Number 61 January 3, 2015

Thomas K. Todsen Field Books — 1967-1981

Kelly W

Allred

20 1 5 Jordan Road, Las Cruces, NM 8800 1 kallred@nmsu.edu

One of New Mexico’s many exceptional amateur

Texas

botanists was Thomas Kamp Todsen (1 9 1 8-20 1 0; see The

El Paso Co.: 6

New Mexico Botanist 53: 1-3. 201 1.). A few years ago.

two of his field books were given to me by his son, Andy,

FB68-81

with the expectation that the information therein might be

Arizona

of interest to New Mexico botanists.

Cochise Co.: 16, 18, 48, 80, 83, 99

The first book, a 4 x 8 inch reporter-style notebook with

New Mexico

top-coil binding, is titled on the front cover, “Botanizing

Catron Co.: 24, 44

Notebook, T.K. Todsen, 1 968.” The notebook is

Colfax Co.: 44

apparently missing one or more initial pages, but the

Dona Ana Co.: 1,2,4, 8, 10, 12, 14,22,24,28,30,34,

second intact page has an unambiguous date of 1 6

36, 38, 40, 42, 46, 48, 50, 52, 62, 64, 68, 70, 72, 74,

September 1 967. Recording ends slightly more than one

76, 78, 86, 88, 89, 93, 94, 95, 96, 97, 100, 102

year later on 28 September 1 968. Notes are recorded on

Grant Co.: 12, 14, 16,26,28,38,40,42,44, 50, 74,81,

only one side of each page. This notebook will be referred

85

to as FB67-68.

Guadalupe Co.: 44

The second book is a 4.5 x 7.25 inch hard-cover Transit

Hidalgo Co.: 2,6, 50, 52, 76, 87, 88, 1 00, 1 04

Book (the cover barely attached by a few threads) with 1 7

Lincoln Co.: 20, 22, 30, 50

preliminary printed pages of “Improved Tables and

Luna Co.: 60, 84

Information.” It is missing the first six pages of notes

McKinley Co.: 32

(snipped out of the notebook), and begins with notes from

Otero Co.: 1 6, 1 8, 20, 22, 26, 30, 34, 36, 48, 60, 62, 74,

1 5 September 1 968, and ends on 26 July 1981. Notes are

141

recorded on both sides of each page. This book will be

San Juan Co.: 68

referred to as FB68-81 .

San Miguel Co.: 24, 26

To help in searching localities, the pages of the field

Sierra Co.: 26, 28, 34, 42, 44, 46, 52, 64, 89, 98, 1 03,

books are indexed by county (United States) or state

105, 106, 107

(Mexico):

Socorro Co.: 72, 78

Taos Co.: 48

FB67-68

Torrance Co.: 42

Arizona

Utah

Cochise Co.: 23

San Juan Co.: 32

New Mexico

Mexico

Bernalillo Co.: 1 9

Sonora: 54, 58, 60

Dona Ana Co.: 2, 3, 4, 5, 6, 7, 9, 11, 12, 18

Grant Co.: 1, 13, 14, 17, 21

Each page of the field books that contained notes was

Hidalgo Co.: 2, 1 6

scanned, and are reproduced on the following pages. A

Lincoln Co.: 1 5, 20, 21

page number was added in the upper corner (left or right).

Los Alamos Co.: 19

to reference the list of counties.

Otero Co.: 14, 1 5

Sierra Co.: 3, 14, 17

Taos Co.: 19

^otanice' est ^cientia ^jyntutnUs quae '^e0etabiHuni co0nitiorem tradit.

— ^/Hnnacus

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Number 62

February 24, 2015

A Comparison of the Mosses Known and Reported for New Mexico with
Flora North America, and a Current Catalog of the Mosses of New Mexico

Kelly W. Allred

Range Science Herbarium, Department of Animal & Range Sciences (emeritus)

New Mexico State University, Las Cruces, NM 88003 kallred@nmsu.edu

The recent completion of the two volumes of Flora North
America that deal with the mosses (BFNA27-2007; BFNA28-
2014) allows for a fuller accounting of the moss flora of New
Mexico. I enumerate, first, mosses new to New Mexico as
recorded in BFNA28 in 2014. (New records from BFNA27-
2007, have already been incorporated into our list.) Second, I
list, with references, moss species that were reported for New
Mexico prior to 2007, but were not included in BFNA27-
2007. Third, I list, with references, moss species that were
reported for New Mexico prior to 2014, but were not included
in BFNA28-2014. Fourth, I itemize mosses that were not
recorded for New Mexico in BFNA27-2007, but have been
found and reported for the state since the publication of that
volume. Lastly, an up-to-date catalog of the mosses of New
Mexico is provided, with validating references.

1.

7

3.

4.

5.

6.

7.

8.

9.

10.

1.

1. New Records for New Mexico from BFNA28-2014

Bartramia stricta Bridel

Brachymeniwn macrocarpiim Cardot : This validates that
our material is not Brachymenium klotzschii as reported in
Conard & Redfearn (1979)

Btyiim argenteum Hedwig var. muticum Bridel
Entodon beyrichii Schwagrichen
Gemmahryum demaretiammi (Arts) J.R. Spence
Genunabtyitm exile (Dozy & Molkenboer) J.R. Spence &
H.P Ramsay

Gemmabryiim kiinzei (Hornschuch) J.R. Spence
Hygrohypnum bestii (Renauld & Bryhn) Holzinger
Hygrohypmim diiriiisciiliini (DeNotaris) D.W. Jamieson
Pohlia bolanderi (Sullivant) Brotherus

II.

Reported for New Mexico Prior to 2007,

but not recorded in BFNA27-2007

Andreaea heinemannii Hampe & Muller Flal. [Mahler
1985]

2. Biyoeryihrophylluin recui'virostrum (Hedwig)

Chen [Bartram 1931; Mahler 1978]

3. Dichodontium pelliicidum (Hedwig) Schimper [Allred et
al. 1998]

4. Dicranella vona (Hedwig) Schimper [Ireland et al. 1981]

5. Dicranoweissia cirrata (Hedwig) Lindberg ex
Milde [Mahler 1978]

6. Didymodon rigidulus Hedwig var. icmadophilus (Muller
Hal.) Zander [Allred et al. 1998]

7. Ditrichum flexicaide (Schwagrichen) Hampe [Allred et
al. 1998]

8. Encalypta rhaptocarpa Schwagrichen [Conard &
Redfearn 1979]

9. Funaria apiculatopilosa Cardot [Worthington 1996]

10. Funaria hygrometrica Hedwig var. calvescens
(Schwagrichen) Montagne [Darigo & Allred 2001]

1 1 . Funaria muhlenbergii Turner [Stark & Castetter 1982]

12. Grimmia elatior Bruch ex Balsamo-Crivelli & De
Notaris [Darigo & Allred 2001]

13. Grimmia orbicularis Bruch ex V^ilson [Mahler 1978]

14. Schistidium ambiguum SuWlvant [Sullivant 1849]

15. Lorentziella imbricata(Mltter\) Brotherus [Allred et al.
1998]

16. Microbryum starkeanum (Hedwig) R.H. Zander
\ar. star keanum [Ireland et al. 1981]

17. Physcomitrium pyriforme (Hedwig) Hampe [Ireland et al.
1984]

1 8. Polytrichastrum sexangulare (Bridel) G.L.

Smith [Mahler 1978]

19. Saelania glaucescens (Hedwig) Brotherus [Bartram
1931; Mahler 1978]

20. Sphagnum fimbriatum [Coop 2002]

21. Tetraphis pellucidaHedwlg [Allred et al. 1998]

22. Tortella humilis (Hedwig) ]eunlr\gs [Ireland et al. 1981]

23. Tortula brevipes (Lesquereux) Brotherus [Allred et al.
1998]

24. Tortula cernua (Huberner) Lindberg [Crum & Anderson
1981]

25. Tortula protobryoides R.H. Zander [Allred et al. 1998]

26. Trichostomum brachydontium Bruch [Ireland et al. 1984]

27. Trichostomum tenuirostre (Hooker & Taylor)

Lindberg [Ireland et al. 1984]

(Continued on page 2)

^ot«mcc' est ^cientia ^oturalis co9mtioreiii tradit.

— ^innncus

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III. Reported for New Mexico Prior to 2014.

but not recorded in BFNA28-2014

1. Anomodonjiilacewn [Allred 2012]

2. (Rohling) Bruch & Schimper [Romig &
Allred 2013]

3. Bartramia ithyphylla [Bartram 1 93 1 ; Mahler 1978]

4. Brachythecium acuminatum [Bartram 1931; Mahler 1978]

5. Brachythecium albicans [Stark & Castetter 1982]

6. Campylophyllum hispidulum [Bartram 1931; Mahler 1978]

7. Campylophyllum hispidulum [Bartram 1931; Mahler 1978]

8. Dicranum tauricum Sapjeg'm [Romig & Allred 2013]

9. Drepanocladus longifolius [Allred 2012]

10. Entodon seducirix [Blisard & Kleinman 201 2]

11. Fabronia pusilla [Mahler 1978]

12. Gemmabryum dichotomum [Allred et al. 1998]

13. Homomallium inciirvatiim [Allred 2012]

14. I/ygrohypnum ochraceum [Mahler 1985]

15. Hylocomium splendens [Bartam 193 1 ; Mahler 1978]

16. llypnum pratense [Allred et al. 1998]

17. Imbribtyum miniatum [Stark & Castetter 1982]

18. Isopterygium tenerum [Allred et al. 1998]

19. Lesciiraea saxicola [Mahler 1985]

20. Leskea angustata [Worthington 2012]

21. Orthotrichum cupiilatum [Worthington 2012]

22. Orthotrichum laevigatum [Allred et al. 1998]

23. Orthotrichum pollens Bridel [Romig & Allred 2013]

24. Oxyrrhvnchium hians [Allred 2012; Conard & Redfearn
1979; Grout 1928-40]

25. Philonotis capillaris [Mahler 1978]

26. Philonotis fontana (Hedwig) Bridel var. americana
(Dismier) Flowers ex Crum [Flowers 1973]

27. Philonotis fontana Bridel var. pumila (Turner)

Bridel [Bartram 1931; Mahler 1978]

28. Plagiomnium ciliare [Allred 2012]

29. Plagiomnium ellipticiim [Allred et al. 1998]

30. Plagiomnium medium [Allred 2012]

31. Plagiomnium rostratum [Bartram 193 1 ; Mahler 1 978]

32. Platydictya subtilis [Allred 20 12]

33. Platygyrium repens [Allred 2012]

34. Pohlia obtusifolia [Worthington 2012]

35. Pseudocampylium radicale [Mahler 1985]

36. Pseudoleskea inciuxata [Mahler 1985]

37. Pseudoleskeella rupestris [Allred 2011]

38. Ptychostomum pallescens [Allred et al. 1998]

39. Ptychostomum pseudotriquetrum [Bartram 1931; Mahler
1978]

40. Ptychostomum turbinatum [Bartram 1931; Mahler 1978]

4 1 . Pylaisia selwynii [Allred 2011]

42. Rhizomnium magnifolium [Crum & Anderson 1981]

43. Rhizomnium pseudopunctatum [Mahler 1985]

44. Sciuro-hypnum plumosum [Mahler 1985]

45. Sciuro-hypnum populeum [Bartram 1931; Mahler 1978]

46. Scleropodium obtiisifolium [Allred 2012]

47. Sphagnum girgensohnii Russow [Romig & Allred 2013]

48. Stereophyllum radiculosum [Grout 1945; Mahler 1978]

49. Tortella fragilis (Flooker & Wilson) Limpricht [Romig &

Allred 2013]

50. Tortella tortuosa (Fledwig) Limpricht var. fragilifolia
(Juratzska) Limpricht [Romig & Allred 2013]

5 1 . Warnstorfia exannulata (Hedwig) Loeske [Romig & Allred
2013]

52. Warnstorfia fluitans [Price 2011]

IV. Not Recorded in BFNA27-2007. but Reported for New

Mexico Since the Publication of that Volume

1. Cn/w/o /art/o//a(Kindberg) W.B. Schofield [Kleinman

2012]

2. Distichium inclinatum (Hedwig) Bruch & Schimper [Allred

2012]

3. Dz7r/c/zMwgrac//e (Mitten) Kuntze [Allred 2011]

4. Grimmia americana BarXram [Allred 2011]

5. Grimmia trichophylla Grev. [Worthington 2011]

6. Microbryum starckeanum (Hedwig) R.H. Zander
var. fosbergii (Bartr.) Zander [Allred 2011]

7. Ptychomitriiim serratum Bruch & Schimper [Worthington

2011]

8. Rhabdoweisia crispata (Dickson ex Withering)

Lindberg [Allred 2011]

9. &/z/VZ(7zww r/wprert'/ (Theriot) W. A. Weber [Allred 2011]

10. Schistidium viride H.H. Blom & Darigo [Allred 2011]

11. Trematodon longicollisMichaux [Allred 2011]

1 2. Tuerckheimia svihlae (Bartram) Zander [Worthington 20 1 2]

13. Weissia muhlenbergiana (Swartz) W.D. Reese & B.A.E
Lemmon [Allred 2011]

Lists II and III comprise 79 taxa that had been reported for
New Mexico in the literature, nearly all with validating vouchers,
but were not noted in Flora North America as occurring in the
state. In a few cases, the identifications in the reports may have
been in error; in most cases, the BFNA authors were apparently
unaware of the published reports, and did not examine specimens
from New Mexico herbaria nor inquire with New Mexico
bryologists. Some of these discrepancies, however, can serve as
alerts for species needing verification.

The last official accounting of the mosses of New Mexico
was 314 years ago (Allred 2011). Below is presented a current
catalog of the mosses of the state, complete with synonyms,
index, and literature citations. The number of families has
increased from 38 to 41, genera from 130 to 139, species from
302 to 343, and total taxa from 312 to 353.

The catalog of mosses, as well as other pertinent information
about the bryophytes of New Mexico, may be found at
floraneomexicana.org.

I am pleased to acknowledge the frequent and agreeable
assistance of Karen Blisard, Russ Kleinman, and Kirsten Romig
in finding and studying the mosses of New Mexico.

(Continued on page 3)

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MOSSES OF NEW MEXICO -
FAMILY LIST

Total families = 41
Total genera =139
Total species = 343
Total taxa = 353

Family AMBLYSTEGIACEAE

Amblystegium

flmiatHe (Hedwig) Bruch, Schimper, & Guinbel = Hygroamblystegium
varium (Hedwig) Mdnkemeyer

humile (Beauvois) Bruch, Schimper, & Gumbel = Hygroamblystegium
varium (Hedwig) Monkemeyer var. humile (Beauvois)
Vanderpoorten & Hedenas

jiiralzkanum = Amblystegium serpens (Hedwig) Bruch & Schimper
noierophi lum (Su]]]vant & Lesquereux) Holzinger= Hygroamblystegium
varium (Hedwig) Mdnkemeyer

ripanum (Hedwig) Schimper = Leptodictyum riparium (Hedwig) Wamstorf
serpens (Hedwig) Bruch & Schimper vax juratzkammi (Schimper) Rau &
Hervey = Amblystegium serpens (Hedwig) Bruch &
Schimper

serpens (Fledwig) Bruch & Schimper [Bartram 1931, Mahler 1978]
Amblystegium juratzkanum Schimper

Amblystegium serpens (Hedwig) Bruch & Schimper var. juratzkanum
(Schimper) Rau & Hervey

tenax (Hedwig) C, Jensen = Hygroamblystegium varium (Hedwig)
Mdnkemeyer

vartum (Hedwig) Lindberg = Hygroamblystegium varium (Hedwig)
Mdnkemeyer

Calliergon stramineum (Bridel) Kindberg = Straminergon stramineum (Bridel)
Hedenas

Campyliadelphus

chrysophyUuS (Bridel) Kanda [Bartram I93l; Mahler 1978]

Campylium chrysophyUum (Bridel) J Lange

Campyliiim

chrysophyUum (Bridel) J Lange = Campyliadelphus chrysophyllus (Bridel)
Kanda

htspidulum (Bridel) Mitten = Campylophyllum hispidulum (Brid.) Hedenas
radicale{P Beauvois) Grout = Pseudocampylium radicale (P.

