Uwiv.oF TORONTO LIBRARY Digitized by the Internet Archive in 2010 with funding from University of Toronto http://www.archive.org/details/mossesofalaska00Ocard be) Wil i Wis ve Ea\b), 7 \ n A ae Pr | a a i) ul ne ay i ie a he } a ie : WAT in . i} Bh) , ‘ i i) a "ih. un aa yon THE MOSSES OF ALASKA The following paper on the Mosses of Alaska, by J. Cardot and I. Thériot, was originally published in the Proceedings of the Washing- ton Academy of Sciences, vol. Iv, pp. 293-372, July 31, 1902. It is here reprinted from the same electrotype plates, so that it may be quoted exactly as if it were the original. The original pagination has been preserved and transferred to the inner or hinge side of the page, where it is enclosed in brackets, thus [294]; while the consecutive pagination of the present volume has been added in the usual place. In the plates the original numbers and running headline, slightly abbreviated, have been preserved [in brackets], while the volume designation and serial plate numbers have been added in the usual place. The original text references to the plates are unchanged. The present headpiece and title have been substituted for the running head- ing of the Academy’s Proceedings and the original title, which was: Papers from the Harriman Alaska Expedition. xxix. The Mosses of Alaska. No other alterations have been made. The authors desire to record the following corrections : Page 255 [295]: Second line from bottom, for Jolycarfam read polycarpum. Page 278 [318]: Eleventh line from bottom, omit ‘ pl. vit, fig. 1.’ Eighth line from bottom, omit ‘ pl. v1, fig. 2.” Fourth line from bottom, omit ‘ pl. v, fig. 3.’ Page 288 [328], seventh line from bottom, for ‘ Barclay’ read Boulay. Page 303 [343]: Omit line of synonymy under ‘ Hypnum sarmentosum beringianum.’ Seventh line from bottom, omit ‘in litt.’ Omit line of synonymy under section ‘Calliergidium.’ The following species should be added to those contained in the paper: Edipodium grifithsianum Schwgr., a species not heretofore recorded as American, is added by Mrs. E. G. Britton, from Kadiak (Trelease, 1416). EpItTor. (252) THE MOSSES OF ALASKA BY J. CARDOT AND I. THERIOT INTRODUCTION Tue following catalogue of the mosses of Alaska and some adjacent islands is based primarily upon the collection made by the Harriman Expedition in 1899. For completeness, how- ever, species previously reported from the region are also included. The whole number here enumerated is 280, of which 124 are new to Alaska and 46 are new to science. The 29 new species and 17 new varieties, except for three species of Bryum, are here first described. The mosses brought back by the Harriman Expedition were collected’ by Wm. H:. Brewer, W- R: Coe, LJ. Cole; F. V. Coville, T. H. Kearney, De Alton Saunders and Wm. Trelease. Previous collections were made by W. H. Dall, 1867 ; Krause brothers, 1882; W. G. Wright, 1891; Jas. M. Macoun, 1891- 2; B. W. Evermann, 1892; C. H. Townsend, 1893-95 (Exp. of U.S. S. Albatross); W. M. Canby, 1897; W. H. Evans, 1897; W. A. Setchell, 1899; F. C. Schrader, 1899. [293] _ (353) 254 CARDOT AND THERIOT . [294] Subclass ANDREZALES. Family ANDREZACEZ. Andreea petrophila Ehrh. in Hann. Mag., 1784, p. 140, and in Beitr. Isp p aLO2s From Orca (Trelease, 2245), Hall Island (Trelease, 2127), St. Matthew Island (Trelease, 2168, 2530). New to Alaska. Andreza petrophila sylvicola Bryol. eur., vi, p. 13, pl. 2, e. From Hall Island (Trelease, 2527). New to Alaska. Andreea parvifolia C. Miill. in Flora, 1887, p. 219. From upper part of Dyea valley (Krause brothers). Andreza papillosa Lindb. in Oefv. af Vet. Ak. Forh., xxm, p. 557. From St. Lawrence Bay, and Plover Bay, Siberia (W. H. Dall). Subclass BRYALES. Family WEISIACEZ. Gymnostomum curvirostre scabrum Lindb. Muse. scand., p. 22. From Port Wells (Trelease, 1834). New to Alaska. Anectangium compactum Schw. Suppl. I, 1, p. 36, pl. x1. From White Pass, 1,900 feet (Trelease, 2309); Orca (Trelease, 2259, 2260 in part). Anectangium compactum alaskanum yar. noy. Habitu robustiore, foliis madore magis patulis, longioribus latiori- busque, cellulis majoribus (mediis 6-9 » latis, loco 4-6) reteque magis opaco distinctum. From Port Wells (Trelease, 1832). Dicranoweisia crispula Lindb. in Oefv. af Vet. Ak. Forh., 1864, p. 230. From White Pass, 3,000 feet (Trelease, 2492); Port Clarence (Trelease, 2119); Hall Island (Trelease, 2129, 2131, 2134)}3 St. Matthew Island (Trelease, 2153, 2154) ; Attu Island (J. M. Macoun). Most of these specimens have the inner pericheztial bracts shortly acuminate, which relates them to D. contermina Ren. & Card. (D. roelit¢ Kindb.), but the alar cells are usually