f, ' T^r:s^r*£ - FOR THE PEOPLE FOR EDVCATION FOR SCIENCE LIBRARY OF THE AMERICAN MUSEUM OF NATURAL HISTORY si i-n a m a SMITHSONIAN MISCELLANEOUS COLLECTIONS. MONOGRAPHS OF THE D I P T E R A OF NORTH AMERICA. PREPARED FOR THE SMITHSONIAN INSTITUTION BY H. L 0 E W. PART I. EDITED, WITH ADDITIONS, BY R. OSTEN SAC KEN. WASHINGTON: SMITHSONIAN INSTITUTION. APRIL, 1862. ADVERTISEMENT. THE present publication is the first part of a work on North American Diptera in process of preparation by Dr. H. Loew, of Meseritz, Prussia (one of the highest living authorities on the subject), undertaken at the especial request of the Smithsonian Institution.' The materials have been derived principally from the collection of Baron R. Osten Sacken, of the Russian Legation in Washington, kindly intrusted to the author for examination. As explained by Dr. Loew, the work will appear in monographs of genera and families, sufficient materials being at hand for illus- trating particular groups only, without relation to their systematic sequence. The Institution is under obligations to Baron Osten Sacken for editing the work, adding species described by Dr. Loew subse- quent to the reception of his manuscript, and for correcting the proofs. He has also added a monograph of the Cecidomyidce, a group of much interest, and one to which it was considered of importance to call the early attention of investigators. JOSEPH HENRY, Secretary S. I. SMITHSONIAN INSTITUTION, WASHINGTON, March, 1862. ACCEPTED FOR PUBLICATION, JCLT, 1861. PHILADELPHIA: COLLINS, PRINTER. PREFACE, THE impulse to write on North American Diptcra was given to me by Baron Osten-Sacken, who, first by sending me rich collections of such Diptera and finally by intrusting me with the greatest part of his own Diptera collected in North America, has enabled me to undertake this task, and, I hope, with some success. If my observations had been written in German, and published in any of our German Transactions, I should have had good reason to fear that the results would not become sufficiently known in North America, and would at all events be longer in obtaining access there. I resolved, therefore, to give them in English, and the Smithsonian Institution in Washington has added to the many proofs it has already given of an energetic furtherance of any studies relative to the natural history of North America, the liberal resolution to print rny paper on North American Diptera at its own expense. If these papers, according to my intention, contribute to the increase of the study of this interesting order of insects, the principal thanks are due to the Institution and to Baron Osten-Sacken. As for the English text, I use, it is true, the assistance and advice of a friend who is well acquainted with the English language ; should, nevertheless, some roughness occur, I beg that it may be attributed to some supplementary alterations of my own, which circumstances prevented me from submitting to him. I hope that shortcomings of this kind will be kindly over- looked, provided my descriptions be not deficient in precision and clearness. As I do not wish to remain the sole describer of N. A. Diptera, but hope soon to have many fellow-laborers, I take the liberty of pointing out briefly what, in my opinion, should be chiefly borne in mind in making and publishing such descriptions. "Without any disposition to find fault with others, I believe I may be permitted some remarks, since for the last twenty Tl PREFACE. years I have been occupied with the study of Diptera, and have been obliged to spend many hours in identifying (how often fruit- lessly!) the published descriptions. What renders the identi- fication of a very great number of the existing descriptions so very difficult, is the inexactness of the system used. For however natural the axiom may appear, that a new species is only to be located in the genus to which it really belongs, it is so little re- spected by most dipterological writers, that a long list could be made out of the instances in which they have sinned against it; in- deed the number of cases, in which a new species has been placed in a wrong family, is not small. It is not even always sufficient to place it in the right genus, for as soon as this genus is at all numerous in species, or the species are difficult to distinguish, the peculiar group of the genns to which the new species belongs should be pointed out, and if among the species already well known there are any very similar to those described, they ought of necessity to be specially mentioned. Consequently only those entomologists will publish new Diptera with success, who are completely acquainted with the system of this order of insects, whereas he who has a defective knowledge of it, far from advancing science, lays impediments in its way. The first task, then, for those who intend to come forth with satisfactory papers on the field of Dipterology, will of course be to acquire a most complete and sure knowledge of the system. As an introduction to the following essays of a more mono- graphic character, will be found a short sketch of the terminology of Diptera, as well as one of the dipterological system. The latter afforded me an opportunity of giving an outline of the North Ame- rican dipterological fauna, as far as known to me at present. An elaborate classification, equally detailed in all its parts, would require not only a larger amount of materials than I have at my disposal, but also, in order to be intelligible, a considerable num- ber of plates. I am compelled, therefore, to give up such an un- dertaking for the present, I hope, however, to be able to execute it at some future time. Although I trust that my short sketch will prove of some help to the student, by furnishing him occasion- ally a useful hint, or guiding him aright in general, it will be readily understood that in the prosecution of the study he will require more detailed information. I will, therefore, briefly indicate the works in which he may find it : Meigen's Zweiflugelige Insecten (7 vols. PREFACE. Vll 8vo.) is still the best work, exhibiting the dipterological system, lu order to obtain information on the progress which science has made since Meigen's age, this work may be followed by the study of Walker's Diptera Britannica. The excellent plates by Mr. Westvvood, and the systematic arrangement prepared for the most part by Mr. Haliday, give to this work a value not shared by Mr. Walker's other publications. Xext to these I would name Mr. Macquart's Dipteres Exotiques, a work which, notwithstanding the errors in many of its figures and the carelessness of nearly all the descriptions, affords a great deal of useful information about the systematic arrangement of Diptera. After having acquired a general knowledge of the system by the study of these three works (or, if not familiar with the German language, of the two latter only), the best plan will be to con- centrate special study on one, or, at the utmost, on a few families of Diptera, and to consult the monographic papers relative to them, which are not difficult to procure, in order to obtain a com- plete and sure knowledge of characters within a more limited field. For even the smallest field will always be found wide enough to afford opportunities for the most interesting discoveries. This mode of obtaining a knowledge of the system capable of serving as a solid foundation to valuable publications is certainly a long and tedious one. It may be considerably facilitated, however, by the use of a well determined collection of typical specimens of all the families and genera, and it will afford me much pleasure to ex- tend all assistance in my power to those who may prefer the latter course ; for both my wish to become better acquainted with the Diptera of North America and their desire to study the system might well coincide to supply the wants of both parties. I am always ready to send in exchange for well preserved N. A. Diptera forwarded to me (address Mr. H. Loew, Meseritz, Prussia) a reasonable equivalent in accurately named representatives of the genera. I should probably be obliged, in most cases, to send only European specimens, whereas, perhaps, it would seem more de- sirable to have N. A. species. But the number of species occur- ring in perfect identity both in Europe and Xorth America is so surprisingly large, and, besides, there are so many N. A. spe- cies exceedingly resembling well-known European ones, that the best foundation for the study of N. A. Diptera would be a com- plete collection of European species. It will be very useful to Vlll PREFACE. both parties, if those who desire an exchange would remember that the smaller and smallest species possess the greatest interest for me, and if they would at the same time point out to me such families as they are chiefly desirous of knowing. Moreover, it will be necessary to agree about the way in which the equivalents might be forwarded. In case I receive no such direction, I shall suppose I am at liberty to send them by the kind intervention of the Smithsonian Institution at Washington, through which I beg all consignments intended for me may be forwarded. I have no doubt as to a successful issue to my labors, if I con- tinue to receive the same liberal and generous assistance from the Smithsonian Institution and from Baron Osten-Sacken which I have enjoyed from the beginning, especially if this commencement contributes to increase the number of those interested in the study, and ready to promote it by the cornmnnication of species taken by them, in the same way in which Messrs. Rob. Kennicott, S. H. Scudder, A. S. Packard, Edw. Norton, and others, have furnished materials for the excellent paper of Baron Osten-Sacken on the Lirnnolidse of North America. II. LOEW. MESERITZ, 3 Oct. I860. TABLE OF CONTEXTS. PAGE Advertisement ... . iv Preface ........ v Table of Contents ....... ix Ou the Terminology of Diptera . . . xi SKETCH OF THE SYSTEMATIC ARRANGEMENT OF DIPTEKA . 1 I. Neiuocera ...... 5 II. Brachycera ..... 15 III. Coriacea .... 48 Ox THE NORTH AMERICAN TRYPETIDAE 49 1. Extent of the family .... 49 2. Its division into Trypetina and Dacina . 51 3. Its natural character . . . . .52 4. Its relationship .... .56 5. North American species hitherto recorded . 57 6. Systematic arrangement of the species described, with their synopsis . . . . .61 7. Description of the species . 64 Appendix I .... 91 Appendix II ... 94 Appendix III . . . . . .99 ON THE NORTH AMERICAN SCIOMYZIDAE . ] 03 ON THE NORTH AMERICAN EPHYDRINIDAE . . . 129 Notiphiliua ... . . 131 Hydrellina . ... 149 Ephydrina . . . - . 158 X CONTENTS. PAGE ON THE NORTH AMERICAN CECIDOMYIDAE. By R. Osten Sacken . 173 On the classification of the family . . .173 On its habits ...... 179 On the North American Ceciclomyiae at present known 186 Index ...... .207 Additions and Corrections . . 220 ON THE TERMINOLOGY OF DIPTERA. DIPTERA have so much in common with other orders of insects that the terms applied to the latter, which I may consider as gen- erally known, may frequently be used for the former. I have there- fore merely to explain those terms which, on account of the peculiar organization of Diptera, are either applied solely to the insects of this order or are used in a more or less modified sense. It is well known, how little the various authors agree in the choice of these terms, and how many of them seem to find pleasure in departing as much as possible from the terms used by their predecessors. This is a great evil, aggravating the difficulty of understanding Dipterological publications, and impeding the progress of Dipter- ology. It would take too much space to explain all the terms used by different authors, and I confine myself to those only which seem the most necessary and which I have used in this publication. The following considerations have guided me in their choice. I think it a duty of a later author to accommodate himself to the usage of his predecessors, especially those who have written stand- ard works, and at the same time to reconcile them as much as possible where they differ from each other. The indispensable innovations should be introduced only gradually and in conformity with the established usage, since in such matters an agreement about the terms chosen is more important than the mode of selecting them. Meigen, Wiedemann, and Fallen in earlier times, Zetter- stedt and Macquart more recently, have in that respect a claim to our attention. It has therefore been my object to assume the position of an arbiter between them, and to avoid such terms as depart entirely from the adopted usagfe, except in cases in which I might differ so much from my predecessors as to become unintel- ligible if restrained by their terminology. The head has a hinder plane opposite to the thorax, called occi- Xll ON THE TERMINOLOGY OF DIPTERA. put (occiput)] that region of it lying over the junction of the head is the nape (cervix). The part of the head which reaches from the antennas as far as the occiput and is limited laterally by the compound eyes, is the front (frons), the upper part of which is the crown (vertex), the limit between the front and the occiput having the name of vertical margin (maryo verticalis). The middle of the front being often of a more membranaceous substance and sometimes differing io color from its borders, is called the frontal stripe (vitta frontalis). On the crown, there are the simple eyes (ocelli), being usually three in number and forming a triangle, sometimes on a sharply defined triangular space, the ocellar trian- gle (triangulum ocellare). Most of those Diptera which undergo their metamorphosis within the larva-skin possess, immediately above the antennae, an arcuated impressed line, which seems to separate from the front a small piece usually of the form of a cres- cent, the frontal crescent (lumtla frontalis). The impressed line itself, which continues over the face nearly as far as the border of the mouth, is called the frontal fissure (fissura frontalis). It owes its origin to a large bladder-like expansion which exists at this place in immature imagos, and which helps them in bursting the pupa case. The frontal fissure of course is the true anterior limit of the front, and the frontal crescent in fact belongs to the face; however, on account of its usual situation, it is commonly considered as a part of the front. In many genera the eyes of the males meet on the front, so as to divide it into two triangles, the superior of which is called the vertical triangle (triangulum verticale), the inferior the anterior frontal triangle (triangulum frontale anterius), or simply the frontal triangle (triangulum frontale). The anterior portion of the head reaching from the antennae to the border of the mouth or oral margin (peristomium) is called the face (fades). In most Diptera it is divided into three parts adjoining each other, the limits of which depend on the situation which the frontal fissure, continued to the oral margin occupies in the developed imago ; the form and mutual proportion in size of these parts are of the highest value in the classification and distinction of the species of Diptera. Beneath the antennas there are in many Diptera longi- tudinal holes for their reception, the antennal furrows (fovece an- tennales) ; the antennae lie in them while the insect is still in the pupa case, sometimes even after its exclusion. That part of the head which lies on the side beneath the eyes is the cheek (gena). ON THE TERMINOLOGY OF DIPTERA. Xlll The compound eyes are sometimes encompassed in a larger or smaller part of their circumference by a ring, somewhat swollen, and separated more or less distinctly from the remainder of the surface of the head ; it is called the orbit (orbita), the successive parts of which may be called the anterior (orbita anterior sivefa- cialis), inferior (inferior s. genalis), posterior (posterior s. occipi- talis), superior (superior s. verticals), and frontal (frontalis) orbits. An orbit is also often spoken of, where no ring is distinctly set off from the rest of the surface of the head ; in this case a distinct color or some peculiar structure mark the nearest surroundings of the eyes. The oral parts of Diptera, destined for sucking, are called the sucker or proboscis (proboscis). They are either inserted at the end of a more or less cylindrical prolongation of the head, called the snout (rostrum), or project from a wide aperture often occupy- ing a great part of the under surface of the head, called the mouth hole (cavitas oris). The common, fleshy root of the oral parts is connected by a membrane with the border of the mouth. This membrane often has a fold, sometimes of a quite horny substance, and is then called the clypeus (clypeus s. prcelabrum) ; it is either entirely concealed by the anterior border of the mouth and is then usually movable, or it projects over it as a ridge and is then usually immovable. The largest of the oral parts in most Diptera is the fleshy under lip (labium), consisting of the stem (stipes) and the knob (capitulum labii) formed by the two suctorial flaps (labella). Besides the under lip, the palpi (palpi) are most perceptible and must be noticed in the description of the species. The remaining oral parts are generally rather small and stunted, having the form of bristles or horny lancets; they are considered as being the tongue (lingua), under jaws (maxillce), upper jaws (mandibulae) , and up- per lip (labrum), the latter shutting the under lip from above. These parts are not easily applicable in distinguishing species. The thorax of Diptera as well as that of other insects consists of three segments, the prothorax, the mesothorax, and the meta- thorax. But in the order of Diptera the development of the meso- thorax exceeds so much that of the two other portions, that it forms by far the largest part of the whole thorax, and in the description of Diptera is exclusively designated with the latter name, while other names are given to the prothorax and metathorax, when some particular part of them is to be characterized. The protho- XIV ON THE TERMINOLOGY OF DIPTERA. rax being generally very little developed, sometimes forms a neck- like prolongation which bears the head, and is then called the neck (collum). Sometimes the fore corners of the mesothorax or the shoulders (humeri) are covered by a lobe of the prothorax (lobus prothoracis humeralis), distinctly separated from the mesothorax; but it is not unusual for this lobe to be so soldered to the meso- thorax that it is not possible to discover a distinct limit between them, except in general, by their color or hairs; it is then called the shoulder callosity (callus humeralis). The prothorax sometimes also applies closely to the anterior border of the mesothorax, and has then the name of collar (collare). The mesothorax frequently has a transverse furrow (sutura transversalis) crossing the middle of its upper side and ending on each side a little before the base of the wing; its presence or absence as well as its form furnishes characters important in the classification of Diptera. On each side of the breast — the breast side (pleura) — there is beneath the shoulder a spiracle (stigma prothoracis) still belonging to the pro- thorax. To the back of the mesothorax applies the scutcheon (scutellum), separated from it by a furrow. Beneath the scutellum a part of the metathorax is to be seen, called metanotum, generally descending obliquely, often very convex, and on each side with a more or less inflated space, called the lateral callosity of the meta- notum (callus metanoti lateralis). The poisers (halteres) have their origin beneath this callosity, and before either of them we see the spiracle of the metathorax (stigma metathoracis). The mem- branous covers which in many Diptera are found above this spi- racle, have the name of covering scales (tegulte).* The abdomen is the third of the three principal parts of the body, but we usually so call its upper side only, the name of belly (venter) being given to the under side. The segments of the ab- domen are of course, as in the other orders of insects, counted from the front to the back; but the anterior ones are often soldered together, while the posterior ones are stunted, and by their con- cealed situation withdrawn from the eye; much caution is therefore required in counting them. The statements about their number are frequently rather arbitrary and conventional, and often require an explanation. At the end of the abdomen we see in the male the appendages destined to take hold of the female in the copula * Some authors call them squamae. — 0. S. ON THE TERMINOLOGY OF DIPTERA. XV (hypopygiwm), in the female the organ for laying the eggs (ovi- positor) • the former, if they have the form of pincers and are not bent under the belly, are called the pincers (forceps), the latter according to its form either the borer (terebra) or the style (stylus). Both organs are of the greatest importance in the distinction of species in many families, and their structure being generally very complicated and varying much in different families, deserves a most attentive study. The neuration of the wings of Diptera forms so essential a foun- dation of their systematical arrangement and is so useful for the distinction of species, that its thorough knowledge and a scrupulous and accurate denomination of its single parts and of their mutual arrangement is quite indispensable. Our first and most important task will be to ascertain which parts of the neuration of the wings correspond to each other in the different families, since this is the only way to obtain a terminology in which corresponding things are designated by the same names, and which, therefore, is not liable to misinterpretation. At a first and superficial glance, the neuration of the wings shows so different a structure in the various families of Diptera, that it seems impossible to reduce it to a common type. But, on a closer examina- tion, we find that we can make out without much difficulty a common type existing in its greatest simplicity and plainness in the Muscidse. The framework of the whole neuration of the wings is formed by the longitudinal veins (vents longitudinales), which are connected with each other by the transverse veins (vence transverse s. venulce). The longitudinal veins spring from four trunks, issuing from the base of the wings; the first and fourth trunks being the least de- veloped, the second and third must be taken for the main trunks, and consequently the longitudinal veins originating from them, for the main longitudinal veins of the wing. To the anterior of these two main trunks belong three longitudinal veins, the foremost of which runs first parallel to the anterior border of the wing and joins it at a greater or less distance from the tip of the wing ; it is called the first longitudinal vein (vena longitudinales prima). The second longitudinal vein proceeds from the first generally be- fore the middle of the wing, and reaches the anterior border of the wing nearer to the tip. In a similar way the third longitudinal vein has, again, its origin from the second. To the second main trunk also belong three longitudinal veins, which are counted XVI ON THE TERMINOLOGY OF DIPTERA. from the front to the back, and are called the fourth, fifth, and sixth longitudinal veins. The hindmost vein of the anterior main trunk and the foremost vein of the posterior main trunk, i. e. the third and fourth longitudinal veins, are connected by a transverse vein situated about the middle of the wing and called the small or mid- dle transverse vein (vena transversa minor s. media). Using this transverse vein as a starting-point, we cannot easily have any doubt about the position of each of these six main longitudinal veins. The remaining neuration takes place in the following way: The first of the four trunks emits a usually rather stout vein, forming the anterior border of the wing ; it either runs round the whole border of the wing, attenuating a little towards its end, and is called the marginal vein (vena marginalis), or it only reaches as far as the fourth or third, sometimes even the second or first longitudinal veins, and is then generally called the costal vein (vena costalis s. costa) ; both these expressions can be used as identical without any fear of incorrectness. The costal vein is sometimes interrupted in one or more places, thus forming a number of successive portions, a structure most characteristic in several families. Besides this vein, a second one proceeds from the foremost of the four trunks, which, from its being frequently absent, is not counted with the other longitudinal veins, but is called the auxiliary vein (vena aux- iliaris); not far from its base, it is connected by the transverse shoulder vein (vena transversa humeralis) with the costal vein. A total or partial absence of the auxiliary vein, its structure, and the peculiarities of situation which it has relatively to the first longi- tudinal vein, are very characteristic marks for the distinction of families and genera. The first and second longitudinal veins are usually simple, the third being frequently forked ; this furcation arises from its emitting beyond the small transverse vein an anterior branch, which gene- rally runs to the border of the wing between the second and third longitudinal veins, and is therefore called the anterior branch of the third longitudinal vein. Both branches together form the fork of the third longitudinal vein, and that part of this vein which lies between the small transverse vein and the point of forking of both branches is called the handle (pedunculus) of this fork. Some- times the foremost branch of the third longitudinal vein is con- nected with the second longitudinal vein by a transverse vein, or it runs into the second longitudinal vein instead of running to the ON THE TERMINOLOGY OF DIPTERA. XV 11 border of the wing, and thus has the appearance of a transverse vein. In most Diptera there is no other connection between the third and fourth longitudinal veins except the small transverse vein, and we can cross the wing between the third and fourth longitudinal veins in its whole length without meeting another vein but the small transverse vein. But in some families the fourth longitudinal vein, abandoning towards its end its former direction, turns to the third longitudinal vein and reaches it either at its end or a little before it, constituting thus a1 'second connection: there is a rarer case when that connection is effected by a transverse vein placed distinctly on the fourth longitudinal vein. A third connection between the fourth longitudinal vein and the anterior main trunk is formed in some families in the neighborhood of the base of the wing; often there is only a transverse fold running obliquely from the fourth longitudinal vein to the first; in some families it incras- sates into a transverse vein. The three longitudinal veins belonging to the second main trunk usually begin to diverge quite near the base of the wing; the hind- most of them, i. e. the sixth longitudinal vein, is often distinctly seen as the continuation of the common trunk, while the two ante- rior ones uniting with their bases seem to form a kind of loop which touches the main trunk only at one point. As frequently, the fifth longitudinal vein represents a distinct continuation of the main trunk; in some families all the three longitudinal veins appear in equal distinctness as its branches. Between the fourth and the fifth longitudinal veins there are in general two transverse veins, which divide the space of the wing, included by the fourth and fifth longitudinal veins, into three parts. The first of these transverse veins is the anterior basal transverse vein (vena transversa basalts anterior s. venula basalts anterior), the absence of which is charac- teristic for some families; the second is usually the longest trans- verse vein of the wing and is of the highest systematic value ; it is called the posterior transverse vein (vena transversa posterior s. venula posterior). Not unfrequently another vein starts from its middle, running to the border of the wing; it cannot be considered as a longitudinal vein, and is called the anterior intercalary vein (vena intercalaris anterior). It must not be confounded with a branch emitted in some Diptera from the posterior side of the fourth longitudinal vein before its tip. B xviii ON THE TERMINOLOGY OF DIPTERA. Near the base of the fifth longitudinal vein rises the posterior basal transverse vein (vena transversa basalis posterior s. venula basalis posterior), usually a short transverse vein, running to the sixth longitudinal vein, but frequently meeting it only in a later part of its course at a very acute angle, or even reaching the bor- der of the wing, without having met that vein; in all these cases it divides all the space of the wing lying between the fifth and sixth longitudinal veins into two parts. In several families there is, immediately beyond this transverse vein, another vein, the posterior intercalary vein (vena inter calaris posterior), which proceeds from the fifth longitudinal vein and runs to the border of the wing ; sometimes it meets the fifth longitudinal vein before reaching the border. In that part of the wing which is behind the sixth longitudinal vein, are spread the branches of the hindmost of the four trunks ; it is entirely wanting in many Diptera, and exists in most of them only in a rudimentary state; therefore generally only one branch, or at the utmost two weak ones, not reaching the border of the wing, are perceptible; they are called the axillary veins (vena ax- illares). Where the hindmost trunk is well developed, these veins become complete longitudinal veins; they best preserve the same name, but may be numbered as the seventh, and, where two exist, as the seventh and eighth longitudinal veins without any fear of misinterpretation. In the case of such a great development of this trunk, the foremost of the veins belonging to it is generally connected near its base with the sixth longitudinal vein by a trans- verse vein. It results, from the foregoing exposition, that the anterior part of the wing is divided by the three longitudinal veins belonging to the anterior main trunk, and the posterior by the three longitudi- nal veins belonging to the posterior main trunk, each into three sections, an exterior one, a middle, and an interior one, while the three sections of the anterior part of the wing are separated from those of the posterior part by a middle stripe or baud which ex- tends from the base of the wing to its tip. It would be an easy task to invent fit names for these principal parts of the surface of the wing, from which convenient expressions would result for their single parts or the cells of the wings. But it seems to me that the introduction of such a new nomenclature would hardly promote our principal end, the agreement of authors in the use of termino- ON THE TERMINOLOGY OF DIPTERA. XIX logical expressions, since it is not so much the nature of the re- ceived terms as the consent in their application which we must aim at. I therefore think it most advisable to retain such names for the denomination of the wing-cells, as are already in general use. But in adopting these names I cannot but mention that many of them do not seem to be well chosen, and that I accept them only with the intention of bringing about a terminology generally agreed upon. I shall, therefore, call the cells belonging to the first section of the wing the costal cells (celhdse costales}, those of the second the marginal cells (celhdse marginales), and those of the third the sub- marginal cells (cellules submarginales). The latter are of the greatest importance for characterizing families and genera, as well as for the distinction of species. When the second and third longitudi- nal veins are simple, and the third anterior section is consequently undivided, there exists only one submarginal cell; but when the third longitudinal vein has a branch running to the border of the wing, we count two such cells, an anterior and a posterior one ; when the anterior branch of the third longitudinal vein is also connected with the second longitudinal vein by a transverse vein, the number of submarginal cells amounts to three, among which that, formed by the inner part of the anterior submarginal cell, is called the interior submarginal cell ; when the anterior branch of the third longitudinal cell assumes the form of a transverse vein running to the second longitudinal vein, only an interior and an exterior submarginal cell are distinguished. Among the existing names, none is. well applicable as a common denomination either to the cells belonging to the middle of the wing or to those of each of the two first sections of the posterior part of the wing ; I am compelled, therefore, though not without reluctance, to give up the application of such names. Among the cells of the portion just mentioned, there are three that have gene- rally been too little noticed in the description of the neuration of the wing. Their different forms give very good characters, the more so as, on the whole, the differences, which the neuration shows in the neighborhood of the base and costal border, have always a higher systematic value than those occurring near the tip or the posterior border of the wing. Those three cells are placed nearest to the base of the wing ; the first of them belongs to the middle of the wing, and reaches as far as the small transverse XX ON THE TERMINOLOGY OF DIPTERA. vein ; the second belongs to the first section of the posterior part of the wing, and extends as far as the anterior basal transverse vein ; the third belongs to the second section of the same part of the wing, and joins the posterior basal transverse vein. These three cells may, in general, be called the three basal cells (ceUulae basales*). The foremost of them is generally much longer than the two others, a proportion which is usually indicated by the expression of "one large and two small basal cells;" against this mode of ex- pression nothing can be objected, since it implies no uncertainty. It is, however, a little puzzling to invent a convenient term, when the posterior basal transverse vein, instead of running to the sixth longitudinal vein, assumes the character of a longitudinal vein, and runs to the border of the wing, so that the hindmost basal cell joins the border of the wing. Not only in this case, but also when the hindmost basal cell, though closed, is distinguished from the second basal cell by a much more considerable length, it is usually named the anal cell (cellula anah's), and then, consequently, two basal cells are considered to be present. Badly chosen as the term "anal cell" may be, it is, nevertheless, so settled that it will be difficult to remove it by the introduction of a more convenient one. In certain families the great and very symmetrical development of the three basal cells is characteristic; they are then called the ternated cells (cellulse ternatse), which term, though expressive of the thing, seems to be superfluous. One of the most important cells is that belonging to the first section of the posterior part of the wing, and extending from the anterior basal transverse vein to the posterior transverse vein, and bearing the little transverse vein on'its anterior margin ; it is generally called the discoidal or discal cell (cellnla discoidalis). When the anterior basal transverse vein is wanting, which is characteristic in many families and genera, this cell coalesces with the second basal cell, which then must be considered as a part of the discoidal cell ; if the posterior trans- verse vein has disappeared, there is no discoidal cell at all. In those Diptera which possess the anterior intercalary vein, sometimes the part of the posterior transverse vein situated before or behind this intercalary vein is wanting, and in that case the existence of a discal cell is granted, which, in the former instance, is considered as anteriorly opened, in the latter, as posteriorly opened. The second cell of the middle of the wing opening in its border, and those of the two first sections of the posterior part of the ON THE TERMINOLOGY OF DTPTERA. wing, are called the cells of the posterior margin, or posterior cells (cellulx posteriores), and numbered as first, second, etc., beginning with that which belongs to the middle of the wing, and is limited at its base by the small transverse vein. It is evident that in all Piptera there are really only three posterior cells. They exist in their typical simple form in the Muscidse. The first of them be- longs to the middle of the wing, the two others to the first and second section of the posterior part of the wing. The first is usually subject to no partition, but is sometimes closed before reaching the border. The second is frequently divided in two portions by the presence of the anterior intercalary vein, and this happens whenever the fourth longitudinal vein emits a hind branch before its end ; it even forms three portions when this branch exists along with the intercalary vein. In the genera having a posterior intercalary vein, a bipartition of the third posterior cell occurs. Though it would be very convenient to speak in all cases of only three such cells, and to point out in the way indicated the mode of their further partition, yet the ruling usage does not admit of this, but counts all these portions as successive posterior cells, whence their number sometimes amounts to six. When the second posterior cell and the discoidal cell are united in conse- quence of the absence of the posterior transverse vein, the cell formed in this way retains the name of second posterior cell. The cells belonging to the third section of the posterior part of the wing. are not, usually, completely separated from each other, and then are frequently termed the false cells (cettulse spurias); a better term for them might be that of axillary cells (cellulse axil- lares). They are numbered in the direction from the sixth longi- tudinal vein towards the posterior angle of the wing. As for the expressions costal border, tip, posterior border, pos- terior or anal angle of the wing, they are understood by everybody. The posterior angle is terminated by the axillary incision (incisura axillaris) towards the base of the wing. The wings of many Dip- tera are provided with a lobiform appendage, the alar appendage (alxla), reaching from the axillary incision to the innermost base ; it must not be confounded with the covering scale that lies above the poisers, and which has often been called by the same name. In order to understand a very intricate neuration and reduce it to the simple tjpe, we must take care not to assume for parts of the same main vein all those ramifications which run in one direc- XX11 ON THE TERMINOLOGY OF DIPTERA. tion. As threads loosely drawn up in a frame, when strongly strained by transverse threads of different length, must necessarily adopt an angular direction, so do longitudinal veins, in consequence of a varied situation and the length of the transverse veins. The outline of the wing, the length of the longitudinal veins, the situ- ation and length of the transverse veins, as well as the area of the two membranes of the wing, stand in such a relation to each other that the wonderful effect of their hardening after the exclusion of the insect will be a surface more or less even, but in every case fit for the performance of flight, the main agents of which are appa- rently the anterior part of the wing, as being more stout and rigid, and its posterior part, which, being in most cases united with the former by the small transverse vein only, is more movable, and acts as an inclined plane, propelling the insect during the motions of the wings both up and down. It is in such genera only as Syritta, Bombylius, Nemestrina, the habits of which require not so much a rapid locomotion, as a constant hovering over a spot, that a multiplied connection of the anterior and posterior part of the wing by transverse veins restrains the mobility of the posterior part of the wing, and renders its propelling effect less sensible. A correct understanding of a very intricate neuration is in many cases by far not so difficult as that of a very incomplete one. The latter will be best obtained by observing, that in such wings the three anterior trunks of the veins are not only incompletely de- veloped, but also crowded together on the anterior part of the wing, an uncommonly large space being allotted to the fourth trunk. In this way, the striking narrowness of the anterior and middle parts of the wing and the extraordinary dilatation of the posterior part, find their explanation. Sometimes a closer exami- nation of the surface of the wing will yield a useful result by our observing the different kind of hair peculiar to the veins, and which remains, though the veins themselves are wanting. All Diptera with very incomplete neuration are bad fliers, since the greater flexibility of the posterior part of their wings can but imperfectly compen- sate the propelling effect of this part when sustained by a stronger neuration. The legs of diptera, like those of the other orders, consist of four principal parts, called the hips (coxse), thighs (femora), shanks (tibiae), and feet (tarsi). The hips consist. of two joints; the second, smaller one, is called trochanter. The feet are gene- ON THE TERMINOLOGY OF DIPTERA. XX111 rally five-jointed ; the first joint is called metatarsus. At the tip of the last joint there are two claws (ungues), and under each of them there is generally a membranaceous appendage called pul- villus. Besides these appendages, many families have between them a third single appendage of similar structure, which is called empodium; in other families this organ is bristle-like, or altogether wanting. I have little to say about the expressions for the different cha- racters of the surface and the clothing of the parts of the body of Diptera; I will observe only that the gradations hoary (pruinosus), dusted (pollinosus), mealy (farinosus), or pubescent (pubesccns~), hairy (pilosus'), bristly (setosus), etc., in their application must be judged more according to a relative than an absolute scale, viz., in a family that has coarse hair the same is called hairy, which in another with fine hair is termed bristly, and so in similar cases. If we were not willing to do so, expressions would fail to point out the existing differences. XXIV ON THE TERMINOLOGY OP DIPTERA. 1. Wing of Ortalis. a. Transverse shoulder-vein (vena transversa humeralis). b. Auxiliary vein (vena auxiliaris). c. d, e,f, ff, and h. First, second, third, fourth, fifth, and sixth longitudinal veins (vena longitudinales prima, secuuda, tertia, quarta, quiuta, et sexta). i. Small or middle transverse vein (vena transversa minor s. media). fc. Hinder transverse vein (vena transversa posterior). I, m, n, o. Costal vein (vena costalis). p. Anterior basal transverse vein (vena transversa basalis anterior). q. Posterior basal transverse vein (vena transversa basalis posterior). r. Rudiment of the fourth trunk. S. Axillary incision (incisura axillaris). A, B, and G. First, second, and third costal-cells (cellulae costales prima, secunda, et tertia). D. Marginal cell (cellula marginalis). E. Submarginal cell (cellula submarginalis). F. G, and)/. First, second, and third posterior cells (cellula; posteriores prima, secunda, et tertia). /. IMscal cell (cellula discoidalis). K. First or large basal cell (cellula basalis prima s. major). L. Second basal cell, or anterior of the small basal cells. Jf, Third basal cell, or posterior of the small basal cells. N. Anal or axillary corner of the wing (angulus analis s. axillaris). 0. Alar appendage (alula). 2. Wing of Emjiix. t. Anterior branch of the third longitudinal vein (venae longitudinalis tertiae ramus anterior). «. Anterior intercalary vein (vena intercalaris anterior). 3. Wing of Dasypogon. t. Anterior branch of the third longitudinal vein. u. Anterior intercalary vein. v. Posterior intercalary vein. DIPTERA OP NORTH AMERICA i. SKETCH OP THE SYSTEMATIC ARRANGEMENT OF DIPTERA : WITH AN ENUMERATION OF THE GENERA HITHERTO RECORDED AS' FOUND IN NORTH AMERICA. OUR knowledge of the Dipterological Fauna of North America has lately made rapid progress by the great attention paid to it by Baron Osten Sacken during his residence in Washington. As a preliminary to further investigations, he prepared, in 1858, for publication by the Smithsonian Institution, a Catalogue of the then described North American Diptera, which had the great and essential merit of nearly entire completeness. It cannot but be considered as a wise precaution that he did not enter upon a criti- cal examination of the published species, as he well understood that such an examination could only be the work of the combined efforts of many persons, and the fruit of a long toil of years, and that consequently undertaking it would have indefinitely retarded the publication of such a catalogue, so desirable for the advance- ment of North American Dipterology. The impulse caused by Baron Osten Sacken's Catalogue is already evident, and it has proved a welcome and valuable assistance 'to every one attempting a more thorough study of North American Diptera, by an intel- ligible arrangement of the already published species, not only sparing him much laborious research, but also giving him the cer- tainty of not overlooking a species already described. But although this Catalogue presents a survey of all papers hitherto published, and of the contributions of each author, it does not, and according 1 2 DIPTERA OF NORTH AMERICA. to its plan could not, afford a survey of the North American Fauna Dipterologica, corresponding to the present state of sys- tematic Dipterology; on the contrary, sketching such a survey is one of the tasks to which it looks forward to as one of the first fruits of its publication. It would be quite impossible to draw such a systematic survey of the hitherto known North American Diptera from the Catalogue itself, since it comprises the publica- tions of the authors of different times and countries, of writers who had the most different systematic ideas and points of view, and since, in consequence of its plan, it could not but include such papers as are devoid of any solid knowledge of systematic Dipter- ology— to which, above all, the descriptions of Rob. Desvoidy, and, in a still higher degree, those of Mr. Walker belong. Conse- quently a survey of those families and genera which North America really possesses, is to be acquired in no other way than from the in- spection and careful investigation of the species themselves. The rich collections of Baron Osten-Sacken have enabled me to examine a number of North American species sufficient to allow me to ven- ture an essay of the kind indicated. In this survey I have adopted for the North American Fauna the same area as that of Baron Osten Sacken's Catalogue, the materials upon which I establish my work corresponding to this area. Still it cannot be denied, as far as I am able to judge, that this area, in its southern extent, reaches beyond the limits of the North American zoological province. In order to give a true, though of course not complete sketch of the North American Dipterological Fauna, I can, besides such species as I know by my own inspection, have regard only to those the systematic location of which is in no way doubtful. For many years past all Diptera have been divided into two large sections, Nemocera and Brachycera. In the Diptera of the first section the antenna, having the fundamental form of a thread, consists of many joints, two of them being called the joints of the scapus, the following those of the flagellum. The latter are all of the same structure, although this structure varies in different spe- cies. The first joint of the flagellum, i. e., the third of the whole antenna, is never so distinguished in size or structure that one might consider the succeeding joints as its accessorial appendages, nor is the connection of the joints (with the exception of the Bibi- onida, Mycetophilidce, and a few others) such as might lead us to DIPTERA OF NORTH AMERICA. 3 consider them as one, divided into several annuliform segments. In the second section, the Brachycera, the two joints of the scapus are likewise separated; the third joint, or first of the flagellum, usually differs by its remarkably developed size and its anatomical structure, causing it to be considered as a sensorial organ about the nature of which entomologists are not yet agreed. The suc- ceeding joints of the flagellum are much reduced in size, generally very few in number, and often of unequal number in nearly related genera, or even in species of the same genus. They even disappear entirely in some genera (e. g., in Scenopinus). If they are extant, they have usually the form of a style or bristle, the position of which, according to its nature, is in fact apical, although, from the development of the under side of the third joint, the bristle often seems inserted on its back, or even, in some instances, in the immediate vicinity of the base itself. In the genera, in which the .first joint of the flagellum is not of a remarkable size, the following joints are generally more numerous, and either all or the greater part of them share the peculiar organization showing their function to be that of a sensorial organ. They are applied at the same time so closely to the first joint of the flagellum, that we are compelled to consider all of them together as one, divided into several segments, or the terminal one as a style or bristle of a single joint, formed by the other joints of the flagellum. Consequently the essential difference between the sections Nemocera et Brachycera is this, that in the latter the number of joints of the flagellum is not only smaller, but also that the lower joint, sometimes a few joints, always the lower ones, rarely all, have a more distinct de- velopment, and at the same time a peculiar anatomical structure undoubtedly proving their function to be that of a sensorial organ. It cannot be denied that those families of Brachycera in which several of the joints of the flagellum are so soldered together as to form one compound and annulated mass, stand nearest to the section of Nemocera, and that amongst these families the Xylopha- gidce must be placed immediately on the limit of both sections. It is more difficult to point out a family of Nemocera, which comes nearer to the section of Brachycera than all the others; in general the families of Rhyphidce and Bibionidce may be considered as those to which this station must be assigned. It is- a fact that some dis- coveries made in modern times have obliterated to a certain degree the sharpness of the limit which was considered to exist between 4 DIPTERA OF NORTH AMERICA. the sections of Nemocera and Brachycera. The fact known long ago, that in some genera of Stratiomydcc and Tabanidce the joints of the flagellum not being closely compressed, do not form a com- pact joint, has been rather neglected in this respect, perhaps be- cause the Stratiomydee and Tabanidce, by their whole organization, are rather remote from the Nemocera, and have so strikingly pecu- liar characters as individual families, that their comparison with the Nemocera has scarcely been thought of. The case was dif- ferent when* a similar structure was observed in the family of Xylo- phagidce. After I had myself first pointed out the genera Electro, and Chrysothemis, discovered by me in Prussian amber, Mr. Hali- day found the still more surprising North American genus Rachi- cerus. I shall have hereafter to mention a second North American genus of Xylophagidce, which has the flagellnm of the antennae not annulated, but really many-jointed. All these facts, however, are not sufficient to compel us at present to give up the separation of the Nemocera and Brachycera. Many authorities have likewise objected to uniting under the head of Brachycera all those families which cannot be referred to the Nemocera, especially and with the fullest reason, to the union of the Hippoloscidce with, the other Brachycera, since both the history of their development arid their internal and ex- ternal anatomy essentially differ from them. They can only be considered as a third section, co-ordinate to the Nemocera and Bracliycera, and having the same systematic value, and may be named Coriacea, or they may be opposed to the other two together as equivalent, and consequently be comprised under the name of Eproloscidea, that of Proboscidea being left to the two other sec- tions. I intend to follow the first of these two arrangements. Of the families which I shall hereafter enumerate as belonging to the Bracliycera, the Plioridce alone have occasioned some doubts about their title to this place, founded, if I judge correctly, on the abnormal structure of their antenna? ; these are considered as one-jointed, with the terminal bristle consisting of several joints. .Judging by the anatomical structure, I for my part am unable to see in the joint, which is pre-eminently developed and forms the main body of the antennae, more than the first joint of the flagel- lum, its peculiarity arising from the soldering together and stunting of the*two joints of the scapus, the covering of which is moreover less horny than in the other families of Brachycera. This differ- CULICIDAE — CHIRONOMIDAE. 5 ence, surprising as it is, does not seem to me to have systematic importance enough to require a separation of the Phoridce from the remaining Brachycera, and the less so as similar deviations, though not nearly of so striking a nature, also occur in other fami- lies. I mention, as an instance, the remarkably stunted second joint of the antennae in the genus Haltericerus Rond. among the Doli- chopidce. I. NEMOCERA. FAM. I. CULICIDAE. Charact. — Ocelli none. Thorax without transverse suture. Costal vein continued round the margin of the wing, fringed with scales ; veins in their last subdivisions more than six in nuniher. This family, rich in species, comprises only a small number of genera. As such, the old well-known genera CULEX, AEDES, ANOPHELES, and CORETHRA are to be named first, being those among which Meigen has distributed the European species. To them may be added the genera MEGARHINA, PSOROPHORA, and SA- BETHES, separated from Culex by Rob. Desvoidy, the two last being scarcely tenable, whereas MegarJiina is acknowledged as holding good. The genus MOCHLONYX, established by me, is very near to Corethra, differing, however, by the abbreviation of the first tarsal joint. Species of the genera CULEX and ANOPHELES occur over all parts of K A., whereas MEGARHINA and PSOROPHORA are only represented by single species belonging, as it seems, more to the South, as is also CORETHRA by C. punctipennis Say. FAM. II. CHIRONOMIDAE. Charact. — Ocelli none. Thorax without transverse suture. Wings with- out vein along the posterior margin ; costal vein ending near the tip of the wing. This family is not much richer in genera than that of the C'ulicidee, but far more so in species. It contains the old and well-established genera CHIRONOMUS, TANYPUS, and CERATOPOGON, to which have been added the genera HYDROB^INUS Fries (= Corynocerus Ruthe), 6 DIPTERA OF NORTH AMERICA. DIAMESA Meig., CORYNONEURA Winn., and CLTJNIO Hal., which is so remarkable by the habitation of its larva. The species of Cera- topogon exhibit a good deal of varied organization. A division into a number of smaller genera, which is indispensable, has been attempted, but not executed in a satisfactory manner, and the genera LABIDOMYIA Steph., CULIOCIDES Latr., PALPOMYIA Meig., SPH^ROMIAS Steph., and PRIONOMYIA Steph. can be only consi- dered at present as sub-genera of Ceratopogon. The genus THA- LASSOMYIA Schin. has been separated from Chironomus. Also the genus MACROPEZA Meig. must be united with the Chironomidce, and Macquart is right in having done so in his " Dipteres exo- tiques." The genus ORPHNEPHILA Hal. (= Thaumaka Ruthe = Chenesia Macq.) differs from all the other Chironomidce by the veins of the wings running without attenuation to, and the costal vein being continued round, the posterior border. If we do not establish a separate family for it, its proper place will be here, but as an anomalous genus. The genera CHIRONOMUS, TANYPUS, and CERATOPOGON are largely represented in N. A. ; the most interesting are the species of Cera- topogon. Heteromyia Say belongs here. Of the remaining genera, I have seen only one N. A. species, which belongs to the genus ORPHNEPHILA, and does not seem to differ from 0. testacea Ruthe. FAM. III. CECIDOMYIDAE. Charact. — Ocelli often none. Thorax without transverse suture. Coxae not elongated, femora not thickened, tibiae without spurs. Wings having only few longitudinal veins. It is rather difficult to define sharply this most interesting family, and consequently to characterize it exactly. It contains a very large number of extremely delicate and elegant minute species, remarkable by long and easily rubbed off hairs on the wings and the other parts of the body. The limits between the families Ceci- domyidce (Gall-gnats) and Mycetophilidte (Fungus-gnats) are not very easily fixed, since Zygoneura Meig. shows a combination of the characters of both; the coxas being far less elongated and the spurs of the tibise far shorter than in any other genus of Myceto- philidce; moreover, the antennae are moniliform with verticillate hairs, as is frequently the case in the Cecidomyidce and never so CECIDOMYIDAE. 7 among Mycetophilidce. But the total habitus of the Zygoneurce being more like that of the former than of the latter, and the tibial spurs being so very short, that in some species they can only be discovered by the closest scrutiny, I think I may be justified if I add them to the Cecidomyidce, though in many respects they agree with the genus Sciara, which has its natural place among the Mycetophilidte. The whole family is divided into two sections. The first of these, the CECIDOMYINA, have on the wings four longitudinal veins, the two last of which often coalesce in the beginning of their course, or are more or less incomplete. They have no ocelli, and the first joint of their tarsi is much shortened. The genera belonging here are : HORMOMYIA Loew, DIPLOSIS Loeiv, CECIDOMYIA Latr., As- PHONDYLIA Loew, DIRHIZA Loew, COLPODIA Winn., EPIDOSIS Loew, ASYNAPTA Loew, LASIOPTERA Meig., and CLINORHYNCHA Loeiv. In the genera of the second section, the ANARETINA, between the second and third of those veins of the wings which the first sec- tion possesses, another longitudinal vein is inserted, being simple only in Campylomyza, while it is furcate in all the other genera ; the first tarsal joint is not shortened, and in all genera, with the single exception of Cecidogona, there are distinct ocelli. To this section belong: CAMPYLOMYZA Meig., CECIDOGONA Loew, ANA- RETE Hal., CATOCHA Hal. (= Macrostyla Winn.), LESTREMIA Macq. (= Diamesa Meig.), and ZYGONEURA Meig. I have omitted here the genera HETEROPEZA Winn. and SPANI- OCERA Winn., not having had an opportunity of examining speci- mens. Heteropeza seems to harmonize in many points with the genera of the first section, but differs very strikingly by the totally different structure of its tarsi. Rondani has established in this family a good number of genera, which are, however, quite un- available, since the observations on which they are founded are too inexact. • Yery little information has thus far been published respecting the Cecidomyidce of N. A. Most of the species sufficiently described belong to the genus Gecidomyia in its restricted sense, as is now in use ; viz : Gee. destructor Say, salicis Fitch, and tritici Kirby ; Gee. grossularice Fitch ought, as it seems, to be referred to the genus ASPHONDYLIA ; some fine species of the genera DIPLOSIS and LASIOPTERA occur there. Out of the second section I have 8 DIPTERA OF NORTH AMERICA. seen species of the genera CAMPYLOMYZA, ZYGONETJRA and LES- TREMIA. Of a new genus belonging to the same section, I have seen only one incomplete individual. FAM. IV. BLEPHAROCERIDAE. Charact. — Ocelli three. Wings very ample, naked (/. e. with hairs only perceptible under a very highly magnifying lens), with cracks caused apparently by folding ; no discoidal cell. Posterior tibiae with stout spurs, anterior tibiae unarmed. The genus BLEPHAROCERA Macq. cannot, except by the utmost constraint, be included in any of the existing families. Its nearest relation is the Ceylanese genus TANYRHINA Loew. I unite these two genera in one small family, the name of which I derive from the older of the two. The Blepharoceridce differ from the Cecido- myidce by the stout terminal spur of their posterior tibias, from the Mycetophilidce by their coxae not being elongated, from the Bibionidce by the want of an empodium and pulvilli, and by the very little development of their prothorax. In the form and tissue of their wings they are most nearly allied to the Simulidce, but by the existence of ocelli, and by the long slender legs, they seem to me to differ from them too much to be reunited with them. The neuration of their wings is rather similar to that of the Cecido- myidce ; but Blepharocera has some more longitudinal veins, and thus its neuration resembles that of Macropeza. Besides the longi- tudinal veins, the wings show some fine cracks, perfectly similar in both, and looking as though produced by the expansion of the wings, which had previously been folded ; this mark is peculiar to them, pointing to some peculiarities in their transformation un- fortunately still unknown ; some certainty about the place due to them in the system may, therefore, be expected from the knowledge of their earlier stages. I know only one N. A. species of BLEPHAROCERA very much resembling that species which is spread over a great part of Eu- rope. Blepharocera americana Walk, neither belongs to this genus, nor even to this family. PSYCHODIDAE — T1PTJLIDAE. FAM. Y. PSYCHODIDAE. Charact. — No ocelli. Body with, long, coarse hairs. Thorax without a transverse suture. Tibiae without spurs. Wings everywhere with long coarse hairs, many longitudinal veins, and only a few trans- verse veins ; no discoidal cells. The genera of Psyclwdida, on account of their neuration, form two sections ; in the first, the PSYCHODINA, there are, between the furcate longitudinal veins common to all genera, two simple longitudinal veins. The genera of this section are : PSYCHODA Latr., PERICOMA Hal., ULOMYIA Hal. (= Saccopteryx Hal. ol.), POSTHON Loew, and NYGMATODES Loew (= Nemapalpus Macq.). In the second section, the PHLEBOTOMINA, we see only one longi- tudinal vein between the two furcate veins. The genera belonging here are: PHLEBOTOMUS Rond. (= Hoemasson Loew), TRICHO- MYIA Hal. (= Phalcenomyia Loew), SYCORAX Hal., DIPLONEMA Loew, and PHIUEMATUS Loew. • The small number of N. A. Psychodidce I have seen, belong without exception to the genera PSYCHODA and PERICOMA. In Europe there have been discovered besides the genera Ulomyia, Phlebotomus, Trichomyia, and Sycorax. From the smalluess and fragility of Psychodidce, it may easily be conceived why so few N. A. species have still been detected. FAM. VI. TIPULIDAE. Charact. — No ocelli.* Thorax with a V-shaped transverse suture. Legs very elongated ; the basal cells of the wings reaching beyond the middle ; discal cell existing in most of the genera. The want of ocelli, the considerable length of the legs as well as of the basal half of the wings (the latter cause producing a prolongation of the basal cells beyond the middle of the wing), are the most essential characters of this family. Moreover, the Y-shaped transverse suture of the thorax is of the greatest value, since only the anomalous genera DIXA Meig. and CHIONEA Dolman are destitute of it. It does not seem natural to attribute to the former genus, on account of this circumstance, any other place * Except in Trichocera, where they exist. 0. S. 10 DIPTERA OF NORTH AMERICA. but among the Tipulidce. The abnormal structure of the thorax in the wingless genus Chionea is less surprising, its form depending chiefly on the situation and size of the alary muscles, and, however it may differ in some respects from all the other genera of Tipu- lidce, it would be still more inconveniently located in any other family. Whether the genus POLYMERA Wied., which is distin- guished by its moniliform antennae and the basal cells not reaching to the middle of the wings, may be placed among the Tipulidce, appears doubtful. I should have thought it belonged to the tribe PsycJwdidce, had not Macquart figured the thorax of Polymera fusca with a distinct Y-shaped transverse suture. The variety of forms prevailing in the family of Tipulidce has caused the foundation of a rather large number of genera. In order to facilitate the description of the new species and the iden- tification of the described ones, a considerable increase of the number of genera is indispensable. On the whole, the Tipulidce may be divided into two sections, the TIPULINA having long, and the LIMNOBINA having short palpi. This division, indeed, is no natural one, since some genera with ^ long palpi agree in all the rest of their organization more with the genera of the second than of the first section. The genera of Tipulidce hitherto established, as I know them by sight, or, in part, by the definitions of their authors, may be dis- tributed as follows. To the Tipulinn belong: TIPULA Linn., PRIONOCERA Loeiv, PACHYRHINA Macq., NBPHROTOMA JTe^., CTE- NOPHORA Meig. , DOLICHOPEZA Meig. , OzoDiCERATI/acy., CTENOGYNA Macq., GYNOPLISTIA Westw., PTILOGYNA Westw., MEGISTOCERA Wied., APEILESIS Macq., PTYCHOPTERA Meig.; MACROCHILE Loew, PROTOPLASA 0. S., PTEROCOSMUS Walk., HESPERINUS Walk., BIT- TACOMORPHA Westw., and RHAMPHIDIA Meig. To the section of Limnobina must be referred : LIMNOBIA Meig., GLOCHINA Meig., RHIPIDIA Meig., GERANOMYIA Hal. (= Aporosa Macq.), DICRA- NOMYIA Steph., ANTOCHA 0. S., ELEPHANTOMYIA 0. S., LIMNOBIO- RHYNCHUS Westw. (= Toxorhina Loew), DICRANOPTYCHA 0. S., TEUCHOLABIS 0. S. , ERIOPTERA Meig. , SYMPLECTA Meig. , CRYPTO- LABIS 0. S., GONOMYIA J/e#., GNOPHOMYIA 0. S., CLADURA 0. S., TRICHOCERA Meig., CYLINDROTOMA Meig., ANISOMERA Meig., AR- RHENICA 0. S., ERIOCERA Macq., DICRANOTA Zett., ULA Hal., AMALOPIS Hal., TRICYPHONA Zett., EVANIOPTERA Guer., PEDICIA Lair., LlMNOPHILA Macq., EPIPHRAGMA 0. S., DlCRANOPHRAGMA TTPULIDAE. 1 1 0. S., IDIOPTERA Macq., LASIOMASTIX 0. S., DACTYLOLABIS 0. S., PRIONOLABIS 0. S. ; as Limnobina may also be mentioned the fossil genera which have been found in Prussian amber : TRICHONEURA, CALOBAMON, HAPLONEURA, CRITONEURA, TANYMERA, TANYSPHYRA, STYRINGOMYIA, ATARACTA, and ALLARITHMIA. As genera of doubtful location we may add the genus POLYMERA Wied., and the anomalous genera DIXA Meig. and CHIONEA Dalm. We know as genera of Tipulina occurring in N. A. the follow- ing : TIPULA, PACHYRHINA, CTENOPHORA, PTILOGYNA, PTYCHOP- TERA, PROTOPLASA, HESPERINUS, BITTACOMORPHA, and RHAMPHI- DIA. About the genera and species of Limnobina indigenous in N. A., Baron Osten Sacken, in the Proceedings of the Academy of Natural Sciences of Philadelphia, 1859, has published a detailed and valuable paper, which enters into a more complete and tho- rough exposition of the system of this section than is possible for me to give here. I must, therefore, refer to this paper. The species enumerated in it, most of which are new, belong to the following genera: Limnobia, Rhipidia, Geranomyia, Dicrano- myia, Antocha, Elephantomyia, Lirnnobiorhynchus, Dicranoptycha, Teucholabis, Erioptera, Symplecta, Cryptolabis, Gonomyia, Gno- phomyia, CJadura, Trichocera, Anisomera, Arrhenica, Eriocera, Dicranota, Via, Amalopis, Pedicia, Limnophila, Epipliragma, Dicranophragma, Idioptera, Lasiomastix, Dactylolabis, and Prio- nolabis; also Dixa and Cltionea are recorded as N. A. genera. Note. — The special attention which I have, for several years, paid to the family of Tipulidce may serve as an excuse for my expressing here my views on its distribution. Although these views are founded merely on the study of the species of this continent, the new character which I introduce for the definition of the two principal sub-families may also prove useful for the classification of the Tipulidce of other countries. I divide the American species of Tipulidce, at present known, into three sub-families, as follows : — I. The auxiliary vein ends in the first longitudinal vein; besides the hu- meral cross vein, there is no other cross vein between the auxiliary vein and the first longitudinal vein or the costa ; last joint of palpi very long, filiform, generally longer than the three preceding taken together. TIPULINA. II. The auxiliary vein ends in the costa; there is a cross vein between it and the second longitudinal vein, generally more or less approxi- mated to the tip of the Jirst longitudinal vein, sometimes more 1 2 D1PTERA OF NORTH AMERICA. removed from it towards the base of the wings ; palpi in most cases short ; last joint either very short, or, if elongated, hardly longer than the three preceding taken together LIMNOBINA. III. Sixth longitudinal vein (anal vein of the former authors) obsolete. PTYCHOPTEKINA. The two first large groups are further distinguished by the structure of the genital organs of the male, which, in most of the Limnobina, ai-e represented by a forceps, consisting of two movable, fleshy lobes, with some delicate horny appendages ; whereas in the Tipulina the forceps is a very compound organ, consisting of manifold horny pieces, which, being inclosed between the dorsal and ventral plates of the two last abdominal segments, produce the club-shaped appearance peculiar to the tip of the £ abdomen of this sub-family. Thus founded, not on a single character, but on a combination of cha- racters taken from, various organs, the definition of the two great sub- families hardly leaves any doubtful case among the Tipulidx which I know of. If one character fails to give a satisfactory result, the others will generally remove at once any doubt as to the relationship of the spe- cies. Thus, the last joint of the palpi of Pedicia is unusually long ; but the auxiliary vein, ending in the subcosta, and the presence of a cross vein between it and the second vein, immediately refer it to the Limnobina, where this genus naturally belongs on account of its habitus. (I have neglected to examine this last joint in fresh specimens of Pedicia, but it appears very probable that its disproportion with the other joints is far from being so striking as is the case in the Tipulina.*) In Rhamphidia, the last joint of the palpi is represented by former writers to be elongated. I had 110 occasion to ascertain, on living speci- mens, how far this is correct. But the presence of the cross vein places this genus among the Limnobina, where it naturally belongs by its habitus. And even if this character should not be considered as sufficient, on account of the extreme shortness of the cross vein, placed at the very tip of the auxiliary vein, the structure of the male genitals removes all doubt. In some Pachyrhinae there is, near the tip of the auxiliary vein, a blackish dot, which might perhaps be mistaken for a cross vein. But should it even be considered as a rudiment of one, the length of the last joint of the palpi, the structure of the male genitals, etc., assign its place among the Tipulina, where its habitus most evidently refers it. In the singular genus Antocha 0. S. the costa and the auxiliary and the first longitudinal veins coalesce insensibly together, so that there is no room left for a cross vein. In this case, the shortness of the palpi and the structure of the male genitals decide of its location among the Lim- nobina. I refer to my sub-family of Ptychopterina the genera Ptijclwptcra, Biltaco- morpha, and Protoplasa (with its congener Macrochile Loew). As to the distribution of the other genera among the two remaining sub-families, I agree with Mr. Loew, with the following exceptions: — MYCETOPHILIDAE. 13 JRhamphidia, as shown above, is more related to the Limnobina than to the Tipulina; by all means it ought not to be separated from Elfphanto- myia, as Mr. Loew does it. In my paper on the Limnobina of this country, I have explained the close relationship of both. Elephant omyia is nothing but a Rhamphidia with an enormously prolonged rostrum, the development of which has also modified the character of the palpi inserted at its tip. (Geranomyia, with its long rostrum and stunted palpi, stands precisely in the same relation to Dicranomyia.) Gynoplistia Westw. (an Australian genus) and Polymera Wied., both of which I know only from plates and descriptions, belong, I presume, to the Limnobina. Hesperinus, Walk, belongs to iheSibionidas (see my note in that family). About Pteror.osmus Walk. I have no opinion whatever, not having seen it, and not being able to establish any opinion on the description. To the list of genera already found in North America, I have to add Dolichopesct, Neplirotoma, and probably Cylindrotoma, as I possess a species apparently closely allied to the latter. Finally, it is the place here to notice that Mr. Westwood (Lond. and Edinb. Philos. Magaz., 1835) has described a Gynoplistia annulata from North America. As it is hardly probable that an Australian genus should also be represented on this con- tinent, it is to be presumed either that the genus is different, or that the statement is based upon an error of locality. Gynoplistia has pectinated antennae in both male and female. OSTEN SACKEK. FAM. VII. MYCETOPHILIDAE. Charact. — Ocelli three or two, in the latter case often hardly perceptible. Thorax without a transverse suture ; wings without discal cell. Coxse much elongated ; all the tibise with spurs. On the whole, the Mycetophilidce are so easily known that it would be superfluous to give any more details about them. The genus differing the most from the rest is Sciara, which shows some affinity with the Cecidomyidce. The genera hitherto introduced in this family are the following: CORDYLA Meig. (= Brachypaljms Macq.), MYCETOPHILA Meig., AZANA Walk., LEJA Meig., BOLETINA Staeg. (= Leptomorphits Walk.), SCIOPHILA Meig., SCIOBTA Loew, TETRAGONEURA Winn., GNORISTE Meig., ASINDULUM Lair. ( = Macrorhyncha Winn.), CEROPLATUS Fair., LEPTOMORPHUS Curt., DIADOCIDIA Ruth. ( = Macroneura Winn.), ACLADA Loew, MYCETOBIA Meig., PLESIAS- TINA Winn., DITOMYIA Winn. (Symmerus Walk.), PLATYURA Meig., PLATYROPTILON Westw., MACROCERA Meig., BOLITOPHILA 14 DIPTERA OF NORTH AMERICA. Meig. (= Messala Curt.), HETEROTRICHA Loew, DIANEPSIA Loew, SCIARA Fabr., and the genus DIOMONUS Walk., which is unknown to me. I have not mentioned the genus SYNAPHA- Meig. , because it seems to have been founded on an individual of a species of Leja, which possessed an irregularly formed neuration ; at least as far as I know, no second specimen of Synapha has been captured since Meigen's time, while a similar anomaly of neuration of the wings has been observed several times in other Diptera. Our knowledge of N. A. Mycetophilidceis exceedingly incomplete. I have seen species of the genera MYCETOPHILA, BOLETINA, Scio- PHILA, TETRAGONEURA, PLESIASTINA, DITOMYIA, PLATYURA, MA- CROCERA, BOLITOPHILA, and SCIARA. Besides these, the existence of CEROPLATUS seems to be certain, and the genus DIOMONUS, which I bave never seen, is founded on a N". A. species. FAM. YIIL SIMULIDAE. Charact. — Ocelli none. Thorax without transverse suture. Wings with very short hair only visible under a very high magnifying power ; legs short, tibiae without spurs ; posterior tibire and first joint of the hind tarsi dilated. The present family comprises only the genus SIMULIUM Latr., rich in species and which cannot be placed in any other family. It does not seem to be less rich in species in N. A. than in Europe. FAM. IX. BIBIONIDAE. Charact. — Ocelli three. Thorax without transverse suture ; prothorax much developed. Wings without discal cell; coxae not prolonged ; empodium proportionally long, whereas the pulvilli are wanting in some of the genera. The family of Bibionidce is divided into two sections sharply separated from each other, and which it would be proper to con- sider as distinct families. In the SCATOPSINA, which form the first section, the palpi are very short, the pulvilli wanting, the tibias without spurs ; the genera belonging to them are : SCATOPSE Geoff}'., ASPISTES Meig., ARTHRIA Kirby. To the second section, the BIBIONINA, belong: DILOPHTJS Meig., BIBIO Geoffr., PENTHE- RHYPHIDAE — XYLOPHAGIDAE. 15 TRIA Meig., CRAPITULA Gimm., PLECIA Wied., EUPEITENUS Macq., PACHYNEURA Zett., and SPODITJS Loew. Species of the genera SCATOPSE, ARTHRIA, DILOPHUS, BIBIO, PLECIA, and EUPEITENUS are known to occur in JST. A. Note. — At the time when this was written by Mr. Loew, neither he nor I possessed specimens of the genus Hesperinus, which its author, Mr. Walker, referred to the Tipulidse. Having obtained specimens since, collected by Mr. R. Kennicott near the Great Slave Lake, I found that Hesperinus be- longs to the Bibionidas, and is apparently synonymous with Spodius Loew. Accordingly, Hesperinus Walk, is to be added to the genera of this family occurring in N. A., and stricken out from among the Tipulidae. 0. S. FAM. X. KHYPHIDAE. Charact, — Ocelli three. Thorax without transverse suture ; wings with a perfect discal cell ; empodiurn similar to a pulvillus ; pulvilli want- ing. Of this family also a single genus, RHYPHUS Meig., is known, which has representatives in Europa, Asia, and N. A. Observation — There is a genus Epidapus Hal., remarkable for having no wings and no poisers, which I have omitted in the pre- ceding enumeration of families, because I do not know it. It is quite impossible to place it among the Mycetophilidce, as Walker does, if we characterize the families as we have done. It rather seems to find its place among the Cecidomyidce ; but there is nothing decisive to be said without the examination of fresh specimens. II. BRACHYCERA. FAM. XI. XYLOPHAGIDAE. Charact. — The three basal cells very prolonged, the third longitudinal vein furcate ; both intercalary veins always present ; the marginal vein encompassing the whole wing ; the third joint of the antennae annu- lated or divided into separate joints, always without style or termi- nal bristle. Tibiae with spurs ; the empodium very developed and pulvilliform. The genera belonging here are: XYLOPHAGUS Meig., PACHY- STOMUS Latr., SUBULA Meig., ELECTRA Loew, CURYSOTHEMIS 16 DIPTERA OF NORTH AMERICA. Loew, RACHICERUS Hal, COENOMYIA Latr., and ARTHROPEAS Loew. The new genera CYCLOTELUS, PHYCUS, and DIMASSUS, es- tablished by Walker as belonging to the Xylophagidce, belong in fact to the Therevidce; likewise NONACRIS must be removed here, but Walker's observations on its characters are far too superficial to admit of any certainty in fixing its place ; also DIALYSIS on ac- count of the hairy, bristle-like antennal tip ascribed to it by Walker might seem to be erroneously located among the Xylophagidce, the characters of which he appears not to have understood. Ccenomyia Latr. has often been separated from the Xylophagidce and considered as forming a distinct family: Ccenomyidos, or for- merly Sicarii. This seems to have been caused by the body of Ccenomyia. being stout, whereas that of Xylophagus and Subula is of a slender form. Moreover, the different form of the palpi, which in Ccenomyia are rather cylindrical and ending in Xylophagus and Subula in a button-shaped thickening, have been made use of to justify the separation. But within a recent time forms of Cceno- myidee have been discovered in which the structure of the body and palpi is such as to form a link between them and the Xylopha- gidce; from this, as well as from the agreement of their other essential characters, results the necessity of reuniting them. In case the separation should be maintained, Arthropeas ought to be placed among the Ccenomyidce. The family of Xylophagidce may be divided into three sections : CCENOMYINA, RACHICERINA, and XYLOPHAGINA. The Ccenomyina are characterized by their robust structure, the third joint of the antennae being annnlated and pointed towards its end, the palpi being cylindrical. The genera comprised here are CCENOMYIA and ARTHROPEAS. In the Rachicerina the third joint of the antennas is divided into separate and frequently very numerous articula- tions, and the palpi are rather club-like; the body is less heavy than in the Ccenomyina, but less slender than in the Xylophagina. The genera ELECTRA, CIIRYSOTHEMIS, and RACHICERUS belong here. The Xylophagina have the slenderest bodies; the third joint of the antenna? is annulated and never strikingly pointed ; the palpi have at their end a button-shaped thickening. The genera SUBULA, XYLOPHAGUS, and PACHYSTOMUS may be referred here. I am acquainted with N. A. species belonging to the genera CCENOMYIA, ARTHROPEAS, RACHICERUS, SUBULA, and XYLOPHAGUS. STRATIOMYIDAE. It Two of the species of RACHICERUS cannot be well placed in this genus without a modification of its characters. Observation. — I have to mention here the genus BOLBOMYIA, which I established on two fossil species found in Prussian amber. When I published in 1850 my observations on the Dipterological Fauna of amber, I thought it would be best placed among the Xy- lopliagidcK. But I perceive from a N. A. specimen belonging to Bolbymyia that its claim to that place is more than doubtful, and at the same time that it is quite as difficult to assign it a fit place elsewhere. FAM. XII. STRATIOMYIDAE. Charact. — Three basal cells much prolonged ; veins of the two main trunks very crowded anteriorly; both intercalary veins usually existing; costal vein reaching only to the middle of the wing. Third joint of the antennse annulated, sometimes divided into several portions. Tibise without spurs ; empodium much developed, pulvilliform. This family, rich in various forms, may be divided into five sharply circumscribed sections. The first is that of the BERIDINA, easily distinguished by the abdomen not showing five segments, as in the other sections, but seven, a difference caused only by the smallness of the two last segments and their concealed situation in the other sections. The Beridina have often been placed in the family of Xylophagid.ee, but figure more naturally among the Stra- iiomyidce. The genera belonging to them are : METOPONIA Macq. (= Inopus Walk.), BERIS Latr., ACTINA Meig., EXODONTHA Rond., ACANTHOMYIA Sch., DiPHYSA Macq. , CAMPEPROSOPA Macq., perhaps also EXOCHOSTOMA Macq. ; also the genus CHIROMYZA Wied., which does not differ from Xenomorpha Macq., may be re- ferred to them. The second section is that of SARGINA, rather agreeing in the form of the body with the Beridina,, and even with the Hermetina, but differing from the former by the abdomen consisting apparently of five segments, and from the latter by the eyes of the males being much more approximated than those of the females. As genera of this section may be mentioned CACOSIS Walk., ACROCHYOTA Wied., EUDMETA Wied., ANALCOCERUS Loew, SALDUBA Walk., TOXOCERA Macq., HOPLISTES Macq., RAPHIOCERA .Mzcg.jBASENTiDEMA Macq., DICRANGI'HORA Macq., CHRYSOCHLORA Macq., PTECTICUS Loew, MEROSARGUS Loew, PEDICELLA Big., 2 18 DIPTERA OF NORTH AMERICA. CHRYSONOTUS Loew, SARGUS Fabr,, CLORISOMA Rond., CHRYSC- MYIA Macq., and MICROCHRYSA Loew. The third section, HERME- TINA, is well characterized by the elongated abdomen, the eyes, which are equidistant and very remote in both sexes, and the peculiar structure of the antennae, the third joint of which is transformed into a ciliated lamel. The genera HERMETIA Latr., THORASENA Macq. belong to them. The fourth is formed by the ODONTOMYINA, which are distinguished from the foregoing by their broad body and from the following section by the less convex ab- domen and especially by the neuration, the longitudinal veins of the Odontomyina being more crowded anteriorly, the discal cell being smaller, hexagonal or pentagonal, never large or subquad- rate ; moreover, both intercalary veins are usually present, while the posterior one is almost always wanting in the Pachygastrina. The following genera may be referred to the Odontomyina: CY- PHOMYIA Wied., "GnoRDONOTA Gerst., EUPARYPHUS Gerst., PYCNO- MALLA Gerst., ALLIOCERA Saund., STRATIOMYS Geoff"., ODONTOMYIA Meig., INERMYIA-B^., NEMOTELUS Geojfr., OXYCERA Meig., HETE- ROXYCERA Big., EPHIPPIUM Latr., CLITELLARIA Meig., CYCLOGAS- TER Macq. (= Lasiopa Brull.), ARTEMIDA Walk., AISSA Walk., METABASIS Walk., PROMERANISA Walk. The fifth section is that of the Pachygastrina ; it is distinguished by the longitudinal veins being less crowded towards the costal border, by the magnitude and quadrangular form of the discal cell, the almost general want of the posterior intercalary vein, the short, generally much inflated, abdomen, and its segments soldered together in some genera. The genera belonging here are : PACIIYG ASTER Meig., LOPHOTELES £oew, STERNOBRITHES Loeiv, PLATYNA Wied., BIASTES Walk., PTILOCERA Wied., CHATJNA Loew, BLASTOCERA Gerst., SPYRIDOPA Gerst., PANACRIS Gerst,, NERTJA Walk., CULCUA Walk., EVAZA Walk., ANACANTHELLA Macq.; perhaps also PHYLLOPHORA Macq., and ANISOPHYSA Macq. To which section of the Stratiomyida the genera Solva, Amp- salis, Tracana, Rosapha, Tinda, Saruga, Gabaza, Adraga, and Obrapa, lately formed by Walker, are to be referred, the extreme vagueness of the characters ascribed to them does not allow me to determine. The N. A. species which are now known to me belong to the following genera: I. Beridina: METOPONIA, ACTINA ; II. Sargi- ACANTHOMERIDAE — TABANIDAE. 1 9 na: SARGUS, MICROCHRYSA : III. Hermetina: HERMETIA : IY. Odontomyina: CYPHOMYIA, EUPARYPHUS, STRATIOMYS, ODONTO- MYIA, NEMOTELUS, OXYCERA, CLITELLARIA ; V. Pachygastrina : PACHYGASTER, CHAUNA. FAM. XIII. ACANTHOMERIDAE. Charact. — Basal cells much prolonged ; longitudinal veins not crowded together anteriorly ; two intercalary veins always present ; marginal vein running round the whole border of the wing. Oral parts with four bristles, even in the male. Third joint of the antennae annu- late. Tibiae without spurs ; ernpodiura developed to a pulvillar form. This small family contains only the two genera ACANTHOMERA Wied. and RAPHIORHYNCHUS Wied. It differs from the Stratio- myidce by the longitudinal veins not being crowded together ante- riorly, by the marginal vein encompassing the whole border of the wing and by the fourth cell of the posterior margin being closed. It differs from the Tabanidce in the form of the oral parts and by the tegulre being very little developed; as to the oral parts, I have no absolute opinion of my own, but must rely on the communica- tions of others. No species of this family has been as yet discovered in N. A.. FAM. XIV. TABANIDAE. Charact. — Three basal cells much prolonged ; third longitudinal vein fur- cate ; two intercalary veins always present; marginal vein running round the whole border of the wing ; tegulse rather large. Proboscis of the male with four, of the female with six bristles. Third joint of the antennas annulate, rarely divided into distinct joints, always without style or bristle ; empodium much developed and pulvilli- form. The Tabanidce are easily distinguished from the foregoing fami- lies by the structure of the oral parts and by the size of the tegulos. On account of the presence or absence of spurs at the end of the posterior tibia? they may be divided into the sections of PANGONINA and TABANINA ; the former often, but not always, possess ocelli, whereas, according to the observations hitherto made, they are always wanting in the latter section. To the Pangonina belongs, firstly, the genus PANGONIA Lair, 20 DIPTERA OP NORTH AMERICA. with the genera separated from it and partly connected with each other by passages : Philoliche Hffgg. , Dicrania Macq. , PeJeco- rhynchus Macq., Cadicera Macq., Nuceria Walk., Melpia Walk., Scaptia Walk., Tacina Walk., Phara Walk., Clanis Walk., Osca Walk., Scione Walk., Ph'nthina W silk. , ScarphiaWalk., and Lilcea Walk.; moreover, the genera: SCEPSIS Walk., SILVIUS Meig. , ME- SOMYIA Macq., TABANOCELLA Big., ECTENOPSIS Macq., RHINOMYZA Wied., ERODIORHYNCHUS Macq., GASTROXIDES Saund., PRONOPES Loew, CHRYSOPS Meig., and NEMORIUS Rond. The Tabanina comprise the genera: TABANUS Linn., with The- rioplectes Zell., DICHELACERA Macq., LEPISELAGA Macq., SELA- SOMA Macq., HADRUS Pert., DIABASIS Macq., ACANTHOCERA Macq., DASYBASIS Macq., HEXATOMA Meig., and HJEMATOPOTA Meig. The N. A. species which we are acquainted with belong to the genera: I. Pangonina: 1. PANGONIA Latr., 2. SILVIUS Meig., 3. CHRYSOPS Meig. II. Tabanina: 4. TABANUS Linn., 5. LEPI- SELAGA Macq., 6. DIABASIS Macq., 7. EJEMATOPOTA Meig. FAM. XV. LEPTIDAE. Charact. — Three basal cells much prolonged ; third longitudinal vein fur- cate ; two intercalary veins always present ; marginal vein running round the whole border of the wing. Third joint of the antennae simple, with a simple or thickened styliform bristle. Tibiae with spurs ; empodium much developed, pulvilliform. This family is very easily distinguished from the foregoing families by the simple third joint of its antennae. A division into sections has not been attempted yet, and would be useless for the small number of genera hitherto known. The genera belonging to this family are as follows: DASYOMMA Macq., CHRYSOPILA Macq., TRIPTOTRICHA Loew, LEPTIS Fair., VERMILEO Macq. (= Psammorycter Blanch.), ATHERIX Meig., NODUTIS Meg. (= Ibisia Rond.), and SPANIA Meig. ( — Ptiolina Zett. = Leptipalpus Rond.). The location here of the genus SynecJtes is one of the many errors which we meet with in the writings of Mr. Walker. I know N. A. species belonging to the genera : 1. CHRYSOPILA Macq., 2. LEPTIS Fabr., 3. TRIPTOTRICHA Loew, 4. ATHERIX Meig. Mr. Walker has also recorded a species of the genus Spania Meig. CYRTIDAE — HIBMONEURIDAE. 21 FAM. XVI. CYRTIDAE. Charact. — Thorax and abdomen inflated. Eyes occupying the greatest part of the head. Tegulae vaulted, exceedingly large. Wings naked, with variable neuration, sometimes very intricate, some- times very incomplete ; the basal cells, when present, are of consi- derable length. Terminal joint of the antennae simple. Tibiae without spurs; empodium much developed, pulvilliform. This family is divided into the two sections of CYRTINA and ONCODINA. In the former section the veins of the wings are strong and well developed, and the neuration is usually rather complicated. It contains the genera: CYRTUS Latr., PTEROPEXUS Macq., EPI- CERINA Macq., PANOPS Lam. (= Mesophysa Macq.), LASIA Wied., EULONCHUS Gerst., PSILODERA Griff. (—- Mesocera Macq.), PTERO- DONTIA Griff., ASTOMELLA L. Duf., PHYLLIS EricllS., OCN^EA Ericlis. ( — Eriosoma Macq. = Exelasis Walk.), PIALEA Erie/is., OBSEBIUS Cost. (= Pithogaster Loew), PHYSEGASTER Macq. The section Oncodina is distinguished by the anterior veins of the wings alone being completely developed, whereas the posterior ones are not only very incomplete, but also disappear gradually, and frequently are not completely connected. The genera which belong here are: ONCODES Latr. (==. Henops Meig.), TERPHIS Erichs. and PHILOPOTA Wied. The N. A. species known to me belong to the genera: I. Cyr- tina: 1. CYRTUS Latr., 2. OCN.SA Erichs., 3. PTERODONTIA Griff., 4. ACROCERA Meig., 5. a genus hitherto unnamed, and related to Obsebius, 6. EULONCHUS Gerst., occurring in California. II. On- codina: 7. ONCODES Latr. FAM. XVII. HIRMONEURIDAE. • Charact. — Three basal cells much prolonged ; veins of the wings varying ; third longitudinal vein furcate; the two intercalary veins present. Third joint of the antennae simple ; terminal bristle simple or simi- lar to a style, and consisting of several joints. Tibiae without ter- minal spurs ; empodium pulvilliform, but more frequently minute as well as the pulvilli. This family, usually called Nemestrinidce, must be divided into the two sections HIRMONEURINA and RHYNCHOCEPHALINA. The first comprises the genera: HIRMONEURA Meig., EXERETONEURA 22 DIPTERA OF NORTH AMERICA. Macq., COLAX Wied., TRICHOPSIDEA Westw., and SYMMICTUS Loew, all of which are characterized by their very short proboscis. To the second belong the genera: FALLENIA Meig., NEMESTRINA Wied., MEGTSTORHYNCHUS Macq., TRICHOPHTHALMA Westw., and RHYN- CHOCEPHALUS Fisch. We are only acquainted with a single N. A. species belonging to HIRMONEURA Meig. FAM. XVIII. MID A SID AE. Charact. — Three basal cells much prolonged ; third longitudinal vein fur- cate ; posterior intercalary vein always present, whereas the ante- rior one is often wanting ; veins of the wings varying ; wings naked. Antennse clavate with the third joint consisting of several distinct segments. Under lip fleshy. Ernpodium very little developed. To this family belong the genera: MIDAS Fair., CEPHALOCERA Latr., RHOPALIA Macq., and DOLICHOGASTER Macq. — POMACERA Macq. may also be placed here till its true place is found. The N. A. species hitherto known belong only to the genus MIDAS Fabr. FAM. XIX. ASILIDAE. Charact. — Three basal cells much prolonged. Third longitudinal vein of the wings furcate, the two intercalary veins always present. Third joint of the antennae simple; under lip forming a horny sheath; empodium similar to a horny bristle. This family, rich in species of the most varied forms, is divided into three sections. The first of them is that of the DASYPOGO- NINA, differing from the two others by its second longitudinal vein running into the border of the wing, whereas in the others it unites with the first longitudinal vein before the border of the wing. The considerable number of genera requires a further division into two subordinate groups, the first of which comprises those genera in which the anterior tibias end in a hooked spine, whereas the genera of the second portion have no such spine. Consequently the genera belonging to the first group of Dasypogonina are as follows: DASYPOGON Meig., SAROPOGON Loew, LASTAURUS Loew, MORIMNA Walk., CYRTOPHRYS Loew, LAPARUS Loew, BRACHYRHO- PALA Macq., CHEILOPOGON Bond., LAGODIAS Loew, and PEGE- . ASILIDAE. 23 SIMALLUS Loew. Those of the second group are : MICROSTYLUM Macq., MEGAPOLLION Walk., XIPKOCERUS Macq., DOLICHODES Macq., DISCOCEPHALA Macq,, SENOBASIS Macq., PLESIOMMA Macq. , STENOPOGON Loew, BATHYPOGON Loew, HABROPOGON Loew, HOLO- TOGowLoew, ERiopOGONZoew, HETEROPOGON Loew, ISOPOGON Loew, OLIGOPOGON Loew, STICHOPOGON Loew, SAROPOGON Loew, DICRA- NUS Loew, TRICLIS Loew, EUARMOSTUS Walk., PROLEPSIS Walk., CODULA Macq., CABASA Walk., PHELLUS Walk., PHILAMMOSIUS Walk., GASTRICHELIUS Rond., DACTILISCUS Rond., ELASMOCERA Rond., PHENEUS Walk., CROBILOCERUS Loew, PYCNOPOGON Loew, ANAROLIUS Loew, ACNEPHALUM Macq., SISYRNODYTES Loew, RHA- DINUS Loew, CERATURGUS Wied., DIOCTRIA Meig., TERATOPUS Loew, DASPLETIS Loew, LAPHYCTIS Loew, SCYLATICUS Loew, II Y- PENETES Loew, SPANURUS Loew, RHABDOGASTER Loew, DAMALIS Wied., LEPTOGASTER Meig. (=GonypesIjsAlT.), EUSCELIDIA Westw., and LASIOCNEMUS Loew. The second section of the Asilidce are the LAPHRINA; it agrees with the third in the second longitudinal vein running into the first, but differs from it in the style of the antennae either being thick and stout, and generally only rudimentary, or entirely want- ing, whereas the antennas of the third section possess a distinct terminal bristle. The genera of the second section are: LAPHRIA Meig., LAMPRIA Macq., HOPLISTOMERA Macq., MzGAPODA.Macq., RHOPALOG ASTER Macq., MICHOTAMIA Macq., ATOMOSIA Macq., LAXENECERA Macq., TAPINOCERA Macq., PHONEUS Macq., LA- PHYSTIA Loew, NUSA Walk., SCANDON Walk. , DASYLLIS Loew, LA- MYRA Loew, LAMPROZONA Loew, DASYTHRIX Loew, THEREUTRIA Loew, AMPYX Walk., CORMANSIS Walk., CHCERADES Walk., Acu- RANA Walk., PSEUDORUS Walk., POGONOSOMA Rond., and DYSERIS Loew. The third section is that of the ASILINA, which is characterized by its second longitudinal vein running into the first, and by its antennae having a distinct terminal bristle. The genera belonging to this section are: MALLOPHORA Macq., PROMACHUS Loew, ALCI- MUS Loew, PHILODICUS Loew, CRASPEDIA Macq. (= Blepharotes Westw.), POLYPHONIUS Loew, APOCLEA Macq., ERAX Macq., ERIS- TICUS Loew, PROCTACANTHUS Macq. , STENOPROSOPIS Macq. , SYNOL- cus Loew, DYSCLYTUS Loew, LOPHONOTUS Macq., TRICHONOTUS Loew, DASOPHRYS Loew, PROTOPHANES Loew, DYSMACHUS Loew, EUTOLMUS Loew, MACHIMUS Loew, MOCHTHERUS Loew (==• Helta- 24 DIPTERA OF NORTH AMERICA. moneura Big.), STILPNOGASTER Loew, EPITBIPTUS Loew, ITAMUS Loew, TOLMERUS Loew, CERDISTUS Loew, PROAGONISTES Loew, ASILUS Linn., ECCOPTOPUS Loew, RHADIURGUS Loew, PAMPONERUS Loew, ANTJPHRISSON Loew, ECHTHISTUS Loew, ANTIPALUS Loew, PHILODICUS Loew, LECANIA Hacq., ATRACTIA Macq., and OMMA- TIUS Wied. Most of the N. A. Asilidce, but by no means all, may be placed in the genera hitherto established. I give, as far as I am able to do so under s'ich circumstances, the following list of genera known to me as occurring on that continent: — I. Dasypogonina: 1. DASYPOGQN Meig., with several species which will require the formation of some new genera; 2. MICRO- STYLUM Macq., 3. DiscocEPHALA Macq., 4. PLESIOMMA Maeq., 5. STENOPOGON Loew, 6. BATHYPOGON Loeiv, 1. LASTAURUS Loew, 8. EUARMOSTUS Walk., 9. PHENEUS Walk., 10. STICHOPOGON Loew, 11. CERATURGUS Walk., 12. LEPTOGASTER Meig. The X. A. spe- cies recorded as belonging to DIOCTRIA Meig. are no Dioctrice at all; at least DIOCTR. OCTOPUNCTATA Say is by no means a true Dioctria. II. Laphrina: 13. MEGAPODA Macq., 14. LAPHRIA Meig., 15. ANDRENOSOMA Rond., 16. LAMPRIA Macq., 17. ATOMOSIA Macq.t 18. LAPHYSTTA Loew. lll.'Asilina: 19. MALLOPHORA Macq., 20. PfiOMACHUS Loew, 21. ERAX Macq., 22. PROCTACANTHUS Macq., 23. MOCHTHERUS Loeiv, 24. EPITRIPTUS Loew, 25. OMMATIUS Wied. FAM. XX. THEREUIDAE. Charact. — Three basal cells much prolonged ; the two intercalary veins present ; third longitudinal vein furcate. Antennze with a terminal style of variable form, sometimes wanting. No empodium. Under lip fleshy. The principal genera belonging to this family are: XESTOMYZA Wied., FARYPHORA Loew, CIONOPHORA Egg., EXAPATA Macq., THEREUA Latr., ECTINORHYNCHUS Macq., ANABARHYNCHUS Macq., TABUDA Walk., CYCLOTELUS Walk., PHYCUS Walk., and DIMASSUS Walk. The N. A. species with which I am acquainted may be conve- niently placed under the genus THEREUA Latr. In case the genus BOMBYLIDAE. 25 PsilocepJiala Zett,, which does not appear to be well founded, should be admitted, some species with naked faces would be located in it. FAM. XXI. BOMBYLIDAE. Charact. — Three basal cells much prolonged ; anterior intercalary vein present almost without exception, the posterior always wanting ; third joint of the antennae simple; empodium quite rudimentary. This, again, is a family exceedingly rich in the most varied forms. A distribution into several tribes would therefore be very useful ; the two sections hitherto adopted, one of which comprises the genera grouped round the genus Bombylius, having a long proboscis, while the second consists of genera more allied to the genus Anthrax, having a short proboscis, do not appear sufficient to embrace all the forms which have hitherto been discovered. I am unable to give a better distribution, and I think it will not be possible to do so until the number of sections is increased to at least five or six. The genera of Bombylidce are as follows: BOM- BYLIUS Linn., EURYCARENUS Loew, TRIPLASIUS Loew, SYSTOZCHUS Loew, SPARNOPOLIUS Loew, DISCHISTUS Loew (= Bombylisoma Rond.), PARISUS Walk., CHORISTUS Walk., HETEROSTYLUM Macg., LASIOPROSOPA Macq., ADELIDEA Macq., ACREOTRICHUS Macq., APATOMYZA Wied., THLIPSOMYZA Meig., AMICTUS Wied., MEGA- PALPUS Macq., PHTHIRIA Meig., CYCLORHYNCHUS Macq., DASYPAL- pus Macq., CROCIDIUM Loew, GERON Meig., APOLYSIS Loew, OLI- GODRANES Loew, MuLio Latr. (= Glossista Rond.), CHALCOCHITON Loew, CALLOSTOMA Macq., SERICOSOMA Macq., TOXOPIIORA Meig., ENICONEURA Macq., LEPIDOPHORA Macq., CORSOMYZA Wied., ECLI- MUS Loew, SYSTROPUS Wied., DOLICHOMYIA Wied., Usio Latr., PLATYPYGUS Loew, CYRTOSIA Perr. , PLEAS Latr., CYLLENiAZortr., LAGOCHILUS Loew, ANISOTAMIA Macq., LOMATIA Meig., ONCODO- CERA Macq., PLESIOCERA Macq., LIGYRA Neivm. , ANTHRAX Scop., ARGYROMCEBA Schin., NEURIA Newm., COMPTOSIA Macq., LITO- RHYNCHUS Macq., SPOGOSTYLUM Macq., ENICA Macq., TOMOMYZA Wied., ARGYROSPILA Rond., EXOPROSOPA Macq., AUTONIA Loew. The N. A. Bombylidce which I have seen may be distributed amongst the following genera: 1. BOMBYLIUS Linn. , 2. SYSTCECHUS Loew, 3. SPARNOPOLIUS Loew, 4. LEPIDOPHORA Westw., 5. TOXO- PIIORA Meig., 6. GERON Meig., 7. SYSTROPUS Wied., 8. PLOAS 26 DIPTERA OF NORTH AMERICA. Latr., 9. ANTHRAX Scop., 10. ARGYROMOZBA Schin., 11. EXOPRO- SOPA Hacq. Moreover, Macquart has founded bis genus Oncodocera on a N. A. species, and described a N. A. species among his Anisotamice, though it seems to be an alien there. Mr. Walker described some N. A. species, which he placed under the genera Apatomyza Wied. and PhtJiiria Meig. FAM. XXII. SYRPHIDAE. Charact. — Three basal cells much prolonged ; third longitudinal vein sim- ple ; a spurious longitudinal vein {vena spuria) between the third and fourth longitudinal veins ; fourth longitudinal vein united at its end with the third ; no intercalary veins. Hypopygiuni unsym- metrical ; no empodium. This is one of the most extensive families and includes about eighty genera, the enumeration of which seems to be superfluous here. A distribution into sections, however desirable, proves exceedingly difficult. To divide the family into genera with an antennal bristle and genera with a terminal style would be no great gain, since the number of the latter is very small. I know the following genera to occur in N. A. : VOLUCELLA Geoffr., TEMNOCERA St. Farg., MICRODON Meig. (= ApJiritis Latr.), CERiA^r., SERICOMYIA Meig., TROPIDIA Meig., SYRITTA St. Farg., XYLOTA Meig., MALLOTA Meig., BRACHIPALPUS Macq., MILESIA Latr., SPHECOMYIA Latr., SOMULA Macq., CHRYSOTOXUM Meig., MlXTEMYIA Macq., MALLOTA Meig., HELOPHILUS Meig., ERISTALIS Latr., PLAGIOCERA Macq., OCYPTAMUS Macq ., BACCHA Fair., SPHEGINA Meig., RHINGIA Fair., ORTHONEURA Macq. (= Cryptineura Big.), PARAGUS Latr., CHEILOSIA Meig., CflRYSO- CHLAMYS Bond., DOROS Meig., DIDEA Macq. (= Enica Meig.), MELITHREPTUS Loew (= Sphserophoria Macq.), MESOGRAMMA Loeio, SYRPHUS Fair., SCJEVA Fair., PLATYCHEIRUS St. Farg. It results from the remarks of some authors that species of the genera: Pipiza Fall., Chrysogaster Meig., EpistropJie Walk., Po- lydonta Macq., and Merodon Latr. occur with certainty in N. A. The genus Chymopkila Macq. founded on a N. A. species must be entirely blotted out from the list of genera. For it is evident that Bigot is right in stating that the specimen on which it was MYOPIDAE. 27 founded was a composition of a body of a Microdon with the head of a Conops. The genus Toxomerus Macq. has not been mentioned in the above list, it being quite untenable. I judge Dimeraspis Xewm. to be identical with Microdon. Psarus has been omitted, because the species described under this name must be placed in other genera. The statement of Eumerus Meig. occurring in N. A. is founded merely on an observation of Walker, and therefore requires further confirmation. Macquart records a 1ST. A. species of the genus Psilota Meig., but this genus having been misunder- stood by most authors, I do not venture now to mention it among those truly represented in N. A. FAM. XXIII. MYOPIDAE. Charact. — Three basal cells large, the third closed, more or less remote from the posterior border ; all longitudinal veins simple ; no inter- calary vein. Eyes in both sexes broadly separated ; proboscis, with few exceptions, much prolonged ; rnaxillse small ; the third joint of the antennae with an apical style or a thick dorsal bristle. Hypo- pygiurn symmetrical, turned under the abdomen. Empodiuin wanting. Omitting the untenable genera into which the genus Conops has been subdivided by Rondani and the genus Myopa by Ferris, we mention here the genera: CONOPS Linn., PLEUROCERINA Macq., ZODION Latr., MYOPA Latr. and STACHINIA Macq. This family has been divided by some authors into two families: Conopidse and Myopidse, the former containing those genera which have an apical style on the antennae, the latter being characterized by a dorsal bristle of the antennae. I cannot approve of this divi- sion at all, since the difference between a style and a bristle, and the difference of an apical and a dorsal position, according to all experience, only furnishes characters of very inferior value for the systematic arrangement, as we see in the families Stratiomydae, Bombylidse, Syrphidde, Hybotidse, Dolichopidse, etc., where this organ is sometimes apical, sometimes dorsal. "We might as well form two families on account of the proboscis being either straight or geniculated. But the conspicuity of the difference in the struc- ture of the antenna? may serve to form two sections in the family, CONOPINA and MYOPINA, the former of which would contain the genera Conops and Pleurocerina, the latter the genera Zodion, Myopa, and Stachynia. 28 DIPTERA OF NORTH AMERICA. The N. A. species which I know belong to the genera: 1. Co- NOPS Linn., 2. ZODION Latr., 3. MYOPAZafr-., 4. STACHYNIA Macq. FAM. XXTV. PIPUNCULIDAE. Charact. — Three basal cells much prolonged, the hindmost closed near the border of the wing ; third longitudinal vein simple, the fourth sometimes almost entirely wanting, sometimes furcate when perfect ; no intercalary vein. Head almost entirely occupied by the eyes, front and face exceedingly narrow ; antennae with a basal bristle. Hypopygium unsymrnetrical. Tibiae without spurs ; empodiuni wanting. This family only comprises the three genera: NEUROPHOCERUS Zett., PIPUNCULUS Lair, and CHALARUS Walk. The N. A. species known to me belong all to the genus PIPUNCULUS Latr. FAM. XXV. SCENOPINIDAE. Charact. — Three basal cells very large ; the third closed rather far from the border of the wing; third longitudinal vein furcate ; no inter- calary vein ; third joint of the antennae without style or bristle. No empodium. This family possesses so many peculiarities that it is very difficult to find a fit place for it among the other families, though it ex- hibits much affinity with some of them. I would especially point out the Bornbylidce as deserving a closer comparison in order to investigate their true relationship. At present it seems best to follow those authors who have considered the genus SCENOPINUS as the type of a separate family. Some species of SCENOPINUS occur in X. A. FAM. XXVI. PLATYPEZIDAE. Charact. — Three basal cells rather large, the hindmost always ending acutely, at more or less distance from the border of the wing ; third longitudinal vein simple ; no intercalary vein. Antennae with an apical bristle. Hypopygium symmetrically turned under the abdo- men. Middle tibiae with spurs ; empodium wanting. The genera which belong to this family are : PLATYPEZA Meig., CALLOMYIA Meig., OPETIA Meig., and PLATYCNEMA Zett. LONCHOPTERIDAE — HYBOTIDAE. 29 I know only one species of PLATYCNEMA, one species of CALLO- MYIA, and two species of PLATYPEZA occurring in N. A. FAM. XXVII. LONCHOPTERIDAE. Charact. — Three basal cells of moderate size, of nearly equal length ; fourth longitudinal vein furcate and united with the fifth near the base. Antennae with an apical bristle. Empodium wanting. This family is also founded on a single genus which cannot be placed in any other family. Though in LONCHOPTERA the basal cells are by no means large, yet their structure and the great deve- lopment of the sixth longitudinal vein seems to prove that this family should be reunited with one of those already mentioned. However, by the form of its neuration and its anal parts it differs so widely from them, that it is very difficult to state in what their affinity consists. Mr. Walker has lately added the genus CADREMA to the family of Lonchopteridce. LONCHOPTERA is found in N. A . FAM. XXVIII. HYBOTIDAE. Charact. — Three basal cells complete, rather large, the third only a little shorter than the second ; posterior transverse vein of the base generally running perpendicularly or at a somewhat acute angle into the sixth longitudinal vein, and thus not being parallel to the posterior border of the wing ; third longitudinal vein frequently furcate ; anterior intercalary vein often wanting, posterior never present. First joint of the antennae not much shortened, the third more frequently with a bristle than with a style, the bristle some- times dorsal instead of being apical. Empodium membranaceous and linear. The three families : Hybotidse, Empidse and Tachydromidse run into each other so insensibly, that it is very difficult to indicate sharp limits between them. If we select this or that character as being of greater importance, we shall always obtain a different result as to these limits. I maintain the family Hybotidse only with the view of falling in with the usual arrangement, since I am fully satisfied that there is no sharp limit to be drawn between it and that of the Empidse. For neither the more convex thorax, nor the horizontal direction of the proboscis, nor the form and position of the palpi, nor the simplicity or furcation of the third longitudi- 30 DIPTERA OF NORTH AMERICA. nal vein, nor the presence or absence of the anterior intercalary vein are characters, on which — whether we use them singly or in any combination — we can found a satisfactory or sharp defini- tion of both families. The resemblance of some Hybotidse with some Bombylidse cannot be denied, but their place will never be doubtful if we consider, that in the Bombylidse the third basal cell is open or only closed near the border of the wing, while in all Hybotidae it always remains remote from that border. To the family Hybotidse may be referred : BRACHYSTOMA Meig., HYBOS Fabr., SYNECHES Hal. (= Pterospilus Rond. = Harpamerus Big.), SYNDYAS Loeiv, STENOPROCTUS Loew, ACARTERUS Loeiv, MEGHYPERUS Loew, OEDALEA Meig., EUTHYNEURA Macq. (= An- thalia Zett.), OCYDROMIA Meig., TRICHOPEZA Rond. on account of of its near relation with Brachystoma and LEPTOPEZA Macq. on account of its resemblance with Oeydromia, The two last genera might as well be placed among the Empidse, since they agree with them in having the posterior basal transverse vein parallel to the border of the wing. The 1ST. A. species which I possess belong to the following genera: BRACHYSTOMA Meig., HYBOS Fabr., SYNECHES Walk., SYNDYAS Loew, and LEPTOPEZA Macq. Mr. Walker also describes a species which he believes to belong to the genus OCYDROMIA Meig. FAM. XXIX. EMPIDAB. Charact. — Three basal cells complete, rather large, the third shorter than the second ; posterior basal transverse vein parallel to the border of the wing ; third longitudinal vein frequently furcate ; anterior in- tercalary vein present, the posterior wanting. First joint of the antennae not much shortened, third joint with an apical bristle sometimes resembling a style. Empodium membranaceous and of a linear form. The genera belonging to this family are : EMPIS Meig., PACHY- MERIA Macq., ERIOGASTER Macq. , APLOMERA Macq., RHAMPHO- MYIA Meig., HILARA Meig., RAGAS Walk., GLOMA Meig., MICRO- PHORUS Macq., HORMOPEZA Zett., ITEAPHILA Zett., MICROSANIA Zett. , and CYRTOMA Meig. The X. A. species known to me belong to the genera : EMPIS Meig., PACHYMERIA Macq., RHAMPHOMYIA Meig., HILARA Meig., TACHYDROMID AE. 3 1 and CYRTOMA Meig. Mr. Walker records a N". A. species belong- ing to the genus GLOMA Meig. FAM. XXX. TACHYDROMIDAE. Charact. — The hindmost basal cell not always present, the second basal cell sometimes coalescent with the discal cell ; when present they are of a tolerable size, but the hindmost is always remote from the border of the wing ; third longitudinal vein sometimes furcate ; an- terior intercalary vein sometimes present, the posterior never. First joint of the antennse very much shortened so that the antenna may easily be taken for biarticulate. Empodium membranaceous and of a linear form. To this family belong the genera: HEMERODROMIA Meig. with CHYROMANTIS Rond. and MANTIPEZA Rond., PHYLLODROMIA Zett., TACHYDROMIA Fabr., DRYODROMIA Rond., ELAPHROPEZA Macq., PLATYPALPUS Macq., PHOROXYPHA Rond., SCTODROMIA Hal. ( — Microcera Zett.), ARDOPTERA Macq. (= Leptosceles Hal.), CLINO- CERA Meig., with which HELEODROMIA Hal. (= Paramesia Macq.,) and WIEDEMANNIA Zett. may be properly reunited. As genera occurring in N. A. I enumerate : HEMERODROMIA Meig., TACHYDROMIA Fabr., PLATYPALPUS Macq., ARDOPTERA Macq., and CLINOCERA Meig. If we may trust Mr. Walker's statement, the genus DRAPETIS Meig. also occurs in N. A. Observation. — With the Tachydromidse ends the series of those families of Brachycera which in the greater development of their basal cells differ from the following, and, with the exception of the Lonchopteridse, form a rather natural series, if a linear arrangement may be spoken of as a natural one. The peculiarities, which also characterize this series of families, and any of which, combined with that just mentioned, suffice to place a family under this head, are the following: 1. the third joint of the antennas is composed of a number of joints more or less soldered together ; 2. the third longitudinal vein is furcate ; 3. one intercalary vein or both are present; 4. the empodiurn is considerably developed. These pecu- liarities characterizing the whole series of families hitherto spoken of are much lessened in the Hybotidse and Empidse, and still more so in the Tachydromidse. Tseniaptera, Dacus and other genera with the third basal cell more developed, are placed by most of the authors lower down in the series of families which follow. In 32 DIPTERA OF NORTH AMERICA. order to retain the accustomed arrangement as much as possible, I shall leave them in their usual places, though it would seem, that a more natural arrangement might be obtained, were they added to the above families which have the basal cells prolonged. FAM. XXXI. DOLICHOPODIDAE. Charact. — First basal cell rather short, the second united with the discal cell, the third small ; auxiliary vein running in the first longitudi- nal vein ; third longitudinal vein simple, the fourth sometimes fur- cate ; no intercalary vein. Hypopygium symmetrical, bent under the abdomen. Empodium small, membranaceous, of a linear form. The principal genera are: PSILOPUS Meig., SYBISTROMA Meig , HYPOPHYLLUS Hal,, HERCOSTOMUS Loew, HYGROCELEUTHUS Loeiv, DOLICHOPUS Jfe^.,TACHYTRECHUS Stann., ORTHOCHILEZo^., GYM- NOPTERNTJS Loew, LYRONEURUS Loew, PLAGIONEURUS Loew, LIAN- CALUS Hal., SCELLTJS Loew, HYDROPHORUS Fall, CAMPSICNEMUS Hal, THINOPHILUS Wahlb., PEODES Loew, ACHALCUS Hal., SYS- TENTJS Loew, SYNTORMON Loew, SYNARTHRUS Loew, PORPHYROPS Meig., RHAPHIUM Meig., XIPHANDRIUM Hal., HALTERICERUS Rond., SMILIOTTJS Loew (= Machserium Hal.), ARGYRA Macq., LEUCOSTOLA Hal., NEMATOPROCTUS Loeiv, SAUCROPUS Loew, XAN- THOCHLORUS Loew, SYMPYCNUS Loew, TEUCHOPHORUS Loew, ANEP- sius Loew, EUTARSUS Hal, DIAPHORUS Meig., CHRYSOTUS Meig., CHRYSOTIMUS Hal, MEDETERTJS Fisch., APHROSYLUS Hal The 1ST. A. genera which I am acquainted with are : PSILOPUS Meig., HYGROCELEUTHUS Loew, DOLICHOPUS Meig., TACHYTRECHUS Sfann., GYMNOPTERNUS Loeiv, PLAGIONEURUS Loew, LIANCALUS Hal., SCELLUS Loeiv, CAMPSICNEMUS Hal., SYNARTHRUS Loew, PORPHYROPS Meig., ARGYRA Macq., LEUCOSTOLA Hal, SAUCROPUS Loew, XANTHOCHLORUS Loew, DIAPHORUS Meig., LYRONEURUS Loew, CHRYSOTUS Meig. , and MEDETERUS Fisch. FAM. XXXII. OESTRIDAE. Charact. — Antenna inserted in rounded pits ; the middle part of the face exceedingly narrow ; the opening of the mouth very small ; the oral organs rudimentary. Tegulfe large. This family has often been considered as very distant from the fol- lowing, but the late discoveries have brought to light forms which DEXIDAE. 33 are more nearly related to them. The following genera may be taken for those which constitute the family: TRYPODERMA Wied. (= Cuterelra Clark), CEPHALOMYIA Latr., CEPHENEMYIA Latr., HYPODERMA Clark, GASTRUS Meiy., AULACEPHALA Macg. and CTENOSTYLUM Macq. A thorough limitation of these genera is still wanted, and the name of Oestrus, instead of being dropped, as we see it done by some authors, may perhaps be again restored to its former rank. I have seen N. A. species of the genus TRYPODERMA, and others of the genera CEPHALOMYIA and GASTRUS, introduced in America from Europe. There is no doubt that species of HYPODERMA occur there also. FAM. XXXIII. DEXIDAE. Charact. — Bristle of the antennae hairy or pectinated. Thorax short. First posterior cell of the wing slightly opened, sometimes closed. Tegulae large. Legs long. The family Dexidse agrees with the Tachinidee, Sarcophagidse, Muscidse, and Anthomyidse, in having the tegulse larger than any of the following families of the Brachycera. These five families have been therefore united under the name of Muscarise calyptratse, and contrasted with the following, called Muscarise acalyptratse. There is no possibility, it seems, to discover any other constant character ; that which appears the most serviceable was pointed out to me by Mr. Haliday; it is the transverse suture of the thorax being usually of the same depth on its whole extent in the Muscarise calyptratse, whereas in the Muscarise acalyptratse it is generally distinct at each side and imperceptible on the middle of the thorax. But as some families among the so-called Musca- rise acalyptratse have the tegul«3 so well developed as to resemble those of many Anthomyidx, a high importance cannot be attached to that subdivision. Should it be maintained, the Oestridse ought to be placed among the Muscarise calyptratse. The four families : Dexidse, Tacldnidse, Sarcophagidse, and Mus- cidse, agree in the first posterior cell being very much narrowed or closed at the end, and differ in this from the family Anthomyidse. The former have, for this reason, been comprised under the common name of Creophilse, in opposition to the latter, which 3 34 DIPTERA OF NORTH AMERICA. received the name of Anthophilse. The differences in the organi- zation of the Muscarise calyptratse are much smaller than those of any two families among the first series of Diptera brachycera, which ends with the Tachydromidse, with the sole exception of the group formed by the Hybotidae, Empidse, and Tachydromidse, in which a similar relationship exists. Consequently the families in question here owe their existence much more to the immense number of species and genera than to a real necessity, based on differences of structural characters. Hence it is much more diffi- cult to define their limits, and one must already be well acquainted with a great number of forms, in order to attempt to point out with certainty the right place for new ones. In the limitation of these families I have made use of what has been said about them in Walker's British Diptera; for however insufficient I may find it, I know of nothing better to be pnt in its place. In the family of Dexidse a number of genera have been already formed; as they still require considerable sifting and a much sharper limitation than they have at present, it seems useless to enumerate them here. The N. A. Dexidse known to me cannot all be placed in the genera hitherto erected. The species about whose position there is no doubt belong to the genera: PROSENA St. Farg., MICROPH- THALMA Macg., DINERA Rob. Desv., and ESTHERIA Rob. Desv. There is also no doubt about TRICHODURA Macq. and MEGAPROSO- PUS Macq. occurring in N. A. FAM. XXXIV. TACHINIDAE. Charact. — Bristle of the antennae bare or with a very short pubescence. Thorax short. First posterior cell closed or only slightly opened. Legs short. The immense extent of this family renders the formation of sec- tions indispensable. It is best divided into four sections, which might perhaps be raised into families. The two first of them are the Tachinina and Ocypterina, both of which differ from the two last by their abdomen being beset with long bristles. All TACHININA have an oval abdomen, or when it is nearly cylindrical, its first segment is much shortened. The abdomen of the OCYPTERINA is always of a slender cylindrical form, and its first segment elongated. SARCOPHAGIDAE — MUSCIDAE. 35 V The third section, the GYMNOSOMINA, has a broad front ?nd a vaulted abdomen. The fourth is that of the PHASINA, having a very narrow front and a flat abdomen. My knowledge of the N. A. Tachinidae is a very incomplete one. I know, however, the following genera: I. Tachinina: 1. DEJEANIA Hob. Desv., 2. ECHINOMYIA Meig., 3. JURINEA Rob. Desv., 4. HYSTRICIA Macq., 5. MICROPALPUS Macq., 6. GONIA Meig., 7. NEMOR-ffiA Rob. Desv., 8. BLSPHARIPEZA Macq., 9. BELVOISIA Rob. Desv., 10. TACHINA Meig., 11. CHRYSOSOMA Macq., 12. METOPIA Meig., 13. MILTOGRAMMA Meig., 14. ILLIGERA Rob. Desv., 15. MASICERA Macq. II. Ocypterina: 16. OCYPTERA Lair. III. Gym- nosomina: 17. GYMNOSOMA Fall. IV. Phasina: IS. PKASIA. Latr. 19. HYALOMYIA Macq., 20. TRICHOPODA Latr. FAM. XXXY. SARCOPHAGIDAE. Charact. — Bristle of the antennae plumose or hairy, with the apex bare. First posterior cell only slightly opened or else closed. Tegulffi large. Legs stout. All the N". A. species I have seen belong to the genera : SARCO- PHAGA Meig., PHRYSSOPODA Rob. Desv., and CYNOMYIA Meig. FAM. XXXVI. MUSCIDAE. Charact. — Bristle of the antennse entirely plumose or pectinated. Body never slender; thorax short. First posterior cell only slightly opened or else closed at the border of the wing. Tegulae large. Legs stout. This family contains two sections : the MUSCINA with plumose antennae, and the STOMOXYNA with pectinated antenna. The N. A. species which I have examined belong to the genera: MUSCA Linn., POLLENIA Rob. Desv., CYRTONEURA Macq., PYREL- LIA Rob. Desv., LUCILIA Rob. Desv., CALLIPHORA Macq., and STOMOXYS Geoffr. The number of species which N. A. has in common with Europe is exceedingly striking in this particular family. 36 DIPTERA OF NORTH AMERICA. FAM. XXXVII. ANTHOMYIDAE. Charact. — Thorax with a complete transverse suture. Fourth longitudi- nal vein straight or nearly so, hence first posterior cell fully open. Tegulse rather well developed, though in many cases of no large size. The riches of the N. A. Fauna in this family have been very little explored. I know species of the following genera only : ANTHOMYIA Meig., HOMALOMYIA Bouch., HYLEMYIA Macq., ARI- CIA Rob. Desv., LISPE Lair., and CCENOSIA Meig. The notices of Mr. Walker about the occurrence of some species of Eriphia and of one Dialyta appear to me very uncertain. FAM. XXXVIII. CORDYLTJRIDAE. Charact. — Neuration of the wings complete ; both posterior basal cells of considerable size ; auxiliary vein well separated from the first lon- gitudinal vein ; first longitudinal vein bare. Whole lateral border of the front bristly; anterior border of the mouth with strong, usually numerous vibrissae. Tibise with spurs. With the CordyluridaB we begin that division of Diptera which is called acalyptratse, and the systematical arrangement of which is still and will be an unsolved problem, till their structure has been much more thoroughly studied than has been hitherto the case. In the present state of our knowledge their subdivision into a greater number of families seems to be the most advisable course to pursue. As for their exterior, the Cordyluridse mostly approach to the Anthomyidve, and namely to the species of the genus Coenosia, but the smaller size of their tegulas and the less incomplete develop- ment of the transverse suture on their thorax serve to distinguish them. On the other side they are closely allied to the Helomyzidse, in which, however, the front bears bristles on its upper half only, the two posterior basal cells are smaller, and the costa of the wings is always bristly. N. A. possesses species of CORDYLURA, some of them very inter- esting, and a number of SCATOPHAG^E among which several coin- cide with European species. HELOMYZIDAE — SCTOMYZIDAE. 37 FAM. XXXIX. HELOMYZIDAE. Charact. — Neuration of the wings complete ; costa bristly; first longitudi- nal vein not abbreviated, but bare ; the auxiliary vein is often rather approximated to it. Front bristly on its upper half only ; a stout bristle at each side of the anterior border of the mouth. All the tibiae with spurs and outwards before their tips with a more or less developed erect bristle. The close relation of the Helomyzidse to the Gordyluridse in- duces me to assign them a place here, although the consideration of the smaller size of their two posterior basal cells would remove them to a more distant place, in the neighborhood of the Geomy- zidse and Heteroneuridse. In fact both families are related to the Helomyzidse. ; but they differ from them by their having the first longitudinal vein abbreviated and the auxiliary vein lying close by it, and besides the Heteroneuridse have the peculiarity of the costa of the wings being without bristles. The known N. A. species belong to the genera HELOMYZA and SCIKENOMYZA. Some of them are likewise identical with European species. , FAM. XL. SCIOMYZIDAE. Charact. — Neuration of the wings complete ; two posterior basal cells of rather considerable size ; auxiliary vein well separated from the first longitudinal vein. On the lateral border of the front before the vertical bristles there are two bristles, one behind the other ; face proportionately long without distinct furrows for the antennae; bor- der of the mouth sharp, without vibrissje. Middle tibiae with a greater number of bristles at the tip ; all the tibiae on the outside before the tip with a small upright bristle. I know N. A. species that belong to the genera SEPEDON, TETA- NOCERA, and SCIOMYZA. Some of them are most nearly related to European species, others seem altogether identical with them. If we place, and we may well justify our doing so, the genus DRYO- MYZA among the Sciomyzidse, it must also be named as a genus represented in N. A. ; one of the two species of this genus occur- ring there does not seem to differ from the European Dryomyza anilis Fall. The genus ACTORA Meig., which agrees with the Sciomyzidse in many characters, may be referred to them without any great difficulty; but on account of its deviation in the struc- 38 DIPTERA OF NORTH AMERICA. ture of the face, the character of the family would have then to be slightly altered. Mr. Walker has described a N. A. species which he contends to belong to ACTORA. FAM. XLI. PSILIDAE. Char act. — Body elongated, with short hairs and almost without bristles. Neuration of the wings complete ; the auxiliary vein lies close by the first longitudinal vein, but diverges from it at its end and runs towards the border of the wing ; by a transverse fold most charac- teristic ill this family running from the tip of the auxiliary vein as far as the base of the third posterior cell, the outward end of the auxiliary vein is obliterated ; the posterior basal cells are very large. Front with only a few bristles in the neighborhood of the crown ; face receding ; opening of the mouth small and with no bristles at its border. Only the middle tibiae have spurs, and all the tibia? are without erect bristle on the outside. This family is represented in N". A. by the genera LOXOCERA, PSILA, and CHYLIZA. The N. A. species, which induced Mr. Walker to form a new genus Prochyliza, placed by him close by CJiyliza, belongs to some other family. FAM. XLII. MICROPEZIDAE. Charact. — Body slender, elongated, with very short hairs and very scarce bristles. Legs proportionately short ; only the middle tibiaa have spurs, these being generally very small and weak ; no small erect bristle on the exterior side of the tibise. Neuration of the wings complete ; first longitudinal vein bare ; the auxiliary vein is very close by it and diverges from it towards its end only ; the two posterior basal cells are very large. Front with some bristles in the neighborhood of the crown only ; bordering of the mouth without vibrissse. Last segment of the abdomen of the female prolonged into a blunt, cylindrical tube. The family Micropezidse comprises genera which differ among each other, both in the form of the head and the structure of the antennas and oral parts. The head is sometimes rounded, some- times more elongated ; the bristle of the antennas is generally dorsal, but in some genera apical ; the clypeus sometimes very much de- veloped, sometimes only rudimentary; the palpi sometimes large, sometimes small, but never rudimentary. The clypeus being very much developed and the proboscis very much thickened in the ORTALIDAE. 39 Tseniaptera Macq. , this family approaches so much to the Ortalidse, that the bareness of the first longitudinal vein, the difference in the structure of the female ovipositor, and the corre- sponding difference in the structure of the male appendages, must be considered as the chief characters, which distinguish it from them. The Sepsidee, too, are rather nearly related to the Micro- pezidse, but are distinguished from them not only by the structure of the female ovipositor, but also by their palpi being always rudi- mentary. The N. A. species I know of are: one true CALOBATA, nume- rous T^NIAPTER^E, and two MICROPEZ^E. Whether the N. A. species which Mr. R. Desvoidy refers to the genus NERIUS, really belong to it, appears to me most doubtful, as they seem to be Tceniapterce all together. The genus LISSA Meig. occurring like- wise in N. A., in most characters agrees with the Micropezidte, and may provisionally be placed among them, till a more convenient place in the system will be pointed out for it. The genus EUME- TOPIA erected by Mr. Macquart on a N. A. species, is also related to the latter, and may likewise obtain here a provisional place. Both these genera differ from the great bulk of the Micropezidce by having the legs less slender, the tarsi less abbreviated, and the last segment of the abdomen not prolonged so as to form a cylin- drical tube. FAM. XLIII. ORTALIDAE. Charact. — Neuration of the wings complete ; auxiliary vein separated from the first longitudinal vein and running to the border of the wing in tlie usual way, under an acute angle and remaining perfectly distinct in its whole length ; third longitudinal vein generally with coarse hairs ; two posterior basal cells large, and the outward one frequently prolonged in an acute angle. Front with bristles on the upper part only ; no vibrissse at the border of the mouth ; clypeus commonly very much developed, and proboscis often very much thickened. Middle tibire alone with spurs ; no tibise with an erect bristle on the exterior side before the tip. Ovipositor of the female rather flattened and horny, consisting of three elongated segments, forming three drawers like those of a telescope, and end- ing in a simple point. The family of Ortalidce is exceedingly rich in variously shaped organizations, which caused a considerable increase of genera in 40 D1PTERA OF NORTH AMERICA. it. Unfortunately most of them are founded on characters so va- riable that they are of very little use, and it seems best to retain the old ample genera. We feel the more compelled to do so, as many species existing in N. A. cannot be referred to any of the modern genera. The whole of the Ortalidce may conveniently be divided into two sections : the first, which may be named TETANOPINA, has the front more prominent, the face receding, the opening of the mouth rather small, the clypeus less developed, and the proboscis less thick ; in the second, the ORTALINA, the front is not prominent, the clypeus very much developed, the opening of the mouth much wider, and the proboscis much thicker. The N. A. Fauna possesses in PYRGOTA a genus of the first sec- tion, particularly striking, and even somewhat deviating. Among the other N. A. Ortalidce known to me there is only one species belonging to CEPHALIA, whereas all the rest belong to the genus ORTALIS, if we take it, as Meigen did, in a wider sense, and are distributed especially among the genera Geroxys, Ortalis, Rivellia and Delphinia, of modern authors. FAM. XLIY. TRYPETIDAE. Charact. — Neuration complete ; the end of the auxiliary vein runs steeply to the border of the wing and becomes obsolete ; first longitudinal vein always with bristles, the third frequently, the fifth sometimes ; two posterior basal cells rather large, the hindmost is often prolonged to a point. Front on each side with two rows of bristles, one of which is more above and interiorly, the other below and exteriorly. Border of the mouth with no vibrissse. Clypeus none or rudimen- tary. Proboscis never incrassated. Only the middle tibiae with spurs ; all tibiae without erect bristle on the outer side before the tip. Ovipositor horny, consisting of three elongated retractile segments like the drawers of a telescope, the last of which ends in a simple point. They are divided into two sections, DACTNA and TRYPETINA. In the former the female abdomen, before the ovipositor, has ap- parently only four segments, the fifth segment being diminutive and entirely concealed under the fifth ; in the Trypetina the five segments are all equally developed. As the Dacina, 'represented in Europe only by Dacus Olece, LONCHAEIDAE — SAPROMYZIDAE. 41 which lives on the olive-tree, and Petalopliora capitata, exclusively dependent on the lemon-tree, are in all respects strangers in the European Fauna, so they appear to be no natives of N. A.; no species of this division has hitherto been noticed there. The Try. petina, on the contrary, are represented there by numerous, partly very handsome species, all belonging to the genus TRYPETA in Meigen's and Wiedemann's sense. FAM. XLV. LONCHAEIDAE. Charact. — Neuration complete ; the auxiliary vein runs to the border of the wing in the usual way, under an acute angle and without be- coming obsolete, and is very near to the first longitudinal vein ; this vein is bare ; the two posterior basal cells are small. Front at each side with a single row of bristles ; border of the mouth with- out vibrissse ; clypeus rudimentary. Middle tibiae with spurs ; all tibiae without erect bristle on the exterior side before the tip. The ovipositor of the female consists of three joints and is rather horny, quite flattened, and ends in a simple point. They are divided 'into the PALLOPTERINA having more slender legs and a broader front, and the LONCH^INA with stouter legs and a more narrow front. There is only one species in N. A., that I know of, belonging to the genus PALLOPTERA : of the second division I have several species of LONCH^EA, a part of which seem to be identical with European species. FAM. XLVI. SAPROMYZIDAE. Charact. — Neuration complete ; auxiliary vein of the usual structure, fre- quently very much approximated to the first longitudinal vein ; costa of the wings without bristles or marginal spine ; longitudinal veins without peculiar hairs ; posterior basal cells small. Front with a single row of bristles on each side ; no vibrissse on the border of the mouth ; clypeus rather rudimentary. Only the middle tibiae have terminal spurs ; all tibiae with a small erect bristle on the exterior side before the end. Ovipositor of the female not horny. IS". A. has numerous species of the genera SAPROMYZA and LAUXANJA, and a few species belonging to PACHYCERINA Jlfacq., a genus detached from Lauxania. 42 DIPTERA OF NORTH AMERICA. FAM. XLVII. PHYCODROMIDAE. Charact. — Thorax, scutellum and abdomen flat ; pleura excised above the coxae. Front bristly ; border of the mouth hairy, with no distinct vibrissae. Legs stout, tibiae with spurs and each with an erect hair or small bristle on the outside before the tip ; the first joint of the posterior tarsi not abbreviated ; last joint of all tarsi enlarged, with stout claws and long pulvilli. Neuration of the wings complete ; auxiliary vein distinct in its whole length ; costa without bristles ; basal cells not small. A certain resemblance with the Borboridce can by no means be overlooked; however, the Phycodromidee are readily distinguished by the completeness of the auxiliary vein, the absence of the vibrissse so remarkable in those, by the first joint of the posterior tarsi not being abbreviated, and by the increased size of the last joint of all tarsi. They appear to have more true relation to the Helomyzida, but from these too they are sufficiently distinguished by the costa of the wings having no bristles and the border of the mouth having close hairs, but no real vibrissoe. Of this family I have seen only one COZLOPA captured in N. A. It was remarkable by the exceedingly strong spines of its legs. FAM. XLYIII. HETEROXEURIDAE. Charact. — Neuration of the wings complete, but the first longitudinal vein rather short, and the auxiliary vein very much approximated to it ; costa without bristles ; basal cells small. Front with long bristles ; border of the mouth with a vibrissa at each side ; clypeus not deve- loped ; palpi broad and proportionately large. Legs, and especially the tarsi, slender ; middle and posterior tarsi with spurs ; all the tibiae without erect bristle on the exterior side before the tips ; claws and pulvilli very small. I know five N. A. species of this family, four of which belong to the genus HETERONEURA Meig. ; the fifth cannot be conveniently placed in any of the genera as yet established. OPOMYZIDAE — SEPSIDAE. 43 FAM. XLIX. OPOMYZIDAE. Charact. — Front with stout bristles above ; clypeus rudimentary ; border of the mouth either pubescent or with long hairs, the foremost of which sometimes forms a distinct vibrissa. Proboscis short ; palpi rather small. Middle tibiae with a distinct, posterior tibias with a very short spur ; the exterior side of the tibiae without erect small bristle before the tip ; claws and pulvilli small. Wings elongated and narrow, with no bristles at the costa; the axillary incision and alulae are either wanting or very diminutive. First longitudinal vein much abbreviated ; the auxiliary vein becomes obsolete before reaching completely the first longitudinal vein ; the latter emits, shortly before its end, towards the costa, a branch, which may be considered as the end of the auxiliary vein ; basal cells small. No species belonging to this family has as yet been noticed in X. A. FAM. L. SEPSIDAE. Charact. — Head rounded ; front bristly; border of the mouth more or less hairy, the foremost hair often imitating a vibrissa ; clypeus rudi- mentary ; proboscis short ; palpi exceedingly small or wanting. Abdomen tapering towards the base. Middle tibiae with distinct spurs ; claws and pulvilli small. Neuration of the wings complete; the auxiliary vein distinctly separated from the first longitudinal vein ; the two posterior basal cells rather large. The most essential character of this family is the rudimentary condition of the palpi. With this exception its characters are rather similar to those of the Micropezidce. The genus Cephalia approaches very much the Sepsidce in structure, but its incrassated proboscis, its large and broad palpi, and its considerably developed clypeus prevent it from being reunited with them ; it must, there- fore, remain among the Ortalidce. The species of Sepsidce occurring in N. A. belong to the genera NEMOPODA and SEPSIS, and are, in part, identical with European species. 44 DIPTERA OF NORTH AMERICA. FAM. LI. DIOPSFDAE. Charact. — Neuration of the wings incomplete from the absence of the foremost of the two small basal cells ; the auxiliary vein very much approximated to the first longitudinal vein. Head prolonged in two lateral apophyses bearing the eyes ; front bristly only on the upper part ; border of the rnouth with no vibrissse. Anterior femora incrassated. One species, or — if the second one, described by Dr. A. Fitch, shpuld really prove different — two species of the genus SPHYRACE- PHALA Say have hitherto been found in N. A. FAM. LIT. PIOPHILIDAE. Charact. — The auxiliary vein, on its whole length, is coalescent with the first longitudinal vein ; with this exception the neuration of the wings is complete. Front with some small bristles above only ; bor- der of the mouth with a vibrissa on each side ; clypeus rudimentary; legs rather stoat, almost of the structure of those of the Sciomyzidce ; middle tibiae with spurs ; all the tibiae without erect bristle on the exterior side before the tip. The three N. A. species of PIOPHILA which I have seen are quite identical with European ones; a fourth differs so much from all the known PiopMlce in the form of the head and the structure of the antennas, that it must be considered as the type of a new genus. It seems to be the same species on which Mr. Walker has founded his genus PROCHYLIZA; if that be really so, he would, by assigning it a place immediately by Chyliza, have shown that he had been fully mistaken about its true relation. FAM. LIII. EPHYDRINIDAE. Charact. — Face convex, with no distinct furrows for the reception of the antennae and without vibrissae, though frequently beset with hairs or bristles ; clypeus very much developed ; opening of the mouth large ; proboscis incrassated with a swollen chin. Neuration of the wings incomplete ; the auxiliary vein distinct only at its base ; the fore- most of the two small basal cells reunited with the discal cell. Middle tibiae with spurs. They are divided into three sections : Notiphilina, Hydrellina, and Ephydrina. The Notiphilina are characterized by the second GEOMYZIDAE — DROSOPHILIDAE. 