the leaves and moss
and remain motionless until reassured by her brood call.

PHOTOGRAVURE 17. SIKHIM—IN THE SACN TS OF “HE BLACK-BACKED
KALEEGE : : : . Facing page 34
Photograph by Wilkam Beebe.

On the sheltered open slopes, tree ferns filter the sunshine through emerald filigree. In the
neighbouring forests we find damp moss hanging from bark, twig and leaves, and the hoofs of deer
and the feet of pheasants sink deeply into soft mould. These kaleege are confined to a tiny bit of
the earth’s surface in Sikhim and Western Bhutan, but dangers are numerous and they are on guard
every moment of the day, watching for snakes, civets, or even little mild-mannered mountain bears,
who know how delicious a meal these birds furnish.

PyHotocravureE 18. WHERE HORSFIELD’S KALEEGE MAKES ITS HOME
Photograph by William Beebe. facing page 46

These pheasants wander in families or small companies through the dense jungles, coming

now and then upon beautiful streams, flowing slowly and quietly around the roots of great trees, or

pouring in a headlong cascade over a ledge of rocks. Here the birds slake their thirst and then pass

on again into the forest. I once watched a pair of birds scratching and pecking on a hillside among

green begonias and jack-in-the-pulpits, making the leaves and twigs fly in all directions. The

sound could be heard for many yards, but the vigilance of the birds never relaxed, and at the
slightest suspicion of danger they were off like two meteors.

PHOTOGRAVURE 19. NORTHERN BURMA—YUNNAN BORDER—HOME OF THE
HYBRID KALEEGE : : ; . Facing page 84
Photograph by Wiliam Beebe.
Standing on the high divide which shunts its eastern waters into China and its aegis into
the great rivers of Burma, a great, tumbled, irregular mass of mountains and valleys is seen. All
are forest, clad with bamboo, oaks and hard wood, and here, hidden beneath that vast extent of
many-tinted foliage, I found the northern Kaleege pheasants.

LINE OF KACHIN PHEASANT TRAPS

The Kachins and other native tribes use a deadly method of trapping. A low fence of split
bamboo runs uphill and down for a mile or more, with dead falls every few feet. When this is in
full operation, few pheasants fail, sooner or later, to be caught, and whole districts are thus cleared
of these splendid birds.

PHoroGRAvuRE 20. A SANCTUARY OF THE HYBRID KALEEGE . Facing page 86
Photograph by William Beebe.

The natives of northern Burma captured many specimens of Hybrid Kaleege for me, taking
them either in deadfalls or in nooses. But there was one spot near their villages where they would
never trap or kill. This was the sacred spot or grove devoted to the Nats or evil spirits. And here
the pheasants were safe from molestation, and here they scratched for food and roosted high at night.

X1i LIST OF PHOTOGRAVURES

PHOTOGRAVURE 21. HOME OF RIPPON’S SILVER AND OF EASTERN HYBRID
KALEEGE : j ' . Facing page
Photograph by Wilkam Beebe.

Three of these rare hybrids crossed this trail one day, headed into a wild, deep gorge. The
mountains rose high on all sides, except to the northward, where the purple distance ended in the
jagged ranges of the unexplored tri-corner of Tibet, Yunnan and Burma.

Parrakeets and flycatchers screamed, and at last I heard the tremulous wing-whirr of a
pheasant. But it was an hour later before the timid birds appeared—clad in ebony and white,
walking slowly downward, on their way to the water, at the bottom of the gorge.

PHOTOGRAVURE 22. HOME OF THE WESTERN HYBRID KALEEGE Facing page
Photograph by William Beebe.

In western Burma, where the ranges of Horsfield’s and the Lineated Kaleege approach or touch,

the jungles and lesser growth are inhabited by pheasants which show all degrees of intergradation.

Their voice, habits, eggs and young are all more or less alike, but on the plumage of the adult birds
is written their blood relationship to one or the other of their parents or more remote ancestors.

PHoTocravurE 23. HAUNTS AND BREEDING-GROUND OF THE LINEATED
KALEEGE 5 : , . Facing page

Photograph by William Beebe.

In central Burma, east of the Irrawaddy, we find a dense growth of small oaks, chestnuts and
pines. Blackberries ripen at the end of the rains, and the Lineated Pheasants come into the grassy
fields for berries and grubs. In the early morning it would not be difficult to approach closely,
were it not for the flocks of blossom-headed parrakeets which fly off screeching, spying one from
afar and warning every creature within hearing.

PHoToGRAvuRE 24. CHINESE HAUNTS OF THE SILVER PHEASANT Facing page

Photograph by William Beebe.

In Fokien, near the borders of Kiangse, the pheasants of “day and night” live among wild
ravines and mountains. There are no large trees, and the dwarf bamboos and shrubs have been cut
again and again by the yellow men for fuel. But here, in company with shrikes and mynas, and a
flock of magpies, I found the Silver Pheasant near the rivers.

PHoToGRAVURE 25. HAUNTS OF THE MALAYAN AND BORNEAN CRESTLESS
FIREBACKS . . ‘ ; . Facing page

Photograph by William Beebe.

In both countries these pheasants live in dense jungle, where they have access to open clearings,
or at least the trails made by wild animals. Here they strive to escape their enemies, snakes and
civet cats, which lie in wait or creep silently through the undergrowth. And here they nest,
although up to the present time no white man has seen nest, eggs or young, so timid are they, and
so skilful in hiding their home among the swamps and tangles.

PHOTOGRAVURE 26. HOME OF THE MALAYAN CRESTED FIREBACK

Photograph by William Beebe. Facing page
In the low, dense jungle of the Malay States, in thickets of thorny tangles and enmeshed
vines, these birds live. To study them one must fight hosts of leeches and mosquitoes, mud and
rain, or, when the sun appears, the steaming heat of these breathless places. Here the bulbuls sing
sweetly, and gorgeous butterflies flap slowly past, and here these pheasants scratch among the dead
leaves for food, or walk in the shallow water of jungle creeks, drinking and catching tiny creatures

in the sand and gravel.

PHoTocGRAvuRE 27. HOME OF THE BORNEAN CRESTED FIREBACK

Photograph by William Beebe. Facing page
The home of this bird was reached by means of a seventy-foot Dyak war canoe, in which I was

paddled up the rivers to the inland jungles. Here, in the haunts of gaily coloured pittas, of
glittering sunbirds and the glorious tropical orchids, these birds live and court their mates and rear
their young. Often the first hint of their presence is the call of the cock, a low, mumbled
Um—um! Um—um! followed by a single, sharp, keen whistle, which cuts through the warm,

quiet air like a knife.

88

56

64

104

124

132

LIST OF PHOTOGRAVURES xii

PHOTOGRAVURE 28. BORNEAN HOME OF THE WHITE-TAILED WATTLED
PHEASANT ‘ : . Facing page 150

Photograph by William Beebe.

On the slopes of the rolling hills of central Sarawak, in the warm jungles, sloping down to the
yellow rivers, these magnificent pheasants have lived for untold centuries. Only by days of the
most patient watching can they be seen, and more often a fleeting glimpse is all that rewards a
painstaking stalk.

Beneath the tangles of soft-fronded climbing ferns, or the solid phalanx of bamboos, the birds
may remain hidden a few yards distant, and never be discovered. The young birds are less wary,
and, trusting more to their inconspicuous colouring, often come into more open spaces, to scratch
for food or to preen their plumage.

PuotocravureE 29. WHITE-TAILED WATTLED PHEASANT . . Facing page 160

Photograph by William Beebe.

In the immature White-tailed Pheasant (left) the wattles are short and inconspicuous, In the
adult (right) they are long and wrinkled, and at the moment of courtship capable of being
considerably elongated.

The head-hunting Sea Dyaks of Sarawak are splendid hunters, and will spend days and nights
stalking and trapping pheasants and other birds. With the crudest kind of traps they would
capture the birds alive and bring them to me unharmed in wicker quakes.

PuotocravurRE 30. HOME OF THE RED JUNGLEFOWL ; . Facing page 174

Photograph by William Beebe.

In northern Burma I found the wild Junglefowl coming out to feed along the trails which lead
from village to village. They cluck and scratch among the turf, and take dust baths exactly like
our domestic fowl, and I have often found it difficult to shoot them, as the action seemed like
unsportsmanlike slaughter in a barn-yard. At the first hint of danger, however, they lower their
tails and run headlong, like pheasants, into the nearest underbrush.

The nests are hidden away in the clumps of bamboo, a mere hollow being scratched out, or the
eggs deposited on the dry leaves.

PHOTOGRAVURE 31. ROOSTING AND FEEDING-PLACES OF THE RED
JUNGLEFOWL . ; : . Facing page 182

Photograph by Wilham Beebe.

These birds roost high up, often on the half-bent curve of a tall bamboo, whose smooth stem
ensures safety from arboreal carnivores. As many as thirty birds have been seen roosting close
together.

Just as tame fowl are fond of the company of barnyard cows, so the wild Red Junglefowl are
often seen in the vicinity of wild cattle, and I have seen them again and again feeding about buffalo
wallows. It is almost impossible to flush these birds. They invariably choose to escape by
running swiftly away, yet, when surprised by a dog, they have the ability of rising as quickly and
strongly as partridges.

PHoToGRAVURE 32. JAPANESE LONG-TAILED FOWL . , . Facing page 104

Photograph by William Beebe.

Of all curious breeds of domestic fowl, none are more remarkable than the Japanese birds,
whose upper tail-coverts sometimes reach a length of over twenty feet. The feathers are kept
wrapped in soft paper, and the birds themselves spend a most uninteresting life in a high, narrow
box, fed by hand and cared for by special attendants.

PuHotoGRavurE 33. HOME OF THE CEYLON JUNGLEFOWL . Facing page 216

Photograph by William Beebe.

Along the coast of Ceylon, where the eucalyptus, acacias and mesquite abound, these
Junglefowl are abundant. The region is park-like, the thorn-bushes and trees alternating with
glades or larger open plains, dry and sandy, or with pools and grass. Here they must ever be on
the watch for leopards and civet cats, and at night they sleep in dense foliage on lofty limbs, well
out from the trunk, so that an enemy approaching along the branch would at once be detected.

XIV LIST OF PHOTOGRAVURES

PHotocravureE 34. NEST AND EGGS OF THE CEYLON JUNGLEFOWL

Photograph by William Beebe. Facing page

The nest is usually placed on the ground at the foot of a tree, and only the leaves and grass
already present are used as lining. The hen sits very closely, and deserts her eggs only when
danger is actually upon her. Her colours harmonise perfectly with the hues of the dry vegetation,
but the eggs, when exposed, are conspicuous. Five or six is the usual number, and they are as
small as the eggs of bantams, and sometimes slightly spotted.

A native Vedda, one of the aboriginal Cinghalese, was skilful in tracking down the birds and
their nests, and found one of my first nests, which the chicks had just left.

PHOTOGRAVURE 35. CEYLON JUNGLEFOWL . : ; : . Facing page
Photograph by Wiliam Beebe.

In the more arid places Junglefowl often suffer from the attacks of ticks, which attach themselves
to their comb in large numbers, and although they will sometimes preen one another’s plumage, they"
seem never to attempt to rid their companions of these pests.

The eggs vary from white to a rich cream colour. Usually they are unspotted, but occasionally
eggs will be found which are thickly covered with dots and spots of brownish red.

In trampled places in the thorn-brush, feathers are sometimes found, showing where severe
battles have taken place between rival cocks. But this photograph reveals a real tragedy. A cock
Junglefowl had been feeding on the insects which it had scratched from a nest of termites, when a

civet cat or similar enemy had pounced upon the bird, plucked out many of the larger wing feathers
and carried it away.

PuorocravurE 36. HAUNTS AND ROOSTING ISLAND OF THE JAVAN
JUNGLEFOWL . : . : . Facing page

Photograph by William Beebe.

This bird haunts the semi-arid coastal lands of Java, but, not satisfied with its proximity to the
booming surf of the tropical sea, I saw two cocks and a hen walk down to the beach, catching
insects as they went, and after some hesitation rise into the air as easily as quail, and fly out to a
tiny mangrove islet a hundred yards off-shore. Here they found a safe roost for the night, returning
to their feeding-grounds in early morning.

PuHotocRAvuRE 37. ROOSTING CLIFF AND HAUNTS OF THE JAVAN
JUNGLEFOWL . : : : _. Facing page

Photograph by William Beebe.

Jagged ridges of weather-worn coral and stony fields covered with a scanty growth of grass ;
spiny cacti sprouting everywhere, and an occasional palm or bunch of feathery bamboo—such is the
home of the Javan Junglefowl along the coast. In one place I found a number roosting in a
limestone cavern beneath the level of the ground. The birds make their way to the more fertile
spots near the creeks and search for insects along the rice-fields and crops of ground pea.

PuotocRavurE 38. HYBRID JAVAN JUNGLEFOWL : , . Facing page
Photograph by Wiliam Beebe.

The wild Junglefowl do not interbreed with the native poultry, but in captivity the two species
will occasionally cross. Some of these hybrids are huge creatures, with enormous pendant combs,
and beautiful plumage ; others are small and bantam-like with absurdly short legs. They are great
favourites with the Javanese, who keep them on triangle perches or in ingeniously woven quakes or
cages. The dominant character of these birds is the voice, which lacks cadence or definiteness, but
consists of a scream which must carry for at least a mile. This is the stimulus to much betting, and
owners of powerful-lunged birds often make large wagers on the vocal powers of the rivals.

222

224

250

252

260

PHOTOGRAVURE 309.

HYBRID

LIST OF MAPS

JAVAN

BAMBOO CAGE
Photograph by William Beebe.

The hybrid Junglefowl are sometimes valued as high as six hundred gulden.
wealthy natives erect a very tall bamboo pole in the compound, with a primitive sort of pulley near
the top. The fowl in its basket is attached to a rope and pulled up, high above all the surrounding
trees, to remain throughout the day, sending forth at frequent intervals its loud, piercing scream.

JUNGELEFOWL IN ITS”) AERIAL
: ; ; : ; facing page

It begins its call even while the basket is whirling around on its jerky ascent.

Mar VI. SHOWING THE DISTRIBUTION OF THE KALEEGE
Mar VII.

Map VIII.

»”

”

3)

MAPS

CRESTLESS FIREBACK
PHEASANTS

GRES PED: FIRE BACK
PHEASANTS :

WHITE-TAILED WATTLED
PHEASANT

JUNGLEFOWL

The chiefs and

XV

262

Facing page 4

100

114

144

170

Ne a es

GENNAEUS
KALEEGE PHEASANTS

Family PHASIANIDAE
Subfamily PHASIANINAE

Genus GENNAEUS

Tuis group is the largest component of the old genus Fuflocamus, which
formerly included Dzardigallus, Lophura and Acomus. It seems to me to
consist of two well-marked subgenera, Gexuaeus and Hverophasis, the former with
seven species, the latter with two. The species most familiar to those who keep
pheasants or have opportunity of visiting collections is doubtless the Silver Pheasant,
or Kaleege. One of the native names throughout the East is Kaleege or Kalij, and
I have chosen to use this to characterize the pheasants of this genus.

The least specialized type of Kaleege Pheasant is quite fowl-like, especially as to
shape of body and carriage of the tail, the latter consisting of sixteen feathers,
laterally compressed. These are strongly graduated in order from without inward,
the central pair being the longest, and at least three times the length of the outer-
most. There is great variation in the actual and relative length of the tail as a
whole.

The sexes are very dissimilar in colour, and both have crests usually long and
flowing (Gennaeus), rarely short, lying close to the head (Azerophasis). In the birds
with long crests, these may end in a point, or else be diffuse, covering the nape
and hind neck. The crest is usually disintegrated. In the subgenus Hervophasis
many of the contour feathers tend to convexity at the tip. Aside from this, there
is no specialization of feather structure.

The bill is moderately stout, strong and curved, being used as a pecking organ
in assisting the scratching function of the feet. The face is entirely bare, the skin
being covered with finger-like papillae, and showing a tendency to the formation of
three pairs of free wattles, one in front of and above the eye, the second back of
the eye at the side of the occiput, and the third at the gape. In related genera
these are carried to the point of highest development in Lodiopfhasis. The ist

primary is considerably shorter than the 2nd, which is equal to the gth or roth;
3

4 A MONOGRAPH OF THE PHEASANTS

the 5th or 6th is slightly the longest. Thé tarsus is much longer than the middle
toe and claw, and is armed in the male with a pair of stout, curved and moderately
long spurs. The feet and legs are strong, and vary greatly as to colour, being
scarlet, greenish, whitish or brown.

A generalized brownish olive is the dominant colour of the females, more or less

variegated with buffy yellow, white and black. The secondary sexual characters of

the males, with the exception of the spurs, consist only of increased size and more
specialized colour and pattern. This latter shows great variation, what may be
taken as the most generalized being an equal black and white vermiculation, produc-
ing a grey effect. On the one hand, we find a disappearance of the white, giving us
a series of black, or metallic glossed birds, with solid patches of white, and greyish
under-parts; and again a dominance of the white, the black lines being greatly
reduced. Finally the metallic hue may be varied so that the bird is brilliant in blue,
green and dull crimson. In the metallic group the feathers of the rump tend to be
truncated, with a disintegrated fringe, which may be of a special colour or even
structurally specialized. This hints strongly of the still more modified rump in
Lophura and Acomus.

GENNAEUS
Type
Gennaeus Wagler, Isis, 1832, p. 1228 , : : . % . G. nycthemerus
Nycthemerus Seeincen Class. Birds, II. 1837, s coal : Pe é by te 45
Alectrophasis Gray, List Genera Birds, ed. 2, 1841, p. 78 . : . : . G. cuviert
Grammatoptilus Reichenback, Nat. Syst. Vogel, 1852, p. 30 . 3 ! . G. lineatus
Fierophasis Elliot, Mon. Phas. 1872, p. 2, text to pl. XXV. . n . . G. swinhoie

For many reasons the genus Gexuaeus is the most interesting in the family Phasz-
anidae, and especially because of the puzzling nature of many of the forms. In the field,
my experience with these birds extended from the haunts of a@/bocristatus in Kashmir
to mycthemerus in Fokien, and from orsfeldi in Upper Burma to “meatus in Lower
Burma. I collected, or made notes upon, many of the forms, not only of adults in
full plumage, but of moulting birds and those in immature dress. Forearmed with
the details of what had already been published on the subject, I was able to search
more intelligently and with more direct design than would otherwise have been the
case. More than this, however, was the aid given by the English sportsmen in
Burma, who in some cases furnished me with data and specimens which were of
the greatest help. In one or two instances these were the very men who had
collected the original types, so their assistance cannot be over-estimated. It seems
quite certain that any future investigation of this group will but slightly affect the
general results I have reached. I feel all the more certain in stating this belief
because I find myself in almost perfect agreement with the general conclusions of
Prof. Alessandro Ghigi, at least as regards the method of evolution of these birds.
It is most significant that while the conclusions of this Italian ornithologist were
reached by a study of the phenomena of experimental hybridism, and those of mine
by independent observation of wild shot individuals, our results differ only in very
minor details. His paper to which I refer is Ricerche di Sestematica Sperimentale
sul geneve Gennaeus Wagler+

1 Memoria R. Accademia Scienze dell’ Instituto di Bologna (Bologna, 1909).

ee
og

MAP VI.

Bombay®

L BENGAL :
7 Madras f aA PAS PHILIPPINE SV res

ian as {sta | | fe ra
. ‘ le s °

ARABIAN SEA

EQUATOR

INDIAN OCEAN

JAVA SEA’ L0f
ee A

East of Greenwich nile) . + 120°

HFEGC.Witherby, Publishers. Stanford's Geograph! Estabt.

MAP SHOWING THE DISTRIBUTION OF THE KALEEGE.

Region 1. Gennaeus lineatus. Region 6. Gennaeus nycthemerus.
5 2 us horsfieldi. ise aly “f whiteheadi.
4 bk F, melanonotus. 18% = edwardsi.
A aed! 5 leucomelanus. MPa 5; swinhoei.
5 ., albocristatus.

The barred zones show where connecting hybrids occur.

KALEEGE PHEASANTS 5

The tangle which has arisen about this genus centres in Burma, and, in a word,
the whole question resolves itself into whether the numerous forms which have been
described from that country are valid species and subspecies or only hybrids, the
naming of which might be continued indefinitely without benefit to the understanding
of the genus.

In order even to begin to clear up the problem it is necessary to have a definite
starting-point, and for this I have chosen the Lineated Kaleege (G. “imeatus). Geo-
graphically and pigmentally this species occupies a central position. From the point
of view of colour and pattern it is the most generalized of all the species, while its
uniformity over a considerable range leaves no doubt as to its right to true specific
recognition. The black and white on the upper plumage is broken up into fine
vermiculation, the two being about equal in extent, the specialized colour extremes
of solid whites and blacks being thus lacking. A further hint of its more general-
ized type is the superficial resemblance which it bears to the related tropical genera,
Acomus and Lophura.

From this point, even a superficial survey of the genus shows two, and very
probably three, main lines of divergence. To the north and westward along the
Himalayan terai extends a succession of dark-feathered birds, in which the whites
are reduced or concentrated on certain limited portions of the plumage. From
east to west these are 4orsfieldi, melanonotus, leucomelanus and albocristatus. In
all four there is a small amount of variation, but, zzfer se, no actual crossing
has ever been recorded in a wild state, nor did most careful search and inquiry
on my part reveal a single instance. I have taken this-up in detail in my mono-
graph descriptions, and there is no need to reproduce it here.

It is important to note that the distinctions between the Himalayan Kaleege
Pheasants have been shown by Ghigi to behave in experimental crossing exactly as
though they have been derived by mutation, not blending as do the characters of
the Burmese birds.

To the north-east of /neatus we find a second-line of Gennaeus Pheasants,
of which the most widely distributed is uycthemerus, the Silver Kaleege. On the
island of Hainan is an offshoot from this, whzfeheadi, quite close to the typical
species, but, of course, worthy of specific recognition on account of its complete
geographic isolation.

A third line, quite distinct from “xeatus, is found to the south-east in Annam.
This is edwardsz, whose connection with /xeafus, at least as a direct offshoot, is at
present somewhat obscure. It is known only from a few specimens from Annam, and
its rather close relationship with swzwhoez is of great interest. The latter inhabits the
island of Formosa, and with its tropical, 4comus-like brilliance of colour differs much
from zycthemerus, the only species at present inhabiting south-east China. Edwards?,
however, shows a very probable former derivation, less specialized, and much nearer
the supposed centre of origin.

After carefully going over all the data available, I cannot see that there are any
other species worthy of recognition on grounds similar to the nine Kaleege I have
mentioned. .

Centring our attention again on Burma and the adjacent Chinese states, we

6 A MONOGRAPH OF THE PHEASANTS

find that most of the country is occupied by three species, Aweatus, horsfieldi and
nycthemerus. Now in all the genera of pheasants there is, as far as I know, no case
of two congeneric species occupying the same territory. This was not true as long
as veevest and soemmerringit were included in Phasianus, a very obviously erroneous
association, which I have attempted to better.1 We find that, in addition to the nine
Gennaeus 1 have already mentioned, there have been described under binomial names
no fewer than thirty-six forms. Almost without exception these are all found within
the range of one or the other of the three above-mentioned, widely-distributed species,
and not only this, but they occur along the borders where the three approach one
another. After studying these forms from every possible point of view, there is no
doubt whatever in my mind but that all are natural hybrids.

After the three main types of Kaleege became differentiated, they seem to have
approached one another again, working along the narrow, intersecting valleys of Central
and Upper Burma, where cross-breeding took place, apparently as freely as it does in
captivity. The English ornithologist, Mr. Eugene W. Oates, towards the end of his
life became interested in the Burmese Kaleege, and gathered all the specimens possible.
He had no belief in the hybridizing of these birds, and so strong was this attitude, that
the naming of new forms became an obsession with him. He grew intolerant of
criticism and welcomed neither argument nor proof, however convincing, that any
of his species were other than normally evolved forms. But aside from this his work
was sincere, and however limited his ability to view the subject from more than one
angle, he at least laboured to gather together a splendid collection, and we are greatly
indebted to him for giving this unusual phenomenon such prominence.

Of the thirty-six so-called species Oates is responsible for nineteen, a goodly
percentage of which are based on single specimens.

The conclusions which Ghigi draws from his experiments seem important enough
to quote in full, both the original and the translation :

“Returning to the birds forming the subject of the present study, if we proceed
to examine G. leucomelanus and G. horsfieldi, or rather this last one and G. Lineatus,
we see that they differ in a complexity of characters which, in the hybrids, are so
arranged as to produce an intermediate form, or else they transmit them in a different
association from that existing in the progenital species. Granted that these intermediate
forms, or with characters associated in a different manner, may be constant in their
descent, it is clear that two distinct species have the power to give origin to a new form
by crossing.

“Tf however, we consider G. leucomelanos and G. muthura, or else the first and
G. albocristatus, it is clear that from hybrids between them we are not able to obtain
new forms, as the difference consists in each case of only one pair of characters,
antagonistic and unresolvable. The hybrids of the first pair will have the white fringe
on the rump or will not be provided with it; those from the second pair will have the
crest black or will have it white; they will belong then either to one or the other
species, and even if they should be intermediate in the sense of having the white fringes
not as large as in /eucomelanos, or the crest not as light as in a/bocristatus, it is plain
that they cannot give origin to any new subspecies, because from what I have shown

1 Beebe, Preliminary Pheasant Notes, “ Zoologica,” I. No. 15, p. 283.

KALEEGE PHEASANTS 7

in my researches, and from what is established in the numerous hybridological literature
of to-day, these hybrids, which differ by only two antagonistic characters, give place in
successive generations to the separation of such characters according to the formula of
Mendel, returning thence to the pure progenital species.”

Riportandomi agli uccelli che formano oggetto del presente studio, se noi prendiamo esaminare il
G. leucomelanus ed il G. horsfieldi, oppure quest’ultimo ed il G. 4éneatus, rileviamo che essi differiscono par un
complesso di caratteri i quali, negl’ibridi, si fondono in maniera da produrre una forma intermedia, oppure si
trasmettono in associazione diversa da quella esistente nelle specie progenitrici. Dato che queste forme intermedie
od a caratteri diversamente associati, siano stabili nella loro discendenza, ¢ chiaro che a due specie distinte compete
la facolta di dare origine per incrocio a nuove forme.

Se invece consideriamo il G. encomelanos ed il G. muthura, oppure il primo ed il G. albocristatus, & chiaro
che dalle unioni provocate fra di loro non si possono ottenere nuove forme, in quanto la differenza consiste per
ciascun caso in una sola coppia di caratteri antagonisti ed indecomponibili. Glibridi della prima coppia avranno
le frange bianche sul groppose o ne saranno sprovvisti; quelli della seconda coppia avranno il ciuffo nero o lo
avranno biancastro; apparterranno dunque nettamente o all’una od all’altra specie, e quand’anche essi fossero
intermedi nel senso di avere le frange bianche non cosi ampie come nel dewcomelanus, od il ciuffo non cosi chiaro
come nell’albocristatus, & evidente che essi non possono dare origine ad alcuna nuova sottospecie, perché da quanto
ho esposto nelle mie ricerche, e da quanto si rileva nella numerosa letteratura ibridologica odierna, questi ibridi
i quali differiscono per due soli caratteri antagonisti, danno luogo nelle generazioni successive alla separazione di
tali caratteri secondo la formula mendeliana, ritornando quindi alle specie progenitrici pure.

The genus Gexnaeus, as I conceive it, is thus a fairly well-marked group inhabiting
moderate elevations in mountainous countries along the southern Himalayas, and
eastward throughout Burma and South China. Nine valid species of Kaleege are
recognized in this work, as follows:

White-crested Kaleege . . : : 3 t : . Gennaeus albocristatus (Vigors).
Nepal Kaleege : : : ‘ t 2 : : . Gennaeus leucomelanus (Latham).
Black-backed Kaleege . : , : ; : : . Gennaeus melanonotus (Blyth).
Black-breasted Kaleege . 2 : ! : : : . Gennaeus horsfieldi (Gray).
Lineated Kaleege . : . . : : , ; . Gennaeus lineatus (Vigors).
Chinese Silver Kaleege . : : : ‘ : : . Gennaeus nycthemerus (Linn.).
Hainan Silver Kaleege . : : ; ; 4 . . Gennaeus whiteheadi Grant.
Edward’s Kaleege . P 5 ; : : ; : . Gennaeus edwardsi Oustalet.
Swinhoe’s Kaleege . : : 4 : : ; ; . Gennaeus swinhot (Gould).

This is by far the most conservative opinion which has been expressed upon this
group. In 1895 Ogilvie-Grant, with very little material, admitted eight species and three
subspecies, Professor Ghigi allowed twenty-four species, Sharpe in the ‘‘ Hand-List of
Birds” recognized sixteen, and Baker in his 1915 revision lists eight species and eight
subspecies. In my first review of the group (“ Zoologica,” No. 17, 1914) it seemed to
me that nine species and four subspecies were legitimate, but on a final examination
of my material, taking feather by feather, wings, tail and contour plumage, and with the
addition of a number of recently collected specimens, I find too great variation in the
intermediate forms to permit consideration of them as other than hybrids. Two
male /neatus from the heart of the supposed oatesz territory and a pair of mated
birds, ¢ 4ueatus and ¢ sharpec from southern Siam helped to bring me to this
conclusion.

The phenomenon of the hybrids is, however, of as great interest as if they had
been geographical subspecies.

Throughout my explorations in the Gexxaeus country, from the humid, hot terai
of Kashmir and Garhwal, to the coastlands of Fokien, I was constantly impressed with

8 A MONOGRAPH OF THE PHEASANTS

the apparent quick reaction of the plumage pigment to the environmental conditions.
At the time I wrote of this in my descriptions again and again, and I have allowed
these interpretations to stand in the monograph text. But I cannot accept this
explanation as covering the entire question. The same thing has struck Baker, and he
tends to give humidity and other meteorological factors considerable weight. Speaking
of the Himalayan birds, of the Lineated and the Silver Kaleege respectively, he says,
“Thus we find that the black and very dark birds inhabit areas of dense forest at com-
paratively low elevations where there is a heavy rainfall, and which are therefore well
provided with rivers, lakes and swamps. Next we see that the grey birds inhabit
hills of moderate height covered with mixed forest, bamboos and grasslands, and with
a moderate rainfall. Thirdly, we obtain the white birds only in hills and plateaux at
a considerable elevation where the grass-covered and open country exceeds in extent
the forest, and where the rainfall is slight or even scanty.

“We thus have it demonstrated that great humidity and heat, with its constant
tropical growth of vegetation, induces black in the plumage of the birds of this genus,
whereas the coldness of the higher mountains, combined with a drier atmosphere and
its consequent higher forests and more open grasslands, induces white. These four
factors, temperature, humidity, elevation, and vegetable growth, we shall find, therefore,
are the principal ones governing not only the differentiation of the species, but also
of the intermediate subspecies.”

Believing thoroughly, as I do, in the darkening effect of humidity, it seems
reasonable that this explanation is correct. But I am convinced that it only partly
explains the changes in coloration, and that we shall have to await the discovery of
unknown factors to be certain. I have kept Himalayan and Silver Kaleege in captivity,
under the same extremes of humidity, for many years, have cross-bred and bred them
pure, and I have never noticed any change in successive moults or successive generations,
either toward melanism or albinism. As I shall mention elsewhere I have shot wild
birds from the same flock which differed widely from one another, and in Fokien I
have found Silver Kaleege living in numbers in low, coastal, humid country. While
these arguments are only partial, yet they make us wary of accepting a wholesale
explanation of the Kaleege pigmentation. It is a problem of transcendant interest,
which it is hoped may be elaborated before these birds become extinct in their native
haunts.

KEY TO GENNAEUS

I. Colour dominately black and white (males).
a Lesser and median wing coverts black and white.

a Black and white equal ; : : : : : : : : : . lineatus.
&' White dominant over black.
a’ (Outer web of outer tail feathers streaked with white : ; : : . nycthemerus.
6 Outer web of outer tail feathers wholly black 5 : : : : . whitehead.
b Lesser and median wing coverts glossed with blue.
c’ Crest white . ; : . : : , j : ; : : : . albocristatus.

ad Crest black.

te

c’ Under parts greyish white.
a” Rump fringed with white . 5 i : . . : : 4 . Leucomelanos.
0” Rump not fringed with white. . . 3 : : : ; . melanonotus.

a’ Under parts black , ; ‘ : ; ; : Ane : . _ horsfieldi.

KALEEGE PHEASANTS

c Lesser and median wing-coverts glossed with green.
e’ Scapulars greenish- blue
J Scapulars crimson
II. Colour dominately brown eeseeles),
a Feet scarlet.

7

a Crest well developed.
a” Lower parts entirely olive brown
d Lower parts variegated with light, dark ona Back Sires
o’ Crest very short.
c’ Primaries uniform brown ;
ad” Primaries barred with chestnut and ieee

b Feet greenish, brown or whitish.

c’ Back with light V-shaped stripes; lower parts with white and black V-shaped

stripes :
dad Back uniform; lower parts ils fehtee Sates Bevis:

at

Eee OLest Rrniont and rigid
J” Crest refulgent.

at

a" More greyish brown (Western Himalayas)
5 More blackish brown (Nepal)

Ls

C More rufous brown (Sikhim)

VOL, II

edwards.
swinhoet,

nycthemerus.
whiteheads,

edwardst.
swinhoet.
lineatus.
horsfieldi,
albocristatus.

leucomelanus.
melanonotus.

WHITE-CRESTED KALEEGE

Gennaeus albocristatus (Vigors)

NAMES.—Specific: albocristatus, L. albus et cristatus, white-crested. English: White-crested Kaleege.
French: Faisan a huppe blanche. German: Weisshaubenfasan. Native: Kalij or Kaleege (Kumaon and
Garhwal); Kookera, Kala-Murgha, Meerghi-Kaleege (Hills north of Mussooree); Kaleysur (male) Kalaysee
(female), (Pahari Hindi); Kullu, Mandi, Suket; Kolsa (Western Punjab).

BRIEF DESCRIPTION.—Male: Long, hairy crest white ; upper parts and throat glossy steel-blue ; mantle and
upper tail-coverts narrowly, lower back and rump broadly margined with white; fore-neck and breast dirty white,
shading on the remaining under parts into whitish-brown lanceolate feathers. Female: Hairy crest brownish-grey ;
plumage in general reddish-brown, brighter on the rump and under parts, the upper parts finely mottled with black
and edged with grey ; wing-coverts and under parts edged with white ; under parts with white shafts. In general
paler than the other Himalayan Kaleege,

RANGE.—Lower and middle ranges of the Western Himalayas.

LEE, BERD STINE ES EvAGaNGEsS

Down from the Tibetan snow-peaks a chilly wind came roaring through the spruces.
In its wake swirled a dense cloud of pearly-grey, dimming the foliage, obliterating the
distant slopes. Then, peal after peal of thunder reverberated through the mountains.
Between two rough-barked deodars I was crouched in my green umbrella tent on the
watch for kaleege or koklass pheasants. As the wind howled past, the tree-trunks
creaked and groaned, dead twigs hurtled down upon the canvas, and I hung tightly on to
the pole to keep the tent from blowing bodily away. As the last echo of thunder died
out, the loud call of a koklass pheasant came from far off among the deodars and below
me a kaleege called. Then the storm passed quickly. The grey haze across the western
valley, etched with long, sloping lines of rain, showed how closely I had escaped a
drenching. A glow of light touched a rocky peak to the north, and with terrific speed
the sunlight swept toward me, down to the tumbling stream and up again, bursting upon
me with a glow of warmth which effaced every trace of chill from the air.

Not as yet knowing the habits of the pheasants, I had sought to cover as wide a
field as possible during my first day or two of watching. I had pitched my observation
tent upon a low saddle, dropping away rather steeply on each side, and rounding upward
across the valleys, with their long, dark slopes of spruce and oak. The hillsides were
partly bare and grassy, partly covered with low, shrubby growth. The shrubs were
bright with panicles of bell-shaped blossoms and the setting of green grass held here and
there golden patches of buttercups, picked out with the white lace-work of saxifrage.

At this season the pheasants were still courting; the titmice were still in the
midst of housekeeping, but other young birds had already left their nests. Two such
families—utterly unlike one another—showed themselves near my tent. Up the trunk
of a spruce crept two bits of feathered bark, Himalayan tree-creepers, one quickly and

searching every cranny, the other slowly, rather uncertainly. Not a sound did the young
1f@)

WHITE-CRESTED KALEEGE II

creeper utter, but from time to time fluttered his wings in that never-to-be-mistaken
pleading for food of a young bird.

The second nursery scene was enacted upon a lofty bough of a dead spruce. A
series of complaining, feline mews attracted my attention, and there perched a great,
full-feathered, baby nutcracker, flapping its wings and endeavouring to look starving and
dependent. One of the parents approached, and the mewing increased to a perfect
medley of cries, screams and croaks—all merging at last into a contented mumbling
guregle as the food was crammed down its throat. Here we had the greatest contrasts of
creation : the defenceless, weak creatures of the wild, who trust to silence, dull feathering
and cautious, mouse-like creeping up of tree-trunks to preserve them from their enemies,
and the blatant, care-free ones who, from their combative ability, know that they may mob
a hawk with impunity, and hence may cry aloud their every emotion, perch in full sight
of the world and yet live. |

A sudden sharp cry and much craning of neck on the part of one of the parent
nutcrackers led my eye downward, and cutting a new loophole in the tent, I was
surprised to see a big bear nosing unconcernedly among the buttercups and turf half-
way down the slope. This was the second time I had encountered bears in my search
for pheasants. As he dug deeper and the earth tickled his nostrils he uttered sundry
snorts and coughs, but appeared to find edible somethings which compensated him for
the discomfort. He moved slowly on and upward until he was not more than forty feet
from the tent, and I reached out behind and worked my three-barrelled gun into a more
_ convenient position.

The use he made of his senses was interesting. Although fully half of my tent was
in view and far from matching the dark green of the deodar needles, yet the bear saw
nothing unusual, even when staring straight at me. Ever and always his mobile
snout curved upward from his teeth, and his nostrils quivered, but even those wonderful
organs told him nothing of my whereabouts, the breeze blowing down the valley, carrying
every scent before it. His sleek coat glistened in the sunlight, bowing out in smooth,
rolling curves as with each motion the great muscles rippled under the skin. Now and
then he seemed to scent some particularly delicate morsel, and in his eagerness to root
it out he moved slowly about the hollow, almost encircling it, with his muzzle buried
deep within it. A flat hind foot would be lifted and held trembling as he lunged and
lunged with bare teeth to scrape the tuber or succulent root.

The sun had dropped low toward the jagged ridge top when a White-crested
Kaleege appeared—the first I had ever seen in its native wilds. I knew it even when,
fifty yards away, it stepped for only a moment into view. A breath of air caught the
long, filmy crest and splayed it out—glistening white, at this distance, against the dark
of the plumage. The bird vanished, then reappeared higher up. Suddenly a second
black bird beat swiftly along the slope toward me, dodging through the rose and
raspberry bushes, and sailing gracefully to the ground. Just before it alighted the
white crest and black feathers flashed out—a second cock kaleege. The moment it
stopped it straightened up, and with strong, vigorous beats it sent forth its challenge, a
deep, reverberating wing-drum, woof/-woof-woof-woof! woof! woof !

All this took place just below and within a few yards of the bear. The bird looked
at the great black form intently, but showed no symptoms of terror, and a moment later

12 A MONOGRAPH OF THE PHEASANTS

the bear discovered the pheasant, stretched out his nose, sniffed, and went on with his
grubbing. The bird went behind a bush, and immediately there came forth a great
scratching and scuffling among the dead leaves. I marvelled at the recklessness of this
usually wary bird, but the noise was explained when a single small babbler appeared,
still scratching up the debris, or picking the leaves up one by one and hurling them over
its back, after the manner of ant-thrushes in the South American jungles. This one
small bird made a far greater disturbance than the pheasant and bear together.
When the kaleege again came out it walked slowly past the bear, making a slight
detour, but within twenty feet, and actually, when a few yards uphill, leaped upward at a
panicle of blossoms and caught some insect. So accurately do the wilderness folk gauge
one another’s possibilities of harmfulness !

But at this point the pheasant saw my tent, and though there was no sudden alarm,
yet a long, searching scrutiny left a suspicion, which dominated all other feelings, and
with tail and head raised, with graceful, dignified steps, it made its way ten yards or more
along the slope of the hill before it ventured to pass the tent. On the summit the bird
hesitated a moment, standing silhouetted against the distant peaks and the sky, a
beautiful symbol of alert eyesight and poised body.

In the never-ceasing warfare of the Himalayan jungles, this individual was a
success ; it had pitted its weak form, but keen eyes and ears, against the powerful
muscles and delicate power of scent of the beasts of prey. As this thought came to
mind a fox barked in the distance. The bear gave a last snort, left his field of manna
and ambled slowly over the ridge beyond me. He had not descended twenty feet before
he stopped, as if blocked by some insurmountable, material wall. For one instant his
nose was stretched out toward me, distorted with the agony of supreme endeavour, and
then, whirling in his tracks, he fled over rocks and turf, tearing through shrubs and
bushes headlong, recklessly, from the dread hidden danger. I wished him luck in his
strange life—a creature who by teeth and relationship should be meat-eating and
animal-slaying, but who prefers berries and roots and a fat, easy life, seldom or never
molesting the hunter unless wounded or in defence of cubs. How did the pheasant
know this? How too did the pheasant read danger in the first retrogade movement of
the bear, for no warning scent had come to its all but dead nostrils, and only the vaguest
of suspicions through eyesight? Yet at the sudden turn of the bear the kaleege leaped
into the air with a single subdued note, and after a few rapid wing-beats sailed in a
beautiful long, descending curve far down into the valley, almost to the stream for which
it had been headed. ‘Thus was a bird’s thirst quenched the quicker, and that of a bear
delayed by my hidden presence on the ridge top.

In the shadow of the hill I began my long walk back to camp, passing through the
favourite feeding-grounds of the White-crested Kaleege. The slight breeze had died
down, and as I reached the zone where rain had fallen, the air became heavy with scent.
In passing through the pleasant gloom of the conifer forest each glade seemed to have
its particular dominant perfume, governed by the predominating blossom of this lush
season of the year. One was starred with a myriad long-stemmed strawberry blooms ;
in another the pale blue faces of a multitude of violets showed faintly—more noticeable
by the sweetness of their odour than their colour. The dominant hue of the forest
blossoms was white, and in another glade was a host of large anemones, many splashed

WHITE-CRESTED KALEEGE 13

with bluish on the under side of their petals. Elsewhere, and thriving as well in the
shade of the spruces as in the glades, were mysterious, fairy star blossoms, flowering
apparently on the dead stem of last year—a strange resurrection—while the new, downy
leaves sprouted independently on a new stem from the ground. These too all had their
individual odour.

Most fragrant and evident to this dulled sense of ours were the spicy perfumes of
the trees themselves. Who can describe these, or who, in the case of the spruce, needs to ?
It was the same bracing, keen-cut scent that greets one in the Canadian wilderness ;
that crosses the bow of one’s sampan in the rivers of eastern China; that soothes one to
sleep in the Rockies, or that is borne on the velvet breeze from a Florida ‘‘ hummock.”
The silver fir sent down this Himalayan valley a strong, resinous odour, as penetrating
as the points of its blue-green needles, while the fragrance of the spruce was fully as
aromatic, but less pungent. One could call it almost sweet, were it not for the perfume
of the real flowers. The deodar—tree of God—had a fainter fragrance, but none the less
Spicy and resinous.

And so, as the dusk of twilight approached, I made my way past the scratched-up
turf of the glades, where, during the day, the pheasants had sought and found their food,
who now, their thirst quenched, were seeking their safe roosting-places high up among
the branches of the spruce or fir. A few would succumb before the dawn to the sudden
attack of marten or weasel, others would awaken in early morning, send forth their
challenge and begin anew their daily life.

GENERAL DISTRIBUTION

This kaleege is confined to the Western Himalayas, inhabiting the lower and
middle ranges from Hazara on the west, through Kashmir, Garhwal and western
Kumaon, but probably not actually found in Nepal itself. On the south it is found in
the outer range of the Siwaliks at a thousand feet elevation, up to a height of ten
thousand feet near the snows of the mid-Himalayas.

GENERAL AccounT.—The preference which the White-crested Kaleege shows for
the vicinity of native villages, together with its comparatively unsuspicious nature,
has tended to make this quite the best known of Western Himalayan pheasants.
Several accounts have been written of it, those of Wilson and Baldwin being perhaps
the best. I have added to the observations which I was able to make those which
could be obtained only by some one resident throughout the year in the haunts
of this bird.

There are few types of localities within its range which are not at one time or
another frequented by the White-crest. On the other hand, after finding the exact
kind of forest which in one place is beloved by this bird, the same species elsewhere
may be found confined to an entirely different vegetation zone. As with other game-
birds in these mountainous regions, the annual migration is little more than one of
altitude. In the autumn and winter almost all the birds will be found low down |
on the outlying flanks of the Siwaliks, perhaps a thousand feet or less above sea-
level, or at the bottom of the lower valleys in the more central ranges of Garhwal.
Here they frequent old grain-fields near water, or camping-places near more or less

14 A MONOGRAPH OF THE PHEASANTS

frequented roads. In the summer they work upward to an elevation of nine or ten
thousand feet. Only a few individuals, however, attain the latter height, but I have
found them in abundance at seven and eight thousand feet.

Their fearlessness in approaching the habitations of man is in striking contrast to
the habits of most of the pheasant family. Even though in any one region there
may be comparatively few White-crested individuals, yet the frequency with which
they show themselves at the edges of grain-fields and along roads may give the
impression of a very common bird, whereas in an Impeyan country, while there may
be many more individuals within as limited a zone, yet it is seldom that one is able
to catch even a glimpse of more than a single bird.

Especially in the winter months, the kaleege is to be looked for in the rather
open forest, along the borders of the more dense jungles, in overgrown thickets near
long-deserted cultivated patches, ruined cowsheds and even inhabited native hamlets.
In fact it is seldom or never that ‘the bird is found in remote, isolated forests.
Where the White-crest is, there, or in the vicinity, one may almost certainly expect
to find the presence of man, or at least some trace of his former occupancy. One
writer has well summed up the favourite haunts of the bird as “among the thick
clumps of bushes and shrubs near the banks of rivers, in low valleys through which
streams of water run, and on the slopes of hills where there is plenty of low bush
cover, especially thorny thickets bordering on cultivation. In the early morning the
vicinity of an old, deserted cowshed is a sure resort of this bird if anywhere in the
neighbourhood. I have flushed this pheasant and the common red jungle-fowl
from the same description of cover at the foot of the hills.”

I have found the low mountain bamboo or ringal to be a favoured cover during
the day, and in the interior of Garhwal it frequents damp evergreen forest, and the
proximity of streams at the bottom of shady ravines. Elsewhere I have found it
at six thousand feet in a semi-arid country, partly bare, partly covered with dense
euphorbias, with daisies and edleweiss-like blossoms relieving the stretches of barren
rock. Still higher it was in dense undergrowth in coniferous forest, varied with
oak saplings, raspberries and wild rose. And finally I have flushed the White-
crested Kaleege from an almost pure culture zone of rhododendron forest. The
typical forest floor where these kaleege love best to wander and feed is carpeted
with a thick matting of dead leaves, through which sprout maiden-hair ferns and
many other species; coarse and feathery-fronded brakes; ill-smelling filmy white
flowers, and lilies-of-the-valley giving forth the most delicate of perfumes; Solomon's
seal, and a host of tiny deodar shoots.

On the whole this pheasant cannot be said to be gregarious. But monogamous
they certainly are—despite more than one accrediting of polygamy. I have never
seen more than a pair together at the breeding season. At other times single cocks
are not uncommonly to be met with. When three, four or more are seen associated
the majority will be found to be birds of the year. Even when ten or a dozen
pheasants are flushed from a single bit of woods, the association is usually accidental,
or due to a local abundance of food. At a high altitude I once watched for several
days a little open platform, jutting out from an evergreen forest, which was
frequented by kaleege. On four consecutive days there were four, eight, four and

a i ti ie

PHOTOGRAVURE 16

=

GARHWAL—THE HAUNTS OF THE WHITE-CRESTED KALEEGE

Deropars, spruce and silver firs cover many of the slopes of lower Kashmir and Garhwal, and offer
sanctuary to White-crested Kaleege Pheasants. In the pleasant twilight of these forests, each glade seems
to have its own dominant perfume. One is starred with a myriad, long-stemmed strawberry blooms; in
another the pale blue faces of a multitude of violets show faintly. Shrubs are hung with bright panicles
of bell blossoms, and the background of ferns is here and there picked out with the white lacework of
saxifrage. Here we find the nest and eggs of this Kaleege, brooded by the patient, brown mother, and
here the little chicks, at her alarm note, dive among the leaves and moss and remain motionless until

reassured by her brood call.

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PHOTOGRAVURE PLATE |6

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GARHWAL= THE HAUNTS OF THE WHITE-CRESTED KALEEGE

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WHITE-CRESTED KALEEGE 15

sixteen birds, all busily feeding. The turf was dying from the attacks of innumer-
able grubs, and it was apparently these which attracted the birds. Of the sixteen
seen at once, three were full-grown males, all the rest apparently birds of the year,
with many scattered brown juvenile feathers remaining in their plumage. These
had not as yet paired off, the breeding season not having begun at this high altitude.
A few hundred yards away a single adult pair of pheasants were feeding by
themselves for three out of the four days.

When feeding as these were doing, or when, passing slowly through forest, the
pheasants are suddenly disturbed by catching sight of a man, they usually run
swiftly off, and seldom fly unless come upon suddenly or rushed by dogs, and even
in the latter case in dense undergrowth they often choose to lie close and risk
discovery until the danger becomes acute. White-crests are never as shy as
Tragopans or Impeyans, and where they are not constantly annoyed or shot at by
natives or sportsmen, they are as tame as the most amateur hunter Sahib could
desire. When walking up a ravine or up the slope of a hill, if pheasants are flushed
by dogs somewhat above, they will often fly into the trees just over the sportsman’s
head, and be so occupied with watching the dogs that several may be shot one after
the other. “When flushed from any place where they have sheltered, whether on
the ground or aloft, they fly off to some distant cover, and alight on the ground in
preference to the trees.” In the case of the sixteen birds which I have mentioned
as being observed feeding together, at my first shot all flew uphill some forty or
fifty yards. This is quite an unusual habit, and it is seldom that they will thus put
forth the necessary exertion of actually beating upward. When suddenly alarmed
they usually fly down the slope, or as often on a level along the hillside. The
kaleege flaps rather heavily, and with rapid beats at the beginning of the flight,
but soon acquires terrific speed—greater, several sportsmen have estimated, than the
burst of a rocketing English pheasant. Referring again to the sixteen pheasants,
five minutes later, when flushed by my native boy, all flew downhill and alighted
in trees some distance below me.

When surprised at a distance of twenty or thirty feet in dense underbrush, they
never attempt to fly, but invariably run quickly away, with neck outstretched and
tail lowered. Wholly undisturbed, and walking along slowly, not feeding, the White-
crested Kaleege cock has a splendid carriage, head, crest and tail raised, lifting the
feet high and daintily, and occasionally uttering a murmuring sound.

The variety in the many written descriptions of the notes of this pheasant is
probably as much due to the fact that they really have an extremely varied voca-
bulary, as well as to the lack of attention which the average sportsman gives to vocal
or other manifestations of a game-bird when it gets up before his gun.

When flushed by a man and actuated by only a comparatively moderate degree
of terror, I have always known the birds to utter a rather low whistled clucking,
very unlike the noisy koklass, which scream or squawk on any provocation. When
flushed by a dog and thoroughly alarmed they begin to cluck the instant they are
a-wing, and gradually gain in rapidity and loudness of utterance, the notes being
sibilant—se ! se! se! se-se-se-se-se-se—Si1p! Sip! Sip! Sire! Srp! One writer says,
~ “Their call is a loud whistling chuckle or chirrup; it may occasionally be heard from

16 A MONOGRAPH OF THE PHEASANTS

the midst of some thicket or coppice at any hour of the day, but is not of very
frequent occurrence. It is generally uttered when the bird rises, and if it flies into
a tree near, often continued some time. When flushed by a wild cat or other small
animal, this chuckling is always loud and earnest.” Again we read that “the call
of the bird, which may be heard at all times of the day, is a sharp fwut! twut !
twut! sometimes very low, with a long pause between each note, then suddenly
increasing loudly and excitedly. Generally speaking, when uttering this cry, which
at times might be mistaken by any one unacquainted with it for that of some small
bird, the kaleege is alarmed by a prowling marten, or a hawk hovering overhead,
perhaps a dog, but still oftener it is heard when a pair of cocks are about to
engage in mortal combat.” The call of the younger birds is noticeably harsher, less
sibilant than that of the adult pheasants. Aside from the vocal utterances uttered
under stress of excitement or fear, the kaleege is rather a silent bird.

The present genus is characterized by a kind of instrumental music, closely
akin to the drumming of the American ruffed grouse, but this whirring, produced
by the rapid vibration of the wings, is chiefly of the nature of a challenge, so that
I shall treat of it more in detail under the notes on courtship and fighting.

The food, as far as I could ascertain, is obtained only upon the ground. A
very reliable observer, however, reports that he has twice seen them perched in
bushes, feeding upon berries. Unlike the Impeyan, the pheasants of this genus are
essentially scratchers, and, like the common fowl, dig vigorously first with one, then
the other foot, sending the earth flying, and in productive spots the kaleege dig
hollows of considerable depth and extent. Their food has been rather indefinitely
and comprehensively stated to consist of grubs, roots, insects, seeds, berries, leaves,
acorns, soft roots and the young shoots of shrubs.

In Garhwal, in the month of May, I found these White-crested Kaleege feeding
chiefly upon small moths of several species. A gvotis consanguinea Moore, was by
far the commonest species, together with several small Geometridae. The following
is a detailed summary of the food of four pheasants, shot on different days and in
different places, but within a radius of twenty miles.

1.—é Shot 9.30 am. Crop empty; gizzard contained a comminuted mass of food,
including an earwig, a small centipede, many small bits of moss and
numerous young fern-shoots.

2.—$ Shot 6 p.m. Crop full, contained 1 spider, 3 small centipedes, 1 moth, 2
small beetles—a Tenebrionid of the tribe Aelofini, and an Elater—2o
small snails of several species (Vanina glauca Benson, Nanina percotteti
Pfr., Cyclotus strangulatum Tutton); 11 green leaves of Savrcococca
pruniformis,; 1 dead leaf, 3 roots and a large leaf-stalk of a fern, probably
an <A spidium,; 30 three-cornered brown seeds of some conifer; 12 pea-like
seeds; 18 white Daphne blossoms; several leaves of a Smilax; 1 large
leaf and many round brown fruits of a Vitis; besides a mass of com-
minuted vegetable matter. The gizzard contained similar food, much
comminuted.

3-—8 Shot 4.30 p.m. Crop contained 38 moths—(all but one were Agvotis consan-

WHITE-CRESTED KALEEGE 17

guinea Moore); 1 black beetle; 2 small, round snail-shells, and a small
amount of vegetable matter. The gizzard contained remains of about a
dozen moths; 1 large turreted snail-shell, and considerable comminuted
vegetable matter, chiefly leaves.

4.—9. Shot 6 p.m. Crop contained 14 small moths ; remains of moths and comminuted
vegetable matter in gizzard.

As I have observed in writing of the koklass pheasant, the higher forests of Garhwal
were, during the month of May, alive with small moths, and during the early morning
and evening the White-crested Kaleege spent much of their time in scratching among
the leaves of the forest floor for these fuzzy but apparently highly appetizing insects.
Even at midday I have seen a female pheasant thus engaged, but without much success,
while just before dusk I have observed a pair of kaleege rushing here and there after
each turn of a leaf, in pursuit of the moths as they rose in short, irregular flights. Twice
under such conditions, several white-throated laughing thrushes were close at hand,
profiting by the efforts of the pheasants, snatching those insects which flew upward and
beyond the reach of the kaleege. As in so many cases elsewhere, these thrushes were
very frequently the companions of the pheasants when they were on the move, feeding
slowly through open forest, and more than once they spoiled many an hour’s careful
stalking, by spying me out when as yet the pheasants suspected nothing. It tested all
one’s power of self-control at such a time to keep from emptying one’s gun-barrels into
the flock of jeering birds, as they flew close about one’s hiding-place, uttering without
ceasing their creaky and intensely irritating see-e-e-e-e-e-p |! each utterance rising to a
most disagreeable, shrill, rasping climax. |

Whenever I was able to study a single bird or a pair for any length of time, I was
impressed with the extremely regular habits or diurnal round of life which they exhibited.
I became acquainted with two male birds which lived within a few hundred yards
of one another and yet never associated closely. But both fed all day up and down the
opposite slopes of a narrow, forested, dry ravine, and in late afternoon, almost
synchronously, worked slowly upward and, still a hundred yards apart, passed over the
sharp ridge, down to drink at the stream which flowed at the bottom of this small
adjoining valley. They then turned some distance back upon their trail up the slope
again to roost an equal distance apart in some dense-foliaged conifers. By pitching my
observation tent just between the points at which the two birds traversed the saddle of
the ridge, I was able to watch and time each bird. They crossed on the way to drink
and to roost regularly at 4 p.m., varying not ten minutes during seven days, and in the
mornings of at least six days they re-crossed to the daily feeding-ground at 8.10 a.m.,
never varying more than seven minutes from this mean. On two very unusually warm

days the birds delayed their time of passing into the roosting ravine by thirty-five
minutes, the two cocks crossing at the very same minute. This short period of
observation suggests how much could be learned of interest about these and other birds
by a longer period of intensive study, the results of which would be of far greater value
than any amount of casual, half-accurate notes.

I found these pheasants roosting in trees from twenty to thirty feet above the

ground, and have no doubt that many spend the nights on much higher perches. In all
VOL, I D

18 A MONOGRAPH OF THE PHEASANTS

cases the birds were not near the main trunk, but a foot or more out on some horizontal
bough, and in all instances but one the bird was alone.

I have mentioned the laughing thrush as an effective, if wholly unintentional,
benefactor of the White-crested Kaleege. As to enemies, one can certainly include the
pine marten, wild cats and several species of hawk eagles, Wzeraétus and Spizaétus, and,
perhaps worst of all, as far as the despoliation of the nests is concerned, the grey langur
monkeys, which swing through the conifers like a rushing hurricane, leaping and hurling
themselves from tree to tree as recklessly as if the tall, narrow spires of the firs made as
easy travelling as the wide-branched deciduous forests of lower altitudes. I once found
a few bloody tail-feathers of a cock kaleege beside a log with no hint of the murderer—
probably some owl—and twice I found remains of broken egg-shells near nests with the
clearly-marked footprints of the langurs all about.

Man is, undoubtedly, the worst enemy the White-crested Kaleege has at present,
and the bird is becoming scarce in many of its haunts which are accessible to the
multitude of military and civilian sportsmen. The birds learn after a while, however, to
gauge the danger of their opponents, and then, we read, “they are not birds that, as a
rule, afford much sport; you may see a dozen together feeding in the early morning on
one of the ‘perows’ or encamping grounds, in the Siwaliks of the Dhtn, and you may
bag a couple; but even with good dogs to help you, they run so fast and fly so far that
long and weary will be your hunt before you bag a second couple out of that same dozen
after you have once fired. In fact, in such places, unless one has been marked into some
neighbouring tree, when you will generally get a shot, it is best to go on sharp, as a
quarter of a mile further on, on frequented roads like this, you will meet with others
along the track, to which the horse-droppings, containing undigested grain, attract
them. I have in old days shot four or five brace in an hour in the early morning on the
road and ‘perows’ when encamped in the Mohan or Lal Darwaza Pass, through which
runs the main road to Dehra and Mussooree.

‘Generally in the hills you may pick up three or four birds in a day, by beating all
likely-looking patches of cover near fields, but it is rare with this species to make a good
bag. There are, however, places where you may come across the kaleege almost as thick
as pheasants ina Norfolk cover. Such places there used to be close to Bhim Tal and
Naukuchia Tal, small lakes not far from Naini Tal, but at a much lower level, and at the
former of these I once, early in November, killed eleven and a half brace in less than
three hours.” Many more records of a bird shot every eight minutes for three
consecutive hours will leave but few of these unfortunate pheasants for the delight of
posterity.

Speaking of the kaleege in the lower ranges, Wilson says: “In the lower hills, in
the absence of larger game, this bird may serve to while away a few hours of the
sportsman’s time in almost every place where there is wood or jungle; narrow, well-
wooded ravines and thickets of low jungles are the places in which to seek it. A good dog
is essential ; and without one, though a bird may be occasionally picked up, it is hardly
worth while going out. In travelling in the interior a dog used to hill-shooting should
always, if available, be brought ; and with its assistance a few kaleege may be bagged in
some of the coppices and jungle passed through almost every day’s march, till the regions
where larger game is expected are reached.” .

WHITE-CRESTED KALEEGE 19

The method of escape of these birds in the face of attack by sportsmen is varied, and
shows considerable intelligence. When a dog arouses an experienced cock bird, instead
of losing its head and flying off at once, perhaps to within easy range of the gun, it will
sometimes fly up into a tree, uttering its loud clucking all the while. From this point
of vantage it will look about in all directions to see if danger other than the dog
menaces. Often a single glance will reveal the sportsman, and the bird ‘is off like a flash
to safety in the opposite direction.

Again, where the bird is shot at a great deal, it often flies into a tree at the very first
approach of the dog, and there remains, crouched motionless in some fork of a branch
or in a dense bit of foliage, or pressed close to the trunk itself, so that it is impossible to
find it. One may walk around and around, throw stones, halloo at the top of one’s voice,
and when at last the quick eyes of a native gun-bearer detect it, the pheasant seems
simultaneously to sense its discovery, and, like an arrow, it drops from its perch on the
opposite side of the tree and is away off down the valley, silently and on scaling wings,
before a warning shout can be uttered or a gun raised.

In the reserved forests the kaleege are included in the law which is meant to protect
all pheasants during the breeding season, and are supposed to be shot by Europeans only
from September 15 to the end of December. As I have said, the kaleege pheasants often
enter the grain-felds and feed on the corn, and in return they are snared in large
numbers, by baiting with small heaps of grain placed in the fields in which they have
previously been observed.

The White-crested Kaleege breeds throughout its range, from the Terai at a thousand
feet elevation, throughout the outer Siwaliks, and northward almost to the snows. The
highest recorded nest was at a height of nine thousand five hundred feet. At the foot of
the hills and lower valleys the hen pheasant begins to deposit her eggs in April. Higher
up the breeding season does not commence until May, and this is delayed at still higher
altitudes until the middle of June. April 4 and June 20 are the extreme dates at which
fresh eggs appear to have been found.

The kaleege are monogamous, and each pair keeps by itself. The cocks are very
pugnacious, and fight fiercely, especially in the early morning. One writer tells of
shooting a male bird, which lay on the ground fluttering in its death struggles, when a
second cock kaleege dashed out of the jungle near by and attacked the dying bird with
the greatest fury, although the sportsman was standing close at hand.

The drumming with the wings, which I have already mentioned, is heard chiefly in
the early part of the breeding season, and from observation there is no doubt that it is
a warning or challenge to other males within hearing. It has been well compared to the
rapid shaking in the air of a piece of stiff cloth. We have certain proof that it is merely
a defiance, and not a call to attract the females, by observing the result of the native
trapping in Burma with a decoy male of one of the allied pheasants of this genus. In
this case, male after male replies to and rushes at the drumming bird, but no female is
ever thus enticed within reach of the snares.

In regard to this drumming Baldwin says, “We had been sitting motionless for,
I suppose, half-an-hour, when I was startled, all of a sudden, by the loud drumming
noise I have already described, close at hand. The sound came from behind, and on
looking over my shoulder, my companion, with a smile, pointed out the drummer. An

20 A MONOGRAPH OF THE PHEASANTS

old cock kaleege was squatting on the stump of a fallen tree, and with its feathers all ruffled
and tail spread, was causing this extraordinary sound by rapidly beating its wings
against its body.” This latter assertion is an error, corresponding to the long
controversy which waged about the method of production of the drumming of the
American ruffed grouse: whether by beating against a hollow log, or against the
inflated sides of its own body, or merely by the impact of the wings upon the air. The
latter was soon found to be the sole method, and the same is true of the White-crested
Kaleege, and indeed of all the members of this extensive group of Gexnaeus pheasants.

I have seen several of them drumming in a wild state, and all of the more commonly
kept captive species, and the method never varies. The bird stands rather erect, with
head and neck stretched out, and with a single motion raises and half opens its wings,
and imparts to them so rapid a vibration that they seem but a grey haze against the
body. There is never any question of their touching the body—the hazy arc is not of
wide extent—somewhat less, in fact, than the corresponding phenomenon in the ruffed
grouse—and its limits are very distinctly outlined. The sound is produced by the air
rushing through the tensely-strung flight feathers. As I shall have occasion elsewhere
to relate, the drumming sometimes has a remarkable ventriloquial quality, and is most
difficult to locate. The sound is a deep, resonant woof-woof-woof-woof/ sometimes
drawn out into a reverberating, drum-like roll, woof-r-v-v-r-r-r-r-v-woof !—a sound which
is like an electric shock to the lover of the wilds where these birds live. I cannot
believe that the same sudden leap of pulse and thrill of joy at the sound is as keen in
the sportsman, to whom its chief import is the hope of a shot, as when it stands chiefly
for the sheer joy of life in the wilderness and the excitement of close association with all
these splendid wild creatures.

The statement that the drumming is heard only at the breeding season is not quite
true, although there is no doubt that its real function is performed at that period. But
very rarely, even in the autumn or winter, when a cock bird is suspicious of danger and
yet has neither seen nor heard anything definite—at that moment of inexplicable keenness,
when some psychological sense of which we as yet know nothing makes the bird /ee/
that all is not right, the bird will sometimes drum, almost or quite silently. The
feathers are apparently held loosely, their edges pliant or separated, for while the wings
vibrate as rapidly as ever, yet no sound comes forth. I have observed this several times,
and in more than one species. The impulse in such a case is probably purely nervous
and unconscious. The bird is tense, every sense on the alert, knowing that all is not
right, yet with no hint of the character or direction of the danger, and as yet unwilling,
and knowing not in what direction, to flee. Fear not yet predominating, impatience and
suspicion produce an irritated, unsettled state of mind which impels it to resort to the
challenge action, but, tempered with caution, the action alone is manifested, the sound is
repressed. If I have read the bird’s emotions aright—and such explanation seems well
within the mental plane of the pheasant—it shows an interesting correlation of widely
separated seasonal activities with a passing, transient emotion. .

The nest itself, like that of most pheasants, is scarcely worthy of the name, although
in some instances there seems a decided attempt to gather materials. This usually takes
the form of a slight pad or mat of dead leaves, fine grass and moss, and in a depression
in the centre the eggs are laid. In one instance, where the bird had gathered a greater

WHITE-CRESTED KALEEGE 21

quantity of debris than usual, the nest measured nearly a foot across, four inches deep
outside, while the central depression was six inches wide and nearly two inches deep.
When the ground is covered with a thick carpet of dead leaves, no additional matter is
gathered, or again, where a slight hollow, either natural or scratched in the ground by the
birds, is utilized, a mere lining skin of grass or leaves is present. The situation of the
nest is varied, and conforms only to very general requirements. The more usual locations
are as follows:

1.—Placed under the shelter of tufts of large, overhanging ferns ; coarse, tall grass, or
the slender dwarf ringal or hill-bamboo, on rather open hillsides.

2.—Among sapling growths with plenty of underbrush, close under the shelter of an
overhanging stone, thick, low bush or tuft of grass.

One unusual situation is noted by Captain Cock, who says that the eggs are laid, “as
a rule, on the ground under a rock or bush; but I have taken a nest on a large bough of
a tree, in a hollow on the upper side of which the eggs were placed.” There seems
absolutely no significance in this as hinting of a tendency toward arboreal habits like
those of the Tragopans, as the Geznaeus group of pheasants is essentially a terrestrial
one in every way.

The experience of many observers shows that the number of eggs ranges from eight
to fourteen, with the more usual number about nine. The hen incubates for a period of
about twenty-six days, and during this period her dependence upon her leaf-brown
coloration is almost absolute. A dog may approach, and detect her, not by scent, but
by blundering upon the sitting bird, and may seize her before she will leave. Several
writers and sportsmen testify to having approached with great care and actually placed
their hand upon and caught the hen as she sat rigid—held to her home by her mother
love, a performance, let us hope, which was followed by the setting free immediately
of the too-trusting bird. The fact is of great interest, however, as emphasizing the
unconscious faith which the bird places in its hues of brown and buff. The cock bird is
never far away, and when the hen is threatened by a dog or other animal, the male will
fly into a tree and cackle loudly.

At the end of the long period of incubation the chicks emerge, escaping through the
neatly-severed top of the egg, and leave the nest almost at once. When the brood leaves
the vicinity, the male at once attaches himself to his family, and during the period of
growth of the chicks the two parents are invariably found close together with them.
Later, when the wings of the young birds allow them to fly up into the branches, the
entire family roosts together in the same tree, and not until the birds of the year are
full-sized do they drift apart from their parents and begin to forage for themselves. It
is now that the young pheasants of several families seem to feel a drawing toward one
another, and although they usually roost singly, they may often be found feeding
together or working through the undergrowth in company in the daytime.

Unusually late broods which have been observed are unquestionably due to the first
nest having been destroyed by hail or monkeys or some other agency, and not to the fact
that this pheasant ever rears two broods in one season.

The eggs are regular ovals, only moderately elongated, and quite pointed at one end,
bearing in shape a closer resemblance to the eggs of the European partridge and peahen

22 A MONOGRAPH OF THE PHEASANTS

than to those of the common pheasant. The shells are glossy, some specimens more so
than others, and the surface is generally very closely and finely pitted with minute pores,
like those of the peafowl’s egg... Occasionally these will be conspicuous, but usually very
close inspection is necessary to observe them. There is considerable variation in colour,
from pale creamy or buffy white to a warm, rich, reddish buff. They are wholly
unmarked, except that it is not rare to find small dots of white lime occurring irregularly
here and there on the shell. The eggs measure from 46 to 52 mm. in length, and from
31 to 39 in breadth.

CAPTIVITY

As regards the White-crested Kaleege in captivity, the first record of the species
breeding seems to be in the London Gardens in 1859, although a bird deposited eggs in
the same institution two years earlier. The birds lay quite readily, and the chicks are no
more difficult to rear than those of allied species. The adult birds live well, the average
life of twelve individuals being three years, while one individual lived six years and four
months.

DETAILED DESCRIPTION

ApbuLT Mate.—From the occiput springs a long, hairy, greyish-white crest,
sometimes reaching the length of 110 mm. These feathers are normal in character
and dark brown in colour for a short distance at their base—as long as the aftershaft—
and then the white colour appears and the barbs become very scanty and exceedingly
elongated, and for the distal four-fifths no shaft is distinguishable, the plume being
composed of fifteen or twenty filamentous, hair-like barbs.

The remainder of the crown and neck all around, mantle, back, scapulars, wing-
coverts, rump and tail-covets are brownish black, shading into a metallic purple and
steel-blue gloss on the visible parts of the feathers. Entire mantle and scapulars with
each feather terminally edged with whitish. Posteriorly the mantle tends to show an
excess Of the basal dark brown and a diminution of the metallic gloss. Entire back
and rump with a very broad terminal white fringe, averaging 7 mm. in breadth. The
longer upper tail-coverts are very narrowly margined with white.

In typically patterned, fully adult birds the feathers of the wings are almost
unmarked, the invariable exception being the tertiaries, or inner secondaries, which are
conspicuously mottled along the terminal border with white. The green-glossed wing-
coverts are often edged with purplish. The secondaries are glossed only on the outer
web, and the primaries are uniformly dull brown, their coverts being darker. The tail
is black, glossed with purplish blue.

Sides of the face, chin and throat dull brownish black. The narrow glossy band on
the lower throat gives place quite abruptly to the long, greyish-white lanceolate plumage
of the lower parts. The dark brown basal portions are ill-concealed on account of the
narrow, tapering character of the feathers, and the glistening white shaft is conspicuous
even on the pale distal portion. Posteriorly the basal brown increases, until the flanks
and under tail-coverts are quite brownish black, with only faint indications of whitish at
the tips. The under tail-coverts are strongly glossed with greenish.

Bare facial and loral area scarlét, covered with numerous elongate finger-like papillz

WHITE-CRESTED KALEEGE 23

and dotted sparingly with minute degenerate featherlets, these being especially abundant
on the lower eye-lid. Irides warm hazel brown. Mandibles pale greenish horn colour,
darker at the base. Legs and feet rather variable, drab grey to pale whitish, usually
more or less tinged with brown or olive. Weight 2 lbs. to 2 lbs. 6 ozs.

Length, 608 to 735 mm., averaging 670; extent, 730 to 800; bill from nostril,
19 Mm. ; wing, 220 to 250; tail, 250 to 330; tarsus, 75; middle toe and claw, 60; spurs,
straight and rather slender, 15 mm.

VARIATION.— The variation in the plumage of the adult male of the White-crested
Kaleege is of special interest, because of the geographical position of the species. It is
at the extremity of the westward, finger-like extension of the kaleege pheasants along the
southern slopes of the Himalayas. Thus, unlike its allies, which are found in inter-
mediate positions, the White-crested Kaleege can have affiliations in only one direction
and with but a single species, the Nepal kaleege (G. /eucomelanus). While variation is
present and is quite strongly pronounced yet it points to no indication of incomplete
separation or of present crossing between the two species. Again, this variation is as
apparent in specimens from the extreme west as from those of the eastern part of the
range. ‘There is no gradation as the two species approach one another, such as we find
so clearly in the koklass pheasants (Pucrasia) of this same region.

The variation is in part and very subordinately individual when it is persistent
throughout life, but on the whole it is due toage. A fewof the most persistent characters
hark back through all the present kaleege plumages to a more primitive patterning, such
as is to-day found in Burmese and Chinese members of the genus. The whitish
vermiculation of the central tail-feathers is worthy of mention here, as it persists in
very old individuals, strongly developed, not only on the extreme concealed basal
portions, but occasionally well up toward the centre of the feathers, beyond the longest
tail-covert. The other variations are detailed in the description of the first-year male
plumage.

ADULT FremMALe.—The long occipital crest is brownish grey in colour, and less
specialized than in the male. It is shorter, seldom over 70 mm. in length, and the vane,
while very narrow, is normally feather-like throughout. Upper body plumage cold
reddish brown, the feathers of the crown, neck, mantle, back and rump conspicuously
edged with grey. The reddish hue increases in intensity posteriorly, and from the
mantle backward is finely vermiculated with black. On the wing-coverts the margins
are decidedly white, forming distinct tranverse bars across the wing. The flight-feathers
are vermiculated only on the exposed portion of the outer webs, the remainder being
brownish-black.

The middle pair of tail-feathers is black, thickly vermiculated with reddish or rufous,
these wavy lines becoming paler and coarser on the inner webs. Traces of this reddish
vermiculation are found on the external margin and tips of the next two or three pairs,
the lateral rectrices being otherwise black, strongly glossed with green on the outer web.
Chin and throat whitish, shading gradually into the warm reddish brown of the lower
plumage, the black vermiculation being less distinct than on the upper plumage, but
with conspicuous and wide margins of white.

24 A MONOGRAPH OF THE PHEASANTS

Facial skin scarlet, with a scattering of minute black featherlets. Bill yellowish or
greenish horn, darker basally. Irides hazel. Legs and feet brownish, sometimes very
pale. -Claws black. Weight, 1 Ib. 6 ozs. to 2 Ibs. Length, 500 to 730, averaging
600 mm. ; extent, 620 to 680; wing, 225; tail, 210; tarsus, 67; middle toe and claw,
55; bill from nostril, 17 mm. The spurs in the female are sharp, slightly elevated
scalules,

CHICK IN Down.—The crown is “chocolate brown, with the sides of the head and
crown rufous, pale on former, rich and somewhat chestnut on the latter; ear coverts
dark brown; upper plumage brown, minutely freckled with black, each feather with paler
edging, a conspicuous white spot at the tip, and a broad sub-terminal bar of black edged
with rufous ; wing-coverts like the back. Lower plumage dull pale brown, the feathers
with whitish shafts and pale edges.”

JUVENILE PLumMAcE.—The juvenile plumage is of a very indefinite buffy brown,
indistinctly mottled with blackish, darker, more greyish above, paler brown below.
Most of the mantle feathers and all of the wing-coverts have a broad terminal band of
pale brownish white, and a still wider sub-terminal cross bar of black, one of the most
characteristic juvenile marks of this genus. The sexes are distinctly marked in the
juvenile plumage, the males being much darker and the females more of a rufous, reddish
brown.

First YEAR MALe PLuMmaceE.—As in all the members of this genus, the young bird
moults directly from the juvenile garb into the adult plumage. There is, however, much
variation in these first year’s feathers, owing to the advanced or retarded condition of
pigment formation in the blood of each individual. I have known of instances where
the moult resulted in fully adult patterns and coloration, where the crest was almost as
white as it is ever found, and where the wings appeared metallic and unmarked. In two
of these cases the birds were in captivity, well nourished by an abundance of food, but
the moult retarded by unseasonable cold weather. The changes in the blood were
therefore apparently completed, and the transition from immature to fully mature was
clean and abrupt.

On the other hand, in both wild and captive individuals we often find a partial
assumption of adult patterns at this moult, although this is apparent only in more or
less inconspicuous characters—which, however, are full of significance and interest.

The most noticeable of such characters are the absence, or extremely poor develop-
ment, of metallic gloss on the plumage, and the presence of whitish vermiculation on the
wing-coverts, secondaries, upper tail-coverts and tail-feathers. The mantle shows the
absence of gloss more commonly than other parts of the plumage. The entire exposed
outer webs of the greater coverts and secondaries are sometimes thickly vermiculated
with pale brown during the first year of the bird’s life, and the same is true of the three
or four middle pairs of tail-feathers, except that in this case the vermiculations are
greyish white and much coarser. As I have said, this character occasionally persists
even in very old individuals.

The young female in first-year plumage closely resembles the adult. The most

WHITE-CRESTED KALEEGE 25

constant differences are a richer, more rufous red of the plumage in general, thus
approximating the Nepal kaleege, and a greater extent of white in the under plumage.

SYNONYMY

Phastanus hamiltonit Gray, in Griff. ed. Cuv., III. 1829, p. 27; id. Ill. Ind. Zool., I. 1830-32, pl. 41.

Phasianus albocristatus Vigors, Proc. Zool. Soc. London, 1830, p. 9; Vigors, Proc. Zool. Soc. London, 1832,
p. 16; id. Phil. Mag., 1831, p. 60; Gould, Cent. Birds Himal., 1832, pls. 66, 67 (text); Vigne, Proc. Zool. Soc.
London, 1841, p. 6 (Alpine Punjab) ; Stewart, Zoologist (3), X. 1886, p. 321.

Gennaeus albocristatus Wag). Isis, 1832, p. 1228 ; Grant, Cat. Game-birds, XXII. 1893, p. 298 ; Grant, Hand-
book Game-birds, I. 1895, p. 258 ; Nehrkorn, Katalog der Eiersammlung, 1899, p. 193 ; Rothschild, Bull. Brit. Orn.
Club, XIV. 1904, p. 58; Beebe, Zoologica, I. No. 17, 1914, p. 320; Baker, Jour. Bombay Nat. Hist. Soc, XXIII,
1915, p. 666 ; Baker, Jour. Bombay Nat. Hist. Soc., XXV, 1917, p. 164.

Euplocomus leucomelanos Gray, List of Birds, 1844, pt. II]. p. 25 (part).

Euplocomus leucomelas Hodgs. in Gray’s Zool. Misc., 1844, p. 85 (part no. 80 2); id. Icon, ined., in Brit. Mus,
Gall. pl. 14, no. 80 ¢. ;

Euplocomus albocristatus Hutton, Jour. As. Soc. Bengal, XVII. 1847, pt. 2, p. 693; Irby, Ibis, 1861, p. 235
(Kumaon); Adams, Wanderings of Naturalist in India, 1867, p. 92; Blyth, Cat. Mus. As. Soc., 1849, p. 244.

Euplocamus albocristatus Adams, Proc. Zool. Soc. London, 1858, p. 499 (Simla, Cashmere) ; Sclater, List of
Phas., 1863, p. 9; Gray, List Gallinae Brit. Mus., 1867, p. 32; id, Hand-list Birds, II. 1870, p. 260; Elliot, Mon.
Phas, II. 1872, pl. 18 (syn. part); Garrod, Proc. Zool. Soc. London, 1873, pp. 468,640; Hume and Inglis, Stray
Feathers, V. 1877, p.42; Hume and Marshall, Game-birds India, I. 1878, p. 177, pl.; Sclater, List Animals in Garden,
Zool. Soc. London, 1883, p. 487; Marshall, Ibis, 1884, p. 423 (Chamba); Tegetmeier, Pheasants, 1904, p. 227 ;
Oates, ed, Hume’s Nests and Eggs, III. 1890, p. 413; Evans, Ibis, 1891, p. 77; Mitchell, Proc. Zool. Soc. London,
1QII, p. 522.

Gallophasts albocristatus Mitchell, Proc. Zool. Soc. London, 1858, p. 544, pl. 148, fig. 1 and pl. 149, fig. 3;
Adams, Proc. Zool. Soc. London, 1859, p. 186; Jerdon, Birds India, III, 1863, p. 532 (part); Blyth, Ibis, 1867, p.
153; Tytler, Ibis, 1868, p. 203 (Simla to Mussoorie); v. Pelzeln, Ibis, 1868, p. 321 (Koteghur) ; Beavan, Ibis, 1868,
p. 380 (Simla); Brooks, Ibis, 1869, p. 60 (Baini Tal, Almorah); Hume, N. & E. India Birds, 1873, p. 526;
Marshall, Birds’ Nests India, 1877, p. 59; Scully, Stray Feathers, VIII. 1879, p. 346; Leverkihn, Jour. fir Orn.,
1890, p. 195.

Gennaeus albicristatus Blanford, Fauna Brit. India, Birds, 1V.1898, p. 89; Oates, Game-birds India, I. 1898,
p. 324; Sharpe, Hand-list Birds, I. 1899, p. 35; Oates, Cat. Birds’ Eggs Brit. Mus., I. 1901, p. 54; Rattray, Jour.
Bombay Nat. Hist. Soc., XVI. 1905, p. 663 (Nesting at Murree, 6000 feet) ; Ward, Jour. Bombay Nat. Hist. Soc.,
XVII. 1907, p. 244 (absent from Kashmir vale; found in Murree Road and Kishtwar); Magrath, Jour. Bombay
Nat. Hist, Soc., XVIII. 1908, p. 298 (Thandiani, Hazara); Ghigi, Mem. Acad. Bologna (6), V. 1908, p. 145;
“Pine Marten,” Jour. Bombay Nat. Hist. Soc. XIX. 1910, p. 797; Magrath, Jour. Bombay Nat. Hist. Soc., XVIII.
1908, p. 298.

Euphocanus albocristatus Anderson, Jour. Bombay Nat. Hist. Soc., IV. 1889, p. 59.

VOL Il - gE

NEPAL KALEEGE

Gennaeus leucomelanus (Latham)

NAMES.—Specific: Jleucomelanus, from Greek jevxdc white, and pédag black. English: Nepal Kaleege
Pheasant. Native: Kalich or Kalij (Perbuttia), Rechabo (Bhutia), Nepal.

BRIEF DESCRIPTION.—Male: Similar to the white-crested kaleege (G. albocristatus), but with the crest
somewhat shorter and bluish black instead of whitish, and the white fringe on the feathers of the back and rump
narrower. Female: Differs from most females of the white-crested kaleege in being generally darker, especially
below where the ground colour of the plumage is dark reddish brown.

RANGE—Nepal.

GENERAL DISTRIBUTION

ELSEWHERE I have discussed the standing of this bird, and here need only
emphasize the fact that the Nepal Kaleege is a perfectly well-marked species. Although
showing ‘a certain amount of variation, there is no indication of true hybridization with
neighbouring species.

This is emphasized by its range, which is quite distinct. It does not reach the
Arun River, in the eastern part of Nepal, and does not appear to have been taken west of
Jemlah, in the extreme west of the state. In the Dun of south Nepal it is found well
down in suitable localities and northward to about mid-Nepal. Altitudinally it probably
seldom exceeds an elevation of nine thousand feet. Its entire range may thus be
included in a long, narrow rectangle of about three hundred and fifty by fifty miles.

GENERAL ACCOUNT

To the west of Nepal I had watched and studied the white-crested kaleege in the
oak forests of Garhwal ; east of Nepal many days were spent in the moss-hung jungles
where the black-backed kaleege lived, but because my skin was fair, not yellow ; because
my eyes had not that slight Mongolian slant which marks the Ghurka or native
Nepalese, I was never able to see in life the Nepal Kaleege. For this bird keeps well
within the boundaries of this sturdy little kingdom, and no Caucasian is permitted to
enter its maze of tumbled mountain ranges. The one exception is the solitary
Englishman who acts as Government agent at Khatmandu. ~ But even he may not leave
the vicinity of this particular valley. It is upon the specimens and facts obtained by
these agents that we must base our meagre knowledge of this kaleege, which, however,
appears to differ in no particular, in its general mode of life, from its allies to the
west and to the east.

By the courtesy of the late Maharaja of Nepal, I was able to obtain several recently

killed pairs of these birds, which furnished data as to plumage and food.
26

PLATE XXII

NEPAL KALEEGE

Gennaeus leucomelanus ( Latham)

No white man has ever seen this bird in its native haunts, for it is confined to the southern part of Nepal,
where no Caucasian is admitted. It lives in the moss-hung oak and spruce forests which cover the maze of

tumbled mountain ranges of this little kingdom, and is trapped wholesale by the Nepalese shepherds.

Hee STATS

NSOTT DAT |
*QrisiT) GONE Mastied |

Jaqe1to,issq aigdigee.o

orlqerle saslaqoVl ody vd olsastory’ Baqdext ai b

PLATE XXIll.

NEPAL KALEEGE.

NEPAL KALEEGE 27

The crops of two of these pheasants contained the following food :

Male: 3 flat seeds; 1 long stem of a fern ; many small leaves.

Female: 20 termite workers; 1 large black ant; 1 small mollusk; 12 fruits of
Eurya sp.; several unknown seeds and berries ; numbers of leaves.

A female shot on May 31 would shortly have laid at least four eggs.

Scully says that these kaleege are fond of thick forest and are “usually seen in
pairs or in parties of from three to ten, often feeding on the ground near cultivated
patches at the borders of forest.”

“These birds seem very fond of perching on trees, and it is usually in this position
that one comes across them in forcing one’s way through forest which has dense
undergrowth. On such occasions the kaleege first gives notice of its whereabouts by
whirring down with great velocity from its perch, and then running rapidly out of sight
to the shelter of some thicket.- In the winter the birds roost on trees at the foot of the
hills, and the plan for making a bag is to post oneself about sunset under some trees
which they are known to frequent, and await their coming. The birds are then soon
heard threading their way through the jungle towards their favourite trees, and at once
fly up and perch. When once settled for the night in this way they are not easily
alarmed, and I have shot four or five birds in quick succession before the rest of the
party would clear out to quieter quarters. Occasionally, too, one can get a shot at the
kaleege as they cross a hill-path through the forest on their way to or from some stream.

“Great numbers of the Nepal Kaleege are snared and brought into Khatmandu for
sale. The birds bear confinement in the valley very well, and I reared several chicks to
maturity.”

The subject of the validity of the Nepal Kaleege as a distinct species may be
summed up in a few words. More than fifty-five years ago ornithologists were inclined
to admit but a single kaleege. Later, two were allowed, and Blyth added another, but
was convinced that the Nepal bird was a hybrid between the white-crested and
black-backed kaleege.

Now, however, that we have abundant material and more certain knowledge of the
range of these birds, we can say with certainty that there are four well-defined species of
kaleege extending along the southern slopes of the Himalayas and forming a linear
series stretching from Kashmir to Burma, where the genus spreads out into the
wonderfully interesting and puzzling series of wild hybrids marking the transition into
the lineated kaleege and the silver pheasants.

DETAILED DESCRIPTION

ApuLT Mare.—A long, hairy crest, as in the white-crested kaleege. Upper
plumage in general black, with a strong metallic gloss. This is deep steel-blue on
the crown, crest and neck, while the mantle, back and rump show considerable
purple, and the wing-coverts and secondaries are predominately green. The shafts
of the mantle and back are white. Mantle, scapulars, some of the tertiaries and
long upper tail-coverts very slightly fringed with white, usually in the form of a
scanty mottling. This increases on the back and rump, where it forms a solid fringe
of white, about one-half as wide as in G. albocristatus.

28 A MONOGRAPH OF THE PHEASANTS

Tail-feathers black, glossed with purplish green, and with the one or two middle
pairs more or less vermiculated with buff. Chin, throat and ear-coverts dull black.
Under-parts of greyish white; lanceolate feathers usually darker than in a@/bocristatus,
the basal dark brown being much more in evidence. Abdomen and flanks dull
blackish brown, the under tail-coverts glossed with green.

Facial skin scarlet, rather thickly dotted with minute featherlets. Irides dark
hazel. Bill greenish horn colour, dusky at base of culmen and around nostrils. .
Feet brownish grey; spurs dusky. Weight 1 lb. 12 ozs. to 2 lbs. 4 ozs.

Length, 580 to 660 mm.; expanse, 660 to 750; wing, 220 to 235; tail, 240 to
280; tarsus, 72; middle toe and claw, 60; bill from nostril, 18.

ADULT FEMALE.—Rich warm brown, with the feathers tipped with greyish white
on the upper plumage, and more widely with white below. Upper plumage from
the mantle backward, including wings, finely vermiculated with black. The rump and
upper tail-coverts contrast with the central tail-feathers in being of a lighter, more
buffy brown, the vermiculations of the latter being predominately rufous on the
outer, and paler buffy white on the inner webs. Otherwise, as in the female of
albocristatus. Weight 1 lb. 6 ozs. to 1 Ib. 12 ozs.

The colours of the eye and fleshy parts are the same as in albocristatus.
Length, 480 to 510 mm.; expanse, 630 to 680; wing, 210; tail, 200; tarsus, 65; middle
toe and claw, 56; bill from nostril, 17.

The female Nepal Kaleege has, on the whole, the vermiculations of the middle
tail-feathers less coarse than in adlbocristatus, while the grey margins of the upper
plumage are broader than in melanonotus. In the series of perfectly fresh skins of
the three species, /eucomelanus is intermediate in general tone of brown colouring,
_albocristatus being paler and melanonotus darker. Elsewhere I have described how
radically the skins of these birds change in time, so that this distinction seems to
be lost in museum skins.

Cuick IN Down.—The only information we have of a wild caught Kaleege of
this age, is the description given by Dr. Scully of a chick captured in Nepal on
June 10. The wing-feathers were just sprouting, and measured 50 mm. in
length. Head rufous brown, body above dark brown, below buff. Each feather of
the new juvenile scapulars and wing-coverts had a buff tip and a sub-terminal black
bar. Legs and feet orange; bill greenish-yellow horn.

JUVENILE PLUMAGE.—Young birds of three months resemble the female in
general, with the bill pale at the tip, the facial skin pale fleshy red and the legs and
feet pale brown. In this plumage the black sub-terminal bars on the upper plumage
are the most conspicuous character.

First YEAR PLuMAGE.—The adult plumage is assumed at the age of about five
months, but traces of the juvenile brown often remain on many of the new feathers
of the upper plumage and wings.

NEPAL KALEEGE 29

EARLY HISTORY

Under the name of Coloured Pheasant, Latham mentions this bird as early as
1787, and three years later he gives it the title Phastanus leucomelanus. Jerdon
confused it with albocristatus, but its occurrence in ornithological literature has been
rare, and even to-day we know but little of its haunts or habits, owing to the
complete isolation and restriction of its native country of Nepal.

SYNONYMY

Coloured Pheasant Latham, Gen. Syn. Suppl. I. 1787, p. 210; id. Gen. Hist., VIII. 1832, p. 203.

Phasianus leucomelanos Latham, Ind. Orn, II. 1790, p. 633; Bonnat. Tabl. Encycl. Meth. I. 1791, p. 189;
Temn. Pig. et Gall., II. 1813, p. 353, III. 1815, p. 670; Viellot, N. Dict. d Hist. Nat., XI. 1817, p. 40; Vigor, Proc.
Zool. Soc. London, 1832, p. 16.

Nepaul Pheasant Latham, Gen. Hist., VIII. 1823, p. 202.

Euplocomus leucomelanos Gray, List of Birds, Pt. III. 1844, p. 25 (part).

Euplocomus leucomelas Hodgs. in Gray’s Zool. Misc., 1844, p. 85; id. Icon. ined. in Brit. Mus. Gall., pl. 14, fig. 79.

Gallophasis leucomelanos Gray, Genera Birds, III. 1845, p. 498; id. Cat. Hodgs. ed., I. 1846, p. 124; Hutton,
Jour. As. Soc. Bengal, XVII. Pt. 2, 1848, p. 694.

Gallophasis albocristatus Jerdon, Birds India, III. 1863, p. 532 (part).

Euplocomus leucomelanos Gray, List Gallinae Brit. Mus., 1867, p. 33; id. Hand-list Birds, II. 1870, p. 260.

Euplocamus leucomelanos Hume, Stray Feathers, VII. 1878, p. 428; Hume and Marshall, Game-birds India,
I. 1878, p. 185, pl.

Gallophasts leucomelanus Scully, Stray Feathers, VIII. 1879, p. 345 (Nepal).

Gennaeus leucomelanus Grant, Cat. Birds, Brit. Mus., XXII. 1893, p. 300; Grant, Hand-book Game-birds, I.
1895, p. 262; Oates, Man. Game-birds India, 1898, p. 329; Blanford, Fauna Brit. India, Birds, IV. 1898, p. 90;
Donald, Jour. Bombay Nat. Hist. Soc., XII. 1899, p. 577 ?(Kashmir); Sharpe, Hand-list Birds, I. 1899, p. 35;
Beebe, Zoologica, I. No. 17, 1914, p. 320; Baker, Jour. Bombay Nat. Hist. Soc., XXIII. 1915, p. 667; Baker, Jour.
Bombay Nat. Hist. Soc., XXV. 1917, p. 172.

BLACK-BACKED KALEEGE

Gennaeus melanonotus (Blyth)

NAMES.—Specific : melanonotus, from the Greek yuélavo (from Léhac) black, and vé@yvo¢g back. English:
Black-backed, Black-headed or Sikhim Kaleege Pheasant. French: Euplocome melanote. German: Schwarz-
riickenfasan. Native: Karrik-pho (Lepcha) Sikhim; Kirrik (Bhutea) ; Muthoora (Bengali).

BriEF DESCRIPTION.—Male: Resembles the male of the Nepal Kaleege (G. leucomelanus), but with no
white terminal fringe to the feathers of the upper plumage; these feathers are also highly glossed, the wings with
green, the back with steel-blue. Female: Resembles the female of the Nepal Kaleege, but is still darker, the
dark red becoming more of a dark brown or black, and the light tips of the feathers thus appearing clearer
white.

TyPE.—¢ Marked Sikhim, G. B. Mainwaring; 9 and juv. marked Sikhim, A. S. B. (E. Blyth), In the
Indian Museum, Calcutta.

RANGE.—Native and British Sikhim and Western Bhutan.

THE BIRD IN ITS HAUNTS

THE terrible silence of fear closed down upon the jungle. A myriad creatures
breathed, panted or held their breath, while a laughing thrush voiced its terror in a wild
outburst of shrieks, as it fled headlong. For several minutes the moss-hung forest gave
forth not a whisper of life. Only the slow flapping of great butterflies showed that any
living thing still existed within its shadows.

I lay prone upon a huge flat-topped boulder on a bed of damp moss, and here the
laughing thrush had flown down and alighted close to my shoulder. Startled for the
moment, I turned suddenly, with the result I have related, of sending the thrush into a
frenzy of fear and of silencing every creature within earshot.

The Sikhim jungle around me was the home of the Black-backed Kaleege, and from
earlier scouting I knew that at this moment there were several pairs within a few hundred
yards, and a nest of these pheasants within a still shorter distance. The particular day
of which I write was a brilliant one in early spring, but in these mid-Himalayas the air
was cool and, although it was high noon, scarcely a ray of sunlight penetrated to where
I lay upon the floor of the jungle. Even when the alarm of the frightened thrush had
passed, the woods lay quiet, with only the distant sibilant tones of the tiny bush-
warblers.

From base of trunk to topmost twig, every tree was draped with a thick coat and
with pendants of moss—long, streaming tassels of green and brown, which softened every
outline, emphasized every knot> The tiniest two-leaved shoot, just broken from its
acorn, bore its burden of fairy filament, which would increase as the plant grew, asking
no sacrifice of sap or light, but only a support upon which the moss could ripen its lowly
spores and whence it could with wider vantage shed them abroad.

My great boulder jutted out from the jungle floor, lichen-painted and moss-softened

and for ever shadowed by the dense foliage overhead. Before me seven great oaks
30

PLATE XXII

BLACK-BACKED KALEEGE

Gennaeus melanonotus (Blyth)

Deep in the mossy, humid forests of Sikhim I have watched a pair of these pheasants picking among the
fallen leaves, and murmuring to each other in low musical tones. Later the hen made her way to her nest
among the ferns at the base of a great tree, and the cock mounted slowly, branch by branch, to a lofty perch,

and night settled quietly down over the Himalayan wilderness.

oc grioms ‘gnidsiq atasesodg oaody go

ieow t6a'Gd vner tod oboe ee Sek

soreg gol & oF t
pttaiee. pan

Phin
Bln

=

- Bry

ey,
co

PLATE XXIII.

tj
O
ri
[1]
i
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ui
al
ni
Ne
O
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e
Ns
O
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i
a0

BLACK-BACKED KALEEGE 31

encircled a little sombre glade, all leaning slightly toward one another as if met in some
intimate solemn conclave. A checkering of twilight sifted through their green, swaying
curtains, and now and then a shining drop of moisture fell from some moss stalactite,
glistened for a moment as it passed through a ray of light, and silently vanished as it
struck the sponge-like carpet of the glade. Just beneath me this carpet was variegated
with tufts of graceful ferns, while tangled among the moss filaments of the boulder were
masses of acorn conglomerates—a dozen great caps grown together, some still filled with
fruit, others empty—gone to fulfill their destiny, whether it be in the fulness of time to
bring into being an oak to replace these giants, or to be crunched by a passing bear or
swallowed by pheasant or jay. In the interstices of the boulder’s steep sides, clinging to
scanty bits of black mould, stood little jacks, much like the jacks-in-the-pulpit of our
American woodlands, but gay with stripes of maroon and pink.