Beauvois) Vanderpoorten & Hedenas
Stellatllin (Hedwig) C. Jensen [Conard & Redfeam 1979; Lawton 1971]

Campylophyllum

hispidulum (Bridel) Hedenas [Bartram 1931; Mahler 1978]

Campylium hispidulum (Bridel) Mitten

Conardia

compacta (Muller Hal.) Robinson [Mahler 1978]

Rhynchostegiella compacta (Muller Hal.) Loeske

Cratoneuron

commutatum of American reports = Palustriella falcata (Bridel) Hedenas
deciptens of American reports = Palustriella talcata (Bridel) Hedenas
flHcinum (Hedwig) Spruce [Bartram 1931; Mahler 1978]

Drepanocladus

aduncus (Hedwig) Wamstorf [Bartram 1931; Maliler 1978; Wynne 1944]
Drepanocladus aduncus (Hedwig) Wamstorf var kneiffti (Schimper)
Moenkemeyer

Drepanocladus aduncus (Hedwig) Wamstorf wax. polycarpus
(Blandow ex Voit) Roth

longifolius (Mitten) Brotherus ex Paris [Allred 2012]
arfuticiM (Hedwig) Wamstorf var. kneiffti (Schimper) Moenkemeyer =
Drepanocladus aduncus (Hedwig) Wamstorf
aduncus (Hedwig) Wamstorf wax . polycarpus (Blandow ex Voit) Roth =
Drepanocladus aduncus (Hedwig) Wamstorf
sendtneri (Schimper) Wamstorf : this species not yet known from North
America [BFNA28]; reported by various authors, our
material is referred to Drepanocladus aduncus (Hedwig)

Wamstorf;

Drepanocladus sendtneri (Schimper) Wamstorf var. wilsonii
fuitans (Hedwig) Wamstorf = Warnstorfia fluitans (Hedwig) Loeske
sendtneri var. xvilsonii= Drepanocladus sendtneri Wamstorf
uncinatus (Hedwig) Wamstorf = Sanionia uncinata (Hedwig) Loeske
Hygroamblystegium

fluviatile (Hedwig) Loeske = Hygroamblystegium varium (Hedwig)
Mdnkemeyer

orthcladon (P. Beauvois) Macoun & Kindberg = Hygroamblystegium
varium (Hedwig) Monkemeyer

noterophilum (Sullivant & Lesquereux in Sullivant) Wamstorf =

Hygroamblystegium varium (Hedwig) Monkemeyer
Holzinger

tenax (Hedwig) Jennings = Hygroamblystegium varium (Hedwig)
Monkemeyer

varium (Hedwig) Mdnkemeyer var. humile (Beauvois)

Vanderpoorten & Hedenas [Vanderpoorten 2014]
Amblystegium humile (Beauvois) Bruch, Schimper, & Gumbel
Leptodictyum humile (P Beauvois) Ochyra
varium (Hedwig) Mdnkemeyer var. varium [Bartram 1931; Mahler
1978]

Amblystegium fluviatile (Hedwig) Bruch, Schimper, & Gumbel
Amblystegium varium (Hedwig) Lindberg
Hygroamblystegium noterophilum (Sullivant & Lesquereux)
Wamstorf

Hygroamblystegium orthocladon {? . Beauvois) Macoun & Kindberg
Hygroamblystegium tenax (Hedwig) Jennings
Leptodictyum trichopodium (Schulz) Wamstorf
luridum (Hedwig) Jennings wax. julaceum (Schimper ) Podpera =
Hygrohypnum luridum (Hedwig) Jennings

Hygrohypnum

bestii (Renauld & Bryhn) Holzinger [Jamieson 2014]

Hypnum molle (Hedwig) Loeske subsp bestii Renauld & Bryhn
duriusculum (DeNotaris) D.W. Jamieson [Jamieson 2014]

Limnobium duriusculum DeNotaris
luridum (Hedwig) Jennings [Bartram 1931; Mahler 1978]

Hygrohypnum luridum (Hedwig) Jennings wax. julaceum (Schimper )
Podpera

molle (Hedwig) Loeske [Bartram 1931; Mahler 1978] Note: This species is
not included for NM in BFNA28, and has been confused with H.
bestii and H. duriusculum; warrants verification.
ochraceum (Turner ex Wilson) Loeske [Mahler 1985]

Leptodictyum

humile (P Beauvois) Ochyra = Hygroamblystegium varium (Hedwig)
Monkemeyer var humile (Beauvois) Vanderpoorten &
Hedenas

riparium (Hedwig) Wamstorf [Poor 1966]

Amblystegium riparium (Hedwig) Schimper
trichopodium (Schulz) Wamstorf= Hygroamblysteium varium (Hedwig)
Monkemeyer

Limnobium duriusculum DeNotaris = Hygrohypnum duriusculum (DeNotaris)
D.W. Jamieson

Palustriella

falcata (Bridel) Hedenas [Allred etal 1998]

Cratoneuron commutatum of North American reports
Cratoneuron deciptens of North American reports
Palustriella commutata of North American reports
Palustriella deciptens of North American reports
commutata of North American reports = Palustriella falcata (Bridel)
Hedenas

deciptens of North American reports = Palustriella falcata (Bridel) Hedenas
Platydictya

jungermannioides (Bridel) Crum [Mahler 1978]
subtilis (Hedwig) Crum [Allred 2012]

Pseudocampylium

radicale {?. Beauvois) Vanderpoorten & Hedenas [Mahler 1985]
Campylium radicale (P Beauvois) Grout

Sanionia

(Continued on page 4)

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uncinata (Hedvvig) Loeske [Bartram I931; Maliler 1978; Wynne 1944]
Drepanocladus tincinalus (.Hedw]g) Warnstorf
Straminergon : go to Calliergonaceae
Tomentypnum

nitens (Hedvvig) Loeske [Cmm & Anderson 1981]

Warnstorfia : go to Calliergonaceae

Family ANDREAEACEAE

Andreaea

heinemannii Hampe & Muller Hal. [Mahler 1985]

Andreaea rupesiris sensu Mahler ( 1 985)

Family ANOMQDONTACEAE

Anomodon

attenuatUS (Hedwig) Hiibener [Bartram 1 93 1; Mahler 1978]
w//IOf (Hedvvig) Fiirnrohr [Ireland et al. 1984]
rostratus (Hedwig) Schimper [Bartram 1931, Mahler 1978]

Family AULACOMNIACEAE

Auhicomniiim

palustre (\\QdW\g) Schwiigrichen [Bartram 1931; Mahler 1978]

Aidacomnium palustre (Hedwig) Schwagrichen var. Unbricatum
Bruch & Schimper

palustre (Hedwig) Schwagrichen var. Unbricatum Bruch & Schimper =
Aulacomnium palustre (Hedwig) Schwagrichen

Family BARTRAMIACEAE

AnacoUa

laevisphaera (Taylor) Flowers [Mahler 1978]

AnacoUa subsessilis (Taylor) Brotherus
menziesii (Turner) Paris [Weber et al 2003]

subsesstUs (Taylor) Brotherus = Anacolia laevisphaera (Taylor) Flowers
Bartrama

hrevifolia Bridel [Allred et al 1998;]
ithyphylla Bridel [Bartram 1 931; Mahler 1978]
potosica Montagne [Allred et al 1998]
stricta Bridel [Gnffm ill, D.G, 2014]

Philonotis

CCtpillaris \-,\ndhQVg [Mahler 1978] Note; According to BFNA28-2014, this

species is associated with an oceanic climate; warrants verification.

fontana (Hedwig) Bridel var. americana (Dismier) Flowers ex
Crum [Flowers 1973]

fontana (Hedvvig) Bridel var. caespitosa (Juratzka) Schimper [Stark
& Castetter 1982]

fontana (Hedwig) Bridel var. fontana [Bartram 1931; Mahler 1978]
fontana (Hedvvig) Bridel var. pumila (Turner) Bridel [Bartram 1931;
Mahler 1978]

Philonotis tomentella Molendo
niarcllica (Hedvvig) Bridel [Bartram 1931; Mahler 1978]
tomentella Molendo = Philonotis fontana (Hedwig) Bridel var, pumtia
(Turner) Bridel

Family BRACHYTHECIACEAE
Brachyth eciastrum

collinum (Schleicher ex Muller Hal.) Ignatov & Huttunen [Bartram
1931; Maliler 1978]

Brachythecium collinum (Schleicher ex Muller Hal.) Schimper
Hypnum collinum Schleicher ex Muller Hal.
fendleri (Sullivant) Ochyra & Zarnovviec [Bartram 1931; Mahler 1978]
Brachythecium fendleri (Sullivant) Jaeger
Brachythecium utahense James
Leskea fendleri Sullivant

Vc/M/mwOT (Hedwig) Ignatov & Huttunen [Bartram I931; Mahler 1978]
Brachytheciastrum velutinum (Hedwig) Ignatov & Huttunen var.

salicinum (Schimper) Ochyra & Zarnowiec
Brachythecium petrophilumRS. Williams
Brachythecium siiberythrorrhizon Renauld & Cardot

Brachythecium velutinum (Hedwig) Bruch & Schimper
Brachythecium velutinum (Hedwig) Bruch & Schimper var venustum
(De Notaris) Arcangeli
Hypnum velutinum Hedwtg

velutinum (Hedwig) Ignatov & Huttunen var. salicinum (Schimper) Ochyra
& Zarnowiec = Brachytheciastrum velutinum (Hedwig)
Ignatov & Huttunen

Brachythecium

acuminatum (Hedwig) Austin [Bartram 1931; Mahler 1978]

Leskea acuminata Hedwig

albicans (Hedwig) Schimper [Stark & Castetter 1982]

Hypnum albicans Hedwig

collinum (Schleicher ex Muller Hal.) Schimper = Brachytheciastrum

collinum (Schleicher ex Muller Hal ) Ignatov & Huttunen
digastrum Muller Hal ex Kmdberg = Brachythecium laetum (Bridel)
Schimper

erythrorrhizon Schimper var. erythrorrhizon [Bartram 1931, Mahler
1978]

fendleri (Sullivant) Jaeger = Brachytheciastrum fendleri (Sullivant) Ochyra
& Zarnowiec

laetum (Bridel) Schimper [Worthington 1996]

Brachythecium digastrum Muller Hal ex Kindberg
Brachythecium oxydadon (Rr\de\) Jaeger
Hypnum laetum Bridel

latifolium Kindberg = Sciuro-hypnum latifolium (Kindberg) Ignatov &
Huttunen

nelsonii Grout = Sciuro-hypnum latifolium (Kindberg) Ignatov & Huttunen
oedipodium (Mitten) Jaeger = Sciuro-hypnum oedipodium (Mitten) Ignatov
& Huttunen

oxycladon (Bridel) Jaeger = Brachythecium laetum (Bridel) Schimper /n
BS.G,

petrophilum R S. Williams = Brachytheciastrum velutinum (Hedwig)
Ignatov & Huttunen

plumosum (Hedwig) Bruch & Schimper = Sciuro-hypnum plumosum
(Hedwig) Ignatov & Huttunen

populeum (Hedwig) Bruch & Schimper = Sciuro-hypnum populeum
(Hedwig) Ignatov & Huttunen
rivulare Bruch & Schimper [Bartram I931; Mahler 1978]
rofaea/TM/M DeNotaris [Allred 2012]
ruderale (Qr'\de\) W.R. Buck [Grout 1926; Mahler 1978]

Brachythecium stereopoma (Spruce ex Mitten) Jaeger
Brachythesium wootonii Grout
Hypnum ruderale Bridel

salebrosum (Weber & Mohr) Bruch & Schimper [Bartram I93i;
Lawton 1971; Mahler 1978; 1985]

Hypnum salebrosum Weber & Mohr
stereopoma (Spmce ex Mitten) Jaeger = Brachythecium ruderale (Bridel)

W R, Buck

suberythrorrhizon Renauld & Cardot = Brachytheciastrum velutinum
(Hedwig) Ignatov & Huttunen
utahense James = Brachythecium fenderi (Sullivant) Jaeger
velutinum (Hedwig) Bruch & Schimper = Brachytheciastrum velutinum
(Hedwig) Ignatov & Huttunen

wootonii Grout = Brachythecium ruderale (Bridel) W R Buck
Eurhynchiastrum

pulcltellum (WedWxg) Ignatov & Huttunen var. pulchellum [Bartram
1931; Mahler 1978]

Eurhynchium diversifolium Schimper
Eurhynchium pulchellum (Hedwig) Jennings
Hypnum pulchellum Hedwig
Rhynchostegium pulchellum (Hedwig) Robinson
Eurhynchium diversifolium Schimper = Eurhynchiastmm pulchellum (Hedwig)
Ignatov & Huttunen

hians (Hedwig) Sande Lacouture = Oxyrrhynchium hians (Hedwig) Loeske
pulchellum (Hedwig) Jennings = Eurhynchiastrum pulchellum (Hedwig)
Ignatov & Huttunen

praelongum (Hedwig) Schimper = Kindbergia praelonga (Hedwig) Ochyra
Kindbergia praelonga (Hedwig) Ochyra : Reported tentatively by Worthington
(1996), but not by BFNA28, awaits verification

(Continued on page 5)

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(Continued from page 4)

Eurhynchiiim praetongum (WtAwig) Schimper

Oxyrrhy n ch i u m

Ilians (Hedwig) Loeske [Allred 2012; Conard & Redfeam 1979; Grout 1928-
40]

Eurhynchiiim hians (Hedwig) Sande Lacouture
Plaiyhypnidium aquaticum (A Jaeger) M. Fleischer= Rhynchostegium
aquaticum A. Jaeger

Rhynchostegiella compacia (Muller Hal ) Loeske = Conardia compacta (Muller
Hal ) Robinson

Rhynchostegium

aquaticum A. Jaeger [Allred etal 1998]

Plaiyhypnidium aquaticum (A. Jaeger) M. Fleischer
Rhynchostegium riparioides of American authors
piikhellum (Hedwig) Robinson = Eurhynchiastrum pulchellum (Hedwig)
Ignatov & Huttunen

riparioides of American authors = Rhynchostegium aquaticum A Jaeger
serrulatum {Hedwig) Jaeger & Sauerbeck [Stark & Castetter 1982]
Hypnum serrulatum Hedwig
Sleerecleus serrulatus (Hedwig) Robinson
Sciuro-hypnum

latifolium (Kindberg) Ignatov & Huttunen [Allred et al 1998]
Brachyiheciiim latifolium Kindberg
Brachyihecium nelsonii Grout

oedipodium (Mitten) Ignatov & Huttunen [Allred et al 1998]
Brachyihecium oedipodium (Mitten) Jaeger
plumosum (Hedwig) Ignatov & Huttunen [Mahler 1985]
Brachyihecium plumosum (Hedwig) Bruch & Schimper
populeum (Hedwig) Ignatov & Huttunen [Bartram 1931; Mahler 1978]
Brachyihecium populeum (Hedwig) Bruch & Schimper
Scleropodium

cespitans (Muller Hal ) Koch : Specimens from NM identified as this have
been corrected to Brachyihecium.
obtusifolium (MiUen) Kindberg [Allred 2012]

Steerecleus serrulatus (Hedwig) Robinson = Rhynchostegium serrulatum
(Hedwig) Jaeger & Sauerbeck

Family BRUCHIACEAE

Trematodon

longicollis Michaux [Allred 20i i]

Family BRYACEAE

Anomobryum

julaceum (Bridel) Schimper [Allred 2012]

Brachymenium

vmosulum Cardot [Spence 2007] = Plagiobryoides vinosula (Cardot) J R.
Spence

klotzschii (Schwagrichen) Paris : Reported by Conard & Redfeam (1979)
but this species is found in Central and South America.
macrocarpum Cardot [Spence 2014]

systylium (Muller Hal.) Jaeger = Leptostomopsis systylia (Muller Hal.) J.R.
Spence