45 joint of the antennse being nnguiculate. The Hydrellina and Epliydrina, in which that joint is not unguiculatecl, differ from each other by the former having the eyes hairy and the latter bare. The N. A. species hitherto recorded have been so badly charac- terized that there is no possibility to decide to which section, and of course far less to which genus they belong. The species which I am acquainted with and have described in the following pages, are distributed among the three above named sections as follows: 1. Notiphilina: 1. DICH^ETA Meig. with two European species; 2. NOTIPHILA Fall, five species ; 3. PARALIMNA Loew, one species ; 4. PSILOPA Fall, five species ; 5. DISCOCERINA Macq. five species. II. Hydrellina: 6. HYDRELLTA Desv. six species; 7. PHILYGRIA Stenh. three species. III. Ephydrina: 8. OCHTHERA Latr. four species, one of which is identical with a European species; 9. BRACHYDEUTERA Loew, one species ; 10. PARYDRA Stenh. two species; 11. EPHYDRA Fall, one species; 12. SCATELLA Desv. three species, one of which cannot be positively distinguished as yet from a European species. FAM. LIV. GEOMYZIDAE. Charact. — Front with stout bristles above ; border of the mouth with vi- brissae. Clypeus rudimentary. Middle tibiae with spurs ; all the tibiae with a small erect hair on the exterior side before the tip. Wings with bristles on the costa ; first longitudinal vein exceed- ingly .abbreviated, and the auxiliary vein so approximated to it that it is distinctly separated from it only towards the base ; the two posterior basal cells very small. I know only one species of this family indigenous in N. A. and belonging to the genus DIASTATA. Mr. Walker records an insect which he believes to be likewise a Diastata. FAM. LV. DROSOPHILIDAE. Charact. — Front with bristles above ; face with distinct sub-antennal fur- rows ; at the border of the mouth there is a feeble, frequently rather indistinct small vibrissa. Middle tibiae with very feeble spurs ; on the exterior side of the tibiae there is either a very small or no erect bristle before the tip. Wings without bristles on the costa; the 46 DIPTERA OF NORTH AMERICA. first longitudinal vein is exceedingly abbreviated ; of the auxiliary vein there is only a rudiment ; the discal cell is usually, but not in all genera, united with the foremost of the two small basal cells. Claws and pulvilli very small. Numerous species of DROSOPHILA are found in N. A., some of which are perfectly identical with European species, and one STE- GANA, the difference of which from the European Stegana hypolenca is at least liable to doubt. FAM. LYI. OSCINIDAE. Charact. — Front without bristles, the crown having only a few short ones ; border of the mouth without vibrissse, which, however, are repre- sented sometimes by a small hair on each side. Middle tibise with small spurs ; all the tibije without erect bristle on the exterior side before the tip. Costa of the wings without bristles. The auxiliary vein is completely wanting ; the anterior of the two small basal cells is united with the discal cell, the posterior one is totally wanting. The N. A. species of this family known to me are distributed among the genera CHLOROPS, CRASSISETA, SIPHONELLA, MEROMYZA, and OSCINIS. The species described by Wiedemann under the name of Homalura plumbella likewise belongs to the genus Sipho- nella. Macquart has established a genus, ECTECEPHALA, on a N. A. species, and he says it is nearly related to Platycephala and Eurina; if that is really the case, it must also be recorded here. FAM. LVII. AGROMYZIDAE. Charact. — Front with strong bristles ; border of the mouth with a vibrissa on each side. Middle tibise with a terminal spur ; all the tibise on the exterior side without erect bristle before the tip. Wings with- out bristles on the costa; first longitudinal vein very short, and the auxiliary vein connected with it at the tip ; basal cells existing, but small; posterior transverse vein generally far distant from the bor- der of the wing. The N. A. species which I have seen belong to the genera AGRO- MYZA, LOBIOPTERA, and MlLICHIA. PHYTOMYZIDAE — ASTEIDAE — BORBORIDAE — PHOR1DAE. 47 FAM. LVIII. PHYTOMYZIDAE. Charact. — Front bristly ; border of the mouth with vibrissse on each side. Middle tibiae with spurs; all the tibiae without erect bristle on the exterior side. Wings without bristles on the costa ; first longitu- dinal vein very short ; auxiliary vein connected with it at the tip ; basal cells existing, but small ; posterior transverse vein wanting. The genus PHYTOMYZA is represented in N. A. FAM. LIX. ASTEIDAE. Charact. — Front bristly above ; border of the mouth with a vibrissa at each side. Middle tibiae with spurs ; all the tibise without erect bristle on the exterior side. Wings without bristles on the costa ; first longitudinal vein exceedingly short ; auxiliary vein connected with it only at the tip; second longitudinal vein very short; two posterior basal cells as well as the posterior transverse vein want- ing. No N. A. species of this small family is as yet known. FAM. LX. BORBORIDAE. Charact. — Thorax, scutellum, and abdomen flat ; front bristly ; face exca- vated, with a vibrissa on each side of the border of the mouth ; clypeus developed; first joint of the posterior tarsi abbreviated. Neuration of the wing incomplete, only a commencement of the auxiliary vein being at best visible ; the hindmost two basal cells are not complete in all genera. N. A. seems to possess numerous species of the genus BORBORUS, which have not yet been carefully compared with the European species. One species taken in Cuba is identical with an African one. FAM. LXI. Charact. — Antennae apparently single jointed, with a long bristle. Wings with several stout veins running into the costa, and three or four weak ones, which run across the surface of the wings and are not completely connected with the hindmost of the stout veins, from which they appear to issue. Femora flattened. Many species of PHORA seem to occur in N. A. ; their form, as far as I am acquainted with them, differs in no way from that of the European ones. 48 DIPTERA OF NORTH AMERICA. III. COEIACEA. FAM. LXII. HIPPOBOSCIDAE. Charact. — Head flattened; first joint of all the tarsi, or at least of the an- terior and middle tarsi, abbreviated. N". A. possesses species of the genera HIPPOBOSCA Linn., MELO- PHAGUS Latr., ORNITHOMYIA Leach, and OLFERSIA Wied., several of which are perfectly identical with European species. FAM. LXIII. NYCTERIBIDAE. Charact. — Head not flattened ; first joint of all the tarsi rather long or very long, in comparison with the following. One STREBLA only and a species belonging to a new genus are known to me as occurring in N. A. II. ON THE NORTH AMERICAN TRYPETIDAE. 1. Extent of the family TRYPETID.E. In stating that the family of Trypetidce comprises the genera Trypeta Meig. and Dacus Wied. we define its limits as exactly as is possible before having developed its character. The genus Trypeta was founded by Meigen in Illiger's Magazine II, 277, 94. Shortly after, the same genus was published in Schrank's Fauna Boica under the name of Trnpanea, and still later, it appeared in Latreille's writings, in a more vague circum- scription, under that of Tephritis. The number of species belonging to it has so much increased since the time of its creation, and so considerable differences in their organization have been observed, that not only the limits of the genus have become a little uncertain, but also the necessity of a division into smaller genera was felt, and more than one attempt to satisfy this want has been made. The first attempt, abortive both from the choice of unfit charac- ters and from the vagueness of the observations used as foundation for the characters, was made by Robineau Desvoidy, who distri- buted the species known to him among the genera Ensina, Stylia, Oxyna, Oxyphora, Terellia, Forellia, Xyphosia, Sitaria, Orettia, Tephritis, Urophora, Aciura, Prionella, Sphenella, Urellia, Acinia, and Nceeta, to which his genera Acidia and Strauzia must be also added. Subsequently Macquart reunited these genera into five : Uropho- ra, Terellia, Tephritis, Acinia, and Ensina, to which he added the genus Ceratitis M'Leay, which he had previously described himself under the name of Petalophora. Later, in the "Dipteres exotiques," he added Acanthoneura, Campylocera, Meracantha, Toxura, and Epicerella; the four last, however, if we may depend on his de- scriptions and figures, must be placed among the Ortalidce. Mr. Walker, in the " List of the Diptera of the British Museum," adopted the genera of R. Desvoidy, after modifying the characters 4 50 DIPTERA OF NORTH AMERICA. of several of them, and retained the genera Anomoia and Euleia, which he had previously founded himself; besides, he erroneously brought again among the Trypetidce the genus Camptoneura, which Macquart had formed on Trypeta picta Wied., and correctly placed among the Ortalidce. The most recent attempt at a detailed classification of the Eu- ropean species of the old genus Trypeta is that given by Rondani in his "Prodromus Dipterologice italicce." He retains — though in a much altered sense — the genera of R. Desvoidy : Oxyna, Uro- phora, Rivellia, Tephritis, Acinia, Aciura, Terellia, and Orellia, and adopting the genus Ceratitis M. Leay and Myopites Breb., he creates the following new genera: Goniglossum, Carpomyia, Gera- jocera, Chetostoma, Epidesmia, Myoleja, SpathuKna, Dithryca, and Oplocheta. But these genera are less fit for reuniting what is really allied, than for isolating out of their nearest relationship such species as are distinguished by any specific peculiarity and for crowding them inordinately together. The dichotomic division of genera from single characters without any indication of the true generic distinctions, renders it impossible to refer to them the other species described by authors, and it is not at all sufficient for this purpose to name a typical species, especially as some of these typical species have not yet been described, and the correct- ness of the names of the others is not proved. Moreover, the characters ascribed by Rondani to the single genera are not all quite certain, and some of them, for instance the scutellum of Myoleja, which is said to have two bristles, the scutellum of Cera- titis six bristles, appear to be errors of the observer. If we add to what we have said already that the genera Xar- nuta, Themara, Calantra, and Aragara, erected by Walker in the "Proceedings of the Linnajan Society," with some probability belong here, and that perhaps the genus Dasyneura Saund., and Rachiptera and Elaphromyia Bigot are Trypetidce, both the va- riety of the forms belonging to the genus Trypeta Meig. and a picture of the chaotic state into which their arrangement has been thrown will be sufficiently illustrated. The genus Dacus, restricted by Meigen to the Dacus Olece Fabr. (the renowned blight of the olive) and used by Wiedemann in a wider sense, is nearest related to the genus Trypeta Meig. ; Fabricius, who formed this genus, comprises so different species in it that we may scarcely consider it as a creation of his. How- DIVISION OF THE FAMILY INTO TRYPETINA AND DACINA. 51 ever striking the difference may be between the greatest part of the species of Trypeta and the larger naked species of Dacus, yet some of the latter approach very much to the larger species of the polymorphous genus Trypeta, and show the near relation of both genera. Wiedemann, misled by some Trypetce, had become uncertain about the limits between the genera Dacus and Trypeta, or he would not have placed the large Brazilian Trypeta parallela among Dacus. One of the surest marks for separating both gen- era is furnished by the structure of the female abdomen, which in Trypeta shows five, in Dacus four segments before the borer, the fifth being very short and concealed under the fourth. None of the other characters, however marked they may appear, is so con- stant as this. Macquart has already justly observed that the whole of the first group of Dacus Wied. is not only a stranger to this genus, but cannot even remain in the same family with it; therefore giving it the generic name of Senopterina (which must be mended into Stenopterina}, he assigned it its right place in the Ortalidce, as will be detailed in the sequel. Among the new gen- era introduced by Macquart, Leptoxys and Enicocera, perhaps also Cardiacera, may be very nearly related to the genus Dacus, which cannot be, however, asserted positively, on account of the insuffi- ciency of Macquart's statements and the incorrectness of his figures. The genus Bactrocera, founded by Guerin, seems also to belong here. The same, perhaps, may be said of the genera Rioxa and Strumeia, formed by Walker in the "Proceedings of the Linnean Society," while the genus Dasyneura of Saunders, which Walker in the "List of the Diptera of the British Museum" places near Dacus, seems to stand much nearer to Trypeta. The species of the genus Trypeta and those smaller genera which either have been comprised in Trypeta or founded in its neigh- borhood, together with the species really belonging to Dacus and the smaller genera subordinate to or co-ordinate with it in a simi- lar way, form the family Trypetida, one of the group of closely related families of the Acalyptera which are characterized by their corneous ovipositor. 2. Division of the family into TRYPETINA and DACINA. A division in two groups may be established as above indi- cated. The two groups would be : Trypetina, with five distinct segments of the female abdomen, and Dacina, with apparently four 52 DIPTERA OF NORTH AMERICA. segments. The latter, moreover, have some peculiarities in the structure and neuration of their wings, which, however, allow of no very sharp limitation. Most frequently a dilatation of the second basal cell and of the space between the third and fourth longitudinal veins (in consequence of which the second longitudinal vein is pushed towards the costal margin) and the posterior angle of the anal cell extended in a long point, are the most striking peculiarities in the structure of the wings. But a sharp limitation of the two groups is perhaps not to be urged too much, as transi- tions from the one to the other are certainly not wanting, and another division quite as useful seems to be possible. In the mean time the groups Trypetina and Dacina, as we have defined them, are characterized as well as our purpose requires. We have now to fix the relation of the family Trypetidce to the nearest families, particularly to that of Ortalidce, since almost all authors have mixed the species of these two families. For this purpose we want only to find out the natural character of the family, which cannot be obtained but from a close examination of the greatest possible number of species and from a careful appreciation of the system- atic value to be assigned to the observed peculiarities and differ- ences. 3. Natural character. After examining nearly 300 species from different parts of the world, I believe I may speak as follows about the organization of the Trypetidce. The bare eyes, in both sexes, are separated by the front, which is of equal breadth or only a little narrowed anteriorly. The mid- dle of the front is not sharply separated from the lateral lists, but has often a different color. The front is even, usually with an almost microscopic, rarely with a longer pubescence, sometimes it is totally bare. On its vertical border it always bears two very strong bristles, rather distant from each other. Two short callo- sities, usually little perceptible, run from them, converging but faintly anteriorly, and bearing one or two bristles directed upwards. On the vertical border itself there are two bristles, each near the upper angle of the eye, and in the middle behind the ocelli there is another pair of bristles, sometimes very stunted. This is also the case with a pair of bristles directed anteriorly, and inserted be- tween the ocelli. More anteriorly on the front there are, at each side NATURAL CHARACTEE. 53 of the orbit, two or three stouter bristles, but generally less stout in the Dacina than in the Trypetina, whereas in the males of some Trypetina they are thickened into spines, or even inserted upon lateral processes. The little crescent cut off by the frontal fissure is often very distinct, though never very large. The antenna? are directed downwards, the third joint elongated or long ; the bristle nearly bare, or with a very short pubescence ; only in a few spe- cies it has longer hairs, but is never pectinated. The face shows below each antenna a flat excavation, more deepened in those few species which have a keel in the middle of the face ; the anterior border of these excavations forms a more or less distinct eleva- tion, and sometimes almost a keel. The cheeks are more or less hairy; in a few Trypetina the furthermost hairs almost have the appearance of vibrissas, which, however, are never present. All more naked species have also less hairy cheeks. The proboscis is never much thickened ; the suctorial flaps are sometimes very short and rather broad, sometimes rather long, sometimes of an extraordinary length, as for instance in the species of Myopites Breb., where they seem to become much stouter, which was the reason why a species of Myopites was placed by Fabricius in Sto- moxys. The prolongation of the suctorial flaps is proportionate to that of the stem ; not unfrequently in most nearly related species the structure of the proboscis seems to be very different ; but on a closer examination this difference appears not to be essential, and cannot even always be used in characterizing the small genera into which the genus Trypeta Meig. has been divided. Among the Dacina I have never seen a species with a very prolonged pro- boscis. The oral cavity is large, sometimes very large and widened, and not seldom a little prolonged at its anterior border ; its form depends very much on that of the proboscis ; for in those species which have a very long proboscis, the anterior border of the mouth is usually also much more projecting. The palpi are either more applied to or more laid upon the labium, when it is retracted into the mouth-hole ; their form is more or less spatulate, and generally more elongated in those species which have a long pro- boscis than in those with a short one. The clypeus owing its origin to a duplication of the skin which connects the stem of the labium with the border of the mouth is narrow, and being con- cealed within the mouth-hole can only be seen anteriorly in those 54 DIPTERA OF NORTH AMERICA. species in which the anterior part of the peristoma is more drawn upwards. There is no striking peculiarity in the structure of the thorax. The transverse suture, in agreement with nearly all the Acalyptera, is distinct in the neighborhood of the lateral border, and totally obsolete in the middle of the thorax. The bristles of the thorax not only offer good specific characters, but sometimes also prove very fit for the separation of genera. In that respect the bristles of the middle of the upper side of the thorax deserve attention ; in those species in which their number is the most complete there are three pairs, one before the suture, the second behind the suture, the third a little before the scutellum. More frequently only the second and third pair of these bristles are present, sometimes only the third ; in almost all African species of Dacus they are all wanting. Besides these bristles of the middle of the thorax there are two rows of bristles on each side; the exterior row consists of four bristles, the first of which stands on the humeral callosity, arid is often wanting in the Dacina; the second has its place before the transverse suture ; the third, which is often much weaker than the others, in the lateral dilatation of the transverse suture ; the fourth above and a little behind the base of the wing. The inte- rior row consists of three bristles only, corresponding to the three last bristles of the exterior row, but is placed a little more back- wards than these. The scutellum, which is more or less convex, generally bears four stout bristles, but in many Dacina and some Trypetina only two; there is sometimes on each side a weaker bristle between the stout ones. The abdomen of the male shows only four distinct segments, the last of which is more or less elongated. The abdomen of the female has five segments before the borer, and the last of them in the Trypetina is always distinct, whereas in the Dacina it is very small, and so concealed under the fourth segment that the abdomen of the female seems to consist only of four segments. That seg- ment which is usually numbered the first, and will also be numbered so in the following descriptions, seems to be composed of two segments soldered together. In many Dacina we also see the next segments more or less completely coalescent on their upper side. The borer of the female is always of a corneous substance ; it is formed of three segments, which are retractile like the drawers of a telescope, and often very long; the last ends in a simple more or NATURAL CHARACTER. 55 less sharp point ; the first segment is either more conical, or more cylindrical, and then usually thick at its base, or it is quite flat; in most species it is hairy, in others it is beset with hairs at the tip only; in others again is quite bare ; its length varies exceedingly in the various species ; the second and the third segments of the borer are always bare. To the length of the borer of the female corresponds that of the thread-like penis of the male. The hairs of the body are sometimes fine and short, sometimes coarse and long ; in the latter case the posterior margin of the abdominal segments is generally beset with bristles, which in the species having fine and short hairs, are either totally wanting or are pre- sent only at the posterior borders of some segments, most frequently on that of the last. The legs are always of moderate length, and of a rather robust structure ; they are beset everywhere with short hairs, which be- come longer on the upper side of the posterior tibia? of some species. There are usually some longer bristles on the under side of the anterior femora, and frequently also on their upper side ; similar bristles exist on the posterior femora of several species, and some- times even on the middle femora. The tips of the middle tibiae are always spurred ; otherwise, the tibite have no bristles. It is very characteristic for all the Trypetidce that the erect bristles are totally wanting, which some allied families, for instance the Sapromyzidce, possess on the outside of the tibia?, not far from the tip. The first joint of the tarsi is always prolonged. The claws and pulvilly are small, and of equal form in both sexes. The neuration of the wings is that of Acalyptera in its highest perfection, and shows many characters peculiar to this family. The auxiliary vein is separated from the first longitudinal vein, though often approximated to it, especially in some Dacina; it never runs in the usual way, that is, at an acute angle and with equal distinctness as far as its end or even incrassatiug towards the border of the wing, but turns suddenly towards it, and, at the same time, becomes much more indistinct, the more so as the space between its end and that of the first longitudinal vein is incrassated. At the place where it runs into the border of the wing, the latter bears a small marginal spine, quite indistinct in many species, and which cannot, therefore, be considered as one of the characters of the Trypetidce. Generally the whole length of the first longitudinal vein is beset with bristles ; this is also most 56 DIPTERA OF NORTH AMERICA. frequently the case with the base, or a greater part, or even the whole length of the third longitudinal vein ; much more seldom there are bristles on the fifth longitudinal vein. The two small basal cells are proportionately large; the posterior of them, i. • 23. interrupta Macq. seems to be an Ortalis related to Herina rufitarsis Macq., if it is not a mere variety of this species, so variable in the color of its body ; moreover, it is so vaguely described that it is not possible to say anything with certainty about it. 24. latipennis Macq., described by Macquart under the name of Platy- stoma latipennis; it is, however, certainly a Trypeta, and I hope not to be mistaken in identifying it with Tryp. sparsa Wied. ; the representation of the head is certainly nothing but the invention of the draughtsman, or a foreign head had been glued to the speci- men. 25. lichtensteinii Wied. I have seen this beautiful species about six- teen years ago, and made a sketch of the picture of the wing, which I give in Tab. II, fig. 25. The bristle of the antenna is thickened at its base in a rather striking manner. 26. longipennis Wied. will be more accurately described in the sequel. The name of it is ascertained from the inspection of the originals. It is surprising that Wiedemann does not mention the thickening of the frontal bristles of the male, though the males in his collection show it. Perhaps he had specimens enough to satisfy himself that this peculiarity is not constant. 27. marginepunctata Macq. is unknown to me. 28. melliginis Fitch belongs to the Ortalidse, and is Herina rufitarsis Macq. 29. mevarna Walk., a Trypeta which has the apex of the wings only reticulated, and is allied probably to the European Tryp. stellata Fiissl. Among the below described species Trypeta Solaris may have the greatest resemblance to it. 30. mexicana Wied. seems to be a Trypeta; the original perhaps exists in the Berlin Museum. It is none of the species known to me. 31. narytia Walk. I believe it also to be a Trypeta; it is likewise not among my species. 32. novaeboracensis Fitch is the same species as Tryp. sparsa Wied. and caliptera Say. 33. nigriventris Macq. probably a Trypeta of the group of Tryp. rotundi- ventris Fall. 34. obliqua Macq., a Trypeta, which seems to be nearly allied to Tryp. suspensa from Cuba and Tryp. unicolor from New Granada, but differs from both by its small transverse vein having an inclined position, and the first hyaline band running uninterruptedly from the border of the wing to the anterior of the two small basal cells, whereas in those species it is interrupted not far from the costal border. 60 DIPTERA OF NORTH AMERICA. 35. obliqua Say seems to be related to the European Tryp. Arctii Deg. and the below described Tryp. palposa from North Wisconsin ; the latter has on its abdomen four rows of black spots, whereas Tryp. obliqua Say has only two. 36. ocresia Walk, apparently related to Tryp. unicolor from New Granada, but it cannot be identified with this or any other species known to me. The description given by Walker is very vague. 37. picta Fabr. a Camptoneura and consequently an Ortalideous species. 38. quadrifasciata Macq. I believe it to be a Trypeta which I do not possess. 39. quadrifasciata Walk, belongs to the Ortalidx and is Herina rufitarsis Macq. 40. quadrivittata Macq. belongs to the Ortalidx. 41. scutellaris Wied. I have seen the typical individuals of this beau- tiful species in the Berlin Museum sixteen years ago. If I recollect right, there were bright bands of a more black than brown color on the two last segments only of the abdomen in the male, but on all segments in the female. I was surprised to see in the female the markings of the wings (Tab. II, fig. 27) more extended than in the male (Tab. II, fig. 26). 42. scutellata Wied. a Trypeta quite unknown to me. 43. septenaria Harr. must be omitted as being undescribed. 44. solidaginis Fitch has been amply described in the sequel. 45. sparsa Wied. I give a detailed description of it. 46. tabellaria Fitch, not among the Trypetx known to me, nor does it seem to belong to them, but is probably an Ortalida. 47. tribulis Ilarr. not described and therefore to be omitted. 48. trimaculata Macq. is the same variety of Tryp. longipennis Wied. which Walker has described under the name of Tryp. cornifera. 49. trifasciata Harr. must be omitted as being undescribed. 50. villosa R. Desv. may be a Trypeta, but is so badly described that there is scarcely a possibility to recognize it. The result of the remarks given about the enumerated 50 species will consequently be as follows : — 1. Five species must be omitted, because they have never been described: albiscutellata Harr., cinctipes Ilarr., septenaria Harr, tribulis Harr., and trifasciata Harr. 2. Fourteen species must be blotted out as identical with species previously described: arcuata Walk., armata R. Desv., caliptera Say, cornigera Walk., cornifera Walk., fimbriata Macq., flavonotata Macq., inermis R. Desv., latipennis Macq., marginepunctata Macq., melliginis Fitch, novcebora- censis Fitch, quadrifasciata Walk., and trimaculata Macq. ON THE SYSTEMATIC ARRANGEMENT OF THE SPECIES. 61 3. Of the remaining thirty-one species seven, the five first with all certainty, the two last with great probability, must be placed among the Ortulidfe. These are : antillarum Macq., flexa Wied., fulvifrons Macq., picta Fabr., quadrivittata Macq., interrupta Macq., tdbellaria Fitch". 4. Consequently twenty-four N. A. Trypetce have been described, namely: acidusa Walk., asteris Harr., avala Walk., beau- voisii R. Desv., comma Wied., culta Wied., dinia Walk., electa Say, fucata Fabr., lichtensteinii Wied., longipennis Wied., mevarna Walk., mexicana Wied., Narytia Walk., nigriventris , Macq. , obliqua Macq., obliqua Say, ocresia Walk., quadrifasciata Macq., scutellaris Wied., scutellata Wried., solidaginis Fitch, sparsa Wied., and villosa R. Desr. 5. Of these twenty-four species I possess four only [now five. — 0. S.], which I shall fully describe hereafter; they are: electa Say, longipennis Wied., solidaginis Fitch, sparsa Wied. [and obliqua Say. — 0. S.]. Moreover I have seen in other col- lections four species ; they are : comma Wied., culta Wied., lichtensteinii Wied., and scutellaris Wied. As an addition to my paper I subjoin the descriptions which Wiedemann has given of them, and accompany them with drawings of the wings, which, however, on account of their being only fugitive sketches, have not the same claim to correctness as the figures of the wings of the other species. 6. On the systematic arrangement of the species to be described. Besides the above mentioned four species I have to describe nineteen new ones, which I leave all united in the genus Trypeta. Though they differ in their organization, I think my course is both reasonable and proper. It has been already sufficiently shown, how very uncertain the limits of the family Trypetidce are. For the immediate purpose, it will be quite sufficient if insects of other families are no longer mixed with these. The number of accurately known species must increase considerably, before a convenient classification can be thought of. The smaller genera hitherto founded on the various forms of the Trypetina are partly formed on European species, partly estab- lished in a very superficial manner on single species of other parts 62 DIPTERA OF NORTH AMERICA. of the globe. Among the former there are some, which are avail- able or may be rendered available by removing the aberrant species from them ; the remaining genera either have no claim to the names of genera, or are understood by different authors in so different a sense as to render their adoption more perplexing than useful. But a few of the available genera are represented in North Ame- rica. The genera created for single species have usually been established on account of a single striking character, no informa- tion being given about the other characters ; so it will be next to impossible to place new species in such genera without incurring the risk of gross mistakes. Such being the case, I will be justified, I think, in comprising all species under the head of Trypeta, in the sense of Meigen and Wiedeinaun. I should be glad indeed if by the communication of numerous species I was enabled to divide the N". A. Trypetina into smaller genera. To obtain numerous species is only possible by breeding them, which is a very easy task ; for the larvae are easily discovered ; they live in stalk-galls, or in berries and berry- like fruits ; but most frequently in the flower-heads of Composites, among which they prefer the Cynarocephalce to all others. To prevent any misunderstanding I finally have to observe, that in the following descriptions, by the length of the borer I always meant the length of its first joint only, which is also comprised in the indication of the length of the females. The length of the whole borer depends so much on the more or less extension of its three segments, that no certain measure of it can be given. Synopsis of the species described in the sequel.* , ( Wings pictured (reticulate or banded). I Wings not pictured, hyaline. 26 0 ( Wings banded. 3 ~ i (. Wings reticulate. 15 „ ( Third longitudinal vein with bristles. 4 ( Third longitudinal vein without bristles. 10 , ( Abdomen black. 1 discolor, n. sp. 1 Abdomen yellow. 5 * If a species is not found among those enumerated in this synopsis, before pronouncing it to be new the Appendices I and II should be con- sulted. This table contains only species described from specimens, and not merely quoted from other works. 0. S. SYNOPSIS OP THE SPECIES DESCRIBED IN THE SEQUEL. 63 , J Face very receding. 2 longipennis Wied. i I Face not receding. 6 /> ( Back of the thorax not striped. 7 ( Back of the thorax striped. It - ( Abdomen with black dots. obliqua Say.* I Abdomen without black dots. 8 ( The band rising over the posterior transverse vein is connected with the preceding on the posterior border. 3 fratria, n. sp. 1 The baud rising over the posterior transverse vein is not connected L with the preceding. 4 suspensa, n. sp. (The two middle bands of the wing diverge towards the posterior border. 5 unicolor, n. sp. 1 The two middle bauds of the wing are converging towards the poste- rior border. 6 electa Say. Thorax and abdomen differing in color. 7 iusecta, n. sp. Thorax and abdomen of the same color. 11 f Color of the body yellow. 12 \ Color of the body black. 14 , 0 f Abdomen with black dots. 8 palposa, n. sp. I Abdomen without black dots. 13 , „ f Basal third of the wing hyaline. vernoniae, n. sp.f ( Basal third of the wing pictured. 9 suavis, n. sp. ,. ( Scutellum yellow. 10 cingulata, n. sp. I Scutellum black. 11 polita, n. sp. , - f Wings much widened. t 16 I Wings not widened. 17 ITip of the wings with an uninterrupted white seam. 12 sparsa Wied. Tip of the wings with an interrupted white seam. 13 rotundipennis, n. sp. , M ( Proboscis geniculated. 18 I Proboscis not geniculated. 19 , „ f Stigma with a limpid drop. 14 clathrata, n. sp. ( Stigma without limpid drop. 15 humilis, n. sp. , „ f Reticulation of the wing not radiating at its tip. 20 I Reticulation of the wing radiating at its tip. 21 20 ( Front exceedingly broad. 16 solidaginis Fitch. ( Front of moderate breadth. 17 seriata, n. sp. 01 ( Wings reticulate only on the apical half. 18 Solaris, n. sp. ( Wings reticulate on their whole surface. 22 22 f The whole reticulation equally broken. 19 sequalis, n. sp. I The reticulation broken much less in the middle. 23 no ( Abdomen yellow. 24 ( Abdomen black. 25 * See Appendix III. f Ibid. 64 DIPTERA OF NORTH AMERICA. 9< f Reticulation paler in the middle. 20 festiva, n. sp. I Reticulation everywhere of the same color. 21 bella, n. sp. f The curvature inside of the first posterior cell considerable. „ ! 22 latifrons, n. sp. j The curvature inside of the first posterior cell very small. 23 melanogastra, n. sp. 26 j Stigma fuscous. albidipennis, n. sp.* 1 Stigma pale. alba, n. sp.f 7. Description of the species. 1. T. discolor LOEW. £ . (Tab. II, fig. 1.) — Lutea, abdomine nigro, alarum fasciis quatuor obliquis fuscanis, prima et secunda antice, tertia et quarta postice connexis, vena longitudinal! tertia setosa, venulis transversis valde approximatis. Luteous yellow with the abdomen black ; wings with four very oblique brownish bands, the two first being anteriorly, two last posteriorly connected ; third longitudinal vein with bristles and the two transverse veins approximate. Long. corp. 0.13. Long. al. 0.15. Yellow with a rather glossy black abdomen, which color becomes more blackish-brown near the base. Front proportionately rather narrow ; three bristles at each side are of a browner color and directed anteriorly. Antennae yellowish, not reaching as far as the border of the mouth ; bristle of the antennae thin, apparently naked. Face descending rather straight downwards, but little excavated. Opening of the mouth proportionately large. Pro- boscis short, palpi somewhat prominent. Upper side of the tho- rax with short yellow hairs and yellowish-brown bristles; it has no pale stripes, but there is an obsolete, paler, longitudinal stripe between it and the pleurae. Pleurae of the color of the upper side of the thorax, with yellowish bristles ; the neighborhood of the coxss is blackish. Scutellum with four bristles. Abdomen with short black hairs. Legs yellow, anterior femora with ochra- ceous bristles on the under side. Wings glassy with four very oblique brown bands, which are partly tinged with brownish- yellow on the inside. The first band begins at the base of the wing, where it is connected with the second, runs over the anal cell as far as the base of the third posterior cell, and, including the fourth longitudinal vein, projects a little, whereupon leaving the latter, it bends towards the posterior border of the wing, which it * See Appendix III. t Ibid. TRYPETIDAE. 65 reaches in the middle between the tips of the fourth and fifth lon- gitudinal veins; its color is yellowish-brown near the base of the wings and dark brown beyond the basal cell. The second band runs first from the base of the wing to near the tip of the first longitudinal vein, then crosses the wing obliquely, on the border of which it includes the tip of the fifth longitudinal vein; the two transverse veins are included by it in such a way that their anterior ends are placed exactly on the outside border of the band ; the color of this band is brownish-yellow with darker brown edges which gradually overcome the lighter color at the posterior end ; also that portion of it which covers the stigma and the space imme- diately beneath it, is dark brown. The third band begins on the costal border immediately behind the stigma and reaches the pos-. terior border immediately behind the tip of the fourth longitudinal vein; it is brownish-yellow, edged with dark brown, the posterior end being likewise dark brown. The fourth band begins a little before the tip of the second longitudinal vein and runs on the bor- der of the wing as far as the] fourth longitudinal vein, where it is connected with the third band; its color is dark brown, being brownish-yellow only at its anterior end. The two transverse veins are very near each other and very steep ; none of the longitudinal veins is extraordinarily arcuated ; the second, third, and fourth longitudinal veins diverge a little at their tips; the bristles of the third longitudinal vein are very distinct and reach as far as its tip. Hah. Cuba. (Riehl.) 2. T. loiigipeimis WIED. £ & £. (Tab. II, fig. 2 £ , 3 £.)— Flava, angusta, rivulis fasciisque alarum augustarum flavo-fuscanis, facie valde recedente. Yellow, slender; the narrow wings with brownish-yellow rivulets and bands; the face much receding. Long. corp. 0.17 — 0.26. Long. al. O.i:2—0.30. SYN. Trypeta longipennis WIEDEMANN, Auss. Zweifl. II, 483, 12. Strauzia armata R. DESVOIDY, Myod. 719, 2. (£ .) Slrauzia inermis R. DESVOIDY, Myod. 718, 1. ( £ .) Tf'phritis trimaculata MACQUART, Dipt. exot. II, 226, 8. Tab. XXXI, fig. 3. Trypeta cornigera WALKER, List. Brit. Mus. IV, 1010. Trypeta cornifera WALKER, List. Brit. Mus. IV, 1011. Very variable both in size and in the color of its body and wings, yet readily recognizable. In the palest individuals the whole body is yellow, only a very small dot immediately above the base of the 5 66 DIPTERA OF NORTH AMERICA. wings and the tip of the borer being black. In darker individuals, the following markings appear successively: 1. A black double spot in the middle of the anterior border of the thorax ; 2. A black spot on each side of the scutellum ; 3. A broad black stripe on each side of the metanotum ; 4. The black posterior portion of the lateral stripes ; 5. The black anterior portion of the lateral stripes ; 6. The double stripe in the middle of the thorax, abruptly ending in its centre. The head is rather bright yellow ; front very prominent, face much receding ; opening of the mouth not widen- ed ; palpi and proboscis short, yellow, the bristle with a very short pubescence. Frontal bristles black, the superior ones considerably stout, two of them on each side in the male assume the shape of straight spines, a little incrassated at their tips ; in smaller males, however, these spines are not seldom only little stouter than in the females, and of the ordinary form. The lateral border of the thorax and the superior border of the pleurae may have a pale yellow color in life ; in dry specimens they are very whitish. From the pale yellow scutellum a broad pale yellow stripe extends to about the middle of the thorax. Hairs and bristles of the thorax black. Scutellum with four black bristles. Abdomen proportionately very narrow, with rather long black hairs. Borer about half as long as the abdomen, with the tip only blackened. Legs yellow, anterior femora with black bristles on the under side. Wings narrow and very long, more elongated and pointed in the males than in the females, but not always in the same degree ; the brown- ish-yellow stripes and bands leave the following clear spots : 1. A space near the costal border between the transverse humeral vein and the tip of the auxiliary vein, and having usually a brownish spot in its middle ; 2. An oval space immediately below the stigma between the third and fourth longitudinal veins ; 3. A tri- angular space immediately beyond the tip of the first longitudinal vein, and reaching from the costal border as far as the third longi- tudinal vein ; 4. An arcuated band running obliquely from the costal border between the two transverse veins as far as the fifth longitudinal vein ; 5. A triangular space on the posterior border filling up the second posterior cell, with the exception of an edge along the veins ; 6. An arcuate oblique sernifascia beginning on the posterior border before the tip of the fourth longitudinal vein, and running as far as the third longitudinal vein ; 7. The alary appendage, the posterior angle of the wing, and a large space TRYPETIDAE. 67 adjoining it, and lying before the last longitudinal vein. The transverse humeral vein, and the space of the costa near it, are usually black as well as the space of the latter, where the costal spine is inserted. The brownish-yellow bands of the wings have narrow brown edges, and are in a greater or less extent brown near the tip and the posterior border of the wings. There are specimens in which the bands are much more extended, but those having a part of them obsolete are more common ; this fading of the picture of the wings is most frequent in the neighborhood of the posterior border. The posterior transverse vein is a little oblique ; the tips of the third and fourth longitudinal veins are curved. The circumference of the wings is not always the same in the males ; those the frontal bristles of which are most thickened appear to have the most prolonged and pointed wings. Hal. Middle States. (Osten-Sacken.) Observation. — I have had an opportunity of examining the typical individuals of Tryp. longipennis Wied. 3. T. fratria LOEW. 9. (Tab. II, fig. 4.) — Tota lutea, thorace non vittato, alarum rivulis fasciisque luteo-fuscanis, maculam ovatam pellu- cidam in posteriore cellulse discoidalis parte iucludentibus, vena longi- tudinal! tertia setosa. Totally luteous yellow ; the thorax without stripes ; the wings with brown- ish-yellow rivulets and bands, including an ovate pellucid spot in the posterior part of the discal cell ; the third longitudinal vein with bris- tles. Long. corp. 0.22. Long. al. 0.22. Rather dark yellow with the scutellum paler and an almost whitish-yellow, not sharply limited stripe, running from the shoulder to the base of the wing ; the metanotum at each side with a dot- like black spot. Front of moderate breadth. Antenna? yellow, little longer than half the face, with the bristle apparently bare. Face receding only a little, and slightly excavated below the an- tennce. Proboscis short ; palpi slightly projecting. Bristles of the thorax black. Hairs of the abdomen short and black. Borer very short, not flattened, concolorous with the abdomen. Legs yellow, tibia? and tarsi paler than the femora; anterior femora with some black bristles on the underside. The picture of the wings is yellowish-brown, and of the same form as that of the European Tryp. Heraclei Linn. The part of it adjacent to the base of the wings reaches from the costal border as far as the dark brownish 68 DIPTERA OF NORTH AMERICA. stigma, having, however, between the transverse humeral vein and the tip of the auxiliary vein a rather large and almost hyaline space ; it includes between the third and fourth longitudinal veins an oval transparent spot near the base of the discal cell ; it covers the whole of the two posterior basal cells and fills up the two first thirds of the discal cell, running then in a darker color behind the fifth longitudinal vein as far as the tip of this vein, from whence forming a band, it rises above the posterior transverse vein and is connected with the remaining picture in the neighborhood of the small transverse vein. From the latter place a band runs obliquely to the costal border, where it seams the tip of the wing and pro- ceeds as far as the tip of the fourth longitudinal vein ; on the third longitudinal vein it emits a parallel branch running to the posterior border. The last portion of the third longitudinal vein is only slightly curved ; the posterior transverse vein is slightly oblique ; the small transverse vein is perpendicular and more than one and a half of its length from the posterior transverse vein. Hal>. United States. (Osten-Sacken.) Observation 1. — Tryp. fratria resembles exceedingly the yellow variety of Tryp. Heraclei Linn., and agrees with it especially in the picture of the wings ; but it differs from it by its shorter and pro- portionately broader wings, by the greater distance between the two transverse veins, and the curve of the last portion of the third longitudinal vein, which is less considerable. I found also the borer of the palest females of Tryp. Heraclei always black, whereas its color in Tryp. fratria. agrees with that of the abdomen. 'Observation 2. — I have been of the opinion for some time that this species might be Tryp. varipennis Macq., but after a closer examination I find this not admissible, since in Macquart's figure (Dipt. exot. II, 3, Tab. XXXI, f. 1) the band rising from the posterior border and seaming the posterior transverse vein includes a large clear space behind the fifth longitudinal vein, which does not exist in Tryp. fratria, and the penultimate band is united with the first near the second longitudinal vein, whereas in Tryp. fratria this union takes place at the third longitudinal vein. The state- ment of the metanotum of Tryp. varipennis being black, whereas in my specimen of Tryp. fratria it is marked with a black dot on each side only, could not be considered as decisive for separating the two species, since the species of this group are very variable in their colors. TRYPETIDAE. 69 4. T. Slispensa LOEW. £ . (Tab. II, fig. 5.) — Tota lutea, thorace non vittato, alarum fasciis fuscanis obliquis, postice divergentibus, vena longitudinal! tertia setosa. Totally luteous yellow ; the thorax not striped ; the wings with oblique brownish bands diverging posteriorly; the third longitudinal vein with bristles. Long. corp. 0.21. Long. al. 0.22—0.23. Dark yellow. Front purer and paler yellow; frontal bristles black, rather short, and not very stout. Antenna yellowish, almost as long as the face, with the bristle very thin, and having a very delicate and short pubescence. Face a little receding, with pro- portionately rather deep furrows for the reception of the antennae. Opening of the mouth rather widened ; border of the mouth sharp. Proboscis rather thick, with the suctorial flaps a little prolonged. Palpi broad. Upper side of the thorax without stripes; its pube- scence yellow and exceedingly short, bristles black. Scutellum with four black bristles. Metanotum colored alike with the rest of the body. Hairs of the abdomen short and pale, but the bris- tles at its end black. Legs yellow ; anterior femora not very stout, with some black bristles on the under side. Wings not very long; their markings are mostly yellowish-brown, and leave the following hyaline spots : 1. A small triangular one on the costal border immediately behind the tip of the first longitudinal vein, reaching as far as the third longitudinal vein, and joining there a hyaline spot which lies below the stigma between the third and fourth longitudinal veins ; 2. An oblique band slightly curved, which rises on the posterior border, near the tip of the last longitudinal vein, and ascends between the transverse veins as high as the third longitudinal vein ; 3. A triangular spot of the posterior border, occupying the greater part of the second posterior cell, and reach- ing with its apex to a little beyond the fourth longitudinal vein ; 4. An oblique band which begins at the posterior border, imme- diately beyond the tip of the fourth longitudinal vein, and ascends as high as the second longitudinal vein, so that the two oblique clear bands almost meet with their anterior ends. The small transverse vein is at the end of the second third of the discal cell, and, like the posterior transverse vein, has a slightly oblique posi- tion ; the end of the fourth longitudinal vein is distinctly curved forwards ; the posterior angle of the anal cell is drawn out into a long point. Hob. Cuba. (Poey.) 70 DIPTERA OF NORTH AMERICA. <5. T. imicolor LOEW. £. (Tab. II, fig. 6.) — Flava, thoracis vittis scutelloque inulto pallidioribus, fasciis alarum fusco-fiavescentibus, pos- tice divergentibus, vena longitudinal! terti&, setosa. Yellow, the stripes of the thorax as well as the scutelluni much paler ; the brownish-yellow bands of the wings diverging posteriorly ; the third longitudinal vein with bristles. Long. corp. 0.26. Long. al. 0.27. Rather pale yellow. Front a little brighter, of moderate breadth ; frontal bristles black, the superior ones rather stout. The yel- lowish antennas are nearly as long as the face, bristle very short, beset with a very short and delicate pubescence. Face a little re- ceding, the furrows for the reception of the antenna? proportion- ately rather deep. Opening of the mouth rather widened, border of the mouth sharp. Proboscis rather thick, with the suctorial flaps slightly prolonged ; palpi broad. The upper side of the thorax, above the base of the wings, shows a stripe running from the suture to the posterior border of the thorax, and has in the described specimen rather a whitish color, which seems to have been pale yellow in the living insect ; of the same color are the shoulder and the space behind it, the scutellum and a large spot above the poisers ; a broad stripe of the same color seems to run from the middle of the posterior border of the thorax to nearly its middle. The dark stripes usual in other species are indicated by rows of blackish spots; they may, however, have become visible only after the drying up of the insect. The short hairs of the thorax are pale yellowish, the bristles black. Scutellum with four black bristles. Metanotum with a black stripe on each side. Abdomen with pale, very short hairs and brownish-black bristles at its end ; last segment a little prolonged, with an indistinct brown longitu- dinal line on each side. Legs yellow; anterior femora with some brown bristles on their under side. Wings rather large; the bands are brownish-yellow with brown edges, entirely brown near the posterior border and the tip of the wing; the clear spaces which they leave are as follows : 1. A very oblique one, interrupted on the third longitudinal vein, with its anterior end forming a trian- gular spot, placed beyond the tip of the first longitudinal vein, and running through the base of the discal cell as far as the base of the posterior basal cell ; 2. A band, having the form of an S, rising on the posterior border, near the tip of the last longitudinal vein, and, after running between the transverse veins, ascending as high as the second longitudinal vein, from whence it turns again TRYPETIDAE. 7 1 to the posterior border, which it joins iu the neighborhood of the tip of the fourth longitudinal vein ; 3. A large triangular spot of the posterior border, which occupies a great portion of the second posterior cell, and with its tip reaches beyond^the fourth longi- tudinal vein. The stigma is long and rather dark brownish. Transverse veins straight and steep ; the fourth longitudinal vein is distinctly curved forwards at its end, as in Tryp. paraUela Wied.; the posterior angle of the anal cell is drawn out into a large point. Hob. New Granada. (Schott.) 6. T. electa SAY. 9. (Tab. II, fig. 7.) — Flava, thoracis vittis scutel- loque multo pallidioribus, alls fuscario-fasciatis, vena longitudinal! tertia setosa, tibiis posticis nigro-ciliatis. Yellow, the stripes of the thorax as well as the scutellum much paler ; the wings with straight brownish bands ; the third longitudinal vein with bristles and the upper side of the posterior tibiae ciliated with black bristles. Long. corp. 0.29. Long. al. 0.29. SYN. Tnjpeta electa SAY, Journ. Acad. Philad. VI, 185, 1. Tephritis flavonotata MACQUART, Dipt. exot. Suppl. V, 125. Tab. VII, fig. 9. This very distinct species has in the picture of the wings a great resemblance with the European Tryp. alternata Fall. Yel- low. Front of middling breadth ; frontal bristles black, rather stout. Face slightly receding, with rather deep furrows for the reception of the antennas. Antenna? yellow, reaching only a little beyond the middle of the face ; the bristle bare and very thin. Opening of the mouth large, but not widened ; border of the mouth rather sharp, but not projecting. Proboscis small ; palpi rather broad. On the upper side of the thorax there is a pale yellow (almost ivory color in the dry specimen) stripe running from the shoulder to the base of the wing, and a second above the base of the wing running from the suture as far as the posterior border of the thorax. Another stripe of the same color runs from the middle of the posterior border to beyond the middle of the thorax, where it is gradually pointed and obliterated; the stripe lying above the base of the wing is interiorly edged with black in the described specimen. Hairs of the thorax short, pale yellowish ; bristles black. Scutellum of the color of the pale stripes of the thorax and a black spot on each side of the base of the lateral border ; it has four bristles. Pleura with a pale yellow longitudinal stripe in its middle and two black little spots above it. Abdomen yellow, 72 DIPTERA OP NORTH AMERICA. with short black hairs and black bristles on the posterior borders of the two last segments ; the last segment has a dot-like black spot on each side near the base. Borer short, thick, not flattened at all, a little rounded at its end, of the color of the abdomen, and covered with black hairs. Legs yellowish ; anterior femora with black bristles on their under side ; the under side of the middle and posterior femora with some black bristles only near the tip, the bristles being shorter on the middle femora; the pos- terior tibiae on their whole upper side are densely fringed with rather long black bristles, which afford a very characteristic mark of this species. Wings hyaline, with brown bands. The first of these bands is the least regular, and runs from the transverse humeral vein as far as the end of the anal cell, the posterior angle of which is drawn out into a long point ; this anal cell, as well as the basal cell lying before it, is brownish-yellow. The second band, beginning with the short stigma, runs over the small trans- verse vein, and, after crossing the discal cell, reaches the posterior border, on which it is connected more or less distinctly with the posterior end of the following band. The latter begins on the costal border before the tip of the second longitudinal vein, and after running over the posterior transverse vein in a straight direc- tion, reaches the posterior border of the wing. Its anterior end is perfectly connected with the last band, which seams the wing as far as a little beyond the tip of the fourth longitudinal vein. Be- vween the second and third bands there is still a yellowish-brown line drawn perpendicularly from the costal border to the third longitudinal vein. Transverse veins straight; the small transverse vein is a little beyond the middle of the discal cell. Hab. Florida. (Osten-Sacken.) 7. T. insecta LOEW. £. (Tab. II, fig. 8.) — Thorace nigro, capite, abdomine pedibusque luteis, alarum nigrarum incisuris marginalibus guttulisque inter venarum longitudinaliurn tertiam et quartam tribus vel quatuor pellucidis, vena longitudinal! tertia nuda, setis scutelli duabus. Thorax black ; head, abdomen, and legs luteous ; wings black, with limpid incisions on the borders, and three or four limpid drops between the third and fourth longitudinal veins ; third longitudinal vein bare ; scutellum with two bristles. Long. corp. 0.14. Long. al. 0.14. Of the group of the European Tryp. rotundiventris Fall. Head dirty yellow. Front rather narrow, more so towards its anterior TRYPETIDAE. 73 end ; frontal bristles brown ; the row of rather long bristles at the posterior orbit whitish. Antennas yellowish, nearly as long as the face. Cheeks descending only a little below the eyes. Thorax and scutelluru black, the neighborhood of the coxa3 brown. Hairs of the thorax short, pale yellowish, bristles brown. Scutel- lum with two long brown bristles. Abdomen brownish-yellow ; borer not quite so long as the two last segments united, much broader at the base, broad and abrupt at the end, flattened, cou- colorous with the abdomen. Legs yellow. Wings black, with pellucid spots. On the costal border there is a small spot before the transverse humeral vein ; a similar spot projecting a little from the first longitudinal vein lies beyond the transverse humeral vein, and a smaller one between it and the tip of the auxiliary vein ; it is followed by two triangular spots of equal size, the first of which is immediately beyond the tip of the first longitudinal vein, and touches the third longitudinal vein at its junction with the small transverse vein. On the posterior border of the wing there are six clear excisions, the two first of which coalesce with the grayish hyaline axillary angle of the wing ; the third reaches the fifth longitudinal vein ; the fourth lying behind the tip of the fifth longitudinal vein is more cloudy than the others, and goes a little beyond the fifth longitudinal vein ; the fifth accompanies the steep posterior transverse vein at its hind side, and reaches as far as the fourth longitudinal vein ; the sixth has a more inclined position and a sharper tip, with which it reaches the fifth longi- tudinal vein. In the middle of the broad first posterior cell there is a considerable hyaline drop ; a much smaller drop is seen at the anterior side of the fourth longitudinal vein, in the middle of its penultimate portion, and one or two hardly visible drops at the antepenultimate portion of this vein. The second longitudinal vein is slightly undulating, and diverges more than usually towards the end from the third longitudinal vein ; the posterior angle of the anal cell is drawn out into a prolonged point. Hob. Cuba. (Poey.) 74 DIPTERA OF NORTH AMERICA. S. T. palposa LOEW. £ • (Tab. II, fig. 9.) — Flava, abdomine punc- torum nigrorum seriebus quatuor picto, alls hyalinis luteo-fasciatis, vena longitudinal! tertia nuda. Yellow, abdomen with four rows of black dots, wings hyaline with luteous bands and the third longitudinal vein bare. Long. corp. 0.26 — 0.27. Long. al. 0.26. It is nearly allied to Tryp. arctii Deg., lappce Cederli, etc. Yellow; front brighter yellow, rather broad; frontal bristles black- ish, the small bristles of the posterior orbit whitish. Antennas yellow, descending a little beyond the middle of the face, their third joint rather broad ; the bristle of the antennas apparently bare, with the base slightly incrassated. Face a little receding, excavated in the middle, so that the border of the mouth is pro- jecting ; the furrows for the reception of the antennae rather flat. Opening of the mouth very wide; proboscis thick and short; palpi broad, rather large, with some small black bristles. Thorax with a large glossy black spot in the middle of its anterior side ; its upper side blackish, with the exception of the lateral and posterior borders, and of a large triangular spot which rises from the poste- rior border ; hairs whitish-yellow ; bristles blackish ; two of the latter, inserted before the scutellum, are placed on larger black dots, and two before them on smaller clots. Scutellum with four bristles, yellow, only a little blackened at the base of the lateral border. Metanotum black. Pleura? with some brown spots, and above the posterior coxas with a small black spot. The third, fourth, and fifth segments of the abdomen have each at its anterior border four small black spots, the intermediate ones being more approximated; the fifth segment is much prolonged, and has, moreover, a black spot in each posterior corner. The rather coarse hairs of the abdomen are yellowish on the first segments and the anterior part of the middle ones, the remaining ones are black. Legs yellow; anterior femora with black bristles on the under side. The color of the picture of the wings is brownish-yellow in my specimen, which is apparently a little faded ; it extends on the costal border from the base to the tip of the first longitudinal vein, and, on the first portion of this extent, reaches as far as the fourth longitudinal vein, and on the second half as far as the third only. From the tip of the first longitudinal vein a band runs over the small transverse vein ; a second band runs from the costal border over the posterior transverse vein ; these two bands become more TRYPETIDAE. 75 obsolete towards the posterior border, and almost coalesce in its neighborhood. The first of them is also connected with a stripe which edges the fifth longitudinal vein. The edge of the tip of the wing is perfectly connected with the second band, and reaches a little beyond the tip of the fourth longitudinal vein. The anal cell is brownish-yellow. Transverse veins steep ; the small trans- verse vein a little before the last third of the discal cell. Hob. Northern Wisconsin. (Kennicott.) 9. T. SURTis LOEW. $ . (Tab. II, fig. 10.) — Pallide flava, unicolor, alarum hyalinarum litura basalt fasciisque tribus nigricantibus in formam literse S confluentibus, vena longitudinal! tertia nuda. Pale yellow, unicolorous ; wings hyaline, with a blackish basal stripe and three blackish bands confluent in an S-shaped mark ; third longitudinal vein naked. Long. corp. 0.20. Long. al. 0.21. Of this species, very conspicuous by the peculiar picture of its wings, I unfortunately possess only one individual, much injured in carrying. It is everywhere pale yellow, and its thorax and scutellum have no trace of a paler picture. Hairs very short, whitish-yellow on the upper side of the thorax, rather blackish on the pleurae ; bristles all black. Scutellum with four bristles. Wings hyaline ; the veins at the base of the wing yellowish ; a blackish not very striking stripe runs from the tip of the basal humeral vein to the posterior angle of the anal cell, which is drawn out into a point. The remainder of the picture of the wings con- sists of three very broad, rather blackish bands ; the first runs from the black stigma, widening gradually perpendicularly to near the posterior border, where it is connected with the second, which rises over the posterior transverse vein as far as the costal border, and connects there completely with the third band which seams the tip of the wing. The connection of the first and second bands is somewhat interrupted by a clear incision reaching from the poste- rior border a little into the discal cell. Above the end of this incision there is another clear spot. Stigma small ; none of the longitudinal veins unusually curved ; the small transverse vein is somewhat before the middle of the discal cell and below the very tip of the first longitudinal vein ; the posterior transverse vein is only a little arcuated ; the two transverse veins are steep, not perfectly perpendicular. Hub. Middle States. (Osten-Sacken.) 76 DIPTERA OP NORTH AMERICA. 1O. T. Ciligulata LOEW. $>. (Tab. II, fig. 11.)— Nigra, capite pedibusque luteis, thoracis raargine laterali scutelloque flavis, margine segmentorurn abdominaliuin singulorum postico albido, alarum hyali- narum fasciis quatuor punctoque apicali nigris. Black with the head and legs luteous, the lateral borders of the thorax and the scutellum yellow, the posterior borders of the abdominal seg- ments whitish ; wings hyaline with four bands and an apical dot black. Long. corp. 0.22. Long. al. 0.20. It belongs to the relationship of the European Tryp. cerasi Linn. (= signata Meig.), a group which must not be confounded with that of Tryp. solstitialis Linn., closely alike in its coloring. Black. Head rather dark yellow, front brighter yellow, of middle breadth, with rather long black bristles. Antennae reaching to a little beyond the middle of the face, last joint rather narrow, and with the anterior corner rather sharp. Face straight, descending, with moderately deep furrows for the reception of the antennae ; border of the mouth by no means prominent. Proboscis and palpi short. Thorax black ; the humeral callosity and a longitudinal stripe running from the latter to the base of the wing are bright yellow. The bristles of the thorax and the four bristles of the yellow scutellum black ; the base and greatest part of the lateral border of the latter black. Metanotum and abdomen glossy black, the latter with broad whitish (perhaps more yellow in life) edges of the posterior borders and black hairs. Borer exceedingly short. Legs dark yellow; femora a little brownish at the base; the anterior femora with small brown bristles on the under side; the two posterior tibia? with short black bristles on the upper side. Wings somewhat broad, especially in the neighborhood of the base, with four black bands and a little black spot at the tip. The first band runs from the basal humeral vein to the posterior angle of the anal cell, which is drawn out into a point. The second is broadest, running from the black stigma beyond the fifth longi- tudinal vein, and ending abruptly in the middle between this vein and the border of the wing. The third, which runs over the pos- terior transverse vein, is also rather perpendicular, and completely reaches the posterior border of the wing. The fourth band is perfectly united with the third on the costal border, and reaches the posterior border behind the tip of the fourth longitudinal vein, so that it has a rather oblique position. The small apical spot TRYPETIDAE. 77 includes the tip of the third longitudinal vein. Both transverse veins are straight and perpendicular. Jfab. Middle States. (Osteu-Sacken.) 11. T. polita LOEW. 9. (Tab. II, fig. 12.)— Atra, nitida, capite pedibusque flavis, scutello tuniido, alarum albido-hyaliuarum macula basali atra fasciisque tribus latissimis fusco-nigris. Deep black, shining ; head and legs yellow, scutellum inflated ; wings whitish-hyaline with a basal black spot and three very broad brownish- black bauds. Long. corp. 0.25. Long. al. 0.17 — 0.18. Belongs to the relationship of the European Tryp. Wiedemanni Meig., the species of which chiefly agree in their inflated scutellum and short wings, while they differ among each other much in the structure of their face. Front bright yellow, beautifully yellowish- brown above, considerably broad ; frontal bristles black. Antennas yellowish, descending to the middle of the face, and having a black bristle, the pubescence of which is exceedingly short and hardly visible. Face whitish-yellow, a little receding, its middle rather flat ; border of the mouth not prominent at all ; opening of the mouth rather small ; proboscis and palpi short. The inferior part of the occiput is whitish-yellow, the superior blackish. Thorax rather convex, altogether glossy black, bare, but the broad lateral stripes are bordered everywhere with a row of yellowish short hairs, and the broad middle stripe is divided by a longitudinal row of such hairs. Bristles black. Scutellum shining black, very con- vex, as if inflated. Metanotum black, with an indistinct whitish reflection. Pleurse shining black, with a few stiff yellowish hairs and some black bristles. Abdomen black ; the hairs rather stiff, whitish on the posterior part of the first segment ; on the second and third segments they are black, except the hindmost ones of the posterior border, which are whitish; on the two last segments they all are whitish. Borer shining black, flattened, pointed, abundantly as long as the abdomen, with very short black hairs. Legs dirty fuscous-yellow; femora not much incrassated, the anterior ones with a few black hairs on the under side. Wings short and rather broad, having the transverse veins very approximated and perfectly perpendicular; they are rather whitish, with very broad brownish- black bands. Their innermost base is yellowish, then follows a large triangular rather deep black spot, which reaches from the costal border as far as the axillary incision of the wing, and only 78 DIPTERA OF NORTH AMERICA. little exceeds the basal cells. The two first black or brownish- black bauds are united on the costal border, so as to form an inverted Y; the second of them runs from the stigma over the transverse veins, and has a more inclined position than the first ; the isolated third band has a position similar to that of the second, and seams the tip of the wing some distance beyond the tip of the fourth longitudinal vein, without coalescing anywhere completely with the border of the wing. ffab. Mississippi. (Schauni.) Washington. (Osten-Sacken.) Observation. — The Brazilian Urophora connexa Macq. (Dipt, exot. Suppl. Ill, 64, Tab. VII, fig. 10) has the picture of the wings rather similar to those of Tryp. polita. But being greenish- black, and having the last band of the wings completely connected with the preceding, and running straight on with the costal border, but not reaching the tip of the third longitudinal vein, it is evi- dently different from Tryp. polita. 12. T. sparsa WIED. £ and £. (Tab. II, fig. 13.)— Fusca, alis latis- simis, rotundatis, nigris, albido-guttulatis et mavgine apicali albo ornatis. Brown ; wings very broad and rounded, black with small drops, and the apical border whitish. Long. corp. 0.27 — 0.30. Long. al. 0.26. SYN. Trypeta sparsa WIEDEMANN, Auss. Zweifl. II, 492. Trypt'ta caliptera SAY, Journ. Acad. Phil. VI, 187, 3. Platystoma latipennis MACQUART, Dipt. exot. II, 3, 200. Tab. XXVI, fig. 8. Acinia novseboracensis FITCH, First Rep. etc. 67. Of a brownish-red, sometimes more brown color. Front broad, brownish-yellow, frontal bristles black; the bristles of the poste- rior orbit whitish-yellow. Antennas descending below the middle of the face, their bristles with a short, but distinct- pubescence. Face excavated in the middle, and marked with two large deep black dots ; another small deep black spot is between the antenna and the anterior angle of the eye. Eyes with three very distinct transverse bands. Palpi dark brown, usually blackish at the tip ; suctorial flaps a little prolonged. On the upper side of the thorax there are usually two or three darker longitudinal lines, and a broad dark-brown edge of the lateral border. The short hairs of the thorax and scutellum are whitish-yellow, the bristles of both black. Of the four bristles placed in the middle of the thorax, the two first are near the transverse suture. Pleura above with whit- TRYPETIDAE. 1 9 ish-yellow, below with black hairs, the bristles mixed among them black. The abdomen usually bears two rows of large blackish spots, of a rather quadrangular form, leaving between them a brown middle streak, and not completely reaching the posterior borders of the single segments ; sometimes they extend so much, that only the posterior borders of the segments retain a paler color, whereas the whole surface is blackish. Borer blackish-brown, sometimes with a red spot on each side, a little pointed, rather flat; its first segment is a little longer than the two last segments of the abdomen taken together. The hairs of the abdomen are mostly black, a few light ones being among them. Legs paler reddish- brown, the anterior femora often partly blackened, with some black bristles on the upper and under sides. Wings exceedingly broad, with the uninterrupted whitish seam of the tip forming a narrow crescent ; on their surface there are numerous clear drops rather equally scattered, but totally wanting in that part of the black color which adjoins the white crescent, as well as before the first longitudinal vein. At the tip of the first longitudinal vein, a small, clear, but little distinct stripe is seen. The small transverse vein straight and perpendicular, the posterior one a little curved and steep. Hob. Northern Wisconsin. (Kennicott.) Observation. — I possess a specimen, which is distinguished by its much paler, almost dull testaceous color, its less enlarged wings and the somewhat larger size of the clear drops, but as to the other characters agrees so perfectly with the ordinary specimens of Tryp. sparsa Wied. that I do not venture tp declare it a different species. 13. T. rotimdipeiinis LOEW. £ . (Tab. II, fig. 14.)— Fusca, alis latissimis, rotundatis, nigris, albido-guttulatis et in marginibus ante- riore et apicali maculas minutas albidas gerentibus. Brown ; wings broad and rounded, black, with very small whitish drops in the middle, and small whitish spots on the costal and apical borders. Long. corp. 0.28. Long. al. 0.26. Of this species I have only one specimen, which is unfortunately so much injured in the journey as to prevent me from giving a full description. However, as it is very nearly related to Tryp. sparsa Wied., it will be recognized even from my incomplete de- scription. The color is the same ; the wings are still shorter and broader, especially the cell which lies before the first longitudinal 80 DIPTERA OF NORTH AMERICA. vein is much broader ; the third longitudinal vein is much more undulated, and the last portion of the fourth longitudinal vein is shorter than the posterior transverse vein, whereas it is a little longer in Tryp. sparsa. None of the drops on the wings of Tryp. sparsa is of a more considerable size than the others, whereas Tryp. rotundipennis has such a drop between the third and fourth longitudinal veins, opposite to the base of the discal cell ; the drops are in general equally distributed in Tryp. sparsa, they are much more irregularly grouped in Tryp . rotundipennis ; the size and number of the drops diminishes a little from the base towards the apex in Tryp. sparsa; their number only, not their size diminishes in Tryp. rotundipennis ; their size increasing towards the posterior border in Tryp. sparsa, this is not the case in Tryp. rotundipennis. In Tryp. sparsa the white crescent seaming the apex of the wing is entire, in Tryp. rotundipennis it is dissolved into several spots. Finally, the anterior border of the wings of Tryp. rotundipennis bears a row of small clear spots, whereas Tryp. sparsa has no trace of them. Hab. Middle States. (Osten-Sacken.) 14. T. clatlirata LOEW. 9. (Tab/II, fig. 15.)— Caua, capite pedi- busque flavis, femoribus litura nigra signatis, alls rare reticulatis, stig- mate atro albo-guttato, peristonrio modice producto, proboscide breviter geniculata. Whitish-gray ; head and legs yellow ; wings with a diffuse reticulation, and the black stigma including a limpid drop ; oral border moderately prolonged, proboscis shortly geniculated. Long. corp. 0.12 Long. al. 0.13. Head yellowish ; the lateral borders of the front, the face, and the much descending cheeks whitish. The bristles on the front black, on the sides of the vertex and posterior orbit white. The face with rather deep subantennal furrows prolonged to the oral border, which is not very projecting. Antennae fulvous, rather short ; the anterior corner of the third joint a little acute ; the second joint with very short black hairs ; antennal bristle black and moderately long, with the pubescence scarcely visible. Tho- rax and scutellum whitish gray, with short pubescence and black bristles. The scutellum bears four bristles, the two apical, ones being much shorter and less stout than the lateral ones ; its tip is sometimes yellow. The whitish-gray abdomen has two rows of very TRYPETIDAE. 81 distinct black spots. The hair and even the bristles on the hind border of the last segment are white. The flat ovipositor is glossy- black, as long as the two last segments of the abdomen and mode- rately tapering towards the tip. The legs and anterior coxae are dark yellow, with white hairs ; the thighs with a blackish stripe somewhat covered with whitish dust. Wings not very narrow, hya- line, with the base very slightly yellowish ; their black reticulation is not very dense and not very delicate, extending towards the base of the wing as far as the base of the discal cell, and dissolving towards the axillary angle into some scattered spots ; stigma black, with a whitish dot; the middle and posterior transverse veins rather approximate. Hob. Middle States. (Osteu-Sacken.) Observation. — The proboscis of this species being short with the suctorial flaps but moderately prolonged, attention is to be paid to its true place, which is among the species with a geniculated pro- boscis. 15. T. lllimilis LOEW. £ . (Tab. II, fig. 17.) — Cinerea, capite pedi- busque flavis, femoribus nigris, alis rare reticulatis, stigmate atro non guttato, peristoniio valde producto, proboscide geniculata. Cinereous ; head and legs yellow, with the femora black ; reticulation of the wings diffuse, and the black stigma including no limpid drop ; oral border much prolonged ; proboscis geniculated. Long. corp. 0.09. Long, al. 0.1. Among the kindred of the European Tryp. elongatula Loew, but in its habit more resembling Tryp. absinthii Fabr. Head yellow, considerably prolonged anteriorly. Front yellow, dis- tinctly margined with white at the orbit ; frontal bristles black. Face excavated with the anterior border of the mouth much pro- longed. Antennae bright yellow, proportionately large and broad, reaching as far as the prolonged border of the mouth ; the ante- rior corner of the third joint a little pointed ; the second joint with very short black hairs ; antennal bristles very long, black, with a very short and hardly visible pubescence. Thorax and scutellum yellowish ashy-gray with short whitish-yellow hairs and black bristles. In the middle of the thorax there are four bristles, the two foremost being very near the suture. Scutellum with only two long bristles rather distant from the tip and not close by its lateral border. Metanotum black with grayish pollen. Abdomen ashy- 6 82 DIPTERA OF NORTH AMERICA. gray with two rows of rather large blackish spots. Its hairs are whitish yellow ; some black bristles on the posterior border of the prolonged last segment. Tip of the first joint and the second joint of the coxaa rather dark yellow. Femora brownish-black, with dark yellow tips. Tibiaa and tarsi dark yellow. Wings pro- portionately rather long and narrow. Their reticulation is black, rather diffuse and coarse ; the base of the wing as far as the base of the discal cell has no reticulation. The black stigma includes no clear dot. Moreover, the reticulation of the wings is somewhat variable. Transverse veins rather near each other. Hab. Cuba. (Poey.) Observation. — I have a female likewise captured in Cuba, which, , I think, belongs to the present species. It resembles the male in everything but the femora, which are darkened to a much smaller extent, and not with black, but brown. The borer is black, flat, about as long as the two last segments of the abdomen taken together. 16. T. soliclaginis FITCH. £ and ?. (Tab. II, fig. 16.)— Rufo ferruginea, capite pedibusque flavioribus, fronte latissima, setis scutelli valde convex! duabus, alis fusco-reticulatis, incisuris una anteriore, duabus posterioribus apiceque hyaliuis, parcissime fusco-maculatis. Brownish-ferruginous with the head and legs more yellow ; front very broad; scutellum very convex with two bristles. Wings reticulated with fuscous having one limpid space at the costa and two at the posterior border scarcely dotted with fuscous. Long. corp. 0.26. Long. al. 0.26. SYN. Acinia solidaginis FITCH. First Rep. 66. This remarkable species, which, according to Mr. Fitch, produces round galls on the stems of Solidago,has no near relations among the European Trypetce. In consequence of the extraordinary breadth of its front, the breadth and convexity of its thorax, and the in- flation of its large scutellum, it has the appearance of a large Lipara. It is brownish ferruginous. Head more yellow, face almost whitish. The bristles of the unusually broad front are black, smaller and weaker than in most other species, so that one might easily be tempted to refer the species to the Ortalidce, if the structure of the auxiliary vein did not prove that it belongs here. Face deepened in the middle, prominent again underneath. An- tennae yellow, short and broad, the third joint having a rather dis- TRYPETIPAE. 83 tinct, but not sharp anterior corner ; bristle of the antennae with a hardly visible pubescence. Opening of the mouth very wide ; palpi and proboscis short. Thorax very convex and broad ; on its upper side in the middle usually a double stripe ending abruptly behind, the posterior part of the lateral stripe and a longitudinal streak above the base of each wing, black. The short hairs of the thorax are whitish yellow, its delicate bristles black. Scutel- lum very convex, quite blunt; the posterior pair of bristles which in most species exists at the tip, is here always wanting so that there is only one bristle on each side near the lateral border ; a second weaker bristle is seldom inserted immediately beside it. Abdomen broad, especially in the male, which has also the last segment a little prolonged and rounded. Borer of the female a little longer than the two last segments taken together, moderately broad, and quite flat, red, blackened at its extremity. Legs rather dirty yellow, femora more brownish. Wings rather large and of more equal breadth than usual. The reticulation of the wings is almost umber brown with small pale brownish drops and hyaline marginal spots very rarely dotted with brown ; the first of these spots is triangular and extends from the posterior border to inside of the discal cells ; the second is much smaller, but also of tri- angular form, and reaches with its tip to the fourth longitudinal vein ; the third forms a margin along the apex of the wing, reach- ing from the tip of the second longitudinal vein to the tip of the fourth longitudinal vein ; the last spot forms a small oblique tri- angle extending from the costal border to the third longitudinal vein, and lying immediately beyond the tip of the first longitudinal vein. A costal spine does not exist. The first longitudinal vein is more hairy than bristly; the transverse veins are perpendicular; the small transverse vein is almost at the end of the second third of the discal cell ; the hind angle of the anal cell has only a short point. Hab. New York. (Dr. Fitch.) Washington. (Osten-Sacken.) Observation. — This species has so many peculiarities, that it might easily be considered as the type of a new genus. The broad front, broad and convex thorax, a scutellum having only two bris- tles, the first longitudinal vein alone being hairy, and the absence of the costal spine, would be its most essential characters. 84 DIPTERA OF NORTH AMERICA. 17. T. seriata LOEW. £ . (Tab. II, fig. 18.) — Flava, alis concoloribus, per maculas minntas nigricantes, seriatim dispositas, reticulatis et nigro- limbatis. Yellow ; wings of the same color, margined with black and reticulated with small blackish spots, arranged in longitudinal rows. Long. corp. 0.24. Long. al. 0.26—0.27. Rather bright yellow, quite unicolorous, the hairs and bristles also yellow, the latter, however, appearing brown when seen in n certain light. Front rather broad. Face descending rather straight, strongly excavated, however, in its middle. Proboscis short ; palpi rather broad and short. Scutellum with four bristles. The wings are proportionately long, and of very equal breadth. Their yellow ground color is rather obsolete in the middle line of the cells and on the posterior border. The reticulation of the greater part of the wings is effected by blackish angular specks arranged in two rows between each two veins ; only the axillary angle has a connected blackish-gray reticulation formed by clear drops. Immediately before the tip of the auxiliary vein begins the black margin of the wing, which encompasses the apex and pro- ceeds to the axillary angle, growing gradually paler in its progress, and meeting several interruptions ; the most remarkable of these interruptions are a rather hyaline spot immediately beyond the tip of the first longitudinal vein, and a row of similar round spots along the posterior border, the two first of which in the second posterior cell, the following more frequent towards the axillary angle. The third longitudinal vein bears very distinct bristles from its base as far as the small transverse vein. The small trans- verse vein is nearly at the end of the second third of the discal cell. The posterior transverse vein is not cjuite perpendicular. The hind angle of the anal cell is prolonged into a moderately long point. Hob, Middle States. (Osten-Sacken.) IS. T. Solaris LOEW. 9 . (Tab. II, fig. 19.)— Cana, capite pedibusque luteis. puncto humeral! et altero ante alarum basim flavis, alis albohya- linis, macula magna subapicali nigra, biguttata et radios octo emittente, ornatis. Whitish gray ; head and legs luteous, a dot on the shoulder and another before the base of the wings yellow ; wings whitish hyaline, with a sub- TRYPETIDAE. 85 apical black spot including two limpid drops and emitting eight rays to the border of the wing. Long. corp. 0.17. Long. al. O.lfJ — 0.17. One of the group of the European T. stellata Fuessl., cometa Loevv, gnaphalii Loew, etc., and very much resembling these species. Head yellow; front rather broad; frontal bristles black- ish ; the bristles of the posterior orbit whitish. Front a little prominent, face slightly receding and a little excavated in the middle, so that the borer of the mouth projects again. Antennae rather broad, reaching down to beyond the middle of the face, with the bristle having a very short, hardly visible pubescence. Open- ing of the mouth very large ; proboscis and palpi short. Thorax whitish-gray with a pale yellow dot at the shoulder angle, and a second immediately before the base of the wing. The short hairs of the upper side of the thorax are whitish, the bristles blackish ; of the four bristles in its middle, the first pair is very near the suture. Scutellum with only two long bristles. Abdomen whitish- gray at the base, more ashy gray towards the end, with short, whitish-yellow hairs, the hind border of the last segment having black bristles. Borer shining black, flat, tapering towards the end, nearly as long as the three last abdominal segments taken together, with black hairs. Legs dark-yellow. Wings whitish hyaline before the tips, with a large radiating spot, incumbent to the costal border ; this black spot includes two clear drops, one of which on the costal border immediately behind the tip of the second longitudinal vein, the second between the two transverse veins at the anterior side of the fourth longitudinal vein. The first ray runs from the anterior end of the small transverse vein to the stigma, in which it vanishes ; the second is shorter and reaches the costal border between the tip of the first longitudinal vein and the black spot itself; the third and fourth rays run to the tip of the wing, reaching it at the tips of the third and fourth longitu- dinal veins ; the fifth and sixth cross the second posterior cell ; the seventh includes the posterior transverse vein and reaches the pos- terior border of the wing, whereas the eighth reaches only to the fifth longitudinal vein. The small transverse vein lies outside of the black spot ; yet in its whole neighborhood the surface of the wing is brownish, and a small gray spot lies immediately before it. Transverse veins approximated, perpendicular ; the small trans- verse vein rather far beyond the tip of the first longitudinal vein. Hob. Georgia. (Osten-Sacken.) 86 DIPTERA OF NORTH AMERICA. 19. T. aequalis LOEW. ^ . (Tab. II, fig. 20.) — Flava, unicolor, alis sequaliter fusco-reticulatis, guttis hyalinis plerisque majoribus. Yellow, unicolored, with the brown reticulation of the wings very uniform and most of the limpid drops of considerable size. Long. corp. 0.22— 0.23. Long. al. 0.23. Totally yellow, also the hairs and bristles, only the anterior frontal bristles and those at the hind border of the last abdominal segment being brownish. . Front rather broad and short. Face descending nearly straight, excavated a little above ; the border of the mouth not projecting. Antenna yellow, slightly descend- ing beyond the middle of the face, third joint rather narrow, bristle with an extremely short, hardly visible pubescence. Opening of the mouth small, a little prolonged anteriorly, so that its form is almost triangular. Proboscis and palpi of middle size. The first pair of the bristles inserted in the middle of the thorax, is very near the transverse suture. Scutellum with four bristles. Legs a little more slender than usual, tarsi longer; anterior femora rather thick, with remarkable, pale yellowish bristles on the under side. Wings of rather equal breadth, hyaline with a brownish reticulation of unusual uniformity. Most of the drops forming it are rather large ; those on the costal border are more oblong, and separated by short blackish-brown rays ; the color of the reticulation near the border of the wings is considerably darker than in the middle of the wing. The small transverse vein lies far beyond the tip of the first longitudinal vein, and a little beyond the third fourth of the discal cell ; the posterior transverse vein is a little oblique. Hob. Illinois. (Kennicott.) 20. T. f estiva LOEW. ^ and 9. (Tab. II, fig. 21.) — Flava, unicolor, terebra fceminje valcle elongata fusca, alis hyalinis insequaliter reticulatis, in apice radiatis, pictura in basi et disco flavescente, prope marginem an- ticum et in triente alarum apicali nigro-fusca. Yellow, unicolored ; the borer of the female brown and very prolonged ; reticulation of the wings unequal, radiated in the apex, yellowish at the base and in the disk, dark fuscous near the borders and on the apical third of the wing. Long. corp. $ 0.17—0.18, J 0.20—0.23. Long, al. 0.22. Rather bright yellow ; hairs and bristles almost all of the same color, only the anterior frontal bristles as well as the bristles of TRYPETIDAE. 81 the hind border of the last abdominal segment brownish, and the short hairs in the middle of the abdomen mostly blackish. Front of middle breadth. Face rather narrow, descending almost per- pendicularly, slightly excavated ; border of the mouth not promi- nent. Antennae yellow, reaching a little beyond the middle of the face ; the third joint not broad, with the bristle having a very short, hardly visible pubescence. Opening of the mouth rather large, raised a little anteriorly. Proboscis and palpi of middle size. Cheeks descending a little beneath the eyes. The foremost of the two pairs of bristles inserted in the middle of the thorax is close by the transverse suture. Scutellum with four bristles. Abdo- men sometimes brownish-tawny, leaving the hind borders of the segments paler ; this color seems to result from desiccation, since in some individuals the abdomen is uniformly yellow. The borer is conical, narrow, not flattened, nearly as long as the four last ab- dominal segments taken together, blackish-brown in well-colored individuals, red with black extremity in more recent individuals. Wings hyaline, the reticulation being blackish-brown, paler and yellowish-brown near the base and in the middle of the wing. In the middle of the wing there are only a few drops of considerable size, four of which are remarkable for their regular position and a more whitish appearance ; one of these drops is above, the second before, the third behind the small transverse vein, the fourth in the discal cell nearly before the posterior transverse vein. The reticu- lation sends the following blackish-brown rays to the border of the wing : 1. A narrow one to the middle of the exterior cbstal cell ; 2. A narrow one to the tip of the auxiliary vein ; 3. A broader one, the end of which is sometimes separated as a spot, to the middle of the stigma, and another being sometimes confluent with it, to the end of the first longitudinal vein ; 4. A narrow one rising from the first of the four drops enumerated above; 5. A very broad one reaching the border of the wing between the fore- going ray and the tip of the second longitudinal vein ; 6. A ray running to the tip of the second longitudinal vein ; 7. A ray end- ing between the tips of the second and third longitudinal veins ; 8 and 9. Two rays running to the tips of the third and fourth longitudinal veins ; 10 and 11. Two rays crossing the second pos- terior cell, the second of which joins the reticulation, which is formed by a few large drops, and fills the hind part of the wing as far as the axillary incision. Small transverse vein a little inclined 88 DIPTERA OF NORTH AMERICA. exteriorly, placed at the end of the second third of the discal cell ; posterior transverse vein steep. Hob. Pennsylvania. (Osten-Sacken.) 21. T.liellaLoEW. -fcand?. (Tab. II, fig. 23.)— Flavo-cinerea, capite, pedibus abdomineque luteis, hoc apicem versus nigricante, alis nigro- reticulatis, in apice radiatis, guttis disci paucissimis, pone venulam transversam nulla. Yellowish gray ; head, legs, and abdomen yellow, the latter blackened towards the end ; reticulation of the wings radiating at the apex, black, with very few drops in the middle ; no drops at all beyond the small transverse vein. Long. corp. £ 0.12 — 0.13. J 0.13 — 0.15. Long. al. 0.11—0.12. Head rather pale yellow, front and antennas sometimes darker, the former being of middle breadth, slightly narrowed anteriorly. Face rather narrow, nearly perpendicular, slightly excavated, the anterior border of the mouth again projecting a little. Antennas yellow, reaching to nearly the border of the mouth; the third joint somewhat broad, with the bristle having an extremely short, hardly visible pubescence. Opening of the mouth very large, a little raised anteriorly. Proboscis and palpi rather large. Cheeks descending a little beneath the eyes. Thorax yellowish-gray. Scutellurn of the same color, pale yellow at the tip, to a larger or smaller extent, with four bristles. Metanotnm black, but dark gray from its being dusted with paler. Abdomen dark yellow, black towards the extremity ; well preserved specimens show distinctly that this black color is produced by each of the last segments having two large blackish spots, which leave an intermediate streak, and the posterior border yellow; in most specimens these black spots are not distinct, or only the hind borders of the last segments are paler. Borer black, rather broad, narrower towards the end, flat, little longer than the two last segments taken together. Legs yellow. Hairs of the whole body and all bristles yellowish, only the very short hairs of the borer being black. The reticulation of the wings is blackish-brown, leaving only the innermost base of the wing free, with the exception that it has some blackish spots. In the middle of the wing there are so few clear drops, that the black color is not only continuous, but also occupies most part of the surface ; in the sub-marginal cell there is only one clear drop, near the hind side of the second longitudinal vein and a little beyond the small transverse vein. Between the third and fourth longitu- TRYPETIDAE. 89 dinal veins there is only a single clear drop on the anterior side of the antepenultimate portion of the fourth longitudinal vein. The want of that drop which is usually inside of the said interval be- yond the small transverse vein, is very characteristic of this species, as is also the considerable depth and blacker color of the convexity existing there. In the discal cell there is always one drop on the hind side of the penultimate portion of the fourth longitudinal vein, and one or more such drops on the anterior side of the fifth longitudinal vein. The third posterior cell and the axillary angle of the wing have a reticulation produced by a few, propor- tionately large, hyaline drops. The reticulation of the wings emits ten rays to the borders, corresponding to those of Tryp. festiva, except that the first ray of the latter species is wanting totally in Tryp. bella, and the two rays described under No. 3 in Tryp. festiva are reunited into one single ray in Tryp. bella ; the last ray in Tryp. bella is usually connected again with the remain- ing reticulation ; in this case the second posterior cell also con- tains a separated clear drop. Posterior transverse vein not quite perpendicular. Hob. Washington. (Osten-Sacken). New York. (Dr. Fitch.) Observation. — Among the specimens forwarded by Baron Osten- Sacken there was one bearing the name of Acinia bella Fitch. I have therefore adopted this name for this fine new species. J^fote. — Very common on Ambrosia artemisicefolia. — 0. S. 22. T. latifrons LOEW. 9 . (Tab. II, fig. 22.)— Obscura, capite, tibiis tarsisque flavescentibus, fronte latissima, scutello convexo biseto, alis latiusculis, parce et satis sequaliter nigro-fusco reticulatis et in apice breviter radiatis, bulla cellula posterioris primse permagna. Obscure ; head, tibiae, and tarsi yellowish ; front very broad ; scutellum very convex, with two bristles ; wings rather broad, with the blackish reticulation rather uniform, but little crowded, and emitting short rays at the tip ; the first posterior cell with a very large convexity. Long, corp. 0.30. Long. al. 0.27. The single specimen of this species which I have seen being oily, I can say nothing certain about the color of its body. On the upper side of the thorax there is a broad, simple intermediate stripe, and on each side a bipartite lateral one, moreover a darker streak above the base of the wing. Metanotum black, shining. The last abdo- minal segment shining blackish-brown. Head yellowish ; front 90 DIPTERA OF NORTH AMERICA. * exceedingly broad; frontal bristles black; on the lateral border there are only two, bent anteriorly. Face perpendicular, deeply excavated in the middle, gradually projecting again below. An- tennae yellowish, short, very broad, the third joint having a distinct, though not sharp anterior corner, the bristle with an exceedingly short pubescence. Cheeks rather broad. Opening of the mouth large; proboscis short; palpi very broad and projecting much over the border of the mouth. The short hairs of the thorax are whitish-yellow, the bristles black ; the anterior one of the two pairs of bristles inserted in the middle of the thorax is very near to the suture. Scutellum very convex, with only two bristles. Borer shining black, a little longer than the three last abdominal seg- ments taken together, conical, not flattened at all, a little inflated at the basal half. Femora almost black at the base, further on brown, yellow at the tip; tarsi and tibiae yellow, the latter brown- ish-yellow towards the base. Wings rather broad, covered entirely with a black reticulation ; the drops in it are more numerous, larger, and hyaline on the borders of the wings, much more scarce, smaller, and mostly yellowish-brown in their middle. There are eight short, blackish-brown rays on the portion of the costal bor- der lying beyond the tip of the first longitudinal vein and at the tip of the wing. Between the third and fourth longitudinal veins there is only a single hyaline drop before the small transverse vein. Transverse veins perpendicular; the convexity of the first posterior cell very large and deep, rendering thereby its surroundings rather uneven. Hob. Carolina. (Zimmermann.) T. melaiiogastra LOEW. $ and £ . (Tab. II, fig. 23.)— Flavo- cinerea, abdomine nigro, capite pedibusque flavis, alis hyalinis, nigro- reticulatis et in apice breviter radiatis, guttis disci paucissiinis, bulla cellulse posterioris primse minima, scutello biseto. Yellowish-gray; abdomen black, head and legs yellow; wings hyaline with a black reticulation, short apical rays, few discal limpid drops, and a small convexity in the first posterior cell ; scutellum with two bristles. Long. corp. £ 0.09— 9 0.12. Long. al. 0.12. Yellowish-gray, with a black abdomen. Head yellow; front bright yellow, of middle breadth ; frontal bristles brown, but ap- pearing yellow in a reflected light. Face rather narrow, descend- ing straight, excavated in the middle, gradually projecting again TRYPETIDAE. 91 beneath. Antennae yellowish, rather broad, with the third joint having a distinct anterior corner, and the bristle with a short, hardly visible pubescence. Eyes almost round. Cheeks narrow. Opening of the mouth large ; proboscis and palpi short. The short hairs of the thorax pale yellowish, the bristles brownish; the first of the two pairs of bristles in the middle of the thorax very near the transverse suture. Scutellum of the color of the thorax, but usually yellow at the tip, with two bristles. Abdomen and metanotum black, rather glossy; the short and scattered hairs of the former pale yellowish. Borer flat, shining black, nearly as long as the abdomen. Legs yellow. Wings hyaline, with the reticulation blackish and a little interrupted, leaving only the innermost base of the wing free. The following hyaline spots produce the reticulation on the borders of the wing : one before the tip of the auxiliary vein; two between the tips of the first and second longitudinal veins, and sometimes a drop immediately before the tip of the second longitudinal vein, five oblong incisions limit- ing the four rays emitted to the tip of the wing ; a clear drop touching the border beyond the tip of the first longitudinal vein, often confluent with a drop lying immediately above it, and forming thereby an incision ; three drops between the tips of the fifth and sixth longitudinal veins, the intermediate of which is the largest ; a drop immediately before the tip of the sixth longitudinal vein, and one in the axillary angle. On the middle of the wing there are the following rather large clear drops : one on the hind side of the second longitudinal vein, a little beyond the small transverse vein; the others on the anterior side of the fourth longitudinal vein, one being before, the other behind the small transverse vein; one on the anterior side of the fifth longitudinal vein, and a little before the small transverse vein, having sometimes a small drop on each side ; a very large drop in the middle of the third posterior cell, and a very minute one at the innermost base of this cell. Hob. Cuba. (Poey.) APPENDIX I. I give here as an appendix a translation of Wiedemann's descrip- tions of four species, which I have formerly seen, but have not before me at present. 92 DIPTERA OF NORTH AMERICA. 1. T. liclitensteinii WIED. (Auss. Zweifl. II, 497, 31.) £ . (Tab. II, fig. 25.) Clay-colored ; wings with, the tip, a square spot on the hind border, an oblique band and some dots, brown. Long. corp. 0.2. Front and antennae rather bright yellow, all the other parts more or less honey-yellow, turning to clay color on the thorax. The fuscous color of the apex of the wing extends farther along the anterior than on the posterior border, and forms a little tooth on the fourth longitudinal vein ; a large square fuscous spot is situated on the hind border, and includes the posterior transverse vein. Between this spot and the brown apex there is a triangular almost hyaline space, having a very limpid drop in each corner, and including a small fuscous spot on the hind border of the wing. The anterior corner of the square fuscous spot is connected with the stigma by an oblique fuscous band, including the small trans- verse vein ; there are besides a small fuscous dot above the fifth longitudinal vein, and another slightly larger below this vein, and also a brown margin of the small basal transverse veins ; some pure limpid drops of a rather large size are seen near the borders of the larger brown spots. Hah. Mexico. Observation. — The bristle of the antennae is thickened near the base in a striking and peculiar manner. In each of the sinuses of the large hyaline spots of the wing there is a large whitish drop, not a clear one, as Wiedemann states. 2. T. scutellaris WIED. (Auss. Zweifl. II, 484, 13.) £ and £. (Tab. II, figs. 26, 27.) Thorax with the lateral border spotted with black ; scutellum polished, brown with a reddish stripe ; abdomen marked with blackish-brown bands ; wings with brown bauds and spots. Long. corp. 0.26. Antenna pale ochreous ; front isabella-colored, with the upper part gray. Middle of the thorax grayish, with two darker stripes and some little dots. Scutellum polished, with the lateral borders dark brown and the middle reddish. Abdomen very pale gray at the base, with a fuscous band, a little interrupted in its middle ; the third, fourth, and fifth abdominal segments each with a similar band at the base, but more interrupted in the middle and attenuated TRYPETIDAE. 93 towards the sides. Ovipositor broad, red on each side of the base, the remainder gray, with the tip fuscous. Wings with a brown band running over the basal transverse veins, but not attaining the posterior border; and with a complete band before the middle transverse vein, emitting another oblique band which crosses the two transverse veins and runs to the posterior border. Tip of the wing brown. The costal portion of the space, included by the second band and the brown of the tip, is tinged with brown on its basal half, whereas its apical half is yellow and spotted with brown before a small hyaline margin of the brown apex ; one of the brown spots in the yellow half reaches the third longitudinal vein. Hal). Mexico. Observation. — The number and size of the black spots on the lateral border of the thorax is rather variable. In the female the abdomen has alternately gray and black bands ; in the male only the two last segments of the abdomen are shining black, with the exception of the posterior borders. I am not sure whether this species is a real Trypeta, several characters seeming to prove that it belongs to Ortalis. Many years have elapsed since I have seen it ; besides, at that time I was not quite certain about the true limit between the Trypetid. (Loew, Berl. Entom. Zeitschr. 1861, p. 345.) — Albida, alis concoloribus imrnaculatis, fronte, pleuris, scutello, segmeritorumque abdominalium singulorum margine postico palMe sul- phureis, facie cum antennis, pedibus et terebra obscurius flavis. Whitish, wings whitish, without spots, front, pleurre, scutellum and the posterior border of the abdominal segments pale sulphur-yellow ; face, antennae, legs and borer of the J darker yellow. Long. corp. 0.13 — 0.17. Long. al. 0.15— 0.1G. Antennas of moderate length ; tip of the third joint round, bristles with a very short pubescence. Eyes large, almost round; cheeks moderate. Oral opening moderate, rounded, proboscis not geniculated, palpi short. Scutellum flat, with four bristles. Borer of the 9 flattened, ferruginous-yellow, longer than the three last segments of the abdomen taken together. Wings whitish, all the veins very pale, and, except the first longitudinal one, bare ; pos- terior angle of the anal cell acute. Hub. Pennsylvania. (Osten-Sacken; taken on Vernonia novce- boracensis, iron weed, together with the two following species, and with T. obliqua Say; in August.) 19. T. alMdipennis LOEW. £ and 9 . (Loew, Berl. Entom. Zeit- schr. 1861, p. 34.3.) — Nigro-cinerea, thoracis dorso albicante, capite, tho- racis vittft laterali scutelloque sulphureis, alarum albidarum stigmate fusco, terebrS, fbeminje atra. Blackish cinereous, thorax more whitish above, head, a lateral stripe on the thorax, and the scutellum sulphur-yellow, wings whitish, stigma fuscous, borer of 9 black. Long. corp. 0.17 — 0.20. Long. al. 0.18— 0.19. Antennae moderate, ferruginous-yellow, tip of the third joint rounded, bristle almost bare. Eyes large, almost round ; cheeks moderate; proboscis not geniculated, palpi short. Thorax dusted TRYPETIDAE. 101 with whitish above, with a short, whitish pubescence and black bristles; scutellum with four bristles. Abdomen with black hair. Borer of 9 flattened, black, with black hair ; a little longer than the three last joints of the abdomen taken together. Feet brown- ish-black, trochanters, knees, tip of the tibiae and the tarsi yellowish ferruginous. Wings whitish, stigma brown, all the veins, except the first longitudinal one, bare, pale yellow on the basal part of the wing, brown beyond it. Hub. Pennsylvania. (Osten-Sacken, on Vernonia, in August.) 2O. T. verilOlliae LOEW. £ and 9 . (Loew, Beii. Entom. Zeitscli. 1861, p. 346.) — Helva, metanoto nigro, capite, thoracis vitta laterali, superiors pleurarum parte et scutello pallidius, antennis, terebra, pedi- busque obscurius flavis ; alarum dimidio apicali fasciis tribus fuscis subreticulato, prinia inconipleta et obsoletiore, secunda integra, tertia postice abbreviata. Pale yellowish, metanotum black, head, lateral stripe of the thorax, upper part of the pleurae and scutellum pale yellow, antennae, borer and legs darker yellow ; apical half of the wing subreticulated with three brown bands, the first of which incomplete and less apparent, the second entire, the third abbreviated posteriorly. Long. corp. 0.18 — 0.22. Long. al. 0.17—0.18. Head yellow, orbit of the eyes narrow, with a silvery reflection. Antennae ochreous, third joint oblong, bristle almost bare. Oral opening rather large, rounded, proboscis not geniculated, palpi moderate. Eyes large, oblong. Thorax above clothed with a short, whitish pubescence, and with faintly brownish bristles. Scutellum flat, with four bristles. Metanotum black, dusted with whitish. Abdomen yellowish luteous (ex helvo luteum), with black hairs on the lateral margin and on the last segments ; remaining portion with yellow hairs. Borer of 9 ochraceous, shining, some- what flattened, equal to the three last abdominal segments taken together, clothed with soft, blackish hair. Legs fulvescent. Wings subhyaline, subreticulated with fuscous by means of three irregular transverse bands and some small apical spots ; the first band, which is much abbreviated posteriorly, starts from the infus- cated base of the stigma and runs obliquely towards the central transverse vein and frequently becomes obsolete, leaving, however, a brown cloud on the transverse vein ; the second band is narrow and straight, extending from the costa to the posterior margin ; the third band is unequal, abbreviated posteriorly, and coalescent 102 DIPTERA OF NORTH AMERICA. with the spots on the costa. First longitudinal vein beset with bristles, the others naked. Hub. Pennsylvania. (Osten-Sacken, on Vernonia, in August.) Note. — The first of the bands on the wings terminates posteriorly in a faint brown line, running along the discal cell and parallel to the longi- tudinal veins ; the second and third bands being more or less coalesceut, the space between the second and the tip of the wing may be described as brown, with five round, hyaline spots (one between the costa and the second longitudinal vein ; the second, just below the first, between the latter and the next vein ; the third between the same veins, but nearer to the tip of the wing ; the fourth on the fourth longitudinal vein, just behind the posterior transverse vein ; the fifth and largest at the tip of the wing, between the third and fourth longitudinal veins) ; second posterior cell hyaline, except a margin along the veins, which is clouded. The brown is more or less intense in different specimens, and hence the hyaline spots, especially the posterior ones, are sometimes less apparent. — 0. S. III. ON THE NORTH AMERICAN SCIOMYZIDfl. The family Sciomyzidce is principally based on the three genera: Sciomyza Fall., Telanocera Dum. and Sepedon Latr., all occurring in North America, and on the genus Thecomyia Perty. The attempt to subdivide the second of these genera has been made in various ways, but without success. I omit, therefore, to mention the genera thus formed, especially on account of the scantiness of my materials. Sciomyza has been also subdivided into smaller genera, namely: Graphomyza Macq., Pelidnoptera Rond., Gtenulus Rond. and Calobaea Zett., the last of which, differing from all the other Scio- myzidce. by its much smaller basal cells, may be considered as an osculant genus. Some of the species placed by Meigen in Scio- myza belong neither to that genus nor to the Sciomyzidce at all. The characters distinguishing the Sciomyzidce from all the other Acalyptera are as follows. The anterior frontal border more or less prominent ; face receding, proportionately long, with the oral border sharp ; no distinct furrows for the reception of the anten- na? ; no vibrissa? on the oral border ; the front with two bristles, one behind the other on each side before the lateral bristles of the vertex; the costal vein of the wings uninterrupted, without spine, reaching to the fourth longitudinal vein ; the auxiliary vein dis- tinctly separated from the first longitudinal vein on its whole length ; the two basal cells much developed, rather large, smaller only in the osculant genus Calobaea Zett. The legs have short hairs and very few bristles, and are of moderate length and rather stout, but not clumsy ; all the anterior legs, especially their tibia? and tarsi, are more developed than in the allied families ; all the tibia? on their outside before the tips have a small, erect, more or less distinct bristle ; the intermediate tibia? have a certain number of stout bristles at the tip ; the fore and hind tibia? have a single weak bristle. 104 DIPTERA OF NORTH AMERICA. Synopsis of the North American genera. J The third joint of the antennae not circular. I The third joint of the antennse circular. SCIOMTZA Fall, f Antennae only a little prolonged ; front not excavated ; eyes not pro- tuberant. TETANOCEKA Dum. 2 "j Antennse much prolonged ; front excavated ; eyes protuberant. SEPEDON Latr. Gen. I. SCIOMYZA FALL. Only three 1ST. A. species of this genus have been described, all by Mr. Walker. His description of Sciom. antica is made in so careless a manner that it is quite impossible to recognize it, and Sciom. parallela Walk, seems to be no Sciomyza at all. 8ci.om.yza nigripalpa Walk, is certainly not among the three species known to me. The scantiness of my present material scarcely warrants my undertaking to describe the N. A. species of Sciomyza, but I will present what I have to say about them, in connection with the Sciomyzidse generally. There is no reason for discussing here the smaller genera sepa- rated from the old genus Sciomyza, or to point out the subdivi- sions to be made, since the three species known to me belong all to the group of the typical species of Sciomyza. Synopsis of the Species.* l f Wings spotted. 1 nana Fall. I Wings not spotted, the transverse veins only clouded with brown. 2 9 J The two bristles on each side of the front extant. 2 obtusa Fall. \ The foremost of the lateral frontal bristles wanting. 3 pubera, n. sp. 1. S. il tvn a FALL. £ and £ .— Cinerea, thorace vittato, alis nigro-macu- latis. Gray, with the thorax striped, and the wings spotted with blackish. Long, corp. 0.1—0.13. Long. al. 0.11—0.14. SYN. Sciomyza nana FALLEN, Sciom. 15, 12.— MEIGEN, Syst. Beschr. VI, 18, 19. — ZETTERSTEDT, Dipt. Scand. V, 2109, 18. Quite agreeing with the European specimens. Ashy gray. Front opaque yellow, with the ocellar triangle and the lateral * The fourth species, added when the manuscript was already in press, is not included in this synopsis. — 0. S. SCIOMYZA. 105 stripes reaching ns far as the middle of the front, yellowish-gray. Antenna? yellowish-ferruginous, usually paler at the base, with the blackish-brown bristle beset with a short pubescence. Face whitish. Upper side of the thorax with four brown longitudinal lines, the two intermediate ones approximated and confluent with their hind ends, the two lateral ones narrower and less complete. Scutellum with a broad brown middle stripe. Pleura? brown, in the middle with a broad longitudinal stripe pollinose with yellow- ish, and a similar, but more indistinct longitudinal stripe more underneath. Abdomen brownish-gray, pollinose with paler on the lateral border, the posterior corners of the segments being whitish. Forelegs black, with the coxa? and the last joint of the tarsi whitish, and the extremity of the knees brownish-yellow. Middle and hind legs brownish-yellow, with the tips and upper side of the hind femora brownish black ; tips of the middle and hind tibia? black, the last joints of the middle and hind tarsi brownish. The dark color is sometimes more, sometimes less extended on the posterior legs than is described here. Wings hyaline, slightly grayish ; the costal border is margined with blackish, from the tip of the first as far as the tip of the second longitudinal vein ; from the end of this margin a blackish transverse band runs as far as the fourth longitudinal vein ; between it and the small transverse vein there are two small blackish spots ; the small transverse vein is clouded with blackish ; the posterior transverse vein is a little curved and marked with a larger blackish spot at its anterior end, and a smaller at its posterior end, both of which but rarely coalesce so as to form a complete margin. Hal. Middle States. (Osten-Sacken.) 2. S. olitusa FALL. £ . — Fusco-cinerea, antennarum seta plumata, venis alarum transversis fusco-liinbatis. Grayish-brown, the antennal bristle plumose, the transverse veins clouded with blackish-brown. Long. corp. 0.22. Long. al. 0.22. SYN. Sciomyza obtusa FALLEN, Sciom. 13, 4, var. a. — MEIGEN, Syst. Beschr. VI, 12, 6.— ZETTERSTEDT, Dipt. Scand. V, 2099, 10. I see no difference between the single N. A. individual I possess and that European species which is generally considered as the true Sciomyza obtusa Fall. But to prevent misunderstandings I must observe that there exists another species hitherto undescribed, differing from Sciom. obtusa Fall, by its antennas having a shorter 106 DIPTERA OF NORTH AMERICA. pectinated bristle, but otherwise resembling that species so much that it is commonly confounded with it. Grayish-brown. Front opaque yellow near its anterior border, remainder yellowish-ferru- ginous ; the ocellar triangle and the lateral stripes reaching beyond the middle of the front are yellowish-gray. Antennae yellowish-ferruginous, with the bristle dark brown, yellowish- brown at the base, and having black hairs of moderate length. Face yellow. Upper side of the thorax, with the exception of the lateral borders, more brownish than gray, with darker brown longitudinal lines, the two intermediate ones being darker and more distinct, the lateral ones doubled behind the suture. Scutellum flat, yellowish-brown, pollinose with grayish-yellow. Pleurae dark brown, with a broad, more chestnut brown lougitudinal stripe running from the shoulder to the base of the wing, and having underneath a hardly distinct longitudinal stripe formed by paler pollen. Legs rather dark brown, especially the anterior ones. Tips of the fore and hind tibiae black ; tarsi blackish towards the end. Wings grayish-brown, clouded with rather smoky brown near the costal border ; transverse veins clouded with blackish- brown ; the posterior transverse vein is slightly oblique and straight. Hab. Illinois. (Kennicott.) it 3. S. pilliera, LOEW. ^ — Fusco-cinerea, seta antennarurn breviter plumata, metatarso antico albo, veuis alarum trans versis fusco-limbatis. Grayish-brown, the antennal bristle brevi-plumose, the first joint of the anterior tarsi whitish, and the transverse veins clouded with blackish- brown. Long. corp. 0.21. Long. al. 0.21. Front opaque, sordid yellow near the anterior border, remainder more yellowish-ferruginous, with the ocellar triangle and the late- ral stripes brownish-gray, the latter reaching to the middle of the front, the foremost bristle wanting. Antennae ferruginous, bristle brown, with a short pubescence. Face pollinose with white. Upper side of the thorax grayish-brown, with but little distinct darker brown longitudinal lines. Scutellum flat, a little paler than the upper side of the thorax. Pleurae rather dark fyrown, pollinose with whitish, without distinct longitudinal stripes. Ground color of the abdomen almost brownish-black ; on each segment there is a large triangular, not pollinose, spot, its tip reaching as far as the hind border of the segment, the remainder TETANOCERA. 107 0 of the segment is covered with whitish pollen, which is much more dense on the hind border, and makes it appear quite pale, whereas on each side, near the lateral border, there is a vestige of a less pollinose, dark spot. The exterior genitals of the male are yellow- ish-brown. Legs almost blackish-brown, the intermediate ones, as well as all knees, part of the hind femora, ami a great part of the hind tibias, more yellow; anterior coxa3 yellowish, with a whitish reflection ; first joint of the fore tarsi whitish ; the follow- ing four black ; the intermediate tarsi have a rather pale brownish- yellow ground color, rendered much darker by their short black hairs, their two last joints appearing brownish on the upper side ; the hind tarsi are like the intermediate ones, but have the three last joints blackish. Wings grayish-hyaline, with the transverse veins margined with blackish, the posterior transverse vein straight and quite perpendicular. Besides the want of the foremost lateral bristle on the front, this species is distinguished from the other species of Sciomyza by its abdomen having the black hairs denser, longer, and finer than those. Rob. Middle States. (Osten-Sacken.) 4. S. llictifera LOEW. ^ . (Translated from Berl. Entom. Zeitschr. 1861, p. 345, by Bar. 0. Sacken.) — Nigra, thorace cinereo, facie alba, antennis et fronte fulvis, coxis anticis albis, pedibus nigris, tarsis poste- rioribus sordide albis, alis nigrieantibus, margine costali nigro-liiubato. Black, tborax cinereous, face white, antennae and front fulvous, fore coxae white, feet black, posterior tarsi of a dirty whitish, wings tinged with blackish, costal border margined with black. Long. corp. 0.13. Long, al. 0.1. Small, black. Face, cheeks, and the inferior part of the occi- put, white. Antenna and front fulvous, lateral stripes of the latter abbreviated anteriorly ; ocellar triangle and the upper part of the occiput cinereous. Thorax dark cinereous, with black hair. Fore legs black, their coxaa white, with a silvery reflection; posterior feet black, trochanters and tarsi dirty whitish, their apex black. Wings of moderate size, tinged with blackish, with a rather broad black margin near the costa. Hob. Pennsylvania. (Osten-Sacken.) Note. — In younger specimens, the black color of the intermediate pair of feet is more or less brownish. 108 D1PTERA OP NORTH AMERICA. Gen. II. TETAWOCERA DUM. Among the recorded N. A. species of Tetanocera, Tet. boscii has been characterized so insufficiently by Rob. Desvoidy, that there is no possibility of identifying it. Tet. canadensis, described by Macquart, is also unknown to me. Tet. guttularis Wied. is mentioned by Macqnart as-a native of North America ; but I must consider this statement as a mistake, since the characters he gives do not agree with the description of Tet. guttularis Wied. ; but what, species he has taken for Tet. guttularis has not as yet been made out. As to the other described species, the following paper will give all the necessary information : — Synopsis of the Species.* -j _[ Wings reticulated. 2 I Wings not reticulated. 11 2 f Bristle of the antennae plumose with black. 3 I Bristle of the antennae plumose with white. 6 Q f Front with three shining stripes. 4 I Front without shining stripes. 5 i Posterior transverse vein oblique and curved. 1 clara, n. sp. Posterior transverse vein almost perpendicular and straight. 2 valida, n. sp. 5 I Femora spotted. 3 pictipes Loeic. \ Femora quite unspotted. 4 pallida Loeiv. P f Thorax without stripes. 7 ( Thorax with stripes. 8 Femora entirely yellow. 5 flavescens Loew. Femora very brownish at the base. 6 arcuata Loew. f The brown margin of the costal border of the wings interrupted by clear 8 •! spots. 9 I Costal border of the wings with an uninterrupted brown margin. ' 10 !The reticulation of the wings shows double bands consisting of spots arranged by pairs. 7 combinata Loew. The reticulation forms no double bands at all. 8 sparsa, n. sp. 10 f Posterior transverse vein straight. 9 costalis, n. sp. <- Posterior transverse vein much curved. 10 saratogensis Fitch. f Posterior transverse vein curved and rather steep. 11 plebeja, n. sp. 11 -j Posterior transverse vein curved in the shape of an S, and very ob- lique. 12 plumosa Loew. * The two species (Nos. 13 and 14), added when the manuscript was already in press, are not included in this synopsis. — 0. S. TETANOCERA. 100 1. T. clara LOEW. 9- — Seta anteimarum nigro-plumosa ; vittse in fronte Isevigatse tres ; alse parce et grosse reticulatse, vena transversa pos- teriore obliqua. Bristle of the antennae plumose with black, front with three shining stripes, reticulation of the wings sparse and coarse, posterior transverse vein oblique. Long. corp. 0.32. Long. al. 0.32. A beautiful large species. Pale yellow. Face white, not much receding. Palpi and proboscis whitish-yellow. Front rather dark ochreotis, with three very shining longitudinal stripes ; the middle one distinctly widened towards its anterior end, the lateral ones near the borders of the eyes and reaching only very little beyond the foremost frontal bristle, hence not much transgressing the middle of the front. Antenna? ochreous, the two first joints short, beset with black hair, the third a little longer than the two first taken together, moderately broad and only moderately pointed ; the black bristle with dense, very long, black hairs. Neither the lateral borders of the front, nor the yellow occiput have black spots. Thorax yellowish, with two brownish middle stripes sepa- rated by a broad line, and on each side with a more indistinct and less complete lateral stripe. Scutellura with brownish middle and yellowish borders. Pleura whitish-yellow, with a narrow brown longitudinal stripe on their superior border. Abdomen without distinct markings. Legs whitish-yellow, the end of the tarsi only a little blackish ; posterior femora with a short, not very -close pubescence on their under side, and only one or two longer black bristles on the second third. Wings large and rather broad, some- what tinged with tawny ; the whole stigma and the broad clouds of the small and of the posterior transverse veins brownish-black; also the tip of the wing margined with brownish-black ; before the second longitudinal vein there are some small, rather indistinct, brownish-black spots, and about four or six larger and darker transverse spots between the second and third longitudinal veins, running from vein to vein, the last of which are most distinct, and include small rudiments of veins rising from the second longitudinal vein ; between the third and fourth longitudinal veins there are, beyond the small transverse vein, two or three brownish-black transverse streaks running from vein to vein; on the posterior side of the fourth longitudinal vein there are only two very small brownish-black spots, one before, the other behind the small transverse vein, which, I suppose, are not always present. The 110 DIPTERA Or NORTH AMERICA. remainder of the wings is unspotted. The posterior transverse vein is oblique and moderately curved. Hob. Trenton Falls, JX". Y. (Osten-Sacken.) 2. T. valitla LOEW. 9 • — Seta, antennarum iiigro-plumosa ; vittre fron- tales tres Irevigatae ; aL-e parce et grosse reticulatfe, vena transversa poste- riore perpendiculari. Bristle of the antennae plumose with black, front with three shining stripes, reticulation of the wings sparse and coarse ; posterior.transverse vein almost perpendicular. Long. corp. 0.29. Long. al. 0.29. Pale yellow. Face yellowish-white, only moderately receding ; palpi and proboscis whitish-yellow. Front orange-yellow, with three very bright longitudinal stripes; the middle stripe not dis- tinctly widened towards its anterior end, the lateral ones reaching a little beyond the foremost frontal bristle, which is inserted rather lower than in the preceding species, so that the lateral stripes reach a little nearer to the anterior border of the front. Antennas ochreous, having the two first joints short, with black hairs ; the third a little longer than the two first taken together, moderately broad and only a little pointed ; the bristle of the antenna? with dense, very long, black hairs. There are no black spots on the lateral border of the front, nor on the yellow occiput. Thorax yellowish, with indistinct brownish longitudinal stripes. Pleura? whitish-yellow, with a narrow brown longitudinal stripe on their superior border. Abdomen without distinct markings. Legs whitish-yellow, with the tips of the tarsi a little blackish ; poste- rior femora with short, not very dense hairs on the under side and only two longer bristles on the second third. Wings proportion- ately a little smaller than in the foregoing species, somewhat tinged with tawny. The stigma, smaller than in the foregoing species, is brownish-black ; the transverse veins and the tip of the wings are clouded with brownish-black ; before the second longitudinal vein there are six or eight small but distinct brownish-black spots; be- tween the second and third longitudinal veins there are four or five darker ones running from vein to vein, the last of which are more dis- tinct; between the third and fourth longitudinal veins there are, be- hind the small transverse vein, four or five brownish-black transverse streaks ; at the posterior side of the fourth longitudinal vein there are two rather large brownish-black spots, one before, the other, larger one behind the posterior transverse vein ; on the anterior TETANOCERA. Ill and posterior sides of the fifth longitudinal vein there are some small brownish-black alternating spots ; the remainder of the wing is unspotted; the posterior transverse vein straight and rather perpendicular. Observation. — Although this species is very similar to the fore- going, and I have only a single individual before me, its specific distinctness seems to be beyond doubt. The straight and rather steep posterior transverse vein, the lateral frontal stripes reaching farther forwards and the middle frontal stripe not being dilated anteriorly afford the best characters for distinguishing Tet. valida from Tet. clara. 3. T. pictipes LOEW. ^ and £. — Seta antennarum nigro-plumosa ; vittffi frontales Isevigatse nullse ; ala? confertim guttato-reticulatse ; femora maculata. Bristle of the antennae plumose with black, front without shining stripes ; wings densely reticulated with confluent fuscous spots and limpid drops, femora spotted. Long. corp. 0.21—0.26. Long. al. 0.23—0.26. SYN. Tctanoccra pictipes LOEW, Wien. Ent. Monatsch. Ill, 292. Front almost more pale brownish than yellow, opaque, beset, on its anterior part, with sparse short black hair, rising from hardly visible dark dots ; hardly a trace of an excavated middle stripe ; all that gloss which other species possess is totally wanting, and only a fine whitish dusted longitudinal line is visible. Each side of the front near the border of the eye a brownish-black dot, and more forwards between the antenna? and the anterior corner of the eye another small brown or blackish-brown spot. The face is silvery white, and recedes only moderately; its middle is marked with a very small black spot ; on the cheeks there is a brown or blackish-brown longitudinal streak. The first and second joints of the antennae are yellowish-brown ; the third is more yellowish-fer- ruginous, scarcely longer than the second, not much pointed, its upper side being distinctly excised; the black bristle has a brown- ish-yellow base, and a scanty, rather long black pubescence ; some individuals differ by the color of the antennae being quite ferrugi- nous brown. The upperside of the thorax is quite opaque, brown- ish cinereous, closely covered with small dark brown dots, which coalesce to larger spots, forming four rows, and having a rather variable size. Scutellum gray, in the middle brown, with small dark brown dots, on the borders with four black dots bearing the ordinary bristles, the hindmost of which are far longer than 112 DIPTERA OF NORTH AMERICA. the others. The ground color of the abdomen is more blackish than that of the thorax, the posterior and the lateral borders of the segments being usually more brown ; besides, the abdomen is covered with a rather light dust and beset with small brown dots coalescing near the lateral border into a row of obsolete spots, and in the middle of each segment into a longitudinal spot, so that a dark middle stripe, interrupted by the incisions, is formed. The sixth segment, being clavate in the male, has a large blackish- brown spot on each side, leaving in the middle a grayish or whitish mark, resembling, as it were, a cup. The ground color of the femora is little visible, being covered with light dust and speckled with black dots ; immediately before the tip they are surrounded with an almost black more or less visible ring, and an almost con- colorous spot before this ring on the under side. The tibite are yellowish-brown, with the tips blackish- the tarsi have the same color as the tibiae, but are generally a little paler; usually the ante- rior ones have the three last joints, and sometimes a great part of the first, blackened, whereas in the remaining tarsi only the two last joints, or even the last alone, is blackish. The wings are more guttated than reticulated ; the color of the posterior part is more gray; immediately along the longitudinal veins, and in the neigh- borhood of the costa, it is much darker and almost brown. The largest drops, the color of which is almost white, are scattered over the posterior part of the wing ; on the anterior part they are placed near the longitudinal veins ; on the costa, between the tips of the first and second longitudinal veins, there are only three small clear quadrangular spots. Hal). Washington. (Osten-Sacken.) Observation. — A series of specimens enables me to compare this species with the closely allied European Tet. umlrarum Linn. The resemblance of both is so great that I cannot but suspect that they are identical. ]STo difference of structure existing between them, the larger size and browner color of Tet. pictipes alone afford a constant distinguishing character. Future observations will per- haps enable us to decide whether Tet. pictipes is merely a climatic variety of Tet. nmbrarum or a different species. Note. — I possess .a specimen from Great Slave Lake, H. B. T., and have seen another from Maine, both perfectly agreeing in size and color with the European specimens. Is this fact to be considered as a proof of the identity or of the diversity of T. pictipes and T. umbrarum? The answer to this question appears to me far from certain. — 0. S. TETANOCERA. 113 4. T. pallida LOEW. ^ and 9 . — Seta antennarum iiigro-plumata ; vittre frontales Isevigatse nulhe ; alje confertim guttato-reticulatre ; fe- mora iiumaculata. Bristle of the antennae plumose with Mack, front without shining stripes, wings rather densely reticulated with dark spots and limpid drops, femora quite unspotted. Long. corp. 0.29. Long. al. 0.27. SYN. Tetanocera pallida LOEW, Wien. Ent. Monatsch. Ill, 294. Yellowish-brown, opaque. Front more yellow, opaque, without black spot near the orbit, the excavated middle stripe very narrow, not glossy ; between the antennae and the anterior corner of the eyes there is a small brown spot. Face white, considerably reced- ing, excavated in its middle more than in most other species. An- tenna? yellow with the third joint hardly as long as the second, its superior edge not distinctly excised, and its end very little pointed ; the antennal bristle blackish, with the base only yellow, and the blackish hairs rather long, but not very close. Upper side of the thorax not punctured, with four complete brown longitudinal stripes, and on its posterior half immediately beside the lateral stripe, a fine, less distinct, brown, longitudinal line. Scutellum with a brown middle stripe. Pleurae with a broad brown longitu- dinal stripe at the superior border, the remainder being everywhere whitish hoary. Abdomen unicolorous, with a blackish middle line, and on each side a brown linear stripe, all of them interrupted at the incisions. Legs yellowish, with the tips of the tarsi a little blackish. Wings somewhat yellowish towards the base with the reticulation moderately close and rather guttated, darker brownish- black at the costal and apical border; before the second longitudi- nal vein there are about six small clear dots, which do not reach the costal border itself; the small transverse vein is some distance before the middle of the discal cell, and the posterior transverse vein is very distinctly curved. Hob. Middle States. (Osten-Sacken.) 5. T. flavescens LOEW. £ . — Seta antennarum albo-plumosa ; thorax punctulatus; alje confertim guttato-reticulatte ; femora tota pallide fla- vescentia. Bristle of the antennse plumose with white, thorax punctured, wings densely 8 114 DIPTERA OF NORTH AMERICA. reticulated with dark spots and limpid drops, femora quite yellowish. Long. corp. 0.33. Long. al. 0.26. SYS. Tetanocera flavescens LOEW, Stett. Ent. Zeit. VIII, 123. — LOEW, Wien. But. Monatsch. Ill, 291. Pale yellowish-brown, somewhat shaded into testaceous. Face white, rather considerably receding beneath. Antennae yellow ; the third joint, when viewed sideways, nearly as long as the broad second joint, not distinctly excised on its upper side, little pointed; the antenna! bristle yellowish with close, white plumation of mode- rate length. Front yellow, opaque, with the excavated, polished middle stripe distinctly tapering anteriorly ; on each side of the orbit there is an oblong oval black spot of rather considerable size and another more anteriorly, between the antenna? and the anterior corner of the eyes. Upper side of the thorax marked with close small brown dots and besides with four rather incomplete rows of small dark brown spots far distant from each other. Scutellum rather convex, glossy, almost blackish-brown, a little dusted with whitish near the base, and having a terminal dot formed of whitish dust. Pleura? with a conspicuous, parallel, brownish-red longitu- dinal stripe at the superior border, below which they appear paler from their whitish dust. Abdomen with a dark middle line and near each lateral border a broad, brown one, all of them inter- rupted at the incisions ; the fourth and fifth segments bear each a glossy yellowish-brown- spot more distant from the border than the lateral lines. Legs brownish-yellow with the tips of the tibia? and the whole of the tarsi appearing darker in consequence of the greater density of the black hair, whereas in reality the two last joints only of the anterior and posterior tarsi are blackened ; the under side of the posterior femora is beset with very numerous short and many longer black bristles. Wings rather broad and obtuse, with the whole surface coarsely and rather uniformly reti- culated, so that there are no fascia? ; some larger brown spots on the costal border, but no clearer spots on the apical border ; the small transverse vein is very far from the discal cell, and the pos- terior transverse vein is only little curved and rather steep. Hab. Carolina. (Zimmerman.) Observation. — When naming this species I overlooked the fact that Rob. Desvoidy already has a Tet. flavescens. Consequently I should have altered the name, had I not before me a larger number of specimens of Tet. arcuata proving that this species is rather vari- TETANOCERA. 