As I lay among such surroundings, hoping for a view of some of the jungle inmates,
the dim light occasionally grew more dim—still more diffuse. Then there reached my
ears the indistinct murmur of wind through moss, and following a sudden shower of
drops from the saturated foliage, there came through the glade billow after billow of
cloud, faintly veiling the jungle vista with blue. It had come down the valley from the
snows high overhead. From the glaciers it brought a cold, humid chill, but on its way
thither it swept through the higher forests of magnolias, and from the great swaying
blossoms on the mountain sides a mile or more above me, and many miles distant, it
gleaned a burden of perfume, and now the air of the glade about me was saturated with
the sweetness.

As quickly as it came, the chill wind passed, the clouds sifted onward through the
waving moss and the sun shone out, bringing a new wave of warmth from the valley
below which penetrated even to my damp couch.

A long, low mound in the furry carpet marked a tree, fallen years ago, and now,
rotten and giving of its substance to nourish its children sprouting on all sides.
Suddenly above this appeared the small, trim head of a hen pheasant. She reached up
snatched an insect from a twig, scanned the glade for a moment and disappeared. The
ferns closed over her, and once, a few feet farther along the log, a fern trembled for a
moment as she brushed against it. In a few minutes I knew she must be safely
ensconced on the seven eggs which I had already located close to the distant end of the
fallen tree. There for the remainder of the afternoon she sat closely—patiently warming
the chicks into life—her mottled plumage one with the browns of moss and sodden leaf.

A dainty green-backed titmouse flew to a twig on a level with my eye and, filled
with the joy of spring, burst into song. He raised his crest, threw back his head and
shouted cheep-a! cheep-a! cheep-a! cheep-a! again and again. No answer came, but
he did not lose heart. He had another song in his humble repertoire, and suddenly he
changed to a high, metallic Leep-heep ! heep-heep ! heep-heep ! trembling with emotion
the while. Surely if any lady titmouse was within hearing, she could not but be moved !
Far up among the foliage many small birds were twittering, and some distant note
seemed to carry a meaning to him, for after a moment of listening he was off like a shot.

Then two more Himalayan lives touched mine for a moment of time, to diverge
for ever almost at once—and two as different as one could imagine. A yellow-backed
sunbird appeared before me, as suddenly as if from the clear air, as beautiful as if from

32 A MONOGRAPH OF THE PHEASANTS

some unknown fairyland. Perched in a glow of sunlight almost within arm’s reach, the
feathered atom was ablaze with metallic colour—from its beak to toe one’s eye registered
green, maroon, olive, bright yellow, green and black, while its breast was of brightest
lemon stained with crimson. A shuffling among the ferns drew my eyes away and
slowly there lumbered past, seen dimly beyond one side of the glade, a half-grown bear.
The black beast made no commotion, pushing quietly through the under-brush and
soon: passing from view. When I looked back the sunbird, too, had vanished. Although
the bear never crossed my path again, it was far different with the hen Kaleege, as I had
occasion to discover a few days later.

The rest of the afternoon passed quietly, although I once had a thrill when two deer
came close to me and crossed the glade, with many stops for a nibble at fern-top or moss.
One was rufous, one was brown, a sambur doe and nearly grown fawn. Their eyes were
lustrous, and together with their ears and nostrils never for an instant ceased their
vigilant watch for danger. I was hidden and above reach of scent, but not a rustle of
squirrel or bird but was noticed, not a movement of shadow or quiver of mossy bough
but was tested with sight and scent and hearing. Their life seemed one great fear—one
never-ending watch for death. And here was I, the type of the most terrible of all their
enemies, my gun ready, but with my mind far more murderously inclined towards any
of my fellow men who could at such a time have shot them, than toward the wonderful
creatures themselves.

The most conspicuous and jolliest of the lesser tenants of this Kaleege jungle were
golden-winged laughing thrushes, sturdy birds, clad in grey-striped browns, set off with
patches of black, white and greenish gold. They were usually quiet vocally, but for ever
making a great racket among the leaves and debris of the jungle floor. They worked in
pairs, and from the sound of their progress might have been a whole flock or herd of
creatures. Now and then they would leap to a low twig and burst into a rollicking duet
—a sudden, startling mutual guffaw of loud, harsh notes. It seemed as out of place amid
the quiet of these dim aisles as the antics of a clown in a cathedral.

The setting sun found many loop-holes in the canopy of moss, and the glade became
brighter than at midday, the long golden shafts reaching far in through the jungle,
turning the moss to golden lacery and the ferns to yellow-green filigree. New bird
voices came to my ears—two sad half-tones reiterated until they seemed to embody all
the sorrow of the tragedies of the wilderness. The first call of the coming night silenced
the voicing of the day’s sadness; the deep, gruff 400 ! Hoo/ of an owl presaging the still
more terrible intimacy of life and death when the sunshine had gone.

Some time later the whirr of wings drew my attention down vista, and there a pair
of Black-backed Kaleege balanced on a low, swaying branch. For a brief space they
conversed in undertones, each murmuring in the manner so characteristic of all this
group of pheasants. The brown hen almost immediately scaled down to a mossy log,
took as before a single comprehensive look about, and dropped down in the selfsame
place among the ferns, going on to her eggs.

The-cock bird ascended the maple sapling, branch after branch, and then crossed to
an oak and continued to climb his arboreal ladder until he had almost reached the level
of my eye. Walking out on a good-sized branch on the opposite side of the trunk he .
stood for some minutes, looking down, behind, upward, in every direction, murmuring

BLACK-BACKED KALEEGE 33

all the while; his vespers an invocation against the dangers of the night. He then
plucked idly at a strand of swaying moss near him, re-arranged a wing feather and
settled down for the night, with a last, low chuckle—one among a hundred shapeless
bunches of moss. I dimly sighted his scarlet face along the sights of my gun barrel, but
would pull trigger neither for science nor dinner. When the light had grown so dim
that he merged with the moss fantasies about him, I crept backward, slipped down the
moss-deadened surface of the great rock, painfully stretched my stiffened limbs, and made
my way silently toward camp.

A week before I was watching blood partridges feeding amid the snow, with no hint
of spring in the air; a few days later the mating call of the Satyra Tragopan had come
to me amid a glory of rhododendron blossoms; here a jungle home had already been
established, and as I walked toward the glow of the camp-fire, and glanced backward at
the dark, tangled mass of jungle, I knew that I shared a secret with two of its lesser folk,
one sitting close upon her treasures, deep hidden beneath ferns and foliage; the other
overhead, helpless to ward off any serious danger, but drawn by love of mate to perch
close by, where he might at least watch and perhaps warn of impending harm.

Thus passed an unforgettable day in the haunts of the Black-backed Kaleege.

GENERAL DISTRIBUTION

The Black-backed Kaleege has probably the smallest range of any of its group.
It was formerly not uncommon in all suitable localities in Native and British Sikhim,
and in western Bhutan. Exactly how far it extends into the latter country is uncertain,
but at all events it does not pass the Monas River. In the south it is found, at least in
the winter, fairly in the Terai, and from here northward to mid-Native Sikhim, and up
to six and eight thousand feet elevation. :

I was unable to find any trace of this species in extreme eastern Nepal, although
formerly it has been killed on the slopes of Mount Tonglo, on the Singhaleela Range
dividing Nepal from Sikhim.

GENERAL ACCOUNT

The shadows of dense forest growth are the favourite haunts of the Black-backed
Kaleege, and the increased melanism of its plumage, in both male and female, well reflects
its preference for humid, mossy jungles. In ravines, on broad, steep slopes, or on flat
valley terraces it may be found, provided always there is dense cover. The foliage of
tea bushes is said to suit the bird, so that this sort of change from the primeval forest
does not seem to exterminate or drive it away at once, as is the case with almost all
other pheasants.

It thrives as near sea-level as the base of the hills will permit, and is found breeding
in cool oak forests at seven or even eight thousand feet elevation. There is but slight
altitudinal migration, the birds at the higher elevations being forced downward by winter
storms, but those living lower down appearing to be resident throughout the year.

The birds seldom leave the cover of their favourite forest undergrowth, except
occasionally in the morning and evening, when they come into the open or upon a road

to feed. One author speaks of this kaleege as at no time shy, but morning and evening
VOL, II F

34 A MONOGRAPH OF THE PHEASANTS

feeding as fearlessly as domestic fowls, allowing one to approach within a few steps.
This must have been in some arcady where firearms were unknown, for all the kaleege
which I observed were tame only as long as they were absolutely ignorant of one’s
presence. After that they vanished with all the speed which feet and wings could provide.

When undisturbed, these pheasants are never found perching in the day-time, but
feeding or resting on the ground. If approached, but not too suddenly alarmed, their
mode of escape is invariably on foot, running swiftly through the underbrush, making
hardly a sound, and crouching so low that it is only by the merest trembling of low
plants and ferns that their course may be traced. When suddenly flushed on a hillside
they take to wing and scale downwards, or on the level along the hillside forest, skilfully
dodging the tree-trunks, and alighting fifty yards or more away. By circling the spot of
alighting they may often be again flushed. When flushed under similar conditions by
a dog or wild animal, they will usually fly up into a neighbouring tree and survey the
source of danger before leaving the vicinity.

They are usually rather silent birds, although fond of murmuring to each other and
to themselves. Besides this indistinct murmur, which is given with closed beak, the
birds have what seems a contented cluck, a muffled tone, also through the nostrils,
sounding as if coming from deep within the breast, and uttered when they are quietly
feeding or scratching and wholly unsuspicious. This cluck is somewhat more metallic and
sharper than the similar note of a domestic fowl. When suspicion comes and a warning
is intended, a series of these clucks is run rapidly together, ending with a loud, shrill
screech or chirp, uttered with the beak wide open. This implies a strong suspicion of
danger, as I have many times observed, and either dies away, if there is no further
indication of impending trouble, or gives place to the silent cursorial escape, or the
frantic cackling of unrestrained terror accompanying winged flight. When alarmed
and suddenly flushed they utter a variety of notes, a loud, sharp outburst of cries,
a harsh cackle; Aookvi ! kookvi! kookri! being perhaps the one most commonly heard
and the one easiest of phonetic transcribing.

In common with all his allies, the cock drums with his wings, more especially as a
challenge during the breeding season, but also during other seasons of the year. By the
natives this sound is thought to be a certain sign of rain. Its frequency just before the
season of the rains may be due to the synchronous occurrence of the season of courtship,
and in many instances at least, when the cock drums before a rainstorm, it is in direct
response to the thurder. Many times after a roll of thunder I have heard drum after
drum from first one kaleege, then another. Indeed, all pheasants seem to react
nervously at such a time of electric tension, and the species which lack the alar
manifestation of excitement crow or cackle their loudest as the reverberations of
thunder die away. It is almost unnecessary again to refute the many statements
which have been made in regard to the method of drumming of this and other
kaleege pheasants. The bird never strikes its sides, as does the domestic cock, but
drums by quivering its wings through a narrow arc, between its sides and the level
of its back, the sound resulting from the air rushing between the large flight feathers.

There are few edible morsels which come amiss to the Black-backed Kaleege. Its
food is varied in the extreme, and includes fruit, seeds and insects of all kinds. It will
pursue flying grasshoppers, moths, and I have even seen them leap into the air after the

PHOTOGRAVURE 17

Sikhiw-—-IN THE BRAUNTS OF THE BLACK-BACKED KALEEGE

On the sheltered open slopes, tree ferns filter the sunshine through emerald filigree. In the neigh-
bouring forests we find damp moss hanging from bark, twig and leaves, and the hoofs of deer and the
feet of pheasants sink deeply into the soft mould. These kaleege are confined to a tiny bit of the earth’s
surface in Sikhim and Western Bhutan, but dangers are numerous and they are on guard every moment

_of the day, watching for snakes, civets, or even little mild-mannered mountain bears, who know how
delicious a meal these birds furnish.

yao “nt escayy

av TRS hia ih
at Seems (a: conten
¢ ooo ot bniatrsleh eae ss :

PHOTOGRAVURE PLATE |7

L
3)

SIKHIM+- IN THE HAUNTS OF THE BLACK-BACKED KALEEGI

BLACK-BACKED KALEEGE 35

little drab forest butterflies. Especially do they love the grubs and larve of dung insects
and those which live in rotten wood, and they even swallow the small millipede pill-bugs,
armour and all. They will sometimes repair day after day to the same stump or fallen
tree, and labour with beak and claw to penetrate into the tunnels of the hidden insects.
Yams are also a favourite article of diet, and the pheasants will often dig quite deep holes
and trenches in the mould of open forest glades in search of these fleshy tubers. When
they go into the small, field-like places along the edge of the forest, the kaleege find an
abundance of seeds of weeds, grain, and other plants, and, in addition, they eat the
young tops of ferns and nettles, many kinds of berries, and the fruit of a Polygonum and
a raspberry which abound in Sikhim. A female which was brought to me by a native
shikaree had eaten many small bits of green moss, some very small fern-sprouts and a
small turreted snail, while the stomach contained many of the reddish hairs characteristic
of begonia leaves. A male bird shot in the late evening had been feeding exclusively on
insects, all but one of which were moths. Among these was one large one (4rcle
caerulea Guerrée), the remainder being smaller and of as yet undescribed species. With
these was an enormous orange-headed wasp (Vespa magnifica Smith).

The kaleege roosts in trees, and when once found it is easily re-located by the
abundant sign on the ground beneath. It is seldom that a bird roosts alone, two usually
being found sitting close together, while after the breeding season the young roost with
their parents for many weeks. Night after night the birds return to the_identical spot,
until, as certain sportsmen relate with evident pride, the roosting-place is discovered,
when “‘as many as six may be shot from the tree” by the intrepid Nimrod!

Whether this kaleege ever associates with the small deer of the jungle I do not
know. I saw nothing of the kind. It seems well provided with enemies. A few days
after I had watched the nesting pair of birds in their haunts I passed close to the log
where the hen should have been sitting. A glint of white drew my attention, and parting
the ferns I found the seven eggs*clawed out, and fragments of shell and yolk all about,
covered now with a host of hungry ants. The havoc had been wrought the night before,
and a little careful search showed that the self-same baloo was the villain—either the
same bear which had passed me two days before or another of the same size. The
footprints and signs were unmistakable. In his case, at least, there was no malice
aforethought ; he had doubtless been searching for grubs and tubers and berries, and had
ambled aimlessly up to the fallen tree. But the sudden apparition of his great paw,
which he had rested on the soft moss, had sent the mother pheasant in swift, terrified
flight. The sudden roar of wings must have brought memories of other ovivorous days,
and we can imagine how eagerly he sniffed the eggs and clawed them out into a broken
pile, licking the shells and his dripping paws with satisfaction. Thus had seven little
pheasant lives come to nought. Two more days passed and then, for several mornings,
I heard an intermittent drumming from this same patch of forest. It was undoubtedly
the first hint of the founding of a new home.

On the last day which I spent in this glade of the kaleege I had another most vivid
example of the dangers to which these wild creatures are subject. I reached my favourite
boulder at noon and lay for several hours watching the life of the jungle. At first it was
unusually warm and quite breathless—the forest fairly steamed in the unwonted heat.
Then a cool breeze sprang up, followed by a sudden bank of dark clouds well above the

36 A MONOGRAPH OF THE PHEASANTS

valley. From these a terrific burst of hail descended without warning. The foliage and
moss were torn to shreds as by shot. The pain of the impact was so great that I
crouched close between an out-jutting bit of rock and a sloping tree-trunk. Holding my
hand out for a moment, it stung and pained as if lashed by a whip. In a very short
time the pellets of ice were piled up three to five inches, and untold numbers of forest
creatures must have perished miserably. The ferns about the old nest of the kaleege
were lashed flat. Two nests, the one of a warbler and the other a flycatcher’s, which I
had been watching, were literally beaten from their supports and their contents crushed.
Every blossom was in shreds, not a leaf remained whole, and the forest, from the
peace and warmth and life of the full flush of spring, took on the death-like aspect of
winter.

While the storm lasted the cold was intense and the downpour was intermittent ;
first hail in sheets for several minutes, then blue sky and a momentary gleam of the sun
through a rent in the swirling grey clouds; next a fierce downpour of rain, changing
almost at once to the cruel ice. Such storms kill dogs, fowls, geese, cattle and even
men, and the destruction of pheasants and their eggs and young must be enormous.
Immediately after this terrific storm, a pair of dainty verditer flycatchers and a tiny
white butterfly flew across the glade, showing that somewhere there were havens of
safety where even the most delicate craft might weather out the gale. Owls and small
martens are probably the worst enemies of the kaleege among the living inhabitants of
the forests. .

The natives say that the males are very pugnacious and fight fiercely during the
season of courtship. Certain it is that the birds remain together throughout the year,
apparently paired for life, and that the male stays in the immediate vicinity of the nest
during incubation, and later takes his full share in finding food for, and defending
the chicks.

The nesting season varies with the altitude. Lower down, in the foot-hills, the
eggs may be deposited as early as March, while mid-April is the usual season for the
majority of the kaleege. Again, near the upper limits of the pheasants’ range, eggs
have been found as late as the end of July.

No nest worthy of the name is ever built. On the tea plantations the eggs are laid
in clumps of grass at the foot of the tea-bushes, while in the forest itself a hollow is
scratched, or the dead leaves and moss merely pressed down by the weight of the bird’s
body, often under the shelter of a projecting rock, or, as in the case of the nest of which
I have already written, hidden by ferns and close to a great fallen tree.

I have known of five eggs which had been incubated for at least ten days, and which
thus would seem to compose a full set, while ten appears to be the largest set recorded.
Six to eight are by far the more usual numbers. Unusually fortunate covies including
eight young have been observed, but two, three, or at the most four, are the more usual
number of survivors of the nesting perils.

The eggs vary considerably in shape, some being broader, more round, while other
slender ones are more perfect ovals. In tint, too, there is very wide variation, from pale
pinkish creamy, and very pale caf¢ au Jait to a very rich dark coffee colour. The tint
appears to be quite constant in the eggs of the same individual, one pheasant having a
whole nestful of pale eggs, while another will lay eggs invariably of a warm, darkish

BLACK-BACKED KALEEGE 37

brown. The eggs vary from 45 to 50 mm. in length, and from 36 to 39 mm. in breadth,
the average being about 48 by 37 mm.

Formerly the Black-backed Kaleege were abundant, but constant snaring and
shooting in and out of season has sadly depleted their numbers. As early as 1885 it
was said of the kaleege about Darjeeling that while they used to be very common, now
few are seen, and in 1900 the same author (W. P. Masson) adds that, owing to
the forest chowkidars snaring the birds all the year round, the kaleege is now one of the
rarest birds near the hill station. This is true of all the more frequented parts of the
bird’s range, although it is still fairly common in many isolated forests. Its haunts are
all so accessible, however, and the range as a whole is so limited in extent, that the close
season must be enforced rigorously if the species is not to become extinct. The fact that
it thrives on the tea estates is a point in its favour, for the owners can forbid all killing
or snaring on their grounds with much more hope of enforcement than could possibly be
the case with the successful carrying out of any general law.

On the other hand, it is almost impossible to keep the coolies from appropriating
every egg which they find, and so, in spite of protection of the old birds, they are
constantly growing scarcer, and as the majority of these kaleege live in places which will
sooner or later be turned into the lucrative tea estates, there seems little ultimate hope
for the species.

CAPTIVITY

A number of Black-backed Kaleege were among the survivors of the large pheasant
collection which was received in London in July 1857, and they bred in the gardens of
the Zoological Society the following year. Since then this species has been reared in
captivity many times in most of the large public collections and by many private
individuals. Two rather remarkable crosses have been produced of this species with the
Impeyan and with Reeves pheasant.

Its care in captivity differs in no way from that which should be accorded to the
common silver pheasant, of which I have treated in detail as typical of this entire genus.
The period of incubation of the Black-backed Kaleege is about twenty-four days. Of
eight individuals confined in the London Zoo of which a record has been kept, the
average length of life was three years and nine months, while the longest-lived kaleege
survived for a period of thirteen years and five months, this being the record for longevity
for the whole genus.

DETAILED. DESCRIPTION

ApuLT Mare.—Crest shorter than in the white-crested kaleege, but of the same
filamentous, hairy character. Crown, crest and neck all around glossy steel blue.
Mantle and back bluish purple, rump with more greenish reflections, all the feathers
with a convex fringe of deep, rich purple. Wing-coverts and secondaries glossed with
greenish. Primaries dull brown, secondaries dull brownish-black on the concealed
parts of the webs. Upper tail-coverts and tail-feathers metallic purplish-green on the
exposed portions of the webs, dull black elsewhere. [Ear-coverts, chin and throat
dull black, the latter merging almost at once into the lanceolate, almost pure white,

38 A MONOGRAPH OF THE PHEASANTS

under plumage. This is whiter than in either albocristatus or leucomelanus as far
back as the mid-abdomen, where the feathers lose their tapering character. The flanks
and posterior portion of the abdomen are brownish-black. All the feathers of the
lower plumage have white shafts, conspicuous only on the darker feathers. The
under tail-coverts are slightly glossed with greenish.

The facial skin is scarlet, with almost no trace of featherlets except on the fleshy
grey lower eye-lid. In fully adult males the gular wattles are quite well developed.
The mandibles are yellowish, or more usually greenish horn colour, darker at the
base and about the nostrils. The irides are a warm hazel brown. Legs and feet
pale horny brown. Weight, 2 lbs. 6 ozs. to 2 Ibs. 12 ozs. Length, 530 to 630;
expanse, 670 to 730; wing, 235; tail, 265; tarsus, 75; middle toe and claw, 65; bill
from nostril, 19 mm.

In an extreme type of coloration, which, however, is rare, there is no trace of
white on the upper plumage, the purple fringe being immaculate. In the greater
proportion of individuals many of the feathers of the back, rump, inner wing-coverts
and tertiaries have a conspicuous mottled white terminal fringe. Blyth’s type now
in the Indian Museum, Calcutta, is a good example of this. A few of the scapulars
show faint traces of terminal white, increasing on the lesser coverts to a very distinct
band, and reaching its maximum on the middle coverts, where, besides the terminal
band, there is a white mottling for two or three millimetres farther up on the webs.
On most of these coverts the white is so pronounced that it is plainly visible twenty
feet away. On the rump feathers the white is present as a faint, light band, most
strongly developed on the most posterior rump feathers. Now and then a bird will
be found with the mantle tipped with shaded grey as in albocristatus. This appear-
ance of grey or silvery white on the mantle feathers is always correlated with a
change in form as well, the feathers being pointed and narrowed in exact proportion
to the amount of light colour; and truncated and broadened as there is less, or a
total lack of the whitish. These are undoubted indications of close relationship with
the allied kaleege, but I know of no specimen indicating true sub-specific connection
or hybridism, and do not believe that the several species now intergrade or cross
in a wild state.

Rather curiously, the traces of the atavistic light vermiculation on the middle
tail-feathers are found on fewer individuals and, on the whole, less strongly
developed than in albocristatus and Jeucomelanus, in which it is so common a
character in both young and adult male birds.

Aputt FreMmaLre.—Fresh skins of this species show a very decided difference
from the females of the Nepal kaleege, in the very pronounced melanism of the
entire plumage. Whereas the latter would be described as rufescent, me/anonotus is
distinctly black. There is, as in all of these kaleege, considerable variation, but the
lightest and brownest of a large series which I collected in various parts of Sikhim,
are distinguishable at a glance from the fresh Nepal specimens.

The crown and crest are dark brown, while the chin, throat, lower face and
neck all around are grey, paler below, but cold grey on the side and hind neck. The
entire upper plumage in dark specimens is a blackish brown, with well-marked grey

BLACK-BACKED KALEEGE 39

terminal margins, these becoming wide white bands on the wing-coverts, forming
very conspicuous bars. In browner individuals the dorsal plumage is indistinctly
mottled with dark reddish-brown and black, the warmer tone becoming strongest on
the rump. Here too the terminal band loses its pure grey character and becomes
increasingly buffy posteriorly.

In the vermiculation of the middle tail-feathers se/anonotus has the red darker,
more rufous and less buffy than in J/ewcomelanus, and, in the great majority of
individuals, with a decided preponderance of greenish glossed black on the outer
webs, in some specimens the rufous markings being almost obliterated. Then again
in melanonotus the lateral tail-feathers are almost wholly unmarked greenish, lacking
the paler vermiculation which in /eucome/anus often extends over the margins of
several pairs.

The lower plumage of a typical fresh melanonotus female shows a strongly
contrasted black and white coloration, the latter colour forming a very. wide
terminal band and grading into the black through a narrow buffy zone. In very
brown individuals this buffy zone gives place to a more or less extensive area
of rich dark rufous, merging abruptly into the black of the remainder of the
feather.

The facial skin is dark red, dotted with numerous featherlets, and the eye is dark
hazel. The general degree of melanism is reflected very exactly in the pigmentation
of the mandibles. In the lightest, reddest individuals the basal black covers only
about half of the upper mandible, and is broken by two large lateral patches on the
culmen, where the greenish horn predominates. In the dark birds, on the other
hand, the upper mandible is almost or quite black, only the extreme tip showing the
paler hue. The legs and feet are darker than in /eucomelanus. Weight, 1 |b. 14 ozs.
to 2 lbs. 4 ozs. Length, 450 to 530; expanse, 630 to 680; wing, 220; tail, 200;
tarsus, 70; middle toe and claw, 57 mm.

VARIATION.—I wish here to speak of a most interesting and unexpected change
in colour which I have observed in museum skins of female kaleege, an alteration
which is especially noticeable in this very dark-plumaged bird. The extreme is to be
observed in Blyth’s type which is in the Indian Museum, Calcutta. In the catalogue
of the birds of the Asiatic Society this specimen is annotated, ‘‘Mr. Webb, 1845,”
so that we may assume that it is about seventy-five years of age. The difference
between this and the very brownest bird which I could secure in Sikhim is startling.
Although, as far as I could learn, it has never been exposed to the light, yet it is
paler than the palest of any fresh kaleege of any species, while when compared with
a typical black melanonotus female, the two would never be suspected of close
relationship. This is an extreme case, but I have carefully compared my fresh ‘skins
of the three Himalayan kaleege females with those in the museums of London,
Berlin, Paris and elsewhere, and invariably I have found a greater or lesser degree
of paling—an apparent breaking down of the black pigment into a more primitive
reddish brown. Even when this is slight it is sufficient to obscure the distinctness
of the various species, which in comparatively fresh wild birds is quite obvious.

The same is true in a lesser degree of the males, and I have a number of

40 A MONOGRAPH OF THE PHEASANTS

museum specimens which show the browning of all the black, especially of the
primaries and the edges of the contour feathers, and which is due to some factor
other than direct light, although this will bring about the same result more quickly.

Cuick IN Down.—Chicks three or four days old collected in Sikhim May 20,
have the crown, neck and mantle down warm rufous. A black line extends obliquely
downward and backward from the posterior edge of the eye to the neck, widening
somewhat on the ear-coverts and dying out on the hind neck. The remainder of
the upper parts is dark chocolate brown, almost black, with two wide, lateral, pale
buff stripes from the scapulars to the tail down. The face is pale buffy white, and
the under parts creamy white, tinged with pale rufous on the sides.

The bill is pale greenish or yellowish horn, darker at the base and along the
culmen. Irides dark olive-hazel. Legs and feet reddish flesh colour.

Bill from nostril, 6 mm.; length, 90; wing, 45; tarsus, 22; middle toe and claw, 20.

JUVENILE PLUMAGE OF Ma.e.—The sexes are distinguishable in this plumage,
the male being blacker, while the female has more brown or reddish in the feathers.
The male has the upper part of the head, neck and fore-mantle uniform dark smoky
brown, almost unmarked. From the mantle posteriorly over the anterior back and
wings, appears the regular juvenile pattern of a sub-terminal black bar. This is
bordered by faint grey on the mantle and back, and is thus rather inconspicuous,
but on the wing-coverts a whitish terminal shaft-spot is developed, and the black is
bordered proximally and distally by wide reddish-brown bands, which set it off in
sharp contrast. On the greater coverts the terminal white increases. The lower back
and rump are rather indistinctly patterned, a dull buffy mottling near the tip of the
feathers. The flight feathers are dark brown, strongly vermiculated with reddish
brown on the exposed webs.

The rectrices are narrow and pointed; black or slighted glossed with greenish ;
the central pair, and..the next pairs marginally, coarsely vermiculated with buff
and greyish.

The chin and throat are pure white, shading very gradually into the lower
plumage. As we leave the white area a blackish mottling obscures more and more
of the feather, until only a central noded streak remains. This is the pattern of the
side and lower neck. The remainder of the ventral plumage is a confused but
coarse mottling of dark brown and buffy white, the margins of the feathers being
pure white. On the abdomen the disintegrated plumage is wholly white, the flanks
being dark brown. The facial skin is fairly free from the down and is fleshy pink;
the legs and feet fleshy horn colour. :

Bill from nostril, 14 mm.; length, 390; wing, 165; tail, 150; tarsus, 60; middle
toe and claw, 50.

JUVENILE FEMALE.—The rufous down of the head is replaced by a buffy brown,
faintly barred with black. The hind neck is rather grey, with broad black tips, and
on the mantle, reddish brown bands appear, which. frame and emphasize the black,
here sub-terminal, band. On many of the feathers this band is constricted at the

PLATE XXIV

PEUMAGES OF THE BLACK-BACKED KALEEGE

Gennaeus melanonotus (Blyth)

Even when young Black-backed Kaleege have moulted late, the shape, pattern and colour of the adult

plumage are not fully attained until after the first year.

Hines
Fic.

ire

ik

2.

Chick in down four days old, collected in Sikhim, May 2oth.

The sexes are clearly distinguishable in the juvenile plumage, the female showing much
warmer, more buffy tones, especially on the head, back and tail. In this individual, the
down is still present on the face, concealing the reddish skin beneath.

The juvenile male is darker throughout, with narrower white tips to the feathers. The
incoming dark, central tail-feathers are not clear black, but coarsely vermiculated with

grey.

1 16 nields “bas mating sqrt

al
ae ia ois CT Rag a

Ried CE

blo ash Te Se ai Anas
§ aF rts = Le? ; ryEe 7 ag aS
oui gitiwere blampt, on woe

ns ee ofl ab _ odin elisala 918 2

‘ ele Sih das
oald bee sbi gibai ih. Abe: list hae aad bead ae A, wieaR

Ors =;

PLATE XXIV.

PLUMAGES OF THE BLACK-BACKED KALEEGE.

ea Sa
wg

See

BLAGK-BACKED KALEEGE Al

shaft, and with the framing of reddish, presents a decidedly occellated appearance.
On all the wing-coverts the black becomes very broad, distinct and unbroken, and
the feather is terminated by an almost equally wide and very conspicuous band of
pale creamy buff.

The secondaries are uniformly vermiculated with reddish brown and black on
the exposed parts of the webs, while the primaries have only a very narrow edging
of the buff markings. The lower back and rump are almost unmarked. The central
pair of tail-feathers are mottled and vermiculated equally with black and a richer
chestnut than is found elsewhere on the plumage. ‘The lateral rectrices are irregularly
and sparingly mottled with reddish brown.

In a female a week younger than the juvenile male described, the pale facial
down is still present, so the reddish colour of the facial skin of the female at this
age is not so apparent as in the other individual. The chin and throat are pure
white, the ear-coverts and throat band grey. The lower plumage of buffy brown,
indistinctly and faintly mottled with darker, begins abruptly at the grey of the neck,
paling to a buffy white on the belly and abdomen, and becoming an indefinite dark
brown on the sides and flanks. Fleshy tints as in male.

Bill from nostril, 11 mm.; length, 300; wing, 150; tail, 110; tarsus, 40; middle
toe and claw, 35.

Blyth’s type of a quarter-grown female, in the Indian Museum, shows almost as
extreme fading as does the adult female type, being of a pale reddish brown, paler
even than any fresh individual albocristatus which I have seen.

First YEAR PLumMAGE.—In the male of melanonotus, as in the other kaleege,
the bird may or may not bear traces of its immaturity throughout its first year of
life, in the form of greyish vermiculations on the coverts, secondaries and central
tail-feathers. In the juvenile plumage there is no hint of the ultimate lanceolate
form of the breast feathers, and the incoming plumage, if acquired early, is pre-
dominately dark, and not nearly so narrow and tapering as the succeeding plumage
of the second annual moult will be.

EARLY HISTORY

In the ‘General History of Birds,” Dr. John Latham in 1822 described a bird,
calling it the Chittygong pheasant. His summary of knowledge of the bird occupied
two lines: ‘“Inhabits India; is a native of the Chittygong Hills, and known by the
name of Muthurau.” Gray in 1829 calls the bird Phaszanus muthura, while in his
“Genera of Birds” Gray gives the genus as Gallophasis. It is doubtful whether this
species is indicated, especially as it was said to be ‘the size of a turkey.” So the name
melanotus, given by Blyth in 1848 to what was undoubtedly this form, is usually
accepted as the earliest authentic name. It was 1857 before the first living specimens
were received at the London Zoo."

SYNONYMY

? Chittygong Pheasant Latham, Gen. Hist. VIII. 1823, p. 200.
? Phasianus muthura Gray, in Griff. ed. Cuv. III. 1829, p. 27.
? Gallophasis muthura Gray, Gen. B, III. 1845, p. 498.
VOL. II G

42 A MONOGRAPH OF THE PHEASANTS

Euplocomus melanotus (Blyth), Hutton, J. As. Soc. Beng. XVII. Pt. 2, 1848, p. 694; Blyth, Cat. Mus, As.
Soc. 1849, p. 244.

Gallophasis melanotus Hodgson, Icon, ined. in Brit. Mus. Gall. App. pl. No. 937; Gray, Gen. B. III. App.
1849, p. 24; Mitchell, Proc. Zool. Soc. 1858, p. 545, pl. 149, Fig. 2; Sclater, Proc. Zool. Soc. 1859, p. 205 ; Gray, Cat.
Hodgs. ed. 2, 1863, p. 68 ; Jerdon, B. Ind, III. 1863, p. 534; Brehm, Der Zoologische Garten, 1864, p. 197; Hume,
N. and E. Ind. B. 1873, p. 527; Marshall, B. Nest Ind. 1877, p. 59; Masson, Game-birds of Darjeeling, 1885, p. 16.

Euplocamus melanotus Sclater, List of Phas. 1863, p. 9 [Sikhim and Bhotan]; Gray, List Gallinae Brit. Mus.
1867, p. 33; id. Hand-list B. II. 1870, p. 260; Elliott, Mon. Phas. II. 1872, pl. 119; Hume and Inglis, Stray
Feathers, V. 1877, p. 42 ; Sclater, Ibis, 1892, p. 86; Tegetmeier, Pheasants, 1904, p. 227; Mitchell, Proc. Zool. Soc.
London, 1911, p. 522.

Gallophasis melanonotus Beavan, Ibis, 1868, p. 381 (Darjeeling, Interior of Sikhim, 2,000-7,000 feet) ; Bulger,
Ibis, 1869, p. 170 (Darjeeling).

Euplocomus melanonotus Hume and Marshall, Game-birds India, I. 1878, p. 191, pl.

Euplocamus melanonotus Sclater, List Animals in Garden, London Zool. Soc. 1883, p. 486; Evans, Ibis, 1891,
P. 77-

Gennaeus muthura Grant, Cat. Game-birds, XXII. 1893, p. 301; Nehrkorn, Katalog der Eiersammlung, 1899,
p- 193; Rothschild, Bull. Brit. Or. Club. XIV. 1904, p. 58; Ghigi, Mem. Acad. Bologna (6), V. 1908, p. 145.

Gennaeus melanonotus Grant, Hand-book Game-birds, I. 1895, p. 263 ; Blanford, Fauna Brit, India, Birds, IV,
1898, p. 91; Oates, Game-birds India, I. 1898, p. 331; Sharpe, Hand-list Birds, I. 1899, p. 35 ; Oates, Cat. Eggs Brit.
Mus. 1. 1901, p. 54; Beebe, Records Ind. Mus. V. 1910, p. 268 ; Beebe, Zoologica, I., No. 17, 1914, p. 320; Baker,
Jour. Bombay Nat. Hist. Soc. XXIII. 1915, p. 668; Baker, Jour. Bombay Nat. Hist. Soc, XXV. 1917, p. 175.

PLATE XXV

_ BLACK-BREASTED KALEEGE

Gennaeus horsfieldi (Gray)

sts and the bamboo thickets of north-eastern | Burma, I first heard the bubbling
ae f In ae or in small flocks they work ibe: through the ferns and over

A grouse-like whirr

PLATE XXV.

BLACK-BREASTED KALEEGE.

BLACK-BREASTED KALEEGE
Gennaeus horsfieldi (Gray)

NAMES.—Specific: orsfieldi, named for Dr. Thomas Horsfield, an English naturalist, who carried on
scientific work in the East. English: Black-breasted, Purple or Horsfield’s Kaleege. French: Faisan huppifére
de Horsfield ; Houppifére pourpré. German: Stahlblaues Fasanhuhn. Native: Dooreek (Dibrugarh) ; Durug,
Dirrik (Garo Hills); Motoora (Khasi, Sylhet); Muthura (Chittagong).

BRIEF DESCRIPTION.—Male: Entire plumage black, glossed with purplish or steel-blue; the feathers of
the lower back, rump and upper tail-coverts margined with white; the lower plumage only slightly pointed.
Female: Olive-brown ; tip of crest rufous; chin and throat whitish ; wings and ventral plumage tipped with paler
brown ; central tail-feathers deep ferruginous ; in general more rufescent than the other Himalayan Kaleege.

RANGE.—Eastern Bhutan, Assam and Upper Burma.

THE BIRD IN ITS HAUNTS

My first experience with this bird in Burma, east of the Irrawaddy, was most
interesting. We were encamped near the small Shan village of Wau-hsaung, at the
foot of the mountains, which rose range beyond range eastward until they culminated
at the Chinese frontier, forty miles away.

The banks of the Namli River were covered with dense undergrowth, through
which the great spreading horns of many water buffaloes had worn a maze of trails.
The clumps of bamboo had been browzed back until they had taken the form of huge
umbrellas. The rains had not yet ceased, and butterflies in scores of species and
individuals filled the forest and the narrow lanes.

In a slight mist I walked a hundred yards into the undergrowth, bending low to
accommodate myself to the buffalo tunnel trails. Not far from the stream I found
a flock of small birds, several species of babblers, silver-eared mesias, yellow-backed
sunbirds and a single scarlet minivet, flashing like a flame against the dark leaves.
I shot a gorgeous sultan-bird, and was wrapping it up when there came to my ears,
like an electric shock, the inimitable bubbling murmur and cackle of a kaleege
pheasant. I sank down upon the mud, and found that the sound came from the steep,
thicket-covered bank leading down to a slimy backwater of the stream.

The note of these birds is one of the few in nature which sounds distant when
uttered close by, and besides raising and cocking my gun I dared not move. I greatly
desired a specimen, to see if the birds from this comparatively low elevation possessed
any of the remarkable variability of those higher and farther east.

Five minutes passed, and no further sound came from the birds. A family of
babblers passed, fortunately without discovering me; a big green barbet began his
rolling, croaking song overhead, and a pair of pygmy falcons, swallow-like in size
and colour, perched on a slanting bamboo and caught butterflies and grasshoppers.

A flurry of rain and a gust of wind sent down leaves and twigs, and filled the air
43

44 A MONOGRAPH OF THE PHEASANTS

about me with the sweet odour of the large pink blossoms which draped the trees
overhead.

I fixed my eye on a rotten log, and strived at the same time to take note of the
line of underbrush each side. I longed to stretch my 20° of vision to the 85° of
a hare!

Without a note or sound of approach I suddenly saw a crested head shoot up
from behind the log, just along the barrel of the gun, and not six feet from the end.
The patch of scarlet face skin shone for an instant as brilliantly as the flaming ginger
blossom at my feet. I pulled trigger and a cloud of leaves, splinters and earth flew
up and two male pheasants rose, flew ten or fifteen feet and settled again. I lay quietly,
and, to my surprise, the birds came nearer, cackling and gurgling excitedly. Now
they shifted to the right, then to the left, always out of sight down the hill.

I knew the slightest movement of mine would alarm them, so I flattened down
into the spicy forest ooze and waited. They would advance a few yards, cackle, then
give their wing whirr softly and return. Twice when a little distance away they had
encounters—feinting like game-cocks and striking with their spurs. But no real harm
was intended, and the mock battles seemed but the result of their excitement. Finally,
they made a complete circle to higher ground, discovered me and ran rapidly off.

For at least five minutes the two birds had remained within twenty-five feet of
where the shot had been fired. Creeping to the log, I peered over and saw my bird
lying dead a few feet down the slope, killed not by my shot, but by the shower of debris.
It was a Black-breasted Kaleege, typical in every feather. .

GENERAL DISTRIBUTION

The range of the Black-breasted Kaleege covers Eastern Bhutan, Assam and much
of Upper Burma, including Sylhet, Cachar, Hill Tipperah, Chittagong, North Arrakan
and Manipur. To the north it ranges throughout the terai and foot-hills of Bhutan
from 91° E. Long. east to at least beyond Sadiya. To the west it does not reach the
Brahmapootra, although this is probably due only to lack of favourable country. It
touches the coast in Chittagong and Northern Arrakan, extending southward to 20° or
perhaps 19° N. Lat. It thence ranges diagonally north-eastward until it reaches its
most easterly point just east of the Irrawaddy, at about 26° N. Lat. and almost
987 =, sone:

GENERAL ACCOUNT

The Black-breasted Kaleege is one of the most interesting of its group. It is the
first of the kaleege which are strung in a line along the Himalayan terai and foot-hills
from east to west, Zorsfieldi, melanonotus, leucomelanus and albocristatus. The Black-
breasted has a wider range than any of the others, and is notable as being one of the
three Genxnaeus which meet in Burma, hybridize and give rise to the numerous forms
to which names have been given indiscriminately.

Curiously: enough it is the darkest of the allied Himalayan kaleege, while it
touches and crosses with the Silver forms which are the whitest. Its haunts are at -
lower altitudes than the others, and much more humid in character. Its home range,

BLACK-BREASTED KALEEGE 45

like that of most of its genus, is restricted, the individuals having regular habits, and
following much the same route to water and feeding-grounds every day.

During my brief stay I could detect no signs of seasonal migration, and many
observers reported to me that there is none. The birds are so omnivorous that they
are not affected by the local ripening of any certain fruit or berry, and seem to pursue
the even tenor of their life from season to season, merely retiring at the breeding season,
for the purpose of nesting, to deeper, more isolated portions of the jungle.

In Upper Burma I found the Black-breasted Kaleege in bamboo jungle, near the
bottoms of rather steep valleys, and along the trails which wound about the slopes,
never over a thousand feet elevation. In Cachar this species is said to favour low
forests near rivers, while occasionally they venture into rice clearings. In Sylhet they
live on hillocks, overgrown with cane, bamboo and tree jungle, isolated by small
streams, and here in the thick underbrush the birds spend the day. Wherever reported
they seem to delight in coming into roads and trails morning and afternoon to scratch
and feed, keeping keenly on the lookout for danger.

Finding that the Black-backed Kaleege were abundant near Wau-hsaung I
remained there some time studying them, and found their habits to differ in no way
from those more in the centre of distribution, to the westward, in Assam. This region
was only a day’s march east of the Irrawaddy, across from Myitkyina, the terminal
of the railroad in Upper Burma, and about seven hundred and fifty miles north of
Rangoon.

I have already recounted meeting the trio of kaleege, one of which I shot. I had
had abundant opportunity to observe the two remaining kaleege pheasants, and noted
that one was dark, and, like the one I secured, typical Aorsfeldi. The third was
decidedly light, showing distinct white vermiculations on the upper parts and along
the central tail feathers. When at last I secured it, I found that it corresponded most
closely to what has been called odscurus. All were young males, just completing their
moult into fully adult plumage.

These birds proved to be fairly numerous and not difficult to observe. But one
factor made the work exciting—the buffaloes. A half-dozen gigantic, black mothers
roamed about the woods, each having in tow a callow, whitish calf. Huge as they were,
these creatures made but little noise when walking slowly over the mud, and more than
once I had hastily to take to trees. Later in the same day, when I shot my first
kaleege, I was crouched close to where a bird was scratching loudly. I had signalled
to my companion to circle around ahead of me to force the pheasant into view.
Hearing no response I crept back to the lane, and saw him perched some ten feet from
the ground, while an irate water buffalo parent tore up sod and snorted angrily below.
On the other hand, the presence of so many of these creatures made it an easier matter
stalking the pheasants, the birds being accustomed to their movement through the
underbrush. :

Several tall trees, covered with ripened fruit, stood near the river-bank, and all day
were filled with birds, a constant rain of berries and pits falling through the leaves.
In. a few days I counted forty species of birds feeding on this fruit, and early one
morning, when the fog from the mountains was too thick to see through, I heard, as
I walked beneath, the unmistakable note of kaleege. Two birds—male and female—

46 A MONOGRAPH OF THE PHEASANTS

were perched twenty feet up in the tree. They flew as soon as the fog thinned,
and I found by the sign that their occupation of the tree had been for roosting
purposes.

The kaleege, for the most part, roosted at a moderate height in trees at some
distance from the stream. Night after night they would return to the same tree, but if
frightened would resort to some other place. In two such instances, the birds, pre-
sumably the same pair in each case, again returned to the first roost after an interval of
one or two nights.

The kaleege showed little fear of the native Shans, who worked about their homes
in the village, or with huge axes and tiny bows went into the forest for wood. I have
seen a pheasant quietly scratching within ten feet of a trail along which Shans were
passing from time to time, and close to the sound of chopping.

Early morning was the best time to observe the birds. Just when the mists were
beginning to clear away, but long before the sun reached the mountain tops, if one walked
along the trail, the trees and bushes appeared much as near a New England road. Vines
clambered high over the trees, and one saw leaves which simulated wild grape, witch
hazel and alder.

A bulbul would open the chorus with a phrase of clear, sweet notes, and a gang
of laughing thrushes usually followed with a united guffaw, jay-like. Before the full
morning songs of the smaller birds greeted the sun, doves’ voices were supreme—coos,
high and low, from all directions. Then, softly, as from a distance, came a louder, muffled
tone, and the sudden, unexpected break into a cackle which never failed to thrill me. After
a space arose a mellow whirr—the sound of wings in rapid motion, and yet how unlike
the sound of flight! The ear at once detected the difference—one the wing-beat set to
music, the other the more usual hum of utility. The latter marks the acme of evolution
through untold generations of awkward flapping reptiles; the former reaches the ideal—
an approach to our own ability of adapting earthly evolved structures to psychic enjoyments.

The wing-beat of the kaleege during the period of courtship is beyond doubt a
challenge—cocks will respond with foolhardy impetuosity to a poor imitation. But
I have known it to be produced by a hen, and have seen the chicks run to her at the
sound. I have seen it used, as I have narrated, by the survivors in an attempt to
summon a fallen member of the flock. When it is produced in early morning by
yearling birds at the end of the rains, I accredit it without hesitation to the same
outburst of feeling which prompts the bulbul’s song at this non-breeding season—a
symbol of sheer exuberant spirits.

At this time of year (November) the kaleege were in pairs—feeding and roosting
together—or in parties of three to five young birds, the sexes sometimes distinct, although
occasionally, at a distance from Wau-hsaung, I saw them mingling together.

There were one or two tigers in the vicinity of Wau-hsaung, but I suppose the birds
had little to fear from these large carnivora. But some species of small cat fed regularly
on the chocolate squirrels, with a narrow white ventral band, which were so abundant here.
I found their tracks often near the haunts of the pheasants, and once I came across the
remains of a kaleege strewn about a favourite drinking-place of the cats. Twice I heard
pheasants in what must have been sheer extremity—that cry of despair which, even when
it emanates from a domestic fowl, serves to silence every bird within hearing. This cat,

PHOTOGRAVURE 18

WHERE HORSFIELD’S KALEEGE MAKES ITS HOME

Tuersr pheasants wander in families or small companies through the dense jungles, coming now and
then upon beautiful streams, flowing slowly and quietly around the roots of great trees, or pouring in a
headlong cascade over a ledge of rocks. Here the birds slake their thirst and then pass on again into
the forest. I once watched a pair of birds scratching and pecking on a hillside among green begonias
and jack-in-the-pulpits, making the leaves and twigs fly in all directions. The sound could be heard for

many yards, but the vigilance of the birds never relaxed, and at the slightest suspicion of danger they
were off like two meteors.

bas wor ‘Boies ited
ot lageh hes Pee

EN ge

Ma ntie

PHOTOGRAVURE PLATE 18

ss x"),

ee |
YJ L/S Si)

OME.

a[=44
IL

MAKES. ns

?

HORSFIELDS KALEEGE

WHERE

BLACK-BREASTED KALEEGE 47

whatever the species, is doubtless the worst foe of the kaleege. When I saw how it was
possible, by crawling flat upon the ooze, to creep close to a bird intent upon demolishing
a termite nest, I realized how the dread feline—mottled and small—could creep easily
within striking distance. And yet the birds were ever on the alert, the head, with its
brilliant facial skin, shooting up and turning in every direction between each hasty
peck.

Even more than the other kaleege the Black-breasted is voluble, and seldom an hour
passes when it can resist cackling or wing-beating. This habit, and taking into con-
sideration the constant noisy scratching among dead leaves, makes the locating of these
birds a rather easy matter. And yet they are not as sociable as others of the genus, and
six or eight is a very unusual number to find together. Usually a pair is seen alone,
although when following some easy route to water several groups will sometimes drift ©
together and not separate until roosting time.

The food taken varies greatly, not only in different localities, but from week to week
on the same slopes. I have shot one of four birds which had spent the day feeding on
berries, and which had its crops crammed with these alone, and the following day I
secured another member of the flock, disturbing them at a banquet of termites, and
finding the bird’s crop distended with these long-suffering insects. In connection with
this food, R. A. Clark of Cachar says that he once watched an encounter between a
Black-breasted Kaleege and a red junglefowl ‘for the possession of a white-ant hill
from which the winged termites were issuing. I watched the contest for a quarter of an
hour, by which time both birds were exhausted, when the kaleege fled, leaving the jungle
cock in possession. On another occasion I came across a pair of male kaleege fighting
amongst a lot of ferns. They were so taken up with their own affairs that they did not
notice my having approached to within fifteen yards. I let them go on for ten minutes,
and then went up and caught both. They were quite exhausted. The feathers from the
head and neck had all been knocked off, and the latter were bleeding in both birds.”

Several accounts agree in confirming the pugnacious character of these kaleege, and
Inglis says of the species in Cachar that when he once winged a female, ‘“‘she got into
the jungle on the slope of a teelah. It was very steep, and I sent a man to catch the
bird. No sooner had he caught her than the cock came rushing at him. He made a
thrust at it, but couldn't hit it. The bird then retired, but only to renew the attack a
second time. It did this three or four times whilst the man was bringing down its mate.
In the end I shot it also.”

An interesting case of a cock kaleege caring for his brood of newly-hatched birds is
recorded by H. W. A. Watson. He says, “‘I saw no signs of the hen, though I watched
the cock for several minutes. Probably she was absent looking for food. The cock
was very aggressive, and ran around demonstrating, often coming within ten yards of
me. The chicks were hiding in the leaves, one within a few inches of my feet.”

I have seen them scratching about the shallows of small, sluggish jungle streams,
and in more than one bird have found the remains of small crayfish, usually only the
cephalothorax and the abdomen, the birds apparently discarding the less nourishing
limbs and other portions. The crayfish in the gizzard were always accompanied by a
quantity of small quartz pebbles. In Burma a large fleshy fruit is often eaten, pecked
into small pieces before being swallowed, while small black seeds and black ants are

48 A MONOGRAPH OF THE PHEASANTS

two other favourite items of diet in the order of their abundance. Bamboo shoots,
earthworms, small centipedes and moths are also eaten freely.

The breeding season in Upper Burma corresponds to that in other parts of the
range—April and May, with delayed broods extending into June. The nest is a hollow
in the ground, usually at the foot of a tree, the eggs sometimes laid on the moss of the
forest floor or again on a layer of dead leaves. Four to six eggs are laid, but two to three
seems to be the usual number of young reared to maturity. The eggs show considerable
variation, from pale buffy white to a rich café au lait, and even a still warmer, redder tone.
Almost all are speckled with dots of white calcareous matter, which are more conspicuous
on the darker specimens. They are usually a rather broad oval, smooth and somewhat
glossy, and measure from 35 to 38 mm. in breadth, and from 46 to 52 mm. in length, with
an average of 36 by 49 mm.

Horsfield’s Kaleege is not uncommonly exported from India, and breeds readily in
confinement. They live well, and in the case of seven individuals in the London Zoo
the average length of life was almost four years, while one bird lived for nine.

Stuart Baker probably knows the Black-breasted Kaleege better than any living
white man, and also knows how to put his knowledge upon paper, and I cannot refrain
from quoting, from his account in the 1917 volume of the ‘‘ Journal of the Bombay Natural
History Society,” both his description of the haunts of the bird and of his most successful
day’s hunt. ‘The nest is nearly always placed in forest, and the class of forest most
often chosen is the damp evergreen forest met with everywhere along the foothills and
broken ground bordering the higher ranges of the Himalayas. Inside these mighty
forests, composed of an endless variety of trees, mostly tall and mostly covered with a
luxuriant motley of parasites of all kinds, but also with a plentiful undergrowth of canes,
brambles and other plants, the Black-breasted Kaleege has its favourite haunts.
Occasionally in their inner depths one may come across tiny green glades in the general
dense undergrowth. Here the vivid green moss seems even more green than elsewhere,
forming a springy carpet; ferns grow here and there over its surface, and the sun only
comes to it in dappled, quivering patches through the branches high overhead. Such
spots are much beloved by the Kalij Pheasant, and many a time have I come across its
nest in the bushes immediately surrounding them. Comparatively open spots of this
description attract numerous insects, and I am afraid it is these, rather than their special
natural beauty, which induces the pheasants to commence their domestic duties within
easy reach of them. The nest itself is more often than not placed in some tangle of
bushes, briars or canes at the foot of one of the bigger trees, well concealed from
inquisitive friends and enemies, and in some position less moist than its surroundings.
Ravines with mossy, fern-covered sides are often selected, and in such places a rock or
boulder may form its principal shelter. As a work of art the nest is a failure: a heap
of leaves and rubbish scratched into a heap, with a rough depression in the middle for the
eggs, is the limit attained, and Mother Nature herself, and not the birds, is responsible for
all the collecting that has been done. The great buttresses of the cotton-tree (Bombax
malabarica), which project on all sides from the main trunk, form recesses into which
the winds from every quarter blow their quota of fallen leaves and other oddinents, and
thus become splendid places in which birds may lay their eggs, and many a nest have I
seen, both of this pheasant and of other game-birds, in these cosy corners.

BLACK-BREASTED KALEEGE 49

“As far as I remember, six brace of Kalij Pheasant is the biggest bag I have ever
made of these birds, and this was made over practically the same ground as that which
is described in the article on jungle-fowl shooting in the last number of the ‘ Journal.’

“As a rule, when shooting along these mustard fields bordering the Kopali River
one got two to three jungle-fowl to one pheasant, but this time the reverse was the case.

““My companions (on the occasion of which I write) consisted of a Mikir tracker and
a second Mikir with my rifle, and an odd man to work as beater, jungle cutter, luncheon
carrier, etc. This was quite enough for all the beating we should require, for the strips
were mostly narrow bits along the banks, shut in on one side by very heavy evergreen
forest, and on the other by the shingly banks of the stream. Starting just after day-
break, our first 72am, or open bit of cultivation, was occupied only by some jungle-fowl,
who were the first to detect our meeting, and gave no chance of a shot, but as we passed
through the next bit of forest, a hasty but lucky shot right and left brought down two fat,
bronze-backed imperial pigeon, and some quarter of a mile further on we came to a
second mustard field. In this we could see some birds feeding about half-way: down,
though we could not make out exactly what they were in the tall mustard. Leaving my
men under cover, I went forward inside the jungle about twenty yards, and when I
judged I had gone far enough, notified them with a low whistle to come on. Sneaking
along just inside the fringes of forest, I kept about the same distance, or a little more,
ahead of the men, and when, frightened by the latter, the pheasant got up, had an easy
right and left, adding two more pheasants to the bag. My next shot was at a hen
jungle-fowl, as she scurried through the mustard into shelter, just giving me a momen-
tary glance as she left the former. Picking her up, we then went on through a patch of
semi-burnt cane and grass, missing an old cock jungle-fowl as he got up with a cackle
and tremendous flurry on the far side of some charred canes.

‘Two more long snap-shots at running birds are tinkered, and then I have a bit of
luck, for we step out of the jungle into a mustard field, right into a family party of Kalij,
who are scratching round for food just outside the forest. Quite overcome by the sudden-
ness of our appearance, the birds, six in number, take at once to flight, scattering in all
directions, but two drop to my shots, and two others who have gone towards the stream
are turned back by the sight of some men fishing, and wheel around to their original
cover, giving me splendid overhead shots, and both coming down tremendous thumps
in response thereto. One of these, however, is a runner, and for some time evades my
men in their efforts to catch her, until at last a luckily flung stick bowls her over as she
dodges from one bush to another.

“So we wander on, now through a bit of virgin forest, now through a mustard field,
and then through a piece of burnt grass, the black surface showing the pugs of a tiger
who has passed by some two or three hours earlier. We follow them to the stream
where the tiger has had his drink, and then return to the forest, in which we soon loose
all trace of him.

“Changing my rifle once more for the smooth bore, and leaving stripes to rest in
comfort, we turned and worked down the opposite side of the stream towards camp.
The first birds we put up are some jungle-fowl who have been drinking before retiring
into the forest for their midday siesta. As they fly past us I knock the tail feathers out

of the cock leading them with a real bad shot, and bring down a second with a straighter
VOL. II H

50 A MONOGRAPH OF THE PHEASANTS

one. The tailless cock and the others have gone down in the jungle just ahead of us, so,

spreading out, we walk them out toward the next yum, every now and then hearing them

as they hurry forward, rustling over the fallen leaves. Before we get to the edge of the

jungle they have cleared off, without giving another chance, but a single cock Kalij, which

has tarried a little longer than the others, gets up just as we too emerge from the trees,

and is promptly bowled over and gathered. The mustard here is so high—up to our
waists—that we may find some birds lying up in it, and accordingly we work through it

in line, myself on the inside next the forest, and a few yards ahead of the men. Within

the next few minutes two birds run through the mustard, and gain the jungle in front of
me without giving a chance, and then a barking deer jumps up and comes bounding past
me about forty yards off, barking as he starts, and barking again as he gains cover and

stands inside, defying me. He is still barking as we pass where he stands, and I can

hear the stamp of his forefeet, before panic again seizes him and he dashes further away

into the depth of the forest. Nothing more shows up until I have reached the end of the

mustard, and stand on the yard or two of bare ground which separates it from the

nearest trees. As the men come nearer two or three jungle-fowl flit across it, and then

there is a tremendous commotion as nearly a dozen birds, jungle-fowl and pheasants

mixed together, rise into the air. A hasty shot at one of the former not only knocks it

over, but also accounts for an unlucky hen pheasant which has come into the line of fire,

and a second shot brings to bag another hen pheasant, which falls, a cloud of flying

feathers, with a bang right on to the man with my luncheon-basket. After he has righted

himself and collected my scattered provisions, we proceed on our way, and by noon, when

we stop to have a rest and lunch, I have managed to bag twelve pheasants, six jungle-

fowl, three imperial pigeon, and a couple of very evil-smelling white-eyed pochard, which, -
however, are not disdained by the coolies.”

DETAILED DESCRIPTION

ApuLt MaLe.—The plumage in general is black, strongly glossed with purplish
blue, especially on the upper surface, the posterior ventral plumage being almost dead
black without the metallic gloss. The only decided colour character is found on the
feathers of the back, rump and shorter upper tail-coverts, which have a very conspicuous
white fringe of a normal width of about 5 mm. Throughout most of the western part
of the range this varies but little, but to the eastward in many individuals we find a
tendency toward the disintegration of this fringe. It either becomes narrower, or else the
basal part is broken by dark mottling. Correlated with this there is often an extension
of white as a very narrow terminal border over the inner, median wing-coverts and
tertiaries and the longest tail-coverts. About fifty per cent. of the individuals have the
extreme hidden bases of the median tail-feathers obscurely but thickly vermiculated with
dirty white. This is almost invariably the case with birds from northern Burma east of
the Irrawaddy, where the vermiculation often extends an inch or two beyond the visible
portion of the feathers. The variations occurring along the geographical junction with
nycthemerus and lineatus are multitudinous, and often asymmetrical, reflecting the
widespread hybridization with these species.

The crest is long, rather thin and partly disintegrated. The primaries are brown ;

BLACK-BREASTED KALEEGE 51

the secondaries and tail-feathers black, glossed with blue only along the visible margins.
The bill is usually greenish horn colour, paler toward the tip and darker basally; facial
skin scarlet ; irides brownish hazel ; legs and feet variable, sometimes leaden blue, again
light horn colour, brown or fleshy. Length about 580 mm. ; bill from nostril 18; wing
221; tail 230; tarsus 75; middle toe and claw 60; spurs about 22 mm. The tail is the
most variable of all these measurements, averaging longer in eastern birds. Weight
from 2% to 34 lbs.

ApvuL_tT Femare.—As the females of albocristatus are paler of hue, and those of
melanonotus more melanistic, so the females of Zorsfeldi tend toward erythryism, being
on the whole more rufescent than the others, but great variation exists between fully
adult birds shot in the same district.

In general the colour is a rich olive-brown ; tip of the crest rich rufescent ; sides of
the head and neck greyish toward the tips; chin and throat whitish; body and wings
decidedly rufescent ; wing-coverts and ventral plumage tipped with paler colour, some-
times very conspicuously ; ventral surface with brownish-white shaft-streaks ; rump and
upper tail-coverts paler olive brown; centre tail-feathers and often the longest tail-
coverts deep ferrugineous, unmarked, or finely vermiculated with dusky. Primaries
brown, darker on inner webs, tinged with olive on the outer webs; secondaries and
tertiaries much more rufescent, either plain or finely speckled with paler toward the tips.

Facial skin crimson; irides hazel; bill greenish, darker at base; legs and feet
variable as in the male. Length, 530 mm.; bill from nostril, 18; wing, 210; tail, 210.
tarsus, 72; middle toe and claw, 55 mm.

YounG Matrr.—Much like the female, sometimes indistinguishable, sometimes
darker. The succeeding moults gradually eliminate the brown and replace it with
black pigment, and a bird of the second year may be fully adult in appearance or
with every feather stained with immaturity.

EARLY HISTORY

Latham introduces us to this kaleege in 1823, under the name of the Sylhet
Pheasant. His only remarks are, ‘‘ Inhabits India, brought from Sylhet, in the Province
of Bengal, by Sir J. Anstruther.” Gray, six years later, repeated the description and
gave to the bird the name of /athami. As his description is not exact, he doubtless
had in his possession one of the wild hybrids. G. R. Gray named it Gallophasis
horsfieldi in 1845, and gave an excellent plate of the cock. The first live birds were
placed on exhibition twelve years later.

SYNONYMY

Sylhet Pheasant Latham, Gen. Hist. VIII. 1823, p. 208.

?Phasianus lathami Gray in Griff. ed. Cuv., III. 1829, p. 26,

Gallophasis horsfieldi Gray, Gen. Birds, III. 1845, p. 498, pl. CXXVI; Mitchell, Proc. Zool. Soc. London,
1858, p. 544, pl. 148, fig. 2, pl. 149, fig. 1; Sclater, Proc. Zool. Soc. London, 1859, p. 205 [24 days incubation] ;
Sclater and Wolf, Zool. Sket. 2, 1861, pl. 39.

Euplocomus horsfieldi Blyth, Cat. Mus. As. Soc., 1849, p. 244; Hume, Stray Feathers, VII. 1878, p. 429.

52 A MONOGRAPH OF THE PHEASANTS

Euplocamus horsfieldi Sclater, List of Pheas. 1863, p. 9 [Assam, Sylhet]; Gray, List Gallinae Brit. Mus., 1867,
p. 33; Gray, Hand-list Birds, II. 1870, p. 260; Elliot, Mon. Phas. II. 1872, pl. 20; Hume and Inglis, Stray
Feathers, V. 1877, p. 42 [Cachar]; Hume and Marshall, Game-birds India, I. 1878, p. 198, pl.; Fasson, Stray
Feathers, 1X. 1880, pp. 203, 205 [Chittagong Dist.]; Sclater, List Animals in Gardens Zool. Soc., 1883, p. 486 ;
-Hume, Stray Feathers, XI. 1888, p. 303 [N. Arrakan, Manipur, Assam, Sylhet and Cachar]; Oates, ed. Hume’s
Nests and Eggs, III. 1890, p. 416; Mitchell, Proc. Zool. Soc. London, 1911, p. 522.

Euplocomus cuviert Hume and Marshall, Game-birds India, I. 1878, pl. only.

Gennaeus horsfieldt Grant, Cat. Game-birds Brit. Mus., XXII. 1893, p. 302; Grant, Hand-book Game-birds,
I. 1895, p. 269; Oates, Game-birds India, I. 1898, p. 334; Blanford, Fauna Brit. India, Birds, [V. 1898, p. 92;
Sharpe, Hand-list Birds, I. 1899, p. 35 ; Nehrkorn, Katalog der Eiersammlung, 1899, p. 193; Inglis, Jour. Bombay
Nat. Hist. Soc., XII. 1899, p. 676 (Hylakandy Dist., Cachar); Baker, Jour. Bombay Nat. Hist. Soc., XII. 1899,
p. 487 (No. Cachar); Oates, Cat. Birds’ Eggs Brit. Mus., I. 1901, p. 55; Oates, Ibis, 1903, p. 102 (Bhutan and
Assam to Chittagong, Manipur and Upper Burma); Rothschild, Bull. Brit. Orn. Club, XIV. 1904, p. 58 (hybrids) ;
Baker, Jour. Bombay Nat. Hist. Soc., XVII. 1907, p.9g71 (Khasia Hills) ; Ghigi, Mem. Acad. Bologna (6), V. 1908,
p. 144; Harrington, Jour. Bombay Nat. Hist. Soc., XIX. 1909, p. 309 (Bhamo Dist.); Finn, Game-birds India
and Asia, 1911, p. 71; Beebe, Zoologica, I. No. 17, 1914, p. 320; Watson, Jour. Bombay Nat. Hist. Soc., XXIII.

1914, p. 159 (male caring for young); Baker, Jour. Bombay Nat. Hist. Soc., XIII. 1915, p. 669; Stevens, Jour.
Bombay Nat. Hist. Soc., XIII. 1915, p. 723.

Euplocamus horsfieldi Tegetmeier, Pheasants, 1904, p. 227.
Gennaeus prendergasti Oates, Jour. Bombay Nat. Hist. Soc, XVII. 1906, p. 10; Ghigi, Mem. Acad. Bologna
(6), V. 1908, p. 144.

Gennaeus batemani Oates, Jour. Bombay Nat. Hist. Soc., XVII. 1906, p. 11; Ghigi, Mem, Acad. Bologna (6),

V. 1908, p. 145; Harrington, Jour. Bombay Nat. Hist. Soc., XX. 1910, p. 327.
Gennaeus mearst Oates, Ann. Mag. N. H. (8), V. 1910, p. 164.

Gennaeus horsfieldt horsfieldi Baker, Jour. Bombay Nat. Hist. Soc., XXV. 1917, p. 181 (Description, habits).

PLATE XXVI

LINEATED KALEEGE

Gennaeus lineatus (Vigors)

‘epitome af the sone life of the Lineated, as well.as of most other pheasants.

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PLATE XXVI.

LINEATED KALEEGE.

LINEATED KALEEGE

Gennaeus lineatus (Vigors)

NAMES.—Generic: Gennaeus, from the Greek yevvaroc, noble or proud, probably with reference to the bird’s
carriage. Specific: Lzmeatus, Latin, striped or lined. English: Lineated or Vermicellated Kaleege. French:
Faisan de Regnant; Houppifere rayé. German: Strichelfasan; Graufasan. Vernacular: Yit, Kayit, Kanayit
(Burmese) ; Sinlouk (Talaing); Poogik, Phooyk (Karen).

BRIEF DESCRIPTION.—Male: Upper plumage vermiculated finely on exposed parts, more coarsely on
basal areas, with black and white in equal amounts; long crest and under parts black, sides of breast and abdomen
with white shaft-stripes ; central tail-feathers creamy-white on inner web and at tip. Female: Olive-brown above
with pointed white shaft-stripes on the neck and mantle, usually split basally into a V-shape, sometimes simple
lines ; secondaries mottled with buff on outer webs; chin and throat greyish white; under plumage chestnut ; all
the feathers with white shaft-stripes, and dark cross-bars on the abdomen; middle tail-feathers buff, mottled with
black on outer web; others barred with black, white and chestnut.

RANGE.—Valley of the Irrawaddy in Central and Lower Burma, eastward to the Shan States and
north-east Siam, and south some distance into Tenasserim.

GENERAL DISTRIBUTION

Tue centre of distribution of the Lineated Kaleege is central and southern Burma,
especially east of the Irrawaddy. While its northern boundaries fade gradually into the
maze of hybrids, apparently typical specimens have been taken as far north as 24° north
latitude. From here southward the Irrawaddy may in general be taken to mark its
limits, although many birds have been shot on the western slopes of the river valley,
especially from 21° southward. At 18° three typical males have been secured on the
western slopes of the Arakan mountains. On the east it ranges well into the Southern
Shan States, probably touching the range of xycthemerus at many points, and has been
recorded east of the Salween at about 20° north latitude. How far into Siam it ranges
we do not know, but at least as far as 12°. To the south we may include all of the
Pegu east of the Irrawaddy and Tenasserim as far as Tavoy in 14° north latitude.

Within this area, often at the higher altitudes, averaging about two or three
thousand feet, appear occasional individuals of what has been called sharpez, a form
which I cannot even tentatively admit as a species.

GENERAL ACCOUNT

Like most of the kaleege pheasants, the Lineated is not a bird of widely ranging
habits. When it finds satisfactory haunts, with food, water, good roosting trees and
freedom from molestation, it will spend its entire time within a remarkably limited
area, disappearing into the deeper, more isolated parts of the jungle only at the breeding
season. Also, like its congeners, it prefers mountainous or at least hilly country to flat
plains or grassy prairies, although it lives at moderate elevations and never ascends to

the heights that the silver pheasant or some of the Himalayan kaleege attain. Dry,
53

54 A MONOGRAPH OF THE PHEASANTS

well-drained terrains, coupled with a fairly cool climate, are ideal conditions for the
Lineated Kaleege, cpen forest in the neighbourhood of water affording them all that any
pheasant could desire.

This bird has been recorded from an elevation of over four thousand feet and from
sea-level, but these are above and below its usual haunts, and the great majority of the
species doubtless spend their lives between a thousand and twenty-five hundred feet.

Leaving the fast-drying plain of the Irrawaddy, with its cactus and water-lilies, the
latter striving to ripen their seeds before their lagoons disappear, the former storing
every drop they can absorb against the coming dry season, we approach the purple out-
lying hills of the Shan States. -These rise abruptly from the plain, usually as a small,
transverse spur jutting out from the regular north and south range. One passes many
small rivulets and waterfalls pouring down to the plain.

Even at a thousand feet the flora has completely changed. At thirty-six hundred
feet (Maymo) a dense growth of small oaks and chestnuts alternates with the curious
“Nelly” fruit-tree, larch-like at a distance, and what appear to be familiar hickory-nut
trees. Pines scent the air, and occasionally we come upon a whole forest of them.
Blackberries are ripening at the end of the rains, and now the Lineated Pheasants come
occasionally into the less-frequented lanes and roads, picking the berries from the bushes
or scratching up the mould at their base for grubs. This is always in the early morning,
and close approach would not be difficult were it not for the blossom-headed parrakeets
which fly off screeching, spying one from afar and warning every pheasant to take to
shelter.

Lineated Kaleege are not gregarious, even to the extent of the white-crested species,
but, on the other hand, they are by no means solitary birds. At times other than the
breeding season isolated individuals, especially old cocks, may be found feeding and
roosting alone, and hens may occasionally appear to be living by themselves. But the
family ties and the lure of certain routes to water are two factors which serve to bind
together, however loosely, the birds of related blood or of a limited neighbourhood.
When concentrating on these birds, one may come across as many as fifteen in a day’s
search, but this is unusual. As in the case of most of the other kaleege the family ties
are strong, and the usual parties of three to seven are often composed of parents with the
brood of the year. |

A reliable index of the amount of sociability among most birds is the rare or more
frequent use of the voice, and with this in mind it is surprising that kaleege are not
more gregarious than they are. They may truly be called voluble, and seem to enjoy
exercising their vocal organs at all times of the day. 1 have heard them thus conversing
or calling, both on their perches before descending and after retiring, on low branches
when resting in the heat of the day, when feeding, and when on their way to water.
The loud call, which is also the note of suspicion and even alarm, is the familiar, long-
drawn-out, liquid gurgle, preceded by several guttural or mumbled sounds. When
carefully stalking birds, and this loud, startling cry comes from the underbrush appar-
ently at one’s elbow, the effect is always that of an electric shock, and the one note is
almost as difficult to vocalize as the other. Major Nisbett has sent me the following as
approximating his idea of it—

Wurk-urk-urk-uk-si—te! urk-urk-uk-si—te !

‘LINEATED KALEEGE BE

This is the utmost that our alphabet will achieve, and if one has listened to a captive
kaleege of any species, he will instantly appreciate the attempt.

When the pheasants are suddenly alarmed so that they take to wing, their outcry is
a shrill squeaking, much like rubbing a wet cork on a bottle. This is more recognizable
as the syllables Whe-ech-cheech-che-ech! .

The most interesting phase of their volubility is when they are feeding quietly
together or preening their feathers. On several occasions when I had cached myself
with such care that my presence was wholly unknown to the birds, I was able to listen
to the intimate conversation of a family. Even my coarse hearing could discover the
modulations and inflexions, but to no human ear is given understanding or appreciation
of the extent of the ideas or sensations thus conveyed. I remember one bird, busily
scratching and feeding, which appeared to be talking wholly to itself. No other was
within several yards. As it scratched the mould and leaves it was silent, then, when it
began to search for the edible results of its energy, it murmured continuously in a low
undertone, which rose and fell, gained in strength and timbre, or actually broke into
short, isolated grunts, according to the excitement of the discovery and chase of the
insects and other creatures upon which it was feeding.

While the Lineated is not a stronger flier than others of its family, it rises easily,
and is able to continue for a sufficient distance to carry it well beyond ordinary danger.
When flushed by the sudden rush of a dog, or an undetected approach of a beater, the
bird will shoot across a valley with strong, rapid wing-beats. Whenever possible it
directs its flight downhill and scales as much as the force of gravitation will permit.
The tail is usually held low, and only when suspicious of danger does the bird draw
itself up to full height on its very toes, and search every direction.

DAILY ROUND OF LIFE

A night spent in slumber on the branch of a tree; an early morning drink and
search for food; a midday siesta; again a feeding period, and in the cool of early
evening a leisurely stroll to water, and thence to roost—such is an epitome of the daily
life of the Lineated, as well as of most other pheasants.

These birds are omnivorous, and they seem to thrive equally well on a wholly
vegetable diet, or one which embraces only insect and other animal food. ‘They are
inveterate scratchers, and to listen for the loud rustling of leaves is one of the commonest
modes of detecting their presence. When a family of pheasants is accustomed to haunt
any definite portion of jungle, daily inspection of this area will reveal the considerable
amount of energy expended in this search for food. If this investigation be carried on
at midday, when the birds are resting quietly, there will be little chance of disturbing
them or driving them elsewhere. They seem to have favourite spots in the open jungle
floor, and day after day will enlarge the borders of this scratched-over area, which is
doubtless chosen because of the abundance of some particular food. Another habit,
which I have both observed and have had reported to me, is that of scratching around
stumps or the trunks of fallen decayed trees, and often the bark itself will be detached in
large flakes, probably by strenuous pecks of the bird’s bill.

Nisbett and Payne report that the birds fatten very noticeably when they take to
feeding on certain foods, such as when there is an unusually good crop of a small species

56 A MONOGRAPH OF THE PHEASANTS

of chestnut. Among other vegetable items found in their crops are small, tuber-like
roots and jungle yam. If these are small they are swallowed whole; if of larger size
they are pecked to pieces, a habit which is very common among the argus pheasants.
Besides these the Lineated Kaleege eats jungle berries and seeds, fruits, flower petals,
fern fronds, young leaves, grass and bamboo sprouts. Nisbett says that he has never
found any grain in their crops, although he has known them to feed on the edges of fields
containing rice, maize and millet. As to this habit of approaching human habitations
or areas of cultivation, the habits of the Lineated seem to differ in various localities.
Some observers report it as among the shyest of birds, never being found within several
miles of any native hut or village, while elsewhere it seems to spend much of its time
feeding near fields. It is generally agreed that it does not object to unfrequented trails,
and a half-dozen sportsmen have spoken of finding the best shooting while walking
along these jungle paths. The insects taken most frequently are the non-flying, jungle-
inhabiting forms, such as caterpillars, grubs, and of course the ubiquitous termites or
white ants.

The pheasants roost in small trees and on slanting bamboos, usually selecting
places which would be difficult of access to civet cats and arboreal carnivores. The
finding of regular roosting-places proved to be only a little more difficult than the
locating of birds’ nests in general. The search was made possible by the conspicuous
sign, the good-sized piles of which often indicated the long-continued use of one
particular spot on a certain branch. In the vicinity, sometimes in the same tree, were
usually the roosts of other birds, two or three close together, or singly. In these
separate instances the number was too great for a single family, indicating that the
fragile flocking instinct which drew the birds together on their search for water, was
maintained throughout the hours of sleep. The roosts were almost always on rather
slender bamboos or tenuous branches, perhaps guarding against the unheralded approach
of a carnivore by the vibration of the thin branch.

Personally I failed to observe any convoying of Lineated Pheasants by jay-thrushes
or other birds, but both Payne and Phillips have reported this to me as a common
association, a fact of no unusual moment when we remember the universality of its
occurrence among other species of the genus. As to enemies, I can present no direct
data, but it is safe to mention wild cats, civets and martens as in the front rank.
Their method of escape from danger varies with the character of the attack. A sudden
alarm by dog or man will send the birds up and away at once, flying above the trees if
out over a valley, or low and swiftly between the trunks in the jungle itself. When
prepared for the appearance of a dog, the pheasants fly at once into the nearest tree, and
sit there cackling softly, watching their disturber until he goes out of sight, or ready to
fly at once at the approach of his master. In any event the birds are exceedingly
difficult of observation by direct approach, but extremely easy if one conceals oneself in
the line of their advance down a hillside or nulla.

HOME LIFE

Although within the twenty-five hundred feet of elevation, and the ten degrees
of latitude of distribution the Lineated Kaleege encounters many changes of climate,

ae ee URE

HAUNTS AND BREEDING-GROUND OF THE LINEATED KALEEGE

In central Burma, east of the Irrawaddy we find a dense growth of small oaks, chestnuts and pines.
Blackberries ripen at the end of the rains, and the Lineated Pheasants come into the grassy fields for
berries and grubs. In the early morning it would not be difficult to approach closely, were it not for the
flocks of blossom-headed parrakeets which fly off screeching, spying one from afar and warning every

creature within hearing.

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LINEATED KALEEFGE 57

the breeding season varies but little. The average is about the end of the first week
in April, while two weeks in each direction from this date will cover the majority
of the more advanced and delayed nesters. March 16, as recorded by Bingham, is
early, while an August record doubtless reflects the unseasonable efforts of a bird whose
first nest had been destroyed. I have no faith in the occasional assertion that this
Species rears two broods.

The loud call, while heard throughout the year, is more frequently uttered
before and during the breeding season, and very probably plays a part, however
subordinate, in challenge and courtship. The wing music is of much greater
importance, and is pre-eminent as a challenge. It is a very simple matter to lure
cocks within sight, by even a rather crude imitation. The usual native custom is
rapidly to twist a small stick, with a bit of stiff leaf or cloth inserted at the top.
While, as I have already said, this wing-beating seems to express many emotions,
that of arousing jealousy of a rival is the dominant one.

When about to give this instrumental challenge the bird stands very erect, with
tail rather depressed, and, instantly spreading wide its wings, rapidly whirrs them
back and forth throughout a rather small arc. It is a very restricted segment of a
circle when compared with the similar, but greatly exaggerated effort resulting in the
drumming of our ruffed grouse. The wing-whirr of the kaleege—and this may be
taken as true of all the species—is markedly ventriloquial. When it seems low and
subdued it yet has a penetrating quality which will carry it far through jungle and
underbrush.

I have heard it’ both subdued and again loud and reverberating, but never
to the extent described by Colonel Tickell, who says, “The noise in question is
the most extraordinary and the most unnatural—that is to say, the most unbird-
like, I have ever heard. I was one day, in the cold season of 1859-60, looking out
for a rhinoceros in the hills which skirt the eastern limits of the Tenasserim pro-
vinces. Some very recent marks of the animal were pointed out to me by my Karen
guides, and, following the traces through the jungle down the hillside, I was at last
brought up by a profound ravine. While some of my party left me to reach the
bottom of this dell by a more circuitous and practicable route, and I remained perched
on a steep declivity, a singular reverberating sound reached my ears, proceeding
apparently from the deep valley below me. It was a tremulous, subdued noise, as
if the mountains were shuddering in an ague fit, and I, who was thinking of
nothing but rhinoceroses at the time, and had made up my mind to see a host of
them emerge from the dense jungle as the result of so strange a symphony, was
utterly amazed by my Karen companions telling me the noise was made by the
‘Yits’ (hill pheasants). I could not help smiling at such a_ singularly literal
illustration of the fabled mountain in labour with the xascitur vidiculus mus
enacted by these funny birds. I have only on that occasion heard this extraordinary
sound, though for weeks at a time journeying and living in forests abounding in
hill pheasants.”

The tremulous quality identifies it at once as the challenge of a kaleege, but it
would require a vivid imagination to picture a rhinoceros as its author. While the

kaleege as a whole are pugnacious, yet there seems no reason to think that the
VOL, II i

58 A MONOGRAPH OF THE PHEASANTS

Lineated excels in this respect. In regard to captive birds I have found that it is
wholly a matter of individuality. Some males will not tolerate a female of their
own kind at any time of year, while others show but little objection to the
presence of other cocks even at the breeding season. No one seems to have wit-
nessed a battle between wild birds, but sorely wounded specimens have been found
under circumstances which left no doubt of the vadson ad’étre. As the result of
widespread inquiries among sportsmen I could only elicit the information that the
fighting took place chiefly in March and early April, and in morning and late
afternoon. In addition to the whirring of the wings, Major Nisbett tells me that
the birds make a clapping noise by striking the backs of the wings together.
This performance, so characteristic of fowls and pigeons, I have never heard from
any kaleege, and it certainly must be but rarely produced.

There can be little doubt that the Lineated is normally monogamous. The
frequent association of the birds in isolated pairs is strong circumstantial evidence,
and the final proof is the not infrequent record of both parents seen with a single
brood. On the other hand, if females are numerous, a cock is not unwilling to
consort with two or three.

I did not find the nest of this kaleege, and my information is derived from
the data of Majors Nisbett, Harington and other excellent observers. Nests have
been found throughout the entire extent of elevation from sea-level up to over
two thousand feet. The location is typically phasianine—on the ground in the forest.
It may be in a clump of dense bamboo or lower undergrowth, close beside a dead, fallen
tree or at the base of a live one. The slight hollow may be a natural one, in which
case the natural forest debris of dead leaves, grass or moss is left and gradually felted
down into a dense lining by the weight of the bird’s body, or it may be scratched —
out by the hen bird, when the eggs are deposited on the earth itself. Six to eight
are the more usual numbers. In the Petchaburi River country, south-west of Bangkok,
Siam, Gairdner found a nest of eight eggs on April 3. Judging by the largest
number of young seen together, five seems to be the largest brood reared to
maturity, while two or three is the usual brood. The eggs are regular ovals, some-
times quite broad, and with considerable gloss. They vary from rich reddish-buff
to buffy cream, and the pores are always lighter and often pure white. They measure
from 44 to 50mm. in length, and from 35 to 38 in breadth; the average being 47
by 36 mm.

The period of incubation, judging by captive birds, is twenty-five days. The
chicks are strong and run at once. Oates says that he was ‘fortunate enough to
capture portions of four broods. It is astonishing in what a short time the little
birds make themselves invisible. It is difficult to secure more than two out of one
batch. It is a case of pouncing on them at once or losing them. The mother is
a great coward, running away at the slightest alarm, and thus contrasting very
unfavourably with the jungle fowl, which keeps running round and round the intruder
with great anxiety till her young ones are in safety.”

This latter statement, I think, is based on an individual performance, as my
experience with closely related species of kaleege seems to indicate that the hens
are quite as brave and as good mothers as those of any of the family. The young

LINEATED KALEEGE 59

appear to remain with the parents throughout the autumn and winter, wandering
off by themselves only at the approach of the next breeding season. The annual
moult occurs about October or November, and while the young birds may breed in
the second year, it is sometimes the fourth annual moult before all signs of
juvenility are eliminated from the plumage.

RELATION TO MAN

Although partial to lonely trails which pass through the jungle, and occasionally
feeding along the borders of cleared fields, yet Lineated Pheasants do not like the
haunts of mankind, and prefer the undisturbed quiet of isolated regions. They will
not, however, desert a place at the first appearance of civilization, and may be shot
for a year or two at comparatively short distances from recently-built rest-houses.
They remain even longer in the vicinity of a new native hamlet, where, in the end,
the trapping of the birds results in an even more thorough elimination.

Kaleege are taken both in spring-traps and by means of decoy birds.. These
are surrounded by scores of nooses, and the approaching cocks seldom avoid
entangling themselves before they reach their supposed rival. I have already spoken
of the decoying of wild birds by means of artificially imitating the wing-whirr.
Besides the bamboo-stick-and-leaf method, this is achieved by means of a bit of
cloth held in the two hands and repeatedly snapped taut. Sometimes two methods
are combined, as when the hidden native whirls his stick and thus stimulates
the tethered tame bird to wing-whirring, which sound, in turn, attracts the wild
cocks.

English sportsmen consider the kaleege as legitimate game, and many are shot
both in season and out. The latter reprehensible deed carries its own punishment,
for very often the birds are exceedingly lean and tough. In the autumn and winter,
after a diet of acorns, they leave little to be desired as an addition to the camp
mess,

Accounts differ widely as to the behaviour of the kaleege when hunted, some
sportsmen saying it always escapes by running, others that it flies at once. In
underbrush, when approached by a man, it certainly always escapes on foot, but the
approach of a dog in almost any kind of cover is sufficient to send it up at once
into a tree.

While the general range of the Lineated Kaleege is being continually restricted
by wholesale shooting and trapping, yet it is protected in all Government reserved
forests between March 1 and October 1, and even outside there are many officers and
other sportsmen who are as conscientious as if they were on preserves at home.

Full-blooded Lineated Kaleege are rarely to be procured alive outside of Burma,
the birds generally showing atypical characters, being captured probably on the
boundary of the range of the species, where hybridism has taken place. The first
were exhibited in London in 1864. Of nineteen individuals of which records were
kept, the average duration of life was a year and ten months, while the extreme
was six years and four months. Lineated Kaleege breed readily, and, of course,
cross with any species of the genus.

60 A MONOGRAPH OF THE PHEASANTS

DETAILED DESCRIPTION

ApuLT MALe.—Top of the head and elongated, stiffened crest black, glossed
on the exposed portions of the feathers with metallic purplish blue. Contour feathers
of the upper plumage finely vermiculated with black and white in equal proportions,
so finely on the exposed portions that the general appearance is of a uniform grey.
The areas thus characterized are the back and sides of the neck, mantle, back,
rump, shorter upper tail-coverts, scapulars, lesser and medium wing-coverts. The
alternate lines extend obliquely across the vanes. Basally they become coarser, and
the black increases until it is dominant. Primaries and their coverts blackish brown,
with irregular, confused white vermiculation. Secondaries black, with narrow but
stronger white lines, those on the outer web often split longitudinally. Tertiaries
gradually merge into the greyish vermiculation of the back. |

Tail of sixteen feathers, moderately long, curved and compressed. Central pair
pale buffy-white, except for the basal three-fourths of the outer web, which is
vermiculated with fine black lines. On the succeeding feathers this black increases,
until about the third pair it becomes dominant, especially on the outer web. On the
outer, shorter rectrices the white is reduced to irregular coarse vermiculations on
both webs. ‘The tips of the second and third pairs remain clear buffy white, however.

Chin, throat, lower neck and remaining under-plumage black with a faint
bluish gloss. Sides of the neck and breast with wide shaft-stripes, confined almost
altogether to the outer webs of the feathers. Mandibles greenish horn colour,
irregularly blackening toward the base. Nasal cere greyish. Entire facial skin bare,
and brilliant scarlet in colour, covered with numerous finger-like papillae. At the
breeding season this brilliant ornament is swollen and extended into three well-
marked wattles or projections, one superior and anterior to the eyes, a second at
the extreme posterior portion, and the third gular, hanging down below the gape.
Lower eyelid pale bluish. Iris hazel-red in fully adult birds at the breeding
season. Legs and feet usually flesh-colour, spurs darker. Weight, 2$ to 3 lbs.
Length, 630 to 740 mm.; bill from nostril, 23; wing, 250; tail, 290; tarsus, 85;
middle toe and claw, 65; spurs, 15 mm.

VARIATIONS.—The commonest variation among fully adult birds from well within
the limits of distribution, is in the amount of striping on the under surface.
From being confined to the sides of the neck and breast, this may extend over all
the ventral plumage. The markings of the central tail-feathers are never exactly
alike in any two birds. In fully adult males from localities in the very centre of
distribution I have seen the entire webs covered with the markings, while from a
near-by locality the birds would show the more normal, clear, whitish inner web
and tips. When we approach the boundaries we at once begin to see the effects
of hybridization—with horsfeldi on the north and west, and xzycthemerus on the
east, and the variations as a consequence become innumerable.

Aputt FEMALE.—Top of head and long crest rufous brown, mottled with black ;
remainder of the upper plumage olive-brown, very faintly mottled with black ; all of
the feathers of the upper and side neck and mantle with a conspicuous white shaft-

LINEATED KALEEGE 61

stripe, slightly bordered with black; many of the stripes are split with brown basally ;
inner webs of the primaries and secondaries dark brown; outer webs of the primaries
olive-brown, and those of the secondaries mottled, and with coarse, broken vermicu-
lations of paler buff along the margins.

Middle tail-feathers yellowish-buff; on the inner webs this colour is clear, with
sparse basal dotting of dark brown; the outer webs are brokenly vermiculated with
dark brown, basally these marks becoming heavy, oblique lines of chestnut, bordered
with black; the second pair shows this latter pattern intensified, while in the next
four pairs the buffy background clears to white, and the equality of pigmentation
gives the effect of successive oblique bars of chestnut and white, the former bordered
strongly with black; the outermost, shortest rectrices are dominately chestnut, with
irregular mottling of dark brown on both webs.

Chin, throat and upper neck greyish white; remainder of under-parts brownish-
chestnut, deepest on the lower breast; all of the feathers with a narrow shaft-stripe
of white, usually bordered with black, and with the feathers, especially of the posterior
portions, more or less distinctly cross-barred with black.

Mandibles horny brown, with a varying amount of black pigment, especially
toward the base; facial skin scarlet, covered rather sparsely with fleshy papillae and
a very few minute degenerate feathers, and not developed into wattles; irides hazel-
brown; feet and legs flesh-colour or brownish; spurs often present, but very small.
Weight, 2 to 2} lbs. Length, 550 mm.; bill from nostril, 20; wing, 225; tail, 225;
tarsus, 75; middle toe and claw, 55; spurs never more than 3 mm., and usually a
flat scalule.

Cuick IN Down.—Pale rufous on the forehead, warmer on the crown, with a
reddish-chestnut post-orbital line. Reddish-brown above, with two lighter, lateral
lines, paling into buffy-white below, the breast being warm rufous.

IMMATURE Matr.—Like female, changing slowly to the adult male plumage,
which it attains usually in the third year.

EARLY HISTORY

In 1823 Latham gave a good account of this kaleege, which he called the
Lineated Pheasant. He says the description was given him by Dr. Buchanan, and
was taken from a bird in an aviary in India. Its home was uncertain. No further
reference is found until 1831. Vigors than called the bird Phascanus lineatus, having
in hand a specimen sent from the Straits of Malacca. Lesson, in the same year,
re-described it under the name Faisan de Reynaut (Phastzanus Reynaudit), and in
his sole comment, “du Pegu,” gave us the first correct information as to its home.

Bélanger, in his ‘“‘ Voyages aux Indes Orientales,’ in 1834, gave two plates of the
Lineated Kaleege.

SYNONYMY

Lineated Pheasant Latham, Gen. Hist., VIII. 1823, p. 201.

Phasianus lineatus Vigors, Phil, Mag., 1831, p. 147; Vigors, Proc. Zool, Soc., London, 1831, p. 24; Jardine
and Selborne, III. Orn., 1836, n.s. pl. 12.

62 A MONOGRAPH OF THE. PHEASANTS

Phasianus reynaudit Lesson, Traite d’Orn., 1831, p. 495; Lesson, in Belang. Voy. Ind. Orient, 1834, p. 276,
pls. 8, 9 (Pegu).

Gennaeus lineatus Wagler, Isis, 1832, p. 1228 ; Gould, Proc. Zool. Soc., London, 1859, p. 150 (Tavoy); Oates,
Stray Feathers, V. 1887, p. 164; Anderson, Anat. and Zool. Researches, I. 1878, p. 669; Grant, Cat. Birds Brit.
Mus., XXII. 1893, p. 304; Grant, Hand-book Game-birds, I. 1895, p. 272; Blanford, Fauna British India, Birds,
IV. 1898, p. 92; Oates, Game-birds India, IV. 1898, p. 351; Sharpe, Hand-list Birds, I. 1899, p. 36; Nehrkorn,
Kat. der Eiersammlung, 1899, p. 193; Oates, Cat. Eggs Brit. Mus., I. rgot, p. 55, pl. VI., fig. 5; Rippon, Ibis,
1901, p. 556 (Low Elevations of Southern Shan States); Oates, Ibis, 1903, p. 100; Ghigi, Mem. Acad. Bologna,
6 (V.), 1908, p. 140; Evans, Jour. Bombay Nat. Hist. Soc., XVI. 1905, p. 520; Finn, Game-birds India and Asia,
1911, p. 73; Wall, Jour. Bombay Nat. Hist. Soc.,, XXI. 1912, p. 460; Hopwood, Jour. Bombay Nat. Hist. Soc.,
XXI. 1912, p. 1215; Beebe, Zoologica, I. No. 17, 1914, p. 320; Gardner, Jour. Nat. Hist. Soc. Siam, I. 1914,
p. 40; Baker, Jour. Bombay Nat. Hist. Soc., XXIII. 1915, p. 675.

Phasianus fasciatus McClell, Calcutta Jour. Nat. Hist., 1842, p. 146, pl. III. (Arrakan ?).

Gallophasis lineatus Gray, Gen. Birds, III. 1845, p, 498; Hume, Stray Feathers, II. 1874, p. 482,
(Tenasserim).

Gallophasis fasciatus Gray, Gen. Birds, III. 1845, p. 498.

Euplocomus lineatus Blyth, Cat. Mus. As. Soc., 1849, p. 244 (part); Gray, Hand-List Birds, II. 1870,
p. 260.

Euplocamus lineatus Sclater and Wolf, Zool. Sket. 2, 1861, pl. 38; Sclater, List of Phas., 1863, p. 8, pl. 9
(Tenasserim and Pegu); Gray, List Gallinae Brit. Mus., 1867, p. 34; Elliot, Mon. Phas., II. 1872, pl. 23; Hume,
N. and E. India B., 1873, p. 525; Gould, Birds Asia, VII. 1875, pl. 14; Hume, Stray Feathers, III. 1875, p. 165,
(Pegu Hills, east of the Irrawaddy R.); Fielden, Stray Feathers, III. 1875, p. 168; Hume and Marshall; Game-
birds India, I. 1878, p. 205, pl.; Anderson, Anat. and Zool. Researches, I. 1878, p. 669 (West of Irrawaddy
above Mandalay); Hume and Davison, Stray Feathers, VI. 1878, p. 436 (Tenasserim); Bingham, Stray
Feathers, IX. 1880, p. 195 (Thoungyeen jungles); Oates, Stray Feathers, X. 1882, p. 236 (Pegu); Oates, Birds
Burmah, II. 1883, p. 316 (East slope Arrakan Hills (?), Siam, Irrawaddy Valley to Bhamo); Oates, ed. Hume’s
Nest and Eggs, III. 1890, p. 416; Mitchell, Proc. Zool. Soc., London, 1911, p. 522.

Lophophorus cuviert Hume, Stray Feathers, III. 1875, p. 166.

Nycthemerus lineatus Blyth and Walden, Cat. Mamm. and B. Burma, 1875, p. 149 (Karen Hills).

Gennaeus lineatus lineatus Gyldenstolpe, Swedish Exped. Siam, 1916, p. 157; Baker, Jour. Bombay Nat.
Hist. Soc., XXV. 1918, p. 326.

- CHINESE SILVER KALEEGE

Gennaeus nycthemerus (Linnaeus)

the beautiful bird of “day and night,” ranges across the whole of south
or er almost to the sea-coast of Fokien. It is found in open forest, on clear,

‘rivers. While the Silver Kaleege is common in captivity, yet no white na has ever seen

chinese have developed a taste for their flesh and trap and shoot the bird extensi

ma

adi. mo} 2 ined to vlodw adtzeoi8) eager” givin.

bar ehid Sonia Ucols ag 2916) maqo ai Pruiol 2 aT

stilw oa tay: ,yiivisyso Cai. Hommosndi lags,

4 : P $ope cele. Sag Dae pan, Bits, DB,
ioft sof ballid yRomiot sow <isdmwn evoariond

fioiz3. bud si joode bas gsi bas deaf tis 102 stent s

PLATE XXVII.

CHINESE SILVER KALEEGE.

i”

CHINESE SILVER KALEEGE

Gennaeus nycthemerus (Linnaeus)

NAMES.—Specific : mycthemerus, from Greek yvy), stem of vvé, night and #yéoa day, an apt simile in the
case of its jet black and pure white plumage. English: Silver Pheasant; Chinese Silver Kaleege ; French:
Faisan argenté ; German: Silberfasan; Vernacular: Ing-ky (Silver fowl) ; Paé-ky (White fowl) Chinese.

BRIEF DESCRIPTION.—Male: Top of the head, long, hairy crest and under parts black, glossed with
purple; upper parts white, the feathers peppered, or on the wings lined, with four to six black, converging
lines ; tail very long, centre feathers white, others with numerous oblique lines. Female: Crest blackish-brown;
throat brownish-white; entire body plumage and central tail-feathers olive-brown, finely mottled with dusky
lines ; outer tail-feathers black, with irregular oblique white and olive lines.

RANGE.—South China, from central Yunnan east to Fokien.

PLE SBIR TIN TiS WikDs HOME

Ir was the ides of March among the tumbled mountain ranges of Fokien. A
week’s rain had freshened the landscape and swollen each bud and lichen, and under
the heat of the midday sun the myriad lives in egg and chrysalid were stirring rest-
lessly. When I left the river bank in early morning the wind blew cold, but after a
climb of a thousand feet the sun’s warmth seemed that of midsummer. Although
frost was almost unknown, yet the air was filled with springy odours, and to the
eye Nature revealed herself as awakening after a long winter's sleep.