more distinct than on the moss from Oregon and Idaho. Moreover, the comparison with numer- ous specimens from different regions of Europe and North America proves that the characters on which D. conterména has been established [295] THE MOSSES OF ALASKA 255 are too variable and insufficient to establish a specific distinction. D. contermina must therefore be considered as only a variety of D. crispula, and the specimens from Alaska and the islands of Bering Sea are nearly all intermediate between the type and this variety. D. obliqua WKindb., which has been recorded from Alaska, is un- known to us; but, from the description, it is probable that it, like- wise, is only a form of D. criéspula. (Cfr. Kindberg, Notes on Canadian Bryology, 1893, and Eur. and N. Amer. Bryinee, p. 210.) Rhabdoweisia fugax Br. eur., fasc. 33-36, p. 4, pl. 41. From Kodiak (Trelease, 2217, 2218). New to Alaska. Rhabdoweisia fugax subdenticulata Boul. Musc. dela France, p. 543. From Juneau (Brewer and Coe, 6992). New to Alaska. Another species of Weisiacee, Oreowersta serrulata Sch., has been recorded from Nulato by J. T. Rothrock and by Lesquereux and James (Manual, p. 58). Family DICRANACEZ. Cynodontium torquescens Limpr. Laubm., 1, p. 288. From Port Clarence (Trelease, 2101, 2102, 2525). Number 2101 has the peristome smooth or nearly so; it is C. swd- alpestre Kindb. in Mac. Cat. Can. pl., v1, Musci, pp. 17 and 257. Cynodontium treleasei sp. nov. (Pl. xi, figs. 17-7.) Monoicum, densiuscule cespitosum. Caulis erectus, brevis, 3-4 mil- lim. altus. Folia siccitate crispata, madore patentia, 2-3 millim. longa, e basi oblonga sat subito constricta, longe et anguste acuminato-subulata, apice sinuato denticulata, marginibus planis et integris, costa percur- rente, cellulis inferioribus rectangulis, 2-3 long. quam lat., superi- oribus irregularibus, plerisque subquadratis, opacis et papillosis, 9-15 p longis, 8-9 latis, cellulis alaribus majoribus, subinflatis, lutescenti- bus. Flos masculusin ramo breyi. Folia perichztialia intima vagi- nantia, longe acuminata. Capsula in pedicello brevi, 7-8 millim. longo, nutans vel inclinata, breviter ovato-conyexa, interdum strumulosa, levis vel vix striatula, operculo longe oblique rostrato, basi crenulato. Annulus distinctus. Peristomium elatum, intense purpureum, 0.5 millim. altum, valde papillosum, dentibus irregulariter bi-trifurcatis. Spore leves, 16-18 » crasse. From Port Wells (Trelease, 2268, 2271). This species is only comparable with C. polycarfam Sch., from which it is easily distinguished by its smaller size, its shorter, smooth or 256 CARDOT AND THERIOT : [296] hardly striate capsule, its more papillose, longer and brighter purple peristome, its shorter leaves with a thinner subula, and its upper cells smaller and less distinct. Cynodontium polycarpum alaskanum var. nov. A forma typica differt foliis apice tantum denticulatis, marginibus minus late et minus longe revolutis, reteque levi vel sublevi, cellulis superioribus paulo majoribus et distinctioribus (20 X 134, loco 14 x 11); a var. daxtrete Dix. foliis angustioribus et rete basilari densiore distinctum; ab Oxcophoro suectco Arn. et Jens. differt foliis inferne revolutis cellulisque alaribus indistinctis vel parum distinctis. From Juneau (Trelease, 2176) ; Cape Fox (Trelease, 2374) ; Indian Camp, Yakutat Bay (Brewer and Coe, 645). The type has been indicated for Alaska by Kellogg and by Les- quereux and James (Manual, p. 58). Cynodontium virens Sch. Br. eur. Coroll., p. 12. From Haenke Island (Coville and Kearney, 1110). A doubtful specimen from St. Matthew Island (Trelease, 2155). Cynodontium virens serratum Sch., loc. cit. From Haenke Island (Coville and Kearney, 1111); Egg Island (Coville and Kearney, 1016, 1017); Port Wells (Trelease, 2290; Brewer and Coe, 654); St. Matthew Island (Trelease, 1891). Cynodontium wahlenbergii Hartm. Flor. scand., ed. 10, p. 113. From Cape Vancouver (J. M. Macoun); Port Wells (Trelease, 1830, 2288 in part, 2289); Port Clarence (Brewer and Coe, 669) ; Hall Island (Trelease, 1882, 1898, 1899, 2130, 2132); St. Lawrence Island (Trelease, 1895, 1896, 1897, 2124) ; St. Matthew Island (Tre- lease, 1892, 1906, 2156, 2157, 2162). Number 2130 is a small form with short leaves, forma brevzfolia. Dichodontium pellucidum Sch. Br. eur. Coroll., p. 12. From Hidden Glacier Inlet in Yakutat Bay (Trelease, 1816, 2154 in part); Disenchantment Bay (Brewer and Coe, 639 in part) ; Muir Glacier (Trelease, 1752 in part) ; Port Wells (Trelease, 1831) ; Una- laska Island (J. M. Macoun). Dichodontium pellucidum fagimontanum Sch., loc. cit. From Juneau (Trelease, 2171); Muir Glacier (Trelease, 1909 in part). Numbers 1816, 2154 in part and 1831 are forms passing to var. Sagimontanum. [297] THE MOSSES OF ALASKA 257 Dichodontium pellucidum kodiakanum var. nov. (Pliexnryfig. 