Bryum

affine Lindberg & Amell = Bryum creberrimum Taylor
algovicum Sendtner ex Muller Hal. = Ptychostomum pendulum Homschuch
andicola Hooker = Rosulabryum andicola (Hooker) Ochyra
argenteum Hedwig var. argenteum [Banram 1 931; Mahler 1978]
argenteum Hedwig var. muticum Bridel [Spence 2014]
bicolor Dickson = Bryum dichotomum Hedwig
billarderi Schwagrichen = Rosulabryum andicola (Hooker) Ochyra
bolanderi Sulhvant = Pohlia bolanderi (Sullivant) Brothems
caespiticium Hedwig = Gemmabryum caespiticium (Hedwig) J.R Spence
capillare Hedwig = Rosulabryum capillare (Hedwig) J.R Spence
creberrimum Taylor = Ptychostomum creberrimum (Taylor) J R. Spence &

H P Ramsay

cyclophyllum (Schwagrichen) Bruch & Schimper = Ptychostomum
cyclophyllum (Schwagrichen) J.R. Spence
demaretianum Arts = Gemmabryum demaretinaum (Arts) J R, Spence
dichotomum Hedwig = Gemmabryum dichotomum (Hedwig) Spence &

Ramsay

drummondii Muller Hal. = Pohlia drummondii (Muller Hal ) Andrews
exile Dozy & Molkenboer = Gemmabryum exile (Dozy & Molkenboer)

J R. Spence & H P Ramsay

gemmiparum De Notaris = Imbribryum gemmiparum (De Notaris) J R
Spence

incrassatoUmbatum Cardot = Plagiobryoides incrassatolimbata (Cardot)
J.R. Spence

kunzei Homschuch = Gemmabryum kunzei (Homschuch) J R. Spence
laevifilum Syed = Rosulabryum laevifilum (Syed) Ochyra
lanatum(P. Beauvois) Bridel [Allred 20ii]
lisae De Notaris var. cuspidatum (Bmch ex Schimper) Margadant =
Ptychostomum creberrimum (Taylor) J R. Spence &

H P.Ramsay

lonchocaulon Muller Hal. = Ptychostomum lonchocaulon (Muller Hal ) J R
Spence

miniatum Lesquereux = Imbribryum miniatum (Lesquereux) J R Spence
muehlenbeckii Bmch & Schimper = Imbribryum muehlenbeckii (Bruch &
Schimper) Pedersen

pollens (Bridel) Swartz = Ptychostomum pallens (Bridel) J.R Spence
pallescens Schleicher ex Schwagrichen = Ptychostomum pallescens
(Schleicher ex Schwagrichen) J R. Spence
pendulum (Homschuch) Schimper = Ptychostomum pendulum Homschuch
pseudotriquetrum (Hedwig) Gaertner et al. = Ptychostomum
pseudotriquetrum (Hedwig) J R Spence
pyriferum Cmndwell & Whitehouse = Bryum valparaisense Theriot
systylium Muller Hal. = Leptostomopsis systylia (Muller Hal ) J R Spence
tortifolium Funck in Bridel = Bryum cyclophyllum (Schwagrichen) Bruch
& Schimper

turbinatum (Hedwig) Turner = Ptychostomum turbinatum (Hedwig) J R.
Spence

uliginosum (Bridel) Bmch & Schimper = Ptychostomum cemuum
Homschuch

valparaisense Theriot = Gemmabryum valparaisense (Theriot) J R. Spence
Gemmabryum

caespiticium {Hedwig) ] R. Spence [Allred etal. 1998]

Bryum caespiticium Hedwig
demaretianum (Axis) J R. Spence [Spence 2014]

Bryum demaretianum Arts

dichotomum (Hedwig) Spence & Ramsay [Allred etal. 1998]

Bryum bicolor Dickson
Bryum dichotomum Hedwig

exile (Dozy & Molkenboer) J.R. Spence & H.P Ramsay [Spence
2014]

Bryum exile Dozy & Molkenboer
AMrtzd (Homschuch) J.R. Spence [Spence 20i4]

Bryum kunzei Homschuch

valparaisense (IherioJ) ] R. Spence [Ireland etal 1981. 1984]

Bryum pyriferum Crundwell & Whitehouse
Bryum valparaisense Theriot

Imbribryum

gemmiparum (De Notaris) J.R. Spence [Ireland et al. 1981]

Bryum gemmiparum De Notaris
miniatum (Lesquereux) J.R. Spence [Stark & Castetter 1982]

Bryum miniatum Lesquereux

muehlenbeckii (Bruch & Schimper) Pedersen [Ireland et al I98i]
Bryum muehlenbeckii Bruch & Schimper
Leptobryum go to Meesiaceae
Leptostomopsis

systylia (Muller Hal.) J.R. Spence [Mahler 1978]

Brachymenium systylium (Miiller Hal.) Jaeger
Bryum systylium Muller Hal
Plagiobryoides

incrassatolimbata (Cardot) J.R. Spence [Dango & Allred 200i]

Bryum incrassatoUmbatum Cardot
vinosula (Cardot) J.R. Spence [Spence 2007]

Brachymenium vinosula Cardo
Pohlia : go to Mielichhoferiaceae

(Continued on page 6)

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(Continued from page 5)

Ptychostomum

Hornschuch [Mahler 1978]

Bryuni idiginosum (Brldel) Bruch & Schimper
cre6rfr/-mH//n (Taylor) J.R. Spence & H.P. Ramsay [Banram 1931,
Mahler 1978]

Bryum affine Lindberg & Arnell
Bryum creberrimwn Taylor

Bryum lisae De Notaris var. ciispidalum (Bruch ex Schimper )
Margadant

cyclophyllitm (Schwagrichen) J.R. Spence [Stark & Castetter 1982]
Bryum cyclophyllum (Schwagrichen) Bruch & Schimper
Bryum tortifolium Funck in Bridel
lonchocaulon (Miiller Hal.) J.R. Spence [Mahler 1978]

Bryum lonchocaulon Muller Hal
pallens (Bridel) J.R. Spence [Allred et al. 1998]

Bryum pallens (Bridel) Swartz

pallescens (Schleicher ex Schwagrichen) J.R. Spence [Allred et al
1998]

Bryum pallescens Schleicher ex Schwagrichen
pendulum Hornschuch [Bartram 1 931; Mahler 1978]

Bryum pendulum (Hornschuch) Schimper
Bryum algovicum Sendtner ex Muller Hal.
pseudotriquetrum (Hedvvig) J.R. Spence [Bartram 1931; Mahler 1978]
Bryum pseudotriquetrum (Hedvvig) Gaertner et al.
tU rhi natu m (Hedwig) J.R. Spence [Bartram 1931, Mahler 1978]

Bryum turbinatum (Hedwig) Turner
Rhodobryum

ontariense (VAndhetg) Kindberg [Mahler 1985]
roseum (Hedwig) Limpricht : Reported by Stark & Castetter ( 1 986), but
identification corrected to Rhodobryum ontariense
(Kindberg) Kindberg

Rosulabryum

andicola (Hooker) Ochyra [Mohamed 1979]

Bryum billarderi Schwagrichen
CapiUare (Hedwig) J.R. Spence [Bartram 193l; Mahler 1978]

Biyum capiUare Hedwig
laevifilum (Syed) Ochyra [Allred 20i i]

Bryum laevifilum Syed

Webera nutans Hedwig = Pohlia nutans (Hedwig) Lindberg
Family BRYOXIPHIACEAE

Bryoxiphium

/mn’c^/cMW (Bridel) Mitten [Allred etal 1998]

Family CALLIERGONACEAE

Straminergon

stramineum (Bridel) Hedenas [Flowers 1973]

Calliergon stramineum (Bridel) Kindberg

Warnstorfia

exannulata (Hedvyig) Loeske [Romig & Allred 20i3]
fluitans (Hedwig) Loeske [Price 20ii]

Drepanocladus fluitans (Hedwig) Wamstorf
Hypnum fluitans Hedwig

Family CLIMACIACEAE

CHmacium

americanum Bridel [Mahler 1985]

dendroides (Hedwig) Weber & Mohr [Bartram 193 1; Mahler 1978]
Family DICRANACEAE

Brothera

leana (Sulliyant) Muller Hal. [Kleinman et al. 2011]

Dichodontium

pellucidum (Hedwig) Schimper [Allred et al. 1998]

Dicranella

var/aJHedwig) Schimper [Ireland etal 1981]

Dicranoweisia

cirratafWedwig) Lindberg ex Milde [Mahler 1978]
crispula (Hedwig) Lindberg ex Milde [Banram 1931; Mahler 1978]
Dicranum

acutifolium (Lindberg & Arnell) Jensen (Ireland 2007]
brevifolium (Lindberg) Lindberg [Ireland 2007]
fuscescensfnmeryzx.fuscescens [Allred etal 1998]
montanum Hedwig [Banram 1931; Mahler 1978]
muehlenbeckii Bruch & Schimper [Banram I93i; Mahler 1978]
rhabdocarpum Sulliyant [Banram 1931; Mahler 1978]
scoparium Hedwig [Allred etal 1998]
tauricum Sapjegin [Romig & Allred 20 1 3]

Oncophorus

Virens (Hedwig) Bridel [Banram 1931; Mahler 1978]
wahlenbergii Bridel [Allred et al. 1998]

Paraleucobryum

enerve (Thedenius) Loeske [Banram I93 1; Mahler 1978]
longifolium (Hedwig) Loeske [Banram 1 931; Mahler 1978]
Rhabdoweisia

crispata (Diekson ex Withering) Lindberg [Allred 20i i]
Symblepharis

helicophylla Montagne = Symblepharis yaginata (Hooker) Wijk &
Margadant

vaginata (Hooker) Wijk & Margadant [Ireland et al. 1984, Mahler 1978]
Symblepharis helicophylla Montagne

Family DITRICHACEAE

Ceratodon

purpureus (Hedwig) Bridel suhsp. purpureus [Banram 193 1; Mahler
1978]

purpureus (Hedwig) Bridel subsp. stenocarpus (Bruch &
Schimper) Dixon [Darigo& Allred 2001]
stenocarpus Bruch & Schimper = Ceratodon purpureus (Hedwig) Bridel
subsp. stenocarpus (Bruch & Schimper) Dixon

Distichium

capillaceum (Hedwig) Bruch & Schimper [Banram 1931, Mahler 1978]
inclinatum (Hedwig) Bruch & Schimper [Allred 2012]

Ditrichum

/Zex/caH/e (Schwagrichen) Hampe [Allred etal. 1998]
gfflc/7e (Mitten) Kuntze [Allred 20ii]

Saelania

glaUCeSCensfHedwig) BrotheruS [Banram 1931; Mahler 1978]
Didymodon trifarius (Hedwig) Rohling

Family ENCALYPTACF.AF.

Encalypta

ciliata Hedwig [Banram 1931; Mahler 1978]
rhaptocarpa Se\\w'agric\\er\ [Conard & Redfeam 1979]
vulgaris Hedwig [Chamberlain 1925; Banram 1931]

Family ENTODONTACEAE

Entodon

beyrichii Schwagrichen [Buck 20i4]

cladorrhizans (Hedwig) Muller Hal : Reported by Mahler (1978), but

identification corrected to Entodon schleicheri (Schimper)
Demeter [Darigo & Allred 200 1 ]
concinnus (De Notaris) Paris [Banram I93i; Mahler 1978]

Entodon orthocarpus (Bridel) Lindberg
orthocarpus (Bridel) Lindberg = Entodon concinnus (De Notaris) Paris
schleicheri (Schimper) Demeter [Buck & Crum 1978]
seductrLx (Hedwig) Muller Hal. [Blisard & Kleinman 2012]
sullivantii (Muller Hal.) Lindberg [Cmm & Anderson 1981]

Family FABRONIACFAF

(Continued on page 7)

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(Continued from page 6)

Fabronia

ciliaris (Bridel) Bridel [Bartram 1931; Mahler 1978]

Fabronia ciliaris (Bridel) Bridel var wrightii (Sullivant) Buck
Fabronia gymnostoma Sullivant & Lesquereux ex Sullivant
Fabronia octoblepharis (Schleicher) Schwagrichen
Fabronia wrightii Sullivant

ciliaris (Bridel) Bridel var. wrightii (Sullivant) Buck = Fabronia ciliaris
(Bridel) Bridel

gymnostoma Sullivant & Lesquereux ex Sullivant = Fabronia ciliaris
(Bridel) Bridel

octoblepharis (Schleicher) Schwagrichen = Fabronia ciliaris (Bridel) Bridel

pusilla Raddi [Mahler 1978]

wrightii Sullivant = Fabronia ciliaris (Bridel) Bridel

Family FISSIDENTACEAE

Fissidens

bryoidss Fledwig [Grout 1928-1940; Mahler 1978; Stark & Castetter 1982]
Fissidens bryoides Hedwig var. pusillus (Wilson) Pursell
Fissidens bryoides Hedwig var. viridulus (Swartz) Brotherus
Fissidens exigitus Sullivant forma emarginatus Grout
Fissidens viridulus (Swartz) Wahlenberg
bryoides Hedwig var pusillus (Wilson) Pursell = Fissidens bryoides
Hedwig

bryoides Hedwig var viridulus (Swartz) Brotherus = Fissidens bryoides
Hedwig

crispus Montagne [Darigo & Allred 2001; Maliler 1978]
exiguus Sullivant forma emarginatus Grout = Fissidens bryoides Hedwig
var. bryoides

fonlanus Steudel [Weber 1973]

limbatus Sullivant ; Reported by Mahler (1978), but our material is referred
to Fissidens crispus Montagne [Darigo & Allred 2001]
liltlei (Williams) Grout [Allred 1998; Williams 1936]

Moenkemeyera /;«/e( Williams
obtusifolius fNWson var. acuminatus GxoviX [Eckel 1986]
obtusifolius Wilson var. apiculatus Grout ; Reported by Stark & Castetter
(1982), but identification corrected to Fissidens
sublimbatus Grout by Ron Pursell. [Darigo & Allred
2001]

sublimbatus Grout [Pursell 2007)

viridulus (Swartz) Wahlenberg = Fissidens bryoides Hedwig
Moenkemeyera littlei Williams = Fissidens littlei (Williams) Grout

Family FONTINALACEAE

Fontinalis

aiUipyretica Hedwig [Mahler 1978; Welch 1963]
hypnoides HarXman [Allred etal 1998]

neomexicana Sullivant & Lesquereux [Bartram 1931; Mahler 1978;
Sullivant & Lesquereux 1856]

Family FUNARIACEAE

Entosthodon

attenuatus (DiGsSon) Bryhn [Miller & Miller 2007]
rubiginosus (Williams) Grout [Ireland et at 1984]

Funaria

apiculatopilosa Cardot [Worthington 1996]

Funaria orcuttii Bartram

calvescens Schwagrichen = Funaria hygrometrica Hedwig var. calvescens
(Schwagrichen) Montagne
flavicans Michaux [Miller* Miller 2007]

hygrometrica Hedwig var. calvescens (Schwagrichen) Montagne

[Darigo & Allred 2001]

Funaria calvescens Schwagrichen

hygrometrica Hedwig var. hygrometrica [Bartram I93l, Mahler 1978]
Funaria hygrometrica Hedwig var. utahensis Grout
hygrometrica Hedwig var. utahensis Grout = Funaria hygrometrica
muhlenbergii Turner [Stark & Castetter 1982]
orcuttii Bartram = Funaria apiculatopilosa Cardot
Physcomitrium

p>’f//£>r/Mef_(Hedwig) Hampe [Ireland etal 1984]
Pyramidula

tetragona (Bridel) Bridel [Goffmet 2007]

Family GIGASPERMACEAE

Lorentziella

imbricata (Mitten) Brotherus [Allred et at 1998]

Family GRIMMIACEAE

Coscinodon

calyptratus (Hooker) C. Jensen ex Kindberg [Bartram 1931; Hastings
1999; Mahler 1978]

Grimmia calyptrata Hooker

cribrosus (Hedwig) Spruce [Hastings 1999; Ireland et at 1984]

Grimmia cribrosa Hedwig

wrightii Sullivant in Gray = Jaffueliobryum wrightii (Sullivant in Gray)
Theriot

Grimmia

affinis Hoppe & Homschuch ex Homschuch = Grimmia longirostris
Hooker

alpestris (Weber & Mohr) Schleicher [Hastings & Greven 2007]