The hills, ravines and mountains surrounded us, their steep, rounded outlines broken
here and there by weathered cliffs of blackish rock. A soft coat of green covered
the slopes, but nowhere were the dark spires of forest. Cultivation must be all but
impossible here, but the hordes of yellow men of this and countless past generations
had gleaned again and again, and left only sprouting pines and brushy bamboo. Grass
was almost absent, its place taken everywhere by a low, coarse fern brake, covering
densely every available inch from water’s edge to mountain top.

I crouched and watched the south side of an open valley. Along the bottom
flowed a quiet green stream; here and there were stubble-filled rice fieldlets, most
of them of only a few yards extent.

Two beautiful shrikes clung to a reed near by, and now and then flew into a
clump of brush, searching for insect food like vireos. They were at last driven away
by a flock of black mynas, which whirled down to the fields and began to feed.

Now an interesting sequence of happenings delighted me. A flock of a dozen
magpies was searching busily through the ferns on an overhanging bank. Occasionally
one flew down and grubbed out a hollow with its beak among the newly-ploughed
clods of a small rice terrace below. It dropped something into the hole and carefully
covered it up before flying up again. After two or three magpies had repeated this,

a hen Silver Kaleege walked slowly out into the field, followed by another hen and a
63

64 A MONOGRAPH OF THE PHEASANTS

cock, and all scratched vigorously where the magpies had been delving. Good-sized
black objects were found and beaten vigorously before being swallowed.

For half an hour this went on, the magpies continuing to bring down objects
and bury them; and the kaleege to dig them up and devour them. Neither species
paid any attention to the other. While this was going on, a creamy white, weasel-like
animal appeared, winding along below the bank. A bit of earth dislodged by a
magpie fell almost on the creature, and it turned and dashed back to shelter.

In order to solve the problem of the magpies and pheasants I crept nearer, but
missed in a try for one of the latter. With the second barrel I secured a magpie
which had just started to fly down to the field with one of its treasures in its beak.
I ran to the fallen bird and found a partly-crushed cock-chafer, or Jung-bug, lying
close to it. In the mouth of the bird was a leg of the beetle, and in its crop eight
more of the insects. The magpies had been feasting to repletion on the newly-emerged
beetles and then, for reasons best known to themselves, had chosen to work hard at
burying all the others they could find. The pheasants had come into the field to
scratch for food and, having uncovered several of the newly-buried toothsome morsels
in the tracks of the magpies, had seemed to realize that these birds were in some way
connected with this manna, and for the last ten minutes hardly had the magpie started
back for the bank when one of the pheasants unearthed its cache.

GENERAL DISTRIBUTION

Except on the extremes of its range, little is known as to the distribution of the
Chinese Silver Pheasant. Throughout Fokien it is common except immediately on
the coast. There are definite records for north-west, central and south-east Fokien.

As to the remaining territory in the eastern and central parts of its range, Robert
Swinhoe has given us the only definite information. He says, “It is found in the
wooded mountains of the following provinces: Fokien, Canton, Kwangse and
Kweichou.” This brings us to the Yunnan border, -where I have evidence of its
occurrence in numbers. It is certainly found in central, and more rarely in western
Yunnan, and also in the eastern part of the Southern Shan States. This is fifteen
hundred miles to the westward of Fokien. It must occur along the south China coast,
for in the Island of Hainan we find a closely related form. Elsewhere we may trace
its range by the line of hybrids which have resulted from its crossing with horsjieldi
and /ézeafus. In Yunnan it grades westwardly into darker forms, while in Burma,
the Shan States and Annam, we find such scattered hybrids as the so-called vujifes,
andersont and bel, all of which show an unmistakable preponderance of xycthemerus
blood.

GENERAL ACCOUNT

Like the golden pheasant, the Chinese Silver Kaleege is one of the best known
of its group in captivity. The two are still further alike in their isolation in a wild
state. While we may purchase a pair of Silvers for a very small sum and be quite
certain of rearing numerous young birds to maturity, there is no authentic record of
the finding of a nest of the wild birds, and in fact, only the most meagre notes even
of distribution.

PHOTOGRAVURE 2

Chatbhoh “HAUNTS. OF THE SILVER: PHEASANT

In Fokien, near the borders of Kiangse, the pheasants of ‘“day-and-night” live among wild ravines
and mountains. There are no large trees, and the dwarf bamboos and shrubs have been cut again and
again by the yellow men for fuel. But here, in company with shrikes and mynas, and a flock of magpies,

I found the Silver Pheasant near the rivers.

ss HHUVAADOTOHY

Page
Ie.

<

PHOTOGRAVURE PLATE 24.

a

CHINESE FAUNES OF THE SILVER PHEASANT.

CHINESE SILVER KALEEGE 65

In spite of numerous repetitions of the fact, there seems to be no definite record
of a Silver Kaleege in Chekiang. The Abbé David asserted as long ago as 1877 that
it was becoming very rare in its wild haunts, but gave no reason for the statement.
In Fokien several naturalists have found it in numbers, and back from the sea-coast
I observed it in several isolated parts of the country.

There is considerable shooting and trapping of the birds on the part of the
natives, but no systematic collecting for their feathers as I noted in Yunnan. One
method reported by the Chinese is to build a rough shelter of boughs and to scatter
around it quantities of grain. When the birds have become accustomed to come day
after day for food, the hunter conceals himself in the shelter and shoots the kaleege
as they come to the food. The dead birds are not disturbed, and by remaining quietly
concealed, it is said that as many as eight may be obtained in a single morning. I
have described a similar disregard for the noise of gun-fire on the part of related
species of Burmese kaleege. The Chinese hunters are also reported to shoot the
birds by torchlight from their roosts. It is said that the civil mandarins wear a
Silver Kaleege embroidered on the heart and back badges of their official dresses to
indicate their rank. It is certainly one of the favourite subjects for decorative
painting and embroidery among the Chinese.

The insight which I could gain into their lives showed that their habits differed
in no way from other kaleege. I never saw a bird at mid-day, but in early morning
and late afternoon their noisy scratching would reveal their whereabouts, and a
careful stalk would sometimes give a chance for a shot or a glimpse of white and
black plumage. I observed no regularity of movements, no accustomed trail toward
water, and so was unable to head them off or intercept them. They seemed to wander
about with but little definite direction in view, and I could locate no roosts.

The food of those I examined consisted chiefly of insects, such as grubs and
wire-worms, together with a less quantity of various kinds of berries, and occasionally
flower petals and leaves. La Touche tells of four eggs brought in by a Chinaman
said to belong to the Silver Kaleege. This was in Central Fokien. Two of the eggs
were broken. The others measured respectively 48 by 37 and 48 by 32 mm. In
captivity the Chinese Silver Kaleege is one of the readiest breeders. Indeed no fewer
than seventy-two fertile eggs have been laid by three hens in a season. The eggs
are broad ovals, rather glossy and without pigment markings. They vary in ground
colour from pale to warm reddish buff, usually café aw lait, and sometimes with
small white spots of lime. They measure from 41 to 54 mm. in length, and 38 to 41 in
breadth. The average of many eggs is 39 by 51 mm. From four to six eggs seem
to form a complete set, and the period of incubation is twenty-five or twenty-six days.

There is great variation in length of life, some birds dying with no apparent
ailment, while others live in perfect health for many years. Records made in the
London Zoo show that of thirty-eight individuals the average length of life was a
little less than two years, while the maximum was nine and a half years. There
are other authentic records of cock birds which have lived twelve, eighteen and even
twenty-one years. |

Birds both living and dead are brought into the large Chinese cities for sale,

such as Foochow, Amoy and Canton. But, as Swinhoe says of the Canton birds,
VOL. II K

66 A MONOGRAPH OF THE PHEASANTS

all have been trapped or shot, and I doubt if the kaleege are reared except very rarely
by the Chinese.

In Europe and the United States, however, it is one of the commonest of its
family with dealers, and is one of the first to be obtained and bred by the amateur.
While these birds are beautiful and hardy, their pugnacious habits make it inadvisable
to confine them closely with other birds, while even if given the freedom of coverts,
they will usually drive off the other species of pheasants. When a pair is allowed
to pair and nest, the cock is very faithful in attendance upon the female and in defence
of the young, and this unquestionably points to the monogamous habits of the wild
birds. The cock, of course, usually has nothing to do with incubation, although there
is actually a record of one bird sitting on the eggs. Its mate had laid four eggs, but
failed to hatch young. When a second batch was laid, the cock, in fully adult plumage,
began to incubate, and so successfully that three young Silvers were hatched, and
were reared by the combined efforts of both birds. This is very unusual, however.
But the cock vigorously opposes any threatened danger, and will fearlessly launch
himself at the head of any one of whom he has suspicions, and has been known to
follow a person indoors to continue his assault. The hens are equally brave, and will
drive away cats and dogs whenthey are with a brood of young chicks.

As furnishing sport in private preserves they are unsatisfactory, as they are
difficult to flush, preferring to escape on foot, and even when they take to wing they
merely skim the underbrush, offering little chance for other than snap shots, and these
dangerous to beaters and to fellow sportsmen. Birds shot under such circumstances
have been described as very inferior in flesh, but two which I had cooked in the field
left nothing to be desired. The chief objections to their artificial establishment as
game-birds are their tameness and their habit of coming close to houses and gardens.

The wing whirring and courtship are like those of the other kaleege, and the former
habit is not confined to the breeding season, but seems to function as a warning signal,
and probably in other ways as well. The usual kaleege, broken, semi-liquid, semi-harsh,
guttural cry is characteristic of this species.

When courting there seems to be usually a quite definite utterance, combined
with a wing whirr. The cock approaches the hen slowly and in an indirect, sidling
manner. He then stops suddenly, faces her, stands erect and utters a two-syllabled
note, something like ofv-chac/ short and sudden, the accent on the first syllable.
This is instantly followed by an equally short and sharp whirr, thus o/v-chac (whirrrrr |)
Then the cock runs or walks swiftly around the hen, with tail and wings spread
widely, showing them laterally or frontally, according to his position relative to her eyes.

When the wing whirr is given as a note of suspicion or upon some similar
provocation, it is usually double, thus, whirr! whirrrrrrrrrrrr! a short, quick beat,
followed by a long roll, during which the half-open wings vibrate back and forth in
a maze of motion within a small arc at the sides of the body, the bird standing
very erect.

As a contrast to my observations made on this pheasant in the extreme east of its
range in Fokien, I am glad to quote a few paragraphs written by a correspondent
of Stuart Baker’s, concerning the bird in the extreme west, where indeed the frequent
infusion of orsfieldi blood has led to the name vipfonz.

CHINESE SILVER KALEEGE 67

This observer writes that “in spite of its being found generally in grass, rather
_ than in heavy trees or bush cover, it is not an easy bird to find, and still less easy
to bring to bag when once found. One imagines that such a magnificent bird must
be extremely conspicuous wherever found, but such is by no means the case, and I
have more than once stared at a motionless bird some seconds before I could: make
it out. The stunted and thinly foliaged oaks, which are scattered about at some
distance from each other, give such a queer dappling of light and shade under the
blazing Indian sun that the outline of even glaring white objects cannot be made
out at once, and the broken black and white of the pheasant’s back assimilates well
with the waving grass and the shivering, broken shadows of the oak-leaves.
Every breath of wind which stirs grass and leaves alters your view, and it is not
until the bird rushes headlong away in the open, or skulks, head and tail down like
some wild beast, into the nearest raspberry tangle, that you grasp the fact that you
have let a pheasant get away.

‘“‘Of-course, once they are on the wing they can be seen and heard from a great
distance, but even under these circumstances I have been sometimes so struck with
their beauty that I have failed to fire until too late.

“One of my first encounters with these birds was when, working over the crest
of a grass ridge with my Sepoys, we suddenly put up a covey of full-grown birds,
and I was so fully occupied in watching these streaks of silver loveliness that I omitted
to fire at all, and the whole lot—I think there were seven or eight—disappeared
unharmed down the hill into a ravine with trees and dense undergrowth.

“Often we used to hear these pheasants moving in front of us as our scouts
worked through the grass on either side of our track. The main body of our men were
following, but we very seldom put them up within sight. When we were working
uphill they continued to run ahead of us until they had crossed the ridge or crest
of the hill to our front, and then, when out of sight, they took to wing with much
fluster and noise.

“We noticed they always ran uphill and flew down, and always seemed to make
for the highest point in the vicinity before taking to flight.

“As, on the occasion of which I just wrote, we several times came on conveys
of full-grown cock birds without a single hen anywhere near that we could see, it
may have been that the hens skulked away on foot, but I think not, for the sound
of the running birds could be followed very clearly when the grass and fallen leaves
were dry and rustly.

“They crowed much like the common English pheasant, but a shorter, deeper
sound. I never saw them crowing, but more than once put up cock birds from
spots where I had heard a vigorous crowing and flapping of wings going on the
moment before.”

DETAILED DESCRIPTION

ApuLT Marz.—Top of the head and long nuchal crest black, strongly glossed
with purplish blue; the crest is flowing, disintegrated and directed backward; it
measures sometimes 100 to 110 mm. in length; the length and disintegration is
due, not to the elongation and degeneration of barbules on many barbs, but to the

68 A MONOGRAPH OF THE PHEASANTS

specialization of a few terminal ones. In a single feather 85 mm. in length, the shaft
is only a third of this length, supporting some forty pairs of short barbs. Then the
shaft breaks gradually into a score of barbs, which are tremendously lengthened, and
which form the terminal two-thirds of the feather; these are clothed to their very tip
with very short, barbicel-less barbules. The transition is very abrupt—there are no
barbs of median length; for example, the last of the short barbs may be 6 mm. in
length, while the succeeding one is 60 mm., fully ten times as long.

Ear-coverts, sides and back of the neck, and central pair of tail-feathers pure
white; remainder of the upper plumage of body and wings white, dotted and lined
with black. Mantle, back, rump and all but longest upper tail-coverts, each with five
or six narrow, regular, black, concentric lines. In some cases these are solid throughout;
or again, they are broken into dots near the apex of the feather, giving the appearance
of a distinctly patterned but peppered upper surface; the scapulars, wing-coverts,
tertiaries and inner secondaries are similar, but the black lines are fewer in number,
much coarser and stronger and less regular. This irregularity increases as we proceed
outward on the flight feathers; on the outermost secondary the outer web shows
three wide, almost longitudinal lines of black, which are continued around for some
distance on the inner web, but part-way down the vein they disintegrate into an
irregular mottling; the primaries show almost a reversed condition, the more complete
lining being on the inner web, the outer showing only mottling.

The tail is very long, slender, tapering and graduated. Central feathers pure
white ; lateral ones show a gradual increase of irregular, oblique, black lines; these
first appear basally, on the outer web, and increase in strength and extent, but decrease
in number until on the outer pair we find only three pairs of broad black concentric
lines; the long upper tail-coverts are similarly entirely covered with numerous, rather
thin regular lines.

Chin, throat, fore-neck and entire ventral plumage black, glossed with purple;
the transition from the upper to the under plumage on the sides of the breast is very
abrupt, the bordering feathers usually having the more ventral web black and the
upper web white.

Entire face and lores bare, the skin strongly papillated and brilliant blood-red ;
at the breeding season the three pairs of wattles, so typical of this genus, are strongly
developed; irides red brown; bill usually greenish horn, but varied with darker; legs
and feet scarlet. The spurs are stout, strong, slightly curved and very sharp, very
efficient weapons.

Chinese Silver Kaleege vary enormously in their measurements. The birds I
have from Fokien are the largest. Those from Yunnan show the greatest variation.
I think that even in some specimens which in pigment and pattern are quite typical,
hybridism is shown by the small size in general and the shortness of the tail, approach-
ing 4inmeatus in these respects.

A fully adult large-sized male which I obtained in central Fokien shows the
following measurements: length, 1112 mm.; bill from nostril, 21; wing, 285; tail, 633;
tarsus, 90; middle toe and claw, 70; spurs, 20 mm. I have measured several birds
which equalled these measurements, two from Yunnan. The first, however, was a cock
from north-west Fokien, which excelled by far any other Silver Kaleege I have ever

CHINESE SILVER KALEEGE . 69

examined. Its total length was no less than 1365 mm; bill from nostril, 23; wing,
290; tail, 860; tarsus, ror; middle toe and claw, 75; spurs, 20 mm. The tail, 860 mm.
(over 34 inches), was a marvel of grace, curving upward in a low gradual arc.

VARIATION

We can hardly deal with variation without encroaching upon the domains of
hybridism. However, in birds from the heart of the range, or from Fokien, we find
that in only about fifty per cent. are the central tail-feathers wholly immaculate.
Even the extreme specimen from north-west Fokien whose remarkable size I have
noticed has very pronounced black spotting on the basal half of the outer web. I
have already written of the variation in size, and in the strength of the black dorsal
lining; there are hardly two individuals exactly alike. All, however, lie within such
limits that there is no hesitation in distinguishing a pure-blooded bird from the mere
hybrid forms. I have obtained birds of the typical zycthemerus plumage well within
the ranges of the so-called rzphonz, rujipes and sharper. A number of skins have
come to my attention, taken well within the range of the Silver Pheasant, in which
the legs and feet, instead of being scarlet, were white or greenish, the birds otherwise
being typically zycthemerus. A captive bird of this description has lived for many
years in one of the London parks.

ADULT FEMALE.—The upper plumage a rich golden brown, peppered with darker.
The head is brown, the crest feathers becoming blackish toward the tip. Tail
variable, the outer rectrices blackish, with irregular broken lines of white or buffy-
white, the four central rectrices ashy-brown with irregular dark brown bars and lines.
Below ashy-brown, paler on the chin and throat, and on the lower breast, which
shows more or less mottling and barring with brownish. These markings increase
in strength and size posteriorly.

Bill dull greenish, paler at the tip; facial skin scarlet or dull crimson; irides red-
brown ; legs and feet clear coral red. Bill from nostril, 20 mm.; wing, 225; tail, 285;
tarsus, 88; middle toe and claw, 70 mm.

Cuick In Down.—Head pale orange buff or golden brown, darkening on crown,
becoming narrower and darker brown on nape, and changing to a very wide area of
blackish-brown on the back. This blackish or chocolate brown is split by two pale,
buffy white lines down each side. Shoulders more reddish brown; a short, narrow
black line extends back from the eye; beneath pale yellow buff, changing to white
posteriorly ; facial area well indicated by very short down. Bill greenish yellow;
iris pale hazel; legs and feet red. The newly-hatched Silver is about a third larger
than a golden chick.

JuveNILE PLumace of SEVEN Werks Brrp.—Top of head from crown to nape
brown, so broadly tipped with black that it appears of this colour. Facial area
almost bare, sparsely covered with pale buff down; ear-coverts brown; upper parts
sandy brown, finely vermiculated with black; wing-coverts and secondaries the same,
but with narrow buff tips to the greater coverts. Scapulars and inner coverts with

70 A MONOGRAPH OF THE PHEASANTS

a blackish, subterminal band as well; black markings on outer secondaries very
coarse; primaries barely edged with brown. Chin and throat white; breast sandy
brown, with whitish centres to the feathers, giving a streaked appearance. This area
increases posteriorly, so that the abdomen appears whitish, coarsely but indistinctly
cross-barred with pale brown. The tail is the most striking part of the bird, and,
except for its very small size, closely resembles the tail of the adult hen. There are
eight pairs of rectrices, averaging a full third shorter than those of the hen. The
outer six pairs are clearly patterned in black and white; a black background, with
from six to twelve wide white cross-bars; a brownish border mottling appears on
the third pair, which on the second almost eclipses the cross-barring; the central
pair is cinnamon brown, obliquely vermiculated with black. The longest tail-feather
measures 200 mm.; wing, 170; bill from nostril, 9.

First YEaR MALe.—Similar to the female, but without the ventral markings,
the lower surfaces being almost uniform buffy brown.

The sequence of moults varies greatly in individuals, but in American-reared
birds the young males change into the plumage of the adult male, and thence, in
April of the second year, begin to acquire adult male characters.

EARLY HISTORY

As long ago as 1740, Eleazer Albin gave a plate and a quaint description of the
White China Pheasants, of which he says: ‘This bird I saw at a Lady’s at Enfield
where I made a drawing from it. I do not find this bird described in any author.”
Seven years later George Edwards’enlarges on this, and gives a plate fairly accurate,
at least in form, of the Black and White Chinese Pheasant. His comments are as
follows: ‘These curious birds were kept many years by Sir Hans Sloans, at his
house in London, where they hatched young ones and brought them to maturity.”
He criticizes Albin’s figure, and says that in return for his corrections and additions
he hopes “the curious will not think my labour lost——” It is probable that he
only saw this bird in a yard walking, and took his sketch from it there, so he could
not be so particular as I had the opportunity of being; these birds being the property
of my good patron, whose house I frequented; and had opportunity often to repair
my draughts by strictly examining them in their minute parts, not only while they
were living, but after they were dead.”

In 1766 Linnaeus gave it the specific name which it bears to-day.

SYNONYMY

White China Pheasant Albin, Nat. Hist. Birds, III. 1740, p. 35, pl. XX XVII.

Black and White Chinese Pheasant Edw. Nat. Hist. Birds, I]. 1747, pl. 66.

Phasianus nycthemerus Linnaeus, Sys. Nat. I. 1766, p. 272; Gmelin, Sys. Nat. I. 1788, pt. II. p. 743; Latham
Ind. Orn. II. 1790, p. 631; Bonnat. Tabl. Encycl. Méth. I. 1791, p. 187, pl. 89, figs. 1 and 2; Hays, Osterl. Menag
1794, p. 13, pls. 13 and 14; Timminck, Pig. et Gall. II. 1813, p. 281, III. 1815, p. 665; Vieillot, N. Dict. d’Hist.
Nat., XI. 1817, p. 40; Steph. in Shaw’s Gen. Zool. XI. 1819, p. 234; Griff. ed. Cuv. III. 1829, p. 23; Less. Traité
d’Orn., 1831, p. 495; Schinz, Nat. Abbild. Vog., 1833, p. 248, pl. 95; Jard. Nat. Lib. Orn., IV. 1834, p. 207, pl.
XVIII.; Schinz, Nat. Vog., 1853, p. 148, pl. 71.

Fatsan notre et blanc de la Chine D’Aubent, Pl. Enl. pls., 123, 124; Buff. Hist. Nat. Ois., I]. 1771, p. 359.

Pencilled Pheasant Latham, Gen. Syn., II. 1783, pt. II. p. 719; id. Gen. Hist. VIII. 1823, p. 199.

?Phasianus crawfurdi Gray, in Griff. ed. Cuv. III. 1829, p. 27 (¢ juv.).

CHIEN SES SLE VERS KAVEEGE 71

Gennaeus nycthemerus Wagler, Isis, 1832, p. 1228; Grant, Cat. Birds British Museum, XXII. 1893, p. 307;
Grant, Hand-book Game-birds, I. 1895, p. 277; Rickett and La Touche, Ibis, 1897, p. 602 [Kuatun, Fokien] ;
Oates, Game-birds India, I. 1898, p. 363; Sharpe, Hand-list Birds, I. 1899, p. 36; Rickett, Ibis, 1900, p. 59
[Ching Fung, Fokien]; Grant, Ibis, 1900, p. 606 [S. E. of Bhamo, S. Yunnan]; Oates, Cat. Birds Eggs British
Museum, I. 1901, p. 55; Oates, Ibis, 1903, pp. 94, 97, 98, 99 [cf. with jones]; Ghigi, Arch. Zool., I. 1903, p. 325;
Giinther, Proc. Zool. Soc. London, 1904, p. 371 [hybrid]; Rothschild, Bull. Brit. Orn. Club, 1904, p. 58 [hybrids];
La Touche and Rickett, Ibis, 1905, p. 58 [Central and N.W. Fohkien]; Ghigi, Mem. Acc. Sci. Bologna, 1909, ~
p. 264; Martens, Jour. fiir Orn. 1910, p. 449 [Fokien] ; Finn, Game-birds India and Asia, 1911, p.74; Ingram, Nov.
Zool., XIX. 1912, p. 270; Finn, Avic. Mag. (3), III. p. 102 [variation]; Beebe, Zoologica, I. No. 17, 1914, p. 320;
Bangs and Phillips, Bull. Mus. Comp. Zool. Harvard, LVIII. 1914, p. 269 (Mengtsze) ; Baker, Jour. Bombay Nat.
Hist. Soc., XXIII. 1915, p. 679.

Euplocomus nycthemerus J. E. Gray, Ill. Ind. Zool., II. 1834, pl. 38, fig.2; G. R. Gray, List of Birds, pt. III.
Gall. 1844, p. 25; Blyth, Cat. Mus. As. Soc., 1849, p. 244; Gray, Hand-list Birds, II. 1870, p. 260; Hesse, Jour.
fiir Orn., 1907, p. 217.

Nycthemerus argentatus Swains., Class Birds, I]. 1837, p. 341.

Gallophasis crawfurdi Gray, Genera Birds, III. 1845, p. 498.

Gallophasis nycthemerus Gray, Genera Birds, III. 1845, p. 498.

Euplocamus nycthemerus Gould, Birds Asia, VII. 1859, pl. 17; Sclater, List of Phas., 1863, p. 8 [Southern
China]; Swinhoe, Proc. Zool. Soc. London, 1863, p. 307; Gray, List Gallinae Brit. Mus., 1867, p. 33; Swinhoe,
Proc. Zool. Soc. London, 1871, p. 399; Elliot, Mon. Phas., II. 1872, pl. 21; David and Oustalet, Ois. Chine, 1877,
p. 416[S. China, Fokien, Chi-kiang]; Gurney, Ibis, 1888, p. 230 (female assuming a male plumage). Evans, Ibis,
1891, p. 77 [length of incubation]; La Touche, Ibis, 1892, p. 494 [Yen-ping-fu hills, west of Foochow]; Rickett,
Ibis, 1894, p. 225 [Yen-ping-fu]; Gurney, Ibis, 1899, p. 39 [longevity]; La Touche, Ibis, 1900, p. 49 [Kuatun,
Fokien] ; Tegetmeier, Pheasants, 1904, p. 222; Horne, Pheasant-keeping, p. 37.

Gennaeus argentatus Ghigi, Rend. R. Acc. Sci., 1908, p. 1 [Development of sexual characters].

Gennaeus nycthemerus nycthemerus Baker, Jour. Bombay Nat. Hist. Soc., XXV. 1917, p. 338.

HAINAN SILVER KALEEGE

Gennaeus whiteheadi Grant

NAMES,—Specific: whitehead, after John Whitehead, who lost his life in the expedition on which he
discovered this species. English: Hainan or Whitehead’s Silver Kaleege.

BRIEF DESCRIPTION.—Male: Most nearly allied to xycthemerus. Nape and upper mantle pure white;
remainder of upper plumage with two or usually a single, arrow-shaped submarginal black band, increasing in width
toward the tail; wing markings are strong black lines, few in number; central tail-feathers usually white or with
a few isolated black lines; these increase in width and number toward the outer pairs, which are predominately
black. Female: Resembles more closely the female of “eatus than that of mycthemerus. Crown and nape mottled
brown ; neck and mantle black with wide white shaft-streaks; under parts with the white dominant, the black
reduced to a wide margin ; lateral tail-feathers chestnut.

Types.—In British Museum of Natural History.
RANGE.—The Island of Hainan.

GENERAL ACCOUNT

In the year 1899 John Whitehead, an English collector, visited Hainan, and
discovered the present species of kaleege. Less than eight weeks after securing it he
died of fever. The following extracts from his journal give us all that we know in
regard to the haunts of this splendid bird—

“March 25th. Started for the mountains again to- -day, arriving at 5 p.m. at a small
village, where we camped for the night. Saw a few birds, a Zewscurus in the river, and
a number of fericrocoti, also a few Paleornis javanicus. 1 see neither the white crow
nor the magpie so far inland. Francolins are still abundant. We meet every day
Chinese carrying bundles of skins of deer, monkeys, and pangolins, so I am sure we are
not far from some large forests.

“26th. Our Chinese porters refuse to go any further to-day; they say they have
no rice, but the truth is they are done up and want to rest. The Mandarin sent two
soldiers with us, so that we should be looked after; they have proved most useful.
Some Chinese wanted to prevent our going through their village yesterday: one of
them was, I think, drunk, and nearly had a row with us. The country is slightly less
barren, but still a miserable place for collecting in. Rain all night. The hills have been
so enveloped in clouds that I have not yet been able to see the mountain-tops. I hope
to reach our destination some time to-morrow.

“orth. Instead of reaching our destination to-day, the Chinese have taken us a
day’s journey away from the mountains, as they say there is no road this (the north) side,
so we have had to tramp over bare hills in a broiling sun. My old trouble, dysentery, is
a great worry tome. The Lois, or Lt, as the Chinese call them, are, so far as the men
go, nearly like the Chinese. The hair is at times a pigtail tied in a knob, but more often
a knob of hair tied in front, which looks like a unicorn’s horn. Their villages are small,

the houses of grass and sloped like an arch, and they live on the ground. The women
72

PLATE XXVIII

HAINAN SILVER KALEEGE

Gennaeus whitehead: Grant

Tus bird is very close to the Silver Kaleege, but as it is found. only on an island off the south
China coast, it must be recognized as a distinct species. There is less white on the outer tail-feathers
than in the silver bird, while the female is actually more like the corresponding sex in the Lineated than
in the Silver Kaleege. Mr. Whitehead, who obtained the first specimens, died before he left the country,

and since that time only Japanese collectors have taken the bird.

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PLATE XXVIII.

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HAINAN SILVER KALEEGE.

HAINAN SILVER KALEEGE 73

are quite distinct from the Chinese, wearing very short skirts of blue, or with a red-and-
white pattern.

“28th. Still on the march. Arrived at Lu village in the evening, but the men
seemed so disagreeable that we went about half a mile away and camped. They then
followed to see all they could, and were amazed at our guns. Bought half a pig for
$1.50.

“soth. Left early, as I wanted to camp near the mountain. Sent Andres and
Juan off at 4 a.m. to look fora spot. Nearly had a row with our Chinese porters, but
formed up in their rear, so that they had to go on, after using a good deal of badlanguage
at us. Have struck a nice place for a camp, close to a fine rocky stream not far from the
mountains. Hear lots of birds. Am very seedy with dysentery, Juan with fever. Busy
building a house all day.

“30th. Busy house-building. Caught some beautiful butterflies in the river. Saw
a fine kingfisher, I think /cedo euryzona, also some monkeys.

“31st. This being Good Friday we have resolved not to begin collecting to-day—
unlucky—but to finish off the house. It rained this morning and was very cold before
daylight. A cloudy day, saw no butterflies. Begin collecting to-morrow.

“ April 1st. At last we were able to go shooting. Andres brought in four birds.
Hills very steep, sand and rocks; this is the reason why they are left alone by the
natives. Sand-flies awful. Butterflies numerous in the river-bed. Saw a large king-
fisher and some squirrels.

‘Natives visit us daily : fowls 10 cts., ducks 25 cts., eggs 4 ct.

“and. Very seedy with dysentery. Juan shot a silver pheasant which may be new,
and a fine paradise flycatcher. Heat and sand-flies awful.

“3rd-1gth. All ill with fever and dysentery.

“to9th-30th. The bird collection is going along slowly—my men being all ill one
day or another ; one or two have been ill every day with fever. I am perhaps in better
health than I have been for some time. LU women catching caterpillars for food. It
rains heavily and thunders every afternoon, and our life here, owing to fever, is most
miserable.

‘““May tst-6th. All my men ill with bad fever. The two Chinese and myself do
not suffer much.

“oth. I saw a Li with some rat-traps identical with those made by the Kina Balu
Dusans. Changed into my new house. Very fine for the last few days, norain. Men
(Philipinos) hopelessly ill, and have done nothing for a fortnight. Sent the Chinese
soldier out to shoot for me ; want to see if he is worth anything.

“oth. Three of my Manila men left me to-day of their own accord. I trust they
will reach Hoihow safely. It makes me very sad all this illness. I gave them the
option of going, as I should be very sorry to be the cause of the death of any one of
them.

“toth. To-day is very wet; the Manila men will have a sad time of it, I expect.
My boy Juan is still with me, not because he cares about staying with me, but because
he was too ill to start with the others. The Chinese soldier is doing my shooting and
gets on very well.

“yith-16th. Am trying to work on with one boy, who is not much use at
VOL. II L

74 A MONOGRAPH OF THE PHEASANTS

bird-skinning. The Lt are beginning to bring things, which is a good job, so perhaps
I shall get on after all.

“20th. Attacked again by fever—very seedy—useless. John Afar also.

“21st-23rd. Nearly dead with fever—no food—no depression of temperature.

“24th May.” (Diary ends.)

His Chinese servants carried Mr. Whitehead’s body and all his luggage and
collections back to Hoihow, a journey which occupied nineteen days, and there the
British Consul forwarded the collections to the British Museum. Three males and a
female were secured.

Since then a Japanese has sent a large series to Tring, where I was able to study
them.

DETAILED DESCRIPTION

ApuLtt MaLe.—Under parts wholly dead black, with a steel-blue gloss on the lateral
plumage, and narrow white shaft-streaks on the extreme sides ; forehead brownish black,
merging into the metallic steel blue crown, and the long, flowing, hair-like occipital crest
of the same colour ; a few feathers just behind and concealed beneath the crest are black
and white, but the ear-coverts and the entire back and sides of the neck, and much of the
mantle, is snow white.

On the mantle a very faint brown mottling of two concentric lines appears on
each web, increasing in blackness and solidness posteriorly; at first both pairs are
equal, but on the mid-back the inner pair shows a tendency to close in on the shaft, and
to reassume the mottled character; this pattern either remains to the rump or else the
inner pair coalesces and forms a black shaft-line, leaving but a single concentric pair
on the white webbing; the longer tail-coverts show a strong tendency to oblique
cross-barring.

The wing markings stand out from the innermost least covert as very wide and
strong black lines. These are not concentric, but in the shape of one or two oblique
V-shaped cross-bars, the inner scapulars with as many as four, and the outer with six,
while a few more are added to the primaries.

The central rectrices may be pure white, or may have five or six isolated black lines
on the outer web, running parallel to the shaft and margin; the second pair is usually
white as to the inner web, with numerous very coarse oblique bars on the outer; the
succeeding lateral feathers show a greater and greater extent of black, until it occupies
by far the major part of the outer web. A greater contrast to the lineated pheasant
could hardly be well imagined.

Iris brownish-yellow; wattles and facial skin scarlet ; bill greenish-white ; legs
coral-red. Length, g00 mm.; wing, 248; tail, 520; tarsus, 91 ; middle toe and claw,
61; spurs, 18 mm.

ApuLt FemaLe.—Crown and nape brown, indistinctly mottled with darker and
with a pale shaft-stripe; there is an abrupt change on the hind neck and mantle to black
feathers with a wide, elliptical, tapering shaft-stripe ; on the lower mantle this changes as
abruptly again to a finely vermiculated rufous buff, the shaft-stripe persisting as a narrow,
pale buff streak, dying out on the shorter tail-coverts ; wing-coverts like the back and

HAINAN SILVER KALEEGE 75

rump, but with the shaft-streaks paler and larger, dying out on the longest coverts
and inner secondaries; secondaries show coarser mottling, and are very dark on the
inner web, which in the primaries is plain brownish black. Central rectrices rufous,
vermiculated with black. Lateral rectices chestnut, almost free from dark mottling.

Chin and throat white, changing abruptly to the mantle pattern, although the white
is so predominant that I should invert the description of the pattern, and say that the
ventral surface was white with very wide black margins; on the side neck this merges
insensibly into the mantle; on the sides and under tail-coverts the black increases in
_ width.

Iris light brown ; upper mandible greenish brown, lower dull green; naked skin on
face red; legs and feet bright scarlet. Length, 520 mm.; wing, 205; tail, 228; tarsus,
76; middle toe and claw, 53 mm. .

First YEAR PLuMAGE Mare.—lIs similar to the female, except that the crest of
steel-blue is just making its appearance, and the black of the mantle and hind neck is
replaced with olive; the body plumage as a whole is darker, and more rufous, while the
central rectrices are chestnut, like the lateral ones, and faintly marked with numerous
fine, wavy, oblique bars. The secondaries are similar to the central rectrices.

This, I think, unquestionably shows the derivation of the species from some more
finely marked mainland form in the direction of /zxeatus.

The chin and throat are black, with broad tips and a shaft-streak of white, and the
ventral surface shows a greater amount of black than do the females. Wing, 230 mm.;
tail, 258; tarsus, 86; middle toe and claw, 60; spur, 8 mm.

SYNONYMY

Gennaeus whiteheadi Grant, Ibis, 1899, p. 586 (n. sp. Five-Finger Mountain in Hainan); Grant, Ibis, 1900,
pp. 460, 461, 503, pl. XXXIV. (desc. and coloured plate); Ghigi, Arch. Zool. I. 1903, p. 327; Ghigi, Memoria R.
Acc. Sci. Bologna, 1909, p. 264; Hartert, Novitates Zoologicae, XVII. 1910, p. 191 (Mt. Wuchi) ; Beebe, Zoologica,
I. No. 17, 1914, p. 320; Baker, Jour. Bombay Nat. Hist. Soc. XXIII. 1915, p. 686.

EDWARD’S KALEEGE

Gennaeus edwardst Oustalet

NAMES.—Specific: edwardst. English: Edward’s or Annam Kaleege.

BRIEF DESCRIPTION.—Male: Cap white; body metallic purplish black, with pronounced terminal fringe
on rump and tail-coverts ; wing-coverts shining green with disintegrated fringe ; wings and tail black ; facial skin,
legs and feet scarlet. Female: Brownish-grey ; paler on the neck, and becoming more rufous on scapulars and
wing-coverts ; tail blackish; face and legs scarlet.

TyPeEs.—In Paris Museum.

RANGE.—Province of Kuang-tri in Annam,

GENERAL ACCOUNT

ALTHOUGH this very distinct species has been known for nineteen years, absolutely
nothing has been recorded in regard to its haunts or habits.

In the Museum d’histoire naturelle in Paris there are four specimens, three
males and a female, the only ones of this species which have been thus far collected.

Oustalet, in his description of the species, as given in the ‘ Bulletin Museum
d’histoire naturelle” in 1896, adds: “This exceedingly remarkable species, which the
Museum has acquired from P. Renauld, consisting of the skins of two adult males,
a young male and a few odd bits of plumage, forms in certain respects a transition
between Gennuaeus with roof-shaped tail, and Acomus with a flattened tail.” I am
quite certain that the mounted bird which Oustalet considers a young male is a
fully adult female. The details of coloration are rather those of a female than a
male in first-year plumage. There are absolutely no signs of immaturity in the
plumage, and this, in connection with the nodule-like spurs, would indicate that the
bird, which was unsexed by the collector, is a full-grown female. I have described
it as such. The final suggestion made by Oustalet is an error, since the tail of
Acomus is as distinctly roof-shaped or laterally compressed as that of Genxmnaeus.
The relation with swzhodc which I have mentioned elsewhere is very apparent, as
in the crest, tail, specialization of feather structure, and other characters.

DETAILED DESCRIPTION

ApuLt Matse.—Facial skin corrugated and scarlet; forehead, narrow superciliary
(above facial skin), posterior crown, chin, upper neck and ear-coverts brownish-black ;
stiffened crown feathers moderately elongated into a crest, white, tipped with black ;
the sharp transition into the short, black, posterior crown is by a few lines of short
feathers, white crossed by several black bars; the black feathers merge at once into
the typical body plumage; this is shining purplish blue, the former tint predominat-

ing on the neck, breast, sides and mantle, and changing almost insensibly into a
76

PLATE XXIX

EDWARD'S KALEEGE

Gennaeus edwardst Oustalet

AttTHouGH known for a quarter of a century, absolutely nothing has been recorded of the habits of
this very distinct and beautiful pheasant. Four specimens have been obtained from the province of

Kaung-tri in Annam.

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EDWARD’S KALEEGE.

EDWARD'S KALEEGE 77

purer metallic blue posteriorly. The most striking difference lies in the fringe.
While a broad, disintegrated fringe exists on the mantle, it is hardly apparent, but
on the lower back, rump and tail-coverts the fringe becomes convex, reflecting the
light from a different angle. So although the entire feather is actually shining blue,
a side view shows the convex portion of the tip as deep velvety black, in contrast
to the remaining basal and extreme terminal parts. The tail-feathers have a faint
bluish lustre on the outer webs, but on the: whole appear dull black in contrast to
the rump and tail-coverts.

The wings are rendered distinct by all the coverts being green, with convex
fringe, as on the rump. The post-fringeal part is rather less metallic in colour.
The illusion of a black centre to the shining green fringe is most persistent, and
only when the wing is looked at in perspective, held away from the light, does the
green sheen creep over the entire fringe. In this position the post-fringeal part
becomes black. The black-centred, green tips of the coverts, arranged in row after
row across the median part of the wing, are very conspicuous. The inner second-
aries are tinged with a greenish-blue lustre, but the remainder are brownish-black,
while the primaries are lighter brown. The belly, thighs, vent and under tail-
coverts are dull brownish-black.

Length (Oustalet), 580 mm.; bill from nostril, 16; wing, 220; tail, 240; tarsus,
76; middle toe and claw, 51; spur, 15 mm.

ADULT FEMALE.—The plumage is very sombre, consisting of various shades of
brown, with almost no striking or distinguishing character. The crown is rather
dark, paling into an ashy grey on the neck. The entire under parts are unmarked
greyish-brown, while the upper plumage is warmer. A mottled fringe is found on
the feathers of the back, which on the wing-coverts, secondaries, rump and _tail-
coverts becomes a fine, dark, inconspicuous vermiculation, evenly distributed over
the entire feather. The wing-feathers and scapulars are more rufous than the rump,
which tends slightly toward pale buffy.

The central two or three pairs of tail-feathers are tinged on the outer web with
an inconspicuous dark purplish-brown, the remainder of the tail being brownish-
black as in the male.

Bill from nostril, 15 mm.; wing, 200; tail, 193 (feather tips missing); tarsus, 68;
middle toe and claw, 46. Spur, a sharp flattened nodule.

SYNONYMY

Gennaeus edwardsi Oustalet, Bull. Mus. Paris, 1896, p. 316 (n. sp. Kuang-tri, Annam); Oustalet, Arch. Mus.
Paris (4), I. 1899, pl. 10 (figure) ; Ghigi, Arch. Zool. Ital., I. 1903, p. 325 (Relation to szzhoet); Ghigi, Mem. Acc.
Sci. Bologna (6), VI. 1909, p. 260 (in genus Hzerophasis, with swinhoii ; Beebe, Zoologica, I. No. 17, 1914, p. 320;
Baker, Jour. Bombay Nat. Hist. Soc., XXIII. 1915, p. 657.

SWINHOE’S KALEEGE

Gennaeus swinhoiw (Gould)

NAMES.—Specific : szvzzhozz, after Robert Swinhoe, the discoverer of the species. He was for many years the
British Consul in Formosa. English: Swinhoe’s or Formosan Kaleege. French: Faisan de Swinhoe. German:
Formosa-Fasan. Vernacular: Wa-koé (Chinese, adult male); Aw-kak or Awbay-kak (immature black-tailed male),

BRIEF DESCRIPTION.—Male: Facial skin scarlet ; head, chin and throat black ; crest, mantle and central
tail-feathers white; scapulars dark metallic crimson; most of the wing-coverts with metallic green fringe ; remainder
of plumage glossed with purplish blue. Female:. Crest short, black, tipped with reddish-brown ; upper plumage
and wings black, thickly mottled with reddish-brown; mantle, back and wing-coverts with a conspicuous arrow-
shaped buffy mark ; flight-feathers black, banded with rufous and whitish-buff; ventral plumage rufous-buff,

obliquely barred with black ; central tail-feathers irregularly barred and mottled with black and white; other
rectrices dark chestnut.

RANGE.—The mountain forests of Formosa.

GENERAL DISTRIBUTION

We have no definite information as to the distribution of Swinhoe’s Kaleege,
except that it is not found near the coast of Formosa, but only in the wooded
mountains of the interior, and from the few certain records appears to occur from
five to seven thousand feet, being most abundant in the central, and rare in the
northern portions of the island.

GENERAL ACCOUNT

My account of the haunts and habits of Swinhoe’s Kaleege is second-hand, as
I had no opportunity of visiting Formosa and studying the bird in its native home.
For many years Mr. Robert Swinhoe was the British Consul in Formosa, and it
was due to his efforts that the first specimens of this pheasant were made known to
science. In regard to this species, which was described in 1862 by Gould, and
named after the discoverer, Mr. Swinhoe from time to time sent the following notes—

“T was informed by my hunters that a second species of pheasant, which was
denominated by the Chinese colonists W4-koé, was found in the interior mountains,
that it was a true jungle bird, frequenting the wild hill-ranges of the aborigines, and
rarely descending to the lower hills that border on the Chinese territory, and that in
the evening and early morning the male was in the habit of showing himself on an
exposed branch or roof of a savage’s hut, uttering his crowing, defiant note, while he
strutted and threw up his tail like a rooster. I offered rewards and encouraged my
men to do their utmost to procure me specimens of this bird, and I was so far
successful that I managed to obtain a pair; but in my trip to the interior it was in
vain that I sought to get a view of it in its native haunts, and to make acquaintance
with it in a state of nature.

“The female was brought to me on the ist of April, soon after it was shot,
78

PLATE XXX

SWINHOES KALEEGE

Gennaeus swinhow (Gould)

In the deep inland forests of the mountains of Formosa this splendid -bird makes its home. None
but the natives have observed or captured it, and there is no record of the nest and eggs being found,

but the bird lays readily in captivity and is not rare in zoological gardens.

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PLATE XXX.

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th

SWINHOE’S KALEEGE.

“SWINHOE’S KALEEGE 79

the heat of the weather compelling the hunters to skin it before they could reach
me. It was, however, quite fresh enough to enable me to note the tints of its soft
parts. The fresh skin of the male arrived on the 11th of April. My hunters
had taken this bird alive, but it battered itself so that they were obliged to kill it
to save its feathers.

“ . . On the 8th of December I procured a fine male of Euplocomus swinhoti ©
in beautiful plumage. It was brought to me from a distant station in the interior
of this island, and I forwarded it to Hong-Kong, whence it was shipped to Dr.
Squire, at Calcutta, for the London Gardens. I trust it may arrive all safe. This
bird is rare, and extremely difficult to procure, as the mountain travelling here is
far from safe. My chief bird-hunter was nearly murdered and robbed of fifty pounds,
the other day, while in search of deer and this pheasant.

“T have seen many males with a plumage intermediate to that of the adult and
their own sex and that of the females. This plumage is carried through the winter ;
but it varies in its resemblance to the one sex or the other. I thought at first that
such birds were melanite varieties, especially as the Chinese distinguished them by
a distinct name, 4w-bay-kak (black-tailed male); but I consider now that they are
only young males in the transition plumage, which they carry till the next vernal moult.

“T have been so unsuccessful in getting live examples of Euplocamus swinhoit
home, that I have had an aviary built here, and stocked it chiefly with birds of this
species. I intend to keep them for some time, to get them into thorough condition,
and then try further shipments. Those I have, though several months in confine-
ment, are still shy birds, and skulk in holes the greater part of the day. They
frequently utter a plaintive note ‘co-co-co-coo, the last a low wail, almost im-
possible to syllable. One fine skin of a hen I have got has a snow-white patch on
the crown, and a few white feathers on the side of the jaws. In other respects it
is normal.”

While nothing has been recorded of the nesting habits of the wild bird, its eggs
are well known, captive hens occasionally laying, but seldom rearing their young.
Swinhoe reports that a hen in his possession laid an egg every two or three days,
the first being deposited on March 17th.

The eggs of Swinhoe’s Kaleege are regular ovals, smooth, rather glossy and
unmarked. The shell varies from pale reddish-buff to creamy-white. They measure
from 48 to 55 mm. in length, and from 37 to 39 in breadth; averaging 51 by 38 mm.
Swinhoe gives 61 by 48 as the measurements of an egg, and adds that it is buffy-
cream, very minutely dotted with white.

CAPTIVITY

Saint-Hilaire has given us some interesting records of Swinhoe’s Kaleege in the
Jardin d’Acclimatation in Paris. The first specimens were received in 1866. The
following year twelve young birds were hatched, and in 1868 seventeen. In 1869 the
birds began to lay March 15th and ended June 25th. From four adult hens sixty-nine
eggs were obtained, from which fifty young birds were hatched, but on October 15th only
fifteen, eight males and seven females, remained alive.

80 A MONOGRAPH OF THE PHEASANTS

The price for the first pair of Swinhoes was between four and five thousand francs
(about $900). In 1868 the price per bird had fallen to three hundred and fifty francs
($70), and the following year it averaged two hundred and fifty francs ($50).

Of thirty-two Swinhoe's Kaleege which have lived in the London Zoo the average
length of life was one year, although one bird reached the age of five and a half years.

As we have seen, Swinhoe’s Kaleege does not attain its fully adult plumage until
the second year, but the birds will breed the first year while still in immature dress.
Thus, of forty-one eggs from three young hens, twenty-nine, or about seventy per cent.,
were fertile. This, however, is exceptional, and the males seem to be much more inclined
to breed the first year. As with golden and silver pheasants, this precocious breeding
seems to increase with successive generations in captivity. Whereas wild trapped birds
of the first generation will produce but few fertile eggs in their first year, succeeding
generations prove much more willing and successful in this respect.

VARIATIONS.—An interesting variety, or perhaps true mutation, comparable with
the black-shouldered peafowl and the black-throated golden has been described by Ghigi.
This was one of four Swinhoe’s Kaleege hatched from a brood of seven eggs, the parents
being normal in every way. The bird in question was yellowish white.

DETAILED DESCRIPTION

ApuLT MALe.—Posterior crown and nape with a flowing white crest. Back of neck
and entire mantle down to the mid-back white. Remainder of head and entire under
parts black, with a strong purplish sheen on all exposed portions of feathers, becoming
dull brownish black on posterior abdomen. Scapulars deep crimson, forming very
conspicuous patches on each side of the white mantle. Wing-coverts, lower back, rump
and tail-coverts black, all with a specialized terminal band. In the wing-coverts this
is metallic green, while in all the other feathers it is a brilliant, deep purple. The tail
feathers are black, with the central pair and outer part of the second pair white.

Facial skin scarlet, covered with roughened papillae and a scattering of small
feathers ; legs and feet deep red ; irides bill yellowish horn.

Length, 790 mm.; bill from nostril, 22; wing, 250; tail, 410; tarsus, 93; middle
toe and claw, 70; spur, 35 mm.

The specialized terminal fringe on the wing-coverts, lower back and tail-coverts
corresponds in gennaeus edwardsi, and is characterized by three abrupt changes in the
vane, first, the total loss of all barbicels, second, a pronounced flattening of the barbules,
and an alteration in the angle of attachment of these barbules to the barbs—the two
rows being raised, so that, instead of lying on opposite sides in a flat plane, they form
an obtuse angle with each other.

AvuLt FemMaLte.—There is a short inconspicuous crest, which, with the remainder
of the head and neck, is greyish brown, mottled indistinctly with black; mantle,
scapulars and wing-coverts buffy-brown, with large, arrow-shaped centres of rufous buff,
margined with black. Lower back, rump and upper tail-coverts black, finely vermic-
ulated with buff, becoming greyer posteriorly. Primaries dark brown, secondaries
black, the former barred with rufous, the latter with buff. Chin and throat greyish

SWINHOE'S KALEEGE 81

brown. On the upper breast a buffy shaft stripe appears, which widens posteriorly until
the dominant colour of the lower breast and abdomen is rufous buff, with irregular,
black, V-shaped markings. The colour of the ventral: plumage is variable, some birds
being much redder than those of equal age and purity of blood. Central tail-feathers,
like the upper coverts, with indistinct black bars strongly marked with pale grey. Outer
rectrices dark chestnut, mottled irregularly on the inner webs with black.

Facial skin, legs and feet dull scarlet ; irides brown ; bill yellowish brown.

Length, 505 mm.; bill from nostril, 18; wing, 240; tail, 200; tarsus, 80; middle
toe and claw, 50; spurs, a low sharp nodule.

First YEAR Mare.—In the post-juvenile plumage the crest is only partly white,
being thickly barred with black. The white in the mantle is apparent only as a wider or
narrower shaft-stripe, the remainder of the feathers being chestnut and black. The
metallic tips of the wing and back plumage are often lacking, or present only in the tail-
coverts, while these feathers in general and the central rectrices are chestnut, more or less
mottled with black. There is no blue sheen on the ventral plumage. No two males are
alike at this stage, and in a bird whose moult has been delayed by cold weather I
have seen every adult character well developed, and only to be distinguished from
full-plumaged males by the impurity of the whites and a slight excess of chestnut.

Cuick IN Down.—Top of head from forehead to nape orange rufous, with a dark
central. crown line; facial area over and around eye whitish buff, with a black line
obliquely downward and backward from the lower posterior rim of orbit over the
ear-coverts to the lower nape; upper body, including sides and wings, dark chocolate ;
two creamy white lines begin faintly on the mantle and suddenly become broad and
distinct on the lateral upper back, where they are strengthened into a spot by an
adjoining patch on the inner wing edge. Posteriorly there is a short break, and then
the typically pheasant-chick character follows, of two broad lines down each side of the
lower back ; these widen interiorly at the end just above the tail, but do not join. The
tail down is rufous, as is also an indistinct line connecting the thighs with the tail ; lower
side, neck, outer thighs and breast also rufous, the latter being less pure in colour; chin,
throat, and abdomen pure creamy white ; across the down of the middle coverts is a very
broad bar of buffy white.

Bill from nostril, 3 mm.; wing, 33; tail (down); tarsus, 22; middle toe and claw,
22 mm.

TweLvE Day Cuicx.—Head and neck as before; wings have increased in length,
and the tail, though short, has appeared ; scapulars show as two lines of new feathers
down the back, while the ventral surface shows two corresponding lateral lines extending
from the lower neck to the thighs, all else being down.

The oth and toth primaries are only a short way out of the sheaths, while the others
show a very equal growth. Measurements of the roth and 8th measure respectively 12
and 63mm. The appearance of the wing with the few first feathers is much simpler
than that of two weeks later. The primaries are dark brown, with pale buff mottlings

on the outer webs, This mottling is more extensive on the secondaries, showing a
VOL. II . M

82 A MONOGRAPH OF THE PHEASANTS

tendency to form a dark line along the outer margin. The greater coverts (which are all
that have appeared) show a wide pale buff tip, with two lateral, subterminal, round spots
of black.

Bill from nostril, 6 mm.; wing, 96; tail, 33; tarsus, 28; middle toe and claw,
28 mm.

Five Weeks OLp Curcx.—A chick of this age is clothed in juvenile plumage
except for the chin, throat and facial area, which are still in down. Culmen, 7 mm.;
wing, 119; tail, 76; tarsus, 35; middle toe and claw, 33 mm.

E1guHt WEEKs OLD Birp—In full juvenile plumage. Facial area showing only a
few scattered down featherlets ; entire upper head and neck and the sides of latter dark
seal brown, paling gradually toward the throat into white on the chin; feathers of chin
and throat, sides of lower face and neck recurved, standing up separately. General tone
of upper parts very dark brown, finely mottled with paler buff. The mantle and
scapulars marked with an expanded, subterminal, elongated shaft-spot of clear buff.
The back, rump and central pair of tail feathers show only the fine vermiculations, these
being grey rather than buff. The buff shaft-spot is present on all the lesser and median
coverts, becoming the terminal band of the greater coverts, but all the wing-coverts are
sharply marked off from the feathers of the adjacent portions of the body by the two
subterminal black spots. Toward the greater coverts these spots join and form a solid,
conspicuous band of black just posterior to the terminal buff. This gives a handsome
barred appearance, several lines of the successive buff and black. On the innermost
secondaries the buff bar disappears, while the black pushes forward and forms a long,
narrow margin. The primary coverts, like their flights, show no definite characters,
being irregularly mottled with rufous and buff.

The 8th, 9th and roth primaries are still growing strongly, showing that there is
considerable delay in the growth of the first as well as the two outer ones. Numbers 4,5,
6 and 7 are full-grown juvenile primaries. Number 1 is new, measuring 94 mm. out of
the sheath, No. 2 is a bare 13 out of its sheath, while No. 3 has apparently just fallen
out.

The juvenile primaries are curved, narrow and chiefly dull brown with little
mottling. The new feather is broad, very dark, and with rufous marking on both
webs, roughly longitudinal on the outer web and irregularly transverse on the inner.

There are six pairs of rectrices. The outer five pairs are much alike, rich rufous,
mottled irregularly with black, while the central pair is cold, dark brown with greyish
mottlings. There are faint indications of six or more pale lateral bars. The two
central pairs are still growing strongly. All are juvenile. The outermost (full-grown)
feather measures 81 mm.; the central (growing) ones 126 mm.

The under parts show a warm orange hue, beginning as a shaft-stripe below the
grey of the lower throat, which rapidly enlarges on the breast and belly, where the
black lateral marking is reduced considerably. The orange pales on the thighs and rear
under parts.

Bill from nostril, 10 mm.; wing, 147; tail, 127; tarsus, 50; middle toe and claw,
43, tam. |

SWINHOE’S KALEEGE 83

SYNONYMY

Euplocamus swinhott Gould, Proc. Zool. Soc, London, 1862, p. 284 [Formosa] ; Sclater, List of Phasianide,
1863, p. 7; Swinhoe, Ibis, 1863, p. 401 [Formosa, notes of discovery]; Gould, Birds of Asia, VII. 1864, pl. 16;
Swinhoe, Ibis, 1865, pp. 353, 538 [Plumage and notes]; Swinhoe, Ibis, 1866, pp. 133, 308, 404 [Notes on plumage] ;
Swinhoe, Ibis, 1867, pp. 232, 409 [Notes in captivity]; Sclater and Wolf, Zoological Sketches, 2, 1867, pl. 37;
Gray, List Galline British Museum, 1867, p. 34; Touchard, Bull. Soc. Acclim., 1870, p. 417 ; Swinhoe, Proc. Zool.
Soc. London, 1871, p. 399; Elliot, Monograph Phasianide, II. 1872, pl. 25 [subgenus Hierophasis|; David and
Oustalet, Oiseaux de Chine, 1877, p. 417, pl. 102 ; La Touche, Ibis, 1895, pp. 329, 338 [Bangkimtsing, South Cape] ;
La Touche, Ibis, 1898, p. 373 [Central Formosa]; Tegetmeier, Pheasants, 4th ed., 1904, p. 227 ; Mitchell, Proc.
Zool. Soc. London, 1911, p. 522 [inability].

Euplocomus swinhoé Saint Hilaire, Bull. Soc. d’Acclim. (2), VII, 1870, p. 131 [Breeding in Jardin D’Acclim.].

Euplocomus swinhoet Horne, Pheasant-keeping, p. 42 [frontispiece].

Faisan swinhoe Touchard, Bull. Soc. d’Acclim. (2), VIII. 1870, p. 417.

Euplocomus swinhaet Mairet, Bull. Soc. d’Acclim. (2), VII. 1870, p. 348.

Euplocomus swinhow Gray, Hand-list of Birds, II. 1870, p. 260.

Gennaeus swinhow Grant, Cat. Birds Brit. Mus, XXII. 1893, p. 309; Grant, Hand-book Game-birds, I. 1895,
p. 278; Nehrkorn, Kat. der Eiersammlung, 1899, p. 193; Ghigi, Arch. Zool., I. 1903, p. 324 [plumages] ; Grant
and La Touche, Ibis, 1907, p. 276 [Racu Racu Mts. 7000 feet; Ho Ho Mts. 5000 feet]; Ghigi, R. Acc. Sci. Inst.
Bologna, 1908, pp. 1-14 [Mutatian in captivity] ; Ghigi, Memoria, Sci. Inst. Bologna, 1909, p. 260 [genus Hzerophasis];
Ghigi, Monitore, Zool. Ital., XX. 1909, p. 95; Finn, Game-birds of India and Asia, 1911, p. 79; Beebe, Zoologica,
I. No. 17, 1914, p. 320; Baker, Jour. Bombay Nat. Hist. Soc. XXIII. 1915, p. 687.

Gennaeus swinhoet Sharpe, Hand-list of Birds, I. 1899, p. 36; Oates, Cat. Eggs Brit. Mus. I. rgo1, p. 56;
Rothschild, Bull. Brit. Orn. Club, XIV. 1904, p. 58 [Hybrids with G. nycthemerus, L. diardi and L. ignita].

(G) Hrerophasis swinhoet Ghigi, Riv. ital. Orn., I. 1912, p. 272.

WILD KALEEGE HYBRIDS

KALEEGE HYBRIDS IN THEIR WILD HOME

STANDING on the high divide which shunts its eastern waters into China, and
its western into the great rivers of Burma, one sees, in the’ direction of the latter
country, a great tumbled, irregular mass of mountains and valleys. All is forest,
clad with bamboo, oak and other hard woods, and it was here, hidden beneath that
vast extent of many-tinted foliage, that I found pheasants which, from the point of
view of their origin, were the most remarkable of their family.

Elephants and mules were the commonest means of transport, and I found it
necessary to take an escort of six Gurkhas. The Kachin tribes hereabouts are
nominally safe, but the individual components of these tribes are uncertain quantities.
As still-hunting was my method of finding and observing these pheasants, and as I
always carried a ‘303 rifle cartridge in the third barrel of my gun, I worried little
about human enemies and only twice was even threatened with any molestation. A
few miles to the north, however, the wild tribes are wholly independent, and work
their pleasure upon strangers.

We spent many delightful days in the study of these birds, glorying in the
wonderful scenery and magnificent climate, after many months of hot, steaming,
tropical jungles. The early November mornings were keen and clear, and every
valley and depression was always filled to overflowing with a calm, waveless lake of
cloud, while the farthest Yunnan peaks were of a deeper purple than ever painter
dared put to canvas. The sweetness of the chorus of bulbuls was the major theme
at this hour, with a minor accompaniment of distant cooing doves.

But pheasants were difficult to find in the morning, and one might wander
about for hours with never a glimpse of them.

About three o'clock in the afternoon of one of my first days in this region,
when the sun still held back the sting of the coming night air, I left camp and
turned down one of the old native trails. The ground was littered with dried leaves,
and the wind soughing through the bamboos gave an added hint of autumn even
in this southern latitude. The rains were just over and the foliage was bright and
clean. I crept as quietly as possible down to the very bottom of a deep ravine
which the sun’s rays had already left. I knew that the pheasants were certain,
sooner or later, to come down to this level for their evening drink. Near the low
murmur of the rivulet I seated myself and began my vigil. For an hour I sat thus,
making certain that the birds had not yet come down. Through the curtain of lofty
ginger stalks overhead I could see drongos darting here and there after insects.

Small flycatchers and babblers passed in flocks, drinking and flitting upward again.
84

PHOTOGRAVURE 1

NORTHERN BURMA—YUNNAN BORDER—HOME OF THE HYBRID KALEEGE

Sranpinc on the high divide which shunts its eastern waters into China and its western into the
great rivers of Burma, a great, tumbled, irregular mass of mountains and valleys is seen. All are forest,
clad with bamboo, oaks and hard wood, and here, hidden beneath that vast extent of many-tinted foliage,

I found the northern Kaleege pheasants.

EINE, OF KACHIN PHEASANT TRAPS

Tuer Kachins and other native tribes use a deadly method of trapping. A low fence of split bamboo
runs uphill and down for a mile or more, with deadfalls every few feet. When this is in full operation,

few pheasants fail, sooner or later, to be caught, and whole districts are thus cleared of these splendid
birds.

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PHOTOGRAVURE PLATE

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NORTHERN BURMA-YUNNAN BORD

WILD KALEEGE HYBRIDS 85

Mosquitoes rose in clouds and pestered me sorely. Once the low tree-ferns on the
opposite bank were shaken, and through the deeper shade of their fronds I saw a
small tiger-cat passing, slowly, sinuously. He, too, sensed that pheasants come here
to drink.

Knowing from the silence that they were not yet among the bamboos above, I
crept on up the valley. Tree-vines had hung their great masses of bloom overhead,
and graceful wisteria-shaped flowers lightened the gloom with their pink and salmon
petals, and spread far their musky odour—that of.hemiptera. Some four-footed
creature dashed from my path and, marking its fright, left another sharp stratum of
musk upon the air. |

I came upon a maze of footprints, where pheasants had that morning crossed
the muddy rim of the pools, and here I turned upward. I know of no more difficult
feat than attempting to climb noiselessly up a steep bank through clumps of bamboo,
the ground covered with the driest of sheaths and leaves. Finally I passed the
grave of a Kachin chief, covered by an oval, thatched hut and a curious ornament
of dyed bamboo. Just beyond I reached the mule trail, which at this point cut into
the bank of the upper slope. Still hearing nothing, I climbed half-way to the summit
of the ridge, here an open growth of oaks, when suddenly a shift in the breeze
brought to my ears a loud scratching and rustling among the fallen leaves beyond
the summit. I was exposed to full view, so with all possible speed I backed down
the slope on hands and knees, crossed the trail and ensconced myself in a small
thicket, which gave me full view of the oak slope which I had just left.

For half an hour I heard nothing, then a leaf flew upward from a tangle of
vines, and a sturdy form leaped high over a log into view. It was not a pheasant,
but a big, black-gorgeted laughing thrush. Another and another hopped down the
slope, now hidden by tree-trunks or bushes, now standing out in full silhouette.
There were sixteen in all, spread out in a segment of a circle, and chuckling low
to themselves at every succulent morsel. They are splendid, sturdy birds, jay-like
from beak to claw, now holding a wormy acorn and pounding away as hard as a
woodpecker, then, ant-thrush like, picking up leaves and throwing them far over
their backs. I was absorbed in watching their gradual approach when a jungle-fowl
crowed loudly in the valley beyond the ridge, and brought my mind sharply back to
pheasants. I was keenly disappointed at having apparently missed my birds, and
half rose to go. At my first motion a laughing thrush set up a truly jay-like yell,
and answers came from a score of throats, guffaws and peals of loud laughter which
no real jay could ever produce. I sat quiet, their alarm passed, and they began to
sail overhead down the valley. Not being certain at this time of the species I fired
and secured one.

I waited five minutes and heard not a.sound, save the calls of the laughing
thrushes far down below me. Rising stiffly, and slowly moving out into the trail, I
began to reload, when half-way up the slope a black head and neck shot up, and
the warning or suspicion cry of a kaleege pheasant rang out sharp and shrill.

I dropped flat upon the trail, and wriggled back over the edge into my thicket
again. Not a cluck or call came from the slope above, but little by little a low
sub-sound of rustling leaves, and in ten minutes the ground over which the laughing

86 A MONOGRAPH OF THE PHEASANTS

thrushes had passed was being quartered by eleven splendid pheasants. With
balanced glasses I could see every feather. Four were adult cocks, four more were
hens, while the other three were nearly grown young males. Without doubt four of
them comprised a still united family of the present year, while five others seemed to
represent another. To my surprise I could easily distinguish between three of the
male birds. A solitary cock was the lightest of all, one of the young males appeared
as dark as a black-breasted kaleege, while its brother was lightly vermiculated. I
have described these in detail elsewhere. I watched the dainty birds, stepping high,
like thoroughbreds, snatching an insect or leaping at some morsel on a leaf overhead,
or picking up an acorn, ever alert and watchful. I remained as still as the tree-
trunk at my back, and the birds descended half-way down the slope toward me.

Then two Kachin women, with silver cylinders and tassels in their ears, and
great baskets on their backs, came along, chattering loudly. They halted when they
saw me, and despite all my motions stood stupidly gaping at me for several minutes
before they plodded on their way. The pheasants had, of course, retreated to cover,
and when, twenty minutes later, they returned they were spread out more irregularly.
I secured the light-coloured old male, which I had seen once before, while the others
passed me on either hand, together with a jungle-cock, which in bearing and gait
was not to be compared with the far more elegant and graceful pheasants. Except
for a short, sharp alarm note and five minutes of silence, the rest of the flock paid
no attention to the roar of the gun. As I had opportunity to notice on many other
occasions, if one shoots from a thicket and makes no movement after firing, the
birds seem to have no sense of direction of the danger, and are but little affected
by the sight of their dead companion. When headed down toward water I have
never known a flock to be turned back by shots fired in this way, and have secured
as many as four from the same ambush.

As I shall describe in detail, even the two birds of the year which I later
secured, although appearing exactly alike at a distance of forty feet, turned out to
differ sufficiently to fit them to two of the so-called species of the closet taxonomists.

The following day the same route was followed by both laughing thrushes and
pheasants, and on each of the succeeding six days, when my observations ceased.
In no fewer than eight other flocks, or more properly families, of pheasants in the
hills farther to the east I found the same interesting relation between the two
different groups of birds.

Early in the morning the birds worked uphill toward the higher, warmer ridges
rather irregularly and at no special time, early or as late as nine o'clock, as the
fancy or abundance of food influenced them. At this time they kept together in
small family parties, uniting with others only when starting down for the evening
drink. Mid-day was spent in dense bamboo thickets or tangles of thorn palms,
where observation of them was almost impossible. I once watched three birds
apparently picking ticks from one another's heads, and even from under the uplifted
wings, at full noon, in the dense shade of some fallen vines. Toward two in the
afternoon of a partly cloudy day, or about three if the sun shone warmly and
uninterfuptedly, the pheasants began calling to one another in, undertones—sweet
notes which much resemble the voice of our own bluebird, without, however, the

PHOTOGRAVURE 20

A SANCTUARY OF THE HYBRID KALEEGE

Tue natives of northern Burma captured many specimens of Hybrid Kaleege for me, taking them
either in deadfalls or in nooses. But there was one spot near their villages where they would never trap
or kill. This was the sacred spot or grove devoted to the Nats or evil spirits. And here the pheasants

were safe from molestation, and here they scratched for food and roosted high at night.

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PHOTOGRAVURE PLATE 20

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SACRED NAT GROVE, WHERE THE HYBRID KALE

WILD KALEEGE HYBRIDS 87

plaintive tremolo. The young birds—nearly in adult plumage as they were—now and
then varied these sweeter notes with chick-like seeps! and peeps !

Gradually working together, with the laughing thrushes drifting along like
scattered leaves or bounding with high, strong leaps over the low bushes and logs,
all united in a loose flock, and began feeding slowly downward, usually over a
southern slope. The greater activity of the thrushes usually carried them several
yards in advance before they had gone far, but many times I watched the birds at
a distance, and saw them keep together for a thousand feet or more of descent.
In such a case I would locate the flock as it crossed an open space well up on the
mountain, and making a detour, and concealing myself far below in the line of their
descent, I would be almost certain to intercept them before they reached water.
The thrushes were almost wholly insectivorous, while the pheasants chose animal
and vegetable food in equal quantities.

Whether the relation is mutually helpful in any way or not, it certainly exists.
And, as I have said elsewhere, while the association may be due solely to the social
love of birds, it is certainly true that the laughing thrushes many times give the
pheasants warning of danger visible from trees, which the latter on the ground could
never detect until too late; then, while the pheasants scratched or pecked to pieces
some fallen log, I have again and again seen several thrushes stand around, now
and then springing into the air to seize an insect which had escaped by flight from
the larger birds.

Two species of laughing thrushes are thus found in intimate association with
the pheasants, the black-gorgeted and the black-throated. These were usually in
separate flocks of from six to eighteen individuals, but now and then I observed
both species feeding together. When a bare patch of ground or a wide trail is
encountered, where they are suspicious of danger, the pheasants cross it by a quick
dash, the laughing thrushes by a single scaling flight.

After drinking at the rivulet or pools in the ravine bottoms, by which time it
is almost dusk, the birds fly up into trees to roost for the night. I have never
actually seen this, being defeated either by the sudden descent of darkness or the
wary scouting of the birds before they retire. But again and again I have heard
the heavy, hollow beating against air and leaves as the birds reach a first and often
a second branch before they settle down, after mumbling a few smothered, low notes.
By aid of strong moonlight I have seen them sound asleep, seldom alone, but two
or three close together, a few feet from the trunk, head under wing.

The few crickets whose instruments were not yet silenced by the chill of the
autumn night still shrilled faintly; small owls hawked about after droning beetles ;
a podargus fanned my cheek like the ghost of a bird, and far off in the blackness,
toward the wild Chinese mountains, came the moan of a leopard. As I turned home-
ward, a wind—first prophet of to-morrow’s storm—rattled the bamboos, drawing
forth weird sounds which seemed to verify the Kachin’s belief in the spirit mats
which wander along every trail at night searching for evil to do. For this reason
these wild hillmen will never travel at night, and as I trudged toward camp from
the sleeping pheasants, I knew that whatever dangers the darkness hid at that hour,
it was from animal not human foes,

88 A MONOGRAPH OF THE PHEASANTS

GENERAL ACCOUNT

After finding and studying the Himalayan kaleege, I later visited Burma, and there
in the south first met with /eatus. My next studies took me to Mandalay, eastward
beyond Maymyo, and northward toward the Ruby Mines District and Bhamo. I then
spent considerable time in the Myitkyina District far up the Irrawaddy, and among the
Shan Mountains to the east and south, to beyond the Yunnan border. Here, thanks to
the exact details given me by Major Nisbett, I was able to collect and observe the birds
at the very localities where the types were collected.

From this latter very limited locality six forms of Gewuaeus had been recorded. Of
these I found no perfectly typical specimens, but pheasants which approximated four,
besides a fifth which was not supposed to occur here. In addition I collected birds
which, if judged by the characters used in separating the already named forms, would
supply at least four additional species.

The species supposed to inhabit this region are the following :—

1. affinis. (1 3 from the River Namli; two thousand feet; east of Myitkyina.)

2. granti. (1 $ from Puntum; six thousand feet; eight miles east of Sadon.)

3. nisbetté:, (An imperfect ¢ from Mt. Kachin ; twenty-five hundred feet ; five miles east
of Sadon.)

4. cliffordt. (The district in general.)

. batemant. (The district in general.)

. horsfieldi. (The district in general.)

OV U1

At the locality from which afzuzs is recorded, the lower River Namli, I found only
horsfieldi, which was dominant and almost typical, and a vermiculated bird associated
with it which closely resembles the description of odscurus, but differed from it as
much as do the most nearly related named species. The two forms were living in close
association, the first flock containing three birds, all males, two of which were Lorsfielad.
In fact this was about the numerical percentage of the latter in this vicinity. On two
occasions I saw a distinctly lighter bird in company with orsfieldi, but I failed to secure
it. My next stop was at Pungatong, some twenty miles farther east, at an elevation of
about four thousand feet. Here I located a flock of eleven kaleege, or what were probably
two families of four and five respectively, and two single cock birds. Each afternoon
these would unite and take the same route to water, down a gently sloping hillside
covered with light forest, across the trail and on down to the stream at the bottom of a
rather steep ravine. I watched these birds carefully day after day until I could actually
recognize the individuals, in spite of the fact that they were usually well looked after by
a mob of laughing thrushes. Then I began shooting, and secured no fewer than eight
birds out of the eleven. This comprised all of one family of two parents and two almost
full-grown young males in first year plumage, the adult male parent of the second family
with a young female of the year, and both of the solitary cock birds—fully adult. The
tale of individuals was as follows, using Oates’ nomenclature.

Family Number One :—
1. ApuLT Marz.—On the whole, this bird resembled Zorsfeld, but it had the narrow
rump fringe and lengthened tail of datemanz, the intense blue, non-vermiculated

PHOTOGRAVURE 21

HOME OF RIPPON’S SILVER AND OF EASTERN HYBRID KALEEGE

Turse of these rare hybrids crossed this trail one day, headed into a wild, deep gorge. The mountains
rose high on all sides, except to the northward, where the purple distance ended in the jagged ranges of

the unexplored tri-corner of Tibet, Yunnan and Burma.
Parrakeets and flycatchers screamed, and at last I heard the tremulous wing-whirr of a pheasant.

But it was an hour later before the timid birds appeared—clad in ebony and white, walking slowly

downward, on their way to the water at the bottom of the gorge.

1s HAUVAADOTOHT

hey *,- vel

-Ansenod e

(lwole gaidisw plist

HOM]

4

RIPPON’S SILVER ANI

PHOTOGRAVURE PLATE 2

[=

WILD KALEEGE HYBRIDS 89

rump of mearsz, and, in addition, the scarlet legs and feet of uzsbett:. The
central tail-feathers were sparingly but strongly vermiculated with white for
most of their length, the terminal fifth being clear black.

2. ADULT FEMALE.—This bird, which was constantly associated with the above cock (in
fact, I secured both with one shot), was not distinguishable from females of
horsfield, except for the somewhat longer tail. Well grown as were the two
young birds, I saw this female more than once allow them to take food away
from her, and from this and other actions, besides the constant association of
the four birds before they joined the others for the afternoon’s descent to water,
there is not the slightest doubt that this was a single family of kaleege.

3. Frrst YEAR Mare.—Typical Aorsfeldi as found in Manipur, with tail of normal
length, but with the basal vermiculation on the inner rectrices visible for a half-
inch beyond the upper tail-coverts.

4. First YEAR Mare.—Between odscurus and davisonz, with characters of each
form.

The two following birds I judged to be parent and offspring :—

5. ADULT MALE.—Superficially close to “nxeatus, with somewhat coarser vermiculations,
and with the outer webs of the outer tail-feathers almost unmarked black, as in
andersout. The central tail-feathers were vermiculated throughout, with no
hint of a pure white area. There was no trace of a rump fringe.

6. First YEAR MaLe.—This bird does not correspond to the description of the females
of any form. The general colour above is olive brown, very minutely vermicu-
lated with black, except along the margins of the feathers, where, the black dying
out, the pure olive brown shows paler and clearer. The primaries are brownish
black, densely mottled with greyish-brown on the outer webs. The secondaries
are similar, with the colour of the outer webs changing into that of the coverts
and remainder of the upper plumage.

The chin and throat are white, tinged with brown along the margins. On the
under parts the light colour persists as a well-defined buffy shaft-stripe. The
rest of the feather is clear olive-brown, with but little mottling except on the
centre of the lower breast and abdomen, where the entire feather is irregularly
blackened especially along the enlarged shaft-stripe. The under tail-coverts
are the blackest of all the contour feathers, most of them showing brown only
along the margin. The outer and median tail-feathers are chestnut, obscurely
but coarsely mottled with black. Toward the central pairs the chestnut changes
to a brown, and the darker colour becomes a coarse vermiculation, the lines
irregular and lying obliquely to the shaft.

The facial skin was scarlet ; the irides light hazel; legs and feet pale, neutral, leaden
grey.

I have gone into this in detail to show what variation I found among the females of
this group of pheasants.

7. ApuLT MaLe.—One of the solitary males was Aorsfe/d: in length of tail and general
markings, except that there was more vermiculation on the central tail-feathers
than I have seen in any bird from Assam, and the inner wing-coverts were

strongly margined.
VOL. I N

go A MONOGRAPH OF THE PHEASANTS

8. ApuLT Mare.—The second male I shot on sight, and at such close range that I
nearly blew it to pieces. At first glance it seemed to be a silver pheasant
(xycthemerus), and stood out sharply from all its companions on both the
occasions when I watched it working downhill. The second time I had my
gun and secured it at once. It proved to be a very dark representative of what
has been called »pponz, but differing in having greenish, instead of scarlet legs
and feet.

All this astounding variety of kaleege I found within two miles of the dak bungalow
at Pongatong, associating together, and, as I have said, with satisfactory evidence of
being in families. Even if the birds were of no immediate relationship, the fact of their
remarkable variation is none the less indicative of hybridism. This is typical of what I
found to exist in other parts of Burma. The Arakan country and Annam I did not
visit, and hence I cannot speak at first hand with regard to the birds which inhabit those
regions.

All the specimens which I gathered in northern Burma tend to exhibit this
individual variation and blending of character, and in all my observations there is
nothing to show any pronounced uniformity in the forms I have mentioned. But while
thus being compelled to consider these as unworthy of specific validity, there is an
interesting phase of the subject in regard to certain of the other forms. Some of these
pheasants, which apparently owe their peculiar colour and pattern to the crossing of two
feral species, seem to have found more or less isolated regions where they have become
more or less established. It is difficult to know exactly how to treat these, and in my
first review of this genus (“ Zoologica,” Review of the Genus Geznaeus) I admitted
tentatively four forms or sub-species-or fixed hybrids. But, as I have already said, more
thorough examination of all the specimens available has shown me that it is a case of
degree, not of kind, and as long as specimens typical of one of the three main species
lineatus, nycthemerus and horsfieldi have been recorded from the heart of the inter-
vening areas containing the hybrids, I see no reason to consider any of them as worthy
of a name.

The sphere of influence existing between Aorsfieldi on the north-west and /ineatus
on the south-east is populated by an interesting series of hybrids, which, except in the
case of the irregular mountain chains and cross valleys, show a fairly gradual gradation
from the one to the other parent species.

I have chosen Cuvier’s kaleege as the wild pheasant hybrid representing one of the
first links in this chain of cross-breeding between true orsfieldi and fineatus. In shape
and in general colouring it is similar to orsfield, being mostly bluish black with
a white rump-fringe. But the feathers of the upper body-plumage, wings and tail are
all finely and regularly pencilled with wavy white lines. ‘The markings are like those of
the Chinese silver kaleege reversed. The female resembles the female of Lorsfield, but
the tail feathers are dominately rufous, mottled with black, the outer pairs being black
toward the tips, and elsewhere pencilled with fine white lines. The average would seem
to be 75% of horsfieldi, and 25% of Lineatus blood. Specimens more or less like this
are found scattered through the middle and northern Arakan Hills, extending into

Chittagong.

PHOTOGRAVURE 22

HOME OF THE WESTERN HYBRID KALEEGE

In western Burma, where the ranges of Horsfield’s and the Lineated Kaleege approach or touch, the
jungles and lesser growth are inhabited by pheasants which show all degrees of intergradation. Their
voice, habits, eggs and young are all more or less alike, but on the plumage of the adult birds is written

their blood relationship to one or the other of their parents or more remote ancestors.

ari. auod. | BogoIgqs

sisdT .noitsbsrsiaini .
neitire ei ebrid alubs sri Yo 9

PHOTOG RAVURE

“HSaaTva4 ClIadgAn

TS GIANT “GLE, EEO

t

AW

WILD KALEEGE HYBRIDS QI

The first mention of this form appears in the ‘Calcutta Journal of Natural History”
in 1842, where it was described as a hybrid between phastanus fasciatus and P. leucomelanos.
Temminck, in the first edition of his Plauches Coloriees, distinguished it by the name of
lophophorus cuvieri. Shortly after my tentative acceptance of this form as a sub-species
in my 1914 review of the group, I realized that it was only another of the unusual types
of wild hybrids, and Baker has independently come to the same conclusion.

Passing over the wz//amsz stage we come, as we go southward, to the forms of
which that named vafsez may be taken as an average. This hybrid may be said to show
25% of horsfieldi and 75% of “meatus, and centres around the southern Arakan ranges.
The white rump fringe is faint, but still distinct, and the dorsal vermiculations are only
slightly fainter than those of /ézeafus. The outer webs of the central rectrices have
got rid of much of their dark pigment, and are quite white. The female in both colour
and pattern is thoroughly correlated with the stage of the male.

Ogilvie-Grant first recognized this as a sub-species in 1893, and Baker still clings
to it. But in one specimen of oafsez which I secured, there is the bilateral asymmetry
which indicates hybridism, and /meatus has been shot farther north than any oafse7 thus
far discovered. Less than a dozen specimens have been found, and they vary enough
inter se to remove all doubt as to the true character of this one, among many forms.

RIPPONI-SHARPEI-NYCTHEMERUS HYBRIDS IN THE FIELD

My first meeting with any true silver kaleege was, as I have related, in Burma,
some distance from the Yunnan frontier, when from a flock of much darker birds
I shot one which belonged in the nxycthemerus-ripponi-rujipes group, but which was
not wholly typical of any good species or named hybrid. After crossing the Yunnan
border, when the last outlying English fort had been left behind, I found traces of an
even whiter bird. These were in the form of stray feathers in the sprung dead-falls
of the Kachin trappers. They belonged to birds which had been caught, but discovered
and eaten by civet cats before the hillmen had had opportunity to go their rounds.

Later I saw a flock of these birds and shot one, which was typical r7pfouz. On
this occasion I was part way down the slope of a wild gorge, on a late afternoon in
December. The mountains rose high on all sides except to the northward, where the
long slopes dove-tailed one another, punctuated by two majestic peaks, and ending
in the purple distance in the jagged ranges of the unexplored tri-corner of Tibet,
Yunnan, and Burma. No huts were in sight, but from a side valley came faintly the
weird, nameless mouthings of a tongueless Kachin cur.

The hillside jungle showed a dozen shades of green, with here and there a blur of
delicate pink, marking the unseasonable blossoming of a wild cherry tree. One close
overhead radiated a hundred sprays of coral bloom, all ahum with insects drawn by
the nectar, while the branches themselves trembled with the perchings of many fly-
catchers, attracted in turn by the insects.

Now a hoarse chorus came from a dense flock of parrakeets, and soon afterward
the tremulous wing whirr of a kaleege. This sent me down at once on my knees
among the scanty dwarf bamboo and everlastings, alert for another hint of the birds
which I was seeking. A crash of leaves and twigs drew my attention to the right along

Q2 A MONOGRAPH OF THE PHEASANTS

the slope, and I grasped my three-barrelled gun. A brace of tigers had taken toll of
mules not far away, but this alarm ended in silence and mystery.

An hour passed with no sound of pheasants to reward my patience. Then a cloud
blotted the sun, and a chill followed the warmth and stilled the humming insects.
A wind rose, and even when the long, slanting rays at last shone forth, the cold of night
still filled the air. I crept slowly to a projecting spur, and found the wind blowing
freshly on the other side. The moment it blew against my face I heard, a short distance
away, the scratching which always meant pheasants of some kind. Twice a bird whirred,
and then, walking slowly downward, on the way to the water at the bottom of the gorge,
there appeared a stately kaleege, clad in ebony and silvery white. This was a time
when I wanted the bird more than any hint of its life-history which observation in the
failing light might give, so I raised my gun and fired. Two more birds burst forth
from the feathery bamboo, and on wide-spread vibrating wings flew and finally scaled
down the great gorge, until their white forms were swallowed up in the darkness far
below. Only a few weeks before, I had watched the blackest of all the kaleege not far
from the banks of the Irrawaddy, and here were the outposts of the silver clan, which
stretched on and on to the eastward until fairly stopped by the shores of the Pacific.

This ended my personal experience with this form. I had known that it (+ jomesz)
was believed to range widely over Yunnan and the Shan country. Not, however, until
I had access to an unexpected and unusually large amount of material was their wide
distribution proved beyond doubt. For a year or more a Chinaman had assiduously
collected silver kaleege pheasants in various parts of Yunnan and the Northern Shan
States, and when he had gathered six large bales, he boxed them up, labelled them
‘ducks’ feathers,” and shipped them via Bhamo to Rangoon, en route to the milliners of
Europe. But the Custom officials at Rangoon, having had previous experience with
Chinamen and with ducks’ feathers, investigated, and in place of the feathers of domestic
ducks, found hundreds of skins of silver kaleege, with a scattering of Lady Amherst and
Burmese bar-tailed pheasants. The bales were promptly confiscated and condemned,
and at the moment when awaiting destruction I was fortunate enough to come across
the great mass of skins. I began at once to set official machinery in motion, and with
the help of a very amiable collector of customs and Dr. Annandale of the Indian
Museum, the entire lot was turned over to me. I spent considerable time in studying
the fragments, and later the best skins were picked out and sent to me.

I found that in one bale about twenty-five per cent. were pure zycthemerus, while
sixty per cent. were equally typical ~zffonz, the remaining fifteen per cent. showing
intermediate grades between the two. Later I compared this lot with several ~pponz
skins and found them about identical, though fluctuating slightly in the direction of
whiter zycthemerus, or with the blacker shades of the so-called vajfipes.

Since the above was written I have examined the specimens in the other bales, with
still more interesting and significant results, About thirty-five per cent. were xycthe-
merus, fifteen per cent. might be rufifes, another fifteen were sharper, twenty per cent.
approached vipponz, five per cent. were close to Aorsfieldi, while the remainder were
anomalous.

I have presented considerable data based on observations in the field to show how
variable were birds even in the same flock in certain parts of Northern Burma. I do

RIPPON’S HYBRID SILVER KALEEGE

thas be ay given a name (Gennaeus ripponi), but no two birds are alike, and there is no
it represents individuals which have a very large percentage of the blood of the Silver
tively small amount of Black-breasted ancestry. I found a typical bird in a flock of
Burma. | They showed great variation, and could not be classified with any specific
nly one approached the description of this hybrid form. 7 ;

ae :

“sr

¥ + bong J

PLATE XXXIl.

RIPPON’S HYBRID SILVER KALEEGE.

WILD KALEEGE HYBRIDS 93

not intend to go into similar detail in regard to all the kaleege which have received
names, and one instance must suffice. I have taken a series of specimens in the vajfifes-
sharpet group, the chief hybrid buffer forms occurring between /zweatus on the west
and zycthemerus on the east, and examined them carefully with a view to recording the
variation, not only in the adult birds, but in immature pheasants, in juvenile and first
year plumage. This series of some thirty-five birds is chiefly from my own collection
and from the Tring Museum. The localities show that the variation is absolutely not
dependent on environment, some of the extremes coming from the Ruby Mines district,
while equally variable specimens were collected far to the southward.

DETAILED DESCRIPTION OF KALEEGE OF THE RUFIPES-SHARPEI GROUP, RANGING
FROM MOGOK IN THE RUBY MINES DISTRICT TO PAZAUNG IN THE SOUTHERN
SELACING aS ICASE ETS:

Apu_t Marre.—Facial skin well developed, scarlet; forehead, chin and throat dull
black; long flowing crest shining steel blue; longest feathers 80 mm. Entire ventral
surface dead black down the middle of breast and belly; the flanks, thighs, and under
tail-coverts with wide, shining, steel-blue margins. Ear coverts white, spotted and
indistinctly banded with black.

Feathers of nape immediately behind crest white, with an excess of black in the
form of a wide, elongated shaft-patch, a concentric line, and a narrow margin. On the
feathers of the anterior upper neck the black decreases in amount, but has the same
pattern. As we go back, the number of black lines increases. The method of this
increase is by the expansion of the shaft-spot into an elongated kite-shaped band, when
at once another spot of black appears in the centre, which a few rows posteriorly will in
turn expand into a more or less circular band and give rise on the shaft at its centre to
the anlage of the succeeding black line.

By this process of increase we find on the mantle five concentric lines on each web
(one marginal), each pair joining on the shaft, besides a narrow shaft-streak, within the
inner or fifth. There is no suggestion of convergence of the lines proximally, as
they run straight backward, and disappear at once in the monochrome grey of the
basal down.

On the midback a sixth line has appeared, there being six well-developed concentric
pairs, and a seventh or even eighth on the rump (always counting the narrow black
marginal fringe as a true line).

On the median and longer tail-coverts we find a new method of linear formation
in progress, namely, the insertion of new lines between the old, appearing first as mere
mottled streaks, and, in addition, the coalescing of lines near the posterior margins, the
lines now lying obliquely to the shaft and not parallel, the outermost. (as well as the
marginal) extending unbroken the entire length of the feather, but the inner ones dying
out or coalescing with others.

On the central rectrices we find this pattern carried at once to an extreme, but with
such variation that the feathers may be said to be black, banded obliquely with white.
The amount of white increases from the outer feathers inward, and in some specimens
the inner webs of the central rectrices are almost wholly white.

The length of the tail is extremely variable, and I have recorded the length of the

94 A MONOGRAPH OF THE PHEASANTS

full-grown, central tail-feathers of two adult males shot on the same day, as 380 and
475 mm. respectively.

A much simpler pattern than that of the mantle and back is seen on the wing-
coverts. On the lesser and median feathers we find two concentric rings and a black
margin, the black lines on the latter being very wide. But the greater coverts show a
sudden change to oblique or almost transverse cross barring.

On the scapulars there are four or five concentric black bands, but quite suddenly,
on the innermost secondary, we find the transition to oblique cross-barring, and with
this, strong, dark, linear mottling in the narrow white interspaces, as if hinting of linear
increase as in the rectrices. Thus on the outer secondary we find thirteen or fourteen
broad black cross-bars separated by narrow interspaces, all of which are split by a
mottled dark line. The primaries show so little white that one must reverse the colour
value in describing the pattern. The ground-colour is not black, however, but dark
brown, across which numerous narrow, white, oblique bands extend on both webs, each
white band split by a mottling or solid bar of brown.

The above description represents one type of pattern. Another, equally common,
shows no dark bar splitting up the cross white bars. A third shows the primary
markings so numerous, broken, fine, and wavy that they are rather vermiculations than
bars, and defy counting. Bill from nostril, 20 mm. ; wing, 240 to 280; tail, 360 to 530;
tarsus, 90; middle toe and claw, 70; spurs, 28 mm.

IMMATURE Ma.es.—A young bird which shows no adult characters is one of the
severest and plainest of birds. Forehead and crown brown, unmarked. Occipital
feathers rather elongated into a short crest. Feathers of neck all around show a large,
central, pale buff area with numerous wavy cross-bars.

Posteriorly on the mantle, back, rump and wings, this central area merges again
with the rest of the feather. The ground-colour is a rufous buff, vermiculated with
oblique, fine, wavy black lines. The secondaries are plain dark brown on most of the
inner web, and the primaries are wholly brown. The delayed ninth and tenth primaries
show irregular white wavy streaks on the inner web, the only hint of the adult plumage.
The other rectrices are very irregularly marked with rufous mottling on dark brown.
The lower throat and breast show distinct traces of the central lighter area, but all the
remaining under surface is identical with the back. |

This is the normal plumage of this moult, but very. few individuals show it in.
perfection. In fact, no two young males are alike, and when the moult has entirely
ceased we find all kinds and conditions of infusion of adult patterns and pigment,
grafted upon correspondingly more or less strongly marked juvenile plumage.

The adult character which shows the most pronounced precocity is the black ventral
surface, appearing usually as a broad band from the throat to the under tail-coverts.
The feathers are as often parti-coloured as black, the brown pigment manifesting itself
most frequently as a mottled shaft-stripe and a wide margin. With this character we
find correlated a considerable development of the steel blue crest, these feathers being
sometimes quite pure, without admixture of brown, this latter colour, when it occurs,
being confined to the terminal third of the feathers. The elongated juvenile crest
growing side by side detracts from the adult appearance of the flowing blue feathers.

OATES'S HYBRID KALEEGE

rd, which some ee have considered to be a species (Genndews oatest), shows great
> dozen skins I have examined, We must either give a half-dozen new names to them
“th m as hybrids. Oates’s Kaleege represents birds with the blood beg of the Black-breasted

ed Kaleege, fe the latter greatly dominant.

iso¥g eworle.,(sisno p ammenin) + esinage By a on 1 borobignan ors | etdglein ‘gmoz oid:

month

PLATE XXAll.

OATES’S HYBRID KALEEGE.

WILD KALEEGE HYBRIDS 95

The wings are quite conservative and are usually wholly juvenile. A slight advance
is sometimes seen in the form of a well-marked sub-terminal band, extending entirely
around the visible margin, with a terminal band of buff. This gives the wing a very
elegant appearance, the monochrome of vermiculated rufous brown broken by the
distinct outlining of each feather. In such a wing the adventitious growth of a second-
ary presents a startling intrusion of black and white, developed, curiously enough, only
on the visible portion, the remainder of the feather being quite juvenile.

The tail is perhaps the most variable factor of all, and hardly any two birds have it
alike. The central rectrices in particular vary from warm buff, everywhere thickly
vermiculated with wavy, dark brown lines, to the other extreme, which is pure white,
with thick, wavy, oblique, black lines on the outer web, the inner being white except for
a few linear mottlings near the shaft.

Whatever the condition of the inner rectrices, the outer are invariably more juvenile,
more tinged with brown, the outer pair being sometimes quite rufous and brown, thus
reflecting vividly the gradual moult from the outside inward.

Two or three immature males which have acquired adult tails show as much
variation as exists in fully adult birds; from an excess of black banding over all the
feathers to the excess of pure white over the inner webs of the central pairs. So this is
wholly zzdividual variation.

In birds acquiring the full adult moult, the numbers of concentric black lines on
the dorsal plumage, in number and variability, correspond exactly with those in the
adult bird.

The young birds show all shades of feet and legs, from pale brown through flesh
colour to deep rich scarlet.

ApuLT FEMALE.—Upper parts brownish buff, everywhere very faintly mottled with
darker. Feathers of rear crown and occiput rather more mottled and elongated into a
crest. Wings the same, becoming a shade more rufous on the secondaries. Primaries
plain dark brown on the inner webs. Rectrices present considerable variation, some
with the lateral feathers showing much more contrasted markings than others. In the
more common type the central rectrices have a ground-colour of pale buff, finely
vermiculated with datk brown, while as we proceed outward, the feathers increase in
intensity of rufous colouring, and the markings become less alike on the two webs,
the outer webs being mottled, the inner obliquely banded with pale buff, these markings
being indistinctly outlined in black.

Chin and throat white, the feathers posteriorly with a rapidly increasing central
spot and margin of brown, until on the breast the white is reduced to one or two con-
centric rings. This pattern, but with considerable variation, covers the entire ventral
surface. From two to four buffy-white and dark-brown concentric rings or oblique
cross-bars show all variations, from a rarer pattern of very regular, concentric markings
to the more usual appearance of broken, spotted, irregular cross-bands. This is wholly
individual.

In well-authenticated skins I find none of the radically characteristic homogeneous
types of females which have been accredited to these and related so-called species of the
group. The females of typical sarpez near the Ruby Mines district are most assuredly

96 A MONOGRAPH OF THE PHEASANTS

not like “eatus, and of those from farther south I have never seen two quite alike. Bill
from nostril, 20 mm.; wing, 250; tail, 268; tarsus, 85; middle toe and claw, 63 mm.
Spurs sharp, but very low nodules.