2925) Magnitudine D. flavescenté Lindb. simile, 5-8 centim. altum ; folia subintegra, apice late obtuso tantum sinuolata, rete vix papilloso. From Kodiak (Trelease, 1848). D. pellucidum serratum Sch. (D. flavescens Lindb.) has been recorded from Alaska by Kindberg. Aongstroemia longipes Br. eur., fasc. 33-36, p. 3, pl. 1. From Muir Glacier (Trelease, 2422, 2466, 2468); Hidden Glacier Inlet, in Yakutat Bay (Trelease, 2519). New to Alaska. Dicranella crispa Sch. Br. eur. Coroll., p. 13. From Yakutat Bay (Trelease, 2334); St. Lawrence Island (J. M. Macoun). Dicranella grevilleana Sch., loc. cit. From Port Clarence (Trelease, 2103). New to Alaska. Dicranella rufescens Sch., loc. cit. From Prince of Wales Island (J. M. Macoun). Dicranella heteromalla Sch., loc. cit. From Juneau (Trelease, 2180; Setchell, 1235); Farragut Bay (Coville and Kearney, 470) ; Kodiak (Trelease, 2206, 2213, 2214) ; Douglas Island (Trelease, 2405, 2407, 2411); Prince of Wales Island (J. M. Macoun) ; Yes Bay (Gorman, 182 in part, 183). Dicranella heteromalla orthophylla Lesq. & Jam. Manual, p. 67. From Sitka (Trelease, 2367); Kodiak (Trelease, 2197); Doug- las Island (Trelease, 2411). Dicranella heteromalla latinervis var. nov. A forma typica differt foliis brevius subulatis costaque latiore, circa 14 basis occupante. From Douglas Island (Trelease, 2389). D. squarrosa Sch. was recorded from Alaska, teste M. W. Harrington, by Lesquereux and James, and D. subulata Sch. and D. polarts Kindb. from the islands of Bering Sea, teste Macoun, by Kindberg. The specimen received by us as D. subulata, from St. Lawrence Island is D. crisfa. We have not seen any specimens of the other two species. Dicranum anderssonii Sch. Syn., ed. 1, p. 689. From Port Wells (Trelease, 2277). New to Alaska. 258 CARDOT AND THERIOT ; [298] This moss undoubtedly belongs to D. anxderssontd Sch. (Arctoa anderssoniz Wich. in Flora, 1859, no. 27). Inhis second edition of the Synopsis, Schimper reunites it to D. hyferboreum, from which, however, it seems sufficiently distinct by its smaller, sub- globose capsule, turbinate after the fall of the lid, very shortly pedi- cellate and almost always surrounded and surpassed by the perichetial leaves, and by the cells of the exothecium being smaller, with more solid and more colored walls. According to C. Jensen (Bryo- phyta of the Faeroes, in his Botany of the Faeroes, p. 159), the capsule of Dicranum anderssonti should be destitute of stomates, but in the Port Wells specimen all the capsules bear several stomates at the base of the very short neck. Dicranum starkei Web. & Mohr, Bot. Taschb., pp. 189, 471. From Yakutat Bay (Trelease, 2059) ; Disenchantment Bay (Brewer and Coe, 635). New to Alaska. Dicranum albicans Br. eur., fasc. 43, Suppl., pl. r. From Yakutat Bay (Trelease, 2059 in part). New to Alaska. A form of greener tint than usual and with homomallous leaves. Dicranum strictum Schl. Pl. crypt. helv., cent. m1, no. 26. From Yakutat Bay (Trelease, 2331, 2336). Dicranum subflagellare sp. nov. (Plcxmx figs 325°.) A D. flagellari proximo differt defectu flagellarum, foliis erectis subfalcatis angustioribus parum flexuosis, brevioribus (2—2.5 millim.), acumine canaliculato non subtubuloso, marginibus dorsoque subintegro cellulisque inferioribus angustioribus, superioribus multo minoribus. Specimina pauca, sterilia. From Kodiak (Trelease, 1899). Dicranum elongatum Schl. Pl. crypt. helv., cent. m1, no. 27. From Port Clarence (Trelease, 1867 in part, 2117, 2118) ; Kodiak (Trelease, 2503); St. Lawrence Island (Trelease, 1892, 1894, 1897 in part); St. Matthew Island (Trelease, 2170). Number 2503 is a short, stunted form, with nearly the facies of D. migquelonense Ren. & Card. Dicranum groenlandicum Brid. Mant. musc., p. 68. Bryol. uniy., I, p. 460. From the Yukon River (W. H. Dall); Port Clarence (Trelease, 1867 in part). New to Alaska. [299] THE MOSSES OF ALASKA 259 Dicranum fuscescens Turn. Muse. hib., p. 60, pl. 5, f. 1. From Yes Bay (Gorman, 184); Juneau (Brewer and Coe, 6994, 400); Skagway (Canby, 478); Wrangell (Trelease, 2317); White Pass, 3,000 ft. (Trelease, 2313); Farragut Bay (Trelease, 2416; Brewer and Coe, 617); Yakutat Bay (Trelease, 1763, 2340); Point Gustavus (Coville and Kearney, 777) ; La Perouse Glacier (Trelease, 2498); Sitka (U. S. S. Albatross, 53,55; Canby, 458, 463; Trelease, 2359); Hot Springs (Trelease, 1914); Kodiak (Trelease, 1853) 3 Douglas Island (Trelease, 1908, 2392) ; Port Wells (Trelease, 2282, 2288); Orca (Trelease, 