OCCASIONAL PAPERS

OF THE

California Academy of Sciences

No. 66, 13 pages, 13 figures.

June 27, 1968

Descriptions of New Species of
Gastropods from Clipperton Island

By

Leo G. Hertlein and Edwin C. Allison

M

rviurine Bioiogicuf Laboratory

LIBRARY

JUL 9 1968

WOODS HOLE, mss.

SAN FRANCISCO

PUBLISHED BY THE ACADEMY
1968

OCCASIONAL PAPERS

OF THE

CALIFORNIA ACADEMY OF SCIENCES

No. 66, 13 pages, 13 figures. June 27, 1968

Descriptions of New Species of
Gastropods from Clipperton Island

By

Leo G. Hertlein='= and Edwin C. Allison**

A collection of molluscan shells from Clipperton Island in the eastern Pacific
(Latitude 10°17'00" N., Longitude 109° 13'00" W.) assembled by the junior
author in 1958 (see Allison, 1959) contained a number of species of gastropods,
many of which were recorded in an earlier paper by the present authors (Hertlein
and Allison, 1966). Species of gastropods not included or cited with provisional
identification in that paper have received additional study and, as they ap-
parently are undescribed new forms, descriptions of them are furnished in the
present paper.

Acknowledgments are due several individuals who aided us in the present
study. Dr. Alison Kay, Department of Science, University of Hawaii, compared
specimens of the new Colubraria with others in the Bernice P. Bishop ^Museum
in Honolulu, Hawaii. Dr. and Mrs. G Dallas Hanna, California Academy of
Sciences, aided in the measurement of specimens and in the retouching and ar-
rangement of the illustrations. Mr. Allyn G. Smith, of the same institution, aided
with the identifications and contributed constructive criticism of the manuscript;
Mr. Maurice Giles, California Academy of Sciences, prepared the photographs
used to illustrate two of the species. The line drawings reproduced as text figures
were made by Mrs. May Bios, scientific illustrator, University of California. An
exchange of specimens with Dr. William K. Emerson and Mr. William Old, Jr.,
American Museum of Natural History, made available specimens of Colubraria
for comparative study.

The holotype specimens of the new species and all other specimens illustrated
in this paper are in the University of California Museum of Paleontology, In-
vertebrate Type Collection, Berkeley, California. Paratypes, where available, are
in the California Academy of Sciences, Department of Geology, Type Collection.

* Curator of Invertebrate Paleontology, California .Academy of Sciences.
** Associate Professor, Department of Geology, San Diego State College.

CALIFORNIA ACADEMY OF SCIENCES

[Occ. Papers

Description of Species

Cyclostremiscus solitarius Hertlein and Allison, new species.

(Figure 1.)

Cyclostrcnia cingidijera A. Adams, Hertlein and Allison, The Veliger, vol. 9, no. 2, p. 1.^9,

1966.
Not Cyclostrema cingidifera A. Adajvis, Proc. Zool. Soc. London for 1850, p. 43, issued

November 12, 1850. "Hab. Dumaguete, island of Zebu, 4 fathoms; H. C. (Mus. Cuming.).'

Description. Shell very small, discoid, nuclear whorl followed by 2 regularly
expanding rounded whorls; spire deeply depressed, umbilicus broadly funnel-
shaped; last whorl with a sharp, serrate, spiral keel on the periphery, two similar
ones, one a short distance above, the other below the central keel, also on each
side above the margin of the umbilicus is a spiral row of fine nodes and at the
margin of the umbilicus there is a trace of a very fine spiral threadlet; the sur-
face between the keels is sculptured with very fine spiral striae, and on the early
portion of the first whorl there are faint transverse markings between the basal
and the succeeding keel; peristome circular, the margin fairly thick, slightly
flaring; operculum circular, the exterior with faint traces of concentric lines.
Maximum diameter .74 mm., minimum diameter .57 mm., maximum height of
bodv whorl .38 mm.

Figure 1. Cyclostremiscus solitarius Hertlein and Allison, new species. Holotype no. 37121
(UC), from Clipperton Island, la, basal view; lb, apertural view; maximum diameter 0.74 mm.