In conclusion, the kaleege pheasants forming the genus Genuaeus seem to me to
warrant the following disposition :—

FULL SPECIES
lineatus
horsfieldi
melanonotus
leucomelanos
albocristatus
nycthemerus
whtiteheadt

HYBRIDS

affines

_andersont

annamensis
assimilis
atlayt
batemant
belt

edwardst clifford

swinhoet cuveere
davisont
elegans
grant
havingtont
Jonest-rippont
macdonaldt
MLCAYSL
misbettt
oatest
obscurus
ommansyt
prendergasti
rippont
rujipes
sharpet
wickhamet
qwelliamst

SUMMARY OF NAMED HYBRID GENNAEUS

afinis Oates, Ann. Mag. Nat. Hist. (7), XI. 1903, p. 231; single male killed by Major Nisbett, river Namli
east of Myitkyina.

andersont Elliot, Proc. Zool. Soc. London, 1871, p. 137; type in Indian Museum, Calcutta; several
specimens; Anderson in Yunnan; Rippon at Warar Burn, 6000 feet, in Kachin Mountains, 30 miles east
of Bhamo.

annamensis Grant, Bull. Brit. Orn. Club, XIX. 1907, p. 13; type in British Museum; three specimens ;
Vassal, Bali region, interior of Annam.

assimilis Oates, Ann. Mag. Nat. Hist. (7), XIV. 1904, p. 286; types in Oates Collection, British Museum ;
six specimens, Ruby Mines District.

atlayt Oates, Ann. Mag. Nat. Hist. (8), V. 1910, p. 162; types in Oates Collection, British Museum; seven
specimens, Atlay, Ruby Mines District.

batemani Oates, Jour. Bombay Nat. Hist. Soc.; types in Oates Collection, British Museum ; eight specimens,
Districts of Katha, Myitkyina and Bhamo.

beli Oustalet, Bull. Mus. Nat. Hist. Paris, 1898, p. 258; types in Paris Museum; several specimens, Bel,
Eastern Annam between Hue and the mountains.

clifford: Oates, Ann. Mag. Nat. Hist. (7), XIV. 1904, p. 286; types in Oates Collection, British Museum ;
six specimens, Myitkyina District, east of Irrawaddy.

cuviert Temm., Planches Coloriees, V. pl. 10, 1820 ; about a dozen specimens; the hills of northern Arakan
and irregularly to the north-east.

davisont Grant, Cat. Birds British Museum, XXII. 1893, p. 304; type male in British Museum; type
female in Oates Collection, British Museum ; about ten specimens, vicinity of Bhamo and in Yunnan.

elegans, listed in Oates Collection.

granti Oates, Ann. Mag. Nat. Hist. (8), V. 1910, p. 163; type in British Museum; single male; Nisbett,
Puntum, east of Sadon.

WILD KALEEGE HYBRIDS 97

haringtoni Oates, Ann. Mag. Nat. Hist. (8), V. 1910, p. 162; types in Oates Collection, British Museum ;
single pair ; Harington, Milum Kha, Bhamo District.

jonest Oates (rippont), Ibis, 1903, p. 97; types in British Museum ; distribution same as ripponz.

macdonald: Oates, Jour. Bombay Nat. Hist. Soc., XVII. 1906, p. 10; type male in Oates Collection, British
Museum ; specimens from Chin Mountains, from Mt. Victoria to Fort White.

mearst Oates, Ann. Mag. Nat. Hist. (8), V. 1910, p. 164; type male in Oates Collection, British Museum ;
several ; Mears from Sylhet; Bateman from Kamaing, Myitkyina District ; also from North Khasi Hills, Manipur,
Tippera, and Goalpara in Assam.

nisbettt Oates, Ibis, 1903, p. 99; type male in British Museum; incomplete skin of male; Nisbet, five miles
east of Sadon, 25,000 feet.

oatset Ogilvie-Grant, Cat. Birds British Museum, XXII, 1893, p. 306; types in British Museum; eight
specimens ; central and southern hills of Arakan.

obscurus Oates, Ann. Mag. Nat. Hist. (7), XIV. 1904, p. 283; type female in Oates Collection, British
Museum; specimens from Katha District.

ommansyt, listed in Oates Collection.

prendergastt Oates, Jour. Bombay Nat. Hist. Soc., XVII. 1906, p. 10; specimens from North Arracan.

rippont Sharpe, Bull. Brit. Orn. Club, XIII. 1902, p. 29; fifteen specimens, Northern and Southern Shan
States and Yunnan.

rufipes Oates, Manual Game-birds India, Part II. 1899, p. 495; type female in Oates Collection, British
Museum; specimens from the Chin Mountains.

sharpec Oates, Manual Game-birds India, Part I. 1898, p. 357; type in British Museum; seven, Southern
Shan States and Salwin country to the north of the river.

wickhamt Oates, Manual Game-birds India, Part II. 1899, p. 495 ; type female in Oates Collection, British
Museum ; specimens from the Chin Mountains.

welliamst Oates, Manual Game-birds India, Part I. 1898, p. 342; over thirty specimens in Oates Collection ;
between Chin Mountains and the Irrawaddy; Williams, from Kalewa; others at Chindwin, near Wuntho,

VOL. II ; re)

ACOMUS

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MAP SHOWING THE DISTRIBUTION OF THE CRESTLESS FIREBACK PHEASANTS.

Region 1. Acomus erythrophthalmus.
si Be a pyronotus.

ACOMUS
CRESTLESS FIREBACK PHEASANTS

Family PHASIANIDAE
Subfamily PHASIANINAE

Genus ACOMUS

Tus small genus of pheasants is unquestionably rather closely related to the
crested firebacks (Lophura) and more distantly to the silver pheasants (Gennaeus),
but instead of lumping all three, and thereby relegating their differences to the obscurity
of subgeneric rank, I consider that the groups are sufficiently distinct to warrant giving
to them full generic value.

The chief point of difference between comus: and Lophura is the extremely
masculine pattern and colour of the plumage of the females in the former genus, together
with the less important total absence of crest in both sexes and the very small size.

The side of the head is bare and covered with fine papillae ; the wattles are very small,
but the bill is of moderate size; the tarsus is extremely long, its length relative to that
of the middle toe and claw being 3:2; the primary formula shows the 1st much shorter
than the 2nd, which is about the length of the 1oth, the 6th being slightly the longest.
The tail is composed of fourteen feathers, quite short and compressed laterally, like that
of a domestic hen. The third pair is the longest, the outer being considerably ‘the
shortest of the seven. Spurs are present in both sexes. The general colour is bluish-
black, the males having white vermiculations or streaks, highly iridescent back and
rump, and buff tail-feathers.

The Crestless Firebacks number two well-known distinct species and a third
extremely doubtful one. For reasons given elsewhere I shall not recognize this latter
form in my treatment of the genus. They are confined to the southern part of the
Malay Peninsula, Sumatra and Borneo.

Malayan Crestless Fireback : : : : : Acomus erythrophthalmus (Raffles).
Bornean Crestless Fireback : : : : Acomus pyronotus (Gray).
KEY TO ACOMUS

I, Plumage not uniform bluish-black (males)

(a) Breast almost uniformly dark. : : : : ‘ : erythrophthalmus.
() Breast with conspicuous white shaft streaks : s . — pyronotus.

(il. Pappagecmieny sissy GueGa. Gun sane Leen { erythrophihalnus.
pyronotus.

(Pyronotus females average slightly larger, and show somewhat stronger bluish gloss, but variation is too

great to render these characters of diagnostic value.)
IOI

MALAYAN CRESTLESS FIREBACK
Acomus erythrophthalmus (Raffles)

NAMES.—Generic: Acomus, dxopos, hairless, referring to the absence of a crest. Specific: evythrophthalmus,
eov0odc, eruthros, reddish ; dp#aludc, opthalmos, eye ; with reference to the red facial skin. English: Malayan
Crestless Fireback, Rufous-tailed Pheasant; French: Faisan a queue rouge; German: Gelbschwanziges
Fasanhuhn; Native: Burong mérah mata (Red-eyed Bird); Kuang bertam or Mata mérah (Malay).

BRIEF DESCRIPTION.—Male: Head dark brown ; lower back bronze-gold, changing posteriorly into bronze-
red ; tail buff; remaining plumage steel-blue, freckled on the back, sides and wings with white; face scarlet.
Female: Head dark brown; entire plumage, wings and tail shining steel-blue ; flight feathers and belly dull
brownish-black.

RANGE.—Southern part of the Malay Peninsula and Sumatra.

GENERAL DISTRIBUTION

Tue Malayan Crestless Fireback inhabits the southern part of the Malay Peninsula.
It has been recorded as far north as Kedah (6° N.), but is apparently abundant neither
there nor in Perak. As we go southward we find more and more evidence of its
occurrence, especially in the low country of the west coast, throughout Selangor, Negri,
Sembilan, Malacca and Johore. I found no trace of this species in any of the central
mountains or elevated regions of the peninsula, nor on the east coast, although indeed
much of the latter region is still unexplored. My studies of the bird were made chiefly
in northern Johore.

The locality “Java” is an error, and no confidence can be placed in the old skin
labels ‘‘ Singapore” and ‘‘Penang.” These were the first settled parts, and in those days
stood for the whole region, just as the word “India” meant anything from Ceylon to
Tibet. There is no foundation for belief that this species ever inhabited either of these
islands. In Sumatra it is found along the eastern coast, but does not extend into the
mountains of the interior.

GENERAL ACCOUNT

A careful search through ornithological literature shows that our ignorance of the
life and habits of this bird is complete. Nota single fact has been recorded, and as far
as I know, only native Malays have heretofore had the good fortune to catch even
a glimpse of them in their native jungles. I went to Johore expecting to find this
species the commonest of all the pheasants of that region, always, of course, excepting
the junglefowl. But I found that while they shared with the latter the habit of
frequenting the vicinity of native villages and the more open jungle where the
half-domesticated buffaloes roamed, they were exceedingly shy, and I actually had
as much difficulty in getting even the barest outline of their life-histories as in the case

of the much rarer argus and bronze-tailed peacock pheasants.
102

PLATE XXXIII

MALAYAN CRESTLESS FIREBACK

Acomus erythrophthalmus (Raffles)

Tues birds are exceedingly pugnacious, and I have even seen hens fighting madly with each other.
Both sexes are armed with spurs. They live along the small creeks which wind through the low jungle,
and haunt the water-holes of the buffalo. The sound of warning or challenge, uttered when two cocks

threaten one another, is a deep, hoarse drawl, almost a snarl, sounding as much like a small cat as a pheasant.

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MALAYAN CRESTLESS FIREBACK.

MALAYAN CRESTLESS FIREBACK 103

My first glimpse of the Malayan Crestless Firebacks was wholly unexpected, and
came while I was completely absorbed in other forms of life.

For three hours or more, since early dawn, I had been creeping on all fours, or
progressing in a still lowlier position, stalking and observing junglefowl. [I had
started inland from the bank of a small river in central Johore, and had become rather
confused and uncertain as to the most direct route back. I knew, however, that
a sight taken from any tall tree would set me right, so I chose a hidden resting-place
in a shady, partly dry spot, beneath a small forest of giant caladium-like growths, and
became absorbed in a pair of tumble-bugs revolving their diminutive globe. They had
gleaned from the sleeping-place of a mouse-deer and had kneaded the clay and
prepared their tunnel. Their methods differed in no way from their kindred all over the
world. Exactly thus had I seen them push and tug on the sand of Egypt before the
tombs of Sakkara; thus had they done before me in a Virginia pasture and a New
England roadside. 1 found later that more than one pheasant had accepted these hard,
horny morsels as food, breaking them in pieces, but swallowing hard elytra and thorax
as well as soft body.

My reverie was suddenly interrupted by a wasp which hovered so close and so
angrily that I left at once. A tropical wasp nearly two inches in length is as dangerous
to health as a strong attack of fever. To my dismay I discovered behind me a herd of
water buffaloes coming slowly along one of their trails. I shifted quickly to a position
between two small trees, either of which I could mount swiftly. A person who has once
been charged by buffaloes puts dignity out of mind at once on their appearance. For ten
or fifteen minutes they grazed near me, fortunately keeping up wind.

Then came a sight for which I was wholly unprepared. Trotting along the trail
came a string of birds—a glance showed they were Crestless Firebacks, twenty-two in all,
about an equal number of males and females. They passed not more than fifty feet
from where I crouched, straight through the herd and on out of sight. This was well
worth the siege of the buffaloes, which I raised by an elaborate circling, and, knowing they
had come from the river, I followed on their back track, and in twenty minutes was at
the camp, eating breakfast. Why so many birds should be together I could not tell,
unless it is the habit for several families to unite after the young birds have reached full
size.

I had one other glimpse of these birds in this locality. I had gone far beyond the
buffalo ground into light jungle. Green, black-faced babblers were everywhere, hanging
to the leaves and twigs in their strenuous search for insects. Long-billed sunbirds
dashed about me, uttering sharp fszs, and a little squeaking on my part drove them to
the highest pitch of excitement. Now and then two or three hornbills passed high
overhead, with their alternate flap and soar like pelicans. One of the most characteristic
sounds of the Malayan jungles is the deep, soft, whoof! whoof! whoof! whoof !
whooooooo000f! whoof! whoof! whoof! whoof! whoooooof! of their great pinions.

A number of gibbons had just finished their morning’s chorus, and were swinging
about the trees just ahead, making a tremendous racket, while at the same time they fed
on some gourd-shaped fruit. Now and then a head would swing up into view, eye me
keenly and hurl a few hoots at me. They kept my advance under most careful
observation. A junglefowl scampered away with a fowl-like, broken cackle of fright.

104 A MONOGRAPH OF THE PHEASANTS

A few yards ahead I heard now and then a muffled scratching of leaves. Fortune
favoured me, and after many minutes of most painstaking stalking, I reached a good
point of observation. Fora long time I could make out nothing but a confused dark
mass tumbling about—first falling over one side, then the other. My low-power glasses
at last showed me it was two Fireback hens waging a battle as fiercely as any
game-cocks,

Breathlessness was the only cause of stopping, and even when both seemed com-
pletely exhausted, they clung together, and from time to time gave wholly ineffectual
pecks at one another. Their wings dragged, their feathers were ruffled, and they were
altogether most disreputable-looking females. Finally, thinking they were so spent that
I could catch them with my hands, I walked toward them. But though their combative
strength was spent, they had reserve enough to run quickly off when they saw me
coming.

The comparative similarity in colour of the sexes of this species, and the presence
of spurs in both, would indicate that courtship was a more or less reciprocal affair, and
the fight of the hen birds which I witnessed would also support this conclusion. On
the other hand, there are good reasons for supposing that this species is more or less
polygamous, and if such is the case, any such theory would fail to explain the facts.

Further observation showed that these birds were more numerous along the smaller
creeks which traversed the low country, and that they haunted the water-holes of the
buffaloes in early morning. They have apparently no regular place of fighting, but pitch
into one another whenever and wherever the spirit moves them.

Birds which become badly injured in these encounters or are otherwise disabled or
weakly have short shrift. Leeches and ticks do their work even before the keen noses of
carnivores find them out. One recently dead bird with a spur-thrust through its breast
I found with a circle of gorged leeches lying about, and the ants were assembling in
hundreds, eager to play their part in the drama of dissolution.

Even continued trapping will not drive these birds away, and a pot-hunter or
trapper could destroy every member of a flock.

The note of suspicion or half-alarm of the Malayan Fireback is a sharp, explosive
kak! followed by the throaty, frog-like gulp, characteristic of all the Gexuaeus, uttered
once or twice. Then the £a#! again, and the subdued bubbling or gurgling sounds.
The note of wild despair when attacked by some animal, or being removed from a trap
by a native, is, in the old bird, a hoarse, long-drawn cry ; in the young a shrill whistling.
The sound of warning or challenge when two cocks threaten one another is a deep,
hoarse drawl, almost a snarl, which one would attribute rather to a carnivore than to
a small pheasant.

The fowl-like appearance which the roof-like, compressed tail gives to this Fireback
is well indicated by the Malayan name. The natives never call it Awang (pheasant),
but always durong (bird) or ayam (fowl).

Although this pheasant is not uncommon in some places, and, as we have seen, even
frequents the vicinity of isolated huts or native hamlets, yet its nest has never been
found, nor, as far as I know, have we any record of its eggs or young. I was unable to
learn anything authentic concerning its nidification, although the natives were ever
ready to furnish me with any information (doubtless made to order) which they thought

PHOTOGRAVURE 25

HAUNTS OF THE MALAYAN AND BORNEAN CRESTLESS FIREBACKS

In both countries these pheasants live in dense jungle, where they have access to open clearings, or
at least the trails made by wild animals. Here they strive to escape their enemies, snakes and civet cats,
which lie in wait or creep silently through the undergrowth. And here they nest, although up to the
present time no white man has seen nest, eggs or young, so timid are they, and so skilful in hiding their

home among the swamps and tangles.

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WH

MALAYAN CRESTLESS FIREBACK 105

would please me. To test the elasticity of such facts I led a Malay headman to assure
me one day that the duvong mérah mata \aid but two eggs and in a tree! while on the
following day, when I expressed disappointment that they deposited so few, he informed
me in a burst of confidence that many of the hens laid five times that number! A female
which I shot well to the southward was about to deposit an egg, while two others were
well developed. The shell was plain buffy-white in colour, with very minute pits.

The birds spend the night well up in trees, and when a native discovers such
a roosting-place he is well-nigh certain of capturing the birds, as he will set scores of
nooses in all the grassy runways leading to the tree, and the habit of returning night
after night to the same place ensures their capture. They seem to feed at all times of the
day, not only early in the morning and in the evening, as with most pheasants. In the
game and buffalo trails they scratch like fowls, making a great noise and rustling of
leaves. At such times, however, unlike the Himalayan pheasants, they seldom utter
a sound, and every moment or two stand erect, motionless and silent, watching,
listening, for the dangers which threaten every inmate of the jungle. It is said that
the females are more numerous than the males, but this I cannot confirm. The sexes
have been approximately equal in the covies or flocks which have come under my
observation.

The food which I have examined in the crops of the birds consists of animal and
vegetable matter in about equal proportions. Termites, as usual, are very frequently
eaten, ticks and grubs less often. Small hard berries, with but a thin skin over
the very large stone, formed the food of two male birds; poor sustenance, one would
think.

They seem to be stupid birds, and I have never seen one in captivity which showed
any relaxation of the shyness or unreasoning wildness of a feral bird. Those which
I sent to America were the only ones of their genus to reach that country. In Europe
they are better known, and some eighteen or twenty individuals have been received at
the London Zoo. These, on an average, have lived about a year and eight months,
although one bird remained in the collection for more than nine years.

DETAILED DESCRIPTION

ApuL_t Mate.—Forehead, crown, chin, throat and ear-coverts (the only cephalic
feathered areas) dull dark brown, the feathers of the occiput often tipped with chestnut
or with a slight whitish freckling. Neck and breast black, glossed with purplish blue.
Mantle, upper back and scapulars similar, but freckled or irregularly banded with wavy
bars of white; the gloss dies out gradually on the wing-coverts and secondaries, but
the freckled barring persists strongly; the primaries are plain, dark brown. Visible
area of lower back, consisting of a wide, terminal, disintegrated fringe, fiery bronze-gold,
basally each feather changing abruptly into a narrow bronze-red and then to a white-
freckled, steel-blue zone. Posteriorly the gold decreases rather suddenly and the
bronze-red takes its place, the basal freckling disappearing at the same time, leaving
the clear steel-blue. The deep metallic red or maroon persists strongly up to the
longest upper tail-coverts, which are wholly dull steel-blue. The tail is uniform pale

rufous-buff, with the basal part of the feathers black. The sides of the body, flanks,
VOL. II P

106 A MONOGRAPH OF THE sPHEASANTS

and under tail-coverts are heavily glossed with blue, and the former show slight traces
of the white freckling. The mid-ventral plumage is mostly dull blackish.

Irides hazel-brown; facial skin vermilion; nape wattle is sometimes fairly well
developed, and in a breeding male the facial shield of skin stands up ridge-like above
the crown feathers; mandibles pale horn, blackish toward base; legs and feet bluish
grey. The single pair of spurs are long, stout and curved, and average 25 mm. in
length. Measurements are, bill from nostril, 17; wing, 245; tail, 162; tarsus, 79;
middle toe and claw, 51 mm.

VARIATIONS OF PLUMAGE

ApuLT Mare.—About fifty per cent. of adult erythvophthalmus males have the
shafts of the ventral plumage, especially on the breast, conspicuously white, thus
approximating the very specialized corresponding feathers of the Bornean pyvonotus.

As regards the status of comus inornatus. In 1879 Salvadori described a bird
from Mount Singalan, on the west coast of Sumatra, as a new species of Crestless
Fireback. Since then a second similar specimen has been obtained. Both are said
on good evidence to be males, and yet curiously enough are almost identical in colour
with the females of the two other species of 4comus, the chief point of distinction
being the scaly appearance of the dorsal surface, owing to the steel-blue colour being
confined chiefly to the distal fringe of the feathers.

Upon examination of many scores of specimens of evythrophthalmus collected by
myself and in various museums, I find so much variation to exist that I cannot bring
myself to accept zzorvafus even as a sub-species. Both Johore and Sumatran specimens
show all gradations between complete gloss and a narrow terminal fringe, and as to
the freckling and iridescent back and rump I have two black-tailed males trapped in
the same line of nooses in north Johore with four perfectly normal males, which show
only the barest trace of white, while the dorsal colour is absent from one and extremely
imperfect in the other. ‘In fact, when skinning the first I sexed the bird merely as
a matter of habit and was most surprised to find that the individual was a male, with
full-developed generative organs, and yet in the plumage of a female. I can offer as
explanation only that this most interesting phenomenon is doubtless correlated with
the very unusual approximation of colour in the sexes of this genus, a character so
unlike their nearest relatives—the crested firebacks (Lopiura). The close association
of the two phases, and their occurrence in both the Malay Peninsula and Sumatra
would seem to indicate that its occurrence is no more than a variation such as the
so-called Lophophorus chambanus in a wild state, or Chrysolophus obscurus or Pavo
nigripennis in captive birds.

ApuLT FrmMaLe.—Forehead, crown and occiput dark brown, chin, throat and
ear-coverts paler, smoky grey. Upper plumage black, strongly glossed with steel-blue
on the exposed portions of the feathers. The gloss dies out on the secondaries and
tail-feathers, which are black, the primaries being brown. The under-parts show very
little gloss, this being confined to the breast, sides and under tail-coverts, the remainder
being dull, brownish black. Many females show a small round white spot or mottled
patch on the feathers of the crown and side of the nape. Correlated with the other

MALAYAN CRESTLESS PIREBACK - 107

masculine characters, such as colour in general and extent of bare facial area, the
spurs are well developed in this sex. They differ from those of the male, however, in
being almost or quite straight, and average 9 mm. in length. The well-developed,
carunculated facial area is blood red; eyes hazel, legs and feet blue-grey. Measurements
are, bill from nostril, 16; wing, 221; tail, 142; tarsus, 71; middle toe and claw,
46 mm.

JUVENILE PLumAGE.—The general colouring of this age recalls the corresponding
plumage of Lofiura, and indeed of the females of that genus; black and varying shades
of chestnut and rufous, the latter colour being confined more or less to the terminal
parts of the feathers in the form of mottling or bands. In the succeeding moult this
is eliminated from the plumage, except on the back and rump of the male, where it
increases in area and assumes the fiery iridescence of those regions. The chin and
upper throat are pure white, and the face is thinly feathered, and dull pinkish
in hue.

First YEAR Matse.—The species attains the general adult colouring at the first
annual moult, but just as the breeding season varies so widely in various localities,
so the condition of the bird at this moult shows unusual variation. In a typical male
bird of four or five months of age the spurs are already 7 mm. in length. The papille
on the bare facial area are scarcely apparent, being represented by rectangular scales
or flattened nodules. There is a scanty growth of very short, minute, but perfectly
formed black feathers over this area. On the body plumage as a whole there is
considerably less steel-blue gloss than in the adult, the under-parts being almost wholly
plain dull brown. ‘The outer primaries are delayed, and show considerable buff mottling
along the margins. The feathers of the back are very impure in colour, much mottled
with black, and the fiery hue is almost absent, rather a dark chestnut or maroon. The
rump is still darker. Of especial interest is the abundant white freckling on the breast,
and the unusually white shafts of the ventral plumage, although even at this age this
character is exceedingly variable. The tail moult produces extremes in colour and
pattern, from a first-year set of rectrices which are almost wholly black, to the clear,
pale buff of the adult. Almost invariably, however, the two central pairs, coming in
later than the others, show a closer approximation to the specialized colouring of the
adult. The young of this undergrowth-haunting bird seems to suffer the loss of many
wing and tail feathers, so that we find them in all stages of half-growth. At this moult
the measurements average, bill from nostril, 16; wing, 227; tail, 145; tarsus, 79;
middle toe and claw, 50 mm. .

EARLY HISTORY

Sir Thomas Raffles described a collection of animals and birds from Sumatra made
through the East India Company, and read the paper before the Linnzan Society of
London, December 5, 1820. This was the first authoritative description of this species.
Subsequently many new generic and specific names were founded on misconceptions
of the status of male and female birds, and disregard of previous appellations.

108 A MONOGRAPH OF THE PHEASANTS

SYNONYMY

Phasianus erythrophthalmus Raffles, Tr. Linn. Soc., XIII. 1822, p. 321 [Sumatra].

Rufous-tailed Pheasant Latham, Gen. Hist., VIII. 1823, p. 204.

flouppifer diardid Guérin-Méneville, Icon. Reg. Anim. 1828-38, Ois., p. 26, pl. 43, fig. 2 [Java].

Louppifer erythrophthalmus Hartert, Nov. Zool., 1X. 1902, p. 537 [Pahang].

Phasianus erythrophthalmus Gray, in Griff, ed. Cuv., II. 1829, p. 28,

Phasianus purpureus Gray, I1I., Ind. Zool., I. 1830-32, pl. 42.

Alectrophasis erythrophthalmus Gray, List Gen. B. 1841, p. 78; id. List of B., Part III., Gall. 1844, p. 26;
Hume, Str. F., VIII. 1879, p. 68 [Malacca]; Kelham, Ibis 1881, p. 531 [ Perak].

Alectrophasis purpureus Gray, List Gen. B. 1841, p. 78.

Gallophasis erythrophthalmos Gray, Gen. B., III, 1845, p. 498.

Gallophasis purpureus Gray, Gen. B., III. 1845, p. 498.

Euplocomus erythrophthalmos Blyth, Cat. Mus. As. Soc. 1849, p. 245.

Euplocamus erythrophthalmus Sclater & Wolf, Zool. Sket. 2, 1861, pl. 34; Sclater, List of Phas. 1863, p. 7,
pl. 8; Sclater, List Animals London Zoo, 1883, p. 484; Mitchell, Proc. Zool. Soc. HONE, joy GAA,

Euplocamus erythrophthalmos Gray, List Gallinae Brit. Mus. 1867, p. 35; id. Hand-l. B., II. 1870, p. 259.

Euplocamus purpureus Gray, List Gallinae Brit. Mus. 1867, p. 36; id. Hand-l. B., II. 1870, p. 259.

Euplocomus erythrophthalmus Elliot, Mon. Phas., II. 1872, pl. 28.

Acomus tnornatus Salvad. Ann. Mus. Civ. Genov., XIV. 1879, p. 250; id. P.Z.S. 1879, p. 651, pl. XLVIII
[Mount Singalan, west coast of Sumatra]; Biittikofer, N. Leyd. Mus., IX. 1887, p. 77 [Padang, W. Sumatra] ;
Grant, Cat. Birds Brit. Mus., XXII. 1893, p. 285 ; Grant, Hand-book Game-birds, I. 1895, p. 242.

Acomus erythrophthalmus Grant, Cat. Birds Brit. Mus, XXII. 1893, p. 283; Grant, Hand-book Game-birds,
I. 1895, p. 241; Sharpe, Hand-list Genera and Species Birds, I. 1899, p. 34; Robinson, Jour. Fed. Malay States
Museums, I. 1906, No. 4, p. 128.

PLATE XXXIV

BORNEAN, CRESELESsS. FIREBACK

Acomus pyronotus (Gray)

Low, wet jungle along the coast of Borneo is the home of this bird. I have seen a pair feeding
beneath a berry tree, upon the fallen fruit, together with the insects which have collected. A small bird
called out in alarm at the sight of some danger, and both birds squatted flat among the begonias. A
scale-bird sang, and confidence was at once restored. A few minutes later a great branch crashed to the

ground, but the birds hardly glanced up, so perfectly did they judge between actual danger and a

harmless jungle happening.

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PLATE XXXIV.

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BORNEAN CRESTLESS FIREBACK.

BORNEAN CRESTLESS FIREBACK
Acomus pyronotus (Gray)

NAMES.—Specific: pyronotus, from Greek zp, fire, and véros, back. English : Bornean Crestless Fireback ;
Bornean Rufous-tailed Pheasant. Native: Singgier.

BRIEF DESCRIPTION.—Male: Head brown, upper neck grey ; lower neck and mantle grey, breast black, all
with white shafts ; lower back golden, changing posteriorly into bronze red, and on the tail-coverts into purple.
Posterior under plumage black, tail buff. Female: Head brown; entire plumage black, heavily glossed on dorsal
surface with steel-blue.

RANGE.—Sarawak, Borneo.

DISTRIBUTION

THE type specimens of this Fireback are labelled “China,” but we now know that it
is confined to the Island of Borneo, and very probably to the north-western portion,
included in the boundaries of Sarawak. Even in this limited area the bird is the
rarest of all the pheasants, and I could learn nothing of it in the interior. Along the
coastal zone, within twenty or thirty miles of salt water, where there is solid ground,
this bird may be looked for.

GENERAL ACCOUNT

The low-lying jungle in Sarawak is the home of the Bornean Crestless Fireback.
The ground for the most part is damp and soggy, and covered thickly with a carpet of
large and small dead leaves in all stages of decay. In every depression water stands,
and every leaf holds at its point a diamond pendant, which drops at longer or shorter
intervals. Even with the most careful stalking and patient watching it is most difficult
to ferret out the life secrets of these wary Firebacks, but first let us see amid what scenes,
what vegetation and animal life they make their home. The pheasant people of whom
we are writing are altogether ground hunters, except at night, when they roost among
the branches of some densely foliaged tree. So we will leave the detailed examination
of the ground until later, and concern ourselves now with the mid and upper jungle.

Vines, saplings, and sprouting palms struggle fiercely for light and air, and always
the vine is the victor, climbing to health and strength on its opponent's trunk and branches.
The trunks are decorated with beautiful scarlet and white lichens, and creeping vines clutch
the bark in their upward course like enormous vegetable millipedes. Thorns and briers
are everywhere ; the terrible spiny palm with its rows upon rows of bristling chevaux-
de-fvise, and innocent-looking vines which, cat-like, sheathe their claws in thick down.
The most insidious is a climbing rotan palm, which sends up its graceful fronds high
in air among the trunks, then from the tips, yards away, drops a wire-like strand to the

ground, covered throughout with the sharpest of tooth-like thorns. This is weighted
109

110 A MONOGRAPH OF THE PHEASANTS

with scores of small orange globes, the fruit, and when one runs against the stem, one
must never pull and lose one’s temper, but carefully disentangle the ingenious trap.

Face-high are myriads of young plants, all shooting up manfully, hoping ever for the
death and fall of one of the giants high overhead—their only hope for continued life. No
two trunks are alike. One has many deep depressions in which water lodges, and affords
a home for hosts of creatures—tree-toads lay their eggs therein, little water-striders
circle about ; each is a tiny ocean with its shallows and its depths, its everlasting game
of life and death. Other trunks are clothed with leaves of creeping, clinging vines,
some dark, lustrous, and covered with a thick mat of long hair, others shiny, wax-like.

Hardly a leaf in the whole jungle is perfect. Some are covered with a beautiful
tracery of fungi; on other plants a score of leaves may present similar patterns of
holes or scallops, showing where some insect had attacked them while yet in the
enveloping bud. j

The jungle growth brings death to our notice much more often than life. True, the
new shoots are here, there, everywhere ; but they grow quietly, and their drooping leaves
are not conspicuous. But on every shrub, or vine, or tree, hang dead or dying leaves.
Thousands are barely attached, ready at the next breath of air to fall. The commonest
~ sound in the jungle—either in a wind or in a dead calm—is the flick, flick, of falling
leaves, the sharper crack and thud of fruit or twigs, and now and then the long-drawn-
out crash of a tree itself.

Small need have the denizens of the jungle to adopt protective colouring, at least
from our eyes. For every conceivable pattern—brilliant or dull—on scale, or fur or
feather, there are a score to match it or go one better among the living and dead vegeta-
tion. A kallima butterfly flits by, the very embodiment of a dead and withered leaf.
Sometimes it alights in orthodox kallima fashion on the stem of a twig, again on a
trunk, again on the surface of some great dead leaf. This one never alighted on a green
leaf, at least in the twenty or twenty-five times I watched it. But our eyes are not those
of keen, hungry, flycatching reptiles, birds, or beasts, so who are we to say this wonderful
colouring is not useful to the uttermost ?

If we are near some berry or fruit tree, we will never be out of the sound of birds’
voices. Usually it is a flock of beautiful green fruit pigeons, which slap their wings
noisily, and almost always a few of the brown-eared chat-thrushes or scale-birds. Their
call-note is sweet and tender as that of a bluebird; their note of alarm is the same, with
an element of harshness predominating. Suddenly every bird in the berry tree may
scream out in dire fear, and with a swish and rush a hawk will fall from the blue sky to
the topmost branches—successful or not, we cannot see.

At such a time I have watched a pair of Crestless Firebacks feeding beneath a berry
tree, upon the fallen fruit, together with the insects which have collected. At the sound
of general alarm from the tree-tops, both birds squatted flat among the begonias, and
with hardly a wink of the eyelids remained motionless until the upper jungle was again
vocal with subdued chirps and calls. A scale-bird sang, and confidence was at once
restored. Hardly had they begun anew their search for food when a good-sized branch
crashed to the ground close at hand. It was a sudden, loud, startling sound to my ears,
but beyond a quick glance the pheasants paid no attention to it; such a sound was to
them a harmless jungle theme, holding no chord of tragedy.

BORNEAN CRESTLESS FIREBACK III

While the pair of Firebacks kept close together, they apparently had no great
affection for one another, and not only occasionally pecked peevishly at each other, but
twice backed away and made half-hearted cuts with their spurs.

In the jungle we are certain, sooner or later, to hear a loud woof! woof! woof!
woof ! and a great black hornbull rushes over. It alights a little distance away, and we
hear the most amazing outcry of complaining creaks and rasping whines. It is, perhaps,
Madame H. in her mud-guarded hollow tree, scolding her husband for a belated djeuner
of berries |

A few subdued crashes announce other actors, and a troop of gibbons swings up.
Their method of progress is the very poetry of monkey travel, hand over hand, each
swing furnishing momentum for the next. Then a sudden stop, and a long thin arm
or leg reaches out and draws in some fruit or berry. A round dark face peers down, a
mother slaps her whining baby clinging to her fur, and the troop is off, almost
noiselessly, except when one lands with a crash after a leap of twenty feet from one tree
to another.

Then a great zebra civet passes close to us with squirrel-like leaps and sudden
rushes. As one sits quietly, birds pass or alight near by every minute or two. In an
island as large as Borneo, which contains no fewer than five hundred and fifty-five
species of birds, the variety which is seen day after day is bewildering. One can never
be sure what will be the next to appear. A certain species may alight close by, fly off
and never be seen again. Others may be observed frequently in many different places.

I have spent a week searching unceasingly for Crestless Firebacks and neither
heard a note nor seen a feather. I have squatted in a most unlikely spot, with my back
against the great buttressed base of a mighty tree, and had two females come into view
almost at once and remain for ten minutes. Not only this, but on the following day
these same birds were in the same place at the corresponding hour. Such perversity of
fate or luck governs the observation of rare pheasants |

The natives trap these birds whenever possible, but only once have I seen their
feathers in head-dresses, and it was seldom that a Dyak could recognize a picture of one,
while the delineation of a crested fireback, argus, or white-tail wrought him to a
high pitch of enthusiasm at once. Their nest, eggs, and young still remain a mystery.
Unlike their Malayan brethren, I never found them near the Dyaks’ communal homes,
but merely by accident in the depth of the jungle.

DETAILED DESCRIPTION

Aputt MaLre.—Crown very dark brown, sometimes with whitish shafts; chin and
throat paler brown. A small patch of bright steel-blue on the nape. Neck blue-grey
all around, paler on the sides and beneath, where the shafts are shining white. - Mantle
black, very finely vermiculated with bluish-grey, with shining white shafts. Back and
wings glossy steel-blue, more sparsely vermiculated, the markings becoming more or
less concentric grey lines on the larger coverts and outer webs of the secondaries.
Primaries plain dull brown.

Lower back iridescent golden bronze. Posteriorly this changes suddenly into rich
dark shining maroon, and this almost at once into the dark steel-blue of the remaining

112 A MONOGRAPH OF THE PHEASANTS

rump and the coverts. Tail pale buff, with the basal half black. On the outer feathers
the buff is sometimes confined, even on old birds, to a small terminal patch on the
outer web.

The remaining under-parts slightly glossed with dark steel-blue, and all under tail-
coverts plain steel-blue ; lower belly dull brown, with enlarged flattened shafts of most
conspicuous shining white. The shape of the breast feathers is wholly unlike that in
erythrophthalmus, being rounded in the latter, but pointed in Ayvonotus, with the shafts
very unlike. In exythrophthalmus the rhachis is dark and normal in shape, tapering,
and splitting up as usual into terminal barbs. In the Bornean bird the rhachis for
the terminal third is expanded and flattened, and of a shining ivory white. In its
enlarged condition it extends clear beyond the terminal barbs as a strong white spine.

Bill yellow, black at base, scarlet facial skin covered with finger-like papillae and a
sparse scattering of hair-like, degenerate, simple featherlets.

The spurs are curved, and from 15 to 33 mm. in length. The measurements are:
bill from nostril, 18; wing, 235; tail, 148; tarsus, 81; middle toe and claw, 51 mm.

Immature males have the lower back much darker, the golden is more tarnished,
mixed with reddish.

ADULT FEMALE.—Crown dark brown. Chin, throat, ear-coverts and sides of neck
paler, smoky brown. Nape, belly and flight-feathers dull brownish black. The
remainder of the plumage black, glossed, especially strongly on the upper surface, with
dark steel-blue.

Upper mandible dark, lower yellowish. Bare facial area scarlet. Iris dark brown.
Spurs straight, 15 mm. in length. Bill from nostril, 15; wing, 220; tail, 135; tarsus, 76 ;
middle toe and claw, 48 mm.

EARLY HISTORY

Temminck was the first naturalist to give this bird a name, but as he never published
it, it counted for naught. J. E. Grey, in his “Illustrations of Indian Zoology,” figured
it, but gave it the same title as the Malayan bird, which by a printer's error reads
erythrothalmus. In 1844, G. R. Gray called the species Ayvonotus, but gave no descrip-
tion of it, so it remained for Dr. Sclater nine years later to finally correlate the bird with
its scientific name and description. Elliot says that a male bird was living in the
London Zoological Gardens in 1870, but I can find no other record of it.

SYNONYMY

Euplocomus erythrothalmus (sic) Gray, (nec Raffles), Ilust. Ind. Zool. II. 1834, pl. 38, fig. 1.

Alectrophasis pyronota Gray, List of B., pt. II]. 1844, Gall. p. 26.

Gallophasis pyronotus Gray, Gen. B. III. 1845, p. 408.

Alectrophasis personatus Temm. MS., Bonap. C.R. XLII. 1856, p. 879.

Euplocamus pyronotus Sclater, P.Z.S. 1863, p. 120; id. List of Phas. 1863, p. 8; Gray, List Gallinae Brit.
Mus. 1867, p. 36; Blyth, Ibis, 1870, p. 174; Elliot, P.Z.S. 1871, p. 138. :

Euplocomus pyronotus Gray, Hand-list of Birds, II. 1870, p. 259; Elliot, Mon. Phas. II. 1872, pl. 29; Salvad.
Ucc. Born. 1874, p. 307; Sharpe, Ibis, 1877, p. 23 (Bintulu); Blas, Verh. Ges. Wien, XXXIII. 1883,
p. 68 (Lihong Bahaja) ; Everett, List B. Born. 18809, p. 199.

Acomus pyronotus Gray, Hand-list of Birds, II. 1870, p. 259.

Phastanus personatus Blyth, Ibis, 1870, p. 174.

Euplocomus personatus Temm. MS., Elliot, Mon. Phas. IJ. 1872, text to pl. 29.

Acomus pyronotus Grant, Cat. Birds Brit. Mus, XXII. 1893, p. 284; Grant, Hand-book Game-birds, I. 1895,
p. 242; Sharpe, Hand-list of Birds, I. 1899, p. 34.

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MAP SHOWING THE DISTRIBUTION OF THE CRESTED FIREBACK PHEASANTS.

Region 1. Lophura diardi
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¥5 3. * ignita.

LOPHURA
CRESTED FIREBACK PHEASANTS

Family PHASIANIDAE
Subfamily PHASIANINAE

Genus LOPHURA

Tue Crested Firebacks have often been placed in the same genus with the silver
pheasants, and indeed Swinhoe’s pheasant bridges them rather closely, but the three
species of Lophura are so well marked, and their resemblances zuzter se are so pro-
nounced, that for convenience it seems much better to recognize them as a distinct
genus. The males are characterized by the shining iridescent feathers of the lower back
and rump, and by the elongated crest, the shafts bare at the base, the tips expanded into
a brush of hair-like barbs. The crest is less developed or absent in the females.

The tail is quite long and composed of sixteen feathers, compressed laterally as in
the crestless firebacks and junglefowls. The third pair of rectrices is slightly longer
than the two central pairs and more than twice the length of the outer pair. The 5th
and 6th primaries are slightly the longest, the 1st being shorter than the 2nd, which is
equal to the roth. A large bare area on the face may be red or blue, and a pair of gular
wattles is more or less highly developed. The tarsus is nearly twice as long as the
middle toe and claw. The males are armed with a pair of stout spurs, absent or very
small in the females.

The term Lofhuva was proposed by Fleming in Philos. Zool. in ihe year 1822.
None of the subsequent generic synonyms call for comment save Diardigallus. This
was instituted by Bonaparte for the Siamese Fireback, and this bird has been considered
sufficiently distinct by a number of authors to be placed in a genus of its own. Any
decision in such an instance can only be made as logical as possible, reviewing the
relative distinctions and resemblances, the relation and value of characters of the bird
as a whole. The crest and fiery back seem in this case to outweigh the other colour and
pattern differences, although avardz is certainly well worthy of pronounced sub-generic
separation, if only on account of the rather unique female plumage.

LOPHURA

Type.
Lophura Fleming, Philos. Zool. II. 1822, p.230 . : . L, ignita.
Euplocomus Temm. (nec Latr., Lep. 1809) Pl. Col. V. ie to Lophophors, 1830) 2b Be Pt
Macarineya Less., Traité @’Orn. 1831, p. 492 . : : : 3
Spicefer Kaup, Das Thierreich, I]. Theil I. 1836, p. 78 . : : : ; "
Lophalector Cab. in Ersch. u. Grub. Encycl. sec. 3, XXII. 1846, p. 116 : ’ ‘ ¢ a
Lophora Gray, Hand-list Birds, II. 1870, p. 259, No. 2378 : . : : .
Diardigallus Bonap. C. R. XLIII. 1856, p. 414 ; 5 E ; ; ; ‘ a th5 CREE.

115

116 A MONOGRAPH OF THE PHEASANTS

or

The genus Lophura, as I limit it, includes three species, the most aberrant of which
is diavdt, The remaining two almost exactly parallel the two species of Acomus in
distribution.

Siamese Crested Fireback 5 : ; : : . Lophura diardi (Temm.).
Malayan Crested Fireback . : : ¢ : . Lophura rufa (Raffl.).
Bornean Crested Fireback : : . ; . Lophura ignita (Shaw and Nodd).

KEY TO LOPHURA
I. Mantle not reddish-brown or chestnut (males).

a Breast grey : ; : : : ; ; c 3 ; . aiardt.
b_ Breast purplish-blue.
a Belly black; central rectrices white : ; i ; , ; . rufa.
& Belly bronze-gold ; central rectrices buff ; : ; : : . ~ignita.
II. Mantle reddish-brown or chestnut (females).
c Wings black barred with buffy white . . . . . . . . diardi.
d Wings chestnut.
c Outer rectrices chestnut . i : : , : : : : . rufa.

@ Outer rectrices black ; ; 5 j : : : : ; . zenita.

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PLATE XXXV

SIAMESE CRESTED FIREBACK.

SIAMESE CRESTED FIREBACK
Lophura dard: (Yemminck) :

NAMES.—Generic: Lophura; from the Greek Adpovgos, bushy or crested tail; Specific: diardz, named for
the French explorer, M. Diard; English: Siamese or Diard’s Fireback Pheasant; French: Faisan Prelat;

German: Praelat; Native: Kai-pha.

BRIEF DESCRIPTION.—Male: Head and throat black, long tufted crest steel-blue ; neck, mantle, breast and
wings grey, finely vermiculated with black ; wing-coverts with a black, white-edged band near the tips; mid-back
shining gold ; lower back and rump bronze-red ; under-parts and tail black, with steel-blue gloss; face scarlet.
Female: Head and neck brown; mantle, lateral tail-feathers and under-parts chestnut, the lower breast and belly
margined with white; scapulars, wings, central tail-feathers, lower back and rump black, the two latter areas
mottled, the others banded with whitish. Face scarlet.

RANGE.—Southern Shan States, Siam, Cambodia and Cochin China.

GENERAL ACCOUNT

Wiru its striking contrasts of colour—shining gold and bronze-red—set off by cold
grey, together with the long, curved tail-feathers and waving, plumed crest, the Siamese
Fireback is one of the handsomest of pheasants. Many are trapped by the Siamese and
brought down to the capital, Bangkok, whence they find their way to many of the
Zoological Gardens and private aviaries of the world. Yet the most recent volume
treating of the pheasants of the Far East begins and ends with the statement that
‘Nothing is known of the habits of this splendid Fireback!”

_ Iwas not able to observe it in its home, and can rely only on the meagre information
which I obtained from residents, whose chief knowledge of the bird was that it was
almost impossible to see and shoot, but when procured was a superb addition to the
daily “chow” !

Bamboo thickets appear to be its favourite haunt, as two of my correspondents give
this as a reason for the impossibility of following up a bird. It leaves these dense
impenetrable tangles in the early morning and comes into more open jungle, preferably
near the banks of rivers, to feed and drink. A sportsman who was accustomed to sit up
in wait for tigers, more than once saw a pair of Diard’s Firebacks come forth from the
close-growing ranks of great bamboos and scratch about in the forest mould.

The natives report it as a stupid bird, and very easily trapped with simple hair or
wire nooses set in runways, or about some food strewn upon the jungle debris. When
a camp or more permanent building is erected in the forest, if the locality has been a
favourite feeding-place of Firebacks, the birds do not desert at once, but after the first
alarm has passed they return, and occasionally in early morning are seen near the
clearing. This is especially true if cattle or buffalo are kept, the birds finding much
insect food in the vicinity of these animals. .

The Siamese Fireback has been hatched and reared several times in the gardens

of the London Zoological Society, but apparently no record has been kept of either eggs
117

118 A MONOGRAPH OF THE PHEASANTS

or chicks. Of eight individuals which have lived in these gardens, the average length
of life was three years and four months, while one reached the age of nine years and
three months. An egg collected in Siam is bluish white, with a faint reddish shade,
and measures 48 by 38 mm.

In captivity these birds soon become tame and familiar, and I have never seen a
cock which would not soon come up to its keeper and take meal-worms from his fingers.
At the same time they are very strong, and with their spurs are able to do great damage
to other birds. If allowed to run freely with poultry they will engage every cock in
combat, and no matter how skilful or strong a game-cock may be, it has no chance
against this active Fireback of the Siamese wilderness. A writer in ‘The Field” tells
us that even when the opponents of such a captive pheasant were double its weight they
had no chance against it. He goes on to say that “ At length its pugnacious propensities
became such a nuisance that my neighbour's servants seized it one day and cut down its
long, sharp spurs to mere stumps. The poor bird, however, though shorn of its strength,
lost none of its spirit, and assailed its gallinaceous enemies as recklessly as ever. These
soon found out that its weapons were gone, and plucking up courage, paid it off with
interest for former injuries ; in a week or two my Diardigallus had scarce a feather left
on it.” The same writer says, that when in good health, the Siamese Fireback incessantly
uttered a sweet, mellow cooing and clucking.

DETAILED DESCRIPTION

ADULT MALE.—Feathered parts of the head, chin and throat black; crest of ten
or more feathers, black glossed with steel-blue. These feathers are 85 mm. long, with
thickened bare shafts for about two-thirds of the entire length, the extremity being
formed of a long tuft or bunch of disintegrated barbs. The neck and the entire mantle
and breast, together with the scapulars, wing-coverts and secondaries (these latter chiefly
on the outer webs), show a beautiful even vermiculation of white and black, producing
almost a monochrome grey. The scapulars, lesser and median wing-coverts show a
black sub-terminal band, narrowly edged with white. Mid-back, shining gold on the
broad, disintegrated terminal fringe which forms the visible portion of the feathers,
then an equal area of pale buff, the remaining basal portion vermiculated grey like
the mantle; lower back, rump and shorter tail-coverts with a terminal fringe of rich,
dark, shining crimson, the remainder of the feathers being glossy steel-blue. Longest
tail-coverts purplish-green with a rather narrow terminal fringe of clear, shining sage
green. Primaries brownish-black, faintly mottled with grey. Belly, lower sides, flanks
and under tail-coverts black, with a broad terminal fringe of steel-blue. Tail-feathers
black, glossed with purplish-blue on the outer webs, greenish on the inner. The
rectrices are strongly curved, the three inner pairs being remarkably long and pointed.
Bill yellowish, darker toward the base; facial skin, wattles, legs and feet, scarlet; spurs
and claws whitish horn colour. Bull from nostril, 18 mm.; wing, 233; tail, 322;
tarsus, 101; middle toe and claw, 58; spur, 20.

ADULT FEMALE. Top of the head and neck brown, shading into smoky grey
on the chin and throat. Mantle clear, dull chestnut, posteriorly becoming more and
more mottled with black, then changing rather abruptly into the pattern of the

SIAMESE CRESTED FIREBACK 119

remaining upper parts. Mid-back black, coarsely barred with buffy white; lower back,
rump and tail-coverts similar, but the barring disintegrates into a coarse mottling.
Scapulars, wing-coverts and secondaries black, strikingly barred with white, the centre
of the bars showing considerable black mottling. The exposed portions of the greater
coverts and secondaries show much greenish gloss. On the primaries the bars become
buffy or rufous-brown, and are much broken. Two central pairs of rectrices closely
resemble the secondaries, but with the eight or more bars much diffused with mottling.
Third pair of tail-feathers chestnut, with a great deal of black barring and mottling
on the inner webs; the remaining rectrices and under tail-coverts clear chestnut.
Entire ventral surface reddish-chestnut, the breast almost monochrome, but with a
white marginal fringe appearing on the lower breast, and increasing in extent over
the remaining posterior ventral plumage. Upper mandible dark, lower chiefly yellowish
horn colour; irides hazel brown; facial skin, legs and feet scarlet ; claws whitish horn.
Spurs low, sharp scalules. Bill from nostril, 16 mm.; wing, 223; tail, 202; tarsus, 81;
middle toe and claw, 46.

The female dard is interesting as exhibiting the same differentiation of pattern
in the tail as is shown by the males of /comus and the other species of Lophura.

JUVENILE PLuMacE.—Crown (lacking a crest), sides of head and back of neck
dark, smoky brown, merging into the mantle, which is rufous, slightly mottled with
dark. On the scapulars a black spot occurs, and on the mottled grey wing-coverts
this increases until the pale buff colour is confined to the tip and to several narrow
transverse bars. On the inner secondaries these become very distinct, each feather
showing six or more straight, narrow cross-bars on a black field. The tail-feathers
are chestnut, marked and barred with blackish. The ventral plumage is dull brown,
faintly freckled with whitish. The specialized fiery back is only faintly indicated at
this age: a slight golden fringe much freckled and dulled with black and white, while
the bronze-red rump is barely suggested by a rather mottled chestnut fringe.

First YEAR PLumace.—The wing of the young male Fireback [Coloured Plate
XXXVI, fig. 4], when in half-way transition between juvenile and first-year plumage,
is exceedingly interesting. One such wing showed primaries Nos. 9 and 1o full grown,
but with the sheath still clinging to their bases as plain evidence of their delayed
growth. Primaries 1 to 6 inclusive were new, 1 to 3 full grown, while Nos. 4, 5
and 6 were new feathers in active growth, measuring respectively 149, 124, and 99 mm.
out of their sheaths. Nos, 7 and 8 were still unshed.

It should be remembered that the wing-feathers of the adult male, both primaries
and secondaries, show no complex patterning. The former are brownish-black faintly
flecked with grey; the secondaries are all evenly and minutely vermiculated (chiefly
on the outer webs) with white. Keeping this in mind, such a wing as that which
we shall describe is most interesting and significant. The gradation of specialized
pattern revealed by the delayed growth of the outermost primaries, together with the
presence simultaneously of pure juvenile and typical adult feathers, gives us, upon
the surface of the single wing, a review of all the ontological pattern changes through
which the young bird passes as it gradually attains the adult state.

120 A MONOGRAPH OF THE PHEASANTS

Primary No. 7 may be taken as typical of the real juvenile plumage. It is dark
brown, with faint, wholly irregular mottling of dull rufous on the outer web.

Primary No. 8 (the third from the outer), which, as usual, was somewhat delayed
in its appearance, has pale buff markings on both webs, in the form of bands oblique
to the shaft, about nine altogether, wavy and mottled, but distinct. Nos. 9 and 10
very evidently appeared synchronously and are almost identical. The pale buff, mottled
bands persist only on the inner part of the inner webs, while in place’of the bands
elsewhere is now a faint, diffuse cloudiness of pale blue-grey, which has spread until
the interspaces have become reduced to roundish, dark brown spots, very regular and
sharply delimited on the outer webs. There are six or seven on each web.

Passing inward to the newly growing feathers, in No. 6 we find a further
development of this pattern. The vane in general is very dark brown, entirely, evenly
and finely flecked with minute white dots, except for the terminal 38 mm., where are
three pairs of markings, half broken oblique bars, half marginal ocelli.

On the succeeding inner primaries the white peppering or vermiculation is
unbroken.

As usual in this stage of phasianine wing-moult, the outermost secondary (No. 1)
is still unshed, although much worn and somewhat broken. It is brown, with pale
buff cross-bars. The next seven secondaries are new, having been shed successively
inward, the eighth being very short. The succeeding five are unshed, showing the
typical juvenile, buff-banded pattern.

In a second immature male, with the wing-moult somewhat farther advanced, the
long-delayed first or outermost secondary has been shed and the new feather is half
grown. In colour it is adult, evenly freckled with bluish white, while the succeeding
four secondaries all show distinct bands or ocelli on the outer webs. The sixth
secondary is typically freckled. So the delayed outer secondary skips the pattern of
the next four inner ones, its pattern being of the same adult type infused into the
sixth and succeeding ones.

The moult of the primaries in this individual must have occurred rather earlier
than usual, before all the juvenile pigment was eliminated from the blood, for the
innermost, or No. 1, is quite buffy and banded, and indeed not until the fifth primary
is reached do we find unadulterated, pure adult freckling.

The tail moult of these birds, proceeding regularly from the outside inward, is
very readily followed by means of the radical change in colour and pattern; the juvenile
rectrices being rich chestnut, banded with black, especially heavily on the inner webs,
while the new feathers are jet black with a slight greenish gloss.

SUMMARY

The reason I have treated in such detail this transition from juvenile to adult
plumage, is that it seems to indicate the suppression of a specialized plumage pattern
which is visible only through the accident of delay in certain wing-feathers. The flight-
feathers of the adult, both primaries and secondaries, show no complex pattern. The
former are brown, faintly flecked with grey, the latter evenly vermiculated with white.
The very distinct bars and ocelli shown by feathers of intermediate growth [see Coloured

PLATE XXXVI

PLUMAGES OF THE BORNEAN AND SIAMESE CRESTED FIREBACK

Lophura wnita and Lophura diardi

Fic. 1. Chick of Bornean Crested Fireback, Lophura tguita.
Fics. 2 and 3. Bornean Crested Fireback, Lophura z¢ntta.
Juvenile plumage of the female (2), and male (3).
Fic. 4. Siamese Fireback, Lophura diard:.
Transition plumage between juvenile and first year, showing sequence and nodes of moults

and the unusual pattern complexity of the immature wing-feathers.

‘IVEXX BTAIT

[pete umn eweermaaetgermcey

Sieh,
ene

Sins

slem bas {s} ae ais to ogenuulq slinovisl

cd ti “s2omaie nt of :
2 bbe

r cinieneT

PLATE XXXVI.

We ROR
AAT arc&,
Ni thes:
wg Not

od ,

PLUMAGES OF THE BORNEAN & SIAMESE CRESTED FIREBACKS.

SIAMESE CRESTED FIREBACK 121

Plate XXXVI, fig. 4] are comparable neither with the buffy markings of the juvenile
plumage nor the unbroken design of the adult pigmentation. The coverts, however,
as we have seen, retain in the adult a terminal band of black. As far as I know, there
is no specimen of the chick in down nor of early complete juvenile plumage extant
in collections, and only two or three immature males in any museum.

EARLY HISTORY

When Gould, in 1860, published Vol. VII. of his “ Birds of Asia,” he was able to
figure the male of this species from a specimen which Sir Robert Schomburgk, British
Consul-General for Siam, procured alive from a small collection of animals kept at
a Wat or Siamese temple. Gould added as the supposed female a figure drawn by
Mr. Crawford from a bird obtained at Ava, Burma. This is undoubtedly a very poor
representative of a young male or female of some form of Gennaeus hybrid. The
elongated tail-feathers of the male are wrongly represented as turning outward like
those of a black cock, instead of closely opposed and sweeping gracefully around and
downward. Elliot's plate of the male bird, while correcting this error, is totally wrong
as representing these feathers as short and but little curved. The female is recognizable,
but has the neck pale buff, not chestnut, and much of the mantle barred instead
of plain.

SYNONYMY

Euplocomus diardi Temm. MSS., Bonap. C. R., XLIII. 1856, p. 415; Gray, Hand-list of Birds, II. 1870,
. 260.

c Diardigallus praelatus Bonap. C. R., XLIII. 1856, p. 415; Schl., Hand-l. Dierk, I., p. 379, Atlas Aves,
1857, pl. V. fig. §5 ; Gould, B. Asia, VII. 1860, pl. 21 [male only].

Gallus diardi Temm. MS., Schl., Hand-l. d. Dierk, I. 1857, p. 379, Atlas Aves, pl. V. fig. 55.

Diardigallus fasctolatus Blyth, Jour. As. Soc. B., XX VII. 1858, p. 280.

Diardigallus crawfurdi Schomb., Proc. Zool. Soc. 1862, p. 250.

Euplocamus crawfurdi Sciat., List of Phas, 1863, p. 6.

Euplocamus praelatus Sclat., List of Phas. 1863, p. 6, pl. 6 [Siam, Shan States]; Schomb., Ibis, 1864, p. 259
[E. Lao Country]; Sclat. & Wolf, Zool. Sket. 2, 1867, pl. 35 ; Gray, List Gallinae Brit. Mus. 1867, p. 34; Elliot,
Mon. Phas., II. 1872, pl. 24; Mitchell, Proc. Zool. Soc. 1911, p. 522.

Lophura diardi Grant, Brit. Mus. Cat. Birds, XXII. 1893, p. 290; Grant, Hand-book Game-birds, I. 1895,
p. 247; Nehrkorn, Kat. der Eiersammlung, 1899, p. 193; Gyldenstolpe, Kungl. Svensk, Hand-l. 50, No. 8, 1913,
p. 67; Gyldenstolpe, Jour. Nat. Hist. Soc. Siam, I. 1915, p. 235; Baker, Jour. Bombay Nat. Hist. Soc. XXVI.
p. 75 [desc. and habits].

Diardigallus diardi Sharpe, Hand-list of Birds, I. 1899, p. 35.

VOL, II R

MALAYAN CRESTED FIREBACK
Lophura rufa (Raffles)

NAMES.—Specific: rufa, from the Latin rufus, red, with reference to the colour of the fiery back. English :
Malayan Crested Fireback ; Vieillot’s Fireback. French: Faisan de Vieillot. German: Rothriickenfasan. Native:
Pégar, or Ayam Pégar (Malay and Sakai) ; Ayam siul [Whistling fowl] (Malay).

BRIEF DESCRIPTION.—Male: General plumage black, glossed with purplish-blue; lower back and rump
fiery bronze-red ; feathers of the sides and flanks with white or chestnut shaft-stripes ; middle pairs of tail-feathers
white. A tufted crest with long bare shafts. Facial skin and wattles blue; feet red. Female: Rich chestnut
above, faintly mottled with black; throat white; breast chestnut edged with white ; remaining under plumage dark
brown, widely margined with white. Crest shorter than in male. Facial skin blue; feet red.

- RANGE—From southern Siam and Tenasserim southward ; Malay Peninsula and Sumatra.

GENERAL ACCOUNT

Tuis is decidedly a low-country bird, and I doubt if it ever ascends to the higher
altitudes of the Malayan or Sumatran mountains. In Siam I know of records only
from the extreme south, and it is improbable that it overlaps the range of dard. Even
in the low Malayan coastal country of the west and the central Pahang region drained
by the tributaries of the river of the same name, where I have observed this species,
its distribution is far from regular. Great stretches which seem eminently suited to
its requirements are apparently uninhabited by it, where the natives have no
knowledge of the bird. Then again, not far distant, every child will recognize its
picture and shout “Ayam pégar!” and a few days of careful search will reveal
traces of a family or flock. Nevertheless, in spite of its comparative abundance in
some areas, my search for the Malayan Crested Fireback proved one of the most
trying of my experiences with wild pheasants. By the calendar it was still the dry
season; by the rain gauge it was the height of the rainy period. In the open forest
where we expected to find this species we neither saw nor heard it.

Years ago, Malays or Sakais made numerous clearings throughout many parts
of the jungle, planted some crop, and then deserted the clearing for another. Many
of these were in valleys, close to the bed of small streams. Such places are now
covered with a tangle of thorny palms and enmeshed vines, and in these all but
impenetrable thickets, the beautiful Crested Fireback elects to spend most of his time.
All trace of the former savage cultivator of the soil has vanished—hut, tools and
pottery. But now and then, from the heart of some such tangle as I have described,
a pair of house crows will fly up croaking hoarsely. Through all the years they or
their descendants have clung to this bit of human handiwork, deep in the jungle
and far from the haunts of living human beings. They form a parallel to the little
Guiana house wrens which one finds still haunting old, long-deserted and overgrown
huts in the heart of the British Guiana forest.*

1 Beebe, “Our Search for a Wilderness,” 1910, p. 307.
122

-MALAYAN CRESTED FIREBACK

: Lophura rufa (Raffles)

wee

Reng stall Bad ce most wearisome wait. They are found in pairs or in inal families, and

larmed_go off with a headlong rush eeinotas the underbrush in preference to pang flight.

ADAGAAIA aareaa ‘ AYA awe
ASM uae

yodT dnswotleveis’ ods

ai Wussd: tial eT dst: we ‘ald 8 sible 10° BR

bas eoilietey linet itan on varia tt ‘haut os yor h ©

jd of Saitiaqtis oe Ob Sonmalarg o ar idetdsobay, an

> 4 z. ‘

ee iy ha y
ees % by

PLATE XXXVII.

ss >:
2 gene a
crn

, MALAYAN CRESTED FIREBACK.

MALAYAN CRESTED FIREBACK 123

One day’s experience on a small unnamed tributary of the Pahang River, in the
heart of that central Malayan State, is typical of many short trips which we made
in search of this bird. The sun rises through the fogs which have drifted down
from the mountains, and we find the air of early morning cool and invigorating.
We follow a narrow game trail for a mile or more, and then turn aside into a
creek bed. The moisture on the foliage soon drenches us, and we tramp through
mud and water with as little noise as possible. In one likely spot we wait for an
hour, but see nothing of the birds. We have heard one call far ahead of us and
so keep on in hope. The sun seems to have leaped overhead and the whole jungle
steams with heat. From now until we return we pant for breath, as if exercising
in a hot-house or Turkish bath. Whenever we stop to listen, a myriad tiny gnats
dance close before our eyes. Mosquitoes worry us on all sides—almost all Stegomyia,
which we have roused from their resting-places on the leaves. One cannot but think
what a terrible scourge to the natives of all this region yellow fever would be.
Nothing could confine its ravages.

At last we reach one of the old over-grown clearings, and begin to cut our
way carefully, foot by foot, pressing the cables of serried, recurved teeth to one side.
When we can go no further we stop exhausted. Two or three species of bulbuls
are singing close by and a pair of big racket-tailed drongos swoop back and forth
overhead. It is impossible to sit quiet on account of the flies and mosquitoes. We
clear a circle of leaves and sit back to back, flicking away the leeches which are
ever humping over the ground toward us. An hour passes, and only the tracks in
the mud near by and the begonias torn up by the powerful scratching of the birds,
hint of their earlier presence.

Once, from a great distance, we hear the clear trumpet of an elephant and,
shortly afterward, a cloud effaces the sun and a wind rustles the palm fronds. This
increases to a gale, and instead of panting with still, humid heat we are shivering
in the coolness. The birds have vanished, two long-tailed monkeys dash wildly from
tree to tree, passing as swiftly as we could run.

Then comes the rain—sudden, windless, torrential. At first we seek shelter
beneath the camera cloth, but it is of no more use than so much mosquito-netting.
Protecting the camera as best we may, we plod slowly back. Twenty yards from
where we have been sitting, three Firebacks rush out from a clump of large-leaved
plants and, with necks low, scatter in as many directions. This is our reward for
the day’s work.

I saw this species, however, a number of other times—once a pair unalarmed,
scratching near an old decayed log, and feeding greedily on the termites which they
uncovered. On another occasion, the glimpse was again but momentary, a sudden
rush and rustle through the underbrush, never flying, but always with keen wit
trusting to their great speed.

Many years ago Davison wrote that “these birds frequent the thick evergreen
forests in small parties of five or six; usually there is only one male in the party,
the rest being females, but on one or two occasions I have seen two males together ;
sometimes the males are found quite alone. I have never heard the males crow, nor
do I think that they ever do so; when alarmed, both males and females have a

124 A MONOGRAPH OF THE PHEASANTS

peculiar sharp note, exceedingly like that of the large black-backed squirrel (Scdurus
bicolor). ‘The males also continually make a whirring sound with their wings, which
can be very well imitated by twirling rapidly between the hands a small stick, in a
cleft of which a piece of stiff cloth has been transversely placed. I have often
discovered the whereabouts of a flock by hearing this noise. They never come into
the open, but confine themselves to the forests, feeding on berries, tender leaves, and
insects and grubs of all kinds, and they are very fond of scratching about after the
manner of domestic poultry, and dusting themselves. When disturbed, they run
rapidly away, not in different directions, but all keeping much together; they rise
at once before a dog, getting up with a great flutter, but when once well on the
wing, fly with a strong and rapid flight. They seldom alight again under a couple
of hundred yards, and usually on the ground, when they immediately start running.

“TI noticed on one occasion a very curious thing. I had stalked an argus, and
while waiting to obtain a good shot, I heard the peculiar note, a sort of chukun,
chukun, followed by a whirring noise made by the male Fireback, and immediately
after saw a fine male Fireback run into the open space, and begin to chase the argus
round and round its clearing. The argus seemed loath to quit its own domain, and
yet not willing to fight, but at last, being hard pressed, it ran into the jungle. The
Fireback did not attempt to follow, but took up a position in the middle of the
clearing, and recommenced the whirring noise with its wings, evidently as a challenge,
whereupon the argus slowly returned, but the moment it got within the cleared space
the Fireback charged it, and drove it back into the jungle, and then, as before, took
up his position in the middle of the space and repeated the challenge. The argus
immediately returned, but only to be again driven back, and this continued at least
a dozen times, and how much longer it would have continued I cannot say, but a
movement on my part attracting the birds’ attention, they caught sight of me, and
instantly, before I could fire, disappeared into the jungle. The argus never made the
slightest attempt to attack the Fireback, but retreated at once on the slightest
movement of the latter towards it, nor did I see the Fireback strike the argus with
either bill, wings or spurs.”

Nothing is known of the nesting of this bird in a wild state, but it has
occasionally laid eggs in captivity. The average size of three such eggs is 536 by
39'4 mm. The surface is smooth, almost without gloss, and the colour is in general
a pale, dull buff.

DETAILED DESCRIPTION

Mave.—Short contour feathers of the head, chin and throat black; neck, mantle,
scapulars, shorter tail-coverts, breast and sides, black with a wide terminal fringe of
purplish blue. Wiing-coverts similar, but with the fringe shining steel-blue. On the
greater wing-coverts, inner secondaries and longer upper tail-coverts the gloss extends
more or less over the entire surface, and on the strongest marked feathers is slightly
greenish. When the feathers show wear, however, this gloss is the first character to
suffer, and the vane becomes dull blackish. Back and rump fiery bronze-red, the
anterior portion with a decided golden sheen. This iridescence is confined to the
visible portion of the feathers, namely, a very broad, terminal, rather disintegrated

PHOTOGRAVURE 26

HOME OF Ith MALAYAN CRESTED FIREBACK

In the low, dense jungle of the Malay States, in thickets of thorny tangles and enmeshed vines, these
birds live. To study them one must fight hosts of leeches and mosquitoes, mud and rain, or, when the
sun appears, the steaming heat of these breathless places. Here the bulbuls sing sweetly, and gorgeous
butterflies flap slowly past, and here these pheasants scratch among the dead leaves for food, or walk in

the shallow water of jungle creeks, drinking and catching tiny creatures in the sand and gravel.

AAUVARDOTOHT

sy

4

only bedesmas

atle asd. 30
aucapiog ft spade:
ff Alaw 90. Baal ey

26.

PHOTOGRAVURE PLATE

a

aS)

a9)

NVAVTIVIW G2HL 40 ZNO

T
ae

T
aly

MALAYAN CRESTED FIREBACK 125

margin, which is followed successively by an equally wide band of chestnut, and a
narrow zone of steel-blue; the remainder being dull black. Primaries and secondaries
are brownish-black unmarked. Two inner pairs of tail-feathers and the inner web and
tip of the third, pure white; the remaining rectrices dull jet black. Feathers of the
side, breast, sides of body and the flanks with a conspicuous shaft-stripe on the visible
terminal third occasionally expanded distally, pure white in some cases, or chestnut,
or a mixture of the two. Abdomen, thighs, under wing and tail-coverts sooty black,
with no metallic gloss, or a small amount on the under tdil-coverts.

Mandibles, spurs and claws yellowish horn, the former darker at the base; irides
red; facial area bright smalt blue; legs and feet red, especially strong on the front
of the tarsi and toes, the posterior surfaces out of the breeding season being reddish
white. Weight, 5 lbs. Length, 720; extent, 940; bill from nostril, 23; wing, 286;
tail, 266; tarsus, 109; middle toe and claw, 65. Spurs straight and stout, average
length, 38.

MALE PLUMAGE VARIATIONS.—Although this species does not show any greater
variations from the normal than many of the other pheasants, yet atypical individuals
have caused great confusion. After careful comparison of all the specimens I could
find in the museums of the Far East, of Europe and of America, together with those
I was able to collect, I can find no sufficient grounds for separating the Sumatran from
the Malayan birds, nor of recognizing the single aberrant captive specimen from an
unknown locality, now in the Leyden Museum, as distinct from the Bornean crestless
fireback.

If, on the other hand, such characters as have been utilized in the separation
of these two forms are recognized as valid, then I should logically be compelled to
distinguish three or four other “species” based on. equally variable, and, it seems to
me, insufficient characters. I have in mind one male bird with half the mantle of clear
chestnut, and another full-grown male with the central rectrices half chestnut instead
of pure white. It is wholly impossible to separate the Sumatran birds on the grounds
that the shaft-stripes of the side and flank feathers are predominately buffy or chestnut
rather than white. Over fifty per cent. of the Malayan birds show considerable chestnut
on these feathers, and an adult male sent from Johore to the Raffles Museum, Singapore,
for mounting, exhibited a greater amount of chestnut-red flank markings than I have
ever seen on any Sumatran bird. Such being the case I see no logical possibility of
distinguishing more than a single species of this northern Crested Fireback. As to
sumatrana as defined by Biittikofer,* the variation zzter se of the five male specimens
which he lists is such as to give but slight value to the status of this form. I have
seen at least a half-dozen adult specimens in the museums of the East and elsewhere
which show as much variation in the chestnut and white of the rectrices and the amount
of red on the flanks as in the above-mentioned five birds. As these specimens were
divided between Sumatra and Pahang, I see no course open but to consider them as
aberrant variations of vzfa in the direction of zgn7ta.

FeMALE.—Head, neck, mantle and lesser wing-coverts uniform chestnut. Re-
maining parts of the upper plumage and wings finely vermiculated with black, this

1 “Notes from Leyden Museum,” XVII. 1895, p. 177.

126 A MONOGRAPH OF THE PHEASANTS

colour becoming more abundant posteriorly and giving the feathers a general darker
tone. Flight-feathers brownish-black, the primaries faintly and the secondaries heavily
mottled and streaked with chestnut on the outer webs. Longest upper tail-coverts
like the secondaries. Tail-feathers chestnut-brown, the two central pairs mottled with
black. In many specimens these markings are in the form of narrow, widely separated,
but distinct transverse bands. Chin and throat creamy white. Lower neck and breast
chestnut, margined with white on the sides. . Lower breast, abdomen, sides, flanks and
thighs brownish-black, often mottled with chestnut, with a wide margin of white all
around. Lower abdomen with the brown reduced to a central mottling, the remaining
portion of the downy feathers white. Under tail-coverts brownish-black, with dull
rufous margins. Irides red; facial skin blue, not as bright as in the male; bill and
cere dark, the edges of lower mandible and tip of the upper paler horn colour; feet
and legs pale flesh, redder on the frontal scales. Weight, 3} lbs. Spurs sharp, but
very small, the longest only 4 mm. Length, 602; extent, 806; bill from nostril, 18;
wing, 255; tail, 207; tarsus, 95; middle toe and claw, 55.

FEMALE PLUMAGE VARIATIONS.—The females show considerable variation, the
chief being where, on the one hand, the entire dorsal surface is rich chestnut, mottled
on the wings, back and rump with black; or again birds with the back, wings and
rump pale buff, heavily vermiculated with black, the mantle being quite evenly dotted
with black. These are the extremes, and we find all sorts of intergradations. Then
too in a small percentage of specimens the white edging of the ventral plumage is
found on the feathers of the upper neck and mantle. These variations are due neither
to age nor locality.

NataL Down.—Short facial down pale buffy white; forehead pale buff, warming
into rufous buff on the crown and rufous on the nape and hind neck. A narrow black
line begins at the lower posterior edge of the orbit (with a tiny bit of black on the
upper posterior margin), and extends backward and downward across the facial area
and ear-coverts, merging with the rich orange rufous of the hind neck. The back is
dark chocolate brown, the wing-coverts more chestnut, with a broad buff bar across
the centre of the wing. Two well-marked buffy white lines begin on the scapulars
and extend backward parallel with the wide chocolate dorsal area, to the rump. Outside
of these in turn two chocolate bands, equal to the buff in width, extend backward from
under the posterior dorsal edge of the wings, turning at last rather abruptly downward
to the flanks. The inferior border of the facial area, the breast, sides and flanks are
strongly tinged with buff, while the remaining ventral surface is creamy white. Iris
dark hazel brown. The bill from nostril is 4 mm.; wing, 25; tarsus, 24; middle toe
and claw, 18.

JUVENILE PLuMAGE, Mare.—An interesting stage is shown by a bird with
abundant juvenile plumage, but showing active traces of the first-year or post-juvenile
moult. This moult, as we shall see in the description of the succeeding bird, seldom
results in perfect adult hues and patterns; and although the bird may breed the coming
season, its tail-feathers are seldom pure in colour, and its iridescence is not as strong
as it will be after the second annual moult.

MALAYAN CRESTED FIREBACK 127

A male at this stage shows little or no signs of moult on the head and neck, the
juvenile feathers being dull blackish-brown, with very faint rufous tips. The lower
hind neck is paler brown, with here and there a very slight metallic glint. The mantle,
back and coverts are rufous with obscure dark brown mottlings. On these areas there
_ is a thin scattering of new jet-black feathers with a shining blue-green border. The
longer juvenile wing-coverts are dark rufous-brown, with the rufous mottlings confined
to the terminal border. The flight-feathers are all dull brown, unmarked.

The juvenile rectrices are narrow and curved, dark rufous with almost obsolete
mottling of darker colour. In the specimen which we are describing there are fourteen,
being moulted from without, inwards. The outer four are new, growing, dark feathers,
becoming successively shorter as we proceed inward, until the fourth from the outer is
very short. The next inner pair is wholly absent, apparently just shed, while the two
central pairs are brown, narrow juvenile feathers, projecting well beyond the others. Now
it is just these three inner pairs which will show the white in the adult plumage, so here
(and in all the other moulting individuals examined) we have a very marked delay in
the appearance of the central pairs, just as in /¢/ag¢nis we had a delay in the outer pairs.

The facial area is very sparsely covered with minute, brush-like feathers. Chin
and throat white; ear-coverts, sides and lower neck like the upper. The ventral surface
shows no signs of moult as yet. The feathers are dark brown, with a narrow white
lateral fringe on the breast, increasing on the sides of the body, and especially on the
belly, where superficially it eclipses the darker central portion. The moult of the
primaries is proceeding regularly outward, all but the external five pairs having been
shed. The spurs are sharp, but very minute. The bird measures, bill from nostril, 18;
wing, 215; tail (juvenile), 122; tarsus, 74; middle toe and claw, 38.

IMMATURE MALE, CoMPLETING First ANNUAL Mou.t.—A bird at this stage
has more tiny, brushy featherlets on the facial area than the fully adults. In a typical
individual the crest has begun to sprout, being about one-half grown. The dorsal
plumage shows only about half the full amount of terminal iridescence, while the fire
of the back is broken by black mottling, and an infusion of dull chestnut at the basal
’ border. The longest tail-coverts are still unshed juvenile, being dark brown with
mottled rufous tips instead of shining purplish-green.

The tail-feathers have been renewed and are full grown. The two central ones
have a conspicuous black shaft-stripe for the basal three-quarters of the feather, while
the basal fourth is wholly brown. ‘The next pair is similar, except that black mottling
covers the basal third of the outer web. The third pair has the white inner web clouded
with dark, and a patch of chestnut at the tip of the outer web. The next has a patch
of white and chestnut at the tip of the inner web, while the fifth shows only a tinge
of chestnut in the same place. The two outer pairs are normal black.

The under-parts are almost wholly dead black. On the sides are a few purplish-
blue tipped feathers, with a narrow shaft-streak of bright chestnut, terminating in a
patch of mottling of the same colour. The outer two primaries are growing; all others
have been renewed. This bird measures, bill from nostril, 23; wing, 253; tail, 195;
tarsus, 106; middle toe and claw, 61. The spurs are sharp, stout and short and
measure 10 mm. in length.

128 A MONOGRAPH OF THE PHEASANTS

As to the change of colour without moulting which some authors describe at this
age of the Crested Fireback, I can find not the least confirmation in the examination
of museum skins, and in the case of the Bornean Crested Fireback, which I have bred
under careful observation, I have discovered nothing except proof that the change in
pigmentation, iridescence or pattern takes place by moult and renewal alone, except as
I have stated before, where excessive wear destroys the high metallic gloss.

IMMATURE FEMALE.—Two immature females show the nape heavily edged with
black, and the mantle edged and mottled with more or less solid black pigment. The
banding is especially noticeable on the middle wing-coverts, and the secondaries are
very heavily mottled with the same colour, On the. lower parts, the black almost
obscures the rufous of the breast.

SYNONYMY

Phastanus ignitus Raffles (nec Shaw and Nodder), Trans. Linn. Soc., XIII. 1822, p. 320 (Sumatra) ;
Daniell, F.Z.S., 1882, p. 24; Elliot, Ibis, 1878, p. 412.

Phasianus rufus Raffles, Trans. Linn. Soc., XIII, 1822, p. 321 (Sumatra); Gray, in Griffiths ed. Cuv., III.
1820, p. 28.

Phasianus castaneus Gray, in Griffiths ed. Cuv., III. 1829, p. 28 (Penang).

Gallus macartneys Schinz (nec Temm.). Nat. abild. Vog., 1833, p. 28, pl. 93 (Sumatra).

Euplocamus ignitus Gray, Ul. Ind. Zool., I. 1834, pl. 39; Blyth, Cat. Mus. As. Soc., 1849, p. 243 (Sumatra) ;
Blyth and Wald., Cat. Mamm. and Birds, Burma, 1875, p. 149 (Tenasserim River); Elliot, Ibis, 1878, p. 124.

Euplocamus vieillotti Gray, List Gen. E. 2nd ed., 1841, p. 77; Gould, Birds Asia, VII. 1852, pl. 15 (Malacca);
Hume, Stray Feathers, II. 1874, p. 481 (Tennasserim) ; Hume, Stray Feathers, III. 1875, p. 324 (Tennasserim) ;
Hume and Marsh, Game-birds India, I. 1878, p. 213; Hume and Dav., Stray Feathers, p. 438 (Pakjan) ; Elliot,
Ibis, 1878, p. 413; Kelham, Ibis, 1881, p. 532 (Perak) ; Oates, Birds of Burma, II. 1883, p. 320 (L. Tennasserim).

Euplocamus rufus Hume, Stray Feathers, V. 1877, p. 121.

Euplocamus sumatranus Dubois, Bull. Acad. Belg. (2), XLVII. 1879, p. 825 (Sumatra).

Lophura rufa Ogilvie-Grant, Cat. Birds Brit. Mus., XXII. 1893, p. 268 ; Ogilvie-Grant, Man. Game-birds, I.
1895, p. 244; Blanf., Fauna Brit. India, IV. 1898, p. 87; Oates, Man. Game-birds, I. 1898, p. 379 ; Sharpe, Hand-
list Birds, I. 1899, p. 34; Oates, Cat. Eggs Brit. Mus., I. 1901, p. 52; Robinson, Jour. Fed. Malay St. Mus., I.
1906, p. 129; Robinson, Jour. Fed. Malay St. Mus., ILI. 1906, p. 56 (Pahang) ; Robinson, Jour, Fed. Malay St.
Mus., V. 1913, p. 16; Stuart-Baker, Jour. Bombay Nat. Hist. Soc., XXVI. No. I. 1918, p. 10.

BORNEAN CRESTED FIREBACK

Lophura ignita (Shaw)

hee movement ve eae back detaches itself from the spots of sunlight, and the blue-black
rom the dark shadows, and the bird stands out in its full glory.

WIVAXX aTAIG ~

ve 7 mn

5 AMAIA aarendd.

in: (way a Ae
babeda (llerie sit ii Bost SF yo Baines shiriny “yq tase

lo yliotwh s,aeagt moddls,asery : div. .boisiogsh

8 of Se0lyogle enw ti sy! Send “AYE “Svitein 6: fot’ 1aF 4S “eAsoWw ee [x 5 A

Stray Feathers, 11/9944) p. 481 Cheba @]
is wd “Skadup ol Iq Sud" Sat tht .

Soa § Fels.
dosld-suld-gdt-bas vadgiltineddatog# vadarsnoit Qsesi. 2 abisiti 90

te 2
he enay fs o
re

PS ERE ae ee ee 2 4 ; iy
Wt ati ni dug abnsia brid Sd). bag

i iol Hi aii

PLATE XXXVIII.

BORNEAN CRESTED FIREBACK.

BORNEAN CRESTED FIREBACK
Lophura tgnita (Shaw)

NAMES.—Specific: zgnzta, from the Latin zgzzs, fire, with reference to the fiery metallic back and abdomen
plumage. English: Bornean Crested Fireback; Fire-bellied Pheasant. French: Faisan a dos roux. German:
Borneo Fasan. Native: Sempidan (Sea-Dyak).

BRIEF DESCRIPTION.—Male: Plumage in general black, glossed over most of the feathers with purplish blue.
The back, rump, the entire belly and sides fiery bronze-red. Two inner pairs of tail-feathers, and inner webs of the
third pair rich buff. A long, bare-shafted, tufted crest. Facial skin blue, legs and feet reddish. Female: General
colour above rich dark chestnut, faintly mottled with black, neck paler chestnut; chin and throat white; breast
chestnut, edged laterally with white ; remaining under-parts dark brown, widely margined all around with white,
Facial area blue; legs and feet reddish.

RANGE.—Borneo in general ; introduced into the Island of Banka.

BEES BLN: Nes. HAW INES

THE home of the Bornean Crested Fireback in the interior of Sarawak is amid
low, swampy undergrowths, where the soggy, matted leaves give forth no sound to
the step of bird or animal. In such a spot in the afternoon when the round sun-
spots of noon have given place to long pencils of light, the splendid Firebacks come
forth from the maze of calamus and other thorn-plants among which they spend the
heat of the day and make their way to the little gravel water-courses which
meander down to the rivers.

I remember one such little glade, densely embowered with overhanging
vegetation. A rivulet trickles through it and between it, and the river is a dense
impenetrable tangle of calamus, bamboo and thorny fern. Here a family of Fire-
backs elected to spend the moulting season. Although not far from the cleared
ground around a native Dyak house, they have chosen wisely, for just beyond is a
Dyak burial-ground, mounds covered with matting and a palm-thatch shelter. On
the mat are dishes, and some Chinese jars of considerable beauty, filled with food
and water for the comfort of the dead. No one but a white “Tuan” who fears not
evil spirits would come here, so the pheasants are safe.

In such a place I watched silently on a certain 2oth of July. I was perched
between two great trunks, eight or ten feet from the ground, partly hidden by a
drapery of lianas. A pair of rufous cuckoos were toying with small twigs and sticks
just overhead, hinting that they have not reached the stage of complete cuculine
parasitism, but had in preparation the flimsy type of nest which some of this family
still fashion. They took no notice of me in their curious squawks and trills of
love-making—sounds so unlike their usual monotonous note. A broad-bill, with
plum-coloured waistcoat, black head and collar, and dark wings with yellow trimmings,
watched for and caught all the insects which flew past a branch in front. A crash

of twigs and small branches drew my eyes upward, and there I saw a tupaia or
VOL. II 129 S

130 A MONOGRAPH OF THE PHEASANTS

tree-shrew stalking its prey. How like and yet unlike a squirrel, moving at times
with the same jerky starts and flicks of tail, yet with a caution which told of the
hunter as well as the hunted. Now he spied what appeared to be a great beetle,
and slowly and imperceptibly crept forward. Finally, he leaped upon and secured
the insect, slipped, hung by one little paw, pulled himself up, spied me and burst
into a series of squirrel-like barks, which in depth of profanity, if not exact tone, did
credit to a Canadian red squirrel.

Sunbirds flitted everywhere, seeking for their insect prey in all likely and
unlikely places. A shama thrush flew to a branch and scanned me closely, with now
and then a graceful lift to: the tail. What wonderful organs of expression are tails
to birds of the thrush family! They take the place of eyebrows, hands, and mouth
in this capacity.

The tupaia had vanished after its vituperative outburst, but I was now delighted
to see on a slanting trunk a tiny tupaia, moving with infantile slowness, out on
perhaps its first excursion. It needed little teaching, for it munched greedily at
every ant which crossed its path. With a rush came a parent tupaia, seized its off-
spring by the neck or an ear, and rushed back with it, and up to a great mass of
felt-like bark and leaves, into which it carried the youngster.

The shama sang softly beyond the glade, and I answered with the opening notes
Whew! whe-whe-whe-whe-o! wheo-wheo! and with a flash of brown and white the
bird came in swift flight to a branch on my right. The white feathers of its lower
back were raised and its flirting tail spelled keenest excitement. Soon with high,
far-off ventriloquial notes, sounding twenty yards away, it ran down its liquid scale
and bubbled into the melody of which our cage-birds give but a parody. Before it
flew off, a half-grown youngster added his saucy note to that of his parent. His
wings were spotted; on his breast was a bar of black, and his throat was of
uncertain mottled hue. But he flirted his half-grown tail and scolded right shamily!

What absurd similes come to one in such an isolated place! Many of the tall
lily-like leaves about me had been injured by insects while they were yet in the bud,
and now that they were unfolded, the rows of larger or smaller holes were carried
completely across the leaf—exactly like the perforated music rolls of a pianola piano!

The ground on the little slope above which I was perched was partially dry,
and through the leaves flat, spreading fronds of moss-like selaginella extended. As
with so many of the low-growing plants, these were variegated, the tips of each
frondlet being pale green in fine contrast to the deep shade of all the rest.

Two small, brightly-coloured pittas passed through the glade, now close together,
throwing the leaves over their backs and eagerly seizing some morsel, now leaping
like jerboas over logs and branches.

A tricolour squirrel sprang from bush to bush behind me, and then down across
the rivulet by an aerial bridge. In passing he jostled the stem of a giant calamus,
and now from the depths came a weird rhythmic sssk! sss !—indescribably unreal,
a sound of some fairyland. Eyes and ears refused to locate it except far away, yet
I knew that at each surge of sound a thousand thousand tiny ants were shaking
with all their might, vibrating the hollow stems in which they dwelt. It was a
most soothing sound, and I sat for some minutes listening abstractedly. Then I

BORNEAN CRESTED FIREBACK 131

suddenly saw a bird where none was a moment before. It was broad daylight and
the sun was shining brightly through the foliage, but the splendid cock Crested
Fireback within twenty feet of me was with difficulty detached from his surroundings.
The iridescent blue and coppery-yellow were like deep shadow and a sun-lit patch
of leaves. Quiet as the tree-trunk near him, for a moment he scanned every inch
of the glade. By some miracle I escaped his eye, and then he walked slowly down
to the rivulet, drank, wiped his bill and picked at the clay bank on the farther side.

As he walked on, three young males and two females followed. Their short
spurs proclaimed the youth of the cock birds, and while doubtless one of the brown
hens was his mate, which it was I could not tell.

After all had drank, two of them daintily catching the moisture dripping from
the moss-ferns, all scattered along the rivulet gravel and began scratching in the
shallow water. Now and then they seized something—whether tiny crayfish or insect
larve I could not tell.

Once the broad-bill fluttered against a dead hanging vine and loosened several
large dried leaves which fell noisily. The cock uttered a mumbled Um-um! um-um !
and then a single sharp, keen whistle, which cut through the quiet air of the glade
like a knife. One by one the pheasants passed from view down the little ravine. I
had been vouchsafed a glimpse of a family of wild Firebacks and had not alarmed
them. The fates had been kind.

When the last of the big, beautiful troop vanished, the glade seemed deserted
indeed. In vain the sunbird sang his lay at my very elbow; even a mellow-
plumaged little trogon on a bough near by could hold my attention but a moment.
The King, with his Queen and Princes had gone—what were commoners now! I
slipped to the ground and stretched my cramped limbs, and began to creep into the
maze of thorny vines from which the birds had first emerged. Three steps on hands
and knees and the vague, indefinite dream-sound arose, this time above and around
me. Now that it was closer I seemed to detect a sinister note in its softness.
Instead of a single tone, it seemed a myriad blended into one, with hundreds
fraying out at the beginning and end. A sharp pain on my wrist caused me to
start involuntarily, and looking up I sawan army of vicious jaws and stings waiting
for my nearer approach. No human being could force his way to the centre of that
clump of harmless-looking underbrush. ‘Truly the secrets of this family of pheasants,
at least in this little ravine, were safe for the present !

GEOGRAPHICAL DISTRIBUTION

_ The Bornean Crested Fireback seems to be very generally distributed throughout
that great island, and wherever I have travelled it has always been by far the least
rare of all the Bornean pheasants. We know of specimens from many parts of
Sarawak, from FHlopura, North Borneo; Pleyharie, South Borneo; and Mount
Kenepai, West Borneo, besides many other localities. Its occurrence on the island
of Banka, separated from Sumatra only by a narrow channel, must be looked upon
undoubtedly as due to introduction by man. Otherwise it is difficult to explain its
absence from the intervening island of Billiton, on which I found no trace of the

132 A MONOGRAPH OF THE PHEASANTS

bird, together with the fact that the general fauna of Banka is wholly Sumatran
and not Bornean.

Such being the case, we can readily conceive of the possibility of hybridization
taking place either in birds kept captive by natives or as a result of a stray bird
finding its way into Sumatra and crossing with the indigenous 7wz/a. ‘This offers an
alternative explanation of such aberrant specimens as have given rise to the various
discussions as to the number of species in this group such as Bittikofer’s smatrana.
From my own observations I am satisfied that they may all be resolved into
variations of one or the other of the two closely related species rufa and Zgnita.

GENERAL ACCOUNT

I spent some time in and near the Dyak graveyard, which I have mentioned in
the bird in its haunts, and concentrated my attention on the family of Crested Firebacks
which there made their home. On the third day I saw them together and missed one
of the young males. The following day I stumbled by accident upon a few remaining
bones, including a leg and many feathers of this very bird, close to the bank of the
river. Of course it was impossible to tell the cause of death. A host of little workers
struggling with the last remaining tough tendons on the leg-bone, and two well-worn
paths to a near-by nest, showed that at least the ants had profited by this windfall.
Apart from this, tragedy did not visit the pheasants during the week or two during
which I watched them.

They roosted together in a tree of medium size, heavily draped with hanging vines
and surrounded by dense bamboo thickets. Once only was I able to see their forms
dimly silhouetted against the sky. Three were perched close together, the others
somewhat scattered. Early in the morning—how early I never learned—they left their
roost and went towards the river, at least I found them riverwards on three separate
occasions early in the forenoon. From the tracks I learned that they had no regular
route, but sometimes followed the gravelled rivulet, or again went through the marshy
forest on either side. Concerning their forenoon occupations I could obtain no
information, although I should say that they fed and scratched near or along the
river bank.

During the heat of midday, twice they returned to the shelter of the bamboo
undergrowth near their roost, once they remained quietly in the shade of a tree on
the very bank of the river, and this, while several Dyaks were working on a canoe
in full sight, but some distance up-river on the opposite sandbank. On the other
days they evaded all my efforts to find them. I had chosen my look-out wisely,
although it was the general jungle life which came to the river that I had hoped. to
see, and the pheasants came rather as a surprise. A thick snarl of grape-like vines
had grown entangled among the branches of a stout tree, and with a little cutting
and arranging I was able to lie outstretched well above the high bank, partly over it
and partly over the water itself. The shadow from the thick mesh of bare stems had
discouraged the growth of foliage beneath, so that my view downward through the
interstices was almost clear. Below me was a narrow cleft in the bank where the jungle
rivulet trickled into the waters of the river. Bulbuls and purple butterflies amused

PHOTOGRAVURE 27

nove OF ete BORNEAN CRESTED FIREBACK

Tui home of this bird was reached by means of a seventy-foot Dyak war canoe, in which | was
paddled ip the rivers to the inland jungles. Here, in the haunts of gaily coloured pittas, of glittering
sunbirdsand the glorious tropical orchids, these birds live and court their mates and rear their young.
Often tk first hint of their presence is the call of the cock, a low, mumbled Um—um/ Um—um!

followed by a single, sharp, keen whistle, which cuts through the warm, quiet air like a knife.

sniisitile Yo @Ellic bowoloo ylise to aigusi. ody ai:
3 Ag ine

b gee ad &

-BauOY risds sot _bns eossen tiod? 44309. brs <oyil ebid, sen

esw | doidw oi sonsa isw AsyG tose

BORN

BAN CRI

Bey Jk

ar

PHOTOGRAVURE PLATE 27.

Libs

ii

yet ee
a
Ha ;

BORNEAN CRESTED FIREBACK 133

me for a time, and I had made some interesting notes on river birds, when an old
wild sow with a half-grown young one came slowly along the edge of the water.
Hardly had they appeared when the Dyaks on the other shore had occasion to hammer
smartly on their upturned canoe. The mud-caked porcine matron whirled facing
the sound, erected her tousled ears, and concentrated her whole being in obtaining
information of the danger through her quivering nostrils. They told her nothing,
and her little pale-winkered eyes evidently carried no image of the distant Dyaks to
her brain. The sound was not repeated, but the memory of it remained vivid, and
the habit of caution of a lifetime and of untold other lifetimes in the past, so deeply
implanted in that porcine brain, conquered, and she turned with a low grunt, and with
her offspring at her heels trotted back out of sight. My pheasants were the next. to
appear, as I have said, wholly unexpectedly, and after pecking vaguely about for a
time, they collected in a compact group in the dense shade of a river plant, and there
for half-an-hour held low converse with each other, preening their plumage the while;
at least they kept up a low confidential clucking or mumbling, which, to my ears,
indicated supreme contentment. When they moved off into the jungle again, I
descended, as I was slowly baking in the sun, and the network of liana stems on which
I lay was fairly hot to the touch.

I caught many another sight of the birds on succeeding days, but apart from the
ever-vivid thrill which their appearance produced, I found them as uninteresting as
domestic fowls. They seldom uttered a note; they pecked and scratched, and ate and
preened and slept! It was the season of moult and I could expect little more from
them than I observed. Once only I heard the full vocal and wing challenge—a subdued
wooonk-k ! (whirrrrr)—wooonk-k ! (whirrrrr). Indeed, I got more from them in the
way of practice in stalking and knowledge of what I could do when in the presence
of wild pheasants without being observed, than I acquired valuable facts of their life-
history.

I was impressed again and again with their invisibility when perfectly quiet and
the ease with which one could distinguish them even in the dimmest recesses of the
forest when any part of their body was in motion. Thus I came to realize the
tremendous importance of their method of movement—quick, sudden jerks of head and
neck—a single motion if possible—in snatching a beetle or other insect, and then a
period—a much longer extent of time—of absolute immobility. This reduced the
perceptive periods of danger to the shortest space of time, and gave to the birds all
possible opportunity for the detection of danger, they themselves at the same time
being least perceptible to possible watching enemies.

Watching the full-grown cock Fireback feed for ten or fifteen minutes at a time—
as I did once—I was impressed with the fact that such a period is divided into two
very unequal parts, first, the search for, snatching and swallowing of food, and second,
the looking out for danger. It seemed as if a few seconds less to the first division
would result in starvation; perhaps a single second deducted from the latter might
bring destruction! Never for a moment was there any relaxation; any sensible period
of transition from one to the other. It was complete concentration on an insect,
discovering it, seizing it, and then at the very moment of swallowing, the head and
neck would shoot up, and the keen eyes were on the watch; twin scarlet search-lights

134 A MONOGRAPH OF THE PHEASANTS

of intelligence, with no need for the head to turn, as would be necessary in ourselves
or in such a pursuer as a civet cat, but so set by nature on opposite sides of the head
that they took in almost full measure of the surrounding circle of three hundred and
sixty degrees of possible danger. When I gave up expecting battles and courtship
displays and other intimate doings of the lives of my well-armed but moulting
pheasants, I gave close attention to such little details as I have mentioned above, and
soon saw that there was a perceptible difference in the behaviour of the old and the
young males. The latter were apparently full-grown in size, and as their moult was
about over, they differed in general appearance hardly at all from their parent. Only
the short spurs, and a few remaining juvenile feathers stamped them for what they
were. But a more intensive study showed many traits which set them apart, although
no signs of affection or of actual dependence were ever noticed. These young pheasants
were far from being as much on the alert as the old bird. They had many moments
of abstraction, of idly doing nothing, and although they invariably ended by shooting
up the head and neck to full attention, yet the few seconds which preceded this would
have quickly spelled death if such a method was persisted in after they began to shift
for themselves.

This observation led me to wonder if I could not distinguish between the females
by some such mental field mark. So the next time I saw the birds I devoted myself
to solving this point. I succeeded at the expense of much bodily suffering. To be
able to get near the birds I crawled under one of the Dyak grave mattings, and bending
it over in front of my face I had an ideal point of vantage.