115 able, and that therefore its distinctness from Tet. flavescens is not quite certain. Should the identity of both species be proved, the name of "fawescens" as being preoccupied must be dropped, and that of Tet. arcuata adopted for the species ; should, however, future observations prove their distinctness, it will then be time enough to choose another name for Tet. flavescens. 6. T. arcuata LOEW. £ and 9 • — Seta antennarum albo-plumosa ; thorax punctulatus ; alse confertim guttato-reticulatse ; femorae antica basim versus fusca. Bristle of the antennre plumose with white ; thorax punctured, wings densely reticulated with dark spots and limpid drops ; anterior femora brown towards the base. Long. corp. 0.18 — 0.3. Long. al. 0.2 — 0.23. SYN. Tetanoccra arcuata LOEW, Wien. Ent. Monatsch. Ill, 292. So similar to the foregoing that it is very easy to confound them, and after the detailed description which I have given of Tet. flavescens, it will be quite sufficient to point out the characters by which Tet. arcuata. differs from it. It is always a little smaller, sometimes much smaller than Tet. flavescens ; its second antennal joint, too, seems to be comparatively smaller and narrower, and the small brown spots on the upper side of the thorax beside the small dots, are distinctly smaller. The anterior femora are to a consider- able extent brown at the base ; their tips as well as the base of the posterior femora are not seldom very brownish ; the anterior tibia? are blackened to a certain extent, and the posterior tibias have this color at their very tips ; the three last joints of the anterior tarsi are blackish. Hub. Middle States. (Osten-Sacken.) Observation. — The more specimens of Tet. arcuata I was able to examine, the more it appeared doubtful to me whether Tet. fla- vescens ought not to be taken merely for an exceedingly large and pale variety of Tet. arcuata. The only difference existing in the structure is, as it seems, the somewhat smaller breadth of the second joint of the antenute in Tet. arcuata; this is a very trifling one, and perhaps only a character belonging to smaller specimens. The narrow arcuated band running over the posterior transverse vein of the wings, by which the first specimen which I received was distinguished, was either more indistinct or quite wanting in the specimens sent to me afterwards. 116 DIPTERA OF NORTH AMERICA. "7. T. comliiiiaf a LOEW. £ . — Seta antennarum albo-plumosa ; thorax vittatus ; alse maculato-reticulatse, maculis fascias duplicatas efficienti- bus, limbo marginis antici obscuro nullo. Bristle of the antennae plumose with white, thorax striped, wings reticu- lated with dark spots forming double bands, costal border without dark margin. Long. corp. 0.26. Long. al. 0.22. SYN. Tetanocera combinata LOEW, Wien. Ent. Monatsch. Ill, 295. Reddish-brown, more acorn-colored on the abdomen. Front dark yellow with the middle stripe broad, impressed, polished, narrowed anteriorly, a small black dot being on each side near the orbit, and a second more anteriorly between the antennae and anterior corner of the eye. The face white, not much receding, rather ex- cavated in the middle. Antenna? yellow, third joint short, with the upper edge margined with blackish and not distinctly excised ; the antenual bristle yellowish at the base, with a white plumation of moderate length. Upper side of the thorax almost brownish- ferruginous, with broad lateral borders dusted with whitish ; the two longitudinal stripes in the middle are blackish and covered with dense white dust. The scutellum is of the same color as the upper side of the thorax, and has the sides dusted with whitish. The pleura? too resemble the upper side of the thorax in their color, appearing however, with the exception of a longitudinal stripe on their superior part, of a paler shade, on account of their whitish dust. Immediately before the poisers there is a very conspicuous, rounded, brownish-black spot. The abdomen is more acorn-colored ; it has a black middle stripe not sharply bordered and interrupted at the incisions, and a rather broad, polished, brownish-red stripe at some distance from each lateral border. Legs brownish-yellow, the tibia? a little darker than the femora ; the tarsi blackish towards their tip. Wings rather yellowish, having the reticulation formed by narrow, gray stripes in the middle of the intervals and by brown spots reaching from the longitudinal veins as far as these stripes; the brown spots are arranged so as to form distinct double bands running across the wing ; the last of these bands is at the very tip of the wing, the penultimate runs between it and the pos- terior transverse vein ; the antepenultimate runs over the posterior transverse vein itself; there is besides, anteriorly, the beginning of a double band before the penultimate band ; the small transverse TETANOCERA. 117 vein is a little before the middle of the cliscal cell ; the posterior transverse vein is a little curved and rather steep. Hub. Middle States. (Osteu-Sacken.) 8. T. sparsa LOEW. £ and 9 • — Seta antennarum albo-plumosa ; thorax vittatus ; alse maculato-reticulatse, fasciis duplicatis nullis, limbo marginis antici obscuro nullo. Bristle of the antennae plumose with white, thorax striped, wings reticu- lated with dark spots forming no double bands, costal border without dark margin. Long. corp. 0.24. Long. al. 0.22. Yellowish-brown, opaque. Front dark yellow, with the middle stripe broad, impressed, polished, not narrowed anteriorly ; a small black clot is on each side in the neighborhood of the orbit, and a second is more anteriorly between the antennae and the anterior corner of the eye. Face white, not much receding, rather exca- vated in the middle. Antennae yellow ; the third joint nearly as long as the second, a little excised on the upper side, rather pointed ; the bristle of the antennae with the base yellow, its pubescence whitish. Upper side of the thorax with two brown middle stripes connected posteriorly and separated anteriorly, and not reaching the anterior border of the thorax; two broader lateral stripes are of the same color, but not so distinct. The scutellum also is of the colour of the thorax, but dusted with whitish on the sides. Pleurae paler than the upper side of the thorax, having, towards their superior border, a brownish-red longitudinal stripe continued as far as below the poisers. The abdomen has a very indistinct, dark middle line interrupted at the incisions ; on each side, at a distance from the lateral border, a brighter stripe not differing sensibly in color from the general color of the abdomen. Legs pale yellowish; tibiae not darker than the femora; tarsi blackish towards their tips. Wings only little yellowish; the reticulation is formed by very narrow, gray stripes running in the middle of the intervals, and by brown spots reaching from the longitudinal veins as far as the stripes; the brown spots are arranged so as to form no double bands, showing only the anterior indistinct begin- nings of some narrow simple bands, the number of which is three, besides the narrow margin of the tip of the wing. The small trans- verse vein is a little before the middle of the discal cell ; the pos- terior transverse vein is only very little curved and rather steep. Hub. Middle States. (Osten-Sacken.) 118 DIPTERA OF NORTH AMERICA. Observation. — Tet. sparsa differs too much from Tet. combinat'a in the markings of the wings to be considered as identical with it. I have, however, to remind, that they agree much more in their struc- ture than is usual in nearly-related species of this genus, and that my opinion, formed on very scanty materials (1 9 of Tet. combi- nata, 1 £ and 1 9 of Tet. sparsa), cannot but have a very second- ary weight. Nor will the difference in the markings of the thorax, however striking they may seem, solve the question, since, in some- what immature specimens, they always appear inconstant, and are often variable even in quite mature ones. 9. T. COStalis LOEW. £, . — Seta antermaram albo-plumosa ; thorax vittatus ; alae grosse inaculato-reticulatse, margine antico auguste nigro- limbato, vena transversa posteriore perpendicular!, recta. Bristle of the antennse plumose with white, thorax striped, wings reticu- lated with coarse dark spots and the costal border having a narrow black margin, posterior transverse vein perpendicular and straight. Long, corp. 0.17. Long. al. 0.17. Yellowish-brown, somewhat tinged with reddish, opaque. Front yellow, with the middle stripe broad, excavated, and polished, not tapering anteriorly ; on each side near the orbit there is a small black dot, and more anteriorly a second larger one between the antennse and the anterior corner of the eye. Antennse yellow, with the third joint a little longer than the second, but little excised on the upper side, not much pointed. Antennal bristle with the base only yellow and covered with a close white pubescence. Face white, rather considerably receding. Upper side of the thorax dusted with whitish on the lateral borders; in the middle, there are two complete longitudinal stripes, formed of whitish dust and bordered with brownish streaks, which are not quite distinct and interrupted in the middle of the thorax. Superior part of the pleurse with a brownish-red longitudinal stripe continued to beneath the poisers. Scutellum yellowish. Abdomen unicolorous, having no trace of darker stripes in the described specimen. Legs whitish-yellow, not distinctly blackened at the end of the tarsi; under side of the posterior femora with small short bristles. Surface of the wings a little yellowish, with the stigma and a margin of the costal border black ; this margin runs from the stigma as far as the fourth longi- tudinal vein, being very narrow as far as the tip of the second longitudinal vein, and then broader; there are about six or seven TETANOCERA. 119 small clear spots on the anterior side of the second longitudinal vein ; the remainder of the coarse reticulation is little connected, and formed of rather sparse small blackish spots; the small trans- verse vein is a little before the middle of the discal cell ; the pos- terior transverse vein is distinguished by its being perfectly straight and perpendicular. ffabt Illinois. (Ostcu-Sacken.) 1O. T. saratogensis FITCH. £ and 9 . — Seta antennaruni albo-plu- mata ; thorax vittatus ; alse confertim striato-reticnlatae inargine antico late fusco-limbato, vena transversa posteriore flexuosa. Bristle of the anteunse plumose with white, thorax striped, wings striped with gray and reticulated with darker dots, costal border having a broad, blackish-brown margin, posterior transverse vein undulating. Long, corp. 0.2—0.22. Long. al. 0.18—0.20. SYN. Tetanocera saratogensis FITCH, Report I, 68. — LOEW, Wien. Ent. Mo- natsch. Ill, 296. This species, which is readily distinguished from all the N". A. species known to nie by the costal border of its wings having a broad blackish- brown margin reaching as far as the second longitu- dinal vein, has been accurately described by Dr. A. Fitch in>the above quoted place. It strikingly resembles the European Tet. pratorum Fall. After examining a great number of specimens of the two species, I found it impossible to discover any constant differ- ence in their structure; but the brownish-black color of the costal border of the wings as well as the spots along the longitudinal veins in all the N. A. specimens reach farther towards the base of the wing than is the case in any European one, besides, in the former, the color of the club-like male genitals is very dark, whereas in the latter it is much lighter and almost yellowish. Whether the two hoary stripes of the thorax really are more distant in Tet. saratogensis, as they seem to be, I dare not pronounce with cer- tainty, since some specimens from Northern Europe approach in this respect the American ones. Moreover the space between those stripes in N. A. specimens as well as in European ones, is sometimes altogether yellow, sometimes bordered with distinct brown longitudinal lines, sometimes entirely browf. Hob. Middle States. (Osten-Sacken.) Mr. Loew referring to Dr. Fitch's description, without giving one of his own, I reproduce the former here. — 0. S. 120 DIPTER A OP NORTH AMERICA. The head above is golden yellow with two small rusty stripes on its fore part, a black spot at base and dot each side anteriorly, almost in contact with the eye, and a second one, also black, on the anterior margin, between the eye and the antennae. Face silvery white. Antennae light yellow, second joint longer than broad, with fine short black bristles along its upper and under edge ; third joint tinged with brown, narrow and curved, its upper side being concave, its lower side convex, and nearly parallel with the upper side, but slightly narrowed towards the apex, which is rounded ; seta yellowish white, plumose. Thorax pale dull yellow, with a faint darker stripe each side of the middle, which stripes have an ash gray reflection when viewed from the front ; clothed with a short black beard and a few long black bristles. Scutel ash gray with two nearly erect black bristles each side. Poisers yellowish white. Abdomen dusky, clothed with a short black beard, hind edges of the segments pale dull yellow. Legs pale yellow, with a fine black beard, and the spine-like bristles at the end of the shanks^black. Wings iridescent, smoky brown on the outer and apical margins, hyaline towards the axilla, the space between divided into niimerous square hyaline spots by dusky longitudi- nal stripes, one stripe being placed in the middle of each cell and sending short transverse branches to the veins at regular intervals ; veins and veinlets black. 11. T. pletoeja LOEW. •£, and £. — Seta antennarmn nigro-plumosa ; alee noil reticulatse, vena transversa posteriore modice arcuata, subper- pendiculari. Bristle of the autennse plumose with black, wings not reticulated, with the posterior transverse vein moderately arcuated and nearly perpendi- cular. Long. corp. 0.28 — 0.29. Long. al. 0.3. Yery allied and similar to the following species, but certainly different. Acorn-colored. Front yellow ; the usual black spot near the orbit totally wanting. The excavated polished middle stripe of the front is not enlarged anteriorly ; the polished lateral stripes are very broad and glossy, yet do not extend far beyond the middle of the front. Antennas dark yellow, the third joint a little longer than the second, rather distinctly excised on its upper side ; antennal bristle black with long and very close black hairs. Face considerably retreating, yellowish with white reflection. Upper side of the thorax, scutellum and pleurte quite as in the following species. Abdomen brownish-yellow without darker middle line nor lateral strip«s in the pair I have before me ; but in well colored specimens they may exist. Legs as in Tet. plumosa. Wings brownish-yellow margined with blackish-brown on the whole of the costal border, beginning at the base of the stigma and being TETANOCERA. 121 extended at the apex to a little beyond the tip of the fourth longi- tudinal vein ; there are besides grayish stripes between the longi- tudinal veins; the transverse veins are margined with dark brown- ish-black ; the small transverse vein is in the middle of the discal cell ; the posterior transverse vein is distinctly curved, but not in. the shape of an S, and has a rather steep position. Hab. Middle States. (Osten-Sacken.) Observation 1. — This species is very similar to the European Tel. elata Fabr., but differs by its posterior transverse vein, which, although more arcuated, has a steeper position, by its antennae being a little broader and the black hair of its antennal bristle being much closer. Observation 2. — From Tet. plumosa this species not only differs by the form and situation of the posterior transverse vein, but particularly by the considerable breadth of the polished lateral stripes of the front, which in Tetanocera plumosa are exceedingly narrow. 12. T. plumosa LOEW. % and £ . — Seta antennarum nigro-plumosa ; alse non reticulatse, vena transversa posteriore biflexa et valde obliqua. Bristle of the antennae plumose with black, wings not reticulated, with, the posterior transverse vein biarcuated and very oblique. Long. corp. 0.31—0.39. Long. al. 0.28—0.35. SYN. Tetanocera vicina MACQUART, Dipt. exot. II, 3, 180, Tab. XXIV, fig. 7. Tetanocera plumosa LOEW, Stett. Ent. Zeit. VIII, 201. — LOEW, Wien. Ent. Zeit. Ill, 296. Tetanocera struthio WALKER, List of Dipt. IV, 1086. A species of the relationship of Tet. arrogans, elata, etc. Acorn-colored. Front yellow, with a small brown dot instead of the ordinary lateral black spots on each orbit ; the excavated pol- ished middle stripe of the front being of middle breadth, and but little enlarged anteriorly; the polished lateral stripes very narrow and obsolete. Antennas dark yellow ; the third joint as long as the second, rather distinctly excised on its upper side ; antennal bristle black with long and very close black hairs. Face rather* receding, yellowish with white reflection. Upper side of the thorax with the lateral border broad, dusted with white and the three central longi- tudinal stripes likewise covered with whitish dust and leaving be- tween them two complete narrow reddish-ferruginous stripes which unite on the hind border of the thorax and run over the scutellum. 122 DIPTERA OF NORTH AMERICA. Pleurae with a narrow ferruginous longitudinal stripe on the supe- rior border. Abdomen acorn-colored, with a darker middle stripe interrupted at the incisions. Legs brownish-yellow, tibias usually a little darker than the femora, tarsi blackened towards the tips. Wings brownish-yellow, margined with blackish-brown on the whole of the costal border, beginning from the base of the stigma and ending a little beyond the tip of the fourth longitudinal vein ; moreover, there are usually dark gray stripes between the longitu- dinal veins ; the transverse veins are margined with dark brownish- black ; the small transverse vein is placed a little before the middle of the discal cell ; the posterior transverse vein is very arcuated in the shape of an S, and has a very oblique position ; in most spe- cimens the fourth longitudinal vein has some small stumps, most of which are emitted from its inner side, each being inclosed in a dot-like brownish black cloud. Such specimens resemble very much the European Tet. aratoria Fabr. Hob. Sitka. Middle States. (Osten-Sacken.) Observation. — The name of Tet. vicina is preoccupied by R. Desvoidy. 13. T. trianglllaris LOEW. J. (Translated from Berlin. Entom. Zeitsch. 1861, p. 344, by B. Osten-Sacken.) — Glandicolor, thorace bili- neato, frontis opacse vittulis lateralibus Isevigatis, subobsoletis, fossula media distincta, nou dilatata, subtriangula, antenuarum seta pilis longis, rarioribus, nigris plumata, alis lutescentibus, unicoloribus, venis trans- versis fusco-limbatis, posteriore subrecta. Acorn-colored, thorax with two lines, front opaque, its lateral stripes shining, almost obsolete, the furrow in the middle distinct, not dilated, subtriangular, anteunal bristle with long, rare, black hairs, wings lutes- cent, unicolorous, transverse veins margined with fuscous, the posterior one almost straight. Long. corp. 0.26 — 0.27. Long. al. 0.27. Occiput yellow, with a large shining-white spot. Front bright yellow, entirely opaque, the ordinary lateral stripes shining, narrow, almost obsolete ; the furrow in the middle distinct, shining, not dilated, very shortened, subtriangular. Antennas rather short, ochraceous, last joint a little longer than the two preceding taken together, ferruginous at the tip, bristle with long but rather rare, black hairs. Face shining-white, immaculate. Thorax above with two darker, very approximated, lines. Abdomen unicolored. TETANOCERA. 123 Legs luteous, three last joints of the fore tarsi, one of the inter- mediate ones, and two of the hind ones, blackish ; hind femora of the male beset below with black bristles. Wings lutescent, unico- lorous, fore and apical border not clouded, transverse veins mar- gined with brown, the posterior one almost straight. Nab. North Red River. English River. (R. Kennicott.) Observation. — Yery like Tet. sylvatica Meig., but still nearer allied to Tet. unicolor Loew, distinct from the former by the frontal furrow, which is very much abbreviated here and by the anterior margin which is not shining; from the latter, which it also exceeds in size, by the striped thorax, and the narrower and less obsolete frontal furrow. 14. T. rotimdicornisLoEW. % and 9. (Translated from the Berlin. Entom. Zeitschr. 1861, p. 344, by B. Osten-Sacken.)— Ex helvo glandi- color, frontis opacse vittulis lateralibus ordinariis sublsevigatis, distinctis, fossula media sequali usque ad anteriorem frontis inarginem producta, antennarum articulo tertio ovato, superius non excise, seta pilis nigris longis, sed raris plumata, alis lutescentibus, costse dimidio secundo et apice anguste fuscis, veuis trausversis fusco-limbatis, posteriore sub- recta. Pale yellowish acorn-colored, the ordinary lateral stripes of the opaque front rather shining, distinct, the intermediate furrow equal, extended to the anterior margin of the front, third joint of the antennae ovate, not excised superiorly, bristle with long black, but rare, hairs, wings lutes- cent, the posterior half of the costa and the apex with a narrow brown- ish cloud, transverse veins clouded with brownish, the posterior one almost straight. Long. corp. 0.26. Long. al. 0.22—0.26. Occiput yellow, with a heart-shaped shining-white spot. Front bright yellow, opaque, the ordinary lateral stripes shining, rather broad, the intermediate furrow shining, of equal breadth, entire. Antennae short, ochraceous, third joint ovate, not excised supe- riorly, the bristle with long, black, but sparse hair. Thorax above with four darker lines, the intermediate ones entire, the lateral ones interrupted, not seldom all four obsolete. Abdomen unicolor- ous with a rather obscure lateral vitta, which is generally obsolete in the male. Legs luteous, last joints of the tarsi black, hind femora of the male beset below with black spines. Wings lutes- cent, the posterior half of the costal border with a paler, the apical border with a more saturated infuscation, transverse veins clouded with fuscous, the posterior one nearly straight and perpendicular. 124 DIPTERA OP NORTH AMERICA. Hub. North Red River. English River. (Robt. Kenoicott.) Observation. — Stumps of veins proceed sometimes from the pos- terior side of the fourth longitudinal vein. Gen. III. SEPEBON LATE. All the described species of the genus Sepedon agree much in their structure and are very readily and sharply distinguished from those species of Tetanocera which approach them in the whole structure of the body, as, for instance, Tet. obliterata Fall, and gracilis Loew. The most striking difference consists in the form of the heads with the protuberant eyes, the excavated front and in the second antennal joint being very much prolonged and attenu- ated. Among the four ]S". A. species known to me three fully agree as to these characters with the described species, but the fourth considerably differs from them by having the second anten- nal joint, though much prolonged, not attenuated, but broad as in the species of the genus Tetanocera, while its head is in every respect that of a Sepedon, so that, if we will not form a new genus for it, it cannot by any means be placed in Tetanocera; I consider it as the type of a new group of Sepedon. Synopsis of the Species. , J Second joint of the antennae broad. 1 fuscipennis LOEW. I Second joint of the antennae narrow. 2 ( Hind femora slender and very much prolonged, with a brown ring be- o I fore the tip. 2 macropus WALK. j Hind femora rather much thickened, with no brown ring before the (. tip. 3 1 Under side of the hind femora in the male with a deep excision and two teeth. 3 armipes LOEW. Hind femora of both sexes simple. 4 pusillus LOEW. 1. S. fuscipennis LOEW. £ and 9 . — Rufo-brunneus, secundo an- teuuarum articulo'lato. Chestnut-brown, second joint of the antennae broad. Long. corp. 0.26 — 0.27. Long. al. 0.27. SYN. Sepedon fuscipennis LOEW, Wien. Eut. Monatsch. Ill, 299. Of a rather dark chestnut color. Head a little paler. Front with a very wide excavation and an oblong black spot on each side near the orbit, and a deep black dot-like one below each antenna SEPEDON. 125 and a little removed from the orbit. Antennae of the color of the head ; the second joint, although elongated as in the other species of Sepedon, is broad as in Tetanocera and with black hairs ; the third joint a little darker, rather acutely ovate, blackish at the tip. Antennal bristle white at the tip, and with a white pubescence, its two first joints and the base of the third being black. Upper side of the thorax in well-preserved specimens with a brown longitudinal stripe on each side ; its middle is fine whitish hoary, and marked with four darker lines not quite reaching the hind border of the thorax, the innermost being by far more distinct. The scutellum, too, the greatest portion of the pleura? and the coxae are whitish hoary. Abdomen rather glossy, with a very slight vestige of whitish hoar. Legs pale chestnut-brown, hind femora with the apical half darker, and the greatest part of the under side beset with rather scattered black spine-like bristles ; the anterior and posterior tibiae at the tips and the tarsi blackish, the middle tarsi chestnut-brown at the base. The very delicate black hairs on the upper side of the posterior tibise are much longer in the male than in the female, and in general longer than usual in the species of this genus. Wings clouded with dark smoky brown, more yellowish-brown towards the costal border; the transverse veins with narrow blackish margins ; the posterior transverse vein is considerably arcuated and has a very oblique position. Hub. Middle States. (Osten-Sacken.) 2. S. macropus WALK. £ . — Testaceus, antennarum articulo secundo tenui, femoribus posticis longissimis, gracilibus, fusco-anuulatis. Yellowish-red, second antennal joint narrow, posterior femora very long, slender, and marked with a brown ring. Long. corp. 0.3. Long. al. 0.35 lin. SYN. Sepedon macropus WALKER, List Dipt. IV, 1078. Yellowish-red, with the inferior portion of the pleura as well as the hips having a bright white reflection. Front without lateral spots. Face glossy ; below each antenna is a black spot distant from the orbit and surrounded with a white reflection on the pol- ished face ; another spot with bright white reflection runs from the under side of the eyes down the cheeks. Antennas yellowish- brown ; the second joint is slender and dark brown towards the 126 DIPTERA OF NORTH AMERICA. tip ; the third joint blackish with the antennal bristle beset with a very short pubescence. Thorax with a rather distinct reddish ferruginous middle stripe continuing over the scutellum (it may be more marked in better preserved specimens). The abdominal segments each with a browner margin of the hind border. Legs very elongated with very short spine-like hairs; the bristles on the under side of the very long, straight, not thickened hind femora are likewise very short; the hind tibiaB slender and almost straight. Color of the legs brownish-yellow, the tips of all femora being brown and the posterior femora having besides a brown ring on their last third ; the anterior and middle tibise have only brown tips ; the posterior tibias are quite brown with a very broad brownish-yellow ring before the tip; middle tarsi yellowish-brown, more blackish-brown towards the tip; hind tarsi quite dark brown. Wings clouded with brown ; posterior transverse vein slightly curved, not very steep. This description has been taken from a rather old specimen. Hob. Jamaica; (Walker.) Cuba; (Poey.) 3. S. armipes LOEW. £ and 9. — Brunnens, antennarum articulo secundo tenui, femoribus posticis incrassatis, in niare subtus profunde excisis et prope basim bidentatis. Brown, second antennal joint slender, hind femora thickened, in the male with a deep excision on the under side and two teeth near the base. Long. corp. 0.18. Long. al. 0.18. SYN. Sepcdon armipes LOEW, Wien. Ent. Monatsch. Ill, 298. Not dissimilar to Sep. spinipes, but darker and somewhat smaller. Head pale yellowish, with the front and upper part of the occiput brown, the former having a black spot on each side near the orbit and a black dot below each antenna a little removed from the orbit. The two first joints of the antennas brownish-yellow, the second attenuated as in most species of this genus; the third more or less lanceolate, black with the base only yellow. The first and second joints of the antennal bristle dark yellow, the third with the base blackish, the remainder being white with very short white pubescence. Upper side of the thorax finely hoary in the middle, on which some dark longitudinal lines are visible. Pleu- ra3 dusted with white. Abdomen rather glossy, pale chestnut- brown or almost yellowish-red at the tip. Legs brownish-yellow, the femora paler, especially towards the base; the hind femora are SEPEDON. 127 somewhat spotted with brown at the tip ; their under side has a rather deep excision, and immediately before this excision a coarse, almost two-headed, hook, and farther to the base a smaller obtuse tooth, the space between the excision and the tip being beset with short black bristles. The femora of the female are simple. The posterior tibiae of the male are much arcuated at the beginning of their last third, the two first thirds being almost quite straight ; the tibia? of the female are of a very similar struc- ture, but not quite so much curved, by which character it is most readily distinguished from the female of Sepedon pusillus, the pos- terior tibia? of which have a slighter and much more uniform arcuatiou. The anterior tibia? are gradually blackened towards the tip, and also the anterior tarsi are rather black, the second and third joints only being paler than the rest ; the middle tibia? show no trace of black, but the tips of the middle tarsi are blackish as well as those of the hind tarsi. Wings smoky gray, more yellowish-brown on the costal border; the transverse veins with narrow black clouds ; the posterior transverse vein is but little curved, and rather steep. Hob. Middle States. (Osten-Sacken.) 4. S. pusillus LOETV. £ and £ • — Brurmeus, anteunarum articulo se- cundo tenui, femoribus posticis incrassatis, in utroque sexu simplicibus. Brown, second antennal joint slender, femora thickened, simple in both sexes. Long. corp. 0.18. Long. al. 0.18. • SYN. Sepedon pusillus LOEW, Wien. Ent. Monatsch. Ill, 299. So very similar to the preceding species that a short enumera- tion of its differences will be sufficient to characterize it. The male is most readily distinguished from that of Sep. armipes by its hind femora being simple. The females of the two species are rather more difficult to separate ; the most certain difference is given in the form of the hind tibia?, which in Sep. pusillus are slightly and uniformly bent in their whole length, whereas in the female of Sep. armipes the arcuation is not only more considerable, but also affects chiefly the last third. All other differences are either uncertain or trifling; namely, the structure of the third joint of the antenna? does not afford any available character for distin- guishing the two species, in opposition to what I was inclined to suppose when I had only a few specimens of both of them. Hob. Middle States. (Osten-Sacken.) 128 DIPTERA OF NORTH AMERICA. Observation. — The genera Actora and Dryomyza, differing in some characters from the true Sdomyzidce, are not comprised in the above exposition. I have not seen the N. A. Actora, pub- lished by Mr. Walker as Actora ferruginea. The two N. A. spe- cies of Dryomyza I know, fall both into that section which is characterized by a hairy third longitudinal vein ; the first is, as it seems, identical with the European Dryomyza anilis Fall. ; the second, though resembling in its colors Dryomyza flaveola Fabr., may easily be identified by the subjoined description. Dryomyza simplex LOEW.— Pallide flava, nitens, fronte et an- tennis saturate flavis, opacis ; alse cinereo-liyalinse, vena longitudinal! tertia pilis longis vestita. Long. corp. 0.28. Long. al. 0.31. Polished, pale yellow. Front deeper yellow, opaque, with the short pubescence and the bristles black. Antennas concolorous with the front ; antennal bristle with rather long and black hairs. Wings limpid, with a distinct yellowish-gray tinge ; the third longitudinal vein beset with long hairs; the posterior transverse vein obsoletely clouded with grayish, the small transverse vein not clouded. Hob. Middle States. (Osteu-Sacken.) IY. ON THE NORTH AMERICAN EPHYDRINIDAE, The family of Ephydrinidce is taken here altogether in the extent which was given to it by Stenhammar, the diligent monographer of the Swedish species, and by Walker in his work on the British Diptera, the latter founded on Haliday's valuable observations. The characters easily distinguishing the EpJiydrimdce from all the other families of Diptera acalyptera may be set down as fol- lows: Face more or less, often considerably convex; either with- out any impression at all beneath the antennas, or moderately im- pressed, but never provided with membranous antennal furrows. Antennas short, first joint small ; antennal bristle either nearly bare, or pubescent, or pectinated on the upper side only. Oral cavity rounded, in most of the genera of considerable size; cly- peus distinct, in some genera retracted in the oral cavity, in the remaining genera prominent over the oral margin, in some of them of a rather large size; palpi small; mentum short, more or less in- crassated. Thorax rather quadrangular; scutellum proportionately large. Abdomen of very variable form, consisting of six seg- ments in the males; the females have one short, and generally not distinctly visible, segment more. The sixth segment being al- ways small and generally much concealed under the fifth, the struc- tural relations depend on the conformation of the five first seg- ments; of these the first is often much shortened and sometimes nearly connate with the second, a circumstance which has led authors to omit it in the enumeration or to count the two first segments for one; this is to be borne in mind in order to understand their descriptions; in mine, I have always counted the first seg- ment as distinct, however difficult it may be to observe it. The fifth segment is also of very variable structure, generally nearly equalling the foregoing in size, rarely considerably longer in both sexes or in the males, still more seldom much smaller, in which case the fourth segment, especially in the males, is longer than 9 130 DIPTERA OF NORTH AMERICA. usual. The hypopygium of the male, which attains a considerable size in some species only, is turned down, and generally, together with the small sixth segment, encompassed by the lateral border of the fifth abdominal segment bending down over it. The anal ex- tremity of the female, except in a few species, is entirely retracted ; for this reason, the sexual difference of many species is somewhat difficult to ascertain on examining single dry specimens. Wings on their whole surface covered with microscopical hair; the costal vein consists of three parts, the first of which reaches from the base to a little beyond the transverse humeral vein, which, quite in its neighborhood, runs over to the costa; the second from thence to the tip of the first longitudinal vein, where the third begins. These three parts of the costal vein are not to be confounded with the three segments of the costal vein so frequently used in charac- terizing the species; the latter are reckoned from the base of the wing to the tip of the first longitudinal vein, from theuce to the tip of the second longitudinal vein, and from this to the tip of the third longitudinal vein. The auxiliary vein is distinct only at its very base and then coalesces with the first longitudinal vein ; the second basal cell, i. e. the anterior of the two small basal cells, unites with the discoidal cell, the ordinary separating vein disappearing, so that the discoidal cell apparently reaches very far towards the base of the wings; it is not at all unusual, that on a closer examina- tion a rudiment of the obliterated transverse vein may be seen ; the posterior of the two small basal cells or anal cell is generally imperfect and very small. Alulae small. Legs slightly bristled ; in some genera the species have some longer bristles on the upper side of the intermediate tibias, which in all genera are provided with spurs. Most of the species, if not all, live in the neighborhood of water or in moist places. I have observed many years ago, that the food of several species chiefly consists of Infusoria. The larva? of those European species, the metamorphosis of which is known, live, at least by far most of them, in water, some exclusively in water which is very decidedly salt; two of them are leaf-miners. General division of the Ephydrinidae. In order to facilitate the determination of the already numerous genera of Ephydrinidce it will be useful to subdivide this family in several sections, which may be done as follows: — NOTIPHILINA. 131 Second joint of the anteunrc unguiculated at the end, or at least, the upper side of the intermediate tibiae beset with a few strong bris- 1 -I ties. I. NOTIPHILINA. Second joint of the antennae not unguiculated, the intermediate tibia? I never have long bristles on the upper side. 2 !Eyes hairy, oral cavity never of considerable size, anterior femora never incrassated. II. HYDRELLINA. Eyes naked, oral cavity generally very large. III. EPHYDRINA. The second joint of the antennae is called unguiculated (unguicu- laturri) if it bears on its end a bristle directed forward, whether it be thickened and long, or thin and short. In the latter case it is often difficult to perceive, particularly in the genera Paralimna and Corylhophora, which in the whole structure of their heads approach very much some genera of Ephydrina; yet the presence of several long bristles on the upper side of the intermediate tibiae and the color and markings of their abdomens point out too evidently their relation to the species of the widely spread genus Notipkila, to be overlooked. The hairy eyes will be sufficient in general to enable us to recognize the Hydrellina ; in those genera, the species of which have densely pilose eyes, the hairs are often exceedingly short; but their presence even then is easily known by the whitish reflection shown by the outline of the eyes. In the genera with scattered hairs on the eyes there are some spe- cies in which it is very difficult to perceive the single small hairs; in order to distinguish them with certainty from the Ephydrina, it is to be borne in mind that in the latter the eyes are much more rounded, that their faces are narrowest just where the antennaa are inserted, and considerably increase in breadth immediately below, whereas the Hydrellina have always more oblong eyes, and their faces have their least breadth beneath the antennae; the oral cavity also is never so strikingly wide as in most genera of Ephy- drina; moreover the clypeus in those species of Hydrellina which, on account of the indistinctness of the pubescence of the eyes, might be taken for Ephydrina, is very little developed. I. NOTIPHILINA. The second joint of the antenna) distinctly unguiculated, or the presence of some long bristles on the upper side of the interme- diate tibite will refer any species to the present section, the genera 132 DIPTERA OF NORTH AMERICA. of which have, without exception, an antenual bristle with long pectinations on the upper side. Division 1. The costal vein reaches to the third longitudinal vein, f Abdominal extremity of the male with elongated bristles. DICHAETA Meig. j Abdominal extremity of the male without elongated bristles. NOTIPHILA Fall. Division 2. The costal vein reaches to the fourth longitudinal vein. , ( Upper side of the intermediate tibiae with some long bristles. 2 I Upper side of the intermediate tibise without long bristles. 3 f Wings with a costal spine, posterior transverse vein perpendicular, legs not prolonged, clypeus very prominent. PARALIMNA Loew. 2 •! Wings without costal spine, posterior transverse vein oblique, legs pro- longed, clypeus hardly projecting beyond the oral margin. |^ CORYTHOPHORA Lo6W. Abdomen sharply edged, apparently three-jointed in both sexes on account of the minuteness of the first and fifth segments. TRIMERINA Macq. \ Abdomen not sharply edged, fifth segment not so strikingly short- (. ened, or only so in the males. 4 4 f Abdomen broad. DISCOMYZA Meig. \ Abdomen not broad. 5 {Superior half of the face not carinated, third joint of the antennae more or less oblong. PSILOPA Fall. Superior half of the face distinctly carinated. 6 SEyes oblong, cheeks not descending much beneath the eyes. DISCOCERINA Macq. Eyes rounded, cheeks descending very much beneath the eyes. 7 7 f Clypeus very prominent beyond the oral margin. ATHYROGLOSSA Loew. \ Clypeus projecting very little beyond the oral margin. HECAMEDE Hal. Of the enumerated genera I know Dichceta, Notiphila, Paralimna, Discomyza, Psilopa, andDiscocerina as occurring in North America. Gen. I. DICHAETA MEIG. This genus is closely related to the genus Notiphila. Both are distinguished by the remarkable stout spine of the second joint of the antennae ; the face is perpendicular and only moderately con- vex, the clypeus small and scarcely prominent beyond the border of the mouth; moreover, in both, the intermediate tibia? are beset on the upper side with some long and stout bristles, and the thick- ened costal vein terminates already at the tip of the third longi- tudinal vein. The characters distinguishing both genera from each DICHAETA. 133 other are as follows : The species of Dichceta have longer and stouter bristles ; the mystacidal bristles on the side of their face there are less numerous, but much longer and stouter; moreover, in the males of Dichceta the penultimate segment of the abdomen, on its posterior edge, is provided with a transverse row of very long bris- tles, and on the tip of the last segment, above the anal opening, there are two bristles inserted near each other and curved upwards, which is never the case in Notiphila. Only two European species of Dicli&la were hitherto known. The two species occurring in North America are identical with them. Synopsis of the Species. ( Last segment of the abdomen in the male prolonged in a conical point. j 1 caudata Fall. j Last segment of the abdomen in the male not prolonged in a conical point. 2 brevicauda Loew. 1. I>. caudata FALL. £ and £ . — Nigricans, segmento abdominal! penultimo in mare setis decem elongatis arrnato segmentoque ultimo in couum producto. Blackish, the penultimate segment of the male abdomen with ten long bristles, the last segment couically prolonged. Long. corp. 0.17. Long, al. 0.17. This well known species is distinguished from the following by its somewhat larger size, the greater number and the length of the bristles on the posterior border of the penultimate segment of the male abdomen, the conical prolongation of its last segment, and the much greater length of the two stout bristles inserted on the tip of this prolongation. Hob. Middle States. (Osten-Sacken.) Note. — A detailed description of this species is to be found in Meigen, Zweifl. VI, p. 62.— 0. S. 2. D. Tbrevicailda LOEW. £ . — Nigricans, segmento abdominal! pe- nultirno in mare setis sex elongatis armato segmentoque ultimo breviter acuminato. Blackish, the penultimate segment of the male abdomen with six long bris- ' ties on the posterior border, the last segment but little pointed. Long, corp. 0.16. Long. al. 0.16. The differences pointed out in the description of the foregoing species will be sufficient to distinguish this. The greater part of 134 DIPTERA OF NORTH AMERICA. the tarsi is dull red, as in Dick, caudata, but generally somewhat paler. Hab. Middle States. (Osten-Sacken.) Note. — This species has been described for the first time by Mr. Loew in 1860 in his paper: Die Europseischen Ephydriniden (in Loew's Neue Bei- traege, VII, p. 5).— 0. S. Gen. II. WOTIPHOLA FALL. The characters of this genus result from what has been said about Dich(Eta. Those which distinguish it most easily from the following genera are the intermediate tibiae being provided on the upper side with single long bristles, and the thickened costal vein terminating at the tip of the third longitudinal vein. We need scarcely mention that but few of the species described by the older authors as Notiphilce, belong to it, in the restricted sense necessary here. Taken in this sense, about sixteen European species have to be referred to it, some of which, however, are not yet duly established. In North America it seems to be repre- sented by numerous species which, compared to those of Europe, show nothing heterogeneous in their organization. The same observation may be made with regard to the South African spe- cies which occur on the Cape of Good Hope. Synopsis of the Species. , f Palpi blackish. 1 scalaris, n. sp. I Palpi yellowish. 2 0 f Antennae quite black. 2 bella, n. sp. i 1 Third joint of the antennre red at the base. r Upper side of the thorax with a broad brown lateral stripe. 3 \ 3 vittata, n. sp. <• Upper side of the thorax without broad brown lateral stripe. 4 f Abdomen with irregular brownish-black semifasciae, each formed of 4 ! two spots. 4 carinata, n. sp. Abdomen with two brown spots on each of the intermediate segments. 5 unicolor, u. sp. i 1. Hi. scalaris LOEW. % and £-. — Obscure ciuerea, facie augusta fulvescente, antennis palpisque nigris, abdomine fasciis nigris, linea longitudinali cinerea interruptis picto, pedibus nigris, geuibus tarsisque posticis testaceis, tarsis anticis testaceo-anmilatis. Dark ashy-gray, the small face yellow, antennas and palpi blackish ; ab- donieu with two transverse fasciae, interrupted by a gray middle line ; NOT1PHILA. 135 legs blackish, knees and hind tarsi yellowish, anterior tarsi aunulated with yellowish. Long. corp. 0.13. Long. al. 0.16. Very much resembling Notiph. uliginosa Hal. (which is identical with Notiph. tarsata Stenh.), but its narrower face distinguishes it from that and all the related European species. Palpi blackish. Antennae entirely black ; front with yellowish-brown on black ground. The same is the case with the upper side of the thorax, which has no broad longitudinal stripes, but only a faint trace, often indistinct, of five fine, brown longitudinal lines. The scutellura is colored as the upper side of the thorax, but generally with a rather lighter gray border and longitudinal line. Abdomen gray, with broad brownish-black fasciae occupying more than the anterior half of each segment, and being interrupted by a gray middle stripe ; the last segment in the male is almost entirely black, and has a gray middle stripe on its anterior half. Femora and tibia? black ; the knees and the extreme tips of the tibiae brownish-yellow ; the fore tarsi black, having the innermost base of each joint yellowish ; the posterior tarsi yellowish, with the tip brownish. Wings pel- lucid brownish-gray, with brown veins ; the second segment of the costal vein being nearly twice as long as the third. Halt. Middle States. (Osten-Sacken.) 2. IV. "bella LOEW. £ and 9 . — Cinerea, antennis totis nigris, palpis flavis, vitta thoracis utrinque laterali, pleurarum superiore, scutellique margine laterali hrunneis. Ashy-gray ; antennae entirely blackish, palpi yellow ; a longitudinal stripe on each side of the upper side of the thorax, a longitudinal stripe on the pleurse, and the lateral edge of the scutellum, brown. Long. corp. 0.14. Long. al. 0.17. Face yellowish. Eye-rings rather broad. Cheeks descending considerably beneath the eyes. Antennae entirely blackish. Front gray, viewed sideways rather whitish ; the divided black middle stripe is more or less covered with thick light-gray dust, which is sometimes of a yellowish tinge, sometimes more light-gray ; near the lateral border \_ofthe thorax*! — 0. £.] there is a broad, well-defined, dark-brown longitudinal stripe. Such a stripe runs on the upper part of the pleurae from the shoulder to near the base of the wing. The brown color of the lateral border of the scutellum continues on the posterior border of the thorax as a short beginning of a stripe. Abdomen, with four rows of long, triangular, blackish-brown spots, 136 DIPTERA OF NORTH AMERICA. the interior of which are a little longer that the exterior ones. Coxae and femora black, covered with light-gray hoar ; the tips of the latter brownish-yellow. Tibiae and tarsi rather pale brown- ish-yellow, the hind tibiae with a broad, the middle and fore tibiae with a narrow blackish-brown ring, which is sometimes wanting ; the last joint of the tarsi brownish. In the male, the middle femora, on the under side, are beset with short, but very thick, black hair, the middle tibiae on the under side fringed with very close, short, black pubescence. Wings grayish, proportionally long and narrow ; veins brown ; the second segment of the costa nearly thrice as long as the third. Hob. Middle States. (Osten-Sacken.) 3. IV. vittata LOEW. 9 • — Fusca, facie laete ochracea, antennarum articuli tertii basi sordide rufa ; thoracis vitta laterali, pleurarum supe- riore scutellique inargine laterali obscure brunneis, abdoiniue macularum nigrarum seriebus quatuor picto, fenioribus nigris, tibiis late nigro-auuu- latis. Brown ; face bright ochraceous, third joint of the antennse dull red at the base ; a longitudinal stripe on each side of the upper side of the thorax, a longitudinal stripe on the pleurae and the lateral border of the scutel- lum dark-brown ; abdomen with four rows of black spots ; femora black, tibiae with a broad black ring. Long. corp. 0.16. Long. al. 0.18. The most robust among the known North American species. Face of middle breadth, rather bright yellow. Palpi yellow. An- tennas black, third joint at its base dull red for a considerable dis- tance. Front, thorax, and scutellum dusted with brown. The upper side of the thorax has on each side, near the lateral border, a broad, well-defined, dark-brown stripe, and, moreover, on its middle, some much less distinct brown longitudinal lines. Pleura grayer than the upper side of the thorax, above with a broad, dark-brown, longitudinal stripe, running from the shoulder to the base of the wing, and another incomplete brown longitudinal stripe immediately above the longitudinal suture. Lateral border of the scutellum blackish-brown. Upper side of the abdomen dusted with gray and having four rows of black spots, those of the two interior rows being longer and more triangular, those of the exte- rior rather shorter and more trapezoidal. Femora black; tibia? and tarsi yellowish, the former with a broad brownish-black fascia, which, on the anterior tibiae, leaves only the base and tip free ; the tarsi, on account of tjieir hair, appear darker than they really are. NOTIPHILA. 137 Wings distinctly tinged with brown ; veins brown ; the second segment of the costal vein scarcely twice as long as the third. Hab. Middle States. (Osten-Sacken.) 4. W. carinata LOEW. 9 • — Cinerea, facie angusta concolore, antenna- rum articuli tertii basi rufa, thoracis dorso scutelloque brunneis, abdo- mine fasciis interrupts nigro-brunneis, postice ernargiuatis, picto, pedibus nigris, tibiis tarsisque posterioribus testaceis, tibiis posticis nigro-annu- latis. Ashy-gray, the narrow face ashy-gray ; base of the third joint of the an- tennae red ; upper side of the thorax and scutellurn brown ; abdomen with interrupted blackish-brown fasciae, which are emarginated poste- riorly ; legs black, middle and posterior tibiae and tarsi brownish-yellow ; hind tibiae with a black ring. Long. corp. 0.13. Long. al. 0.16. Face gray, rather narrow, indeed remarkably narrower and with a more extended and sharper keel on its upper part than in the similar European species Notiph. annulipes Stenh. and Notiph. dorsata Stenh. Palpi yellow, antennae black ; the third joint with the basal half yellowish-red. Front, upper side of the thorax, and scutellum, grayish-brown, or even almost brownish-yellow ; upper side of the thorax without lines or stripes. Pleura ashy-gray, brownish above. Abdomen on the basal half of each segment with two blackish-brown sernifascias, emarginated posteriorly, which on the last segment dissolve themselves more or less into the two spots composing them. Anterior legs entirely black, only the knees and the extreme tips of the tibiae being brownish-yellow. Middle and posterior tibiae and tarsi brownish-yellow; hind tibiae with a brownish-black band; tips of the tarsi brownish. Wings of a rather dull gray, veins brown ; the second segment of the costal vein a little more than twice as long as the third. Hab. Middle States. (Osten-Sacken.) 5. N. linicolor LOEW. £ . — Flavo-cinerea, facie laetius flava, abdo- minis segrnentis duobus intermediis brunneo-bimaculatis, femoribus nigris, genibus, tibiis tarsisque flavo-testaceis, anticis fuscanis. Yellowish-gray, face of a brighter yellow ; the two middle segments of the abdomen each with two brown spots ; femora black, knees, tibiae and tarsi brownish-yellow; the fore ones more brownish. Long. corp. 0.13. Long. al. 0.16. Entirely yellowish-gray. Face bright yellow, rather broad ; cheeks descending beneath the eyes a little more than usual. Palpi dark yellow. Third joint of the antennae with the basal half red- 138 DIPTERA OF NORTH AMERICA. dish-yellow. Thorax without lines or stripes. Abdomen very unicolorous, having only on each of the two middle segments two triangular brown spots of middle size ; of the two exterior rows of spots, which generally occur on the abdomens of the Notiphilte, nothing is to be seen here. Femora black, appearing gray in consequence of their being dusted, with yellowish tips. Tibia3 and tarsi brownish-yellow ; fore tibiae towards their tips and fore tarsi brownish on their whole extent ; the posterior tarsi only with their last joint brown. The short hair, resembling fringes, on the under side of the middle femora and tibite is rather thin. Wings rather sandy-yellowish, particularly at the base, the second segment of the costal vein is a little more than twice as long as the third. This species resembles most the European Noliph. guttiventris Stenh., but is easily distinguished from it by its smaller size and more yellow color, by the cheeks descending deeper be- neath the eyes and by the much less spotted abdomen. Gen. III. PARALIUftfA LOEW. The characters of this genus, of which I hitherto only know South African and North American species, are the following. Structure, colouring, and markings as in Notiphila. Eyes much rounded ; front and face very broad, the latter slightly convex ; eye-rings broad ; cheeks descending very deeply beneath the eyes ; clypeus prominent ; palpi narrow ; terminal bristle of the second joint of the antennas very small and hardly visible ; the third joint of the antennas very distinctly hairy on its upper side and tip ; the antennal bristle with long rays. Structure of the thorax, scutel- lum, and abdomen as in Notiphila. Middle tibia? on their upper side with three long bristles, the first being very near the base, the second immediately before the middle, and the third not far from the end. Wings as in Notiphila, only with the exception of the thickened costal vein being extended to the tip of the fourth lon- gitudinal vein. 1. P. appemliculata LOEW. £ and 9 . — Bruunea, fronte, thorace scutelloque obscurius punctatis ; facie fasciisque interruptis abdoininis nigri canis ; palpis, antennis pedibusque nigris, tarsorum anticorum posticorurnque basi rufa ; alis cinereis, venis transversis et venula ap- pendicea e penultimo venae quarta? segmento ascendente nigro-limbatis. Brown, front, thorax and scutellum with darker dots ; the face and the interrupted fascia? of the black abdomen are grayish-white ; palpi, an- PARALIMNA. 139 termse, and tarsi black, the fore and hind tarsi red at the base ; wings gray, the transverse veins and an additional veinlet being placed on the anterior side of the penultimate segment of the fourth longitudinal vein, are bordered with black. Long. corp. 0.16 — 0.18. Long. al. 0.16 —0.18. Face dusted with grayish white, usually with some more brown- ish spots, sometimes with a more yellowish-gray tinge ; it is slightly convex, but not even, eye-rings broad ; viewed laterally, they show at their upper end two black spots, united by a white transverse line, which disappear when viewed in other directions. The very projecting clypeus is of the same color as the face. Proboscis thick and black ; palpi narrow, rather long and black. Antenna black, the third joint in certain directions with a whitish-gray reflection; the hair on the upper side and the tip remarkably long. Front brown, anteriorly with some small dots, further back with some nearly black spots. Thorax and scutellum brownish, with numerous close, small, dark-brown dots ; pleura? also dotted. Ab- domen brownish-black, rather opaque, on the posterior border of each segment with a whitish-gray, very opaque fascia, a little widened on its middle, and intersected by a brownish-black middle stripe. The two halves of the gray fascia of the second segment are sometimes connected on their posterior margins ; the fifth abdominal segment of the male is a little longer than the preceding. Legs entirely black, the fifth joint of the fore and hind tarsi clothed with shining felt of a bright reddish-yellow ; the first joint of the fore tarsi at its base, and the first joint of the hind tarsi almost to its tip, are usually red ; paler specimens have also the first joint of the middle tarsi red ; in darker ones the first joint of the anterior tarsi is entirely black. Wings gray, veins brown ; the second half of the costal vein, the end of the third longitudinal vein, and nearly the whole fourth and fifth lon- gitudinal veins, rather black ; the transverse veins and a small stump, emitted by the fourth longitudinal vein about the middle of its penultimate segment, black and narrowly bordered with black ; the second segment of the costal vein more than twice as long as the third. Hal). Middle States. (Osten-Sacken.) Georgia. 140 DIPTERA OF NORTH AMERICA. Gen. IV. DISCOMYZA MEIG. The characters of this genus are as follows. Head more or less orbicular, with very sharp borders of the vertex ; second joint of the antennas unguiculated, the third oblong, with long pectinations of the terminal bristle. Face not keeled, rather convex, receding again towards the edge of the mouth, uneven, on the sides with coarse warts and wrinkled. Clypeus entirely concealed. Abdo- men flat, broad, on account of the shortening of the first segment apparently consisting of four rather equally broad segments. Wings proportionately broad, third and fourth longitudinal veins parallel at their ends. There were only two species hitherto known; the following North American species deviates a little by its head not being so strikingly orbicular, and by its abdomen not being so broad, but more flattened. 1. I>. l>alioptera LOEW. £. — Nigra, tliorace punctulato, antennis genibusque rufis, tarsis posterioribus flavescentibus, alis fusco-maculatis. Black, with dotted thorax ; antennae and knees yellowish-red, middle and posterior tarsi yellowish ; wings dotted with brownish-black. Long. corp. 0.15. Long. al. 0.14. Head shining black, really not so orbicular as in Discom. in- curva, but the vertical border likewise very sharp. Front ante- riorly with two rather flat depressions, placed near each other; the more shining lateral border of the front rather wrinkled. Antennae yellowish-red, the upper border of the second and third joints a little darker ; the antennal bristle with long pectinations. The middle of the face narrow and rather transversely wrinkled ; its lateral parts with coarse warty wrinkles ; the eyes surrounded with a fine white line. The upper side of the thorax and scutel- lum appear to be dusted with white, but have a rather indistinct, exceedingly fine and close punctuation, leaving only small traces of the white dust. On the pleurae, where the punctuation is more distinct and much coarser, the whitish dust is more visible. Abdo- men black, rather shining, exceedingly flat, narrower than in Dis- com. incurva, the cause of which may be that the upper horny plates of the abdomen are turned clown to an unusual extent ; the last segment of the abdomen rather smaller than the preceding PSILOPA. 141 ones. Legs black, knees yellowish-red; middle band of the posterior tarsi pale yellowish, having the last joint rather blackened. Poi- sers whitish with darker petiole. Wings short and broad, clouded with grayish ; the small transverse vein is below the tip of the first longitudinal vein ; the posterior transverse vein rather distant from the margin of the wing and rather oblique ; the two last segments of the fourth longitudinal vein of equal length ; the second seg- ment of the costal vein less than twice as long as the third ; the transverse veins with broad brownish-black borders ; a spot of the same color lies between the third and fourth longitudinal veins a little before the posterior transverse vein ; a larger spot of the same color lies before it on the costal margin, reaching to the third longitudinal vein and being connected with an equally large, blackish-brown spot on the apex of the wing, which almost attains the fourth longitudinal vein. Hob. Cuba. (Poey.) Gen. V. PSII.OPA FALL. The characters of the genus Psilopa are as follows. Second joint of the antennae with a stout spine ; third oblong, the bristle with long pectinations. Face on its upper part without any keel, slightly convex everywhere, not wrinkled on its sides, receding to- wards the opening of the mouth. Clypeus either quite concealed or scarcely projecting beyond the oral margin. Middle tibia? with- out long bristles on their upper side. The costal vein thickened and attaining the fourth longitudinal vein. This genus is represented in Europe by about twelve species known with more or less certainty. Its representatives in North America seem to be more numerous; a number of them are distin- guished from the European ones by a more robust structure and a more strikingly dusted appearance, and by their faces being not exactly smooth and their cheeks descending a little deeper beneath the eyes ; but neither the number of the species of this group hitherto known is large enough, nor are the characters such as to render a generic separation necessary. On the contrary, it will be sufficient for the present, to put these species together as a sub- division of the genus Psilopa. 142 DIPTERA OF NORTH AMERICA. Synopsis of the Species. Division 1. — Middle of the face slightly convex without any elevation on it. , ( Thorax finely aciculate. 1 aciculata nov. sp. ( Thorax polished. 2 {Posterior part of the thorax and scutellum hronze-colored. 2 scoriacea nov. sp. The whole body pure black. 3 atra nov. sp. Division 2. — Middle of the face slightly convex with some flat longitudi- nal impressions. , ( Abdomen black. 4 umbrosa nov. sp. ( Abdomen steel-colored. 5 caeruleiventris nov. sp. 1. P. aciculata LOEW. J. — Thorace scutelloque nigris, transverse subtiliter aciculatis, capite abdoruineque aut ex cupreo aut ex viridi senescentibus, antennis flavis, pedibus nigris, tibiarum apice tarsisque flavescentibus, basi alarum sublutesceutium nigra. Thorax and scutellum black, transversely with fine scratches ; head and abdomen either coppery or greenish brassy ; antennae yellow ; legs black, tips of the tibiae and tarsi yellowish ; the rather yellowish wings with the base black. Long. corp. 0.09. Long. al. 0.1. Antennae entirely reddish-yellow. Front and face shining, either dull coppery or even almost metallic black, or metallic green. The thorax and the proportionately large scutellum black, hardly brassy, everywhere covered with close and exceedingly fine scratches. Abdomen polished, shining, the color varying in the same way as that of the face. Legs shining black, knees in- distinctly yellowish-brown ; tips of all the tibiae yellowish as well as all the tarsi ; last joint of the tarsi blackish at its tip only. Poisers brownish-black. Wings rather clay-colored, blackish at the base ; this blackening of the costal margin reaches a little be- yond the middle of the first segment, on the disk of the wing only as far as the basal transverse veins ; towards the posterior margin it extends in such a way, as to occupy half of the corner of the wing lying behind the fifth longitudinal vein, but it becomes at the same time very pale. Hab. Cuba. (Poey.) 2. P. scoriacea LOEW. 9- — Atra, nitida, colore in posteriore tho- racis parte scoriaceo, in scutello obscure seneo, proboscide pedibusque nigris, tarsis posticis fuscis, alis cinereo-hyalinis. Black, shining ; the posterior part of the thorax scoriaceous ; scutellum PSILOPA. 143 dull brassy ; proboscis and legs black, hind tarsi brown ; wings grayish- glassy. Long. corp. 0.1. Long. al. 0.13. Shining black. Head proportionately broad, shining black ; above the antennas with a small, dilated spot dusted with whitish. Face broad, shining black, viewed laterally, more brownish-black; viewed from above it appears as if dusted with white. Anten- nae deep black j antennal bristle with long pectinations. Probos- cis and palpi perfectly black. Thorax shining black, scoriaceous posteriorly; immediately before and on the flat scntellum the color is more of a dull brassy green. Abdomen shining black, slightly dusted. Legs black; the posterior tarsi appear rather dark brown to the naked eye, viewed through a lens their color is almost dull whitish, the dark appearance being caused by the black hair. Poisers white. Wings grayish glassy ; the second segment of the costal vein is not half as long again as the third. Hal. Xew York. (Schaum.) 3, P. atra LOE\V. £ . — Atra, nitida, proboscide halterumque capitulo albidis, basi tarsorum posticorum rufa, alis hyaliuis. Shining black, proboscis and knob of the halteres whitish ; base of the posterior tarsi red ; wings glassy. Long. corp. 0.12. Long. al. 0.16. Shining black ; head broad, front shining black ; the small white-dusted spot above the antennas and the whitish, exceedingly fine dust of the broad, shining black face are scarcely perceptible. Proboscis yellowish-white. The palpi seem to be black. Antennas black ; the bristle with very long pectinations. Abdomen shining, scarcely with a trace of dust, rather narrow and flat, apparently consisting of four segments, the first being very much shortened ; even the sixth, however, is perceptible. Legs black ; middle and hind tarsi red at the base. Halteres with blackish petiole and white knob. Wings glassy, slightly grayish ; the second segment of the costal vein not quite half as long again as the third. Hub. Middle States. (Osten-Sacken.) 4. P. limbrosa LOEW. 9- — Nigra, fronte, thoracis dorso et scutello polline brunneo-cinereo tectis, facie griseo-pollinosa, antennis tarsorurn- que basi ex rufo flavis, alis adversus rnargiuem anteriorem nigricantibus, halterum capitulo albo. Black; front, upper side of the thorax and scutellum dusted with ashy- gray with a fuscous tinge, face dusted with whitish-gray ; antennae and 144 DIPTERA OF NORTH AMERICA. bases of all the tarsi reddish-yellow ; wings blackened towards the costa ; knob of the poisers white. Long. corp. 0.13. Long. al. 0.14. Black ; front, thorax and scutellum thickly dusted with ashy- gray with a fuscous tinge. Antennas reddish-yellow ; the bristle with scattered pectinations. Face thickly dusted with whitish- gray, rather of uniform breadth, with more numerous bristles on the sides than in most other Psilopce, generally slightly convex, with the lower part a little projecting, but towards the border of the mouth again remarkably receding, rendering thus the opening of the mouth smaller than is usual in this genus. Proboscis and tarsi blackish. Pleura? brownish-black, shining. Abdomen shin- ing black with very little visible dust. Legs shining black ; tarsi reddish-yellow, the three last joints of the foremost, and only the two last of the hindmost ones being black. Poisers with brown pe- tioles and white knob. Wings clouded with grayish-black, becom- ing gradually black towards the costa; the posterior transverse vein scarcely perceptibly margined with blackish; the second segment of the costal vein more than half as long again as the third. Hob. Cuba. (Poey.) 5. P. caeruleiveutris LOEW. 9- — Capite thoraceque cinereis opacis, abdomine chalybeo nitido, antennis palpisque' flavis, pedibus nigris, tarsis posterioribus rufis, alis hyalinis vitta latissinia atra, rnar- gini antico contigua, pictis. Head and thorax opaque ashy-gray, abdomen shining steel-blue ; antennse and palpi yellow ; legs black, posterior tarsi red ; wings hyaline, with an exceedingly broad, deep black longitudinal stripe on the costa. Long, corp. 0.15. Long. al. 0.16. Front ashy-gray. Antenna? reddish-yellow. Face whitish-gray, slightly convex, with rather slender small bristles on the sides ; oral cavity small; clypeus projecting a little beyond the border of the oral margin. Palpi pale yellowish, a little broader than usual. Upper side of the thorax ashy-gray without any gloss. Pleurae thinly dusted with grayish, and consequently blacker and rather glossy. Scutellum dusted with brown, shining black at the tip. Abdomen shining steel-blue ; the first segment but little shortened, the second as long as the first, the three following longer ; even the narrow sixth segment is distinctly visible. Legs black, the foremost with dark brown knees ; middle and hind tarsi yel- lowish-red with blackish-brown tip ; the fore tarsi have only DISCOCERINA. 145 the base brownish-red. At the costal margin of the wing there is a broad, deep-black band, running from the base to the tip of the third longitudinal vein ; its posterior limit runs from the base of the wing along the middle of the discoidal cell to nearly the pos- terior transverse vein, recedes from it suddenly almost to the third longitudinal vein, follows this vein first at a little, then gradually at a greater distance, and lastly turns to its tip ; the posterior part of the wing is rather dull glassy, and almost grayish at the axillary angle ; the veins in the latter are brownish, those in the blackish parts of the wing black. The place of this species in the system is very deceptive ; for at a superficial view the thickened costal vein seems to reach only the third longitudinal vein ; but this deception arises from its color being black as far as the third longitudinal vein, and very pale between this and the fourth. Hob. Cuba. (Poey.) Geii. VI. DISCOCERI1VA MACQ. The second joint of the antenna has a distinct spine, the third is rounded ; the bristle pectinated. The face on its upper part is distinctly keeled, in the middle more or less inflated, receding again towards the border of the month. Clypeus projecting very little beyond the border of the mouth or entirely concealed ; cheeks moderately descending beneath the eyes. The costal vein attains the fourth longitudinal vein. Upper side of the inter- mediate tibiae without stout bristles. Discocerina stands between the genera Psilopa Fall, and Hecamede Hal., being distinguished from Psihpa by the more rounded form of the third joint of the antenna? and the keel on the upper part of the face ; from Heca- mede chiefly by the cheeks descending much less beneath the eyes. I can describe only two North American species of this genus, yet four are known to me as occurring in Europe. [Three more species, reproduced below, have been published by Mr. Loew since. — O. S.] 1. D. lacteipennis LOEW. ^ . — Ciiierascens, opaca, antennis, geni- bus, tibiarum apice tarsisque flavis, alis albidis, vena costali atra. Opaque, ashy gray ; antennae, knees, tibise at the tips and tarsi yellow ; wings whitish with deep black costal vein. Long. corp. 0.11. Long. al. 0.12. Very similar to the European Hecamede costata Loew, but easily 10 146 DIPTERA OF NORTH AMERICA. distinguished by its cheeks descending much less beneath the eyes. Front with yellowish-gray dust. Antennas reddish-yellow, the third joint a little infuscated on its apical margin ; bristle with a few rays. Face a little more yellowish than the front, distinctly keeled on its upper half, then moderately convex, and receding a little towards the border of the mouth. Eye-rings downwards rather broad; the uppermost of the bristles, inserted near the eye-rings, is more removed towards the middle of the face. Cheeks remarkably de- scending beneath the eyes for a Discocerina. Palpi pale yellowish. Upper side of the thorax and scutellum rather light ashy-gray. Pleurce more whitish-gray. Abdomen light ashy-gray, appearing, on account of the shortness of the first segment, to consist of four segments, the last of which is at least as long again as the penulti- mate. Femora and tibiae black, the former with the extreme tips, the latter with the base and tip yellowish to a greater extent. Tarsi yellowish with blackish tips. Halteres whitish. Wings whitish, especially if viewed in an oblique direction. Costal vein black, the other veins remarkably paler; the second segment of the costal vein is about half as long again as the third. Hob. Washington. (Osten-Sacken). 2. D. parva LOEW. 9 . — Obscure cinerea, opaca, abdomine nigricante ; antennis, genibus, tibiarum apice tarsisque flavis, alls cinereo-liyalinis. Dark ashy gray, opaque, abdomen rather black ; antennae, knees, tibise at their tips and tarsi yellow ; wings grayish-hyaline. Long. corp. 0.07. Long. al. 0.09. Though similar to Discoc. lacteipennis, it is easily distinguished by its much smaller size, nearly black abdomen and grayish-hyaline wings not showing any trace of whitish color. Antennae brownish- yellow, second and third joints brownish on the upper margin ; bristle with four or five long rays. Face dusted with whitish-gray, very distinctly keeled on its upper half, farther beneath rather con- vex, and receding a little again towards the border of the mouth ; in proportion to the size of the insect, it is narrower than in Dis- coc. lacteipennis. Eye-rings exceedingly narrow, not becoming broader downwards. Among the bristles standing in its neighbor- hood, the uppermost is a little more advanced towards the middle of the face. Cheeks descending only a little beneath the eyes. Palpi brownish-yellow. Upper side of the thorax and scutellum blackish ashy gray, opaque ; the pleura? likewise. Abdomen gray- DISCOCERINA. 147 ish-black, opaque, almost pure black and shining towards the end; first segment not strikingly shortened. Femora and tibiae black ; knees, tibiae at their tips and tarsi yellowish. Halteres white. Wings grayish-hyaline, with blackish-brown veins; the second seg- ment of the costal vein is at least half as long again as the third. Hab. Washington. (Osten-Sacken.) 3. I>. orliitalis LOEW. £ . (Translated from Berl. Entom. Zeitschr. 1861, p. 354, by R. Osten-Sacken.) — Cinerea, opaca, colore thoracis magis in ochraceum, abdominis in nigrum vergente, antennis rufis, oculorum orbita Candida, alis hyalinis, segrnento costali secuudo tertii longitudi- nem modice superante. Cinereous, opaque, color of the thorax merging in ochraceous, that of the abdomen in black, antenna rufous, orbit of the eyes shining white, wings hyaline, second costal segment a little longer than the third. Long. corp. 0.065. Long. al. 0.07. Head obscure cinereous, opaque, orbit of the eyes rather broad, not dilated below the eyes, shining white. Face rather broad, the upper half distinctly keeled, the lower half convex, about six small bristles each side, which are more distant from the orbita3 than is generally the case in this genus. Cheeks moderately narrow. Antennre rufescent, third joint rather obscure. On the upper side of the thorax and on the scutel the cinereous color merges in ochraceous ; pleurae somewhat hoary. Abdomen darker than the rest of the body, black towards the apex, very slightly glossy. Femora black, hoary, with a whitish pollen ; tibiae and tarsi yel- lowish, the former with a broad brown ring, the latter with the tip brown. Halteres white. Wings pure hyaline ; second segment of the costa a little longer than the third. Hob. Washington. (Osten-Sacken.) » 4. D. simplex LOEW. (Translated from Berl. Entom. Zeitschr., p. 355, by R. Osten-Sacken.) — Cinerea, opaca, antennis nigris, setis faciei utrinque duabus, genis latioribus, tarsis 'flavescentibus, apicem versus nigris, alis hyalinis. Cinereous, opaque, antennae black, two bristles each side of the face, cheeks rather broad, tarsi flavescent, black towards the apex, wings hyaline. Long. corp. 0.07. Long. al. 0.09. Yery like D. lacleipennis, but easily distinguished by its black 148 DIPTERA OF NORTH AMERICA. antenna?, the smaller number of facial bristles and their different position, finally, by its hardly whitish wings. Cinereous, opaque. Front rather broad, a little darker than the remainder of the body, with an impressed longitudinal line on each side ; frontal lunule very narrow, whitish pollinose. Antenna? black, a whitish polli- nose dot on the upper edge of the second joint, bristle pectinated with four or five long hairs. Face moderately broad, gibbous, its upper half distinctly keeled, the lower one convex, receding at the aperture of the mouth. Two bristles on each side of the face, approximated to the eyes. The very narrow orbit of the eyes becomes broader on their under side. Cheeks broader than in most Discocerince. The cinereous color merges into yellowish on .the thorax ; on the upper side of the abdomen, especially towards the apex, it becomes more distinctly hoary. Feet concolorous to the rest of the body, hoary with a whitish pollen ; anterior tarsi yellowish, blackish towards the tip; hind tarsi entirely obscure. Halteres white. Wings hyaline, slightly tinged with an impure whitish, costal vein not incrassated. Hub. Maryland. (Osten-Sacken.) 5. I>. leiicoprocta LOEW. 9 • (Translated from Berl. Entom. Zeitschr. 1861, p. 355, by R. Osten-Sackeii.) — Cinerea, abdominis atri segmento ultimo niveo, alls hyalinis. Cinereous, abdomen black, its last segment snow white, wings hyaline. Long. corp. 0.064. Long. al. 0.07. Front brownish-cinereous, opaque. Antenna? rufous, third joint fuscous, bristle pectinated with five or six long hairs. Face yel- lowish-white, its upper half keeled, the lower one convex, furnished on each side with three strong bristles. The narrow cheeks, as well as the whole orbit of the eyes, are whitish. Upper side of the thorax blackish-cinereous, opaque, with short black hairs. Pleura? whitish pollinose. Scutelluna^concolorous with the thorax. Ab- domen black, opaque, last segment rather short, shining white. Fore coxa? black, with a white reflection, yellowish at the tip; trochanters yellow ; femora black, cinerascent with a whitish pollen, tip yellow ; fore and hind tibia? black, yellow at basis and apex ; the intermediate ones entirely flavescent ; all with a whitish re- flection on the upper side; tarsi yellow, last joint blackish. Wings hyaline, the third segment of the costa is equal to two-thirds of the length of the second. Hob. Maryland. (Osten-Sacken.) HYPRELLINA. 149 II. HYDRELLINA. The tribe of Hydrellina is characterized by the hairy eyes, the absence of a spine on the second joint of the antennas, and the absence of long bristles on the upper side of the middle tibia?. The eyes in some genera are covered with very short, close hairs ; in other genera these hairs are only scattered, but much longer. Haliday restricts the Hydrellina to the genera Glenanthe, Hydrel- lia, and Atissa. It seems that some other genera, as Philygria, Hyadina, and Axysta can by no means be separated from the Hydrellina, to which they are much more closely related than to the Epliydrina by their whole organization, and chiefly by the structure of the head. The hairs on the eyes of some species of the three last named genera being very sparingly scattered, and therefore difficult to observe, perhaps it will not be superfluous to remark that in the Hydrellina the eyes are always longer and the face is narrowest beneath the eyes, whereas in all Epliydrina the eyes are rounder, the horizontal diameter being sometimes even longer than the vertical, so that the antennas stand where the eyes are most approximated, and the face increases much in breadth immediately below them. Moreover, in the Hydrellina the hole of the mouth is never strikingly widened, and the face downwards never projects much, whereas the great width of the oral cavity and the great projection of the inferior part of the face is a most striking character for the Epliydrina, excepting only the genera Pelina and Ochthera, A confusion between the two last named genera with any genus of the Hydrellina is suSiciently prevented by the entire bareness of their eyes. The genera of Hydrellina may be arranged as follows: — Division 1. Eyes with exceedingly close hair. , ( Antennal bristle with a short pubescence. GLENANTHE Hal. ( Antennal bristle pectinated. 2 2 ( Face convex. HYDRELLIA Desv. I Face impressed. ATISSA Hal. Division 2. Eyes with scattered hair. , ( Face with bristles on both sides. PHILYGKIA Stenh. \ Face almost bare. 2 f Costal vein running to the tip of the fourth longitudinal vein. HYADINA Hal. I Costal vein running to the tip of the third longitudinal vein. AXYSTA Hal. 150 DIPTERA OF NORTH AMERICA. Xorth American species of the genera Hydrellia and Philygria only are known to me. Gen. I. HYDREIXIA R. DESV. The species of Hydrellia are very easily recognized by the very short but exceedingly close pubescence of the eyes, and by the pectinations of the antennal bristle. The other characters are : Second joint of the antennae not unguiculated ; face rather narrow and perpendicular, slightly convex, receding a little towards the border of the mouth ; opening of the mouth not widened ; cheeks descending very little beneath the eyes. Legs rather slender; middle tibiae on their upper side without bristles ; costal vein ex- tending to the tip of the fourth longitudinal vein. Synopsis of the Species.* , f Anterior coxae yellow. 1 ischiaca, n. sp. I Anterior coxse blackish. 2 2 ( Face snowy white. 2 hypoleuca, n. sp. I Face not white. 3 o ( Face brownish-black, opaque. 3 obscuripes, n. sp. ( Face yellow. 4 r Face dark yellow, narrow, much dilated below. 4 scaptilaris, n. sp. 4 < Face pale yellow, rather broad, but little widened below. I 5 valida, n. sp. 1. H. iscliiaca LOEW. 9- — Subsenescenti-fusca. antennis nigris, facie punctoque frontali albis, thoracis margine antico pleurisque canis, pedi- bus ex fusco nigris, coxis anticis, genibus, tibiarum apice tarsorumque basi ex rufo flavis. Somewhat brassy brown, antennae black, face and frontal dot white ; ante- rior border of the thorax and pleura? whitish-gray, legs brownish-black, anterior coxje, knees, tips of the tibiae and base of the tarsi reddish- yellow. Long. corp. 0.1. Long. al. 0.11. Face of medium breadth, slightly dilated below, without keel; the ground color in the middle is more pronounced, giving it rather a brownish aspect ; on each side of the face there are four little bristles, one above the other. Palpi yellow, cheeks a little de- scending. Antennae entirely black ; the bristle in the described specimen has seven rays. Front proportionately broad, dusted with brown; the dot immediately above the antennae white. Upper * The species No. 6 has not been included in this synopsis. — 0. S. HYDRELLINA. 151 side of the thorax dusted with brown ; its anterior border, shoul- ders, lateral border and pleurte grayish-white with white dust. Scutellum like the upper side of the thorax, only a little more glossy. Abdomen almost blackish-brown, not very shining, but also little dusted. Legs brownish-black; fore coxae yellow, some- what infuscated at the base ; all the knees brownish-yellow ; end of the foremost tibias for a little distance and the end of the mid- dle and hindmost as far as the middle, reddish-yellow; fore tarsi only at the base, middle tarsi as far as the middle, the hind tarsi as far as the end of the fourth joint, reddish-yellow. Poisers yellow. Wings a little grayish ; second segment of the costal vein nearly half as long again as the third. Hydrell. iscliiaca is rather similar to the European species: fulviceps Stenh., pilitarsis Steuh , and laticeps Stenh. ; from the first it is sufficiently distinguished by its broader face being dusted with whitish ; from pilitarsis like- wise by its broader and whitish-dusted face, and moreover by the pure white color of the frontal dot, the much whiter dust on the shoulders, anterior and lateral borders of the thorax and pleura?, and finally by the much more extensive pale color of the legs ; from laticeps by the rather less breadth of the front and face, by the wings showing no whitish appearance in any direction, and by the less extensive pale color of the legs. Hub. Middle States. (Osten-Sacken.) 2. H. liypoleuca LOEW. 9 • — Subsenescenti-fusca, antennis nigris, facie punctoque frontali candidis, thoracis margine antico et niargine laterali, pleuris ventreque albo-pollinosis, pedibus nigris, nietatarsis posticis rufis. Somewhat brassy -brown, antennae black, face and frontal dot pure white ; anterior and lateral borders of the thorax, pleurae, and the whole under side of the abdomen dusted with white ; legs black, first joint of the hind tarsi red. Long. corp. 0.11. Long. al. 0.12. Very similar to the European H. incana Hal., which Mr. Haliday thinks to be the same as H. ranunculi, previously de- scribed by him. Face snowy white, not very narrow, underneath broader, slightly keeled in its whole length, beset on each side with three small bristles. Palpi yellow. Cheeks descending but little below the eyes. Antennae black ; antennal bristle in the described specimen with five rays. Front dusted with brown, opaque; anterior border, but especially the shoulder and lateral 152 DIPTERA OF NORTH AMERICA. border dusted with whitish ; the foremost beginning of a grayish- white middle line is indistinctly indicated. Pleurae dusted with whitish. Scutellum dusted with brown and opaque. Upper side of the abdomen only a little dusted, and therefore a little greener and less opaque. The under side and the part of the upper ab- dominal plates which is turned downwards, covered with white dust ; this dust extends to the upper side of the abdomen on the poste- rior part of each segment. Legs black; first joint of the middl'e and hind tarsi yellowish-red; first joint of the fore tarsi brown at the base. Poisers yellow. Wings hyaline, rather grayish; the second segment of the costal vein distinctly half as long again as the third. This species diifers from H. incana by the pure white dust on the anterior and lateral borders of the thorax, as well as on the pleurae. Hab. Middle States. (Osten-Sacken.) I 3. H. ol>scuriceps LOEW. $, . — Subsenesceriti-fusca, abdomine magis virescente, anteunis nigris, facie brunneo-nigra, puncto frontal! albido, pleuris cinereis, pedibus ex fusco nigris, tarsis posterioribus in basi nigris. Brassy brown, abdomen more greenish, antennse black ; face brownish- black with a whitish frontal dot ; pleurse ashy gray ; legs brownish-black, base of the middle and hind tarsi red. Long. corp. 0.1. Long. al. 0.1. Face rather narrow above, a little widening underneath, not keeled, of a brownish-black velvety color; on each side there are three small bristles, one above the other. Palpi yellow. Cheeks descending very little below the eyes. Antennas black, in some directions with a whitish reflection ; antennal bristle in the described specimen with six rays. The dot immediately above the antennae dusted with whitish, but not strikingly so. Front and upper side of the thorax dusted with brown, opaque; anterior and lateral borders of the thorax as well the shoulder without pale dust. Pleura pale ashy gray, more brown above. Scutellum of the same color as the upper side of the thorax. Abdomen brownish metal- lic-green, somewhat glossy; first segment much shortened, second and third of equal length, fourth a little longer, fifth as long as the second and third together, rather broadly truncate at its end, somewhat convex. Legs brownish-black ; first joint of the pos- terior tarsi yellowish-red; the first joint of the foremost tarsi brownish-red only at the base. Poisers yellow. Wings hyaline, a HYDRELLINA. 153 little grayish; the second segment of the costal vein scarcely half as long again as the third. Not possessing any of the few Eu- ropean HydrellifB with dark colored faces, I cannot point out bow H. obscuriceps differs from them. Hal. Middle States. (Osten-Sacken.) 4. II. scapularis LOEW. 9 • — Subsenescenti-fusca, antennis nigris, facie ochracea, puncto frontali pallidius flavo, interdum albido, thoracis margine antico, hunieris pleurisque albo-pollinosis, pedibus nigris, me- tatarsis posticis rufis. Brassy-brown ; antennae black, face ochraceous, frontal dot paler yellow, sometimes whitish, anterior border of the thorax, shoulders, and pleurae dusted with white; legs black, first joint of the hind tarsi red. Long. corp. 0.1. Long. al. 0.12. Very similar to H. hypoleuca, notwithstanding the different color of its face, but certainly not a variation in color of that species. Face a little narrower above than in the latter, quite as broad underneath, thus appearing more dilated below, likewise keeled on its whole length, but more distinctly and a little less obtusely; on each side of it there are three small bristles; its color is dark ochraceous. Antennas black; bristle with five or six rays. Front dusted with brown, opaque, narrower than in H. hypoleuca; the dot immediately above the antenna? is dusted with paler yellow or whitish. Thorax dusted with brown, opaque, the dust not being so thick as to prevent its color from inclining a little to greenish ; its outermost anterior border and the shoulders are dusted with whitish ; the dust of the pleurae is of the same color. Upper side of the abdomen greener than that of the thorax, slightly dusted, but also slightly glossy; its under side and the part of the upper abdominal plates which is turned downward, are but thinly dusted with whitish. Legs black; first joint of the posterior tarsi yellow- ish-red ; first joint of the anterior tarsi brown at the base. Poisers yellow. Wings glassy, rather grayish; the second segment of the costal vein more than half as long again as the third. Hob. United States. (Schaum.) 5. H. valida LOEW. 9. — Inter majores sui generis; glauco-cinerea, tota opaca, facie latiuscula pallide ochracea, antennis pedibusque nigris, basi tarsorum omnium rufa. Belonging to the largest species of this genus ; greenish-gray, opaque every- 154 DIPTERA OP NORTH AMERICA. where ; face rather broad, pale ochraceous ; antennae and legs black ; base of all the tarsi red. Long. corp. 0.12. Long. al. 0.15. Face rather broad, becoming a little broader upwards, only slightly keeled above, ochraceous, with three small bristles on each side. Palpi yellow. Cheeks slightly descending. Antennae black; antennal bristle in the described specimen with five rays. Front greenish-gray and opaque in consequence of its grayish dust; the dot above the antenna? has a still duller yellow color than the face and is not conspicuous. Upper side of the thorax and scutellutn greenish-gray and opaque from its whitish-gray dust. Pleura? a little paler greenish-gray. Abdomen grayish-green, opaque, with the fifth segment considerably longer than the fourth. Legs black; tarsi yellowish-red as far as the end of the first joint; also the knees, chiefly those of the hind legs, are of this color. Poisers yellow. Wings relatively to the length of the body, large, hyaline ; the veins in the neighborhood of the base pale ochraceous ; the second segment of the costal vein about twice as long as the third; the posterior transverse vein does not stand quite perpen- dicularly to the longitudinal axis of the wing, but is slightly oblique. Hah. Middle States. (Osten-Sacken.) 6. H. formosa LOEW. 9 • (Translated from Berl. Entom. Zeitschr. 1861, p. 355, by R. Osten-Sacken.) — Atra, thoracis dorso, abdorninis apice marginibusque nitidis, fronte, thoracis macula lateral! permagna scutel- loque aterrirnis, opacis, facie, puncto frontali, pleurarurnque vitta supe- riore, candidis, femoribus nigris, tibiis tarsisque pallidis, alis cinereo- hyaliuis. Dark, thorax above, tip of the abdomen and its borders shining ; front, a large spot on the side of the thorax and scutellum deep black, opaque ; the face, a dot on the front and a band on the upper side of the pleurae shining white, femora black, tibiae and tarsi pale, wings cinereous-hya- line. Long. corp. 0.057. Long. al. 0.064. Face moderately convex, not keeled, bright shining white. Cheeks very narrow, black. Front and superior part of occiput deep black, velvety, with a striking shining white frontal mark. Antennas black, third joint rufous, margined with black above, bristle pectinated with scattered black hairs. Thorax very shining above, on each side a large, deep black, velvety lateral spot. HYDRELLINA. 155 Scutellum deep black, opaque, with a narrow subcinereous border. Pleurae black, with a broad, shining white band above. Abdomen black, opaque, the apical half as well as the lateral borders shining. Femora black ; tibiae and tarsi pale yellowish, the upper edge of the former with a white reflection, terminal joint of the latter black. Halteres pale lemon-yellow. Wings cinereous-hyaline, second segment of the costa somewhat longer than the third ; third longitudinal vein ending at the very tip of the wing ; posterior transverse vein occupying the middle between the basis and the tip of the wing. Hab. Pennsylvania. (Osten-Sacken.) Observation. — This species, although very much like Philhygria picta Fall, and the allied species, proves to be a true Hydrellia on account of the short and very dense pubescence of its eyes. Gen. II. PHILYGRIA STENH. Haliday has employed for this genus the name Hydrina, given by Rob. Desvoidy; but as this name, being derived from Hydra, is also used in the family of Polypi, it seems more advisable to adopt for it the name Philygria of Stenhammar ; otherwise this name would not be used at all, the two other genera, which joined with -the present, form the genus Philygria of Stenhammar, being already possessed of their authorized names, Hyadina and Axysta. The genus Philygria, taken in the present sense, may be character- ized in the following manner: Second joint of the antennas not unguiculated; antennal bristle with a short pubescence. Eyes dis- tinctly hairy, rather rounded, but higher than broad, slightly pro- minent. Face descending obliquely, narrowed upwards, receding a little towards the mouth, the anterior border of which is a little pointed; on both sides there are distinct bristles. Clypeus unde- veloped ; mentum rather thickened ; cheeks slightly descending. The costal vein attains the fourth longitudinal vein ; the posterior transverse vein is rather distant from the border of the wing. 1. I*, fiiscicarnis LOEW. Fusco-cinerea, abdoniine nigricante, facie flavescente, pedibus nigris, tarsis rufis, in apice fuscis. alis cinerascen- tibus, cellula discoidali et gutta pone venam transversam posteriorem limpidioribus, venis transversis fusco-limbatis. Brownish-gray ; abdomen black'isli, face yellowish ; wings gray with the discoidal cell and a drop behind the posterior transverse vein more hya- 156 DIPTERA OF NORTH AMERICA. line ; transverse veins margined with blackish-brown. Long. corp. 0.9. Long. al. 0.11. It lias a certain resemblance with Philygr. femorata Stenh. and interrupta Hal., namely, the form of its face is almost as in the latter, and likewise more yellowish on the middle, whitish on the lateral borders and towards the cheeks. Antennas blackish, ap- pearing whitish-gray in certain directions, brownish on the inferior border only, when held against the light. Thorax grayish-brown, on its anterior border dusted with whitish-gray; its darker stripes are obsolete, but more visible in the neighborhood of the anterior border, where they extend a little into the brighter gray color. Scutellum as the upper side of the thorax. Pleurae gray. Ab- domen grayish-black, more black towards the end, not glossy. Legs black, tarsi yellowish-red as far as the fourth joint. Wings grayish with a hyaline spot behind the posterior transverse vein and with a rather clearer discoidal cell ; the clearer color of the latter is only seen if the light shines through the wing and the wing is looked at in an oblique direction, while the clear spot be- hind the posterior transverse vein is distinctly seen in every direc- tion. The two transverse veins have only a very narrow and ill-defined dark margin, and the posterior transverse vein is a little less distant from the posterior border of the wing than is usual in this genus ; the second longitudinal vein being very long, the second segment of the costal vein is more than twice as long as the third. Hob. Middle States. (Osten-Sacken.) 2. P. opposita LOEW. % . (Translated from Berl. Entom. Zeitschr. 1861, p. 356, by R. Osten-Sacken.) — Ex cinereo fusca, abdomine atro nitido, alarum venis longitudiualibus secunda, terti& et quarta nigro- punctatis, venis transversis late nigro-limbatis. Cinereous-brown, abdomen black, shining, the second, third and fourth longitudinal veins of the wings spotted with black, transverse veins broadly clouded with black. Long. corp. 0.07 — 0.09. Long. al. 0.095. Cinereous-brown, opaque. Ocellar triangle large, concolorous, rather indistinctly separated from the remainder of the front. Two basal joints of the antennae black, the third black, with the bases and the apical half impurely rufous. Face narrow, black, with a HYDRELLINA. 157 whitish pollen, its middle portion flavescent below. Facial orbital of the eyes narrow, with a white reflection. Thorax obscure, cinereous brown above, with very narrow, obsolete darker lines. Pleuras dark cinereous. Scutellum concolorous with the thorax. Abdomen black, very glossy, a large obscurely cinereous opaque basal spot, not attaining the posterior margin of the second seg- ment. Legs yellowish ferruginous, last joint of tarsi black, base of femora sometimes fuscous. Wings cinereous hyaline, veins black ; short stumps of veins clouded with black, proceed from the second, third, and fourth longitudinal veins ; the third vein emits four such stumps, all of which, except the last, are opposed to similar stumps on the second vein ; the last segment of the fourth vein generally emits two stumps; the ordinary transverse veins are broadly clouded with black. The second segment of the costa is almost twice as long as the third. Hab. Pennsylvania. Washington. (Osten-Sacken.) Observation. — Phil, opposita is very like P. punctato-nervosa Fall., but distinguished by a more brown color, a more narrowed face, darker legs and antenna, a larger portion of the abdomen colored with black and by a smaller number of dots on the wings. The facial orbits of the eyes, which are much narrowed in P. opposita, evidently prove it to be a distinct species. Note. — Some specimens have^/tfe stumps on the third vein, opposed to four on the second, and more than two stumps on the last segment of the fourth vein. 0. S. 3. P. deMlis LOEW. % . (Translated from Berl. Entom. Zeitschr. 1861, p. 356, by R. Osten-Sacken.) — Nigro-cinerea, opaca, antennis totis nigris, fronte atra opaca, triangulo ocellari maxinio, nigro-cinereo, ultimo abdo- minis segrnento atro, nitido, pedibus obscuris, genibus, tibiarum anteri- orum apice, tarsisque flavescentibus, horum apice nigro, alis cinereo- hyalinis, circa venas transversales infuscatas limpidioribus. Blackish-cinereous, opaque, antennae entirely black, fro-nt black, opaque, ocellar triangle very large, blackish-cinereous, last segment of the ab- domen black, shining, feet obscure, knees, tip of the anterior tibiae and tarsi yellowish, tip of the latter black, wings cinereous-hyaline, with clearer spaces round the infuscated transverse veins. Long. corp. 0.05. Long. al. 0.064. Blackish-cinereous, opaque. Front black, with a very narrow white marginal line on each side and the rather large ocellar 158 DIPTERA OF NORTH AMERICA. triangle, blackish-cinereous. Antennas entirely black. Face yel- lowish, orbits narrow, whitish. Thorax above with very narrow almost obsolete lines. Abdomen a little darker and less opaque than the thorax, last segment black, smooth. Legs blackish, knees and tip of the anterior tibiae yellowish, hind tibiee either alto- gether blackish, or marked with a narrow, very obsolete pale ring, tarsi yellowish, their last joints blackish. Halteres impure white, knob somewhat darker. Wings cinereo-hyaline, with clearer spaces round the infuscated transverse veins, second costal seg- ment almost twice as long as the third. Jfab. Pennsylvania. (Osten-Sacken.) Observation. — This species is very like Philygr. femorata Stenh., but distinguished by entirely black antennse, by a less obtuse an- terior angle of the ocellar triangle and by a conspicuously longer second costal segment. III. EPHYDRIXA. The JEphydrina are well characterized by their quite naked, prom- inent, and usually much rounded eyes, by the second joint of their antennae not unguiculated, and by the middle tibiae without spinous bristles on their upper side. By the genus Pelina they are nearest related to the latter genera of Hydrellina. The mentum is much enlarged and swollen in almost all the genera, the oral cavity generally of large width. The genera with less widely opened mouth, as Pelina and Ochthera, so manifestly bear the chief cha- racters of Ephydrina, that no doubt can arise about their systema- tic position. The genera of Ephydrina hitherto established may be arranged as follows : — Division 1. Clypeus prominent. , ( The small basal cells of the wings complete. CANACE Hal. \ The small basal cells of the wings wanting. 2 n ( Oral cavity proportionally narrow. ( Oral cavity exceedingly wide. 4 „ ( Fore femora not thickened. PELINA Hal. \ Fore femora much thickened. OCHTHERA Lair. f Costal vein attaining the third longitudinal vein. 4 -| BRACHYDEUTERA Loew. (. Costal vein attaining the fourth longitudinal vein. 5 EPHYDRINA. 159 fFace on each side with a long bristle; lateral border of the mouth without bristles. PARYDRA Stenh. [ Face on each side with several long bristles ; lateral border of the mouth with bristles. HALMOPOTA Hal, Division 2. Clypeus retracted in the oral cavity. 1 Claws almost straight, pulvilli indistinct. EPHYDRA Fall. Claws curved, pulvilli distinct. „ ( Oral border quite bare. ILYTUEA Hal. \ Oral border with bristles. 3 o ( Antennal bristle bare. TICHOMYZA Macq. ( Antennal bristle not bare. 4 A ( Antennal bristle pubescent. SCATELLA R. Desv. (. Antennal bristle pectinated. C^ENIA R. Desv. The North American Ephydrina known to me belong to the five genera : Ochthera, Brachydeutera, Parydra, Ephydra, and Scatella. Gen. I. OCHTHERA LATK. One of the most distinct genera of Ephydrina. Front very broad ; antennal bristle above, with three rays. Face above moderately broad, with two furrow-like longitudinal impressions approaching each other very much on the middle, then diverging from each other as they descend, and finally continued in a direc- tion parallel to the lateral border of the mouth ; on the surface of the face there are some fine and short hairs, but no bristles at all. The face and cheeks descend very deep beneath the large promi- nent eyes, but are again contracted sensibly towards the opening of the mouth, rendering it smaller than in any of the other genera of Ephydrina. Clypeus having the form of a small flat lamella, projecting beyond the anterior border of the mouth. The fore coxaB a little prolonged ; the fore femora exceedingly swollen, furrowed on their under side for the reception of the curved tibise, which terminate in a spine, and beset with a few small bristles ; the first joint of the hind tarsi more or less thickened. The costal vein of the wings reaches to the fourth longitudinal vein ; the second segment of the costa is proportionally very long; the pos- terior transverse vein is very oblique ; the third and fourth longi- tudinal veins converge rather remarkably towards their ends. Observation. — Th. Say has described a fly as Ochthera empifor- mis ; but on a closer consideration of his "observations on the an- terior femora, the color of the insect, and its small size, it becomes 160 DIPTERA OF NORTH AMERICA. indubitable that he has been deceived relatively to the true charac- ters of the genus Ochthera, and it is to be supposed that his OcJdhcra empiformis is an insect belonging to the Tachydromidse. Synopsis of the Species. ( Face with deep black furrows and dots. 1 exsculpta, n. sp. ( Face without black furrows and dots. 2 9 ( First joint of the hind tarsi but little swollen. 2 mantis Deg. i First joint of the hind tarsi much swollen. 3 „ ( Tarsi black, face broad. 3 rapax, n. sp. ( Tarsi red, face narrow. 4 tuberculata, u. sp. 1. O. exscillpta LOEW. £. — Fades angustissiuia, lineis punctisque exsculptis ornata ; tibiae anticae rufae. Face exceedingly narrow, with shining-black furrows and impressed black dots ; fore tibiae red. Long. corp. 0.16. Long. al. 0.16. A readily distinguished species, not quite equalling the three following in size. Front narrower than in all the other known species, almost entirely covered with a large shining spot having the form of a regular trapezium, near which the color is velvety- black at the borders of the eyes and brownish on the anterior cor- ners of the front. Eyes larger and longer than in the other spe- cies. Face unusually narrow, dusted with yellow ; a shining black furrow runs from the tubercle placed in the middle of the face to the border of the mouth, and has on each side a similar furrow, the under part of which is laterally continued in a parallel direction to the border of the mouth ; the lateral parts of the face have some impressed, rather coarse, dots. Clypeus sensibly smaller than in the other species. The fore femora black ; the fore tibia? and tarsi red, the first joint of the latter a little longer and a little less pear-shaped than in the other species. The middle and the hind legs black ; the tips of the knees and the tibia? on their first third red ; the first joint of the middle tarsi red as far as the tip, the following joints being so only at the base ; the first joint of the hind tarsi is very little swollen, the second and following joints red at the base. Hob. Cuba. (Poey.) OCHTHERA. 161 2. O. mantis DEG. £ and 9 • — Pedes nigri, tibiis intermediis non dilatatis. tarsorum intermediorum basi rufS,, metatarso postico modice incrassato. Legs black ; middle tibiae not enlarged, middle tarsi red at the base, first joint of the hind tarsi only little swollen. Long. corp. 0.24 — 0.25. Long. al. 0.2. I am quite unable to distinguish this species, so common in the Middle States of the Union, from the European Ochlh. mantis. It is true, indeed, that in most American specimens the eyes are a little more distant from each other than in the European; but this difference in some cases disappears entirely; nor do the European specimens altogether agree in this respect. The color of the face is likewise as variable as in the European specimens. As mark- ings, distinguishing this species from the two next ones, which resemble it very much, the following may be noted : The ground color of the legs, in well-colored specimens, is black, only the mid- dle tarsi being red from their base for a very variable extent. The middle tibia3 are considerably narrower than in Ochth. rapax and tuberculata, and entirely dusted on their anterior side; the first joint of the hind tarsi is very little swollen and rather long. Halt. Middle States. (Osten-Sacken.) 3. O. tuberculata LOEW. $. — Pedes nigri, tibiis intermediis subdila- tatis, tarsis omnibus obscure rufis, metatarso postico nigro, valde iucras- sato. Legs black; middle tibiae a little enlarged ; all the tarsi dark red, the first joint of the hind tarsi black and very much swollen. Long. corp. 0.18. Long. al. 0.17. Yery similar to Ochth. mantis, but its face is considerably nar- rower in its upper part, and the elevation in the middle of it forms more distinctly a small double knob. The middle tibia? are broader, on their outer edge sharper, and polished on a great part of its anterior side ; the knees of the hind legs and all the tarsi brown- ish-red, the last joint of the latter more brownish ; the first joint of the hindmost tarsi black, mueh swollen. Hob. Illinois. (Schaum.) 11 162 DIPTERA OF NORTH AMERICA. 4. O. rapax LOEW. ^ . — Pedes riigri, tibiis intennecliis subdilatatis, tarsorum iutermediorum basi rufa, metatarso postico valde incrassato. Legs black, middle tibise a little enlarged, middle tarsi red at the base, first joint of the hind tarsi much swollen. Long. corp. 0.16. Long. al. 0.17. Yery similar to Ochth. tuberculata in the form of the legs, only the middle tibise are of a less equal breadth, but become sensibly broader towards their end. It is also very easily distinguished from Ochth. tuberculata by its broader and shorter face showing only an exceedingly flat elevation in the middle. Legs entirely black, only at the base of the first joint of the middle tarsi there is a slight red tinge; the first joint of the hindmost tarsi is still a little shorter and thicker than in Ochth. mantis. The wings have rather a more distinct blackish-gray clouding than in the other species. It differs from Ochth. mantis by its shorter face, the flat- ter elevation in the middle of it, and the much shorter and thicker basal joint of its posterior tarsi. Hob. Carolina. (Zimmermaun.) Gen. II. BRACHYDEUTERA LOEW. Eyes naked, proportionately rather large. Front exceedingly broad. Second joint of the antennae not unguiculated, as large as the third, the latter rounded ; antennal bristle with unusually long rays. Upper part of the face deeply impressed on both sides, and with a keel, resembling a nose, in the middle; the lower part of it is very prominent. The anterior end of the oral margin very much ascending and allowing the convex clypeus to appear. Be- sides, the whole face is quite bare, with the cheeks descending but very little beneath the eyes. Legs quite bare, rather slender and long; anterior tarsi elongated and exceedingly slender; claws small and delicate, pulvilli rather indistinct. Costal vein of the wing reaching only to the tip of the third longitudinal vein; second longitudinal vein exceedingly short and curved towards the costa like an arch, so that the third segment of the costa is several times longer than the second; the small transverse vein is unusually dis- tant from the base of the wing; the posterior transverse vein is at a little distance from the border of the wing and has a nearly per- pendicular position ; the last segment of the fourth longitudinal vein is much attenuated. BRACHYDEUTERA. 163 1. B. diniicliata LOEW. 9- — Superius brunnea, inferius tota Can- dida. On the upper side brown, on the whole under side white. Long. corp. 0.13—0.14. Long. al. 0.14—17. ' Dark brown and entirely opaque on the whole upper side. In well preserved specimens there are, on the upper side of the thorax, two somewhat grayish-brown, approximated, longitudinal lines, which commence at the anterior end of the thorax and stop before reaching the posterior end; between them there is the trace of a fine pale middle line, which becomes more distinct at the posterior end of the thorax and is continued through the scutellum; there are besides two other longitudinal lines, which, being nearer the lateral border and interrupted in the neighborhood of the suture, are not truncated posteriorly and continue indistinctly on the lateral borders of the scutellum. Some specimens show very faiut 'traces of these markings of the thorax. The keel, resembling a nose, on the upper part of the face is dark brown ; the remainder of the face together with the cheeks, and the inferior half of the occiput, breast, and pleurae, as well as the part of the upper abdominal plates which are much turned downwards, are almost silvery white; this color on the last abdominal segments ascends a little to the upper side of the posterior borders. Legs in well-colored indi- viduals blackish-brown, only the apical third of the femora and the first half of the posterior tibia? being more or less reddish-brown; in less distinctly colored specimens often only the tips of the tibia? and the tarsi are blackish-brown, all the remainder being brownish- yellow. Wings hyaline with brownish-black veins, sometimes more clouded with grayish in the neighborhood of the costa ; the third segment of the costa is twice and a half or three times longer than the second. Hob. Washington. (Osten-Sacken.) Observation. — A female sent by Poey from Cuba differs from those received from Baron Osten-Sacken by its brown wing-veins and clay-yellow legs, the tarsi only being of a dark-brown color ; but it is only a paler colored specimen of Brachyd. dimidiata, which became still paler in the course of time. 164 DIPTERA OF NORTH AMERICA. Gen. III. PARYDRA STENH. Form of the body short and stout. Thorax and scutellum very convex. Front very broad. Antennal bristle on the upper side with a short pubescence, which in some species is difficult to perceive ; bare towards the end. Face very broad, not so much vaulted as in the true Ephydrce, but with a convexity descending more obliquely, on each side with a very characteristic, long, curved, hair-like bristle, beneath which are some shorter hairs, hardly per- ceptible in some species. Clypeus prominent. Cheeks descend- ing beneath the eyes. Lateral borders of the mouth quite bare. Mentum exceedingly thickened. Legs short and rather clumsy. The small transverse vein of the wings is behind the middle of the discoidal cell, consequently proportionately far from the base of the wing ; the posterior transverse vein is not very near the border of the wing, and has a more or less oblique position ; the alula is strikingly large. The species of this genus may be divided into two sections, the first of which comprises thickly hairy species with very convex faces. In North America only naked species, belonging to the second section, have been as yet discovered ; they are very similar to the European species of this section ; however, they appear to have more plastic differences than these, and to be consequently more easily distinguished from each other. This can be said at least of the four species known to me, none of which is provided with the small appendage of the second longitudinal vein distinguishing some of the European species. Synopsis of the Species.* , ( Scutellum with conical warts. 2 I Scutellum without conical warts. 3 2 f Scutellum with two warts. 1 bituberculata, n. sp. 1 Scutellum with four warts. 2 quadrituberculata, n. sp. o / Face nearly perpendicular. 3 breviceps, u. sp. I Face descending obliquely. 4 paullula, n. sp. * The species No. 5 has not been included in this synopsis. — 0. S. PARYDRA. 165 1. P. Mtllbercillata LOEW. £ and 9. — Ex brunneo senescens, alis cinereo-hyalinis, venis transversis nigro-limbatis, scutello bituber- culato. Brassy-brown, wings grayish with black margins of the transverse veins ; scutellum with two warts. Long. corp. 0.17 — 0.18. Long. al. 0.17. Yery similar to Parydra aquila Fall, in size, form, and color. Face proportionally not very prominent, dusted with brown ; the characteristic bristle on each side proportionally slender ; upwards near it, but a little more towards the middle of the face, is a rather distinct, impressed spot; the shorter hairs inserted beneath it are hardly perceptible. Orbitae and cheeks very broad ; clypeus very prominent. Antennae black ; antennal bristle hair-like and bare towards the end, stouter about the middle, and with a short pubes- cence on the upper side. Upper side of the thorax with rather indistinct stripes ; the rows of fine punctures, including the stripes, a little more distinct than in the other species. Scutellum at the tip with two not approximated warts, bearing at the end the two usual small bristles of the scutellum. Legs dark, with only the tarsi usually red with black tips ; there are individuals with much darker tarsi ; the white reflection at the base and tip of the tibiae not very striking. Wings clouded with grayish, having brownish- black veins and black margins of the transverse veins, in the neighborhood of which the surface of the wings is more distinctly hyaline ; the second segment of the costa is nearly twice as long as the third ; the ends of the third and fourth longitudinal veins parallel. There are some specimens, the faces of which are dusted with dull whitish ; but these certainly belong to the same species. Hob. Middle States. (Osten-Sacken.) 35. P. quadritll"bercillata LOEW. £ and £>. — Nigro-senea, alis hyalinis, veuis transversis interdum nigro-limbatis, scutello quadritu- beruulato. Brassy-black, wings hyaline, transverse veins sometimes margined with blackish ; scutellum with four warts. Long. corp. 0.17. Long. al. 0.17. Similar to the preceding species in color, but a little blacker, not quite equalling it in size. Face generally dusted with white, the dust less frequently quite yellowish on the upper part; the under part of the face projects somewhat less than in Parydra Ut,u- berculata; the characteristic bristle on each side is very slender and 106 DIPTERA OF NORTH AMERICA. rather short ; no impressed spot in its neighborhood ; the short small hairs beneath it are scarcely visible ; eye-rings and cheeks very broad, but the latter a little narrower than in Parydr. bitu- berculata. Antenna? black ; the bristle towards its end excessively slender, being stouter to about its middle, and provided on its upper side with a hardly distinguishable pubescence. Thorax rather indistinctly striped ; the two longitudinal lines formed by fine scarcely visible punctures. Scutellum on its tip with two very approximated conical warts, on the tips of which are the two small bristles usually inserted at the end of the scutellum ; on each side there is a similar tubercle, ending likewise in a small bristle. Tibise and tarsi usually brownish-red, with blackened tips ; but there are specimens with the tibiae quite black and the tarsi brown only at the base, the remainder being quite black ; only in recently devel- oped specimens the anterior side of the tibia? is dusted with white on their whole length ; this white dust is generally interrupted behind the middle of the tibiae. Wings proportionally a little longer than in the other species ; the second segment of the costa is about one-half longer than the third; the last segment of the fourth longitudinal vein is unusually long, showing the trace of a slight convergency towards the third longitudinal vein ; the fifth longitudinal vein is truncated immediately behind the posterior transverse vein ; otherwise the wings are hyaline with a very faint grayish tinge ; the veins are brownish-black as far as the base, or frequently brown or brownish-yellow in the neighborhood of the base ; sometimes this brownish-yellow color on the costal vein ex- tends to far beyond the middle of the wing ; the transverse veins in most specimens are not margined, or show only a trace of black- ish-gray clouding ; but sometimes they have rather broad blackish margins, the surface of the wing being clearer in their neighborhood ; these margins are found particularly in specimens which have a blacker coloration and almost entirely black legs. The deviations are more remarkable than those occurring in the other species of Parydra; but there are various transitions between them, which make it improbable that there is more than one species. Hab. Middle States. (Osten-Sacken.) PARYDRA. 161 3. P. toreviceps LOEW. 9 . — Nigro-senea, facie subperpendiculari, scu- tello inutico, venis alarum transversis obscure linibatis. Blackish-aeneous, face rather perpendicular, scutellum without warts ; transverse veins of the wing margined with obscure. Long. corp. 0.16. Long. al. 0.16. Blackish-aeneous. Face dusted with brown, less projecting than in any other Parydra known to me, consequently almost quite perpendicular. Orbita? excessively narrow ; the characteris- tic small bristle on each side of the face is of moderate length and rather slender ; beneath it there are a few shorter distinctly visible hairs. Clypeus very narrow ; cheeks broad. Antennae black ; the bristle rather slender even at its basal half, hair-like towards its end, with a short but distinct pubescence on its upper side reaching beyond the middle. Thorax rather indistinctly striped ; scutellnm without tubercles, as is the case in the European species. Femora black. Tibiae reddish-brown, with a little white reflection at the base and tip, but in the specimen now before me it is too rubbed off to afford any certainty about its extent and nature. Abdomen rather shining, almost with a band of whitish-gray hoar on the posterior border of each segment. Wings rather tinged with grayish, having blackish margins on the transverse veins, the surface of the wings being more hyaline in their neighborhood. The second longitudinal vein is considerably shorter than in the two preceding species, in consequence of which the second segment of the costa is but little longer than the third ; the ends of the third and fourth longitudinal veins are parallel ; the fifth longitu- dinal vein curves a little posteriorly at the second half of the dis- coidal cell. Hob. Middle States. (Osten-Sacken.) 4. p. paullula LOEW. 9. — Omnium minutissima, facie proclivi, genis angustis. Very small ; face projecting obliquely, cheeks very narrow. Long. corp. 0.06—0.07. Long. al. 0.06—0.07. A very small species, of which I have only a single somewhat immature specimen, so that I am unable to say more of its colors than that they appear to differ little from those of the other species. Face descending obliquely, and therefore rather projecting with its lower parts ; the characteristic bristle on each side rather long ] 68 DIPTERA OF NORTH AMERICA. and unusually near the border of the mouth. Clypeus and cheeks excessively narrow. Antennae black, the bristle with fine pubes- cence to beyond the middle. Scutellum without marginal tuber- cles ; the two small bristles on its tip rather distant from each other. Wings in better colored individuals undoubtedly with much gray clouding and blackish margins of the transverse veins, in the neighborhood of which the surface of the wings is more glossy; the second segment of the costa is only a fifth longer than the third ; the ends of the third and fourth longitudinal veins with a trace of a slight divergency ; the fifth longitudinal vein extends to the border of the wing. This species is very similar to Parydra pusitta Meig. 5. P. a"b"breviata LOEW. % . (Translated from Berl. Entom. Zeitschr. 1861, p. 357, by R. Osten-Sacken.) — Minuta, antenuis tibiisque ferru- gineis ; alse infuscatse guttis aliquot liyalinis distinctissimis ornatae, segmento costse tertio secundi longitudinem paulo excedente, venis lon- gitudinalibus tertia et quarta distincte divergentibus. Small, antennse and tibiae ferruginous ; wings infuscated, with several hyaline, very distinct dots, third segment of the costa somewhat exceed- ing the second in length, third and fourth longitudinal veins distinctly diverging. Long. corp. 0.07. Long. al. 0.07. Olivaceous. Antennae obscure ferruginous, the two first seg- ments and the upper edge of the third, black ; the whole bristle has a short pubescence above. Face moderately sloping, the or- dinary bristle on each side is not more approximated to the peri- stoma than in most of the congeners. Cheeks narrow. Scutellum not tuberculated. Legs black, knees, tibiee and base of tarsi fer- ruginous ; the whitish pollen, generally extant on the tibiae of the allied species, is wanting here. Wings rather short, distinctly infuscated, marked with seven rather large hyaline spots ; second longitudinal vein with a very short, hardly perceptible appendage ; third and fourth veins diverging near the apex ; second costal segment almost equal in length to the third. Hub. Pennsylvania. (Osten-Sacken.) EPHYDRA. 169 Gen. IV. EPHYDRA FALL. The hairy, exceedingly vaulted, and very projecting face, the very large opening of the mouth with ciliated border, the con- cealed clypeus, the nearly straight and rather long claws, and the indistinct pulvilli, characterize the genus Ephydra. The bris- tle of the antennae is usually pubescent, sometimes almost pecti- nated with short rays. The genera nearest related to Ephytfra are Ccenia and Scatella, the claws of which are curved and the pulvilli distinct. The genus Tichomyza is not so near to the genus Epltydra and may be easily distinguished from it by its unusually large pulvilli. 1. E. atro-virens LOEW. £ and £ . — Obscure viridis, nitida brun- neo-pollinosa, antennamm articulo tertio unipili, seta brevissime pube- rula; £ quinto abdominis segmento prsecedente breviore, hypopygio brevi, marginem segmenti quart! posteriorem non attingente. Dark green, glossy, dusted with brown; third joint of the antennae with a hair, the terminal bristle with very short pubescence ; £ , fifth segment of the abdomen shorter than the fourth, hypopygium short, not reach- ing the posterior border of the fourth ventral segment. Long. corp. 0.17 —0.18. Long. al. 0.17—0.18. Exceedingly similar to the European Ephydra micans Hal., so that I am unable to distinguish the female of the two species, but the much shorter hypopygium of the male characterizes the species as a distinct one. Dark metallic green, very shining, but with brown hoar on the front, thorax, and abdomen, which, distinctly appearing on an oblique inspection of these parts of the body, makes them appear brown and opaque; this brown color is least visible on the abdomen. AntennaB black ; third joint on its outer side near the base with a single bristle-shaped hair, which is longer than the joint itself; antennal bristle only with very short pubes- cence. The front and the sloping space extended between the antennae and the highest elevation of the face are shining green or bluish-green. Face dusted with white, which, according to the observations made in the allied species, may not be a constant marking; border of the mouth in both sexes with short and rather fine cilia, quite as in Ephydr. micans Hal. The ground color of the legs is greenish-black, covered with dust, shining blackish- green on the rubbed parts. Wings clouded with blackish-gray. Hob. Middle States. (Osten-Sacken.) 170 DIPTERA OP NORTH AMERICA. Gen. V. SCATELLA ROB. DESV. This genus contains only smaller and generally not, metallic species. Front and face very broad ; eyes rounded ; face usually very convex, hairy and bristled ; border of the mouth ciliated ; opening of the mouth wide; clypeus concealed; cheeks moderately broad; mentum swollen. Second joint of the antennas not ungui- culated ; antennal bristle with fine, usually very short pubescence. Claws curved, pulvilli distinct. Costal vein of the wings reaching to the tip of the fourth longitudinal vein ; the small transverse vein .generally almost exactly beneath the tip of the first longitudi- nal vein ; the posterior transverse vein not approaching the border of the wing. The nearest genus is Ccenia, differing, however, from Scatella by the pectinated bristle of its antenna?. 1. S. favilBiicca LOEW. 9- — Cinerea, facie alba, alis cinereo-hyalinis, obsolete quadriguttatis. Ashy-gray ; face white ; wings grayish-hyaline with four indistinct clear drops. Long. corp. 0.12 — 0.13. Long. al. 0.13. This species resembles most the European S. sorbillans Hal., which is identical with S. argyrostoma Stenh., but differs from it by its more considerable size, more roughly haired face and gray color of the dust on the posterior part of the cheeks, on the infe- rior part of the occiput, on the pleura? and under side of the abdo- men, on all which parts it is whitish in that species; S. favillacea wants also the clear drop lying beyond the posterior transverse vein in S. argyrostoma. The upper side of the whole body is covered with grayish-brown dust, which on the middle of the tho- rax and on the scutellum does not conceal the shining of the ground color; the large spot lying on the middle of the front is shining greenish. Face very convex, dusted with snowy white, with rather rough hair and the usual row of curved upwards bristles. Open- ing of the mouth wide, with distinct black cilia on the borders. Antennas black ; the pubescence of the bristle is a little longer and more distinct than in most species of this genus. Upper side of the thorax not distinctly striped. Pleura? dusted with yellowish- gray, on their superior border with rather brown dust. Under side of the abdomen, femora, and upper side of the tibia? with gray dust. Wings clouded with grayish, having black veins ; of the five clear drops, peculiar to so many species of this genus, that SCATELLA. 171 lying beyond the posterior transverse vein is entirely wanting, and the remaining are rather indistinct ; the second segment of the costa is at least four times as long as the third; the third and fourth longitudinal veins are parallel towards their ends. Hub. Middle States. (Osten-Sacken.) 2. S. lllgeiis LOEW. £ and J> . — Nigra ; alse iiigricantes, guttis hyali- nis quinque in disco duabusque obsoletioribus in apice pictse. Black ; wings blackish with five clear drops in the middle and two more indistinct ones towards the tip. Long. corp. 0.11. Long. al. 0.13. It differs from the European ^ca^. stagnalis only by somewhat more acute wings, its somewhat less convex face, and the stripes of the thorax being a little more distinct ; perhaps on examining a larger number of specimens it may prove to be only a variety of it. Black ; face with brownish-gray dust, rather convex, only a little impressed beneath each antenna, hairy and bristled, with distinct black cilia on the border of the mouth. Antennae black ; the bristle with an excessively short, but yet distinct pubescence. Cheeks exceedingly narrow. Front dusted with grayish-brown, the spot on the middle of it a little glittering. Upper side of the thorax likewise dusted with brown, but not without all gloss, with two distinct whitish-gray longitudinal stripes, but little distant from each other and beginning on the anterior border, but not reaching nearly to the posterior border ; besides there are two short lateral stripes of the same color, beginning at the shoulder- corner. Scutellum of the color of the upper side of the thorax, only a little more glossy. Abdomen black, rather glossy towards the end : the fifth abdominal segment of the male is almost twice as long as the fourth. Legs entirely black. Wings clouded with black, having five glassy drops on the middle, in the usual position ; besides there is an obsolete spot, forming an indistinct clear drop near the border of the wing between the tip of the second and third longitudinal veins, and another still less perceptible spot in the cloudy color beyond the tip of the third longitudinal vein. Hob. Middle States. (Osten-Sacken). 172 DIPTERA OF NORTH AMERICA. 3. S. Obsoleta LOEW. 9 . (Translated from Berl. Entom. Zeitschr. 1861, p. 358, by R. Osten-Sacken.) — Opaca, capite toto, scutello pleurisque ex flavo, pectore et abdomiue ex cano cinereis, antennis pedibusque nigris ; alse hyalinse, dilute cinerascentes, guttis lirnpidioribus quinque obsoletis- simis. Opaque, the whole head, scutellum and pleurae yellowish-cinereous, pectus and abdomen hoary-cinereous, antennas and feet black ; wings hyaline, with a pale cinereous tinge ; five almost obsolete clear spots. Long, corp. 0.07. Long. al. 0.09. Head altogether yellowish-cinereous, antennae black, face very vaulted,* peristoma ciliated with moderate hairs. Thorax conco- lorous with the head, pectus subglaucous. Scutellum yellowish- cinereous. Abdomen hoary-cinereous, subglaucous, opaque. Legs altogether black, slightly pollinose with white. Halteres impure yellow, stem brown. Wings hyaline, tinged with very pale cine- reous, marked with five clear very obsolete spots ; transverse veins not infuscated ; second costal segment more than thrice longer than the third. Hob. Washington. (Osten-Sacken.) * The original has fornicatus, which means forming a rounded arch with an empty space below. — 0. S. V. ON THE NORTH AMERICAN CECIDOMYIDAE. BY BARON R. OSTEN-SACKEN. It is a peculiarity of the family of Cecidomyidcs that its natural history has always been studied in close connection with its classi- fication. This is owing chiefly to the fact that the gall, the produce of the insect in its first stage of life, is generally a more striking object in nature than the insect itself. The latter small, tiny, dif- ficult to preserve on account of their extreme delicacy, still more difficult to distinguish from their congeners on account of the uni- formity of their appearance and coloring, would afford a very un- satisfactory object of study, unless in connection with the varied deformations which their larvte produce on plants. The study of this family, different in this respect from most of the other families of insects, cannot be prosecuted apart from the observation of living nature, and for this very reason will always be a monopoly of the naturalist so situated as to afford such observations. The aim of the present paper is to direct the attention of American entomologists to this most interesting subject, by giving an account of the observations already made on the North Ameri- can Cecidomyidce, as well as a general introduction to the study of the habits and the classification of this family. The latter has been extracted chiefly from the two following admirable monographs: — LOEW, Dr. H. Dipterologische Beitrage, Part fourth, Posen, 1850, with a plate. • (Contains a monograph of the European Cecidomyidee.} WINNEETZ, J. Beitrag zu einer Monographic der Gallniiicken. In the Lin- naea entomologica, Vol. VIII, Berlin, 1853, with four plates. I. On the classification of the CECIDOMYID^;. In the sketch of a systematical distribution of the Diptera, given by Prof. n. Loew in this volume, he has mentioned the difficulties attending a sharp definition of this family, and has shown that it may be naturally divided in two sections. The species embraced in the first section, which he calls Cecido- myina, have four longitudinal veins on the wings, the last two of 174 DIPTERA OF NORTH AMERICA. which often coalesce in the beginning of their course, forming a more or less distinct fork. They have no ocelli, and the first joint of their tarsi is ranch shortened. The second section, which Prof. Loew calls Anaretina, has one longitudinal vein more, which is inserted between the second and third veins of the first section; this supplementary vein is simple in Campylomyza and furcate in all the other genera. The first tarsal joint is not shortened, and in all the genera, with the excep- tion of Cecidogona, there are distinct ocelli. The first section, which contains all the gall-producing Cecido- myidce at present known, comprises two genera of Meigen and a third genus, discovered by Mr. Winnertz, and of which but a sin- gle species is described. These three genera are easily distin- guished by the neuration of their wings, which are always pubes- cent, and may be characterized as follows : — Cecidomyia Meig. Three or four longitudinal veins ; in the first case the third vein is forked, thus representing the third and fourth veins, which are coalescent in the greater part of their extent (figs. 1, 2, and 4; in some rare cases a branch of this fork or the whole Fig. 1. Fig. 2. Fig. 3. Asynapta. Epidosis. fork becomes obsolete, as in fig. 3) ; in the second case all the four veins are simple (fig. 5). Surface of the wings hairy ; margins with long cilia. Antennae long, moniliform or cylin- drical, generally verticillate, seldom without verticils, from 13 to 36-jointed. CECIDOMYIA. 1Y5 Spantocera Winn. Three longitudinal veins, which are all sim- ple (uot forked); the first close by the costa, the second at some distance from it, but reaching the margin of the wing- before its tip (fig. 6). Hairs on the surface of the wing scaly. Antenna? filiform, 13-jointed, joints elongated, Spauiocera. cylindrical, with a short pubescence and without verticils. Lasioptera Meig. Three longitudinal veins, the first and second of which run very near the costa and are so closely approximated as to be hardly discernible (fig. 7). Wings rather short and broad. An- Fig> 7* tennfe from 16 to 26-jointed ; joints subglobular, sessile, with short verticils. (The sub-genus Clinorhyncha Lw. has been formed of the Lasiopterce, having the mouth prolonged in a rostrum.) The considerable number of. species contained in the genus Ceci- domyia Meig. and the great variety of their structure have made a subdivision necessary. The following tabular arrangement of the sub-genera now adopted has been extracted, with a few modi- fications, from Mr. Winnertz's work, although the subdivision itself is chiefly due to Mr. Loew. CECmOMYIA MEIG. I. Wings with three longitudinal veins, the third either forming a fork (figs. 1, 2, and 4), or becoming more or less obsolete towards the tip (fig. 3).* A. Cross-vein placed between the root and the tip of the first longitudinal vein, as in figs. 1 — 3 : (in this section the cross-vein is frequently almost obsolete.) Cecidomyia Loew. The second longitudinal vein reaches the mar- gin of the wing a little before its tip (although in most cases this distance is very short, as in fig. 1). Generally the same number of joints in the antenna of the J1 and 9; joints either * In examining the wings of the Cecidomyiae, care must be taken not to mistake for a vein a longitudinal fold which generally exists between the second and third longitudinal veins. 176 DIPTERA OF NORTH AMERICA. pedicelled or sessile (sometimes pedicelled in the <£ and sessile in the 9 ; sometimes of the same structure, pedicelled or ses- sile, in both sexes). Diplosis Loew. The second longitudinal vein reaches the margin of the wing at or beyond its tip (fig. 2). Antennae of the male 26 (2 + 24) jointed, sometimes with one rudimental joint more; joints pedicelled, simple joints alternating with double ones (Tab. I, f. 11 and 12), seldom all joints simple. Antennae, of the 9 14 (2 + 12) jointed, sometimes with one rudimental joint more; joints pedicelled, cylindrical. Asphondylia Lw. The second longitudinal vein reaches the margin of the wing a little beyond its tip (as in fig. 2). Antennae of both sexes with the same number of joints; the latter, cylin- drical, sessile, with a short pubescence and without verticils. (A single European species is known.) Hormomyia Lw. The second longitudinal vein reaches the margin of the wing either at or beyond the tip. Thorax more or less gibbose, frequently extending over the head in the form of a hood. Joints of the £ antennas pedicelled ; those of the 9 pedicelled or sessile. Colpodia Winn. The second longitudinal vein forms a curve before the cross-vein and joins the margin a little beyond the tip of the wing (fig. 3). Cross-vein rather large, oblique. (A single European species is known in the female sex only; the joints of its antennae are pedicelled. This sub-genus, which is un- known to me, must be very difficult to distinguish from Epi- dosis.) B. Cross-vein very oblique, originating at the root of the first longitudinal vein (fig. 4).* Dirhiza Lw. Second longitudinal vein hardly undulating before the cross-vein; joints of the antennae sessile or almost sessile in both sexes. (A single species is known.) * The sections A and B, as denned by MM. Loew and Winnertz, seem to be somewhat difficult to distinguish. According to the latter, the cross vein in the section B almost assumes the appearance of an intercalary longitudinal vein; it begins at the root of the first longitudinal vein, runs, although very indistinct, alongside of it and then turns obliquely towards the second longitudinal vein, which thus almost appears to be its continu- ation, or, in other words, to have two roots. CECIDOMYIA. 