Holotype no. 37121, University of California Museum of Paleontology, In-
vertebrate Type Collection, from Locality B-4241, outer portion of intertidal
bench with prominent Pocillopora, brown algae, and slight amounts of coralline
algae, Clipperton Island.

Remarks. The generic assignment of this new species may be open to ques-
tion, but it bears a general resemblance to some species of Cyclostremiscus. The
general shape and character of coiling resembles that of the form described as

Xo. 66 1 HERTLELX & ALLISON: NEW CLIFPERTUN ISLAND GASTROPODS 3

Cyclostremella venezuclana Weisbord (1962, p. 136, pi. 9, figs. 7, 8; pi. 12,
figs. 20-22), from strata of late Cenozoic age. However, according to Moore
(1966, p. 480) the Venezuelan species is not referable to the genus Cyclostremella.
The illustration given by Moore of Cyclostrema humilis Bush, the type
species of Cyclostremella, reveals the presence of a slight depression below the
suture and the operculum is somewhat elongated and paucispiral. The whorls
of the new species from Clipperton Island are rather evenly rounded and the
operculum is circular in outline.

Amphithalamus trosti Hertlein and Strong.

(Figure 2.)

Amphithalamus trosti Hertlein and Strong, .\llan Hancock Pac. Exped., vol. 2, no. 12,
p. 228, pi. 21, fig. 2, Augu.st 21, 19.^9. "Dredged in from .? to 9 fms. in Bahia Honda,
Panama."

Figure 2. Amphithalamus trosti Hertlein and Strong. Hypotype no. 37122 (UC), from
Clipperton Island. Length LIS mm.

One minute, glassy shell found in shell washings closely resembles Amphi-
thalamus trosti. It differs from the type specimen of that species in that the whorls
are slightly more rounded and impressed at the suture. Only a faint trace of a
peripheral threadlet is present but this feature apparently varies somewhat in a
series of specimens from the type locality of A. trosti. The similarity of the
present specimen to the Panamanian shell is so great that we assign it to that
species.

Heliacus infundibuliformis strigatus (Hanley).

(Figure 3.)

Solarium injundibiiliforme variety ( ?) strigata Hanley, Thes. Conch., vol. 3, Solarium, p. 243,
pi. 254, {Solarium, pi. 5), fig. 94, 1863. Expl. to pi. 5, "Sandwich Isles (var. Chemnitzii).''

CALIFORNIA ACADEMY OF SCIENCES

[Occ. Papers

Not Solarium chemnitzii Kiener, Spec. Gen. et Icon. Coq. Viv., Fam. Turbinacees, Solarium,
p. 12, pi. 4, fig. 8, 1838-1839. "Habite la mer des Indes I'ile Bourbon et la mer de la
Chine." According to Tryon, 1887 this = Trochus injundibulijormis Gmelin, Linnaeus
Syst. Nat., Tom. 1, Pars. 6, p. 3575, 1791. "Habitat rarissimus. . ." (Ref. to Chemnitz,
Bd. 5, pi. 173, figs. 1706, 1707, 1781. "Ihr Vaterland ist mir unbekannt.")

Tonnia inju)idihulijorme Gmelin var. strigala Hanley, Tryon, Man. Conch., vol. 9, p. 20,

pi. 6, fig. 99, 1887. (No locahty cited.)
Heliacus infundibulum strigata Hanley, Hertlein and Allison, The Veliger, vol. 9, no. 2,

p. 140, 1966. CHpperton Island.

A specimen from Locality B-4237 (A-588-1), beach on CHpperton Island,
9.5 mm. in maximum diameter and 5.5 mm. high, appears to be referable to the
form described by Hanley as "Solarium injimdibuUjorme variety (?) strigata^
No specific locality was assigned to this variety but localities cited for S. in-
jundibuUjorme were, "Philippines; Lord Hood's Island; Sandwich Isles (var.
Chemnitzii) ."