But I soon found that one may not lie quiescent for hours upon a Dyak’s grave
with impunity. I had disturbed a populous city of ants—innocuous as to bites or stings,
or I should not have been able to lie still for a moment—but afflicted with an inordinate
curiosity as to the meaning of this strange intruder. They certainly left no portion
of my clothing unexplored, and it took all the will-power I possessed to restrain myself
and disregard the scrambling host. Then I realized that certain nerves in my body
were going to sleep under the unaccustomed pressure, and it was not until I rose and
shook the dirt from my person that I learned the fact that every centre of drowsy
sensation had been a hungry leech who had feasted to repletion while my attention
was distracted by the pheasants. But with all the discomforts of my afternoon’s vigil
I felt well repaid for succeeding in my desire, and the pleasure of having penetrated
ever so slightly into the mental life of these splendid birds soon eclipsed all
remembrance of my small but active tormentors. The first time the birds approached
I feared they would pass me some distance away, too far for careful watching, but
luckily one of the young males pursued some flying insect in my direction, and then
found a feast of plenty, and his vigorous efforts in scratching among the leaves attracted
the attention of the remaining birds, and all were soon gathered in the same spot.
I had an excellent chance of observing the two females, and within a minute’s time
I had fully made up my mind which was the older—the probable other parent. Even
she, however, did not quite equal the adult male in constant alertness. I fancy that
this was not individual, but sexual, a phenomenon which I afterwards noticed more
especially in the behaviour of the white-tailed pheasants (Lodcofhasis) in the face of
danger. ‘The hen’s very protective plumage seems to demand less constant watchfulness.

BORNEAN CRESTED FIREBACK 135

The young female—having the added cause for negligence of parent protection, was
positively regardless of what might threaten. She gave herself wholly to the pleasure
of eating, with only now and then a perfunctory glance around, with no motionless
periods whatever, but scratching vigorously all the while. It is exceedingly interesting
that while in the adults continual consciousness of danger should be so acute,
nevertheless it should be manifested in so defective a manner in nearly grown birds,
in which total independence has not as yet called out the need for the perfect exercising
of this function.

I have mentioned several times the habit of scratching among the leaves for
insects, and that I saw the birds again and again taking small insects or other little
creatures from leaves. When the birds are feeding on the move, to or from the river,
they seem chiefly to look for such food, but when more or less settled down in one
spot, I noticed them more often plucking at the leaves and young grass-like growths.
The crops of a number of birds indicated that seeds and leaves were the main source
of food supply, while it was only occasionally that insects actually predominated. The
most common proportion was a goodly number of seeds of various kinds, a less amount
of leaves and comminuted vegetable matter of a similar nature, with a scattering
indication of small insects. These latter consisted almost entirely of ants and small
beetles, either small iridescent flower beetles, or grubs and ground coleoptera. An
acorn-like fruit was found in several cases, called by the natives dompelt.

CAPTIVITY

One day on the Upper Rejang River in Sarawak, a Dyak brought me two Crested
Firebacks, a cock and a hen, in coarse woven native ‘‘quakes” or round baskets. They
had been trapped together at the edge of a paddy field, and the savage said through
the interpreter that they were “man and wife.” I took this as a good omen, and as
it was near the end of my stay I did not sacrifice them for the sake of science, but
placed the birds in more comfortable quarters, and in the semi-darkness of half-covered
cages began to accustom them to a diet of paddy and what other vegetables I could
procure. They soon became rather quiet, and in fact within a week would show but
little fear even when the food dishes were being cleaned and filled. The cock, at first
pecking fiercely at my hand from anger, soon appreciated that each time I approached,
a large fat insect was dropped within his cage, and he soon took the grasshoppers and
beetles direct. They were such model captives that they had a most wholesome effect
on my other birds, and ‘did much to quiet two particularly nervous Argus females and
a covey of wood quail. On the down-river journey they were perched high up on the
matting roof of the war canoe, and made the journey safely. Finally at Singapore they
started for their final destination in most elaborate crates of teak-wood fashioned by
the skilled hands of Chinese carpenters.

Several. weeks later, three of the Bornean Crested Firebacks were turned out in
a roomy aviary at Bernardsville, New Jersey, where they stretched their wings, and
preened their plumage in comfort. The ‘‘man and wife,” if such they had been, would
have none of each other, but the splendid cock bird cast his affections upon a young
hen which was in the same lot. These paired off, and made things so unpleasant for

136 A MONOGRAPH OF THE PHEASANTS

the other pheasants in the paddock that all had to be removed. The hen then made
a rough hollow under a dense bush, lined only with the bent, dead grass stems which
happened to be growing on the spot. Here at intervals of one and two days she laid
six eggs, and on these she sat faithfully for twenty-four full days. This period of
incubation lasted from July 21 to August r4. The cock never took his turn upon the
nest, but was very faithful, fairly wearing a path in his circling guard duty several
yards distant from the spot. I never saw him approach closely to the nest, but he
was very pugnacious, and never failed to make a pretence of attacking any one who
approached the place where his mate was sitting.

The five young chicks which hatched, left the nest within a few hours after
emerging from the shell, and kept very close to their mother, hardly leaving the shadow
of her body for the next two days. As soon as the chicks had left the nest the cock
joined his mate, and from this time onward they were constantly together. The cock
took his full share in feeding the young birds, calling them to him and giving them
small morsels as frequently as did the hen. Whenever they crowded under him,
attempting to brood, however, he apparently did not understand what was expected
of him, as he would walk slowly away, lifting his feet very high so as not to crush
the little downy creatures. His attitude towards danger also changed with the sudden
increase of his assiduity in caring for his offspring. When any one approached he
slunk away, holding body and tail low, making himself as inconspicuous as possible.
The chicks at such a time behaved much like young grouse or quail, separating at the
first note of danger from the mother, pushing deep into the grass, or squatting flat
as their little bodies would allow if no shelter was near. The hen pheasant presented
a complete antithesis to these tactics. Instead of slinking off or of attempting to hide,
she ruffled up every feather on her body, her wings were half spread and all a-tremble,
her red eyes dilated and fairly blazed with righteous fury. Then with shrill squeaks—
apparently too full of emotion to voice her feelings in any louder way—she would
fairly hurl herself at the intruder, sometimes striking one headlong as high as the
knees ; then, falling to the ground, she would cling with feet and claws to one’s shoes,
wings spread and drawn around in a semi-circle, like an infuriated owl, pecking with
all her might at shoes and ankles. She presented a picture of absolute fearlessness
and mother fury. Pushed away with one foot, she returned at once with undiminished
vigour, and not until one had beaten a retreat did she slowly and reluctantly return
to where her young had implicitly obeyed her warning, remaining crouched and
silent.

In about a week the young birds were able to use their wings with considerable
skill, and from this time on, when alarmed and given the danger signal, they flew off
in various directions, radiating out on every side except that of approaching danger
to the limits of their paddock before settling and crouching. In two or three weeks
the hen accepted the occasional visits of human beings as a necessary, normal event,
and the moment she ceased warning her brood, just so quickly did they, too, cease
to show fear, and all became reasonably tame. The year following this, the same cock
and hen bred again. After the first autumn moult the young birds are superficially
hardly distinguishable in colour of body plumage from their parents.

BORNEAN CRESTED FIREBACK 137

DETAILED DESCRIPTION

ApuLT Ma.e.—Feathered parts of the head, with the exception of the crest, very
dark dull brown. Crest composed of a number of feathers, each showing a dense tuft
of long, loose terminal barbs at the end of an elongated bare shaft. The neck plumage
is also loose and rather disintegrated, and, like the crest, is strongly glossed with
steel-blue. The breast, mantle, scapulars, wing-coverts, inner secondaries and upper
tail-coverts are dull black on the basal portion, and strongly glossed with metallic
purplish-blue over the distal part of the feathers. Most of the feathers of the dorsal
plumage show a more or less distinct terminal fringe with steel-blue sheen, this
character being especially marked on the upper tail-coverts, where the fringe is quite
long and disintegrated. When a single upper covert is examined, the loose terminal
fringe seems to be separated from the main proximal part of the feather by a narrow,
velvety black line. Manipulation, however, shows that this is merely an optical effect
caused by a changed incidence and refraction of the rays of light. This results from
a sharp upward bend in the vane as a whole—a narrow transverse wrinkle clearly
evident both to the sight and touch. Posterior to this bend the feather is normal,
closely connected into a solid vane. That portion of the barbs forming the fringe,
from near the bend to the tip of the feather, while possessing the usual number of
barbules, are wholly without barbicels, the barbs thus standing apart in the loose
detached fashion so characteristic of this portion of the feather in many pheasants.

Back and rump fiery metallic coppery bronze, decidedly darker than in the Malayan
bird, more red than golden. Examining a typical feather from the lower back we
find it measures about 80 mm. in length. Of this forty, or a little over half, consists
of the basal disintegrated grey down. Then the contour portion begins abruptly, a
zone of some 20 mm. dark brown changing to black, the distal portion glossed with
steel-blue. The terminal 20 mm. is divided into a shorter portion of dark, dull maroon,
giving place abruptly to the long (7 mm.) fringe of glowing bronze red. There is no
trace of the fost-jimbrie wrinkle. Proceeding outward, the secondaries lose first the
bluish gloss, and then change from dull dead black to dark brown. The primaries
are still paler, sooty brown. The two inner pairs of tail-feathers and the inner web
and tip of the third are clear rich dark buff. The bases of these inner feathers are
dusky, and all the remaining rectrices are unglossed dead black.

Beneath, the purplish-blue breast zone ends abruptly, and the entire lower breast,
abdomen, sides and flanks are of the same fiery metallic bronze red as the back, except
lighter. In extremely coloured individuals the feathers of this entire area lack black
pigment, the main portion of the vane being dark chestnut. Usually, however, the
feathers down the mid-line are at least half black, and variations of this colour may
be found up to where the chestnut and metallic red colour is confined to flank patches.
I collected one specimen of this extreme variation in the hinterland of Sarawak. The
amount of black pigment present in these ventral feathers is wholly individual. In
two male birds from the same set of eggs, hatched and reared by their mother, at the
moult into the first-year plumage, one showed solid chestnut on the posterior ventral
surface with only extreme basal black mottling; the other had only the margins

chestnut—comparable in extent with the white margins on the ventral plumage of the
VOL, II T

138 A MONOGRAPH OF THE PHEASANTS

female. And this indeed only on the sides and flanks, the mid-breast and abdomen
feathers being wholly black or with two small lateral spots of chestnut. Thighs and
under tail-coverts dull black, glossed with greenish-blue on the terminal portion of the
feathers.

Irides red; legs and feet in the breeding season reddish, fading at other times
to a fleshy pink; facial skin bright blue at the period of courtship, considerably paler
during the rest of the year; mandibles horny white; spurs and claws yellowish horn,
The spurs are unusually strong, long and slightly curved, reaching a length of 35 mm.
or more. Weight about 44 lbs. Length, 650; extent, 845; bill from nostril, 21;
wing, 271; tail, 253; tarsus, 111; middle toe and claw, 58 mm.

VARIATION IN MaLe PLumace.—To show the variability of even a single character
in birds supposed to be sufficiently homogeneous and distinct to form a esparate species,
let us consider the white colour of the tail-feathers of the males upon which Biittikofer

bases his sumatvana.

His specific characterization is ‘Similar in colour to ZL. iguz¢a, with the exception
of the centre tail-feathers, which are pure white, with a faint ochrous tinge on the basal
part as far as it is hidden by the upper tail-coverts” (‘‘ Notes from Leyden Museum,”

XVII. 1895, p. 178).

(a) Leyden Museum specimen.—Central pair of rectrices and inner web of second pair white.—No locality.

(6) Leyden Museum specimen.—Two central pairs of rectrices and inner web of third pair white—Palembang,
Sumatra.

(c) An “albinisme” male from Amsterdam Zoological Garden.—No mention of white on tail-feathers.

(zd) Type male in Brussels Museum.—Dubois describes the tail colour as follows: “Queue noire, les quatre
rectrices médianes d’un blanc roussatre, la barbe interne de la paire suivante également d’un blanc
roussatre ” (“ Bull. Ag. Belg.” (2) XLVIL., p. 825). The ochraceous hue of the basal concealed portion
of the feathers is so pronounced that Dubois considers the whiter exposed area has been faded out
to its present colour.

(e) Vienna Museum specimen.—Said to agree with the two Leyden specimens, which, as they differ in the
amount of rectrice white, is obviously impossible.

(f) British Museum specimen.—This much-described individual (Sclater, “Proc, Zool. Soc.,” 1863, p. 119;
Grant, “ Brit. Mus. Cat. Birds,” XXII. 1893, p. 289; Grant, “ Hand-book Game-birds,” 1895, p. 247)
is classified by Bittikofer under his swmatrana. Of the white in the tail Sclater says, “vectr. 4 mediis
albis” ; Grant, “middle pair of tail-feathers white,” but both are wrong, for as the inner and third
pairs are white and the second pair is missing, the inner three pairs of rectrices were undoubtedly
originally of this colour. The other character of red upon the flanks seems to be almost as variable
and from the fact that in this British Museum specimen the golden hue of the sides is almost white
on the anterior portion, I should be inclined to think that the “albinisme” of specimen ¢ was but an
extreme aberration due to hybridism or abnormal variation.

ADULT FEMALE.—Entire upper surface, neck all around and breast chestnut, the
fore-neck and occasionally the hind neck and upper mantle with white lateral margins
to the feathers. Crest with terminal plumes as in the male, but with much shorter bare
basal shafts, and slightly darker chestnut than the head. Short feathers of the head
and upper neck, chin and throat somewhat recurved. The chin and throat are white.
From the rear mantle and wing-coverts backward there is a constantly increasing
mottling and vermiculation of black pigment, causing the chestnut to appear darker
and darker as we proceed toward the tail. On the outer webs of the secondaries and
primaries, and on the shorter and median upper tail-coverts, the chestnut and black are
of about equal proportions. The inner webs of the flight feathers are dark brown,

BORNEAN CRESTED FIREBACK 139

paler in the primaries. The tail-feathers are quite variable; sometimes the two central
pairs and the longest tail-coverts are heavily freckled or mottled with dark chestnut—
though this is always subordinated to the black (the reverse pattern of the anterior
dorsal surface)—while again there will be hardly a trace of the lighter colour, all the
rectrices being uniformly black.

The entire lower breast, abdomen, flanks and anterior thighs sooty black, broadly
margined all around with white. In the disintegrated, downy feathers of the lower
abdomen the white margin encroaches upon the black centres, turning them impure
brown and restricting them to a small central area. The under tail-coverts and the
posterior sides of the thighs are dull black, tipped more or less strongly with dark
chestnut.

Irides red; legs and feet yellowish-white; facial skin blue, duller than in the
male; mandibles horny-white; spurs sharp but abortive, only about 3 mm. in length.
Weight 34 lbs. Length, 580; extent, 778; bill from nostril, 18; wing, 258; tail, 190;
tarsus, 89; middle toe and claw, 50.

NataL Down.—Four day old chick. Warm rufous on crown and nape, becoming
black on scapulars and entire back. Rufous buff bar across centre of wing. Latero-
dorsal longitudinal bands almost obsolete. Face pale rufous, a black line obliquely
back from the orbit across ear-coverts, ending just beyond. Chin, throat and belly
buffy white, a strong orange-buff tinge across breast and down sides and flanks.
Primaries just appearing at the tips of the sheaths.

A THREE WEEKS’ OLD CHICK is much the same, except for great development
of wings which cover the entire body, extending well beyond the tail. Coverts and
flight-feathers dark brownish black, the former with buffy tips and a jet-black
sub-terminal band. The inner secondaries with very slight buff edgings, other flight-
feathers unmarked. At this age, seven primaries and eight secondaries are functional.
Primary No. 8, although over 13 mm. in length, is still ensheathed, and Nos. g and 10
are as yet invisible.

The tail-feathers have just broken their sheaths, bearing aloft the pale rufous
down.

JuvenILE Prumace.—In a chick of seven weeks—four weeks older than the one
above—considerable change has taken place. The entire body and the rear crown are
clad in short but fairly complete juvenile plumage, while the head and neck are still
downy.

On the rear crown, sharply limited on all sides, on or just anterior to the place
where the adult crest will appear, is a dense triangular patch of loose chestnut feathers,
somewhat shaded with black. The entire hind neck is downy, but the mantle and
back are densely covered with a growth of rich chestnut, black mottled feathers. At
this stage the sexes are distinguishable, the males having the dorsal plumage much
darker than the hens.

The eight primaries are growing strongly, the eighth still lagging far behind, about
half as long as the rest, while Nos. 9 and 10 are just detectable as two minute sheaths

140 A MONOGRAPH OF THE PHEASANTS

about 6 mm. in length. The tail of sixteen almost black feathers is growing rapidly,
and from the outer pairs inward, the three outer pairs having already completed their
growth.

A male in a somewhat later stage shows the full juvenile plumage, the whole top
of the head from the cere back now being feathered. Down the nape and hind neck,
however, there is only a very narrow plumaged area, two lines of feathers only, all
the rest of the hind and side neck being still covered with a dense growth of the orange
rufous down. The crown feathers are black with very wide chestnut tips, which,
however, are absent on the nape and narrow nuchal line, these showing slight bluish
glints among the black. The dorsal body plumage—mantle, back and rump—is basally
dark brown, with an indistinct black sub-terminal bar, and a wide dull rufous mottling
at the extremity.

The scapulars and wing-coverts show this pattern to a very pronounced degree,
and thus present a very different appearance, the characters most apparent being a
rich rufous tip or terminal band, followed by a black band or two lateral black ocelli,
The inner secondaries show a very slight rufous marginal mottling, but the flights and
rectrices are otherwise plain brownish black.

The under-parts exhibit a fairly dense growth of white (eatHers on the chin and
throat, but the lower neck has only a very sparse covering of feather sheaths, which
show the skin almost bare. The ventral plumage is of black-centred feathers with the
loose webbing margined with grey or white—a most indescribable mass of downy
plumage with no regular character or pattern.

The post-juvenile moult has just begun in the primaries. No. 1 is well grown,
and Nos. 2 and 3 have just been dropped. Nos. 9 and 10 are still growing, and even
No. 7 is not yet dry.

The tail is about in the same condition, or a trifle more delayed. All the juvenile
feathers are full grown, but the outer pair have been dropped, and the new black
incoming feathers already measure 33 mm., with about 12 out of the sheath.

TRANSITION BETWEEN JUVENILE AND First YEAR PLuMAGE.—Three males
collected in August are all in about the same condition, half moulted into adult.
dress.

. In all these birds the long, rufous crown feathers stop abruptly at the hinder
crown, their rear edges overhanging the shorter posterior feathers like a cap. From
directly at their posterior edge the first few crest feathers are sprouting. The occiput,
nape, chin, throat and neck feathers are short and recurved; those of the occiput,
ear-coverts and side neck are dark brown; the chin and throat white, with a sprinkling
of new black feathers. The nape and hind neck show a wide border of shining greenish
blue, not the purplish blue of the fully adult. The change shown by the moult of
the wing-coverts and flight-feathers is abrupt, from the faded brown, mottled-tipped
old feathers to the new brownish black ones. The under-parts show as yet little change
from the chaotic, near-female, brown, grey and white of the juvenile plumage, except
along the sides, where the golden rufous shows as two broad solid bands.

In all three individuals the stage of the wing moult is almost identical. The
inner five primaries are new, No. 5 being still ensheathed, while the outer five are

BORNEAN CRESTED FIREBACK I4I

full grown but unshed. The outer secondary is still unshed; the second is full grown.
The next five are new, more recent as we proceed inward, until the fifth is merely a
short unbroken sheath; the next inner six are unshed.

The persistence of the first secondary is very remarkable. Not only in these birds,
but in another of seven months, which in many respects is full plumaged, the right
outer secondary still persists, the left having just been shed. The succeeding nine
secondaries are full grown, the next three being, like No. 1, still old, unshed
feathers.

Another focus of delayed growth is found in the central rectrices. In the three
males just described, the three inner pairs of tail-feathers are still unshed, juvenile
feathers, although a@// of the others are new and well advanced. In all there is a
regular gradation in growth from the outer to the fifth pair, the latter being only
25 or 30 mm. beyond the tip of the sheaths. The narrow, brown, worn tips of the
three inner pairs extend some 50 mm. beyond the longest new feathers, giving all
the birds a most curious appearance. At this age the spurs are about 5 mm. in length,
sharp, thin, triangular scales.

The measurements of the five stages of growth in male birds which I have
described, together with those of the adult male are as follows, tabulated for easier
comparison.

Bill from Middle Toe
nostril Wing Tail Tarsus and Claw
Four day chick . ; : : ; : . 6 ZO — 23 16
Three weeks chick . : : : j . 8 78 IO 25 18
Seven weeks juvenile ‘ : ¢ : ' II 129 66 35 25
Older full juvenile . : . , ‘ s 15 210 137 68 46
Transitional, juvenile to first year . ‘ : 18 235 Juv. 168 86 51
New 127
(Not full grown)
Adult male... : 7 é : ; ‘ 21 271 253 III 58

IMMATURE FEMALE.—The sexes are well marked first in the early juvenile plumage
when the mantle and breast begin to show a preponderance of dark colour in the male,
or rufous in the female. The head, nape, mantle and lower throat in the latter sex
moult into almost solid chestnut, shading into white on the upper throat and chin.
The black ocelli or sub-terminal band on the scapulars and wing-coverts are much
more conspicuous than in the male, owing to their pale, rich chestnut setting. The
secondaries also show the chestnut mottling much plainer in the female sex.

The ventral surface of the immature female is more definitely coloured, the regular
pattern showing dark brown centres, broad white lateral fringes and an equally
well-marked terminal band of chestnut. This pattern extends well down the sides
of the body, but on the lower breast the white begins to eclipse all other hues, until
on the belly it wholly replaces them.

One young female shows the inner six pairs of rectrices still unshed, while the
two outer pairs are represented by tiny sheaths. In another the three central pairs
are still unshed. All show the same gradual moult from outside inward as in the
males.

142 A MONOGRAPH OF THE PHEASANTS

SYNONYMY

Phasianus ignitus Shaw and Nodder, Nat. Misc., IX., pl. 321; Gray, in Griff. ed. Cuv., III. 1829, p. 30.

fire-Backed Pheasant Lath., Gen. Syn. Suppl., II. 1801, Pp. 274,

Gallus macartneyi Temm., Pig, et Gall., II. 1813, p. 273; Temm., Pig, et Gall., III. 1815, p. 663.

Gallus macartnyi Steph., in Shaw's Gen. Zool., XI. 1819, p. 218 [Part].

Flouppifer ignitus Guérin-Méneville, Icon. Rég. Anim. Ois. 1829-38, p. 26, pl. 3, fig. 3.

Euplocomus macartneyi Temm., Pl. Col. V. in text of genus Lophophorus, 1830, p. 3.

Euplocomus ignitus Gray, List of Birds, Part III. 1844, p. 26; Muller, Verhandl. Land-en, Volkenk, 1839-44,
p. 376 [Banjarmasin]; Low, Sarawak, 1848, p. 411; Sharpe, Ibis, 1879, p. 270 [Lawas and Mengalong Rivers] ;
v. Pelz, Verh. Ges. Wien, XXIX. 1880, p. 531; Sharpe, Proc. Zool. Soc., London, 1881, p. 800 [Sandakan] ;
Nichols, Ibis, 1883, p. 90 [Silam]; Elliot, Auk, VIII. 1891, p. 15 [Kinabataugan R., N.E. Borneo].

Gallophasis ignitus Gray, Gen. Birds, III. 1845, p. 498.

Euplocamus nobilis Sclater, Proc. Zool. Soc., London, 1863, p. 118, pl. XVI.; Sclater, List of Phas. 1863,
p. 7; Gray, List Gallinae Brit. Mus. 1867, p. 351; Sclater, Proc. Zool. Soc. London, 1868, p. 261; Gray, Hand-
list Birds, IT, 1870, p. 259; Elliot, Proc. Zool. Soc., London, 1871, p. 138; Walden, Ibis, 1872, p. 382; Elliot,
Mon. Phas. II. 1872, pl. 27; Salvad., Ucc. Borneo, 1874, p. 306; Elliot, Ibis, 1878, p. 414; Sharpe, Ibis, 1879,
p. 234 [Sarawak]; Sclater, Ibis, 1880, p. 371; Guillem., Proc. Zool. Soc., London, 1885, p. 416 [Silam, Sandakan] ;
Everett, List of Birds, Borneo, 1889, p. 199 [N. and S. Borneo].

Lophura ignita Ogilvie-Grant, Cat. Birds Brit. Mus. XXII. 1893, p. 288.

LOBIOPHASIS
WHITE-TAILED WATTLED PHEASANT

MAP IX.

100°

tee Mee

TS

Canton
SRN I a Wate Oe Gelade Ob
,

of
g

OF

F BENGAL
7 Madras i

Sa

EQUATOR

INDIAN OCHAWLN

ARABIAN SEA

=

East of Greenwich
H.F.& G. Witherby, Publishers. / Stanford's Geograph! Estab*.

MAP SHOWING THE DISTRIBUTION OF THE WHITE-TAILED
WATTLED PHEASANT.

Lobiophasis bulweri.

LOBIOPHASIS
WHITE-TAILED WATTLED PHEASANT

Family PHASIANIDAE
Subfamily PHASIANINAE

Genus LOBJOPHASTS

Tue remarkably specialized Bornean pheasant which alone forms this genus
finds a quite natural position at the extremity of specialized radiation of the Gexnaeus-
Lophura group. As the maximum of plumage development has found vent in the
secondary wing-feathers of the argus and the tail-coverts of the peacock, so in the
Bornean Wattled Pheasant it is the tail itself which, in number of feathers, has
exceeded all bounds for the family, and in the male consists of more than thirty
feathers.

No crest is present, but much of the head is bare of feathers, with the addition,
in the male, of three pairs of highly developed wattles, a small pair on the lores on
each side of the base of the culmen, a much larger pair on each side of the occiput,
and an extremely long, more or less pendant, pair on each side of the throat. All
three pairs of wattles are visible in the female in a rudimentary condition.

The 1st primary is very much shorter than the 2nd, which is equal in length to
the roth. The 5th is the longest in this series.

The tail in the fully adult male is composed of thirty to thirty-two white feathers,
the variation in number having nothing to do with age, but being a wholly individual
character. The central pairs are extremely curved, and more than twice the length of
the outer pairs. On many of the latter the web is defective or absent from much
of the terminal portion of the shaft, which is thickened and spine-like. In the tail
of the female there seems to be always twenty-six feathers, of more equal length
than in the male.

The spur is short and stout in the male, rudimentary in the female; the tarsus
is considerably longer than the middle toe and claw.

LOBIOPHASIS
Type.
Lobiophasis Sharpe, Ann. Mag. Nat. His. (4), XIV. 1874, 'p. 373 A . Lz bulwert.

The genus consists of but a single species, the White-tailed Wattled Pheasant,
Lobiophasis bulwert Sharpe, and there is little chance that a second exists. As far
as we know, it is confined to the central parts of Borneo.

VOL. II 145 U

WHITE-TAILED WATTLED PHEASANT
Lobiophasis bulwert Sharpe

NAMES.—Generic: Lobzophasis, from the Greek JoBdc, lobe or wattle, and gac.tavds, pheasant, from the very
highly developed cephalic wattles. Specific: du/werd, after His Excellency H. F. Bulwer, Governor of Labuan.
English: White-tailed Wattled or Bulwer’s Pheasant. Native: Blaiou (Kayan and Dyak, Sarawak); Bau-eu
(Dutch Borneo); Bagier (circa Mount Dulit).

BRIEF DESCRIPTION.—Male: Neck all around and upper breast dark crimson ; remaining plumage black,
all the feathers narrowly margined with steel-blue. Upper tail-coverts and tail pure white. Bare skin of head and
wattles bright blue; legs and feet red. The immature male has the tail coverts and tail chestnut. Female:
Brownish-~-buff above, finely mottled with black ; lower plumage rufous similarly mottled ; tail clear chestnut.

RANGE.—Central Borneo.

THE WHITE-TAILED PHEASANT IN ITS HAUNTS

As a rule animal life is not abundant in the Bornean jungle, compared with
the great primeval forests of South America. Occasionally, however, I was treated
to most interesting sights, and such days will always remain most vividly impressed
on my mind. Once, far up near the foot-hills of the mountains, on a tributary of
the upper Rejang in Sarawak, I had pushed on around a sharp bend in the river,
and was preparing to work up through the rapids ahead, when one of my Kayan
paddlers said through the interpreter that he and his companions had heard of
the death of a sub-chief at a village on a side branch of the river, and they would
all like to go and pay their respects to the deceased—in words, half English, half
Malay, to that effect. I always made it a rule to comply with all the little cere-
monies of my savage helpers, and to allow any tribal customs, religious or otherwise,
and in return I found they always gave me most honest, efficient service. I was
rather disappointed at having to stop, but as the afternoon was half over we could
not have made much more progress against the boiling, yellow rapids just ahead.
So I gave my permission, and the canoes were quickly beached on a sandbar.

The Kayans with their great truncated, sword-like knives cut down the under-
growth and erected two little raised shelters for me and my Malay servants, well
out of the reach of any possible sudden rise of water. Having made certain that
Tuan needed nothing more that they could provide, they unpacked their small bundles
of possessions and proceeded to adorn themselves with what pieces of black cloth
they possessed, and arrayed at last, they dropped swiftly out of sight down-stream,
and I was left canoeless in the heart of Borneo. I knew I could trust the savages,
however, and began at once to cast about for a way of getting into the forest which
began at the very bank of the river. The undergrowth was extremely dense, but at
the second circling I intersected a low game-trail. I fear it was but a wild-boar

path, and a “pig-high” trail was never intended for the passage of a six-foot
146

PLATE XXXIX

WHITE-TAILED WATLED PHEASANT

Lobtophasis busert Sharpe

Percuep in a tiny half-cave in the rocky side da deep gorge in central Bees eched! ihe
flying lizards, gorgeous as butterflies, pass swiftly from|ree to tree, while long-tailed paradise flycatchers
swooped after flying insects, in and out of the shadow\overhead. Without warning there stepped into
view three chevrotain, the tiniest of jungle deer, and wi them a quartet of White-tailed Pheasants, one
a fully adult male with great, spreading tail sweeping Ce hen walked Stone, beneath me, and:

still unalarmed, passed from view in the direction of the Ver.
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WHITE-TAILED WATTLED PHEASANT 147

human being. Now semi-erect, now quadrupedal, I made my way on and on,
passing for the first hundred yards through old, long-overgrown clearings. These
are characterized by trees of medium size and a very dense undergrowth of rotan-
like growths, spiny palms and, in every spot where sufficient sunlight filters through,
by the climbing fern which raises its fronds stratum upon stratum, resting upon the
stems and branches of any surrounding growths. Then a dimness settled down upon
the trail, I was able to walk in a less crouched attitude, and soon the undergrowth
thinned and I found myself surrounded by great mossy, lichened trunks—the remains
of the primeval forest of this great island.

I turned off the trail between the low, irregular walls of a gully, down which a
small brook filtered noiselessly through and over great pads of moss. Some distance
up-stream I found recent evidences of scratching among water-worn pebbles and
coarse damp gravel. This seemed encouraging, and as the sun was getting low I
looked about for a place to conceal myself. The gully narrowed and deepened into
a gorge, and presently showed where a side stream, a still finer unit in the plexus
of tributaries, entered. Ten or fifteen feet from the ground I found a little half cave,
in front of which leaned an old tree. The stones were quite dry, and here I ensconced
myself, crushing a score of pink begonia blossoms as I wriggled in. Somewhere
above me a dyal-bird was singing its sweet, long-drawn-out notes, and the steady
throb of great crickets shrilled close at hand among the crevices of rocks. After a
few minutes of waiting, the loud woof! woof! of hornbills’ wings came up the little
gorge, and several birds alighted out of sight and barked at each other for a while.
Then I heard the low crackling of leaves and twigs, and to my amazement I saw
cattle passing almost under me, splashing through the shallows. There were three,
one a bull almost black with all four feet white, the other two being cows. I knew
them for banteng, the wild Bornean cattle, and watched them pass slowly from sight
with the keenest interest.

For a long time I thought that I had seen my last game. Only pittas hopped
here and there, and an occasional barbet or broad-bill swooped across the space in
front of me. Once a flying lizard, with wings brilliant as those of a butterfly, glided
from a tree high overhead to a trunk only a few feet away, where it clung and
watched me. Its tail and hinder body were pressed closely to the bark, but the
forelegs held the shoulders and head at full length, while from the throat the long,
conical gular pouch inflated and deflated nervously. A family of paradise flycatchers
flew over, greatly excited, crests erected and scolding hoarsely. Then came the sight
of the afternoon. My eyes were resting idly on the big elephant-ear leaves at the
edge of the forest-trees when there stepped out the daintiest of tiny deer, a chevrotain,
then another and another. After a momentary sniffing at the spoor which the greater
hoofed animals had left, they stepped ahead and, close behind them, showing no fear,
came four White-tailed Pheasants. There was one fully adult male with sweeping
white tail, and a female, apparently its mate, while the other two were immature
birds of the year. I was already flattened to the rock as closely as possible, but
now I even half closed my eyes, dreading lest their sharp vision should search me
out. But it was another proof of how little these lesser jungle folk, while in the
forest, look for danger from above. Indeed, the slender stems of the great arum-like

148 A MONOGRAPH OF THE PHEASANTS

plants and elephant-ears could hardly support a dangerous foe, while the absence of
low branches left merely the smooth or mossy trunks which gave foothold only to
squirrels and woodpeckers. It was upon the surrounding jungle that they concen-
trated their sight and heating, and, as in my posts of observation in the densely
foliaged deodars and firs in the Himalayas, here, too, I was wholly safe from detection.
When the deer and the pheasants reached the stream they all drank deeply, selecting
clear pools well away from the disturbing footprints of the banteng. Never did all
seven drink at once; always one or two—a chevrotain with nostrils outstretched
and quivering, or a pheasant with head raised on high, each with a pair of bright,
unwinking eyes. For twenty seconds at a time the male White-tail would hold such
a position unmoving—a position of which it has been stated that “The picture in
the ‘ Birds of Asia’ gives a wrong idea of the carriage of the bird, and I very much
doubt whether it ever sits up in the way there depicted,” as if any bird whose habit
it was to skulk could not as well raise itself on tiptoe to gain a wider view-point.
During the many minutes which I was fortunate enough to keep these birds in
sight, I was impressed most profoundly with the constant, never-failing alertness
and sense of danger. I realized as never before how, in this jungle, which in com-
parison with South American forests seemed almost barren of organisms, life could
be lived safely by these weaker creatures only by an unceasing watchfulness. Even
when the birds began scratching, they were always on the lookout, and only when
for a fraction of a second they snatched a morsel did they cease their watch.

I was interested to see them scratch not only in the gravel and mud, apparently
uncovering tiny worms, but also in the shallows, making the water fly in all directions,
and then snatching eagerly at something edible which swam in the water. Some-
times the old female would take her stand in an inch or two of water, and her two
nearly-grown young would reap the benefit of her exertions, all three pursuing the
minute worms and crayfish and aquatic insects; for I later found that all three groups
contributed to the diet of these pheasants. The musk-deer seemed to enjoy standing
with all four feet in the water, and they busied themselves not so much with brows-
ing, as with their toilet. With their tiny hind hoofs they scratched and rubbed all
parts of their head and shoulders. Perhaps leeches were tormenting them or flies
had bitten them, as they did ourselves morning and evening upon the river banks.
There was a very obvious association and an apparent mutual understanding between
mammal and bird which brought vividly to mind the similar relation between the
musk-deer and kaleege pheasants of the Himalayas. The pheasants walked in and
out among the deer unheeded, and the deer, hardly a yard distant, sniffed unconcerned
at the muddy water sullied by the vigorous scratching of the birds. The bonds
between the various members of the White-tail family were very loose, indifference
expressing the attitude, especially on the part of the parents, although all kept more
or less closely together. But there was no attempt at feeding, or of any motion that
I could detect, except that of tolerance as of one bird to another in a flock. The
male invariably took the lead, and usually passed ahead of the others from one pool
or spot of gravel to another. The deer, although they appeared first in the open
part of the gorge, seemed rather to follow than to lead, and more than once passed
on, following and keeping close to the vicinity of the pheasants. It was remarkable

WHITE-TAILED WATTLED PHEASANT 149

how similar had been the first glimpse I had had of the Bornean crested firebacks—
feeding along a gravelly rivulet in a family group. Apart from the wonderful
opportunity thus offered, I was delighted with the picturesqueness; the deep, dark
gorge, with the green tangle up-stream opening into the clear space between the
rocky walls. Begonias dotted the rocks, a tiny pale blue blossom covered much of
the banks, and through this wild glade to see the tiny, graceful deer picking their
way, and the trim, wary birds in absolute freedom, wholly unconscious of being
watched, was a sight never to be forgotten.

It was all the more a remarkable sight (although later I was to have even more
intimate views of this pheasant) when we read of the experiences of other observers
in these jungles. Finsch says, “Bei der Undurchdringlichkeit des Dschungel ist
es z. B. unmoglich die hier lebenden prachtvollen Wildhuhner (d7gus, Lobtophasis
und Lofhura) zu schiessen, deren Stimme man wol hort, die man aber fast nie
zu Gesicht bekommt.”

So absorbed was I in not losing a movement, that I did not notice that the day
was passing, and suddenly a terrific clap of thunder crashed overhead. The storm
was going around, however, the black masses of cloud just visible through the trees
piling one over the other. The dimness of twilight fell, but soon the wonderful
Bornean afterglow changed the clouds to greyish pink and filled the whole glade
with the rich rosy hue. I knew how fleeting was this last light of day, and as the
pheasants had vanished, I dropped from my cave and made my way back to camp
as quickly as possible, reaching my hammock just as darkness fell. I watched
carefully, but caught not a glimpse either of deer or pheasants.

GENERAL DISTRIBUTION

The oft-quoted geographically and grammatically ambiguous statement that the
White-tailed Pheasant is “only found on the mountains” is untrue. There is no
doubt but that it does range upward on the lower slopes, and has been recorded from
Kalulong, Mulu, Dulit and other mountains, although I know of no record of over
twenty-two hundred feet elevation. On the other hand, it is not uncommon in low,
rolling forest country in Sarawak, many miles from any mountain or range of mountains.
Indeed, the natives of the lower zones are much more familiar with the B/azou, as they
call it, than many of the tribes from nearer the central ranges. It is extremely local
in distribution, a fact for which I could discover no explanation, and in passing down
a stream, bordered with homogeneous forest, the bird would be well known to the
natives of one section and wholly unknown to the next few tribes, although all were
skilful trappers and acquainted with all the edible pheasants and similar birds in their
neighbourhood. I could find no reason for the assertion that these birds dwell always
“far from water.” On the contrary, during the dry season I found them invariably
frequenting the water-courses, usually the larger streams and rivers, while at the
beginning of the rainy season, they, together with most other jungle animal life, came
less often to the river banks, finding, I suppose, sufficient moisture for their needs in
temporary jungle pools.

In British North Borneo White-tailed Pheasants are not uncommon along the

150 A MONOGRAPH OF THE PHEASANTS

upper Kimabatangan River. In Sarawak, they have been found in the interior from
Brunei on the north, to the mid Rejang region in the south, while in Dutch Borneo
we have several records (Biittikofer), and the upper Mahakam (Nieuwenhuis). So
while these birds do not apparently occur on the coast, they seem to be very generally—
if locally—distributed over much of the island as a whole, more especially in the
northern and central areas.

GENERAL ACCOUNT

There is no doubt that the White-tailed Pheasant is almost as particular as the
argus as to the character of its jungle home. Almost all the rivers and larger streams
of this region show the former occupancy of natives by the wide reaches of second
growth on each bank. Even when this has grown up to some semblance of the original
primeval growth, it is seldom that these pheasants are content to make their home
here. They prefer the deeper, uncut dry jungle, but morning and evening they have
no objection to crossing overgrown, or even cleared areas in order to reach the water.
As regular as do the big game, so these birds come twice each day to drink. The
argus and the White-tail will, however, sometimes return and inhabit such overgrown
country if it be wholly deserted by natives, while the crested fireback seems actually
to prefer such territory rather than to avoid it. Both the White-tail and the argus
require rolling country, and I have never known of one of the former being found in
the low, level, half-marshy regions which are the delight of the firebacks.

The first news of this little-known pheasant which I received on reaching Sarawak
was that there was a live male in the possession of an American missionary at Sibu
on the Rejang River. Although the few specimens in museums have been taken
mostly in the extreme north of the colony—on the Baram and Lawas Rivers, I decided
on following up the rumour of this live bird. When I reached Sibu by one of the
little fortnightly steamers; I found that the bird had recently died, and after an attempt
had been made to skin it, had been thrown away. The memory of the man who had
buried it was poor, and after a thorough digging up of numerous plots of ground by
sundry Dyak and Chinese convicts, we gave up the search for the skeleton.

The bird had been caught by the Dyaks two days’ journey by canoe up the river,
so I went on to the last outstation, Fort Kapit, where I found myself just within the
zone of White-tailed Pheasants. From here I pushed on in a seventy-foot canoe, and
later in smaller craft up the Rejang, Balleh and Mujong Rivers, and found this Species
local but not rare, and distributed over much of central Sarawak, as frequently in the
low-lying but rolling hilly country as near the foot-hills of the mountains. Its supposed
rarity seems due to several causes: its extremely local distribution, the fear of the
head-hunting Dyaks, and especially the unconquerable tendency of these natives to
instantly kill and eat every animal, bird, snake or other creature which shows enough
flesh to make it worth while.

One can draw a close analogy between the White-tailed Pheasant and the Dyaks
themselves. Both are governed by the presence of jungle. When it is cut down and
the land impoverished by a crop, the Dyaks move on up-river to more primitive jungles.
The White-tails prefer the open undergrowth of the older forests to the dense thorns

PHOTOGRAVURE 28

BORNEAN HOME OF THE-WHITE-TAILED WATTLED PHEASANT

On the slopes of the rolling hills of central Sarawak, in the warm jungles, sloping down to the
yellow rivers, these magnificent pheasants have lived for untold centuries. Only by days of the most
patient watching can they be seen, and more often a fleeting glimpse is all that rewards a pains-
taking stalk.

Beneath the tangles of soft-fronded climbing ferns, or the solid phalanx of bamboos, the birds may
remain hidden a few yards distant, and never be discovered. The young birds are less wary, and, trusting

more to their inconspicuous colouring, often come into more open spaces, to scratch for food or to preen

their plumage.

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WHITE-TAILED WATTLED PHEASANT I5I

of second growth, and so keep just ahead or inland of the devastating path of the
Dyaks. Both seem to cling closely to the rivers and creeks. Away from these aquatic
highways one may travel for many days and meet neither pheasants nor men.

My first meeting with the living White-tailed Pheasant gave me an idea of the
keenness of observation of some of the Dyaks. On July 12, 1910, I sent my Eurasian
taxidermist and collector on up-river to interest a distant village of Sea-Dyaks in
trapping Llazou for me. I had collected several argus and fireback pheasants, together
with a zebra civet and a “moon rat,” and was engrossed in preparing these specimens,
when two little Dyak boys came to the camp carrying a wicker cage. I gave but a
glance at the birds through the small interstices, and seeing what I took to be a pair
of immature crested firebacks, I refused to purchase them, especially as they wanted
an outrageous price. They carried the birds away, but while I was at tiffin they
returned with a Dyak from the nearest village, who said the birds were “ B/azou wives.”
I laughed at this, but looked again at them, and a glance at the ventral surface showed
that the savage was right and that they were undoubtedly immature male or female
White-tailed Wattled Pheasants. The three pairs of wattles were plainly distinguish-
able, and the paleness of the head left no doubt. One of the birds, a young female,
soon died, but the other lived, was perfectly tame, and for weeks thrived on its paddy.
Indeed, I was able to ship it from Singapore in good condition, and it lived on board
ship until within sight of New York City.

From this time on we had no trouble in collecting adult and first-year birds, but
in several weeks’ search observed no individual in the chestnut-tailed plumage of the
second year. In July the pheasants were all moulting heavily, and they showed no
signs of a recent breeding season. Indeed, the age of the immature birds indicated that
the nesting season, at least for these individuals, is about April in this part of Sarawak.

The White-tailed and argus pheasants living, as I have said, usually in the
vicinity of the larger rivers and streams, may be found in the first, or at most the second
line of hills stretching back from the banks of the upper reaches of the rivers. Their
regular habit of leaving these dry, hilly jungles and making their way to the water
to drink morning and evening, unfortunately facilitates the trapping of these splendid
birds. The Dyaks cut a line of dense underbrush and pile it along a ridge of hills
parallel with the river bank, packing it closely, fence-like, uphill and down, winding
along for a half or even a whole mile. Every ten feet or thereabouts a converging
semi-circle of bamboo-sticks leads to an opening in this brush fence, and in this is
spread a noose attached to a bent, living sapling. Each Dyak in some of the villages
has from twenty to eighty of these traps, and goes his round every morning. If
the birds are not promptly found and removed, they are invariably devoured by civet
cats, although these creatures themselves are often caught, together with binturongs
and porcupines.

Practically nothing is known about the nesting habits of this pheasant. Three
Dyaks of as many different tribes assured me that it laid only two eggs, and on two
separate occasions 1 saw what was very apparently a family of four—parents and two
young, so that some measure of credence may be given to this number. The only egg
of which I have knowledge is the one said to be of this species collected near Mount
Dulit on the Baram River, Sarawak, in September, by A. H. Everett, and now in the

152 A MONOGRAPH OF THE PHEASANTS

British Museum. It is a regular broad oval in shape and of a pale pinkish cream
colour, being thus hardly distinguishable from the egg of the Bornean crested fireback.
It measures 51 X 40 mm.

It is but a truism to state that these pheasants are skulkers and runners rather
than fliers. What ground bird whose home is amongst jungle undergrowth could be
aught else? When approaching these birds with the greatest caution I have very often
been able to catch a momentary glimpse of them, the white tail of.the male advertising
him at once. When several of both sexes were together it was the males which first
fled, the brown immature males and females delaying to have a second look at me,
and then usually disappearing in a different direction. If when all had vanished I
squatted and remained motionless, in ten or fifteen minutes I would hear what I judge
is the covey call, a penetrating, rather metallic; Look/ kook ! uttered by both sexes.
This would invariably become subdued by distance, the birds evidently converging at
some far-off spot. But if the first place of meeting with the birds was a favourite
feeding-ground, as evidenced by the abundant signs of scratching, one might be
reasonably certain of seeing the birds at the same place on another day. Only in the
morning or toward evening, however. Search as I might, I could never discover a
single individual at other times of the day.

The natives, both Kayans and Dyaks, told me that the birds had regular roosting-
places, and that they set their snares about these trees. One young man—a “grey
Dyak” as I called him, as he was one of those whose skin from head to foot was like
grey powder, a common uncontagious disease among these people—undertook to lead
me to one of these roosting-places. After a toilsome tramp of several miles through
the low jungle we came to a series of steep hills, and near the summit of one of these
elevations my guide suddenly stooped and picked up a tuft of feathers and part of
the foot and leg of a White-tailed Pheasant. The bird had been caught in a snare
which now dangled overhead from the tip of a sapling, and had been devoured by a
zebra civet, so the Dyak said, indicating the species of the marauder by pointing to
the bit of skin dangling at the back of his loin-cloth. A few paces farther he pointed
to the ground, where there was abundant sign, showing that some good-sized bird or
birds had been accustomed to spend the night in the branches overhead. The tree was
slender and very smooth barked, and it stood quite isolated, its branches free from
contact with those of the surrounding growths—a well-chosen roost for protection
against night attacks. I asked the Dyak later how he discovered and recognized such
roosts, and his answer was that the finding was by accident, but that such sign on
a steep hillside could be made only by Blazou or Ruoi—the White-tailed or the argus,
but that the latter never roosted in small trees, only in large ones.

The spurs of the male White-tailed Pheasants are moderately developed, so that
a certain amount of pugnacity is implied. Hewett states that they “are very
pugnacious, and that their heads are consequently often raw and scarred, while they
would attack any other birds put in their cages.” I saw no evidence of the scarring
on the face and wattles of the birds which came under my observation, and those
which I had in captivity were gentle, although it is true that they had no opportunity
to attack birds other than their own species. I suppose that as it was not the breeding
season they were quieter than they might have been six months before.

WHITE-TAILED WATTLED PHEASANT 153

It is probable that the breeding season of these pheasants is rather elastic, and,
like most tropical birds, may in different localities extend over a good portion of the
year. Certain it is that in July in Sarawak I obtained birds of five or six months
of age, and during the same week saw a pair of adult males sparring with one another
and assiduously courting a female. This was the most important observation which
it was my good fortune to make concerning these pheasants in Borneo.

Instead of sunshine, or at least a thin haze of clouds during the day, balanced by
a terrific downpour of driving rain throughout the night, the reverse one day took
place, and after I had made an early start and was several long miles from camp, a
sudden deluge descended upon me, and in a few minutes I was thoroughly drenched.
I was encumbered with nothing but my stereo glasses, as I had planned a day of pure
observation. Plodding on through the dripping undergrowth I came to where a great
tree had fallen-—a tree whose life must have stretched back well beyond the time when
white men first set foot in Borneo. It had been strongly buttressed in all directions,
but a stroke of lightning had run down the trunk and splintered one of the great basal
supporting walls of living wood, and thus unsupported, the mighty mass had given
way. It had cut a wide swathe for itself, carrying down all the lesser growth in its
path. Under the shelter of two of its remaining upreared buttresses I found ample
protection, and here I remained for many hours watching the jungle life of a rainy day.
None but small folk came near, however, until well after midday, when the sun at last
broke through, although the saturated forest continued to rain down drops for many
a minute after the whole sky above had cleared. ‘The first intimation I had that
pheasants were about was a sharp, nervous kak! kak! which came from the other
end of the mass of shattered foliage. By slow and painful manceuvring I was able
to peer out unseen and to detect a male White-tail stalking slowly along at full height,
concentrating his attention upon a clump of maroon-leaved plants near by. Whatever
it was which had excited his interest or suspicion he was soon satisfied that all was
well, and he began feeding unconcernedly, scratching in the mould, or now and then
picking some insect from a leaf. For seven or eight minutes I watched him, and at
last dared to shift my glasses and rest them upon a ledge of bark, so that I was able
to see even the contraction and dilation of his pupil as he remained motionless for a
moment. The splash of falling drops, the twitter of small flycatchers, even the sudden,
wholly unexpected and startling notes of a barbet did not distract his attention a
moment. But always he was alert, always a momentary snatch at some morsel of
food was followed by an instant of concentrated listening, a quick, comprehensive glance
which took in all the adjacent shrubs which might shelter a foe.

A change at last came over him; without seeming alarmed, his attention became
focussed more and more in a particular direction. Straining all my senses, I could
neither hear nor see anything—but he knew of something which was as a fourth
dimension to my dulled faculties of perception. Finally the bird ceased all attempts |
at feeding and stood immovable, never taking his eye from the edge of the jungle.
At that exact spot, after several minutes of further waiting, a pair of adult White-tails
emerged rather precipitately, the female first, as if the male had been pursuing her. On
the instant both caught sight of the waiting bird, and both uttered a low, startled faz !

In a moment, however, the alarm passed and the female came diagonally across the
VOL, II x

154 A MONOGRAPH OF THE PHEASANTS

clear space, taking ants or other insects from leaf or ground as she came. The male
birds stood for some time apparently watching one another, then my pheasant began
approaching the female with quick, short steps. A moment later and the second male -
drew slowly nearer. I was breathless with excitement. My mind ran over the few
meagre padded lines which represented our knowledge of these wary birds, and here
the good fortune of the unusual diurnal rainfall had given me this wonderful oppor-
tunity. The second male approached steadily and slowly, with no noticeable change
in his demeanour, but my bird became more and more excited as he neared the female.
His wings were slightly lifted from the sides of the body, although the fluffing out of
the plumage left no space beneath them. The head was well up and drawn back, the
wattles seemed very much elongated, but a careful look through the stereos showed
the wrinkles still evident, the apparent enlargement being due to the straightening of
the occipital and gular pairs, so that they no longer hung flaccid and limp. Now the
bird came close to the little brown hen, who edged away but without showing alarm,
and the moment for which I had been waiting came at last. Two complete turns my
male made around the hen, circling her irregularly but completely, his attitude being
a stiff strutting with his magnificent tail spread to the full. It was a lateral display,
and yet I could detect no depressing of the plumage on the side toward the object of
his emotion nor yet an elevating of the feathers on the farther side. The bird simply
progressed evenly, symmetrically, head pointing forward, but the tail-feathers forming
almost a semi-circle of immaculate brightness. High over the back curved the inner,
wide-vaned plumes, almost touching the upraised neck, while the remaining pairs
arched around down to the very feet, in fact, again and again they were pushed forward,
actually concealing the scarlet legs. The gait was peculiar, a few quick steps, then
immobility and a statuesque pose, again the few steps and again the hesitancy.

At each rest the bird seemed to become actually larger, the feathers of the body
fluffing out so evenly and slowly that one could not avoid the idea of solidity.
Simultaneously, the skin of the face seemed to puff out, although this appearance
may have been caused by a change in colour intensity. But there was no question
as to the muscular extension of the wattles; they visibly lengthened, rather spas-
modically, reaching their greatest extent at the moment when the bird ceased its walk.
I would place this extreme distention at about half the length again of the retracted
organ. This fact observed, I centred my attention upon the tail, and when the bird
slowly turned away from me I saw that the outer feathers were spread well apart
as well as downward, so that from the rear view it appeared as if the bird were
walking between the opposite sides of its tail, The normal carriage is with the
opposite sides of the tail closely appressed at a very acute angle; at this time the
posterior view showed them spread widely at the base—tent-like. As the feathers
pushed forward or were relaxed an audible rustling was produced, and when the
bird passed over dried leaves this sound increased; at times there were a dozen or
more leaves literally raked up by the stiff, recurved outer quills of the tail-feathers.
The supreme moment came when the hen stopped for a moment and her suitor
halted squarely in front facing her. With what wattle or tail accompaniment this
new, supreme phase would have been enacted I shall never know, for male number
two now approached, and both at once rushed at each other with half-spread wings,

WHITE-TAILED WATTLED PHEASANT 155

and for a moment pecked viciously, but showed no inclination to use their spurs.
Then the zeztgezst of the jungle, in its mysterious way sent forth to these wild
creatures the knowledge of danger, and with a single impulse all three fled, my
friend in one direction and the others near where they had emerged. There was
no apparent choice of companion or direction—the birds simply ran swiftly for the
nearest shelter, heads outstretched, tails low and close shut, the male as different
from his courtship attitude of a minute before as can be imagined.

Thus passed my only opportunity for observation of the intimate life of these
splendid birds. This was sufficient, however, to confirm my suspicion that the worn
and abraded condition of the tail-feathers was not due wholly to congenital defects.
Judging by the recklessness of passage over sodden leaves, twigs and irregularities
in the debris of the jungle floor, which I observed during the courtship, it is only
remarkable that anything remains intact but the abraded stumps of the outer tail-
feathers. I have discussed the interesting correlation between the inherited defective
condition of the rectrices and this ontogenetic abuse of these long-suffering feathers
elsewhere.

What I saw of the courtship of the White-tailed Pheasant leads me to think
that the general method at the start is a lateral, asymmetrical display, but the
finale is as certainly frontal. Whether the tail at such a moment is spread outward,
so as to form a white background, must be left to conjecture. If so, the courtship
as a whole is quite similar to that of the peacock pheasants. The short and inde-
cisive combat which I witnessed was suggestive only in the fact that no attempt
was made to use spurs, and this, taken in connection with the extreme variation and
at most only moderate development of these organs, would indicate that the White-
tails are much less given to fighting than the terribly armed fireback pheasants.

As to the food of the White-tailed Pheasant, from examination of the crops of
recently killed birds, I found that they were equally insectivorous and frugivorous.
Ants formed by far the dominant item in their diet, a small black species being the
most abundant, while small crickets and other orthoptera were also not uncommon.
Termites were second in point of numbers, and twice I actually saw the birds feed-
ing about the earthen tunnels of these so-called white ants. Minute round black
seeds, and large pieces of some fruit of a nut-like consistence were also a favourite
article of diet. The Dyaks called this cacos, and said it was not edible for man.
These natives were also unanimous in saying that when a certain fluviatile fruit
became ripe, these pheasants and many of the smaller mammals came to the river
banks in much larger numbers than usual for the purpose of feeding on this dainty.
I could discover neither the fruit nor the trees which bore it.

Until we know far more than we do at present concerning the causes of the
colours of organisms, we can do little else than narrow down the various questions
and theories by the process of elimination. In the White-tailed Pheasant, for example,
we have a bird which spends its life in the dense jungle, among thick underbrush,
where the light is constantly dim and shaded. No one will dispute that the sombre
colours of the female are protective among such surroundings, and even the more
conspicuous second-year male, with his blue-edged body pattern and chestnut-tail,
would merge with almost any situation in these tropical forests. But for the fully

156 A MONOGRAPH OF THE PHEASANTS

adult bird we can find no such background. Shining white spots of such size are
not scattered about upon the jungle debris, and whether stalking beside a brook in
an open glade, or half-hidden by climbing ferns, the White-tailed Pheasant in the
glory of his full beauty is striking and conspicuous anywhere. Twice I had oppor-
tuuity to see both males and females alarmed, and both times it was the white-
tailed adult which fled first; swiftly, without a backward glance or hesitation; the
duller individuals waiting a moment to see if the danger was imminent. Once a
solitary female squatted for an instant, before my hand touched a twig and sent her
too flying headlong, threading the fern-stems as if they were no obstruction. This
is the best proof we have, and I consider that many of our ultimate conclusions
must rest upon just such evidence. The realization of danger is most acute in the
bird whose plumage is inevitably conspicuous; the sense of trust to assimilation
with the surroundings—wholly instinctive though it be—indicates with absolute
certainty that many, many times such behaviour had enabled the little brown hen
to escape observation. The red eye, feet and legs and purplish wattles detract, if
anything, from the general conspicuousness of the pheasant. There are myriads of
red or pink or purple jungle shades—dead or insect-eaten leaves, lichens, fungus,
moss, bark, begonia stems, and scores of other unnamed and unnameable. But as
for the flaunting white tail, that splendid badge of reward for three years’ successful
striving against the thousand and one forest dangers, it mimics nothing, it flares
out like a heliograph; the brave owner finds his food, his mate, and lives his life
in spite of it, with only his keen eyes and ears and his swift scarlet feet as counter-
balancing assets.

And how little all these count for when pitted against the native savages. The
rainy season has come to an end—the jungle pools are dried up, and every day the
White-tail must make longer and longer pilgrimages to the low-lying marshy country.
One day he turns his steps toward the river itself, walking steadily over hill after
hill, at last turning into a narrow valley. Halfway down a pile of brush obstructs
his path, and strive though he will, he can find no way past it. A few yards
farther and an opening appears, just wide enough for him to step through. His
scarlet toes press upon the loose bamboo sticks lying at the mouth, there is a sudden
snap, the whole world revolves and high in the air the splendid bird dangles, swaying
back and forth from the bent sapling. Perhaps death comes swiftly from a passing
civet cat, perhaps the following morning the Dyak trapper goes his rounds. But
no matter whether soon or late, the days, months, years of watchfulness, from the
chick which crouchéd and watched the civet slink past, to the white-tailed bird
which yesterday out-saw and out-ran the swift-footed moon rat itself—all have been
in vain. The Dyak squaw strips the shining plumage from the body, or sells it for
a few beads to a passing Chinaman; the White-tailed Pheasant has run its race.

CAPTIVITY

The first knowledge we have of these birds in captivity is of the pair which
were sent to Sourabaya, Java. The male died in that island, but the female reached
the Amsterdam Zoological Gardens, being so weakened by the journey, however, that

WHITE-TAILED WATTLED PHEASANT 157

it lived only a few weeks. Treacher says that he found difficulty in keeping these
pheasants alive, but Hewett’s experience was otherwise, as he tells us that they
thrive in captivity on paddy, boiled rice and fruit. He goes on to say, however, that
he could never keep any of the hens alive, as they refused food and seemed to
mope, and would die in a day or two. The hens I had in captivity were, on the
contrary, more gentle and ready to feed than the cock-birds, and it was a hen which
lived on board ship from Singapore until actually within sight of the harbour of
New York. Such isolated and conflicting experiences are of little value in forming
any opinion, and doubtless represent merely individual physique or temper on the
part of the birds. It is my opinion that when a systematic attempt is made to
procure a number of these birds alive, they will be found to be no more delicate
than argus pheasants.

DETAILED DESCRIPTION

ApuLtt MALE.—Feathered parts of head and upper neck dull black, with slight
steel-blue terminal margins. Chin and throat thinly feathered, plain black. Neck
all around and breast shining crimson, most of the feathers with a narrow steel-blue
margin. A typical feather from these areas is divided into three equal parts, the
basal third of fluffy grey down; a middle third of normal black vane, and a distal
third of shining crimson. At the beginning of this latter area the barbules become
very abruptly shortened, lose all trace of barbicels, and throughout its length are so
imperfect and minute, that to the eye the shafts of the barbs appear bare. At the
very tip the barbules are slightly longer, forming the exceedingly narrow terminal
steel-blue band.

Posterior to the neck and breast (and also somewhat with advancing age) the
crimson disappears apparently rather abruptly, the dead black mid-area pushing up
and taking its place, while the steel-blue band widens—this being the general
character of all the body plumage, except that the blue terminal band is absent from
the primaries, and outer secondaries together with their coverts, the abdomen, thighs
and the extremely short under tail-coverts. The steel band is much narrower on the
ventral than on the dorsal plumage.

The primaries are dark brown, not dead black, and in a considerable percentage,
more than forty per cent., of wild shot birds, they show as an interesting variation
the presence of more or less white. This may be present on as many as nine of
the primaries, occurring chiefly on the basal third of the feathers. This colour is
usually quite pure on the outer two or three feathers, but becomes mottled and
clouded with dark brown as we pass inward. It is rather asymmetrical in occurrence
in the total: amount on each feather of a corresponding pair in the two wings.
None of the adjoining coverts, either upper or lower, show any traces, and indeed
the majority, some sixty per cent., of the adult males which I have examined show
no trace whatever of this alar-whitening. Nevertheless, its occurrence again and
again in birds collected in different parts of Borneo stamps the character as one, while
wholly individual, yet in no sense pathologically albinistic; very probably correlated
in some way with the white colour of the tail-feathers.

158 A MONOGRAPH OF THE PHEASANTS

I have said that the disappearance of the crimson is apparently abrupt, because
this is true only from a superficial point of view. The crimson persists on almost
all the body plumage, much darkened, however, and purpled by the encroaching black.
A feather from the mid-back shows a shining blue terminal band of 5 mm. in
width. This exhibits the peculiar wrinkle or upward bending of the vane which
I described in Lofhura. Bordering on this blue band is a well-marked zone of dark
_purplish-crimson, which merges into the proximal dull black.

The change in colour of the crimson zone is most interestingly correlated with
a corresponding alteration in structure. The barbules are almost as short as in the
pure crimson nuchal region, but with the blackening of the bright colour barbicels
appear, and throughout the entire area-from four to six of these tiny hooklets are present
on every barbule. Thus the barbs of this zone tend to cohere to one another, while
the shortness of the barbules results in a much closer approximation of these barbs than
usual. This results in a springing apart of the web in a number of places, and produces
the curious split-up appearance of the feathers of the White-tailed Pheasant, a character
I have observed in no other bird. The terminal part of each feather is usually split
apart into eight or ten divisions, each consisting of a greater or less number of barbs
very closely cohering to one another.

The feathers of the rump are short and truncate, very sharply set off from the
upper tail-coverts. These latter are long, somewhat curved and pure white, so that
in both form and colour they resemble the tail-feathers themselves.

As the tail of this unusually interesting pheasant is its chief glory and the most
specialized in colour and number of feathers of any tail in the family of pheasants, it
is well worth considering in some detail. In the fully adult male the entire tail is pure
white, and at this age the usual, normal number of feathers—certainly the maximum
number—is thirty-two, just 7wece as many as in the somewhat allied genus Lophura.
It is not often that the complete number is present, one or more usually having been
pulled out in some way. After very careful examination of the roots of the tail-feathers
and the surrounding skin by the complete relaxation of many adult male birds, I am
_convinced that it is not uncommon for thirty to be the total number. It is a very easy
matter to detect the location of missing feathers, and in a number of cases where thirty
feathers were present, or that number as represented by the total number of follicles,
I have satisfied myself that there was absolutely not a single feather missing, nor any
additional unoccupied follicles. In such a specialized organ as the tail of this pheasant
such variation is surprising, and is of especial interest in view of the variation in
number of rectrices in the species of Cvossoptilon. Thirty-two, however, may be
taken as the maximum, usual number of tail-feathers in the adult male White-tailed
Pheasant.

As we shall see, the number of rectrices in the chestnut tail of the second-year
male is twenty-four, so that-in the succeeding or third annual moult there is an increase
of eight feathers. When the tail of a freshly killed pheasant is spread widely the
feathers seem to be in quite regular alignment, but as the tail is allowed slowly to
close, an irregularity becomes apparent, and we find that this occurs solely among the
central eight or ten pairs. Comparison of fully adult birds with those in the chestnut-
tail plumage, and of individuals moulting from one to the other show clearly that the

WHITE-TAILED WATTLED PHEASANT 159

additional tail-feathers are all more or less central and make their appearance anterior
to the regular line of rectrices, sometimes forcing these out of alignment. In other
words, the additional feathers are derived from the distal row of upper tail-coverts,
lengthened and coloured to similate rectrices in the third annual moult. Indeed, as
we have seen, even the remaining lesser and median coverts are all affected by the
same tendency, and completely isolated by form and colour from the feathers of the
rump. I shall speak further of this transformation when describing the second-year
plumage.

Specialization is not confined to the number of rectrices, but has very noticeably
affected their structure. The vanes of all the feathers, while to a certain extent normally
cohesive, yet show a downy character and softness of texture wholly incompatible with
any great usefulness in flight. One can see at a glance that the tail is avowedly an
ornament, with but slight power to perform its more normal function of a rudder.
Beginning with the central pairs the feathers show splendid wide, symmetrical webs,
and are similar in character for the inner nine or ten pairs, gradually, however, becoming
less curved. The outer six, or almost invariably seven pairs become, quite abruptly,
very different in character. They show almost no curvature, the outer five pairs being
quite straight, the outer webs becoming exceedingly short, and although the feathers
rapidly decrease in length toward the outer pairs, yet the shafts of all are much enlarged,
stiff and spine-like. The most interesting character is the degeneration of the webbing
on these outer seven pairs. This is faintly indicated on some of the more central
feathers, whose tips may be rather worn and frayed for 20 mm. or more, but on the
outer, stiff-shafted feathers it reaches an extreme. Long before I observed the courtship
(p. 154) I suspected that some such manceuvring must take place to account for the
worn and broken candition of the tips of the outer quills, but in the case of this
pheasant (as I have shown in the racket tails of the motmot, “ Zoologica,” New York
Zoological Society, No. 5, 1910, p. 141), environmental manipulation only goes hand
in hand with congenital structure. There is great variation in this congenital weakness
of the web or in its complete absence. In an unsheathed feather sprouting among these
outer rectrices, the shaft may be wholly bare for 20 or 30 mm., followed by an equal
area of strongly attached barbs. Or the web may be present for a third of the length
of the feather and complete down even to the barbicels, and yet its point of attachment
near the shaft so weak that the whole vane tears away with the removal of the enfolding
sheath.

In the full-grown outer tail-feathers of the pheasant, examined in the living bird
or in a recently shot or trapped specimen, we find the inner web fully developed, while
the outer is very greatly reduced, up to the very root of the rachis. This reduction
occurs in two decided nodes or steps. Taking the average of a score of adult males,
it is first especially noticeable on the fifth from the outer pair where (all measurements
are 125 mm. from the tips of the feathers) the outer web is 12 mm. in width as compared
with 25 mm. on the sixth pair of feathers. On the fourth pair, the outer web narrows
suddenly to only 4 mm., and from this the narrowing is very gradual down to 2°5 mm.
on the outer pair of rectrices.

On the terminal portion of the outer feather a most dishevelled and imperfect
condition of the vane is found. Usually, for the length of 80 to 100 mm. from the tip,

160 A MONOGRAPH OF THE PHEASANTS

the shaft is entirely bare, while beyond this point the web, both outer and inner, is
broken by bare patches and is generally mussed and wrinkled for often half the entire
length of the feather. The frequent bending or breaking of the barbs midway of their
length is explained by the occurrence of numerous fault-bars on these outer feathers.
The tips of the longest tail-feathers—the sixth and seventh from the outer pairs—are
frequently themselves bent or broken, and all the shaft tips are soiled and unsightly.
Not only are the shafts of the outer rectrices exceedingly thick and strong, but they
have actually downward curving tips—almost woodpecker-like—curving in the direction
from which the friction and attrition against the ground and leaves must come. From
the sixth pair inward this character is wholly lacking.

When the scattering of barbs left here and there in the heart of the denuded area
is examined, the congenital weakness is plainly evident. It is a constricting or rather
flattening of the extreme base of the barbs, together with the loss of all the barbules at
that point. In every way it is directly comparable with the corresponding phenomenon
in the freshly-grown central rectrices of the motmot.

There seems to be some grounds for supposing that in these pheasants the absence
of pigment in the tail-feathers is at least indirectly correlated with the downy character,
and perhaps a tendency to basal weakness of the barbs, for in many specimens of the
Malayan crested fireback the central rectrices show a very similar denudation of
the shaft, not extending beyond the white area which includes the entire webs of the
two central pairs and the inner web of the third.

Even in the chestnut rectrices of the second-year male White-tail, the phenomenon
is foreshadowed, the outer webs of the outer one or two pairs of rectrices sometimes
showing considerable denudation, while the tips of several of the feathers may be bent
or broken. .

In the tail of the adult male White-tail, the seven external pairs of rectrices seem
to be much more intimately connected with the complicated caudal musculature than
the succeeding inner ones, the relation being especially close with the depressing
muscles. So it seems certain that this structure is connected with the greater wear
and tear which the congenitally weakened webs of these feathers invariably show,
causing them to be more active in the downward and somewhat outward spreading
which is a feature apparently of both challenge and courtship.

Face and bare skin of the side crown bright purplish-blue, duller on the wrinkled
retracted wattles. Skin of chin, visible through the scanty feathering, back to the
base of the gape, pink. The two anterior pairs of wattles are pointed; those on each
side of the occiput are bifurcated, this terminal area being distinctly dark in colour and
of a wholly different structure from the rest of the wattle. The wattle as a whole
consists of transverse bands of smooth blue skin, which wrinkles closely together.
The bifurcated black tip is covered thickly with small and irregular papilla. The
irides are very deep, strong carmiine; legs and feet carmine, less intense than the
eyes.

Mandibles black except distal half of lower and the tip of the upper mandible,
which are pale horn colour. Weight of an adult male 23 to 34 lbs. Length, 790 mm. ;
bill from nostril, 21; wing, 260; tail, 460; tarsus, 92; middle toe and claw, 63. elite
spurs of this pheasant are comparatively short and weak, but exceedingly variable in

PHOTOGRAVURE 29

Wet e-TAILED WATTLED PHEASANT

In the immature White-tailed Pheasant (left) the wattles are short and inconspicuous. In the adult
(right) they are long and wrinkled, and at the moment of courtship capable of being considerably
elongated.

The head-hunting Sea Dyaks of Sarawak are splendid hunters, and will spend days and nights
stalking and trapping pheasants and other birds. With the crudest kind of traps they would capture the

birds alive and bring them to me unharmed in wicker quakes.

"gs AMUVAHDOTOHS

Sr. Rae ;

" esouaigenosnt

yldgrsbiano:

aidyin brg..eveb chaage.dlix

onfj. stuiqed blvow: godh eqeit ie °

LIVING AND

DEAD WH

ITE-TAILE!

DYAK. HUI

NTERS ANI

D PHEASANTS.

PHOTOGRAVURE PLATE 29

WHITE-TAILED WATTLED PHEASANT 161

size. In fully adult males they vary from 7 to 10 mm., while in several birds of the
same age I have seen them merely undeveloped flattened nodules.

There has been much misconception as to the length of the wattles, both in their
retracted and extended condition. I have examined living adult males carefully when
they were in a state of emotional rest, and then seen the wattles somewhat extended
when the birds became alarmed. Again I have seen the wattles still further lengthen
when two males were beginning to show off before a female, and finally, after manipula-
tion of the wattles in both living and freshly-killed birds, I realized that the amount
of possible lengthening and expansion has been greatly exaggerated. In the Museum
in Kuching, Sarawak, are two adult male White-tails said to be mounted by Hose.
The wattles of these birds have been stretched to their utmost while pliable and pinned
in this position, thus giving a most remarkable appearance to the head. After watching
the movement of the wattles in live birds and dissecting the musculature in dead
specimens, I am certain that any such extension as the taxidermist has produced in
these cases is wholly false. I give in tabular form the wattle measurements in
millimetres of these falsely-stretched ones ; of the natural normal length of the retracted
wattles, and of what I consider to be the greatest length possible when the bird is
showing off before the female.

Artificially Normatly Normally

stretched retracted extended
Loral : : 17 6 II
Occipital . : 90 13 24.
Gular ‘ : 107 32 56

Thus the bird is unquestionably able to increase the length of its wattles nearly
100 per cent. in the case of all three pairs.

ApULT FrmMaALeE.—Entire upper body plumage brownish-buff vermiculated with
black, except on the hind neck, where the feathers are almost wholly dull, brownish
buff. The wing-coverts are slightly more rufous. The black secondaries have very
coarse vermiculations in the form of numerous wavy, oblique lines of a conspicuous
rufous buff colour on the outer web and a very indistinct dim rufous on the inner.
The primaries are sometimes indistinctly mottled with rufous buff on the outer web,
or they may be plain rufous brown on webs. The upper coverts and tail are rich
chestnut, sometimes irregularly mottled with black, more rarely plain chestnut. The
tail is paler beneath, with very conspicuous brownish-white shafts, while the usual
number of rectrices is twenty-six. Chin and throat whitish, shading into yellowish
buff on the side and lower neck. Under-parts quite uniform, pale rufous buff, faintly
mottled with dark brown. The pale brown shafts are quite conspicuous. All three
pairs of wattles are clearly distinguishable in this sex; the loral as minute nodule8, the
two other pairs as short flaps about 3 mm. in length. The occipital pair show no
evidence of bifurcation. The spurs are sharp-pointed but minute. Facial skin clear
blue; iris, legs and feet bright red; mandibles black, the tips paler horn colour.
Weight, 24 lbs. Bill from nostril, 19 mm.; wing, 230; tail, 175; tarsus, 70; middle
toe and claw, 50.

CHANGE OF CoLouR IN ADuLT MusEum Sprcimens.—The adult White-tailed

Pheasants undergo considerable change in colour in the course of years even in dark
VOL. II Y

162 A MONOGRAPH OF THE PHEASANTS

skin cases, but more especially when exposed to the light for the same length of time.
A male bird which has been mounted in the Sarawak Museum for twenty years has
all the dark brown primaries and coverts and the black ventral plumage changed to
pale brown, the shafts keeping much lighter than the vanes. The upper plumage has
held true, but the black feathers of the head and neck have turned as light brown as
those of an average female.

The photochemical changes which have taken place are even more extreme in the
plumage of a mounted female. In a bird which has been exposed to only a moderate
amount of light for about eighteen years, the head and neck have become quite white,
basally somewhat mottled with brown, while all the upper plumage has become
extremely faded on all the uncovered portions of the feathers. The exposed parts of
the primaries are a dirty white, and the two or three pairs of central rectrices, which
are marred by numerous fault-bars, are irregularly whitened in the vicinity of these
imperfections. In both this mounted specimen and in skins which have been kept
in light-tight drawers, the blue facial skin of the females has given place to a fleshy
tint which would readily be mistaken for a faded red. The entire aspect of the birds
is changed, and when we remember the similar case of a radical alteration in the female
type of Gennaeus melanonotus (p. 39), we should realize the danger of naming species
from old skins, and should constantly keep in mind the danger of possible photophobic
plumage in using old type specimens for purposes of comparison.

NataL Down.—The chick in the down bears a striking resemblance to the young
of the Bornean crested fireback, except that the general tone is warm rufous instead
of darker. The upper parts are rich rufous, brighter on the head; the face and under
down whitish buff, with a black bar through the face and an indistinct wash of warm
buff over the breast.

JUVENILE PLumace.—Here, too, as the dominant character, we find the pair of
sub-terminal black ocelli which is so pronounced in the corresponding plumage of the
species of Lofhura. As usual, the down persists on the head and neck long after
the remainder of the body is clothed in the contour feathers of the juvenile plumage.
The ventral plumage of this very transient garb is quite downy and rather characterless,
being of a pale buff with more or less indications of transverse darker markings, and
the mantle and back, while of firmer vaned feathers, are greyish-buff mottled with dark
rather than distinctly patterned. The scapulars and middle wing-coverts, however,
show this mark at its fullest development, the feather being a rather even vermiculation
of rufous and black up to the elongate, solid black ocelli, framed very effectively basally
with a black-lined space of rufous, and distally by the solid, broad, golden-rufous
terminal band. On the lesser and greater coverts there is less development of pattern,
a narrow, black-lined terminal rufous bar taking the place of the ocelli. The wing
and tail-feathers are quite dark chestnut, with considerable variation as to black
mottling, some individuals showing a large amount, while in others the tail is very dark
but wholly unmarked. The rectrices are short, narrow and quite curved, the outer
ones being almost sickle-shaped, an interesting reversal of the conditions in the adult
male, where the outer ones are straight and the central ones curved,

WHITE-TAILED WATTLED PHEASANT 163

First YEAR PLumMacE.—The change from the juvenile to the plumage of the
first year is marked chiefly by the complete loss of the ocellus pattern. The change
in the dorsal plumage of the body is very slight.’ Instead of the black vermiculated
pale buff feathers we find warm reddish-buff tones, the contrast showing the new
feathers as generally darker and warmer, with a decided terminal fringe. This,
however, has no tinge of the ultimate steel-blue, but is rather a purplish or wine colour,
rather indefinite in tone, but clearly distinct from the remainder of the feather. This
is especially marked on the lower mantle and back. The new secondaries are much
darker, showing a reduction in the rufous vermiculation, this being confined to the
outer two-thirds of the outer web. ‘The tail-feathers are more variable, always pre-
‘dominately chestnut, but in some individuals with a great deal of black mottling. They
are considerably larger and broader than the juvenile rectrices.

The change on the ventral surface is from pale buff to darker rufous feathers, with
more distinct irregular, narrow cross-bars. Anteriorly, on the upper breast, the rufous
darkens, and in certain lights a distinct crimson tinge may be observed on the fringe,
prophetic of the strong crimson collar of the succeeding plumages. The dorsal head
plumage comes in darker, but the chin and throat are still pale ashy brown. ‘The facial
area loses some of the tuftlets of tiny feathers, and the wattles become distinct fleshy
lobes. In no newly-moulted bird in first-year plumage have I observed any trace of
steel-blue margins, only the darkened or crimson-tinged fringe which I have mentioned.
Weight, 12 lbs. Length, 488; bill from nostril, 17; wing, 215; tail, 145; tarsus, 85;
middle toe and claw, 60 mm.

YounG MALE IN TRANSITION, MOULTING INTO SECOND YEAR PLUMAGE. —A
bird shot in June is in full moult. The wattles are not much further developed than
in the younger birds, but the face is somewhat more bare of feathers. At this moult
the juvenile ninth and tenth primaries are renewed for the first time. In this individual
the thirteenth secondary of the right wing and two back feathers have been lost by
accident and renewed a month or more earlier, and the new feathers show narrow but
clear-cut steel-blue margins, very conspicuous among the rufous and black vermiculated
feathers, the blue colour evidently becoming strongly developed in any feather which
grows in a few months after the moult from the juvenile.

SEcoND YEAR Martre.—The male White-tailed Pheasant after its second annual
moult bears quite a close resemblance in body plumage to the adult, but still carries
a chestnut tail. The principal difference in the contour plumage is the much less
development of the convex, terminal band of shining blue. The crimson of the neck
and anterior body consequently is less concealed, and stands out very conspicuously.
In fact, the rich crimson or wine colour is at this stage very pronounced on all the
dorsal plumage, the terminal blue bands being very narrow. The feathers of the crown
are dark purplish-black, considerably mottled with rufous. The wing-feathers show
no trace of white. As early as two months after this second annual moult is completed,
if any of the chestnut rectrices or upper tail-coverts are lost, they will be replaced by
pure white feathers, or with the white marred only by a terminal patch of dark brown.
And if this be an outer tail-feather, the web will be very defective and much of the

164 A MONOGRAPH OF THE PHEASANTS

shaft bare, showing that the congenital degeneration is evident long before the time
for the normal growth of this plumage. The size of this adventitious feather is not
correlated with the adult colour, but is nicely adjusted to the length of the chestnut
feathers growing on either side. Thus while the normal chestnut outer tail-feather of
the second-year plumage is about 115 mm., and the corresponding white feather of a
fully adult bird 220 mm. an adventitious outer rectrice appearing during the second
year will be nearer the lesser than the greater length. Thus we see that the static
pigmental and structural change in the blood long precedes that of size, and whether
it is a direct adaptation or not, the danger to the bird at this age of undue con-
spicuousness together with a hindrance of the normal use of the tail in flight, is certainly
avoided. The presence of a single long, projecting white feather would assuredly be
a considerable handicap to the immature pheasant. It was from exactly such a case
as this—the asymmetrical appearance of a second-year tail with an adventitious white
feather or two, that confirmed Sharpe’s suspicion that the chestnut-tailed bird was only
the immature of the white-tailed adults.

Throughout this year’s growth there is considerable increase in the size of the
wattles, and after they have reached a very moderate length the occipital wattles show
their characteristic, darkened, bifurcated tips. But it is not until several months after
the next moult into fully adult plumage that they attain full size, at the period of the
next breeding season.

Bill from nostril, 19; wing, 248; tail, 188; tarsus, 88; middle toe and claw,
63 mm.

PaRASITES.—White-tailed Pheasants appear to be particularly subject to the
presence of Mallophaga, and seldom do we find a bird without numbers of the empty
flattened egg-cases tightly attached to the barbs of the ventral plumage. I have seen
some cases where there were so many hundred of these, and so evenly distributed, that
they gave the appearance of some normal structure of the feathers themselves.

EARLY HISTORY

It is remarkable how long this splendid species of pheasant evaded discovery
throughout the first fifty years when collectors were gathering specimens of vertebrates
from various parts of Borneo. Wallace never apparently heard of it, and as late as
1874 Count Salvadori did not include it in his “Catalogo sistematico degli Uccelli di
Borneo.” In December of that same year, however, Sir Hugh Low obtained a specimen
in the mountains bordering the upper Lawas River in the Sultanate of Brunei, and
gave it to Sir Henry Bulwer, Governor of Labuan. The latter gentleman forwarded
and presented it to the British Museum, and Sharpe at once described it, naming it
after the donor. In 1875 Gould figured this specimen in his “ Birds of Asia.”

Two years later a pair of Wattled Pheasants was obtained alive, the male dying
in Java, the female reaching Amsterdam, but surviving only a few weeks. Sclater
figured the female in the “ Proceedings of the Zoological Society” for 1876. In the
following year a pheasant was obtained by a succeeding Governor of Labuan from the
Lawas River with rich chestnut tail-feathers, and the concensus of opinion of Sharpe,

WHITE-TAILED WATTLED PHEASANT 165

Sclater and Gould was that it was undoubtedly a new species to which the former gave
the name of castaneicaudatus. Sharpe’s taxonomic intuition, however, led him to add,
“I must confess that the smaller size of the wattles and the plumed head induced me
to consider this new bird as being probably Z. du/werz in an intermediate stage, perhaps
in the second year.”

Subsequent specimens proved the entire truth of this suspicion, and thus we
account for the only synonym which has been applied to this species—a most welcome
simplicity of nomenclature after such unfortunate taxonomic tangles as surround the
specific identity of the species of Lophura !

SYNONYMY

Lobiophasis bulwert Sharpe, Ann. Mag. Nat. His., [4] XIV. 1874, p. 373 [Lawas Mts., Sarawak]; Gould,
Birds of Asia, VII. 1875, pl. 13; Sclater, Proc. Zool. Soc. 1876, p. 465, pl. XLIV.; Sharpe, Ibis, 1879, p. 267;
Burbridge, Gardens of the Sun, 1880, p. 61; Treacher, Ibis, 1888, p. 413 [Kinabatangan]; Everett, List Birds

Borneo, 1889, p. 198; Sharpe, Ibis, 1892, p. 422 [Mount Dulit]; Grant, Cat. Game-birds Brit. Mus. 1893, p. 292;

Hose, Geogr. Jour. 1893, I., p. 201; Hose, Ibis, 1893, p. 442; Sharpe, Ibis, 1893, p. 548 [Mount Kalulong];
Sharpe, Ibis, 1894, p. 544 [Mount Mulu, 1000-2000 ft.]; Grant, Hand-book Game-birds, I. 1895, p. 249; Sharpe,
Hand-list Birds, I. 1899, p. 35 ; Biittikofer, Notes Leyden Museum, X XI. 1900, p. 271 [Mount Liang Koeboeng];
Oates, Cat. Eggs Brit. Mus., I. 1901, p. 53, pl. V., fig. 2; Finsch, Notes Leyden Museum, XXVI. 1905, p. 142
[Upper Mahakam, Dutch Borneo]; Beebe, Zoologica, I., No. 15, 1914, p. 273 [sequence of plumages].

Lobiophasts castaneicaudatus Sharpe, Proc. Zool. Soc. 1877, p. 94 [Lawas River]; Gould, Birds of Asia,
VII. 1877, pl. 12; Sharpe, Ibis, 1879, p. 267.

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"THE JUNGLEFOWL

—_—

GALLUS
THE JUNGLEFOWL

Family PHASIANIDAE
Subfamily PHASIANINAE

Genus GALLUS

TuHRouGH the form of the domestic fowl, this group of birds is familiar to more
people than any other which comes within the scope of this monograph. The breed
known as the black and red game is very close to the wild Red Junglefowl, which may
stand as the type of the genus.

Admitting four species as comprising the genus, we find that they are birds of
medium size, characterized in the cocks by the following features. The bill is stout,
fairly short and curved, and the feet are strong and pre-eminently fitted for scratching:
There is a large, erect, fleshy comb on the top of the head, extending from the base
of the bill to behind the eyes, with the margin serrated or entire. The sides of the
face, chin and throat are bare, either with two pairs of wattles situated below the
ears and on each side of the throat, or a single median wattle down the middle of
the throat.

~The tail is composed of seven or eight pairs of feathers. This organ is strongly
laterally compressed, and slightly graduated except for the central pair of rectrices,
which are much longer than the others, soft in texture and curved, resembling the
upper tail-coverts. These are about twice as long as the second pair, and four times
the length of the outer rectrices. The first primary is considerably shorter than the
tenth, the fifth being the longest. The tarsi are longer than the middle toe and armed ©
with well-developed spurs.

The feathers of the rump are long and lanceolate, and the hackles of the neck are
of the same character, or if truncate have a specialized curve in the vane near the tip,
and show iridescent colouring.

All four species will cross with one another, and the hybrids are more or less fertile
among themselves. The moult is typically Phasianine, that of the tail being from the
outer rectrices inward. In the females the comb is rudimentary, while the wattles,
specialized hackles and central rectrices and spurs are lacking. Thus we see that the
secondary sexual characters are the comb, wattles, hackles, central tail-feathers and
spurs. ;

The only one of these characters which may be taken to distinguish the genus
VOL, II 169 WE

170 A MONOGRAPH OF THE PHEASANTS

is the comb. The compressed tail is found in Lophura, Acomus, Lobiophasis and
others ; the lateral wattles and the hackles in Chrysolophus, and the median wattle in
Tvagopan.

The four species fall into two quite distinct but unequal divisions, good subgenera
they might be called; first, gallus, lafayetti and sonnerati, and second, varius. These
have been considered as two genera (Gal/us and Creagrius) by a few writers, e.g. Ghigi,
1903. Varius possesses the peculiarities of a smooth-edged comb, a median throat
wattle, truncated neck hackles and an extra pair of rectrices. Taking the group as a
whole, however, the hiatus between the four species and the nearest related genera
seems much greater than between the two groups themselves. So I choose to keep
them together. And here comes in the question of logicality; whether by doing this
I have not been somewhat illogical in comparison with other generic divisions. This
is of not the slightest moment to me. No two genera of the Phasianidae or any other
group of organisms, as now isolated by time and space on the earth, are separated from
each other by exactly the same intervals of character distinction. Classification, we all
admit, is merely the make-shift, incident upon, and made necessary or indeed possible
by, our ignorance of intervening forms. Hence relative clarity of interrelations is its
sole aim. In this instance the genus Ga//ws, considered as embracing all four forms,
expresses much more exactly the homogeneity of the quartet as a whole than would
the isolation of varius, such segregation setting it as far apart from Gallus as is the
genus Chrysolophus.

There is no doubt that the Red Junglefowl alone is the direct ancestor of all of
our domestic poultry, so this question is removed from the discussion. Study of the
plumage of the four species shows a tangle of characters, which can be logically oriented
only when we think of all four birds having descended from some form quite different
and much more generalized than any of them are to-day. For example, taking the
cocks first, the ventral plumage brings ga//us and varius close together; the dorsal
surface shows a close similarity between gad/us and lafayetti. Sonnerati, while it has
a general body plumage of a much more generalized pattern and coloration than any
of the others, has hackles and wing-coverts more specialized than in the other three,
the peculiar sealing-wax-like spots deserving of as great distinction as some would give
to varius for its peculiar characters.

The hens, on the contrary, show propinquities entirely unlike those of the respective
cocks. The ventral plumage associates closely /afayettd and sonnerati, while the
dorsal patterns and colours indicate an affinity between gal/us and sonnerati, and the
generalized black and white wing-bars link /afayett7 and varius.

GALLUS
Type.
Gallus Linnaeus, Faun. Suecica, 1746, p. 61 : : : : : . G. gallus.
Alector Klein, Hist. Av. Prodr. 1750, p. 111 . f . : : . G. gallus.
Creagrius Gloger, Hand- u. Hilfeb. 1842, p. 387 . : . ; : . G. varius.

This well-marked group of four species is widely distributed throughout India,
Burma and the Malay Peninsula, Ceylon and Java. Where it occurs outside of these
regions it has probably been introduced by man. The four species are as follows—

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MAP SHOWING THE DISTRIBUTION OF THE JUNGLEFOWL.

Region 1. Gallus gallus.
96 D *) lafayetti.
9 3. i. sonnerati.
4. 3 varius.

”

THE JUNGLEFOWL 171
Red Junglefowl . : ; : ‘ : ‘ : : . Gallus gallus (Linnaeus).
Ceylon Junglefowl . ; ; 3 : : ; : . Gallus lafayetti Lesson.
Gray Junglefow!l . : ; ; ‘ ; ; : ; . Gallus sonnerate Temminck.
Javan Junglefowl : ; ; ; ; : : : . Gallus varius Griffith.

KEY ORG AEE GS

I. Comb and spurs largely developed; middle pair of tail-feathers elongated (males).
a A wattle on each side of the throat; tail of fourteen feathers.

a Lower plumage black : : : : 3 ; : ; ; : ‘ . gallus.

6’ Lower plumage orange-red ‘ : ; . lofayetii.

c’ Lower plumage black, edged with ey ta with a bites here sis : ; . sonneratt,
b A median throat wattle; tail of sixteen feathers . : : . varius.

I, Comb rudimentary ; spurs aieeaes middle tail-feathers of normal shes (erales)!
a Lower back reddish-brown, anes mottled with black.

a Breast pale reddish- zone ; gallus.

o’ Breast white, fringed and marked WR Slade.
a’ Outer web of secondaries widely barred with buff and black é : b . lafayette,
6” Outer web of secondaries brown, finely mottled with black . 3 : 1 . sonnerate.

b Lower back black, fringed and centred with buff . : : 3 : ia . varius.

RED JUNGLEFOWL
Gallus gallus (Linnaeus)

NAMES.—Generic and specific: Gallus, Latin gal/ws,a cock. English: Red or Wild Junglefowl. German:
Kaminhuhn ; Bankivahuhn; Wildhuhn. French: Cog bankiva. Native: Jungli murgh 2, Jungli moorghi °, Bun
moorghi (Upper India) ; Kukar, Kukra, Bun-kookkoor, Bunkokra (Bengali, Lower India and Assam); Nagtse-ja,
Natsu-pia (Bhutan); Tang-kling, Pazok-tchi (Lepcha, Sikhim); Beer-seem (Koles); Gera-gogor ¢, Kuru ?
(Gonds) ; Lall (Chanda); Tanghet or Tanquet (Burma) ; Ajam-utan (Celebes). In the Malay States the following
names are current: Manuk (Negritos); Nanak (Northern Sakai tribes of Upper Perak); Dénak, Puk Dénak
(Central Sakai of Batang Padong); Ayam Dénak, Ayam Mérih (Forest fowl, Besisi of Selangor and Negri
Sembilan) ; Pok Brugak (Jakun of Pahang); Ayam Utan, Ayam Dénak, Ayam Borga (Junglefowl, Decoy fowl
and Wildfowl respectively ; Malay).

BRIEF DESCRIPTION.—Male: Crown, neck and hackles golden brown to orange-red, passing into straw
yellow on the longer hackles. Mantle, smaller and greater coverts metallic green or purple. Scapulars, back and
median wing-coverts dark maroon, shading into orange-red on rump. Exposed parts of secondaries cinnamon.
Tertiaries and tail metallic green. Lower parts black. Face and throat thinly feathered; a serrated comb and
two gular wattles. Female: Rusty red on head, shading into orange on neck and pale yellow on mantle with
median black stripe. Upper parts reddish-brown, finely mottled with black. Secondaries mottled with pale

brown on outer webs. Fore-neck chestnut. Rest of under-parts pale light red. Face thinly feathered, comb
very small.

RANGE.—North-eastern and Central India, south through Siam, Cochin-China and the Malay Peninsula to
Sumatra. Introduced in many other islands of the East Indies.

TE BIRD UN: ALS: WiahisD TOME

Far up in the wilderness of Northern Burma, close to the Chinese border, I left
camp one morning before dawn, shivering in the cold December dusk. As I brushed
against the wet undergrowth, some bird now and then uttered a startled chirp, or, not
yet fully awakened, began a sleepy refrain. No other sign of life came from the jungle.
The air was heavy with the overpowering sweetness of a multitude of white blossoms
draped on vines from tree to tree. Within the next two hours I sought and found the
place where a flock of kaleege pheasants passed over the ridge every morning from
roost to water.

On my return the sun was already well up and the early morning chorus in full
swing. At this season the jungle trails are alive with butterflies. As one walks along
they rise in myriads, alighting on one’s clothes and hat, brushing one’s face like a flurry
of leaves in autumn. Scores of species of all the spectrum’s hues are seen within the
space of a few yards.

Birds are eagerly hunting their breakfast, but snatching every spare moment for
song or cry. Rollers fly past, flashing their brilliant heliographs of blue from beneath
the dull brown wing-coverts. As we approach a straggling Shan village, the under-
growth thickens and small birds increase. A flock of at least fifty silver-eared mesias
drifts past, then a gorgeous yellow and black Indian oriole, and a jewelled sunbird

shoots up like a meteor.
172

PLATE XL

RED JUNGLEFOWL

Gallus gallus (Linnaeus)

To the human race this is the most important wild bird living on the earth, for it represents the
ancestor of all varieties of domestic fowl. It ranges from the border of Kashmir to Singapore, and is
found in the wildest regions, as well as close to native villages, with the fowls of which the wild birds
frequently cross. Its crow is bantam-like, and sounds strangely out of place when heard in deep jungle.

These Junglefowl are usually monogamous, the hens lay from five to eight eggs.

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RED JUNGLEFOWL 173

From the distant village comes the long-drawn, wavering crow of chanticleer,
and then pandemonium in the shape of two hundred mules loaded with rice and
ammunition, and guided by an escort of swarthy Punjabis. The jingling of bells,
screaming of coolies, and undisguised complaints of the mules themselves fill the air
with a continuous din, as I crouch unseen in a clump of bushes near the trail which
leads to China. Finally, the last falsetto shriek dies out in the distance, and a dyal-
bird sends up his clear, sweet phrase, clearing the jungle air and giving it over again
to the sounds of the wilderness.

Two hundred yards farther on, the edge of the jungle opens upon the encampment
of the preceding night, a disordered litter such as only ten score of army mules can
leave. The scavengers are already at work, doing their part to re-sweeten the field
again. Forty great black pigs, several vultures with others circling downward each
minute, a score.of many-hued doves and dyal-birds, all at work, the first snuffling
among the remains of the forage bags, the latter daintily catching flies—those scavengers
of a still lesser world of life.

Suddenly in the very midst of it all my eye caught a bit of incongruity. Walking
with quick, nervous steps among the lethargic pigs and the crowding, scratching hens
was a wild Red Jungle cock. Every poise of body, every turn of the head spelled
caste. Never for an instant was he off his guard. Only at the rarest intervals did he
snatch some morsel of food.

Once a native cock approached with ruffled hackles; the wild cock gave a sudden
turn in the direction of the threat and the coward took to instant flight. My glasses
showed the sleek, slender body, the dark, trim legs and the sinister black curving spurs
capable of dealing death with a single thrust. Once, and once only, did the jungle bird
forget himself fora moment. He crossed the path of a brown hen, much too big for
any strain but that of the farmyard, yet without the white and blacks which marked
the other hens. She had retained the jungle tints, and as if he recognized it, the wild
cock turned aside and made one quick, brilliant obeisance. A narrow circling-turn
sideways to the hen, inner wing dropped, back flattened, hackles and tail spread. Just
for a moment he was agleam, the sun reflecting metallic red, green and purple from his
plumage. Then a great lumbering pig interrupted my view, and when seen again the
bird was quietly making his way through the throng, unnoticing, but not unnoticed.
Not a cock or a hen but ceased feeding or scratching when he passed near. They
recognized him as something alien, perhaps as superior, certainly to be respected, for
they took no liberties with him.

All this came clearly through my glasses, and then, when about to walk on, I saw
the jungle cock give one leap and vanish behind a low bush. The other creatures paid
no attention to the cause of the alarm, and only when I stealthily stood upright did
I perceive the native chowkidar, who had left the dak bungalow and was making his
way down to the river.

The cock crept carefully off toward the jungle, with never a stop for even a snatch
at a morsel of food. Ten minutes later, some distance to my left, I heard his sharp,
decisive double crow, so free of quaver or drawl, so like to the wild call of his pheasant
kindred : cock-ka-charr !