177 Epidosis Lw. Second longitudinal vein sinuose before the cross- veiu (fig. 4) ; joints of the antennae pedicelled in both sexes; their number variable. II. Wings with four longitudinal veins (fig. 5). Asynapta Lw. The cross-vein is sometimes like that in section A, and then the second longitudinal vein is not sinuated ; some- times as in section B ; then the second longitudinal vein is sinuated, like in Epidosis (fig. 5) ; in this case also the collare is a little prolonged. The classification of the section Anaretina Loew, is very imper- fect ; almost nothing is known about their habits, and even their position in the system is doubtful. Mr. Loew considers them, at least provisionally, as a sub-section of the Cecidomyidee, whereas Mr. Winnertz prefers to isolate them as a distinct family, placed between the Cecidornyidae and the Mycetophilidte, and having many points of relationship to both. Following the authority of Mr. Loew in this volume (p. 7), I will confine myself to the enumeration of the genera which he refers to this section, adding only short sketches of their characters as I find them in the former writers. I. Ocelli extant ; Wings bare or almost bare ; third longitudinal vein forked, the two following veins simple. Antennae 16-jointed; ^ verticillate, joints pedicelled; £> pubes- cent, joints sessile; branches of the fork of the 3d longitudinal vein very arcuated at base (fig. 8, wing). ZYGONEUKA Meig. Antennae 9-jointed, short, slightly pubescent ; joints subsessile, sub- globose (fig. 9, wing). ANAEETE Hal, Wings pubescent ; Third longitudinal vein forked. The upper branch of the fork forms a double curve, almost in the shape of an S ; (see Plate I, fig. 13.) TRITOZYGA Lw. The upper branch of the fork forms a single smooth curve ; £ antennae 16-jointed, verticillate, joints pedicelled; £ antennae 10-jointed, pilose, joints moniliform (fig. 10, wing). CATOCHA Hal. 12 178 DIPTERA OF NORTH AMERICA. Fourth longitudinal rein forked ; antennse 11-20-jointed ; £ moni- liform, pilose ; joints pedicelled ; 9 subrnoniliform, joints sessile, pubescent (fig. 11, wing). CAMPYLOMYZA Meig. II. Ocelli wanting ; third longitudinal vein forked ; first longitudinal vein very short ; wings pubescent ; antennse £ moniliforin, verticillate ; £> submoniliform, pubescent. Antenna; 16-jointed (fig. 12, wing). LESTREMIA Macq. Antenna; 11-jointed. CECIDOGONA Lw. Fig. 8. Fig. 9. Fig. 10. Campyloniyza. Lestremia. For further details as well as for the references, see Walker, Diptera Britannica, Vol. Ill, which also contains beautiful figures of all the genera. As to the new genus Tritozyga Lw., formed on an American species, Mr. Loew thus characterizes it in a manuscript note of his : — " The whole structure of its body shows the nearest relation to Gampylomyza; the form of the legs and wings is as in that genus; the wings (Tab. I, fig. 13) have the same short pubescence and cilia, and the three ocelli are just as distinct. The differences are the following. 1. The vein, which in the second section of Cecidomyida; is added to the number of the veins of the first section, is not simple, but forked, in the new genus, and therefore approaches in some measure the genus Anarete; 2. The very thick longitudinal vein is not forked as in Campylomyza, but simple. The new genus cannot be confounded with Anarete, the species of which have a much more slender structure, a very elongated first joint of the tarsi and the third longitudinal vein of the wing bipartite as far as the base. From Lestremia and Cecidogona it differs in quite a similar manner, and besides by the presence of ocelli, which are wanting in both genera. The antenna? are mutilated in the single specimen which I have before me (a male from the District of Columbia), therefore I can say nothing of the number of their joints ; their CECIDOMYIA. 179 structure is as in most Campylomyzce. The number of the joints of the antennae being of a higher value, among the Gall-gnats, for the distinction of species than for that of genera, since almost every genus comprises species with very different numbers of joints of the antenna?, I abstain from giving a name to the species known to me so incompletely ; as to the genus, which can never be mis- taken, I propose for it the name of Tritozyga." II. On the habits of the CECIDOMYIDJS. The food of the larva? of Cecidomyia is of a vegetable character. A few apparent exceptions will be mentioned below. They furthermore seem to live in preference on living plants; neverthe- less several species of the subgenera Epidosis and Diplosis, have been reared by Mr. Winnertz from decaying wood ; Gee. fuscicollis Meig. (?) has been reared by Bouche from decaying bulbs of tulips and hyacinths. (Instances like that of Gee. bicolor Bouche, found in dung during winter, must be received with caution, as the larva? may have gone there for transformation only.) Although the majority of these larva? attack the soft and green parts of plants, some of them live under the bark of trees, in the cones of pines (Cec. strobi Kalt. ) or in fungi '(Diplosis polypori Wz., Asynapta lugubris Wz., etc.) Again, most of the larva? are monophagous, that is, each species lives exclusively on a certain species of plant, or, at least on closely allied plants; Mr. Winnertz remarks that even those found under the bark of trees follow the same rule. Exceptions are Cec. sisymbrii Schr., which, according to Mr. Winnertz, inhabits in May and June a gall on Berberis vvlgaris, and from June till No- vember a somewhat different gall on Nasturtium sylveslre (Winn. 1. c. p. 209 and 231) ; Gecid. arcuata Wz., has been found in the pappus of different syngenesists, in decaying wood and fungi. Besides these, there is a class of larva? which live as guests or parasites in galls formed by other Cecidomyia (Cec. acroplnla Wz. and pavida Wz. live socially in the deformed buds of Fraxinns excelsior; Diplosis socialis Wz. inhabits the gall of Lasioptera rubi ; Dipl. tibialis Wz., has been reared from the same gall with Cec. salicina Schr., etc.); or by Acari (Cec. peregrina Wz., and similar cases, observed by Loevv.) Some even live in the society of Aphides. According to Mr. Winnertz the larva? of the sub- genus Diplosis principally, share these parasitical habits; even 180 D1PTERA OF NORTH AMERICA. those living under the bark of trees or in fungi are seldom found alone, but for the most part in the society of other larvae (Winn. 1. c. p. 206). Thus, the larva of a species of Diplosis has been found in a stem of Saroihrium scoparium, together with larvae of Hyhsrnus and Apion. Among the larvae with an exceptional mode of life, those should be mentioned which live on the surface of the plant, as that, ob- served by Mr. Loew on the leaves of Veronica Beccabunya, or those of Diplosis ceomatis Wz., and D, coniophaga Wz. found on the leaves of a rose-bush overgrown with the fungus Ceoma miniatum, on which they feed. The American species, probably also a Dip- losis, which I have called Cec. glutinosa, and which will be described below, has a similar mode of existence on the surface of hickory leaves. The greater number of larvae penetrate inside of the plant, so as to be concealed from view during their development. Their presence is generally indicated on the outside of the plant by some deformation. Every part of the plant, from the root to the flower and the fruit, is liable to such attacks. But each species of Ceci- domyia always attacks the same part of the plant, and deforms it in the same way. (Exceptions seem to be rare; Cec. tremula Wz. has been reared from two galls of different shape, both found on the poplar ; the insects differed only in size). The deformations thus produced are very numerous, and several unsuccessful attempts have been made to classify them according to the nature of their origin and their shape. At one extreme of the series is the true gall, a vegetable growth of constant and de- finite form, attached to the plant by a very small portion of its . surface and not otherwise deforming that part of the plant (of this class are, for instance, the numerous galls, described below, ou the leaves of the hickories) ; at the other extreme is the simple defor- mation, folding of a leaf, swelling of a leaf-rib, arrest of the growth of a bud or a stalk, etc. The egg of Cecidomyia is elongated, rounded at both ends, orange-yellow, or whitish. The time within which the larva is hatched is very different, and depends on the state of the weather; in a great heat, the hatching sometimes takes place within a few hours ; generally a few days are required. Mr. Loew supposes, however, that the species having but one yearly generation remain much longer in the state of eggs. CEC1DOMYIA. I Si When first hatched, the larva is colorless, transparent, with a translucent green, yellowish or red stomach; later in life it assumes different shades of red (orange, pinkish, cochenille-red) or becomes yellow or whitish; the color of the same species may also some- what change with age. All these larvce have the extraordinary number of fourteen joints, thus affording an apparent exception from all other larvaj of insects, which, as a general rule, have thirteen joints. The supernumerary fourteenth joint is placed between the head and the first thoracic (stigma-bearing) segment. It may be considered either as a part of the head, or as a prolongation of the first thoracic segment. Number and position of the stigmata are normal ; one pair on the first thoracic segment, and eight pairs on the first eight abdominal segments, so that the ninth or last segment bears none. Sometimes the last pair of stigmata is removed from its usual lateral position, more towards the middle of the segment. In one case (Gee. pini Deg. and the American G. pint inopis) this last pair is placed apparently on the last segment; but this segment is in reality the eighth, the ninth segment being in this larva unusually small and concealed under the eighth. The stigmata are horny, more or less nipple-shaped projections. The skin of most larvas appears finely chagreened under a strong magnifying power; in some cases it is perfectly smooth. The dorsal segments of Gee. sarotliamni Lw., G. genista; Lw., etc. are uneven ; those of G. craccce Lw., G. quercus Lw., G. fuscicollis Bouche, etc. are furnished with bristles or sparse hairs ; those of G. entomophila Ferris with hairs arranged in regular rows ; those of Gecid. pini Degeer, and of two larva? which I found in this country {Gee. pini inopis 0. S. and Gee. glutinosa, nov. sp.), have rows of fleshy, setiferous caruncles along the back. (It is to be noticed here that both Degeer and Dufour, in describing such larva?, mistook the back for the venter, and described these carun- cles as pseudopods. See Deg. Mem. VI, Tab. XXYI, fig. 9 — 19, and Dufour, Ann. Soc. Ent, de Fr., 1838, p. 293). The last abdominal segment is smooth and rounded, or furnished with two setiferous tubercles {Gee. pini), sometimes uneven and bristly, or excavated, or armed with a pair of horny processes, frequently curved upwards. Dufour saw a larva use these pro- cesses for leaping. The structure of the head and of the organs of the mouth is but imperfectly known. What Mr. Katzeburg saw (see his paper 182 DIPTERA OF NORTH AMERICA. in Wiegmann's Archiv, vol. vii. p. 233, with a plate) and what I have found confirmed by my own observations, may be reduced to the following : The horny parts of the head consist of a ring with two processes extended backwards; a soft, fleshy swelling which protrudes through this ring is taken by Ratzeburg for the labium ; two openings in the tipper part of the ring emit a pair of two- jointed organs which this author and L. Dufour believed to be palpi, but which I would rather consider as rudimental antennas, especially on account of their position on the upper side of the head. (Laboulbene and Perris entertained the same view.) On the under side of the body, at the juncture of the first thoracic segment with the supernumerary (14th) segment, there is a horny, more or less elongated piece, projecting with its anterior part, whereas its posterior end is concealed under the skin of the first thoracic segment, and more or less translucent. This organ, the use or the homology of which is unknown, is peculiar to the larva? of Cecidomyia, and seems to be seldom wanting. (I found under the bark of a tree a full-grown larva which, for its structure I believe to be a Cecidomyia, although it showed no trace of this breastbone.) It may be that this organ is used for locomotion, although I hardly would consider it as homologous to the pseudo- pods of the larvas of Chironomus and Ceratopoyon. If the sup- plementary (14th) segment be considered as a part of the head, this breastbone might be taken for the menturn, in^analogy to the horny mentum of the larvae of the Tipularice. The form of this organ is variable in different species ; sometimes it ends anteriorly in two points, with an excavation between them ; sometimes in one elongated point ; or it is serrated, etc. The remaining part of the under side of the body sometimes shows other organs of locomotion. The larva of Gee. entornophila, according to Perris, has three slender, elongated, pointed, sub- corneous, approximated projections in the middle of every ventral segment. Gee. fu&cicollis Bouche (Bouche, Naturg. der Ins. p. 25), has a pair of elongated, pointed pseudopods under each thoracic segment, and three such pseudopods under each abdomi- nal segment. Bouche's figure of the latter closely resembles Perry's figure of the pseudopods of Gee. entomophila. The motions of the larvaa, except those few, living on the surface of the leaves, are generally slow ; but those which change their abode before assuming the pupa state become very active about CEC1DOMYIA. 183 that period. Winnertz observed an extraordinary activity in some such larvae after a thunder storm ; they left their hiding-places un- der ground, and crawled about restlessly for some time ; they did the same after every thunder storm, some of them even two mouths after having left their galls. The larvae of several species, for instance, Cec. loti, Gee. pisi, and Cecid. rumicis, have the power of leaping. Mr. Loew remarks that all such larvae belong to the sub-genus Diplosis. Cec. popidi Duf. performed its leaps by straining the horny hooks at the tip of its abdomen against the under side of the thoracic segments. (Dufour, Ann. Sc. Nat., 2e se"r. XVI, p. 257.) " The want of horny organs of mastication," says Mr. Winnertz, "authorizes the supposition that a lesion of the plant does not take place ; it is much more probable that the larva has the power of producing in the plant some peculiar irritation, which causes an overflow of the sap necessary for its food. How little the larva requires for its support is evident from the circumstance that it attains its full growth and development in a gall just large enough to inclose it, a gall apparently hermetically closed, for the most part with hard walls, which do not show the least sign of internal lesion. It seems even as if a certain amount of moisture alone was sufficient to sustain these larva?, especially when a great num- ber of them live socially in the same gall (from ten to fifteen larva? in the pea-sized bud of Cardamine pratensis; from fifty to sixty in another kind of gall, etc.). Another proof of the small quan- tity of nourishment required by these larvae is, that no excrements are to be found in their place of abode." "The only exception known to me of this extreme frugality," says the same author, "are the larvae of two species which live on the leaves of the white rose, attacked by the fungus Ceoma minia- tum. These larvae not only lick the sap exuding at the bottom of the heaps of spores, but they also greedily consume the spores themselves, and their intestinal canal is always filled with them." The observation of Vallot (Mem. de Dijon, 1827, p. 95), that a larva of Cecidomyia (C. acarivora) found on the surface of the leaves of Chelidonium feeds by sucking Acari, as yet requires con- firmation. Winnertz saw Cecidomyia-\arv& living as guests in deformations produced by Acari, greedily lick their hosts, but he never found in such galls an empty skin of an Acarus. As to the larvae of Cecidomyia inhabiting galls, produced by other species 184 DIPTERA OF NORTH AMERICA. of the same genus, it is a question, according to Winnertz, whether they take the same food with their hosts, or live on their excre- ments. Ferris (Mem. de Lille, X, p. 274, with figures) found Cecid. entomophila in an insect-box, living on the excrements of the minute Acari abounding in such boxes ; he compares them, apparently with good reason, with the larvsa of Cecidomyia found under the bark of trees, among the excrements of the xylophagous insects. These larvae underwent their transformation in the cor- ners of the box. It is very probable that the larvae of Cecidomyia, like most of the dipterous larva?, do not undergo several moultings. I do not find any mention about it in the authors. Only Dr. Harris states that C. tritici casts off its skin before going under ground for transformation. Before assuming the pupa state, some larvae of Cecidomyia leave their galls and abscond themselves under ground, under dry leaves or moss, or under the bark of trees. Other larvae, on the contrary, undergo their transformation within their gall.* In both cases the pupoa are frequently, although not always, inclosed in a cocoon. Winnertz positively denies that the larvse spin this cocoon ; ac- cording to his observation, the latter is, so to say, exuded by the larva. He found that larvse which had fastened themselves to a leaf, were encircled within twenty-four hours by a white halo, con- sisting of tiny thread-like particles, which seemed jo grow some- what like crystal-needles ; the larva during this time remained perfectly motionless. The cocoon is perfected within a few days, and even then, under a strong magnifying power, no genuine thread is perceptible. The mode in which the papa state is assumed has been de- scribed by Dr. Harris in a posthumous paper published in the Proceedings of the Boston Soc. of Nat. Hist., 1860, p. 179. "The approaching change is marked by an alteration of the color of the anterior segments of the larva, which (in the case of Cecid. salicis Fitch) from orange become red and shining, as if distended by blood. Soon afterwards, rudimentary legs, wings, and antennae begin, as it were, to bud and put forth, and rapidly grow to their full pupal dimensions, and thus the transformation to the pupa is The larva of Cecid. terminalis Lw., according to Winnertz, varies in its habits. It sometimes goes under ground, and sometimes traiisforms within the willow leaves deformed by it. CECIDOMYIA. 1 85 completed." The peculiarity of this process is, that the transforma- tion is undergone without shedding the larva skin, and, as the same observation has been repeated by Dr. Harris on the larvae of C. destructor Say and G. tritici Kirby, it is very probable that it ap- plies to all the larvae of the genus. I do not find this fact men- tioned in the European authors. Instead of a cocoon, the pupa of Cec. destructor Say is inclosed in an oblong, brown case, which is nothing but its own hardened pupa-skin. "The larva of this insect, says Dr. Harris (I. c.), when it has come to its growth, remains fixed and motionless on the culm of the wheat. Its body contracts and soon takes the form and color of a flax-seed. While this change is going on externally, the body of the insect gradually cleaves from its outer dry and brownish skin. When this is carefully opened, the included insect will be seen to be still in the larva state. It does not change its condition until a few days before it discloses the winged insect," etc. Cecid. graminicola Kalt. and another Cecidomyia, mentioned by Dr. Fitch as forming an imbricated gall on Agrostis lateriflora, undergo a similar kind of transformation, their pupae being inclosed in the dry larva-skin. However different the mode of transformation of Cecid. pini Deg., C. pini maritimce Duf. , and Cec. pini inopis 0. S. may ap- pear, the pupoa, of which are inclosed within a cocoon of resin, it is in perfect analogy with the preceding instances. The process by which the cocoon is formed is exactly the same as that described above by Dr. Harris. The larva of the American species C. pini inopis 0. S. observed by me in the environs of Washington, fastens itself to a pine leaf and remains motionless until the resinous sub- stance which it exudes abundantly, begins to harden ; the larva then gradually frees itself from the contact of the cocoon-like case thus formed. It is very probable that this cocoon is nothing but the outer larva-skin, saturated with resin. The pupae of Cecidomyice show a close resemblance to those of the Tipvlce fungicolce, especially those of Sciara. As in the latter genus, the bases of the antennas are often produced in points ; these frontal projections are sometimes long, approximated, and resemble horns (Cec. sarothamni, see Winnertz, 1. c. Tab. I, f. 6, or Cec. verbasci Dufour, Ann. Sc. Nat., 3e se'r., Yol. IY, p. 5-24, with figures, 1845); in other cases they are smaller and at some 186 DIPTERA OF NORTH AMERICA. distance from each other (Cec. salicina, 0. veronica, etc., see Wz., 1. c. f. 3, 4). Behind these horns, two pairs of bristle-like pro- cesses may be observed in most pupae. The first pair is also on the head, close by the horns, the second on the thorax. Both vary in size and strength in different species. The second, tho- racic pair, has been taken by some authors for a spiracle. These projections and horns, especially the frontal ones, aid the pupa in working its way through the gall or from underground, before entering its last stage of existence. The dorsal segments of the abdomen are, for the same purpose, frequently rough with spines. The tip of the abdomen is sometimes smooth; in other instances it bears a few bristles. After the exclusion of the perfect insect, the pupa-skin remains frequently hanging on the outside of the gall. Some species of Cecidomyia have only one, others more than one yearly generation. The summer generation of the latter kind remain but a short time in the pupa state; the winter generation much longer. The Iarva3 of Lasioptera resemble those of Cecidomyia in their structure as well as in their habits. They frequently have the same reddish color and the peculiar breast-bone. L. rubi Ileeger and the N. American L. vitis O. S. produce swellings in the stem of the plants which they inhabit. L. pusilla Heeger forms galls on the leaves of Sonchus, L. cerris Kollar on those of the oak ( Quercus cerris). III. On the North American CECIDOMYIA hitherto observed and their galls. The species of N. A. Cecidomyice at present known, may be distributed into three categories, according to the extent of our knowledge concerning them. About the species of the first cate- gory, nothing but the description of the perfect insect is extant, its habits remaining unknown ; as to those of the second category we are acquainted with the first stages of their existence, especially with the deformations they produce, without knowing the perfect insect; finally, to the third category belong those, the habits of which, as well as the perfect insect, are described. The following is a synopsis of the species recorded by previous authors, as well as of those mentioned in this paper: — CECIDOMYIA. 187 I. Perfect insect described, habits unknown. C. ornafa Say, Long's Exped. App. p. 357. Wied. Auss. Zw. I, 22, 2. C. caliptera Fitch. C. cerealis Fitch. }• Dr. A. Fitch, Essay on the Wheat-fly, etc. C. tergata Fitch. C. thoracica Fitch. J C. spongivora Walk. List of Dipt. Brit. Mus. I, 30. Campylomyza scutellata Say, Journ. Acad. Phil. Ill, 17, 1. Wied. Auss. Zw. I, 22, 1. Tritozyga, sp. Lw. (see p. 178). Diplosis maccus Lw. The following note on this new species was furnished by Mr. Loew : — " Gall-gnats cannot be recognizably described from single dried specimens, unless they are distinguished by some striking peculiar- ities. I feel no temptation at all to describe species which have no such peculiarities, and allow myself an exception only with the following Diplosis on account of its remarkable beauty. D. macciis LOEW. £ and J. (Tab. I, figs. 11 and 12.)— Flavida, thorace fusco-vittato, antennarnui artieulis nigris et pallidis alternanti- bus, alis violaceo-maculatis, tibiis tarsisque nigro-annulatis. Yellowish, thorax with fuscescent stripes ; the joints of the antennae alter- nately black and whitish ; wings with violet-blue spots ; tibiae and tarsi annulated with black. Long. corp. 0.08. Long. al. 0.11. Yellowish ; the joints of the antennae alternately black and yellowish-white, the simple joints being black, the double joints yellowish-white ; also the hairs of the black joints are black, and those of the light ones light. Thorax with three brown longitu- dinal stripes coalescing anteriorly, the intermediate one reaching only to the middle of the thorax, the lateral ones running as far as its posterior margin. Abdomen without dark bands. Coxa3 yel- lowish. Fore and middle femora black on the upper side and tips, the outermost extremity of the tip being yellowish ; hind femora with a black line not reaching far beyond the middle, and with the tips black. Tibiae black ; anterior ones with a very broad yellowish ring beyond the middle, the hind ones with such a ring at their base and a second ring beyond the middle. Anterior tarsi black on the first, short joint, at the base of the second and at the tips of the second, third and fourth joints ; the hind tarsi have the same markings with the exception of the base of the second 188 DIPTERA OP NORTH AMERICA. joint, which is not black. Poisers yellowish, the base of the knob blackish. Wings yellowish, appearing almost golden yellow in an oblique direction, with bright spots of a violet reflection. Before the second longitudinal vein there are two such spots, the first immediately beyond the tip of the first longitudinal vein, the second between the first and the tip of the second longitudinal vein. Between the second and third longitudinal veins there are three violet spots, the first of which is the largest ; it is situated under the first costal spot and runs far towards the base of the wing in the form of a wedge without sharp limitation ; the second is the smallest, and is placed below the yellowish space between the two costal spots ; the third is a double spot almost S shaped, and united to the second costal spot with its anterior end. Behind the third longitudinal vein the violet color prevails to such an extent as to leave only two golden spots, one of which is placed behind the anterior branch and the other immediately behind the posterior branch of the third longitudinal vein. The cilia of the wings are quite pale yellowish, but blackish where the violet spots reach the margin of the wing. Hob. Washington. (Osten-Sacken.) This species resembles very much the European Diplosis pavo- nina Loew, but is easily distinguished from it by the smaller ex- tent of the violet color of the wings, the smaller extent of the black color of the legs, and the sharper limitation of both colors. Whether the male of Diplosis pavonina has likewise the joints of the antenna? alternately dark-colored I do not know, as I did not succeed in discovering it; judging, however, by the appearance of the antennas of the female, this does not seem to be the case." II. Galls or larva? known, perfect insect unknown. (The descrip- tion of these galls and larva? is given below, under the indi- cated numbers.) On hickories, Carya, of different kinds, seven species, besides one belonging to the third category. (Nos. 1 — 8.) On the golden-rod, Solidago, of different kinds, two species, besides other two belonging to the third category. (Nos. 9 — 12.) On Vaccinium (or Gaylussacia ?), one species. (No. 13.) On the scrub pine (Pinus inops), two species. (Nos. 14 and 15.) On the red maple (Acer ruhrum"), one species. (No. 17.) On the ash (Fraxinus arnericana), one species. (No. 18.) On the oaks of different kinds (Qaercus'), four species. (Nos. 19 — 22.) CECIDOMY1A. 189 On the wild grape (Vitis~), one species (No. 24), besides another belonging to the third category. On the hornbeam (Carpinus americana*) one species. (No. 25.) On the tulip-tree (Liriodendron tulipiferum),iwo species. (Nos. 26 and 27.) On the willow (Salix), one species (No. 28), besides one belonging to the third category. On Impatiensfulva, one species. (No. 30.) On the blackberry (Rubus i-illosus), one species. (No. 31.) On Agrostis lateriflora (?), one species. (No. 32.) III. Perfect insect described, and its habits known. On the cereals (wheat, rye, etc.). f About the habits of these well-known insects, see C. destructor Say. j Dr. Harris's Treatise, etc., and Dr. Fitch's papers : C. tritici Kirby. The Hessian Fly (Trans. N. Y. State Agric. Soc., I vol. VI), and The Wheat Fly (ibid. vol. V). C. culmicola Morris. See Dr. Harris's Treatise, p. 465. On the locust (Robinia pseudoacacia) . C. robinix Hald. Amer. Jouru. Agric. and Sc., vol. VI, 193. Harris, Treatise, etc., p. 452. (Haldeman's paper is also reproduced in the Proc. Boston Soc. of Nat. Hist., vol. VI, January, 1859.) The larva lives upon the leaves, the margins of which it deforms into a roll. It is evident, from Mr. H.'s description of the perfect insect, that it belongs to the sub-genus Diplosis. (See also Fitch, Reports, vol. II, No. 332.) C. pseudoacacix Fitch, Reports, vol. II, No. 331. The larvse injure the tender young leaflets near the tip of the stem, causing them to be folded like a little pod (in July and August). They transform under ground. On the gooseberry (Ribes uva crispa). C. grossularise. Fitch, Reports, vol. I, p. 176, and vol. II, No. 150. The berries turning red prematurely and becoming putrid, contain the bright yellow larvze. Dr. Loew thinks that the perfect insect be- longs to the sub-genus Asphondylia. (See p. 7.) On the willow (Salix rigida and S. lucida). C. salicis Fitch, Am. Quart. Journ. Agric. and Science, vol. I, p. 263. (See also Dr. Harris's paper in Proc. Bost. Soc. Nat. Hist., vol. VII, January, 1860.) The gall is a woody tumor, surrounded by the dry and brittle terminal bud, at the tips of the twigs. It contains but a single larva. The name of the species must be changed, as there is already a European C. salicis. I propose to call it C. riyidie. On the alder (Alnus serrulata). C. scrrulatce 0. S. (See below, No. 16.) On the hickory (Can/a). IHplosis caryae. 0. S. (See No. 1.) On the wild grape. 190 DIPTERA OP NORTH AMERICA. Lasioptera vitis 0. S." (See No. 23.) On the golden rod (Sottdago). C. solidaginis Lw. (See No. 9.) C. hirtipes 0. S. (See No. 10.) On Chrysopsis mariana. C. chrysopsidis Lw. (See No. 29.) In comparing this list with similar enumerations existing for European Cecidomyice, but few cases of analogy will be found. Such cases are, for instance, the habits of C. pini inopis, nov. sp., which correspond exactly to the European C. pini Degeer ; the gall of G. strobilo'ides, nov. sp., on the willow, which is repre- sented in Europe by G. strobilana Brerai; the analogy between the gall on Fraxinus americana (No. 18) and that of C. bolularia Wz. of the European ash, is more doubtful ; likewise that between the deformation of C. erubescens, nov. sp. (No. 20) on the oak leaves, and a similar deformation described by Mr. Loew (C. quercus Lw.) Two galls occur on the American wild grape, whereas none has been discovered on the European grape; likewise, although eight galls are already known to occur on the hickory (Cctrya), none is recorded as belonging to the European walnut (Jng/ans). Al- though galls have been found on the European maple, alder, and blackberry, they are different from those recorded below on the American species of these trees and shrubs. Robinia, Lirioden- dron, and in some degree Solidago, being peculiar to America, their galls could not, of course, be expected to be found in Europe. I will proceed now to give a condensed description of the ob- servations which I had occasion to make on Cecidomyice during my residence in this country. Those observations were made in the environs of Washington, unless otherwise mentioned. I have followed a practice adopted in Europe, in giving names to species known only on account of the deformations they produce, the per- fect insect not having as yet been reared. This affords the advan- tage of being able to designate each described gall by a fixed name. In order, however, to distinguish such species from those the gall-fly of which has been reared and described, the first are simply put down as new species (n. sp.), whereas the names of the authors have been mentioned after the specific names of the latter (Lw. or 0 S.) CECIDOMYIA. 191 1-8. On hickories (Carya) of diiferent kinds. The numerous galls of Cecidomyice occurring on the hickory are found indifferently on the various species of this tree. I have noticed also that whenever a spot is found where one of the galls occurs in abundance, some of the other kinds are sure to be found. Thus the galls of Gee. holotricha and those of Diplosis caryce, or those of the latter with the galls of Cec. tubicola are frequently met with on the same leaflet. These galls may be distributed as follows (the numbers from 1 to 8 corresponding to those of the descriptions given below) : A. True galls, fastened to the under side of the leaf and breaking off easily. a. Bare. 1. Snlglobular, with a small nipple at the tip, diam. 0.05 to 0.1. 2. Elongated onion shaped, a little larger than the preceding. 3. Conical, contracted at base, blood red or purplish. 4. Cylindrical, erect, inserted in a cylindrical socket, b. Pubes- cent. 5. Subglobular, with a nipple at tip (or short onion shaped), pubescent with ferruginous. 6. Subglobular, without nipple at tip, finely downy. B. Other deformations. 7. Swelling of the midrib at the base of the leaf. 8. Yellow spots on the leaves ; larva living on the leaf, not within it. 1. Diplosis caryce O. S. Gall Subglobular, smooth, seedlike, 0.05 to 0.1 in diameter, with a small nipple at the tip. In sum- mer they are yellowish-green and their shell is soft; in winter they become brownish, and the shell, although thin, is hard and woody. They begin to grow in June. I gathered them in October, when the larva was full grown. Each gall contains a single larva; it is white, and stouter in proportion to its length than most larva? of Cecidomyia. The breast-bone has two sharp points anteriorly, with an excavation between them; the tip of the last abdominal segment has no horny processes. It undergoes the transformation within the gall. The pupa resembles, by the structure of its head, that of C. sarotJifnnni Wz., figured by Mr. Winnertz in his monograph (/. c. tab. I, f. f>); namely, the pointed projections at the basis of the antennas are closely approximated and not remote, as in other species. After having kept these galls on moist sand all winter, I ob- tained the fly in April. (Description drawn from a fresh specimen.) D. caryae 0. S. £ and 2.— Antenna; pale; J 26-jointed; alternate joints a little larger than the intermediate ones; verticils 192 DIPTERA OF NORTH AMERICA. moderate; pedicels between the joints rather short; 9 14-jointed, joints subsessile ; front and mouth pale ; collare with a blackish edge posteriorly, ending on both sides in a short, black streak on the pleurae ; thorax pale, with three broad, almost contiguous blackish or grayish stripes ; the intermediate one is subcuneiform and slightly capillary towards its posterior end, which, for this reason, appears slightly bifid ; it does not reach the scutellum ; the lateral ones are rounded anteriorly, narrowed posteriorly, and end just before the scutellum in a short, black streak, communicating with a brown triangle on the side of the scutellum, so that the latter, being pale itself, is inclosed on both sides by the black streaks and the brown triangles; a couple of black dots are visible on the pleurae ; a pale brown spot on the pectus, between the first and the second pair of coxae ; a brown spot at the basis of the lialteres, which are pale; abdomen reddish, hardly darker laterally, and with a tuft of hair on each side, near the posterior margins of the segments ; legs pale, with a minute, appressed black pubes- cence, which makes them appear blackish; wings immaculate; the second longitudinal vein joins the costal at the apex of the wing or immediately beyond it ; cross-vein indistinct or none. 2. C. carycecola, n. sp. Gall somewhat larger than the preced- ing, elongated onion-shaped, with the tip prolonged in a point, pale green. Found through the summer either in separate clusters, or mixed with other galls, for instance that of C. holotricha. 3. C. sanguinolenta, n. sp. Gall conical, narrowed at the basis, blood red or purplish, about 0.15 high and 0.12 broad. I found them for the first time about the middle of July. At this time they were solid inside, except a narrow hollow near the basis which contained the small, somewhat yellowish larva, with a distinct, pointed, spear-shaped breast bone. These galls occur in nume- rous clusters on the same leaflet. 4. C. tuMcola, n. sp. Gall narrow-cylindrical, erect, about 0.15 or more long. They break off easily, being inserted in a small protuberance on the leaf, with a sharp-edged socket in the centre, in which the cylinder fits exactly. Their color, when ripe, is more or less brownish, pale greenish at base. They are hollow inside and contain in October a whitish larva with a breast bone ending anteriorly in a single, elongated point. They generally occur in clusters. Some of these galls are found covered with a viscous fluid. CECIDOMYIA. 193 Early in summer I frequently found a gall of the same form, but smaller, generally reddish at the tip and easily distinguished by the absence of the basal piece in which the other is inserted ; it is simply fastened to the leaf by a minute pedicel. Besides, it occurs always singly, frequently on the edges of the leaves, whereas the other gall is for the most part found in clusters. Is it the same species? 5. C. holotricha, n. sp. Subglobular, pubescent, onion-shaped galls. Diam. up to 0.1 or a little more. They resemble the galls of D. caryce in shape, but are somewhat larger and covered with a pubescence which is pale when the gall is young and growing, and becomes rust-colored in the stage of ripeness. I have observed two modes of occurrence of these galls ; either they are scattered in numbers, as many as a hundred on the same leaflet, or they grow in a row along the mid-rib of the leaflet ; in the latter case they are generally larger, and being packed close together, assume an irregular shape. It is very probable that these two forms belong to two different species, and in this case I would retain the above name to the first form. Galls of the first form begin to grow in June ; in September and October I found the white larva apparently full grown. The breast-bone has one elon- gated point anteriorly and two projections on both sides, about the middle. At the same time I find in my diary that in some of these galls (it is not distinctly stated which), I had found a pale orange, apparently full grown larva, with the breast-bone ending anteriorly in two triangular points with a rectangular excision between them. 6. C. persicoldes, n. sp. Gall round, 0.1 to 0.2 in diameter, smooth, without nipple-shaped tip, yellowish or red, clothed with a delicate down like that of peach, and looking somewhat like a diminutive fruit of this kind. I found these galls more seldom than the others. 7. C. cynipsca, n. sp. Rounded, irregular, hard swelling on the under side of the hickory-leaf, on the midrib, near the base of the leaf, about half an inch long. When I found it (in July) it was pale yellowish, and contained, in several small hollows, minute whitish larva?, with a breast-bone narrowed anteriorly and ending in a point. 8. C. ylntinosa, n. sp. The small yellowish-orange larva forms no gall, but lives in the open air on the under-side of the leaf, to which it is attached by a viscous substance probably secreted by 13 194 DIPTERA OF NORTH AMERICA. the leaf. The presence of the larva is indicated on the other side of the leaf by a round yellow spot. The structure of the larva is peculiar : it has rows of fleshy, pointed tubercles along its back, like the larva of C. pini inopis (described below), with which it agrees in some respects in its habit of fastening itself to the sur- face of the leaf by means of a viscous substance. 9. C. solidaginis Lw. Gall on Solidago produced by the arrest of the growth of the stalk, which causes the leaves to accumulate round the same spot and thus to produce a large imbricated de- formation. It begins to appear already in July, but the flies escape only late in the fall. The following description of gall and fly have been prepared by Mr. Loew : — " The gall (Tab. I, fig. 8) represents a globular head of the size of 1| to 2 inches formed by hundreds of leaves, the exterior ones being only little altered, the interior ones becoming more and more narrow ; on a closer examination we easily perceive that this structure results from the coalescence of several deformations at the tips of abortive twigs ; in a specimen which I dissected I counted five such shortened twigs. At the top of each twig there is a single gall, without compartment, somewhat of the shape of a very small seed, and having in its interior a cavity widened a little underneath. The tip of one of them (Tab. I, fig. 10) showed at its end three small convergent lobes, giving it the appearance of being produced by three coalescent leaves. I could not discover this structure in the others ; I found only a rounded, rather irre- gular opening at the tip. The insect which produces this defor- mation likewise belongs to the genus Cecidomyia in the restricted sense. C. solidagiiiis LOEW. £ and 9. (Tab. I, fig. 4— 7.)— Fusca, ab- domine fasciis rufis et nigris picto ; antenuarutn flagellum in mare arti- culis 20 vel 21, in foemina circiter 18 ; ake pilosse, nigricautes, venula transversa nullo, ; terebra foemiiise modice elongata. Fuscous, abdomen with black and red bands ; flagellum of the antenna? with 20 or 21 joints in the male, with about 18 in the female; wings hairy, blackish, without transverse veinlet ; borer of the female mode- rately long. Long. corp. ^ 0.1G, J 0.17. Long. al. £ and 9 0.16 — 0.17. Thorax with the pleura? sometimes brown, sometimes dark fus- cous, with black hairs. Abdomen of the female with distinct CECIDOMYIA. 195 black and red transverse bands, the latter less distinct in the male ; hairs of the abdomen blackish with a lighter reflection. Antennae of the male with 20 or 21 brown flagellar joints with rather long peduncles, the uppermost being much smaller than the preceding ; the verticillate hairs very long and rather light. The female has generally some flagellar joints less, and its joints are round, with shorter hairs and without any peduncle. The female ovipositor has a very moderate length and is little pointed. Legs of the female black without white reflection. Legs of the male much longer and more slender than those of the female; hind tibia? and tarsi everywhere with a white reflection, which, on the fore and middle tibia? and tarsi, is chiefly seen on the under side. Poisers black. Wings blackish on account of their close and long hairs ; between the first and second longitudinal veins no transverse vein is visible; the second longitudinal vein towards its end is very little curved exteriorly ; the anterior branch of the third longitudinal vein is distinct and nearly straight." (Descrip- tion drawn from dry specimens.) 10. C. hirtipes 0. S. Rounded gall at the tip of stunted stalks of Solidago, sometimes nearly an inch in diameter, smooth, brown- ish on the outside, solid inside, containing several larvae in different compartments. I found them in August, and obtained the fly on the 17th of September. C. hdrtipes 0. S. 9. — Antennas reddish-black, 22-jointed, joints short, subcylindrical, almost subglobular, gradually decreasing in size towards the tip, separated by pedicels which are shorter than the joints, yerticillate-pilose ; head dark reddish with black hairs on the vertex; eyes contiguous on the front; thorax blood-red, its back blackish, the usual three stripes being almost coalescent and separated by rows of erect black hairs ; collare blackish above ; pleura3 blood-red, with indistinct black dots ; scutellum and meta- thorax red, the first with black hairs; halteres reddish at base, the club deep black ; abdomen red, upper side of the segments with a blackish, apressed, rather sparse pubescence ; coxa? reddish, feet deep black; wings with a dense, blackish pubescence; costa black, especially along its middle portion ; second longitudinal vein reaches the margin at or close by the tip of the wing ; cross-vein indistinct. (Description drawn from a fresh specimen.) 11. C. carbonifera, n. sp. Pale, circular spots, surrounded by 196 DIPTERA OF NORTH AMERICA. a purplish-black ring, on the leaves of Solidago; under each spot, inside of the leaf, several larva?. I found them commonly in Au- gust, and observed that the hollow space within the leaf was fre- quently filled with a hard, black substance, not unlike charcoal. 12. G. racemicola, n. sp. Bud-shaped gall among the racemes of Solidago. It has about 0.1 in diameter, is green, and looks exactly like a bud, but is easily distinguished from the buds of Solidago by its stout, rounded form. Each gall contains a single reddish larva. Not rare in September. 13. 0. vaccinii, n. sp. Gall on the leaf of Vaccinium (or Gay- lussacia ?), in the shape of a cock's comb, I found near Wash- ington, in October, one single leaf with two galls of this kind, arising from the central rib. The largest of the galls was about 0.15 high and 0.2 broad about the middle. They were green, and resembled pretty much a cock's comb, or, still better, an oyster, fastened by its hinge. After having been kept for some time on moist sand, both burst open exactly like the valves of a shell, and a reddish larva escaped from each. Both wandered for some days in the bottle in which I kept them, and inclosed themselves after- wards in delicate semitransparent cocoons, formed above the sur- face of the sand, between some chips of paper which I had provided for them. Unfortunately, both died without undergoing their final transformation. 14. C. pini inopis, n. sp. Resinous cocoon on the leaves of the scrub pine (Pinus inops}. Similar cocoons have been observed on the European pine, and described a century ago by Degeer. Ratzeburg, in his Forst-Insecten, describes and figures the same cocoon, as well as the larva and the perfect insect, C. pini Deg.* Dufour (in the Ann. Soc. Entomol. de France, 1838, p. 293) gives an account of a Gecidomyia with precisely similar habits, which he observed on the South European pine (Pinus maritima), and which he called G. pini maritime. The larva producing these cocoons is remarkable for two rows of oblong, pointed, fleshy protuberances along its back, and a similar row on each side. (See Ratzeburg, Forst-Insecten, III, Tab. x, f. 14, L.) Early in April I saw some of these larvae emerge from a small hollow between two terminal buds, where they had probably spent the winter, and crawl along the leaves, * See the same figures of cocoon and larva in Wiegm. Archiv, etc., vol. VII, p. 233. CECIDOMYIA. 191 aided in this by a resinous substance which they exuded abund- antly. Having readied a certain height on the leaf, they stop and remain quiet till the resinous substance covering them becomes hard and assumes the shape of an oblong, whitish, semi-transparent cocoon. Then the larva may be seen moving to and fro inside of this cocoon. I did not succeed to rear the fly from the cocoons which I brought home, and when I returned to the same spot in the woods about a month later, the cocoons were already empty. According to Ratzeburg's statement the European species spends the winter in the cocoon. The American species, as just shown, forms its cocoon only in the spring. As, nevertheless, it may be identical, or at least closely allied to G. pini Degeer, I subjoin here the description of the latter, translated from Mr. Winnertz's mo- nograph. (Compare also Ratzeb. 1. c. Ill, p. 159.) C. (Diplosis) pini Degeer. £ Antennae somewhat longer than the body, brown, basal joints yellow, verticils snow-white ; joints strong, stout, on short pedicels, double joints three times as long as the pedicel, the last joint with a very small, nipple-shaped pro- jection ; hypostoma and front pale reddish or brown; palpi reddish- yellow; thorax brownish-black or black, with two rows of white hairs from the collare to the scutellum, and one row from the shoul- der to the origin of the wing ; pectus blackish ; pleurae reddish- brown ; halteres white ; abdomen reddish-brown, with white hair, forceps blackish ; feet brown with white articulations, under side silvery-white ; posterior feet with a silvery-white reflection when viewed in a certain light; wings milky white, with a white pubes- cence, the costal and the two first longitudinal veins brown, the third longitudinal vein pale ; transverse vein pale, but distinct, very oblique, situated a very short distance beyond the middle of the first longitudinal vein ; third longitudinal vein straight, turning towards the posterior margin in an obtuse, rounded angle ; the second longitudinal meets the costal immediately beyond the tip of the wing. 9 Antennae a little more than half as long as the body, brown with gray verticils, basal joints yellow; joints of the flagellum about five times as long as the pedicel, last joint ending in a small bud-shaped appendage ; hypostoma reddish-yellow ; front reddish- brown ; palpi and thorax as in <^; halteres brown ; abdomen red- dish-brown, with short whitish hairs, more dense and with a silvery reflection laterally ; ovipositor short, yellow, with two small oval 19S D1PTERA OF NORTH AMERICA. laraels ; femora and tibiae and the anterior pair of tarsi superiorly black or black-brown ; inferiorly white with a silvery reflection ; the posterior tarsi have the two basal joints black or black-brown superiorly, silvery white inferiorly; the three last joints are silvery white, sometimes with blackish articulations ; wings gray, irides- cent with a dense, blackish-brown pubescence and brown veins ; cross-vein distinct, very oblique, situated a little before the middle of the first longitudinal vein; second longitudinal as in £ , the third likewise, although almost perpendicular to the posterior mar- gin. Length g 9 0.1 to 0.15. 15. C. brachynteroides, n. sp. Swelling at the basis of the leaves of the scrub pine (Finns inops). In consequence of this swelling the pairy leaves diverge, their bases coalesce, and the sheath at the basis of the bunch bursts. In July these swellings contain several small reddish larvae; in winter I found them empty, but having observed some larva? hanging on cobwebs near these galls, I conclude that they undergo their transformation under ground, and were caught in these cobwebs in the attempt to leave the gall. The habits of this Cecidorayia seem to be very like those of C. brac/tyntera Schwagr. living at the base of the pairy leaves of the European Pinus sylvestris. Still, the latter produces no gall or swelling whatever, and causes the leaves only to wither ; it also goes under ground for transformation. (See Ratzeburg, Forst-In- secten, Vol. Ill, p. 160.) 16. G. serrulalce 0. S. Deformed terminal buds of the common alder (Alnus serrulata). The buds appear enlarged, rounded, pointed at the tip, having from three to five lines in diameter. In autumn they are greenish ; in winter withered, brown, and frequently covered with a whitish efflorescence. Each gall contained in October from two to six red- dish larva?, lodged in the same compartment. In winter the galls are found empty, as the larvae go under ground. By keeping some of these galls, gathered in October, on moist earth, I obtained the fly in the following April. It belongs to the sub-genus Cecidomyia Loew. The description has been drawn from fresh specimens. C. serrulatae 0. S. 219 175 176 Gen. XL. CHRYSOTIMUS . 220 178 1. pusio Lw. 9 221 178 2. delicatus Lw. 9 • 222 179 181 Gen. XLI. XANTHOCHLORDS 223 182 1. helvinus Lw. 9 • 224 183 183 Gen. XLII. SAUCROPUS . 224 184 1. dimidiatus Lw. £ 225 2. rubellus Lw. 9 . 226 185 3. superbiens Lw. £ . 9 » 227 4. tenuis, nov. sp. 9 228 185 187 Gen. XLIII. PSILOPUS 229 188 diffusus Wied. £ . 9 . 235 189 guttula Wied. $ . 9 237 191 1. dimidiatus Lw. £ 246 193 2. sipho Say, $ . 9 247 3. scaber Lw. £ 250 194 4. patibulatus Say, £ . 9 . 251 5. melampus Lw. $ . 9 . 253 196 6. pilosus Lw. $ 256 196 7. jucundus Zw. $ . 9 258 TABLE OP CONTENTS. PAGE 8. ciliatus Lw. £ . . 260 9. comatus Lw. £ . 9 • 262 10. chrysoprasius Wk. £ . 9 265 11. scobinator Lw. £ . 9 • 268 12. caudatulus Lw. £ .. 271 13. calcaratus Lw. £, . 272 14. inermis Lw. £ . . 272 PAGE 15. scintillans Lw. $ . 9 . 273 16. pallens Wied. $ . S? • 275 17. variegatus Zw. 9 . 278 18. tricolor Zw. 9 . . 280 19. psittacinus Lw. £ . 9 • 281 20. tener Lw. $ . 9 . .284 21. filipes Lw. 1 . .286 APPENDIX. Species described by previous authors, and not contained in this Monograph .......... 289 Fabricius, Systema Entomologies Musca longicornis . « Fabricius, Entomologia systema- tica .... Musca longicornis . Say, Jour. Acad. Nat. Sc., Phil. Dolichopus unifasciatus . obscurus femoratus Chrysotus nubilus . concinnarius . abdominalis Medeterus lateralis . punctipennis . Dolichopus abdominalis . Wiedemann, Aussereurop. Zwei- fliigel. Insecten . Psilopus macula Sayi . longicornis caudatus . virgo femoratus . mundus Dolichopus obscurus Say . Macquart, Suites & Buffon, Dip- teres .... Psilopus radians 289 Macquart, DiptSres Exotiques . 297 289 Psilopus sipho 297 radians 298 Chrysotus viridifemora 298 289 Dolichopus heteroneurus . 299 289 Walker, Insecta Saundersiana . 300 289 Psilopus lepidus £ . 300 289 Medeterus exustus 9 300 290 viridiflos 9 301 290 Dolichopus bifrons 9 301 291 censors 9 302 291 contingens 9 • 302 291 hebes 9 302 291 ineptus 9 303 292 maculipes 9 • 303 292 pulcher £ . 9 • 304 varius 9 304 Walker, List of Dipt. Brit. Mus. 304 292 Psilopus delicatus 9 304 292 gemmifer £ 305 293 chrysoprasi 305 294 suavium 306 294 amatus £ . 9 306 295 inficitus 307 295 nigrofemoratus . 307 296 albicoxa £ . 9 307 296 Chrysotus incertus 308 Porphyrops pilosicornis . 308 Medeterus glaber . 309 297 chrysologus 9 309 297 alboflorens 9 310 TABLE OF CONTENTS. PAGE Dolichopus affinis £ . 9 • . 310 Dolic cupriuus Wiecl. . 311 lamellipes $ . 9 . 311 ciliatus £ . 9 . 312 ' adjacens 9 • . 312 coercens 'J, . . 312 Orthc finitus £ . 313 distractus . 313 Wall discesus 9 • . 314 I contiguus £ . 314 exclusus 9 • . 31-t confiuis 9 . 315 conterminus £ . 315 separatus 9 . 316 PAGE Dolichopus terminatus 9 • 31 1! seqiiax . , . 317 soccatus . . :!17 rernotus . . 317 irrasus . . . 31S Orthochile derempta . . 318 Walker, Trans. Entorn. Soc. . 319 Psilopus ungulivena . 319 solidus . . 319 peractus . .319 haeretictis . . 320 peruiodicus . 320 SUPPLEMENT. I. General remarks on the Dolichopodidse of North America . II. Description of some Species communicated after the volume had gone through the press ........ Gen. II. DOLICHOPUS . . 323 43. dorycerus Lw. £ . 326 44. spleiididulus Lw. £ . 327 45. paljestricus Lw. £ . 9 . 328 . 9 . 46. melanocerus Lw. •£, . 9 47. quadrilamellatus Lw. Gen. III. GYMNOPTERNPS 19. politus Lw. £ . 9 24. pusillus Lw. 9 . 27. chalcochrus Lw. 330 331 332 334 334 335 26. coxalis Lw. $ 27. meniscus Lw. £ . 28. humilis Lw. £ . 9 29. exiguus Lw. 9 Gen. V. PELASTONEURUS . 6. lamellatus Lw. 9 7. abbreviates Lw. 8. alternans Lw. 9 • Gen. VI. PORPHYEOPS 5. longipes Lw. $ . 321 323 335 336 336 337 337 338 33S 339 340 340 EXPLANATION OF THE PLATES . ...... 341 REMARKS ON THE GENERIC CHARACTERS EXPLANATORY TO THE PLATES . 345 INDEX ............ 357 DIPTERA OP NORTH AMERICA. PART II. VI. ON THE NORTH AMERICAN DOLICHOPODIDAE. INTRODUCTION. THE Dolichopodidas are among those families of diptera which show but so few points of relationship to others that it is impos- sible to name any family of a particular affinity to them. A superficial glance, it is true, seems to point out the EpTiydrinidse and the genera of the Tachydromidse in the vicinity of Clinocera as the nearest to the Dolicliopodidse ; but a closer examination will show that, instead of a true relationship, there are merely some points of external analogy between them, points common to most insects (and not only diptera) preferring to live in the vicinity of water. The want of a true relationship becomes immediately apparent when any portion of their inner or outer organization is submitted to a comparative scrutiny. The neura- tion of the wings of the Dolichopodidse alone is so peculiar that it is sufficient in itself to distinguish this family not only from the Epliydrinidae and the above-mentioned group of Tachydro- midse, but also from all other families of the order. This neura- tion, together with the brilliant-green color of the greater part of the species, renders the Dolichopodidse, one of the most easily recognisable families of diptera. The peculiarities of the neuration, which is shaped in general 1 2 DIPTEEA OP NORTH AMERICA. [PART II. according .to the type of the Diptera acalyptera (compare Mono- graphs, etc., I, page xxiv, fig. 1), are the following in this family : The costal vein extends as far as the tip of the fourth longitudinal vein ; the auxiliary vein does not run towards the costa, but to- wards the first longitudinal vein, stopping quite far from its tip, and either coalescing entirely with it or disappearing without having joined it. The third longitudinal vein emerges from the • second not far from its root, and both veins form at this place a more or less knot-shaped swelling; the small cross-vein (often quite obsolete) is at, or close by, this swelling, so that the first basal cell is unusually short ; the discoidal cell, much narrowed towards its base, is coalescent with the anterior of the two small basal cells in consequence of the absence of the vein separating them. The posterior small basal cell (the so-called anal cell) is small and rounded at the end. The sixth longitudinal vein does not generally reach the border of the wing ; sometimes it is en- tirely wanting, and in this case the anal cell becomes indistinct. The alar appendage (alula) is so narrow that it may be considered as wanting. Although all Dolichopodidse agree in the above-described pe- culiarities of neuration, still -this same neuration affords very good and abundant characters for the distinction of the genera and the species ; the- length and direction of the first longitudinal vein vary considerably, the position of the posterior transverse vein also ; the most striking characters, however, are taken from the structure of the last portion of the fourth longitudinal vein, which is subject to considerable variation, being sometimes parallel to the third longitudinal vein, sometimes more or less convergent with it, sometimes straight, often curved or bent abruptly in the middle, or even broken. In the latter case, the angle formed by the strong flexure or fracture sometimes bears a rudiment of a branch-vein, which in some cases (as in most Psilopus) is large enough to make the vein appear forked. For the purpose of distinguishing the Dolichopodidse from the other families of diptera the characters taken from the neuration are sufficient, and it is superfluous to have recourse to others taken from the rest of the body. It is different, however, if we intend to establish the natural character of this family. I will confine myself here to characters taken from the external organi- zation, and as to the internal one, I will merely notice in passing INTRODUCTION. 3 that, in accordance with the external structure, it forms a striking contrast with the organization of all the other dipterous families. The head of the Dolichopodidse in general is more or less he- mispherical. Its sides are occupied by the large, ovate, hairy eyes (naked only in Medeterus), which usually reach so far down that in most genera there are no cheeks (genee) at all, and that on the under side of the head, between the eyes, there is just space enough for the large opening of the mouth, which, in some spe- cies, even encroaches upon a considerable portion of the occipital side of the head. The usually rather broad front bears three ocelli, protected by strong bristles, and several bristles in both corners of the vertex ; the remainder of its surface has no bristles. The frontal fissure and lunule* are indistinct. The antennae are three-jointed, of very different structure in different genera, nay even often in the sexes of the same species ; the second joint is sometimes rudimentary. The arista is two-jointed, seldom alto- gether bare, generally with a very short pubescence, more seldom fringed with longer hairs or even plumose ; its dorsal or apical position affords very constant distinctions between different gene- ra ; its length and the other characters belonging to it differ sometimes even in the two sexes of the same species. The face is usually vertical and smooth ; it has often on its inferior third an elevated transverse swelling, or at least, on each side, near the orbit of the eye, a nodnle-like elevation ; seldom does the face reach down to the inferior angle of the eye ; it is always without any bristles, although sometimes it is hairy ; its lower edge is rather sharp, and does not coalesce with the border of the mouth ; from under this edge project the single-jointed palpi, incumbent upon the proboscis (except in Aphrosylus, where they hang down on its sides) ; they are usually scale-shaped, more seldom lancet- shaped (for instance, in Orthochile) or round, pallet-shaped (in Diostracus) ; in the females they are usually considerably larger than in the males ; sometimes however (in Diostracus), the oppo- site is to be found. The proboscis is generally short and (with the exception of Aphrosylus and Orthochile) stout ; its opening is wide, more or less surrounded by the protuberant suctorial flaps, which, by clap- * For the explanation of these terms, compare Monographs, Vol. I, p. xii. —0. S. 4 DIPTERA OF NORTH AMERICA. [PART II. ping together, shut the opening ; the species of the genera with a large proboscis, especially their females, can altogether engulf small insects within their proboscis, as if they swallowed them, whereas they only suck them out and throw the remainder away. The horny parts of the mouth are short. The stout labrum has on its lateral edge, which is bent downwards, several deep incisions, thus forming strong teeth. The tongue is longer than the labrum, and lies under it without being encased in it. The upper side of the thorax is convex, with bristles inserted in rows, between which there are more or less numerous short hairs ; in some genera there is a flat, somewhat concave declivity before the scutellum. Prothorax and mesothorax are very coalescent ; no distinct collare is perceptible, and the humeral callosities are not sharply limited ; of the ordinary transverse suture only the begin- ning is indicated on both sides by a large, more or less triangular impression. Scutellum with two, four, or six strong bristles ; more- over, its surface is sometimes hairy. The elongated prothoracic stigma is closed. Immediately above the basis of 'the fore coxae there is a strong bristle, seldom several. The metathoracic stigma is unusually small. The epirnera of the metathorax have a very large development ; they embrace often the basis of the abdomen from the side to a considerable breadth, and extend as a narrow stripe along the basis of its upper side. The abdomen of the female has seven segments, the last two of which are generally very narrow and contracted under the pre- ceding ones, so that only five segments appear on the outside ; the seventh segment, in the female, bears on its end two small lamellae connected with it by an articulation and fringed with hairs ; above them there is, in many species, a fan-shaped row of short, stiff, thorn-like bristles ; in other species the number of these thorns is reduced to four or two ; sometimes they are altogether wanting ; the latter is especially the case with those species which prefer dry localities, or also those, the hind part of the thorax of which has a flat usually concave declivity ; this difference in the structure of the ovipositor has certainly some relation to the difference of the locality where the eggs are deposited ; it is possible that a complete fan of thorn-like bristles constitutes a peculiarity of the species the larva? of which live under ground, whereas it is wanting when the eggs are laid in decayed wood. The abdomen of the male can also be considered as consisting INTRODUCTION. 5 of seven segments, if the hypopygiurn be considered as a single segment ; of the sixth segment only the upper half is distinctly developed, but it is much narrower and shorter than the surface of the immediately preceding segments. When this sixth segment is concealed under the upper half of the fifth, or when it is strik- ingly different by its coloring from the preceding segments and resembles in this respect the seventh, then the male abdomen appears only five-jointed. The seventh segment consists of the proportionally large hypopygium, the structure of which, near its basis, is not quite symmetrical. It consists of a rather stalk-like basal part and of a club-shaped posterior part ; the stalk-like, part is often very much abbreviated, and then not easily perceptible ; the club-shaped part is movable towards it ; the junction of both parts lies always above, on the left hand side of the club-shaped portion. The hypopygium is more or less inflected under the abdomen, sometimes imbedded in an excavation of the venter proportionate to its size. At the tip of the inflected hypopygium, below, are inserted, by means of articulations, three pairs of appendages of manifold structure. The outer pair of these appendages is gene- rally larger than the two others, and must be considered as organs of a great irritability, as the roots of the hairs on their inside are connected with nerves. The basis of this pair is linked to the hypopygium by a free articulation. The two. other pairs, attached by a less free articulation, can be considered as organs for the purpose of seizing, clutching ; they lie within the former pair and are not seldom of a very complicated structure,. Sometimes one or the other of these pairs exceeds in length the outer pair. In- nermost between them, coinciding with the middle line of the clutching apparatus, a single appendage is inserted, which also appears to be linked at its basis. On the upper side of the hypo- pygium, which is the side turned towards the abdomen, there are two more appendages, usually in the shape of a gutter ; they have no link at their insertion. Above this gutter, or, more seldom, within it, is another appendage, in the shape of a spike, which in some cases is somewhat button-shaped at the tip ; it is pierced lengthwise by a channel, and incloses more or less completely the penis, which protrudes considerably during the act of copulation. This organization is most distinctly developed in the genus Doli- chopus, whereas it is more difficult to observe in the genera with 6 • DIPTERA OP NORTH AMERICA. [PART II. a small and imbedded hypopygium. In some genera the one 01 the other of the paired organs are so small as to escape notice. A further and more thorough investigation has as yet to show whe- ther in some genera, and this seems, for instance, to be the case with Nypophyllus, a still larger number of appendages does not occur, and whether in general the organization of all the genera can be reduced to the common type, described above. Of the first abdominal segment often only the upper half is perceptible in the male and the female ; this is especially the case in those genera in which the metathoracic epimera cover a con- siderable breadth of the basis of the abdomen-; in other genera the inferior half of the segment is much shortened, and in a few cases only equal in length to the upper half. In several genera the lateral margin of the second and of the following segments shows deep punctures, arranged in a longitudinal row. The fore coxae, which are somewhat distant from the middle ones and placed much higher than those, are also longer, and reach as far as about the middle of the latter. The feet are in genera^ slender, the hind pair generally longer and stouter than the others ; the femora are often rather strong. Besides short hairs, the feet bear usually a number of stiff bristles, especially the tibiae. The pulvilli are only of moderate size ; the empodium is linear and always distinct; the claws (ungues) are plain and small. We have already explained above what is necessary for the understanding of the very characteristic neuration of the wings. The surface of the latter is microscopically hairy upon its whole extent. The tegujse are ciliated with bristle-like hairs, and some smaller ones besides ; they are simple, as the ^nferior duplication is wanting or only apparent as a narrow membranous stripe ex- tending towards the corner of the scutellum. The sexes of the same species show, besides the difference in structure of the genital organs, other important differences in their organization ; the coloring of the same parts of the body is often different. Here, as in many other families, it is in the male sex that these peculiarities assume the character of variously modified ornaments, and it seems impossible to trace any relation between them and the sexual functions or the sustenance of the male. The only exception in this respect is afforded by the considerable dif- ference in the structure of the parts of the mouth, which is much more developed in the females, in accordance with their greater INTRODUCTION. 7 need of food, and consequently their more predaceous habits. The plastic differences distinguishing the male sex from the other may be defined as follows : The eyes of the male are generally some- what larger, the face and sometimes also the front narrower ; the contiguity of the eyes in the male is not frequent ; still it takes place in some Diaphorus above the antenna?, and in some Chry- sotus below them. The third joint of the antenna? of the male is usually somewhat longer, sometimes much longer than in the female; likewise the antennal arista of the male is. often much prolonged, sometimes extended into a club or button at the tip or enlarged in the shape of a laniel, whereas in the female the arista is much shorter and quite plain. The feet of the females are, almost without exception, plain ; those of the male often differ consider- ably from them, and have various handsome ornaments, principally on the fore and middle tibia?, and the fore and middle tarsi ; the femora as well as the hind tibia? and tarsi very seldom show any- thing but a plain structure. To these differences in the structure qf the feet must be added those derived from the hairs and bristles which they bear; in the female these are usually more sparse, shorter, and coarser ; in the male, closer, more delicate, and longer ; sometimes also they assume in this sex some peculiar modified structure. Even the ungues of the male are sometimes of a pecu- liar irregular shape ; the pulvilli are in some cases (as in Diapho- rus) larger in the male than in the female. The wings of the male often differ from those of the female in the outline and the neura- tion, those of the latter being in general more plain, and repro- ducing in their neuration the characters common to the genus ; whereas the wings of the male show in both respects more specific peculiarities. These consist usually in characteristic sinuses of the posterior margin and in a stronger sweep of flexure of the longitudinal veins ; sometimes the anterior margin also shows a peculiar curve in its outline, or a local thickening, or an elegant fringe of hairs, all of which do not exist in the female. The hairiness of the eyes, as well as the hairs and bristles on the other parts of the body, is frequently more dense, often con- siderably longer in the male than in the female. The very minute and dense tomentum with a silvery reflection, which adorns the abdomen and the thorax in most species of Argyra, also forms spots on the thorax of some other genera (as Pelastoneurus) and, in the species of most genera, is perceptible at least on the lateral 8 DIPTERA OP NORTH AMERICA. [PART II. margins of the abdomen, is likewise generally more extended and much denser in the males than in the females. The sexual differences in the coloring may be described as fol- lows : The eyes of the male are frequently of a different color than those of the female, particularly so, for instance, in the genus Diaphorus. The enlargement of the antennal arista in the male often has a different coloring, usually white. The color of the face in the male is generally of a purer, often a brighter shade. In many species, the halteres of both sexes are colored differently — for instance, in an entire group of species of Psilopus, they are black in the male and yellow in the female. Even the cilia of the tegulas have in a few instances (as in some species of Dolichopus) a different coloring in the two sexes. Differences in the coloring of the feet are not rare ; they are especially striking in the genus Psilopus, some species of which have altogether yellow feet in the female, and black femora in the male ; in others, the pale coloring of the feet is more extended in the female than in the male. The ornaments of the feet, peculiar to the males, also differ in their coloring from the corresponding parts of the feet of the other sex, being usually black, sometimes whitish, or with a handsome silvery reflection. Even without displaying any peculiarity of structure, the feet of the male have sometimes white or silvery spots, which are wanting in the female. The wings are in some cases pictured in the male and not in the female. Such are some species of Dolichopus, Tachytrechus and Systenus, the males of which have a black or white spot at the tip of the wing or in its proximity, whereas the female does not show any trace of such a spot. The same peculiarity occurs also in some other genera. These, often so conspicuous differences between the sexes of the same species, sometimes render the recognition of their specific identity somewhat difficult. In order to proceed in such cases with some degree of certainty, it is necessary to pay a particular attention to those characters which are usually common to both sexes. The most reliable characters of this kind are : the hairs on the antennae, especially on the first joint ; the shape of the second antennal joint ; the position of the arista ; the color of the cilia on the inferior orbit ; that of the cilia of the tegula?, notwith- standing some exceptional cases of its diversity in the two sexes. In most genera, to these characters may be added the coloring of the feet and of the halteres as well as the neuration of the wings. INTRODUCTION. 9 The first two of these characters will hardly ever mislead, if it is borne in mind that they have no value in the genus Psilopus, especially in the subdivision with black cilia on the tegulge. As to the latter character (neuration) it should be remembered that the peculiarities strikingly developed in the males as specific marks are but slightly indicated in the female, and can be perceived only by a very close observation. The habits of the Dolichopodidse are, as far as known, generally predaceous. Most of them hunt for smaller diptera or other insects with soft bodies and suck them out. They are usually found in damp places, covered with a rich vegetation ; many are principally found on the leaves of aquatic plants, on stones, partly overflown with water, on darns and near waterfalls ; some of them are able to run rapidly over the water even when it is rippled by wind (Hydropho'rus) ; others are fond of salt or brackish waters (Aphrosylus, Thinophilus and some Hydrophorus) ; the species of Hedeterus prefer dry situations and are found on stumps of trees, fences, etc., even in very dry and hot weather. Little is as yet known about their mode of transformation. Most species live as larva? under the ground ; some are found in the earth collected in hollow, rotten stumps ; others (as Systenus) in wood undergoing a process of dry decaying. After all that has been said above, the natural characters of the Dolichopodidse may be put down as follows : Generally metallic green, brisk and restless diptera of small or medium size, preda- tory on other insects and living principally in damp situations ; the male sex .are principally distinguished from the females by differences in the structure of the feet; the larva? living under ground or -in decaying wood. Head hemispherical, eyes large, usually not contiguous in both sexes, hairy, fenced in-, along the hind border with a row of bristles or hairs. Front, with bristles on the vertex only, and with three ocelli. Antenna? stretched out straight, with a two-jointed arista. Face, without mystacine bristles ; its inferior border not merging into the lateral border of the mouth. Oral opening occupying the whole underside of the head, and often also a considerable portion of its hind plane. Proboscis short and stout, concealed above by the single-jointed, usually scale-shaped palpi, with a wide opening which can be shut by the protruding suctorial flaps. Labrum short and stout, with coarse tooth-like excisions on the sides ; lingua much more slender 10 DIPTERA OF NORTH AMERICA. [PART II. and somewhat longer. Prothorax and metathorax very coalescent, with rows of bristles ; the usual transverse suture indicated only on the sides. Abdomen with seven segments, only five of which are visible on the outside in the female, whereas in the male the sixth segment is generally perceptible, and the seventh consists of the hypopygium, usually inflected under the abdomen and com- posed of two consecutive parts, bearing at the end eight paired and two single appendages. Wings microscopically hairy on their whole surface ; auxiliary vein not running towards the anterior margin ; anterior basal cell very short ; discoidal cell coalescent with the second basal cell ; posterior basal cell very small ; alula rudimentary ; egutlas distinct, simple, ciliated with long hairs. TABLE FOE THE DETERMINATION OP THE GENERA. 11 TABLE FOR THE DETERMINATION OP THE GENERA. 1 ( First antennal joint hairy above. 2 ( First antennal joint glabrous above. 16 2 f Hypopygium disengaged. 3 I Hypopygium more or less imbedded. 14 o ( First joint of the hind tarsi bristly. 4 I First joint of the hind tarsi not bristly. 5 Face descending as far as the inferior angle of the eye. Gen. I. HYGROCELEUTH0S. Face not descending as far as the inferior angle of the eye. Gen. II. DOLICHOPUS. , ( Palpi of the male unusually large. Gen. XII. DIOSTRACUS. I Palpi of the male small 6 (The last portion of the fourth longitudinal vein is parallel, or almost so, to the third longitudinal vein. Gen. III. GYMNOPTERNUS. I The last portion of the fourth longitudinal vein is distinctly convergent (_ towards the third longitudinal vein. 7 f The end of the fourth longitudinal vein is abruptly, or at least steeply deflected anteriorly. 8 8 The end of the fourth longitudinal vein is only gradually deflected I. anteriorly. 9 Arista with the usual pubescence ; the end of the fourth longitudinal vein, beyond the angular flexure, runs in a curve. Gen. IV. PARACLIUS. Arista short-plumose ; the end of the fourth longitudinal vein, beyond the rounded flexure, runs in a straight line. Gen. V. PELASTONEURUS. / The face reaches down to the inferior corner of the eye. 9 < Gen. VI. TACHTTRECHUS. ' The face does not reach down to the inferior corner of the eye. 10 -.^ ( Proboscis and palpi very much prolonged. Gen. VII. ORTHOCHILE. 1 Proboscis and palpi not prolonged. 11 n ( Scutellum hairy. Gen. VIII. STBISTROMA. I Scutellum not hairy 12 , 2 ( Hypopygium sessile. Gen. IX. HERCOSTOMDS. I Hypopygium pedunculated. 13 , o ( Second antennal joint of usual shape. Gen. X. HYPOPHYLLUS. \ Second antennal joint rudimentary. Gen. XI. HALTERICERUS. 12 DIPTERA OF NORTH AMERICA. [PART II. ,4 ( Abdomen of male laterally compressed. Gen. XIII. ANEPSIUS. I Abdomen of the male not compressed. 15 ( Second antennal joint of the usual transverse shape. ,, J Gen. XIV. AEGYEA. j Second antennal joint with a thumb-like projection over the inside of L the third. Gen. XV. SYNTOEMON. Third autennal joint in both sexes, or at least in the male, prolonged, pointed, and with an apical arista. 17 16 -{ Third antennal joint short even in the male, and if it should be somewhat prolonged, then neither pointed nor with an apical, but at the utmost with a subapical arista. 23 /•Second antennal joint with a thumb-like projection over the inner 17 < side of the third. Gen. XVI. SYNAETHBUS. v Second antennal joint without a thumb-like projection, transverse. 18 Posterior transverse vein distant from the margin of the wing ; palpi -, n incumbent. 19 Posterior transverse vein approximated to the margin of the wing ; palpi hanging down. Gen. XXII. APHEOSYLUS. 19 $ Hypopygium pedunculated, free. Gen. XVII. SYSTENUS. I Hypopygium sessile, more or less imbedded. 20 2Q ( The male abdomen has five segments. Gen. XXI. SMILIOTUS. ( The male abdomen has six segments. 21 f Third antenual joint prolonged also in the female. 21 j Gen. XVIII. RHAPHIUM. v Third antennal joint of the female not prolonged. 22 f Third antennal joint of the male very much prolonged (small, less 09 J hairy species). Gen. XIX. XIPHANDEIUM. i Third antennal joint of the male moderately prolonged (larger, more t hairy species). Gen. XX. POEPHYEOPS. oo f Fourth longitudinal vein forked. Gen. XLIII. PSILOPUS. ' Fourth longitudinal vein simple. 24 {Upper side of the thorax convex behind. 25 Upper side of the thorax behind with a somewhat concave depres- sion. 40 or ( Fifth longitudinal vein altogether wanting. Gen. XXXVIII. ACHALCUS. v. Fifth longitudinal vein distinct. 26 f Distance of the posterior transverse vein from the margin of the wing equal to its own length or longer. 27 j Distance of the posterior transverse vein to the margin of the wing shorter than its own length. . 38 f Posterior transverse vein unusually oblique. 27 \ Gen. XXXIV. PLAGIONEUBUS. I Posterior transverse vein but little oblique. 28 r Hypopygium distinctly bent under the venter. 29 28 -I Hypopygium not distinctly bent under the venter or entirely imbed- 1 ded. 30 TABLE FOR THE DETERMINATION OP THE GENERA. 13 f The face of both sexes very broad, not narrowed superiorly. Gen. XXIII. THINOPHIUTS. 29 j The face of both sexes rather narrow, somewhat narrowed superiorly. Gen. XXIV. PEODES. ! Outer appendages of the hypopygium long, filiform. Gen. XXV. NEMATOPROCTUS. Outer appendages of the hypopygium not long, nor filiform. 31 (Third joint of the male antennae conspicuously large. Gen. XXVI. LEUCOSTOLA. Third joint of the male antennae small. 32 oo ( Pulvilli of the male fore tarsi conspicuously enlarged. 33 I Pulvilli of the male fore tarsi not, or very slightly, enlarged. 34 oo f Pulvilli of the male fore tarsi not prolonged. Gen. XXVII. ECTARSUS. I Pulvilli of the male fore tarsi prolonged. Gen. XXVIII. DIAPHORUS. 04 ( Arista altogether or almost altogether apical. 35 1 Arista dorsal. 36 OK ( Wings of considerable size (larger species). Gen. XXIX. LYROXECRPS. I Wings of small size (smaller species). Gen. XXX. CHRYSOTUS. (Feet of the male with isolated, strong, spine-like bristles. Gen. XXXI. TEUCHOPHORUS. Feet of the sexes without isolated, strong, spine-like bristles. 37 „- ( Face not narrowed above. Gen. XXXII. SYMPYCNUS. ( Face considerably narrowed above. Gen. XXXIII. CAMPSICNEMUS. !A11 femora slender, abdominal segments with bristles before the hind margin. Gen. XXXV. LIANCALUS. Fore femora incrassated towards the basis. 39 OQ I Fore tibiae with long thorns. Gen. XXXVI. SCELLITS. I Fore tibiae with very short little thorns. Gen. XXXVII. HYDROPHOKUS. 4~ ( Arista apical, or at least subapical. 41 v Arista distinctly dorsal. 42 r Third and fourth longitudinal veins strongly convergent. 41 1 Gen. XXXIX. MEDETERUS. I Third and fourth longitudinal veins parallel. Gen. XL. CHRYSOTIMUS. >„ ( Male abdomen with six distinct segments. Gen. XLI. XANTHOCHLORUS. (. Male abdomen with five distinct segments. Geiit XLII. SAUCEOPDS. . 14 DIPTEKA OP NORTH AMERICA. [PART II. SYSTEMATIC ARRANGEMENT OF THE GENERA. I. First joint of the antennae with hairs on its upper side. A. Hypopygium disengaged. A. Palpi of the male small. 1. First joint of the hind tarsi bristly. Gen. I. Hygroceleuthus. Gen. II. Dolicliopus. 2. First joint of the hind tarsi not bristly. a. Third and fourth longitudinal veins parallel or sub- parallel. Gen. III. Gynmoptermis. 6. Third and fourth longitudinal veins convergent. Gen. IV. Paraclius. Gen. VIII. SyMstroma. Gen. V. Pelastoneurus. Gen. IX. Hercostomus. Gen. VI. Tachytreclms. Gen. X. Hypophyllus. Gen. VII. Ortliocliile. Gen. XL Haltericerus. B. Palpi of the male conspicuously large. Gen. XII. Diostracus. B. Hypopygium more or less imbedded. Gen. XIII. Aiiepsius. Gen. XV. Syntormon. Gen. XIV. 4rgyra. II. First joint of the antennae entirely bare on its upper side. A. Third antennal joint in both sexes, or at least in the male, elon- gated, tapering in a point, with an apical arista. A. Posterior transverse vein distant from the margin of the wing, palpi incumbent. Gen. XVI. Synarthrus. Gen. XIX. Xiphandrium. Gen. XVII. Systemis. Gen. XX. Porpliyrops. Gen. XVIII. Rhaphium. Gen. XXI. Smiliotus. B. Posterior transverse vein approximated to the margin of the wing ; palpi hanging down. Gen. XXII. Aphrosylus. SYSTEMATIC AKRANGEMENT OP THE GENERA. 15 B. Third antennal joint short even in the male, or if somewhat elon- gated, then neither pointed at tip nor with an apical, but, at the utmost, with a subapical arista. A. Fourth longitudinal vein simple. 1. Upper side of the thorax convex posteriorly. a. Fifth longitudinal vein distinct. a. -Distance of the posterior transverse vein from the hind margin of the wing equal to its own length or longer. Gen. XXIII. Thinopliilus. Gen. XXIX. Lyronetirus. Gen. XXIV. Peodes. Gen. XXX. Clarysotus. Gen. XXV. Nematoproctus. Gen. XXXI. Teuchopltorus. Gen. XXVI. Leucostola. Gen. XXXII. Sympycuus. Gen. XXVII. Eutarsus. Gen. XXXIII. Campsicneinus. Gen. XXVIII. Diapfiioriis. Gen. XXXIV. Plagioneurus. 0. Distance of the posterior transverse vein from the margin of wing shorter than its own length. Gen. XXXV. Liancalus. Gen. XXXVII. Hydrophortis. Gen. XXXVI. Scellus. 6. Fifth longitudinal vein altogether wanting. Gen. XXXVIII. Aclialcus. 2. Upper side of the thorax posteriorly with a flat, somewhat concave, declivity. Gen. XXXIX. IHedeterus. Gen. XLI. Xantkochlorus. Gen. XL. Clirysotimus. Gen. XLII. Saucropus. B. Fourth longitudinal vein forked. Gen. XLIII. Psilopus. HYGROCELEUTHUS. IT Gen. I. HYGROCELEITTHUS. The genera Hygroceleuthus and Dolichopus are the only ones which have spine-like bristles on the first joint of the hind tarsi, and this character helps to distinguish them most easily from all other Dolichopodidse. They have also the following characters in common : first antennal joint beset with hairs above ; third joint not prolonged ; arista dorsal ; hypopygium entirely disengaged ; its outer appendages proportionally large or moderately large, lamelliform. The difference between the two genera is to be found in the length of the face, which, in Hygroceleuthus, reaches down to the inferior corner of the eye, and which is shorter in Dolichopus. In the typical species of Hygroceleuthus the first, and sometimes also the second joints of the antennae are consider- ably longer than in Dolichopus. As on one side the European Hygroc. Diadema Hal. approaches Dolichopus by the structure of its antennae, so, on the other side, some species of Dolichopus, for instance, D. plumipes Scop., a species common to Europe and North America, are related to Hygroceleuthus by the somewhat greater length of their face. I know as yet only four species of Hygroceleuthus; of the three typical species, one inhabits northern and middle Europe; the second Siberia ; the third North America. The fourth species of the genus is spread over all Europe. The name Hygroceleuthus (t>y£oxaji£i£oj, living in the wet) has been given to the genus on account of the species being found in moist localities. 1. H. latipes LOEW. % and 9- — Aeneo-viridis, antennis rufis, ciliis oculorum inferioribus pallidis, tegularum ciliis nigris, tarsis intermediis maris compressis. Brassy green ; antennae red ; cilia of the inferior orbit pale ; cilia of the tegulse black ; middle tarsi of the male compressed. Long. corp. 0.26. Long. al. 0.23. SYN. Hygroceleuthus latipes LOEW, Neue Beitr. VIII, 5. 2 18 DIPTERA OF NORTH AMERICA. [PART II. Green, shining. Antennse reddish-yellow, with a black supe- rior edge and with the tip of the third joint black ; first joint nar- row and elongated. Face white, yellowish above, much broader in the 9 than in the J\ Cilia of the inferior orbit pale. Fore coxa3, tip of the middle and hind COXSB and feet yellow ; tarsi black from the tip of the first joint ; the first joint of the fore tarsi some- times altogether dusky ; the root of the second joint of the hind tarsi, on the contrary, is pale. On the upper side of the middle tibias a bristle is conspicuous by its greater length ; there are two bristles before the tip of the hind tibias. Tegulse with black cilia. Wings tinged with brownish ; fourth longitudinal vein not broken ; hind transverse vein straight and steep. Four last joints of the middle tarsi compressed in the <£ , beset on the upper side with incumbent black hairs ; the fifth joint much narrower than the pre- ceding ones. Costa with a stout swelling near the tip of the first longitudinal vein. Lamellae of the hypopygium of moderate size, white, bordered with black at the tip, jagged and fringed with 'black bristles. Hob. North Red River. (Kennicott.) Gen. II. DOLICHOPUS. The principal characters of the genus Dolichopus, as it follows already from what has been said about it in the genus Hygro- celeuthus, are : the presence of hairs on the upper side of the first antenna! joint, the shape of the third joint, which is hardly ever very much elongated, the. dorsal position of the arista, the en- tirely disengaged hypopygium, the lamelliform shape of its rather large outer appendages and the presence of spine-like bristles on the first joint of the hind tarsi. The genus Dolichopus, established by Latreille already in 1796, is the oldest of the family. It comprised at that time all the Dolichopodidse, so that all the other genera have been gradually formed by the separation of some groups and by further subdivision of the latter. In the sense in which this genus was adopted by Wiedemann and Meigen, it still included the present genera Gym- nopternus, ParacUus, Pelastoneurus and Tachytrechus, besides some isolated species belonging to other genera, which had been erroneously located in it. (Such was the case, for instance, with D. adustus Wied., which belongs to Lyroneurus.} The defini- DOLICHOPUS. 19 tion of Dolichopus, in its present limited sense, was given by me in 1857. Still, even now, this is one of the largest genera of this family. Although possessing abundant and striking specific dif- ferences, its numerous species show at the same time so much agreement in their general organization, that a subdivision into smaller genera is impossible at present, and in future an attempt of that kind will require a great deal of caution. One is easily tempted here to establish generic groups founded upon characters of a purely specific value. The genus Rhagoneura, formed by Rondani, is due to a mistake of this kind. To found a new genus on D. ziczac, which species apparently requires it on account of some differences in the neuration and in the structure of the an- tennae, seems hardly worth while, as this is as yet the only species showing such differences. The geographical distribution of the genus Dolichopus cannot be inferred from the data found in older authors, as this genus con- tained at that time very heterogeneous elements. Neither can Mr. > Walker's superficial publications on exotic species be considered in this respect as a source of information, as it is impossible to tell from his descriptions which of his species belong to Dolichopus in the restricted sense. Those species from the southern hemis- phere, and fi'om the southern parts of the northern, which I had occasion to examine at different times, proved not to belong to Dolichopus in that sense. Hence we may safely conclude that the genus Dolichopus, in the large majority of its species, belongs to the cold and temperate zones of the northern hemisphere ; it can, at least, be positively asserted that the number of species, in Europe as well as in America, goes on diminishing toward the South. The name of the genus (SOM^OJ, long, and rtov;, foot) has refe- rence to the length of the feet of its species. Thirty-one North American species of Dolichopus have been described by former authors ; some of them, however, do not belong to this genus in the restricted sense adopted here. Of these species three have been described by Say, one by Zetterstedt, one by Macquart, and twenty-six by Walker. The descriptions given by Mr. Walker are, for the most part, very bad, and the worst are those published in the Diptera Saundersiana, as they con- tain only such characters as are common to all the species, or at least to entire groups, without paying the least attention to those 20 DIPTERA OF NORTH AMERICA. [PART II. marks which serve to distinguish one species from another. The eighteen descriptions contained in the List of Diptera of the British Museum mention at least occasionally such specific cha- racters, and may therefore C9ntain some data for identification; but their great defect is that Mr. Walker has not given a complete description of those peculiar marks which distinguish the males of many species, so that, from his silence about these marks, one can- not with safety conclude that they are Teally wanting. This accu- racy is absolutely necessary in order to make a description avail- able. Among the forty-one North American Dolichopodidse now known to me, I recognize with certainty only a single species already described before (besides the three species which North America has in common with Europe, D. plumipes Scop., D. brevipennis Meig., and D. discifer Stann.); it is theZ). cuprinus Wied. (= cupreus Say). This striking result induces me to give a separate account of all the other species published, by former authors, in the order of their publication. 1. obscurus Say. This species, also described by Wiedemann, evi- dently belongs to the genus Gymnopternus and will be discussed there. Wiedemann's collection affords no light upon this species. It contains, it is true, two specimens of a Dolichopus marked obscurus, but this is no other than a species of Tachytrechus, from the Cape, described by Wiedemann himself under the name of obsccenus. It is probable that Wiedemann named this species, as he was in the habit of doing, as soon as he received it in his collection, and that when he described it afterwards, he changed its name to obscoenus, on account of the already existing obscurus Say, neglecting, at the same time, to change the etiquette in his collection. 2. abdominalis Say. The abdomen is said to be reddish. If Say means a reddish, non-metallic color, then it is a distinct species, entirely unknown to me, which will be easily recognizable even if it is no true Dolichopus, as may very probably be the case. If, how- ever, he means a metallic, coppery-red coloring, then the descrip- tion is too unmeaning to pronounce even about its belonging to Dolichopus in the restricted sense ; as to the identification of the species, it is altogether out of question. 3. groenlandicus Zett. A true Dolichopus with black as the prevailing color of its feet. It is not among the number of the species known to me. 4. heteroneurus Macq. is either a Pelastoneurus or a Paraclius. 5. bifrons Walk. Dipt. Saund. It may be inferred, from the peculiar coloring of the face of this species, that it is a Pelastoneurus, DOLTCHOPTTS. 21 although no mention is made of the peculiar course of the fourth longitudinal vein, which characterizes this genus. 6-9. censors, contiguus, hebes, ineptus, all four described by Walker in the Dipt. Saund. from female specimens. The descrip- tions are so wretched that the identification is impossible. 10. maculipes Walk. Dipt. Saund. The spots on the tibise, mentioned by Walker, seem to indicate that this is a Pelastoneurus. But the peculiar course of the fourth longitudinal vein, the chief character of the genus, is again not mentioned. 11. pulcher Walk. Dipt. Saund. I take this to be a true Dolichopus, although the datum of the fourth longitudinal vein beyond its flexure not converging, but being parallel to the third, seems t» in- dicate a Gymnopternus. It belongs to the species with black femora, but among the species of this description which I possess from North America, there is none to which Mr. Walker's description of the fourth longitudinal vein is applicable. 12. varius Walk. Dipt. Saund. A very distinct species on account of its spotted wings and which will be easily recognizable even if, as it seems probable, it does not belong to the genus Dolichopus. 13. affinis Walk. This and the following species have been described by Mr. Walker in the List of Diptera, etc. All belong very probably to the genus Dolichopus, and to the subdivision with yellow feet, except the cases which I have expressly mentioned below. It is not said whether the tegulse of D. affinis are ciliated with black or pale hairs. If the latter is the case, and if, as it appears from Mr. Walker's data, the hind tarsi are altogether black, my D. splendidus might alone be taken in consideration ; but it is hardly possible that it should be this species, as Mr. Walker's description of the color- ing does not apply to it, and as the hind femora of the £ of D. affinis Walk, are ciliated with only a few hairs, whereas in D. splendidus these hairs are very numerous. If D. affinis has black cilia on the tegulse, then D. discifer and lobatus could be thought of, but the hind femora of their males are entirely destitute of bristles, so that the description of D. affinis cannot be applied to either of .them. 14. lamellipes Walk. It has the inferior orbit ciliated with black. Among the North American species of Dolichopus with pale-colored feet D. pachycnemus and D. brevipennis alone partake of this cha- racter. The other data of Mr. Walker do not apply at all to these species, as neither of them has a white face, the first antennal joint red and the third joint very long. 15. ciliatus Walk. Very poorly characterized. I suppose that the cilia of the inferior orbit are pale. If the tegulse are likewise ciliated with pale hairs, then D. variabilis and D. luteipennis might be taken in consideration. But D. variabilis has no golden-yellow face, its fore tarsi are not "dark tawny," but always black from the tip of 22 DIPTERA OF NORTH AMERICA. [PART II. the first joint ; besides, its male has ciliated hind femora, which Mr. Walker does not ascribe to his species. D. luteipennis has a white face and the last joint of its fore tarsi black ; its wings are yellowish and not gray, the veins are luteous and not black ; finally - the hind femora of its male are al^o ciliated. If the tegulse of D. ciliatus are ciliated with black, then we might perhaps identify it with D. vittatus ; but the size of the latter is too large to admit of this identification, and its face, instead of being golden-yellow, is whitish ; its fore tarsi are not dark tawny but always black from the tip of the third joint. 16. adjacens Walk. Very poorly described from a female specimen, so that it would be entirely useless to attempt its identification. 17. coercens Walk. Mr. "Walker says that the coxae are blackish-green towards the basis. This character, combined with the others which are given, excludes at once all the species with the tegulse fringed with black, which are known to me. If the tegulse are fringed with yellow, this species might be taken for D. longimanus; but the last joint of the fore tarsi of the latter is only moderately enlarged, assuming the shape of a very small lamella, and its hind tarsi are colored black in a rather striking manner from the very root of the first joint ; whereas Mr. Walker says that in his species they are pitch brown towards the end. Under such circumstances the des- cription of D. coercens Walk, cannot possibly be referred to D. longimanus. 18. finitus Walk. If the tegulse are fringed with pale, then, among the species known to me, D. longimanus, D. splendidus and D. batillifer have to be taken in consideration. The face of D. longimanus is not whitish, but pale ochre-yellowish in the £ and yellowish gray in the 9 • the cilia of the inferior orbit are not white, but yellowish, and the hind tarsi are not pale at the basis, as in D. finitus, but en- tirely black. D. splendidus is distinguished from D. finitus by its hind tarsi, which are not pale at the base, and by the ciliated hind femora of the male. The face of D. batillifer is not white ; the hind femora of the male are ciliated and the hind tibise incrassated in a very striking manner; therefore this species is also distinct from D. finitus. If the cilia of the tegulse of D. finitus are black, then we might compare D. discifer and lobatus with it. The hind tarsi of D. discifer are not pale at the base, and the first joint of the an- tennse is tinged with black only on its upper edge, so that it is hardly probable that Mr. Walker should have overlooked its striking red coloring. This also excludes this species from the identification with D. finitus. The hind tarsi of D. lobatus likewise are not pale at the base, and the first joint of the antennse is red, and only somewhat dusky on its upper edge, so that it has also to be considered as different from D. finitus. DOLICHOPUS. 23 19. distractus Walk. Walker says nothing about the sex of the de- scribed specimen ; it seems to have been a female. The descrip- tion does not afford any data for even an approximative identifica- tion. 20. discessus Walk. Mr. Walker gives a very unsatisfactory descrip- tion of a female. It is a rather surprising datum that the upper side of the thorax has two longitudinal coppery-red stripes, whereas all the species known to me, and marked with stripes of this color, have always three, that is, a narrow intermediate one, and two broad lateral ones. This character might perhaps serve as a clue for identification. 21. contiguus Walk. Nothing is said about the color of the cilia of the inferior orbit. If they are black, then it is certain that D. contiguus is not among the species from North America known to me. If they are pale, it still remains to be known of what color are the 'cilia of the tegulze about which Mr. Walker is also silent. Sup- posing that they are pale, then D. splendidus would have to be taken in consideration ; but its male has fringed hind femora and cannot therefore be identified with D. contiguus. If the cilia of the tegulae are black then we would have to compare D. discifer and lobatus. But the last joint of the fore tarsi of D. discifer is not at all much enlarged, and forms only a small pallet ; therefore it can- not be taken for D. contiguus. D. lobatus has a very much enlarged last joint of the male fore tarsi, but as the first joint of the antennae is red and only slightly dusky on the upper edge, and as the lamellae of the hypopygium have a broad black margin, whereas Walker describes the lamellae of D. contiguus simply as whitish, as the wings of D. lobatus, in the male sex, are distinguished by their peculiar shape, which is not mentioned in the description of D. con- tiguus, on account of all this we cannot consider these species as being identical. 22. exclusTis Walk. Very poorly described from a female, so that all effort to identify the species must remain fruitless. 23. confinis Walk. The same may be said of this species. 24. conterminus Walk. Mr. Walker does not mention the color of the cilia on the tegulae, and this renders the identification impossible. If they are black, then the species is not among those known to me. D. discifer and D. lobatus are here again the species with which Mr. Walker's description agrees in most particulars, but both are easily distinguished by the absence of fringe on the hind femora of the male. If, on the contrary, the cilia of the tegulae of D. conterminus are pale, then D. splendidus might possibly be identified with it, although it must be admitted that the coloring of this species as well as some other characters, somewhat disagree with Mr. Walker's description. 24 DIPTERA OF NORTH AMERICA. [PART II. 25. separatus Walk. Bad description of a female, rendering identifica- tion impossible. 26. terminatus Walk. Also a female described. The only species to which it may possibly be referred is D. chrysostomus, and I would have no doubt about it if I knew that the cilia of the tegulse of D. terminatus are black. But Mr. Walker's description is silent about this ; neither does it mention the very striking black incisures which the abdomen of D. chrysostomus shows ; finally, the third joint of the antennae of the latter species is pointed at the tip. Con- sidering all this, it would be premature to assume the. identity of these species. Those who. in the determination of the species, rely upon possibilities and vague reasonings, would perhaps do so. And doubtless it is that the adoption of this synonymy would be more reliable than that of any other of Walker's species with one of those described by me. 27. sequax Walk. Mr. Walker says that this species has small tufts' of black hairs at the basis of the middle tibiae. He does not mention, however, whether this peculiarity is to be found in the male alone, or in both sexes. At all events this is a very unusual distinction among the species of Dolichopidae, which will render the identifica- tion easy. This species is not among those described by me. 28. soccatus Walk. The sex of the described specimen is not men- tioned ; it seems to have been a female. The description is too in- complete to allow recognition. 29. remotus Walk. Description of a male with plain tarsi. The cha- racters given allow a comparison merely with D. incisuralis, the male of which, however, has fringed hind femora and cannot there- fore be identical with D. remotus. 30. D. irrasus Walk. This is a small species, distinguished by its dark blue color and unusually short abdomen, which is certainly not to be found among the species known to me. It seems probable that it is no true Dohchopus at all. The slender result of the foregoing discussion of Mr. Walker's species is, that there are only two among them which, with cer- tain problematical admissions, may perhaps be identified with species described by me, namely, D. conterminus Walk, with my D. splendidus, and D. terminatus with D. chrysostomus. To facilitate the determination of the species I give, first, a dichotomic table. I found it impossible to bring it down to the single species without making use of characters peculiar to the DOLICHOFUS. 25 male sex only. The systematic arrangement which follows the dichotomic table subdivides the genus in groups, founded upon easily perceptible characters common to both sexes. I think it will be useful to retain these groups, or, at least, the principal among them. Table for determining the Species. , f Prevailing color of the feet black. 2 I Prevailing color of the feet yellowish. 9 9 ( Cilia of the inferior orbit black. 3 v. Cilia of the inferior orbit whitish. 4 o f Face ochre yellowish. 1 gratus Lw. ( Face silvery white. 2 laticorais Lw. . f First joint of hind tarsi with numerous bristles. 3 setifer Liu. \ First joint of hind tarsi with but few bristles. 5 _ ( Hind tibiae black only at the tip. 6 \ Hind tibiae quite black. 7 f The black at the tip of the hind tibiae rather extended and not very • sharply limited. 4 albiciliatus Lw. i The black at the tip of the hind tibias but little extended and sharply I limited. 5 xanthocnemus, n. sp. H ( A considerable extent of the tip of the femora yellow. 6 tetricus, n. sp. I The extreme tip of the femora only somewhat yellow. 8 R f Lamellae of the hypopygium pointed. 7 acuminatus Lw. \ Lamellae of the hypopygium rounded ovate. 8 ovatus Lw. q ( Cilia of the inferior orbit black. 10 I Cilia of the inferior orbit pale. 11 ^0 ( Fore coxae blackish. 9 pachycnemus Lw. I Fore coxae yellow. 10 brevipennis Meig. , , f Tegulaa with pale cilia. 12 I Tegulse with black cilia. 23 ,„ f Antennae black, at the utmost, the first joint almost red. 13 I Antennas altogether, or at least their larger portion, yellowish red. 21 .„/ Fore coxae dark beyond the middle. 11 longimauus Lw. I Fore coxse pale. 14 , . f Tip of the hind tibiae decidedly blackish. 15 I Tip of the hind tibias not or very slightly infuscated. 16 1 , ( Fore tarsi only ferruginous-brownish. 12 brevimanus Lw. I Fore tarsi blackened from the tip of the first joint. 13 socius Lw. ,„ ( Hind tarsi entirely black. 17 X Basis of the hind tarsi pale to a considerable extent. 19 , - f Hind femora of the male not ciliated. 14 uudus, n. sp. \ Hind femora of the male ciliated. 18 26 DIPTERA OF NORTH AMERICA. [PART II. ' Hind femora of the male with sparse and rather short cilia. 15 subciliatus, n. sp. Hind femora of the male with very long and dense cilia. 16 splendidus Lw. , Only the last joint of the fore tarsi of the male is enlarged. 19 J 17 batillifer Lw. ( The two last joints of the fore tarsi of the male are enlarged. 20 0(, f Hind femora of the male ciliated. 18 eudactylus Lw. I Hind femora of the male not ciliated. 19 tonsus Lw. o-i f Last joint of the fore tarsi of the male enlarged. 20 tener Lw. I Fore tarsi of the male plain. 22 22 / Wings hyaline with a grayish tinge. 21 variabilis Lw. (• Wings hyaline with a yellowish tinge. 22 luteipennis Lw. go f Fourth longitudinal vein broken. 24 X Fourth longitudinal vein not broken. 28 24 f Antennae black. 23 ramifer Lw. I Antennas yellowish-red. 25 f Fourth longitudinal vein broken twice at right angles. 24 bifractusiw?. 25 | The inferior angle of the fourth longitudinal vein sharp, the superior v one rounded. 26 2/. f Tarsi of the male plain. 25 vittatus Lw. I Tarsi of the male enlarged at the tip. 27 27 f Hind femora of the male ciliated. 26 cuprinus Wied. I Hind femora of the male not ciliated. 27 longipennis Lw. /-Antennae red, at the utmost the third joint somewhat or altogether 28 ] blackened at the tip. 29 I. Antennae black, at the utmost the first joint partly red. 36 ! Humeral callosity of the same color with the upper side of the thorax. 30 Humeral callosity yellowish. 35 OQ f Arista much enlarged towards the tip in the male. 28 hastatus, n. sp. v. Arista of the male not enlarged. 31 o-j f Last joint of the fore tarsi not enlarged in the male. 32 I Last joint of the fore tarsi enlarged in the male. 33 First joint of the male middle tarsi feathered. 29 plumipes Scop. First joint of the male middle tarsi not feathered. 30 fulvipes Lw. {Last joint of the male fore tarsi with a lamelliforni appendage. 31 sexarticulatus, n. sp. Last joint of the male fore tarsi without lamelliform appendage. 34 04 f Last joint of the male fore tarsi small. 32 ruficornis Lw. I Last joint of the male fore tarsi large. (40 lobatus Lw.~) „, f Fore tarsi of the male plain. 33 scapularis Lw. I Fore tarsi of the male enlarged at the tip. 34 funditor Lw. og / Face dark golden-yellow. 35 chrysostomus Lw. \ Face not golden-yellow. 37 SYSTEMATIC DISTRIBUTION OP THE SPECIES. 27. „- f Tip of the hind femora infuscated above. 38 I Tip of the hind femora not infuscated above. 39 OQ 5 Fore femora on the under side with a black stripe. 36 praeustus Lw. (. Fore femora without black streak. 37 comatus Lw. OQ f Fore tarsi black only at the tip. 40 I Fore tarsi black from the tip of the first joint. 41 "Two last joints of the male fore tarsi but little enlarged, feathered with black. 38 scoparius, n. sp. Two last joints of the male fore tarsi enlarged, not feathered. 39 discifer Stann. £, / Last joint of the male fore tarsi very much enlarged. 40 lobatus Lw. I Fore tarsi of the male plain. 42 42 / Bristles of the hind tibiae longer than usual. 41 setosus Lw. *- Bristles of the hind tibiae not longer than usual. 42 incisuralis Lw. Systematic distribution of the Species. I. Prevailing color of the feet black. A. Cilia of the inferior orbit black. I. gratus Lw. 2. laticornis Lw. B. Cilia of the inferior orbit whitish. 3. setifer Lw. 6. tetricus, n. sp. 4. albiciliatus Lw. f 7. acuminatus Lw. 5. xanthocnemus, n. sp. 8. ovatus Lw. II. Prevailing color of the feet yellowish. A. Cilia of the inferior orbit black. 9. pachycnemus Lw. 10. brevipennis Heig. B. Cilia of the inferior orbit pale. A. Cilia of the tegulae pale. 1. Antennae black, the first joint at the utmost partly red. II. longimanus Lw. 16. splendidus Lw. . 12. brevimanus Lw. 17. batillifer Lw. 13. socius Lw. 18. eudactylus Lw. 14. nudus, n. sp. 19. tonsus Lw. 15. subciliatus, n. sp. 2. Antennas entirely, or their greater portion, yellowish red. 20. tener Lw. 22. luteipennis Lw. 21. variabilis Lw. 28 DIPTERA OP NORTH AMERICA. [PART II. B. Cilia of the tegulse black. 1. Fourth longitudinal vein broken. a. Antennae black. 23. ramifer Lw. b. Antennae yellowish-red. 24. bifractus Lw. 26. cuprinus Lw. 25. vittatus Lw. 27. longipennis Lw. 2. Fourth longitudinal vein not broken. a. Antennae red, at the utmost the third joint at the tip almost entirely red. a. Humeral callosity of the same color with the thorax. 28. hastatus, n. sp. 31. sexarticulatus, n. sp. 29. plumipes Scop. 32. ruficornis Lw. 30. fulvipes Lw. B. Humeral callosity yellowish. 33. scapularis Lw. 34. funditor Lw. b. Antennae black, at the utmost theifirst joint partly red. 35. chrysostomus Lw. 39. discifer Stann.' 36. preeustus Lw. 40. lobatus Lw. 37. comatus Lw. 41. setosus Lw. 38. scoparius Lw. 42. inclsuralis Lw. DESCRIPTION OF THE SPECIES. 29 DESCRIPTION OF THE SPECIES. I. PREVAILING COLOR OP THE FEET BLACK. A. Cilia of the inferior orbit black. 1. I>. grains LOEW. $ . — Ex viridi chalybeus, pedum nigrorum tibiis anterioribus totis tibiarumque posticarum dimidio basali flavis, facie ochracea, ciliis oculorum inferioribus, tegularumque ciliis nigriis. Bluish-green, feet black, the four anterior tibiae entirely, the two hind ones upon the basal half yellow ; face ochre-brownish ; cilia of the inferior orbit and of the tegulae black. Long. corp. 0.23—0.24. Long. al. 0.21. SYN. Dolichopus gratus LOEW, Neue Beitrage, VIII, 11, 1. Bluish-green. The narrow face ochre-brownish. Antennae black ; the first joint rather narrow. Front metallic bluish-green. The cilia of the inferior orbit black. Lamella? of the hypopygium whitish, of moderate size, on the upper and the apical margin with a moderately broad black border ; the apical margin somewhat jagged and fringed with numerous black bristles. Femora black, with yellow tip ; the hind femora rather stout, provided with a bristle before the tip ; upon the under side ciliated with long black hair ; the four anterior tibia? and tarsi yellowish, the latter ones but little darker towards the tip ; hind tibia? somewhat thickened, the apical half and the whole of their hind edge black, the remain- ing parts yellowish ; hind tarsi quite black, the first joint with few bristles. Tegulse with strong black cilia. Wings hyaline, the costa is thickened before the tip of the first longitudinal vein, and only very gradually attenuated beyond ; the end of the fourth lon- gitudinal vein converges towards the third. Hob. Trenton Falls, West Point, Palisades, etc., N". Y. (Os- ten-Sacken.) 2. D. laticornis LOEW. %, . — Viridis pedum nigrorum tibiis, excepto posticarum apice, flavis, facie alba, ciliis oculorum inferioribus nigris, tegularum ciliis albis. Green, feet black ; tibiae, excepting the tip of the hind ones, yellow ; face 30 DIPTERA OP NORTH AMERICA. [PART II. white ; cilia of the inferior orbit black ; cilia of the tegulae white. Long, corp. 0.17. Long. al. 0.15. STN. Dolichopus laticornis Loew, Neue Beitr. VIII, 12, 2. Face white, rather broad for a male. Antennas black; the third joint large and broad, ovate ; arista inserted upon its second third. Front metallic green. The cilia ^of the inferior orbit black. Lamella? of the hypopygium rather small, of a trapezoidal form, the upper and apical margin are but very little bordered with black ; their margin is not jagged but only fringed with minute black hairs. Tip of the coxa? yellow. Femora black, with a faint greenish reflection and yellow tip. Tibiae yellowish. (The middle tibiae are wanting in the described specimen.) The tip of the hind tibise, which are not thickened at all, is black. Fore tarsi from the tip of the first joint black ; hind tarsi entirely black; the first joint is beset with a few thorn-like bristles. Hind femora before the tip with a stout "bristle. Tegulse with whitish cilia. Wings hyaline ; the costa but scarcely thickened about the tip of the first longitudinal vein ; the end of the fourth longitu- dinal vein converges towards the third. Hob. Connecticut. (Norton.) B. Cilia of the inferior orbit pale. 3. D. setifer LOEW. £ . — Obscure viridis, pechim nigrorum tibiis an- terioribus posticarumque dimidio basali flavis, facie alba, ciliis oculorum inferioribus tegularumque ciliis albidis, metatarso postico valde setoso. Dark green ; feet black, the four anterior tibise and the basal half of the two hind ones yellow ; face white ; the cilia of the lower orbit and of the tegulse whitish ; the first joint of the hind tarsi with many thorn-like bristles. Long. corp. 0.17. Long. al. 0.15. SYN. Dolichopus setifer Loew, Neue Beitr. VIII, 12, 3. Face silvery-white, narrow. Antennae black, third joint short. Front- metallic green. The cilia of the inferior orbit whitish. The lamellae of the hypopygium whitish ; their long apical margin is finely jagged only below and fringed with long bristles, above only finely hairy ; its lower corner has a narrow black border. The black femora show a green reflection and have a clayish-yel- low tip ; the hind femora have a bristle before their tip and are ciliated on their under side with long black hairs ; the four anterior tibiae are clayish-yellow ; the first joint of the four anterior tarsi DESCRIPTION OF THE SPECIES. 31 is of the same color; its tip and the remaining joints are black. The hind tibia? are black ; upon their upper side, from the base beyon'd the middle, clayish-yellow ; towards their tip but little thickened ; the first joint of the hind tarsi is covered with many thorn-like bristles. Tegula? with whitish cilia. Wings entirely hyaline, towards the base somewhat wedge-shaped, with a black spot which occupies their very tip. The costa is hardly thickened near the tip of the first longitudinal vein ; the end of the fourth longitudinal vein converges towards the third. Eab. District Columbia ; Trenton Falls, 1ST. Y. (Osteu-Sacken.) 4. D. altoiciliatlis LOEW. 9 • — Obscure viridis, peduin nigrorum tibiis, posticarum apice tarnen excepto, tarsorumque anteriorum basi flavis, facie alba, ciliis oculorum iuferioribus albis, tegularum ciliis nigris. Dark green ; feet black, tibise, excepting the tip of the hind ones, and the basis of the four anterior tarsi, yellow ; face white ; cilia of the inferior orbit white ; cilia of the tegulse black. Long. corp. 0.20. Long. al. 0.20. SYN. Dolichopus albidliatus LOEW, Berl. Ent. Monatschr. VI, 211, 59. Dark green. Face proportionally rather broad, white. An- tennas black, third joint short. Front metallic green. Cilia on the inferior orbit white. Femora black, with a greenish reflection, their extreme tip brownish-yellow ; on the under side of the hind femora the hairs are somewhat longer than in the related species, and of such a kind as to lead to the supposition that the male has the hind femora ciliated with black. Tibia? yellow, the tip of the hind ones black ; this black coloring occupies on the anterior side perhaps the fourth, on the hind side nearly the third part. On the four anterior tarsi the four last joints, including the tip of the first joint, are black ; the hind tarsi are entirely black ; their first joint is as long as the second, upon its upper side with two bris- tles, and upon its under side with one. Cilia of the tegula? black. Wings hyaline. Hob. Illinois. (Le Baron.) 5. D. xantliocnemus, n. sp. £ and 9- — Obscure viridis, pedum nigrorum tibiis, posticarum apice taruen excepto, tarsorumque anteriorum basi flavi^, facie alM, ciliis oculorum inferioribus albis, tegularum ciliis nigris, fernoribus maris posticis albo-ciliatis. Dark green ; feet black, tibiee, excepting the tip of the hind ones, and the 32 DIPTEEA OF NORTH AMERICA. [PART II. base of the four anterior tarsi, yellow ; face white ; cilia of the inferior orbit white ; cilia of the tegulse black ; hind femora of the male ciliated with white. Long. corp. 0.15 — 0.16. Long. al. 0.17 — 0.18. In the color and even the structure of the body this species re- sembles D. albiciliatus; but it is much smaller. Pace white, that of the male rather narrow, that of the female proportionally rather broad. Palpi blackish at the base, at the tip more yellowish and with a somewhat whitish reflection. Antenna? black ; third joint rather short, in the g somewhat larger than in the 9. Front metallic green. Cilia of the inferior orbit black above, white on the side and below. Coxas black, trochanters brownish-yellow. The fore coxa? dusted on their fore side and beset with black hair. Femora black, with brownish -yellow tip, the hind ones before the tip with a bristle. The hairs of the femora are black, but there are on the .under side of the four anterior femora of the male some delicate and short white hairs. The under side of their hind femora is ciliated with very long white hairs ; upon the under side of the anterior femora of the female the hairs are closer than in the male, and show a whitish appearance only in a certain direction, whilst in another direction they appear rather blackish. Tibise yellowish ; the hind tibiae are colored with black at the tip ; this black coloring has but a moderate extent and is rather sharply limited. Tarsi plain in both sexes; on the four anterior ones the tip of the first joint and the four following joints are co- lored black ; the hind tarsi are entirely black ; in the male their first joint has more thorn-like bristles than in the female. Cilia of the tegulse black, but mixed with minute white hairs, as it is also the case with the related species, for instance with D. albici- liatus. Wings grayish hyaline ; in the male the anterior margin has at the tip of the first longitudinal vein a small knot-shaped swelling. The white lamellae of the hypopygium are of a rounded- ovate form, bordered with black, jagged on the upper and apical margins and fringed with black bristles. Nab. Sitka. (Sahlberg.) Observation. — Between the diagnosis of this species and that of D. albiciliatus there is no difference with regard to the female sex except in the size. Indeed the females of both species are very much alike. In order to distinguish them it will be well to bear in mind that the female of D. xanthocnemus is not only DESCRIPTION OF THE SPECIES. 33 much smaller, but that the incisures of its abdomen are less blackened, that the black coloring on the. tip of its hind tibia? is less extended and more sharply limited, and that finally the short hairs upon the under side of its fore and middle femora have, in a reflected light, for the most part a whitish appearance, whilst in the female of D. albiciliatus this is the case only in the proximity of the base of the fore femora. 6. D. tetricilSj n. sp. ^ and 9 . — Obscure viridis, rarius cupreus, antennis pedibusque nigris, apicali femorum triente testaceo, inferioribus oculorum ciliis flavicantibus, alis ciuereis. £ . Facie exochraceo-cine- rea, lamellis hypopygii albidis, ultimo tarsorum anticorum articulo dilatato, tibiis posticis crassiusculis. 9 • Facie albida, pedibus simpli- cibus. Dark green, seldom coppery ; antennae and feet black, the last third of the femora brownish-yellow ; cilia of the inferior orbit yellowish ; wings grayish. £ . Face yellowish-gray ; lamellae of the hypopygium whitish ; the last joint of the fore tarsi enlarged : hind tibiae somewhat thickened. 9- Face white; feet plain. Long. corp. 0.18 — 0.20. Long. al. 0.20 — 0.22. Dark green, bright; the last segments of the abdomen are usually dark bronze-colored, and sometimes the color of the body is everywhere very coppery. Antennae black ; the third joint almost round, still with a sharp projection at the tip. Front me- tallic green. The face of the male not very narrow, yellowish gray ; the face of the female much broader, whitish. Palpi black, dusted with whitish on the edge, particularly in the female. Cilia of the inferior orbit- yellowish, sometimes nearly whitish. Hypopygium with elliptic, whitish lamella? of moderate size, which are bordered with black on the upper and apical margin ; the latter is somewhat jagged. Coxse black ; fore coxa? with black hairs and bristles, only very slightly dusted with white. Feet black; the last third of the fore and middle femora, as also somewhat more than the last quar- ter of the hind femora, reddish luteoua-yellow or almost yellowish- red ; the hind femora before the tip with *a seta, not ciliated upon their under side, even in the g . The tibia? show only at the very extreme base a lighter color ; in the 9 all tibia? are of a plain structure, in the J* the hind tibia? are conspicuously thickened. The 9 has plain tarsi, in the male the last joint of the fore tarsi is flattened from the side, and enlarged above into a lobe ; the first joint of the hind tarsi in both sexes is but little fringed with 3 34 DIPTERA OP NORTH AMERICA. [PART II. thorn-like bristles. Cilia of the tegulse black. Wings grayish with brownish-black veins ; the last section of the fourth longi- tudinal vein has a rather strong flexure in the middle, but from that point runs nearly parallel with the third longitudinal vein ; the g has no swelling of the costa at the tip of the first longi- tudinal vein. Eab. Fort Resolution, Huds. Bay Territory. (Kenuicott.) T. I>. acuminatlis LOEW. £ and 9 • — Obscure viridis, pedum ni- grorum tibiis anticis ex flavo fuscis, facie alba, ciliis oculorum inferiori- bus albis, tegularum ciliis nigris, lamellis hypopygii magnis, acutis. Dark green ; the feet black, fore tibise brownish-yellow ; face white ; cilia of the inferior orbit white, of the tegulse black ; the large lamellae of the hypopygium pointed at the end. Long. corp. 0.15. Long. al. 0.14 — 0.15. SYN. Dolichopus acuminatus LOEW, Neue Beitr. VIII, 12, 4. Male. Dark green. Face narrow, white. Antennae black; their third joint short. Front metallic green. Lamellae of the hypopygium white, large, spatule-shaped, pointed, so that the upper and the lower margin strike together and there is no dis- tinct apical margin ; the upper one has a narrow black border and is ciliated with minute black hairs. The black femora with a bluish-green reflection, and their extreme tip brownish-yellow ; fore tibise upon the greatest part of the upper side brownish-yel- low, dark brown beneath ; fore tarsi brownish-black with yellow base ; middle tibise and middle tarsi brownish-black, still the base of the latter yellowish-brown ; hind tibise and hind tarsi entirely black, the latter sparsely bristly upon the first joint. Cilia of the tegulse black. Wings hyaline ; the costa at the tip of the first longitudinal vein not thickened ; the end of the fourth vein con- verging towards the third; the hind transverse vein somewhat less steep than in D. ovatus. Female. Very much like the male. The white face much broader than in the male, still not so broad as in the female of D. albiciliatus. The yellow coloring upon the upper side of the fore tibise is not only generally lighter than in the J*, but also often distinctly observable upon the first half of the upper side of the middle tibise. Hab. Washington. (Osten-Sacken.) Illinois. (Le Baron.) DESCRIPTION OF THE SPECIES. 35 8. D. ovatlis LOEW. £ . — Obscure viridis, pedum nigrorum tibiis an- ticis ex flavo fuscis, facie alba, ciliis oculorum inferioribus albidis, tegu- larum ciliis nigris, lamellis hypopygii parvis, rotundato-ovatis. Dark green ; feet black, fore tibise brownish-yellow ; face white ; cilia of the inferior orbit whitish, of the tegulse black ; lamella? of the hypopygium small, rounded-ovate. Long. corp. 0.15. Long. al. 0.14 — 0.15. SYN. Dolichopus ovatus LOEW, Neue Beitr. VIII, 13, 5. Face narrow, white. Antennas black, third joint short. Front metallic green. Cilia of the inferior orbit whitish. Lamella? of the hypopygium white, rather small, roundish-ovate, on the upper and apical margin with a narrow black border, on the latter split into a bristle-like lobe and fringed with black bristles. Feet black ; the femora with a somewhat greenish reflection ; their ex- treme tip brownish-yellow ; the root of the fore and middle tarsi are of the same color. Fbre tibite upon the greatest part of the upper side brownish-yellow, beneath dark brown. Middle tibios and middle tarsi brownish-black. Hind tibiae and hind tarsi black. Cilia of the tegulse black. Wings hyaline ; the costa at the tip of the first longitudinal vein not thickened. The end of the fourth longitudinal vein converges towards the third ; the hind transverse vein straight and steep. . Hab. Middle States. (Osten-Sacken.) Observation. — The ? of D. ovatus, which is not known to me, must resemble very much that of.D. acuminatus. Besides the but slight discrepancy in the color of the feet, the difference con- sists probably in the coarser hairs on the fore coxae ; at least the hairs in the J* of D. ovatus are coarser and also longer than in that of D. acuminatus. The difference in the direction of the hind transverse vein, in the hiales of both species, is not sufficient to be relied upon for an easy discrimination of the 9 of the same species. II. PREVAILING COLOR or THE FEET YELLOWISH. A. Cilia of the inferior orbit black. 9. I>. pachycnemus LOEW. £ and 9. — Aeneo-viridis, antennis, oculorum tegularumque ciliis nigris, coxis omnibus totis obscuris, pedi>- bus flavis, dimidio tibiarum posticarum apicali, tarsis intermediis inde ab articuli primi apice, posticisque totis nigris. £ . Tarsis anticis attenuatis, articulis tribus ultimis atris, compressis, 36 DIPTERA OF NORTH AMERICA. [PART II. duobus ultimis valde dilatatis, femoribus posticis nigro-ciliatis, tibiis posticis incrassatis. 9 . Pedibus simplicibus, tarsis anticis inde ab articuli primi apice nigris. Metallic green ; antennae, cilia of the inferior orbit and of the tegulse black ; all coxae entirely dark ; tarsi yellow ; the whole second half of the hind tibiae, the middle tarsi from the tip of the first joint and the whole hind tarsi black. $ . Fore tarsi attenuated; the three last joints black, compressed, the two last joints much enlarged; hind femora ciliated with black; hind tibiae incrassated. 9- Feet plain, fore tarsi from the tip of the first joint blackened. Long. corp. 0.25—0.26. Long. al. 0.22. SYN. Dolichopus pachycnemus LOEW, Neue Beitr. VIII, 13, 6. Dark metallic green, bright. Face of the £ narrow, ochre-yel- low; face of the 9 broad, grayish-yellow. Antenna? entirely black ; the third joint ovate. Front bright, generally for the most part steel-blue. The cilia of the inferior orbit black. Thorax with a rather broad brass-colored middle line. Abdomen coppery towards the end. All the coxae black, only their extreme tip somewhat brownish-yellow. Feet dark yellow; hind femora on the extreme tip darker, before the same with a strong bristle ; all tibiae with numerous bristles \ hind tibiae upon the entire second half black; middle tarsi from the tip of the first joint black; hind tarsi entirely black. Tegulae with black cilia. Wings some- what shorter than usual, in the 9 somewhat more dusky than in the male, especially somewhat darker towards the fore margin. Male. Fore tarsi not very much longer than the fore tibia? ; their first and second joints slender, stalk-like, yellow ; the first one much longer than the second ; the following joints black, flat- tened ; the third somewhat broader towards the end ; the fourth much enlarged, of a nearly triangular form, beset upon the upper side with short, close, minute black hairs ; the fourth joint is like- wise much enlarged, still not as broad as the third, and of a more ovate form. Hind tibiae much thickened, stoutest in the middle, and marked upon the upper side with a narrow pale line running from the middle to the tip ; hind femora ciliated beneath with long black hairs. Lamellae of the hypopygiurn dingy yellow, with a broad black border, of moderate size and of a rather round shape, jagged on the apical margin and fringed with black bristles. Female. Feet plain ; the fore tarsi rather blackish already from DESCRIPTION OP THE SPECIES. 37 • the tip of the first joint, which is sometimes the case in £ , but only in very dark-colored specimens. Hob. Middle States. (Osteu-Sacken.) Illinois. (Kennicott.) 1O. E>. brevipeimis MEIG. £ and 9 . — Aene"o-viridis, antennis, oculorum tegularumque ciliis nigris, coxis anticis praeter basim pedi- busque flavis, triente tibiarum posticarum apicali, tarsis intermediis inde ab articuli primi apice, posticisque totis nigris. £ . Tarsis anticis attenuatis, articulis duobus ultimis atris, compressis, ultimo eximie dilatato, femoribus posticis pallide-ciliatis. 9 . Pedibus simplicibus, tarsis anticis inde ab articuli primi apice nigris. Metallic-green ; antennae, cilia of the inferior orbit and of the tegulje black ; fore coxae, excepting the basis, and feet yellow ; the last third of the hind tibiEe, the middle tarsi from the tip of the first joint and the whole hind tarsi black. $ . Fore tarsi attenuated, the two last joints black, flattened, the last one extremely enlarged ; the hind femora ciliated with pale hairs. 9 . Feet plain ; the fore tarsi from the tip of the first joint black. Long. corp. 0.24— 0.25. Long. al. % 0.22; 9 0.24—0.25. STN. Dolichopus plumitarsis (var. 5.) FALLEN, Dol. 10, 4. Dolichopus brevipennis MEIGEN, Syst. Beschr. iV, 89, 27. — STANNIUS, Isis, 1831, 60, 12. — ZETTERSTEDT, Ins. Lapp*. 700, 8. — STAEGEB, Kroyer Tidaskr. IV, 23, 13. — ZETTEKSTEDT, Dipt. Scand. II, 603. — WALKER, Dipt. Brit. I, 160, 10. Dark metallic green, bright. Front green. Antennae black. Face of the g grayish-yellow; that, of the 9 grayish- white, slightly yellowish, much broader than in the £ . Palpi brown. Cilia of the inferior orbit black. Hypopygium black. Lamellae large, oblong-ovate, dingy-yellowish, with a broad black border, on the apical margin somewhat jagged and fringed with crooked black bristles. The hind coxas blackish, only on the extreme tip yellow. Fore coxae yellow, hairy with black ; on the outside of the basis with a somewhat triangular greenish-black spot. Feet yellow. Hind femora before the end only with one bristle ; in the J* they are sparsely ciliated beneath with very long, yellowish hairs. On the hind tibias the last third at least is black ; hind tibiae of the g perceptibly stronger than those of the 9 , but not near so strong as those of the male of D. pachycnemus ; they have no large bare spot upon their hind side, but at the end of the upper side a straight, pale, longitudinal line occupying the whole of their last third. The fore tarsi of the J* are about 1 £ DIPTERA OP NORTH AMERICA. [PART II. the length of the tibiae; their three first joints are style-like, very attenuated, generally brownish-yellow ; their two last joints are black, flattened from the side, the penultimate but little, the last joint on the contrary very much enlarged. The fore tarsi of the 9 are plain, blackish from the tip of the first joint. Middle tarsi of both sexes plain, black from the tip of the first joint ; upon the second half of the upper side of their first joint a single stronger bristle is inserted. Hind tarsi always altogether black. Cilia of the tegulas black. Wings of the J1 somewhat less grayish than those of the 9 , proportionally somewhat shorter and more pointed, with a strong swelling of the costa near the tip of the first longi- tudinal vein. The hind transverse vein almost perpendicular; the last portion of the fourth longitudinal vein not broken. Sab. Fort Resolution, Hudson's Bay Territory. (Kennicott.) Observation. — The identity of Mr. Kennicott's specimens with the European D. brevipennis may be considered as doubtless, after the most careful comparison. B. Cilia of the inferior orbit pale. A. Cilia of the tegulse pale. 1. Antennae black, at the utmost the larger portion of the first joint red. 11. D. longimanus LOEW. £ and 9 — Aeneo-viridis, facie maris pallide ochracea, fceminae ex flavo cinerascente, antennis nigris, ciliis oculorum inferioribus teguhirumque ciliis flavicantibus, coxarum anti- carum dimidio basali obscuro, pedibus flavis, tarsis posticis totis nigris ; maris ultimo tarsorum anticorum articulo valde dilatato femoribusque posticis ciliatis. Metallic green ; the face of the % pale ochre-yellowish, that of the 9 yel- lowish-gray ; antennae black ; cilia of the inferior orbit and of the tegulze yellowish ; basal half of the fore coxae dark ; feet yellow with entirely black hind tarsi, last joints of the % fore tarsi enlarged and $ hind femora ciliated. . Long. corp. 0.26 — 0.27. Long. al. 0.27 0.28. SYN. Dolichopus longimanus LOEW, Neue Beitr. VIII, 14, 7. Bronze-green, rather bright. Face of the £ narrow, pale ochre-yellow; face of the 9 rather broad, light yellow-grayish. Antennae entirely black, lower corner of the third joint with, a scarcely perceptible lighter coloring ; third joint of the antenna in the £ short ovate, in the 9 almost round. Front bright, green or blue-green. The cilia of the inferior orbit yellow. DESCRIPTION OF THE SPECIES. 39 Upper side of the thorax somewhat pruinose, with a coppery middle stripe and with a somewhat coppery-colored mark on each side before the transverse suture, sometimes coppery on a larger extent. Basal portion of the fore coxae blackish beyond the mid- dle ; on the middle and hind ones this coloring extends almost as far as the extreme tip. Feet yellow ; fore tarsi black at the tip only, middle ones from the tip of the first joint ; hind tarsi entirely black ; the hind femora before the tip with a bristle. Tegulse with yellowish cilia. Wings rather large, hyaline ; the fourth longi- tudinal vein not broken. Male. The lamellae of the hypopygium whitish, on the upper margin with a narrow, on the apical one with a broader black border, ovate ; their apical margin is fringed with black bristles and jagged on its lower part. Fore tarsi slender and nearly twice as long as the tibiae ; the four first joints yellow ; first joint as long as four-fifths of the tibias ; second joint half as long as the first ; the third one but little shorter than the second ; the fourth only about half as long as the third ; the fifth joint but little longer than the fourth, black, flattened, beset upon the upper margin with appressed black hairs. Hind femora upon the second half of the under side closely ciliated with yellow hairs. Hind tibiae somewhat stout, without being actually thickened ; they are glabrous upon the anterior half of their hind side. Costa near the tip of the first longitudinal vein with an elongated swelling. Hab. English river. (Kennicott.) West Point, N. Y. (Os- ten-Sacken.) 12. D. Ibrevinaanus LOEW. £ . — Aeneo-viridis, facie alba, antenna- rum nigrarum articulo primo subtus rufo, ciliis oculorum inferioribus albis, tegularum ciliis pallide flavicantibus, pedibus flavis, coxis anticis concoloribus, tarsis anticis ex flavo pallide ferrugineis, intermediis inde ab articuli primi apice posticisque totis cum tibiarum aplce nigris. Metallic-green ; face white ; the first joint of the black antennse upon the under side red ; cilia of the inferior orbit white, on the tegulae yellowish ; the fore coxse and feet yellow ; the fore tarsi pale rusty-brownish ; mid- dle ones from the tip of the first joint and the whole hind ones, including the tip of the tibiae, black. Long. corp. 0.17. Long. al. 0.17. SYN. Dolichopus brevimanus LOEW, Neue Beitr. VIII, 14, 8. Metallic-green, bright. Face white. Antennae black; the whole lower margin of the first joint red ; third joint ovate, not DIPTERA OF NORTH AMERICA. [PART II. rounded at the tip. Front bright bluish-green. Cilia of the in- ferior orbit white. •• Lamellae of the hypopygium white, of medium size and of ovate form, upon the upper and apical margin with a narrow black border, and fringed with black bristles; on the apical margin somewhat jagged. Fore coxae yellowish-white, with- out minute black hairs upon the anterior side ; middle and hind coxaa blackish with whitish-yellow tip. Feet light yellow ; hind femora beset with somewhat longer minute black hairs, without being actually ciliated. Hind tibiae plain, upon the hind side without glabrous stripe, at the tip blackish. Fore tarsi only as long as the tibiae and but little darker than those, yellow- brownish; middle tarsi black from the tip of the 'first joint; hind tarsi entirely black. Tegulae with pale yellowish cilia. Wings hyaline, towards the fore margin with a faint brownish-gray tinge ; costa near the tip of the first longitudinal vein although somewhat stouter, not actually thickened; the fourth longitu- dinal vein not broken. Hob. Washington. (Osten-Sacken.) • 13. 