1918, 1925, 2262; Setchell, 1213); New Metlakatla (Coville and Kearney, 364.) ; Plover Bay, Siberia (Trelease, 1865). The Yes Bay specimen was associated with Mntum glabrescens Kindb. and Scapanza sp. The Kodiak specimen (1853) is a paludal form, with entire, shorter, erect leaves. We must point out too a sterile form, collected on Unalaska Island by Mr. C. H. Townsend (U.S. 8. Albatross exped., 43), which is very near D. muehlenbeckii Br. eur. var. brevifolium Lindb. The same form was found by Trelease on Hall Island (1907). It differs from the European plant chiefly in having its leaves less flexuous when dry. Many authors separate D. congestum Brid. from D. fuscescens Turn. but as it is almost impossible to find two descriptions of D. congestum which agree, we can infer that it is one of those species which every author understands in his own way, that is to say a very bad species; and we deem it preferable to merely unite it with D. fuscescens. Dicranum dipteroneuron C. Mill. in Flora, 1887, p. 221. From valley of the Takhin River (Krause brothers). Dicranum scoparium Hedw. Fund. musc., uy, p. 92, pl. 8, f. 41, 42. From Point Gustavus (Coville and Kearney, 772 in part.) A paludal form. Dicranum neglectum Jur. Laubm. fl., p. 47. Limpr. Laubm.,1, p. 353. D. spadiceum ZETT. Musc. pyren., p. 30. From Port Clarence (Trelease, 1868) ; Sturgeon River Bay, Kodiak (Trelease, 1929); Unalaska (U. S. S. Albatross exped., 7a); St. George Island (U. S. S. Albatross exped., 57); St. Lawrence Island (Trelease, 1873, 1874 in part, 1902); St. Matthew Island (Trelease, 1886, 1888 in part, 1980) ; Hall Island (Trelease, 1988, 1990, 1991). New to Alaska. 260 CARDOT AND THERIOT : [300] Dicranum howellii Ren. & Card. in Bot. Gaz., 1889, p. 93, pl. xu, B. From Skagway (Canby, 483); New Metlakatla (Trelease, 1906) ; Point Gustavus (Coville and Kearney, 755); Virgin Bay (Trelease, 2308); Sitka (Setchell, 1255; Trelease, 2360); Kodiak (Trelease, 2202, 2223, 2504); Unga (Saunders, 2294, 2295). New to Alaska. Dicranum bonjeani De Not. apud Lisa Elencho, p.29. Epil., p. 616. From Sturgeon River Bay, Kodiak (Trelease, 1854). A form nearly allied to var. schlotthauwert Barnes by its short and entire leaves. The type was recorded from Sitka and Nulato, teste Rothrock, by Kindberg, Mac. Cat. Can. pl., v1, Musci, p. 32. Dicranum majus Sm. FI. brit., m1, p. 1202. From Point Gustavus (Coville and Kearney, 754) ; Orca (Setchell, 1215); Sitka (Trelease, 1953; U.S. S. Albatross exped., 61); Hot Springs (Trelease, 1952). Number 1215 Setchell, is a slender form. Dicranum bergeri Bland. Musc. frond. exs., 111, no. 114. From Kodiak (Trelease, 1675a in part). Dicranum molle Wils. is recorded from St. Paul Island and D. angustifolium Kindb. from Unalaska by Kindberg, in Mac. Cat. Can. pl., vi, Musci. D. muchlenbeckit has also been recorded from Alaska. Dicranodontium longirostre Br. eur., fasc. 41, p. 2, pl. I. From Orca (Trelease, 1839). New to Alaska. Dicranodontium aristatum Sch. Syn., ed. 1, p. 695. From Hot Springs (Trelease, 1809). New to Alaska. Slightly different from the European plant, the costa being rather less broad, but one fifth or one fourth the width of the leaf base, and having a less rough subula. Décranum virginicum Aust. ( Cam- pylopus virginicus Lesq. & Jam.) is intermediate between the Euro- pean and Alaskan forms. Mr. Kindberg has, teste Macoun, reported from Alaska, Campylopus schimperi Milde. Family SELIGERIACEZ. Blindia acuta Br. eur., fasc. 33-36, p. 3, pl. 1. From Juneau (Coville and Kearney, 582) ; Port Wells (Trelease, 2274, 2275, 2276, 2280, 2291 in part); Hall Island (Trelease, 1881). New to Alaska. Number 2280 is a form with long innovations, surpassing the capsule. Blindia acuta flexipes Ren. & Card., in Rev. bryol., 1892, p. 79. Port Wells (Trelease, 2286). New to Alaska. [301] THE MOSSES OF ALASKA 261 Family DITRICHACEZ. Ceratodon purpureus Brid. Br. univ., 1, p. 48o. Very common and yariable. From Juneau (Setchell, 1233; Co- ville and Kearney, 589; Trelease, 2175, 2177); New Metlakatla (Trelease, 2240); Wrangell (Trelease, 2316); Port Clarence (Tre- lease, 2106, 2107, 2108, 2109, 2110, 2111, 2112; Brewer and Coe, 671); Cape Fox (Trelease, 1934); Yakutat Bay (Trelease, 2326, 2332); Disenchantment Bay (Coville and Kearney, 1065; Brewer and Coe, 637); Muir Glacier (Trelease, 1801, 1804, 2443, 2446, 2447, 2450, 2465); Orca (Trelease, 2255, 2256); Bogoslof volcano (Coville and Kearney, 2614 in part); Sitka (Coville and Kearney, 868; Trelease, 2361; Canby, 460); Kodiak (Trelease, 2189, 2198, 2212, 2224, 2226); Douglas Island (Trelease, 2393, 2404); Prince of Wales Island (J. M. Macoun); Bering