Figure 3. Heliacus infundibuliformis strigatus (Hanley). Hypotype no. 37123 (UC),
from beach on CHpperton Lsland. 3a, apical view; 3b, apertural view; 3c, basal view; maxi-
mum diameter 9.5 mm.

Hanley's original description follows: "In the variety (?) strigata (f. 94)
there are 3 approximate narrower, and 3 broader belts under the lower peripheral
one: the threads, if any, are most minute: there are 4 or 5 ridges on the umbilical
wall." The coloration was mentioned by Tryon as, "Periphery and the rib below
it ornamented with white and smoky-brown spots."

The present specimen from CHpperton Island agrees so well with the variety
described by Hanley that we refer it to that form.

The form described as Solarium chemnitzii by Kiener was reported from
Hawaii by Hanley. Later, Tryon relegated Kiener's species to the synonymy of the
earlier "Trochus" injundibulijormis Gmelin and considered Hanley's "Solarium
injundibulijorme var. (?) strigata'' to be a valid variety. Tryon also suggested
the possibility that (imelin's species might be identical with the earlier
Turbo crenellus Linnaeus, 1767, and later Bayer (1944) also followed this assign-
ment. Neither H. injundibulijormis nor H. i. strigatus was reported from Hawaii

Xo. 661 HERTLELX & ALLISON: NEW CLIPPERTUS ISLAND GASTROPODS 5

in recent papers by Bryan (1958, suppl. p. 5 ) nor by Robertson ( 1967, p. 247).
However, Dr. Alison Kay informed us (written communication, 5 November
1967) that one shell referable to H. injundibidijormis was dredged (by the Pele)
off Oahu in 40 fathoms.

Odostomia (C^hrysallida) limbaughi Hertlein and Allison, new species.
(Figure 4.)

Description. Shell minute, ovately conic, vitreous. Nuclear whorl about
half immersed at a right angle in the succeeding post-nuclear whorl, and bearing
5 spiral lirations. First and second post-nuclear whorls straight sided, slightly
shouldered at the summit and sculptured with a row of nodules of which about
18 are present on the penultimate whorl. Spiral sculpture consists of flattened
ribs which are separated by incised spiral grooves, the first post-nuclear whorl
with 4, the second 6, the penultimate with 8 and the body whorl with 8 of which
the last 4 are narrower. The periphery and base of the last whorl is well rounded.
The aperture is large, broadly ovate, and angulated posteriorly; outer lip thin,
the base slightly reflected. Length 1.47 mm., maximum diameter .72 mm.

Figure 4. Odostomia (ChrysalUda) limbaughi Hertlein and .\llison, new species. Holo-
type no. 37124 (UC), from Clipperton Island. 4a, apertural view; length 1.47 mm.; 4b, view
of nuclear whorl, greatly enlarged.

HoLOTYPE no. 37124, University of California Museum of Paleontology, In-
vertebrate Type Collection, from Locality B-4241, outer portion of intertidal
bench with prominent PociUopora, brown algae, and slight coralline algae, Clip-
perton Island. E. C. Allison, collector.

This new species is quite different from any species known to us. It bears
a general resemblance to Odostomia (ChrysalUda) sapia Dall and Bartsch (1909,
p. 167, pi. 18, figs. 3, 3a) described from San Diego, California. The new species
differs from 0. (C.) sapia in that the two post-nuclear whorls are straight sided
rather than rounded and these are sculptured with more numerous spiral ribs.

This species is named for Conrad Limbaugh, leader of the E.xpedition to
Clipperton Island during which the type specimen was collected.

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[Occ. Papers

Turbonilla (Pyrgisculus) clippeitonensis Hertlein and Allison, new species.
(Figure 5.)

Description. Shell elongated, nuclear whorls about one and a half, smooth,
the nucleus about one-half immersed, the axis at almost a right angle to that of
the succeeding whorls; post-nuclear whorls 4, decidedly rounded and constricted
at the sutures, sculpture consisting of about 16 slightly curved axial riblets on
the first whorl and 18 or 20 on the following whorls: axial sculpture crossed by
spiral threads of nearly the same strength as the axials, a fine node is formed
at the intersection of the axial and spiral sculpture; the intercostal spaces each
with a slightly spirally elongated pit, forming spiral series; base of last whorl
sculptured with fine spiral lines; aperture ovate, the anterior portion very
broadly rounded. Length 1.1 mm., maximum diameter .43 mm.

