MALAC QL 401 .N68 v.13 no.l NvVAPEX Trimestriel de la Société Belge de Malacologie association sans but lucratif Quarterly of the Belgian Malacological Society VOL. 13 (1) 2012 10 MARS SOMMAIRE Articles originaux - Original articles C. Vilvens New species and new records of Seguenzioidea and Trochoidea (Gastropoda) from French Polynesia 1 R. Houart Description of a new species in the Siratus pliciferoides group (Gastropoda: Muricidae) from the Philippines 25 E. Rolân & F. Rubio A new species of the genus Leucorhynchia (Gastropoda, Turbinidae) ffom West Africa 29 D. P. Cilia A new Javan species of Agaronia Gray, 1839 (Neogastropoda, Olividae) 33 R. Houart Description of Muricopsis (Muricopsis) gorii (Gastropoda: Muricidae: Muricopsinae) from Southern Sâo Tomé 37 E. Rolân & H. G. Lee Rissoinaparkeri (Mollusca: Rissooidae): a curious Caribbean species of uncertain status 43 NOVAPEX/SOCIETE C. Vilvens Prochaines activités 1 C. Delongueville, R. Scaillet • Relations trophiques entre quelques Pyramidelloidea ■yjjjjP et leurs hôtes 3 M. Alexandre L'écho des réunions : R.Scaillet & C. Delongueville : "De l’informatique à un outil pratique de détermination" (10/12/2011) 7 (suite du sommaire en dernière page de couverture) ISSN 1375-7474 Avec le soutien de EwôII Sale Stewart Llbrary Périodique trimestriel Bureau de dépôt 1370 Jodoigne Wallonie SEP 1 7 2013 Academy of Naturel Sciences of Philadelphia COTISATIONS / MEMBERSHIP 2012 Membres résidant en Belgique (NOVAPEX et les numéros hors série) Membre effectif.40 € (sans le service du bulletin) Personne appartenant à la famille d'un membre effectif et ayant même résidence.15 € Versement à effectuer auprès de la Banque ING, au n° 363-0831752-17 de M. Marc Alexandre, rue de la Libération, 45, 6182 Souvret. 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(Ail bankcharges to be paid by customer) malâc MOI ,M(o& V-1 b I Suivant un accord avec la SIM (Societa Italiana di Malacologia), nos membres européens qui souscrivent également à cette société pour 2012, peuvent payer leur(s) cotisation(s) à la SBM pour les deux sociétés. Les membres européens de la SIM peuvent faire de même chez eux. By common agréé ment with SIM, our European members that subscribe to that society for 2012 can pay to the SBM the membership fees for both societies. The European members ofthe SIM can do the same paying to their society. Dans ce cas, vous obtiendrez une réduction de 2 € pour la SIM et de 2 € pour la SBM, soit : In this way y ou can hâve a discount of 2 € for the SIM and of 2 € for the SBM : SIM (Bol. Malacologico + Notiziario SIM + SBM (Novapex + Novapex/Société + Hors Séries) Suppléments).€ 58,00 € 53,00 (Belgian members: € 38,00) Editeur responsable: C. Vilvens, rue de Hermalle, 113, B-4680 Oupeye President honoraire ( Honorary President) • M. R. Duchamps, av. Mozart, 52, 1190 Bruxelles Vice-President honoraire (Honorary Vice-President) • M. E. Waiengnier t, rue Camille Wollès, 42, 1030 Bruxelles CONSEIL D’ADMINISTRATION President Vice- president & Bibliothécaire ( Vice-president and Librarian) TRESORIER (Treasurer) SECRETAIRE (Secretary) Administrateur (Novapex Manager) • M. C. Vilvens, rue de Hermalle, 113, 4680 Oupeye 04/248.32.25 • M. E. Meuleman, rue du Chafour, 4, 4171 Poulseur 04/380 55 16 • M. M. Alexandre, rue de la Libération, 45, 6182 Souvret 071/46.12.88 • Mme A. LanglEIT, av. Cicéron, 27, bte 92, 1140 Bruxelles 02/726.17.61 • M. R. Houart, St. Jobsstraat, 8, 3400 Landen (Ezemaal) 016/78.86.16 Internet : http://www.societe-belge-de-malacologie.be e-mail : vilvens.claude@skynet.be ou e.meuleman@skynet.be Les articles et textes présentés dans cette revue réflètent l’opinion personnelle de leur(s) auteur(s) et non pas nécessairement celle de la Société ou de l'éditeur responsable. Tous droits de reproduction, de traduction et d'adaptation des articles publiés dans ce bulletin, réservés pour tous pays. _ AU rights of reproduction are reserved with the written permission ofthe board. _ Publications precedentes/Former publications: - Bulletin Mensuel d'information (1966-1971) - INFORMATIONS de la Société Belge de Malacologie (1972-1985) - ARION (1986-1999) -APEX (1986-1999) SOCIETE BELGE DE MALACOLOGIE C. VlLVENS NOVAPEX 13(1): 1-23, 10 mars 2012 New species and new records of Seguenzioidea and Trochoidea (Gastropoda) from French Polynesia Claude VlLVENS Rue de Hermalle, 113 - B-4680 Oupeye, Belgium Scientific Collaborator, Muséum national d'Histoire naturelle, Paris. vilvens. claude@skynet. be % LIBRARY ^■PHILADEI-P^^ KEYWORDS. Gastropoda, Société, Australes, Tarava, Calliotropidae, Chilodontidae, Cantharidinae, Calliostomatidae, Margaritinae, Calliotropis, Herpetopoma, Thalotia, Calliostoma, Gaza , new species. ABSTRACT. New records of eight known Seguenzioidea and Trochoidea species from French Polynesia area are listed, extending the distribution area of some of them. Seven new species are described and compared with similar species: Calliotropis amtnos n. sp., Herpetopomapoichilum n. sp., Thalotia tiaraeides n. sp., T. khlimax n. sp., T. polysarchosa n. sp., Calliostoma (Fautor) lepton n. sp., Gaza polychoronos n. sp. RESUME. De nouveaux relevés de huit espèces connues de Seguenzioidea and Trochoidea provenant de Polynésie Française sont listés, étendant ainsi faire de distribution d'un certain nombre d'entre elles. Sept nouvelles espèces sont décrites et comparées avec des espèces similaires : Calliotropis ammos n. sp., Herpetopoma poichilum n. sp., Thalotia tiaraeides n. sp., T. khlimax n. sp., T. polysarchosa n. sp., Calliostoma (Fautor) lepton n. sp., Gaza polychoronos n. sp. INTRODUCTION The deep-water fauna of the south-westem Pacific is poorly known and not very documented. In the last décades occurred only a few books (Salvat & Rives, 1975; 1990) or papers (Cernohorsky, 1980; Trôndlé & Boutet, 2009; Vilvens, 2009b) about Polynesian shells; because some of these shells also occur in Hawaiian Islands, two recent books about Hawaiian malacofauna (K.ay, 1979; Severn, 2011) can sometimes be useful. At the présent time, one can mention that a team of malacologists (with J.Letourneux and R.Gourguet as contacts) is planning a compendium of the French Polynesian sea shells. This explains the huge interest of deep-water material from French Polynesia brought recently in MNHN by the two campaigns BENTHAUS and TARASOC. The BENTHAUS expédition was conducted in November 2002 aboard R.V. A lis by IRD (Institut de Recherche pour le Développement) with B. Richer de Forges as mission leader. It surveyed the Austral Islands from Southwest to northeast (in an area covering 21°-29°S to 153°-140°W) from 60 to 1350 meters depth, with the aim to study the fauna living in these waters, especially zoo-benthos organisms like molluscs and crustaceans, and to perform seamounts cartography of this area. Dredging has been performed at 134 stations and trawling at 19 stations. More recently, the TARASOC campaign was conducted in September and October 2009 aboard R.V. Alis by IRD and MNHN (Département Systématique & Evolution) with P. Bouchet as mission leader. The aim was to explore the fauna living on the benthos of Société Islands and the Tarava seamounts and to study the relations between isolation of these seamounts and their biodiversity, in partnership with MarBOL (Marine Barcode Of Life) project and CenSeam (Census of Marine Life on Seamounts) program. The Tarava Seamounts are 700 km long, are much older (about 35 to 50 million years) than Société seamounts and hâve some submarine summits reaching 600 to 900 m depth; they hâve been discovered in 1996 by the ZEPOLYF (Zone Economique de la Polynésie Française) program and no biological survey was ever undertaken in this area. During this campaign leaded in an area covering 15°- 20°S to 155°-148°W, 213 dredging and trawling operations hâve been performed. The présent paper reports new records and new species of Seguenzioidea and Trochoidea species collected during these two campaigns. C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia 10 ° kahangci. Groupe 'apten trio nul 140° I _ alu tu A Eiaos' NukuJhra d uaM d£ wna r 130 e 75555$ axa Fatu-Hiva, 5^85 .. „ 1 ’tA s 4 1 x 23 7 mnr 1 3 rwi ~~' 7 mm; 10 ' 12 ‘ Parat yP e MNHN I4-.5. CW»**, derMosa Vilvens, TJff ” mb " i “ l — • stn CP992], 22.3 x 25.4 mm. ’ vanuatL h 748-775 m [MUSORSTOM 8, 8 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 9 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Agathodonta sp. Figs 28-30 Cantharidus marmoreus (Pease, 1868) Figs 31-32 Material examined. French Polynesia, Société Islands. TARASOC: stn DW3499, 17°41 ’S, 149° 1 T W, 550-700 m, 1 dd. Comments. This single specimen is so broken and and the whorls are so eroded that it is very difficult to give reliable a possible description and comparison with known species. Superfamily TROCHOIDEA Rafinesque, 1815 Family TROCHIDAE Rafinesque, 1815 Subfamily TROCHINAE Rafinesque, 1815 Genus :Trochus Linnaeus, 1758 Type species: Trochus maculatus Linnaeus, 1758 (by o. d.) - Recent, Indo-Pacific. Trochus niloticus Linnaeus, 1767 Trochus marmoreus Pease, 1868: 287, pl.24 fig. 9. Type locality: Tuamotu Archipelago, Paumotus. Cantharidus marmoreus - Cemohorsky, 1980: 113, figs 1-3. Calliostoma marmoreum — Kaicher, 1979: card#2102. Material examined. French Polynesia, Australes Archipelago. BENTHAUS: stn DW 1884, 27°54'S, 143°33'W, 570-620 m, 1 dd sub. - Stn DW 1885, 27°52'S, 143°33'W, 700-800 m, 1 dd. - Stn DW1926, 24°38'S, 146 o 0EW, 50-90 m, 3 dd. - Stn DW1932, 24°41’S, 146°02'W, 500-800 m, 2 dd. - Stn DW 1933, 24°4ES, 146°0rW, 500-850 m, 3 dd. - Stn DW 1936, 24°40'S, 145°57'W, 80-100 m, 1 dd. Distribution. French Polynesia, Tuamotu Archipelago, intertidal (dead) (Letoumeux et al., p.c.); Australes Archipelago, 90-700 m (dead). Trochus niloticus Linnaeus, 1767: 1227, no. 579. Type locality : Indian Océan. Trochus niloticus - Cemohorsky, 1972: pp 38-39 pl-8, fig 1 Trochus niloticus - Salvat & Rives, 1975: 256, Trochus niloticus - Kaicher, 1979: card#2175. Trochus niloticus - Poppe, Tagaro & Dekker, 2006: 80-81, figs 70-71, pi. 33, figs 4-8. Material examined. French Polynesia, Australes Archipelago. BENTHAUS: stn DW1913, 27°02'S, 146°00'W, 120 m, 1 dd juv. - Stn DW1917, 27°03'S, 146°04'W, 50-60 m, 4 dd juv. - Stn DW2015* 22°38'S, 152°50'W, 250-280 m, 2 dd juv. Distribution. Indo-Pacific, low subtidal; French Polynesia, Australes Archipelago, 60-250 m (dead juv). Subfamily CANTHARIDINAE Gray, 1857 Preliminary comment. The discrimination between the various généra belonging to Cantharidinae {sensu Hickman & Mc Lean, 1990) is not very clear, especially using only conchological features (e.g. shape of the whorls, round aperture or with a vertical columella, angle between inner and outer lip, tooth on the columella or not, base with a gentle or a steep slope, umbilicus présent or absent). The new species described here are placed into one of the existing généra, although some features don't fit exactly with the features of the chosen genus, because it seems useless to add to confusion by creating new généra. Genus Cantharidus Montfort, 1810 Type species: Trochus iris Gmelin, 1791 (by o.d.) = [ lmax opalus Martyn, 1784 (by s.d., 1847) - Recent New Zealand. Cantharidus sp. Figs 43-44 Material examined. Australes Archipelago. BENTHAUS: stn DW 1939, 23°50'S, 147°42'W, 100 m, 1 dd. Comments. This single specimen (7.3 x 5.6 mm) is highly eroded and lias a broken protoconch : it is very difficult to describe accurately the spiral cords ontogeny. Genus Jujubinus Monterosato, 1884 Type species: Trochus matoni Payraudeau, 1826 = Trochus exasperatus Pennant, 1777 (by s.d. Pilsbry, 1889)] - Recent, north -eastern Atlantic. J ujubinus geographicus Poppe, Tagaro & Dekker, 2006 Figs 39-42 Jujubinus geographicus Poppe, Tagaro & Dekker, 2006: 88-89, pl. 38, figs 2. Type locality : Philippines, Balicasag Is., 80-150 m. Archipelago. BENTHAUS: stn DW 1880, 27°55'S, 143°30'W, 90-94 m, 3 dd, 1 dd sub. - Stn DW 1927, ]46 ° 02 ' W ' 95-105 m, I dd. - Stn DW1932, ôUio’ ,46 ° 02 ’ W ’ 500-800 m, I dd. - Stn DW 1958, | 49°30'W, 80-150 m, I dd sub. - Stn DW2013, 22°39'S, 152°50'W, 80-93 m, 1 dd ?nn t 7 ib r ti0n ' Phili PP' nes > 80-150 m (Poppe et al., ^006); French Polynesia, Australes Archipelago, 93- m (dead); French Polynesia, Tuamotu Archipelago, 0-100 m (dead) (Letoumeux et al., p.c.); French Polynesia, Société Islands, 0-85 m (dead) (Letoumeux et al., p.c.). 10 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 Comments. Regarding the huge gap between the type locality (Philippines) and French Polynesia, one can wonder to find this species in Australes Archipelago stations. But no clear conchological distinctions can be made between santples from the two areas, except that some Polynesian shells are less elevated, the subsutural spiral cord can be divided into two cords on the last whorls and the colour of the whorls is orange based rather then red wine. Genus Thalotia Gray, 1847 Type species: Monodonta conica Gray, 1827 [=Trochus pictus Wood, 1828] (by o.d.) - Recent, Southern Australia. Thalotia tiaraeides n. sp. Colour Figs F1-F2, Figs 33-34, Table 3 Type material. Holotype (6.4 x 6.4 mm) MNHN (24969). Paratypes: 3 MNHN (24970, 24971). Type locality. French Polynesia, Australes Archipelago, Président Thiers Bank, BENTHAUS, stn DW 1932, 24°41’S, I46°02'W, 500 800 m. Material examined. French Polynesia, Australes Archipelago. BENTHAUS: stn DW1923, 27°01’S, 146°05'W, 360-840 m, 1 dd. - Stn DW1932, 24°41'S, 146°02'W, 500-800 m, 1 dd (holotype). — Stn DW 1955, 23°19'S, I49°26'W. 750-850 m, 1 dd sub. - Stn DW1958, 23°20'S, 149°30'W, 80-150 m, 1 dd sub. - Stn DW 1999, 22°25’S, 151°22’W, 270-500 m, 1 dd. - Stn DW2003, 22°28'S, 151°19’W, 250-330 m, 1 dd (paratype). - Stn DW2018, 22°37'S, 152°49’W, 770-771 m, 1 dd. - Stn DW2020, 22°37’S, 151 0 49'W,’ 920-930 m, 2 dd (paratypes). Distribution. French Polynesia, Australes Archipelago, 150-920 m (dead). Diagnosis. A small white Cantharidinae species with a moderately elevated spire, a subsutural angulate keel making shoulder and a thick peripheral keel; 6 main granular spiral cords on last whorl with a thick peripheral keel; slightly convex base with up to 10 Iow, nearly smooth spiral cords; narrow and deep umbilicus, partly covered by a columellar expansion. Description. Shell of small size for the genus (height up to 6.6 mm, width up to 6.4 mm), as high as wide or slightly higher than wide, rather thick, conical with a subsutural angulate keel making shoulder with gentle slope; spire moderately elevated, height 0.9x to 1,2x width, 3.3x to 3.6x aperture height; umbilicus narrow and deep, partly covered by a columellar expansion. Protoconch approximately about 200 pm (eroded or damaged on ail samples), without visible varix. Teleoconch up to 6.1 convex whorls, bearing 6 main, similar in size spiral granular cords and peripheral keel produced by three close thin spiral cords. Suture visible, not canaliculated. First whorl convex, sculptured by 4 smooth primary spiral cords and weak, low, wide axial folds; PI, P2 and P3 similar in size and evenly pinkish coloured, abapical cord P4 thinner with regular orange dashes; distance between P3 and P4 greater than distance between other cords. On second whorl, axial folds thinner and making PI, P2 and P3 subgranular. On third whorl, axial sculpture weakening; shoulder appearing with a round keel made by PI at fîrst adapical quarter; PI stronger than other cords; S2 appearing rather quickly as strong as P2 and P3; strong suprasutural angulation above P4. On fourth whorl, keel much more angulate; S3 appearing above suprasutural angulation, quickly as strong as P3; Tl appearing at end of whorl on shoulder between suture and PI, quickly as strong as P2 but weaker than PI; T2 appearing between suprasutural angulation and P4, both cords thinner than other cords; ail spiral cords granular. On fifth whorl, S3 stronger than P3, S2 and P2; T3 appearing between angulation and T2, similar in size to T2 and P4; beads of cords prosoclinely elongated. On last whorl, additional Ti appearing by intercalation; P4, T2 and T3 very close, hard to distinguish and forming a thick peripheral keel. Aperture subquadrangular, slightly transversally elongated; outer lip thickened inside, meeting inner lip with obtuse angle. Columella straight, almost vertical, with a strong tooth at abapical third. Base weakly convex to almost fiat, with 8-10 smooth to subgranular, similar in size spiral cords; distance between cords smaller than cords. Umbilicus fairly narrow (diameter about 9% of shell width) and deep, funnel shaped with thin axial ridges inside, partly covered by columellar expansion. Colour of protoconch and teleoconch first whorl deep pink, next whorls orange brown with wide, irregular axial white patches; spiral cords of three last whorls alternating pinkish white and orange dashes; umbilical area and inner part of base light pink, spiral cords of outer part alternating orange and pink. TW H w HA H/W H/HA holotype 6.1 6.4 6.4 1.8 1.00 3.56 paratype 1 5.8 5.9 6.4 1.8 0.92 3.28 paratype 2 6.1 6.2 5.8 1.8 1.07 3.44 paratype 3 5.7 —- 6.6 5.5 1.9 1.20 3.47 Table 3. - Thalotia tiaraeides n. sp.: Shells measurements in mm for types. C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Discussion. Regarding the straight, vertical columella, the rather sharp basal columellar tooth, the obtuse angle between inner and outer lip and the narrow umbilicus, and taking in account the shape of the new species similar to the one of Thalotia ( Odonîotrochus ) chlorostoma (Menke, 1843) (despite a very different size), the genus Thalotia (as commented by Wilson, 1993) seems the more appropriate for this new species. The new species is rather close to Jujubinus hubrechti Poppe, Tagaro & Dekker, 2006 (figs 35-36) from Philippines, but this similar in size species has a more elevated spire, a much stronger adapical spiral cord on the shoulder with elongate beads, a much stronger peripheral spiral cord with stronger, much rounded beads and different colours (green background colour instead orange brown). Etymology. Tiara shaped (Greek : napa and eiôqQ, adjective agreeing with a neutral noun - with reference to the general shape of the shell reminding an ancient tiara. Thalotia khlimax n. sp. Colour Figs E1-E2, Figs 37-38 Type material. Holotype (6.2 x 5.1 mm) MNHN (24972). Paratype (5.4 x 4.7 mm) MNHN (24973). Type locality. French Polynesia, Australes Archipelago, BENTHAUS, stn without data. Material examined. French Polynesia, Australes Archipelago. BENTHAUS: stn DW 1885, 27°52'S, 143°33'W, 700-800 m, 1 dd. Stn DW 1903, 27°27’S, 144°04'W, 400-800 m, I dd sub. - Stn DW1923, 27°0rS, 146°05 f W, 360-840 m, 1 dd (paratype). - Stn DW1961, 23°2FS, 149°34'W, 470-800 m, 1 dd. - Stn with no data, 1 lv (holotype), 1 dd sub. Distribution. French Polynesia, Australes Archipelago, 700-800 (dead). Diagnosis. A small white Cantharidinae species with a moderately elevated spire and a small subsutural Figures 16-30. Scale bars = 1 mm except scale figs 18- angulate keel making shoulder; 7 main granular spiral cords on last whorl, the two abapical cords slightly thinner, doser and separated from the other cords by a weakly concave area; slightly convex base with up to 10 low, smooth to subgranular spiral cords; moderately wide, deep umbilicus. Description. Shell of small size for the genus (height up to 6.2 mm, width up to 5.1 mm), higher than wide, rather thin, conical with a small subsutural angulate keel making shoulder with gentle to almost horizontal slope; spire moderately elevated, height l.lx to 1.2x width, 2.8x to 3.9x aperture height; umbilicus narrow and deep. Protoconch about 100 pm, exserted, without visible varix. Teleoconch up to 6.1 convex whorls, bearing 7 spiral granular cords; 5 adapical cords similar in size and 2 th inner, close peripheral cords with a concave area between two groups. Suture not canaliculated, poorly visible. First whorl convex, sculptured by 3 primary spiral cords P2, P3 and P4 similar in size and evenly pinkish coloured; PI appearing at mid whorl, similar to other cords. On second whorl, ail cords subgranular; PI, P2 and P3 thicker, P4 thinner; shoulder appearing with a round keel made by P2 at First adapical quarter; distance between P3 and P4 greater than distance between other cords; S4 appearing at end of whorl, similar in size and shape to P4. On third whorl, S3 appearing, quickly as strong as P3; additional Tl appearing at mid whorl on shoulder between suture and PI. On fourth whorl, PI thickening; keel more angulate and moving adapically to PI; Tl similar in size to PI; ail spiral cords granular; distance between S3 and P4 still greater than distance between other cords, distance between P4 and S4 smaller than distance between other cords. On last whorls, T2 appearing between Tl and PI, similar in size to Tl; beads ot ail cords rounded, without interspaces between; distance between PI, P2, P3 and S3 similar to cords; no interspace between P4 and S4; area between S3 and P4 concave, wider than cords, with prosocline, racher thick threads. - 100 pm. 16-19. Herpetopomapoichilum n. sp, Société Islands. 16-17. Holotype MNHN P4966Ï 4SS ™ DWM^Ïst^mm 041 ’ ^ X2 ' 5 ^ ^ MN ™ (24967) ’ 52 °' 572 m m [TARASOC, stn VÜVenS & Hér ° S ’ 2 °° 3, Caled ° nia ’ T ° Uh ° area ’ 5 °^ 62 m [MONTROUZIER, f^H erpet0pomacom, Z atum (Pcase ’ 1861 >’ Société ls| ands. 493-540 m [TARASOC, stn DW3429], 3.0 x ] 4 ~ 2 ^J aCeUchelus f° veolatus (A ' Adams ’ 1851 >’ Société Isla "ds, 380-758 m [TARASOC, stn DW3390], 3.6 x 26-27. Vaceuchelus sp., Tuamotu Archipelago, 976-997 m [TARASOC, stn DW33491 4 ? v ^ -r 28-30. Agathodonta sp., Société Islands, 550-700 m [TARASOC, stn DW3499], 5 0 x4 0 mm h™’ 12 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 13 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Aperture subcircular; outer lip with a weak peripheral angulation and meeting inner lip with obtuse angle. Columella straight, almost vertical, with weak swelling (holotype) or blunt tooth (paratype) at abapical third. Base weakly convex to almost fiat, with 10 smooth to subgranular, similar in size spiral cords; distance between cords smaller than cords. Umbilicus moderately wide (diameter about 15% of shell width) and deep, funnel shaped with thin axial ridges inside. Colour of protoconch pinkish white; teleoconch first whorl pinkish mauve; next whorls orange brown with irregular axial white and pink patches; periphery altemating orange dashes and narrower axial pinkish white fiâmes; umbilical area and inner part of base nacreous pink, spiral cords of outer part altemating orange and pink. Discussion. For the same reason as for Thalotia tiaraeides n. sp., the genus Thalotia seems the more appropriate for this new species. Thalotia khlimax n.sp. may be compared to T. tiaraeides n. sp., but this similar in size species has a wider shoulder, a thick peripheral keel and an umbilicus partly covered by a columellar expansion. The new species may be compared to Kanekotrochus vietnamensis Dekker, 2006 from Vietnam, but this species is much greater (height about 15 mm) for a same number of whorls, a more depressed spire and no umbilicus. Etymology. Staircase (Greek : %À.ijiaÇ) - with reference to the staggered shape of the shell. Thalotiapolysarchosa n. sp. Colour Figs D1-D2, Figs 45-50, Table 4 Type material. Holotype (4.5 x 4.0 mm) MNHN (24974). Paratypes : 3 MNHN (24975). Type locality. Australes Archipelago, BENTHAUS, stn DW 1894, 27°40'S, 144°22'W, 100 m. Material examined. French Polynesia, Australes Archipelago. BENTHAUS: stn DW 1894, 27°40'S, 144°22'W, 100 m, 6 dd (holotype and paratypes), 1 dd sub. - Stn DW 1939, 23°50'S, 147°42'W, 100 m, 1 dd, 2 dd sub. - Stn DW1946, 23°49'S, 147°4TW, 100- 200 m, 1 dd. - Stn DW2018, 22°37'S, 152°49'W, 770-771 m, 4 dd. French Polynesia, Société Islands. TARASOC: stn DW3420, 16°46'S, 151°04’W, 550 m, 1 dd juv. - Stn DW3429, 16°43'S, 150°38'W, 493-540 m, 2 dd juv. - Stn DW3434, 16°42'S, 151°03'W, 700-785 m, 1 dd juv. - Stn DW3435, 16°41'S, 151°02'W, 500-612 m, 5 dd juv. - Stn DW3451, 16°53'S, 151°2TW, 440-490 m, 1 dd juv. - Stn DW3458, 16°46'S, 151°23’W, 573- 611 m, 1 dd. - Stn DW3459, 17°28’S, 149°48'W, 485- 560 m, 1 dd sub, 3 dd juv. - Stn DW3460, 17°28'S, 149°50'W, 660-680 m, 4 dd juv. - Stn DW3476, 17°29'S, 149°45'W, 435-490 m, 7 dd sub. - Stn DW3481, 17°29'S, 149°45'W, 610 m, 7 dd juv. - Stn DW3482, 17°29'S, 149°45 r W, 440 m, 2 dd. Distribution. French Polynesia, Australes Archipelago, 100-770 m (dead); Société Islands, 440- 700 m (dead). Diagnosis. A small reddish or orange brown Cantharidinae species with a moderately elevated spire with a weak suprasutural angulation; 8 main granular spiral cords on last whorl, the two abapical cords roundly granular, the médian cords nearly smooth and the subsutural cords granular with axially elongated beads; base moderately convex with about 12 low, smooth cords; moderately wide, deep umbilicus partly covered by a columellar expansion. Description. Shell of small size for the genus (height up to 4.6 mm, width up to 4.0 mm), higher than wide, rather thin, weakly cyrtoconoidal with an angular periphery; spire moderately elevated, height l.lx to 1.2x width, 2.8x to 3.8x aperture height; umbilicus moderately wide and deep. Protoconch about 100-120 pm, exserted, with a thin varix. Figures 31-44. Scale bars = 5 mm. 31-32. Cantharidus marmoreus (Pease, 1868), Australes Archipelago, 700-800 m [BENTHAUS, DW1885], 6.5 x 4.1 mm; 33-34. Thalotia tiaraeides n. sp, holotype MNHN (24969), Australes Archipelago, 500-800 m [BENTHAUS stn DW 1932], 6.4 x 6.4 mm. 35-36. Jujubinus hubrechti Poppe, Tagaro & Dekker, 2006, holotype NMP, Philippines, Balicasag, 80-150 m, 7.0 x 6.2 mm - Photographs courtesy of Conchology, Inc. 37-38. Thalotia khlimax n. sp., holotype MNHN (24972), Australes Archipelago (BENTHAUS, stn with no data], 6.2 x 5.1 mm. 39-42 .Jujubinus geographicus Poppe, Tagaro & Dekker, 2006. 