Island (J. M. Macoun) ; Hall Island ( Trelease, 2139); St. Paul Island (Trelease, 1860, 2086) ; Plover Bay, Siberia (Trelease, 2097, 2531, 2532, 2536, 2537, 2546; Coville and Kearney, 1862). ‘Ceratodon heterophyllus Kindb. in Ott. nat., v, p. 179. Macoun, Cat. Can. pl., v1, Musci, p. 261. From St. Paul Island (J. M. Macoun; B. W. Evermann; Tre- lease, 2062, 2070, 2071, 2072, 2075). Distichium capillaceum Br. eur., fasc. 29-30, p. 4, pl. 1. From White Pass (Trelease, 2310) ; Port Wells (Brewer and Coe, 653; Coville and Kearney, 1291 in part; Trelease, 2278, 2291); Orca (Trelease, 1837, 1838, 2260 in part); Yakutat Bay (Brewer and Coe, 644); Port Clarence (Trelease, 2104). Ditrichum homallum Hpe. in Flora, 1867, p. 182. From Kodiak (Brewer and Coe, 658 in part). New to Alaska. A slender etiolated form, of which we find only some stems among other mosses; leaves erect, costa narrower, basilar cells broader. Ditrichum flexicaule densum Sch. has, teste Rothrock, been recorded from Alaska by Lesquereux and James, and D. glaucescens Hpe., teste Macoun, from Unalaska, by Kindberg. Kindberg has also, teste Macoun, described a Lefto- trichum tomentosum from St. Paul Island. Family POTTIACEA. Pottia heimii beringiana var. nov, (BI Sscivaifie:2 ao*.)) A forma typica differt foliis brevioribus, limbo lutescente circum- ductis, costa breviter excurrente mucronatis reteque magis opaco valde papilloso. 262 CARDOT AND THERIOT F [302] From St. Matthew Island (Trelease, 2151 in part). We found only a few stems of this moss, mixed with Barbula brachypoda Card. & Thér. and a Bryum. By the pellucid margin of the leaves, it is nearly related to P. odbtus¢folia C. Miill. (P. heimti arctica Lindb.), but it is easily distinguished from it by its pointed leaves. It also closely resembles Desmatodon systyliotdes Ren. & Card., from Labrador, which is probably also a Pottza of the same group, but it differs from this species by its ovate, shorter and proportionately wider capsule, its longer and more finely beaked lid, its shorter and more briefly acuminate leaves, and finally by its areola- tion formed of larger and less obscure cells. According to J. M. Macoun P. hezmit typica has been recorded by Kindberg from St. Matthew Island and from Bering Island. Didymodon rvubellus Br. eur., fasc. 29-30, p. 3, pl. 1. From Juneau (Trelease, 2178) ; Port Wells (Trelease, 2272, 2283) ; Agattu Island (U. 8. S. Aléatross exped., 36); St. Matthew Island (Trelease, 2143, specimen in bad condition, and determination doubt- ful) ; Bering Island (J. M. Macoun). Didymodon baden-powelli Kindb. Ott. Nat., v, p. 179. Macoun, Cat. Can. pl., v1, Musci, p. 262. From St. Paul Island (J. M. Macoun). Contrary to Kindberg’s description, the leaves are entire or nearly so on the specimen we have seen. It is probable that this moss is but a form of D. rubellus. Trichostomum cuspidatissimum sp. nov. (Pl. xin, fig. 4°—*.) Dioicum, elatum, compacte cespitosum, fusco-lutescens. Caulis erectus, circa 5 centim. altus, dense foliosus, rufo-tomentosus, ramis erectis numerosis. Folia siccitate crispata, madore erecto-arcuata, ad apicem caulis et ramorum congesta, fragilia (acumine sepe effracto), e basi ovata sensim et longissime acuminata, 3-3.5 millim. longa, 0.45- 0.7 lata, marginibus planis, superne inflexis, papillis prominentibus crenulatis, costa angusta, 80-90 » basi lata, in cuspidem acutissimam longe excurrente, cellulis inferioribus rectangulis, 5-6 long. quam lat., lutescentibus, superioribus plerumque quadratis, 10-12 y latis, opacis, grosse papillosis. Cztera desunt. From Hall Island (Brewer and Coe, 674). A fine species, easily distinguished from the large forms of 7. mata- ble Bruch, and its var. cuspidatum Limpr. (Z. cuspidatum Sch.), [303] THE MOSSES OF ALASKA 263 by the form of the leaves, and more particularly by its narrower costa and its upper cells which are much larger and more distinct (10-12 » instead of 6-8). Trichostomum sitkanum sp. nov. (Pl. x1v, fig. 17.) Dioicum? Cespites densiusculi, superne virides, intus nigrescentes, inferne terra obruti. Caulis 2-3 centim. altus, parum divisus, laxe foliosus. Folia mollia, sicca cirrato-crispata, madore patula, inferiora 2.5-3 millim. longa, superiora majora, 4—4.5 millim. longa, e basi longe subvaginante lineari-lanceolata, marginibus planis integris, costa sat valida, basi circa 110 p» lata, in mucronem lutescentem breviter ex- currente, cellulis basis subvaginantis rectangulis, hyalinis, 4-6 long. quam lat., cateris quadrato-hexagonis, 12-14 » latis, valde papillosis. Cetera desunt. From Sitka (Trelease, 2370). The aspect, the form of leaves, the looser basal areolation and the upper