Grimmia ungeri of New Mexico reports
alpicola of American authors = Schistidium agassizii Sullivant &
Lesquereux

alpicola Swartz ex Hedwig var. rivularis = Schistidium rivulare (Bridel)
Podpera var rivulare

ambigua (Sullivant) Sullivant = Schistidium ambiguum Sullivant
americana Bartram [Allred 20 1 1 ]
anodon Bruch & Schimper [Bartram I931; Mahler 1978]
apocarpa Hedwig var. ambigua (Sullivant) G, Jones in Grout = Schistidium
ambiguum Sullivant

apocarpa Hedwig var. apocarpa = Schistidium apocarpum (Hedwig) Bruch
& Schimper

apocarpa Hedwig var. conferta (Funck) Sprengel = Schistidium confertum
(Funck) Bruch & Schimper

apocarpa Hedwig var. stricta = Schistidium apocarpum (Hedwig) Bruch &
Schimper

arizonae Renault & Cardot [Hastings & Greven 2007; Weber et al. 2003]
bemoullii Mueller Hal. : Reported by Weber et al (2003), but duplicate
specimens were considered to be Grimmia ovalis
(Hedwig) Lindberg (Hastings & Greven 2007), awaits
verification.

calyptrata Hooker = Coscinodon calyptratus (Hooker) C Jensen ex
Kindberg

catalinensis Bartram = Grimmia longirostris Hooker
converta Funck var. obtusifolla Schimper = Schistidium apocarpum
(Hedwig) Bruch & Schimper

cribrosa Hedwig = Coscinodon cribrosus (Hedwig) Spruce
crinitoleucophaea Cardot [Hastings & Greven 2007]

Grimmia poecilostoma Cardot & Sebille
elatior Bruch ex Balsamo-Crivelli & De Notaris [Dango & Allred
2001]

donniana Smith : Reported by Mahler (1978), but material re-identified as
Grimmia longirostris Hooker [Darigo & Allred 2001]
laevigata (Bridel) Bridel [Mahler 1978]
longirostris Hooker [Dango & Allred 200 1]

Grimmia affinis Homschuch
Grimmia catalinensis Bartram
montana Bruch & Schimper [Darigo & Allred 200 1]
orbicularisfQruch ex Wilson [Mahler 1978]
ovalis (Hedwig) Lindberg [Bartram 1931; Mahler 1978]
pilifera Beauvois [Darigo & Allred 200 1 ]
plagiopodiaWcdWlg [Mahler 1978]

poecilostoma Cardot & Sebille = Grimmia crinitoleucophaea Cardot
pulvinata (Hedwig) Smith [Mahler 1978]
raid Austin = Jaffueliobryum raui Theriot
sessitana De Notaris [Hastings & Greven 2007]

(Continued on page 8)

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Grimmia tenerrima Renault & Cardot
tenenima Renault & Cardot = Grimmia sessitana De Notaris
fexica/ia Greven [Greven20i0]
trichopliylla Grev. [Worthington 2011]

ungeri of New Mexico reports = Grimmia alpestris (Weber & Mohr)
Schleicher

wrightii (Sullivant in Gray) Austin = Jaffueliobryum wrightii (Sullivant in
Gray) Theriot

Jaffueliobryum

raw/ (Austin) Theriot [Ireland et at 1981]

Grimmia rant Austin

wrightii (Sullivant in Gray) Theriot [Bartram I93i, Maliler 1978]
Coscinodon wrightii Sullivant in Gray
Grimmia wnghtti (Sullivant in Gray) Austin
Schistidium

alpicola of Amer. authors = Schistidium rivulare (Bridel) Podpera
ambigUUm Sullivant [Sullivant 1849] Note: The type is a Fendler specimen
from Santa Fe; essentially ignored by bryologists since its
publication; possibly conspecific with Schistidium confertum
(Funck) Bruch & Schimper
Grimmia ambigua (Sullivant) Sullivant

Grimmia apocarpa Hedwig var ambigua (Sullivant) G. Jones in
Grout

apocarpum (Hedwig) Bruch & Schimper : reported by Bartram (1 931) and
Mahler ( 1 978), but this species occurs in norther U S. and
Canada (McIntosh 2007), our material is perhaps referred
to Schistidium ambiguum Sullivant or Schistidium
confertum (Funck) Bruch & Schimper
Grimmia apocarpa Hedwig
Grimmia apocarpa Hedwig var. stricta
Grtmmina converta Funck var obtusifolia Schimper
flfro/MycMAM (Schimper) Limpricht [McIntosh 2007]
confertum (Funck) Bruch & Schimper : not reported for NM, but material
previously referred to Schistidium apocarpum (Hedwig)
Bruch & Schimper may belong to this species; awaits
verification

Grimmia apocarpa Hedwig var. conferta (Funck) Sprengel
dupretii (Theriot) W.A. Weber [Allred 20i i]
frigidum H.H. Blom [McIntosh 2007]

rivulare (Bridel) Podpera var. latifoltum (Zetterstedt) Crum & Anderson =
Schistidium rivulare (Bridel) Podpera
rivulare (Bridel) Podpera [Bartram I93I; Mahler 1978]

Grimmia alpicola of Amer. authors
Grimmia alpicola Swartz ex Hedwig var. rivularis (Bridel)
Wahlenberg

Schistidium alpicola of Amer authors

Schistidium rivulare (Bridel) Podpera var latifoltum (Zetterstedt)
Crum & Anderson

viride H.H, Blom & Darigo [Allred 20i i]

Family HEDWIGIACEAE

Braunia

secunda (Hooker) Bruch, Schimper, & Gumbel [Allred etaf 1998]
Hedwigia

ciliata (Hedwig) P. Beauvois [Bailram 193l; Mahler 1978]

Family HYLOCOMIACEAE

Hylocomium

splendens (Hedwig) Schimper [Bartram 1931; Mahler 1978]

Pleurozium

schreberi (Bridel) Mitten [Mahler 1985]

Family HYPNACEAE

Breidleria pratensis (Koch ex Spruce) Loeske = Hypnum pratense (Rabenh.)
Spruce

arcuata (Lindberg) Loeske = Hypnum lindbergii Mitten
Calliergonella lindbergii (Mitten) Hedanas = Hypnum lindbergii Mitten
HomomalUum

adnatum (Hedwig) Brotherus : Plants previously thought to be

Homomallium adnatum have all been verified by Bruce
Allen (MO) as Homomallium mexicanum Cardot
incurvatum (Schrader ex Bridel) Loeske [Allred 20I2]
mexicanum Cardot var. latifolium Cardot [Bartram I93i; Mahler 1978]
mexicanum Cardot var. mexicanum [Bartram I93i; Mahler 1978]
Hypnum

albicans Hedwig = Brachythecium albicans (Hedwig) Schimper
coUinum Schleicher ex Muller Hal = Brachytheciastrum colhnum
(Schleicher ex Muller Hal ) Ignatov & Huttunen
cupressiforme Hedwig var. subjulaceum Molendo [Bartram 1 93 1 ;
BFNA28; Little 1937]

fluilans Hedwig = Warnstorfia tluitans (Hedwig) Loeske
lindbergii Mitten [Bartram 1931; Mahler 1978]

Breidleria arcuata (Lindberg) Loeske
Calliergonella lindbergii (MiP&n) Hedenas
molle (Hedwig) Loeske subsp bestii Renauld & Bryhn = Hygrohypnum
bestii (Renauld & Bryhn) Holzinger
pallescens (Hedwig) P. Beauvois [Stark & Castetter 1982]
pratense (Rdihenh.) Spruce [Allred etaf 1998]

Breidleria pratensis (Koch ex Spruce) Loeske
pidchellum Hedwig = Eurhynchiastrum pulchellum (Hedwig) Ignatov &
Huttunen

revolutum (Mitten) Lindberg [Bartram 1 93 1; Mahler 1978]
ruderale Bridel = Brachythecium ruderale (Bridel) W R. Buck
salebrosum Weber & Mohr = Brachythecium salebrosum (Weber & Mohr)
Schimper

serrulatiim Hedwig = Rhynchostegium serrulatum (Hedwig) Jaeger &
Sauerbeck

vaucheri Lesquereux [Ireland et al. 1981]

velutinum = Brachytheciastrum velutinum (Hedwig) Ignatov & Huttunen
wahlenbergii F. Weber & Mohr = Pohlia wahlenbergii (F Weber & Mohr)
Andrews

Isopterygiopsis

pulcliella (Hedwig) Iwatsuki [Allred et al. 1998]

Isopterygium pulchellum (Hedwig) Jaeger
Plagiolhecium pulchellum (Hedwig) Schimper
Isopterygium

homomalltfolium Redfeam = Pseudotaxiphyllum homomallifolium
(Redfearn) Ireland

pulchellum (Hedwig) Jaeger = Isopterygiopsis pulchella (Hedwig) Iwatsuki
tenerum Mitten [Allred et at 1998]

Platygyrium

fuscoluteum Cardot [Mahler 1978; Reese 1999]
repens (Bridel) Schimper [Allred 20i2]

Pseudotaxiphyllum

homomallifolium {KeAtearn) \re\nnA [Ireland etaf 1984]

Isopterygium homomallifolium Redfearn

P}’laisia

polyantha (Hedwig) Schimper [Bartram 193l; Mahler 1978]

Pylaisiella polyantha (Hedwig) Grout
selwynii (Kindberg) Crum, Steere, & L.E. Anderson [Allred 20ii]
Pylaisiella polyantha (Hedwig) Grout = Pylaisia polyantha (Hedwig) Schimper
Taxiphyllum

deplanatum (Bruch & Schimper ex Sullivant) Fleischer [Bartram
1931; Mahler 1978]

taxirameum (Mitten) Fleischer [Mahler 1978]

Family LFSKF.ACF.AF.

Bryohaplocladium

angustifolium (Hampe & Muller Hal.) Watson & Iwatsuki = Haplocladium
angustifolium (Hampe & Muller Hal ) Brotherus
microphyllum (Hedwig) Watson & Iwatsuki = Haplocladium microphyllum
(Hedwig) Brotherus

Claopodium

pellucinerve (Mitten) Best [Mahler 1978]

Haplocladium

(Continued on page 9)

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angiistifolium (Hampe & Muller Hal.) Brotherus [Allred et al 1998]
Bryohaplocladhim angiistifolium (Hampe & Muller Hal.) Watson &
Iwatsuki

microphyllum (Hedwig) Brotherus [Bartram 1931, Mahler 1978; Stark &
Castetter 1986]

Biyoliaplocladium microphyllum (Hedwig) Watson & Iwatsuki
Leptopterigynandrum

austroalpinumMviWQr Hal. [Allred etal 1998]

Lescuraea

arizonae (Williams) P Wilson & Norris = Pseudoleskeella arizonae
(Williams) Lawton

incurvata (Hedwig) Lawton = Pseudoleskea incurvata (Hedwig) Loeske
saxicola (Schimper) Milde [Mahler 1985]

Leskea

acuminata Hedwig = Brachythecium acuminatum (Hedwig) Austin
angUStataTay\0T [Worthington 2012]

fendleri Sullivant = Brachytheciastrum fendleri (Sullivant) Ochyra &
Zarnowiec

Leskeella

nervosa (Bridel) Loeske [Stark & Castetter 1986]

Pseudoleskeella nervosa (Bridel) Nyholm
nen’osa (Bridel) Loeske var. sibirica (Arnell) Brotherus = Pseudoleskeella
rupestris (Berggren) Hedenas & Soderstrom
williamsll Best = Pseudoleskeella tectorum (Funck ex Bridel) Kindberg
Lindbergia

brachyptera (Mitten) Kindberg [Bartram 1931; Mahler 1978]
mexicana (Bescherelle) Cardot ex Pringle [Bartram 1949]
Pseudoleskea

incurvata (Hedwig) Loeske [Mahler 1985]

Lescuraea incun’ata (Hedwig) Lawton
radicosa (Mitten) Macoun & Kindberg var. radicosa [Worthington
1996]

Pseudoleskeella

rtWzOA/fle (Williams) Lawton [Bartram 1931; Maliler 1978]

Lescuraea arizonae (Williams) P Wilson & Norris
nervosa (Bridel) Nyholm = Leskeela nervosa (Bridel) Loeske var. nervosa
rupestris (Berggren) Hedenas & Soderstrom [Allred 201 1]

Leskeella nervosa (Bridel) Loeske var. sibirica (Amell) Brotherus
Pseudoleskeella sibirica (Arnell) P. Wilson & Norris
sibirica (Arnell) P Wilson & Norris = Pseudoleskeella rupestris (Berggren)
Hedenas & Soderstrom

tectorum var flagellifera (Best) Amann = Pseudoleskeella tectorum (Funck
ex Bridel) Kindberg

tectorum (Funck ex Bridel) Kindberg [Bartram 1931; Mahler 1978]
Leskeella williamsii Best

Pseudoleskeella tectorum var. flagellifera (Best) Amann
Family MEESIACEAE

Leptobryum

pyriforme (HedWig) Wilson [Bartram I931; Mahler 1978]

Family MIELICHHOFERIACEAE

Polilia

bolanderi (SuWivani) Brotherus [Shaw, A.J. 2014]

Bryum bolanderi Sullivant

cruda (Hedwig) Lindberg [Bartram 193 1; Mahler 1978]

Mnium crudum Hedwig

drummondii (Muller Hal.) Andrews [Stark & Castener 1982]

Bryum drummondii Muller Hal
elongata Hedwig [Allred etal. 1998]
nutans (Hedwig) Lindberg [Bartram 1931; Mahler 1978]

Webera nutans Hedwig

obtusifolia (Bridel) Koch [Worthington 2012]
tundrae A.i. Shaw [Allred 2012]

wahlenbergii (F. Weber & Mohr) Andrews [Ireland etal. 1981]

Hypnum wahlenbergii F Weber & Mohr

Family MNIACEAE

Mnium

affine Bland, ex Funck var. rugicum (Laur.) Bruch & Schimper =
Plagiomnium ellipticum (Bridel) T Koponen
arizonicum Amann [Bartram 193l; Mahler 1978]
blyttii Bruch & Schimper [Bartram I931; Mahler 1978]
crudum Hedwig = Pohlia cruda (Hedwig) Lindberg
marginatum (Dickson ex Withering) Bridel ex P. Beauvois
[Bartram 1 931; Mahler 1978]
titomsonii Schimper [Bartram 1931; Mahler 1978]

Plagiomnium

ciliare (Muller Hal.) Koponen [Allred 2012]
cuspidatum (Hedwig) Koponen [Bartram 1931; Mahler 1978]
ellipticum (Hridei) Koponen [Allred etal. 1998]

Mnium affine Bland, ex Funck var rugicum (Laur ) Bruch &
Schimper

medium (Bruch & Schimper) Koponen [Allred 20I2]
rostratum (Schrader) Koponen [Bartram I93i; Mahler 1978]

Pohlia : go to Mielichhoferiaceae

Rhizomnium

magnifolium (Horik.) T. Koponen [Crum & Anderson I98i , as Mnium
punctatum]

pseudopunctatum (Bruch & Schimper) Koponen [Mahler 1985]
punctatum (Hedwig) T. Koponen [Allred et at 1998]

Family NECKERACRAF,

Metaneckera menziesii (Drummond) Steere = Neckera menziesii Drummond

Neckera

menziesii Drummond [Cardot 19 lO; Mahler 1978]

Metaneckera menziesii (Drummond) Steere
Porotrichum neomexicanum (Cardot) Wagner
pennata Hedwig [Bartram 1931; Mahler 1978]

Neckera pennata Hedwig var tenera Muller Hal.
pennata Hedwig var. tenera Muller Hal. = Neckera pennata Hedwig
Porotrichum neomexicanum (Cardot) Wagner = Neckera menziesii Drummond

Family ORTHOTRICHACFAF

Orthotrichum

alpestre Hornschuch ex Bruch & Schimper [Mahler 1978]
anomalum Hedwig [Bartram 1931; Mahler 1978]
cupulatum Hridei [Worthington 2012]
diaphanum Hridei [Mahler 1978]

liallii Sullivant & Lesquereux in Sullivant [Bartram 1931; Mahler 1978]

laevigatumZeXtersiedi [Allred etal. 1998]

obtusifolium Qridei [Mahler 1978]

pollens Bridel [Romig& Allred 2013]

pellucidum Lindberg [Worthington 2012]

pumilum Swartz [Mahler 1978]

pycnophy Hum Schimper [Vitt 1973]

rupestre Schleicher ex Schwagrichen [Bartram I93i; Mahler 1978]
tenellum Bruch ex Bridel : Reported by Mahler (1978), but this species
occurs in the West Coast states and Nevada, and is not
likely in NM; our material most likely referred to
Orthotrichum pumilum Swartz.