39-40. Australes Archipelago, 95-105 m [BENTHAUS: stn DW1927], 5.3 x 4.1 mm; 41-42. Holotype NMP, Philippines, Mactan Island, 80-150 m, 4.9 x j. 8 mm - Photographs courtesy of Conchology, Inc. 43-44. Cantharidus sp., Australes Archipelago, 100 m [BENTHAUS: stn DW 1939], 7.3 x 5.6 mm. 14 C. VlLVENS NOVAPEX 13(1): 1-23, 10 mars 2012 15 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Teleoconch up to 6.2 convex whorls, bearing 8 spiral granular cords, in two areas of 6 and 2 cords, separated by a weak angulation; most adapical and two most adapical cords granular, other cords nearly smooth. Suture not canaliculated, poorly visible. First whorl convex, sculptured by 4 primary spiral cords poorly marked, more or less similar in size. On second whorl, spiral cords wider, very low, distance between cords much smaller than cords; P4 slightly stronger, weakly subgranular; other cords smooth; S3 appearing at end of whorl. On third whorl, angulation appearing between S3 and P4; S3 quickly similar to P3. On fourth whorl, ail cords more elevated; S2 appearing at begin of whorl, quickly similar to P2; P4 dividing into two cords, subgranular at mid whorl, adapical cord slightly thinner; distance between S3 and two abapical cords greater than distance between other cords; SI appearing at end of whorl by séparation from PI. On fifth whorl, PI and SI weakly subgranular; two P4 clearly granular with round beads. On last whorls, four adapical cords granular with prosocline elongated beads; P3 and S3 still nearly smooth; two P4 roundly granular, similar in size, producing a week peripheral keel; Tl appearing under Discussion. Regarding its small size (the well formed columellar tooth implies that the studied specimens are, at least, shells of young adults) and the general shape ot its shell, the new species can only be compared to similar in size known species from Philippines or French Polynesia. Among them, Thcilotia polysarchosa n. sp. is rather close to Jujubinus geographicus Poppe, Tagaro & Dekker, 2006 (Figs 35-36 ) from Philippines and French Polynesia (new record in this paper), but this similar S3; T2 possibly appearing between two P4 on large specimens. Aperture subelliptic, slightly prosoclinely elongated on large specimens; outer lip with a weak peripheral angulation and meeting inner lip with obtuse angle. Columella straight, slightly opisthocline to vertical, with one blunt tooth at abapical third. Base moderately convex, with 12 nearly smooth spiral cords; distance between cords smaller than cords; size of cords decreasing from outer part of base to inner part. Umbilicus moderately wide (diameter about 15-16% of shell width) and deep, partly covered by expansion of columella, funnel shaped with thin axial ridges inside. Colour of protoconch pinkish white to deep pink; teleoconch colour pattern rather variable : ground colour reddish brown to orange brown or pinkish brown, irregular axial white, light brown or pinkish patches; periphery altemating orange and pinkish white dashes; umbilical area and inner half of base nacreous pink or light red, spiral cords of outer half altemating pink and red or orange and pink, or almost uniformly pink. in size species is slightly more elevated, has a more conical shape, 7 (not 8) main granular spiral cords ail with rounded beads, a more pronounced suprasutural keel and a subcircular aperture without transversal élongation. Etymology. Obese (Greek : Tco^uaapxoQ, with reference to the cyrtoconoidal shape of the moderately elevated shell. TW H W HA H/W H/HA holotype 6.2 4.5 4.0 1.3 1.13 3.46 paratype 1 6.0 3.7 3.2 1.3 1.16 2.85 paratype 2 6.0 4.6 3.8 1.2 1.21 3.83 paratype 3 5.9 3.9 3.3 1.1 1.18 3.55 Table 4. - Thalotiapolysarchosa n. sp. : Shells measurements in mm for types. 45-50. Thalotia polysarchosa n. sp. 45-48. Australes Archipelago, 100 m rBENTHAUS sfn nwi rcmi- dï dh Holotype MNHN (24974). 4.5 x 4.0 mm. 47-48. Paratype MNHN(24975 3 7 x 3 2 mm49 Z f K Islands, 440 m [TARASOC, stn DW3482], 5.2 x 4.1 mm. 1 * 12 ^ S ° C ' ete Marshall, 1995. 51-52. Tarava Seamounts 670-757 m DW809], 1 l.èT\S2m^' '°' 9 X 101 mm; 53 ' 54> Southem New Caledonia, 650-730 m [BATHUS 3, stn ^^r^s\'^r < i.ri i ‘ ,,ype mn ™ ^ t “ 3 ■ «4» m s 5 - sSrstom para,ype mnhn - New ca " donia - a, °" * s " rprise - 16 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 17 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Genus Calliotrochus Fischer, 1879 Type species: Turbo phasianellus Deshayes, 1863 (by monotypy) = Margarita marmorea Pease, 1861 - Recent, Indo-Pacific. Calliotrochus marmoreus (Pease, 1861) Margarita marmorea Pease, 1861: 435. Type locality: Sandwich Islands. Turbo phasianellus - Deshayes, 1863: 74, n°216. Gibbula marmorea - Kay, 1979: 51, fig 14-E. Calliotrochus marmoreus - Severn, 2011: 48, pl. 8, fig T Material examined. French Polynesia, Tarava Seamounts. TARASOC: stn DW3302, 19°15'S, 150°57'W, 600-660 m, 9 dd, 2 dd juv. - Stn DW3316, 19°14'S, 151°33'W, 519-520 m, 5 dd juv. - Stn DW3318, 19°15'S, 151°3FW, 557-569 m, 1 dd juv. - Stn DW3333, 18°45'S, 152°18'W, 795-975 m, 1 dd juv. - Stn DW3336, 18°23’S, 154°06’W, 573-619 m, 9 dd. - Stn DW3337, 18°23'S, 154°05'W, 571-614 m, 1 dd juv. French Polynesia, Tuamotu Archipelago. TARASOC: stn DW3349, 15°05'S, 148°03'W, 976-997 m, 1 dd juv. French Polynesia, Société Islands. TARASOC: stn DW3420, 16°46'S, 151 o 04'W, 550 m, 1 dd juv. Distribution. Indo-Pacific (from northern Zululand to Hawaii and French Polynesia), shallow waters (Herbert, 1998); Hawaiian Islands, intertidal (Severn, 2011); French Polynesia, Tarava Seamounts, 520-795 m (dead); French Polynesia, Tuamotu Archipelago, 976-997 m (dead); French Polynesia, Société Islands, intertidal (living), 10-550 m (dead) (using data from Letoumeux et al., p.c.). Family CALLIOSTOMATIDAE Thiele, 1924 Subfamily CALLIOSTOMATINAE Thiele, 1924 Genus Calliostoma Swainson, 1840 Subgenus Fautor Iredale, 1924 Type species : Ziziphinus comptus A.Adams, 1855 (by o.d.) - Recent, Southern Australia. Calliostoma (Fautor) paradigmatum Marshall, 1995 Figs 51-54 Calliostoma (Fautor) paradigmatum Marshall, 1995b: 395-397, figs 13-15, 119, 155. Type locality: Southern New Caledonia, 505-550 m. Calliostoma (Fautor) paradigmatum - Vilvens, 2005: 2 . Calliostoma (Fautor) paradigmatum — Vilvens, 2009a: 132, figs 25-26. Material examined. Tarava Seamounts. BENTHAUS: stn DW3300, 19°19 f S, 151°00'W, 670- 757 m, 1 dd. - Stn DW3302, 19°15'S, 150°57'W, 600- 660 m, 2 dd juv. Distribution. Off Ile Surprise, northern New Caledonia, 585 m (living); South of lie des Pins, Southern New Caledonia, northern Norfolk Ridge, 470-795 m, living at 550-795 m (range computed using data of Marshall, 1995); Tonga, 342-500 m (dead); Tarava Seamounts, 660-670 m (dead). Comments. Regarding the huge gap between the type locality (Southern New Caledonia) and French Polynesia, one can wonder to find this species on Tarava Seamounts stations, although some dead specimens were found in Tonga Archipelago (Vilvens, 2005). But the BENTHAUS adult shell shows ail the characteristics of the New Caledonian species, especially the uniform white colour, the saine ratio H/D (about 1.08), a spiral cord Pl commencing only at 2 nd whorl, S2 appearing at 3 rd whorl and SI appearing at 4 th whorl, a base with 14 spiral cords with the innermost stronger and granular. The only différences for the Polynesian samples are a spiral cord P4 emerging later (instead of présent immediately but partly covered by next whorl), a spiral cord S3 absent (but this can happen, flde original description) and a smaller protoconch (only less than 300 pm for the Polynesian adult specimen). Calliostoma (Fautor) lepton n. sp. Colour Figs C1-C2, Figs 55-56 Type material. Holotype (12.6 x 10.8 mm) MNHN (24976). Type locality. French Polynesia, Tuamotu Archipelago, TARASOC, stn DW3370, 15°39'S, 146°52'W, 315 340 m. Material examined. Tuamotu Archipelago. TARASOC: stn DW3370, 15°39'S, 146°52'W, 315-340 m, 1 lv (holotype). Distribution. French Polynesia, Tuamotu Archipelago, 315-340 m (dead). Diagnosis. A rather small yellowish light brown Calliostoma species with a moderately elevated spire, coeloconoidal in shape in upper part and cyrtoconoidal in lower part, with a subangular periphery; aperture and columella highly nacreous; 6 main granular spiral cords on last whorls, axial threads visible between cords; base moderately convex base with about 12 granular cords; no umbilicus. Description. Shell of rather small size for the genus (height 12.6 mm, width 10.8 mm), slightly higher than wide, rather thick, coeloconoidal in upper part, cyrtoconoidal in lower part, with a subangular periphery; spire moderately elevated, height 1.2x width, 3. lx aperture height; anomphalous Protoconch about 250-300 pm (apex eroded), with a 18 C. VlLVENS NOVAPEX 13(1): 1-23, 10 mars 2012 Teleoconch up to 6.7 convex whorls, with 6 main, granular, similar in size spiral cords and an additional peripheral cord on last whorl; beads of ail cords rounded conical, bluntly pointed. Suture not canaliculated, poorly visible. First whorl convex, sculptured by thick axial folds and 3 primary spiral cords PI, P2 and P3 more or less similar in size, rather low, granular by intersection with folds; distance between folds 1.5x size of folds; distance between cords similar in size to cords. On second whorl, spiral cords stronger and wider, with well rounded beads; P3 slightly stronger than other cords, with rounded conical beads; axial folds more prosocline. On third whorl, beads of ail cords bluntly pointed; P4 emerging from suture at end of whorl, beaded, weaker than other cords. On fourth whorl, P4 stronger, similar in size and shape to other cords; distance between P4 and P3 smaller than cords, distance between other cords similar in size to cords; S2 appearing at end whorl, thinner than other cords; axial folds weakening into axial threads. On fifth whorl, SI appearing at mid whorl; S3 absent. On last whorls, ail cords similar in size except P4 slightly stronger; ail cords with conical, blunt pointed beads; axial threads weak but still visible; Tl appearing between P2 and S2; S4 visible on last whorl, peripheral, thinner than other cords; distance between ail Pi and Si similar in size to cords, except S4 doser to P4; T2 appearing between SI and P2. Aperture subelliptic, slightly prosoclinely elongated; outer lip with nacreous thickened inside, with fine ridges at rim, without angle at meeting with thick inner. Columella weakly arcuated, slightly oblique, thick and nacreous, flaring at lower part, without tooth. Base moderately convex, with 12 granular spiral cords; subquadrate beads of cords made by thin axial threads across base; distance between cords smaller than cords; size of cords increasing in width from outer part of base to inner part. No umbilicus (closed by columellar expansion). Colour of protoconch and teleoconch yellowish light brown; base shiny white; aperture and columella nacreous. Discussion. The new species is rather close to Calliostoma (Fauîor) necopinatum Marshall, 1995 (Figs 57-58) from New Caledonia, but this similar in size species has a different, narrowly conical, weakly cyrtoconoidal shape, more convex last whorls, a spiral cord S3, a wider protoconch (400-420 pm) and a subquadrangular thin aperture without ridges inside. Etymology. Flazelnut (Greek : Z£7rcov), used in apposition as a noun - with reference to light hazelnut colour of the shell, reminding of a eut hazelnut. Genus Thysanodonta Marshall, 1988 Type species : Thysanodonta aucktandica Marshall, 1988 (by o.d.) - Recent, New Zealand. Thysanodonta cf aucklandica Marshall, 1988 Figs 59-60 Thysanodonta aucklandica Marshall, 1988: 217-219, figs 3A-C. Type locality: off Auckland Islands, New Zealand, 549 m. Material examined. Tuamotu Archipelago. TARASOC: stn DW3389, 14°55'S, 148°15'W, 889 m, 1 dd. Comments. This single specimen (6.8 x 5.5 mm) has the spiral cords ontogeny of T. aucklandica (P1-P2- S2-P3-S4 with P4 peripheral on last whorl), but has only 7 spiral cords on the base (instead of 10 fide original description Marshall, 1988). This specimen, with First whorls rather eroded and an indistinct suture, could be subadult because its teleoconch has only 5.3 whorls. Additional material is clearly needed to décidé if this is a different species or not. Family TllRBINIDAE Rafinesque, 1815 Subfamily MARGARIT1NAE Thiele, 1924 Genus Gaza Watson, 1879 Type species: Gaza daedala Watson, 1879 (by o. d.) - Recent, Fiji Islands. Gaza p oly ch oronos n. sp. Colour Figs B1-B2, Figs 61-67, Table 5 Type material. Holotype (14.5 x 18.4 mm) MNHN (24977). Paratype (12.9 x 16.5 mm) MNHN (24978). Type locality. French Polynesia, Société Islands, Bora Bora, TARASOC, stn DW3418, 16°33’S, 151°48'W, 580-618 m. Material examined. French Polynesia, Société Islands. TARASOC: stn DW3418, 16°33'S, 151°48'W, 580-618 m, 1 lv (holotype). - Stn DW3497, 17°43'S, 149°14'W, 365-850 m, 1 lv (paratype). Distribution. French Polynesia, Société Islands, living at 580-618 m. Diagnosis. A rather small white Gaza species with a moderately elevated, more or less conical spire, a rounded periphery; First whorls smooth, last whorls with numerous spiral cords, granular on adapical half and almost smooth on abapical half; granular callus covering a fairly wide umbilicus. Description. Shell of rather small size for the genus (height up to 14.5 mm, width up to 18.4 mm), wider than high, thin, conical, with a rounded periphery; spire moderately elevated, height 0.7x to 0.8x width, 1.8x to 2.2x aperture height; fairly broad umbilicus. Protoconch of about 200-250 jum, of 1.25 whorl, low, smooth, glossy, without terminal varix. 19 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia Teleoconch up to 5.8 convex whorls, bearing numerous spiral, similar in size cords; adapical cords granular, abapical cords smooth; on last whorls, thin, prosocline axial threads between spiral cords. Suture visible, not canaliculated. First whorl convex, almost smooth with growth Unes poorly visible. On second whorl, low, poorly marked, axial folds appearing under suture. On third whorl, axial folds stronger, weakly prosocline, ending on adapical part with a rounded bead; 4 smooth, very thin, spiral cords appearing on abapical half. On fourth whorl, axial folds thinner, becoming thin threads, more numerous and more prosocline, covering ail adapical half; subsutural beads resolving in a subsutural, roundly granular, spiral cord; at begin of whorl, 4 thin spiral cords appearing on adapical half, increasing in number by intercalation under suture, giving a reticulate pattern by intersection with axial threads; abapical cords nearly smooth to subgranular. On last whorls, number of spiral cords reaching 16 to 18; cords evenly spaced, distance between cords similar in size or slightly greater than cords, very different in shape on the two halves: cords of abapical half rather thin, almost smooth or slightly subgranular, elevated, with thin axial threads between them; cords of abapical half thicker, lower, granular, Aperture well rounded, slightly deflected on adult shell, with srnall, inner denticles ail around outer and inner lip; outer lip slightly flared at rim, inner lip meeting outer lip with a weak angle on subadult specimen. Columella arched, without tooth, with a basal flaring on subadult sample. Base convex, with up to 30 spiral cords; up to 50 axial threads around umbilicus; distance between spiral cords smaller than cords; cords of two outermost thirds smooth, thin; cords of innermost third wider, granular by intersection with axial threads. Umbilicus fairly wide (diameter 9% to 10% of shell width), deep, funnel shaped, with steep sloping, slightly concave walls, covered by a moderately thin columellar callus (broken on paratype) covered by numerous, irregular in size and shape granules; thin, weak axial threads inside. Colour of teleoconch nacreous white with pinkish sheen; protoconch pinkish brown (holotype). TW H W HA H/W H/HA holotype 5.8 14.5 18.4 6.7 0.79 2.16 paratype 5.6 12.9 19.5 7.1 0.66 1.82 Table 5. - Gazapolychoronos n. sp.: Shells measurements in mm for types. Discussion. The new species is provisionally placed into the Gaza genus, regarding its spiral sculpture, its rounded, prosocline aperture and the umbilical callus, although it has an only weakly descending suture and a poorly reflected peristome (features used by Simone, 2006). The new species is rather close to Gaza daedala Watson, 1879 (Figs 68-69) from Fiji, but this similar in size species has much thinner, more numerous spiral cords on the whorls (25 spiral lines in penultimate whorl), has a higher H/W ratio (1.24), a body whorl with a weak spiral carina between middle and lower thirds and a smooth, not granular callus. Etymology. Bead-shaped object (Greek : xopovoQ and numerous (Greek : ttoAajÇ, àu), used as a noun in apposition - with reference to the granular umbilical callus. Acknowledgements I would like to thank P. Bouchet (Muséum national d'Histoire naturelle, Paris) for reading the manuscript, constructive advice and access to the malacological resources of the MNHN, and V. Héros (MNHN) for her help in finding various scientific papers. Also, I am very grateful to A. Salvador (Natural History Muséum, London) for her help for her help to search material in the NHMUK collections and to get photographs of types and information about them. T also would like to thank G. Poppe and J. Sarino (Conchology, Inc., Philippines) for the kind permission to use the types photographs they hâve sent. Figures 59-69. Scale bars = 5 mm. 59-60. Thysanodonta cf aucklandica Marshall, 1988, Tuamotu Archipelago, 889 m [TARASOC stn DW33891 6.8 x 5.5 mm. nwi Caypofyctonmos n. sp., Société Islands. 61-64. Holotype MNHN (24977), 580-618 m [TARASOC, stn n-A n AO^ 1 " 63 ' l4 ' 5 X 18,4 mm; 64 ‘ Detail of the granular callus. 65-67. Paratype MNHN (24978), 520-572 m [TARASOC, stn DW3497], 12.9 x 16.5 mm. 68-69. Gaza daedala Watson, 1879, holotype NHMUK (1887-2-9-346), Fiji, Kandavu, 1128 m, 20.6 x 17.0 mm lotographs courtesy of Phil Crabb, NHMUK Photographie Unit. 20 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 21 C. VlLVENS Seguenzioidea and Trochoidea from French Polynesia REFERENCES |papers and books ] Adams, A. 1853. Contribution towards a monograph of Trochidae, a family of gastropodous Mollusca. 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New species and new records of Calliostomatidae (Gastropoda: Trochoidea) from New Caledonia and Solomon Islands. Novapex 10(4): 125-163. Vilvens, C. 2009b. A new species of Calliostoma (Gastropoda: Trochoidea: Calliostomatidae) from Tahiti. Novapex 10(3): 109-113. Williams, S. T., Karube, S. & Ozawa, T. 2008. Molecular systematics of Vetigastropoda: Trochidae, Turbinidae and Trochoidea redefined. Zoologica Scripta 37(5): 483-506. Williams S.T., Donald K.M., Spencer H.G. & Nakano T. 2010. Molecular systematics of the marine gastropod families Trochidae and Calliostomatidae (Mollusca: Superfamily Trochoidea). Molecular Phylogenetics and Evolution 54: 783-809. Wilson, B. 1993. Austral ian Marine Shells. Prosobranch gastropods — part one. Odyssey Publishing, Kallaroo, Western Australia. 408 pp. 44 plates. 22 C. VlLVENS Novapex 13(1): 1-23, 10 mars 2012 [web sites ] Rapport d'activité IRD http://www.brest.ird.fr/us 191 /flotte/rapports/ activite_flotte_2009.pdf (access on 2011 -12-31 ) Census of Marine Life on seamounts (CenSeam) program http://censeam.niwa.co.nz/ (access on 2012-01-08) IFREMER - TARASOC campaign http://www.ifremer.fr/sismer/FR/catal/campagne/ campagne.htql?crno=9100040 (access on 2011-12- 31) IFREMER - BENTHAUS campaign http://www.ifremer.fr/sismer/FR/catal/campagne/ campagne. htql?cmo=2100100 (access on 2011 -12-31 ) Marine Barcode Of Life project. http://www.marinebarcoding.org/ WoRMS (World Register of Marine Species) http://www.marinespecies.org (access on 2011-12- 31) 23 R. HOUART Novapex 13(1): 25-28, 10 mars 2012 Description of a new species in the Siratus pliciferoides group (Gastropoda: Muricidae) from the Philippines Roland HOUART Research Associate Institut royal des Sciences naturelles de Belgique Rue Vautier, 29, B-1000 Bruxelles, Belgium roland.houart@skynet.be KEYWORDS. Philippines, Gastropoda, Muricidae, Siratus n. sp. ABSTRACT. A new species of Siratus is described from Balut Island, Philippines. It is compared with Siratus pliciferoides (Kuroda, 1942). The synonymy of Siratus pliciferoides is reviewed and commented. RESUME. Une nouvelle espèce de Siratus est décrite de l'île de Balut aux Philippines. Elle est comparée à Siratus pliciferoides (Kuroda, 1942). La synonymie de Siratus pliciferoides est passée en revue et commentée. INTRODUCTION The Siratus pliciferoides group contains four names usually considered to be monospecific: Chicoreus pliciferoides (Figs 4-7, 15) was described by Kuroda (1942: 81) as a substitute name for Murex pliciferus Sowerby, 1841 which was preceded by, and thus a homonym of, M. pliciferus Bivona-Bemardi, 1832. The species occurs widely in an area consisting of the Philippines, the China Seas, Taiwan and Southern Japan. Murex (Siratus) propinquus was described by Kuroda & Azuma in Azuma (1961: 300) for a quite similar shell from Japan but with a comparatively lower spire, a shorter siphonal canal and a broader last teleoconch whorl. This form is currently known from Western Australia as well as from the Philippines, Taiwan and Japan (Figs 8-10). A third name, Siratus hirasei was given by Shikama (1973: 5) who described another form from Japan, with broad varices, short spines and broad, straight, short siphonal canal as illustrated here (Figs 11-12). Finally, S. vicdani was described by Kosuge (1980: 55) who separated it from both S. alabaster (Reeve, 1845) and S. pliciferoides. It differs from the latter by having angulate shouldered spire whorls and more strongly webbed varices (Figs 13-14). Siratus pliciferoides being the oldest available name, ail the other names were considered objective junior synonyms by Vokes (1971: 86), Fair (1976: 69) and Radwin & D’Attilio (1976: 107), for Siratus propinquus only, the other shells having been described after Vokes (1971). Fair (1976) and Radwin & D’Attilio (1976) probably were not aware of the description of Siratus hirasei in a little-known Japanese publication. Houart (1992: 110) and Merle et al (2011: 100) also considered ail these names as being conspecific. Since 1992, 1 had the opportunity to examine numerous specimens of ail these forms from the geographical distribution area and 1 am less certain about some of these names being synonym of S. pliciferoides. Maybe eventual DNA researches will confirm or invalidate the validity of these names. Another new species, probably part of this group, is here described for the first time from the Philippines. Abbreviations Repository EGS coll. : Collection of Evelyn Guillot de Suduiraut IRSNB: Institut royal des Sciences naturelles de Belgique, Bruxelles, Belgium. MNHN: Muséum national d'Ffistoire naturelle, Paris, France. RH coll. : Collection of the author. Terminology used to describe the spiral cords (after Merle, 1999 and 2001) (Text Fig. 1) P: primary cord; s: secondary cord; t: tertiary cord; ad: adapical (or adapertural); ab: abapical (or abapertural); IP: infrasutural primary cord (primary cord on subsutural ramp); adis: adapical infrasutural secondary cord (on subsutural ramp); abis: abapical infrasutural secondary cord (on subsutural ramp); PI: shoulder cord; P2-P6: primary cords of the convex part of the teleoconch whorl; sl-s6: secondary cords of the convex part of the teleoconch whorl (example: si = secondary cord between PI and P2; s2 = secondary cord between P2 and P3, etc.); ADP: adapertural primary cord on the siphonal canal; MP: médian primary cord on the siphonal canal; ABP: abapertural primary cord on the siphonal canal; ads: adapertural secondary cord on the siphonal canal; ms: médian secondary cord on the siphonal canal; abs: abapertural secondary cord on the siphonal canal. 