1>. socilis LOEW. $ .— Aeneo-viridis, facie alba, antennarmn ni- grarum articulo primo subtus rufo, ciliis oculorum inferioribus albis, tegularum ciliis dilute flavicantibus, pedibus flavis, coxis anticis con- coloribus, tarsis anterioribus inde ab articuli primi apice, apice tibiarum posticarum tarsisque posticis totis nigris. Metallic-green ; face white ; the first joint of the black antenna upon the under side red ; cilia of the inferior orbit white ; cilia of the tegulse pale yellowish ; fore coxae and feet yellow, the four anterior tarsi from the tip of the first joint, the tip of the hind tibiae and the whole hind tarsi black. Long. corp. 0.17. Long. al. 0.17. SYN. Dolichopus socius LOEW, Berl. Entom. Zeitschr. VI, 211, 60. Metallic-grpen, bright. Face white. Antennas black; the whole lower margin of the first joint red, the third joint ovate, not rounded at the tip. Front bright, bluish-green. Cilia of the in- ferior orbit white. Lamellae of the hypopygium white, of medium size, ovate, on the upper and apical margin with a narrow black border and fringed with black bristles, on the apical margin somewhat jagged. Fore coxae yellowish-white, without minute black hairs upon the front side ; middle and hind coxaa blackish, with yellowish tip% Feet pale yellow ; hind femora upon the un- der side with but extremely short hardly visible hairs ; hind tibiaa , DESCRIPTION OP THE SPECIES. 41 blackish at the tip ; upon the hind side without glabrous stripe ; fore tarsi but little longer than the tibiae, rather slender, blackened from the tip of the first joint, still so that the extreme basis of the second and sometimes even that of the third joint remain pale ; the last joint of the fore tarsi is light rusty -brownish ; middle tarsi from the tip of the first joint black ; hind tarsi entirely black. Cilia of the tegulse pale yellowish. Wings hyaline, towards the fore margin with a faint brownish-gray tinge ; the costa near the tip of the first longitudinal vein only very little stouter, but not thickened ; fourth longitudinal vein not broken. Hob. Illinois. (Le Baron.) Observation. — In order to .distinguish D. socius from the very like D. brevimanus, it is only necessary to pay attention to the difference in the length and coloring of the fore tarsi and also to the different nature of the hairs on the under side of the hind femora. From D. subciliatus, nudus and splendidus, D. socius differs by its hind tibias, which are distinctly blackish at the tip, whereas in D. subciliatus and nudus they are only slightly infus- cated at the tip, and altogether yellow as far as the extreme tip in D. splendidus. From D. subciliatus and D. splendidus it further differs by the red coloring of the lower margin of the first joint of the antennas and by the hind femora of the £ which are not ciliated ; from all three above named species it differs by the plain fore tarsi of the £ . 14. D. nudus, n. sp. £ and £ . — Viridis, nitidus, coxis anticis pedi- busque flavis, apice tibiarum posticarum subfusco, antennis prseter in- . ferum articuli primi marginem tarsisque posticis nigris, ciliis oculorum inferioribus tegularumque ciliis flavicantibus. £ . Tarsis anticis elongatis tenuibus, articulo ultimo dilatato atro, femori- bus posticis non ciliatis. 9 . Pedibus simplicibus, tarsis anticis inde ab articuli primi apice nigris. Green, bright ; fore coxae and feet yellow ; tip of the hind tibiae brownish ; hind tarsi black; lower margin of the first joint of the black antennae red ; cilia of the inferior orbit and of the tegulse yellowish. £ . Fore tarsi elongated, slender, their enlarged last joint black ; hind femora not ciliated. 2 . Feet plain ; fore tarsi from the tip of the first joint blackened. Long. corp. 0.21—0.22. -Long. al. 0.25. Male. Metallic-green, bright. Front metallic-green. An- tennae black ; the under side of the first joint yellowish-red ; third 42 DIPTERA OF NORTH AMERICA. [PART II. joint rather blunt at the tip. Face ochre-yellow ; palpi yellow. Cilia of the inferior orbit pale yellowish. Hypopygium black ; lamellae of medium size, ovate, white, with a rather narrow black border, on the apical margin jagged and fringed with black bristles. Four posterior coxae blackish with yellow tip. Fore COXJE yellow, only somewhat blackened at the extreme basis, beset upon the an- terior side with delicate black hairs, which reach nearly to their base. Feet pale yellow. Hind femora before the tip with a bris- tle, upon the under side only with very short minute pale hairs. Hind tibiae stout but not exactly thickened, upon the first half of the hind side without hairs, at the extreme tip brownish. Fore tarsi twice as long as the tibiaj ; their four first joints very thin, yellow ; first joint nearly as long as the three following ones together ; the third somewhat shorter than the second ; the fourth hardly half as long as the third ; the fifth joint black, flattened, broad, still not as large as in D. batillifer ; upon its upper margin it is beset with appressed minute black hairs. Middle tarsi from the tip of the first joint blackened ; hind tarsi entirely black. Cilia of the tegulae yellowish-white. Wings hyaline ; near the tip of the first longitudinal vein with a long but not very thick swelling, which gradually merges into the costa ; fourth longitudinal vein not broken. Female. Wings and feet plain, fore tarsi from the tip of the first joint blackened. All the rest as in the male. Hob. Fort Resolution, Hudson's Bay Territory. (Kennicott.) Observation. — The resemblance of the present species with D. subciliatus is so great that, as both occur in the same region, I was for a long time in doubt whether I should consider them as two species. As the unfringed under side of the hind femora of my two g of D. nudus does not seem to be rubbed off; as, in both sexes, the first joint of the antennae is tinged with red on the under side, and as there are some other 'differences besides (which will be seen by the comparison of the descriptions of both species), I am led to the conclusion that there is a specific difference between D. nudus and D. subciliatus. It is very striking how closely allied the species of some groups of North American Dolichopus are. 15. D. Sllbciliatlis, n. sp. £ . — Viridis, nitidus, coxis anticis pedi- busque flavis, apice tibiarum posticarum subfusco, antennis tarsisque DESCRIPTION OF THE SPECIES. 43 posticis nigris, ciliis oculorum inferioribus tegularumque ciliis flavican- tibus. . Tarsis anticis elongatis tenuibus, articulo ultimo dilatato atro, feruo- ribus posticis rare flavo-ciliatis. Green, shining ; fore coxse and feet yellow ; tip of the hind tibise brownish ; antenuse and hind tarsi black ; cilia of the inferior orbit and of the tegulse yellowish. £ . Fore tarsi elongated, slender ; the enlarged last joint black ; the hind femora sparsely ciliated with yellow. £ ....... Long. corp. 0.21—0.22. Long. al. 0.25. Metallic-green, bright. Front metallic-green. Antennae black ; on the under side of the first joint I am not able to discover any lighter coloring ; third joint rather blunt at the tip. Face gray- ish-yellow, nearly ochre-yellow. Palpi yellow. Cilia of the in- ferior orbit pale-yellowish. Hypopygium black ; lamella? of medium size, ovate, white, with narrow black border, on the apical margin somewhat jagged and fringed with black bristles. The four poste- rior coxa? blackish with yellow tip. Fore coxa? yellow, only at the extreme basis somewhat blackened, upon the anterior side beset with very delicate black hairs, which do not reach to their .-base. Feet pale yellow. Hind femora with a bristle before the tip, upon the greater part of the under side sparsely ciliated with moderately long yellowish hairs. Hind tibia? stout, but not exactly thickened, not hairy upon the first half of the hind side ; brownish at the extreme end. Fore tarsi twice as long, as the tibia? ; their four first joints very slender, yellow ; the first joint nearly as long as the three following ones together ; the third somewhat shorter than the second ; the fourth hardly half as long as the third ; the fifth joint black, flattened, broad, still not as large as in D. batillifer ; upon its upper edge it is beset with appressed minute black hairs. Middle tarsi from the tip of the first joint blackened. Hind tarsi entirely black. Cilia of the tegula? yel- lowish-white. Wings hyaline, somewhat grayish, at the tip of the first longitudinal vein with a long but not very stout swelling, which gradually merges into the costa ; fourth longitudinal vein not broken. Hob. Fort Resolution, Hudson's Bay Territory. (Kennicott.) Observation. — The necessary data for the distinction of this species from the foregoing, is contained in the description of the latter. 44 DIPTERA OF NORTH AMERICA. [PART II. 16. D. splendidus LOEW. £ . — Aeneo-viridis, Isetissime cupreo mi- cans, pedibus flavis, coxis anticis tibiisque posticis totis 'concoloribug, antennis tarsisque posticis nigris, ciliis oculorum inferioribus tegula- rumque ciliis flavicantibus. £ . Tarsis anticis modice elongatis, articulo ultimo dilatato atro, femoribus posticis confertim flavido-ciliatis. 9 Metallic-green, with, a brilliant coppery-red reflection ; feet and fore coxae yellow, hind tibiae not darker at the tip ; antennae and hind tarsi black ; cilia of the inferior orbit and of the tegulae yellowish. £ . Fore tarsi but moderately elongated ; the enlarged last joint black ; hind femora closely ciliated with yellowish. 9 Long. corp. 0.24. Long. al. 0.23—0.24. SYN. Dolichopus splendidus LOEW, Neue Beitr. VIII, 14, 9. Bronze-green, with a brilliant coppery-red reflection, very bright. Face pale-yellowish. Antennae entirely black ; the third joint short-ovate. Front bright green with a coppery-red reflec- tion. Cilia of the inferior orbit pale yellowish. Upper side of the thorax with an almost purplish spot on each side before the transverse suture. Lamellae of the hypopygium dingy whitish, rather large and of an oval form, on the upper and apical margin narrowly bordered with black, on the latter jagged and fringed with black bristles. The four posterior coxse blackish, yellow only at the extreme tip. Fore coxse entirely yellow, beset upon their anterior side with rather minute black hairs, which do not reach to their base. Feet yellow. Hind femora before the tip with a bristle and upon the greater part of their under side closely ciliated with very long yellowish hairs. Hind tibiae although stout, but not exactly thickened, without hairs upon the anterior half of the hind side. Fore tarsi not quite 1^ times the length of the tibiae ; the four first joints yellow ; the first to the third stalk-like, still not quite as slender as in the previous and in the three fol- lowing species ; the fourth joint somewhat broader, particularly towards its tip ; the first joint nearly as long as the three following taken together, the third somewhat shorter than the second and the fourth distinctly shorter than the third ; the fifth joint black, flat- tened, broad, particularly towards its tip, still by far not as large as in D. batillifer; upon its upper margin it is beset with appressed minute black hairs. Middle tarsi blackened from the tip of the first joint ; hind tarsi entirely black. Cilia of the tegulae whitish. Wings hyaline, a little grayish, of a rather equal breadth; the DESCRIPTION OF THE SPECIES. 45 costa near the tip of the first longitudinal vein with an elongated but not very stout swelling; the fourth longitudinal vein not broken. Hob. Illinois. Observation 1. — I believe I possess also the 9 of this species in a single specimen, and would not doubt it at all if the tip of the hind tibiae did not show a rather distinct brown coloring, which is not perceptible in the g. All the other 'marks in the 9 are precisely as they might be expected in the 9 of the above described tudinal vein not broken. Hale. Lamellae of the hypopygium whitish, of moderate size and of a rather oval shape ; on the upper and apical margin they are narrowly bordered with black ; on the latter very much jagged and fringed with black bristles. Hind femora very closely ciliated with yellow hairs on the middle of the under side. Fore tarsi about l£ times the length of the tibiae ; their three first joints slender and stalk-like, the first as long as the second and third taken together ; the second about 1|- times the length of the third ; the fourth joint somewhat more than half as long as the third and somewhat broader ; upon the whole under side and upon the upper side, excepting the tip, whitish ; the fifth joint somewhat longer than the third and fourth taken together, flattened, extremely broad, black with a silk-like reflection ; upon its outside this reflec- tion takes a handsome silvery hue when viewed in a very ob- lique direction. Hind tibia? considerably thickened, the greatest thickness somewhat before the middle ; the anterior half of their hind side without pubescence. The costa near the tip of the first longitudinal vein with a rather elongated swelling. Hob. West Point, N". Y. ; Illinois. (Osten-Sacken.) Con- necticut. (Norton.) IS. U. eildactylus LOEW. £ and 9 • — Aeneo-viridis, pedibus flavis, coxis anticis tibiisque posticis totis concoloribus, antennis tarsisque posticis nigris, horum basi flava, ciliis oculorum inferioribus tegula- ruinque ciliis flavicantibus, maris duobus ultimis tarsoruni auticorum articulis dilatatis et femoribus posticis ciliatis. Metallic-green ; fore coxae and feet yellow ; hind tibise not darker at the tip ; the black hind tarsi with yellow root ; cilia of the inferior orbit and of the tegulse yellowish ; the two last joints of the fore tarsi of the % enlarged and the hind femora ciliated. Long. corp. 0.26. Long. al. 0.25. SYN. Dolichopus eudactylus LOEW, Neue Beitr. VIII, 16, 11. Metallic-green, bright. Face of the g narrow, ochre-yellow- ish ; the face of the 9 broader and yellowish-white. Antenna entirely black, the lower corner of the first joint hardly somewhat lighter ; third joint short. Front green, or bluish-green, bright. Cilia of the inferior orbit yellowish. The more bronze-colored middle line of t^e thorax usually but little distinct ; on each side DESCRIPTION OP THE SPECIES. 47 before the suture a bronze-colored spot. Fore coxse yellow, upon the anterior side with black pubescence. The four posterior coxge yellow only at the extreme tip. Feet yellow ; hind femora with a bristle before the tip. The color of the fore tarsi changes from brown into black towards the tip ; middle tarsi blackened from the tip of the first joint ; the hind tarsi are of the same color. Cilia of the tegulse yellowish. Wings grayish-hyaline, the fourth longitudinal vein not broken. Male. The larnellre of the hypopygium whitish, of medium size and of ah elongated, ovate shape, on the upper and apical margin narrowly bordered with black, on the latter much jagged and fringed with partly black, partly yellowish bristles. Hind femora upon the under side sparsely ciliated with very long yellowish hairs. Hind tibia? although somewhat stout, but not thickened ; the two- thirds of their hind side from the base are without hairs. Fore tarsi over 1^ times the length of the tibiae ; their three first joints slender, stalk-like ; the second joint measures nearly three-fourths of the length of the first, and the third more than three-fourths of the length of the second ; the fourth joint is almost as long as the third, flattened somewhat broader at the end, still even here not half as broad as it is long, white, with a handsome silvery reflec- tion ; upon its upper margin with hardly visible short minute black hairs ; the fifth joint is hardly shorter than the fourth, flat- tened, and somewhat broader than the previous one, black, beset upon the upper margin with short appressed minute black hairs. The costa near the tip of the first longitudinal vein with a not very considerable but quite distinct swelling ;" the margin of the wing between the apex and the tip of the fifth longitudinal vein is less rounded than usual, so that there is a trace of a shallow sinus. Hab. New York. (Osten-Sacken.) Mass. (Sanborn.) 19. I>. tonsils LOEW. £ and 9 . — Aeneo-viridis, pedibus flavis, coxis anticis tibiisque posticis totis concoloribus, antennis tarsisque posticis nigris, horum basi flava, ciliis oculorum inferioribus tegularumque ciliis flavicantibus, maris duobus ultimis tarsorum anticorum articulis dila- tatis et femoribus posticis non ciliatis. Metallic-green ; fore coxae and feet yellow, the hind tibiae not darker at the tip ; antennse and hind tarsi black, the latter at the basis yellow ; cilia of the inferior orbit and of the tegtilse yellowish ; in the £ the two 48 DIPTERA Or NORTH AMERICA. [PART II. last joints of the fore tarsi enlarged, and the hind femora not ciliated. Long. corp. 0.26. Long. al. 0.25. SYN. Dolichopus tonsus LOEW, Neue Beitr. VIII, 16, 12. Metallic-green, bright. Face of the J* narrow, more light ochre-yellowish than golden-yellow ; the face of the 9 broader and yellowish-white. Antenna? entirely black, the lower corner of the first joint only lighter ; the third joint short. Front green, or bluish-green, bright. Cilia of the inferior orbit yellowish. A bronze-colored middle line of the thorax is not distinct ; on each side before the transv.erse suture a bronze-colored spot. Fore coxas yellow, upon the anterior side with rather delicate minute black hairs, which gradually disappear towards their base. Hind coxa? only at the extreme tip yellow. Feet yellow ; hind femora with a bristle before the tip. The coloring of the fore tarsi in the 9 gradually changes towards the tip into brown and black ; mid- dle and hind tarsi black from the tip4of the first joint, still the base of the second joint yellow. Cilia of the tegula? yellowish. Wings grayish-hyaline ; the fourth longitudinal vein not broken. Hale. Lamellae of the hypopygium whitish, of medium size and of an elongated ovate form, on the upper and apical margin nar- rowly bordered with black, on the latter much jagged and fringed with partly black, partly yellowish bristles. Hind femora not ciliated upon the under side. Hind tibia? of ordinary stoutness ; their whole hind side uniformly beset with minute black hairs. The fore tarsi more than 1^ times the length of the tibise ; the three first joints slender, stalk-like ; the second joint is equal to about two-thirds of the length of the first, and the third to about three- fourths of the length of the second ; the fourth joint is nearly as long as the third, flattened, at the end somewhat broader, still even here not half so broad as long, white, with a handsome sil- very reflection ; upon its upper margin with hardly visible and short minute black hairs ; the fifth joint as long as the fourth, flat- tened, and somewhat broader than the previous one, black, beset upon its upper margin with short and appressed minute black hairs. Costa near the tip of the first longitudinal vein without swelling ; between the apex of the wing and the tip of the fifth longitudinal vein the margin shows a shallow sinus. Hob. Washington. (Osten-Sackeu.) Observation. — The 9 of the three species described above are DESCRIPTION OF THE SPECIES. 49 very easy to confound. The characters by which they can be dis- tinguished are as follows : 1. for D. batillifer 9 : The costa near the tip of the first longitudinal vein with a slight trace of a swell- ing, perceptible only to a very attentive observer ; this swelling is rather elongated ; the apex of the wing somewhat broader than-Jn the 9 of the two other species ; the fore coxae beset with rather coarse minute black hairs reaching almost to the base. 2. for D. eudactylus 9 : The costa near the tip of the first longitudinal vein with a slight trace of a swelling, which has only a very trifling extent in length ; the apex of the wing somewhat narrower than in D. batillifer, but somewhat broader than in D. tonsus ; the fore coxae covered with rather coarse minute black hairs reaching till very near the base. 3. for D. tonsus 9 : The costa near the tip of the first longitudinal vein without any trace of a swelling ; the tip of the wing somewhat narrower than in the two other spe- cies ; the black pubescence of the fore coxae more delicate and not reaching as far towards the base as in the two latter species. Whether the slight differences in the coloring of the tarsi, percepti- ble in the specimens of these species compared by me, are sufficient for their sure distinction, can only be decided by the examination of a larger number of specimens. 2. Antennse altogether or for the most part yellowish-red. 2O. I>. tener LOEW. £ . — Viridis, facie ex flavo albida, antennis rufis, oculorum ciliis inferioribus tegularumque ciliis albidis, pedibus pallide flavis, ultimo tarsorum anticorum articulo maris modice dilatato, nigro. Green ; face yellowish-white ; antennae red ; cilia of the inferior orbit and of the tegulse whitish ; feet pale yellow ; the enlarged last joint of the fore tarsi of the male black. Long. corp. 0.15. Long. al. 0.17. SYN. Dolichopus tener LOEW, Neue Beitr. VIII, 17, 13. Green, bright. Face yellowish-white. Antennae yellowish-red, the apical margin of the third joint blackened ; arista with a very short but still perceptible pubescence. Front metallic-green, bright. Cilia of the inferior orbit whitish. Upper side of the thorax not very bright. Lamellae of the hypopygium white, rounded, slightly bordered with black, on the apical margin some- what jagged and fringed with black bristles. Coxse whitish-yellow, the middle ones upon the outside partly gray ; the fore ones have, besides the black bristles near their tip, only a very short and deli- 4 50 DIPTERA OP NORTH AMERICA. [PART II. cate white pubescence. Feet whitish-yellow ; hind femora before the tip with a bristle, and upon the second half of their under side ciliated with six to seven very long yellowish-white hairs. Fore tibiae long and slender ; hind tibiae rather stout, but not thickened, only at the base of the hind side somewhat glabrous. Fore tarsi filiform, over If times the length of the tibiae ; the four first joints pale-yellowish, their relative length about as 5 : 4 : 3 : 1£ ; their fifth joint but little shorter than the fourth, somewhat flat- tened, black, beset upon its upper side with rather appressed black hairs. Middle and hind tarsi from the tip of the first joint brown- ish. Cilia of the tegulae yellowish-white. Wings hyaline, rather narrow ; the costa near the tip of the first longitudinal vein with a distinct swelling ; the third longitudinal vein not broken ; the hind transverse vein perpendicular and straight. Sab. Chicago. (Osten-Sacken.) 21. D. variaMlis LOEW. $ and $ . — Laete viridis, facie maris pal- lide aurea, prope os albida, facie fceminse tot§, alba, antennis rufis, ocu- lorum ciliis inferioribus albidis, tegularum ciliis flavicantibus, pedibus flavis, alis cinereo-hyalinis. Handsome green ; the face of the £ pale golden-yellow, whitish below ; the face of the J white ; antennae red ; cilia of the inferior orbit whitish, cilia of the tegulse yellowish ; feet yellow ; wings grayish-hyaline. Long, corp. 0.19. Long. al. 0.19. SYN. Dolichopus variabilis LOEW, Neue Beitr. VIII, 17, 14. Handsome green, bright. Face of the g more pale gold-colored than ochre-yellowish, near the oral border whitish ; face of the ? proportionally narrow, white. Antennas yellowish-red ; their third joint short, ovate, generally red only at the base and on the under side, otherwise brownish-black, sometimes brown only at the tip or entirely reddish-yellow. Front bright green. Fore coxae yellow- ish-white, beset, besides the black bristles near their tip, with very delicate minute whitish hairs. Middle and hind coxae of the same color, but colored with black upon the greater part of the outside. Feet yellowish ; hind femora before the tip with a bristle, in the J1 ciliated upon the under side with very long pale-yellowish hairs. Fore tarsi of the $ about \\ times the length of the tibiae ; those of the 9 hardly as long as the tibiae, blackened from the tip of the first joint, plain also in the $. Middle tarsi of the same color as the fore tarsi. Hind tibiae also in the £ not stout, but DESCRIPTION OP THE SPECIES. 51 upon their basal half on the inside glabrous. Hind tarsi usually quite black ; sometimes their first joint, with the exception of the tip, is only brownish or even yellowish ; more seldom the basis of their second joint has also the same lighter coloring. Cilia of the tegulse yellowish. Wings grayish hyaline, of rather equal breadth ; the costa has in the £ near the tip of the first longitudinal vein a slight swelling ; the fourth longitudinal vein is not broken ; some- times the apical portion of the anterior part of the wings shows a somewhat stronger grayish tinge. Hob. New York. (Osten-Sacken.) Observation. — D. variabilis is more variable in the coloring of the posterior tarsi than is usually the case with the species of the genus Dolichopus. I have received a female as belonging to the variety of D. variabilis, having light feet, which has the tegulae ciliated with black, and differs besides from the other females un- doubtedly belonging to D. variabilis, by the more clayish-yellow color of its wings. I cannot take it for the 9 of the present spe- cies. In the coloring of the wings and in several other characters it approaches very much D. luteipennis, but as its hind tibiae are without spots, it may perhaps not even belong to this species. I possess also some other females which I can only distinguish from the above described 9 of D. variabilis by their black ciliated tegulas. It seems therefore that either the females vary in the coloring of the cilia, or that we have here two exceedingly similar species. 22. D. luteipennis LOEW. £ . — Laete viridis, facie albida, antennis rufis, oculorum ciliis inferioribus albidis, tegtilarum ciliis flavicantibus, pedibus flavis, tarsis maris simplicibus, alls lutescentibus. Handsome green ; face whitish ; antennae red ; cilia of the inferior orbit whitish ; cilia of the tegulae yellowish ; feet yellow ; the fore tarsi of the £ plain ; wings yellowish. Long. corp. 0.19. Long. al. 0.19. SYN. Dolichopus luteipennis LOEW, Neue Beitr. VIII, 18, 15. Handsome green, moderately bright. The color of the face whitish, only upon its upper part somewhat more yellowish. An- tennae yellowish-red ; their third joint short-ovate, perceptibly in- fuscated at the tip. Front bright, green-blue. Cilia of the in- ferior orbit whitish. Upper side of the thorax dusted, and hence somewhat dull, with a rather distinct brassy -yellow middle line. The pubescence of the abdomen is whitish, not only on the lateral 52 DIPTERA OF NORTH AMERICA. [PART II. margin, as in the resembling species, but also upon the greater part of the upper side. Lamellae of the hypopygiuni of ordinary size, rounded-ovate, on the upper and apical margin narrowly bor- dered with black, the latter jagged and fringed with black bristles. Fore coxae yellowish-white, and except some black bristles near their tip, beset with only very delicate minute whitish hairs. Mid- dle and hind coxae of the same color, the former darker only at the basis. Hind femora before the tip with a black bristle, upon the under side ciliated with about 6 to T yellowish hairs ; hind tibiae rather stout, but not thickened ; in veiy dark colored specimens sometimes near the tip and also upon the fore and hind side with a narrow brown streak ; upon their hind side only a short glabrous stripe, which does not reach to their middle ; fore tarsi slender, plain, about 1^ times the length of the tibiae ; only their last joint black. Middle tarsi from the tip of the first joint brownish, to- j wards the end blackish-brown, in paler specimens generally but moderately brownish. Hind tarsi generally entirely black, some- times paler towards the end of the first joint or also at the basis of the second joint ; in the palest specimens brownish only towards the end. Cilia of the tegulte yellowish. Wings rather dis- tinctly clayish-yellow ; veins clayish-yellow ; the fourth longitudi- val vein not broken ; the costa near the tip of the first longitudi- nal vein with a little swelling. Hob. Washington. (Osten-Sacken.) Observation. — With regard to a female which may be taken for that of the present species, the necessary remarks have been made in the observation to the previous species. However un- usual the inconstancy in the coloring of the tarsi of D. luteipennis and of D. variabilis may be, I have no doubt that these varieties do not represent different species. At least the most attentive ex- amination of the specimens distinguished by the coloring of their feet, did not lead to the discovery of the slightest difference in the plastic characters in one as well as in the other species. B. Cilia of the tegulse black. 1. Fourth, longitudinal vein broken. a. Antennae black. . D. ramifer LOEW. £ and 9 • — Obscure viridi-aeneus, thoracis dorso aeneo-nigro, facie alba, autennis nigris, ciliis oculorum inferioribus DESCRIPTION OP THE SPECIES. 53 albidis, tegularum ciliis nigris, pedibus flavis, alarum ven& longitudinal! quarta appendiculata. Dark bronze-green, upper side of the thorax bronze-black ; face white ; an- tennae black ; cilia of the lower orbit whitish ; cilia of the tegulse yel- lowish ; the fourth longitudinal vein with a stump of a vein. Long, corp. 0.13. Long. al. 0.13. SYN. Dolichopus ramifer LOEW, Neue Beitr. VIII, 19, 16. Dark bronze-green, the upper side of the thorax more bronze- black. Face of the g nearly as broad as that of the 9, in both sexes white. Antennas black ; the lower corner of the first joint colored somewhat lighter ; in the g the third joint is rather long elliptic with a pointed tip, and the apparently bare arista inserted quite near the tip ; in the 9 it is perceptibly shorter, and has a less pointed tip, to which the arista is still more approximated. Front bluish-black, very bright. Cilia of the inferior orbit whitish. Upper side of the thorax bronze-black, or more dark bronze-green. Abdomen somewhat coppery. Fore coxas dark yellow, at the extreme basis somewhat blackened, beset upon the fore side with short minute black hairs. Middle and hind coxag blackish, dark yellow only at the tip. Feet dark yellow ; the fore and middle tarsi from the tip of the second joint black ; the hind tarsi, including the tip of the hind tibiae, black. The hind femora somewhat broad, before the end with a bristle. Cilia of the tegulse black. Wings hyaline-gray ; the fourth longitudinal vein broken, so that its inferior angle is a right one and the superior is rounded, the former is supplied with a rather long stump of a vein. Male. Lamellae of the hypopygium white, rather small, round- ish, at the upper and apical margin only with an extremely nar- row dark border, on the latter but very little jagged and fringed with black bristles. Hind femora very sparsely ciliated with moderately long whitish-yellow hairs. Hind femora rather stout, the greater part of their hind side glabrous. Costa with a small swelling at the tip of the first longitudinal vein. Hob. Nebraska. (Dr. Hayden.) Lake Winnipeg. (Kennicott.) New Rochelle, N. Y. (Osten-Sacken.) b. Antennae yellowish-red. 24. I>. MfractllS LOEW. £ and 9- — Aeneo-viridis, thoracis dorso pollinoso, opaco, abdomine cupreo-micante, facie albida, interdum ex flavo cinerea, antennis runs, ciliis oculorum inferioribus albidis, ciliis tegularum nigris, alarum venS, longitudinal! quarta appendiculata. 54 DIPTERA OF NORTH AMERICA. [PART II. Bronze-green ; upper side of the thorax not very bright, dull ; abdomen with a coppery-red reflection ; face whitish, sometimes more yellowish- gray ; antennae red ; cilia of the inferior orbit whitish ; cilia of the tegulse black ; the fourth longitudinal vein with a stump of a vein. Long. corp. 0.22. Long. al. 0.17. SYN. Dolichopus bifractus LOEW, Neue Beitr. VIII, 19, 17. Bronze-green, but little shining. Face whitish, in not recently excluded specimens generally more yellowish-gray ; in the J* it is not much narrower than in the 9 . Antennae red ; their third joint ovate, somewhat broad ; at the point of insertion of the arista it is somewhat swollen and blackened ; its apical half is often brownish. Arista with a very short but distinct pubescence. Front bronze-green, but opaque, on account of a very delicate light brownish-gray dust. Cilia of the inferior orbit whitish. Upon the upper side of the thorax the ground color, although bronze- green, is quite opaque, on account of a dense brownish-gray dust, which in fresh, not denuded specimens, altogether conceals it. Abdomen brighter metallic-green, in not recently excluded speci- mens rather coppery. Coxae and feet yellow ; the middle coxae up to the tip gray ; the front side of the fore coxse sparsely beset with delicate minute black hairs, glabrous towards the basis. Hind femora before the tip with a bristle. Fore tarsi brownish, only the last joints really black. Middle tarsi from the tip of the first joint blackish ; the extreme tip of the hind tibiae, including the whole hind tarsi, black. Cilia of the tegulse- black. . Wings with a rather dark grayish tinge ; on the anterior margin and along the veins generally somewhat brownish ; the fourth longitudinal vein broken twice at right angles ; both angles sharp ; as an ex- ception, the upper one sometimes slightly rounded ; at the lower angle there is generally a stump of a vein, whilst the upper one for the most part has none. Male. Lamellae of the hypopygium white, of moderate size, rather rounded, on the upper and apical margin narrowly bor-- dered with black, on the latter somewhat jagged and fringed with black bristles. Fore tarsi 1£ times the length of the tibiae ; first joint somewhat longer than the second and third taken together ; fourth and fifth joints black, somewhat flattened, the upper edge of the fifth bearded with close black hairs. Hind tibiae plain, their hind side not glabrous. Hah. Chicago. (Osten-Sacken.) Nebraska. (Dr. Hayden.) DESCRIPTION OF THE SPECIES. 55 25. D. vittatus LOEW. % . — Aeneo viridis, thoracis linea media vit- tisque lateralibus orichalceis, facie albida, antennis rufis, ciliis oculorum inferioribus albidis, tegularum ciliis iiigris, pedibus flavis, tarsis maris siinplicibus, alarum vena longitudinali quarta fracta et appendiculata. Bronze green ; middle line and the two lateral stripes of the thorax brassy yellow ; face whitish ; antennae red ; cilia of the inferior orbit whitish ; cilia of the tegulse black ; feet yellow ; tarsi of the £ plain ; fourth longitudinal vein broken and furnished with a stump. Long. corp. 0.26—0.27. Long. al. 0.25—0.26. SYN. Dolicliopus vittatus LOEW, Neue Beitr. VIII, 20, 18. Bronze green, shining. Face rather broad for a g , whitish. Antennae red,, apical half of the third joint somewhat infuscated ; arista with a short, distinct pubescence. Front metallic green, ' or greenish-blue, shining. Cilia of the inferior orbit yellowish- white. Upper side of the thorax green or blue, with a conspicu- ous, almost golden-yellow or more copper-colored middle-line, and with similar lateral stripes. Lamella of the hypopygium white, of moderate size, narrow-ovate in shape, with a narrow black margin on the upper and the apical edge ; the latter jagged and fringed with black bristles. Coxa? and feet pale yellowish ; the fore coxae are beset on their anterior and inner side with numerous, on the outer side with very scarce, small, black hairs, besides the white pubescence which clothes them; the middle coxse upon the greater part of their outside blackish. Hind femora before their end with a bristle. Hind tibiae of the ordinary size, upon their hind side with a glabrous stripe, which extends beyond their second third. Fore tarsi plain, only very little longer than the tibiae, from the middle of the third joint black ; the middle and hind tarsi are black from the tip of the first joint. Wings grayish-hyaline, towards the fore margin somewhat more brown ; costa near the tip of the first longitudinal vein with a very thick swelling ; fourth longitudinal vein broken ; superior angle of the fracture rounded, inferior one with only a short stump. Hob. Chicago. (Osten-Sacken.) 26. D. cuprinus WIED. £ and 9- — Aeneo- viridis, thoracis linea mediS, vittisque lateralibus orichalceis, facie ex flavo albida, anteunis rufis, ciliis oculorum inferioribus albidis, tegularum ciliis nigris, 'pedi- bus flavis, tarsorum anticorum apice in mare dilatato, alls basim versus non angustatis, ven& longitudinali quarta fracta. 56 DIPTERA OF NORTH AMERICA. [PART II. Metallic green, middle line and lateral stripes of the thorax brassy yellow ; face yellowish-white ; antennae red ; cilia of the inferior orbit whitish, cilia of the tegulse black ; feet yellow ; tip of the fore tarsi of the £ en- larged ; wings towards the base not narrowed ; fourth longitudinal vein broken. Long. corp. 0.25 — 0.26. Long. al. 0.24. SYN. Dolichopus cupreus SAY, Journ. Ac. Philad. Ill, 86, 9. Dolichopus cuprinus WIEDEMANN, Zweifl. II, 230, 1. — WALKEK, List III, 660.— LOEW, Neue Beitr. VIII, 20, 19. Metallic green, saining. Face whitish, with a more or less yel- lowish tinge, particularly that of the J\ which is considerably narrower than that of the 9 . Antennae yellowish-red, the apical half of the third joint often infuscated. Arista with short but distinct pubescence. Front shining, bluish-green. Cilia of the inferior orbit whitish-yellow. Upper side of the thorax green, often bluish-green, seldom blue, with a conspicuous yellow brass- colored, sometimes copper-colored middle line and with similar lateral stripes. Abdomen usually more bronze-green or coppery. Coxse and feet pale yellowish ; fore coxae only on the inner margin of their anterior side with sparse minute black hairs, which are more distinct in the 9 than in the J* ; middle coxoa upon the greater part of their outside, blackish. Hind femora with a bris- tle before the tip. Fore tarsi from about the middle of the third, middle and hind ones from the tip of the first joint, blackened. Cilia of the tegulas black. Wings tinged with gray, towards the fore margin more grayish-brown, not more narrowed than usual towards the base ; the fourth longitudinal vein broken, so that its inferior angle is a sharp right one and the superior is rounded ; the former is supplied with a short stump of a vein. Male. Lamellae of the hypopygium of moderate size and of elongated-ovate form, white, narrowly bordered with black on the upper and apical margin ; the latter jagged and beset with black bristles. Hind femora upon the second half of their under side sparsely ciliated with very long yellowish hairs. Hind tibiae not thickened, upon their hind side with a glabrous stripe extending beyond the middle. Fore tarsi not one and a half times the length of the tibiae, two first joints stalk-like ; first joint nearly one and a half times the length of the second ; three last joints slightly flat- tened1, third joint upon its upper side very densely fringed with longer, the fourth with somewhat shorter black hairs ; the three DESCRIPTION OP THE SPECIES. 57 last joints of the tarsi are about as long as the second ; the two last ones as long as the third. Hob. Middle States.' (Osten-Sacken.) Nebraska. (Dr. Hayden.) Observation. — The determination of this species, the most common in the Middle States, is not doubtful when we compare the descriptions of Say and Wiedemann, which serve to complete each other. The 9 is easily distinguished from that of D. longi- pennis by its less narrowed basis of the wings. Its distinction from the as yet unknown 9 of D. vittatus must be very difficult, unless perhaps the proportional length of their feet affords an avail- able mark of distinction. Two males measuring only 0.23 in length resemble in all plastic characters the £ of D. cuprinus so much that I take them merely for a large variety. A single, un- fortunately not well-preserved . hastatus, nov. sp. £ and f. — Viridis, antenuis rufis, arti- culo tertio ex parte nigro, ciliis oculorum inferioribus flavis, tegularum ciliis nigris, pedibus flavis, tibiis intermediis maris ante apicern per- spicue, foeminse obsoletissime albido-notatis, tarsis intermediis maris apicem versus compressis, foeminae subconipressis, alarum vena longi- tudinal! quarta non fracta. Green ; antennae red ; third joint partly black ; cilia of the inferior orbit yellow, cilia of the tegulse black ', feet yellow ; middle tibiae before the tip with a white spot, which is very distinct in the $, , and indistinct in the 9 ; middle tarsi strongly flattened towards the tip in the £ , and more slightly in the J ; fourth longitudinal vein not broken. Long, corp. 0.22. Long. al. 0.22—0.23. This species, distinguished by many peculiar marks, resembles in the structure of the £ arista D. Sagittarius Loew, from Sibe- ria. Bronze-green, shining. Face of the <£ narrow and brassy- yellow ; the face of the 9 is much broader, and has a grayish-yel- low tinge. Palpi yellow, in the 9 towards the basis blackish. Antennae reddish-yellow ; their third joint short and rather rounded ; its apical half black or brown ; the arista of the 9 is plain and rather stout ; that of the £ is longer and more slender, enlarged at the tip into the shape of a lancet-like lamel pointed on both sides, the tip of which has a dingy-whitish coloring. Front shining bluish-green. Cilia of the inferior orbit yellow. Fore coxae yellow, upon the anterior side with a delicate black pubes- cence. Middle and hind coxae grayish-black, only the extreme tip and the trochanter yellow. Feet yellow. Hind femora before the tip with a bristle. Middle tibise with black tip, and before it, in the J1 , upon the upper side with a whitish spot, of which there is hardly a trace in the 9 . Hind tibiae at the tip blackish-brown, in the £ upon the hind side with a narrow, linear, hardly distinct glabrous stripe. The tarsi have the tip of the first joint, the four following joints, and besides, the base of the first joint of the mid- dle tarsi, black ; the three last joints of the middle tarsi of the £ are distinctly, although not very strongly, flattened ; this is also the case in the 9 , but less perceptibly so ; this part of the middle tarsi viewed from the side seems distinctly stouter than 60 DIPTERA OP NOETH AMERICA. [PART II. when it is seen from above or below. The first joint of the hind tarsi with only two thorn-like bristles. Cilia of the tegulaa black. Wings grayish-hyaline ; the last segment of the fourth longitudinal vein is inflected before its middle but not broken. The wings of the J* are somewhat narrower than those of the 9 , and have be- fore the anal. angle a very remarkable large bisinuated excision, so that the angle assumes the shape of an independent lobe-like appendage. Lamella? of the hypopygium of moderate size, rounded-ovate, whitish, with a rather broad black border, on the upper and apical margin jagged and fringed with black bristles. Nab. Sdtka. (Sahlberg.) 29. D. pllimipes SCOP. $ and ?. — Viridis, humeris concoloribus, antennis rufis, articulo tertio prseter basim nigro, facie aurea, ciliis ocu- lorum inferioribus flavis, tegularum ciliis nigris, pedibus flavis, articulo tarsorum intermediorum primo maris setulis nigris pennato, alarum vena longitudinal! quarta non fracta. Green, including the humeral callosity ; third joint of the red antennae with the exception of its basis, black ; face golden-yellow ; cilia of the inferior orbit yellow ; cilia of the tegulae black ; feet yellow ; first joint of the middle tarsi of the £ feathered with black bristles ; fourth lon- gitudinal vein not broken. Long. corp. 0.17 — 0.16. Long. al. 0.18. SYN. Musca plumipes SCOPOLI, Ent. Cam. 334, 895. Dolichopus pennitarsis, FALLEN, Dolich. 11, 16. — MEIGEN, Zweifl. IV, 90, 29.— MACQUART, Suites, I, 446, 34.— STANNIUS, Isis, 1831, 63, 14. ZETTERSTEDT, Ins. Lapp., 709, 9. — STAEGER, Kroyer's Tidsskr. 30, 21. — ZETTERSTEDT, Dipt. Scand. II, 541, 35. Dolichopus plumipes WALKER, Dipt. Brit. I, 162, 16. Metallic-green, sometimes somewhat coppery. Antennae yel- lowish-red, third joint, excepting its basis, blackened ; that of the g ovate, with a rather sharp angle at the end ; that of the 9 shorter. Front metallic-green. The face reaches lower here than in most other species of Dolichopus ; that of the g is rather nar- row and golden-yellowish ; that of the 9 is broad and pale-gray- ish, dusted with whitish-yellow. Palpi yellow. Cilia of the in- ferior orbit yellow. Fore coxae yellow, with a black pubescence anteriorly ; middle and hind coxae blackish, the extreme tip and the trochanter yellow. Feet yellow. Hind femora with a bristle before the tip. The middle tibiae are but slightly infuscated near the tip ; the tip of the hind tibiae is somewhat blackish, this color occupying but a narrow extent. Fore and hind tibiae plain in DESCRIPTION OP THE SPECIES. 61 both sexes ; the middle tibiae only in the . sexarticulatlis, n. sp. £. — Aureo-viridis, nitidus, anten- nis rufis, ciliis ooulorum inferioribus flavicantibus, ciliis tegularum nigris, coxis omnibus pedibusque flavis, tarsis posterioribus iride ab articuli primi apice ex fusco nigris, alarum vena longitudinal! quarts, non fracta. £ . Hypopygii apice flavo, lamellis albis late nigro-limbatis, tarsorum an- ticorum articulis tribus primis elougatis, flavis, articulis ultiinis duobus brevibus compressis, paenultimo toto atro, ultimo in basi atro, in apice niveo et appendicula lamellifornii, cum articulo ipso concolore, instructo. 9 Gold-green, shining ; antennae red, cilia of the inferior orbit yellowish, on the tegulas black; all coxae and feet yellow, the middle and hind tarsi from the tip of the first joint brownish-black ; the fourth longitudinal vein not broken. $ . Tip of the hypopygium yellow, lamellae white with a broad black border ; the three first joints of the fore tarsi elongated, yellow, the two last ones short and broadly flattened ; the penultimate entirely black ; the 'last one black at the basis, snow-white at the tip, provided with a lamelliforrn appendage of the same color with this joint. 9 Long. corp. 0.22. Long. al. 0.22. Golden-green, rather shining, still the thorax rather distinctly dusted with ochre-yellow ; front steel-blue ; scutellum greenish- DESCRIPTION OF THE SPECIES. 63 blue. Antennae yellowish-red, the short third joint more brown- ish-red. The narrow face ochre-yellow. Cilia of the inferior orbit pale yellowish. Cilia of the tegulas black. Hypopygium of moderate size, the basal half metallic-green or green-blue, the apical half yellow ; the lamellae rather large and broad, with a rather broad black border, fringed with black on the upper mar- gin only ; jagged as usual, on the apical margin and beset with crooked black bristles. Fore coxas entirely yellow, with a delicate and short pubescence ; near the tip with black bristles. Middle and hind coxae of the same color, still the former more or less blackish at the basis. Femora and tibia? yellow ; hind femora upon the front side before the tip with a stout bristle ; all femora glabrous upon their underside. Fore and middle tibiae plain, rather slender, moderately bristled. Hind tibiae very stout, thickened upon the basal half and with a large glabrous spot upon their hind side. Fore tarsi slender and nearly twice as long as the tibiae ; the elongated, but plain, three first joints yellow, of much decreasing length, the fourth joint deep black, short, broadly flattened, triangular ; the fifth joint of the same shape, hardly some- what longer than the fourth, deep black at the basis, snow-white at the broad tip ; the ungues, pulvilli and empodium inserted at its lower end are of the ordinary structure ; at the upper end there is a distinct elliptical laruel, which, like the joint itself, is deep black at the basis, and snow-white at the tip. Wings grayish- hyaline, rather narrow towards the basis ; near the tip of the first longitudinal vein there is but a slight trace of a swelling of the costa ; the latter, however, is rather stout from this point to the tip of the wing ; .the last segment of the fourth longitudinal vein is only moderately inflected upon its middle. Hob. District Columbia. (Osten-Sacken.) 32. D. ruficornis LOEW. £ . — Viridis, humeris concoloribus, anten- nis rufis, oculorum ciliis inferioribus albis, tegularum ciliis nigris, pedi- bus flavis, articulo tarsorum antieorum ultimo dilatato, alarum vena longitudinal! quarta non fracta. Green, including the humeral callosities ; antennae red ; cilia of the inferior orbit white ; cilia of the tegulae black ; feet yellow ; last joint of the fore tarsi enlarged ; fourth longitudinal vein not broken. Long. corp. 0.19. Long. al. 0.18. SYN. Dolichopus ruficornis LOEW, Neue Beitr. VIII, 21, 21. 64 DIPTERA Or NORTH AMERICA. [PART II. Green, rather shining. Antennae red ; third joint somewhat longer than the two first taken together, of rather equal breadth ; arista distinctly pubescent. Front bright bluish-green. Cilia of the inferior orbit whitish. Humeri of the same color as the remainder of the upper surface of the thorax. Upper side of the abdomen more golden green ; the whitish pubescence on its sides occupies more space than usual. Lamellae of the hypopygium of the ordinary size and rather rounded in shape, white, with a rather narrow black border along the upper and apical edge ; the latter jagged and fringed with black bristles. Coxse and feet pale yel- lowish ; anterior side of the fore coxae only with a very delicate whitish pubescence ; middle coxae with a blackish spot on their outside. Hind femora with a bristle before the tip and sparsely ciliated with long yellowish-white hairs on the latter half of their under side. Hind tibise of ordinary thickness, on their hind side only with a very short glabrous stripe near the basis. Fore tarsi more than once and a half the length of the tibia?, slender ; the four first joints stalk-like, yellowish ; the two first joints, taken together, are somewhat longer than the tibia ; the joints diminishing in length and stoutness from the first to the third ; the fourth joint is equal to about one-third of the length of the third and is only slightly stouter than the latter ; the fifth joint is black, as long as the fourth, somewhat flattened and bearded on its upper side with dense, somgwhat incumbent, short, black hairs. Wings rather yellowish-gray, somewhat narrowed near the basis ; the fourth longitudinal vein not broken ; costa only slightly thickened at the tip of the first longitudinal vein. Hob. Middle States. (Osten-Sacken.) Observation. — If in determining a female specimen, this' species is hit upon, and if its wings, instead of being yellowish gray, are distinctly gray, then it will be necessary to compare what has been said about such females in the observation to the twenty- first species. #. Humeral callosity yellowish. 33. D. scapularis LOEW. ^ and 9 • — Viridis, humeris flavis, facie albida, antennis rufis, oculorum ciliis inferioribus albidis, eiliis tegula- rum nigris, pedibus flavis, tarsis maris simplicibus, alarum vena longi- tudinal! quarta non fracta. Green with yellowish humeri ; face whitish ; antennae red ; cilia of the DESCRIPTION OP THE SPECIES. 65 inferior orbit whitish. ; cilia of the tegulse- black; feet yellow; tarsi of the £ plain; fourth longitudinal vein not broken. Long. corp. 0.25 — 0.26. Long. al. 0.25—0.26. SYN. Dolichopus scapularis LOEW, Neue Beitr. VIII, 22, 22. Bright-green or bluish-green, the abdomen more golden-green, especially towards its tip, sometimes rather coppery. Face whitish, in the J* generally somewhat yellowish upon the upper half. Front shining, generally bluish-green, seldom green or blue. Antennae yellowish-red ; third joint short-ovate ; generally slightly infuscated at the tip ; arista distinctly pubescent. Cilia of the inferior orbit yellowish-white. Humeral callosity yellowish. The callosity between the root of the wing and the scutellum and generally also the margin of the latter are of the same color. Coxae and feet pale yellowish ; the front side of the fore coxae is beset with delicate whitish hairs ; there are some short minute black hairs on their inner side only, which however are not always easily discernible in the g . Middle coxae upon their outside with an elongated blackish spot. Hind femora with a bristle before the tip. The two last joints of the fore tarsi of the ? are black- ish-brown, still the brown color often begins already before the tip of the third joint and the tips of the first and second joints are also often somewhat infuscated ; in pale-colored specimens the coloring of the middle and hind tarsi is the same, whilst in darker ones the distinct brown color begins already in the middle of the first joint. Cilia of the tegnlae black. "Wings grayish, towards the fore margin somewhat yellowish-brown ; fourth longitudinal vein not broken, still the inferior angle of the flexure is some- times not rounded. Male. Lamellae of the hypopygium of moderate size, rather rounded, on the upper and apical margin with a rather broad black border, on the latter, jagged and fringed with black bristles. Hind femora upon the second half of their under side ciliated with moderately long yellowish hairs. Hind tibiae of the usual thick- ness, upon the hind side with a glabrous stripe, which reaches nearly to their middle. Fore tarsi somewhat over once and a quarter the length of the tibiae, their joints decreasing in length, the last one somewhat more pale at the tip. Costa near the tip of the first longitudinal vein with a very slight swelling. Hob. — Middle States ; District Columbia (in June ; Osten- Sacken); Illinois. 5 66 DIPTERA OF NORTH AMERICA. [PART II. 34. D. fimditor LOEW. £ and J . Viridis, liumeris flavis, facie albida, antennis rufis, ciliis oculorum inferioribus albidis, tegularum ciliis nigris, pedibus flavis, tarsorum anticoruin articulis duobus ultimis in mare dilatatis, alarum venS, longitudinali quarta non fracta. Green, with yellowish humeri ; face whitish ; antennae red ; cilia of the inferior orbit whitish ; cilia of the tegulse black ; feet yellow, the two last joints of the fore tarsi of the £ enlarged ; fourth longitudinal vein not broken. Long. corp. 0.25. Long. al. 0.25. STN. Dolichopus funditor LOEW, Neue Beitr. VIII, 22, 23. Bright green or blue-green, the abdomen more golden-green, especially towards its end, sometimes rather coppery. Face whitish, in the g generally somewhat yellowish upon the upper half. Front shining, generally blue-green, seldom blue or green. Antennae yellowish-red ; third joint short-ovate, generally slightly infuscated at the tip ; arista with a distinct pubescence. Cilia of the inferior orbit yellowish-white. Humeral callosity yellowish. The callosity between the root of the wing and the scutellum is usually of the same color and generally also the margin of the latter. Coxae and feet pale yellowish ; the front side of the fore coxae is beset with delicate white hairs ; on their inner side there are also some short black hairs, which however are not always distinct in the g . Middle coxae with an elongated blackish spot upon their outside. Hind femora with a bristle before the tip ; two last joints of the fore tarsi black-brown in the 9, still this brown color often begins before the tip of the third joint and the tip of the first and second joint also are usually somewhat infus- cated; pale colored "specimens have the same coloring on the middle and hind tarsi, whilst in darker specimens a distinct iiifus- cation already begins in the middle of the first joint. Wings grayish, somewhat more yellowish-brown towards the fore margin ; fourth longitudinal vein not broken. Male. Lamellae of the hypopygium of medium size, rather rounded, white, on the upper and apical margin with a narrow black border, on the latter jagged and fringed with black bristles. Hind femora upon the second half of the under side ciliated with not very long yellowish hairs. Hind tibiae somewhat more slender than in the g of the previous species, upon their hind side with a glabrous stripe extending beyond their middle. Fore tarsi once and a quarter the length of the tibiae ; the three first joints of moderate size and of decreasing length ; fourth joint flattened, DESCRIPTION OP THE SPECIES. 67 black, on its upper edge fringed with close black hairs ; the fifth joint still more flattened, ovate, snow-white, upon its upper side with very short and fine minute snow-white hairs. The costa at the tip of the first longitudinal vein only with an insignificant swelling Hab. Middle States. (Osten-Sacken.) Observation. — Of this and of the preceding species I possess so many specimens taken together that I can entertain no doubt about having the ? of both before me. Unfortunately I found it impossible as yet to discover any reliable character to distinguish these females. &.' Antennae black, at the utmost the greater part of the first joint red. 35. D. chrysostomus LOEW. £. — Viridis, facie aurea, antennis nigris, ciliis oculorum inferioribus albidis, tegularum ciliis nigris, pedi- bus flavis, tarsis maris simplicibus. Green ; face golden-yellow ; antennas black ; cilia of the inferior orbit whitish ; cilia of the tegnlse black ; feet yellow ; tarsi of the $ plain. Long. corp. 0.18. Long. al. 0.17. STN. Dolichopus chrysostomus LOEW, Neue Beitr. VIII, 23, 24. Green, shining. Face narrow, dark golden yellow. Antennae altogether black ; third joint elongated-ovate with a pointed tip ; arista rather slender with a somewhat imperceptible pubescence. Front shining blue-green. Cilia of the inferior orbit whitish. Upper side of the thorax but little dusted ; on each side, at the transverse suture, with a coppery-brown spot. Abdomen with rather apparent dark incisures, which, in fully colored specimens, are margined with coppery-red. The lamellae of the hypopygium are large, rounded, yellowish-white, with a not very narrow black border on the upper and the apical edges ; the latter is jagged and fringed with black bristles. Fore coxae yellow, somewhat blackened only at the base, clothed anteriorly with short black hairs, middle and hind coxae blackish, their extreme tip only yel- low. Feet yellow. The hind femora with a bristle before the tip and with short yellowish hairs on the underside, although not ciliated with them in the true sense of the word. Hind tibiae not stout ; their posterior side without glabrous stripe. Fore tarsi plain, but little longer than the tibiae, gradually infuscated towards the tip. Middle and hind tarsi infuscated from the tip of the first joint ; however the second and third joints of the middle and the 68 DIPTEKA OP NORTH AMERICA. [PART II. second joint of the hind tarsi, except its tip, are still rather pale. Tegulae with black cilia. Wings tinged with gray ; more brown- ish-gray along the anterior border ; costa with a hardly perceptible thickening at the tip of the first longitudinal vein ; fourth longi- tudinal vein not broken. Hob. Washington, D. C. (Osten-Sacken.) 36. lJ. praeustus LOEW. £ . — Aeneo-viridis, nitens, facie ex cinereo ochracea, antennis nigris, ciliis oculorum inferioribus albidis, tegularura ciliis nigris, pedibus testaceis, litura femorum anteriorum, femorura posticorum apice, tarsis anterioribus inde ab articuli primi apice, tarsis posticis totis cum tibiarum posticarum apice nigris, alarum ex cinereo hyalinarum apice nigro. * Bronze-green, shining; face grayish ochre-yellow; antennae black; cilia of the inferior orbit whitish ; cilia of the tegulse black ; feet luteous- yellowish ; a stripe on the fore-femora, the tip of the hind femora, the four anterior tarsi from the tip of the first joint, and the hind tarsi altogether, as well as the tip of the hind tibiae, black ; the tip of the grayish-hyaline wings black. Long. corp. 0.21. Long. al. 0.21. STN. Dolichopus prseustus LOEW, Berl. Ent. Zeitschr. VI, 212, 62. Bronze-green, shining. Face grayish-yellow. Antennae alto- gether black ; their third joint short. Front metallic green, some- what dusted and therefore but little shining. Occipital bristles very long. Cilia of the inferior orbit whitish. Last segments of the abdomen generally somewhat colored with coppery. Lamellae of tH*e hypopygium of medium size only, broad, with very rounded upper border, white, bordered with black, bristly along the upper and apical edge ; the latter somewhat jagged. Coxae blackish, their tip brownish-yellow ; the fore coxae with a grayish-white dust and hairy with black. Feet brownish-yellow ; the fore femora on the under side with a brownish-black longitudinal stripe ; the hind femora before the tip with a black bristle and tinged with black on the upper side of the tip ; the delicate hairs on their under side are somewhat longer than in many other species, although they cannot be called cilia. The hind tibiaa are blackened towards the tip, and have, towards the end of the upper side, a rather dis- tinct dimple or impression ; their hind side has no glabrous stripe. Tarsi plain, the four anterior ones from the tip of the first joint, the two hind ones altogether black. Cilia of the tegulae black. Wings grayish hyaline, the extreme tip of the wing black ; the DESCRIPTION OP THE SPECIES. 69 third longitudinal vein is somewhat directed backwards towards its end, so that its tip is nearer to the tip of the fourth vein than is usual in other species ; the last portion of the fourth longitudi- nal vein is almost straight ; on the spot where its usual flexure is situated, the surface of the wing is distinctly convex ; costa but indistinctly thickened at the tip of the first longitudinal vein. Hob. Illinois ; (Le Baron. ) Observation. — It is to be presumed, judging from the analogy of similar European species, that the 9 of D. praeustus has no black spot at the tip of the wing. 31?. D. comatus LOEW. £ and 9 • — Viridis, facie Candida, antennis nigris, ciliis oculorum inferioribus albis, tegularum ciliis nigris, pedibus flavis, feinoruni posticorum apice superne nigro, alarum vena longitu- dinal! quart§, non facta. Green, face snow-white, antennae black ; cilia of the inferior orbit white ; cilia of the tegulse black ; feet yellow, the tip of the hind femora black above ; the fourth longitudinal vein of the wings not broken. Long, corp. 0.17. Long. al. 0.16. SYN. Dolichopus comatus LOEW, Neue Beitr. VIII, 23, 25. Green or bronze-green, shining. Face snow-white, that of the 9 rather broad. Antennae altogether black, the third joint short- ovate ; arista with an almost imperceptible pubescence ; cilia of the inferior orbit white. Front shining green. Abdomen with distinct dark incisures. Fore coxae yellow, blackened only at their very base ; in the % their inner side and their tip only, in the 9 almost the whole anterior side is beset with small blackish hairs. Middle and hind coxae blackish, their very tip only yellow. Feet yellow ; the hind coxae with a bristle before the tip and tinged with brownish-black on the upper side of their tip. Hind tibiae slender, their tip black. Fore and middle tarsi black from the tip of the first joint ; hind tarsi altogether black. Cilia of the tegulae bback. "Wings tinged with gray; fourth longitudinal vein not broken. Male. Lamellae of the hypopygiurn hardly medium-sized, ovate, whitish, on their apical edge with a vestige only of a narrow black border and very little- jagged, ciliated with hairs, most of which are pale. Hind tibiae on their hind side without any bare stripe. First joint of the fore tarsi a little longer than the four following ones taken together ; the latter are deep black and somewhat flat- 70 DIPTERA OF NORTH AMERICA. [PART II. teuecl. The bristles on the outer side of the middle tibiae are very prolonged and become very slender towards their tip. The first joint of the middle tarsi is of considerable length, ciliated on its upper side with about nine or ten very long, bristle-like black hairs. Hob. Pennsylvania; Maryland; District Columbia. (Osten- Sacken.) 3§. D. scoparilis, nov. sp. •£>. — Viridis, facie Candida, antennis nigris, articulo priino infra rufo, ciliis oculorum inferioribus pallidis, ciliis tegularum nigris, pedibus flavis, tarsis ex parte nigris, articulis antico- rum ultimis duobus subdilatatis et utrinque nigro-pennatis, vena ala- rum longitudinali quarta non fracta. Green, face snow-white, antennae black, the first joint upon the under side red ; cilia of tlie inferior orbit pale ; cilia of the tegulse black ; feet yel- low, tarsi partly black ; two last joints of the fore tarsi somewhat thick- ened, feathered with black on both sides ; the fourth longitudinal vein of the wings not broken. Long. corp. 0.24^ Long. al. 0.24. Bright metallic green. Antennas black, the lower edge of the first joint red ; third joint short. Face snow-white ; palpi brown- ish-yellow, the basis black. Cilia of the inferior orbit yellowish. Front metallic green. Fore coxa3 yellow, blackened at the base only to a moderate extent, upon the front side with a black pubes- cence. Middle and hind coxse blackish, only the extreme tip and the trochanter yellow. Feet yellow. Hind femora before the tip with a bristle. Tibiae plain, with rather numerous black bristles ; hind tibiae at the tip not blackened, upon the hind side without glabrous spot. Fore tarsi from the tip of the third joint black, not quite once and a half the length of the tibiae ; their two last joints are only very slightly enlarged, but . closely feathered with bristle-like minute black hairs upon the front and hind side, so that they seem to be rather broad. Middle and hind tarsi plain, from the tip of the first joint black ; upon the upper side of the first joint of the middle tarsi, not far from the tip, there is a stout black bristle. Cilia of the tegulas black. Wings grayish-hyaline, of the usual form ; fourth longitudinal vein not broken and the fore margin, near the tip of the first longitudinal vein, not thickened. The lamellae of the hypopygium of the only specimen which I possess are almost destroyed; I am able to state only that they are yellowish. Hab. Maine. (Packard.) Mass. (Sanborn.) DESCRIPTION OF THE SPECIES. 71 39. D. discifer STANN. £ and 9 . — Viridis, facie alba, antennis ui- gris, ciliis oculorum inferioribus albis, tegularum ciliis nigris, pedibus flavis, femorum posticoruin apice concolore, tarsorum anticorum articulo ultimo nigro, in mare modice dilatato, alarum vena longitudinal! quarta non fracta. Green ; face white, antennae black ; cilia of the inferior orbit white, cilia of the tegulse black; feet yellow, tip of the hind femora not blackened; the last joint of the fore tarsi black, in the £ moderately enlarged; fourth longitudinal vein of the wings not broken. Long. corp. 0.25 — 0.26. Long. al. 0.24. STN. Dolichopus patellatus MEIGEN, Syst^Beschr. IV, 86, 22. Dolichopus discifer STANNIUS, Isis 1831, 57, 10. Dolichopus confusus ZETTERSTEDT, Ins. Lapp., 709, 7. Dolichopus patellatus STAEGEK, Kroyer's Tidskr. IV, 21, 12. Dolichopus discifer ZETTERSTEDT, Dipt. Scand., II, 533, 28. WALKER, Dipt. Brit. I, 163, 20. Dolichopus tanypus LOEW, Neue Beitr. VIII, 24, 26. Bright green. Face white, in the £ upon the upper half gene- rally more yellowish-white. Antennas black ; first joint upon the under side red, third joint elongated-ovate, rather large; arista with a very short, but distinct pubescence ; it is inserted beyond the middle of the third joint. Front shining green. Cilia of the inferior orbit white. Fore coxse yellowish, beset upon their front side with delicate white hairs, and only on the inner side in the 9 with some black hairs. Middle and hind coxae blackish, at the tip yellowish. Feet yellowish; hind femora with a bristle before the tip. Hind tibiae somewhat brownish-black only at their ex- treme tip, particularly on the inner side. Fore tarsi, although darker from the tip of the first joint, but only the last joint black ; middle tarsi from the tip of the first joint black ; hind tarsi en- tirely black, seldom only brown at the base of the first joint. Cilia of the tegulas black. Wings grayish hyaline ; fourth longi- tudinal vein not broken, towards its end somewhat more converging with the third than in most of the other species. Male. Lamellae of the hypopygium not very large, elongated ovate, white, on the upper and apical margin with a very narrow black border, on the latter somewhat jagged and fringed with black bristles. Hind femora not ciliated. Hind tibiae slender, without glabrous spot upon their hind side. Fore tarsi exceed- ingly slender and elongated, the four first joints yellow, still some- what darker beyond the tip of the first joint, each following joint 72 * DIPTERA OF NORTH AMERICA. [PART II. more slender than the previous one ; the first joint is equal to three-fourths of the tibia, and is somewhat longer than the second and third taken together ; second till fourth joints but very little decreasing in length ; fifth joint deep black, somewhat flattened, so that it appears like a small ovate disk ; at its extreme basis it is colored with yellowish-white. Wings towards the basis rather narrow, though of the usual form. Costa near the tip of the first longitudinal vein with an almost imperceptible swelling. Hob. English River ; Red River. (Kennicott.) Sitka. (Sahl- berg.) White Mountains, N. H. (Osten-Sacken.) Observation. — A very close examination renders it certain that this species, as it appears widely spread in North America, is identical with the European D. discifer. I had overlooked this identity, while describing it from American specimens, as D.. tanypus. 4O. I>. lobatus LOEW. $ . — Viridis, facie dilute lutescente, antennis nigris, inferioribus oculorum ciliis fiavicautibus, tegularum ciliis nigris, pedibus flavis, femorum posticorum apice concolore, tarsis anticis inde ab articuli primi apice nigricantibus, articulo ultimo nigro, in mare latis- simo. Green; face pale luteous-yellow ; antennae black; cilia of the inferior orbit yellowish, cilia of the tegulse black ; feet yellow, the tip of the hind femora not darker ; fore tarsi from the tip of the first joint black- ish ; the last joint black, very much enlarged in the £ . Long. corp. 0.27. Long. al. 0.26. SYN. Dolichopus lobatus LOEW, Neue Beitr. VIII, 24, 27. Bright green. Face pale yellow, rather whitish below. An- tennae black, first joint red with blackish upper edge ; third joint short-ovate. Front shining, green. Cilia of the inferior orbit yellowish. Lamellae of the hypopygium rather large, ovate, white, on the second half of the upper margin and on the apical margin with a rather broad black border, jagged on the latter and beset with black bristles. Fore coxse yellow, upon the front side with minute yellowish hairs, only on their inner side also with a few minute black hairs. Middle and hind coxae blackish, at the tip yellow. Feet yellow. Hind femora not. ciliated ; before the tip with a bristle. Hind tibiae somewhat thickened about the middle and colored with darker yellow upon their second half; their hind side without glabrous stripe. Fore tarsi hardly once and a half DOLICHOPUS. 73 the length of the tibise ; three first joints stalk-like and very slen- der;, from the tip of the first joint black-brown; first joint some- what longer than the second and third taken together ; the third only half as long as the second ; fourth joint very short, somewhat broader than the previous one, brownish-black ; fifth joint black, nearly as long as the second, flattened, very much enlarged, so that it has an almost semi-obcordate shape ; the close black pubes- cence of its upper edge makes it appear still larger and broader. Middle tarsi from the tip of the second joint black. Hind tarsi entirely black. Wings gray, towards the fore margin more gray- ish-brown, narrow; towards the base the hind margin has two very remarkable sinuses, a longer one between the' fifth and sixth longitudinal veins, and a shorter one behind the sixth longitudinal vein, so that there is a lobe between them ; the anal angle of the wing also projects considerably as a rounded lobe ; the fourth lon- gitudinal vein only with a slight flexure, somewhat more converg- ing towards its end with the third than is the case in the related species ; the costa at the tip of the first longitudinal vein with a rather imperceptible swelling. Hob. English River. (Kennicott.) Observation. — I believe I know also the 9 of this species. It differs from the 9 of D. discifer, by its somewhat larger size, its somewhat more yellowish face, and by the fore tarsi being not only shorter, but also tinged with black already from the tip of the first joint. The fore coxae have, upon the greater part of their anterior side, some minute black hairs. Although the fore coxse of the 9 have in many species a more extended black pubescence than the g , the difference between this 9 and the above described J* is more striking than usual. This circumstance will render it some- what doubtful that the two sexes really belong together, until a positive observation settles the question. 41. D. setosus LOEW. 1 . — Viridis, nitidus, facie et inferioribus ocu- lorum ciliis albis, antennis tegularumque ciliis nigris, coxis anticis pedibusque flavis, tarsis anterioribus inde ab articuli prirni apice tar- sisque posticis totis cum tibiarum posticarum apice nigris ; femora pos- tica pilis flavis ciliata ; tibia? posticse setis longis armatse, alarum vena longitudinalis quarta non fracta. Green, shining ; the face and the cilia of the inferior orbit white ; the ' antennae and the cilia of the tegulse black ; fore coxae and feet yellow, the four anterior tarsi from the tip of the first joint and the whole hind 74 EIPTERA OF NORTH AMERICA. [PART II. ones, including the tip of the hind tibise black; hind femora ciliated with yellowish hairs ; hind fibiae armed with long thorn-like bristles ; fourth longitudinal vein of the wings not broken. Long. corp. 0.23 — 0.24. Long. al. 0.25. SYN. Dolichopus setosus LOEW, Berl. Ent. Zeitschr. VI, 213, 63. Green, shining. The narrow face white. Antennae black, the lower edge of the first joint brownish; third joint ovate, not rounded at the tip. Front rather dark green, but little shining. Cilia of the inferior orbit whitish. Lamellae of the hypopygium of medium size, broad, rather rounded, white with narrow border, fringed on the upper and apical margin with black bristles, the latter but little jagged. Fore coxse pale yellowish, dusted with white; their short pubescence near the tip and upon the inner half of their front side black. Middle and hind coxset blackiah, with pale yellowish tip. Feet pale yellowish ; the hind femora have but one bristle before the tip and are ciliated with long yellowish hairs upon the under side ; the hind tibise are black at the tip and have upon their upper, as well as upon their under side, longer bristles than usual ; I am not able to distinguish a glabrous spot upon their hind side, but at the tip of 'the upper side there is a short pale line. Fore and middle tarsi from the tip of the first joint, hind tarsi entirely, black. Wings grayish hyaline ; costa at the tip of the first longitudinal vein distinctly, but not strikingly thickened ; the last section of the fourth longitudinal vein mode- rately inflected upon its middle ; the hind transverse vein perpen- dicular. Hab. Massachusetts. (Le Baron.) 42t D. incisuralis LOEW. £ and J. — Viridis vel aeneo-viridis, facie alba, antennis nigris, ciliis oculorum inferioribus albidis, tegula- rum ciliis nigris, pedibus flavis, femoruni posticorum apice concolore, tarsis anticis inde ab articuli primi apice nigris, in mare simplicibus, alarum vena longitudinal! quarta non fracta. Green or bronze~gre«n ; face white ; antennae black : cilia of the inferior orbit whitish, cilia of the tegulae black, feet yellow, the tip of the hind femora not darker; fore tarsi from the tip of the first joint black, plain evefc in the ^ ; fourth longitudinal vein not broken. Long. corp. 0.17. Long. al. 0.16. gYN. Dolichopus incisuralis LOEW, Neue Beitr. VIII, 25, 28. Green or bronze green, well preserved specimens purer green, GYMNOPTERNUS. 75 shining. Face white. Antennae black ; the inferior edge of the first -joint red or reddish-brown, which, however, is not distinctly perceptible in some specimens ; third joint short ; arista with a hardly perceptible pubescence. Cilia of the inferior orbit white. Front green. Abdomen with remarkably distinct black incisures. Fore coxae whitish-yellow, only at the extreme basis somewhat blackened ; their front side is beset upon its basal half with numer- ous black, very delicate and rather sparse hairs. Middle and hind coxae black, only at the extreme tip somewhat yellowish. Feet yellowish. Hind femora before the tip with a bristle. Fore and middle tarsi blackened from the tip of the first joint ; still the whole first joint is also somewhat dusky. Hind tibiae with a black tip ; hind tarsi entirely black. Cilia of the tegulae black. Wings grayish ; fourth longitudinal vein only with a slight flexure and towards its end somewhat more than usually converging yith the third longitudinal vein. Hale. Lamellae of the hypopygium of medium size and of a rounded-ovate form, white ; on the upper and apical margin with a rather narrow black border, on the latter jagged and fringed with black bristles ; hind femora ciliated with moderately long and very delicate pale hairs. Hind tibiae slender, plain, upon their hind side without glabrous stripe. Fore tarsi plain, about once and a quarter the length of the tibia? ; their first joint is longer than the two following, but somewhat shorter than the three following taken together. Costa at the tip of the first longitudi- nal vein with a very short but distinct swelling. Hob. Trenton Falls, N. Y. (Osten-Sacken.) Gen. III. GYMNOPTERNUS. The following characters of the genus Gymnopternus are to be observed: The first joint of the antenna? is hairy upon the upper side, the third almost never remarkably elongated ; arista dorsal. The hypopygium is entirely disengaged, the exterior appendages are latuelliforin and of moderate size. The first joint of the hind tarsi is shorter than the second and not provided with bristles. The third and fourth longitudinal veins of the wings are parallel or almost so. The last of these characters is applicable to all the North Ame- rican species of Gymno2)ternits known to me. Among the species 76 DIPTERA OP NORTH AMERICA. [PART II. of the old world which have been placed into the genus Gynmop- ternus there certainly is a whole group of closely related species the third and fourth longitudinal veins of which decidedly con- verge. However, as this group must necessarily be separated from the genus Gymnopternus, it could not prevent me from adopting the parallelism of the third and fourth longitudinal veins as cha- racteristic marks of Gymnopternus. Precisely this character dis- tinguishes in the easiest manner the species of Gymnopternus from those of the following genera, which, like Gymnopternus, have the upper edge of the first joint of the antennae hairy, but are without bristles upon the first joint of the hind tarsi. Most of the species of Gymnopternus are small and have shorter antennas, but a more distinctly pubescent arista than the species of Dolichopus. They are much poorer in suitable plastic charac- ters for^the distinction of the species than the latter. The feet of the males are very seldom ornamented. The color of the cilia of the inferior orbit is also of the highest importance for the determination of the species of the present genus. Unfortunately it cannot be so easily observed as in the species of Dolichopus. Its discrimination in some species, of which I have only single specimens, was totally impossible ; as to others, I often remained uncertain. In the former case I have been silent about their color, and in the latter I did not use any positive expressions. Under these circumstances 'it was impossible to use the color of the cilia of the lower orbit as a basis for a subdivision. This is, however, but of little moment, as the cilia of the lower orbit seem to be black in almost all North American species of Gymnopternus. Another important mark for the dis- tinction of the species is the hairy or glabrous surface of the scu- tellum, provided there is a sufficient number of well preserved specimens ; otherwise, if the specimens are few or not well pre- served, this mark will be rather uncertain. I did not wish to omit characters of this kind altogether, but have to request the reader not to place too much confidence in them, especially when my expressions seem to imply doubt. The same rule applies to the form of the lamellae of the hypopygium. In 'many species they have the form of an erect crescent, fastened by its .lower point. This form will only then be recognized, when they are not closely applied to the hypopygium with the concave side ; if the latter is the case, then they appear only as small lamellae, with GYMNOPTERNUS. 77 rounded ends, and the length of which is greater than their breadth. I have, then, called them rounded, without further remarks about their form ; to prevent mistakes, however, I must state that this applies only to the convex edge. I have omitted other characters in the descriptions, because they are common to all North Ameri- can species known to me ; for instance, the presence of only one bristle at the end of the hind femora, etc. This genus derives its name (yv^oj naked, and ttttpvij the sole) from the absence of bristles upon the first joint of the hind tarsi, whereby it differs fro^n the genus Dolichopus, to which its species formerly belonged. The species described by Say as Dolichopus obscurus seems to be a Grjmnopternus. I do not know of any -other species of North American Gymnopternus, described by a previous author. This undoubtedly arises from the circumstance that the species of Gym- nopternus, on account of their small size and their apparent insig- nificance, have been less noticed by collectors. The number of species known to me shows that North America is very rich in species of this genus. To produce a really satisfactory treatise on the subject would require much more material than that over which I could dispose, because the positive discrimination and exact delineation of the characteristics of the species present many difficulties. I will give now a dichotomic table for the purpose of deter- mining the species, and a synopsis of the systematic arrangement. A-s will be seen from the latter, the bulk of the species known to me, are very nearly related and form but a single group; whereas but a small number show characters which isolate them from the others. i Table for the determination of the Species. 1 ( Color non-metallic. 1 flavus Lw. I Color metallic. 2 9 ( Third joint of the antennae with an elongated point. 2 subulatus Lw. \ Third joint of the antennae without elongated point. 3 „ ( Prevailing color of the feet black. 4 I Prevailing color of the feet yellow. 6 . ( Third joint of the antennas remarkably hairy. 3 scotias Lw. \ Third joint of the antennae with scarcely perceptible hairs. 5 , ( Wings grayish hyaline. 4 barbatulus Lw. \ Wings somewhat tinged with blackish. 5 tristis, n. sp. 78 DIPTERA OP NORTH AMERICA. [PART II. g f Tip of the hind femora blackish. 6 ezilis Ln\ I Tip of the hind femora not blackish. 7 - f Thorax dark violet. 8 I Thorax not violet. 9 r, ( Coxae up to the tip somewhat blackish. 7 spectabilis Lw. \ Coxae yellowish-white. 8 albiceps Lw. !Fore coxae up to the tip blackish. 9 subdilatatus Liu. Fore coxae entirely yellow, or at the utmost somewhat infuscated near the base. 10 r Hind tarsi from the tip of the first joint black. 10 leevigatus Lw. 10 -| Hind tarsi towards the tip but little dusky, at the utmost brownish, I never black. 11 ,, f Antennae entirely black. 12 1 Antennae partly red. 16 , £ f Lamellae of the hypopygium black. 11 frequens Lw. I Lamellae of the hypopygium not black. 13 , „ ( Lamellae of the hypopygium dark yellow. 12 lunifer Lw. X Lamellae of the hypopygium white. 14 (Interior appendages of the hypopygium penicillate. 13 fimbriatus Lw. Interior appendages of the hypopygium not penicillate. 15 Third and fourth longitudinal veins but slightly converging. 14 despicatus Lw. Third and fourth longitudinal veins altogether parallel. 15 difficilis Lw. ! Middle and hind coxae from the basis distinctly blackish. 17 Middle and hind coxae yellow, or, at the utmost, the former with a grayish tinge. 19 , - f Lower part of the face of the 9 distinctly hairy. 16 nigribarbus Lw. <• The lower part of the face not hairy. 18 , of Antennae small. 17 parvicornis Lw. I Antennae of tolerable size. 18 opacus Lw. -i q / Venter and posterior margin of the pleurae not yellow. 20 1- Venter and posterior margin of the pleurae yellow. 21 2Q 5 Thorax brightly shining, front white. 19 politus Lw. C Thorax rather dull, front gray. ' 20 debilis Lw. o-i f Hypopygium remarkably stout and large. 21 crassicauda Lw. I Hypopygium of the usual size and thickness. 22 22 f Antennae very small. 22 minutus Lw. I Antennae of middle size. 23 ventralis Lw. 15 -I \ GYMNOPTERNUS. 79 Systematic arrangement of the Species'. I. Coloring of the body non-metallic. 1. flavus Lw. II. Coloring of the body metallic. A. Third joint of the antennae with an elongated point. 2. subulatus Lw. B. Third joint of the antennae without an elongated point. A. Prevailing color of the feet black. 3. scotias Lw. 5. tristis, n. sp. 4. barbatulus Lw. B. Prevailing color of the 6. exilis Lw. 7. spectabilis Lw. 8. albiceps Lw. 9. subdilatatus Lw. 10. laevigatus Lw. 11. frequens Lw. 12. lunifer Lw. 13. fimbriatus Lw. 14. despicatus Lw. feet yellow. 15. difficilis Lw. 16. nigribarbus Lw. 17. parvicornis Lw. 18. opacus Lw. 19. politus Lw. 20. debilisZw. 21. crassicauda Lw. 22. minutus Lw. 23. ventralis Lw, 80 DIPTERA OP NORTH AMERICA. [PART II. DESCRIPTION OF THE SPECIES. I. COLORING OF THE BODY NON-METALLIC. 1. G-. flavus LOEW. £ and £> . — Flavus, abdominis segmentis interme- diis plerumque virescentibus. Yellow ; the middle segments of the abdomen usually greenish. Long, corp. 0.10—0.11. Long. al. 0.12—0.13. SYN. Gymnopternus flavus LOEW, Neue Beitr. VIII, 28, 1. Pale yellowish. Face whitish. Antennae dark yellow, the third joint with a very pointed brownish or blackish tip and with rather distinct hairs, which are visibly shorter in the female. Arista black with an almost imperceptible pubescence. Front and occiput of a greenish color, but thickly dusted with yellow so as to appear dull and altogether light greenish-gray. Cilia of the inferior orbit white-yellowish. Thorax entirely yellow, not unfre- quently with a slight trace of a greenish lustre, its bristles black, the small hairs pale, scutellum provided with two black bristles, otherwise glabrous. Abdomen with yellow hair, the stout hairs on the incisures somewhat darker, but not black ; the middle and sometimes also the posterior segments of the abdomen show a greenish lustre; hypopygium yellow, lamella? small, yellowish- white, without a dark margin, thinly ciliated with short yellowish hairs. Feet white-yellowish, their scanty bristles black ; the smaller hairs yellowish. Cilia of the tegula? yellow. Wings towards the anterior margin yellowish, otherwise more yellow- grayish. Hob. Pennsylvania. (Osten-Sacken.) • • II. COLORING OF THE BODY METALLIC. A. Third joint of the antennae with an elongated point. 2. €r. Sllbulatus LOEW. % . — Viridis, thorace subopaco, antennarum articulo tertio acutissimo, hirto, seta subapicali instructo. Green; thorax rather dull, the third joint of the antennae very pointed, GYMNOPTERNUS. 81 roughly hairy, with a subapical arista. Long. corp. 0.13 — 0.14. Long., al. 0.16. SYN. Gymnopternus subulatus LOEW, Neue Beitr. VIII, 29, 2. Green, made dull by pale gray-brownish dust, especially upon the thorax. Face gray-whitish. The first joint of the antennae black-brown, the second red, the third dark brown, at the root red, unusually long and sharply pointed, and covered with much longer hairs than is the case with the other species of the same genus. The black arista has a hardly perceptible pubescence, is scarcely somewhat longer than the third joint of the antennae, and is inserted about its last third, so as to be nearer to the tip than is the case with the other species. Front, in consequence of a thick covering of dust, dull greenish-gray. The color of the cilia of the inferior orbit cannot easily be recognized, how- ever only the lowest of them may possibly be of a pale color. Thorax and scutellum, on account of a thick covering of dust, pretty dull gray-green ; the scutellum bears, as usual, the two black bristles, and seems otherwise to be entirely without hairs. Abdo- men more green, and brighter than the thorax. The black hypo- pygium rather stout, with small yellowish lamellae, which are cili- ated on the margin with short black hairs and have no dark edge. Interior appendages simple, provided with one hair upon the upper side and with two hairs upon the point, before it is bent down. Coxae and feet pale yellowish, fore coxae with black hair. Cilia of the tegulJB black. Wings somewhat yellow-grayish, large and broad, especially towards the tip ; the anal angle rounded off. Hah. Trenton Falls, N. Y. (Osten Sacken.) B. THIRD JOINT OP THE ANTENNAE WITHOUT ELONGATED POINT. A. Prevailing color of the feet black. 3. G. scotias LOEW. £ and $> . — Atro-virens, pedibus nigris, troehan- teribus, genibus, tibiis, tarsorumque anteriorum basi flavicantibus, tertio antennarum articulo hirto, facie non pilos&. A • Black-green, feet black ; trochanters, knees, tibise, root oT the four anterior tarsi yellowish, the third joint of the antennae roughly hairy, face not hairy. Long. corp. 0.13 — 0.14. Long. al. 0.14 — 0.15. SYN. Gymnopternus scotias LOEW, Neue Beitr. VIII, 29, 3. Dark black-green, or almost metallic black. Face and front gray. Antennae entirely black, the third joint elongated, ovate, 82 DIPTERA OP NORTH AMERICA. [PART II. not very broad, pointed at the end, with longer hairs than in most of the other species ; the arista is inserted in its middle and has a rather indistinct pubescence. Cilia of the inferior orbit black. Scutellum with the usual two bristles ; otherwise I cannot per- ceive any hairs upon its surface. Feet black. Trochanter with the extreme tip of the first joint of the coxa, tip of the femora, the tibiae, and the roots of the four anterior tarsi, yellowish, but, on account of the density of the short black hairs, of pretty dark appearance. The hind side of the hind tibiae is clothed towards its end with dense black hairs, so that it appears pretty black ; the root of the hind tarsi is brown. Cilia of the pale yellowish tegulaa black. Halteres yellow-whitish. Wings gray-blackish, a little darker towards the anterior margin. The small lamellas of the hypopygium are black. Hob. English River. (Kennicott.) 4. G. barbatulus LOEW. $ and 9. — Atro-virens, pedibus nigris, trochanteribus, genibus, tibiis (excepto tamen posticarum apice) tarso- rumque anteriorum basi flavicantibus, alls ex cinereo-hyalinis, infera faciei parte nigro-pilosa. Black-green ; feet black, trochanters, knees, tibiae (with the exception of the tip of the hind ones) and the root of the four anterior tarsi yellowish, wings grayish-hyaline, the lower part of the face with black hair. Long, corp. 0.12. Long. al. 0.12—0.13. SYN. Gymnopternus barbatulus LOEW, Neue Beitr. VIII, 29, 4. Dark black-green, face gray-white, the inferior part of it some- what swollen transversely, and with small sparse black hairs. Antennas entirely black, their third joint broad, pretty rounded, and only with short, scarcely perceptible hairs. Pubescence of the arista extremely short, hardly perceptible. Front dark me- tallic green ; the dust on its surface can only be perceived in an oblique direction. Cilia of the inferior orbit black. Besides the usual two bristles upon the scutellum, there are a few short, extremely slender, and therefore scarcely perceptible hairs. Feet black ; the tip of the first joint of the coxas, the trochanter, the tip of the femora, the tibiae, and the root of the four anterior tarsi yellowish, the tip of the hind tibiae to a moderate extent black. The cilia of the yellow tegulae black. Halteres white- GYMNOPTERNUS. 83 yellowish. The wings dusky with gray ; the small lamella? of the hypopygium brown. Hob. Middle States. (Osten-Sacken.) 5. G. tristis, n. sp. % and 9 • — Atro-virens, pedibus nigris, genibus, tibiis tarsorumque anteriorum basi testaceis, tertio antennarum articulo nudo, alis nigricantibus. Black-green ; feet black, knees, tibiae and the root of the four anterior tarsi brownish-yellow, the third joint of the antennae bare ; wings black- ish. Long. corp. 0.13—0.15. Long. al. 0.14 — 0.15. Resembles much not only the G. scotias, but also G. barbatulus. Black-green, sometimes more black (like ore). Face of the "£ black, of the 9 black-gray, the latter much broader than in the $ ; upon its lower part, in the 9 , several hardly perceptible black hairs, which I did not observe upon the face of the £ . Antennae entirely black ; the third joint broad, rather short, however some- what longer in the £ than in the 9 , bare, that is to say, only with the usual microscopic pubescence, which is very difficult to ob- serve. The rather strong arista is also covered with this almost imperceptible pubescence. Front dark metallic green ; the rather whitish dust upon it becomes visible, when viewed in an oblique direction. Cilia of the inferior orbit black. The scutellum has besides the usual two bristles, also some shorter hair. Feet black, tip of the coxae and trochanters in well matured specimens hardly much paler; tip of the femora, the tibiae, and the root of the four anterior tarsi brownish-yellow ; tip of the hind tibise brownish, the root of the hind tarsi sbmetimes brown. In less matured specimens the lower side of the femora is mostly pitch- brown. Cilia of the tegulse black. The small brownish-black lamellae of the hypopygium are crescent-shaped, and adhere with the concave side to the hypopygium, so that their true form cannot be easily perceived ; on their convex side they are fringed with small blackish hairs, but not jagged. The wings are comparatively long, distinctly tinged with smoky black; the third and fourth longitudinal veins show towards the end an indication of a slight convergeucy; the hind transverse vein is comparatively distant from the margin of the wing. Hob. Sitka. (Wahlberg.) Observation 1. — Gymn. tristis is distinguished from Gymn. 84 DIPTEEA OF NORTH AMERICA. [PART II. barbatulus by its larger size, its longer and darker wings, and by a darker and less hairy face ; the male further differs by the greater length of the lamellae of the hypopygium. From G. scotias it dif- fers by the third joint of the antennas, which has not the long hairs, so apparent in G. scotias. Observation 2. — A male from the same locality shows a con- siderably stronger convergency of the third and fourth longitudinal veins, coincides, however, so much in all the other characters with the rest of the males, that I cannot consider it for more than a variety, although a very striking one, of G. tristis. B. Prevailing color of the feet yellow. 6. G. exilis LOEW. £ . — Viridis, pedibus flavis, coxarum intermedia- rum basi femorumque posticorurn apice nigricantibus, tarsis fuscis. Green, with, yellow feet, the basis of the middle coxae and the tip of the hind femora blackish, tarsi brown. Long. corp. 0.10. Long. al. 0.11. SYN. Gymnopternus exilis LOEW, Neue Beitr. VIII, 30, 5. Green or 'bluish-green, not very bright. Face and front light grayish. Antennae brownish-black ; the second joint and the root of the third reddish-brown ; the third joint comparatively rather large, not very broad in proportion to its size, not rounded at the tip, distinctly hairy ; the pubescence of the not very long arista is difficult to perceive. Cilia of the inferior orbit black. Tho- rax, in consequence of a light cover of dust, somewhat dull, and grayish-green. In one specimen only, I perceive upon the scutellum, besides the usual bristles, a few small hairs, which are rather indistinct. Feet pale yellowish. Middle coxes upon the outside distinctly blackened beyond their middle. Hind coxse darkened only at the basis. Tip of the hind femora distinctly blackened upon the upper side. Fore and middle tarsi infuscated from the tip of the first joint; hind tarsi black-brown to the same extent. The row of short small bristles which is usually found upon the upper side of the fore tibiae in the species of Gymnop- ternus is less developed here than in most of the other species. Cilia of the tegulae black. Wings gray. The small lamellae of the hypopygium yellow, fringed with rather apparent, small black bristles ; their form is rather kidney-shaped, still they have in the lower corner a very small, somewhat protruding black flap ; the GYMNOPTERNUS. 85 interior appendages of the hypopygium bear a few hairs before the tip. Hob. Pennsylvania. (Osten-Sacken.) 7. G. spectaMlis LOEW. 9 . — Thorace violaceo, antennis nigris, coxis nigricantibus, pedibus flavis. Thorax violet, antennae black, coxae blackish, feet yellow. Long. corp. 0.17. Long. al. 0.17. SYN. Gymnopternus spectabilis LOEW, Neue Beitr. VIII, 30, 6. Is among the largest North American species of this genus known to me. Face and front with an almost silvery-white dust, though upon the latter the dust is less thick. Antennae altogether black ; third joint short ; the arista is somewhat stout at the basis and has a plainly perceptible pubescence. Cilia of the inferior orbit black. The upper side of the thorax metallic violet, the scu- tellum likewise ; the latter has some short hairs in the middle. Abdomen blackish metallic green, bright. All the coxae up to the extreme tip blackish. Feet yellow, tarsi from the tip of the first joint infuscated; the usual row of bristles upon the upper side of the fore tibise complete and distinct, though the single bristles are comparatively not long. Cilia of the tegulee black. Wings tinged with gray-brown, towards the anterior margin a little browner ; the third and fourth longitudinal veins perfectly parallel. Hab. New York. (Osten-Sacken.) Observation. — Had Mr. Wiedemann not stated the size of Do- lichopus obscurus Say to be l£ line, I would have most certainly believed that my G. spectabilis is the D. obscurus of Mr. Say. Nevertheless the statements of MM. Wiedemann and Say about D. obscurus do not apply so closely to G. spectabilis as to waive such an important difference and to consider both species as one and the same. 8. G. alMceps LOEW. $ . — Thorace violaceo, antennis rufis in apice fuscis, coxis pedibusque flavis. Thorax violet, the red antennae brown at the tip ; coxae and feet yellow. Long. corp. 0.17. Long. al. 0.17. SYN. Gymnopternus albiceps LOEW, Neue Beitr. VIII, 30, 7. Face very broad, more so than that of G. spectabilis, snow- white. Antennas dusky red ; third joint small, rounded, dark 86 DIPTERA OP NORTH AMERICA. [PART II. brown upon the apical half. Arista with a comparatively long and striking pubescence. Front covered with a snow-white dust. Cilia of the inferior orbit black. Thorax and scutellum metallic violet ; no hairs are perceptible upon the surface of the latter. Abdomen metallic dark green. Coxoa and feet yellowish ; middle coxae upon the outside with a grayish streak. Tarsi from the tip of the first joint gradually blackened. The usual row of bristles on the upper side of the fore tibiae is extant and complete ; the single bristles, however, comparatively short. Cilia of the tegulce black. Wings tinged with grayish-brown, a little more brown towards the anterior margin ; the third and fourth longitudinal veins very slightly converging towards the end. Hob. Middle States. (Osten-Sacken.) 9. Cr. silTjdilatatus LOEW. £ . — Viridis, antennis nigris, coxis n?gri- cantibus, anticarum apice pedibusque flavis, maris tarsorum anticorum articulo ultimo depresso, subdilatato. Green ; antennae black ; coxse blackish ; tip of the fore coxse and the feet yellow; the last joint of the fore tarsi of the £ flattened and a little enlarged. Long. corp. 0.13. Long. al. 0.13. SYN. Gymnopternus subdilatatus LOEW, Neue Beitr. VIII, 31, 8. Metallic green, rather bright. Face covered with a whitish dust. Antennae entirely black ; third joint short, rather rounded ; arista with a scarcely perceptible, extremely short pubescence. The cilia of the inferior orbit seem to be black ; upon the scutel- lum, besides the usual two bristles, a few small hardly perceptible hairs are inserted. The rather large lamellae of the hypopygium are more kidney-shaped than crescent-shaped, upon their lower side brownish-yellow, upon the upper part brownish-black, closely fringed with black bristle-like hairs ; the interior appendages are simple. The fore coxas blackened as far as the middle, middle and hind coxae almost as far as the tip. Feet yellowish, a little more slender than in the allied species. The hairs on the hind femora are also blackish upon their under side, and more distinct than in the related species. The usual row of bristles on the upper side of the fore tibiae is extant, but the single bristles are very short. Fore tarsi from the tip of the first joint strongly iufuscated, towards the tip black, very slender, but hardly longer than the tibiae. Their first joint is as long as the two following taken together ; the last joint is flattened and a little enlarged, GYMNOPTEENUS. 87 the pulvilli also larger than usual. The middle and the hind tarsi strongly infuscated from the tip of the first joint, towards the tip black. Cilia of the tegulce black. Wings tinged with blackish-gray. Hab. Middle States. (Osten-Sacken.) Observation. — A single female specimen agrees with the just described male of G. subdilatatus in the color of the coxae, and cannot therefore belong to any of the other species known to me ; I am prevented, however, from taking it for the 9 of G. subdila- tatus on account of the more clumsy shape of the feet. 1O. Cr. laevigatlis LOEW. % . — Viridis, thorace subcserulescente, niti- dissimo, antennis parvis nigris, articulo secundo et tertii basi obscure rafis, coxis anticis totis pedibusque pallide flavis, tarsis posticis inde ab articuli prinri apice nigris, lamellis hypopygii pallide flavis, appendi- cibus interioribus siruplicibus. Green, with, a somewhat violet, very bright thorax ; the small antennae black, the second joint and the root of the third dusky red ; the whole fore cox£e and the feet yellow, the hind tarsi from the tip of the first joint black ; the lamellae of the hypopygium pale-yellow ; the interior appendages simple. Long. corp. 0.12. Long. al. 0.12. SYN. Gymnopternus lasvigatus LOEW, Neue Beitr. VIII, 31, 9. Green, bright. Fa.ce and front covered with whitish dust. An- tennae small, black ; the second joint and the root of the third dusky red. Arista with a short but distinct pubescence. The cilia of the inferior orbit seem to be black. Upper side of the thorax bluish-green and very bright. Upon the surface of the scutellum, besides the usual bristles, there are a few quite imper- ceptible little hairs. The small lamella? of the hypopygium are light-yellowish, with a scarcely perceptible blackish border and crescent shaped. Coxae and feet white-yellowish ; the middle coxae on the outside almost as far as the tip, and the hind coxae at the root, blackened. The hairs on the feet are somewhat coarse, and the usual row of bristles on the upper side of the fore tibiae con- sists of comparatively long and rather strong bristles. Hind tarsi black from the tip of the first joint ; fore and middle tarsi infus- cated from the same joint. Cilia of the tegula? black ; wings tinged with blackish-gray ; the end of the third and fourth longi- tudinal veins parallel. Hab. Middle States. 88 DIPTERA OP NORTH AMERICA. [PART II. Observation. — If the coloring of the tarsi should not prove con- stant, the distinction from G. parvicornis would be rather diffi- cult. It would then be necessary to observe that the feet of the present species are decidedly somewhat more clumsy and covered with coarser hair, and that the row of bristles on the upper side of the fore tibia? consists of somewhat longer bristles. The con- formity of both species in the structure of the antennas and of the appendages of the hypopygium is striking. G. laevigatus cannot be confounded with any other species, 11. G. frequens LOEW. £ and 9. — Obscure viridis vel aeneo-viridis, antenuis nigris, facie et fronte ex albo cinereis, pedibus flavis, latnellis hypopygii nigris. Dark-green or bronze-green ; antennae black ; face and front whitish-gray ; feet yellow; lamellse of the hypopygium black. Long. corp. 0.12. Long. al. 0.12—0.15. SYN. Gymnopternus frequens LOEW, Neue Beitr. VIII, 32, 10. Blackish-green, recently developed specimens rather bluish- green, more aged specimens darker bronze-green. Face and front covered with a whitish-gray dust. Antennas black, the third joint quite small ; arista with a short but distinct pubescence. Cilia of the inferior orbit black ; upon the surface of the scutellum there are, besides the two bristles, several short hairs. Coxae and feet yellow ; middle coxae almost on their whole outside blackish, or at least brownish ; the fore coxae show only at the extreme basis traces of a brownish tinge, such as is often also perceived on the hind coxa?. The usual row of bristles on the upper side of the fore tibiae is distinct and dense. The hind tarsi become, from the tip of the first joint, more and more brown, their tip is black- brown. The fore and middle tarsi are infuscated in a similar manner, but less dark. Cilia of the tegula? black. Wings tinged with blackish-gray, the third and fourth longitudinal veins with a slight trace of convergency. The lamella? of the hypopygium black, quite rounded at the end, fringed with black hairs ; the in- terior appendages not bristly. Hob. Middle States. (Osten-Sacken.) Observation. — G. frequens is, among the kindred species, the only one whose males have black lamellce, and thus is easy to recognize. Female specimens occur which have the dust upon face and front much whiter ; in other respects they a,re like the GYMNOPTERNUS. 89 other females. Whether they 'are, as I suppose, merely a variety of G. frequens, or whether they belong to another closely re- lated species can only be determined by further observations. The changes in size of the present species are not so striking as would appear from the measurements given above, because the larger specimens are always females, which, in this species, more than usual exceed the males in size. It will be quite difficult to distinguish the female of G. lunifer from that of G. frequens. 12. G. lunifer LOEW. £ . — Obscure viridis vel aeneo-viridis, anten- nis nigris, facie et fronte cinereis, pedibus flavis, lamellis hypopygii obscure luteis. Dark-green or bronze-green, antennae black, face and front gray ; feet yel- low; lamellae of the hypopygium dark-yellow. Long. corp. 0.13 — 0.14. Long. al. 0.13— 0.14. SYN. tiymnopternus lunifer LOEW, Neue Beitr. VIII, 32, 11. Dark-green, rather bright, face and front with whitish-gray dust. Antennae rather short, entirely black, the third joint small, not rounded at the end. Arista with a short but distinct pubes- cence. Cilia of the inferior orbit black. Upon the scutellum, besides the bristles, a few not easily perceptible hairs ; the lamellae of the hypopygium a little larger than those of the kindred species, crescent-shaped, but rounded on the upper end, so as to become somewhat kidney-shaped, and thus to approach the shape of the lamellae of G. subdilatatus. They are of a dingy brownish-yellow color, and upon the upper margin somewhat blackish. Their black fringe is not so strong as that of G. subdilatatus. Fore coxse dark yellow, a little brownish at the extreme basis ; the mid- dle and hind coxas black almost up to the extreme tip. Feet somewhat dark yellow, rather slender, hind femora somewhat iu- fuscated on the upper side towards the tip. The usual row of bristles on the upper side of the fore tibiae is complete. Tarsi brownish towards the tip. Cilia of the tegulse black ; wings tinged with brownish-gray. Hob. New York. (Osten-Sacken.) 13. G. fimbriatus LOEW. $ . — Viridis, pedibus et coxia flavis, coxis intermediis, apice excepto, nigricantibus ; appendicibus hypopygii inte- rioribus elongatis et penioillatis. 90 DIPTERA OP NORTH AMERICA. [PART II. Green, feet and coxse yellow, the middle coxae however, with the excep- tion of the tip, blackish ; the interior appendages of the hypopygium elongated, hairy, penicillate. Long. corp. 0.10. Long. al. 0.11. STN. Gymnopternus fimbriatus LOEW, Neue Beitr. VIII, 32, 12. Rather light-green, bright. Face and front covered with a white-grayish dust. Antennae entirely black, short, the third joint rounded. Arista with an extremely short and very imper- ceptible pubescence. Cilia of the inferior orbit black. The upper side of the thorax moderately bright. Upon the scutellum only traces of very imperceptible hairs. The lamella? of the hypopygium whitish-yellow, crescent-shaped, ciliated with stiff black hairs ; the interior appendages somewhat elongated, with a brush-like tuft of long hairs at the end. Coxa? and feet yellow, more slender than those of the next following species ; most of the outside of the middle coxa? blackish ; the fore and hind coxae hardly somewhat blackened at their extreme basis. Tarsi somewhat infuscated towards the tip, especially the hind ones. The usual row of bristles on the upper side of the fore tibia? is extant ; the single bristles of middle size. Cilia of the tegulse black. Wings tinged with gray. Hob. Maryland. (Osten-Sacken.) 14. G-. despicatlts LOEW. £ . — Viridis, antennis nigris, facie et fronte albido-pollinosis, pedibus flavis, tibiis posticis prope apicem supra paulo lougius pilosis, quam in speciebus ad quas accedit ; alarum venia longitudinalibus tertia et quarta subconvergentibus ; lamellis hypopygii pallide flavescentibus. Green, antennae black, face and front covered with a whitish dust ; feet yellow ; hind tibiae on the upper side towards the end with longer hairs than in the allied species ; the third and fourth longitudinal veins of the wings show a slight convergency ; lamellae of the hypopygium pale yel- low. Long. corp. 0.12. Long. al. 0.12. SYN. Gymnopternus despicatus LOEW, Neue Beitr. VIII, 33, 13. Green, rather bright. Face and front covered with whitish dust. Antennae entirely black and only of middle length ; the third joint rather rounded at the tip. Arista with a very short, hardly perceptible pubescence. Cilia of the inferior orbit black. Upon the scutellum of the described specimen there are, besides the two bristles, only a few small, pale hairs on the mar- gin. Lamella; of the hypopygium pale-yellowish, crescent-shaped, GYMNOPTERNUS. 91 fringed on the margin with small black hairs. The interior ap- pendages plain. Cox9B and feet yellowish. Middle coxa? on the outside as far as somewhat beyond the middle, blackish. The usual row of bristles on the upper side of the fore tibiae is com- plete, and consists of comparatively large bristles. The usual short hairs upon the last third of the upper side of the hind tibia? are not only a little denser, but also visibly longer, than in the allied species. Tarsi towards the end a little blackish. Cilia of the tegulae black. Wings tinged with gray. Third and fourth longi- tudinal veins towards the end a little more approximated, arid therefore a little more converging, than in the allied species. Hob. Middle States. (Osten-Sacken.) 15. G. difficilis LOEW. $ . — Viridis, antennis nigris, facie et fronte albido-pollinosis ; pedibus flavis ; tarsis apicem versus dilute infuscatis, alarum venis longitudinalibus tertia et quarta perfecte parallelis ; lamellis hypopygii pallide flavescentibus. Green, antennae black, face and front covered with a whitish dust, feet yellow, tarsi brownish only towards the tip, third and fourth longitudi- nal veins perfectly parallel ; lamellae of the hypopygium pale-yellowish. Long. corp. 0.12. Long. .al. 0.12. SYN. Gymnopternus difficilis LOEW, Neue Beitr. VIII, 33, 14. Is so extremely like the preceding species, that the statement of the differences will be sufficient for its recognition. They consist in the following : the usual row of bristles on the upper side of the fore tibias consists of much smaller bristles. The hairs on the upper side of the hind tibiae are, towards their end, less dense and long ; the third and fourth longitudinal veins are perfectly parallel towards their end, and all the longitudinal veins have a paler coloring. Hab. New York. 16. G. nigritoarbus LOEW. $ . — Nigro-seneus, thorace cserulescente, subopaco, inferS, faciei parte pilis nigris barbata. Bronze-black, thorax rather blue and somewhat dull, the lower part of the face bearded with black hairs. Long. corp. 0.09 — 0.10. Long. al. 0.12. SYN. Gymnopternus nigribarbus LOEW, Neue Beitr. VIII,. 33, 15. Bronze-blackish, thorax rather blue, and rather dull on account of a dense, brown-gray dust. Face covered with a whitish-gray dust, rather broad, upon its inferior portion convex and beset with 92 DIPTERA OF NORTH AMERICA. [PART II. a short but distinct and rather striking pubescence. Antennas small, black-brown, second joint and the root of the third red ; the third joint rounded at the end and beset with very distinct but not long hairs. Arista with a comparatively long, very dis- tinct pubescence. The front seems in most directions light brownish-gray ; in others nearly whitish. Cilia of the inferior orbit black. Upper side of the thorax rather blue and dull on account of a gray-brownish dust. Scutellum apparently glabrous upon its upper surface. The color of the abdomen varies between bronze-black and bronze-green. Fore coxa? yellow-brownish at the base ; middle coxae almost up to the tip, hind coxas about as far as the middle, blackish. Feet yellowish. Tarsi moderately infuscated towards the tip. The usual row of bristles on the upper side of the fore tibise, consists of comparatively short bristles. Cilia of the tegulse black. Wings with a rather strong blockish-gray tinge. The third and fourth longitudinal veins parallel towards the end. Hob. Pennsylvania. 17. G. parvicorsiis LOEW. $ . — Viridis, thorace cserulescente, niti- dissimo, antennis parrls, nigris, articulo secundo et articuli tertii basi rufis ; coxis anticis totis, pedibusque pallide flavis, tarsis apicem versus infuscatis, lainellis hypopygii pallide flavicantibus, appeudicibus inte- rioribus simplicibus. Green, the bluish-green thorax very bright ; the small antennae black, the second joint and the root of the third red, the whole fore coxae and the feet pale yellowish ; tarsi towards the tip brownish ; the lamellae of the hypopygium pale yellowish ; the interior appendages plain. Long, corp. 0.12. Long. al. 0.12. SYN. Gymnopternus parvicornis LOEW, Neue Beitr. VIII, 34, 16. Green, bright, face and front with a white dust. Antennas small, black ; second joint and the root of the third red ; third joint remarkably small and not rounded at the tip. Arista with a short but distinct pubescence. Cilia of the inferior orbit black. Upper side of the thorax bluish-green and very bright. Upon the surface of the scutellum there seem to be, besides the usual bristles, a few small hairs. The small lamellae of the hypopygium are whitish-yellow with a scarcely perceptible black margin, cres- cent-shaped. Coxse and feet whitish-yellow ; the middle coxas on the outside almost up to the tip, and the hind coxae at the root, QYMNOPTERNUS. 93 blackish. The hairs on the feet are scarcely so rough as usual, and the row of bristles on the upper side of the fore tibiae consists of rather short bristles. Tarsi but slightly infuscated towards their tip. Cilia of the tegulae black. Wings with a blackish-gray tinge. Third and fourth longitudinal veins, towards the end, parallel. Hob. Middle States. (Osten-Sacken.) Observation. — The striking resemblance of this species to G. laevigatus has already been noticed above. If the small bristles on the upper side of the fore tibia? were not visibly shorter in this species, I would suppose it to be only a variety of G. laevigatus with much paler tarsi. IS. G> opactis LOEW. % . — Viridis, modice nitens, facie et fronte polline ex albo-cinereo vestitis, aiitemris majusculis, fusco-nigris, arti- culo secundo et articuli tertii basi rufis ; pedibus cum coxis flavis ; ooxis intermediis, posticarumque basi nigricantibus ; alis ex flavo dilutissiine cinerascentibus, lamellis hypopygii parvis, pallide flavescentibus. Green, only moderately shining ; face and front with a whitish-gray dust ; antennae rather large, brownish-black; second joint- and root of the third red ; coxae and feet yellow, middle coxae and the base of the hind cox33 blackish ; wings altogether pale yellowish-gray ; the small lamellae of the hypopygiurn pale yellowish. Long. corp. 0.12 — 0.13. Long. al. 0.13. Sra. Gymnopternus opacus LOEW, Neue Beitr. VIII, 34, 17. Green, only moderately shining ; the dust upon the face seems to be whitish-gray, but in an oblique light it has a more pure white appearance. Antennas brownish-black ; second joint and root of the third red; third joint of a considerable size, rather broad, forming a sharp angle at the tip, beset with not very long but very distinct hairs. Arista with a rather short but very dis- tinct pubescence. Front with a yellowish-gray dust. Cilia of the inferior orbit black. Thorax dull On account of a yellowish- gray dust. I cannot discover any hairs upon the scutellum. La- mellae of the hypopygium small, pale-yellow, short and sparsely fringed. Pleurae without yellow coloring on the posterior margin. Cox33 and feet pale-yellowish. Middle coxa? on the outside almost up to the tip and hind coxa? at the basis, of a dark color. Tarsi hardly infuscated towards the tip. Cilia of the tegula? black. Wings with a slight gray-yellowish tinge. The third 94 DIPTERA OP NORTH AMERICA. [PART II. and fourth longitudinal veins towards their end almost entirely parallel. Hob. New York. (Osten-Sacken.) Observation. — A single ? which I possess I believe to be that of the present species. The circumstance that the single bristles of the row on the upper side of the fore tibia? are some- what stronger, the third joint of the antenna? much shorter and the arista more distinctly hairy than those of the above described g , cannot justify any doubts, as the females of nearly all the species differ in this way from the males. The only objection which might be raised against their belonging together, is the more whitish color of the dust upon face and front. 19. G-. politus LOEW. 9 • — Viridis, nitens, faciei albse parte infera subtiliter pilosa, fronte albo-pollinosa, antennis ex fusco rufis, apicem versus fuscis, coxis pedibusque pallide flavis, alis majusculis ex fusco cinereis, appendicibus analibus duabus styliformibus. Green, bright, the lower part of the white face with delicate hairs, front with a white dust, the brownish-red antennae brown at the tip ; coxae and feet pale-yellowish ; wings somewhat large, brown-grayish ; at the end of the abdomen two styloid appendages. Long. corp. 0.14 — 0.15. Long. al. 0.14—0.15. STN. Gymnopternus politus LOEW, Neue Beitr. VIII, 34, 18. Green, very bright ; the moderately broad face and front covered with white dust ; the lowest part of the face beset with minute pale hairs and a few blackish ones. Antenna?, at least for a female, of middle size, dusky brownish-red ; third joint with short but dis- tinct hairs, towards the end blackish-brown and the tip sharply angular. Cilia of the inferior orbit black. Thorax only d little dusty. Upon the scutejlum I cannot perceive any hairs at all. The anal appendages distinguish themselves from those of the related species by consisting of two short black styles. The pos- terior margin -of the pleura? is not yellow. Coxa? and feet pale- yellowish. Tarsi but little infuscated towards their tip. The usual row of bristles on the upper side of the fore tibia? rather prominent. Cilia of the tegula? black. Wings rather broad and pretty strongly tinged with brownish-gray. The third and fourth longitudinal veins towards their ends with a slight indication of convergency. Hob. New York. (Osteu-Sacken.) GYMNOPTERNUS. 95 Observation. — It is remarkable that the circle of short thorns on the tip of the abdomen of the female, which belongs to allied spe- cies, is wanting here. This species, however, cannot be located in any other genus ; on the contrary, it coincides most perfectly in all other respects with the species of Gymnopternus. 2O. G. detoilis LOEW. £ and 9 . — Viridis, modice nitens, facie et fronte cinereo-pollinosis, antennis rufis apicem versus nigris, coxis pedi- busque pallide flavis, alls ex flavo ciuereis, laraellis hypopygii parvis, pallide flavescentibus. Green, moderately shining ; face and front grayish-dusty, the red antennae black at the tip ; coxae and feet pale-yellow ; wings yellowish-gray ; the small lamellae of the hypopygium pale-yellowish. Long. corp. 0.12. Long. al. 0.12. STN. Gymnopternus debilis LOEW, Neue Beitr. VIII, 35, 19. Green, only moderately shining ; face covered with white-gray- ish, front with yellow-grayish dust. Antennae red, of middle size ; third joint quite distinctly hairy, at the tip black-brown and pro- vided with a sharp angle ; arista with a short, but distinct pubes- cence. As far as I can perceive, the ciliaa of the inferior orbit are black. The upper side of the thorax, on account of some yellow-grayish dust, rather dull. The scutellum seems to be bare. Yenter not yellow. Pleurae wholly gray, without yellow posterior margin. Coxa? and feet pale yellowish. Tarsi towards their end a little infuscated. The usual row of bristles on the upper side of the fore tibiae rather prominent. Cilia of the tegu- laa black. Wings only with a slight yellow-grayish tinge. Third and fourth longitudinal veins towards their end almost entirely parallel. The small lamellce of the hypopygium are pale-yellow- ish and fringed with little black hairs. Hob. Pennsylvania. (Osten-Sacken.) Observation. — G. debilis can easily be distinguished from G. opacus by its smaller size and smaller antenna? ; likewise from G. crassicauda by not having a yellow venter and the pos- terior margin of the pleurie not being yellow. From G. politus it differs by a more light-green color, less brightness, smaller size, &c. &c. G. crassicauda LOEW. £ and 9 •— Viridis, modice nitens, an- tennis rufis apicem versus fuscis, pleurarum margine postico, ventre, coxis pedibusque pallide flavis, hypopygio maris valde incrassato. 96 DIPTERA OP NORTH AMERICA. [PART II. Green, moderately shining ; the red antennae brown towards the tip ; the posterior margin of the pleurae, the venter, the 00x33 and the feet pale- yellowish. The hypopygium of the $ very much thickened. Long. corp. 0.15. Long. al. 0.14 — 0.15. SYN. Gymnopternus crassicauda LOEW, Neue Beitr. VIII, 35, 20. Green, moderately shining ; face and front with some whitish dust ; in well preserved 9 a delicate and pale-colored pubescence can be seen upon the lower part of the face. Antennas red, of very moderate size, the third joint at the end rounded and infus- cated. Arista of the J* with a short but distinct, that of the 9 with a comparatively long and very striking pubescence. Cilia of the inferior orbit black. Thorax somewhat dull from grayish dust. The hairs upon the scutellum delicate and rather diffi- cult to perceive. Pleurae gray, their whole posterior margin (epimera metathoracica) yellowish. Tenter yellow ; -upon the anterior segments of the abdomen this color extends somewhat upon the upper side ; in well preserved specimens, however, it is concealed by a whitish dust ; in the 9 this yellow coloring some- times extends further, so that there is upon the first segment a complete, and upon the second an interrupted yellow band. Coxa? and feet pale-yellowish. Tarsi towards the end scarcely a little infuscated. Cilia of the tegulse black. Wings with a slight yel- low-grayish tinge. The third and fourth longitudinal veins to- wards the end almost entirely parallel. The hypopygium of the g uncommonly thickened. The very small crescent-shaped lamellae have a yellowish coloring and a fringe of short, delicate and sparse hairs. Hab. New York. (Osten-Sacken.) . G. minuf us LOEW. £ . — Viridis, antennis parvis rufis, pleurarum margine postico, ventre, coxis pedibusque pallide flavicantibus, coxis anticis denudatis, hypopygio non incrassato. Green, the small antennas red, the posterior margin of the pleura, the ven- ter, the coxse and the feet pale-yellowish ; fore coxae bare ; the hy- popygium not thickened. Long. cprp. 0.10. Long. al. 0.11. SYN. Gymnopternus minutus LOEW, Neue Beitr. VIII, 35, 21. Green, quite bright, face with a dense, front with a thin whitish dust. Antennae brownish-red, small, the third joint at the end dark-brown. Arista with a short but distinct pubescence. Cilia of the inferior orbit black. Thorax a little dull from a white- PARACLIUS. 97 grayish dust. Scutellum with a few short hairs. Posterior mar- gin of the pleurae yellow. Tenter yellow. On the anterior seg- . ments of the abdomen the lateral margins are also colored with yellow. Hypopygium of the usual shape. The small, delicate lamellae yellowish, sparsely ciliated. Coxae and feet white-yellow- ish. The fore coxae have upon their anterior side no black, but throughout only extremely delicate small whitish hairs, so as to appear glabrous, which constitutes a very striking character of this species. The tarsi towards their end are scarcely somewhat infuscated. Hob. Middle States. (Osten-Sacken.) 23. G. ventralis LOEW. $ . — Viridis, modice nitens, antennarum articulo tertio latiusculo, rotundato, pleurarum margine postico, ventre, coxis, pedibusqu'e pallide flavis, hypopygio maris non incrassato. Green, moderately shining, third joint of the antennae rather broad, rounded ; posterior margin of the pleurae, venter, coxae and feet pale-yellowish ; hypopygiurn not thickened. Long. corp. 0.13 — 0.14. Long. al. 0.15. SYN. Gymnopternus ventralis LOEW, Neue Beitr. VIII, 36, 22. Green, not very bright. Face with a whitish, front with a pale yellow-grayish dust. -Antennae brownish-red, of moderate size ; the third joint, which is rounded, is rather broad ; arista with a rather short but distinct pubescence. Cilia of the inferior orbit black. Thorax quite dull on account of a yellow-grayish dust. Scutellum with delicate but distinct hairs. The entire pos- terior margin of the pleurae yellow. Tenter yellow. Hypopygium of the usual form. The small yellowish lamellae of middle size, rather sparsely ciliated. Coxae and feet pale-yellowish. The hairs on the anterior coxae are partially blackish, but so delicate that they might be easily overlooked. The tarsi towards their end are only little infuscated. Cilia of the tegulae black. Wings with a yellow-grayish tinge ; the third and fourth longitudinal veins parallel towards their end. Hob. New York. Gen. IV. PARACLIUS. » When I adopted, in the fifth part of the Neue Beitrage, the genus Gymnopternus, and distinguished it from the related genera, it was done merely upon the basis of an investigation of European species, so that I had only these species -in view when 98 DIPTERA OP NORTH AMERICA. fPART II. I defined the characters of this genus and its differences from the neighboring genera. America possesses species which necessarily come within the definition of the genus Gymnopternus, as under- stood in that publication, but which, at the same time, differ too much from all other species of this genus, to find a natural place among them. The most striking, although perhaps not the most important, character whereby these species differ from the others, is the course of the last segment of the fourth longitudinal vein. At or beyond its middle it is suddenly deflected anteriorly, and its end is so near the end of the third longitudinal vein that the first posterior cell appears almost closed. In order to separate these species from the genus Gymnopternus, I have added above to the characters of Gymnopternus the complete, or at least nearly com- plete, parallelism of the third and fourth longitudinal veins. A more minute examination of the species in question shows that they should form two, or perhaps more correctly, three groups ; still, before we are able to judge with certainty about it, our as yet im- perfect knowledge of the species will require a considerable in- crease. In the meantime, however, if we draw our attention to the character which distinguishes all these species from the other Gymnopternus, that is, to the course of the last segment of the fourth longitudinal vein, we will soon find among these species two principal modifications of this course. In one case the deflec- tion of the fourth segment at or beyond its middle takes place in a steep curve forward, and the vein then runs in a straight direc- tion to the margin of the wing, which it reaches very near the tip of the third longitudinal vein. In the other case the last segment of the fourth longitudinal vein forms beyond its middle a but slightly rounded angle, and thence, in the form of a curve, the concavity of which is turned backwards, it runs to the margin of the wing, which it likewise reaches in the immediate neighborhood of the third longitudinal vein. Those species which show the first of the above mentioned neurations, possess, moreover, many other characters in common, which distinguish them from the species of Gymnopternus, and thus they form the genus Pelastoneurus. As the most important of these characters may be mentioned the feathered arista, the broad face, which is common to both sexes, strongly convex upon its lower part, and provided with a sharp, curved inferior margin ; also the elongated and distinctly peduncu- lated hypopygium. On the contrary all those species, in which the PARACLIUS. 99 end of the fourth longitudinal vein forms a curve, the concavity of which is turned backwards, and which in this respect differ more than the others from the species of Gymnopternus, approach them very closely in the structure of the face, and are easily distinguished on that account from the species of Pelastoneurus ; the structure of their face would be indeed altogether like that of the species of Gymnopternus, if the face was not somewhat less broad and more narrowed below. The structure of their hypopygium also ap- proaches more to that of the species of Gymnopternus, than to Pelastoneurus, the hypopygium not being elongated and being provided with a shorter peduncle, so as to appear sessile or almost sessile. The outer appendages of all the species have more of the usual form of a shell than those of the species of Pelastoneurus. While there is a great uniformity with regard to all the above men- tioned characters, this is not the case with the structure of the antennae. In both species, which I describe below, the third joint of the antennae is rounded and the arista more or less distinctly hairy, but not feathered. In the species from Surinam, which I have described in the Wiener Entomol. Monatschr. as Gymnop- ternus leucospilus, the third joint of the antennae is longer, quite distinctly excised on its upper side, and has, like the species of Pe- lastoneurus, a distinctly feathered arista. Whether this difference in the structure of the antennas is sufficient to form two genera of these species, I am unable to decide, on account of the scanty ma- terial at my disposal, but I rather incline to that opinion. Doli- chopus heteropterus Macq. undoubtedly belongs to this group, but whether it is more related to the two species described below, or to G. leucospilus, cannot be decided without the comparison of the specimen ; however, according to the statements and the drawing which Mr. Macquart furnishes of the structure of the antennae, the former seems to be the case. Mr. Bigot founded upon it a separate genus, which he calls Paracleius, and distinguishes it from the genus Gymnopternus by the latter having the third joint of the male antennae slightly excised on the upper side, and the fourth longitudinal vein bent, while in Paracleius the third joi-nt of the male antennae is not excised on the upper side, and the fourth longitudinal vein is strongly bent. My experience does not, as yet, allow me to agree with this mode of subdivision ; I therefore cannot adopt the genus Paracleius in the sense of Mr. Bi^ot. Nevertheless, I see no inconvenience in retaining the 100 DIPTERA OF NORTH AMERICA. [PART II. newly coined name, with the usual latinized modification in Para- clius, for the new genus which I intend to establish and to define here. As our principal aim at present is an available generic distribution of the already known North American species, I will merely have the latter in view in establishing the characters of Paraclius, and leave out G. leucospilus for the present. The discovery of a larger number of related species will have to decide whether the character of Paraclius is to be modified so as to admit species like G. leucospilus, or whether a new genus is to be founded for such species. The following are the characters of the genus : The first joint of the antennas hairy on the upper side ; third joint of the antennae rounded ; arista dorsal, with the ordinary pubescence, not feathered. Face of very moderate breadth, narrowed towards the bottom, not convex in its lower part and not reaching to the inferior corner of the eye. The first joint of the hind tarsi without bristles. The last segment of the fourth longitudinal vein, beyond its middle, is bent forward in a rounded angle, thence running in a curve, with the concave side turned backwards, towards the margin of the wing, and reaching it quite near the tip of the third longitudinal vein, so that the first posterior cell has but a small opening. Hy- popygium entirely disengaged, not prolonged, with a very short pedicel, so as to appear sessile or nearly so ; the exterior append- ages lamelliforin. The characteristic differences between Paraclius on one side and Gymnopternus and Pelastoneurus on the other, will be easily understood from the foregoing. Besides the species of the latter two genera, there, are those of the genus Hercostomus, which re- semble the species of Paraclius; but in this genus the last seg- ment of the fourth longitudinal vein only very gradually approaches the third longitudinal vein, without any vestige of an angular flexure, and reaches the margin of the wing not so near the third longitudinal vein. Only American species of -Paraclius are as yet known. The name of the genus (from xapa — xheiu, I close), means that the first posterior cell of the species is almost closed. PARACLIUS. 101 Table for the determination of the Species. Arista with long hairs ; first segment of the costa not swollen. 1 arcuatus Lw. Arista with short hairs ; first segment of the costa strongly swollen. 2 albonotatus, n. sp. Description of the Species. 1. P. arcuatus LOEW. $> . — Obscure viridis, pedibus flavis, ex parte fuscis, alis nigricantibus, primo costae segmento non incrassato. Dark green, feet yellow, partially dark-brown, wings blackish, first seg- ment of the costa not thickened. Long. corp. 0.12. Long. al. 0.11. SYN. Pelastoneurus arcuatus LOEW, Neue Beitr. VIII, 39, 4. Green, bright. Face narrow, still narrower below, covered with thick white dust. Front green, rather dull from a whitish dust. Antennas comparatively small, black, the third joint rounded. Arista with comparatively long hairs, but not feathered. Cilia of the inferior orbit whitish. The upper side of the thorax is upon the first two-thirds of a dark bronze color and less bright, upon the last third of a magnificent golden green color and very bright ; the triangular impression on each side near the transverse suture is thickly covered with white dust ; a small spot in the vi- cinity of the posterior corner of the thorax is dusted in a similar manner. In looking at the thorax from behind, a deep black stripe-like double spot above the root of the wing becomes appa- rent. Scutellum rather bright, of copperish color with a green middle line. The extreme tips of its lateral corners are deep black, and the hairs on its upper side are particularly distinct. Abdomen bright, rather dark green, along the incisures blacker, the lateral margins of the single segments with not very distinct spots of whitish dust. Pleura greenish-black, and rather gray from a thin whitish dust. Fore coxse yellowish-brown, towards the tip lighter ; middle and hind coxae as far as the tip black. Feet brownish-yellow ; fore and middle femora on the upper side brownish, hind femora rather dark brown upon their whole latter part. Hind tibisB, with the exception of the root, dark brown. Fore and middle tarsi from the tip of the first joint dark brown ; the whole hind tarsi blackish-brown. Cilia of the tegulse black. Wings blackened, towards the anterior margin darker ; the last segment of -the fourth longitudinal vein is suddenly bent forward 102 DIPTERA OF NORTH AMERICA. [PART II. almost at a right angle, and this segment forms a curve, the con- vexity of which is turned backwards. Hob. Cuba. (Poey.) 2. P. altoonotatus, n. sp. £ and 9- — Obscure viridis, pedibus totis nigris, alls nigricantibus, primo costae dimidio valde incrassato. Dark green, feet entirely black, wings blackish, the first segment of the costa very much thickened. Long. corp. 0.17 — 0.18. Long. al. 0.17. Dark green, sometimes more bronze-green. Face of the £ narrow, of the 9 a little broader ; in both sexes it is covered with a snow-white dust. Palpi brownish-black. Antennas entirely black, of very moderate size, the third joint rounded ; arista with the usual short pubescence. Front covered with a rather dense white dust. Cilia of the inferior orbit whitish. Thorax dark me- tallic green, sometimes, with the exception of the posterior part of the upper side, of a dusky bronze-color. The pleurse and the triangular lateral impression on the transverse suture are covered with a bright white dust. Scutellum of the same color as the upper side of the thorax. Abdomen on the posterior margin of the single segments usually rather blackish-blue-green, upon t^e remaining part of the segments more golden-green or coppery ; on the lateral margin covered with white dust. Hypopygium disengaged, sessile, greenish-black ; lamella only of very moderate size, rounded, brownish-black. Coxae and feet black, the former covered with black hairs, the latter with a greenish lustre ; fore tibiae only with a simple row of bristles ; middle and hind tibiae with numerous bristles. Tegulae blackish-brown, with black cilia. Wings of moderate size, of an elongated-oval shape, blackened ; the last segment of the fourth longitudinal vein about its middle is bent forward at an obtuse angle, and its tip, which reaches the margin quite near the third longitudinal vein, forms a curve, the concavity of which is turned backwards. In the $ the portion of the costa which lies before the end of the first longitudinal v^in shows a very strong swelling ; in the 9 this swelling is much weaker, but still of a rather conspicuous size. Hab. New Orleans. PELASTONEURUS. 103 Gen. V. PELASTONEIJRIJS. The characters of the genus are the following: First joint of the antennas short, hairy on the upper side ; third joint rounded ; arista dorsal, distinctly feathered. Face in both sexes compara- tively broad, upon its lower part strongly convex; its lower margin is sharp, and forms a curve. Proboscis stouter than in Gymnopternus, and approaching in its structure the species of Medeterus. The first joint of the hind tarsi without bristles. The last segment of the fourth longitudinal vein turns forward at or beyond its middle in a strong curve, and runs then almost in a straight line towards the margin of the wing, which it reaches closely in the vicinity of the tip of the third longitudinal vein, so that the first posterior cell is almost closed. The hypopygiurn is entirely disengaged, very much elongated, pedunculated, with ^lainelliform black appendages, which, in most of the species, have a very elongated form, and are of a more tough substance than in the allied genera. The next related genus is Paraclius. The differences of both have already been detailed above. Pelastoneurus can hardly be mistaken for any other genus. ' As yet, only American species of Pelastoneurus have been made known. Among the species described by former authors, Dolichopus maculipes Walk., andi>. bifrons Walk., seem to be- long here. The name of the genus (from *f?ia!w, I approach, and vevpov, the nerve) has reference to the position and the peculiar course of the last segment of the fourth longitudinal vein. Table for the determination of the Species. , ( Cilia of the inferior orbit black. 2 1 Cilia of the inferior orbit pale. 4 , Thorax with a large spot of white dust on the posterior margin. 2-1 1 longicauda Lw. ( Thorax without a spot of white dust on the posterior margin. 3 o ( Wings blackened. 2 lugubris Lw. \ Wings gray. 3 laetus Lw. . ( Fore coxae blackened at the basis. 4 vagans Lw. Fore coxae not blackened at the basis. 5 cognatus Lw. •• 104 DIPTERA OP NORTH AMERICA. [PART II. Systematic arrangement of the Species. I. Cilia of the inferior orbit black. 