Zygodon

viridissimus (Dickson) Bridel var. rupestris Lindberg ex Hartman

[Bartram 1931; Mahler 1978]

Zygodon viridissimus (Dickson) Bridel var. vulgaris Malta
viridissimus (Dickson) Bridel var. vulgaris Malta = Zygodon viridissimus
(Dickson) Bridel var. ruprestris Lindberg ex Hartman

Family PLAGIOTHECIACEAE

Plagiotliecium

denticulatum (Hedwig) Schimper [Ireland et al. 1981]

(Continued on page 10)

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(Continued from page 9)

/ae/HW Schimper [Mahler 1978]

pulcheUutn (lledwig) Scliimper = Isopterygiopsis pulchella(Hedwig)
Iwatsuki

Family POLYTRICHACEAE

Atrichum

rupestns Hedwig : Reported by Mahler (1985), but not included in North
America by Smith-Merrill (2007); awaits verification,
selwynii Austin [Mahler 1985; Smith-Merrill 2007]
teneliu m (RohVmg) Bruch & Scliimper [Romig& Allred 2013]
undulatum (Hedwig) P Beauvois : Reported by Bartram (1931) and

Mahler (1978), but scarcely in North America fide Smith-
Merrill (2007), specimens await correct determination.

Meiotrichum

lyallii (Mitten) G.L. Smith [Conard & Redfeam 1979; Smith-Memll 2007]
Polytrichiim lyallii (Mitten) Kindberg

Pogonatum alpinum (lledwig) RhI. = Polytrichastrum alpinum (Hedwig) G.L.
Smith

Polytri c has! rum

alpinum (Hedwig) G.L. Smith var. alpinum [Bartram 1931; Mahler
1985]

Pogonatum alpinum (Hedwig) Rhl
ohioense (Renauld & Cardot) G.L. Smith [Smith-Memll 2007]
sexangulare (Bridel) G.L. Smith [Mahler 1978]

Polytrichiim sexangulare Bridel

Polytriclium

juniperinum Hedwig [Bartram 193 1; Mahler 1978]
lyalliifMitten) Kindberg = Meiotrichum lyallii (Mitten) G.L. Smith
piliferum Hedwig [Mahler 1985; Smith-Memll 2007]
sexangulare Bridel = Polytrichastrum sexangulare (Bridel) G L. Smith

Family POTTIACEAE

Aloina

rigida (Hedwig) Limprieht yar. rigida [Delgadillo-M. 2007; Mahler 1978]

Anoeciangium obiusifolium (Brotherus & Paris in Cardot) Grout = Molendoa
sendtneriana (Bruch & Schimper ) Limprieht

Astomum phascoides (Drumm ) Grout = Weissia phascopsis Zander

Barbula

acuta (Bridel) Bridel = Didymodon rigidulus Hedwig var. gracilis
(Schleicher ex Hooker & Greville) Zander
acuta (Bridel) Bridel) var bescherellei (Sauerbeck ex Jaeger) Crum =

Didymodon rigidulus Hedwig var. gracilis (Schleicher ex
Hooker & Greville) Zander

bescherellei Sauerbeck = Didymodon rigidulus Hedwig var gracilis
(Schleicher ex Hooker & Greville) Zander
bolleana (Miiller Hal.) Brotherus [Eckel 1988; Zander 2007]

Barbula ehrenbergii (Lorentz) Fleischer
convoluta Hedwig var. convoluta [Zander 2007]
ehrenbergii (Lorentz) Fleischer = Barbula bolleana (Muller Hal.) Brotherus
gracilis Schwagnehen = Didymodon rigidulus Hedwig var. gracilis
(Schleicher ex Hooker & Greville) Zander
indica (Hooker) Sprengel var. indica [Zander 2007]
re/7e.ica (Bridel) Bridel = Didymodon ferrugineus (Schimper ex Bescherelle)
M.O Hill

sinensis Muller Hal. = Syntrichia sinensis (Muller Hal ) Ochyra
spiralis (Schimper) ex Muller Hal. = Pseudocrossidium replicatum (Taylor)
Zander

unguiculata Hedwig [Bartram 1931; Mahler 1978]

Bryoerythrophyllum

recurvirostrum (Hedwig) Chen [Bartram 1931; Maliler 1978]

Bryoerythrophyllum recurvirostrum (Hedwig) Chen var. dentatum
(Schimper) Crum et al,

Didymodon recurvirostrum (Hedwig) Jennings
Weissia recurvirostra Hedwig

recurvirostrum (Hedwig) Chen var. dentatum (Schimper) Crum et al. =
Bryoerythrophyllum recurvirostmm (Hedwig) Chen

Chenia

leptophylla (MiiWex Had.) Z-andet [Ireland etal 1984]

Chenia rhizophylla Sakurai) Zander
Tortula rhizophylla (Sakurai) Iwatsuki & Saito
Crossidium

aberrans HoMitger & Bartram [Ireland etal I98i]
crassinervium (De Notaris) Juratzka var. crassinervium [Stark &
Castetter 1986]

seriatum Crum & Steere [Delgadillo-M. 1996]

Crumia

latifolia (Kindberg) W.B. Schofield [Kleinman 2012]

Desmatodon cernuus (Huberner)Bmch & Schimper = Tortula cernua (Hubemer)
Lmdberg

convolulus (Bridel) Grout = Tortula atrovirens (J.E. Smith) Lmdberg
guepinii Bruch & Schimper = Tortula guepmii (Bmch & Schimper)
Brotherus

heimii (Hedwig) Mitten = Hennediella heimii (Hedwig) Zander
latifolius (Hedwig) Bridel = Tortula euryphylla Zander
latifolius (Hedwig) Bridel var muticus (Bridel) Bridel = Tortula hoppeana
(Schultz) Ochyra

obtusifolius (Schwagrichen) Schimper = Tortula obtusifolia (Schwagrichen)
Mathey

plinihobius Sull & Lesquereux = Tortula plmthobia (Sullivant &
Lesquereux) Austin

Didymodon

anserinocapitatus (X.-j. Li) Zander (Zander* Weber 1997]
acutus (Bridel) Saito = Didymodon rigidulus Hedwig
australasiae (Greville & Hooker) Zander [Stark & Castetter 1986; Ireland
et al. 1984]

Husnotiella torquescens (Cardot) Bartram
Trichoslomopsis australasiae (Greville & Hooker) Robinson
australasiae (Hooker & Greville) Zander var. umbrosus (Muller Hal )
Zander = Didymodon umbrosus (Muller Hal ) Zander
bracityphyllus (Sullivant) Zander [Zander & Ochyra 200 1 ]

Didymodon luridus of NM reports
fallax (Hedwig) Zander [Zander 2007]

fallax (Hedwig) Zander var reflexus (Bridel) Zander = Didymodon
fermgineus (Schimper ex Bescherelle) M O. Hill
/e/TM^t/ieMS (Schimper ex Bescherelle) M.O. Hill [Bartram I93i;
Mahler 1978]

Barbula reflexa (Bridel) Bridel

Didymodon fallax (Hedwig) Zander var. reflexus (Bridel) Zander
luridus of NM reports = Didymodon brachyphyllus (Sullivant) Zander
nevadensis Zander [Zander 2007]

recurvirostrum (Hedwig) Jennings = Bryoerythrophyllum recurvirostrum
(Hedwig) Chen

revolutus (Cardot) Williams [Stark & Castetter 1982]
rigidulus Hedwig var. gracilis (Schleicher ex Hooker & Greville)
Zander [Bartram 1931; Mahler 1978; Stark & Castetter 1986]
Barbula acuta Bridel

Barbula acuta (Bridel) Bridel) var. bescherellei (Sauerbeck ex Jaeger)
Crum

Barbula bescherellei Sauerbeck
Didymodon acutus (Bridel) Saito

rigidulus Hedwig var. icmadopitilus (Muller Hal.) Zander [Allred et
al. 1998]

rigidulus Hedwig var. rigidulus [Bartram I93i]
tectorum (Muller Hal.) Saito [Zander & Ochyra 2001]
topliaceus (Bridel) Lisa [Bartram 1 93 1; Mahler 1978]
trifarius (Hedwig) Rhl. = Saelania glaucescens (Hedwig) Brotherus
(Ditrichaceae)

umbrosus (Muller Hal.) Zander [Zander 2007]

Didymodon australasiae (Hooker & Greville) R.H. Zander var
(Muller Hal ) R H Zander

vinealis (Bridel) Zander var. luridus of NM reports = Didymodon
brachyphyllus (Sullivant) Zander
vinealis (Bridel) Zander var. vinealis [Stark & Castetter 1982]
Eucladium

verticillatum (Bridel) Bruch & Schimper [Bartram I93i; Mahler 1978]
Weissia verticillata Bridel

(Continued on page II)

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Gymnostomum

aeruginosum Smith [Banram 1931; Mahler 1978]

Gymnostomum calcareum of NM reports
calcareum of NM reports = Gymnostomum aeruginosum Smith
Hennediella

heimii (Hedwig) Zander var. heimii [Bartram 1931; Mahler 1978]
Desmatodon /le/wn (Hedweg) Mitten
Pottia heimii (Hedwig) Hampe

Husnotiella torquescens (Cardot) Bartram = Didymodon australasiae(Greville &
Hooker) Zander

Hymenostomum tortile (Schwagrichen) Bruch & Schimper = Weissia condensa
(Voit) Lmdberg

Hymenostylium

recurvirostrum (Hedwig) Dixon var. recurvirostrum [Banram 1931;
Mahler 1978]

Gymnostomum recurvirostrum Hedwig

Leptodontium

y/exv/oZ/Mw (Withering) Hampe in Lindberg [Mahler 1978]
Microbryum

davallianum (Smith) R.H. Zander var. conicum (Schwagrichen) R.
H. Zander [Reported tentatively by Worthington 1996]

Pottia texana Wareham

starckeanum (Hedwig) R.H. Zander var. fosbergii (Bartr.) Zander
[Allred 2011]

starkeanum (Hedwig) R.H. Zander var. starkeanum [Ireland et at
1981]

Pottia anzonica Wareham var mucronulata Wareham

Molendoa

sendtneriana (Bruch & Schimper ) Limpricht [Ireland et at 1981; Stark
& Castetter 1982]

Anoectangium obtusifolium (Brotherus & Paris in Cardot) Grout
Neocardotia subnigra (Mitten) Theriot & Bartram = Rhexophyllum subnigrum
(Mitten) Theriot ex Hilpert

Oxystegus tenuirostris (Hooker & Taylor) A J E Smith = Trichostomum
tenmrostre (Hooker & Taylor) Lindberg
Phascum cuspidatum Hedwig = Tortula acaulon (Withering) Zander
Pleurochaete

luteola (Bescherelle) Theriot [Mahler 1978]

Pleurochaete squarrosa of New World reports
squarrosa of New World reports = Pleurochaete luteola (Bescherelle)
Tlieriot

Pottia arizonica Wareham var mucronulata Wareham = Microbryum

starkeanum (Hedw ) R H Zander [Ireland et at 1981]
bryoides (Dickson) Mitten = Tortula protobryoides R H. Zander
heimii (Hedwig) Hampe = Hennediella heimii (Hedwig) Zander
nevadensis Cardot & Tlieriot = Tortula nevadensis (Cardot & Theriot) R.H
Zander

texana Wareham = Microbryum davallianum (Smith) R H Zander var.

conicum (Schwagrichen) R. H. Zander [Reported
tentatively by Worthington 1996]

Pseudocrossidium

crinitum (Schultz) Zander [Mahler 1978]

Pseudocrossidium aureum (Bartram) Zander
Tortula aiirea Bartram

aureum (Bartram) Zander = Pseudocrossidium crinitum (Schultz) Zander

obtusulum (Lindberg) Crum [Zander 2007]

repUcatum (Taylor) Zander [Mahler 1978, as Baibula spiralis]

Barbula spiralis (Schimper) ex Muller Hal.

Pterygoneurum

cavifolium (Ehrh ) Juratzka = Pterygoneurum ovatum (Hedwig) Dixon
ovatum (Hedwig) Dixon [Mahler 1978]

Pterygoneurum cavifolium (Ehrh.) Juratzka
subsessiie (Bridel) Juratzka var. subsessile [Ireland et al. 1981]
Rhexophyllum

subnigrum (Mitten) Theriot ex Hilpert [Banram 1931; Mahler 1978]
Neocardotia subnigra (Mitten) Theriot & Bartram

Stegonia

lafifolia (Schwagrichen) Venturi ex Brotherus var. latifolia [Zander

2007]

Syntrichia

bartramii (Steere) Zander [Mahler 1978]

Tortula bartramii Steere in Grout
caninervis Mitten [Flowers 1973, as Tortula bistratosa Flowers]

Tortula bistratosa Flowers
Tortula caninervis (Mitten) Brotherus
chisoa (Magill, Delgadillo, & Stark) Zander [Magill et al 1983; Stark &
Castetter 1986]

Tortula chisoa Magill, Delgadillo, & Stark
fragilis (Taylor) Ochyra [Bartram 1 93 1; Mahler 1978]

Tortula fragilis Taylor

inermis (Bridel) Bmch = Tortula inermis (Bridel) Montagne
laevipila Bridel [Bartram 1 93 1; Mahler 1978]

Tortula pagorum (Milde) De Notaris
montanaNees [Allred etal. 1998; Mishler2007]

Tortula intermedia (Bridel) Berkeley
Tortula montana (Nees) Lindberg

norvegica Weber & Mohr [Allred et al 1998; Flowers 1973]

Tortula norvegica (Weber & Mohr) Wahlenberg ex Lindberg
obtusissima (Muller Hal.) Zander [Ireland et al I98i]

Tortula obtusissima (Muller Hal ) Mitten
pagorum (Milde) Amann = Syntrichia laevipila Bridel
papillosa (Wilson) Juratzka [Weber 1957]

Tortula papillosa Wilson

papillosissima (Coppey) Loeske [Flowers 1973]

Tortula papillosissima (Coppey) Brothems
Tortula ruralis (Hedwig) Gaertner var hirsuta (Venturi) Paris
ruralis (Hedwig) Weber & Mohr [Bartram 1931; Mahler 1978]

Tortula ruralis (Hedwig) Gaertner

Tortula ruralis (Hedwig) Gaertner et al. var. crinita De Notaris
sinensis (Muller Hal.) Ochyra [Mishler 2007]

Barbula sinensis Muller Hal.