25 R. Houart Siratus evelynae n.sp. Text Fig. 1. Spiral cords terminology (paratype MNHN) SYSTEMATICS Family MURLCIDAE Rafinesque, 1815 Subfamily MURICINAE Rafinesque, 1815 Genus Siratus Jousseaume, 1880 Type species, by original désignation: Purpura Sirat "Adanson" Jousseaume, 1880 (= Murex senegalensis Gmelin, 1791), Recent, Brazil. Siratus evelynae n. sp. Text Figs 1-2, Figs 1-3 Type material. Philippines, Digos, Davao del sur, coral rubble, 150-200 m, by tangle nets, holotype IRSNB IG.32073/MT2574. Paratypes: Philippines, Balut Island, south of Mindanao, in 400 m, by tangle net, 1 MNHN IM- 2010-19527; Digos, Davao, in 400 m, by tangle nets, 2 RH coll.; Davao, coral rubble, sandy mud, 200-400 m, by tangle net, 1 RH coll, 1 EGS coll. Distribution. Currently only known trom the type material: Philippines, south of Mindanao, live in 150- 400 m, by tangle nets. Description. Shell medium sized foi the genus, up to 112 mm in height (paratype EGS). Height/width ratio 2.0-2.2. Slender, biconical, broadly ovate, heavy, very weakly spinose, tuberculate. Subsutural ramp broad, strongly sloping, convex. Creamy white or light tan with tan, brown or dark brown spiral cords. Aperture white. Spire high, acute, with a teleoconch of 8 broadly convex, weakly shouldered, nodose whorls. Suture weakly adpressed. Protoconch unknown. Axial sculpture of teleoconch whorls consisting of high, broad, rounded ribs and high, narrow, rounded varices. Two first teleoconch whorls eroded. Three varices and two intervarical broad ribs from third to last whorl. Last whorl occasionally with two broad and one narrow ribs on last portion of whorl, between second and apertural varix. Spiral sculpture of low, narrow, primary, secondary, tertiary cords and narrow threads. Spiral sculpture of shoulder ramp of last teleoconch whorl consisting of adis, IP and abis with a tertiary cord between each. Convex part of last teleoconch whorl of PI (shoulder cord), t, si, t, P2, t, s2, t, P3, t, s3, t, P4, s4, P5, s5, P6, t, s6, t, ABP, abs, MP, ms, ABP, abs. Few threads and secondary cords occasionally of same strength and height; occasionally with a few additional narrow threads. Aperture moderately large, roundly ovate. Columellar lip naiTOw, smooth, with strong, low pariétal tooth at adapical extremity. Lip adhèrent. Anal notch deep, narrow. Outer lip erect, crenulated with numerous, elongate, narrow denticles within, many of them split. Siphonal canal long, 39-43% of total shell length., broad adapically, abruptly tapered and very narrow abapically, weakly recurved dorsally. Operculum dark brown, ovate, inverted tear-shaped, with apical nucléus and 9 or 10 concentric ridges at outer surface. Attached surface with 10 growth lines and broad, callused rim. Radula unknown. Figures 1-15 1-3. Siratus evelynae n. sp. 1-2. Philippine, Digos, Davao del sur, coral rubble, 150-200 m, holotype IRSNB IG.32073/MT2574,83.3 mm; 3. Balut Island, south of Mindanao, in 400 m, paratype MNHN IM-2010-19527, 81.7 mm 4-7. Siratus pliciferoides (Kuroda, 1942). 4-5. Northeast of Taiwan, RH, 134 mm; 6. Philippines, Bohol, RH, 128.4 mm; 7. Japan, Tosa, RH, 113.5 mm; 8-10. Siratus pliciferoides "form" propinquus Kuroda & Azuma, 1961; 8-9. Northeast of Taiwan, RH, 106.4 mm; 10. West Australia, Port Hedland, RH 93 2 mnr 11-12 Siratus pliciferoides "form" hirasei Shikama, 1973, Japan, Minabe, RH, 96.1 mm; 13-14. Siratus pliciferoides "form" vicdani Kosuge, 1980, Philippines, Mactan, RH, 38.4 mm; 15. Siratus pliciferoides (Kuroda 19421 Philippines, juvénile, RH, 38.2 mm. h 26 R. HOUART Novapex 13(1): 25-28, 10 mars 2012 27 R. Houart Siratus evelynae n.sp. Remarks. Siratus evelynae is probably related to the Siratus pliciferoides group, however it differs consistently from S. pliciferoides and related forms in being more strongly biconical, in having a last teleoconch whorl with a more sloping shoulder ramp, giving the shell a more rounded shape vs shouldered in S. pliciferoides and related forms, and in starting a broad siphonal canal but promptly tapering to become very narrow at its abapical extremity. Siratus evelynae also differs in having a spineless shell except for a sériés of short, acute, broadly open, webbed spines abapically. The new species also has a lower spire compared to the shell height, being ot 27- 28% of total shell height in S. evelynae n. sp. vs 37- 39% in S. pliciferoides and 41-46% in Siratus propinquus or the "propinquus" form, whatever it may be. The two other names are strongly related to S. pliciferoides and do not need to be compared here, although illustrated (Figs 11-14). Etymology. I am very happy to dedicate this new species to Evelyn Guillot de Suduiraut, wife of the late well-known Emmanuel Guillot de Suduiraut. Acknowledgements I am greatly indebted to Evelyn Guillot de Suduiraut and to her daughter Jackylen to having brought my attention to this interesting species and for the gift of the holotype. Many thanks also to John Wolff, Lancaster, Pennsylvania, USA, for checking the English text and to Claude Vilvens, Oupeye, Belgium for critical comments. REFERENCES Azuma, M. 1961. Descriptions of six new species of Japanese marine Gastropoda. Venus 21(3): 296-303. Fair, R.Fl. 1976. The Murex Book, an illustrated catalogue of Recent Mûrie idae (Muricinae, Muricopsinae, Ocenebrinae), Sturgis Printing Co., Honolulu, Hawaii: 1-138. Flouart, R. 1992. The genus Chicoreus and related généra (Gastropoda: Muricidae) in the Indo-West Pacific. Mémoires du Muséum national d'Histoire naturelle , (A), 154: 1-188. Kosuge, S. 1980. Descriptions of three new species of the Family Muricidae (Gastropoda: Muricacea). Bulletin of the Institute of Malacology, Tokyo 1(4): 53-58. Kuroda, T. 1942. Two Japanese murices whose names hâve been preoccupied. Venus 12(1-2): 80-81. Merle, D. 1999. La radiation des Muricidae (Gastropoda : Neogastropoda) au Paléogène: approche phylogénétique et évolutive. Paris. Unpublished thesis, Muséum national d'Histoire naturelle : i-vi, 1-499. Merle, D. 2001. The spiral cords and the internai denticles of the outer lip in the Muricidae: terminology and methodological comments. Novapex 2 (3): 69-91. Merle, D., Garrigues, B. & Pointier, J.P. 2011. Fossil and Recent Muricidae of the World -Part Muricinae- Ed. Conchbooks, D-55546 Hackenheim: 1-648. Radwin G. & D'Attilio, A. 1976. Murex shells of the world. An illustrated guide to the Muricidae. Stanford University Press, Stanford: 1-284. Shikama, T. 1973. Description of new marine Gastropoda from the East and South China Seas. Science Reports oj the Yokohama National University 20: 1-8. Vokes, E.H. 1971. Catalogue of the genus Murex Linné (Mollusca: Gastropoda. Muricinae, Ocenebrinae. Bulletin of American Paleontology 61 (268): 1-141. 28 E. Rolan & F. Rubio NOVAPEX 13(1): 29-32, 10 mars 2012 A new species of the genus Leucorhynchia (Gastropoda, Turbinidae) from West Africa Emilio ROLÂN Museo de Historia Natural, Campus Universitario Sur, 15782, Santiago de Compostela, erolan@emiliorolan.com Federico RUBIO Pintor Ribera, 4-16 a , 46930 Quart de Poblet (Valencia) federubio@ono.com KEYWORDS. Turbinidae, Leucorhynchia , West Africa, new species. ABSTRACT. A new species of Leucorhynchia is described from Principe Island, West Africa. It is compared with L. lirata (E.A. Smith, 1871). INTRODUCTION The genus Leucorhynchia Crosse, 1867 was considered of Indo-Pacific distribution until it was reported from the West African coast by Adam & Knudsen (1969), who described a new species: Leucorhynchia bicarinata and figured two more species previously included in the genus Ethalia : Leucorhynchia plicata (Smith, 1871) and L. lirata (E.A. Smith, 1871). Rubio & Rolan (1991) illustrated specimens of Leucorhynchia bicarinata and L. lirata collected in Sâo Tomé and Principe, giving from the latter one drawing of the soft parts and SEM photographs of the protoconch and radula. At the same time they included in the same genus L. punctata (Jousseaume, 1872), previously placed in the genus Teinostoma A. Adams, 1853. So far, there are four recognized species within the genus Leucorhynchia: L. caledonica Crosse, 1867 (type species of the genus), of Indo-Pacific distribution, the other three having a West African distribution: L. punctata Jousseaume, 1872, L. lirata (E.A. Smith, 1871) and L. bicarinata Adam & Knudsen, 1969 In the sédiments recently collected at the island of Principe by the ïtalian malacologist Sandro Gori, during a visit to the islands of Sào Tomé and Principe, some shells appeared to belong to a new species, which is described in the présent work. SYSTEMATICS Family TURBINIDAE Rafinesque, 1815 Subfamily SKENEINAE Clark, 1851 Genus Leucorhynchia Crosse, 1867 Type species: Leucorhynchia caledonica Crosse, 1867. Recent. New Caledonia. Leucorhynchia gorii spec. nov. Figs 6-7, 9, 11 Type material. Holotype (Fig. 7) deposited in the Muséum National d’Histoire Naturelle, Paris (MNHN 24633). Paratypes: Museo de Ciencias Naturales of Madrid (MNCN 15.05/60001,1 shell) (Fig 6); Museo de Historia Natural de Santiago de Compostela (MHNS 100556, 1 shell); collection of Sandro Gori (1 shell). Type locality. Tinhosa Pequena, 25 m, Island of Principe, Republic of Sâo Tomé and Principe. Distribution. Only known from the type locality. Description. Shell minute, lenticular, solid, and cream in colour. Protoconch with only one smooth whorl and a diameter of 227-230 pm. Teleoconch with a little over two whorls, almost lacking sculpture at the beginning, but on which some sulci shortly appear; they are undulating at first and after the first whorl are of circular shape; they become more irregular and on the last half-whorl they turn into smooth cords, a little irregular, separated by spaces almost of a similar width and where it is possible to see under magnification, tubercles and lines in an axial direction are observed. In the ventral side there is a well opened umbilicus, within which there are four prominent spiral sulci. In the border of the umbilical infundibulum short axial ribs are formed. The spire ends in a circular and very regular aperture with a narrow and free peristome, which extends until getting in contact with the previous whorl. Dimensions. Holotype 1.5 mm in diameter; paratypes are of a similar size. 29 E. Rolan & F. Rubio A new Leucorhynchia from West Africa Discussion. Leucorhynchia gorii spec. nov. shows an apparent similarity with L. lirata , because of its rounded form, its small protoconch, the beginning of the spiral sculpture with cords with dépréssions, which later disappear. But L. lirata has a larger shell (up to 3.5 mm, as opposed to 1.6 mm the new species may reach). The nurnber of whorls in the teleoconch of an adult L. lirata is about 3 !4, while L. gorii has no more than 2 l A whorls. The spiral whorls which form the sculpture of the teleoconch, after the First whorl, hâve smaller interspaces in L. lirata and wider in L. gorii. The most important différence between these two species is on the base where L. lirata has a very close umbilicus while that of L. gorii is wider showing the spiral cords on its inner part. The juvéniles of L . lirata hâve a wider umbilicus but are similar in size to L. gorii ; the aperture of L. lirata has quicker development and the umbilicus is bordered by very different elevated cords. Etymology. The new species is named after Sandro Gori who collected the sédiments where the shells were found. ACKNOWLEDGEMENTS To Jésus Méndez of the Centro de Apoyo Tecnolôgico a la Investi gaciôn (CACTI) of the University of Vigo, where the SEM photographs were made. Antonio A. Monteiro revised the English. REFERENCES Adam, W. & Knudsen, J., 1969. Quelques genres de mollusques prosobranches marins inconnus ou peu connus de l’Afrique occidentale. Bulletin Institut royal des Sciences naturelles de Belgique , 44(27): 1-69. Rubio, F. & Rolan, E., 1991 “1990”. Aportaciones a los conocimientos sobre los micromoluscos de Âfrica Occidental. 2. Archaeogastropoda de Sâo Tomé y Principe. Iberus , 9(1-2): 209-219. Figures 1-7 1-5. Leucorhynchia lirata Adam & Knudsen, 1969. 1-2. Adult shell, 2.9 x 3.0 mm, Tinhosa Pequena, Principe I.; 3-5. Juvéniles, 1.9, 1.8 and 1.54 mm Tinhosa Pequena. 6-7. Leucorhynchia gorii spec. nov. 6. Paratype, 1.6 mm (MNCN 15.05/60001); 7. Holotype, 1.5 mm (MNHN 24633). 30 E. Rolan & F. Rubio Novapex 13(1): 29-32, 10 mars 2012 31 — E. Rolan & F. Rubio A new Leucorhynchia from West Africa Figures 8-11 8-19. Protoconchs of Leucorhynchia species 8. Leucorhynchia lirata Adam & Knudsen, 1969; 9. Leucorhynchia gorii spec nov 10-11 . Microsculpture of Leucorhynchia species 10. Leucorhynchia lirata Adam & Knudsen, 1969; 11. Leucorhynchia gorii spec nov 32 D. P. ClLIA Novapex 13(1): 33-36, 10 mars 2012 A new Javan species of Agaronia Gray, 1839 (Neogastropoda, Olividae) David P. CILIA 29, Triq il-Palazz 1-Ahmar, Santa Vénéra SVR1454, Malta dpcilia@gmail.com. KEYWORDS. Olividae, Indian Océan, Java, Indonesia, Agaronia johnabbasi sp. nov. ABSTRACT. A new species of olivid neogastropod from West Java, Agaronia johnabbasi sp. nov., is described according to conchological characters. It is distinguished from congeners by means of its distinctive morphology and colouration. INTRODUCTION The highly evolved gastropods in the family Olividae are found circumglobally in predominantly tropical or subtropical waters. Ail inhabit soft substrates including sand and silt, though the depths at which different species may be found vary from the littoral to the sublittoral. The genus Agaronia Gray, 1839 is sometimes regarded as forming part of the subfamily Agaroniinae Olsson, 1956 (Olsson, 1956; Ponder & Warén, 1988; Vaught, 1989; Sterba, 2004); though in Bouchet & Rocroi (2005), this taxon is synonymized with Olivinae Latreille, 1825. The majority of species are concentrated along the west African and east American coasti ines, with some Indian Océan représentatives that include the species described herein (see Table 1 for a complété list of recent species). Abbreviations Descriptive: D - diameter; H - height. Repositories: DC: collection of author, Santa Vénéra, Malta; GP: collection of Guido T. Poppe, Cebu, Philippines; JA: collection of John Abbas, Jakarta, Indonesia; MNHN: Muséum national d’Histoire naturelle, Paris, France; NMNH: National Muséum of Natural History, Mdina, Malta. MATERIALS & METHODS Seven shells were collected by fishermen with bottom trawling nets from the eastem bay in Pangandaran, West Java, in late 2009, at a depth of about 60m. Their shell morphology was compared and contrasted with that of the five known sympatric congeners. Maximum diameter and height of the shells, the former incorporating the lip, were measured twice using a dial caliper with a resolution of 50pm, the average of the two measurements was then calculated and noted. SYSTEMATICS Family OLIVIDAE Latreille, 1825 Subfamily OLIVINAE Latreille, 1825 Genus Agaronia Gray, 1839 Type species by monotypy Voluta hiatula Gmelin, 1791, West Africa Agaronia johnabbasi sp. nov. Figs.1-6 Type material. Collected by fishermen, c. -60m on silty substrate, east Pangandaran Bay, Java, Indonesia, October 2009. Holotype MNHN 23267; paratypes: DC R.GA 1000-1; GP unreg.; JA unreg.; NMNH unreg. Type locality. East Pangandaran Bay, Java, Indonesia. Description. Dextral solid shell, of maximum height of about 40mm and a height/diameter ratio of about 2.8. Spire tall and conical, with a slightly concave profile and featuring a spiral callus occupying about half the height of each whorl. Widest band of the body whorl a uniform dark-orange brown, sometimes with a purplish tinge intensifying towards the suture. This colouration is continuons along the spire, contrasting with the orange-brown spiral callus. Thin wash of white callus présent on the columellar side of the aperture close to the filament channel. Just above the postfasciole, a thin pale band is discemible. The postfasciole and the fasciole are of the same colour and are considerably paler than the main part of the body whorl, with the former featuring a few very fine spiral threads and numerous darker growth striae which do not extend upon the fasciole. The belt and the base of the columellar callosity are beige. Outer lip thin, with its outline recurved, and, at the proximity of the siphonal canal, almost parallel with the columellar side of the aperture. Dimensions. See Table 2 for details. Remarks. In colouration, the new species is remarkable in its distinct lack of pattern (except for the growth lines typical of ail Agaronia) and, unusually for the genus, the colouration of the postfasciole and fasciole, which are lighter in colour than the main part of the body whorl. The type locality of the new species, Pangandaran Bay yields examples 33 D. P. ClLIA A new Javan species of Agaronia of ail other four Indian Océan représentatives of the genus (J. Abbas, pers. comm. XI 1.2009) (refer to Table 1). Of these, Agaronia nebulosa (Lamarck, 1811) may easily be distinguished by a reticulated pattern over a (generally) very pale background combined with a thick pariétal callosity, while Agaronia gibbosa (Born, 1778) features réticulation, a significantly lower H/D ratio, a strongly marked postfasciole and a thick pariétal callosity. The lighter, occasionally non-reticulated form flavescens Melvill, 1904 has a colour which is relatively close to that of A . johnabbasi, but the dimensions and the geographical provenance (South India) set it apart at first glance. Agaronia lutaria (Rôding, 1798) is larger and more slender, with a narrower aperture and strong réticulation. The différences from A. johnabbasi are herein illustrated by a juvénile specimen of the saine size as two of the paratypes (Figs 7-8). A doser relative of A. johnabbasi is Agaronia johnkochi Voskuil, 1990 (Figs 9-10). This is suggested by the absence of réticulation, a similarly curved outline of the outer lip, a similarly proportioned spire callus which does not bulge out disproportionately towards the aperture, and the strong démarcation of colour, in perpendicular direction to the filament channel, between the pigmented spiral and the whitish pariétal calluses. However, the spire of A. johnabbasi is higher and more acutely angled than that of A. johnkochi , while the spiral callus is lighter in colour, contrasting less with the rest of the whorl. The widest part of the shell of A. johnabbasi is doser to the apical part of the shell, the pariétal callus is thinner, and the oblique striae on the pillar structure are thinner and doser to each other, with no chromatic aberrations between them; ail these features are consistent and may be used for morphological différentiation between the two species. The gracile shell of A. johnabbasi is also similar to that of the West African Agaronia hiatula (Gmelin, 1791), but the outer lip is recurved and therefore the aperture is narrower. Etymology. The species is named after John Abbas (Jakarta, Indonesia) who initially recognized the species as a new one, kindly bringing it to the author s attention. ACKNOWLEDGEMENTS The author is indebted to John Abbas for the specimens sent for examination, including comparative material, Eddy Wilmet (Mechelen, Belgium) for comparative material, Jon Camilleri (Birkirkara, Malta) for the photography of specimens, Roland Houart (Landen, Belgium) for éditorial comments and recommendations and John J. Borg (NMNH), Philippe Bouchet & Virginie Héros (MNHN), Guido Poppe & Sheila Tagaro (conchology.be) for technical assistance. REFERENCES Bouchet, P. & Rocroi J.-P., 2005. Classification and nomenclator of gastropod families. Malacologia , 47(1-2): 1-397. Marrat, F. P., 1871. Oliva , Bruguière. Thésaurus Conchyliorum , 4: 1-46 + pis. 342-351. Olsson, A. A., 1956. Studies on the genus Olivella. Proceedings of the Academy of Nat lirai Sciences of Philadelphia , 108: 155-225. Petuch, E. J., 1987. New Caribbean molluscan faunas. pp. i-v + 1-154 + Al-A4 + 29 pl. Virginia (Coastal Education and Research Foundation). Ponder, W. F. & Warén, A., 1988. Classification of the Caenogastropoda and Heterostropha - A list of the family-group names and higher taxa. Malacological Review , Supplément 4: 288-326. Sterba, G. H. W., 2004. Olividae a collectors guide. pp. 1-172. Hackenheim (Conchbooks). Teso, V. & Pastorino, G., 2011. A révision of the genus Olivancillaria (Mollusca: Olividae) from the southwestern Atlantic. Zootaxa , 2889: 1-34. Vaught, K. C., 1989. A classification of the living Mollusca. pp. i-xii T 1-195. Florida (American Malacologists, Inc.). Figures 1-10 1-6. Agaronia johnabbasi sp. nov. East Pangandaran Bay, Java, Indonesia. 1-2. Holotype, MNHN 23267, 38mm; 3-4. Paratype 1, DPC R.GA1000, 41 mm; 5-6. Paratype 4, NMNH unreg., 31mm. 7-8. Agaronia lutaria (Rôding, 1798), East Pangandaran Bay, Java, Indonesia, 31mm (juvénile). 9-10. Agaronia johnkochi Voskuil, 1990. East Pangandaran Bay, Java, Indonesia, 47mm 34 D. P. ClLIA NOVAPEX 13(1): 33-36, 10 mars 2012 35 D. P. ClLIA A new Javan species of Agaronia Name and authority Distribution Oceanic division Agaronia acuminata (Lamarck, 1811) West Africa East Atlantic A. adamii Terzer, 1992 Philippines West Pacific A. annotata Marrat, 1871 West Africa East Atlantic A. cauta (Marrat, 1871) West Africa East Atlantic A. gibbosa (Born, 1778) India to Malaya East Indian Océan A. griseoalba (von Martens, 1897) Central America (west) East Pacific A. hiatula (Gmelin, 1791) West Africa East Atlantic A. hilli Petuch, 1987 Central America (east) West Atlantic A. jesuitarum Lôpez, Montoya-Maquin & Lôpez, 1988 Central America (west) East Pacific A. johnabbasi sp. nov. Indonesia East Indian Océan A. johnkochi Voskuil, 1990 Indonesia East Indian Océan A. leonardhilli Petuch, 1987 Central America (east) West Atlantic A. lutaria (Rôding, 1798) Indonesia East Indian Océan A. nebulosa (Lamarck, 1811) Indonesia East Indian Océan A. nica Lôpez, Montoya-Maquln & Lôpez, 1988 Central America (west) East Pacific A. propatula (Conrad, 1849) Central America (west) East Pacific A. razetoi Terzer, 1992 West Africa East Atlantic A. steeriae (Reeve, 1850) South America (east) West Atlantic A. testacea (Lamarck, 1811) Central America (west) East Pacific A. travassosi Morretes, 1938 South America (east) West Atlantic Table 1. Recent species of Agaronia Gray, 1839 based onMarrat (1871), Petuch (1987), Sterba (2004), Teso & Pastorino (2011) and the présent research. Bold type indicates species in close geographical proximity to the new species. Holotype MNHN 23267 Paratype 1 DC R.GA1000 Paratype 2 JA unreg. Paratype 3 JA unreg. Paratype 4 NMNH unreg. Paratype 5 GP unreg. Paratype 6 DPC R.GA1001 mean H 38 41 40 33 31 31 28 35 D 14 15 14 12 11 11 10 12 H/D 2.7 2.7 2.9 2.8 2.8 2.8 2.8 2.8 l'able 2. Dimensions of the type sériés of Agaronia johnabbasi sp. nov. (7 specimens) and their mean value Measurements are in millimétrés. 