cells more papillose, easily distinguish this plant from 7. cuspi- datissimum Card. & Thér. It is more closely connected with 7. bambergert Sch., but the latter has the costa shining on the back in a dry state, and the hyaline cells of the base going up along the borders of the leaf, as in Barbula tortuosa Web. & Mohr. Desmatodon latifolius Br. eur., fasc. 18-20, p. 5, pl. 1. From Unalaska (J. M. Macoun). Barbula brachypoda sp. nov. (Pl. x1v, figs. 37+.) Monoica, laxiuscule cespitosa, viridis. Caulis 5-10 millim. altus, erectus, simplex furcatusve. Folia mollia, sicca erecta, madida erecto- patentia, elliptica vel subspathulata, 2.5—3 millim. longa, 0.6—1 millim. lata, late et breviter acuminata obtusa, subobtusa acutave, marginibus e basi ad medium usque revyolutis, deinde planis et pro more limbo lutes- cente e 3-4 seriebus cellularum composito limbatis, costa angusta (lat. 50/4), paulo sub apice evanida, rete levi, in dimidio inferiore laxissimo hyalino, cellulis rectangularibus, 66-SS » longis, circa 22 latis, superne valde chlorophylloso, cellulis inequalibus quadrato-rotundatis vel brevis- sime rectangulis, e costa ad margines sensim minoribus (majoribus 25 longis, 16 latis, minoribus quadratis, 144 latis). Flos masculus infra femineum situs. Folia perichetialia caulinis majora, acuminata, intima angustissima. Capsula in pedicello brevi, crassiusculo, pallido, 264 CARDOT AND THERIOT [304] 5-7 millim. longo, o.2 millim. crasso, erecta, cylindrica, circa 2 millim. longa, operculo conico tertiam partem capsule wquante. Annulus latus, distinctus. Peristomium papillosum, membrana _ basilari brevi, dentibus semel vel bis convolutis. Spore leves, 12-16 » crassa. From St. Matthew Island (Trelease, 2151 in part, 2166). This species, which belongs to the section Cumnezfolie Sch., is easily distinguished from &. cunezfolia Brid. by its leaves revolute be- low, its longer lower cells, the upper more chlorophyllose with thicker walls, its shorter seta, its broad annulus, etc. Kindberg has established (in Revue bryologique, 1896, p. 22) a B. subcunetfolia from Alaska, which, by some characters, seems to be near to our B. brachypfoda; but his description is so incomplete that we can neither ascertain whether this B. swbcunezfolza is identi- cal with the plant here described, nor, with still more reason, mention the characters which might distinguish it from the former, Barbula saundersii sp. nov. (Pl. xvi, figs 165) Dioica? laxiuscule cespitosa, olivaceo-viridis. Caulis brevis, 5-8 millim. altus, simplex vel parce divisus. Folia sicca incurvata, madida erecta vel erecto-patentia, 1.5-2 millim. longa, 0.8 lata, e basi ovata breviter acuminata, obtusa vel brevissime mucronata, marginibus e basi usque ad 24 valde revoluta, superne concava, costa valida, tota fere longitudine squaliter crassa (0.1 millim.), dorso valde promi- nente, percurrente, rarius paulisper excedente; rete levi, cellulis in- ferioribus laxis, rectangulis, lutescentibus, 4o longis, 12 latis, se- quentibus quadratis, 12-13 latis, parietibus incrassatis, mediis et superioribus minutis, 7-8 latis, parum distinctis. Folia perichetialia caulinis majora, sicca erecto-incuryata, madida erecta, appressa. Cap- sula in pedicello purpureo, circa 10 millim. longo, siccitate sinistror- sum torto, erecta, oblonga vel subcylindrica, 1-1.25 millim. longa, operculo conico longissimo, capsulam zquante. Peristomium pur- pureum, papillosum, membrana basilari brevi, 30 alta, dentibus semel vel bis contortis. Spore leves, 14-16 crasse. From Hidden Glacier Inlet, Yakutat Bay (Trelease, 2514). In habit recalls the smallest forms of B. ungutculata Hedw., from which it differs by the shorter and proportionately broader leaves, which are shortly ovate-lanceolate, not or hardly mucronate, etc. By the form of the leaves, it is also connected with B. drachyphylla Sulliv., but the latter has the stems much longer and the basal areolation of the leaves quite different. [305 ] THE MOSSES OF ALASKA 265 Barbula treleasei sp. nov. (Pl. xv, fig. 2*—9.) Dioica? dense cespitosa, lutescenti-viridis. Caulis 1-2 centim. altus, erectus, divisus. Folia siccitate crispatula, madore erecto-pa- tentia, circa 1.5 millim. longa, 0.6 lata, ovato-lanceolata, breviter acuminata, integra, marginibus e basi longe revolutis, costa valida, rubella, percurrente, basi 80 » crassa, cellulis inferioribus rectangulis subhyalinis, levibus, mediis superioribusque minutis (diam. 5 4), quadratis, valde papillosis, parum distinctis. Folia perichetialia multo longiora et latiora, sat subito constricta, longe acuminata, madore arcuato-patula, intima subvaginantia, in dimidio inferiore hyalina. Capsula