Timmiella

anomala (Bruch & Schimper ) Limpricht [Stark & Castetter 1986]
Tortella

fragilis (Hooker & Wilson) Limpricht [Romig & Allred 2013]
/jMW/7/S.(Hed wig) Jennings [Ireland et at 1981]
tortuosa (Hedwig) Limpricht var. fragilifolia [Romig & Allred 2013]
tortuosa (Hedwig) Limpricht var. tortuosa [Stark & Castetter 1982]
Tortula

aurea Bartram = Pseudocrossidium crinitum (Schultz) Zander
acaulon (Withering) Zander [Ireland et at I981; Zander* Eckel 2007]
Phascum cuspidatum Hedwig

atrovirens (J.E. Smith) Lindberg [Stark & Castetter 1986]

Desmatodon convolutus (Bridel) Grout
bartramii Steere in Grout = Syntrichia bartramii (Steere) Zander
bistratosa Flowers = Syntriehia caninervis Mitten
Am’/pes (Lesquereux) Brotherus [Allred etal. 1998]
caninervis (Mitten) Brotherus = Syntrichia caninervis Mitten
CerrtMa_(Huberner) Lindberg [Crum & Anderson I981]

Desmatodon cernuus (Hb.)Bruch & Schimper
chisoa Magill, Delgadillo, & Stark = Syntrichia chisoa (Magill, Delgadillo,
& Stark) Zander

etiryphylla Zander = Tortula hoppeana (Schultz) Ochyra
fragilis Taylor = Syntrichia fragilis (Taylor) Ochyra
guepinii (Bruch & Schimper) Brotherus [Mahler 1978]

Desmatodon guepinii Bmch & Schimper
hoppeana (Schultz) Ochyra [Mahler 1978; Zander & Eckel 2007]
Desmatodon latifolius (Hedwig) Bridel

Desmatodon latifolius (Hedwig) Bridel var muticus (Bridel) Bridel
Tortula euryphylla Zander
inermis (Bridel) Montagne [Mahler 1978]

Syntrichia inermis (Bridel) Bmch
intermedia (Bridel) Berkeley = Syntrichia montana Nees
montana (Nees) Lindberg = Syntrichia montana Nees
mucronifolia Schwagrichen [Bartram 1931; Mahler 1978]

(Continued on page 12)

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nevadensis (Cardot & Theriot) R.H. Zander [Ireland et al. 1984]

Potna nevadensis Cardot & Theriot
norvegica (Weber & Mohr) Wahlenberg ex Lindberg = Syntrichia
norvegica Weber & Mohr

obtusifolia (Schwagrichen) Mathieu [Bartram 193I; Mahler 1978]
Desmatodon (Schwagrichen) Schimper

obtusissima (Muller Hal.) Mitten = Syntrichia obtusissima (Milller Hal.)
Zander

pagonim (MWde) De Notaris = Syntrichia laevipila Bridel
papillosa Wilson = Syntrichia papillosa (Wilson) Juratzka
papillosissima (Coppey) Brotheais = Syntrichia papillosissima (Coppey)
Loeske

Tortula niralis (Hedwig) Gaertner var. hirsuta (Venturi) Paris
plinthobia (Sullivant & Lesquereux) Austin [Mahler 1978]
Desmatodon plinthobiiis Sullivant & Lesquereux
protobryoides R.H. Zander [Allred et al. 1998]

Pottia bryoides (Dickson) Mitten

rhizophylla (Sakurai) Iwatsuki & Saito = Chenia leptophylla (Muller Hal.)
Zander

niralis (Hedwig) Gaertner et al. var. crinita De Notaris = Syntrichia ruralis
(Hedwig) Weber & Mohr

niralis (Hedwig) Gaertner var. hirsuta (Venturi) Paris = Syntrichia
papillosissima (Coppey) Loeske

ruralis (Hedwig) Gaertner= Syntrichia ruralis (Hedwig) Weber & Mohr
Tortula ntra/w (Hedwig) Gaertner et al. var. crinita De Notaris
Trichostomopsts australasiae (Greville & Hooker) Robinson = Didymodon
australasiae (Greville & Hooker) Zander

Trichostomum

Trichostomiim svihlae Bartram = Tuerckheimia svihlae (Bartram) Zander
brachydontium_^mch [Ireland et al. 1984]

Cmpu/H/M Bruch [Ireland etal, 1984]
planifolium (Dixon) R.H. Zander [Stark 1996]

Trichostomum swelii (Bartram) Stark
Weissia planifolia Dixon
Weissia sweetii Bartram

sweetii (Bartram) Stark = Trichostomum planifolium (Dixon) R.H. Zander
tenuirostre (Hooker & Taylor) Lindberg [Ireland et al. 1984]
Oxystegus temiirostris (Hooker & Taylor) A.J.E. Smith
Weissia tenuirostris Hooker & Taylor
Tuerckheimia

svihlae (Bartram) Zander [Worthington 2012]

Trichostomum svihlae Bartram

Weissia

andersoniana Zander = Weissia ligulifolia (Bartram) Grout
condensa (Voit) Lindberg ; Not known from North America [Zander
2007], reports of this species in NM refer to Weissia
controversa Hedwig

Hymenostomum tortile (Schwagrichen) Bruch & Schimper
Weissia tortilis (Schwagrichen) Muller Hal.
controversa Hedwig [Bartram 1931; Mahler 1978]

Weissia condensa auct., non (Voit) Lindberg
Weissia viridula Hedwig

glauca Bartram = Weissia ligulifolia (Bartram) Grout
ligulifolia (Bartram) Grout [Ireland et al. 1981; Stark & Castetter 1982]
Weissia andersoniana Zander
Weissia glauca Bartram

muhlenbergiana (Swartz) W.D. Reese & B.A.E Lemmon [Allred
2011]

phascopsis Zander [Eckel 1986]

Astomum phascoides (Drummond) Grout [not Weissia phascoides
(Wilson) Muller Hal ]

planifolia Dixon = Trichostomum planifolium (Dixon) R.H. Zander
recurvirostra Hedwig = Bryoerythrophyllum recurvirostmm (Hedwig)
Chen

sweetii Bartram = Trichostomum planifolium (Dixon) R.H. Zander
tenuirostris Hooker & Taylor = Trichostomum tenuirostre (Hooker &
Taylor) Lindberg

tortilis (Schwagrichen) Muller Hal. = Weissia condensa (Voit.) Lindberg
verticillata Bridel = Eucladium verticillatum (Bridel) Bruch & Schimper

viridula Hedwig = Weissia controversa Hedwig

Family PTYCHOMITRIACEAE

Ptychomitrium

serratum Bruch & Schimper [Worthington 20i i]
sinense (Mitten) A. Jaeger [Ireland et al. 1984; Reese 1998]
Ptychomitrium leibergii Best

leibergit Best = Ptychomitrium sinense (Mitten) A. Jaeger
Family RHYTIDIACEAE

Rhytidium

rugosum (Hedwig) Kindberg [Bartram 193 1; Mahler 1978]
Family SELIGERIACEAE

Blindia acuta (Hedwig) Bruch & Schimper : Allred 9880 was identified as this,
but the specimen is corrected to Hymenostylium
recurvirostrum (Hedwig) Dixon

Family SPHAGNACEAE

Sphagnum

fimbriatum Wilson [Coop 2002]
girgensohnii Russow [Romtg & Allred 2013]

Family SPLACHNACEAE

Tayloria

Hornschuch [Cmm & Anderson 1981]

Family STEREOPHYLLACEAE

Stereophyllum

radiculosum (Hooker) Mitten [Grout 1945; Mahler 1978]

Family TETRAPHIDACEAE

Tetraphis

pellucidaWeAwg [Allred etal. 1998]

Family THUIDIACEAE

Abietinella

abietina (Hedwig) Fleischer [Bartram 1931; Mahler 1978]

Thuidium abietinum (Hedwig) Schimper
Thuidium

abietinum (Hedwig) Schimper = Abietinella abietina (Hedwig) Fleischer
delicatulum (Hedwig) Schimper yar. radicans (Kindberg) Crum et
al. [Bartram 1931; Mahler 1978]

Thidium philibertii Limpricht

philibertii Limpricht = Thuidium delicatulum (Hedwig) Schimper var.
radicans (Kindberg) Crum et al.

Family TIMMIACEAE

Timmia

austriaca Hedwig [Bartram 1931; Mahler 1978]

bavarica Hessler = Timmia megapolitana Hedwig var. bavarica (Hessler)
Bridel

megapolitana Hedwig var. bavarica (Hessler) Bridel [Mahler 1978]
Timmia bavarica Hessler

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(Continued on page 13)

.>u (

Ik. V . Ml

\

■ fi..

(Continued from page 12)

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(Continued on page 14)

I. •/ \vVi -t "U”. '3|

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• •• • : -i '

/

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(Continued from page 13)

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"The New/ Mexico Botanist" is a digital newsletter to help
make information about the flora of New Mexico readily
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Submissions will be reviewed by the editor, who may ask

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Submissions may be sent to the editor at kallred(g)nmsu.edu.
You may be placed on the emailing list by sending your name
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is available online at floraneomexicana.org.

- Kelly Allred, editor

is the natuml science thnt trnnsmits the UnowledQe of plants.

— ^innneus

Lu£3fH£RT' MERi^

library

FEB 2 5 2015

new YORK
pnTANICAL GARDEN

Number 63

March 16,2015

A Review of Carex in New Mexico: Initial Findings

James McGrath

20 Robin Ct., Edgewood, NM 87015 sedgesf(/!s\\CD.coin

Max Licher

Box 1456, Sedona, Arizona 86339 in I icher('i;/ \vi ldapache.net
William R. Norris

Department of Natural Sciences, Western New Mexico University, P.O. Box 680,
1000 W. College Ave., Silver City, NM 88062 norris\v'r/!\vnnui.edii

Glenn Rink

Denver Herbarium, Biological Sciences Department, Northern Arizona University
P.O. Box 5640, Flagstaff, Arizona 8601 1-5640 faroutbotanv'frigma il.com

During January 8-10, 2014 four botanists convened at the
University of New Mexico Herbarium (UNM) to review New
Mexico plant specimens in the genus Carex. The purpose of
the review was 1) to annotate misidentified specimens, and 2)
to clarify the status of Carex species known to occur in the
state of New Mexico. This initial 3-day event launched
formation of the Southwest Carex Working Group
(SWCWG).

SOUTHWEST C/4/?£^X WORKING GROUP
(SWCWG)

The four authors of this paper are all experienced
botanists with diverse backgrounds who have over the years
developed a particular interest in sedge species in the genus
Carex while working in states other than New Mexico. James
(“Jim”) McGrath has been studying Carex species since his
days mapping wetlands in Yellowstone National Park in the
mid 1990's. McGrath has specialized in wetlands and has
discovered and documented the existence of six Carex species
previously unrecorded in New Mexico, originally reported
under Plant Distribution Reports in The New Mexico Botanist
newsletter, and later included in the Flora Neomexicana series
(Allred 2012). In 2013 he co-presented (with Bill Norris and
Russell Kleinman) a Carex identification workshop in New
Mexico sponsored by the Native Plant Society of New
Mexico. Glenn Rink and Max Licher are based at the Deaver
Herbarium at Northern Arizona University in Flagstaff They
have studied Carex extensively in Arizona and neighboring
states. Their studies have included reviewing the Carex
collections in 20 herbaria and examining nearly 6,000 Carex
specimens during the past four years. The culmination of their

efforts has resulted in a document entitled ""Carex of
Arizona” (Rink and Licher 2015 - currently submitted for
publication). This document includes a key to Arizona Carex
species, full descriptions of each species and discussions of
various taxonomic, identification, ecological, and habitat
issues pertaining to these species. William R. (“Bill”) Norris
has studied Carex extensively in Iowa, where he has
annotated more than 12,000 Carex specimens, prepared an
unpublished dichotomous key to Iowa's 120 Carex taxa, and
co-presented six Carex identification workshops. Norris is
currently a biology professor at Western New Mexico
University in Silver City. He is the author of the Carex key in
Flora Neomexicana III (Allred & Ivey 2012), a manual
dedicated to the identification of New Mexico flora.

Botanist Charles (“Chick”) Keller is also a member of
SWCWG, but he has not been as deeply involved in specimen
annotation. Like McGrath, Keller discovered several Carex
species previously unrecorded in the state of New Mexico and
he published these in The New Mexico Botanist under Plant
Distribution Reports. Keller is the founder of and primary
contributor to the Jemez Mountain Herbarium in Los Alamos,
New Mexico.

NEW MEXICO SPECIMENS REVIEWED AND

ANNOTATED AT SIX REGIONAL HERBARIA

Our 3-day event at the UNM herbarium was just the
beginning of our review of New Mexico Carex. During
2014 through January 2015, New Mexico Carex specimens
deposited in the Biology Herbarium (NMC) and the Range
Science Herbarium (NMCR) at New Mexico State

(Continued on page 2)

^otdnice' cst ^cicnti« atuvnUs ^egetabilium co^nitiorem trndit.

— ^innncus

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University, and at the University of Texas at El Paso
Herbarium (UTEP), were reviewed and annotated by Licher,
Rink and Norris. All four of us reviewed and annotated New
Mexico Carex specimens at San Juan College (SJNM) in
Farmington, New Mexico in early January 2015. Licher and
Rink reviewed a similar collection at the Brigham Young
University Herbarium (BRY) in Provo, Utah. Licher and
McGrath e.xtended our study of New Mexico Carex to the
field by making the first Southwest Carex Working Group's
sedge collecting trip to the Vermejo Park Ranch in Taos
County about 25 miles northeast of Questa, New Mexico in
late July 2014.

ANNOTATIONS - RULE OF THREE

During our review of UNM, NMC and SJNM specimens,
we adopted a Rule of Three regarding the annotation of
specimens. That is, we affix an annotation label listing our
four names to a given specimen sheet if three of us agree on a
determination. We decided that if three of us agreed on a
determination, then confirmation by a fourth reviewer was
unnecessary. This procedure evolved because sometimes it
was logistically difficult for all four of us to review every
single specimen.

GOALS

It is our intent to improve the understanding of the large
and complex genus Carex, which is the second most diverse
genus (after Astragalus) in the New Mexico vascular plant
flora. It is our intention that this paper will be the first of a
series of publications focused on New Mexico Carex taxa. In
this paper we first summarize our findings during reviews of
the UNM, NMC and SJNM collections based on databases
that we developed for those collections. Secondly, we identify
and discuss additions, deletions and nomenclature upgrades to
the New Mexico flora based on our review of all of the above-
described herbaria. In this discussion, we clarify the status of
several Carex taxa previously thought to exist in New Mexico,
but for which verification of their existence in this state is
needed.

Future publications will focus on specific sections of genus
Carex as described in Volume 23 of Flora of North America
(Ball and Reznicek 2002). Each publication will focus on a
particular section or sections of genus Carex. By focusing on a
few closely related species we can highlight the
morphological, ecological and habitat characteristics that
distinguish these species from one another as well as from
other species of Carex.

REVIEW OF UNM, NMC, AND SJNM COLLECTIONS
- A SUMMARY

UNM

We annotated 714 specimens at UNM representing 48
Carex taxa In this process, we determined 26 specimens
(3.6%) for which there was no specific epithet. Furthermore,
we annotated 17 specimens as Carex sp. because of their
immature and/or fragmentary condition that precluded their
accurate determination to species level. Among these 17

specimens were 15 that we downgraded from a previous
species determination. We determined that 51 of 671
specimens (7.6%) which had a previous determination were
misidentified. An additional 27 specimens are currently under
review by Norris and eight highly questionable specimens
were sent to North American Carex expert Anton A. Reznicek
(University of Michigan) for determination. The most
commonly misidentified specimens were Carex emoryi, C.
uebrascensis, C. petasata, C. praegracilis and C. siccata.

Finally, we set aside for later review all UNM specimens
representing taxa in Carex section Acrocystis (i.e., Carex
geophila, C. inops subsp. heliophila, C. pityophila, C. rossii)
and many specimens in Carex section Ovales because they
were too difficult for us to confidently determine in a short
time period.

NMC

The Southwest Carex Working Group reviewed all 159
Carex specimens in the NMC collection. We upgraded the
nomenclature of 24 specimens (15.1 %), and corrected the
identification of 39 (24.5%) misidentified specimens. The
most commonly misidentified species were Carex inops
subsp. heliophila, C. occidentalis, C. praegracilis, and C.
wootonii. We could not reach consensus on determinations of
1 1 specimens due to immature or poor quality specimens or to
taxa that are not clearly defined. Nor could we reach
agreement on the identity of two additional specimens, which
will be sent to A. A. Reznicek for determination.

SJNM

We annotated 166 Carex specimens representing 40 taxa
during our review of the Carex collection in the San Juan
College Herbarium. We provided nomenclature upgrades for
eight (4.8%) specimens and we decided that two specimens
were not identifiable and annotated them as Carex sp. We
determined that 63 of the remaining 164 specimens (38.4%)
were misidentified. The most commonly misidentified species
were C. microptera, C. occidentalis and C siccata. Less
commonly misidentified taxa included C. bella, C. douglasii,
C. diiriiiscida, C. inops subsp. heliophila and C. praegracilis.