36 R. HOU ART NOVAPEX 13(1): 37-41, 10 mars 2012 Description of Muricopsis (Muricopsis) gorii (Gastropoda: Muricidae: Muricopsinae) from Southern Sâo Tomé Roland HOU ART Research Associate Institut royal des Sciences naturelles de Belgique Rue Vautier, 29, B-1000 Bruxelles, Belgium roland.houart@skynet.be KEYWORDS. Sâo Tomé, Muricidae, Muricopsinae, Muricopsis n. sp. ABSTRACT. A new species of Muricopsis is described from Sete Pedras Island, Sâo Tomé. ït is compared with Muricopsis matildeae Rolân & Fernandes, 1991. RESUME. Une nouvelle espèce de Muricopsis est décrite de l'île de Sete Pedras à Sâo Tomé. Elle est comparée avec Muricopsis matildeae Rolân & Fernandes, 1991. INTRODUCTION A few muricopsine species were described recently from Sâo Tomé and Principe and from Annobon Island by Rolân & Fernandes (1991), Houart & Rolân (2001), Houart (2005), Rolân & Gori (2007) and Houart & Gori (2008). Four Recent species and one subspecies of Muricopsis s.s. were described from Sâo Tomé: M. (M.) matildeae Rolân & Fernandes, 1991, M. (M.) rutilus mariangelae Rolân & Fernandes, 1991, M. (M.) delemarrei Houart, 2005, M. (M.) hemandezi Rolân & Gori, 2007, and Muricopsis (M.) testorii Houart & Gori, 2008. Two other species are known from two closely situated islands: Muricopsis (M.) pnncipensis Rolân & Fernandes, 1991- which occurs only in Principe and M (M.) annobonensis Houart & Rolân, 2001 which is endemic to Annobon Island. The new species described herein was recently discovered east of Sete Pedras Island (Fig. 1). Abbreviations IRSNB: Institut royal des Sciences naturelles de Belgique, Bruxelles, Belgium. MNHN: Muséum national d'Histoire naturelle, Paris, France. PR: Collection Peter Ryall. RH: Collection Roland Houart. SG: Collection Sandro Gori. Tenninology used to describe the spiral cords and the internai denticles of the outer lip (based on Merle 1999 and 2001) (Fig. 2). P: primary spiral cord; SP: subsutural cord; P: shoulder cord; P2-P6 : primary cords of the convex part of the teleoconch whorl; ADP: adapical primary cord on the siphonal canal; MP: médian primary cord on the siphonal canal; ID: infrasutural denticle of the aperture; DI to D5: abapical denticles of the aperture Fig. 1. Map of Sâo Tomé and location of the type locality of Muricopsis (M.) gorii n.sp. 37 R. Houart A new Muricopsis from Sâo Tomé Fig. 2. Muricopsis (Muricopsis) gorii n. sp. Spiral sculpture and apertural denticle morphology (paratype SG, 9.6 mm). SYSTEMATICS Family MURICIDAE Rafinesque, 1815 Subfamily MURICOPSINAE Radwin & D'Attilio, 1971 Genus Muricopsis Bucquoy & Dautzenberg, 1882 Subgenus Muricopsis Bucquoy & Dautzenberg, 1882 Type species by original désignation: Murex blainvillei Payraudeau, 1826 (= Murex cristatus Brocchi, 1814). Recent; Mediterranean. Muricopsis (Muricopsis) gorii n. sp. Figs 1,2, 3-8 Type material. Holotype MNHN 24275, 8.6 mm, 1 paratype PR, 9.6 mm, 1 paratype RH, 9.6 mm, 2 paratypes SG, 8.5 mm and 9.9 mm, Sâo Tomé, east of Sete Pedras, 30 m, collected on small rounded stones. Distribution. Currently only known from Sete Pedras, Sâo Tomé, in 30 m. Description. Shell small for the genus, up to 9.9 mm in height at maturity. Height/width ratio 1.9 - 2.1. Slender, biconical, weakly spinose, nodose, lightly built. Subsutural ramp broad, weakly concave, weakly sloping on spire whorls, more strongly sloping on last teleoconch whorl. Bright orange, weakly darker coloured on subsutural ramp, outer edge of columellar lip and apertural denticles lighter coloured or white. Figures 3-15 3-8. Muricopsis (Muricopsis) gorii Houart, n. sp., East of Sete Pedras, Sâo Tomé, on small rounded stones, 30 m 3-4. Holotype MNHN 24275, 8.6 mm; 5-6. Paratype SG, 9.9 mm; 7-8. Paratype RH, 9.5 mm. 9-14. Muricopsis (Muricopsis) matildeae Rolân & Fernandes, 1991. 9-10. Lago Azul, Sâo Tomé, 15 m, SG, 9 mm; 11-14. Ponto de Diego Vaz, Sâo Tomé, 12 m SG 11-12 9 6 mm; 13-14. 10.4 mm. 15^ Muricopsis (Muricopsis) testorii Houart & Gori, 2008. Northwest Sâo Tomé, Lagoa Azul, 00°24.4T N, 06°36.18' E, offshore, 37 m, on dead corals, holotype IRSNB IG.31 042/MT 1977, 18.8 mm. 38 ■ R.Houart Novapex 13(1): 37-41, 10 mars 2012 39 R. Houart A new Murïcopsis from Sâo Tomé Spire high with 1.5 protoconch whorls and teleoconch up to 4 or 5 broad, convex, strongly shouldered, nodose whorls. Suture adpressed. Protoconch small, whorls smooth with a narrow, strong, single keel adapically. Protoconch of holotype with a second weak shallow keel abapically, width 600-650 pm. terminal lip narrow, weakly raised, curved. Axial sculpture of teleoconch whorls consisting of moderately high, strong, broad, nodose ribs and varices. First and second whorl with 7 or 8 ribs, third and fourth with 6 or 7 ribs, last whorls with 4 or 5 broad, moderately high varices, each with 5 small, broad, short open spines at intersection of P1-P5 spiral cords with axial varices. Spiral sculpture of moderately high, narrow primary cords: first teleoconch whorl with visible SP, PI and P2, second with SP, PI, P2 and P3 partially covered by succeeding whorl, and several threads between primary cords. Last whorl with SP, P1-P6, ADP and occasional MP. Presence of several narrow threads between each pair of cords. P1-P4 of approximately similar size and strength, P5 and P6 narrower. Aperture small, ovate. Columellar lip broad, lip strongly flaring abapically, adhèrent at adapical extremity, smooth. Anal notch deep, narrow. Outer lip erect, smooth, with 5 strong denticles within: ID, Dl- D2 fused, D3, D4 and D5. Siphonal canal short, narrow, straight, weakly tapered abapically, narrowly open. Operculum and radula unknown. Remarks. Muricopsis (M.) gorii n.sp. is closely related to Muricopsis (M.) matildeae which lives in Sao Tomé and Principe and which, together with its typical form (Figs 9-10), shows a variety of shell colours from light orange to dark brown, occasionally with a white siphonal canal (Figs 11-14). However, ail these forms are sympatric and the shell morphology is quite similar. Not any of these could be definitely separated from the typical shell. Muricopsis (M.) gorii , in addition to having a bright dark orange colour, differs from M. matildeae in having a more strongly shouldered and angulate shell; higher and comparatively broader axial ribs on the 3 or 4 adapical teleoconch whorls and still higher and more conspicuous on the last teleoconch whorl but comparatively narrower than in M. matildeae ; in having a less sloping subsutural ramp from second to last teleoconch whorls, and in having blunt, broad, short spines at intersection of axial and spiral sculpture while absent or low in M. matildeae. Muricopsis (M.) testorii Houart & Gori, 2008 (Fig. 15), another species described from Sao Tomé is also orange coloured but differs in many ways (size and shell morphology) and doesn't need to be compared here. The other Muricopsis s.s. species described from Sâo Tomé, Principe and Annobon are very different and don't need to be compared either. Figures 16-19. Protoconchs (scale bar: 0.5 mm) 16-17. Muricopsis (Muricopsis) gorii n. sp. 16. Protoconch; 17. Protoconch and First teleoconch whorls. 18-19. Muricopsis (Muricopsis) matildeae Rolân & Fernandes, 1991. 18. Protoconch; 19. Protoconch and first teleoconch whorls. 40 R. HOUART NOVAPEX 13(1): 37-41, 10 mars 2012 Etymology. I am particularly pleased to naine this new species in honour of Sandro Gori (Livorno, Italy) who kindly donated the types and other material and who discovered many new taxa during his field trips in Sâo Tomé and Principe. Acknowledgements I am very grateful to Sandro Gori for the opportunity he gives me to study his material and for donating specimens. Thanks also to John Wolff, Lancaster, Pennsylvania, USA, for checking the English text. REFERENCES Houart, R. 2005. Description of a new species of Muricopsis (Gastropoda: Muricidae: Muricopsinae) from Sâo Tomé, West Africa. Novapex 6(4): 119-122. Houart, R. & Gori, S. 2008. Description of a new Muricopsis species (Muricidae: Muricopsinae) from Northwest Sâo Tomé. Novapex 9(4): 149- 153. Houart, R. & Rolân, E. 2001. A new Muricopsis (Gastropoda, Muricidae) from Annobôn Island, Eastem Atlantic. Novapex 2(2): 61-66. Merle D. 1999. La radiation des Muricidae (Gastropoda : Neogastropoda) au Paléogène: approche phylogénétique et évolutive. Paris. Thèse de doctorat du Muséum national d'Histoire naturelle: i-vi, 1-499. Merle D. 2001. The spiral cords and the internai denticles of the outer lip in the Muricidae: terminology and methodological comments. Novapex 2(3): 69-91. Rolân, E. & Fernandes, F. 1991. Muricopsis (Risomurex) (Gastropoda, Muricidae) de las islas de Sâo Tomé y Principe (Golfo de Guinea, Africa Occidental). Apex 6( 1 -2): 11 -20. Rolân, E. & Gori, S. 2007. A new species of Muricopsis (Muricidae: Muricopsinae) from Sâo Tomé Island. Novapex 8(1): 23-26. 41 E. Rolan & H. G. Lee Novapex 13(1): 43-44, 10 mars 2012 Rissoina parkeri (Mollusca: Rissooidae): a curious Caribbean species of uncertain status Emilio ROLÂN Museo de Historia Natural, Campus Universitario Sur 15782 Santiago de Compostela, Spain Harry G. LEE 4132 Ortega Forest Drive Jacksonville, FL 32210-5813, USA KEYWORDS. Gastropoda, Rissoina , Grand Cayman, Caribbean. ABSTRACT. Two shells of a Rissoina with planktotrophic development found in Grand Cayman, show the problematic of its diagnosis. Recently malacologist David Kirsh collected two specimens of a Rissoina in the waters of Grand Cayman Island. Each shell possess a planktotrophic protoconch (Figs 1-3) which character is not consistent with any known congener from the Caribbean (see Rolan & Fernândez-Garcés, 2010). The shells are coincident in size and sculpture with the figure and description of Rissoina parkeri , but hâve one more teleoconch whorl than the holotype of that species. This can explain why these two shells are a little larger (6.2 and 6.0 mm) than the holotype (4.4 mm). Olsson & Harbison (1953: 327-328, pl. 48, fig. 5) described the fossil species Rissoina parkeri from Shell Creek, Florida, USA. The type specimen is a little immature, with the lip not fully developed. It was deposited in ANSP (19434). In the searchable database of the Florida Muséum of Natural History (FLMNH) there is a shell with Catalog Number 26545 from Glades County, Neogene of Florida. On the Internet database Malacog 4.1.1 (Rosenberg, 2009) R. parkeri is tentatively treated as a synonym of Rissoina krebsii (Morch, 1876). However, on examination of these two species it is évident that the latter (Fig. 5) has more ribs and spiral cords and, most important perhaps, a paucispiral protoconch (Fig. 6). Another similar species is Rissoina multicostata (C.B. Adams, 1850) (Figs 7-8), which likewise has many more ribs and cords but a paucispiral protoconch (FigT 9). The only problem is that the species R. parkeri was considered a fossil while the two Kirsh shells hâve the appearance of Recent mollusks. Given a protoconch of more than 2 Vi whorl s, one must présumé that if it is a Recent species it would be known from other countries in the Caribbean and this is not presently the case. Another possibility is that this material, despite bearing the appearance of Recent shells, may be from a submarine Quarternary deposit. Acknowledgements The scanning électron micrographs were made by Jesüs Méndez in the Centro de Apoyo Cientifico y Tecnolôgico (CACTI) of the University of Vigo. To David Kirsch for the loan of the specimens studied. REFERENCES Olsson, A.A. & Harbison, A. 1953. Pliocène Mollusca of Southern Florida, with spécial reference to those from North Saint Petersburg. Monographs of the Academy of Natural Sciences of Philadelphia, 8: vii + 459, 65 pis. Rolan, E., & Fernândez-Garcés, R. 2010. New information on the Caribbean Rissoina (Gastropoda, Rissoidae) of the group R. sagraiana-cancellata, with the description of a new species. Iber us, 28(1): 79-89. Rosenberg, G. 2009. Malacolog 4.1.1: A Database of Western Atlantic Marine Mollusca. [WWW database (version 4.1.1)] URL http://www.malacolog.org/ . Specifically: < http://www. malacolog.org/search. php?syns : =Show+S vnonvms&showsvnonvms=true&mode=wheresearch &wherebit=+and+(+w.originalgenus+%3D+'Rissoina , ++or+w.currentfullname+like+ , Rissoina+%25 , +or+w. originalcombination+like+ , Rissoina+%25'+or+w.origi nalcombination+like+ , %25(Rissoina)%25')++ > 43 E. F. Garcia Rissoina parkeri a curious Carribean species Figures 1-9 1-4. Rissoina parkeri Olsson & Harbison, 1953 1-2. Shells, 6.2, 6.0 mm, Grand Cayman (coll. D. Kirsch); 3. Protoconch; 4. Microsculpture. 5-6. Rissoina krebsii (Morch, 1876) 5. Shell, 4.6 mm, Cienfuegos, Cuba (MHNS); 6. Protoconch. 7-9. Rissoina multicostata (C.B. Adams, 1850) 7-8. 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Vilvens, rue de Hermalle, 113, B-4680 Oupeye, Belgium or by e-mail: vilvens.claude@skynet.be NOVAPEX/SOCIETE (suite) C. Vilvens & Quoi de neuf ? Bourse d'Antwerpen 8 R.Scaillet, Quelques nouvelles publications C. Delongueville & 1. Plusieurs livres 10 C. Vilvens 2. Des documents électroniques 14 E. Meuleman Nous avons reçu 16 C. Vilvens && Nous avons reçu sur le Web 34 C. Delongueville & Les grandes marées de 2012 39 R. Scaillet IJ MALAC QL 401 .N68 v.13 no.2 NvVAPEX Trimestriel de la Société Belge de Malacologie association sans but lucratif Quarterly of the Belgian Malacological Society VOL. 13(2) 2012 10 JUIN SOMMAIRE Articles originaux - Original articles E. Rolân, R. Fernandez - Garcés & C. Redfern New records and description of four new species of the genus Agathotoma (Gastropoda, Mangeliidae) in the Caribbean 45 T. Ibarrola, F. Rubio & E. Rolân Some new information on Calliotropis ottoi (Philippi, 1844) (Vetigastropoda: Seguenzioidea: Calliotropidae) 63 A. Wakefield A review of the Marginella bicatenata Sowerby, 1914 complex (Gastropoda: Marginellidae) with the description of a new southeast African Marginella species 69 R. Houart & J. Colomb Description of a new species of Siratus (Gastropoda: Muricidae) from Guadeloupe, Lesser Antilles 75 R. Le Béon Une nouvelle Marginella (Gastropoda: Marginellidae) de la côte occidentale d’Afrique 79 J. Trôndlé & J. Letourneux Description de Turbo fakaauensis n. sp. (Mollusca: Gastropoda: Turbinidae) du Pléistocène de Niau, Tuamotu (Polynésie française) 87 NOVAPEX/SOCIETE C. Vilvens C. Delongueville & R. 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All rights of reproduction are reserved with the written permission of the board. Public a tions precedentes/Former publica tions: - Bulletin Mensuel d'information (1966-1971) - INFORMATIONS de la Société Belge de Malacologie (1972-1985) - ARION (1986-1999) -APEX (1986-1999) SOCIETE BELGE DE MALACOLOGIE E. Rolân, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 New records and description of four new species of the genus Agathotoma (Gastropoda, Mangeliidae) in the Caribbean Emilio ROLÂN Museo de Historia Natural, Campus Universitario Sur, 15782 Santiago de Compostela, Spain Raül FERNÂNDEZ-GARCÉS Centro de Estudios Ambientales (CEAC), Division de Gestion Ambiental (DGA), calle 17, esquina Ave 46, Cienfuegos, Cuba Colin REDFERN 7475 Estrella Circle, Boca Raton, Fia. 33433, USA KEYWORDS. Gastropoda, Mangeliidae, Agathotoma , Caribbean, new records, new species ABSTRACT. The species collected in the Caribbean and supposedly belonging to the genus Agathotoma are revised. Some new records are reported and four species new to science are described. RESUMEN. Se revisan las species recolectadas en el Caribe y supuestamente pertenecientes al género Agathotoma. Se refieren algunas citas nuevas y se describen cuatro especies nuevas parta la ciencia. INTRODUCTION The genus Agathotoma Cossmann, 1899 was discussed by Powell (1966), who provided a description defining ail the characters of the genus. He also listed the type species and gave examples of fossil and Recent species. In a recent paper (Fernândez-Garcés & Rolân, 2010) it is mentioned that in the révision of the Turridae by Powell (1966), 561 taxa names at genus or subgenus level are mentioned, from which 89 are in Mangeliinae. More recently new généra hâve been described, but unfortunately these descriptions are based on minimal morphological characters with no reference to soft parts, radula, DNA or the most important characters. In our opinion it is inadvisable to create new généra without first undertaking the broader studies that are necessary to give stability and future permanence to these new taxa. Expérience indicates that families with many généra based on minimal morphological characters will hâve to be restructured in the future, creating many synonyms and causing many changes. In West Africa there are also species which hâve been considered to belong to the genus Agathotoma as well as to the genus Pyrgocythara Woodring, 1928 (see Rolân & Otero-Schmitt, 1999). The latter genus has been employed for some species recently described (Fernândez-Garcés & Rolân, 2010). For the above-mentioned reasons we will présent ail the species studied in this paper in a well-established genus, avoiding the use of other more recently described généra, for which we do not hâve complété information. Abbreviations ANSP: Academy of Natural Sciences of Philadelphia. BMSM: Bailey-Matthews Shell Muséum, Sanibel, Florida. IES: Instituto de Ecologia, La Habana MCZ: Muséum of Comparative Zoology, Harvard. MNCN: Museo Nacional de Ciencias Naturales, Madrid. MHNS: Museo de Historia Natural, Santiago de Compostela. MNHN: Muséum national d'Histoire naturelle, Paris. NHMUK: National History Muséum United Kingdom, London. USNM: National Muséum of Natural History, Washington. CCR: collection of Colin Redfern, Boca Raton, Florida. CDK: collection of David Kirsh, Durham, North Carolina. CFG: collection of Fernândez-Garcés, Cienfuegos. sp: specimen with soft parts, s: shell. f: fragment, j: juvénile. 45 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species o f Agathotoma SYSTEMATICS Superfamily CONOIDEA Fleming, 1822 Family MANGELIIDAE P. Fischer, 1883 Genus Agathotoma Cossmann, 1899 Type species: Mangelia angusta Jan, Miocene, Europe. Agathotoma candidissima (C.B. Adams, 1845) Figs 1A-F, 10A, 11A-B, Table 1 Pleurotoma candidissima C.B. Adams, 1845. Proc. Boston Soc. Nat. Hist, 2: 4. Mangelia badia Reeve, 1846. Remarks in Krebs (1866): 396. Pyrgocythara coxi in Warmke & Abbott, 1961: plate 25, fig. T. Type material. Holotype in MCZ (186002), represented in Clench & Turner (1950, pl. 30, fig. 5) and in Williams (2006: 5510). Type locality. Jamaica. Other material examined. Cuba: 2 s, Guajimico, intertidal (MHNS); 2 s, Rancho Luna, Cienfuegos, 24 m (CFG); 1 s, Bajo de Sancho Pardo, 15 m (MHNS); 3 s, Rancho Luna, 20 m (MHNS); 3 s, Faro de los Colorados, Cienfuegos, 40 m (MHNS); 1 s, 1 j, Naranjo, Guajimico, Cienfuegos (MHNS); 6 s, 4 j, Itabo, Gavilân, Cienfuegos. 22°00 , 890”N, 80°24'832 , ’W, 10 m (MHNS). Nicaragua: 2 f, Cayo Witties, 12 m (MHNS). Bahamas: Abaco: 14 s, 1 j, Sandy Point, intertidal (CCR); 1 s, 1 sp, Treasure Cay, 1 m (CCR); 5 s, Chub Rocks, 10 m (CCR); 27 s, 5 j, off Chub Rocks, 23 m (CCR). Description. See C.B. Adams (1845) and Clench & Turner (1950). The protoconch has a diameter of about 400 pm, with about 2.25 whorls; the First part (about 1.75 whorls) is smooth but immediately small ribs begin in the suture and are complété on the last half whorl. The colour is white or cream, sometimes with two brown lines in the middle of the last whorl (Fig. 10A). Dimensions: Up to 5 mm in length. Distribution. Throughout the Caribbean. Some references can not be included due to doubtful détermination of the species. Remarks. This is the older species in this group described trom the Caribbean. The First problem is its generic placement. In a recent work by Fernândez- Garcés & Rolân (2010) this species was referred to the genus Pyrgocythara Woodring, 1928. However this is open to discussion, as no more information on soft parts is availabié. In the présent work we hâve decided to place it provisionally in Agathotoma, following most authors and databases. We hâve had many doubts about the assignment of A. candidissima made by several authors, as for example Leal (1991: pl. 24A), and also Garcia (2008: fig. 23) who reproduced the figure of the previous author. These Figures presented a shell with the upper extreme of the axial ribs very prominent which is not in accordance with the figure of the type represented in Clench & Turner (1950); on the contrary these shells are more coincident with the morphology of the species we will mention below as A. castellata. Pyrgocythara coxi Fargo, 1953 appears in Williams (2006) as a synonym of A. candidissima. We think that it is not, as the holotype of A. coxi shows axial ribs narrower than those on the typical A. candidissima. In De Jong & Coomans (1988: Figure 604A, p. 264) the shell presented as A. candidissima is not this species and the shell Figured in Fig. 604B is also dubious. The shell Figured here (Figures IA, 10A) has the typical profile. The species is correctly represented on some internet sites - (Conchologist Forum and Fémorale, for example). Agathotoma ecthymata Garcia, 2008 Figs 2A-G, 10B-D, 1 IC, Table 1 Agathotoma ecthymata Garcia, 2008. Novapex, 9(1): 3,figs. 10-18. Type material. Holotype in ANSP (416412) (represented in the original description). Type locality. Abaco, Bahama Islands. Other material examined. Cuba: 4 s, Canon de la Bahia, Cienfuegos, 12 m (CFG); 3 s, Los Laberintos, Cienfuegos, 20 m (CFG); 6 s, Rancho Luna, Cienfuegos, 24 m (CFG); 9 s, 2 j, Rancho Luna, Cienfuegos, 15 m (MHNS); 4 s, 3 j, Rancho Luna, Cienfuegos, 18 m (MHNS); 1 j, Cayo de Sancho Pardo, 12 m (MHNS); 1 s, Punta del Diablo, Gavilân, Cienfuegos, 21°57796”N, 80°24784”W, 20 m (MHNS); 11 s, 3 j, Bajo de Sancho Pardo, 15 m, (MHNS); 2 s, La Habana, 5 m (MHNS); 2 s, Itabo, Gavilân, Cienfuegos. 22 o 00'890”N, 80 o 24'832”W, 10 m (MHNS). Nicaragua: 4 s, 5 j, Cayo Witties, 12 m (MHNS). Mexico: 4 s, 2 j, Puerto Morelos, Yucatân, 12 m (MHNS). Bahamas: Abaco: 12 s, Treasure Cove, intertidal (CCR); 5s, 1 sp, Treasure Cay, 1 m (CCR); 4 s, Shell Island, Guana Cay, intertidal (CCR); 1 sp, Fish Cays, 3 m (CCR); 1 j sp, Guana Cay, 3.5 m (CCR); 2 sp, 1 j sp, Scotland Cay, 2.5 m (CCR); 1 sp, Don’t Rock, 3.5 m (CCR). Description. See Garcia (2008). In the original description the protoconch is mentioned as having 1.5 whorls, but following the Verduin method (Verduin, 1976) which we usually employ, it has only a little 46 E. Rolân, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 Figure 1. A-F. Agathotoma candidissima (C.B. Adams, 1845). A. shell, 4.7 mm, Faro de los Colorados, Cienfuegos, Cuba (MHNS); B. shell, 3.4 mm, Maria la Gorda, Cuba (MHNS); C-D. protoconch; E-F. microsculpture and detail. 47 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma more than one whorl and does not reach 1.25 whorls. Also Garcia (2008) did not mention the dimensions of the protoconch, which we calculate to be between 320 to 340 pm in the figures. The protoconchs of our material are approximately 350-370 pin in diameter. Distribution. In Garcia (2008) it is recorded from: Bahamas, Belize, British Virgin Islands, Colombia. This is the first record for Cuban, Nicaraguan and Mexican waters. Remarks. Most of our material is consistently smaller than the shell figured by Garcia (2008), but the microsculpture of the protoconch and the teleoconch seems to be similar. We show the microsculpture with better detail (Figs. 2F-G). A . candidissima is more elongate, less robust, the shoulder is rounded and not so angular, and the protoconch has almost 2.25 whorls. Agathotoma apocrypha (Garcia, 2008) Figs 3A-F, 11D, Table 1 Suturocythara apocrypha Garcia, 2008. Novapex , 9(1): 3, figs. 33-36. Type material. Holotype in ANSP (416415) (represented in the original description). Type locality. Bahia de Campeche, SW Gulf of Mexico. Other material examined. Cuba: 10 s, Faro Luna, Cienfuegos, 22 m (CFG); 1 s, Faro de los Colorados, 30 m (MHNS); 6 s, Rancho Luna, Cienfuegos (MHNS); 1 s, Bajo de Sancho Pardo, 12 m (MHNS); 1 s, Punta del Diablo, Gavilân, Cienfuegos, 21°57’796 ,, N, 80°24’784”W, 20 m (MHNS). Mexico: 3 s, Puerto Morelos, Yucatân, 12 m (MHNS). Description. See Garcia (2008). The protoconch is mentioned in the original description as having 1.25 whorls, which agréés with our shells. In that work the dimensions of the protoconch were not mentioned, which we calculate to be about 470 pm in the figures. The protoconchs of our material are approximately 400-430 pm in diameter. Otherwise, the microsculpture seems to be similar, although the zigzag Unes are not as well-defined as in our shells. The microsculpture of the teleoconch is shown in our plates. Distribution. In the original description, Garcia (2008) records this species from the southwestem Gulf ol Mexico, the southeast coast of Florida and south of Belize. This is the first record for Cuban waters. Remarks. In spite of the différences in the size of the protoconch and the microsculpture, we consider our material to be the same species. The shells are very similar to the type species of the genus, Agathotoma angusta (Jan) Bellardi, 1848 represented in Powell (1966: pl. 15, fig. 15), being even more similar to the type species than the shells of Agathotoma candidissima. We figure with better detail the microsculpture of both the protoconch and the teleoconch in Figures 3E-F. The species presented above are very different in profile and microsculpture, also with protoconch différences. The most similar protoconch is that of Agathotoma ecthymata but the microsculpture has minute tubercles aligned spirally. Agathotoma castellata (E.A. Smith, 1888) Figs 4A-G, 10E-F, 11E, Table 1 Pleurotoma (Mangilia) castellata E.A. Smith, 1888: 312-313. Pyrgocythara candidissima in RlOS (1994): 175, fig. 806. Type material. The lectotype of A. castellata is in BMNH (1854.6.30.136-7) and was designated by Kilburn (1993: fig. 71); also it was represented by KAICHER (1984), as Agathotoma costellata (sic), and in Williams (2006). Type locality. Unknown. Material studied. Cuba: 4 s, Faro de los Colorados, Cienfuegos, 20-35 m (MHNS); 5 s, 2 j, Rancho Luna, 20-35 m (MHNS); 5 s, Faro Luna, Cienfuegos, 22 m (CFG); 2 s, Los Laberintos, Faro Luna, Cienfuegos. 