M^-

5b

Figure S. Turbonilla (Pyrgisculus) clip pert onensis Hertlein and Allison, new species.
Holotype no. 37125 (UC), from Clipperton Island. Sa, apertural view; length 1.1 mm.;
5b, detail of sculpture greatly enlarged.

Holotype. University of California Museum of Paleontology, Invertebrate
Type Collection, no. 37125, from Locality B-4241, Clipperton Island.

Remarks. This new species is provisionally placed in the subgenus Pyrgis-
culus Monterosato because of its general similarity to the species described as
Dunkeria paucilirata Carpenter, a species placed in Pyrgisculus by later authors.
The general shape of the rounded whorls, strongly constricted at the suture, and
the strong axial riblets, are similar to those features on that species.

The shell of the new species differs decidedly from the species described by
Carpenter in the more closely spaced axial riblets, in that the .spiral threads are
nearly as strong as the axials and in that pits are present in the intercostal spaces.

Xo. 661 HERTLELX & ALLISON: NEW CLIl'PERTU.X ISLAND GASTROPODS 7

The strong spiral sculpture on the present species is similar to that of some
species of Pyrgiscus Philippi.

Duukeria paucilirata Carpenter (1856, p. 534) was described from a single
specimen from "Mazatlan; 1 sp. off Chama,'" iVIexico. It was illustrated later
under the name of TiirboniUa (Pyrgisculus) paucilirata (Carpenter) by Dall and
Bartsch (1909, p. 129, pi. 12, fig. 10) and more recently as Chcmnitzia (Dun-
keria) paucilirata (Carpenter) by Brann (1966, p. 72, pi. 47, fig. 536). The
spiral sculpture of that species is much finer than the axial and no pits are visible
in the intercostal spaces shown in the illustrations.

Colubraria ochsneri Hertlein and Allison, new species.

(Figures 6, 7, 8.)

Colubraria species, Hertlein and Allison, The Veliger, vol. 9, no. 2, p. 140, 1966.

DESCRrpTiON. Shell elongately turreted, of medium thickness, about six and
a half gently convex whorls, each with a prominent varix, the varices on every
other whorl are rudely aligned in apertural view; protoconch and nuclear whorls
lacking on holotype; about 12 well developed, closely and evenly spaced radial
threads appear on the fourth whorl and these increase to about 28 or 30 on the
last whorl, these are crossed by concentric threads, about five on the fourth
whorl and seven or eight on the penultimate whorl, a double row is present an-
terior to the suture on the last whorl and these are followed by about 20 or more,
evenly spaced but somewhat finer toward the base; a node is present where the
radial and concentric threads cross, the whole forming granular, reticulate sculp-
ture; the interspaces between the radials and between the concentric threads are
ornamented with a few minute concentric striae, about six on the body whorl, sepa-
rated by narrower grooves parallel to the major threads: aperture elongately oval,
posteriorly ending in a narrow notch in the parietal callus; outer lip with a varix,
the inner margin with fine, paired striae, the pillar excavated in the middle, the
inner lip formed by a broad, smooth, white callus which at base projects out-
ward as a narrow plate; the anterior canal is rather narrow, slightly recurved;
color of shell light brown with a darker medial concentric band, above and below
which on the body whorl there is a row of small brown spots, the varices are
light in color but with brown spots sometimes corresponding to the brown bands.
Dimensions of the holotype; length (apex lacking), 29.8 mm., maximum diam-
eter, 11.4 mm.

The protoconch of a paratype, 23.2 mm. in length, is slightly convex, brown,
followed by 2 smooth, glassy whorls, the latter one developing very fine longi-
tudinal threadlets which are crossed by concentric threadlets forming decussate
sculpture, with very fine beads at their intersections. In general appearance,
these features resemble those of Colubraria jordani Strong illustrated by Campbell
(1961, fig. 1).