in pedicello 10-11 millim. longo, erecta, oblongo-cylindrica. Cetera desunt. From Juneau (Trelease, 2179, 2181). This moss has the aspect of a slender B. fallax Hedw.; but the texture of the leaf base, formed of rectangular subhyaline cells, clearly separates it. On the other hand, it differs from 2. véxeals Brid. and allied forms by the much shorter leaves. Barbula rigens sp. noy. (Pl. xv, fig. 1°79.) Rubella, laxe cespitosa vel aliis muscis gregarie intermixta. Caulis gracilis, erectus, rigidulus, divisus, 1-2 centim. altus. Folia sicca crispatula, madida erecto-patentia, stricta, breviter lanceolato-linearia, 1.25-1.5 millim. longa, 0.3 lata, marginibus integris medium versus paululum revolutis, costa valida, tota fere longitudine zqualiter crassa, diam. 56 », biconvexa, percurrente vel in mucronem brevem excurrente, cellulis inferioribus rectangulis, hyalinis, plerumque levibus, superiori- bus opacis, indistinctis, quadrato-rotundatis, utraque pagina dense papillosis, mediis circa 11 y latis. Cetera ignota. From Orca (Trelease, 2260; mixed with Déstichium capillaceum and Anactangium compactum). This species belongs to the group of B. rig¢dula Mitt., from which it is easily distinguished by its stiff stem and leaves, the latter being shorter and very briefly acuminate and by its much more papillose areolation. Barbula cylindrica Sch. in Hedwigia, 1873, p.47. Syn.,ed. 2, p. 208. From Prince of Wales Island (J. M. Macoun). Barbula fragilis Br. eur., fasc. 62-64, Suppl., pl. 4. From Port Wells (Coville and Kearney, 1291 in part). 266 CARDOT AND THERIOT ; [306] Barbula aciphylla Br. eur., fasc. 13-15, p. 42, pl. 26. Yakutat Bay (Trelease, 1746 in part); Muir Glacier (Trelease, 1802). New to Alaska. Number 1802 is a rather badly characterized form, which can be ascribed to B. aczéphylla, but which has also some relationship with B. ruralis Hedw. Barbula ruralis Hedw. Fund., ny, p. 92. From Agattu Island (U. 8. S. Aldéatross Exped., 42). A form having the facies of B. muelleri Bruch. "Mr. Kindberg has described from Alaska a B. subcunetfolia and a B. ruralis subsp. alaskana, of which we have seen no specimens. Family GRIMMIACEZ. Grimmia apocarpa Hedw. Descr., 1, p. 104, pl. 39. From Juneau (Trelease, 2183); Muir Glacier (Trelease, 2432) ; Kodiak (Trelease, 2215); Hall Island (Trelease, 2128); St. Paul Island (Trelease, 2079). Grimmia apocarpa gracilis Web. & Mohr, Taschenb., p. 131. From White Pass (Trelease, 2493) ; Muir Glacier (Trelease, 1789, 2454); Hidden Glacier Inlet, Yakutat Bay (Trelease, 1790, 2058) ; Kodiak (Trelease, 1788) ; Hall Island (Trelease, 1880). Grimmia apocarpa alpicola Hook. & Tayl. Muscol. brit., p. 87. From Portage Bay (U. S. S. Albatross Exped.) ; Cape Fox (Tre- lease, 2386); St. Paul Island (Trelease, 2080). A form allied to this variety by the dimension of its spores, but differing from it by its sharp pointed leaves, the upper ones ending in a hyaline point, was collected at Wrangell (Canby, 471) ; Cape Fox (Trelease, 2385) and at Hot Springs, near Sitka (Trelease, 2495). Grimmia apocarpa rivularis Web. & Mohr, Taschenb., p. 129. From Muir Glacier (Trelease, 2424, 2494); Yakutat Bay (Tre- lease, 2324). Grimmia conferta Funck, Moostaschenb., p. 18, pl. 12. From St. Paul Island (Trelease, 2470@); St. Matthew Island (Tre- lease, 2167). Seems to belong to G. conferta by the short, hemispherical capsule, but the peristome is wanting. Grimmia maritima Turn. Muscol. hib., p. 23, pl. 3, f. 2. From Virgin Bay (Trelease, 2303) ; Port Wells (Trelease, 2279) ; Yakutat Bay (Trelease, 2325); Kodiak (Trelease, 2205, 2216); Agattu Island (U. S. S. Aléatross Exped., 62). New to Alaska. [307] THE MOSSES OF ALASKA 267 Grimmia torquata Grev. Scot. crypt. fl., Iv, p. 199. From Kodiak (Trelease, 2203, 2204). Grimmia elatior Br. eur., fasc. 25-28, p. 17, pl. 10, forma? From Yukon River (W. H. Dall). New to Alaska. We find only some stems of this moss, with a single capsule, mixed with Polytrichum yukonense Card. & Thér., and their determina- tion remains rather doubtful. Kindberg has recorded from Alaska G. agassizi? Sulliv. and Lesq. and from Unalaska G. crassinervis C. Miill. Rhacomitrium patens Hiib. Muscol. germ., p. 198. From Unalaska (J. M. Macoun). Rhacomitrium sudeticum alaskanum var. nov. Forma minor, habitu varietati fexe//um Boul. similis, sed foliis subepiliferis vel apiculo hyalino omnino destitutis costaque validiore distincta. ; Hidden Glacier Inlet, Yakutat Bay (Trelease, 2508 in part). Rhacomitrium aciculare Brid. Mant., p. 80. From Juneau (Coville and Kearney, 573); Kodiak (Trelease, 1849). Rhacomitrium nevii Wats. Bot. Calif., m, p. 381. From Juneau (Trelease, 