22°02'039”N, 80°25'792 ,V W, 10 m (MHNS); 1 s, Comodoro Beach, 8-10 m (MHNS). Bahamas: Abaco: 17 s, Sandy Point, intertidal (CCR); 1 j, off Guana Cay, 60 m (CCR); 7 s, 2 j, off Chub Rocks, 23 m (CCR); 1 s, Sandy Cay, 7 m (CCR). Description. Shell broadly ovoid, spire with turreted profile, solid, pointed, cream in colour with 4-5 irregular spiral brown bands sometimes faded partially or totally. Protoconch with 2.25 whorls, a diameter of about 500 pm, a nucléus of about 100 pm, 1.5 smooth whorls and the last 3 A bearing about 20 curved axial ribs. Teleoconch of about 3-4 whorls, ail having a very prominent subsutural shoulder with the upper part of the whorls almost horizontal. Each whorl has about 7-8 almost orthocline or slightly opisthocline axial ribs, more prominent on the shoulder and much naiTower than their interspaces. Spiral sculpture of very numerous and variable cordlets (about 14-16 on the first whorl), 20-22 on the second, 22-23 on the third and more than 60 on the body whorl, sometimes with intervening cordlets developing. Under high magnification the cordlets can be seen to be rugose, bearing numerous rounded rough nodules which 48 E. Rolân, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 Figure 2. A-G. Agathotoma ecthymata Garcia, 2008. A-C. shells, 3.0, 3.1, 3.4 mm, Los Laberintos, Cienfuegos (MHNS). D-E. protoconch. F-G. microsculpture and detail. 49 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma Figure 3. A-F. Agathotoma apocrypha (Garcia, 2008). A-C. shells protoconchs; F. microsculpture. 3.8, 3.7, 3.5 mm, Cienfuegos (MHNS); D-E. 50 E. Rolân, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 Figure 4. A-G. Agathotoma castellata (E.A. Smith, 1888). A. shell, 5.2 mm, Abaco, Bahamas (CCR); B-C. 3.9, 4.0 mm, Faro de los Colorados (MHNS); D-E. juvénile, 2.6, 3.2 mm, Rancho Luna (MHNS); F. protoconch; G. microsculpture. 51 EmilioRolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma overlap the interspaces; in the interspaces there are numerous irregular minute microtubercles. Aperture elongate-oval, extended by a wide and short siphonal canal; on the upper part a relalively deep sinus, with a tubercle on its lower edge. Microsculpture of the shell extends onto a wide thickened peristome. Dimensions: The lectotype is 6.5 mm in length. The material studied from Cuba and Bahamas is smaller. Distribution. This species lias been recorded from the Caribbean (Rosenberg, 2009) and from Cuba in the présent work. It was also recorded (as A. candi dis sim a) from the Bahamas (Redfern, 2001: fig. 537A) and Brazil (Leal, 1991: pl. 24, fig. A), with the latter figure reproduced by Rios (1994: 175, fig. 806) and Garcia (2008: fig. 23). Remarks. It is évident that this species is rather variable: 1- The axial ribs of the lectotype of A. castellata are very narrow (interspaces 4-5 times wider) while in most of our shells the interspaces are only 2-3 times wider. 2- Usually the ribs extend above the level of the suture, a feature which is more évident in juvénile shells. 3- The lectotype oï A. castellata measures 6.5 mm in length for a shell with 4 teleoconch whorls, while we hâve shells with the same number of whorls and a length of only 4-5 mm. 4- The siphonal canal of A. castellata is relatively shorter in larger shells. With regard to the confusion of the présent species with A. candidissima , we must point out that the latter is narrower, more elongate, the rounded upper part of the axial ribs is less pronounced , the axial ribs never extend above the suturai level and the colour is whitish with narrow brown bands. Both species are présent in Cuba and we hâve never tound any intergradations between them, and so we consider them to be different species. Agathotoma kirshi spec. nov. Figs 5A-H, 11F, Table 1 Type material. Holotype (Figs. 5A-D) deposited in BMSM (17948) (Ex-CCR 10498). Paratypes: CDK (1 s) (Fig. 5E), Pélican Point, North Caicos, intertidal; MNHN (24829, 1 s) (Fig. 5F), from Sandy Cay, Abaco, Bahamas, 7 m. Type locality. Chub Rocks, Abaco, Bahamas, under a rock, 12 m. Other material examined. Nicaragua: 1 s, 1 f Cavo Witties, 20 m (MHNS). Description. Shell with ovoid elongate profile, slightly turreted, rather solid, pointed, white in colour with 2 spiral bands formed by isolated brown blotches, mainly évident on the ribs. Protoconch with only a little more than 1 whorl, with a diameter of about 480 pm, being totally covered by very small rounded tubercles spiral ly aligned. Teleoconch of about 5-5.5 whorls, with a rounded shoulder close to the upper suture. Each whorl has about 7 almost orthocline or slightly opisthocline ribs, more prominent on the shoulder and narrower than their interspaces. On the last A whorl, the axial ribs may be absent. Spiral sculpture of very numerous and variable cordlets (between 15-20 on the first whorl), with additional intervening ones developing, numbering about 24 on the second, 27 on the third and more than 70 on the body whorl. Under high magnification the cordlets can be seen to be rugose, bearing numerous rounded rough nodules that are axial ly elongate and overlap the interspaces; in the interspaces there are numerous irregular minute microtubercles. Aperture oval elongate, extended by a wide and short siphonal canal; on the upper part a shallow sinus. Peristome wide, with the same microsculpture as the shell. Dimensions: The holotype is 8 mm in length. The soft parts examined in a specimen from Abaco (Bahamas) (Figs. 5 B-D) are whitish and slightly translucent, with a few milk-white spots at the end of the siphon and with the eyes very close to the end of the tentacles. Distribution. Known from the Bahama Islands and Turks and Caicos Islands. A shell from Nicaragua is probably this species, but is not in good enough condition tor a definitive comparison. Remarks. Agathotoma kirshi spec. nov. is the largest of the species studied in the présent work. Axial ribs may be obsolète on the last whorl. The persistence of a previous apertural thickening seems to be frequent. Agathotoma candidissima is smaller, shorter, without former varices, the protoconch has more whorls, the colour is whitish. Agathotoma ecthymata is more robust, the shoulder is more prominent, the axial ribs wider, the last whorl straighter towards the base, the protoconch is narrower and less elevated. Agathotoma apocrypha is smaller, the axial ribs more numerous and wider, the microsculpture has cords altemating with cordlets, the protoconch has zigzag fines. Agathotoma castellata is smaller, relatively wider, with the axial ribs prominent and surpassing the level of the suture; the protoconch has 2.5 whorls. Etymology. Named after David Kirsh, who collected the shell from North Caicos and loaned it to us for study. 52 E. ROLÂN, R. FERNAnDEZ-GARCÉS, C. REDFERN NOVAPEX 13(2): 45-62, 10 juin 2012 Figure 5. A-H. Agathotoma kirshi spec. nov. A-D. bolotype, 8 mm (BMSM); A. shell; B-D. holotype with soft parts; E. paratype, 9 mm, North Caicos (CDK); F. paratype, 7 mm, Sandy Cay, Abaco, Bahamas (MNHN); G. protoconch, from figure F; H. microsculpture. 53 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma Agathotoma asthenika spec. nov. Figs 6A-E, 10G, 10K-L, 1 IG, Table 1 Type material. Holotype (Fig. 6A) in the MNCN (15.05/60008). Paratypes in the following: MNHN (24830, 1 s); frora the type locality; MHNS (100567, 1 s), IES (1 s); CFG (1 s) and CCR (1 s), both from Faro de los Colorados, Cienfuegos, 40-65 m; USNM (1 s), La Habana, 12 m. Other material examined. Cuba: I s, Rancho Luna, 22 m, broken during the study; 3 s, Los Laberintos, Faro Luna, Cienfuegos 22 m (CFG); 5 s, Los Laberintos, Faro Luna, Cienfuegos, 22°02 , 039”N, 80°25 , 792”W, 10 m (CFG); 4 s, Ij, Bajo de Sancho Pardo, 15 m (MHNS). Type locality. Los Laberintos, Faro Luna, Cienfuegos, 30 m, Cuba. Description. Shell ovoid-elongate, rather solid, pointed, white with some tan areas between the ribs and also in the suturai area. Protoconch of a little more than 2 whorls, the first one and a quarter whorls smooth and the last half axially ribbed. Diameter of about 410 jim. Teleoconch of about 4 whorls, with a slightly prominent shoulder a short distance below the suture. Each whorl has 8-9 prominent orthocline or slightly opisthocline ribs, a little prominent on the shoulder and narrower than their interspaces. Spiral sculpture of very numerous and irregular cordlets (about 10 on the first whorl, about 11-12 on the second and 40 to 55 on the body whorl). High magnification reveals very fine Unes in the interspaces which form axially elongate nodules when Crossing the cordlets. In the interspaces there are also numerous irregular micro-tubercles. Aperture oval elongate extended by a wide and short siphonal canal and with a deep sinus on its upper part. Peristome wide, with the same microsculpture as the shell. Dimensions: The holotype is 3.9 mm in length . Other material studied is between 3-4 mm in length. Distribution. Only known from Cuba. Remarks. A. candidissima is larger, a little wider, more solid, the shoulder is less pronounced and rounded, the colour is white sometimes with faded spiral bands and lacks subsutural colour. A. candidissima is a slightly variable species, but it is not possible that these shells could represent a population of narrow shells because typical shells of both species hâve been collected in the same area without intergradations. Agathotoma ecthymata is wider and more robust, and has a paucispiral protoconch. Agathotoma apoctypha is wider, mainly on the base, the spiral sculpture has larger cordlets and the piotoconch is short and strongly sculptured by zigzag cords. J o o Agathotoma castellata is wider, with axial ribs which are more elevated in the subsutural area. Etymology. The spécifie name is derived from the Greek word “asthenikôs ” which means “thin, sickly” alluding to its narrow profile. Agathotoma eduardoi spec. nov. Figs 7A-G, 10H, 10M-N, 11H, Table 1 Type material. Holotype (Fig. 7A) in the MNCN (15.05/60009). Paratypes in the following: MNHN (24831, 1 s, Fig. 7B, 11D); MHNS (100568, 5 s, Fig. 7C, 11C), IES (1 s); CFG (1 s) ail paratypes from Rancho Luna, Cuba. Other material examined. Cuba: 2 s, Itabo, Gavilân, Cienfuegos, 22°00'890”N, 80°24'832”W, 10 m (MHNS); 1 s, Rancho Luna, Cienfuegos 20 m (CFG). Type locality. Cayo Witties, 12 m, Nicaragua. Description. Shell ovoid-elongate, rather solid, pointed, whitish in colour. Protoconch with a little more than one whorl, appearing smooth, but magnification reveals numerous spirally aligned tubercles; diameter of about 380-400 pm. Teleoconch of about 4 whorls in most mature specimens, with a slightly prominent shoulder a short distance below the suture. Each whorl has prominent orthocline or slightly opisthocline ribs, nine on the first whorl and about ten on the subséquent whorls, the ribs being narrower than their interspaces. Spiral sculpture of very numerous and irregular cordlets (about 10 on the first whorl, about 14 on the second and between 45-50 on the body whorl). High magnification reveals very fine Unes in the interspaces which form axially elongate nodules when Crossing the cordlets, with an irregular microsculpture. In the interspaces there are also numerous irregular micro-tubercles. Aperture oval elongate extended by a wide and short siphonal canal and with a deep sinus on its upper part. Peristome wide, with the same microsculpture as the shell. Dimensions: The holotype is 4.2 mm in length. Other material studied is between 2-4 mm in length. Distribution. Known from Cuba and Nicaragua. Remarks. The différences with the species previously mentioned are very similar to those discussed for A. asthenika spec. nov., as both shells are very similar except for the different shorter protoconch. Agathotoma candidissima is larger, a little wider, more solid, the shoulder is less pronounced and rounded, the colour is white sometimes with faded spiral bands, lacks subsutural color, and has a protoconch with more whorls. 54 E. ROLÂN, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 Figure 6. A-E. Agathotoma asthenika spec. nov. A. holotype, 3.9 mm, Los Laberintos, Faro Luna (MNCN); B. juvénile, 3.0 mm, Maria la Gorda (MHNS); C-D. protoconch; E. microsculpture. 55 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma ramr™ 7 ; Spec - nov - A holol yi>'- 4 - 2 mm, Cayo Witties, Nicaragua (MNCN); Nie— Vr ^ a’- r J mm (MHNS) ’ from Ranch0 L ““. 1 MO m; D. protoconch from Witties, icaragua, E-F protoconchs from Rancho Luna, Cuba. G. microsculpture. 56 E. Rolân, R. Fernândez-Garcés, C. Redfern NOVAPEX 13(2): 45-62, lOjuin 2012 Figure 8. A-D. Agathotomaprominens spec. nov. A. holotype, 4.1 mm, (MNCN); B. paratype, 4.3 mm (MNHN), Faro de los Colorados; C. protoconch; D. microsculpture. 57 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma Agathotoma ecthymata is wider and more robust, the shoulder more angular, and its protoconch is rather similar but a little narrower and its tubercles are more dense. Agathotoma apocrypha is wider, mainly on the base, the spiral sculpture has larger cordlets and the protoconch is sculptured by zigzag cords. Agathotoma castellata is wider, with axial ribs which are more eievated in the subsutural area and the protoconch has more whorls. Agathotoma kirshi spec. nov. has a larger shell, wider and more sculptured, and the axial ribs do not hâve a clear shoulder. Agathotoma asthenika spec. nov. has a rather similar shell, but it has a very different protoconch with a little more than 2 whorls, the first one smooth. Etymology. The spécifie name honours Eduardo Nâpoles Fernandez, grandson of the second author. Agathotoma prominens spec. nov. Figs 8A-D, 10I-J, 111, Table 1 Type materlal. Holotype (Fig. 8A, 101), in MNCN (15.05/60010); Paratypes: MNHN (24832, 1 s, Fig. 8B, 10J), MHNS (100569, 3 s), IES (2 s), USNM (1 s), CFG (2 s), ail from the type locality; CFG (3 s) Canon de la bahia de Cienfuegos, 12 m; (2 s), Faro Luna, 20 m; CCR ( 1 s), Bajo de Sancho Pardo, 15 m; MHNS (100569, 2 s), from Cayo Maria la Gorda, 12 m; (100569, 1 s), Cayo Witties, 12 m, Nicaragua. Type locality. Faro de los Colorados, Cienfuegos, 20 m, Cuba. Other material examined. Cuba: 2 s, 1 j, Itabo, Gavilân, Cienfuegos. 22°00 , 890”N, 80°24'832”W, 10 m (MHNS); 1 s, 5 f, Rancho Luna, Cienfuegos, 10-20 m (MHNS). Nicaragua: 1 f, Cayo Witties, 12 m. Bahamas: Abaco: 1 s, Sandy Cay, 7m. Description. Shell with rhomboidal profile, solid, pointed, cream with three irregular spiral brown bands. Protoconch with 2.25 whorls, a diameter of about 510 gm, a nucléus of about 100 pm, the last Vi whorl with 11 curved axial ribs, the rest being smooth. Teleoconch of about 3.5 whorls, which hâve a prominent and angular shoulder just below the suture. Each whorl has prominent, almost orthocline ribs, prominent on the shoulder and narrower than their interspaces. Spiral sculpture of numerous and variable cordlets (about 15 on the first and more than 60 on the body whorl). High magni fi cation shows the cordlets to be rugose, bearing numerous axially elongate nodules which mainly overlap the interspace below. Interspaces sometimes with fine growth Unes and numerous irregular minute micro-tubercles. Aperture oval elongate, extended by a short siphonal canal that is wide at the base but constricted above; also with a deep sinus on the upper part. Peristome wide, with the same microsculpture as the shell. Dimensions: The holotype is 4.1 mm in length. One paratype reaches 5.2 mm in length. Distribution. Only known from Cuba, Nicaragua and the Bahamas. Remarks. Comparison may be made with the following species: Agathotoma candidissima is usually larger, a little wider, more rounded in profile, the shoulder is less pronounced and not angular, the colour is white sometimes with faded spiral bands. Agathotoma ecthymata is wider at the base, more robust, the ribs are a little prominent but less angular, and has a paucispiral protoconch. Agathotoma apocrypha is narrower, wider at the base, the shoulder less prominent and the spiral sculpture has larger cordlets; the protoconch is short and strongly sculptured by zigzag cords. Agathotoma castellata is usually smaller, narrower, the shoulder is less prominent and the ribs are more eievated; the colour does not form spiral bands. We include images (Figs. 9A-B, 10O) of Glyphoturris rugirima (Dali, 1889) with a detail of the microsculpture (Fig. 9C). It can appear similar to the species here described, but differs due to its very irregular and prominent spiral cords. Etymology. The spécifie name is derived from the Latin word “ prominens ”, prominent, alluding to the projecting angulation on the axial ribs. Agathotoma sp. Fig 10P-Q Remarks. Also we found a large shell that is apparently différent from ail those previously studied, but it could be a gerontic form of A. candidissima. We wait for more material in order to reach a conclusion. CONCLUSION The Caribbean is an area very rich in biodiversity. It is expected that more species from many groups will be described in the future. In the group that we hâve examined in the présent work the profile of the shells, in particular the shape of the upper part of the ribs, was considered to be very important and was consistent in many samples. Foi comparison, we show in Figure 11 how the ditteient profile of the shells dépends on the shape of the upper part of the axial ribs below the suture. As a séparation for the species mentioned in the présent work we add a Dichotomous Key on the basis of the most important différences. 58 E. Rolân, R. Fernândez-Garcés, C. Redfern Novapex 13(2): 45-62, 10 juin 2012 Figure 9. A-C. Glyphoturris rugirima (Dali, 1889). A-B. shell, 5.0, 4.8 mm, Rancho Luna, 10-30 m (MHNS); C. microsculpture. Dichotomous key for the species of Agathotoma in the présent work| 1 -Protoconch with only 1-1.3 whorls. -Protoconch with at least 2 whorls. 5 2 -Microsculpture of the protoconch with zigzag Unes. apocrypha -Microsculpture of the protoconch with tubercles. 3 3 -Protoconch with cylindrical profile . kirshi -Protoconch with rounded dôme profile. 4 4 -Protoconch with dense microsculpture of tubercles. ecthymata -Protoconch with microsculpture of tubercles in lines. eduardoi 5 -Axial ribs with pronounced angulation a short distance below suture . 6 -Upper part of axial ribs rounded near suture. 7 6 - The angulation is < 90° on first whorl and around 90° on the following ones . prominens - the angulation is about 90° on first teleoconch whorl and > 90° on the following. asthenika 7 - Axial ribs elevated subsuturally but not surpassing level of suture; the profile of the ribs is vertical . candidissima - Upper curvature of axial ribs frequently surpassing level of suture; the profile of the ribs is inclined . castellata 59 Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma Table 1 protoconch whorls protoconch microsculpture protoconch diameter inpm upper part of ribs height/width ratio A. candidissima 2-2.3 smooth+axial about 400 rounded 2.6 A. ecthymata 1-1.3 dense tubercles 350-370 lower angulation 2.2 A. apocrypha 1-1.3 zigzag lines 400-430 rounded 2.6 A. castellata 2-2.3 smooth+axial about 500 rounded high 2.3 A. kirshi 1-1.3 aligned tubercles about 480 slightly rounded 2.4-2.7 A. asthenika 2-2.3 smooth+axial about 410 lower angulation 2.8 A. eduardoi 1-1.3 aligned tubercles 380-400 rounded 2.6 A. prominens 2-2.3 smooth+axial about 510 sharp angulation 2.1 Acknowledgements The authors thank the persons who hâve read this manuscript and hâve given us their opinions, allowing us to make corrections. Jésus Méndez and Inès Pazos made the SEM photos in the Centro de Apoyo Cientifico y Tecnolôgico a la Investigaciôn (CACTI) ofthe University of Vigo. The optical photographs were made in the Departamento de Xenética of the same University. REFERENCES Clench, W.J. & Turner, R.D. 1950. The Western Atlantic Marine Mollusks described by C.B. Adams Occasional Papers On Mollusks , 1(15): 233-403. Conchologist forum: http://zl4.invisionfree.eom/Conehologist_Forum/a r/t586.htm (accessed Dec. 2011) De Jong, K. M. & Coomans, Fl. E. 1988. Marine gastropods from Curaçao, Aruba and Bonaire. E. J. Brill, Leiden, 261 pp. Fémorale: http://www.femorale.com.br/shellphotos/detail.asp ?species=Agathotoma+cf.+candidissima+(C.B.Ad ams%2C+1845) (accessed Dec. 2011) Fernândez-Garcés, R. & Rolân, E. 2010. Two new species ofthe genus Pyrgocythara (Gastropoda, Conidae) from Cuba. Gloria Maris, 49(3-4): 68- Figure 10. A-Q. Colour figures of Agathotoma. A. Agathotoma candidissima (C. B. Adams 1845) 4 7 mm Faro de los Colorados, Cuba (MHNS). B-D. Agathotoma ecthymata Garcia, 2008; B 3 7 mm Rancho I una (MHNS); C-D. 5.1 mm, Faro de los Colorados (MHNS). E-F. Agathotoma castellata (E A Smith 1 8S84 "1 4 , 0 mncn;,v mh h S) rsnrr r F,gs - r* G Asa,ho,oma “te ™«. .2 19 mm (MNCN) (figured in Fig. 6A). H. Agathotoma eduardoi spec. nov., holotype, 4 2 mm (MNCN) (t.gured in Fig. 7A). I-J. Agathotoma prominens spec. nov.; I. holotype, 4.1 mm (MNCN)- I naratvne 4 1 rŒr "rf n S; « n B) , K 'V **«r?*°& s p ec - no.! 3 4™’: (MHNS) (Shell figured in Fig. 6B). M-N. Agathotoma eduardoi spec. nov.; M. paratype 3 3 mm (MHNS)' » m t have been figured in Figs - 7b - c) - °- mg™ dIh x 1889), 5 mm. Rancho Luna, 10-30 m (MHNS); P-Q. Aga,kcoma s p. 8.4 mm. Rancho Lufa.TSo m (MH 60 E. Rolân, R. Fernândez-Garcés, C. Redfern NOVAPEX 13(2): 45-62, 10 juin 2012 61 1 mm Emilio Rolân, R. Fernândez-Garcés, C. Redfern New records and new species of Agathotoma Garcia, E.F. 2008. Eight new molluscan species (Gastropoda: Turridae) from the western Atlantic, with the description of two new généra. Novapex, 9(1): 1-15. Kaicher, S. D. 1984. Gard Catalogue of World-wide Shells. Pack 39. Turridae 1. Kilbum, R. N. 1993. Turridae (Mollusca: Gastropoda) of southem Africa and Mozambique. Part 6. Subfamily Mangeliinae, section 2. Armais of the Natal Muséum , 34(2): 317-367. Leal, J.H., 1991. Marine Prosobranch Gastropods from Oceanic lslands off Brazil. W. Backhuys, Oegstgeest. 419 pp. Powell, A.W.B. 1966. The molluscan families Speightiidae and Turridae. An évaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Muséum , 5: 1-184; 23 pis. Redfern, C. 2001. Bahamian Seashells. A thousand species from Abaco, Bahamas . Bahamianseashells.com, Boca Raton, 280 pp. Rios, E. 1994. Seashells of Brazil. 2nd Edition. Museo Oceanogrâfico, Fundaçâo Universidade do Rio Grande: Rio Grande. 368 pp. Rolân, E. & Otero-Schmitt, J. 1999. The family Turridae s. 1. (Molluscs, Neogastropoda) in Angola. 