Holotype, no. 37126 and paratypes 37127, 37129, University of California

CALIFORNIA ACADEMY OF SCIENCES

[Occ. Papers

Museum of Paleontology, Invertebrate Type Collection, from Locality B-4237
(x\-588-l, Allison field station number), beach at Clipperton Island. A para-
type, no. 37128 from Locality B-6101 (A-588-5), approximately 100 meters off
edge of reef flat and 30 meters inshore from outer edge of submerged terrace
(depth, 10-12 meters) opposite United States Naval Observatory marker on
north side of island.

Paratypes also are in the California Academy of Sciences, Department of
Geology, Type Collection.

^*

6a

6b

8

Figures 6, 7, 8. Colubraria ochsneri Hertlein and Allison, new species. From Clipperton
Island. 6, Holotype no. 37126 (UC), length (apex lacking) 29.8 mm.; 6a, apertural view;
6b, dorsal view. 7, paratype no. 37127 (UC), length (apex lacking) 27.3 mm., apertural view.
8, paratype no. 37129 (UC), view of apical whorls approximately X 19.

Remarks. About 25 specimens of this new species, 12 to 30.5 mm. long
(apex lacking on most specimens) have been available for study. There is some
variation in the degree of coarseness of the granular sculpture on various speci-
mens but this may be the result of erosion.

The general form and sculpture of this new species is similar to that of
Triton nitidulus Sowerby (1833, p. 71; Reeve, 1844, Triton, species 70, pi. 17,
fig. 70), originally described from "Hab. ad Insulam Annaa," "Found on the
reefs. - G.B.S." The new species differs from Colubraria nitidula in the smaller
size, less elongate form, shorter anterior canal and proportionately wider aperture.

Colubraria ochsneri, new species, appears to be distinct from any species of
Colubraria described from the eastern Pacific (see Campbell, 1961). The ex-
terior of the shell of the new species bears finer sculpture than the "narrow form"
of C. lucasensis Strong and Hertlein from the Galapagos illustrated by Emerson
and D'Attilio (1966, p. 176, pi. 26, figs. 1-4) and the anterior canal is slightly
shorter. A difference in detail of sculpture is that on C. ochsneri, new species,

No. 66] HERTLEIN & ALLISON: NEW CLIPPERTOX ISLAND GASTROPODS 9

the interspaces between the node-bearing concentric spirals are sculptured with
fine spiral striae of nearly equal size whereas on C. lucasensis the corresponding
spiral striae are somewhat coarser and among them one thread is coarser than
the others.

This species is named for Washington Henry Ochsner who collected marine
shells on Clipperton Island in 1905 during the Expedition of the California
Academy of Sciences to the Galapagos Islands.

Latirus clippertonensis Hertlein and Allison, new species.

(Figures 9, 10.)

Latirus aff. L. socorroensis Hertlein and Strong, Hertlein and Allison, The Veliger, vol. 9,
no. 2, p. 140, 1966.

9 10

Figures 9 and 10. Latirus clippertonensis Hertlein and Allison, new species. 9, holo-
type no. 37130 (UC), from Clipperton Island. Length {apex lacking) 50. S mm.; 10, paratype
no. 37131 (UC), length 42.8 mm.

Description. Shell elongately turreted, yellowish-white covered with a
golden-orange periostracum; SVi whorls present, the apex lacking; whorls regu-
larly increasing in size, concave above a subangulate shoulder, below this gently
convex, the last whorl nearly straight sided and sloping steeply anteriorly; axial
sculpture consists of 9 rounded ribs on the penultimate whorl and 10 on the last
whorl, these fade out toward the base on the last whorl; spiral sculpture on the
early whorls consists of 2 raised cords, 3 on the penultimate whorl and 6 on the
last whorl, followed by 3 or 4 widely spaced ones on the canal; the spiral cords
are faint in the interspaces but where they cross the radial ribs they form
elongated nodes, very prominent on the shoulder; aperture ovately elongate,
inner margin of outer lip with fine spiral ridges, a calcareous node on the parietal
wall near the posterior end of the aperture, columella with 3 plaits near the

10

CALIFORNIA ACADEMY OF SCIENCES

fOcc. Papers

lower end, canal narrow; siphonal fascicle well developed. Dimensions of holo-
type: length (apex lacking) 50.5 mm., maximum diameter, 23.5 mm.