After a hasty cup of cocoa and a cracker I started on a stalk, and creeping up to

174 A MONOGRAPH OF THE PHEASANTS

the top of the ridge in the still early morning sunlight, I found me a lowly place beside
a small fallen log. The wood upon which I rested was torn with the shot which had
laid low a Horsfield’s kaleege two days before, when we needed fresh meat for the
pot, but no pheasants were hereabouts this day. I idly watched a huge grasshopper
creeping slowly up a dead sapling, and unconsciously made a sudden motion to brush
away a cloud of mosquitoes which had found me out. At once a great outcry arose
only a few yards away down the slope—a jungle hen had perceived that something was
wrong on my side of the thicket, but whether a—to her—harmless water-buffalo or a
real danger she evidently could not make up her mind.

By hitching along and wriggling worm-like over a few feet of wet ground I came
within sight of the bottom of the gully. Several large clumps of bamboo were browsed
to hour-glass shape by the—to me—fearfully dreaded self-same buffalo. Just below the
yellow-green foliage was a wallow, partly filled with water, and at the edge of this,
scratching daintily in the damp soil, was my jungle cock of the mule-field and his
mate.

The cock allowed no fall of leaf or twig to escape him, and it was interesting to
watch how, every second or two, he systematically swept the sky and the woods all
about. The hen, evidently relying on his alertness, devoted all her attention to feeding,
and her chuckle of content as she drew forth a large, protesting worm was delightful
to hear.

When a squirrel rushed through the bamboos and loosened a bunch of large leaves
which eddied downward, the jungle hen gave her loud, strident cackle, cut! cut! cut !
cut ! cut-dd-cut! exactly as a domestic hen announces an egg. In this case it was
a startled exclamation of suspicion, given rapidly and sharply.

From some distance within the bog a forest cat was yowling querulously, but the
junglefowl paid no heed until the sound ceased. Then, although they fed for a few
minutes longer, much of their attention was concentrated in the direction from which
the sounds had come. At last both listened intently, their heads drawn up to full
height, and both started up the bank toward me.

But, as so often before in my experience, the clan of wood creatures came to one
another's aid in an eminently effective manner, although so wholly unconscious of any
altruism. A wretched chocolate squirrel and a quartet of laughing thrushes made my
ears tingle with their expletives. Scarcely had they begun their tirade at me when the
junglefowl made a quick dash down one side of the gully and passed out of sight
for ever. .

GENERAL DISTRIBUTION

The present range of the Red Junglefowl is very extensive, but we must draw
a sharp line between what appears to be the original area of distribution, and the
recent extensions brought about by man. In northern India the bird extends along
the terai and the southern outer ranges of the Himalayas, from the head of the
Assam Valley beyond Sadiya, westward through southern Sikhim, Nepal and the
Siwaliks of Garhwal to Kashmir. In the Punjab it occurs only in isolated, especially
suitable districts, so sporadically, in fact, that the western extension from its more

HOME OF THE RED JUNGLEFOWL

Burma I found he wild Junglefowl coming out to feed along the trails which lead from
. They cluck and scratch among the turf, and take dust baths exactly like our domestic
ave often found it difficult to shoot them, as the action seemed like unsportsmanlike slaughter

eee ae eo of danger, however, they lower their tails and run headlong, like

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RED JUNGLEFOWL 175

extensive eastern distribution takes the form of a long, narrow finger laid along the
southern Himalayan slopes. It thus rather closely parallels the range of the four
Himalayan species of kaleege pheasants.

Although absent from the level, alluvial, semi-arid plains of Upper India drained
by the Ganges, it reappears to the south in the northern and eastern portions of the
Central Provinces and on southward as far as the Godavari River, especially toward
the east coast, thinning out toward the west, and finally disappearing as we approach
the line of central India.

Eastward we find it common, throughout the hilly portions of Bengal, the
Sunderbans and the whole of Assam, including the Khasi, Géro and Naga Hills,
Cachar and Sylhet. The connection across the Ganges River is a rather narrow one,
as the bird is not found in the semi-arid parts of Behar to the north, nor in the
southern delta district. Still farther east we find its centre of distribution throughout
Burma, including Pegu and Tenasserim, western Yunnan, Siam, Cochin China, and
southward in the Malay Peninsula more particularly in the western half, and generally
distributed in the island of Sumatra. It is absent from the island of Singapore.

In all this area I believe the Red Junglefowl to be more or less indigenous
with slight changes in the general contour of the range, but showing no radical or
important increases since mankind began exerting a direct effect on the environment
and range of the species.

Severtsov and David state that the bird occurs in Western Turkestan. This is
an error, except as domestic fowls are found associated with the natives. As the
result of careful inquiry, I am very certain that the Red Junglefowl does not cross
the Himalayas at any point, neither into Persia, Yarkand, Afghanistan or Tibet.

It is entirely absent from Borneo, but has been recorded as a more or less
feral inhabitant of Java, Timor, Lombok, Celebes, the Philippines, BalAbac, Palawan
and Hainan. Also in many of the isolated archipelagos and islands of the South
Seas, such as Tahiti, Tonga, Viti, New Caledonia, the Great and Little Cocos,
Ponapé, etc.

A few quotations giving the condition of the Junglefowl in some of these out-
lying islands, will be of value in supporting the supposition that they are all
domestic birds run wild. There is no question about this fact in such isolated
islands as Tahiti, but the fact of interest here is the apparent beginning of reversion
to the wild colour type.

TAHITI

“The common dunghill fowl is found wild in the forests here. Some of the
residents think it is a Junglefowl, peculiar to the country, but, upon examination, I
have not been able to perceive any material difference between it and the domesti-
cated bird, and therefore incline to the belief that it is the common species returned
to its original habits. In my excursions, I have killed about a dozen of them.
Their plumage is generally more rich and brilliant than that of the domesticated
bird, and there is not so much variety in the colour of different individuals. Their
flesh is exquisite. They are very shy, running away with singular rapidity, and
concealing themselves on the approach of the sportsman, When flushed, they fly

176 A MONOGRAPH OF THE PHEASANTS

with great vigour and swiftness, and where the trees and bushes are not too dense,
afford a very good mark” (Townsend, “Journey . . . to the Sandwich Islands,” etc.
1839, p. 289).

PONAPE

“Of the specimens from this locality, the male shows no difference from
specimens from Sumatra, except that the primaries are not cinnamon, but dull rusty
brown, and the wings are a trifle longer. The female differs a good deal... I am
inclined to believe that the differences which this bird shows are due to a domestic
state” (Finsch, “Proc. Zool. Soc.,” 1877, p. 780).

SuLu IsLAND

“This jungle-cock, although very numerous on Sulu Island, is but rarely seen,
owing to its haunting the thicker jungle and being very shy... . After a few days’
captivity they become even tamer than domestic fowls, and are freely crossed with
the latter by the natives” (Guillemard, “ Proc. Zool. Soc.,” 1885, p. 272).

And again, “Junglefowl are plentiful on Kushai and Ponapé, but on both
islands they are not native species, but the offspring of imported tame birds, which
have run wild and have returned to the original plumage of Gallus ferrugineus.”

PELEW AND Uap IsLANDS

“Captain Peters sends a single female (from Pelew), but unfortunately he does
not state whether the Junglefowl lives in a wild state in the islands, or as in Uap,.
as Mr. Kubary remarks, only in a half-wild state.”

WETTER, NorTH OF TIMOR

“Wild fowls were met with in great numbers on Wetter, but the great variation
in the males and the geographical distribution at once suggest that they are feral.
Some of these birds, in fact the majority, are indistinguishable from Gallus
ferrugineus from India and China, though some have very short tarsi. Probably
either wild-caught G. /errugineus have been introduced, or more likely a race of
domestic fowl descendant from Gallus ferrugineus has run wild and thus the jungles
are inhabited by an apparently quite wild fowl” (Hartert, “ Novitates Zoologicae,”
XI. p. 176).

NEGROS

“The collection contains a female Junglefowl which is entirely black, with
greenish-metallic reflection.

‘At first I much doubted if such a melanic specimen was likely to be of wild
origin. Mr. Keay, however, assures me that the bird was not a roving specimen.
It was seen for several days on the margin of the forest, in company with a
normally-plumaged cock, and was finally secured by Mr. Keay with some difficulty,
as it was very wary and smart on the wing. When shot it towered to a height of
over 150 feet, and then fell dead.

‘Mr. Keay is convinced that it is a genuinely wild Junglefowl; and, if measure-

RED JUNGLEFOWL 177

ments are any guide in the case, the specimen is certainly only of normal dimensions,
the wing being 7°5 inches. On the other hand, I am unaware that this species is
variable in plumage, and can find no mention of its being liable to melanism”
(Clarke, ‘‘ Ibis,” 1900, p. 360).

JAVA

My own researches in this island have led me to the certain belief that the Red
Junglefowl is a recent introduction, an intruder into the territory of the true Javan
bird (Gallus vartus).

Many more instances might be adduced of quite clear proof of the running wild
of domestic fowls, and while I shall allude to this in another connection, I mention
it here only to emphasize the importance of trying to limit the range of this bird
to the natural, normal boundaries. I admit Sumatra as a valid locality, because of
the corresponding distribution of several species of pheasants, such as the Malayan
crestless fireback (4comus erythrophthalmus) and the Malayan argus pheasant
(drgusianus argus).

I have mentioned the fact that the Junglefowl occurs in the Sundarbans, which
is the name in use for the extreme delta region of the Hoogly and other rivers
which are derived from the Ganges. Concerning these birds Mr. Rainey writes:
“T may add that the Junglefowls in the Sundarbans appear to be descended
immediately from domestic fowls, which used to be let out there in considerable
numbers by superstitious wood-cutters to propitiate the sylvan deities—a practice
still prevailing to some extent—and I have shot these birds there in different stages
of transition. This is interesting, as we evidently thus find the domestic fowl
reverting to its pristine condition, for the Red Junglefowl is undoubtedly the origin
of our tame varieties of fowl. I had a couple of chicks produced from eggs of wild
birds set under the domestic fowl, and they remained contentedly in the poultry-
yard, and freely bred—they were both hens—with the tame fowl. The progeny were
in appearance midway between their parents, and exactly similar to some I had shot
in the Sundarbans. About that time the cyclone of 1867 swept over the place I
was residing at, and of course put a premature end to the varied denizens of the
poultry-yard, hybrids included. I soon afterwards left my abode in the wilds of the
Sundarbans, and have had no opportunity since of continuing the experiment.”
There seems little doubt that all the birds of this isolated, unique region are actually
domestic birds run wild, so that even on the mainland, adjoining the natural haunts
of the birds, we may have, as in this case, a zone stocked by man, where the birds
have become almost indistinguishable from the feral individuals elsewhere.

GENERAL ACCOUNT

Of all the pheasants, indeed of all the birds in the world, the Red Junglefowl
stands first in importance to mankind on the earth. From this species, and this
alone, all the forms of domestic fowls have taken their origin, and hence from the
point of view of economic utility this bird assumes a position of great interest.

With this point of view in our general account of the species we are not directly
VOL. II AA

178 A MONOGRAPH OF THE PHEASANTS

concerned, but it is well to state the fact at once, so that the details of its life
history may not be forgotten when we come to compare the feral and the domestic
birds, and to attempt to trace the part which the cock and the hen have played in
the history of mankind.

It is difficult to correlate the habits or to present them in a uniform, inclusive
summary in the case of a bird such as this, which ranges from the borders of
Kashmir to the southern limits of Johore. I met with the Red Junglefowl in many
countries and watched it under many varying conditions, and in most cases I have
found it necessary to do away with the convenient generalizations which are so
easy to make in the case of pheasants of more restricted range, living under more
homogeneous ecological conditions.

I believe the home range of the Junglefowl is of quite limited extent. I have
been told by tea-planters of a family of these birds which inhabited a certain bit of
bamboo jungle month after month, apparently never leaving it except to make short
excursions for water and food. And these sedentary habits are, I believe, more
pronounced in this group of birds than in most of the more typical pheasants.
During the breeding season the birds which frequent some given area are seldom
seen, retiring into the deeper, denser parts of the jungle. But there is no extensive
wandering. Where wild birds get into the habit of feeding upon crops, or associating
with domestic fowls, they correspondingly limit their home range even more, and
when not molested, their comings and goings can be timed as accurately as in the
case of the barnyard fowls themselves.

Taking three thousand feet as an average maximum height, we find the Jungle-
fowl living at a comparatively low elevation. The great proportion are not exposed
to any but slight changes in annual temperature, and hence have no cause to make
even limited seasonal migrations. The birds which range up the valleys and southern
slopes of the Himalayas, are found breeding at a maximum height of about five
thousand feet, while occasional records show that exceptional individuals may wander
upward two thousand feet higher. These, of course, are affected by the approach of
cold weather, and as I observed in Garhwal, descend several thousand feet to warmer
altitudes, reascending when the warm weather again returns. So we see that with
these local exceptions, no regular seasonal migration is to be found among Red
Junglefowl.

While the name Junglefowl is quite correct, bamboo-fowl would be even more
appropriate, as they are especially fond of this type of vegetation, and as I look back
over my memories of these birds, any visualizing of them is invariably accompanied
by a background of the tall, curving stems and the soft foliage of bamboo. —

Rarely I have found them in heavy tropical forest devoid of bamboo, and in
the vicinity of the semi-arid plains they must needs be content with the shelter
afforded by clumps of acacias and other similar plants.

Throughout India the Junglefowls live of necessity more or less close to
cultivated districts, and indeed, even in the less settled parts of Burma and the
Malay States, I found them far more often in the vicinity of native villages than in
the trackless depths of the jungle. This fact must be taken into consideration in
speaking of their comparative abundance. On the whole Red Junglefowl show a

RED JUNGLEFOWL 179

remarkably even distribution. In a country suited to them and within their general
range they are almost certain to be found. That is to say, there is not the inex-
plicable zzter habitat hiatuses which characterize some of the true pheasants.
Exceptions, of course, are to be found where they have been exterminated by human
agency from some intermediate tract of country. Considered in general they may
certainly be said to be abundant, probably the most abundant of the so-called game-
birds in the East, although it is hardly fair to make this comparison, owing to the
constant recruits to their ranks from half-wild native fowls.

Red Junglefowl are sociable birds, and, except during the breeding season, when
association is either for fighting or for pairing, they are often found in good-sized
flocks. They are found very frequently in pairs, and it is seldom that I have seen or
have heard of a solitary cock or hen.

As often as I heard the Red Junglefowl in various Eastern countries it was always
with a start at such familiar, barnyard sounds coming from the jungle, where sometimes
I knew there were no human dwellings within a distance of many leagues. When
in search of a specimen or a bird for the mess, more than one has escaped with its
life by giving utterance to such domestic sounds that I unconsciously withheld
my fire.

The crow of the Red Junglefowl, like the details of plumage and colour of wattles,
shows quite marked variation, which is rather individual than characteristic of the
birds of any one district. This is undoubtedly due to the infusion of a domestic strain
in many apparently purely feral birds. I have several times had the opportunity of
watching and hearing Junglefowl crow, in districts far from cultivated areas, the birds
showing by their low-hung tails and lank, rakish carriage that they were in appearance
at least truly wild.

On one such occasion I had concealed myself at earliest dawn in a thicket, on
either side of which ran a trickle of water. This was in the heart of the Pahang jungle,
and my choice of this particular place was influenced not only by the fact that for
several days I had heard a Junglefowl calling near by, but also that it was free from
the hordes of leeches which infested the surrounding foliage, and which made stationary
observation elsewhere an impossibility.

I became aware of the bird first by an excited cackle, which I feared was caused
by the discovery of my presence. But through the leaves I saw that its attention was
directed elsewhere. Its suspicion was soon allayed, and it came quietly on to the edge
of the dark jungle rivulet, where, after a momentary delay, it leaned down, swiftly
scooped up a beakful and again froze into a statuesque attitude of keen scrutiny. Seven
times the bird drank, then turned aside, wiped its beak upon a pebble and instantly
flew up to a bare branch about twelve feet from the ground, its wings beating rather
noisily.

Three or four minutes passed. Then the bird flapped its wings three or four times,
giving forth resounding smacks, and at once crowed, then listened intently. Again
and again the challenge rang through the jungle, without, however, any further beating
of wings. Once a distant crow was heard in answer, and this caused an excited reply
of several utterances in quick succession, followed by an equally long period of waiting.
After a half-hour the bird descended and vanished into the forest. Its cry as I wrote

180 A MONOGRAPH OF THE PHEASANTS

it down on the spot was Evr-er-érk—er! LEvr-ev-évyk—er! The accent was all on
the third note, while the fourth was sometimes very weak, and at a distance would have
been inaudible. It seemed more like an involuntary intake, or recovery of breath, than
an actual integral part of the call.

This bird was unquestionably challenging. Every motion, every attitude attested
this, and on another occasion I observed the same thing. _

But at another time in the terai I had the good fortune to watch and listen to
a bird which was crowing and yet not challenging. I knew he had a mate near by
with a nest, although I did not actually locate it until a day or two later. The Jungle
cock was on a fallen log, and his whole demeanour was of assured peace. He was alert,
but only as any wild bird is whose life is one long fight against danger. His crowing
was intermittent, uttered at a lower pitch, and did not have the tang and abruptness
of the challenge. It had a hint of the domestic drawl, and between each effort he sang
to himself the low, content song with which we are so familiar in the barnyard hen—
Waadaadaak—waak—wadaadaak ! The whole seemed to me a real song, to be
compared with the tempered notes of a thrush who perches near his mate on her nest
and gives this vent to joy in the success of his life.

The difference in the apparent human phrasing of birds’ songs when heard near
by and at a distance is very apparent in the case of the Junglefowl. In Garhwal I have
more than once mistaken the Kok! kok! koklass! of the koklass pheasant for the
Junglefowl and wce versa, when the notes were softened and mellowed by filtering
through the mist of a long length of valley. Others also have noticed this
resemblance.

More commonly heard than the 4rr-er-évk ! is a less wild, less distinctive call,
that which I take to be influenced by some domestic strain, although uttered by birds
which, in habits, are as truly feral as any bird of the jungle. When heard near at hand,
this sounds to our ear like Cak-ka-chirr ! or Cock-ka-charr—ca! This bears a fair
comparison to the crow of some breeds of bantams, and an additional interesting fact
is that the domestic birds whose crow most closely resembles that of the wild Jungle-
fowl are those whose plumage is nearest the red and black of their feral ancestors.
Even the crow of the wild birds quoted above is always, however, distinct, being shorter,
more viril and wholly lacking the final drawl which seems to hint of the degeneration
of the domestic cock.

The cackling of the wild hen, as I have already said, is given in moments of excited
suspicion. I do not know whether the direct stimulation of having deposited an egg
also inspires this note or not. I have never heard it in a captive wild-caught bird, but
several times I have known the first generation of captive-bred birds to give voice to
it. I am inclined to believe that it is a call to the cock, giving notice that the nest
duties of his mate are for the moment over. I have known a peahen to leave her nest
with a half-running, half-flying rush, and to call loudly until her mate came, when they
went off quietly feeding together.

The squawk of a wild bird when picked from a trap or a wounded one when caught
in the hand is indistinguishable from the note of terror of the domestic fowl, but it is
never continued for minute after minute as in the latter, but uttered once or twice, and
then the bird awaits its fate in silence. The cackle of the wild hen is sharp and shrill,

RED JUNGLEFOWL 181

and the cackle of fear uttered as birds are suddenly flushed is as much pheasant-like as
it resembles the corresponding utterance of a barnyard hen fluttering over a fence from
a pursuing dog. The peeps of wild chicks differ in no way from those of the fluffy
yellow offspring of our domestic hen.

In gait and carriage there is much difference between individual Junglefowl, the
same influence being at work here as in the case of the voice. The birds which haunt
the vicinity of villages have usually much more of what we are pleased to call a proud
carriage than the real jungle individuals. The tail is held higher and the movements
are slower and more dignified, as we interpret them. But once the real fowl of the
deep jungle is seen it will not be forgotten.

I have seen many so-called Junglefowl in captivity and they satisfied all the
requirements of the casual observer, who would remark that such splendid carriage
was well worthy of the ancestor of our common fowls, and similar sentiments. But
the pair of birds which arrived at the London Zoo in 1912 were almost the first real
feral Junglefowl I have ever seen in captivity. Dignity was absent; the carriage was
that of an untamable leopard; low-hung tail, slightly bent legs; heads low, always
intent, listening, watching; almost never motionless, but winding in and out of the
shrubbery, striving to put every leaf possible between themselves and the observer!
To my mind, they fulfilled every ancestral requirement much more satisfactorily than
the usual Junglefowl type. It would take more than one generation to tame them.
They were wild as the pheasants of the Himalayas. Dignity and high-bred carriage
they doubtless had, but it was reserved for their kind; for the combats and the
courtships of their own haunts, not struttings and steppings while mankind looked
on. All this is not a sentimental point of view, but a very real distinction.

I have walked along a trail and suddenly come in sight of several Junglefowl, who
looked at me, drew up to their full height and hurriedly enough, but with raised tails,
took to the jungle. And again I have walked behind a soft-padding army elephant
in Northern Burma and caught a momentary glimpse of birds, slinking swiftly from
view with outstretched necks and trailing tails, or hurling themselves headlong into
the underbrush. There is no dignity in such birds in the loose, undesirable meaning
of the word as we use it, but there is caste, high caste, the feeling of undefiled lineage,
untainted by the degeneracy of a captive strain.

The flight of wild Junglefowl is direct and swift. When amid dense undergrowth
they are able to rise almost vertically until clear of the vegetation and then fly strongly
to a neighbouring tree. This is their procedure when escaping from a sudden rush
of dogs, but they discriminate sharply if men disturb them, and without pause fly as
rapidly as possible out of sight, if on a slope invariably downhill. I once saw a cock
fly across a valley some half-mile in width, and as long as it remained in sight the
flight was as strong as a pheasant’s, a number of quick wing-beats being followed by
a few seconds of scaling. Ordinarily where the bird is in sight from its rise until it
alights, especially if its course, as usual, be downward, the wing-beats are rapid and
continuous at first, while the latter half of the flight consists wholly of a motionless
scaling with bowed wings.

This facility of flight seems all the more striking when we see it displayed by the
village fowls of many parts of the East. When, in riding over the trails or Government

182 A MONOGRAPH OF THE PHEASANTS

roads in the Malay States, one comes suddenly upon a group of fowls, instead of
blundering ahead or with frightened squawks and frantic, awkward beating of wings
flopping toward their native coops, they often rise swiftly, and with the greatest ease
fly out over the steep slope, all together like a flock of doves, then circle around and
come back to the road some distance behind. This reveals clearly the recent admixture
of wild blood, and is correlated with a much lower fertility than exists among more
carefully-bred domestic strains.

DAILY ROUND OF LIFE

The food of Junglefowl, in keeping with their other rather generalized habits, is
of an omnivorous character. They are, however, graminivorous on the whole, and
in the crops of birds which I have shot at a considerable distance from cultivation
I have always found vegetable matter predominating. Small nuts and seeds of
unknown plants are often taken in considerable quantities, besides which the young
shoots of bamboo and other grasses, leaves, petals and wild seeds of all kinds are eaten.
As with all other pheasants, the larvae and eggs of white ants or termites are enjoyed,
and the birds will go to considerable trouble to scratch and peck away the hard earthen
tunnels and mounds of these insects to expose the nurseries with their abundant
tenants. A certain discrimination seems to be exerted in the choice of pebbles, which
are swallowed, and the gizzard is often partly filled with bright and shining bits of
stone. Seventeen sapphires have been taken from a single hen Junglefowl.

The greater number of birds which make their home in the immediate vicinity
of native villages and cultivated fields lose no opportunity to make the most of such
advantages. Associating so freely with the domestic fowls, they are thus protected to
a great extent from direct attack with guns and traps, the natives not wishing to sacrifice
their own useful fowls. After the harvest has been gathered, Junglefowl in early
morning or late evening may be seen gleaning in the grain-fields, in loose flocks of
ten to twenty birds. The damage they do, however, is not great, and their wariness
makes it an easy matter to drive them from the fields.

Besides fattening on the millet and rice when these grains become ripe, Junglefowl
share throughout the year the habits of domestic birds in haunting cow-sheds, and the
sign of animals which travel over the jungle roads and trails. The easiest way to catch
at least a glimpse of these birds in any locality which they are known to inhabit is to
walk quietly along the roads near the forests in the cool of early morning. Even where
wild cocks and hens form close associations with domestic birds they usually keep to
their feral habits in the matter of feeding. In the higher parts of their range in the
Himalayas they may occasionally be seen at midday with the village fowls. But this
is very unusual, and the early morning and the evening are the only times when,
throughout their range in general, they are seen feeding. During the heat of the day
they retire to the deepest, coolest shade in the vicinity, and during this time can only
be discovered or dislodged by a dog, or by the accident of stumbling upon them. At
such a time they are very disinclined to take to wing, and unless actually forced to fly
they invariably make their escape by swift running and dodging around and through
the undergrowth,

PHOTOGRAVURE 31

ROOSTING. ees DING PLACES OF THE RED JUNGLEFOWL

Turse birds roost high up, often on the half-bent curve of a tall bamboo, whose smooth stem ensures
safety from arboreal carnivores. As many as thirty birds have been seen roosting close together.

Just as tame fowl are fond of the company of barn-yard cows, so the wild Red Junglefowl are often
seen in the vicinity of wild cattle, and I have seen them again and again feeding about buffalo wallows.
It is almost impossible to flush these birds. They invariably choose to escape by running swiftly away,

yet, when surprised by a dog, they have the ability of rising as quickly and strongly as partridges,

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Unlike a number of the true pheasants, Junglefowl are essentially gregarious in
their roosting. In Tenasserim this seems to be carried to the extreme, as the following
note suggests: “On one occasion, near Pahpoon, I counted thirty males and females
seated side by side on one enormous bent bamboo. Mr. Hildebrande was with me,
or I should not have ventured to record the fact. I counted them carefully through
my binoculars. They were at the other side of the Younzaleen, I guessed about seventy
yards off; I loaded a large duck-gun with big shot, fired at the lot and—apparently did
not touch one.”—(Davison, in Hume’s “‘ Game-birds,” I., p. 225.)

When watching a small flock of bronze-tailed peacock pheasants in Pahang, I saw
four Junglefowl go to roost across a deep, narrow mountain valley. Although within
easy shot they did not mind my presence, apparently considering that my only method
of attack was to descend and ascend the extremely deep and steep slopes. They had
chosen a stubby tree growing obliquely out of the cliff side and directly overhanging
the great depth beneath. Far down below the water could be heard, and high overhead
the wind was raging, but the four birds crept silently along branch after branch,
ascending slowly until they reached one which was protected by an overhanging mass
of foliage and vines, and here, after a prolonged scrutiny of me, they settled down close
together ; a family evidently, cock and hen and two nearly grown young birds. Twice
I have known of cocks roosting singly, night after night in the same place, but in both
cases the hen was sitting on her eggs near by. I do not think the cocks often crow
from their roosts before they descend in the morning. The several times that I have
surprised birds as they sent forth their challenge, it has been uttered either from the
ground, a fallen log or a low perch other than their regular roost. When they begin
their evening calling, they are usually upon the ground, and the crow is certainly not
uttered after they have retired to their roost.

The roosting-place most often chosen is near the top of a horizonally bent bamboo,
a position liable to sway wildly in a wind, and, one would think, offering but a poor
foothold at the best. It must, however, be as safe a place as could be selected, few
creatures being able to climb the thick, smooth butts of the stems. When an ordinary
tree is utilized, its position, either isolated in the jungle, or, as I have described, over-
hanging a deep chasm, aids in making the roosting-place less liable to direct attack
from creatures of the night. That most excellent observer, Mr. Osmaston, writes me
that ‘usually several birds roost fairly close together in one or in adjacent trees.” In
countries other than Garhwal, all I have seen were grouped in one tree, sitting as close
to one another on the same branch as they could crowd.

The only intimacies which I have observed in the life of these birds are with their
domestic kindred of the villages. In regard to this relationship I shall have more to say
farther on. With this exception, I have observed only the casual company of babblers,
who sometimes accompany the Junglefowl down a long slope on their way to water,
and are ever ready to sound the alarm when their keen eyes and more elevated position
have enabled them to detect one’s presence.

Their enemies vary, of course, with the locality, but members of the mink family, the
smaller cats (such as fre/is chaus), hawks and eagles are unquestionably the greatest
dangers with which these birds have to contend. In all my watching, I never observed
a tragedy, although more than once I saw individuals suddenly terrified by some menace

.

184 A MONOGRAPH OF THE PHEASANTS

concealed from my point of view. Mankind is probably the bird’s greatest enemy at
present, in spite of the fact that the great majority of the individuals which live in his
vicinity are becoming tainted with domestic blood.

The senses of sight and hearing are those which protect the Junglefowl from its
enemies, the former dominant, while the ears are yet so keenly attuned that the least
crack of a twig will often send the bird in headlong flight. Never have I seen a wild
bird off its guard for a moment, and although I have lain prone and had a cock come
within ten feet, yet it was only because I was perfectly hidden and motionless. On this
and on other occasions I have seen the bird under observation become suspicious, and
even finally take alarm when I was absolutely certain that through none of its five
senses had it received warning of my presence. There seemed to be an intuition, a
mental sensing of concealed danger, an indefinite conviction which gradually increased
in power and assumed control of the bird’s emotions, in spite of the fact that it had as
yet no knowledge of the location or character of the peril. In such case it was as likely
to make its escape by passing close to my concealed position as in the opposite
direction.

On the general character of the country, the lay of the land and the type of
vegetation, as well as the suddenness and degree of extremity of attack, the mode of
escape depends. When surprised by dogs or suddenly come upon by a man, especially
when a steep, open hillside or plain lies ahead, the birds rise at once and with
strong wing-beats swiftly skim down the slope. Where only a dog is concerned,
they take to the nearest trees, crane their necks down at their disturber and cackle
querulously and noisily. When taking to wing, no utterance other than a sudden
terrified cackle is given. But there is no hint of the weak muscles of the barnyard
degenerate, and I have told already how they have the power of mounting some
distance almost vertically to clear themselves of surrounding dense vegetation.

If conditions are right they are as ready to escape on foot, running with head
and tail low, with long, swift strides which take them almost instantly from view.
They are adepts at dodging, and in an almost open field I have seen a cock dash
into the shelter of a bush so quickly that the eye could hardly follow. Waiting a
favourable opportunity it would make a dash for another clump of foliage, and so
on, until the shelter of the jungle was reached. I have seen nearly grown birds,
which were not accompanied by their parents, squat, and once I think a solitary hen
bird adopted the same tactics until she made certain that I was approaching,
when she fled at full speed. I have never seen even an attempt at crouching in his
tracks on the part of a cock.

Of considerable interest is the fact recorded by several reliable observers that
the hen with chickens will act like a wounded bird, as in the case of the ruffed grouse
and many other species. Osmaston writes me: “The hen simulates a wounded
condition, whilst the young crouch motionless and are almost impossible to discover
among dead leaves.” This act seems best interpreted on the whole as an involuntary
convulsion of fear; since we can hardly credit the bird with even an instinctive mimetic
intention ; and certainly not with a conscious desire to reproduce the actions and postures
of an injured and dying bird, which she herself has very probably never had the
opportunity of witnessing. Natural selection, I think, can easily explain the

RED JUNGLEFOWL 185

development of this phenomenon, while at the same time I wish to give full credit to the
bird for sheer bravery and extreme parental devotion. For while the action in itself may
not be voluntary, yet when it has failed, and the bird, as I have seen in the case of
many species, realizes that the danger is becoming acute, and she returns and actually
buffets the assailant—this, it seems to me, rises to the level of voluntary initiative.
The spasm has passed, and the bird, with all the strength and power in her body, strives
to divert attack from her offspring.

I have never observed even a trace of this simulation of disability on the part
of domestic hens, so this must be a very recent acquisition in the Junglefowl, whereas
we all know of a hen’s conscious bravery in the face of attack by hawk or dog. In
domesticity this latter trait is evidently of value, while the other ruse, like the power of
strong flight, is unnecessary, and hence has degenerated and been lost.

We have seen that the distribution of the Red Junglefowl has been considerably
increased through the agency of man. Swinhoe has given us an account of his
_ finding the bird in the Island of Hainan.

“T fully believed that Hainan would yield a pheasant of some kind, and I never
ceased inquiring of the natives whether ‘ Hill-fowl’ (the usual expression for pheasants)
were found among their woods. The reply was in the affirmative, and I eagerly
looked out for them. Judge my disgust, then, when I discovered that the wonderful
‘Hill-fowl’ was only a wretched Junglecock. Though greatly disappointed, in heart
I was still pleased to make the acquaintance of a bird that I had not met before.
On the 18th of February I was rambling in the early morning at Shuy-wei-sze
(Central Hainan), and stepped through a hedge into a field, at the further end of
which was growing a patch of sweet potatoes. A rustic who was with me pulled me
by the sleeve and cried ‘Twa-kai’ (Hill-fowl). I turned, expecting of course to see
a pheasant. The clodhopper was pointing at an ordinary-looking rooster standing in
the middle of the field with body erect and tail decumbent. Seeing me take no
notice, the native cried out again, ‘Shoot, it is a wild bird, and not a barn-door
fowl.’ I looked again and saw the bird moving away behind a hedge. When I got
in view of him again, he ran for the hedge with all speed. There was no waddle
in his gait, and I then saw that it was a wild bird. I fired and ran up. The
wounded bird flapped and tumbled about precisely as a cock does when its neck is
broken and it is thrown down to die. My heart misgave me; I thought I had shot
some poor peasant’s fowl; but I was wrong—every one I met told me that it was a
genuine wild fowl, and on carefully examining it I found it to be so. Unfortunately
neither its hackles nor its tail were fully developed. Its comb was small, and its
spurs wart-like. Its iris was orange-yellow. Skin under the ear cream-white, purplish
pink at its forward corner. Bill ochreous, brown on culmen. Legs brownish-grey,
tinged with purple. Its. intestines measured 35 feet; and its stomach contained
sweet potatoes, vetches, and stone-grits. Its flesh was juicy and delicious eating, and
quite different in flavour from that of the Barn-door. Its festes were of an enormous
size; so that there can be no doubt the bird was breeding. On the 27th of February
I visited the same field again; and on the same spot where I first saw the cock,
appeared the hen. Several natives were with me. They shouted, ‘H6 ho Taw-kai’
(look, look! Hill-bird); and at the noise the hen rushed into the hedge, and we tried

VOL, II BB

186 A MONOGRAPH OF THE PHEASANTS

to beat her out in vain. She seemed to be of a deep brown colour, and in running
kept her head low and her tail partly erect.

“At Lingshuy (S.E. Hainan) we found grave-mounds on the edge of the jungle
strewn with cocks’ feathers, as if the wild fowl were in the habit of meeting on the
mounds to fight. At Yu-lin-kan (S. Hainan) I heard them repeatedly chuckling in
the jungle quite close to me; but there was no getting a shot at them. In the dense
woods about Nychow (S. Hainan) they were particularly common, and we heard and
saw them often. When put up in the open, they make at once for the covert, flying
heavily, with the body and tail nearly perpendicular. I saw a Le man put a cock
bird up; and marking it drop into the wood, I hastened to the spot. It gave a
crow, ‘Tok-tok tok tok chea’—as a domestic hen does when frightened. My follower
raised it from the thick bush with a stone; it flew a short distance, and fell again
into the thicket. Our party returned to the boat without a Junglefowl, and we saw
no more of them in the course of our cruise.”

HOME LIFE

The breeding season of the Red Junglefowl in the northern sub-Himalayan part
of their range extends from February, when the males begin their challenging, combats
and courtship, until the end of May, when the few hardy individuals which make
their home well up in the hills, deposit their eggs. In central Burma eggs have been
taken from March to June. As we follow the birds southward throughout their
range, there is, as usual among birds in general, an increasing laxity in the precise
limits of the nesting months, and in the Malay Peninsula young birds have been
recorded from February to late August.

In the first few pages of this monograph I have mentioned the courtship of the
wild Red Junglefowl. Even in the degenerate domesticated rooster we still see hints
of this feral chivalry, but as with flight and many habits of life this centuries-old
inmate of our barn-yards has let his manners become sadly slipshod. Too often his
procedure is to sidle up to a hen, and make only one or two half-hearted circlings,
with wing awkwardly drooped, an atavistic effort to reveal beauties of plumage which
are no longer his. Then if the hen, obeying her innate impulse, shows but slight
interest, his patience is at an end, and the courtship degenerates into a mere rough-
and-tumble pursuit and capture by sheer force. We see much the same thing in
the house sparrow.

One noteworthy thing about the courtship of the domestic cock is the rattling
croak or harsh crooning sound he often makes when going through the display. The
wild birds are absolutely silent when they are courting, uttering a cluck now and
then between displays, but giving no hint of the rather coarse garrulity which
characterizes their captive relation.

The little definite information we have of wild birds shows that Junglefowl differ
in no way from pheasants in general in the reliance they place in persistent showing
off to produce some effect on the hens. Hypnotic rather than sentimental though
this probably is, yet it is an effect gained solely through emotional channels. The

RED JUNGLEFOWL 187

manoeuvres themselves are not of the highest order, corresponding to the first stage
—the lateral display—of such birds as peacock pheasants and tragopans.

The assertion has been often made and with some truth that some of the court-
ship attitudes are similar to those assumed in fighting, the intention of the assertion
being to belittle any specialized effort on the part of the male to catch directly the
eye of the hen. This statement is now quoted and repeated in the most sweeping
manner by every dilettante writer on birds, until it requires some refutation to bring
it within logical bounds. The lateral display of the Junglefowl has nothing to do
with the fighting position of these birds, which is essentially frontal. Even when
two cocks are approaching one another and circling for an opening, the resemblance
between the two motor phenomena is extremely remote.

The nodes of brilliant plumage in the wild Junglecock are on the back, wings
and tail. Hence it is these areas which are of prime importance in courtship display.
The dull under-parts are concealed. The bird endeavours to manceuvre into a
position almost in front of the hen, and then to render himself as asymmetrical as
possible. The inner wing and tail is spread and lowered almost or quite to the
ground. The back is slanted and the inner side depressed as much as possible.
The hackles are raised and spread a little, hanging far down on the side toward the
hen. When the extreme posture is assumed, the bird takes short, rapid steps in a
half circle, bringing up on the opposite side. After a few seconds of quiet, he may
retrace his steps and make a return circle, displaying the beauties of the opposite
side, or, if the hen has advanced, he may walk quietly around behind her and display
again from the same side. In any case we are reminded strongly of the display of
the golden pheasant. I have seen this courtship on the part of a wild bird, when
it had no inkling of my presence, and although I have watched many domestic cocks
going through the same performance, it has, even at its best, lacked the patience,
the thoroughness and the vigour of the courtship of the wild bird.

Junglefowl are notoriously pugnacious. This is attested by the great length of
spurs and indirectly by the hackles, which appear to function especially as a protective
ruff or a false shield. The beak-to-beak position of the fighting birds is well known to
every one, and at such a time the hackles are erected until they form a perfect halo about
the head, hiding from view all the vulnerable parts of the body.

Hume has an interesting note about the combats and fighting arenas of wild Jungle-
cocks. He says, ‘‘ No one especially notices the extreme pugnacity of these birds in the
wild state, or the fact that where they are numerous they select regular fighting-grounds
much like ruffs.

“Going through the forests of the Siwaliks, in the north-eastern portion of the
Saharanpur district, I chanced one afternoon, late in March, on a tiny open grassy knoll,
perhaps ten yards in diameter and a yard in height. It was covered with close turf,
scratched in many places into holes, and covered over with Junglefowl feathers to such
an extent that I thought some Bonelli’s eagle, a great enemy of this species, must have
caught and devoured one. Whilst I was looking round, one of my dogs brought me
from somewhere in the jungle round a freshly-killed Junglecock, in splendid plumage,
but with the base of the skull on one side pierced by what I at once concluded must
have been the spur of another cock. I put up for the day at a Bunjara Perow, some two

188 A MONOGRAPH OF THE PHEASANTS

miles distant, and on speaking to the men, found that they knew the place well, and one
of them said that he had repeatedly watched the cocks fighting there, and that he would
take me to a tree close by whence I could see it for myself. Long before daylight he
guided me to the tree, telling me to climb to the fourth fork, whence, quite concealed,
I could look down on the mound. When I got up it was too dark to see anything, but
a glimmer of dawn soon stole into the eastern sky, which I faced; soon after crowing
began all round, then I made out the mound dimly, perhaps thirty yards from the base
of the tree, and forty from my perch; then it got quite light, and a few minutes later
a Junglecock ran out on to the top of the mound and crowed (for a wild bird) vocifer-
ously, clapping his wings, and strutting round and round, with his tail raised almost
like a domestic fowl.

“And here I should notice that although, as has often been noticed, the wild cocks
always droop their tails when running away or feeding—in fact almost whenever you see
them—yet I believe, from what I then and once subsequently saw, that, when challenging
rivals, they probably always erect the tail, and I know (having twice so surprised them
before they saw me, when watching for cheetul and sambhur from a machdu, near water
in the early morning) that when paying their addresses to their mates they do the same
during the preliminary struts round them.

“TI learnt so much and no more; there was a rush, a yelp; the Junglecock had
vanished, and I found that one of my wretched dogs had got loose, tracked me, and was
now careering wildly about the foot of the tree.

“Next day I tried again, but without success. I suppose the birds about had been
too much scared by the dog, and I had to leave the place without seeing a fight there ;
but putting all the facts together, I have not the smallest doubt that this was a real
fighting arena, and that, as the Bunjara averred, many of the innumerable cocks in the
neighbourhood did systematically fight there.

“Only a week later I shot two cocks, who were tumbling head over heels, a
confused mass, with wings and legs interlaced in an incredible manner, and on several
other occasions, when watching and waiting, concealed and in silence, for larger game, I
had witnessed desperate battles between cocks who Hepp to meet, attracted by each
other’s crows and flapping of wings, near my tree ambush.”

I have never been fortunate enough to see two wild cocks fanned but I have
three separate times chanced upon what appeared to be fighting-places. In all cases
these were in small glades in bamboo jungle, and in one instance a half-decomposed
cock bird was lying in the centre. On two other occasions I have found where a Jungle-
fowl had been plucked and doubtless eaten by an eagle or owl, and there was no doubt
as to the distinction between the two situations. The widespread abundance of hackles
and body feathers, the tracks, the sign, all indicated a much-frequented arena. My
attention was attracted to two of the three places by the constant crowing in early
morning, and this crowing did not die out little by little, with longer and longer
intervals between the challenges, but suddenly, with suggestive abruptness, as might
occur if a rival appeared on the scene. I believe that the combatants in these places
are chiefly young or unmated birds, and that hens are to be found somewhere in the
vicinity, although my belief in this is based chiefly on my observations of other
species of Junglefowl. The mated birds keep together and do not range far afield,

RED JUNGLEFOWL 189

and in their case combats are probably confined to encounters with trespassing strange
males.

A fact of passing interest, which I shall touch upon in more detail elsewhere, is the
isolation and accentuation by breeders of the mental character of combativeness. The
mentality of the domestic game-cock is as much a product of artificial selection as is the
physical character of a crest in the Polish fowl and the physiological function of increased
fertility.

In spite of many exceptions, I believe that the normal, or at least the much more
common state of mated life among Red Junglefowl is that of monogamy. Birds living
in open jungle, isolated from the fowls of natives, are almost invariably seen in pairs,
except when families are flocked together, and several sportsmen who have hunted this
bird emphasize this fact. In more than one instance where I have found a single cock
associated with several hens, it has been in the immediate vicinity of villages, and I
credit this apparent polygamy to the influence of native fowl blood. Among domestic
birds polygamy is, of course, the rule. Just as the power of flight and speed has
degenerated in our barnyard fowls, so the normal pairing relation has been upset, and
mating is indiscriminate and all but lacking in courtship.

The site chosen for the nest is again, to a certain extent, an index of the absolutely
feral nature of each respective pair of Junglefowls, or reveals the demoralizing effect of
crossing with native birds. Twice in widely separated countries I have discovered,
with the aid of expert native hunters, far from human habitation, the nests of wild hens.
They were hidden deep in thickets, and protected by a very dense growth of bamboo in
one case, and brake in the other. There would have been no possibility of discovery
except by the accidental flushing of the hen. In four instances of nests near villages, on
the other hand, none were thoroughly concealed, and in two cases it was impossible to
decide whether or not the nests were those of extremely wild native hens or rather tame
feral jungle birds.

The nest is, of course, on the ground, usually well toward the centre or denser
portion of a thicket, usually of bamboo. It is variable as to character; sometimes
merely a hollow scratched out of the earth, without any lining; again the leaves which
were already on the ground remain, and are matted down by the eggs and pressure of
the bird’s body. More rarely a considerable heap of grass and leaves is collected,
forming a real nest, unlike the case of most of the pheasants. Another variation which
has been described is where the bird, beside making a nesting hollow, also scrapes up
the earth outside and around this nest, thus forming an appreciable rim, which protects
the contents, keeping the lining firm and the eggs secure.

Hume speaks of finding a Junglefowl’s nest almost every day during a month’s
shooting in May along the southern slopes of the Siwaliks, with the aid of dogs and
men. Once six were found near the Bhing-ka-khol within a radius of two hundred
yards. ;
The time of year of laying and the altitudinal range within which breeding takes
place have already been discussed. The number of eggs is not very constant, but five
to eight is the usual number. Eleven have been known to be deposited by a single hen
at one laying, but the record of a nestful of fourteen probably represents the joint efforts
of two birds.

190 A MONOGRAPH OF THE PHEASANTS

In appearance the eggs are, of course, like niniature hens’ eggs. The shell is smooth
and fine of texture, with considerable gloss. Near native villages the shells partake of
the character of those of the domestic birds. Usually they are of a pale buff or yellowish
café au lait colour, sometimes paler, whiter, or again darker. In length they vary from
4o to 53 mm.; and in breadth from 32 to 39. The average Red Junglefowl egg
measures 46 by 35 mm.

The period of incubation of Junglefowl is the same as in their domestic descendants,
about twenty-one days, and the chicks are strong and very wild, able to leave the nest,
to follow the parent and to hide at her command almost as soon as their down is dry.
When the eggs of really wild birds are hatched under domestic hens, and the chicks are
confined with other fowls, they will accept captivity in the end, and will remain and
breed. But they always choose to roost in trees adjacent to or within the village, rather
than spend the nights in coops, or on the rafters, or under the floor-beams of the
Burmese or Malayan houses. If, however, the chicks are permitted at once to run in the
jungle, they will drift away from the hen, and through fear will one by one be lost.
Nevertheless, I repeat that it is most interesting that the chicks, or even the adults of
real feral Red Junglefowl, unlike the other closely related species of Gallus, are sus-
ceptible of what is practically domestication in one generation, unlike any other member
of the pheasant family. In this it parallels the mallard duck, and to a less extent certain
South American birds, such as the trumpeters, cariamas, and chachalacas. ‘These latter,
however, refuse to breed in captivity, but within the space of their own life-time become
perfectly tame, and accept man as protector and friend while enjoying full liberty at the
very edge of their native jungle.

The call-notes of the chicks and the warning and calls of the hens differ but little
from the corresponding utterances of domestic birds. The cock, as far as I know, never
goes near the nest, nor does he take any part in incubation, although he is frequently
seen in association with the hen when she goes off to feed. When the brood is hatched,
or at least when the chicks are a week or more old, the cock is found with them,
scratching and helping to find food. This I have verified personally. When alarmed,
however, he is the first to dash away, carrying out the rule of pheasants in general, that
susceptibility to suspicion, fear, and flight is in direct relation to the excess of colour or
conspicuousness of plumage. When threatened with danger the hen leaves her brood
with a single cackle, sharp and incisive, which sends every chick to cover, where they
remain motionless until they hear her reassuring notes. I once was fortunate enough
to have a hen and her brood of four young pass close beneath the tree in which I was
concealed on the watch for fireback pheasants. Some animal of which I could not catch a
glimpse rushed suddenly through the ferns near by, and the hen rose and whirred
swiftly into the jungle in the opposite direction. By my watch, over twelve minutes
passed before the bird reappeared, although after the first rush there was no further hint
of danger. Then she came silently, cautiously, picking her way back through the under-
brush, strangely enough from the side whence the sound had come, having described a
hemicircle in her flight and return. When a few feet away she uttered several low
crooning notes, walking slowly, and before she disappeared I made out the forms of
several of the little chicks. Where they had been or when they answered her reassuring
call, I could not tell. Their markings and colours were perfectly protective; when

RED JUNGLEFOWL 191

motionless they were bits of the jungle floor mosaic of various pigments, lights and
shadows, and with no tell-tale scents to betray them, were safe from all discovery during
the absence of the mother.

In the crop of a two weeks’ chick I found a bit of a scarlet earthworm, several small
grubs, anda piece of grass blade. Beyond this I have no data as to the food of the
young birds. Judging from the scratching proclivities of both sexes, I believe that
insects form an important part of their food. As I know that the cocks share in the
defence and feeding of the brood, it is reasonable to suppose that he roosts with them
from the time when they are first able to flutter up toa limb. Certainly when they are
half grown he is found with them or on a neighbouring branch every night. The
dangers to which the eggs and very young chicks are subject are those which ever
threaten ground-nesting birds; snakes, lizards, small carnivores, and other predaceous
mammals, and the small, jungle-frequenting hawks and owls being the worst.

RELATION TO MAN

The details of the past and present relation which obtains between the Red
Junglefowl through its descendants and mankind, could easily be elaborated into an
entire volume of this monograph. In economic importance the domestic fowl takes
rank with the cow, sheep and pig. Many accounts have been written on this subject,
one of the best still being Darwin’s chapter published more than forty-five years ago.

Sometime in the far distant past, exactly when we shall probably never know, some
primitive race of mankind captured living Junglefowl and tamed them. The birds
learned that intimate association with man ensured comparative freedom from attack
by beasts of the jungle and a more or less regular supply of food. Certainly their lives
are a great deal longer than among their wild kindred. There are records of domestic
cocks twenty-four, twenty-five and even thirty years old. So in numberless races and
varieties we find the domestic fowl as the companion of man distributed over the whole
world, with the exception of the high mountains and the polar regions, where it is
infertile, and, therefore, cannot be bred. It everywhere betrays its southern origin by
its love for warm, sunny places, its dislike of cold, and the infertility in chilly regions
above mentioned. The early history of mankind and the fowl, as far as we know it,
I have detailed elsewhere.

After the first domestication, and throughout succeeding years, during generation
after generation, these tamed fowls of the jungle made three separate appeals to man.
First, he reared them and found their flesh and eggs goodly food, and little by little
learned to breed birds which either were large in size, and thus produced an unusual
amount of flesh, or by selection, increased feeding, and in other ways brought about
an enormously increased egg productivity. In the second place the natural pugnacity
of these birds appealed to the primitive instincts of savage man, and many generations
of him must have given much thought to the sport of cock-fighting to have brought
into existence the specialized race of game-cocks with its peculiar structure and
abnormal pugnacity. Thirdly, when the plasticity of form and structure of Junglefowl
bred in captivity became evident through the variation of successive generations, the
aesthetic side of mankind seized upon this as a sort of living, organic potter’s clay, and

192 A MONOGRAPH OF THE PHEASANTS

with this material, breeders and fanciers from early in human history to the latest
exhibitors at our modern poultry shows have exercised their ingenuity in the production ©
of beautiful, bizarre or monstrous races of domestic fowls. All this is of interest to
us only as showing the possibilities inherent in a Species of wild bird when it has
become adapted to domestication. Emulation and competition have become so great
that in England alone, twenty poultry shows are sometimes held within the space of
a week, and prize fowls may fetch at auction from five hundred to seven hundred and
fifty dollars.

The plasticity of domestic fowls is very remarkable, and has furnished the
mutationists with considerable evidence. The unnatural conditions of their lives,
however, doubtless accounts for much of this, and in Nature, for example, among feral
Junglefowl there is unquestionably much less of this abnormal variation. Davenport
says (chapter on Mutation in ‘“ Fifty Years of Darwinism,” 1909, p. 171) that within
a period of four years ‘in the egg, unhatched, I have obtained Siamese twins, anteriorly
duplex individuals with shortened upper jaw . . . and chicks with thigh-bones absent.
There have been reared chicks with toes grown together by a web, without toenail, or
with two toenails on one toe; with five toes, six toes, seven toes, or three toes; with
one wing lacking or both absent; with two pairs of spurs; without oil-gland or tail
(though from tailed ancestry); with neck nearly devoid of feathers; with cerebral hernia
and a great crest; with feather-shaft curved; with barbs twisted and dicotomously
branched, or lacking altogether. Of the comb alone I have a score of forms: single,
double, triple, quintuple and walnut, V-shaped, cup-shaped, comprising two horns, or
four or six, absent posteriorly, absent anteriorly, and absent altogether.” And in
addition to the increase in egg production, of which we shall speak shortly, we find eggs
varying much in colour, and of all sizes and shapes, from sausage-like to a complete
horseshoe. Some of these abnormalities are very patently, wholly inimical to the life
of the organism, but others have been developed and increased by selection until they
have come to be characteristic of races and breeds known over all the world.

Game fowls are more like wild Red Junglefowl than any other domestic race,
differing chiefly in being larger and in carrying the tail more erect. When these birds
were bred for fighting only, the general proportions of the wild Junglefowl were not
departed from, but in recent years, stimulated by the desire to excel in competitive
poultry shows, breeders have wrought remarkable changes in this race, especially as to
extreme length of neck and legs. Within a comparatively short time birds had been
evolved with the breast over a foot from the ground. The sickle feathers of the tail
have been shortened, and the entire plumage rendered firm and closely pressed to the
body. These abnormal proportions, of course, unfit the birds for fighting. On the
other hand, as in many other races, one strain of the original type of game bird has been
reduced in size through selective breeding, and game bantams of exceedingly small size
have been produced. In this race as a whole there is a curious tendency towards the
assumption of female plumage by the cock, and such breeds are known as “‘henny” game.

Years ago Temminck gave the name Gad/us giganteus to a fowl of great size from
the Malay States. This is now well known as a race appropriately called Malays, which
are much larger than the game, and with very long and heavy legs. In fact so
disproportionately long are these limbs that the bird often rests its entire tarsus upon

RED JUNGLEFOWL 193

the ground, looking like a cassowary, or, with the tail-feathers trailing, actually like a
kangaroo. The plumage is so scanty that,on the neck, breast and legs the skin shows
through in many places as patches of bright red, this colour being apparently the effect
of its direct exposure to light and air. These birds are excellent for table use, having
an abundance of fine flavoured flesh, especially on the breast. In the aseel, the
pugnacity of the cocks has been handed on, not only to the hens, but even to the chicks,
who fight with one another at every opportunity.

The birds known as Cochin China fowls, including brahmas and langshans, have
been evolved along lines of great size and weight. Their name is a misnomer, as they
originally came from Shanghai. The cocks sometimes reach sixteen or seventeen
pounds. The plumage is soft and downy, and the feathering extends down to the toes,
the wings and tail being so small as to be wholly functionless. In fact the feathering
on the legs and feet has been carried to such an extreme that the normal scalation has
given place to almost wing-like masses of quills, six and even eight inches in length.
Brahmas did not come from the vicinity of that river, but were produced in the United
States, by crossing a variety of Malay fowl with Cochins. By crossing the latter with
Dominiques the Plymouth Rock was produced, a very prolific layer.

Spanish fowls have giveh off leghorns, Andalusians, Minorcas and others. In the
black Spanish, the originally small spot of white on the ear lappet has spread over the
entire face, and the lappet itself increased in size until it is sometimes seven inches in
length. and four in width. These birds and their subraces have been bred for egg
production, and correlated with this the incubating instinct has been lost. In the place
of the nestful of four to eight eggs which their feral ancestor laid once, or at most twice

‘during the season, these domestic egg machines will lay well over two hundred eggs
in a year. In Australia, four hundred and fifty hens laid over eighty thousand, nine
hundred and fifty eggs, while in America there is a record of a single yard of six
hundred fowls which averaged in the year one hundred and ninety-six large, perfect
eggs for each bird!

The falsely named Hamburghs and the crested fowls are also non-incubating,
abnormal producers of eggs. The combs of the former are double. In the latter, on
the contrary, the great development of the crest has almost, and in some cases wholly
obliterated the comb. The wattles too are absent in some forms, their places being
taken by tufts of feathers. Almost the only changes produced by man in the
domesticated races of fowl are those of general proportion and colour. But in the
crested fowl the cranium itself has undergone a remarkable change, the front part of
the skull forming a prominent rounded swelling, which contains a large part of the
brain. Only occasionally is this remarkable structure wholly ossified.

The Dorking fowl has an extra toe on each foot. This is probably a monstrosity
and not a reversion to the real pentadactyl foot of the ancestors of birds. In one of
the latest contributions (Ghigi, ‘Sul Significato Morpologico della Polidattilia nei
Gallinacei,” 1901) to this subject, the author’s summary is as follows: ‘Summing up,
it seems to me conclusive that the extra toe of chickens, homologous with that
sometimes observed in other birds, must not be attributed to a doubling of the first
toe, but to the teralogical development of an organ (fvea//uce) which in the ancestors

existed in a rudimentary state.”
VOL, II” . GG

194 . - A MONOGRAPH OF THE PHEASANTS

Sebright bantams are in some ways the most artificial of all domestic breeds, for
the cock has completely assumed the characters of the hen. The markings of the
plumage, the short, pointed hackles and the short, stiff, central tail-feathers, show an
extreme specialization and departure from the ancestral conditions. In the silky fowls,
barbicels are almost absent from the feathers, giving the plumage a loose, silky or
hair-like texture. In the best-known breed of this race the feathers are pure white,
while the skin, the connective tissue separating the muscles and the periosteum covering
the bones, are a deep blue-black. When one has travelled in Malaysia and purchased
these birds for the first time for the table, the sight of the black tissue on their flesh
is most startling. They are nevertheless delicious eating. This character is not
appreciated by the general purchaser at home, so these birds have little value for the
market. In fact even in crosses with other races, while the silky character of the feather
is soon lost, the dark skin and other tissue is dominant and will do much to depreciate
the selling value of the crosses.

Frizzled fowl, or ‘ pine-apple hens” as they are called in China, show the character
of every feather, even the flight feathers and tail, turning outward from the body. One
sees them everywhere in the East, especially in India, and indeed they are adapted only
to a warm, dry climate, owing to the lack of protection which their plumage affords, the
rain and cold easily reaching their skin. A radical, inheritable alteration of structure
is found in the rumpless fowls, in which the coccygeal vertebrae are absent. This
character is dominant, so that by crossing, any race of rumpless fowls may be produced.
Dumpies or creepers are fowls with legs so short that they can progress only with
difficulty. Bantam is a term applied to very diminutive fowls of any race. Even
Cochins have been bred down to less than a pound weight. The Japanese have a
bantam with enormous tail and comb, very prominent breast and absurdly short
legs.

While shape is an easy factor to alter in the breeding of fowls, segregation of
colour, unlike the condition existing in doves, offers much greater difficulty. The
Japanese bantam is an unusual exception to the possibility of this localization of
colour—the bird in general being white, while the tail is black. Darwin accounts for
this by what he calls analogous variation, finding its explanation in the patterns and
colours, not only of the direct ancestor, but of other members of the genus and related
groups. Thus although the wild rock dove has not a white head, other species of
doves have, and this character is easy to obtain by selective breeding. A white head
is unknown among any species of wild Junglefowl or of pheasants, and a white-headed
domestic race has never to my knowledge been obtained.

The long-tailed fowls of Japan are popularly known as Yokohama, or Shinowara or
Phoenix Fowls. Much has been written about this breed, and in this literature there is
more of error than of truth. Authors have taken as their theses the supposed secret of
production of the long tail-feathers by constant stroking and gentle pulling, stimulating
in this way the abnormal growth. When one visits the home of these birds in Japan,
it is found that such methods exist only in the ingenuity of their Caucasian inventors.
As a matter of fact there is no secret connected with these birds, and their great length
of feathers is as normal a product of development as the crests of other breeds. It has
been proved that constant gentle pulling and stroking of the feathers has a certain effect

PHOTOGRAVURE 32

JAPANESE LONG-TAILED FOWL

Or all curious breeds of domestic fowl, none are more remarkable than the Japanese birds, whose
upper tail-coverts sometimes reach a length of over twenty feet. The feathers are kept wrapped in soft
paper, and the birds themselves spend a most uninteresting life in a high, narrow box, fed by hand and

cared for by special attendants.

PHOTOGRAVURE PLATE 32

“TMOF CHTIVI-ONO

r

NVdAV EL

RED JUNGLEFOWL 195

in making the feather grow somewhat more rapidly, but this is only an unimportant
indirect stimulation, not used by the Japanese. The finest specimens come from
Shinowara, a small village three miles from Kochi, in the Province of Tosa on the
Island of Shikoku. Prof. Ijima of Tokyo University told me that tradition in regard to
these fowls goes back many hundred years, but their origin is wrapped in unsolved
mystery. It is said that the breed was known in Corea before a.p. 1000. In former
years the Japanese breeders were encouraged by the Daimyo or Governor of the
Province to produce birds with especially long feathers, as these were of importance in
heraldry and also used as spear decorations by high officials. Recently, however, owing
to lack of government support and popular interest, the industry is on the wane, as but
few purchasers are to be found. At present, a good pair of birds with the cock’s feathers
six or eight feet long may be purchased for two hundred yen; one hundred dollars in
our currency.

For the first two years the birds are kept like ordinary fowls, but after this time the

cocks are confined in a narrow box-like cage, on a perch from which they cannot descend
nor can they turn around on it. From this time there appears to be no moult of the
tail-coverts and central rectrices, and within from four to six years these feathers have
attained an enormous length. Cocks are used for breeding only six or seven times, and
are then valueless, as their long feathers drop out. This irregularity of moult is the
explanation of the phenomenon.
_ The lower rump feathers sometimes reach a length of three or four feet, while fifteen
or twenty of the upper tail-coverts themselves attain a much greater development. The
roots of these feathers become correspondingly thicker and stronger as the weight
increases. The central sickle-shaped rectrices attain the greatest length, but the
remainder of the tail itself is of normal length.

The box in which the bird is confined varies in height according to the length of
the feathers. As these increase in length they are carefully wrapped in mulberry paper,
thus protecting them from dirt and from any danger of being abraded.- The birds are
fed with great care on nourishing food, and it is said that under the best treatment
a growth of six inches a month has been obtained. The longest recorded feather
measured twenty feet and two inches. The bird remains contentedly enough on its
perch until taken out for exercise, which, when it approaches relative perfection, is done
only every other day. The fowls are very tame, and stalk solemnly about, an attendant
holding up their enormous train until they are replaced in their exhibition box. Once
or twice a month the bird, together with its tail-feathers, is carefully washed with warm
water. I could not learn definitely of its diet, but unhusked rice, cabbage, and other
vegetables are given and all the water which the bird will drink. The birds are white
and of various colours, but usually of the game type. The hens are very ordinary-
looking, and lay about twenty-five or thirty eggs each year. Inbreeding has made this
race of fowls exceedingly delicate, and birds under a year old cannot be transported with
safety. Second year birds are the best for purchasing, as-in older specimens the
lengthened feathers render shipment most difficult. The fertility is very low, three
fertile eggs out of twenty being considered an excellent average. But within the last
few years crossing with black minorcas and white leghorns has been attempted, and the
non-moult of the posterior dorsal feathers is found to be a dominant factor in the hybrids,

196 A MONOGRAPH OF THE PHEASANTS

while the evils of inbreeding are banished. One sees traces of long-tailed-fowl blood
even in the fowls of Kagoshima, where I saw many cocks walking about whose
tail-coverts were so long as to drag for several inches on the ground.

In the Malay States most of the aborigines, even the small nomadic tribes, keep
domestic fowls, which they breed and eat. They also occasionally have captive pheasants,
but do not eat these, but keep them for barter with the Malays. Any wild Junglefowl,
however, which they happen to shoot or snare they consider good food. The usual
method of trapping of these natives is to place numerous nooses woven of hair or
vegetable fibre on the ground and bait the place with grain.

Another way which I observed at Kuala Tembeling is by means of an ayam déndk
or decoy bird, usually a hybrid cock or hen tethered near the traps or snares. This same
method is used elsewhere in Assam, Siam and the Sundarbans. An open space near
the forest is usually selected, the decoy bird tied to a stake and surrounded with open
nooses and scattered grain. The challenge of the tame bird, which is trained to call often
and loudly, attracts the wild cocks, who rush up to give battle, when they are almost
certain to become entangled. Or if a hen approaches, she is attracted by the grain and
also falls a victim. In Celebes the decoy fowl is called wawansa/ in the language of the
Alfaros, and to catch a Junglefowl in this manner is called mawansal. The natives of
this island do not attempt to tame the wild-caught birds, but use them only as food.

Still another variation of the decoy trapping is practised in Sulu, where the domestic
bird is tethered in the centre of a circle of spring nooses. Many wild birds are thus
captured, but only cocks. Even where grain is added to the charm of the calling bird,
the females remain aloof.

In northern Burma the bamboo fence traps, with their scores of deadfalls, account
for many hundreds of Junglefowl in the course of a year. In other places I have seen
separate nooses set for these birds, each attached to a bent sapling and baited with
grain.

Besides using decoy birds the Malays imitate the crowing and the flapping of the
wings of the Junglecocks, and they taught me so that several times I was able to
inveigle the birds close to me by making them think that a challenging bird was
concealed in my cover.

It is difficult to say whether Junglefowl in general are decreasing or are at least
holding their own. If we consider only the isolated, really pure strains, they are
unquestionably becoming fewer in numbers, but those which haunt the vicinity of
villages and cultivated districts, although shot frequently and suffering from the many
enemies, such as snakes and small carnivores, which make life a burden for the village
fowl in India, yet are constantly gaining recruits from the ranks of the domestic birds.

In the reserved forests in the sub-Himalayan region the birds seem to be fully
holding their own, as there is little shooting there and the native foresters have little
time for snaring and trapping. In these Government Reserves the birds are protected
from all shooting from the first day of March to the middle of September. The most
common time for Junglefowl drives either on foot or from elephants is from November 1
to the end of February.

To the southward, in the Central Provinces of India, the close season for this
species is from March 1 to November 30.

RED JUNGLEFOWL 197

JUNGLEFOWL AS GAME-BIRDS

The ornithologist is indeed rash who ventures to tell in his own words the delight
which the sportsman takes in his pursuit of game. Let him, unhampered by scientific
fetters, relate what pleasure the wild Red Junglefowl can give to the huntsman, whether
he pits his own woodcraft and ability singly against these wary birds or fills his gamebag
more easily by the assistance of dogs or a line of native beaters.

“When detached clumps of jungle or small hills occur in a jungly district where
these fowls abound, very pretty shooting can be had by driving them by means of dogs
and beaters; and in travelling through a forest country, many will always be found
near the roads, to which they resort to pick up grain from the droppings of cattle, etc. ;
dogs will often put them up, when they at once fly on to the nearest trees. Young
birds, if kept for a few days, are very excellent eating, having a considerable game
flavour.’—(Beavan).

‘Sometimes when thus beating for Junglefowl you meet with odd surprises. It
was in April 1853, in the good old days of palki dak, from Meerut to Mussooree. Three
nights we used to make of it when ladies were of the party, and the close of the second
night brought us to the Kheree Dak Bungalow, in broken jungly ground just south
of the Siwaliks. After breakfast I went out to look for Junglefowl, luckily with a rifle
(a heavy 2 oz. band spherical ball) in case of seeing cheetul. We beat a lot of low
jungle grass and scattered bushes, and I had got a partridge and a Junglehen, when I
turned into a very likely-looking nalla, about eighty feet deep, with sloping, well-grassed
sides, and at the bottom a narrow, perpendicular-sided water channel about four feet
deep and three feet wide, cut through the boulder clay. In this channel I walked, with
one or two men along the slopes on either side, and one or two above, all a little behind
me. Suddenly there was a shout on my left, and instantly a tremendous grunting. As I
seized my rifle from the shikari behind me, four black heads showed through the grass
immediately above me. I could not get out of the wretched water-course, which was
nearly up to my armpits, and without one second’s hesitation one of the bears (the old
female as it proved) came down upon me like a thunderbolt. I got my first barrel off
when she was about ten yards from me; the second let itself off as her chest struck the
muzzle, and then I was knocked over, half-stunned and nearly crushed to death. I
don’t know exactly how it all happened, but I found myself on my face, hardly able to
breathe; my head, arms, and body pinned down by the massive, motionless (lucky for
me) corpse of lady Bruin. Seeing that the bear was quite dead, my shikari and a good
pahari bearer I had soon pulled her off and released me, a mass of blood, a good deal
cut and bruised, but not really hurt; my first bullet had gone straight through her from
stem to stern (2 oz. hardened bullet and six drams of powder), the other had gone right
through the heart and come out behind the ribs on the left side.

“Tt will be well for griffs (as I then was) to bear in mind that, in the sub-
Himalayan ranges, at any rate, where Junglefowl are common, there bears and tigers are
not unlikely to be met with, and that they should never beat for Junglefowl in such
situations on foot without a rifle in trustworthy hands behind them, and never allow
themselves to be caught in such a trap as that in which I had stupidly placed myself.

‘‘In the autumn, after the millet-fields have ripened, they grow very fat on this

198 A MONOGRAPH OF THE PHEASANTS

grain, and the birds of the year are then vea//y good eating, but, as a rule, the birds one
shoots (be it confessed with shame, for it owght to be a close season) from March to June,
tiger-shooting in the Terai, when, the day’s sport over, one turns homeward towards the
tents, are no whit better than ordinary village fowls.” (Hume).

“The best shooting I ever got at this species was at Jalpaiguri, where the nallas, or
beds of stream, in the neighbourhood, which are common in that country, and filled with
jungle, gave cover to very many of these birds. When put up by beaters they fly out at
a considerable pace, and require a good knock-down blow to bag them. They run, too,
a great deal. In the Manbhoom district the native shikaris used to get many of them
by placing corn near some water in the half-dried-up beds of streams, and then
shooting them when they came there both in the early morning and evening to eat and
drink.” —(Beavan).

“This bird must be sought in all jungly country which is partly cultivated; and
where paddy-fields extend in long strips into the forest, two sportsmen walking on each
side just within the cover, with a line of beaters between them, can enjoy very pretty
shooting. The fowls rise from the stubble and fly into the wood, passing overhead, and
the sport resembles pheasant-shooting in England, the flight and size of the birds being
pretty similar. When the fields have been cleared of the fowls, the shooting may be
continued with success in the woods, if they be pretty open, and the sportsman furnished
with spaniels, the sight of which forces the birds to tree, from whence very pretty snap-
shots may be obtained, as they will often rest on a high branch till the sportsman has
arrived underneath before taking wing again. Both cocks and hens make a desperate
cackling and flutter when thus roused up by dogs, and I know of no shooting which
requires greater nerve and steadiness. If there are no dogs the birds will not tree,
but run slyly and silently along and are seen no more, unless you be mounted on an
elephant, when it is easy enough to fof them, should you be so minded, as they skulk
under the brushwood.”—(Col. Tickell).

“In some places, where the borders of the forest are much broken and irregular,
and the villagers have cultivation here and there between patches of wood and bushes, I
have seen capital bags made by a couple of guns, three or four beaters, and a few
‘bustling spaniels. The plan is this: to beat out strips and patches separately, and make
a corner here and there, placing the guns in the first instance between the patch of
standing crop about to be beaten and the forest towards which the Junglefowl when
flushed are certain to make. The birds, finding their retreat cut off, and pushed hard
by men and dogs, are forced to take flight, and when well on the wing offer as fine a
shot as a sportsman could desire.”’—(Capt. Baldwin).

“The Red Junglefowl is found in the Garo Hills, and in all the Assam plains
districts. I shot the bird beyond Sadiya the other day. It is a permanent resident in
Assam ; it is found in bamboo and tree jungle, and is very often numerous near villages.
In the low hills near Ststing, in the Mymensing district of Bengal, I have had excellent
sport with these birds. I had the hills thoroughly beaten by beaters, the guns being
carefully posted across the line of flight of the birds. I have shot ten to twelve couple
in an hour’s shooting in this way. The best time for this sport is just at the season
when the cold weather rice crop is ready for the sickle; say, during the month of
December and early in January.’—(Col. Williamson).

RED JUNGLEFOWL 199

“The best way of shooting these birds here in the Sundarbans is by proceeding
morning and evening along the edge of the forest, between it and the rice-fields. The
sportsman will thus flush two or three coveys of them, and secure a few brace. The
largest bag that could be obtained by a single gun would hardly be more than three or
four brace.”—(Mr. Rainey).

_ “The Burmese Junglefowl is similar to those in Assam and over the greater part of
India, but perhaps a little smaller. It is a game little fellow, and at times affords
excellent sport. I used to take up my position on an elephant and have them beaten up
towards me, and once or twice I found them as plentiful as pheasants in a preserve in
England. Once on the wing, they are very swift and strong fliers, and require a good
blow to knock them over.—(Pollok).

“In Perak I found Junglefowl breeding from March to July ; and the young, when
three or four weeks old, were capital eating—far better than the full-grown bird, which
has but little more game flavour than the ordinary domestic fowl.

“At the first glimmer of daylight, and again towards evening, the Junglecocks
may be heard in all directions, crowing loudly, and by very careful stalking may
occasionally be got at; but I found far the most successful plan was, either early in the
morning, or else about sunset, to sneak quietly along the edges of clearings and patches
of cultivation, which at those times the Junglefowl frequent in search of food; and in
this way, by dodging from bush to bush, I frequently rendered a good account of them.
But it required the most careful stalking, as on the slightest alarm the: birds ran into
the thick jungle, where it was almost useless to follow them, Once or twice I shot them
in thick cover, by letting my dog hunt them up into the trees, which they did not leave
till 1 was within range.

“In Province Wellesley, the Malays decoy Junglefowl by imitating the crowing
and flapping of the wings of the cock, when the birds coming to accept the supposed
challenge are shot.”—(Kelham).

From personal experience in the actual jungle in the Malay States and elsewhere,
I can assure the sportsman that hunting the Junglefowl is far from being a simple
matter of dogs or beaters. There may be many all around you, and yet if you are not
willing to abase yourself to the very earth and stalk them with all the woodcraft which
you possess, and then shoot the bird as it stands or perches, you need not expect
success. Of the Philippine Junglefowl we read, “ They are difficult to shoot, as they
are very wary, and I have hunted for hours with birds crowing all about me without
getting a shot.”

COCK-FIGHTING

Cock-fighting has been carried on through much of the historical period, and the
literature of nations contains many interesting accounts of this sport. The mention
of one must suffice for us.

The seriousness with which cock-fighting is undertaken by the natives of India is
shown by a most interesting and amusing treatise called Murgh-Nama, or “ Extract
on Cocking” as we might translate it. This was written by Nawab Yar Muhammad
Khan of the Rampur State many years ago, and formed a part of the Sayd-gah-z
Shawkati, an Urdu work on sport. I give a few extracts from this, translated by my

200 A. MONOGRAPH OF THE PHEASANTS

friend Lieut.-Colonel D. C. Phillott, for the Asiatic Society of Bengal, the translation
being as close as the English language will permit.

‘The cocker must know that there are in Hindustan four breeds of fowl. First,
there is the fez, or common barn-door fowl, kept for domestic purposes; second, the
ghagas or cross between the game-fowl and the ew? third, the arnatak, a breed in
which the skin, bones, tongue, eyes, and blood are all black; this breed is useless for
fighting ; and fourth, the as#/ or Indian game-fowl, which is kept for fighting only.

“The points of the Indian game-cock are: beak white, shanks white, eyes white
and blood-shot (if the white of the eye be slightly yellowish it does not signify), the
comb of medium size, the jaw and cheeks large and hard and with little flesh, the bones
of the neck small, the whole neck being like an iron bar, the pope’s nose large, the tail-
feathers small, and the jaws and wings not fleshy. Its crow is not as long as that of
the barn-door cock. The best breed is obtained from Haiderabad, Deccan.

“When the game-cockerels are a year old, the cocker should keep them apart,
feeding them on dajva or millet wetted in water. Should éajva not be obtainable, he
must substitute wheat-flour mixed with butter. In either case he must give them but
little water to drink. When the cockerel is in high condition, he must foment it with
butter (the hand is warmed before the fire, the palm and fingers dipped in butter and
pressed on the joints), and begin feeding it on the yolks of two eggs to begin with.
The following is the method of feeding on the yolks of eggs :—

“Take the yolk of one egg, one fo/@ (about 200 grains) of butter, one masha of
Sambhar salt, and four ma@shas of pepper-corns; mix together and half-fry, and give
to the cockerel, giving it water every third or fourth day. This will make it strong
and lusty.

“Another receipt. Take the yolks of 200 eggs, of musk 2 mdashas, of good
ambergris 2 mashas, of saffron 2 mashas, of white-shifted sugars 5 sevs, of almond
kernels 2} sevs, of kernels of the Edible Pine 14 sevs, of pista kernels 2 sevs, of
walnut kernels 1 sev, of butter 6 sevs, and of fine wheaten flour 5 sevs. First fry
the flour well in the butter, and then cast in the shifted sugar. Next add all the
ingredients (with the exception of the saffron), having previously pounded them
well. Then add to the mixture the saffron well rubbed down in “ewva water.”
The first day give one Zo/@ as a dose, and on the top of that a feed of wheaten flour
unmixed with butter, so that the grease may be removed from the cock’s throat. Then
put on the muzzle, either the leather-muzzle or the string-muzzle (so that it may
neither pick up any grit nor drink) and let it loose to exercise itself. There are two
times for exercise, first early in the morning till seven o'clock, when the cockerel should
be fed, spouted (spouting is spraying the bird with water from the mouth, @ da a Chinese
laundryman) with water, and confined under a square or around coop. At four o'clock
it must be released, spouted as before, and muzzled and freed for exercise till five
o’clock. The cock should be studied, to see whether it has increased in strength or not.
If the cock is lusty and strong, more Za/wa should be given to it and less flour. If it
is not strong and has become fat, it should, at night, be dry-fomented with a pad. If
fat has collected on the /angot (stomach), it must be reduced by dry fomentation, and
the cock must be kept confined in a warm place away from cold air. In the morning
it should be fought for two Aant—{do pani: a match is made for many faz or intervals

RED JUNGLEFOWL 201

for rest. The first day, the first faz is 20 minutes, the second 30, the third 4o, the
fourth 50, and the fifth 60. On the second day (or any Aan? after the fifth) every panz
is 60 minutes. On the first day, the match may commence at any time between two
and four o'clock, usually the second hour. On the second day the match must com-
mence at two o'clock. An antagonist failing to come to the scratch when time is called,
or, to turn up at the correct time loses a Jau?. Say A. and B. make a match for five
pani. A.sees his cock getting the worst of the encounter; he claims a fawz. The fight
is stopped for twenty minutes while both pitters spout their cocks, etc. A., however,
has now only four pai to his credit, while B. has still five. The cocker who first loses
his five Aavz is counted the loser, and has to pay. Though the duration of each Janz is
fixed by scale, no limit is fixed for each round in fighting. Written rules are not drawn
up; custom alone rules|—with a dal6a—|dalba is a weak barn-door cock or an inferior
game-cock, kept for a young game-cock to bully. Cu¢tha@ is a quail kept for a fighting
quail to bully|to see whether it has improved in condition or not. If it has improved,
then keep it at this weight and condition. If it has disimproved, fight it again, fat, thin,
and medium, and observe in which condition it fights best and keep it in that condition.
Every week increase the number of rounds with the da/éa by one, till the number of
eleven Jani is reached, which is called by cockers /am.

‘An ointment for dressing is compounded—{I have omitted the tedious details of
weights and measures (W.B.)|—of zeodary, cassia bark, rennet, pomegranate-rind,
white cumin seeds, Indian madder, gall oak, betel-leaf juice, and a ‘sufficiency of
double-distilled wine.’ Grind these all up, beat in the wine and betel-leaf juice and
anoint the cock’s face. |

“Just before battle give a cock the following: syrup of sweet pomegranates,
preserved apples and quinces, barberries, cucumber seeds, endine chicory, and de/ fruit.
Grind, add the pomegranate syrup; roll into pills, cover with gold and silver leaf, and
give to the cock; after one ghavi [about half-an-hour] go to the cock-pit and fight your
cock. If you find a lack of fire in the cock, give it half a fresh ja/edz [a sweetmeat made
of milk, flour and sugar], pepper-corns, and the egg of a red fowl, mixed with the juice
of garlic and the juice of green ginger. Next, tying up the spurs with cloth, and then
spouting the bird, make it fight. It then rests with the cocker to make or mar the
bird.

“In the author's opinion the best birds have thick, powerful beaks, white, as
described above; the eyes white, like lustrous pearls; large jaws and head; a small
comb; the ¢#fau bone is conspicuous; the neck short and the neck-bones small, fine,
and the ridges small, fleshless, and strong as an iron rod; and the back broad like the
stone of a hand-mill; the feathers spotted, and beautiful, like the eye of a peacock’s
feather. The bird should be handsome, and shapely, and active, and quick as a cobra
in movement ; and in fighting it should be Zama-gir [attacking any part of an adversary’s
body with the beak], and retiring after a blow, so as to avoid its adversary’s counter;
and should it receive a blow, it should so retaliate as to lay its adversary at its feet in
the throes of death, fluttering as though its throat had been cut.

“To make the young cockerel strong and fit for battle. When the cockerel is four
months old the cocker should separate it from the mother and make it familiar with
him, and give it daily two almonds mixed with @/a@ [coarse wheaten flour], and feed it

VOL. II DD

202 A MONOGRAPH OF THE PHEASANTS

on pila, or salan |salan, anything eaten with bread or with rice, and specially vegetables
cooked with meat, such as curry, etc. Game-fowls are fed from the hand, and are not
allowed to pick up food from the ground, lest the beak should become worn. Some
game-fowl were so wedded to the hand that they will starve rather than pick up food
from the ground]|—etc., z.e., whatever the cocker eats himself, and give it kalwa, of
yolks of eggs.

“In the morning give half a belly-full of the 4a/wd, and on the top of it half a
belly-full of balls of coarse wheaten flour. After exercising the cock, put on the
muzzle, spout water over its face, shampoo (by grasping and pressing all the limbs
and joints) with the hand, and confine in a ga/gul or square coop, releasing it at four
in the afternoon. After a time look and see if it is thirsty, and if it is, take it out
of the coop and give it water. At four o'clock release it. If it is thirsty it will drink.
If it is not thirsty and won't drink, then again spout it-and shampoo it, and exercise it
for an hour. After that bring it into the house, and confining a hen in a small basket-
cage, let loose the cockerel that it may see the hen and run round and round her, and by
the exercise increase his strength, but do not let him mate with the hen. After running,
lift up the cockerel, and shampoo him. In the evening see whether the cock has
digested its food. If it has, then in the evening too give it Zalwd, etc., as before. If,
however, the cock has not digested its morning meal, give it a few whole pepper-corns ;
the cocker had, too, better continue to give it pepper-corns in the evening, and keep it
awake by lamp-light for an hour, tickling it under the pope’s nose to make it oil itself.
Then setting it down on a swing made out of a child’s cot, swing it for an hour, or else
place it on a perch. Then confine it somewhere in a safe place in a basket-cage. In
the morning at prayer-time [z.e. about an hour before sunrise] take it out of the coop,
give it water to drink, and then spout it, and, putting on the muzzle, let it exercise itself
for a full hour. After this feed it. You must gradually increase the amount of Za/wa, and
decrease the amount of flour, feeding and treating the bird as already described for forty
days. Once a week too, at night, you must foment it with a damp, hot pad, fomenting
those parts of the body that have accumulated fat. The places to foment are, first from
the head to the neck, or rather to the shoulders, next the two wings on the top, next the
thighs inside and outside, also the hip-joints and the loins, and underneath between the
legs, omitting the lower stomach, and also the breast. The object of fomenting is to
make flesh and joints hard and strong.

‘“When the cockerel is ten months old, fight it one fax7 with a dalba of its own
age, but first put the muzzle on to the da/éa. Afterwards spout and suck the cock’s beak
and face well, to remove blood-spots, and damp a rag in water and clean the inside of
the throat with it.”

The dalba is muzzled and hobbled before being pitted against its aristocratic
antagonist.

One writer states that a cockerel should not be fought till he is one year old, and
that he should then be pitted against one of his own age for five minutes the first day,
ten the second, and fifteen the third, and then for as long as desirable; and that at the
expiration of the fixed time the birds should be separated and their beaks sucked and
blown upon. (If a child falls and hurts itself, Indians generally blow upon the seat of
injury with the idea of cooling the spot and lessening the pain.)

RED JUNGLEFOWL 203

“ Tf the jaws be fleshy, or if the wattles are large, and it is desired to remove them,
then extract the oil of 642/awan and rub it on the cock’s face, and if it please God this
defect will be removed and will not return. If the comb is too large and it is desired to
cut it off, then, after fighting, cut the comb immediately to the size desired, applying
to the wound the white web of a small spider, or else staunch with small feathers plucked
from under the cock’s wings: until a recovery takes place do not spout the cock. Then
take a quarter of a piece of tumeric and add half a nux vomica nut and a little edible
lime and heat over the fire, and plaster the mixture thickly on the cock’s face ; if it is the
cold season, keep the cock in the sun, if the hot, in the shade. The cock must not be fed
during the day, but in the evening after fomentation, when it should be fed on “a/wa or
on bread dipped in fresh milk. In the evening the cock should be fomented with a pad
dipped in a liquid decoction.

“ After fifteen days, when the cock has recovered from its wound, let it be fought for
two fant with the unmuzzled da/ba, and act as described above. After twenty-two days
let the trainer fight his cock three az and act as before. After forty days let him fight
four fanz, and lastly, after an interval, five fauz. After fighting five Jazz, he should
foment the cock’s face with a small bag of Za/wa, and dipping a pad into the hot liquid
application, he should foment the loins, the hip-joints, both the shoulder-joints, the
breast, and between the legs as described above. After this evening fomentation, he
should feed the cock on Za/wa, or on bread dipped in fresh milk. In the morning he
should again foment in the same manner, and, mixing some of the best szzssz in wine,
should heat it on the fire and paint the cock’s face thickly with it. Increase gradually
the number of Aauz up to eleven, acting as just described. Then fight your cock for a
wager with some one, having first put it on dry food, and please God the cock of your
antagonist will be unable to face yours.

‘Just before fighting your cock, you (being in a state of ceremonial purity) should
repeat once into each of its ears the words, ‘“‘ Allah the Immutable,” and exhale your
breath, and if it please God Most High your cock will never deteriorate.

“Should your cock show signs of flight, then give it this corrective :—

“Musk, fabashiy, small cardamoms ; pound and put in a paper. Take of juice of
Indian sweet fennel seed two ounces and put it in a vessel. When you see that your
cock intends flight, first give it two ma@shas of the powdered medicine, and then two
tolas of the juice. Your cock will go mad and will never quit the field.

“When you wish to cease fighting your cock, and to pair him with a hen, do so, but
pair him with a mature hen, but this must be in the warm days of spring. You must
not mate your cock with a young hen.”

LEGENDS AND SUPERSTITION

Since the first domestic cock crowed, mankind has admired, hated or feared it, has
thought of it with reverence, with dislike or with superstition. We have space for
mention of but one or two of the legends which have arisen concerning it.

One Al-Qazwini, who died about 1283, wrote an Arabic work, the 47a 7b-ul-
Makhlugat, or the ‘World of Wonders,” and has this to say about the domestic cock :—

‘The cock is the most lustful and vainest of birds. It heralds the dawn. One of

204. A MONOGRAPH OF THE PHEASANTS

the strangest things about it is that it knows the watches of the night, and apportions
the times for its night-crowing according to the length or shortness of the nights ; for
instance, if the night is fifteen hours in length, he crows, at stated intervals, the same
number of times as he does in a night of nine hours; and this he does by a God-given
instinct. It is related that the Prophet (may the Peace and Blessing of God be upon
him) said, ‘God the Most High has created a cock beneath His Throne, with wings that
can extend beyond the East and the West; and towards dawn he spreads his wings,
and flaps them, and raises his voice in praise of Him, crying, ‘‘Glory to the Most Holy
King,” and when he has ceased, all the cocks in the Earth join in returning that cry,
flapping their wings in like manner.’ It is said that the chief caller-to-prayer amongst
the cocks is that breed that has long wattles and a castellated comb. The cock has a
sense of jealous honour about his wives, and he is generous to them, and cares for them.
It is a belief that, should a man rise from sleep at cock-crow, he will be fresh and bright
all day. A white cock puts to flight the lion. The best of cocks is the game-cock. Its
points are: a red comb, a thick neck, small and black eyes, sharp claws, a loud cry.
A cock is unselfish to his hens; he takes a grain in his beak and casts it to them. It is
said that he does this in the time of his youth, when his passions dominate him ; but
that when he ages he no longer does so. The cock defends his hens from the attack of
an enemy, and at night collects them in a safe place and stands guard at the door.
They say, too, that the cock lays one egg in his lifetime, called in Arabic dayzat '-'ugr,
and that it is very small. The following lines are by the poet Bashshar :—
“* Thou hast visited me but once in all this time.
Make not thy visit rare like the egg of the cock.’

“Tt is a belief that one who slaughters a white cock with a divided comb will suffer
loss in his possessions and in his house, and also that the Devil never enters a house in
which such a cock is to be found.. As for the properties of the several parts of the cock,
if the comb be dried and pounded and given to drink, bad habits will depart from a
man. The smoke of the dried comb of a white or of a red cock does good to a madman.
The gall applied as a collyrium to the eyes, cures dimness of sight, or a film over the
eye. Some physician has said that the gall must be placed in a silver vessel and used
continuously to obtain a cure. Polonias has said that the gall of a cock, mixed with
mutton-broth and taken in the morning on a fasting stomach, is a cure for loss of
memory. If the wing-bone be bound on one suffering from intermittent fever, the fever
will depart. Ifa rider ties that bone on his loins he will suffer no fatigue. The blood,
used as a collyrium, is beneficial for film over the eyes. If the blood drawn in a cock-
fight be mixed with food and given to a number of people, it will cause dissension
amongst them. If you take a cock’s blood and mix it with honey and place it on the
fire, the application will increase a man’s virility. If you take the dried flesh of a cock
and pound it with equal quantities of gall-nuts and sumach, and make pills the size of
peas, and administer them with a draught of water to one that has a pain, he will be
relieved on the spot. In the stomach of the cock there is a pebble, sometimes sky-blue ©
in colour and sometimes crystal, which, if suspended around the neck of a madman,
cures him.”

RED JUNGLEFOWL 205

THE COCK IN HUMAN HISTORY

Several years ago I had the enviable opportunity of sharing with Dr. John P. Peters
an attempt to trace the relation of the cock to mankind from ancient historical times.
There seems little doubt but that the original home or centre of distribution of the Red
Junglefowl, the most anciently inhabited part of its present range, lies to the east and
south, in the Burmese-Malayan portion of its habitat, rather than further east in China,
or to the west in the Indian region. Distributional study of other birds, such as the
Himalayan kaleege pheasants, makes it certain that these originated in Southern Burma
and have since migrated westward, along an elongated sub-Himalayan finger, as far as
Kashmir. The same holds true of a number of other-forms of life, both mammalian
and avian. There is no doubt that the Red Junglefowl is of tropical or sub-tropical
origin. Neither it nor its domesticated descendants can bear extreme cold; and the
elaborately specialized exposed comb and wattles could have been evolved only in a
warm country. ‘The three other species of Junglefowl are all tropical, and the affinities
of the group among the other pheasants are altogether with Southern Chinese and
Malayan genera.

Like the mallard duck.and the rock dove, the Red Junglefowl has, for times long
antedating human history, been a species especially susceptible to domestication, and
with early civilizations such as arose in China and India, it is not surprising that
domestication and variational breeding began before the proofs of hieroglyphics and
written record.

Tradition carries back the domestication of the cock in China to as early as
1400 B.c., and the modern name for the bird, £2 or Raz, can be traced to the Chou
Dynasty, which extended from 1122 to 249 B.c. In a very ancient dictionary £2 is
defined as “the domestic animal which knows the time.” In the /7-ya, a glossary of
the time of Confucius, it again is found. While there is evidence that in very early
times the cock was an object for sacrifice in China, yet those people were quick to see
the utilitarian value of the fowl, and to breed several distinct varieties. The Cochin
China fowl is one of the most extreme, mutation-like of the early established breeds.

In India it is necessary to depend upon literary rather than monumental evidence
of the early occurrence of the cock. In the Indus period of the Aryan invasion, the time
of the Rig Veda, there is no mention of the cock, but about 1000 B.c., when the Aryans
had reached the Ganges, in the Atharua and the Yajur Vedas, the cock is well known.
This supplements our zoological distribution theory, and also emphasizes the significance
of the fact that Solomon’s Pheenicians, who found and imported the peacock from that
part of India which they could reach by sea, did not find the cock.

In the White Yajur Veda the name of the cock, kukkuta, is onomatopoetic. We
read, ‘Then cut a cock whose tongue is sweet with honey; call to us manly vigour ;
may we with thee in every fight be victors.” And a verse attributed to Chanakya, about
320 B.C., says you may learn four things from a cock :—

1. To fight; 2, to get up early; 3, to eat with your family; 4, to protect your
spouse when she gets into trouble.

If, as seems certain, the cock moved northward and westward, against the line of
Aryan invasion, it should have reached Bactria and Persia at a very early date, and

206 A MONOGRAPH OF THE PHEASANTS

judging by the part which the cock plays in Persian religion and mythology, and the
manner of reference to it in Zoroastrian literature, its advent must have antedated
considerably 600 B.c. .

The first mention of the cock in Grecian literature is by Theognia about 525 B.c.,
but his image occurs on coins from the temple of Artemis at Ephesus of at least 700 B.C.
In the Metropolitan Museum in New York is a Corinthian Alabastron with a well-
executed cock, dated 650-600 B.c. As to the Chinese the fowl was primarily an edible
bird, and to the Persians an object of sacrifice, so to the later Greeks his fighting
qualities appealed strongest. To them he was primarily a game cock, and cock-fights
were the commonest representations on coins. While so common during the middle
and later periods, the cock does not enter into Greek mythology, neither is it mentioned
in either Hesiod or Homer, nor in those widespread legends or folklore tales which
originated in the early period of Greek development.

The establishment of the cock in Greece was only a slight advance compared with
the steady progress along the line of the Iranian invasion, which carried the bird through
Bactria and Persia on into Scythia and Europe, stretching across finally to the British
Isles and spreading down from Gaul into Central Italy. Long before Greek colonists
carried the bird to south Italy, it had passed on to the northward, and was being carried
southward through Italy on the line of an independent advance. The first European
distribution of the cock was overland rather than by sea, or by coastal colonists. The
Romans found it well established in Gaul, England, and among the Germans.
The Greeks knew it as the Persian bird, the Romans called it ga//us after their return
from Gaul. Czesar tells of the religious significance of the cock among the Gauls and
of its religious importance. Indeed we realize that no nation has more right to its
national bird than France, by constant association and symbolism from the earliest times
down to le chanticler daujourd’ hui, the singer, the herald of the dawn.

The Greeks carried the cock southward to the Phoenician cities, but only at a late
date did it become well established on the Syrian mainland. As early as 700 B.c. the
Assyrians and Babylonians received the fowl from the Medes and Persians, where it had
been known since 1000. We do not know where the Aramzans first obtained the cock
—those natives of the lands stretching from the western frontiers of Babylonia to the
highlands of western Asia. But from them in the intercourse of later times the cock
was introduced to the Jews in Palestine about 200 B.c.

Beside the dove and peacock there is no mention of domesticated fowls in the Old
Testament, with one or two very doubtful exceptions. Certainly during all the time
covered by that period the cock was quite unknown to the people as a whole. Both the
New Testament and the Talmud mention the cock, as in Mark xiii. 35: “Watch ye
therefore ; for ye know not when the master of the house cometh, at even, or at midnight,
or at the cock-crowing, or in the morning.” And in Berakhoth 60 b: “ Praised be thou,
O God, Lord of the world, that gavest understanding to the cock to distinguish between
day and night.” To-day the cock is the one sacrificial animal of the Jews; at Yom
Kippur as an atonement offering—a cock by a man, a hen by a woman. And indeed in
ancient Palestine there was at first considerable opposition to the ordinary economic
breeding of fowls, by those who considered the birds as special objects of religious
significance.

RED JUNGLEFOWL © | 207

Strangest of all is the historical distribution of the cock in Egypt. The Egyptians,
who reached far and wide through their dependencies for strange creatures and objects,
whose artists crystallized in exquisite sculptured bas-relief and paintings even the
separate species of ducks and geese, doves and quail with which they were familiar,
could never have failed to depict the characteristic mien, or the comb, hackles or tail of
the cock, had they known that bird. Yet before the Roman period there is no image
on Egyptian monuments, nor word or sign for cock in their language. It is strange
that no hint of chickens or casual fowls should have been brought by the Persians or
the Grecian mercenaries of Psammetichus. However we may fail to explain the silence
of all the dynasties, no cock or hen is known to have reached Egypt at an earlier date
than 50 B.C.

DETAILED DESCRIPTION

ADULT Marr.—As I have emphasized in the course of narration of this species, the
constant interchange of blood, the continued crossing of the wild Red Junglefowl with
native birds, has brought about an amount of variation in both habits and coloration
which is unique among. pheasants. The following descriptions of the birds refer to
specimens which appear to be pure-blooded, from birds which were secured at a
considerable distance from human habitation.

Top of the head, neck and upper mantle orange red or dark orange, the longer
hackles paling posteriorly into orange or orange-yellow. This latter colour is confined
to a very wide disintegrated border, the concealed central, solid portions of the vane
varying from slightly darker to a smoky brown. These hackles are slender and greatly
elongated, covering and concealing the real feathers of the mantle and upper back. Mid-
mantle dull brown, sometimes showing a purplish sheen. Lower mantle and lesser
coverts glossed with purplish blue or green. A line of feathers bordering the entire
ventral border of the hackles, scapulars, central line of the lower back and median
wing-coverts rich dark maroon red, shading into orange-red or orange on the long, soft,
hackle-like feathers of the sides of the back, rump and upper tail-coverts. Most of the
maroon feathers have a concealed zone of iridescent green. Greater primary coverts dull
brown; secondary coverts black, strongly glossed with purplish green. Flight feathers
brownish black, the outer margin of the primaries pale brownish buff and the outer half
of the outer webs of the secondaries chestnut-buff anteriorly, deepening into tawny
toward the tips of the feathers. This colour gradually disappears from the inner
secondaries, and is replaced on the tertiaries by a gloss of green. Longer upper
tail-coverts solidly glossed with green; the central tail-feathers lack this sheen along the
shaft and toward the tip, while it dies out on the lateral rectrices, leaving them dull dark
brown. There is no green gloss on the underside of any of the feathers. Ventral surface
of the neck, breast and all the under parts uniform brownish black, faintly glossed with
greenish. A fleshy, bare, deeply notched comb arises from the forehead and crown.
There are two rounded gular wattles, and a large lappet growing just beneath the
opening of each ear.