2. Subfamily Mangeliinae Fischer, 1883. Argonauta , 13(1): 5-26. Rosenberg, G. 2009. Malacolog 4.1.1 : A Database of Western Atlantic Marine Mollusca. [WWW database (version 4.1.1)] URL http://www.malacolog.org/. Smith, E.A. 1888. Diagnoses of new species of Pleurotomidae in the British Muséum. Annals and Magazine ofNatural History , 6(2): 300-317. Verduin, A. 1976. On characters, variability, and distribution of the European marine gastropods Bittium latreillii (Payraudeau) and Bittium lacteum (Philippi). Basteria. 40: 133-142. Warmke, G. L. & Abbott, R. T. 1961. Caribbean Seashells. Livingston Publishing Company: Wynnewood, PA. 348 pp. Williams, M. 2006. Shallow water Turridae of Florida and the Caribbean. Published privately. Not paginated. Figure 11. A-I. Comparative profile of the different species. A-B. Agathotoma candidissima (C. B. Adams, 1845); C. Agathotoma ecthymata (Garcia, 2008); D. Agathotoma apocrypha (Garcia, 2008); E. Agathotoma castellata (E.A. Smith, 1888); F. Agathotoma kirshi spec. nov.; G. Agathotoma asthenika spec. nov.; H. Agathotoma eduardoi spec. nov.; I. Agathotomaprominens spec. nov. 62 T. IBBARROLA, F. RUBIO, E. ROLÂN Novapex 13(2): 63-67, 10 juin 2012 Some new information on Calliotropis ottoi (Philippin 1844) (Vetigastropoda: Seguenzioidea: Calliotropidae) Teodoro IBARROLA Instituto Espanol de Oceanografia, Vigo teo.ibaiTola@hotmail.com Federico RUBIO Pintor Ribera, 4-16 a 46930 Quart de Poblet, (Valencia), Spain, federubio@ono.com Emilio ROLÂN Museo de Historia Natural, Campus Universitario Sur 15782, Santiago de Compostela, erolan@emiliorolan.com KEYWORDS. Vetigastropoda, Calliotropidae, Calliotropis ottoi , synonymy, new data. ABSTRACT. Some specimens of the deep water species Calliotropis ottoi (Philippi, 1844) collected in Hatton Bank were studied. The shell, operculum, radula and soft parts are illustrated. The synonymy of this species with other taxa from the Atlantic is discussed. RESUMEN. Se estudian algunos ejemplares de la especie de aguas profundas Calliotropis ottoi (Philippi, 1844) recolectadas en el Banco Hatton. Se muestra concha, opérculo, radula y partes blandas. Se discute la sinonimia de esta especie con otros taxones del Atlântico. INTRODUCTION Calliotropis ottoi (Philippi, 1844) is a species from the Atlantic deep water, but there is a record of Martens & Thiele (1904) referring this species (and presenting the radula) from the Indian Océan. Hickman & McLean (1990) gave information on the soft parts, operculum and radula on the basis of the study of the American populations. Vilvens & Swinnen (2008: figs. 29-33) made a révision of the genus Calliotropis from central eastem Atlantic, showing the original figures of Calliotropis ottoi (Philippi, 1844) from the original description and from Martens & Thiele (1904) as well as a shell from lceland. Colman & Tyler (1988) gave information on the reprodution of this species. Some specimens of Calliotropis ottoi (Philippi, 1844) were collected by rock dredge by the Spanish Océanographie Institute Ecovul/Arpa 2007 Survey on board R/V Vizconde de Eza on Hatton Bank area (NE Atlantic) at 59°17N, 17°33W in 962 meters. Some additional information on the morphology of this species is given in the présent work. SYSTEMATICS Remarks The genus Calliotropis Seguenza, 1803 has been placed in several subfamilies as Margaritinae, Angariinae, Monodontinae, Calliostomatinae and Solariellinae. According to Hickman & McLean (1990) it should be placed in Eucyclinae Koken, 1897, in the tribe Calliotropini Hickman & McLean, 1990. Following Bouchet & Rocroi (2005) and Vilvens (2007) this genus should be placed in the family Chilodontidae Wenz, 1938, in the subfamily Calliotropinae Hickman & McLean, 1990. Williams et al (2008), based on DNA sequences placed Calliotropidae Hickman & McLean 1990 in Seguenzioidea Verrill, 1884. Genus Calliotropis Seguenza, 1903 Type species by original désignation: Trochus ottoi Philippi, 1844, Pliocène, Pleistocene, Italy. 63 T. IBBARROLA, F. RUBIO, E. ROLÂN New information on Calliotropis ottoi Calliotropis ottoi (Philippi, 1844) Figs 1-13 Trochus ottoi Philippi, 1844: 227, pl. 28, fig. 9. Description. Shell (Figs 1-4): Vilvens & Swinnen (2008) commented in the remarks the most important differential characters with other species. Soft parts (Figs 6-10): Foot elongate, sole very rough. The cephalic lappets are relativelly small and simple. The cephalic tentacles are large and elongate with an irregular ciliate surface. The cephalic membranes are of medium size and with a continuons border. Buccal margin with latéral extensions, mouth small. The neck lobes are of medium size and very digitate. The tip of the snout are amply expanded latérally. The epipodial tentacles are numerous: Two in medium position and three other ones of different size located near of the operculum insertion. Between the tentacles and at the base of some of them sensorial organes tuberculiformis, remind small tentacles. Propodium with latéral extensions. Eyes black, pedunculate. Operculum (Fig. 5): yellowish, transparent, with short growing edge. Radula (Figs 11-13): The rachidian tooth has a plate from which a sharp-pointed curved part appears on the upper area. The rachidian tooth (Fig. 12R) is slightly smaller; then the latéral teeth (Fig. 12L) are three, also curved on their upper part, where some small cusps can be seen near the sharp-pointed part; they are not so large as those shown by Hickman & McLean (1990: fig 47). The marginal teeth (Fig. 12M) are numerous and similar, narrow, elongate and curved at their upper part. The existence of lateromarginal plate is confirmed but difficult to be seen. Remarks. Figures of this species has been published by Philippi (1844) and by Vilvens & Swinnen (2008) where the figures of Martens & Thiele (1904) are also shown. Discussion. A problem not yet resolved is the synonymy of this species: Warén (1991) mentioned with doubts that Margarita regalis Verrill & Smith, 1880 could be the same species. He also presented Solariella infundibulum Odhner, 1912 as a synonym. About the identification of the species we hâve studied, il we follow the key given by Vilvens & Swinnen (2008) we could hâve doubts between Calliotropis ottoi and C. mogadorensis (see Locard, 1898). Some characters, as the existence of a spiral coid into the umbilicus which has been mentioned for this last species is not sure and seems to be a variable character, as well as the first row of nodules in the subsutural area, variable in oui* own material. The most definitive decision was based in the origin of the material, very far from Morocco and close to the references from North Europa. It is possible that Calliotropis mogadorensis (Locard, 1898) could be another synonym of C. ottoi. Hickman & McLean (1990: 84, figs D-E) figured the soft parts of Calliotropis species, showing the existence of a short number of epipodial tentacles at each side: Four in Calliotropis carlotta (Dali, 1902) and three in C. regalis (Verrill & Smith, 1880). Both are very different from the species here studied in the number and position of the epipodial tentacles. In spite of that, the position of the soft parts was examined in retracted animais, we can see in our material up to eight epipodial tentacles at each side. This seems to mean that this character is very variable into the genus. Another question is that C. regalis was also considered a synonym of the species here studied. ACKNOWLEDGEMENTS The authors thanks Jésus Méndez of the CACT1 (University of Vigo) for the SEM photographs of the radula. The optical photographs were made in the Department of Xenetic of the University of Vigo. REFERENCES Bouchet P. & Rocroi J.-P. (eds), 2005. Classification and nomenclator of gastropod families. Malacologia 41 (1-2): 1-397. Colman, J.G. & Tyler, P.A. 1988. Observations on the reproductive biology of the deep-sea trochid Calliotropis ottoi (Philippi). Journal of Molluscan Studies 54: 239-242. Hickman, C.S. & McLean, J.H. 1990. Systematic révision and suprageneric classification of Trocacean Gastropods. Natural History Muséum of los Angeles County. Science sériés 35, 169 pp. Locard, A. 1898. Expéditions Scientifiques du Travailleur et dit Talisman pendant les années 1880, 1881, 1882, 1883. Mollusques Testaces. \o\. II. Masson et Cia, Paris. 515 pp, 18 pis. Martens, E. von & Thiele, J. 1904 "1903”. Dis beschalten Gastropoden der Deutschen Tiefsee- Expedition, 1898-1899. A Systématise h geographischer Teil Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expédition auf dem Dampfer "Valdivia" 1898-1899, 7(A): 1-146. Philippi, R.A. 1844. Enumeratio molluscorum Siciliae cum viventium tn tellure tertiaria fossilium, quae in itinere suo observavit. Vol. 2. Eduard Anton, Halle [Halis Saxorum] iv + 303 p., pis 13-28. SStS/iw Ef li ° tropis ot,oi ( PhiliPPi, 1844). 1-4. shell, 14.4 height x 16.5 mm diameter, Hatton Bank, 59 17N/17 33 W; 5. operculum; 6-7. soft parts. 64 T. IBBARROLA, F. RUBIO, E. ROLÂN NOVAPEX 13(2): 63-67, lOjuin 2012 65 T. IBBARROLA, F. RUBIO, E. ROLÂN New information on Calliotropis ottoi Vilvens, C. 2007. New species and new records of Calliotropis (Gastropoda: Chilodontidae: Calliotroponae) from Indo-Pacific. Novapex , Hors Série 5: 1-72. Vilvens, C. & Swinnen, F. 2008. New records of Calliotropis (Gastropoda: Chilodontidae) from the central eastem Atlantic. Novapex 8(1): 17-32. Warén, A. 1991. New and little known Mollusca from Iceland and Scandinavia. Sarsia 76: 53-124. Williams, S.T., Karube, S. & Ozawa, T. (2008) Molecular systematics ol Vetigastropoda: Trochidae, Turbinidae and Trochoidea redefined. Zoologica Scripta 37: 483-506. Figures 8-10 Calliotropis ottoi (Philippi, 1844). Several positions of the soft parts (preserved in alcohol). CT: cephalic tentacles, F. foot, M. mouth; NL: neck lobes; O: operculum; ET: Epipodial tentacles; SO: sensorial organes. 66 T. IBBARROLA, F. R.UBIO, E. ROLÂN Novapex 13(2): 63-67, 10 juin 2012 Figures 11-13 Calliotropis ottoi (Philippi, 1844). Radula. 11. general view; 12. details under magnification, showing details of the different teeth: R: rachidian tooth; L: latéral teeth; M: marginal teeth; 13. general view under magnification. 67 A. Wakefield Novapex 13(2): 69-74, 10 juin 2012 A review of the Marginella bicatenata Sowerby, 1914 complex (Gastropoda: Marginellidae) with the description of a new southeast African Marginella species Andrew WAKEFIELD 14 Forest Side, Buckhurst Hill, Essex, IG9 5SL, UK. bmw.awake@btintemet.com KEYWORDS. Marginellidae, South Africa, Marginella bicatenata , M. tomlini , M. lemaitrei, M. seccombei , species complexes, new species. ABSTRACT. The taxa Marginella bicatenata , Sowerby, 1914, M tomlini Shackleford, 1916 and M. lemaitrei Liltved & Millard, 1994, ail rare deep-water South African marginellids from off the southem Cape to KwaZulu-Natal, are revised from their type material and additional lots from both public and private collections. M. tomlini is proposed to be a junior synonym of M. bicatenata , whereas M. lemaitrei is considered to be a sibling species. Marginella seccombei n. sp., a benthic species from KwaZulu-Natal often confused with M. bicatenata on account of its superficially similar shell pattern, is described. INTRODUCTION In 1914, George Brettingham Sowerby (III) described a new species of Marginella from a then unique shell, lacking data, which had been discovered in the collection of a Mr. M. Denans. Since Denans had collected extensively in Sénégal, Sowerby erroneously assumed the type locality to be Gorée. The otherwise plain whitish shell was named Marginella bicatenata on account of its double row of dark spots, one at mid-body and the other at the shoulder. Two years later Lewis J. Shackleford, without reference to Sowerby’s taxon, described Marginella tomlini from a shell with a similar pattern, only this time its provenance was known with accuracy to be off Cape St. B laize, on the Southern Cape of South Africa. This shell was dredged from 105 fathoms (équivalent to 192 m) indicating a deep water habitat on the Agulhas Bank. Since that time, and especially in recent years with increased sampling of South African benthic environments down to several hundred métrés and more, further specimens of shells resembling the type specimens of both taxa hâve corne to light and hâve found their way into both private and public collections. There has also been relatively recent taxonomie activity in this small complex of similar shells, with the description of Marginella lemaitrei Liltved & Millard, 1994. Despite the continuing scarcity of specimens available for study sufficient material now exists to make a preliminary review of the complex. M. bicatenata ranges from Africa’s southem Cape, northeast to central KwaZulu Natal, with the shells of this species exhibiting morphologie différences at each end of the géographie range of distribution. These différences are exemplified by the type specimens of M. bicatenata and M. tomlini. A new deep water species from Natal, bearing a superficially similar pattern of a double row of markings at the shoulder and mid-body, does not however appear to be directly related to M. bicatenata. It is described herein as M. seccombei n. sp. Materials and Methods Ail specimens from private collections and ail of the material of the new species described herein were obtained via suppliers who sourced material from South African commercial fishing vessels as dead dredged empty shells. Curators of national muséums in the United Kingdom and South Africa permitted the use of images and data of their examples of the species under study. Since live animais are as yet unknown, the species were treated conchologically. The author used a Nikon D300 SLR, a 60mm AF Micro Nikkor 1:2.8D lens and ring flash for images of M. bicatenata and M. seccombei n. sp. taken by the author. Images are shown at the same relative scale. Abbreviations NMW: Amgueddfu Cymru (National Muséum, Wales), Cardiff. NM: KwaZulu (Natal Muséum), Pietermaritzburg. SAM: Iziko (South African Muséum), Cape Town. AWC: Andrew Wakefield Collection, UK. TMC: Tony McCleery Collection, UK ad.: adult juv.: juvénile sh.: dead collected shell n. sp.: new species SYSTEMATICS Family MARGINELLIDAE Fleming, 1828 Subfamily MARGINELLINAE Fleming, 1828 Genus Marginella Lamarck, 1799 69 A. Wakefield A review of Marginella bicatenata Type species: Voluta glabella Linnaeus, 1758, by monotypy. Marginella bicatenata G.B. Sowerby (III), 1914 Figs 1-21 Marginella bicatenata Sowerby (III), 1914: 147, pl. 19, fig. 7. Marginella tomlini Shackleford, 1916: 193, text figs 3, 4. Type material. Marginella bicatenata ,1 ad sh., holotype, preserved dry, 13 x 7 mm, Gorée, col 1. Tomlin (ex. Coll Denans), NMW.1955.158.01434, (Figs 1,2). Marginella tomlini , 1 ad. sh., holotype, preserved dry, 18 x 10 mm, off Cape St. Blaize in 105 fms, SAM A3704 (Figs 13-15). Other material examined. 6 ad and 1 juv. sh, from KwaZulu-Natal, preserved dry: 12.8 x 6.9 mm, 29.825° S 31.2383°E, NM ref. no. D3819 (Figs 3,4); 13.9 x 7.5 mm, 30.0132° S, 31.06°E, NM ref. no. DI 159 (Figs 5,6); 15.7 x 8.5 mm, 30.0067° S 31.05°E, NM ref no. DI 094 (Figs 7,8); 13.5x6.6 mm, juv., NM ref. no. E8656 (Figs 9, 10); 11.7 x 6.8 mm, 30.0182° S 31.0533°E, NM ref. no. D800 (Figs 11, 12); 14.6 x 8.8 & 12.8 x 7.4 mm, 30.1067° S 31.0133°E, NM ref. no. D1946. 2 ad sh., from the Southern Cape région, preserved dry: 17.8 x 9.95 mm, Agulhas Bank, dredged, depth unrecorded, AWC; 16.2 x 9.33 mm, Southern Agulhas Bankat 100m, AWC (Figs 16, 17). Type Locality. Gorée, Sénégal (in error) Descriptive notes. Shell thin, smooth, length 11.5-18 mm, W:L ratio 54-60% (mean 56%), strongly biconic to elongated biconic. Spire elevated, of 4.5 whorls including paucispiral protoconch, stepped (mainly between penultimatc and last adult whorl) to straight sided. Shoulder smoothly rounded to slightly angulated, shell tapering to narrow, slightly truncated anterior end. Creamy white, pale straw to pale grey body whorl, with two spiral rows of blurred charcoal grey spots: anterior row of 6-8 spots emerges from aperture immediately posterior to 4 th plication, ending at anterior l/6 th of lip; posterior row of 7-10 (on body whorl) spots at shoulder level. Posterior row continues onto spire to reach plain, glassy protoconch. Suture not impressed. Lip, plications, base of columella white. Aperture approximately 1.5 x as wide as labial varix. Lip smooth, straight to gently convex, thickened externally as a single moderately strong varix with a smooth rolled edge. Varix groove présent externally. Siphonal notch absent, posterior notch weak, lip thins slightly in posterior l/6 th before inserting to body whorl at shoulder at level of posterior row of spots, or just anterior to it. Lip continues round anterior end, thinning out completely to join base of columella at end of First plication. Columella with 4 moderately strong, thin, single plications, gradually increasing in séparation from l sl to 4 lh . First two oblique, 3 and 4 becoming more horizontal. Plications fill ovei /i but less than 2/3 of aperture. Columella slightly concave in région of plications. Pariétal surface smooth, weakly convex to straight. Anterior and posterior callus absent. Animal unknown. Distribution. Off central KwaZulu-Natal to Cape St. Blaize, southem Cape, South Africa, depth range 100- 200 métrés. Remarks. The species is listed by Tomlin (1917: 253) who records the holotype of M. bicatenata as being présent in the Tomlin collection. The Melvill-Tomlin Collection was received by Amgueddfu Cymru (NMW) in 1955, and enquiries hâve confirmed that the holotype is présent in the collection. It is biconic in profile (Figs 1, 2) and from the ventral view has three large spots at the shoulder and four at mid-body. It clearly is the shell depicted and described by Sowerby. Similarly robust, relatively small, dirty-white shells with charcoal coloured spots and with a variety of strong morphologie features such as stepped spires, and strongly angulated shoulders are lound at the north-eastern end of the range, off KwaZulu-Natal (see the specimens from the NM in Figs 3-12). As the holotype exhibits these characters it is much more likely to hâve originated from Natal than from the Southern Cape, and the original type locality has always been erroneous - this is emphatically not a West African shell. Millard (1981, fig. Da, p. 6, 7) figured a specimen présent in the SAM (ref. A3704), citing it as the type of M. bicatenata . As noted above, the holotype of M. bicatenata is in the NMW. The specimen to which Millard was referring is in fact the holotype of M. tomlini Shackleford, 1916 (Figs 13-15). The original description of it featured a quality photograph of the holotype - one of the earliest original descriptions to do so - and its identification as the holotype of M. tomlini is beyond any doubt. Shackleford recorded the type locality as off Cape St. Blaize, southem Cape, South Africa, C N. by E. Vi E., distant 68 miles - 105 fathoms {sic) 9 . M. tomlini is considered to be a junior synonym of M. bicatenata , and represents the form found at the Southern end of the range. These shells are larger and thinner walled than their northem counterparts, and they also inhabit relatively shallower waters in the south. This trend is observed in most other species of South African marginellidae with a similar distribution pattern. This progressive morphological change has been attributed to many factors, one being the availability of food (the cooler waters ot the Cape are rich in nutrients compared to the warmer waters of KwaZulu-Natal). One ot the NM specimens (Figs 9, 10) of M. bicatenata is in exceptional condition and is very well marked, and even shows extra fine spiral lines of spots 70 A. Wakefield Novapex 13(2): 69-74, 10 juin 2012 between the two main ones. Most specimens of M bicatenata are dead collected and worn - it may be that fresh specimens hâve traces of extra fine spiral Unes in the surface layers of the shell. These markings are not as defined or as bold as those found in M. lemaitrei. M. bicatenata is very distinctive and is unlikely to be confused with anything else. M. nevillana Kilbum, 1977 (7.6 x 4.5 mm) as yet only known from its holotype from the eastem Cape, is much smaller than M. bicatenata , has eight rows of large charcoal grey spots on a white background, a much thicker shell and because of its denticulate lip is better placed in the genus Glabella Swainson, 1840 (in the G. obtusa Sowerby, 1846 complex). Marginella lemaitrei Liltved & Millard, 1994 Figs 22-26 Marginella lemaitrei Liltved & Millard, 1994: 3, 4, fig 1 . Type material. 1 ad. sh., holotype, 18.4 x 10.0 mm, ex. pisce , trawled approx 100m, Cape St. Blaize, Southern Cape, South Africa, SAM A3 7572 (Figs 22- 24) ; 1 ad sh., paratype, 17.1 x 8.9 mm, ex. pisce , trawled approx 100m, Cape St. Blaize, Southern Cape, South Africa, NM E7213/T 184. Other material examined. 1 ad. sh., unmeasured, no data, photographed by Markus Lussi (Figs 25, 26). Specimen figured on p. 6, 7, fig. Db, in Millard, 1981 (21x11mm), ‘ ex . pisce ’ off the Cape. Specimen image by Brian Hayes (2011). Type locality. Cape St Blaize, Southern Cape, South Africa. Distribution. Restricted to the Agulhas Bank off the Southern Cape, South Africa. Descriptive notes. Shell length 17.1 — 21.0 mm, W:L ratio 52-54% (mean 53%), straw coloured to greyish, thin, smooth, satin, elongate-biconic, of 4.5 whorls including paucispiral protoconch, weakly shouldered, tapering to a narrow anterior end. Suture not impressed. Lip with smooth, evenly thickened, white margin, smooth internally, with extemal varix groove. Plications strong, evenly spaced plications filling over half of aperture. First two oblique, third and fourth straighter. Columella slightly concave. Aperture narrowing posteriorly, otherwise of even width. 4-8 spiral rows of fine grey spots and streaks on body whorl. Holotype has 8 rows; 4 anteriorly, 2 at mid- body, 2 on the spire. Paratype has 4 rows with less frequent, more blurred markings. Axial pattern of pale grey narrow to wide flammules on body whorl. Remarks. According to Liltved and Millard (1994: 4), M. lemaitrei occurs sympatrically with M. bicatenata (‘M tomlini form’) off the Southern Cape. In fact the types of M lemaitrei and the holotype of M. tomlini are syntopic. The types of M. lemaitrei were taken from the stomach contents of the fish Congiopodus tonms Walbaum. M. lemaitrei differs from the ‘M. tomlini form’ of M bicatenata in that it is generally slightly larger, more slender (less shouldered), has multiple rows of dashes and dots rather than two, and often has an additional axial pattern of pale grey narrow to wide flammules. The slender thin shell and axial flammules are also characters seen in Marginella diadochus Adams & Reeve, 1848 and therefore the possibility of an ancestral Iink of the M bicatenata group to the M. mus ica group should be considered. Marginella seccombei n. sp. Figs 28-39 Type material. 5 ad. sh., dredged, preserved dry, off Central KwaZulu-Natal in 150m; holotype, 12.61 x 7.46 mm, NM W8641/T2985 (Figs 27-30); paratype 1, 13.29 x 8.37 mm, NM, W8642/T2986, (Figs 31, 32); paratype 2, 12.48 x 8.28 mm, AWC (Figs 33, 34); paratype 3, 10.88 x 6.35 mm, TMC (Figs 35, 36); paratype 4, 13.75 x 8.48 mm, NMW.Z.2012.016.00001 Type locality. Off central KwaZulu-Natal, South Africa. Other material. 