Holotype no. 37130, University of California Museum of Paleontology,
Invertebrate Type Collection, from Locality B-6120 (A-588-25, Allison field
station number), off edge of submerged terrace at a depth of 40-45 meters, on
the north side of Clipperton Island opposite the west end of main washout area.
A paratype no. 37131 and two additional imperfect specimens, from Locality
B-4237 (A-588-1, Allison field station number), shells washed onto the beach,
Clipperton Island.

Remarks. This form, as is true of a number of species and subspecies de-
scribed from Clipperton Island, although resembling its nearest congener, de-
velops shell characters differing sufficiently to separate it from the typical form.
The present specimens are quite similar to some forms of Latirus socorroensis
Hertlein and Strong (1951, p. 76, pi. 26, fig. 8), from Clarion and Socorro islands,
Revillagigedo islands, Mexico, but they differ especially in their more elongate
outline, longer and wider anterior canal and coarser spiral ribbing which is more
sharply nodulose posteriorly.

Succinea atollica Hertlein and Allison, new species.

(Figure 11.)

Figure 11. Succinea atollica Hertlein and .Allison, new species. Holotype no. 37132
(UC), from Clipperton Island. Length 11.1 mm.

Description. Shell elongately ovate, thin, translucent, upper whorls light
amber colored, the last whorl grayish-yellow with irregularly spaced brown lines
paralleling the lines of growth; spire acute, about one-half or slightly less the
total length of the shell; whorls 4V2, broadly convex, sculptured only with lines
of growth and on the last whorl there are shallow longitudinal depressions parallel

No. 66J HERTLEIN & ALLISON: NEW CLIPPERTON ISLAND GASTROPODS 11

to the lines of growth; suture rather deeply incised; columella with well de-
veloped callus which is appressed and decidedly reflected in the middle but be-
coming thinner and gradually disappearing before reaching the base of the inner
lip; aperture elongately ovate about '-> (or slightly more) the length of the shell,
the greatest expansion before reaching the anterior margin. Length of shell 11.1
mm., maximum diameter 5.76 mm., length of aperture 6.47 mm., width of aper-
ture 3.72 mm.

Holotype no. 37132, and paratypes nos. 37133, 37134, 37135, University of
California Museum of Paleontology, Invertebrate Type Collection, from Locality
B-4241 Clipperton Island. Paratypes also in the California Academy of Sciences,
Department of Geology, Type Collection.

Remarks. About 30 well-preserved specimens from Locality B-4241, also
many weathered specimens from Locality 4237, beach, Clipperton Island.

The new species described here bears a close resemblance to the form de-
scribed as Succinca luteola var. subtUis von Martens (1898, p. 331, pi. 19, fig. 2)
from Vera Cruz, Mexico. It differs from the east Mexican form in the slightly
more slender outline, in the narrower body whorl, and in that the aperture is
about one-half rather than five-ninths the length of the shell.

Opeas opai anum (Pfeiffer).
(Figure 12.)

Biilimus oparanus Pfeiffer, Proc. Zool. Soc. London for 1846, p. 34, issued May, 1846.
"From the island of Oparu ; found in earth at roots of Plants (H. Cuming, Esq.)."

Opeas oparanum (Pfeiffer), Pilsbry, Man. Conch., ser. 2, vol. 18, p. 183, pi. 22, figs. 1, 2,
3, 12; pi. 24, fig. 39, 1906. "Hawaii, Marquesas, Paumotu, Society." Bartsch and
Rehder, Smithson. Miscell. Coll., vol. 98, no. 10 (Pub. 3535), p. 16, 1939. Clipperton
Island. Hertlein and Allison, The Veliger, vol. 9, no. 2, p. 140, 1966. Clipperton Island.