The bill is dark brown, sometimes reddish toward the base in full-coloured males,
and paler at the tip of the lower mandible. The legs and feet are lead-coloured or slaty,
sometimes rather brown, purplish or tinged with greenish, while again they may be pale

208 A MONOGRAPH OF THE PHEASANTS

slaty grey, irrespective of age, sex or locality. Iris light to rather deep orange-red. The
bare skin of the sides of the head, skin, throat and upper part of the neck in front,
smooth and of varying shades of red. The large erect comb, thin and deeply notched
above, and the two gular wattles vary from deep dull red to bright crimson. When
a large series of birds with definite locality labels is examined it is evident that the ear
lappets tend to be whitish or pinkish-white in Indian birds, and red, like the wattles and
comb in birds from Burma and the Malay Peninsula. The proportion is about forty
per cent. of each, leaving twenty per cent. which are neutral or actually negative. I have
seen many Junglefowl from Pahang, Johore and Java, which had the ear-lappets pearly
white, and, on the other hand, I have shot birds in southern Garhwal with the lappets
showing no trace of light colour, being indistinguishable in hue from the comb and
wattles, even when these were very bright in colour. In northern Burma I have
secured two adult Junglecocks feeding together with several hens in a part of the jungle
far distant from any native fowl which respectively showed the two extremes in colour
of lappets. Where white is present, it is like enamel or polished ivory, and changes to
violet or blue where it merges into the red of the face or upper part of the lappet. It is
true that Indian birds, especially those from the dryer, semi-arid regions, are noticeably
pale, while the Junglefowl from the terai and the Malay States are richer, with the red
more brilliant. I do not, under the circumstances, however, consider it right to give
these subspecific designation. There is a small area in Pahang where the Junglefowl
are paler than any Indian birds, probably due to the infusion of some pale domestic
strain. And again we find many accounts written by sportsmen which present such
facts as those of Kelham in the “Ibis” of 1882: ‘‘ Whether or not the Malay species,
Temminck’s G. dankiva, is really distinct from the Indian, it is hard to say; but if it is
distinct, both kinds are certainly found in the Malay countries; for while stationed in
Perak I shot, out of the same tract of jungle, unmistakable specimens of G. ferrugineus,
with the rich golden hackles and white ear-patches, also birds of far darker, in one case
almost black, plumage. But the wild Junglefowl interbreed so much with domestic
roosters from the villages, that I cannot help thinking these dark-coloured birds to be
the results of such intercourse, particularly as many of them, though very unlike the
typical G. ferrugineus, are not like one another, varying much in the intensity of their
colouring.”

Weight, 1 lb. 12 ozs. to 2 Ibs. 5 ozs.

Length, 630 to 700 mm.; culmen from nostril, 16; wing, 235; tail, 350; tarsus, 77;
middle toe and claw, 55. Spurs averaging 33 mm., sharp, slender and much curved.

VARIATION AND MOULT

The hen-coloured game cocks are, of course, congenitally so coloured, but it is not
unusual, as in other true pheasants, for the cock bird ontogenetically to assume the
plumage of the hen. This is due to some injury to the sexual organs, and is one of the
most inexplicable phenomena known to science. Its solution will mark a decided step
in problems of evolution, and perhaps in the origin of life. Recent experiments by
Morgan have indicated that the sexual organs of the hen-coloured Sebright bantams
contain luteal cells which are usually characteristic only of ovarian tissue. This, together

RED JUNGLEFOWL 209

with the well-established acquisition of the cock’s plumage in sterile hens, shows that
these cells exercise a very important suppressive effect on the secondary sexual characters
of the male bird.

Perhaps the most interesting thing about the moult of Red Junglefowl is the
partial eclipse plumage into which the cock enters immediately after the breeding season.
The general facts are the same as those given under Ga//us sonnerati (p. 245). The long,
red and yellow hackles are shed, while the remainder of the worn plumage remains intact.
They are at once replaced by short, dusky brown or black feathers, not like the hackles
of the hen, but very similar both to the hackles of the juvenile plumage and to the
normal feathers of the mid and lower mantles, which in the full-plumaged cock are
concealed from view by the overlapping, long lanceolate hackles. This eclipse plumage
has sometimes a gloss of purplish or purplish blue. The long central tail feathers are
sometimes shed simultaneously with the hackles, more often they fall out a little later.
In two or three months the regular annual moult begins and the eclipse hackles are shed
with the rest of the plumage, and in the autumn the cock is seen in all his resplendent
colours, ready for another year of virile life.

No good reason has been advanced to explain this curious partial moult. In ducks
the cause for the assumption of the dull female garb during the period of helplessness,
consequent upon the simultaneous shedding of the flight-feathers, is very apparent.
But this time of the year presents no unusual dangers to the Junglefowl, nor is the bird
less able to care for itself than at other seasons. For the present we must be content
to state the mere fact that these birds expend a considerable extra outlay of vitality in
growing a localized patch of feathers, which lasts only a few weeks, and which in
character is both atavistic and like the generalized feathers of the upper back. The
time of assumption of this generalized neck plumage varies with the locality. From
the terai south of Darjeeling I have a bird in full moult, shot on the twentieth of May,
but this is unusually early even for the northern range. In the Malay States, August
seems the usual period. Sportsmen who shoot birds in this condition usually consider
them as abnormal, perhaps hemaphrodite individuals.

I believe that in birds of strictly wild blood and pedigree, this eclipse moult is
regular, but in several cocks which I have shot, and in many more which I have trapped,
I have noticed a great irregularity and even asymmetry in this moult, due, I am
convinced, to the infusion of the blood of native village birds.

JUVENILE PLumace.—In this phase the neck and rump-hackles are short and
broader, with considerable variation, due to the earlier or later moult of the individual
bird. The colours of these feathers are brighter and more distinct, the dark centres
being larger, and the edges yellow, rather than orange. The flight feathers and their
coverts are dark cinnamon, mottled and finely vermiculated with black.

ApvULT FremMaLe.—The variation in the plumage of the hen is, within narrower
limits, correspondingly as great as in the cock, and, of course, attributable to the same
causes.

The crown of the head is rusty red, shading into orange on the neck and pale
yellow on the mantle, all the feathers with a wide black stripe down the centre. On

VOL, II EE

210 A MONOGRAPH OF THE PHEASANTS

the mantle this stripe is strongly glossed with green. Remainder of the upper plumage,
beginning rather abruptly behind the mantle feathers, pale reddish or buffy brown,
finely mottled and vermiculated with black, with a conspicuous, narrow, yellowish-white
line down the shaft. Flight feathers brownish-black, the primaries very narrowly edged
with buff. The visible portion of the secondaries (four-fifths of the outer web) vermi-
culated with pale reddish-brown. ‘Tertiaries evenly vermiculated, with distinct metallic
green sheen on the darker portions. Tail-feathers with rufous brown mottling, confined
to the edges of the feathers, and considerable green gloss. Comb sometimes a minute
fleshy knob; again a low, inconspicuous notched ridge. Wattles absent, and ear-lappets
greatly reduced. Face and upper neck chestnut. Lower neck and breast light red,
shading posteriorly, and on the sides, into pale buffy-brown, each feather with a pale
shaft. The feathers have more or less dark mottling, especially those of the sides,
where it takes the form of irregular, concentric bands. The breast and mid-belly is
usually monochrome. Under tail-coverts blackish-brown, with slight brownish
mottling.

The bill of the hen is usually paler than that of the cock ; but otherwise the colours
of the horny and fleshy parts are the same in both sexes.

Weight, 1 lb. 2 ozs. to 1 lb. 10 ozs.

Length, 420 to 460 mm.; culmen from nostril, 15; wing, 185; tail, 153;
tarsus, 60; middle toe and claw, 50. Spurs absent.

CuIcK IN Down —Head fawn-coloured or yellowish buff; a wide maroon or
chocolate coronal patch, pointed in front and narrowly bordered with black. A dark
brown line extends through the eye, and back over the ear-coverts. The dark crown is
continued down the neck and over the back, but is split on the sides of the lower back
and rump by two wide creamy-buff lines, these also outlined in black. Under-parts
creamy white, the breast sometimes deep buffy. Bill, yellow; legs and feet, yellowish
green.

HYBRIDS

Gallus pseudhermaphroditus of Blyth was based upon an abnormal domestic
specimen, said to bea male. The bird had the comb, wattles, spurs and tail of a cock,
with a hen-coloured body plumage.

“Gallus stramineicollis, type Sharpe. Sulu Islands.” ‘Thus is labelled the type
in the British Museum. It is a very large hybrid or domestic variety, with a small
rose comb; bare face, chin, throat and lower neck; hackles, mantle, neck, lower back
and rump, straw colour with narrow green shaft-stripe. Feet and legs very thick and
heavy. The spurs are abortive. Culmen from comb, 18 mm.; wing, 230; tail, 410;
tarsus, 96; middle toe and claw, 70.

SYNONYMY

Phasianus gallus Linn. S. N., XII. 1766, p. 270; Raffl. Trans, Linn. Soc., XIII. 1822, p. 319 [Sumatra].
‘Grande Caille de la Chine Sonn. Voy. Ind. Orient., II. 1782, p. 171.

Hlackled Partridge Latham, Gen. Syn., II. 1783, p. 766, pl. 66; id. Gen. Hist., VIII. 1823, p. 307, pl. 129.
Letrao ferrugineus Gmelin, S. N., I. 1788, pt. I1., p. 761.

RED JUNGLEFOWL 211

Perdix ferruginea Latham, Ind. Orn. II. 1790, p. 651.

Coturnix spadicea Bonnat, Tabl. Encycl. Méth,, I. 1791, p. 218. ,

Gallus bankiva Temminck, Pig. et Gall. II. 1813, p. 87; III. 1815, p. 654; Steph. in Shaw’s Gen. Zool., XI.
1819, p. 198; Horsf, Tr, Linn, Soc., XIII. 1822, p, 185 [Java]; Vieillot, Tab]. Encycl. Méth,, I. 1823, p- 364;
Griff, ed. Cuv., III. 1829, p. 20; Guérin, Icon. Reg. Anim., Ois. 1829-38, p. 26, pl. 42, figs. 2, 2a; J. E. Gray, Ill.
Ind. Zool, I. 1830-32, pl. 43, fig. 3; Lesson, Traité d’Om, 1831, p. 491; Schinz, Nat. Abbild. Vog, 1833, p. 342,
pl. 94; Jard., Nat. Lib. Orn. IV. 1834, p. 175, pl.; Hodgson, in Gray’s Zool. Misc. 1844, p. 85; Gray, List of
Birds, Pt. III., Gall. 1844, p. 27; id. Gen. B., III. 1845, p. 499 ; id. Cat. Hodgs, ed. I. 1846, p. 125 ; Schinz, Nat. Vog.
1853, p. 145, pl. 69; Adams, Proc. Zool. Soc. 1858, p. 498 [Simla, S. Cashmere]; Saint-Hilaire, Bull. Soc. d’Acclim.
1859, p. 13 (Origin in History) ; Wallace, Proc. Zool. Soc. 1863, p. 486 [Lombock, Timor]; Sclater, List of Phas.
1863, p. 10; Sclater, Proc. Zool. Soc, 1863, p. 122; Blyth, Ibis, 1867, p. 156; Sclater, Proc. Zool. Soc. 1869, p. 628
[breeding in London Zoo]; Melliss, Ibis, 1870, p. 103 [domestic, St. Helena]; Wald. Tr. Zool. Soc., VIII. Pt. II.
1872, p. 86 [Celebes]; Hartlaub and Finsch, Proc, Zool, Soc. 1872, pp. 89, 103 [Pelew Is.]; Garrod, Proc. Zool.
Soc. 1873, pp. 627, 640 [thigh muscles]; Garrod, Proc. Zool. Soc. 1875, p. 341 [plantor tendons]; Walden, Trans.
Zool. Soc., 1X. 1877, p. 223 [Luzon, Guimaras]; Tweeddale, Proc. Zool. Soc, 1878, p. 953 [Zamboanga]; Tweed-
dale, Proc. Zool. Soc. 1879, p. 73 [Basilan]; Meyer, Ibis, 1879, pp. 138, 146 [Menado and Limbotto, Northern
Celebes; Posso, Central Celebes; Togian Is. habits]; Garrod, Proc. Zool. Soc. 1879, pp. 377-378 [trachea] ;
Sharpe, Trans. Linn. Soc., N.S. Zool., I. 1879, p: 348 [Panay, Philippines]; Gadow, Proc. Zool. Soc. 1882, pp.
318-319 [tarso-metatarsus muscle]; Kelham, Ibis, 1882, p. 1 [Perak]; Guillemard, Proc. Zool. Soc. 1885, pp. 248,
250, 272 [Sulu; habits]; Sundevall, Ibis, 1886, p. 455 [No. of arm-remiges]; Blas., Ibis, 1888, p. 374 [Palawan];
Shufeldt, Journ. Comp. Med. and Surg., Oct. 1888, p. 1 [anatomy and osteology]; Blas., Ornis. 1888, p. 317;
Everett, List of Birds, Borneo, 1889, p. 199 [Balabac, Palawan]; Sharpe, Ibis, 1890, p. §7 [Palawan]; Whitehead,
Ibis, 1890, pp. 40, 57 [Palawan]; Hartlaub, Abh. nat. Ver. Bremen, XII. 1893, p. 324 [Hainan] ; Sharpe, Ibis, 1894,
pp. 241, 257 [Sulu and Bongao Is. Sulu Arch.]; Everett, Ibis, 1895, p. 32 [Balabac Is.]; Hartert, Novitates
Zoologicae, V. 1898, pp. 61, 69 [Marianne Is,]; Finn, Jour. As, Soc, Bengal, N.S, LXXI. 1902, p. 85 [hen-
feathered cock]; Goodchild, Bird Notes, IV. 1905, p. 66 [compared with Black-red Game]; Lawrie, Ibis, 1909,
p. 192 [Increased fertility of domestic fowl]; Robinson and Kloss, Ibis, 1910, p. 672 [Trang]; Stubbs and Rome,
Zoologist (4), XVI. 1912, p. 1 [prehistoric origin of fowl].

Gallus ferrugineus Blyth, Ann. Mag. N. H., XX. 1847, p. 387, 1. 1848, p. 455; id. Cat. Mus. As. Soc., 1849,
p- 242; Adams, Proc. Zool. Soc., 1859, p. 185 [S. Cashmere]; Irby, Ibis, 1861, p. 234 [the Terai of Oudh and
Kumaon]; Jerdon, Birds of India, III. 1863, p. 536; Gray, List Gallinae Brit, Mus., 1867, p.37; Blyth, Ibis, 1867,
p. 154 [general remarks]; Beavan, Ibis, 1868, p. 381 [Darjeeling, Sikhim and Bhootan terai, Umballa ; valley of
the Salween and in Tenasserim, Maunbhoom]; Brooks, Ibis, 1869, p. 60 [Naini Tal, Almorah]; Bulger, Ibis, 1860,
p. 170 [Leeboug, Sikhim]; Swinhoe, Ibis, 1870, p. 357 [Hainan]; Gray, Hand-list of Birds, II. 1870, p. 261;
Swinhoe, Proc. Zool. Soc., 1871, p. 399; Elliot, Mon. Phas., II. 1872, pl. 32; Hume, N. and E. Indian Birds, 1873,
p. 528; Ball, Stray Feathers, II. 1874, p. 426 [Chota Nagpur]; Hume, Stray Feathers, II. 1874, p. 482 [Pahpoon,
Tenasserim, Amherst]; Blyth and Walden, Cat. Mammals and Birds, Burma, 1875, p. 149 [Tonghoo]; Hume,
Stray Feathers, III. 1875, p. 171 [Upper Pegu]; Armstrong, Stray Feathers, IV. 1876, p. 338 [Irrawaddy Delta,
China Bakeer]; Finsch, Proc. Zool. Soc., 1877, p. 780 [Ponapé]; Hume and Inglis, Stray Feathers, V. 1877, p. 44
[North-eastern Cachar]; Oates, Stray Feathers, V. 1877, p. 164; Wardl.-Ramsay, Ibis, 1877, p. 468 [Karen-nee,
Burma, Nest]; David and Oustalet, Ois. Chine, 1877, p. 420 [Cochin China, Hainan]; Marshall, B. Nest India,
1877, p. 59; Hume and Marshall, Game-birds of India, I. 1878, p. 217, pl.; Anderson, B. W. Yunnan, 1878,
p. 669 [E. of Kachin Hills, Upper Burma]; Hume and Davison, Stray Feathers, VI. 1878, pp. 442, 521 [throughout
Tenasserim]; Ball, Stray Feathers, VII. 1878, p. 225 [from the Ganges to the Godaveri]; Hume, Stray Feathers,
VIII. 1879, p. 68 [Malacca, Wellesley Province]; Scully, Stray Feathers, VIII. 1879, pp. 348, 367 [Nepal]; Salvad.
Ann. Mus. Civ. Gen., XIV. 1879, p. 251 [Sumatra]; Anderson, Zool. Results, Yunnan, 1879, p. 669 [Bhamo] ;
Bingham, Stray Feathers, IX. 1880, p. 195 [Thoungyeen Valley, Tenasserim]; Fasson, Stray Feathers, IX. 1880,
p. 205 [Chittagong]; Finsch, Ibis, r88z, p. rr4 [Kushai and Ponapé Is., Caroline Arch.]; Mill., Jour. f. Orn., 1882,
p- 432 [Salanga Is.]; Kelham, Ibis, 1882, pp. 1-2 [Perak, W. Malay Pen. habits]; Nicholson, Ibis, 1882, p. 65
[Gounong Trang and Kitta Djama, S.E. Sumatra]; Oates, Stray Feathers, X. 1882, p. 236 [Pegu]; id. Birds
British Burmah, II. 1883, p. 322 [Siam, general account]; Nicholson, Ibis, 1883, p. 255 [Tandjong Djete, near
Lake Ranau, 1700 ft. Central Sumatra]; Marshall, Ibis, 1884, p. 423 [Chamba]; Biittikofer, Notes Leyden Museum
IX. 1887, p. 78 [Padang, W. Sumatra]; Taylor, Stray Feathers, X. 1887, p. 531 [Orissa]; Hume, Stray Feathers
XI. 1888, p. 304 [Assam, Sylhet, Cachar, Manipur and British Burmah]; Hartert, Jour. f. Orn. 1889, pp. 379, 434;
Tegetmeier, Ibis, 1890, pp. 304-327 [domestic breeds]; Oates, ed. Hume's Nests and Eggs, III. 1890, p. 417
[nesting habits]; Hartert, Jour. f. Orn., 1891, p. 301 (Mindoro); Laurie, Jour. Bombay Nat. Hist. Soc., VI. 1891,
pp. 93, 98; Elliot, Auk, IX. 1892, p. 164 [hybridism and reversion]; Styan, Ibis, 1893, p. 435 [Hainan]; Blanford,
Jour. Bombay Nat. Hist. Soc., IX. 1894, pp. 186, 188; Clarke, Ibis, 1895, p. 479 [Is. of Negros]; Hartlaub, Abh.
nat. Ver. Bremen, XIV. 1898, p. 371 [Hainan]; Blanford, Fauna Brit. India, Birds, IV. 1898, p. 75 [General
account]; Baker, Jour. Bombay Nat. Hist. Soc., XII. 1899, p. 487 [North Cachar]; Inglis, Jour, Bombay Nat.
Hist. Soc., XII. 1899, p. 676 [Hylakandy Dist., Cachar]; Clarke, Ibis, 1900, p. 360 [melanistic specimen from
Negros]; Rippon, Ibis, 1901, p. 556 [Southern Shan States, up to 3500 feet]; Nisbett, Jour. Bombay Nat. Hist.

212 A MONOGRAPH OF THE PHEASANTS

Soc., XIII. 1901, p. 525 [colour of legs of domestic birds]; Hartert, Novitates Zoologicae, IX. 1902, p. 218
[Sumatra]; Meyer, Notes Leyden Museum, XXIV. 1904, p. 235 [South-east Celebes]; Hartert, Novitates
Zoologicae, XI. 1904, p. 174 [Wetter Is. N. of Timor]; Macdonald, Jour. Bombay Nat. Hist. Soc., XVII. 1906,
p 496 [Myingyau Dist. Burma]; Baker, Jour. Bombay Nat. Hist. Soc., XVII. 1907, p. 764 [Singapore]; Ward,
Jour. Bombay Nat. Hist. Soc., XVII. 1907, p. 944 [Kashmir and Jammu]; Baker, Jour. Bombay Nat. Hist. Soc.,
XVII. 1907, p. 971 [Khasia Hills]; Macgrath, Jour. Bombay Nat. Hist. Soc., XVIII. 1908, p. 298 [Thandiani,
Hazara Dist.]; Harington, Jour. Bombay Nat. Hist. Soc, XIX. 1909, p. 309 [Bhamo Dist.]; Harington, Jour.
Bombay Nat. Hist. Soc., XIX. 1909, p. 365 [Rangoon Dist.]; Hartert, Novitates Zoologicae, XVII. 1910, p. 191
[Hainan]; Stubbs and Rome, Zoologist (4), XVI. 1912, p. 1 [prehistoric origin of fowl].

Gallus domesticus Tegetmeier, Proc, Zool. Soc., 1861, p. 102 [cock assuming hen plumage]; Lubbock, Ibis,
1862, p. 180 [possible bone found in Swiss lake-dwellings] ; Sundevall, Ibis, 1886, pp. 435, 439, 441 [arm muscles] ;
p. 455 [No. of arm-remiges]; Miller, Der Zoologischer Garten, XXXIV. 1893, p. 146 [cosmopolitan distribution] ;
Gurney, Ibis, 1899, p. 39 [extreme age]; Rothschild, Bull. Brit. Orn. Club, XIV. 1904, p. 58 [hybridism].

Gallus giganteus Tegetmeier, Ibis, 1890, p. 308 [domestic Malay fowl].

Gallus gallus Grant, Cat. Birds Brit. Mus., XXII. 1893, p. 344; Bourns and Worcester, Minn. Acad. Nat.
Sci. Acc. Papers, I. 1894, p. 29 [Philippines]; Oates, Jour. Bombay Nat. Hist. Soc., X. 1895, p. 112 [Ruby Mines,
Burma]; Grant, Ibis, 1895, p. 471 [North Luzon]; Hartert, Novitates Zoologicae, III. 1896, p. 164 [Bulekomba,
South Celebes]; Hartert, Novitates Zoologicae, III. 1896, pp. 564, 598 [Lombok]; Grant, Hand-book Game-
birds, IT. 1897, p. 48 [General account] ; Hartert, Novitates Zoologicae, V. 1898, p. 124 [Timor]; Sharpe, Hand-list
of Birds, I. 1899, p. 39; Nehrkorn, Katalog der Eiersammlung, 1899, p. 194; Whitehead, Ibis, 1899, p. 493
[Philippines, from Fuga Is. to Basilan]; Grant, Ibis, 1900, p. 606 [Wei-yiian, South Yunnan]; Bonhote, Proc. Zool.
Soc., 1901, p. 78 [Khota Bharu, Kelantan; Kwala Selama, Perak]; Oates, Cat. Birds’ Eggs Brit. Mus., I. 1gor,
p. 59; Ghigi, Mon. Zool. Ital., XIV. 1903, p. 319 [Generic character]; Finn, Avic. Mag. N. S., III. 1905, p. 102
[Red replaced by yellow and white]; Grant, Ibis, 1906, p. 504 [Mt. Apo, S.E. Mindanao, 8000 feet]; Robinson
Jour. Fed. Malay States Museums, I. 1906, pp. 125, 129 [Malay Peninsula south of Kra]; Robinson, Jour. Fed.
Malay States Museums, II. 1907, p. 67; Robinson, Jour. Fed. Malay States Museums, IIJ. 1908, p. 55 [Pahang
River]; van Oort, Notes Leyden Museum, XXXII. 1910, p. 110 [Western Java]; Stubbs and Rome, Zoologist (4),
XVI. 1912, p. I [prehistoric origin of fowl]; Peters, Jour. Amer. Oriental Soc., XXXIII. 1913, p. 363 [history of
distribution]; Beebe, Zoologica, I. 1914, p. 271; Morgan, Carnegie Insti. of Wash., 1919, Pub. No. 285.

Gallus pseudhermaphroditus Finn, Jour. As. Soc. Bengal, N. S., LX XI, 1902, p. 85 [hen-feathered cock].

Gallus bankiva bankiva Baker, Jour. Bombay Nat. Hist. Soc., XXV. 1917, p. 18 [desc, habits].

Gallus bankiva ferrugineus Baker, Jour. Bombay Nat. Hist. Soc. XXV. 1917, p. 3 [desc. habits].

Gallus ferrugineus ferrugineus Robinson and Kloss, Rec. Indian Mus., XIX. p. 14.

Gallus ferrugineus murght Robinson and Kloss, Rec. Indian Mus., XIX. p. 15.

Gallus ferrugineus bankiva Robinson and Kloss, Rec. Indian Mus., XIX p. 15.

PLATE XLI

CEYLON JUNGLEFOWL | |
Gallus lafayettt Lesson

sk of early dawn the Junglefowl begin to leave their roosts and make their way through
ts of south Ceylon to open glades. Here they send forth their loud crow, chuck-George-

re they battle with each other, armed with sharp spurs, and here they scratch vigorously for

id grubs. ~ Sometimes a dozen birds can be heard crowing in various directions, but after sunrise

| with the coming of heat the birds seek shelter under the dense foliage.

ast aoe

seliae sotte tod oo ‘BuOrgy nt mareey ‘a

sake ie

silo. gactob, ds robs be

PEATE. Alen

“IMOSJATONONP NOTAAO

ve

30h:

3:
2

CEYLON JUNGLEFOWL
Gallus lafayetti Lesson

NAMES.—Specific: /afayettz. English: Ceylon Junglefowl. French: Coq Lafayette. Native: Wali-kukuld.

BRIEF DESCRIPTION.—Male: Hackles of neck and mantle and all but the largest wing-coverts pale straw
to deep golden yellow, with a black or chestnut central stripe; back and rump orange-red, with much of the
visible portion glossy violet ; secondaries purplish, and tail-coverts and tail greenish-blue; greater coverts partly
blue, partly chestnut; a violet patch on the lower neck; remainder of under parts bright orange-red; with the
chestnut stripe becoming dominant on the belly, and black on the lower belly and flanks ; comb red, with a cefitral
yellow patch ; face, throat and wattles red; iris yellow, legs and feet pink or yellow. Female: Crown brown,
nape rufous; mantle, back and wing-coverts brown or greyish, vermiculated with black ; secondaries with irregular
bars of black, buffy-white and chestnut; tail chestnut, mottled with black ; upper breast mottled black and brown,
with white shaft-stripes, which become dominant on the lower breast, with black margins and cross band ; iris
yellow ; legs and feet brownish-yellow.

RANGE.—The Island of Ceylon.

2HE BIRD IN Its HAUNTS

Ir is two o'clock in the morning in the wilderness of South Ceylon, and darkness
and silence claim the whole world. From the porch rafters of a native house my
hammock sways gently, and I twist around and peer out between the meshes. - For a
very long time there is absolute silence. Then far off there comes a series of falsetto
shrieks and wild, inarticulate cries, as a native urges his bullocks along the distant trail.
Now and then the wheels give forth a whining creak, and again silence settles down.
The Cinghalese driver dozes, the wheels are quiet, but still the path of the cart is marked
through the distant darkness. As it jolts past the acacias, a line of slumbering bulbuls
is roused to semi-consciousness and a minute of silvery song. After a phrase or two
each bird sleeps again, and the sweet refrain is taken up by the next in line. Always, so
insistently that our conscious hearing fails to record it, sounds the dull klonk, klonk, of
the bullock bells, those wooden notes of warning to the leopards slinking hungrily along
the opposite side of the thorn scrub.

The first sound of dawn comes from the little house-lizards, as they cease their
night's hunting and retire to safe nooks and crannies. All join in a chorus of sweet,
thin notes, an acceleration of tones as of a marble dropped and settling to rest upon a
silver table. About six o'clock the first bulbul awakens, and soon afterwards the tailor
birds add their vibrations—seef-o ! seet-o ! seet-o! Now with a rush comes the full
matin orchestra, and from all directions sounds the daily medley of croaks, whistles,
warbles and coos. The crows vocally monopolize a certain period of the dusk.
A lazy kwa, kwa, near at hand; then a distant Awa! kwao/ and every black fellow
within hearing at once takes up the variation and bandies it back and forth. Before
sunrise the first Junglefowl challenges chuck / George-Joyce/ and is promptly answered,
and by listening carefully and orienting the various crows and their directions, ten or

twelve may be individually distinguished.
213

214 A MONOGRAPH OF THE PHEASANTS

I clap my hands and receive no response. Again, and a sleepy “Yes, Sahib!”
comes from the rear of the house. When I am dressed, the very lightest of a chofa
hazri is brought, and then I am ready. The Tamil tracker falls in ahead, and with an
occasional flash from my electric light, we make our way slowly along a narrow, winding
animal trail. There is little danger of getting off the trail, for the least touch of the
surrounding vegetation reveals a myriad punishing thorns.

After we have gone a mile or more I send him off to a clump of trees near the water-
pools to see if he can bag a peafowl, and I continue my way in the dim light of early
dawn. With a snarl a civet cat leaps to one side from my path, “civeting” in his
excitement until the whole air is poisoned with the pungent musk.

Junglefowl were now calling close at hand, and as the light increased I began to
efface myself as much as possible. I crawled to the top of a low, rolling bit of turf,
and surprised a second large cat rooting for grubs. The animal slunk away silently.
In front lay an acre or two of jungle-locked glade, bounded with tall euphorbias and
acacias, with a few grassy knolls and a scattering of cock-spur thorn-bushes. As the
sun rose I watched quietly and listened to the booming of the distant breakers.
A tremulous, vibrating thread of sound arose close at hand, and I soon traced this to
a high-backed tortoise, gorgeously marked with a radiating pattern of gold. He
clambered awkwardly into view, waddled onward a few steps, reared up his leathery
head, and with swelling throat sent forth the penetrating trill. At the very moment, as
if Esop’s fables were to be re-enacted here in this Ceylonese wilderness, a great hare
leaped from a thorn bush, cleared a segment of the glade in three bounds, and vanished
on the opposite side. Patiently I waited for more wild creatures, watching a pair of
bee-eaters flycatching from a high acacia. A snort came from behind, and twisting my
head, I was just in time to see a pair of axis deer vanish among the bushes. Then I
was thrilled by a short, broken crow. Now that I heard it so close at hand, no human
words fitted it; there was no George Joyce nor any other human appellation in it, nor
was it at all like the crow of a domestic chanticleer, but a true wilderness voice, the
challenge of the Ceylon Junglefowl.

Soon a form emerged from the shadows, and regardless of leeches and ticks, I
flattened myself on the turf and strained through my glasses. Until the bird reached
the first knoll it looked like a dark kind of pheasant, its head and tail carried low, and
watchful of everything on land and in the air.

I could distinguish every detail of feather, every glance of the eye. There was
naught of the domestic fowl in that carriage. Then he rose to full height and gave his
call. The tail was raised, the head lifted high, the wings a trifle free of the body; and
then the low catch-syllable and the ringing double-note. The cock advanced to a newly
erected turret city of termites, and after a careful survey of every point, began vigorously
scratching. Fora full two minutes he made no attempt to feed, but with strong efforts
jerked the earth backward with his strong claws, working devastation in the insect
metropolis. Then he began picking, picking rapidly, glancing up every few seconds.
He uttered a low chack, chack, in the tone with which, in fowl language, a hen is
summoned. But before any Junglehen could answer, I unintentionally put an end to
further observation. At full tension for many minutes, as I had been, I must have
jumped a foot as a sudden sw-e-e-e-e-ef of wings roared in my very face—sudden and

CEYLON JUNGLEFOWL 215

sharp as rifle shots. Close overhead, so that I was fanned by the disturbed air, a half-
hundred golden plovers dashed like flying bullets. I have seldom been so momentarily
alarmed. Frightened at the plovers or at my sudden motion, a Junglehen ran out from
a bush at my right, cleared another bush without a movement of her wings, and melted
into the dark shadows of the underbush. A glance at the cock showed a fleeing figure,
running with remarkable speed for the nearest shelter.

Only the pair of green bee-eaters remained undisturbed, perched side by side on an
isolated thorn twig; first one, then the other, hurtling upward, somersaulting after an
insect, and dropping back to its mate.

GENERAL DISTRIBUTION

The Ceylon Junglefowl is confined to the island of that name—the oval pendant to
the peninsula of India, measuring roughly one hundred and fifty by two hundred and
fifty miles. It may be said to be generally distributed over the island, being more or less
scattered through the dry jungly districts of the low country, and diffused throughout
the hills of the Southern and Central Provinces. It is rather rare in the damper, more
humid jungles of the maritime portions of the Western Province where Colombo is
situated, and the south-western district, and is not common even in the forests of the
interior. It is occasionally brought into Colombo and Galle by natives, but very seldom
indeed into the former town. In the jungles of the Hewagam and Kayigam Korales its
note is seldom or never heard, but further inland, in the Three and Four Korales,
in Saffragam, and in the Pasdum Korale, one may often listen to its well-known cry.
Likewise in the hilly jungles of the south-western district it is not met with near the sea ;
it is found about Oodogamma, and further up at the base of the ranges it becomes more
plentiful. On the eastern slopes of the Morowak Korale, where a drier climate prevails,
it finds a more congenial home, and along the Wellaway river, and from that eastward
itis numerous. In the maritime portion of the south-east it abounds, delighting in the
dense Euphorbia scrubs along the sea-coast. From this section of the country round
the east coast to the north of the island it is very numerous, and inhabits all the
northern forests, extending down the west side as far south as the Kurunegala district.
In the hills it is resident, and breeds commonly up to about five thousand feet. On the
Nuwara-Elliya plateau and up on the Horton Plains it is sometimes abundant during
the north-east monsoon, coming up from lower down on the hills, and probably, to some
extent, from the low country, to feed on the berries of the wz//oo. It is probable that
some may remain throughout the year in these uplands.

GENERAL ACCOUNT

The -home range of the Ceylon Junglefowl varies greatly. It is undoubtedly
strictly resident throughout the year in most of the dry maritime districts, but in the low
hills and the jungles which fringe the slopes of the higher mountains of the interior it
may show a marked seasonal migration. ‘This is induced, not of course on account of
change of temperature, but owing to the fruiting of certain food plants localized in well-
defined districts, as I have already mentioned in the case of the wz//oo berry. Aside
from this there is little or no seasonal shifting, and although when the breeding season

216 A MONOGRAPH OF THE PHEASANTS

is past the birds are not nearly as concerned in keeping to their own particular beat, yet
they do not appear to wander far. Four birds—two pairs—have been known to inhabit
a circumscribed area of semi-desert scrub for a year and a half, breeding some two
hundred yards apart, and almost never leaving this zone, which was bounded by two
small streams. The young appeared to leave when full grown, and to settle elsewhere.

While I have stalked and watched Ceylon Junglefowl in the forests of the hills,
I became most intimate with the bird in the semi-arid scrub of the south-east coastal
lands. Here the vegetation is eucalyptus, acacias, and mesquite, with introduced cacti
now forming one of the dominant floral features. The region as a whole is park-like,
the thorn bushes and trees forming glades, sometimes dry and sandy, or again with little
pools of water, and a thick carpeting of coarse grass. Into these open spaces the
Junglefowl come to feed and exercise, leaving the denser coverts, in which they roost and
to which they retire during the heat of mid-day.

In such a region, well up on the south-east coast, I found them very abundant in
March. As I have said, it was possible to hear as many as a dozen birds calling, as one
listened in the early morning, and during a walk of several miles in any direction their
numbers did not seem to decrease. As one approached any given bird it would take
alarm and cease crowing, and not infrequently I would catch a momentary glimpse of it
as it dashed away, but a little way farther on, the voice of another bird would come into
range, then others to right and left, and standing quietly one could count as many
individuals as from the first point of listening.

I have never seen more than five birds together at one time, and of these, three were
birds of the year, very evidently the brood of the remaining pair. From all that I have
heard I believe these Junglefowl to be rather unsocial, except that several hens with
their broods seem to flock together at times, the association being, however, somewhat
loose except during the early life of the chicks.

The voice of the Ceylon Junglefowl is very characteristic, and not to be confused
with the notes of any other bird in the island. When softened and modified by distance,
the crow has a reasonably close resemblance to the syllables George Joyce, but when
heard at close quarters the quality which hints of an insistent summons to this
particular gentleman disappears, and a third distinct note becomes audible. The
syllables and the tempo which close attention suggests, is as follows: tsek, ——, ——,
craw, crot! The dashes represent fifths equal in time to the duration of each of the
three notes. The whole utterance is snapped off short at the end and the accent falls
upon the final syllable. There is little variation, and while I have been able to
recognize each one of three or four out of the dozen which I could hear from my
hammock each morning, it was only by such means as a very slight difference in accent
or a slurring over of the middle tone. When suspicious, but not certain of danger, the
cock utters the ¢se# alone. When alarm begins to be felt this changes to clock, clock.
This note, too, is uttered when they hear any unusual noise as they are about to go to
roost. A bird in captivity, which when approached is so timid that it rushes nervously
about looking for some way of escape, usually at the same time utters this sound. The »
call of the cock to the hen is a low chack, chack, sometimes followed by a muffled
tremolo of content. The cry of utter despair when the bird is suddenly flushed by dog or
man, and takes to its wings in extremity of terror, is the usual loud gallinaceous cackle—

PHOTOGRAVURE 33

HOME OF THE *CEYEON JUNGLEFOWL

AtonG the coast of Ceylon, where the eucalyptus, acacias and mesquite abound, these Junglefowl are
abundant. The region is park-like, the thorn-bushes and trees alternating with glades or larger open
plains, dry and sandy, or with pools and grass. Here they must ever be on the watch for leopards and
civet cats, and at night they sleep in dense foliage on lofty limbs, well out from the trunk, so that an

enemy approaching along the branch would at once be detected.

ya

PHOTOGRAVURE PLATE 33

inlOuMUs <Onn aMel

CEYLON JUNGLEFOWL,

[E

CEYLON JUNGLEFOWL 217

a series of harsh, shrill cries which convey their meaning to every creature within earshot.
The challenge is accompanied, especially in the early morning, by a clapping together of
the wings, and this appears to me to stand for a much more serious challenge than when
the crow alone is uttered. I shall speak of this in greater detail under courtship and
fighting. we ;

These Junglefowl are extremely adverse to flying, and invariably try to escape on
foot. I once saw a hen rise and partly fly, partly scale a distance of seventy-five yards
before settling again into the scrub—the longest flight of which I have record. More
fortunate observers say that they are strong on the wing, but all agree that they are
flushed with difficulty. There seems to be no racial weakness of wing power, only the
flat country which they prefer offers no incentive to exercise of this function. I have
already spoken of their swiftness of foot. Their gait when unalarmed is, as in the other
Junglefowl, dignified and alert. The tail is usually held quite low, except during
challenge, courtship, and approach to battle, when it is raised as high as in any domestic
cock.

DAILY ROUND OF LIFE

The food is varied, consisting usually of insects, wild seeds and grain, while
whenever possible the birds will enter cultivated fields and by scratching do considerable
damage to the newly-planted grain. The following list shows the contents of the crops
of eight birds from South Ceylon, shot in March :—

1. Male. Nine large reddish berries with purple pulp ; several scarab beetles.

2. Male. A few green seed-pods ; 5 grass seeds; 23 termites.

3. Male. One hundred and fifty green seed-pods; 7 small mollusks of three
species ; several hundred termite workers.

Male. One large tick.

Male. Many grass seeds still on panicles; 1 termite; 2 small mollusks.

Male. One large tick; many grass seeds; several scarab beetles.

. Female. Twenty-five seed panicles of grass; 1 small beetle:

. Female. In the gizzard were 6 small, smooth, reddish stones, about 6 mm. in

diameter, and a mass of black ants. The crop contained several hundred

termite workers; 1 brown and 2 black beetles; 1 grasshopper; 1 small wood

roach; several brown wood lice and centipedes; a small snail; a small

hemipterid insect, many green seed-pods, and several petals of flowers.

PESUIMON GTN

of

The big ticks in the crops of Numbers 4 and 6 were alive and distended with blood,
which they were apparently drawing from the lining wall of the crop. Termites take
first place in the animal diet of Junglefowl, as in almost all tropical gallinaceous birds.
It is a general belief that the young cannot be reared except on these insects, and in the
birds which I was able to examine, it formed fully four-fifths of the food of the chicks.

I have already mentioned the love which Junglefowl have for the berries of the
nilloo or cone-head plants, a species of Strobclanthes. Whenever this fruits among the
undergrowth of the hill forests it tempts the birds from far and near, and they flock
thither, coming even from the lowlands at the foot of the hills. This maturing of the

seed in quantity takes place at very irregular occasions, sometimes it is said, at intervals
VOL. I FF

218 A MONOGRAPH OF THE PHEASANTS

of four to six years. One observer writes me that the x//oo shrub seeds only once in
eleven years, after which it dies down. But this happens in the various districts in
different years, hence the possibility of the seed being found ripe at much shorter
intervals of time. Whenever this takes place, there is a widespread migration of the
birds. They begin to ascend in December, January, and February, and disappear from
the hill jungles about June.

A very general idea obtains in Ceylon that this fruit has the effect of stupefying the
birds. A probable explanation is the excessive fattening effect of the w2//oo, the birds,
after feeding for some time, becoming veritable feathered balls of obesity. This, in
many cases, would result in the effect described of the birds becoming sluggish and
inert. It is true that the Junglefowl in the Horton Plains and about Nuwara Elliya do
become affected, and are sometimes so fat or intoxicated, as the case may be, that they
may be knocked down with a stick. One writer tells of several cock Junglefowl,
apparently in perfect health, skulking under a bush at one’s approach and allowing
themselves to be caught in the hand. The bronze-winged doves, which are as fond of
the zz/loo as are the Junglefowl, never show any similar symptoms. Botanists know of
no narcotic properties of this plant, however, and it may be that the birds eat some
noxious fungus or other vegetable growth in the woods where the wz//oo thrives. The
Singhalese also believe that the eating of these berries at this season causes blindness
among the Junglefowl. ‘About that season of the year,” says Bligh, “if village fowls
be brought to the hills they rarely escape a serious eye disease, which rapidly spreads
through a given district, and in many cases they become totally blind in two or three
weeks. This is the disease which the Junglefowl evidently catch. A dog of mine
caught a Junglecock with one eye lost, and apparently from this cause.”

The feeding habits of the birds are the surest indications of their presence, when no
vocal utterance gives us a hint of their proximity. This may be purely circumstantial,
as their tracks and evidences of scratching along roads or about the nests of white ants,
or it may be the actual noise of this latter habit. In the dryer north-eastern forests and
the semi-arid coastal regions, the fallen leaves do not decay as soon as in the humid
jungles, and among these, harbouring a multitude of insects and seeds, the birds scratch
vigorously in the same manner as the domestic fowl. The crackling and rustling
of the leaves may be heard at some distance, and forms a very easy way of locating
the birds. This by no means, however, ensures an approach within sight, for after
every effort which they make, the birds listen intently, and are able to detect the very
slightest noise. Many times, after the most careful stalk, I have utterly failed to catch
even a glimpse of the bird. Occasionally, aided by an intervening mound and shield of
leaves, one is able to watch the author of the noise, as, first with one foot, then the
other, he sends leaves and déérzs flying.

This feeding begins in the early morning and continues until the heat begins to be
oppressive. The birds then retire to the shade of some dense-foliaged tree or under-
growth, and either dust themselves, or doze, or strive to rid themselves of ticks and
other parasites. This latter is a very important operation, as I shall have occasion later
on to show.

The jungle and the coastal semi-desert regions swarm with hostile creatures, eager
for prey such as the Junglefowl, but most of these are nocturnal. So before dusk has

CEYLON JUNGLEFOWL 219

settled down the birds go to roost. They seem to choose a perch as high as possible,
but always within the foliage itself, never upon an exposed limb at the top of the tree,
and never close to the main trunk. The first precaution may be to guard against owls
or the buffeting of the elements, the second, which is common to many birds, is
undoubtedly for the purpose of being warned of the approach of any arboreal carnivore
by the vibrating of the branch along which this approach must be made. In the dry
coastal zone, where there are few high trees, the birds may be found only eight or
ten feet above the ground in some acacia. But such a tree is always well isolated, in
two instances to my knowledge standing alone in a glade, and thus fairly well protected
against chance ascent by small carnivores.

In very wet weather the birds retire to trees, usually perching on the lower branches,
waiting for the downpour to be over, but this is only a temporary return to roost, and any
tree convenient to the feeding-ground is acceptable. The birds are extremely attached
to the chosen roosting-place, and the regular roost is resorted to night after night, as is
proved both by the abundance of sign and by the similarity of location as indicated by
the early morning challenge. I found where one Junglefowl—a cock—spent night after
night, and one mid-day I went to the tree and with as little noise and disturbance as
possible sawed off and carried away the limb on which the bird was accustomed to roost,
the sign locating this beyond doubt. The bird spent at least the following two nights
on the nearest adjoining branch, after which I did not visit the place.

These wild fowl are usually found roosting singly, and when a pair or rarely four
birds are associated, they perch on different branches or even adjoining trees. Once I
found where such a quartette, probably two birds of the year with their parents, roosted
on three trees growing close together. Fresh sign each morning showed that all four
perches were occupied.

I have never observed any close association between the Ceylon Junglefowl and such
birds as babblers, as is so common with other related species. All that I have seen have
been alone, feeding or dusting themselves or on the lookout for danger, without any
commensal or other relations with creatures of their environment. Exception may be
made as regards domestic cattle, as the birds are sometimes seen feeding among a herd
of these animals, doubtless benefiting by the insects which are disturbed as the big
herbivores walk slowly along. They come out occasionally and feed with domestic
fowls, although there is no such widespread intimacy as exists between the Red
Junglefowl and the native poultry.

During the zz//oo harvest, doves in large numbers are found feeding in close company
with the Junglefowl, coming up-country until the whole jungle resounds with their
cooing, but there is no actual competition in this case, as there is always a great surplus
of food, and the association is in no way of mutual benefit (unlike the company of the
babblers in other species) and results only from the unusual abundance and localization
of food.

A captive cock Junglefowl made great friends with a Ceylon green parrot which
was kept in the same run, and for months the two birds were inseparable. They fed
together from the same dish, and much of the time the parrot sat contentedly upon the
back of the cock, while at night they roosted side by side.

Enemies they have in plenty: large hawks and snakes, civet cats, all the small

220 A MONOGRAPH OF THE PHEASANTS

carnivores, and especially the mongoose. I have more than once encountered one of
these latter animals stalking, as was I, a Junglefowl busy with its scratching. A
reliable observer told me that once when lying in wait for deer he had seen a mongoose
kill a hen Junglefowl and then make a systematic search for all her brood, four or five
in number, of about two weeks’ old chicks.

The hearing and the-sight of these birds is amazingly acute. They will become
suspicious at the least unusual rustle of an approaching footstep, while leaves and twigs
may be falling in all directions and the wind making sudden rustlings and noises, to
which they pay not the slightest attention. While the wild fowl in early morning come
occasionally out into the roads and trails, or venture into grassy glades, yet it is always
with an eye to cover. I have approached a crowing bird without attempting to conceal
the sound of my footsteps and had it retreat slowly before me for several hundred feet,
crowing from time to time as it went. In such a case one need never expect even the
slightest glimpse of the bird, as it cunningly keeps under cover or dodges behind some
intervening mound or grassy hillock. Of their pietc cues for escape by fleetness of foot
rather than by flight I have already spoken.

Junglefowl learn by experience to distinguish between degrees of danger. In a
place where one could be sure of finding a half-dozen birds along a mile or two of jungle
road in early morning, if I walked on foot the birds would dive into cover as soon as
ever they caught sight of me, even when still a hundred yards away. But going over
the same route in a bullock cart, and sitting under the cover alongside of the driver, the
birds would reluctantly leave off their feeding and walk into the undergrowth only when
we had approached within thirty or forty yards. They had evidently had occasion to
fear neither the slow-moving carts nor the bullocks nor drivers.

In examining the specimens of Junglefowl which I shot day after day there came to
my notice evidence of a factor in the lives of these birds of which we know but little.
This was the mass of ticks which covered much of the combs of the birds. Parasites
such as these probably always cause great discomfort, and doubtless, in many cases, bring
death to a larger number of wild creatures than we realize. It is only by the close
examination of specimens as soon as shot that we can learn anything definite concerning
such phenomena as this, which may be of the utmost importance in limiting distribution,
both local and general, and in affecting the vitality of the individuals. In affected cocks,
the posterior and inferior borders of the comb were usually hidden by a mass of these
parasites. In one bird I took the trouble to count the ticks on one side, and found two
hundred and sixty-three, of all sizes and ages. When the ticks were removed by a wash
of formaline, this part of the comb was seen to be raw, and in one case a secondary
bacterial infection seemed to have set in.

HOME LIFE

The breeding season of this species has the tropical character of elastic limits, and
eggs seem to be laid during almost every month in the year. I doubt if the explanation
of two broods serves to account for this. A more reasonable interpretation might be the
destroying of many nests by enemies and the consequent delay in the nesting. February
to May seems to be the more usual period, but eggs in November and, on the other

CEYLON JUNGLEFOWL 221

hand, in September are far from unknown. In the Batticaloa district it is said that eggs
are found every month except in November, December and January. In the province of
Noa, April, May and June are given as the most favoured months; in the northern
parts of the island nests are found from February to August more often than at other
times.

The Ceylon’ Junglefowl are very pugnacious. The clapping of the wings and the
crowing are the preliminaries to battle, although much of this demonstration may be
considered in the nature of bluff or of warning. One correspondent, Mr. Thomas, writes
me that he considers the clapping as answering the purpose of a long stretch immediately
after awakening, and to support this theory, makes the comment that it is seldom or
never heard except before the first crow or challenge uttered in the early morning.
Referring to a Junglefowl cock which he kept for some time in captivity and from which
he bred a number of hybrid birds, he notes that if unexcited the cock would never clap
his wings after the first call. On hearing another cock, however, he would do so once
or twice, “ probably in the nature of a stretch to feel if he was in fighting trim.” I must
take exception to this view, owing to the facility with which cocks may be called up by
imitating the clapping of the wings, even when an attempted imitation of the crow is
not added. Indeed, by use of the latter challenge alone, it is a much more difficult
matter to entice the challenging cock within shooting distance. I shall give more in
regard to this aspect later. |

It is true that the clapping is usually confined to the first few crows. Whether
this had its origin in the desire to stretch the wings in preparation for a flight to the
ground from the elevated roost, we shall probably never know. To-day, at least, it is
an important and significant part both of the audible challenge and defiance, and actually
more potent than the crow itself to precipitate an encounter.

The cocks inhabiting the same general region, if mated, undoubtedly have a more
or less definite territory over which they hold sway, and into which the advance of a
strange cock is considered an intrusion to be disputed with all their powers of offence,
and if necessary with their life. Cocks settled on adjacent beats or preserves do not
advance to the attack at one another's early morning challenge, whether of clapping,
crowing, or both together. Each, however, reacts at the very first sound, and a few
seconds after the first bird awakens and challenges, every cock within hearing has thrown
off his slumber and returns the defiance with all the force of his lusty lungs. A roving,
unmated cock is detected at once, and he must either make his way silently across
country, or be distinctly looking for trouble if he challenge anywhere in the vicinity of a
settled cock.

The first one or several calls are given from the roost itself. The bird then
usually descends to a low branch, and for a variable period of time devotes itself to
crowing. In this situation the observations of several writers agree perfectly with
my own. The bird walks or struts back and forth along the branch, lowering and
raising his head. When crowing, the head and neck are stretched far upward. Where
the bird has no room for lateral movement it nervously raises first one, then the
other foot, during the intervals of crowing. It seems usually to have some particular
rival within hearing, for whose challenge it listens. As soon as it has finished a
crow, it steps or moves about, but the instant the clap or the crow of the other bird

222 A MONOGRAPH OF THE PHEASANTS

is heard, it freezes into an attitude of concentrated attention, and gives voice again
the moment the rival has ceased.

When their mate has a brood of chicks, I am told they fly down at once to the
ground, where they crow from time to time as they search for food. When thus
engaged even continued and exact imitation of the wing clapping will not entice
them away.

Many days of observation in the dry coastal belt of country show that the
Junglefowl begins to crow at the very first hint of dawn, about 5.15 a.m., and at
5.30, just defore sunrise, their chorus is at its height. Half an hour later it has
begun to diminish, both as to number of birds and frequency of utterance. The
birds are now apparently feeding, but there is no actual cessation until about nine o'clock.

Toward evening a second challenge period occurs, never, however, as enthusiastic
or persistent as that in the morning, and this is stopped by dusk, when the birds
reach their roosting-places. I have heard what was probably a final crow, sounding
forced and sleepy, with but little of defiance in its drawled utterance. On cloudy
days, hardly an hour passes without a crow from one direction or another, and now
and then an emotional outbreak will occur and spread from cock to cock until the
chorus is almost as great as at daybreak. This is of rare occurrence, however, and
lasts but a short time, when silence again ensues.

I have several times found scattered feathers and the entangled tracks of the
feet of Junglefowl, indicating that a battle had recently taken place. Where the
ground is soft such evidence is quite conclusive if no tracks other than those of
the birds themselves are visible. But I have never found a disabled bird nor blood
on the feathers. My most direct evidence was the condition of some of the cock
birds which I shot, their damaged hackles, half-healed scars on the breast and sides,
and torn combs showing that they must have been through some severe contests.
The length and sharpness of the spurs of this species make it probable that such
encounters must frequently end fatally. There are several accounts of fights between
wild Junglefowl and native cocks, the former always coming out victorious.

Ceylon Junglefowl are both monogamous and polygamous, with no apparent
preponderance one way or the other. I have known of two hens undoubtedly mated
to a single cock, and again I have found three different pairs of birds, each pair
feeding and roosting in such isolation that there was little doubt that they were
mated. Three is the largest number of hens reported to me as constantly seen with
a single cock during the breeding season. Captive Junglefowl are always attentive
to hens, feeding them, and paying much court to them. The few times that they
have mated in captivity it has always been with but one out of a number of domestic
hens. When, however, several hybrid hens have been introduced, the wild cock has
mated with them without delay.

The sites of nests are various. I found two nests, one close to the trunk of a
small tree, protected by overhanging vines, and the other far under the drooping
twigs of a cock-spur thorn-bush near the edge of a glade. Other situations on the
ground may be in a clump of grass or bamboo, in the shelter of the turrets of a
white ants’ nest when surrounded by grass, or in the hollowed base of a rotten
stump of a tree. Mr. Thomas has found a nest hidden under an old log, and also

PHOTOGRAVURE 34

MiSs eon EGGS lOro Prin CEYLON JUNGLEFOWL

Tue nest is usually placed on the ground at the foot of a tree, and only the leaves and grass already
present are used as lining. The hen sits very closely, and deserts her eggs only when danger is actually
upon her. Her colours harmonize perfectly with the hues of the dry vegetation, but the eggs, when
exposed, are conspicuous. Five or six is the usual number, and they are as small as the eggs of bantams,
and sometimes slightly spotted.

A native Vedda, one of the aboriginal Cinghalese, was skilful in tracking down the birds and their

nests, and found one of my first nests, which the chicks had just left.

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PHOTOGRAVURE PLATE 34

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CEYLON JUNGLI

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CEYLON JUNGLEFOWL 222

one in the summit of a decayed stump, seven feet from the ground. Even more
surprising are several nests recorded by Parker in the tops of trees, and in deserted
squirrel nests. One was in a well-built nest in a sapling, at thirty feet elevation,
doubtless the deserted home of a crow or hawk, which had been appropriated by the
jungle hen. How the young chicks which emerged from the three eggs reached the
ground in safety is impossible to conceive. There is hardly any doubt but that their
little necks were broken; a tragic penalty imposed by the foolishness of the mother
in choosing such a site. The dry leaves with which the more usual ground sites
are carpeted form a splendid background for the protection of the sitting hen, the
colours of whose plumage harmonize perfectly with the various browns and russets.
She sits very closely and deserts her eggs only when danger is quite upon her.
There is no attempt at actual nest-building, and it is seldom that any lining, other than
that already present, is added. The hollow in which the eggs rest is sometimes a
natural one, sometimes scratched out by the bird, in which case it bears traces of
the excavation, and is usually devoid of any lining, the eggs resting upon the bare
earth or sand.

The number of eggs varies from two to eight. There seems to be no definite
variation as to number in nests of low- or of up-country birds, although there seems
to be a slight percentage of increase as we ascend. Four seems to be the most
usual number. I have found five in one nest, all with a slight peculiarity of pig-
mentation which confirmed their being laid by a single bird. In the zz//oo season on
Horton Plains eight to ten have been found in one nest, but two hens unquestion-
ably united in such cases. We can readily imagine that the unusual altitude and
general environment, as well as the change in food and the close association of such
large numbers of Junglefowl, would doubtless affect, at least to a slight extent, the
regularity of the birds’ habits. The suggestion that the fat condition of the birds
at this period may have conduced to an increased fertility is extremely improbable,
as we know that excess of fat in domestic birds often brings about a cessation of
laying altogether.

The eggs themselves vary widely. They are smooth and glossy ovals, and may
be either of a uniformly unspotted white, or buff, or rich café au lait colour. Or
these ground colours may be present in eggs uniformly spotted with brownish red
or burnt umber. Usually this marking is absent or very sparse at the small end,
while the rest of the shell shows an even, minute speckling, or this reddish pigment
may increase toward the obtuse end until it is all but confluent at that point. The
eggs vary from 33 to 36 mm. in breadth, and from 45 to 48 mm. in length, averaging
35 by 47 mm.

Two or three seems to be the usual number of young hatched, or at least
brought to maturity. Seven is the largest brood recorded as following one hen.
The wings of the chicks develop very rapidly, but for at least two weeks they are
dependent altogether on their marvellous powers of concealment. When alarmed,
the hen utters a loud, sharp clock /! and makes her way slowly away from the source of
danger. I have never seen her, nor have I any record of her pretending to be
injured, but the rapidity of her exit from the scene of concealment of her brood is
adapted to the exigency of the danger. She may half fly, half run if a dog fly at

224 A MONOGRAPH OF THE PHEASANTS

her, or she may run slowly with many backward glances and occasional stops before
she disappears in the underbrush. After the first note of warning she is silent, and
upon the instant of utterance the chicks vanish, either squatting in their tracks or
taking a short run to the shelter of a mass of dead leaves or a fallen branch.
Here they remain until the hen returns. I once sat quietly for an hour, close to
where the chicks had disappeared, without hearing or seeing a sign from any of
them. The hen was in the neighbourhood, but did not show herself. In some
way she knew of my continued presence, but only twice, during the entire period,
did she repeat her warning clock! When the young are three-quarters grown,
there comes a time when they do not seek to escape by squatting or hiding, while
yet their independence is not developed strongly enough to send them off at the
first alarm. More than one observer has noted that when the hen of such a brood
has been shot, the young would continue to run back and forth in the vicinity,
calling loudly, and loath to leave their parent.

The food of the chicks consists chiefly of termites, as well as minute insects
scratched from among the leaves.

In unfrequented districts, where the natives do not trap or molest the birds,
the hens with their broods show less fear of the approach of man, and even when
the cocks are present, they may be approached within gunshot. The cock is always
the first to leave, the hen and her brood following when one’s steps have brought the
birds a few yards nearer. During the months when the young Junglefowl remain
with the parents, both of the latter are found together, although the cock never
incubates or approaches the nest closely. He remains in the vicinity, however, and
crows at intervals throughout the period of incubation. Several people told me the hen
cackled or called when she either deposits an egg or leaves her nest to feed, but of
this I have no direct confirmation, and place no reliance upon it. She goes to the cock,
however, at such a time, and the two usually feed together, after which she returns
to her eggs.

RELATION TO MAN

The Junglefowl seems to have entered quite deeply into the life of the Singhalese
in past times, for both the wild bird, the domestic descendants of the Red Junglefowl
and the peafowl are found as ornaments in graves and elsewhere. We find these
represented in all periods, that of bronze work, about five hundred years ago; later,
when brass became the general medium, during the eighteenth and nineteenth
centuries, and in the comparatively modern and recent period of pottery. The old
cocoanut-oil lamps, used in the temples during the bronze and brass periods, are
- more often than otherwise surmounted by figures of these birds.

To-day the Junglefowl seems to be holding its own in many parts of Ceylon,
in spite of more or less trapping throughout the year. It adapts itself in a measure
to deforestation, and is able to live in coffee plantations or in scrub of reasonable
density in the absence of jungle. In the hill districts, if anywhere, the bird is
becoming scarcer. A few instances have been brought to notice where these birds
systematically injure newly-planted crops, so that they had to be shot or frightened

PHOTOGRAVURE 35

CEYLON JUNGLEFOWL

In the more arid places Junglefowl often suffer from the attacks of ticks, which attach themselves to
their comb in large numbers, and although they will sometimes preen one another’s plumage, they seem
never to attempt to rid their companions of these pests.

The eggs vary from white to a rich cream colour. Usually they are unspotted, but occasionally eggs
will be found which are thickly covered with dots and spots of brownish red.

In trampled places in the thorn-brush, feathers are sometimes found, showing where severe battles
have taken place between rival cocks. But this photograph reveals a real tragedy. A cock Junglefowl
had been feeding on the insects which it had scratched from a nest of termites, when a civet cat or
similar enemy had pounced upon the bird, plucked out many of the larger wing feathers and carried

it away.

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PHOTOGRAVURE PLATE 39

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CEYLON JUNGLEFOWL 225

away, but as a whole these birds may be considered a negligible quantity in their
power to injure man’s cultivations.

The Junglefowl, from the variety of its food, has caused some sportsmen to
be enthusiastic over the delicacy of its flesh, while others describe it as tough, tasteless,
and scarcely edible. It seems, however, to be hardly worth shooting for the table
when doves can be procured. At any rate, that was my experience in south Ceylon,
and I have heard from many Europeans in various parts of the island that only
the natives find them worth eating. This fact, together with the religious aversion
of the Buddhists to taking life, are points in favour of the continued existence
of the Junglefowl for many years to come. While their feathers are sometimes
used in tying fishing flies, and occasionally exported for millinery purposes, these
uses do not threaten their extinction.

In the recent game laws, the Junglefowl does not seem to be included in the
term ‘‘game,’ which embraces four species of deer, besides ‘‘ peafowl, grey partridge,
painted partridge and Ceylon spur-fowl.” We find them mentioned, however, in a
clause which reads: “Shooting at night and netting game or Junglefowl, unless
trespassing upon cultivated land, is prohibited under a penalty of one hundred
rupees, or three months, or both.” So we find the birds totally unprotected by
law, and yet holding their own in most places.

The natives of Ceylon have many ways of trapping Junglefowl, both by snares
and by spring-traps. The most effective method in use in the hill jungles is the
construction of a miniature fence of small sticks, nine or ten inches high and fifteen
or twenty yards long. Several gaps are left near the centre, six or eight inches
wide. For several days food is scattered about on both sides of the fence and
around the gaps. Then, when the birds become accustomed to feeding regularly
hair nooses, attached by a catch to a bent spring, are cae over the gaps, and
several birds are certain to be caught.

Another native method of trapping is by the use of a net, about six feet high
and twenty yards long, which is stretched in a semicircle across a gully. When it
is in position, a score or more Singhalese form a line and work down toward the
gully through the jungle, noisily driving all the terrestrial birds and animals ahead
of them. In this way many species of small mammals and birds, including Junglefowl,
are entangled and captured.

The most widespread way of shooting Junglefowl, practised both by Europeans,
Singhalese and Tamils, is to attract them by imitating the sound of the clapping
of the wings which often precedes the vocal challenge. When the sportsman has
made his way into a good position as close as possible to several crowing birds
without having aroused their suspicions, he crouches low on the ground and begins
to call. If a native, the hollow sound of the clapping wing, wop! woh! wop! wop!
is made by striking the thigh with the hollowed hand. Europeans produce an
imitation of equal excellence by making a pad of a pocket handkerchief, holding it
in one hand, and striking it with the palm of the other, both hands being hollowed.
It is only when the cock thinks its preserve is being encroached upon, or if a
wandering, unmated bird be near, that it will heed the challenge. But when once

aroused, its anger increases as it approaches, and it is apt to burst into sight without
VOL, II GG

226 A MONOGRAPH OF THE PHEASANTS

warning. Only a moment is permitted to the sportsman, as the keen sight of the bird
takes in the situation at a glance, and like a flash it turns and vanishes beyond recall.

CAPTIVITY

Even in Ceylon, Junglefowl do not thrive well in captivity, and there are only a
few instances of birds having been reared from the nest and kept for years in good
health and feather. They must have plenty of cover as well as a chance to be in the
sunlight as much as they choose, together with an abundance of insect food. If captured
adult, the first is the critical week, as on the slightest provocation the birds dash them-
selves again and again against the top of their run. A domestic chicken (not a
full-grown fowl) placed in the run with them is a most effectual help in calming the
wild-caught birds and inducing them to eat. Low country and medium elevations
offer the best climates for keeping these birds, as captive Junglefowl feel the cold over
four thousand feet rather keenly, and besides, insect life is less abundant at the higher
altitudes.

It is seldom that any details are available in connection with captive birds, newly
caught in their native land, so the following notes! seem to me of particular interest in
showing the unexpected psychological acceptance of captivity on the part of some of the
wild Ceylon Junglefowl.

“During the experiments carried on by members of the Ceylon Poultry Club with
the Ceylon Junglefowl, several interesting incidents occurred which are worth recording.
The following notes record some of these incidents. They occurred mainly in the experi-
mental run put up by Mr. Clement Johnson, who was the only experimenter who
succeeded in producing hybrids, some thirty chicks in all, from a mating of a jungle-
cock with a domestic hen.

‘At one time he secured two jungle hens, which he placed in a large covered-in
run with a jungle cock and two domestic hens. These hens tamed down wonderfully
quickly, and were great friends with the cock. After a time one jungle hen developed
gapes, so it was caught and set at liberty. Writing of this hen, Mr. Johnson says:
‘The jungle hen that I released interests me greatly. Its one object is to get back into
the pen. It walks round and round outside or perches on the top. Any sudden or
unusual noise alarms it, and it flies or runs into cover. On the other hand, you can
approach within a few yards’ length of it, when it just calmly walks out of your path like
a very tame domestic fowl, no hurry or flurry about it at all. It avoids fowls that cross
its path. Since its release the jungle cock inside the run calls more or less all day long,
and is undoubtedly distressed at seeing this hen at liberty outside his run. He gets
frantic when she makes a run and disappears from view. I will give her a week or ten
days’ liberty, and then drive her back into the run again.’

“This hen, after haunting the scene of her captivity for many days, disappeared one

night. It is presumed that she was destroyed by one of the jungle cats that patrol the
- neighbourhood. .

“A little later the second jungle hen developed chicken-pox. Fearing that infection

would spread, Mr. Johnson had this hen also caught and liberated, but she likewise

1 Thomas, “Spolia Zeylanica,” VII, 1911, p. 159.

CEYLON JUNGLEFOWL 227

refused to depart from the scene of her captivity. But as her removal was deemed
necessary, she was caught and taken across a ravine and liberated in the jungle some
quarter of a mile away. Next day, however, she turned up again, trying to get into the
run. She was caught a second time and taken further afield and liberated. After this,
as she did not return, it was thought she had gone for good. However, some days
later she was back again. She was now quite cured of the chicken-pox, having
evidently cured herself in the jungle, either by eating some herb or by living in
surroundings natural to her. After this she continued to live in the garden outside
of the run, and used to walk about with some of the young hybrids which Mr. Johnson
had bred, roosting at night in the branches of a tree along with the hybrids. The fact
of consorting with the wild hen rendered these hybrids a little less tame than usual.
This hen eventually made a nest in the garden and laid three eggs and sat on them.
As she was running with immature hybrid cockerels, and had always rejected their
advances, these eggs were not expected to be fertile. They were, however, removed
from the nest and set under a domestic hen, and, as expected, all proved infertile. There
is little doubt that, if Mr. Johnson had not left for England at this period, this jungle
hen would shortly have produced fertile eggs by running with the more matured hybrid
cockerel in the garden, and he would have produced the unique cross of hybrid cock and
jungle hen.

“When her own eggs were removed from the jungle hen’s nest, they were replaced
by three eggs laid by the domestic hen running with the jungle cock, and these she
incubated. Just at the time of hatching one egg got broken in the nest; it was an
addled one. This attracted thousands of ants to the nest, which not only drove off the
sitting hen, but killed and partly devoured the two chicks just hatched from the other
two eggs. It would have been a strange sight to have had a jungle hen strutting about
the garden with some hybrid chicks.

“Jungle hens have never bred in captivity. Mr. Johnson’s opinion is that this hen
would never have bred with the hybrid cock or any other cock if it had been confined
within wire-netting walls.

“On the other hand, the late Mr. Young of Udabagie had two ate hens in
captivity for considerably more than one year, and they were mating up with a domestic
cock, and Mr. Young was very hopeful of producing hybrids from this mating, when
his tragic death by lightning put a stop to the experiment.

“Tt was just at this period that Mr. Johnson left Ceylon for England. Before he
left this jungle hen was enticed into the run and caught, and with the jungle cock
was sent to Mr. G. C. Bliss at Atagalla. The cock did not take kindly to the close con-
finement necessary while his big run was being put up in the new locality, and began
to sicken; when turned into the big run he did not recover, so he was let out and .
given his liberty. At night time, however, he returned to this run (in which the
jungle hen had been also placed) and was allowed to go in. Next morning he was
found dead. Thus, after a captivity of sixteen months, ended the life of a most
interesting bird—the progenitor of all the thirty hybrids that were produced during
the experiments. This jungle cock mated only with the one domestic hen, and would
have nothing to do with any other hen, in fact he drove them all away. Even when his
own particular hen had been removed for a month owing to illness, he still would have

228 A MONOGRAPH OF THE PHEASANTS

nothing to do with any other. After the death of this cock the jungle hen became
excited and wild, so she was given her liberty, and flew away to be heard of no more.
The history of this hen is surely unique. She had lived either in the experimental run
or in the garden just outside it for eighteen months.”

The Ceylon Junglefowl has bred at least once in the London Zoological Gardens,
in the year 1874, but it is unusual for the hens even to lay in captivity, and this record
is almost unique. Of eleven individuals which have been confined in those Gardens,
one lived for three years, the average length of life of all being a year and a half.

DETAILED DESCRIPTION

ApvuLT Mavre.—Ear-coverts greyish-white; feathers on the top of the head and
occiput deep rufous orange; hackles on the back and sides of the neck, upper mantle
and lesser wing-coverts straw yellow or golden orange, with a conspicuous black shaft-
stripe. Posterior part of the mantle, scapulars, median wing-coverts and lateral feathers
of the lower back and rump orange-red, this colour being confined to the wide, visible,
disintegrated fringe, the remaining part of the feather being maroon or chestnut. On
the central feathers of the lower back and rump the deep orange-red is confined to a
narrow fringe, the rest of the visible part being richly iridescent violet, shading into
bluish posteriorly. On the upper tail-coverts this in turn changes into a bluish-green
gloss, which characterizes the central rectrices, and the outer web of the succeeding two
or three pairs, the remainder of the tail being dull black. The primaries are dull
brown; the secondaries and their greater coverts black, with considerable purplish-blue
gloss. The inner coverts are sometimes margined on the outer web with dark
chestnut.

A conspicuous patch of feathers at the edge of the bare throat rich, iridescent violet
or purplish-blue; chest, breast and sides orange-red like the lower mantle and upper
back, the central maroon streak less prominent; belly chestnut, mottled and tipped with
black; flanks; thighs and under tail-coverts chiefly black, the latter glossed with
greenish.

Bill horny brown, anterior half of lower mandible yellow; iris straw yellow; comb
bright purplish red, with a large wing-shaped, central, yellow spot. This spot is very
bright and arises abruptly just over the eye, but shades off at its margin into the colour
of the comb. When the comb measures about 80 x 30mm., the yellow spot will cover
an approximate area of 50 x 25mm. In some birds the yellow is so abundant that the
general aspect is of a yellow comb with a red border.

Facial skin, throat and wattles usually clear bright red. Occasionally the wattles
will show a central yellowish spot, and in these individuals the throat too is yellowish.
Legs and feet pink in full-coloured birds, or in others mottled with white or yellowish,
or wholly waxy yellow; claws dark brown; spurs black or yellowish at base; weight
2} to 24 Ibs.

Culmen from nostril, 16mm.; length, 660 to 725; wing, 230; tail, 340; tarsus, 80;
middle toe and claw, 60; spurs long and curved, 37 mm.

CEYLON JUNGLEFOWL 229

VARIATION AND MOULT

In Ceylon one hears a great deal about the difference in the species of Junglefowl
which inhabits the hills and mountains of the interior, or of variations in birds
from the several provinces. The following quotation from Price will show what I
mean by this. ‘It may interest you to learn that there is, or was until lately, a white
jungle fowl hen to be seen in company with other ordinary jungle fowl at a place on the
Anuradhapura-Puttalam road, about fourteen miles from Anuradhapura. When I saw
it first I mistook it for a paddy bird, and it would have escaped identification as a
jungle fowl if it had not been in the company of another pair. I got quite close to it
and saw that it was undoubtedly a jungle hen, almost pure white, except for a few grey
feathers on the neck and head. Mr. Vigors, the Government Agent, saw the same bird
shortly afterwards at the same place. At the time I saw this bird I was not aware that
such a variation in colour was so uncommon, or I might have made an exception to my
general principle of sparing curiosities. Both Mr. Vigors and I thought it would be
better to leave it where it was, to see if it would have any effect on the colour of the
other birds about that locality. Down in the south of Sabaragamuwa and in Humban-
tota there is an inclination for the birds to be brown, especially the hens, which are very
dark, in many cases with black spots on tail and”° wings. The tendency in the open
scrub country of the Northern Province and in Tamankaduwa, in the North-Central
Province, is for the birds to be red in colour; the cocks, of course, always have a lot of
red about them, but it is more brilliant and lighter up north, and exactly to the same
extent is the hen light in colour. I suppose it is the same influence which makes the
wandaroo monkey almost black down south, while he is almost white up north.”

I lost no opportunity of obtaining direct and first-hand evidence of this, and came
to the final conclusion that if any such distinction actually exists, only series of several
hundred birds with exact data of age, locality and season could prove it. I was told
that the voice of coastal birds differed from those of the interior, which I proved beyond
doubt to be an error. The plumage of the low country birds was said to be lighter and
paler, and the birds somewhat larger, with a greater development of comb and wattles.
The series which I was able to collect in the coast districts of southern Ceylon ran the
full gamut of these variations, and the same is true of the birds which I obtained from
high levels. I believe that all altitudinal or regional variation is lost in the very
considerable individual variation.

The regular annual moult takes place usually in September and October.

ApDULT FEMALE.—Crown dark brown, nape rufescent, the feathers sometimes with
distinct black margins. Neck and mantle dark orange or rufescent brown, sometimes
with indistinct black mottling, or again, with the black arranged in two wide, irregular,
concentric bands. These feathers are sometimes fringed with quite bright golden-brown.
Remainder of upper parts and wing-coverts finely mottled reddish brown and black,
usually with a hair-like, whitish shaft-streak. These streaks are always absent from the
lower back and rump, which are of a deeper rufous tone. Primaries dark brown with a
series of pale buff spots on the outer web; secondaries and their coverts black with
mottled buff cross-bars on the outer webs, the black areas often with an irregular basal

230 A MONOGRAPH OF THE PHEASANTS

wedge of chestnut springing from the shaft. Inner web plain brownish-black or mottled
with buff. The larger black areas are often glossed with purplish-blue.

Tail rich rufous, mottled with black. The black on the central feathers sometimes
takes the form of two broad longitudinal bars on the inner web and a succession of spots
on the outer web, lengthening basally into irregular transverse bars. These black areas,
when well developed, are glossed with purplish-green. The remaining rectrices may be
evenly mottled, or may show black spots or bars of similar character.

Face scantily clothed with pale brown ; chin and throat with whitish feathers ; upper
breast mottled black and rufous-brown, with broad buff shaft-stripes. These increase in
area and become white on the lower breast. The remaining under plumage is dominately
white, with a fringe and a wide cross-bar of black, the latter margined basally with buff.
Under tail-coverts like tail. Comb, wattles and spurs rudimentary.

Upper mandible brownish-black, yellowish toward the tip; under mandible chiefly
yellow ; iris lemon yellow; face, comb and wattles purplish-red ; legs and feet pinkish to
waxy or brownish yellow. Weight 14 to 12 lbs.

Bill from nostril 15 mm. ; length, 400 to 450; wing, 190; tail, 125; tarsus, 65; middle
toe and claw, 53 mm.

VARIATION

Hardly any two hens are alike in coloration, and besides this, a bird in well-worn
plumage presents a very different appearance from one which has recently completed its
moult. The rufous and buff in the plumage change by wear and abrasion to greyish
white, this alteration being particularly noticeable in the wing-bars. These areas of
lighter pigmentation seem to offer much less resistance to wear and tear, and at the end
of the season the light bars in the wing will have lost almost all their barbules and
appear quite white, while the intervening areas of black show but little destruction to the
components of their portion of the vane. I have shot three females on successive days
within an area of some few hundred yards, all birds which had recently bred, which
exhibited in their plumage the extremes of abundance and absence of black patches in
the tail feathers, and with mantle feathers of very different shades and patterns.

Cuick IN Down.—Triangular crown patch narrowly margined with black, chestnut,
extending down the neck and spreading over the upper plumage. A wide black line
begins in front of and above the eye and extends back as a superciliary across the
ear-coverts and on until it joins the nuchal black. Remainder of the top of the head
buff, becoming whiter on the face and pure white on the chin and throat. The dark
upper down is bordered laterally on the back and rump by a black-margined pale buff
stripe ; breast tinged with reddish-brown; remaining under parts creamy white. Upper
mandible black, with the tip and all of the lower mandible yellow. Legs and feet
yellowish. Bill from nostril, 6 mm.; length, 90; wing, 50; tarsus, 22; middle toe and
claw, 20 mm. .

JUVENILE PLumMAGE.—This state of plumage of the young birds is singularly like
that of the adult female, differing chiefly in the ventral white being less pure, more mixed
with the browns, and the rufous tending rather to greyish or sandy.

The down of the head and neck persists, as usual, long after the entire body is

CEYLON JUNGLEFOWL 231

clothed in contour plumage. The mantle is an indefinite mixture of sandy buff, reddish
brown and black mottling, with the shaft areas of the lighter tint. The remainder of the
dorsal plumage is much disintegrated, with terminal brownish mottling. The scapulars
and many of the coverts are tipped with buffy white. The primaries are dark brown,
with sandy or whitish mottling on the outer webs; the secondaries are heavily banded,
as in the adult female, but with purer whites, differentiated from the chestnut bands
which separate the buffy white from the black bands. Toward the tip of the inner
secondaries and tertiaries, the black becomes restricted to round marginal ocelli on each
web, and this ocellated appearance is even more pronounced on the larger wing-coverts,
where the black eye-like spots are framed in buffy-white. There is an unusual amount
of variation in this plumage, however, extremes being found both in lowland and upland
birds, in those from the arid south of the island and the more humid north. For
example, the tail-feathers in the juvenile plumage of either sex may be rich chestnut,
evenly mottled and vermiculated with black, or each web may have a longitudinal black
band down the centre, or again, there may be as many as six black cross bars, equalling
the intervening chestnut spaces.

The chin and throat are pure white; the breast, like the mantle, with the pale buffy
shaft-stripes much clearer and more conspicuous. The breast and belly are whitish,
the brown and buff mottlings dying out into an indefinite brownish tinge. The bill, legs
and feet are yellowish, the upper mandible sometimes quite dark.

First AnNuAL Movutt.—A series of birds taken when this moult is nearly or
quite completed shows no two individuals alike. The females, having comparatively
little change of colour and pattern to undergo, usually moult into quite fully adult dress,
but the males range all the way from a richly chestnut bird mottled with black, to
individuals with but little remaining traces of juvenile pigment. The barred, secondaries
are always lost, but the feathers are usually heavily mottled with brown and rufous ;
while the mantle shows all stages between rich chestnut, olive-yellow and the straw
yellow of the adult. From the birds which I have seen, the upland birds seem to moult
into cleaner, clearer colours, the lowland individuals usually retaining more of the
immature, generalized patterns.

The succeeding moult takes the bird into the complete adult plumage.

HYBRIDISM

Several Englishmen in Ceylon, notably Messrs. Thomas and Johnson, have gone
to great pains to gather accurate data concerning the reputed sterility of the hybrids
between the Ceylon Junglefowl and domestic poultry. They hoped to bring proof that
the former had played an actual part in the origin of our barnyard fowls. This, I think,
is not necessary. The red junglefowl seems to supply all the requirements necessary
-as the direct ancestor of our domestic stock, and these pypuidizatien experiments are of
purely scientific rather than of atavistic interest.

The percentage of fertility of the hybrids when bred zuzter se has been exceedingly
low. Only one egg in each lot laid has been fertile, and of all the eggs produced (about
one hundred in the course of a year) only two chicks have been hatched, these living

232 A MONOGRAPH OF THE PHEASANTS

twelve and eighteen days respectively. The following summary gives us the general
results of the Ceylon experiments :—
The experiments have shown that

1, The hybrids are not always sterile when bred zyer se.

2. The hybrids are not sterile when bred back to the domestic parent (ze. hybrid
cock with domestic hen).

3. There is some indication that the hybrids may sometimes be fertile when mated
back to the jungle parent (¢.e. Junglecock and hybrid hen).

4. Jungle hens have never laid in captivity.

While, without much difficulty, it is possible then to obtain hybrids from a cross
between the wild Ceylon Junglecock and a domestic hen, yet these hybrids bred
together are, to all intents and purposes, sterile, while the hybrids of the wild red
junglefowl are as fertile as either of the parents.

I believe that very rarely the wild Junglecock crosses voluntarily with the native
poultry, and here and there hybrids resulting from such interbreeding are to be found
in the native villages. But that this is a frequent or widespread occurrence seems to
me an error based on groundless assertions of the natives. Many so-called hybrids were
brought to me and proved to be nothing but domestic birds.

EARLY HISTORY

The first mention of this species in scientific literature seems to be in the year 1831,
in Lesson’s 7vazté a’ Ornithologie, where he names the bird Cog Lafayette, “gut,” as
Des Murs says, ‘“‘zudique suffisamment influence des préoccupations politiques du
moment.’ Lesson has this to say of it: “ Gallus Lafayetu: Cog sauvage de Ceylan,
Gal. de Paris. Deux petits barbillons a la mandibule inférieure; plumes de la
collevette effilées, gaune dor, avec une flamme brune au centre, le thorax recouvert de
longues plumes étroites vouge doré, flammées de now, bas-ventre now, queue courte,
brune; un demi-collier violet sous la peau nue du devant du cou. Habite Ceylan.
(Leschenault).” .

About the same year, Gray, in the volume of illustrations which he gives from the
collection of Major-General Hardwicke, figures (pl. 43, Fig. 2) the female. This he calls
Lord Stanley's Hen, Gal/us stanleyi, Gray, but gives no text of explanation. In 1849
Des Murs in his /uconographie Ornithologique gives an excellent plate of the cock. He
says this bird had been donated to the Paris Museum in 1822 by Leschenault, who
brought it from Ceylon, and through some curious mischance had passed unnoticed for
nine years until discovered by Lesson. Des Murs, of course, did not recognize the hen
in Gray’s plate as the female of this Junglefowl, so he ends his account: ‘“‘// est
a vegvetter que lon ne connaisse pas la femelle du G. Lafayeti.” .

The earliest account of the general habits—a good one, too—is that of Layard, in
the Annals and Magazine of Natural History for 1854.

CEYLON JUNGLEFOWL 233

SYNONYMY

Gallus lafayetit Lesson, Traité d’Orn. 1831, p. 491; Gray, Gen. B. III. 1845, p. 499; id. List Gallinae Brit.
Mus. 1867, p. 39.

Gallus stanleyi J. E. Gray, III. Ind. Zool. I. 1830-32, pl. 43, fig. 1; Blyth, Cat. Mus. As. Soc. 1849, p. 243;
Layard, Ann. Mag. N. H. (2), XIV. 1854, p. 62; Sclater, List of Phas. 1863, p. 10; Blyth, Ibis, 1867, pp. 154, 307;
Gray, Hand-list of Birds, II. 1870, p. 261; Holdsw., Proc. Zool. Soc. 1872, p. 468; Legge, Ibis, 1875, p. 400; Hume
& Marshall, Game-birds India, I. 1878, pl.; Sclater, Animals in Gardens Zool. Soc. London, 1883, p. 488 ; Mitchell,
Proc. Zool. Soc. 1911, p. 522 [viability in captivity].

Gallus lafayettit Des Murs, Icon. Orn. 1849, pl. 18; Elliot, Mon. Phas. II. 1872, pl. 33; Hume, N. & E. Ind.
Birds, 1873, p. 530; Hume & Marshall, Game-birds India, I. 1878, p. 241, pl.; Hume, Stray Feathers, VII. 1878, p. 429;
Legge, Birds Ceylon, III. 1880, p. 736, pl. ; Oates, ed. Hume’s Nests and Eggs, III. 1890, p. 422; Blanford, Fauna
Brit. Ind. Birds, IV. 1898, p. 77 ; Thomas, Spolia Zeylonica, IV. 1907, p. 19 [hybridization] ; Ryan, Spolia Zeylonica,
V. 1907, p. 16 (Dimbula) ; Finn, Game-birds India, 1911, p. 19; Henry, Spolia Zeylonica, VII. 1912, p. 59
[Tamblegam ; habits].

Gallus lafayette Grant, Catalogue Game-birds Brit. Mus. XXII. 1893, p. 348; Butler, Jour. Bombay Nat.
Hist. Soc. X. 1896, p. 311 (Tungala, Wva; habits); Grant, Hand-book Game-birds, II. 1897, p. 53; Oates,
Game-birds India, I. 1898, p. 375; Lewis, Ibis, 1898, pp. 339, 550 [Sabaragamuwa Province]; Ghigi, Mon. Zool.
Ital. XIV. 1903, p. 319 (generic characters) ; Storey, Hunting and Shooting in Ceylon, 1907, p. 69.

Gallus lafayetted Sharpe, Hand-list of Birds, I. 1899, p. 39; Oates, Cat. Eggs Brit. Mus. I. 1901, p. 60, and
pl. VII. fig. 8.

Ceylon Junglefowl Thomas, Spolia Zeylonica, VII. 1911, p. 159 [notes on captivity].

VOL, II HH

GREY JUNGLEFOWL
Gallus sonnerati Temminck

NAMES.—Specific: sonnerati, after M. Sonnerat who first described the cock and hen. English: Grey or
Sonnerat’s Junglefowl. French: Cog Sonnerat. German: Sonnerat-huhn. Native: Komrd, Mt. Abu; Jungle
Murght (Hindustani) Central India; Ran-kobada [male], Ran-kobadi [female] {Mahrati) Sahyadri Range; Parda

Komrt (Gondhi) Chanda District; Kombadi, Deccan; Adanikode (Telegu); Katu-koli (Tamil); Kol, Kad-koli
(Canarese), Mysore.

BRIEF DESCRIPTION.—Male: Hackles on neck and mantle fringed with grey, with black centres along
which are several yellow or whitish spots like sealing-wax; median and greater wing-coverts very similar; back,
lesser coverts and under parts brown or black, with a greater or less amount of grey, forming a fringe and central
stripe ; rump and tail-coverts purplish, tail glossy green. Female: Upper mantle rusty, with broad, buffy-white
shaft-stripe ; upper parts brown, mottled with black, the light shaft-stripe disappearing posteriorly ; under parts
predominantly white, fringed and sparsely mottled with black.

RANGE.—Western, Central and Southern India.

GENERAL DISTRIBUTION

Tuis Junglefowl is generally distributed in Southern India. Its northern boundary
may roughly be defined thus: the valley of the Godavari from its mouth on the eastern
coast up to the northern tributary, the Indravati. After continuing some distance up
the valley of this latter river, the northern line of distribution leads irregularly north-
westward through Bhandara, Seoni and Pachmarhi to the westward-flowing Nerbudda.
About one hundred and twenty-five miles from the Arabian Sea, the birds cross the
river and extend in a narrow finger northwards through the Vindhya and the Mahikanta
Mountains to Mt. Abu, and thence are found in few and fewer numbers some
distance farther north among the Aravalli Hills the northernmost record being about
26° ANE at.

Southward its haunts are bounded only by the sea, the fifty miles of water between
the mainland and Ceylon forming a barrier which divides two distinct species of
Junglefowl. Like the other birds of its genus, the Grey Junglefowl is not a lover
of flat, open, cultivated tracts. Hence within the limits of Southern India there are
large areas from which the bird is wholly absent. In jungle, or in hilly, broken
country they are very widely distributed, with centres of abundance in the larger ranges,
such as the Western Ghats, the Satpuras, the Nilgiris, Puhneys, Shervaroys and
Anamalais.

GENERAL ACCOUNT

The home range of the Grey Junglefowl seems to be rather limited, and in the
few cases reported to me of an individual bird being observed for any length of time,
the wanderings have been very circumscribed. Davidson writes: ‘“‘They are very

punctual in their appearance at particular feeding-grounds, and when one or more are
234

PLATE XLII

GREY JUNGLEFOWL

Gallus sonnerate Temminck

Tue Junglefowl of southern India range from the sea-shore to an altitude of five thousand feet.
Mated pairs appear to remain together throughout the year. At the breeding season the birds retreat to
distant parts of the jungle, but at other times they often feed openly in trails leading out of villages,
especially where cattle are pastured. The spots on the hackles of the cock are like drops of sealing-wax,

and unlike any character found in other Junglefowl.

“

WK aTAUS

AonimintaT Wooten,

rle-g98 9

dost basavodi avit lo obutisls
ot jsorter ebtid 9d) noesse gnibesid ont GA;
259

slliv lof ive yaibsal) elie si, ylasgo. bos

q
xpw-unilsos to -eqorfioaih ote: alo

PLATE XLII.

GREY JUNGLEFOWL.

GREY JUNGLEFOWL 235

met with in any particular spot, they are certain, if not disturbed in the interim, to be
found again in the same place at about the same hour the next or any subsequent day
on which they may be looked for. There was one particularly fine and remarkably shy
and cunning old cock that frequented an open glade in the forest (above the Government
Cinchona Plantations at Neddivuttum) in the sovuzug, whereas in the evening he always
came into the plantation and wandered about under the cinchona trees, and along the
plantation roads. He never, to my knowledge—and I must have seen him fifty times
at least—came into the plantation in the morning, or into the glade in the evening.
There was no doubt as to this being the same bird that frequented the two places
(nearly a quarter of a mile distant), for he was the largest, handsomest, and to judge
from his spurs, the oldest cock I ever saw. ‘I loved that cock as a brother I did, and
at last J circumvented and shot him.” They are occasionally found among evergreen
forests in the mountains, and near the edge of the plains I have surprised them among
semi-arid vegetation, cacti and euphorbias. They are found in greater numbers in light
deciduous jungles, but their favourite haunts are hillsides covered with bamboo jungle,
whether small and thin in character or among thick, giant stems.

Where the jungle reaches the shore, these birds may be found within sound of
the breakers at sea-level, and from here they range upward to four and five thousand
feet. Indeed on Dodabetta Mountain, in the Nilgiri Hills, individuals have been seen
at seven thousand feet elevation. In spite of this very considerable altitudinal range,
there is no seasonal migration, owing to the tropical latitude, there being not sufficient
extremes of temperature to induce any change of locality. There is, however, in many
places a very marked seasonal wandering in search of food, or rather in search of some
variety of food which becomes locally abundant at a certain period of the year. Thus
we read that while usually the birds are evenly scattered over a given district, when a
certain tract of bamboo comes into seed, the Junglefowl will collect in large numbers,
again dispersing when the food is consumed. Hume quotes an authority who says:
“T remember on one occasion when the undergrowth of the S/o/as about Pykarra
(which consists almost entirely of S¢vodclanthes sf.) seeded, the Junglefowl congregated
there in the greatest numbers, I mean by hundreds, and were excessively numerous for
more than a fortnight, when they gradually dispersed, owing, I believe, not so much
to the seeds having all been eaten, as to what remained of them having sprouted and
so become uneatable.”

Where quiet jungle roads are found, frequented by cattle and mules, these birds
will often be found in the early morning, and as many as fifteen have been seen within
the space of a few miles. The Grey Junglefowl is decidedly not a gregarious bird,
however, and I have never heard of a real flock or covey, except as above mentioned
when numbers are associated temporarily for the purpose of enjoying some favourite
article of food. When several are found together, it will be noticed that there are
almost never more than two adults, presumably the parents of the remaining more or
less immature birds. Generally they are found singly or in pairs. As with many
birds, I believe that the pairs are associated throughout the year, although they may
feed day after day by themselves. It may be that there is some sort of reassociation,
however, during the hot weather and rains, for at such times the cocks seem to be
solitary, while when one hen is flushed, several others are often to be found close by.

236 A MONOGRAPH OF THE PHEASANTS

The voice of the Grey Junglefowl is very characteristic. The crow of the cock
must either show great variation in different parts of its range, or, what is more
probable, the ears of the people who have described it have been at fault, deceived by
the call of other birds of the jungle, or by the differences caused by distance, certain
notes being thus altogether inaudible, or much altered as to time and pitch. During
the breeding season, especially when the weather is cloudy, the cock may call at intervals
throughout the day, but as a rule in the cold weather its challenges are confined to
early morning and toward sunset. The crowing seems to extend beyond the actual
limits of breeding in any one locality, being heard sometimes from October or November
until May, during the period of most perfect plumage. The utterance is not difficult
to imitate, and with a very little practice the bird may be made to answer again and
again. I have never, however, been able to draw the bird within sight or even very
far in my direction, as is so easily done with other species.

Disregarding several published descriptions, which fall very wide of the mark, we
find that one reliable author transcribes it as a broken, shrill a&-d-ak-Rhee ! uttered
very deliberately and at intervals. This is certainly a very unusual type of crow, if
it be correct.

A very careful observer writes me as follows: ‘The crow of the cock (preceded on
the ground by a flapping of the wings as a rule, but not always) is Kaaa, a-a-doodle-a,
the first note a scream, but not always uttered; in fact there seems to be no rule as
to when it precedes the crow and when it does not. There follow two short, staccato
notes, a double, rather lower note, well described by ‘doodle, which is the emphatic
syllable of the crow; then another short a, followed, sometimes, by an indrawn note
which is not heard except at close quarters. Perhaps in the crow as represented above,
the final a would be better represented by da. The call is uttered, as a rule, from trees
in the early morning, sometimes even at night, generally when there is moonlight, but
occasionally even when there is pitch darkness, and is then, as far as I know, never
preceded by a flapping of wings. The key of the voice is somewhat sharper and higher,
more staccato than that of the common farmyard fowl. The flapping of the wings
always precedes the vocal part of the challenge.”

‘The crow of the male,” says Davidson, “is very peculiar, and might be syllabled,
‘ Kuck-kaya-kya-kuck, ending with a low, double syllable, like ‘Kyukun, kyukun,
repeated slowly and very softly, so that it cannot be heard except when one is very
close to the bird.” This is the best published transcription with which I am familiar.
Sitting with note-book in hand within a comparatively short distance of a crowing bird,
and then checking this with repeated observations of the challenge of this Junglefowl
in captivity, I find the average crow to be as follows, except that perhaps an octave
higher would be nearer the pitch:

SN

ide’ = poe ==
—

The alarm call consists of three short notes in a high key, like fuck-tuck-tuck or
kuck-kuck-kuck, the middle syllable, or rather word, for the notes are quite separate,
emphasized and somewhat higher than the other two. This call means that the cock
or hen (hers is very slightly different) sees something it does not like, or cannot quite

GREY JUNGLEFOWL 237

make out. When put.up by an enemy they go off with a loud, frightened kakakakakak !
with the @ as in the word “that.” This outburst is often continued until they alight
in a distant part of the jungle.

| There is nothing of especial interest about the gait of these birds, which differs in
no respect from that of other Junglefowl. The tail is carried low, except when the bird
is courting, or approaching a rival. When it runs at full speed the tail fairly trails
and the head is also lowered until in a straight line with the body. When listening
for danger, the bird stands so erect that the line of its neck and back is almost vertical,
the longer tail-feathers pressing against the ground. The flight is rapid and strong.
When thoroughly alarmed by the sudden appearance of a sportsman, the birds will
often run with head and tail held low, dodging about any convenient tufts of vegetation,
until at some distance, when they rise with rapidly beating wings and fly for several
hundred yards before seeking safety in some dense bit of jungle.

DAILY ROUND OF LIFE

Grey Junglefowl come into the open, as upon roads, or at the edge of fields, or in
open patches in the jungle itself in early morning, retiring to cover as the heat increases.
During mid-day they remain quietly on the ground or on low branches, seeking food
rather casually, preening their plumage, or indulging in the luxury of dust baths.
Toward evening they again resume activity, coming again into the open, or descending
to some favourite stream to drink. In cloudy weather, however, they move about in
the open throughout the day.

The food of the Grey Junglefowl is varied in character, consisting in general of
various seeds and grain, small fruits and berries, with occasionally leaves and flower
petals, while insects of almost any order, but especially termites, are acceptable.
Sometimes their crops will be crammed with nothing but grass seeds, or again millet
from the fields of the natives, or after land has been burnt over, they enjoy the tender,
juicy shoots of the newly-sprouted grass.

They roost in trees, and when deer-hunting at daylight sportsmen have often
flushed them from their perches. They usually select some bent bamboo stalk, or again
frequent a clump of dense evergreens, perching high up and always in thick jungle.
One correspondent writes me that they always perch singly, even although there may
be several in the same tree or in neighbouring ones.

It is the impression of some observers that the Grey Junglefowl are less courageous
than the red species. ‘They are so extremely wary,” to quote Davidson, ‘“‘ where birds
and animals of prey are concerned, and wander such short distances from the edges of
cover, that I think very few of them fall victims to any enemy but man. There are
plenty of the Bonelli eagles and some hawk-eagles too in the Nilgiris, but I do not
think that these ever succeed in capturing Grey, as they do elsewhere red, Junglefow] ;
at any rate, I have never once seen the feathers of sonnevati strewed about, as I have
those of ferrugineus in Burma.” It is probable that this is an error, and I see no
reason to think that they enjoy any greater immunity from attack than any other species
of bird living under corresponding conditions. In fact the proclivity of red junglefowl
to live in the vicinity of native villages, and their consequent familiarity with mankind,

238 A MONOGRAPH OF THE PHEASANTS

probably renders them immune to the attack of many wild creatures who fear to
approach so closely to civilization, and also tends to make us more familiar with the
pugnacity which is so very general a character of birds of this group. To the north
at least, in Kanara, their enemies are numerous enough, including leopards,. cats of
several species and hawk-eagles. Pythons must prove a danger at times, and mon-
gooses may be one of the most dreaded enemies. In Kanara the crested hawk-eagle
kills large numbers. :

Feeding in the same thickets or jungle with Grey Junglefowl, one often finds
spurfowl, bustard-quail, and other small gallinaceous birds, none of which appear to be
in any way actual competitors. On the other hand, we find, as among so many
pheasants, a quite close association between Junglefowl and various species of babblers,
which at times are found feeding and actually mingling with a family of half-grown
Grey Junglefowl. The bond in this, as in all similar cases, is facility in obtaining food,
the scratching of the fowls sending many insects flying and hopping, enabling the lesser
birds to seize them before they can escape. In return, the babblers all unconsciously
render full payment, by keeping a keen look-out for danger from their more elevated
perches, and often spoil a long, careful stalk.