1 ad. sh. unmeasured, off Durban, KwaZulu-Natal, photographed by Markus Lussi (Figs 37, 38) Distribution. Off Central KwaZulu-Natal in 100-150 métrés. Description. Shell moderately sized (L=12.6 mm), W:L ratio 58 - 66 % (mean 61.4%), solid, biconic, of 4 Vi whorls including paucispiral protoconch, posteriorly tapering to moderately wide columella base. Pale straw coloured with two spiral bands of very short axial ly oriented red-brown dashes encircling body whorl. Anterior row appears darker due to dashes being doser together. Dashes regularly spaced and shaped ventrally, becoming more separated and irregular from mid-dorsum to lip. Anterior (mid-body) row of 24 closely spaced dashes emerges immediately posterior to 4 th plication. Posterior row of 20 more spaced marks at shoulder on body whorl, continuing for another 2 % whorls onto spire. Lip, base of columella and plications white. Shoulder rounded, spire moderately elevated, suture présent, spire whorls shouldered. Posterior end of lip inserts at shoulder at level of posterior spiral band. Columella slightly concave anteriorly, pariétal surface very slightly convex. Aperture widest in middle third, twice as wide as the lip, narrowing posteriorly and anteriorly to as wide as the lip. Lip evenly convex in 71 A. Wakefield A review of Marginella bicatenata profile, smooth, denticles absent, external varix strong with varix groove externally. Posterior notch absent, siphonal notch weak. Four strong pilications, first two oblique, third and fourth straighter, round crested, spacing between increasing from tirst to fourth, ends terminate at level of lip except for first which sweeps round to join with base of columella and anterior end of lip. Animal unknown. Etymology. The species is named after Alan Seccombe (Cape Town), who first drew the authors attention to this species. Remarks. The type sériés (Figs 27-36) displays the variability in this species. The number ol short axial dashes in the two bands varies slightly, from 19 to 23 in the anterior band and from 19 to 24 in the posterior band. Whilst many shells are marked only with the two main spiral rows, others hâve extra rows ol tiner markings. Paratype 2 (Figs 33, 34) is exceptionally well marked and reveals these extra spiral lines of extremely fine dots and dashes on the body whorl, with 22 fine spiral dotted lines between the two main bands, but also several anteriorly, and posteriorly onto the spire. Paratype 1 (Figs 31,32) has these extra line spiral lines of dashes and dots more organised so they line up in an axial pattern. DISCUSSION Although the colour pattern of M seccombei apparently links it to the M. bicatenata complex, there are several characters which when studied in detail, provide us with an indication that M seccombei is not closely related to it at ail. Firstly, the shells of the M bicatenata complex are ail relatively thin and hâve lower W:L ratios; 54-60% (mean 56%) for M bicatenata and 52-54% (mean 53 % ) for M lemaitrei. The W:L ratio of M. seccombei is much higher at 58- 66 % (mean 61.4%) and is clearly a more robust, stockier shell than those of the M bicatenata group. Secondly, the double row of reddish-brown axial dashes in M. seccombei , is fundamentally different in its genesis from the double row of smudged charcoal grey spots of the other species. This strongly suggests that M. seccombei has a different phyletic lineage, and that its pattern should be regarded merely as a convergent shell character with the M bicatenata group. "We must look elsewhere for links to its related species. When the species in the M. ornata complex express spiral lined patterns, they tend to do so as bands of small axially orientated markings. This is very évident on reddish specimens of M. ornata Redfield, 1870 and also in M. beltmani Fïart, 1993 and M. peelae Bozzetti, 1993. M seccombei also shares the same general morphology of the shells in the M. ornata complex, with their stocky outline and convex, often stepped later spire whorls. On purely conchological grounds therefore, a relationship with the M. ornata complex would seem justified although the lack of external lip markings in M seccombei , compared with strongly marked lips of those in the M ornata complex, cannot be ignored. Looking then to more northerly species for allies of M seccombei , the pattern and morphology of two deep- water Mozambique species M. verdascai Hayes & Rosado, 2007 and M. monicae Bozzetti, 1997 appear close. M verdascai , though much smaller at a length of 5.6 - 6.8 mm (in the type sériés), has a very fine spiral lined pattern very much like that seen in the better preserved specimens of M seccombei. Figures 1-38 1-21. Marginella bicatenata , G.B. Sowerby (111), 1914; 1-2. M. bicatenata , Holotype, 13x7 mm, Coll. Tomlin (NMW 1955.158.01434); 3-4. Off Durban, Natal,12.8 x 6.9 mm, 29.825°S 31.2383°E (NM D3819); 5, 6. Off Durban, Natal, 13.9 x 7.5 mm, 30.0132°S 31.06°E (NM DI 159); 7, 8. Off Durban, Natal, 15.7 x 8.5 mm, 30.0067°S 31.05°E (NM DI 094); 9, 10. Off Durban, Natal, 13.5 x 6.6 mm (NM E8656); 11,12. Off Durban, Natal, 11.7 x 6.8 mm, 30.0182°S 31.0533°E (NM, D800); 13- 15. M tomlini , Holotype, trawled off Cape St. Blaize, Southern Cape, South Africa in 192m, 18x10 mm (SAM A3704); 16, 17. Southern Agulhas Bank, Southern Cape, South Africa, 16.2 x 9.33 mm (AWC); 18-22. Sériés demonstrating pattern and morphologie variability, data unknown (photo Lussi). 22-26. Marginella lemaitrei Liltved & Millard, 1994; 22-24. Holotype, Cape St. Blaize, Southern Cape, South Africa, 18.4 x 10.0 mm (SAM, A37572); 25, 26. Data unknown (photo Lussi) 27-38. Marginella seccombei n.sp., off Durban, KwaZulu-Natal, dredged dead in 150m 27-30. Holotype, 12.61 x 7.46 mm (NM, W8641/T2985); 31, 32. Paratype 1. 13.29 x 8.37 mm (NM, W8642/T2968); 33, 34. Paratype 2. 12.48 x 8.28 mm (AWC); 35, 36. Paratype 3. 10.88 x 6.35 mm (TMC); 37, 38. Data unknown (photo Lussi). 72 A. Wakefield Novapex 13(2): 69-74, 10 juin 2012 73 •• A. Wakefield A review of Marginella bicatenata However, it lacks the two main spiral bands of brown axial markings and it has a weakly denticulate lip. Although smaller than M. seccombei, M monicae is a shell of comparable shape though it is thicker and slightly pyriform. Although mainly lacking a colour pattern, it has a faint trace of dark markings (not clear spots or axial streaks) at the shoulder and at the anterior end in the same place as the spiral rows in M seccombei. It appears to be the species closest morphologically to M. seccombei discovered to date. M tuguriana Lussi,1993, another benthic species occurring in northern Natal and possibly ranging further north into Mozambique, also has a faint anterior band, but its extremely biconic morphology, thicker shell and labial markings distinguish it clearly from M. seccombei n. sp. Acknowledgements Dr. Igor Muratov (KwaZulu-Natal Muséum) kindly provided the images of M bicatenata (Figs 3-12). Linda Davis (Collections Manager, KwaZulu-Natal Muséum), Liz Hoenson (Iziko, South African Muséum), and Harriet Wood (Amgueddfu Cymru, National Muséum of Wales) generously provided access to type material and/or information. 1 would like to express spécial thanks to Alwyn Marais (Centre for Molluscan Studies, South Africa) for the images of the holotype of M tomlini (Figs 13-15), and to Markus Lussi (Durban) for his opinions on the distribution of M. bicatenata, for his excellent images (Figs 16-21, 25, 26, 37 and 38) and for being my referee. A final thanks to Alan Seccombe (Cape Town) for providing the author with type material ot M seccombei. REFERENCES Cossignani, T. 2006. Marginellidae and Cystiscidae of the World. L’informatore Piceno, Ancona. ISBN 88-86070-10-1. Hayes, B., 2011. www.gastropoda.com (date of access 28/02/2012) Liltved, W.R., & Millard, V.G. 1994. Five New Species from Southern Africa. World Shells, 10: 3- 10 . Millard, V. G., 1981. Marginellidae of South Africa. The Strandloper , 206: 1-12, 2 pl. Shackleford, L.J., 1916. Two New Species of Marginella from South Africa. Annals of the South African Muséum , 9 :193-194. Sowerby, G. B. IIÏ, 1914, Descriptions of New Mollusca from New Caledonia, Japan, Philippines, China and West Africa. The Annals and Magazine of Natural History , ser. 8, 14(84) : 475-480, pl. 19. Tomlin, J. R. Le B., 1917. A Systematic List of the Marginellidae. Proceedings of the malacological Society of London. XII, 242-307. 74 R. Houart & J. Colomb Novapex 13(2): 75-78, 10 juin 2012 Description of a new species of Siratus (Gastropoda: Muricidae) from Guadeloupe, Lesser Antilles Roland HOUART Research Associate Institut royal des Sciences naturelles de Belgique Rue Vautier, 29, B-1000 Bruxelles, Belgium roland.houart@skynet.be Jacques COLOMB 82, rue A. Daudet F- 13013 Marseille, France j acquescolomb@wanadoo. fr KEYWORDS. Gastropoda, Muricidae, Lesser Antilles, Martinique, Siratus , new species. ABSTRACT. Siratus michelae is described from six specimens dredged in about 70 m depth off Martinique, French Antilles. It is compared with S. cailleti (Petit, 1856) and S. kugleri (Clench & Pérez Farfante, 1945), both also occurring in the same area, but at greater depth. INTRODUCTION There are currently 27 Recent species of Siratus of which 24 live throughout the Western Atlantic and three in the Indo-West Pacific (Houart, 2012). Ail the species are listed by Houart (2010) and most of them are illustrated by Merle, Garrigues & Pointier (2011). The Western Atlantic species were also commented and illustrated by Vokes (1965 and 1990a), while nine species from the Western Atlantic were named since 1990 by Vokes (1990b), Houart (1999 and 2000), Merle, Garrigues & Pointier (2001) and Merle & Garrigues (2008 and 2011). The most recent species was described from the Dominican Republic and Martinique. Abbreviations Repository MNHN: Muséum national d'Histoire naturelle, Paris, France. RH coll. : Collection of Roland Houart. JC coll.: Collection of Jacques Colomb. Terminology used to describe the spiral cords and apertural denticles morphology (after Merle, 1999 and 2001) (Figs 1-2). P: primary cord; s: secondary cord; t: tertiary cord; ad: adapical (or adapertural); ab: abapical (or abapertural); IP: infrasutural primary cord (primary cord on subsutural ramp); adis: adapical infrasutural secondary cord (on subsutural ramp); abis: abapical infrasutural secondary cord (on subsutural ramp); PI: shoulder cord; P2-P6: primary cords of the convex part of the teleoconch whorl; sl-s6: secondary cords of the convex part of the teleoconch whorl (example: si = secondary cord between PI and P2; s2 = secondary cord between P2 and P3, etc.); ADP: adapertural primary cord on the siphonal canal; MP: médian primary cord on the siphonal canal. Aperture: ID: Infrasutural denticle ; DI to D6: Abapical denticles SYSTEMATICS Family MURICIDAE Rafinesque, 1815 Subfamily MURICINAE Rafinesque, 1815 Genus Siratus Jousseaume, 1880 Type species, by original désignation: Purpura Sirat "Adanson" Jousseaume, 1880 (= Murex senegalensis Gmelin, 1791), Recent, Brazil. Siratus michelae n. sp. Figs 1-2, 3-4, 7-12 Type material. Holotype MNHN 25131 Paratypes: 4 JC coll., 1 RH coll. Type locality. Martinique, Atlantic coast, 70 m. Distribution. Currently only known from the type material: Martinique, Atlantic coast, alive at 70 m. Description. Shell small for the genus, up to 41.4 mm in height at maturity. Height/width ratio 2.1-2.2. Broadly ovate, nodose, lightly built. Subsutural ramp broad, strongly sloping, weakly concave. Light brown or greyish-brown with lighter coloured or creamy white axial nodes. Spiral cords usually topped with fine dark brown lines. P3 and P5 almost entirely white, more obvious on axial nodes. Area between P3 and P5 lighter coloured. MP spine creamy white. 75 R. Houart & J. Colomb A new species of S ira tus from Guadeloupe Protoconch light or dark brown. Aperture bluish- white, pariétal tooth white, edge of outer lip with dark brown spots between apertural crenulations. Spire moderately high with 1.75 protoconch whorls and 6-6.5 broadly convex, weakly shouldered, nodose whorls. Suture weakly adpressed. Protoconch small, whorls rounded with a weak, narrow, single keel abapically (Fig. 4). Maximum width 700-900 pm, height 700 pm. First whorl small, 300-350 pm width. Terminal lip shallow, délicate, opistocyrt. Axial sculpture of teleoconch whorls consisting of high, narrow, rounded ribs and high, strong, narrow, rounded, weakly spinose varices. First whorl with 11- 13 ribs, second whorl with 5-9 ribs, starting varices, third whorl with 3 low varices and 2 intervarical ribs of similar height, fourth to last whorl with 3 high, narrow, rounded varices and 3 narrow, high intervarical ribs. Spiral sculpture of high, strong, naiTow, nodose, primary, secondary and tertiary cords. First whorl with P1-P3, second and third with IP, Pl- P3, fourth with adis, IP, abis, PI and P2, P3 occasionally covered by subséquent whorl, fifth whorl with adis, IP, abis, PI, si, P2, last teleoconch whorl with t, adis, IP, abis, PI, si, P2, s2, P3, s3, P4, s4, P5, s 5 , P6, s6, ADP, ads, MP. Intersection of axial ribs and varices with spiral cords torming bioad, high, strong nodes. Spiral cords PI and MP giving rise to small, acute spine. Aperture narrow, ovate. Columellar lip narrow, with narrow, elongate folds adapically and abapically with 3-5, elongate, strong, oblique knobs, increasing in strength abapically. Rim partially erect, adhèrent at adapical extremity. Strong pariétal tooth at adapical extremity. Anal notch deep, narrow. Outer lip erect, weakly crenulated, with weak or strong, low, narrow denticles within: ID, D1-D6. ID, D1-D4 split at inner edge of outer apertural lip (Fig. 2). Adapical and abapical denticles higher and stronger, D3 and D4 low. Siphonal canal long, 49-50% of total shell height, narrow, straight, strongly dorsally bent, narrowly open. Operculum brown, ovate, with subapical nucléus and 9 or 10 broad, concentric ridges. Attached surface with about 6 or 7 growth lines and broad callused rim. Etymology. The new species is named for Michèle Colomb, the wife of the second author. Figs 1-2. Siratus michelae n. sp. spiral cords and apertural denticles morphology (paratype RH coll.). Remarks. Siratus michelae n. sp. can be compared with two species only, that also occur off Martinique, although at depth of about 150-200 m. Siratus cailleti (Petit, 1856) (Figs 5-6, 13-16) differs in having a larger shell with a larger and broader protoconch (Figs 5-6), with a width of 1000 pm and a height of 800-900 pm,.lacking the fine abapical keel, in having only two, broader and higher intervarical îibs on penultimate and last teleoconch whorls, and in having a relatively broader siphonal canal. Siratus kugleri (Clench & Pérez Farfante, 1945) (Figs 17-22) most ol the time also has a shell with three 76 intervarical ribs on four or three abapical teleoconch whorls, but the protoconch is larger and also lacks the abapical keel, the shell is relatively larger with a much broader last teleoconch whorl, a broader aperture and a relatively broader siphonal canal. Merle et al (2011, pl. 24, fig. 12) illustrated a shell in MNHN as a syntype of M similis Sowerby, 1841. Murex similis is an earlier name for S. kugleri but a junior homonym of M similis Schrôter, 1805, thus invalid. Siratus kugleri was named as a replacement name for M. similis Sowerby. The specimen from MNHN was brought to the muséum by Sowerby in R. Houart & J. Colomb Novapex 13(2): 75-78, 10 juin 2012 1879 and is most probably the shell that he illustrated himself (Sowerby, 1879: pl. 3, figs 22 & 23 only), but not a syntype. Apparently M. similis Sowerby was described from a single specimen. The shell originally figured by Sowerby (1841: pl. 189, fig. 70) from the Jane Saul collection is in the University Muséum of Cambridge and was catalogued as the holotype by Bishop&Way (1976: 43) (Figs 17-18). The shell morphology and the colour in S. michelae n. sp. differentiate it definitively from any other Western Atlantic Siratus species. Figures 3-6. Protoconchs (scale bars: 500 pm) 3-4. Siratus michelae n. sp., paratype RH coll.; 5-6. Siratus cailleti (Petit, 1856), Martinique, 250 m, JC coll. (specimen illustrated Figs 15-16). Acknowledgements We are very grateful to Richard Preece (University of Cambridge) for the picture of the holotype of Murex similis Sowerby, 1841, to Virginie Héros (Muséum national d'Histoire naturelle, Paris) for the images of one of the syntypes of Murex cailleti and for information about M. similis deposited in MNHN, to Virginie Héros, Richard E. Petit (North Myrtle Beach, South Carolina, U.S.A.) and Charlie Sturm (Carnegie Muséum of Natural History, Pittsburgh, PA, USA) for bibliographical research and to John Wolff, Lancaster, Pennsylvania, USA, for checking the English text. Thanks also to Alan Beu (Institute of Geological & Nuclear Sciences, Lower Hutt, New Zealand) and to Gregory S. Herbert (Department of Geology, University of South Florida, USA) who reviewed this paper. Jacques Colomb also thanks Régis Delannoye, diver and seashell collector from Martinique for his help in many ways. References Bishop, M.J. & Way, K. 1976. Type specimens in the Jane Saul collection, University Muséum of Zoology, Cambridge. Journal of Conchology. 29(1): 41-46. Houart, R. 1999. Two new species of the genus Chicoreus (Siratus) (Gastropoda : Muricidae) from the western Atlantic. The Nautilus 113(4): 121- 126. Houart, R. 2000. Description of two new species of Chicoreus (Siratus) (Gastropoda, Muricidae) from Honduras and Nicaragua. Novapex 1(3-4): 75-82. Houart, R. 2010. Siratus Jousseaume, 1880. Accessed through: World Register of Marine Species ( WoRMS) at http://www.marinespecies.org/aph ia.php?p=taxdet ails&id=405258 on 2012-02-01. Houart, R. 2012. Description of a new species in the Siratus pliciferoides group (Gastropoda: Muricidae) from the Philippines. Novapex 13(1): 25-28. Merle, D. 1999. La radiation des Muricidae (Gastropoda : Neogastropoda) au Paléogène: approche phylogénétique et évolutive . Paris. Unpublished thesis, Muséum national d'Histoire naturelle : i-vi, 499 pp. Merle, D. 2001. The spiral cords and the internai denticles of the outer lip in the Muricidae: terminology and methodological comments. Novapex 2(3): 69-91. Merle, D. & Garrigues, B. 2008. New muricid species (Mollusca, Gastropoda) from French Guiana. Zoosystema 30(2): 517-526. Merle, D. & Garrigues, B. 2011. Description of four new species of Muricidae (Mollusca, Gastropoda) from the Philippines and the Caribbean area. Zoosystema 33(4): 557-575. Merle, D., Garrigues, B. & Pointier, J.P. 2001. An analysis of the sculptural pattern of the shell in Caribbean members of Chicoreus ( Siratus ) Jousseaume, 1880 (Gastropoda, Muricidae), with description of a new species. Zoosystema 23(3): 417-431. Merle, D., Garrigues, B. & Pointier, J.P. 2011 . Fossil and Recent Muricidae of the World -Part Muricinae- Ed. Conchbooks, D-55546 Hackenheim, 648p. Sowerby, G.B. 1834-1841. The Conchological Illustrations , Murex , Sowerby, London: pis 58-67 (1834); pis 187-199 + catalogue: 1-9(1841). Sowerby, G.B. 1879. Thésaurus conchyliorum , vol. 4, pts. 33-34: 1-55, pis. 380-402, London. Vokes, E.H. 1965. Cenozoic Muricidae of the western Atlantic région, pt. II. Chicoreus s.s. and Chicoreus (Siratus). Tulane Studies in Geology andPaleontology. 3(4): 181-204. Vokes, E.H. 1990a. Cenozoic Muricidae of the western Atlantic région. Part VIII - Murex s.s., Haustellum, Chicoreus, and Hexaplex\ additions and corrections. Tulane Studies in Geology and Paleontology. 23(1-3): 1-96. Vokes, E.H. 1990b. Two new species of Chicoreus subgenus Siratus (Gastropoda: Muricidae) from northeastern Brazil. The Nautilus 103(4): 124-130. 77 R. Houart & J. Colomb A new species of Siratus from Guadeloupe Figures 7-22 7-12. Siratus michelae n. sp., Martinique, Atlantic coast, 70 m. 7-9. Holotype MNHN 25131,39 mm; 10-11. Paratype JC coll, 41.4 mm; 12. Paratype RH coll., 41.4 mm. 13-16. Siratus caUleti (Petit, 1856). 13-14. Guadeloupe, syntype MNHN0062, 53.4 mm; 15-16. Martinique, 250 m, JC coll., 59.8 mm; 17-22. Siratus kugleri (Clench & Pérez Fartante, 1945). 17-18. Locality unknown, holotype of Murex similis Sowerby, 1841, Cambridge Mus., 48.6 mm; 19-20. Guadeloupe, lie de la Désirade, 250 m, RH coll., 51.3 mm; 21-22. St. Bartelemy, Banc de Rabet, 150 m, RH coll., 62 mm. 78 R. Le Beon Novapex 13(2): 79-85, 10 juin 2012 Une nouvelle Marginella (Gastropoda: Marginellidae) de la côte occidentale d’Afrique Roger LE BEON 157 avenue docteur Schweitzer, 83160 La Valette du Var, France r 1 ebeon@orange. fr MOTS-CLES. Gastropoda, Marginellidae, Afrique de l’Ouest, Groupe Marginella glabella , nouvelle espèce, sympatrie. RESUME. L’auteur décrit une espèce du genre Marginella Lamarck, 1799 : Marginella pseudodesjardini sp nov. draguée au large de la côte ouest africaine, dans une zone s’étendant du nord de la péninsule du Cap Vert (Sénégal) à la Côte d’ivoire, au niveau circalittoral. Le nouveau taxon est comparé à des espèces similaires : Marginella sebastiani Marche-Marchad & Rosso, 1979, M. desjardini Marche-Marchad, 1957, M. glabella Linnaeus, 1758 et M. psendosebastiani Mattavelli, 2001, dont la distribution est partiellement sympatrique. ABSTRACT. The author describes a species belonging to the genus Marginella Lamarck, 1799: Marginella pseudodesjardini sp. nov., dredged off the west African coast, along an area spreading from North the peninsula of Cap Vert (Sénégal) to the Ivory Coast, at the circalittoral level. The new taxon is compared to the similar species M. sebastiani Marche-Marchad & Rosso, 1979, M. desjardini Marche-Marchad, 1957, M. glabella Linnaeus, 1758 and M. pseudosebastiani Mattavelli, 2004, ranging partially in sympatry. INTRODUCTION Le présent article est consacré à la description d’une espèce nouvelle dans le groupe Marginella glabella. Le groupe M. glabella est constitué par un ensemble d’espèces de taille relativement importante dont les coquilles lisses et globuleuses de 1 à 8 centimètres présentent une coloration rosée à rouge soutenu et ornées de taches pâles en forme de points plus ou moins importants : Marginella. glabella L., 1758, est distribuée du Maroc à la Guinée y compris dans les archipels des Canaries et du Cap Vert. Marginella. desjardini Marche-Marchad, 1957 est distribuée du sud de la Mauritanie au nord du golfe de Guinée. Marginella sebastiani Marche-Marchad et Rosso, 1979 est distribuée du Sénégal à la Guinée. Marginella psendosebastiani Mattavelli, 2001 est distribuée du sud de la Mauritanie jusqu’à l’extrême nord du Sénégal. On peut aussi rapprocher à cet ensemble Marginella aurantia Lamarck, 1822 et Marginella lamarcki Boyer, 2004, dont les tailles sont néanmoins radicalement inférieures, la silhouette de la coquille étant nettement plus cylindrique et le système de décoration en nappes blanches étant très différent du système de décoration ponctué ou tacheté rencontré dans le reste du groupe M. glabella. On écarte aussi de cette étude la comparaison avec Marginella goodalli Sowerby, 1825, caractérisée par une coquille de forme trapue à l’épaule anguleuse. On écarte enfin de cette étude la comparaison avec Marginella irrorata Menke, 1828, dont les caractéristiques sont très proches de celles de Marginella glabella mais dont la zone de distribution est limitée au sud marocain et au nord mauritanien, ne présentant ainsi aucune sympatrie avec l’espèce nouvelle décrite ici. Abréviations MNHN: Muséum national d’Histoire naturelle de Paris, France. AT: Collection Alex Trencar MCA: Collection Marie Christine Aron RLB: Collection Roger Le Béon Matériel. Cinq spécimens à l’état sec, ont été réunis par l’auteur. Les données morphométriques complémentaires proviennent des collections de M. Alex Trencar (11 échantillons), Mme Marie Christine Aron (14) et M. Gilles Granpoder (1). La provenance est donnée de l’Afrique de l’ouest, sous-produit des dragages halieutiques intensifs pratiqués depuis de nombreuses années dans la zone du plateau continental. M.C. Aron signale que ses exemplaires proviennent plus précisément de Côte d’ivoire au large d’Abidjan. Par ailleurs, l’exemplaire présenté de 79 R. Le Beon Une nouvelle Marginella façon erronée comme M. desjardini par Marche- Marchad (1957 : fig. 6) a été collecté au large de Dakar. Enfin, l’auteur a pu identifier de nombreuses illustrations de coquilles de cette marginelle dans différents ouvrages et sur des sites web dédiés à la conchyliologie, où elle est généralement désignée comme M. desjardini et plus rarement comme M sebastiani. Trente-quatre échantillons ont ainsi pu être exploités pour les données morphométriques. Faute d’avoir obtenu des exemplaires vivants ou conservés dans l’alcool, les parties molles n’ont pu être étudiées. Historique. L’auteur a signalé dans un article antérieur (Le Béon. 2010) la découverte de trois exemplaires de forme et décor identiques qui, au premier abord ressemblaient à Marginella sebastiani, mais surtout à Marginella desjardini. Constatant aussi que ces coquilles étaient souvent confondues dans les publications avec Marginella desjardini , fauteur a proposé de distinguer cette forme sous la dénomination de Marginella cf. desjardini. A la suite de cet article, des données relatives à plus de 30 exemplaires provenant de diverses collections ont été rassemblées . L’étude de ce matériel a conduit l’auteur à émettre l’hypothèse de l’existence