ADDITIONS AND DELETIONS TO THE NEW
MEXICO FLORA

Carex kellogii W. Boott - Nomenclatural Upgrade

A recent publication by Dragon and Barrington (2009)
recognized Carex lenticiilaris Michaux var. lipocarpa (Holm)
L. Standley as Carex kelloggii W. Boott. We have accepted
this nomenclatural change. Dragon and Barrington (2009)
recognized other varieties of Carex lenticiilaris, but we have
encountered none of these in our review of Carex collections
in New Mexico. The treatment of Carex Section Phacocystis
by Standley et al. (2002) reports that the other varieties of C.
lenticiilaris occur in geographic regions (Pacific northwest.
Pacific coast states, northern U.S and Canada) far distant from
New Mexico.

A collections search on SEINet (2014), an on-line
searchable database of plant specimens housed in more than

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100 herbaria mostly in the southwestern United States,
revealed only eight Carex kelloggii specimens collected in
New Mexico. Our review of UNM specimens revealed several
specimens identified as what we now recognize as C kelloggii
to be other members of section Phacocystis, especially C
aquatilis Wahlenberg var. aquatilis, C. nebrascemis Dewey
and C. eniotyi Dewey. Carex kelloggii is cespitose, whereas
the latter two species are long-rhizomatous. C. kelloggii
normally has only one staminate spike, while C. aquatilis var.
aquatilis and C emoryi typically have more than one
staminate spike (Rink and Richer 2015; Standley et al. 2002).

Carex aureolensis Steudel (Verified as part of the New

Mexico flora) and

Carex frankii Kunth (Dropped from the New Mexico flora)

Several sedge specimens originally determined as Carex
frankii Kunth and collected in New Mexico are housed in five
western herbaria (UNM, NMC, BRY, UTEP and UCR -
University of California, Riverside). All of these specimens
have been reviewed by us except for two deposited at UCR.
However, the UCR specimens are duplicates of specimens that
we reviewed at other herbaria. Based on our review of these
specimens we conclude that all New Mexico specimens
originally determined as Carex frankii Kunth are, in fact,
Carex aureolensis Steudel. Hence, we conclude that Carex
frankii Kunth does not exist in New Mexico. Our conclusion is
consistent with the treatment of Carex section Squarrosae by
Ford and Reznicek (2002), who present the distribution of C.
frankii as extending no further west than Oklahoma, Kansas
and Nebraska and who show the distribution of C. aureolensis
to include the state of New Mexico. Allred (2012) has
recognized C. aureolensis as part of the New Mexico flora,
citing Ball and Reznicek (2002), which includes Ford and
Reznicek (2002).

Ford and Reznicek (2002) provided characteristics that
distinguish C. aureolensis from C. frankii. Carex aureolensis
is long-rhizomatous, while in contrast C. frankii is cespitose
and short-rhizomatous. C. aureolensis is also distinguished
from C frankii by slightly wider and translucent pistillate
scale bodies (0.4-0. 9 mm wide compared to 0. 1-0.4 mm wide)
and wider staminate scales. According to a SEINet (2014)
collections search and a review of UTEP Carex collections, C.
aureolensis has been found only at a few springs in southern
New Mexico: Sitting Bull Falls in the Guadalupe Mountains,
Oak Springs in Carlsbad Caverns National Park, both in Eddy
County, and at a spring in the Florida Mountains of Luna
County.

Carex stricta Lamarck - Dropped from the New Mexico

Flora

In their treatment of Carex section Phacocystis in Flora of
North America vol. 23, Standley et al. (2002) indicated at the
end of their description of Carex senta Boott that “Collections
from New Mexico identified as C. senta are C. strictaC They,
therefore, exclude New Mexico from the geographical range
of Carex senta Boott. However, in this same publication they
contradict the above statement by excluding New Mexico

from the geographic range for Carex stricta. Since the above
publication, Norris (2012a) verified the existence of C. senta
in New Mexico based on a specimen collected in Grant
County with determination verified by A. A. Reznicek. Later,
Norris (2012b) reported that Carex senta is “typically
encountered and fairly common along high energy streams and
rivers in the Gila National Forest...”

The presence of C. stricta in New Mexico has been an
unanswered question since the publication of Flora of North
America Vol. 23 in 2002. Norris (2012b) included C. stricta in
his treatment of New Mexico Carex, basing his decision on an
on-line collections search (SEINet, accessed late 2011) that
revealed specimens determined as C. stricta were “collected in
northern (border of small pond, Mora; in canyon. Union) and
southwestern (sandy gravel bar, Catron) counties in New
Mexico.” However, one or more members of SWCWG has
reviewed Carex specimens collected in New Mexico at UNM,
NMC, NMCR, SNM (Dale A. Zimmerman Herbarium at
Western New Mexico University), UTEP and BRY herbaria.
We have found no specimens identified as C. stricta nor any
specimens of C. stricta misidentified as other taxa.
Furthermore, a collections search on SEINet (2014) failed to
reveal any C. stricta specimens collected in New Mexico. In
fact, SEINet revealed only two C. stricta specimens collected
in the entire state of Texas. An examination of the distribution
of C. stricta on maps found on The PLANTS Database
(USDA, NRCS 2014) indicated that C. stricta is known no
further west than central Texas and western Kansas. C. stricta
is apparently unrecorded in Oklahoma (Standley et al. 2002;
USDA, NRCS 2014). Standley et al. (2002) reported that C.
stricta may be “the most common wetland sedge in eastern
North America,” but it is apparently very uncommon to absent
in the southern U.S.

McGrath searched the New Mexico Biodiversity
Collections Consortium website (NMBCC 2007) in late
December 2014 and found records of seven C. stricta
specimens that matched the habitat and location information
described by Norris (2012b) in his treatment of New Mexico
Carex. However, a search of the determination history of these
specimens revealed that five of them had subsequently been
annotated to C. aquatilis in 1998 by Miriam Fritts and we
agreed with the Fritts determinations during our early 2014
review of UNM Carex specimens. We annotated the other two
such C. stricta specimens to C. emoryi and C. lenticularis var.
lipocarpa (C. kelloggii) during our review of NMC specimens
in early 2014.

Based on the above evidence, we conclude that Carex
stricta does not now exist nor has it ever existed in New
Mexico. We base our conclusion on 1) no evidence of
specimens documenting the existence of this species in New
Mexico, 2) an apparent error by Standley et al. (2002)
resulting in contradictory information regarding the
distribution of C. stricta in New Mexico, and 3) the likelihood
that New Mexico is substantially outside the range of C.
stricta.

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Carex biixbaiimii Wahlenberg, Carex magelhmica Lamarck

subsp. irrigua (Wahlenberg) Hiitonen, and Carex

saxatilis Linnaeus - Additions to the New Mexico Flora

Specimens of all three of these taxa were collected by
Keller, Roy Greiner and McGrath in the San Pedro Parks
Wilderness in Rio Arriba County, New Mexico in mid-August,
2012. The identities of the Carex biabaumii Wahlenberg,
Carex magellanica Lamarck subsp. irrigua (Wahlenberg)
Hiitonen, and Carex saxatilis Linnaeus specimens were
confirmed by Norris and the identities of the C. biabawnii and
C. saxatilis specimens were also confirmed by Reznicek. All
three taxa are represented in the key to New Mexico Carex
developed by Norris (2012b) because they are among more
than 20 Carex taxa that have been collected in adjacent
counties in Arizona, Utah and/or Colorado, implying that they
might occur in New Mexico.

Chick Keller has for several years hypothesized that the
flora of the San Pedro Parks Wilderness is a relict left over
from the Ice Age. The San Pedro Parks Wilderness is at
approximately 10,000 feet elevation and contains numerous
springs and associated wetlands. In addition, two UNM
specimens collected from what we now recognize as the San
Pedro Parks Wilderness in the mid 1960's have been verified
by Reznicek as Carex macloviana d'Urville. The recent
documentation of C. macloviana, C. magellanica subsp.
irrigua and C. saxatilis at high elevation sites in the San Pedro
Parks Wilderness supports Keller's hypothesis because the
range of each is centered in Colorado and states further north
(Ball and Reznicek 2002).

An additional specimen most recently determined as Carex
sp. by Julian Starr was annotated to Carex saxatilis Linnaeus
by SWCWG in 2014. Paula Durkin collected this specimen on
September 7, 1997 at Glacier Lakes on the Vermejo Park
Ranch in northern New Mexico. Licher and McGrath
attempted to return in late July 2014 to the Glacier Lakes to
collect and photograph C saxatilis and C. limosa Linnaeus.
Durkin, now a wetland specialist with the Colorado
Department of Transportation, graciously provided an aerial
photo showing the sampling sites where she thought she made
collections on behalf of the New Mexico Natural Heritage
Program in the UNM Biology Department. The sampling sites
are approximately 500 feet south of the New Mexico-Colorado
state line. During their July 2014 visit to Glacier Lakes, Licher
and McGrath failed to re-locate C. saxatilis at either of the two
presumed sampling sites or in two wetlands on the Colorado
side of the Glacier lakes area. It may have been too early to
find this species in suitable collecting condition given that
Durkin's specimen was collected about six weeks later in the
growing season than our visit.

We found a record of another Carex saxatilis specimen
supposedly collected in New Mexico during a SEINet
Collections Search (SEINet 2015). However, Cortez, Colorado
botanist Marian Rohman, the collector of this specimen
(SJNM catalog # 63928, Marian Rohman 3473. San Juan
College Herbarium), has reported (personal communication
2015) that the locality information for this specimen reported
on SEINet is incorrect, and that the specimen was, in fact,

collected in the San Juan Mountains of Colorado.

Details of the collected specimens as well as the annotated
1997 specimen follow;

Carex magellanica Lamarck subsp. irrigua (Wahlenberg)
Hiitonen {Cyperaceae, boreal bog sedge): Rio Arriba
County: San Pedro Parks Wilderness, Santa Fe National
Forest, Vega del Oso; very seepy wetland in heavily
grazed meadow with Eleocharis; sec 26, T22N, RIE,
36.10872, -106. 80833 (datum: NAD 27 CONUS), 10,420
feet, 16 August 2012, J. McGrath 1199 (UNM). Verified
by SWCWG. [first report for New Mexico].

Carex buxbaumii Wahlenberg {Cyperaceae, Buxbaum's
sedge): Rio Arriba County: San Pedro Parks Wilderness,
Santa Fe National Forest, Rio de Las Vacas, near trail.
36.105556, -106.823611, sec 27, T22N, RIE, 10,350 feet,
17 August 2012. C. Keller & R. Greiner 03530h (Jemez
Mountain Herbarium). Verified by W. R. Norris and A.A.
Reznicek. [first report for New Mexico]

Carex saxatilis Linnaeus {Cyperaceae, rock sedge): Rio
Arriba County: San Pedro Parks Wilderness, Santa Fe
National Forest, Rio de Las Vacas, streamside near trail.
36.105556, -106.823611, sec 27, T22N, RIE, 10,300 feet,
17 August 2012. C. Keller & R. Greiner 03529h (Jemez
Mountain Herbarium). Verified by W.R. Norris and A.A.
Reznicek [first report for New Mexico].

Carex saxatilis Linnaeus {Cyperaceae, rock sedge): Taos
County: Vermejo Park Ranch, Glacier Lakes; fine sandy
loam, fen, Carex limosa-Eleocharis quinqueflora
herbaceous vegetation. Associated species: Carex
aquatilis, Carex limosa, Eleocharis quinqueflora. 11,560
feet, 7 September 1997. P. Durkin 97MB024-G5 (UNM).
Determination by SWCWG. [first report for New Mexico].

The Carex tetrastachya - Carex crus-corvi Conundrum
Our 2014 review of 159 Carex specimens at the New
Mexico State University Herbarium (NMC) revealed
specimens of two Carex species for which there are no other
records in the state of New Mexico. These are Carex
tetrastachya Scheele and Carex crus-corvi Shuttleworth. Label
data for specimens of both species indicate that these sedges
were collected from Bosque del Apache National Wildlife
Reftige, Socorro County, in central New Mexico. Carex
tetrastachya has been recognized as part of the New Mexico
flora since Miriam Fritts annotated the specimen to this
species in 1998. During our 2014 review of NMC Carex
specimens, we annotated a specimen identified as Carex
stipata Muhlenberg ex Willdenow var. maxima Chapman to C.
crus-corvi Shuttleworth (subsequently verified by Reznicek).
However, closer examination of the two specimen labels raises
considerable doubt about the authenticity of these two
collections.

As it turns out, NMC also contains two additional
specimens representing these same two species, but which

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were collected by apparently the same collector (at least two of
the four specimens) at “Hageman Refuge” near Denison,
Texas. “Hageman Refuge” is Hagerman National Wildlife
Refuge, which is located about 75 miles north of Dallas,
Texas. The information on the labels of the four specimens is
minimal and sometimes incomplete.

The detailed information on the four specimens follows,
but we will first discuss the Carex tetrastachya specimens.

Carex tetrastachya Specimens

Unknown collector 153. No date of collection. Bosque del
Apache NWR, Socorro County, San Antonio, NM. NMC
catalog # 42333. Original determination Carex brittoniana
Bailey by F. J. Hermon. Determined as Carex tetrastachya
Schule (C brittoniana L.H. Bailey) by M.C. Fritts in 1998
and by A. A. Reznicek in 2012. Determined Carex
tetrastachya Scheele by SWCWG in 2014.

F.J. Herman 153. 03 May 1955. Hageman Refuge,
Grayson County, Denison, TX. NMC catalog # 39572.
Original determination Carex brittoniana Bailey.
Annotated to C. tetrastachya Schule (C. brittoniana) by
M.C. Fritts in 1998. Annotated to Carex tetrastachya
Scheele by J. McGrath Jan 2015 and by other members of
SWCWG via examination of digital images of the
specimen.

Note that the New Mexico C. tetrastachya specimen label
does not provide the collector or date of collection. However,
the Texas specimen label has the same collector number as the
New Mexico specimen. This coincidence suggests that F.J.
Herman was the unknown collector and that the collection date
for the New Mexico specimen was sometime in 1955. The
coincidence of the collector number and the incomplete label
information on the New Mexico specimen also raises doubt
that the specimen was actually collected at Bosque del Apache
National Wildlife Refuge. It is possible that the specimen was
collected at Hagerman National Wildlife Refuge and that
somehow during processing the specimen was mixed up with
material collected from Bosque del Apache National Wildlife
Refuge. A SEINet (2015) collections search revealed that the
New Mexico specimen is the only such record of this species
in New Mexico. According to a PLANTS database (USDA,
NRCS, 2015) distribution map, C. tetrastachya is known no
further west than west central Texas.

Ccirex crus-corvi Specimens

F.J. Herman 157. No date of collection. Bosque del
Apache National Wildlife Refuge, San Antonio, NM.
NMC catalog # 41697. Original determination Carex
stipata var maxima Chapm. and annotated by M. C. Fritts
in 1998 as Carex stipata Muhlenberg var. maxima
Chapman. Determined as Carex crus-corvi Shuttleworth
by SWCWG 2014 and verified by A.A. Reznicek via
examination of digital images of the specimen in 2014.

Unknown collector. #151. 03 May 1955. Wet Place
Hageman Refuge, Denison, TX. NMC catalog # 39573.
Original determination Carex stipata var. maxima
Chapman by F.J. Herman. Annotated to Carex stipata
Muhlenberg var. maxima Chapman by M. C. Fritts in
1998. Determined as Carex crus-corvi Shuttleworth by
McGrath in 2015 and by other members of SWCWG via
examination of digital images of the specimen.

Note that, like the New Mexico C. tetrastachya specimen,
the New Mexico C. crus-corvi specimen label lacks critical
information, including the date of collection. Note also that the
Texas C. crus-conn specimen was collected in the same
location and on the same date as the Texas C. tetrastachya
specimen by presumably the same collector. However, the
Texas Carex tetrastachya specimen lists F. J. Herman as the
apparent collector, but the Texas Carex crus-corvi specimen
specifically states “Det. By F. J. Herman.” The incomplete
label of the New Mexico C. crus-corvi specimen again raises
doubt that the specimen was actually collected at Bosque del
Apache National Wildlife Refuge. The similarity of the
collector's name “F.J. Herman” to that of F. J. Hermann, the
author of “Manual of the Carices of the Rocky Mountains and
Colorado Basin” (Hermann 1970) suggested that perhaps F. J.
Hermann was the collector of these specimens. We consulted
A. A. Reznicek of the University of Michigan Herbarium
(where Hermann's field notes are housed), who informed us
that F. J. Hermann collected specimens 153 and 157 in 1926 in
Ann Arbor, Michigan. Reznicek speculated that perhaps F.J.
Hermann determined the identity of the subject specimens
collected by someone else, and that during the processing of
the specimens he (Hermann) was incorrectly listed as the
collector of these plants, with his name misspelled. A
PLANTS Database distribution map reveals that C. crus-corvi
is not known further west of east central Texas (USDA,
NRCS, 2015).