Figure 12. Opeas oparamim (Pfeiffer). Hypotype no. 37136 (UC), from Clipperton
Island. Length 9.3 mm.

12

CALIFORNIA ACADEMY OF SCIENCES

[Occ. Papers

Many specimens of Opeas oparanum were taken on Clipperton Island, where
they were living commonly in the protection of rock and plant debris. This
species is rather widely distributed in the Polynesian region.

POmalogyra species.

(Figure 13.)

A few minute, flatly coiled, translucent brown shells of gastropods, some
bearing microscopic threadlets on the periphery, are present in shell washings
from Clipperton Island. These were taken on "the outer portion of the inter-
tidal bench with prominent Pocillopora, brown algae, and slight coralline algae."

13a

13c

13b

Figure 13. ?Omalogyra species. Hypotype no. 37137 (UC), from Clipperton Island.
Maximum diameter 0.425 mm., height 0.18 mm. 13a, apertural view; 13b, basal view; 13c,
apical vie\\'.

These specimens were examined by Allyn G. Smith who pointed out their
similarity in general appearance to some families of land snails such as the Endo-
dontidae. However, in view of the locality, the lack of soft parts and the oper-
culum, he agreed that these shells may be referred questionably to ?Omalogyra
Jeffreys, a genus of minute marine snails.

REFERENCES
.Allison, E. C.

1959. University of California Scripps Institution of Oceanography. "Invertebrates" in
Progress Report. Field Report IGY Clipperton Island Expedition, August-
September, 1958. SIO Reference 59-13, pp. 3-4, May 1.
Bayer, Ch.

1948. Catalogue of the Solariidae in the Rijk.'^museum van Natuurlijke Historic (Leiden).
III. TorinJa. Zoologische Verhandeling, no. 4, pp. 1-44.

No. 66 1 HERTLKIN & ALLISON: NEW CLIPPERTOX ISLAND GASTROPODS \?,

Brann, Doris C.

1966. Illustrations to "Catalogue of the Collection of Mazatlan shells" by Philii) P.
Carpenter. (Paleontological Research Institution, Itliaca, New York), j)]). 1-
111, plates 1-60, .April 1.
Bry.an, E. H., Jr.

1958. Provisional check list of gastropods recorded from the Hawaiian chain. Hawaiian
Shell News, vol. 7, no. 2, Supplement pj), 4-5, December 3.

C.'VMPBELL, B. G.

1961. Colubrariidae (Gastropoda) of tropical West America, with a new species.

Nautilus, vol. 74, no. 4, pp. 136-142, pi. 10; figs. 1-5 in te.xt, .April 6.
Carpenter, P. P.
1855-

1857. Catalogue of the collection of Mazatlan shells in the British Museum collected by
Frederick Reigen. (Published by the trustees of the British Museum), pp. I-
XVI, 1-552. July, 1855 to June, 1857.
Dall, W. H., and p. Bartsch

1909. A monograph of West American Pyramidellid mollusks. United States National
Museum, Bulletin 68, pp. I-XII, 1-258, pis. 1-30, December 13.
Emerson, W. K., and A. D'Attilio

1966. Remarks on Colubraria soverbii (Reeve, 1844) and related species (Mollusca:
Gastropoda). The Veliger, \ol. 8, no. 3, pp. 173-177, pi. 26, January 1.
Hertlein, L. G., and E. C. Allison

1960. Gastropods from Clipperton Island. The Veliger, vol. 3, no. 1, pp. 13-16, July 1.
1966. .Additions to the Moiluscan fauna of Clipperton Island. The Veliger, vol. 9, no. 2,
pp. 138-140. October 1.
Hertlein, L. G., and A. M. Strong

1951. Descriptions of three new species of marine gastropods from west Mexico and
Guatemala. Bulletin of the Southern California Academy of Sciences, vol. 50,
pt. 2, pp. 76-80, pi. 26, May-.August, issued October 15.
Martens, E., von
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