1. longicauda Lw. 3. lastus Lw. 2. lugubris Lw. II. Cilia of the inferior orbit pale. 4. vagans Lw. 5. cognatus Lw. Description of the Species. I. ClLIA OP THE INFERIOR ORBIT BLACK. 1. P. longicauda LOEW. £. — Aeneo-niger, facie argenteo micante, sub antennis triangulum nigrum gerente, ciliis oculorum inferioribus nigris. Bronze-black ; face with a silvery lustre, with a black triangular spot under the antennae ; cilia of the inferior orbit black. Long. corp. 0.17. Long. al. 0.16. SYN. Pelastoneurus longicauda LOEW, Neue Beitr. VIII, 37, 1. Face, for a £ , extremely l?road, the inferior two thirds of it are strongly convex and have a bright silvery-white reflection, which shows a somewhat olive-brown appearance only in a certain ob- lique light ; the upper, flat portion of the face has in each lower corner a deep, triangular spot with a silvery lustre ; that triangu- lar part of it, which is not covered by this spot, appears deep- black, when seen from above ; seen from below, it appears less dark and somewhat dusty. The lower margin of the fac,e is very sharp. Palpi large, on the outside with a silvery-white lustre and covered with black hairs. Front shining blackish. Antennae brownish black ; the under side of the first and second joints brownish-red ; the rounded third joint rather large. Arista rather short, very much thinner towards the tip, and upon the last two thirds with short feathery hairs. Cilia of the inferior orbit black. Thorax bronze-black, rather shining, with an almost imper- ceptible white dust ; the upper side of the thorax shows five spots covered with snow-white dust, namely one on each side near the transverse suture, one in the shape of a dot, on each side above the root of the wing near the posterior corner, and finally a large triangular spot in the middle of the hind margin ; the velvet-black stripe-like double spot immediately above the root of the wing, so common in the species of this genus, is very distinct here. PELASTONEURUS. 105 Coxae and pleurae black, with a silvery lustre. Scutellum ' with velvet-black lateral spots and with a velvet-black middle-stripe, smooth steel-blue between the corners and the middle stripe. Ab- domen with a violet lustre. Hypopygiuru black, upon the under side covered with snow-white dust, pedunculated, not very stout, but very long, so that it reaches as far as the basis of the abdo- men ; the comparatively small lamellae brownish-black ; the inte- rior appendages slender, black, provided at the tip with not very numerous but long hairs in the shape of a brush. Feet brownish- yellow, hind femora blackened on the upper side of the extreme tip ; the bristles on the upper side of the tibiae are inserted upon irregular small black-brown spots ; tarsi dark brown, paler at the basis. Cilia of the tegulae black. Wings comparatively small and narrow, tinged with blackish-gray and darker towards the end of the anterior margin. Hob. New York. (Osten-Sackeu.) Observation. — The punctation of the tibiae may remind one of Dolichopus maculipes Walk. This species is described as only one and a half lines long and bronze-green, and with all the femora having black tips. No mention whatever is made in Mr. Walker's description of the very peculiar structure of the face and of the striking white spots on the thorax, which are peculiar to Pelastoneurus longicauda. Under such circumstances the identification of these two species is impossible, the more so as all the species of Pelastoneurus are very much alike and as the spots on the tibiae are a character which frequently occurs in this genus. 2. P. lugllbris LOEW. 9 . — Niger, tliorace fusco-pollinoso, opaco, ciliis oculorutn inferioribus nigris, alls nigricantibus. Black, thorax covered with, brown dust, dull ; cilia of the inferior orbit black ; wings blackish. Long. corp. 0.11. Long. al. 0.10. SYN. Pelastoneurus lugubris LOEW, Netie Beitr. VIII, 38, 2. Black. Face black, with a whitish, not shining dust, and with a blackish-brown, not well defined middle stripe ; upon the larger, inferior part but moderately convex. Palpi rather large, black, on the outside with a thin whitish dust and black hairs. Front dull, brownish-black. Antennae reddish-brown, the basis of the first and the larger part of the third joint brownish-black ; the third joint is small and rather rounded, beset with short but distinct 106 D1PTERA OF NORTH AMERICA. [PART II. hairs. Arista towards the tip with short feathery hairs. Cilia of the inferior orbit black. The spot on each side near the transverse suture of the thorax is covered with white dust, however this dust is not distinctly seen in every direction ; the usual deep-black stripe-shaped double spot immediately above the root of the wing becomes distinctly visible, when looking at the thorax from be- hind ; likewise the usual little white spot in the neighborhood of the hind corner is seldom distinctly seen and is always very small. On the posterior margin of the thorax there is no spot with white dust. Scutelluin bluish-black with velvet-black lateral corners. In well preserved specimens there is a middle-stripe with grayish- white dust. Pleurse black, gray on account of a thin whitish dust. Abdomen bronze-black, each segment on the lateral margin with a small spot covered with white dust and not visible in every direction. Fore coxae brownish-yellow, with an almost imper- ceptible, very thin covering of white dust. Middle and hind coxas black. The color of the feet is rather variable ; usually they are yellowish-brown, the upper side of the anterior femora, the tip of the hind femora, likewise all the tibias and tarsi black-brown ; nevertheless there are specimens in which they are more of a brownish-yellow color and where the tip of the femora and the tarsi, with the exception of their roots, are blackish-brown, while the upper side of the tibise is indistinctly spotted in consequence of the brownish color of the places of insertion of the bristles. Cilia of the tegulse black. Halteres blackish. Wings rather small, narrowed towards the basis, distinctly blackened, darker towards the anterior margin ; in more faded specimens dark mar- gins appear around the veins, as is also the case in the other species of the same genus. Hob. Trenton Falls, N. Y. (Osten-Sacken.) 3. P. laetus LOEW. % . — Viridis, ciliis oculorum inferioribus nigris, fronte et dimidio thoracis posteriore violaceis, alis ex fusco cinereis. Green, cilia of the inferior orbit black ; front and hind part of the thorax violet ; wings brownish-gray. Long. corp. 0.12 — 0.13. Long. al. 0.12 —0.13. SYN. Pelastoneurus Isetus LOEW, Neue Beitr. VIII, 38, 3. Dark-green, rather shining. Face considerably broad and covered with a dense snow-white dust, the inferior third convex. PELASTONEURUS. 107 Palpi of moderate size and yellowish color, covered on the outside with snow-white dust. Antennae yellowish-red, the third joint a little longer than broad, at the tip altogether rounded, its apical half blackish-brown ; arista towards the tip with short feathery hairs. Cilia of the inferior orbit black. Front violet, sometimes almost steel-blue. Upper side of the thorax green with a thin gray-brownish dust, upon the larger portion of the posterior part violet ; the spot on each side of the suture is covered with white dust ; the usual deep black, stripe-shaped double spot, immedi- ately above the root of the wing is very distinct ; the small dot of white dust in the vicinity of the hind corner, however, is seldom distinctly visible ; on the posterior margin of the thorax there is no spot of white dust. Scutellum shining black-green with deep- black lateral corners ; only in faultless specimens there is a middle stripe of white-grayish dust, surrounded by a more black color- ing. Pleurae black, gray on account of whitish dust. Each seg- ment of the abdomen has on the lateral margin a spot of white dust, which is not very sharply defined, and the sixth, small seg- ment, is entirely covered with whitish dust. Hypopygiuni shortly pedunculated, greenish-black ; on the under side gray from pale dust ; it reaches with its tip as far as the middle of the abdomen ; the long brownish-black lamellae are narrow, at the end gently bent upwards, reaching the basis of the abdomen ; the slender interior appendages are also blackish-brown, beset at the tip with a few long hairs. Fore coxa? yellowish, middle and hind coxa? blackish almost as far as the tip. Feet pale-yellowish ; the tip of the hind femora is not of a dark color, and the bristles on the upper side of the tibia? are not inserted on dark spots ; middle and hind tarsi, with the exception of the roots, black-brown ; fore tarsi brown only at the tip. Cilia of the tegulae black. Wings with a more brownish-gray than blackish-gray tinge and darker towards the anterior margin. Hob. Georgia; District Columbia. (Osten-Sacken.) II. ClLIA OP THE INFERIOR ORBIT PALE. 4. P. vagans LOEW. % and 9 • — Obscure viridis vel nigro-seneus ; antenuarum basi rufa, ciliis oculorum inferioribus pallidis, coxis anticis, excepto apice, nigris, alls cinereis. Dark-green or bronze-black ; the root of the antennae red ; cilia of the in- 108 DIPTERA OF NORTH AMERICA. [PART II. ferior orbit pale ; fore coxae with the exception of the tip black ; wings gray. Long. corp. 0.15 — 0.16. Long. al. 0.15. SYN. Pelastoneurus vagans LOEW, Neue Beitr. VIII, 39, 5. Dark-green or blackish bronze-colored, moderately shining. Face broad, with a snow-white dust, in the 9 with a broad gray- brownish middle stripe, which is wanting in the £ ; its lower part convex. Palpi rather large, blackish, yellowish at the tip, on the outside covered with a dense snow-white dust and black hairs. Front covered with a brown dust, seldom entirely concealing the ground color, which is steel-blue, except in the vicinity of the upper corners where it is violet. Antennae not very long, the third joint, however, which is rounded and distinctly hairy, is rather large ; their color is red ; the upper side of the first and the greater part of the third joint are black-brown ; sometimes the upper side of the second joint has the same color. Arista with rather long feathery hairs. Cilia of the inferior orbit whitish. The color of the upper side of the thorax, in recently excluded specimens, is more green, and shows then two longitudinal lines of a violet color, which increase in breadth backwards and become visibly divergent; in more faded specimens this color is more dark bronze-black, and of the two violet longitudinal lines only the hind part is often perceptible, which then becomes more extended. The spot of white dust on each side, near the suture, and the usual deep-black double spot immediately above the root of the wing, are very striking ; upon the hind corner, which is of a brighter green color, there is a spot of white dust in a diagonal direction ; how- ever, it is very difficult to perceive. The extreme tip of the lateral corner of the scutellum appears black, the elevated middle stripe is usually green, and the slight depression on each side of the latter more bronze-colored. I cannot perceive any hairs upon the sur- face of the scutellum. Abdomen bronze-green, often somewhat copper-colored ; the white dust in the vicinity of the lateral mar- gin does not form any distinct spots. Coxoe black with a bright snow-white lustre ; fore coxae yellow at the tip to a rather large extent, the middle and hind ones only to a very small extent. Feet somewhat brownish-yellow ; fore tarsi only at the tip, middle and hind tarsi from the tip of the first joint, blackish-brown. Cilia of the teguloa black. Wings in recently developed specimens slightly tinged with gray, in faded ones visibly darker. The TACHYTRECHTJS. 109 elongated and slender hypopygium is pedunculated and of a black color ; the long and narrow lamellce are of an equal breadth, black, fringed with rather long black hairs ; the interior appendages small, without hairs at the tip. Hob. Middle States. (Osten-Sacken.) 5. P. coguatlis LOEW. 9- — Obscure seneus, antennarum rufarum apice fusco, ciliis oculorum inferioribus pallidis, coxis anticis totis flavis. Dark bronze-colored ; the tip of the red antennae brown ; cilia of the infe- rior orbit pale ; fore coxae entirely pale. Long. corp. 0.12 — 0.13. Long, al. 0.10. Of this species I know only the 9 , which is a little larger than that of the preceding species, and is distinguished from it by the paler antennae, by the feathery hairs of the arista being a little longer, by the depression on each side of the suture, of the thorax which is dusted with white only in the interior corner, and by the entirely yellow fore coxse. The' specific distinctness cannot be called in doubt. Hob. Middle States. (Osten-Sacken.) Gen. VI. TACHYTRE CHITS. The vertical diameter of the strongly pubescent eyes is very large, and, on that account, the head very high. The face is com- paratively narrow, becomes gradually broader towards the bottom, and reaches altogether the lower corner of the eyes. Palpi of very moderate size, also in the female. The first joint of the antennsa hairy on the upper side ; the third joint of moderate size, rounded or ovate. Arista dorsal, with an almost imperceptible microscopic pubescence or apparently bare. Cilia of the whole orbit particu- larly long. Feet rather slender ; femora not very strong ; the fore femora towards the basis a little thickened. The first joint of the hind tarsi not bristly. Wings comparatively small ; the last seg- ment of the fourth longitudinal vein converges gradually towards the third longitudinal vein, so as to reach the margin of the wing only at a moderate distance from this vein and before the tip of the wing ; upon its middle there is a more or less distinct flexure, which is sometimes more considerable in the g than in the 9 , but is never very strong. The hypopygium is entirely disengaged, with lamelliform rounded exterior appendages of moderate size. 110 DIPTERA OP NORTH AMERICA. [PART II. The habitus of the species belonging to the genus Tachytrechus is very peculiar, so that they cannot be easily mistaken or con? founded with species of another genus. It is difficult to give an adequate expression to such peculiarities of the habitus' in the characteristic of a genus. All that has been said above about the peculiar structure of the head, deserves in this respect especial attention. A particular mark, which distinguishes the genus Tachytrechus from all the other related genera is, that the face reaches as far as the inferior corner of the eye. The species of Tachytrechus known at present are found in Europe, Asia Minor, Africa and North America. The name of the genus (from taxvt, rapid, and tpeXu, I run), has reference to the habit of many species to run along sandy and muddy banks. Table for the determination of the Species. , f Antennae for the most part dark yelfow. 1 moechus Lw. I Antennre altogether black. 2 2 f Tibiae brownish-yellow almost to the tip. 2 vorax Lw. \ Tibiae black, with a greenish reflection. 3 angustipennis Lw. Systematic arrangement of the Species. I. The second joint of the antennae rudimentary. 1. moechus Lw. II. The second joint of the antennae of the usual structure. 2. vorax Lw. 3. angustipennis Lw. Description of the Species. I. THE SECOND JOINT OP THE ANTENNAE RUDIMENTARY. 1. T. moechus LOEW. $ and 9- — Viridi-aeneus, antennis maxima ex parte flavis. $ . Setae antennalis tenuissimae apice in lamellam atram dilatato, pedibus flavis. 9 • Seta antennali simplice, pedibus nigro et testaceo variegatis. Bronze-green, antennae mostly yellow. ^ . The tip of the very slender arista enlarged into a black lamella, feet yellow. J . Arista simple, feet partly black, partly brownish-yellow. Long. corp. ' 0.24—0.26. Long. al. 0.22—0.23. SYN. Tachytrechus moechus LOEW, Neue Beitr. VIII, 40, 1. TACHTTRECHUS. Ill Male. Face very long and narrow, more broad below, almost t golden-yellow, but without any lustre. Palpi small, blackish. Antenna?, in consequence of the rudimentary condition of the second joint, apparently two-jointed, as in the male of the genus Haltericerus ; the first joint elongated and somewhat swollen, of a bright dark-yellow, bare on the under side, on the upper side covered with black hairs ; the rudimental second joint of the same color ; the third joint also extremely small, rounded or somewhat kidney-shaped, brownish-black and only at the root yellow. The arista very slender, bare, half as long as the thorax and abdomen taken together ; it is black, only at the extreme tip white, and ends in a small, deep-black, rather rounded lamella, which is white at its extreme, somewhat attenuated, basis. Front metallic- green, rather without lustre, covered with brown-gray dust, which is only visible when viewed from the side. Cilia of the posterior orbit black above, yellowish below. Thorax metallic-green, usually with a more bronze or copper-colored or even violet mid- dle line ; it is rather shining, covered, however, with a distinct brownish-yellow dust. Scutellurn and abdomen have the same color and dusted covering. Hypopygium pedunculated, black, upon the lower side more black-green, and covered with yellow dust ; the yellow, rather rounded lamella? with not very long black hairs. Pleurae with a covering of thick dark-yellow dust upon bronze-green ground. Fore coxae bright yellow with almost gol- den-yellow dust, without any lustre and with some delicate sparse black little hairs. Middle and hind coxae blackish, gray on ac- count of a yellowish dust. Feet bright yellow ; middle and hind tarsi from the middle of the first joint brownish-black ; fore tarsi almost imperceptibly flattened ; in a certain direction they show a bright snow-white lustre, which reaches as far as the root upon the fore tibias. Hind femora before the tip with a single bristle. Cilia of the tegulas black. Wings tinged with gray. The tip of the fourth longitudinal vein is near the tip of the third. Female. It differs remarkably from the male in color. Face very narrow for a female, a little broader below, pale gray-yellow- ish, seldom white-grayish, and then at least upon its inferior part yellowish. Palpi small, blackish. First joint of the antennas much smaller than that of the male, less swollen, and more of a reddish-yellow color ; the second joint of the same color and less abortive than in the male ; the third joint a little larger than that 112 DIPTERA OF NORTH AMERICA. [PART II. of the male, brownish-black, reddish-yellow only on the inferior side of the basis, rounded. The black arista plain, bare, gradually thinner towards the end. Front and cilia of the posterior orbit the same as those of the male. Upper side of the thorax less green, more bronze-colored, the dust more brown, and the middle line, which is of a different color, more distinct. The abdomen is more of a bronze color ; the dust upon it, however, is rather whitish. The dust on the pleurae is also more whitish than yellow. The fore coxae are likewise blackened as far as the extreme tip ; femora green-black ; their tip to a considerable extent with a yellowish- brown tinge, which extends further on the lower side than on the upper side ; hind femora before the tip only with one bristle. Fore tibiae usually brownish-yellow, with a very thin whitish pruinose covering ; fore tarsi black, with the exception of their extreme basis. Middle and hind tibiae usually dark brown, with yellowish- brown basis and with black tip. Hind tarsi brownish-black. Cilia of the tegulae black. Wings tinged with black. Hob. Trenton Falls, K Y. (Osten-Sacken.) Observation. — A genus, based upon the present very remark- able species, would be entitled to the same claims as the genus Haltericerus, with which it nearly coincides in the structure of the antennae. I consider the establishment of such a genus as un- necessary, as this species agrees in all other respects with the already known species of Tachytrechus, which, however, are as yet not very numerous. II. THE SECOND JOINT OP THE ANTENNAE OP THE USUAL STRUCTURE. • 2. T. vorax LOEW. $ and £> . — JEneus, abdomine seneo-viridi, anten- nis pedibusque nigris, femorum apice tibiisque prseter apicem flavis. $ . Apice alarum gutta Candida niaculaque adjecta atra ornato. 9 • Alis immaculatis. Bronze-colored, abdomen bronze-green, antennae and feet black, tip of femora and the tibiae, with the exception of the tip, black. $ . Tip of the wings with a snow-white drop, and with an adjoining deep- black spot. $ . Wings spotless. Long. corp. 0.26 — 0.27. Long. al. 0.23—0.24. STN. Tachytrechus vorax LOEW, Neue Beitr. VIII, 41, 2. Face narrow, but considerably broader than that of the previous TACHYTRECHUS. 113 species, broader inferiorly, in both sexes covered with a pale yel- low dust, and without lustre. Palpi small, black. Antennae of the usual form, black. Arista in both sexes plain and bare. Front covered with a dense yellow or brown dust. Cilia of the posterior orbit above black, below white. Upper side of the thorax with a gray-yellowish or brownish-yellow dust upon a metallic-green or partially copper-colored and lustrous ground, very dull. The scutellum has a similar coloring, still its ground color can some- times be distinctly recognized. Abdomen green and coppery, dull with a gray-whitish dust. Pleurae and coxa? grayish-green, on account of a whitish dust upon green ground. Femora dark metallic-green, thinly pruinose with whitish, their tip brownish- yellow ; hind femora before their tip with a row of four bristles ; tibia? brownish-yellow ; the tip of the fore and hind tibia? blackened, the tip of the middle tibia? usually only brown. Tarsi black, plain also in the male ; the fore tarsi usually brownish-yellow only at the extreme root, the middle tarsi, however, brownish-yellow upon the first half of the first joint. Cilia of the tegula? black. Wings of the male narrow, hyaline, scarcely tinged with gray, at the tip with a small spot, the first two-thirds of which are deep-black, the last third, however, appears snow-white when seen against the light. Wings of the female not quite so narrow as those of the g , dis- tinctly tinged with gray, with a slight dark shadow around the hind transverse vein. The end of the fourth longitudinal vein in both sexes is less approximate to the end of third longitudinal vein, than in the previous species. The short peduuculated hy- popygium of the J* is black, upon the inferior side more greenish- black, but gray from a pale dust ; the lamella? are black, of mode- rate size only, rather rounded, and covered with black hairs. Hab. District Columbia. (Osten-Sacken.) 3. T. angustipennis LOEW. £ . — Viridis, antennis nigris, pedibus totis ex viridi nigris, alis immaculatis, basim versus attenuatis. Green, antennae black ; the whole feet greenish-black ; wings spotless, nar- rower towards the basis. Long. corp. 0.22. Long. al. 0.19. SYN. Taclnjtrechus angustipennis LOEW, Berl. Ent. Zeit. VI, 213, 64. Green and but little shining, on account of being rather densely covered with a fine dust. Palpi black, with a gray-yellowish dust. Face pale, ochre-yellow, dull. The rather small antenna? 8 t 114 DIPTERA OF NORTH AMERICA. [PART II. Mack. Front dull from being covered with an ochre-yellow dust. Cilia of the lateral and inferior orbit whitish.. The upper side of the thorax is covered with a grayish ochre-yellow dust, so as to make the green ground color but little apparent ; up1 on its middle there are two brown longitudinal lines, which diverge a little behind and' are very much shortened ; some portions of the usual lateral stripes are also visible, and the single bristles are inserted upon brownish-black spots. Scutellum dull, usually more brown than the upper side of the thorax. Pleurae greenish-gray. Abdomen green, covered with a rather thick whitish dust, which gives it a somewhat checkered appearance ; viewed from another point, the middle line and the posterior margins of the single segments ap- pear almost black. Hypopygiurn black, with whitish dust ; the lamellae are of moderate size, rounded oval, with short hairs, which are black on the upper and apical margin, and whitish on the lower margin. Coxae black, with yellow- whitish dust ; fore coxas beset with extremely short, delicate and sparse hairs ; be- sides, on the inner side of their basis there are a few stiif black hairs, and towards the tip a few black bristles. Feet black ; fe- mora and tibia? with a metallic-green lustre ; on the front side of the hind femora there is a single black bristle, rather distant from the tip ; very characteristic are the bristles on the upper side of the hind tibiae, which have shorter bristles in the vicinity of the basis and a longer one near the tip, otherwise of the usual shape ; upon the middle, however, there is a row of three solitary remark- ably flattened bristles. All the tarsi plain. Cilia of the tegulae black. Wings narrow, towards the basis remarkably narrowed, grayish-hyaline ; the second portion of the marginal cell more dis- tinctly dusky, the posterior transverse vein with a somewhat darker margin ; the costa distinctly thickened upon the middle of its first segment. Hob. District of Columbia. (Osten-Sacken.) Gen. VII. QRTHOCHILE. The following are characters of the genus OrtliocMle : Probos- cis slender, elongated and directed straight downwards. Palpi likewise very elongated. The first joint of the antennae distinctly hairy on the upper side, the second transverse, the third not elon- gated. Arista dorsal, with an extremely short, almost impercepti- ORTHOCHILE — SYBISTROMA. 115 ble microscopic pubescence. The inferior corner of the eye dis- tinctly encased in the narrow cheeks. Hypopygiura entirely dis- engaged, sessile, its exterior appendages lamelliform. The first joint of the hind tarsi without bristles. The last segment of the fourth longitudinal vein converges towards the third, although but gradually, still sufficiently so as to reach the margin of the wing quite far from its tip, in the immediate vicinity of the tip of the third longitudinal vein. The whole structure of the body approaches rather closely that of the species of ffercostomus, still the species of Orthochile distinguish themselves sufficiently by the extraordi- nary elongation of the proboscis and of the palpi, and by the pre- sence of narrow cheeks ; besides, the tips of the third and fourth longitudinal veins lie more closely together and farther from the tip of the wing, than it is the case with any of the species of Hercostomus. The described species of this genus are found in Europe and in Asia Minor. The North American species, which Mr. Walker described as Orthochile derempta, cannot, by any means, be an Orthochile, as its arista has an apical position. In what genus it is to be located, or "whether a new genus is to be created for it, cannot be determined from the very imperfect state- ments of Mr. Walker, as he does not even state the sex of his spe- cimen, nor whether the first joint of the antennae is bare or hairy, whether the first joint of the hind tarsi is bristly or without bris- tles, and whether the fourth longitudinal vein converges towards the third or not. The name of the genus (from 6p£oj, straight, and ^saoj, the lip) has reference to the form of the proboscis, by which the species of this genus can easily be distinguished. Gen. VIII. SYBISTROMA. The following are the most important characters of the genus Sybistroma : Face not reaching as far as the lower corner of the eye, very narrow in the male, very broad in the female. The first joint of the antennas distinctly hafty^bn the upper side ; the second joint of the antennce transverse ; the third narrow and somewhat long in the male, broad and short in the female. Arista sub- apical ; in the male it is very long, its first joint longer than the second and thickened at the end in the shape of a knot ; the second has at its end a lamelliform enlargement. Scutellum very dis- 116 DIPTERA OF NORTH AMERICA. [PART II. tinctly hairy. Hypopygium entirely disengaged, on a short pe- duncle ; its exterior appendages lamelliform. The first joint of the hind tarsi without bristles. First posterior cell narrow, towards its end very narrow ; nevertheless the last segment of the fourth longitudinal vein only very gradually approaches the third longitudinal vein. The next related genera are Hercostomus and Hypophyllus.; their species were formerly located with Sybistroma. From both these genera Sybistroma differs, besides the peculiar structure of the antennas of the male, by the very distinct hairs on the scutel- lum. This genus was hitherto confined to the European Sybistroma nodicornis, unless perhaps Sybistroma Dufourii belongs to it. The derivation of the name of this genus is not clear to me. That it should be derived from clfivvij, the spear, and otpZfia, the couch, is not probable, at least it would then be a very unsuccess- ful composition. Gen. IX. HERCOSTOMUS. I have' established the genus Hercostomus in the fifth number of the " Neue Beitrage," upon the species Sybistroma cretifer H&l., fulvicaudis Walk., and longiventris Loew. These species agree in the following characters ; in the distinct pubescence of the otherwise plain arista ; in the structure of the proboscis ; in the distinct hairs upon the surface of the suctorial flaps ; in the glabrousness of the scutellurn, and in the apparently sessile hy- popygium of the male. Otherwise they approach the species of Hypophyllus very much, show however among themselves some marked differences, so that their consolidation into one genus may be considered as provisional, and will certainly have to be modified when a more considerable number of species will be known. In order to give more homogeneity to the genus Gymnopternus, it was necessary, as I have remarked before, to exclude all those species the third and fourth longitudinal veins of which are de- cidedly convergent. These species, however, approach the above named species of Hercostomus more than the species of any other genus. I see no difficulty at present to unite them with the genus Hercostomus, which is not ripe as yet for further subdivision on account of the insufficiency of our knowledge of its species. HERCbSTOMUS. 117 The characters of the genus may be defined as follows : An- tenna? of ordinary structure ; the first joint hairy on the upper side ; the second joint of the antennas transverse ; the third joint not elongated ; arista dorsal, of the usual plain structure. Scutellurn without hairs. Face not reaching as far as the in- ferior orbit. Hypopygium on a very short peduncle, so as to appear sessile ; exterior appendages lamelliform ; interior append- ages but little developed. The first joint of the hind tarsi without bristles. The first posterior cell narrowed towards its end ; tlje last segment of the fourth longitudinal vein only gradually ap- proaches the third longitudinal vein. The differences from the genera Gymnopternus, with its third and fourth longitudinal veins parallel, from Pelastoneurus with its feathered arista and the fourth longitudinal vein strongly in- flected forwards, Paraclius with the end of the fourth longitu- dinal vein angularly inflected forwards and then running towards the margin of the wing, in the shape of a segment of a circle, are self-evident. The species of Hercostomus differ from those of Hypophyllus by their arista, whieh is plain in both sexes, by the apparently sessile hypopygiuin and by the lesser development of its interior appendages. Up to the present time only European species have been made known ; I am now enabled to add to them a North American species. The name of the genus (from jpxoj, wall, fence, and otopa, mouth) has reference to the oral opening, surrounded, fence-like, by the suctorial surface covered with rows of hairs ; this being the case with those species on which I had originally established this genus. 1. H. ii til color, n. sp. £ . — Obscure viridis, nitidus, antennis, ocu- lorum tegularumque ciliis pedibusque totis nigris, alis ciuereis, lamellis hypopygii ovatis, nigricantibus, in disco sordidissime ex albidis. Dark-green, bright ; antennae, cilia of the inferior orbit and of the tegulas, also the feet, black ; wings gray, lamellae of the hypopygium oval, blackish, upon their middle very dingy whitish. Long. corp. 0.11 — 0.12. Long. al. 0.13. Dark metallic-green, almost black-green, bright. Front me- tallic-green. Antennae black ; third joint oval, at the tip only with a blunt point. The color of the narrow face seems to have been 118 DIPTERA OP NORfH AMERICA. [PART II. originally gray. Cilia of the inferior orbit, as far as I can dis- tinguish, black. Scutellum without hairs. Hypopygium black ; its lamella? rather large, oval, narrowed at the root, fringed with black hairs ; they have a blackish appearance, are however really black only on the margin, while in the middle, at least when seen in a certain direction, they look dingy whitish. Coxa? and feet black, the latter plain ; femora with a greenish reflection ; the upper side of the fore tibia? only with two small bristles. The yellowish-white tegula? have black cilia. Wings grayish hyaline with rather delicate black veins ; they are comparatively long and narrow and have a very regular elongated elliptic outline. The last segment of the fourth longitudinal vein is especially long, with an imperceptible sweep and approaches in its entire course gradually the third longitudinal vein, so that their ends are not very distant from each other. Hab. Port Resolution, Hudson's Bay Territory. (Kennicott.) Gen. X. HYPOPHYIXUS. This genus may be characterized in the following manner : The first joint of the antennae distinctly hairy on its upper edge, the second joint of the antennae transverse, the third not elongated ; the arista dorsal, very bare, rather strong as far as its end ; its first joint in the male remarkable either for its great length or its incrassated tip. The face of the male very narrow, especially below ; the face of the female broader, sometimes much broader. Scutellum not hairy. Abdomen stretched out, that of the male rather strikingly pointed at the end. The entirely disengaged hypopygiuin pedunculated, usually of a yellow color ; its exterior appendages lie on its under side, stretched out alongside of each other, and are small, narrow, elongated lamella? ; the interior ap- pendages are remarkable for their extraordinary development, are much longer than the exterior ones, either strap-shaped or broader at the end and beset with long hairs. Feet comparatively long and slender ; the first joint of the hind tarsi without bristles and shorter than the second. The last segment of the fourth longi- tudinal vein has only a very gentle sweep and very gradually ap- proaches the third longitudinal vein. The narrow, stretched-out shape of the body, the peculiar struc- ture of the arista, the long pedunculated yellow hypopygiuin and IIALTEIUCERUS. 119 the peculiar structure of its appendages, distinguish this genus sufficiently from all the other related genera. Only European species of Hypopliyllus are as yet known. The name of the genus (from vno, under, and $\>MOV, the leaf) has reference to the mode of life of the species, found in shady places on bushes and herbs and running on both sides of the leaves. Gen. XL HALTERICERUS. This genus was established by Mr. Rondani, in the year 1844, in the Xth volume of the Annali delle scienze Naturali di Bo- logna, under the name of Ludomcius, which afterwards, in the first volume of the Prodromus Dipterologise Italicse, he changed into Haltericerus. His statements with regard to the characters of the genus are not sufficient and not altogether correct. I am able to complete and to correct them as follows, from two Span- ish species of my .own collection, which, according to Mr. Hali- day, are also found in Upper Italy. The face of the male very narrow, that of the female compara- tively very broad ; in both sexes it does not quite reach the lower margin of the eye. Palpi small. The first joint of the antenna; of the male large and very much inflated, in the female much smaller and less inflated, in both sexes however beset on the upper side only with extremely short, rather imperceptible hairs. The second joint of the antennae in both sexes very small, rather rudi- mental, somewhat imbedded into the first joint and only distin- guishable by the bristles with which it is fringed at its end. The third joint of the antenna in the females of all species appears to be rounded ; in the males it has. either a more elongated or almost a conical form. The arista of the female is plain, dorsal, two- jointed, its first joint short. The arista of the male is also two- jointed ; its first joint is filiform and very much elongated, the ab- breviated second joint forms a flat lamella ; the position of the arista in the males with an oval third joint is distinctly subapical, in the other species it is apical or appears to be so. The neura- tion of the wings resembles that of the species of Systenus, the last segment of the fourth longitudinal vein being gently, but still sufficiently inflected forward to approach with its end closely to that of the third longitudinal vein. Feet slender, with scarce bristles ; the first joint of the hind tarsi without bristles and much 120 DIPTERA OF NORTH AMERICA. [PART II. shorter than the second. Hypopygium entirely disengaged and pedunculated ; its exterior appendages are lamelliform, the inte- rior ones much less developed than those of Hypophyllus. The species of Haltericerus are related to the species of the genera Hypophyllus and Hercostomus ; they differ from them sufficiently by the rudimentary second joint of the antennae and also the structure and position of the arista. They have also some resemblance to the species *of the genus Systenus, the first antennal joint of which, however, has no hairs on the upper side and the arista is distinctly apical in both sexes. The name of the genus (from dxr^p, the poiser, and xspas the horn) has reference to the remarkable structure of the arista of the male. As yet, only the three above named species of Haltericerus, which belong to the Fauna of Southern Europe, are known. Gen. XII. mOSTRACUS. The North American species, for which I have established this genus, resembles Thinophilus. In the structure of the abdomen it reminds me of Aphrosylus, with which it also agrees in the struc- ture of the hypopygium. It differs from both of these genera by the distinct hairs on the upper side of the first joint of the antenna?. The following may be considered as the characters of the pre- sent genus : Face in both sexes broad ; the palpi in the male of extraordinary size, in the female much smaller, and in both sexes loosely recumbent upon the proboscis. Antennae small ; first joint hairy ; second joint transverse ; third joint extremely small, dis- tinctly covered with hairs, and with a dorsal bristle. Thorax, scutellum and feet only with short and very scarce bristles. Ab- domen with short hairs, without any longer bristles before the in- cisures. First joint of the hind tarsi without bristles, considera- bly longer than the second. Wings long and narrow ; the poste- rior transverse vein somewhat close to the margin of the wing ; the last segment of the fourth longitudinal vein without any flexure, but only a little curved, converging somewhat towards the third longitudinal vein and ending beyond the tip of the wing. The fifth segment of the abdomen of the male is rather narrow ; the small sixth segment partially concealed under the former ; the DIOSTBACUS. 121 hypopygium .short, stout, rather disengaged ; its outer appendages are small lamella?. The genus Diostracus is so peculiar that more detailed state- ments are unnecessary to distinguish it from other genera. Its most striking character is the peculiar disk-like form of the palpi and their very considerable enlargement in the males. The name of the genus (from fii'j, twice, oarpaxov, potsherd) has reference to this character. 1. D. prasinus LOEW. $ and J . — Prasinus, subopacus, abdomine nigricante, pedibus flavis, maris tertio tarsorum anticorum articulo com- presso et in margine supero pilis nigris barbato. Leek-green, somewhat dull, with a blackish abdomen and yellow feet ; third joint of the fore tarsi of the male compressed and bearded on the upper margin with long hairs. Long. corp. 0.18 — 0.20. Long. al. 0.23 —0.24. SYN. Diostracus prasinus LOEW, Neue Beitr. VIII, 44, 1. Male. Face for a male very broad, metallic-green or blue- green, sometimes copper-colored, with a rather distinct transverse swelling, and by far not reaching the lower margin of the eyes ; dusted with grayish-yellow. Palpi of unusual size, rather rounded, loosely recumbent upon the proboscis, yellow, covered on the upper side with a thick snow-white powder, and with a delicate, almost imperceptible, white pubescence. Proboscis for a male unusually large and stout, brown. The small antenna? brownish-yellow, most of the third joint brown, the dorsal arista blackish-brown, long and not very strong, with a short, but distinct pubescence. Front above broader, metallic-green or blue-green, seen obliquely, somewhat darker, without dust. Cilia of the upper orbit black, of the lateral and inferior orbits yellow. Upper side of the thorax of a saturate leek-green or parrot-green coloring and with very little lustre ; on its anterior margin there is some gray-whitish pollen, which is interrupted by 'the anterior end of a not very striking and not far-reaching dark-colored middle stripe. The usual black bristles on the upper side of the thorax are few and short ; there are no hairs upon it. If examined from behind, a stripe- like dark scarlet-brown spot, immediately above the root of the wing, may be noticed ; on the posterior margin of the thorax there is also a more distinct covering of brown-grayish dust. Scutellum with the usual bristles, otherwise bare, somewhat short, usually of a more 122 DIPTEEA OF NORTH AMERICA. [PART II. dusky color than the upper side of the thorax. Pleurse with grayish dust upon blackish-green ground. The metathorax is of unusual length and slopes but very gradually, so that the length of the thorax, as compared with that of the abdomen, is unusually large. The color of the abdomen is dusky blackish-green and but little metallic ; the hairs are short and only on the posterior margin of the first segment there are some black hairs of greater length. The black hypopygium is short and stout, sessile, but rather disengaged ; its small outer appendages are lamelliform, blackish-brown and hairy. Fore coxoe long, pale yellow ; on the front side they are beset with so short and delicate white little hairs that they appear gla- brous ; at their tip there are black bristles. Middle and hind coxae yellow, often brownish as far as the tip, especially on the outside. Feet yellow, rather long ; middle and hind femora very slender ; fore femora considerably stronger ; all the tibiae on the under side very bare, otherwise beset with quite short little black hairs ; fore tibiae rather stout, somewhat compressed and curved inwardly, colored with brownish-black upon the latter half of the upper side and most of the hind side thickly bearded with long yellow hairs. Middle and hind tibiaa plain, beset only with few and weak bristles, infus- cated at the end. Fore tarsi black, only at the basis of the first joint brown ; the first joint only a little longer than the second ; the second at the tip with a vestige of a slight compression ; the third joint strongly compressed, broad, bearded on the upper edge with stiff black bristle-like hairs ; the two last joints very short, and of the usual form. Middle tarsi somewhat longer than the tibiaa, the first joint about as long as the other three taken together, yel- lowish-brown with black tip ; the last four joints are black and the middle tarsi on the hind side rather thickly covered with long hairs. Hind tarsi black, about as long as the tibioa, of plain struc- ture and not unusually hairy ; the first three joints are of gradually diminishing length, the fourth about half as long as the third, and the fifth again somewhat longer than the fourth. The teguloe have brown margins with yellowish cilia, which assume, in some directions, a brown tinge. Wings grayish hyaline, long and nar- row with rather strong brownish-black veins ; the first longitudinal vein reaches far beyond the third part of the anterior margin ; second longitudinal vein straight ; the third longitudinal vein at its end only very gently curved backwards ; the posterior trans- verse vein lies far beyond the middle of the wing. ANEPSIUS — ARGYRA. 123 _ Female. Face broader than that of the male, with a more developed transverse swelling, and covered with dust of much more gray color. Palpi much smaller than those of the male, only about half as large, blackish with yellow-grayish dust, and with a considerable covering of pale brown hair, which in another direc- tion appears to be entirely black. The sixth segment of the ab- domen, although very short, still distinctly perceptible. The hairs on the anterior side of the fore coxae longer and coarser, yellowish. Tibia? and tarsi simple and with the usual short hairs ; the joints of the fore tarsi gradually diminishing in length. The wings usually a little more tinged than those of the male. Hob. New York. (Osten-Sacken.) Gen. XIII. ANEPSIUS. The genus Anepsius shows the closest relation to the genus Systenus, from which it differs only by the structure of the first joint of the antennae. Its characters are the following : The first joint of the antennae hairy on the upper side, the second transverse, the third rather large ; the arista inserted on its upper side, quite near the basis. The abdomen of the male appears compressed from the side. Hypopygium short, not entirely imbedded ; its outer appendages very small. The first joint of the hind tarsi without bristles. Wings not enlarged towards their basis ; the last segment of the fourth longitudinal vein not inflected and parallel to the third vein. The name of the genus (from avsfyos, cousin) has reference to its intimate relationship with Systenus. No species of Anepsius has yet been found besides those known from Europe. Gen. XIV. ARGYRA. The species of Argyra • are easily distinguished by the dense silvery-white dust, which almost in all the species covers head and abdomen, in many also thorax and scutellum. The majority of the species of the genus Leucostola resemble in this respect those of Argyra, and differ from them only by the first joint of the antennae of the former being entirely bare, while in the species of Argyra it is covered with hair on the upper side. The following are the most important characters of the genus Argyra : Second 124 DIPTERA OP NORTH AMERICA. [PART II. joint of the antennae transverse; third in the male rather large, bare ; the apparently bare and distinctly two-jointed arista is in- serted close to the tip of the antenna. Wings broad, the posterior angle rather strongly projecting ; the first longitudinal vein is rather more distant from the margin of the wing than in most of the other genera, and is longer than usual ; the fourth longitudinal vein is inflected forward before the middle of its last segment, thence however it is quite parallel, or almost parallel, with the third longitudinal vein ; the posterior transverse vein is not ap- proximated to the margin of the wing. The first joint of the hind tarsi without bristles. Hypopygium small, imbedded ; its outer appendages are two very small, narrow lamellfe directed downwards ; the interior appendages are of rather simple struc- ture and often not distinctly perceptible. The name of the genus (from apyvpoj, silver) has reference to the beautiful silvery lustre of most of the species. The hitherto known species are distributed over Europe, a part of Asia and North America. I know seven North American species, of which the first has a hairy scutellum and therefore belongs to the relationship of the European Argyra diaphana. The other six species have no hairs upon the scutellum ; the upper side of the first joint of the antennas is, in some of them, so scarcely provided with hairs that they can easily be mistaken for species of Leucostola. Table for the determination of the Species. , f Scutellum hairy. 1 albicans Lw. I Scutellum not hairy. 2 0 f Feet mostly black. 3 i I. Feet entirely or mostly yellow. 4 f The entire fore tibiae and a part of the four posterior tibiae yellow. 2 mgripes, nov. sp. 1 All the tibiae entirely and the root of the fore tarsi yellow. 3 albiventris, nov. sp. , } The whole feet yellow. 5 I Not the whole feet yellow. 6 f The first joint of the hind tarsi only with the usual very short hairs. 4 minuta Lw. 1 The first joint of the hind tarsi with longer hairs than usual. 5 calcitrans Lw. „ I Tjp of the hind femora not black. 6 calceata Lw. I Tip of the hind femora black. 7 cylindrica, nov, sp. ARGYRA. 125 Systematical arrangement of the Species. I. Scutellum distinctly hairy. t 1. albicans Lw. II. Scutellum without hairs. A. Abdomen somewhat conical, white, glittering. 2. nigripes, nov. sp. 5. calcitrans Lw. 3. albiventris, nov. sp. 6. calceata Lw. 4. minuta Lw. B. Abdomen entirely cylindrical, without white glitter. 7. cylindrica, nov. sp. i Description of the Species. I. SCUTELLUM DISTINCTLY HAIRY. 1. A. alllicans LOEW. £ and 9. — Scutello piloso, tibiarum postica- rum apice tarsisque posticis nigris. Scutellum hairy, tip of the hind tibize and the hind tarsi black. Long, corp. 0.23—0.24. Long. al. 0.23—0.24. SYN. Argyra albicans LOEW, Neue Beitr. VIII, 45, 1. Male. Covered all over with glittering, silvery-white dust. Face and front of middle breadth, silvery-white. Palpi black. Pro- boscis brownish-black. Antenna? black, the third joint more brown-black ; arista distinctly inserted before its end. Cilia of the superior orbit black, the hair-like cilia of the lateral and infe- rior orbits snow-white. Upper side of the thorax and Scutellum shining green, still so that the silvery-white dust seems to cover the ground color, even if looked upon from different sides. The scutellum has upon its upper side, besides the usual bristles, some very distinct little black hairs. Groiind color of the abdomen greenish-black, the second and third segments have very large, rounded, transparent yellow lateral spots ; otherwise, the whole abdomen is also covered with thick silvery-white dust ; its hairs are almost exclusively black. The small and narrow lamella? of the hypopygium are brownish-yellow with black tip and with a black pubescence. Fore coxse yellowish with white dust and with black bristles and little hairs. Middle and hind coxa3 black with white dust; their hairs and bristles black. Feet yellow with black hairs ; upon the under side of the fore and middle femora are in- 126 DIPTERA OF NORTH AMERICA. [PART II. serted black hairs of greater length than those on the under side of the hind femora, which are brownish-black at the tip. Tibia? moderately provided with bristles, the hind tibise blackened at the tip. Fore tarsi towards the end only slightly infuscated ; the first joint is at least 1± the length of the four following joints together, upon the under side with a row of delicate, but rather long little hairs, which may be easily overlooked. Middle tarsi from the tip of the first joint blackish-brown ; however, the root of the next following joints somewhat paler ; the first joint is at least by one- third longer, than the following four joints together. Hind tarsi entirely black, first and second joint of about the same length, the following of a decreasing length. Cilia of the tegulse, which have a black margin, yellowish-white. Wings somewhat grayish with blackish-brown veins ; the last segment of the fourth longitu- dinal vein is strongly bent forward before its middle. Female. Face broader than that of the male, and the palpi much larger, the latter with a distinct covering of white dust. The third joint of the antennae, as usual, much smaller than that of the male. The dust upon the thorax and the scutelluni less thick, so that the green ground-color is distinctly visible in every direction. The ground-color of the abdomen is shining and metallic-green, the dust upon it is confined to the anterior half of the segments,, extending only upon the last segment as far as the posterior mar- gin ; the yellow spots of the second segment are about of the same size and nature as those of the male, those of the third segment occupy only the anterior corners and are much smaller. The hairs upon the under side of all the femora are short and the hind femora are very slightly infuscated at the tip only. All the rest as in the male. Hob. District Columbia. (Osten-Sacken.) Observation 1. — Notwithstanding the difference in the coloring of the hind femora, the, female agrees with the above described male so much, thai I cannot have the least doubt of its being the other sex of Argyra albicans. Observation 2. — Argyra albicans can easily be distinguished from the European Argyra diapliana by its somewhat smaller size, the paler color of its fore coxae and of the cilia of the tegulse. Nevertheless it resembles it very much. As Fabricius, in his Systema Antliatorum, states America to be the home of his Musca diaphana, it might well be supposed that he meant Argyra albi- ARGYRA. 127 cans or some similar American species, and that Meigen was mis- taken when he believed it to be identical with the common European species. Such a supposition, however, would be erroneous. Fabricius in his older works (of which at present I cannot com- pare only the Mantissa II), mentions everywhere Europe as the habitat. The statement of the Systema Antliatorum is, there- fore, either a mere mistake, or Fabricius confounded later an American species with the European one. Even in the latter case the name cannot be transferred upon the American species. The first, however, seems to be more probable, because Fabricius in the Systema Antliatorum, quotes his former works without the least hesitation, and declares that America is the habitat of this species, without mentioning at all that he is thus in contra- diction with his own previous statement. II. SCUTELLUM WITHOUT HAIRS. A. Abdomen somewhat conical, glittering with white. 2. A. TI i gripes, n. sp. •£ . — Ex viridi Isete chalybea, abdomine albo •micante, fronte et facie atris, velutinis, pedibus nigris, tibiis anticis totis, reliquis ex parte flavicautibus. Green-blue, with the abdomen glittering white ; front and face velvet- black ; feet black, the fore tibise entirely and the four posterior tibize partially yellowish. Long. corp. 0.17 — 0.18. Long. al. 0.17. Green-blue. Front and face velvet-black, without pale dust. Palpi and proboscis black. The first joint of the antenna? with a comparatively long and close pubescence. Upper side of the thorax and of the scutellum shining ; the latter, as far as percepti- ble, without hair. Abdomen without transparent yellow spots upon the anterior segments, and with a thick glittering-white dust, which becomes thinner only towards the basis of the abdomen. The small lamellse of the hypopygium are brownish-black. Coxje black ; fore coxa? with coarse black hairs. Femora brownish- black, the extreme tip of the four anterior ones yellow ; they have only short hairs and short bristles. Fore tibia? yellow, on the upper side with less numerous, but stronger hair-like bristles, on the hind side with more numerous but more slender hair-like bris- tles of considerable length. Middle tibia) upon the whole upper side yellowish, upon the under side blackish-brown ; they have upon the under side, upon the middle of the anterior side and 128 DIPTERA OP NORTH AMERICA. [PART II. upon the first half of the upper side rather strong bristles. Hind tibiae at the root and almost upon the whole second half black, otherwise brownish-yellow and not incrassated. Fore tarsi yellow, somewhat infuscated towards their end, of plain structure, only the first joint upon the under side provided with a few bristles. Middle, tarsi brownish-yellow at the basis, further brownish-black ; the first joint has upon the first half of its under side rather numer- ous black bristles. Hind tarsi black, plain. Cilia of the tegulse black. Wings hyaline, only a little tinged with gray. Hab. Sitka. (Sahlberg.) Observation. — Of this and of the next following species, as also of Argyra cylindrica, I possess only single specimens, damaged by mould, which crumbled to pieces in the attempt of cleaning ; however, as these three interesting species come from a country which is so little explored with regard to Dipterology, and as the important specific characters could be determined, I did not hesi- tate to describe them. I must, however, request not to attach more weight to my statements about color, diffusion of the white tomentum upon abdomen and thorax, and about the appendages of the hypopygium, than the circumstances should warrant. » 3. A. alMventris, n. sp. £ . — Viridis, nitens, abdomine albomi- cante, fronte et facie nigris, albido-pollinosis, pedibus nigris, tibiis om- nibus totis tarsorumque anticorum basi flavicantibus. Green, shining, abdomen white, glittering ; front and face blackish, but with whitish dust ; the whole tibia and the root of the fore tarsi yel- lowish. Long. corp. 0.18 — 0.19. Long. al. 0.18. Metallic-green. Front and face appear, when looked upon in most directions, almost whitish-gray on account of the dust which covers them, but are black. Palpi and proboscis black. An- tenna? smaller than those of Argyra nigripes ; the first joint is comparatively short and sparsely beset on the upper side with rather short hairs. Thorax and scutellum shining metallic-green ; but when looked upon from the front, the covering of white dust becomes distinctly visible. Scutellum without hairs. Abdomen covered with a thick dust having a white lustre ; its second and third segments have, on the lateral margin, a large yellow transparent spot. Venter mostly yellowish. The small lamellae of the hypo- pygium are brownish-black. Coxa? black ; fore coxa? with long ARGYRA. 129 black hairs. Femora brownish-black with yellow tip ; their black hairs are comparatively long, especially upon the under side of the four anterior femora. Tibia? yellow, only the extreme tip of the hind tibiae blackish-brown ; the fore tibiae are beset with four or five bristles only upon the upper side ; the small bristles upon the middle and hind tibiae are likewise but short and very scarce ; the hind tibiae are not in the least thickened. The four anterior tarsi • are brownish-yellow at the basis, a little further blackish-brown, of plain structure, the first joint upon the under side without bristles. Hind tarsi entirely black. Tegulse mostly black with black cilia. Wings hyaline, a little more distinctly, tinged with gray than in A. nigripes. Hab. Sitka. (Sahlberg.) 4. A. minilta LOEW. £ . — Scutello nudo, pedibus totis pallide flavi- cantibus, metatarso maris postico simplici, brevissirne piloso. Scutellum bare ; the whole feet pale-yeUowish ; the first joint of the hind tarsi of the male simple, with very short hair. Long. corp. 0.16. Long, al. 0.14—0.15. SYN. Aryyra mimita LOEW, Neue Beitr. VIII, 46, 2. Male. Face very narrow, silvery-white. Palpi black. Pro- boscis brownish-black. Antenna black, third joint more blackish- brown ; arista distinctly inserted before its tip. Front silvery- white. Cilia on the upper orbit black, very short and delicate ; cilia of the lateral and inferior orbits white. Thorax and scutel- lum shining green, covered with white, not very thick dust, which conceals the ground color at the utmost only in the vicinity of the shoulders. Scutellum, with the exception of the usual bristles, bare. The ground color of the abdomen appears to be blackish- green, is however so thickly covered with white dust that it can- not be distinctly perceived ; the second segment of the abdomen has on each side a very large rounded yellow lateral spot ; the third segment has a similar spot, but smaller. The very small and narrow lamellae of the hypopygium are yellowish-brown and but little hairy. Fore coxa? yellowish-white, with white hairs and black bristles. Middle and hind coxae also yellowish-white, but blackened from the basis to a considerable extent ; hairs and bristles black ; feet pale-yellowish, only the end of the hind femora has a vestige of infuscation on its upper side. The hairs upon 9 130 DIPTERA OF NORTH AMERICA. [PART II. the feet are not entirely black ; the fore femora have upon the hind side more, the hind femora on the under side less numerous blackish hairs of greater length. Fore tarsi not infuscated ; the first joint has scarcely 1^ the length of the four following joints put together. Only the last joint of the middle tarsi is somewhat infuscated. The first joint is fully as long as the others together. • The last joint of the hind tarsi is brown ; the first joint is of sim- ple structure and beset with the usual short hairs, scarcely a little longer than the second, but considerably thicker. Cilia of the tegulffi yellowish-white. Wings hyaline, scarcely a little tinged with yellowish-gray, with brownish-yellow veins ; the last segment of the fourth vein is very abruptly bent forward before its middle. Hob. District Columbia. (Osten-Sackeu.) 5. A. calcitraiHS LOEW. ^ . — Scutelloc nudo, pedibus totis pallide flavicantibus, metatarso postico maris paulo incrassato et valde liirto. Scutellum bare ; the whole feet pale-yellowish ; the first joint of the hind tarsi of the male somewhat thickened and covered with rough hair. Long. corp. 0.14 — 0.15. Long. al. 0.11. SYN. Argyra calcitrans LOEW, Neue Beitr. VIII, 46, 3. Male. Face narrow, silvery-white. Thorax and proboscis black. Antennas black, third joint more blackish-brown ; arista distinctly inserted before its end. Front silvery-white. Cilia of the upper orbit black, very short and delicate ; cilia of the inferior and lateral orbits white. Thorax shining green, covered with thick white dust, so as to make the ground color invisible in some directions. Scutellum also shining green, or blue-green, with less dust, and bare with the exception of the usual bristles. The ground color of the abdomen seems to be blackish-blue, but cannot be distinctly seen on account of the thick white dust which covers it ; the second segment is yellowish and transparent, with a black- ish border on the posterior margin and with a blackish middle line, which is sometimes wanting ; the third segment is of a simi- lar color, only the margin on the posterior border and the middle line are broader, though the latter is sometimes interrupted. The email lamellae of the hypopygium are brown. Coxaa and feet white-yellowish ; fore coxaB with delicate white little hairs .and delicate black bristles ; middle coxae on the outside with a black- ish spot ; hind coxa? scarcely a little blackened on the extreme ARGYRA. 131 basis. Fore and middle femora without any longer hairs on the under side ; the hind femora have a row of black bristle-like hairs, inserted rather more on their hind side than upon the under side. Tarsi towards their end not infuscated, but only very little darker ; the first joint of the fore tarsi is 1^ the length of all the following joints taken together, on the under side with a row of delicate hairs ; the first joint of the middle tarsi not quite as long as all the other joints together ; hind tarsi unusually short, the first joint not much shorter than all the others, a little thickened, on the under side with rather long bristle-like hairs ; the second and third joints of the hind tarsi of about equal length. The cilia of the tegulae, which have a blackish margin, are white-yellowish. Wings some- what tinged with yellowish-gray, with yellowish-brown veins ; the last segment of the fourth longitudinal vein is but gently inflected forward. Hob. New York. (Osten-Sacken.) 6. A. calceata LOEW. £. — Scutello nudo, pedibus pallide flavis, tarsis posticis nigris. Scutellum bare, feet pale-yellow, with black hind tarsi. Long. corp. 0.16. Long. al. 0.15. SYN. Argyra calceata LOEW, Neue Beitr. VIII, 47, 4. Female. Face, for a female, of moderate breadth, silvery- white. Palpi rather large, black, with almost silvery-white dust ; proboscis brownish-black. Antennae black ; third joint small, with an acute angular tip ; arista distinctly inserted before its end. Cilia on the upper orbit black, extremely short and delicate. Cilia of the lateral and inferior orbits white. Thorax shining green, in the vicinity of the shoulders rather thickly covered with white dust, so as to induce the belief that the males are altogether covered with silvery-white dust. Scutellum also shining green and with the exception of the usual bristles, bare: Ground color of the abdomen greenish-black ; the second, third and fourth seg- ments yellow, with the exception of the extreme anterior margin, the posterior margin and a narrow, not always distinct middle line ; on the sides of the segments and towards the tip of the ab- domen there is some white dust. Coxae and feet pale-yellowish ; fore coxa? with delicate whitish hair and with black bristles ; mid- dle coxae with a gray spot on the outside ; hind femora scarcely a little darker at the tip, hind tibia; at the tip not of a dark color. 132 DIPTERA OF NORTH AMERICA. [PART II. Tarsi comparatively long ; fore tarsi scarcely infuscated towards the end, only the last joint brown, the first joint a little longer than the others taken together. Middle tarsi from the tip of the first joint strongly infuscated ; the four last joints together as long as the first one. Hind tarsi entirely black, the first joint a little shorter than the second, the following joints decreasing in length. The cilia of the tegula?, which have a blackish border, are pale. Wings tinged more with brownish than yellowish-gray ; veins rather dark brown ; the last segment of the fourth longitudinal vein, before its middle, only slightly inflected forward. Hob. Middle States. (Osten-Sackeu.) Observation. — It is impossible to mistake A. calceata for the female of one of the two preceding species for the following rea- sons : — 1. On account of the greater extent of yellow color upon the abdomen, which is not so extensive in the females of the species of Argyra as in the males. 2. ' On account of the black coloring of the whole hind tarsi. It is probable that the coloring of the abdomen of the male of this species resembles that of the male of Leucostola cingulata. B. Abdomen entirely cylindrical, •without white lustre. 7. A. cylindrica, n. sp. £ . — Viridis, nitens, abdomine vix obso- letissime albido pollinoso, coxis'anticis pedibusque flavis, femorurn pos- ticorum apice, summo tibiarum posticarum apice, tarsis denique omni- bus inde ab articuli prirni apice nigris. Shining-green, the abdomen has scarcely a trace of whitish dust ; fore coxae and feet yellow ; tip of the hind femora, tip of the hind tibiae and all tarsi, with the exception of their root, black. Long. corp. 0.23. Long. al. 0.22. Metallic-green, and differing from most of the species of by the cylindrical form of the abdomen and the almost entire absence of all dust. Face with a white reflection. Palpi and proboscis brownish-black ; the hair-like cilia of the inferior orbit pale-yellowish. Thorax and scutellum shining, the latter without hairs. Abdomen entirely cylindrical, not stouter about the basis, metallic-green, without any transparent spots upon the anterior segments and covered with a thin, almost impercep- tible, grayish-white dust. Fore coxa3 yellowish, with some very scattered black hairs, and, at the tip, with longer black bristles. SYNTORMON. 133 Middle and hiiid coxa? blackish-brown with yellowish tip. Feet yellowish, tip of the hind femora brownish-black to a considerable extent. The black hairs on the feet are somewhat sparse and only of middle length ; on the under side of the fore femora there are but a few black hairs of greater length ; upon the under side of the middle femora they are in greater number ; on the under side of the hind femora only those hairs which are near the tip have a somewhat greater length. Fore tibia? with somewhat longer hairs on the under side, on the upper side with but two or three bristle-like hairs of greater length. The middle and hind tibia? with but few and very short little bristles ; the extreme tip of the latter is infuscated. (Fore tarsi wanting.) Middle tarsi plain, black from the tip of the first joint ; the first joint somewhat longer than the four following together ; on the under side with but two very short black little bristles ; hind tarsi black ; the first joint up to the middle brownish-yellow. Tegula? with a broad black margin, with pale-yellowish cilia. Wings hyaline, some- what tinged with brown, the anal • angle less protruding than in most of the other species ; the ueuratiou shows nothing unusual for the genus. Hob. Sitka. (Sahib erg.) Observation. — Although the imperfect state of the above de- scribed specimen did not afford a thorough examination of the hairs upon the upper side of the first joint of the antenna?, still I believe to have satisfied myself of their existence. Gen. XV. SYNTORMON. The generic character is as follows : First joint of the% antenna? with hairs on the upper side ; the second reaching on the inner side of the third in the shape of a thumb ; third joint of the an- tenna? elongated and pointed in the male and shorter in the fe- male ; the position of the arista apical. Scutellum bare. The first joint of the hind tarsi without bristles and a little shorter than the second. The hypopygiuni small and imbedded, with very small, often not distinctly perceptible appendages. The next related genus is Synarthrus, the species of which differ from the species of Syntormon only by the absence of hair upon the first joint of the antenna?. Although this character may appear trifling to those who have not studied the family of the 134 DIPTERA OP NORTH AMERICA. [PART II, Dolichopodidse carefully, nevertheless its observation is important, as it is of very great service for the specific distinction as well as for the generic location of such females of different species which resemble each other very closely. The name of the genus (from awtoppou, I connect by inserted pins) has reference to the characteristic formation of the second joint of the antenna?. As yet only European species of Syntormon have become known. Gen. XVI. SYN ARTHRUS. The following are the most important characters of this genus : The first joint of the antenna? without hairs ; the second reaching more or less on the inner side of the third, usually forming in the males a thumb-like projection, and in the females a more rounded lobe ; the third joint of the antennae in the males elongated and pointed ; arista apical or so near the extreme tip of the antenna?, as to be taken for such. Scutellum usually bare, first joint of the hind tarsi without bristles. The hypopygium small, imbedded, with very small, often not distinctly perceptible appendages. That Synarthrus differs from Syntormon only by the absence of hair upon the first joint of the antennae, has already been stated. Of the three species described below, two undoubtedly belong to the genus Synarthrus. This cannot be said with regard to the third species, Synarthrus barbatus. Its position in the genus Synarthrus can only be a temporary one, brought about by the difficulties of placing it into another genus. It is sufficiently dis- tinguished from the other species of Synarthrus by the peculiarity alone, that the second joint of the antennae encroaches only very little on the inner side of the third. In its general appearance it approaches the species ofPorphyrops very closely, so that I leave it undecided whether it would not be better located there. The size of the pulvilli of the fore tarsi betrays a relationship with Eutarsus and Diayjhorus, the structure of the antennae, however, does not allow its location in these two genera. To erect a new genus does not seem advisable, as the species shows close rela- tionship in various directions. The name Synarthrus (from avv, together, and dp^pov, joint) has reference to the peculiar mode of connection between the second and the third joints of the antenna?. SYNARTIIRUS. 135 .The species already known belong to Europe and to North America. Table for the determination of the Species. 1 f Posterior margin of the pleurje yellow. 1 palmaris, nov. sp. I- Posterior margin of the pleurae not yellow. 2 2 f Hind femora yellow. 2 cinereiventris Lw. I Hind femora green. 3 barbatus Lw. Systematical arrangement ofJhe Species. I. Lower half of the occiput only with the usual cilia. 1. palmaris, nov. sp. 2. cinereiventris Lw. II. Lower half of the occiput with a strong beard. 3. barbatus Lw. Description of the Species. I. LOWER HALF OP THE OCCIPUT ONLY WITH THE USUAL CILIA. 1.- S. palmaris, n. sp. £ and £ • — Viridis, pleurarum margine pos- tico, coxis, pedibusque flavis. £ . Tarsis intermediis apicem versus dilatatis. 9 . Tarsis simplicibus. Green, the posterior margin of the pleurae, coxse and feet yellow. £ . The middle tarsi towards their end enlarged. $> . Tarsi simple. Long. corp. 0.13. Long. al. 0.14. Hale. It can be easily recognized by its striking resemblance to the European Syntormon tarsatus Fall. Rather dark bronze- green, little shining ; the abdomen usually rather copper-colored. Antenna? black ; first joint on the upper side entirely bare ; the second with a long thumb-like projection which overlaps the inner side of the third ; third joint- rather long, pointed, with much hair ; the arista has not precisely an apical but a somewhat sub- apical position, as it is the case with Syntormon tarsatus. Face covered with whitish dust, narrow. Palpi and proboscis black. Front steel-blue. The delicate cilia on the inferior and lateral orbits pale. Scutellum of the same color as the upper side of the thorax, arid beset, besides the usual bristles, with a few short hairs. Pleura? grayish-green, their posterior margin yellowish. The sides of the abdomen near its basis somewhat yellowish and trans- parent; on the posterior margin of its first segment there are long 136 DIPTERA OF NORTH AMERICA. [PART II. black bristles, on the posterior margin of the other segments there are only very short bristles. Venter as far as the tip yellowish. Hypopygium rounded, small, rather imbedded ; its blackish ap- pendages very short and therefore not distinctly perceptible. Coxa? and feet yellowish ; fore coxoe only at the tip with a few black bristles, otherwise in front with short and very delicate white hairs. The short hairs upon the feet are black ; under side of the fore femora glabrous ; middle femora upon the under side with a row of short black bristles. Fore tibice on the upper side with a dense row of black hairs, upon the first quarter of the hind side with a single very small black bristle. Middle and hind tibiae only with a few short black bristles ; the end of the latter is some- what thickened and of a brownish-black color. Pore tarsi plain, from the tip of the first joint blackened ; the first joint not quite as long as the other four taken together. The first joint of the middle tarsi nearly as long as the other four taken together, stalk- like, a little thicker at the end and of a whitish color ; the second joint very much flattened, whitish, the apical margin bordered with black ; the third and fourth joints also flattened, and, with the fifth joint, which is not flat, of a deep black color. Hind tarsi as far as the middle of the second joint brown, then black ; the first joint is a little stouter and shorter than the second and has upon the middle of its under side a black bristle which is a little curved backwards. Wings hyaline, tinged with brownish-gray ; the third and fourth longitudinal veins show towards their end a slight convergency. Female. The third joint of the antennas rounded-oval, very much shorter than in the male ; the second joint of the antennae overreaches the inner side of the third only by a rounded lobe. Face less dusty than in the male, very broad, by far not reaching the lower corner of the eye ; its lower part is very convex and protrudes in the shape of a roof, whereby the mouth becomes very large. The black palpi considerably larger than in the male. Feet plain, the hind tibia3 not thickened at the end and only very slightly infuscated. Fore tarsi from the tip of the first joint gra- dually becoming more infuscated ; middle tarsi from the tip of the first joint blackened, though the root of the second joint is some- what paler ; hind tarsi at the root brown, black towards the end. The little black bristles upon the under side of the middle femora are missing. The rest as in the male. Hob. Sitka. (Sahlberg.) SYNARTHRTJS. 137 Observation. — Syntormon tarsatus has upon the upper side of the first joint of the antennae, quite in the vicinity of the basis, a few almost imperceptible little hairs, which I ani unable to per- ceive in the present species, so that I ana compelled to locate it into the genus Synartlirus. It is further distinguished from Syn- tormon tarsatus by the shorter arista and by the broader wings, which are less narrow towards the root. Besides, in the male the tip of the hind tibia? is less thickened, the second joint of the mid- dle tarsi is broader, of a whitish color, and has only at its tip a black margin, while in the male of Syntormon tarsatus it is less broad, yellowish upon the middle of the root and black upon the whole apical half. The relation which exists between Synarthrus palmaris and Syntormon tarsatus is exactly the same as that be- tween Synarthrus pallipes and Syntormon Zelleri. 2. S. ciBiereiveiltris LOEW. 9 • — Viridis, cosis anticis pedibusque flavis. Green, fore coxae and feet yellow. Long. corp. 0.13. Long. al. 0.13. STN. Synarthrus cinereiventris LOEW, Neue Beitr. VIII, 48, 1. Female. Shining metallic-green. Face very broad ; the infe- rior portion protrudes considerable in the shape of a roof, so that the palpi are rather concealed and the mouth becomes unusually large ; the blue-green ground color of the face is rather covered by a gray-whitish powder. Antennae black, rather large for a female ; the second joint overreaches the inner side of the third with a broad lobe ; the third joint is short, but very broad, beset with very short but distinct hair ; the arista is distinctly longer than the antennae. The front in the middle is almost blackish- green, about the antennas steel-blue, o*n the upper corners rather violet and may possibly vary a little in its coloring. Cilia of the upper orbit black, those of the lateral and inferior orbits white. Thorax shining metallic-green, with very thin and almost imper- ceptible, nearly whitish, dust. Abdomen also shining metallic- green, on the lateral margin with but little perceptible whitish powder. Tenter whitish-gray. Fore coxse yellow, with delicate whitish hair and no black hairs or bristles. Middle and hind coxa? blackish with yellow tip, the former green on the front side, and, like the fore coxa?, beset with delicate whitish little hairs. Feet yellow ; tarsi gradually dark brown towards their end, but 138 DIPTERA OF NORTH AMERICA. [PART II. so that the first joint, with the exception of its tip, is dark yellow, and the second joint, with the exception of its tip, is brownish- yellow. The hind tarsi are visibly shorter than the hind tibia?, and the first joint is longer than the second. The yellowish tegu- IJB have a narrow dark-brown margin and yellowish cilia, which in some directions assume a very dark color. Halteres yellowish. "Wings grayish ; the last segment of the fourth longitudinal vein not distinctly inflected ; posterior transverse vein steep. Hob. Middle States. (Osten-Sacken.) II. LOWER PART OP THE OCCIPUT" WITH A STRONG BEARD. 3. S. "barbatus LOEW. $ . — Viridis, pedibus flavis, femoribus posticis viridibus. Green, feet yellow, hind femora green. Long. corp. 0.12 — 0.13. Long. al. 0.12. SYN. Synarthrus barbatus LOEW, Neue Beitr. VIII, 48, 2. Shining metallic-green. Face green, and dull on account of a moderately thick yellowish-white dust, which however does not conceal the ground color. Palpi a little larger than those of the males of Synarthrus in general ; they have also a more disengaged position, a bright yellow color, and are beset with a few black little hairs. Antennas black ; the second joint reaching only a little on the inside of the third, almost of transverse form ; the rather strikingly hairy third joint is broad only about the basis and becomes elongated into a narrow and very long point; the arista is entirely apical and only half the length of the third joint. Front metallic-green, little shining. Cilia of the upper orbit black, short and delicate ; cilia of the lateral and inferior orbits very long, white ; they fortii with the hair on the lower part of the occiput a striking beard. Thorax and scutellum metallic-green, the former less shining on account of a cover of pale dust, the lat- ter sometimes more bluish-green. Abdomen metallic-green. The last segments assume in some directions a rather black-green color. The small imbedded hypopygium black ; its outer append- ages have the form of small lamella? and are of a brownish color ; the inner appendages are not distinctly perceptible. Fore coxae blackish, on the front side greenish-blue ; their delicate hairs are principally white, but towards the basis of the coxae there are also some black hairs ; on the tip are black bristles. Middle and hind SYNARTIIRUS — SYSTENUS. 139 coxa: black ; the former with black hairs and bristles. Fore feet yel- low ; femora incrassated towards their basis, on the upper side with a narrow brownish-black longitudinal stripe, reaching to the ex- treme tip ; the under side is sparsely fringed with short black hairs, and where these end, there are three black bristles turned somewhat backwards. Fore tibine covered with thick, rather coarse, but short hairs. Fore tarsi from the tip of the first joint, which is about as long as the others, strongly infuscated. The first and second joints beset .on the under side with short bristle- like hairs ; pulvilli rather large. Middle feet yellow ; femora thickened about their basis ;' the tibia3 have besides the bristle at the tip only one on the upper side, not far from the basis ; the tarsi are very much infuscated from the tip of the second joint. Hind femora metallic-green with yellow tip, at the end of the under side with a few black bristles. Hind tibia? yellow, only sparsely bristled. Hind tarsi yellow at the basis, from the tip of the first joint black-brown ; their joints decrease in length, the first considerably longer than the second. The cilia of the whitish tegulas show in most directions a brownish-black coloring, while in some, they appear with a yellowish lustre. Halteres yellowish. Wings grayish hyaline with brownish-black veins ; the posterior transverse vein is straight and has a very steep position ; the last segment of the fourth longitudinal vein shows upon its first third an almost imperceptible flexure, otherwise it very little approaches the third longitudinal vein, runs parallel with it and terminates precisely in the apex of the wing. Hob. Middle States. (Osten-Sacken.) Gen. XVII. SYSTENUS. This genus, hitherto confined to European species only, may be characterized in the following manner : First joint of the antennae without hairs on the upper side ; the second joint transverse ; the third large, longer in the male than in the female, in both sexes broad at the basis, ending into a point, distinctly hairy ; arista completely apical. Feet rather slender, in both sexes plain and b'eset only with very few and short bristles. The first joint of the hind tarsi without bristles and scarcely half the length of the second. The sixth longitudinal vein of the wings distinct. The abdomen of the male towards the tip and laterally much com- 140 DIPTERA OF NORTH AMERICA. [PART U. pressed, its last segment narrow. The basal portion of the entirely disengaged and inflected hypopygium forms a long peduncle ; the outer appendages are rather filiform and whitish, the inner appendages also. The last segment of the fourth longi- tudinal vein with more or less flexure. The name of the genus (from avv, together, and attvos, narrow) has reference to the very pointed shape of the third joint of the antennae, which is peculiar to both sexes. Gen. XVIII. RHAPHIUM. The first joint of the antennae has no hairs on the upper side ; the second is transverse ; the third glabrous, very narrow, in both sexes very much elongated, though in the male more so than in the female. Arista entirely apical, bare ; its first joint somewhat elongated in the male. Scutellum glabrous. Hypopygium small, rounded, rather imbedded ; its outer appendages more filiform than lamelliform ; the inner appendages small ; the first joint of the hind tarsi without bristles. The genus Rhaphium stands in next relation to the genera Porphyrops and Xipliandrium. "With the species of Porphyrops it shares the larger size, the greater number of bristles on the feet, the broader wings, less narrow towards the basis. With the species of Xiphandrium it has in common the great elongation of the second joint of the antennas, the lesser density of hair on the occiput, and the shorter hair upon the coxas and feet, also that the last segment of the fourth longitudinal vein is less inflected. From both these genera, however, Rhaphium is distinguished by the somewhat longer first joint of the antenna?, which, particularly in the male, is rather swollen ; by the great narrowness and the glabrousness of the third joint, which is uncommonly elongated not only in the male but also in the female ; by the glabrousness of the arista, and finally, by the more n.arrow and more pointed palpi of the female. The other genera most closely approaching Ehaphium, as Systenus, Synarthrus and Smiliotus, cannot be easily confounded with it, the hypopygium of the male in the spe- cies of Systenus being very much pedunculated, the second joint of the antennae in the species of Synarthrus reaching over the third, and the abdomen in Smiliotus having only five segments. RIIAPHIUM — XIPIIANDRIUM. 141 i The name of the genus (from '^lov, small needle) has reference to the shape of the antenna?, which distinguishes it. Only a single European and one North American species, de- scribed below, are known at present. 1. R. lllgllbre LOEW. 9 . — Ex viridi nigrurn, nitens, pedibus nigris, anterioruin tibiis intermediorumque femoribus hiteis. • Greenish-black, shining ; feet black ; the four anterior tibi?e and the middle femora dusky yellow. Long. corp. 0.16. Long. al. 0.16. SYN. Rhaphium lugubre LOEW, Neue Beitr. VIII, 49, 1. Of a bright metallic, but very dark greenish-black color. Face moderately broad, with a bright lustre of silvery-white powder upon black ground. Palpi black with white powder. The black antennae very long, narrow and glabrous ; the apical bristle short and bare. Front shining black. Cilia of the upper orbit black, delicate, short ; cilia of the lateral and inferior orbits white. Thorax bright, with an almost imperceptible gray-whitish dust. The scutellum has no hair with the exception of the usual bristles. Abdomen bright, only on the lateral margin with distinct white powder ; the hair upon it is black. Coxae black with white dust ; the front side of the fore coxae with white hair and black bristles. Anterior femora black with luteous tip ; fore tibia? luteous, only with two bristles on the upper side ; fore tarsi black, the first joint as far as the tip, luteojis. Middle feet luteous, tarsi from the tip of the first joint blackened. Hind feet entirely black, only the knees yellow ; the first joint of the tarsi scarcely a little longer than the second. Cilia of the brown tegulaa whitish. Wings blackish, on the fore margin and along the veins darker ; the last segment of the fourth longitudinal vein only in the middle gently inflected forward. Hob. Carolina. Gen. XIX. XIPHANDRIUM. The genus Xiphandrium comprises small species, of slender form, with little hair. The first joint of the antennae has no hair on the upper side ; the second is of a transverse form, the third rather narrow, in the male very much elongated and in the female much shorter ; the hairs upon it are distinct, especially on the under side. Arista entirely apical, comparatively short, with 1 42 DIPTERA OF NORTH AMERICA. [PART II. scarcely perceptible short pubescence. The lower part of the occiput only with the usual fringe of cilia. Scutelluni without hair. Coxae and feet comparatively bare, the latter with very few bristles ; the first joint of the hind tarsi without bristles. Wings not very broad, somewhat narrowed towards the basis ; the last segment of the fourth longitudinal vein little inflected ; hypo- pygium small, rounded, rather imbedded, its outer appendages more filiform than lamelliform ; the inner appendages small. The next related genera are Rhaphium and Porphyrops. The characters of the genus Rhaphium have already been described. The species of Xiphandrium differ from the species of Porphyrops by their smaller size, more slender form, less hair, especially on the lower part of the occiput and on the coxse ; the third joint of the antennas of the males is more elongated, more distinctly hairy, particularly on the whole lower side ; the arista is beset with a comparatively short but easily perceptible pubescence ; the wings are less broad and towards their basis still narrower. It cannot be mistaken1 for the genera Systenus, Synarthrus and Smiliotus, as they differ from Xiphandrium and Rhaphium by the same distinctive marks. The name (from fi'o$, sword, and oU^p, man) has been given to this genus on account of the sword-shaped antennae of the male. Out of Europe, no species of Xiphandrium are as yet known. I have a female from North America, which probably, belongs to this genus. As it has lost its antennas, its systematical location could not be ascertained. Gen. XX. PORPHYROPS. The genus Porphyrops comprises species of at least middle size, rather stout shape and very hairy. The first joint of the antennas has no hair on the upper side ; the second is transverse ; the third is moderately elongated in the male and shorter in the female ; the hairs upon it are very short and the whole under side (with the exception of a few species) is entirely bare. Arista altogether apical with an almost imperceptible and very short pubescence. The under side of the occiput more or less densely fringed. Scutellum without hairs. Coxae and feet with rather much hair and bristles. First joint of the hind tarsi without bristles. Wings comparatively broad, towards the basis only a PORPHYROPS. 143 little narrower ; the last segment of the fourth longitudinal vein moderately inflected. Hypopygium small, rounded, rather im- bedded, its outer appendages almost in all the species more fili- form than lamelliform ; the outer appendages small. With regard to the mutual relation of the three kindred genera, namely, Ehaphium, Xiphandrium and Porphyrops, all the re- quired information has already been furnished, so that a repetition of their distinctions is not necessary. I have vindicated the name of Porphyrops for this genus in the fifth volume of the Neue Beitriige. An entirely unfounded opposition had been raised against it. The following are the reasons by which I have been governed with relation to the name Porphyrops. Meigen, in the fourth volume of his works, has taken a wider view of the genus Porphyrops and divided it into three sections : 1. With a subapical arista. 2. With an apical arista ; and, 3. With an arista inserted dorsally on the third joint, near the basis. In his seventh volume he adopts the genus Argyra, which M. Macquart had in the meantime established for the first of the three divisions ; the third division he unites with Medeterus ; and for the only remaining second division, which principally contains species of the present genus, he retains the name of Porphyrops ; at the same time he unites with them the species of his genus Ehaphium (that is the genera Ehapliium and Xiphandrium in the sense adopted above). As I cannot agree with this reunion, I am com- pelled to retain the name of Porphyrops for the genus, which embraces most of the species contained in Meigen's second division and this is the present genus. The name Porphyrops (from mop^vpa, scarlet, and pd$, the froth, and av^du, I rob) has reference to the habit of these species to pursue their prey along the shores of a surging sea. Gen. XXIII. THItfOPHILlIS. Character. The face in both sexes broad, not reaching as far as the lower eye-corner, and ending below at an obtuse angle. THINOPHILUS — PEODES. 149 Palpi in both sexes large, reposing upon the proboscis. Antennas very short ; the first joint without hair, the second short, trans- verse, longer above than below ; the third joint circular, the bare arista dorsal. Wings of equal breadth ; the posterior transverse vein is distant from the margin of the wing more than its own length ; the last segment of the fourth longitudinal vein is rather parallel with the third. The abdomen of the male has six seg- ments ; the hypopygium small, rather imbedded, and somewhat turned inside ; its exterior appendages are narrow, parallel lamellae. The abdomen of the female shows five segments. Femora rather strong, the first joint of the hind tarsi without bristles. The species of Tliinophilus are easily distinguished from the species of Diostracus by the absence of hair upon the first joint of the antennas ; from the species of Peodes they differ by the structure of the face and of the hypopygium. They live principally along the shores of the sea, whence their name (from £i'j, down, sand hill, and $0,0 j, friend). As yet only European species have been made known. Gen. XXIV. PEODES. Clmracter. Face not reaching as far as the lower corner of the eye, rather narrow in both sexes, though a little broader in the female, and more enlarged below ; the lower margin is straight. Palpi in the female considerably larger than those of the male and in both sexes reposing upon the proboscis. Antennae very short ; the first joint without hairs ; the second short, transverse ; the third joint rounded ; arista dorsal, only with a short pubes- cence. Wings of uniform breadth ; the posterior transverse vein is removed from the margin of the wing more than its own length ; the last segment of the fourth longitudinal vein parallel with the third. The abdomen of the male has six segments ; the short, stout hypopygium is not imbedded, but disengaged and a little inflected ; its exterior appendages are two small parallel lamella; alongside of each other and with long hair ; the interior appendages form a remarkably large curved forceps. The abdo- men of the female shows five segments. The femora are rather strong ; the first joint of the hind tarsi has no bristles. Peodes is closely related only to Tln'/io/JiUus ; the differences become evident when the characters of both genera are compared. 150 DIPTERA OF NORTH AMERICA. [PART II. The name of the genus (from ttsufys, provided with a large penis) has reference to one of its most striking characters. Only one European species has as yet been made known. Gen. XXV. NEMATOPROCTUS. Character. Antennae short ; first joint without hair ; second joint short, transverse ; third joint small, in the male not larger than in the female, rounded ;. the rather long arista entirely dor- sal. The face, much narrower in the male than in the female, does not reach as far as the inferior corner of the eye. Palpi re- posing upon the proboscis, those of the female much larger than those of the male. Front of equal breadth. ^ The lower part of the occiput distinctly bearded. Eyes very hairy, especially towards the lower corner. Scutellum not hairy. The abdomen of the male has six segments; the small rounded, and a little im- bedded, hypopygium is at its tip ; the exterior appendages are long and filiform, the interior appendages very short and usually not distinctly perceptible. The female abdomen shows five segments. Wings a little narrowed towards the basis, the first longitudinal vein not elongated ; the posterior transverse vein distant from the margin of the wing by more than its own length ; the last segment of the fourth longitudinal vein very gently inflected and towards the end parallel with the third. Feet rather strong ; pulvilli of the fore tarsi not enlarged ; the first joint of the hind tarsi without bristles. The species of Nematoproctus mostly resemble in their habitus the species of Porphyrops ; they differ from them, however, by an altogether different structure of the antenna? ; the third joint of the male being also very short and the position of the arista com- pletely dorsal. They are less closely related to the species of Argyra, to which they were formerly reckoned, as the first joint of the antenna? is without hairs, the third joint of the male is not en- larged, but as small as that of the female, the arista not subapical but entirely dorsal, the first longitudinal vein of the wings not elongated, further, the exterior appendages of the hypopygium have not the shape of short lamellce, but of long threads. The species of Nematoproctus agree with the species of Leuoostola in the glabrousness of the first joint of the antennae : otherwise they LEUCOSTOLA. 151 differ from them precisely in the same manner as from the species Argyra. The name of the genus (from W^o, the thread, and rcpuxto;, the anus) has reference to the thread-like form of the exterior ap- pendages of the hypopygium. The species hitherto known are all European. Gen. XXVI. D.EUCOSTOI.A. The species of Leucostola, like those of Argyra, have usually upon the abdomen, and often also upon the thorax, a thick bril- liant silvery dust, which renders them easy to recognize. The first joint of the antennae is entirely hairless, the second is trans- verse, the third bare, rather large in the male ; the apparently bare and distinctly two-jointed arista is very near the tip of the antenna. Wings broad, the posterior angle rather protruding ; the first longitudinal vein is farther from the margin of the wing than in most of the other genera, and is also much longer than usual ; the fourth longitudinal vein is inflected forward before the middle of its last segment, thence, however, again parallel with the third ; the posterior transverse vein is not approximated to the margin of the wing. The first joint of the hind tarsi has no bristles. Hypopygium small, imbedded ; its exterior appendages are two small narrow lamella?, bent downward ; the interior ap- pendages are of rather simple structure, and often not distinctly visible. The close relationship of the genus Leucostola to that of Argyra can be easily perceived by a comparison of their characters. There is scarcely any difference between them, but that the first joint of the antennas of Leucostola is entirely without any hair, while in Argyra it is distinctly covered with hair. The name of the genus (from teuxos, white, and {to^, dress) has reference to the beautiful silvery lustre which covers the abdomen and sometimes also the thorax of most of these species. The known species belong in part to Europe an/1 in part to America. 152 DIPTERA OF NORTH AMERICA. [PART II. JL. L,. cinglllata LOEW. 0 . — Viridis, Izete splendens, abdoiaine non pollinoso, cingulis flavis. Green, brightly shining, abdomen without wMte dnst and with yellow stripes. Long. corp. 0.19. Long. al. 016. SYN. Leucostola cingulata LOEW, Neue Beitr. VIII, 53, 1. Male. Face very narrow, silvery-white. Palpi snow-white. Proboscis brownish-black. Antennae black ; the third joint black-brown ; the arista inserted close to its tip. Front black with silvery-white dust. Cilia of the upper orbit black, extremely short and delicate ; cilia of the inferior and lateral orbits snow- white. Thorax metallic green, very bright, dusted only on the lateral margin. Scutellum of the same color, without hair, besides the usual bristles. Abdomen without any white dust ; the first segment mostly black ; the second yellowish, transparent, with metallic black border on the posterior margin and with an indis- tinct blackish spot near the middle of the anterior margin ; third segment also yellowish, transparent, and at the basis with a rather broad violet-black transverse stripe, bisinuated on the hind side, and with a narrow blackish-green border on the posterior margin ; the coloring of the fourth segment of the abdomen is the same, but the stripe on the fore-margin and the posterior border are broader, so that the yellow part of the segment has the appear- ance of being divided into two spots ; the fifth segment has no yellow color, but is violet-black near its basis and black-green towards the tip. The same color prevails on the sma.ll hypopy- gium, the extremely small lamella? of which are brownish. The rather long but not very coarse hair upon the abdomen is chiefly black, only upon the fore part of the first segment and upon the venter it is yellow-whitish. COXEB and feet pale-yellowish ; fore coxte with white hair and yellowish-white little bristles. Middle and hind COXOB near the basis a little blackened and also fringed with pale hairs and bristles. Femora slender, the hind ones slightly infuscated at the tip on the upper side. The hair upon the femora is«blackish on the upper side and near the tip, whitish near the basis and on the under side ; the very delicate whitish hair on the under side of the fore and middle femora is rather long-. Fore tibiaa without any bristles ; middle and hind tibiae only with very few delicate and short bristles. Fore and middle tarsi slen- der, but not very long ; the first joint about as long as the others ETJTARSUS. 153 taken together. The first joint of the hind tarsi is shorter than the second and a little stouter, especially towards the tip ; it is fringed with the usual short hair, which is however unusually dense upon the latter part of the under side. Cilia of the black margined tegulse yellowish-white. Wings short and broad ; the posterior transverse vein strikingly far distant from the margin of the wing ; the last segment of the fourth longitudinal vein be- fore its middle gently inflected forward. Sab. District Columbia. (Osten-Sacken.) Observation. — A rather badly preserved female, which I pos- sess, I suppose to be that of.Leucostola cingulata; it shows, how- ever, somendifferences, so that it may belong to some other closely related species. It differs from the above described male by the following characters : The face is comparatively broad and covered with a dense glittering silvery-white powder. The white palpi are larger than those of the male. The third joint of the antennae is very short ; the position of the arista almost apical. The ab- domen is yellow with the exception of the last segment which is of a metallic-green color, shows however, when looked upon in an oblique direction, an indistinct greenish lustre ; its penultimate and antepenultimate incisures are greenish-black towards the lateral margin. The posterior margin of the pleura? is not gray, but yellow. Among the whitish hair of the fore coxae there are a few light- brown bristles, which, in a certain direction, have a black appear- ance. The tip of the hind femora is not darker and the first joint of the hind tarsi is of the usual plain structure, however compara- tively of the same length as that of the described males. Other- wise it coincides with it perfectly. The only character which makes it doubtful whether they belong together is the yellow color on the hind margin of the pleurae in the female. Hob. District Columbia. (Osten-Sackeu.) Gen. XXVII. EUTARSUS. Character. Face narrow, especially in the male ; a little broader upwards. Palpi very small. The first joint of the an- tenna? without hair, the second much overreaching the third ; the third rounded ; the arista dorsal, though rather closely approxi- mated to the tip of the third joint, distinctly two-jointed. The elongated, cylindrical abdomen of the male consists of six seg- 154 DIPTERA OP NORTH AMERICA. [PART II. inents ; the hypopygiurn is very small, entirely imbedded and without bristles ; its appendages are extremely small, almost com- pletely hidden. The female abdomen shows only five segments. Feet rather long, moderately bristly ; the first joint of the hind tarsi without bristles, much shorter than the second ; the pulvilli on the fore tarsi of the male are enlarged, but not elongated. Wings of rather uniform breadth or narrower towards the root ; the first longitudinal vein not elongated ; the last segment of the fourth longitudinal vein though somewhat inflected, still rather parallel with the third. Eutarsus appears to be nearer related to Diapliorus than to any other genus. The striking smalluess of the palpi, the face narrowed below, the hypopygiujn covered with but delicate hairs, the very small and almost completely con- cealed appendages of the hypopygium and the not elongated pul- villi of the fore tarsi in the male distinguish Eutarsus from Diapliorus sufficiently. From Nema^roctus, whose exterior appendages of the hypopygium are long threads covered with hair, Eutarsus is distinguished by the small and concealed ap- pendages of the hypopygium. The species of Saucropus cannot be confounded with the species of Eutarsus on account of the en- tirely disengaged hypopygium. The genus has been founded upon the well known European Eutarsus aulicus Meig. The name (from'tu, handsome, and fapao$, foot) has reference to the structure of the feet of the male. As yet no North American species is known to me, but I know a species from Venezuela, the hind tarsi of the male of which have a remarkable structure, and of which I furnish a description. 1. Et eques, n. sp. £ . — Chalybeus, modice nitens, abdominis ex seneo uigri maculis lateralibus, ventre pedibusque flavis, femoribus pos- ticis supra tibiisque posticis totis fuscis, tarsis anterioribus, prseter basim, posticis totis, nigris, articulo horum primo brevissiino, quarto in aculeuui producto. Steel-blue, moderately shining; lateral spots of the bronze-black abdomen, venter and feet yellow ; upper side of the hind femora and the whole hind tibiaj brown ; the four anterior tarsi with the exception of the root, and the whole hind tarsi black ; the first joint of the latter very short, the fourth joint produced into a point. Long. corp. 0.27. Long. al. 0.26. Male. Front shining steel-blue, not very broad, not excavated on the vertex. Antenna? only of moderate length, black ; the EUTARSUS. 155 narrow and a little elongated first joint is entirely bare on the upper side, on the lower edge of a rather distinctly reddish-yellow color ; the third joint is short ovate ; the arista is inserted upon the back of the third joint, nearer to its basis than it is the case with Eutarsus aulicus. . The face is very narrow immediately below the antennae, and grows more and more so till it becomes cuneiform, so that the large eyes are entirely contiguous on a large extent. Palpi very small ; proboscis rather small. Thorax steel- blue or violet, little shining, around the shoulders more blue-green and more densely covered with dust ; the extreme corner of the shoulder brownish. Scutellum of the same color as the upper side of the thorax and with two bristles. Metathorax and pleurae black-green, the latter with whitish dust and a yellow hind mar- gin. Abdomen cylindrical, blackish bronze-colored ; the second segment has near the basis a narrow yellow transverse stripe, which is somewhat dilated near the lateral margin ; and upon the hind corners a large yellow spot ; there are similar spots, but diminishing in size, upon the other segments. The small hypo- pygium is rounded and rather imbedded ; its appendages are short, indistinct lamellae. The short hair upon the abdomen is black ; longer black bristles only on the hind margin of the first segment. Coxae and feet yellow ; fore coxa? with pale hair, at the tip only with a few thin black little bristles ; middle coxas with a large black spot, which covers about three-quarters of their outer side ; the hind coxae with a small blackish spot. The hind femora on the upper half of their hind side blackish-brown ; fore and mid- dle tibiae only at the extreme tip slightly infusoated ; the hind tibiae totally black-brown. The fore tibiae are without bristles ; the middle and hind tibiae are but sparsely beset with short bris- tles. Fore and middle tarsi from the tip of the first joint black- brown, plain ; the first joint of the fore tarsi is somewhat shorter than the following taken together, the first joint of the middle tarsi longer than the others taken together. The hind tarsi are black, very much shorter than the hind tibiae and of a very peculiar struc- ture ; the first joint is remarkably shortened, the second 4^ the size and the third 2^- the size of the first ; the fourth joint has only the length of the first, its end, however, is produced into a stout thorn, so that with, it this joint is not much shorter than the third ; the fifth joint is not at the end of the fourth, but attached on its under side at the place where the fourth joint begins to be pointed ; 156 DIPTERA OF NORTH AMERICA. [PART II. on both tarsi it is rather singularly at right angles with the fourth joint, is somewhat longer than the third joint and has the form of a slender club. The grayish-hyaline wings are rather long and narrow ; their anal angle is rather rounded off ; the last segment of the fourth longitudinal vein runs upon its middle over a dis- tinct convexity of the wing and is there a little inflected ; towards its end it does not much approach the third longitudinal vein, which is here very gently curved backwards ; the sixth longitudi- nal vein becomes entirely indistinct at a considerable distance from the margin of the wing. Hob. Venezuela. (Moritz.) Observation. — Although the present species differs in some re- spects from Eutarsus aulicus, still it coincides in many important characters with it, so. that it cannot be located into any other genus, unless, rather prematurely, a new genus is created for it. Eutarsus aulicus approaches the forms which prevail in the genus Diaplwrus more than the above described species. Gen. XXVIII. DIAPHORUS. At the time of its adoption, and long after, the genus Diaphorus was considered as one which was very distinct from the genus Chrysotus. The characters which Meigen uses for their dis- tinction are the following : for Chrysotus, arista apical, the eyes of the male contiguous under the antennae, and the wings some- what divaricated when in repose ; for Diaphorus, arista dorsal, eyes of the male contiguous on the front, and the wings reposing upon each other when at rest. The more species of these two genera have become known and the more carefully they have been examined, the more unsatisfac- tory Meigen's characters have been found. North America is particularly rich in species belonging here, but showing important structural deviations. Besides the species showing a different structure of the thorax, and which were formerly classed with Chrysotus, but are united now in the genus Chrysotiimis, the other species of Chrysotus, agreeing in their habitus, are'divided in such, the males of which have eyes contiguous under the antenna;, and in such, where the eyes are separated by the sometimes broad face ; the position of DIAPIIORUS. 157 the arista is likewise by no means always apical ; on the contrary, it is even 'subapical in the majority of the species. The same happens to be the case with Diaphorus, where the eyes of the males are contiguous upon the front in some of the species only, while in others they are separated by the broad front ; the arista likewise is not always distinctly dorsal, but in numy species subapical, and in some truly apical. Thus none of the distinctive marks, which Meigen had established for these two genera, holds good, except for Chrysotus, the wings divaricated in repose, and the parallel wings for Diaphorus. As this charac- ter can only be observed on living specimens, it is, even if proved to be correct, entirely insufficient for a systematical distinction of both genera. In order to escape this difficulty there are two different ways to be followed : either the species, with the eyes of the males not separated upon the front, must remain with Diaphorus, and those the eyes of which, in the male, are contiguous below the antennas, must go with Chrysotus, and a new genus must be established for the species of Diaphorus and Chrysotus, the males of which have the eyes distant above as well as below the antennas. The other way to follow would be to discover characters for the distinction of Chrysotus and Diaphorus better than those which Meigen had chosen. The first of these two alternatives is liable to serious objections. The establishment of three genera would disconnect the relation naturally existing between the insects forming them, a relation based upon their general habitus, and their distinction would con- sist in a character pertaining merely to the male. Therefore nothing remains but to try the other way. If we compare first the species of Chrysotus, in which the eyes of the males are contiguous under the antenna?, w.ith those of Dia- phorus, where the eyes are not separated on the front, we will observe the following distinctions : The structure of the body of the species of Diaphorus is more slender, the abdomen especially is comparatively narrower, and more stretched out ; the hypopy- gium of the male has on the hind side four bristles of rather striking size ; the feet are longer, the pulvilli of the fore tarsi in the male are not only enlarged, but considerably elongated (with the exception of D. nigricans Meig.) ; the wings of Diapho- 158 DIPTERA OF NORTH 'AMERICA. [PART II. rus are comparatively larger, and have a different outline, because the anal angle is more protruding. If, separating the typical species of Diaphorus from the typical species of Chrysotus, we follow out these characters through a series of those species, winch, notwithstanding the separation of their eyes upon the front, are placed into the genus Diapliorus, on account of their general habitus which approaches the typical species of Diaphorus, we find that those among the above men- tioned marks of distinction, which are peculiar only to the males, hold also good among these species ; at least I do not know of any species which, being placed on account of its general habitus among the species of Diaphorus, had not on the pos- terior end of the hypopygium bristles of greater length and remarkable strength, or in which the pulvilli of the fore tarsi of the male were not elongated. It is different, however, with those marks of distinction which belong to both sexes, as also with the more slender form of the body, the greater length of the feet, and the more projecting anal angle of the wings of the species of Diaphorus ; each of these charac- ters gradually fades away from species to species so that, taken singly, these characters are utterly insufficient to decide whether a species belongs to Diapliorus or to Chrysotus. As it happens, however, that where one character decreases, another one becomes more salient, it follows that in their totality they are sufficient to distinguish the females of both genera, with the exception, perhaps, of a few isolated cases. In doubtful cases it will be well to compare the description of the species of both genera. The character of the genus Diaphorus may, therefore, be estab- lished, as follows : Form of the body rather elongated. Eyes of the male never contiguous on the face, in some species sepa- rated upon the front, in others 'not. Antenna? short, the first joint bare, the second transverse, the third short, distinctly hairy, with a dorsal or subapical, rarely with an apical arista. Wings rather large, with strongly projecting anal angle, and thus usually somewhat broader towards the basis ; the posterior transverse vein is either a little beyond, or in, or before the middle of the wing, never close to the margin of the wing ; the last segment of the fourth longitudinal vein is almost straight, or but gently inflected, seldom interrupted, so that its end is thus more approxi- mated to the third longitudinal vein, without, however, converging DIAPHORUS. 159 towards it. Feet rather long, but not very slender ; the first joint of 'the hind tarsi without bristles ; the pulvilli of the fore tarsi elongated in the males of all species ; in the males of some species the same is the case with the pulvilli of the middle tarsi ; in some with the pulvilli of all the tarsi. The hypopygiuni small, imbed- ded, on the posterior end with stouter bristles. The name of the genus (from Sia^opo;, different) signifies nothing more but tBat the species on which this genus was established were remarkably different from the species of previous genera, and is therefore not at all characteristic. The known species of Diaphorus are found in Europe, Asia, Africa and America. Table for the determination of the Species. < Color of the body non-metallic, black. 1 opacus Lw. \ Color of the body metallic-green. 2 ( Feet entirely yellow. 2 mundus Lw. ( Feet not entirely yellow. 3 ( Tegulss with black cilia. 4 I Tegulse with whitish cilia. 6 ( Eyes of the male contiguous. 3 spectabilis Lw. \ Eyes of the male not contiguous. 5 f All the tibiae yellow. 4 sodalis Lw. 5 -I Only the first half of the four anterior tibise yellow. 5 lamellatus, nov. sp. fLast segment of the fourth longitudinal vein not internipted. 6 leucostomus Lw. j Last segment of the fourth longitudinal vein interrupted. 7 interruptus Lw. Systematical arrangement of the Species. I. The eyes of the male contiguous upon the front. 1. opacus Lw. 3. spectabilis Lw. 2. mundus Lw. II. The eyes of the male not contiguous upon the front. A. The last segment of the fourth longitudinal vein not interrupted. 4. sodalis Lwf 6. lamellatus, nov. sp. 5. leucostomus Lw. B. The last segment of the fourth longitudinal vein interrupted. 7. interruptus Lw. 160 DIPTEEA OF NORTH AMERICA. [PART II. Description of the Species. I. THE EYES OF THE MALE CONTIGUOUS UPON THE FRONT. 1. I>. opacus LOEW. £ . — Totus niger, tibiis piceis. Entirely black, tibiae pitch-brown. Long. corp. 0.12. Long. al. 0.12— 0.13. I SYN. Diaphorus opacus LOEW, Neue Beitr. VIII, 56, 1. Male. Entirely black. Face with the palpi and the proboscis black, entirely glabrous. Antenna? black ; third joint small ; posi- tion of the arista more subapical than dorsal. The eyes are com- pletely contiguous on the upper part of the front ; immediately above the antennas a brownish-black, opaque, triangular spot lies between them. Upper side of the thorax and of the scutellum covered with brown dust and opaque. The dust upon the black pleurae is more gray-brown and less distinct. The abdomen shining black, covered with black hair ; the stronger bristles on the posterior part of the hypopygium very striking ; its exterior appendages very small, black ; coxae and femora black and with black hair ; fore and middle femora on the under side with a row of sparse, erect, not very long black hairs ; on the under side of the hind femora there are similar black little hairs, which are less erect and somewhat longer only towards the end. Fore and mid- dle tibiae more yellowish-brown; hind tibise dark-brown. Fore tarsi slender, the first joint as long as the following three together; a great part of the first joint is yellowish-brown, its tip with the rest of the joints black-brown ; pulvilli not very much enlarged and only moderately elongated. Middle tarsi black-brown with yellowish-brown basis ; hind tarsi entirely black-brown. Halteres and tegulaa black ; the cilia of the latter also black. Wings smoky- blackish, towards the anterior margin darker ; they become visi- bly broader towards the basis ; posterior transverse vein but little before the middle of the wing ; the first longitudinal vein reaches almost as far as the middle of the anterior margin and is some- what distant from the latter. Hob. New York. (Osten-Sacken.) Observation 1. — D. opacus is very closely allied to the Euro- pean D. nigricans. As I have only one specimen of the former, I am unable to prove the coincidence of both species in all the DIAPIIORUS. 1C1 plastic characters as fully as it is necessary when an American species is to be recognized as identical with a European species. Neither can I mention any reliable characters for the distinction of both species. I believe that the examination of a larger num- ber of specimens will establish their identity. Observation 2. — I believe I know the female of D. opacus, am however not certain, on account of the smaller length of the first longitudinal vein and the paler coloring of the hind tibia?. The proboscis of this female is remarkably stout and protruding with a flattened tip ; palpi rather large and broad. Face with an almost imperceptible grayish dust, with a distinct transverse swell- ing upon its middle ; front with gray-brown dust and with a trans- verse furrow below its middle ; all the tibia? yellowish-brown, wings tinged with a dusky blackish color, though not so much as in the above described male, and towards the fore margin not much darker ; the first longitudinal vein reaches about as far as the middle between the extreme root of the wings and the end of the second longitudinal vein. All the rest as in the male. Hab. Pennsylvania. 2. D. mundus LOEW. % and 9 • — Laete viridis, pedibus totis flavis. Light metallic green, all the feet yellow. Long. corp. 0.12 — 0.13. Long, al. 0.12—0.13. SYN. Diaphorus mundus LOEW, Neue Beitr. VIII, 57, 2. Light metallic green. The face has a blue-green, but little shining, ground color, which is rather concealed by the distinct white dust ; it is somewhat broader in the female and has a dis- tinct transverse swelling. Palpi yellowish-brown, much larger -in the female and blackened to a considerable extent towards the basis. Proboscis dusky yellow or yellowish-brown. Antennas brownish-yellow ; the small third joint more infuscated ; the arista almost apical ; the eyes of the male meet completely on the upper part of the front, while they are separated directly above the antenna? by a triangular spot of white dust. The front of the female is of entirely uniform breadth, only very little exceeding the breadth of the face, has a blue-green, scarcely a somewhat shining ground color, and is covered with yellowish dust. The cilia of the upper orbit are black and, on account of their brevity, 11 1G2 DIPTERA OF NORTH AMERICA. [PART II. but difficult to perceive ; the cilia of the lateral and inferior orbits are whitish. Upper side of the thorax and of the scutellum pale- green, with metallic lustre, covered with rather thick ochre-yellow dust. Abdomen somewhat darker metallic green, often more gold- green, or coppery-brownish. The bristles at the end of the hypo- pygiurn are not very long ; its exterior appendages brownish and very small. Coxae and feet yellow ; the first two-thirds of the middle coxae blackened, the hind coxae near the basis infuscated. The hair on the feet appears black ; when seen by a reflected light it changes on the tibiae to brownish and on the larger portion of the femora to fallow-yellowish; the root, a part of the under side of the femora, also the fore coxae are distinctly fallow-yellow- ish. The tarsi are scarcely a little infuscated towards their tips, though their last joint is of a dark brown color. In the male the pulvilli of the fore and middle tarsi are considerably enlarged and elongated ; the pulvilli of the hind tarsi are much less so. Halteres and tegulse yellowish with black-brown cilia, which in some directions have a yellowish lustre. Wings grayish-hyaline, on the anterior half usually somewhat yellowish with yellowish- brown veins ; they are rather large and towards the basis a little broader, but their greater breadth is not so near to the posterior margin as in the previous species ; the posterior transverse vein is not nearer to the extreme tip of the wing than it is to its root and the first longitudinal vein reaches scarcely above the first third of the length of the whole wing. Hob. Pennsylvania. 3. D. spectalbilis LOEW. % . — Aeneo-viridis, femoribus nigris, tibiis tarsorumque omnium basi flavis, ciliis tegularum nigris, oculis maris in fronte contiguis. Bronze green, femora black, tibise and the root of all the tarsi yellow, cilia of the tegulse black, the eyes of the male contiguous on the front. Long, corp. 0.13—0.17. Long. al. 0.15—0.16. SYN. Diaphorus spectabilis LOEW, Neue Beitr. VIII, 57, 3. Dark metallic-green, thorax and abdomen more bronze-green, the former sometimes more coppery. Face blue-green with thick white dust, which conceals considerably the ground color. Palpi and proboscis black. Antennas black, small ; arista almost com- pletely apical. The eyes meeting upon the front to a large extent .and separated only by a very small triangular spot, immediately DIAPHORUS. 163 above the antennae. Cilia on the upper orbit black, on the lower and lateral orbits white. Upper side of the thorax and scutellum covered with yellow-brownish dust. The stouter bristles on the posterior end of the very small hypopygium rather striking ; its exterior appendages brown and very small. Coxae black and with whitish dust ; the hair on the fore coxa? appears in most directions black, in others fallow-brownish. Femora black, somewhat with a green reflection ; the tip of the fore and middle femora brownish- yellow ; their black hair, even on the under side, neither of con- siderable length nor density. Tibiae brownish-yellow, rather slender, with a few bristles. Fore tarsi very slender, from the tip of the first joint black-brown, though the basis of the second joint is again paler than the tip of the first ; their first joint is as long as the other four together ; the pulvilli are very much enlarged and elongated. Middle tarsi of an entirely similar structure, of the same color, but their pulvilli are somewhat less enlarged ; hind tarsi from the tip of the first joint black-brown, their pulvilli but very little enlarged ; the yellowish tegulae with blackish cilia, which assume a yellowish glitter in a reflected light. Halteres yellowish, the tip of their knob usually somewhat infuscated. Wings tinged with gray, with blackish-brown veins, along which, in faded specimens, there are blackish-brown margins ; they are broad, though their greatest breadth is not very close to the pos- terior angle ; the space between the third and fourth longitudinal veins is rather wide ; the last segment of the latter shows a very gentle flexure ; posterior transverse vein rather long and somewhat beyond the middle of the wing ; the first longitudinal vein reaches somewhat beyond the first third of the whole length of the wings and is not very far distant from the anterior margin. Hob. District Columbia. (Osten-Sacken.) II. THE EYES OP T.HE MALE NOT CONTIGUOUS UPON THE FRONT. A. The last segment of the fourth longitudinal vein not interrupted. 4. D. sodalis LOEW. £ and 9 • — Aeneo-viridis, femoribus nigris, tibiis omnibus tarsorumque anteriorum basi flavis, tarsis posticis totis ex nigro fuscis, ciliis tegularum nigris, oculis maris in fronte separatis. Bronze-green ; femora black ; all the tibiae and the basis of the four ante- rior tarsi yellow ; all the hind tarsi black-brown ; cilia of the tegulae 164 DIPTERA OF NORTH AMERICA. [PART II. black ; eyes of the male separated upon tlie front. Long. corp. 0.14 — 0.15. Long. al. 0.15. STN. Diaphorus sodalis LOEW, Neue Beitr. VIII, 58, 4. Rather dark metallic-green, the abdomen and sometimes also the thorax more bronze-green, the scutellum in one specimen violet upon its middle. Face of the male of uniform breadth, green-blue with thick white dust ; the face of the female visibly broader, more green, with less dust, below the middle with a transverse swelling. Palpi of the male whitish, only near the extreme basis somewhat blackish ; the palpi of the female much larger, only at the tip dusky-whitish, otherwise blackish. Antennas small, black, with an apical arista. Front of the male of uniform breadth, somewhat exceeding the breadth of the face, blue with yellow- brownish dust, and green and shining upon the vertex. The front of the female is considerably broader, more green and less thickly dusted, the dust has in the vicinity of the antennae a whitish appearance. Cilia of the upper orbit black, those of the lateral and inferior orbits whitish. Thorax distinctly, but not very thickly, dusted. The coarser bristles on the posterior end of the small hypopygium are less striking ; the very small exterior appendages are brownish-black. Coxa3 black with whitish dust ; the fore coxae indistinctly dingy white-yellowish at the tip ; the sparse hair on the front side pale, but the bristles black at the tip. The tro- chanter of the fore and middle feet dusky-yellowish, of the hind feet more brownish. Femora black with green lustre ; their black hair is comparatively short ; only on the under side of the femora of the male, very closely to the tip, a few somewhat longer bristle- like hairs. The tips of the four anterior feihora and the tibias are brownish-yellow, but the tip of the hind tibiae is rather dark- brown. The fore tarsi of the male are slender and elongated, brownish-yellow at the root, then gradually becoming more infus- cated ; their first joint is about as long as the two following taken together ; the pulvilli are but moderately enlarged and elongated. The middle tarsi of the male are of the same color as the fore tarsi and of the same structure, but the first joint is almost as long as the following four taken together, and the pulvilli are not so much enlarged and less elongated. The fore and middle tarsi of the female correspond in color with those of the male, are, how- ever, considerably shorter, have no enlarged pulvilli, and the first DIAPHORUS. 165 joint of the fore tarsi is about equal in length to all the other joints together. The short hind tarsi are in both sexes dark black-brown. Cilia of the white-yellowish tegulie black. Halteres white-yellow- ish. Wings tinged with gray, with black-brown veins, which are margined with dusky in faded specimens ; they are rather large and broad ; their greatest breadth is close before the middle ; the posterior transverse vein lies in the middle between the extreme root and the extreme tip of the wing ; the fourth longitudinal vein is somewhat distant from the third one ; the first longitudinal vein is comparatively close to the margin of the wing and scarcely reaches one-third of the length of the wings. Hob. New York. 5. D. lameliatus, nov. sp. £ . — Aeneo-viridis, pedibus nigris, tibiarum anteriorum diraidio basali fla^o, ciliis tegularum nigris, oculis maris in fronte separatis. Bronze-green ; feet black ; basal half of fhe four anterior tibise yellow ; cilia of the tegulse black ; the eyes of the male separated upon the front. — Long. corp. 0.13. Long. al. 0.14. Dark metallic-green, the color of the scutellum sometimes more blue and that of the abdomen more coppery. Face with thick whitish dust. Palpi small, whitish and fringed with a few black hairs. Antennas small, black ; the arista subapical. Eyes separated ; front much narrower than the face, broader below than above, and covered with thick white dust. Cilia of the upper orbit black, of the lateral and inferior orbits whitish. Thorax, though with dis- tinct, but very thin brownish-yellow dust.' On the posterior end of the hypopygium there are six bristles of remarkable strength. The brownish-black exterior appendages' are of much larger size than in the kindred species ; they are elongated spatule-shaped, very narrow at the root, rounded at the tip and fringed with blackish hairs. Coxas and feet black ; the trochanter of the fore coxae, the extreme tip of the four anterior femora and basal half of the four anterior tibise yellow. The hair upon the feet is black, on the under side of the hind femora elongated and more dense towards their tip. The pulvilli of the fore tarsi are rather un- commonly elongated, while those of the middle tarsi exhibit only a small elongation and those of the hind tarsi none at all. Cilia of the white-yellowish tegulre black. Halteres white-yellowish. Wings gray with brownish-black veins, rather large and broad ; 106 DIPTERA OF NORTH AMERICA. [PART II. they have their greatest breadth somewhat before their middle ; the posterior transverse veitf is in the middle between the extreme root and the tip of the wings ; the first longitudinal vein runs at least as far as the third of the length of the wing. Hob. Middle States. (Osten-Sacken.) • 6. D. leucostomus LOEW. $ and J. — Laete viridis, thorace et scutello interdurn caerulescentibus, tegularum ciliis albidis, ven§, longi- tudinal! quarts, non interrupta. Light green, thorax and scutellum sometimes more blue ; cilia of the teguloe whitish, the fourth longitudinal vein not interrupted. Long. corp. 0.09—0.10. Long. al. 0.12. SYN. Diaphorus leucostomus LOEW, Neue Beitr. VIII, 58, 5. Male. Light green, metallic, shining, thorax and scutellum sometimes sky-blue. Face for a nlale very broad, deepened length- wise, without transverse swelling, so thickly covered with snow- white dust, that the ground-color becomes invisible. Palpi protruding, much larger than in the males of other species ; proboscis very small, black. Antenna black, larger than in other species ; the third joint is particularly distinguished by its more considerable size and is extended at the end into a short point ; arista inserted on the upper side before the tip of this point, but so much bent downward as to be easily mistaken for being apical. Front of uniform breadth, scarcely exceeding that of the face, blue with white dust, which is thicker near the antennae and is almost totally wanting upon the vertex. Cilia of the upper orbit black, those of the lower and lateral orbits whitish. Thorax with thin gray-whitish dust. . The bristles on the posterior margin of the small hypopygium rather long and strong; its outer appen- dages are not distinctly visible. Coxae black, the foremost dusky- whitish at the tip, on the front side rather bright blue-green and fringed with whitish hair, but without black hairs or bristles. Femora metallic blue-green with yellowish tip, with very short hair. Tibia? and tarsi yellowish, the latter towards the end gradu- ally somewhat darker, but only their last joint brown ; fore tibite without strong bristles, middle and hind tibiae with a stronger bristle on the exterior edge of their upper side. and not far from the root ; the hind tibiae, on the exterior edge of the upper side, are sparsely beset with shorter and weaker bristles. Fore tarsi DIAPHORUS. 1C 7 long and slender ; their first joint about as long as the other three together, the pulvilli considerably enlarged and elongated. Middle tarsi like the fore tarsi and of a similar color, but their first joint about as long as the other four together, and the pulvilli not quite so large and not quite so elongated as on the fore tarsi. Hind tarsi more infuscated, only the root of the first joint brown- ish-yellow. Tegulse whitish with whitish cilia. Halteres also whitish. Wings hyaline, but very little tinged with gray ; veins brown-black ; the posterior transverse vein rather exactly in the middle between the root and the tip of the wing ; the first longi- tudinal vein reaches but very little beyond the third part of the length of the wings. Female. Face very little broader than in the male, with thick white powder, although appearing gray on account of the appa- rent dark ground-color ; it is somewhat deepened upon its larger upper part and gently convex upon its smaller lower part ; both parts are divided by an imperfect transverse swelling. Palpi whitish, near the basis somewhat gray. Antennae considerably smaller than in the male, the third joint much smaller, rounded, with an almost imperceptible angle below the insertion of the arista. Front more broad and more green than in the male. Tarsi shorter and usually somewhat more infuscated than those of the male. The pulvilli not enlarged. Hob. Maryland. (Osten-Sacken.) Observation 1. — I believe that I am not mistaken with regard to their belonging together, but I rather preferred to describe them separately: If, contrary to expectation, they should prove as not belonging together, then the name must remain to the male, which I consider as typical. Observation 2. — D. leucostomus approaches in its entire habitus several species which I believe must be referred to Chrysotus more than any other species of the genus Diaphorus, known to me. The elongation into a point of the third joint of the antenna in the male seems to indicate a relationship to Synarthrus barbatus ; neverthe- less the latter differs materially by its narrow, not deepened face, by the conspicuously elongated third joint of the antennas and also by the entirely apical insertion of the arista. 168 DIPTERA OP NORTH AMERICA. [PART II. B. The last segment of the fourth longitudinal vein interrupted. "7. D. iuterruptus LOEW. £ . — Obscure virescens, inodice nitens, femoribus et tibiis concoloribus, genibus testaceis, tarsis fuscis, vena alarum longitudinal! quarta interrupta. Dark green, moderately shining ; femora and tibiae also green ; knees brownish-yellow ; tarsi brown ; the fourth longitudinal vein interrupted. —Long. corp. 0.23. Long. al. 0.20. SYN. Diaphorus interruptus LOEW, Wien. Ent. Monatschr. V, 37, 9. — LOEW, Neue Beitr. VIII, 59, 6. Male. Rather dark green, not very shining. Face of uniform breadth with the front, very broad for a male, covered with thick whitish dust, so that the ground color becomes invisible, moder- ately deepened and without a transverse swelling. Palpi and proboscis black. Antennae black ; their first joint somewhat longer than in other species ; the third joint rounded ; position of the arista distinctly dorsal. Front with thick dusky-whitish powder, so as to conceal the ground color. Cilia of the upper orbit black ; the cilia of the lateral and inferior orbits are whitish and form a considerable fringe. Upper side of the thorax and of the scutel- lum dark-green and dull from grayish dust. Abdomen more shining-green, with extensive but less thick whitish dust and on the anterior part of the segments coppery to a large extent. The four stout bristles on the posterior end of the small and imbedded hypopygium are very prominent. Coxae black ; the fore and middle coxa? on the front side more black-green and fringed with black bristles. Femora metallic green, stout, beset with dense and coarse black hair, on the under side with numerous, but not strong black bristles. Knees yellowish-brown. Tibiae on the under and front side black-brown, on the upper and hind side dark metallic-green, of strong structure and with unusually strong bristles. Tarsi black-brown, the root of the anterior ones and the under side of all the others more brownish-red ; all tarsi are stouter and less elongated than in the other species known to me, also with more hair ; the pulvilli are all very much enlarged and elongated. Tegulaa yellowish with pale-yellowish cilia. Wings hyaline, scarcely a little tinged with gray, alternately with yellow and brown veins ; first longitudinal vein somewhat distant from the margin of the wing and reaching about as far as the middle of the wing ; the third longitudinal vein is very close to the second LYRONEURUS. 