This Junglefowl needs little help, however, in the matter of detecting danger, for
its eyes and ears are of the keenest, and it will even discover one when perched
motionless in a tree. In such a position I have found it possible to watch many
species of pheasants without attracting their attention or even arousing their suspicions,
but the keen, suspicious nature of the Grey Junglefowl has always been more than
equal to any ruse of mine.

I have myself noticed, and we find the fact recorded by several writers, that the
cock is much more shy than his mate. This is in line with my observations upon other
pheasants, where the more conspicuous the difference in colour between the sexes, the
more pronounced is the amount of wariness and boldness respectively, displayed in the
face of approaching possible enemies.

Where they have never been disturbed they are not especially wary, and will even
take their time in leaving some attractive bit of food. Even in such a case, however,
a good gun-shot distance marks the deadline at which they disappear into the nearest
cover. When they have been disturbed or shot at for a few weeks, they become
exceedingly shy and wary, and it requires the best woodcraft to catch even the most
fleeting glimpse of them.

When not feeding they do not usually wander very far from cover, and a fraction of
a second suffices for them to dash out of sight. Unless actually pursued, they do not
go very far away. If the observer passes on and pretends to leave the spot, slipping
behind a tree-trunk at the last moment, and waiting quietly, the birds will soon reappear
close to the place where they vanished. If the danger seems real, and they think that
they are being sought, they will continue to thread their way silently, deep into the
heart of the jungle, often, after they have gone some distance, flying up and concealing
themselves in the dense foliage of a tree.

GREY JUNGLEFOWL 239

HOME LIFE

There are records of these birds breeding from October to July. As for example:
Nilgiris, March and April; Ootacamund, October, November, and December ; Coonor,
May and June; Satara, March and April; Mysore, November to July; Mt. Abu,
March to mid-June; Kanara, October to April. In fact, there is one record from South
Travancore of a nest and eggs found on August 20.

The courtship of the Grey differs in no essential particular from that of the red
junglefowl. The approach, the beginning, and the climax, are all similar.

Despite some arguments to the contrary, they are fully as pugnacious and
courageous as the red junglefowl, and their well-developed spurs show that their
encounters must be far from harmless jousts. A writer from the Nilgiris says that
their pugnacity is something incredible. ‘‘On one occasion, when my brother was out
shooting, he heard in the jungle near him the peculiar under-toned notes that the cocks
emit when fighting. After a few minutes the sound ceased, and on reaching the spot
whence the sounds had proceeded, he found two Junglecocks dead, and one of his dogs
by them. On examining the birds, both their heads proved to bear the marks of the
dog’s teeth, which could only be accounted for by the supposition that they were so busy
fighting that they failed to observe the dog’s approach, and were so close by pecking each
other's heads that the dog seized both heads at once; for if she had seized only one, the
other bird would have been out of reach before she could have made a second bite.”
The encounters usually take place in the morning, soon after the challenges begin.
A mated bird seems to have its own particular beat and sends its alar and vocal
challenges from its perch, awaiting the acceptance and approach of a rival. The
unmated birds are probably those who are the actual aggressors and whose advance
precipitates the battle. ‘This explains the fact of two birds, within a short distance of
each other, crowing and challenging continually day after day without attack. Both
are probably mated, and concerned only with defending their chosen section of jungle
and their mate. The monsoon period is the time of these encounters, while at other
seasons, when, as I have related, many cocks and hens are temporarily associated in
feeding, there is no hostility exhibited between the cocks.

While a cock may mate with two or three hens under certain circumstances, I
believe this to be unusual, and that monogamy is the normal condition. This view is
strengthened by the fact that while the cock has nothing to do with incubation, and in
reality keeps at a considerable distance from his sitting mate, yet when the young are
hatched he is frequently seen with them, taking his share in finding food. Such a habit
could hardly have become established were he polygamous.

Within narrow limits the nest itself is rather variable. It may be a mere natural
depression in the ground, devoid of any lining whatever, or it may contain the dé6ris of
leaves, grass or moss which existed in it when chosen by the bird. Usually the parent
scratches out a hollow, almost always close against the trunk of a tree or under the
shelter of some bush, which is lined with dead leaves and feathers. There are a number
of records of the eggs being laid on the top of dead, half-decayed stumps, from two to
four feet above the ground. Indeed, this seems to be a habit which is on the increase,

240 A MONOGRAPH OF THE PHEASANTS

and is interesting as a radical departure from the pronounced terrestrial nidification of
this group. :

When all the records of number of eggs are compared, there seems to be a very
significant gradation from north to south, marking a regular increase. A correspondent
writes that in Kanara there are never more than four in a nest, and usually three ; this
test being applied to many nests found early in the season, and with eggs about to
hatch, thus freeing this record from the complications both of a second laying or an
incomplete set. South of Mt. Abu, six to seven seems to be the average number, while
in the.Nilgiris and southward, eight or nine to thirteen are recorded. Whether this is
due to an increase in the number of dangers or not, is an interesting speculation. In
connection with this, it is significant to note that the birds from the northern part of the
range seem to run larger and heavier than those from farther south. Two observers at
extremes of the Junglefowl’s range tell of the hen covering the eggs with dried leaves
when she goes away to feed, but I can obtain no recent verification of this, while a third
man, writing from Poona, emphasizes the close resemblance of the eggs to the dried
leavés among which they lie.

There is great variation in the colour and size of the eggs, and when the extremes
are compared it would never be supposed that they were laid by the same species of
bird. At one extreme we have an egg of a long, oval shape, with a pale, creamy-white
shell like that of a domestic hen; the other extreme shows a broad oval, of a deep rich
buff or café au lait, and a coarse texture, deeply and densely pitted, like the shell of a
peahen’s egg. Every intermediate type is met with, some of which are everywhere
thickly speckled with brownish red. The eggs vary from 43 to 51 mm. in length, and
31 to 38 in breadth, averaging 47 by 34 mm.

Several observers write me that in the north, at least, there is some evidence that
two broods are reared inaseason. I think this rather doubtful, and probably due to
finding late broods consequent upon the first laying of eggs being destroyed. In the
north, two seems to be the usual number of young reared to maturity, while in the
south, I have had three reports of half-grown young in families of four, four and six
respectively.

The young. are able to look out for themselves soon after hatching, hiding at their
mother’s note of warning by squatting or scurrying under leaves or into clumps of
bamboo. Not many days pass before their tiny wing-feathers are sufficiently well grown
to support them. Even before they are strong enough to rise from the ground they will
clamber, with the aid of feet and wings, up into shrubs and bushes, and from this point
of vantage leap boldly into the air, and go off with a diminutive whirr like quail, getting
up speed quickly, keeping up the pace for a few yards and then dropping exhausted into
the undergrowth. Mr. T. R. Bell, of Kanara, has had Junglehens come up when he had
sat for a long time, near where several little chicks were hiding, and pretend to be
maimed, and when this failed, erecting their hackles and showing considerable anger.
Only once was the cock seen at such a time, and he kept at a safe distance. The notes
of the hen as she calls her chicks is like the voice of the domestic bird, a soft tooh-took-
took, and the cock uses the same utterance at times. The Jeep, peep, of the chicks is the
same infantile note as in our barnyard birds. The young birds remain with,the parents
at least until they have attained fully adult plumage, and how much longer is not known.

GREY JUNGLEFOWL 241

The crop of one young chick was filled with soft vegetable matter, some tiny beetles,
and a single small moth. The colours and pattern of the young bird, yellowish-buff
with the characteristic broad black line through the head and eye, is of great value in
breaking up the general outline and approximating the hues of the jungle dédrzs. This
down gives place quickly to the juvenile plumage, and the first annual moult brings the
bird into adult garb. About June or July the old cocks begin to moult their hackles,
and possibly, sometimes, their longest tail feathers, the former being replaced at once by
a short temporary plumage. Later in the year, these short eclipse feathers are replaced
with the resplendent whitish or yellow hackles, and by October the bird is in perfect
feather, and clears his throat for the first challenge.

RELATION TO MAN

Natives capture the Grey Junglefowl both by pegging out decoy birds, which challenge
and thus attract other cocks, and by means of nooses and spring traps. When a gun is
available, the jungle natives put a plan into operation which seldom fails of its purpose.
They make a blind of brush and leaves near a spring of water, and wait patiently
hour after hour until the birds come to drink. When there are other springs in the
neighbourhood they hang pieces of white cloth about to frighten the fowl away, and
thus make certain of their visiting the spring where they are keeping watch. Large
numbers are killed annually in this way in the north. Although the birds are invariably
thin, and consequently tough, the flesh is eaten by the natives. The hackles are valued
by the makers of artificial flies for fishing, and great quantities are exported to Europe
for that purpose.

In the Ghats, while the birds are still numerous in places, their numbers have
been greatly decimated by the numerous snares which the natives set around every patch
of rice. The jungles and dense bamboo clumps offer a safe haven for this species, and
as long as such exist, it is in no immediate danger of extermination. It does not adapt
itself to deforestation, however, and when any considerable area is brought under
cultivation, the Grey Junglefowl retreats to the nearest dense coverts. The game laws
in the Nilgiris are very generally respected by white sportsmen, and prohibit Junglefowl
shooting from April 1 to September 30.

The best time for shooting Grey Junglefowl is from November to the end of
March, as during that time their plumage is in the most perfect condition. Indeed,
the birds are worth little else than the beauty of their feathers, for the flesh, as I have
said, is very dry and hard, and even the breast possesses little real flavour. Although at
times food is as abundant as could be desired, and their crops will be filled to bursting
with succulent grain and seeds, yet these birds seem never to become fat, but always,
like an athlete, to be in perfect condition for exercising their senses and their muscles to
the utmost in detecting and avoiding danger.

“Their great timidity and watchfulness,” says Davidson, “result in their yielding
much less sport than the Red Junglefowl. You may get these latter in standing crops
and in many other similar situations without any extraordinary precautions, but the
Grey Junglefowl never goes more than a few yards inside the fields, and if a stick

cracks, or a sound is heard anywhere within fifty yards, he vanishes into the jungle,
VOL. I i

242 A MONOGRAPH OF THE PHEASANTS

whence it is impossible to flush him. Only when beating the narrow, well-defined belts
of tree jungle that run down the ravines on the hillsides in the Nilgiris, and which we
there call s/olas, is anything like real sport to be got out of them. Then, indeed, the gun
at the tail end of the skola may get three or four good shots in succession, as they rise at
the end of the cover and fly off with a strong, well-sustained flight to the next nearest
patch. Even thus, working hard and beating sola after shola, a man will be lucky to
bag five or six brace in a day.

“The reason is, that all the well-defined so/as which can be thoroughly beaten are
in the higher parts of the hills, where the birds are comparatively rare, while when you
get lower down, where the birds are plentiful, the jungles are so large that they cannot
be effectively worked. If you merely want to £7// the birds, you might get perhaps ten
or a dozen in a short time poking along some of the roads, but they afford no sport
thus, only a series of pot shots.

“T remember once watching an old cock that my dogs had driven up into a tree.
For some time I peered round and round (the tree was a large and densely-foliaged one)
without being able to discover his whereabouts, he all the while sitting silent and
motionless. At last my eyes fell upon him, that instant he hopped silently on to
another bough and from that to another, and so on with incredible rapidity, till,
reaching the opposite side of the tree, he flew out silently, of course never giving me a
chance at a shot.”

A sportsman near Mt. Abu says that the cordon system of driving “is usually
adopted in shooting them here. The guns are placed behind screens made previously
by the ‘shik4aris,’ at the ends of patches of jungle the birds are known to affect, and the
beaters are sent round to drive the birds up to them, forming a semi-circular line to
prevent the birds escaping at the sides. It is very poor sport, you seldom or never get
a flying shot, and when you do, the jungle is so thick that it is about ten to one you
miss. The birds, especially the old cocks, are remarkably wary, and the moment they
hear the beaters they begin to run, stopping about every fifty yards to listen.

‘They have a very quick eye, and alter their course immediately if they see or hear
the slightest thing in front. The only way, therefore, when you know a bird is coming,
is to raise your gun silently to your shoulder, turn very quietly in the direction from
which it is coming, and remain perfectly motionless, and as soon as ever the bird gets
within shot, fire.

“T have shot them with dogs, but that is equally poor sport. As soon as the
Junglefowl sees the dog, he flies up into a tree and squats upon a bough until you
dislodge him from his supposed place of security with a charge of shot.”

Dr. G. H. Krumbiegel, superintendent of the Mysore Museum, writes me that in
Mysore the Grey Junglefowl is protected from March 1 to September 1, but that
natives shoot them throughout the year. The method usually adopted is to sit in
ambush over a water-hole, or feed the birds regularly at a certain spot, and when they
are well together feeding, to “ brown” the lot. A shikari will sometimes call up a cock
by imitating the hen. The sound is a very soft note, and one would scarcely think the
cock would hear it thirty or forty yards away, but he rushes along impetuously within
half a minute. The Kurbars track Junglefowl, but the people who destroy them to the
greatest extent are the Pardees, a wandering tribe in Mysore, whose only occupation is

GREY JUNGLEFOWL 243

poaching. The Kurbars use dead-falls of sorts, and also drive stakes about fifteen
inches high into the ground close together to form a circle about two feet in diameter,
in which grain is placed, and the birds learn to feed through two openings which are
left open for the purpose. After a while, a very ingenious arrangement made of bamboo,
on the principle of a scissor trap, is placed at the opening, into which the bird gets its
head and is killed. The Pardees approach the feeding birds under cover of a bull, and a
line of nooses having been set, they mill around until the birds are close to the line.
They then suddenly leap out and scare the birds, and in their rush to escape many are
caught by the neck or leg. Any one who has tried to get even within gunshot of such
birds will realize the skill which these natives must possess to successfully carry out this
method of attack.

CARIN la Va

The Grey Junglefowl was first bred in Europe in 1862, when thirteen hybrids with
the red species were reared in the London Zoo. Since then it has been bred many
times in various public and private aviaries, both pure and as crosses. Of twelve birds
of which records were kept in the London Zoo, one notable individual lived twelve years
and four months, the average length of life of the others being two years and a half.
The period of incubation lasts from eighteen to nineteen days. A pair of these birds
bred successfully in the aviary of F. E. Blaauw in Holland. The hen laid three
successive times, in clutches of four eggs each, and from these ten birds, five cocks and
five hens, were reared. The first two layings were taken away and set under bantam
hens, while the last lot of four was left to the mother, who sat faithfully and hatched
them all. The richly-coloured chicks thrived on ants’ eggs, and assumed their adult
plumage in the first autumn.

When the eggs of wild Junglefowl are brought in and hatched under a domestic
hen, the chicks remain contentedly with the mother until they are able to fly, when they
roost at night by themselves on some bush or low tree. When a few months old they
invariably retreat to the jungle, and do not return. Grey Junglefowl seldom become
tame in captivity, certainly not more so than the average pheasant. While there is no
doubt that the red species is the direct ancestor of all of our poultry, the rather close
relation of the grey bird is shown in the facility with which it crosses with the
descendants of its generic relative. In the native villages of Kanara and elsewhere, it is
not a rare sight to see hybrids which possess more or less perfectly the bright sealing-
wax hackles of one parent, while they have inherited sufficient domestic instincts to
induce them to remain with the other inmates of the compound.

DETAILED DESCRIPTION

ApvuLtt Mare.—The feathering of the head is confined to a narrow line of reddish,
rather recurved feathers between the comb and eye, and a small rounded tuft of silvery-
white feathers covering the ear. The face, chin, and throat appear naked, but are evenly
but thinly covered with a scanty growth of filo-plumes, each a simple hair-like shaft, or
tipped with several rudimentary barbs. Immediately back of the comb the feathers take
on the character of the hackles as a whole. ‘The smallest of these show a small spot, flat

244 A MONOGRAPH OF THE PHEASANTS

and enamel-like, framed in black, with the base of the feather white. As we proceed
backward over the neck, the feathers gain in complexity. The longest hackles are well
over 100 mm. in length, and strung along the centre of the vane they show as many as
four distinct, specialized, enamel-like patches. This is in no sense an enlargement of
the rhachis or shaft, but a solidification or cornification of the vane as a whole. The
barbs all run into this area, and distally emerge from it, unchanged as to their position
in the vane as a whole. The sealing-wax simile is a perfect one. If one took any
normal feather and dropped four beads of sealing-wax at intervals along the shaft,
allowing them to spread over about half the width of the vane, the hackle spots would
be perfectly imitated. The terminal waxy spot soon becomes frayed and split, and when
the wear and tear thus becomes apparent, it presents the appearance of the margin of a
palm frond.

The hackles have a wide, disintegrated fringe of grey, the remainder of the feather
being dark-brown or black. The concealed spots are always pure white, but the
terminal wax may be either white or tinged with pigment until it is of a deep yellow
ochre. The free unmodified part of the vane may be grey-fringed to the tip, or it, too,
may be heavily stained with yellow. The extremes of these two colours are birds very
unlike in appearance, but the variation is dependent neither upon age nor locality. There
is sometimes a hint of a proximal fifth area of specialization, and often this is so rudi-
mentary that the modification has affected pigment only, the node being white, but
wholly normal in structure. The dark portions of most of the longer hackles are
strongly glossed with violet and purple.

The hackles, besides extending backward over the whole mantle plumage, are
continued around the sides of the bare neck until they almost meet in front on the
lower neck.

On the lower hind neck the hackles give place abruptly to the normal plumage of
the mantle. These and the majority of the body feathers and the lesser wing-coverts are
dark-brown or black—dull in some individuals, in others glossed with purple—narrowly
edged with grey, and with still narrower white shaft-stripes. There is sometimes con-
siderable grey mottling in the dark area, regardless of age, but usually this is lost in the
fully adult bird.

The purple gloss is accentuated on the rump feathers, and in addition a series of the
small, wax-like ornaments appears, and the visible fringe is deep orange-red. The
shorter upper tail-coverts are bronze-purple, the longer ones purplish-blue, with a
marginal band of greenish. The tail-feathers are black glossed with bluish-green, on
both webs in the central, on the outer web alone of the lateral rectrices.

On the median wing-coverts occurs another extremely specialized zone; if anything
even more extreme than that of the hackles. The enamelled area is so extensive that in
full-plumaged birds it forms a broad, solid zone across the wing. The proximal part is
formed by feathers which have but the merest tip of waxy structure, while in the
posterior part of this wing area the feathers are solidly cornified for some 30 mm. of
their length, the entire distal part of the vane being thus affected, and the colour being
a rich ochre. Where any part of the visible portion of the vane is free, it is orange-red.
The greater primary coverts are dull, dark-brown, while those protecting the secondaries
are glossed with purple, with only a white shaft-mark to hint of the specialization in the

GREY JUNGLEFOWL . 245

adjoining feathers. Flight-feathers brownish-black, the primaries faintly edged with
pale grey, and the secondaries glossed with greenish. /

Under-parts like the back, but with much more greyish-white on the fringes and
shaft-stripes. Indeed, posteriorly, the black becomes restricted to two broad longi-
tudinal bands. A patch of feathers on the sides of the belly and the flanks shows a
tendency to form another zone of specialization, but here we find only an abundance of
the orange-red pigment, with little or no cornification of the vane.

Comb, wattles and bare skin of head, chin; and throat red, very intense at the
season of courtship, and often extending far down the neck under the clothing of hackles.
Upper mandible dark-brown or black, paler at the tip; lower mandible yellowish horn.
Irides varying from clear red, through reddish-orange to yellow. When in full colour
the legs and feet are salmon red, and the claws black. After death and at other than
the breeding season, a yellowish hue predominates.

Length, 700 to 800 mm.; culmen from nostril, 16 ; wing, 240; tail, 380; tarsus, 77 ;
middle toe and claw, 64. Spur long, slender distally and much curved, 30 mm. in
length. Weight 1 lb. 9 ozs. to 2 Ibs. 9°5 ozs.

VARIATION AND ANNUAL MOULTS

In examining a large series of skins, some from areas of dense, humid jungle, and
others from semi-arid regions, I can detect no correlated difference in colour. I have
already spoken, however, of the considerable variation in individual cocks, regardless of
age or locality. This is most pronounced in the hackles, in which the terminal enamelled
spot may be pure white or rich yellow ochre. Then, too, in cocks of equal age, the
body plumage may sometimes lack the iridescent gloss, while the amount of white varies
within wide limits. The absence of gloss and excess of white are also, however, marks
of immaturity.

In June or July, when the duties of the breeding season are quite over, the hackles
begin to be dropped. ‘This moult of the hackles is of great interest as being an eclipse
moult of short duration. From the forehead back to the lower neck, and around, almost
meeting on the ventral part of the neck, the shed hackles are at once replaced with black
feathers. Those on the head have white, hair-like shaft-stripes, while the larger ones
are faintly glossed with purple. ‘There is absolutely no trace of the specialised waxy
spots, and the whole aspect of the bird is altered. These black feathers are retained
until the regular annual moult sets in, some three months later, when they, together
with the entire body and wing plumage, are shed, and replaced with resplendent new
feathers. The early shedding of the long, central tail-feathers is irregular. They are
often broken off at this season after the wear and tear of the period of battle and of
breeding, so that they do not project beyond the other shorter tail-feathers. In ten
specimens in eclipse plumage, six had the feathers thus broken, and hence apparently
moulted ; in two others they were present and full length, although much worn, the
eclipse hackle plumage in this case already showing signs of being shed. In two other
birds the central tail-feathers were shed; but so were one or two of the other rectrices
adventitiously from one side, showing that the regular rectrice moult, from outside in,
had not yet begun. Thus there seems no ground for belief in the widely-repeated

246 A MONOGRAPH OF THE PHEASANTS

statement that the central pair of tail-feathers is shed simultaneously with the hackles.
After two or three months, in the course of the autumn, all the oe ornaments are
again assumed by a second moult of these feathers.

ADULT FEMALE.—Feathers on the head brown or dull orange, shading into rusty
or rufous on the neck and mantle, and paling into buffy-white on the rear mantle.
These feathers have all two broad, longitudinal lines of black extending down the vane.
All the remaining upper plumage, together with the wing-coverts and visible portion of
the secondaries, is a fine mottling or vermiculation of sandy or rufous brown and black,
the back, the lesser and median coverts with a conspicuous, hair-like, white shaft-streak.
Primaries plain dull brown. Tail-feathers black, slightly glossed with greenish, the
central pair with a marginal mottling of reddish-brown.

The face is covered with a scanty growth of brown featherlets, the chin and throat
with a somewhat denser plumage of white. The pattern of the ventral plumage is of a
central area of white and a wide black or dark-brown border. The white increases
posteriorly, until on the mid-belly it obliterates the dark marginal colour. There is
considerable mottling, however, of the white area, in the form of two irregular longi-
tudinal lines in the centre of the vane. The under tail-coverts are dark-brown, with a
small, terminal, white shaft-spot.

The comb is very rudimentary, 3 or 4 mm. in height, slightly notched. The spurs
are short, rounded nodules. Upper mandible dark horny-brown ; lower yellowish-white.
Irides red to yellow; facial skin reddish, never as bright as in the cock. Legs and feet
brownish yellow, darker on the toes. Length, 380 mm.; culmen from nostril, 19 ; wing,
200; tail, 145; tarsus, 63; middle toe and claw, 50 mm. Weight 1 lb. g ozs. to
1 lb. 12 ozs.

Cuick IN Down.—The lores of a week-old chick, and a line backward around the
edge of the eyelids, and on over the ear-coverts and around to the nape, dark chocolate
brown. A \-shaped mark, with the apex on the mid-crown, and extending backward
to the nape, where it meets the two lateral facial lines, is dark chocolate brown, with a
narrow bordering line of black. All the rest of the head, neck and breast yellow buff.
Chin and throat creamy white. The chocolate band is continued in an even width down
the back to the rump. The side mantle, anterior wings and sides are buffy brown or
fawn colour. The new sprouting scapulars and wing-coverts are tipped with greyish-
white. From the mid-back to the rump, the central chocolate band is flanked by two
creamy white stripes of somewhat lesser width, and outside of these we find still another
pair of narrow chocolate lines. The under surface and thighs are creamy white. The
wing-feathers are pale sandy peppered with darker brown.

Culmen from nostril, 5°55 mm.; wing, 55; tail, 18; tarsus, 20; middle toe and
claw, 19 mm.

JUVENILE PLumAGE.—When the down of the head is shed, it is replaced by a
fairly dense growth of brownish-black feathers. The comb at this time is small and
rounded, exactly like that of the adult hen. The head and neck hackles of this plumage
show no hint of the succeeding specialization, nor do they resemble the corresponding
feathers of the female, but they are very much like the post-nuptial eclipse into which

PLATE XLIII

PLUMAGES OF THE GREY JUNGLEFOWL

Gallus sonnerati TVemminck

Tue chick in down has very distinct patterns and coloration, and the wing-feathers sprout
rapidly so that it can fly a short time after hatching.

The juvenile plumage of both sexes resembles that of the adult female, with usually a hint of
the sealing-wax spots on the median wing-coverts of the cock.

A full-grown cock in the eclipse plumage has the specialized neck hackles replaced with black

ones. This partial moult lasts only for three months after the breeding season.

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iv

PLATE XLIII.

PLUMAGES OF THE GREY JUNGLEFOWL

GREY JUNGLEFOWL 247

the adult cock moults every year. They are black, with a faint violet gloss, a wide
greyish or rusty fringe and a very narrow shaft-streak. The lower mantle, back and
rump show somewhat the same general pattern, but lack the gloss and have much more
adventitious mottling. The grey fringe is much whiter.

The primaries are plain brown, but the visible parts of the greater coverts and the
secondaries are thickly mottled and vermiculated with sandy brown. The median
coverts are mostinteresting as showing well-developed waxy spots of yellow ochre, and
terminal shading of reddish-orange, hinting that this zone of specialization is older than
the others. The tail-feathers, like the secondaries, are much mottled. The under parts
are very similar to the plumage of the adult cock, with the colours less distinct and
separate. There is sometimes a slight tinge of orange on the flanks.

In the full-grown juvenile bird, both the irides and the feet and legs are yellowish-
brown. The upper mandible is brownish, duller than in the adult, and the lower
mandible yellowish. The comb and facial skin are pink. Length, 380 mm.; culmen
from nostril, 14; wing, 200; tail, 125; tarsus, 70; middle toe and claw, 55 mm.

Post-JUVENILE OR First ANNUAL Movutt.—With this moult the adult plumage
is attained, the chief difference from the juvenile plumage being the acquisition of the
specialized hackles and central tail-feathers. There is a great deal of variation, however,
in the amount of clearing up of the plumage, and in birds which have moulted early
the waxy hackles are very imperfectly developed and the general body plumage and
wing-feathers much mottled with brown or sandy.

EARLY HISTORY

Under the name Cog e¢ Poule sauvage des Indes we may recognize the Grey
Junglefowl in Mon. Sonnerat’s Voyage aux Indes Orientales et a la Chine, which was
“Fait par ordre du Rot, depuis 1774 jusguwen 1781.’ He gives a long, rambling
description of both cock and hen, and a black-and-white engraving vaguely showing the
sealing-wax hackles, but nothing of the habits or life of the bird. His final conclusion
is the very erroneous one that this Junglefowl represents the progenitor of our breeds
of domestic fowls. Other authors simply copy Sonnerat or enlarge on his facts with
sad effect. All agree with his theory of descent, until Temminck in 1813 proves
conclusively that this is an error, and names the bird in question after its first
describer Gallus Sonneratit.

SYNONYMY

Cog et Poule sauvage des Indes Sonnerat, Voy. Ind. Orient, II. 1782, p. 148, pls. 94, 95.

Wild Cock Latham, Gen. Syn., II. 1783, p. 698.

Phasianus gallus Scop. (nec Linn.), Del Flor. et Faun. Insubr., pt. II. 1786, p. 93; Gmelin, S.N. I. pt. II,
1788, p. 737; Latham, Ind. Orn., II. 1790, p. 625; Bonnat. Tabl. Encycl. Méth., I. 1791, p. 180, pl. 86, figs. 4, 5.

Sonnerat’s Wild Cock Latham, Gen. Hist., VIII. 1823, p. 181.

Gallus sonneratti Temminck, Pig. et Gall. II. 1813, p. 246, III. 1815, p. 649; Steph. in Shaw’s Gen. Zool.,
XI. 1819, p. 200, pl. 12; Temminck, Pl. Col., V. 1823, pls. r and 2, Nos. 232, 233; Vieillot, Gal. Ois., II. 1825, p.
26; Griff. ed. Cuv., III. 1829, p. 19; Lesson, Traité d’Orn, 1831, p. 492; Sykes, Proc. Zool. Soc., 1832, p. 151
[W. Ghauts]; Schinz, Nat. Abbild. Vog., 1833, p. 243, pl. 94; Jard. Nat. Lib. Orn. 1834, p. 186, pls. XI. and
XII.; Gray, List of Birds, pt. III. Gall. 1844, p. 27; id. Gen. B., III. 1845, p. 499; Blyth, Ann. Mag. N. H., XX.

248 A MONOGRAPH OF THE PHEASANTS

1847, p. 388, I. 1848, p. 455 ; id. Cat. Mus. As. Soc., 1849, p. 243 ; Schinz, Nat. Vég., 1853, p. 145, pl. 69; Burgess
Proc. Zool. Soc., 1855, p. 29; Sacc, Rev. Zool. (2), XIV. 1862, p. 11, pl. 3; Sclater, Proc. Zool. Soc. London, 1862,
p- 186 [first breeding in captivity]; Jerden, Birds India, III. 1863, p. 539; Sclater, List of Phas., 1863, p. 10 ; Bulger,
Proc. Zool. Soc. 1866, p. 571 [Nilghiris]; Ornithogonomon, Field, XXVIII. 1866, p. 183 [general account] ;
Tegetmeier, Field, XXVIII. 1866, p. 191 [characters of hybrids]; Gray, List Gallinae Brit. Mus., 1867, p. 39;
Blyth, Ibis, 1867, pp. 154, 307; Sclater, Proc. Zool. Soc. London, 1869, p. 628 [breeding in London Zoo]; Gray,
Hand-list, B. II. 1870, p. 261; Elwes, Ibis, 1870, p. 528 [Cardamum Hills]; Elliot, Mon. Phas. I]. 1872, pl. 34;
Nathusius, Jour. fiir Orn., 1872, p. 329 [shell characters]; Lloyd, Ibis, 1873, p. 4or [S. Konkan]; Hume, N. and
FE. Ind. B., 1873, p. 531; Butler, Stray Feathers, 1V. 1876, p. 5 [M. Guzerat and Aboo]; Fairb., Stray Feathers,
IV. 1876, p. 262 [Khandala to Goa]; Hume, Stray Feathers, IV. 1876, p. 404 [Travancore]; Butler, Stray
Feathers, V. 1877, p. 222; Fairb., Stray Feathers, V. 1877, p. 409 [Palani Hills]; Marshall, B. Nest Ind., 1877,
p. 59; Gould, B. Asia, VII. 1877, pl. 56; Hume and Marshall, Game-birds India, I. 1878, p. 231, pl. ; Davidson
and Wend., Stray Feathers, VII. 1878, p. 86 [Deccan]; Butler, Cat. B. Sind., 1879, p.53 [Aboo]; MclInroy, Stray
Feathers, VIII. 1879, p. 493 [Mysore]; Vidal, Stray Feathers, IX. 1880, p. 76 [S. Konkan]; Butler, Stray
Feathers, IX. 1880, p. 205 [Belgaum], p. 421 [Deccan and S, Mahratta]; Schmidt, Proc. Zool. Soc. London, 1880,
p. 315 [viability in Frankfort Zoo]; Davidson, Stray Feathers, X. 1882, p. 316 [W. Khandeish]; Davison, Stray,
Feathers, X. 1883, p. 409 [Nilghiris, Wynaad and S. Mysore]; Sclater, List Animals in Gardens Zool. Soc.
London, 1883, p. 488; Swinhoe and Barnes, Ibis, 1885, p. 131 [Central India]; Taylor, Stray Feathers, X. 1887,
p. 464 [Manzeerabad, Mysore]; Terry, Stray Feathers, X. 1887, p. 479 [Palani Hills]; Oates, ed. Hume’s Nests
and Eggs, III. 1890, p. 420; Barnes, Jour. Bombay Nat. Hist. Soc., VI. 1891, p. 2 [Mt. Aboo: habits]; Laurie,
Jour. Bombay Nat. Hist. Soc., VI. 1891, pp. 93, 98 [laws for protection]; Grant, Cat. Birds Brit. Mus., XXII.
1893, p. 350; Grant, Hand-book Game-birds, II. 1897, p. 55; Davidson, Jour. Bombay Nat. Hist. Soc., XII.
1898, p. 63 [North Kanara]; Oates, Game-birds India, I. 1898, p. 371; Sharpe, Hand-list Birds, I. 1899, p. 39
[South and Central India Peninsula]; Nehrkorn, Kat. Eiersammlung, 1899, p. 194; Betham, Jour. Bombay Nat.
Hist. Soc., XIII. 1900, p. 383 [Poona: Nest and eggs]; Oates, Cat. Birds’ Eggs Brit. Mus., I. 1901, p. 60 [Eggs] ;
Ghizhi, Mon. Zool. Ital., XIV. 1903, p. 319 [generic characters]; Jukes, Jour. Bombay Nat. Hist. Soc., XIX., 1909,
p. 216 [laws for protection]; Finn, Avic. Mag. (3), I. 1910, p. 129; Mitchell, Proc. Zool. Soc. London, Ig1t, p.
522 [viability in London Zoo]; Stubbe ahd Rowe, Zoologist (4), XVI. 1913, p. 13 [prehistoric origin]; Ghighi,
Mem. R. Accad. Bologna (7), III. 1916, p. 1 [crossing with domestic fowl]; Baker, Jour. Bombay Nat. Hist. Soc,,
ZORIW OM, hs Bile
Phasianus tndicus Leach, Zool. Misc., II. 1815, p. 6, pl. 61.
Sonnerat’s Junglefowl Blaauw, Avic. Mag. N. S., VII. 1908, p. 37 [breeding in captivity].

REAWE env’

JAVAN JUNGLEFOWL

Gallus varius (Shaw and Nodder)

Earty in the morning, before the sun appeared, picking my way quietly through a tangle of cactus
on the low-lying Javan coast, with a shrike sitting in every tree, with bulbuls singing from every thicket,
there would come across the valley a sharp, crisp, virile chaw-au-auk/ the challenge of the Javan Jungle-
fowl. With erect iridescent comb and plumage glittering with dew, the splendid wild fowl was leading

his family from their roosting-place to the nearest pool of water for their sunrise drink.

AG ate BT AIG

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PIEJNIUE OIE AW.

JAVAN JUNGLEFOWL.

JAVAN JUNGLEFOWL
Gallus varius (Shaw and Nodder)

NAMES.—Specific: varius, from the variegated character of the plumage. English: Green, Fork-tailed, or
Javan Junglefowl. French: Coq ayamalas. Native: Ayam oetan (Javanese Malay); Ayam alas, Pitté wono
(Javanese) ; Ayam leuweung (Sudanese) ; Baki koek, Bekikko (Javanese); Kasintoe (Sudanese). The two latter
names are also applied to Gallus gallus. Uybrids between Gallus varius and native poultry are known as Bekisar
or Kakok.

BRIEF DESCRIPTION.—Male: Rounded comb green and purplish red; single, median throat wattle
brilliantly coloured red, yellow and blue. Top of head, neck and upper mantle of short, square-tipped feathers,
black, edged with greenish-bronze; lower mantle shining green, fringed with black; lower back and rump
elongate, narrow feathers, black, edged with pale yellow ; lesser and median wing-coverts similar, but with an orange-
red fringe ; tail glossy green ; flight feathers and lower plumage black. Female: Head, neck and mantle sandy
brown, with an indistinct, concentric black band ; rest of upper parts black, usually glossed with green, irregularly
barred and margined with buff; bars on secondaries buffy white. Throat white, under parts pale buff with darker
margins and indistinct mottlings.

RANGE.—The islands of Java, Madura, Kangean, Bawean, Bali, Lombok, Sumbawa, Flores and Alor.

THE BIRD IN ITS WILD HOME

THREE birds flying out to sea: such was my sensational first meeting with the
Green Javan Junglefowl, amid a scene of rare beauty—the perfect close of a tropical day
on the very north-east tip of the Javan coast.

For hours I had motored over dusty roads in the shade of mile after mile of great
tamarinds, and now, in late afternoon, I was sitting spellbound by the beauty of the
sunset. I had camped on the tip of a little promontory jutting out into the Java Sea.
Behind was an all but impassable chevaux-de-frise of cactus, and overhead and on to the
east, rank after rank of feathery palms and bamboos. I was almost suspended in air, for
directly beneath me were great caves hollowed out by the water, in and out of which
dashed tiny swiftlets—makers of the edible birds’ nests. Seaward lay fair gardens—
every limestone pool left by the tide a mass of colour, blossoming coral—pink, green and
brown; great serpent starfish, like animated sessile Compositae; wandering patches of
lettuce—the green-leafed nudibranchs. Beyond all this, five rows of great breakers
boomed, on the waste of jagged rocks.

On either hand a deep bay swept inward, tracing its lines by the white of the waves,
and to the west, just out of the path of the setting sun, a tiny islet of mangroves raised
its head bravely far from shore.

Between the crashes of the waves I could hear the wheenk ! of a Javan nighthawk,
the sharp twitters of the swiftlets, and now and then the plaintive whistle of a distant
sandpiper. A host of little bats appeared and gyrated about my head; and then, high
up in mid-air, great flying foxes passed slowly over. Herons, no less graceful, began to
collect from far and near in threes and fives, flying low, and headed straight for the

VOL. II 249 K K

250 A MONOGRAPH OF THE PHEASANTS

mangrove islet. As they crossed the sun’s path, they became silhouettes of ebony ;
when they swung up to their roost, they showed as purest ivory against the dark foliage.
Forty, fifty, seventy came, all concentrating on one or two of the trees.

Of the colours of such a scene none may write. The maze of rose, salmon, scarlet,
violet, mauve, and the hundred unnamed tropical tints, succeeding one another and
staining sky and sea and land, defy pen as well as brush.

The afterglow of softened tones of orange and yellow had come, glowing strongest
from the east, as if a new dawn had begun, when three birds walked quietly out of the
bushes, and picked their way over the high tide drift-line of shells and corals. They
were Green Junglefowl—two cocks and a hen, all young birds of the year. Even in this
faint light I could catch an occasional metallic glint from the plumage of the leader as he
scratched half-heartedly here and there among the shells. The small size of his comb
and tail and the buff marks on his wings clearly betrayed his immaturity.

The hen stabbed at a prickly pear fruit, and buried its beak deep in the rosy juice;
another chased a flying insect. After a few minutes one of the cocks crouched and
sprang into the air with a low cackle, and his two companions were on the wing a
moment later. Up and up they went, out over the breakers, straight, as only birds can
fly, to the mangroves. They almost vanished from my straining eyes before they landed,
but I could see a branch bend beneath their weight before the afterglow snuffed out like
a candle and the faint silver of the crescent moon began its dance on the ripples. The
night wind arose, swept through the palm fronds and clattered their frilled edges
together like a myriad castanets; the air was filled with the aromatic incense of leaves
which I had crushed underfoot, and the last sound in my ears was the lulling crescendo
boom of the breakers.

GENERAL DISTRIBUTION

The Javan Junglefowl bears the distinction of being the only member of the group
included in this monograph whose home lies altogether south of the equator. It
resembles its congener, the Ceylon junglefowl, in being confined chiefly to one large
island.

It is a bird of the drier coastal belt, extending inland along the lower valleys, and
in some cases ascending the mountains to a considerable height, but always in places
where the configuration of the land results in a lessened degree of humidity. I have no
reliable record of its occurrence over twenty-three hundred feet, and at this point it is
very rare, and shifts downward at the beginning of the rains.

Besides this. altitudinal, the species has a marked longitudinal distribution in Java.
It is found in greatest abundance on the east and north-east coasts, becoming more rare
- aS we approach the western end of the island, where in many places it is altogether
absent. As one of many examples of the data on which I base this statement, I found
that on most of the big estates in the Preanger the red junglefowl is well known, while
the Javan bird had never been seen or heard of. This is in country lower than one
hundred feet elevation, from the sea-coast to a distance of forty miles inland.

This eastern concentration of numbers becomes an actual centre of distribution
when we add to this area the two easterly-stretching chains of small islands: Madura
and Kangean to the north, and Bali, Lombok, Sumbawa, Flores and Alor to the east.

PHOTOGRAVURE 36

PAUNT Sean ROOSTING ISLAND OF THE JAVAN JUNGLEFOWL

Tuts bird haunts the semi-arid coastal lands of Java, but, not satisfied with its proximity to the
booming surf of the tropical sea, I saw two cocks*and a hen walk down to the beach, catching insects as
they went, and after some hesitation rise into the air as easily as quail, and fly out to a tiny mangrove
islet a hundred yards off-shore. Here they found a safe roost for the night, returning to their feeding-

grounds in early morning.

“iwotaronut

oft ob Wintizorg:

eb aosenl ynidotss ;

ovoighicin Yait's OF He yi bus disup i236: ini 9
-gnib3ot Si) ‘Or dniaiaet adtdia Sat ‘bruot yotlt

PHOTOGRAVURE PLATE 36

HAUNTS AND ROOSTING ISLAND OF THE JAVAN JUNGLEFOWL.

JAVAN JUNGLEFOWL 251

The sixth and last bit of dry land inhabited by the Javan Junglefowl is the isolated islet
of Bawean, due north of eastern Java.

Rather curiously, the distribution of this Junglefowl in the small island of Madura
is similar to Java—the birds being found only in the eastern part.

GENERAL ACCOUNT

To the visiting ornithologist, Java is one of the most difficult countries in the East
in which to get orientated for any definite work. The island is so densely populated
that it would seem a matter of but a few casual inquiries to learn much of the habits, or
at least of the haunts, of so conspicuous a bird as the Green Junglefowl. I had indeed
no lack of proffered help, but all of so indefinite a nature that it was more of an
embarrassment than assistance. The very names were most confusing, all, at various
times and places, being interchangeable, both the terms for the red and the Javan
Junglefowls and the hybrids between the latter and domestic poultry. When one was
certain that one had mastered the local names for these species, an enthusiastic
Dutchman or Javanese would go to great pains to point out some water fowl as the
bird intended. Many well-intentioned people sent me such a résumé of the Junglefowl’s
habits as the following : “They nest in holes in trees, laying eighteen eggs, white, spotted
with red. The round eggs produce hens, the long eggs only cocks. They fight all day
for their food. They never moult their feathers !” !

Our ignorance of the Javan Junglefowl has been so great that I was particularly
anxious to make our knowledge of the life history of this interesting bird as complete as
possible, especially as with the increasing density of population it cannot survive many
more decades. In English ornithological literature we find practically nothing, and in
the Dutch reports the facts are so vague or so conflicting that I have been able to place
no reliance upon them.

There is no doubt that the Javan Junglefowl does occur in the wooded mountains
where I have myself seen it, but it is of uncommon occurrence, and this rarity is not
because of the difficulty of finding it in the dense forests, but an actual status. In the
interior a favourite haunt is a deserted coffee plantation, partly overrun with jungle.
As we approach the drier coastal area, especially towards the east, the bird becomes more
abundant, and in several different places, within sound of the waves beating on the coral
reefs, I have found it in large numbers. In such localities one is at once struck by the
general superficial resemblance between the country and the corresponding haunts of
the Ceylon junglefowl. Here, in the region most favoured of the Javan bird, is no hint
of jungle. Leaving behind the lofty, humid, dense-jungled mountains of the interior,
we find a narrow coastal belt of low, rolling, or level country, covered for the most part
with a stunted, semi-desert vegetation. So much alike is all this region, that an account
of one locality, that in the vicinity of Patjiran, may well stand as representative of the
typical home of this Junglefowl.

My studies of this species began early in September. At this season the whole
earth cried out for the rains which had not yet begun. The rainy season is usually
November, December and January, but there is often more or less condensation
of moisture in October.

252 A MONOGRAPH OF THE PHEASANTS

Leaving the rest-house, or pasanggrahan, 1 made my way through cactus hedges
and across ploughed fields, and climbed a steep limestone ridge three or four hundred
feet high. At the summit was a huge, balanced boulder, and from its rounded, weather-
worn back I could overlook many miles of country. Here I began my systematic study
of the wild life of the Javan Junglefowl, after having wasted weeks of fruitless search in
the west of the island following up false reports.

These limestone ridges are very characteristic of this north-eastern coast, forming
sometimes low lines of hills extending inland, often at right angles to the shore.
Between these run small streams, and up to the very slopes of the ridges almost every
patch of ground is given over to rice, tapioca, ground bean, Indian corn, or castor oil.

In the case of most of the birds of the family of pheasants treated of in this
monograph, one is concerned chiefly with their wild surroundings, but the Javan coast
is almost a solid band of cultivation, and we are forced to link this Junglefowl very
closely-with this human environment. Farther inland a few stragglers may live in
comparative isolation, but here, in its typical home, one is never out of sight of the native.
fields. The dull, native Javanese think of little else besides their fields, and crave no
flesh besides their dried fish, so the Junglefowl are seldom disturbed.

At this season the grass was dry, the bushes and vines crackled at a touch, and even
the cactus pads looked yellow and shrivelled. As I climbed the ridge, the sun beat down
with terrific force. Even the butterflies—the yellows and the reds—often sought the
shade of tiny stones. Everywhere on the ridge the steel-grey weathered limestone
cropped out in a myriad points and sharp edges. One’s shoes were soon cut up; a
single sweep of the butterfly-net reduced it to tatters. For the vegetable life, too, as if
following out some law of this region, Nature provided a thousand spikes and thorns
and briers.

Toward the summit of the ridge I began to feel the cool sea breeze, which brought
no moisture, but which, because of its very dryness, made the great heat easy to endure.
The rocks stood up more boldly the higher I ascended, and great water-worn fissures
and crevices appeared. In one place I found a well-worn little path, feathers, and
abundance of Junglefowl sign. Here I could look down and inward twenty feet or
more, and see the light from the other side of the ridge shining through. So somewhere
in the cool ramifications of this rocky maze these birds, as I subsequently made certain,
spent the heat of the day, and here some of them roosted at night.

From his lookout on the summit of the ridge, a Junglefowl, emerging at dawn from
this roosting-cavern, would see the steep slope dropping rapidly away, a mass of cacti
draped with a tangle of briers; stubby treelets like Japanese stunted conifers; and
aromatic shrubs and herbs, smelling of mint and pennyroyal. Like a myriad toadstools,
the sago and cocoanut palms dot the fields, and now and then a feathery wisp of giant
bamboo, at this distance appearing exactly like a graceful clump of ferns. Winding up
and down the slopes, enclosing only fields of pointed coral rock, are frail stone walls,
built of heaped-up bits of angular limestone, all half-hidden by sprawling cactus.

Farther to the west, in a broad, low valley between two ridges, I found that Jungle-
fowl abounded. And here I spent many hours, concealed either in my umbrella tent, or
behind a sheltering blind of cactus. At early dawn the birds would work down from
the ridges, in small parties of two to five, passing in single file through the corners of

PHOTOGRAVURE 37

HAUNTS, OF PEE fAVAN JUNGLEFOWL

al and stony fields covered with a scanty growth of grass; spiny
ional palm or bunch of feathery bamboo—such is the home of
In one place I found a number roosting in a limestone cavern
rds make their way to the more fertile spots near the creeks and

crops of ground pea.

LIQUUL WAV AL: aH

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; ebay : ig as
Mmovéa Sagleorul §.mt peileoor iad reauvn.

o Qheepn of the Gulierliner veucish Bicwote tien
, si 1g!

PHOTOGRAVURE PLATE 37

aS

, ROOSTING CLIFF AND HAUNTS OF THE JAVAN JUNGLEFOWL

JAVAN JUNGLEFOWL 253

the fields. In the lowest part of the valley was the growing rice, and here they would
scurry swiftly along the enclosing embankments, frightening up the large doves as they
went. Now and then one would take a low, short flight across a stretch of rice, the
passing shadow of the bird sending out a flurry of scaling, twittering sandpipers.

Beyond the rice-fields were ground pea and Indian corn, and after hesitating a while
in the dense tangle along the fences, the Junglefowl would gradually spread out and
feed. This tangle was composed chiefly of small aromatic plants and the orange-
flowered lantana. ‘The cactus disappears before we reach these low, moist places.

Many of the Junglefowl spend the day in the shade of these tangles, and come out
again in late afternoon. A few return when the sun gets high, to the cactus-guarded
limestone caves on the ridges, to come down again in the evening for water and a short
period of feeding. A few hungry youngsters may be found feeding out late in the
morning in the hot sun, but evening is the time when all are visible. In a short walk
across two fields, I disturbed thirteen, shooting three in the plumage which I wished
to study.

Early in the morning, even before I had reached my place of concealment, I
would find a shrike in the top of almost every conspicuous bush, while bulbuls
babbled everywhere from the thickets. The shrill voices of little Java boys driving
cattle and goats would reach me from the dusty road. One or two native roosters
would crow in their lazy, blatant way, then, from half-way up the ridge, there would
come the sharp, crisp, virile Chaw-aw-awk ! of the Green Junglefowl, and I knew
the splendid birds were on the move. The sun rises from behind a bank of cloud,
and a half-dozen white herons fly past up the valley. Overhead, watchful above a
green, unhealthy-looking pool, sits a little maroon kingfisher, perching motionless
until the Junglefowl have passed with low clucks on their way to the tangles at
the edges of the valley fields.

Such in general is the haunt of these wild fowl. But unless we knew to the
contrary, we should never suspect a Junglefowl of inhabiting such a dreary, rocky
waste. Having nothing to fear from the natives in the way of guns, and, at least
in this part of the country, but seldom trapped, the birds were not unusually wary,
and when they were feeding, one could, with but little woodcraft, approach to
within range, whether of gun or of field-glasses. By locating their general range
and making as thorough a canvass as possible on one morning, I learned that at
least twenty-seven birds were living on or between the three limestone ridges nearest
my headquarters. Besides which I had shot eight others. There was, then, a total
of thirty-five Junglefowl which roosted or fed within considerably less than a square
mile of territory, and probably many others escaped my rough census.

In September, these birds were in pairs or families, at least as regards their
roosting habits. In the daytime, when many of them would drift down into the
cultivated valley lands, there would at times be six or eight birds feeding close
together, half of which might be adult. But when they separated, never more than
a pair of old birds would go off together, either alone or with a following of well-
grown young of the year.

I saw considerable evidence that the same individuals roosted in the same place
each night, and worked over much the same ground during the daily feeding, and I

254 A MONOGRAPH OF THE PHEASANTS

am inclined to think that the home range of these birds on the coast is extremely
limited. On the other hand, forest wardens, the only reliable natives with whom I
came in contact, informed me that in the rainy season the number of Junglefowl
was augmented by birds from the interior, which came down from the hills for
some reason toward the coast, and retired at the beginning of the dry season. As
even at the height of three thousand feet there would be no temperature change
sufficient to account for this general shifting, we must credit it to a direct food
stimulus, or a desire to escape from the excessive humidity of the hills at this time
of the year.

Although the breeding season had scarcely begun, I found the cocks very voluble,
crowing not only in early morning, but even when feeding, and in the presence of
other adult cocks. These would sometimes crow af one another, the challenge
and defiance ending, however, with the vocal outburst. Later, the affair would
doubtless have entailed more serious results. .

I was able to master the essentials of the wild fowl’s vocabulary from my blind.
The note of content, when lazily feeding, or when stretched out on their side flicking
the dust over and through their wings, was a slowly uttered, drawn-out wah, wih, wak,
wak. ‘Nhen several captive cocks were placed in their respective baskets close to
one another, they would talk for an hour at a time, the tone being a most irritating,
rasping drawl, which seemed never to cease, but as soon as one bird got out of
breath, it was instantly taken up by a second, and so on ad nauseum. When
suspicion of danger came to the wild birds, and the alarm brought them to full
attention in readiness to escape, the note was the syllable chof/ chop! chop! or
op! op! uttered many times, sharply and in quick succession. When in this
emotional state, the cocks held their head high, the position distending the brilliant
throat wattle into a gorgeous sheet of colour. The tail was, at the same time,
lowered, until the feathers fairly dragged on the ground. Another mood, apparently
when the bird is certain of impending danger and all too willing to escape, but does
not know from which quarter it threatens, is indicated by a series of disagreeable,
shrill, metallic tones, drawn out like the content note of a domestic hen, but of
the same timbre as the despairing wail of a captured fowl: awk-awwwwk-aaaawwwk !
The note of utter despair when a bird is cornered and makes a final wild effort at
escape, or when a captive bird is seized by the legs, is a long-drawn-out wail, almost
peacock-like, louder and quite different in quality from the last call described,
au—waaaaak ! and uttered only once or twice, after which the bird is silent.

The notes of the female are somewhat different. The call-note, which is usually
answered by other birds within hearing, sounds like chak, chak, chak, repeated
slowly six to ten times. The call of suspicion or uneasy curiosity is a high, strident
ak-kak-kak-kak-kak-kak, kept up at a rapid rate from ten seconds to half a minute.
I was told by natives in widely-separated parts of Java, that the hen invariably
made an outcry when she left her nest to feed—whether after laying an egg or not,
I could not learn. This note was given me as kowak, kowak, kowak, kokOwak, and
such was the general agreement, independently confirmed, that I am inclined to
give it some credence; at least I present it for what it is worth.

The imitations of the crow of the wild cock, which were given me in various

JAVAN JUNGLEFOWL 255

parts of Java, seemed to have: run the gamut of the vocal utterances of all the island
avifauna, from the hoarse croak of a cormorant to the scream of the peacock. The
Sudanese name for the Junglefowl is onomatopoetic, spelled in many different ways,
but most commonly Tjangehgar. To the natives the crow appears to sound some-
thing like “jang-eh-wear. This crow is three-syllabled, but with no decided break
or stop as in that of the common fowl. It is a high, shrill, metallic vocal explosion,
chaw-au-dwk ! as well as I can write it. These syllables are the result of listening
to, and recording on the spot, the crows of about a score of wild birds. Usually
the notes are on one tone, sometimes an individual will slightly raise the last two
syllables, or all three may form a gradually ascending scale, separated hardly by
half tones.

When neither rival nor hen is in sight, the tail droops even more than usual,
the body is held upright, the head stretched ae and forward, but with the
wattle only slightly extended.

The Javan Junglefowl are strong on the wing, although, like the other species
of the genus, they will not use these organs unless forced to, but trust to their
swiftness of foot to take them out of any ordinary danger. Several times I have
cornered one or more birds on a steep cliff and made them fly into the valley. They
rise easily, beating rapidly at first and then descending in a long, scaling curve to the
fields below. In the case of the birds which roosted on the islet off shore, there was
a voluntary flight of seventy-five yards twice each twenty-four hours, over the shore
and the water, sometimes against a stiff breeze, which made it necessary for the
birds to beat very rapidly and continuously the entire distance.

The general carriage, of course, varies with the speed and the mood of the
bird. The tail is held much lower than in the domestic cock; in the normal
walking position, the long central feathers sweep the ground, drooping on each
side, so as to hide completely the real tail-feathers. When running at full speed,
the head and tail are almost in alignment, the latter streaming straight out behind.
In crowing, the tail may be held fairly high or drooping, and when approaching a
rival, or beginning to display before a hen, this organ is raised to the usual carriage
in a domestic cock.

Even a slight lowering or raising of the head makes a remarkable difference in
the general appearance, owing to the respective retraction or distension of the throat
wattle. The comb is, of course, always displayed, but whereas usually only a bit of
red skin is visible above the feathers of the neck, when the bird stands fully erect
the blue and yellow areas of skin become very conspicuous, equalling the scarlet of
the chin region in the brilliance of their hues. |

DAILY ROUND OF LIFE

As in so many of the tropical pheasants, termites or white ants enter largely
into the diet of Javan Junglefowl. Every dry and rotted stump on the slopes of the
limestone ridges sheltered large numbers of these insects, a few pecks of the bird’s beak
being often enough to expose hundreds. Four birds examined as soon as shot, collected
at different times and places, showed that they had been feeding as follows :—

250 A MONOGRAPH OF THE PHEASANTS

A. Male, adult.—Fifty-two termites, 1 black ant, 1 small hemipterous insect, 6 seed
cases of /Zyftis sp.,and many berries and leaves of the widespread Lantana
mixta.

B. Male, adult.—Several hundred termites, 6 black ants, 1 mosquito, 2 small
spiders, 1 green coleopterous grub, 15 seed pods of the ground pea, several
hundred seeds of weeds, chiefly a harsh spiny grass, Roffbadia sp.

C. Male, juvenile-—Forty-eight termites, many leaves and small white flowers and
several hundred seeds of weeds and grass (Rottbadia).

D. Female, adult. About 200 termites, 34 large black ants, 1 coccinellid beetle;
the juice of cactus fruit, many good-sized pieces of tapioca root from the
whittlings of a native woman in a field.

As in the coastal district of Ceylon, the flora of this part of Java was, in a sense,
a false one, the A/yffzs and the cacti, forming such dominant characters, being both
introduced by mankind from South America. The aromatic character of both the
Flyptis and Lantana not only scented the air when one pressed into a thicket and
bruised some of the leaves, but was transferred to the flesh of some of the individual
Junglefowls, adding a not unpleasant natural spice. An abundant source of food was
the fruit of the cactus. The blossom of this imported plant was pale lemon yellow
with a tinge of orange, but the succeeding fruit or “ prickly pear” was stout and rose-
red, containing a very juicy pulp of the same colour. I have shot more than one bird
whose bill, forehead, nostrils and chin were stained red from the juice of this fruit.

Besides the above detailed examples I have found representatives of every principal
order of insects in the crops of these birds, in addition to the remains of many
marine or littoral creatures such as small crabs, shrimps and mollusks.

The roosting habits of the Junglefowl were of great interest, and I spent consider-
able time in tracing out the various places where the birds spent the night. I found
three separate roosts on low branches in a mass of cacti and other thorny plants, where
one or more birds sought safety from nocturnal enemies. Another bird, or perhaps a
pair, as the sign seemed to indicate, roosted on a low, sweeping bamboo which had been
bent far over, near the foot of a cliff, and partly shielded from above by a dense growth
of grass which sprouted from its precarious hold in a niche of the cliff side.

I have already alluded to the strange introduction which I had to this bird, the
sight of three—two cocks and a hen—flying straight to sea for some seventy-five yards
to an isolated mangrove islet. It may be that this habit was the result of the birds
being once driven off shore by some sudden danger at low tide. I noticed that in the case
of both these and other individuals, much time was spent about the tidal pools search-
for small organisms. At low tide half the distance to the island became bridged by
bare coral reefs, so that if the birds first found their way to the mangrove islet at low tide
they would have had much less distance to fly than when I saw them at sunset taking
off from the very edge of the shore itself. It was the more remarkable that all three
birds were young of the year, evidently of the same brood, whose wing-feathers were
still in moult. This I ascertained by examination of one of the cocks. The other birds
were not disturbed by the loss of their fellow, and continued to fly out as usual night
after night.

JAVAN -JUNGLEFOWL 257

The last roosting-place which I shall mention was almost equally striking and
unexpected. I have already spoken of its location, an underground cavern near the
summit of one of the limestone ridges. With the aid of an electric flash lantern I once
made my way to this spot after nightfall, and dropping a bit of stone into it I awakened
the inmates and heard the unmistakable voices of several Junglefowl, but could see
nothing. One day, by means of much painful squeezing, I managed to enter the outer
part of this cave, and by a most uncomfortable twisting upside down I could see into a
still deeper portion. Here were several broad, sharp-angled ledges, and the feathers
and sign beneath them showed that they must have been used for a long time. They
were so isolated from the floor that nothing without wings could have reached them,
and here the birds spent their nights in safety. Scores of tiny bats were hanging in
festoons on the walls, so that the cavern afforded a shelter to wild creatures both by
day and night. Judging from the number of Junglefowl which every afternoon made
their way up the slopes of this ridge, there must have been at least a dozen which found
nightly shelter somewhere near the summit. The only reasons, however, that I have
for thinking that most of these may have roosted in the cave are, first that I was unable
to find any other roosting-place on this entire ridge, and again judging by the abund-
ance of feathers and sign in even the partial view which I could get of the interior of
the cave.

The Junglefowl of the coast seemed to be peculiarly isolated. The other birds of
this vicinity had nothing in common with them, and if they were seen anywhere in the
neighbourhood of one another the association was of the most casual character. The
natives, as I have said, bothered the birds in this region hardly at all and apparently
knew but little of trapping. Their four-footed enemies are very numerous, and the
mongoose, martens and snakes doubtless make life one great need for vigilance and
watchfulness. Wild boars are said to do much damage in ferreting out the eggs and
devouring them. It was doubtless the fear of such enemies that drove some of
the Junglefowl near Patjiran to utilize such unusual roosting-places. It was rather
significant that the forest guard near this place told me that it was in the rainy season
that the small four-footed carnivora are most abundant and do most damage to
poultry. This was also the season when the Junglefowl from the interior come toward
the coastal lowlands.

I was not fortunate enough to obtain any insight into the actual tragedies of life of
these birds. Often, however, while I was watching them through my glasses quietly
feeding, a nearby rustle would send them fleeing headlong through the tangle, some-
times directly past my blind. Again, when the wind filled the air with rustling sound
of leaves and bamboo stems, the birds would pay no attention whatever. I could
never discover the author of any of the sounds which alarmed them so. Only when I
myself came upon them suddenly, or pursued them to the edge of some small patch of
cover did I see them take to wing to escape. At all other times their sturdy legs were
trusted to carry them into safety. I never saw females or young birds in their native
haunts squat to avoid observation. When confined in small native quakes or baskets,
and alarmed at one’s approach, the male, if a newly-caught bird, would invariably
dash madly about and try to force his way out, while the newly captured females often

squatted until one had approached quite closely, when they too fluttered wildly.
VOL. II orp,

258 A MONOGRAPH OF THE PHEASANTS

HOME LIFE

The breeding season varies considerably in the various portions of the bird’s
haunts, and even in one locality several months may intervene between the laying
of neighbouring pairs. I believe that this and not a double brood explains the
presence of chicks of various ages in any one place. It usually corresponds with some
portion of the dry season, from June to November, more often in the first month or
two of the East Monsoon, for then the abundance of insect life and the newly sprouting
plant life is propitious for the rearing of the hungry young birds. In Patjiran the
extremes of the breeding season must extend at least from June until November,
judging by the ages of the young birds which I secured. In the interior they are
reported to nest usually in June, and I believe this is correct.

The cocks are said to fight fiercely during the breeding season and assiduously to
court their mates. Their method of courtship does not differ from that of the red
junglefowl, except that the head is brought more prominently into display. They
seem to be fully aware of the beauty of their comb and wattle, and when the cock is
strutting before the hen and flattening himself obliquely toward her, the head is always
stretched out and leaned well in her direction. At such times the wattle is stretched
to the utmost, and temporarily the head of the bird appears merely a small median
connection between the two great parti-coloured sheets of skin—the comb and
wattle.

I have seen the Junglefowl only in pairs, and I have watched many of these day
after day. Other observers report that the species is sometimes polygamous, a cock
having as many as four hens in his harem.

The nest is usually a hollow in the ground hidden amongst dense vegetation of
some kind. Several, however, have been authentically reported as being placed in the
shelter of a clump of parasitic ferns growing at some height against the trunk of a tree,
and again, more than one has been found in the heart of a tree-fern’s top, surrounded
by the curving fronds, with a lining of the soft red down from the stalks of the
ferns.

The only nest which I found was one which had retained its traces from the
preceding breeding season, preserved by the dryness of the air and the security of its
location. I had made my way to a new ridge of limestone a mile or more inland, and
with great difficulty had forced a path through briars and cacti to a sheltered wall of
jagged rock, the whole surface of which was gutted and sculptured and whetted to
razor edges by the wind and water. Close above a ledge, breast high, several large
oak-leaf ferns were growing, strongly rooted to the face of the rock, and just beneath
one, on the ledge itself, I found considerable old sign, weather-worn feathers in the
deep crevices, and a number of pieces of dried egg-shell. Having satisfied myself that
this was without doubt the remains of a Junglefowl’s nest, I broke down several thorny
plants in order to set up my tripod. A thousand bits of flame burned my flesh, and
regardless of sharp rocks and cactus needles, I leaped down from the ledge covered with
ahost of fireants. After I had freed myself from them I carefully approached, ‘but an hour
afterwards the whole ledge was still alive with them, and I had to content myself with
a distant photograph. Thus was the nest of one Junglefowl protected from disturbance.

JAVAN JUNGLEFOWL 259

It is not an uncommon occurrence for tropical birds of many species to thus gain
protection, although in this case the association may have been accidental. They them-
selves seem to be immune, unless some accident should occur such as the breaking of
an egg, when the terrible ants, attracted by the contents, would probably force the bird
to leave her nest. And woe to any young birds which happened to be touched by the
broken yolk.

This species seems to deposit an unusually large number of eggs, six or twelve,
with the node of average about eight. This is the consensus of statement of many
foreign plantation owners whose facts in other departments of knowledge I have found
to be quite reliable.

I know of no greater number of young seen with their parents than four, and
several times I have observed what were apparently a brood of the year numbering only
two. The voice of the chick does not differ from the plaintive peep of a domestic bird.
The young remain with the old birds for at least six or seven months, feeding and
roosting in close association with them. The number of times when I saw both parents
with their broods leads one to suspect that monogamy is a more common habit than
polygamy. Termites form the staple food of the very young birds—at least, this was
true in the case of three nestlings which were sent to me in spirits for study.

From what I saw of the Javan Junglefowl in the small island of Madura, it differs
in no particulars from the same species on the larger island. It is found only in the
eastern part, and I found it to be especially abundant where there was much outcropping
limestone, and less so where this gave place to earth and jungle. I saw a family of
birds with an interesting history. A pair had been kept for several years in semi-
captivity by the proprietor of a big, rambling hotel at Manding. They disappeared for
a few weeks, and then returned to feed as usual on the scraps from the kitchen,
accompanied by three young birds—another proof of monogamy.

The eggs are very rare in collections, and the published descriptions and measure-
ments show such discrepancy that it is probable that the eggs of Gaus gallus or of
hybrids have sometimes been confused with those of varius. Four eggs in my
possession, taken near Patjiran, are buffy-white with a faint tinge of yellowish, of a
rounded oval shape, the shell being glossy, with very inconspicuous punctures. They

measure 43 x 34, 44 x 34, 45 X 35, and 46 x 35 mm., averaging 34°5 x 44°5.

RELATION TO MAN

The relation of the Javan Junglefowl to mankind is of a most peculiar character.
The birds are of no direct economic importance. The contents of their crops show that
on the whole they are more useful than injurious, feeding chiefly on noxious insects and
the seeds of weeds, and taking but small toll from the grain-fields. The Javanese are a
dull, lethargic race, and possess none of the interest in their surroundings which is
exhibited by many other races of Eastern people. So that both in the coastal lowlands
and in the mountains but little heed is paid to the wild fowl, the plantation owners alone
shooting the birds now and then. There are no laws protecting them, but the fear of
native uprisings has put such a high licence price both on bringing a gun into the
country and carrying it when landed, that the Junglefowl have little to fear from powder

260 A MONOGRAPH OF THE PHEASANTS

and shot. One finds, here and there, captive birds carefully tended and often tamed.
These, however, are exceptional. The great desiderata among the dl aveuicee are hybrids
between the wild Junglecock and a domestic hen.

In spite of reports which came to me frequently, I found no direct proof that the
Javan Junglefowl, in a wild state, interbreeds with the native fowls. Even in captive
birds this very seldom occurs, and several ingenious methods are resorted to in order to
produce the greatly valued hybrids. In some cases the Junglecock is placed in a quake,
or round beehive-shaped basket, from which the bottom is removable, and is then
excited by the gradual approach, in a second basket, of a wild hen. To allow the two
birds to mate would be quite useless, for the wild hen seldom or never lays in captivity.
When the cages are close together and the cock is crowing and showing off his brilliance,
exerting every effort to attract the attention of the female, she is suddenly replaced with
a domestic hen, coloured as similarly as possible, and the cock is allowed access to her
cage. Were this strategy not resorted to, he would have nothing to do with the
domestic bird. Most of the inhabitants of the small Kangean islands seem to be
engaged in this curious industry, and the numerous hybrids which one sees in all parts
of Madura and Java come from these places. Hardly a house is to be seen which has
not from two to six of these hybrids or dezsars, each in its separate basket, hung on
branches in the compound or in the shelter of some outhouse. I secured evidence
which seemed satisfactory that the Kangean islanders had been engaged in this hybrid-
making for well over two hundred years. At present about fifty def/sars are exported
to Madura and Java each month.

The more usual method of capture in Kangean is by means of a long, narrow net,
which is suspended between the bushes of a hedge and a rice-field. The Junglefowl
come down in late afternoon and pass by certain favourite openings through the bushes
into the fields. Just before dusk the hinder side of the net is lowered to the ground,
and then several natives rush with cries and much beating of sticks through the fields.
This frightens the birds, which never rise if they can avoid doing so, but with lowered
heads run swiftly for the shelter of the bushes. Well knowing that their only safety lies
in this direction, they strive to push on even after they encounter the net. When well
entangled the front of the net is dropped and the birds are caught.

The usual food of both wild birds and hybrids in captivity is bananas and unhulled
rice, while cooked rice is now and then given as a special delicacy.

The vazson a’étre for these hybrids and their subsequent history is as interesting as
their origin. Whatever characters of pattern and pigment are dominant or recessive,
that of the voice is decidedly in the former class. I do not recall hearing any wild bird
of equal size, unless it be the chachalaca in Mexico, which excels these hybrids in vocal
power. They are able to give utterance to a sound, a scream, which must carry for
nearly a mile. There is no cadence, no rhythm, no pleasing quality whatever. It is
simply a blatant utterance, as strange as the idea of a hybrid itself must always be to us.
Some excel others in power and endurance, and there is likewise a great variety in
colours, due to the inherent strain of the mother. It is difficult to think that even the
Javanese, with their ideas of music, so unlike our own, can derive pleasure from this
sound, but be that as it may, they find its quality and continuity, characters well adapted
for gambling upon, and prize fowls are matched with as great interest and excitement

PHOTOGRAVURE 38

HYBRID JAVAN JUNGLEFOWL

owl do not interbreed with the native poultry, but in captivity the two species will
me of these hybrids are huge creatures, with enormous pendant combs, and beautiful
nall and bantam-like with absurdly short legs. They are great favourites with the
on triangle perches or in ingeniously woven quakes or cages. The dominant
is the voice, which lacks cadence or definiteness, but consists of a scream which
. mile. This is the stimulus to much betting, and owners of powerful-lunged birds

on the vocal powers of the rivals.

tw: Pea) 92. si ion

Repiul hitrowog Ae: erScWee | ee “ating sabi ‘OF li cen

D JAVAN

JUNG!

LEFOWL.

PHOTOGRAVURE PLATE 3O

JAVAN JUNGLEFOWL 261

as are fighting cocks in other countries. The hen hybrids are killed, as they are of no
value, but the cock birds when full grown are placed in quakes and are peddled in all
the Javanese towns, a basket on either end of a pole balanced over the owner’s shoulders.
Thus they are hawked about. The common kind—common in possessing no especially
favoured type of plumage and only a mediocre crow, are sold for six or ten guldens.
Pure white birds with better (or worse!) voices bring fifteen, while for a black bird of
equal vocal ability thirty guldens are asked. Bekisars which have the head, comb and
wattle like the ajam oelas or Javan Junglefowl, and the remainder of the plumage of the
normal bekisar type, are very rare, and are known as vatum#a. One in Madura, just
brought from Kangean, was offered me for seventy-five guldens. A gulden is equal to
4o cents. or 1s. 8a. Connoisseurs recognize and appreciate fine points which mean
nothing to an outsider, and I heard of a native chief who paid as much as two hundred
guldens for a greatly prized bird, while there is a record of six hundred guldens paid for
two birds, jet-black as to plumage, comb, face, wattles, legs, feet, and even iris, with
very strong voices.

A poor native who possesses a hybrid must content himself with keeping it near or
in his house in its quake, and there listening to its constant outcries, or matching it and
backing it against the vaunted crowing of some neighbour's fowl. But the native chiefs
make the most of their high-priced bird’s vocal ability.

In Soemanep, Madura, the last independent sultan died in 1885. His son has been
made Regent under the Dutch authority, and this prince, whose name is Roden Ario
Manahoe-Hoesoemo, is a connoisseur of hybrids or bekisars. He told me that the
character most valued in the crowing of these birds is the loudness and the piercing
quality; the crow must also be long-drawn-out and monosyllabic.

A special apparatus is used to induce these birds to exercise their lungs. This
consists of a very tall bamboo, which is erected in the compound, with a primitive sort
of pulley as near the top as possible. The bekisar in its basket is attached to the end
of a rope and pulled up to the pulley high in air, where it remains during the day,
crowing lustily hour after hour. The prince had a prize bird pulled up for my benefit,
and the bird began its crowing while the cage was whirling around on its jerky ascent.
A village sometimes presents a curious aspect, and the visitor is startled by the
sight of the baskets suspended to scores of swaying bamboos, while one’s ears are
assailed throughout the day by the terrible raucous outcries from mid-air. This loud
and penetrating crowing, while primarily valued for the means of gambling, is supposed
to bring good luck to the house over which the vibrations pass. The wild Javan
Junglefowl are not admired, because their crow, while of the correct timbre, is very
faint compared with the lusty cries of the big hybrids.

I learned through interpreters, that among the poorer classes another standard of
vocal excellence is rife; where birds with a short, abrupt crow, more like that of the
wild Javan bird, but with a persistence which would drive a white person insane, are
valued over other individuals. In a smaller number of cases I found that birds with
voices of indifferent power and persistence were kept solely for the beauty of their
plumage, as indeed, in this respect, they far excel any breed of native poultry.

The hybrids soon become tame, and are allowed to run freely about among the
domestic poultry. The plumage of these hybrids is so unlike the colours and patterns

262 A MONOGRAPH OF THE PHEASANTS

of either of the parent species that individuals which found their way to Europe in
former years were considered to be distinct species. Thus we find such synonyms as
Gallus temminckiu Gray; G. eneus Cuvier; and G. violaceus Kelsall. The most
striking character is the metallic gloss of the upper plumage, fringed with golden yellow.
Even in the pure white birds this iridescent fringe persists, and is wonderfully beautiful
in the sunlight.

As regards the fertility of the hybrids, our ideas would be chaos if we had to depend
upon the statements of the Javanese. Throughout the west of the island one hears
everywhere not only that the bekisars are sterile, but that they will not survive a
single mating with a domestic hen. In the east such ideas are laughed at, and the
birds are said to cross readily. I have proved the truth of the latter theory to my own
satisfaction, and have bred many kekoks, as they are called. On the other hand, this.
second generation in Madura is known to many natives as dehikkos, which name in
Java is applied in the west to the Red and in the East to the Javan Junglefowl. Such
is the confusion of names, coupled, as I have said, with widespread ignorance of the
occurrence, appearance or habits of the wild birds. This second generation of cocks,
between the bekisars and domestic hens, reverts to the red junglefowl type of plumage,
or at least loses much of the green and violet iridescence of the upper plumage, while
retaining the large size of the bekisars. I shall have more to say about this farther on.

As we have seen, the Javan Junglefowl bears captivity remarkably well in its native
islands. Wild as the cocks naturally are, they are susceptible of taming to such an
extent that they may be taken out of their baskets and handled without showing any
signs of fear. The aberrant general character of this species makes it certain that this
is merely an incidental characteristic of the bird, and has nothing to do with the
tameness of domestic poultry. No one, I think, has seriously considered the Javan bird
as sharing in the direct ancestry of our breeds of fowls.

These birds are not common in zoological gardens, but sailors occasionally bring
cock birds to Europe and America. Of eleven Javan Junglefowl which have been kept
in the London Zoo, one lived for five years and two months, while the average length
of life was about two years.

The difficulty with which hybrids are produced in Java diminishes under other
conditions. In the New York Zoological Park adult Javan Junglefowl cocks have
mated readily with bantam hens, and as many as five out of a clutch of six eggs have
proved fertile and the chicks reared to maturity.

DETAILED DESCRIPTION

ADULT MarLet.—Comb, face, wattle and fore-neck bare of feathers. Ear-coverts
dull brown; top of head, back and sides of neck and upper part of the mantle, corre-
sponding to the elongated hackles of other junglefowl, composed of short and square-
tipped feathers, with the visible portion raised into a convex transverse band, the vane
itself being thus moulded into a rounded terminal ridge. This is exactly similar to the
corresponding structure in the feathers of Lofhura and Lobiophasis. The base of the
feathers is dull brownish black, part of this zone showing when the feathers lie naturally.
Just before the distal curved section, the black shows a very faint violet gloss, then

PHOTOGRAVURE 39

HYBRID JAVAN JUNGLEFOWL IN ITS AERIAL BAMBOO CAGE

Tue hybrid Junglefowl are sometimes valued as high as six hundred gulden. The. chiefs and wealthy
natives erect a very tall bamboo pole in the compound with a primitive sort of pulley near the top. The
fowl in its basket is attached to a rope and pulled up, high above all the surrounding trees, to remain
throughout the day, sending forth at frequent intervals its loud, piercing scream. It begins its call even

while the basket is whirling around on its jerky ascent.

T.&
O73 Ah) Aca te

HYBRID JAVAN JUNGLEFOWL IN ITS AERIAL ]

BOO CAGE.

PHOTOGRAVURE PLATE 59

: wes
¥ aa
Ay

JAVAN JUNGLEFOWL 263

suddenly becomes a brilliant purplish-blue. The convex band itself is a rich, glowing,
greenish bronze, the feather terminating in a very narrow fringe of black. These colours
are wholly absent from the under surface, and, of course, from their character, disappear
when viewed by transmitted light.

On the mid-mantle the purplish-blue becomes so extreme that it replaces the
proximal black in the visible portion of the feathers. The lower mantle changes rather
abruptly, the feathers losing their truncate character and becoming rather obtusely
pointed. Simultaneously the curved ridge becomes obsolete and the black fringe
increases. The purplish-blue disappears and the whole of the visible feather is
uniformly golden green. Just basal to the black fringe the vane is sometimes tinged
with bronze. The lower back and rump form another abrupt colour area, with almost
no transition pterylum. The feathers are narrow and greatly elongated, finger-like, with
rounded ends, and in colour dead black, with a conspicuous fringe all around of golden
yellow. The shorter upper tail-coverts are similar, but with a large central area of bluish
green. In the longer feathers of this series the fringe becomes obsolete and the green
gloss dominates the whole feather. The lesser and median wing-coverts are much like
the back feathers in shape, hackle-like, but the disintegrated fringe is longer and of
a rich, deep, reddish orange. On the inner greater coverts this pales to the golden
yellow of the back hackles. Greater coverts and secondaries black, more or less glossed
with bluish green. Primaries dull blackish brown. Tail-feathers black, glossed with
greenish blue, especially on the outer webs. Entire under parts dead black; under
tail-coverts slightly fringed with yellowish buff.

This Junglefowl has the long, curved, central tail-feathers, but lacks the elongated
flowing neck hackles of the other species. Its most striking secondary sexual character
‘is the pigmentation of the comb and wattle. The former is very large, with the upper
margin entire, and no traces of the deep notches of the other species. That is, the usual
comb is thus entire, but perhaps ten per cent. of the cocks show very faint serrations on
the posterior half. In one wild shot bird, for instance, I found, on close examination,
five well-marked teeth, one of which was double-notched, all near the posterior upper
angle. In three females I have observed minute serrations along the summit of the
rudimentary comb. ‘The size of the comb often results in its leaning far over to one side,
especially at other than the breeding season. The central basal part is greenish or
greenish blue, this colour changing abruptly into a violet red, which extends over the
whole remaining four-fifths of the comb. In some individuals this is almost clear deep
red; again the violet, peach-bloom sheen may cover almost all of it. The rim is usually
pure blood red. The lower eyelid is dull violet in the centre, changing into pale yellow
around the edge. When the eye is open this colouring is invisible, but when drawn up
over the eye, the lid stands out clearly against the facial red.

The whole unfeathered part of the crown, lores, the face to the edge of the feathering
all around, including the small, sub-auricular wattles, chin, throat and anterior two-thirds
of the median, throat wattle, deep blood red. The wattle begins well up between the
lower mandibular rami and extends far down on the neck, hanging downward as a large
median fold of skin. In fully adult birds at the breeding season this may be two and a
half inches in length. The free ends of the feathers of the neck, truncated though ‘they
are, reach cape-like over the bare throat, and when the head of the bird is drawn down

264 A MONOGRAPH OF THE PHEASANTS

the wattle is almost completely hidden except for a portion of the anterior red. When
courting or preparing to fight, or when the head is stretched out for any reason, the full
glory of pigmentation of the wattle excels even that of the comb. Beginning at the
lowest fold of the small facial, sub-auricular wattles, a band of intensely brilliant chrome
yellow extends downward, gradually widening, to beyond the attached end of the wattle
and almost to the posterior free end. The remainder of this organ, the lower outer
portion, is a rich azure blue, sometimes with a hint of greenish. The relation of the
three colours to one another is interesting, and also the character of the skin itself in the
respective zones of pigmentation. The yellow is abruptly marked off from the other two
colours, while the red and the blue show a considerable area of gradual transition, a
violet which appears darker than either colour. The yellow area is the most folded and
creased, the lines being transverse, longitudinal and diagonal, forming a complex
network. The red hardly shows its rather longitudinal wrinkling, while the blue is
conspicuously marked with dark wrinkles running at right angles to the edge of the
wattle. In fully adult birds the face and entire wattle are free of feathering, the last
traces being a small patch of black featherlets on the throat and wattle in the red area
close to the yellow.

Iris lemon, orange or pale straw yellow. Mandibles usually pale horn-colour. The
base of the upper mandible may be black or dark reddish and the whole under mandible
yellowish or ivory horn. Legs and feet varying from fleshy pink in the breeding season,
to quite pale whitish. Or the posterior half of the tarsus, the toes and the spurs may be
dark brown.

Length, 700 mm.; bill from nostril, 16; wing, 225; tail, 325; tarsus, 73; middle
toe and claw, 60; spurs, slender and curved, about 25 mm.:in length. Weight,
1 lb. 12 ozs.

AvULT FEMALE.—Top of the head, neck and upper mantle sandy brown, the latter
feathers with a wide, black, sub-marginal, concentric band, enclosing an area of rather
rufous tone. The remaining upper plumage and the smaller coverts are black with buff
edges and an irregular transverse band of rufous buff, many of the feathers with a
hair-like, white shaft-streak. Much of the black, especially on the scapulars and wing-
* coverts, is glossed with iridescent green. Secondaries brownish black, barred on the
outer webs with buffy white. Primaries and alula plain dull brown. Tail-feathers
brownish black, with a few irregular marginal spots of buff, separated with dark areas
indistinctly glossed with greenish.

Lores, broad superciliary stripes and the rather scanty feathering on the sides of the
face pale buff; chin and throat white. Ventral plumage brownish buff, the breast
feathers margined with dark brown, and the feathers in general sparsely and irregularly
mottled with the same colour, especially on the sides and flanks, where the dark colour
forms irregular bars.

Iris pale straw yellow. Skin of the face as showing through the sparse feathering,
and the small comb reddish pink. Throat fully feathered, but a low, permanent fold of
skin represents the wattle of the adult cock. Legs pale flesh colour.

Length, 400 mm.; bill from nostril, 15; wing, 195; tail, 115; tarsus, 58; middle
toe and claw, 48; spurs rudimentary. Weight, 1 Ib. to 1 Ib. 4 ozs.

JAVAN JUNGLEFOWL 265

Cuick In Down.—Upper parts from crown to tail, and the wing down, dark
chocolate-brown ; head, side neck and breast, brownish; line from lores through eye and
ear-coverts back to the nape chocolate or black. A creamy buff line on either side of the
dorsal brown along the sides and rump, bounded outside by a black line. Under parts
creamy white. Iris pale hazel. Bill, legs and feet pinkish white or light horn colour.

The difficulties under which one has to work in the study of such an important
member of the Javan fauna as the Junglefowl, even in its native land, is well shown by
the specimens in the Buitenzorg Museum. There were six males all told, very badly
mounted, two adult birds, one nearly so, and three immature cocks. No females, chicks,
or birds in juvenile plumage. Of the three young birds of equal age, one had yellow
glass eyes, another jet-black beads, and the third rich hazel irides. The combs were
enclosed or replaced with thin wooden pointed sheaths. If such diversity, due to lack
of definite observation, exists here in the very haunts of this insular species, we, thirteen
thousand miles away, can readily understand the total lack of accurate details up to the
present time.

JUVENILE PLumacE.—The sexes in this plumage are almost indistinguishable,
judging from the small series I was able to collect. The cocks are more heavily pig-
mented with black, the general type of pattern and colouring being, however, much like
the adult female. The feathering on the face is already scanty, and, together with the
chin and throat, is pure white. The feathers of the top of the head, neck, and upper
mantle are all normally rounded in shape, and in colour predominantly black, with a
rufous-buff fringe and a narrow shaft-stripe. On the lower mantle, back, and rump this
shaft-stripe widens and becomes a conspicuous buffy-white.

The lesser coverts are like the upper mantle, and the median like the back with the
addition of a broad, transverse band and wide tip of buff or whitish. ‘The secondaries
and their coverts are banded or spotted with whitish and rufous on the outer webs, as in
the adult female. The primaries are plain brown. The tail-feathers are dark-brown,
irregularly barred or edged with buff.

The white feathers of the throat extend well down on the neck in a \V-shape,
covering the area which in the adult is bare and occupied by the median wattle. The
ventral plumage is reddish-brown, mottled and spotted with black. On the flanks are
found faint traces of the sub-terminal concentric band which characterizes the adult
female. The under tail-coverts are black, with a fringe of buff.

The iris is pale hazel, and the spurs and comb are rudimentary. The facial skin as
seen through the feathering is flesh colour, and the legs and feet are pinkish.

First ANNUAL OR PosT-JUVENILE Mou tt.—In all the specimens which I have
seen, this moult leaves the bird in a very imperfectly adult plumage, and it is probable
that not until the second annual moult does the Javan Junglefowl ever come into its
fully adult dress.

The typical change is as follows, although no two specimens are exactly alike, and
indeed this is the most variable period of the bird’s life as regards its external characters.

The brown head, neck, and mantle feathers are replaced with black ones, with a slight
VOL. II M M

. 266 A MONOGRAPH OF THE PHEASANTS

green gloss near the tip, the fringe, however, retaining its rufous-brown colour. The
convexity is hardly apparent, and the shape is less truncate than in the adults. The new
upper-mantle plumage has only a faint gloss of green, this appearing first as two sub-
terminal foci or eyes in the centre of the webs. The hackles of the rump are transitional
between the juvenile and adult, broad, with stubby, tapering extremities and a very wide
disintegrated fringe of pale yellow. The black centre is often mottled with buff and
brown. The coverts are even less advanced, and in place of the orange red, show only a
faint rusty-yellow fringe. The upper tail-coverts are all bordered with yellow-buff, and
very imperfectly glossed with green. The tail-feathers have buff edgings, and the
secondaries a series of large, reddish-buff spots on the outer web, the whitish bars and
spots having been eliminated. The new ventral plumage shows a diminishing of the
red-brown, and a corresponding increase of black.

This is the more usual style of moulting, where a gradual transition carries the
bird only part way to the final colouring and patterning. A week one way or the other
would doubtless make considerable difference in the retention or elimination of juvenile
characters. In fact, we have absolute proof of this in the secondaries taken as a whole
in a moulting bird of this age. The sequence of moult being from the outer to the inner
feathers, the new outer feathers are well marked with juvenile characters, while as we
proceed inward, the adult colours become more and more pronounced, the manufacture
of the immature buffy giving way to the melanism of the adult.

Now and then, in fact in about twenty-five per cent. of the young cocks which I
have seen, a very interesting condition is found, usually confined to the tail and its
coverts, never occurring on the wings, but occasionally on the body plumage. The
change from youth to adult is sudden: twenty-four hours at most sees all the pigmenta-
tion altered, all the imperfect patterning of the juvenile repressed and succeeded at once
by perfect gloss and specialized shape of the adult cock’s plumage. Widespread as this
appears to be, it is assuredly abnormal, for the plumage suffers, and the majority of the
feathers soon break at the point of demarcation. In such a bird, some of the side coverts
may be of very recent growth, and hence adult in colour; other old lateral coverts are as
completely juvenile, but the recently sprouted feathers will have been caught half-way.
The terminal half of each feather is the purely juvenile, or feminine, mottled brown.
Then comes an abrupt change, and without transition appears the adult condition, in
width, curvature, colour and size of shaft and web, strong metallic gloss and all. The
sudden increase in the weight of the feather and the diameter of the shaft has resulted
in a top-heavy weakness, and all but one of the feathers affected have broken off, the
break occurring in the shaft, and not in the barbs. The point of chief interest in this is
not the details of the phenomenon itself, as it is clearly abnormal, but the possibility, as
shown by the twenty-four-hour growth of the webs, of the entire secretion of the bird’s
pigment changing within so short a space of time.

Abnormal though it is, this condition is at least understandable, but in two of the
birds in the Buitenzorg Museum, and in several which I have shot, a state of affairs still
more remarkable is found. The sequence is reversed. That is, the terminal portion of
the feather, which of course first grew out, is altogether adult, black, and iridescent
green, while the last two-thirds to appear are quite juvenile. Here the adult pigmenta-
tion must have come into action and then abruptly, within the space of a day, have

JAVAN JUNGLEFOWL 267

ceased or been turned into another channel, while the juvenile colours again flowed and
functioned.

In following the development from chick to adult, confining ourselves always to
corresponding centres of plumage, we find two paths of pigment evolution: first, sandy-
brown, red-brown, dark-brown, black, and iridescent green; and second, whitish-buff,
pale straw, yellowish, orange-yellow, orange and deep orange-red. This is well illus-
trated by the inner secondaries, which, in the retention of juvenile characters (permanently
in the female, and longer than any other area in the developing cock) are the most
generalized feathers of the plumage. They are mottled and barred with sandy-brown
and whitish-buff. If we proceed from them mantlewards, we find them leading through
feathers with more and more black until this culminates in the extreme of green
iridescence. Covertwards, the pale edges increase and change through the yellows to a
rich orange-red.

HYBRIDS

I have given what data I could in regard to the origin and uses of the bekisars or
hybrid cocks between the wild Junglecock and domestic hen. In coloration there are
two general types, derived through the mother from the red and the cream-coloured
Malay game. These two colours are always most apparent and the last to disappear on
the wing-coverts. The extreme of the red type is where the violet and green iridescence
have spread over the body until they almost or quite obliterate all other pigments.
I secured a young cock in which the red is reduced to narrow fringes on the coverts
and rump. The bill, face, feet, and legs, and most of the comb and wattle are jet black.

The extreme of the cream type is pure white throughout. I have a poor specimen,
which is whitish in general, with golden-yellow fringe on the hackles, but dirty cream
on the coverts and back, with the wings and tail part white, part metallic green.

The neck hackles of the first generation hybrids are much like those of varius, but
with narrow margins of buff or red. The sub-auricular flaps or wattles are often of large
size. The green area on the comb is usually replaced with the red, and the yellow
on the wattle with bluish white. In pure white birds this area may be sulphur-yellow.
The irides of all are pale, but orange instead of yellow. The shape and colour of the
comb are like those of the domestic cock, while the median wattle is as in varius. After
death, however, the basal part of the comb, which is greenish in vavzus, at once turns
fleshy white, while the remainder holds its deep red hue for many days before slightly
fading. Occasionally a bird will be found with both comb and wattle as in varius, and
exaggerated both in size and colour. The rarity of such types of birds is attested by
the high price at which they are held.

The general character of the crow and the power of flight are both like those in the
wild fowl. In general size the hybrids excel both parents.

The second generation are known in many places as #ekoks. This is probably
a contraction of dekékko, a name which in Western Java is given to the wild red
junglefowl. The aptness is apparent when we see that in the majority of these kekoks
the plumage reverts to the gallus type. The hackles become long and lose much or
all of the violet. While the median wattle may be well developed in life, its two
dermal walls are attached so loosely that after death it splits, and the throat shows

268 A MONOGRAPH OF THE PHEASANTS

merely a wrinkled skin, with no hint of the wattle, owing to the non-adhesion of the
two walls. —

The crow in this generation usually returns to the three- or four-syllabled call of
the domestic rooster. I was given a small, white kekok whose pedigree was:

Gallus ey 3 Cream Malay Game 2
|

|
Cream Bekisar ¢ White Japanese Bantam Q

| | |
White Kekok ¢

This bird was especially interesting as regards its wattles and its crow. Across
the front of its chin and neck was a flap of skin on which were strung five structures,
the median wattle, outside of which were two larger gular wattles, while near the outer
ends of the flap were the well-developed sub-auricular wattles. The crow was quite
like the challenge of the wild Javan bird—an abrupt three-syllabled call, but at the
end the beak of the bird remained open, and it strained ludicrously forward in a final
silent exhalation, as if giving a long-drawn-out finale to the last syllable. Its audible
crow was that of vavzus, the duration of posture and effort was clearly that of gallus !
The kekoks cannot be classified, and some individuals which I have seen vary even
assymetrically, having parti-coloured adventitious feathers cropping out here and
there.

SYNONYMS BASED ON BEKISARS

Gallus aeneus Cuvier, is one of the types with violet upper plumage, margined with
golden yellow; comb with minute teeth ; a small median wattle.

Gallus temminckit Gray, is a red phase of hybrid. All the plumage margined with
golden chestnut; six coarse teeth in comb; well-developed median throat wattle, and
small gular wattles.

Gallus violaceus Kelsall, has the violet gloss dominant; a toothed comb, a good-
sized median wattle.

EARLY HISTORY

The début of this bird in ornithological literature occurs in the form of a weirdly
coloured plate given by Shaw and Nodder in their “ Naturalist’s Miscellany” of early
but unknown date. The general characters are correct, however, and the entire comb
and median wattle leave no doubt as to the identification. They call it Phascanus
varius or variegated pheasant, and describe it as a ‘“‘black pheasant, with red front,
glossy-green neck and back, and compressed ascending tail with the coverts hanging
down on each side.” Aside from a more detailed but no more correct description, we
are told that ‘this beautiful bird seems to be a nondescript, of which the present figure
is the first that has been presented to the public. . . . The native country of this species
is perhaps not clearly ascertained, but it is probably an Indian bird.”

Temminck in his Wzstoive Naturelle Générale des Pigeons et des Gallinacés in 1813
gives us the first authentic note. “‘ Ce¢te belle espece est tres abondante dans les grandes

JAVAN JUNGLEFOWL 269

Jfovéts de lintérieur de Vile de Java, elle se tient communément pendant le jour a la
listéve des bois; son naturel est farouche, attentif a se cacher au moindre indice de
danger, tl est difficile de Capprocher pour la tuer.’ And up to the present we have
known but little more concerning the habits of this bird,

SYNONYMY

Phasianus varius Shaw and Nodd, Nat. Misc., X., pl. 353 [text].

Gallus varius Griff. ed. Cuv., III. 1829, p. 21; Lesson, Traité d’Orn., 1831, p. 492; Gray, List of Birds,
Pt. III., Gall., 1844, p. 27; id. Gen. B., III. 1845, p. 499; Sclater, List of Phas., 1863, p. 10; Blyth, Ibis, 1867,
pp. 307, 308; Gray, List Gallinae Brit. Mus., 1867, p. 40; id. Hand-list of Birds, II. 1870, p. 261; Elliot, Mon.
Phas., II. 1872, pl. 35; Kelham, Ibis, 1881, p. 3 [Singapore?]; Hume, Stray Feathers, X. 1887, p. 68 [Johore,
Malay Pen.]; Grant, Cat. Birds Brit. Mus., XXII. 1893, p. 352; Vorderman, Bijtrage tot de kennis der Vogels
Kangean-Orde, 1893, p. 204 [production of hybrids]; Hartert, Novitates Zoologicae, 1896, p. 554 [Bali, 2000 feet] ;
Hartert, Novitates Zoologicae, 1896, p. 574 [Sambawa, up to 3000 feet]; Hartert, Novitates Zoologicae, 1896,
p. 598 [Lombok]; Grant, Hand-book Game-birds, II. 1897, p. 59; Sharpe, Hand-list of Birds, I, p. 39;
Hartert, Novitates Zoologicae, 1898, p. 49 [South Flores]; Hartert, Novitates Zoologicae, 1898, p. 465 [Alor];
Hartert, Novitates Zoologicae, 1898, p. 476 [Sumba]; Vorderman, Java-Vogels, II. 1898, No. 119; Nehrkorn,
Katalog Eiersammlung, 1899, p. 194; Hartert, Novitates Zoologicae, 1902, p. 421 [Kangean Is.]; Ghigi, Mon.
Zool. Ital., XIV. 1903, p. 319 [generic character] ; Koningsberger, De Vogels van Java, II. 1909, p. 42; van Oort,
Notes Leyden Mus., XXXII. rg10, p. 111; Mitchell, Proc. Zool. Soc., 1911, p. 522 [viability in captivity] ;
Vorderman, Nat. Tijds. Ned. Ind., XLIV., p. 231 [description of female]; Vorderman, Nat. Tijds. Ned. Ind., LIL,
p. 204 [hybridization]; Vorderman, Nat. Tijds. Ned. Ind., LIX, p. 225 [description of male].

Gallus furcatus Temminck, Pig. et Gall., II. 1813, p. 261; III. 1815, p.662; Steph. in Shaw’s Gen. Zool., XI.
1819, p. 215; Vieillot, Tabl. Encycl. Méth., I. 1823, p. 364; Temminck, Pl. Col., V. 1829, pl. 4 [No. 483]; Gray,
Ill. Ind. Zool., I. 1830-32, pl. 43, fig. 2; Schinz, Nat. Abbild. Vég., 1833, p. 242; Jard, Nat. Lib. Orn., IV. 1834,
p. 184, pl. X.; Schinz, Nat. Vog.; 1853, p. 145; Bernstein, Jour. fiir Orn., 1861, p. 188 [habits and nesting];
Wallace, Proc. Zool. Soc., 1863, p. 486 [Lombock, Flores]; Finsch, Notes Leyden Mus., XXII. 1901, p. 302.

Creagrius varius Gloger, Hand-u. Hilfsb., 1842, p. 387; Ghigi, Mon. Zool. Ital., X1V. 1903, p. 322 [generic
characters].

Gallus javanicus Horsfield, Trans. Linn. Soc., XIII. 1822, p. 185 [Java].

Gallus violaceus Kelsall, Jour. Straits Asiatic Soc., XXIV. 1891, p. 167; Kelsall, Jour. Straits Asiatic Soc.,
XXV. 1894, p. 173.

END OF VOL. II

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