d’une nouvelle espèce (Le Béon, 2011), dont la description fait l’objet du présent article. SYSTEMATIQUE Famille MARGINELLIDAE Sous famille MARGINELLINAE Genre Marginellidae Fleming 1823 Espèce type par monotypie: Marginella glabella Linnaeus, 1758 Marginella pseudodesjardini n. sp. Figs 9-12 Matériel type. Flolotype MNHN 23796, Sénégal, 62,01 mm x 28,8 mm. Paratype 1, Sénégal, 60,2 mm x 28,7 mm. Paratype 2, Sénégal, 49,15 mm x 23,6 mm. Paratype 3, Sénégal, 45,1 mm x 21,5 mm (paratypes coll. RLB). Localité type. Sénégal. Distribution. L’holotype et les paratypes sont originaires du Sénégal. Les autres coquilles dont la localisation est certaine sont : La coquille (Fig. 6) figurée dans l’article de Marche- Marchad & Rosso (1979) provenant de "Dakar, Sénégal". Le lot de 14 coquilles de M.C. Aron provenant de la région d’Abidjan. L exemplaire que signale Fabio Mattavelli, d’une taille exceptionnelle de 74 mm, provenant des environs de l’île de Gorée au Sénégal. Un exemplaire de 54 mm signalé sur le site de Fabio Mattavelli proviendrait de Casamance au sud du Sénégal. La zone de distribution de Marginella pseudodesjardini n. sp s’étend donc au moins de la presqu’île du Cap Vert, où elle vit en sympatrie avec Marginella sebastiani et Marginella desjardini , jusqu’à la Côte d’ivoire. Description. Coquille épaisse, brillante, biconique, étroitement fusiforme. La spire est formée de 5 à 6 tours au profil légèrement courbe, la courbure étant plus marquée du côté adapical. Le dernier tour de spire est légèrement convexe dans le bas. Il est très développé et égal en hauteur à plus de 4 fois le reste de la coquille. La suture est légèrement oblique à gauche et bien accusée. Le dernier tour va également en s'atténuant vers le bas jusqu'au canal abapical (antérieur) qui est tronqué. Sommet obtus, mamelonné, bien distinct. La plus grande largeur est située au tiers postérieur du test. Ouverture égale à plus des 2/3 de la hauteur totale, sensiblement plus étroite vers le haut, légèrement élargie vers le bas. Labre fortement marginé à l'extérieur, souvent épaissi à l'intérieur, sauf vers son bord adapical où cet épaississement s'atténue, laissant une légère gouttière en relation avec le canal. Sa jonction avec la zone adapicale se situe entre un et trois millimètres au- dessus de la suture du dernier tour. L’épaississement porte vers le milieu une dizaine de denticulations obtuses dont la base commune contribue encore à épaissir le milieu du bord interne du labre donnant à celui-ci sur sa face interne un dessin en forme de S inversé et très étiré. Ces denticulations disparaissent ou s'atténuent aux deux extrémités de la coquille. Le profil extérieur du labre est très courbe dans sa partie adapicale puis presque rectiligne sur les 2/3 suivants pour finir avec une courbure moins accentuée dans sa partie abapicale. La columelle est oblique et à peu près rectiligne, munie de 4 plis en forme de lamelles qui convergent vers l’intérieur, presque horizontales pour les deux postérieures et presque verticales pour les deux adapicales. Bord columellaire étalé et bien limité vers l'avant, se reliant avec la marge postérieure. La coloration consiste en un fond jaune pâle lavé de beau rouge carnéolé sur lequel se détachent trois zones rouges plus sombres alternées avec trois zones plus claires. L’ensemble du dernier tour est irrégulièrement maculé de taches blanc-crème. La première zone rouge, sous la suture du dernier tour, est ornée de macules en forme de flammes longitudinales qui sont prolongées sur le reste du dernier tour par des macules vaguement quadrangulaires qui semblent approximativement organisées en lignes longitudinales. Cette zone apparaît aussi sur les tours de la spire. 80 R. Le Beon Novapex 13(2): 79-85, 10 juin 2012 Variations. Elles portent sur la forme générale plus ou moins fusiforme mais qui semble remarquablement stable avec une variation de l’élongation (rapport de la longueur maximum sur la largeur maximum) très faible. - La taille peut varier de 38 mm de haut pour les échantillons observés, allant jusque 74 mm. - La surépaisseur du labre plus ou moins marquée, jusqu’à 20 dents, mais parfois quasi obsolètes pour les exemplaires plus matures. - La densité plus ou moins forte des flammes sous la suture, mais toujours existantes. - Le nombre et la forme des macules et la coloration du dernier tour plus ou moins foncée qui peut être de couleur jaunâtre uniforme par atténuation relative des zones rouges. Remarques. Cette marginelle de grande taille pour le genre, est très proche de Marginella desjardini pour sa taille et sa forme, mais aussi de Marginella sebastiani. Elle est aussi proche de Marginella pseudosebastiani surtout pour son décor presque identique. Un examen croisé des caractéristiques principales de ces espèces permet néanmoins de distinguer la forme Marginella pseudodesjardini avec certitude. Les caractéristiques comparées sont: • L’élongation et la taille de la coquille ainsi que l’aspect et la répartition des macules sur le décor du dernier tour, qui constituent les caractères principaux. • L’épaisseur et la forme du labre ainsi que la position de la jonction entre le labre et la suture du dernier tour, qui constituent des caractères secondaires. • La présence plus ou moins marquée de flammules au niveau de la suture et le dernier tour qui constitue un caractère secondaire. 1. Elongation SYNTHESE ELONGATION L/I Taille maximum échantillons en millimètres Espèce Nbre échantillons L/l min L/l moyen L/l max Dispersion* M. sebastiani 29 1,6 1,82 2,03 0,43 71,4 M. pseudosebastiani 18 1,7 1,85 1,94 0,24 54,05 M. glabella 19 1,7 1,84 1,97 0,27 58,12 M desjardini 21 1,91 2,17 2,44 0,53 72,8 M. pseudodesjardini 34 1,94 2,07 2,35 0,41 74 * caractérise par espèce, l’amplitude de la variation de l’élongation moyenne. Les mesures comparatives révèlent la proximité des ensembles M sebastiani-M. pseudosebastiani-M. glabella : 1,82 /1,85/1,84, et M desjardini-M, pseudodesjardin i: 2,17/2,07, avec une hiérarchie croissante M sebastiani , M. glabella , M. pseudosebastiani , M ps eudo desjardini, M. desjardini. Même si certains résultats sont peu divergents quand on considère les ensembles ainsi mis en évidence, les données lissées par contre, révèlent une claire divergence entre les cinq formes considérées. M. desjardini et M pseudodesjardini se distinguent fortement des trois autres espèces par le rapport L/l et divergent entre elles pour ce rapport. Pour ce même rapport les trois espèces M. sebastiani , M. glabella , M. pseudosebastiani sont pratiquement indiscernables. 81 R. Le Beon Une nouvelle Marginella Elongation L/l 1,50 0 10 20 30 N° échantillons classés par longueur — M. Sébastiani - M. pseudosebastiani - -à. - M. pseudodesjardini - m - M. desjardini M. glabella Linéaire (M. sebastiani) Linéaire (M. pseu doseb astia ni) Linéaire (M. pseu dodesjardini) Linéaire (M. desjardini) Linéaire (M. glabella) 40 2. Macules blanchâtres sur le dernier tour M. sebastiani Grosses. Répartition aléatoire sur tout le dernier tour. Forme ronde. M. glabella Petites, parfois organisées en treillis. Parfois obsolètes. M. pseudosebastiani Petites. Répartition aléatoire sur tout le dernier tour. Forme ronde. M. pseudodesjardini Quadrangulaires. Répartition aléatoire sur tout le dernier tour. Parfois en lignes longitudinales M. desjardini Quadrangulaires. Répartition aléatoire sur tout le dernier tour sauf sur les trois bandes claires. Parfois en lignes longitudinales L’absence de taches sur les trois bandes claires chez Marginella desjardini est un critère très discriminant. Elle est la seule du groupe à avoir cette caractéristique. 3. Surépaisseur médiane du labre M. sebastiani Quasi inexistante, mais présences de denticulations M. glabella Inexistante, mais présences de denticulations M. pseudosebastiani Inexistante M. pseudodesjardini Assez fréquente mais de faible ampleur M. desjardini Très fréquente et parfois très marquée Ce critère secondaire permet de séparer les couples M. sebastiani!M. pseudosebastiani et M. desjardini!M. pseudodesjardini. 82 R. Le Beon Novapex 13(2): 79-85, 10 juin 2012 4. Position de la jonction entre le labre et la suture du dernier tour. Jonction du labre sur la spire 2,5 2 1,5 E E t 03 O ‘ - M.sebastiani —□ - M.desjardini —sir— M. pseudodesjardini — — Logarithmique (M. sebastiani) “■ ■ Logarithmique (M. desjardini) . Logarithmique (M. pseudodesjardini) Ce critère permet de séparer aisément Marginella desjardini des deux autres espèces. 5. Flammes au niveau de la suture M. sebastiani Rarement présentes et alors sur les grosses tailles et en petit nombre M glabella Très fréquentes M. pseudosebastiani Très fréquentes M. pseudodesjardini Fréquentes sauf parfois sur les grosses tailles M. desjardini Très fréquentes sauf sur certains spécimens anormaux. 6. Comparaison synthétique des données morphométriques de M. pseudodesjardini avec M. sebastiani , M. glabella , M. pseudosebastiani et M. desjardini M. pseudodesjardini - M. sebastiani L’élongation de M pseudodesjardini est nettement plus forte : 2.07 pour 1.82. Le labre de M. pseudodesjardini est souvent plus épaissi. Les flammes sont souvent absentes chez M. sebastiani. I es macules sont rondes et plus grosses chez M. sebastiani M. pseudodesjardini-M. glabella L’élongation de M. pseudodesjardini est nettement plus forte : 2.07 pour 1.84. Le labre est dépourvu de surépaisseur chez M. glabella. Les macules sont plus petites et souvent organisées en treillis chez M. glabella. M. pseudodesjardini-M. pseudosebastiani L’élongation de M pseudodesjardini est nettement plus forte : 2.07 pour 1.85. Le labre est dépourvu de surépaisseur et de denticulations chez M. pseudosebastiani. I es macules sont plus petites et rondes chez M. pseudosebastiani. M. pseudodesjardini-M. desjardini L’élongation est légèrement plus faible chez M. pseudodesjardini : 2.07 pour 2.17. La distance entre la jonction du labre et la suture de la spire sur le dernier tour est nettement plus faible chez M. desjardini : 0.5 pour 2. Les macules sont absentes sur les trois bandes claires du dernier tour chez M. desjardini. 83 R. Le Beon Une nouvelle Marginella Mattavelli (2001) a décrit M. pseudosebastiani de Mauritanie. Elle est très voisine de Marginella glabella dont elle partage en partie le biotope. A partir de mes articles (Le Béon, 2010 et 2011) et de ses propres recherches il publie un nouvel article (Mattavelli, 2011) où il propose trois hypothèses concernant Marginella pseudodesjardini : • En premier lieu il proposerait qu'elle soit adoptée en tant que sous-espèce de Marginella pseudosebastiani. • Deuxièmement, en se fondant sur une hypothèse d’hybridation entre Marginella pseudosebastiani et Marginella pseudodesjardin i ou entre Marginella sebastiani et Marginella desjardini, il proposerait Marginella klepton pseudosebastiani • En troisième lieu enfin, il considérerait Marginella pseudodesjardini comme une espèce valide. Concernant la première hypothèse seuls les critères de décor convergeants entre Marginella pseudosebastiani et Marginella pseudodesjardini permettent de soutenir cette hypothèse. Par contre l’élongation et la tendance à l’épaississement du labre chez Marginella pseudodesjardini permettent de séparer les deux formes sans intermédiaires avérés. La deuxième hypothèse ne serait démontrée que si les zones de dispersion de ces espèces étaient confondues ou au minimum contigües, ce qui ne semble pas être le cas. La troisième proposition est donc la plus probable tant que les hypothèses d’hybridation ne sont pas démontrées. Remerciements Mes remerciements vont à M. Jack Basset (Rennes, France) pour la fourniture de l’holotype, M. Alex Trencar (Istres, France), Mme Marie Christine Aron (Le Puy Sainte Réparade, France), M. Gilles Granpoder (Les Paluds de Noves, France) et Jean François Michard (Garéoult, France) pour la fourniture de données morphométriques et d’images, ainsi qu’à Mme Virginie Héros (Muséum national d'Histoire naturelle, Paris) pour la documentation fournie, M. Franck Boyer (Sevran, France) pour ses conseils éclairés et M. Flavio Mattavelli (Gorgonzola, Italie) pour l’intérêt qu’il a porté à cette étude et les images fournies. Un remerciement particulier pour M. Roland Houart (Landen, Belgique) pour sa grande patience. REFERENCES Le Béon R. 2010. Marginella cf. desjardini. Xenophora 129: 28-30. Le Béon R. 2011. Marginella cf. desjardini (suite). Xenophora 133: 17. Marche-Marchad, I. & Rosso, J.-C., 1979. Une nouvelle marginelle de la côte occidentale d'Afrique: Marginella sebastiana sp. nov. (Gastropoda, Marginellidae). Bollettino Malacologico 15(7-8): 197-208. Marche-Marchad, I. 1957. Description de cinq gastropodes marins nouveaux de la côte occidentale d'Afrique. Bulletin du Muséum National d'Histoire Naturelle 2° série, 29(2): 200- 205. Mattavelli, F. 2001. Marginella pseudosebastiani. Malacologia 34: 3-8. Mattavelli, F. 2011. Marginella sp. oppure kl. pseudodesjardini?Malacologia 70: 16. Figures 1-22 1. Marginella glabella 52.2 mm, Mauritanie, coll. RLB; 2. Marginella pseudosebastiani 49 mm, Mauritanie, coll. RLB; 3. Marginella sebastiani 52.8 mm, Sénégal, coll. RLB; 4. Marginella pseudodesjardini 60.2 mm, Sénégal, paratype 1, coll. RLB; 5. Marginella desjardini 55.5 mm, Sénégal, coll. RLB; 6-7. Coquille figurant dans l’article de Marche-Marchad & Rosso (1979). 8-9. Holotype de Marginella sebastiani .MNHN 55.4 mm; 10-11. Holotype de Marginella desjardini. MNHN 70.8 mm 12-19. Marginella pseudodesjardini 12-13. Holotype (MNHN 2396) (longueur x largeur) 62,01 x 28,8 mm. Hauteur du dernier tour 49,46 mm, hauteur de l'ouverture 48,7 mm; 14-15. Paratype 1 (Coll. RLB) 60,2 x 28,7 mm; 16-17. Paratype 2 (Coll. RLB) 49,15 x 23,6 mm; 18-19. Paratype 3 (Coll. RLB): 45, 2 x 21, 5 mm. 20-22. Jonction de la lèvre par rapport à la suture du dernier tour 20. Marginella pseudodesjardini ; 21. Marginella desjardini ; 22. Marginelli sebastiani. 84 • • R. Le Beon NOVAPEX 13(2): 79-85, 10 juin 2012 J. Trôndle & J. Letourneux Novapex 13(2): 87-90, 10 juin 2012 Description de Turbo fakaauensis n. sp. (Mollusca: Gastropoda: Turbinidae) du Pléistocène de Niau, Tuamotu (Polynésie française) Jean TRÔNDLÉ Attaché au Muséum national d'Histoire naturelle Département Systématique et Évolution 55, rue de Buffon, 75005 Paris, France j. trond le@orange. fr Jean LETOURNEUX Mahina, Tahiti, Polynésie française natual ey la@ma i 1. pf MOTS-CLEFS. Mollusca, Gastropoda, Turbinidae, Polynésie française, Pléistocène. KEYWORDS. Mollusca, Gastropoda, Turbinidae, French Polynesia, Pleistocene. RÉSUMÉ. Une nouvelle espèce Turbo fakaauensis du Pléistocène est décrite de Niau, Archipel des Tuamotu (Polynésie française) et est comparée à Turbo crassus Wood, 1828, espèce proche actuelle de l'Indo-Ouest Pacifique. ABSTRACT. Turbo fakaauensis n. sp. is described from Niau, Tuamotu Archipelago (French Polynesia) dated Pleistocene and is compared with Turbo crassus Wood, 1828, a quite similar présent Indo-West Pacific species. INTRODUCTION Les tests de Turbo fakaauensis n. sp. ont été extraits de sables coralliens détritiques prélevés en bordure de lagon de l'atoll de Niau. L'altitude moyenne de l'atoll est actuellement de 7,5 m et les sables récoltés correspondraient à un niveau marin du dernier interglaciaire (Pléistocène, environ 125 000 ans), de 6 à 10 m plus élevé qu'actuellement (Trôndlé & Salvat, 2010). Dans ces sédiments détritiques 12 familles de bivalves et 33 familles de gastropodes sont représentées. Au total 121 espèces de mollusques (29 bivalves et 92 gastropodes) ont été répertoriées et sont en cours d'étude. Chez les bivalves ce sont les Tellinidae qui dominent avec 8 espèces et chez les gastropodes ce sont les Cerithiidae avec 11 espèces. Les Turbinidae ne sont représentés que par deux espèces et une seule dans le genre Turbo. Deux nouvelles espèces de mollusques gastropodes ont déjà récemment été décrites Strombus blanci Trôndlé & Salvat , 2010 et Terebra niauensis Trôndlé & Letourneux, 2011. Abréviations BMNH: The Natural History Muséum, London, U.K. CRIOBE: Centre de Recherches Insulaires et Observatoire de f Environnement, Moorea, Polynésie Française. EPHE: École Pratique des Hautes Études, Perpignan, France. MNHN: Muséum national d'Histoire naturelle, Paris, France. ZSM: Zoologische Staatssammlung München, Deutschland. JL: Collection Jean Letourneux. JT: Collection Jean Trôndlé. PF: Polynésie française. SYSTÉMATIQUE Famille TURBINIDAE Rafinesque, 1815 Genre Turbo Linnaeus, 1758 Espèce type: Turbo petholatus Linnaeus, 1758, par désignation subséquente, Montfort, 1810. Turbo fakaauensis n. sp. Figs 1-2, 5-6 Matériel type. Atoll de Niau, Tuamotu, Polynésie française, 16°08’S, 146°20’W, holotype, 102 mm, MNHN 24607 (Figs. 1-2). Paratypes: 49,5 mm, MNHN 24608 (Fig. 5); 103,5 mm, CRIOBE (Fig. 6); 84,6 mm, JL; 80 mm, JT. Localité type. Atoll de Niau, Tuamotu, Polynésie française, 16°08’S, 146°20’W. Description de P holotype. Coquille lourde, épaisse, turbinée et globuleuse, mesurant 102 mm de hauteur et 87 mm de diamètre. Le test usé est de couleur blanche. La spire est peu élevée. L'apex est érodé. Le 87 J. Trôndle & J. Letourneux Turbo fakaauensis n. sp. dernier tour est ample et marqué d'une zone sous- suturale légèrement concave, absente des premiers tours. L'ouverture est nacrée, large, arrondie, bordée par un bourrelet columellaire et un large cal obturant l'ombilic et se projetant vers l'avant. L'ouverture, au péristome brisé dans sa partie antéro-exteme, mesure 74 mm de hauteur et occupe ainsi plus de la moitié de la hauteur totale de la coquille. Le test est parcouru par de nombreuses rides circulaires fines, une cinquantaine sur le dernier tour. Des stries de croissance bien marquées près de l'ouverture ne sont à peine visibles qu'à fort grossissement sur le reste du test. Distribution. Uniquement connue de la localité type. Remarques. La taille maximale observée est de 103,5 mm (paratype, fig. 6) chez un individu dont l'apex est érodé et la partie antérieure de la coquille tronquée. La protoconque est absente chez les cinq exemplaires examinés. Aucun opercule n'a été récolté. Un des paratypes (Fig. 6) présente 4 cavités et 1 perforation sur la face ventrale du dernier tour, alignées parallèlement à la columelle. Ces cavités, d environ 10 mm de diamètre et 10 mm de profondeur s’élargissent très légèrement de la surface du test vers le fond. Des empreintes en relief identiques au fond de toutes les cavités permettent d’affirmer que ces dernières ont été creusées par des bivalves lithophages [Lithophaga (Mytilidae) ou Gastrochaena (Gastrochaenidae)]. La perforation correspond à un orifice situé en arrière des cavités. 11 espèces de lithophages (8 Mytilidae et 3 Gastrochaenidae) ont été répertoriées en Polynésie française (Trôndlé & Boutet, 2009); aucun de ces mollusques perforants n’a été trouvé dans le gisement de subfossiles du pléistocène où ont été récoltés les tests de Turbo fakaauensis n. sp. Turbo fakaauensis n. sp. est proche, par son allure générale, de Turbo crassus Wood, 1828 (Fig. 3-4), absent en Polynésie française. Cependant, d'aspect moins massif, T. crassus est de taille plus modeste, de 60 à 80 mm, et possède une spire plus élevée. L'épaule de T. crassus est marquée d'un épais cordon chez l'adulte, caractère que l'on ne retrouve pas chez T. fakaauensis. Par ailleurs la sculpture du test de T. crassus est constituée de fortes cordes spirales, d'épaisseur inégale, entrecoupées de fines stries axiales, alors que le test de T. fakaauensis est parcouru par de nombreuses rides spirales sensiblement d'égale importance. L'imposant cal columellaire se projetant en avant de l'ouverture chez T. fakaauensis est absent chez T. crassus . Le récent inventaire des mollusques de Polynésie Française (Trôndlé & Boutet, 2009) fait état de la présence de 4 espèces du genre Turbo : Turbo argyrostomus Linnaeus, 1758 est une espèce pouvant atteindre 100 mm, mais dont le test est parcouru d'épais cordons circulaires souvent squameux chez l'adulte, caractère absent cher T. fakaauensis. Turbo petholatus Linnaeus, 1758, possède un test lisse et brillant, dépasse rarement 65 mm en Polynésie, mais peut atteindre exceptionnellement 100 mm. Turbo mannoratus Linnaeus, 1758, introduit en Polynésie française dans les années 60, est une espèce de grande taille atteignant 220 mm de hauteur qui présente à un stade juvénile une certaine ressemblance avec T. fakaauensis. Cependant T. mannoratus est dépourvu de sculpture spirale et est parcouru de très nombreuses stries de croissances bien marquées, caractères que l'on ne retrouvent pas chez T. fakaauensis, Par ailleurs son aspect général est moins globuleux et la spire est plus élevée. Enfin, un épais cordon noduleux orne très tôt l'épaule de T. mannoratus. Turbo setosus Gmelin, 1791 peut atteindre 95 mm, mais à la différence de T. fakaauensis son test est sculpté d'épais cordons circulaires, alternant avec des cordons plus fins. Alors que les sédiments bioclastiques de Niau n'ont révélé qu'une seule espèce de Turbo , les études sur la faune malacologique du Pléistocène de l'Indo- Pacifique mentionnent la présence fréquente de plusieurs espèces du genre Turbo: Tongatabu [Ostergaard , 1935 (T. argyrostomus, commun, T. crassus, T. petholatus)], Hawaii [Ostergaard, 1939 (7’. intercostalis, commun, T. sp.)], Henderson [Spencer & Paulay, 1989 (T. argyrostomus, commun)], Guam [Ladd, 1966 (T. argyrostomus, T clvysostomus)], Vanuatu (Nelles Hébrides) [Ladd, 1966 (T. argyrostomus , T chrysostomus ), Ladd, 1982 (T. petholatus )], Okinawa [MacNeil, 1960 (T. petholatus , T. argyrostomus)]. Etymologie. L'espèce est nommée d'après la localité type, Fakaau (nom Paumotu de l’atoll de Niau). Remerciements Nous remercions Axel Alf (ZSM) et Suzanne Williams (BMNH), spécialistes des Turbinidae, pour leurs avis sur cette nouvelle espèce, Philippe Maestrati (MNHN) pour les photos et la réalisation de la p ancie, Piene Lozouel (MNHN) pour ses remarques dans le manuscrit et ses commentaires, Bernard Salvat (EPHE) pour ses commentaires sur les cavités de lithophages sur un des tests et la relecture du manuscrit, ainsi que ceux qui d'une façon ou d'une autre nous ont aidés à réaliser cette étude: Philippe Boutet (PF), Robert Gourguet (PF). REFERENCES Ladd, H.S., 1966. Chitons and gastropods (Haliotidae îoug Adeorbidae) ffom the western Pacific islands. G eological Survey Professional Paper, 531: 98p. 88 J. Trôndle & J. Letourneux Novapex 13(2): 87-90, 10 juin 2012 1-2,5-6. Turbo fakaauensis n. sp.; 1, 2. Holotype MNHN 24607, 102 mm; 5. Paratype MNHN 24608 49 5 mm (sculpture spirale); 6. Paratype CRIOBE, 103,5 mm (cavités dues à des lithophages). 3-4. Turbo crassus Wood, 1828, 72 mm, îles Salomon (photos Axel Alf). 89 J. Trôndle & J. Letourneux Turbo fakaauensis n. sp. Ladd, H.S., 1982. Cenozoic fossil mollusks from western Pacific islands; Gastropods (Eulimidae and Volutidae through Terebridae). Geological Survey Professional Paper , 1171: 100p. MacNeil, F.S., 1960. Tertiary and quaternary Gastropoda of Okinawa. Geological Survey Professional Paper, 339: 148p. Ostergaard, M.J., 1935. Recent and fossil marine Mollusca of Tongatabu. Bernice P. Bishop Muséum , Bulletin 131: 3-59. Ostergaard, M.J., 1939. Report on fossil Mollusca of Molokai and Maui. Occasional Papers of Bernice P. Bishop Muséum , Honolulu, Hawaii, 15(6): 57- 77. Spencer, T. & Paulay, G., 1989. Geology and geomorphology of Henderson Island. Atoll Research Bulletin , 323: 18p. Trôndlé, J. & Boutet, M., 2009. Inventory of marine molluscs of French Polynesia. Atoll Research Bulletin , 570: 87p. Trôndlé, J. & Letourneux J., 2011. Description de Terebra niauensis n. sp. (Mollusca: Gastropoda: Terebridae) du Pléistocène de Niau, Tuamotu (Polynésie Française). Novapex, 12(3-4): 87-90. Trôndlé, J. & Salvat, B, 2010. La thanatocènose du lagon de Fatoll de Niau (Polynésie française) avec la description d’une nouvelle espèce de Slrombus (Mollusca, Gastropoda, Strombidae). Zoosystema, 32(4): 613-623. 90 NOTE AUX AUTEURS r • ' aipc I •affiliation à la Société n'est pas obligatoire pour les auteurs. La publication des articles de maximum 12 pages imprimées en double interligne est numéro chaque page sera facturée au prix de 40,00 €. Les articles de taille supérieure peuvent être scindés sur plusieurs numéros. 9ra méros hors série sont publiés irrégulièrement. 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