1G9 and ends long before the tip of the wing, although its end is very much curved backwards ; the posterior transverse vein is very short and lies much before the middle of the wing, so that the last segment of the fourth longitudinal vein becomes uncommonly long ; the latter diverges very much from the third longitudinal vein, is entirely interrupted upon its second third and the last third, which is remarkable by its slenderuess, is pushed forward towards the third longitudinal vein. Hob. Cuba. (Poey.) Observation. — The interruption of the last segment of the fourth longitudinal vein is particularly striking in this species ; a trace of it is also found in some of the European species. The position and the course of the third longitudinal vein recall the neuration of Lyroneurus. Gen. XXIX. LYRONEURUS. The following are the characters of the genus Lyroneurus: The body is elongated. Eyes upon front and face widely sepa- rated in both sexes. Antennas short ; the first joint not hairy, the second transverse, the third short, rounded, distinctly hairy ; arista apical. Wings very large, at the tip broad and very ob- tuse ; the posterior transverse vein does not approach the margin of the wing ; the third longitudinal vein very close to the second and very much turned backward at the end ; the space between the third and fourth longitudinal veins remarkably broad ; last segment of the fourth longitudinal vein distinctly inflected. Feet rather long, but not very slender ; first joint of the hind tarsi without bristles. Pulvilli of the fore tarsi in the male not elon- gated. Hypopygium small, imbedded, at the posterior end with four strong bristles ; its appendages are very small and hidden. The genus Lyroneurus is by far the next related to the genus Diaphorus. A. more minute examination of the American spe- cies of Diaphorus has satisfied me that this relationship is greater than I supposed, when establishing the genus Lyro- neurus (Wien. Ent. Monatsch. I, 37). The larger size of the wings, which are very broad at the tip, the greater breadth of the space between the third and fourth longitudinal veins, the distinct flexure of the last segment of the fourth longitudinal vein and the not elongated pulvilli of the fore tarsi in the male, these are the 170 DIPTERA OF NORTH AMERICA. [PART II. characters which distinguish Lyroneurus from DiapTiorus. The last of these characters is decisive for the maintenance of the genus Lyroneurus, as the elongation of the pulvilli of the fore tarsi in the male cannot be dispensed with in the character of the genus Diaphorus, without rendering the limit between Diaphorus and Chrysotus entirely uncertain. The genus Lyroneurus, to which also belongs D. adustus Wied., seems to contain only American species; they appear to be particularly numerous in South America. The name of the genus (from xvpa, the lyre, and vsvpov, the nerve) has reference to the lyre-shaped position of the third and fourth longitudinal veins. / 1. L,. caerulescens LOEW. £ . — Viridis, thorace et abdominis dorso cseruleis, femorum apice tibiisque totis testaceis, tarsis ex nigro fuscis. Green, thorax and dorsum of the abdomen sky-blue ; tip of the femora and the whole tibiae brownish-yellow; tarsi black-brown. Long. corp. 0.22. Long. al. 0.25. SYN. Lyroneurus cserulescens LOEW, Wien. Ent. Mon. I, 39. — LOEW, Neue Beitr. VIII, 60, 1. Light metallic-green, most of the upper side of the thorax, of the scutellum and the greater part of the upper side of the abdo- men sky-blue or violet. Face of considerable and uniform breadth and so thickly covered with gray- whitish dust that no trace of the blue-green ground-color is left. Palpi black, with white-gray dust and with strong black hairs. Antenna? black, short. Front of uniform breadth, equal to that of the face ; the dust upon it is so thick that hardly a trace of the ground color is left. Cilia of the upper orbit black, cilia on the lateral and inferior orbits white and forming a rather thick beard. Thorax pale green, upon the middle line and upon the whole hind part sky-blue, or shifting to violet. The rather thick dust on the upper side of the thorax has a gray-brownish tinge. Scutelluin blue or violet with gray-brown dust, on each side with a stronger and with a weaker bristle, on the surface bare. Pleurae green with rather thick gray-white dust. Abdomen cylindrical, metallic-green; its upper side shining blue or violet from the middle of the second segment to the tip. The hair upon the abdomen is black ; the bristles on the hind margin of the single segments are but of moderate length. CIIRYSOTUS. HI The lateral margin of the abdomen shows a distinct gray-whitish dust. The small hypopygiuni is completely imbedded ; it has (as I now perceive on a well-preserved specimen) four strong bristles on the hind margin, like the hypopygiuni of the males of Diapho- rus ; the appendages of the hypopygiuni are extremely small, and completely hidden. Coxae black, somewhat shifting to green, ren- dered gray by a covering of dust ; the fore coxa? at the tip are of a dusky-yellowish color, and beset with black bristles, while on their front side there is some pale hair. Femora green, not strong and fringed with comparatively short black hair. On the four anterior femora the tip to a considerable extent, and on the hind femora only the extreme tip are of a brownish-yellow color. Tibiae brownish-yellow, the tip of the hind tibiae strongly infus- cated, all the tibiae with but few bristles. Tarsi black-brown, the first joint of the middle tarsi up to its first third, that of the fore tarsi up to the middle, yellow-brownish. Fore tarsi but little longer than the fore tibiae, and their first joint not quite so long as the rest ; their pulvilli not enlarged. Tegulae white-yellowish with brownish-black cilia. Wings grayish hyaline with a greasy lustre ; veins brown ; the first longitudinal vein lies close to the margin of the wing and reaches only a little over the first quarter of the length of the wings ; the costa is rather stout, particularly near the end of the second longitudinal vein ; the posterior trans- verse vein is straight and is pretty much in the middle between the root and the tip of the wing. Hob. Mexico. • Gen. XXX. CHRYSOTUS. The genus CJirysotus contains on the average only small spe- cies. The eyes of the males in many species meet upon the front ; in males of other species they are separated. The front becomes broader towards the vertex in most of these species, and more so than is the case with the species of Diapliorus. Antenna? very short ; the first joint without any hair ; the second transverse ; the third rounded, often rather kidney-shaped, sometimes pointed and distinctly hairy ; only in one species, which, on account of the agreement of all the other characters, I have located with Chry- sotus, the third joint of the antennae is considerably longer, almost of the same shape as in most of the species of Argyra. The tvvo- 172 DIPTERA OF NORTH AMERICA. [PART II. jointed arista has an apical or a subapical position. Hypopygium imbedded, distinctly hairy ; its outer appendages have a lamelli- form structure and are usually concealed. Feet rather short and comparatively strong. The first joint of the hind tarsi without bristles. Wings in comparison rather broad and very rounded at the tip ; the small transverse vein in most of the species very far distant from the margin of the wing ; the last segment of the fourth longitudinal vein straight, parallel or almost parallel to the third longitudinal vein. I have already expressed myself in detail (see Diaphorus) about the difficulty attending the distinction of the genera Dia- phorus and Chrysotus. I have pointed out that I consider the smaller size, the less slender form, the comparatively smaller size of the wings, the smaller length of the feet, the not elongated pul- villi of the fore tarsi in the male and the absence of stronger bris- tles on the posterior end of the hypopygium, as those characters, which enable us to distinguish the species of Chrysotus from Diaphorus. The range of the genus Chrysotus is known to be Europe, Asia, Africa, and America. The name of the genus (from xpvour6$, gilded) has reference to the gold-green color of many species. Mr. Say has described three North American species of Chry- sotus ; hardly one of the three probably belongs to this genus, as Mr. Say seems to have misunderstood its characters ; Chrysotus nubilus may be a Hedeterus, C. concinnarius is perhaps a Dia- phorus, and C. abdominalis is probably a Chrysotimus. In the next* place Mr. Macquart has described a female as Chrysotus viridifemora ; if face and front are covered with white dust, as it seems to follow from his description, this character and the color of the feet may perhaps help to recognize the species. Finally, Mr. Walker has, in his usual careless manner, published a Chry- sotus incertus, which probably will remain incertus forever; as he has not even stated the sex of the specimen described, his descrip- tion is of no use whatever. I know, thus far, twelve North American species of Chrysotus, of which five are represented in both sexes, three only in the male, and four in the female sex ; two of the latter offer but so little peculiar characteristics, that I hesitate with their publication, while the two others are striking enough to preclude the possi- bility of a mistake. CIIRYSOTUS. 1 I 3 » Table for the determination of the Species. , ( Third joint of the antennse elongated. 1 cornutus, nov. sp. (. Third joint of the antennse not elongated at all. 2 q J Femora of dark color. I Femora of pale color. 11 „ ( Ground-color of the palpi pale. 4 I Ground-color of the palpi blackish. . ( Palpi white. 2 longimanus Lw. \ Palpi yellow, near the basis blackish. 3 validus Lw. , ( Cilia of the tegulae black. 6 I Cilia of the tegulae pale. 8 fi ( Eyes of the male separated. 6 vividus, nov. sp. 1 Eyes of the male contiguous. 7 f Four anterior tibiae of the male and all the tibiae of the female yellow. 4 obliquus Lw. | Both the fore tibiae of the male and the four anterior tibic-e of the female yellow. 5 affinis Lw. o ( Wings of the male with thickened costa. 9 1 Wings of the male without a thickened costa. 10 Q ( Costa of the male very much incrassated. 7 costalis Lw. \ Costa of the male moderately incrassated. 8 subcostatus, nov. sp. ' Smaller species, with about three bristles on the upper side of the hind tibiae. 9 discolor Lw. Larger species, with about five bristles on the upper side of the hind tibiae. 10 auratus Lw. ,, ( Antennas entirely black. 11 pallipes Lw. 1 First joint of the antennae red. 12 picticornis, nov. sp. Systematical arrangement of the Species. I. Third joint of the antenna elongated. . 1. cornutus, nov. sp. II. Third joint of the antennae not elongated at all. A. Femora of a dark color. A. Ground-color of the palpi pale. 2. longimanus Lw. 3. validus Lw. B. Ground-color of the palpi blackish. 1. Cilia of the tegulae black. 4. obliquus Lw. 6. vividus, nov. sp. 5. affinis Lw. 2. Cilia of the tegulae pale. 114 DIPTERA OF NORTH AMERICA. [PART II. a. Costa of the male incrassated. 7. costalis Lw. 8. subcostatus, nov. sp. b. Costa of the male not incrassated. 9. discolor Lw. 10. auratus Lw. B. Femora of a pale color. 11. pallipes Lw. 12. picticornis, nov. sp. Description of the Species. I. THIRD JOINT OF THE ANTENNA ELONGATED. 1. C. corimtus, nov. sp. £ . — Obscure viridis, paulo nitens, tertio antennarum articulo eloiigato, oculis infra antennis contiguis, tegularum ciliis femoribusque nigris, tibiis testaceis, tarsis anterioribus iude ab articuli primi apice, posticisque totis fuscis. Dark green, little shining ; the last joint of the antennae elongated, eyes meeting below the antennae ; cilia of the tegulse and femora black ; tibiae brownish-yellow ; the four anterior tarsi from the tip of the first joint and all the hind tarsi brown. Long. corp. 0.09. Long. al. 0.09. Dark green, moderately shining. Antennae black, the third joint uncommonly elongated for a Chrysotus, almost of the same shape as in the species of Argyra, only somewhat longer, with a blunt tip ; arista apical. Front black-green. ' Coxaa and femora black, the latter with brownish-yellow tip. Tibiae brownish-yel- low, the hindmost black-brown at the tip and fringed on their upper side with a moderate number of bristles. The four anterior tarsi are black-brown from the tip of the first joint ; the hind tarsi are of a black-brown color. Cilia of the tegula? black. Wings of the usual structure peculiar to the genus Chrysotus, hyaline with a gray tinge ; the small transverse vein is before the middle of the wing and is short.- Hob. Illinois. (Le Baron.) Observation. — This species is distinguished from all other spe- cies of the genus Chrysotus in a striking manner by the unusual elongation of the third joint of the antennas, agrees, however, com- pletely in all other respects. I have hesitated to establish a new genus upon it, as but a single character constitutes its difference from Chrysotus, which character belongs probably only to the male. CHRYSOTUS. 175 II. THE THIRD JOINT OF THE ANTENNAE NOT ELONGATED AT ALL. A. Femora of a dark color. A. Ground color of the palpi pale. 2. C. loilgimanus LOEW. £ . — Viridis, nitens, palpis albis, tarsis anterioribus tenuibus et longis. Green, shining, palpi white, the four anterior tarsi long and slender. Long. corp. 0.14. Long. al. 0.14. SYN. Chrysotus longimanus LOEW, Neue Beitr. VIII, 62, 1. Shining green. Face of uniform, and for a male, rather consi- derable breadth ; the covering of white dust does not entirely con- ceal the green-blue ground color. Palpi white, not very broad, but for a male rather long. The third joint of the antenna? not large, rather kidney-shaped ; the position of the arista rather pre- cisely apical. Front metallic-green, very little dusty and towards the vertex very little enlarged. Upper side of the thorax covered only with a thin brownish-yellow dust. Coxae and femora black with a blue-green metallic lustre, which is more distinct on the femora ; the tip of the fore coxa? yellow, the tip of the posterior coxae dusky-brown. On the anterior feet the tip of the femora, the tibia? and the greatest part of the first joint of the tarsi are yellow ; the remainder of the unusually slender and long fore tarsi is dark brown ; the hind tibite are yellow, but distinctly infuscated at the tip ; the hind tarsi are dark brown. The hair upon the feet is short and the bristles very scarce. Cilia of the tegula? pale. Wings distinctly tinged with gray and with rather black veins. The posterior transverse vein is more distant from the root of the wing than in most of the other species. Hob. Middle States. 3. C. validus LOEW. f . — Aureo-viridis, nitens, palpis flavis, basim versus nigricantibus, pedibus flavis, femoribus posticis excepto apice nigris, viridi-micantibus, feinoribus anticis nigro lituratis. Golden-green, shining, palpi yellow, towards the basis blackish ; feet yel- low, hind femora with the exception of the tip black, with greenish lus- tre, the anterior femora striped with black. Long. corp. 0.13. Long, al. 0.13. SYN. Chrysotus validus LOEW, Neue Beitr. VIII, 63, 2. Golden-green, shining. Face for a female of moderate breadth ; 176 DIPTERA OP NORTH AMERICA. [PART II. its covering with whitish dust does not entirely conceal the blue- green ground-color ; the usual transverse swelling lies, as in most of the other species, usually rather far below its middle ; palpi for a female of moderate size, yellowish and blackish at the root. The third joint of the antenna comparatively not large, rather rounded. Front metallic-green, with thin dust and towards the vertex a lit- tle enlarged. Upper side of the thorax only with thin, brownish- yellow dust. Coxae and hind femora black with metallic-green lustre, which is most distinctly seen on the latter. The tip of the fore coxae yellow ; fore feet yellow ; femora on the upper side with a black longitudinal stripe, the tarsi infuscated from the tip of the first joint ; middle feet entirely yellow ; the tarsi infuscated only from the tip of the first joint ; on the hind feet the extreme tip of the femora, the tibiae and the greater part of the first joint of the tarsi are of a pale color. The hair upon the feet is short and rather delicate, the bristles only few, though the little bristles on the hind tibiae are rather long. The pale hairs on the cilia of the tegulae seem to have,, in some directions, a dark appearance. Wings only slightly tinged with gray, with dark brown veins. The posterior transverse vein approaches but little the root of the wing, and is somewhat farther from it than in most of the other species. Hab. Middle States. (Osten-Sacken.) B. Ground color of the palpi blackish.. 1. Cilia of the tegulae black. 4. C. Ofcliquus LOEW. % and £ • — Viridis, nitens, tegularum ciliis nigris, femoribus nigris viridi-micantibus, summo anteriorum apice flavo. £ . Oculis contiguis, tertio antennarum articulo obliquo, tibiis anteriori- bus flavis. 9 . Tibiis omnibus flavis. Shining-green ; cilia of the tegulse black ; femora black, with green lustre ; the extreme tip of the four anterior femora yellow. $ . The eyes contiguous ; the third joint of the antennae oblique ; the four anterior tibiae yellow. 9 . All the tibiae yellow. Long. corp. 0.09 — 0.10. Long. al. 0.09 — 0.10. SYN. Chrysotus obliquus LOEW, JMeue Beitr. VIII, 63, 3. Male. Eyes completely contiguous upon the face ; the small triangular spot between them, immediately below the antennae, is CII11YSOTUS. 177 covered with brown-gray dust ; the very small palpi black. The third joint of the antennae comparatively with long hair, rather large, oblique, its upper margin much more arched than the lower margin, which is almost straight ; the arista is subapical. Front metallic-green, without a distinct trace of dust and becoming broader upwards. Thorax shining green, sometimes rather gold- green. Coxa? and femora black with metallic-green lustre; on the four anterior feet the extreme tip of the femora, the tibia3 and the greater part of the first joint of the tarsi are yellow, the fol- lowing part of the tarsi black-brown ; the hind tibia? and hind tarsi are decidedly of a brown-black color. The hair upon the feet, though somewhat rough, is short, and even on the hind tibice of very moderate length ; the hind femora have on the under side before the tip but a few bristle-like hairs ; the bristles on the tibise are very scarce ; the pulvilli are very small even on the fore tarsi. Cilia of the tegulae black, though some of them exhibit in a re- flected light a yellow-brownish glitter. Wings somewhat tinged with gray ; veins black ; the posterior transverse vein very close to the root of the wing ; the last segment of the fourth longitu- dinal vein parallel to the third and ending a little before the tip of the wing. Female. Face of moderate breadth with grayish-white dust upon rather black ground ; the usual transverse swelling is far below its middle. Palpi blackish, the third joint of the antennas smaller than that of males, and its oblique form' less striking. The brownish-yellow dust on the upper side of the thorax some- what more dense than in males. The color of the four anterior feet like that of the males ; the hind tibiae yellow with a somewhat dusky tip ; hind tarsi dark In-own, the first joint is sometimes more yellowish-brown near the basis. Wings like those of the male, only the anal angle somewhat more protruding. Hob. New York. (Osten-Sacken.) 'Observation. — I have no ground for doubting that these two sexes belong together, as all those characters which distinguish the male from the female are within the range of the sexual dis- tinctions peculiar to this genus, and the agreement of all the other characters is very striking. 12 178 DIPTERA OF NORTH AMERICA. [PART II. 5. C. affinis LOEW. £ and 9 • — Viridis, nitens, tegularum ciliis nigris, femoribus nigris viridi-uaicantibus, summo anteriorum apice ex flavo piceo. £ . Oculis contiguis, tertio antennarum articulo subobliquo, tibiis anticis ex flavo piceis. 9 . Tibiis anterioribus ex flavo piceis. Shining green, cilia of the tegulae black ; femora black with green lustre ; the extreme tip of the four anterior femora yellow-brownish. £> . Eyes contiguous ; the third joint of the antennae but little oblique ; the fore tibiae yellow-brownish. J. The four anterior tibiae yellow-brownish. Long. corp. 0.09 — 0.10. Long. al. 0.10. SYN. Chrysotus affinis LOEW, Neue Beitr. VIII, 64, 4. Male. It is so much like the male of C. obliquus, that a state- ment of the differences will be sufficient for its recognition. The third joint of the antenna? is visibly smaller and less oblique ; the hind femora have upon the second part of their under side a greater number of bristle-like black hairs ; finally the hair upon the hind tibia? is much longer ; the feet are considerably darker ; fore tibiae more yellowish-brown than yellow and towards their tips distinctly dusky ; middle tibia? often almost brown-black, but always with a yellowish-brown basis ; middle tarsi entirely black- brown ; fore tarsi only near the basis of a pale brown color. I have not discovered any other differences. Female. I have only a single female, which I think belongs here. It resembles the female of the preceding species very much, only .the fore tibia? are more of a brownish-yellow color and dusky towards the tip ; the middle tibia? are still darker than the fore tibia? and the hind tibia? are like those of the male, black. The third joint of^the antenna? is somewhat smaller than in the females of C. obliquus. Hob. Middle States. (Osten-Sacken.) 6. C. Tividus, nov. sp. •£ . — Viridis, nitens, ciliis tegularum nigris, femoribus nigris viridi-micantibus, genibus tibiisque flavis, maris oculis distantibus et costa alarum non incrassata. Green, shining, cilia of the tegulae black, femora black with green lustre, knees and tibiae yellow ; in the male, the eyes separated and the costa not thickened. Long. corp. 0.09. Long. al. 0.09. Metallic-green, bright, but on the upper side of the thorax with a rather thick brownish-yellow dust and therefore more dull. The CHRYSOTUS. 179 eyes are separated by the face, which is comparatively broad for a male ; the face has a rather distinctly impressed middle line, is of a metallic-green color, but opaque on account of the cover of whitish dust. Palpi black, their whitish powder only becomes visible when seen in a very oblique direction. Antennae black, of middle size, their third joint is somewhat longer than in most of the other species, hairy, and of a somewhat irregular form, because that part, where the arista is inserted, is somewhat produced in the shape of a lobe. Front metallic-green, dull on account of brownish-yellow dust. Coxae and femora black, the latter with a green metallic lustre ; the second joint of the fore coxae, the tips of all the femora, all the tibiae and all the tarsi as far as the tip of the first joint, yellow, the end of the feet brownish-black. Hairs and bristles upon the feet very short, the bristles also very few in number. Pulvilli of the fore tarsi rather small. Cilia of the tegula? black. Wings somewhat grayish, with a rather protruding anal angle ; the costa shows no thickening ; the last segment of the fourth longitudinal vein is scarcely a little inflected, parallel with the third and ends immediately before the extreme tip of the wing. Hob. Illinois. (Le Baron.) Observation. — It is not necessary, when identifying this species, to pay too much attention to the shape of the third joint of the antennae, as it sometimes changes its form, especially when recently developed specimens dry up. The separated eyes, the black cilia of the tegulae, the color of the feet, and the costa without a thick- ening, are characters which prevent its being confounded with any other species known to me. * 2. Cilia of the tegulse pale, a. Costa of the male thickened. ?. C. costalis LOEW. £ and $ . — Viridis, polline confertissimo ex fusco cinereo opacus, femoribus nigris, genibus tibiisque flavis, maris oculis distantibus etjnedia costse parte valde incrassata. Green, opaque on account of a very thick brownish-gray dust ; femora black ; knees and tibiae yellow ; the eyes not contiguous in the male and the middle of the costa thickened. Long. corp. 0.09 — 0.10. Long. al. 0.09. STN. Chrysotus costalis LOEW, Neue Beitr. VIII, 64, 5. Male. Ground-color metallic-green, or blue-green, but so thickly covered with brown-gray dust as to conceal this color. 180 DIPTERA OF NORTH AMERICA. [PART II. Face broad for a male, though a little narrower downward ; the dust is of about the same color as the rest of the body, usually, however, somewhat paler. Palpi black, of middle size. The third joint of the antenna small, kidney-shaped, with an entirely apical arista. Front quite opaque on account of its thick dust. The metallic-green ground-color of the upper side of the thorax becomes more visible only when seen from behind. Scutellum and abdomen less thickly covered with dust than the thorax, so that their metallic ground-color becomes more apparent in most direc- tions. The hair upon the abdomen appears, in a reflected light, of a pale-brownish color. Coxa? and femora black, without a dis- tinct green lustre, the second joint of the fore coxae, the extreme tip of all femora, all the tibia? and all the tarsi as far as the tip of the first joint, yellow ; the end of the tarsi dark brown. The hair and bristles upon the feet very short everywhere, the bristles also very scarce ; the pulvilli of the fore tarsi rather small. Cilia of the tegulas pale. Wings somewhat grayish, with a rather protrud- ing anal angle and of more uniform breadth than in most of the other species ; the fore margin of the wings shows a strong black thickening, which commences abruptly at the end of the first lon- gitudinal vein, becomes then gradually thinner and disappears already before the end of the second longitudinal vein ; the last segment of the fourth longitudinal vein is not inflected at all, parallel with the third and ends rather exactly in the extreme tip of the wing ; the posterior transverse vein is almost at an equal distance from the extreme root and from the tip of the wing. Hob. Florida. Female. It resembles the male very much, only the dust upon the whole body is more thick and the green ground-color of the abdomen less bright. The face is not very broad for a female ; its covering of thick dust has the same color as that on the rest of the body ; although the usual transverse swelling lies somewhat below the middle of the face, it is considerably higher than in the females of all the other species known to me, so that the face is divided by it into two almost equal parts. The wings have the same shape of equal breadth as in the male, show however no trace of a thickening on the fore margin. Hob. Maryland. (Osten-Sacken.) CHRYSOTUS. . 181 8. C. Sllbcostatiis, nov. sp. £ . — Viridis, polline raro ex fusco cine- reo aspersus, femoribus nigris, viridi-micantibus, geuibus tibiisque flavis, ruaris oculis distantibus et media parte costje modice incrassata. Green, sparsely covered with, brownish-gray dust ; femora black, with green lustre ; knees and tibiae yellow ; the eyes in the male separated, and the costa moderately thickened. Long. corp. 0.09. Long. al. 0.09. Metallic-green or blue-green, upon the thorax with not very thick brownish-gray dust and therefore less shining. Face for a male rather broad, towards' the bottom somewhat narrower, with very thick brownish-gray dust. Palpi near the root and margin blackish, upon the middle more brownish ; it was not possible to distinguish their color with certainty in the described specimens. Third joint of the antennae very small ; arista apical ; front quite opaque on account of a covering of brownish-gray dust. The scutellum and especially the . abdomen have very little dust and are shining. The hair upon the abdomen is black. Coxce and femora black, the latter with a very bright green metallic lustre ; the second joint of the fore coxse, the tip of all femora, the tibite and all the tarsi as far as the end of the first joint, yellow ; the tip of the tarsi dark brown. The hairs and bristles upon the feet everywhere very short, the bristles at the same time very scarce. Pulvilli of the fore tarsi rather small. Cilia of the tegular whitish. Wings grayish with a rather protruding anal angle ; the anterior margin of the wings shows a not very strong, but distinctly visible thickening, which begins abruptly at the end of the first longitu- dinal vein and thence gradually decreases towards the tip of the wing ; the last segment of the fourth longitudinal vein is scarcely a little inflected, parallel with the third, and ends a little before the extreme tip of the wing ; posterior transverse vein rather equidistant from the root and the tip of the wing. Hob. Illinois. (Le Baron.) Observation. — G. subcostatus is easily distinguished from the male of G. costalis by the smaller antennae, the less thickened costa, the thinner cover of dust, and the green metallic lustre of the femora; from all other species it is 'distinguished by the thickening of the costa. 182 DIPTERA Or NORTH AMERICA. [PART II. I. Costa of the male not thickened. 9. C. discolor LOEW. £ and . — Viridis, nitens, femoribus concolori- bus, genibus, tibiis venisque alarum flavis, maris oculis distantibus et abdomine violaceo. Shining green, also the femora ; knees, tibiae and veins of the wings yel- low ; eyes of the male separated and its abdomen violet. Long. corp. 0.09—0.10. Long. al. 0.10—0.11. SYX. Chrysotus discolor LOEW, Neue Beitr. VIII, 65, 6. • Male. Shining green. The face rather broad for a male, a little narrower towards the bottom, with yellow-grayish or rather white-grayish dust upon green ground. Palpi 1'ather small, black. The third joint of the antennae not very large with an apical arista. Front with rather thick brownish-yellow dust upon green ground. The upper side of the thorax and the scutellum bright golden- green, the posterior end of the former and the latter sometimes more blue-green. ,The brownish-yellow dust on the upper side of the thorax is distinct, but not sufficient to conceal the ground- color. The upper side of the abdomen is bright violet, the basis of the first segment and the lateral margin steel-blue or blue-green. Coxa? black-green. Femora dark metallic-green. The tip of all the femora and the tibia? yellow ; the four anterior tarsi become dusky from the basis so gradually that it is difficult to state where the iufuscation begins ; on the hind tarsi the yellow coloring ex- tends much farther, so that only the last joints exhibit a distinct dusky tinge. The pulvilli are rather large, especially on the fore tarsi. The hairs and bristles upon the feet are everywhere very short, on the tibia? and tarsi yellowish, with the exception of the stronger bristles at the tip of the middle tibia?. The cilia of the tegula? are pale. Wings hyaline, scarcely a little tinged with gray, with luteous veins, which become a little more dark towards the tip of the wing ; the last segment of the fourth longitudinal vein is parallel to the third and ends exactly at the tip of the wing ; the posterior transverse vein lies rather exactly in the mid- dle between the extreme root and the -tip of the wing ; the anal angle of the wings is rather protruding. Female. It differs from the male by the following marks : The face broader, but not too much for a female ; the usual transverse swelling is far below its middle. The third joint of the antenna? somewhat smaller than that of the male. The abdomen golden- CHRYSOTUS. 183 green, without any trace of a violet coloring. The pulvilli of all the tarsi are very small. Hab. Middle States. (Osten-Sacken.) 10. C. auratus LOEW. 9 • — Viridis,thorace et scutello auratis, polline lutescente subopacis, femorum nigrorum apice tibiisque omnibus flavis, facie latiuscula albido-pollinosa, palpis nigris. Green, thorax and scutellum* gilded, somewhat dull on account of a luteous dust ; the tip of the black femora and all the tibise yellow ; the rather broad face covered with white dust ; palpi black. Long. corp. 0.11. Long. al. 0.11. SYN. Chrysotus auratus LOEW, Neue Beitr. VIII, 65, 7. Metallic-green. Face even for a female rather broad, with gray-white dust ; the usual transverse swelling lies very far below its middle. Palpi black. The third joint of the antennse rather large for a female, with comparatively long hair. Front golden- green, rather dull on account of a yellowish dust, only a little broader upwards. The upper side of the thorax greenish-golden, but thickly covered with yellow dust and therefore opaque. Color and dust of the scutellum the same as those of the thorax. Abdo- men of a purer metallic-green and with less dust, near the basis usually more golden-green. Coxas black, the extreme tip of the first joint and the second joint of the fore coxa? yellow, the second joint of the hind coxse yellowish-brown. Femora black with in- distinct green or bronze-colored metallic lustre ; the tip of the four anterior femora is to a larger, and that of the hind femora to a smaller extent, yellow. The tibise and the tarsi have the same color, but the latter, towards their end, gradually become dusky. The hair upon the feet is everywhere very short, and the bristles very scarce. Cilia of the tegulae pale. Wings somewhat grayish with brown veins ; the posterior transverse vein lies about midway between the extreme root and the tip of the wing. Hab. ]XTew York. (Osten-Sacken.) B. Femora of a pale color. 11. C. pallipes LOEW. $ and 9. — Viridis, nitens, coxis anticis pe- dibusque flavis. Shining green, the fore coxae and the feet yellow. Long. corp. <0.09 — 0.10. Long. al. 0.10—0.11. BYN. Chrysotus pallipes LOEW, Neue Beitr. VIII, 66j 8. 184: DIPTERA OP NORTH AMERICA. [PART II. Both sexes resemble each other very much. Metallic-green, shining. The eyes of the male meet almost completely, so that the face appears small, linear ; in the female it is broader and has the usual transverse swelling far below its middle. The dust upon it is white in both sexes. The palpi are rather small and covered with whitish dust so as to conceal the ground-color, which appears to be more yellowish than blackish. The third joint of the an- tennas is not large, even in the males, and but little oblique. Front green, rather dull from whitish dust. The upper side of the thorax has a distinct whitish dust, which however does not conceal the ground-color. The black hair upon the abdomen is somewhat longer than in most of the other species. The whole fore coxas, the tip of the middle and hind coxae and the whole feet are yel- low, even the tarsi are only slightly dusky towards the tip. Cilia of the tegulse pale. Wings a little grayish, with brownish or brown veins ; the posterior transverse vein lies considerably nearer to the root than to the tip of the wing. Hob. Middle States. (Osten-Sacken.) . C. picticorniSj nov. sp. ^ . — Minutus, viridi-aeneus, modice nitens, oculis infra antennas contiguis, priino antennarum articulo rufo, coxis anticis pedibusque flavis. Small, bronze-green, moderately bright ; the eyes contiguous below the an- tennae ; first joint of the antennae red ; fore coxae and the feet yellow. Long. corp. 0.08. Long. al. 0.08. Very small, bronze-green, moderately shining. The eyes con- tiguous -below the antennae. The antennae small, the first joint red, the following two black, the third rather small and a little oblique ; arista apical. Fore coxa? and feet somewhat brownish- yellow. The tip of the hind femora on the upper side and all the tarsi from the tip of the first joint, brown. The hind tibiae are fringed on the upper side only with a few delicate, not very con- spicuous bristles, and show a dusky tinge at the tip. The color of the cilia of the tegulas I cannot positively state. Wings of the usual structure, grayish-hyaline with brownish-black veins ; the posterior transverse vein is rather short and lies before the middle of the surface of the wings. Hob. Illinois. (Le Baron.) Observation. — This description is made only after a single -spe- TEUCHOPHORUS. — SYMPYCNUS. 185 cimen, and will probably require some correction ; the color of the first joint of the antenna? is so characteristic for this species, that there is no probability of its being mistaken for another. Gen. XXXI. TEUCHOPHORU§. • The genus Teuchophorus remains hitherto confined to but a few European species. They rather resemble the small species of Chrysotus, but are easily distinguished from them by the follow- ing characters : Antennas smaller; arista distinctly dorsal. The abdomen of the male somewhat compressed laterally. The pos- terior transverse vein, which is far distant from the margin of the wing, has an extremely steep position, so that its posterior end is farther from the root of the wing than its anterior end. The feet of the male are fringed with isolated, strong, stiff bristles, and its hind tibiae are curved and adorned in various manners. Besides, in all the hitherto known species, the costa of the male is thickened in the same manner as that of the previously described C. costalis. The other characters of the genus coincide with those of Chry- sotus. The name of the genus (from *tvx°s, armor, and lp«, I bear) has probably reference to. the peculiar organs with which the male is provided. Gen. XXXII. §YMI»YCtf*JS. Character. Small, but little shining species, of a rather slender shape. The face is not narrower upwards. Antenna? rather small, in the female shorter than in the male ; the first joint with- out hairs ; the arista is inserted upon the edge of the third joint in the vicinity of its basis. The metathorax is not unusually pro- truding nor elongated. The abdomen of the male is more or less compressed laterally. The hypopygium is small, more or less im- bedded ; its outer appendages small, sometimes not distinctly visi- ble. The fourth longitudinal vein, towards its end, is perceptibly, although only slightly, approximated to the third and very little convergent towards it ; it ends somewhat before or into the tip of the wing ; the posterior transverse vein before or upon the middle of the wing, distant from its margin ; the sixth longitudinal vein becomes indistinct long before it reaches the margin of the wing. 186 DIPTERA OF NORTH AMERICA. [PART IT. The feet are sparely fringed with bristles ; the hind tarsi shorter than the hind tibise and their first joint without bristles. The genus Sympycnus stands in the closest proximity to Anep- sius, is, however, very easily distinguished from it by the glabrous- ness of the first joint of the antennae. Among the genera which have no hair upon the first joint of the antenna, Xanthochlorus, Teuchophorus and Campsicnemus are the next to it ; they differ from Sympycnus by the following characters : 1. Xanthochlorus by the depression on the posterior end of the thorax and the pre- vailing yellow color of the body and of the bristles upon the thorax ; 2. Teuchophorus by the steeper position of the posterior transverse vein, the thickening of the costa in the male, the isolated and strikingly strong bristles upon the feet and the entirely hidden hypopygiuin ; 3. Campsicnemus by the elongated metathorax, by the last segment of the fourth longitudinal vein, which is parallel to the third and ends always beyond the middle of the wings and by the face of the male, which is very narrowed upwards, &c. The name of the genus (from ovp.*vxvos, crowded together) has reference to the crowding together of the ends of the fourth and the third longitudinal veins, whereby Sympycnus is distinguished from Campsicnemus and many other related genera. I am only acquainted with species. from Europe, Africa and North America; the majority of the North American species differ from the European and from the South African species by the fourth longitudinal vein ending exactly into the tip of the wing, while in the other species, this end is distinctly before the tip. As they agree in all other details of organization, there is no ground for a generic separation, but it would be advisable to form of them a group within the genus Sympycnus. The charac- ter of the genus Sympycnus, as hitherto established, requires, with regard to these species, a slight modification, which I have already introduced. Table for the determination of the Species. ! Fourth longitudinal vein ending before the tip of the wing. 1 tertianus, nov. sp. Fourth longitudinal vein ending into the tip itself. Antenna entirely black. v 2 frontalis Lw. Antennae pale near the basis. ( Thorax with dark longitudinal lines. 3 lineatus Lw. \ Thorax without dark longitudinal lines. 4 nodatus Lw. ( \ SYMPYCNUS. 1ST Systematical arrangement of the Species, I. The fourth, longitudinal vein ending before the tip of the wing. 1. tertianus. II. The fourth longitudinal vein ending into the tip itself. 2. frontalis Lw. 4. nodatus Lw. 3. lineatus Lw. Description of the Species. I. THE FOURTH LONGITUDINAL VEIN ENDING BEFORE THE TIP OF THE WING. 1. S. tertianus, nov. sp. £ and 9 • — Ex cinereo virescens, sub- opacus, thorace non lineato, duobus primis antennarum articulis, pal- pis, ventre, coxis pedibusque dilutissime flavicantibus, tarsis inde ab articuli primi apice nigricantibus. £ . Articulo tarsoruna posticoruni tertio abbreviate et prope apicem pilis paulo longioribus hirto. 9 . Pedibus sirnplicibus. Gray-greenish, rather dull ; thorax without aark lines ; the first two joints of the antennae, palpi, venter, coxae and feet pale-yellowish ; the tarsi from the tip of the first joint blackish. £ . The third joint of the hind tarsi shortened and rough on account of some longer hairs near the tip. Long. corp. 0.10. Long. al. 0.10. Dark grayish-green, rather dull. The front seems to be of the same color. Face grayish. Palpi and the first two joints of the antennae white-yellowish ; the third joint blackish, in both sexes rather rounded. Arista in both sexes plain. Tenter whitish-yel- low as far as its tip. The hypopygium, as in the other species of this genus, rounded and semi-imbedded ; its lancet-shaped exterior lamellae small, but distinctly perceptible and of rather dark color. The posterior margin of the pleurae and the coxse white-yellowish ; the four posterior coxae near the basis more or less infuscated. Feet white-yellowish, with black, somewhat scattered hairs, on the middle and the hind tibia? with a few black bristles. The tarsi from the tip of the first joint very much infuscated, the hind tarsi from the same spot almost entirely black. The joints of the fore tarsi of decreasing length in the female, while in the male the third and fourth joints are of about the same length. The joints of the middle tarsi are of decreasing length in both sexes. The first joint of the hind tarsi is, in both sexes, a little shorter than 188 DIPTERA OP NORTH AMERICA. [PART II. the second ; the following joints are, in the female, of decreasing length ; in the male the third joint is somewhat shorter than the fourth, and at its end, on the posterior side, beset with longer black hairs. Wings towards the basis very much narrowed ; the posterior transverse vein is before the middle of the disk of the wing, but rather exactly in the middle between the extreme root and the tip of the wing ; the fourth longitudinal vein ends some- what before the extreme tip- of the wing ; in the female, however, at a very small distance from it. Nab. Sitka. (Sahlberg.) II. THE FOURTH LONGITUDINAL VEIN ENDING INTO THE TIP OF THE WING. 2. S. frontalis LOEW. £ and 9 . — Nigricans, fronte Isete violaceo splendente, antennis totis nigris. Blackish. ; the front bright violet ; the antennae entirely black. Long. corp. 0.11. Long. al. 0.12— 0.13. . Sympycnusfrontalis LOEW, Neue Beitr. VIII, 67, 1. Face in the female of moderate breadth, in the male below very narrow, towards the antennae broader, with white dust, so that the blue ground-color becomes very little visible. Antennae black, larger than in the next following species ; the first joint rather long ; the third joint only with "a very short pubescence, larger and ovate in the male, smaller and rather rounded in the female. Front bright steel-blue or violet. Cilia of the inferior orbit whitish. Upper side of the thorax dull on account of a thick gray-brownish dust, nevertheless the green or blue ground-color is distinctly visible through the dust. The scutellum is of the same color as the upper side of the thorax, and has no hairs besides 'the usual bristles. Abdomen black or greenish-black, the second segment usually with a complete or almost complete yellowish transparent transverse band, the third segment with one, which is interrupted in the middle ; moreover the first and fourth segments are usually yellowish-transparent on the lateral margin. The venter is always white-yellowish. The hypopygiurn, of the same color as the abdomen, is somewhat larger than in the other species of this genus known to me, and but very little im- bedded ; its outer appendages are so small and hidden that I can- not distinctly perceive their shape. The posterior margin of the SYMPYCNUS. 189 pleune, ail the coxae and the feet yellowish. Fore coxae only with pale hairs. The femora have, besides the usual small bristles im- mediately before the tip, no other bristles. The fore 4ibia3 are without bristles,- the middle and hind tibia? with but few bristles. The fore tarsi are more or less ,infiiscated towards their tip ; in the female the joints are of decreasing length, the first nearly as long as the three following together ; in the male, on the contrary, the first joint is extremely shortened and not quite as long as the last one, the second almost as long as the two following together, the third considerably shorter than the fourth joint, which latter is fringed on its upper side with little curved hairs. The joints of the middle tarsi, which towards their end become more and more dusKy, are of decreasing length in the female ; in the male their first joint is considerably longer than the following four joints together, the second as long as the following three together, the third and fourth of almost equal length, but very short, on the anterior side bearded with delicate little fringe-like hairs ; the fifth joint is somewhat more slender than the two preceding joints and almost as long as these taken together. The hind tarsi, from the tip of the first joint, are more or less infuscated, the first joint much shorter than the second, and the following joints of decreas- ing length in both sexes. The tegula? with pale-yellowish cilia. "Wings rather long and narrow, but moderately pointed towards the root, in the female less tinged with gray than in the male ; the fourth longitudinal vein is parallel with the third and ends rather exactly into the tip of the wing ; the posterior transverse vein is perpendicular and lies before the middle of the wing. Hob. Pennsylvania. (Osteu-Sacken.) 3. S. lineatlis LOEW. £ and 9 • — Cinereus, fronte nigra, antennaruni basi, scutelli margine, ventre pedibusque flavis. Gray, front black ; the root of the antennae, the margin of the scutelluin, the venter and the feet yellow. Long. corp. 0.10 — 0.11. Long. al. 0.11—0.12. STN. Sympycnus lineatus LOEW, Neue Beitr. VIII, 67, 2. Brownish-cinereous, opaque. Face covered with whitish dust upon black ground ; in the female it is rather narrow, in the male so much narrowed that the eyes are contiguous on the lower part of the face. Palpi rather blackish. Antennas rather short, the 190 DIPTERA OP NORTH AMERICA. [PART II. third joint smaller than in the next preceding species, in the male elongated-ovate, in the female considerably shorter, in both sexes with a basal arista ; the first joint is always of a yellowish color, the two following paler or darker brown, sometimes rather black- ish. Front black. Cilia of the inferior orbit pale. Upper side of the thorax brownish-cinereous, entirely opaque ; the humeral corner usually brownish-yellow ; on its upper side there are several dark longitudinal lines, the more distinct of which are usually a delicate middle line and two stronger lines alongside of it ; the latter bear the middle rows of bristles. These lines disappear, however, when looked upon from another direction, and are not equally distinct and sharply defined in all specimens. Scutellum on the middle of the upper side gray with a metallic-blue lustre, on the margin yellow. Pleurae gray, their inferior portion more yellow. The metathorax blackish-gray. Abdomen in well colored specimens brownish-gray, in less matured ones more yel- lowish-brown ; the whole venter always pale-yellowish. The small hypopygiuin is mostly shining black and rather imbedded; the outer appendages are larger than usual in the species of this genus and have almost the form of small filiform lamellae ; their color is a dingy yellowish-brown. Coxae and feet yellowish. Femora only with the usual bristles immediately before the tip. Fore tibiae without bristles. Middle and hind tibia? with a moderate number of short black bristles. The first joint of the fore tarsi about as long as the three following, which are of decreasing length, most of the fourth and the whole fifth joint black -brown. Middle tarsi from the tip of the first joint more or less dusky ; their first joint about as long as the four following together, which are of decreasing length. The first joint of the hind tarsi much shorter than the second, the following of decreasing length. Tegulae with brown margin and with pale-yellowish cilia. Wings tinged with gray, in the male with a cuneiform tapering towards the basis ; this is not the case in the females ; the fourth longitu- dinal vein is parallel with the third and ends rather exactly into the tip of the wing ; the posterior transverse vein is perpendicular and lies rather upon the middle of the wing in the female, consi- derably beyond it in the male. Hab. Virginia ; New York. (Osten-Sacken.) SYMPYCNUS. 191 4. S. nodatllS LOEW. £ and 9 . — Cinereus, margine prirni antenna- ruin articuli infero, fascia abdominis basali, niaculisque lateralibus, ven- tre et pedibus flavieautibus, femorum posticorum apice nigro, seta an- tennali maris capituluni minutuni apicale gerente. Gray, the lower margin of the first joint of the antennae, a band near the basis of the abdomen, lateral spots upon it, venter and feet, yellowish ; the tip of the hind femora blackened ; the arista of the male at the tip with a small button. Long. corp. 0.10 — 0.11. Long. al. 0.11 — 0.12. SYN. Sympycnus nodatus LOEW, Berl. Ent. Zeitsch. VI, 215, 68. Face of the male very narrow, upwards a little broader, with white dust ; the face of the female much broader, not perceptibly narrowed below, and much less thickly dusted, so that the black ground-color is distinctly apparent. Antennas of a stouter struc- ture, and even a little larger than in S. frontalis ; the first joint rather long, upon the under side always of a yellowish color, some- times yellow, with the only exception of its upper edge, in which case the yellow coloring extends also on the lower edge of the second joint ; the third joint in the male is broad ovate, in the female rounded. Arista basal, in the female somewhat shorter and plain, in the male longer and enlarged at the tip into a small button. The front is gray, but appears in some directions almost black. Upper side of the thorax brownish-gray, opaque, without distinct longitudinal lines, with black bristles. Scutelluni usually darker than the upper side of the thorax and without hairs, except the usual bristles. Pleura? whitish slate-gray, their posterior margin yellowish. Abdomen of a bronze-black, moderately bright coloring, sometimes with a green or blue metallic lustre ; upon the second segment there is a very broad, usually inter- rupted, transverse baud ; upon the third segment there is also a transverse band, usually interrupted in the middle, and consisting of two yellow lateral spots ; the fourth segment has usually a yel- low spot on each side. The hypopygiurn, being of the same color as the abdomen, is of a similar structure as in S. frontalis ; its small, not easily perceptible, appendages are black. Coxa? and feet yellowish. The fore coxa? are covered only with whitish hair, nevertheless the stronger hairs on the tip assume sometimes a blackish appearance. The femora without any other bristles but those small ones, usually found before the tip ; the hind femora are of a brownish-black color to a rather large extent, though their extreme tip is again paler. The fore tibias have only a single 192 DIPTERA OF NORTH AMERICA. [PART II. small bristle, which is on their upper side, not far from the root ; the middle tibice have only a few bristles ; the hind tibi® have a larger number of bristles, but they are mostly very short ; in the male there is one bristle on the under side, not far from the basis, which is remarkable for its length. The fore tarsi have 1^ the length of the tibiae ; in the male the first joint is very much shortened and not as long as the last one, the second is as long as the three following together, and usually a little blackened on the extreme tip, the three last ones ave black and very little decreasing in length ; the last one with somewhat shorter hairs than the two preceding ; in the female the fore tarsi are gradually of a darker black-brownish color towards the tip, and their joints are of a de- creasing length, the first somewhat longer than the two following, but not quite so long as the three following together. The mid- dle tarsi of the male but little exceed the tibiae in length ; their first joint is of a plain structure, but almost 1^ the length of the other joints and blackened at its tip ; the last four joints are black ; the second is as long as the last three together, gradually enlarged towards its end, and at the end on the outside, in consequence of the greater length of the appressed black pubescence, elongated into a kind of projection ; the third and fourth joints have on the posterior side a few erect crooked hairs. The middle tarsi of the female are plain, scarcely shorter than those of the male, towards the tip gradually of a darker black-brown color ; their joints are of a decreasing length ; the first is longer than the following two, but shorter than the following three together. The hind tarsi are of the same structure in both sexes, shorter than the tibias ; their last four joints black ; the first joint is -scarcely longer than the third, the second at least as long as the third and fourth together. Hal- teres yellowish. Tegulas with a narrow black margin ; their cilia appear yellowish in a reflected light, seen towards the light, however, blackish, in the female even often black. Wings grayish hyaline, in the male somewhat wavy on the posterior margin and towards the basis much more pointed than in the female ; the fourth longi- tudinal vein is parallel to the third and ends precisely at the tip of the wing ; the posterior transverse vein is perpendicular and lies upon the middle of the wing. Hab. Illinois. (Le Baron.) CAMP3ICNEMUS. 193 Gen. XXXIII. CA]»PSIC]VEMU§. Character. The first joint of the antennae without hairs on the upper side ; the third more or less pointed, distinctly hairy ; the arista inserted on its back, near the basis. Face upwards very narrow, especially in the male. The inetathorax is elongated ; the abdomen flattened. The small hypopygium imbedded ; its appendages extremely small. Feet slender ; the first joint of the hind tarsi without bristles. The males are usually remarkable by the peculiar structure of their tibise and often also of the tarsi. The last segment of the fourth longitudinal vein, about its first quarter, runs over a distinct convexity of the wing, is parallel to the third longitudinal vein and ends beyond the tip of the wing. The genus Campsicnemus possesses so many remarkable cha- racters that the species belonging to it cannot either be mistaken or confounded with species of another genus. The name of the genus (from jea/w^tj, the cur-ve, and xvr^tj, the tibia) was given be- cause the males of .many species are distinguished by the peculiar curvature of their middle tibise. The species hitherto known belong to Europe and North Ame- rica. Description of the Species. 1. C. hirtipes LOEW. £ and 9 • — Obscure olivaceus, facie ochracea, alis infuscatis, coxis anticis pedibusque ex testaceo rufis, tarsis prseter basim ex fusco nigris. £ . Primo tibiarum intermediarum dimidio incrassato, tarsisque anticis setas longiores gerentibus. 9- Pedibus simplicibus. Dark olive-brown, face ochre-colored ; wings blackish-gray ; fore coxae and feet brownish-red ; tarsi black-brown with the exception of the root. £ . The thickened basal half of the middle tibise and the fore tarsi beset with long bristles. 9- Feet plain. Long. corp. 0.08. Long. al. 0.13. SYN. Campsicnemus hirtipes LOEW, Neue Beitr. VIII, 68, 1. Metallic olive-brown. Face very narrow, yellowish-brown, with a bright golden-green spot immediately under the antennae, which is not always easily discernible. Antennas entirely black ; their third joint in the male long and pointed, in the female short 13 ' 194 DIPTERA OF NORTH AMERICA. [PART II. and small. Front black with a violet, often very bright lustre, immediately above the antennae with a pale copper-reddish spot. The cilia on the inferior orbit pale. The upper side of the thorax often shows violet reflections. The scutellum shining violet, rarely shining black with an indistinct violet lustre. The color of the abdomen is usually more greenish-black. Fore COXES brown- ish-yellow, near the basis blackened ; middle and hind coxae black with brownish-yellow tip. Feet yellowish-red or brownish-red. The extreme tip of the knees blackish-brown and the tarsi from the tip of the first joint black. Sometimes the upper side of the femora is distinctly infuscated. Cilia of the tegulas black. Wings tinged with blackish-gray and with black veins ; the convexity of the wings lies before the first quarter of the last segment of the fourth longitudinal vein. Male. Its fore femora are thickened as far as their middle, and on the under side, precisely at the end of this thickening, they are densely bearded with stiff little bristles ; the fore tibias are visibly stouter than in the female, beset on the under side with numerous and erect, on the upper side with less numerous and less erect bristles ; fore tarsi with unusually long hairs, especially on their first two joints. The structure of the middle femora is similar to that of the fore femora, though their thickening is less strong and reaches as far as the tip ; the thick beard on the under side, formed of short stiff bristles, is thus brought nearer to their end. The middle tibice are of rather irregular structure ; from the basis to about their middle they are distinctly thickened and fringed on the upper side with a few long black bristles, on the under side they are provided with a small tubercle, beset with short bristles. That part of the middle tibias, which is not thickened, is of a plain struc- ture, though beset on the under side with a row of rather long, black, bristle-like hairs. The middle tarsi and the whole hind feet are of a plain structure. Nab. Pennsylvania. (Osten-Sacken.) 2. C. claudicans, nov. sp. £ and 9 • — Olivaceus, facie ochracea, alis infuscatis, punctum nigrum in ultimo venae quartae segmento geren- tibus, coxis anticis ex testaceo fuscis, pedibus ex rufo testaceis. $ . Tibiis intermediis crassissiinis, varis, supra nigro-spinulosis, tarsis in- termediis totis nigris, articulo prirao crasso, recurve, supra nigro-setoso, tarsis anticis posticisque simplicibus, inde ab articnli primi apice nigris. •J . Pedibus simplicibus, tarsis omnibus inde ab articuli primi apice nigrig. CAMPSICNEMUS. 195 Olive-green ; the face ochre-yellow ; wings infuscated, with a black spot upon the last segment of the fourth longitudinal vein ; fore coxae red- dish-brown ; feet brownish-red. 'J, . Middle tibiae very stout, crooked, on the upper side with small black spines ; the middle tarsi entirely black ; their first joint thickened, curved upwards, on the upper side with black bristles ; fore and hind tarsi plain, from the tip of the first joint black. . All the feet plain ; all the tarsi from the tip of the first joint, black. Long. corp. 0.09. Long. al. 0.14. Olive-green ; thorax more bright than the abdomen, near its fore margin with a few strikingly green reflections. Front black- ish-blue. Antennae entirely black ; the third joint small and not pointed. Face brownish ochre-yellow. Palpi ochre-brownish. Proboscis black. Cilia of the posterior orbit black above, below pale. Fore coxae brownish-red, sometimes yellowish-red, with a white reflection near the root and a large part of the outside brown ; the four posterior coxae grayish-black, the trochanters brownish-black. Feet yellowish-red ; the knees, especially those of the hind feet, more or less infuscated ; all the femora, the fore and hind tibiae, as also the fore and hind tarsi are plain in both sexes, the latter blackened from the tip of the first joint. The middle tibia3 and middle tarsi are only in the female of the same plain structure and of the same color, while they are distinguished in the male by a very different structure ; for its middle tibiae are uncommonly stout and somewhat curved ; the greatest thickness is in their middle ; the last two thirds of the posterior side are excised and provided with a brown stripe ; before this excision, almost on the upper side of the tibia, there is a longitudinal row of black spine- like bristles, which does not occupy, however, the basal third and the apical one-fourth of the tibia ; the second half of the tibia bears upon the other two sides a few long black bristles ; the middle tarsi of the male are entirely black ; their first joint is somewhat curved upwards at the tip, stout and near the extreme basis a little more swollen, on the upper side excised furrow-like and fringed with a row of black bristles ; the following joints are plain, the second not quite so long as the third, and at the extreme basis sometimes of a yellowish-brown color. Wings with a dis- tinct smoky black tinge and with a small blackish spot upon the last segment of the fourth longitudinal vein. Hab. Sitka. (Sahlberg.) 196 DIPTERA OF NORTH AMERICA. [PART II. Gen. XXXIV. PLAGIOlVEtTRUS. This genus has been established by me (in the Wien. Entom. Monatschr. I, 43) on the species described below, and known as yet only in the female sex, but tie extraordinary and peculiar cha- racters of which rendered the establishment of a new genus neces- sary. The whole habitus approaches the species of Gymnopter- nus and Pelastoneurus most, differs, however, from both by the . first joint of the antennae being entirely without hairs and by the posterior transverse vein having an unusually oblique position ; from Gymnopternus it differs moreover by the course of the last segment of the fourth longitudinal vein, which is almost like that of the genus Pelastoneurus. The peculiarities of the female seem to indicate that the hypopygium of the male is disengaged. The establishment of the characters of this genus on so scanty materials presents many difficulties. I believe, however, that the following may be regarded at least as a temporary definition. The first joint of the antennae without hairs, the second not reach- ing thumb-like over the third, on the upper side much longer than on the under side ; the third joint short, without distinct hair and with a dorsal arista. The posterior transverse' vein very oblique ; the last segment of the fourth longitudinal vein runs much forward in its second portion and ends in the vicinity of the third longitudinal vein, so that the first posterior cell becomes very narrow near its end. The first joint of the hind tarsi is without bristles. The name of the genus (from ftxayioj, oblique, and vtrpov, the nerve) has reference to the extraordinary obliqueness of the pos- terior transverse vein. 1. P. univittatlis LOEW. $> — Viridis, thoracis vitta media abdo- minisque fasciis latis purpureis, antennis pedibusque nigris, femoribus virescentibus, genibus tibiarurnque anteriorum basi testaceis, alis cinereis. Green, middle stripe of the thorax and broad bands of the abdomen of a purple color ; antennae and feet black, the knees and the basis of the four anterior tibiae dusky yellow ; wings gray. Long. corp. 0.25. Loug. al. 0.23—0.24. SYN. Plagioneurus univittatus LOEW, Wien. Ent. Mon. I, 43. — LOEW, Neue Beitr. VIII, 69. PLAGIONEURUS. 197 Face not very broad for a female, somewhat elevated, upon its larger upper part with an impressed middle line ; the smaller, con- vex lower part is separated from the upper part by a transverse swelling, incomplete in its middle ; the dense, almost silvery-white dust, conceals a great deal of the ground-color of the face. An- tennae black. The third joint with a short, but sharp tip ; the arista rather strong, with a very short but distinct pubescence. Front bright metallic blue-green. The cilia on the upper orbit black, on the entire lateral and lower orbits white. Thorax me-, tallic-greeu, only on the fore and lateral margin with a little whitish dust, upon the middle with a not very sharply defined longitudinal stripe, which in some directions appears more black, in others more brown and purple, sometimes of a beautiful cinna- mon-brown color. Scutellum metallic-green, with the usual two strong bristles, otherwise bare. On the segments of the abdomen the two first thirds have a dark, the last third a more pale color ; the color of the former part shifts from black, through bronze- brown into a beautiful dark violet ; on the last third the color is chiefly metallic-green, nevertheless it changes on the anterior part into steel-blue, and on the hind margin of the segment into golden- green or- almost a coppery color ; on' the lateral margin the last third of the segments is covered with white dust. Coxa3 black with a rather dusky-green lustre ; the foremost with white dust, clothed with delicate white little hairs, and at the tip with a few black bristles. Feet black ; femora with green lustre ; knees brownish-yellow ; this coloring extends on_the fore tibice as far as the middle, on the middle tibiae as far as the first third, while on the hind feet it is confined to the tip of the knee. The femora have on the under side from the basis almost as far as the tip, erect, but short, whitish hairs ; otherwise their hair is black ; mid- dle and hind femora have on the front side before the tip a few insignificant black bristles. The hair upon the tibiae is altogether black, very short, only on the upper side of the hind tibiae some- what longer, so that its great density is easily perceived ; all the tibiae are beset with short and not very numerous black bristles. Wings tinged with smoky gray, the veins brown-black ; the pos- terior transverse- vein is so very much oblique as to run parallel to the hind margin of the wing ; the last segment of the fourth longitudinal vein approaches the margin of the wing rather closely, without changing its course, then, however, it suddenly turns 198 DIPTER A OF NORTH AMERICA. [PART II. towards the front, so as to end rather far from the tip of the wing in the vicinity of the third longitudinal vein. Hab. Cuba. (Riehl.) Observation. — This species is also found in Brazil. Gen. XXXV. MAWCALUS. . The genus Liancalus shows the closest relationship to the genera Scellus and Hydrophorus. It agrees with them in the following characters : The body in general is beset with neither numerous nor long bristles. Wings elongated ; the posterior transverse vein very closely approximated to the margin of the wing ; feet elongated and slender ; the first joint of the hind tarsi on the upper side without bristles, not shorter than the second, but in 'the majority of the species, longer. Face in both sexes broad, provided with a small tubercle upon the lowest third of each side of the orbit, and with an indistinct swelling running from one tubercle to the other. Antenna? rather short, the first joint without hairs ; the apparently bare arista dorsal, distinctly two- jointed. The hypopygium of the male imbedded. The above mentioned three genera differ sufficiently from the other genera of the Dolichopodidx by the above stated characters, which they have in common. The genus Liancalus in particular, however, differs from Scellus and Hydroplwrus in the following points : 1. All the femora are slender and unarmed, while the genera of Scellus and Hydrojihorus have the femora very much thickened toward the basis, which at least in the males, is armed on the under side ; 2. The segments of the abdomen are beset with bristles before the posterior margin, which is not the case in the species of Scellus and of Hydrophorus. The genus Liancalus contains as yet only three European and one North American species. They form two groups ; in the first the scutellum has only four bristles and the exterior appendages of the hypopygium are more lamelliform, while in the second the scutellum has six bristles and the exterior appendages of the hypopygium are filiform. To the first group belongs Liancalus lacustris Scop, and leucostomus Loew, to the second L. virens Scop, and the following North American species. The name of the genus (from JIEI'OJ, smooth, and avxd^, arm) LIANCALUS. 199 has reference to the unarmed fore feet, by which it is distinguished from the next related genera of Scellus and Hydrophorus. 1. L. gemialis LOEW. £ and 9 • — Viresceus, thoracis lineis quatuor abdominisque fasciis obscuris, pedibus ex nigro viridibus, genibus liavis, alis maris macula apicali nigra, guttam candidam includente, ornatis. Greenish, four lines upon the thorax, and the bands upon the abdomen dark, feet blackish-green with yellow knees ; the tip of the wing in the male with a black spot, which contains a snow-white drop in its centre. Long. corp. 0.26—0.28. Long. al. 0.31. SYN. Liancalus genualis LOEW, Neue Beitr. VIII, 70, 1. Closely related to the European L. virens Scop, and very much like it, though differing from it in the neuration and the picture of the wings, as also by the greater length of the filiform append- ages of the hypopygium. Blue-greenish, somewhat gray from pale dust. Face green or blue with rather whitish dust, which, how- ever does not conceal the ground-color. The large black palpi, fringed with black and comparatively long hair, when seen from the side, usually appear entirely gray-yellowish on account of the dust, with which they are covered. Front green and somewhat spotted with whitish dust. Antenna? entirely black. The cilia of the posterior orbit black above, whitish below. The upper side of the thorax has two narrow, linear longitudinal stripes, separated by a reddish-gray middle line ; these stripes are rather black in fully colored specimens ; there are besides two lateral stripes, the posterior part of which is bifurcated near the trans- verse suture. Thus, not much is left of the beautiful blue-green color, except two broad longitudinal stripes, bearing the stronger bristles. Scutellum with six bristles, as in L. virens. The ab- domen is provided with broad, copper-colored or bronze-brown, sometimes almost black, transverse bands, on the posterior margin of the segments ; on the edge of these bands the ground-color of the abdomen changes often into yellowish-green. Coxae, femora and tibiae metallic black-green ; the knees yellow ; the tarsi black. Fore coxae elongated, cylindrical, on the front side with long whitish hair, at the tip only with a few small black bristles. In the male the second joint of the fore tarsi is uncommonly short- ened and somewhat thickened, so as to be the shortest of all joints, and almost as broad as it is long. Cilia of the tegulce whitish. 200 DIPTERA OP NORTH AMERICA. [PART II. Wings with black veins, hyaline, in the female with but few irre- gular gray spots upon the apical half, in the male moreover near the tip with a few grayish-black longitudinal stripes, and on the tip itself with a black spot, containing near the end of the fourth longitudinal vein anteriorly a round drop, which, -the light falling through it, has a snow-white reflection ; on the anterior margin of this drop, in the black, there is always a small paler spot. The outline of the wings in the male differs considerably from that of the female, being not only narrower, but also sinuated on their whole posterior margin in a peculiar manner. The ex- terior appendages of the short black hypopygium are two very long threads, which reach back almost as far as the basis of the abdomen, and which are beset on their whole length with very long pale hairs. Hab. Middle States. (Osten-Sacken.) Gen. XXXVI. SCELLUS. Character. First joint of the antennae comparatively narrow, bare ; the second short ; the third rounded, elongated only in a hitherto undescribed European species, not excised on the edge. Arista dorsal, apparently bare, distinctly two-jointed. The front on the vertex but little deepened, a little narrower anteriorly. The eyes much higher than broad, encased below by the linear cheeks. Face of middling breadth, very long, reaching somewhat below the lower corner of the eye ; its lowest sharply-edged part is separated from the narrowly-margined eyes by an incision, which turns away from the eyes on its upper end. Palpi recum- bent, of middling and about equal size in both sexes. Proboscis stout. Upper side of the thorax upon its middle with but short bristles. Scutellum flat, with two bristles. Abdomen with- out .bristles and only with scattered and very short hairs. The abdomen of the male has five segments ; the first four are normally developed, while the fifth is usually shortened, often also of a different color ; the following segment is formed by the short, half-imbedded hypopygium. At its lower end there are two small, dark lamellae, directed obliquely downwards, which lie so close together as to present the shap6 of a stout, dentiform pro- jection ; besides these, the comparatively thick penis, bent down- ward and curved, may be seen, but no other appendages. Between SCELLTJS. 201 the fourth and fifth abdominal segments of the male, however, two long, mostly pale-colored tape- or thread-like appendages protrude, which are turned either backward or outward ; their place of in- sertion seems to forbid us to take them for representatives of the ordinary external appendages of the hypopygium ; thus, we are led to regard as such the previously mentioned inferior appendages ; if we do this, thea the representatives of the interior appendages will be wanting, unless these same tape- or thread-like appendages are taken for them. If the point of insertion of the tape-like appendages was really at the place where they first appear on the outside, then the question would be solved, as in such a case they could not be considered as appendages of the hypopygium ; that however, this is not the case, and that they rather originate much further inside, and proceed from there upwards between the fourth and fifth abdominal segments, before they reappear on the surface, can be distinctly seen in many specimens ; to ascertain their true place of insertion requires the anatomical examination of fresh specimens, for which I have no opportunity at present. The form and position of the hypopygium and of its appendages in the males of Scellus has so many peculiar features, that it is difficult to arrive at a conclusion about the true meaning of its diifereut parts. The female abdomen consists of five normally developed segments, followed by one segment more, which is shortened, retracted, and of a different color ; the extreme, somewhat opaque tip of the female abdomen is beset with black bristles. The feet are generally bare, middle and hind feet much longer than the fore feet, and, except the thickening of their femora, which belongs to the males of some species, they are more slender than the fore feet ; fore femora thickened towards the basis, on the under side with nume- rous bristles ; fore tibias on the under side with bristles, elongated at the end into a large tooth, which is still larger in' the males than in the females ; the males have, moreover, a strong spine on the inside, not very far from the basis ; the middle tibia3 of the male are variously decorated with long curly hairs and stiff bris- tles, while those of the female are plain ; the hind tibiae and the feet in both sexes plain, the joints of the latter of decreasing length ; the empodium distinct. Wings long and narrow ; the posterior transverse vein oblique and close to the margin of the wing ; the third and fourth longitudinal veins converging, the 202 DIPTERA OF NORTH AMERICA. [PART II. sixth almost entirely obliterated or existing only as but a short rudiment. In the genus Liancalus we have already mentioned the charac- ters which this genus has in common with Scellus and Hydro- phorus, as also those which distinguish Scellus from Liancalus; to the latter may be added the presence of the two appendages, peculiar to the males of Scellus. The presence of these append- ages also distinguishes the species of Scellus from Hydrophorus, where they are entirely wanting. Moreover the under side of the fore femora and of the fore tibise in the species of Scellus is beset with long spines, catching into each other when the knee is bent, while in the species of Hydrophorus there are at the utmost some spine-like bristles on the under side of the fore femora near the basis, otherwise the under side of the fore femora and of the fore tibiae is only beset with very short thorn-like bristles. The name of the genus (from OXEMOS, with crooked feet) has reference to the peculiar structure of the fore feet. I know as yet only six species of Scellus, of which three are peculiar to North America, one is common to Europe and North America, and two are exclusively European ; one of the latter species, occurring in Sweden, is as yet undescribed. Table for the determination of the Species. ( Wings entirely blackened. 1 exustus Walk. \ Wings not entirely blackened. 2 o ( Apical half of the wings black. 2 spinimanus Ztt. \ Apical half of the wings not black. 3 „ ( Wings entirely tinged with blackish gray. 3 avidus, n. sp. I Wings scarcely a little tinged with gray. 4 filiferus, n. sp. Description of the Species. 1. S. exustus WALK. £ and 9 • — Thoracis dorso seneo-nigro opaco, abdomine cupreo, latera versus viridi, nitidissimo, halteribus nigris, alis nigricantibus adversus costam nigris, lamellis analibus maris albis, in basi nigris, apicem versus flavis, in summo apice puncto nigro notatis. The upper side of the thorax bronze-black, opaque ; the abdomen copper- colored, laterally green, very bright ; halteres black ; wings blackish, towards the fore margin entirely black ; the anal appendages of the male are white, near the root black, towards the tip yellow, at the extreme tip with a black spot. Long. corp. 0.22. Long. al. 0.26. SCELLUS. 203 SYN. Medeterus exustus- WALKEK, Dipt. Saund. 211. Scellits exustus LOEW, Neue Beitr. VIII, 71, 1. Male. Black. The face rather narrow, opaque from a bright ochre-yellow dust. Antennae black. Front covered with white dust. The middle of the upper side of the thorax is, at least in my specimen, black, opaque, and exhibits some traces of gray dust ; towards the lateral margin it is more bright and shows a less distinct coppery reflection ; on the lateral margin itself there is a broad longitudinal stripe covered with white dust. Scutelluni with two bristles, opaque upon the middle, with a thin, almost im- perceptible coat of white dust, bright on the sides. Pleura bronze-black, on the upper half with a dusky copper-colored reflec- tion, on the lower half with a thin gray-whitish dust. Abdomen brilliant coppery-red, in a certain light it appears brass-colored upon the posterior segments, in an oblique direction even green ; its first segment almost reddish-violet. The upper appendages, peculiar to the males of Scellus, are of a very considerable length, white, near the root black, somewhat enlarged at the tip, curved towards each other and of a yellow color, at the extreme tip black and provided with a tuft of pale hairs, which are turned back- wards. Coxa? black, with a thin white-grayish dust, the foremost with extremely short pale hairs, with a few stiff black little hairs and near the tip with.a few black bristles. Feet black, the femora more metallic green-black, with coppery reflections ; the fore femora but short, very much thickened, toward the basis on the whole under side beset with bristles of different length, on the an- terior side with a row of stiff black bristles ; middle femora elon- gated, thin, gently curved, on the under side almost entirely bare ; the hind femora near the basis of the under side are enlarged into a large blunt appendage, beset with large black spines, beyond this appendage there is an arch-like excision ; then again they are stouter and beset on the under side with black bristles. The fore tibiae, which are comparatively stout, bear on the front side, not far from the basis, a stout black thorn, their tip is elongated into a coarse tooth and their under side, which is beset with black bristles, has somewhat before this tooth a small excision ; middle tibiae long and rather slender ; their first half has only three short bristles ; the second is fringed on the front side with a row of short black bristles ; upon the posterior side somewhat beyond the 204 DIPTERA OF NORTH AMERICA. [PART II. middle, there are a few long black bristles, and between these and the tip of the tibia, some long, curly black hairs. The hind tibiae are much stronger than the middle tibia?, their first half is stouter than the second and the front side before the tip is armed with a strong black bristle. Tarsi plain, their joints of decreasing length, the first joint of the middle tarsi with a few bristles. Halteres brownish-black. Wings blackish, all their veins broadly margined with black ; the margins of the costa and of the first four longitu- dinal veins are entirely confluent, so that the anterior part of the wings appears altogether black ; upon the middle of the posterior transverse vein and upon the curve of the last segment of the fourth longitudinal vein there is a black spot ; the costal cell is of a dark brown color. Female. The only female which I possess, is not as well pre- served as the described male, especially the characters of the face cannot be recognized with certainty ; I would therefore recall the circumstance, that the face of most of the females of Scellus is less yellow than in the males. The first joint of the antennae in the female is considerably shorter than that of the male. Fore femora and fore tibia? less stout, though the tip of the latter has also a dentiform, but less stout elongation ; their under side has no excision before this tooth and the front side of fhe tibite no thorn. Middle femora not curved, on. the larger half of their under side with a few sparse bristles. Middle tibia? plain, upon the first half with a considerable number of black bristles, upon the latter part of the posterior side without the curly hairs which are found in the male. Hind femora plain, slender, towards the tip but very little stronger, upon the second part of the under side with about six rather strong b^ack bristles. Hind tibia? without the strong bristle which^ in the males, exists at the tip of the front side. Hob. Middle States. (Osten-Sacken) ; Illinois. (Le Baron.) 2. S. spinimanus ZETT. £ and J . — Thoracis dorso obscure zeneo, albido-pollinoso, opaco, abdomine ex viridi cupreo, iiitido, balteribus subfuscis, alia nigris, basi et costse dimidise limbo subalbidis, margins postico toto cinereo, lamellis analibns maris albis, basim versus infra nigro-marginatis, apicem versus flavis, in surumo apico puncto fusco notatis. Tbe upper side of the thorax dusky bronze-colored, with whitish dust ; SCELLUS. 205 abdomen coppery-green, bright ; halteres brownish ; wings black, though the root and the margin of half the costa is whitish, the whole posterior margin gray ; anal appendages of the male white, towards the basis on the under side with a black margin, towards the tip yellow, at the ex- treme tip marked with a brown spot. Long. corp. 0.15. Long. al. 0.19. STN. Hydrophorus notatus ZETTEKSTEDT, Ins. Lapp. 701, 4, in obs. Hydrophorus spinimanus ZETTEKSTEDT, Dipt. Scaud. II, 445, 5. Male. Blackish bronze-colored. The face is comparatively a little broader than in S. exustus, covered with bright ochre-yellow dust and opaque. Antennas black. Front with whitish dust. The ground-color of the thorax is of a coppery-bronze ; upon the middle of the upper side more of a blackish-bronze, but almost everywhere so thickly covered with dust that the coppery lustre is only very little perceptible ; the dust on the upper side of the thorax is snow-white, upon the two longitudinal stripes near the lateral margin it is less thick, so that the coppery reflection of the ground-color is more distinct ; upon the middle there are two nar- row, dark longitudinal lines, close to each other, which do not reach as far as the posterior margin of the thorax. Upon the pleurte the color of the dust is more yellowish. The scutellum has two bristles, is rather opaque, with a thin whitish dust. Abdo- men green,- mostly with a coppery lustre, which becomes much more bright near the lateral margin. The anal appendages are of middling length, white, near the basis on the lower margin with a narrow black border, and on the upper margin usually marked with a blackish spot ; beyond the middle they are inflected up- wards and gradually assume a yellow color ; their extreme tip is marked with a small brown spot and bears a small tuft of delicate pale hairs, which are turned backwards ; about the middle of the interior margin there is a similar pubescence ; between them, towards the anal region, there is a small tuft of delicate whitish hairs. Coxse bronze-black ; the four anterior with yellow and the two hind ones with a rather whitish dust ; the fore coxae with very short and delicate pale hairs, near and upon the tip with a few black bristles. Feet black, femora and tibise more black-green, the former bright coppery.' Fore femora short, towards the basis very much thickened, beset on the under side with bristles of dif- ferent length, on the front side with a row of stiff black bristles. Middle femora long, stronger than in S. exustus and more curved, on the latter half of the under side with erect black bristles. Hind 206 DIPTERA OF NORTH AMERICA. [PART II. femora of a plain structure, not stouter than the middle femora ; their under side has only close before the tip, a few black bristles. The comparatively stout fore tibiae have on their front side, not far from their basis, a stout black thorn ; their tip is elongated into a very stout tooth, before which the under side of the tibiog, which is beset with strong bristles, has a small excision. Middle tibise not quite so long and slender as those of S. exustus, on the upper side only with three or four short bristles, on the under side with a r,ow of extremely long, straight, erect black bristles, and on the hind side with long curved hairs, which latter are more dense near the tip and curl up to the shape of a lock of hair. Hind tibia3 plain, scarcely stouter than the middle tibia? ; on their upper side, not far from the basis, there is a strong bristle, and on the latter half a few small bristles ; the under side is beset with short small bristles, which are isolated upon the first part and closer together and in more regular order upon the second half; at the end of the under side there is a considerable number of less strong and less short bristles, of which the last is distinguished by its greater length ; on the outside of the tip of the tibiae there are several short and one longer and curved bristle, which has almost the thickness of a thorn. Tarsi plain, the joints of decreasing length, the first joint of the fore and of the middle tarsi with more, that of the hind tarsi with less bristles. Halteres yellowish-brown, the lower part of the knob more dark. The wings of uniform breadth and at the end more rounded than in the other species ; an un- commonly large black spot covers their apical half with the excep- tion of a broad gray border on the posterior margin, and extends as a broad cloud along the fifth longitudinal vein almost as far as the anal cell ; inside of the discoidal cell it is somewhat paler, otherwise, however, so dark that the two black spots, peculiar to this genus, upon the posterior transverse vein and upon the last segment of the fourth longitudinal rein, can only be perceived when the wing is held towards the light ; the anterior part of the wing from the basis as far as the middle is dingy- whitish hyaline ; the anal angle and a broad border along the posterior margin are more hyaline-gray. Hab. Fort Resolution, Hudson's Bay Territory. (Kennicott.) Female. It is distinguished from the male by the following characters : Face with pale yellow-grayish, front with brown dust. The middle of the upper side of the thorax with yellow-brownish SCELLUS. 207 dust, marked with a few spots of whitish dust. Its lateral stripes are covered with brown, the edge of the lateral margin, however, again with gray-whitish dust. The abdomen is more green, less coppery and less bright than in the male. The dust upon the pleurae and upon the fore coxae is less yellow. The fore femora are of a similar structure as those of the male ; the fore tibiae with- out a thorn on the inside, elongated at the tip in a much smaller and sharper tooth, before which there is no excision ; otherwise the feet are plain, the middle and hind femora straight and much more slender than in the male ; middle and hind tibiae only sparely beset with scattered bristles. Observation. — As I do not possess a North American female of &. spinipes, I have prepared the above description from Swedish specimens. 3. S. avidws, nov. sp. £ . — Thoracis dorso seneo-nigro, nitido, mar- giiie et linea media cinereo-pollinosis, pleurarum plaga supera, abdo- mineque ex viridi laete cupreis, nitidissimis, halteribus albis, alis cine- reis, punctis duobus nigris, altero in vena transversa posteriore, altero in ultimo venae longitudinalis quartse segmento ; lamellis analibus maris albis, basi et apice tamen nigris. Upper side of the thorax bronze-black, shining, its margin and a middle line dusted with gray powder ; a large spot on the upper part of the pleurae and the abdomen bright greenish copper-colored, very shining ; halteres white ; wings gray with two large dots, one on the posterior transverse vein, the other upon the last segment of the fourth longitu- dinal vein ; the anal appendages of the male are white, but their basis and tip are black. Long. corp. 0.17. Long. al. 0.21. Male. Face somewhat broader than in the previous two spe- cies, dusted with bright ochre-yellow powder, opaque. Front with white dust. Antennae black. Most of the upper side of the thorax bright bronze-black with faint violet reflections ; its whole margin has a rather broad border dusted -with a whitish-gray pow- der, and therefore opaque ; there is also a narrow middle line, which is much abbreviated behind and likewise dusted with a white-grayish powder. The upper part of the pleurae, from the shoulder to the root of the- wings, is entirely without dust, metallic greenish copper-colored, very much shining ; as I have only a sin- gle specimen, I am unable to judge with certainty whether this large shining spot is also present in fresh specimens, as I believe it to be, or whether in the above described specimen it is merely 208 DIPTERA OF NORTH AMERICA. [PART II. rubbed off ; .the other parts of the pleurae are dusted with gray. The scutellum, which has two bristles, is greenish bronze-colored and opaque. The very shining abdomen is of a bright coppery color, but assumes, when its surface is looked upon in a very ob- lique direction, a green or at least brassy-yellow color. The tape- like anal appendages are very long, somewhat blackened at the basis, and still more so at the tip which is turned upwards, and there beset with a small tuft of blackish hair directed backwards ; otherwise their margins are not hairy, although there is a single black bristle where the blackening of the tip begins on the under side in the vicinity of its inner margin. Near the anus between the above mentioned two appendages there are a few small black hairs. Coxa3 greenish-black, with white-yellowish dust ; the fore- most have besides some pale hairs, almost imperceptible on account of their shortness and delicacy, a few black bristles before and upon the tip. Feet black with a metallic-green reflection, which gra- dually disappears upon the last joints of the tarsi. Fore femora towards their root not so much thickened as in the two preceding species, beset with strong black bristles of different length on the under side, on the front side with a sparse row of short black bris- tles. Middle and hind femora long, slender, straight, of a plain structure, beset on the second half with a moderate number of short, scattered black bristles. The moderately stout fore tibia? bear on their anterior side, not far from the basis, a short black thorn and are prolonged at their tip into a large, somewhat clumsy but sharp tooth, before which the under side of the tibia, provided with bristles, has a very small excision. The middle tibise are long and slender ; on the upper side they are beset with only three, on the anterior side with about seven scattered bristles of very moderate length ; on the other half of their hind side they bear long curly black hairs. Hind tibia? slender, straight, rather long, only on the second half with a few isolated black bristles ; the bristles on the outside of their tip are also only short. Tarsi plain, their joints of decreasing length ; the first joint of the fore and middle tarsi on the under side with numerous black bristles, the first joint of the hind tarsi only with a few and much shorter bristles. Wings hyaline-gray, somewhat darker towards the tip on account of the gray margin of the second, third and fourth longitudinal veins ; upon the posterior transverse vein and upon SCELLUS. 20'.) • the middle of the last segment of the fourth longitudinal vein there is a gray-blackish spot of considerable size. Hab. Fort Resolution, Hudson's Bay Territory. (Kennicott.) 4. S. filiferus, nov. sp. £ . — Thoracis dorso cinereo, opaco, abdomine cupreo, cinereo-pollinoso, subopaco, halteribus albidis, alls hyalinis in basi subalbidis, apicem versus cinereo-striatis punctisque duobus nia- jusculis nigricantibus, altero didymo in vena transversa posteriore, altero simplici in ultimo venae longitudinalis quartse segmento ; lamellis analibus maris augustissimis albis, in basi nigris, in summo apice fla- vicantibus. Upper side of the thorax gray, opaque ; abdomen copper-colored with gray- ish dust, rather opaque ; halteres whitish ; wings hyaline, whitish near the root, with gray stripes towards the tip ; upon the posterior transverse vein with a double blackish spot of considerable size, and with a simi- lar, but single spot upon the last segment of the fourth longitudinal vein ; the anal appendages .of the male are very narrow, white, black at the basis and yellowish at the extreme tip. Long. corp. 0.15. Long, al. 0.20. The narrow face is ochre-yellow. Antennas black. Front with white dust. Thorax blackish bronze-colored with copper-colored reflections ; on the upper side with thick whitish dust, which almost conceals the ground-color, opaque ; upon the pleurae with a somewhat thinner dust of the same color. Upon the middle of the upper side there are two narrow parallel lines of a darker color, which do not reach as far as the posterior margin of the thorax. The scutellurn, which has -two bristles, is of the same color as the pleura?. The ground-color of the abdomen is like • that of the thorax, is, however, more distinct, not being so thickly covered with dust ; towards the lateral margin of the abdomen, where the dust almost entirely disappears, there is a bright cop- per-colored lustre. The anal appendages of the male, which are turned upwards at their tip, are not ribbon-like, as in the pre- viously described three species, but filiform, white, black at the root, at the tip pale-yellowish to a small extent ; on the middle of their exterior margin there is a dense beard of delicate little white hairs ; there is no tuft of hairs at their tip. Between them, in the anal region, only a moderate number of short delicate little hairs, which may be easily overlooked, are inserted. Coxae of a black- ish-bronze color with whitish dust ; the fore coxa? have, besides 14 210 DIPTERA OP NORTH AMERICA. [PART II. the pale, and, on account of its shortness and delicacy, almost im- perceptible pubescence, quite a number of black bristles. Feet black, the femora with a more coppery, the tibia? with a more metallic-greenish tinge. Fore femora rather strongly thickened towards the basis, on the under side with strong black bristles of different length, on the front side only with an imperfect row of short black bristles. Middle femora moderately thickened and strongly curved, on the under side of the basis with two or three not very long, on the apical half with a large number of long, straight and erect bristle-like black hairs. Hind femora plain, of middling stoutness, on the under side with bristle-like short hairs ; on the upper side with two longer black bristles, of which the larger is not far from its tip. Fore tibia? strong, with a large and sharp thorn on the front side, not far from the basis ; elongated at the tip into a large clumsy tooth ; the under side of the tibia, pro- vided with bristles, has no distinct excision before this tooth. Middle tibise long, straight, a little stronger in the neighborhood of the basis ; the first two thirds of their under and posterior side are fringed' with long curly black hairs, the end of the under and front side, however, with long and stiff black bristles ; on the under side, where these bristles begin, there is between them a somewhat curved black thorn. Hind tibia? straight, on the under side with short but very strong black bristles, one of which, being not far from the tip, is remarkable for its greater length ; on the outside of the tip of the tibia? a few shorter and one somewhat longer curved bristle are inserted. Tarsi plain, their joints of decreasing length ; the first joint of the fore and middle tarsi on the under side with black bristles, which are of considerable length near the basis of the tarsi, but decrease in length very rapidly so as to be very short on the larger portion of the joint ; the under side of the first joint of the hind tarsi is everywhere beset with short black bristles. Halteres pale-yellowish, but the basis of the knob some- what brownish. "Wings hyaline ; almost the whole basal third appears somewhat whitish when seen in a certain direction ; gray- ish stripes along the second half of the second and third longitu- dinal veins, likewise along the tip of the last segment of the fourth and the greater portion of the fifth longitudinal vein ; there is also a gray streak between the third and fourth longitudinal veins ; upon the posterior transverse vein there is a blackish-gray double spot, and upon the middle of the last segment of the fourth longi- HYDROPHORUS. 211 tudinal Vein there is a larger rounded blackish-gray spot, which is less sharply denned on the side turned towards the tip of the wing. Hob. Fort Resolution, Hudson's Bay Territory. (Kennicott.) • Gen. XXXVII. HYDROPHORUS. Character. The first joint of the antennae short, bare ; the second very short ; the third rounded, more or less distinctly notched under its tip ; arista dorsal, two-jointed, apparently bare. Front a little excavated on the vertex, narrower anteriorly. Eyes large, higher than broad, encased below in the cheeks, which in their structure resemble those of Orthochile, and are broader than in the other genera. The face reaches somewhat below the lower corner of the eyes ; it is scarcely narrower in the male than in the female, its two upper thirds are usually a little concave ; the lowest third is separated from the upper one by two knotlike elevations near the eyes, is convex and ends in a sharp rounded edge. Pro- boscis of moderate thickness ; palpi incumbent, in both sexes rather small. Upper side of the thorax only with moderately long hairs upon its middle. Scutellurn rather flat, though some- what elevated along its middle line, in all species known to me with four bristles. Abdomen broad and rather flat, very short, without bristles ; it has five segments in both sexes. The hypopy- giurn of the male is small and imbedded ; at its lower end there are two small, dark-colored lamellae, obliquely turned down, and so closely approximated that they seem to form but one clumsy dentiform protuberance ; usually no other appendages are seen besides these. Feet generally bare ; middle and hind feet much longer and more slender than the fore feet ; fore femora gradually rather thickened towards the root ; on their under side, either in both sexes or at least in the male, with short, thornlike bristles ; sometimes they are beset with a few longer and thornlike bristles ; fore tibiae on the under side densely beset with very short thorn- like bristles ; middle tibiae never beset with longer hairs even in the male ; tarsi plain, their joints very much decreasing in length; the empodium very distinct. Wings long and narrow ; the pos- terior transverse vein close to the margin of the wing ; the third and fourth longitudinal veins usually somewhat converging towards their ends ; in some species they are parallel ; the sixth longitudi- nal vein usually apparent as far as the middle of its course. 212 DIPTERA OF NORTH AMERICA. [PART II. Hydrophorus differs from Scellus by the fore femora and fore tibije not being provided on the under side with long thorns, catching between each other, by the tip of the fore tibiae not being elongated into a clumsy projection, by the middle tibia3 of the male not being adorned with long hair, by the hypopygium not having those peculiar pale-colored appendages, which distinguish the species of Scellus, &c. Liancalus has no notches on the third joint of the antennae, its fore femora are not thickened and un- armed, and it has bristles before the incisures of the abdomen, so that no mistake can occur between the species of Hydrophones and Liancalus. The name of Hydrophorus (from i'Sup, water, and qipftv, to carry) has been bestowed upon these insects with reference to the ability of many of the species to run even upon agitated waters. The species of Hydrophorus known to me are distributed over Europe, Northern Asia, Africa and North America. There is no perceptible habitual difference between the species from the dif- ferent parts of the globe. Table for the determination of the Species. , ( Face with white dust, shining green above. 1 innotatus, nov. sp. ( Face with brownish-yellow dust, opaque above. 2 ~ ( Third and fourth longitudinal veins convergent. 2 pirata Lw. \ Third and fourth longitudinal veins parallel. 3 parvus Lw. Description of the Species. 1. H. innotatus, nov. sp. £ and J. — Olivaceo-jeneus, scutello virescente, abdomine viridi, femoribus tibiisque viridibus, tarsis nigris, facie supra viridi-splendente, polline pleurarum albido, venis alarum cinerascentium atris, longitudinalibus tertia et quarta apicem versus paulo couvergentibus. Metallic olive-brown with greenish scutellum and green abdomen ; femora and tibiae green ; tarsi black ; face shining green above ; pleurse with whitish dust ; wings gray with black veins ; the third and fourth longi- tudinal veins somewhat converging towards their end. Long. corp. 0.13. Long. al. 0.22. The dust upon the face is yellow-brownish immediately below the antenna? ; otherwise everywhere white in the male, but so thin upon the entire upper part of the face that its metallic-green color becomes distinctly perceptible ; in the female the face is covered with white dust only along the sides, upon the middle with HYDROPHORUS. 213 brownish dust. Antennas entirely black. Front greenish-black, opaque ; seen in an oblique direction the covering of brown dust upon it becomes perceptible, which otherwise is visible only on the anterior margin. The ground-color of the occiput is green and but little covered with brownish-gray dust. The cilia of the upper orbit are, as usual, black, the dense and hair-like cilia on the late- ral and inferior orbits, however, rusty-yellowish. The upper side of the thorax metallic olive-brown ; the hindmost part with a metallic-green reflection ; the dust upon it is brown. Scutellum with four bristles, shining, rather green, though somewhat copper- colored upon its middle. Pleuras and coxse with whitish dust. Abdomen metallic-green, shining, especially on the sides ; the short hair upon it is blackish, upon the sides of the first segment, however, fallow-yellowish. Fore coxse on their front side with a very short and delicate whitish pubescence, beset on the upper half of their exterior side with a moderate number of compara- tively long black bristles ; a few of them are also at the tip. Femora slender, green, with an almost imperceptible grayish dust ; the fore femora as usual thickened towards their basis, and beset on the under side near the basis with four or five rather long thornlike bristles ; besides these there is, nearer to the anterior margin, an apparently incomplete row of very short bristle-like little hairs, which are but difficult to perceive. Tibia? dark green, the foremost on their under side uniformly fringed with very short, black thornlike bristles. Tarsi black. Cilia of the tegulas yel- low. Halteres with a dusky yellow peduncle and with blackened knob. Wings very long, tinged with gray, not darker towards the anterior margin and with veins which are black up to the ex- treme root ; the end of the third longitudinal vein is somewhat curved backwards so as to converge distinctly towards the fourth longitudinal vein ; no dark spot is to be seen neither upon the fourth longitudinal vein, nor upon the convexity, crossed by the last segment of the fourth longitudinal vein. Hab. Sitka. (Sahlberg.) Observation. — Notwithstanding the not unimportant difference in the coloring of the dust on the lower part of the face between the two sexes described above, I have no doubt that both belong together. From the other North American species, known to me, H. innotatun differs by the shining green color of the upper part of the face. Among the European species it can only be 214 DIPTERA OF NORTH AMERICA. [PART II. compared to balticus Meig., alpinus Wahlb. and callostomus Lw. It can easily be distinguished from balticus by the dark knob of the halteres ; from alpinus by the want of a curved thorn at the end of the fore tibiae ; from callostomus by its more con- siderable size and a comparatively narrower face. 2. H. pirata LOEW. £o$, yellow, and z*-"P°s, green) has reference to the peculiar color of the species. Besides a few European species, only the following North 224 DIPTERA OF NORTH AMERICA. [PART II. American species is known, which may perhaps be identical with one of the European species. 1. X. lielvinus LOEW. £ . — Flavus, fronte, facie et setis thoracis nigricantibus. Yellow, front, face and the bristles of the thorax blackish. Long. corp. 0.11. Long. al. 0.13. STN. Xanthochlorus helvinus LOEW, Neue Beitr. VII, 75, 1. Altogether yellow. Front and face blackish with whitish dust, which is more distinctly visible in an oblique direction. The arista, the bristles on the vertex and on the upper side of the thorax, black, but shifting into brown in a reflected light. A small blackish spot on the pleura?, immediately below the root of the wing. Hab. Chicago! Observation. — This species resembles the European X. tenellus Wied., very much, and is probably a slight variety of it. No plastic distinctions at all are to be seen ; the only difference which I can perceive, is the darker color of the bristles upon the vertex and on the upper side of the thorax. The comparison of the male is necessary in order to determine whether this sj>ecies can be con- sidered as a variety of X. tenellus. Gen. XLII. SAUCROPUS. The species of this genus, which older authors have united with Porphyrops, agree in many characters so much, and differ by these characters so sharply from the related species of the Doli- chopodidse, that their claim to form a separate genus cannot be questioned. The following are the principal characters of this genus : First joint of the antenna? without hair on the upper side ; arista dorsal. The thorax with a sloping surface upon the mid- dle of its posterior end. Feet very long and slender ; hind tibiae elongated ; the first joint of the hind tarsi without bristles, shorter than the second. Abdomen elongated and narrow ; especially in the male. Hypopygium disengaged, short and stout, inflected, with short, very little developed appendages. Color of the body principally, or at least partially, yellow. Hairs and bristles chiefly black. SAUCROPUS. 225 The characters by which the genus Saucropus is distinguished from the related genera, need not be repeated here, as they have already been sufficiently explained among the characters of those genera. The name- of JSaucropus (from aavxp6$, delicate, and rtov;, foot) has reference to the great sleuderness of the feet, which distin- guishes all the species of this genus. The known species are distributed over Europe, America, and South Africa. The American species resemble more those of Europe than those of South Africa ; in the latter the third and fourth longitudinal veins are parallel, while in the European species the last segment of the fourth longitudinal vein approaches the third longitudinal vein, although very gradually, still, in gene- ral, very strongly. li S. dimidiatus LOEW. £ . — Pallide flavus, thoracis linea media et macula postica nigris, abdomine nigrofasciato, setis -coxarum anteriorum albidis. Pale-yellow, thorax with a black middle line and with a black spot upon the posterior end, abdomen with black bands ; the anterior coxae with whitish bristles. Long. corp. 0.18. Long. al. 0.17. SYN. Saucropus dimicliatus LOEW, Neue Beitr. VIII, 75, 1. Pale-yellowish. Face very narrow, white. Antenna? bright- yellow. Front black with white dust ; frontal bristles black. Cilia on the posterior orbit as well as the two stronger bristles behind the upper corners of the eyes, yellow-whitish. Upper side of the thorax reddish-yellow ; the middle line and the sloping sur- face on its posterior margin are black. Scutellum whitish-yellow with two strong black bristles ; on the outside of each of these bristles there is a very small delicate hair, easily overlooked, which in all our European species is much stronger. Metathorax black ; the pleura? have a small black dot above the middle coxa?. Abdo- men with three black transverse bands, the first in the vicinity of the basis of the second segment, the two following near the basis of the third and fourth segments ; the two last are somewhat emarginated on the middle of their posterior margin. Hypopy- gium short and clumsy, shining black ; its small exterior appen- dages are white. Coxa? and feet pale-yellowish ; on the fore and middle coxa? there are altogether no black, but only whitish bris- 15 226 DIPTERA OF NORTH AMERICA. [PART II. ties ; the hairs on the front side of the fore coxae are also whitish ; on the outside of the hind coxa? there is a solitary black bristle. Tibias and tarsi elongated and very slender; middle and hind tarsi strongly infuscated from the tip of the first joint ; the fore tarsi from the tip of the first joint become likewise more dark, though their color changes much more gradually into brown. Cilia of the tegulae whitish. Wings hyaline with a slight gray-yellowish tinge ; the end of the fourth longitudinal vein approaches rather strongly the end of the third. Hob. Florida, District of Columbia. (Osten-Sacken.) 2. S. ruTiellllS LOEW. £ • — Pallide flavus, thoracis macula postica, abdomiuis fasciis setisque coxarum nigris. Pale-yellow ; thorax with a black spot upon the posterior side ; abdomen with black bauds ; coxae with black bristles. Long. corp. 0.25. Long, al. 0.23. SYN. Saucropus rubellus LOEW, Neue Beitr. VIII, 76, 2. Pale-yellowish. Face narrow, white. Antennse bright-yellow. Front black with white-yellowish dust. The frontal bristles and the two stronger bristles behind the upper corners of the eyes, black. Cilia of the orbit yellowish-white. Upper side of the thorax reddish-yellow, the sloping spot on its hind margin black. Scutellum yellowish with two strong black bristles ; on the outside of each there is a very minute delicate hair, easily overlooked. Metathorax brown only on the upper margin and upon the. mid- dle line. Pleura? with a black dot above the middle coxae. Ab- domen with four black transverse bands of uniform breadth, of which the first near the anterior margin of the second segment, the others on the anterior margins of the following segments ; the last of these bands is sometimes indistinct. Coxse and feet pale- yellowish. The bristles on the fore coxae black ; the hair on the front side blackish, towards the basis of the coxae pale ; middle coxa? with black bristles and hairs ; the hind coxa? on their out- side with a single black bristle. Tibia? and tarsi elongated, very slender ; the tarsi from the tip of the first joint infuscated. The cilia of the tegulae yellowish. "Wings with a very distinct gray- yellow tinge ; the end of the fourth longitudinal vein rather strongly approaches the end of the third one. Hah. Virginia. (Osten-Sacken.) SAUCROPUS. 221 3. S. superfoiens LOEW. £ and 9 • — Ex glauco viridis, abdomine L-ete ffineo-viridijlbasim versus flavo. Grayish-green, the abdomen shining metallic-green, towards the basis yellow. Long. corp. 0.13 — 0.14. Long. al. 0.15. SYN. Saucropus svperbiens LOEW, Neue Beitr. VIII, 76, 3. Face of the male entirely linear ; that of the female also very narrow, snow-white. Palpi and proboscis dark -yellow. Antennae bright reddish-yellow ; . Bristles upon head, thorax and abdomen, slender, of medium length; the first joint of the fore tarsi indistinctly ciliated with very short minute bristles, middle tarsi plain. Long. corp. 0.18 — 0.23.' Loug. al. 0.20—0.24. SYN. Psilopus comatus LOEW, Neue Beitr. VIII, 89, 7. Male. Bright shining-green ; thorax and scutellurn (in the described specimen) more steel-blue, and the head handsome purplish-blue. Palpi black ; proboscis brown. Face without hairs, broader than usual; its small under side is dusted with grayish-white, the upper part more convex and, from a side view, more projecting than usual. The second joint of the antenna? with a small number of rather long, but not very stout bristles ; the third joint ovate ; arista apparently apical, but, in fact, only subapical, still considerably shorter than the body. Front with a scattered, very delicate white pubescence ; the black bristles, inserted upon the ocellar tubercle and in the immediate proxi- mity of the upper corner of the eye are hair-like and of a quite unusual length. The black bristles upon the upper side of the thorax and of the scutellum are of the same hair-like description ; also those inserted before the incisures of the abdomen, the latter are only of a still more striking length than the former. Upon the upper side of the abdomen there seem to be, besides the long bristles before the incisures, only single short hairs of a black color ; upon the first segment, however, as also on the lateral margin and on the venter, there is a very long whitish pubescence ; long, hair-like, black bristles are also to be found on the hind margin of the segments of the venter, similar to those upon the upper side of the abdomen. No black bands are perceptible at the base of the abdominal segments. The hypopygium is partly destroyed in the described specimen ; it seems that it was brown- ish-black, and that it had horn-shaped yellow appendages. Coxa> black, the foremost with a long white pubescence and with some black bristles. Femora black, with a green reflection ; the very extreme tip of the foremost ones yellow, all fringed with a scattered erect pubescence of a white color, which is everywhere of g, con- siderable length, but longest upon the under side. Tibia; and tarsi very slender, yellowish ; the usual black hairs rather scattered and very short. The fore tibia? upon the first two-thirds of their 2G4 DIPTERA OF NORTH AMERICA. [PART II. upper side, fringed with a row of hair-like bristles, which are alternately of remarkable length, and the last of which much ex- ceeds all the others in length. Middle tibia3 with a moderate number of hair-like black bristles, which are also distinguished by their unusual length, especially those upon their upper side. Hind tibiae only upon the upper side with short bristles, which are but little distinguished from the usual minute hairs. Fore tarsi at least twice the length of the tibia?, very slender : their first joint somewhat longer than the four following taken together, upon its upper side with a regular row of propor- tionally very long hair-like black bristles ; the following joints of rapidly decreasing length ; the third, at the upper side of the tip, with a single bristle-like black hair ; the fifth joint blackish-brown. Middle tarsi very slender, about once and a third the length of the tibias, from the tip of the third joint blackish-brown ; their first joint about over 1^ the length of the four following taken to- gether, the length of which is quickly decreasing ; upon its upper side it is regularly ciliated with bristle-like hairs; this fringe con- tinues over the upper side of the three following joints, but there it is shorter, more delicate, and closer ; the last joint is not ciliated, but has upon its upper side a short, appressed, snow-like pubes- cence, which is not very distinct. Hind tarsi not quite as long as the tibiae ; their first joint yellowish-brown, distinctly longer than the following taken together ; these are blackish-brown and of decreasing length. Halteres yellowish ; tegula? with a broad black margin and long black cilia. Wings somewhat narrow, hyaline, with blackish-brown veins, not ciliated on the fore mar- gin, the first longitudinal vein reaches nearly to the middle of the fore margin ; the anterior branch of the fourth longitudinal vein diverges at a rather acute angle and turns towards the margin at a rounded right angle, reaching it before the apex, near the tip of the thhxl longitudinal vein ; hind transverse vein very oblique, little sinuated. Female. It resembles the male very much. Face somewhat broader. Bristles on the second joint of the antenna?, the arista, the bristles upon the ocellar tubercle, at the upper corner of the eye, upon ,the upper side of the thorax, and upon the scutellum shorter ; the bristles before the incisures of the abdomen are very consider- ably shorter ; on the venter, as it seems, there are none at all. The black pubescence upon the upper side of the abdomen is less PSILOPUS. 265 scarce than in the male, if it has not been perhaps somewhat rubbed off in the latter ; the whitish pubescence of the venter much shorter. Feet of the same coloring as in the male ; the whitish pubescence of the femora is much shorter, still upon their under side of considerable length for a female ; upon the under side of the fore femora, near the basis, there are also some stiff, nearly bristle-like whitish hairs of a remarkable length. Pubescence and bristles of the tibiae generally like those of the male, still all bristles are not so long, and upon the upper side of the fore tibiae, the alternating shorter bristles are very small. Fore tarsi hardly once and a half the length of the tibiaa ; their first joint much longer than the following taken together, fringed upon the upper side with a regular row of short black bristles, upon the under side with only a few still shorter black bristles ; the following joints of decreasing length, from the second to the fourth brown, the fifth black. Middle tarsi distinctly longer than the tibia3, not ciliated upon the upper side ; the first joint distinctly longer than the four following taken together ; the latter black-brown and of decreasing length. Hind tarsi brownish-black, much shorter than the tibia3, the first joint .hardly somewhat longer than the following taken together. Wings as in the male, only somewhat shorter, also proportionally less narrow. Hab. Middle States. (Osten-Sacken.) Observation. — Whether the distinctly darker color of the tarsi of the 9 is only an accidental, individual deviation or not, cannot be judged from a single specimen of each sex. 2. Arista distinctly dorsal and not particularly elongated. a. Fore tibiae only in the ^, , fore and middle tibiEe in tlie 9 > yellow. 1O. P. clirysoprasius WALK. £ and 9. — Viridis, vel ex viridi chalybeus, nitidus, alis immaculatis, facie nuda. £ . Abdomine fasciis latiusculis nigris ornato, halterum nigrornm capitulo fusco, pedibus nigris, tibiis anticis flavis, tarsorum intermediorum articulo primo superne pilis erectis rigidis ciliato. 9 . Abdominis fasciis nigris, angustissimis, lialteribus flavis, pedibus nigris, tibiis anterioribus flavis. V Green or greenish-blue, shining ; wings without picture, face without hairs. £ . Abdomen with rather broad black bands ; knob of the black halteres brown ; feet black, the fore tibiae yellow, the first joint of the middle tarsi upon the upper side ciliated with upright stiff hairs. 2GG DIPTERA OF NORTH AMERICA. [PART II. $>. Abdomen with extremely narrow bands ; halteres yellow; feet black, fore and middle tibire yellow. Long. corp. 0.19 — 0.23. Long. al. 0.18 . —0.20. SYN. Psilopus chrysoprasi WALKER, List, etc. Ill, 646. Psilopus chrysoprasius LOEW, Neue Beitr. VIII, 90, 8. Male. Very shining, green, the head, the hind part of the thorax, the scutellum, and the anterior segments of the abdomen more steel-blue, the last segments of the abdomen sometimes golden-green. Palpi black ; proboscis brown. Face without hairs, thinly dusted with white ; its upper part rather convex. The bristles on the second joint of the antennae rather long ; the arista dorsal and of ordinary length. Front with a long whitish pubescence and the usual black bristles upon the thorax and the scutellum of a rather considerable length. The black bands of the abdomen are narrow upon the anterior segments and of con- siderable breadth upon the posterior ones. The black pubescence of the abdomen is rather short ; upon its first segments, as also upon the anterior part of the lateral margin and the venter there is a delicate white pubescence, while the hind part of the venter has a black pubescence of considerable length. The black bris- ' ties before the incisures of the abdomen are numerous, but rather short. The small hypopygium black, its larnellffi dark-brown. Coxae black, the foremost with a considerable white pubescence and a few black bristles. Femora black with a greenish reflection, upon the under side with a long, upright, whitish pubescence. Fore tibiae brownish-yellow, fringed with only a moderate number of short black bristles. Middle tibia? brownish-black, sometimes nearly black, beset upon the front side with a regular row of black bristles, otherwise only with a small number of them ; upon the upper side with rather bristle-like black hairs. Hind tibias black with coarse black hairs, upon the outside fringed with a regular longitudinal row of black, only moderately long, bristles. Fore tarsi rather slender, about once and a third the length of the tibiae ;' the first joint brownish-yellow, much longer than the following taken together, upon the hind side with three or four stout black bristles; the following joints blackish-brown, from the second to the fourth of rapidly decreasing length, the fifth as long as the fourth. Middle tarsi about once and a half the length of the tibia3 ; the first joint black-brown, nearly twice the length PSILOPUS. 26T of the following taken together, upon its upper side with a regular fringe of stiff, perpendicularly erect, bristle-like, minute hairs, upon the under side with about seven minute black^ bristles ; the joints of the tarsi from the second to the fourth very rapidly decrease in length, so that the second is still somewhat longer than the third and fourth taken together ; the fifth joint not shorter than the fourth ; the ciliatiou of the first joint of the tarsi continues also over the upper side of the second and third joints, but consists there of little hairs of gradually dimin- ishing length, so that finally it becomes almost imperceptible. Hind tarsi black, perceptibly shorter than the tibiae ; their first joint somewhat longer than the following taken together, the length of which is gradually decreasing. Halteres blackish, the knob brown or dingy brownish-yellow ; the tegula? with a broad black border and long black cilia. Wings hyaline with black veins ; the first longitudinal vein reaches nearly to the middle of the wing ; the anterior branch of the fourth longitudinal vein diverges from it under a nearly right angle, and turns afterwards in a curve towards the margin, which it reaches rather far from the apex, immediately near the tip of the third longitudinal vein ; hind transverse vein rather oblique, hardly somewhat sinuated ; the space between the margin of the wing and the third longitu- dinal vein, beyond the tip of the first longitudinal vein, is some- what more grayish than the remaining surface of the wing. Female. Yery like the g . The face but little broader. The arista, the bristles of the second joint of the antennae, those upon front, thorax and scutellum, as also before the incisures of the abdomen, are much shorter. The white pubescence on the under side of the femora also shorter, still of a considerable length for a 9. Fore tibise as in the g , still the bristles shorter. Middle tibice brownish-yellow, with scattered black bristles of medium length. Fore tarsi but little shorter than in the g , however of the same structure and coloring. Middle tarsi plain, brownish- black, not quite once and a half the length of the tibiae ; their firsi joint not ciliated. Halteres light-yellowish with blackish peduncle. Wings as in the J\ still without the gray shade, which is to be found in the latter beyond the tip of the first longitudinal vein. Hab. Cuba. (Poey.) 268 DIPTERA OP NORTH AMERICA. [PART II. 6. In the males all the tibiae, in the female also all femora yellow. 11. P. SCoMliator LOEW. £ and 9. — Parvus, nitidisimus, viridis vel ex viridi chalybeus, facie pilosa, alis immaculatis, anteriore vense longitudinalis quartae ramulo arcuatim ducto, halteribus flavis. £, . Abdomine fasciis nigris angustis ornato et in apice pilis solito paulo longioribus vestito, femoribus nigris, viridimicautibus, summo femorum anteriorum apice tibiisque omnibus flavis, tibiarum posticarum apice tarsisque omnibus ex fusco nigris, primo tameu tarsorum anticorum articulo flavo, calcare tibiarum iutermediarum longissimo, primo tarso- rum intermediorum articulo setulis incurvis scabro. 9 . Fasciis abdominalibus nigris nullis, coxis anticis, femoribus tibiisque omnibus flavis, extreme tibiarum posticarum apice tarsisque omnibus ex fusco nigris, basi tamen tarsorum anteriorum flava. Small, very shining, green or greenish-blue ; face hairy • wings without picture, the anterior branch of the fourth longitudinal vein arch-like ; halteres yellow. £ . Abdomen with narrow black bands, at the tip with a somewhat shorter pubescence than usual ; femora black, with a green lustre, the extreme tip of the four anterior femora and all tibiae yellow, the tip of the hind tibiae and all tarsi brownish-black, still the first joint of the fore tarsi yellow, the spur of the middle tibiae extremely long ; the first joint of the middle tarsi rasp-like, being beset with crooked bristles. 9 • Abdomen without black bands ; fore coxae, all femora and all tibiae yellow ; the extreme tip of the hind tibiae and all tarsi brownish-black* still the basis of the fore and middle tarsi yellow. Long. corp. 0.15 — 0.16. Long. al. 0.15.— 0.16. SYN. Psilopus scobinator LOEW, Neue Beitr. VIII, 91, 9. Male. Handsome shining, green, sometimes partly steel-blue. Palpi black ; proboscis brownish-yellow. Face with a long whitish pubescence and not very closely dusted with white. The second joint of the antenna? with moderately long bristles ; the arista itself only of the usual length. Tbe delicate scattered pubescence of the front generally yellowish, more seldom whitish ; the usual black bristles upon the ocellar tubercle and in the upper eye- corner rather long. The bristles upon thorax and scutellum are also rather long. The hind part of the abdomen is more pointed in the shape of a cone than in most of the other species ; the black bands at the basis of the segments of the abdomen are narrow, broader only upon the segments. The short pubescence of the abdomen is black; upon the first segment, as als"o on the anterior part of the lateral margin and on the greater part of the venter it is. PSILOPUS. 209 whitish ; the black bristles before the incisures are not remarkably long ; the bristle-like hairs at the extreme tip of the abdomen are rather long. The small hypopygium is black, with extremely small black appendages. Coxa? black, dusted with white, the fore- most with a white pubescence and a few black bristles. Femora black with a green reflection ; the anterior ones light yellow at the extreme tip ; upon the under side of all there is a delicate, erect, rather sparse pubescence of medium length. Tibiae yellow ; the hindmost colored with blackish-brown to a rather considerable extent at the tip. Fore tibia3 upon the upper side with about four short black, minute bristles, upon the hind side with about six still shorter ones. Middle tibiae upon the first half of the hind side with three not very long black bristles ; a perceptibly longer bris- tle is to be found upon the first quarter of the front side, another near its end ; immediately before the end on the under side an unusually long, straight, diverging, black bristle assumes the shape of a rather striking spur of the tibia. On the hind tibia?, besides the black bristles inserted at the tip, there is but one bristle de- serving to be noticed, upon the first third of the outside. Fore tarsi slender, somewhat longer than the tibia? ; their first joint yellow, only at the extreme tip brownish-black, nearly once and a half the length of the following taken together ; the latter are brownish-black ; from the second to the fourth of rapidly decreasing length, the fifth as long as the fourth. There are no bristles nor any unusual pubescence on the fore tarsi. Middle tarsi brownish- black, considerably shorter than the tibia? ; their first joint about once and a quarter the length of the following taken together, its whole length upon the under side fringed rasp-like with short black bristles, crooked downwards, and inserted in a row on both sides ; the four following joints of decreasing length. Hind tarsi black, hardly two thirds of the length of the tibia? ; their first joint little longer than the following taken together ; the second to fourth joint of rapidly decreasing length ; the two last joints of an equal length, somewhat flattened. Halteres yellowish with blackish peduncle ; tegula? with black cilia. Wings hyaline with black veins ; the end of the first longitudinal vein is a considerable dis- tance before the middle of the wing ; the anterior branch of the fourth longitudinal vein diverges from it at an angle which is very nearly a right one, and turns then arch-like towards the margin, which it reaches rather far before the apex, quite near the tip of 270 DIPTERA OF NORTH AMERICA. [PART II. the third longitudinal vein'; hind transverse vein nuderutely ob- lique and nearly straight. Female. Face somewhat broader than in the J*. All bristles perceptibly shorter. Abdomen; in the usual position of the seg- ments, without black bands. Pore coxae yellow, at the extreme basis often colored with gray ; their white pubescence much shorter and the black bristles more striking. All the femora altogether yellow, fringed upon the under side with but very short, erect, whitish hairs. Tibiae quite yellow, the hindmost somewhat infuscated only at the extreme tip. The bristles of the tibia? are quite like those of the J1 , still some of the bristles are missing, or at least shorter. Tarsi plain, somewhat shorter than in the J1 ; the first joint of the middle tarsi brownish-yellow, upon the under side with but a few very short black bristles. Hob. New York, Illinois. (Osten-Sacken.) Observation 1. — With P. scobinator begins a series of very closely resembling species, which agree in a remarkable manner nut only in size and coloring, but also in the majority of the plastic characters. The structure of 'the middle feet in the J* shows differences, which leave no doubt as to their specific dis- tinctness. The distinction of their females offers such difficul- ties, that I am unable to overcome them with the materials at my command. I have received P. scobinator iu so large numbers, that the 9 belonging to this species is surely abundantly repre- sented among them ; but whether I have not confounded with it females of the three following species, I am not able to tell. Some of the females, I suspect, belong to P. caudatulus, others may be P. inermis, but I am not positively certain about any one of them. Positive characters for the distinction of the females of these species can only be acquired by the observation of the species in life. Observation 2. — There is no doubt that P. femoratus Say belongs to the present group. Whether it is one of the species known to me, and which of them, I cannot decide, as Say does not mention any of those plastic characters by which alone they can be distinguished, and as the J1 specimen sent by Say to Wiedeman has not been preserved in the collection of the latter. PSILOPUS. 271 12. P. cailclSLtulllS LOEW. £ . — Parvus, uitidissimus, viridis vel ex viridi chalybeus, facie pilosa, alls irumaculatis, aiiteriore veuse lougi- tudinalis quartse raraulo arcuatira ducto, halteribus flavis. 'J, . Abdomine fasciis nigris angustis ornato et in apice pilis solito multo longioribus vestito, femoribus uigris, viridirnicantibus, suinino femorum anterioruru apice tibiisque omnibus liavis, apice tibiaruni posticarum tarsisque omnibus ex fusco uigris, tarsorum anteriorum basi taineii flava, calcare tibiarum intermediarum lougissimo, priiuo tarsorum iiiter- inediorurn. articulo setulis incurvis scabro. 9 Small, very shining, green or bluish-green ; face hairy ; wings without picture ; the anterior branch of the fourth longitudinal vein arched ; halteres yellow. £ . Abdomen with narrow black bands, at the tip with much longer hairs than usual, femora black, with a greenish reflection, the extreme tip of the four anterior femora, and all tibiae yellow ; the tip of the hind tibire, and all t£,rsi brownish-black ; still the first joint of the fore tarsi yellow, the spur of the middle tibia very long; first joint of the middle tarsi beset with crooked bristles, rasp-like. 9 Long. corp. 0.15. Long. al. 0.15. SYN. Psilopus caudatulus LOEW, Neue Beitr. VIII, 93, 12. It resembles the P. scobinator so much, that the statement of the slight differences will be perfectly sufficient for its recognition ; the hairs at the tip of the abdomen are much longer, more bristle- like, and when well preserved, are of the same length as the four last segments of the abdomen, whilst in P. scobinator, the two which are longest among them are but little longer than the last segment. Hab. Missouri. (Schaurn.) Illinois. (Le Baron.) Observation. — The name which I have given to this species is intended -to remind of P. caudatus Wied., which undoubtedly belongs to the same group ; but, by its larger size, it seems to be different from the species known to me. In the Neue Eeitra . Alarum costa nee concava, nee ciliata. Golden-green, moderately shining, proboscis, palpi, the two first joints of the antennae, the basis of the abdomen, the venter, the fore and hind coxae, as also the feet, yellow ; the middle coxse gray with yellow tip. £ . The anterior margin of the wings concave, with a fringe of short cilia. J . The anterior margin of the wings neither concave nor ciliated. Long, corp. 0.20—0.22. Long. al. 0.19—0.20. SYN. Psilopus psittacinus LOEW, Neue Beitr. VIII, 96, 16. Male. Golden-green, moderately shining. Proboscis and palpi yellow. Face greenish-blue, closely dusted with yellowish-gray, reaching much downwards, without hairs. The two first joints of the Antennae yellowish, the second with very short minute black bristles, The arista dorsal, moderately long. Front blue or bluish-green, with a grayish-yellow or nearly whitish dust, without 28?; DIPTERA OF NORTH AMERICA. [PART II. Lairs, its usual black bristles of a very moderate length. Thorax, scutellum and abdomen with a slight yellowish dust, which moderates the lustre of these parts. The black bristles of the thorax of moderate length. The scutellum has only the pair of bristles at the tip, which is of a considerable length ; immediately near each bristle of this pair a much shorter black hair is inserted towards the outside. Pleura? black with a green reflection and with a grayish-white dust ; their hind margin gray or at least only in part yellowish. Abdomen green, towards its end generally some- what gilded ; the first segment yellowish, at its basis blackish, on the hind margin metallic-green ; on the second segment the ante- rior margin and corner, sometimes also the lateral margin, yellow ; the third segment has generally, on the anterior part of the lateral margin, an elongated yellowish spot. The scattered pubescence of the abdomen is rather delicate and long ; upon the upper side it is black, whitish only upon the first segment ; the pubescence of the venter is whitish near its basis, but becomes gradually darker towards the end. The black bristles before the incisures of the abdomen of a very moderate length. The external append- ages of the hypopygiuni small and very narrow, brownish-yellow. Fore and hind coxae pale yellow, the former with a shorter and more delicate whitish pubescence and with a few light-yellowish bris- tles; middle coxa? gray with pale-yellowish tip. Feet yellowish. Femora slender, upon the under side sparely fringed with exceed- ingly short, minute whitish hairs ; besides, upon the second half of the under side of the middle femora there is a sparse row of minute black hairs. Tibia? slender and long ; the usual black pubescence of the fore and hind tibia? is very short and appressed, that of the middle tibia? somewhat longer and more erect, there- fore almost fringe-like. The fore tibia?, with the only exception of a rather small minute black bristle, inserted. upon their outside, near the basis, are entirely without bristles. The middle tibias bear no bristles whatever, distinguished from the other pubescence. Hind tibise upon the front side, not far from the basis, with a rather apparent little bristle, whilst those upon the upper and under side are less numerous, extremely small and therefore more difficult to perceive. Fore tarsi slender, nearly twice as long as the tibiae ; their first joint for itself alone somewhat longer than the tibia, the following ones of decreasing length, the last one brown. Mid- dle tarsi slender, about once and a half the length of the tibia? ; PSILOPUS. 283 the first joint distinctly shorter than the tibia, upon its front side somewhat sparsely ciliated with oblique niiiiute black hairs ; the following joints only with the usual short pubescence and of de- creasing length, the last one dark brown. The length of the hind tarsi somewhat exceeds three-fourths of the tibia? ; their first joint is only somewhat longer than the following taken together ; these are of decreasing length, somewhat infuscated, still only the last one dark brown. Halteres light yellow ; tegula? with extremely narrow black margin and with yellowish-white cilia. Wings with brown veins ; the anterior margin is gently sinuated upon its larger second half, and forms before the tip of the second longitu- dinal vein a projecting angle, so that the whole wing acquires a quite unusual axe-like shape ; the whole anterior margin, as far as that angle, is delicately and equally ciliated ; the third longitudinal vein, near its end, is turned back very abruptly and unusually far ; the anterior branch of the fourth longitudinal vein diverges from it under a rather acute angle and turns then in a curve towards the margin, which it reaches not far from the tip of the third lon- gitudinal vein ; posterior transverse vein rather remarkably oblique, gently sinuated in the shape of an S. Female. It resembles the male very closely. Besides the lesser length of the bristles on the whole body, and besides the somewhat lesser length of the feet, which are of the same color as in the J1 , there are only the following differences : the whole hind margin of the pleura? is colored with yellow. The middle tibia? are without the longer and erect pubescence of the^, but their pubescence is short and appressed as on the other tibiae ; there are, however, upon the front side near the basis one, and at the tip of the tibia? a few more distinct minute black bristles, besides some smaller ones upon the hind side. The anterior margin of the wings is neither concave nor ciliated, the wings therefore of the usual form ; the third longitudinal vein, near its end, is much less sud- denly and much less strongly curved backwards; the course of the anterior branch of the fourth longitudinal vein is somewhat less in a curve, and the sinuosity of the hind transverse vein not so strong. Hob. Florida. (Osten-Sacken.) 284 DIPTERA OF NORTH AMERICA. [PART II. B. Fore femora upon the under side with ihornlike bristles in loth sexes. 1. All the coxae entirely yellow. . P. tener LOEW. £ and 9 • — Laete viridis, proboscide, palpis, priniis duobus antennarurn articulis, pleurarum margine postico, abdo- minis basi, coxis pedibusque flavis, ciliis tegularum albicantibus, alarum ven§, transversa posteriore valde obliqua. £ . Femora antica setulis subtribus armata ; appendices hypopygii majus- culas flavse. 9 . Femora antica setis quatuor validis armata. Light green, proboscis, palpi, the two first joints of the antennae, the hind margin of the pleura, the basis of the abdomen, the coxae and feet yel- low ; cilia of the tegulse whitish ; hind transverse vein of the wings very oblique. £ . Fore femora generally with three minute, bristles ; the rather large appendages of the hypopygium yellow. 