Based on the poor label information, the confusion
surrounding the identity of the collector, the absence of
additional specimens and the fact that the New Mexico
specimens of both C. tetrastachya and C. crus-corvi are from a
locality substantially outside the normal range of both species,
we conclude that their presence in New Mexico is highly
doubtful both at present and in the past. We conclude that
Carex tetrastachya Scheele should be dropped from and Carex
crus-corvi Shuttleworth should not be added to the New
Mexico flora pending verification of their existence in New
Mexico.

[Footnote: Carex stipata Muhlenberg ex Willdenow var.
maxima Chapman ex Boott, the original determination for the
two Carex crus-corvi specimens described above, has never
been recognized as part of the New Mexico flora (Allred 2012;
Allred and Ivey 2012; Martin and Hutchins 1980, Wooton and
Standley 1915). The distribution of this taxon was recognized
no further west than Alabama by Standley (2002). A SEINet
(2015) collections search revealed no other specimens of this
taxon west of the Carolinas and Florida.]

(Continued on page 6)

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VERIFICATION OF TAXA IN THE NEW MEXICO
FLORA

Carex petasata Dewey

Carex petasata Dewey has been a source of confusion in
New Mexico. Allred (2012) and Norris (2012b) indicate this
species as part of the New Mexico flora. At the same time
Allred (2012) states that ”ln naming Carex wootonii,
Mackenzie (1915) argued that his new Carex wootonii
replaced what had been called Carex petasata in NM; merits
verification.” Mastrogiuseppe et al. (2002) in Volume 23 of
Flora of North America do not include New Mexico as part of
the range of distribution for Carex petasata.

Our review of the collections at UNM and NMC revealed
that only two out of 12 C. petasata specimens were previously
identified correctly as C. petasata. Our ability to accurately
identify C petasata derives from Licher and Rink's extensive
study (herbarium specimens and in the field) of C. petasata
and the very similar Carex wootonii in Arizona.

Both of these species are in Section Ch’ales, which is
characterized by gynaecandrous spikes and winged perigynia
(Ball and Reznicek 2002). Both species have large perigynia
(more than 5.5 mm long), have ovoid or obovoid to fiisiform-
shaped spikes and occur in mountain meadows and clearings
within forests (Rink and Licher 2015). However, these two
species each also occur in other, non-overlapping habitats: C.
petasata in non-montane grasslands (Mastrogiuseppe et al.
2002) and C wootonii in forests. The distribution of C.
wootonii includes southeastern and east central Arizona and
adjacent southwestern and west central New Mexico.
However, C. wootonii has been found as far north as the
Kaibab Plateau in Arizona (Rink and Licher 2015) and the
Valles Caldera National Preserve in New Mexico, but it is
unknown in Utah (SEINet 2015). During our review of SJNM
specimens in January 2015, we annotated a specimen collected
in Colorado as C. wootonii - a state record, since SEINet
(2015) reveals no other C. wootonii specimens collected in that
state. C petasata occurs in northern Arizona and northern
New Mexico and ranges as far north as Alaska (SEINet 2015;
Mastrogiuseppe et al. 2002). The general region of overlap
between the distributions of C. wootonii and C. petasata is east
central Arizona and west central New Mexico and the northern
portions of both states for as far north as the range of C.
wootonii extends. In New Mexico both species are known to
occur in the Valles Caldera National Preserve and the area
around Mount Taylor (SEINet 2015).

Key differences between the two species as derived from
Rink and Licher (2015) are described below:

Carex petasata

Carex wootonii

Perigynia light olive green,
contrasting less with tan pistil¬
late scales

Perigynia forest green, con¬
trasting sharply with brown
pistillate scales

Perigynia usually with evident
nerves

Perigynia usually nerveless

Hyaline margins on pistillate
scales 0.2-0. 8 mm wide

Hyaline margins on pistillate
scales 0-0.4 mm wide

Spikes narrowly fusiform to
narrowly obovoid with many
staminate flowers at base of
each spike

Spikes broadly fusiform to
ovoid with few staminate flow¬
ers at the base of each spike

Lowest inflorescence bract
never longer than inflorescence

Lowest inflorescence bract
sometimes longer than inflo¬
rescence

Near the conclusion of Licher and McGrath's sedge collecting
trip to the Vermejo Park Ranch in July 2014, Licher indicated
he wanted to walk up a grassy valley east of the broad Costilla
Creek valley. He thought he might find Carex petasata
because the valley was the right habitat. McGrath did not
accompany Licher, but several days later Licher reported that
he found C. petasata at about 9,650 feet elevation in that
grassy valley.

Carex macloviana d'Urville

McGrath (2003) reported finding Carex macloviana
d'Urville at timberline on the eastern shore of the uppermost
lake of the Latir Lakes, Taos County, New Mexico. However,
this single specimen collected by McGrath was determined by
Reznicek and is housed in the University of Michigan
Herbarium (MICH).

Following our review of UNM specimens, Reznicek
further verified the existence of C. macloviana as part of the
New Mexico flora by annotating two UNM specimens as
Carex macloviana d'Urville. These specimens were originally
collected in the San Pedro Parks Wilderness by Annehara
Fleck in the mid-1960's. These specimens are as follows:

Rio Arriba County: San Pedro Cabin. [36 deg. 6 ' 59” N,
106 deg. 49' 24” W, ca 3150 m - hand written in pencil]
(10,335 feet). 11 July 1964. A. Fleck s.n. (UNM).
Determined C. macloviana by A. A. Reznicek 2015.
Originally determined C. festivella Mackenzie presumably
by collector in 1964, det. C. microptera Mackenzie by
M.C. Fritts 1998, det. C. macloviana d'Urville by A. Hipp
2012.

Rio Arriba County: San Gregorio. 19 September 1964. A,
Fleck s.n. (UNM). Determined C. macloviana by A. A.
Reznicek 2015. Originally determined C. festivella
Mackenzie presumably by collector in 1964, det. C.
macloviana d'Urville by M.C. Fritts 1998, det. C.
macloviana d'Urville by A. Hipp 2012.

Presumably, Fleck's collection of C. macloviana at San
Gregorio refers to San Gregorio Reservoir in the San Pedro
Parks Wilderness.

(Continued on page 7)

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(ColUimied from page 6)

CURRENT STATUS OF NEW MEXICO CAREX

With this report we add three new Carex taxa to the New
Mexico flora: C. hiixbaiimii Wahl., C. magellanica Lam.
subsp. irrigiia (Wahlenberg) Hiitonen, and C. saxatilis
Linnaeus. We also verify the existence of Carex anreolensis
Steudel, C. macloviana d'Urville and C. petasala Dewey in the
New Mexico flora. We provide evidence to drop three species
from the New Mexico flora: Carex frankii Kunth, C. stricta
Lamarck, and C. tetrastachya Scheele. In the case of C.
tetrastachya, poor labeling of a single specimen collected
substantially outside its normal range of distribution precludes
it from being eonsidered part of the New Mexico flora.
Similarly, a single poorly labeled specimen of Carex crus-
corvi Shuttleworth also collected well outside its normal range
of distribution precludes its addition to the New Mexieo flora
at this time. Therefore, the existence of the latter two species
in New Mexieo must be verified beyond the above-mentioned
poorly labeled specimens. Finally, we provide a nomenelatural
upgrade for one species. Carex lentictilaris Michaux var.
lipocarpa (Holm) L. Standley is now recognized as Carex
kelloggii W. Boott.

Based on the findings reported in this paper and the species
reported in Allred (2012), there are apparently 92 Carex
species documented in New Mexico. Additional field searches
will very likely add more Carex species to the New Mexieo
flora. However, at least 10 species in the New Mexico flora
are based on very few records, sometimes as few as a single
specimen. In fact, we have encountered no records of
specimens collected in New Mexico for one species, Carex
lillra L.H. Bailey (reeognized as such by Rink and Licher
2015, listed as Carex spissa L.H. Bailey in Allred 2012 and
Allred and Ivey 2012). However, we have had an
undocumented report of this species occurring in the
Peloncillo Mountains of southwestern New Mexico (Richard
Worthington - personal communication to McGrath in 2003).
The few documented New Mexico records of some Carex
species (e.g., C. blanda Dewey, C. rosea Schkuhr ex
Willdenow, C. leptalea Wahlenberg, C. garberi Fernald) are
substantially outside their normal range of distribution (Norris
2012b; USDA, NRCS 2015; SEINET, 2015). Additional field
work and tracking down and examination of plant specimens
w ill be necessary to determine whether these species actually
exist within the state. Two other Carex species tentatively
recorded from New Mexico {Carex tenera Dewey, Carex
lahnensis Smiley) do not match precisely the expeeted
characteristics of these two species despite their determination
by Carex expert A. A. Reznicek. Furthermore, one Carex
species (Carex pityophila Maekenzie), currently included in
the New Mexico flora (Allred 2012) and under current
systematic study elsewhere, may be merged with another
taxon in Carex section Acrocystis sometime in the future
(Bruce Ford, University of Manitoba Herbarium, Derick
Poindexter, University of North Carolina Herbarium - personal
communication). We also suspect that Carex chihuahuensis
MacI.enzie may eventually become subsumed into Carex alma
L. II. Bailey, but more research is required to substantiate this

supposition.

ACKNOWLEDGMENTS

We thank Tim Lowrey, Curator of the UNM Herbarium,
Donovan Bailey, Curator of the NMC Herbarium, Ken Heil,
Curator of the San Juan College (SJNM) Herbarium, Amy
Ganguli, Curator and Kelly Allred, Emeritus Curator of the
NMCR Herbarium, Richard Worthington, Curator of the
UTEP Herbarium, and Leigh Johnson, Curator of Vascular
Plants at the Stanley L. Welsh Herbarium at Brigham Young
University for granting us permission to review specimens.
We also thank Phil Tonne, Collections Manager of the UNM
Herbarium and Lillis Urban, Postdoctoral Curator at NMC
herbarium for facilitating our review of the Carex specimens
and patiently answering our questions. Additionally, we thank
Russell Kleinman of Western New Mexico University and Phil
Tonne for photographing specimens that facilitated our review
via the internet when personal specimen examination was not
possible by some of us. We also extend our thanks to the
Vermejo Park Ranch for allowing Licher and McGrath to
make plant eollections on the Ranch and to Les Dhaseleer,
Natural Resource Manager, and Tanner Laird for their
guidance and assistance during the collecting trip to the ranch.

LITERATURE CITED

Allred, K.W. 2012. Flora Neomexicana I: Annotated
Checklist. An Introduction to the Vascular Plants of New
Mexico, with Synonymy and Bibliography. 2"‘* ed. Range
Science Herbarium, Dept, of Animal and Range Sciences,
New Mexico State University, Las Cruces, NM. Published
by the author and available at www. Iulu.com.

Allred, K.W. and R.D. Ivey. 2012. Flora Neomexieana III: an
Illustrated Identification Manual. Published by the authors
at www.lulu.com.

Ball, P.W. and A. A. Reznicek. 2002. Carex Linnaeus, pp. 254
-572. IN Flora of North America Editorial Committee, ed.
2002. Flora of North America North of Mexico Vol. 23.:
Magnoliophyta: Commelinidae (in part): Cyperaceae.
Oxford University Press, New York, NY.

Dragon, J.A. and D.S. Barrington. 2009. Systematics of the
Carex aquatilis and C lenticularis lineages:
geographically and ecologically divergent sister clades of
Carex section Phacocystis (Cyperaceae). Am. Jour. Bot 96
(10): 1896-1906.

Ford, B.A and A. A. Reznicek. 2002. Carex Linnaeus sect.
Squarrosae J. Carey, pp 518-519. IN Flora of North
America Editorial Committee, ed. 2002. Flora of North
America North of Mexico Vol. 23.: Magnoliophyta:
Commelinidae (in part): Cyperaceae. Oxford University
Press, New York, NY.

Hermann, F.J. 1970. Manual of the Carices of the Rocky
Mountains and Colorado Basin. Agricultural Handbook
No. 374. USDA Forest Service. 397 pages.

Mackenzie, K.K. 1915. Two new sedges from the
southwestern United States. Smithsonian Misc. Collections
65 (7): 1-3.

Martin, W.C. & C.R. Hutchins. 1980. A Flora ofNew Mexico,

(Continued on page 8)

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Vol. 1. Reprinted 2001, Bishen Singh Mahendra Pal Singh,
Dehra Dun -248 001 India and Koeltz Scientific Books,
Koenigstein, Germany.

Masirogiuseppe, J., P.E. Rothrock, A.C. Dibble and A. A.
Rjznicek. 2002. Carex Linnaeus sect. Ovales Kunth, pp.
322-378. IN Flora of North America Editorial Committee,
ed. 2002. Flora of North America North of Mexico Vol.
23.: h'iagnoliophyla: Commelimdae (in part): Cyperaceae.
Oxford University Press, New York, NY.

McGrath, J. 2003. Plant Distribution Reports. The New
Mexico Botanist 25:7.

NMBCC. 2007. The NMBCC Web Site. New Mexico
Biodiversity Collections Consortium, New Mexico, USA.
A ailable at http://NMBiodiversity.org. (Accessed: late
December, 2014).

Norris, W.R. 2012a. Plant Distribution Reports. The New
Mexico Botanist 57:3. May 30, 2012.

Norris, W.R. 2012b. Carex., pp. 580-592. IN Allred, K.W. and
R. D. Ivey. 2012. Flora Neomexicana III: an Illustrated
Identification Manual. Published by the authors and at
V vw.lulu.com.

Rink G. and M. Licher. 2015. Carex of Arizona: Cyperaceae
- Sedge Family: Part 1: Family Description, Key to the

genera and Carex L. Canotia, submitted for publication.

1 1 5 pages.

SEINet (SOUTHWEST ENVIRONMENTAL

INFORMATION NETWORK), SEINet. 2014-2015.
http//:swbiodiversity.org/seinet/index.php. Accessed in
December 2014 and January-February 2015.

Standley, L.A. 2002. Carex Linnaeus sect. Viilpinae (Heuffel)
H. Christ, pp. TTh-H^. IN Flora of North America
Editorial Committee, ed. 2002. Flora of North America
North of Mexico Vol. 23.: Magnoliophyta: Commelinidae
(in part): Cyperaceae. Oxford University Press, New York,
NY.

Standley, L.A., J. Cayouette and L. Bruederle. 2002. Carex
Linnaeus sect. Phacocystis Dumortier, pp. 379-401. IN
Flora of North America Editorial Committee, ed. 2002.
Flora of North America North of Mexico Vol. 23.:
Magnoliophyta: Commelinidae (in part): Cyperaceae.
Oxford University Press, New York, NY.

USDA, NRCS. 2014, 2015. The PLANTS Database (http://
plants.usda.gov, 30 December 2014, 16 January 2015, 7
February 2015). National Plant Data Team, Greensboro,
NC 27401-4901 USA.

Wooton, E.O. and P.C. Standley. 1915. Flora of New Mexico.
Contr. U.S. Natl. Herb 19: 1-794.

"The New Mexico Botanist" is a digital newsletter to help
make information about the flora of New Mexico readily
available to the botanical community. The newsletter focuses on
floristic and taxonomic information of interest to the botanists of
the state, such as new state records, nomenclatural notes and
explanations, brief taxonomic summaries, reports of inventories,
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announcements and reports of meetings and conferences, and
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Submissions will be reviewed by the editor, who may ask

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- Kelly Allred, editor

J^otnny Is the nnturnl science thnt trnnsinlts the hnoivleilgc of plonts.

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