2174); Atka Island (J. M. Macoun). Rhacomitrium fasciculare Brid. Mant., p. So. From Portage Bay (U. S. S. Albatross Exped.); Yakutat Bay (Trelease, 1785, 2322); Cape Fox (Trelease, 2377); Muir Glacier (Trelease, 1781, 2455) ; Kodiak (Trelease, 1786, 2193) ; Sitka (U. S. S. Albatross Exped., 47); Hot Springs (Trelease, 1769, 2346, 2350). Number 2322 is a forma mor. Rhacomitrium tenuinerve Kindb. Rev. bryol., 1896, p. 19. R. fasciculare var. haplocladon K1NDB. Not. on Can. bryol., 1893. R. microcarpum var. palmeri KINDB. apud Macoun, Cat. Can. pl., v1, Musci, R. pu en Rev. bryol., 1896, p. 19. From St. Paul Island (J. M. Macoun); St. Matthew Island (Tre- lease, 1885, 2169); Pribilof Islands (Palmer, 1891). R. tenutnerve and palmert of Kindberg surely constitute but one species, which differs from A. fasctculare Brid. principally by its weak, flat costa, disappearing far from the point. Kindberg is mis- taken inattributing to #. Ja/mer¢ a percurrent or subexcurrent costa. 268 CARDOT AND THERIOT : [308] On the original specimen collected by Palmer, and which Kindberg himself formerly communicated to us, the costa has exactly the same length and the same structure as in A. tenuznerve. The latter is a form with long simple or hardly branched stems, whereas PR. palmeri is a shorter and more ramulose form. Rhacomitrium cyclodictyon sp. noy. (Plate xv, fig. 37-%.) Dioicum? parvum, dense cespitosum, atrofuscum. Caulis depressus, ramosissimus, ramis confertis, erectis, brevibus, 3-5 millim. longis. Folia siccitate suberecta vix flexuosa, madore erecto-patentia, 1.25 mil- lim. longa, 0.5 lata, ovato-lanceolata, mutica, integerrima, inferne mar- ginibus revoluta, costa sat tenui, 35-40 » crassa, paulo sub apice eva- nida, rete subequali, cellulis infimis juxta costam paucis rectangulis vel sublinearibus, haud sinuosis, omnibus czteris rotundatis vel brevissime ovatis, 8-12 latis, parietibus incrassatis, levibus sed valde convexis, ita ut papillas maximas emulent. Folia perichetialia multo majora, e basi subvaginante sensim et longe acuminata, madore erecta. Cap- sula in pedicello brevi, purpureo, demum nigricante, siccitate sinistror- sum torto, 5 millim. longo, erecta, anguste cylindrica, 1.5 millim. longa, 0.3 crassa. Spore minute granulose, diam. 16-17. Cetera ignota. From Muir Glacier (Trelease, 2431). A most remarkable species, which cannot be mistaken for any other on account of its characteristic areolation very different from that of all known species of the genus Rhacomitrium. Rhacomitrium heterostichum Brid. Mant., p. 79. From Hot Springs (Trelease, 1773, 1774); Kodiak (Trelease, 1776); Orca (Trelease, 1961 in part). Rhacomitrium heterostichum affine (Schleich.) Card. and Thér. From Unalaska (Trelease, 2296). Rhacomitrium lanuginosum Brid. Mant., p. 79. From Juneau (Setchell, 1240) ; New Metlakatla (Trelease, 1949) ; Virgin Bay (Trelease, 1775); Sitka (Trelease, 1772); Kodiak (Brewer and Coe, 655); Hall Island (Trelease, 1777, 1778); St. Matthew Island (Trelease, 1856). Number 1949 is the form falcata Boul. Numbers 1777, 1778 and 1856 belong to a form stricta. (Branches rigid when dry, subdisti- chous ; leaves erect-appressed. ) [309] THE MOSSES OF ALASKA 269 Rhacomitrium canescens Brid. Mant., p. 78. From Orca (Setchell, 1211) ; Muir Glacier (Trelease, 17644, 1766, 1767, 1768, 2423, 2429, 2456); Hidden Glacier Inlet, Yakutat Bay (Trelease, 1780 in part); Disenchantment Bay (Trelease, 1779 in part) ; Unalaska (J. M. Macoun). Rhacomitrium canescens ericoides Br. eur., fasc. 25-28, p. 12, pl. 8, fig. From Yakutat (Trelease, 1794); Disenchantment Bay (Trelease, 1065 in part, 1770, 1779 in part, 2505, 2506; Brewer and Coe, 639, 640); Hubbard Glacier (Coville and Kearney, 1071, 1073 in part, 1065 in part); Hidden Glacier Inlet (Trelease, 1771, 1780 in part) ; Russell Fiord (Coville and Kearney, 995); Muir Glacier (Trelease, 1764, 1765, 2418, 2430, 2464); Muir Inlet (Coville and Kearney, 636) ; Point Gustavus (Coville and Kearney, 776). Numbers Trelease, 1780 in part, and Coville and Kearney, 1073 in part, constitute a form efzlosa or subepilosa. Coscinodon pulvinatus Spreng. has, teste M. W. Harrington, been recorded from Alaska by Lesquereux and James. Family ORTHOTRICHACEZ. Amphoridium lapponicum Sch. Syn., ed. 1, p. 247. From Orca, 1,200 ft. (Trelease, 2246); Port Wells (Brewer and Coe, 651) ; Yakutat Bay (Trelease, 2323). Amphoridium mougeotii Sch. Syn., ed. 1, p. 248. From Juneau (Coville and Kearney, 577). New to Alaska, Ulota drummondii Brid. Bryol. uniy., 1, p. 299. From Kodiak (Trelease, 2209); Unga (Saunders, 2292). Ulota phyllantha Brid. Mant., p. 113. From Bailey Harbor (U. S. S.