9. Fore femora with four stout bristles. Long. corp. 0.16 — 0.17. Long, al. 0.22. SYN. Psilopus tener LOEW, Berl. Ent. Zeitschr. VI, 217, 71. Light metallic-green, rather shining. Proboscis and palpi yel- low, the latter beset with a few minute bristles, which have in the $ a whitish, in the 9 a black color. The glabrous face, covered with white dust, is very broad, particularly upon its upper part, which is very convex. Antenna small, the two first joints yellow, the second beset with a few short, minute black hairs upon the upper side ; upon the under side with a few short, minute white hairs ; the very small third joint is generally blackish, still it has sometimes a much lighter coloring. The front is generally sky- blue, and, besides the usual black bristles, without pubescence. Upper side of the thorax sometimes more blue than green, rather distinctly dusted and beset with but a moderate number of black bristles. Pleuras, in consequence of a close white dust, grayish- green. The scutellum, which has two bristles, is sometimes green, sometimes purplish-blue. Abdomen shining light-green, some- times rather golden-green ; its basis is in the 9 always, in the male generally, colored with yellow ; the black bristles before the hind margin of the single segments are proportionally short. Hypopygium blackish-brown and dusted with white ; its append- ages are pale yellow ; the external ones are narrow lamellae and are fringed with a blackish pubescence, which is somewhat longer PSILOPUS. 285 at their tip ; the inner ones are stile-like, distinctly longer than the outer ones and fringed upon their upper side with some few minute light hairs ; the stile-like central organ is as long as the inner appendages and is smoothly curved downwards. The coxae and the long, yery slender and very glabrous feet are yellowish. The fore coxae of the £ have a proportionally long, rather close and delicate whitish pubescence ; in the 9 there are, instead of the longer hairs, bristles of a white-yellowish coloring. The fore femora of the £ have upon the under side near the basis, three thin yellowish bristles of decreasing length ; in the 9 there are in their stead four stout yellowish bristles. The hind femora of the £ have upon the upper side near the basis a few long and very delicate white hairs, which are not present in the 9 . The plain and slender fore tarsi are in both sexes much longer than the tibia? ; they are still much longer in the £ than in the 9 ; the first joint of all tarsi is very elongated, particularly in the £ ; on the fore tarsi it is about twice as long as the following joints taken together, on the middle tarsi about three times as long, besides they are distinguished by their slenderness, and apparently complete glabrousness. The hind tarsi in both sexes are nearly as long as the tibia?, their first joint in the £ somewhat longer than the fol- lowing joints taken together, but in the 9 somewhat shorter. Tegulse on the margin with only a single quite small black dot, ciliated with whitish. Wings grayish hyaline, long and narrow, towards the basis still more narrowed, particularly in the % ; their anterior veins are yellow, the posterior ones more infuscated ; the anterior branch of the third longitudinal vein is very long, rather flat and but little curved ; the posterior transverse vein is far remote from the margin and has a very oblique position. The £ possesses, as a particular distinctive mark, upon the under side of the basis of the wing, a crooked, black, rather stout thorn, in- serted near the anterior margin. Hob. Pennsylvania. (Coll. Winth.) Observation. — The P. delicatus of Mr. "Walker, who described a 9 , has an entirely green abdomen, can therefore not be tener, as the basis of the abdomen of its 9 is always colored with yellow. 286 DIPTERA OF NORTH AMERICA. [PART II. 2. Middle and hind coxae blackish. 21. P. filipes LOEW. £ . — Gracilis, longipes, nitidus, capite, thorace scutelloque chalybeis, abdomine viridi, proboscide, duobus primis an- tennarum articulis, coxis pedibusque flavis, femorum intermediorum basi nigra, tibiis intermediis tarsisque omnibus fuscis. Slender, long-legged, shining, head, thorax and scutellum steel-blue, ab- domen green ; proboscis, the two first joints of the antennae, coxa3 and feet yellow, still the basis of the middle femora blackish ; middle tibiae and all tarsi brown. Long. corp. 0.26. Long. al. 0.26. SYN. Psilopus filipes LOEW, Neue Beitr. VIII, 99, 18. Of slender shape and very long-legged, shining. Head pur- plish-blue ; proboscis yellow, palpi brown, both beset with minute light hairs. Face without hairs, very broad, covered with dense yellowish dust ; its upper part is quite unusually convex. The two first joints of the antennae yellow, the second fringed with short minute white-yellowish' hairs ; the small third joint rounded, brownish-black; arista dorsal, proportionally of insignificant length. Front without hairs, with grayish-yellow dust, which does not cover the shining purplish ground-color upon the middle ; the usual black bristles of moderate length. Thorax and scutel- lum shining purplish-blue, with a yellowish dust, which does not cover the lustre of the ground-color. The black bristles of the tho- rax rather long. The scutellum has but two bristles, still near the lateral corners, on its margin, some long hairs are inserted. Ab- domen shining green with a very slight yellowish dust ; if the light falls upon it from behind, narrow black fasciae on the anterior mar- gins of the segments become apparent, which are invisible, if the light falls from the opposite side. The delicate pubescence of the abdomen is whitish ; upon the upper side of the two first segments, on the lateral margin and on the venter very long, otherwise short; black bristles of moderate length are only to be found before the hind margin of the third and of the following segments. Hypopygium blackish-green with white dust ; the external ap- pendages stile-like, but little shorter than the inner ones, their basal half light-brownish and hairy, their apical half white and glabrous, their extreme tip black ; the inner appendages form a brownish-black forceps. Fore coxae yellowish with a yellowish- white pubescence, which has on their outside a rather unusual length ; there are no bristles upon it. Middle and hind coxse PSILOPUS. 2 87 blackish. Feet yellow, very long and beset with but exceedingly short, for the greatest part minute light-colored hairs. All femora slender ; the foremost ones tapering towards the end, the middle ones from the basis as far as the first third, black ; fore femora upon the first half of the under side with six perpendicular, erect, long, yellow, spine-like bristles ; hind femora upon the first third of the hind side fringed with long, but very delicate whitish hairs. All tibiae very long and slender, fore and middle tibia? without bris- tles ; the latter, excepting the basis, are rather dark-brown, very elongated, and gradually tapering towards their end. Hind tibias with rather numerous, very short black bristles upon the under side. Fore tarsi brown, very slender, over once and two-thirds the length of the tibia? ; their first joint is distinctly longer than the tibia ; the following joints are of a decreasing length, and the last one somewhat flattened. Middle tarsi much longer than the fore tarsi, but not quite as long as the middle tibia?, filiform ; the first joint brownish-black, about twice and a half the length of the following taken together, with hairs of such shortness that it ap- pears entirely bare unless very closely examined ; the four follow- ing joints light-brownish, still the extreme tip of the second and of the almost equally long third joints brownish-black ; the two last joints are again of rather equal length, but, taken together, are only about as long as the second joint ; the second and third joints with a distinct black pubescence, the fourth joint with a still longer pubescence ; the small ungues are sharp and the pulvilli very short. Hind tarsi brownish-black with yellowish basis, not quite as long as the tibia? ; their first joint somewhat longer than the four following taken together ; the joints from the second to the fourth' of rapidly decreasing length ; the fifth joint about as long as the fourth. Halteres pale-yellowish ; tegula? with whitish cilia. Wings very long and narrow, with brown veins ; the third longitudinal vein is only slightly turned' backwards near its end ; the anterior branch has its origin in the middle between the hind transverse vein and the margin ; it diverges from this vein at an obtuse angle and turns in a very flat curve towards the margin, which it reaches somewhat before the extreme apex and not far from the tip of the third longitudinal vein ; the hind transverse vein is unusually distant from the margin, has a rather oblique position and is not distinctly inflected. Hab. Middle States. (Osten-Sacken.) APPENDIX SPECIES DESCRIBED BY PREVIOUS AUTHORS AND NOT CONTAINED IN THE PRESENT MONOGRAPHS. Fdbricius, Systema Entomologise. Page 783. Must a longicornis. Antennis setariis, pilosis, asneo-nitens, pedibus nigris. Hab. America. Parva, statura M. angulatse.1 Antennae nigras, pilosae setaque unica longitudine fere corporis. Corpus viridi aeneum, nitidum, pedibus solis nigris. Alas hyalines. Cauda uncinata. Fabricius, Entomologia Systematica, Vol. IV. Page 341. Musca longicornis. Antennis setariis elongatis pilosa aeneo-nitens, abdomine ob- scuriore. Hab. Americse insulis. Statura parva M. unguTatse. Antennas elongatse nigrae, pilosae setaque unica longitudine fere corporis. Corpus viridi seneum, nitidum, pedibus solis nigris. Cauda uncinata. Say, Journ. Acad. Nat. Sc. Philad. Vol. III. Page 85. Dolichopus imifasciatiis. Bluish-green ; a white band at the base of the abdomen. Hab. Pennsylvania. Body bluish-green, polished, slender ; antennas, palpi and pro- 1 Misprint for ungulatx. '19 ( 289 ) 290 DIPTERA OF NORTH AMERICA. [PART II. boscis whitish ; scutellum blue ; wings immaculate ; feet whitish ; tergum, first segment and half of the second whitish, posterior half of the second segment and third segment much tinged with blue, remaining segments green. Length — one-fourth of an inch. Central nervure of the wing furcate, the exterior branch widely angulated and terminating near the tip of the preceding nervure, which is curved very considerably inwards, towards its tip. Page 85. Dolichopus obscurus. Blackish-brassy ; wings dusky ; feet pale Hob. Pennsylvania. Head dark silvery ; antenna? black-brown ; mouth blackish ; thorax and scutellum dark-brassy ; wings dusky ; feet white, a lit- tle dusky on the tarsi ; poisers white ; tergum rather darker than the thorax. Length — less than three-twentieths of an inch. The central nervure of the wing is nearly rectilinear, being hardly perceptibly reflected. Page 86. Dolichopus femoratus. Green ; tibiae and tarsi whitish. Hob. Pennsylvania. Body brilliant green, with bluish reflections ; front pruinose ; antennae blackish ; proboscis yellowish ; wings hyaline ; scutellum blue ; thighs green and excepting the posterior ones, whitish at tip, tibiae white, tarsi dusky ; tergum, ultimate joints cupreous at their bases. Length — three-twentieths of an inch. The brilliancy and shade of green in this insect are similar to D. sipho; when living, and in the sun's rays, it resembles bur- nished gold, nervures nearly as in sipho. Say, Journ. Acad. Nat. Sc. Philad. Vol. VI. Page 168. Psilopus femoratus. This brilliant species varies in the color of its thighs, which in my description are stated to be green ; a specimen taken in Indiana 'has whitish thighs. APPENDIX. 291 ' Page 168. Clirysotus nubilus. Blackish ; feet dull honey-yellow. Hob. Indiana. Body blackish ; head dull plumbeous ; thorax cinereous, with three brown lines ; wings immaculate ; poisers white ; feet dark honey-yellow ; thighs black at base and above. Length — rather over one-tenth of an inch. Page 168. Chrysotus conciimarius. Green-brassy ; tergum blue towards the tip. Hob. Mexico. Head violaceous, with a cinereous reflection ; palpi with a cine- reous reflection ; antennae black ; thorax green with a gray oliva- ceous reflection ; wings hyaline, obsoletely tinged with yellowish on the costal margin ; poisers white ; tergum with a gray reflec- tion, brassy-green at base, and violaceous towards the tip ; thighs green ; tibiae whitish. Length — one-fourth of an inch. Page 169. Chrysotus abdominal is* Green, polished ; feet white. Hob. Indiana. Body bright green, brilliant ; hypostoma purple ; antennae yel- low ; thorax immaculate ; poisers yellow ; tergum, first segment at base with an obsolete yellowish line ; feet white ; venter white, at tip blackish purple. Length — £ one -tenth of an inch. * Page 169. Medetems lateralis. Tergum pale, with a lateral series of polished spots. Hob. Indiana. Head silvery ; proboscis et antennas yellowish, seta of the latter with the first joint very short ; eyes (when recent) green polished, with a cupreous reflection ; thorax green, somewhat pruinose, with a dorsal rather compound vitta ; wings hyaline ; poisers whitish ; tergum dull yellowish, with a series of brassy spots on each side, posterior two largest ; feet and venter whitish. Length — nearly three-twentieths of an inch. The dorsal vitta is impressed behind. 292 DIPTEKA OF NORTH AMERICA. [PART II. Page 170. Medeterus pimctipennis. Thorax variegated ; wings with brown spots. Hob. Mexico. Thorax olive-brown, trilineate ; middle line slender, dull yel- lowish, obsoletely zigzag ; outer lines cinereous with black points; scutel brown, cinereous in the middle ; wings hyaline, with many irregular fuscous spots, hardly to be traced into four bands ; poisers yellow ; tergum cupreous, posterior margins of the seg- ments blackish ; feet white ; tarsi blackish. Length — nearly one-fifth of an inch. Page 170. Dolieliopus abdominal!*. Green, abdomen rufous. Hob. Indiana. Head silvery ; antennae, first and second joints black, third ; thorax polished green ; wings hyaline ; abdomen, excepting the terminal joint, rufous ; halteres white ; pleurae and pectus blackish, pruinose ; feet white ; tarsi dusky. Length — less than three-twentieths of an inch. Wiedemann, Aussereuropaische Zweyfluglige Insecten, Vol. II. Page 219. No. 12. Psilopus macula. Yiridaureus ; antennis nigris, pedibus flavis ; alis macula magni fusca. Griingolden, mit schwarzen Fiihlern, gelben Beinen und einenj grossen schwarzlich-braunen Fliigelflecke. Lange 3 Linien 9« Yon der Krabbeninsel in Westindien. Untergesicht an den Fiihlern schon stahlblau, weiter unten griingolden, uberall aber in gewisser Richtung fast silberweiss schimmernd. Stirn griingolden, in's Stahlblaue fallend. Riicken- schild ebenso, ganz vorn wenig weiss-schimmernd. Brustseiten ebenso, doch uberall stark weiss-schimmernd. Hinterleib griin- golden, an. den Einschnitten schwarz, in gewisser Richtung auch wohl kupferrothlich. Beine gelb ; hinterste Schienen und Fiisse allmahlig braun. Fliigel wasserklar ; der Fleck liegt langs der Rippe und nimmt von ihr selbst bis Zwei drittel der Fliigelbreite ein, er liegt von Wurzel und Spitze gleichweit entfernt, nimmt etwa die Zwei mittlern Yiertel der Fliigel ein, und hat einen ge- APPENDIX. 293 rundeten Innenrand. (In Hornbeck's Sammlung zu Kopen- hagen.) (Translation.) — Golden .green, with black antennae, yellow feet and a large broWnish-black spot on tlie wings ; length three lines, 9 • From the Crab Islands (West Indies). Face bright steel-blue near the antennae, golden-green farther below, everywhere with a silvery reflection in a certain light. Front golden- green, With a bluish reflection ; thorax likewise, but with a silvery reflec- tion anteriorly ; jjleurse of the same color, altogether with a whitish reflec- tion. Abdomen golden green, black at the incisures, in a certain light reddish-coppery. Feet yellow ; hind tibiae and feet gradually infuscated. Wings hyaline ; the dark spot is along the costa and extends to about two- thirds of the breadth of the wing ; it is equally distant from the base and the tip, and occupies about two-fourths of the length of the wing in the middle ; its inner border is rounded. (Hornbeck's Collection in Copen- hagen.) Page 219. No. 13. Psilopus Say i. jEneus ; antennis, abdominis basi, incisuris pedibusque flavis. Erzgriiu, mit gelben Fiihlern, Hinterleibswurzel, Einschnitten und Beinen. Liinge 2f Linien $ . Aus Pennsylvanien. Dolichopus unifasciatus Say, Journ. Acad. Phila'd. Ill, 85, 2. Fiihlerwurzel, aber auch wahrscheinlich das verloren gegangene Endglied, gelb. Uutergesicht lebhaft erzgriin, unten weuig weiss- schimmernd ; Stirn smaragdgriin, mit Metallglanz. Riickenschild in's Griingoldeue iibergeliend ; die griingoldenen Brustseiten weiss bereift. Erster Hinterleibsabschnitt uberall, zweiter an der Wurzel, folgeude an den Einschnitten gelb ; die Hauptfarbe des 'Hiuterleibes ist an der Wurzel smaragdgriiulich erzfarben, was an der Spitze ins Griiugoldene iibergeht. Fliigel sehr licbt gelblich, mit gelben Adern. Beine gelb ; vorderste Fiisse uberall, hintere nur an der Spitze sckwarzlich-braun. (Im Philadelphischen Museum.) (Translation.) — Bronze-green, with yellow antennae, basis of the abdo- men, incisures and feet ; length two and two-thirds of lines, £ . Penn- sylvania. SYN. D. unifasciatus Say, Journ. Ac. Phil. Ill, 85, 2. Basis of antennae and probably also the last joint (which is broken) yellow. Face bright bronze-green, with a slight whitish reflection below. Front emerald-green, with metallic reflection. Thorax verging into golden- green ; the golden-green pleurae whitish-pruinose. First abdominal seg- ment yellow, the root of the second and the incisures of the following, also 294 DIPTERA OF NORTH AMERICA. [PART II. yellow. The principal color of the abdomen is bronze emerald-greenish towards the base, changing into golden-green towards the tip. Wings very light yellowish, with yellow veins. Feet yellow, fore tarsi altogether blackish-brown, posterior ones only at the tip. (Museum of Philadelphia.) Page 220. No. 14. Psilopus Ion gi corn is FABR. ./Eneus ; abdoniine incisuris atris, alls limpidis. Erzgriin, mit tief schwarzen Hinterleibseinschnitten und was- serklaren Fliigeln. Liinge 2f Linien. Aus Westindien. Untergesicht erzgriin, kaum ain untern Theile in gewisser Richtung weisslich schimmernd. Stirn erzgriin. Riickenschild und Hinterleib griingolden, dieser an der Wurzel der Abschnitte tief schwarz. Fliigel ungefarbt ; Schwinger gelb. Beine schwarz an den Schenkeln fast metallglanzend an den vordersten Schienen lehmgelb. (In der Fabricius, schen Sammlung.) (Translation.) — Bronze-green; abdomen with deep black incisures ; wings hyaline ; length, two and two-thirds of lines. West Indies. Face bronze-green, with a slight whitish lustre on its lower part. Front bronze-green. Thorax and abdomen golden-green ; the latter deep black at the root of the segments. Wings hyaline ; halteres yellow. Feet black, almost with a metallic lustre on the femora ; fore tibiae luteous yel- low. (Fabricius's collection.) Page 224. No. 23. Psilopus caudatus. Thorace aeneo-viridi, abdomine viridaureo ; incisuris atris, tibiis flavis. Mit erzgriinem Riickenschilde und griingoldenem, schwarz eingeschnittenem Hinterleibe und gelben Schienen. Lange 2 bis 2£ Linien. Von Savannah. Deni P. longicornis verwandt. Fiihler schwarz ; Untergesicht erzgriin, silberweisslich schimmernd ; stirii griingolden, am Weib- chen mehr als am Mannchen. Riickenschild des Mannchens hin- ten in's Stahlblaue iibergehend, am Weibchen durchgehends griin- golden. Brustseiten silberweisslich, Hinterleib griingolden, mit an der Wurzel tief schwarzen Abschnitten. After des Mannchens mit sehr langen Haaren besetzt. Fliigel wasserklar ; Schwinger gelblich ; Schenkel des Mannchens erzgriin : vordere mit ledergelber Spitze, Schienen sammtlich ledergelb. Fiisse schwarz. Am Weibchen sind auch die Schenkel gelb. (In Westermann's Samm- lung.) APPENDIX. 295 (Translation.) — Thorax bronze-green; abdomen golden-green, with black incisures ; tibiae yellow ; length from two to two and one-fourth lines. (Savannah.) Related to P. longicornis. Antennae black. Face bronze-green, with a silvery-white reflection ; front golden-green, more so in the 9 than in the £ . Thorax of £ more steel-blue posteriorly ; of the 9 altogether golden- green. Pleurae silvery-white. Abdomen golden-green, with the segments deep-black at the basis. Hypopygium beset with black hair. Wings hyaline ; halteres yellowish. Femora of the male bronze-green ; the fore- most with the tip yellowish ; all the tibiae leather-yellow. Tarsi black. The femora of the 9 also yellow. (Westermann's collection.) Page 224. No. 24. Psilopus Virgo. ^Eneo-viridis ; pedibus laete flavis. Erzgriin, mit schon gelben Beinen. Liinge 2 Linien, 9 . Von New York. Fiihler schwarz. Untergesicht griin, mit silberweissem Schim- mer. Mittelleib schon erzgriin. Hinterleib griingolden. Fliigel ungefarbt, Adern brauu ; Spitzenquerader keinen winkeligen Bogen bildend. Schwinger und Beine lebhaft gelb, in's Rostgelbe fallend. Hinterste Fiisse schwiirzlichbraun. (In meiner Samrn- luug.) (Translation.') — Golden-green with handsomely yellow feet ; length two lines, 9. (New York.) Antennae black. Face green, with a silvery-white reflection. Thorax handsome green. Abdomen golden-green. Wings not pictured ; veins brown ; upper branch of the fourth longitudinal vein not forming an angular curve. Halteres and feet bright-yellow, somewhat reddish-yellow. Hind tarsi blackish-brown. (In my own collection.) Page 226. No. 2§. Psilopus femora* us* jEneo-viridis, antennis nigellis ; pedibus flavis, femoribus basi virentibus ; alis limpidis. Lebhaft erzgruu, mit schwarzlichen Fiihlern, gelben Beinen, an der Wurzel griiulichen Schenkeln und wasserklaren Fliigeln. Lange 1| Linien. Aus Pennsylvanien. Auch diese kleine Art zieht der Farbe nach stark in's Griin- goldene, zuweilen an Stirn uiid Riickenschild iu's Blauliche. Un- tergesicht und Brustseiten weisschimraernd oder bereift. Riissel gelblich. Fliigel ohne alle Zeichnung. Beine bleich gelb. 296 DIPTERA OF NORTH AMERICA. [PART II. Schenkel griinlich, die vordern mit gelber Spitze. (In meiner Sammlung.) (Translation.) — Brilliant -bronze-green, with blackish antennae, yellow feet, femora greenish at the basis, and hyaline wings. Length one and one-half 'lines. (Pennsylvania.) The color of this small species also approaches the golden-green ; front and thorax are sometimes bluish. Face and pleurae have a white reflec- tion or are pruinose. Proboscis yellowish. Wings not pictured. Feet pale-yellow. Femora greenish, the anterior ones with yellow tip. (In my own collection.) Page 227. No. 3O. Psiloptis mundiis. Omnino chalybeus, vena apicali in angulum obtusum flexa. Ueberall stahlblau, mit stuinpfwinkelig gebogener Spitzen- querader. Lange 1^ Linien. Von Savannah. Fiihler schwarz ; Untergesicht uud Stirn salt stahlblau, der unterste Theil jenes nur sehr wenig weiss-schimmernd. Mittel- und Hinterleib satt stahlblau, an den Seiten hin und wieder griin- lich, doch so, dass ich kaum glaube, dass es ganz griine Abiinder- ungen gebe. Fliigel ungefarbt. Die Spitzenquerader bildet keinen so gleichformig gekriimmten Bogen, wie bei P. virgo, sondern raacht einen stumpfen Winkel. Schwinger gelb. Beine schwarz, an den Schenkeln stahlblaulich. (In meiner Sammlung.) (Translation. ) — Altogether steel-blue, with the anterior branch of the fourth longitudinal vein angularly curved. Length one and one-half lines. (Savannah.) Antennae black ; face and front saturate steel-blue ; the lower part of the former with only very little white reflection. Thorax and abdomen saturate steel-blue, in some places on the sides greenish, still so as to make me doubt whether there are entirely green varieties. Wings not pictured. The branch of the fourth vein is not regularly arched as in P. virgo, but forms an obtuse angle. Halteres yellow. Feet black, femora steel-blue. (My own collection.) Page 232. ]Vo. 6. Dolicliopus obs curias SAY. JrSneo nigellus ; capite niveo-micante ; alis infumatis, pedibus flavis. Erzgriinlich-schwarz, mit schneeweiss schirnmerndem Kopfe, rauchgraulichen Fliigeln und gelben Beinen. Lange 1^ Linien, 9. Aus Pennsylvanien. Fiihler tief schwarz ; Untergesicht und Stirn schwarz, beide APPENDIX. 297 schneeweiss schimmernd. Riickensckild aus dem Schwarzlicheu in's Griine und rothlich stahlblaue spieleud ; Brustseiten weiss- bereift. Hinterleib metallisch schwarzlick, in's duster griinliche ziehend. Fliigel salt rauckgrau ; Schwinger uud Beine gelb (Im Philadelphisclieu Museum.) (Translation.) — Blackish bronze-green ; head with a snow-white reflec- tion ; wings smoky-gray ; feet yellow. Length one and one-half lines, J . (Pennsylvania.) Antenna deep-black ; face and front black, both with a silvery- white reflection. Thorax changing from the blackish into green and reddish steel-blue ; pleurae pruinose with white. Abdomen metallic blackish, with a dusky greenish reflection. Wings saturate smoky-gray ; halteres and feet yellow. (Museum of Philadelphia.) Macquart, Dipteres, Suites a Buff on, Vol. 7. Page 450. No. 6. Psilopus radians. Long 2^ lig. D'un vert ou d'un bleu violet, tres brillant. Antennes noires ; deuxieme article garni de soies allonge'es, rayon- nant a 1'entour ; style fort allonge. Bord des segmens de 1'abdo- men noir ; organe copulateur peu epais. Pieds noirs ; jambes ante'rieures et intermediates jaunatres. Bord exterieur des ailes un peu brunatre vers 1'extremite. De PAmerique septentrionale. (Cabinet de M. Perc'heron.) (Translation.) — Length two and one-half lines. Purplish-green or blue, very brilliant. Antennae black ; second joint fringed with elongated, radiating bristles ; arista very long. Borders of abdominal segments black ; genital organs not very stout. Feet black ; fore and middle tibise yellowish. Anterior margin of the wing somewhat brownish towards the tip. North America. (Mr. Percheron's collection.) Macquart, Dipteres exotiques, Vol. II. Page 119. No. 11. Psilopus siplio. Lsete seneus. Alis abbreviato-bifasciatis. Antennis nigris. Pedibus flavis ; femoribus nigris g ; tibiis posticis nigris g. (Tab. 21, fig. 1.) Wiedemanu a decrit cette espece comme ayant les pieds jaunes dans les deux sexes. Suivant nos observations sur un assez grand nombre d'iudividus, les femelles seules ont les pieds de cette couleur, a 1'exception des tarses noirs ; les males ont les cuisses noires, ainsi que les jambes poste'rieures. 298 DIPTERA OF NORTH AMERICA. [PART II. Cette espece est commune et repandue dans uue grande partie de PAmerique. Nous en avons vu des individus de la Pensyl- vanie, de Cuba, de la Guyane et des differentes parties du Bresil. (Translation.') — Bright bronze-green. Wings with two abbreviated fasciae. Antennae black. Feet yellow. Femora black ( £ ) ; hind tibiae black ($). (Tab. XXI, fig. 1.) Wiedemann described this species as having yellow feet in both sexes. According to my observations on a considerable number of specimens, the females alone have the feet of this color, with the exception of the black tarsi ; the males have black femora and hind tibiae. This species is common in a considerable part of America. We have specimens from Pennsylvania, Cuba, Guyana, and from different parts of Brazil. Page 121. IVo. IS. Psilopus radians. Laete-asneus. Antennarum articulo secundo setis elongatis radiatis ; stylo longissimo. Long. 2£1. J. D'un vert ou d'un bleu violet tres brilliant. Antennes noires; deuxieine article garni de soies allongees, rayonnant a 1'entour ; style fort allonge. Bord des segments cle 1'abdomen noirs ; organe copulateur peu epais. Pieds noirs, janibes posterieurs et inter- mediaires jaunatres. Bord exterieur des ailes un peu brunatre vers 1'extremite. De PAmerique septentrionale. (Cabinet de M. Percheron a Paris.) {Translation.') — Bright bronze-green. Second joint of the antennae with elongated, radiating bristles ; arista very long. Length two and one-half lines, $ . Purplish-green or blue, very brilliant. Antennae black. Second joint fringed with elongated, radiating bristles. Style very long. Margin of the abdominal segments black. Hypopygium not stout ; feet black ; intermediate and hind tibiae yellowish. Anterior margin of the wings somewhat brownish towards the tip. North America. (Collection of M. Percheron in Paris.) Macquart, Dipteres exotiques, Suppl. IV. Page 124. No. 2. Clirysotus viridifemora. Yiridi aureus. Antennis nigris. Pedibus rufis, femoribus viridibus. (Tab. 12. fig. 3.) Long, une 1. 9. APPENDIX. 299 Face et front verts, a duvet blaiic. Antennes et style noirs. Thorax et abdomen d'un vert clore. Cuisses d'un vert brillant, a genoux jaunes ; janibes fauves ; tarses d'un jaune brunatre ; posterieurs bruns, a premier article jaune. Ailes claires. De 1'Amerique septentrionale. (Collection des M. Hoffmeister de Nordhausen.) (Translation.} — Golden-green. Antennae black. Feet rufous, femora green. (Tab. XII, fig. 3.) Length one line, $>. Face and front green, with white down. Antennse and arista black. Thorax and abdomen golden-green. Femora brilliant-green ; knees yellow ; tibiae fulvous ; tarsi brownish-yellow ; the hind ones brown, with the first joint yellow. Wings hyaline. North America. (Collection of Mr. Hoffmeister in Nordhausen.) Page 128. No. 5. Dolicliopus lieteroneurus. .J^neo-viridis. Thorace vittis violaceis. Pedibus flavis. Alis cellula postica prima subclausa. (Tab. 12, fig. 10.) Long. 1^ 1.. Palpes noirs. Face et front larges, d'un vert noiratres, a le'ger duvet gris. Antennes : les deux premiers articles fauves ; premier un peu allonge et menu ; troisieme ovale, assez large, noir, a base fauve ; style noir, peu allonge. Thorax d'un vert fonce, a bandes violettes. Abdomen vert ; ventre a duvet blanc. Pieds jaunes, hanches ante'rieures noiratres ; un peu de brun h 1'extremite des cuisses ; tarses bruns. Ailes assez claires ; premiere cellule poste- rieure presque ferme'e ; deuxieme nervure transversale eloiguee du coude. De 1'Amerique septentrionale. (Collection de M. Hoffmeister de Nordhausen.) (Translation.) — Golden-green. Thorax with violet stripes ; feet yellow. Wings with the first posterior cell almost closed. (Tab. 12, fig. 10.) Long. lin. one and one-half. Palpi black. Face and front broad, blackish-green, with a slight gray down. Antennse : two first joints fulvous : the first somewhat prolonged and slender ; the third oval, rather broad, black, with fulvous basis ; arista black, not very long. Thorax dark-green, with violet stripes. Abdomen green, venter with whitish down. Feet yellow, anterior coxae blackish ; tip of femora somewhat infuscated ; tarsi brown. Wings rather hyaline ; first posterior cell almost closed ; second transverse vein at some distance from the flexure of the fourth vein. Bab. North America. 300 DIPTERA OP NORTH AMERICA. [PART II. Walker, Insecta Saundersiana. Page 207. Psilopus lepidus MAS. Yiridis, abdominis segmentorum marginibus anticis nigris, antennis pedibusque nigris, alis limpidis fusco-bifasciatis. Bright-green, beset with black bristles ; head blue, tinged with purple, adorned in front with white bloom ; eyes bright-red ; mouth black ; feelers black ; sixth joint as long as the chest ; chest not shining, tinged with blue ; breast with a hoary covering ; abdomen tapering from the base to the tip, narrower and much longer than the chest ; a black baud on the fore border of each segment ; legs black, clothed with biack hairs and bristles ; wings colorless, adorned with two dark-brown bands which are united on the fore border ; tip cross-vein forming a right angle, whence* it is indistinctly waving to the tip of the wing ; lower cross-vein very slightly waving ; wing-ribs and veins black. Length of the body two and one-half lines ; of the wings six lines. Mexico. Page 211. Medeterus exustus FEM. Nigro-aeneus, capite antico viridi ; abdomine purpureo cupreo, antennis pedibusque nigris, femoribus seneis, alis cinereis, costa venarumque marginibus fuscis. Allied to M. notatus. Bronze-black, beset with black bristles, which form a cross-row behind the head, adorned beneath with a whitish covering ; head green in front, clothed beneath with black hairs ; peristoma prominent ; eyes red ; facets of the fore part rather large ; feelers black ; abdomen obconical, coppery with a slight purplish tinge, not longer than the chest ; legs long, black, thickly clothed with black down, beset with a few black bristles ; thighs brassy, armed with black spines ; foot-cushions dark-tawny ; wings darkish-gray, very dark-brown beneath the fore border and along the borders of the veins in the disk ; wing-ribs and veins black; poisers dark-tawny with pitchy knobs. Length of the body two and one-half lines ; of the wings six lines. Bolton, North America. APPENDIX. 301 Page 212. Medeterus viridiflos FEM. Fulvo-viridis, thoracis disco fusco, antennis-pedibusque nigris, femoribus viridibus, alis subcinereis ad costam subfuscis. Pale grassy-green, tinged with tawny above and with whitish bloom beneath, beset with a few black bristles ; head brown about the eyelets, beset behind the eyes with a row of black bristles, clothed beneath with white hairs ; eyes red, thickly clothed with short, white hairs ; feelers black ; disk of the chest brownish ; abdomen obconical, longer than the chest, clothed with very short white hairs ; legs black, clothed with very short black hairs, beset with black bristles ; thighs green, with which color the shanks are also tinged ; wings slightly gray, tinged with pale-brown beneath the fore border ; wing-ribs and poisers tawny ; veins black, tawny at the base Length of the body one and one-half lines ; of the wings four and one-half lines. North America. Page 212. DoiicSnopus Mfrons FEM. ^Eneo-viridis, capitis vertice cyaneo, abdominis segmentorum marginibus anticis cupreis, anteunis fulvis, pedibus flavis, alis subcinereis. Green, beset with black bristles, adorned beneath with a white covering ; crown of the head blue, tinged with green and purple ; an olive stripe between the feelers and the epistoma ; eyes red ; mouth pitchy. Feelers tawny ; third joint pitchy towards the tip ; sixth black, feathered ; a brassy tinge on the chest ; abdomen obconical, longer than the chest, coppery on the fore border of each segment whose sides are adorned with a white covering ; legs yellow, clothed with very short black hairs, which as usual are most frequent on the feet ; thighs stout, shanks beset with black bristles ; wings very slightly gray ; wing-ribs tawny ; veins black, tawny at the base, poisers yellow. Length of the body one and one-half to one and three-fourth lines ; of the wings three and one-half to four lines. United States. 302 DIPTERA OF NORTH AMERICA. [PART II. Page 213. Dolicliopus censors FEM. JSneo-viriclis, vertice purpureo, thorace antico cyaneo, abdominis suturis nigris, antennis pedibusque fulvis, alls subcinereis. Green, beset with black bristles, adorned beneath with a whitish covering ; crown purple ; eyes black ; mouth tawny ; feelers tawny ; sixth joint black, feathered with much shorter hairs than those of D. bifrons; chest brassy-green, blue in front; abdomen obconical, clothed with short, black hairs, a little longer than the chest, sutures of the segments blackish ; legs tawny, clothed with very short black hairs ; shanks beset with black bristles ; wiugs grayish ; wing-ribs tawny ; veins black, tawny at the base ; poisers tawny with yellow knobs. Length of the body one and one- fourth line ; of the wings three lines. United States. Page 213. Dolicliopus contingens FEM. Viridis, vertice purpureo, antennis pedibusque fulvis, alis subcinereis. Green, beset with black bristles, adorned beneath with a whitish covering ; crown purple ; eyes black ; mouth tawny ; feelers tawny, sixth joint black, feathered like that of D. consors ; abdo- men obconical, clothed with short black hairs, a little longer than the chest ; legs tawny, clothed with very short black hairs ; shanks beset with black bristles ; wings grayish ; wing-ribs tawny ; veins black, tawny at the base ; tip cross-vein less angular than that' of D. consors ; poisers dark-tawny. Length of the body one and one-fourth line ; of the wings three lines. United States. Page 213. Dolicliopus lieoes FEM. ^Eneus, vertice cyaneo, abdominis segmentorum marginibus posticis viridibus, antennis pedibusque fulvis, alis cinereis costa venarumque marginibus fuscis. Brassy, beset with black bristles, adorned beneath with a whitish covering; crown of the head blue; eyes red; feelers tawny, sixth joint black, feathered with very short hairs ; abdo- men obconical, longer than the chest ; hind borders of the seg- ments green ; legs dark-tawny, clothed with very short black APPENDIX. 303 hairs ; shanks beset with black bristles ; wings gray, brown beneath the fore border, and along the borders of the veins ; wing- ribs and veins black ; poisers tawny with pitchy knobs. Length of the body one and one-fourth line ; of the wings three lines. United States. Page 214. Dolichopus ineptus FEM. JEneus, vertice purpureo, abdominis lateribus albo maculatis, apice viridi, antennis pedibusque fulvis, tarsis subpiceis, alls cinereis fusco vittatis. Brassy, beset with black bristles, adorned beneath with a whitish covering ; crown purple ; eyes red ; feelers tawny, sixth joint black, feathered with moderately long hairs ; abdomen obconical, longer than the chest, green at the tip ; a white spot on each side of every segment ; legs dark-tawny, clothed with very short black hairs ; shanks beset with black bristles ; feet almost pitchy ; wings gray, tinged with brown along the third and fourth longitudinal veins ; wing-ribs tawny ; veins black ; poisers tawny, with ferruginous knobs. Length of the body one and one-fourth line ; of the wings three lines. United States. Page 214. Dolichopus maculipes FEM. Aeneo-viridis purpureo varius, antennis fulvis apice piceis, pedibus fulvis, tibiis nigro maculatis, tarsis piceis, alis cinereis costa nervorumque marginibus fuscis. Brassy-green, beset with black bristles, adorned beneath with a whitish covering ; crown adorned with blue and purple ; eyes red ; feelers tawny ; third joint pitchy ; sixth black, feathered with moderately long hairs ; disk of the chest partly purple ; abdomen obconical, a little longer than the chest, clothed with ' short black hairs ; legs tawny, clothed with very short black hairs ; shanks beset with black bristles ; feet almost pitchy ; a black spot on the tip of each thigh ; five or six black spots on each shank, these spots are most distinct on the hind legs ; wings gray, brown beneath the fore border and along the borders of the veins ; wing-ribs and poisers tawny ; veins black. Length of the body one and one-half line ; of the wings three and one-half lines. United States. 304 DIFTERA OF NORTH AMERICA. [PART II. Page 215. Dolicliopus pulchcr, Mas. et Fern. Cyaneo-viridis, antennis uigris, femoribus viridibus, tibiis ful- vis, tarsis piceis apice nigris, alls limpidis. Bright green with a bluish tint, beset with black bristles, adorned beneath with a whitish covering ; eyes bright red ; feelers black; sixth joint bare; abdomen of the male cylindrical, of the female obconical, clothed with short black hairs, a little longer than the chest ; scales of the male white, bordered with black ; legs tawny, clothed with very short black hairs ; thighs green ; shanks beset with black bristles ; feet pitchy, black towards the tips ; trochanters of the male yellow, of the female tawny; wings colorless ; wing-ribs tawny ; veins black ; fourth longitudinal vein not as usual converging to the third after its curve, but almost parallel to it ; poisers yellow. Length of the body 1^ line ; of the wings. 3 lines. United States. Page 215. Dolichopus varius, Fern. ^Eneo-viridis cyaneo varius, abdomine fasciis albidis ornato, antennis pedibusque fulvis, tarsis nigris, alis cinereis, costa macu- lisque quinque fuscis. Brassy-green, tinged with blue, beset with black bristles, adorned beneath with a whitish covering ; eyes red ; feelers tawny ; sixth joint black, feathered with moderately long hairs ; abdomen obconical, longer than the chest, adorned with bands of whitish hue ; legs dark tawny ; clothed with short black hairs ; shanks beset with black bristles ; feet black ; wings gray, brown beneath the fore border, adorned with four or five brown spots ; wing-ribs pitchy ; veins black ; poisers tawny. Length of the body l£ line, of the wings 3 lines. United States. Walker, List of Dipterous Insects in the Collection of the British Museum, Part III. Page 645. Psilopus delicatus, n. s., Fern. Viridis, gracilis, abdomine aureo-viridi, antennis fulvis, articulo tertio nigro, pedibus flavis, alis limpidis. Body slender, bright green, clothed with black hairs and bris- Ai-PENinx. 305 ties ; crown of the head blue ; eyes bright red ; mouth yellow ; feelers tawny ; third joint black; bristle black, shorter than the chest ; abdomen golden-green ; legs pale yellow, long and slender, clothed with short black hairs ; feet pitchy towards the tips ; wings colorless ; wing-ribs and poisers tawny ; veins pitchy. Length of the body 2^ lines, of the wings 5 lines. a. New York. Presented by the Entomological Club. b. New York. Presented by E. Doubleday, Esq. Page 646. Psilopus gemmifer, n. s., Mas. Yiridis, cyaneo purpureoque varius, anteunis nigris, pedibus flavis, alis limpidis, costa apicem versus nervisque transversis fusco-nebulosis. Body bright green, beset with black hairs and bristles ; head covered in front with silvery down ; eyes red ; mouth tawny ; feelers black, as long as the head and the chest, disk of the chest bluish-green, tinged with purple ; scutcheon purple ; sides and breast covered with silvery bloom ; abdomen slender ; tip bluish purple ; appendages dark tawny ; legs yellow, beset with black hairs and bristles, which are most thick on the feet ; four hinder hips green ; thighs clothed with white hairs ; feet towards the tips and hind feet pitchy ; wings colorless, clouded with pale brown towards the tips of the fore borders and along the cross-veins ; wing-ribs tawny; veins black; poisers tawny. Length of the body 2i lines, of the wings 5^ lines. a. Trenton Falls. Presented by E. Doubleday, Esq. Page 646. Psilopus cJirysoprasi, n. s. Aureo-viridis, capitis vertice purpureo-cyaneo, scutello abdo- minisque basi purpureis, abdominis segmentorum suturis asneo- purpureis, antennis nigris, pedibus piceis, femoribus viridibus, tibiis anterioribus tarsisque anticis fulvis, alis subciuereis. Body golden-green, beset with black bristles ; head purplish- blue on the crown, slightly covered with white down in front ; eyes bright red ; mouth and feelers black ; scutcheon purple ; ab- domen adorned with purple towards the base ; sutures of the seg- ments brassy-purple ; legs pitchy, thickly clothed with short black- hairs ; hips and thighs green ; hips slightly covered with white bloom, thighs fringed with white hairs ; fore shanks pale tawny ; middle shanks and fore feet dark tawny ; wings slightly gray ; 20 306 DIPTERA OF NORTH AMERICA. [PART II. wing-Tibs and poisers pitchy ; veins black. Length of the body 2 lines, of the wings 4^ lines. a. "West Indies. From Mr. Children's collection. Page 648. Psilopus siiavium, n. s. Yiridis, capitis vertice cyaneo-purpureo, abdomine apicem ver- sus purpureo, antennis pedibusque nigris, feraoribus viridibus, alis limpidis, fusco bifasciatis. Body bright green, beset with black bristles ; head bluish-pur- ple on the crown, clothed with white down in front ; eyes red ; mouth pitchy ; feelers black ; bristle nearly as long as the chest ; breast and sides of the chest covered with a white bloom ; abdo- men purple towards the tip ; legs black, beset with black hairs and bristles ; hips and thighs green ; hips covered with a white bloom ; thighs clothed with white hairs ; wings colorless, adorned with two brown bands, which are joined together on the fore border and more slightly on the disk, but do not reach the hind border nor the tip ; veins black ; wing-ribs and poisers pitchy. Length of the body 2^ lines, of the wings 5 lines. a. Jamaica. From Mr. Grosse's collection. Page 648. Psilopus amatms, n. s., Mas. et Fern. Yiridis, capite purpureo, abdomiue nigro-fasciato apice pur- pureo, antenuis nigris, pedibus piceis, fernoribus viridibus, alis sub- cinereis, costa apicem versus nervisque transversis fusco nebulosis. Body bright green, beset with black hairs and bristles ; head purple, covered in front with white down ; crown of the male adorned with a green spot on each side ; eyes red ; mouth and feelers black ; bristle a little shorter than the chest ; hind part of the chest tinged with blue and purple ; sides and breast covered with white down ; abdomen at the tip purple in the male, bluish- purple in the female ; hind borders of the segments in the male adorned with black bands ; legs pitchy, beset with black hairs and bristles; hips and thighs green, the former covered with white down ; wings slightly gray, indistinctly marked with brown towards the tips of the fore borders and along the cross-veins ; wing-ribs pitchy ; veins black ; poisers of the male pitchy, of the female tawny. Length of the body 1^ — If line, of the wings 3| — 4 lines. a. New York. Presented by E. Doubleday, Esq. b. Trenton Falls. Presented by E. Doubleday, Esq. APPENDIX. 307 Page 649. Psilojms i illicit us, n. s. Yiridis, capite purpureo, abclomine purpureo,* abdomine pur- pureo-cyaueo, fasciis nigris, auteunis pedibusque nigris, alis sub- cinereis fusco bifasciatis. Head and chest beset with black bristles ; head purple, fringed about the mouth with hoary hairs ; eyes red ; mouth and feelers black ; bristle a little longer than the chest ; chest green ; sides and breast covered with whitish down ; abdomen deep purplish- blue ; sutures of the segments black ; legs black, clothed with black hairs and bristles ; wings slightly gray, adorned with two brown bands, which are united on the fore border, but do not reach the hind border; wing-ribs and veins black; poisers pitchy, with tawny knobs. Length of the body 2J? lines, of the wings 5 lines. a. Mexico. Presented by E. P. Coffin, Esq. Page 650. Psilopus nigrofemoratus, MSS. Cyaneo-, aut aureo-viridis, antennis nigris, capite duplo longi- oribus, pedibus nigris tibiis fulvis, alis limpidis. Head and chest bright bluish-green, armed with black bristles ; head covered in front with white down, clothed beneath with white hairs ; eyes red ; mouth tawny ; feelers black, about twice the length of the head ; breast and under side of the abdomen covered with white bloom ; abdomen golden-green, blue at the base, coppery at the tip ; legs black, beset with a few black bristles ; shanks tawny, with black tips ; wings colorless ; wing-ribs tawny ; veins black ; poisers vellow. Length of the body 1^ line, of the wings 2^ lines. Tar. 3. Chest golden-green, bluish-green behind ; abdomen coppery-green ; tips of the thighs and the whole of the shanks tawny. Var. y. Abdomen bright green or bluish-green ; a bronze band on the fore border of each segment. a. North America. Presented by the Entomological Club. b. Nova Scotia. From Lieut. Redman's collection. Page 651. Psilopug allbicoxa Mas. et Fern. Cyaneo-, aut cupreo-viridis, antennis nigris, capite duplo longi- oribus, pedibus flavis, tarsis posticis nigris, alis subcinereis. Head and chest armed with black bristles ; head bluish-green, * Evidently a misprint in the original. 308 DirTERA OF NORTH AMERICA. [PART II. covered in front with white down, clothed beneath with white hairs ; eyes red ; mouth tawny ; feelers black, about twice the length of the head ; palpi black ; chest of the male bluish-green, sometimes black towards the tip, of the female bright green or coppery-green ; sutures of the segments sometimes black ; breast and under side of the abdomen covered with white down ; legs yellow, adorned with rows of minute spines, clothed with a few white hairs, and beset with a few black bristles ; four hinder hips green ; tips of feet black ; hind feet black, first joint brownish ; wings slightly gray, wing-ribs tawny ; veins black ; poisers yellow. Length of the body 1^ line ; of the wings 2^- lines. a. North America. Presented by the Entomological Club. b. Ohio. Presented by the Entomological Club. c. Massachusetts. From Prof. Sheppard's collection. d. Nova Scotia. From Lieut. Redman's collection. Page 651. Chrysotus iiicertus, n. s. Yiridis, antennis nigris, femoribus viridibus, tibiis fulvis, lapice tarsisque obscurioribus, alis limpidis. Allied to C. femoralis. Body green, beset with black hairs and bristles ; eyes red ; mouth and feelers black ; hips and thighs green ; shanks tawny ; feet and tips of shanks dark tawny ; wings colorless ; wing-ribs pitchy ; veins black ; poisers tawny. a. United States. . Presented by the Entomological Club. Page 653. Porplayrops pilosicomis, BAKNSTON'S MSS. JEneo-viridis, antennis nigris, pedibus fulvis, tarsis piceis, femoribus posticis apice fusco maculatis, alis limpidis. Body brassy-green, beset with black bristles ; eyes dark-red ; mouth dark tawny ; feelers black ; bristle downy, proceeding from the base of the third joint and more than twice its length ; breast and sides of the chest covered with a white bloom, which appears also on the chest, but is there very slight ; legs tawny, clothed with short black hair, beset with a few black bristles ; feet pitchy ; a small brown mark on the tip of each hind thigh ; fore hips at the base and the other hips green and covered with white bloom ; wings colorless ; wing-ribs tawny.; veins black ; poisers yellow Length of the body l£ line ; of the wings 2£ lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. APPENDIX, 309 Page 655. Medeterus glaber, BARNSTON'S MSS. Viridis, thoracis disco nigro-ceneo, abdominc seneo-viridi, anten- nis nigris, pedibus viridibus, tarsis nigris, alls cinereis, fusco biraa- culatis. Body green ; head and chest beset with a few black hairs ; head covered with white bloom in the male, with golden bloom in the female ; eyes red ; mouth and feelers black ; disk of the chest bronze-black ; sides covered with tawny bloom ; breast covered with white bloom ; abdomen brassy-green, covered above with short tawny hairs ; under side covered with white bloom ; legs bright green, rather stout, clothed with short black hairs and bristles ; hips covered with white bloom ; feet black ; wings gray ; each with two small brown spots, one on the cross-vein, the other on the fourth longitudinal vein, a little before half the distance between the cross-vein and the tip of the wing ; wing-ribs pitchy ; veins black ; poisers tawny. Length of the body If line ; of .the wings 5 lines. a. St. Martin's Falls, Albany River, Hudson's Bay, Presented by Gf. Barnston, Esq. Page 655. Medeterus clirysologus, BARNSTON'S MSS., Fern. Nigro-asneus, antennis nigris, pedibus viridibus, tarsis nigris, alis cinereis, fusco bimaculatis, ad costam sulbfuscis. Body brassy black ; head covered with golden bloom, which is paler and brighter towards the mouth ; eyes dark-red, covered with white down ; mouth and feelers black ; sides of the chest covered with tawny bloom ; breast and under side of the abdomen adorned with white bloom ; a row of black punctures on each side of the abdomen, as in other species ; legs green, clothed with black hairs and bristles ; hips and thighs covered with white bloom ; feet black ; wings gray, brown along the fore borders, each with two darker brown spots, like those of M. glaber, but larger and more distinct ; wing-ribs tawny ; veins black ; poisers pitchy. Length of the body 1^ line ; of the wings 3 lines. a. St. Martin's Falls, Albany lliver, Hudson's Bay. Presented by G. Barnston, Esq. 010 DIPTERA OF NORTH AMERICA. [PART II. Page 656. Medeterus atJjoflorens, n. s., Fern. ^Eneus, fulvo pubescens, subtus albus, abdoinine cupreo-viridi, anteunis iiigris, pedibus viridibus, tarsis piceis, alls cinereis fusco subvittatis. Head and chest bronzed, beset with a few black bristles, thickly covered with tawny bloom ; eyes dark red, covered with white down ; mouth and feelers black ; abdomen rather light green, mingled with copper-color, thinly clothed with short black hairs, not longer than the chest ; hind chest, breast, and under side of the abdomen covered with white bloom ; legs long, slender, green, slightly covered with tawny bloom, beset with short black hairs and bristles ; feet pitchy towards the tips ; claws black ; foot-cushions pale yellow ; wings gray, very slightly clouded with brown along the borders of the veins ; wing-ribs pitchy ; veins black ; poisers tawny, pitchy, and covered with white bloom towards the tips. Length of the body 1^-H line ;. of the wings 4-4^ lines. a. Nova Scotia. From Lieut. Redman's collection. Page 659. DolicBaopus affiiais, HALIDAY'S MSS., Mas. et Fern. Cyaneo-, aut cupreo-viriclis, antennis nigris, pedibus flavis, femo- ribus posticis tarsisque fulvis, tarsis anticis, mari. apice nigris dilatatis, tarsis posticis nigris, alis limpidis. Male. — Head and thest green, armed with stout black bristles ; head fringed behind with pale tawny hairs, covered in front and beneath with white bloom; eyes red, covered with white down; feelers black ; chest bluish-green on the disk ; abdomen coppery- green, clothed with short black hairs, covered with white bloom beneath and on each side, where there is a row of black punctures ; appendages pale yellow ; legs yellow, clothed with short black hairs ; four hinder hips green ; hind thighs tawny, furnished with a few tawny hairs ; shanks and feet armed with black bristles ; feet tawny, darker towards the tips ; hind feet black ; tips of fore feet black, widened ; wings colorless ; wing-ribs and poisers yel- low ; veins pitchy. Female. — Body coppery-green ; legs tawny ; shanks darker than the thighs ; hind feet pitchy. Length of the body 2-2£ lines; of the wings 4-5 lines. a. Nova Scotia. From Lieut. Redman's collection. APPENDIX. 311 Page 660. Dolichopus cuprinus? WIED., Auss. Zweif. ii, 230, 1. D. cupreus ? SAY, Journ. Acad., Phila., iii. 8, 6. Aureo-viridis, thorace vittis tribus cupreis, antennis fulvis apice fuscis, pedibus flavis, tarsis apice nigris, anticis apice nigro-fasci- culatis, alis subcinereis. Body green, covered above with golden down, beneath with white down ; head and chest armed with stout black bristles ; hind part of the head fringed with white hairs ; eyes red ; mouth yellow ; feelers tawny, covered with short black hairs ; their tips brown ; bristle pubescent, pitchy, longer than the rest of the feelers ; chest adorned with three coppery stripes ; abdomen golden green, clothed with short black hairs, white beneath and on each side, where it 1ms a row of black punctures ; appendages yellow ; legs yellow, clothed with black hairs ; four hinder hips green ; feet towards the base and shanks armed with black bristles ; four hinder feet black towards the tips ; tips of fore feet adorned with tufts of black hairs ; wings slightly gray ; wing-ribs tawny ; veins pitchy ; poisers yellow. Length of the body. 2|- lines ; of the wings 5 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. • Page 660. Dolicliopus lamellipes, BARNSTON'S MSS., Mas. et Fern. Yiridis aut viridi-cupreus, abdomine subtus albo-pubescente, antennis nigris basi runs, articulo tertio mari. longo, pedibus fulvis, tarsis nigris, intermediis basi fulvis, anticis mari. apice dilatatis, alis limpidis. Body green ; head and chest armed with black bristles ; head covered with silvery down, fringed behind with black hairs ; eyes red ; mouth tawny ; feelers black ; first joint pale red ; third joint very long ; abdomen clothed with black hairs, adorned with a coppery tinge towards the tip, which is black, covered with white bloom beneath and on each side, where there is a row of black punctures ; legs pale tawny, four hinder hips green, clothed with black hairs and bristles ; feet black ; tips of fore feet widened ; middle feet tawny towards the base ; wings colorless ; wing-ribs tawny ; veins pitchy ; poisers pale tawny. Female. — Third joint of the feelers short, nearly round. ' Length of the body 2^-3 lines ; of the wings 5 lines. 312 DIPTERA OF NORTH AMERICA. [PART II. Yar. j3. Chest and abdomen coppery. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 661. Dolicltopus ciliatus, BAKXSTON'S MSS., Mas. et Fern. Yiridis, capitis fronte aureo-pubescente, antennis fulvis, arti- culo 3° supra nigro, pedibus fulvis, tarsis piceis alis subcinereis. . Body green ; head covered in front with golden down ; eyes red ; mouth black ; feelers tawny ; third joint nearly oval, black from near the base to the tip above, and from half its length to the tip beneath ; Tbristle black ; breast and sides of the chest and of the abdomen covered with white bloom ; appendages of the abdomen tawny ; scales white ; legs tawny ; feet pitchy ; fore feet dark tawny ; wings slightly gray ; wing-ribs tawny ; veins black ; poisers yellow. Length of the body 1^-lf line ; of the wings 3 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 661. Dolicltopiis adjaceiis, n. s., Fern. ^Eneus viridi varius, capite cyaneo-viridi, antennis nigris, pedi- bus fulvis, tarsis posticis piceis, alis subcinereis. Head bluish-green on the crown, covered in front with yellowish white down, fringed along the eyes with hoary hairs ; eyes red ; mouth pitchy ; palpi tawny ; feelers black ; third joint very short ; chest and abdomen brassy, mingled with green ; breast and sides of the chest covered with gray bloom, which also appears beneath the abdomen, but is more slight ; legs tawny ; hips green, covered with gray bloom ; fore hips mostly tawny ; feet darker than the shanks, especially towards the tips ; hind feet pitchy ; wings slightly gray ; wing-ribs and poisers tawny ; veins black. Length of the body 3 lines ; of the wings 6 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 661. Dolichopus coercens, n. s., Mas. Yiridis, capite cyaneo-viridi, thoracis lateribus cupreo etcyane< ornatis, abdomine cupreo vario, apice nigro, antennis nigris, pedi- bus fulvis, alis limpidis. APPENDIX. 313 Head bluish-green, covered iu front with pale tawny down, fringed along the eyes with white hairs ; eyes bright red ; mouth pitchy ; feelers black ; third joint very short ; chest green, slightly tinged on each side with blue and copper color ; disk sometimes bluish-green ; breast covered with hoary down ; abdomen green, with a coppery tinge here and there ; tip black ; appendages pale tawny ; scales white, bordered with black ; legs tawny ; hips, towards the base, green, and covered with a white bloom ; shanks beset with black bristles ; feet pitchy towards the tips ; fore feet slender, pale tawny ; their tips black, and much widened ; wings colorless ; wing-ribs and poiser tawny ; veins black. Length of the body 3 lines ; of the wings 5|- lines. a. New York. Presented by the Entomological Club. Page 662. Dolichopus finitus, n. s., Mas. Viridis, thoracis lateribus abdomineque cupreo variis, hujus lateribus basi cyaneo-viridibus, autennis nigris, pedibus fulvis, tarsis apice nigris, tarsis anticis apice latis, tarsis posticis nigris, alis subcinereis. Body green ; head covered in front with white down ; fringed along the eyes with white hairs ; eyes red ; mouth pitchy ; palpi tawny ; feelers black ; third joint rather large ; chest with a slight coppery tinge on each side, which, like the breast, is slightly covered with hoary bloom; abdomen tinged with coppery color, and with a slight blue hue on each side towards the base ; append- ages at the tip tawny ; scales white, bordered with black ; legs tawny, middle feet towards the tips, and hind feet, excepting the base, black ; tips of fore feet black and somewhat widened ; wings slightly gray, wing-ribs and poisers tawny; veins pitchy. Fern. — Feet black, tawny at the base. Length of the body 3 lines ; of the wings 5 lines. a. New York. Presented by the Entomological Club. Page 662. Dolichopus distractus, n. s. Viridis, abdomine cupreo, antennis nigris, articulo 1° subtus fulvo, pedibus fulvis, tarsis apice piceis, alis subcinereis. Body green ; head covered in front with white down, clothed on each side of the eyes with white hairs ; eyes bright red ; feelers black ; first joint tawny beneath ; third joint short and broad ; 314 DIPTERA OP NORTH AMERICA. [PART II. abdomen copper colored ; legs tawny ; feet pitchy towards the tips ; wings slightly gray ; wing-ribs and poisers tawny ; veins black. Length of the body 2^- lines ; of the wings 5 lines, a. New York. Presented by E. Doubleday, Esq. Page G62. Dolichopus discessws, n. s., Fern. Cyaneo-yiridis, thorace cupreo bivittato, abdomine ffiueo-viridi, autennis pedibusque fulvis, tarsis posticis piceis, alis subcinereis. Body bluish-green ; head covered in front with white down ; eyes bright red ; mouth and feelers tawny ; bristle black ; chest adorned with two bright copper-colored stripes ; sides and breast covered with white bloom ; abdomen green, brassy here and there, especially towards the tip ; sides and under side covered with white down ; legs tawny ; tips of feet and hind feet, except the base, pitchy ; wings slightly gray ; wing-ribs and poisers tawny ; veins black. Length of the body 2^ lines ; of the wings 5^- lines. a. Massachusetts. From Prof. Sheppard's collection. Page 663. f&olichopus contigmis, n. s., Mas. Aureo-viridis, thorace viridi-cyaneo, lateribus purpureo variis, abdomine cyaneo et cupreo vario, antennis nigris, pedibus fulvis, tarsis anticis apice nigris latis, tarsis mediis piceis, basi fulvis, tarsis posticis nigris, alis limpidis. Head golden-green, covered in front with pale tawny down ; eyes bright red ; feelers black ; third joint oval ; chest greenish- blue, with a slight purple tinge on each side ; abdomen golden- green, slightly bluish and coppery here and there ; breast and under side of the abdomen covered with hoary bloom ; tip black ; appendages tawny, scales whitish ; legs tawny ; four hinder hips mostly green, and tinged with hoary bloom ; tips of fore feet black, much widened ; middle feet pitchy, tawny at the base ; hind feet black ; wings colorless ; wing-ribs and poisers tawny ; veins pitchy, tawny towards the base. Length of the body 2 lines ; of the wings 4 lines. a. New York. Presented by the Entomological Club. Page 663. Dolichopus excltisiis, n. s., Fern. Cupreus, nonnunquam viridi varius, abdomine purpureo-cupreo, antennis nigris, articulo primo subtus fulvo, pedibus fulvis, tarsis nigris, alis subcinereis. APPENDIX. 315 Body coppery, sometimes varied with green ; bead covered in front with white bloom ; eyes red ; mouth pitchy ; palpi tawny ; feelers black ; first joint tawny beneath ; third joint oval, as long as the first and the second ; breast covered with gray bloom ; abdomen purplish copper-color, covered beneath with gray bloom ; legs tawny ; hips coppery ; fore hips mostly tawny ; feet black ; wings slightly gray ; wing-ribs tawny ; veins black ; poisers pale tawny. Length of the body 2 lines ; of the wings 4^ lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 664. Dolichopus coiifm is, n. s., Fern. J^neus, viridi varius, capite viridi, antennis nigris, pedibus fulvis, tarsis piceis, tibiis posticis apice tarsisque posticis nigris, alis cinereis. Body brassy, mingled here and there with green ; head green, covered in front with white down, clothed along the sides of the eyes with white hairs ; eyes red ; mouth pitchy ; feelers black ; third joint nearly oval, rath er' short ; breast covered with white bloom ; legs tawny ; feet pitchy, tawny towards the base ; hind feet and tips of hind shanks black ; wings gray ; wing-ribs and poisers tawny ; veins black. Length of the body 2 lines ; of the wings 4 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 664. Dolichopus contemiinus, n. s., Mas. Tiridis, thoracis disco abdomineque aureo-viridibus, hujus apice seneo, antenuis nigris, pedibus fulvis, tarsis anticis apice nigris latis, tarsis mediis apice posticisque piceis, alis limpidis, fern. tarsis piceis basi fulvis, tarsis posticis nigris. Body bright green ; head covered with tawny down, fringed along the sides of the eyes with white hairs ; eyes bright red, covered with white down ; mouth pitchy ; palpi tawny ; feelers black ; third joint rather large ; abdomen and disk of the chest golden-green ; breast and sides of the chest covered with hoary bloom, which also slightly tinges the under side of the abdomen ; tip of the abdomen brassy ; appendages tawny ; scales white, slightly bordered with black ; legs pale bright tawny ; four hind hips mostly green, and covered with a white bloom ; thighs fringed o 16 DIPTERA OF NORTH AMERICA. [PART II. with white hairs, fore feet slender, with black and much widened tips ; middle feet pitchy towards the tips ; hind feet pitchy ; wing colorless ; wing-ribs tawny ; veins black ; poisers pale tawny. Fern. — Feet pitchy, tawny towards the base ; hind feet black. Length of the body 2 lines ; of the wings 4 lines. a. New York. Presented by E. Doubleday, Esq. &. North America. Presented by the Entomological Club. Page 665. Dolicltopus separatus, n. s., Fern. Viridis, abdominis apice cupreo, anteunis fulvis, articulo tertio apice nigro, pedibus fulvis, tarsis nigris, alis subcinereis. Body green ; head covered in front with tawny down ; eyes red ; mouth pitchy ; palpi tawny ; feelers tawny ; third joint oval, black towards the tip ; bristle black ; breast, sides of the chest and under side of the abdomen covered with white bloom ; abdomen coppery towards the tip ; legs tawny ; four hind hips green, covered with white bloom ; feet black ; wings slightly gray ; wing- ribs and poisers tawny ; veins black. Length of the body 2 lines ; of the wings 4 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 665. Dolicliopiis termiraatus, n. s., Fern. Aureo-viridis, capite cyaneo-viridi, abdornine cupreo basi viridi, antennis nigris, pedibus fulvis, tarsis piceis basi fulvis, alis sub- cinereis fusco subvittatis. Head bluish-green, covered in front with golden clown, fringed on each side with white hairs ; eyes bright red ; mouth pitchy ; palpi tawny ; feelers black ; third joint nearly oval ; chest golden- green ; sides and breast covered with hoary bloom, which also appears beneath the abdomen ; abdomen copper-color, gf een at the base ; legs tawny ; four hind hips green, covered with white bloom ; feet pitchy, tawny towards the base ; wings slightly gray, indistinctly tinged with brown along the borders of the veins ; wing-ribs and poisers tawny; veins pitchy. Length of the body 2 lines ; of the wings 4 lines. a. North America. Presented by the Entomological Club. APPENDIX. 317 Page 666. Dolictiopus sequax, n. s. Cyaneo-viridis, thorace aeneo-viridi, abdominis apice aeneo, an- tennis fulvis, articulo tertio nigro subtus fulvo, pedibus fulvis, tarsis nigris anticis piceis, alis linipidis. Body bluish-green ; head covered in front with golden down ; eyes red ; mouth pitchy ; feelers tawny ; third joint black, tawny beneath towards the base ; bristle black ; chest with a slight brassy tinge ; breast covered with a hoary bloom ; abdomen bluish-green ; tip bronzed ; appendages tawny ; scales white, with dark borders, under side slightly covered with hoary bloom ; legs tawny, feet black ; four hind hips mostly green, covered with white down ; middle shanks with a slight tuft of black hairs at the base ; fore feet pitchy, tawny at the base ; wings colorless ; wing- ribs and poisers tawny ; veins black. Length of the body 1| line ; of the wings 3 lines. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by Gr. Baruston, Esq. Page 666. Dolicliopus soccatus, BAENSTON'S MSS. JEneus, capite viridi, abdomine cupreo basi viridi, antennis fulvis, articulo tertio nigro subtus fulvo, pedibus fulvis, tarsis nigris, alis subcinereis. Head green, covered in front with hoary down, fringed along the eyes with whitish hairs ; eyes red ; mouth tawny ; feelers tawny ; third joint black, very short, tawny beneath till near the tip ; bristle black ; chest brassy ; sides and breast covered with hoary bloom ; abdomen coppery, green at the base ; legs tawny ; feet black ; wings slightly gray ; wing-ribs tawny ; veins black ; poisers yellow. Length of the body 1| line ; of the wings 3^- lines. Yar. J3. Body brassy, tinged with green. a. St. Martin's Falls, Albany River, Hudson's Bay. Presented by G. Barnston, Esq. Page 666. Dolicliopus remottis, n. s. ^Eneo-viridis, capite thoracisque lateribus cyaneo-viridibus, ab- domine basi viridi, antennis nigris, pedibus fulvis, tibiis posticis apice tarsisque posticis- nigris, alis limpidis. 318 DIPTERA OF NORTH AMERICA. [PART II. Head bluish-green, clothed in front with white down ; fringed on each ' side with white hairs ; eyes red ; feelers black ; chest brassy green, bluish-green on each side ; breast covered with hoary bloom ; abdomen brassy, green at the base, tinged with green on each side, covered with white bloom beneath ; tip black ; appendagefs tawny ; scales white, bordered with black ; legs tawny ; tips of feet pitchy ; hind feet and tips of hind shanks black ; wings colorless ; wing-ribs and poisers pale tawny ; veins black. Length of the body 1^ line ; of the wings 3 lines. a. North America. Presented by the Entomological Club. Page 667. Dolichopus irrasus, s. n., Fern. Cyaneus, abdomine aeneo, antennis nigris, pedibus fulvis, tarsi? piceis, alis cinereis fusco subvittatis. Body deep blue, beset with black hairs and bristles ; head covered in front with a silvery bloom ; eyes red ; mouth and feelers black ; -chest covered with ferruginous bloom ; sides and chest covered with white bloom ; abdomen dark bronze, slightly covered with white bloom, not longer than the chest ; legs tawny, clothed with black hairs and bristles ; feet pitchy ; wings gray, brownish along the borders of the veins ; wing-ribs and vein? black ; fourth longitudinal vein slightly bent ; poisers dark tawny. Length of the body 1 line ; of the wings 2 lines. a. Florida. Presented by E. Doubleday, Esq. Page 667. Ortliocfaile derempta, n. s. Yiridis, thoracis disco cupreo, abdomine purpureo, basi apiceque cyaneo-viridi, lateribus aureo-viridibus, autennis nigris, pedibus fulvis, femoribus viridibus, alis subcineris. Body green ; head and chest beset with black bristles ; eyes and mouth black ; feelers black ; third joint very short, round ; bristle proceeding from its tip ; disk of the chest copper-colored ; abdomen purple, clothed with black hairs, bluish-green at the base and at the tip, golden-green along each side ; legs tawny, clothed with short black hairs ; hips and thighs green ; wings slightly gray ; wing-ribs and poisers tawny ; veins pitchy. Length of the body 1^ line ; of the wings 3 lines. a North America. Presented by the Entomological Club. APPENDIX. 319 Walker, in the Transactions of the Entomological Society, Tom. IV. Page 149. Psilopus ungulivena. Mas. — Laste virfdis, antennis nigris thorace longioribus, thorace subcyanescente, abdomine subaurato, pedibus testaceis, alls sub- cinereis, venis nigris. Male. — Bright green; antennae black, much longer than the thorax ; thorax slightly bluish ; abdomen somewhat gilded ; legs testaceous, long, slender ; wings grayish ; veins black, fore-branch of the praebrachial vein very much bent, nearly rectangular ; discal transverse vein very deeply undulating. Length of the body 4^ lines ; of the wings T lines. United States. Walker, in the Transactions of the Entomological Society, Tom. V. Page 287. Psilopus solid us. Fcem. — Cyaneo-viridis, robustus, subtus albido-tomentosus, an- tennis pedibusque nigris, abdominis lateribus basi cupreis, alis subcinereis, fasciis duabus (la media lata, 2a apicali latissima) nigris antice connexis, halteribus testaceis. Female. — Bright bluish-green, stout, with whitish tomentum beneath ; antennae and legs black ; abdomen bright cupreous on each side at the base ; wings slightly grayish, with a broad black band in the middle and a very broad apical black band, the two bands connected in front ; fore branch of the prasbrachial vein almost rectangular ; discal transverse vein straight, oblique ; halteres dull testaceous. Length of the body 3 lines ; of the wings 7 lines. Mexico. Page 287. Psilopus peractus. Fcem. — Viridis, robustus, subtus • albido-tomentosus, capite cyaneo, antennis, pedibus halteribusque nigris, abdomine aeneo- viridi, alis subcinereis, venis nigris. Female. — Green, stout, with whitish tomentum beneath ; head blue; antennae and legs black; abdomen seneous-green ; wings 320 DIPTERA OF NORTH AMERICA. [PART II. grayish ; veins black ; fore-branch of the prsebrachial vein obtusely rectangular ; discal transverse vein oblique, almost straight ; hal- teres black. Length of the body 2^ lines ; of the wings 4 lines. Mexico. Page 287. Psilopus haeretious. Fcem. — Purpureo-niger, latus, nitens, subtus albido-tomentosus, capite, antennis pedibusque nigris, abdomine nigricante purpureo, alis subcinereis, venis nigris. Female. — Purplish-black, broad, shining, with whitish tomen- tum beneath ; head, antennae and legs black, the latter rather stout ; thorax rather thickly beset with black bristles ; abdomen blackish-purple ; wings slightly grayish ; veins black ; fore-branch of the pra3brachial vein rectangular, but with the angle somewhat rounded ; discal transverse vein oblique, nearly straight. Length of the body If line ; of the wings 3^ lines. Mexico. Page 288. Psilopus permodicus. Mas. — Aureo-viridis, gracillimus, antennis pedibusque flaves- cente albis, alis limpidis, venis halteribusque pallidis. Male. — Golden-green, very slender ; antennas and legs yellowish- white ; wings limpid ; veins pale ; fore-branch of the preebrachial vein obtusely rectangular ; discal transverse vein oblique, straight ; halteres very pale. Length of the body If line ; of the wings ^ lines. Mexico. SUPPLEMENT TO THE MONOGRAPH ON NORTH AMERICAN DOLICHOPODIM.1 I. General Remarks on the Dolichopodidse of North America. THAT the North American fauna of Dolichopodidse is an ex- ceedingly rich one, is proved by that portion of it upon which I have based the present publication. I am satisfied that this fauna far exceeds the European fauna in the variety of forms and in the number of species. I take the following points to be peculiar to this fauna : 1. The apparently rather numerous species of Pelastoneurus ; 2. The re- markable abundance of closely allied species of true Gymnopter- nus; 3. The number of species of Chrywtus distinguished by a variety of plastic characters, which is not generally the case in this genus ; 4. The abundance of species of Diaphorus and of forms related to this genus. Our knowledge of the genera occur- ring in North America is too limited, yet, to indicate the absence of some of them as being peculiar to the fauna. A very striking circumstance connected with the North Ameri- can fauna of Dolichopodidx is, that precisely in those points which we have just enumerated as peculiar to it, this fauna shows the most remarkable analogy to the remains of the fossil fauna of the same family preserved in amber. In both, there is the same abundance of species of genuine Gymnopternus, difficult to dis- 1 The volume had already gone through the press when the present sup- plement was sent in by Mr. Loew. It contains descriptions of the new species discovered mostly by me during the summer 1863. The (ii'nrral Remarks, prefixed to this Supplement, form au important addition to the preface of this volume (page iii — vi). 0. S. 21 ( 321 ) 3:22 DIPTERA OF NORTH AMERICA. [l-ART II. tinguisli on account of their close resemblance ; in both, the same frequence of species of Chrysotus, and not only the same varicu- of plastic specific characters among them, but even a most strik- ing conformity in the nature of these characters ; in both, nume- rous species of Diaphorus and of forms related to them. A cer- tain coincidence is even perceptible among those genera, which hitherto are not represented either in the North American or in the amber-fauna. It must be added, however, that the latter shows nothing like the great abundance of the North American fauna in species of genuine Dolichopus. It would be difficult at present to make any satisfactory state- ment as to the relation in which the North American fauna of Dolichopodidse stands to that of any other zoological province, as, with the exception of the European fauna, our knowledge of other faunas is not sufficient for this purpose. From what we know, however, we distinctly perceive that the North American fauna closely approaches the European and the North Asiatic faunae in the species of the genera Hygroceleuthus, Dolichopus, Tachytre- chus, Campsicnemus, Scellus, Hydrophorus, Liancalus, Chrysoti- mus, and Xanthochlorus, whereas its coalescence with the South American fauna is apparent in the species of Paraclius, Pelasto- neurus, Lyroneurus, and Plagioneurus. The species of genuine Gymnopternus, so numerous in North America, are but scantily represented in Europe. The North American species of Diapho- rus agree in part with the European, in part with the South American species. The North American species of Argyra, Por- phyrops, Leucostola, and Liancalus do not show any striking difference from the European species of these genera, but just as little from the South American species. Of such species, as are common to Europe and North America, the following have hitherto come under my observation : Doli- chopus brevipennis Meig., Dolichopus plumipes Scop., discifer Stann., Scellus spinimanus Zett., and Wied. The first four of these species belong altogether to specific types commonly represented on both continents ; but this is not the case with Psilopus pallens. This species unques- tionably belongs to the circle of European types of Pxilopus, whereas all the North American Psilopus at present known closely approach the types of their South American brethren. It seems, therefore, not altogether unnatural to suppose that this SUPPLEMENT. S23 species, which, according to Baron Osten Sacken's statement, is not uncommon in the lower parts of New York City, should have been accidentally imported in ships from the south of Europe. As species common to both continents may perhaps be also re- garded Diaphorus nigricans Meig. and Xanlhochlorus tenellus Wied. ; the North American Diaphorus opacus might be con- sidered as identical with the first, Xanthochlorus helvinus with the second of these species ; certainty about this point, however, can only be acquired by the close comparison of a larger number of well-preserved specimens of the two American species. II. Description of some Species communicated after the Volume had gone through the Press. Gen. II. DOLICHOPUS. Corrected Table for determining the Species. f Prevailing color of the feet black. 2 *. Prevailing color of the feet yellow. 9 „ ( Cilia of the inferior orbit black. 3 " \ I Cilia of the inferior orbit whitish. 4 ., f Face ochre-yellow. 1 gratus Lw. o s t Face silvery white. 2 laticornis Lw. f First joint of the hind tarsi with numerous bristles. 3 setifer Lw. I First joint of the hind tarsi with a few bristles. 5 f Hind tibiae black only at the tip. (j t Hind tibiae entirely black. 7 f The black color at the tip of the hind tibiae is rather extended ajid not very sharply limited. 4 albiciliatus Lw. 1 The black color at the tip of the hind tibiae but little extended and sharply limited. 5 xanthocnemus n. sp. ,. f A considerable extent of the tip of the femora yellow. 6 tetricus n. sp. I The extreme tip of the femora only somewhat yellowish. 8 „ f Lamellae of the hypopygium pointed. 7 acuminatus Lw. ( Lamellae of the hypopyginm rounded ovate. 8 ovatus Lw. Q f Cilia of the inferior orbit black. 10 ' I Cilia of the inferior orbit pale. 12 ,(. ( Fore coxae blackish. 9 pachycnemus Lu-. * Fore coxas yellow. ] 1 -., f The first two joints of the antennae yellow. 43 dorycerus /.,,-. I The whole antennae black. 10 brevipennis Men;. , „ f Tegulae with pale cilia. 13 I Tegulse with black cilia. 2G 324 DIPTERA OF NORTH AMERICA. [PART II. , o / Antennre black, at the utmost the first joint almost yellowish-red. 14 t Antennae, altogether or at least their larger portion, yellowish-red. 24 . f Fore coxfe dark at the base, beyond the middle. 11 longimanus Ln\ 14 < -.f (. Fore coxae pale. f Tip of the hind tibiae distinctly black. 1 Tip of the hind tibiae not, or very slightly infuscated. 17 , fi f Fore tarsi only ferruginous-brownish. 12 brevimanus Lw. I Fore tarsi blackened from the tip of the first joint. 13 socius Lw. .. „ f Hind tarsi entirely black. I Basis of the hind tarsi to a considerable extent pale. 22 1 „ f Hind femora of the male not ciliated. 14 nudus Lw. I Hind femora of the male ciliated. ("The enlarged last joint of the fore tarsi of the male on the outside with a white reflection. 45 palsestricus Lw. j The enlarged last joint of the fore tarsi of the male on the outside without a white reflection. 20 9n C'Hind femora of the male very densely ciliated. 16 splendidus Lw. (. Hind femora of the male sparsely ciliated. 21 /-Hind tibiae not infuscated at the tip ; the fourth joint of the fore tarsi of the male somewhat broader than the preceding. 44 splendidulus Lw. Hind tibiae somewhat infuscated at the tip; the 4th joint of the fore tarsi of the male not broader than the preceding. 15 subciliatus Lw. f Only the last joint of the fore tarsi of the male enlarged. 22 -^ 17 batillifer Lw. (. The two last joints of the fore tarsi of the male enlarged. 23 9q ( Hind femora of the male ciliated. 18 eudactylus Lw. I Hind femora of the male not ciliated. 19 tonsns Lw. 9 , f Last joint of the fore tarsi of the male enlarged. 20 tener Lw. I Fore tarsi of the male plain. 25 nr- f Wings hyaline with a grayish tinge. 21 variabilis Lw. \ Wings hyaline with a yellowish tinge. 22 luteipennis Lw. Fourth longitudinal vein broken. 27 Fourth longitudinal vein not broken. 31 9_ ( Antennas black. 23 ramifer Lw. \ Antennae yellowish red. 28 f Fourth longitudinal vein broken twice at right angles. 24 bifractus Lw. 28 •! | The lower angle of the fourth longitudinal vein sharp, the upper one rounded. 29 9q | Tarsi of the male plain. 25 vittatus Lw. \ Fore tarsi of the male enlarged at the tip. 30 21-! f 1 SUPPLEMENT. 325 OA ( Hind femora of the male ciliated. 26 cuprinus Wied. tj\J S I Hind femora of the male not ciliated. 27 longipennis Lw. (Antennae red, at the utmost the third joint at the tip, or its larger portion, blackened. 32 Antennae black, at the utmost the first joint, in part, red. 39 09 f Humeral callosity of the same color as the dorsum of the thorax. 33 "i ( Humeral callosity yellowish. 38 ! Arista of the antennae of the male very much enlarged at the tip. 28 hastatus Lw. Arista of the antennae of the male plain. 34 04 ( Last joint of the fore tarsi of the male not enlarged. 35 ' Last joint of the fore tarsi of the male enlarged. 36 f First joint of the middle tarsi of the male feathered. I 29 plumipes Scop. 1 First joint of the middle tarsi of the male not feathered. 30 fulvipes Lw. f Last joint of the fore tarsi of the male with a lamelliform appendage. 31 sexarticulatua Lw. i Last joint of the fore tarsi of the male without lamelliform appendage. [ 37 o- f Last joint of the fore tarsi of the male small. 32 ruficornis Lie. ( Last joint of the fore tarsi of the male large. 40 lobatus Lw. 00 f Fore tarsi of the male plain. 33 scapularis Lw. \ Fore tarsi of the male enlarged at the tip. 34 funditor Lw. on f Antennae entirely black. 40 ' First joint of the antennae partly red. 43 ,Q ( Hind femora not blackened at the tip. 41 I Hind femora blackened at the tip. 42 ^ f Hind tibiae not blackened at the tip. 35 chrysostomus Lw. ( Hind tibiae blackened at the tip. 46 melanocerus Lw. f Anterior femora without dark streaks on the under side. J 37 comatus Lw. 1 Anterior femora with dark streaks on the under side. L 36 praeustus Lw. 40 f First joint of the hind tarsi yellow, with the exception of the tip. 44 ( First joint of the hind tarsi entirely black. 45 Lamellae of the hypopygiuni ochreous-yellow, not double. 38 scoparius Lie. 44 -j Lamellae of the hypopygium ochreous yellow, double, that is, having an inner pair of flabs besides the outer ones. L 47 quadrilamellatus Lw. f Tips of the hind tibiae at the utmost somewhat blackened on the in- 45 \ side. 46 I Tips of the hind tibiae distinctly black. 47 326 DIPTERA OF NORTH AMERICA. [PART II. f The first joint of the antennse red on the under side only. 39 discifer Lw. 46 ^ The first joint of the antennae red, with the exception of its upper side. 40 lobatus Lw. f Hind tibise of the male with bristles of unusual length. 41 setosus Lw. j Hind tibise of the male with bristles of ordinary length. 42 incisuralis Lw. 43. D. dory cerus LOEW. $ . — yEneo-viridis, oculorum tegularumque ciliis nigris, primis duobus antennarum articulis, coxis anticis pedi- busque saturate flavis. $ . Seta antennarum lamellifera, tarsorum auticorum articulis ultimis quatuor dilatatis, atris. 9 Metallic green ; cilia of the posterior orbit and of the tegulse black ; the first two joints of the antennae, the fore coxse and the feet saturate-yellow. £ . Arista expanded into a lamella at the end, the last four joints of the fore tarsi enlarged, deep black. 9 Long. corp. 0.26. Long. al. 0.26. SYN. Dolichopus dorycerus LOEW, Berl. Ent, Zeitschr. VIII, 93, 85. Male. Bright, bronze-green, usually with extensive coppery- red reflections. Antenna? small ; the first and second joints, which are very much obliterated, of a saturate-yellow color ; the third joint, which is round, and the arista, black ; the latter bears at its tip an elliptical black lamella. The face more ochre-brown than ochre-yellow ; the cilia on the posterior orbit altogether black. Hypopygium black ; the lamellae of middling size, dingy white with a rather broad black margin, jagged on the edge and beset with black bristles, on the upper margin with black hairs. Fore coxse with black hairs. Feet saturate-yellow ; hind femora before the tip usually with two, sometimes with one bristle ; the hind tibiee have upon their hind side, before the middle, a small brown callus, and are blackened at the extreme tip on the inside. Fore tarsi about as long as the tibise ; the first joint slender, stalk- like, considerably longer than the following four joints together, dark yellow, blackened only at the extreme tip ; the following SUPPLEMENT. 327 four joints velvet-black, strongly compressed from the sides ; the three last ones are expanded on the upper side into long lobes, which are velvet-black on the third and fourth joints; on the fifth joint the lobe is black only at the base, otherwise whitish. Middle and hind tarsi, from the tip of the first joint, black. Wings grayish-hyaline with dark-brown veins, tinged with clay- ish-yellow in the costal, marginal, and subinarginal cells, the costa only slightly mcrassated at the tip of the first longitudinal vein ; the tip of the third longitudinal vein strongly deflected backwards ; the last segment of the fourth longitudinal vein not broken ; the posterior margin of the wing has a deep sinus before the unusually protruding anal angle ; the latter is again sinuated, so as to appear bilobed. Nab. Glen-House, White Mountains, New Hampshire, July 2, 1863. (Osten-Sacken.) 4-4. I>. spleitdidulus LOEW. £ . — Viridis, nitidus, coxis anticis pedibusque flavis, tibiis posticis totis concoloribus, antennis tarsisque posticis nigris, ciliis oculorum inferioribus tegularunique ciliis flavi- caiitibus, alarum vena longitudinal! quarta non fracta. £, . Tarsis anticis elongatis, articulo quarto prsecedentibus latiore, quinto conipresso atro, femoribus posticis minus coufertini flavo-ciliatis. 9 Green, shining, fore coxse and feet yellow ; the hind tibiae not blackened at the tip ; antennae and hind tarsi black ; cilia of the inferior orbit and of the tegulae yellowish. £ . Fore tarsi elongated, fourth joint broader than the preceding; the fifth joint laterally compressed, black ; hind femora ciliated with rather sparse yellowish hairs. 9 Long. corp. 0.22. Long. al. 0.22-0.23. SYN. Dolichopus splendidulns LOEW, Berl. Eut. Zeitschr. VIII, 91, 82. Male. Metallic green, bright, shining. Face rather bright- yellow. Antenna? altogether black ; the third joint short-ovate. Front shining green. Cilia of the inferior orbit pale-yellowish. Larnellte of the hypopygium broad, ovate, whitish ; on the upper and the apical margins with a very narrow blackish border ; apical margin jagged and beset with black bristles. The four hind coxa? are blackish, only at the extreme tip yellow. Fore coxae yellow, somewhat blackened only at the extreme basis, beset with short 328 DIPTERA OF NORTH AMERICA. [PART II. black little hairs almost upon the whole front side. Feet yellow. The hind femora before the tip with a bristle, upon the greater part of the under side sparsely ciliated with rather long yellowish hairs. Hind tibia; of ordinary strength, not infuscated at the tip, with a long glabrous streak upon the hind side. Fore tarsi abundantly one and a half the length of the tibias ; the first four joints yellow, on the inside with a somewhat whitish reflection; stalk-shaped from the first to the third joint ; the fourth joint late- ! rally compressed, somewhat broader than the preceding, especially towards the tip ; the first joint nearly as long as the three follow- ing together ; the fifth joint black, compressed, broad, especially towards the tip, beset on the upper side with closely appressed little hairs. Middle tarsi blackened from the tip of the first joint. Hind tarsi altogether black. Cilia of the tegulse whitish. Wings hyaline, somewhat grayish, of rather uniform breadth ; the costa at the tip of the first longitudinal vein with a weak and very short swelling ; the fourth longitudinal vein not broken. Hab. White Mountains, New Hampshire, July, 1863. (Osten- Sackcn.) Observation. — This species has an extraordinary resemblance with D. splendidus on one side, and with D. subciliatus on the other. It4differs from D. splendidus, with which it agrees more with regard to the structure of the fore tarsi, by the less densely ciliated hind femora, and by the smaller extent of the incrassa- tion of the costa. D. subciliatus has longer and more slender fore tarsi, the fourth joint of which is as slender as the preceding ; it has the hind tibiae infuscated at the tip ; the cilia of its hind femora are not only more scarce but also shorter, finally the swell- ing of the costa is more extended. Moreover, not only D. splendi- dus, but also D. subciliatus are considerably larger than D. splen- didulus. This character will enable us to distinguish the female of D. splendidulus from that of D. splendidus, as well as from the female of D. subciliatus ; the two latter, however, cannot be confounded on account of the different color of the hind tibite. 45. D. palaestrictis LOE\V. £ and 9- — ^Ineo-viridis, pedibus flavis, coxis anticis tibiisque posticis totis concoloribus, aiitennis tarsisque posticis nigris, ciliis oculormn inferioribus tegularumque ciliis flavi- cantibus, vena alarum longitudinal! quarta iion fractft. £ . Ultimo tarsoruui auticoruni articulo aduioduui dilatato, nigro, in latere SUPPLEMENT. ,'llM) externo albo-micante ; femorum posticorum ciliis flavicantibus, non confertis. 9 • Pedibus simplicibus. Metallic green ; feet yellow ; fore coxae and the whole hind tibife of the same color; antenme and hind tarsi black; cilia of the inferior orbit and of the tegulse yellowish; the fourth longitudinal vein not broken. % . The last joint of the tore tarsi very much enlarged, black, on the out- side with a white reflection ; the yellowish cilia of the hind femora sparse. 9 • Feet plain. Long. corp. 0.24. Long. al. 0.23. SYN. Dolichopus palaestricus LOEW, Berl. Entom. Zeitschr. VIII, 92, 84. Metallic green, bright. Face of the male narrow, more pale ochre-yellowish than golden-yellow ; the face of the female broader and paler. Antenna? entirely black ; the third joint of the male ovate, that of the female shorter. Front green, bright. Cilia of the inferior orbit yellowish. Fore coxae yellow, on the front side with a short black pubescence. The four posterior coxa3 yellow only at the extreme tip. Hind femora with a bristle before the tip. Fore tarsi of the female and middle tarsi in both sexes blackened from the tip of the first joint. Hind tarsi black, ex- cepting only the extreme basis, which is yellowish-brown. Cilia of the tegulse yellowish. Wings grayish hyaline ; fourth longi- tudinal vein not broken. Male. Lamellae of the hypopygium whitish, of moderate size and oval form ; on the upper and apical margin they have a nar- row black border, the latter is jagged and beset with black bristles. Hind femora sparsely ciliated with yellow hairs. Fore tarsi once and a half so long as the tibia? ; the first four joints yellow, with a white reflection on their sides, slender, stalk-like ; the first joint as long as the three following together, the second abundantly one and a half so long as the third ; the third some- what broader than the preceding, especially toward its tip ; the fourth considerably shorter and broader than the third ; the fifth joint laterally compressed, very much enlarged, black, with a silky reflection ; on the outside this reflection sometimes appears almost silvery. Hind tibiee somewhat thickened ; the two thirds of their hind side without any pubescence. The costa at the tip of the first longitudinal vein with a rather elongated swelling. Hab. New Hampshire. (Osten-Sacken.) 330 DIPTERA OF NORTH AMERICA. [PART II. Observation 1. — D. palsestricus is very much like D. batillifer. It differs from it in both sexes by the somewhat larger antennae, and principally by the hind tarsi, which are black as far as the extreme basis ; moreover the male has distinctly shorter fore tarsi and their first three joints are somewhat stouter ; the cilia of the hind femora are more scarce ; the hind tibiae are less thickened, and the glabrous spot on their hind side is longer. The female may be distinguished from the somewhat uncertain female of D. splendidus, and from the as yet unknown female of D. splendidu- lus by the pubescence on the sides of the abdomen, which is, to a greater extent, of a pale color ; from the female of D. nudus it differs by the under side of the first joint of the antennae not being red. Observation 2. — The discovery of the present species makes it necessary to mention, in the diagnosis of D. batillifer, the pale color of the first joint of the hind tarsi, and the very dense fringe of cilia on the hind femora of the male. 46. D. melanocei'lis LOEW. 'J, and 9. — yEneo-viridis, antennis nigris, inferioribus oculoruin ciliis flavicaiitibus, ciliis tegularum uigris, coxis anticis pedibusque flavis, tarsis anterioribus hide ab articuli primi apice, tibiarurn posticarum apice tarsisque posticis totis uigris. •J, . Facie subaurea, tarsis simplicibus, femoribus posticis flavo-ciliatis. 9 . Facie albicante, femoribus posticis non ciliatis. Metallic green ; antennae black ; cilia of the inferior orbit yellowish ; cilia of the tegulse black : fore coxje and feet yellow ; the four anterior tarsi, from the tip'of the first joint, the tip of the hind femora and the whole hind tarsi black. ^, . Face almost golden-yellow ; hind femora with yellowish cilia. $ . Face whitish ; hind femora not ciliated. Long. corp. 0.20. Long. al. 0.20. SYN. Dolickopus melanocerus LOEW, Berl. Ent. Zeitsch. VIII, 93, 86. Male. Metallic green, bright. Front bright green. Antemife entirely black, rather large ; the third joint elongated-ovate, rather of equal breadth. Face rather narrow, golden-yellowish, but not shining. Cilia of the inferior orbit yellowish. Lamelke of the hypopygium of medium size, ovate, whitish, with a narrow black border, jagged on the apical margin and beset with blnck bristles. Fore coxse yellow, somewhat blackened at the extreme basis, and clothed on the front side with a black pubescence. Feet yell^tv ; hind femora before the tip with a bristle, ciliated un SUPPLEMENT. 331 the under side with scattered yellowish hairs. Hind tilme at the tip, to a considerable extent, black; on the hind side with a gla- brous streak, which reaches from the basis up to the tip. Fore and middle tarsi blackened from the tip of the first joint ; hind tarsi altogether black. Cilia of the tegulee black. Wings with a rather dark-gray tinge, and with black veins ; the costa has, at the tip of the first longitudinal vein, a very short knot-like swelling ; the fourth longitudinal vein is not broken. Female. The plastic characters, which .distinguish the male are wanting here, otherwise it resembles the male very much. The antennas are considerably shorter and their last joint is much smaller. The face is very much broader, grayish-white, with but little admixture of yellowish. Hob. Canada. (Couper.) Observation. — The male cannot be mistaken for any other species. The female differs from that of D. comatus by its more considerable size, darker wings, and the absence of a dark tip on the hind femora. It cannot be mistaken for the as yet unknown female of D. chrysostomus, on account of the extended black color of the tip of its hind tibiaa. All the other species, with the females of which it could be confounded, have the antennas not entirely black. . D. qiiadrilamellatlis LOEW. $ and 9 . — Viridis, nitens, an- tennis nigris, margine infero articuli primi rufescente, facie alba, inferi- oribus oculorum ciliis albidis, ciliis tegularum nigris, coxis anticis pedibusque flavis, tarsis posterioribus inde ab articuli primi apice nigris, alarum vena longitudinal! quarta non fracta. £ . Duobus ultimis tarsorum anticorum articulis depressis, atris ; lamellis hypopygii ochraceis, bilobis. 9 . Tarsis anticis inde ab articuli primi apice nigris. Green, shining ; antennae black ; the inferior margin of the first joint red- dish ; face white ; the cilia of the inferior orbit whitish ; cilia of the tegulse black ; fore coxae and feet yellow, the four posterior tarsi from the tip of the first joint black ; the fourth longitudinal vein not broken. $ . The two last joints of the fore tarsi flattened, black ; lamellae of the hypopygium ochre-yellow, bilobed. 9 . Fore tarsi from the tip of the first joint black. Long. corp. 0.27. Long. al. 0.2(i. SYN. Dolichopus quadrilai . — ^Eneo-viridis, nitens, anteunis nigfis, pedibus flavis, coxis omnibus, praeter apiceni, lamellisque hypopygii uigris. Metallic green, shining ; antenna black ; feet yellow, all the coxse with the exception of the tip and the lamellae of the hypopygium black. Long. corp. 0.15—0.16. Long. al. 0.15—0.16. Metallic green, shining. Antennae entirely black ; the third joint short, with a not easily perceptible pubescence. Front covered with white dust. Face of the £ ochre-yellow, sometimes rather brownish-yellow, that of the 9 grayish-white ; cilia of the inferior orbit black. Upper side of the thorax covered with gray or brown-gray dust, which is distinctly visible, when the upper side of the thorax is looked at in an oblique direction. Fore coxae blackened as far as the middle or nearly as far as the tip ; the four hind? coxa? yellow only at the tip. Feet yellow ; tarsi from the tip of the first joint strongly colored with brown or blackened, the crescent-shaped lamella of the hypopygium black. Wings tinged with blackish-gray and with brownish-black veins. Hab. District of Columbia ; Kow York. (Osten-Sacken.) Observation. — The female has, in the plastic characters, much in common with the female of G. spectabilis, of which I have now three specimens ; but the latter is somewhat larger, and the third and fourth longitudinal veins of the wings seem to be somewluit closer to each other ; I cannot, therefore, believe that G. spectabi- lis is only a variety in color of G. chalcochrus, and hope that my "view will be sustained through the discovery of the still unknown male of G. spectabilis. 26. c- antenna £ , and d. wing $ . 2. Doliciiopus FCTJJDITOR Lw. £ . a. head £ , 6. head 9 i c- antenna £ , and c?. wing 9 • 3. Rliagoneurtis POLYCHROMES Lw. £ . a. head £ , 6. antenna £ , and c. wing 9 • 4. Gymnopternus LUNIFEK iw. £ . rt. head £ , and b. head 9 °f Gymnopternus crassicauda Lw. — c. an- tenna £ of Gr. lunifer. — f/. antenna £ of G. subulatus Lw. — e. wing 9 °f G. crassicauda. 5. Pelastoneurus VAGANS Lw. ^ . a. antenna £ , i. head ^ , c. head 9 > and rf. wing ^ . 6. Tacliytrechus VORAX Lw. ^ . a. head £ of T. vorax. — 6. head £ of T. moechus Lw. — c. antenna £ of T. vorax. — d. antenna £ of T. moechus. — e. wing 9 of T. vorax. 7. Paraclius ALBONOTATUS Lw. £ . a. head 9 > &• antenna 9 > au(i c- wing 9 of P. arcuatus Lw. 8. Ortliochile SOCCATA Lw. ^ . a. and 6. head £ , c. antenna £ , and d. wing £ of same. 9. Hercostomus UNICOLOR Lw. % . a. antenna £ , and b. wing of same. PLATE IV. 10. SyMstroma NODICORNIS Meig. £ . — a. and b. head 9 , c. head £ , d. antenna 9 > e- antenna $ , and /. wing 1 of the same. ( 341 ) 342 DIPTERA OF NORTH AMERICA. [PART II. 11. Hypoplayllus DISCIPES Ahr. £ . a. head 9 , b. antenna 9 , c. antenna £ , and d. wing £ of the same. 12. Haltericerus EUCERUS Lw. £ . a. head £ , 6. antenna 9 , c. antenna £> , and d. wing £ of the same. 13. Diostracus PKASINUS Lw. $ . a. antenna £ , 6. head £ , c. head 9 ) and d. wing £ of the same. 14. Anepsius FLAVIVENTKIS Meig. £ . a. head $ , 6. antenna $ , and c. wing $ of the same. 15. Argyra ALBICANS Lw. $ . a. head $ , 6. head 9 > c- antenna £ , d. antenna 9 , and e. wing £ of the same.1 16. Syntoriuou METATHESIS Lw. £ . a. head % , ft. head 9 > c- antenna £ from the outside, e?. antenna 9 from the inside, e. antenna $ inside, and /. wing $ of the same.2 17. Synartlirus PAOIARIS Lz«. $ . a. head 9 > and b. head $ of £. pallipes Fabr. — c. antenna $ , and f/. wing £ of 5. palmar is. IS. SysteilMS SCHOLTZII i?<;. ^ . a. head £ of 5. bipartitus Lw. — &. antenna ^ of 5. Scholtzii. — d. wing $ of 6. head £ , c. head 9 ) d- antenna 'J, , and e. antenna 9 of the same.—/, antenna 9 of & lugubre. — g. wing 9 of R. longicorne. 20. Xipiianclriiim QUADRIFILATUM Lw. -J, . a. head'J,, and b. head 9 of X. caliginosum Meig. — c. antenna ^ of X. quadrifilatum, — d. antenna £ , e. antenna 9 , and f, wing 9 of X. caliginosum. 21. Porpliyrops MELAMPUS Lw. ^. a. head ^ , 6. head 9 > c- antenna ^ , d. antenna 9 > and e. wing $ of the same. 1 The antennal arista of the £ was made by the engraver a little too long, that of the 9 a little too short. 2 Owing to a mistake, which was discovered too late, the posterior trans- verse vein is wanting in fig. /in some of the impressions of this plate. EXPLANATION OF THE PLATES. 343 22. Smiliotus MARITIME Hal. £ . a. antenna £, of *S. thinophiius Lw. — b. antenna £ , c. head $ , c/. head 9 , and e. wing £ of S. maritimse. 23. Aplirosylus RAPTOR Walk. £ . a. antenna $ , 6. and c. head $ , d. wing £ of the same. 24. Tliiiiophilus FLAVIPALPIS Zett. £ . a. antenna $, ?>. head £ , c. head 9> d. wing 9 of the same. 25. Peodes FORCIPATUS Lw. $ . a. head£, 6. head 9> c- antenna £, d. wing £ of the same. 26. NematoproctUS DISTENDENS Mfig. £ . a. head $ , b. head 9 > c- antenna £ , rf. wing $ of the same. 27. Leucostola CINGULATA iw. £ . a. head £ , b. antenna $ , and c. wing $ of the same. PLATE VI. 28. Eutarsus AULICOS Met p. £ . a. head £> , 6. antenna £ , and c. wing £ of the same. 29. Diapliorus SPECTABILIS Lw. % . a. head £, , 6. head 9 °f -O- sodalis Lw. — c. head ^ , and c/. an- tenna £ of ZX spectabilis. — e. wing 9 of Z). interruptus Lw. 30. Liyroiieurus C.ERULESCENS Lw. 9 • a. head 9 > b. the same from the side, c. antenna 9 , d. wing 9 of the same. 31. ClirysotllS OBLIQUUS Lw. £ • a. head 9> and 6- head £ of C. obliquus.—c. head $ of (7. vividus Lw. — rf. antenna ^ of C. vividus. — e. antenna £ of C. obliquus Lw.—f. antenna $ of C. cornutus Lw. — . ^ . a. head ^ , 6. head 9 , c. antenna 9 > ^- antenna $ of £. nodatus. — e. wing -J, of S. tertianus Lw. 35. L.iaiical«is GENUALIS Lw. ^ . «. antenna 'J, , 6. head 9 , <-'• head ^ , and d. wing 9 of the same. 36. Plagioneiirus TTNIVITTATDS Zw. 9. a. head 9 > &• antenna 9 > au(i c. wing 9 of the same. 344 DIPTERA OF NORTH AMERICA. [PART II. PLATE VII. 37. Scellus AVIDUS Lw. £ . a. head £ , b. antenna £ , and c. wing $ of the same. 38. Hydropltorus INNOTATUS Lw. £ . a. antenna ^ , b. head 9 » c- wing £ of the same. 39. Medeterus DIADEMA Linn. £ . a. and b. head £ , c. antenna £ , and d. wing 9 of the same. 40. AchalcUS FLAVICOLLIS Meig. 9 • a. and b. head 9 > c- wing 9 > an(i d. wing 9 °f the same. 41. Xantlioclllorus HELVINDS Lw. $ . a. head £ , 6. antenna $ , c. wing $ of X. ornatus Hal. 42. Clirysotimus PUSIO Lw. % . a. head 9 , b. antenna $ , c. wing £ of C. molliculus Fall. 43. Saucropus DIMIDIATUS Lw. ^ . a. antenna 9 from the outside, b. antenna £ from the inside, c. head 9 » d. head $ , and e. wing $ of the same. 44. Psilopus FILIPES Lw. $ . a. head £ of Ps. filipes. — 6. head $ of Ps. scobinator Lw. — c. head £ of Ps. pollens Wied. — cZ. antenna £ of Ps. scintillans Lw. — e. antenna £ of Ps. pilosus Lw. — -f. antenna £ of Ps- comatus Lw. — jr. antenna £ of an undescribed species from Ceylon, related to Ps. globulifer Wied. — h. wing $ of A. psittacinus Lw. — i. wing £ of Ps. scobinator. REMARKS ON THE GENERIC CHARACTERS, EXPLANATORY TO THE PLATES. ALTHOUGH the present publication on North American Dolichopodidas was based upon a considerable number of species, it can be safely assumed that these species do not represent all the Genera occurring in North America. I have, therefore, added the characters of even those genera of the family, representative species of which have not yet been found on that continent, and I hope that this addition will prove useful to those desirous of studying this family in detail. In order to facilitate the recog- nition of generic characters, five plates, drawn for this purpose, have been added to this volume. But as on these plates every genus is represented only by a single species, this might easily give rise to the mistake that specific marks belonging to that particular species are generic characters. In order to prevent this, I have deemed it advisable to append to the plates the following explanatory remarks on the generic characters, which should always be consulted in determining species. The figures of the antennae show that the first joint in No. 1 — 16 is dis- tinctly provided with bristles on the upper side ; in No. 17 — 44, on the con- trary, it is glabrous. This distinguishes the two principal divisions of the Dolichopodidse. Among the species belonging to the FIRST PRINCIPAL DIVI- SION the genera numbei'ed from 1 to 12' have a completely disengaged, elongated hypopygium ; those numbered 14 — 1(3 a small, rounded, more or less imbedded one ; in No. 15 the hypopygium is short and sessile, but not imbedded, so that this genus (Diostracus) forms a transition from one of these two subdivisions to the other, and may be included either in the first or in the second. I have given preference to the first arrangement, but have separated this genus from all the others of this subdivision on account of the very large size of the palpi of the male. The other genera of the First Sub-division stand much nearer to each other in their organization ; they may, however, be easily arranged into two groups according to the circumstance whether the first joint of the hind tarsi is provided with bristles on its upper side (No. 1 — 3), or is without such bristles (4 — 16). To the first of these groups belong : Hygroceleuthus (1), Dolichopus (2), and Rhagoueurus (3). The difference between 1 These numbers refer to the plates. ( 345) 346 DIPTERA OF NORTH AMERICA. [PART II. HygroceleuthuS and Dolichopus consists in the structure of the head, which is higher and narrower in Hygroceleuthus, especially in the males, and in the length of the face, which reaches farther down in Hygro- celeuthus aud gives to the head a different profile. The genus Rhago- neurus differs from Tachytrechus and Dolichopus by the feathered arista of the antennae and the different structure of the male abdomen, which is a little compressed laterally, as also by the fracture of the last segment of the fourth longitudinal vein peculiar to it, and in which it is not equalled by any of those species of Dolichopus which have the last segment of the fourth longitudinal vein also fractured ; in the structure of the head Rha- goneurus agrees with Dolichopus, in that of the hypopygiuin with Doli- chopus and Tachytrechus.' 1 Mr. Rondani has established the genus Rhageneura on those European species of Dolichopus, the fourth longitudinal vein of which is broken. This genus cannot be retained in this sense for several reasons. 1. Because these species do not differ at all from the other species of the genus in the other parts of their organization. 2. Because the fracture of the fourth longitudinal vein of these species is very different in kind as well as in degree. 3. Because this fracture in some species is sometimes present, sometimes not. I have already stated above, on p. 19, that Dol. ziczac Wied. requires the establishment of a new genus. At that time I knew only this one species of the newly proposed genus, and considered therefore its establishment as premature. Since then I have become acquainted with several species, so that I feel prepared now to introduce this new genus, and take occasion at the same time to furnish a description of the typical species which I have figured. I propose for this new genus the name of Rhageneura, introduced by Rondaui in a somewhat different sense, modifying it only in the more correct Rhagoneurus. The characters of this genus are : First joint of the antennae with bristles on the upper side ; the arista of the antennae feathered or hairy; first joint of the hind tarsi with a bristle ; fourth longitudinal vein twice broken at right angles, with a considerable stump of a vein at each angle of the fracture ; abdomen of the male a little compressed laterally ; hypopygium entirely disengaged, as in Dolichopus. The figured species is the following : — Rhagoneurus pohjchromus nov. sp. £ and J . — Viridis, thorace violaceo- et cupreo-variegato, fronte violacea, antennis runs, pedibiis flavis, alis cinereis, venis transversis non infuscatis. ^ . Facie ochracea, hypopygii margine supero et apice flavis, lamellis parvis, albicantibus, teuuissime nigro-marginatis. 9 • Facie albida. Green with violet and coppery spots on the thorax ; front violet ; an- tennas red ; feet yellow ; wings gray ; the transverse veins without a dark margin. 1 . Face ochraceous ; upper margin and tip of the hypopygium yellow, the small lamellae whitish, with a very narrow black margin. REMARKS ON THE GENERIC CHARACTERS. 347, The peculiar ornaments, which the figured males of Hygroceleuthus and Dolichopus possess, as well as the strong swelling of the costa before the tip of the first longitudinal vein in Hygroceleuthus are not generic, but merely specific characters which, moreover, do not belong to the females. To the second yroup belong the genera G-ymnopternus (4), Paraclius (7), Pelastoneurus (5), Tachytrecuus (G), Orthochile (S), Hercostomus (9), Sybistroma (10), Hypophyllus (11), and Haltericerus (12). The genus Gyninopternus, if understood in the limited sense adopted above, differs from all the other genera of this group by the parallelism of the third and 9 . Face whitish. Long. corp. 0.17. Long. al. 0.17. Shining metallic green. Front bright, violet-blue, antennse yellowish- red ; the pubescence on the upper side of the first joint rather short ; the third joint rounded-ovate, however but little rounded at the end ; shorter in the female than in the male, in both sexes blackened to a very small extent on the upper margin and at the extreme tip. The arista of the antennae in both sexes with a considerable feathery pubescence. Fnce of the male not very narrow, ochre-brownish ; the face of the female broad, whitish. Cilia of the inferior orbit yellowish. Upper side of the thorax with copper-colored spots and with violet-blue reflections ; the former are more striking in the male than in the female, while the latter are visible either on the hind part of the dorsum of the thorax only, or spread on its middle more towards the front. Scutellum violet-blue with blue-green margins. Abdomen metallic green, often coppery upon most of the upper part, black at the incisures, covered on the sides with a rather strikingly white dust ; the hypopygium rather small, its second segment yellow along the margin, which is turned towards the venter, and at the tip; its external lamella? scarcely of middling size, yellowish-white, with a very narrow black margin, on the edge of the margin a little jagged and beset with crooked black bristles. Coxae yellow ; the foremost are beset, besides the usual black bristles, with short and fine black hairs ; the outside of the middle coxae is almost entirely covered by a large gray-black spot. Feet yellowish, even plain in the male ; hind femora with a bristle before the tip ; fore tarsi brownish-yellow, about as long as the tibia? ; middle and hind tarsi infuscated from about the tip of the first joint, brownish-black towards the end ; the hind tibiae of the male without glabrous spot on the hind side ; the first joint of the hind tarsi in both sexes has only one or two strong bristles on the upper side. Tegulse with black cilia. Wings tinged with dark gray with brownish-black veins ; the last segment of the fourth longitudinal vein is interrupted twice at right angles, and has, at each interruption, a long stump of a vein, as the other species of this genus ; transverse veins without any trace of dark margin ; in the male the costa has a slight swelling immediately before the end of the first longitudinal vein. Hub. Ceylon. 348 DIPTERA OF NORTH AMERICA. [PART II. fourth longitudinal veins of the wings ; it stands in rather close relation to the genus Dolichopus, which belongs to the first group, difl'ers from it, however, not only by the want of bristles on the first joint of the hind tarsi, but also by the smaller size of its species, the shortness of the first joint of the antennae, and usually by the more distinct pubescence of the arista ; moreover the lamellae of the hypopygium are usually smaller and not jagged at the end. The genus Paraclius is distinguished by a pecti- nate or subpectinate arista of the antennae, by the face, narrowed below even in the female, and by the peculiar course of the fourth longitudinal vein, the end of which forms a curve with its concavity turned backwards. Closely related to Paraclius is Pelastoneurus; the distinction between both genera is easy, if attention is paid to the essentially different course of the fourth longitudinal vein, to the very much more broad face, which is also more convex upon its lower part, and to the not sessile but peduu- culated hypopygium of the species of Pelastoneurus, which have the feathered arista of the antennas in common with the species of Paraclius. The genus Tachytrechus is very easily recognized by the great length of the perpendicular diameter of the very hairy eyes, by the face, which is narrower towards the middle but broader towards the bottom, and reaches entirely as far as the inferior margin of the eyes ; the palpi are also com- paratively small in the female, and the fourth longitudinal vein has before the middle of its last segment a gentle flexure, from which it converges towards the third longitudinal vein ; the figures of the antennae of the male of T. moechus, which remind of the structure of the antennae of Haltericems, are an exception to the rule ; however a similar structure occurs by way of exception in the genus Dolichopus. The genus Ortho- chile is so much distinguished by the structure of the proboscis and of the palpi as also by the distance of the tips of the third and fourth longitudinal veins from the tip of the wing, that it cannot be mistaken for any other genus. The genus Hercostomus, to which is to be added a large portion of those species which in my previous works I have com- prised in the genus Gymnoptemus, embraces a variety of organizations and is evidently capable of a subdivision into several genera ; in its whole structure it shows the greatest resemblance to Gymnoptemus, but in all the species the third and fourth longitudinal veins, towards their ends, are rather strongly converging, while in the species of Gymnoptemus they are either entirely parallel, or show but a very slight trace of convergency ; the hypopygium has, in its structure, a great similarity with that of Doli- chopus, but in some species the internal appendages are elongated, brush- like and hairy, like those of Hypophyllus and Haltericerus, which, how- ever, is also the case with some few species of Gymnoptemus. The genus jaybistroma, which hitherto remained confined only to a few species, is nearest to the genus Hercostomus, differs, however, by the scutellum which is much clothed with hair, by the peculiar structure of the face of the female which is much protruding below, and by the peculiar structure of the arista of the male-. The species of Hypophyllus are easily distinguished REMARKS ON THE GENERIC CHARACTERS. 349 by the slender structure of the whole body, by the length of their slender feet, by their pedunculated, usually yellow hypopygiuui, which is pro- vided with narrow, linear external appendages and elongated, more or less penicillated, internal appendages, likewise by the always very pro- minent development of the first joint of the arista of the antennae, which is different, however, in different species ; the scutellum is glabrous and the lower part of the female face not protruding. The spocies of Haltericerus are more robust than the species of Hypophyllus ; the face of the male is very narrow, that of the female very broad ; the second joint of the antennae in both sexes, especially in the males, is rudi- mentary, and the arista very elongated in the latter and enlarged at its tip into a lamella ; the hypopygiurn, attached to a long peduncle, and its appendages, bear the greatest resemblance to those of the species of Hypophyllus. To the above-named genera of the first group is appended Diostracus (13), as an anomalous genus. The very broad face in both sexes, the palpi, much larger in the male than in the female, the very small third joint of the antennae, the incrassated fore femora, the rounded, but not imbedded hypopygiurn, with its very small appendages and the long nar- row wings, with the posterior transverse vein very close to the margin — all these characters render this genus very easy to recognize. To the Second Sub-division, which is distinguished by the small, more or less imbedded hypopygium, belong the following genera : Anepsius (14), Argyra (15), and Syntormon (16). In Anepsius the second joint of the antennas has the usual transverse form, and the arista is inserted very close to its basis ; the third joint of the antennae shows a distinct pubes- cence ; the first longitudinal veiu is short, the fourth parallel with the third ; the abdomen of the male is laterally compressed. The relation- ship between Anepsius and the genus Synipycnus, which belongs to the second principal division, cannot be mistaken, though the distinct pubes- cence of the first joint of the antennae distinguishes it very easily from the other. The genus Argyra is distinguished by the transverse form of the second and the rather considerable size of the third joint of the antennae, its subapical arista, the broad wings — particularly towards the basis, the length of the first longitudinal vein, its distance from the costa and the inflection of the fourth longitudinal vein ; to these characters may be added, in most of the species, the delicate, but striking silvery white tomentum, spread over a large part of the body. Argyra is closely related to Leucostola, the first joint of the antennae of which, however, is glabrous on the upper side. The genus Syntormon differs from all the other genera of the whole first principal division by the form of the second joint of the antennae, which, on its inner side, overlaps the third joint in the shape of a thumb ; the face of the male is narrow, that of the female broad and protruding below like a roof: the arista of the antennas is com- pletely or almost completely apical; the third longitudinal vein is parallel or almost so. 350 DIPTERA OF NORTH AMERICA. [PART II. The SECOND PRINCIPAL DIVISION of the Dolichopodidse is divided, according to the shape of the third joint of the antennae, into two suit-divisions; the genera of the first sub-division (No. 17-23) have this joint, either in both sexes or at least in the male, pointed and provided with an apical arista ; in the genera of the second sub-division (No. 24-44), it is short in both sexes, and if it should be somewhat elongated in the male, it is not pointed, and the arista is either dorsal, or, at the utmost, sub- apical. The genera of the First Sub-division are divided into tioo groups. To the first group belong those genera, the posterior transverse vein of which is distant from the margin of the wing. They are the following : Synarthrus (17), Systenus (18), Rhaphium (19), Xiphandrium (20), Porphyrops (21), and Siniliotus (23). Synarthrus shares with Syntormon not only the peculiar structure of the second joint of the antennae, but also the remain- ing characters, with the single exception of the first joint of the antennae, which is glabrous. Systenus strikingly differs from all the other genera of the group by the pointed abdomen and the very pedunculated hypo- pyeium of the male ; to its distinguishing characters may also be reckoned the distinct pubescence of the third joint of the antennae, and the remark- ably sharp dividing line between the lower part of the face and the upper one. Rhaphium stands in close relation to the two following genera, is however distinguished from them by the very elongated antennae, even in females, and by the still more considerable stoutness of the- female pro- boscis ; moreover from Xiphandrinin it is distinguished by a less slender structure of the body, a much stronger pubescence, and a stronger flexure of the fourth longitudinal vein. The species of Xiphandrium have, like the species of Porphyrops, only in the males an elongated third joint of the antenna, in the females it is short ; their difference consists in a greater sleuderness, less pubescence, and less flexure of the fourth longitudinal vein ; moreover the largest species of Xiphandrium are scarcely equal in the length of their body to the smallest species of Porphyrops. The dis- tinguishing characters of the genus Porphyrops become manifest from what has just been stated about the two preceding genera. The genus Smiliotus has the antennae of equal size and form in both sexes ; they have on the under side, from the tip almost to the base, a distinct ex- cision ; the face of the male is broad, and its palpi are as large as those of the female ; finally, the abdomen shows externally one segment less than the related genera ; the feet are comparatively short and rather vigor- ous ; the small hypopygium is deeply imbedded and its appendages are often difficult to perceive. To the second group, which is characterized by a close proximity of the transverse vein to the posterior margin of the wing, belongs the only genus Aphrosylus (23) ; the third joint of its antenna? is but little elongated, but pointed ; the pendant palpi are larger in the male than in the female ; the proboscis is bent towards the chest, the face is narrower above, and REMARKS ON THE GENERIC CHARACTERS. 351 the eyes are excised near the antennae ; the feet have rough bristles, and the first joint of all the tarsi is much longer than the second. Among all genera of the Second Sub-division Psiloptis is distin- guished by the slender structure of its body, and especially of its feet, its very broad and more or less excavated front, the smallness of its antennae, and finally by its fourth longitudinal vein, which is provided with a pos- terior branch. I have assigned to it a position altogether at the end of the second sub-division, and will revert to it there. The other genera of this sub-division may be distributed into two groups according to the struc- ture of the thorax. To the first group belong those genera, the upper side 1 of the thorax of which is convex, as far as the scutellum (No. 24-39) ; to the second those, where the upper side of the thorax, before the scutellum, bears an inclined, more or less concave, area (No. 40-43). The genera belonging to the first group, and possessing a sixth longi- tudinal vein, form the first sub-group ; those where this vein is wanting, form the second sub-group. The first sub-group contains either such genera as have the transverse vein not close to the posterior margin of the wing, or such where an approximation of that kind takes place in a striking manner. The genera, where there is no striking approximation of the posterior transverse vein to the margin of the wing, are: Thiuophilus (24), Peodes (25), Nemato- proctus (2(5), Leucostola (27), Eutarsus (28), Diaphorus (29), Lyroneurus (30),Chrysotus (31), Teuchophorus (32), Sympycnus (34), Campsicuemus (33), and Plagioueurus (3ti). Thinophilus has the small and but little imbedded hypopygium in common with Peodes (in all the other genera that belong here, it is much more imbedded) ; it differs from Peodes by the structure of the face, which, in both sexes, is broader below, and has an angular margin ; by the large palpi of both sexes, and the very small size of the interior appendages of the hypopygium ; the first two joints of the antennae are small, the third somewhat in the shape of a lens, and the arista dorsal ; the last segment of the fourth longitudinal vein, towards its end, is parallel, or almost so, to the third longitudinal vein ; the feet of the male are usually variously adorned. The structure of the body of Peodes resembles in general that of Thinophilus ; the structure of the antennae and of the wings is also nearly the same, only the parallelism of the third and fourth longitudinal veins is still more complete than in most of the species of Thinophilus ; both of the interior appendages of the hypopygium form a large horny forceps ; the tarsi of the male of the only species hitherto known are plain. The species of Nematoproctus, in the whole structure of the body, are nearest the species of Porphyrops ; they can be, however, easily distinguished from them by the rounded, and in both sexes small, third joint of the antennae, and by the arista, which is inserted in the neighborhood of the basis. They are less closely related to the genus Leucostola, because the elongation of the first longitudinal vein, the broad shape of the wings, and the subapical position of the arista are wanting in them ; the same characters and the glabrousness of the 352 DIPTERA OF NORTH AMERICA. [PART II. first joint of their antennae distinguish them from the species of Argyra. The genus Leucostola stands to Argyra in the same relation as Synar- thrus does to Syntormon, that is to say, it differs from it only by the first joint of the antennae being without hairs ; the considerable size of the third joint of the antennae, the subapical position of the arista, the great breadth of the antennas towards the basis, the considerable length of the first longitudinal vein and its great distance from the costa, are characters shared by Argyra ; most of the species of Leucostola ha^e also, like the species of Argyra, a delicate, almost silvery white tomentum, which covers a considerable portion of the body. Eutarsus is very near to Diaphorus ; the difference is, that the former has the third joint of the antenna? some- what larger and that the transverse vein is a little more removed from the posterior margin of the wing ; moreover the face of the males of Eutarsus is narrower, the bristles of the imbedded hypopygium are less striking, the first joint of the hind tarsi is shorter, and the pulvilli of the fore tarsi are, although also enlarged, not so elongated as in Diaphorus. The genus Diaphorus is represented in North America by particularly varied forms ; its distinction from Eutarsus has been explained above ; from Lyroneurus it differs by the altogether different neuratiou of the wings, from Chrysotus by the greater slenderness of the whole body, and especially of the feet, and the comparatively longer and generally also proportionally narrower •wings ; moreover the structure of the head is different, as, while the eyes of the males of Diaphorus are either contiguous above the antennae or at least are separated by an equally broad front, those of the males of Chrysotus are sometimes contiguous below, but never above the antennae, and the front is always considerably widened above ; this difference in the structure of the front exists also in the females of both genera; finally, tlie males of Diaphorus distinguish themselves by the remarkable prolongation of the fore tarsi or of the fore and hind tarsi, and by the particularly striking manner in which the small imbedded hypopygium is provided with bristles, which characters are not found in the species of Chrysotus ; the neuration varies in the different species of Diaphorus in consequence of several modifications in the position of the posterior transverse vein ; in no species, however, is this vein approximated to the posterior margin of the wing ; the end of the last segment of the fourth longitudinal vein is sometimes cut off from the preceding part of it and pushed nearer to the anterior margin of the wing, so that both parts are either completely sepa- rated from each other or at least connected by an indistinct rudiment of a vein, as the illustration of the wing of Diapli. interruptus shows it. The genus Lyroneurus has a superficial resemblance to Diaphorus in the structure of the body, differs, however, strikingly by the large wings, with a more or less apparent greasy lustre, by the costa, which is usually thickened in both sexes, or at least in the male, to a very great extent ; by the end of the third longitudinal vein being strongly turned backwards, and by the wide space between the fourth and the third longitudinal veins ; the hypopygium is usually less provided with bristles than in the species REMARKS ON THE GENERIC CHARACTERS. 353 of Diaphorus ; the males of Lyroneurus, known to me, show no elongation of the pulvilli of the fore tarsi. The genus Chrysotus contains only small, mostly bright-green species, which cannot be mistaken for any other of the following genera, and which are sufficiently distinguished from Dia- phorus by the already mentioned characters; the statements made above in regard to the shape of the front, the absence of bristles upon the hypo- pygium, and the absence of the elongation of the pulvilli on the fore tarsi of the male, will help to recognize whether a given species, unless its female alone is known, is to be reckoned to Chrysotus or to Diaphorus ; as to the female, the structure of the front, the form of the wings, and the greater or smaller slenderness of the feet, have to guide us in its location. Striking is the very different form of the third joint of the antennae of the different species of Chrysotus in North America. The species of Teucho- phorus resemble more or less the smallest species of Chrysotus, are how- ever very easily distinguished in the male sex by the somewhat laterally compressed abdomen, by the feet which are beset with single, scattered, strong bristles, and by the crooked and variously adorned hind tibiae ; moreover, in all the known males of Teuchophorus there is a large black swelling on the costa before the tip of the first longitudinal vein, which is altogether an exception in the genus Chrysotus. More attention is neces- sary in order to recognize the females of Teuchophorus ; the best guide in this case is the steep position of the posterior transverse vein, the anterior end of which is nearer to the basis of the wing than the posterior ond, and the comparatively small antennae, the arista of which is somewhat less approximated to the apex than in most of the species of Chrysotus. The genus Sympycnus contains only small, mostly but little shining species, which are characterized by the smallness and the distinct pubescence of the third joint of the antennae, the insertion of the arista in the vicinity of its basis, the laterally compressed abdomen of the males, the not elongated metathorax, and the wings being more or less narrowed towards the basis ; they have the greatest analogy to the species of Anepsius, the first joint of the antennae of which, however, is clothed with hair; from Campsicnemus they are distinguished by the not elongated metathorax, by the abdomen, which is longer and not flattened from above, and by the face, which is narrowed towards the bottom ; the end of the fourth longi- tudinal vein converges somewhat, in the European species, towards the third longitudinal vein ; in the North American species it is parallel with it or almost so. (The want of the small transverse vein, and of the sixth longitudinal vein, which occurs in a number of the impressions of Tal>. VI, are merely mistakes of the engraver, which were discovered too late for correction.) Campsicnemus is distinguished by its face attenuated upwards, the distinctly elongated metathorax, and the short and, in both sexes, much flattened abdomen ; the third joint of the antenna) is small, or rather small, and in most of the species somewhat pointed ; the dorsal arista is inserted in rather close proximity to the basis ; the fourth longi- tudinal vein is always parallel to the third, and runs before its middle over 23 354 DIPTERA OF NORTH AMERICA. [PART II. a distinct convexity of the wing. The males are usually distinguished by striking ornaments on their feet, especially on the middle feet. The species of Flagioneurus are altogether distinct on account of the very diagonal position of the posterior transverse vein ; otherwise their neuratiou is very near to that of Pelastoneurus, while the glabrousness of the first joint of the antennae renders it utterly impossible to mistake them for any specie's of the latter genus. To the second sub-group, embracing those genera the transverse vein of which is strikingly approximated to the posterior margin of the wing, belong the following genera: Liaucalus (35), Scellus (37), and Hydro- phorus (38). Liancalus is easily distinguished from Scellus and Hydro- phorus by all the femora being slender and unarmed. In the species of Scellus the fore-femora are beset on the under side with rather strong bristles, catching into a row of similar bristles of the fore tibiae, and the third and fourth longitudinal veins are rather strongly convergent toward their ends, while the species of Hydrophorus have only on the under side of the fore-femora, towards the basis, a few elongated thorn-like bristles ; the under side, however, not only of the fore-femora, but also of the fore tibiae, is beset only with short bristles, and the third and fourth longitudinal veins are parallel towards their ends, or almost so. To the next following genus, Achalcus, I have assigned this position on account of the general structure of its body, which led me to suppose the existence of a relationship between it and the following genera ; I also presume, therefore, that the structure of the dorsum of the thorax, which I have not been able to ascertain positively in any of the few specimens belonging to me, will, when found out, justify the location I have assigned to the genus. In general the species of Achalcus may be easily dis- tinguished, as the total absence of the sixth longitudinal vein is a cha- racter peculiar to them ; the species hitherto made known are of a nou- metallic color, and of a very small size. The genera belonging to the second group, the thorax of which has on its posterior part a more or less concave, sloping surface, are the following : Medeterus (39), Chrysotimus (41), Xanthochlorus (42), and Saucropus (43). Medeterus differs essentially from the other three genera by the apical position of the arista, the very large size of the proboscis, and the totally disengaged hypopygiurn. The genera Chrysotimus and Xantho- chlorus contain only small species, and are easily distinguished by their coloring, which is either entirely yellow, or in part yellow, in part metallic- green ; Chrysotimus is easily distinguished from Xanthochlorus, by the subapical position of the arista and by the much smaller and imbedded hypopygium ; with Xanthoehlorus the position of the arista of the antenna is dorsal, and the swollen and rather disengaged hypopygium is directed backward so as to assume the appearance of a prolongation of the abdomen of the male. The species of Saucropus are of a more considerable size than the species of Chrysotimus and Xanthochlorus, and their feet are comparatively longer ; the second joint of the antennae somewhat differs REMARKS ON THE GENERIC CHARACTERS. 355 from the usual transverse form, as, on the inside, it reaches a little over the third joint ; the arista is dorsal ; the sixth longitudinal vein runs as far as the margin of the wing ; the course of the last segment of the fourth longitudinal vein varies in the different species, nevertheless it always shows a distinct convergency towards the third longitudinal vein ; the first joint of the hind tarsi is always considerably shorter than the second. The color of all the species is either entirely yellow, or reddish-yellow, or partly so. The genus Psilopus (44), closing the series, has the last segment of the fourth longitudinal vein provided with a posterior branch ; this cha- racter is not wanting in any of the American, European, or African species that are known to me ; in some South Asiatic species, however, it is repre- sented only by a fold of the wing, which sometimes is very indistinct. Besides, the species of Psilopus are very easily recognized by the slender- ness of the whole body and especially of the feet, by the very broad, more or less excavated front, by the small antennae, provided with a long, thin, dorsal or subapical, rarely almost entirely apical arista, and by the entirely disengaged hypopjginm. In the living state they keep the wings divari- cated, which gives them an entirely different aspect from all other genera. The males have their feet variously adorned, and in some species they are distinguished by a structure of the wings peculiar to them. INDEX. Achalcus, 217 Agonosonia, 230 Anepsius, 123 Aphrosylus, 148 Argyra, 123 Argyra albicaiis Lw., 125 albiventris Lw., 128 calceata Lw., 131 calci trans Lw., 130 cylindrica Lw., 132 diaphana Fbr., 126 minuta Lw., 129 nigripes Lw., 127 Campsicnemus, 193 Cainpsicnemus claudicans Lw., 194 hirtipes Lw., 193 Chrysotimus, 220 Chrysotimus delicatus Lw., 222 pusio Liu., 221 Chrysotus, 171 Chrysotus abdominalis Say, 172, 291 affinis Lw., 178 auratus Lw., 183 concinnarius Say, 172, 291 cornutus Lw., 174 costalis Lw., 179 discolor Lw., 182 incerus Wlk., 172, 308 longimanus Lw., 175 nubilus Stny, 172, 291 obliquus Liu., 176 pallipes Lio., 183 picticornis L?«., 184 subcostatus Lw., 181 validus £w., 175 viridifemora Mucq., 172, 298 vividus Lw., 178 Condylostylus, 230 Dasypsilopsus, 230 Diaphorus, 156 Diaphorus interrnptus Lw., 168 laniellatus Lw., 165 leucostomus Lw., 166 mundus Lw., 161 opacus T^y., 160 sodalis Zrzu., 163 spectabilis iw., 162 Diostracus, 120 Diostracus prasinus Lw., 121 Dolichopus, 18, 323 Dolicliopus abdominalis Say, 20, 292 acuminatus Lw., 34 adjacens Wll:, 22, 312 adjustns Wied., 16 affinis Wife., 21, 310 albiciliatus Lw., 31 batillifer ZM;., 45 bifractus Lw., 53 bifrons IF//'., 20, 301 brevimanus Lw., 39 brevipennis Meiy., 37 curysostonius Zw\, 67 ciliatus IF//,-., 31, 312 coercens IF//,-., 22, 312 comatus Lw., 69 confinis IF//-., 23, 315 con f us us Ztt., 71 consors Wlk., 21, 302 conterminus JF//\, 23, 31 5 contingens Wlk., 21, 302 contiguus Wlk., 23, 314 cupreus Say, 56 cuprinus IF/er/., 55, 311 discessus Wlk., 23, 314 disci fer Stain., 71 distractus IF//t., 23, 313 dorycerus Lw., 326 eudactylus Lw., 46 ( 357 ) 358 INDEX. Dolichopus exclusns Wlk., 23, 314 femoratus Sat/, 290 finitus Wlk., 22, 313 fulvipes Lw., 61 funditor Lw., 66 gratus Lw., 29 groenlandicus Ztt., 20 hastatus Lw., 59 hebes Wlk., 21, 302 heteroneurus Macq., 20, 299 incisuvalis Lw., 74 ineptus JF//L, 21, 303 irrasus Wlk., 24, 318 laniellipes IF/A:., 21, 311 Laticornis Lw., 29 lobatus Lw., 72 longimanus Lw., 38 longipennis Lw., 57 luteipeniiis Lu;., 51 maculipes Wlk., 21, 303 melanocerus Lw., 330 nudus Lw. , 41 obscurus Say, 20, 290, 296 ovatus Lw., 35 pachycnemus, Lw., 35 palsestricus Lit?., 328 patellatus Meiy., 71 pennitarsis Fall., 60 pluniipes Scop., 60 pluiuitarsis Fa//., 37 prseustus Lw., 68 pulcher Wlk., 21, 304 quadrilamellatus, Lw., 331 ramifer Lw., 52 remotus Wlk., 24, 317 ruficornis Lw. , 63 scapula.ris Lw., 64 scoparins Lw., 70 separatus Wlk., 24, 316 sequax VF7J-., 24, 317 setifer Lw., 30 setosns Lw., 73 sexarticulatus L?r., 62 soccatua Wlk., 24, 317 socius Lw., 40 splendidus Lw., 44 splenclidulus Lw., 327 subciliatus Lw., 42 tanypus Zw., 71 tener Lw., 49 terminatus PF/^.,24, 316 tetricus Lw., 33 to7isus Lw., 47 unifasciatus •?«(/. 289 variabilis Lw., 50 Dolichopiis varius IF//,-., 21,304 vittatus iw., 55 xanthocnemus Lw., 31 ziczac IFi'ec?., 16 Eurostomems, 230 Eutarsus, 153 Eutarsus eques Lw., 154 Gyrnnopternus, 75, 332 Grymnoptemua al biceps Lw., 85 barbatulus, Lw., 82 chalcochrus £?/' ,6tffo**t- PI. 17 f A if for J M Z*> jT HI*'- '. i A